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TRANSACTIO^JS
ENTOMOLOGICAL SOCIETY
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n,g,t,7.dh,G00glc
TRANSACTIONS
ENTOMOLOGICAL SOCIETY
LONDON
1917.
LONDON:
SOLD AT THE SOCIETY'S ROOMS, 11, CHANDOS STREET
CAVXHDIBH SQO&RE, W.,
AND BY LONGMANS, GREEN, AND CO..
PATBRHOSTBR BOW, K.C. ; AND KBW lOHK.
1917-1918.
n,g,t,7.dh,G00glc
DATES OF FUBLICATION IN PARTS.
Part I. (TnASe , p. 1-287, Proc. i-xxiii) wns published 24 Nov., 1917
„II,III1V. ( „ 269-426, „ xxxiii-xovi) „ 16 May, 1918
264401
n,g,t,7.dh,G00glc
ENTOMOLOGICAL SOCIETY OF LONDON
POUKDBD, 1633.
Inoobporatu) bt Rotal OoAnut, 1S86.
PATRON: niS MAJESTY THE KINO.
OFFICERS and COUNCIL for the SESSION W7-19J8.
preelDent.
C. J. QAHAN, M.A., D.Sr.
T. A. CHAPMAN, M.D.. F.Z.S.
G. B. LONOSTAFF, M.A., M.D.
The Hon. N. C. ROTHSCHILD, M.A., F.L.S., F.Z.S.
Qieasuier.
ALBERT HUGH JONES.
Secietarfee
CoHHANDXR JAMES J. WALKOi,^ M.A., R.N., F.L.a
Trb Rbt. OEOROE wheeler, M.A., F.Z.S.
Ifbrnrun.
GEORGE CHARLES CHAMPION, F.2.S., A.L.S
9tbet AemlKM of Oouncll.
A. W. BACOT.
- E. C. BEDWELL,
E. A. COCKAYNE, M.A., M.D.
W. C. CRAWLEY, B.A.
H. WILLOUGHBY ELLIS, F.Z.S.
J. C. F. FRYER, M.A.
S. A. NEAVE, M.A., B.Sc., F.Z.S.
R. M. PRIDEAUX.
A. E. TONQE.
Retident Librarian.
GEORGE BETHELL, F.R.HiST.S.
n,g,t,7.dh,G00glc
( vi )
Xaelneea aii& VubUcations Commfnee.
ROBERT ADttlN.
0. T. BETHUNE-BAKER.
JOHN HABTLET DURRANT.
REV. F. D. MORICE.
. And the Executive Officers of the Council.
ecftleb ttatlonal aommlttce of jentomologlcal DomeiiclatuK.
Q. T. BETHUNE-BAKER.
DR. C. J. OAHAN.
DR. K. JORDAN.
L. B. PROUT.
REV. GEORGE WHEELER
JOHN HARTLEY DURRANT, S«r«ton/-
n,g,t,7.dh,G00glc
CONTENTS.
B7 B.
I Id the Britiah Huaeam. Bj
BowLAND E. Tdbnib,^.Z.S.,P.EB.
III. New or littt^tenmi Hettroctra from Hadacaaou. B7 Sir
Obobob H. Kmucit, F.E.8.
IV. Additjons to tha koowladge of the Citoniidat of Britiah India.
Bj Olitu B. JiHSON, F.B.8
Collece, Cambridge (Lieut. B.A.H.O.)
■ in BritUh Qaiasa. B7
VIL The oondition of the sealea in Uw leaden malea of Agriaia
Aitii, Bott., and in other Lycaenida. Bjp B. A. Coctxtsm,
D.lI.,P.R.C.P.,TemporarT Surgeon, R.N
VHI. On Mv uid little-known Lai/riidat froni Tropical America.
Br Obobob CUBLEa Chupion, F.Z.S
IX. A Reriaion of tbo genua Tarucui. Bv O. T. BcrHDHi-BiiiB,
F.Lil., F.Z.8 :
X. Kotea on mnw BritUb Quluu Himetoptera (eicluaife of the
FarmieiAu). By ti. B. Bodb(M, B.A.. Dip. Agric. (Cantab.),
F.Z.8,, F.B.S-.Ooi'eronient Economic Biologiat, Department
of Science and AgTiciittore, Britiah Guiana •
XL On a Collection ol Butterfliea taken Id Eaat Africa bj Mr.
W. A. Lambom. Bj H. Kltbihoh*!!, M.A., D.Sc. With
notea on the Fitrinat. by Dr. F. A. Diibt, F.R.S . and
deacription of a new form of F. dardaiuu 1 , by Prof. E. B.
PoDLTuH, F.HS ;
iM-,i,A J h, Google
XIII. A PrelimiDBry Catalogue of Britiah Cieidomgidae (Diptrra),
with Kpecial reterrore to the ti&ll-midgf* of Ibe North of
GogLuid. By RicKiBD 8. BlaI•AL^ F.L.S^ and J. W.
HisLOP Habrison. D.Sv. _
ProcMdtDg* tor laiT
Aonual Meeting ...
Balance Sheet
Preaident'a Aildreai
Gsneral Index
EXPLANATION OF PLATES, TRANSACTIONS.
PUtel.
8eepa«« e5 Plate XL
PUte II.
86 Plate XII.
Plate III
92 PUte XIII.
Plate IV
83 Plate. XIV-XX.
Plate V.
S9 Plates XXI-XXIII
PUleVI
100 Sketch Map
Plate* VII-X.
. 1S2
PROCEEDINGS.
Plate A.
See page «ii.
PlaU* B. C
., page >Ll»ii.
Plato D
E
F
„ pigea Ixim-UxT.
Plates a
H
J
„ page* «, iri.
n,g,t,7.dh,G00glc
ENTOMOLOGICAL SOCIETY OF LONDON.
HONORARY FELLOWS.
Marked * hare died during the year.
Klectlon.
I9O0 AUBIVIU.IUS, Professor Cliristoplier, StvcMtiim.
1915 Beblrsk, Professor Antonio, Wa Boiaana, 19, Fireate, Italy.
1905 BoLiTAR, Ignaoio, itfiuai lUKional de Biatoriu naltiral, Hipodrinno,
17, Madrid.
1911 CouBTOCK, Prof, J. H., OiTwli Vniver>itij, Itha/M, New York, U.S.A.
1894 FoRETi, Profeseor Aitgnete, M.D., Ckigay, p}-u ilorgef, Swilierland.
1912 • FREV-QcnsNEIt, Dr. Eniilu, La Bageruit, Gtnive, SwitztrUind.
1898 Orabsi, Profeaaor BdUista, The Universitg, Home.
1915 Howard, Dr. L. O., Nutiontd M^aeum, WaiJtinglon, U,S.A.
1914 Lambere, ProfesHorA., Brvxellet.
1908 Oberthur, Charles, Rennet, lUe-el-VHaiiie, France.
1913 Tian-ShaK8KI, a. P. SemeuolT, VaitUi Oitrot, 8 tin., 39, PtliiHjrad,
Ruaaia,
1911 Wasmann, Fr. Erich, S.J.,Vuil:enhiirg{L.) Iifimlht>Kolhg,Hollattd.
SPKCIAL LIFE FELLOWS.
1916 (1894) MiALL, Louis Oompton, F.R.S., (Cuuscil, 1903,
Norttm Way N., Lelduvorth.
1916 (18B8) Yehburt. Colonel John W., late R.A., F.Z.S., (Cot
1896, 1903-5), 2, Byder-dreel, tSl. Jamet'i. S.W.
Marked * have died during the year.
Marked f hare coiiipouiuUd for their Anuual Siihscripti-ni.
D*U "f _ .
1914 A114IB, E. W,, B.A., Tvr/aiib, Cairo, Egypt.
1913 Ahamb, B. G., 16, Fernahaiv-road, CheUea, S.W.
1877 Adams, Frederick C\<M]HtTOtii,T.Z.S., 50, Athleij-ga,-de»ii, Vidoria
ttrtet, S.W. 1.
u,y,i,Ajh,Googlc
( - )
1902 Adkih, B«naiah Wlutley, TiKnototth, Hopt-park, Bromiey, Kent.
1885 Adkin, Robert, (CoDNCT^ 1901-^2. 1911-13), Hodedm, Meadt,
1904 AOAii, E. A., La Haul, Dominica, B. W. Indies.
1915 AlTAR, K. S. Padmanabha, Trivandrain, Travancore, India.
1914 AtiAR, T. V. Ramakriahna, B.A., F.Z.S., I7t« AgrieuJtHral OoOege,
Coimbatore, 8. India.
1912 Allen. J. W,. M.A., 266, Waie»dtn-laHe, London, N.W. 2.
1911 Ahdersom, T. J., SnUnnotogical Laioralory, Kabeti, Brit. E.
AJrica.
1910t Andreifbs, H. E., 8, Nmih Grove, Uighgale, N.
1899 Andrews, Henry W.. Sttiley, WtUing S.O., Kmt.
1901 Ansino, Williom, 39, Lime Street. E.C. 3.
1908 1 Antram, Charlea B., Somerdait EUale, Ootacamund, Nilgiri EiiU,
S. India.
1913 Abmytaob, Edward 0., Petiroae, Uiiivek, Potindj^tock, Bade.
1907 Arnold, 0., U.Sc., A.B.C.S., Curator, Bhodetia JUtiwum, Bviawayo
S. Africa. '
1899 1 Arrow, Qillrert J., (Codncil, 1905-7), 9, BoBsdoie-roiMf, Futaes,
S.W. 15 ; and BrUidt If.weum (Nalnral Sistonj), (VOTftweU-rood,
S.W. 7.
1911 Ashbv, Edward Bernard, 36, B^thlrode-nxid, Houtulow, MiddUtex.
1907 t AsHBY, Sidney R., 39, Park-lnne. Wembley, Middlttex.
1886 Athorb, E. a., 48, Hightlreel, King's Lijnii.
1913 AviNOFF, Audrii, Liteyny, 12, Petrograd, Svma.
1914 AwiT[, P. R., Medical Entomologist, c/o Qrindlay & Co., Bonkera,
26, WestmoiiaTid-itreet, 0(Jc«((a.
1901 Bacot, Arthur W., (COONCII, 1916- ), York Cottage, York-hm,
Loti^htor^ Sttei.
1904 1 Baonall, Ricijard S., Penthaw Lodge, Penshaw, Ihirkam.
1909 Baowell-Purefoy, CapL Edward, Bail Farleigk, Matditmie.
1916 Balfoor, Miss Alice, 4, Carllon-gardeiu, S.W., and Whittingehame,
PretlorAirk. Scotlaiid.
1913 Ballaxd, Edward, Qovt. Entomologist, Agricidlmxd College and
BeKurch Institute, Cminbatore, Madras, S. India.
1886 Bankea, Eustace R., M.A.
1890 Barclat, Francis H., F.G.S., The Warreti, Ci-omer.
1886 Baroaqli, Marcliese Piero, Fiona S. Maria, Palazzo Tempi No. 1,
Florence, Ikily.
1895 Barker, Cecil W., 244, Esstmcood-road, Durban, Natal, SoMh
Africa.
1902 Barraod, Philip J., Cheater Coltaije. Benhillroad, Svtton, Sitrrey.
1907 Bartlett, n. Frederick D., 1, Myrtle-road, BoMrnenunith.
1894 t Bateson, Prof. Williimi, M.A., F.R.S., Fellow of St, John's College,
Cambridge, The JUiinw House, Meiiou, Surrry
1908 Bayfohd, E. O., 2, Bockiiigham-street, Banidey.
ll,g,l,.eJh,GOOglC
( "i )
1901 Batnb, Antinr P., c o Mevrs. FTeeninn, Oul/c-afrMt, Pramtingkmt,
Snffalk.
1912 Batnes, Edw&rd Stout Augustus IM, fTonndt-ifml, Bettaion-
a^Hare, &W. t.
I896tBUKE, Pn>r. T. Hndson, B.Sc., •F.aS.G.. (V.-Prb., 1910;
COUKCIL, 1909-11), 10, lU^eiU Terraef, Ediubrtr^.
1908 Bkk, Bicbaid, ITecfcVU, r;^ Park. Teoril.
laOo Bbdford, The Dnke of, E.G., Pr«a. Z.a, etc., ira6i(r» AbUy, BttU.
1912 Bedpobd, Oenld, Entomologist to tbe L'nion of South Afric*,
Veterinary Bacteriological Lkboratonr, Oiulttlepvort, Prtloria,
TVaiunxiJ.
1913 Bediwbd, Capt. Hugh Wonen, Churrh FrtU^ HorUy.
1899 Bkdwsll, Ernest C, (Cor\ciL, 1917- ), Bnuj-irn, firigA (on- nxuf,
Cowltdon, iSurrrif.
1914 Benderitteb, Eugene, 11, Rtit SI. Jiirqtin. Le Main. France.
I901 Bbsotsso.n-, Simon, Ph.D., Lecturer, Unir*railii of Land, Sinden ;
Curator, Entomological Collection of the University.
1915 Bbsham, Prof. William BlaiUud, M.A., D.Sc., F.B.S.; I7iiiPfr«(y
of Otago, Dunedin, Xew Ztalaitd.
1906 BBNT»L^ E. E, The TotMn, Beybrid/ff:, S»K:t.
19)3 Bb8T-Gakdhbr, Charlea C, Rooktcvod, ^'rt^lh, Glamorgtw.
1885 Beibune-Baker. Geoige T.. F.L.S.. F.Z.S., (Pres.. 1913-14;
V.-Pres.. 1910-11,1915; CoDNcit, 1895, 1910-I5:., IS.C/arcnrfoH-
mad, EdgbatloK, B'lrtaiitgham,
1891 Blaber, W. H., F.L.S., 34. Crommtirood. Hot*, Brijk(on.
1901 Black. Jamen E., F.L.S., Nelhtrcroft, FttUe.-.
1904 Blair, Kenneth G., S3, n>)( IIUI, Hl<^te, N. 6.
1885 BuTHVAIT, Lt.-Col. Linley, F.L.S., Emjlc Houit, Balheailon,
Balh.
1904 Bliss, Maurice Fredericlc, M.H.C.S., L.R.C.P., 26 WoodrilU-
gardeiu. Eating, W. 5.
1916 Bococc, Charlea Uanslope. The Elms, Ashley, XriDiunkel.
1912 Bodkin, G. C, Govt Entonjologiat. Gtonitlotrn, Britiuli Gniaita.
1903 BoGDB, W. A., Tht Uaiik Hoiite, Watchel.
1911 BolLEAU, H., 99, B'ltde la C>Ue St.. ThiboiiU, Boit de Cofomtw,
iSeiiM, Frantf.
1891 BooTB, George A., P.Z.S., M.B.O.U., The Ho-mihi;-. Kirkham,
LaiKt.
1902 BOSTOCK, E. D., OuJton Cross,. Stone, S(<i/a.
1913 BOWATER, Captain WiUiaiii, 20, Ru*Mllroa<J, Moide;i, Birmingham.
1888 BOWBR, Benjamin A., Langlrji, Willow Grote, Cliistehuisl.
1894 1 Bowles, K Aiigustiw, M.A., M'lihMloit Hmisr, WiiUham Cros/.
1912 1 BowRINO, C. Talbot, Hoihov; Hainan, S. CT.iiici.
1916 Bos, Leonard Charies, F.B.H.9., Dominion Expt rim fatal St.itioH,
Fredericlon, Nea Bruntioictc.
1910 BOTD, A. Whitworth, The Attun, Allrincham, Chesliiie.
1905 Bracken, Charles W., B.A., 5, Carfrae Ten-ace, Upson, Flyno\>l''.
ll,y,l,/eJh,G00glc
( xii )
1917 BRBiJeit, Dr. H. O., Ph.D., Director of the Transvaal Musei^.ti,
PrctoTui, Transnuil, 3. Afrim.
1904 BBiDdKMAN, Commander Tlie Hon. Ricljard O. B., R.N., 44, Lownda-
upiare, S. W. 1 ; and c/o Commander-in-Chief, Capt of Oood Hope
Station, c/o G.P.O.
IBTO Briqgs, Thomaa Henry, M.A., Roek Ilonie, Lyamoiilh 8.0.,N.IkvoH.
1894 Briqht, Percy M., Clierilon, 26, Portcheater-roail, Bovnirmoutli.
1909 Bbittbn, Harry, Myrtle View, Windmill-road, Beadington, Ox(m.
1902 Brouobton, Major T. Delves, R.K, Mhoie, India.
1878 Bbodn, Major Thomaa, Chev. Legion of Honour, Sfoutit Albert,
Aitcl-liiiiil, N(vy Zealanil.
1904 Brown, Henry H., Toioer Einue, 8, BntnUJUJd-terraet, Edinburgh.
1910 Brownc, Horftce B., M.A., Park Htirsl, Morley, Yorka.
1911 Brutzkr, Rev. Henry W.,GceatBotnIen Viairagf, Mai^t Harboro'.
1909 Bryant, Gilbert E., Fir Grore, Edut, Surrey.
ISftSfBuCHAN-HsPDORN, Sir Arcliibnld, Burt, J.P., D.L., SmeaUm-
Ilepburn, Prtitoukirk.
1917 BuLKLny, Dr. George Oranville, M.D., F.S.A., Ryt Croft SouiK,
Maiu:he»ttr-rnad, Bury, Lane*.
1916 BruNlON, Prof. E,, La Lticiolc, Aix-en-Provaice, Vranee.
1907 Bdueid, Arthur, F.S. A., Wimboro, Midtomer Norton, SotnerKtAire.
1896 1 Burr, Malcolm, D..Sc., F.L.S., F.Z.S., F.G.S., A.R.S.M., (V.-Prm.,
1912 ; Council, 1903, 4, 1910-13), The DuUh Cuttage, Woking.
1909 BDERUwa, The Rev. C. R. N., The Vicarage^ Mwhitig, Stanfont-le-
Hope, fissar.
lB68tBCTLBR, Artiuir G., Ph.D., F.L.S., F.Z.a, (Sbc, 1875; CoOMCIL
1876), Tbe Liliei, Beekenhamroml, Becktnham.
1683 BuTLKR, Eilward Albert, B.A., B.Sc, (CouNcn, 1914-16), 14,
Drt^jtds-road, Uurnsey, N. 8.
1902 BuTLKR, William E., ifu^ing Emar, Oxfmd-road, Beading.
190& BUTTBRFIELD, Jos. A., B.Sc., Oi-mesby, 21, DorvUleroad, Lee, S.E.
1914 t BuTTERFiBLU, Roase, Curator, Corporation MvKam, KeighUjf, Yorkt.
I912tBnxToiJ, Patrick Alfred, M.B.O.U., Fairhilt, Tot^ridge; and 40,
Gidogtm Plfice, Londtm, S.W.
1904 Byatt, Horace A-, B.A., The Palace, Malta.
1917 Cambron, Dr. Alfred E., M.A., D.Sc, The Sntoinotogical Branch,
VtjHiHm^nt of Agrieidlurt, 0»ch(v(, tVixurf-t.
1902 Caueron, Malculin, M.B., li.N., 7, Blei«ngltm-road, Lee, S.E.
1885 CAypBELi^ FraiiciB Mniile, F.L.S., F.Z.S., etc., BrynUwyd«yn,
MachytiUeth, Mniitgomerydi'ire.
1898 C'ANDiiZB, L^on, Mont St. Martin 7S, Liige.
1680 Cansoalb, W. D., S«h«.v Hani; Simlh Norieooti, S.E. 25.
1889 Cant, A., 33, Feftingroad, Pu/iwi/, S.W.j and c/u Fredk. Du Can*
Godniaii, Esq., F.H.S., 45, PonZ-rfrerf, S.W. 1.
1910 Carmer, E. Wace, M.D., F.R.S.E., Momiimside, GranvilU-road,
Dorridge, and The University, Birmingham.
ih, Google
( "ii )
1692 Cabpinter, The Hunble. Mrs. Beatrice, 28, (frmrenor-road, S.W. 1.
1910 Carpenter, Geoffrey D. H., D.M., B.Ch., Vgnnda Medical Sereice,
Uganda ProtectoiaU.
1895 Cahpentkr, Prof. Oeorge H., B.A., B.Su., Royni CoUfge of Science,
Dublin.
1898 Carpbntbb, J. H., Bedcot, Belmont-iwid, Leiitherhead.
191fi Carb, Professor John Wesley, M.A.. F.L.S., F.G.S., Professor of
Biolt^y, Vniverritij CtiUfgt, Nottingham.
19ia Carr, William, B.Sc, Station-road, Bentham, Lanctiater.
1911 Uabson, Oeoige Moffatt, Ei)tomolo>;itit to the GoveiDtnent of New
Guinea, Pm't Mortshy, Papuii, rid Avatralia.
1895 Cabtbr, Sir Gilbert, K.C.H.O., GreyHiffe. toirer IVarbeny-road,
Tanfuay.
1912 Carter, Henry Frani'is, AMiHtant Letlurer and Deinonstnitor in
Uedical and Economic Enlonioliigy, LIrerpool Srhool of Tropical
Mtdieitie, Vmrersily of Liverpool.
190fl Carter, H. J., B.A., Ganmrillah, Kintore-slreH, Wnkroongn,
Syditey, N.S. W.
1900 Cakteb, J. W.. 15, WeUfidd-road, Heaion, Bradford.
1889 1 Cave, Charles J. P., DiUham Park, Ptterffieid.
1900 Chamberlain, Neville, WedboMrne, Edgbastoii, Biiiiiingham.
1871 CHAMP10N,GeorgeC.,F.Z.S., A.L.S., Librarian, 1891- (COUNCIL,
1875-7); liuilhereule, llondl, Wokii>g;&nA 45, f.m(-ir(cee(,S.W. 1.
1914 Champion, Harry Geoiye, B.A., As»islnnt Conservator of Forests,
W. Almora, U.P., India.
I89t Chafhan, Thoinas A]){eriion, M.D.. F,Z.S., Vice -Prrsi dent,
(V.-Pbeh., 1900, 1904-5, 1908, IDlti ; Ciiuncil, 1898-1900,
1903-5, 1907-9, 191fr- ), BHida, ReigaU.
1897 CUAn-NBR, Misa EUiel F., Forett Bank, L>jndhurat S.O., Hantt.
1913 Cheavin, Harold S., F.R.M.S., F.N.P.S., The (ilenroy Hotel, S2,
Wobam Place, Ku^M-nquuTe, W,C.
1902 Chbbbhan, E. M.
1889 Cheibtt, William M., M.A., F.L.S., Waltrijalf, Bingiixirth.
1914 CBR¥HT4^ R. Neil, B.Sc., 377, Fimt-arenae, Oltatm, Canada.
1909 Clark, Major C. Turner, F.Z.S., UiUtred, tit. A^guatitt^t-avenuf,
8. Croydim.
1908 Clark, Edgar L., Laureslim, llidije Koad, Overpmi, Vurhan,
Natal.
1914 Clbare, Ii. D,, Dcpt. of Scienoe and Agriculture, Qeorgetoim,
BritiA Guiana.
1914 Cleouorn, Miss Maude Lina West, F.L.S., 5, Alipore-road,
CaleulUi, India.
1908 Clditebbdck, Charles Q,, HeaOi^ide, 23, BeathviUe-road, Glonctaltr,
1908 Clotterbock, P. H., Indian Forest Deparliaent, Naini 2'al, United
Provinces, Inditi.
1904 CoCEATNE, Edward A., M.A., M.D., (Couscil, 1915- ), 16, Cam-
bridge'tquare, London, W. 2.
l„y,l,Ajl^,COOl^lC
< '" )
1917 CocKBRELii, Prof. T. D. A., Uiiirtraity of Colorado, Colorado, U.S.A.
1917 C0CK8, Frederick, 26, Crotim-street, Reading.
1914 CoLEHAN, Leslie C, Dept, of Agriculture, Bangalore, Mgaore, India.
1899 CoLUN, Jauiea K, F.Z.S., (V.-Pres., 1913; Council, 1904-6,
1913-1.^), Silfsfx Liidge, Neicmarket
1906 COLLINGB, Walter E., D.Sc. (St. And,), M.Sc. (Birm.), F.I--S.,
Research Fellow of the University of Si. Aadrewfi, The Galty
Marine Litboraiory, St. Andreu», Scotlatid.
1813 COHEY, MisB Blanohe A., The Poplam, Pwklechiirch, OloucesUrAire.
1916 CoRNFORD, The Rev. Bruce, 13, Bavelock-road, Porttmoath.
1911 Cotton, Sidney Howard, 1a, Oieslerfield-atreet, Maiifair, W. 1.
1913 Coward, Thomax Alfred, F.Z.S., 36, George- street, Manditster.
1895 Crabtreb, Benjamin Hill, Crinyte Lodge, Leventliulnic, Maiichetltr.
1913 Craog, Cajjt. F. W., M.U., I.M.S., i/o MeBsrs. Cos & Co., ffomfey-
road, Madras, India.
1909 Crawlby, W. C, B.A., (Codncil, 1917- ), 29, Holland Parkroad,
W.4.
1890 Cbewe. Sir Vauncey Harpnr, Bart., Calke Abbey, Derby fhtre.
1880 1 Crisp, Sir Frank, LL.B.. B.A., J.P.
1907 Ckopt, Eilward Oetaviua, M.D., 28, Clurtndon-rixid, Leeds.
1908 COLPIN, Millais, M.B,, F.R.C.S., The Pak,ce Hold, Shanghai.
1908 Cdrtis, W. Parkinson, Aysgnrth, P,«Je, Dorse*.
1901 Dadd, Eiiivard Mftrtia(ftu/tfel/e)i Internment Gamj/).
1900 Dalglish, Andrew Adie, 7, Keir-tfrfet, PnUnkshidds, Glasgow.
1907 Uambs, Felii L., 10, LorlzingslTaate, BeHin-Lichterfelde.
1886 Danmatt, Walter, 6V, LaMrenee, Gaitxd-road, Lee, 8.E.
1911 Davet, H. W., Inspector of Department of Agriculture, Mdboume,
Victoria, Awtlralia.
1913 Davidson, Jamea, M.St., Imperial Cdlrge 11/ Science and Teeh-
nolojfy. South E'ensinjrfoii, S.W.
1906 Davidson, James D., 32, Druinsheiigh Gardens, Edinburgh.
1918 Davis, Frederick Lionel, J.P., SLB.C.S., L.lt.C.P., Belize, British
EonduToa.
1910 DAW80.V, William Qeon{e, Manor Hviise, Upper Wi-k, Woi-ctiter.
1903 Day, F. H., 26, Cnrroekderrace, Carlide.
1898 Day, G. O., Sahiahlon, Ditneau') Stalion, Vanconvei- Inland, Britith
Colundiia.
1912 Dkwitz, Dr. John, Devunt-let- Fonts, Mttz, Loi-raine.
1913 Dickinson, Barnard Oimiston, B.A., Beech Bill, Xewport, Salop.
1917 DiCKSEE, Arthur, 24, Lij/ord-road, Wandsirvrfh Coiumon, W. 18.
1675 Distant, William Lucas, (V.-Prb8., 18B1, 1900 ; Sec, 1878-
80; Council, 1900-2), Glenni-Ie, 170, Birc)Mw,er-rv'id, South
Norwood, S.E. 25.
1887 DiiEY, Frederick Aii^^UBlus, M.A., M.D., F.E.S,, Fellow and Bursar
of Wadhani College, (Pre«., 1909-10; V.-Pres., 1904-S, 1911 j
CODNCIL, 1895, 1904-6), Wailham College, Oxford.
n,g,i,.rjh,G00glc
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1909 t DOBBOS', Thomas, 399, Ploddrrlanr, Famtmrih, nr. Bolton.
1905 DoDD, Frederick P., Kuranda, via Oairtu, Qutendand.
1912 DoiG, Capt. Kenneth Alan Crawford, R.A.M.C, M.aC.S., LR.C.P.,
c/o Messrs. Holt & Co., 3, W hiUtuUl-plact, London, S.W. I.
1906 DoLLiiAN, Hereward, Hort Hmttt, Nemton-grote, Btdford-park, W. i.
1903 DoLLHAH, J. C, Bore Souat, NtiBton-grore, Btd/ordparl; W. 4.
1906 DONCABTBR, Leonard, M.A-, Tht Unitertity Mfueum of Zoology,
Cambridge.
1B91 DoHiBTBOBPK, Horace Si. John R., F.Z.S., (V.-Pres., 1911 ;
CoraciL, 1899-1901, 1910-12), DurandtMiorpe, 19, nailetcrtt-
road, Putruy,S.W. 15.
1913 Dow, Waller James, 5, Grtat GAttgestretl, Wtstminster, S.W. 1.
1910 DowhE8-Shiw, Rev. Archibald, Scollon Rictory, Gaiiuborough.
1884 Drocb, Hamilton H. C. J., F.Z.S., (Council, 1903-6), TrtfuiU
Lodge, 3, Nor/M-road, N.W. 8.
1900 Drdkt, W. D., Olamufon, Laton-ivad, Boating*.
1894 Dddokom, O. C, Director General of the Dept. of Agriculture,
iltadi, Cairo.
1913 DrFFiKLD. Charles Albaa William, Stowting Beiiory, Hyttie, and
Wyt College, Kent.
1906 DoKinriBLD Jombb, E., Gattro, BeigaU-
1883 DuRRANT, John Hartley, (V.-Prb8., 1912-13; CollKCl^ 1911-13),
Jfe./™, 17. Buritock'toad, Putney, S.W. 15; and BritiA MitM^m
(Ifalurat Biatory), Cromnxllroad, South Kenainglon, S.W, 7.
1910 Eales-Wbiib, J. Ciishnj, 47, Chetter-lerrare, Butonaqimre, S.W. t.
1912 Earl, Herbert L., M.A., 1 2, AnondaU-road NuHh, Soiilhport, Lana.
1885 Eatoh, Tlie Bev. Alfred Edwin, M.A., (Council, 1877-9), Rich-
mond Villa, Norlhaia S.O., N. Dtron.
1902 Edelstrn, Hdbert M., The Elnu, Forty Bill, E'\/ietd, Middlesex.
1911 Edwards, F. W., KiTtgamar, Comwall-road. Barrow.
1886 Edwards, James, Coleabnme, CheUenham.
1884 Edwards, Stanley, F.L.S., F.Z.S., (Council, 1912-14), 15, St.
aermant-plaee. BUckhmtk, S.E. 3.
1913 Edwards, William H., Nattirtd UvA<ny Xkpt., The Mu»r><m,
Birmingham.
1916 ErPALATOUN, Hiaean, Chftuhrah-arentis, Cairo, Eyifjit, aiul 8.E.
AgrictMural CoQege, Wye, Kent.
1900 EmOTT, E. A.. 16, Bdaist Orere, UompsUad, N.W.
1900 Ellis, H. Willoiighby, F.Z-S. (CovKCI^ 19ie- ), 3. Lauuisler-
}iaee, BeUin Park, N.W. 3.
1903 Eltrihoham, Harry, M.A., D.Sc, F.Z.S., (V.-Pbes., 19U;
CoUKCIL, 19)3-15), IToorf/ioiwe, Slroxid, Gtimceslfrshire ; and Bope
DepartmeiU, Cnirtraity Mvatum, Oxford.
1878 Elwes, Henry John, J.P., P.R.S., F.L.S., F.Z.S., (Pres,, 1893-4 ;
T.-Pbim., 1889-90, 1892, 1895; Council, 1888-90), Coltabome,
Ch^tKham.
iM-,i,A J h, Google
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1914 Ehhftt, Capt. Charles P., 1, High Gliff Villa, Felii»lowv.
1903 Etheridob, Robert, Citrator, A'tttralian Mntenm, Sydniy, N.H. W.
1908 Eustace, EiiBtace Maltabone, M.A., Wi-liingUm CMege, Berk*.
1909 EvANB, Frank J., Superintendent of Agriculture, Calabar, EaMem
Fiwintt, S. Nigtria.
1917 Farquhaksoh, Charles C^ilvie, M.A., B.Sr., Government Agri-
cultural Department, hfoor Plantation, Ibadtin, Nigeria,.
1907 Fbather, Walter, Kihivtzi, Brilith Enat Africa,
1900 Fblthah, H. L. L., MtrriinfUe Buildings, S'lmmands-istrett,
JofianHtibitrg, Traiuroal.
1861 Fbnn, Chflrieo, Evenden Houte, Burnt Anli Hill, I.tt, S.E. 12.
1910 FKslEe, A., M.D., 170, ffoHh Grange Gron-Arenut, Paiadeua,
Califoraia, U.S.A.
1889 Fernald, Prof.C. H., c/o H.T. Fern aid, Esq.. A mherst, Masx., U.S.A.
1900 FiHTB, J. Digby, F.L S., Boi/,' Modei-n Schod, Letdt.
1900 Flbmvhg, The Rev. Canon W. Westropp, M.A., Oxi/fin, FmOaiii, en.
WaUrfoiiI.
1898 Flbtchbr, T, Bainbrigge, R.N., Agricultitml Kesearch IiutiMt,
FiUKi, Bihar, India.
1683 1 Flbtcher, William HuDatid B., M.A., Aldwkk Mamir, Bngnur.
1905 Flokrsheiii, Cecil, 1(1, KetiMngton dturl JtfutMtuiu, 8.W.
1885 FoKKER, A. J. F., ZierXhxee, Zeeland, NelhrniaiuU.
1914 FoHDHAM, William John, M.B.C.S., L.R.C.P.. The TiUa, Bnhrilh,
Selbii, Yorbi.
1913 FoBTKR, Arthur H., M.B.CS., L.R.C.P.(Eng.), M.K.O.U,, Sutiex
Hmue, Hitchiu, HrrU.
1900 FOULKBS, p. Hedworth, B.Sc, Ha^'per- Adunui Agri-mltuini CalUgt,
Neioport, Salop.
1893 FoUHTAINB, Miss Marfjaret, 1727, Wilcnx-ntenue, Lot AngeUt,
Cali/.irniti, U.S.A.
1880 Fowler, Tlie Rev. Canon, D.Sc, M.A., F.L.S, (Pbes., 1901-3;
V.-Frbs., 1903 ; Skc., 18«(i-96), Eiiiiev Virar.ige, near Reading.
1908 Fraber, Frederick C, Caiit., H,U, I.M.S., i/o The Ent. See of
IB96 Frbke, Percy Evans, Soulhpi/iiit, Lima-road. Fotkesloiie.
1888 Fremun, H. Stitart, M.R.C.8., L.R.C.P., 3tarhmth, Nelhtratreet,
North Fiurhley.
1910 Fribbt, G, E., 31, Daniley-road, Oraveaemf.
1908 Frooqatt, Waller W.,F.Ii,S., Government Entomologist, 138, (?«(ti'ge-
itreet, Sydney, New Soiilh Wales.
1891 Frohawk, F, W., Stanley Ilimx, Fark-road, WaUington, Suirey.
1906 1 Frt, Harold Armstrong, F.O. Bux 46, Johanneaburg, Transvaal
Coloiiy.
1900 Frier, H. Forteacne, The Prtoiij, OiatUris, Cambs.
1907 Frter, John Claud Fortescue, M.A., (Codncil, 1916- ), Board
of Agriculture and Fitherttt, NorthfUiAertand'avenue, W.C, 2.
ih,Cooi^lc
J87« Poller, Tlie Eev. Alfred, M.A., 3V Lodgt, 7, SyitfAam-kiU,
S^detAam, S.E. 36.
1898 FuLLEB, Claude, Government Entoniolt^lst, PielermarilzbiiTg, NataL
1687 Uahan, Charlea Joflepb, M.A., D.Sc., Prbsidkkt, (7.~Prbb., 1918;
Sue., 1899-1900; Council, 1893-5, 1901. 1914- ), B, LontdaU-
road, Bt4ford Park, W. 4 ; and Brilith J/nmuhi {Natui-al
Hutory), (7rOTlWPeH-r.Hu/, S.W. 7.
1890 Qabdnib, John, Laarei Lodgty Hart, Weti Harllepool.
1901 1 Oakdkir, WiUoughby, F.L.S., F.S.A., Dtgantoy, N. Waltt.
1913 DB Gavk, J. A., Kin^a College, Lagos, S. Nigeria,
1899 Geldaht, William HortiD, &[.A., 10, Chadiington-Toad, Oxford.
1913 Gibb, Uchlan, 3d, BlactJitalh Park, Btackheath, S.E. 3.
1906»tGiBBa, Arthur Ernest, F.L.S., F.Z.S., (CouNCl^ 1912-14, 1917X
Houndtpath, Upptr Marthormt^-road, St. Albant.
1915 Gibbon, ArtJior, Bntomologieat Branrh, Dept. of AgncuUvrt,
OUaioa, Canada.
1908 GlrFARD, Walter M., P.O. Box 308, Honolvlu, Baunii.
1907 GiLU, U«nT7 Marray, Mead Keeper uf Zoologiral Gardens, Sovih
Perth, W. Aiatralia.
1904 QiLUAT, FnDcis, B.A., Windham dxtb, St. Jama'i-tquart,
PtemdiUy, S.W. 1.
1914 GoDrBET, B. J., Education Dept., Batigkok, Siam.
1866 1 GoDHAS, Frederick Du Cane, D.CL., F.It.S., F.L.S., F.Z.S.,{PbM.,
1891-2 ; V.-Pbeb., 1882-3, 1886, 1889-90, 1903; CodkCTL, 1880-
1, 1900), South Lodge, Lower Betdiiig, HoriAam; and 45, Pont-
itreet, S.W.
1886*tO<x>DRiCH, Capt. A. Mainwaring, Bridington Hmite, nr. BritM.
1904 Goodwin, Edward, Canon Court, Watei-ingbitrt/, Kent.
1898 Gordon, J. G. McU., Coiaemahie, Whauphilt S.O., Wtgloaiisliiri:.
1898 Gordon, R. S. G. McH., Drumblair, Invemen.
18S5 GoRiiAM, The Bev. Heniy Stephen, F.Z.S., (Codncil, 1882-3), High-
croft, Great Malvern.
1913 GoDOH, Lewis, Ph.D., Entomologist to the Govt, of Egypt, Depl. of
AgriniUure, Cairo.
1909 GOWDKV, Carlton C.,' B.Sc., Bioiogmd Laboratory, P.O. Box. 5,
Kampalu, Uganda, E. Africa.
1914 Gravslbt, F. H., The Indian Mtuenm, Cak'Uta.
1911 Grates, Capt P. P., Tnr/auh, Cairo, Bgijpt.
189ltGRE>N, E. Emeat, F.Z.S., (V.-Preb., 1915; Council, 1914-16),
Wai/i End, BewA uwtnw, CanAerieg.
1910 Green, Herbert A., The Cerdral fire Station, Durban, Natal.
1894 Obebn, J. F., F.Z.S., 49, Fictoria-road, Eeneington, W. 8.
1893 1 Grbenwood, Henry Powy^ F.L.S., WhiUsbury Hovae, Sati^Hry.
1886 Griffiths, G. C, F.Z.S., PerAurit, 3, Lagh-r-xtd, Ciiftou, BriMtol.
1894 QRiuaBAW, Percy H., Royal Seottiih Mueeuui, Edinburgh.
1905 Grist, Charlea J., Tht Crofi, Carol Gretn, SerkitciU, Coventry.
1906 Gurnet, Gerard H., Ketwiek Haii, Nonvich.
iM-,i,A J h, Google
19)0 Gdbnbt, WiUiam B., Aut. Gov't. Entomologist, Departfiunt of
AffrieuUim, Sydnty, Australia.
191S Hackkr, Henry, BatttTfieidartet, Botoen Bridge-road, Britbant,
Quetmlandi
1906 Hall, Arthur, 7, Fark-lane-mantion*, Croydon.
1890tHiL^ Albert Eraent, e/a City Librarian, Surrey-Ortet, Sheffield.
1885 Hau^ Thomas William. Slaahope, The Crtieent, Croydon.
1912 Hallttt, Howard Mountjoj, 64, lVe»a>ourn*-Toad, Penartlt,
Qtamorgantkire.
1898 Hamltn-Harkib, R., D.Sc., P.L.S., F.Z.S., FB.M.S., Director of the
QueenaUnd Mnsenm, SL Bonatu, Wilston, Britbone, Aiutralia.
1915 Hanm, Albert Harry, 22, Southfietdroad, Oxford.
1891 HamFSOW, Sir George Fiancie, Bart, B,A., F.Z.S., (V.-Pbbs., 1698 ;
CocxciL, 1890-8X 62, Slanhopt-gardent, S.W. 7-
1891 Hanbdby, Frederick J., F.L.S., BroekhurtI, E. GrintUad.
1905 1 Hancock, JoMph L., 5454, Unieerrity-arvniK, Chicago. U.S.A.
1916 HANirmoTON, Frank, c/o Ueaars. Parry & Co., Madra*, India.
1917 Habding, WiUiam G., A*cham, St. Finrmfs Mead^ BatOionmt.
1903 Hakk, E. J., 4, New-i^uare, LtfKoin's Inn, W.C. S.
1904 Harrib, Edward, 50, WiUon.tlmt, FinAury, E.G.
1910 Hahwood, Philip Bernard, 2, Fern miat, M^ford-road, Sudbury.
1910 Hawkshaw, J. C.
1913 1 Hawkbhaw, Oliver, 3, HilUtreet, Maufair, W. 1.
1910 HBDais, Alfred van der, Stvke Botat, Stoke Mandttaie, Buela.
1910 Hbndkrsos, J., e/o Messrs. Osborne & Cbappel, Ipoh, PerA,
Ftderattd Malay Stattt.
1896 Hekon, Francia A., B.A., 9, Park Hmitt, Highbury Pari; N. 5.
1003 Hirkod-Hkmpball, Williaui, W.B.C. Apiary, Old Bed/ord-road,
Z.ut«n, Btdt.
1908 HBwm, C. Gordon, D.Sc., Dominion Entomologist, Ztept. •/
AfricMltuTt, OttavM, Canada.
1913 Hnrnr, John, B.A., Director, Albany Muieum, GrahamtUwn,
S. Africa.
1913 Hill, Geiald F., OotL Entomologist, Port Ztaricin, VorUent
Territori/, Smith Afittralia.
1876 t HlLLUAli, Thomas Stanton, Ba»(gate-»trttt, Ltttt*.
1907 Hoar, Thomas Frank Partridge, Mercio, Albar\y-road, LetgUoa
B'osard,
1917 UocKIN, John W., CuMetlrtet, Launce^on.
1914 HoDaB,Tbe itev. Prebendary Edward Grose. The Vicaragt, Pad-
dington, W. 2.
1912 HoDOE, Harold, St. Jamc, Maruioni. 54, Piecndaiy, W.
1883 HoDSOS, The Rev. J. H., B.A^ B.D., Jlh[iddi,tgto.i, aiflon DrtW,
Lytham.
1903 HoLB, R. S., c'o aieesrs. King and Co., Boinkiy.
1910 HoLTORD, H. C, Slttead Lodge, Godulminy, Sxtney.
l„y,l,Ajh,COOl^lC
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1B87 Holland, The Rev. W. )., D.D., Ph.D., Camt^t JITiiMum,
PitUbargh, Ptmi., U.S.A.
1898 Holiia!.'-Hd!(t, C. B., Asst Entoniologist, Department of Agri-
caltDK, Knala Lumpur, Federalnl Afnlay Slattt.
1910 HoLVES, Edwud Morrell, itiifAmi, Semtoakt.
1901 HopsoN, Montagu F.,L.D.S.,R.C.S.EnK,,F.L.S.,7.ffor(*y-»(rte(,W. 1.
1897 HORKB, Arthur, Bonn- mi -(ml*, Mnrllr, AberdeaiihiTt.
1903 HocoHTON, J. T.. 1, Purtlond-pdur*, WoThop.
1907 t Howard, C. W., Cbnton CArutinn OoUegt, Canton, Otiaa.
1900 UowEB, W. Geor)(e. 432, George-atntt, Dunidin, .V«((r Zwtand.
1907 HoWLBTT, Frank H., M.A., Ws>notidkam, So}iolk.
1865 t HODD, A. E., 106, Ptmbn>ke~road, Cliftou, Briitol.
1888 HoDSOH, George Vernon, BUI View, Karori, Wellinglon, Nam
Zealand.
1907 HuoBSS, C. N., 178, Oarenre OaU-gankBi, Regent'a Parlf. N.W. 1.
1912 UuiE, Miss Lily, HMyKOod, Cotinion-road, Edinburgh.
1917 Udnter, David, M.A., M.B., The Coppice^ Nottingham.
1897 Ihaqe, Prof. Selw)-n,U.A., (Council. 1909-11), 78, ParUunt-roo^J,
Oamden-road, N. 7.
1913 t Imub, a. D., D.Sc, B.A., F.L.S., Bntomdogiml Dept., THt {7m-
vertUy, Manchtrler.
1906 Ibbv, Col. Leonard Paul, Bvinglon-plact, Aihford, Kent.
1907 Jaci£, Rupert Wellstood, Government Entomologist, Department
of Agriculture, Satitbury, Bhodttia.
1917 Jackbon, MiBB Dorothy J., Stcordalt, Evantoa, Bou-Airt.
1907 JiCtSON, P. H., 112, BaUutm-park-Toad, S.W. 12.
1907 Jacobi, ProtesBor A,, I'h.D., Director of the B. Zoological and
Anihrop.- Ethnographical Uuseum, Drttden, Saxony.
1911 Jacx^bs, Capt. J. J., R.E., 15, Paget-tlrert, OiUingham, Kent.
1910 Jacobs, Lionel L., c/o Algoma Steel Corporation, Sault Ste, Maiie,
Ontario, Giruirfa.
1914 • Jannings, Bev. Frederic S. F., fFarmtreorth Rectory, DotwuUr.
1914 Janbe, a. J. T., \>l-are4!t, Ge:iiw, Pretoria, S. Africa.
ISea Janson, Oliver E., 44, Great Ranell-ttreet, Bloomebury, W.C. 1. ;
and Cetlria, Claremont-road, HighgaU, N. 6.
1898 Janbom, Oliver J., Ce»tria, Claremont-road, Higkgate, N. 8.
1912 Jabdinb, Nigel K., SuntnwrAiK Ho^tte, WiBwioroiijA, nr. Aihford,
Kent.
1912 JKMHKrr, C, Withertdaw Hmise, Wye, Kent.
I8S6 Jbnker, James Herbert Augiistue, Eait Oate Houm, Lewei.
1899 Jknninqb, F. B., 152, SUver-iUreel, Vpptr Edmonton, N. 18.
1909 Jepsos, Frank P., Department of Agrictdt^ire, Suva, Fiji Idaad*.
1917 Jermts, Col. Tnrenne, Highcliffe, We»ion-mper-MaTt.
1886 John, Evan, Llantritant S.O., Glamorganthire.
1D07 JoBRBOK, Charles Fielding, West Bank, Didaburyroad. Heaton
iM:,i,A J 1^, Google
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1917 Johnson, Jaxe, 16 and !7, Mnnton-road, Stafford.
1889 JoBNBON, The Rev. W. F., M.A., Acton UecUiry, PoynU Pan,
CO. A nuagk.
1908 JoiCBT, James J„ Tht HUl, WilUy, Sun-ty.
1888 Josis, Albert H., Trkabuhbr, 1904- , (V,-Pres,, 1912, CoONCiI,
1898-1900), Shrttbtamh, Eilhani, S.E. '
I894tJoRDAK, Dr. K., (V.-Prbb., 1909; CoiiNCii,; 1909-11). The
ifmeum, Tring.
1910 Joseph, E, G., 83, OanricanU-ganleni, W, 2.
1910 Jot, Emeat Cooper, Etxr»lty, Dale-road, Furiey.
1902 Joi, Norman H., M.R.C.S., L.RC.P., Theide, Berkt.
1913 JoNK, WUhelm, 68, Sach*i»che-*tra>»e, Berlin, W. 15.
1876 1 Kat, Jolm Dunning, Leede.
1896 1 Kayk, William James, (CocBCl^ 1906-8), Cai-acas, Ditloii BUI,
Surtitai.
1907 Kbllt, Albert Ernest McClure, Division of Entomology, Department
o( Agricultare, Fntoria, S. A/riea.
1690 Kbukice, Sir George H., Whetttone, Somertel-road, BtigbaHon,
Birmingham.
1904 Kershaw, O. Bertram, IngUside, West Wk/clutm, Kent.
1905 KlINES, John Neville, M.A., D.So., 6, Harvey-road, Cambridge.
1900 Kbts, James H., 7, Whimple-itrwi, Ftyntoulh.
1911 Khdnan, Kunui, M.A., Asst. Entomologist to the Govt, ot Mysore,
BanyeUore, South India.
1912 Kino, Haroltl H., Govt. Entomologisl, Gordon College, Kharioma,
Stutan.
1889 Kino, Prof. Jaraea J. F.-X., 1, AthUe Gnrdent-tenace, Kdvituide,
GlaMgow.
1913 KiRBT, W. Egmont, M.D., MUden, 48, Sutton Court-road, Gkiemrk,
W. 4.
1917 KiRKPATRICK, Thos. W., The De<inriij, Ely, and Room 270, iVnr
Office, WhiMiaU, S.W,
1889 Klapaler, Professor Franz, Karlin 263, Prague, Buhemia.
1887tSLSiN, Sydney T., F.L.S., F.ll.A.S., Ilalheilow, RaglaH-rottd,
1917 KuBBEZov, Nicholas J., The Imperial Aeademy of Srienrei,
Petrograd.
1916 Laiko, Frederick, Natwai Hiilnry Mmenm, Oromtcdl-road, S,W. 7.
1910 Lakin, C. Ernest, M.D., F.R.CS., 2, Park-cretcent, Portland.
place, W. 1.
1911 1 Lamborn, W, a., M.R.C.3., LR.C.P., Zomba, Nyas<d<ind, R Africi.
1917 LAHaHAH, Rir Charles, Bart,, Tempo Mniuir, Co. Fermanagh.
191C Latta, Prof. Robert, D.Phil., Univerntij of Glatgow.
1912 Latoor, Cyril Engeihart, Port of Spniii, Trinidad, BrUith Wttt
IndU$.
1895 Lattbb, Oswald H., M.A^ Oharterhovte, Godalming.
iM:,i,A J 1^, Google
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1899 Lea, Arthur H., GoTemmeut Entomologist, Uwtmm, Addaidt,
S. Aa^ralia.
1914 LXBOHMAK, Alleyne, M.A., F.L.S., F.C.3., Cbrpui ChriUi CoUegt,
Oxford; and SI. Uiibcrt't, Mai»-ilrtel, Georgetoini, Britiih
1910 Leioh, R. S., The Univerniy, Maneheder.
1900 Leiqh-Phiujps, Rer. W. J., Bvrtie yicarage, Bridi/wattr.
1003t Lrvett, Tito. B«T. Thomu Prinsep, Frtnehgale, Riehmond, Yoikt.
1876 Lawis, Geoige, P.L.S., (CouNcI^ 1878, 1884). 30. Shvmcliffe-Toad,
PUktttone.
190St Lewis, John Spedaii, ChoM Fami,GruHfiyTdOntn,Sovlh Barrow;
and 277, Oxford^treet, W.
1892 LiOHTFOOT, R. M., Sooth Afriean M>ueam, Cape Tom*, Capt of
QoodHopt.
1914 LiSTKR, J. J., 8t. John'* OuUege, Cambridge; anil Mttion Houm,
Oranichetter, CatahM.
1903 LlTTLBB, Frank M., Box 114, P.O., LaMKt*Uyn, Tamiania.
1865 1 Llewelyn, Sir John Talbot Dillwyn, Bart., M.A., F.L.S.,
PeulUrgare, Saatuea.
1881 fLt^lD. Alfred, F.C.S., 7'lie Dome, Bognor.
1885tLLoyp, Robert Wylie, ^CoDNCl^ 1900-1), I, 5 and 6, Albany,
Pieeadia^, W. 1.
1903 LOTTHonsB, Thoroa!) Ashton, The Ornfi, Linlhorpe, MiitdUibroagh.
1908 LoNGHDOS, D., The Flower Hmae, Southend, Catford, S.E. 6.
1904t LososTAir?, George Bluudell, U.A., M.D., Vic-if-pRRaiDBNT,
(V.-Pre§., 1909, 1915 ; Council, 1907-9, 1915- ), HigUandt,
Putney Neath, S.W. 15.
1999 LouNflBDav, Cliarlee P., B.Sc., Government Entoniologiit, Box 513,
Pretoria, 8. Africa.
1694 Lowe, The Rev. Frank E., M.A,, Si. SUphen't Vieamge, Otienitey.
1893 TjOwkr, Oflwuld B., Piiutrro, So>Uh Auttnilia.
1901 Lower, Rupert S., DaBonporl-Urraee, WatfviUe, So^Uh Avdraiia.
1909* Lucas, Dr. T. P., WakeJUld-buildinga, Addaide-Hred,, Bi-iabant,
Av^Tidia.
1898 LuCAB, Williaui John, B.A., (Council, 1904-6), 86, KnigM* Park,
Ki aj/glou-on- Tha ma.
1903 Ltbll, G., Gvbome, Fictoi-wi, A\^Hndia.
1912 LiLE, Oeorga Trevor, MayfUld, Limfi^d-iixtd, Cambridge.
1909 Lton, Francis Hamilton, 89, Clarence Oategardene, Upper Baker-
Ureet, N.W. 1.
18S7 M'DouOALL, James ThoniBi, Si. Lawrence, Itle of Wight.
1910 Macdodoall, R. Stewart, M.A, D.Sc., F.R.S.K, EditAurgh Uni-
1900 Mackwood, The Hon. F. M., M.L.C., Colombo, Ceylon.
1899 1 Main, Hugh, B.Sc, (Coo kcil, 1908-10), Almondaie, Buckingham-
road, South Woodford, N.E.
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1914 Mallock, J. Rtusel), Sute Entomologiat's Office, Grbana, lUiioi*,-
U.S.A.
1905 Mallt, Charles Wm., H.Sc., Dtpt. of A^ricvltare, Cape Toien,
5. Africa.
1892 MtDBDRtDQE, WillUm, 4, Niavnch-tXKid, Wavertree, Lirtrpoal.
1894 t Marbball, Alick.
1895 Marbhall, Guy Anstruther Knoi, D.Sc., F.Z.S^ (Codkcil, 1907-8),
6, Che*ter-plact. Eydt Park-tquart, W. 2.
1806 Mabsball, p., M.A., B.Sc, F.Q.S., Vnirertity School of Minet,
Dtiiwiin, New Zealand.
1856«t Marbhall, WilliBin, V.M.H., F.RH.S^ Auehim-ailh, Berley S.O.,
Kent.
1897 Mabtihkad, Alfred H., 16, AmmmorcToad, W. 14,
1910'tMABON, C. W., at. Dmi>, Shafit^ury, Dontt.
1895 Mabbet, Herbert, Ivy-Lea, Bumage, Didtbury, Manchttltr.
1B6B Mathkw, OervoM V., F.L.S., Pay raaater-in -chief, R.N., (Council,
IBSTX /■« HauM, Doverooarl, Harieicb.
1887 Uatthbws, Corfndon, SUi\iu«>ay, Ptynuiock, S. Dtwn.
1912 Madlik, SftmArenda, 211, Pu^cadiUy, W.
1900 Maxwsll-Lbpkot, H., Imperud College of Science and Technology,
SmUh KensingUm, S.W.
1916 Mat, Harry Haden, BiackfTiari Houte, Plymo^Oh.
1913 Meadsn. Louis, Melbourne, Dyke-i-oad, Preiton, Brightem.
16BS Mbltill, James Coanio, M.A., F.L.B., Meole Brace Hall, Shrmr*-
1007 Melville, Mrs. Cathariue Maria, Bedrers, Eaa-road, Saltath.
1914 Menon, J. a, B.A., Trichur, Cochin StaU, S. India.
1867 Merrifield Frederic, (Pkes., 190Li-6 ; V.-Pres., 1893, 1907 ; Sbc.,
1897-il ; Council, 1894, 1899), 14, aiftoa-tei-race, Brighton.
1912 Metcalfe, Bev. J. W., The Vicarage, Ottery St. Mury, Devon.
1880 Mbtrick, Edward, B.A., F.R.S., F.Z.S., ThonJianger, Marlborough.
1883 Miles, W. H., c/o E, Step, Esq., 158, Dora-road, WiaibUdon Park,
S.W. 19.
1913 Miller, F. V. Bruce, Liiingston, N. Rho-iesia, Africa.
1906 MiTFORP, Robert Sidney, C.B., Tlwridea, Weybridge.
1914 MlCAK^, Dr. Tsunekata, The Agricidttiral College, Tokyo Imperial
Univereity, Komaba, Tokyo, Japan.
1879 MoNTBiRO, Dr. Antonio AiiguslodeCarvallin, 70, Rita do Aleeiinar,
I90S Montgomery, Arthur Meadowa, 34, Shaliniar Gardens, Pemhridge-
road. North Actmv,Vf. 11.
1899 Moore, Harry, 12, Loieer-road, Rollierkttbe.
1916 Moore, Ralph Headley, B.A., Heathfield, Plynulock, Devon.
1886 MoroaS, a. C. F., F.L.S., 135, Oahvood-cmirt, Ken»i,ujton, W. 14.
1889 t MoBlCE, The Bev. F. D., M.A., Fellow ot Queen's College, Oxford
(pREfl., 1911,1912, V.-Pbes., 1902,1904, 1913; Council, 1902-4),
Brmuieick, Mount Hermon, Wok'mg.
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ISSft t UOBLIT, Claade, F.Z.S., Monk Soham Hoiue, Suffolk.
1907 MoBTUmB, Charlea H., Boytm Cha^ Bufleti, Stttrty.
ISaS UoBTON, Kenneth J., 13, Blaek/ord-road, EdinlmrgK
I&IO MoeSLT, U«rtin E., 21, Altxaitdra-eowt, Qtutt/t-gaU, aW. T
1900 Hosu, JolioB, 69, BtUow^tratte, BeHin.
1883 MoSLBT, S. L., The Muteum and TechnietU OolUge, Htiddtrsfitld
1911 MOBB, Rev. A. Uiles, i/rim, Wxnderrture.
1907 t MoDtTOs, Capt John C, M.A., F.Z.a, 4lh Wiltohire B^, Port
Canning, Singapore, and The Bail, Bradfottl-on-Amn.
1911 MocNSRr, J. Jaduon, S4, Olmcaint-creicent, fdtnfruf^A.
1901 1 UotR. Frederick, H.S.P.A. BxptrimtHt Siatio», HoHoltdu, Oohu,
1912 1 MULLAS, Jal Phiroishah, M.A., F.L.S., F.Z.S., Professor of Biology,
St. Xatrier't CMege, Lamington-twid, Orant Bowl Poit, Bombay,
IsesfHuLLiB, Albert,F.R.G.S., (Council, 1872-3), c/o Hen A. Hiiller-
Hechel, GTtnsaeherdra$»e 60, A««fe, Swit:triand.
1914 Udrbat, George H., Oovernntmt Station, Kikori, DtUa Divvion
1917 McscHAJiP, Percy A. H., QmrUrhouae SdtooL, Qodalmittg.
IDOg HcBHAM, John F., 48, Brook-dnet, Sdby, Torki.
1903 NuvK, S. A., M.A., B.So., F.ZA, (Codncil, 1916- ), 24, de Vere-
gardent, Kensington, W. 8,
1901 NEVtNaON, E. B., ilmiand, Cubham, Surrey.
1907 Nkwhak, Leonard Woods, Bexleg, Kent.
1913 Newhan, Leslie J«hn William, Bemard'tireel, Clarentont,
If. Jurfralia.
1909 Newstkad, Alfred, The Oronenor ATdantm, Oieder.
1890 Newbtead, Eobert, M.Sc., A.L.S., Hon. F.It.H.S., Dutton Memorial
ProfeasoT of Entomology, I^ School o/' Tropieat Medicine, Univer-
»ity of Liverpool.
1914 Nicholson, Charles, 35, The Avenne, Haie-end, Chittgford, B. 4.
1909 Nicholson, Gilbert W., M.A., M.a, (Codnco, 1913-15), Oxford
and Cambridge Club, Pfdl Mall, S.W. 1.
1906 Nil, John Ashbnmer, Tiigale, CratcUff, Sunex.
1916 NoBIRA, Akio, Tehijoji, Otagigua, Kyoto, Japan.
1914 NoBRia, Frederic de U Mare, The Agricultural Department, Kuola
^umptir, Federated Malag Slate*.
I&1& NoBTHOOTE, Dr. A. B., Blenimim Souse, Monigate, Tort.
1878 NornDGE, Thomas, Ashford, Kent
1895 NCBBB, Lt-Colonel C. G., Timuiorlh BaU, Bury St Edmund*.
1877 ObbbthOb, Een^, Retme* {JUe^et-Vilaim), France.
1893-f-OoLE, Bertrams., SteepU AiUm, Oxfordshire.
1910 Oldakek, Francis A., M.A., The Btd Bouse, HasUnure.
1913 OBinsroit, Walter, KiUupahani, HaldummvlU, Cej^an.
u,y,l,7rJM,G00glc
( xxiv )
1B9& Paob, Herbert E., Berb-ote, OeUaay-road, SU Calherme't Park,
8.E. 16.
1916 Palkbr, Arthur Raj'inond, Jn^hoInM, Norton Way, LetektaoHh,
Etrl*.
1915 Paterbon, Edward J., Fairholme, Oroiobormigh.
1911 Fbarbon, Donglas, ChUvidl Htnue, Chitindi, Notts.
1916 Peebles, Howanl M., 13, Chesham-ttrttt, S.W. 1.
1915 PsiLR, Major Harrj Diamond, I.M.S., c/o Alliance Bank </ SimI,
Ftthaiear, Tndia.
1914 Pendlbburi, Major Wm. J. tor Houti, Broadlattds, iShmcabun/,
and StibU OoUegt, Oxjiird,
1883 P^RtNarBi, Louis, D.Sc., F.Z.S., Director, South African Muteutii,
Cope Town, South Affica.
1903 1 Pkreins, B. C. L., M. a., D.Sc., F.Z.S., Park Hill Bouse, Paignton,
Devon ; and Board of Agriculture, Division of Entomology,
HoBoltiin, HauMii.
1879 Perkins, Vincent Bobert, Wotto«-«nder-Eilife.
1907 1 Perkins, J. A. D., 3rd Seaforth Highlanders, Datienham, Malvern.
139T Phillips, Capt. Hubert C, M.RC.S., L.S.A., 37, Prineet-nquare,
Bayswater, W. 2.
1903 1 PHILLirs, Montagu A., F.R.G.S., F.Z.S., Devon$liire Hmttt Prepara-
tory School, Beigate.
1917 PickardCambbiikie, Arthur D., M.A., Balliol C'uUege, Oxford.
1891 Pierce, Frank Nelson, 1, The Elms, Diiiglr, Limrpool.
1903 PiLCBBR, Colonel Jesse Oeoi^, I.M.S., F.H.C.S., 133, Gltnustgter-
road, Kensingloti, S.W. 7.
1913 PLATT, Ernest Edward, 403, Eistnicmd-Toad, Dmian, Niital.
1885 Poll, J. B. H. Neerworl van der, Drieliergtn, Neiheiianda.
1870 1 PoRRiTT, Geo. T., F.L.S., (Council, 1887), Elm Lea, Ifalton,
Hudtteig/ielti.
1884 1 PODLTON, Professor Eilward B., D.Sc, M.A., F.R.S., F.L.S., F.O.S.,
F.Z.S., Hope Professor of Zoology in the University of Oxford,
(PRBa., 1003-4 ; V.-Pbes., 1894-5, 1902, 1905 ; Council, 1886-8,
1892, I89S, 1905-7), Wykefmm Home. Banbury-road, Oxfoj-d.
1905 Powell, Harold, 7, Rue MireiUe, Hyirei ( Var), Franee.
1908 Pratt, William fi., 10, Lion Gate Gaidens, Richmund, Siirrty.
1878 Price, David, 48, Wetl-etreet, Hor.ham.
1908 Prideadi. Robert M., (Council, 1917- ), Woodlands, Brasled
Chart, Strenoaks.
1904 Prisrb, Richard A. R., 9, MeWovnie Jtfnfie, Wed Ealing.
1893 Prodt, Louia Beethoven, (Col'HClL, 1905-7), 84, AlbtH-road,
VaUtott, E. 8.
1910 PoNNETT, Professor BegiRsld Crundall, M.A., Caius GMege,
Cambriilge.
19O0 Bainbow, William J., The Auntralian M^ienm, Sydney, N.S. W.
1918 Bait-Smith, W., Hollybrook, Rose Eeyitortkroad, AUrtUUry,
Monmouththire.
ih,Cooi^lc
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1913 Rao, B. Anuitbasiraiikj, Oundor 0/ Ihe Gonmtnent Mmtntm,
BangaioTt, India.
I»16 Rao, Telaali R«m«r.handim, M.A., Asat. Govt EntomologiEt,
Agnctilhiral OoU^ye, Coiminlore, India.
1907 Ratwakd, Atthor Italic, 91 and 93, SoiUkvati-Mrttl, S.E. 1.
1898 Hbutkr, PnifeBaor Enilo, Helniu^on, Fintaint.
1910 DB Rai-PHIUPK, G. W. v.. Chief Eiaminer of Accounts, North-
Westeni Ry., Abholt-road, Lakare, l¥tdia.
1912 Riley, Kurman Deab^h, 94, Bratf/UU-mnd, l'pi>tr Touting,
&.V. 17. ; and BriliA Mturnm [Xalural Hiaurif}, S. KettnHfftw,
&W. 7.
1906 RiPFON, Cland«, M.A., 28. WiOtonilrrtt, Ox/otd.
1917 ROBBRTS, A. W. Rjnier, M.A., Bolhantdttt Ii/pfriment.U Stiilum,
Harpemdtn,
19(A Robinson, Herbert C, Oiraior of Stale AfiiKHm, Kiutln Liimfnir,
1901 Ronrssos, I«dy, n'ork*v Manor, NoU*.
1869 1 ROBissos.DocoLAS, WillUm Douglas, M.A., F.L.S., F.aO.S.,
(hnJtarrllon, CatlU Doitglat.
1908 RoGBBS, The Rev. K. SL Aubjm, H.A., Cfaorch Mistiooar; Society,
Mombaia, Briti»h Had Afrita.
1912 RoHBir, Kurt, Baron, ZoiAo^\t, iStoatniHimjMnif, MnnitK.
1907 RosENBERo. W. F. H., 57, IhiTttaoA-tm, N.W. 3.
1868 RoTRHKt, Geor^ Alexander Jiiiues, PnnJxiry, IWor-nxxt, Vp^tr
Noraond, S.E.
1888 t RoTHacliILD, The Rigbl Honble. Lonl, U.Sc, F.R.S., F.L.S., F.Z.S.,
(Council, 1900), Zoulogieal Miueum, Tritig.
1894 t RoTUs<;alLI>, The Honble. Nathaniel Charles, H.A., F.L.S., F.Z.S.,
Vicb-Prkbidbnt, (Pkek., I91&-16 ; V.-Pres., 1914 ; Coiscir,
1901, 1013- ), Arnnd^-honse, Keitiinglon P<tlart Oardms, W.8.
1890 RouTLKiKiB, G. B., Turn Lodge, H^^adi Xook, CailiiU.
1913 RowDBN, Alfred Oliver, 3, Ardiibaid-road, Exeter.
1887 Rowland-Brown, Henrj-, M.A., (V.-Prbs., 1908, 1010; Sbc.,
1900-10 ; Council, 1914-16), Oxhey-gr^re, Harrow Wtatd.
1910 RtiDaK, Chaclea Henry.
1892 KussELL, S. O. C, M<mk; Wood, Henthrrnidt, Park-rond, Wi,kin<j.
1905 St. QuiNTIN, W. H., S<ampUm Hall, Jmiugtoa, York.
1906 Sampson, Cobnel F. Winn, Cotuinandant Prisoners of Wnr (Officera)
C^nip, Di0'ryn AM, lAantantuin, Abergdf, X. WiiUs, and
115, TanntJUld-road, SydenJuiin.
1010 Saunders, H. A., Druokfitld'himM, Simiuuje.
1901 ScaAva,Vi.,V.ZS.,U.S.NaliotudM'wum,WaAiiiylon,D.C.. U.S.A.
1007 SCHMAHSHANN, W., Betttah Lodge, London-nad, Biijield, N.
1912 ScHDNCK, Charles A., Eu.'dme, fyallinfiford.
1881 * ScoLLiCK, A. J., Eimsimod, B, M<ddenn>ad, Xew Matden.
191 1 ScOEBR, Alfred George, UOi Crtd, ChUvyoHh, Q»Mford.
l„y,l,Ajh,COOl^lC
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190» Scott, Hugh, M.A^ F.Ua, Curator in Entomology, Vnitemty
JIfiueum of Zoologij, Camiyridge.
1911 Scorr, Percj William Affleck, Chinese Imperial Ciutoms Service,
Hatufluno, China.
11)12 Seitz, t)r. Adalbert, 59, Bttmarekdraue, DarmOadi, Gtrmany.
1911 Sblocs, Cothbert F., M.D., M.R.C.S.. L.R.C.P., SUa/ord, Pain
Bin, Parktloie, Dortet.
1911 t BBN!trrr, Noel StaotOD, 24, d« Vere-ynniett*, Ktnsinglon, W. 8.
1882 Sharp, David, U.A., M.B., F.R.S., F.L.S., F.Z.S., (Pbks., 1887-*;
V.-Pbku., 1889, 1891-2, 1896. 1902-3; See., 1867; CouMcn,,
1893-5, 1902-4), Laumtidd, Broekaihtinl, HnnU.
1902 Sharp, W. E., (Codscil, 1912-13X THe JSiinjjaioir, Crowthwiu,
Berk$.
1915 Sbaw, Dr. A. ElanJ, to R. Kelly, Esq., Solicitor, 59, Siaintton-
ttnel, MHboume, Vklima, AiutmlUi.
1886 Srait, George T. (Librarian of the Liverpo')) Free Public Library),
WiUiaifi Broien-Mtrat, Liverpool.
1905 Shbldos, W. Ueorge, youlgreave, SofdK Croydon.
1900 1 Shbpbeard-Walwtk, H. W., M.A., Dediehinnie, Kadry, Surrey.
1887 1 SlCH, Alfred, (Oouncil, 1910-12), Cornqf Hovm, Chiguiietr, W. 4.
191 1 BlMEB, James A., Mon Rejio, MoiMuim't-lant, Woodford-grttn,Sutx.
1901 SlUlloSDs, Hul^rt W., 12, are-/i ChamUrt, Court HouteAaite,
AucUiiJtd, New Ztnlaiid.
1913 SiTWELI,, Capt. F., Woottrr, Norlhimbtrlaad.
1902 Sladeh, Frederick William I^mbart, Depl. of Agricultvrt, Centrai
ExpeTimtnlal Farm, Oltt.ty,, Canada.
1902 Slopkr. Gerard Orl.y, F.Z.S., J.P., Badminton CTufc. Piixaditty, W. 1.
1907 Sly, Han.lil Baker, Oxford Hook, Eail-drire, Brighton.
1906 SlIALLUAN, Raleigli 8., Eliot Lodge, Albemaiie-road, Beckenham,
Ktnt.
1916 Smart, Capt. H. Douj^la!', H.A.M.C^ Shttlry, Hnddenjidd.
1915 Smith, Adam Charlea, Norton, Momington-road, Woodford Orten.
1901 Smith, Arthur, Count ir Miueam. lAneoln.
1911 Smith, B. H., B.A.. Edgekill, Warlingham, Stirre-j.
1912 Smith, Roland T., 131, Q.ieei.'s-rw.rf, Wimbledon, S.W. 19.
1898 SoPP, Erasmus John Burtjew, F.R.Met.S., 34, FemdaJeroad, Hort.
1B85 South, Richard, (CofNCII, 1890-1), 4, Mtii>fab,inj-eourt, Shoot-up
Hat, Bnmde^wy, N.W. 2,
1916 Sowerbv, Lieut. F. \Y., R.N.I)., Ctedhorp-.', Lincdnshirt.
1908 Sfever, Edward R., Ridgehnrd, Shenley, Htrtt.
I889»STANr)EN, Richard S., F.L.S., (Council, 1906), JVeiriyn, Bomrey,
H-i,d,.
1910 Stanley, The ReT. Hubert George, Marshfitld Vicarage, Cardiff.
1898 Stareh, C. L. B., M.H.C.S., L.R.O.P., TTie Lh,>et,S,caidty Junetiim,
Kent.
1898 Stbbdino, Htiiry, Chasfieood, Ktmtxd Oak-road, Wei/bii'lgr.
1910 Stestos, Rupert, St. Eduyird'i, SI. Mary Church, Torqtioy.
l„y,l,Ajh,COOl^lC
{ sxvii )
1910 SiONEH&u, Hngh Freilerick, Capt Ut Batt. K Surrey Begt,
StoneUigK, Beigate.
1913 Storki, Gilbert, Dept. of AgncydtHre, Caii-o, Eg^fpt.
1915 Stott, Charles Ernest, Eaton, Beigate.
1896 Stbicku-ND, T. a. Geralii, SotUhcotl, Fovlton, Fairford.
1900 Stcdd, E. a. C, P.O. Box 906, Vancourer, BritiA Columbia.
1895 Stodd, E. F., M.A., B.C.L., Oxion, Extltr.
1906 SwiZRSTBA, Corn. J., Ist AssiHtant, Transvmd Mtueum, Pretoria.
18e4 SwiSHOK, Colonel Charles, M.A,, F.L.S., F.Z.8., (V.-Pbim., 1884;
Council, 1891-3 ; 1902-4), 4, aunUritont-roail, iVtit Kensing-
(o», W. 14.
1894 SwiNHOB, Ernest, 4, QuuUivtone-road, Went Ketiaingion, W. 14.
1876 SwiNTON, A. H., Oak Villa, BraiAJietd, Romsey, Banti.
1911 SwiN>-£RTOK, C. F. M., Oitngjotgana, Mel*etUr, S.-E. Ehodtiia.
1910 Tait, Rnlit, jiinr., Boaentalh, Harboroti^road, AAton-on-Merity.
1908 Talbot, G., MonplaUir, Womiletf, Surrey.
1916 Tatcbell, Leonard Spencer, 43, Hiyrall Hall-road, WaruUad, N.E,
1911 Tadtz, p. H., Craidtigh, Pimitr, Hiddltiex.
1911 Tayi^r, Frank H., Aunlralian Institute of Tropical Medicine
P.O. Box 207, ToivmvilU, Qjitendand.
1903 Taylor, Thomas Hnrold, Si.A., Ywt>hire CoiUge, Ltsdi.
1914 Tempkrlby, Reginald, c/o Dr. W. Qrlffitb, 43, Park-iquan, Lttds,
snd L'AuroTt, Vm>ey-la-Tmir, Vaiid, Sailierland.
1910 Thkobald, Prof. F. V., M.A., Wye Ourt, Wye, Kent.
1901 Thoupsoh, Matthew LawHon, 40, Goifonl-tlreet, Middleibrough.
1892 Thornlef, Tlie Rev. A., M.A., F.L.S,, " Ewglunden," Coppict-,-oad,
NoUmghtim.
1907 TiLiTARD, B. J., M.A., B.Sc., F.L.S., Liniiean Macleay Fellow in
Zoology, Knraiida, Mnvnt Errington, Hornitty, Neto South Walea.
1911 Todd, R. Q., 54, Horntey-lane, Highgute, N.
1897 TOMLIN, J. R. le B., MJi., (CoUNCIi. 1911-3), Lakefoot, Hamilton-
road, Beading.
1907 TONGE, Alfred Ernest, (Cocncil, 1915- ), Aincrofi, Beigate, Surrey.
1914 DE LA Torre Bdeno, J. R., 25, Broad-itreet, New y<irk, U.S.A.
1907 Traoardi}, Dr. Ivar, The Univernty, Upiala, Sweden.
1900 Tclloch, Col. B., TIte King's Otm Yorkiliire Light Infantry, o/o
Messrs. Cox & Co.. 16. Charing Cro>», S.W.
1895 1'uHALBY, Henry, CattlelvH, Searle-road, Famham.
1910 TuRATi, Conte Emilio. 4, Piazia S. Alemindro, Milan, Italy.
1898 Torher, A. J., M.D., The Manor War Hoapilal, Epiom, and
Wiekhant Terraee, Briebane, Aiietralia.
1893 Tcrner, Henry Jerome, (Council, 1910-12), 98, Bral-efell-road,
New Orou, S.E. 14.
1906 Turner, Rowland E.,{Council, 1909-10).
1916 Tytlkr, CoL H. C, c/o Mrs. Tytler, Messrs. Griiidliiy & Co.,
Parliament-etreet, S.W. 1.
ihyCOOt^lC
1893 Uiticir, Frederick William, C.M.Z.S., Port of fi^in, Trinidad,
BrilM Wtat Itidia.
1904 t Vaughan, W., The Oid Sectary, BwkingUtn, Bath.
1914 Veitch, itoberi, Entomologist, Natom, Nadi, Fiji Idanil*.
1909 ViDLER, Leopold A., The Oimwiitt Stone Hotue, Ryt.
1911 VlTALIS DE Salvaza, R,, Vientiane, Loot, Indo-Chttui.
1895 Wachbr, Sidney, F.R.C.S., Dane John, Canterlury.
1897 WitHWRioHT, Colbmn J., (Council, 1901, 1912-14), 45, Handtteorih
Wood-roatl, JiamUwiirOi, Birmiaghiuu.
1878 Walker, James J., M.A., ILN., F.L.S., Secbbtary, 1905-
(V.-Pees., 1916; Cocncil, 1894 ; Sac. 1899), Aorangi, Lannlale-
road. ISuiiimerloicn, Oxftn^i.
1012 Wallace, Henry S., 6, Kayllroad ViOai, Sunderland.
1914 Walsh, Mrs. Maria EmeaiTi&, Soekaioemi, Java, Dutch EaH India.
1866 t Walbihhhah, The Rifbt Honble. Lord, (Pres., 1889-90 ; V.-Prks.,
1882, 1888, 1891-2, 1894-5; Co^)NCl^ 1896), Briiiah Miueum
iNatural Uistvry), CromictU-road, S.W. 7.
1910 Ward, John J., Rnginiirbe Home, Somtrtet-road, Coventry.
1908 Warren, Brisbane C S., Hotel Moy, Oberlwfen, Lac de TImine,
Sioitierlunrf.
1901 t Watkrbocsb, Giistaviia A., B.Sc, F.C.S., AUonrie, Stanhtipt-road,
KiUara, JV<io South H'nle*, Aiistialin.
1914 WATERhTON, Itev. James, B.D., B.Sc., 32, Itlaudfmd-road, Beil/ord
Park, W. 4.
IflU Watt, Morris N., St. John') BiU, Wangtmui, Neto Zetilatid.
1893 Wkdd, John Unoper, 89, Dulwkk filta, DuUick, S.E. 22.
lH76t Wkhtkbn, E. Young, '27, Pemhrid^ - aqiiarf, Nolt'mg HUl
Uaie, W. 3.
190G Whkeleb, The Rev. GeoiBe, M.A., F.Z.S., Skckkiari, 1911- ;
(V..PREfl., 1914), 37, aioiK'ealer-pUice, W. 1.
1910 IVhitb, Edward Dnrlon, M.R.C.S., Canlif City Mental Ho^pilal.
Cavdig.
1913 f Whitley, Purcival N., Bi-anlvxiod, H'di/x^; and Nem CoUrge,
Oj-furd.
1913 1 Whittakbh, Oscar, Oi-mulale, Atlilatuh, Aslilon-nimn-Mersey.
191 1 Whittinohah, Rev. Canon W. O., Gtiist'/n Eextory, Uppingham.
1917 Wkkhaii, Rev. Prebendary A. 1'., Eatt Brent Viairage, High-
bridyr, Sonieraet.
1906 WiCKWAR, Oswin S., fhiirlemmil, Gregory-road, Calumho, Cei/on.
1903 Wiggins. Clare A., M.K.C.S., Eiitrbbe, Uganda.
1896 WiLKMAN, A. E., Thatehed Howe Club, at. Jameif- street, S.W. 1,
1910 WiLLcocKS, Frank C, Entomoli^ist to the Kliedivinl Agrictiltural
Society, Cairo, Egijpt.
1911 WiLLUM8,C. B., M.A., JW((./5pat«, rriiiido./, and 20, Saiey-rood,
Birkenhead.
ih,Cooi^lc
( xxix )
1015 WiLLlAMB, Harold Beck, 131, ^wnVnxid, IF"imW«(oti, S.W. 19.
1915 WiSK, Albert F., Lihmnj :/ MrGiU UitirrrsUij, WtMmouiU,
MoatrtiA, Cnnadn.
1894 WOLLKI-DOD, F. H., JtfiH«»nVU P. 0., Aifftla, X.W.T., Oiiio'la.
1905 WooDBRiDCK, Francis Charles, Briar Clax, Latchmort-imtiiw,
Gtrrar<P» Oojw S.O., BwU.
1914 WooDFOBDE, FrsDcis Oanlew, B.A., 2, Jus alrefl, O.rford.
1912 Woodruffe-Peacock, Bev. E. Adrian, F.LS., F.(5.S., Cmlney
Vifarngt, Brigg, Linenliuhire.
less VocDALB, William Henry, F.R.M.S.,31, BtUe lAtalreet, \Voi1ii«glon.
n,g,t,7.dh,G00glc
ADDITIONS TO THE LIBRARY
DuBiNO THE Year 1917.
AiNSLLB (G. G.)- fSeo Luqikbili (P.).]-
Atein (E. E.) and BicoT (A ). The/clstion between the batching of the
egga Bud the developmeiit of the larvae of Sltgomyia fasriala
(Jed/s ealopai), aod the prcMDce of bacteria and yeiala.
[Pataiitologf. Vol. IX, 1917.] The Jvtkors.
Back (E. A.) aad Pehberton (C. E.). The Meloa Flj' in Hawaii [Bacfro-
etra euatrtilat, Coq.).
[n. S. Dept. Agric, Ball. No. 491, June 1917.1
U. S, Dtpt. Agric,
BlCOT (A,). The Use of loieclicideB ugalnet Liee.
[Brit. Med. Journ., Sept. 30, 1916.]
A simple means of BKertaiDing if a sterilising hut is hot enough lo
destroy lies and uiti in clothing and in blankets.
[Brit. Hed. Jouru., Aug. 4. 191T.]
A contributioa t« the BioDomics of Ftdicului hiimanvi (ttilimeyili)
and Pedinilti eapitiM.
[Parasilologj, Vol. XS, IBH.]
The LoUM Problem.
. [See Amin (E. E.).]
Baub(A. C.) and Davidson (W. H,). A further contribution to the study
of £r(o»ina pyricol/i. the woolly pear Aphis.
[Jonm. Agrlc. Besearch, Vol. X, July 1911.]
['. S. Dift. Agric.
[See QuAiNTiucE (A. L.).]
BiVHELEN (J. F. ran). The colour pattern on Diptera wings,
[Koninkl. Akad. Wet. Ameterdam, Vol. XIX. Nos. 9, 10, 1917.1
The Avlhcr.
BoLTOH (H.>. The "Hark Stirrup" collection of Fossil Insects from the
Coal Ueaiures of Comroentry (Allierj. Central France.
[Reprinted from the Memoirs Manchester Museum, Vol. LXI,
No. 2, 1917.] O. C. Champim.
BttTINO (A.). A Generic *;nopsia of the Cociinellid larvae in the United
»Ut« National HaBcuni, itith a description of the larva of
Hgptraipii biikotata. Say.
[Proc. 11. S.Nat, Mui,, Vol, LI, Jan. 1917,]
The SmHkioeian lnstilulim.
BsofH (T,). Descriptions of new genera and species of Coleopten.
PartV.
[New Zealand Institute, Bull, No. 1, June 1917.]
Tit A«thor,
BuBOBSS {A. F.) and ColItns (O, W,). The genua Cilesoma, including
studies of seasonal bistories, hshita, and economic importance
of American species north of Mexico and of several introduced
■peciei
ih, Google
Bdru(H. B.)- PUt-head<dboren(Bui>r«stidlarTie}BffeclipgfoTe(ttr
'd the UnitFil 8' ■
[U. 8. Dept. Agric, Ball. No. 437, J«d. 1817.1
. lit pi. Ai/ric.
CkRPEKTKB (0. H,)* iDJDTioui InKrU iDil othrr AoimtU obwrvtd id
I
Tht Anthar.
CstKpiOH (O. 0.)- ColeoptftB, Heteremer* (eicluciing TeDebrionidae) fiom
.. ~... , ..... . . -undAldBb™.
,., Ser. e, Vol. SIX, Feb. 1917.)
n NfUaUud.
A new Barid from s Coata Rican bromeliad.
[EoL HoDtUr Ha«.. Vol. LIU, 1911.]
Notes OQ Tropicftl Ameiicau Lasriidae, with descriptions of new
[Est. MoDthly Mag., Vol. UIl, 1917.]
New Xylophilii)* fiDDi Aaitrelii, India aud Borneo.
[Eot. HoDtbly Hag., Vol. LIU, 1S)7.]
Notes OD the Coleoptera recorded from " Retin Animd" bj tlie
Rer.F. W. Hope, (11,(21.
[Eot. Honthl; Htg., Vol. LIU, 191T.]
On new and little-known lagriidat from tropical America.
[Trana. Ent. &oc Load., 1917.] Tkt Anlktir.
CHrmKDBH (F, H.). The bone-ndiah Fle»-beetle {Fhyllolrtla amioraciai,
Koch) ; ita lite-hiatoi; and distribution.
[0. 8. Dept. Agric, Bull. No. 53a, June 1917.]
(f.S. Dipt. Agric.
CntTnaL (R. N.). The Wertem Cedar-borer {Trachvkth Bp,),
[Afrit. Qaiette o( Canada. Vol. V, Nov. 1917] •
fanad. Dtpt. Agric.
ClkhmmT (O. B.) and Heinro (W). Control of the Gipsy Math by forrst
management. Part I : The Cip*; Moth in ^yuads. lij G. B.
Clement. Part II : Management of typical wocxl lots iafeitnl
with the Oipay Moth in the Wbile-pine region. By Wiliia
Munro.
[C. S. Dept. Agric, BoU. No. IM, April 1917]
U. S. Dipl. Ayric.
CoaD (B. R.). and HcQiheb (T. P.). Collection oF WecTiis and infenled
Suareg aa a mean* of control of the Cotton Boll-WecTil in the
iwiiHippi.
[XJ. 8. Dept. Agric, Boll. No. 594, Oct, 1917.]
V. S. Dtpi. Agric.
CoCEBSCLt (F. D. A.). Some Beei from Aaatraiia, Tasmania, and the New
Hebcidea.
[Proo. Acad. Nat. Sci. Philad., 1918.]
n,g,t,7.dh,G00glc
( xxxii )
CoatBRBLt (F. D. A). TbB Centinid Bee* oF (he Philippine IsIsikIb.
[Philippine Journ, Sci„ Vol. XI, IfllO,]
A collection of Bees from QueeDsUoil.
[H«moin Queenilaad Mua., Vol. V, July 1916.]
Tbe Fbuiu or Boulder County, Colorado, InsecU. Ill, IT.
[ViiiT. Colorado Studies, Vol. Xlt, No. I, Jan. 1911.}
— — ^— New Tertinry Insecti.
[Proc. U. S. NBt.Miu.,Vol. LIl, Fob. 1817.1
The Dawn of Uintory : k dramH in three acti.
[Amer. Mmeum .louro.. May IPIT.]
DescriptioiK ot Foaii) Inseda.
[Proc. Biol. Sac. WMbiogtou, Vol. XXX, Maf 1911.}
Some Foiwil TmectB from FloiiBhaat. Colorado.
[Proc. U. S. Nat. Mua., Vol. LIII, June 1917]
Some Beea of tbe ^ous Pmtnythin ;Hyin.).
[Ent. Newa, Vol. XXVIIl, July 19i:.]
. — — Arthropodbin Burmeae Amber.
[Psyche, Vol. XXfV, No, 2, 1817.] Tlu A«lk<.T.
CoLB (A. C.) and iHHg (A. D,). Report on m infestation of larvae of the
Antler Moth (Chiiracai gramims, L.) in the Peak District.
[Journ. Boant Agric, Vol. XXIV, Aug. 1817]
Dfpt. Agric. fntom.. ManeknttT.
COLIIHS (C. W.). [See BrnoMS (A. F.).].
C01IB3 (A. F). Notes on a collection of Odonata from Schoolcraft County,
Michigan.
[OccBS. Papers Mua. Zool. ITniv. Micbigan, No. 41, May 1917}
fBi'f, of Michigan.
CtJMHINOg (B. F.). The Louse and its rdation to disease, its iife-bistorj
[British Muaeum (Natural History), Economic Series, No. 2, 1815.]
>— — — The Bed-bug, its habits ami life-bistory, and how to deal with it
(Britiah Museum (Natural History). Economic Series, No. 6, 1911.1
■JhiT,T.Hfa£Fit.3l«i.
CuBBiE (B. P.). Oomphut parrid/iu, a new speciea of Dragon-fly from
Maryland.
[Proc. U. S. Nat. Mui., Vol. LIII, June 1917]
Thi SiiiithfORiaH InMution.
CDaaHAN (R. A.). Eight new spncica of reared Ichneumon- Bias, with notes
# on some other speciea.
[Proc. U, S. Nat. Mus.. Vol. LIII, Aug. 1817.]
A revision ot Hymenopteroui insects of the tribe Crrmaatiiii o!
America north of Mexico.
[Proc. U. S. Nat, Mua,, Vol. LIII, Aug, 19IJ,]
Thi Siititksoiiian loaliltlioa.
DaTioauN (W, M,), [See BiKKn (A. C,),]
D'Ehicebbz i>b Chahkoy (D.). Uotli-horers affecting sugar-cane in
[Dept. Agric. Mauritius, Scient. Series, Bull. No. S, 1817.]
Tht Author.
DiSTiNT (W. L.). Rbyiiohota, Part II : Sub-order Homnptora. Percy
Slulen Trust Expeilition to Indian Ocean in lUOS.
[Trans. Linn. Soc. Loud., Zool. 2nd Ser., Vol. XVII, 1017.]
Jly Exehaugt.
DOOMIH (P.). H^tcrocArfs de I'ArotTiqUB du Sud.
[Faac. XUI. 1817,] Tkt Author.
n,g,t,7.dh,G00glc
DONHTHOBPK <H.)l {Delie/ioderiii) (Hvfwelinea) micfryt, d, sp., » species
a[ Ant new to Scirnt-e : niib s lew noUs on the geuua.
[Botom. Becord, Vul XXIX, 191T.] Thr Juthor.
BDwum (F. W.). HiMqnitues and their retation to diiesBe, tbeir life-
biston. babita and control.
[British Huseum (Nstural History), Economic Series. No. 4, 1914.]
Till TnaUti £Hl. Mvl.
Faqan(H. H.). [SeeRuHwiB (8. A.).].
FouoM (J. W.). North AmeriFaa CoUemboloua iuaecta ot the subtamily
OnycbiuriDse.
[Proc U. S. Nat. Mu« , Vol. LIII, Aug. IBIT.]
The Umithioitias Iitilit^tioH.
FitBD (G. H.J. Obaeriatioiis on the larval anil pupal stages ot Jifriolii
oiinrvi. LiDDKua.
[Add. Applied Biolog}, Vol. Ill, Jan. 1911.1
J>epl. Ayric, t'lllom., MaKchetlrr.
Fox(H.). Field Dote* on VirgioisD Orthoptera.
[Proc. U. 8. Nat. Mus., VoL LII, Haruh 1917.)
Ibt SMitkioAiaa iHUilulion.
FbuOOATT (J. L.). [SteFHOOQiTTOV.W,).].
FBOOflAiT (W. W. and J. L,). Pb^f p-mnggot Flies, No. 3. Beport of work
carried out iu the Itivirius Uistnct, during 1915-16, at the
Oovernment SheefBy Eiperimeiit Station at Woolooodool,
GiHAN (A, B,). Deierlptions of some new parasitic Hymenoptera.
[Proc. U. 6. Nat. Hiu., Vol. I.Ut, Hay I91T.]
Thi Saittluoaia'i Imtitiitha.
Oeteb(E. W.). [8eeQDAiNT*NCK(A. I,.).].
Gibson (A.). A new species of Torlrii of economic importance from New-
foundland (Lepidoptera : Tortricidae) .
[Canad. Hntomologiat, Vol. XLVIII, IS16.]
Tlu Jtithar.
The Bntomological Record for 1918.
[RepriDtcd irom 4Tth Anu. Rept. Ent. iSoc, Ontario, 1916.]
tanarf. Dtpt. Aj/ric.
nlilla, Zell.).
*, I>rpt. Agric.
K <E. B.). Notea on Coccidse accurriog in the Seychelles IsUnds, with
deacriptions of new jipccies.
[Bull. Ent. Keseatch, Vol. VII, Oct. 1918.]
— Obserrations on Brilish Coccidse, with i
No. m.
[Eot. Monthly Mag., Vol. LllI, lOlT.]
n,g,t,7.dh,G00glc
( zxxiv )
Obekn (El B.)- ObwrTktkiM od Britiab Coccidw, with dc*
tpraet. Ho. IV.
[Bot. H<nithl7 H<g., Vol. LIQ, 10171
iiBiw<l'. HO- AGuidetatheLilnmtureof BrituhDipton.
[Proe. Rojril Boc Edinburgh, Snsiau I9I<I-17, Vol. XX, Put 3.
The .IXAor.
IK to (be Kpecimeof and mUned inadclK of inxectaaiid ticks exhjbitn) in
tbc Ceotral HkII, Bntiab Humudi (Natural Hiitorj), illnitrat-
<Dg tbeir importaiKe in Ibe wfrad of diwaae.
[Sped*] Guide No. T. 1916.] T%t Trutlrri Brit. Mts.
Vol. XUX, iei6,l Tki Author,
Hahfson (Kir George). The DrtenninatioD of Generic Type* in tbe
Lepidopten.
[Bolom. N«w«, Vol. XXVJIJ, Dec. IBIT.] Tkt Autkor.
Hawirr (C. G,). Reguiatiooa under tbe Dettnictive Insect and Pett Act,
with iantructious to importen and eiportin of trem, pbntc
and otber nuraery atock.
[Canid. Dept. Agric, Entom. Bnncb, Circular No. 10, 1917.]
Report of the DominioD EDtomoloaiat for tbe year ending March
31,1917.
RegulatiouB under the destructive Inaett and Peat Act, 1917.
[Canad. Dept. Agric, Eutom. Branch, Circular No. lil.lBlT.]
Vamii. itepl. Ayric.
. — [See Report of (be Dominion Entomologiat, OtUwa, 1917.]
HiBsT {%.). Species of Ararhntila and Hyriopoda (scorpiona. epiders, mitea,
tick* and centipede*) injurious to man.
[Britiab Husetim (Natural History), Rconomjc Series, No. 6, 1917.1
The TTttiteri Brit. -Vbi.
UoDoiiss (H. B.). [See Pakrott (P. J.).]
Ihhs (A. D.). Tarred felt "disci" for protecting cabhagea and rela(«d
vegetables from attacks of the root fly {Ch-rleMte /irauicat).
[Joum. Board Agric, Vol. XXIII. March 1917.]
Ilfpl. Auric. KatBm.. Manehtilir.
[See C<>LK (A. C.),]
Islti (Dwight). Control of tbe grape-berry moth in the Erie-Chalauqua
Grape Beit (/•o/ycAra. - - '■' - •
[U. 8. Dept. Agric.,"Kid1. No, 5.W, Aug, 1917.]
U. S, litft. At/rie.
R1K> iJ. Joauuia) Invesligsciones sobre el pHludjomo en Pnebla.
[Piib'tshed by the (^olegiu del K-Uilo di- Puebla [Mexico], 1916.1
Thf .Si«U Calftgt of FutMa.
<an (H. 8.). I'ure lines in the study of genetics in lower orgaoiima.
[Amer. Kal., Vol. XI.V, Feb. 101 1.]
— The Numerical reanlta of iliireriu sy(ti;nis of breeding.
[Geuetic*. Vol. I.Jan. IHlti]
— Hereflity, Variation atid tbe Results uf Selection in the UBi-
parental reproduction of DiMnyiii conma.
[Genetic*, Vol. 1, Sept. 1916.]
n,g,t,7.dh,G00glc
XXXV )
jDniiN09(H. S.). The numerical mulUofJicerwijatomiiof breeding, sith
respect to two pair* of cbsnctmt, linkeil or indepeDdent, with
■pecial relation to tlie effect! of linkace.
[Oenetics, Vol. 11, Hareh ]&1T.]
Hodirjing ftetorii and multiple allelomorph! in relation to the
reMilts or nelectioD. [Baaed on the fruit At I>naimhila.'\
[Amer. Nat., Vol. LI, Ha; ISIT.j
Obeervi d change* in Hereditary characters in Relation lo Brolatimi
[Jouro. Washington Acad. Sci., Vol. VII, Hay ISITJ
The Author.
Jo)iM(D, W.I. TbeEnroHan Barwig<fDn!ci>/n iiiiri<-ii(Dri*<i)and itacontrol.
[U.S. ft'iH.Agrlc., Bull. No. 566, June 1917 1
IJ. S. Dtft. Agric.
Jones {T. H.). The 8ireet-pot»to leaf-folder (PiVktki* trimnelala. Fab.),
[f. S. E>ept. Agtic, Bull. No. «», Nov. 1917.1
('. S. Dcpt. AgrU.
KiBiPtTnicK <R ). The Biology of Watcroorks.
[British Museum (Natural Uirtor;), Bconomic Striea, No. 7. 1917.
Thi TTi-stit> Brit. M«,.
KRjiiTiiO.) [8«eKl'sTen(H. C,)]
Edsteb (H. C.) Die Kafrr Eiiropa's. Kach der Natur bescbrieber. Heft
Li RcB (Q. R.) Two new larial Trrmalodts from Thamnophit utareiajta
and Tkemnopha tauei.
[Occa*. Pnpers Hua, Zoo!. UdIy. Michigan, No. 35, Jan. 1917.]
Unix, of Afiekigaii.
LiTBM>p (F. H.). [Pee Pajiiiott (1'. J, ).]
Lecas (T. p.). Memoir Ipubltnbcd in Brisbane 1917).
tVr«r- »--.(-..-. fo., flriiion*.
Lccl^■lLL (P.) an<l Aikblin (O. tl.). The Lesser I'omaUlk -borer (E/aimn-
palpfs ligioidlvi. Zell.).
[U. 8. Dept. Agric. Boll. No. 539, Sept. 1917.]
v. S. Ihpt. Agrif.
LoNPBICK (W.>. Diptera DaaiciL Parts IV, V, 1912, 1916.
Th, Author.
HarBB (U. O.). Life-history of /'lalella macuUptunii, the diamond-back
[Joum. Aigric. Research, Vol. X, July 1917.
('. S. Dipt. Agric.
UcCrat (A. H.). Spore-formiDg Bacteria of the A|dary.
[Joum. Agifc. Heseircb, Vol. VIII. Marrh 1017.]
C..S. Iltyl. Agric.
HcDoKovaH (F. L.). [Res HcfinKwu (B. A.).]
HcGiHBB(T.F.). [SeeCoAD (B. K.).]
HcObigok (B. A.>, Descriplions of seven new species of Red Spidera.
[Proc. U. 8. Nat, Hus„ Vol. U. Jan. 1917.1
HoJrwoh (E. a.) and HcDonoi^oh (F. L). .The Red Spide
(Tttranychui bimactiatvs, Harvey).
[U. S. Dept. Agric. Bull. No. 4ia, Jan. 1917.]
iM,Googlc
MclMDOO (N. B.I linJ SlKTBBS (A. F.). (jlttMW f.
inKCtiride.
[Jouro. Agric. Kesearcli, Vol, X. Sept. 1917.J
S. Iftpt. Ayri,
HlSRAfC.S.). The ladixu Sugir-cim
[M™. Dept. Agrie. tudia, ._ . .
1917.] JaJiaOffift.
UdbsisoM (H.). Monograph of the Nearctic Hjinenopt^n of the genua
Bmen«, Fabric jus.
[Proc. U. 8. Nat. Mu^.., Vol, Mi, Marcl. I1H7.]
Tht .S«,i,h,o«ian Inititulio,.
Mi-KBO (W.). [See CUOiKNT (G. K.)-]
Mtb»(P. K.), AuAmerieuiJipecieiior the HymenoptiTouB genua H'fimatlta
of Koerster.
[Proc. U. S. Nat. Hint., Vol. LIII, May lfll7.]
A new Ameriran pararile of the Hfuiaii fly | Majiniolii deitmclor,
Say).
[Proc. U. S. Nat. Mus.. Vol. LllI, U*j IPIT.]
Tht .SmithiBiinn /sjliJiKiojt.
N0THAN(H.). Coleopten IlluMraU. Vol. I. No. 3. CaraMdae. 191T.
The Avthor.
OuERTHtiR (riiarlec). fitndeR de I.-pulo|)l4Tologie Company. XIII. XIV,
1U17. Ti/ Autkor.
P«INK (J, H.l. Ad aafnimetriral Biid-louae roiirul on three iliffereut apade*
.MH/iunurton.
R (J. B.). A Kevisioa of tlie Bembicin? wupa of America north of
[Proc. U. S. Nat. Hu«, No. LIl, Feb. 1817.1
Thf SmiihtBuiau liuUulim.
R <R. R.). New flies of the genua Kircni^aga from Quam aud the
Philippiuen.
[Proc. U. S. Nat. Miu., Vol, I,1V. OtI. 1917.]
Tht Satithioaiatt JnsliUition.
1S.S (H, E,|, an.l Latmhop <K,
._, pple orcharrit, II.
[New York Agrir.
PATtH (E. M.). Elm-lenf rosetle awl
[Maine Agric. Biprr. Station
Fiu*B« (L. M.). [See Sanderiion (E. 0,1.]
m(U, B.). [KeeBAcafE. A.).l
',.>and BakemCA.C). A Gintribution to onr KnonledKe
of the Whit« Bies of the *tibfarotly Aleyrodinae (Al«yradidae).
[Proc. V. S. Nat. Mus., Vol. LI, Jan. 1917.]
Thf SmilhtenlnH ftutilHlion.
(JDAiNTAKCE (A. t..l and OifEn IE. W.). Life-hiitory of the Codling moth
(Cari-rxvpia iHimoKr//". L.) in the Peros Valley. New Meiico.
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('. .V. Dipt. Afrie.
ih,Cooi^lc
( XXX vii )
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n,g,t,7.dh,G00glc
TRANSACTIONS
ENTOMOLOGICAL SOCIETY
LONDON
Fob the Year 1917.
I. Descriptions of Soulk American Micro-Lepidoptera.
By E. Meymck, B.A., F.R.S.
[Read December flth, 1916.]
The following descriptioos are from specimens in my -own
collection. They include some further interesting material
from high levels in the Andes (9000-12,000 feet elevation),
amongst which are' examples of the genera Pkalonia,
C'roddosema, Arislotelia, and Phtkorimaea. Altogether six
genera and 102 species are described as new.
PHALONIADAE.
Amallectis, n. g.
Palpi rather long, porrected, triangulsrly rough-scaled. An-
tenaae in S strongly ciliated. Forewinga with 2 from towards
angle, 3 absent, 8 absent, II from middle. Hindwings with 2 fiom
towards angle, 3 and 4 stalked, 6 somewhat approximated, 6 and 7
long-stalked.
Probably a development of Phalonta.
Amallectis devlncta, n. sp.
$■ 13 mm. Head ochreous-whitish, sides more ochreous. Palpi
oehieons-whitish, laterally more ochreoos, with a few dark fuscoua
speoka. Thorai. oohreons -whitish, shouldera marked with dark
TEANS. ENT. SOC. LOKD. 1917. — PART I. (NOV.) B
iM-,i,A J h, Google
■_$ •. '"■ Mr. E. Meyrick's Desenpttmu of
fuMXHU. Abdamen pale greyish, anal toft whitiah. Forewinp
elongate, posteriorlj dilated, coata gently anihed, apex obtnae,
t«rmeii obliquely rounded ; ochreous-whitiah ; a dark fnscoua ipot
mixed with brownish -ochreonB on baae of coeta, and another near
bejond it ; a transvene fascia from middle of costa, costal portion
lather narrow, dark fuscous, mixed with ochreous-brownish, lower
half formed of grej irroration, dilating to dorsum, where it extends
over more than median third, and is marked with dark fuscous; a
clondy grey spot on coeta towards } : cilia ochreous-whitisb.
Hindwin^ whitish, with some cloudy grey stiigulae in diao and
posteriorly; cilia whitish.
Peru, Lima, in August {Parish) ; one Bpecimen.
nuJonU subUmls, n. sp.
$. 16 mm. Head whitish. Palpi 2}, pale oobieoos mixed with
ferruginous. Thorax whitisb-ocbreouB tinged with ferruginous.
Abdomen greyish. Forewinga elongate, rather nairow, posterioily
■lightly dilated, coeta slightly arched, apex obtuse, tfimen hardly
rounded, oblique; whitish-ochreous, almost wholly suffused with
femiginous except towards costa beyond middle; some minute
blackisb strigulae scattered along oosta; a tnodetate suffused deep
ferruginous fascia slightly before middle pardlel to termen, this
fascia and dorsal half of wing beyond it minutely speckled with
black; several deep femiginous spots and mariu on apical fourth
of wing forming a coarse reticulation : cilia whitish-ochreous, su(-
fnsedly barred with grey. Hindwings whitish-grey, IndisliiMtly
strigulated with grey ; cilia grey-whitish.
Peru, Huancayo, 10,650 feet, in July {ParUh); one
specimen.
Phalonla galbanea, n. sp.
S 9' 11-13 mm. Head and thorax pale brownish -ochreous, face
whitish-ochreous. Palpi 2], oobreous-whitish tinged with brownish.
Antennal cUiations of S 1- Abdomen rather dark grey. Forewings
elongate, posteriorly slightly dilated, costa slightly arohed, apex
obtuse, termen rounded, rather oblique; light brownish -ochreous;
basal patch slightly darker, undefined, on costal edge often snfiFosed
with dark fuscous; a rather narrow ochreous- brown postmedian
fascia, obtusely sngulaled and interrupted above middle, its costal
edge sometimes marked with dark fuscous; a more or less indicated
ochreous- brown mark, sometimes with dark fusoous scales, on dorsum
before torous; a rather narrow ochreous-brown fascia from j of
Ij.y.l.Ajl^yCOO'^lL'
South American Mvcro-Le'piiopteTa. 3
coat* to middle of tormeo, and three or four imall indistioot A»A
fnflcooa costal atrigol&e oa or near tbie : cilia pale ochreoua, tsiuMj
spotted with dark fuscous or blBckiah irroration. Hindwings
blackish-grey ; cUia pole grey, vith darker grey subbasal shade.
British Guiana, Bartica and Mallali, frum December
to March {Parish); Peru, Pacaya, in August (Mounsey);
eight specimens.
Phalonla swrifluti, n. sp.
S $. 8-1 1 nuD. Head, palpi, and thorax pale ochreons. Antennal
oiliationB of (J 1. AbdomeD grey. Forewii^ elongate, poet«riorij
BODMwhat dilated, oosta gently arched, apex obtuse, tennen obliquely
nnmded; 7 to oosta; oohreoue, more or less silvery-irideeoent ;
some minate scattered strigulae of black irroration along coeta; a
qiiadnrte deeper spot on costa beyond middle, edged laterally with
■trignlae of black irroration ; some indistinct darker suffusion, with
inegoUr mariu of black irroration, in disc beneath this ; a more or
leas developed biangnlar blotch of black irroration on dorsum before
middle, and a small spot towards tonus; an indistinct spot of
deeper snSnsion above tomus ; an indistinct deeper faaciat« streak
from ^ of costa to middle of termen, its lower portion covered by an
irregular patch of scattered black irroration ; cilia pale oohreouB.
Hindwings with 3 and 4 closely approximated or connate; grey,
thinly scaled in cell, veins darker ; ciiia pale grey.
British Guiana, Bartica and Mallali, from December to
March {Pariah); Dutch Guiana, Paramaribo; Brazil,
Parana; twenty-six specimeoB.
Phalonla enellUea, n. sp.
S- 13-16 mm. Head whitish. Palpi 2), oohreouB- whitish,
tinged or sprinkled with brownish. Antennal ciliations nearly 2.
Thorax whitish, sometimes tinged with ochreous. Abdomen grey, anal
tuft whitish. Forewings elongate, posteriorly slightly dilated, costa
gently aiohed, apex obtuse, t«rmen nearly straight, oblique; pale
brownish-OChreoQB, sometimes more or less whitish-tinged; a faint
brownish basal patch, edge angulated, its lower portion marked by a
more or less indicated darker streak; a moderate brownish post-
median fascia angulated and more or less interrupted in disc, par-
tially sprinkled and marked with dark fuscous on edges, lower portion
forming a semioval spot ; a roundish brown spot on costa at ;), and
one or two dark brown strigulae before and beyond it ; cilia pale
iM-,i,A J h, Google
4 Mr. £. Ueyrick's Descripliont <^
oohreooB. Hindwinga grey; oUift whitiBh-grey, irith grey mibbMal
shade.
Ecuador, Alsusi, 9450 feet, in June (Pamh); five
specimens.
Phklonla mordax, n. sp.
[}. 1&-I7 mm. Head, pali», and thorax whitish, more or leas
sprinkled with fuscoDs and dark fuMoua, palpi 3. Ant«iiiial oilia-
tloDB 2. Abdomen whitish -fosoons. Forewings elongate, posteriorly
■lightly dilated, coata gently arched, apes obtuse, termen ronnded,
obliqne ; whitish, strewn with pale ochreous strigolae more or less
iiTonted with fusoous and dark fuscous; maikiogs brownish
sprinkled with dark fuscous; thiee somewhat oblique spots from
costa at i, middle, and }; three rather iiregnlar oblique fasoiate
streaks from donrum, first at }-, reaching halt aoross wing, second
from beyond middle, similar but more or less interrupted beneath its
apex, third from lomus, more or less obsolete at tower extremity,
reaching f across wing near termen; some small msrginal spots round
apex and termen : cilia whitish, mixed of indistinctly barred with
fuscous. Hindwings pale grey, strigulated with grey ; cilia whitish-
grey, with darker subbasal shade.
Aegentina, R. Parana; four specimens.
Lasiothybis, n. g.
Antennae in ^ ciliated. Forewings with 2 from J, 3 and 4
approximated, 7 to costs, 11 from middle. Hindwings with 3
and 4 connate, 6 hardly approximated, 6 and 7 nearly parallel.
Apparently allied to Pharmacia.
Lasiothyris linutula, n. sp.
S- 9 mm. Head and thorax light ochreous, face whittsh-ochreous.
(Palpi broken.) Anteimal ciliations 1. Abdomen grey, anal tuft
grey-whitisL Forewings elongate, costa slightly arched, apex
obtuse, termen rounded, oblique; light ochreous; costa marked
with some minute indistinct dark fuscous strigulae, and with narrow
elongate spots of ferruginous -fuscous suffusion at \ and middle :
dorsal half suffuspd with ferruginous, more strongly posteriorly,
with a few dark fuscous specks, crossed by some silvery-whitish
iridescence beyond middle ; an oblique ferruginous -ochreous fascia
from } of costa to middle of termen, preceded and followed by
silvery -whitish iridescence : cilia ochreous, towards tips whitish-
tinged. Hindwbigs hyaline with thinly strewn dark groy hoirscales.
Ij.y.l.Ajl^yCOO'^lL'
South American Micro-Lepiioptera. 6
tomrda tennen raSused vitb giey, veins uid t«rmin&I edge d&rk
grey; oiliA whitish-grey.
EcuAiK>R, Huigra, 1500 feet, in June [Pariah); one
specimen.
Ptitbuehnis praepositi, n. sp.
^. 26-30 mm. Head and palpi fuscous. Anteonal ciliationa
1|. Thorax white. Abdomen light grey. Forewingi elongate,
posteriorly considerably dilated, costa anterioriy hardly, poBt«rioriy
moderately arched, apex obtuse, termen slightly rounded, rather
oblique; silvery- white ; a rather broad somewhat oblique fusoous
median fascia, tending to be partially interrupted above middle,
extended along coeta as a suffnsed diminishing band t« base, on
oosta marked with small darker spots or strignlae, in disc with some
silvery iridescence and transverse markings of rained silvery acalee ;
about seven dark fuscous dota on dorsal edge; a. rather curved
traoBverae fuscous streak mixed with dark fuscous above tomua,
and a small spot above upper extremity of this ; a triangular fuscous
apical pateh, partially sprinkled with dark fuscous, extending over
neariy \ of coeta and to below middle of termen : cilia grey (imper-
fect). Hindwings pale grey, coarsely strigulated or marbled with
grey; cilia whitish -grey.
Colombia, San Antonio, 5800 feet, in November; two
specimens.
TORTRICIDAE.
CApnk amett, n. sp.
i^. 17 mm. Head and thorax brown, suSusedly irrorated with
dark grey. (Palpi broken.) Abdomen grey. Porewings sub-
oblong, costa anteriorly moderately, posteriorly slightly arched, with
costal fold reaching nearly to J, apex obtuse, termen straight,
somewhat oblique; ferruginous -brown, irrorated with dark grey
costal fold and about Hvc costal spots suffused with blackish-grey
central fascia and large triangular costal blotch represented by a
broad indelinite general blackish-grey suffusion ; a tuft of raised
scaJee on fold beneath middle of wing, and numerous scattered
raised scales on surface of wing : cilia brownish, basal half spotted
or mixed with dark grey. Hindwings grey, towards apex and upper
part of termen paler and strigulated with darker ; cilia grey -whitish,
with snbbasal line.
Peru, Lima, in August {Parish); two specimens.
iM:,i,A J 1^, Google
6 Mr. E. Mej'iick's Descriptunu <^
Ctpna lllsqusts, n. sp.
$. 14 mm. Head and thor&x whitish-jellow, ant«rioi mugin
of th<n«j( marked with brown. Palpi yellow-whitiBh, with two
tranarerae fuscous ban on second joint, and one on base of tennuutl.
Abdomen giey. Forewings elongate, posteriori^ dilated, oosla
gently arched, apex obtuse, termen rounded, rather oblique; light
brassy-yellow sprinkled with ferruginous, partially euffosed with
eilveiy-white on margins of markings; markings lilac-brown, edged
with some dark brown scales; a fermginous- brown spot along base
of costs, enclosing two pale yellow atrigulac on costs ; a small
ferruginous spot on base of dorsomi an irregular oblique fascia
from \ of costa to below middle of wing, its extremity dibted and
truncate ; a triangular blotch on dorsum about i ; a large spot on
middle of costa and a smaller one beyond it, connected by a thick bar
beneath ; a small costal spot beyond this ; an irregular fascia from
costa before apex to tomua, including a pale yellow strigula on oosta,
its anterior edge triangularly prominent above middle so as joat to
touch angle of the subcoBtal bar and suSused here with fermginous,
its posterior edge connected with termen below middle by a narrow
bar ; some small silvery- white spots along termen and tomus : cilia
pale yellow barred with ferruginous. Hindwings dark grey; cilia
greyish with darker subbaaal shade.
French Guiana, R. Maroni; one specimen. Allied to
alumiiiias.
Pseudatteria forniMU, n. ap.
9. 30-38 mm. Head white spotted with black. Palpi nearly 6,
slender, black, bssal joint white, second joint internally white.
Thorax black, posterior margin of collar, a mark on each side of
back, a dot in middle, and two posterior white. Abdomen black,
segmental margins white. Forewings suboblong, moderately
broad, costa strongly arched, apex rounded -obtuse, termen rounded,
somewhat oblique; orange ; dorsal edge ochreoue- whitish ; a
moderate evenly broad ochreous -white border running all round
costa and termen, with purple-black markings as follows, viz. a
basal bar, eight thick transverse spots crossing it between this and
I of costa, four marginal spots round apex connected anteriorly
by an irregular marking but first or second of these sometimes dis-
cormected, and five semicircular marginal spots on termen, second
conQuent with an anterior spot bo as t« form a bar crossing border;
in one specimen a dot beneath this bar anterioriy, in the other a
small spot above tomal spot : cilia white, barred with purple-black
on markings. Hindwings coppery -orange, dorsum nsnowly suf-
l„y,|,AJh,COOl^lC
South Ammean Micro-Lepidoptem. 7
fnaed iritb darii grey ; oine snuU nmioiroiUAr puiple-black moigiiiaJ
■potB round «pex utd tennen; oilia whitiah, beooming oiwige
tomrds tomtu and duk grey on domun, barred with ptir[Je-
Uackioh on »pat«.
Colombia, San Antonio, 6800 feet, in November; two
CMoeda eheli^rapto, n. sp.
(f. 13 mm. Head whitiah, aides of ototd mixed vith orinuon-
foBcoOB. Palpi vhitiab, eli^tiy aprinkled with crimaon. Ant«uu1
ciUatkniB }- Thor&x pale ochreoua, anterior margin aaffused
with pnrjdiah-bniwTi. Abdomen ochreoua-orange. ForewingB tmb-
obtong, moderately broad, rather dilated posteriorly, coata anteriorly
gently, posteriorly hardly arched, without fold, apex obtiue, t«rmen
•lightly rounded, neariy vertical ; pale glosay yellow-ochreous; an
obliqne dark rcddiah-brown streak from middle of costa reaching
half across wing, snSnsed with dork grey except on costa; a small
dark brown spot on oosta at | : cilia whitish-ochieoua, on upper
haU of termen aufinsed with reddish -blown. Hindwinga oohreotu-
orange ; eilia light ochreoua.
Fbbnch Guiana, R. Maroni ; one specimen.
Caeoeoift MrobktlM, n. sp.
i. 30 mm. Head blackiih-fuaooue on crown, face and palfn
light greyiah-ochreona. Ant«nnal ciliatioiiB I. Thorax light glossy
lilao-foBOOUa. anterior margin blackish-fuBcouB. Abdomen giey.
Foiewings elongate, moderate, posterioriy dilated, ooata gently
•rafaec^ without fold, apex obtuse, termen hardly obliqne, obtusely
bent in middle; rather light purple-fuscous, finely strioUted tiana-
Tersely with dark grey, along costa suffused with grey; an irregular
txansrerse white spot in dise at { ; a large balloon-shaped blaokiah
Uotoh before tennen, extending from near costa | acrosa wing,
partially edged finely with white, especially on upper part anteriorly :
oiUa purple-brownish (imperfect). Hindwings grey, with some faint
da^er striguloe posteriorly ; cilia grey-
CoLOMBiA, San Antonio, 5800 feet, in November; one
specimen.
Tortrli InfawnUiM, Zell.
I think I have correctly identified this as a apecies of
which I have eleven examples taken at Huigra, Ecuador.
4500 feet, in June. If so, it ia a true Tortrix, very like
iM-,i,A J h, Google
8 Hr. E. Heyiick's DtacripUont oj
the North American peritana, bat Zeller ia not strictly
correct in stating that the male has no costal fold; the
costal edge is slightly rolled over on basal fonrth, and the
posterior part of this is furnished with rather long project-
ing scales, but so closely appressed to surface of wing as to
be unnoticeable unless liftt^.
Tortrix homophyla, n. sp.
^. 12 mm. Head, palpi, and iiioras ochreoilB. Antennal oilia-
tiouB 1. Abdomen pale whitiBh-ochreooB. Forewinga elongate,
posteriorly somewhat dilated, costs gently aicbed, without fold,
tffiex obtuse-pointed, t«nnen elightly Biniiat«, rather oblique;
whitish-grey-ochreouB, stiigulatod with light giey, with several
indistinct grey-yellowish striae ; edge of basal patch angulated in
middle, marked with blackish in disc but obsolete towards margins ;
central fascia moderate, oblique, grey, anterior margin straight,
marked with blockish scales, posterior irregular and indefinite;
costal patch large, semioval, dork grey, sending a well-marked dark
grey line to termen below middle, continued a little down it; a
dark grey streak along apical part of termen : cilia whitish-grey-
ochreous. Hindwings ochreous-groy-whitieh, towards apex with a
few grey strigulae ; cilia ochreous-grey- whitish.
Colombia, La Cnimbre, 6600 feet, in May (Parith);
one specimen. Very Bimilsr to preceding, yet easily dis-
tinguished by. sinuate termen of forewings and whitish
hindwings. besides other differences.
Tortrlx eapnostlehs, n. sp.
1^$. 13-16 mm. Head and thomx pale fuscous. Palpi oofareous-
whitish sprinkled with dark fuscous. Antennal ciliations of 3 ''
Abdomen fuscous. Forewings elongate-oblong, costa anteriorly
moderately, posteriorly hardly arched, in (J with edge very narrowly
rolled over from base to i and slightly rough-scaled, apex obtuse,
termen straight, rather oblique; pale ocbreous irrorated with
fuscous; central fascia moderate, oblique, rather dark fuacous,
somewhat narrowed upwards, margins nearly straight, sometimes
irregulariy interrupted above middle ; costal patch rather large,
irregular- trapezoidal, rather dark tuBcous, with an indistinct lino
running from it to termen below middle : cilia pale ocbreous, with
line of fuscous irroration. Hindwings whitish, obscurely strigulated
with groy towards apei and on termen, vein 2 and lower margin of
cell beyond it suffused with grey ; cilia whitish.
Peru, Lima, in August (Parish) ; six specimens.
ih, Google
South American Miero-LepidopUra. 9
Torlrlz Ugnea, n. Bp.
(J$. 16-18 mm. Head, palp, and thorax brown. AntemuJ
aUatioiu of <^ |. Abdomen whitish-gKy. Forewingi sabobknig,
coata vitbont fold, anteriorly strongly arched, posterioriy slightly
sinnato, apex obtnae, termen samewhat bowed, little obtiqae ; brown,
■omewbat tinged with fenuginona, abecorely fltaigalat«d with dark
fuscous, especially on margins; basal pateh obeoorely darker,
p&rtially aufinaed with dark fuscous irroration, especisJIy towards
lower half pOBteriorly, edge angulated in middle; oential fascia
daiker, broad, rather irregular, from before middle of oosta to
dorsum before toruD-, narrower and irregularly mixed with dark
fnsooua irroratioD .owarda oosta; costal pateh darker, flattened-
triangalar, whence a thick streak runs to termen below middle;
a short darker streak along upper port of termen ; cilia whitiah-
oehreons, with dark brown Bubbaaal shade- Hindwings grey-
whitish, strigulated with grey posteriorly; cilia whitish with grey
sabbasal line.
Ecuador. Huigra (4500 feet), Alauai {9450 feet), in June
(Parish) ; six specimens.
Tortrix Ossionlata, n. Bp.
S9- lS-20 mm. Head, palpi, and thorax light brownish closely
irrorated with dark fuscouB. Antcniml ciliationa of S 2- Abdomen
grey. Forewings elongate, pOHteriorly dilated, costa moderately
arched, withont fold, apex obtuse-pointed, termen somewhat
sinuate, oblique ; brown suffusedly irrorated with grey, with inegolar
anastomosing dark fuscous transverse striae ; the confluenoe of these
forma a narrow somewhat curved fascia about |, a narrow irregular
somewhat oblique fascia beyond middle, and a streak from ] of
costa to tomoH, widest on coita : cilia whitish-fuscous, with dark
fuscous subbaaal line. Hindwinga grey-whitish, more or less
strigulated with grey ; cilia whitish, with grey subboaal line.
Peru, Aqualani, in June; four specimens.
Ealla atalodes, ii. sp.
S- 14 mm. Head and thorax silvery-white, face, palpi, and
sbouldera dark fuscous. Antenna! ciliations 1. Abdomen whitish.
Forewings rather elongate, posteriorly dilated, costa slightly arched,
straight in median ai«a, apex obtuse, termen slightly rounded,
obUque; silvery -white, tinged with pale greenish -yellow; a dark
iM-,i,A J h, Google
10 Hr. E. Bfeyrick'B DaenpUom r^
fuBcooB streak along basal fifth of costa, edged beneath b; an un-
defined spot of pale yellow anffusion ; a laige dark fiuoous triangular
bloteb extending on oosta from i to ^, and reaching half aorois
wing; a emaU dark foBoous spot on oosta near apex; some light
giey safhuion along median portion of teimen : oilia light gny,
towards apex and tomus white. Hindwinfp pale grej, towards
base whitish-tinged ; cilia whitish, with two pale grey linee-
CoLOMBiA, .San Antonio, 5800 feet, in November; one
specimen.
Eulia melaneets, n. sp.
$. 19 mm. Head and palpi whitish tinged with fuscous. Thorax
iriiitisb, anterior margin inftuoated. Abdomen pale grey. Fi»e-
wingi elongate, posteriorly slightly dilated, oosta gently arched,
apex obtuse, termen rounded, oblique ; whitish ; costa mariced with
a aeries of small dork grey spots, and in middle with a moderately
large irregular spot; a dark grey patch occupying doraal half of
wing from base to f, towards base extended to 04>sta, posteriorly with
an irregular grey lobe above nearly reaching median costal spot
marked with a subtriangulsj' black spot anteriorly and a black dot
beyond this; a curved sabterminal grey streak, marked with some
small black doU in disc ; a grey niarginal streak round apex and
termen, marked with four small irregular black spots on Ha upper
portion : cilia dark grey irrorated with whitish. Hindwings grey-
whitish posteriorly marbled with grey; cilia whitish, with grey
sabbasal line.
Ecuador, Alaiui, 9450 feet, in June (Pariah); one
specimen.
Cnephisla utou, n. ep.
^. 18 mm. Head and thorax whitish, sides of crown and shoulders
mixed with dark fuscous. Ant«nnal ciliations }. Abdomen grey-
whitish. Forewings elongate, costa gently arched, apex obtuse,
termen straight, rather obUque; ochroous-whitisb, with a few
scattered dark fuscous scales posteriorly, and some dots along coeta;
a Bubtriangular dark fuscous spot on base of costa, and a smaller one
at } ; a quadrate dark fuscous spot on middle of coBtA, an oblique
fascia from dorsum before middle pointed above and reaching more
than half across wing, and a curved elongate spot in disc beyond
middle, these three all connected by a spot of grey suffusion; an
irregular-trapezoidal dark fuscous spot on dorsum at J, and an
insular bent fascia from oosta at ^ directed towards this but only
Ij.y.l.Ajl^yCOO'^lc
South American Micro-L^doptera. II
rnnrhmg half Miross wing : cilia wfaitisb (Impeifeot). HimtwingB
oohieoiu-whitiBh; a brush of very long spreading black haino«JsB
wiUiin odl; cilia vbitish.
Colombia, San Antonio, 5800 feet, in November; one
specimen.
Ow^uuia pneelpna, n. ap.
3. 18 mm. Head and thorax whitish. Palpi whitish, basal
half sprinkled with dark fuscous. Anternial ciliatious J. Abdomen
grey-whitish. Forewings elongate, posteriorly rather dilated, costa
gently arched, apex obtuae, t«nnen nearly straight, rather obliqne;
ochreoaa-wbitlsh, faintly tinged hei« and there with pale reddish-
ochreooB ; markings dark f usooux ; several dote or strigulae on costa,
and a small triangular spot at J ; a quadrate spot on middle of costa,
an oblique fascia from dorsum before middle reaching haJf across
wing, and a mark in disc beyond the interval between them; a
large acute-triangular spot on doraum beyond middle ; a fascia from
} of costa to it of dorsum, anterior edge with an irregular prcuninence
in middle, posterior edge nearly straight : oilia ochreous- whitish
(imperfect). Hindwings ochreous-whitisb, irregularly marbled with
grey; cilia ochreous-whitish.
Colombia, San Antonio, 5800 feet, in November; one
specimen.
Cntpbasia dentati, n, sp.
^ 9- 13-14 mm. Head and thorax whitish, shoulden with some
dark fusoons scales. Palpi dark fuscous, terminal joint and apex
of second whitish. Antennal ciliations of g I. Abdomen whitish-
grey. Forewings elongate, costa moderately arched, apeK obtuse,
termen slightly rounded, lather oblique; oohreous- whitish, tinged
here and there with light brownish ; some blackish dots or small
atiigulae on costa ; a dark fuscous spot on base of costa, and a smaller
one at } ; an oehreous-grey spot on middle of costa; an ochreous-
grey inwardly oblique fascia from costa at }, reaching { across wing,
contracted above middle, becoming pale ochreous in disc; a dark
foeoons oblique fasciate blotch from dorsum before middle reaching
half across wing, its upper extremity denticulate and connected
by an irregular streak with a dark fuscous spot on dorsum at
t ; some blackish strigulae on upper part of termen : cilia whitish,
on upper part of termen suffused with dark grey, on costa barred
with dark fnscous. Hindwings grey-whitish, irregulariy Btrigulat«d
u,y,l,/eJh,G00glc
12 Mr. E. Sbyrick'B Daaiftiaiu tf
withgR?; eaMwUtMh. Fcaewn^ bcoMtb anfliMMl with reddiah-
EcuADOB, Hmgia, 4500 feet, in June (Parish); two
Bpecimene.
Cne^iuis laaam, u. sp.
1^. 12 mm. HcMi and tbonx wliiUsb-ocbreoua tinged with
browniih. Palpi rotber short, whitish, irrorated with blockish.
Antennsl cilistions 1(. Abdomen ochreons-gcej- whitish. Fore-
wings elongate, costs slightly siched, apex obtDse, tecmen elightlj
rounded, oblique ; whitisb-ochreona ; a broad datk grey tiansverse
liand, mixed with reddish' blown and blackish, anterior edge nui-
ning from ^ of coeta to beyond middle of doraiua. rather o
potterioT edge from g of costa t« J of doisiun, slightly c
cilia whitish^ochreons. Hindwings ocbreoiu-whitisb, poeterioriy
iiT^^nUrly roSosed with grey ; cilia ochreous- whitish.
Colombia, I a Ouinbre, 6600 feet, in May (Parisk);
one specimen.
Amorbia heliountha, n , sp.
9. 24-26 mm. Head, palpi, and thorax deep purple-femginoiis,
palpi 2|. Abdomen orange. Focewings oblong, coeta towards
base very strongly arched and somewhat roughened with eoales,
then almost straight, apex obtuse -pointed, termen almost Tertioal,
sinuate below apex, rounded beneath; ferruginous-brown, strigu-
lated with dark grey ; markings formed by darker ferraginous-brown
suffusion, with purple gloss, very undefined, strewn with dark gtey
strigulao and some small groups of bluish-grey scales; basal patch
hardly defined, edge veiy oblique ; central fascia moderate, veiy
oblique; a triangular apical patch, its edge running from J of costa
to tomus, somewhat sinuate : cilia ocbreous-brown, with torruginons-
brown basnl line. Hindwing deep orange; an apical spot of deep
ferruginous suffusion, strigulatcd with dork grey; cilia orange, tips
pale, round apex more or less milfuBed with deep ferruginous ; costa
somewhat sinuate before prominent origin of cilia.
Kbemh UriANA, li. Maroiii; eleven specimens. A
sinjrie qj fr«ni the same locality is probably the other sex
iif this specie^ ; tho costa of {ore\viii<:s is much less strongly
arched, with moderately strong fold fn»m base to f, the
hindwinfts dull light fulvous-ochreous, dee)>er posteriorly,
with similar dark apical spot.
ih,Cooi^lc
South American Micro-Lepittoptera. 13
Spa^CinotUs UlumiiutB, n. sp.
$. 20 mm. Head, palpi, and thorax fenogmoua' brownish, palpi
4. Abdomen onnge. Forewingi oblong, costs anterioriy gtrongly
arched, posteriorly nearly gtnight, apex obtuse, tennen Htraigbt,
vertical ; feimginonB-hrawD with ailveiy-iridesceot-vialet reflections,
strewn with tiansverae anastomosing greyiah-ochreons striolae with
Htrigolse of dark fuscous scales adjoining them ; two slightly unnat«
fine greyish -ochreous lines edged posteriorly with dark fuscous scalu,
fintfromlotcostat« I of dorsum, second from g of oosta to tomns :
cilia brownisb-ochreous, en termen with deep Tiolet-ferruginous
batal line. Hindwings deep ontnge, towards apex and upper part
of l«niien ccppery-tinged ; cilia coppery tinged with fuscous.
French GmANA, R. Maroni; one apecimen.
Spargsnothis sabaelds, n. sp.
S- 15 mm. ; $. 22 mm. ^ad, palpi, and thorax greyish-
ochreoos, slightly sprinkled with dark fuscous, palpi in :? 3, in $ 4.
Antennal ciliations of S 1- Abdomen pale greyish-oohreous. Fore-
wings elongate-oblong, costa anteriorly in 3 strongly arched, ah^tly
bent oyer and roughened with scales to beyond middle, with more
maAed subtriangular projection near base, in 9 veiy strongly
arched, posteriorly nearly straight, apex obtuse, t«rmen slightly
sinuate, little oblique ; biownisb^KhreouB, strewn with ferruginous-
brown strigulae ; some blackish strignlae on costa ; in $ a ferruginous-
brown spot on costa at i; costal pat«h elongate-triangular, ferm-
ginous-hrown, in <J marked with black, in $ narrowly produced
anterioriy to before middle of costa; a wedge-shaped ferruginous-
brown streak along apical part of t«rmen, with some black scales :
cilia light brownisb-ochreous, on termen with basal half ferruginous-
brown. Hindwings light dull fnlTous, faintly darker-strigulated,
in ij nther broadly suffused with grey towards dorsum; cilia
whitish-oohreons.
French Gitiasa, R. Maroni; two Bpecimens.
Peronaa apheDotathra, n. ep.
3. IB mm. Head and thonx light greyisb-ochreous sprinkled
with fuscous. Palpi ^, funcoua, somewhat mixed with pale greyish-
ochreouB- Abdomen grey. Forewings elongate, costa moderately
and evenly arched, apex obtuse -pointed, tennen slightly rounded,
oblique i pale greyish-ochreoue sufFused with light brownish; a
wedge-shaped fuscous and grey blotch extending along basal fifth
iM-,i,A J h, Google
14 Mr. E. Meyiick's DeBcriptioTu o^
of costa, reaching at bow to doTsnin; a dork leaden-grey apical
patch irregularly spotted with dark fuscous, its edge numing from
before middle of costa to tomne, nearly straight ; a tratuvene ridge-
tnft on lower half of wing beyond middle, and another abore
torouB ; gome sraall tufta at f above middle and towards apes :
cilia grey, beneath toniiis pole greyiBh-ochreoos. Hindwings with
6 from middle of transverse vein ; rather dark grey, lighter towaids
base; cilia grey.
BiuTiaH Guiana, Baitica, in Febraary [Pariah]; one
specimen. Two other examples from the same locality
ate in all probability the same species, the chaiacteristic
tufts being in the same position ; in these the forewings are
wholly fuscous with slight greenish tinge, sprinkled with
dark fuscous, and more or less largely sufFused with dark
leaden-grey; in one of them there is a cloudy white dot
on costa beyond middle.
Peronea euprata, n. sp.
$. 13 mm. Head and thorax light greyish, patagia tinged with
, coppery. Palpi 2, grey irrorated with whitish. Abdomen grey.
Forewings elongate, posteriorly dilated, costa anteriorly slightly,
posteriorly moderately arched, apex obtuse, t«rmen nearly straight,
little oblique ; light brownish ; a leaden-grey basal patch somewhat
8I»inkled with blackish, especially on edge towards costs, edge
miming from ^ of costa to g of dorsum, slightly sinuate ; a broad
t«rminal patch of deep coppery-brown suffusion, mixed with pnridisfa-
grey suffusion, towards apex with a somewhat oblique trtuisvene
light greyish streak and tome small spots round apical maigin,
spaoe between these mixed with blackish ; small grey totta at angles
(rf cell : cilia brownish irrorated with grey and grey- whitish. Hind-
wings with 5 from middle of transverse vein; dark grey; cilia
grey, with darker Hubbasal tine.
British Guiana, Bartica, in January {Parish); one
specimen.
Peronea flexillnesua. Walk.
British Gutana, Bartica, in April {Parish) ; ones peci-
men. An undoubted example of this widely distributed
Indo- Australian species, which hae not been previously
recorded from America ; it is probably attached to some
tree or shrub of cultivation.
Ij.y.l.Ajl^yCOO'^lL'
South Amancan Miefo-Lepidopttra.
EUCOSMIDAE.
Spllonotft immliHiis, n. ap.
,}9. 11-13 mm. BmA, palpi, and thonx grey irror&t«d with
whitiih. Antoonae in <J with notuh at J. Abdomen AmA giej.
Forewiuga elongate, po«terioiij alighU; diUt«d, coeta slight!; arched,
in $ with strong fold from base fai middle, apex nhtuae, term w slightly
ronnded, rather oblique; Taiying from li^t grej to pale brownish-
ochreone, more or lees iiTomt«d with whitish, with a few blackish
scales; some more or less developed oblique dark fuscous atrignlae
fromcosta; int^coBtalhalf wh<^7 suffused with dark gtey from base
to}or): cilia rather daik grey. Hindwtngs dark grey; oiliagrey,
with daiker sabbasal shade.
British Guiana, Mallali, in March {Parish); nine
specimens.
SpHonola vlridxns, n. sp.
^ $. 12-13 mm. Head and palpi grey, pale-speokled. Antemute
in S with not«h at \. Thorax grey, sometimee tinged with greenish
or partially aaSused with dark fneoons. Abdomen dark grey,
anal tuft in S grey-whitish. Forewinga elongate, poateiiorl;
slightly dilated, coeta gently arohed, in S v>th strong fold from base
U) beyond middle, filled with whitish hairs, apex obtuse, tennen
slightly rounded, somewhat oblique ; grey suffused with dull greenish,
with scattered blackish scales and short variable marks ; costa with
short oblique blackish marks ; costal | from base to 1 in (^ mora or
less wholly suffused with dark fuscous, this area projecting down-
wards as an obtuse-triangular blotch marked with black crossing
fold beneath middle of wing, preceded by a light grey tuft on fold,
dorsal area beneath this mora or leas tinged or suffused with whitish,
in $ tbe lower portbn and black marks of this araa appear as an
irregular median interrupted longitudinal band ; a black longitudinal
median streak from cell to near termen, and some shorter black
longitudinal marks above and below this posteriorly; lateral
margins of ocellus obecnrely indicated with leaden- metallic : cilia
gray sprinkled with whitish and obscurely barred with dark fuscous.
Hindwinga with 3 and 4 stelked ; dark gray, basal area aubhyaline ;
cilia grey, with darker basal shade.
British Guiana, Baitica, December to Februaty
(Parish) ; twenty-seven specimens.
iM:,i,A J 1^, Google
16 Mi. K. Meyrick's DescrifUons a^
Cbocidosema, Ze!l.
I bad abandoned this genus, which, being originally
founded on the single species plebeiana, seemed to be
unnecessarily separated from Etuosma by a structure which
could be regarded aa specific. As, however, I now describe
iive other species undoubtedly allied to it and possessing
the same diBtinguishing peculiarity of structure (the erect
tuft at the base of hindwin^^ in (J), it becomes worth while
to restore the generic validity of the group, which is
evidently characteristic of South America, the wide dis-
tribution of jdebeiana being doubtless due to artificial
introduction. The interesting features of the genus would
be concealed if merged in the extensive and cosmopolitan
Evcosma.
Crooldoseina rorarli, n. sp.
1^ $. 1&-18 mm. Head, palpi, and thorax grey. Abdomen in ij
pale gnyiab-oohreoOB, basal half ochreoua-whitJBh speckled with
black, in ? light grey. Forewings elongate, posteriorly Bomen^t
dilated, coata gently aiehed. in S ^^^ edge shortly folded over at
baae and large eiect tuft of scales beneath it, apex obtuse, tennen
slightly sinuate in middle, somewhat oblique ; ptJe greyish -ochreous,
in S sprinkled with fuscous and on costal half and towards termen
rather suSuHed with fuscous, in $ with dorsal half suffused with dark
fuscous from near base onwards, posteriorly extending to near
coeta; cosla strignlated with dark fuscous, and on posterior h^f
with pairs of indistinct whitish strigulae, two distinct white strigulae
before apex followed by a small dark brown apical spot; in $ a
quadrate whitish blotch on doraum beyond middle more or less
apparent ; ocellus narrow, speckled with whitish, margined laterally
by leaden -metallic streaks reaching more than half across wing and
above by some more or leea developed irregular blackish maiiings,
its apex connected with that of dorsal postmedian blotch by an
oblique grey-whitish streak cutting off between them a triangular
dark fuscous dorsal blotch, in £ less cleoriy indicated : cilia grey
irrorated with whitish and blackish, towards tomus whitish. Hind-
wings grey-whitish, in i^ towards dorsum speckled with black, lower
margin of cell block, tuft rather short but extending nearly to vein 2,
ochreous-whitisb, at base black, in $ suffused with grey towards
termen; cilia whitish, with light grey subbasal line.
Peru, Matucana, 7780 feet, in July {Parish); six
specimens (1 (^. o 5).
ihyCOOl^lC
Micro-Lepidoptera.
Croeidosenu eosmoptlla, n. np.
^. 17 mm. Head ajid thorax grey mixed with blackkh. (Palpi
broken-) Abdomen mostly black (partly defaced), towardsapexpale
ochieons- ForewingB elongate, poBterioriy somewhat dilated, oosta
gently arched, with nanow (old from base to near middle and rough
scales beneath it basally, apex obtuse, termen slightly sinuate ia
middle, somewhat oblique ; gKJ, mixed with whitish and suSused
with doll greenish, on costal { dark riolet-grey ; coata blsckish, on
posterior half with five pairs of whitish strignlaei basal patch
suffused with blackish, undefined, cut by an irregular suffused white
median streak from base, running int« a large trapezoidal median
dorsal whitish blotch whose upper portion projects posteriorly so as
almost to reach ocellus ; ocellus margined laterally by silvery-metallic
streaks and limited above by a curved irregular outwardly oblique
black streak, int«mally speckled with white and containing two
block dots : cilia grey sprinkled with blackish (imperfect). Hind-
wings dark grey, paler and thinly scaled in cell, veins dark fuscous;
toft long, grey, posteriorly whitish, projecting downwards over an
ochreons-yellow space surrounded with a few scattered black scales ;
cilia grey.
Aboentika, R. Parana, in April ; one specimen.
CrooldOHma Impendens, n. sp.
3 ?. 16-17 mm. Head whity- brownish, crown in J s
suffused with dork grey. Palpi whitish. Thorax ochreous-whitisb,
somewhat strigulated with dark fuscous, or in <} anteriorly and
doisally suffused with dark grey. Abdomen whitish-grey. Fore-
wings elongate, posteriorly somewhat dilated, costa gently arched,
in (5 without fold, apex obtuse, termeu slightly rounded, faintly
sinuate in middle, somewhat oblique; pole brovmish-ochreous; in
(J a basal patch of dork fuscous suffusion extended on costal half
almost to apex, on doraal half reaching to I and followed by an
ochreous- whitish quadrate doisal blotch; in $ whole wing longi-
tudinally streaked with dark fuscous suffusion on veins, most
strongly subdorsally, costal and dorsal edges strigulated with
dark fuscous; ocellos laterally margined with indistinct silvery-
metallie streaks, containing three or four undefined block data near
posterior edge ; cilia whitish -ochreous somewhat mixed with dark
fuscous above middle and on base of lower part of termen, on
coBta and a snbtomal spot dark fuscous. HLndwings whitish-grey
TBANS. ENT. SOC. LOND. 1917. — PAIIT I. (NOV.) C
u,y,i,Ajh,Cooglc
18 Mr. E. Mejrick's Deaeriptiont of
Btngnlated witb grey; tuft in <} larg«> dense, ocbreoiu-wiiituh,
basally BufEuaed with fuacons ; cilia whitish, with grey basal line.
Ecuador, Alausi, 9450 feet, in June (Pnmh); three
speciiQens.
Croddosenui c»Ul<l», n. sp.
,3. 17 mm. Head and palpi dark giey, collar paler. Thorax
greyish -ochreoDB, anteriorly and centrally suffused with daric
purpUsb-fnscouB. Abdomen grey. Forewings elongate, postenoriy
rather dilated, costa gently arched, without fold, apex obtuse, ter-
men slightly sinuat« in middle, little obUque ; fuscous suffused with
purplish -grey ; basal patch suffused with dark purplish-fuscous.
eEtending on costa to middle and on dorsum to |, followed by a
quadrat« whitish donal blotch; posterior half of costa with five
pain of obUque whitish strigulae, last more distinct and followed by
a small dark fuscous apical spot ; ocellus filled witb white inoration,
margined laterally with silvery- metallic streaks, and limited above
by an outwardly oblique curved blackish-fuscous streak : cilia dark
grey with rows of whitish points. Hindwings pale grey, veins and
t«rmen suffused with darker grey ; tuft somewhat shorter and more
extended towards dorsum than in phbeiana, grey-whitish, towards
base suffused with blackish ; cilia grey, with dark grey basal line.
Peri'. Choaica, 2800 feet, in July (Parish) ; one specimen.
Croeidosenu ptiUdelplu, n. sp.
S- 13-14 mm. Head grey, face and palpi whitish. Tbom
dark grey. Abdomen grey, anal tuft whitish. Forewinp elcmgate,
costa gently arched, without fold but witb a toft of scales near base,
apex obtuse, termen sinuate, somewhat obUque; grey, sometimes
tinged witb wbitisb-oclireous in disc poeterioriy ; basal patch darilc
fuscous, extending on coeta to i and on donum to middle, edge
obtusely angulated in middle, costal tnft grey or grey-whitish ; a
subquadrate whitish blotch on dorsum beyond this, posteriorly
soffused ; four or five indistinct pairs of whitish strigulae on costs
posterioriy, with obscure oblique leaden stiigae running from them,
a more distinct pair of white strigulae before apex; ocdtns wlitish,
margined laterally with silvery- metallic streaks, enclosing three or
four linear black dots, and preceded by some ochreons tinge, with
a black subdorsal dash : cilia whitish, speckled with blackish ex-
cept on tomns. Hindwings grey-whitish, veins and terminal ana
suffused with grey, acterioHy semibyaline; tuft large, eit«xiding to
basal portion of dorsum, ochreone-grey-whitiah, with a spot of grey
raffusiMianterioriy; cilia grey-whitish, with faint gteysubbasal line.
l„y,|,AJI^,COOl^lC
South American Micro- Lepidoptera. 19
9- 14-16 mm- DiSecB bom 1^ as follows : heAd mostly ochreoiu ;
palpi whitish, Mcond joint euSused with gray towuda apex ; thorax
p*le ocbnooa, putially mixed with grey; forewinga gt«yiah-
ochreoua, paler or whitish-tinged anterioriy, Mmetimes strigulated
with fnscooB ; lower half of basal patch more or leas mixed with
dark fuBcons, remainder obsolete; doreal whitish blotch obsolete;
other roarkings as in ,} : cilia euffuBed with pale ochreouB.
Ecuador, Huigra, 4500 feet, in June; Perl-, Choaica.
2800 feet, and Lima, 500 feet, in July and Atijrust {Parish) ;
fortv-five specimens. The (J is easily distin^shed from
pM>eiana by the costal tuft, dark thorax and basal patch ;
bttt the 9, which differs much in appearance from the o-
would sometimes be very similar to obscurely marked
forms of that species. I have not, however, yet received
jiebeiatta from South America.
Eneosma proMllosB, n. sp.
3 ?. 10-1 1 mm. Head, palpi, and thorax grey, more or less ino-
lated or soffoaed with ochreonB'grey-whitisb, palpi moderate, aub-
ascending, shortly rongh-scaled. Abdomen pale greyish -ochreous.
Forewinga elongate, pOBteriorly slightly dilated, coeta slightly
arched, in 3 without fold, apes obtuse, termen slightly sinuate,
little oblique; dark fuscous iirorated with whitish; costa on
posterior half with five paira of atrigulae of whitish irroration,
fint and third pairs emitting very oblique blue-leaden strigae,
second of these lunning to termen beneath apex, othera with short
blue-leaden marks; a very undefined oblique patch of whitiah
suSuaion or strigulation from middle of doisum reaching half across
wing, aometimea little indicated; ocellus margined laterally by
erect thick leaden -metallic streaks, and enclosing three black dots :
cilia fuscous, with dark fuscous basal lino edged by whitish suffusion.
Hindwings with 3 and 4 stalked ; whitish-grey, veins and a aufTused
terminal band fuscous ; cilia whitish-grey, with darker basal shade.
Peru, Choaica, 2800 feet, and Lima, 500 feet, in July
and August (PamA); forty-two specimens,
Eaeosma evldens, n. sp.
3% 12-15 mm. Head whitish-ochreous, forehead aad collar
sometimes spotted with fuscoua. Palpi porreclcd, second joint
broadly rongh-acaled, whitiah-ochreoua, more or leas wholly suffused
with fuflcouB except on a median blotch on upper portion, terminal
joint fuscous. Thorax dark fuscous, posterior extremity whitiah-
u,y,i,Ajh,Cooglc
20 Mr. E. MejTick'a Deaeriptiona of
ochieoDB, patsgia more or leas suffused with whituh-ochreous.
Ab^men fuacoua. Forewinga elongate, posteriorly slightly dilated,
ooBta gently arched, in <J without fold, apex obtuse, t«nnen slightly
annate, little oblique i brown, more or lees etrigulat«d with daric
hucoua; dorsal area mote or leas miSused with dark fusooiu, with
some indistinct whitish transveise strigalation towards base, and a
postmedian subquadrate blotch formed of four suboonflnent whitish
atrigae not reaching half across wing ; costa dark fuscous obscurely
strigulated with whitish suffusion, posteriorly with more distinct
paired white strigulae, whence arise violet-leaden oblique strigae,
two beyond middle running respectively to anterior margin of ocelluB
and termen above middle; ocellus margined laterally by thick
leaden-metAlho streaks reaching more than half across wing and
closed above by a curved black streak, internally bUckiBb closely
speckled with white : cilia on upper part of termen blockish speckled
with whitish, on lower part ochreous- whitish. Hindwings with 3
and 4 stalked; rather dark grey; oiliagrey.
British Guiajja, Bartica and Mallali, from December
to March {Parish); Dutch Guiana, Paramaribo; Cuba.
Holguin; twenty-two specimenB.
EucostOA operla, n. sp.
,J. 11 mm. Head and thorax grey. Palpi whitish, second joint
with fuscous median spot and apical band. Abdomen fuscous.
Forewings elongate, posteriorly rather dilated, costa gently arched,
with moderate fold from base to beyond middle, apex obtuse, termen
hardly sinuate, nearly vertical; brownish-grey; basal g obscurely
strigulated with dark fuscous ; a dark brown triangular blotch extend-
ing over posterior halt of costa, its apex reaching to lower angle of
cell, marked with an irregular blackish longitudinal streak across it«
middle, a blackish longitudinal mark on its margin above ocelhia,
and on costa with four pairs of indistinct whitish strigulae whence
rise faint oblique violet-leaden strigae; ocellus margined laterally
by thick silvery -metallic streaks, anterior triangularly dilated
upwards to adjoin costal blotch, internally crossed by five fine
blackish dashes separated with whitish : cilia rather dark fuscous
finely sprinkled with whitish, towards tomiis greyish. Hindwings
with 3 and 4 stalked ; rather dark fuscous, near base thinly scaled ;
cilia grey, with darker subbasal shade.
British Guiana, Bartica, in December (P«mA); one
Bpecim«n.
ih,Cooi^lc
South American Micro-Lepidoptera.
Eneosnu religiosa, n. sp.
S. 12 mm. Head white, aides of crown orinuon-fiucoua. FtJpi
Bubascending, white, second jomt fuscous except rough anterior
scales. Thorax white, patagia with a crimson-fnacous bar. Abdo-
men grey, anal tuft grey -whitish. Forewings elongate, posteriody
dilated, costa gently arched, without fold, apex veiy obtuse, termen
somewhat prominent, slightly rounded; fuscous suffused with dull
crimsoD, mixed with blackish in disc; a broad ochieous' whitish
dorsal band occupying neaiiy half of wing, upper edge very inegular ;
costal edge blackish with paits of whitish striguke , a violet-leaden
striga from ^ of costa to terraen below apex ; ocellus limited beneath
and poaterioriy by short silvery- metallic streaks not united, contMn-
ing three ahort black marks partially surrounded by whitish : cilia
whitish, on upper part of termen with reddish -fuscous subbasal lino,
on tamus spotted with reddish. Hindwinga with 3 and 4 connate;
rather dark grey ; cilia grey, with darker sobbasal shade.
Beitish Guiana, Bartica, in January {Parish); one
specimen.
EncDsina ktrieapUla, n. sp.
$. 12 mm. Head, palpi, collar, and shoulders dark fuscous,
thorax otherwise ochreons- whitish, palpi asoending, shortly scaled.
Abdomen dark grey. Forewings elongate -oblong, costa towards
tase rather strongly arohed, then nearly straight, apex obtuse,
termen hardly sinuate, nearly vertical; pale pinkish-ochreous,
somewhat suffused with whitish anteriorly, and strigulated with
violet-grey ; a, large rounded-triangular dark fuscous bloteh on
dorsum slightly before middle, reaching more than half across wing ;
an oblique ferruginous spot on middle of costa, whence a line of very
tine transverse black and white striolation runs to near termen
beneath apex, thence downwards to middle of termen, closely
followed on subcostal portion by a very fine violet line ; coata
beyond this with three small deep ferruginous marks separated by
white strigulae, and then a wedge-shaped deep fciruginous spot
occupying apical portion and cut transversely in middle by a dark
leaden mark : cilia pale ochreoue. with interrupted violet-groy
baa^ line, round apex dark brown with blackish basal line. Hind-
wings with 3 and 4 connat« ; dork gi«y; cilia grey, tinged with
whitish towards tips.
British Guiana, Baitica, in Febraary (Parish); one
specimen.
iM-,i,A J h, Google
22 Mr. E. Meyrick's Descriptions ^
Euoosma digni, n. sp.
S- 11-12 mm. Head, palpi, and thorax fulvous, palpi ascending,
Bbortly scaled. Abdomen dark grey. Forewings elongate, pos-
teriorly somewhat dilated, costa gently arched, without fold, apex
obtuse, termen nearly straight, almost vertical ; fulvous, irregulariy
ietioulat«d with rather dark violet-fuacous, costal area on posterior
halt suffused with deep ferruginous ; costa strigulated with blackish,
and on posterior j with whitish ; an iiregolar blackish line, angu-
lated in middle and suffused with deep ferruginous, from | of costa
to J of dorsum ; a slender oblique blackish streak from middle of
costa reaching nearly half across wing, somewhat angulated near
its extremity, with a tine acute projection from angle posteriory,
followed by two small block marks; a short oblique blue-leaden
striga running to tennen below apex, finely edged with ochreous-
whitish beneath; ocellus Lmited beneath by a leaden dash and
posterioHy by a leaden -metallio streak edged externally with
ochreoua- white, and enclosing three strong black dots; oilja fulvous,
on t«rmen with a leaden line, on tomus spotted with dark grey at
base. Hindwings with 3 and 4 connate or short-stalked; dark
fuscous ; cilia grey, with darker subbastJ shade.
British Guiasa, Bartica, in December {Pamh); two
specimens.
Eueosraa ortygia, n. sp.
35. 17-18 mm. Head, palpi, and thorax whitish -ochieous
variably spotted or suffused with reddish -fuscous, palpi ascending,
shortly rough-scaled. Abdomen dark grey. Forewmgs elongate,
posteriorly dilated, costa gently arched, without fold, apes obtuse,
lermen straight, nearly vertical; whitish -ochrcous strigulatcd with
rcddlsh'fuscous mixed with dark fuscous, towards costa and pos-
teriorly or sometimes wholly suffused with ochrcous-bronD ; costa
strigulated with blackish and posteriorly with whitish; short
violet-leaden marks from posterior pairs of whitish strigulae, and
a striga from | of costa to termen beneath a]icx; ocellus limited
beneath by a dark fuscous dash and posteriorly by a thick leoden-
metollic streak edged externally with ochreous -whitish, containing
three strong black dots or short dashes : cilia whitish-ochreous with
reddish-brown line, broken up on tornus. Hindwings with 3 and 4
connate; dark fuscous; cilia grey, tinged with whitish towards
tips, with darker subbasal shade.
British Guiana, Bartica and Mallali, from December
to March {Parish); eleven specimens. The allied species
transferrana. Walk. {= descriplana, Walk. = vincentana.
ll,y,l,/eJh,G00t5lc
South American Mtcro-Lepidoptera. 23
WaU.). metaapilana. Walk., and leucomesana, Walk., were
all received from Bartica in good series.
Euoosma prudeiu, n. sp.
t? $. 16 mm. Head in ^ ochieooB-wbitiBh somewhat spotted with
duk fiucfxis, in $ ochieons spotted or almoet wholly suffosed with
dark fnscona. Folpi subsacending, shortly rougb'Scaled, whitish,
wcond joint soffiued with reddish 'fuscous except towards apex,
baw of terminal joint dark fuscous. Thorax whitish more or lees
mixed or suffused with fuscous, anterior fourth dark fuscous. Abdo-
men dork grey, anal tuft in ^ grey- whitish. Forewings elongate,
posteriorly dilated, coeta gently arched, without fold, apex obtuse,
termen somewhat sinuate, vertical; fuscous, sprinkled with dark
fuscons and blackish, in ^ irregularly mixed and sufFusedly striated
with whitish, doisal half in $ suSnsedly striated with violet-leaden-
grey; costa marked with paira of whitish atdgala«, and small
blackish spote between these; edge of hssal patch indicated by
an irregular dark fuscous streak, ongulated in middle, across fold
with a blockish mark edged posteriorly with white ; central fascia
Teiy irregolar, obUque, dork fuscous, suffused, only distinct on costa ;
four bine-leaden oblique marks from pairs of costal whitish strigulae
posteriorly, second running as a striga to termen beneath apex, edged
beneath with white posteriory, others short; termen nonowly
fnlvous -brown, with a fine oblique white stT^:ula before apex;
ocellus Umited beneath by a whitish line and posteriorly by a
leaden- metallio streak edged eztemally with white, and enclosing
two btaok dote : cilia whitish, greyish towards tips on termen, with
a leaden sabbasal line broken on temus. Hindwings with 3 and 4
connate ; dork grey ; cilia grey- whitish, with grey subbasal line.
Peru, Lima, in August {Parish) ; three specimens.
Polyehrosis araucea, n. sp.
^ $. 11—13 mm. Head, paJpi, and thorax ochreous- whitish or
whitish-ochreous. Abdomen grey, anal tuft of ^ grey-whitish.
Forewings elongate, posteriorly somewhat dilated, coste gently
arched, apex obtuse, termen nearly straight, rather oblique; pale
ochreons or whitish-ochreous ; costa with whitish strigulae, more or
less sprinkled with blackish between them ; dorsal edge with very
small blackish strigulae; a more or less developed blotch of darker
suffusion about fold indicating angle of basal patch ; an oblique rather
narrow deeper ochreous or brownish fascia from middle of costa
reaching half aoroea wing, and some undefined sufCusion towards
iM:,i,A J 1^, Google
24 Mr. E. Meyrick's Dexcri'pliont of
domim beyond middle, seldom forming & diatinct tranarene blotch ; '
Hometimes a patch of blAckieh irrorKtion on coata beyond thia, or an
oblique streak of blackislr irroration from above middle of disc to
dorsum before tomns, or more undefined blackish irroration towards
dorsum generally and across wing beyond middle, all very Tariablo
and sometimes wholly absent ; a deeper ochreoua or brownish
transverse spot on tomus, and a wedge-shaped upwards-ohUque
streak from middle of termen, both often very faint; a slender
streak of ftne blackish transverse striation along termen : cilia
whitish or whitish -ochreous, with ochreous subbasal shade. Hind-
wings groy-whitiBh, thinly-scaled, veins and termen suffused with
grey ; cilia whitish, with faint grey aubbasal line.
Ecuador, Duran, low swampy couotr)', in June {Parish) ;
twenty-three specimeriB. Varies remarkably, but easy of
recognition.
Polychrosis oxymochla, n. sp.
3 $. 12-14 mm. Head, palpi, and thorax dark grey. Abdom^
pale grey, anal tuft of ,$ grey-whitish. Forcwings elongate, pos-
teriorly rather dilated, costa gently arched, apex obtuse, t«nnen
almost straight, rather oblique; dark fuscous finely speckled with
whitish, variable in development ; costa strigulated with blaokish-
fuscooa, with some small spots; basal patch dark grey, edge blackisb-
fuBCOUB, angulated in middle, oft«n interrupted above angle, some-
times followed by a fascia of grey-whitish suffusion; an oblique
rather narrow blackish -fuscous fascia from before middle of
coeta, reaching more than half across wing, extremity pointed; a
quadrate blackish -fuscous blotch on dorsum before middle: a
moderate blackish -fuscous fascia from below middle of termen
obliquely upwards towards f of costa but not reaching it : cilia
rather dark fuscous speckled with whitish, with blackish-fuscous
subbasal line. Hindwings grey, lighter towards base, vems and
termen darker-suffused ; cilia pale grey, with dark groy subbasal line.
Perc, Lima, in August {Parish); twenty-eight speci-
mens. Varies considerably in distinctness of niarkin<^:
nearly allied to the preceding, notwithstanding difference
of colouring.
Bactn serla, n. sp.
i. 18-21 mm. Head, palpi, and thorax pale brownish -ochreous.
palpi 3. Ant«nnal ciliations }. Abdomen light grey, anal tuft pale
greyish -ocbreons. Forewings elongate, rather narrow anteriorly,
posteriorly rather dilated, costa slightly arched, apex obtuse-
l„y,l,Ajh,COOl^lC
SoMh Ammoan Micro-Lepidoplera. 25
pomted, tennen faintly sinuate, rather oblique; whitish-ochreoos
tinged with browniah, tending to be nuu^ed with fine fnacoiu lines
on veins, and fine brownish lines between veina; costs obUqnely
Rtriguloted with brownish and fnaoous ; dorsal edge with a series of
cloudy dark fnscous dots or marks ; some dark fusoona irroration
towards lower margin of cell, tending to form dark fuaoous lines on
veins, with a more or less developed irregular dark fuscous spot at
}, sad one or two cloudy 6eak fuscous marks on end of oelli a
more or leas developed triangular spot of fuscous suffusion on
lennen bmeatb apex : cilia whitish-odueooa sufhised with grey,
with obscure dark grey bars. Hindwings whitish-grey or light grey,
somewhat darker posteriorly ; cilia grey- whitish, with grey snbbaml
Peru, Aqualani; three Bpecimens.
Ai^ropIoM fueta, n. sp.
0*9' 13-14 mm. Head, palpi, and thorax fuscous -whitish suf-
fnsedly spotted or marbled with red-brown. Abdomen dark giey,
anal tuft of 3 grey-whitish. Forewings rather elongate-triangular,
costa moderately arched, somewhat bent beyond middle, apex
romided, termen rounded, somewhat oblique; ochreous-whitish to
light of^treoua-brownish, irregutarly strigulated with brown-reddish
and leaden-gi«y; basal patoh and central fascia coarsely strigulated
with black but very indefinite, on coste suffused with dark tad-
brown, edge of basal patch irregular, somewhat curved, central
fascia l»oadeDing downwards, rather oblique ; beyond central fascia
a very irregular blue-leaden stria; posterior half of ooeta black with
vrhite strigulae, whence rise short blue-leaden marks, and a sinuate
bhie-leaden stnga from jj of costa running to termen beneath apex.
snbcoatal area round these deep fulvous or ferruginous; beneath
this a cnrved dark red-brown fasoiate psteh crossed by black lines
on veins extending to near termen below middle, its lower extremity
followed by a blue-leaden terminal mark ; a very fine black terminal
line ; cilia pale red-brownish, deeper towards base, spotted with
lesden-grcy. Hindwings with 3 and 4 connate; blackiah-gi«y ;
cilia grey, with dark grey subbasol shade.
Bnmsii Guiana, Bartica, from January to April
(Parish)', five specimens.
Argyroplooe oataphracta, n. sp.
ij$. 14^16 mm. Head, palpi, and thorax dark fuscous, head
tinged with crimson. Abdomen dark grey. Posterior tibiae in ^
u,y,i,Ajh,Cooglc
26 Mr. E. Meyrick's Desa-tpUom of
clothed with long dense whitiBh haira above. Foiewin^ auboblong,
rather broad, coeta anteriorly strongly, posteriorly slightly arohed,
apex rounded, tennen rounded, slightly oblique; dark crimson,
fuscous; hasal patch, lower half of central fascia, and an imgnlar
patch above tomus marbled with dark blue-leaden; a whitish-
ochroous elongate- triangular patch ezt«nding over poBt«rioi' half
of costs, more ochreonB towards costa, anterior side tongeet, marked
with an iridescent- violet-silvery streak dilated downwards, costs)
edge deep ferruginous with five pairs of oblique white strigulae, a
blue-leaden striga running from fourth pair obliquely across apex,
apex suEFused with deep ferruginous round this; a very fine black
terminal line, partly edged with silvery ant«rioriy : oilia leaden-
grey, basal half deep ferruginous except towards tomfta. Hindwings
with 3 and 4 comiate ; dark grey ; cilia grey, with dark grey haaal
shade; dorsal edge in S thickened and rather contorted, with long
British Guiana, Bartica, from December to February
(Parish); seventeen specimens.
Argyroploce eymotoma, n. sp.
iJQ. 13-14 mm. Head whity- brownish, sides marked with dark
red-brown. Palpi whitish, second joint suffused on lower part with
grey, on upper part with red-brown marked with black. Thorax
whitish mixed withgrey and irregularly spotted with dark red-brown,
Bometimes much suffused with dark fuscous. Abdomen dark grey,
anal tuft of S grey- whitish. Forewings elongate, posteriorly dilat«d,
costa moderately arched, apex rounded-obtuse, termen rounded,
nearly vertical ; red-brownish ; baa^ patch mixed and strigulated with
white, spotted with black on its lower portion, suffused with fuscous
on dorsum, marked posteriorly on costa with a dark red-brown
blotch ; beyond this a somewhat angolated fascia of white striation,
on costa forming a more conspicuous white blotch, followed on
coHta by a dark red-brown blotch, narrow on costa aad produced
posteriorly into a long acute prominence ben^th costs, and on
domum by a triangular brown blotch edged with white; costa
posteriorly dark red-brown with pairs of whitish strigulae, with
more or less orange suffusion beneath these, and a sinuate blue-
leaden striga running from edge of costel blotch to termen beneath
apex, edged beneath with whitish posteriorly; ocellus narrow,
whitish -speckled, containing three black marks and enclosed
laterally by broad' erect spots of pale purplish iridescence : oilia
red-brownish, on termen darker towards base. Hindwings with
Ij.y.l.Ajl^yCOO'^lL'
SotUh American Micro-Lepid&plem. 27
3 and 4 connate, 5 very closely approumatod ; dark grey; oilia
grey, with darker aubfaosal shade.
British Guiana, Bartica, in January and February
{Parish) ; five specimens. Allied to alinmacula, Wals.
Argyroplooe nonwei, n. sp.
1$ $. ft-lO mm. Head, palpi, thorax, and abdomen dark leaden'
grey. Foreiringe eloDgate-triangular, coata gently arched, spei
obtnie, tennen rounded, little oblique ; dark leaden-grey; markinga
blackish-fuscous; small spots along coata, separated by gioupa of
two or three very fine indistinct whitish strtgulae; some scattered
marks and Btrigulae on baaal area; an irregular streak from } of
cOHta to j of domum, sharply angulated in disc; central fascia
narrow, oblique, nearly straight or slightly curved, in one specimen
thicker on lower half 1 ocellas containing three black dots, and closed
above by a rounded -triangular blotch : cilia grey, with dark fuscous
aubbaaal line. Hindwings with 3 and 4 connate; dark fuscous;
cilia whitifih-grey, with dark fusooua subbasal line.
British Guiasa, Bartica, from December to February
(Parish); six Bpecimene.
Argyroploee cyelidios, n. sp.
^. 12 mm. Head, palpi, and thorax grey, patAgia terminating
in expansible tufts of broad scales. Abdomen dark grey. Fote-
wingB elongate, posteriorly dilated, costa gently arched, apex
obtuse, termen rounded, nearly vertical; grey, mostly aulTused
with glossy blue-leaden between the markings; markings blackish-
fuscous, finely edged with white laterally and above ; some small
marks and spots along costa; an irregular subcostal streak from
base to 1 : a large rounded blotch extending on dorsum from near
bsae to middle, and reaching more than half across wing; a smaller
rounded praetomal blotch, not teaching half across wing; an
irregular rather thick bisinuate streak from above middle of disc
to near termen above middle : cilia grey, on termen with blaekish
basal line. Hindwings with 3 and 4 connate; dark fuscous; cilia
whitish-gKy, with dark fuscous subbaa&l line; dorsum with a short
much thickened lobe clothed with soalee.
British Gxhasa, Bartica, in December {Parish) ; one
specimen.
u,y,i,Ajh,Cooglc
28 Mr, E. Meyrick's Deacripliom of
Argyroploce platyzona, n. sp.
t^. 16 mm. Head, palpi, and thorax blackish -fuscoua. Abdomen
dork grey. Fotewingi suboblong, moderately bioad, costa moder-
ately arched, apex obtuse, tennen rounded, little oblique; rosy-
ochreoDB, slightly sprinkled with daik fuscous on veins posteriorly;
coeta with small blackish marks edged with whitish; basal patch
blaokish-fuscoue, edge nearly straight, space between this and cen-
tral fascia suffused with ochreous-whitish, of even width; central
fascia blaokish-fuscous, moderate on casta, becoming very broad
downwaids, anterior edge straight, posterior evenly curved, finely
edged with whitish; a fine black whitish-edged streak along upper
part of termen, with a strong oblong projection inwards beneath
apex; a few leaden-grey strigulae in region of ocellus : cilia dark
grey with blackish basal line, with Ught rosy-ochreous tomal patch.
Hiudwings with 3 and 4 connate ; dark grey ; cilia grey, with darker
basal line.
Feench GriANA, R. Maroni; one specimen.
Argyroploce Impolita, n. np.
S- 24 mm. Head, palpi, and thorax rather dark brownish, tinged
with reddish, especially on patagia. Abdomen grey. Porewinp
lather elongate -triangular, moderate, costa moderately arched, apex
obtuse, tennen haidly sinuate, little oblique ; rather dark brownish
marbled with bluish-leaden on basal half ; basal patch edged with
a suffused dark fuscous streak, obtusely angulated on fold; two
pairs of whitish Btrigolae on costs between this and central fascia,
and a blotch of whitish suffusion beneath these; central fascia
broad throughout, dark fuscous, oblique, foUowed on upper portion
by orange-ochtcoua sufFusion more extended posteriorly beneath
coata, and on tomal area by a patch of whitish suffusion; costa
posteriorly dark fuscous, with pairs of indistinct whitish strigulae;
apical area ochreous-browDish : cilia fuscous, on tornus mixed
with whitish. Hind wings with 3 and 4 connate; grey; cilia grey,
towards tips whitish -tinged.
Colombia, San Antonio, 5800 feet, in November; one
specimen.
GELECHIADAE.
Anomoxena, n. g.
Head smooth; ocelli absent; toitgue developed. Ant«nnae g,
in i^ simple, basal joint moderate, without peoten. Labial palpi
iM:,i,A J h, Google
Soath American Micro-Lepidopiera. 29
Icxig, Tecmred, second joint thiokened with whorls of scaleA TOUghty
projecting beneath or with whorla forming separate aonte projecting
teeth of BCaJea, terminal joint as long as second or somewhat shorter,
slightly thickened with scales, acute. Maxillary palpi minnte,
filiform, appressed to tongue. Posterior tibia« clothed with rough
projecting hairs above aitd beneath. Forewings with 16 furcate,
2 from angle, 2-5 parallel, transverse vein very oblique outwards
from 2 to fi, faint between 5 and 6, 6 and 7 stalked, 7 t« oosta, 8
separate, appnndnutted to 6 at hose, 11 from near middle. Hind-
wingB g, narrow-Uapemidol, apex acute, more or less strougly pro-
dnoed, termen emarginate beneath apex, cilia 4; 2 remote, 3 and 4
nearly parallel, 4 from angle, 4 and 6 somewhat approximated, 6
and 7 approximated at base.
Type sjnnufera. A remarkable genus, diiTeriiig from the
whole of the family in having vein 8 of the forewings
separate instead of stalked with 7; the atructiire is un-
questionable, since all the veins are present, and is alike in
both species. Nor can there be any question that the genus
belongs to this family, and to the most advanced type of
it, since the highly characteristic hindwin;^ (similar to
those of Arislotelia) are foimd in no other. The genus must
be supposed to have arisen from Aristotdia or a form of
similar structure, and the peculiarity of aeuration can be
explained by the reduction and eventual absorption of
the stalk of 7 and 8 and apparently in this way only.
I regard it therefore as a proved instance of a atruc-
tuial change which is certainly rare, and would usually be
difficult of demonstration.
AnomoMDE splnlgers, n. sp.
S $. 10 mm. Head ochreous-wbitish, crown with a few dark
fnscons specks. Palpi whit«, second joint with eight rather long fine
whorl-teeth edged with block above, terminal joint with block rings
near boM and above middle. Antennae white ringed with black,
towards apex with seveiaJ norrow grey black-dotted bands separated
by single whit« rings. Thorax ochreous-wbitish, dorsally finely
striated transversely with blackish. Abdomen blackish-grey.
Forewings narrowly elongat« ■ lanceolate ; pole ochreous irr^ularly
irrorated with blackish; an inwardly obUque slender fascia of
blackish suffusion from costa at jj, not ot scarcely reoching dorsum ;
costal area from this to apex fulvous, without dark irroration. with
four oblique fine wedge-shaped pole ochreous streaks posteriorly
edged with black : cilia grey, round opex whitish with three curved
iM-,i,A J h, Google
30 Mr. E. Meyrick's Descriptions oj
daj-k grey lines and a straight nther oblique black bar at base.
HindwingB dork grey ; cilia gray.
Colombia, La Crumbre, 6600 feet, in May {Parish) ;
three specimens.
Anomoxena tetraxoa, n. sp.
(} ¥. 8 mm. Head and thorax nhitish-ochreous with a few dark
faacoiu specks. Palpi white, second joint with four oonfnaed whorifl
towards apes roughly ptrojecting beneath, irrorsted with dark fuscous
on these, terminal joint with basal and median rings of dark fuscous
iiToration. Antennae white ringed with dark grey, towards apes
with bands of two dark rings separated by single whit« rings. Ab-
domen grey. Forewings narrow-lanceolate ; light ochreous irrorated
with dark fuBcous; a slender inegular inwardly oblique fascia of
blackish sufFusion from f of costa, and undefined blotches at tomus
and apex; costal area posteriorly narrowly orange, with four short
slender oblique pale greyish-ochieous streaks edged posterioriy
with blackish ; cilia light giey, ronnd apex speckled with black,
with a rather oblique straight black basal line acrosa apex. Hind-
wings and oilia light grey.
Ecuador, Huigra, 4500 feet, in June {Parish); four
specimens.
Arlstotollk aulonota, n. sp.
^ $. 7-9 mm. Head and thorax ochreous -white, pata^ dork
fuscous. Palpi white, baaol f of second joint dark fuscous. Abdo-
men grey, apex whitish. Forewings elongate -lanoeolat« ; 7 and 8
out of fl; dark fuscous, slightly pale-freckled except on edge of
dorsal streak ; a rather broad whitish -ochreous streak along dorsum
from base to beyond tornus, posteriorly pointed, upper edge with
two OF three slight irregular prominences ; a small whitish -ochreous
Spot on costa at f ; cilia fuscous, blockish -sprinkled, towards tomua
ochreous -whitish. Hindwings light grey, in j with expansible pencil
of long ochreous- whitish hairs from costa near base; cilia grey-
whitish.
Ecuador, Duran, low countn.', in June {Parish) ; three
specimens.
Ailstottlia plumabt, ». sp.
39' 10-11 mm. Head and thorax glossy grey or dark grey.
Palpi dark fuscous, very finely lined with white irroration, terminal
joint longer than second. Abdomen grey. Forewings elongate.
Ij.y.l.AjhyCOO'^lL'
South American Micro-Lepidoptera. 31
naiTow, costs moderately uched, apex pointed, terraen extremely
obliqnelj rounded; 6 separate; grey, sprinkled with blaokiah and
tinged hen and there with whitish ; pUcal and second diacal stigmata
moderate, achreous, edged with black marks above and beneath,
first diBcal small, block, obliqnely beyond plical; tJiiok cloudy
femiginonB-blBckish obliqne tasciate ban from coeta at J, }, middle,
and }, reaching aboat hall across wing, second terminated by plical
■tjgma, third shorter but with a more oblique projection reaching
to apex of fourth, both these terminated by second discal stigma,
fourth being little oblique; a cloudy darker spot on tomus; an
irregnlor blackish praeapical blotch : cilia light greyish, with two
blackish lines. Hindwings grey; cilia light grey; in ^ with two
or three expansible enormonsly enlarged shuttle'Shaped pearly
Bcalea (i length of wing) from towards base l3dng along oosta above,
and a very large elongate chitinous process projecting obliquely
from costa near base, beneath black and concave to receive a fine
pencil of long hairs, partially covered beneath by a fringe of much
enlarged flat pearly prismatic scales, the remainder covered by a
fringe of very long ecalee projecting downwards from beneath costa
of iorewings on basal |.
British Guiana, Bartica, in December and January
{Parish); eight specimens. The curious and exception-
ally complex apparatus for protecting the scent- producing
haiipencil of the hindwings suggests that the perfume is
in this species unusually volatile and precious.
Arlstotolla eryeina, n. sp.
S $. ^10 mm. Head whitish -ochreous, sometimes sprinlded
with fuscous. Palpi ochreous-nhitish, second and terminal joints
each with two dark fuscous bands. Thorax pole greyish -ochreous
sprinkled with dark fuscous. Abdomen whitish -ochreous more
or less suffused with grey. Forewings elongate, narrow, oosta
slightly aiohed, apex obtuse -pointed, termen extremely obliquely
rounded; 6 separate; pale greyish, snffusedly irrorated with dark
fuscous, irrognlaily tinged and spotted with light rose-pink; a very
obliqne suffused blackish streak across fold about I, and an oblique
blackish streak from costa to pLical stigma, space between these
and towards doisnm anteriorly sometimes suffused with pale ochre,
ous; stigmata black, rather elongate, plical obliquely before first
discal; two or three small pole marks on costa towards ajiex; a
more or le«s developed longitudinal blackish streak from second
discal stigma : cilia pale greyish, with blackish subbaaal line round
apex interrupted with pale losy-tinged spots on tomus, and dark
iM:,i,A J 1^, Google
32 iSr. E. Meyrick's Descriptions of
grey postmedian shade. Hindwings grey, in i^ with a HtreaJi of vwy
fiiw bUckish-grey itriation along eubmedian groove ; cilia light grey.
Forewings in ^ beneath with costal edge slightly refleied and rough-
ened from near base to f, with a more or lesa developed patch of
pale ochreous towards this anteriorly, rest of wing euCFiued with
rather dark fuscous. Hindwings in 3 beneath greyiah^ochreooa,
■uSused with rather dark fuscous towards costa posteriorly, and
with a atieak of very fine blackish striation eJong submedian groove
Ecuador, Huigra, 4500 feet, in June; Peru, ChoHica,
2800 feet, in July (Parish) ; twenty-five specimeiia.
AristolelU cyttaeiMa, n. sp.
(J $. 8-10 mm. Head pale greyish .ochieouB. Palpi ochreoua-
whitish, second and terminal joints each with two dartc fuscous
bands. Thorax pale greyish KKhreous sprinkled with dark fuscous.
Abdomen grey, anal tuft of 3 whitish Kwhreous. Forevringa elongate,
narrow, costa gently arched, apes obtuse -pointed, t«niien extremely
obliquely rounded; 6 separate; pale greyiah-ochreous suffosedly
irrorated with dark fuaooUB, more or less tinged with rosy-crimson,
especially posteriorly; stigmata strong, blackish, somewhat elon-
gate, more or lees accompanied with spots of yellow .ochreous sufiusioo
beneath and second discal also above, pUcol obliquely before Siat
discal, an oblique suffused dork fuscous streak from costa to pliool ;
two small pale spots on costa at | : cilia pale ochreous, with blackish
Bubbaaol line interrupted on tomus with pale sometimes rosy-tinged
spots and grey postmedian shade, towards tomus greyish-tinged.
Hindwings grey; in ^ with expansible fringe of long grey-whitish
hairs from costa near base; cilia light grey. Forewings in 3 with
costal edge on anterior half ochreous- whitish and folded over
beneath.
Colombia, t'ali, 500 feet, in May (Parish) ; thirty
flpecimens.
Arlstotella eynthia, n. ap.
3 $. 10-12 mm. Head whitish-ochieous. Palpi whitish, second
and terminal joints each with base and two t>anda iirorat«d with dark
fuscous. Thorax pale ochteous irrorstod with dark fuscous. Abdo-
men grey, anal tuft of a P^lc ochreous. Forewings elongate, narrow,
costa gently arched, apex pointed, termen extremely obliquely
rounded; fl separate-; white irrorated with dark fuscous, doiaal
half suffused with grey and partially mixed with ochreous; broad
Ij.y.l.AjhyCOO'^lL'
Souih American Mioro-Lepidojilera. 33
blackuh-fnscons obliqoe ban from costa at i and ^ to fold, aooond
m&rgined beoeatb by & brownish -ochreona mafk; a BubtruDguIar
flark tuocoue blotch on costa beyond middle, tenniuated beneath by
a brownish-ochreoOB mark; a BufiuBed dark fuscous apioal blotch,
sometimes connected with preceding in disc but separated from it on
coHta by a whitish spot : cilia whitish, sometimes with faint loej or
ochreous tinge at base, with two wide sometimes oonfloent grey
shades, on coeta wholly dark grey. Hindwings dark slaty-grey,
pale in cell; cilia in <; whitish-ochreoos, in $ light greyish; in ^ on
basal hall of costa with eztremdy long expansible cilia. Forewittge
and hindwings beneath'in both sezes blackish-tinged, (orewings in
^ clothed with ochreona bain in disc.
Pebd, Lima, in August (Parish) ; sixteen specimens.
AristotelU satnrnlna, n. sp.
<J$. 11-12 mm. Head, thorax, and abdomen grey. Palpi
whitish or sometimee niffuaed with grey, second with two dark
fnscons bands, terminal joint considerably longer than second, suSus-
edlj lined with dark fuscous. Foiewings narrowly elongate-lanceo-
late; 6 separate; rather dark fuscous, sometimes whitish-speckled,
dorsal area more or less sufiused or marked with ochreous; obscure
darker oblique bars from costa at } and i to fold, matked with black
on fold, more or less edged with whitish towards one another ; discal
stigmata blackish with a suffused hrowniah-ochreous streak beneath
them ; a suffused pale brownish-ochreous spot on costa at { : cilia
grey or grey-whitish, with two darker grey lines, towards base
sprinkled with blackish or dark fuscous in indistinct bars. Hind-
wings grey; cilia light grey; in (^ with basal portion of costa cloth^
with much longer expansible pale greyish cilia.
Pkbu, lima, in August {Parish) ; ten specimens.
AristotelU perfossa, n. sp.
3 9- 10-12 mm. Head and thorax ochreous- whitish irrorated
with foscons. Palpi whitish, second and terminal joints each with
base, two bands, and tip dark foscous. Abdomen grey, anal tuft
of 3 whitish -ocbreons. Forewings elongate-lanceolate; 6 separate;
whitish irrorated with dark grey; oblique bara of blackish suffusion
from costa at ^ and ^, just crossing fold, plical stigma forming an
elongate black mark on end of second; discal stigmata black, first
obliquely beyond pUcal, second indistiQctly edged with ochreous
beneath ; a suffused white spot on costa at j ; sometimes some indis-
tinct ochreous marking near tomus ; cilia whitish, round apex with
TKANS. ENT. SOO. LOND. 19X7. — PART I. (NOV.) D
iM-,i,A J h, Google
34 T£i. £. Meynck's Deacriptioni of
two dark fnacous Udm. HiiidwiDgigrey; cilia pole greyiob. Hind-
wingi in S bwMth with ezponatble bruah of long light oohieouB Iwiim
beneath basal portion of coato, and ainular hain clothing doisum aod
tennen from base to beyond middle of wing.'
Ecuador, Huigra, 4S00 feet, in June ; Peru, Chosica,
2800 feet, and Lima, in July and August (Parish) ; thirty-
five apecimene.
ArlstotoUa piphfa, n. sp.
S $. 10-12 mm. Head roej-whitiBb, more or teas or eometiiBea
wholly Bufimed with grey. Palpi white, with two banda of seoond
joint, and two bands and anterior edge of terminal joint blackish.
Thorax grey, sometimes rosy-tinged, shoulders darker. Abdomen
grey, apex yellowish or rosy. Forewinga narrowly elongate-lanceo-
late ; 6 separate ; rather dark grey, dorsal area more oi leas mixed
tnffosedly with light crimson-pink and sometimes with light oohie-
ous; blackish oblique bars from costa at | and i, just crosmng fold,
first posteiiorly and second on both sides more or leas distinotly
edged with whitish tinged with criinBon-pink ; a rosy-white mark on
middle of oosta; discal stigmata black, first sometimca oonneoted
with apex of preceding bar, second larger, edged beneath by a light
yeUow-ochieous spot; a pale rose-pink spot on costa at j, white
on costal edge : cilia whitish-grey, with three dark grey lines, tips
whitish. Hindwings grey; cilia light greyish.
Peru, Chosica, 2800 feet, in July {Parish); sixteen
ArlstotoUa aphUIra, n. sp.
3 $- 10-12 mm. Head fuscous more or less mixed with whitish.
Palpi Tety long and slender, whitish, second and terminal joints
each with base and two bands dark fuscous. Thorax fuscoua nuxed
with dark fuscou«, somewhat sprinkled with whitish. Abdomen
grey, anal tuft whitiah. Forewin^ narrowly elongate-lanceolate;
6 separate ; grey sprinkled with dark fuscous and whitish, dorsal area
suffused with brownisb.ochreous ; oblique dark fuscous bars from
costa at i and i to fold, space between these sometimes more mixed
with white; a dark fuscons trapezoidal blotch narrowed downwards
on costa about {, edged beneath by a brownisb-ochreous spot, beycmd
this a white spot on costA, produced along margin of blotch, followed
by a dark fuscous apical blotch : cilia whitish with two grey shades,
on base spotted with dark fuscous irroration. Hindwings grey ; cilia
pale greyish.
Peru, Matucana, 7780 feet, in July (PomA); th»e
l„y,|,AJI^,COOl^lL'
South Anurican Micro-Lepidoplera. 35
Bpeumens. Two other examples (,J$) from La Crumbre,
6000 feet, Colombia, I cousider also identical ; the (J has
the vhite markings towards apex tinged with rosy.
Another ^ specimen from Alauei, 9450 feet, Ecuador, may
probably be the same species, but ia in poor condition.
Arittotella ndleata, n. sp.
S $. 9-10 mm. Head pale ochreona, more or leis Bprinkled with
fnacons. Palpi whitish, Becond joint with three, terminal joint
iritb four dark fnscouB rings, terminal joint longer than second.
Thorax pale ochreons iirorated with dark grey, ahoulders with a dark
foHMHU spot. Abdomen grey, apex pale gteyish^Kihreous. Fore-
wings elongat«-lanc«olat«; 6 closely ap{«ozimat«d to 7; grey irro'
rated wilii darii tnscoiu and aometimea with whitish; a HuSnsed
ydlow-ochreoQS gradually attenuated streak along dorsum to near
^Mx; oblique dark fuscous ban from costa at \ and }, terminated
by himlfiah marks on fold ; a small cloudy whitish mark on middl«
of costa ; discal stigmata blackish, second elongate, edged beneath
by an elongate yelloW'Ochreoua spot ; a whitish spot on costa at },
formed of two subconflnent inwardly oblique strigulae, continued by
an oblique white line to posterior extremity of preceding ochreons
spot ; some white ma^pnal dots or scales round apex : cilia whitish.
grey, round apex white with two Aaxk grey lines, baae sprinkled
with blackish. Hindwings grey; cilia pale greyish.
COLOMBU, Caldas, 4400 feet, in May {Parish); eight
specimens.
Aristotella oribatis, n. sp.
$. li mm. Head whitish. Palpi white, second joint with base
and two bands brownish-ochteous, terminal joint with base and
three bands dark fuscous. Thorax ochrcous-whitish, shoulders dark
fuscous. Abdomen grey. Forewings elongate -lanceolate ; 6 sepa-
rate; blackish -fuscous; a suffused brownish-ochreoua streak along
dorsum from base to apex, occupying about J of wing, partially
eonflnent with a brownish-ochreous longitudinal streak above it from
before middle to }, which is partially edged with white above ; an
oblique white bar from costa at J almost reaching dorsal streak, and
a similar inwardly oblique white bar from costa at J ; a white apical
dot: cilia pale greyish, round apex with four dark grey lines, at
base sprinkled with blackish, above temus with on indistinct whitish
bar. Hindwings light violet-grey ; cilia pale grey.
Fbbu, Oroya, 12,200 feet, in July {Parish) ; one specimen.
iM-,i,A J h, Google
36 Mr. E. Meyrick's Descriptions of
Aristotellft vioaiu, n. sp.
3 $• 10-11 mm. Head grey-whitish. Palpi whitish, Becond
and terminal joinla each with base and two bands dark fuscous.
Thorax grey more or less sprinlded with whitish, shouldera dark
fuacona. Abdomen gnty, apex whitish. Forewings elongate*
laaceolal«; 6 closely approximated basally to 7 ; light grey irrorated
with dark fusoons ; oblique dark fueoons bara from ooata at } tuid }
t«rminated by black marks on fold, apace between these sufiused
with white; a fascia of irregular white suffusion crotaiiig wing in
middle ; discal stigmata black, second forming an elongate mark,
edged beneath by an elongate yellow-oohreous spot; an inwardly
obliqne triangular white spot on costa at }, and a mark on tomna
sometimes connected with it; some undefined ochreous marking
before temna and along termea ; two or three white marginal dote
loond apex : cilia pole grey, round apex white with a blackish
aatemedian line, on termen with ba«al groups of blackish irroration.
on coate dark fuscous, white on costal spot. Hindwings grey;
oUia pale grey.
Peru, Lima, in August {Parish) ; seventeen specimena.
ArlitotelU ephorit, n. sp.
(J. 9 mm. Head white, slightly sprinkled with grey. Palpi
whitish, second and terminal joints each with basal and supramediui
dark fascans bands, terminal joint shorter than second. Thoiuc
whitish iiTorated with giey. Abdomen light giey, anal tuft whitish.
Forewings elongate -lanceolate ; 6 separate; grey sprinkled with
whitish and dark fuscous ; a veiy oblique blackish mark on fold near
base ; an oblique dork fuscous bar from costa at \, terminated by a
blackish mark on told edged beneath with ochreous; discal stigmata
blackish, edged beneath with ochreous, fint beyond plioal; some
blaokish acales above tomus and at apex : cilia tight giey, round
apex sprinkled with blackisL Hindwings light bluish-grey; cilia
pole grey.
Peru, Matucana, 7780 feet, in July {Parish); one
specimen. Distinguished from all the preceding by the
short terminal joint and different marking of palpi.
Arlstotelia eosmographa, n. sp.
3 $, 11-12 mm. Head and thorax brownish-ocfareooa. Palpi
white, second joint with three and terminal joint with four dark
fuscous bonds, terminal joint much longer than second Abdomen
l„y,l,Ajh,COOl^lL'
Souih American Micro-Lepidoplera. 37
gre;, ftpez grey- whitish. Forewings narrowly elongate-UnoeoIate ;
6 aepuate; brownish -oohreons or deep yellow-ochreoua, costal
edge anffiued with dark fuscous; on oblique interrupted silvery-
white streak near base; three white streaks from casta terminated
by nlvery-metallio subdorsal spots, first from i, oblique, edged
anteriorly with blockish suffusion, second from middle, direct,
third from |, inwardly oblique, second and third connected by a
Boffused blaokieh streak in disc, a spot of silvery-white suffusion
beneath costa between first and second connected with costa by a
white strigula; a whit« dot on kpei, and somotimes two or three
on termoi : cilia pale greyish, with indistinct blackish median shade
and several slender wliitish bats. Hindwings slaty -grey; oilia pale
grey.
Peru, Chosica, 2800 feet, in July {Parieh); three
speciinens.
£Upau>sabi3, n. g.
Head smooth; ocelli absent; tongue developed. Antennae ],
in 3 simple, basal joint moderat«, without peoten. Labial palpi
very long, curved, ascending, second joint above in ^ clothed with
long fine expansible hairs, in $ with triangularly expoikded hair-
scales towards apex, beneath smooth-scaled, terminal joint some-
what shorter than second, slender, acute. Maxillary palpi rudimen'
taiy. Posterior tibiae clothed with long rough hairs above. ForC'
win^ with Ibfurcate, 2 from very near angle, 3 absent, 4fromangli
0 approximated, 7 and 8 out of 6, 7 to costs, 11 from middle. Hind'
wings }, elongate -trapezoidal, apex acute, termen obliquely bisinuate
beneath apex, cilia 3 ; 2 remote, 3 and 4 almost coni)at« from angle,
5 approximated, 6 obsolete.
Albed to Compsosaris, which it closely resembles
superficially. Vein 6 ol hindwmgs appears to be truly
oteolete (represented by a weak fold parallel to 7), and
not coincident with 7.
Hapalosaris pelnlaiu, n. sp.
3 9- 9-l\ mm. Head and thorax ochreous-whitish with a few
fuacons specks. Palpi whitish, second joint with three rings of dark
fuscous irroration, terminal joint with two black rings. Abdomen
ochreous-whitish. Forewings elongate-lanceolate; white, irregu-
lariy irrotated with fuscous and dark fuscous, more thinly towards
dorsum anteriorly; an oblique blackish bar from costa at | to
fold; blackish spots on costa before middle and at {; s black
iM-,i,A J h, Google
38 Mr. E. Meyrick'B Beacriptiona of
■nbcostal dot at }; stigmata black, sometimes ringed witb iritite.
pUoal rather obliquely before first discal, an additional dot betweoi
second disoal and dorstuu ; a small blackish spot on donmm befiH«
tomus; three blackish dots on costs towards apex, apical are*
darker-snSused : cilia pale grey, round apex sprinkled witb dark
fuscous. Hindwings light grey, in S thinly scaled and whitish-
tinged ant«riorly i cilia grey-whitish, in 3 whitish -ochreoos on lower
part of termen and dorsum ; in c? a long One pale ochreous-y^owi^
hairpencil from base of wing lying along abdomen.
Colombia, La Cmmbre, 6*500 feet, in May; Ecuador,
IIiiiRra, 4r)(H) feot, hi June ; Peru, Chosica, 2800 feet, in
July (Parish); twenty-six specimens.
Reourvarift xanthotrioha, n. sp.
ij ?. 8-0 mm. Head ochreoua-whitish, sides of crown with some
dark fuscous specks. Palpi whitish, second joint with basal halt
irrorated with dark fuscous and a blackish subapical ring, terminal
joint with two black rings. Thorax whitish irrorated with dark
fuscous. Abdomen grey, anal tuft of J pale ochreons. Forewings
elongate, narrow, cosia slightly arch<rd, apex obtuse- pointed, tennen
slightly sinuate, rathot strongly oblique; whitish, irregnlariy irro-
rated with grey and dark fuscous ; markings suffused, blackish, more
or less confused with the daib iiroration; a spot on base of costa;
a curved oblique scries of thi«e spots (costal, plioal, dorsal) towards
base, followed by a more or less developed narrow white fascia; a
subcostal dot at \, spots on costa at | and f ; stigmata represented
by cloudy dots, plical slightly beyond first discal ; a spot on tomns ;
sometimes an irregular angulated fine whitish transveree line is
traceable beyond this : cilia whitish-grey, sprinkled with daric fus-
cous or black except towards tomus. Hindwings and cilia pale
greyish; costa in £ rolled on anterior portion, with long ozpansible
yellow-ochreous hairpencil from base.
Peru, Chosica, 2800 feet, and Lima, in July and August
(Parish) ; twenty-four specimens.
Recurvaria thiodes, n. sp.
(J. 10 mm. Head pale yellow. Palpi whitish-yellow, second joint
with black snbapical ring, terminal joint with base and supramedian
band black. Antennae dark grey. Thorax pale yellow, with two
minute black marginal Aota posteriorly. Abdomen grey-whitish,
anal tnft whitish -ochreoos. Forewings elongate-lanceolate; pale
sulf^nr-yellow, riightly speckled with fnlvons; aa dongate Uaok
n,g,i,.rjh,G00glc
Sovtk American Mtcro-Lepidoptera. 39
■potklongbaMlaiztlial ooitk; amall bkok seioioval qiota OB ooaU
at } aad f ; {diottl and ■eoond duo«l Btigauta bUok; two or three
HD»11 blkck dots on ooMa towuds Kpex; four narrow aemioTal dsrk
gny nuika alwig termon, with & few bUek aokle* ; oilia grey, on
oo«t« pale jellow. Hindwinga light grey; cilu pale greTioh.
COLOMBU, La Crumbre, 6600 feet, in May (ParwA) ; one
RMnrvarU Mquorea, n. Bp.
(} $. 8 mm. Head, palpi, and thorax ochreom-whitish, shouldera
safiFneed with dark {dboohh. Abdomen in ^ grey-whitiah, in $ grey.
Forewinga lanoeoUto; dark fuaeoiu, sprinkled with whitish; a
cloud; whitish doraal streak from base to tonms, speckled with dark
fDecoos, pointed posteriori^, edge irregnlai : oilia ochreous-whitiah,
round apex and upper part of t«rmen irronit«d with dark fuscous.
Sndwings in i^ pale slaty -grey, in $ grey; oilia ocbieoos-whitiah ;
in 1^ with ochreons-yellawiah expansible hairpencil lying along ooeta
from base to beyond middle.
Pebd, Chosica, 2800 feet, and Lima, in July and Angust
{Parish) ; foui specimens.
OXYLECHIA, n. g.
Head smootb ; ocelli present ; tongue developed. Antennae {, in
1^ simple, baaal joint moderate^ without peoten. Labial palpi very
long, recurved, second joint with short rough tuft beneath and hatn
roughly expanded towards apex above, terminal joint longer than
aeoond, slender, acute. Maxillary palpi nidimentaiy. Posterior
tibiae clothed with hairs above. Forewings with lb furcate, 2 from
Dear angle, 3 absent, 7 and 8 etalked, 7 to costa, 11 from middle.
HindwingB j, elongate-trapezoidal, apex somewhat produced, acute,
termen obliquely bisiauate beneath apex, oilia 4; 2-4 remote, 4
frwn angle, 5 approximated to 4 at base, 6 and 7 connate.
Oxylechla eonOrmata, n. sp.
g. 8 mm. Head white. Patpi white, second joint marked with
Uock towards base, tenninal joint with block submediau ring and
snfnamediai^band. Thorax white, shoulders dark fuscous. Abdo-
men gi«y> segmental margins and anal tuft white. Forewings
elongate, very ikarrow, ooste slightly arched, apex pointed, termen
very obUqndy rounded; white, tinged with ochroous-yellowieh in
dvc, more atnmgly and sprinkled with fuscous posteriorly ; markings
dark fuacouB, with some black scales on their edges, and maigined
iM:,i,A J h, Google
40 Mr. E. Meyrick's Detcrijilions cf
with oleftr iriiite; & baeol patch, edge nmning from ^ of ootta^
atnight, obliqna ; an elongate narrow patch extending along ooata
from before middle to apex, antenoriy pointed and pieoeded by a
slight blackish strignla, beneath with slight proidinenoes at middle
and { of wing tipped with black scales probably repiesenting disoal
stigmata ; a small white costal mark intenecting this at ^ ; natiow
elongate brownish spots sprinkled with black on dorsom beyond
middle and on tamos ; a black apical dot ringed with white : cilia
whitish, roond apex with two dark fuacona linea. Hiadwings
whitish-greyj cilia ochreous-whitiBh.
Colombia. La Cmmbre, 6600 feet, in May (Parigk) ; one
Bpecimen.
Tholekostola, n. g.
Head smooth; ocelli absent ; tongne apparently obsolete. An-
temiae ), in ^ serrulate, simple, bssaJ joint elongate, without pecten.
Labial palpi long, recurred, second joint rather thickened and
roDghened with scales beneath, terminal joint hardly shorter than
second, slightly thickened with scales, acute. Maxillary palpi
rndimentary. Posterior tibiae clothed with long hairs above. Fore-
wings with 16 long-fnrcate, 2 from towards angle, 4 and 5 stalked,
7 and 8 stalked, 7 to costa, 11 from middle. Hindwings f, trape-
Eoidal, apex rather produced, acute, termen obtusely emarginate
beneath it, cilia 2 ; 2 from ) of celt, 3 approximated to 4 at base,
4 and 6 connate or closely approximated from angle, transverse
vein mostly obaolete except near lower extremity, 6 obeolete,
repreeentcd only by a weak fold parallel to 7.
Allied to Phihmmaea.
Ttaolerostola omphaloiM, n. sp.
3 $■ 6-7 mm. Head whitish, crown more or less speckled with
dark foBCOns. Palpi white, with two bands of second joint, and
baaal and median bands of terminal joint of blackish irrorsticm.
Thorax whitish irrorated with dark fuscous. Abdomen light grey,
anal toft of S grey-whitish. Forewiogs rather broad -lanceolate:
whitish irrorated with dark fuscous or blackish ; marf|^ngs formed
by denser blackish irrorstion or suffusion, undefined, vis. a rather
oblique bar from costa at J to fold, a broad rather oblique fascia
before middle, a broad fascia at j including an ochreous spot in
middle, and an apical blotch : cilia pale grey, round apex su&osed
with whitish and speckled with blackish. Hindwings grey; in ^
Ij.y.l.Ajl^yCOO'^lL'
South American Micro-Lepidoptera. 41
an expuiHible pencil of long light gnyiBh-ochieous luitB from
oostA neu base; cilia pale gre;.
Ecuador, Duran, low country, in June {Pariah); nine
specimens.
PhthorinuM triehinupls, n. ap.
S ?■ 9-10 mm. Head and thorax whitiBh-grey-ochreous, more or
lees sprinUed with fnscona. Palpi whitish, Becond joint irrorated
with dork fnacous except apex, terminal joint shorter than second,
with basal and supramedian bands of dark fusooiu or blackish irro-
ration. Abdomen whitish-ochieous, in 3 ^^h broad median dark
fiuoons band. Forewings elot^^te -lanceolate; fl sometimes out
of 7 near base; whitish-grey -ochieous, iiregularly iiTorat«d with
dark foacoos; stigmata dark fuscons, obscure, plical slii^tly before
first discal : cilia oobreous-grey-whitish, round apex irrorated with
dark fuscoos. Hindwinga and cilia ochreous-whitiBh ; in ^ beneath
with an elongate-oval patch of long black hairscoles below cell,
covered by a fringe of very long obliquely directed ochreoos hain
from lower median vein.
Pbbu, Lima, in August {Parish) ; five specimeDs.
Phthorlmae* tonte, n. ep.
(j$. 10-11 mm. Head and thoiax ochieous-whitish irrorated
with grey. Palpi whitish, second joint irrorated with dark fuscous
except apex, terminal joint almost as long as second, with basal and
supramedian bands of dark fuscous irroration. Abdomen grey-
Fore wings elongate -lanceolate ; 6 separate; pale grey iirorated with
darker. Obscurely and irregularly mixed with light ochreoos ; stig-
mata blackish, plical beneath first discal : cilia pale grey, somewhat
sprinkled with black. Hiudwings light bluish-grey; cilia pale
gnyish-ochreoofl ; in ^ with long ochreons-grey-whitisb expassible
haiipencil from base tying along costa.
Peru, Lima, in August (Parish) ; eight specimens.
Phthorimaeft perfldlosa, n. sp.
(J. 10 mm. Head and thorax whitish irrorated with grey. Palpi
whitish, second joint irrorat«d with dark fuscous except apex, ter-
minal joint somewhat shorter than second, with basal and supra-
median bands of dark fuscous irroration- Abdomen dark grey, anal
tuft ochreous-whitish, claapeis very long, clothed with ochieous
whitisb hairs suffused with yellow internally. Forewings elongate,
narrow, coeta gently arobed, apex pointed, tennen very obliquely
iM-,i,A J h, Google
42 Hr. E. Meyrick's Dettrijitions of
ronodMl; 6 Mpttfate; grey-irliitiBh sprinkled with gny and dark
fnsoons, more etrongly iirontod akmg oosU; plical atignu nthar
large, bUckiah : cilia p«le grejiib, aprinUed with blBckiah towaids
baae. Hindwinga bluiah-grej ; a strong oohreoaa-j'ellowiali ezpan-
aible bairpenoil from base lying along coeta to beyond middle ; cilia
pale ochieoufl-grey.
Colombia, Caldas, 4400 feet, and La Crambre, 6600 feet,
in May (Parish) ; two specimens.
nithorimus ernslarls, n. sp.
<}$. 11-12 mm. Head and tboi«x ochreons-wfaiti^ Bometimea
sprinkled with giey. Palpi white, aeoond and tenninal joimU ettcb
with baaal and nipramedion bands of blackish inoiation, terminal
joint somewhat shorter than second. Abdomen ochreons-whitid),
sometimes doraally greyish. Forewings elongate, narrow, ooeta
gently arched, apes pointed, termoi eitremdy obliquely roonded;
S separate; oohreoos- whitish, iiregnlarly and variably sprinkled
with grey, ochreous, and blackish ; small cloudy spots of blaokiri)
irroration on coeta at base and ^, aod a clovdy blaokiah dot on fold
near basei atigmata rather large, black, sometimea tinged with
ochreoos, plical rather obliquely before first diacali sometimes
indications of cloudy dots of blackish irroration on coeta posteriorly
and termen : cilia ochreons-wbitish, on basal half wit^ scattered
dark giey and blackish points. Hindwinga light alaty-grey; cilia
ochreons-wbitish, sometimea tinged with grey.
Peru, lima, in August [Parish] ; fourteen specimeos.
PhOioritiUM densata, n. sp.
,J$. 12-Umin. Head and thorax grey, sometimea sprinkled witli
whitish. Palpi white, second and terminal jointa each with basal
and eupramedian bands of blackish irroration, tenninal joint aome-
wbat shorter than second. Abdomen grey, basal segment aometimea
with two small fulvous spots, segments 2 and 3 with fulvous doraal
patches. Forewings elongate -lanceolate ; 6 separate ; whitish ino-
ntted with dark grey or blackish; several indistinct blackish dots
on baaal area, an obscure rather obli<]ne darker streak from coeta
at i to plical stigma, sometimes edged with whitish antwioriy, pre-
ceded and followed beneath coeta by oobreous marks ; stiginato
moderate or large, black, edged below by ochreooa spots aod some-
limee sonounded by irregular ochreous markings, plical rather
obliquely before first discal; indistinct cloudy whitish opposite cost»l
and tomal marks at j, sometimea united into a aUgbtly angaUt«d
Ij.y.l.Ajl^yCOO'^lL'
South American MicTO-Lepidoplera. 43
shade : cilia light giej, basal half mora or lees sprinkled with whit«
and black. Hindwings slaty-grey ; cilia grey.
Pebu, Lima, in August (Parish); twenty -Beven speci-
PhtttorlmMa gngalls, n. sp.
<}$. 12-13 mm. K^ad and tfaocax dark gcey, more or less
■prinkled witii whitish, face sometimes sofFused with whitish. Palpi
whitish, second uid terminal joints each with basal and sapra-
niediaQ bands of blackish irroration, terminal joint somewhat shorter
than second. Abdomen dark gray, apes tinged with ochieoos.
Forewinp eloi^ate-lanceolate; fl separate; dark fuscous, speckled
with whitish; sometimes short brownish.oohraoQS dashes beneath
oosta at \ and \ ; stigmata small, black, sometimes edged with brown-
ish markings, plical rather obliquely beforo first discal; sometimes
some obscure brownish marking in disc pOBteriorly : cilia light grey,
basal half sprinkled with black. Hindwings slaty-grey \ cilia light
ochreous-grey.
Peru, Lima, in August {Parish) ; five apecimens.
Pbthorlmaea utosenut, n. sp.
1^. 14 mm. Head ochreous -whitish, ciown irrorated with grey.
Palpi whitish, second joint with two suffused [uacous bands, tenninal
joint with two dark fuscous hands. Thorax pale greyish -ochreous,
mostly sufiusedly iirorated with dark grey. Abdomen grey, anal
toft ochraoDB-whitiah with a strong black mark on each side above.
Forewings elongate-lanceolate ; Bseparate; gray ish-ochreoUB, some .
what sprinkled irregularly with dark grey, costa suffused with dark
grey irroration; three or tour cloudy blackish-grey dots towards
base on dorsal half; an oblique transfeTse patch of ochreous .whitish
auEFoaion from costa towards base followed on costa by a small
blackish spot and in disc by an elongate blotch of blackish suffusion ;
stigmata blackish, surrounded by irregular ochreous- whitish suffu-
sion, plic^ obliquely before fint discal. a blotch of blackish suffusion
in middle of disc lying between and beneath discal stigmata; costa
towards apex with three small ochreous- whitish spots interrupting
the dark grey irroration : cilia ochreous.grey- whitish, towards base
spotted with dark grey irroration. Hindwings slaty-grey; cilia
pale gtejiBh.ochreaiiB.
Peed, Matucana, 7780 feet, in July {Parish); one
specimen.
iM-,i,A J h, Google
44 Mr. B. Me3nick'B DeacrijilioM <^
PhtiiorlnuM melanoeunpta, n. sp.
(J. 16 tnm. Head and tbonuc pile grej'iah-ochieoiu Boffnaed with
grey. Palpi whJtiah, second joint Bprinkled with dork fuscoos,
twniiiul joint with two blackish barab. Abdomen blsokiab becom-
ing brawn towards base, aides and anal toft ochreotis-whitish. Foie-
wings elongate-lanceolate ; 6 separat« ; light grejisb-ochreoiu,
irregularly sprinkled with blackish-grey, ooata narrowly snSused with
dark grey irroration; several cloudy black dot« on basal area; a
thick black sufiused streak from coata at } rather obliquely half
aoroes wing, thence abruptly bent and continued through middle
of disc to apex, attenuated posteriorly, neaiiy interrupted by small
pole spota repreeenting discal stigmata, and irregolarly intemptod
near apex : cilia light greyish-ochreous, sprinkled with black towards
base. Hindwings pale slaty-grey; oilia light gieyieh-ochreons.
Peru, Lima, in August {Parish); one specimen.
PhthorinuM kquUlna, n. sp.
S $. 19-22 mm. Head and thorax varying from pale oohreous
to grey. Palpi oohreous- whitish, second joint sprinkled with fuscons.
terminal joint with basal and supramedian rings of daxk fuscous
irroration. Abdomen ochreous- whitish, segments tinged with fuscous
towards base, two basal segments more or less sufFused with yellow-
ochreous. Forewings narrowly elongate-lanceolate ; 6 separate ; vary-
ing from pale ochreous, sprinkled with ferruginous ($) to light fuscous
(ij); A thick black (or in 9 deep ferruginoUB) ratherobliquestieak from
costa at } reaching half across wing, in disc dilated posteriorly, form-
ing base of an undefined cloudy wedge-shaped patch rather darker
than ground colour extending along coeta to } ; indistinct cloudy spota
ol darker suEFusion in disc at middle and j ; seTcral small obscure
cloudy darker spota on costa and termen towards apex : cilia pale
gieyish-ochreous or whitish- ochreous, sprinkled with fuscous ot dark
fuscous. Hindwings pale slaty-grey ; cilia pale greyish-ochreous or
whitish-ochreous.
Peru, Matucana, 7780 feet, and Huancayo, 10,650 feet,
ill July (Parish) ; four specimens.
Phthorimaea absoluta, n. sp.
cJ- 14 mm. Head whitish -grey -ochreous, crown sprinkled with
grey. Palpi stout, ochreous- whitish sprinkled with grey, second
and tenninal joints each with two bands of blackish irroration, second
joint with longer rough scales beneath towards apex. Thorax
l„y,|,AJI^,COOl^lC
South American Micro-Lepidoptera. 46
p«yiih-ochnoiu iirorated with whitish and dark grey. Abdomen
grey, segmental margins and anal tuft oohreooB-whitiah. Fore-
wing) nanowl; elongate-lanceolate; 6 aepante; greyiBh-ochreous
s^inkled with dark grey and whitisb, here and there mixed with
brownish, oosta snSnaedly iirorated with blackish-grey, Teina
poateriorlj otwcarely inorated with dark grey; Bevetal obsonre
blackish dots on basal area ; irregular rather obliqne transverse bars
of blackish snSusion from ooeta at { and |, reachinghaU aoroas wing;
stigmata blackish, plioal obliquely before first disoal ; costal irnira-
tion towards apes interrapted by three or fonr small pale spots :
cilia light ochreons-greyigb, on basal half mixed with ochreoua-
whitish and sprinkled with fuscous and black. Hindwings light
slaty-grey; cilia light ochieous-grey.
Peru, Hiiancayo, 10,650 feet, is July {Parish); one
specimen. Much like urosema, but readily distinguished
by absence of the characteristic black marka on anal tuft.
PhlborlnuM loquax, n. sp.
j$. ^10 nun. Head and thorax grey, sometimes partially
tinged with oohreone, face sometimes whitish. Palpi fuscous
aprinkled with whitish, second and terminal joints each with baaal and
snpramedian bands of blackish irroiation, terminal joint somewhat
shorter than second. Abdomen in ^ dark grey, in $ whitish-giey.
Forewinga elongate-lanceolate ; 6 separate; dark grey sprinkled with
whitish, tinged here and there with oclireous; a black dot beneath
COsta near base, and two or ttiree Other indistinct ones on bssal area ;
» small black costal spot before } ; sometimes ochreons subcostal
dasbee before and beyond this ; stigmata moderate or large, ochreons-
brownish, sometimes accompanied by a few blackish scales, plical
aligbtly before first discal ; a blackish dot on fold beneath middle of
wing ; a small cloudy darker spot on costa at f ; a clondy darker dot
above tomns; a cloudy spot of dork fuscous suffusion on termen
above tomns ; an elongate blackish mark in disc near apex : cilia
light grey, basal half sprinkled with whitish and dark grey- Hind-
wings slaty-grey ; cilia pale ochreous-greyish, in ^ longer and darker
towards basal portion of costa.
Peru, Chosica, 2800 feet, in July {Parish); thirteen
Bpecimens-
Phthorlmaea alritascls, n. sp.
3 $■ 9-10 mm. Head ochreons- whitish irrorated with grey.
Palpi grey, sprinkled with whitish, second and terminal joints each
witii basal and supramedian bands of blackish irroration, terminal
u,y,l,/eJh,G00glc
46 Mr. E, Meyrick'a Descriptions e^
joint nearly as long aa Becond. Thorax grey ipiinkW with ndiltiah.
Abdomen whitiah-grej. Poaterior tibiae in 3 ^^ '^'T '""S
whitish -oohreouB hairs. Forewinge eloDgate-lanoeoUte ; 6 aepante ;
dark grey inorated with white ; a blackish dot beneath costa near
base ; an oblique blackish bar from costa at } to lold ; discal stigmata
blackish, indistinctly edged with ochreons beneath, plioal ochreous,
■lightly before first discal; indistinct whitish opposite ma^s on
ooeta at i and tomua : cilia whitish-grey, irrorated with dark giey
and black round apex and towards base generally. Hindwings pole
slaty-grey ; in d a very long dense black cxpansibls hairpencil lying
along oosta fiom base to ] ; cilia wbitisb-grey.
Peru, Ghoaica, 2800 feet, in July iParish) ; two specimens.
PbthorinuM altlsona, n. sp.
1^. 11-12 mm. Head and thorax dark fuscous, slightly whitish-
sprinkled. Palpi dark grey sprinkled with blackish and slightly
with whitish. Abdomen dark grey. Forewings elongate-lanceolate ;
6 separate; dark fuscous, slightly whitish -sprinkled, doisal area
tinged with oohreoos -brown ; stigmata cloudy, black, plioal obliquely
before first discal ; in one specimen some oohreous-brown streaking
towards coflta posteriorly ; apical area suffused with blackish :
cilia grey, round apex mixed with dark fusoous and somewhat
epriokled with whitish. Hindwings bluish-giey ; cilia light grey-
Peru, Huancayo, 10,650 feet, in July (Parisk); two
specimens.
FbthorimMa eueta, u. sp.
S $. 11-12 mm. Head whitish, sprinkled with dark grey. Palpi
white sprinkled with grey, second joint blackish except towards
apex, terminal joint nearly as long as second, with black basal ring
and supramedian band. Thorax whitish irrorated with dark grey.
Abdomen grey, anal tuft in S ochreous- whitish. Forewings elongate,
narrow, costa gently arched, apex pointed, termen extremely
ofaliqnety rounded ; 6 separate ; whitish irrorated with grey and dark
fuscous ; a small blackish spot beneath costa near base ; a suffused
dark fuscous bar from dorsum at } to above middle ; blackish spots
on costa at } and before J, stigmata cloudy, blaek, plical beneath
fiist discal, second discal elongat«; blotches of darker suffusion on
lomns and at apex : cilia pale grey mixed with dark fuscous.
Hindwings grey, darker posteriorly; cilia light grey.
British Guuna, Bftrtica, in January and February
{Pariah); two specimens.
l„y,l,Ajh,COOl^lC
South American Miorb-Lepidoplera. 47
ntthoiiioMa epitrieha, n. sp.
1^ $. 8-10 mm. Head whitish, more or leea eprinkled with grey.
Palpi whitish, second joint irrorated with dark fusooos except apex,
terminal joint as long as second, with subbasal and Bn|»amedian
bands of dark foscons iiroration. Thorax whitish more or less ino-
rat«d with dark grey. Abdomen grey, in 3 with raised lateral snb-
apical tofts of long grey haira. Forewinge elongate, rather narrow,
ooBta gently arched, apex pointed, tennen veiy obliquely rounded ;
6 separate; grey-whitiah inoiated with dark fuscoua; a blackish
spot or short obliqne bar from coeta near base ; a amall ochreous spot
on fold at J, sometimes edged above and beneath with small blackiBh
q)Ots; a t^ck obllqoe blackish streak from | of coeta to fold;
stigmata ochreoos, more oi lees edged above and beneath with sm^
blackish spots, plica! obliquely before fiist discal, an elongate cloudy
UachJsb spot on costa just above second discal, sometimes edged
boieath by another ochreous dot; undefined spots of blackish
snffnHion <m tomns and at apex : cilia pale grey sprinkled with
black. Hindwings grey, snbhyaline in disc anterioriy and towards
dorsum ; cilia light grey.
BsTTiSH Gdiana, Bartica, fiom December to February
(Parish); ten Bpecimena.
PhthoiimMa InTOlate, n. ap.
ij$. 9-1 1 mm. Head whitish sprinkled with giey. Falpi
whitish, seoond and terminal joints each with subbasal and supra-
median bands of dark fuscous irroration, terminal joint as long as
seoond. Thorax whitiah irrorated with dark fuscous, patagia
sometimes mSused with oclireoiu. Abdomen dark grey. Pore-
wings elongate, narniw, coBta gently arched, apex jwinted, termen
very obliquely rounded ; 8 separate ; grey- whitish or whitish-grey,
iiroi«ted with black; a narrow oblique blackish bar from costa
Dear base and broader one at ), both terminated by small yellow-
ochreouB spots on fold edged beneath by black marks, second spot
representing plical stigma; discal stigmata yellow-ochreous, edged
above and below by black spots, Rrst obliquely beyond plical stigma,
its margin separated by an ochrcous mark from a small blackish spot
on coeta above it, maigin of second usually absorbed in a suhquad-
rate blackish blotch on costa above it, its lower mai^ sometimes
forming a oonaidetable spot ; a well-defined blackish apical blotch :
cilia grey, inorated with black except towards tips. Hindwings
daric grey, subhyaline in disc anteriorly and towsids dorsum ; cilia
Mther dark grey.
iM:,i,A J 1^, Google
48 Mr. E. Meyrick'a Descrijitionb of
British Guiana, Mallali, in March {Parish) ; six speci-
mens. Nearly allied to epUricha, but forewings rather
narrower, yellowish markings clearer and larger, black
markings also larger and better defined, hindwings darker,
abdomen in S without praeapical tults of hair.
Stegasta zygotoiu, n. sp.
<;$. ^11 mm. Head wlute, often irrorated with dark grey.
Folpi white, second joint with ill-defined basal and subapioal buidt
of dark fuscous irroration, terminal joints with two blackish bands.
Thorax dark fuscous, slightly whitish-sprinkled, with small ochreous-
white posterior spot. Abdomen grey. Forewings elongate, narrow,
oosta gently arched, apex pointed, termen very obliquely rounded;
dark fuecons, sometimes sufiused with deep ferruginous- bronze,
slightly whitish-sprinkled ; two connected ochteons-white triangular
blotches occupying doranm from near base to near tomus, sometimes
much sufiused with grey irroration, especially dorsally, first narrowly
reaching costa at \, second reaching neariy half across wing, its apex
oonnected with an indistinct whitish dot on middle of costa hy a
ftunt grey or grey-whitish cloud in which is a more or less strongly
expreoeed blackish dot (first discal stigma); second discal stigma
close beyond this, blackish, sometimes edged below with white or
yellowish, or obsolete ; a slightly inwards-oblique transveise white
spot on coeta at \, landing to connect with a leaden-grey spot on
tomus; apical area sometimes irrorated with grey-whitish : cilia
whitish, with two indistinct lines of blackish irroration. Hindwinga
giey or light grey, darker posteriorly; cilia grey or grey-whitish.
Forewings in $ beneath with an expansible pencil of long whitish
hairs from beneath base of costa, covered by fringe of projecting
scales.
Colombia, Cali, 600 feet, and Ia Crumbre, 6600 feet, in
May ; Ecuadoe, Huigra, 4500 feet, in June ; Peru, Lima,
and Chosica, 2800 feet, in August {Parish); forty-four
specimens. The Peruvian examples have the white mark-
ingB always much mote suffused with grey, whilst in the
others they are nearly clear ochreous-white, but I can find
no rehable distinction between them otherwbe.
GeleeUa lactlconu, n. sp.
fj. 10-11 mm. Head and thorax ochreous-wbitish, shoulders
with a suffused dark fuscous spot. Palpi whitish, basal third of
second joint dork fuacons. terminal joint as long as second,
with blackish subapical band. Abdomen grey, anal tuft ochreous-
ih,Cooi^lc
South American Micro-Lepidoptera. 49
whitish. Forevinga elongate, doitow, costa gently arched, apex
tolerably pointed, termen very obliquely rounded; ochreous-
whitish or yeUoW' whitish; marking fuscous, towards costa iirorated
with blackish ; aa obUqoe wedge-shaped spot from base of casta to
iokl ; a V-shaped marking from costa before middle, iU apex reaeb-
iog fold, marked with black plical and first discal stigmata, latter
obliquely posterior; an irregular fascia from } of costa to dorsum
before tonins, second dieoal stigma fomuDg a black mark on its
anterior ma^in; some more or less indicated blackish irroration
towards apex : cilia ocbreous-whitieh, slightly sprinkled with blaok.
Hindwings light grey; cilia ochreous -whitish.
Pebd, Chosica, 2800 feet, in July (Parish) ; three speci-
mens.
Gelechla litlglosa, n. sp.
ij$. ll-13mm. Head and thorax pale pinkish-fuscous or whitish-
fuscous, mixed with dark fnscous. Palpi fuscous- whitish irroratod
with dark fuscous, on terminal joint sometimes forming two dork
bonds. Abdomen grey. Forewings elongate, narrow, costa gently
arched, apex obtuse -pointed, termen very obliquely rounded;
dark fuscous, variably iirorated (sometimes very slightly) with
whitish -fuacoua or whitish-ochreous ; stigmata doudy, blackish,
plical slightly before first diecal; a small cloudy pale ochreous or
whitjsh-ochreous spot on costa at | : cilia grey, somewhat mixed
with dark fuscous. Hindwings and cilia grey.
Ecuador, Huigra, 4500 feet, in June {Parish) ; thirteen
specimens.
Gflleohla dryobathra, n. sp.
<;. 13-14 mm. Head fuscous, face paler. Palpi pale greyish-
ochreous .irrorated with dark fuscous, terminal joint somewhat
■hcvter than second. Thorax brown or fuscous. Abdomen grey.
Forewings elongate, narrow, costa gently arched, faintly unuate in
middle, apex obtuse, termen very obliquely rounded; dark fuscous,
in one specimen streaked with brownish in disc posterioriy; a
brown baaal patoh occupying about ^ of wing, edge irregularly
curved or bent; stigmata blackish, approximated, plical somewhat
obUqoely before firat discal ; a small pale brownish spot on costa at
} : cilia pale greyish-ochreoue, with two lines of blackish innration,
at base fuscous, on costa wholly fuscous. Hindwings grey; cilia
light greyish-ochreoua.
Colombia, La Crunibre, 6600 feet, in May {Parish) ; two
specimens.
TBANS. EST. 80C. LOND. 1917. — PART I. (NOV.) E
u,y,i,Ajh,Cooglc
60 Mr, E, MeTTick'e Description of
Gehetala eborata, n. sp.
[}$. 14-15 mm. Head oohreous-whito. Palpi whit«, baoe of
■eoond joint black, terminal joint aa long aa second, apical f black
except ertreme tip. Thorax dark aBhy-fuaoouB, with broad oohreous-
white doraal stripe. Abdomen dark grey; ForewingB elongate,
rather narrow, costa gently arched, apes obtuse-pointed, termen
rounded, rather strongly oblique ; dark ashy-fuscons ; stigmata hiaok,
obecm:^, plical rather obliquely before first diacal ; a small cloudy
white dot on costa at } ; sometimes one or two white scales on npper
part of t«rmen : cilia dark grey. Hindwings dark grey ; cilia grey.
CoLOMBU, La Crumbre, 6600 feet, in May {Parish) ; five
specimenB.
Geleohla consona, d. sp.
$. 13 mm. Head ochreous-white. Palpi white, base of second
joint blackish, terminal joint nearly as long as second, anterior
edge blackish on apical half. Thorax ochreous-white, patagia darie
purplish -fuscous. Abdomen grey. Fore wings elongate, narrow,
costa gently arched, apex pointed, termen very obliquely rounded ;
dark purplish-fuscous ; a thick white streak along donom from base
to J, irregularly terminated and uniting with a roundish white spot
in disc beyond middle of wing; a roundish white spot on coeta at
}: cilia grey mixed with blackish. Hindwings grey; cilia light
Ferd, Lima, in August (Parish) ; one specimen. Allied
to the North American unifasdeUa.
Gelechia argosema, n. sp.
(J$. 9-11 mm. Head ochreous-white- Palpi ochreous-white,
basal third of second joint blackish, terminal joint as long as second,
apical I blackish except extreme tip- Thorax ochreous-white,
patagia dark purplish-fuacous. Abdomen grey- Forewings elon-
gate, rather narrow, costa gently arched, apex obtuse-pointed,
termen very obliquely rounded; dark purplish .fuscous; a sub-
triangular transverse ochreous-white spot from doisum before
tomns, reaching half across wing, and a smaller spot from costa at
I ; cilia grey suffused with dark purple-fuscons. Hindwings grey;
cilia light grey.
Ecuador, Huigia, 4500 feet, in June (Parish); five
South American Micro-Lepidoplera. 51
GeheUa pentadon, n. sp.
,J$. 12-14 mm. Head ochreoua-white. Palpi yellow- whitUh,
second joint with baeedark foacous, terminal joint as long aasMond,
with gobapical band and aometimea base blackiab. Thorai: dark
purple-foscona, witJi yellow posterior spot. Abdomen dark grey.
Forewings elongate, narrow, coata gently arched, apex tolerably
pointed, tennen very obliquely rounded; dark pnrple-fuacona;
markingB yellow; a alightly oblique oval transverse blot^sh from
oosta at } reaching { acroas wing; a spot beneath middle of disc;
an irr^ular inwardly oblique transverse blotch bom coata at f,
reaching more than half across wing; a spot on tomus; a small
apot before apex : cilia dark purple-fuscous. Hindwings and cilia
dark grey.
Fkench GciANA, R. Maroni ; three Bpecimens.
Polyhynmo gladlata, □. sp.
^. 9 nun. Head, palpi, and thorax white (partially defaced),
■boulders dark fuscous. Abdomen grey, anal tuft ochreous- whitish.
Forewings elongate, rather narrow, costa alightly arched, apex
pointed, prominent, termen ainnate, oblique: costal half dark fus-
cous, dorsal area ailvery-grey-whitish, these separated by a broad
median longitudinal shining white stfeak from liase to g, pointed
pCNterioriy, lower edge suffused; a very fine white line along costs
from near base to middle, thence diverted aa a alender very oblique
atreak to just above apex of median atreak; an oblique white
strigula from costa about {, whence a rather thick angulated leaden-
metallic line tuns to tomus ; beyond this a small white coatat strigula,
followed by a small white triangular spot; apical and terminal
areas round these markings biownish-ochieous : cilia pale grey,
above apex with two white wedge-shaped marks separated with dark
foscoos, on termen white towards base, with a very abort black
■nbbasol mack beneath apex and two black subbaaal dote below
middle. Hindwings and cilia light grey.
Colombia, Cali, 500 feet, in May (Parish) ; one apecimen.
Polyhymno confllota, n. sp.
<J $. 10-13 mm. Head whitish. Palpi whitish, second joint
fnacotui except apex, scales roughly projecting beneath towards
apex, terminal joint fuscous anteriorly. Thorax fuscous-whitish.
Abdomen whitish-fuscous, anal tuft fuscous-whitish. Forewings
ehmgate, narrow, costa slightly arched, apex produced, acute.
u,y,l,/eJh,G00glc
52 Mr. E. Meyrick's Deacriplvm <^ Miero-Lejndoptera.
termen uniiat«, oblique ; dark fuscous, with cloudy white or whitish
markingi, vie a BubcoetAl line on haMal half, a aupramediaD line to
g, where it meets ui attenuated stre&k from beneath middle of disc.
Mid a confused and ittegular more or less confluent suSu^ion occupj-
ing most of donsl area up to fold; a very oblique white line from
middle of costa to near termen above middle; a white oblique
strigula from coBta at j, whence a fine acutely angulated leaden-
roetalUo line runs to tomos; two white wedge-Bhaped marks from
costa beyond this ; apicalarea tinged with ochreous: cilia lightgrey,
above apex with two white triangular marks separated with dark
fuscous, at apex with a projecting dark fuscous book, beneath apex
white towards base, with a very short black aubbasal mark beneath
apex and dot below middle. Hindwings light slaty-grey ; cilia
pale greyish.
Peru, Choaica, 2800 feet, and lima, in July and August
{Parish); twenty-seven apecimens.
n,g,t,7.dh,G00glc
II. iVeic Specim of Hvmenoptera i» the British Musfom.
By Rowland E. Tueseb, F.Z.S., FES.
Family BETHYLIDAE.
G«nuB DiCBOGENiuv, Stadelm.
DicTogenium, Stadelm., Entom. Nachticht, zz. p. 201,
1894.
Nomineia, Kieff., Ann. Soc. Ent. France- lux, p. 453,
1911.
I consider that these are identical, and that Ashmead's
action in tiansfemng the genus from the Bdkylidae to
his Cosilidae was entirely unjustifiable.
IMen^idwii maxbnaiB, sp. n.
J. Niger; mandibolis apioe fusco-femgineis: alia fnacis, ^lioe
fnsco-hyalinis.
Long. IS mm.
Q. Maodiblee strongly bidentate at the apex, the outer tooth ttte
longest; clypeoa abort, transverse, with a strong median carina.
Flagelhun ptmctnred, hair;, becoming slender towards the apex,
scape smooth and shining, the antennae as long aa the thorac
and median segment combined. Head very large, broader than
the thorax, broadly emarginale posteriorly, closely and coarsely
pUDctured- reticulate; cjee very near to the liase of the mandibles,
separated from the posterior margin of the head by a distance
exceeding their own breadth; the cheeks broader than the eyes,
spanely but dce[dy punctured, bearing a stout, acute, spine. Prono-
tam short, strongly narrowed antcrioriy, the anterior maigin raised,
a deep groove along the posterior margin. Mesonotum coarsely
pnnctored, more sparsely on the middle than on the aides, the
parapetdal furrows shallow; scutellum very sparsely punctured;
mcAopleonie cloaely and coarsely punctured, with a shining, almost
Binooth, area behind the wings. Median segment irregularly
rvgoloee, with two longitudinal csrinae close together, the narrow
space between the carinae transveraelj striated, the sidea of the
■eminent strongly horizontally striated, the apical slope moro finely
TBASS. EKT. SOC. LOND. 1917. — PAST I. (NOV.)
u,y,l,/eJh,G00glc
54 Mr. R. E. Turner on New Spedes oj
tnmevenely striated. Abdomen shining, almoat smooth; the
seventh donal segment Hpaisely and ahallowljr punctuied, brosdly
truncate at the apex; apical ventral segment closely punotuTsd,
hairy, deeply divided longitudinally in the middle. Radial cell
open at the apex, but the radius is indicated by a blown continua-
tion almost to the apex of the wing ; cubitus indicated by a white
scar beyond the fitst transveise cubital nervure, the latter indicated
by a brown scar, the second transreise cubital by a white scar.
Hah. Sierra Leone, TBiuadu (■/. 3. Simpson), June.
There is no tooth on the inner side of the mandibles as
there is in D. conradti, Stadelm., and the carina of the
proatemum is not furcate as in D. rosmarus, Stadelm.
Family MUTILLIDAE.
Ephutomorpha eoDtlgoa, sp. n.
$■ Nigra; mandibulis baai, tubercnliaque antennalibns fusco-
femigineis; thorace rufo-femiginoo ; flagello subtus, tarsisque
brunneo-terrugineis; segraento doraali primo apioe late luteo;
segmento dorsali secundo obscure aeneo-vireecente.
Long. 1 mm. '
$. Antennal tubercles well developed, rounded; second joist
of the flagellum distinctly longer than the third. Eyes oval, with
distinct facets, situated nearer to the posterior margin of the head
than to the base of the mandibles. Head nearly as broad as the
thorax, feebly rounded at the posterior angles, closely and not very
finely punctured. Thorax closely punctured rugose on the anterior
two-thirds, the apical third coarsely rugose, the aides of the apical
slope feebly seirate ; the thorax broadest anteriorly, the anterior
angles feebly rounded, the anterior maigin almost transverse, the
length of the thorax greater by about one quarter than the greatest
breadth, abruptly hut not strongly narrowed behind the middle,
a small tuft of erect white hairs on the mesopleurae. Second dorsal
segment a little longer than the greatest breadth, the sides rather
strongly convex, closely punctured rugulose; with an apical pateh
of white hairs, which is continued as a median band on the third,
fourth and fifth segments. Pygidial area triangular, finely longi-
tudinally striated.
Hob. S. Qdeeksland, Brisbane {Hacker), December.
Nearest to E. mrulerUa, Turn., from S.W. Australia,
but the thorax U less strongly narrowed posteriorly than
in that species, the colour of the thorax is much brighter.
ih,Cooi^lc
Hymenoptera in the British Museum. 55
and the spots of pubescence on the dorsal segments
are different, virvlenla having an apical fascia of white
pnbesoence instead of an apical spot.
Ephntomorpha Inrida, sp. n.
$. Pallide terruginBa; aegmsnto doraali woimdo fascia in«diana
l<»tgittidiiiAli nigift, Mgmentis qninto aextoqne fuscis; eegmento
donali aectmdo macula magna ntrinque fawiaqne angnata apioali,
t«rtioqiie macula medianaalbopilons; area pjgiduJi lat« triangaUri,
delkatisaime punctata.
Long. 5-6 mm.
$. Head and thorax sparaely clothed with pole fulvompobeeoence;
antonnal tuberclee fairiy welt developed, leoond joint of the flagel-
hun very little longer than the thiid Eyea round, Btiongly oonvex,
roach nearer to the poateriot maxpa of the head than to the base
of the mandibles. Head narrower than the thorn, strongly rounded
at the posterior an^ce, finely and closely punctured. Thorax fully
half B8 long again as the greatest breadth, strongly narrowed poB-
terioily, the apioal margin scarcely more than half as wide as the
baaal; the aat«rior margin feebly arched, not quite tranaverae,
rugose, more coanely at the apes than at the base, the posterior
slope coaisely rugose. Second dorsal segment finely longitudinally
tnguloBe ; fully half as long again as the greatest breadth, the sides
very feebly convoi. Sides of the abdomen thinly clothed with
Itrog fulTOOS hairs 1 pygidial area broadly triangular, very minutely
pnnotored. Hind tibiae with one row of four spinea.
Hob. N. Queensland, Townsville (F. P. Dodd), Sep-
tember.
This belongs to the gronp of E. dartoiniana, Andi^, and
E. eccerpla, Tom., but differs from both in colour; from
the formei in the more strongly narrowed apical portion
of the thorax, the longer second abdominal segment and
the sculpture of the pygidial area; from the latter in the
more graduaUy narrowed thorax and in the less convex
sides of the second abdominal segment.
Ephatomorpha dlrarslpes, sp. n.
$. Nigra; vertice lat«, front« lugro Intaminato, thoraceque
fermgineis ; tubeiculis antennalibus, flagello articulo primo, taraiaque
pallide testaceis; segmeoto doraali primo fascia apicali pallide
hitea; calcaribua pollidis: segmentis dorsaibus 2-fi fascia mediana
longitndinali oontinua albido pilosa.
j/»)tf 5 mm.
n,g,t,7.dh,G00glc
56 Bfr. B. E. Turner <m Sew Species <^
$. Antennal tnbercleB wdl devdoped, roonded ; Moond joint of
the flagellnin distinctlj longer than the ttiiid. Head a little
narrower than the thorax, rounded at the posterior angle* ; finrij
and closely punctured on the vertex, more deeply but not so cloaelj
on the front. Eyes nearer to the posterior margin of the head tluD
to the base of the mandiblea, round and etrraigly convex. Thorax
a little le«s than half as long again as the greatest breadth, broadly
rounded on the anterior margin, slightly narrowed poateriorly.
longitudinally rugose striate on the basal two-thirds, coanely mgoae
on the apical third; the posterior dope almost vertioa], coars^
rugose, with two minute lateral teeth close to the basal angjes.
Second donal s^ment findy longitudln^y rugose, about one-thiid
longer than tJie greatest breadth, the sides moderatdy convex.
Pygidial area very nanowly rounded at the apei, findy iMigi-
tudiually striate on the basal half. Hind tibiae with two rows of
four rather feeble spines.
Hab. N. Queensland, Tow-nsville {F. P. Dodd), Novem-
ber and June.
Near E. vittigera, Aiidre. a variety of which occurs in
October in the same locality. It differs in the sculpture
of the thorax, which does not run into striae in rilligera,
in the absence of fulvous hairs on the thorax, in the ferru-
ginous colour of the head and thorax, in the feebler spines
of the hind tibiae, and in the colour of the tarsi ; it is also
a smaller species.
Ephutomorplu unlUnwto, sp. d.
$. Nigra; capite foaco-femiginco, fronts et vertice nigro in-
taminatis; scapo fusco; flagello infra poUide testaceo; tborace
rofo-ferrugineo, brevi, latitudine vix longiore; segmento dorsali
prino fascia apicali pallide lutea; segmentis dorsalibus 2-^ fascia
mediona longitudinali continua albopilosa; area pygidiali late
triangulari, longitudinaliter striata; calcaribus pallidis.
?. Antennal tubercles rather large, with a low transverse carina
between them ; second joint of the flagellum short, no longer than
the third. Eyes round, strongly convex, situated nearer to the
bind margin of the head than to the base of the mandiblea. Head
narrower than the thorax, broadly rounded posteriorly; the front
rugose, the vertex closely punctured. Thorax closely punctured
rugose, scarcely longer than the greatest breadth, slightly narrowed
from the middle both anteriorly and posteriorly, the anterior margin
straight, the posterior slope almost vortical, aballowly nigoee.
Ij.y.l.AjhyCOO'^lL'
Hymenoptera in the British Museum. 57
[deurae almost Hmooth. First dorsal segment short, slightl^r
depressed below the second, which is nearly half as long ag^n as its
greatest breadth, finely and closely ponctoied, the sides somewhat
coDTex. The sides of the abdomen sparsely clothed with long
-whit« haira, intettiperBed with black near the apex. Hind tibiae
wHh two rows ot four spines.
Hab. S. Queensland, Briabane (Hacker), November.
Allied to E. argmteolineala, Audr^, but may be die-
tinguiahed by the colour of the legs, the difltinctly stronger
sculpture of the head and thorax, the slightly different
shape of the latter, and the distinctly longer second
abdominal s^ment.
Ephntomorpha setlsen, sp. n.
$. Nigra; tnberculis antennalibns pallide t«Btaceis; segmento
doisali primo margine apicali patlide tuleo ; segmento dorsali secundo
in medio longitodinaliter late, tertio, quarto, quintoque in medio,
aextoque basi bninneo-ferragineis; tareis fusco-femigineis.
Long. 6 mm.
Q. Clypeus small and flat, truncate at the apex ; antennal tubenlea
iftther large, second joint of the flagellum a little longer than the
thiid, less than twice as long as the first: a low carina from the
anteonal tubercles not reaching the eyes. Head closely and rather
strongly punctured, more sparsely on the front than on the vertex,
sparsely clothed with close lying fulvous bristles, interspersed with
which are a few long black hairs. Eyes round, strongly convex,
aitnated nearer to the hind margin of the head than to the base of
the mandibles ; the bead ronnded at the posterior angles, no wider
tban the thorax. Thorax granulate -rugose, more coarsely pos-
teriorly than anteriorly, nearly half as long again as broad, the
Bides almost smooth, the posterior slope finely rugulose, tiie sides
almost parallel, the anterior angles rounded, the doisal surface
clothed with short setae as the head, but with rather more numerous
k>og black hairs. Dorsal segments 1-5 with a small patch of sparse
white hairs in the middle of the apical maigin; second segment half
as long again as the greatest breadth, the sides convex, closely but
not very coareely punctured, the punctures confluent longitudinally.
Pygidial area triangular, finely longitudinally striated, the striae
not reaching the apex. Base of the abdomen with long whitish
hairs, the hairs on the sides of the abdomen whitish near the base,
mostly black near the apex. Hind tibiae with two rows of four
Hab. S. Queensland, Brisbane (Hacker), November.
iM-,i,A J h, Google
58 Mr. R. E. Turner on New Spedes o§
Family THYNNIDAE.
Zas^thynnas strittUroiu, sp. d.
(J. Niger; mandibnlia, apice excepto, dypeo macula BubapkwU
fnaca vtiinqae, otbitk, macula srcuata inter antemiae, pronoto
ma^ifie antico, ptapleuu, pronoto mvgine postico, tegolis, meso-
□oto macola magna quadrat* postice, meeopleuris antice et maculA
magna postice, sontello fascia latiwima, poatice sd apicem productA,
scat«Ui angulis anticis, postecatello, Bcgmento mediano fascia lata,
longitudinali ]at«ribiisc[ae, Begmentjs donialibiia 1-6, ventnlibtu
2-4 fascia tronsveTsa anguate inteirupta, s^mento primo ventrali
macula mediana, ooxisqne sabtus flavis; aegmento donali sexto
apice, eeptimoqne, ventralibus sexto septimoque pedibosque
fecTugineis ; alia aubbjralimB, venis feimgineis.
$. Nigra; capite femigineo, pedibus bnmneo-ferragineis, femori-
bus nigro-maculatii; Bt«mo, segmentiaqne ventralibna aecimdo
tortioque brunneis; piotborace postice nigro-marginato, ecuteUo,
poatscutello, aegmento mediano lateribus, segmentia doisalibus
1-3 fascia transveiBa undulata, segmentoqae dorsafi quarto faaoia
tranaversa interrupta flavis ; pygidio fusco.
Long. 5, 15 mm; $. 13 mm.
(J. 0ypeas pointed and prominent at the base and connected
by a short carina with the interant«nnal prominence, very broadly
truncate at the apex, the apical angles itot prookinent, Btiongly
longitmliaally atriated. Interant«nnal prominence very broadly
triangular, a low longitudinal carina from the apex reaching more
than half-way to the anterior ocellus. Front rogosel; punctured,
irregularly rugose striato above the base of the antennae; vertex
more epusely punctured. Antennae abont as long as the thorax
and median segment combined ; pronotum rather sparsely and finely
punctured, with a shallow shining groove behind the slightly raised
anterior margin j meoonotum very cloeely punctured between the
deep parapsidal furrows, but much more sparsely in the middle.
Scutellum very broadly rounded at the apex, subtruncate, rather
closely punctured ; median segment oblique from the postscnlellum,
finely rugulose, the middle rather sparsely punctured. Abdomen
somewhat elongate, the basal dorsal segment oblique anteriorly,
■B broad at the apex as the second segment, all the segments rather
sparsely punctured and not constricted; sixth rential segment
with a spine on each side at the apical angles; seventh dorsal
segment produced into a broad flattened lamella, longitndirmlly
striated and veiy broadly rounded at the apex. Hypopygium
obliquely striated above, subtriangular, broad at the base with a
l„y,l,Ajh,COOl^lC
Btftnaioplera in the British Museum. 59
distinct apine on each side at the basal angles, narrowly tounded at
the apex. Maxillae with a few long liain at the base beneath.
Wings sparsely clothed with very short hairs, third abscissa of the
radius mnch longer than the second, first recurrent nervure reoeived
a little beyond two-thiids from the base of the second cubital cell,
second a tittle before one-third from the base of the second cubital
celt.
?. MandibJea acute at the apex; clypeus punctured, without a
carina, truncat* at the apex. Front with a large concave area on
each side touching the inner margin of the eye and extending nearly
*to the vertex, smooth and sliining ; the ridge between the depressions
not quite as wide aa one of them, smooth and shining, with a median
longitudinal sulcoB and with a few punoturee on the sides ; vertex
shining, with a few scattered punctnrea. Head ucarly twice as
broad as long, rounded at the posterior angles. Pronotum broader
tlian the head, the anterior margin straight, slightly narrowed
posteriorly, the liind margin broadly arched, twice as broad anteriorly
as long, shaltowly punctured, with a few large setigerous punctnr«a
on the anterior margin. Scutellum smooth and shining, broadly
truncate at the apex; poBtscutellam plainly visible, transverse.
Median segment very short, the dorsal surface scarcely half as long
aa the scutellnm, widened from the base, almost vertically truncate
posteriorly, the dorsal surface smooth and shining, the face of the
Inmcation microscopically punctured. Dorsal surface of the
abdomen shining, with a few scattered punctures; first s^ment
^most vertically truncate anteriorly, witb a deep groove close to the
apical margin and a few indistinct, short, transverse striae close to
the apical angles ; second segment with numerous transverse striae,
which gradually increase in height from the base to the apex, the
hssal striae strongly curved, those at the apex straight. Vential
segments coamely, but not veiy closely, punctured ; the fifth seg-
ment coarsely longitudinally striated. Pygidium narrow, trans-
versely striated at the base, vertically truncate posteriorly, strongly
compressed before the truncation, the dorsal plate of the truncation
narrowly rounded at the base, broadly rounded at the apex, with
about ten arched striae, longer than the greatest breadth, scarcely
reaching beyond the middle of the truncation; the ventral plate
rather broadly truncate at the apex, finely, but distinctly serrate
at the sides. Basal joint of intermediate tarsi rather broad, with six
*tout spines on the outer maigin.
li(d>. Queensland, Stradbroke Island, Moreton Bay
(ff. Hacker), September 17, 1915.
Thifl is intermediate between Thynnus and Zasjnlo-
ih, Google
60 Mr. R. E. Turner an New Speciet <^
thynnus, and perhaps might be placed in the fonner g«nUB.
The nearest ally seems to be Tkymnts britbanensis, Turn.,
but it is aiso very near Zaspilotkipmns rufolvteus. Turn.
It differs from brisbanensts in the form of the hypopygiiim,
in the more oblique median segment, and in the less trun-
cate first dorsal segment, also in the colour of the legs and
apical abdominal segments. The distinct post^cutellum
of the female is remarkable.
Family SCOLIIDAE.
Stibfamilr ELIDINAB.
Elis (Heu) nursel. Turn.
Plesia nursei, Turn., Ann. and Mag. Nat. Hist. (8), iii,
p. 480, 1909, (J.
Elis (Mesa) nursei. Turn,, Proc. Zool. Soc. London, p. 721,
1912, cj-
$. Nigra, capita thoraceque cntsse, abdomine Bparee panctatis,
pygidio loDgitudinaliter rugose etriato, alls hyalinis, vends pallide
tefitaceiB ; tegulis teataceis, calcaribus albidis.
Long. 13 mm.
$. Clypeus with a low carina, strongly punctured, tlie ^ueal
margin smooth and transverse. Head coarsely punctoied, the
punctures on the front more or less confluent; scape smooth and
shining above, with a few strong piliferooa punctures beneath;
pronotum coarsely punctured, mesonotnm and scutellum rather
more sxHirsely punctured, the parapeldal furrows very distinct.
Median segment more finely punctured, not margined poaterioriy,
the posterior slope rugulose, two convergent longitudinal corinae
from the base close to the middle of the median segment. Abdomen
rather sparsely and shallowly punctured, the punctorefl latber
large; sixth dorsal segment longitudinally rugose -striate, narrowly
rounded at the apex. Basal joint of the hind tarsi with a row of
close set hairs beneath. Second abscissa of the radius veiy Uttle
shorter than the third, first recurrent nervure received just before
the middle of the second cubital cell, second just before three-
quarters from the base of the third cubital cell.
Hab. Simla {Nurse). September.
This is nearest to li. usltdata, Turn., but the sculpture
is much coarser and the sixth dorsal segment much more
narrowly rounded than in that species. E. nursei $ was
n,g,i,.eih,.G00glc
Bymeno^era in the British Museum. 61
identified by Cameron for Colonel Nurse as anthradna,
Sm.; but, as I liave pointed out in other papers, that
insect is an Anthcbcaca and is Australian ; not Indian, as
stated by Smith and Bingham.
Elis (Hesa) heteroehroa, sp. n.
$. Fusoo-fermginea ; mandibuliB apice, flagello, pleuris, St«nio,
Gozia poatioii snbtiu, segmento ntediano, segmento Tentrali primo,
ugmeittisqiie doraalibtu 1-4 margine apicali late nigris; calcaiibuB
albidis; alis pallide flaTO-hyalinia, venia fuBcia; segmento doreali
texto loQgitudinalitei etrkto, apice truncato.
3. Niger; clypeo apice, maiidibnlia apice excepto, maonla
tiansversa atrinque inter antenuaa, pronoto macula transveraa
mediana, tegulis basi, segmentis dorsalibus 1-6 fascia angusta apioali
lateribus dilatata, in medio angusto intemipta, segmentis ventra-
libni 2-6 macula porva apicali ntrinque, coxis subtus, femoribos
uitioii poBticisqne subtus et spice, intermediis apice, tibiis anticia
intennediisque supra, posticis basi, tarsisqoe hie illio infoscatia,
flavis; alia hyalinis, stigmate venisqne nigris.
Long. $, 18 mm; 3, 17 mm.
$. Clypeus sparsely but rather strongly punctured, with a low
median carina which does not reach the broadly truncated apex.
Head coarsely and closely punctured, more sparsely on the vertex
than on the front, antennal tubercles well defined. Scape smooth
above, beneath punctured, with long pale fulvous setae towards
lite apex. Pronatum coarsely and closely punctured, subcarinate
longitudiiiallj in the middle; acutellum and mesouotum rather
more sparaely punctured; pleurae more finely and shollowly
punctured. Median segment subopaqae, with a longitadinal
impressed line on the dorsal surface, the puncturation sh^ow and
obscnie, finer at the base than at the apex, the posterior slope
■hallowly punctured- rugolose, the sides of the segment very finely
and closely striated and clothed with short white pabescence. Abdo-
men shining, sparsely punctured, more closely on the fourth and lif th
than on the basal segments, ventral segments more coarsely punctured
than the dorsaL Sixth dorsal segment closely longitudinally striated,
the apical ■"a.pgin broadly smooth and rather broadly tnmcate at
the apex. Second abscissa of the radius as long as the third, first
nciirrent nervure received close to the middle of the second cubital
ceil, second jost before two-thirds from the base of the third cubital
cell. Basal joint of the hind tarsi with a close row of bairs l>eneath
mingled with a few spines.
j. Clypena finely punctured, clothed with white pubescence,
u„,,i,A J.., Google
62 Mr. R. E. Turner on New Speaes cf
slightl; convex, aubcorinate longitudinally at the base. later-
anteanal prominenoe bilobed ; the front rugose, the vertex punctond.
Head much broader than the thorax i rounded at the poBterior
angles; the posterior ocelli further from the ejee than from each
other; antennae about as long as the head, thorax and median
segment combined. Thorax punctured, more finelj and aparaelr
on the ecutellum than elsewhere, mesopleurae finely rugose; pro-
notum shorter than the mesonotum, the anterior margin straight,
the angles not prominent. Median segment closelj and strongly
punctured. First abdominal segment long; the petiole occupying
rather more than the basal third of the segment, rather short«T than
the baaal joint of the hind tarai; the apical portion moderately
swollen, the extreme apex a little constricted; second segment
flcaicely more than half as long as the first, broadened from the baae,
longer than the apical breadth ; third segment a little broader than
long; basal segment smooth and shining, the remaining segments
gradually becoming more strongly punctured; the punctures on
the sixth dorsal segment large but rather sparse ; seventh segment
smooth at the base, with a few large punctniw before the apex, the
apical emargination narrow, a little deeper than its apical breadth.
Second abscissa of the radius about equal to the third.
Sab. Nyasaland, Mlanje (S. A. Neave), December to
June.
The female is very near E. h^erogamia, Sauss., which
0CCUI8 in the same district, and may prove to be only a
variety of that species, but the difference in the colour
of the legs and abdomen seems to be constant. The nine
females in the collection were taken from January to June,
seven of them in February, specimens of heterogamia being
taken from December to June. The male has the fiist
abdominal segment distinctly more elongate than in
davata, Sauss., and the colour of the nervures is dilTeient.
In some specimens of the male the clypeus is wholly black,
and in some the yellow mark on the pronotum shows a
tendency to disappear.
Ells (Hesa) erythropoda, Turn.
Plesia {Mesa) en/lhropoda. Turn., Ann. and Mag, Nat.
Hist. (8), i, p. 505, 1908, $.
The type was from Lake Ngami. Ulr. Neave has sent
a series of the female from Mlanje, taken from January to
May. In these specimens the head is more sparsely
pimctured than in the type.
ih,Cooi^lc
Bj/meaoptera in the British Museum. 63
Elis (He») arnoldl, sp. n.
$. Nigra; nuuidifatilis dimidio apicoli fusco-fernigineiB ; fla^^o
sabtua fuaco ; pronolo propleurisqoe rufis ; calcariboB albidia ;
nnguicnlia pollide tefitBceia ; alia hyalinis, venia fuacia.
Long. 9 mm.
$. ClfpeuB subcarinate langitudinallj in the middle, the apical
margin tranareiae and slightly depressed. Front oloeelj and
atrongly, vertex and pronotam leaa closely punctured; mesonotum
and scutellum eparaely ponctured. Propleotae sparsely, meeo-
pleurae very doaely and atrongly punctured; median eegmeat
more finely and veiy closely punctured, with the usual Bhallow,
margined, longitudinal groove from the base, the sides of the segment
fiitely and closely striated. Abdomen rather closely and shallowly
punctured on the dorsal surface ; the ventral surface more finely
and closely punctured at the base of the segments, the grooTe be-
tween the two basal ventral segments very deep as in other species
of the genus. Sixth dorsal segment closely punctured at the base,
the punctures towaide the apex intermingled with fine longitudinal
striae, the apical ma^in broadly smooth and pale t«Btaeeous, broadly
rounded at the apex. Third abscissa of the radios twice as long as
the second, recurrent nervures received distinctly beyond the middle
of the second and thiid cubital cells. Basal joint of the hind tarsi
with a Bcopa of fine whitish hairs beneath.
Hab. 8. Rhodesia, Buluwayo {G. Amdd), March.
In colour this species somewhat resembles Anlhobosca
erythronota. Cam., but is generically distmct. The sculpture
of the sixth doiBal segment separates it from all other
African species of Elis, the segment being nearly always
striated; in E. ■peringueyi, Sauss., in which it is punctured,
the punctures are very coarse and sparse.
Pteiombius wiUiamsl, sp. n.
$. Niger; mandibulis ferrugineis; abdomine aegmentis tertio
qnartoqne apice lateribusque,, quinto, sexto septimoque omnino
mfo-femigineis ; alia hyalinis, venis nigris.
Long. 7 mm.
3- Head shining, sparsely and rather finely punctured ; antennal
tubercles well defined, fonnuig short longitudinal carinae. Jlan-
diblea bidentate at the apex; labrum transverse, with an apioal
fringe of hairs. Fronotum distinctly longer than the mesonotum,
tbe anterior ma^in distinctly raised, rather sparsely punctured;
mesonotum rather more strongly punctured ; scutellum as long as
iM:,i,A J 1^, Google
64 Mr. R. E. Turner on New Species of
the meaonotum; median segment very coaraelj rngoee, the ddea
finely and closely striated. Abdomen smooth and shining, petiolate ;
the fint segment u long u the second and third combined, Tciy
nanow at the baae, the apical half moderately swollen and slightly
constricted at the apex ; the second segment twice aa broad at the
apex aa at the base ; the recurved apical spine of the bypopygiom
rather shorL First tecmrent nervure received just before one-
quarter from the base of the second cubital cell, wcond just beforv
one-third from the base of the third cubital celL Third ahscioaa
of the radios as long as the second and first combined, second folly
twice as long as the first.
Hob. British Guiana, Bartica (C B. Williams),
September.
This is distinguished from other known males of the
geuus by the black basal and led apical segments of the
abdomen; the anterior margin of the pronotura is raised,
which is not the case in tflabricollis, DUcke, or confusus, Sm.
Family PSAMMOCHARIDAE.
The two following species belong to the wide-ranging
genus Episyron, in which the tarsal ungues are bifid in
both sexes, the cubitus of the hindwing originating just
before the transverse median nervure, the third cubital
cell much shorter than the second and strongly narrowed
on the radius, the first dorsal segment clothed more or leae
densely with scale-like hairs, and the male antennae rather
slender, with the ioints not arcuate beneath. The labrum
is sUghtly exposed.
1. Episyron hpldohlrtus, Turn.
Anoplius (Episyron) lepidohirtus, Turn., Proc. Zool. Soc.
London, p.' 331, 1910, 2 (J.
This is a true Episyron, nearly related to the European
P. Tufipes, the type of the genus. It is very closely related
to the New Guinea species E. papuensis, Sm.
Hub. N. Queensland, Mackay to Cooktown.
2. Episyron kurandensls, Turn.
AnopUus (Episyron) kurandensis, Turn., Proc. Zool. Soc.
London, p. 333, 1910,?.
ih,Cooi^lc
Hymenoplera in the Briiiah Museum. 66
The Bcale-like haiis on the Bist dorsal segment are not
very stronglj' developed in this species.
Aa6. N. QuEENSLAin), Kuranda.
Genus Psahmochabes, lAtr.
The Anstralian species still included in this genns fall
into several eections, which will doubtless eventually be
raised to generic rank ; but I consider it best to wait until
a more comprehensive revision is possible, in the meantime
defining certun groups of related species. One of these
groups, somewhat allied to Batozonus, Ashm., and Episyron,
8chiodte, has the tarsal ungues of both sexes bifid as in the
latter genus, but has the joints of the fl^ellum in the male
strongly arcuate beneath as in Balozonut, though shorter
than in that genus, and diSers from both in having the
cubitus of the hindwing interstitial with the transverse
median nervure. The median segment of the female is
truncate at the apex and more or less emarginate, with a
distbct median sulcus on the dorsal surface ; and the third
cubital cell is much narrowed on the radius. The sexual
dimorphism in the group is great, much greater than in
Episyron, though less than in Balozonus. The species
included are —
I. ftammoehares eonsimlUs, Sm.
Pompilus consimUis, Sm., Descr. new speues Hymen.,
p. 152, 1879, ?.
?. Nigra; front©, vertice, pronotoqne pilis breviHSimia auran-
tiacis, eabeqasmo«is, etratis, dense obtectia; poatscatella macula
utrinqnfl aegmentoque mediano angolis posticia albo-pUosts; seg-
m«ntu dorsaliboB 2-3 fascia transversa grisea utrinque ; alis fosots,
apice obBcurioribns, veoia fuscis.
iJ. Niger; antennis fusco-femigineia, scapo supra infuscato,
■^mento dorsali aexto albido; s^pnenlo mediano angolia posticis
aJbo-piloHis ; fronts, pronoto, scgmentis doisalibos 1-3 fascia lata
b«sali, qoarto quintoque omnino griseo-pubesoentibua; alls fusco-
hjalinis, apice obscarioribus.
Long. ?, 14 mm ; ,J, 12 mm.
9. Clypeoa widely but vny shallowly emarginate at the apex,
the Ulmtm exposed. Head somewhat flattened, the posterior ocelli
Deariy twice aa fat from each other as from the hind margin of the
head. Second joint of the flagellum lees than half as long again as
TBAlfS. ENT. 80C. LOND. 1917. — PART I. (NOV.) F
iM:,i,A J h, Google
66 Mr. R. E. Turner on Nea Speciea cf
the third. Pronatum aiched poeteriorly, not angnlaUt, almcwt m
long &B the mesonotam ; gcutellum depraoed ftt the sideft. Mediao
segment looger than the meaonotum, Tith a distiiict median aulcoa.
rather Btrongly emarginate at the apei, the apical alape almost
verticaL Second ventral Kgment convex, pygidium smooth.
Basal joint of the fore tarsi with four short spines, all the ungoM
bifid Third cubital cell pointed on the radius, subpetiolate.
cf. Cljpeus broadly and shallowly emarginate, the labniin ex-
posed. Head flatt«ned, the front subcarinate longitadinallj' from
the anterior ocellus but grooved on the lower part towards the base
of the antennae, the posterior ocelli very near Uie hind margin of the
bead ; antennae stout, tapering towards the apex, very little longer
than the thorax and median segment combined, fifth to eleventh
joints of the flageUum strongly arcuate beneath. Hind margin of
the pronotum with a distinct angle in the middle. Median aegmcnt
without a distinct sulcus, the apical slope less abrupt than in the
female and leas deeply emarginate. First abdominal segment long,
second ventral segment convex, with a distinct, though ahallow,
transverse impressed line near the base. Third cubital cell not quit«
pointed on the radius, but the third abscissa of the radius is ex-
tremely short. All the tarsal ungues bifid. The distanoe between
the base of the mandibles and the eyes is considerable.
H(A. Queensland, Mackay iJ'uTner), March ; Brisbane,
{Hncker), January and February.
The naale has not been previously described ; the descrip-
tion is taken from two Mackay specimens; the specimen
from Brisbane is without the transverse groove near the
base of the second ventral segment, but does not differ
otherwise. The female shows no trace of such a groove,
the presence of which in males is most remarkable. The
female has a strong superficial resemblance to Paammo-
chares frontalis, Fabr., which belonj^s to another section
of the genus and has the tarsal ungues unidentate, not
bifid.
2. Pummoohares doddl, Turn.
Anoplius doddi, Turn., Proc. Zool. Soc. London, p. 328,
1910, ?.
j. Niger; fiagello articults secundo tertioque subtus fulvis;
olypeo macula magna utnnqoe, pronoto maipne poateriore, aeg-
mento dorsaU tertio fascia bssall intemipta, segmento dorsali
septimo, calcariis, tibiisque posticis supra albido flavia; alis hyalinia,
fascia apioali fusca, extiemoapice hyalinis. Vor. a
Ij.y.l.Ajl^yCOO'^lL'
BymeKOptera in the British Muwum. 67
Mignlni afHCAlibos, aegmoitoqae secimdo donali dmcuIa baaaii
ntrinqne flftTiB.
Long. S-d mm.
^. ClypooB tnmcftte at the ftpex, the Ubrum expoeed. Fomth
to eloTentb joints of the flagellum strongly uciute beneath; eyM
only narrowly sepuated from the base of the mandibles. Head
aomewhst flattened, the posterior ocelli very near the hind margin
of the head. Hind margin of the pronotum strongly aiohed, not
uigulate. Median segment with a distinct longitudinal sulcus,
shorter than the mesonotum, steeply sloped posteriorly, but not
abruptly truncate, with patches of white pubescence at the apical
angles. First dorsal segment rather densely clothed with scale-like
hairs, as in the genus Epitf/ron, fourth, fifth and sixth segments
covered with grey pubescence; second ventral segment convex,
aerenth with a longitudinal carina. Third abscissa of the radius
about h^ aa long as the second.
ffofc. Queensland, Townaville {Dodd) ; Mackay (Turner),
January to March ; Brisbane and Stradbrolce Island
(Hacker), October to February; Northern Territoby,
Alexandria [Stalker), January ; Hermannsbui^ (Hillier),
April.
Tbe female iaeasily distinguished by the fasciate forewings.
The following species are very closely allied to Episyron,
bat differ in having tbe third cubital eel) much larger than
the second and the cubitus of the hindwing in most speci-
mens interstitial with the trans verae median nervure.
The forewings are crossed by two fuscous fasciae. They
form a generic group.
1. Piammoetiues limpldns. Turn.
Anoplius [Episyron) limpidva. Turn., Proc. Zool. Soc.
London, p. 332, 1910, ?^.
This is not a true Epiayron, and pending a revision,
most be retained in Vsamrnochares.
Hah. N. Queensland, Euranda.
2. Runmoehtru mulrl, sp. n.
$. Nigra 1 mandibulia basi, clypeo dimidio apicali, scapo, pionoto
margine postioo, pedibusque anticis brunnoo-ferrugineiB ; flsgello
snbtns, t«giilisque fnsco-ferrugineis ; alis byalinis, antioia fusco
bifaactatiB.
Long. 8 mm.
$. ClypeuB transverse at the apex, clothed with delicate white
iM:,i,A J 1^, Google
68 Mr. R. E. Turner on Neu> Species of
pnbeaoenae, microKopically panctured, with a veij ap&ise traiu-
Tene row ol large puooturea neat the middle, from each of which
Hprings a long bUok hur. Eyes diatinctlj diveigent towards the
clypeUB ; the posterior ocelli as far from each other as from the eyea ;
a distinct frontal Boloue reaching the anterior ocellua. Antennae
slender, the second joint of the flagellum as long as the fiist snd third
oombined. Pronotnm strongly arched posteriorly, not angulate
in the middle; aontellnm nthet strongly compressed laterally,
the dorsal surface small. Dorsal sutfoce of the median s^ment
shorter than the scut«lliiffl, much broader than long, without a
median sulcus; the posterior slope oblique and clothed with short
whitish pubeecence. Abdomen opaque, the basal dorsal segment
clothed with greyish soale-like hairs; uzth doisal segment finely
punctured and sparsely clothed with long fuscous hairs, elongate
triangular; second ventral segment strongly convex. Third
abscissa of the radius longer than the second, as long as the second
transverse cubital nervure, the cubital margin of the third cubital
cell half as long again as that of the second ; first recurrent nervure
received at two-thirds from the base of the second cubital cell,
seoond just beyond the middle of the third cubital celL Cubitus of
the hindwing interstitiaL Fore tarsi very feebly spined, the basal
joint with three short spines ; hind tibiae feebly spinose. The basal
fascia of the forewing is not very broad and is situated on the outer
side of the basal nervuie, not reachirkg the costa; the second (asci»
veiy broad, crossing the wing, and occupying the whole of the radial
and second and third cubital cells.
Hab. Ahboinj {F. Muir), October.
Easily dUtingiiisheii from limpidus by the colour of the
antennae and legs, but very close in structure.
Another group has the tarsal ungues unidentate in both
sexes; the joints of the Sagellum iu the male not arcuate
beneath; the second cubital cell always longer than the
third on the radius, though not always on the cubitus ; the
median segment with a sulcus, usually strongly developed
from base to apex; and the fore tarsi of the female in most
species strongly spinose. The species are all black, with
white or grey pubescence, rarely with orange close-lying
hairs on the pronotum. To this section belong —
1. hammoohares senex. Turn.
Anopliu3 senex. Turn., Proc. Zool. Soc. London, p. 327,
1910, $.
Hab. Victoria.
n,g,t,7.dh,G00glc
Hymenoptera in the Brituk Mttaeum. 69
2. FsanunoehftTM itevoi, Tuid.
Anoplius atamia. Turn., Proc. Zool, Soc. London, p. 326,
1910, 9.
Sab. N.W. AUSTRAUA.
3. hammoohares ahrlnuuies, Turn.
Anoplitts ahrimanea. Turn., Proc. Zool. Soc. London, p. 326,
1910, 9.
H<A. N. QUEENSLAMD.
4. fsunmoflbaies pinto, sp. n.
$. Nign; alie nigro-vioUcetB ; Bagello aubtns taaoo; nnguioolia
nnidentatia.
(f. Feminae Bimillimag; scapo mbtus bnmeo, orbitia intemis
angnato SA-ria ; ongiiiotiUB nmdentetiB.
Lcmg. ?, 17 mm.; J, 14 taat.
$. Clfpens broadly tnmcate at the apex, the labnim Dot exposed ;
second joiut of the flagellum as long as the first and thiid oombined,
the third distinctly longer tbaa the fourth. Eyes slightly divergent
towards the clypeos, separated on the vertex by a distance equal
to the length of the third joint of the flagellum. Fronotum shorter
than the mesonotum, the posterior margin broadly arched, withont
a distinct angle; scutellom narrowly truncate at the apex. Median
segment broader than long, without tubercles, rounded at the
posterior angles, gradually sloped posteriorly, with a deep median
sulcus from the base. Abdomen opaque, with a few long black
luurs on the sides near the apex; sixth doisal segment rather
broadly rounded at the apex and sparsely clothed with long black
setae. Basal joint of fore taiaus with three moderately long spines
cm the outer margin, hind tibiae spinose, tanal ungues with one
tooth. Third abscissa of the radius a little longer than the first,
and more than two- thirds of the length of the second, submedian
cell equal in length to the median, second recurrent nermre received
beyond the middle of the third cubitjil cell, cubitus of the hind-wing
interstitial with the transverse median nervure.
3- Antennae not stout, the joints very slightly arcuate beneath,
second joint of the flagellum scarcely longer than the third. Clypeus
widely emarginale at the apex, the labnim exposed in the arch of
the emaigination. Median segment as long as broad, without a
u„,,l„eJh,G00glc
70 Mr. R. E. Tomer m New SpecUs of
Bulciu. T&nal nngoes miidenUto, hind tibiae modentely apinoae ;
neoiHtion of wings u in the fenulc
Hob. Mackay, Q. {Turner), 1 ?, November 1897; 1 <?,
February 1899.
5. Fsammoehana frontalis, Fabr.
Sphex frontalis, Fabr., Syst. Entom., p. 349, 1775, 2.
P<mp{lus frontalis, Fabr.. Syst. Kez., p. 188, 1804, °.
The male is simiUr to the female, but has tbe pubescence
on tbe head and pronotum greyish instead of oranp;e.
The seventh dorsal segment is broadly truncate at the
apex, the seventh ventral segment carinate longitudinally
and produced into a short spine at the apex.
Hab. Queensland, Brisbane and Mackay.
6. hammocbantsamlluolnosm, Sm.
Potupihis semilvcluosus, Sm., Cat. Hvm. B. M., iii. p. 166,
1855, 5.
This is a large species, with six long, slightly spatulate.
spines on the basal joint of the fore tarsi.
Hab. Sydney.
7. PsuiinoehaTes berthondi, sp. n.
$. Nigra, fronte, cljpeo lateribuH, pronoto maigmibns, meeonoto
fascia transversa apicali, scuteUo angulis tuisalifaiu, poatscutello
macula magna mediana. maculaque minore utrinque angolia
apicalibus, meeopleuris fascia ofaliqua, eegmento mediana angulis
baBalibuB et apicalibus, aegmento dorsali primo roacula utrinque.
Begmentis dorsatibus 1-6 macula tranaversa apicali utrinque, aeg-
mentiaque ventralibue secundo teitioque macula utrinque anf^ia
apicalibuB albido-pubeacentibus ; alia fuacia. apice obacurioribns,
venia nigria; metatareo antico spinis aeptem longis instmcto.
Long. 23 mm.
$. Clypeua broadly truncate at the apes; second joint of the
flagellum very long, nearly as long as the third and fourth com-
bined; front flat, without a distinct aulcua, posterior ooelii much
nearer to each other than to the eyes. Pronotum strongly arched
posteriorly, not angulat«; scuUllum with a broad donal surface,
not very strongly compresaed laterally. Median aegment aoaicely
longer than the pronotum, much broader than long, the sulcus on
the dorsal suriace very deep, the lateral tubercles not developed.
Ij.y.l.AjhyCOO'^lL'
Hymenvptera in the British Museum. 71
Sixtb aomtl Begment broadly rounded At the apex, with ap&ne
ponctiiTes, each of which bean a long bUck h&ir. Baaal joint of
the fore taisi with eeven long alightly apatulate Bpinea; inter-
mediate and bind tibiae with a patch of grej pnbeaoence at boM
and apex, tore tibiae with a line of grej pubeeoence on the onter
ode. Tanal nngnes nnidentaite. Second abeoisia of the radiua
more than half u long ^;ain as the third, the second cubital cell
as long on the cnbitna as the third ; fint reoturent nerrure received
at about one-sixth from the apex o( the eeoond cubital cell, seoond
close to the middle of the third cnbital cell. Cabitiu of hindwing
intcoMitiaL
Hob, S.W. AusTRAUA, Waroona {G. F. Berthoud),
January.
Thifl 13 a western fonn of semiluauoais, differing little
from that species except in the presence of an additional
spne on the basal joint of the fore tarsi, and in the much
more broadly interrupted abdominal fasciae.
8. ^»mmoeh»res melanehoL'eiis, Sm.
PompUus mdanckolicus, Sm., Trans. Eot. Soc. London,
p. 244, 1868, ?,J.
This species closely resembles semiluciuiMus in colour,
but is much smaller, the joints of the flagellum are less
elongate, the abdominal fasciae continuous, the sixth
dorsal segment much more nairowly rounded at the apex,
the basal joint of the foie tarsi with only three spines,
and the median segment with an oblique band of cinereous
pubescence on each side near the apex. The male has
the apical ventral segment with a low longitudinal carina,
the hairs on the segment and the apical fringe short ; the
sixth ventral segment almost flat, not strongly convex.
H(d>. S.W. Australia, Yallingup (Turner), October to
December; Champion Bay {Du Boulay).
Smith's type is unfortunately lost, but his description
of the species as only 5 lines long agrees with the present
species much better than with the larger allied species
occurring in the same locality. Mv specimens measure
?, ]3mm.i J, 7 ram.
9. Psammochtres vassei, sp. n.
S. DiSeiB from mtlaiKholKaa in the larger bim, the female
mm., in the much broader apex of the aixth dorsal
iM:,i,A J 1^, Google
72 Mr, B. E. Turner cm New Species cf
B^ment, in the more convex baae of the seoond Tentral SegmeDt,
and in the preeence of a fourth spine on the baoal joint of the fore
tUBua.
(J. DiSera from mda»eholicu» in the larget size, 11 mm. ; in the
strongly oompieesed sides of the apical ventral segment, which
leave the mediui carina veiy prominent, the segment is also covered
with long haiis, vhich are especially conspicuous along the carioft
and on the apical fringe ; the sixth ventral segment is more convex
than in melawAolicaa, and the fasciae of cinereous pubescence
which in melancholieat are confined to the three basal segments
extend in vcuwi to the sixth.
H(d>. S.W. AusTRAUA, YalliDgup {Turner), December,
10. Psammocbftres basstanns, sp. n.
$. Ooeely allied to vaseei, but has three spines only on the basal
joint of the fore tarsi, the sixth dorsal segment is quite distinct,
the apical half of the segment being smooth, shining, and slightly
concave in bastianiu, the apex a little produced and not very
broadly roundbd, in vassei the segment is slightly convex throughout,
the apical half subopaque and very closely and minutely punctured
and very broadly rounded at the apex; in mdandioUeui the Mo-
ment is more noirowly rounded at the apex than in either, convex
throi^DUt, the apical quarter shining and almost smooth.
Hob. Tasmania, Eaglehawk Neck {Turner), February,
Unfortunately I did not take the male. The group of
mdaticholictis seems to be very extensive and the species
very closely allied, but differing especially in the apical
segments of both sexes. I have two oi three Eastern
Australian forms, but have not sufficient material from
which to describe them.
II. haminoeliani bbills, Sm.
Pompilus labilis, Sm., Descr. new species Hymen., p. 151,
1879, (J (as ?).
The thorax in this species is without pubescence in the
female, but the pronotum of the male is covered with
close-I}"ing whitish hairs, the head also being covered with
similar hairs in both sexes except on a transverse band
across the ocellar region. The colour of the pubescence
varies from whitish to dull yellowish. The female has
three long spines on the basal joint of the fore taiai.
l„y,l,Ajh,COOl^lL'
Hymenoptera in the BrUish Museum. 73
Hab. Queensland, Townsville {Dodd), January and
February; Nobtherk Territory, Port Darwin ((?, F.
HUl); N.W. Australia, Nicol Bay (Dr. Clement).
12. Fsammocharei baslUcus, ap. n.
$. Nigra, captte Intco pnbescente, laaoia tiaiuverea in regions
oc«llari nnda; s^mento donali faaoia Bngoata apicali BUbint«r-
rupta, secundo tertioque macula transTeraa utrinque oinereo-
pnbeacente ; s^mento mediano Uteribns ante apicem distinctiseinie
tubercnlato.
Long. 19 mm.
$. Veiy near P. lo&ilis, but may be diBtingnisbed from that
■peciee by the absence of the very broad abdominal fascUe, by the
preeeDce of four long spines on the baaal joint of the fere tarai;
tbe median aegment has a very deep median buIcub and a blunt
tabeiole on each side before the apical angles, tbe tubercle being
almost obeolets in labilis; in both the apical dona! segment is
broadly rounded at the apex, but the puncturation of the segment,
though Teiy minute in both, is more distinct in bibilU. Tbe second
abscissa of the radius is scaroely twice as long as the third in baaUieu*,
and in West Australian specimens of labUia, but about four times as
long in Queensland specimens of labUis,
Hab. N. Queensland, Townsvilie {Dodd).
Nearly allied to this group but difiering in having the
submedian cell distinctly longer than the median in both
sexes and the tarsal ungues in tbe male bifid are the two
following species —
1. Fsammoebans unoflanlai. Turn.
Anoplius amoemilus. Turn,, Proc. Zool. Soc. London,
p. 329, 1910, $.
HiA. Queensland, Brisbane and Mackay.
,2. Rammoeharei elatus, Sm.
PompUtts elaius, Sm,, Joum. linn. Soc. Zool,, viii, p. 82,
1864,9.
Hab. Queensland, Mackay and Townaville ; Moluccas,
Morty Island.
Another generic group is easily distinguished by the
coarse granulation of the thorax, especially strong on the
median segment ; by tbe very long submedian cell, which
is much longer than the median ; by the very short third
u,y,i,Ajh,Cooglc
74 Mr. R. E. Turner on New Species of
cubit&l cell, which is twice aa long on the cubitus as on
the radius, but no longer on the cubitus than the length
of the third transverse cubital nervure ; by the red colour
of the thorax and median segment, and by the unidentate
tarsal ungues. To this group belong —
1. Pummoeluns eirthroslelhiis, Sm.
Pompilus erythrosteihua, Sm., Cat. Hvm. B. M., iii, p. 162,
1&55, O.
The male of this wide-ranging species is unknown, but
I have seen a male closely resembling this female, but
with red legs, from Western Australia.
Hab. S.W. AusTRAUA, Yallingup (Turner), November
to January ; Tasmania, Eaglehawk Neck (Turaer),
February; Queensland, Mackay.
2. FsammoflhareB perpuleher, sp. n.
$. Bufo-femigiiiea; oapite nigro; scapo aabtns, maodibnlk
cljrpeoque femigineiH; tarsis infoscatis; alii hyalinis, tertio apioali
taaciaque mediona hiscis.
Long. 7 mm.
S. Second joint of the flagellum as long as the first and thiid
combined; the olypeua and lower part of the front clothed with
silver pubescence. Head finely and very cloeely punctured;
posterior ocelli further from the eyes than from each other. Thorax
closely and very distinctly punctured; pranotum veiy widely
anihed posteriorly ; scutellum subqoadrate. Median segment
coarsely granulate, without a sulcus, with short, spare, white
pubescence. Abdomen shining, the punctures microscopic; sixth
dorsal segment with a few large scattered punctures, broadly
rounded at the apex. Fore tarsi with a short comb, the baaal
joint with four short spinea. Cubitus of the hindwing originating
just beyond the tiauaverse median nervure.
Hab. Queensland, Townsville {Dodd], Aiigust.
Differs from erylhroalethis in the smaller size, the sculp-
ture of the thorax, the colour of the abdomen and legs,
and the much shorter spine.'; of the fore taisi.
Agenloldens ezpulsus, Bp. n.
$. Nigra ; ptonoto margine poetico late, tibiisque posticis macula
basali albidis; calcaribus palUdis; alis hyalinis, ontiois fusoo
bifasoiatis, Tenis nigris.
l„y,l,Ajh,COOl^lC
Bymatoptera in the British Museum. 75
<J. Feminac amilis; tdbiie intermeduB etiom basi albido mftou-
Ifttis; segmraito doiwdi septimo olbido; alia fascia baeali sub-
obaoleta.
Lraig. $, 7-8 mm. ; J, 7 mm.
$. Clypena abort and btoodly truncate at the apex; antennae
abont oa long oa the thorax and median segment oombined, the
second joint of tbe flagellum as long aa the first and third com-
bined. Pronotum broadly acouate poBt«riorly, no't angulatc;
acutellum convex. Median segment alightlj convex, oblique, the
dorsal and poeteiior surfaces. not separated, rather thinly covered
with short white pnbeecence. Abdomen subopaque; the apical
doTBoI segment aabtriongalar, with a few rather long dark hairs.
Torvot ungues iuiidentat«. Cubitus of the hindwing received at a
distance beyond the tranaveise median nervure equal to the length
of the transverse cubital nervure ; submedian cell of the forewing
very slightly longer than the median ; third abscissa of the radius
scarcely more than half as long ss the second, the third cubital
cell shorter on the cubitus than the second. The fascia on the
basal nervnre narrow, that from the radial cell broad, filling the
aeoond and third cubital cells.
3- Tarsal ungues as in tbe female, but the tooth rather nearer
to tbe apex.
S(A. Queensland, Mackay {Turner), November to
March.
This seems to be undoubtedly congeneric with A, kumUis,
Cress., from N. America, the type of Ashmead's genus.
The description of tbe genus, however, is far from accurate.
The cubitus of the hindwing in all species of the genus
known to me originates well beyond the transverse median
nervure, though not so far as in the present species. The
Indian PompUus maadipes, Sm., also belongs to this genus.
Genus Austbosalius, gen. nov.
$. Head small, flattened, rather longer than broad; clypeus
very short, broadly truncate at the apex ; the labrum exposed ,
emsrginate at tbe apex. Pronotum long, sometimes longer than
the mesonotum, sometimes a little shorter; median segment either
verticoUy or obliquely truncat«, opaque and smooth, with a median
sulcus, the sides parallel, without tubercles; abdomen rather
nuTow, convex, the sides of the four basal segments almost parallel ;
the transverse line on the second ventral segment visible, but not
deeply impressed. Fore femora very stoat; fore tarsi without a
iM:,i,A J 1^, Google
76 Mr. R. E. Turner on New Species cf
oomb; hind tibiae with a few gmall spines, not senate. Sooond
absoisM of the radios st least as long as the thiid, the raonmott
nermres received *t the middle of the second and of the thiid
cubital cells; submedian cell of the foi«wing very slightl; longv
than the median; cubitus of the hindwing either intentitial with
01 originating just beyond the transverse median nerrure. Hale
unknown.
Type of the genos Ferreolomorpka artemis. Turn,
1. Auitrosftlius artemis. Turn.
Ferredonwrpha artemis. Turn., Froc. Zool. Soc. London,
p. 325, 1910, $.
Hab. Mackay, Q. (Turner), October to March.
2. Anstrosallus mtllgnuB, Sm.
Salius malignus, Sm., Joum. Froc. Linn. Soc. Zool., iii,
p. 157, 1858, O.
This is the only other species of the genus known to me.
It is much lai^er than artemis, and the mesonotum b much
longer, the colour of the antennae and of the wings is also
different. It has not been recorded from Australia.
ffofi. Asu {Wallace).
Nearly allied to this genus are the South Indian Psettda-
genia rava, Bingh., and an allied species which seems to
be undescribed ; but in which the submedian cell is much
longer, the head rather larcer and the labnim not so
strongly exposed. The tarsal ungues are bifid in these
»>ecie3, not unidentate as in Austrosalivs. For this genus
I propose the name
Cbyptosalius, gen. nov.
Type of the genus PseuSagenia rava, Bingh.
1 . Ciyptosallns nvs, Bingh.
Pseudagenia rava, Joum. Froc. Linn. Soc. Zool., xxv,
p. 426, 1896, 9.
Hab. Bangalore {Bingham) ; Coimbatobe {T. V. Rama
Krishna), Februarj-.
l„y,l,Ajh,COOl^lL'
Bjftnenoptera in the British Mateum. 77
2. Cryptosalins pandiyinni, ap. d.
$. mgn, albopiloea; mandibolis ferrnglDeis, apice nigra; (ui-
tennie tuekque antiois fuBoia; alis flnvahyolinia, &pice late infn-
matiB, venis tcstaceia ; vertioe, pronoto poatice, Begmento mediano
poetice, aegmentis donalibus 1-4 fascia lata apicali, segmentoqae
Bezto, dense albopabeacentibtu; pronoto elongate, meeonoto dnplo
longiore.
Long. 6-8 mm.
$. Head longer than the greatest breadth; clTpens short, the
apical margin wideijr and very shallawly emaiginate. Posterior
oe^ about half as far again from the eyes as from each other;
inner maigina of the eyes parallel ; the front with a distinct bnt
Tuy shallow longitudinal sulcus extending to the anterior ocellus.
P^ont, pronotnm and median segment, except broadly posteriorly,
clothed with veiy short and sparse fulvous pubescence. Antennae
short and stout, not more than twice as long as the head, second
and third joints of the flagellum subequat. Pronotum at least
twice as long as the meeonotiim, nanower than the head, as long
■a the greatest breadth, very slightly narrowed ant«riorIy. Meso-
notora very short; median segment shorter than the pronotum,
obliquely sloped posteriorly, the sides paialleL Second ventral
a^mcakt with a distinct transverse groove near the base. The
whole ventral surface clothed with veiy delicate silver pubescence.
Anterior tibiae produced at the apex without and ending in two
spiues; hind tibiae smooth. Second abscissa of the radius longer
than the third ; first recurrent nervure received close to the middle
of the second cubital cell, second at about three-quarters from the
hue of the third cubital cell; submedian cell much longer than
the median, cubitus of the hindwing inteistitial with the transverse
median nervure.
Bob. CoiHBATOUE, S. India {T. V. Rama Krishna),
January to August.
This ia a much smaller species than C. rava, and may
be distinguished by the very much longer pronotum,
which in rava is much broader than long and no longer
than the mesonotum. The third cubital cell is shorter
than m rava and receives the recurrent nervure nearer to
the apex. There ia some difference in the present species
in the length of the third cubital cell, specmiens t^ken in
July and August having the third abscissa of the radius
as long as oi longer than the second. The type was taken
in January. There ia also some variation in the position
ll,y,l,/eJh,G00glc
78 Mr. R. E. Turner on New Species <f
of the second recurrent nervure. But I regard these as
individual variations only.
Cnrptoehilni commixtBl, Turn.
Cryplocheilus commixius, Tom.. Proc. Zool. Soc. London,
p. 317, 1910, 5 J.
This species belongs to the group in which the male has
the tarsal ungues bifid, while they are unidentate in the
female. This structure is found in a considerable number
of oriental and Malayan species, also in one or two Ethio-
pian species, but not as far as I know in any other Australian
Cryplochilus. SuperficiaUy the species closely resembles
Hanipepsis auatralasiae. but is not as plentiful and has a
more Umited range.
Hab. Mackay and Cairns, Q.
The nearest ally is C. basimacula. Cam., from New
Britain.
Genus Cyphononyx, Dahlb.
Cyphononyx, Dahlb., Hymen. Europ., i, p. 461, 1843.
This genus was founded on the single character of the
bifid tarsal ungues. But I am very doubtful if it will
eventually stand, owing to the fact that a considerable
number of species have males with the bifid tarsal ungues
°\ ^yP^°*^^y^ aid females with the unidentate ungues
of CryptocheUus. At present I retain the name for those
species m which the tarsal ungues are bifid in both sexes.
Taken m this sense there ia only one Australian species of
the genus.
Cfphononyi aspasU, Sm.
Mygnimiaaspasia, Sm., Joum. Proc. Linn. Soc. Zool., iii,
p. 157, 1858, $,
A beautiful large species with yellow wings and the
abdomen glossed with blue.
A ^l^ii/'J^^^Vf^.^V^"'"*' **■ trw"'«-); New Guinea;
Am (WalUKe]; K6 {Stalker).
Cyphononyx vitiensis, sp. n.
Mo. poati^teUo apic«, «gmentu. shdominaUbna tertioTS
qu»rto, qumu. «xtoque. pedibuaque fulvo-fenugmeta; <Ji. ^.
Bymen&ptera in the Brtti^ Museum. 79
auantiftcis, uiticia nutcuU nugns medi&iut fusco-purpuieft, apice
leviter iofuscatis, Tenis ferragineia.
(^. Feminae Bimilia ; meeoDoto fosco-fenugiDeo.
Long. ¥, lS-21 mm.; ^, 18 mm.
$. Clypens bnud and tnuuveise at the apex; labnim Boarc«ly
expoaed, very btoAdly rounded at tbe apex and iritfa a fringe of
long fnlrouB haira. Auteonal prominence somewhat ponect, veiy
feebly bilobed and divided by a strong loagitadinal sulcna. Second
joint of the flagellum a little longer than the firat and third com-
bined- Posterior ocelli farther from tlte eyea than from each
other. Kvnotam short, rounded at the anterior angles, the posterior
margin vei7 broadly arched. Head and pronotum sparsely, mceo-
notnm and acutelhim closely, covered with short dark golden
pubescence, the mesonotum and Bcutellum very finely and cloaely
punctured; scutellum with a flat dorsal surface, broadly rounded
at the apex; poslacntelluni subcariuate longitudinally in the
middle, slightly produced in the middle posteriorly and rounded
at the apex. Median segment with a blunt tubercle on each side
at the base, with a deep median sulcus from base to apex, opaque,
with very minute and indistinct transverae striae; tbe posterior
dope oblique, not abruptly separated from the dorsal surfaoe,
spanely clothed with fuscous hairs. Abdomen highly polished,
with a few small scattered punctures; sixth dorsal segment more
coarsely pnnctuied, narrowly rounded at the apex, densely clothed
with golden pubescence intermin^ed with coarse folvons setae.
Hind tibiae rather feebly serrate; tarsal ungues bifid; spines of
the fore tani rather short. First recurrent nerrure received at
about three-quarters from the base of the second cubital cell, second
just bc&re the middle of the third cubital cell; third abscissa of
the radius nearly half as long again as the second; cubitus of the
hindwing interstitial with the transverse median nervure; sub-
median cell of the forewing longer than the baaaL
<}. Antennae very long, measuring 16 mm., the whole length of
the insect being 18 mm. ; groove near tbe base of the second
ventral segment very distinct, quite as deep as in the female;
seventh dorsal segment very broadly rounded at the apex; sixth
ventral segment widely emarginate at the apex, with a short acnte
spine on each side at the apical angles; seventh ventral segment
flat, subqnadrate, broadly truncate at the apex.
Hob. Fur {British Museum ex Crawley Coll.); Fiji
(R. C. L. Perkins), Fiji, Natova, Nadi (ff. Veilck), October.
Not very nearly related to any other species known to
me, though nearer to the group of C.flavus, Fabr., than
u,y,i,Ajh,Cooglc
80 Mr. R. E. Turner m New Species of
to C. aspasia, Sm., and the related species C. trUrepida,
Sm., and C. cmfector, Sm.
Gentis Hemipbpsis, Dahlb.
Hemipepsis, Dahlb., Hymen. Europ., i, p. 123, ISiS.
I think this is a good genus and distinct from Crypto-
ckeUvs, Panz. After muot consideration I disagree with
Schulz, who uses Shuckard's name Mygnimia for the
genus. This name was pubhshed in 1840, and therefore
has priority over Hemipepsis. But no species are given
in Shuckard's work, and the single character given, the
position of the first recurrent nervure, would apply to other
species not included in Hemipepsis, such as some species
of Cyphononyx, equally well. Thus Shuckard's name
should, in common with other names in the same work,
be treated as a nomen. nudum, and only date from Smith's
publication of the name in 1855. There is only one
Australian species of Hemipepsis.
Hemipepsis austnlutae, Sm.
Mygnimia amtralasiae, Sm., Ann, and Mag. Nat. Hist. (4),
xii, p. 259. 1873.
This seems to me to be the Australian form of the
Indo-Malayan H. atireosericeua, Gufir. It has a wide
range in Australia, especially in the northern half of the
continent.
Bab. Cooktown, Cairna and Mackay, Q.; Hermanns-
burg, Central Austraba; Fowler's Bay, S.A.
Family CRABRONIDAE.
Subfamily AMPVLICINAE.
Ampules orawshayl, sp. n.
$. Tiridi-cyanea: antennk taiaiaque nigria; m&ndibulis feiru-
gineiB; alia tayalinia, venia nigris, onticia in cellulia radiali, oubitali
seouudo discoidEiliqne secundo levit«r infuacatis; abdomine seg-
mentia apicalibua fortiter comprMsia.
Long. 20 mm.
$. Carina of the clypeua sharply bent donnworda near the apex,
produced into a abort blunt apical tooth. Second joint of the
flagellum abont equal to the combined length of the third and
ih,Cooi^lc
Hymenoptera m the BrUish Museum. 81
fourtli; tjee BepBrat«d on the vertex by a distance slightly leea
thma the length of the second joint of the flagellum. Head etrongly
ragoeely poDctured, less coaisely on the vertex than on the front,
the median frontal carina not nearly reaching the anterior ocelitu,
the Utera) carinae curved, not straight, and meeting above the
anterior ocelloB. Fronotum rather long, narrowed anteriorly,
■parsely punctured, produced posteriorly into a low tubercle, with
a median eulciis reaching to the base of the tubercle, the anterior
half of the buIcub with a few short transverse striae. Meeonotnm
and Boutellum Hmooth, with a few scattered punctures. Median
segment broader in the middle than long, the third carina in the
middle as far from the fourth aa from the second; the tnberclefl
at the apical angles elongate triangular, not curved. Second
dorsal segment at least one-quarter longer than the breadth in the
middle, the sides not strongly convex; the three apical eegmente
very strongly compressed laterally. Hind tibiae very sparsely
punctured; fifth joint of the tarsi inserted at the base of the
fourth, the latter not reaching the middle of the fifth joint. Length
of the radius beyond the third cubital nervure less than half of
the length of the second transverse cubital nervure. Three cubital
ceDs.
BtA. Nyasaland, Kondowi, Lower Nyika (R. Craw-
ahay), June.
lit the compression of the apical segments of the abdo-
men thia species resembles A. sUnrica, Fabr. {compressi-
veiUris, Gu4r.) ; but in neuration ia close to aaaimilia, Eohl.,
which, however, differs much in sculpture and other
points.
Ampuht krlsteiiMiil, sp. n.
$. Viridi-cyanea; olypeo antenaisqnenigris; tarais fusois; man-
dibniis, tibiis antiois, femoribusque ferrugineis ; alis byalinia, venia
nigris, cellula radiali leviter infnscata.
Long. 16 mm.
3- Clypeus strongly convex, porrect, the carina not produced
into a tooth at the apex. Second joint of the flagellum as long as
the third and fourth combined; eyes separated on the vertex by
a distance scarcely exceeding the lei^h of the second joint of the
flagellum. Head mgosely punctured, the median frontal carina
ver; short, not nearly reaching the anterior ocellus, the lateral
frontal carinae well developed. Prbnotum shining, feebly and
im^ulariy tiansvetaely striated, with a median sulcus on the
aDt«rior portion, raised inte a tubercle at the apex. Mesonotnm
TRANS. ENT. SOC. LORD. 1917. — PART I. (NOV.)
u,y,i,Ajh,Coot5lc
82 Mr. R. E. Timier on Not Species ^
in the middle and •coteUnm with a few BcAttcred pnnotmca, the
wdee of the meeoDotnm more cloedj ponctared. Medun aegment
as in eoHigera, KohL, but with the taberclea at the apcal an^
much amaller and leaB cnrred. Second doraal segmmt much
broader than long, ahorter than in eotiigtn; second vecti^ aeg-
ra&tt convex at the baae aa in amigtra. Fifth joint of the Um
inH!rt«d cloee to the baae of the fourth joint. Three oabital oeUs,
the length of the radius beyond the junction of the thiid transvtfse
cnbital nermre equal to the length of the secood transveiae cubital
nervure.
S<d). Abyssinia, Uarar {Kristensen).
Very nearly allied to conigera. Kohl., but differs, as
pointed out in the description, also in the much greater
length of the radius beyond the third cubital cell, and in
the narrower apical portion of the dypeus and shorter
frontal carina.
SuhfamUy STIZINAK.
Stizu* jwolfleui, sp. Q.
$. Nigra ; labro, dypeo macnla magna utiinqne, oibitis interton-
bna, scapo subtus, flagello subtus, callii humeralibua, meaonoto
angulia posticia, scutello macula utrinque, postacntello taacia
angusta inteinipta, 8^;mento meditmo angolia posticis. s^menlii
dorsalibua 1~4 mm^iU transversa apicali utrinque, 2-6 atriga
tiansverea apicsli in medio, tibiis anticis, intermediis subtua, t«r-
aisque autiois subtus flavis; tibiis postiais torsisque ferrugineisi
mandibulis fusco-ferrugineis ; alis hyalinis, leviter infuscatis, veiui
fusoO'feiTugineis; segmento mediano angulis lateralibus fortiter
excisia.
Long. 10 mm.
$. Eyes separated at the base of the clypeus by a distance haU
aa gKat again as the length of the scape. Head and thorfkx sub-
opaque, very closely and minutely punctured and covered with
very short fuscous pubescence, which is closer on the meaonotum
than on the head i median segment much more strongly punctured,
thinly covered with short whitish pubescence ; abdomen rather Ice*
strongly punctured than the median segment, with faint blue tints
in certain lights, sixtii dorsal segment very closely punctured.
Second cubital cell not petiolate, the first abscissa of the radius at
least three times as long as the second.
Hab. Fiji, Natova, Nadi (R. Veiuh), October.
This belongs to the group of S. Irtdena, Fabr., but
Ij.y.l.Ajl^yCOO'^lL'
BymmopUra in the Bntuh Museum. 8S
differs in the colouring, especially of the legs, from the
allied speciea. The infuacatioo of the wings is caused by
numerous small hairs.
Subfamily CSABBONISAB.
Entomognalhiu msoiisslmu*, sp. n.
?. Nigra; abdomine pedibusqne femgineis; coxia aotiois, tro-
chuilflribtu antioia, femoribnaqoe antioia rabtus nigiu; tegoli*
tMtaceii; alis faaow, venis nigrU; clypeo orgenteo-pabeMente.
Long. 8 mm.
$. Handiblea deeply eioised on the oater margin, simple at the
apex. Clypeiu raised into a poirect toberole jnat above the middle
of the ajHcal margin, with a very short blunt tooth on each aids
near the apical angles. Eyes hairy, the faeets in front rather
larger than elsewhere. Mead Urge, but not broader than tbe
thorax ; front bioad, the eyes aeparat«d at the beae of tbe dypene
by a diatance equal to tbe length of the scape ; antennae short
and stoat, the third Joint of the flagellnm as Itmg as the second;
ocellar region and vertex veiy coarsely punctured ; ocelli in a very
broad triangle, the poeterior pair far apart, bat nearly as far from
tbe eyes aa from each other, an irr^olu obliqne groove from each
posterior oeellns reaching almost to tbe eye. Fronotum trann-
vene, roonded at tbe anglea, punctured; meeonotum coarsely
pnnctaFed-nigoae ; scntellnm with apatae large pnncturea; meao-
grfenrae coarsely, bat rather sparsely, punctured, with a smooth
groove in front for the reception of tbe anterior femora. Median
segment very short, the dorsal surface only half as long as the
scatellum; Uie basal area very broadly rounded at tbe apex,
with aeveral strong longitudinal oarinae; tbe posterior slope
abrupt, almost smooth, with a broad shallow median groove narrow-
ing towards tbe apex. Abdomen seaeile, tbe basiJ segment the
Imwdeet, the two baaal dorsal segments with spane, but rather
large, puncturea; the remaining segments with small scattered
punctores ; sixth dorsal segment rather narrowly triangular, clothed
with golden pubeaoence. Hind tibiae hairy, almost imarmed, with
feeble aeirationa near the apex; fore tani unarmed. Recurrent
nervure received at three-qoartera from the baae of the cubital cell;
radial cell broadly truncate at the apex, the laansverse cubiUl
nemue received at the middle of the radial cell.
Bab. Ntasaland, Mlange {S. A. Neave), January,
This is very distinct in colour and sculpture from other
species of the' genus, but does not differ much in structural
cnaracteis.
u,y,i,Ajh,Cooglc
84 Mr. R, E, Turner on Neio Species <^ Hymenoptera.
Cnbro veltchi, sp. n.
$. Nigra; flogello, pronoto, callis hnmeralibus, scutollo angulia
tmticis, postecutetloque fa«cia transverBft flit vis; mandibulia &pioe
exoepto, tegulie, segmentk donalibus secundo dimidio basali, aez-
toque, TeDtralibuB primo, secundo dimidio basali, quinto, saztoque,
pedibusque feirogineis ; alia hyalinia, leriter inf uacatis, Tenia foscis.
Long. 11 mm.
$. Mfuidiblee tridentat« at the apex; clypeus porreot, triaoga-
larly emarginate at the apex, not carinata, clothed with ailver
pnbescence. Ejres with veiy taige facets in front, separated at
the base of the clifpena bj a distance equal to the length of the
second joint of l^ flagellnm, a groove along the inner margin of
the eyes near the summit. Second joint of the flagellum neariy
u long aa the first and third combined. Head large and mssuve,
broader than the thorax, minntelj and closely ponctured, the
temples vei; broad and clothed with silver pubescence; ooelU in
a wide triangle, the posterior pair as far from each other aa front
the eyes, fully half as far again from the posterior margin of
the head as from each other. Pronotum transverse, rounded at the
anterior angles; mesonotum very finely pnnctnred-ruguloBe on the
anterior half, more sparsely punctured posteriorly, » broad depres-
sion from the anterior margin to the middle. Scutellum and post-
Bcnlellum shining, with scattered jnmctures ; propleurae vertically
striated; mesopleorae sparsely and rather coarsely punctured,
without a groove for the intermediate or hind femora. Median
segment closely and rather strongly punctured, the triangular
basal area defined by grooves and with a median longitudinal
groove, the sides of the segment closely and rather strongly obliquely
striat«d. Abdomen petiolate, smooth and shining, the first seg-
ment longer than the second and third combined, slender, the
apical half moderately swollen, not constricted at the apex; second
segment nearly as broad at the apex as the third, the second ventral
segment with a amall, oblong, pubescent, opaque mark on each side
near the base; fifth dorsal segment minutely punctured; pygidial
area very narrow, lanceolat«. Recurrent nervnre received just
beyond three-quarters from the apex of the cubital odi ; transverse
cubital nervure received close to the middle of the radial cell.
Hind tibiae serrat«.
Hah. Fiji, Natova (fi. Veitch), April.
The petiolate abdomen gives this inaect somewhat the
appeatance of a Das'jprocltis, but it does not belong to
that genus, but does not seem to be closely allied to any
described Crabro.
l„y,l,Ajh,COOl^lC
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
( 85 )
m. NeiB or liille-known Heterocera/roni Madagascar, By
Sir Geo. H. Keneick, F.E.S.
[Read May 2iid, 1917.]
Plates I-VI.
In continuation of a paper read before the Society on
November 5th, 1913, I now submit a further list of insects
taken in Madagascar by Mr. Felix B. Pratt in 1910,
There still remain the Geometridae to be described, and
among them there appear to be many interesting forms,
I have again to thank Sir George Hampson and
his assistants for their very kind help in the work of
identification .
I wish that it had been possible to compare the insects
with types to be found in Continental cabinets before pub-
lishing these descriptions, but as that is out of the question
for the present, I let these go out as they are.
NOTODONTIDAE.
SUvropos malgasstoa, sp. n. Plate I.
He&d, thorax, bue and shaft of antennae, apper part of palpi
wbit« ; base of palpi and fringes round ejea dark brown ; antennae
cheetnnt; legs white, tani blackish; abdomen, baae and last two
aegments white, remainder grey. Fort-wing white; an oblique
basal black line from coBta to cell; followed by an anlemedian
black line, obliqne with strong angulation in the cell, and a thicken-
ing at inner moi^in, which in the female develops into a ronnd
■pot; the median line is faint and not visible beyond the middle
of the cell; the postmedian line has three distinct angulations
aod the enblerminal is composed of 6 V-shaped marks; there are
4 dark dots on the costa beyond the postmedian line. Hind-tcing
white, slightly fuscous towards the inner maj^, with a dark
anbt«nninal line- Exp. SO mm.
Stanropui lllaolna, sp. n. Plate I.
Head, thorax, and legs pale grey; tibiae darker, antennae dark
brown, abdomen grey, the two last segments lighter. Fort-wing
TRAKS. ENT. SOC. LOND. 1917. — PAKT I. (NOV.)
u„,,i,A J 1^, Google
86 Sir Geo. H. Eenrick m
pale gray Bufiuaed with lilac vitb dark brown linea; baaal line
oblique and difiiued, aiitcmediaii line obliqne and angnlated:
median line faint and int«iiupted; postmedian double, tuignlated.
and not vei; obliqne ; in the female with a round spot jiut before
the inner margin ; eabtenninal line compoeed of 6 V-ahaped mark-
ingB ; a more or lees pronoonced dark shade beyond the antemedian
line, most conspicuous towarda coeta. Bind-mng dark grey, fringe*
pale. Exp. SO mm.
Soalmleaada obliqnllSaciats, sp. n. Plate I.
Head and legs reddish brown, palpi paler, antennae and collar
dark brown; thorax above pinkish; abdomen dark at the basB.
then ochreouB, but tuft blackish. Fore-wing pinkish brown, with
a brownish cloud, more or less intense, from cell to costa; a fine
double line from middle of inner margin to apex; an irxegolar
dark brown transverse line at base, reaching half acroM wing,
followed by an angulated anteniedian tine; the 3 stigmata are of
the ground-colour, but outlined by fine dark lines fringes dark.
Hind-wing pale straw-colour with fringes conoolorous. Exp.
Varies a good deal in intensity of colour, and the stig-
mata in some specimens are more or less black.
Scahnieaada ochteoploU, sp. n. Plate II.
Somewhat like the foregoing, but has no obliqne line; the stig-
mata are barely discernible, and in most specimens the ana at
base of wing and at the angle is spotted with pale ocbrcons. Hind-
vAng much darker than in the preceding species. Exp. 40 mm.
Scalmleanda pratti, sp. n. Plate II.
Head, legs, palpi, and thorax reddish brown, shaft of antennae
pale; abdomen reddish brown, paler beneath, tuft of the same
colour. Fore-wing reddish brown, mottled with pale ochreons,
which forms a patch e;(tending from cell to near apex; a black
spot at base of wing, and a whitish horizontal line extending from
base to near middle of inner margin, where it meets an irregular
transverse pale line extending to costa; a black streak at end tA
cell, and a pale postmedian waved hne with dark border; 8 black
dotfl form a subtermlnal line, and a pale line precedes the maigin.
Hind-wing reddish oohieoua. Exp. 34 mm.
l„y,|,AJI^,COOl^lC
I
: I
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
NttD or lU^hnown Beterocera from Madagtucar. 87
NOCTUIDAE.
TTMhu tTluigabtta, sp. n. Plate II.
Head, lag^ P^lpu ^nd thor&z pi^'l'"'' oohreous, « pole line at
baaa erf collar; abdomai ochiMHis at bsw and Uien pinkish, fore-
whtg pale pinkiah brown, with a CMupioiKnis triangle having its
ftpex on inner mwgiD, and of a dark cheetnnt coloor; inside the
base of the trian^e ia a semioirciilar bond of the ground-colour.
The triangular marking is edged with a fine [riak line, and there is
a straight snbterminal line. Eind-wing whitish oohreous, hinges
oi both wingH pinkiah. Exp. 30 mm.
DueochMto nulgiulea, sp. n. FUte IV.
Bead, legs, palpi, and thorax apple-green with blaok maridngs,
antennae pale brown ; abdomen blackish with pale green tuf ta, the
last s^ment bright gcten. Fort-viing apple-green, with 7 oi 8 black
spote on coBta, a black basal patch and ontemedian, median, and
postmedian interrupted black lines, the latter connected above
die inner margin b; a black dash to the subtemunal line, which is
ontwardfy bordered by a eilrerj white line; there are also black
daahes dong the inner margin, and a row of black dots on the
hind margin. Hind-mttg uniformly smoky with two tiansverse
daricer striae, fringes paler. Exp. 42 mm.
Hnopwlgei mlnuMuli, sp. □. Pi&te I.
Head, l^s, and palpi ochreous with a pink tinge, antennae dark
brown; thorax ochreous with scattered blaok scales; abdomen
ochreons with a smoky tinge. Fore-wing dark brown with oohreous
spots, the whole suffused with a faint purple gloss; a dark basal
line is followed by two ochreous blotches and then an antemedian
line; the postmedian line is parallel with tJte hind margin, and
between them are tiiree ochreous blotches, besides a small one at
end of cell ; both inner and hind margins have ochreous patches on
them. Hind-wing uniformly smoky, fringes paler. Exp. 36 mm.
Hy^per^ea variegata, sp. n. Plate I.
Head and palpi reddish oohreons; antennae pale brown; legs
ochrEons with dark rings on tarai; thorax whitish above, patagia
dark; abdomen dark ochreous paler between segmentfl. Fort-
wtnjr daik yellowish grey, with pale ochreous blotches; a whitish
spot at base, with indications of a dark basal line ; an interrupted
black antemedian line, and a curved and angulated postmedian
line; the stigmata oatlined with black, and the reniform filled with
iM-,i,A J h, Google
88 Sir Geo. H. KeDiick on
white uid oontinaed beyond, making a large round spot. There
are two pole blotohes on the inner margin, and two on the hind
nuugin, with a dark Btieab between them. BtTul-mitg very pale
with dtukf margin, fringee pale. Exp. 40 mm.
Perlgea rubidita, gp. n. Plate I.
Head, aQtonnae, palpi, and tegs reddish, the tarsi ringed with
darker, collar dark with whitish crest; thorax and patagia dark
with whitiah scsJes; abdomen reddish brown with darker creeta
and jellowish tuft. Fore-wing dark brown suSuscd with pink,
coeta broadly covered with whitish scales; a conspicuous curved
central streak from base to costa very dark, and a dark spot on
coBta beyond end of cell. The stigmata are pale, but the ctavi-
form has a dark streak in it ; there is an indistinct pale Bubtermlnal
line, and some dark streaks before the hind margin. Hind-wing
reddish brown. Exp. 42 mm.
Perlgea grlsetta, sp. n. Plate II.
Head, antennae, palpi, and legs reddish, the tarsi ringed with
darker; collar dark brown, patagia pale ochreous; abdomen
yellowish ochreous, crcsla not developed. Fore-iving whitish
ochreous, the stigmata, which ore large, outlined with black, with
a block space between them ; an antemedian line starting from
near base on costa curves outward to inner margin; a postmedian
line is nearly straight, the space between these is filled in with
dark blackish- brown ; there is a wared and interrupted subterminal
line, and some dark streaks near it. Hind-wing unifonnly oohreons,
fringes paler. Exp. 38 mm.
BfaoroMmyra robusta, sp. n. Plate II.
Near to Jf. ttatbrota, Butl., but the wings are aquarer, and the
insect decidedly stouter. Head, legs, palpi, and anl«nnae brown;
thorax and abdomen darker and shaggy. Fore-uring with sealea
slightly raised, giving them a rough appearance, dark brown, but
the coeta bright brown, especially underneath; indications of the
beginning of a dark antemedian line on costa, and a stronger dark
Bubterminat line begins on the coata before the apex and is con-
tinued to vein 4. Hind-wing unifonnly dusky, fringes rather
paler. Exp. 36 mm. One S-
BoroUa carneatlneta, sp. u. Plate I.
Antennae of male ciliated. Head, palpi, legs, and collar yellowish-
oohreous, antennae darker; pats^a ochreous, abdomen smoky.
ih,Cooi^lc
New or little-knoum Heteracera from Madagatcar. 89
toft ochiGOoa. Fon-wing yellowish olive snffiued with {^"t
divided longitndiiuUly into three folds, the coeta aod edge of aeoond
fold pale jrink ; indicsticnu of a dark Gist line ; a triaDgalar olive
blot«h midway along cOBta, a blsck dot on vein 5 forming part of
« postmedisn line ; the hind margin is broadly eoffiued with fosoons,
and there are a few black enbtenninal pointa. Bind-wiag greyish
with faint dark hmnle, fringes paler. Exp. 30 mm.
Borolla pkrvula, sp. n. Plate 11.
Antennae of male nearly umple. Head, palpi, le^e, antennae
and thorax reddish ochreoos, abdomen rather darker. Fore-wing
reddish ochieous, with veins paler, but rather more colour in tbe
tsell ; a short dark streak from the base under the cell, two or three
dark specks along the costa indicating a postmedian line, a small
black dot below the middle of the cell, and another more oonspiouoas
two-thirds along the inner margin from the base. Hind-wing 4rf
male pale ochreoua, of female pale grey. £xp. 26 mm.
Boroli* lithargyroides, sp. n. Plate II.
Head, thorax, palpi, antannae, and legs pinkish oohreous, abdo-
men less pink. Fore-tcing pinkish ochreous, with a centi^ shade
and a triangolar shade below apex; on the central shade at the
«nd of tbe cell is a short white longitudinal streak, and above this
a black dot; both antemedian and postmedian lines aie repre-
sented by a tew black dots, and the subterminal line by a complete
row. Biwi-wing grey, fringes pinkish. Exp. 28 mm. One 2-
Aeosmetlft nulgassiea, sp. n. Plate II.
Head, legs, antennae and palpi grej^sh brown; thorax grey, the
collar divided into two prominences; abdomen greyish oohreous.
Fore-tcing dark grey with a purplish gloss; indications of ante-
and post-median lines, subterminal line paler. Bind-miig pale
grey. Underside whitish, thickly iriorated with grey. Exp.
28 mm.
Parscaroldes pratti, sp. n. Plate I.
Head, antennae, palpi, and legs chestnut; thorax duker, hut
with patagja paler; abdomen yellowish ochreous. Fon-wing: tbe
disc is apaisely coveted with scales, while the margins have a thicker
coat BO as to appear decidedly raised; pale ochreous, darker to-
' wards margin, tbe oosta chestnut, with a few white scales ; the
. inner and bind matguis have a pale band, followed by one nearly
iblack; thereaie two small dark dote at end of cell; fringes purplish.
iM-,i,Ajh,Googlc
90 Sir Geo. H. Kenrick on
Bind-wing ochreom, darker round tbe margiDB, fringea pale. Exp.
38 mm.
TnehM kueopieto, sp. n. Plate II.
This species falls among the section in which the males
have lateral tufts of hair on the abdomen.
Heiuj, palpi, and lege black, vitik whit« spots, anUmnae ohest-
nnt; thorax black with a donal white streak, and white on the
patagia and back of collar ; abdomen ochreona, inclined to foscons
above, the crests darker, tipped with white. Fort-mni/ black,
with white mai^tings, the median area parttj' suffused with lilac,
an irregular three-lobed white blotch at base ; the ant«median line
white, angnlat«d, and numing into a square white spot on inner
margin; the orbicular and reniform stigmata black with whit«
margins. A large white blotch fills the remainder of the wing
beyond the postmedian line, but it is interrupted by a square patch
of black at the apez, with two white spots in it, and by an irregular
black pateh at anal angle which is connected to the central area.
Hind-wing fuscous, but whitish at anal angle. Exp. 40 mm.
Elaeodes proteoldes, sp. n. Plate I.
Head, palpi, antennae, and collar, pale pinkish ochreoua; li^
the same colour, with tarsi ringed with darker, the long bain on
tibiae paler. Fort-icing greenish with purple blotchea; ante-
median and poetmedian lines pale with darker maigins; reniform
and orbicular stigmata with some pink and darker markings, ont-
lined by a pate line; e. row of subterminal black dashes, fringes
spotted. Hind-wing brownish with pale fringe, and paler at tbe
base ; a faintly darker lunule and on the underside a dark stiia.
Exp. 32 mm.
Rjpooalymnla glorlosa, sp. d. Plate IV.
The venation of tbe hind-wing does not agree with HampooD's
figure, p. 186, in this insect there is a distinct vein 6 from just
above the lower angle of cell. Head and thorax greyish ochre-
ODS, legs, antennae, and palpi reddish ochreous. Fore-wing coppeiy
shot with purple; an angulated pale stripe starting from thorai
below costa reaches to inner margin, and turns at the bind margin,
gradually becoming narrower; it is bordered on the outside by a
chestnut semiciroular line; the orbicular and reniform stigmata
are of the some colour, and towards the hind margin on the disc
are two small dark dashea. Hind-wing and fringes pale ochreous.
Exp. 40 mm.
Ij.y.l.AjL.yCOO'^lL'
Nae or liiUt-kniAen Seterocera Jrom Madagatcar. 91
Stietoplen peetlinta, sp. d. Plate IV.
If this insect does not constitute a distinct genns it
shotild be placed in a separate division on account of the
strongly pectinate antennae of the male, and the square-
shaped wings; in other respects it follows the genus.
Be^ and temiiiial joint of palpi ochreom ; l^s, Antennae and
second joint of palpi nther d&rker ; collar reddifib ochreoiu, pstagik
puridiBh brown; abdomm dark giey witb pale tuft. fon-uii»g
dark marbled with grey, green, pnrple and brown ; baaal line dark,
nwrgined witb pater extending to oell; antemedian obacarelf
double, greenisb; metiiBn line sbarp and dork, but not extending
to inner margin ; poetmedian line dark, angulated, and inteiinpt«d,
followed by two brown blotches and a brown subterminal line.
Bind-wing dark grey, aubbyaline at base. What I take to be tile
female has plain antennae, and the patagia and baaal half of Ion-
wing, t<^lfae( witb a postmedian area pale grey. Exp. 31 mm.
Eutetla anreo^eta, ^p. □. Plate IV
Head, antennae, legs, and palpi pinkish grey; thorax and creeta
of abdomen coppery bronze, abdomen pinkish grey. Fort-wing
bronze, with grey lines and chocolate blotches; basal line ill de-
fined, followed by three chocolate patches; an antemedian angu-
lated whitish line ; the orbicular, a darii dot outlined with white ;
the renifonu large, oblique, outlined with white, with a dark edging
of chocolate on the inner aide ; three ill-defined chocolate patches
on the hind margin, and tiocea of a double white subterminal tine;
beiore the middle patch ifl an oblique golden spot, followed by a
minute dot of same colour. Hind-wing whitish, with a broad
fuscous margin. Underude giey, with reddish suffusion towards
the margins, and a white spot on coata before the apex. Exp.
EuteUa oehreoplagata, sp. n. Plate II.
Head, 1^, antennae, and palpi dark fuscous ; abdomen without
cTWta, dark fuscous. Fort-wing purplish grey, with lines much
daiker; basal line broad, antemedian line double, one portion
joining with central line on inner margin; the postmedian line
BugulBt«d below costa, and preceded by a slightly paler patch;
beyond this the apex becomes abruptly pale ochreous, and pene-
trating half of this patch is the subterminal line, consisting of six
dark brown connected spots. Hind-wing purplish grey, inclining to
ochreous at base, with three angulated transverse lines, the outer-
iM:,i,A J 1^, Google
92 Sir Geo. H. Eenrick on
most dark brown, frsngea dork. Undenide foaootu, pale at the
base, the apioal patch dark yellow. Exp. 33 mm.
Phlegetonla pratti, sp. n. Plate III.
Near to artipars, but the male has the tibiae very heavily
fringed with hairs, and the markings are somewhat different.
Head, palpi, U^ and thorax entirely pale ochreout, aatemiae
reddish ocbreoiu; abdomen oohreoua with darker cieots. Fort-
wing pale ochreons, with a faint purplish gloaa and dark bbtches ;
baoal line indicated by a dark dot on the coata, the antemedian is
a dark and ainuoua line in the middle of a brown transverse blotch ;
there is a dark central spot, and a fine indistinct median line, followed
by two others nearly parallel ; the postmedian is a fine dark line,
inwardly angled below cell and on vein 2; on its umer side near
the costa is a pale blotch, surrounded by a dark triangolar patch
on costa in which are three pale costal data; the subterminal line
is fine and black, and there is a ronnd dark dot near the angle on
inner margin. In ^e female the whole wing is darker, and the
markings are obscured by a dark grey central band. Hin^-ving
pale oohreona, with numerous striae and a double dark line from
anal angle to middle of wing, Bubt«rminal line very fine and black.
Exp. 40 mm.
Blenina hyblaeoides, ap. n. Plate III.
Palpi with second and third joint well developed; tennen of
hind-wing very slightly excised. Head, palpi, thorax and 1^
greenish grey, tarsi ringed with black, antennae dark; abdomen
blackish above, with orange margins to the segments, ochreous '
below. Fore-iinng greenish grey with faint pink ceflections; first
line black, joining a black basal streak; antemedian line oblique
and angulated ; three blaok dots at end of cell, followed by a much
angulated postmedian black line, edged outwardly with paler;
beyond this are some irregular dark brown tooth-shaped markings
forming a subterminal line. Hind-wing orange, with broad fuscous
margin and orange fringes ; in one specimen there is a wide black
transverse band quite absent in the other. Underside fore-wing
fuscoue, with paler costal band, interrupted by dark central spot.
Bind-wirtg orange with dark marginal band. Exp. 33 mm.
Chlorozada purpurea, sp. n. Plate IV.
The position of vein 7 in the fore-wing differs from the
type in leaving vein 8 beyond the end of the cell, but this
is hardly sufficient for the creation of a special genus.
u„,,i,A J 1^, Google
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
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i
n,g,t,7.dh, Google
Nete or lit^knowm Heterocera Jrom Madagascar. 93
Anteniue and palpi brown, head and patagiA pale alive; legs
while, abdomen and thorax porplish grey, paler below. Fore-
wing whitish, with a purple cloud occupying the centre of the wing,
and leaving a narrow pale external margin ; traces of a dark basal
blotch edged with while externally; a sinnous fine white ante-
median line edged with darker; two dark dots in the cenlral area,
followed by an oblique fine white angulated poetmedian line; the
mbterminal line takes the form of six dark dots on the pale apical
aiea; there ia a dark terminal line extending half way down the
wing ; fringes pink. Hind-mttg uniformly giey, with pale fringes.
Exp. 26 mm.
Polydesnu reotefueUta, sp. n. Plate IV.
Head and thorax fawn-colour; antennae, legs, and palpi dark
brown with numeroas paJe grey hairs; abdomen fawn-colour above,
the crests darker, paler below. Fore-wing hind-margin aubangular,
fawn-colour; anl«mediaa line whitish edged with darker, much
angulated and interrupted; stigmata repreeenled by three minute
dots, dark, edged with paler; from jnst before the angle to the
oosta there is a group of fine lines, nearly straight; the first is
white, the second dark, the third pink, the fourth wider and dark
brown, the fifth pale oohreous, forked at the top and expanding
into an apit^l patch in which are two dark dote; beyond these
lines there is a broad pink area. Bind-wing yellowish, broadly
margined with fusoous, but again yellowish at the outer margin;
» few whitish dots at the angle. Underside uniformly whitish
diiokly irrorat«d with black, a black transverae stria and central
dot in both winga. Exp. 42 mm.
Polydesma tesselUta, sp. a. Plate IV.
Antennae, bead, and thorax dark brown, palpi brown outside
and ocfareooe inside; abdomen dark brown above, ochreoua be-
neath, legs ochreouB. Fore-wtTig brown with violet shade, the
coata with a seriea of dark brown spots; a baeat patch of ochreoua
jrroraled with black, followed by aD,iiTegular double violet line; an
indistinct daric median line and a strongly marked dark poHtmedian
line; Aeroniform stigma outlined with a pale line, and with internal
lunule; the anbtenninal line much angulated, double, and whitish,
followed by a series of dark marks outlined with paler, and a thin
terminal pale line, fringes rather paler. Hind-tinag umilar to fore-
wing, the pattern of the outer portion forming a continuation of the
pattern of the fiwe-wing. Underside ochreous, with lunules, and a
u,y,l,/eJh,G00glc
94 Sir Geo. H. Kenrick on
enrred poatmedian line in both wings; a row of black doU along
tha maqpn. Exp. SO nun.
Builuu eaUastnths. Plate IV,
After describing and figuring this insect, it was identified
with Baniana caUaxcaniha, Roll., but the figure could not
be withdrawn.
Ptaytometra pratti, sp. n. Plate III.
Head, antennae, palpi, and legs smoky grey, thorax and abdo-
men rather paler. Fore-mng dark grej, with coppery reflecttona
on the disc. Neither lines nor stigmata are visible, hut there is n
fine silveiy line from inner margin near the base extending obliquely
upwards to the cell, where it turns outward to fonn a silvery oblong
spot, in continuation of which, but quit« distinct is a second silvei;
oval spot. Hind-wing ochreous-grey. Exp. 40 mm.
Nlgranuna malgassloa, sp. n. Plate IV.
Head, antennae, palpi, and legs, reddish ochreous; tegulae
oohreous, with two darker transvene bands, patagia dark brown ;
abdomen reddish grey. Fort-mng pale ochreous clouded with
reddish; a oonspicnoua oval basal blotch, edged with paler and not
reaching either to the costa or inner margin ; there is an antemedian
darker fine line, not very distinct, and a similai poetmedian line and
three dark dots at end of cell; on the inner margin theie is a pale dot
near the anal angle, and at the angle another pale dot, from irtiioh
extends upwards a pale subterminal line ; on either aide of this is
a row of black dots. Hind-wing uniformly dark grey, fringes paler.
Exp. 34 mm.
Corgatha sublndlcata, ap. n. Plate III.
Head and thorax pale ochreous, palpi, antennae, and legs reddish
brown; abdomen pale at the base, but darker on the remaining
segments. Fore-wing pale oohreous, clouded with brownish grey;
antemedian line whitish, slightly angulated at cell, and edged with
darker; central shade oblique, straight, and brown, reniform
stigma indicated by a white maigin; poatmedian white, angled
outwards below costa and oblique, terminating in the middle of
inner margin; a fine interrupted snbtenninal line extends front
bdow vein 2 to vein 6 ; beyond this is a distinct pale patch, followed
by an uignlated brown marginal line. Hind-wing pale ochreon^
with two oblique dark lines, the outer one maigined with white.
l„y,l,Ajh,COOl^lC
New or little-known Heterocera Jrom Madagaecar. 95
And bejond this an & few dark dote, fringaa pale brown. Tha
nndenide of both wings mnoh darker than the upper, and the
hind-wing bee a well-defined daA margin. The angle of the hind-
wing ia Btiongly orenulated, poaaiblj a eexual deTelopment. Exp.
»nim.
Corgfttha thyridoides, sp. n. Plate III.
Head, palpi, anUmnae, and thorax leddiah oohreoua; legs reddiab
ocbreone with the tarsi paler; abdomen reddiah above, paler beneath.
Fore-iping pale ochreouB with reddish trrorations; baaal line darker,
extending from ooeta to middle of cell; a dark angulated ante-
median line, followed by a dark central dot, and this again by an
outwardly angled median dark line, bejond which the reniform
stigma is outlined with darker; the postmedian line ia etrongly
angled outwards at coata, and then continued obliquely to middle
of inner margin ; beyond this is a diffused reddish band, after which
tbe wing is paler, with the snbt«rminal line indicated by three black
dots : in the apex there is an indistinct mai^al row of dots. Hind-
winy dull red, slightly ochreous at the base, with two transverse
dark lines beyond which is a paler patch with some grey in it ; the
anal angle of the wing is strongly crenulate, as in vubindieata.
Underside paler, with similar markings. Exp. 28 mm.
This insect ia figiireH by >SaalmulIer. but neither named
nor described.
OEUba vlridaria, sp. n. Plate III.
Head and thorax ochreous, with a faint reddish tinge ; antennae,
l^s, and palpi ratiker darker; abdomen pinkish ochreous. Fore-
wing dark grey, marbled with green with whitish linee ; basal line
indicated by dark dots on costa; an outwardly curved whitish
antetnedian line; the reniform stigma pale margined with darker;
tlie postmedian line whitish, strongly outwardly curved at costa
aad margined on both sides with darker; beyond this the wing is
whitish ochreous, with a half round dark spot on costa before apex
and a grey cloud, with a few black t«nninal dots, before the hind
margin- Hind-win^ uniformly grey with paler fringes. Exp.
Goaepleronla Upunetata, sp. u. Plate III.
Head, palpi, antennae, and legs dull reddish ochreous, tarai
ringed with white. Fort-tping dull reddish, minutely itTorat«d
with ochreous; lines darker, antemedian angulated, sloping out-
wards from ooMa, median line nearly vertical, not reaching above
iM:,i,A J 1^, Google
96 Sir Geo. H. Eeurick on
reniform stigma; poBtmedian line much angnlated from coeta to
vein 3, then turning inwarda abruptly and meeting the median
line; eabterminal line faintly indicatod, pale ochreouB and much
angnlated ; the orbicular stigma a minute white dot, and a aimilar
white dot at the top of the reniform atigma ; fringes reddish. Bind-
wing uniformly greyish ochreona, fringes paler. Exp. 43 mm.
Xanthodesm» notugulita, sp. n. Plate III.
Head, antennae, palpi, legs, and thorax ochreous-orange, abdo-
men dull yellowish. Fort-uing dull orange, the veins all dark
brown ; a short dark basal line, a straight antemedian line sloping
outward from costa, a postmedian line, strongly angnlated on vein 6 ;
from the middle of this a short line extends to the reniform atigma;
the Bubterminal line is curved, and is followed by a distinct ter-
minal line. Hind-wing uniformly dull yellow; fringes of fore-wing,
orange of hind-wing dull yellow. Underside uniformly dull yellow.
Exp. 44 mm.
DELTOID AE.
Singari albonuoiUs, sp. n. Plate III.
Palpi well above vertex, second joint thickly scaled, third joint
bare and sharp; antennae bipectinated, tibiae spurred; veins 3
and 4 from end of cell, S from just above, 6 and 7 from upper end
of oell ; costa nearly straight, hind margin rounded. Head, thorax,
and patagia orange; palpi orange outside and paler inside; legs
oohreous, antennae pale brown. Fore-wing orange irroratod with
darker; a basal dark pateh, followed by an antemedian line curved
inwards, a dark ohbque streak from apex to middle of inner margin;
the orbicular a conspicuous white dot with dark margin ; the reni-
form also white, with dark margin and dark centre. Hivd-mng
ochreouB, the outer portion inclined te orange; a narrow median
line extending te vein 3, fringes narrow, orange. Underude ochte-
ODS, the fore-wing with three dark spots, and the hind-wing with
dark central lunule. Exp. 40 mm.
PYRALIDAE.
Acara pratti, sp. n. Plate V.
I cannot distinguish any generic character to separate
this species from the genus, but the hind-wing is sqiiarer
than in morosella, and vein 2 of hind-wing is not present
iM-,i,A J h, Google
New or little'knoam Beterocera from Madagascar. VI
as shown in the Sgure,* but this is not mentioned in the
description.
Head, lege, and tmtemiae reddish brown ; thorax ntber brightor,
psJpi in female darker; abdomen shining ochreoua with leddiah
BUffosion. Fore-udng unifonnly reddish ochreoua, the co«t« clothed
with ochieone scales, and many tranaverw bars of the same. Hind-
wing yellowish oohreous without markings. Exp. <} CO, $ TO mm.
Haoalla malgassloa, sp. n. Plate VI.
Somewhat like a small species from Natal.
Head white, antennae, palpi and legs dark gny; thorax duk,
with collar and patagia grey ; abdomen ochreoua. Fon-^ng pale
gny, with darker markings; an antemediau and portmedian line
indicated Ity dark dots, a darker patch in the c«ll; snbterminal
Hno a series of dark dashes, fringes gKj. Hind-wing ochreotu.
£xp. 24 mm.
Taenuphoba, gen. nov. (Sub-family Epipaschianae).
Palpi porroct, extending twice the length of the head, seoond
joint fringed with hair above and below, third joint long and naked.
Antennae of both sexes pectinated, but simple at the tip; tibiae
with fairly long spines. Fort-mng vein 6 from end of cell, 7, 8,
and B stalked. Hind-wing 3, 4, and 6 from end of cell ; median
nermre not pectinated; no raised scales below the cell on the
npper side, but on the underside of the fore-wing the scales in the
cell are crowded together, and slightly raised ; the longitudinal fold
below the cell in the fore-wing in the m^e has no scales, but a
namber of vertical striae ; in the female the scales are uomiaL
TienUphon tubnurgtaiats, sp. n. Plate V.
^Ipi and antennae pale ochieous; legs the same, with reddish
hairs on the tibiae ; thorax and patagia reddish ochreoua, abdomen
paler. Fore-wing pole ochreous with fiiscoua inner and hind margin ;
at the base of the costa is a blackish shade, followed by two dark
■pots ; fringe oohreoos spotted with reddish ; at tbe end of the cell
IB a conspicuous black dot, and beyond this a series oi fine black
triangular tnarka margined outwardly with paler. Hind-wing pale
ochreous, with a thin black marginal line and pale fringe. Under-
side pale, broadly margined on tbe costa of both wings with reddish,
and with a dork spot at the end of cell in both wings. Exp. SO mm.
* Hampson's Moths of India.
TEAN3. BNT. SOC. LOST). 1917. — PAST I. (NOV.) H
iM-,i,A J h, Google
dS Siz Geo. H. Kenrick on
Slndris leuoomelas, sp. n. Plate VI.
Head, legs, antennae and palpi olive, patagia the same ; thorkx
and abdomen orange. Fore-wing olive enfiiued with orange, and
with three distinct white blotchea, one roundish on hind margio:
one in the middle of the wing, rounded towards the body, bnt
forming a hroad lateral stripe ontwardly; one at apex just joining
the last, and with three pointed teeth on the ontode; fringes
ochreoUB. Hind-wing uniformly golden. Exp. 38 mm.
LoFHOCE&A, geD. nov. (Sub-family Pyralinae).
Palpi aptnmed, the third joint well developed and acuta, pfo-
bowis prraent; onteniute pectinated in the male, with a bunch of
long haiia about the middle ; venation. Fort-wing 3, 1, and fi from
lower end of oell, 6 from upper end, 7, 8, 9, stalked. Hind-ving 6
from end of cell, 7 ansetemoong with ft.
Lophoean flavlpuiieta, sp. n. Plate VI.
Head, legs, palpi, and tboras black, patagia and collar onnge,
abdomen orange, tuft black. Fore-wing black, with a huge yellow
oblique spot beyond the end of the celL Htitd-wing orange with
broad black border. Underside similar. Ezp. S 34 mm., $ 30 mm.
Fllodes albotermlnalls, sp. n. Plate V,
Paljn, head, antennae, and legs oohreous; antennae barely the
length of the fore-wing; patagia of male long and darker; abdo-
men golden above, paler beneath, tuft conspicuously white, ^ore-
wing ochreoua with golden suSuaion; a dark spot at baae of ooeta,
a dark antemedion line curved ontwatds; a conspionooa oblong,
Innulated, oblique dark spot at end of oell ; a postmedian «.iignlf.tjiH
dark line thickeet at costa; beyond this the wing is greyish; in
some specimens a dark dot in the celL Hind-wing with oottxal spot.
Exp. 36 nun.
modes frlsMlls, sp. n. Plate VI.
Head, palpi, antennae, thorax, and abdomen dark grey; lep
and underaide silvery white. Fort-wing dark grey, with alight
tinge of purple; the antemedian line central shade, and poat-
median line more or leas distinctly divide the area into four nearly
equal portions ; in the cell is a dark spot, and a da^er mark at the
end of the cell. Hind-wing with central spot and tranavene lino
l„y,|,AJI^,COOl^lC
n,g,t,7.dh,G00glc
h,Googlc
New or liule-knoton Heterocera from Madagascar. 99
rather indistinct ; these ma.rfcing« are clearer on the onderBide.
Exp. 32 mm. One $.
Boeehorls rectUlimlli, sp. n. Plate V.
Head, antennae, and palpi pole r>chreoua ; pectus and le^ silvery
white ; thorax ochreona with two dark apats on collar ; abdontMi
with a golden tinge, toft paler. Fore-wing ochreous miffiued with
pale lilac, the ooeta paler; an indiatinct dork basal line, an angu-
lated, broad dark antemedian line, a faint dark maHc at end of
cell; a broad dark nearly Htraight oblique poatmedian line, followed
by a lanab fainter thin parallel line. Hind-teing pole with d^rk
central spot, angulated dork line and brood dark border, strongeet
at apex ; a fine Babt«rminal pale line, fringes darker. Exp. 24 mm.
Boedwria tenuilinwlli, sp. n. Plate V.
Head, palpi, and legs pale ocbreouB, antennae pale brown ; thorax
and abdomen whitish. Fort-wing semi-hyaline with golden sheen,
the veins showing dark ; a dark basal spot, a straight obliqtie ante-
median line, the angle formed by it and the inner maigin being
darker, a very faint dot at end of cell, and a fine, dark, evenly
curved poatmedian line. Hind-vnng pale at base, with an irregular
darker Innnle, and a fine curved median line, the veins darker;
fiingea pole golden. Exp. 23 mm.
Sylepta malgsssiea, sp. n. Plate VI.
Head aod antennae ochreous, palpi, flnt joint white, seocmd joint
ochreona : legs pale ochreona and silvery; thorax and abdomen
ochieoiiB. Fort-wing ffey, with gold and purple sheen ; a whitish
blotch at base extending to first line, which is dark; the cell itself
is whitish, with dork spots at either end ; beyond this is a strongly
angulated dark line, followed at costa by a white blotch with two
teeth ; below tbg cell is another white blotch ; at apex is a small
dark streak. Hind-wing whitish, with dark Innule and angulated
line, tile margin widely bordered with fuscons. Exp. 26 mm.
SyhptB aeutlpeniulls, sp. n. Plate VI.
Head doll orange, palpi darker, antennae pale brown; legs
silvery, the tibiae of the median pair of 1^ with a long toft of
yellow hair tipped with darker; thorax, a.bdomen, and patagia
pale brown, toft blackish. Fort-mng pale brawn with buff patches
iM:,i,A J 1^, Google
100 Sii G«o. H. Eenrick on
— 'the two fitst in the cell, the third very amall at end of cell, the
fourth large, extending in triangular form from the oosta to vein 4,
and margined outride hy a dark line. Hind-wing with two faint
striae enclosing a paler space. Exp. 26 mm.
Glyphodes parftmlralis, sp. n. Plate VI.
On comparing this iosect with Swinhoe's description of
amicalis, and the type of this and of vubamicalis in the
. Natural Hiatory Museum, I am convinced that this is a
diflerent species.
Head, antennae, palpi, and legs whitish; thorax and abdomen
fnscOQS brown, collar and patagia dove^colonr. Fore-wing daric
brown sofEused with purple, with two semi-hjaline bonds with
violet refleotions; the first band oblique, and sharply angulated on
the outAide; the second wide on the oosta, and tapering to a pomt
a little below the third vein ; a taint pole dot at end of oelL Hind-
wing similar, with one trioiigalar traosvetHe hyaline band bordered
outside bj a double oblique dark line, beyond which the diso bos a
greenish reflection; fringes paJe, but not white. Exp. 36 mm.
Pyreosta aureoflnctalls. sp. n. Plate Y.
Head, antennae, and palpi pale brown, legs and nnderaide of
thorax white; thorax and abdomen orange, patagia oanajy-yellow.
Fon-wing semi-hyaline with golden reflections; aotemediaa tine
indieated by two dork dots; a dark dot at end of cell, and two
dote nearer to inner margin, the postmedian indicated bj four faint
dote. Hind-wing similar, with a row of dark marginal dote, fringes
golden. Exp. 34 mm.
Pynnita elutalis, sp. n. Plate VI.
Head, antennae, and palpi dark brown, legs paler, and the tani
ringed with dark brown; thorax and patagia greeniih grey; abdo-
men pale brown, tuft pale. Fort-wing greenish grey mottled with
paler; a cnrved antemedian line darker, a ainaoua median line, and
a postmedian angulated line, curving outwards froro the casta but
not reaching the inner margin. Hiiid-mng pole brown without
mukings, fringes paler. Exp. 32 mm.
pyraiuta marglnesorlptalls, sp. n. Plate VI.
Head, palpi, and antennae dull red, legs red above, silvery below ;
thorax and abdomen above dull ted, paler below, tuft pale. Fort-
n,g,i,.eih,.G00glc
^1 >! I
ZV ^
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
JioD or UOle-hwtm Heteroeem from Madagaicar. 101'
temg nniftMrnly doll pink with Inown inoratiotu ; a few black tcalea
oUiqiKlj jdaced iitdicate tbe nntemediaD line, and there is a dark
maifc at end of cell; tbe snbterminsl line is very distinct, cod-
siating of fine black angolaUona. Hind-ioing aemi-hyaline and
whitish, fringes of both wiu^ pale. Expt 34 mm.
n,g,t,7.dh,G00glc
( 102 )
IV. AddUi&ns lo the knowledge of the Cetoniidae o/" British
India. By Ouvbr E. Janson, F.E.S.
[Bead FebnuTy 7tb, IQIT.]
The basis of this contribution is an interesting and well-
preserved series of Indian Cetoniidae collected by the late
Captain B, Selous, and placed in my hands for identification
by his brother, Di. C. F. Selous, who has most generously
allowed me to retain any of the specimens that I desired
for my collection. A Ibt of the species with the collector's
notes that accompany them is given below, followed by a
description of the veiy pretty new Anatona that he bad the
good fortune to discover,
I have taken this opportunity to describe three other
new Indian species that nave recently come into my posses-
sion from other sources, and of giving references to the
additions that have been made to the family since the
publication, in 1910, of the admirable volume by Mr, G, J.
Arrow, on the Cetoniidae of the " Fauna of British India."
I have also added some notes on a few errors and omissions
in that work, and relative to the subject of this contribution.
List of the Cetoniidae collected by the Late
Captain B. Selous.
1. Anthracophora crucifera, Oliv.
Mhow. " On grass, September 1905," one specimen.
A common and widely distributed species, but not
hitherto recorded from Central India.
2. Analona selousi, n. sp.
Mhow, " Found on Nil Gai dung, July 15th, 1905,"
one specimen.
Of this very distinct and pretty new species a single male
example only was found by Capt. Selous,
3. Adhiessa bagdadensis, Burm.
Quetta. " Found flying and on the ground, March lltb,
1907," two specimens.
TSAHS. ENT. 8O0. LOND. 1917. — ^FABT I. (ifOV.)
Ml. Jaosoa'B Addiliims to Cetoniidae of Bnlish India. 103
A rathet scarce species, occurrine in Armenia, Meso-
potamia, Peisia and Afghanistan, and already recorded by
Arrow from Baluchistan.
4. Protaetia (Polosia) impavida, Jans.
Kashmir. " On flowering bush, July 1906," one
specimen.
This species appears to be confined to the northern parts'
of India and is not common.
5. Protaetia (Potosia) negleda, Hope.
Baodipur Nallah, Kashmir. " On flowering bushes,
July and September 1906," three specimens.
A common and somewhat variable Indian species, with
a range extending northwards to E. Turkestan.
6. Protaetia {Eumimimeltca) terroaa, G. F.
Near Mhow, " July 27th, 1905," one specimen.
7. Protaetia aU>oguUala, Vigors.
Mhow. " On grass and flowering plants and flying to
lamp, June to August, 1905," nine specimens.
A common and generally distributed Indian species
ext«nding into Ceylon, but not before recorded from Central
India. Capt. Selous' series include blue, green and fine
coppery-red varieties.
8. Oxi/cetonia versicolor, !Fab.
Mhow, " On Date Palms, July 7th, 1905," two speci-
mens.
Capt. Selous' specimens of this very variable and widely
distributed species are the typical form (the " var. a " of
Arrow), with a shining upper surface, and the prothorax
and elytra extensively adorned witli red,
9. CMoloba acuta, Wied.
Mhow. " On grass, August and September 1905," four
specimens.
10. Bpicometia kirteUa, Linn.
Quetta. " Eating Iris flowers, March and April 1907,"
nine spedmeus.
This species is an addition to the list of Indian Cetoniidae.
The very doseiy allied E. squalida, Linn,, is recorded by
iM:,i,A J 1^, Google
104 Mr. Oliver E. Janson's Additions to
Arrow from the same locality ; both are common Kuiope&n
species and of wide distribution, ranging through Asia
Minor, Mesopotamia and western Persia to Baluchistan,
where the Indian fauna assumes a very Palaearctic char-
acter. All Capt, Seloua' specimens have a second white
spot on the outer discal costa, a httle before the middle
of the elvtra, that I do not find present in any of the many
.European examples that I have had an opporttmity of
examining.
Anatona salousi, n, sp.
<^ Body of &D oval, compaot and convex form. Caat&nemu,
shining; baae of head, the prothorax and elytra teataoeoos red,
opaque, and with shining oaatoneooB matkinga diepoeed as follows :
On the prothorax a mai^pnal band on each aide, and eight small
apoto, viz. three placed in a nearly r^nlar longitudinal row on each
aide of the disc and two in the middle (both prolonged anlariorly
with a t«ndenoy t« form a median stripe); on the ecnt«llum an
apical spot prolonged anteriorly ; on each elytron eight spots, viz.
one in the middle near the baae, one near the suture and just before
the middle, one about equidistant between the laet and the apex
and close tn the suture, and four adjoining the outer margin, the
first at the lateral sinus, the second and third (both small) behind the
middle and the fourth close to the sutural angle. The pygidium and
the sides of the prothorax, sternum and abdomen with irregular
chalky-white markings. Head somewhat flattened between the
eyes, coarsely and confluently punctured ; clypeus long, narrowed
in front, with a slight median carina and slightly prodnced and
strongly reflexed angles. Ftothoiaz nearly as broad in the middle
as at the base, strongly narrowed in fiont, the base feebly tri-sinnat«
and with the lateral angles rounded; the disc with a very fine and
sparse puncturatioo which becomes coarser and very much closer
in front and towards the sides. &cutellum large, slightly rounded
at the sides, impunctate. Elytra feebly bi-coetat«, with six discd
rows of small and rather remote pnncturcs, the sides with scattered
fine punctures and the apex slightly strigose. Fygidinm spanely
setose and remotely punctured, transversely strigose only in the
middle. Underside thinly clothed with yellowish-grey pubescence
and punctured at the sides ; metastemum with an impressed median
line ; sternal process broad, rounded and with an impre«aed hii8Ut«
line just before the apex ; abdomen with a broad central depression.
Legs stout, anterior tibiae with two large margin^ teeth, and the
apex rather obtuse, all the femora and tibiae thinly fringed with
long golden-grey hair. I«Dgth 17 mm., breadth 9 mm.
l„y,l,Ajh,COOl^lC
the knoteledge 1/ the Celwiiidae of Brititk India 105
Mbow (Type, coll. Janson) ; Poona (c<^. Briiish Museum).
Apart from the very peculiar coloration this pretty
species differs from its nearest ally, A. aiboguUala, fiuim.,
in it£ narrower and less convex foim, in having the pro-
thorax more abruptly narrowed in front, the clypetis more
strongly caiinate and the pygidium strigose only in the
middle. The darker markings on the npper side are pro-
duced by the absence of the lighter coloured opaque
indumentum, that covers the other parts of the stuface,
leaving the derm exposed and with a talc-like lustre.*
I have named the species after the discoverer, the late
Captun R. Selous.
Clerota rtglflca, n. sp.
Black and veiy ahiiiiiig above and below, with <naiige-7ellow
m^rfcingK cmmpiiBuig ft medifto atripe on the bead, a marginal band
on each' Bide and a median atripe on the prothorax, a spot 000a j^ing
neori; the whole of the Bcatellom, a broad and alightlj einooua
longitudiiial stripe on each elytron and a large triangular patch on
each lide of the pygidium. The sidee of the meeotboraoio epimera,
metathoracic epistema and poBt-cozae, and a large spot at the mdea
of the fint to fourth abdominal segmeato are aleo otange-yellow.
Head finely and sparingly pnnotured in the middle, more ooanely
panctored in the lateral fonows, olypeus aligbtly ainuoua at the udes
and distinctly widened in front. Prothorax almoet as bioad ae the
elytra at the baee and obliquely narrowed to the apex, broadly
■nlcate behind, very minntely and remotely punctured at the sides.
Elytia slightly narrowed behind and separately rounded at the apex,
anlcate at the sntnie, the middle third of the disc with several rows
of more or lees obsolete poncturee, the sntural stria feeble and
becoming obsolete before reaching the middle, the apical third of
the sides and the apex very cloeely and deeply strigose. ^rsidium
btoad and transversely convex, closely and concentrically strigose.
Underside of the body almoet entirely smooth, the flanks of the
piDtltorax and sides of the basal abdominal segment feebly strigose ;
Iritish HuBenm, has these shining spaoes covered
to a large extent with a dense white sqaamose or farinose clothing,
which is apparently very lightly attached and eaailj rubbed oS, and
in my type specimen is only present to a small extent on the sides
of the prothorax and on the pygidium. It would most probably
be found that in fteehly emerged examplee the clothing entirely
covered the spaoes, so that the markings of the npperside would be
white instead of oastaoeons as I have described Uiem.
iM-,i,A J h, Google
106 Mi. Oliver E. Jansoo's Additions to
meaosternal proocM vei7 broad at the baM, pointed and slightly
onrved inwarda at the apex; anterior tibiae with the two lateral
teeth and the prodnced apex acute. Length 36 mm., breadth 17 mm.
MouLMEiN, Lower Bunna (Type in coll. Janaon).
This fine species is closely allied to C viltigera, Hope,
but is larger and of a more robust form, the prothorax is
broader at the base and less sinuate at the sides, the elytra
more parallel-sided and longer in proportion to the pro-
thorax, and the aedea^rus of the mate is much broader and
lees constricted before the apex.
The type specimen, a male, was taken flying by the late
Col. Bingliam in August 1894, and I have others from
Penang; Mana Riang, Sumatra; and Euching, Borneo.
This distribution would seem to indicate that it ia a Malayan
^>ecies with a range extending northwards into southern
Burma.
C. bodhiiaUva, Kunck., an evidently closely allied species
from Annam and Tonkin, and only known to me by descrip-
tion, is of a much larger size (44 mm, in length) and luu
yellow markings on the elytra only.
Clerots amwl, n. sp.
$ Shining hiaok, the outer edge of the epimera and posterior
ooxae, and a Hmall spot on the sidee of the second, third and fourth
abdominal aegmenta orange-yellow.
Head stmngly punctured, the pnnoturee of an ovate form in hont
and coaTHei, cloeer and irregularly confluent towards the apex, the
extreme base smooth; clypeus a little narrowed towards the apes,
the apical lobes rounded and with a strongly reflexed margin, the
raised side margins becoming almost obsolete before teaching the
eyes. Prothorax obliquely narrowed from the base, with the aides
slightly sinuons and very narrowly margined, the basal angles
obtuse, the entire surface, exoepting a narrow and rather indistinct
median line, with a very strong punoturation that becomes more
dense as it approaches the sides and strigiforro close to the lateral
margins, the basal lobe broad and slightly depressed but not sulcata.
Scntellum slightly snlcate and smooth in the middle, pnnctnred at
the base, the apex very acute. Elytra gradually narrowed from the
base and sub-truncate at the apex, the basal part very convex and
smooth, strongly punctured on the median third, the pnnctur«e
extending to the hnineral c^osity at the side and forming five or
six tolerably regular rows in the middle, and one row next the suture
that extends to the base, the apical declivity and the stdee, behind
Ij.y.l.Ajl^yCOO'^lL'
the huftdtdge of the Cetoniidae of Britiak India. 107
the middle finely and oloMly strigoae. I^gidinm and tbe aplul
doml Mgmeiit of the abdomen finely and closely atrigofle, the former
Tery prominent And feebly bi-nodoae at the apex. Underside ooanely
bat rather spusely pnnctuied on tbe meta-eoxae and ddee of the
metMteninm ; mesoatemal prooeas gradnallj narrowed to the apex
wbere it ia acnte. Legs closely strigose on the femora and oat«r
side of the tdbiae, anterior tibia with the two marginal teeth and the
prodaoed apex veiy aonte. Length 33 mm., breadth 16 mm.
Ehasia Hili^, Assam (Typ« in coU. Janson).
The strongly raised mai^ns of the ftpical lobes of the
clypeuB form a semicircular rim to a pit-like cavity on
either Bide, and coalesce with the narrow median carina
at the base of the excision : this peculiarity and the atrongly
punctured and non-sulcate prothorax and closely strigose
legs will suffice to at once distinguish G. arrowi from all
the hitherto described members of the genus. In general
aspect it most nearly resembles narrow examples of C.
budda, G. P., that ate without the usual yellow markings
on the uppeTside.
Pseadoelwleothea rltsemae, n. sp.
Body of a narrow oral form and deeply channelled along the
median line of the npperside. Dark grass-green, shining; the
wdee and suture of the elytra with a, reddish Unge in some lights;
head, sides of the prothorax and onderside golden green; antennae,
palpi, tibiae and torsi fatvona tinged in parts with green and coppery
led. Head sparsely punctured, convex and smooth in the middle;
olypeoB a little widened in front, margined at the sides and deeply
notched at the apex. Prothorax strongly narrowed in front, th«
ndes nnnouB and slightly emarginate at the basal half, the basal
an^^ea prodnoed and snb-acnte, smooth on the diao, panctured and
strigose at the sides. Soatellnm panctured at the base only.
Hytaa gradually narrowed posteriorly, separately rounded at the
apex, the mtnral an^^es a little produced and acnte, some scattered
fine punctaree in the sntnral depression and a regular raw next the
mtme, at the sides fonr or five rather confused rows of punc-
tures on the bsaal half and thence irregularly strigose to the apex.
Pygidium prominent ukd closely strigose, deeply grooved and hi-
nodooe at the apex. Undenide of the body almost impunctate;
■terool process broad at the base and obliquely narrowed to a point
at the apex ; abdomen concave in the centre and broadly emarginate
at the apex. Front tibiae slender, curved and withont marginal
teeth; hind tibiae on the innerside with abroad, thin and flattened
u„,,i,Aji^,Coot5lc
108 Mi. Oliver E. Janson's Additions to
appendage carving forwards, and towards ita extremity narrowed
and strongly bent baokwards in the form of a sharply pointed hook.
Length 26 mm., breadth 12 mm.
In the female, besides the usual sexual characl«i8, the prothotax
is broader in middle than in the male, the pygidium is only feebly
bi-nodoBe, the apex of the abdomen is broadly rounded and the last
segment and the apical part of the preceding one have a oouse
setigerooB punotuiation, and the ont«r apioal spine of the hind
tibiae is broad and bi-mncronate.
Ranqoon and Penang (Types, ^, $, in caU. Janson).
This species comes neatest to P. vtrene, Kitsema, but is
smaller and of a much narrower and more parallel-sided
form, and has the piothorax more strongly sinuate at the
sides. The male difieis, moreover, in the form of the
appendage of the hind-tibiae, and the female in having
the pygidium sulcat* and the apical ventral segment of
the abdomen rounded, instead of broadly emarginate as
it is in vireite.
P. riisemae is the first Psetidochdcothea (if kept as distinct
from Pledrons) that has been discovered on the mainland,
the genus being essentially an insular one, with its head-
quarters in North Borneo, and the occurrence of this
species at Rangoon brings it just within the Umits of the
British- Indian fauna. Pledrone tristis, Westw,, b recorded
by Wallace from Penang.
Maaronota baliUifera, Bourg., Bull. Soc. Ent. France,
1914, p. 292.
This is the jiavofasciata. Arrow (nee Moser), and is de-
scribed by Bourgoin as a distinct species, differing in the
structure of the hind-fibiae in the male. It is recorded
from Bhutan and Assam.
The " female " specimen in ray collection referred to by
Arrow (" Fauna Brit. Ind.," Ceton,, p. 52) proves apou
dissection to be a male, and is the tme, fiav^asdata, Moser,
from Tonkin. The females of both species are apparently
still unknown.
Clinleria sternalts, Moser, Deuts. Ent. Zeite., 1910, p. 532.
This addition to the Indian Fauna would appear, from
the description, to come nearest to C. modesta, Blanch., in
coloration, but is stated to have the mesostemal process
broad and flat, a character not found in any other member
of the genus. It comes from Pegu, Lower^Buima.
ih,Cooi^lc
the knowledge of the Cetaniidae of British India. 109
Qlycosia durdi, Pouill., Insecta iv, 1914, p. 187.
From the description and figures this is evidently a
slight vanety of G. luctifera, Fairm., with markings Bimilai
to those of the Indian specimens described by Arrow.
It is also from Bhutan.
The FOLLowiNa Notes refer to the " Fadna of
British India," Getoniinae.
Macronata, p. 41.
In the synonymy and sub-genera CoUodera, Hope, 1831,
is cited as ondescribed ; it was later emended to Cododera,
and characterised by Burmeister, Handb. Ent. iii, 1842,
p. 320.
M. diardi, p. 43.
The two specimens referred to (p. 44) with black elytra
and purple-black prothorax are in my collection, and came
from the late M. Jacoby, labelled " Ceylon (W. Morton)."
This locality is most probably incorrect, as I now have
similar specimens, from the van de Poll collection, from
Mana Riang, Sumatra, and this form has since been de-
scribed as a distinct species, under the name maindroni,
by Boutin, Bull. Soc. Ent. France, 1916, p. 133.
M. mearesi, p. 45.
The type, as cited by Weetwood, was in the Party collec-
tion, which was dispersed by auction-sale, and the par-
ticular " lot " comprising this specimen was bought by
Mr. R. Oberthur, in whose possession it should now be
found.
Cyphonocephtdus amaragdyluB, p. 69.
Reference to Westw., Trans. Ent. Soc. Lond., 1878, p. 29,
pi. 1, %s. 3, 4, is omitted.
TorynorrMna distinda, p. 82.
Var, mlipes, Burm. Handb. Ent. iii, p. 779; Westw.,
Arcana Ent. i, p. 120 and 192, is omitted from the synonymy.
This is the brassy green variety and usually has the legs
tinged with blue.
Rhomborrhina microcephala, p. 87.
Var. staudingeri, Nonf. Stett. Ent. Zeit., 1890, p. 17,
is omitted. Nonfried described this as a variety of micro-
iM-,i,A J li, Google
110 Mr. Oliver E. Janson'a AMUiotu to
cephala, but the descriptioa applies to the typical form,
and the name therefore sinks aa a synonym.
The specimen bearing the " Type " label from the Parry
collection, cited by Westwood, is in my possession.
Helerorrhina sinuatocoUis, p. 96.
The type is incorrectly stated to be in the Paris Museum.
The specimen described and figured by Westwood (as a
variety of degans. Fab.), as cited by him, was in the Parry
collection, and is now in my possession. It bears Pany's
" Type " label, and must be regarded as the type of this
species, Westwood's description taking priority over that
of smaragdina, Burm. [nee G. P.), and it is the type of the
latter that is in the Paris Museum.
AtuUona alboguUata, p. 115.
Reference to Westw., Trans. Ent. Soc. Lond., 1874,
p. 476, pi. 7, fig. i, is omitted.
Glycyphana minima, Bates, Entomologist, xxiv, 1891,
Supp. p. 21.
This species, founded on a single specimen stated to have
been received from Captain G. Young, from the Hill region
of Kulu, Korth- western India, is omitted. Mr. Arrow
informs me that he doubts the correctness of the locality
assi^ed to it, and behoves it will prove to be of Malayan
origm. I have at present seen nothing from India that
agrees with the description.
Ooliathopsis despedus, p. 206.
The type of this species is undoubtedly the specimen in
the Oxford Museum, which I have examined and found to
agree with Westwood's description and figures. It, more-
over, has the mouth-parts extracted and moimted on a
card beneath the specimen, as is usually the case with the
Westwoodian types. I have before stated (Cist. Ent. ii,
1881, p. 610) that Westwood's figures were evidently not
made from the specimen that is in the British Museum.
The examination of the type of despectus has shown me
that cervm, Jans., is not the same species. In comparing
the two forms (the female type in both cases) I find that in
cervus the clypeus is more broadly and strongly refiexed
at the apex (especially at the sides, where it projects in an
obtuse angle), the prothorax is conspicuously larger and
n,g,i,.eih,.G00glc
&e knowledge of the Cetoniidae of British India. Ill
more convex and has the basal angles more prominent;
it is also of an altogether stouter and more robust form
than despetiua, and the mouth-parts differ, as will be seen
on comparing the figures that accompany the original
descriptionB of both species.
The following that have been relegated by the author to
the rank of varieties and synonyms, have, I coosider,
sn£Bcient distinctive features to warrant their retention as
species:
Cypfumocephaltts stmiTagdtdus, Westw., sunk under C.
Uivaceus, Dap., p. 69.
Heterorrhina mitrata. Wall., sunk under Diceros divea,
Westw., p. 72.
Gtycifphana svbcincta, Jans., sunk under 0. torquata, F.,
p. 124.
Protaeiia aerata, Er., sunk under P. orienUdis, G. P„
p. 143.
Clinieria valida, Lansb., sunk under C. auronoUUa, Bl.,
p. 179.
Clinteria decora, Jans., sunk under C. Idvgi, Hope, p. 187.
Agestrata samson, Sharp, sunk under A. orichalcea, L.,
p. 192.
GcAieUhopsis cervus, Jans., sunk under G. despectus,
Westw., p. 206.
n,g,t,7.dh,G00glc
( 112 )
Y, On the Protocerdmim of Micropteryi [Lepidoptera).
By P. A. Buxton, B.A., F.E.S., M.R.C.S., Fellow
of Trinity College, Cambridge (Lieut. R.A.M.C.).
[BMd FebnMff Tth, 1917.]
Plates VII-X.
Table of CoNTENra.
FoBiwoBD us
iNTBODucnon . . . . . .Ill
Thb Fkotoobbbbrdh of MlCaOPTBRYZ —
I. The Neurilemma and Ganglion Cells . . 118
n. The Protocerebral Lobea 120
UL The Mushroom Body 124
IV. The Central Body 133
V. The Bridge 134
VI. The Viaual Centres 136
Vn. The Frotocerebral Tracts 136
SVHMABT 138
TXCHKIQIII —
I. Fixation and Impregnation 141
n. Section Cutting 145
m. Staining 146
IV. Note [Composition of BeagenU) . .148
BiBUOOIUPBT 149
Abboktiations 151
EXFLANATIOir OF Pl^TES 152
FOREWORD
Moke than four years ago I commenced to study the
internal anatomy of Micropteryx {Eriocephala) in the hope
that I might be able to throw some light on the question of
its systematic position. As is well known, most entomo-
logists regard it as a primitive ]>pidopteron (Protolepido-
ptera), though thete ia really gmte as good ground for
regarding it as a Trichopteron.* I am now ia a position
to publish my results only in so far aa they relat« to a
portion of the brain of that insect. This I do with a feeling
* Vidt, however, Dr. Chapman's paper (Trans. Ent. Soc., 1916.
pp. 310-4, pL 81-93, (1917)), which raises ificropferyz to ordinal
rank (order Zeugoptera).
TRANS. ENT, 800. LOND. 1917. — PART I. (NOV.)
l„y,l,Ajh,COOl^lL'
Mr. P. A. BuztoD on Prolocerebrum q^ Micropleryx. 113
that aome apology should be made, because I do not at
any rate describe the whole brain : it is owing to the war
and pressure of other work that there is no likelihood of my
being able to carry my investigations further. This paper
is however complete in itself, and is not of the nature of a
preliminary note. From the point of view of comparative
anatomy, the Frotocerebrum, with which alone this paper
deals, is by far the most important part of the insect brain,
not only on account of its complicated stracture, but also
because of certain qnestions relating to the homologies of
some of its parts. Such questions of homology do not at
present arise in connection with the Deuterocerebrum or
Tritocersbrum, or ventral brain. My hope that my inves-
tigations would throw Ught on the systematic position of
MicTopUryx will not be fulfilled until we can compare the
brain of this insect with a number of other Lepidopterous
and Trichopterous brains, after they have been fully and
properly investigated.
I have made some attempt to render this paper useful
also as an introduction to the study of the brain of insects;
this I think is justified, because it is the first paper on the
Babject published in Great Britain since 1878 (Newton),
and I know by bitter experience how exceedingly difficult
it is to obtain a clear knowledge of the subject from a
variety of papers written by many men in many languages
at different dates. My taak has been rendered difficult
by the small size of Micropteryx, which is, I believe, the
smallest insect of which the brain has been investigated
in any detail.
I take this opportunity of acknowledging how much I am
indebted to many friends in the University's Zoological
Laboratory at Cambridge, particularly to Mr. F. Balfour
Browne for constant criticism and much good advice, and
for reading through the whole paper before it was pubUshed ;
also to Mr. L. A. Borradaile for helping me with the theory
of the segmentation of the head of the Arthropoda ; also
to Dr. D. Keilin of the University of Paris, and of the
Quick Iiaboiato^ and Magdalene College, Cambridge, for
putting at my disposal his deep knowledge of fixing and
staining. Canon W, Brocas Waters gave me a room to
work in, while I was stationed in Bury St. Edmunds on
military duties.
I must also acknowledge my indebtedness to Dr. K. F.
Kiihnle of Stuttgart for his paper on the brain of the
TBAN8. airr, soc. lond. 1917. — pabt i. (nov.) i
114 Mr. P. A. Buiton on the
Earwig and other types (see Bibliography), which gives a
full review ol insect neurology up to 1913, together with a
bibliography and a table of the terminology of the insect
brain, which has materially lightened my labours. This
paper is by far the most important contribution which has
yet been made to insect neurology.
All my material has been collected in the neighbourhood
of Cambridge, and I have worked entirely with Micro-
jjteryx {Eriocephala) • adthdla (Linn.), and not with any
other species.
INTRODUCTION.
The anterior part of the central nervous system of insects
consists of a suprateeophageal portion, which is the biain in
the narrow sense of the wonl. From this the circum-
cesophageal conomissures pass round the (esophagus to the
ventrocerebrura or subc^phageal portion of the brun.
Most authors include this also in the brain of the insect.
The supracesoph^eal ganglion, or brain in the narrow sense
of the word, was foimd by Viallanea to be composed of three
paired elements or neuromeres, which he believed to be
segmental ; these give rise to the parts of the brain to which
he gave the names Protocerebrum, Deuterocerebrum (or
Deutocerebrum) and Tritoceiebnun. We now know that
the Protocerebrum is not a segmental ganglion ; and it
will be convenient at this point to give a short sunmiar>' of
the results obtained by those who have studied the subject
of the segmentation of the head of the Arthropoda, and the
homologies of the various appendages throughout the class
from the point of view of comparative embryology. The
whole matter is one of great difficulty, and has been neg-
lected by insect neurologists; as, however, it is a matter
which bears directly upon the subject of this paper I give
this summary of our knowledge in so far as it affects the
insect protocerebrum.
* The eubj«ct of this paper belongs to the true genus Mieropltrffx
Hlibner. lliiH genus nas been referred to in some writings, e.g,
Meyrick's Handbook, the Cambridge Natural History, etc., ander
iU synonym Erioctphala Curtis : while the leaf-mining eenus erron-
eously called " Micropleryx " in certain of the same works should be
known as EHocrania Zeller.
Sriocephala Curtis = MicTDpteryx Hilbner.
" Micropleryx " auctt- (nee Hiibner) = Sriocrania Zeller.
See Tutt, Brit. Lep., I, pp. 129-137 (1899). and Steudinger-Rebel,
Cat.. U. pp. 246-8(1901).
l„y,|,AjL., Cookie
Protocere6mm <^ Mu^opleryx. 115
The brain of the Arthropoda in its fullest development,
tiiat is to say as exemplified in the brain of the embryo of
Scolopendra (Heymons), consists of the following parts :
au archicerebrum, which is median, unpaired and preoral :
three lobes on each side, the ajiicerebral lobes, the outer
two of which arise from a common rudiment ; these also
are pteoral, and together with the archicerebrum form the
syncerebrum : the preantenuary ganglion, or protocerebrum,
which is the ganghon of the &:st somite, or true segment ;
we believe that this was primitively postotsi, but it is
preoral in all living Arthropoda : the deuterocerebrum or
anteonary ganglion, and the tritocerebnim or premandi-
bular ganglion, which correspond respectively to the second
and tMrd somites. It may be said at once that the deutero-
cerebrum and the tritocerebrum correspond in Heymone'
nomenclature to the organs which I shall subsequently
describe under those names. This is not, however, the case
with the protocerebrum, for that word has been used in a
great vanety of senses. In the development of the insect
head that part of the central nervous system which entomo-
lo^ts generally call the protocerebrum (Viallanes) is
develop^ from the archicerebrum and the syncerebral
lobes ; we do not yet know which parts of the insect brain
correspond to which of these structures, except that the
outer syncerebral lobe gives rise to the optic lobe, and
Haller suggests that the mushroom body is formed from
the archicerebrum. The preantenuary ganglion or proto-
cerebrum of Heymons is not found at all in the insect
head, and is to be carefully distinguished from that part
of the br^Q which is commonly called by that name. The
aynouymy is further complicated because the preantennary
gangUon or protocerebrum of Heymons is the precerebnim
of VerhoelT, and the word " protocerebrum " has been
used ID yet a third sense to denote the procerebrmu of
Heymons, that is the syncerebrum and preantennary gang-
lion (protocerebrum] together. The word protocephalum
has been used by Holste, and perhaps by others, to denote
that part of the brain which is dorsal to the gut in the
insects : »'. e. the syncerebrum of Heymons (the proto-
ceiebnun of insect neurologists since the time of Viallanes),
with the deuterocerebrum, and the tritocerebrum.
I shall continue to use the word protocerebrum in the
sense in which neurologists from the time of Viallanes have
alvays employed it, though I should be glad to avoid a
iM-,i,A J h, Google
116 Mr. P. A. Buxton on the
word to which bo many meanings have been assigned. By
it I mean a mass of nerve tissue arising from that preoral
part of the embryo which is not segmented and which bean
no appendages. It b the nervous element corresponding
to the acron of some embiyologists, and it is not the serial
homologue of the deuteioceiebmm and tritocerebrum
(mesocerebrum and metacerebmm of some writers on the
segmentation of the Arthropoda). The protocerebrum of
insects is, in fact, the synceiebrum of Heymons, unless it
contains some element not yet differentiated as belonging
to the pteantennary ganglion.
In tlus paper I propose as I have said to deal solely with
the protocerebrum. I give, however, the following brief
summary of the function and connections of the other two
BupracBsophageal ganglia. The deuterocerebrum is the
ganglion of the antenna, to which it gives motor and sensory
nerves. The pair of ganglia fomung the deuterocerebmm
are united across the middle line above the cesophagus, and
lie before and below the prutocerebral lobes. They are the
antennaiy or olfactory lobes of some authors. The deutero-
cerebrum gives rise to the paired sympathetic system, which
lies upon the lateral wall of the cesophagus on each side ; thia
consists of two pairs of small gangha with nerves which con-
nect them to each other, and, as is known in some insects, to
the median or tritocerebral sympathetic system. The pair
of ganglia composing the tritocerebrum lie on each side of
the anterior part of the cesophagus and are generally fused
above to the rest of the supraoesophageal brain. The lower
part of the tritocerebrum is the circumoesophageal connec-
tive or commissure. This ganglion supplies the labrum,
but has no paired appendage connected with it in the
insectfi; a band of fibres, the tritocerebral bridge, passes
across from one side to the other beneath the oesophagus.
The tritocerebrum also gives rise to a pair of fine nerves
which run forwards and inwards to the frontal gangUon,
which lies upon the upper surface of the anterior part of the
oesophagus. This is the largest ganglion of the sympathetic
system : from it a fine nerve runs forwards and another
backwards. This latter, the nervus recttrrens of some
authors, connects the frontal ganglion with a short chain of
ganglia lying on the upper surface of the oesophagus, and
from this unpaired sympathetic system the stomodaeum
is innervated.
The suboesophageal ganglia or nerve masses will not
l„y,|,AJI^,COOl^lC
ProtocerArum of Micropteryx. 117
agwn concein us. They are formed by the fnaion of fonr
segmental ganglia, the mandibular, the intercalary, the
mazillaiy and the labial. The intercalary ganglion has
hardly been noticed by insect neurologists ; the correspond-
ing appendage is the maxilliila, which is vestigial or absent
in adult insects ; the ganglion is accordingly ill-developed
or absent. The other three ganglia are mainly if not
entirely motor and sensory centres to their respective
append^es.
AH the nerve centres of insects consist of the following
layers.* They are hounded externally by a neurilemma,
which is a thin syncytial structure. Witlun this, and lying
loosely in a qiiantity of fluid, are the nerve cells, or ganglion
cells. The processes of these, the axons, pass inwards to
form the innermost part of the centre; here they unite
in very lai^e numbers to form the tissue known as axonic
sabstance (or fibrillar material], which consists of innumer-
able axons and their twigs bound together by a varying
amount of neuroglia. Of this axonic substance two types
may be distinguished; the first is that which is called
Piuiktsubstanz, or Marksnbstanz, or neuroepongium ; its
composition was first accurately explained by Viallanes.
Until his day it had been known as a tangled web, bnt
in it he distinguished very fine axis cylinders running
in all directions, and their twigs, and also the neurogfia.
In the second type of fibrillar substance, the Fasersub-
stanz, the axons run in bundles and form welt-defined
tracts in which there b little or no neuroglia. The
distinction between these two types must not be insisted
upon, for every degree of intergradation may be found;
even in the most indisputable Funkteubstanz small tracts
of fibres may generally be detected. Physiologically again
the difference is one of degree, though Fasersubstanz
is mainly a tissue of conduction, Funkteubstanz one of
association, that is to say one in which impulses pass from
one neuron to another.
Two parts of the brain may be connected either by
fusion of their component Punktsubstanz (Versckmdzuvgen,
soudnrei), or by definite tracts of Fasersubstanz {Faaer-
^>erbindungen). This distinction, again, has only a relative
value.
Before we pass to examine the structure of the various
iM:,i,A J. ^, Google
118 Mr. P. A. Buxton on the
parts of the protocetebnim it should be realiaed that this
portion of the brain ia not merely a complex but also an
entity. We might compare it physiologically to the cere-
brum of a vertebrate. Both are known to be the highest
or governing centres of the organism ; both possess a soli-
darity or unity of action ; both consist also of parts, each
of which in its turn is not only an anatomical, but also a
physiological entity. We are quite justified in regarding
the protocerebrum of an insect as the headquarters from
which are directed all those complicated reactions and
instincts of the organism which give to it* activities what
at any rate appear to be purpose, and adaptation to the
surroundings. There is too great a tendency to lay
emphasis on the potentialities for independent action
which are undoubtedly possessed by the lower nerve
centres of the insect. In all but the very lowest insects
there is a marked degree of speciaUsation in the structure
and function of the protocerebrum, and this fact of ita
solidarity should not be allowed to pass from the mind while
we study in detail the structure and perhaps something of
the function of its parts.
THE PROTOCEREBRUM OP MICROPTERYX.
I. The Neurilemma and Ganguon Cells.
The whole brain of Micropleryx, that is to say the axonic
parts and the cells, is included in a limiting membrane or
Neumlemma (Plate X). This is a very fine sheet of
substance which stains well with the acid stains. It is
certainly a syncytium in Mxcropteryx, and probably in all
insects. In places where no ganglion cells intervene be-
tween the neurilemma and the a.xonic part of the brain
the two are closely applied to each other, and the neuri-
lemma can hardly be distinguished, though in material
fixed in the picro-chlor-acetic mixture it can generally be
seen. Occasionally the nuclei of the neurilemma can be
seen even when the layer itself la indistinguishable. The
neurilemma is somewhat thickened mid-dorsally, partly
owing to the fact that a number of tracheal tubes (Plate X)
lie in it in this position, partly owing to the presence of a
number of the nuclei, the celb corresponding to which have
fused to form the syncytium of which the neurilemma con-
sists. These nuclei are elongate and smaller than those of
l„y,l,Ajh,COOl^lc
Pratocer^mim of Micropteryx. 119
" normal " ganglion cells. They ettan deeply with hema-
toxylin. The neurilemma is continuous over the whole
brain dorsally and ventrally and also over the optic lobes ;
it ia continued downwards to cover the ventral parte of the
central nervous system.
The Gakouon Celis.— These are spread over the whole
anterior and superior parts of the brain, in a layer which
reaches its greatest thickness mid-dorsally. The layer is
discontinuoas or absent beneath the protocerebnim. On
the upper side of the brain the cells may be as many aa
15 ce\& deep, particularly near the middle line between
the ocelli. Various types of ganglion cell must be distin-
guished. They all possess a spherical nucleus and a very
small quantity of cytoplasm. The normal cells (j. c.)
cover the protocerebnim above, before and behind. Cells
of this type, which is much the most abundant, are either
motor or else cells of connecting-fibres (Kenyon). The
cells of the mushroom body {mb.c.) are found as a rounded
mass of cells lying just over the head of that organ. Their
nuclei stain heavily with htematoxylin ; they are also
smaller than the normal cells. The fibres from these cells
Ess into the mushroom body. The distinction in size
tween these cells and those of the normal type is not very
great ; it can be best observed in material fixed in Gilson's
fluid. The celb of the optic lobes (o. c.) are still smaller
than those of the mushroom body; and their nuclei
are absolutely spherical and stain very heavily and com-
pletely. No structure within the nucleus can he detected
in ordinary sections and this gives to the masses of cells a
very characteristic appearance. The c}^plasm, per contra,
is scarcely stained at all. These cells form a deep coating
which completely envelopes the three optic ganglia; this
layer is less deep above than below. In Mtcropleryx giant
cells (gi. c.) are found in small numbers round the base of
the mushroom body just where it passes into the proto-
cerebral lobes (fig. 12, etc.). These cells are few in number,
hardly more than a score on each side. Their nuclei are
sphencal and about four times as large as those of normal
cells ; there is a considerable quantity of cytoplasm, which
can be stained with eosin; this distinguishes it at once
from the cytoplasm of the other types of ganglion cell.
The nucleolus is generally clearly seen. Haller states that
these cells are mainly, but not entirely, concerned with
conduction across the middle line, and that their axons
iM-,i,A J h, Google
120 Mr. F. A. Buxton on the
pass to the opposite side of the brain, to the antennary lobe
or head of the mushroom body, or even into the optic
lobe. Inside the brain are found small cells, lying singly
or in ^ups (Plate X). Kiilmle refers to them as
neurogha cells {ng.) and doubtless he is conect in so doing.
They are found particularly in the space surrounding the
central body and in the interval between the two capsules
of that organ ; there are also a few on the surface of the
stem of the mushroom body and in other places (PI. X).
The nucleus of a neuroglia cell is pyriform or elongate and
stains deeply, and its outline is generally irregular. The
nucleus is smaller than that of a " normal " ganglion cell.
The axons from the ganglion cells enter the axonic part
of the brain vertically ; and they are generally united into
small bundles at their point of entrance. It is to this that
Kiihnle gives the name Eitislrommung.
Spherical black granules occur among the cells in material
fixed in osmic acid, or any mixture containing osmic
acid; These granules are not found in material fixed in
any other fluid, and I regard them as unsaturated fat.
They are found among the ganglion cells and are quite
definitely extracellular.
A note on technique is given at the end of the paper;
suffice it to say here the cells may be studied in material
fixed in the picro-chlor-acetic fluid, but that some specimens
may with advantage be fixed in Bouin'a or Gilson's fluids,
especially for the study of the different types of cell.
Trachkation. — In the brains of most insects fine
tracheal trunks can be detected ramifying in the fibrillar
part of the organ. This is not the case in Micropteryx,
perhaps because of the extremely small size of the whole
insect : ao far as I can discover there are no tracheae at all,
either in the ganglion cell layer or the axonic fibrillar part
of the brain. There is a considerable collection of tracheal
trunks (tr.) in the neurilemma which lies over the mid-dorsal
part of the brain (Plate X), and it is at lea.<;t possible that
it is the fimction of these trunks, which are large and
numerous, to oxygenate the brain by diffusion through the
fluid which lies beneath the neurilemma.
II. The Protocerebral Lobes.
The protocerebnmi of insects is generally described as
consisting of the protocerebral lobes and the various
etructures such as the mushroom body, the central body.
l„y,l,Ajh,COOl^lC
Protocerdmim of Micropteryx. 121
the bridge, the ocellarj gloDienilus, and the optic lobes.
The term " protocerebral lobes " is a compreheitaive term
for the great mass of the protocerebrum in or upon which
the other structures lie. The word " lobes " is perhaps
unfortunate, but its use in all papers from an early date to
the present day renders it a classical term. In all insects
the protocerebral lobes {pc. !.) fonn by far the greater part
of the brain ; they are bilaterally symmetrical about the
middle line, but they are not divided from one another by
a raphe. Eiilmle describes their fusion across the middle
line (Versckm^zung), " above," " below," etc. This is per-
fectly accurate, but it would give a clearer impression to
say that the two lobes are united over their whole extent
at the middle line, except that in the centre a space (la
loge, Viallanes) is left in which lie the central body and
ocellary glomenili, and the inner root of the mushroom
body.
The union of the two sides in Micropteryx is complete,
but much less definite posterodorsally. Only a very vague
web of fibres covers the central body in this region. It
would, for instance, be possible for a micro-organism to
swim down from the flmd in which the ganglion cells he
through this web into the space surrounding the central
body. The protocerebral lobes together form a rounded
mass, with it£ longest axis in the transverse direction.
The mass is flattened above, and prolonged downwards to
fuse with the deuterocerebnim and the tritocerebrum.
The dividing line between the deutero- and trito-cerebrum
cannot be accurately determined. In the embryo they Ue
behind one another. In most adult insects, and Micro-
pteryx is DO exception to the rule, the deuterocerebrum is
pushed forwards, and the tritocerebrum fuses directly
with the protocerebrum, at any rate by a small part of its
posterior surface. From the lateral side of the proto-
cerebrum the optic nerve is given off. This connects the
protocerebrum to the optic lobes (medullary masses of the
eye).
The relations of the protocerebral lobes are as follows
(text fig. 1, p. 122) : mid-dorsally lie the rounded heads of the
mushroom bodies, and between them the bridge. SUghtly
in front of this the ocellary nerve is seen, disposed in a
transverse plane with a slight inclination downwards and
backwards. Various organs he within the protocerebral lobes
in a space full of fluid which has been called la loge by
u,y,i,Ajh,Cooglc
122 Mr. P. A. Buxton on the
Tiallanee. These organs axe the central body and the ocellary
glomemluB, and the Btem and inner root of the mushroom
body ; the forward and backward roots are also buried in the
protocerebral lobes, but they are not definitely separat«d
from the surrounding parts by a free space. It may be
mentioned here that the mushroom body system is com-
pletely buried in the protocerebrum except at three points.
The parta which project are the head, the lower end of the
stem, and the tip of the forward root (see pp. 125 aqq.).
TiXT noma 1. — General relationahipB of the parts of the proto-
cerebrum. The organ is divided at tLe middle line and the left
side is shown in the finue. viewed from in front. The cut surface
shows the loge of VioUanes ajid the organs within it. No cella ore
ahown; the whole organ as drawn here consisU of axonic substance.
a, c, i. i, q, Kfet to traote of fibres (see p. 136). br. bridge, d. I.
dorsal protocerebral lobe. hd. head of mushroom body. in. ea,
inner capsule of central body. in. r. inner root of mushroom body.
/. I. lateral lobes of protocerebrum. lo. la logo (Viallanes). mi. L a.
and mi. L p. anterior and posterior parts of middle lobe of proto-
cerebrum. oc gl. ocellary glomerulus, oc. n. ocellary nerve, cm. <n.
outer capsule of central body. #u>. hd. swollen head of ascending
branch <a mushroom body. ew. al. swollen foot of Bt«m of mnshroora
body, to dm. tracts passing from protocerebrum to deuteroceie-
brum. (tt. tumulus. vl.La. anteiior part of veotrolateral lobe.
Ij.y.l.AjhyCOO'^lL'
Protocerebnim of Micropteryx. 123
Special names have been assigned to various parts
of the protocerebmm (refer to PI. VII-IX), Thus doiBally
there is the dorsal lobe (Haupllappe), below this the ventro-
lateral lobe or Nd>enlappe, and midventrally the middle
lobe (Mittdstuck). These parts may all be distiDguished
in Mua-Mderyx, and perhaps the mere shape of the lobes
merits description. The dorsal lobe {d.l.) is the widest
part of the whole brain. In front its superior surface is
flat ; further back there is a specialised rounded projection
in the middle line, to which I give the name Tumulus ((«.).
This lies between the heads of the two mushroom bodies,
and consbts of a very tight homogeneous web of axonic
substance more densely compacted than any other part of
the brain. The portions of the dorsal lobe which lie around
and beneath it are of an extremely loose consistency
(see n. X).
A large lateral lobe (l. I.) is present on each side.* ItA
relations ate shown in figs. 10, 12, 13.
The ventrolateral lob^ (N^yenlappe) consist of two very
definite parts placed one in front of the other. The anterior
ett of the ventrolateral lobe {v.l.l.a.) appears aa a swelling
low the anterior extremity of the stem of the mushroom
body ; in the region beneath the inner root of the mushroom
body the lobe is insignificant; and behind this its posterior
part (v.l.l.p.) appears as a large round lobe above the exit
of the motor antennary nerve from the deuterocerebmm.
The middle lobe (mi. (.}of the protocerebrum lies between
the two ventrolateral lobes, hi most insects it consists
of a single body, shaped like an honr-glass, and lying
transversely between the inner roots. In After opteryz
we can distinguish an anterior and a posterior part of the
lobe. The anterior portion {mi. I. a.) is of the shape of an
hoar-glass, and lies, as it should, between the ends of
the inner roots; it is connected with the anterior part
of the ventrolateral lobe on the same aide by a tract
of nerve fibres (tract h). Behind it there is a transverse
bar of axonic tissue, placed below the ocellary glomeruli
and above the various bands which connect the two anten-
nary lobes or deuterocerebra ; this is the posterior part
{mi. I. p.) of the middle lobe ; to the antennary lobe and
also to the protocerebrum above it this middle lobe is
united by well-marked tracts of nerve fibres {tracts j and k).
u,y,i,A J h, Google
124 Mr. P. A. Buxton m the
Histologically the whole of the protocerebral lobea are
very uniform iu structure, and consist of Punktsubstanz of
a moderate degree of density. The tumulus, however, b
very much closer in sttuctute, and the parts of the proto-
cerebial lobes immediately below and aioimd it are ver}'
loosely formed. The lobes aie penetrated in all directions
by bundles of axons (Faseisul^tanz), some of which are
enumerated below.
III. The Mushkoom Body.
(Stalked body — Packard. Pilz — Kiihule, etc. Gestielte
Korper — Leydig. Lee Corps PSdoncul^s — Dujardin.)
Before I describe the mushroom body of Micropleryx it
may not be out of place to state that the organ consists
typically of a cup-shaped or globular head {calice, PUzhut,
Becker, Jobeei oonvdvtifms) supported by a stem {Slamm, lige,
caulundus, pedunculus) which divides below into a number
of roots or branches. The word Stiel is used by Kiihnle to
denote the stem and roots together. An early worker,
Newton, described the brain of the cockroach. In this
insect the bead and stem of the mushroom body are double,
and Newton named the two stems the cauliculus and
pedunculus, respectively. A small number of insects have
their mushroom body formed on a simpler plan, with only
one head and one stem. It is better, therefore, not to use
the terms cauliculus and pedunculus, which are responsible
for the notion that two supports of the mushroom body are
to be looked for in the typical insect biain. As will be seen
later the homologies of the roots of the organ are very
obscure.
This exceedingly simplified account of the least complex
type of mushroom body may serve to remind the reader
of the essential characters of that organ. A full account
of that of ^orjJcu^, together with a painstaking summary of
previous work, is given by Kuhnle. This is valuable, but
as I shall e7q)lain later I believe that Kuhnle has made
a fundamental mistake in homology. The summary in
Packard's text book is out of date and most difiScult to
understand.
At first sight it appears that the mushroom body of
Microjiteryx is formed on a plan not altogether identical with
that found in other insects. This is not very surprising,
for nothing is yet known of the brains of the Lepuioplera
or Tricko^era. I hope, however, to show that the diffi-
l„y,l,Ajh,COOl^lL'
Prolocerelmim of Mieropteryx. 126
cutties are rather apparent than real, and that Mieropteryx
is leaUv one of thoae organisms froni the study of which
we may draw valuable inference as to the comparative
anatomy of the insect bmia. It is always a most difficult
thing to form a picture of the mushroom body of an insect
when that oi^n is described by another worker. This is
dae, in part at least, to the complexity of the organs which
lie in all three planes of space. I shaJl endeavour to make
my meaning clear by giving several sketches of a mental
reconstruction of the organ in question. Its structure in
Mieropteryx is comparatively simple, and I have not found
it necessary to make a wax-plate model. I have, of course,
most carefully examined sections in all three planes of
space (text figs. 2, 3, pp. 12&-9).
The bead of the mushroom body of Mieropteryx is a single
globular mass of axonic tissue, and belongs to the Rocker
type of Kiihnle. It appears that a similar structure was
described by Fldgel in certain moths; but it is never easy
to understaad his descriptions, for he was much hampers
by the defective methods of his time.
The head of the mushroom body (Ad.) projects conspicu-
oosly into the ganglion cells above and behind. It is not
so large, however, as to make a prominence in the upper
surface of that layer.
From the cells of the mushroom body (p. 119) the fibres
pass into the head of the mushroom body. We must notice
that there are no points at which a number of fibres enter
together ; the entry is general and spread over the whole
surface of the head. Eenyon's application of the Qolgi
method to the brain of the bee mi^es it clear that after
entering the substance of the head the fibres give off a
coUateral branch which in turn divides to form twigs.
These twigs interlace with similar twigs from the collaterals
of other cells to form a glomerular body {Faserhdllchen).
There must be several score of these bodies in the head of
tie mushroom body of Mieropteryx. They are very small
and by no means easily distinguished. After giving off
its collateral the fibre proceeds downward as a component
part of the stem. The fibres do not form a definite tract
within the head, but pass through in a diffuse manner. It
is only when they reach the underside that they unite to
form the stem.
From the inner and inferior aspect of the head of the
muahioom body a band of fibres sweeps downwards and
126 Mr. P. A. Buxton on the
inwards paat the side of the outer capsule of the central bod^.
This band gives off a few fibres to the outer capsule (tract n),
and then passes into that region of the protocerebral lobes
which lies on each side of the central body (tract r) ; thete
the band divides and is lost to sight (PI. VIII, fig. 12, and
PI. X). Such a tract has not been described in other
insects.
The stem (st.) of the mushroom body leaves the ventral
side of the head and passes downwan^ and forwards and
also slightly inwards ; it is a single cylindrical rod of parallel
fibres ; and whatever may be the case in other insects it
is not penetrated by a canal. It may also be noticed that
it is not surrounded by a sheath. (Even if it were I should
not follow Kiihnle in calling the sheath a neurilemma.
A neurilemma is a syncytial layer covering a brain or
ganglion.)
The stem is one of the most striking features of any
section in which it occurs; it stains more heavily than
the surrounding protocerebral lobes, and is a most use-
ful landmark. The stem is a cylindrical structure, and
well-fixed material shows that the greater part of its
shaft is surrounded by .a space which is not developed
at its top or bottom; at these pointe the stem simply
pierces the surrounding parts of the protocerebral lobes.
At its lower and anterior end the stem is swollen and be-
comes superficial, that is to say it is no longer buried in
the protocerebral lobes, but reaches the surface in the
interval between the dorsal lobe and the anterior part
of the ventrolateral lobe. At this point it is covered by a
few nuclei ; some of these are merely nuclei of neuroglia
cells, some are nuclei of true nerve cells, which send their
axons into the foot of the stem. This is certainly remark-
able, but I have satisfied myself that it is the case by
examining a large number of sections through this region.
At the bottom of the stem we should expect the roots to
arise, and actually we find two processes of rather uncertain
homologies, both of which make a marked angle with the
stem. The first runs inwards and slightly backwards, and
is the inner root (in. r. ; innere Wunel of Kiihule). It is
straight and cylindrical and separated from its fellow of
the opposite side by a very distinct part of the proto-
cerebrum, the anterior part of the middle lobe. At its
termination the inner root \a sotiiewhat swollen.
The second structure which leaves the anterior end of
l„y,|,A J 1^, Cookie
Prolocerdmim of Micropteryx. 127
the stem may be called the ascending tnmk (as. tr.). The
homologies of this organ are obscure and will be discussed
later. It runs upwwds for a very short distance and
divides into two portions.
At its point of division it is swoUen. For the moment
let us call one of its branches the ascending and the other
the posterior branch.*
The ascending branch {as. br.) runs up directly to the
surface of the brain and is there swoUen into an acom-
ahaped head. It is covered by a thin layer of ganglion cells
of the normal type, and these send their axons into the
branch. This is a point of interest, for here and also at
the foot of the stem we have a few nerve cells which appear
to belong to the mushroom body. Similar conditions have
been occasiooally described m other insects, notably in
Penplanela by Haller.
The posterior branch ('po. br.) of the ascending tnmk runs
backwards and sHghtly upwards and inwards. Its ter-
minal part is bifid, but the two portions do not diverge from
one another. It is completely enclosed in the protocerebial
lobes, and even in fixed material is not always very easy
to see. It is about two-thirds the length of the stem.
Before we proceed to discuss the homologies of these organs
it is absolutely necessary to grasp their anatomy and
relationships.
Let us now consider the homologies of these three
branches of the stem. The first I have already identi-
fied as the inner root {innere Wurzel of Kiihnle). This
identification rests on its relations to other organs.
The ascending trunk is an organ for which I find no
parallel in any insect brain yet investigated. This b not
very remarkable when we remember that the brains of no
Lepi(kmleron or Truthajpfermt have yet been fully described.
According to a view which I now put forward the ascend-
ing trunk is to be regarded as the united base of the
forward root (vordere Wund) and of the backward root
[riicklaujige Wurzd). Great obscurity exists with regard to
* I nae the term " trunk " and " branch " rather than " root "
in ofder not to commit myself to way view as to homologies which
are folly discuaaed later. The terms are of a provisional nature.
It may be objected that I am adding to the synonymy, but it ia
almost a necessity to have some unequivocal name for an organ
until Um homolc^ies oie fixed. The word " root " I use as a full
equivalent of the German Wvrstl.
iM:,i,A J 1^, Google
128
Mr. P. A. Buzton on the
the homologiea of the ascending and posterior branches of
the ascending trunk.
It is probably best to consider the ascending branch as
forward root {vordere Wurzd, Kiihnle ; tuberade anUrieur,
Tkxt nouBS 2. — Oatline dnwingB of the right mushroom body.
A, seen from the OQt«r side (lateral view). B, &om above. C, from
in front. (The stem and head are behind and not sbown in this
drawing.) The line aa repKsenla the median (sagittal) pluis.
The CWA are not shown ; the whole organ as drawn here consiita
of azonio substance.
at. br. ascending branch, eu. Ir. ascending trunk, hd. head of
mushroom body. in.r. inner root, po.br. posterior branch, tt.
stem. «u>. M swollen head of muebroom body. sut. el. swollen foot
Ij.y.l.Ajl^yCOO'^lL'
Prolixerebrum c^ Micropteryx. 129
Tkxt nauBE 3. — Outline drawing of six longitudinal vertical
Motions (A-F), to show tbe relations of the parts of the mushroom
body to surrounding pc. L, the last being nearest the middle line.
Tbe six sections are not consecutive. Dorsal is to the left; anterior
(cephnUd) towards the top of the page. Only the asonio parts are
here shown, the cells being omittod.
a. tie nerve fibres of troot a (p. 136) and other tracts, passing
from protocerebrum to lower parts of brain, tu, br. ascending
branch, (u. Ir, ascending trunk, b. tract b. br. bridge, d. i.
donal lobe. ti. Einstrommnng (p. 120). Ad. head of mttshroom
body. tit. ea. inner capsule of centrt^ body. tn. r. inner root of
ffiDshroom body, oc gt ocellary glomerulus, oe, n. ocelUry nerve.
OK. ea. outer capanle. pc I. protocerebral lobes, po. br. posterior
brsocb- A stem. mo. nd. swollen head of mushroom body. «c st.
swollen foot of stem. vL I. ventrolateral lohea.
TRANS. BUT. 80C. LOND. 1917. — PABT I. (NOV,
j^, Google
130 Hr. P. A. Buxton on the
Viallanes). This is suggested by its genem! direction and
by the fact that it ends on the surface of the fibrillar part
of the brain, under a thin portion of the ganglion cell layer.
With this we may compare the similar " free " ending of
the tuberciile anUrieur in Acridians (Viallanes, 1887, p. 42,
fig. 46), and of the vordere Wurzel in Apis (Jonescu, p. 137,
Text &g. 10a) and in Tespa {Viallanea, 1886). The author
remarks : " La premiere se parte directement en avarU pour
gagner la surface antirieure du renjlemerU primaire ;' c'est
la come anUrieure"
If, then, the ascending branch is the equivalent of the
vordere Wurzel, we must homologise the posterior branch
with the rucklaujige Wtirzel, or backward root, in considera-
tion of its backward direction and deeply buried termination :
this would probably be accepted were it not that Eiibnie
has asserted that the tubercule anUrieur of Viallanes is the
homologue of the vordere Wun^H and also of the rUcidauJige
Wurzel. Against this I must enter a most emphatic prot«st.
In the first place, there ia an inherent improbability about
it ; we cannot willingly believe that vordere and anUrieur
refer to an organ which is described in other insects as
"running back" {rucMauJige). We surely need good
evidence before we can accept such a statement? In the
face of that improbability Klihnle was dangerously bold in
asserting the homology. When a great many more types
have been investigated we shall be able to bridge many of
the gaps which at present exist in our knowledge. Till
then we can none of us be certain of any but the most
obvious homologies, partly because the described types
are so few, partly because no living man has first-hand
knowledge of more than half a dozen insect brains.
Kiihnle was probably led into this error by the fact that
the majority of insect brains show only two roots, some of
them lacking the forward, others the backward root.
If, however, EUhnle's homology be accepted, the one
which I have suggested must fall; for clearly I cannot
give the terms vordere and rucMaufige to two structures if,
as Kiihnle says, they are in this case synonymous. I do
not wish to press my own convictions unduly ; but at any
rate they are based on considerations of relative position,
that is to say on actual fact.
If, then, Kuhnle is right and I am wrong, we may either
assume that the ascending trunk and its branches (ascend-
ing and posterior) together form the backward {rucklaufige.
ProUx^rebrum cf Micropteryx. 131
vordere) root, or else that the poaterior branch represents
that oi^tiD and that the ascending branch is a new organ.
The first of these views is supported by the fact that the
ascending and posterior branches leave the stem by a com-
mon origin, the ascending trunk. On the other hand, so
far as our knowledge yet goes there is no otlier brain in
which the backwaid root is bifurcated, which b what this
view impUes.
As I have said, it is also possible to regard the posterior
branch as the rCcldaufige Wund {vordere Wnnel, tubercule
aniirieur) and the ascending root as an organ which cannot
be homologised with anything yet described. This is quite
a rational view to adopt, for practically nothing is yet known
abotlt the brain of the Lepidoptera. Personally I do not
see any necessity for dubbing this a new organ, but if
Kijhnle's identification of the vordere with the TiicBauJige
WuTzel is proved correct, then we shall probably find it
necessary to find a name for what I have provisionally called
the ascending branch. An investigation of other types
might yield most fruitful results.
This very small and abstruse point must be settled defi-
nitely before the study of insect brains has gone further.
Unfortunately it is not possible to attack the question from
a comparative standpoint, but it is essential that we should
atart work with our homologies correctly and clearly defined.
There can be no compromise between Kuhnle's view and
my own, and the point at issue is fundamental. I consider
thAt there is, at any rate, very Uttle reason for doubting
my identification of the innwe Wurzel, which agrees with
the views of previous writers.
These conflicting views may be expressed thus : accord-
ing to my view, ascending trunk = common origin of —
1. Ascending branch {vordere Wurzd, etc.)
= Forward root.
2. Posterior branch {rOcklaujige Wurzel)
= Backward root.
Eiihnle, however, asserts that riichlaujige WuTzd=vordere
WvTzel. If this is so, then either —
(i) Ascending trunk and ascending branch and posterior
branch tt^ther = rucklanfge Wurzel, or else —
(ii) Ascending trunk = common origin of (a) posterior
bnuich {riicklaufige Wurzd) and (&) ascending branch
(not homolc^us with anything yet described).
iM-,i,A J h, Google
:C Ifr. p. A. Buxton on tMe
I Safirc* xbaa the dunctos by vhich tbe three roots
su' W jvetOKvd mn these : The inner root (in. r. ; innere
Wtrst^ S^siijf: ttAtnsle itderne, Dujaidin; Balken,
Vijt^' nsB Wlrvxrds and inwards and terminates
iwrwuii u» ^jiale lobe and the inner capsule of the
.ft:~3K! K-«^. ks cod is adjacent to that of its fellow on
^M .'onwr-f sue. It appeus that this root is found in
WKT' ^ :3w<Rs. TW (oHmd root (as. br. ; vordere TFurzel,
"> -JweCT. I'll liii i* K. FlQ^ ; anterior root, Eenyon ;
•■.>^~h< gmit^mr. I>c;>aidin) runs forwards and upwards
v-i twas ' r:w " .-c tiie saiboe ot the ptotocerebial lobes,
- > ~ Rtate ;i>F arur^irc tcOs ax else duectly beneath the
•wj~.«a:Ti«. TV >ttd:van] root (po, br.; rwHaufige
I '!!»'. Sij:Li.w rc» faactrods and terminates in the
"s-s;.',-.'- .- !•?- .-£ li* |wcooei»tgum without ever reaching
•w -«.-ak'», 1: Bar.r hsects either the forward or the
^itsrcK^ T.v*- s ajtiin:.: ; tlis has W Eiihnte to believe
. -i' .i..r :.«» - ■ -ihe :iJ» tm^ oppoitonitT of answeiing
■-? .-wt-- " v'l.t.-'iv^ZTn-^UT be made to the hypothesis
■ - :*» r.,tt«trviii K'^ST is hs tjpical development pos-
■cNM^ •■•>•» 'vfls, h M weD knowB from Eenyon's work
*■- ;^ ^ nw~h>£.iLT^fannaftlie bee, that the axons
* ■ : .- .■ -vw :iw ^cMi }kbdc^ dicltottHnously, and that
-. • '.-I. ;i-> > im ;*ietwv toots of the mushroom body
■ ■ .^ --«• \.* - may be craed that this division of
^ x. ,- »- , » :.>.-i u revtiaKy diaiacteiistic of the
t*— • -!-.•- .ir :.-. c»,>M i:: «oenl. would find its
■«.'4.-. , ^■.'-ss-..: .:. a i:n^j\xd body with two roots.
.*.,-- ^ If !■- »TiT tiat ii>e»e is no diGBcidty in
-- -ki.>^ „ -*. »<fc : : :ir» jtf t: irjies snpjdies two of the
>^ <~% .^. ii~»-. K* fc T&'f :^ dimity lonains whether
1 «*.-•-. . BrjL x-K-fcvc fcx=x-i>es be M»e root or two,
'. ■ ■, - s*,-s- .1- ^!•.T «aT.-cr.T «*a hardly be disputed.
- •i-- . .-V »-, wn fcjnw.vr iiT,.L:.ar wi:h tie division of
■1 *■, ,r^ -.1 :jw ;iTi.;-t> .c .vJKC 2SK^: DOW if the fibres
> ■ , ^v i.v» ( ' vv .■*! iw rr.-cppd in faich » way as to
-■■. .-. 1 r..-.'--*i ii 1 -iwT,xc 'riy jJn-Hild not the fibres
- -.- - V < , ■ .-,■«*.■ » •■ stToiy a}ii»8t any number of
• ^ V-,»» ■>■ \i' ti i>tt» jOMk^ described the mnsh-
sv' \ t ^.ns.!. '.-. «u-. ,1 a TwaJte- both of which
.M^ >% .>^ ■< tivi ■« ifcivc K> trasp the bearing of
~ ?«- ..-.V ;.'i. ^n«>Tu 0W(!O.-e. <f hecDoiogy. The fact
>. -aH.ivtw '■ ;»«e Tons T^ de ■nichiooin body is
ih, Google
Protocerdnntm <^ Micropteryx. 133
not, then, a new discovery, but I trust that I have been
enabled to put the homologies of the matter on a sound
basis.
IV. The Central Body.
The central body of Mieropteryx consists of two capsules ;
the lai^ei of these, the outer {ou. ca.), is superior and
posterior, the smaller or inner capsule {in. ca.) is inferior
and anterior. They are respectively the dussere SchaU
and innere Schale of EiUmle. They lie together in that
space in the middle of the protocerebral lobes to which
Viallanes gave the name la loge; this contains also the
ocellaryglomeruliand the inner roots of the mushroom body
(PI. VIII, figs. 7-10). The space is bounded on all sides by
the protocerebral lobes, and above by the tract/, in front
by the tract c, behind by the tract d, and below by the
double tract e (see p. 137).
Micropteryx is one of those insects in which the central
body is large and the mushroom body comparatively small ;
that is to say, it falls within one of Bretschneidet's lower
The outer capsule is slightly wider than the inner. The
anterior edge of each is in the same vertical plane, but the
outer extends back a considerable distance behind the inner,
and this posterior part of it is very thick; thus the outer
capsule overlaps the inner above and behind and is much
the more bulky of the two. This condition is character-
istic of nearly all the insect biains which have yet been
described. Turning to internal structure we find that the
outer capsule stains rather more deeply with eosin or
oTange G than do the protocerebral lobes. There is no
definite division of either capsule into bodies like the rays
of a fan, a condition which has been described in the brains
of various insects since the time of Dietl. The anterior
part of the inner capsule is, however, divided into a number
■ol small rounded masses arranged in no definite manner
and separated from one another by bands of axons, the
great majority of which pass into the outer capsule. These
maases resemble to some extent the glomerular bodies
{Faaerb&icken) of the antennary lobe. The 8catt«red
neuroglia cells which lie in the space which surrounds the
central body are referred to elsewhere (p. 120). There is
no group of cells which can be said to belong to the central
body either here or in any other insect, and we believe that
u,y,l,/eJh,G00glc
134 Mr. P. A. Buxton on the
the oigan is a reflex centre not connected with any one
motor or aenaory function.
(The ocellary glomeruli, which are sometimes conBtdered
with the central body, are described on p. 135.)
V. The Bridge.
(Die Himbriicke — Kilhnle, etc. Le Pont — Viallanes.
Fibrillar Arch— Kenyon.)
The Bridge (br.) ia a protocerebral structure found in all
insect brains. In Microplerux it occupies its usual position
as a transverse band of axonic material on the superior side
of the prot^cerebrum. It lies behind the nerve to the
ocellus, and between the lieads of the mushroom bodies
(PI. VIII, IX and X).
The bridfre consists in part of Punktsubatanz ; this is
found at either end. The middle of the bridge is much
attenuated and appears to consist entirely of nerve fibres
passing from one aide to the other. The oi^an, then, is
dumb-bell shaped, the swollen ends, or " heads," being
preaumably centres, the narrow waist a decussating tract.
The bridge is covered by some layers of ganglion cella,
which appear to belong to the normal type.*
The axona of these apparently " normal " cells pass down-
wards to several parts of the brain. Some which proceed
from the more lateral celb pass in front of, or behind, or
round the end of the bridge, and enter the dorsal surface
of the protocerebral lobes in a diffuse manner. Other
axona pass through the heads of the bridge and so on-
wards to the lobes of the protocerebrum ; it ia probable that
these give off a collateral while they are within the aub-
atance of the bridge. Other cells, again, send their axons
into the bridge it«elf, where the fibre is lost to sight. It
is probable that some of these fibres cross the middle line.
The whole matter requires investi£;ation by the Golgi
method. We have, then, a number of similar cella, some of
which appear to be associated with the bridge, some with
the dorsal part of the protocerebrum, some, again, with both.
From this I am inclined to aigue that the bridge is of less
importance as a physiological entity than some authorities
have believed; that it is rather of anatomical than of
* In Forficula Ktihnle distinguiBbes several types of cell in this
region, but in every respect the ganglion cells of Micropttryx appear
to have \ery little tendency to be mSeientiated into types.
l„y,l,Ajh,COOl^lC
ProtoeerAntm c^ MicropUryx. 135
phyBiological significance. Against this view is the ad-
mitted fact that the bridge exists as a distinct structuie in
all insect brains which have been investigated.
I can find no visible connection between the bridge and
the nerves auppljring the compound ejes, though a few
fibres of the ocellaiy nerve enter the ends of the bridge.
This perhaps supports Kiihnle and tends to contradict the
contention of Bretschneider and others who regard the
bridge as a centre for the co-ordination of visual impulses.
VI. The Visual Centres.
A. The Ocdiary Apparalut.
In MicTOpteryx paired ocelli are present, but the median
ocellus is not developed here, or in anv other Lepidopteron
or Trichopteron. A stream of fibres, the ocellary nerve
{oc. n.), leaves the back of the spherical chitinoua capsule
in which the ocellus is contained. At the point where the
fibres leave the capsule there is some tendency for the nerve
to break, as it is very much narrowed. The sensory cells
are contained partly in the capsule of the organ, and some
of them lie along the course of the nerve away from the
actual ocellus, and as the nerve proceeds inwards they
become less and leas numerous. The nerve runs straight to-
wards the middle line in a plane slightly anterior to the bead
of the mushroom body (PI. VIIT, fip. 10 and 11). When
it is over the external edge of the central body it bends
backwards, and at this point a few fibres leave it to pass
into the protocerebral lobes. From here it passes backwards
and inwards and continually gives ofi more and more of its
fibres, so that though there is no point at which the ocellary
nerve as a whole passes into the substance of the proto-
cerebral lobes yet the whole nerve ultimately does so. A
few fibres may also be seen to pass into the swollen head
of the bridge.
Two small spherical bodies with rather indefinite margins
are found in the space beneath the outer capsule of the
mnahroom body, posterior to the inner capsule and to
the middle lobe (PI. VIII, fig. 10) ; these are the ocellary
glomeruli (oc. gl.) or " tubemdea du corps cerUral" (Viai-
lanes). In Micropleryx I have been unable to demonstrate
the connection between these structures and the ocellar?
nerve, owing to the diffuse way in which the fibres of the
nerve pass thioogh the dorsal part of the protocerebnun.
136 Mi. F. a. Buxton on the
Fiom a consideration of what is known about similai organs
in other insects I do not consider that much doubt exists
as to the functional connection between these small, round,
deeply placed lobes and the ocellus. It is hardly to be ex-
pected that we should be able to find more than a propor-
tion of the smaller tracts in so minute a structure as the
brain with which we are deahng. Fibres pass from the
ocellary glomerulus to mapy parts of the brain, including
the paired (deuteroeerebral) 83Tnpathetic system (tract «)
and the posterior part of the antennary lobe {tract ()■
B. The Optic Lobes.
Pressure of other work has absolutely prevented my
devoting attention to the optic lobes {o. I.), and much special
study of the literature would be required before I could
hope to treat of them at all adequately; this would
delay the publication of this paper almost indefinitely,
and I have accordingly decided to leave them entirely
undescribed rather than to deal with them in an inadequate
VII, The Fbotocerebbal Tracts.
Authors have frequently attempted to enumerate the
tracts of fibres which connect one part of the brain with
other parts, but they can only detect and describe the larger
bundles and can never even attempt to follow the minute
tracts which ramify in all directions through the Fnnkt-
substanz. Such an enumeration must always be incom-
plete even if one part of the brain is proved to have a score
of paths connecting it with other regions : and even if the
Golgi method is apphed to a very large number of individ-
uals, definitive completeness can hardly be arrived at. In
this present paper I make no attempt to give any complete
list : I only describe a few of the more noticeable tracts
which are useful either as landmarks or else as definite
boundaries to regions.
The following tracts should perhaps be noticed, because
they are important features of the sections in which they
occur, (a) A nnde tract of fibres arising from cells situated
over the dorsal part of the protocerebrum, and passmg
vertically downwards in the middle line over the fiwnt of
the protocerebral lobes : this tract forks below and the
fibres then pass to the antennary lobe of each side, and
ihyCOOl^lC
Prolocer^fTwn of Micropteryx. 137
poesibly also to the tritoceTebnmi and the ventral parte of
the ceotral nervoxis system (PI. VII, figs. 2-4).
(b) A tract of fibres which is the Riechstrang, or Riech-
bafin, of the Gennan autboiB. The cells from which the
tract arises appear to be ganglion cells of the " noimal "
tvpe. and are situated dorsal to the protocerebral lobes
and slightly behind the bead of the mushroom body : the
fibres pass downwards and forwards and slightly inwards,
through that part of the protocerebral lobes which surrounds
the space contMning the central body, etc, ; from here the
fibres are directed downwards and outwards to the anten-
nary lobe. Through most of their course the fibres lie
in a free space. Owing to its diffuse structure, which ren-
ders it difficult to distinguish it when it is cut transversely,
the lower part of this tract can only be distinguished in
longitudinal section (text fig. 3, D and E, p. 129).
(c) A broad anterior commissure (PI. VII, fig. 6).
{d) A deep commissure.
(e) A double ventral commissure (the vordere und hinlera
Brucken der Nebatlappen utUer sick of Kuhnle; PI.
VllI, fig. 9), These three are the most definite tracts
which unite the two sides of the protocerebnim, though it
should be remembered that the lobes are united over the
greater part of their internal aspects, and that a large
number of smaller tracts pass from one side to the other.
These three tracts fonn the boundary of the " loge" in
front, behind and below : the anterior and ventral ones lie
upon the surface of the fibrillar part of the brain, the deep
commissure passes between the two protocerebral lobes
behind the " loge " : the ventral commissure (e) arises on
each side from the anterior part of the ventrolateral lobe.
(/) A small but distinct band of fibres which runs trans-
versely across the upper surface of the " hge." This tract
is the Faaerhofoi Kuhnle, and possibly also the commessvra
protocerdtrale anleriore of Berlese (PI. VIII, fig. 8).
(y) A posterior commissure uniting the two protocerebral
lobes at their most posterior part, where thev overlap the
tritocerebrum (PI. IX, fig. 16).
(h and i) Tracts uniting respectively the anterior part
of the middle lobe to the anterior part of the ventro-
lateral lobe, and the posterior part of the middle lobe to the
posterior part of the ventrolateral lobe of the same side.
(j) A tract running from the posterior part of the middle
lobe to the deuterocerebrum.
u,y,l,/eJh,G00glc
138 Mi. P. A. Buxton on tie
(k) A tract mimmg up from the middle lobe into the
deeper parts of the ptotocerebral lobea which lie lateral to
the " loge."
(l-g) Tracts from or to the central body.
[l] The two capsules of the central body are united
especially along their anterior margin bj fibres which make
the partitions between tlie " Facher " of the inner capsule
(Pl.VIII, fig. 8).
(m and mm) Both capsules are united to the protocerebral
lobes by bands of fibres which leave their anterolateral
margins; the band from the upper capsule (m) passes
upwards, that from the lower [mm) downwatxls to the
ventrolateral lobes (PI. VIII, &g. 7).
(n) The outer capsule receives fibres which leave or
possibly enter the head of the mushroom body on its under
side. This is really a part of tract r.
(o) A few fibres connect the outer capsule to the bridge.
(p) A few also pass from the ocellary nerve to the outer
capsule.
[a) A well-marked band connects the anteonary lobe
with the outer capsule. This band enters that part of
the outer capsule which lies immediately superior to the
posterior part of the inner capsule (PI. VIII, fig. 10).
(r) The tract which has been mentioned on p. 126 as leav-
ing the inner and inferior aspect of the head of the mush-
room body, and passing partly to the outer capsule of the
central body (tract n), but mainly to the deep part of the
protocerebral lobes lateral and posterior to the " loge "
(PI. VIII, fig. 12 ; PI. IX, fig. 13 ; PI. X).
(a and I) These tracts pass from the ocellary glomerulus
to the paired deuterocerebral sympathetic system and to
the posterior portion of the antennary lobe respectively.
(u) This consists of a few fibres which pass down from
the swollen ends of the bridge to the dorsal lobe imme-
diately below and to the tumulus (Plate X).
It is, I believe, generally true that pured organs are
united across the middle line, but I am unable to say
whether this is invariably the case.
Summary.
The protocerebrum of Micropteryx might be described
in the following terms. The neurilemma, which covers
the whole central nervous system in one continuous sheet.
Prolocer^mim <^ Microfteryx. 139
is a thin B3aiC7tium, and bsneath it are found the ganglion
cells and the axonic parts of the nervous syetem. Over the
protoceTebrum the layer of ganglion cells ia deep, and four
types can be distinguished : the normal type, the mushroom
body cells, the cells of the optic lobes, and the giant cells.
Neuroglia cells are found in the substance of the protocere-
bnim in amall numbers. The tracheal system of the
brain is very sightly developed. The protocerebral lobes
are large, and in volume greatly exceed the other parts
of the protocerebrum t<^ether. The various parts of the
protocerebral lobes which have been described in other
insects are all present, though Mie^opteryx presents some
peculiarities, for the ventrolateral lobe and the middle lobe
are each divided into anterior and posterior portions. A
mid-dorsal lobe b also present, and to this I have given the
name tumulus, an organ which has not been described
before. The mushroom bodies are of a small, simple type,
and only one is developed on each side : the head is remark--
able because of the shape, which M that of a rough sphere,
wUhout any approach to theformali<m of a cup. In section
it is seen to contain minute glomerular masses of nerve
fibres, which are regarded as association centres : these
are comparable to similar structures described in the mush-
room bodies of many insects, and also in antennary lobes
and central bodies. The origin of the stem ia below, not
within, the head of the mushroom body, and it runs down-
wards and forwards in a definite space; it ie rod-like,
and not perforated by a canal. The stem divides below
in a complicated manner which does not lend itself to
summarisation. I have suggested several possible homo-
Ic^es for the parts into which the stem divides, and
my own view is that there are three rooU to the mushroom
body in this insect — an inner, a forward, and a backimrd —
aiid that this is the typcal number for the insect brain :
other views are also discussed. I have also given what I
betieve to be the normal relations and characters of
tbeae roots ; and I beUeve that thb part, at any rate, of
my paper has some permanent value. The central body
is Urirge, and consists of two capsules, as usual ; the outer
is the larger. There is no tendency tomards the division
oj either capsule in a fanlike manner, but the inner capsule
contains a number of minute glomerular bodies. The tracts
passing from or to the central body are numerous and some
of them are large. The nerves from the ocelli run inwards
iM:,i,A J 1^, Google
140 Hr. P. A. Buxton on tJie
ociosB the froDt of the bead of the miishioom body and
paaa giadnaUy into the sntstonce of the pTotocerebial
lobes, and a few fibres pass into the bridge. Two smitll
bodies are found beneath the central body, and thcee aie
preeomed to be the ocellary glomeruli of other antboie,
though in the brain of Microptaryx there is no actual evi-
dence of their connection with the oceDary nerve. The
bridge is simple and straight; its ends are rounded and
consist of Punktsubstanz, and into these pass the axons of
a few celk which are situated in the immediate neighbour-
hood ; the middle of the bridge iti formed of a large number
of fibres which pass across the middle line. (I have undei-
lined those characters which appear to indicate that the
brain of Micropteryx belongs to a simple type, bo far as
morphological points are concerned.)
It would doubtless be interesting to compare the simple
brain of this Frotolepidopteron with that of other Lepido-
plera or Trichoptera. This is, however, impossible, except
to a veiy slight degree, because the only work to which we
can refer is the classic paper which Fliigel published in 1878,
and a few lines in Berlese'a text-book. Flogel dealt with the
brains of a numbet of larvae and imagines of Lepidoptera,
and his fullest description is that of the biain of the imago
of Gossus. He devotes his attention to the mushroom body,
which differs from that of Micropteryx in several important
particulars. The head is developed as two cups on each
side, placed in apposition to one another. The two stems
which proceed downwards from these unite to form a
single cylindrical stem which stains deeply and lies in a
space. An inner root is given off, and this occupies tbe
usual position of that organ ; there is also a forward root
which runs up to the surface of the brain and there divides
in a complicated manner which is not further described;
no backward root is described, but it is possible that this ia
represented by one of the branches of the forward root.
This suggestion is an attempt to bring Cossus into line with
Micropteryx, and it may well be correct, for we must remem-
ber that Flogel was hampered by the deficient methods
of bis time, and that he was the earliest insect neurologist
in any true sense of the word,
Berlese describes the brain of Sphinx very shortly. The
protocerebral lobes are large, the mushroom bodies of
moderate size; two pairs are present, which lie one in
front of the other : their stems do not unite. A mass of
l„y,l,Ajh,COOl^lc
Profocerebrum oj Mtcropteryx. 141
very large cells (c«U»I« maeatre) are developed behind and
above the protocerebmm, and tbe fibres from these proceed
over the front of the protocerebmm to the ventral brain
by way of the aeaophageal connections.
TECHNIQUE.
I. Fixation asid Ihfbegkation.
My early work on Micropleryx was all done upon material
which had been fixed and stained by very simple methods.
I became convinced that for insect neurology the employ-
ment of complicated technique was not only desirable
bot necessary. Accordingly I devoted the early spring of
1915 to a somewhat extensive senes of experiments in
staining and fixing the brains of cockroaches (Periplaneta),
my object being to familiarise myself with some forms of
technique which I proposed to apply later to Micropteryx.
I shall describe my methods for both insects together,
though some of them are only applicable to one or other
of the insects.
Owing to the chitinous cuticle of insects it is necessary
to take every care to ensure the penetration of the fixing fluid.
Unless there is good reason to the contrary Micropteryx
should be cut in two with a sharp knife ; only the anterior
end will be preserved and fixed. A cockroach, on the other
hand, should be chloroformed and held between the finger
and thumb, with the head resting on the thumb-nail ; the
epicranium should then be punctured with a small sharp
Imife, and also the eyes if the individual is a lai^e one ;
the same knife should then be used to remove all the mouth
parts and the labrum at one transverse sweep, the thumb-
nail forming a block on which the cutting is done. All this
can be performed without any damage being done to the
brain by pressure. The head is then cut off and placed in
the fixing fluid.
Fixatives. OsHic Acid (osmium tetroxide). — This is
perhaps the most generally used of all fixatives, ever since
the time of Viallanes, who described it as " le riactif le
plus pricieutc que nous possidions pour mettre en ividence
le trajet dea fines." It has been used in strengths of from
i% to 1 %■ Flemming's solution baa also been much
used, and it is probable that its results are slightly better
than those given by osmic acid alone. Bottger recom-
mended its employment for periods of about three weeks ;
iM-,i,A J h, Google
142 Mr. P. A. Buxton on tie
I cannot see that anything is gained by leaving material
in it for so long a time, thou^ it is well known that all
fixatives containing osmic acid penetrate slowly even
through small pieces of tissue. Forty-eight hours is quite
sufflcieot, according to my experience, Borrel'a fluid also
gives good results very similar to those obtained with other
osmic acid fluids. Micropteryx tends to float in this and
other fixatives ; if it cannot be caused to sink with the ud
of shaking it may be lightly painted with 90 % alcohol in
order to reduce the surface tension. All these fixatives
are extremely useful, though they occasionally tend to
shrink the cytoplasm of the la^r nerve cells. The nerve
fibres (axons) stand out from one another with great clear-
ness, and in this way the sections ate well suit«d for study :
they are never distorted, and there is no tendency for
the ganglion cells to break away in masses from the under-
l3^g fibrillar substance. Great care must be exenused
in washing the material veiy thoroughly in water after
fixation, or the staining will be unsatisfactory.
FoBUALiN.— Formalin has been recommended in various
rather high percentages (10 %, 20 %, etc.) by more than one
worker. It is customary to leave the heads in it for some
days. I anticipate that the use of formalin will soon be
discontinued, for though it gives a distinctly good denoon-
stration of the tracts of axons, there is a great tendency for
the formation of vacuoles in the fibrillar substance. The re-
sult of this is that the tracts are pushed to one side and
distorted. This vacuoUaation is not invariable, but it
constitutes a grave defect in the method, which is one that
I found unreliable.
Picro-Chlor- Acetic Mixtobb. — I do not know to
whom we are indebted for this very useful fixative; but
it appears that it has not previously been used by insect
neurologists. My own experience ia that it is the best
general fixative I have ever employed, and I trust that the
workers of the future will be as satisfied with it as I am
myself. It possesses very great powers of penetration,
and can be relied on to fix small insects completely without
decapitation or any other precautions. Inaect histologiats
will find that it is an exceedingly fine preservative of the
details of cell-structure; as far as the brain is concerned
this fluid demonstrat«d the tracts of axons with particular
clearness, and in this respect it does not fall far short of
osmic acid. The nerve cells are also well preserved, and
l„y,l,Ajh,COOl^lC
ProtocerArutn of Microptenfx. 143
all the different types can easily be distinguished, though
for a special study of the cells it is certainly best to have
some material fized for that purpose in Bouin's fluid.
Bouin's Fluid.— This fixative is only of use for a study
of the nerve celb, and for this purpose it ie unrivalled. It
fixes material in such a way that the tracts of fibres cannot
be distinguished at all, but that is immaterial provided it is
realised that the fluid is essentially a special fixative.
AcBTic SuBUHATE SOLUTION.— This is Simply a satu-
rated solution of mercuric chloride in dilute alcohol to which
a small percentage of acetic acid has been added. It has
been used by other authors but there is nothing to
recommend it ; the tracts or bundles of axons are shown in
much the same way as they are in material fixed in the
picn>-chlor-acetic mixture, the cells are shrunken and the
di£Ferent types cannot be distinguished, and the fluid has
poor power of penetration.
PEEEann's Fluid. — This is a fixative with very small
power of penetration, even when used hot. Even if penetra-
tion is secured the tracts of axons cannot be distinguished
from one another, and the cells ate swollen and matted
together.
Gilson's Fluid. — The penetrating power of this fluid
is so great that insects may be fixed in it whole. It is an
exceUeat fixative of ganglion cells, and shows the differences
between the types very clearly : tor this purpose it is
valuable but it fails entirely to define the axons.
BiCHBOMATE. — Potassium bichromate, apart from its
use in the Golgi method, is quite useless as a flxative of
insect nerve tissue, first because details of structure and
the course of axons are not well preserved, secondly because
material so fixed stains most intensely and generally with
the aniline dyes, thirdly because these stains can scarcely
be washed out or difTerentiat«d, and fourthly because of the
great brittleness of sections which have been exposed to
the action of these fluids.
The Golgi Meh^od. — This method has been appUed
by Kenyon to the brain of the bee, and with it be has ob-
tained some very renuirkable results; his original paper
(Kenyon, 1896, I) should be consulted for a full account
of bis procedure. It is almost impossible to apply it to
insects which are not available for the greater part of the
year because it is extremely precarious, and even Kenyon
himself only obtained good results with an occasional
iM-,i,A J h, Google
144 Hr. F. A. BuxtoD on the
qKcimen. I Iiare been unable to use it with any success
upon Micropteryx.
Mbtaluc bfPBEQSATioN.— The object of this method
is to impregnate nerve cells and fibres with actnal metallic
ulver and gold. The silver salt which is invariably used
is the nitrate, and as it gives ezceUent results I have tried
no other salts. The beads are dropped into a solution of
this salt and kept in the dark for a period. I have devoted
some time to discovering the best strength of silver solution
and the period during which the heads shoold be exposed
to its action. I find that the best reanlta are obtained
by dropping them into 1 % silver nitrate in water, and leav-
ing them in the dark for ten days. The ralver is very slow
in penetrating the head, and if a 6 % solution is used there
is great danger that the periphery will be blackened before
the central portions are aSected at all. I believe that
penetration can be accelerated by keeping the whole at
aO'-SS" C. It is probable that the period during which the
head lies in AgNOg is immaterial provided that the fixation
proceeds in the dark and that sufficient time is allowed for
the full and equal penetration of the silver. It was not
found advisable to assist the silver to penetrate more quickly
by employing an alcoholic solution of the salt. I have, for
instance, experimented with a 1 % solution in 30 % alcohol,
following this by the various processes which I describe
below. The impregnation of the various fibres was not
obtained at all, though the various parts of the bnun were
coloured to diSerent degrees. In fact, the alcoholic solu-
tion of silver gave quite a pretty differential stain of no
particular value, but failed utterly to produce the sweeping
black lines which are what is desired.
The head, then, is fixed for ten days in I % silver nitrate
in darkness. It is then washed. A few workeis transfer
it to pyrogallic acid for a day, in order to reduce the silver
and leave it in the tissues in a finely divided state. I am
quite convinced that this is unwise. The reduction may be
done much more evenly by a method which I shall now de-
scribe. The heads are embedded, unreduced, in paraffin,
fixed to the slide in the usual manner, and treated with
xj'lol and descending grades of alcohol. At this stage the
sections are sienna-brown in colour. From "a low grade of
alcohol the sections are moved to distilled water. (I need
hardly say that if the beads or sections are brought into tap
water a fine deposit of chloride will be precipitated which
Prolocer^rum oj Micropttryx. 145
will completely ruin the preparations.) The slides are now
placed in 1-2 % AgNOj and exposed to bright sunlight or
an electric lamp for about ten minutes. After this they
are washed for two minutes in distilled water and placed in
1 % gold chloride for two minutes in a bright light. They
are then again washed and placed in an aqueous solution of
pyrogalUc acid until the reduction is complete, deposits of
metal being left in the fibres. The sections are now brought
up through the usual grades of alcohol, stained for a very
few seconds in orange G, and mounted. These prepara-
tions do not degenerate under the cover-slip in the same
manner as Golgi preparations.
This impregnation is only a modification of one intro-
duced by Ramon y Cajal; a similar method has been
employed by Jonescu.
li it is successful it gives sweeping black lines of axons
running through the brain in the most diagrammatic
manner. It is unfortunately almost inapplicable to sb
small an insect as Mu^opteryx, owing to the fact that the
aqueous silver solution hardly penetrates the insect's
minute neck even after decapitation. I am quite confident
that this method will be found most useful in the study of
the brains of insects which are large enough to admit of
the brain being laid partly bare to the fixing fluid.
II. Section Cutting.
All material should be stored in 90% alcohol, rather than
in a lower percent^e. Excellent material may be com-
pletely ruined if the spirit in which it is kept has ever been
in contact with cork, the tannin of which interferes with
the action of mrist stains : glass-stoppered vessels must
accordingly be used.
Section Cutting, — It is well known that the cutting of
sections through heavily chitinous insects presents great
difficulties. Much may be done to overeome this, but be-
fore discussing methods of softening chitin I should like
to state my firm conviction that the one factor of prime
importance is the microtome knife. In the absence of a
really sharp knife no softening reagents and no care exer-
cised during the euibedding are of the slightest value. The
best softening reagent, so far as my limited knowledge goes,
is spirit soap (German Pharmacopoeia), the use of which was
first advocated by Kurt Bedau. The insects are placed in
TRAHS. BNT. SOC. LOND. 1917.— PART I. (NOV.) L
ih, Google
146 Mr. P. A. BnztoD on the
this for some days, well washed in 70 % and 90 % alcohol,
and then embedded. The chitin is certainly softened by
this solution, but will regain some of its hardness if it is
simply embedded in hot paraffin. It appears that heat,
absolutfi alcohol, and xylol, all ezeicise a marked harden-
ing effect on chitin. It is beat, then, that the head or
insect should be dehydrated as much as possible in 90%
alcohol, left a short time in absolute alcohol, and.cleared in
chloroform. I then place it for some days in a chloroform
solution of paraffin, and finally drop it into the hot parafBn
of the ordmary embedding bath. Here it remains only
long enough for it to attain the temperature of the bath,
and is then removed in the crucible or other vessel in which
the paraffin is contained, and placed under a vacuum pump ;
the pump nill quickly remove the chloroform, most of
which has by now become diffused into the paraf&n. The
mass may then be turned out into a mould and cooled.
Terpineol has also been used as a softening reagent and
it appears quite satisfactory, though I have not much
experience of it.
Celloidin, — I have used this to some extent, though
I no longer do so, because I find it unnecessary if spirit
soap is used as described above. It cannot be relied upon
to penetrate a whole insect unless thin celloidin be employed
for many days.
Both with and without celloidin I have been able to
obtain serial sections of the head of Micropteryx of con-
siderable thinness. I have several series of 35 n, which is
not by any means too thin, because of the smallness and
complex structure of the brain.
Practical experience teaches me that it is never safe to
move either complete brains or sections from absolute
alcohol to x}'lol or vice versa, but that an intermediate
mixture of the fluids should always be employed. Unless
this is done the ganglion cells will frequently break away
from the axonic part of the brain.
III. Staining.
For general study sections should be stAined with Deia-
field's HjEMatoxylin, and orange G (eosin may also
be used, but I think that the orange G gives better result*).
Such sections are excellent for preliminary work, and I
always use this stain as a standard test for a fixative which
is new to me.
l„y,|,AJI^,COOl^lC
Prolocerebntm oj Micropteryx. 147
Another valuable stain is FiCBO-HiGBOsra. This bringa
into special prominence the tracts of fibres which run
throu^ the brain. Counter-staining should be avoided,
and also over-stainiiig, because picio-nigrosin washes oot
only with difficulty in add alcohol. The beet results are
obtained with matenal &ssA in osmic acid or Flemming'a
solution.
Other hmnatoxylin methods have little to leconunend
them. The Weigert-Pal method cannot be used because
it is specific for myelin sheaths, which are never found on
the nerve fibres of insects.
5% IlSMATOXYLIN CONTAININO LITEIUH CABBONATB
is a stain for nerve fibres, but picro-nigroein gives similar
though more distinct results. Staining with Malloby's
afUATOZYLm is very strongly recommended by Kenyon
and by Altens. The process is complicated. At firat
sight the stain appears much too general to be of use.
Every tissue is stained a heavy dark blue, and differentia-
tion in acid alcohol is useless. I have discovered that the
sections may be quickly differentiated in a solution of
sodium bicarbonate in distilled water. The stain becomes
bright sky blue and most of the tissues are partly decolor-
ised. The nerve fibres retain the stain. This complicated
method produces results which are not really superior to
those produced by staining with picro-nigrosin. The blue
colour appeals to be permanent. The cytoplasm of the
fiant cells retains even more of the stain than do the nerve
bres. The ordinary small ganglion cells are very u:uch
decolorised in the alkali.
Mallobt's AmuN Blub. — This has been used by
Bretachneider in his work on the brain of Periplanela. The
method is one of great complexity, especially with the addi-
tion of the modifications which he Introduces. The results
are extremely beautiful, but most workers will probably
consider them hardly worth the trouble and time expended
on them.
Maxn's Stain.— I am much indebted to Dr. D. Keilin
for insisting on my giving a trial to this stain, which
will be found of great value. I find it best to stain first,
Kghtly, with Delafield, but this is not necessary or desir-
able except for nerve cells; the stain is \^'idely known
to insect histologists, and is strongly recommended to
neurologists. It is seen at its best when applied to
noateiial fixed in the fluids of Bouin, or Gilson, or in the
iM-,i,A J h, Google
148 Mr. P. A. Buxton on the
picro-chlor-acetic mixture ; it should not be used upon
OBRiic acid preparationa,
I have tned several stains which have proved more or
less useless, and I mention them below in order to save
others from wasting time upon them.
Methylene Blue and Methyl VroLET. — It appears
that the cells have little affinity for these stains. This is
remarkable when it is remembered to how lai^e an extent
methylene blue has been used as a vital stain for the
nervous systems of the Arthropoda.
Van Gieson's Stain, — This stain is useless because it
colours all the soft parts of the section a uniform jank
colour, without any of the differentiation which it gives
with sections of the tissues of Vertebrates.
Varioua preparations of carmine were tried, because of
its historic interest as the only stain used by the workers
of thirty or forty years ago. It appears to have singularly
little affinity for any part of the brain of Micropteiyx.
The stains on which I ^ace most reliance are Delafield's
hcemaloxylin wUh orange 0 as cownter-stain for preliminary
study, 'picTo-nigrodn and the reduced silver and gold method
for the study of the course of nerve fibres, and Mann's stain for
the nerve cms.
IV. Note. — Some of the fixatives and stains to which I
have had reason to refer are not very well known, and it
will perhaps he helpful if I give their compositions. The
picro-cklor-acetic mixture is 1 % picric acid in absolute
alcohol, 6 parts; chloroform 1 part; formalin (40%)
1 part ; glacial acetic acid J part. Fix twenty-four hours,
then three days in 90 % alcohol. Borrd's fluid consists of
osmic acid (08 04) 1 gm.; acetic acid 10 c.c; platinum
chloride 1 gm. ; chromic acid TS gm. ; and distilled water
175 c.c.
The syiril soap which is recommended as a reagent
for softening chitin is one of the official preparations of
the German Fharmacopceia : 6 gms. of olive oil are
saponified with 7 gnm. of a solution of potassium hydroxide ;
to this is added alcohol 30 gms., water 17c.c. (Kurt Bedau,
Zeitachr. f. wisa. Zoologie, Vol. 97, p. 418, 1910-11).
Of the stains the following should perhaps be described.
The piCTO-nigrosin I used was made up as follows : I vol,
1 % aqueous solution of nigrosin ; 9 vols, saturated aqueous
solution of picric acid. The fact that the various authors
Prolooerebrum of Microfteryx. 149
who have tried this stain give conflicting accounts of its
value is due to the fact that there is no standard composi-
tion for it. MaUory'a htBtnatoxylin. — ^The stain consists
of htematozylin crystals 1 gm.; chloral hydrate 10 gms.;
10% solution of phosphomolybdic acid in water 1 cc;
distilled water 100 c.c. The sections are mordanted in
5 % copper sulphate solution for twenty-four hours, washed
in tap water, placed for i or J hour in the stain diluted
with four times its volume of distilled water, rinsed and
carefully decolorised in a solution of sodium bicarbonate
in distilled water. Bretschneider's application of MaUory's
anilin lAue, with some slight modifications of my own, is aa
follows ; Delafield's hffimatoxyl 1 hour, or until the nuclei
are faintly stained ; wash ; eosin twenty minutes ; wash in
water; 1 % phosphomolybdic acid two minutes. Mallory'a
stain one minute ; wash, dehydrate and mount. The com-
position of the Mallory's stain is anilin blue (water-soluble)
I gm., orange Q (water-soluble) 2 gms., oxalic acid 2 gnis.,
distilled water 100 cc. Mann's stain ; 1 % aqueous sol.
methyl blue 35 cc, 1 % aqueous eosin 35 cc, water 100 c.c.
Stain ten minutes or longer, for it is impossible to oveistain,
tuid then wash in alcohol containing 1 % of pyridin, watch-
ing the process of decolorisation under the microscope;
with practice the right degree of decolorisation can easily
be obtained.
BlBUOaRAPHY.
A very complete bibliography is given by Eiihnle. It
is only necessary for me to give a list of the authors and
papers to which I have referred.
A. Works dealing with the Segmentation of the Arthro-
pod Head, etc
Brauer, a. (1) 1894. Beitrage zur Kentniss der En-
twicklungsgeschicbte des Skorpions. Part
I. Zeitschr. f. wiss. Zool., Bd. LVII.
(2)1895. Ditto. Partll. Op.cit.,Bd.LVlX.
Heymons, R. (1) 1895. Die Embryonalentwicklung von
Dermapteren und Orthopteren. Jena
(Fischer).
(2) 1897. Entwicklungsgeschichtliche Un-
tersuchungen an I^episma sacchanna.
Zeitschr. f. wiss. Zool., Bd. LXII.
(3) 1901. Entw. gesch. der Scolopender.
Zoologica (Chun.), XIll.
iM-,i,A J h, Google
160 Mi. F. a. BuztOQ on the
KoRSCHBLT and Hbidbe. 1899. Textbook of Embry-
ology of Invert«bnite3, Vol. III.
B. Works dealing with the Central Nervous System of
Insects.
Altem, Hans von, 1910. Zur Pkylogenie des Hymeno-
pteiengehinis. Jen. Zdtecbi. f, Katurwiss., 6d.
XLVI, pp. 511-590.
Bbrlbsb, a. (1) 1907. GU Insetti, Vol. I, pp. 569-588.
(2) MonogT. dei Myiientomata. Bedia, Vol.
VI, p. 134.
Bbetschnetder. 1914. Jen. Zeitschr. f. Naturwiss.,
Bd.LI.
BouviBR, M. E. L. Henri Viallanes, sa vie et ses tiavaux.
Ami. de Sci. Nat. Zool., S4r. VII, Tome XV (1893).
(CoDtains a complet« bibliography of the papers of
Viallanes, twenty-three of which deal with the nervous
system of the Arthropoda.)
DiETL, M. J. 1876. Die Oiganisation des Arthropoden-
gehirns. Zeitschr. f. wise. Zool., Bd.. XXVII, p. 489.
Fl6oel, J. H. h. 1878. Ueber den einheitlichen Bau
des Gehims iu den verschiedenen InsectenoidDungen,
Zeitschr. f. wiss. Zool., Suppl.-Bd. XXX.
Haller, B. 1905. Uebei den allgemeinen Banplan des
Tracheaten Syncerebrums. Aich. f. microscop. Anat.
und Entwick., Bd. LXV, p. 181.
HoLSTB. 1910. Zeitschr. f. wiss. Zoologie, Bd. XCVI,
p. 420.
J0NE8CU, C. N. 1909. Vergleichende Uatersuchungen
iiber das Gehim der Honigbiene. Jen. Zeitechr. f.
Naturwiss., Bd. XLV, p. 111.
Kenyon, C. F. (1) 1896. The Brain of the Bee. Joura.
ofComp. Neuiol., Vol. VI.
(2) 1896. The Meaning and Stnicturo of
the so-called " Mushroom Bodies " of
the Hexapod brain. Amer. Nat., Vol.
XXX.
KOhnlb, K. F. 1913. Daa Gehim des gemeinen Ohr-
wurm's (Forfic, etc.). Jen, Zeitschr. f. Naturwiss.,
Bd. L, pp. 147-276.
LowMB. 1893. The Blow-fly, Vol. II, p. 432.
Newton, E. F. 1879. On the Brain of the Cockroach.
Q JJI.S., Vol. XIX.
Packard, A. S. 1903. A Textbook of Entomology.
l„y,l,Ajh,COOl^lC
Protooerebrum of Micropleryx, 151
VuLLANES. (1) 1886. Le Cerveau de la Gu€pe. Ann.
de Sci. Nat. Zool., Sdr. VII, Tome II.
(2) 1887. Etudes hiatologiques et o^^o-
logiqu^ BUT les centres nerveux. V°".
M4moire. Le Cervean du Criquet (Oedi-
poda and Caloptenus). Ann. de Sci. Nat.
Zool., 86t. VII, Tome IV.
ZiEGLER, H. E. 1912. Die Getiinie der Insectea. Naturw.
Wochenschr., Bd. XI.
Abbbbviations Used.
In eveiy case the same abbreviations are used in the
Plates as m the text figures.
The letters a, b, c, a, etc., refer in all cases to tracts.
a. b, c, d, etc. (p. 136).
as. br. = ascflodjQg branch {= forward root) oi mushroom
body.
a^. ^ axonic substance ^ fibrillar substance.
ag. It. ^ ascending trunk of mushroom body.
hr. = bridge.
br. hd. = head of bridge.
d. I. ^ dorsal lobe of protocerebnun.
dm. = deuterocerebrum (= antennary lobe).
d. ty. s=s deuterocerebral or paired sympathetic system.
ei. = Einstrdmmung (see p. 120).
g. c. = ganglion cells of the " normal " type.
gi. c. = giant cells.
hd. = head of mushroom body.
in. ca. =. inner capsule of central body.
in. r. ^ inner root of mushroom body.
1. 1. = hiteral protocerebral lobes.
lo. = " la loge " of Viallanes.
mb. = mushroom body.
mb. c. = cells of musturoom body.
mi. I. = middle lobe.
mi. I. a. and mi. I. p. = anterior and posterior portions of
middle lobe.
mm. = tract mm. (p. 138).
mo. n. = motor antennary nerve.
ng. = nuclei of neuroglia cells.
nl. ^ neurilemma.
nn. ^ nnclei of the neurilenmia.
n,g,t,7.dh,G00glc
152 Exj^naium of Plates.
o. c. = cells of optic lobes.
oc.gl.= ocellarj' glomerulus.
oc. n. ^ ocellary nerve,
0. 1. = optic lobes.
ou. ca. = outer capsule ol central body,
pc. I. = protocerebral lobes.
po. br. = posterior branch (= backward root) of musb-
room body.
«. n. = sensory antennary nerve,
5(. = stem of mushroom body.
sw. hd. = swollen head of ascending branch.
sw. St. = swollen foot of stem.
Im. = tritocerebnmi.
tr. = tracheal tubes.
tu. s= tumulus.
vl. I. ^ ventrolateral lobe of protocerebrum.
vl. I. a. and vl. I. p. = ita anterior and posterior portions.
Explanation of Plates,
PUt«s Vll to IX (figures 1 to 16) Tepresent a series of Tertical
transverse Bections through the protocerebrum of Mieroplerj/x,
pftrtially diagrammaitic. Each figure represents a, BUcoeaHiTe sec-
tion 3'5 li thick, except that single sections are omitted betweem
figures 6 and 7, 9 and 10, 14 and 16, and 15 and 16.
The neurilemma is shown as a black line, the cells of the mush-
room body {mb. c. ) and those of the optic lobes (o. c ) hy black masses,
and the normal ganglion cells {g.c.) by dork grey, except where one
or two occur alone, in which case they are represented as indi-
vidual black dots (e. g. fig. 6). Giant cells {gi. e.) are always drawn
individually. Axonic substance is shown pale grey, except lbs
Stem of the mushioom body, which stains heavily and is coloured
dark, and certain other parts of the protocerebrum which aie
shown in medium grey because they stain slightly moio intensely
than other parts. Bands of fibres are shown as dark lines when
they are striking features of any section.
The upper part of each figure is dorsal, the lower ventraL
Plate X (fig. 17).— This plat« is from a thin section, hence the
amount of free space among the ganglion t^Us. The drawing ro-
Ij.y.l.Ajl^yCOO'^lL'
Trans. Ent. Soc. Land., igi^, Plale VII.
5 to
BUXTON: PROTOCEREBRUH OF MICROPTERYX.
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
Trans. Enl. Soc. Lond., igij, Plate X.
BUXTON : PROTOCEREBRUM OF MICROPTERYX.
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
Explanation of Plata, 153
presents the npper put of the protc>cerebniin (i/. fig. 12), and ahovfl
maahrooni body cells and nonnal ganglion cells lying in a loose
mBBB beneath the nenrilenuna (nJ.), which here contains tracheae
and chaiaoteristic elongate nncleL The minute glomenilar bodice
on the head of the mnshioom bodj are indicated, and also the large
tract r, typical of Faaemibetanz, and the donal lobe, on example
of Fonktaabetanz. Note the maaa of imgolar neuroglial nuclei
bcmeath one head of the bridge.
The plates are in every case drawn with the aid of a drawing
eyeiHece.
n,g,t,7.dh,G00glc
( 154 )
VI. Some Notes on Butterfly Migrations in Brititk Ouiana.
By C. B. WiLUAMS, M.A., F.E.S.
With Sketch Map.
[Read IhTCh IQi. 1017.]
The whole problem of the migration of insects, and of
butterflies in particular, is one of great interest, and, at the
same time, one about which we have so little data that it
is almost impossible at present to make any generalisations.
With the exception of records of such well-known migratory
insects as the locusts, most of the published accounts are
fragmentary, inaccessible or iBolated records for a coimtry
by passing visitors. From the records relating to butter-
flies almost the only common fact which emeigee is that
migrations are particularly abundant in the Pieridae,
records in this group being known from all parts of the
world.
I have recently spent a few months in British Guiana,
and have been fortunate enough to observe two migrations
of the yellow Pierid, Callidryaa eubtde, one of which lasted
for at least t«n days. I also questioned a number of
residents of the district about migrations, which when
large are noticeable even by the non-entomological public,
and find that they are a phenomenon of regular occurrence,
and that almost every one who has resided there for any
length of time has seen the clouds " of yellow butterflies
all flying steadily in one direction." Most of the accounts
were too vague to be of any value, especially with regard
to details ; but from among them I was able to get some in
which the locality and the direction of flight was given
with sufficient appearance of accuracy to make it seem
worth recording. In addition to these I have given ex-
tracts from a few published accounts of migrations in
Britbh Guiana, chiefly from local pubhcations which are
not readily accessible to the entomologist.
The climatic conditions near the coast of British Guiana,
to which district most of the records refer, consist of two
wet and two dr>' seasons each year. In general, FebruaT>'
TSANS. EMT. SOO. LOND. 1917. — PABT I. (NOV.)
ii^,Cooi^lc
Me. C. B. WUiiains' NoUa m BtMerfiy Migrations 155
to April ia more or less dry ; May to the middle of August
wet; the end of August to October the driest aeasou;
and November to Febrnaiy again wet. Furtbei in the
interior the second diy season is lost, and there is only
one wet and one dry season each yeai.
There are given below particulars of sixteen different
migrations (really more, as some of the records refer to a
number of similar migrations seen in the same locality in
different years). All except one of these reler to CaUidryas
eubide. Two are from my own observations, nine are
collected from residents and are, I beheve, quite reliable,
while five have been previously recorded. The exact
locality and direction of all, except one, will be fo\md
indicated on the accompanying map.
1. This migration I saw in the North-west District of
British Guiana within a short distance of the Venezuelan
border. Most of the observations were made on the River
Amka, between the junction with the Barima, of which it
is a tributary, and lasororo about six miles further up the
Aruka. The junction of the two rivers is about twelve
miles in a direct line to the coast, and about nine miles from
Venezuela. The district is chiefly a vast forest swamp,
below the high-tide sea level, with here and there a smtdl
rounded or &t-topped hill, one or two hundred feet high.
The migration consisted almost entirely of the yellow
Rerid CcSUdryaa ei^jule, interspersed with a very small
proportion of at least three other species which I was
unable to catch ; one of them, a large, dark, Fapilio-Iike
insect, may possibly have been Cydimon [Urania] Idlus,
a day'fl3ring moth which is not uncommon in the district,
and which nas been recorded as having migrations of its
OWTl,
I first became aware of the migration on August 1st,
1916, although three days before specimens had been
noticed in the district. It was then two and a half months
after the beginning of the first wet season. The migration
was only at times really conspicuous and never attained
a density which could without exaggeration be recorded
as a " thick cloud." On many days only careful observa-
tion indicated that anything unusual was occurring. In
order to get a comparative idea of the density of the
migration a distance of about two hundred yards was
estimated from the observer in a line across the direction
ot flight (in close spaces this sometimes bad to be reduced
iM-,i,A J h, Google
156
Mr. C. B. \^Uliamfi' Notes on
to one hundred yards or leas), and the number of butterflies
crossing this line per minute was counted.
It will be best to give the actual observations taken
at the time.
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1st Auffu^. — The yellow butterfly, CaUidry<a eiAuIe, waa
migrating in large numbers to-day. They were flying in
an easterly to south-easterly direction almost across what
is, for this district, a rather strong wind. They were first
Ij.y.l.Ajl^yCOO'^lL'
Butlerfiy Mirations in British Outana. 157
noticed crossiDg the Rivei Amka about half a mile from
Issororo. At 1.10 p.m. I counted in a httle over five
nuDUtes sixty-two CaUidryas, one orange butterfly of about
the same size, one smaller yellow species, and two dark
Paptlio-hke insects crossing a space estimated at two hundred
yards (moving as our boat moved). They became rapidly
more common, and from 1.30-1.35 I counted three hundred
and fifty crossing the same estimated distance. They were
then for some distance a bttle less common, but at the
junction of the Aruka and the Barima, where we arrived at
about 2.15, at least a hundred and fifty per minute were
passing. Below this the flight thinned out, and at Mora-
whanna, four miles below the junction, scarcely any were
to be seen. The course of the rivers is far from straight,
but the flight must have been at least five miles across.
About one in a hundred of the butterflies was a bright orange
species, and a smaller proportion of the dark FapUio-like
species ; this latter may, however, have been more common,
as it was difficult to see at a distance.
Catching butterflies while standing up in a small boat
is neither easy nor safe, but I managed to net two specimens,
one male and one female.
Ind August. — We left Morawhanna at 11.30 a.m., and
were in the migration practically the whole way back to
Issororo, where we arrived at 2.30 p.m. The butterflies
were occasional near Morawhanna, almost as common as
yesterday at the junction of the Aruka and Barima, and
frequent all the way up until near the end, when a very
heavy shower sent both ua and them into shelter. Even in
the thinner parts ten to twenty per minute were crossing
every two hundred yards. To-day I caught five specimens,
four males and one female.
3rd August. — At ten o'clock this morning the yellow
butterflies were flying at the rate of two or three per minute
past the bouse (in a clearing about one hundred yards wide
and about half-way up a bill about two hundred feet high).
At 10.15 none were to be seen ; at eleven o'clock they were
again noticed for about five minutes; at 12.30 they were
still passing over three or tour per minute, and again at
two o'clock they were seen occasionally in intervals between
showers. At 3 p.m. a few more were noticed. I caught
to-day one more specimen, a female.
4th August. — At 9.30 a.m. six or seven CaUidryas were
seen passii^ south-east over the house; about 11 a.m. they
iM-,i,A J h, Google
158 Mr. C. B. Willi&ms' Note* m
were noticed whenever the snn was shining; between
11.35 and 11.40 thirty-foor passed over on a hundred yards
line. At 1 p.m. two or three per minute were Bying rapidly
at the foot of the bill, all going south-east. At 2 p.m. they
were still passing, bnt none were seen after three o'clock.
One caoght to-day was a male.
5lh Augtat. — Very wet, r38 inchee of run ; no butterflies
seen.
6th August. — They were first noticed to-day at 11 ajn.;
from 11.30-11.35 sixty-three passed over the garden, 'i. e.
thirteen per minute on a hundred yards line.
llh August. — Iha butterflies are still migrating. At
9.30 a.m. six or seven per minute were passing the house;
at 10.15 a.m. eight a minute were paasmg on a hundred
yards of garden ; and agun at mid-day and at one o'clock
they were still flying. One specimen captured to-day was
a male.
&A August, — At 11.15 a.m. they were again noticed
migrating. The sun seems to make a great diflereace to
the flight. I started counting at 11.16 a.m., and in the
first minute eight passed ; then the sun went in, and in the
neit four minutes only three were seen ; in the first two
minutes of sunshine after this fifteen passed. Do they
follow the patches of sunlight, or settle when a shadow
comes ^ The former does not seem possible, as they have
always been flying across the direction of the nind. At
1.15 occasional specimens were seen crossing the river,
but they were far apart ; at a casual ghince it might not
be noticed, but every ten seconds or so one would come
into view going full speed south-east.
9{A August. — The butterflies were still in migration,
but only occasionally in intervals between heavy showers.
10(i August. — The butterflies were plentiful to-day,
crossing the river at Issororo at least twenty to thirty per
minute on two hundred yards line. Half a mile between
Issororo they thinned out and remained at three or four
per minute right down to the river junction ; below this
to Moiawhanna only very few were seen. I made an
attempt to-day to estimate the speed at which they were
flying; previously I had found it impossible to overtake
them even when on a clear footpath, arid bad considered
their speed as at the very least nine miles per hour. The
river now gave a good opportunity of testing this, as they
were passing directly across it and could be timed from
BuUtrfiy MigraHona in British Quiana. 159
one aide to the other with ease. The river waa at least
a hundred and fifty, and poasibly two hundred yards vide.
Tbiee butterflies were timed and took twenty, twenty-four
and twenty-six seconds to cross. Taking twenty-four
seconds as an average and a hundred and fifty yards as the
distance, this gives twelve miles per hour (if the river were
two hundred yards wide it would oe sixteen miles per hour).
This ia, I think, a not unreasonable estimate.
In flying the insects kept close to the surface over which
they were passing ; over the forest they seemed never to
be more than a few feet above the tops of the trees, and in
any but the smallest clearing they descended rapidly to the
ground and flew between small bushes and trees about four
to eight feet up. On crossing the river they kept still
lower, being seldoni more than three feet above the water
and occasionally almost touching it.
Any attempt to estimate the number of butterflies in a
flight of this kind must of necessity he only an approxima-
tion. Taking ten butterflies per minute per two hundred
yards as an average, this gives 5,400 per hour per mile,
or 135,000 for a day of five hours on a front of five miles.
At the rate noted at the junction of the Aruka and Barima
on the Ist August {150 per minute) 84,000 would pass in a
single hour on a front of one mile.
On the 10th August I left the district, and was later
informed by Mr. A. A. Abraham, Manager of the Govern-
ment Experimental station at Issororo, that the butterflies
disappeared shortly after my departure.
The chief points of the above migration may be summar-
ised as follows : (1) The migration lasted at least ten days ;
(2) both males and females were represented, the former
predominating (my total captures were seven males and
three females). (3) They Hew constantly in a south-
easterly direction at a speed of about twelve miles per
hour across the prevailing north-east trade wind.
2, In the middle of October 1916 Mr. Abraham wrote
to me from Issororo to say, " Since my last note to you [end
of September] I have observed that the butterflies are
returning at the same rate and numbers to the bank of the
river from which they migrated." This is a particulariy
interesting record, as it confirms several other records of
migration in opposite directions in the same locality about
wliich I had been doubtful.
3. In September 1916 I observed a very diffuse niigra-
iM-,i,A J h, Google
160 Mr. C. B. Williams' Notes on
tion at Bartica at the junction of the Esaequebo, Cuyum
and Mazanmi Kivers, and from there down towards the
mouth of the Essequebo. On the 11th September 1
noticed that all the CaUidryas passing over the garden of
the house where I was staying (at H. M. Penal Settlement
on the opposite bank of the Jlazanmi to Bartica) were
flying in the same direction, towards the N.N.W., at full
speed. They were not common, and in twenty minutes
(12.40-1 p.m.) I only counted thirty-nine, i. e. about two
a minute ; during that time not a single one was seen to
settle, or fly in any other direction. They were still
passing over about half an hour later, but after that the
usual afternoon rains came on and they were not seen
again that day. There was a slight east wind blowing at
the time. On the following day (12th September) at
Bartica, several were seen, again all fljing full speed N.N.W.
On the 13th September I descended the Essequebo to the
mouth by steamer, and during the early part of the journey
CaUidryas were fl}'ing N.N.W. at the rate of four or five a
minute on a two hundred yards line. There was then
practically no wind. About 10.30 a.m. the north-eaat trade
wind became stronger, and the flight changed to a westerly
direction (almost directly across the river instead of down
it). This would be the natural resultant of a N.E. breeze
on an attempted flight in a N. or N.N.W. direction,
4. Mr, Withers, Jilanager of Hills Estate, Bartica, who
has resided in the district about seven years, tells me that
he has many times seen the yellow butterfly migrating,
sometimes for days together, alrcays in a N. to N.W.
direction. These flights, he says, usually occur in October.
5. Mr. Frear, Chief of H. M. Penal Settlement, tells me
that about the beginning of August 1916 there was a small
flight about ten miles up the Mazaruni River flying north-
west. This would be about the same time as I saw them
at Issororo, flying in an exactly opposite direction.
6. Mr. Cameron, Acting Chief Engineer of the Govern-
ment Steamer Ser\'ice, tells me that on the 8th September,
1916, he saw a small number of yellow butterflies at
Camaria on the Kiver Cuyuni, about eight miles above the
Penal Settlement, all flying north-west,
7. H. W. B. Moore, in " Timehri," the Journal of the
Royal Agricultural and Commercial Society of British
Ciuiana, 3rd Series, vol. ii (1912). p. -105. says: "In
July ... I ohserved thousands of CaUidryas eiibule . . ,
ih,Cooi^lc
Bviterfly Migrations in Briliah Guiana. 161
flying across the Essequebo Bivei from Wakenaam to Hog
Island and Great Troolie Island [all near the moath of the
river]. They were fiying chiefly in ones, but twos and
threes and higher numbers were not infrequent, whilst
once in a way a flock of twelve to twenty could be counted.
The great majority were males. . , , Soon after Kunibaru
Islands were passed the butterflies were seen to be flying
from the mainland towards Wakenaam. Going on towards
Aoiora and Suddie on the west bank at the mouth of the
river they were seen flying upstream, following, it seemed,
the direction of the wind." All the directions mentioned
are between south and east.
8. Mr. Marshall, who was for many years Manager of
s sugar plantation near Suddie, on the west bank of the
mouth of the Essequebo, tells me that migrations of the
yellow butterfly are a regular phenomenon there, chiefly
in May and June, rarely July, always flying from the N.W.
towards the S.E., wMch is across the prevailing N.E.
wind. They come from the north-west, and frequently
reach the coast just north of the Suddie ; thete they turn
southward along the coast to the mouth of the Essequebo
and then cross over via Tiger Island, Wakenaam, Leguan,
etc. It must have been one of these migratiooB which
Moore describes above (7).
9. Mr, Humphreys, for many years Manager of a sugar
plantation at Anna Regina, whidi is not far from Suddie,
says that he has frequently seen migrations going in a
northerly direction down the Essequebo Kiver and up the
Coast. This is in the exact opposite direction to the last
ivcorder, but there is, I think, no reason to doubt that both
are possible. Mr. Humphreys made the further interesting
remark that the butterflies were sometimes in a long,
nanow band, only ten to twenty feet wide.
10. Mr. Rodway, in " Timehri " (see above), 3rd Series,
vol. i (1911), p. 131, says: " CaUidryas eubide ... is
noted for its extensive flights ; for hours they have been
seen passing over the Demerara River going east, probably
billions in number, and all males. Whence they come and
whither they go is a mystery. Their food-plants [Cassia]
aie common everywhere, and there does not appear to be
any scarcity of females," He refers again to this migration
in "In the Guiana Forest," 2nd edition, 1911, p. 122,
bat with no further particulars.
11. Mr. Feteikin, of the Department of Agriculture,
TEANS. ENT. SCO. LOND. 1917.— PABT I. (NOV.) M
u,y,i,Ajh,Cooglc
162 Mr. C. B. Williama' Notes on
telta me that about 1906 he saw a migmtioa of some
thousands of yellow butterflies crossing the Demetaia
River from PlaDtatioQ Diamond to Plantation Wales,
that 18, from the east to the west bank about six miles from
the mouth of the river. They were fiying with a strong
wind behind them.
12. The Rev. Mr. Salmon tells me that at ^lamar (about
sixty miles up the Demeraia River) at the begimiing of
August 1916 be saw a " procession " of yellow butterflies
between his house and the river. They were flying in
groups of ten to twenty, with a short interval between
each group. The flight was first noticed at mid-day and
lasted for several hours after this. It might have already
been proceeding for some time. They were flying approxi-
mately from N.N.W. to S.S.E.
13. Messrs. Bancroft and Ward described to me a migra-
tion that they bad seen a few days before, about the 20th
September, 1916, at Murirato, about ten miles above
Wismar on the Demerara River. The butterflies were all
flying from the left to the right bank of the river, that is,
from west to east.
14. Richard Schombu^k, in " Reisen in Britisch Guiana,"
Zweite Theil, Leipzig, 1848, p. 157, describes a migration
of yellow butterflies which flew from S.E. to N.W. in the
interior of British Guiana neat Firaia about the 13th
September, 1842. The flight lasted the whole day, and at
mid-day and just before sunset the butterflies settled in
countless numbers on the patches of sand at the edge of
the river. According to the natives they were the butter-
flies which came from certun caterpillius and chrysalides
which they readily ate.
15. Both Moote {I.e.) and Rodway ("In the Guiana
Forest," 2nd edition, p. 122) refer to a record of a migration
by Robert (not Richard) Schombui^k, of which, however,
I have been unable to trace the original. Moore says it was
observed by Sir Robert Schomburgk " on the 18th of
October, 1838, when going up the Esaequebo, and it con-
tinued crossing the coutse of the river for nine houis and
a half, during which time his boat ascended nice miles.
A thousand million b not too high an estimate for the
number of individuals in the swarm."
16. This last record relates not to CaUidryas eubide, but
to another Pierid, Appias tnargarita, a small white species.
Mr. A. Leechman, m the " British Guiana Handbook,"
Ij.y.l.Ajl^yCOO'^lL'
Buttafiy Migrations in Briiish €fu%ana. 163
1913, p. 137, writes that he has " once witnessed a flight
of white butterflies (Apjnas margartia) on the lower left
bank of the Berbice River which lasted for over three days,
and could only be compared during the whole of that time
to a heavy sDowatorm. And the extraordinary thing waa
that they were all flying directly out to sea. Mr. Leechman
has informed me that this occurred in April 1909.
It is difficult even from the above records to get any
indication of what is happening. On the real problem,
why the migration takes place, there is still no light, and
many more correlated observations must be made before
there can he any hope of solving it.* There are, however,
a few points to which attention might be directed. In the
first place, there are here two general directions of migra-
tion for CaUidryas eubtde, roughly from the north-west to
the soath-east, and vice versa. There is as yet no record
of a migration in a north-east or south-westerly direction.
The prevailing wind is the north-east trade wind, so that
the migrations were across the wind. It is impossible to
say if this is the real determining factor in the direction of
the migration or not.
Secondly, all the migrations of CaUidryas in which the
date is recorded took place between May and October.
Thirdly, it seems possible to distinguish between several
different types of migration. At least one can make a
rough bat convenient grouping into three classes. (1) The
thick cloud, " like a snowstorm," a case which immediately
attracts the attention of the ordinary individual and is
most frequently recorded. (2) The diffuse migration,
which may vary from distinctly noticeable to so attenuated
that only a close observer would realise that anything
nnnaual was happening. There is, of course, no strict
line of demarcation between this and the last, and the
edge of a " cloud " migration would probably be difiuse.
A diffuse migration, however, can exist unaccompanied
by a cloud. (3) A particularly interesting form is the
" ribbon " migration, or " procession," in which a narrow
band of butterflies a few feet or a few yards wide flies
across the country in the direction of its length. The edge
of such a migration is well defined. In this connection
it might be mentioned that, even when not on migration,
* firitash Qoiaiia would aeem to be a promuing field for each
inyeatigations, bat they miut be extended over s. series of yaan,
with a number of competent observeni atatioaed over the ooo&trjr.
iM:,i,A J 1^, Google
164 Mi. C. B. Williams' NoUt on Butterfly Miffraiiont.
CaUidryas eubvle has a habit of S.yiag round and round ft
field in short stringB o{ about half a dozen almost head to
tail, and closely following each other's movements. This
habit may throw some light on the formation of the ribbon,
but does not explain the movements of the leader.
f^ally, we have the extremely inteieating question of
the sexes Fepresented in the migration. Rodway records
that all that he saw were males, but I understand that this
was from observing their colour whilst in flight. The male
and female of CaUidryas eubule differ distinctly in colour
and markings, and perhaps any one very famiUar witA
this species could tell them apart in this way. I found it
impossible to do so, and could only tell the sex after capture.
My specimens were, as mentioned before, seven males and
three females. It does seem, then, to be a general rule
that the males predominate, and this is confirmed by obser-
vations in other parts of the world. This branch of the
subject seems to me to be of fundamental importance, for
if the migrations consist so largely of males, miat becomes
of the corresponding females ? Mr. Bodway has bred this
species and bids the two sexes to be more or less in equal
numbers, and this is the general rule for other insects
except in rare cases, such as parthenogenetic reproduction,
whicn seems scarcely feasible here. There remains the
possibility of the males developing more rapidly and
emereing from the chiysaUdes earlier than the females,
but there is no direct evidence for this, and against it is
the fact that, except perhaps at the b^inning of the first
wet season, the successive broods of insects in the tropica
are ill-defined and usually overlap consideraUy. It may
be contended that the females are lees fitt«l for long
flights, being heavily laden with e^. Even if this is so.
we are left with the question, " Why, then, do the males
migrate? "
n,g,t,7.dh,G00glc
Trans. Ent. Soc. Lond.. 1917, Pari I.
p
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( 165 )
VII. The condition of the scales in the leaden males of
Agriades thetis, Rott,, and in other Lycaenids. By
E. A. CoGKAYTre, D.M., F.R.C.P., Temporary
Surgeon, R.N.
[BMtd Uftroh 7tb, 1917.]
Plate XI.
The peculiar colour of the leaden males of Agriades ihetis,
many of which were taken near Folkestone in 1916 and
occasional specimens iu previous years, led me to think
that a microscopical examination of theit scales might be
of considerable mterest.
In the nonual male thetts the wing is covered with lines
of smoky scales, sbott and broad, which have strong
longitudinal ribs and well-marked cross striations. Alter-
nating with the rows of dark scales are rows of longer
scales, which I call for convenience the "colour scales."
These aro also longitudinally ribbed, but have weaker
cross striation, and are yellow by transmitted light and
brilliant blue by reflected light. The blue colour has
generally been regarded as an interference colour due to
the cross striae and not a pigmentary colour, but H. M. Sims
(Canadian Entomologist, 1915, p. 161) considers that it
is dependent on a fluorescent dye. In either case the
dark scales serve to absorb any light which passes through
(he blue scales, and to prevent the coloured scales on
the underside of the wing from being visible on the
upperaide.
In addition to these two kinds of scales small colourless
androconial scales are present in abundance. In the leaden
males of ihetis, the smoky scales and androconia are of
normal shape, size and colour, but all the colour scales
are very thin, and have their distal part rolled up to form
a tube. By reflected light the curled-up edges and tubular
ends of these scales look silvery, and under a low power
of the microscope appear as ghostly triangles overlying
the dark scales, which are much exposed to direct view
and give the leaden colour to the wings.
TEAKS. ENT. SOC. LOND. 1917. — PABT I. (NOV.)
ih, Google
166 Dr. E. A. Cockayne on the eoitdiHon of the scala
Exatniiied under a high power (^ inch oil immenion)
it is seen that they are quite colourless by transmitted
light. The longitudinal nbs are present, though often
crinkled, but no cross striae are present with the excep-
tion of a few imperfect ones at tne extreme base of the
scales.
A few scales were found in which the lateral margins
were turned up, and in which very pale yellow dye was
present, but no cross striation. Unfortunately they were
mounted in balsam, and I could not see whether they
were blue by reflected light. K a scale of this natiue
could be isolated and examined umuoimted it would settle
the controversy as to the cause of the blue colour in the
blue Lycaenida.
Near the base of all four wings in both the specimens
examined, and along the costal maigin of the right fore-
wing in one of them, normal thick blue BceAes were found
with the abnormal ones.
Some of the scales on the fringes were tlumier and more
hair-like than is usual, but the scales on the undersides of
the wings were all quite normal. The peculiarity can
scarcely be due to any pathological condition acting upon
the scales from without. If this were so the neighbouring
dark scales and androconia would not escape, nor would
the scales of the underside be perfect.
No injury, nor any infection by a pathogenic organism,
would be hkely to affect the upper surfaces of aJl four
wings in the uniform and complete way in which it is
almost always affected in these leaden thais. It is much
more likely to be dependent on some inborn error of
development.
The following observations of Mr. G. T. Bethune-Baker
lend strong support to this view. According to this
author the blue scales in Pdyommatia dduB var. vUbUa
are very similar to the abnormal scales of the abemtiou
of thelis. I quote the description of these in his Frea-
dential Address (Proo. Ent, Soc. 1913, p. clviii) : " An
extraordinary character, however, obtains in the ordinary
blue wing scales, the whole of which are curled round
so as to form more or less short tubes ; the process ap-
pears to be that each side of the scales turns over, and
occasionally they meet thus in the centre, bat ■ mote
generally one side will overlap the other and so form a
more or less perfect tube ; by this I mean that the basal
u,y,i,A J h, Google
m the katkn maUs 0/ Agriades tJutit. 167
and ft|HC&l ends nmain open — a tube that is sealed at
each end naturally ceases to be a tube, becoming a
cylinder."
ThiB description agrees very closely witt the condition
met with in the " colour scales " of the leaden thetis, but
in dolus the tubular scales are blue and preBOmably retain
th^ cross striation or the fluorescent dye. Similar rolled-
np scales have been described by Mr. Betbune-Baker in
the " Menalcas " group, in which the wing colour is much
wbit«r than in most I.ycaenids.
I wrote to Mr. Bethune- Baker, who has examined micio-
soopically several leaden aberrations in his collection, and
liaB very kindly allowed me to publish hia notes on the
condition of the scales in them. He has examined one
Lycaetui arion, one Polyommatus icams, two Lycaenopsis
{Cdatlrina) puspa, two Lycaenopsis platitn, and a single
specimen each of three species of Tajuria, an exotic genus.
In the anon, which is a dull bluish colour, the scales
ate on]y curled up in a small percentage, in the majority
bring merely tfain and inclmed to buckle at the edges.
Id the icorus, which is greyer though not extremely leaden
coloured, the scales are thinner and more curled than in
the arion.
In all three species of Tajuria, which are much mote
leaden coloured, the acalee are rolled upwards and inwards
at the sides, the roUing being deeper at the apices, so that
in many tbey have a triangular shape. The rolled-up
Kales are abnormally thin. This thinuess is readily demon-
etiated in one Tajuria, which has one or two spots of
blue on the wings, where the scales are flat, normal in
colour and of much greater density.
The two spedmenB of Lycaenopsis pmpa from Formosa
ahow much the same condition as the Tajuria. The two
L. ptanta from Borneo are quite unusually leaden coloured.
Under an inch objective the " colour scales " are almost
invidble, but imder a 12 mm. objective are seen to be
excessively thin and rolled up absolutely tight, so tight
as to look like a number of short thick pale hairs scat-
tered over the surface of the wings. The fringes are also
abnormal, the usual long, elegant, somewhat fan-shaped
scales being replaced by scales like long thin haiis with
the apex slightly split up.
Breeding e:^riments with leaden thetis would be very
interesting. These leaden aberrations, so widespread
iM:,i,A J 1^, Google
1&8 Dr. E. A. Cockayne on the scales of Agriades thetit.
tkiough the blue Lycaemde, may well be Mendelian
recessives to the nonual blue males. The " dolus " and
" merudcas " group may be examples of Mendelian reces-
sives superseding the normal dominants through whole
Bpeciea, as the recessive form of Callimorpha dominula with
yellow bind-wings has entirely replaced the dominant with
red hind-winga in certain parts of Italy. This suggestion
is one put forward in order that some one may take the
necessary steps to prove or disprove it.
A further point of interest in connection with the leadoi
thetis is that Mr. L. \V, Newman thinks that they are
unusually fragile. It is quite possible that the whole
wing membrane is thinner than that of normal males,
and, if so, it may be correlated with the thiimess of the
" colour scales." I have not been able to satisfy myself
on the point, as I did not wish to destroy a specimen for
this purpose.
Explanation of Plate XI.
1. " Colour " Bcalea of le&den taale AgriadM Aetie.
2. Blue " colour " Boale of normal mole Agriadet lAefw.
3. Smokj (light absorbing) scale of leadeD male Agriadtt lAefio.
4. Andcocoiual ecale of leaden male Agriades IhtUt.
The fine regular striae of the blue scale are onlj rougblf indicated.
The drawings ai« magnified equally.
n,g,t,7.dh,G00glc ■
Trans. F.tit. Sac. Lend., 1917, PlaU XI
E. A. Cockayne, del.
SCALES OF AGRIADES THETIS.
ll,y,l,7rJM,G00glc
n,g,t,7.dh,G00glc
VIII. On new and liUle-knmcn Lsgriidae frmn Tropical
America. By Georob Chablbs Champion, F.Z.S.
[Bead Hucli Tth, 1917.]
Plates XII, XIII.
The present paper gives an account of the unnamed species
of Slalira from S- America, the Antilles, etc., in the British
Museum, supplemented by those in the Hope Museum
at Oxford, and a few Brazilian and Trinidad forms kindly
supplied by Mr. G. E. Bryant. The study of these S.
American insects had to be undertaken under exceptionally
difiBcult circumstances, as not a single definitely-named
representative of any of the numerous species described
by Maklin was to be found in the collections in this country.
It is therefore not improbable that some of them have
been incorrectly identified by me from the descriptions
alone, in which the sexual distinctions are not given, or,
if seen, not recognised as such. The materia] examined
is mainly that obtained by J. Gray, H. Clark, and A. Fry
in the neighbourhood of Rio de Janeiro, and by H. W.
Bates on the Amazons, the specimens from other S.
American localities being few in number. Dr. R. F,
Stthlberg was in Brazil in 1850 and earlier, and a few of
his captures (most of which have been identified in the
Fry collection) were described by Maklin in 1875. H. W.
Bates appears to have paid a good deal of attention to the
genus Statira and ita allies ^ while he was resident on the
Amazons, doubtless on account of the great resemblance
of many of them to genera of Carabidae, e.g. Agra, Casnonia,
Drmniua, etc.
Statira, Lep. et Serv.
Eighty-three members of this genus were recorded
from Central America in the "Biologia" in 1689-1893,
one only of which was known to me at that time to occur
south of Panama. The collections from South America,
> Tb«ae Utter are dMcribed in the "Entoniologitt'B Monthly
Uasftzine," Vol. LIU, pp. 132-154, lSS-196, 218-223, pi. 2, Jont-
October, 1917.
TEANS. BHT. SOC. LOND. 1917. — ^PART I. (NOV.)
l„y,|,AJI^,COOl^lC
170 Mr. G. C. Champion on
etc., aow under ezaminatlon, include repreBentatives of at
least 130 more, showing that hundiads of Bpedes must
exist on the southern continent. The following additions
to the distribution, etc., of six of the Central American
forms require notice : S. detUiculata, Colombia {Mus. Brit.),
an insect with denticulate anterior femora, one of two
placed under Sect. Aa in the table given in the " Biologia,"
the other, S. glabrata, having been found by BioUey as far
south as Costa Rica; S. costarioensia, Colombia; 5. ingens,
Costa Rica {BioUey) ; S. aibolineata, Belize, British Honduras
{Mvs. Brit.); S. nigripennis, Makl. [not Champ.], var. y
(Biol. Centr. Am., iv. 2, pi. 2, fig. 18), from Mexico, has
been named ckampioni by Pic (Melanges ezot.-bntom. iv,
p. 20, Sept. 1912). Amongst the South American Staiirae,
some {S. catenata, etc.) have very remarkable characters
in the legs, antennae, or aedeagus,' or in the clothing of
the under surface, in the males. Other structural peculiar-
ities to be noted are : the presence of a long curved spine
on the narrow basal portion of the anterior femora • in
both sexes of certain species {S. atUuTolis, etc.) ; two rugose
stigmata on the disc of the protborax {S. dtsligma, ,J);
a basal constriction of the anterior tibiae (S. degans, de-
gantuJa, casnonioides) ; a deep, sharply-defined, triangular
or oblong sulcus on the prothorax in front {S. vagegiUtata,
etc.), etc. The apical joint of the antenna, too, is often
greatly elongated in the male, as in Lagria. The species
here enumerated may be grouped thus : —
A. Anterior femon armed with t, long curved
Bpine at base Noo. 1-6.
B. Anterior femora unarmed at base.
1. Species large, shining, oastaneous, with
darkor, submetallic, oatenulato-tuberou-
late elytra, and sharply dentate anterior
tibiae No. 6.
* Dr. Sharp has beem kind enough to examine this organ in two
apeoiee (S. vtridipennis and S. geniculata), and he telu me that
when two projeoting pieces are visible, one will be the conjoined
lateral lobM (t«chiiioiilly "tegmen"), the other the median lobe,
this being really tlie inner one.
* A character evidently noticed by Hope or Weetwood, the
speoimra of iS. cotUineenms in the Osford Huseum having an ante
leg detached and mounted separately to ehow this stnictuie.
was overlooked by Haklin in 8. tutwatie.
ll,y,l,/eJh,G00glc
Hew and liliU-known I/ignidae. 171
2. Speoies large, elongate, mostly i«eembling
the Corabid-goius Agra, the elytra
□snally metallic,* nigro-teaaellftte in 8.
meUagria, and with two or more of the
altoniftte intoTstieeB catanulato-tubenm-
Ut« thiou^oat their Jengtb or at leaat
teworde apes, the posterior femora
bidcntato at base in 8. gemmifer; 3 eome-
times irith BtrDD^y pronounced secondary
sexual ebaracters in the legs or sedeagus,*
or in the clothing of the antconoe or
ventral surface.
a. Elytra mucronate at tip Nob. T-16.
b. Elytra not mucronate at tip Nos. 16-34.
3. S^tcdes large, very elongate, with a long,
conical, red prothoiaz and metallic elytra,
the latter acuminate at apex and with
almost smooth interstices No. 3tS.
4. Species moderately large, narrow, black,
with narrow head, elongate-cylindrical
prothcrax aitd cyaneons elytra, the inter-
stices of latter almost smooth . . ■ No. 36.
fi. SpeoieBmoderatetyloi^, block, with brilliant
green, non-tuberculate, mucronate elytra,
and exti«mely slender antamoe (as in
Othryadet) No. 37.
6. Species moderately large or small, reeemblii^
some of those placed under B 2a, with the
elytra unarmed at tbe tip, blue or green, and
bearing small, scattered, rounded tubercles Nos. 3S-40.
7. Species resembling some of those placed
under B 2b, with uneven, very feebly
striato-punctate, brilliantly metollicelytia,
which are catenulato-tuberculate towards
aides or apes Nos. 41-43.
8. Species moderat^y large or small, the elytra
usually metallic or with metallic lustre,
with simple scattered setigerous impres-
sions to tip (interruptedly catenulate
laterally in 3. impreastpennU).
* 8. agraeformit. Champ., from Panama, belongs te this section.
• Penis-sheath {= conjoined lateral lob» of tegmen, sec. Sharp)
twisted and asymmetric in males of S. eattnala, vmdipemua, geni-
Cfiata, atgmmeiTka and Uaivpta.
iM:,i,A J 1^, Google
172 Mi. G. C. Champion on
a. Prothorax not or obeoletely cantlionlate
oadiM NoB.«^40.
6. Prothorax distinctly canaliculate on due . Noe. 61-6C-
9. Species elongat«, with opaque black head
and prothorax and purplish or dull black
elytra, the latter bearing small rounded
tuberclee, the legs long Nos. 60, 67.
10. Species with a abort, broad head and pro-
thorax, metallic elytra, and stout an-
tonnaa • No. 68.
11. Spcoies cBstoneouH in oolour, with broadly
viridi-vittateolytraandatoutantramae . Noa. 69, 70.
12. Spnoies teBtaceouB or brown, with the elytia
(except in vara.) infuscate or metallic along
the sides, and the antennae slender,
o. Head and prothorax shining .... Noe. 71, 72.
b. Head and prothorax opaque, scabrous . No. 73.
13. Species testaceous, reddish, or brown, with
the legs partly infuscate.
a. Prothorax densely, more ooaisely punctate No. 74.
b. Prothorax finely aoabroso -punctate or
almost smooth Nos. 76, 76.
14. Species teetaoeous or reddish- brown, the
legs included.
a. Prothorax feebly shining; eyes subap-
proximate No. 77.
6. Prothorax opaque, alutaoeous ; eye*
distant No. 78.
Ifi. Speoiee testaceous, with blackish, flavo-
vittate elytia, the upper surface dull . . No. 79.
16. Species oastaneous or pioeous, shining.
a, Eljara with each alternate interstice more
or lees tubetculate throughout .... No. 80.
b. Elytra with scattered setigerous impios-
siona, sometimes nigro-lineate .... Nos. 81-86.
17. Species with red head and prothorax and
blue or green elytra.
a. Anterior tibiae compressed at liase; alMJo-
men black No. 87.
6. Anterior tibiae simple i prothorax (5) with
two scabrous patches on disc . . . No. 88.
* The Mexican S. eroMieomis, Champ, and the ColombiaD
S. validieomit, Makl. belong to this section.
Ij.y.l.Ajl^yCOO'^lL'
N^ea and litUe-hwum Lagri^e. 173
IS. Speoiea Hmall, slender, with opaque bkok
head and prothoiax, blue elytrs, uid
yellow tarsi No. 89.
19. Speoiei Blender, with large bead and narrow
prathoraz, piceous or in part testaceous,
resembling the Carabid-genera Dromiiu
and Casnaniai elytra witli setigerous im-
preaaions, preceded b; a more or leu
distinct tuberole; anterior tibiae oam-
prCKed at bass in iS. eatnonioidta Npa. 9(MI3.
20. Species testaceous or teddish, the elytra
(except in vars.) with metallic or black
markings or tasoiae, these latter sometimes
greatly extokded ox conflnent, the alternate
interstioee, at most, with setigerous im-
pressions, the protborax not incised on
Oie disc in front, sometimes bivittate.
0. Elytra with nnmeroua small tuba;clea . No. M.
b. Elytra not tuberculate Nos. 9C-106.
21. SpeciestestaceouB, withnigro-teesellat«elytra No. 107.
22. Species teataceooB, with a deep incision on
the disc of the prothorax in front, the
elytra (except in vors.) with coalescent
Uook markings or wholly black . . . Nos. 108-110.
2J. Species rufo-t«taceons, shining, with nigro-
birittate prothorax and nigro-fssciate,
closely setose, elytra, each of the inter-
stices of latter seriato -punctate . . . No. 111.
^ Species testaceous, hairy, with coarsely
punctate prothorax and nigro-maculate
elytra, each of the interstices of latter
Beriat«-punet«te Nos. 112, 113,
^ SpedeB slender, tesUuteous, with bead and
prothorax opaque and elytraJ suture, at
least, inf UBoate.
a. Elytra tuberoulat«; head small . . . No. 114.
6- Elytra not tuberculate ; head la^e . . No. Ufi.
S8. Species with prothorax distinctly margined
laterally.'
o- Body obscure testaceous, the elytra fusco-
iHciate l^icaiagoa] No. 116.
0, Body nigro-piceona or black, the prothorax
,.^»ed, the elytra blue No. 117.
varioos Central- and N.-American forms belong to this section.
ih, Google
Mr. G. C. Champion on
174
27. Species anull, Antiuoiform, with short be«d,
small e3rea, Btout ant«iuise, and faintly
striato-punotaM elytra, the elytra with
irregularly distributed, aetigerouB impree-
sions.
a. Body shining black, the prothora
patch on elytra often reddish
b. Body blaotc, the
dytra blue .
e. Body and terminal joint of antennae t«8-
taoeouB, bead and apex of elytra black
28. Species resembling Uiose placed vinder
sect. 27, but with longer, deeply punctate-
striate elytn, teetaeeous, with apical half
of elytra violaceous
29. Species with head considerably developed
behind the eyes, the latt«r small, the prO'
thorax subeylindrical or cordate.
a. Head and protboiax closely, coarsely puno-
tate; antennae stouter; body reddish-
brown, pioeous, or black, head and pro-
thorax aometimee testaceous.
"fElytra more elongate, with
setigerous impressions on alternate inter-
ttElytra shorter, with very few setigerous
impressions; protboiax subcyliiulrical
or cordate ; species small, Anthidform
6. Head and protliOTSx much smoother,
obsoletely punctolate; antranae wry
slender; body obaoore t«ataceous, dytra
black
30. Species small, narrow, aeneo-pieeous, with
very coarsely, rugosely punctured head and
prothorax, small, depressed eyes, and each
elytral interstice seriato-punctat«[Uexjca]
31. Speciea elongate, Strongyliiform, hairy,
acneo-piceous, with very coaieely punctate
head and prothorai and ooarsdy punctato-
striate elytra, the pro thorax subquadrate
32. ^wcies narrow, elongate, shining, hairy,
resembling ColparAruM, with very laige
eyes in (J, a long, smooth prothor&x.
h,Googlc
New and liule-huntm Lagritdae. 175
metoJlio, ooareely punotato-striate elytn,
with inoonapicuous setigerouB itnpreesions,
aad olavate femora No. 130.
33. Species narrow, elongate, leeembling Hoe-
fHonia, with extremalj largo eyea, very
long, snbeemte antennae, an almost
smooth prothorax, andnigro-lineate elytra No. 131.
Section A.
1. Statlra eoitailMiuls. (Plate XII, fig. 1, anterior leg, S)
Suuyra htMrio, Dej. Cat., 3rd. edit., p. 236 (1837).
Statira costaricensia. Champ., Biol. Centr.-Am., Coleopt.
iv. 2, p. 36 (1889).
^. .Anterior femora greatly thickened, abruptly narrowed and
angnlate near the base, the narrow basal portion armed with a long
curved spine; anterior tibiae hollowed beneath, and twisted, befon
tiie apex; ant«anal joint 11 nearly three times as long as 10.
For. f EUytra more ooaiBety pimotat«-Btriate, the punctures on
the diso transverse, the inteisticee slightly oonvez, the black mark-
ing much more extended and subcoalesceat, the strongly angulate
post basal fascia oontinned down the second intfirstice to beyond
the middle and nearly joining the median transverse mark, the
narrow, oblique subapical fascia reaching the suture and continued
akmg it for a short distance, in front and behind.
Hab. Costa Eica; Colombu (ifus. Oxm.; Mus. Brit.,
ex. coU. Le^erte).
Described by me in 1889 from a mutilated example
(without anterior legs) from Costa Biea. A specimen from
Colombia in the Oxford Museum agrees with this insect,
except that the black post-basal mark on the elytra is
wanting. The variety?, ex coll. Lafert^, from that of
Dejean, ia labelled SktUyra kistrio, mihi; it superficially
resembles iS. vagegutUUa, Pic, and S. conspidlUda, Makl.,
specieB wanting the anterior femoral spine. The anterior
femora themselves are more abruptly narrowed before the
base (appearing angulate) than in the allied S. suturalis,
M&kl.
2. Statin flavoslgnata, n. sp. (Plate XII, fig. 2, ^.)
3- Elongate, depressed, shining, pale reddish -brown, the l^s
and under surface teetooeous ; the elytra each with an oblong mark
near tJic suture at about the middle, two others obliquely placed
iM:,i,A J 1^, Google
176 Mr. G. C. Champion on
b«low this, and two more before the apex {these two coatesoent uid
forming an oblique faaoia), yellow, the base also indeterminatety
flATesoent. Head almost smooth, ahallowly foveate in the middle
betwoen the widely separated eyes; antennae moderately long,
joints 7-10 slightly decreasing in length, 11 about three times as
long as 10. Prothonx oblong-suboocdate, slightly narrower than
the head, as long as broad, almost smooth, the basal ma^in not
much raised. Elytra long, twice as brood as tiis prothorax, gradually
widening to the middle, somewhat acuminate at the apex; finely
punotato^triate, the interstices flat, 3, 6, and 0 with scattered
■etigerous punctures, those on 3 est^iding forwards to near thfi
base, the others placed on the apioal half. Anterior femora much
tiiickoned, abruptly narrowed and angulate near the base, the
narrow basal portion armed with a long curved spine; anterior
tibiae hollowed and slightly twisted at the apex beneath.
Length 8), breadth nearly 3 nun.
Hab. EcuADOB (fiuoUey).
One male. In this inaect the yellow markings on the
apical two-thiids of each elytron are arranged into two
oblique fasciae and an isolated submedian spot, the elytra
thetuselvea are much elongated, and the anterior femora
laid tibiae are shaped as in S. costaricsnsia, ^.
3. Statlra splnlgera, n. sp.
S- ElongKte, rather dull, piceo-aastaneous, the antennae, the
margins of the elylnt. the barsi, and the bases of the temoia, obscnrely
rufeaeeat. Head rather nanow, shining, ahnoat smooth, the eyes
•epaniied by more than half the width of one of them; antamse
comparatively short, moderate stout, joint 11 nearly as long sa
T-IO united. Pruthi>nix lon^^er than broad, oUong, oonatricted at
the base, m wide aa the head. alulaMous. oheoletdy punctolate,
Iwhty i.<au^lii.'ukte c>a the di^ antenorty. and with two obliqiie
duliuctly punvturcd lmpi«Esii«9 in the middle before the base,
the ba&al itun:ut not muoh raised. Elytn long, twice as broad as
lh» pivihorax. iirMludltr vitlaicd to the nuddle azkd somewhat
t»vwUy n«Rvw1^t |«wtw^'rly: fini?{y punclato^triale, the inter-
»!».■«« dJutci.t^'iu, »>iiw»!iat cvionM. iUiut od the diic anteriotly, 3
"•nil »i5 *ia>^^v i^uratvd *Kull «t;j»ivtts punetaro and 5 and 9
»iiS\ t*v>i>t ilitw oi:',rts loHttnb ih<f ape:!. JLattnc« femoi* greatly
thi^-tvunl, ^t'rui'il^v iuttv>»«d ti«ar the bwdc the nanow banl
j>i'tt.\-n atUHil *;s>, a U-n^ t.-uiT«il »[>in*.
I*tv:t!i SI, ^^\*n!■.^ Iff mm.
ll,y,l,7rJM,G00glc
New and litAe-hiown Lagriidae. 177
One male. This species has the elytra shaped as in
S. flavosignata \ but the head aod piothorax are narrower,
the antennae are stouter, and the upper surface is duller
and almost unifonnly fusco-castaneous.
4. Statira MuiUioiiieTa, □. sp.
Hoderatelf elongate, Bhining, piceons or reddiah-bioNni, the darker
example with the antennae in great part, the knees and toiai, and
Uke Butuie of the elytra at the base, ferruginous. Head rather
broad, almost smooth, ehollowly foveate in the middle between
tbe widely separated eyes; antennae long, rather slender, joint II
about aa long aa 8-10 united. Prothorai aubcordate, as broad aa
long, about aa wide as the head, moderate]; constricted before the
base; Bparaely, obeoletelj punctulate, the basal margin not much
raised, the diso foveate in the middle behind in one example. Elytra
moderately elongate, much brooder than the prothorax, gradually
widened to the middle and rapidly narrowed posteriorly, trans-
versely depressed below the base; finely puncta to -striate, the
interstices flat, 3 with six and 6 with four widely separated oon-
RjncnouB Hetigeroua punctures, 9 also with two or three punctures
towaids the apex. Ventral segments 2 and 3 with a few fine pili-
gerous punoturee between the usual double series of setigerous
impreaaions running down 1-6. Anterior femora greatly thickened,
abruptly narrowed and angulate near the base, the narrow basal
portion armed with a long curved spine; anterior tibiae hollowed
at the apex beneath.
Length 8A. breadth 3 mm. dJ.)
Hab. Colombia {Mus. Brit. : type) ; Ecdadob {BticHof).
Two specimens, assumed to be males, the one from
Ecuador smoother beneath and somewhat immature.
Less elongate and more shining than 8. spinigera, the head
and piothorax broader, the antennae much longer, the
setigerouB punctures on the disc of the elytra larger, the
anterior femora angulate before the base, as in iS. costari-
centra and S. flavosignata. The Colombian example was
acquired by the Museum in 1871. S. nigella and S.fueca,
M^., from the same country, may be allied forms ">
6. Statlia sutoialia. (Plate XII, £g. 3, anterior leg, ^.)
Statira mtUTolis, Maid., Act. Soc. Fenn. vii, p. 157 (1862).
{$. Antemiae moderately long, joints 3-10 subequal in length,
11 about three times aa long aa 10.
TRAB8. KST. SOC. LOND. 1917. — PABT L (NOV.) N
,i,A J h, Google
178 Mr. G. C. Champion on
$. Antennae ahorter, joints 4~10 decreasing in length, 11 lees
elongate.
Var. Elytra with two or three dark lines on the disc extending
downward from the base.
,Hab. Brazil, Rio de Janeiro, Petropolis, Constancia,
Tijuca.
A leddish-brown, shining, rather convex insect, usually
with the suture or two or three lines on the disc infuscate,
these markings sometimes wanting; the anterior femora
greatly thickened from near the base, and with the narrow
basal portion armed with a long, curved spine {not noticed
by M&klin] ; the anterior tibiae slightly hollowed before
the apex beneath ; the elytral interstices 3, 6, and 9 with
a few widely separated setigerous punctures. The aedeagns
of the male is long and slender, tapering towards the tip.
S. presuturalis, Pic {No. 81, infra), ftom the same region, is
a very similar form, but it wants the anterior femoral spine.
Section B
6. Stattra denttgers, n. sp..
Elongate, broad, widened posteriorly, Bhining ; nifo-castaneouB.
the palpi pioeous, the elytra aenea-piceous, the latter with scattered,
long, erect, bristly hairs. Head rather small, not so broad aa the
prothorax, almost smooth, foveate in the middle between the
eyes, the latter very Urge, somewhat distant; ontcmnae long,
slender, joint 11 equalling 8-10 united, Prothorax slightly longer
than broad, rounded at the sides, strongly oonstrictcd before the
prominent basal margin; sparsely, minutely punctate. Elytra
long, broad, widening to the middle, and there twioe as broad aa
the prothorax, arcuately narrowed posteriorly, and Bcununats at
the apes; closely, finely punctate -striate, the interstices broad, flat
on the anterior half of the diao, 3, 6, 7, and 9 with a series of rather
large setigerous impressions extending throughout their length,
the impressions (except those towards the base) each preceded by a
rather prominrait tubercle, the tubercles gradually heooming longer,
oariniform, and oatenulale towards the sides and apex, the first
(sutural) interstice also with five impressions near the tip. Legs
[posterior pair wanting] rather stout; anterior femora strongly
olavata, sulcata along their outer half beneath, and also finely oiliate;
tibiae pilose within, the anterior pair armed with a sharp triangular
toolh at one-third from the tip.
Length 15, breadth 6 mm. (ij I)
Nea and littte-knoum Lagriidae. 179
Hab. EcUADOB, Paramba {Rosenberg).
One specimen. The ciliate anterior femora and toothed
anterior tibiae (not necessarily (^-characters) bring this
species neai certain Central American forms, S. gUibrata,
Makl. and S. deMieidata, Champ., placed by me at the
head of the genus in the arrangement adopted in the
" Biologia." Compared with S. tiAercuUUa, Makl., the
antennae are longer and more slender, the inter-ocular
fovea is smaller, the prothorax is htrger and smoother, and
the first elytral interetice is tuberculate (instead of un-
armed) at the apex. The general shape is like that of S.
laiicfiUis, Makl.
7. StaUra gemmUer. (Plate XII, fig. 4, posterior leg, 3-)
Suaira gemmijer, Makl., Act. See. Fenn. vii, p. 147 (1862).
Poeterior femora in each sex with a truncate, exteriorly toothed,
prominence at the baae, and a Bmall triangular tooth at about the
tMsal fourth, beneath (fig. 4) ; antenna! joint 11 in i^ about equalling
7—10, in $ 8-10 united; anterior tarsi slightly widened in ^, all the
teni and tibiae a little more hairy in <J than in $; eyee distinctly
more approximate in i^ than in $.
Hab. Brazil {Miis. Brit., Mtis. Oxon.), Rio de Janeiro
{Fry), Constancia {J. Gray and H. Clark, Jan. 1857), Minas
Geraes [Mm. Brit.), Ilha Santo Amaro near Santos [G. E.
Biyanl ; 23. iv. '12).
This is perhaps the finest known species of the genus.
It has very elongate, sharply acuminate, translucent,
aeneous elytra, with the alternate interstices closely
86 riato- tuberculate and catenulate throughout; the pro-
thorax closely punctulate ; the legs very elongate, slender,
the posterior femora feebly bidentate near the base beneath
in both sexes; the tarsi and tibiae hairy; the antennae
rather slender and infuscate. S. gemmijer may be the
S. regina, Lac, of Dejean's catalogue, and his S. gemmata
a var. of S. gaiitndata, Makl.?. Twelve specimens seen,
iacluding three males. The penis-sheath, so far as visible
without dissection, appears to be symmetric.
8. Statin afioldes. (PlateXU.fig. 5, aedeagus, in profile, 3*-)
Statira agroides, Lep. et Serv., Encycl. M6th., Ins. X. p. 480
(1827); Makl., Act. Soc. Fenn. vii, p. 148 (1862).
iM-,i,A J h, Google
180 Mi. G. C. Champion on
Slatyra agroides, Dej. Cat., 3rd edit. p. 236 (1837).
Statira armata, Makl., op. cit. x, p. 636. (1875)
(J. Aedeogus (fig. 6) : basal piece elongate,' produced into a long
oMt, apiniform prooees infericrly, the stout penis-sheatii Btill longer
and with a sagittifcrm hook projecting from the tip (as seoi in profile).
Var. I Castaneous, the elytra black, the aedeagas of S without
projecting sagittiform piece at the tip ( } withdrawn).
Hab.'SiRkm,{exMua. Dejean; Mus. Brit.; Mus. Oxon.),
Kio de Janeiro, Santa Cathanna (Fry), (Ibnstancia {J.
Gray and H. Ctark), Petropolia, Santa Rita and Boa
Sorta {Dr. Sahlberg: types of S. antMla), Eapinto Santo
(DescouTtils).
This insect seems to be fairly common in Brazil, and is
eaiiily recognisable amongst its allies by the mucronate
apices of the elytra, the catenulate, posteriorly tubercutat«
alternate interstices, 1, 3, 5, 7 and 9, and the simple an-
tennae and legs in the two sexes. The terminal joint of
the antennae about equals B~10 united in 2, and is slightly
longer in J. The colour varies from ferruginous or rufo-
castaneous to piceous; the two examples with black
elytra, from Constancia and Espirito Santo, may belong to
a different species. The elytra in S. agroides are said to
have a " bluish-violet reflection which ought to be more
distinct in the living insect," a character of no importance
in this genus. The imperfectly described S. interrupte-
coatata. Fie,* from French Guiana, seems to be an allied
form.
9. Statin longlceps, n. sp. (Plate XII, fig. 6.)
Very elongate, shining; fusco-testaceouB, the bead and antomae
nifcBcent, the latter with joints 1-^ and 11 darker, the eyce, palpi,
and labnim black or piceous, the prothorax (except at the base)
also slightly infuscate ; elytra with a few long, bristly hairs. Head
oblong, narrow, almost smooth, longitudinally grooved between the
eyes, the latter large, somewhat depressed, well separated ; antennae
long, rather slender, joint 11 as long as S-10 united. Prothorax
wider than the head, much longer than broad, gradually narrowed
from the middle forward, and constricted before the base, the
* A large chitinous tube from which the penis^heath is extmded :
it is usually withdrawn into the body, and not visible without
diaaeetton.
* Melanges exot.-entom. xi, p. 19 (Nov. 1914).
u,y,l,/eJM,G00glc
Neu) and Utde-hnmon Latfriidae. 181
nised bsAftl in&rgin termiDating in a stout tubercle on each side;
the gaiUoB with a few, widely Bcattered, minute puscturee. Elytn
long, gmdually widening to the middle, aomtunate pogteriorlj, the
Bjncee muoroiMte ; cIohIj, finely punctato-Htriate, the striae
sinuate on the disc, the intetBticee 1 , 3, 6 slightly widened and each
with ft series of somewhat closely placed, subquadrate, setigerous
impreeeions extending throughout their length, 7 with a series of
similar impressions down the apical half, and 9 with four othen
towards the tip, the int««pacee between them longitudinally raised
en the disc and tuberculate on the apical declivity. L^s very long,
eimple.
Lsngth 12, breadth 3} mm. ($ 1)
Hab. Brazil, Rio de Janeiro {Fry).
One specimen. A close ally of S. agroides, L. and S.
(^ armaia, Makl.), and with the elytra mucronate at the
tip and very similarly sculptured, differing from that
insect in its oblong, narrow h^, with shallow longitudinal
inter-ocuiar groove, the less prominent eyes, and the
laterally tuberculate basal margin of the prothorax. The
catenulate first (eutural) elytra! interstice and mucronate
apez separate S. longtceps from S. catenata, meleagria,
and others.
10. Statint vemeosa, n. sp.
Very elongate, widened posteriorly, moderately shining; nigro-
piceons, the elytra with a faint brassy lustre; the elytra with long,
erect bristly hairs, the antennae, tibiae, and tarsi pilose. Head long,
narrow, almost smooth, shallowly depressed in the middle between
the eyea, the latter large, moderately distant; antennae long,
slender, joint 11 in ,£ about equalling 7-10, in $ 8-10, united.
Protiiocax long, narrow, the sides obliquely convei^iTng from about
the middle forward, and constricted before the raised basal margin ;
cloaely, finely punctate, depressed in the centre at the base and
longitudinally excavate behind the anterior margin. Elytra very
long, widening to beyond the middle and there about three timee
the breadth of the prothorax, mucronate at the tip; closely, rather
findy, crenato -striate throughout, the striae sinuate and arranged
in pairs, the alternate interatioee each with a series of approximate
oblong, flattened spioulae, which become more prominent and
earinifoim towards the sides and apex. Legs very loi^.
LengOi 13, breadth nearly 4 mm. (<;$.)
Hab. Upper Amazons, Ega {H. W. Sates).
u,y,l,7rJM,G00glc
182 Mi. G. C. Champion on
Two specimens, assumed to be sexes, one having a longer
apical joint to the antennae than the other. Closely ielat«d
to S. agraeformis. Champ., from Panama (figured in
B.C.-Am,, Coleopt. IV, 2, pi. 1, fig. 7), the head not so broad,
the eleventh antennal joint in (J shorter, the elytra with
narrower, smaller tubercles (and the striae, in consequence,
less sinuous), the legs infuscate, etc. S. agroidea, S. gem-
mijer, and S. catenata are somewhat similar forms, all of
them superficially resembling the Carabid-genus Agra,
which is numerous in species in the same regions.
11. Statlra mucronata, n. sp.
Elongate, widened posteriorly, Bhiiiiiig; vaiying in oolour from
nigni-piceoiu to outaneouB, the anteonae more or leaa infoHcate and
with at leaat the apical joist ferruginous, the elytra in the darker
examplee braaay or greenish -aeneonB ; the elytra with numerous
bristly hain. Head rather closely punctate, deeply, broadly
foveate betwerai the eyes, the latter very large, somewhat narrowly
separated ; antennae stout, moderately long in S< shorter in $,
joint 11 in ^ equalling 0-10, in $ 8-10, united. Protborax longer
than broad, slightly narrower than the head, feebly rounded at the
sides, oonstricted before the prominrait basal margin; closely,
conapiouously punctate, the interspaces polished, the disc trans-
veraely depressed on each side anteriorly and also excavate in the
centre at the baae. Ellytia long, widening to beyond the middls,
more Uisn twjoe the width of the prothorax, mueronate at the tip;
finely, deeply punota to -striate, the striae strongly undulate towards
the sides and apex, the interstices convex, 3, S, 7, and 9 throughout
their length, and 1 at the apex, with a scattered series of setigeioua
impressions, the spaces between them longitudinally tubereulate,
the tubercles on 7 and 9, and those towards the apex on 1, 3, 6,
shorter and more closely placed than those on the disc Femora
moderately thickened.
Length ia-12, breadth 31-4} i™- (o'$-)
Hab. Brazil, Ceara {Gmnelle).
Three males and three females, two only with metallic
elrtra. Verj- like S. nigrocofntUa, but with the alternate
elrtral interstices 3, 5, 7 and 9 here and there tuberculate
throughout their length, the tubercles on 7 and 9 shorter
and more numerous, much as in iS. tvhercuUUa, Makl.; the
prothorax longer and a Uttle more finely punctate; the
antennae slightly stouter.
Ij.y.l.Ajl^yCOO'^lL'
New and litUe-knoum Lagriidae. 183
12. SUtin nlKroeMTOlea, n. sp.
g. Elongate, broad, widened pmteriorly, Bhining; nigro-piceoos,
llie derentfa antainal joint, the tipa of the toisi, and the head in
one spedmcn, fenuginons or reddish, th« eljij^ blue or bluish-green ;
the elytra wiUi a few bristly hairs. Head bioad, finely pnnctate,
deeply, longitudinally impressed between the eyes, the latter very
Urge, Bomewbat narrowly separated ; ant^mae rather long, slovt,
joint II equalling 7-10 united. Prothorox as wide as tbe head,
about as broad as long, feebly rounded at the sides, moderately
eonstricted before the laterally-piojeGting raised basal margin;
cloaely, ratber coarsely punctate, the intenpacee polished, the dise
depressed in the centre at the base and also on each side anteriorly.
Elytra long, more than twice the breadth of the prothorax, widening
to beyond the middle, transversely depressed below the base, the
apioee mucconate; closely, finely, deeply punctato-striate, the striae
crenate and posteriorly undulate, the interaticee convex throughout,
3, 5, and 7 with from three to five deep setigerous impreasions
towards the apex, 9 with a scattered series of similar impteHsions
along their entire Imgth, and I with three others near tiie tip, the
spaces between them longitudinally tuberoulate and suboatenate.
Iiep long, the femora moderately thickened.
Length II-IU, breadth 31-4 mm.
Hab. Bbazil, Jatahy, Province of Goyaa (Pujol, ex coU.
fry)-
Two specimens, one with the tip of the aedeagus exposed,
agreeing veiy nearly with the description of the Brazilian
iS. nigra, Makl. ; but as the author aays nothing about the
stout antennae, ot the mucronate apices of the elytra,
and states that the brood inter-ocular impression is almost
obeolete and the eleventh antenna) joint (c^) about equals
6-10 united, the Jatahy insect must be treated as distinct.
The allied S. caeUUa, Er., from Peru, should have a smoother
prothorax. 5. pertiana, infra, has more slender antennae;
a non-foveate l^ad, the apices of the elytra obtuse, etc.
13. Statin Intesi, n. sp.
J. Elongate, rather narrow, shining ; piceous, the elytra with a
brassy lustre, the apical joint of the antennae ferruginous ; the
elytra with numerous long bristly bails. Head slightly broader
than the prothorax, closely, finely punctate, with a deep oblong
fovea between the eyes, the latter very large, somewhat narrowly
separated; antennae rather slender, long, joint 11 equalling 6-10
united. Prothorax rather narrow, oblongo-coidate, oonatrioted
iM-,i,A J h, Google
184 Mr. G. C. Champion on
before the prominent hind angles, the basal margin raised, the disc
depressed in the centre behind ; sparselj, finely punctate. Ely^a
long, gradually widened to the middle and there twice as broad as
the prothorai, transversely depressed below the base, acuminate
posteriorly, mucronat« at the tip; closely, finely, deeply punctoto-
striate, the strifte undulate, the interstices more or less convex, 3,
5, 7 and 9 with a scattered series of setigerous imprcBsions, each
of which ia preoeded by a prominent tubercle, 1 also with three or
four tubercles before the spex, 7 and 9 catenulate throughout. Legs
slender [anterior pair wanting].
Length 10^, breadth 3^ mm.
Hab. Upper Amazons Ega {H. W. Bates). '
One male. ThU species agrees with S. (uhercwiota,
B mucronala, and S. nigrocaendea in having the bead
deeply foveate between the eyes and the apices of the
el)i;ra mucronate, differing from the first-named in the
convex iiitetEtices, smaller foveae, prominent tubercles,
and more strongly mucronate apex of the elytra, and from
the two others in its narrower shape, less thickened antennae,
and the more prominent small scattered tubercles on tbe
disc of the elytra.
14. Slatlra euspldata, n. sp.
Elongate, somewhat robust, narrow, widened posteriorly, shining ;
piceouB, the antennae (joints 1-4 excepted) femiginous, the elytra
metallic golden-green, tlie colour changing to golden and cupreous
laterally, and to green across the base ; the elytra with a few long,
briatly hairs. Head about as wide as the prothorax, cloGely, minutejy
. punctate, slightly hollowed betweoi the eyee, the latter large,
somewhat distant; antennae slender, moderately long, joint 11
about equalling 7-10 united. Prothorax considerably longer than
broad, subcylindrical, constricted before the raised, lalerally-pro-
jecting basal margin ; densely, minutely punctate, the diso longi-
tudinally depressed in the centre behind. Elytra long, widening to
the middle and there considerably more than twice the breadth of
the prothorax, acuminate posteriorly, the apices mucronate ; closely,
finely, deeply punctato-striate, tlie interstices more or less convex,
3, 6, and 9 each with a seriee of small setigerous impreesions ex-
tending from a little below the base to the apex (those on 3 closely
placed on the apical half), 1 and 7 atso with three or four impres-
sions before the tip, the spaces between them becoming m6re or leas
raised longitudinally or tuberculate towards the sides and apex.
Length 10, breadth 3) mm. ($ T)
Ij.y.l.Ajl^yCOO'^lL'
Nob and litde-htoum Lagriidae. I8B
EtA. LowsB Amazons, Para {H. W. Batex, ex coU.
Patcoe).
One example. Not xmlike the Peruvian S, vigifUi-
punetaia (No. 46), but with a subcylindrical, subsulcate,
densely, minutely punctate prothorax, and more nnmerous,
much emallei setigerous impreasions on the elytra, those
on the apical half of the third interstice somewhat closely
placed, the elytral apices mucronste. The longer and
narrower prothoraz, smaller head, deeply striate elytra,
with more numerous smaller setigerous impressions in
the iaterstices 3 and 5, separate S. cuspidata from S. semi-
cuprea (No. 47). The sides of the elytra are brilliant
cupreous towards the apex in the present species.
15. Staflnt tubereoUtta.
Staiira tuberculata, Makl., Act. Soc. Fenn. x, p. 637 (1876).
Elongate, ehining; dilute rufo-castaneous, the bead and pro-
thorax often moie or leas iofnacate, the patpi, and sometimee the
antoinae also, piceoua, the upper Burface with a faint brassy luitie
(in Hliklin's var. a the elytra are aeneous or green); the elytra
with scattered long, bristly hairs. Head about as wide as the pro-
Uioiax, almost smooth, deeply foveate in the middle between the
eyes, the latter large and narrowly separated in ^, a little smaller
and more distant in $ ; antennae rather slender, joint 11 in [J about
equalling 5-10, in $ slightly longer than 8-10, united. Piothor&x
a little longer than broad, moderately rounded at the sides, con-
stricted befora the prominent basal margin ; finely punctate, often
with one or two oblique impiessiona on each aide of the disc. Elytra
long, twice as wide as the prothorax, very slightly widening to the
middle, pointed at the apex; finely punotato -striate thiougbout,
tlie striae undulate, the interaticee 3, 6, 7, and 9 each with a scattered
•eoriea of lat^e, deep, subquadr&te, setigerous impressiona, separated
towards the sides and apex by elongate tubercles, those on 7 and 9
shorter and more prominrnt, the eighth interstice nanow and
cariniform. Anterior femora strongly clavate, feebly ciliate.
Length 8<i-l^i' breadth 2i-3i mm. (,?$.}
Hob. Brazil, PetropoUs and Santa Rita {Saklberg :
typee), Rio de Janeiro, Bahia (Fry), Espinto Santo {Des-
courtils), Alto da Serra in San Paulo (Bryant).
Eleven specimens (9 <J, 2 ?) before me are referable to
this species, two of them belonging to the colour var.
a of Maklin. There is also some variation in the shape
iM-,i,A J h, Google
186 Mr. G. C. Champion on
and puncturing of the prothorax, and in the number of
tubeicles and setigeroua impressions on the elytra, the
latter being always large and deep. The apices of the
elytra are pointed or Bubmucronate. The frontal fovea is
also deep. The sexual characters were not mentioued
by the author.
16. Stetira unetarenuw, n. sp.
Very elongate, narrow, ebining; piceoos, the elytra giectuih-
aeneous, the antennae in their outer half and the legs nifo-tcetacwoi
or ferruginous, the knees slightly infuscato. Head rather small,
sparsely, finely punotate, unimprMsed between the eyes, the latter
la^e, narrowly Beparat«d ; antennae moderately long, rather simder,
pilose, joint II equalling 8-10 united. Prothorax longer than
broad, a little wider than the head, feebly rounded at the sides, and
constricted beforo the raise)), laterally-projecting basal margin;
closely, finely, oonspiouously punctate, the disc transversely im-
pressed on each side before and behind the middle. Elytra very
long, twice as wide as the prothorax, somewhat rounded at the side^
gradually widening to the middle, without mucro at the tip; closdy,
finely punctate -striate, the interetices feebly convex, 1, 3, fi, 7, and
9 each with a series of rather small, deep, setigerous impieeeioas
extending throughout their length (olosely placed on 3 and 6, and
scattered on 1, 7, and 9), the spaces between them longitudinally
raised or tuberculate, the tuberolee elongate on 7 and 9. Tibiae
pilose within.
Lengthlli.broadthSlmm. ($1)
Hab. Amazons, Santarem (H. W. Bates).
One specimen, now wanting the elytral setae. Smaller
and less elongate than S. verrucosa, the head narrow, the
prothorax not excavate in front, the elytra narrower, with
smaller elevations on the alternate interstices, the striae
straighter, and the apices unarmed.
17. Statira longleollis. (Plate XII, fig. 7, posterior leg, 3-)
$. Statira longicoUis, Makl., Act. Soc. Femi. vii, p. 151
(1862).
<J. Lower surface of the anterior femora at the base, and that of
the other femora to near the apex, and a broad space down the
middle of the metastemnm and abdomen, thickly clothed with
long, fine, projecting or eemi-ereot, hairs ; antennae cloeely piloee.
ih,Cooi^lc
Nob and Uule-lmown Lagriidae. 187
joint 1 moderately thickened, 10 much shortcT than 9 and angolarly
dilated at the inner apical angle, II {as in $) nearly equalling 8-10
united; anterior tibiae gradually dilated on the inner aide into a
broad, anbangiilar, concave plate; intermediate tibiae hollowed
rithin; posterior tibiae (fig. 7) ezcaTat« along their inner face,
broadly arcuato^emaiginate towards the middle (as seen from above),
and nidoied thence to the apex, appearing strongly sinuate within.
Bah. BRAzn. (Mm. Oxon.), Rio de Janeiro {Fry).
Eight specimens seen, three of which are males. U
correctly identified by me, S. lotigicf^lis is a close ally of
S. mndipennia, Lep. et Serv. (and not of S. geniadata, as
stated by Maklin), from which it differs in the narrower
liead and prothorax in both sexes, and in the following
(^-characterB : — antennae with joint 1 less thickened, and
10 shorter and dentate at the apex within ; intermediate
femora ciliate to near the apex and the ventral surface
more hairy; posterior tibiae strongly sinuate within,
the plate on the anterior pair less angular. The number
and arrangement of the setigerous impressions on the
elftral interstices 3, 5, and d are much the same in the
two species. The colour is variable — ^piceoua or rufo-
CBstaneous, the eljrtrs with an aeneous or greenish lustre,
the two basal. joints of the antennae sometimes infuscate.
18. Statlra mslsagris. (Plate XH, fig. 8, ^.)
ij. Statira meUaffrie, Makl., Act. Soc, Fenn. vii, p. 149
(1862).
Antenna] joint 11 in ^7 equalling 7-10, in $ 8-11), united; inter-
mediate femora in i^ gradually thickened outwards and then abruptly
hollowed before the apex beneath.
Hab. Brazil, Rio de Janeiro {Fry, Mva. Oxon.), Kspirito
Santo {Mm. Brit.).
Eight examples seen. A very elongate, pallid form
allied to S. catenaia, Makl., with the widened, catenulate
alternate elytral interstices 3, 5, 7, and 9 each bearing a
series of somewhat closely placed, subquadrate, black
impressions; the legs and antennae differently formed
or clothed, and the abdomen almost glabrous (the usual
setae only present), in the <J ; the head narrow and sulcate
between the eyes.
iM-,i,A J h, Google
188 Mr. G. C, Champion on
19. Statin oalenata. (Plate XII, figs. 9, ^i 9a, b,
penis'sheatli, ^.)
?. Statira caUnata, Makl., Act. Soc. Feim. vii, p. 148 (1862).
j. Statira plumicomis, Deyr. in litt.
(J. Antennae thickened, joint 1 very stout, 2 quite short, S-10
rapidly decreasing in length, 9 and 10 dentate at the inner apical
angle, 11 very elongate, equalling 7-10 united, 3-10 wth a dttm
fringe of long hairs within ; anterior femora Btnmgly incraMatc^
oiliate at the base beneath i anterior tibiae broadly, angolari;
explanate towards the apex within; int«Tnediate femora oiliate
beneath i intermediate tibiae hollowed along their inner faoe;
posterior femora very etout, ouired, excavate and densely ciUatc
beneath, angularly dilated before the apex ; posterior tibiae sinuond;
bowed, broadly widened in their outer half, excavate and pubescent
within, and furnished with a dense brush of very long hairs at about
the middle of their upper inner edge; metaatemum and abdomen
thiekly clothed with long hairs down the middle; penis -sheath '*
stout, asymmetric, broadly, obliquely, subangularly dilated at the
apex, serrulate along the upper edge.
$. Antennae, slender, simple, joint II ncMly or quite equalling
3-10 united.
HcJ). Brazil, Rio de Janeiro, Santa Catbarina (Fry),
Constancia {/, Gray and H. Clark, Jan. 1857), State of ^
Paulo (QouneUe), Minaa Geraea {Mtis. Brit.).
Nine specimens seen, including three males. The extn-
ordinary combination of characters in the atracture or
clothing of the legs, antennae and ventral suif&ce in this
sex makes S. caienata easily recognisable, at least in the
(J; the ?, however, is very like that of S. geniadata, Maid.,
differing from it in having the alternate elytral interstices
a little broader, 5 and 7 nith more numerous setigerous
impressions. The amount of catenulation of these inter-
stices is somewhat variable in both iasecfaB.
20. Statin rirldlpennls. (Plate XII, figs. 10, lOo,
aedeagus, ^.)
Statira riridipmnis, Lep. et Serv., Encyci. Mdth., Ins x,
p. 480 (1827) ; Makl., Act. Soc. Fenn. vii, p. 149 (1863).
" The oonjoined lateral lobes of the tegmen, seo. St»Tp.
Ij.y.l.AjL.yCOO'^lC
Neu> ani Uoie-known Lagriidae. 189
(J. Antennae a little tbiekcr than in $, joint I Very stout, 10
dcotete at the apex witUn, 11 aeorlj equalling 7-10 united, 3-10
thicklf pilose on their inner side ; anterior femora very stout ;
Ulterior tibiae angularly eiplanate towards the apex within ; inter-
nudiate tibiae hollowed along their innor face; posterior femora
doMdy cUiat« Along their basal half bmeath; paetarior tibiae
Bicarata wiUiin, widened outwards, and broadly, shallow]; emarg^-
ate »t about the middle, appearing sinuate on their inner edge;
metastemum and abdomen thickly pilose down the centra, the fifth
Bcgment bntadly depressed in the middle posteriorly.
Aedeagus {figs. 10, lOa] : basal pieoe stout, curved, boat-shaped ;
pcnk-eheath asymmetric, twisted, obliquely dilated outwards into
U e)ongat«, apoon-ahaped piece, which is angulafe on the right aide
at some distonoe before the tip.
ffoA/BKAZiL {Mua. Brit., Mus. Oxon.), Rio de Janeiro
[fry), Constancia and Tijuca {J. Gray and H. Clark,
Jan. 1857).
The long senes of this species before me vary in colour
from piceouB to rufo-testaceous ; the prothorax and an-
tennae (joints 1 and 2 excepted) are usually red; the
elytra with a tranalucent green, aeneoue or cupreous lustre ;
the legs in most of the specimens testaceous or rufo-
Ustaceoufi, the knees and tarsi sometimes infuscate. The
absence of the setigerous impressions along the seventh
elytral interstice separates both sexes of 5. mridi-pennis
from the very closely allied S. geniculata, Makl.
21. Statint Eenionlata. (Plate XII. figs. 11, 11a, aedeagua, <?.)
^.Statyra gemmata, Dej. Cat., 3rd edit., p. 236 (1837).
StaUra geniotdata, l^kl., Act. Soc. Fenn. vii, p. 150
3- Antennae very long and slender, joint 1 scarcely stouter than
in S, 10 angulate at the inner apical angle, 11 nearly equalling 8-10
nnited; eyes extremely large, subcontiguous; anterior femora very
■tout, almost glabrous; posterior femora ciliate beneath; inter-
"Kdiate and posterior tibiae closely pilose within, the latter Bimple;
■iietagtemiim and abdomen thickly clothed down the middle with
long, erect, fulvous hairs, the fifth s^ment broadly depressed in the
"Wtm posteriorly. Aedeagus (figs. 11, 11a): basal piece long,
curved 1 pcnis-Bheftth dongate, broad, twisted, deeply grooved,
"w apical portion fiddle-shaped.
iM:,i,A J 1^, Google
190 Mr. G. C. Champion on
Hab. Brazil (ex Mas. Dejean; Mua. Oxon.), Rio de
Janeiro, Bahia (Fry), Espirito Santo {Mus. Brit. ; Deectmr-
tila).
Numerous specimens seen, these varyio^ greatly in the
colour of the body and legs, and in the mtensity of the
metallic suffusion of the elytra ; the uitennae are osoaUv
rufo-testaceous with the basal two or three joints infuscate,
rarely black in their basal half. This species, if correctly
identified by me, is closely related to S. viridipennis, difier-
ing from it in the more ot less catenulate elytral iutersticas
1,3,5, 7, and 9 (the catenulation on 7 extendmg along their
entire length), and in the simple anterior and posterior
tibiae, and the slender basal joint of the ant«nnae of tbe
male. The flddle-shaped outer portion of the penis-sheath
(= tegmen), too, is characteristic of the present insect,
22. Statira asymmBtrioa, n. sp. (Plate XII, fig. 12,
penis-sheath, ^.)
cJ. Very elongate, narrow, shiningi pioeoua, the elytra with »
tranalucent aeneoua lustre, the head, antennae (the slightly infiu-
oate basal joint ezcepud), legs, and under surface testaceous or
rufo-t«staoeooa, the elytia with a few long bristly hain. Head
rather small, short, almost smooth, the eyes large, narrowly separ-
ated; antennae very slender, extremely elongate, joint U aboal
equalling S-10 united. Frothorax wider than the head, subooidaU,
Bcaroely longer than broad, the basal maigin prominent ; alutaceous,
spatsely, minutely punctate. Elytra very long, subparallel, Icm
tiian twice the width of the prothorax, rounded at the tip; doasly,
finely punatato-striate, the inteistioee flat on the disc, 3 widentd
and with a series of about twelve, and S and 7 with from 8-10, deep,
subqusdrate, setigerous impreesions.d also with several othera down
the apical half, the spaces between the impressions longitudinally
raised and cateoato-tuberoulate from about the middle to the apei-
Metastemum and ventral s^ments 1-6 hollowed and thickly pihee
down the middle. Penis-sbeath long, broad, asymmetrio, twisted,
arcuately dilated on the right side at some distance before the tpa,
the apical portion spoon'Shaped. L^^ very long; anterior femon
thickened ; posterior femora closely ciliate in their basal half be-
neath; posterior tibiae sinuously compressed, appearing hollovtd
from a little below the base to near the apex, closely pilose vithin.
Length 10, breadth 2} mm.
H(A. Bb&zil, Bio de Janeiro (Fry).
One male. A close ally of S. genwulata, M&kl., difienug
New and UoU-kmnim Lagriidae. 191
from the corFesponding sex of that species in having a much
smaller, Bhoit«r head, a less elongate piothorax, more slender
antennae, sinuously compressed, haiiy posterior tibiae,
and the broad penis-sheath dissimilarly formed. The non-
dilated posterior tibiae, etc., separate S. aaymmarica from
S. tortipes, S. arcuatipes, and other forms with bowed ot
twisted tibiae in the male.
23. Statlnt tortipes, n. sp. (Plate XH, figs. 13, posterior
leg; 13a, penis-sheath, (^.)
Very donate, narrow, moda«tdy ahising; otMcaie teataceona,
the eyes blkck, the elytra in some apecimeoa with a faint aeoeoua
luatie, the latter with a few very long, briatly hairs. Head small,
aJrooBt smooth, obeoletely aulcate between the eyes, the latter large
and modnately diataot; anteonae long, alfliider, joint 11 in (J
equalling S-10 united, very little Bhortcr in $. Prothorax wider
than the head, oblongo .cordate; very sparsely, obeoletely punctate,
slightly depressed in the middle at the ttase. Elytra elongate, aub-
parallel ; closely, finely punot«t« .striate, the interstices almoet flat,
3 with a series of about eight to ten, and S with five or six, setigeions
impressions, 9 also with thiee impressions near the tip, 3 and 5
c*t«Da]at« posteriorly.
^. Femora ciliate beneath, the anterior pair very stout, the inter,
mediate pair clavate, and the posterior pair angularly dilated towards
the apex; posterior tibiae abruptly, bisinuately twisted, broadly
dilated, concave and thickly clothed with fine hairs within, appear-
ing closely ciliate along their inner upper edge (fig. 13); metaster-
num pilose down the middle, the ventral segments 1-3 and 5 (except
on the basal half) also with numerous long erect hairs down the
centre ; penis-sbeath (fig. 13a) twisted, asymmetric, angutate on the
Irftside, the apical portion somewhat shovel .shaped.
$. Hetast«TQum pilose down the middle; anterior and inter-
mediate femora ciliate at the base.
Length 10-11, breadth 3-3^ mm. (,}$.)
Bab. Brazil, Ceara {Gounelie : <J$), Rio de Janeiro
(fry : $), Alto da Sierra in San Paulo (G. E. Bryant, 16.
iii. '12 : $).
One male (somewhat injured by an Ayuhrewts) and
three females from Ceara, and a female from each of the
other localities. An immature-looking insect related to
S. longiccUis, Makl., with a shorter head and prothorax.
smaller eyes, etc. ; the posterior femora angularly dilated
iM-,i,A J h, Google
192 Mr. G. C. Champion on
(as in S. catenata), and the po8t«riot tibiae abruptly, biu-
nuately twisted, pilose, and dilated, in the i^, the tibiae
moie strongly sinuate and more hairy than in the same
sex of iS. arcualipes and S. fusdtarsis, these latter more-
over wanting the pilosity down the middle of the abdomen.
The penis-sheath is asymmetric, and shaped somewhat a«
in S. virtdipennis and its allies.
24. SUtira arouatipei. (Plate XII, figs. U, <^; 14a,
aedeagus, ^J.)
c^. Slatira arcuatipes. Pic, Melanges ezot.-entom. iv, p. 14
(Sept. 1912).
Very elongate, rather narrow, the head uid prothorax dull, the
rest ol the upper surface moderately Bhining ; piceous or nigro-
pioeous, the dytm bluish-green or green, sometimea with cupreous
reflections, the antennae often in part fermginoua; the eljtra with
a few, long, bristjj haira. Heed almost smooth, feebly groOTed
betweoi the eyes, the latter very large and somewhat narrowly
separated; antennae long, slender, shorter in $, joint II in ^ equal-
ling 6-10, in $ about 8-10, united. Frothorax oblongo-ooidate,
rather sparsely, minutdy punctulate. Elytra long, aubparallel in
tkeii basal half in d ; closely, finely puncta to -striate, the inteistiMa
simoet flat on the disc, 3 and 6 tuberculato-catenate towards the
apex, 3, 6, and 7 (thoea on 7 pres^it in one ij-specimen only) with
•ereral widely scattered setigerous impressions, and 9 with three
impressions near the tip.
,}. Intermediate femora gradually widened to near the apex, and
abruptly hollowed thence to the tip, the angle thus fanned clothed
with short hairs externally; posterior femora towards the apexvwy
broadly, arouately dilated, and deeply sulcate beneath, gjabroua;
posterior tibiae sinuously twisted, broadly dilated, hollowed and
pubescent within ; penis-sheath symmetric, long, comprMsed, the tip
triangularly dilated as seen from aboTe, hooked beneath, the tube in
which it is enclosed produced into a long spiniform process on each
Var. Femoraand tibiae, except at their apicee, testaceous. ([^$.)
Length lO-lli, breadth 2i-3^ mm. <iJ?.)
Hab. Brazil. State of San Paulo [type], PetropoUs [J. Gray
and //. Clark, Feb. ISJi). Rio de Jaaeiro (Fry).
Seven males in the Fr}' collection, including two of the
variety with pallid femora and tibiae, are undoubtedly
referable to ^. iireHatipe.s, Vx; and throe females with
l„y,l,Ajh,COOl^lC
Xet and lAde-bmni Lt^idae. 19S
fiimilailr ooknued legs, two of them from Petropolis. most
alao belong beie. These insects were labelled by Fry aa
the sexes of the same species. The variety nearly agreea
with the description of S.fuieitanu. Makl., cf. infn.
25. Statin tudtanis. (I^ate Xn. fig. 15, posterior leg, <5.)
°. Slatva fuacitartis. Maid., Act. Soc. Fenn. x, p. 638
(1875).
Very like 8. aremUipeg, Pic, the bead and piothoiax nigro-pweooB,
the eljtr» brilliant metallic green or braasy-green, the femora (eio^t
Kt Ute apex in one example, $) and tibiae dear rafo-teetAoeoua (o)
or teetAoeoDS ($) ; the apical ant<nnal joint of ^ still moie etongale,
eqnalliiig fi-lO united; the tyta aa large aa in tliat spedee; the
elytial intasticea 3 and S with from three (o five widely separated
setigecouB impreeoicmB down the diac and both tabercnlaU>.catcnate
on the apical decliritjr ; the intennediate and poaterior femois, and
the poeterior tibiae, ebaped exactly aa in the 3 o^ ^- amiatipe*,
except that the posterior tibiae (Gg. Ifi) bear a daiae brush of hairs
at about the middle of the concave inner (ace ; the peDi8.sheath (ao
far as vioit^) shaped aa in S. oreuatipea.
Hab. Brazil, Rio de Janeiro {Fry, J?}, Petropolis
(Dr. SaMbery : type).
A pwr from the Frv- collection are provisionally referred
to this species, the type of which was captured at Petropolis.
The male of S. calenato has a somewhat similar brush of
much longer haiis on the posterior tibiae. S. fugcUarais is
said to have four additional setigerous impressions on the
seventh elytral interstice, and those on the disc large, but
too much importance need not be placed on these characters.
Maklin ignored, or overlooked, the marked sexual peculiari-
ties of many of these insects, and it is therefore impossible
to certainly identify some of his Statirae from the descrip-
tions alone.
26.' StaOra tibialis.
cJ. Siatira tibialis. Pic, M^iangea exot.-entom. iv, p. 14
(Sept. 1912).
,J. Antennae rufo-testaoeoas, with joint 1 1 equalling 6-10 united ;
intermediate femora gradually widened to near the apex, and hol-
lowed thence to the tip, concave along their lower face, glabrous;
posterior femora inloate beneaUi, gradually, arouately dilated on
TRAKS. EST. SOC. WIND. 1917, — PART I. (NOV.) O
iM-,i,A J h, Google
194 Mr. G. C. Champion on
theic lower edge to near the apex ; posterior tibiae aimply sinuate,
broadij dilated, hollowed and spanely pubeaceot within; elytia
metallia green, theint«iBtioe8 3 and 5 with four or five widely eepar-
ated setigeroufi impreasione, becoming taberculato-oatenate towaidi
the apex; penia-aheath simplj acuminate at the tip.
Hab. Brazil, Salobro [type], Ilha Santo Amaro neat
Santos (G. E. Bryant : 4. iv. '12).
A male found by Mr. Bryant ia perhaps referable to this
species, a very close ally of S. arcualipes, according to its
describer. The shape of the penis-sheath (examined in four
out of the seven males seen of the last-named insect) shows
that S. tibialis cannot be a form of S. arcualipes.
27. Statira rimpUdpu, a. sp.
"Very like S, arcuatipet, Pio, the eljtra relatively narrower, nib-
parallel in <J, green or biaaaj, the prothorsx also with an uneoua
luatie, the antennae (the baaal joints exoept«d) testaceous ; antennal
joint II in ^ nearly equalling 7~I0, in $ 8-10, united ; cyee sli^tly
smaller ; prothorex a little less rounded at the aidea, Bubc}riindrieal
anteriorly in some speoimons, distinctly punctate; elytra! inter-
atioee 3 and S with more numerous aetigeious impressions (3 with
fromten to twriTeand5witfaaboutBix),oateiiato-tuber(nilateat the
apex ; legs long and slender, those of the ^ simple, as in $ ; ponia-
sheath ot <J acuminate at tip.
Var. The femora and tibiae, the knees excepted, testaceous.
Length »-I0|, bieadth 2H nun. ((J$. )
Hab. Brazil [Mtis. Brit., Mua. Oxon.), Petropolia and
Conatancia (J. Gray and H. Clark, Feb. 1857 : var.), Sio
de Janeiro {Fry : type).
Eight specimens, three only belonging to the dark-legged
form, one of each of them acquired by the Britbh Museum
in 1871. Extremely like S. arcuatipes, and with the legs
varjTng m colour in the same way ; the legs simple in the
two sexes, the apical joint of the antennae mucn shorter
in the S< >uid the aedeagus not dilated at the tip. Compared
with 5, amoena, Makl., which also haa simple intermediate
and posterior lefjs in the J, the mote numeroua setigerous
impressions on the third and fifth elytral interstices, and
the less elongate apical joint of the j-antenna, will serve
to distinguish the present species. These three forms occur,
with many others, in the vicinity of Rio de Janeiro.
ih,Cooi^lc
New and little-known Lagriidae. 196
28. Statin unoena.
9. Stalira amoena, M&kl., Act. Soc. Fean. vii, p. 152 (1862).
Vw; dong^te, narrow, Bhiniiig; piceous or nigro-piceous, paler
benefttb, the Apical antomal jomt UBnall^ femiginous, the elytra
translucent green, aeaeouB, or ftmeo-cupieooB, the latter with
scattered, long, briatlj' hairs. Head rather small, almost smooth,
the ejea very laige and Habapproximate in ^, more distant in $;
antennae long. Blender, joint lOsnbtriai^nlar and 11 about equalling
6-10 united in ,?, 11 aa long as S-10 in $. Frothorax oblongo-
coidate, oloaelj, minutely punctiilat«,tiie basal margin very promi-
nent. Elytra long, subpaiallel in their basal half, finely punctato-
atriate, Uie interstices flat, 3 with about eight or nine and 5 with sis
or sevea setigerous impressions scatteied between the baee and apex,
and 9 with two or tliree similar impreesiona near the tip, the spaoea
between Uiem longitudinally raised or tubeniulate on the apical
declivi^. ^. Anterior femora thickened, glabrous ; intermediate and
poetcnor femora gradually widened to near the apex, and hollowed
thenoe to the tip; posterior tibiae closely pilose along their outer
half within.
Length 10-12, breadth 2{-3 mm. (<;$.)
Bab. Brazil, Rio de Janeiro (Fry).
The above description ia taken from four males and two
females captuied by Fit. It is one of several extremely
closely allied Brazilian fonna, the females of which are
scarcely distinguishable inter se, though the males poaseas
marked speci&c characters in the structure of the legs, etc.
&laklin*s descriptioii of S. amoena must have been taken
from a ?, and it would apply almost equally well to the
same sex of S. arcuatipes. Pic.
29. Statira mieans.
Suutfra morbiOosa, Dej. Cat., Srd edit., p. 236 (1837).
(J. Stotira micans, Mftkl., Act. Soc. Fenn. vii, p. 153
(1862).
Exttemdy like S. amoena, Makl. (as here identified), bat with
several setigerons impressions on the seventh elytral interstice (alto-
getliei wtuiting in 8. amoena), 3, fi, 7, and ft tuberculato -catenate
tow&rds tJie apex; the head longitudinally grooved or impressed
between the eyee, the latter not bo large in ^ ; the Seventh anlennal
joint of 3 nearly equalling 6-10 united ; the 1^ variable in colour,
simple in 3 ; the aedeagus narrow, acuminate, the sheath straight,
tnincato at tip.
iM:,i,A J h, Google
196 Mr. G. C. Champion on
Sab. Brazii. (ex coU. Dejean), Rio de Janeiro (Frif),
Bahia (Reed), Corcovado (G. E. Bryant).
Seven examples seen, the one from the Dejean collection
being labelled with the MS. name S. morhUlosa. Compared
with the variable S. geniculata, it ia a little smaller and lesB
elongate, the setigerous impressions and tubercles on the
eljrtral interstices 3, 5, 7 and 9 are reduced in number
(especially on 7) ; and in the ^ the abdomen is glabrous
down the middle, the legs are simple, and the penis-sheath
of a totally different shape. Maklin does not mention the
longitudinal inter-ocular groove (also present in his S, rufi-
frons), and the identification of the insect before me with
his species is not certain.
30. Statlra formou, n. sp.
1^. Very elongate, rather narrow, ahining; pioeoiu, the elytra
brilliaiit golden-greMt, cupreous along tbe apical maigin and on the
humeral callus ; the elytra with numerous, and the head and abdo-
men with a few, long, fine, enot bristly hairs. Head rather narrow,
not wider than the piothoiax, with afewminuteaoatteredpuiictuTee,
the eyes veiy large, somewhat narrowly separated ; antennae long,
slender, joint 11 very elongate, equalling 6-10 united. Protboisz
narrow, considerably longer than broad, the sides obliquely conveig-
ing from the middle forward and sinuately compreesed before the
base, the basal margin raised ; sparsely, minutely punctate, the disc
with a posteriorly widened, interrupted, rather broad median chann^,
and an oblique curved depression on each side of the disc, the trans-
verse basal sulcus not extending across the middle. Elytra very
eloi^te, subparellel, rather more than twice the width of the pro-
thorax, fiatt«ned on the disc, and witii an oblong, deep, intra -huTneral
depicesion ; closely, finely, sinuately striate. punctate, the intersticet
flat, 1, 3, 6, and 7 with numerous small sBtigerous impressions scat-
tered throughout their lengtli, 9 also vrith seveial rather large im-
pressions along the apical half, those towards the apex on 7 and 9
each preceded by a tubercle or short carina. Legs very long, slender.
Leogtli 121, breadth Sf mm.
Hab. Ecuador (Buckley).
One male. An Agraeform insect allied to the Colombian
S. steinheili, Makl., with hrilhant golden-green elytra, the
alternate interstices of which have numerous small setiger-
ous impressions scattered throughout their length, the
seventh and ninth cat«nulate towards the tip.
l„y,l,Ajh,COOl^lL'
Nob and H^e-knovm Lagmdae, 197
31. Stalin latieoIUs.
9. Statira hlicoUis, MSkl., Act. Soc. Fenn. x, p. 637 (1875).
Eloogftte, rather faroiid, widened postftriorly, mod^atel^ shmiiig ;
rufo -testaceous, costaneous, or piceous, the dytra with a more or
less distinct aeneous lustre, the setigerouH impressions towards the
apex indicated by darker subquadrate epota in the light-«olonred or
imtnatore individuala, the elytra with a few erect bristly hails towards
the apex. Head closely, finely punctate, the eyea large, somewhat
distant; antennae long, slender, joint 11 in (} as long bb the four or
fire preceding joints united, in $ about equalling S~10. ProtJiorax
as wide as the head, not or very little longer than broad, moderately
ronnded at tbe sides, oonstrioted before the base, the basal roai^
very prominent; densely, finely punctate. Elytra long, widening
to beyond the middle in both sexes, and there more than twice as
broad as the prothorax, the sides arcuately converging thence Ui the
apex; closely, finely puncta to -striate, the interstices broad, almost
flat on the disc, 3 and 9 with four or five, and 6 and 7 with two or
three, deep setigerous impressions on the apical half or third, the
spaces between them more or less rajsed longitudinally or tubercu-
late. Anterior femora moderately clavate, bare.
Length lOi-12, breadth SMi'k mm. (5?.)
Hab. BEA!aL, Santa Rita (Dr. Sahlberg, Aug. 185tt : type),
Petropolis, Conatancia (J, Gray and H. Clark, Jan. and Feb.
1^7), Rio de Janeiro, Espirito Santo (Fry).
This insect is apparently not uncommon in the neigh-
bourhood of Rio de Janeiro, to judge from the numeioua
specimens before me. It is extremely probable that
S. laticollis is a dark ? of the previously described S. rufa,
Makl., and that the latter is an immature <J of the same
species; but as the author makes no comparison between
them, the identificatdoQ is uncertain. The apically tuber-
culate, subcatenulate alternate elytral interstices 3, 5, 7,
and 9, and the finely impressed striae, are characteristic
of the present species. In paUid examples the setigerous
impressions are indicated by dark spots, much as in typical
S. niffroaparsa, Makl. A specimen (9) before me from Rio
de Janeiro, rufous in colour, with testaceous elytra, and a
fovea on each aide of the disc of the prothorax behind the
middle (not mentioned by Makhn), may be referable to
5. rufa, the type of which must be a ij, with a verj- long
eleventh antennal joint.
iM:,i,A J 1^, Google
198 Mr. a. C. Cliampioii on
32. Ststlra vlridltinoU, n. sp.
Elongate, widened poeteriorly, moderately shining ; pioeotu or
nigro-pioeoQH, the femora paler at the baoe, the elytra. with a braaaj
or greenieh Iiutie, Uie antennae (except two or more of the basal
joints) fenuginouB, the elytra with a few brietly haira towards the
apex. Heed densely, minutely punctate, smoother in front, slightjy
hollowed between the eyes, the latter very large, somewhat distant;
antennae long, slender, joint 11 as long as five or six of the preceding
joints united. Prothorax as wide as, or a little wider than, the head,
as broad as long, moderately rounded at the sides, strongly con-
strioted before the raised basal margin; densely, very finely punc-
tate. Elytra long, widening to beyond the middle, anniately nar-
rowed posteriorly; closely, finely punctato-striate, the striae deeply
impressed towards the tip, the Jnterstioee more or lees convex, 3 with
five or six scattered setigerous impreesions down the apical half,
and S, 7, and 9 with from two to four (tboee on 7 sometimes want-
ing) similar impreesions towards the apex, the imprMiions each
pieoeded by a small tubercle.
Length 9-11, breadth 3-4 mm. (<;.)
Hob. Amazons, Santareid, Ega (ff. W. Bates).
Six examplea, poxaibly all males. This is an Amazonian
form of S. lalicoUis, Makl., that requires a distinctive name.
The more deeply impressed striae (especially at the apex)
and the convex interstices of the elytra, the relatively
longer apical joint of the antennae, the blackened basal
joints of the latter, and the darker body, are sufficient to
distinguish S. viriditincta. A still more nearly allied fonn,
S. peruana, occmB at Chanchamayo, Peru.
33. Ststlra ptnuna, n. sp.
Elongate, widened posteriorly, modemtely shining ; nigro-piceous,
the apical joint of the antennae obscure fermginons, the elytra with
a greoi or bluish-groen lustre; the elytra with a few long bristly
haira towards the apex. Head closely, minutely punctate, the eyes
very large, somewhat distant; antennae slender, moderately long,
joint 11 about equalling the four preceding joints united. Pro-
thorax a little wider than the head, scarcely so long as broad, rounded
at the sides, strongly oonstricted before the raised baeal margin ;
closely, minutely punctate. Elytra long, widening to beyond the
middle, and there considerably more than twice the brewltb of the
prothorax, srcuatdy narrowed posteriorly; finely, closely, deeply
punotato-slriate, the interstices convex, 3 with five, and fi, 7, and
ih,Cooi^lc
New and little-hiown Lagriidae. 199
S Moh with three or four, Mattered deep setigwoiu itnpnssiiHtB
tovnrdn the bpex, the apaoea between th«n ItmgitndiiuUly swdUco
and aubcatcnnlate.
Length llf-lS, breadth 4 mm. ($T)
Hab. Perd, Chanchamayo (ex coU. F. Bales).
Two examples, sex not ascertained. This is yet another
form of iS. lattcoUis, approaching S. caeUUa, Kr., and S. nigra,
Hakl., these latter having the ninth elyttal interstice intei-
rnptecUy catenulate for nearly its entire length . The creoate
elytral striae in S. pervana are still more deeply impressed
at the apex than in the Amazonian S.viridUincta, and the
elevations between the setigeroua impressions are strongly
convex longitudinally, instead of shortly tuberculate as in
the last-named insect.
34. Statira raUrons.
9. Slatira ntfifrons, Makl., Act. Soc. Fenn. vii, p. 152 (1862).
Elcnigat«, iBther narrow, shining; obscure tcetaoeoiu above,
darker beneath, the pnithoiaz aeneoua, the reet of the upper
auifaoe (tba front of the head excepted) with a faint aeaeoua luatre ;
the dytn with a. few long, aoatteced, biiatly haira. Head broad,
■pareely, minutely ponctate, longitudinally Boloate and foveat« in
the middle between the eyes, the latter large, moderetely distant;
antennae long, alender, joint 1 1 in (} about equalling 7-10, in $ S-10,
miited. Prothorax aa wide as, or a little wider than, the head,
IcHiger tiian broad, oblongo4ordat«, oonatrioted before the promi-
nent baaal margin; oloaely, finely, oouapiouonsly punctate, the
interapBcee poliahed, the diao aometimea with a faint traoe of a
median channel. Elytra long, twice aa broad aa the prothorax,
gndnally widaied to the middle, somewhat acuminate post«riorly;
doaely, finely punctato-striate, the striae deeply impressed at the
apex, the interstices flat on the dieo, 3 and 5 each with five or six
deep setigeroua impresBions scattered between the baae and apes,
and 9 with three or four (and 7 sometiniea with one) similar impree-
sions near the tip, the interspaces between them raised and sub-
catflnnlate towards the apex. Anterior femora moderately clavate,
bftie.
Length 8^-10, breadth 2}^^ mm.
Hab. Brazil {Mus. Oxon.), Rio de Janeiro {Fry).
Six examples agreeing very nearly with Maklin's descrip-
tion. Smaller and narrower than 8. JcUicollis, Makl., as
here identified ; the head and prothorax more polished,
iM:,i,A J 1^, Google
200 Mr. G. C. CIiampioD on
the former longitudmsUy siilcat« and foveate between the
eyes, the prothoraz oblong, less densely punctate, and
aeneous in colour; the elytra lens dilated, translucent
aeneo-testaceous, with the atriae deeply impreH»ed at the
apex, the aetigeroua impressions on the interstices 3 and
5 extended forward to the base.
35. Statirs sphenodera, n. sp. (Plate XII, £g. 16, 3.)
Very elongate, narrow, shining; pioeouB, the prothorax, under
surface, and sometimea the femora at the base, mfouH or nifo-teetscfl-
0U8, the elytra with e, translucent greenish or cupreous lustre, the
latter with a few brietly hairs towards the apex. Head long, narrow,
somewhat closely punctate, Bhallowly, loDgitudinaily, grooved
between the eye«, the latter very large and subappraximate, the
neck rugose ; antennae very lot^ and slender in both bcsbs, joint 1 1
in S equalling about five, in $ rather more tlian three, of the pre-
ceding joints united. Protboras elongate, narrow, suboonical,
sinuate at the sides before the base, the basal and apioal margins
slightly raised; closely, finely, irregularly punctate. Elytra very
elongate, twice as wide oh the prothorax, subparallel, acuminate at
the apex; finely punctato -striate, the interstices almost flat, 1, 3, 5,
7< and 0 each with two or three setigerous impressions towaids the
tip. Legs very elongate, slender, the femora gradually thickened to
near the apes in both sexee.
Lengt 121-131. breadth 3-3| mm. (SS-)
Hab. Brazil {Mu^. Oxon.], Rio de Janeiro {Fry), Per-
nambuco (ea: coU. F. Bates).
Seven specimens. A very elongate, narrow insect, not
unlike S. longicollis, Makl., and easily recognisable by ita
long, suboonical, red prothoraz, very elongate, slender
antennae, with an extremely elongate terminal joint in i^,
the metallic elytra, with the interstices smooth to near
the apex, and the simple legs in both sexes. The general
facies is that of an Agra.
36. StaHra slenoeaphala, n. sp.
^. Very elongate, narrow, shining; nigro-piceous, the femora
paler at the base, the apical margin of ventral segments 3 and 4
testaceous, the elytra with a oyaneons lustre, the latter with a few
bristly hairs at the tip. Head oblorkg, narrow, sparsely puncUt«,
foveate in tbe middle between the eyes, and with two small fove«e
on each aide adjacent to them, the eyes latber small (as seen from
Ij.y.l.Ajl^yCOO'^lL'
New and UtHe-known Lagriidae. 201
Above), Bomewhftt distant; antemuie long, quite almder, joints 7-10
T^idlr deoiwing in length, II rather longer than 6-10 unitad.
I^thonx a little wider than the head, maoh loi^;er than broad,
Bubcylindrical, slightly narrowed and constrioted before the apex,
and very feebly sinuate at the sides before the base, the basal margin
a little raised, the hind angles not prominent; very sparsely, finely,
irregularly punctate, the disc trsjisversely depressed on each side
before and beJiind the middle, and indistinctly so in tbe centre at
the base. Elytra elongate, scarcely twice the width of the pro-
thorax, subparallel to far beyond tbe middle, rounded at the tip;
closely, finely slriato-punctate, striate at the apex, the interstices
flat, transrereely wrinkled, without definite coarser setigerous im-
pressitms on the alternate inteisticee. Legs long, slender, the
anterior femora feebly clavate.
Length 10, breadth 2} mm.
Bob. Brazil [Mus. Brit.).
One male, acquired by the Museum ia 1871. An i3olat«d
form, tecogniaable by its narrow head, slender antennae,
long, cylindrical, polished prothoraz, and nigro-cyaneous,
long, subparallel, non-foveolat« elytra. It beats some
resemblance to Hypoatatira variicohr, Fairm., from Minas
Geraes,^^ but the present insect cannot be referred to that
genus as defined by its describer.
37. Stollra viridlnltolu, n. sp.
?. Very elongate, somewhat convex, polished; head and pro-
thoraxnigro-piceoUB, the elytra brilliant metallic green, with cupre-
ous or gold^i reflections at the sides and apex, the legs, antennae,
and under surface piceous, the elytra with a few long, bristly haiis.
Head almost smooth (the covered rugose neck excepted), foveste
in tie middle between the eyes, the latter moderately large, separ-
ated by the width of one of them as seen from above; antennae
extrentely slender and elongate, the joints slightly thickened at the
tip, 11 equalting 9 and 10 united. Prothorax about as wide as the
head, not longer than broad, cordate, almost smooth, the basal
margin laterally projecting, raised. Elytra very elongate, at the
base twice as broad as the prothorax, gradually widening to beyond
tbe middle, acuminate posteriorly, the apices pointed; closely,
finely puncta to -striate, the interstices broad, almost flat, 3, 5, and
7 with several widely separated setigerous impressions scattered
iM-,i,A J h, Google
202 Mr. G. C. Champion on
between the (mm and ftpex, 9 also with three impresBiotu towaids the
tip. Legs very long and Blesder, the femora moderately thifkfn"^
Leogth 12, bieadtb 3| mm.
Bab. CoLOMBU (ex ctM. F. Bates).
One female. An insect with brilliantly metallic elytra,
as in S.sumtuosa, M&kl.,and other allied Colombian and
Ecuador forms, the head and piothoraz relatively short,
the elytra very long, posteriorly acuminate, and mucrooate
at the tip, the antennae and legs as slendet aa in OtAryadet
jTogilicomii, Champ., from Panama. The last-named
species has the mandibles tridentate at the tip, which is
apparently not the case in the present insect, so far as can
be ascertained without opening them. S. anguslala, Yic,
from Ecuador, must have equally slender antennae.
38. StaOra aegnte.
cj. Suaira aegrola, Makl., Act. 8oc. Fenn. vii, p. 149 (1862).
S- Antennae simple, joint 11 abont equalling 7-10 (in $ S-10)
imited; anterior femora stout; intermediate femoia gradoally
thickened to beyond the middle and ali^tly hollowed thence to
the apex.
H(A. Brazil (Mxta. Oxon.), Kio de Janeiro {Fry), Espirito
Santo (Schmidt).
Four specimeus before me are probably referable to this
species. They differ from Maklin's description in having
the small rounded tubercles preceding the setigerous im-
pressions placed on the alternate interstices 1, 3, 5, and 9,
mstead of on 1, 3, 5, and 7, as stated. The tubercles are
distributed along the whole length of the three dorsal
iotetstioes mentioned, varying greatly in number, but on
the ninth they are reduced to a short series near the apex.
An elongate reddish insect, with the elytra translucent green
or bluish-green, the prothoraz sometimes infuscate; the
head broad, almost smooth, unimpressed between the eyes,
the latter large and somewhat distant; the antennae
slender; the prothoraz cordate (9) or oblongo-cordste (J),
densely pimctolate, with very prominent basal mai^^n ; the
elytra Imely putictato-striate, the interstices (except just
in front of the setigerous impressions) almost flat; the
femora and tibiae simple in (J. S. n^tnthorax. Pic," from
San Antonio da Barra, Brazil, seems to be an allied fomi
t> Melange* eiot.-entom. iv, p. U (Sept. 1912).
l„y,|,AJI^,COOl^lL'
NeiB and little-knoum Lagriidae. 203
with a subopaque head and prothoraz, and black antennae,
knees, tibiae, and taisi.
39. Stallta sdntiUans, n. sp.
5. ElougBto, tiAiTow, Tery shilling; aeneo-piceous, the head,
uitennM, and lege reddish ; the head and elytra with numeroue very
long, subject bristly hairs, the antennae also setose. Head rather
small, sparsely pnnotnlate.Iongitndinally grooved betweco the eyes,
the Utter large and narrowly separated ; antennae long, slender,
joint 11 extremely dongate, about equalling 3-10 united. Pro-
thorax as wide as the head, much longer than broad, rounded at the
sides, obliquely narrowed from the middle forward, and constricted
before the prominent basal margin ; closely, very minutely punctate.
Elytra long, nearly twice as wide as the prothorax, subparailel in their
basal half, flattened and uneven on tike disc ; very minutely atriato-
punotate, the interstices broad, flat, 1, 3, S, 7, 9 with from six to ten
setigerous impressions scattered between the base and apex, the im-
pressions each preceded by a small tubercle, these beooming slightly
dongated towards the tip. I«gB rather stout, the femora strongly
davate, the tibiae slightly curved at the base.
Length 7, breadth 2-2^ mm.
Hab. Amazons, Santarem, Ega {H. W. Bales).
Two males. A narrow, elongate fonn related to the
Colombian S. mnUuosa, Makl., but mach smaller, with the
a]ncal joint of the (J'-antenna nearly half the length of the
entile organ, and the surface of the elvtra so uneven as to
appear scintillate.
40. Statlra sphenoptora, n. sp.
S- Elongate, narrow, attenuate posteriorly, rather dull, the elytra
and under surface shiniiig; black, the elytra cyaneous on the disc,
aeneous at the sidea, the antennae (joints 1~3 excepted) and legs (the
bLack knees excepted) testaceous, the abdomen piceous ; the elytra
with scattered long, erect, bristly haiis. Head large, broader than
the prothorax, deosely, very finely punctate, t)>e eyee extremely
large, narrowly separated ^ antennae long, slender [joints 10 and 11
mining). Prothorax much longer than broad, oordate, strongly con-
stricted before the raised, laterally-projecting basal margin ; densely
nunulely punctate, traosverBely excavate in the centre at the base.
£lytra moderately long, less than twice the width of the prothorax,
narrowing from the base, unarmed at the tip; very finely, closely,
puiKtat«4triate, the interstices feebly convex, 1, 3, and 6 with a
n,g,i,.rjh,G00glc
204 Mr. G. C. Champion on
MvieB of eight or nine setigerouB impTcaeioDt, ench pieoeded hy a
anudl tubercle, scattered between the ba«e and apex, and 9 with
four ■mall tubercles along the apic«l half. Legs long, slender, the
anterior femora stonter than t^e others.
Length S, breadth 2 mm.
ffoi. Upper Amazons, Ega (ff. W. Bates).
One male. A narrow, graceful form, with a black head
and prothorax, very large, subapproximate evea, sub-
cuneiform, tuberculate, blue elytra, and long, flavescent
legs, the black knees excepted.
41. Statin sumtnoia.
Statira tumiuota, Makl., Ofv. Finska Vet.-Soc. Forh. xx,
p. 350 (1878).
2- Elongate, widened posteriorly, shining j nigro-piceons, the
elytra brilliant greeniah-aeneous, the elytral depieesions and apex
golden-cupreous in certain lighte, the antennae and 1^ rufo-piceoos ;
the elytra with a few long, bristly hairs. Head sparsely, very findy
punctate, longitudinally depressed in the middle between the eye«,
the latter very large, somewhat narrowly separated; antnmae
sl«ider, joint 11 equalling 6-10 united. Prothoraa a little wider
than the head, about as long as broad, moderately rounded at the
sides, constricted before the raised baeal margin ; closely, oon-
spionously punctate, shallowly, interruptedly sulcate down the
middle, and with an oblique depression on each side of the diao
posteriorly. Elytra long, widening to beyond the middle, twice oa
broad as the prothonts, flattened on the disc below the base, and
longitudinally depieesed within the humeri; extremely finely,
shallowly atriato-punctate, the punctures becoming obsolete towards
the apex ; the interstices flat, 1 , 3, 5, 7, and 9 each with a scattered
series of small setigerous impressions, most of which are placed in
large, broad, shallow, coalescent foveae, the interspacea between
those on T and 9 carinate. I^egs long and slender.
Length 12^, breadth 3J mm.
Hab. CoLOtiBiA (ex <M. F. Bates).
One specimen, agreeing well with Maklin's description,
80 far as it goes. Recognisable by the bhltiantly metallic
uneven elytra, with interruptedly carinate seventh and
ninth interstices, the cloaely punctured, interruptedly sul-
cate prothorax, and slender limbs. The femora are probably
abraded.
l„y,l,Ajh,COOl^lC
New and Uule-lmown Lagriidoe. 205
42. Stotirft ehaloopten, n. sp.
Elongate, widened posteriorly, very shining i lugro-piceoue or
black, the elytra brilliant brassy-green, partly cupreous when viewed
in certain lighta, the lags rufo-teetaceouB, the antennae ferruginous
orfnsco-feiTUgiiiouBj the elytra with a few long bristlj hairs. Head
nther narrow, not wider than the prothorax, sparsely, minutely
punctate, obsoletely impressed in the middle between the eyes, the
latter very large, somewhat narrowly separated; antennae long,
slender, joint 1 1 in $ equalling 8-10 united [broken of! in ,}]. Pro-
thorax slightly longer than broad, rounded at the sides, moderately
constricted before the prominent basal margin; sparsely, minutely
punctate (almost smooth in one example], with a deep, oblong-
triangular excavation in the middle at the base and an oblique im-
praesion on each side of it anteriorly. Elytra long, widening to
beyond the middle and there more than twice the width of the
prothorax, tranaversdy exoavate below the base, and also longitudi-
nally deprcBsed within the humeri; extremely finely, shallowly
■triato -punctate, the inlentioee flat, 1, 3, S, 7 and 9 each with a
Nattered series of small, shallow, setigerous imprwaionB, the im-
pressions on 7 and 9, and those towards the apex of I, plaoed in large
shallow foveae. Legs long; anterior and intermediate femora
sparsely oiliate baneatii.
Length lli-12|, breadth 3t mm. ((?$.)
Hab. Colombia {Mm. Brit.).
Two examples, one with the head and prothorax almost
smooth. This is a form of S. sumtuosa, Makl., vith the
elytra deeply excavate below the base, the broad, shallow,
foveifonn depressiona on the disc restricted to the sides
and apex, and the short carinae between them on the inter-
stices 7 and 9 wanting ; the prothorax deeply triangularly
excavate in the middle at the base ; and the legs wholly
red.
43. SUtira eaveniosa, n. sp.
$. Elongate, depieased, widened posteriorly, shining; nigio-
inceouB, the elytra metallic green, the depreasioos, sides, and apex
aaneo-cupreouB, the legs fusco-oastaneous, the mandiblee nifeacent;
the elytra with scattered, erect, bristly haiia. Head a little broader
than the prothorax, veiy minutely punctate, with a shallow fovea
in the middle between the ejee, the latter moderately large, distant ;
aDtmmae slender, moderately long, joint II equalling ft and 10
nikited. Prothorax slightly longer than broad, narrow, oblongo-
eordate, constricted before the prominent basal margin; closely.
iM-,i,A J h, Google
206 Mt. G. C. Champion on
exoeaaiyely minuted; punctate. Elytn long, oomprMHed below tbe
hnmeri, widaiing to bcTCHid tiie middle and then three timw tiie
breadth of the piDtborax, transversely depressed below the beae.
Mid with a deep, oblique poat-baasil excsvation ; minutelj, sinuoual;,
strialo-punctat«, the interatiocB dat, 3, S, 7, and 0 each witii four or
five large, ahallow,8etigeioua impreesioDB on the apical half,l trans-
rereely notched along the sutnte and also with three or four foveae
towards the tip, the larger impresaions each preceded by a small
tuberele or short carina, the carina on 7 extoiding uninterruptedly
forward to the humeral callus. Legs lorkg, slender, the femora
Length 9^, breadth 3 mm.
Hab. Colombia (Mus. Brit.).
One specimen, acquired in 1847. Xear S. imprestipenms,
Makl., with a narrower, oblong, smoother prothoraz, smaller
eyes, and very minutely striato-punctate, uneven elytra,
the large shallow setigeious impiessiona on the apical half
diverting the linear arrangement of the striae, the spaces
between the impressions interruptedly carinate or catenu-
late. S. sumtuoea ia an alUed larger insect.
44. SUtira Impresripennls.
Statira impreanpennie, Makl., Ofv. Finska Vet.-Soc. FSrh.
XX, p. 351 (1878).
Elongate, depressed, widened posteriorly, rery shining; nigro-
pioeous, the head and prothoraz with an aeneous lostre, the elytra
metallic green, brassy at the tip, the tarsi and joint 1 1 of the antennae
fraruginous ; the elytra with numerous very long, erect, bristly hairs.
Head narrower than the prothorax, sparsely, finely punctate, deeply
foveate in the middle betweai the eyee. the latter very large, some-
what narrowly separated; antennae slender, rather short, joint 11
equalling 8-10 united. Prothotax not longer than broad, rounded
at the sides, constricted before the prominent bssal margin ; some-
what closely, finely punctate. Elytra very long, twice as broad as
the prothorax, widening to beyond the middle, broadly depttesed
for some distance below the base ; closely, very finely, aballowly,
punetato-striate throughout, the intetstices flat, 1 (except along
median third), 3, 5, and d with numerous Betigerous impressions
scattered between the base and apex, 9 interruptedly coetate pc«-
teriorly. Legs very long, slender ; anterior and intermediate femora
finely eiliate beneath; tibiae pilose within.
Length lOi, breadth 3i nun. ($1)
u,y,l,7rJM,G00glc
Nea and UtHe-known Lagriidae. 207
Hab. GoLOHBU (Mils. Brit.).
One specimen, almost certainly referable to S. impressi-
pennis, Makl., the type of which was found by Steinheil in
the same country. It is related to his S. aumiuoia and
other Colombian fonns.
45. StaUra eapeovirfdls, n. ap.
Elongate, ntber narrow, depresaed, ahining; black with an
aeneotu lostre, th« dytra brilliantly metallia green or golden-greoi
at the base, cnpreons from tiie middle onwaida, the anUnnal joints
4—11 obeoore ferrugiiKins, the tarai rnfo-teetaceous ; the elytra and
abdomen with a few long bristly hain. Head small, not wider than
the prothoraz, very spaisely, finely pnnctate, de^ly foveate in the
middle between the eyes, the latter larg«, separated by about half
the width of one of them; anteonae atender, joint 11 in $ [that of
^ wanting] sli^tly longer than S--10 united. Fiothoraz narrow,
longer than broad, obliquely narrowed anteriorly, and deeply con-
stricted before the prominent basal ma^in ; sparsely, minutely punc-
tate, depressed in the middle at the base. Elytra long, flattened,
more than twice the width of the prothorax, subparallel in their
basal half, the biuneri prominent and with a deep oblique depreasiaa ;
very finely striata -punctate, the interstices broad, flat, feebly convex
at the apex, 3 with five or six widely aoatterod, 5 and 7 each with
two on the apical half, and 9 with one at the middle, setigerous im-
piessions, each of which is placed in a large shallow fovea, 9 also
widi lOTeral other impressions towards the tip. Le^ long, slender,
anterior femora feebly clavate.
Uaglh 9H01> bieadth 3-3} mm. (3%)
Bab. Peru, Chanchamayo {ex coU. F. Boies).
One pur, the tip of the antennae wanting in one of them.
AbiiUiantlv metaUic insect allied to the Colombian S. sum-
tuota. Maid., with a narrower head and prothorax, differ-
ently sctdptuied elytra, and ruio-teataceoua tarsi. The
large shallow foveae on the apical half of the el3i^ra give
an uneven appearance to the surface; the general shape
is hke that of an Agra, fam. Carabidae.
46. Stattra viglntlpunetata, n. sp.
Elongate, widened posteriorly, shining; black, the elytra metallic
green, cupreous at the sides and apex, the antennae (exoept at the
base) obscDie femiginons, the legs nigro-piceoue ; the elytra with
a few Tery long bristly hairs. Head small, cloaely, finely punctate,
tlie eyes moderately large, distant; antennae slender, joint 11
iM:,i,A J 1^, Google
208 Mr. G. C. Champion on
Bqnalliiig S-IO united. Prothorax about ae nide aa tlie head, con-
iidfliably longer than broad, oblongoHx>rdBte, ooiutricted heton
the moderately raised basal margin ; closely, finely punctate. Elytra
long, widening to bejrond the middle and t^ere more than twice the
width of the ptothomx, flattened below the base; oloeely, finely
punctatO'Striate throughout, the intersticce almost flat, convex at
the tip, 3 with six, and 6 with four, rather large, deep, setigerons
impresBions scattered between the base and apex, 9 also with two
smaller impressions towards the tip. Legs long; anterior femora
moderately ctavate. '
Length 9}, breadth 3 mm. ($ T)
Hab. Peru, Chanchamayo (ex coil. F. Bates).
One specimen. Differs from S. cupreoviridia, from the
same locality, in having the elytra shorter, less brilliantly
coloured, punctato-striate to the apex, and with the
setigerouB impressions smaller, deeper, and otherwise
arranged, the head and prothoiax closely punctate, the
tarsi infuscate. The sharply-defined elytral foveae are
suggestive of those of various species of the Carabid-genus
Pterostickus.
47. StKflra semienprsa, d. sp.
Elongate, widaied posteriorly, somewhat robust, shining ; piceous,
the elytra aeneoos at the base, the oolour ohaitging to cupreous
from about the basal thiid onwards, the cupreous coloration enclosing
an oblong green patch on the disc near the tip ; the elytia with a tew
long bristly hairs. Head short, barely as wide as the prothorax,
closely, finely punctate, the eyes large, distant; aoterinae moder-
ately long, slender [tip broken off]. Prothorax transverse, rounded
at the sides, deeply constricted before the prominent basal margin;
closely, finely punctate. Elytra long, broad, more than twice tha
breadth of the prothorax, gradually widened to the middle, and
somewhat acuminate posteriorly; very finely, closely striato-
punctate, the int«rstices broad, flat, 3 with four or five, and 6 with
three, large setigerous impreasions towards the apex, 9 also with
a aeries of impressions down the apical half, the interspaces between
them cariniform. Anterior femora moderately olavate.
Length 11^, bit^th 3} mm. ($T)
Hab. Pebu, Chanchamayo (ex cdl. F. Bates).
One specimen. Larger than the Central American
S. glabrata, Makl., with more elongate, smoother, partly
cupreous, apically foveolate elytra, more slender antennae,
a less constricted prothorax, simple anterior femora, etc.
n,g,t,7rJM,GOOglC
New and liaIe-]cnoti>n Lagritdae. 209
The elytral sculpture is rather like that of S. cupreoviridu,
the lai^ eetigetons impres^oiiB, however, in the present
species are mostly placed near the apex.
48. Statin splendleans.
$. Siatira apletidicatis, Makl., Act. Soc. Fenn. vii, p. 151
(1862).
Vetylike S. mkant, HaJcl. (No. 29), the altenwtd elytral inWrsticee
not catenulate towards the apcE, I, 3. and 5 with a scattered series
of aetigeroiis impreaaioiiB, those on 7 wanting, S witii two or thiee
impieeaions near the tip; head a little broader, the inter-ocolar
grooveonly just indicated, the eyes men distant; antenna! joint II
in J oqualling 7-10, in $ 8-10, onited ; legs slender, simple in i^ ;
aedeagua slender, thickened and slightly hooked at the tip above.
ffoi, Brazil {Mus. Oxon.), Constancia {J. Gray and H.
Clark, Jan. 1857), Rio de Janeiro {Fry), Minas Geraes
(Mus. Oxon.).
Two males and four females seen.
49. Stittrs TOtundieollls, n. sp.
Uoderatelj elongate, somewhat robust, shining; piceoos, the
bead and prothoraz with on aeneous lustre, the elytra brassy, the
antennae obscure ferruginous, the elytra with a few long, bristly
hairs. Head about as wide as the prothorax, almost smootli, a
tianaverse foveate depression between the eyes excepted, the latter
iatge, distant; antennae slender, joint 11 nearly as long as 7-10
onited. Prothoraz convex, about as broad as long. atroDgty
rounded at the aides, deeply constricted before the prominent basal
margin; sparsely, very minutely punctate. Elytra long, twice
as broad as the prothorax, gradually widened to the middle, and
somewhat acuminate posteriorly; closely, finely, deeply punctato-
striate, the interstices becoming more convex towards the apex,
3 with five, and S witb three or four, widely scattered, setigerous
impressions, 7 also with one at the shoulder and 9 with two impres'
■ions near the tip. Anterior femora clavate; anterior tibiae feebly
Length 8^, breadth 2| mm. {^ T)
Hab. Colombia, Bogota {Mus. Brit.).
One spedmen. Narrower than the Central American
8. ^abrata, Makl., the prothorax not wider than the head,
the anterior femora unarmed, the elytra more deeply
TRANS. ENT. SOC, LOND. 1917. — PART I. (NOV.) P
iM-,i,A J h, Cookie
210 Mr. G. C. Champion on
striate, with the interstices convex at the apex. More
shining than S. jmncticeps (No. 53), the head and pro-
thorax almost smooth, the tatter deeply constricted before
the base.
60. Statin dlviu, n. sp.
Elongate, d«pieeeed, widened poetesnotij, very Hhining ; ofaecuie
teetMMOOB, the hwd, antennae, and palpi in great part pioeons; the
djtta of a brilliMit braaay tint, ting^Bd with cupteons lUong the
inferior maigin, the autnn tMtaoeoiu throughout, the teataoeoui
ooloration gradoallj becoming more extended towaids the apex;
tiie elytra with a few long bristly hairs. Head about aa wide u
the piothorax, eporedy, minutely pnnctete, ahallowly 'loveate in
the middle between the eyee, the latter moderately large, distant:
antennae long, alaider, joint 11 barely the length of 8-10 united.
Prothoraz about aa broad as long, rounded at the aides, constricted
before the raised basal margin ; very sparsely, obeoletely punctulate.
Elytra long, widening t« beyond the middle, and there more thu
twice the width of the pro thorax, subacuminate post«riorly, depieseed
below the bsse, and with oa oblique intra-humeial depression:
closely, minutflty punctato-atriate, the sutural stria deeply im-
pressed at the apex, the interstices flat, 3 with about eight, S wi th fonr
or five, and 9 wi^ numerous, deep setigerous impressions scattered
throughout their length, and 1 with two small impressions near
the tip. Legs long, slender, the femora glabrous.
Length 8^, breadth 3 mm. ($ T)
Hob. Venkzuela (Dyson, in Mm. Bnt.).
One specimen, somewhat immature and discoloured,
but otherwise perfect, acquired in 1847. Smaller than
the Colombian S. impressipennis, Makl., the piothorax
narrower, the elytra testaceous along the suture, with fewer
setigerous impressions on the disc, the femora gtabioiu
( ? abraded), etc. The numerous deep impressions scattered
along the whole length of the ninth elytral interatice is a
marked feature of the present species. The much larger
;S. nubeculosa, Makl. (length 13 mm.), from Colombia, also
has a Butural stripe.
51. Statlra dUnta. n. sp.
Elongate, shining; piceous or mfo-castaneous, the antennas
and legs paler, the femora slightly infusoate, the elytra with s
cupreous or greenish lustre, the latter with a few bristly bain
towaids the apex. Head as brood as the prothorax, sparsely, wj
l„y,l,Ajh,COOl^lc
Hew and little-knoim Lagriidae. 211
minntelr puncUt^ with eevtnJ ooaner panctona between the
ofM, tbe bttot modentely large, diatant; antaDnae almder, long,
joint 11 nesitj or qait« eqnaJling 8-10 nnitcd. Frothor&x aboat as
Intg aa broad, romtded at the sidea, deeplj ocmstrioted before tbe
raised basal nuugin; apandy, very minnteij ponotate. Eljrtn
long, flattened on the disc, twice aa broad aa tbe protborax, grada&Dy
widened to the middle, and somewhat aonminate poaterioriy;
doeely, finely, shallowly punctAto-Btriate to the tip, the inlerstices
broad, fiat, 3, 6, 7, and 9 each with from two to fooi eetigeioDB
impraesiona on tbe apical half (those on 7 wanting in one specimen).
Length 9i-10i, breadth 3-31 !»)>>•
Hab. EcoABOB, Macas (SuoU^).
Three Epecimens, possibly including the two sexes.
Near the Venezuelan S. divisa, the elytra uniformly cotouied,
simply flattened on the disc, the setigerous impressions on
the inteistices 3, 5, and 9 fewer in number and mostly
placed near the apex. The much smoother, polished
prothorax separat«s S. dUula from S. UUtrdlis (No. 71),
timonis, aufffisa (No. 72), caeruUotincta (No. 59), etc.;
and ita larger size and more elongate elytra from S.
tran^ludda (No. 56).
52. Statfrs slmonls.
Statira nmonis, Fairm., Ann. Soc. Ent. Pr. 1892, p. 95.
Elongate, somewhat depressed, widened posteriorly, shining;
nifo-testaoeouB, the eyes black, the elytra with a faint brassy lustre,
(he tatter with a few long bristly hairs. Head not so broad as the
proUiorax lOlosely. piinotulate, transversely impiessed between the
eyes, the btt«r moderately large, distant; antenns« slender, long,
joint 11 not quite equalling S-10 united. Prothorax a little broader
than long, moderately rounded at the sides, deeply constricted
before the raised basal margin; densely, finely punctate, transversely
grooved on each side of the disc just behind the middle, the basal
groove complete. Elytra long, twice the breadth of tbe prothorax,
widening to b^ond tbe middle, the sides arcuately oonvet^ing
thence to the apex; closely, very finely stria to -punotate, the inter-
stioea broad and fiat thronghont, 3 with six (scattered betweoi
the base and apex), 6 with four (on the apical half), and 9 with three
(towards the apex), large, deep setigerous impressions. AaUaioz
femora moderately olavate.
Length 10, breadth 3^ nun. ($t)
n,g,t,7.dh,G00glc
212 Mr. G. C. Champion on
Hah. VEiTBZtmLA {Dyson). San Eeteban, Colonia Tovar
{Simon : type).
One specimen, acquired by the Museum in 1847, agre^g
with the description of S. simonis, Fairm. Neat S. iateralu,
Makl. (as here identified), and separable therefrom by the
broader, laterally-grooved prothoraz, and the very finely,
shallowly striato-punctate elytra, the interaticeB of which
are fiat throughout, 3, 5, and 9 with a few large deep
Betigerous impressions. S. divisa from Venezuela, S.
caendeolinaa, from Colombia (No. 59), and S. testacea and
S. ameotincta, from Central America, are also alUed fonns.
53. Statira functieeps, n. sp.
Moderately elongate, niUTOV, feebly shining ; greenish .aeneoiu,
theontennae, legs, and undersurfaceingreat part or wholly pieeona;
the head and elytra with scattered long, erect, pallid, bristly liain.
Head rather small, somewhat coarsely, closely punctate, smoother
in the middle between the eyea, the latter large and well separated;
antennae slender, long, shorter in $, joint 11 in <} about equalling
7-10, in $ barely as long as 8-10. united. Prothorax obtong, longer
than broad, as wide as the head, constricted before the base, t^e
basal margin moderately raised; closely, conspicuously punctate,
without trace of median channel, the interspaces alutaoeous.
Et]rtrs rather long, about twice as broad as the prothorax, sub-
parallel at the base; closely, finely, rather deeply punctata-striate,
the intersticee becoming oodtcx towanls the apex, 3 with six or
serai, and 5 with four or five, setigerous impressions scattered
between the base and apex, 9 also with two impressions near the tip.
Anterior femora strongly clavate.
Var. a. Antennae (joints 1 and 2 excepted) rufo- testaceous, (o-)
Var. ^. Prothorax and abdomen rufo-piceouB, \egi and antennae
(joints 1 and 2 excepted) rufo -testaceous. ($.)
Var. y. Prothorax, a large apical patch on the elytra extending
narrowly up the suture to the base, antennae (joints I and 2 ex-
cepted), legs, and under surface ferruginous or rufo -testaceous, (o-)
Length 7-fl, breadth 2-3 mm. (,J?.)
H<A. Brazil, Rio de Janeiro (Fry), Conatancia {J. Gray
and H. Clark. Jan. 1857 : type).
Four males and two females, certainly belonging to one
variable species, the darker eitamples ((J$) selected as tyjKB.
More elongate than the equally variable S. incoratmu.
Champ., from Central America, the elytra subparallel in
male, the head and prothorax closely, conspicuously punc-
ihyCOOl^lC
New and liiie-knovm Lagriidae. 213
tate, the eyes larger and more approximate. The longer
elytra, with deeper striae and more numerous setigerous im-
pressions, the more coarsely pnnctured head and prothorax,
the 1att«r without trac« of median groove, easily separate
S- puncticeps from S. sericea (No. 65). The colour of the
antennae and Legs proves to be of no value as a speofic
character in the present species. S. fiavkomis, Makl.,
must be a very (fifferent insect.
54. Statin gritlosa.
Statira gratiosa, Makl., Ofv. Finska Vet.-Soc. Forh, xx,
p. 353 (1878).
S. Elongate, narrow, ghining; rufo-castaneoue, the antauae
(joint 11 excepted) and upper surface darker, the elytra brilliant
violaceo-cupteooB, metallic grerai along the Buture and at the sides,
the green colontion extending across the base. Head smalt, not
wider than the prothorax, coanelj, cloeelj punctate, smoother in
front, the eyes moderately large and separated by about half the
width of one of them; antennae rather slender, joint 11 extremely
dongate, about eqnalling 4-10 united. Prothorax longer than broad,
oblongo-cordate, as wide as the bead, constricted before the rather
prominent basal margin ; ctoaelj, conspicuously punctate. Elytra
moderately l<mg, twice as wide as the prothorax, subparallel at the
base, narrowing from before the middle to the apex, the intra-
humeral depression oblong, deep; minutely striate- punctate, the
pnDctureH becoming still smaller towards the tip, the interstices
qnite flat throughout, 3 with sis large, shallow setigerous impressions
scattered between the base and apex, 9 with two smaller impiesaions
near the tip. Legs sparsely pilose; femora oiliate with long haira
beneath, the anterior pair strongly clavate.
Length TJ. breadth 2^ mm.
Hab. Colombia (Mus. Brit., ex coll. F. Bales],
One male, assumed to be referable to Maklin's S. gratiosa ;
but as the author does not mention the sculpture of the
head and form of the antennae, and describes the body as
black (an unimportant character), the identification is
not altogether certain. S. cupripainis, also found by
Hteinheil in Colombia, is said to have similarly cupreous
and golden-green elytra, with setigerous impressions on the
interstices 3, 5, and 7, an almost smooth prothorax, etc.
The specimen here described is one of two Colombian
insects from the F. Bates collection labelled " S. mridi-
pennis, Dej,," a name used by Serville for a very difieient
iM-,i,A J h, Google
214 Mr. G. C. Champion on
Brazilian fotm. The present species is nearly allied to
the Central-American S. variabilis. Champ., which has
ciliat« ant«rior femora in both sezee.
55. SUtlm dejeani, n. sp.
Slalyra viridipennis, Dej. Cat., 3rd edit., p. 236 (1837)
(nee Serv.).
,J. Elongate, narrow, ahiiJng ; castaneous, the elytra tranBlucent
metallic green, except along the auture posteriorly. Head clceelj,
rather coarsely punctate, the eyes large and somewhat narrowly
separated; antennae with joint 11 neariy equalling 4-10 united.
Prothorax considerably longer than broad, oblongo-cordate, the
basal margin raised; closely, conspicuously punctate. Elytra
moderately long, narrowing from the middle, the oblong intra-
humeral depression shallow ; closely, rather finely punctata -striate,
the Butural stria deeply impressed posteriorly, the inteistices flat,
3 with five widely separated deep setigerous impreBBioiiB scattered
betweoi the base and apex, 9 also with two smaller impressions
towards the tip. Femora sparsely ciliate, the anterior pair olavate.
Length 7}, breadth 2^ mm.
Hob. Colombia, Carthagena (ex coU. F. Bates).
One male, from the Dejean collection, and included
by him under S. viridip&inis. Extremely like S. gratiosa,
but with the elytra translucent metallic green (except along
a common elongate space on the disc posteriorly), more
coarsely punctato-striate, and with the similarly-placed
setigerous impressions deeper and less extended laterally;
the prothorax longer, and less rounded at the sides ; and
the head a little more sparsely punctured; the eleventh
antennal joint slightly shorter. S. terminalis, Makl.,
must be similarly coloured.
56. Statira trausliidda, n. sp.
Moderately elongate, widened posteriorly, shining ; ruf o-teetaceous,
the eyes blaok, the palpi sometimes more or lees inf uacate, the elytra
(the epipleura excepted) suffused with green or golden-green, the
latter with a few long, erect, bristly hairs. Head sparsely, minutely
punctate, obsoletely foveate in the middle between the eyas, the
latter distant, moderately large; antennae rather sIraideT, joint II
in j nearly as long as 8-10 united, in $ a little shorter. Prothorax
almoet or quite as wide as the head, about as broad as long, cordate,
strongly constricted before the prominent hind oitgles, the basal
u„,,i,A J 1^, Google
New and lHUe-lcnovm Lagriidaf. 215
BMrpn feebly isised ; apandy, minDtdy punctata. Elytn modB'-
aMj long, twice u btoad as the prothoiftx, wideoed to the middki,
and mpkUy lufrowed thence to the somewhat oonminate apex,
trMMToraoly dapreeaed below the base; cloaely, finely punctato-
■triate, the intenticn beconting convex towards the apex, 3 witii
' from five to eight, and 6 and 9 with foor or five setigeions impna-
aions {those on 3 and 5 scattered between the base and apex), 1
also with one or two impressions near the tip.
Lengtii «{-71, breadth 21-21 mm. ((}$.)
Hab. Brazil (Mua. Oxon.), Rio de Janeiro [type] (Fry),
PetiopoliB {J. Gray and H. Clark, Jao. 1857), Eapirito
Santo (Degcourtils).
Nine apedmena seen, inclading the three in the Oxfoid
HoBeoin. A rather amall, mfo-testaceous form, with dilute
golden-green or green, very shining elytra, which in both
sexea are widened towards the middle and acuminate
posteriorly, slender antennae, and a relatively narrow
head and prothorax. Maklin's description of S. svbaenea,
from Santa Catbarina, nearly appUes to the present insect ;
but the former is said to have the head shallowly, longi-
tudinally foveate between the eyes, the basal margin of
the prothorax moderately raised, the elytra linear or very
feebly widening in their basal two-thirds, and the five
setigerons impressions on their third interatice restricted
to the apical half. S. cupreolincta. Champ., from Panama,
is a larger and more elongate insect, with a wider pro-
thorax and less uniformly coloured elytra, the latter with
much larger setigerous impressions.
57. Stadia sabglabrata, n. sp.
Moderately dongate, widened posteriorly, shining ; ru{o -testaceous
or testaceous, the eyee black, the elytra aeneo-piceous, the latter
almost gkbrotu. Head spanely, minutely punctate, the eyes
moderately large, distant; anteimae rather stout, comparatively
short, joint 11 not or very little longer than 9 and 10 united. Pro-
thorax as wide SB the head, about as long as broad, cordate, slightly
dilated at the base, the basal margin feebly raised; spaiaely,
minatdy punctate, findy, obsoletely canaliculate on the disc
aaterioTly. Elytra not very elongate, in their widest part twice as
bioad aa the prothorax, gradually dilated to the middle, and arouately
narrowed posteriorly ; closely, finely punctate -striate, the inter-
■ticee feebly convex, 3 with three or four widely scattered setigerous
ll,y,l,/eJh,G00glc
216 , Hi. G. C. Champion on
impreasiona aJong their apical half, and 9 usually with one or tvo
impTOBaions near the tip.
Length 4^-6}, breadth 21-21 mm. (<;$.)
Hob. Amazons, Ega, Santarem [type] {H. W. Boies].
Seven examples, including three males with the aedeagus
partly exposed. This insect is of about the same size and
shape as S. viridicincta (No. 96), differing from it in having
uniformly aeneo-piceous elytra, with the setigeious impres-
sions OQ the disc reduced to three or four on the apical
half of the third interstice, shorter and stouter antennae,
with the terminal joint not much longer than 9 and 10
united in (S, and an obsoletely canaliculate piothorax.
58. Statira caliensls, d. sp.
Moderately elongate, depremed, Bhining ; aeneo-piceous or rufo-
ptoeoua with an aeneous lustre; the elytra with a few long, erect,
bristly hairs. H«ad sparsely, very minutely panctale, transversely
depreesed between the eyes, the hitter lai^ and well separated;
antennae long, rather Bt«ut, joint 11 in (J as long as 7-10, in $ the
length of 8-10, united. Prothorax in (J as hroad as, in $ broader
than, the head, not so long as wide, subcordate, strongly constricted
before the base, the basal margin prominent and laterally projecting;
rather sparsely, minutely punctate. Elytra moderately long, at
the middle twice as broad as the protliorax, gradually widened in
their basal half; closely, finely crenato-striate, the interstices be-
coming convex towards the sides and apex, 3 and 6 with soattered
conspicuous eetigerous impressions along their entire length, 7 also
with one, and 9 with two or three, near the apex. L^s lather
Length S^9, breadth 2^-3 mm. (3^.)
Hab. W. Colombia, Call (Rosenberg).
One pair. Near S. laieralie, Makl. (No. 71), as here
understood but aeneo-piceous in colour, the antennae
stouter, and the el^'tra with a scattered row of setig«Y>u8
impressions along the interstices 3 and 5 from the base to
the apex.
59. SUOm oaeruleotineta, n. sp.
Elongate, somewhat robust, moderately shining; castaneous, the
riytra with a translucent bluish lustre on thedisc, the colour changing
to brassy towards the sidee and apex ; the elytra with a few bristly
hairs. Head nearly as broad as the prothorax, sparsely, minotdy
u„,,i,Aji^,Coot5lc
New and liUle-knovm Lagriidae. 317
punctste, the eyes Ui^, aomewbtit distant; antannae long, Blender,
joint II «a long as 7-10 united. Protboraz rather broad, about u
long M wide, rounded at the eidea, constricted before the prominent
lateratty -projecting baaal margin ; closely, minutely punctate.
Elytra long, broad, slightly widening to the middle, about twice as
wide as the prothorai, somewhat acuminate at the tip; closely,
finely puncta to -striate, the interstices flat, 3 with four or five deep
setigerous impressions scatteied along the apical half, and fi and 9
also with two or three otheis towards the tip, 3 and 5 subcatenulate
posteriorly. Anterior femora clavate, bare.
Length 9i, breadth 3 mm. (St)
Hab. CoLOMBU (Mwt. Brit., ex coU. F. Soies).
One specimeD, from an old French collection. Not onlike
the Brazilian S. rrtfa and S. mfifrons, Makl., the elytra
bluish OD the disc, the setigeroue impressions fewer in
nombei (wanting on the seventh interstice), the head not
sulcate as in ;S. rufifrons, the head and prothorax formed
nnich as in S. laticollis. The Mexican S. caeruleipennis,
Champ., is somewhat similarly coloured.
60. Statin puietlooUls, n. sp.
Moderately elongate, shining; reddish -brown, the femora paler
ftt the base, the eyes block; the elytra with a faint metallic lustre,
and bearing a few bristly hairs. Head broad, sparsely, minutely
punctate, with a shallow fovea between the eyes, an oblong depres-
siott in front of this, and a deep fovea ('/accidental) on the epistoma,
the eyee very large, narrowly separated; antennae moderately
long, tatber stout, joint 11 nearly equalling 7-10 united. Prothorax
•a wide as the head, not longer than broad, subquadnite, strongly
ccoutricted before the base, the basal margin raised ; closely, very
finely punctate, the interspaces polished. Elytra moderately long,
•t the middle twice as wide as the prothorax, somewhat acuminate
posteriorly; closely, finely punctato-striate, the striae deeply im-
piesMd at the apex, the interstices flat on the disc, 3 and 6 with
from six to eight small setigerous impressions scattered throughout
tbeu- Imgtb, 9 also with two othera near the tip.
Length 7}, breadth 2^ mm. (£!)
H<A. Amazons {H. W. Batea).
One specimen, exact locality not given on the label.
Near the variable S. muUiformis, Champ., from Central
America, S. insularis {No. 77), from Trinidad, and S.
(Wruieodnda, from Colombia, difiering from all of them in
iM-,i,A J h, Google
218 Mr. G. C. Ghampion on
the inteiTuptedly aulcate inter-ocular portion of the head
(the deep fovea on the epistoma being pmbablj accidental),
the antennae stoutet and the prothoras more shining and
leas densely punctate than in the first two species, the eyes
more approximate and the prothorax not so convex and
less rounded at the aides than in S. caendeolincta.
61. SUflra ehloropten, n. ap.
Eloag&t«, nth«r oonvox, robust, very ahining ; piceoiu, the dytn
brillimt metdlic green, Uve latter witli a few very long, erect, pallid,
brietl; hairs. Head rather Bmall, narrower than the prothoi
almoHt smooth, the eyes large, distant ; antennae moderately long,
stout, joint 11 equalling 9 and 10 united. Prothorax broad, t
verse, arouately dilated at the sides, narrowed and deeply con-
stricted posteriorly, the baaal ma^in prominRit ; almost amootb,
transversely excavate in the middle in front, and sballowly,
terruptedly grooved down the middle. Elytra long, broad, neariy
twice as wide as the prothorax, subparallel in their basal haK.
with rows of fine, approximate punctura plaoed in deep striae,
the inteiBtioea here and there transversely wrinkled, becoming mora
convex towards the apex, 3. 5, and 9 with several widely sqiarated
inoonspicuous setigerous punctures. Ijegfl rather atoot, anterior
femoia clavate.
Length 10, breadth 3^ mm. ($1)
Hab. Pbbu (ea: coH. Pascoe).
One specimen. This insect has stout antennae and s
short, arcuately dilated prothorax, somewhat as in iS.
eurydera (No. 68), but the general facies is more like that
of the much larger S. ingens, from Central America. The
brilliant metallic green, rather broad elytra, the small head,
and somewhat convex body are also characteristic.
62. StaUra eanalienlata, n. sp.
$. Elongate, shining; nifo-piceous (obscure testaceous in the
immature example), the head and legs paler, the elytra with a dilute
aeneous lustre, the latter with a few long bristly hairs. Heed broad,
almost smooth, the eyu large, distant; antennae short, slender,
joint 11 nearly equalling 8-10 united. Prothorax not quite so broad
as long, a little wider than the bead, transversely cordate, strongly
oonstrioted before the raised basal margin ; very spsiselj, finely,
unequally punctate, interruptedly oanaliculate down the middle,
the groove widened before the base. Elytra very long, twice as
l„y,|,AJI^,COOl^lC
Nea and litUe-lcnown Lagriidae. 219 '
broad aa tiie protboros, gndnaJly widening to the middle, and
Bomenriiat acuminato poateriorly; ctoaelj, finely pnnotAto-afcriate,
the etoiae deeply impteosed at the apex, the intentioea almost fiat
to near tbe tip, 3 with nine or ten, and 5 with five or six, setigenoB
impreeetMw Matt«Ted thronghoat their hngth, 9 alio with two or
three impreMiona towards the apex.
Length ll^^- breadth 3i-4 mm.
Hob. Peru (ex coU. Fry), Chanchamayo (ex coU. F.
Bateg).
Two femaJes, possibly obtuned by the same collector,
Tliamm. Nanower than the Brazilian S. lattco&is, Makl.
(No. 31); the prothorax much smoother, interruptedly
canaliculate on the disc ; the elytra lelatively ]oot;;er, with
a Bcatteied series of setigerous impres^ons along the
interstices 3 and 5, the latter not catennlate at the apex.
The short, convex, conspicuously canaUculate, compara-
tively smooth prothorax also separates the present species
from various other allied species. Compaied with the im-
maculate form of the Colombian S. Irisdlata (No. 103), the
eyes are larger, the antennae shorter and not so slender,
and the elytra less acuminate at the apex, with the seriate
punctures much &ner and the interstices almost flat.
63. Statlra ranteua, n. sp.
S- Elongate,narrow,depnHsed,TeryBhiiiing,theprothoraiduUer;
bead and prothorax aeneooa, the elytra brilliant metallic green,
the antamae, t^s, ajid under surface black; the bead and elytra
with a few long, fine, erect, bristly hairs. Head a little wider than
tbe prothorax, sparsely ptmctulate, foveate in the middle betwe^i
tlie eyea, the latter large and somewhat distant ; antennae slender,
joint 11 equalling S-10 united. PrDthonx longer than brood,
sobo^indrioal, strongly constricted before the prominent baaal
maigin; sparsely, minutely punctate and finely canaliculate, the
iatorspaoes alutAceous. Elytra long, about twice as broad as the
prothorax, snbpaiallel at the base, very little wider at the middle,
the intia-bnmetal depression oblong, deep; very finely, olosdy
punotato-striate, the interstices broad, flat on the disc, convex at
(be apex, 3 with four scattered setigerous impresaions on the apical
half, and 9 with two similar impreesions towards the tip. Lega
long, slender, the anterior pair with strongly clavate femora, curved
tibiae and broadly dilated tarsi, the tarsal joints 2 and 3 ti
Lcngti) S, breodtii 2} mm.
iM:,i,A J 1^, Google
220 Mr. G. C. Champion on
Hab. Brazil, Caraca (P. Germain],
One example. An insect veiy like S. versicolor, with
wholly black legs and antennae, a lees deeplv canalicnlate
prothorax, a relatively shorter eleventh antennal joint in
(J, and briUiant metallic green elytra. Compared with
S. sericea, the elytra are more elongate, the head has a
single fovea between the eyea, and the legs and antennae
are black. The dilated anterior tarsi are obviously indica-
tive of the male sex. S. degans, Makl. (No. 87), is a
somewhat similar differently-coloured form.
64. Statirs vertfeolor, n. sp.
(J. Elongate, narrow, depreeeed, Bhining; piceo-castaneoae, the
head and prothorax with a faint aeneous lustte; the elytra in the
esample selected as type met«llic blue-green in the sutural region,
the colour changing to golden exterior to this, and to cupreous along
the sides and at the apei, in a second specimen aeneo-cupieous ;
the head and ejj^ra with a few long, erect, bristly hairs. He«d
large, a little wider than the prothonix, sparsely, minntely punctate,
flattened, punctulate. and laterally impressed between the eyes,
the latter large and rather distant ; antennae slender, long, joint 1 1
a little longer than S-10 united. Prothorax longer than broad,
oblongo-cordate, constricted before the prominent baaal tnaigin;
sparsely, very minutely punctate, the interspaces alutooeous, tiie
disc strongly, interruptedly canaliculate. Elytra long, twice aa
wide as the prothorax, subparslle] in their basal half, the intra-
hnmeral depression oblong, deep; closely, finely pusola to -striate,
the interstices flat on the disc, convex at the apex, 3 with about
four widely separated setigerous impressions on the apical half,
1 and 9 also with a single impieesion near the tip. Lc^ long,
slender, the anterior pair with the femora moderately clavate
and the tibiae feebly curved.
Length 7t-9, breadth 2i-2i mm.
Hab. Brazil {Mm. Oxon.), Bio de Janeiro (Fry : type).
Two males, one of them immature, with the elytra
almost uniformly aeneo-cupreous and the legs testaceous.
Very near S. sericea, the elytra partly cupreous, the pro-
thorax more deeply canaliculate, the head a little broader,
the terminal joint of the antenna longer. The stron^y
canahculate prothorax, the more metallic elytra, with fewer
setigerous impressions (those on the third interstice not
extending beyond the middle anteriorly), and the absence
Ij.y.l.Ajl^yCOO'^lL'
New and Uule-knoum Lagrii^ae. 221
of the median sulcns on the head, separate S. i-ersicolor
from S. nififrons, Makl. (No. 34).
65. SUtiia serieea, n. sp.
Moderately elongate, narrow, shining, the prothoraT dull ; a«meo-
piceoua, or nifo-castaneoua with a faint aeneous lustre, the elytra
aeneous or golden-green, becoming oupieouB or golden towards the
Bidee and apex, the ant^inae t«etaoeous, the two basal joints, the
eyee, and palpi black or blackish ; the elytra with a few long, erect,
bristly hairs. Head rather broad, very sparsely, minutely punctate,
shallowly bifoveate between the eyes in <J, the latl«r large, some-
what distant; antennae slender, rather long, joint 11 in J aa long as
7-10, in $ equalling S-10, united. Prothorai narrower than the
head, longer than broad, oblongo -cordate, strongly constricted
before the raised, laterally prominent, basal margin; sparsely,
minutely punctate, the interspacea atutaceous, and finely, distinctly
canaliculate down the anterior half. Elytra moderately long, twice
aa broad as the prothorax. gradually widened to the middle, and
obliquely excavate or depreesed within the humeri ; cloe^y, very
finely punctato-striate, the interstices flat on the disc, convex at
the Apex, 3 with four or five widely scattered setigerous impieesions,
0 also with one or two others towards the apex. L^b slender.
Length 6-6J, breadth l,V-2 mm. {(J?.)
Hab. Brazil [Mus. Ozon. : <^), Constancia (J. Gray and
H. Clark, Jan. 1857 : type, cJ), Rio de Janeiro {Fry : $).
Three males and one female. Narrower and less robust
than the Central American S. variabilis. Champ. ; the
antennae more slender, testaceous (the two basal joints
excepted), with a less elongate apical joint in (J; the
head and prothorax smoother and not so shining, the latter
finely canahculate; the elytra cupreous or golden at the
sides, with the seriate punctures finer and the interstices
convex at the apex. S. subaenea, Makl., from Santa
Catharina, and S. gratiosa, Makt., from Colombia, are
similarly coloured forms. The upper surface in the
present insect is not very highly polished, appearing
somewhat sericeous.
66. Statira purpuielpennls.
SUUira purpureipennis. Pic, Melanges exot.-entom. xi.
p. 19 (Nov. 1914).
tfafc. Bbakil, Jatahy, Prov. of Goyas {Pujol).
u,y,l,7rJM,G00glc
222 Mr. 6. C. Champion on
In the Fry collection there is an example ($?) of this
species from the same locality as the type. It is a very
elongate, narrow insect, nigro-piceous in colour, with the
elytra moderately shining, purple ; the abdomen castuteous ;
the head and prothorax opaque, and densely scabroao-
punctulate, the fonner large, the latter longer than broad,
cordat«, with rained, laterally-projecting basal mai^in;
the elytra elongate, subparallel in their basal half, the
alternate interstices 1, 3, and 5 with a complete series,
and 7 with a row down the apical half, of rather cloeely
placed, setigeroufl impressions, each of which is preceded
by a small tubercle, 9 also with three impressions near the
tip ; the antennae long and slender [apical joints wanting] ;
the legs unusually elongate, the antenor femora not much
thicker than the others.
67. StaOrs nmbrosa, n. sp.
Elongate, nktiier luknow, opaque; piceous or oaHta&eoiu, the
femora p&ler at th« base, the elytra black, the latter with numeroiu
VBiy long, ereot, bristly hairs. Head lai^ dauely, Boabrooo.
punotulate, the eyes very lai^ modarately diataut; antatmae
Blender, long, joint II in <} about equalling 6-10, in $ S-IO, onitcd.
Prothonx aa wide as the head, laager than broad, oordaM, atnai^y
oonatriotnd before the raised, Uterally-projeoting hind angles, tlie
tmnsverse basal groove not extending aorasa the diso; dcoady
soabroao-punotulate. Elytra long, aubparallel in their baaal haU,
barely twioe the width of the prothorax; closely, finely punctab>-
striate, the striae deeply impreBsed at the spex, the inleraticea morn
or less convex, alutaoeoua, 1, 3, and 5 with a scattered series of
setigerouB impreesiona extending throughout their kngUi, (he
impressions each preceded by a small tubercle, 9 also with two
impresBions near the tip. legs ve^ long, the tibiae and taisi
pilose; antenor femora atnwgly dsvate, intermediate fentora
thickened before the apex boieath, awl intermediate tibiae widened
from near the base and clueely pubesocnt within, in S- Pmia-
sheath very lorkg, stout, curved, concave above, rounded at tip.
Length 10, breadth 3 mm. (3i.)
Hab. Braziu Ceara {GoaneUe, (J : type), Jatahy, Prov,
of Goyas (Pujol, ?).
Two apeoimens— a J with the penis-sheath ftUly extruded,
the other assumed to be a ^ of the same species. Very tike
iS. purpumpeHHts. l*ic, also from Jatahy, but with sericeous,
opaque, black elytra, and fewer setigeroua impressktnB on
l„y,l,Ajh,COOl^lC
Nea aitd liole-knoiim LagnHae. 223
the inteistiGes I, 3, and 5, those on 7 wanting. The sexes
differ in the form of the intermediate legs, the anterior
femora of the ^ are also stron^y cUvate.
68. Statin euydns, n. sp. (Plate XD, fig. 17, J)
Elongate or moderately «longAt«, robust, ahfaung ; ruf o-cutsiieoiiB,
oastaoeoDB, or piceoiu, the npper anrfsce more or len araieons, the
dftn OBually metaJlic greoi, the spicai joint of the antom&e
femiginoiu in one or two of the darker ezsmplea; the elytn with
a tew long, erect, hriatljt haiia. Head short, bn»d, dauelj, finrijr
punctate, the ejes very large, nanowly sepaiated ; antennae stout,
moderatdy long, joiata 3-10 anbeqaal in length, II in ^ abont
eqnalling 7-10, in $ S-10, united. Prothorax stronf^y ttansveree,
aa wide as tiie head, anmately dilated anteriorly, in some speciiuens
diatincUy angulate at abont one-Uurd from the apex, marpned and
rapidly narrowed thence to the deep boaal groove, the basal ma^in
much TBised and laterally projecting; densely, minutely punotat«,
the narrow intenpacea still more minutely punctate or scabrous.
Elytra lather dongate, one-half broader than the protiiorax,
gradually widened to a little beyond the middle, rounded at the
apex; olosedy, findy punctato-Btriate, the intersticea 3, 5, and 9
with from about six to eight conspicuous setigerous tmpicsaiouH
aoattered between the base and apex.
Length Sf-lO^, breadth 2-3J mm. ( j$.)
Hab. Amazoks, Ega [type], Santarem, Para (H. W. Bates),
Bahia (fry).
Eleven specimens, all but two from the Amazons, these
insects varying greatly in size and in the relative length of
the elytra. S. earydera is a southern form of the Mexican
S. crassicomis. Champ., differing from it in the much
rougher, more densely punctulate prothorax. The pro-
thorax is obliquely compressed along the flanks posteriorly,
so aa to appear marked above.
69. SUtlra latevlUata, n. sp.
$. Eltuigate, narrow, robust, moderately shining; rufous or
caataneous, the eyes black, the palpi piceous, the prothorax slightly
infuscate, or with the sides piceous, the elytra with a broad metallic
green s^pe extending down the out«r half from the base to near
the apex. Head broad, densely, finely punctate, the eyea very
large, narrowly separated ; antennae moderately long, stout, joints
4-10 slightly increasing in length, 11 as long aa 9 and 10 united.
iM-,i,A J h, Google
224 Mr. G. C. Champion on
Prothorax a« wid« as the heftd, much broader than long, tnni-
votmIj coidate, strongly oonatrict«d before the prominoit hind
angles, the basal margin much raised; densely, finely pnnctaU,
the narrow interspa<Ma somewhat scabrous. Elytra long, about
one-halt broader than the prothorax, very gradually widened to
beyond the middle, flattened on the disc, and rounded at the apex ;
closely, finely striato-punctate, the interstices alutaceous, flat, 3,
6, and 9 with one or two inconspicuous setigerous impressions towards
the apex.
Length 6)-8, breadth 2-2} mm.
Hob. Amazons, Para {H. W. Bates).
Two examples, one with the tips of the antennae wanting.
This species has the head, eyes, and antennae shaped as in
S. eurydera, and the body coloured as in iS. vindh-ittata,
all three insects inhabiting the Amazon region. The
brilliant green lateral stripe on the elytra extends downward
to the epipleural margin in <S. kUevitUita. The Colombian
S. medialis, Makl., must be an allied foim, with a longer
and smoother prothorax, the dense puncturing of the latter
giving; it a scabrous appearance in the Amazonian insect."
70. StattTB vlridlTltUta, n. sp.
J. Elongate, narrow, robust, flattened above, shining; rufo-
oaslaneous, the elytra each with a broad golden-green submarginA]
stripe (between the striae 4 and 7) extending from the base to aeur
the apex, and with a few long, erect, bristly hairs towards the tip.
H<«d not wider than the prothorax, sparsely, minutely punctate,
longitudinally impressed between the eyes, the latter very large and
somewhat narrowly sep*nted; aatamae very stout, rather short,
joints 4-10 rapidly decreasing in logth, 6-10 traosveise, II veiiy
elongate, about equalling S-IO united. Prothorax longer than broad.
Rubeylindrical. obliquely narrowed anteriorly and sinuate at the
Rides before the base, the basal margin not much raised ; spandy,
minutely punctate. Elrtra limg. one-half wider than the pro-
tliOMX. nubparallel in their basal half, rounded at the tip; doeely,
Knel,v Hi riato- punctate to near the apex, striate posteriorly, the
inlfrstiom bn^ad, flat on the disc, 3. 5. and 9 with two or three
iiiot>i)apioui>ua uidoly !>catler«d fine, setigerous impressionB, each
111 uliii'h is prr^^lixl bv a minute tubercle. Anterior femora
Mr,..4:l.v.l«v«r.
Uunil. fij. brf«.lii. 21 u.m.
» ^'. n'W*;^..!.'... IV. M>'Ua«es exot.-ent. xxr. p. 20 (Aug. 1917}.
fiMiw Itraait, w i><nii>an>l »ith .■^. -tlutlii. Makl.; but it is impos-
«bl<» t.i Hlt>uiif\ -ii.h iM«sts from ■ ileacriplions abr^g^es."
l„y,l,A J h, Cookie
Hew and Utde-known Lagriidae. 226
Hab. AiuzoNS, Santarem (H, W. BaUa).
One male. A robust, elongate, Bhining, castaneous
insect, with a broad brilliant metallic green submarginal
stripe on each elytron, the antennae very stout, with ths
outer joints tiansveiBe and the terminal one as long as the
preceding six united, the prothorax subcylindrical, the
head rather narrow. It is just possible that S. viridiviOata
may be a male of the Colombian 5. validicomis, Makl.
(described as subcylindrical, with a narrow green sub-
mai^al stripe on the elytra, and very stoat antennae,
the apical joint equaUing 8-10 united) ; but without com-
parison of the types, it would be unsafe to refer insects
from such distant localities to one species. The general
fades is not unlike that of 8. vremturalta, Yic.
71. Statira lalaraUi.
Siat^ lateralu, Makl., Ofv. Finska Vet.-Soc. F6rh., xz, p.
357 (1878).
$. Elongate, depraaaed, widened posterioily, Bhining; testaceous,
the eyM black, the eJytn with a taint, indeterminate, metallio green
marginal stripe extending (rotn the base to about two-thirds of their
length and curving inwards to the middle of the disc poeteriorly;
the eljtiA with a few bristly hairs near tbe apex. Head lai^,
sparsely, minutely punctate, with a shallow arcuate impression
between the eyes, the latter large, well -separated ; antennae slender,
nther short, joint 11 equalling 8-10 united. Prothorax about as
wide as the head, as long as broad, cordate, dilated at the base, the
basal margin prominent; closely, finely, conspicuously punctate.
Elytra elongate, twice as broad as the prothorax, widening to beyond
the middle, and somewhat acuminate at the apex; closely, finely
panetato-striata, the interstices becoming convex towards the apex,
3 only with one or two setigerous impressions near the tip.
Length 9, breadth 3 mm.
Hab. Colombia [type], Ibt^e {Mus. Brit.).
One example, labelled with the MS. name S. scapularis,
Germ. This insect agrees with Maklin's diagnosis of
S. lateralia, but as there are allied forms in Brazil, it is
advisable to redescribe the single specimen before me. 8.
aeneoUncta, Champ., from Mexico and Guatemala, is of
about the same size and shape.
TBANS. BNT. SOC. LOHD. 1917.— PAST 1. (NOV.) <)
ih, Google
226 Hr. 0. 0. Champion on
72. Statin suffusa, n. ap.
$. Elongate, depteased, rather narrow, shiiuDg; piceoua, the
antminae, tarsi, bases of the femora, and elytra teetooeovs, the
elTtra with a broad, uidet«ntimato, metallic green marginal stripe
extending from the base to the apex, the disc with a ver;^ faint
aeneous lustre; the eljtni with a few long, eroot, bristl]' hairs. Head
rather narrow, sparsely, minutely punctate, obsoletely foreate in
the middle between the eyes, the latter large, distant; antennae
slender, rather short, joint U nearly eqnsJling 8-10 nnited. Pro-
thorax a little longer than broad, oblongo -cordate, dilated at the
base, the basal margin feebly raised ; sparsely, minutely punctate.
Elytra elongate, twice aa broad as the prothorax, gradually widened
to beyond the middle, and somewhat aouminate posteriorly ; closely,
finely punctate -striate, the intoraticea becoming convex towards the
tip, 3 and 5 with from six to eight aetigerous impiesaiona scattered
between the base and apex, 1, 7, and 9 also with from one to three
similar impressions on the apical half,
(?. Rufo-teetaoeous, the elytra with a distinct aeneous lustre, the
green maipnal stripe wanting; antennae much longer, joint II tc^
elongate, nearly equalling 7-10 united; alternate elytral interstioes
with fewer aetigeroua impressions — five on 3, and two on 6, those on
I and 7 wanting.
Length 8, bieadtii 2f-21 mm.
Hab. Brazil [type], Parana {ex coU, Fry : (J).
Two specimenft, almost certainly the sexes of the same
species, the Parana male differing as stated above. Very
near the Colombian S. lateralis, Makl., but with a much
narrower head and prothorax, the latter very sparsely,
minutely punctate, the el}^ra with scattered setigerous
impressions, along the interstices 3 and 5, the marginal
stripe (when present) extending to the apez, the apical
joint o{ the antennae (9) shorter. A simiUr variation is
common to other members of the genus.
73. Statira Miwomarglnata, n. sp.
$. Elongate, depneeed, shining, the head and prothorax sub-
opaque; teetaoeDiu,theeyee black, theheadondprothoraxnifeeoent,
the aides of the elytra indeterminately aoieo-piceous from the base
to far beyond the middle, the elytra with a few bristly hairs. Head
d«iselyscabroso.punotulate,the eyes small, distant; antennae short,
slender, joint 11 equalling 8-10 united. Prothorax a little ni
than the head, longer than broad, moderately romided at the si
Ij.y.l.AjL.yCOO'^lL'
Nea mtd liuUrhwvm Lagriidae. 227
feebly conatricted towaida the base, the haa&l margin not raised;
deasely sonbroso-punctnlate. EiytiH long, at the base twice as wide
as the protboTBi, widening to the middle, somewhat acuminate
posteriorly; cloaely, finely, shallowlj punctato -striate, the inter-
stices flat throughout, 3 with four small setigerous impressions scat-
tered along the spical half, 5 also with two other impressions, and
7 with one, towards the tip. Legs slender.
Length 7|, breadth 2) mm.
Hab. Brazil, Eio de Janeiro (Fry).
One specimeD. A small pallid insect, not unlike the
Colombian S. lateralia, Makl., and the Brazilian S. suffvsa;
but with a dull, scabrous head and prothorax, shining,
aeneo-marginate elytra, the latter with the inteistices flat
throngbout.
74. Statin fraohyden, n. sp.
Very elongate, narrow, feebly shining ; piceous, the front of the
head, elytral sutnre, tarsi, bases of femom, and under surface oaa-
t«neoas or ferruginous; the elytra with scattered bristly hain.
Head rather small, closely, coarsely pimctste, longitudinally im-
pressed between the eyee, the latter large, narrowly sepanted;
antennae moderately thickened, rather long, joint 11 equallii^ 8-10
united. Prothorax a little wider than the head, longer than broad,
rounded at the sides, constricted before the raised basal margin ;
coarsely, irregularly, confluently punctate. Elytra very elongate, at
the middle about twice as wide as the prothorax, somewhat acumin-
ate poeteriorly; closely, finely, deeply punctate -striate, the inter-
stices convex, flatter on the disc, 3 and 5 with a series of small
aetigGTDuB Impreesions scattered throughout their length, 9 also with
three impressions near the apex, and 1 with sevenil others near ths
base. Tibiae pilose within.
Length 11, breadth H mm. ($T)
H(d>. Brazil, Jatahy, Prov. of Govas {Pujol, ex cdl.
F. Bates).
One specimen. This insect is of about the same eize
and boild as S. jmrjnireipennis, Pic, from the same locality,
difietiog from it in the small, closely punctured, subsulcate
bead, the coarsely, confluently punctate prothorax, and the
non-tuberculate elytra, the elytra with fewer, differently-
arranged aetigerous impressions, those on the interstices
1 and 7 mostly wanting.
iM-,i,A J h, Google
228 Mr. G. C. ChampioD on
75. Stfttln soabrieoUls, n. sp.
$. Elongate, dull, the eljtra more shining; pioeous, the elytn
brown, the antomae (joint 1 excepted) and legs (the infnecate outer
halvee of the femora excepted) testaceous; the head, elytra, and
under surface with a few long, erect, bristly hairs. Head barely as
wide as the prothorax, densely, minutely punctate, hollowed between
the eyee anteriorly, the latter large, narrowly separated ; antennae
moderately long, quite slender, joint 11 equalling 7-10 united. Pro-
thorax slightly longer than bioad, oblongo -cordate, constricted before
the prominent, laterally-projecting basal margin ; densely, minutely,
soabroao-punctate, thediscwith two small transveraely-plaoed toveae
before the middle ( I accidental). Elytra long, twice as wide as the
prothorax, widening in their basal half, acuminate pOHteriorly;
closely, finely, deeply puncta to -striate, the interatioea convex
throughout, 3, 6, and 9 witJi several small setigerous impressions
aoattered between the base and apex. Femora rather stout.
Length 8}, breadth 3 mm.
Hab. Upper Ahazonb, Ega (H. W. Bates).
One female. A close ally of S. iTistdaria {No. 77), from
Trinidad, with the elytra moie acuminate at the apex,
deeply punctato-striate, and the interstices convex through-
out ; and the femora nigro-piceous in theii outer halves.
The eleventh antennal joint is no doubt equally elon-
gate in c^, to judge fiom its length in 9- The Brazilian S.
flamcomis, Makl., is probably another allied form.
76. Statira convexluuula, n. sp.
$. Elongate, rather convex, shining; testaceous, the head and
prothorax rufescent, the eyes, palpi, femora (except at the base),
and tibiae black or pioeous, the elytra with a few bristly haira. Head
ratlieF small, ikotquitAso wide as the prothorax, densely, very finely
punctate, obsoletely foveolate between the eyes, the latter moder-
ately large, distant ; antennae short, slender, joint 11 nearly equaling
8-10 united. Prothorax oblongo-oordate, constricted before the
baae, the basal margin raised, but not very prominent laterally;
almost smooth, except within the transverse basal groove, which is
shallow in the centre. Eljftra moderately elongate, aboot twice
as wide as the prothorax, narrowing from the middle ; oloeely, finely,
rather deeply punatat« -striate, the interstices feebly convex, 3 and
6 with three or four uiconspicuous, setigerous punctures scattered
down the apical half, and 9 with three others near tiia apex. L^a
rather short, the anterior femora olavate.
Length 3}, breadth 2} mm.
n,g,t,7.dh,G00glc
New and Ultle-kHown Lagriidae. 229
Hd>. Brazil, Bahia (Reed).
One female. Not unlike S. scabricoUis (9), from Ega,
but in great part testaceous and more shining, the eves
smaller and more distant, the ant«nnae shorter, the pro-
thorax almost smooth, the tibiae and the basal halves of
the femora black. S. mdanocephala, Makl., from 0«ntTal
America, is a somewhat similar insect, except that it has
the bead and antennae infuscate or black.
77. Stetira ImnUrls, n. sp.
Elongate, the b«Ml and prothorax sabopaque, the elytn Bhining ;
teetaoeooB, tb« head and prothoraz obBoure ferruginous, the eyes
black i the elytra with a few long, bristly hairs. Head rather small,
closely, minutely punctate, longitudinal ly depressed down the middle
betweoi the eyes, the latter extremely large and narrowly separated
in the two sexes ; antennae long and slender, shorter in $, joint 11
in ,; as long as 6-10, in $ the length of 8-10, united. Prothorax as
iride as (3) or rather wider than ($) the bead, about as broad as long,
feebly rounded at the sides, strongly constricted before the base,
the basal margin prominent and laterally projecting; alutaceous
and closely, very minutely punctate. Elytra long, at the middle
twice as broad as the prothorax, somewhat rapidly widened in their
basal half, and acuminate posteriorly! closely, finely, deeply puno-
tato-striate, the intersticee becoming oonvex towards the sides
and apex, 3, 6, and 9 each with from three to six widely scattered
setigeroue impressions.
Length S-8}, breadth 2i-2i mm. (3^.)
Hab. Tbinidad {G. E. Bryant, iji. 1903; /. Birch, 1904).
Three males and one female. Very like the Antillean
iS./u2va, but with a long apical joint to the antennae, espe-
cially in (7, ft comparatively small bead, extremely large,
subapproximate eyes, and rathei convex, mesially widened,
postenorly acuminate elytra. S. tdefisis. Champ., from
Panama, is not unUke S. irutularis, but the latter has the
elytra less elongate, etc. Mr. Bryant's specimens have
been examined by M. Pic and returned unnamed.
78. Statlta Inlva.
Stalira fulva, Fleut. et SallS, Ann. Soc. Ent. Fr. 1889,
p. 431."
iM:,i,A J 1^, Google
230 Mr. G. C. Champion on
Elongate, somenlut deprewed, th« bead aad piothoiax opaque,
0x0 elytra Bhining; teataceoua, the bead and prothorax obsoura
ferniginouB, the ejee black ; the elytra with a few long, eieot, bristiy
hairs. Head obsoletely punctulate, the eyes large and wdl aepanted ;
antennae long, alender, joint II about as long as 9 and 10 united in
die two sexes. Prothoraz aa wide aa the head, oblongo-oordate,
dilated at the base, densely alutaceoua, the minute scattered pnnc-
turea just traceable. Elytra long, widened to the middle and there
twice as wide as the prothorax, acuminate posteriorly; oloeely,
finely, deeply punctato-striate, the interstices rather convex, 3 and
6 with several widely scattered setigerous impreeaionB, 1, 7, and 9
also with one or two similar impressions towards the apex.
Length Ti-lOi, breadth 2i-3 mm. (^5-)
Hab. Antilles, Guadeloupe [type], Dominica {Mus.
BrU.).
Two specimens from Dominica, one of them received
from Mr. E. F. Becher in 1908, are no doubt referable to
the imperfectly -described S. fnlva from Guadeloupe. In
the Museum there is also another example, without locality,
from the Dejean collection, apparently belonging to the
same species ; it is labelled " S. humeralis ^^ var. ? (Dupont)."
S. ftdva is a form of S. viUala, Champ., from the Lesser
Antilles and Trinidad, with a densely alutaceous, opaque
prothorax, and slightly longer, wholly testaceous elytra.
It has been found " at light " in both islands.
79. Statin vlHata.
Slatira vitUUa, Champ., Tians. Ent. Soc. Lond. 1896, p. 37,
pi. 1, fig. 9 (J).
Hab. Lesser Antilles ; Trikidad.
In the British Museum there is a series of this insect
captured long ago in St. Vincent by Lansdown Gailding,
also two from Trinidad, in Fry's collection.
80. SUtira asperata.
Stalira asperata. Champ., Biol. Centr.-Am., Coleopt. iv, 2,
p. 49, pi. 3, fig. 4 (,?) (1889).
Stalira anliUarum, Champ., Trans. Ent. Soc. Lond. 1896,
p. 36.
Hab. Panama; Lesser Antilles, St. Vincent, Grenada,
" Neo S. hvmeralis, Makl., a Mexican insect.
New and fiule-^notra Lagriidae. 231
Grenadines; Colombia; Venezuela; Tbikidad (&. E.
Bryant); Bbazil, Femambuoo {Fry).
The long series before me connect the Antillean with the
mainland form, Panama examples having conapicuoua
tubercles on the alternate elytral interstices 1, 3, 5, 7 and 9.
One of the Pemambuco specimens is rufo-castaneous with
the elytra piceous, and another is rufo-castaneous with the
sides of the elytra broadly piceous.
81. Statira presnturalb. (Plate XII, fig. 18, $.)
Slatira preauturalis. Pic, L'£change, zxviii, p. 76 (Oct.
1912).
Aatawae not longer in (J Uian in $, rather atont, joint 1 1 equalling
9 and 10 united in the two sexes ; head with a targe shallow inter-
ooolar fovea; prothorax elighttj longer than broad, veiy finely,
spoTMly punot«t«; etTtni depressed below the base, very finely,
closely striato-punctate, the punctures usually placed in fine striae
on the besal half, the interstice 3 with several, and 5 and 9 with one
or two, widely separated setigerous impieesions on the apical half;
body nther convex, castaneous, shining, the palpi and prothorax
sometimes mfuscate, the juxta«utural blackish stripe on the elytra
often oontinoed to the apex, dilated beyond the middle into a com-
plete or interrupted tronaverM fascia, and widened again at theapes;
femora strongly olavate; aede^naof S rather broad and acuminate.
Femora more or lees clavate.
Length SHI' >»»^th 2-3 mm. 139-)
Hah. Brazil {Mus. Brit., Mus. Oxon.), Blumenau {coU.
Pic : type), Bio de Janeiro (Fry).
The long series of this insect before me agree sufQciently
well with Pic's brief diagnosis to render the identification
certain. Froni the somewhat similar 8. siUuralis, Makl.
(No, 6), from Rio de Janeiro, etc., it is at once separable by
the absence of the long spine at the base of the anterior
femora, the stouter antennae, with shorter apical joint in
t^, the longer prothorax, etc. The juxta-sutural stripe is
often dilated into a poet-median fascia and an apical patch.
82. Statira oetollneata, n. sp.
Somewhat robust, moderately elongate, shining; obscure cas-
taneous, the le^ and antomae ferruginous in one speoimoi,
ibe alternate elytral interstices 2, 4, 6, 8 each with a narrow
blackish stripe extending to near the apex; the elytra wiUi a
n,g,i,.rjh,G00glc
232 Bli. 0. C. Champion on
few, erect, briBtly hairs. Head almost emooth, excavate or
foveate in the middle between the eyes, the Uttm moderately
lai^ Hid separated by about the width of on« of tbem ; antennae
rather slender, moderately long, a little shorter in $, joint 11 as
long as 9 and 10 united. Prothorax as wide as the head, aa
long as broad, oblong-ooidate, dilated at the base, very sparsely,
minntely punotulate. Elytra moderately long, broad, twioe ae wide
as the prothorax at the base, flattened on the dlao anterioily,
slightly widening to the middle and a<!ununat« posteriorly, the apioea
distinctly macronata; closely, finely punotato-atriate, the inter-
Btiae« becoming feebly convex towaids the apex, 3 and S each with
about five very widely scattered settgerouB impressions, B also with
two or three similar impresBions towards the tip. Femora more or
leas olavate.
For. The elytra uniformly caatoneous, t^e setigerons impreesiiMU
wanting on the baaal half of the third interstice. {^.)
Length Sf-S^, breadth 2i-2i( mm.
Hab. Brazil, Kio de Janeiro (Fry), Constancia {J. Gray
and H. Clark, Jan. 1857).
Two specimens of the Imeate lonn, assumed to be (? and
$, one of them having longer antennae than the other,
and one of the variety. More elongate than 8. presutundis.
Pic, the head and prothorax a little broader, the elytm
longer, moie acuminate at the tip, and with the alternate
interstices nigro-lineate to near the apex.
83. SUtira triUneala.
Statira trilineata, Makl., Ofv. Finska Vet.-Soc. Forh. xr,
p. 357 (1878).
<}. Antennae rather short and stout, joints 4-10 decreasing in
length, 0 and 10 traoavene, 11 very elorkgate, as l<Hkg ae 6-10 united ;
head deeply foveate in the middle between theeyesj the elytra with
an elongate intra-humeral depression, the scattered setigerons im-
preeeions along the entire length of the alternate interstices 1, 3, 5,
7, and 9 each preceded by a small tubercle; femora more or less
Hab. Colombia {Mus. Hdaingjors : type ; Mus. Brit.].
A male in the British Museum, from the F. Bates col-
lection, is almost certainly referable to this spedes. It
diiTers from the description in having the nometous setiget-
oiis impressions along the alternate elytral interstices
deeply impressed and each preceded by a sm^, tumid,
l„y,|,AJI^,COOl^lC
Nob and tittle-known Lagriidae. 2S3
tubetcniifonn space. There is conaiderable variation in this
respect in the allied S. asperata. Champ. S. triXinetUa is
of a bright nifo-caataneous colour, with a blackish line along
the elytral suture and another down the fifth interstice.
S. prestUuTolis, Pic, is a very similar form.
84. Statira tennis, n. sp.
ModeTat«ly elongate, narrow, Bhming; CMtaneous or obsouie
oostaneouB, the anUnnae in the male piceouB with the apical Joint
f emigiiKius ; Uie elytra with a few very long, erect, bristly hairs.
Hwd ratiiet small, rounded, almost sntootit, the eyee very large and
narrowly separated ; ant^mae {ij) slender, long, joints 7-10 decreas-
ing in length, II extremely elongate, about equalling 3-10 united, ($)
Bhortet and stouter, 11 barely ^e length of 7-10 united. I^thorax
about as wide as the head, longer than broad, oblong, moderately
constricted before the base, sparsely, minutely punctate. Elytra
about twice as broad as the prothorax, moderately long, but little
widened towards the middle and acuminate posteriorly ; very finely,
closely, striato-punotate, the interstices flat, 3 with about six promi-
nent setigerouB impreaaions scattered between the base and apex,
and 6 and 0 eacli witli thi«« or four similar impressions on the apical
half.
Length ef-7j, breadth 1^-2^ mm.
Hob. Amazoks, Ega [type], Santarem {H. W. Bates).
Two males and one female. Not imlike the Brazilian
S. stenodera, but with an extremely elongate apical joint
to the (^-antennae, large, subcontiguous eyes, and much
narrower elytra, with flat interstices. The general shape
is similar to that of the Colombian S. IriliTieata, Uakl., as
here interpreted.
85. Statita atenodera, n.sp.
Moderately elot^te, shining i rufo-caataneous or oastoneous, or
piceo-oaataneouH with the antemiBe and legs in great part reddish,
the elytra often faintly streaked with piceous on the alternate in-
l«TBtioea; the head and elytra with a tew long, erect, bristly hairs.
Head sparsely, very finely punctate, well developed behind the
eyes, and with a large, shallow, punctured fovea between them, tlie
eyes rather smalt (as seen from above) and distant ; antennae slender,
joint 11 the length of 9 and 10 united in the two sexes. Prothorax
slightly longer than broad, nearly as wide as the head, oblong,
modeiately oonstrioted before the base; sparsely, minut«ly pun<!-
tate. Elytra not very elongate, at the base twice as broad as the
iM-,i,A J h, Google
234 Mr. 0. C. Champion on
prothoTAS, gToduall; widened to the middle and npidly narrowed
poatoriorif, muoronate at the tip, tianBTereely depiened on the disc
antoriorlj; oloaely, finely punctato-etriate, the intereticee feebly
oonTBX, 3, S, and 9 with from two to f oor widely Boatt«red setigennu
impieeaioiu. Anterior femora clarate.
Length 6-6}, breadth 2-2t mm. (,?$.)
Hob. Brazil (Mus. Brit., Mus. Oxon.), CoDstancia,
Tijuca {/. Gray and H. dark, Jmi. 1857), Rio de Jafleiro
{Fry), Sao Antonio, Bahia ((lOuneUe).
A long serieB, those in the Oxford Museum in very bad
condition, some of them (immature) obscure testaceous in
colour. Separable from the closely allied S. presuturalis.
Pic, by the relatively narrower bead and prothorax, the
smaller eyes, the more slender antennae, etc. ; from S. octo-
lineata by the much shorter elytra, narrower head and
prothorax, and smaller eyes ; and irora 8. svtwalis, JA&ki.,
by the absence of the anterior femoral spine, etc. This
is one of several members of the genus with the eleventh
antenna) joint similar in length in the two sexee.
86. Statin egaen^ n. sp.
Moderately elongate, widened poateriorly, shining ; rufo- or fuaoo-
castaneoiu, the eyee blaek, the eljrtra and legs (the knees and bases
of the femora eiccepted) nigio-piceoue or piceous, in one specimen
almost oonoolorous witik the rest of the surface; the elytra with
numerous long bristly hairs. Head broad, well .developed behind
the eyes, Bparaely, 6jiely punctate, the eyee moderately large, dis-
tant; antennae slender, comparatively short, joint 11 equalling 9
and 10 united. Protfaoias as wide aa the head, broader than long,
transversely cordate, constrioted in front of the promintDt hind
angles, the basal margin not much raised; sparsely, finely, con-
spicuously punctate. Elytra not very elongate, in their widest part
more iJian twice the bieadth of the prothorax, rapidly dilated to
the middle, and arouately narrowed thence to the apex; somewhat
ooarsdy, closely, orenato-striate, the punotureeon thedisc tranavetBe,
the inleraticee more or lees convex, 1, 3, 5, 7, and 9 each a row of
rather closely placed setigerous impressions extending from the base
to the apex. Anterior femora strongly clavate.
Leagth 9i~l, breadth 2i~2i mm. ($T)
Hab. UppEK AMAZ0N3, £ga {H. W. Bates).
Three specimens, varying in the colour of the elytra and
legs, as is frequently the case in this genus. S. egaensia
resembles S. slenodera in it« general faaes ; but the head
l„y,|,AJh,COOl^lC
Nea and ItUle-known Lagriidae. 235
and prothoiax an much broader, the piotbonuc is trans-
verse, and the alternate elytral mteistices have each a row
of setigerous impressions extending throughout their length,
much as in the Central American S. microps, Champ.
87. Statlra eleguu- (Plate XII, fig. 19, anterior leg, ^.)
Statira d^ans, Makl., Act, Soc. Fenn. vii, p. 156 (1862).
Var. Slatira /estiva, Makl., loc. cit.
Antennae in ^ with joint 11 about as long aa 7-10, in $ as long
aa 8-10, united; anterior femora davate; anterior tibiae abruptly
oaiTOwed and oompreeBed at the base externally, subangulale at
basal third in ^.
Hab. Brazil, Petropolis {J. Gray and H. Clark, Feb.
1857), Rio de Janeiro, San Paulo, Espirito Santo {Mus.
BrU.).
A dozen specimens before me from the above-mentioned
localities agree with Maklin's description of S. degans,
except that they have the prothorax very finely alutaceous
(thus appearing eubopaque), instead of shining, as stated
by him ; but this is probably an error of observation. A
rather narrow, moderately elongate form, with the head
(the eyes excepted), antennae, prothorax and legs testace-
ous or rufo-testaceouB, the elytra uniformly green or bluish-
green, and the metastemum and abdomen black; the
antennae slender ; the eyes la^e, the space between them
longitudinally impressed in the middle; the prothorax
strongly constricted before the base, canahculate on the
disc, and almost impunctate ; the elytra closely, finely
punctato-striate, with seven or eight setigerous impressions
scattered along the third interstice and one or two others
at the apex of the ninth ; the anterior tibiae compressed
at the base. S./estitxt, to judge from the description, is a
variety of S. degana with the head (except in front) and
prothorax piceous. The head in one of the specimens before
me is infuscate, but not the prothorax. These insects might
easily be mistaken for similarly-coloured Carabids.
88. Statint dlstipiu, n. sp. (Plate XII, fig. 20, prothorax, S)
3. Uoderately elongate, narrow, ehiimig; testaceous, the eyes
and abdomen black, the elytra metallio blue, the apical joint of the
antennae and the posterior femora (except at the base) and tibiae
tl^tly iufoBoatei the elytra with a few long, erect, bristly hairs.
iM-,i,A J h, Google
236 Ifr. G. G. Champion on
Head raUier large, sparaely, ehallowly BufafoveoUte between tlie
eyes, the latter large uid somewhat distant; antennae slcoder,
joints 8-10 deorea«ing in length, 11 as long as 7-10 united.
Frotlioraz nanower than the heed, about as long aa broad,
oblongo -cordate, strongiy constricted before the base; sparsely,
minutely punctate, obeoletely canalioulate at the middle of the disc,
and with a la^e, subtriangular, rugose, depressed area on each side
(fig. 20). Elytra moderat«!y long, at the middle about tnice oa wide
aa the prothorax, rounded at the apex, longitudinally deprceaed
within the humeri ; very finely, closely punctato-atriate, the third
interstice with six widely scattered setigeroue iropreesions, the
ninth oiao with two others near the tip. hegp long; anterior
tibiae rounded externally at the base.
Length 7}, breadth 2} mm.
Hab. Peru, Chanchamayo (ex c<M. F. Boies).
One male, probably collected by Thamm. Very like
the Brazilian S. eUgans, Makl., and similarly coloured; but
with the prothorax mote shining, distinctly punctured,
and with a large, subtriangular, rugose, depressed area on
each side of the disc (somewhat as in the Central American
S. fovekoUis, Champ., ^, and possibly confined to that sex),
the posterior femora and tibiae partly infuscate, the anterior
tibiae not compressed at the base.
8d. StaOra oyuoptera, n. sp.
[J. Elongate, narrow, depressed, slender, opaque, the elytra and
the front of the head somewhat shining ; black, the elytra cyaneous,
the palpi at the base, joints 4-11 of the antennae, the tArsi, and in
one apecimen the legs entirely, testaceous; the elytia with a few
long, erect, bristly hairs. Head rather narrow, sparsely, finely
punctate, shallowly aulcate between the eyes, the latter large,
separated by about half the width of one of them; antennae long,
slender, joint 11 extremely elongate, equalling 6-10 united. Ite-
thorax nearly as wide aa the head, much longer than brood, oblongo-
oordat«, feebly dilated at the baee, amooth, the basal margin not
laiaed. Elytra moderately elongate, twice as broad as the pro-
tiiorax, somewhat acuminate posteriorly; closely, finely punctato-
atriate, the punctures transverse on the disc, the inleratioee feebly
convex, 3 with five or six, and 5 with three, widely scattered aetiger-
ouB impreasiona, 1 and 9 also with one or two near or before the tip.
li^B long and slender.
Length OiV-'i^j. breadth 2-2jV mm.
n,g,t,7.dh,G00glc
New and litUe-kiunon Lagriidae. 237
H(di. Brazil {Mus. Brit. : type), Rio de Janeiro (Fry).
Two examples, one with the tarei only, the other (from Bio
de Janeiro) with the legs entirely, teataceoua, showing that
no reliance can be placed on the colour of the legs as a specific
character. A narrow black insect, with the elytra blue and
slightly shining, the prothoraz long, smooth, and opaque,
the antennae and legs long and slender, the antennae
testaceous, with joints 1-3 blackened.
90. Statlra eai&miloldss, n. sp. (Plate XII, fig. 21, 3.)
Uodeiatelj elongate, narrow, depreesed, rather dull, the elytra
Bhining; pioeotu, the head black in one speoimea the femora and
tibueBOmetimeBpalerthan the bodj, the ant«mnae (joint 1 exerted)
and tarai testaceous; the head mmI elytra with very long, ereot,
HCattored bristly hairs. Head large, broad, sparsely, finely punc-
tate, in two speotmens, longitudinally depressed between the eyes,
the latter large and moderately distant in t^, a little smaller in $;
antennae slender, moderately long, joint II in (^ about as long as
7-10, in $ not quite equalling 8-10, united. Prothorax narrow,
longer than broad, oblong -camponulate, sparsely, minutely punctate,
the interapaces alutaceous, the basal margin prominmt. Elytra
moderately long, at the base, twice as wide as the protborax, sub-
pandlel in their basal half in 3, with a very deep, oblique depression
on the disc below the base; shallowly, minutely striato -punctate,
the intATstioes flat, 3 with about five, and 5 and 9 with thi«e or four,
widely scattered conspicuous setigerous impressions, 1 also with a
single impieasion near the tip, Hie imprMsions eaeh preceded by a
minute taberole. Legs slender, long ; ulterior tibiae in both sexes
abruptly rkanowed at the base externally.
Loigth 6[^7i, breadth Ii-2i mm. {3^.)
Hab. Bbazil {Mua. Oxon.), Constancia (J. Gray and B.
Clark, Jan. 1857 : type, <J), Rio de Janeiro [Fry : $).
Five specimens, the one in the Oxford Museum immature
and almost wholly testaceous. Near 5, dromioides, infra,
from which it diSers in having a narrower and smoother
prothoiax, a shorter apical joint to the (^-antennae, deeply
excavate elytra, with fewer setigerous impressions, and
pecuharly formed anterior tibiae, the body also being
uniformly piceous. The large head and narrow prothorax
^ve the insect a Casnoniiform facies. S. el^ana, Makl.,
has somewhat similar anterior tibiae in the two sexes.
iM-,i,A J h, Google
238 Mr. G. C. Champion on
91. Slattn eanvwiiilata, n. sp.
Elongate, lunow, dull; piceoaa, the antennae (joints 1 and 2
ezaept«d} obscure fermginone, the dytra with Mattered loDg, eiect,
black briatty haira. Head large, broad, cloeelj punctate, longitu-
dinally grooved betwem the eyes, the latter large and narrowly
separated; antennae Bhort, rather eleoder, joint 11 about equalling
7-10 united. Prothorax narrower than the head, longer than
broad, campanulate, closely, somewhat coarsely punctate, the basal
margin very prominent. Elytra moderately long, about twice as
broad as the prothorax, subparallel at the base ; closely, finely pnno-
tato-Btriate, the intecBticea with nnmerouB setigerouB imprcBsions,
each of them preceded by a small tubercle — thtee near the apex of 1,
ten scattered between the base and apex of 3, five or aix on 6, and
eight on 9, those on 6 and 9 placed along the apical halt. Legs
moderately slender, roughly punctured and setnloHe, the tibiae
shallowly suicate externally.
Length 8}, breadth ^ mm. ( j T)
Hab. Upper Amazons, Ega {H. W. Bates).
One badly preserved axatnple. iMtgexthtta S.dromimdes,
wholly piceous, the eyes more approximate, the apical
auteimal joint shorter, the prothorax longer and more
coarsely punctate, the legs roughly punctured. The seti-
gerouB impressions on the elytra are similar in number and
position in the two insects.
92. Statira dromloidM, d. sp.
S- Elongate, narrow, depressed, moderately shining; testaceous,
the eyes black, the elytra piceous i the head and elytra with a few
very long, erect, bristly hairs, one behind each eye being oonspico-
ouB. Head large, broad, rather convex, sparsely punctete, longi-
tudinally grooved between the eyes, the latter very large, modetat^y
distant; antennae rather long, alender, joints 4-10 gradually de-
creasing in length, 11 extremely elongate, equalling six or aeven of
the preoeding jointe united. Frothorax much narrower than the
head, a tittle longer than broad, obi ong-campanu late, closely, eon-
apiouously punctate, the interspaces alutaceous, the basal margin
sharply laiaed. Elytra moderately elongate, twice as broad as the
prothorax, not much wider at the middle than at the baae^ narrowly
rounded at the tip; very finely ponctato-striate, the intereticee
alutaceous, almost flat, 3 with about 7 setigeroua impressions scat-
tered between the base and apex, 5 and 9 also with tour similar
impressions along Uieir apical half, and 1 with two or three near the
l„y,|,AJI^,COOl^lC
Nod and liide-known Lagritdae. 239
tip, each of them pieceded by a sin&U tDbe««le. Legs long and
■lender.
Length 6-7, biewlth 2 mm.
Hab. Bbazil, Ceara {GouneUe, ex coU. Fry).
Two males. A narrow, moderately shining, testaceous
form, with piceous, sparsely seriato-tubennilate elytra, a
very broad head, a conspicuously punctured, subcsmpanu-
late prothoTaz, and an extremely elongate apical joint
to the (J-antenna. 5. dromioidea is very Uke various
similarly coloured species of the Carabid-genus Dromius.
93. Stafira ifenoptera, n. sp.
Elongtit«, narrow, the head and protboraz dull, the elytra shining ;
tcstaoeooB or obsonie teataeeoiu, (he eyes black, the aides of the
elytm broadly and indeterminately, the prothorax, and the poaterior
fflmora in t^eir outer half, more or leu inf luoate ; the elytra with a
few very long bristly hairs. He«d large, transrersely convex, much
broader than the prothorax, closely punctate, the eyea Utge, narrowly
separated in ^, moi« distant in $; antennae Blender [outer joints
missing]. Prothonx much longer than bioad, oblong-oampanu'
l»te, constricted before the bii«e, the basal margin raised ; somewhat
closely, conspicuouHly punctate, the intenpacee olntaceous. Elytra
long, narrow, slightly widened at the middle and there twice as
wide as the prothorax, somewhat acuminate posteriorly, deeply,
obliquely depressed on the disc below the base, and also longitudin-
ally hollowed towards the sides just below this; closely, finely
punotato-striate, the interstices almost flat, 3 with five or six setiger-
cms impreesiona scattered between the base and apex, and 1 and 6
with one or two, and 9 with three, similar impreeBiona towards the
tip.
Laigth 6-ef, breadth 1$ mm.
Hab. Amazons, Ega, Santarem {H. W. Boies).
Two specimens, assumed to be <^ and $. A narrow,
fragile form, with a larce head, alUed to S. dTomimdea and
8. casnonioides, the head and prothorax punctured as in the
former, and the elytra deeply excavate below the base as
in the latter; the prothorax is long and narrow, as in
S. dromioides.
94. Statlra qnadristgnata, n. sp.
(J. Elongate, narrow, shining, the eljrtra dull ; testaceous, the
eyes and two brood transTeree fasciae on the elytra (one ba«al, nar-
rowed outwards, the second broader, post-median, straight, neither
iM:,i,A J 1^, Google
240 Mr. G. C. Champion on
reaohing the Butui« or outer margin) black ; the el jtn wi tb scattered
long, erect, bristly haira. Head Bparaely, very minutely punctate,
canaliculate between tlie eyes, the latter very large and narrowly
separated; antennae moderately slender, joint 11 as long as 8-10
united. Prothorax longer than broad, almost aa wide as the head,
subcotdate, dilated at the base, sparsely, very minutely punctate.
Elytra long, rather narrow, rapidly narrowed from the middle, and
pointed at the apex; closely, finely, shallowly punctate -striata, the
alternate interstices 1,3, 6, 7, and 9 with a series of scattered, deep,
setigerous impressions extending from the base to the apex, the
impressions each preceded by a small tubercle. ,
Length 6|, breadth S mm.
Hab. AuAZONS, Santarem (H. W. Bates).
One male. Not unlike S. bryatUi, Pic, fiom Trinidad,
but with the elytral aculpture similar to that of the widely
distributed S. aaperata. Champ. ; the elytra longer and more
acuminate than in the Trinidad insect, and very differently
marked, the tvo broad black fasciae (basal and poet-median)
not leaching the suture and thus forming two large trans-
verse patches on each wing-case.
95. atattra brjranti. (Plate XII, fig. 22, ^.)
Suuira bryatUi, Pic, L'Ek;hange, xzviii, p. 100 (Jan. 1913).
Antennal joint 11 in ^ about as long as 7-10, in $ equalling 8-10,
united.
Hab. Trinidad (0. E. Bryant : iii. 1903).
There are two niales and three females of this specdes in
Mr. Bryant's collection. A narrow, testaceous form allied
to the Central American S. conspiciUata, Makl., the pro-
thoraz with a black submarginal line on each side of the
prothorax, and the elytra with two transverse fasciae
(one just below the base, broad, the other narrower, curved,
median), usually connected along the first interstice, and an
oblong streak on the disc towards the apex (in one specimen
extending forward to the median fascia), blackishormetallic.
The elytra are closely, finely punctato-striate to the tip,
the third and ninth interstices each with from 2-4 scattered
setigerous impressions on the apical third. The prothorax
is rather closely, minutely punctate and more or less dis-
tinctly canaliculate. The description of the similarly
coloured S. quadrimaculaia, Makl., from Colombia, applies
ll,y,l,/eJh,G00glc
New and tittle-knovm Lagriidae. 241
very neariy to the Trinidad insect, except that the pro-
thorax in the former is said to be almost impunctate and
the fine median channel is not mentioned.
96. Stetln vlrldtelnett, n. sp.
Uoderately elongate, narrow, shining; toetaoeouB, the eyes black,
the protliorax wilh a nigro-fuBcoua gubmargmal vitta on each side
(abbreviated or wanting in iinmature exampie«) ; the elytra with
two tranBTeiBC metallio green fasciae of variable extent — ona baaal,
not reaching the humeri, and Bometimea enclosing an oblique tes-
taceous spot on the disc, the other median, angulate, the two aome-
tintee ootmeoted along the suture, and in two apecimena at the sMee
ako ; the elytra with a few long, erect, bristly haiis. Head sparsely,
minutely punctate, tranversely depressed in the middle between the
eyee, the latter large, distant; antennae long, slender, joint 11 in (J
About as long as 7-10, in $ as long as 8-10, united. Prothorax
narrower than the head in ,J, slightly wider in ?, not longer than
broad, cordate, dilated at the base, sparsely, minutely punctate.
Elytra moderately long, gradually widened to tbe middle and there
twice as wide aa the prothorax; closely, finely, deeply punctato-
striate, tbe interaticee somewhat convex, 3 with from aix to eight
oonspiououe setigeroua impressions scattered between the base and
ufiex, 9 also with three similar impreesions towards the tip.
Length 6t-6|. breadth 2-2^ mm. {3<f.)
Bab. AuAZONS, Santarem, Ega (H. W. Bates), Para
{GounsUe).
Nine specimens, varying in the development of the pro-
thoracic and elytral marlangs, and in the puncturing of the
piothoraz. The metaUic coloration on the elytra in the
Para example is so extended as to leave two oblique fasciae
on the disc of each of them, and a common apical patch,
testaceous. It is quite posaible that these insects will
prove to be inseparable from the Colombian S. quadri-
maculala, M&kl., which, to judge from the description, has
a longer, almost impunctate prothorax, fewer setigeroua
impressions on the third elytral interstice (and those
placed towards the apex), and a dark ante-apical epot. S.
bryatUi, Pic, from Trinidad, which ia similarly coloured,
has shorter antennae, a longer and distinctly canaliculate
prothorax, and fewer setigerous impressions on the elytra.
8. viridimitata, Pic, from Cumbase, Peru, must be another
allied form.
TRANS. ENT. 80C. LOND. 1917. — PART I. (NOV.) K
iM:,i,A J 1^, Google
242 Mr. G. C. Champion on
97. Statin eonneu, n. ap. (Plate XIII, fig. 23, 3.)
Rather short, narrow, somewhat convex, shining, glabrous ;
testaceous, the ejee and the base of the antennae, and the elytra with
two transTwse fasciae (one close to the base, the other submedian,
connected along the suture), and in one example (ij) a common,
transverse, subapical patch at th« termination of the narrow Bulural
streak, black or fuscous. Heed large, broad, sparsely, obsoleteiy
punotulate, flattwed between the eyes, the latter Ibi^ and moderately
distant; antAnnae (,}) elongate, slender, joint 11 as long as 9 aitd 10
united, ($) much shorter, joint 11 lees elongate. Prothorai l<mger
than broad, oblongo-subcordate, the basal constriction moderately
deep ; closely, findy punctate. Elytra rather short, twice as wide as
the prothorax, subparallel in their basal half in S> dilated posteriorly
in $, transversely depressed below the base ; closely, somewhat
deeply punota to -striate, the punctures rather coarse on the basal
half, fine on the apical declivity, the interBtioee convex towatds the
sides and apex. Femora strongly clavate.
Length 41-5, breadth li-lln™- (39-)
Hah. Amazons, Santarem [(J. type], Ega [$] {H. W.
Bales).
One pair, the male with the elytial marMngB more
developed than in the female. A small, comparatively short
fonn, not unlike an Anthicua, with the el3^ra fasciate much
as in S. viridicinOa from tlie same Amazonian localities,
and S. bryanti, Pic, from Trinidad. iS. connexa would
perhaps be better placed in the genua Colparthrum; but
till the structure of the mandibles, etc. has been examined,
the insect ia better left under Slalira, The apical joint
of the labial palpi is securiform and rather small; the
femora abruptly clavate.
98. SUtira viridltasdata, n. sp.
EUongata, narrow, shining ; testaceous, the eyta black, the elytra
with two broad, transverse, metallic green fasciae— one baaal, not
induding the humeri, the other post'inedian, the two connected
along the suture, the second extending a short distance down the
first interstice poeteriorly j the elytra with a few long, erect, bristly
hairs towards the apex. Head almost smooth, with a shallow,
transverse, sparsely punctate fovea between the eyes, the latter very
lai^; antennae long and slender, joint 11 as long as 7-10 united.
Prothorai longer than broad, oblongo-cordate, strongly oonstriolcd
before the dilated base ; closely, finely punctate. Elytra long, paralM
l„y,l,Ajh,COOl^lL'
Nae and little-ibunm Lagriidae. 243
their basal half; oloselj, finely, deeply punotato-Btriate, the
interatioee convex beyond the middle, 3 with four deep setigeroiu
lesions scattered along the apical half, 1 with one and 9 with
two similar impressions near the tip. Legs long; anterior femora
strongly cIsTate ; anterior tibiae ronnded estemally at the base.
Length 9}, breadth 2} mm. (^ T)
Sab. Uffeb Auazons, Ega (H. W. Bates).
One specimen. A close ally of S. calophaenoides, infra,
with the broad elytral faaciae green and connected along
the sutoie, the setigeious impressions deep and veiy few in
number, foui only being present on the third interstice.
The elytra are paiallel, not widened towards the middle.
S. vindinataUt, Pic (Melanges ezot.-entom. iv, p. 15), from
Cumbase, Peru, must be very similar; but as nothing is
said about the testaceous humeral patch, and it apparently
has more numerous setae on the elytra, the identification
cannot be accepted without an examination of the type.
99. SlftUra elegaatnls, n. sp. (Plate XIII, fig. 24, <^.)
ij. Moderately elongate, narrow, depressed, shining; testaceous,
the eyes black, the elytra, except the basal margin, an ante-median
tmoBTerse fascia (widened outwards and not reaching the suture],
and the apex, metallic green or bluish green ; the elytra with a few,
long, erect, bristly haira. Head almost smooth, with a shallow,
transTerse, punctured depression between the eyes, the latter
moderately large ; sntemtoe long and slender, joints 7-10 decreasing
in length, 11 extremely elongate, about as long as 4-10 united.
Prothorax narrower than the head, longer than broad, oblongo-
cordate, dilated at the base; sparsely, minutely punctate, in one
specimen obsoletely canaliculate on the disc anteriorly. Elytra
moderately long, subparallel in their basal half, nearly twice as
broad as the prothorax; closely, finely, deeply puncUto -striate,
the interstices convex, 3 with four or five widely scattered setigerous
impressions, and 9 with two similar impiessions at the base. Legs
l<mg, slender; anterior femora stout, clavate; anterior tibiae
obliquely narrowed externally at the base.
length fit-6l, breadth 1^2 mm.
Hob. Ahazons, Ega and Para (H. W. Bates).
Three specimens, showing no variation. Smaller and
more slender than S. vindifasciata; the elytra less elongate,
and with the green coloration more extended, leaving the
basal mai^in, fui inwardly-narrowed, abbreviated, ante-
median fascia, and a small patch at the apex testaceous ;
iM-,i,A J h, Google
244 Mr. G. C. Champion on
the apical joint of the antennae (^) extremely elongate.
The anterioT tibiae are compresBed at the base mnch as
in S. elegant, Makl., but the species seems b«8t placed next
S. viridyasdata.
100. Statira ealophaenoidei, n. sp.
Elongate, narrow, depieeaed, Bhining; tMtaceoua, the eyes and
two broad straight fasciae on the elytra (one basal, the other poit-
median) black; the elytra with scattered, long, erect, bristly hairs.
Head sparsely, minutely ponotate, ehallowly foveate betwetn the
eyes, the latter large and not very widely separated ; antennae very
long and slendw, joint 1 1 in ij nearly equalling the preceding five,
and in $ a little longer than the preceding three, joints united.
Prothorai as wide as the head in $, slightly narrower in ij, long^
than broad, cordate, dilated at the base, sparsely, minutely punctate,
obaoletely canaliculate anteriorly. Elytra moderatdy long, at the
middle twice as broad as the prothorax, gradually widened in their
basal half, flatteoed on the disc, rounded at the apex; closely,
finely crenato-striate, the interstices feebly convex towards the tip,
1, 3 and 6 with scattered setigerous impreeaions throu^iont their
length, and 7 and 9 with three or four similar impressions on the
apical half. Lega long; anterior femora stout; anterior tibiie
rounded estemally at the base.
I.eiigth ^-^, breadth 21-2f mm.
HtA. Peru {ex coU. F. Bates).
Two examples, assumed to be sexes. This insect
superficially resembles a small Calophaena (a Tropical
American genus of Carabidae], some of the species of which
are similarly nigro-bifasciate. It is of about the same aee
and shape as S. elegans, Makl,, the latter having metallic
green elytra, etc.
101. Statin Hgregata, n. sp.
3. Extremely like S. caiophaewndM, and similarly coloured,
except that the transverse black basal fascia on the elytra is broader
than the submedian one; antennae very long and slender, joint II
nearly as long as 6-10 united; the prothorax more narrowed
anteriorly ; the setigerous impressions on the eljrtra fewer in number,
reduced to six along interstice 3, and two or three only towards
the apex of 1, C, 7, and 9.
Length %^, breadth 2} mm.
H(A. Upper Ahazoks, Ega (H. W. BaUs).
u.,.w,,t; Google
New attd liOlfr-inown Lagriidae. 245
One male, sufficiently difierent from ;S. cahphaenoidea
to leqnite a diBtiiictive specific oi varietal name.
102. Stelira uminign, n. sp. (Plate XIII, &g. 25.)
Elongate, narrow, shining ; mfo-teetaceouB, the eyee and the
apical half of the elytra black ; the elytra with a few, long, erect,
briatly hairs. Head almost amooth, shallowly foveate in the iniddle
between the eyee, the latter large and moderately distant ; antennae
long and slender, joint 11 as long as 8-10 united. Prothorax aa
wide as the head, not longer than broad, cordate, deeply oonstricted
before the base, the basal margin much raJEed and laterally project-
ing; almost smooth, obeoletely canaliculate at the apen. Elytra
long, subparallel in their basal half; closely, finely puncta to -striate,
the interstice 3 with aii, and 5 and 9 each with three or four, widely
scattered setigerous impieesione, 1 also with two impiessions at the
apex. Femora more or leas clavate.
L^igth 7}, breadth 2i mm. ($t)
Hob. Upper Auazohs, San Paulo [de Olivenca] (H. W.
Bales).
One specimen. Differs from the allied S. adophaetwides
in having the apical half only of the elytra black, and the
bead and prothoraz shorter and broader, the latter almost
smooth, the elytra with scattered setigerous impressions on
the interstices 3^ 5, and 9.
103. Statira Irlsdlata, a. sp. (Plate XIII, fig. 26, <$.)
S- Very elongate, shining; rufo-teetaceoua, the eyea and three
oblique, angulate fasciae on the elytra (one before and one just
beyond the middle, neither reaching the suture, and one, narrower,
Bnbapical, the last-named continued down the first interstice and
oater margin to near the tip) black ; the elytra with a few, long,
erect, bristly hairs. Head almost smooth, longitudinally grooved
betwetm the eyee, the latter lai^e, moderately distant; antennae
slender, long, joints S-10 decreasing in length, 11 extremely elongate,
as long as 6-10 united. Prothoraz as wide as the head, a little
broader than long, subcordate, dilated at the base; sparsely, finely
punctate at the base and with two deep widely separated f oveae on the
diso poetoriotly. Elytra somewhat convex, very elongate, twice aa
brood as'the prothorax, gradually widened to the middle, acuminate
at the tip: closely, finely crenato-striate from base to apex, the
interstices convex, 3, 5, and 7 with scattered setigerous impreesions
iM:,i,A J 1^, Google
246 Mr. G. C. Champion on
throughout their length, I and 9 also with several aimilar impteB-
sions OD the apical half. Legs veiy long, i«ther alaider.
Var. The upper Burfaee uniformly oastaneoos.
Length HH^. breadth 3)-3i °>°>-
H<A. Colombia, Ibague {Mus. Brit.).
Two specimens, acquiied in 1846. A very elongate,
somewhat convex, shining, rufo-testaceous or castaneouB
insect, with sharply, obliquely, nigro-trisellate elytra
in the form selected as the type, the elytrsl striae
lather deeply impressed and crenato-punctate, the alter-
nate interstices with scattered setigerous impressions. The
two {oveae near the base of the prothorax cannot be acci-
dental. The aedeagus (partly exposed in the type) is
narrowed to the tip, and a pair of narrow carved claspers
are visible beneath the fifth ventral segment. S. triadlala
cannot be identified amongst the 28 Colombian SUUirae
described by Maklin in 1878; it is coloured somewhat as
in Colparthrum gerstackeri, Kirsch. The immaculate form
must come near his S.femtginea : the latter is said to have
a somewhat densely punctured prothorax and the elytral
striae evanescent towards the apex.
104. Statlra puaensis, n. sp.
Moderately elongate, narrow, the head and prothorax subopaque,
the elytra shimng; t«etBceouB, the eyes black, the elytra with a
narrow, nlgro-fuscouB, traneverse, undulate, poet-median fascia —
sometimee extending narrowly forwards along the suture and dilated
anteriorly into a faint triangular scutellar patch, sometimes not
reaching the suture or altogether wanting; the elytra with a few
very long, erect, bristly hairs. Head almost smooth, longitudinally
impressed between the eyes, the latter large and somewhat widely
separated; antennae long, slender, joint 11 in <} aboat equalling
8-10, and in 9 ^ and 10, united. Prothorax as long as broad,
nearly or quite as wide as the head, coidate, dilated at the base;
ajmoHt impunctate. Elytra moderately long, gradually widened to
the middle; very Gnely, oloeely punctato^tiiate, smoother at the
apex, the interstices flat, 3 with about seven conspicuous setigetous
impressions scattered along ita entire length, 6 and 9 with three
or four similar impressions on the apical half, and 1 with one or two
near the tip. Aedeagus of j strongly acuminate at the tip.
Length 6i-6, breadth li-2i mm. (^9-)
Hob. Lower Amazons, Para (H. W. Bates).
iM-,i,A J h, Google
New and litite-kiwwn LagnOae. 247
Five BpeciineD — ^three fas<^ate, two immaculate. A
fono of tne Central American 8. picla, Champ., with the
apical joint of the antennae shorter in both sexes (ju S.
picta, ^, equalling 7-10 united), the head with a deeper
longitiidinal groove between the eyes, and the elytral
markings, when present, showing a tendency to extend
forward along the suture to the scutellar region (instead of
along the outer margin to the humeri, as in 5. picta), the
scattered setigerous impressions along the third interstice
very conspicuous.
105. SUtfnt einnmduflta, n. sp.
3. Moderately elongate, depi«aeed, Bhining; nifo-testaoeooa,
tlie eyes black, the elytia with the suture narrowly, and a common,
transTBTse, curved median fascia, which is continued forwards along
the outer part of the disc to tlie basB, nigro-fuscous ; the elytra with
» few long erect setae. Head broad, spaisel;, minutely punctate,
grooved down the middle betweco the eyes, the latt«r large and
separated by about the width of one of them ; antennae moderately
long, rather Blender, joint II slightJy longer than S-10 united. Pro-
thorax narrower than the head, about as long as broad, auboordate,
dilated^t the baBe ; almost smooth, faintly canaliculate down the
middle anteriorly. Elytra twice as broad as the prothorax, moder-
ately long, gradually widened to the middle, obliquely depressed
below the base ; olosdy, finely punotato-striate, the interetioea 3, 6
and 9 each with about six widely separated setigerous impicsBions,
and 1 and 7 with one or two similar impreesions near the apex.
Length 8, breadth 2) nun.
Hob. Ecuador (Buckley).
f One male. Very like 8. vageguUata, Pic, and its allies,
but wanting the deep lanciform sulcus on the front of the
prothorax, the dark markings on the anterior half of the
elytra partly enclosing a broad, elongate, subquadrate
patch of the testaceous ground-colour on the disc of each
of them.
106. Stattra snUentstrata, n. sp.
Moderately elongate, narrow, shining, testaceous ; the eyes
black, the elytra with a broad basal and a narrow median fascia,
connected along the suture and approaching one another at the sides,
nigro-piceous ; the elytra with a few long, erect, bristly hairs. Head
laige, broad, obliquely narrowed and well developed behind the
eyes, sparsely, obsoletely punctate, the eyes smell, widely eepaislf d ;
u,y,i,Ajh,Cooglc
248 Mi. G. G. Champion on
aatoimae very slender, moderately long, joint 1 1 barely the lengtii of
9 and 10 united. Prothoras narrow, longer tlum broad, coidate,
widened at the base, apaisely, finely punctate. Elytra moderately
riongate, twice as broad as the prothorax, gradually widened to the
middlp, acuminate posteriorly; ahallowly, minately striato-punc-
tate, smooth at the apex, the inteistices flat, 3 and 5 with four or five
widely scattered oonspionous aetigerous impressions, 7 and 9 also
with two or three similar impressions towards the tip.
Lengths}, breadth 1{ mm. ($T)
Hab. AuAZONS, Santarem (H. W. Baiet).
One specimen, imperfect. A very narrow, testaceous
insect appioacIuDg S. vageguUala, Pic, with the elytra
somewhat similarly maculate ; the head large, the eyes small,
the antennae miusually slender, and with a feebly developed
apical joint, the prothorax qmte narrow, punctulate, and
without sulcus in front, the seriate elytral ptmctures very
fine and shallow. The piothorax and elytra aie differently
shaped from those of 8. paraensis.
107. Statir» multino'tata. (Plate XIII, fig. 27, ?.)
Suuira muUinobUa, Pic, Melanges exot.-entom. iv, p. 15
(Sept. 1912).
Ant«mal joint 11 in (^ as long as 7-10, in $ as long as 8-10,
Hab. Bkazil (coU. Pic : type), Minas Geraea (Mus.
Brit.).
Three examples in the Museum, two of which ate from
Minas Geraes, are doubtless referable to this species. A
narrow, elongate insect, 8^9 mm. in length (Pic gives
10 mm.), rufo-testaceous in colour, the elytra flavous, with
the numerous scattered setigerous impressions along the
alternate interstices each placed in a sharply-defined,
subquadrate, blackish or fuscous spot, those on the fiwt
(sutural) interstice confined to the apical half. The eyes
ate large and somewhat narrowly sepatated, the space
between them depressed down the middle.
108. Statira vagegutUta. (Plate XIII, fig. 28, S.)
Stalira vagegiUtaia, Pic, L'Echange, xxviii, p. 75 (Oct. 1912).
Statira vagenolata, Pic, loo. cit. p. 76,
l„y,l,Ajh,COOl^lc
New and litde-lmmon Lagriidae. 249
Var. Piot^orai with a dbttow bUutk vitU on each Bide near the
outer maigiti. Joint 11 of uitamafl in (^ as long aa four or five
of the piecedmg jointa united, in $ about equalling 8-10 united.
Length 5-8}, breadth l|~2f nun. 1,$^)
Bab. Brazil {Mus. Brit., M\ts. Oxon.), Rio de Janeiro
{Fry), Constancia (J. Gray and H. Clark, Jan. 1857),
MatUBinhos [type of 5. rojeywUata] and Sens de Bemarda
[type of S. vagenofata] {cdl. Pic) ; Amazons {H. W. Bates,
ex cdl. Fry).
This seems to be one of the commonest species of the
genus in the neighbouthood of Bio de Janeiro, but it was
apparently unknown to Maklin. The long series before
me, including many specimens belonging to the Oxford
Museum, illustrate the great variation in the development of
the elytra! maridngs, the additional variety here noted (eight
specimens seen) simply differing in having a submarginal
black line on each side of the prothorax. The commonest
form has three dark angulate fasciae on the elytra — one
just below the baae, another at the middle, these connected
along the sntnral and outer margins, and a narrower one
towards the apex, this latter connected with the median
fascia along the suture and outer margin and down the
middle of the disc (the fuscous markings thus enclosing
on each elytron a la^e ante-median discoidal patch, and
two transversely placed oblong marks behind it, of the
yellowiah gronnd-coloot) ; the dark coloration sometimes
eztends over the whole of the apical portion, or it may be
almost entirely or in great part obHterated. S. vageguUata is
Teadily recognisable by the deep, sharply-defined, elongate-
triangular sulcus on the disc of the prothorax in front;
the prothorax iUe\i ia sparsely, very finely punctate, and
usually shining, sometimes opaque. The elytra are finely
ponctato-striate, the interstices 3, 5, 7, and 9 each with from
two to four, and 1 with one or two, scattered setigerous
impressions on the apical half. There is a specimen before
nie from the Dejean collection labelled with the MS. name
S. pkta, Buq. [nee Champ.] and one from the Oxford Museum
is ticketed 8. varians, Hope. The general resemblance of
this insect to a spotted Drmnitis is rather striking.
109. 8taflralnd)leollU,n.Bp. (Plate XIII, figs. 29, antenna;
29a, prothorax, {^.)
(J. Moderately elongat«, narrow, depressed, shining; rufo- or
fiaro-testaoeouB, the lltb anl^awal joint slightly infuscate, the eyes,
iM-,i,A J h, Google
250 Mr. G. C. Champion on
the elytra to a little beyond the middle, esoept around the soutellom
and on an obljqae or subtriangular space on the disc, and in one
speoimm a nanov, tnuuverse oblique mark on Ihe disc before the
apex, black ; the elytra with several erect bristly hairs towaida the
tip. Head broad, sparBcIy, minutely panctat«, obeoletely canalicu-
late between the eyee, the latter Urge, separated by about tjie
width of one of tbem; antennae (fig. 29) elntder, moderately long,
jointe 3-10 rapidly decreasing in length, 7-10 moniliform, II es-
tremely elongate, about as long as 3-10 united. Prothoras (fig. 29a)
narrower tlisn the head, about as long as broad, cordate, dilated
at thebaMi sparsely, minutely punctate, and with a deep lancif onn
BulouH on the disc in front. Elytra moderately long, gradually
widened to the middle; closely, finely punctato-striate, the iuter-
Bticea broad, feebly convex at the apex, 3, G, and 9 with threeor four
scattered setigerous impreaBions on the posterior Iialf, 1 also with a
single impression near the tip.
Var. The prothorax broadly inf usoate along the sides, the blackish
portion of the elytra more extended, reaching the subapioal
mark. (tS.)
Length 6)-6- breadth 11~2 mm.
HrA. Lower Auazons, Santarem {H. W. Bates : type);
Brazil, Pemambnco {Fry : var.).
Described from three specimens, one of those from
Santarem now wantmg the antennae. A very close ally
of S. vagegvUata, Pic, with the apical joint of the ^ antenna
nearly as long as the rest united, and the joints preceding
it moniliform. There is also a corresponduig vanety with
a laterally fusco-vittate prothorax. It ia just possible that
these insects may be forms of S. brasUiemia, ^c (Melanges
exot.-entom. iv, p. 19), which is said to have a long terminal
{'oint to the antennae ; but without comparison, it would
le unsafe to identify them as such.
110. Slatlra xanlhodera, n. sp.
Moderately elongate, shining, the elytra sericeous; black, the
prothorax, the femora at the base, and the tarsal claws testaceous ;
the elytra with a few long, erect, bristly hairs. Head almost smooth,
well -developed behind the eyee, the latter small and widely separated ;
antennae short, not very slender, joint 11 equalling S-10 united.
Prothorax as wide as the bead, transverse, subcordate, dilated at the
base i smooth, with a very deep, sharply defined, oblong excaT&tion
on the disc in front. Elytra moderately long, about twice as broad
as tlie prothorax, gradually widened to the middle, rounded at the
ih,Cooi^lc
New and little-known Lagriidae. 2S1
Rpex; olo«riy, minutelj, very shallowly pimota to -striate, the inter-
stices flat, 3 with five, and S, 7, and 9 with two or three, scattered
setigerouB impreuiona on the apical half, 1 ^bo with a single impies-
aion near the tip.
Length 6, bnadth 2^ nun. ($ T)
Hab. Brazil, Sasta Cathaiina (Fry).
One specimen. This is one of the veiy few species of the
genus known to me with a deep, oblong, buIcob on the disc
of the prothoisx in front. This character brings it near
S. vageguttata, Pic, from which it differs greatly in colour,
and in having short antennae, araaller eyes, and very faintly
striate elytra. The wholly black body and testaceous
prothoraz, too, ate characteristic.
HI. SUtfn flfforato. (Plate XIII, fig. 30, $.)
Statirafyurata, Makl., Act. Soc. Fenn. vii, p. 158 (1862).
IStaiiTa lunidala. Pic, Melanges exot.-entom. iv, p. 18
(Sept. 1912).
Elongate, rather broad, very abioing; nifo-, the elytra flavo-
teataceoiu, the eyee, the prothorax with a submarginal line on each
side, and theelytrawithanaTTow,anguIatep08t'baBalaada broader
carved median fascia — these markings connected along the suture
and outer margin (the eutural stripe extending don-nwards to near
the apex), and the post-basal fascia with a curved ramus extending
forwards along the middle of the disc to the scutellum — black;
the elytra and the sides of the head thickly set with rery long, erect,
eetiform hairs, the l^s, antennae, and under surface also with
(easily abraded) hairs. Head broad, finely punctate, slightly
impressed in the middle between the eyes, the latter separated by
about the width ol one of them; antennae with joint 11 about as
long as three ($) or four of the preceding joints united. Prothoras
a little narrower than the head, as broad as I<Hig, subcordate, dilated
at the base; rather closely, finely, conspicuously punctate. Elytra
twice as broad as the prothorax, long, gradually widened to the
middle ; closdy, finely punotato-striate, the interaticee each with an
irregular row of rather coarse, somewhat closely placed setigerous
impressions extending from the base to the apex.
Length l\-9, breadth 2)-3 mm.
Hab. Brazil, Rio de Janeiro {Fry), Constancia (J. Gray,
Jan. 1857).
The two specimens here described, possibly (J and ?,
agree with Makhn's description of S.fyvrata, from " Brazil,"
iM-,i,A J h, Google
252 Mr. G. C. Champicin on
except in having the apical portion of the elytra flavo-
teetaceous, instf^ of black; but as some of the allied
forma vary in this way, no importance need be attached
to Buch a colour-difference, the five yellow spots on the
basal half being exactly as be describes. S. lunulala, Pic,
from Tijaca (a place near Constancia), which is compared
with my figure of S. coTispicillaia, Makl., in the " Biologia,"
is said to have an irregular black ring enclosing a yellow
patch on the anterior portion of each elytron. 'Die specific
identity of these insects, therefore, must remain in doubt
till types are compared.
112. Staiira annuUta. (Plate XIII, fig. 31.)
Stalira annulata, Makl., Act. Soc. Fenn. x, p. 6i3 (1875).
Elongate, rather nurow, shining; testaceous, the eyes black, the
elytra flavous, with three common, angulate fasciae, connected along
the suture, the first two along the sides also, and the second and third
nearly coalescent along the fifth interstice, the first with a brancli
on the disc extending forwards to the base, nigro-fuscous (the dark
markingB enclosing nine spots, ananged 3, 2, i, of the yellowish
ground -col out] ; thickly clothed, the legs included, with long, soft,
erect hairs. Head sparsely, rather coarsely punctate, the eyes
somewhat narrowly separated ; antennae moderately long and
slmder, joint 1 1 as long as 9 and 10 united. Prothorax longer than
broad, oblong-subcordate, a little dilated at the base ; very coarsely,
sparsely punctate. Elytra long, about twice as wideas the prothoras,
gradually widened to the middle; closely, finely punctato-striate,
the interstices each with on irregular series of closely placed pili-
gerous punctures.
Length 7}-8|. breadth 2^2| mm.
Hah. Brazil, Rio de Janeiro and Espirito Santo (J^ry).
The two specimens, probably ^ and 9> from which the
above description has been taken agree with Maklin's
diagnosis of his S. annulata, from Brazil, except that they
apparently have relatively narrower elytra. The differences
given by bim to separate S. annuUOa and S. jigttrata (as
here recognised) also apply, so that there cannot be much
doubt about the identification of either of them. A fresh
description, however, was required in each case.
113. Statlra quadriplaglata, n. ep. (Plate XIII, fig. 32, ^J.)
1^. Moderately elongate, narrow, shining j testaceous, the eyes
black, the elytra flavous, with a broad, common, triangular soutellar
iVcw and liUle-lmown Lagriidae. 253
pateh (reaching the humeri), a large triangular patch on the onter
pttrt of the disc jiut beyond the middle (extending inwards to the
second stria), and a common apical patch, nJgio-hucouB ; thickly
clothed, the Ic^ included, with long, soft, erect hairs. Head rather
small, coarsely punctate, the eyee large and somewhat narrowly
separated; antennae comparatively short, rather etout, joint II aa
long as S-10 united. Prothorax convex, about sa long aa broad,
somewhat oval , scarcely dilated at the base ; coarsely, rather closely
punctata. Elytra about twice as wide aa the protboras, long,
gradually widened to the middle; closely, finely punctato-striate,
the interstices each with an irregular series of rather closely placed
piligerous poncturee extending from the base to the apex. Aedeagus
slender and acuminate at the tip.
Length 61, breadth 2^ mm.
Hob. Brazil, Rio de Janeiro {Fry).
One male. Closely related to S. annulata, Makl., but
with very differeotiy marked elytra (resembling <S. scutd'
larig. Pic, S. evanescena. Champ., etc., in thia respect),
the ant«miae less elongate, the prothorax not so long, and
still more coarsely punctured. The piligerous punctures
on each elytral interstice are closely placed and extend
from the base to the apex, as in iS. annulata and S.jigurala.
114. Statlra blfureata, n. sp.
$. Elongate, narrow, opaque, the elytra moderately shining;
tMtaoeouB, the eyes black, the elytra with a narrow sutural stripe,
which is obliquely bifurcate before the base and nearly reaches the
apex, and three longitudinal streaks at about the middle (on inter-
atioes 3, 6, and 7, that on 7 longer than the others), nigro-piceous;
the elytra with scattered long, erect, bristly haira. Head slutaceous,
closely, roughly punctulate, the eyes large and separated by about
half the width of one of them; antennae rather slender, joint 11
neariy as long aa S-10 united. Prothorax longer than brosd, as wide
as &e head, cordate, dilated at the base ; closely, very finely scabroso-
punctalate, the basal groove almost obsolete. Elytra long, at the
middle twioe as wide as the prothorax, rapidly narrowed and acumi-
nate posteriorly; closely, very finely punctato -striate, the alternate
interstices 1, 3, 5, 7, and 9 with a series of somewhat closely placed
setigerous impressions extending throughout their length, the
iropreesions each preceded by a small tubercle.
Length 6^. breadth 2 mm.
ffoi. Brazil, Jatahy, Province of Goyas {Pujd, ex coll.
Fry).
iM:,i,A J 1^, Google
254 Mr. G. C. Champion on
One female. Not unlike S. ^uadripkiffiata, the head
and prothoraz opaque and scabroBO-punctulate, the
piothotax longer and with shallower basal groove, the
elytra very differently marked and with more nimierouB
tubercles along the alternate interstices. Pic has described
various SUUirae from the same Province, but the present
insect is not included amongst them': iS.^oyasenm*" (length
9 nam.) also has the elytra testaceous, with a black sutural
stripe enlarged towards the acutellum, but the other
particulars given do not accord with S. infuTcata, and the
tuberolea are not mentioned.
115. Statira tennipes, n. sp.
iS, Elongate, narrow, depiessed, alender, opaque above, ahining
beo«atli; testaceouB, the eyes block, the elytra with the sutuie
narrowly pioeoiu to near the apex, the latter with a few bristly hain.
Haid broad, alutaoeoua, obsoletely oan^iculate betwerat the eyes
and obliquely narrowed behind them, the eyes laige, somewhat
distant ; antennae moderately long, slender, joint 1 1 nearly equalling
7-10 united. Prothoraz much longer than broad, oonaidenbly
narrower than the head, rounded at the sides, the latter deeply
sinuate before the base, the basal margin not raised, the surfaoe
alutaoeous. EUytra elongate, at the middle about twice as wide as
the prothotaz somewhat acuminate poeteriorly ; closely, very finely,
ahatlowly punctato-atriate, the inteisticcs feebly convex, 3 and S
with about six widely scattered setigerouB impreeeions, I also with
three others near the tip, and fl with five impressioni bey<»d the
middle. Legs long and slender.
Length ?{, breadth 2} mm.
Hab. Brazil, Bahia (Reed).
One specimen. This b one of several pecuhar forms
obtained by Reed at Bahia. It resembles S. bifurcata,
from Govas, but has a much broader head, a longer and
smoother prothor&x, non-tuberculat« elytra, with the
autur«> only infuscate. and longer legs.
no. Statin eniaiata, n. sp.
KklnplI•^, niiher broad, mxletatety ahining; teetaceoos, the
lifsid and piMtliorax mMi>h. th* eyes blarlc; the elytra with an
ill-ilrtiiKil. wintmm. ourrrd piet-nwdian faaria, extending forwards
'• M<^«>,w Mot.-nju.m. iv. p. 20 (S^t- 1912).
u,y,l,/eJh,G00glc
New and lioie-knoum Lagriidae. 255
aJoQg the oater margiD to the humeral calhie, a&d th« imee in-
determinately, infiiBcato, the eurfoce bearing a few bristly haira.
Head small, narrower thui the prothonz, closely, finely, con-
Epicuoasly ponctate, unimpiesaBd between the eyes, the latter
moderately large and somewhat distant ; antennae slender, short,
. joint 11 equalling 7-10 united. Prothorax oblongo-eubquadrole,
distinctly margined laterally, slightly hollowed at the sidee anteriorly,
and sinuously narrowed towards the base, the anterior angles
obtuse, prominent, the hind angles very slightly projecting, the
basal margin but little raised, the transTeree groove in front of it
deeply impressed laterally; deosdy, minutely punctate. Elytra
widening to the middle and theif twice as wide aa the prothorax;
finely, closely, deeply punctato-striate, the interstices convex, 3
and 5 with four or five widely scattered setigeious impreasicms,
and land 9 with two or three othersnear the apex. Legs moderately
long-
Length 9, breadth 3 mm. ($ ?)
Hab. NiCARAGDA, Chontales [Janeon).
One specimen, differing from all the Central American
forms enumerated by myself in the " Biolo^ " in 1889,
It is perhaps nearest related to the Mexican S. i«mecr«m,
from which it is separable by the less rugnse, shining head,
the longer, lai^r, and less scabrous prothorax, and the
conmioti, ili-defined, fuscous post-median fascia of the
elj^tra.
117. Statira haltlflnsis, n. sp.
Elongate, rather dull ; nigro-piceous or black, the prolkoiax and
tarsi rufous or rufo -testaceous, the elytra nigro-cyaneous, the
antmnae Bometimee wholly or in part ferruginous ; the elytra with
a few fine, long, erect hairs. Head rather short, narrower than the
prothorax, sparsely, minutely punctate, the eyee large, somewhat
distant; antennae slender, moderately long, joint 11 equalling the
three or four^receding joints united. Prothorax transverse, faintly
margined laterally, rounded at the sides, constricted faefoie the
prominent hind angles, the basal margin raised and the tianBTerse
sulcus in front of it sharply defined; sparsely, minutely punctate.
Elytra long, gradually widened to the middle, and there twice as
«ide as the prothorax; finely, clocely, deeply punctato -striate,
the interstices more or lees convex, 3 with nine or ten, ar^ 5 and 9
with several, sotigerous punctutee scattered between the base and
apex. L^s moderately long.
Length 6i-7i- breadth 2j(-2f mm. {?.)
u,y,i,A"jM,Googlc
25^ Mr. G. C. Champion on
Hab. Haiti, Port au Prince (ex coll. Fry).
Four examples, possibly all females. This species
bears some resemblance to the Antillean S. vittala, Cttamp.,
but it is very differently colonred, and has a shorter head
and prothorax, and the latter finely margined at the sides.
Also approaching S. croceiccHis, Makl. (from Florida,
Georgia, and Alabama), and other N.-American forms.
118. Stallra falfleoUts.
Anthiciis ftdvicdlis, Fabr., Syat. Eleuth. i, p. 290 {1801).i
SlalirafulvicoUis. Makl., Act.jSoc. Fenn. vii, p. 160 (1862).»
Vor. a. Elytra each with a reddiab or tcetaoMiuB apot below the
SlalirafulvicoUis, v. discoidaiis, Pic, Melanges exot.-entom,
iv, p. 13{1912).»
Var. $. Prottaorai black, the elytra eometimee fueoous or pioeous.
Anihicua fusci^cnnis, Fabr., loc. cit.*
S(at\ra ftiscipmnis, Makl., Act. Soc. Fenn. vii, p. 160.*
Sifttira exigua. Makl., Ofv. Fineka Vet.-Soc. Porh. xx,
p. 360 (1878).'
Var. y. Prothonz black; the elytra and abdomen Ustaoeona
at the base, or the elytra (a triangular scutellar patch and the auton
excepted) teetaoeous in their baaal h&U.
Anihiai.i aMominatis. Fabr., loc. cit.'
Statira abdominftlis. Makl., Act. Soc, Fenn. vn, p. 160.'
Var. 3. Prothorax black, the elytra with an oblong yellowiah
mark on the disc before the middle.
Staimi atricotlis. Pic, M4lan<.:es exot.-entom. iv, p. 20
(St'pt. 1912).*
Var. •. Prothorax black, the elytra «Mih with a ydlow atripe on
the disc extending from the base to b^md the middle.
Stitlini f?.itv..'f.i/.i. Pic. M^lances exot.-entooi. xi. p. 18
tXov. nil4>."
rV»r. i. IVxIiorsi. a brD«d ol>tt>n^ stripe od the dkc of eadi
el.vl)vn tciintilitic fn'm a litile below the base to beyood the middle
aitd «luK*.t rtwiiiitw the suture), under sDrfaw. femora, and tibiie
liW anterior (Mir eii-«|>ted> tevtaeevus or ttavoos.
Ij.y.l.Ajl^yCOO'^lL'
Hoe and Uole-biOKn Lagriidae. 267
Hab. South Ahebica u*>^; Colombia *, Bc^ta;
GuUKA, Cayeime ' ; Brazil >° (Miat, in Mia. Oxtm. : -m.
JlavoviOata), Rio de Janeiio, EsiHiito Santo, Babia, Per-
nambaco [var. C]', Amazons, Pais, Kanta [var. a];
Bolivia*.
The muDerons specimens before me oomiect the Daic«d
fonna of this veiy variable Stalira, and it is not impossible
that S. {AtUhicus) ruficoOis, Fabr., which has a red head
and piothorax, appertains to the same species. Makhn
did not identify any of these Fabrician insecta amongst the
material examined by him ; but there can be little doabt
that the three named quoted above refer to one and the
same species, that with a red or yellow prothorax being
apparently confined to Brazil and Guiana. An example of
the var. a (= discoidaiis. Pic) from the F. Bates collec-
tion (from that of Dejean) is labelled with the MS. name
Pedtlus sanguinicoUis." The allied S. irregularis and iS.
aibofasciaia. Champ., from Central America, are almost
eqnally variable in colour. iS./uivtcoJJts is a small, polished,
Aiithiciform insect, wholly or in part black; the elytra
feebly striato-punctate, with an irregular series of numerous
larger, rough, setigerous impressions along each of the
intetsticea; the antennae gradually thickened outwards,
the apical joint about as long as the two preceding joints
united in both (S and $; the head transverse, foveate in
the middle between the eyes, the tatter small ; the tibiae
somewhat curved in <^. There is a broken specimen of the
var. flavovUtata in the Oxford Museom. These forms,
shoold perhaps be removed from the genus Statira.
119. StaUra oyanipennis.
Statira cyanipennis, Mfikl., Ofv. Finska Vet.-Soc. FdrB.
XX, p. 36 (1878) (nee S. cyanipennis, Matd., Act. Soc.
Fenn. vii, p. 591, and x, p. 647).
Statira colombica. Champ., Biol. Centr.-Am., Coleopt.
iv, 2, p. 62, nota (nom. prov.).
Bab. Colombia [type], Bogota (ex coll. F. Baiea).
Two specimens from the F. Bates collection agree with
Uaklin's description, except that they have the abdomen
black, instead of dilute fulvous. They are separable from
typical S. ftdvic(Mis, F., by the prothorax being more
>' C/. B. C-Am., Coleopt. iv, 2, p. 46.
tbams. ent. soc. lond. 1917. — part I. (NOV.) s
iM-,i,A J h, Google
?68 Mr. 6. C. ChampioD on
dilated ftat«rioriy (thus appearing nairowei at the base)
and a little shorter, and tne elytra brilliant cyaneous in
-colour. The first species described by Maklin under the
name S. cyanipetmis (1862) was selected in 1889 as the
type of Sphrafftdcphonu.
120. Staflra eajrsiuMUls, n. sp.
Moderately elongate, somewhat robuat, widened posteriorly, very
■hining; testaceous, the head and palpi, the antennal jointa t and
-4-10. in part or entirely, nearly the apical half of the elytra (a spot
on the outer margin near the tip excepted), and the abdomen in
part, blank, the tibiae and tarsi nigro-piceous ; the elytra and legs
somewhat thickly, and the head sparsely, clothed with long, fine, erect
OT projecting, pallid haiis. Head short, barely as wide as the pro-
thotax, well developed behind the eyes, veiy spatsdy punctulate;
eyes small, widely separated; antennae moderately long, gradnally
thickened outwards, joint 11 equalling 9 and 10 united. Piothorex
almost smooth, scarcely as long as broad, tranaverEely coidate,
strongly constricted before the base, the basal margin raised and
thickened, preceded by a deep groove. Elytra moderately long,
widening to the middle, and there twice as wide aa the prothorax,
somewhat acuminate posteriorly; finely, shallowly, confusedly
punctate, the seriately-arranged punctures barely distinguishable
from those of the interstices. Legs rather stout, the femora clavate.
Length «, breadth 2 nun.
Hab. Guiana, Cayenne {Miis. Brit.).
One specimen ( ? sex), injured by pinning, acquired in
1858. An insect reaembling S. fncol-or. Champ,, from
Panama, except that the head is black, but with the an-
tennae (less the flavoua apical joint) and elytra formed
much as in the Central American S. albofasciata, Champ.
S.fulvicollis, ¥., is also an allied insect. The Peruvian S.
nigroapicoXis, Pic, compared with S. Incolor by its describer,
must be a very different species, simply resembling the latter
in colour.
121. StaUra semlTlolaoea, n. sp.
moderately elongate, robust, very shining; mio-testaeeous, the
vpical half of the elytra and the sutuie narrowly thoioe to tlie
base, Dupreo- violaceous, the eyee, joints 2-7 of the antennae, the
metastenium, abdomen, tibiae, and tarsi black or piceous; the
elytra with a few bristly hairs. Head small, much narrower than
l„y,|,AJI^,COOl^lC
New and l^tU-knovn Lagriidae. 259
the prothorftx, apandy, fiaely pnnctete, foTMte on eaoh •id*'
betwe«n tbe «7ea, and nftnowing behind thno; eyta tmuvuM,
aawU, dist*nt,feeb);ooDTes.iK>trMu;hii)g the bueof the head; an-
tctmae rather short. modeTfttd; stout [joint 1 1 niissiiig). Prothoias
smooth, broader than long, arcuAtely dilated At the sidee, n«iTOw«d
and constricted at the base, the basal margin preceded hj a deep
groove; the disc with a veiy lai^, deep, ronnded excavation on
each side just behind the middle. Elytra modeTstely long, at the
middle about twice as wide as the protborax, conjointly rounded at
the tip; closely, finely, deeply punctato -striate, the interstices
convex, flatter on the disc, 3 with tour setigerous impreesions
scattered between the base and apex, 5 and 9 also with two or three
widely separated impreasions. Legs rather short and stout, the
aJiterior femora strongly clavate.
Length 6}, breadth 2} mm. l^.)
Bab. Upper Amazons, San Paulo [de Olivenca] {H. W.
Bates).
One specimen. Near S. biador. Champ., from Panama,
the protborax more dilated at the sides and with a very
large, deep, rounded excavation on each side of the disc
( ? accidental) ; the elytra conjointly rounded at the apex,
sharply punctato-striate, and unth the apical half and
suture cupreo-violaceous, the setigerous impressions fewer
in number. The apical joint of the antennae wanting in
the type, is probably elongate, at least in the male.
122. SUtlra eribrinpB, n. sp.
Elongate, narrow, widened posteriorly, shining; reddish-brown
or obscure rufo- testaceous, the femora paler at the base, more or lees
infuscate in their outer half, the eyes black; somewhat thickly
clothed with long, fine, erect, bristly hairs. Head large, broad,
well developed behind the eyes, rounded at the sides posteriorly,
closely, coarsely punctate, with a shallow, transverse, inter-ocular
impression, the frontal groove deep and almost straight; eyes
small, convex, widely separated; antennae long, slender, joint II
in both sexes about equalling 9 and 10 united. Prothorax nearly
OS wide as the head, convex, longer than broad, rounded at the sides
anteriorly, feebly sinuate and gradually narrowing towards the base,
the basal margin slightly raised ; closely, coanely punctate. Elytra
rather convex in ^, flatter in 9, moderately long, rapidly widening
to the middle, and there about twice the bre«dth of the prothorax,
atonately narrowed and somewhat acuminate posteriorly, the humeri
obtuse; Boarsely, closely striate -punctate, the punctures transveite
u,y,l,/eJh,G00glc
260 Mr. G. C. Champion on
&nd impinging on the aarrow raised interetioee. tbe allemate iatet-
stioes 1, 3, 5, 7, and 9 each with & row of somewhat oloselj placed,
fine, setigerous imprMsiona. Legs moderately long, the femon
rather stout.
Length 6-6*, breadth If-2i mm. (,J¥.)
Hah. Brazil, Espirito Santo (Fry : ,^, type), Parana
(ex coU. F. Bales : 2).
One pair. A peculiar form related to S. ajlindricdlis,
&Iakl., from Colombia, Venezuela, and Trinidad ; but
much larger and longer than that insect, with a shorter,
broader head, and long, slender antennae, the elytra with
TOWS of closely packed coaiae crenate punctures separated
by narrow raised interstices, the alternate ones each
bearing numerous long bristly hairs, the humeri less
prominent.
123. Statlra melanoptara, n. sp.
9- Elongate, narrow, widened posterioriy, shining; rufo-tes-
tooeoufi, the eyee, palpi, elytra, and legs (the bases of the femom,
and the tibiae and tarsi in part, excepted) black or piceoue, the
abdomen and basal joints of the antennae also in part infuscate;
the elytra with numeroua long, fine, bristly hairs. Head large,
broad, well developed behind the eyes, rounded at the sides pos-
teriorly, closely, rugosely punctate, the frontal gtouve deep and
almost straight; eye« email, convex, widely distant; antennae long,
slender, joint 11 equalling 9 and 10 united. Prothorax slightly
longer than hroad, narrower than the head, oonstricled before the
prominent hind angles, the basal margin moderately raised; rather
coarsely, closely punctate. Elytra long, somewhat convex, rapidly
widening to the middle, somewhat acuminate posteriorly, the humeri
obtuse; closely, deeply, rather coarsely punctato-atriate, the
punctures transverse, crenate, the interstices more or less convex,
OS wide as the striae, 1, 3, 5, 7, and 9 each with a series of numerous,
conspicuous, setigerous impressions.
Length 7, breadth 2j, mm.
Hab. Brazil (ex ct^l. F. Bates),
One specimen, A close ally of S. cribruxpa, with the
elytra and the outer halves of the femora black, the head
not so coarsely punctate, the prothorax less constricted
behind, the elytra with smaller seriate punctures and
broader inteistices, the alternate ones with conspicuous
eetigerous impressions, as large as those of the striae.
l„y,l,Ajh,COOl^lC
ATew and lHOie-known LagrHdae. 261
124. StiOn qrHnditooDis. (Plate XUI, fig. 33, 3.)
Suaira cyUndricdlis, Maki., Ofv. Fioska Vet.-Soc.
Forh. II, p. 361 (1878); F«nn., Ann. Soc. Ent., Fr.
1892, p. 97.
Hab. CoLOMBU [type]; Vbnkzuku, Caracas and San
Esteban (Simon), Cumana (ex cott. F. Bates) ; Trinidad
{F. Birci, G. E. Bryant, cofl. Fry).
A small blackish or niip-piceouB, Anthiciform insect,
with a long head and a narrow snbcylindrical prothorai,
both of which are coarsely, closely punctate ; the antennae
mfo-testaceons to about the middle, thickened and infus-
cate towards the ap<nc, joint 11 stout and about as long
a8 9and 10 united in both seies; the eyes small, prominent,
distant from the base of the head ; the elytra rather short,
comparatively broad, strongly, transversely impressed
below the base, deeply punctato-striate, the interstices
more or less convei, 3, 6, 7, and 9 with several widely
scattered setigerous impressions, 1 also with two or three
others near tne apei. The peculiarly shaped, elongate
head ia not mentioned in the descriptions of either of the
above-quoted authors. A Trinidad specimen is figured.
125. Stafira anthteoides.
St4aira anthteoides, Kirsch, Berl. Ent. Zeitschr. 1873,
p. 412.
Slatira anthteoides, v. alaudingeri. Pic, L'Bchange, ixviii,
p. 76(1912).
Hab. Peru, Chanchamayo {ex coll. F. Bates).
A close ally of S. cylindricoUis, Mnkl., with the upper
surface more shining ; the head larger and broader; the
prothorax deeply constricted before the base, and much
more finely punctate; the elytra varying in colour from
rufescent, a common, elongate, black patch at the base
of the suture excepted, to entirely black (var. staudingeri),
the interstices flatter, the striae shallow.^^
126. Stotira flUcornis, n. sp.
Moderately elongate, ateader, widened posteriorly, Bhining;
obeoure teeteoeouB, the eyes &nd elytrs black, the beaal Jointe of
tbe Bnteniue fosoo-annulate, the tarei slightly infuscate; the
•• This ia the Peruvian insoct mentioned in the " Biologia "
(Goleopt- IT, 2, p. 73) under the description of the genuB Bboaaua.
ih, Google
262 Mr. G. C. Champion on
dytn with a few bristly bain. H«ad aJmoat smootb, veiy laige,
much broader than the prothorsx, greatly derdoped bdund the
small, widely separated, convex eya, and arcuatdy narrowed behind
them, the inter-ocnlar space transvetsdy bifoveate in the middls
posteriorly; antennaeTeryeleoder, rather short (joints ^11 misaing)-
Prothorai narrow, loikger than broad, oblongo -cordate, constricted
befoie the moderately prominent basal margin; closdy, finely
punctate, smoother on the anterior half, the disc obsoletely canalico-
late anteriorly and slightly depressed in the middk at the baae.
Elytra moderately long, at the base twice as broad as the prothoni,
wideoing to the middle, areuately narrowed posteriorly, transTiTEely
flattened anteriorly, and rounded at the tip; very finely striate-
punctate to near the apes, the interstices almost flat, sJutaceous,
3 and S with a sniee of five or six widely soatteied, small setigerous
impiesaiona, 1, 7, and 9 also with two or three impreeeions neaj' the
tip. Legs slender.
Length 6, breadth 2 mm.
HiA. Peru (ex coa. F. Bates].
One specimen. The large, poateriorly developed head
and small ejes bring this species near S. antkicoida,
Kirsch, also from Pern. S.JUicomis, however, differs from
the latter in its much more slender build, the smoother
testaceous head and prothorax, the siibfiliform antennae,
the longer, duller, less convex, non-excavate elytra, and
the pallid legs.
127. SUUm perforata, n. sp.
S' Elongat«, narrow, depressed, shining, somewhat thickly
clothed with soft, fine, semi-erect hairs (now mostly abraded in (he
t3'pe}; piceous, the head and anternioe black, the elytra with a
greenish lustre. Head rather short, coarsely, confluently, rugosely
punctate, the eyes small, widely separated, depressed, the labnun
large; antennae moderately long, thickened outwards, joint 10
transverse, II stout, about equalling 7-10 united. Prothoias
uneren, wider than the head, longer than broad, feebly romided
at the sidee, the latter slightly sinuate before the base, the bassi
margin moderately prominent; very coarsely, oonfluraitly, foveo-
lato-ponctAte. Elytra barely twice the width of the prothoni.
not very elongate, rounded at the apex, depressed below the base;
closely, rather finely crenato -striate, the interstices feebly convex,
narrow, each with a row of piligerous impressions, which are of about
the same size as the transverse punctures of the striae. Lege rather
stout, the intermediate and posterior femora transv««ely strigose
l„y,l,Ajh,COOl^lC
n,g,t,7.dh,G00glc
Trans. Enl. Soc. Land.. 1917, Pi XII.
Tropical Amehican Lagriidae.
, CToo"^'-
Explanation of Plate XII.
Fto. 1(J. Slatira eoilarieenait. Champ. [Costa Rica and Colombia],
anterior 1^.
23. fiavoriffriata, a, ep. [Ecuador].
3^, „ »tituTalit, Mskl. [Brazil], anterior l<^.
4J. „ gemmifer, Mikl. [Brazil], posterior leg.
StJ ,. agroidts, Lep. et Serv. [Brazil], a«deaguB, in profile,
shoning the targe bsaal piece.
6. „ longicept, n. ep. [Brazil].
7|J. „ loagicMU, Makl. [Brazil], posterior leg.
8(J. „ meUagria, MakJ. [Brazil].
9cJ. „ calenata, Hitkl. [Brazil]; On, penia-sheath, in pro-
file; 9 b, ditto, from above.
lOtJ. „ vindi'peimU, Lep. et Serv. [Brazil], aedeagus.
showing the lai^ basal piece; 10a, ditto, in
llt^. „ nenicuiaia, Hiikl. [Brazil], a«deagus, showing the
large basal piece; llo, ditto, in profile.
12cJ. I, agg-Mmetrica, □. sp. [Brazil], penis-sheath,
\Z^. „ iortvpea, n. sp. [Brazil], posterior leg; 13a. penia-
aheath.
I4(J. „ areuatipea. Pic [Brazil]; 14a, aedeagua.
Ifi^- 11 futeilartis, Makl. [Brazil], posterior leg.
16J. „ »phtnocUra, a. sp. [Brazil],
17i}. „ curydera, n. sp. [Amazons].
18$. „ pruuttarttU, Pic [Brazil].
Ids. » *itvm», Makl. [Brazil], anterior \eg.
20(J. „ disligma, n. Bp. [Peru], prothorai.
21(J. „ ca«n<mioi(^, n. sp. [Brazil],
223. „ Slalira bn/anti. Pic [Trinidad],
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
h,Googlc
. OF Plate Xm.
Amazons).
Amazon;].
I'lliimbia].
linuill.
r.nml].
; Ilrazil], antaon*; 29a, {vo-
Colombia, Vcnezada, and'
n,g,t,7.dh,G00glc
Trans. Ent. Sac. Land., 1917, PI. Xllt.
/ V
Tkopical American Laghiid^
,L.oo^k
Explanation of Plate Xm.
Statira Bonncm, n. ep. [Amazons].
elefantiiia,n. ep. [Amazooa].
teminigra, a. sp. [AmsMiw].
IriteBata, n. sp. [Colombia].
muUinUala, Pio [Brazi)}.
vagfipUlala, Pio [Brazil].
ineiticoUit, n.Bp. [Brazil], uitenna; 29a, pro-
fiffurata. Makt. [Braul].
amulaia, M&kl. [Brazil].
qvadriplagiata, n. ap. [Brazil].
cylindHeofiH, Makl. (Colombia, VenezueiR, and.
Trinidad],
„ pUoaa, a. sp. [Brazil].
M AacfflonioulM, n. sp. [Brazil].
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
New and lUUe-hwwn Lagriidae. 263^
on tbeirinnerftuw, the tibiaeBomewhAtcarvedatthebase. AedeaguB
Btoat, Bcumiiwte and curved upwards at the tip.
Length 5J, bieodUi I J mm.
Hab. Mexico {Truqiii, ex coU. Fry).
One male. An isolated form, with the general facies
of an Arthromacra. Smaller and narrower than the
Mexican 5. brevipilis. Champ., the head and prothorax
very coarsely, con fluently punctate, the latter sub-
cylindrical, the elytra more deeply crenato-striate, with
each of the intetstices uniseriato-punctate, much as in
S. nigroaenea, Champ., from the same country. S. perforata
seems to approach the S.-American genua Statiropais,
Borcbm., which is unknown to me.
128. StaUra pUota, n. sp. (Plate XIII, fig. 34, S-)
Elongate, narrow, rather convex, ahining, thicklj clothed, the
legs included, with long, soft, erect or projecting, pallid hairs ; piceous
or ruto-piceous, with a brassy or aeneous lustre above and beneath,
tbe antaina« and legs sometimea reddish. Head short, ooanelj,
coafluently, nigosely punctate, the eyes rather email, separated in
both ae»B by about the width of one of them aa seen from above;
VitMinae rather atout, thickaied outwards, in <J about reaching
the middle of the elytra, in $ much shorter, joint II in :? a little
longer than, in 2 not equalling, 9 and 10 united. Prothorax sub-
quadrate, the sides slightly rounded anteriorly and feebly sinuate
before the base, the transverae basal groove well defined, the margin
feebly raised ; very coarsely, oonfluently, foveolat« -punctate. Elytra
long,subparalleI in their basal half, somewhat acuminate posteriortyj
ooaisely, closely crefiato-atriate, the interstices transversely rugu-
loee (except near the suture), each with an irr^ulai row of inconspicu-
ous piligeroDB pimctures, 4, 6, 8 smoother and subcostate. Ventral
aegmeDta with scattered piligeroua punctures, glabrous along the
median line. Legs moderately long, the femora clavate, the tibiae
roughly punctate. Aedeogus of ij very long, slender, acuminate,
aoeloeed in a long, narrow sheath.
Length THI.>>re(ulth2f-3i mm. (,^?.)
Hab. Brazil {Miers, in Mua. Oxon.), Rio de Janeiro
^F,y).
Four males and three females, varying greatly in size.
The description of S. t^cura, Makl., from Santa Rita,
Brazil, applies to some extent to the present insect : it
is, however, here assumed to have been taken from the
u,y,l,/eJh,G00glc
261 Mr. G. C. Champion on
female of a species of Dtsema. The following is an
extremely closely allied fonn.
129. StaUrs itrongyllold«, n. sp.
(J. Very like S. pih»a and simiUrly coloured, but more spareelf
pilose; the Mitennae more slsDder (joint II misaing); the beftd
sparsely, much more finely punctate, Bmoother between the eyes,
the latter mora approximate; the prothorax shorter, transveiBely
Bubquadnite, the very coarse, irregularly scattered punctures re-
duced in number, separate one from another, the transverse baaal
groove uninterrupted ; theelytralinteratioee smoother, the piligerous
imprSBsions coarser, reduced in number, and placed along 1, 3, G, 7,
and 9, those on 7 and 9 larger and more closely placed, 4, 6, S very
Jittle raised. Aedesgus, so far as visible, very similar to that of
S. piloia.
Length 81^-10, breadth 2i-2,% mm.
Hab. Brazil, Constancia and Petropolis {J. Graff
and H. Clark, Jan. and Feb. 1857).
Two males, one with the femoia black, except at the base.
This is a form of S. pilosa requiring a distinctive name.
The sparser vestiture is not wholly due to abrasion, but
to the more scattered pimctuiing of portions of the surface.
Both these insects, when abraded, are so like some of the
smaller species of SlTongylium {a genus of Tenebrionidae
numerously represented in the same region) that they
might easily be taken for such, if the dilated penultimate
tarsal joint were not noticed.
130. SUtlra gradlls.
^. Statira gracilis. Makl., Act. Soc. Fern, x, p. 646
(1875).
c^. Elongate, narrow, shining; piceous, the elytia aeneous with
the suture castaneous, the antennae (except towards the tip),
lege, and ventral surface ferruginous; somewhat thickly clothed,
the legs included, with long, fine, erect or projecting hairs. Head
broader than the prothorax, rugosely foveolate between the eyes,
the latter larger, eubapproximate ; antauiae long, rather slender,
the joints becoming stouter and subserrate outwaida, II equalling
9 and 10 united. Prothorax smooth, much longer than broad,
oblongo-cordate, constricted before the raised basal margin, the
transverse groove in front of it complete, deep. Elytra long, barely
twice the width of the prothorax, subparallel in their basal half,
l„y,l,Ajh,COOl^lc
iV«u> and lUtU-krwwn Lagriidae. 265
wmminsto posteriorly; coarsely, closely Btriato- punctate, the
ponctum beoomtng obaolete before the apex, the intentioM narrow,
trknavenety wrinkled, 1, 3, 6, 7, and 9 eftoh with a loattered seriei
of Bmall, indiatinot, setigerous impieeHiona. Le^ very long, slender,
bairj, the femon cUvate,
$. Rufo-outaaeoua, the protborax and elytra slightly infuscate,
the latter with a faint metallic lustre; antennae shorter, more
thickened outwards, joint II not equalling 9 and 10 united; head
smaller and narrower, trifoveate between the eyes, the latter
mnch Hinaller, distAnt; elytra gradually widened to the middle,
rounded at the sides posteriorly.
I/ngth 8-8}, breadth 2-2J nun. (^$.)
Hab. Brazil, Petropolis [Dr. Sahlberg, Dec. 1850,
type, (J; J. Gray and H. Clark, Feb. 1857, S), Rio de
-Janeiro (fry : $).
Redescribed from two males from Petropolis, and from a
fem&te taken much later by Fry, the latter almost certainly
belonging to the same speciee. S. gracilis approaches
ColpariKrum, but so far as can be seen without dissection,
the mandibles are not tridentate aa in that genus, and the
apical joint of the labial palpi is narrow.
131. SUtira hMmonloldes. n. sp. (Plate XIU, fig. 35, 3.)
Elongate, rather convex, dull above, Bhining beneath ; testaceous,
the eyes, antennae, palpi, and acutellum, the elytra with the intersticM
-4 and 6 (or t^e greater part of tbeir length, and the suture to near
the tip, the under surface in part (the abdomen excepted), the tarsi,
aod the intermediate and posterior knees, block or pioeous ; the
eljtra with a few bristly hairs. Head alutaceous, the eyes extremely
large, auboontiguous: ant«mae very elongate, rather stout, feebly
serrate, joint II equalling 9 and 10 united. Prothorax wider than
the head, a little longer than brood, rounded at the sides, constricted
before the base, the baaol margin thickened and raised ; alutaceous,
-obsoletely canaliculate anteriorly. Elytra elongate, about one-half
■wider than the prothorax, slightly broader at the middle than at the
base, and produced at the tip, the humeri not prominent ; closely,
finely crenato-stri ate, the interstices convex, 3 and S with sevecal
conspieuoua setigerous impressions scattered between the base and
apex, the striae arranged in pairs, obsolete at the tip. Legs very
■elongote, slender, simple.
Length 10}, breadth 2J mm. (^ ?)
Hab. Brazil, Rio de Janeiro (Fry).
n,g,t,7.dh,G00glc
266 Mr. G. C. Champion on
One specimen, assumed to be J, on account of the very
lar^e eyes and long antennae. This insect has the general
facies of a Haemonia [a auhaquatic genus of Phytophi^a
not known in America from south of Mexico) ; it approaches
the Central American S. albolineala. Champ., but has
stouter and more distinctly serrate antennae, much larger
eyes, an almost smooth prothorax, aod nigro-lineate elytra.
There is no trace of an opaque stigma on the elytra. jS.
haemonioides cannot be included under Disema, certain
species of which are somewhat similar. The other de>
scribed nigro-lineate Slatirae are all very different from the
present insect.
ju^fhabbtical numbered list of the sfgoies of
statisa enumerated in the preceding paqbs
(excluding synonyms), the new forms indicated
by an asterisk.
'aoanthomero, 4
•cnapidata, 14
latJcollii, 31
Mgrota,38
'melanoptera. 123
•cyanoptera, 89
ogroides, 8
cylindricollis, 124
ainoena, 28
•dejeani, 55
mdeagriB. 18
aimulata, 112
•dentigera, fl
mi cans, 29
anthicoides. 125
•diluU, 51
*mucronBta, 11
•diatigma, 88
*divi8a, 50
*aayinmetrica, 22
'dromioidee. 92
*octolineata, 82
•batesi, 13
•egaenaiB. 86
elegans. 87
•paraensifl, 104
•bifurcata. 114
•perforata, 127
bryuiti, 95
•elegantula, 99
*penuu)a. 33
*caeruleotiDcta, 59
•eurydera. 68
•pUoaa, 128
•calieneie. 58
fiaurata. Ill
•fiSconus. 126
presutnraUs. 81
•punctieeps, 53
•puocticoUis, 60
*flavMignata, 2
•oanaUculata, S2
•formo«a.30
•caracana, ea
fulvft, 78
catonaU, 19
fimcitaraia, 25
•rotundicoUis, 49
•cavernosa, 43
gemmifer, 7
ruafrons, 34
•cayonnenaui, 120
geniculata. 21
•chalooptera, 42
gracilia, 130
Kratio8a,54
■circumducta, 105
•aegwgata, 101
•conneia, 97
■semicuprea. 47
'convexiuacula, 76
•incisicol^. 109
•wmiiugra, 102
•cribricep*. 122
•inaulftiis, 77
•8erioea,06
•oTDciaU, 116
latoraliB. 71
aiinoni8,52
'cupreoviridiH, 45
l„y,l,Ajh,COOl^lC
New and litUe-known Lagriidae.
*i[^DOdem, 36
"aphenoptera, 40
^pinigera, 3
■plendicanH, 48
"atsnoceph&ls, 36
*Btenodera, 86
^tenoptera. 93
*atrongylioide8, 129
"■nbtenestrata, 106
*aabglat)rata, 67
*auf^Ba, 72
aumtuoso, 41
*tei»upee, 115
•torn
>, 84
tibialia, 26
*tortipeB, 23
•tracliYdeni, 74
■traoHiucida, 66
trilineata,S3
•trisellaU, 103
tnberculata, 15
*ambroea, 67
vogeguttaU, 108
- 10
•Teraicolor, 64
•vigintipunctata, 46
•yiridicioota, 96
*TiridifafK;Uta, 98
*Tiridiiutens, 37
TiridipenniB, 20
'Tiridltincto, 32
•viridivittata, 70
vittata. 79
'xanthodera, 110
SYNONYMS AND VARIETIES.
■Dtillaium [80]
«nnata[81
*tria)lli8[118]
<;olombica[1181
4i9coidalu[118]
fostiva [87]
fl«Tovittat«[lI8]
hiBtrio [1]
tannlataflll]
morbiliosa [29]
plui»icomis[19]
Tiridipennis [55]
EXPLANATIOK OF PlATES XII, XIII.
[Ste StrplaTialiojta facing the Plate».]
November 24, 1917.
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
IX. A Revision oj the genua Tanicus. By G. T. Bethonk-
Bakeb, F.L.S., F.Z.S.
tR»d April 4th, 1917.]
Plates XIV-XX.
A YBAR 01 two i^o Mr. Rowland-Brown queetianed me
on the distinctness of the Euiopean species of the genus
Tamcus, Moore, and my answer was that the clasping organs
of the males were difierent, and that therefore I considered
the species were distinct. This conversation led me to
look up the whole matter afresh, and in doubtful cases
to make more preparations of the genitalia, thus bringing
to light the fact that one of the Indian or a new species
occurred in Egypt and Algeria as well as the well-known
and first-described species iheophTOStus, Fab.; this dis-
covery involved a more extended research of the Indian
species of this complex and very difficult little genus.
My investigations have caused me great searchings of
heart, not on the distinctness of the species themselves,
but rather on the questions " What is a species? " and
" On what characters are we to form species V It is quite
obvious that in this genus it is scarcely possible from the
pattern alone to decide, in certain cases, which is which,
and yet it is equally certain that Butler's and Moore's
species, which de Nic^ville called so strongly into question,
are quite distinct from theophrastua, their clasping organs
are quite different, an4 they also are difierent from each
other in addition to differing from that species described
by Fabricius.
I have endeavoured to make a table of differences in
the pattern between the species, but in vain, for whilst
there is a general look that enables one to assign a name
to the specimens, and as a rule the assignment is right,
yet it often breaks down ; for instance, I have specimens
from Egypt and from Algeria that I had placed under
theophrastus, but theit clasps proved that they were in
reality a new species altogether; again, specimens from
I^ahe] (Arabia) and from India, that I had no doubt were
nara, proved by their genitalia to be ikeophraslvs. There
"TRANS. ENT. 80C. LOND. 1917. — PARTS II, ni, IV (MAY '1 8) T
u,y,l,/eJh,G00glc
270 Mr. G. T. Bethune-Baker on
is, of course, an explanation so far as the patt«m is con-
cerned, for Lahej and Bhuj vere both dry-season specimens,
and this would account for the pattern being small and
but finely marked, even though the specimens themaelvea
were of an average size. The difBcuJty generally occurs
just with such specimens as these ; localities unfortunately
do not render much assistance, for if we are to rely on the
determinations of observers and authors like Moore, de
Nic4ville, Bingham and others (and I think we can rely
on them, as in some instances I have been able to confirm
their identifications from my own collection), the three
commonest Indian species overlap each other. The whole
matter is very difficult and complex, but it shows that
we must not depend entirely on pattern when dealing
with closely allied species. On the other hand, I believe
that in the formation of genera we ought not to com-
pletely ignore pattern, especially where there is a strong
dominant arrangement of it that enables us to identify
the genus of the species directly it ia seen.
The genus is so closely allied to CasUtlim that reference
to the species usually placed therein became necessary,
and I found that in one species at least — ananda, de Nic^
ville — the genitalia led me to think that it mu^ be included
in Tarucus, whilst an examination of the wing scales con-
finned it, for in Castalius there are no " battledore " scales,
but in Tarticus they are plentiful, as they are also in ananda.
The distribution of the genus is interesting; in the
Palaearctic region three species occur which are confined
in that area to the Mediterranean subregion, two being
peculiar to it; six species occur in the Ethiopian region
(including Socotra therein), of which five are peculiar to
it; in the Oriental region twelve species obtain, only one
of them (T. tkeophraslus) occurring outside its limits, and
this one is the most widely distributed of all, obtaining
in each of the previously mentioned regions ; the Australian
legion is just invaded in its Austro -Malayan or Papuan
subdiatrict, for on its extreme western limits a single
species has been recorded from Celebes, and thb, as would
be premised, is peculiar to that wonderful island.
The genitaha are essentially Lycaenine, and they belong
to the Lampides section; they have, however, certain
characters that are quite peculiar, there is no " furca "
— the usual " furca " is a bifurcate organ arising from the
clasps near the base. In the place of this there b another
ih,Cooi^lc
A Reviaion o^ the genus Tarucus. 271
organ of special design, there is also a long hom-Iike
sderite attached to each clasp arising from the same
point, or very near the same point as the special organ
just referred to, but developed in the opposite direction.
These Bclerites appear to me to be capable of independent
movement; this, howevei", would not prevent combined
action with the other organ should occasion arise; they
lie within the hollow of the clasps just above the lower
mai^ns, and rise slightly upwards and forwards, often
extending beyond their (the clasps) apices. It is difficult
to believe that they can assist the clasps at all in gripping
the abdomen of the female, but they might easily drop
rt below the abdomen and be useful as excitatory agents
tapping each side of the lower area of the stemite;
they might well be called the " virgae excitatae." The
special organ leferted to above can be termed the " tec-
torius " (used for a covering); it seems to me to be different
from the "anellus," being in no sense a cone-like tube,
nor anything like it ; it also quite differs from the " manica,"
and is a distinct and peculiar development that I have not
hitherto met with. The " tectorius " rises from a point
in the very rear of the clasps, expanding immediately
hindwards {i. e. towards the abdomen) into two broad
chitinous curtains — one on each side — lying within the
cingulum; these taper more or less rapidly in different
species and assume different contours. At the penal
aperture the anellus becomes apparent in the shape of a
solid ring, or, as in iheophratlua, a loosely fitting tube
through which the aedoeagus passes; this nng is soldered
on to the lateral curtains of the tectorius, which rises
slightly above the anellus, forming a small hood over it.
The whole organ is capable of considerable movement
fore and aft, and in the latter case can be moved right to
the back of the cingulum, in which case it takes with it
the aedoeagus, which is then thrown right out of its usual
horizontal position into a sharply angled one. It is most
probable that this organ (the tectorius) may be the per-
sistence of a very ancient character; it is very closely
similar to the cingulum or girdle, keeping the aiedoeagus
strictly in position, and it appears to me to be a reasonable
proposition to consider it a reversion to the original
structure when there were two seta of armatures, one around
the anus and another protecting the sexual organs; oi it
may be an instance of the survival of an atavic character.
iM-,i,A J h, Google
272 Mr. G. T. Bethune-Baker on
De Nic^ville, when he dealt with this genua in 1890
(Butt. Ind., Ill, p. 187), wrote, " Omitting T. jdinuis,
which is cAsily recognised and belongs to a. difieient groap,
six fonns of T. theopkraslus have been recorded from
India, and are maintained as distinct species by Mr. Butler.
I am able to recognise as satisfactorily distinct two only of
these forms, T. theophrasltis and T. venosus." The author
then goes on to record the various species in the usual way,
but states definitely under each name that he regards it
as a form of tAeophToalus. The very celebrated French
entomologist, C. Oberthur, also records, in his incomparable
" Etudes de Lepidopt4rologie Comparde," fascicule iv,
p. 158 ef seq., a similar opinion, only he goes further and
treats vawsus also as a form of theophrasltis, and he con-
firms this in vol. x, p. 377. Both Bingham and Swinhoe
have followed de NicSville's lead, going further, however,
in that they give as synonyms all the other species except
venosus. I believe I wrote to my old correspondent, de
Nic6ville, and informed him after the publication of his
third volume that both T. baikanica and some of the
Indian species had genitalia quite different from T, theo-
jArastus. I have long intended to revise the genus and
the time has now come when it ought to be done, because
I have no doubt at all that all the species Butler refers to
will stand as entirely separate species from the European
one. He (Butler), however, was not correct in his idea that
Iheophrastus would turn out to be confined to Europe; it
is quite a common species in many parts of India, and it
occurs in Africa south of the Sahara.
The species group themselves by their genitaUa into
three well-marked sections, viz. the baikanica section,
including nara, callinara, extricalus, aUeralus and venosus;
these all have short, broad clasps that are provided with
longish, strong, horn-like sclerites; theophrasttts will form
another section; it has a long, narrow clasp, and a long
aedoeagus ; whilst sybaris and grammica and quadralus
have short very broad clasps, with no hom-like sclerites,
di&erent in shape to either, but more nearly allied to
baikanica than to theophrastus ; the last of the three,
quadralus, forms the connecting link with Fabricius's
species in the increased length of the clasps and the long
tooth at the apex.
It is interesting and curious to find that both in this
genus and also in Castalivs, the species that have been
Ij.y.l.Ajl^yCOO'^lL'
A Revision ef the genus Tarticus. 273
selected as the types of the genua have both long har-
pagones (clasps), by no means typical of the bulk of the
genera, but in each case there are connecting links between
them. The two genera are evidently closely rehit«d.
They can, however, be separated by two characters; the
species of Tamcua have androconia (i. e. battledore scales),
none of the species of Casbilius have them; there are
only two or three that I have not had the opportunity of
examining, and I believe I am correct in thinking that
those I do not know are not Ukely to have these scales.
The second character is the genitalia, the general form of
the clasps is quite different, the aedoeagus differs also,
whilst the shape and position of the cingulum and tegumen
conErm the previous points; there ia also no tectorius.
It will be seen that my conclusions have been brought
about in the first instance solely by my investigations of
the genitaha; these led to the necessity of re-grouping
most of the species and very many of the individual speci-
mens, and in so doing I have discovered other species,
evidently quite distinct, that had always hitherto passed
under other well-known names. Under these circum-
stances it was manifestly advisable to confirm these points,
and for this I turned to the uidroconial scales that are so
marked a character in the males of the majority of the
Jjycaeninae; these amply confirm the correctness of each
of the specific identifications I had made, and they also
confirm the re-grouping of such species as ananda and
bowkeri, neither of which had hitherto been included in
the genus. The South African species (bowkeri) is, I admit,
abenant ; its pattern differs from the rest of its allies in
certain particulani, its clasps differ also, but the androconia
are so close to theophrasltis that I have included it with
the others. In cases of this kind it appears to me to be
better to do this, indicating the small divergencies, rather
than to create a new genus for a single species that otherwise
groups itself very naturally with its close allies.
Tarueosnara, Eollar. Plates XIV, figs. 1, la;
XV, fig. 1, and XIX, fig. 19.
Lycaena nara, Eollar, Hiigel's " Kaschmir," iv, pt. 2, p.
421 (1648).
The general consensus of opinion on the identification
of this species seems to me to be correct ; it is almost as
iM:,i,A J 1^, Google
274 Mr. G. T. Betbune-Baker on
laree as venoms, but the primaries are more triangular
and look broader at the termen. In colour it is lustrous
violet with narrow dark borders. Below it is white with
blackish spots, the postmedian and subt«rmiii&l lines
being strongly marked and but slightly interrupted at the
veins, the former is sharply turned round (almost angled)
at vein 6 basewards to the costa ; in the secondaries the
postmedian line is continuous and slightly crenulated,
whilst the spots are well separated. I think it would be
well to say that I know of no pattern or mark that holds
true in a long series of any species in this small genus with
the exception of sybaris and its allies; they all seem to
gravitate towards theophrastua, whilst theophrastus in its
dry-season forms gravitates towards one or two of the
Indian species such as caUinara or ejirkatus. I have
never experienced such difficulty in separatilig species
as I have done with these. There are only three that I
have not made mistakes with even after the utmost care,
and had it not been for the very distinctive genitaUa of
each species, I should have found it quite impossible to
decide with certainty what some forms were. In diagno«ng
the pattern I have picked a good average specimen, but
with either the dry or wet season forms the diagnosis
will not be absolute. It is fortunate that the genitalia
are quite distinctive, otherwise it would have been im-
possible to say, as we can say now, that Moore's and
Butler's eyes served them well and correctly, when, without
any knowledge of the reproductive organs, tbey described
the forms they had before them as new spedes.
The genitalia {of fiom) are easily separable from iU near alliea.
The olaaps are subovate with the upper margin slightly flattened,
the rounded front edge being furnished thronghout with a row of
sharp teeth. The hornlike sclerites (virgae eicitatae) are wedge-
shaped, tapeting to a point, and not extending beyond the apex
of the claep. The tegumen is typical of the fiist SBCtJon of the
gcnuH, being deeply divided with only a narrow connecting ridge at
the rear; the falees are shortish and stout. The aedoeagna is ^so
fairly typical of this section, it is moderately stout, shortish, with the
front portion beyond the zone suddenly reduced, and rapidly
tapering to a point The claspe an famished with kuig Strang
bristles, and the tegumen with finer and shorter onea. llie teotorius
is very broad at the base, tapering at its tnat edge rapidly np to
the aneltua. The androconia are rounder and shorter sad anuUler
Ij.y.l.AjL.yCOO'^lL'
A Bevinon ^ the genua Tarvcus. 275
than any of ita allies; the apex or distal extremity being mora
deeply convex; tbeie are fourteen rows of laminft with minute
tabercles, the eighth row being the longest, though the row oa each
side runs it reiy close, the eighth, however, marks the summit of
the csonvex apex; the proximal or basal portion of the scale is
asymmetrical, being more rounded on one aide of the Coot-stalk
than the other.
I should, perhaps, explain that in this genus, as generally
with the Lycaeninae, the foot-stalks are quite straight,
arising from near the centre of the androconia, and that
when I refer to the proximal end or base I always mean
the contour of the scale itself, not the nanow foot-stalk,
Tameiu venosus, Moore. Plates XIV, fig. 2 ; XV, fig. 2 ;
XVIII and XIX, fig. 20.
T. venosus, Moore, P.Z.S. Lond. 1882, p. 245, pi. xii, ff. 6,
6a S.
With the exception of T. balkanica, a much amallei
species, this is the darkest of the genus, and it does not
appear to vary in size like the other species, its <^ average
size being about 26 mm., the $ being slightly larger (both
tiara and iheophrastus reach these sizes not infrequently,
but their average is certainly less) ; the colour of the male
is dull sublustrous violaceous, with a single brown spot
at the end of the cell, which is frequently almost absent
in the secondaries. Moore says it has a broad marginal
dusky border (the type form has the border increasing
from the aper very rapidly in the primaries to a quarter
of the inner margin), but this varies, and occasional speci-
mens occur with barely more than a lineal dark border.
The underside is more nearly related to T. theophTOSttis
than to the others; the pattern being spotted, without
lines, t. e. the rows of spots do not form continuous lines,
the postmedian and submarginal series consisting of uniform
and almost parallel rows of large spots in both wings.
The Cashmire form is spotted above as is balkanica, but
below it is typical.
The genitalia are more nearly allied to ballamica, but are decidedly
larger and more robust. This is very noticeable in the clasps and
«ven more so in the hom-Uke sclprites (virgae excitatae), which are
as large again. The clasps are similar in shape, being broad and
rounded on the upper edge, but being suddenly reduced and excised
u,y,i,Ajh,Cooglc
276 Mr. G. T. Bethune-Baker on
near the apei. The tegumen is hood-ahsped, open aboTC, except
for the rounded ridge at the nor ; the falcea are BmoUer in propor-
tion than in balka»iea. The aedoeagus is short and broad, and is
suddenly leduoed at the lone (for this name see Chapman in these
Transections, 1916, pp. 168, 159), where it topers off to a fine point
with two lai^ comuti in the vesica. The hristles on the clasps are
not plentiful and are fine, those on the tegumen equally fine but
longer than usual. The tect«riu8 is developed on the same linee as
in nara, but is shorter, less ample, and somewhat different in outline.
The androconia are oblong, evenJy rounded distally;
the sides oi the oblong are of unequal length owing to the
proximal end of the scale being quite different on one
side of the foot-stalk to the other ; on the one side it is evenly
rounded, the other being excised, the foot-stalk is not in
the centre and thus causes one side of the base to be
longer than the other. There are sixteen rows of lamina.
Tarueus watorstradtl, Dnice. Plate XVII, fig. 16.
Tantcus waterstradti, Druce, P.Z.S. 1896, p. 585, pi. xxxii,
f . 21 ?.
This species is desciibed from a female, and Druce says
that the upperside is very similar to the upperside of the
9 of T. theopkraslus, Fab.
" Underside perhaps nearest to T. venogtu, Moore. Fore-wing :
basal streak shortar and much broader, and extending down to the
Bubmediau nervure, the streak beyond broader and placed at a
much greater angle, the spote beyond the middle more in Une,
the submarginal row distinctly separated, and the marginal row
smaller. Hind-wing : a brood basal streak from just below the
costal margin to the anal angle ; a broad streak beyond, also from
the costal to the anal mo^n ; then a series of spots as in 7. venotu»,
which are more inclined to run parallel with the streaks; then a
submarginal row of targe distinct spots followed by a marginal
row of small spots, the three upper being simply dots, the three
lower gradually increasing towards the anal angle and dusted
thickly with mel-allic green scales. The ground-colour of both wings
is slightly tinged with yellowish and all the markings are black; the
cilia of both wings black.
•'Kina Balu (Wateretr,), Type, Mus. Stand."
The Bomean spedes is evidently a close ally of venoms,
but the imique type is not available for comparison. I
l„y,|,AJI^,COOl^lC
A Revision of the genus Tarucus. 277
have given a photographic reproduction of Druce's excellent
figure in which the underside showa the pattern as well
as the original drawing.
Tarueua balkanica, Freyer. Plates XIV, figs. 3-36;
XV, fig. 3, and XIX, figs. 21, 22.
L. balkanica, Frr., v, p. 63, pi. 421, ft. 1, 2 (1844).
J. Deep lustrous violet, spotted with black-; in the primary
there is a spot closing the cell, a series of six postmedian spots, the
second, third and fourth irregular and shifted outwards from the
first, the fifth and sixth confluent, shifted veil inwards ; these
occupy the same position &e those on the underside, but are not
merely showing through as they are definitely pigmented on the
upper surface ; there U also a small dark cloud in the tomus of the
primary. The underside is white with the spots almost formed
into lines or dashes; the postmedian line is almost cienulate and
practically continuous in both wings; in the primaries it is curved
to the coata from vein 6 and not infrequently is fractured at that
point; the submarginal series on the primaries is prominent and
generally interHecfed at the veins. The principal distinguishing
feature is its deep violet colour with prominent black spots on the
nppeiBide.
The form from the Transcaspian region is unusually large
and fine and is quite distinctive enough to be described as
a local race; I propose for it the name of T. balkanica
areshaTia, vat. no v.
The blue though dark is much moie lustrous than the fonn from
Asia Minor or Syria, whilst the underside pattern is much more
heavily marked, the postmedian and submarginal bands being
more than as wide ^ain and most commonly taking the form of
broad bands, rather than rows of spots. Again, the siie also is
decidedly larger, my specimens of the type form measuring 2 1-22 mm.
against the Aresh form, 26-29 mm. 1 have a series of a dozen speci-
mens which were captiued for mo at Geok Tepe by my friend.
Captain Malcolm Burr.
The species looked so different that at first I mistook
it for ikeophraslus, and it ia referred to again under that
Ttie genitaUa are usually small and slight, the clasps are brood
and very saddenly excised very near the apex, which is reduced
iM-,i,A J h, Google
278 Mr. G. T. Bethune-Baker on
and has more or lew a Htreight terminaljon, finely seml«d; the
homy solerites are aickle -shaped with the blade portion only moder-
ately curved ; the bristles on the claapa are long but not very numer-
ous; the tegumen is of the usual shape, vith large, strong (propor-
tionately) falces, the bristles being fairly long, but not very numerous.
The aodoeagus is short sjid moderately stout and has the apical
portion reduced as in most other species. The teclorius is of
moderate siEe, of the usual shape, with the anellus rather definite.
The androconia are almost oval, the sides being nearly straight,
but not quite bo; they are evenly rounded distally, but not quite
evenly rounded proximally, possibly caused by the foot-stalk being
given off somewhat away from the centre ; there are fourteen rows
of lamina, the sculpturing of which is rather uneven.
It is intereating to find that the androconia of the Axeah
race are markedly diiTerent to those of the type form ; the
scale ia broader proximally almost evenly oval, but in-
creasing in width to near its distal extremity, which is
evenly but sUghtly rounded. The foot-stalk is given off
almost at the centre, and there are nineteen rows of
lamina which are heavily sculptured. The naming the race
areahana is, I think, fully confirmed by the androconia.
Tuuotu eaUinara, Butler. Plates XIV, fig. 4 ; XV, fig. 4 ;
XVIII and XIX, fig. 23.
T. caUinara, Butler, Ann. and Mag. N.H., vol. xviii, p. 185
(1886).
^. Type form. Both wings lustrous violet blue with a single
dark spot closing the cell in the primaries only. Underside entirely
spotted, not in lines or dashes; the postmedian and terminal series
ofspots are parallel and are composed of definite spots in both wings;
the basal marks of the secondaries are also spots.
T, eaUinara nigra, forma nov.
3. Pale, BubluHtrous lilac with a large dark spot closing the cell
in the primaries only, beyond which are one, two, or more dark
spots, smaller than in balkanica, but quite distinct; the marking
of the underside is finer than in the type form and inclined to resolve
itself more into lines, i. e. the spots are apt to become confluent.
This form seems to be commoner than the type. I
have a aeries from Cutch, from Karachi and Campbellpore.
Ij.y.l.AjL.yCOO'^lL'
A Revision (^ the genus Tarucua. 279
It appeals to me to be prpbable that this species and
extTKotus have been mizecl together not infrequently, ae it
is most difficult to separate the type fonu (i. e. the form that
is not spotted on the upper surface) from extricattts, Butler ;
both species are to be found at Karachi at the same time,
as also is nara, but whether they obtain in exactly the same
locality together I have been unable to find out. I fear
I could only separate the type form of callinara from
txtricalus by an examination of the prehensores. These
in callinara ate very close to balkanica.
The clasps are ver; similar, but are decidedly larger and broader,
the sclerites are also larger and heavier in shape and are not so
■hiuply sickle -shaped ; the falceg are also proportionately larger
and lue without the reduced apical hook; the aedoeagus is, how-
ever, narrower than in balkanica and slightly longer; the bristles
on both the clospe and the t«gumen are much finer and ore less
nnmeroua. The tectorina is very ample and broad at the base and
is curved round at the rear up to the anellus, fronUd it is convex
and is strengthened at its edge by extra thickened cbitin folded
over to the upper edge of the organ.
Th« androconia are of a long oblong shape with atraightish
Rides, a fairly even oval base (with the foot-stalk nearly central),
uul bat slightly curved distally ; there are twelve rows of
lamina, of which the sculpturing is very definite and wide
Tanieiis eztrieatas, Butler. Plates XIV, fig. D : XV fig 5
and XIX, fig. 24.
T. exlricatus, Butler, P.Z.S. Lond., 1886, p. 366, pi. xsxv
f . 2 (J.
3. The type is a diminutive specimen from Campbellpore
dated "31.v.'85," the abdomen is missing, so we cannot
decide the point from the genitaUa.
The colour is hiac blue, of a pale tone, the pattern is
composed of fine lines rather than spots ; the specimen
however, is very small indeed, so that the pattern b com-
preesed into a very small area, and would therefore almost
of necessity faU into Unes. The type is a dry-season speci-
men. Those taken m September and onwards are nuire
violet bine, and the underside pattern, though still fine
iM-,i,A J h, Google
280 Mr. G. T. Bethune-Baker on
resolves itaelf into spots &nd lunulea and is less fine than
in the type.
I see no reasoa to doubt the correctDeea of tb« general identiGca-
tione of Ibis insect; the genitalia are distinct from other species,
they are small, the clasps are broad, rounded on the npper edge
and slightly dentate, the lower apex being produced forward some-
what and is very slightly deiitBt«; the homy sclerites are fine
and curved; the tegumen is similar to the usual pattern, but the
falces are very short and angled sharply at the rear; the aedoeagui
is fine and waved with the pointed apex generally obtaining
in the genus. The tectorius is very simple and of moderate
The uidrocouis are oblong, subovate proiimally, and slightly
curved distally; there aie twelve rows of lamina, the sculpturing
of which is rather small and well separated; the foot-stalk is given
oS rather out of the centre.
Tarnens aitontos, Moore. Plates XIV, fig. 6 ; XV, fig. 6 ;
XVIII and XIX, fig. 25.
T. aUeratus, Moore, P.Z.S. Lend., 1882, p. 245, pi. xii,
ff. 4, ia (J.
^. Uppetside : the blnest of the genus with less violet than any
of its allies; it is perhaps the only one that can truly be termed
blue; the spot closing tho cell, in the primaiies only, is not very
prominent, whilst the spot in the secoadaries at the anal angle is
distinct. The terminal dark lino is linear. The underside is gre3ri£h
rather than white ; the pattern is small, very much broken up,
the spots and dashes being rusty red; in some females this may
become tawny brown.
The genitalia ore distinct and large, the clasps being much the
shape of a ham with the apes sharply serrate at the knuckle end;
the hom-Uke sclerilfs are strnighl, stout and long; the bristles are
long, strong and abundant; the tegumen is not bo deeply divided,
with rather finer and shorter bristles and with the falcee Urge and
very strong; the oedoeagus is short with the front part suddenly
reduced and tapering to the tip. The tectorius is rather small,
of the usual shape with the anellus very pronounced. Tlie andco-
conia are large and broad, somewhat ovale |a«ximally, the base
being evenly oval with the footstalk central; the sides are veiy
slightly curved; the distal curve is slight also and not quite even;
there are sixteen rows of lamina, the sculpturing being wide apart
and very distinct.
ll,y,l,/eJh,G00t5lc
A Revision t^ the genus Tarvcus. 281
Tarneus bengalensfa, sp. nov. Plates XIV, fig. 8 ;
XVl. fig. 8, and XIX, fig. 27.
1^. Both wings pale riolet-blue tinged with lilac, the colour being
BcJid and not showing the nndeniide through, except to & veiy slight
extent. Primaries with a conspicuous spot closing the cell. Ter-
nmutl linM blackish. Underside very similar to mediUmmtae,
bnt tbe small series of spots just bejond the celt in both wings nearer
the cell than in that speciee where they are close to the postmedian
tines. Poetmedian and submarginal Unes parallel and decidedly
broader than in the previous species.
Genitalia nearer to T. aittrattt*, but very different from T. mtditer-
rantae, the clasps are Urge aad are suddenly excavated in a deepish
arc midway along the upper edge, from whence they extend in a
broad curve to the apex, which is straigbtish but sharply aerrated ;
the bom-like scleritea aie very long, rather narrow at the base and
tapering quickly to a fine point, they are curved the revetse way to
those of tJie Bpeciee already mentioned and extend well beyond the
apex of the clasp, the bristles are fine, longish, but few in number;
the tegamen is of the usual pattern, less ample and with very long
faloes; the aedoeagus is quite different from either of the species
referred to previously, being more of the bathtnira pattern ; it is,
however, decidedly longer with the apex suddenly reduced at rather
more than a third from the tip, and having a shorter pointed process
from the point of reduction. The tectoritis is of mcdcrate size and
well developed.
Expanse 25 mm.
HiA. Cauctta.
Type in my collection.
The genitalia are so diflerent to the nearest alhes that
there can be no question as to the advisability of aaming
the insect, even though at present it is unique.
The androconia are broadly oval, the curve iliHtatly being slightly
broader than the proximal section and the sides are also somewhat
rounded ; there are eighteen rows of lamina, the sculpturing being
fine and rather close.
Tarnous medlterraneae, sp. n. Plates XIV, figs, "l-lb;
XVI, fig. 7; XIX, fig. 26.
(J. Upperside lilac blue, with a black bar closing the cell in the
prinuuies only, anal spot in secondaries distinct, terminal borders
very nurowly blackish. Underaide, white with dark markii^s.
u,y,l,/eJh,C00glc
282 Mr. 0. T. Bethune-Bakei m
Primariee with tjie basal and subbasal marks as osual, but well
separated ; a long narrow dash from close to tRe costs across the
end of the cell, directly below which is a broader waved dash, theee
are followed by a subcostal spot with a second spot projected far
out between veins 6 and 6, a dash almost below the subcostal spot
between veins 3 and 6 ; postmedian line continuous, obtusely angled
between veins 6 and 6. Submarginal line consisting of a series of
internervular spots. Secondaries with a basal stripe, directly below
which is an inner marginal spot, a median row of fonr spots below
each other, the two lowest of which may be confluent, a dasb closing
the cell, two spots below the cosia generally united, three united
spots projected outwards between veins 3 and 6, two united spots
below the daah closing the cell, a continuous curved line just beyond
these spots, followed by a series of submai^inal spots with metallic
blue gteen suffusion, the second anal spot being the most prominent.
$. Upperside : both wings brown with whitish traces in the discal
area. Underside as in the male.
Expanse, £ 23-26 mm; $ 22-23 mm.
Bab. EoYPT (Alexandria) ; Ai^iria ; Paleotink.
Types in my collection from Alexandria.
Specimens from Cairo are paler above with finer markings
below, whilst the form from Palestine is much paler above
and ia sbghtly larger also. A pair from Biski« (Algeria),
collected by Eaton in 1895, are more heavily spotted below,
whilst the female is well suffused with blue in the basal
area of the primaries.
Lord Rothschild has in the Tring Museum a series
collected in different parts of Algeria, and several hundred
miles into the Sahara. I shall, however, refer to these
again under the species theopkrastus.
In Section I of the genus this species is an excellent
example of the instability of pattern, specimens from
Alexandria and Cairo difiering to some extent, both difier-
ing more markedly from the Biskra pair, whilst these from
Biskra differ quite perceptibly from those in the Tring
Museum from other Algerian localities.
The genitalia are fortunately easily lecognisable and differ from
others of the genus ; the clasps ate large and broad, rapidly tapering
for the apical third, the whole of this portion beingsharply and deeply
dentate, the ajiex itself consisting of two sharp t«eth ; the hom-like
•olerites are very bioad at their base, tapering narrower for two- thirds
where they are angled downwards and are rapdly reduced to a
ih,Cooi^lc
A Reomon cf the genua Tarucue. 283
fine pcHiit extending to the apei of the claspa; the bristles are
longisb, moderately stoat, but not numerous ; the togumen is fairly
large of the usual pattern with strong falces; the bristles being
finer &nd shorter than thoee of the clasps; this is, hoverer, usual;
the aedoeagus is long and vaved ; the vesica being finely shagreened,
and the tectorius ample and of the usual shape ; it is, 1 think, the
largest in the genus. The androconia are very broad, and were it
not that one side of the proximal extremity is excised, it would
form an evenly-shaped oblong; the foot'Slalk is given ofF centrally;
there are seventeen rows of lamina rather widely separated, whoee
sculpturing is somewhat small.
Section II contains three species, T. grammiea, G. -Smith,
T. starts, Hopffer, and T, madralus, Grant. The first is
a species from Mombasa and from Somaliland, the second
a widely-spread, if local, S9Uth African insect, and the laat
a species from Socotra,
The genitalia in all of these lack the horn-like sclerites
which are so peculiar a character of the first section;
guadralus, however, has developed a tusk-like extension
of the upper apex of the clasps, and thus forms a connecting
link with Section III, containing only the type of the
genua. All three species in Section II have the same type
of aedoeagus as has Section I.
Taraetu grammiea, Grose-Smith. Plates XIV, fig. 9, and
XVI, fig. 10.
Lycaeneslkes grammiea, Grose-Smith, Rhop. Exot., ii,
p. 102, pi. xxiii, ff. 3, 4 (1893).
(J. Both win^ dark brown. PrimarieB with a darker, spot
closing the oell ; secondaries with a, terminal row of Bpot« encircled
with white, more prominently on the inner side. There is a trace
of a similar row of spots on the tecmen of the primaries, but it is
very obscure. Underside white with the markings of the primaries
large. From the spot closing the cell in the primaries there is a
short, broad dash forming an L with the cell spot; the broken series
of marks outside this is imited into an irregular band, very broad
below vein 2; the postmedian series of spots is pushed far out,
near to the eubterminal series, the former being composed of fair-
sized spota increasing in size towards the. inner margin where they
coalesce, the latter consists of six intemervular smaller spots;
the nsual basal dash and subbasal wedge-shaped mark are present.
Secondaries : a basal subcostal dash with a spot below it, followed
iM-,i,A J h, Google
284 Mr. G. T. Bethune-Baker on
by four short distinct dashes; ft short costal and subcostal con-
fluent mark touching the spot closing the cell, below wbich is another
short dash ; outside the cell spot are three irregular confluent spots,
beyond which is the postmedian row of spots moderately evenly
ourred, followed by the subterminal row, some of which are slightly
iridescent.
Genitalia : the clasps are very brood and are evenly curved, but
if flattened somewhat wedge-shaped; the bristles arc fine and not
long; the tegumen is of moderate size, and the falces are small;
the bristles line and shortish; the aedoeagus is shortish, suddenly
reduced all round about the middle, from whence it gradually tapers
in a curve to a fine point. The tectorius is very reduced.
This is a very distinct species and was first described by
Grose-Smith as Lycaenesthea grammica in 1893 (l.c.).
In 1898 Miss Sharpe described, her huisae, as it had not
at. that time been discovered that the former species had
nothing to do with the genus LycaenesOiea ; louisae therefore
falls as a synonym to grammica.
Tarvom sybarls, Hopffer. Plates XIV, fig. 10 ;
XVI, fig. 9, and XX, fig. 28.
Lycaena sybaris, Hopfi., Monats. K. Preuaa. Akad. Wis-
senach.. p. 642 (1855).
3- Blue tinged with mauve in both wings. The primartea with
a black spot closing the cell, and broadish black margins; the
secondaries with a submarginal row of black spots from the uial
angle to the costa, decreasing in size as they approach the costs;
outside this row is a fine white line, which ia succeeded by the black
termen of uniform and moderately narrow width. The fringM
have the basal half black and the outer half gi^y wilh a slight
indication of tessellation. Underside white spotted with black,
the contrast being sharper than in any other species. The primaries
have the usual basal marks, the wedge-shaped mark being genemlly
somewhat L-ehaped ; a large spot closes the cell with a small one
between it and the costa, beyond which is another somewhat larger
one between veins 6 and ft, between veins S and 3 are two conBuent
spots, and two more confluent spots further inwards are below vein 3,
between S and 6 is a single isolated spot, shifted right out on to tbe
poBtmedian curved series of six inlemervular spole, this being near
the margin and very close to the subtcrminal series of intemervular
Bpole. Secondaries with three basal spots and four subbasal spols
below each other ; the upper two basal epote are connected to each
lj,y,l,Ajh,COOl^lc
n,g,t,7.dh,G00glc
Explanation op Plate XIV,
Fio. l-la. Tarueusnara,^, p. 273. ,»
«n(««.,cJ,'p.27V
baltaniea,!^, p. 277.
eallinam, ^, p. 278.
etiricafiM, J, p. 279.
alUmtug, 1^, p. 280.
vtediUrranrae, (J, p. 281.
mtditerraneae, $, p. 281.
6ensaferwM, (J, p. 281.
grammica, p. 283.
a7,baTie,^,p. 284.
7uadralu«, (^, 28S.
Iheophmsliit, ^, p. 286.
anando, ^, p. 289.
ttm'jfceft, (J, p. 294.
dharla, J, p. 291.
efalhratus, $, p. 293.
n,g,t,7.dh,G00glc
Trms-Ert. Socloni ,1917, PI XK
6,
If
31>.
EC Kniyit drt.et chTOBO.
THE GENUS TARUCUS.
U.
12.
12b..
^ 13.
™^
14.
15,
1'
ih, Google
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
Trans. Ent. Soc. Land., igij, PlaU XV.
G. T. B.-B., Photo. Andrt. Sltigk &■ Anglo. Ui.
GENITALIA OF THE GENUS TARUCU5.
l„y,l,Ajh,COOl^lL'
■ Explanation of Plate XV.
IS nam, p. 273.
venogM, p. 275.
balkaniea, p. 277.
cattinara, p. 278.
txtricatua, p. 279.
alttratui, p. 260.
The figures of the genitalia are magnified 30 diameters.
n,g,t,7.dh,G00glc
Explanation of Plate XVI.
7. Tamcut medUtrrantat, p. 281.
bmgalengis, p. 281.
Bffbaria, p. 264.
10. ,. grammica, p. 283.
11. „ quadralva, p. 285.
12. „ Ihfopkraglus, p. 280.
The figures of the genitalia are magnified 30 diameters.
n,g,t,7.dh,G00glc
TVawr. Ent. Soc. Lond., igi?, Plate XVI.
^
<-^
G. T. B.-B., Pkolo. Andrl, SlHgk &■ Anglo. Lti.
GENITALIA OF THE GENUS TARUCUS.
u,y,i,Ajh,Coot5lc
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
TVam. Ent. Soc. Loud., igjy, PlaU XVIT.
M.W
G. r. fi.-S., PAofo. Andri, SUigh &. Anglo. LU.
GENITALIA, etc., OF THE GENUS TARUCUS.
n,g,t,7.dh,G00glc
Explanation of Plate XVII.
Fio. 13. Tarvciuananda, p. 280.
dharia, p. 291.
bowktri, p. 294.
laiteratmdii, p. 278.
fageialua, p. 292.
Itapardag, p. 2!i:t.
The figures of the genitalia arc niagnitied 30 diamelcri>.
n,g,t,7.dh,G00glc
Explanation of Plate XVIIl.
Each Fiovre is named ok the Plate.
Tamcus calUnara, p. 278, upper left-hand figure.
„ venoiiia, p. 275, upper right-hand figure.
„ alUralus, p. 280, lower left-hand figure.
„ Iheopkragtvi, p. 286, lower right-hand figure.
Tlie figures od Plate 3CV11I of the virgae exeiUilae and teettiHvt
are magnified about 66 diameteis.
n,g,t,7.dh,G00glc
Tratu, Ent. Soc Lortd., zgij, Plate XVIIl.
G. T. B.-B.. dtl. Andri. SWgk &■ Anglo. Lid.
VIRGAE EXCITATAE AND TECTORIUS OF THE GENUS TARUCUS
n,g,t,7.dh,G00glc
n,g,t,7fdh,G00glc
n,g,t,7.dh,G00glc
TYans. Ent. Sac. Lend., 1917, PlaU XIX.
G. T. B.-B.. dtt. Andri. Sltigk & Angh. Ltd.
ANDROCONIA (Battledore SoUea) OF THE GENUS TARUCUS.
n,g,t,7.dh,G00glc
Explanation of Platb XIX.
Fia. 19. Tanew nam, p. 273.
80. , „ lenoffMi, p. 275.
21. „ baiJcaniea, p. 277.
22. „ Ixdkaniea aruhana, p. 277.
23. „ ealliTuiTa, p. 278.
24. „ txiriealue, p. 279.
25. „ alteralus. p. 280.
26. „ medilerrantat, p. 281.
27. „ bengalenaia, p. 2S1.
The scales ore magnified 500 diameters and the palpi
diameters.
n,g,t,7.dh,G00glc
EbcPLAMATION OF PLATE XX.
FiQ. 2». Tarueiu tybarit, p. 284.
Vttophnutui, p. i
30.
a«anda, p. 289.
31.
bmritri. p. 294.
32.
33.
rtropArartiijr, p. 286.
34.
(AropAnwhM, p. 286. denuded of it*
scales.
The scales
re magnified
600 diametera and tl>e palpi 30
ameters.
n,g,t,7.dh,G00glc
TraHs. Ent. Soc. Land., igij, PlaU XX.
AvM, SUigh &• Anglo. Lti.
AHDROCONIA (BftttledoK Scales) AND PALPI OF THE
GENUS TARUCUS.
iM-,i,A J h, Google
,t,7.dh,Googlc
A Revision of the genua farucus. 285
other by a fine line ; a. Uigish spot closes the cell, beyond which are
three pain of spots, two oblique ones below the costo, two simiUrly
oblique between veins 3 and 5, and two below each other, shifted
inwards between veins 3 and la ; a spot shifted out close to the post-
median row between veins 6 and 6; the postmedian row of eight
intemervular spots is strongly curved, beyond which is the sub-
terminal row of seven spots, the lowest anal oi» being in the shape
of a short dash, that and the two above have inet«llic blue scales
BQperimposed.
This is also a very distinct species easily separable from
any of the otheTs.
Genitalia : the clasps are broad and wedge-shaped, but do not
taper down to a point, the apex t>eing somewhat rounded ; the
tegumen is deeply excavated as usual, but has lobed cheeks or sides ;
the falces ate of moderate length and strength, and rise from just
below the upper lobes. The aedoeagus is of moderate length and
width, reduced suddenly at the middle by the excision of the upper
half, the lower half tapering laterally to a fine point. The tectorius
is developed, but slightly.
The androconia are of a long, oblong shape, with an asymmetrical
ovate base, the distal extremity being very evenly curved, with the
sides straight and long, the footstallc is given ofi from near the
centra, but not centrally ; there are seventeen rows of lamina, the
sculpturing being smallish but veiy diattnct.
Tarmus quadratus, Grant. Plates XIV, fig. 11 ; XVI, fig. 11.
Tarticus quadratus, Grant, Bull. Liverpool Mua., ii,
p. 10 (1899).
The blue of thia species is very similar to that of T.
ttfitaris with a prominent black spot at the end o£ the cell
in the primaries. The underside is similar in the strong
contrast of the black markings on a very white ground;
the pattern conaiate of spots, but by no means so isolated
as tney are in aybaria. The inner wedge-shaped mark at
the base of the primaries is very large; the postmedian
line in both wings does not consist of isolated spots as in
sybam, but rather of lunules intercepted by the veins;
the subterminal rows of spots in both wings are composed
of isolated spots, but the subbasal and median series of
spots are by no means so isolated as in Hopffer's species.
TRAKS. EMT. 8O0, LOND. 1917. — PABTS !(, Ill, IV (MAY '18) U
u,y,i,Ajh,Cooglc
286 Mr. G. T. Bethnne-Baker on
The genitalia are very distinctive ; the clasps are unusuaJlf broad
and are somewhat excised on the upper margin near the base, as will
be seen from the flgiue (PL XVI, fig. 11), after which they rise in
an even curve, and at a third from the tip they taper off into a
tusk-like projection at the upper apex, below which the front edge
recedes and is faintly dentate lo the lower apex, which is produced
very slightly forwards in n dentate termination; the bristles ar« of
moderate length and are not numerous ; the tcgumen ia of moderate
dimensions, deeply excavated to the back ridge, which is nairow;
the cheeks or sides are almost lobed, and the falcee are stout and of
a fair length ; the bristles are very flue and plentiful on the lobes ;
the aedoeagus is of the usual type, shortish, rather narrow, the
reduction taking place near the centre, from where it gradually
tapers off to the pointed apex. The tectorius is but weakly
developed.
I regret I have been unable to procure specimens of this
insect so as to figure the androconia, the ouly specimens
X have been able to examine being those in the National
Ck>llection.
Taruous Iheophrastus, Fabricius. Plates XIV, figs. 12, 12a;
XVI, fig. 12; XVIII and XX, figs. 29, 32-31.
Hesperia Iheopkrastus, Fabricius, Ent. Svst., iii, p. 281,
No. 82 {1793).
If it has been difficult to give a satisfactory diagnosis
of the pattern in most of the species dealt with in Section I,
it is even more difficult to give one for the type species
of the genus. Even Moore, whose eye for minute differ-
ences in pattern and general aspect was far more keen
and critical than any one I have known, failed with this
species.
The colour of the upperside may be violet blue or lilac
blue. I have specimens from Souk Arras (Algeria) that
are violet blue of a solid texture, and in one case it has a
very large black spot, closing the cell with an angled dash
beyond it. Examples from Asia Minor, in my collection
(exact locality unknown) may be very similar in colour
to the Algerian ones, but more generally they are lilac
blue and are decidedly transparent; all, however, have
a prominent spot Closing the cell. Bhuj (India) and
Labej (Aden) specimens are similar to the lilao ones, but
with a finer mark closing the cell, whilst a pair from Senegal
n,g,t,7rJM,GOOglC
A Revision q^ the gernu Tameue. 287
are of a mach more delicate lilac blue, with a small spot
closing the cell; in all the black t«nneD is moderately
narrow, not Unear.
The underside of the Medit«rranean form is usually
strongly spotted, the spots being often large and very
black, excluding, of course, the basal streal^, that is to
say, the spoU are generally more or less distinct in both
wings. The form from Asia Minor and from India has
the postmedian band in the secondaries as a series of
lunules, or a more or less broad lunular baud. In my
specimens from Senegal the <? pattern is very fbe, but the
$ pattern is heavier and more Uke the type form.
The genitalia are totally distinct in some particulars from all
others in the genns; the clasps are quite diverse, as also is the
aedoeagna, whilst the parts represenling, perhaps, the hom-lite
sclerites rest in an entirely different position. The clasps are long
and of moderately even width, tcnninating in thrtM) tusk-like teeth,
one at the lower eitremity and two deeply -divided at the upper
extremity; the bom-like sclerites, instead of lying along the hollow
of the cl«Bp,*rise immediately from their origins at the extreme
base of the clasp, and occupy a position just above its upper margin
at the rear, they do not extend much beyond the centre; whilst in
the first section they lie almost " perdu " in the hollow of the basal
part of the clasp and rise into prominence well beyond the centre,
and are always within the clasp itself, their apices only extending
occasionally outside. The acdoeagus is very long, of moderately even
and narrow width, tapering slightly to the apes; the vesica iscchinoid
in form bat without the spines, it is a very delicate and beautiful
object. The tegumen is very deeply divided, with narrow wedge-
shaped cheeks which carry prominent falcea; the hristles are fine,
of moderate length and number; there are, however, very few on
the clasps; the tectcirius is well developed, but quite diflerent in
shape to all its allies.
At first sight it would appear that the Mediterranean
specimens with heavily marked undersides would probably
be theophrastus, but that is not really the case, and it is
very far from being so with those that have a finer under-
side pattern. I have a series from the Caucasus that I
bad no doubt were this species — large, heavily-marked
spectraens; but the pattern was very confluent, and this
impelled me to examine the genitalia ; these proved quite
conclusively that the species was T. balkanica, whilst part.
iM-,i,A J h, Google
288 Mr. G. T. Bethime-Baker on
of a series that is in the Tring Museum from Algeria with
a finer pattern, that was flying freely with T. mediterraneae,
also proves (o be balkanica, thus extending the range of
this species in a very unexpected manner.
In the Tring Museum is a large series of specimens
collected in Algeria, with a few from Egypt and Morocco.
They belong to three species; tkeophrastus is the most
abundant, mediierraneae is leas so, whibt balkanica is
much the rarest. There are in all some 477 examples,
and when first I sorted and examined them I must confess
to a sense of complet« bewilderment so far as regards the
first two ; the upperside of balkanica marks it out fairly
distinctiv from the others, but it took a long study before
I was able to sort out with any degree of certainty tkeo-
phraatus and mediierraneae, and in the end I found the
only way of coming to any satisfactory conclusion was to
dissect a good number of specimens ; this Lord Rothschild
kindly let me do. I had considered at first that nearly
all the specimens were my mediierraneae; the genitaUa,
however, proved that the great majority were Aeophrastus.
At Guelt-es-Stel, in the Hautes, there are 164 of Fabricius's
insect to nineteen of my new species ; they were, however,
flying together on the same ground and at the same time,
in June, July and August, though it was in the last month
that they were moat abundant; at Nedroma (Oran)
nine specimens of tkeophrastus only were taken ; at El
Kantara rather more than twice as many tkeophrastus
as of mediierraneae occurred; at the desert post of £1
Hadrada ten of my species were taken and one balkanica,
but no tkeopkrasltts, whilst at Ghardaia, far in the Sahara,
one Iheopkrastus, twenty- one meditenaneae, and five
balkanica were captured, and at St. Oued Mya (Sahara)
eight of the latter and a pair of mediierraneae were collected ;
the other localities yielded much the same results, except
that at Biskra eleven mediierraneae and three iheopkrastus
were taken, and at Batna there were thirteen of the latter
to nine of the former. In Morocco both species occurred,
from the Masser Minea only theopkrastus was sent home,
and from Zoudj-et-Beghal only nCediterraneae; there were,
however, but a few in each instance.
We see therefore, on the whole, that mediierraneae
appears to thrive better than tkeophrastus in the desert
localities and vice versa. The distribution of the species
is so unusually interesting that I have given it somewbafc
ih,Cooi^lc
A Revm<m <^ the genus Tarucus. 289
in detail, though I have omitted a number of localities
from where very few or only one specimen were sent home.
M. Oberthur writes me that he has Iheopkrastus in
considerable numbers from Biskra, Bon Saada, Djurjura
and Sebdou. From the second-named locality there are
two medUerraneae and one tkeophraslm in the Tring
Museum ; no doubt, taking into consideration my dissec-
tions, both species will be found in the Rennes Museum
also.
After my dissections had been done, I sorted the species
in accordance with those results, and they gave me 8
fwrly easy rule of separation. I foimd that all my tfieo-
■phrastus dissections came from specimens in which the
postmedian lines in both wings were separate spota, not
confluent ; in the hind-wing they were frequently more or
less fine lunules, but not a continuous crenulated line ; whilst
in medUerraneae these lines were continuous, sometimes
fine, but often very broad and heavy. This would, there-
fore, seem to be a fairly safe character whereby to separate
the specimens.
I found that this was confirmed in tkeophraslus from
India and other p^rts of Africa, in all of which the post-
median lines are composed of separate spots or lunules.
The androconia are somewhat ovate, but truncated diatally into
a, broad arc, tbe proximal extremity being asymmetrically ovale
with the foot-Btalk given off noD-cenlrally ; there are twelve rows
of lamina widely separated with the sculpturing very defined and
well apart It frequently happens that abnormal androconia are
present, and I have figured one of theMe found on the same wing
vith the normal one of this Bpeciee.
Tamous ananda, de Nic^ville. Plates XIV, fig. 13;
XVII, fig. 13, and XX, fig. 30.
CasUdius ananda, de Nic4ville, Joum. A.8.B., lii, pt. 2,
p. 75, pi. i, fi. 11 ,S, llo ? (1883).
(5. Uppeiside deep luatrous violaceous with a distinct dark border
of moderate width in each of the wings, quite markless except that
the undcrude apola show through indistinctly . Underside, whitish
iFitb dark marks and spots. Primaries with a broad baaal costal
stripe, and an irregular, broad (not wedge-shaped) dash ascending
to it from the inner margin, outdde which is a, sbo^, very broa^
iM-,i,A J h, Google
290 Mr. G. T. Bethtme-Bakei on
da«h from the costa to the end oE tbe cell ; postroeiliiui line composed
of two confluent spots on the coata, a spot between veins 6 and 6
projected right out into the subroarginal row, two confluent spoU
between veins 3 and 6 below the firet two, and two confluent spot«
between veins 1 and 3; theee ore stiift«d inwaids, and tbe lower
spot is quite snuJl, outside these is the submarginol, veiy even row
of six moderate-siied intemervular spots, this being roughl j parallel
with the terminal row of six smaller inteitoervniar spots; tennen
finely dark. Secondaries with a broad basal band right across the
wing; four sabbasal large spots below each other, tbe upper
three more or loss confluent, a spot cloning the cell: poatmedian
series consisting of three pair of confluent spot« followed by one
on tbe inner margin, tbe second pair of spots is shifted outwaids ;
submarginol row composed of seven intemervular spots, the third
from the costs being largo on account of its coalescence with tbo apot
projected out from the fioetniedian series; terminal row composed
of six spots, the two anal ones having a very few superimposed
blue scales that are easily rubbed off; termen finely dark.
$. Entirely brown in both wings; a whitish cloud in the disc^
area, the underside discal pattern showing through. In the secon-
daries there is a prominent terminal row of dark spots edged in-
wardly with pale dashes. Underside like the male.
Genitalia : the clasps are shovel-shaped, broad, with the whole
of the upper and apical margins irregularly and widely serrate, the
apical margin being somewhat concave; the bristles are very fine
and very few; a sclerite arises from the base of each clasp as in
thtophrattut, but is longer and B(«uter, extending along twO'tJiirds
of the upper margin ; the cingulum is ample; the tegumcn is deeply
excised with long, strong falces; the bristles are more numerous
than on the clasps, but are very fine; the aedoeagus is raodcmtely
long, stout, and tapering to a fine tip near the apex; it is provided
with two long sclerit«B, lying internally, one on each side, the vesica
is furnished with two rather long haiiy brush-like processes — like a
fox's tail — which are capable of inflation and thenassumean almost
spherical shape; the tect«rius is small, being reduced to a broad
column thrown right back in the rear of the clasps. The androconia
are proiimatly asymmetrically ovat« with the foot-stalk given off
slightly sideways; the sides are very slightly curved; the distal
extremity being slightly and evenly convex; there are sixteen or
seventeen closely placed lamiita whose sculpturing is irregular and
rather smalL
This well-known Indian species is easily recognised from
all its neighbours but one, for here also I hav« found two
l„y,|,AJh,COOl^lC
A Revision o/" the genus Tarucus. 291
species mixed together that have revealed themselves
by their sexual oi^ans.
I have placed these species in the type section on account
of the position of the sclerites that replace the " vii^ae
excitatae," and on account of the aedoeagus and reduced
tectoiius.
Tuaeas dharta, sp. nov. Plates XIV, fig. 15 ; XVII, fig. 14.
(J. Upperside sublustroua violaceous with brown borders to each
wing of moderate width. Underside whitish with brown markings.
Nroaries with a broad costal band and a broad erect subbasal
dash (not wedge-shaped), both much shorter than in T. ananda
and more separate; a broad dash aciOBS the end of the cell sur-
mounted by a spot shifted somewhat outwards; postmedian lino
consisting of three pair of confluent spots and a single one shifted
well outwards, but not touching the submaiginal series; the
second pair of spota is shifted slightly outwards and the third pair
well inwards; submarginal row composed of six fair-sized inter-
nervular spots followed by the terminal row of six intemervular
daahes ; tennen finely dark. Secondaries with a broad basal
dash, followed by four largish spot« below each other, the upper-
most costal spot being very large ; these spots are almost confluent ;
a leniform mark closes the cell; postmedian series composed of a
pair of confluent subcostal spo<«, three confluent spots shifted
outwards and almost forming a broad dash, a pair of confluent
spota shifted inwards under the renlform spot; postmedian row
composed of eight intemeural spots followed by a trace of a terminal
row of dashes represented by three fine intemeural dashes at the
apex and two spots at the anal angle ; t^rmen flnely dark.
$. Both wings brownish with whitish discal areas and moat of
the undeiside pattern showing through. Frimariea irrorated on the
fold and the discal area with brilliant lustrous blue, and the post-
median TOW of spots showing prominently through. Secondaries
with a trace of the blue in tlie diecal area, and a line series of whitish
intemeural dashes outside the postmedian row of spots showing
through. Underaide precisely like the male.
Expanse, ^ 25; $ 26 ram.
Hab. SiXKm, Darjeeling.
Types in my collection.
This species may be recogniaed from ananda in the
greater separation of the underside pattern throughout,
and especially in the submarginal and terminal lines ; the
female is markedly difierent.
iM-,i,A J h, Google
292 Mr. G. T. Bethune-Baker on
Genitalis : tiw clasps ue of moderate size, broad at the rear,
but tapering gradually to the apex and temuDating in a rounded
extremity; the ",virgae excitatae " are replaced by a peculiar
structure, and is for nearly two-tbirds of tbe basal portion a portion
of the lower margin of tbe clasps, then it suddenly develops into a
longish narrow rod wbich extends well beyond tbe end vi the clasp ;
tbe cingulum u narrow, ■ncieasing in width as it approacbee the
tegnnien, which is of moderate sue, deeply excavated with fairly
developed (aloee ; the aedoeagns is peculiar of moderate uze at the
rear behind the mi^, but for the apical two-thirds it is very narrow
indeed, and tapera gradually into a fine point; tbe tectorius is
reduced, thrown right behind tbe clasps, as in tbe previous species,
having no hood, but being of a different structure, and in a different
position from tbe furca.
There remain four other descriptions to be considered,
Taructis (P/efctW) fasciattts, Bftber, from Banks, Tarucua
Uopardvs, Schultze, from the PhiUppines, Tarucus dath-
ratus, Holland, from Celebes, Tarvcus flumalis, G. -Smith,
also from Celebes. These form a small section wherein
the spotted pattern is transformed almost entirely into a
" banded " pattern, the spots having become confluent
so as to form broad definite bands across the wings, thus
making them separable at a glance from any others of the
genus.
Tarucus fasclatus, Bober. Plate XVII, fig. 17.
Plebeius fasdatus, Bober, Ent. Ver., " Iris," i, p. 194,
pi. ix, f. 15 (1887).
Bober describes the species (freely translated) as
follows : —
" $. Upperside smoky grey with the median ares of all the wings
whitish, the basal area being weakly irrorated with tight blue ; at
the termen of the hind-wings tbe black border spots of tbe under-
side show through; hind-wings with a white terminal line, shortly
tailed at vein 2 (M'). Underside yellowish white with blackish
grey bands, and terminal rows of spote-
" Expanse, 27 mm."
Hab. Banka Island {H. Kuhn).
There is an excellent photograph of the insect (" Iris,"
1887, pi. 9, f. 15, p. 194), so sharp and defined in all
particulars that there can be no difficulty in identifying the
species. I give a reproductioo of it on PI. XVII, fig. 17.
ih,Cooi^lc
A Revmon <^ the genus Tarucus. 293
Taruens eUthralus, HoUand. Plate XIV, fig. 16 $.
T. cUahratus, HoUand, Proc. Boafc. Soc., xxv, p. 71, pi. 5,
f. 8 (1891).
Holland describes his species thus : —
" Near T. {Plebeiug) fasciaiaa, Boeber (' Iris,' vol. i, p. 194), but
smeller and differently marked. Tbe uppenide of the wings is
lilac, with Bmoky grey margins. The black msrkipgs of the under-
Etide are distinctly visible from the upperaide. In faaciaiu* the
submarginal black line is narrow, in dathratnt it ia broad. In the
former the two succeeding black lines are distinct throughout; in
the latter they unite, forming a nido figure of the letter V, There
are many other minor differences which readily reveal themselves
upon a comparison of the two Bpeeles, and which are mote enaily
seen than described. The type, a male, is in my collection."
The habitat is Celebes, and I quite concur in Dr. Holland's
opinion that his species is distinct from Rober's. I have
no doubt, however, that Grose-Smith's T. fluvialis is the
female of Holland's clcUAnUus. Smith gives a long and
careful description of his species, a female, which also
comes from Celebes, saying that it is nearest to T.fasciatus,
Rober, but that the arrangement of the bands on the
underside is quite different, and then he goes on to say
that " it may not improbably be the female of T. dathratus,
HoUand, but that his figure is so diminished that it is
impossible to decide with certainty whether this is the case."
I have enlarged Holland's figure, and find that the pattern
is precisely the same below, only in the original, being a
male, the bands ate narrower than in the Tring tvpe.
The underside shows through in just the same manner,
and I do not think there can be any doubt as to tbe identity
of the two insects.
Tarucus leopardus, Schultze. Plate XVII, fig. 18.
T. kopardus, Schultze, Philippine J. Sci. D., 5, p. 161,
pi. l,f. 9(1910).
The species was described in the Phihppino Journal of
Science. The following is a copy of the description : —
" i^. Uppereide of wings irideBcent, purplish blue with a narrow
dull black line along outer margin; cilia white. Tail black, tipped
with white. Hind-wing with two indistinct submarginal spots at
posterior angle. Underside of wings whit« with a faint ochraceous
iM-,i,A J h, Google
294 Mr. G. T. Bethune-Baker on
tinge fUid nmaerons dark brown markiDgB aa foUown : a narrow
subcostal band from base to. middle of fore-wing, thence obliquely
toward posterior outer angle ; basal area with three triangular
patches, the medial one being most prominent; outer area with
three oblique bars from the costa; the second, which ia the longeet,
reaching vein IV; a prominent, nearly round, poetmedial spot
between veins III and IV ; a submarginal band and a marginal row of
six very small spots, each between two veins, and a veiy fine mar^nol
line. Cilia white. Hind-wing with a baeal bar, six antemedial
transverse streaks, two medial transverse etreake, two postmedial
streaks, and a~ postmedial band from inner margin to vein VI ; a
prominent submarginal band, six submarginal spots and a fine
ant«ciliary line. The submarginal area between veins I and 111 ia
dark ochraceous, and the two black spots upon it ace surrounded
by a highly metallic, green line.
" $. Upperside of wings grayish brown with a bluish iridescence
on basal half. Discal area whitish. All markings on underside
similar to those above, although less diffused. Hind-wing with the
submarginal row of spat^ large and distinct. Underside similar
to that in ij; all markings somewhat larger.
"Length of wing, S i' miUimetera; $ 13'5 miUimct^rs.
" Luzon, Province of Camarinea, Paracale, P.I, (J. P.
Iddings collector). Type (J, $ and co-type No, 12743 in
Entomological Collection, Bureau of Science, Manila, P.I."
From the figure referred to, which I reproduce on PI.
XVII, fig, 18, I believe that this species is allied to
dalhratus, Holland.
This brings us to the last two species of the group,
that I should perhaps have hesitated to include in the genus.
The pattern is different, but the androconia have quite
the same shapes and facies, and the genitalia have the
remarkable form of the " tectoriua " strongly developed,
but modified ; they are, therefore, better placed here than
elsewhere, at least, for the time being. Trimen himself
indicated an alliance between them and sybaris.
Tarueus bowkerl, Trimen. Plates XIV, fig. 14 ;
XVII, fig. 15; XX, fig. 31.
Lycaena bowJceri, Trimen, Trans. Ent. Soc. Lond., 1883,
p. 351.
{J. Both wings on the upperside are lustcous pale violet blue.
PcimatieB with a dark indefinite spot oloaing the cell, with a IoomI
n,g,i,.eih,.G00glc
A Revision o^ the genus Taructis. 295
black tormen and black and white tessellated fringe. Secondaries
with an indefinite spot closing the cell, a very obUque row of dark
poetmedian spots and a complete row of dark eubmarginnl spots,
both these rows occupying the position of the underside pattern,
bat not being merely the showing through of that pattern; a
terminal series of dark spots merging into the black termen around
tbe apex; fringes white with black teesellations at the ends of tho
veins. Underside white with ash brown spots. Primaries with
the snbcoBtal basal stripe, a spot in the middle of tho cell with
two other spots below it, representing the usual wedge-shaped
mark; a good-sized spot closing the cell with one or two above it,
beyond which are three very oblique spots from tho coata, with two
confluent ones below tho lowest of the three, but shifted inwards;
two confluent squarish spote almost below the one closing the cell,
a series of six intenteurol submarginol marks fallowed by a terminal
row of six distinct spots; termen consisting of finoblsckdasheemter-
aected with white intetneurolly. Secondaries : a costel basal spot
below which is the basal stripe which is very irregular, four subbasal
spots below each other with another small spot from the second
connecting it with the spot closing the cell ; postmedian series
{composed of eight spots) widely separated between the second and
the third spots, the lower six forming a very oblique row from vein 6
to the inner margin ; all the spots hitherto referred to in the second-
N7 are somewhat darkly encircled; submarginal row of seven
marks very irregular; terminal row compoi^ of seven distinct
spots, the iiiBt above the apex very small ; termen with dark dashes
intersected with white intemeurally.
Genitalia : these organs are very lai^e indeed when compared
with those of the rest of the genus ; the clasps an long, very broad
at the base, but taper rapidly to a narrowish rounded extremity;
the cingulum is of moderate and even width ; the tcgumen rather
email and less excised proportionately; the falces are highly
developed, being deeply curved and longish, but of only moderate
vidth; the aedoeagns is of moderate size, brood and slightly
increostng in width to the zone, from whence it gradually tapers
for two-thirds of the remaining length; the apical third tapering
very rapidly to a sharp point; the tectorius is ample and well
developed, being somewhat of tho theophrastiis type, but much
more ample basally. The androconia are very similar to those in
iMeophraatus, but somewhat larger altogether, they have thirteen
rows of lamina that are sculptured rather more finely than in
Fabricius's species.
,t,7.dh,Googlc
296 Mi. Bethune-Baker on Revision oj the genus Tarncut.
Tanieus IhMpis, L.
Papilio ihespis, Lumaeus, Mug. Ind. Ner., p. 318, N. 136
{1764) ; id. Syat. Nat., p. 791, N. 236 (1767).
TliU species is very close to T. bowkeri, but difiera in
the following particulars. It is of a decidedly brighter
and clearer blue on the upperside, and has quit« narrow
almost linear black borders, the fringes are longer and
more definitely tessellated ; it has not in the secondaries
a terminal row of spots as obtains in bowkeri. On the
underside the predominant colour is brown, not whitish
as in Trimen's insect. The markings are similar, but
decidedly larger and the terminal row of spots, well marked
in bowkeri, is lacking in theapis. In the primaries the fold
and inner marginal area ia entirely brown, except for quite
a small whitish patch at the tornus ; the tail is Uttle more
than a dentition in the fringe.
The 9 ia almost entirely brown above in both wings
with no white areas, as in bowkeri, but with a sUgbt,
restricted, basal blue suffusion, which, in the secondaries,
extends to the inner marginal area; a small white spot
edges exteriorly the spot ckising the cell and has three
or sometimes two white spots beyond it.
It occurs in Cape Colony, where it is widely distributed
over the eastern and western districts, and has also been
recorded from Natal.
The genitalia are of the eame type as bowkerif the cl&sps being
large and long, but easily differentiated. The tegumen is distinctly
dIfFereot; it ia much smaller, with a veiy narrow apex, and very
much smaller and slenderer side lobes, whibt the falcea ore longer
and much more slender; the aedoeagua originates of moderate aiae,
but very rapidly widens up to the zone, where it is suddenly excised
and tapers more gradually to a point, this front portion being half
as long again as the rear part. The vesica is a beautiful object,
its oriRce being elegantly trumpet- shaped, and being very finely
shagrpcned all over; the virgae excitatae are abecnt, but the
t«ctoTius is developed in a modified fortn, Bomewhat as in bowteri.
Explanation of Plates XIV-XX.
[See Explanation facing the Plates.]
l„y,|,AJI^,COOl^lC
( 297 )
X. Notes on some British Guiana H3nnenopt€ra {exclusive
of the Formicidae). By G. E. Bodkin, B.A., Dip.
Agric. (Cantab.), F.Z.8., F.E.S., Government Eco-
nomic Biologist, Department ot Science and Agri-
culture, Britisb Guiana.
(Pnblisbed by permission of the Director of Science And
Agricultare, British GuianB.}
[lUad December 6tb, 1916.]
Plates XXI-XXIII and Sketch Map.
Up to the present time very little has been known of the
habits and life-histories of the Htpnenoptera of Britbh
Guiana. Schomburgk,* in his " Fauna and Flora of British
Guiana," devotes six pages to the Hymenoplera and gives
a few observations with regard to their biology as observed
by himself, but unfortunately many of the scientific
names are quite impossible to trace. Peter Cameron has
published in "Timehri"'!' a comprehensive list of the
Hymenoplara of this country, with descriptions of many
new species, but no biological notes are attached. Scat-
tered references have appeared from time to time in many
scientific publications, but the majority of these are
descriptions of new species.
The present collection in this laboratory' was commenced
in 1911 and has been formed chiefly by myself as oppor-
tunities have occurred. All the commoner species have
now been collected, and in many cases observations made
on their life-histories and habits. The accompanying map
indicates in red dots the areas where collections and
observations have been made. These necessarily follow
* Schomburgk, B., "Fauna und Flora von British Guiana."
Leipsig, 1948.
t Peter Comeran, Hymeaoptcra of the Georgetown Museum,
" Ttmehri." Joumal of the Royal Agricultural and Commercial
Society, 1911-12. Pt. I. Parasitica, I. pp. 153-186 (1911). Pt. 2.
Parasitica, I, pp. 306-330 (1911). Pt. 3. Marabuntaa or Waspa,
II. pp. 207-231 (1912). Pt. 4. Fosaores, 11, pp. 412-440 (1912).
TRANS. KNT. SOC. LOND. 1917. — PARTS U, UI, IV (MAY '18)
iM-,i,A J h, Google
298 Mr. G. E. Bodkin's Nolea on
the chief commuDication routes either by wat«r, road, or
trail. Vast areas consequently remain untouched.
The collection at present contains 1,600 specimens, of
which 161 species are named. All these identilicationB
have been made either by specialists in the British
Museum, through the co-operation of the Imperial Bureau
of Entomology, or by speciaUsts in the U.S. National
Museum.
The present work clearly shows that in scope it can
hardly pretend to do more than outUne the vast field which
awaits entomologbts in this part of the world.
There is a distinct difference between the Ht/menoptera
of the fiat, cultivated and inhabited coast lands and those
of the forest-clad area. Many species of common occur-
rence on the coast lands are never found in the forests,
and vice versa. The climate of these two areas also varies,
the interior districts having a higher rainfall and a some-
what higher temperature. The trade-winds which sweep
the coast lands most of the year are not experienced to
any extent inland. No opportunity has yet occurred
to investigate the large tracts of savannah lands which
exist at the back of the Colony near the Brazilian
frontier.
The observations on the habits and Ufe-histories are
mostly my own, but a number of interesting notes by the
following gentlemen must be acknowledged : Mr. C. B.
Williams, Mr. L. D. CIeaT«, Jnr., Mr. H. W. B. Moore, and
Mr. A. A. Abraham.
I have endeavoured to give as full information as pos-
sible concerning each species. Where only one specimen
of a species has been collected I have given the locality of
collection.
The Ants have not been included in these notes, as
Mr, W. ('. Crawley has already published • an account of
the Family.
Family APIDAE.
Subfamily 8PHEC0DINAB.
Genus TE.MNO^iOMA, Snuth.
T. (leruginasitm. Smith. Issororo, N.W.D.
• " Ania from Rritiah Guiana," W. C. Crawley, B,A Annals
and Magazioe of Natural History, Ser. 8, vol. xvii. May, 1016.
l„y,l,Ajl^,COOl^lC
some British Guiana Hymenojptera. 299
SubfamUy ANDBENINAE.
Genus Augochlora, Smith.
A. graminea, F. 1 Taken while feeding on flowers
A. ihalia. Smith. V of " woiiralli " (fish-poison plant),
A. calypso, Sm. ) lasororo, N.W.D.
SnMamily PANUJtOlNAE.
Genua Meqalopta, Smith.
M. sodalis, Vachal, Ibsoioto, N.W.D. This species ia
frequently attracted to artificial light.
Subfamily XTLOCOPINAB.
Genua Xylocopa, L.
X. fimbriata, F. The commonest wood-boring bee in
British Guiana. The female is black and the male an
ochreoufi yellow with green eyes. The proportion of
females is greater than males — about 4 to 1. Both sexes
may frequently be seen collecting honey from a number
of commonly occurring flowers, especially those of the
large red Hibiscus and the flowers of the Pigeon Pea.
It possesses a powerful sting, and the flight is accompanied
by a loud buzzing sound. I have never observed the
species in the interior. Any Ary decaying wood is utilised
for nesting purposes, such aa rotten paling-posts or tree-
stumpe. The softer kinds of wood are preferred. I^ogs of
wood infested by these bees soon become literally riddled
with their borings, and large quantities of frass may be
seen piled up at the foot of the log. If such a log is sharply
tapped a shrill buzzing noise may be heard caused by the
contained bees. A log of wood 5 feet long by 2 feet in
diameter on being split open was found to contain 20
imagos and 25-30 larvae and pupae. There were about
three entrance holes, and these led directly into galleries
bored at right angles to the grain of the wood. In such
galleries the cells are formed, usually three or four together,
never more. Each cell is about an inch in length and
about I inch in diameter. The cells are separated from
one another by a partition or " wad " of sawdust cemented
together and hardened by the bee. These partitions are
\ inch in thickness. The galleries and cells are perfectly
smooth and very neatly executed. Each cell is stocked
with a small, fairly solid mass consisting apparently of a
mixture of honey and pollen. It is a dark yellow in
u,y,l,/eJh,G00glc
300 Mr. G. E. Bodkio's NoUsjm
colour with a peculiar though not unpleasant odour.
These masses of foodstuff weigh about 25 grammes, and
an equal quantity is deposited in each celt. On this mass
the egg is finally deposited and gradually incieasea in size
as development proceeds; later, the segmentation of the
future larva may dimly be perceived through the envelop-
ing shell. It is sausage-shaped, slightly curved, and almost
transparent in the earlier stages, I^ength I'Tcm., diameter
0"4 cm. During development one end becomes somewhat
larger than the other. Eventually the extremely thin
pelhcule strips off and the wrinkled larva is exposed,
which starts feeding immediately. The larval excrement
is hard, and formed in short rods black in colour. The
full-grown larva measures 39 cm. in length, and is creamy
white in colour. The larval stage lasts about three weeks,
(iiradually the outlines of the pupa may be seen through
the larval skin, which eventually peels off. The period
between the full-fed larva and the final stripping of the
larval skin is 48 hours. The pupa is at first creamy white,
and in form roughly resembles the future adult insect.
Length of pupa 2*5 cm., breadth Vi cm. Within the firat
week of pupation a general darkening of colour takes
place, the eyes going almost black ; hardening of the
mtegument occurs simultaneously. These two processes
progress rapidly till the perfect insect is formed in about
3 weeks. It then makes its exit from the cell by gnawing
away the wad of hardened sawdust.
X. barbala, F. A fairly common species on the coast
lands.
X. brasHwnorum, F. A not uncommon species within
the forest area. Rockstone, Essequebo River, and H.M.
Penal Settlement, Mazaruni River.
X. aurulenta, F. An micommon species within the forest
area. Rockstone, Essequebo River,
SuMaraUy PR080PID1NAE.
Genus Caupoucana, Spinola.
C. eximia. Smith, Essequebo River, in vicinity of
SubfamUy ANTHOPHOSINAE.
Genus Eucera, Scopoli,
E.fesliva, Sm. Vicinity of Georgetown.
u,y,l,7rJM,G00glc
s&me Britiah Guiana Hymenoptera. 301
Genus Exomalofsis, Spioola.
E. globosa, F. Botauic Gaidens, Georgetown.
Genus Epicharir, Klug.
E. rttslica, Oliv. Rockstone, Eseequebo River.
Genus Mbutoma, Latr.
M. englossoides, Lep. From cultivated Cotton blosBoms,
Georectown. This bee has a curious habit of clinging to
the edges of blades of Para grass, with its mandibles finnly
embedded in the tissues. The reason for this is not
apparent. They remain quite motionlesa in this position,
and at times may be seen in considerable numbers. Fairly
common about coast lands.
Genus Centris, F.
C. longimana, Lep. A common species both on the
coast lands and in the interior. It is attracted to strong-
smelling substances such as salt fish, and will folluw boats
with this substance on board for long distances up the
rivers. Frequently seen about buoys moored in the centre
of the tidal passages in the big rivers and elsewhere. It
is apparently attracted there by the excreta dropped by
sea-birds on these objects. It has a swift, noisy flight.
C. tanipes, F. Fairly common on the coast lands and in
the iatenor. Has been taken at flowers and also in the
act of collecting soft mud from a pathway.
C. tersicdor, F. Taken on one occasion while atlracted
to artificial light. Appears to occur only in the interior.
C. personata, Sm. It has similar habits to V. longimatKi,
I«p. Taken under similar conditions in the North-west
District.
C. Utbrosa, Friese. An uncommon species. H.M. Penal
Settlement, Mazaruni River.
Genus Tbygater, Hohnb.
T. rubrifola, Sra. Rockstone, Essequebo River,
Genus Euglossa, Latr.
E. dimidiaia, F. A fairly common species in most
parts of the Colony. Observed on several occasions to
collect mud from a pathway, A somewhat clumsy insect,
TRAHS. ENT. SOC. LOND, 1918. — PARTS U, ni, IV. (MAY '18) X
iM-,i,A J h, Google
302 Mr. G. E. Bodkm'a Notes on
easy to capture, but when alarmed assumes a rapid flight.
Frequently observed Aying about and alighting upon the
bark of certain trees, though on closely inspecting the bark
no feature which might attract the bee conld be observed.
E. cor data, F. The commonest Euglossa in British
Guiana. Observed in all areas visited. It will construct
its somewhat sticky nest in all sorts of curious places,
such as the inside of a disused reel of cotton, interior of
empty cartridge case, eye-piece of polariscope, keyholes,
small cavities in timber, etc. It also takes over the
disused celb of Scdipkron fislulare, Dahlb. In the forest
areas I have observed thb bee to construct its nest on the
under surface of a leaf. The nest varies in the number
of its cells, some only containing two or three, others as
many as six. The cells are roughly about 1 cm. in length
and about 5 mm. in breadth, with the ends neatly rounded.
Each cell is stored with a viscid mixture of honey and
pollen, on which the larva feeds. The waifs of the cells
are thin and soft, being constructed of some dark vegetable
substance. The whole exterior is sticky. This insect has
an extremely rapid, darting flight, and is particularly fond
of the white, sweet -smelling flowers of a certain species of
prickly wild Solanaceous plant. While the bee is within
the white blossoms it emits an exceedingly shrill buzzing
sound, which is often difficult to locate.
E. surinamensis, L. Another very common species of
Euglossa throughout British Guiaua. It is greatly attracted
to the flowers of the same Solanaceous plant as attracts
E. cordala, L. Its loud buzzing and somewhat slow flight
makes its presence conspicuous. The nest is constructed
sometimes in artificial holes in timber, in cavities in the
trunks of trees, and at times on the t^p of roof-beams.
The substance of the nest is thin flakes of bark, or some-
times flakes of white plaster from houses, exceedingly
strongly cemented together. Several cells are usually
formed united in an irregular mass. Each cell is stored
up with a quantity of honey and pollen of moderately
hard consistency. The cells are about 15 cm. in length,
with a diameter of about 1 cm., ovoid in shape, with a
perfectly smooth lining.
E. nigrita, Lep, Not a particulariy common species.
So far only observed on the coast lands. On one occasion
the bees were observed to be nesting within a hollow beam
in the large dining-hall of one of the largest hotels ia
l„y,l,Ajh,COOl^lc
some British Guiatui Hymenoptera. 303
Georgetown. The bees passed to and fro, apparently quite
Tega^ess of the proximity of human beings.
E. pUiventris, Gu^r. An micommon species from Upper
f^ssequebo River. Attracted to flowers of the wild Sola-
naceous plant previously mentioned,
£. analis, Lep. An uncommon species. A large nest
consisting of many ovoid cells was once taken from the
soil on the East Coast of Demerani. The cells were hard,
dark in colour, and joined together. Only one bee hatched
from this nest.
E. caycnnensia, Lep. {= fandata, Jjep.). A fairly common
and widely distribute species throughout the Colony.
Nesting habits not observed.
E. decorata, Sm. An uncommon species from the interior.
Essequebo River.
Genus Exaerete, Hoffm.
E. smaragdina, Gu4r. A conunon species in some parts
of the interior, especially the N.W.D. All of my specimens
were collected while flying about piles of cordwood from
which a strong-smelling sap was exuding, and on which
the bees were feeding. Occasionally seen on the coast
lands and in the Botanic Gardens, Geoi^etown. Nesting
habits unobserved.
Subfamily MEQACBILINAB.
Genus Meuachile, Latr.
M. Idbiiarsis, Smith. An uncommon species with a wide
distribution,
M, Uinata, F. This insect has only once been collected,
and that beneath the Government laboratory, George-
town, The mud cells were situated within a disused
J-inch iron pipe. The nest was cylindrical in shape and
contained about four cells. The nest was very firmly
cemented together, making the whole structure exceed-
ingly strong and hard to break. This bee is well known
in India.*
SubfamUy COBLIOXTNAS.
Genus Coelioxys, Latreille.
C. siviiUima, Smith. Apparently a widely distributed
though not common species. Nest observed in cyhndrical
borings in a wooden post.
* Lefroy, H. Maxwell, " Indian Insect Life," p. 219.
ih,Cooi^lc
304 Mi. G. E. Bodkin's NoUs on
SnblwoUy BOMBINAE.
Genus Bombus, Latr.
B. cay^nensis, F. A common species found only in the
interior.
Genus Melipona, 111.
M. inlerrupa, Latr, This species ia widely distributed
and common. The specimens in the collection were all
t&ken while they were collecting soft mud from paths.
M: pallida, Latr. A fairly common species in the
interior, A small neat on one occasion observed in a
rotten log of timber with a small circular hole formed of
wax for exit. When the nest is disturbed they swarm
out and attack by biting the exposed parts of the head
and neck, emitting at the same time a shrill buzeing
sound. These small bees possess a peculiar odour which
ia characteristic.
M. favosa, F. A common species on the coaat lands,
where it is known as the Courida Bee, from a supposed
habit of collecting honey from the flowers of the maritime
Courida {Awennia nitida). The nests are found in hollow
trees, etc., especially in old Courida trees, but they have
been found in disused drain-pipes and other unlikely
places. This bee possesses no sting, and I have never
observed it to attack in any way; when the nest is dis-
turbed the bees swarm out, but do not demonstrate their
resentment in any more practical manner. This bee is
frequently domesticated, and when the nests are found in
the field they are removed, taken home, and put in wooden
boxes with a small exit hole. They thrive in captivity.
The honey which they produce is thin and of a somewhat
insipid flavour; mixed with other ingredients it is utilised
by the natives as a cure for cold in the throat or cheat.
The honey is stored in egg-shaped cells constructed of wax,
about lJ-2 inches in length; similar cells are also con-
structe<i containing nothing but wax. The cells contain-
ing the larvae are 8 mm. in length and about 5 mm. in
breadth.
M. dacipes. F. A common species in most parts of
the Colony, The nests are usually constructed in hollow
decaying logs. The following notes on the nesting habits
of this species were made from a nest which was found in
a hollow log of Tr>-sil wood (Penladetkra filamattoaa).
The extreme length of the hollowed-out portion containing
Ij.y.l.Ajl^yCOO'^lL'
some British Guiana Bymenoplera. 305
the nest was 32 inchea. Three distinct layers of various
kinds of cells were observed on splitting open the log
longitudinally. The layer next to the entrance consisted
of wax cells about \ inch in length and J inch in diameter;
ovate in shape. The length of this layer was 8 inches.
These wax cells were yellowish in colour and contained
solid wax. To the taste the wax had an exceedingly bitter
Savour, and a sour smell which seemed to pervade the
whole neat. The layer next to the wax cells contained
honey cells; tbis layer was 9 inchea in length. The cells
themselves were slightly smaller than the wax cells, same
shape and dark brown in colour. The contained honey
was thick, very sweet and possessed quite a good flavour,
but if the slightest trace of the wax coating of the cell
got mixed with the honey the bitterness of the wax entirely
obliterated the pleasant flavour of the honey. The next
layer consisted of the cells with embryonic bees in various
stages of development; it was 9 inches in length. These
cells were about \ inch in length and dull yellow in colour.
In shape they were cylindrical. Apparently the nest was
entirely enclosed, with the exception of the eat. This
exit was by no means a conspicuous object, as it was
constructed of wax much the same colour and texture as
the surrounding bark. It was slightly raised above the
surface of the bark, and roughly cone-shaped with a very
small exit hole. Attention was drawn to the presence of
this nest by the bees hovering around the exit. Thb
species is by no means so pugnacious as some of the other
species of Melipona.
M. recurva, Sm. An imcommon species from the N.W.
District.
M. lin&ila, 1/ep. Bartica, Essequebo River.
M. jiavij>enni3, Sm. Taken on one occasion while
attending flowers of Guava.
M. varia, Jjcp. A nest of this species observed in a
large greenheart beam supporting the hotel at Rockstone,
Essequebo River. A non-pugnacious species. The entrance
to the nest is funnel shaped and constructed of wax,
M. guianae, Ckll. A most pugnacious species. If the
nest is only slightly disturbed the bees swarm out and
attack the intruder by biting the exposed parts of the
head and neck, at the same time emitting a shrill buzzing.
The nests (which are large) are usually constructed in the
branches of trees at some distance from the ground. The
u,y,i,Ajh,Cooglc
306 Mr. G. E. Bodkin's Nc^es m
bees posBCBs a peculiar smell, due to the character of the
substance which they collect on their hind-legs. A lai^e
nest observed on one occasion on the branches of a Pimento
tree {Pimenta ^cinalis).
M. rufiventris, Lep., var. Jlavolineala, Friese. Tumatu-
mari, Essequebo River.
Mdipcma amaUhea, F. A common and widely distributed
species. May be observed on the blossoms of most garden
plants, and is particularly fond of feeding on overripe
fruit. It may also frequently be seen collectint; mud
from damp paths, creeks, etc. Known locally as " Tarbaby
bees." It has a habit of collecting the scrap-rubber from
recently tapped trees of Hevea brasili^isis.
M. mfivenlris, Lep, Rockstone, Essequebo River.
M. daliatorreana, Friese. East Coast, Demerara.
M. mulata, Lep. Upper Demerara River.
Genus Api.s, Ijinn.
A. mfHifera, L. Only a small number of hives of the
domestic bee are kept in the Colony, and these are prin-
cipally owned by the Chinese and Portuguese. The honey
produced is of good quality and very sweet. Fresh stock
IS usually imported from the United States. They are
mostly " Italian bees." The Wax Moth (GaUeria mdlonetta,
L.) is common and causes much damage.
Family VESPIDAE.
Subfamily VESPINAE.
Genus Polistes, Latr.
P. pacificus, F. Essequebo Coast, An uncommon
species.
P. analis, F. A fairly common species in the interior.
P. versicdoT, OUv, A common species on the coast
lands.
P. goddii, Ducke. A rare species in the interior.
P. canadensis, Tj., var, amazonicvs, Schulz. The com-
monest species of Pdistes in the Colony, occurring every-
where. Unless severely molested it rarely attacks people,
but its sting is both powerful and painful, and causes a
severe swelling. lian^e nesta are rarely seen, the usual
number of cells being about twenty. Beneath bridges,
homes, on the rafters, under the eaves, beneath the plat-
form of railway stations, behind pictures, etc., are favourite
ih,Cooi^lc
sfime British Guiana Htfmenqplera. 307
nesting-places for this species. It appears to prefer
domestic habitations for its nesting- places. The short
wooden bridges which spaa the navigation trenches on
sugar estates are always thickly infested beneath with the
nests of these insects, and passing beneath such bridges
in a small boat is always an exciting and quite occasionally
ft painful experience. In dwelling-houses they are always
a source of danger, especially when children are about.
The local name is " marabunta." Destroying marabunta
nests with a wad of dried palm leaves attached to a long
pole soaked in kerosene and ignited is an interesting
operation for an onlooker at a respectful distance. At
times these wasps will remain quite motionless in an alert
position OD their nests for hours together, as though on
guard. From the economic standpoint they are exceed-
ingly useful, for they may often be seen hunting for and
consuming the larvae of various agricultural peats, espe-
cially the Rice Worm (Lapkt/gma frugiperda, S. & A.).
The nests are irregular in structure and not strongly made,
for pieces are frequently falling from the nest. The flight
of this insect is somewhat clumsy.
P. crinitus, Feltou. An uncommon species on the coast
lands.
Genus Polybia, Lep.
P. fulvofasciala, de G, (= phlhiska. V.). A common
species on the coast lands ; the nests are frequently found
attached to the under surfaces of leaves, especially those
of the mango tree. Large nests are seldom encountered.
P. occidentalis, Oliv. An exceedingly common species
throughout the coast lands, and at times encountered in
the interior. The nests are found attached to the under
surfaces of many species of palms. The nests are never
lai^e, more or less circular in shajw, and constructed of
exceedingly light and fragile material. This small wasp
is not unduly pugnacious, and seldom attacks unless the
nest is damaged; it is thus frequently encountered when
felling the branches of young coconut palms. The sting
has no great lasting effects, but the first shock is exceed-
ingly painful. On one occasion a gardener who was
trimming a hedge of Barbados Cherry (Malphigia glabra)
brought into the laboratory a good-sized nest of this
species which he had very carefully removed with some
of the wasps in sUn on the outside of the nest. The local
iM-,i,A J h, Google
308 Mr. G. E. Bodkm'a NoUs on
name is " honey marabunta," A native method of
destroying such nests is to seize the nest quickly and
firmly with both hands and then crush it. An individual
possessed of large and tbick-akinned hands is likely to be
the most successful.
P.faslidiomscula, Sauss., var. sampaioi, Bucke. Appears
to take the place of the foregoing in the interior, where it
is common. It has never been taken on the coast lands.
The nest is often met with attached to the under surfaces
of palm leaves.
P. fasei/Ua, Lep. A species by no means of infrequent
occurrence both on the coast lands and interior. A email
nest was taken on one occasion attached to the floor-boards
beneath a house ; the structure is somewhat peculiar
(see photo). Length about 2^ inches; length of single
cell I inch.
P. fxtscicomis, I^ep. A rare species from the interior.
P. chrysoOiorax, Web. A fairly common species only
encountered on the coast lands. Large pendant nests nearly
a foot in length and roughly cylindrical are constructed ;
the one actually observed was attached to a bush only a
short distance from the ground. This nest was imfor-
tunatejy destroyed in an attempt to secure it.
P. diniidiata, Oliv. Rockstone, Essequebo River.
P. conslTuclrix, Sauss. Tumatumari, Essequebo Eiver.
P.jurinei, Sauss. Isaororo, N.W.D.
P. seric&i, Oliv. Tuniatimiari, Essequebo River.
P. sycophanta, Gribodo, An uncoDUUon species with a
wide distribution.
P. vdulina, Ducke. Issororo, N.W.D.
P. TfQecta, F. A species with a wide distribution, but
uncommon.
P. obidensis, Ducke. Puruni River.
Genus Protopolybia, Ducke.
P. labariosa, Sauss, Rockstone, Essequebo River.
Genus Mbtapolybia, Ducke.
M. pediculata, Sauss. A fairly common species which
builds its nests on beams beneath bouses. The nest,
which is constructed of papery material, is usually about
5 inches in diameter and about 1 inch in thickness und
of a flattened irregular appearance. It is by no means a
l„y,l,Ajh,COOl^lL'
softie British Guiana HymaiojAera. 309
conspicuoiis object, and is often of much the same colour
AS the beam to which it is attached. This insect shows
no hostile tendencies, and the taking of the entire nest is
quite a safe undertaking. It has only been observed in
the interior.
Genus Charterou-s, I^peletier.
C. chartarius, Oliv. A fairly common species in tbe
interior. The nest of this species is a well-known object
and has been previously described by a number of authois.
They are prized by many colonists as " curios," and fetch
a very fair price in Geoi^etown. These nests may some-
times be seen attached to the branches of trees overhang-
ing the river. The taking of the neats is by no means an
easy matter, as the insects resent any interference with
their home. Nests, however, which overhang the river
may be taken by getting a native to ascend quietly the
particular tree to which the nest is attached and with one
well-directed blow of a sharp cutlass sever the branch
bearing the nest so that it falls into the river, whence it
is eventually retrieved as soon as the wasps have left.
An unsuccessful blow, however, spells disaster, and there
is a case on record where such an incident occurred, and
the unfortunate native rather than suffer jumped into the
river many feet below and thus evaded the infuriated
wasps.
Genus Chartercinus, Fox.
C paUiAHineaius, Cameron. Issororo, N.W.D.
Genus Nectarina, Shuckard.
N, hilineolala. Spin., var. moinana, Sauas. A species so
far only taken in the Botanic Gardens, Georgetown. The
nest was found attached to the end of a dried " arrow "
of sugar-cane. Greatest diameter about IJ inches — depth
3 inch.
N. satteUuris, F. Issororo, N.W.D.
N. lecheffuana, Latr. Issororo, N.W.D.
Genus Synoeca, Saussure.
S. suriTtama, h. A common species on the coast lands
and occurs at times in the interior. The nests of this
species, which are irregular in shape, may usually be seen
attached to trees in the Botanic Gardens, Georgetown.
ihyCOOt^lC
310 Mr. G. E. Bodkin's Notes on
The whole of one Bide of the nest is attached to the tree,
and the exi«mal wall exhibits a slight " ribbing." The
nests are often a foot or more in length. The sting of this
. species is particularly formidable, and it does not hesitate
to use it when occasion arises. The adult wasps may
often be seen feeding on decaying fruit which has fallen
on the ground, and they frequently visit certain spedes
of flowers. I have taken specimens of this insect with
the " polhnia " of a species of Euphorbiaceous plant
attached to its legs; this is by no means a common
occurrence.
8. irina. Spin. An uncommon species in the interior.
Genus Apoica, Lepeletier.
A. pallida, Oliv. A common species on the coast lands.
The nest, which is invariably attached to a tree and never
far from the ground, ia disc-shaped. The under surface
consists of innumerable cells, the ends of which are exposed.
The under surface is invariably crowded with adult wasps,
which attach themselves each to a particular cell and re-
main motionless, an aspect is thus presented of a cluster of
wasps adhering to the under surface of the nest ; in this
position the bright yellow tips of their abdomens are very
conspicuous. They do not readily move from this position,
and if slightly disturbed will only make a slight move-
ment. Once while observing a large nest of this species
I happened to approach rather too near, when one wasp
left the nest and stung me on the forehead immediately
between the eyes, and returned at once to its original posi-
tion. The effect of the sting was as though a blow had
been delivered, and in a short space of time a swelling the
size of a fowl's egg made its appearance, accompanied by
considerable pain. The species is readily attracted to
artificial light.
A. pollens, F. Issororo, N.W.D,
Genus Mischocyttahus, Saussure.
M. labialtis, F. A fairly common species on the coast
lands. Only small nests are constructed.
Subfamily EUMENIDINAB.
Genus Montezumia, Sauss.
M. leprieiiri, Spin. (1841) {= M. rodwayi. Cam. (1911).
An uncommon species taken in the vicinity of Georgetown,
n,g,i,.eih,.G00glc
same British Guiana Hymeno^era. 311
M. nigriceps. Spin. One of the commonest species of
Htftnem^era on the coast lands. Also a species of par-
ticular economic value, as it hunts for and destroys the
larvae of many agricultural pesta, including the Rice Worm
(Laphygma frvgiperda, S. & A.) and the Para-grass Worm
(Mods repanda, F.). This wasp may frequently be seen
Bearchtng for its prey amongst grass and tall rice. A large
pendant, irregularly -shaped nest is formed, which is some-
what fragile in its structure. These nests contain a large
and active community.
M. infemalis. Spin. An uncommon species on the coast
lands. According to C. B. WiUiams, who has observed
the habits of this wasp, it constructs burrows in the clay
banks of the canals or trenches to be found on all sugar
estates. Green caterpillars are stored up in the nest, and
an egg is deposited, which is hung from the roof of the
burrow by a long stalk. The opening of the burrow ia
small in comparison with the size of the wasp.
M. infundHndiformis, F. Issororo, N.W.D.
Genus Eumekes, Latreille.
E. canaliadata, Oliv. An exceedingly common species
to be met with mostly on the coast lands. Its dome-
shaped mud cells, in small colonies of live or six and some-
times more, are objects of common observation attached
to the rafters beneath houses and in sheltered spots on
walls and palings. These mud cells (see photo) are usually
about \ inch in height and with a diameter of about \ inch.
Some cells possess a kind of " neck " at the top of the
cell, giving it the appearance of a squat-shaped earthen
bottle. These cells are stored with Lepidopterous larvae
usually slightly over an inch in length and, of course,
paralysed by the sting of the wasp at the time of capture.
From four to six such larvae are stored. The wasp larva
becomes mature in ten days to two weeks from the time
of emergence from the egg. The pupal stage lasts shghtly
over a week. The adult wasp emerges from the cell by
biting a circular hole in one side. These empty cells are
afterwards utilised by a species of Trj/patylon for its nest,
and stored with spiders. A small species of black ant
(Cremaetogaster, sp.) finally colonises these disused cells.
In constructing such cells the adult wasp holds the piece
of moist and plastic mud in position on the half- constructed
nest by means of the two front pairs of legs, and the deli-
iM-,i,A J h, Google
812 Mr. G. E. Bodkin's Notes on
cate process of moulding the mud around the edge of the
nest 18 performed with the mandibles, which are long and
well suited to the work. During the operation the long
antennae are bent downwards and kept rapidly moving
about the work as though guiding it and preserving the
symmetry of the structure. A small species of Chrysid
was bred on one occasion from the mud cells of this
Eumenes.
E. calUmorpha, Sauss, Vicinity of Georgetown.
Genus Pachymenes, Sauss.
P. pallipes, Oiiv. Ondemeeming, Essequebo.
Genus Zethus, Fab.
Z. mexicanus, L., var. hiffubris, Perty, An imcommon
species on the coast lands.
Z. gigaa, Spin. Issororo, N.W.D.
Z. sichdianus, Sauss. Inhabiting disused borings in
timber, Courantyne Coast, Berbice.
Genus Odynerus, Latr.
0, nasidens, Latr. Courantyne Coast, Berbice.
O. davilineatus, Cameron. An uncommon species in the
interior.
FOSSORES.
Family MUTILLIDAE
Genus Mutilla, L.
M. (ThaumalmmUilUi) parallela, Khig. A fairly common
species on the sandy soils of the interior.
M. mediata, F. From Courant\-ne Coast, Berbice. Taken
while issuing from some disused borings in timber.
Genus Thaumatomutilla, Andr6.
T. ocellaTis, Klug. Tumatumari, Essequebo River.
Family SCOLIADAE.
Subfamily TIPHIINAE.
Genus Tiphia, Fab.
T. paralkla. Smith. The larva of this wasp is parasitic
on the larva of Dysdnelus indeniatus, Burm. (Coieoptera) ;
fairly common on the coast lands.
l„y,i,AJL.,COO'^lC
8<mte Britith Guiana Bymeno^era. 313
Genus Dielis, Sausa.
D. doTsata, P. Of common occurrence in most parts of
the Colony.
D. hyalina, Lep. (= D./altax, Sauss.). Fairly commoo
in most parts.
D, vari^ata, P. Issororo, N.W.D,
Genus Eus, F.
E. Jlavopicta, Smith. Turkeyn, East Coast, Demerara.
Family RHOPALOSOMIDAE.
Genus Ruofalosoha, Schulz.
R. gutanertfe, Schulz. At light, Rockstone, Essequebo
Biver.
Family PSAMMOCHARIDAE.
Genus Pskudaoenia, Kohl.
P. cmnparala, Sm. Tumatumari, Essequebo River.
P. chiorosoma, Sm. Punmi River.
SnbfttmUy PEPSIN AE.
Genus Cryptochilus, Panz.
C. puTpuT^pes. A common species on the coast lands.
Genus Pepsis, Fab.
P. tinctipennis, Smith. Issororo, N.W.D.
P. sappkiria, P. de B. Rockstone, Essequebo River.
P. jucunda, Mocs. Rockstone, Essequebo River.
P. chlorolica, Mocs. Ondemeeminp, Essequebo.
P. nigreacens, Smith. Rockstone, Essequebo River.
P. dimidiata, F. Rockstone, Essequebo River.
P. ametkystina, F. A common species in moat parts of
the Colony.
P. aeladonica, Dahlb. Issororo, N.\V.D.
Subfanuly PSAMMOCHABINAE.
Genus Pompilogaster, Ashm.
P. philaddphka, Lep. A common species on the coast
n,g,t,7.dh,G00glc
' 311 Mr. 6. E. Bodkin's Notes on
Snbfanulr SPBECISAE.
GeDUS SCELIPHEON, Klug.
S. fislulare, Dablb. This species is widely distributed.
Its mud nests are common objects on palings, beneath
houses, and at times within the bouse iteelf, behind pic-
tures, etc. The nesta are somewhat irregular in shape and
may contain as many as eight or ten cells. Sometimes
only two cells are constructed together, when the archi-
tecture is then better demonstrated. The usual procedure
is to build one cell and attach it firmly, and then to build
other cells around it. Each cell is stored with spiders,
from sixteen to twenty according to the size of the spiders.
As soon as the requisite number of spiders has been obtained
the cell is closed up. The egg, according to C. B. WiUiams,
is deposited on dorsal surface of the abdomen of the first
spider placed in the ceil. The following observations by
Mr. Williams on the habits of this wasp are of interest.
A wasp which was engaged in constructing its mud cells
was observed, and its movements recorded as follows—
12.5' 30" p.m. gone for mud,
12.9' 10" p.m. returned,
12.10' 15* p.m. gone,
12.17' 58' p.m. returned,
12.19' 20' p.m. gone again,
12.22' 20' p.m. returned,
12.23' 10' p.m. gone.
At 12.37' 45' the wa.ip entered its nest and remained
inside; 12.38' 50' the wasp flew away, and on inspection
the neat was found to contain a spider (Fam. Argiojndae —
Ga-ileracatUhinae) with an egg on dorsal surface of abdomen.
This spider was eventually removed by myself. At
12.51' 15' the wasp returned with another spider, and
went away 12.02' 15'. At 4.10 p.m. the nest was almost
completely closed. The egg of this species is sausage-
shaped and slightly curved, dull yellowish white in colour,
with small almost transparent areas towards each end;
length 3'4 mm. Young larvae when first hatched out are
almost 4 mm. in length. They soon attach themselves
to a spider, which they proceed to consume, leaving only
the legs. Development is completed in about two weeks.
The larva then commences to spin its cocoon, which
occupies several days. The cocoon is dark brown in
l„y,l,Ajh,COOl^lC
some British Guiana Hymenoptem. 315
colour, and of a papery consistency, easily broken. Pupa-
tion lasts a little over a week, and the adult wasp then bites
its way out through the mud wall of the nest, leaving a
neat circular hole. The small green metallic bee, Euglossa
cordata, L., often utiUses these disused cells for its nest.
A fungus occasionally destroys the stored spiders and
finally the young larvae. Ko actual parasite of the species
has been observed up to the present. It appears, however,
that only about 60 % of the stored celb produce an adult
wasp,
S. figvlus, Dahlb. Closely allied to the above species,
and with similar habita and distribution, though not so
numerous.
Genus Sphex, L.
S. ichneumoneus, L. A common and widely distributed
species throughout the Colony. For its subterranean nests
it usually chooses a dry piece of soil, often in the centre
of a path or other places where people are frequently
passing. While constructing the initial excavation for its
nest it is a most conspicuous object, moving rapidly about
on the surface of the soil. The following observations
made by Mr. C. B. WilUams are of interest. " In this
instance the insect was boring into soft pegass soil on
swampy land at the foot of a hill. High spring tides
from the river invariably cover the land. TTie burrow
was commenced about 1.40 p.m., and at 1.45 p.m. it was
about ^ inch deep. At 2.19 the wasp entered the hole
backwards, remaining below a few minutes, and then
came out again. Burrowing continued till 4 p.m. These
nests are stored with four or five short-winged grass-
hoppers, either nymphs or brachypterous species; the
neat is not closed up between the insertion of each
Sphex (Isodonlia) coslipatnis. Spin. Issororo, N.W.D,
S. {Harpaclopus) ihomae, F. An uncommon species
which does not occur on the coast lands. Constructs
burrows in soil and stores them with a single grasshopper,
which is placed head first in the nest. The egg is attached
at the base of the grasshopper's hind-leg (C, B, Williams).
Genus Podium, F,
P. ^tUiculalum, Sm. An uncommon though widely
distributed species.
n,g,i,.rjh,G00glc
316 Mr. G. E. Bodkhi'a NaUs on
SnblamUy PHILANTHISAB.
Genus Cerceris, Latt,
C. [Trackyjms) mexicanvs, Cameron. Aii
species. Taken while emeri^ting from its burrow in sandy
soil at the base of a tree. Plantation Bath, Berbice.
Subfanuly BEMBICINAE.
Genus Bembidula, Burm.
B. dtsdsa, Tasch. Upper Demerara River.
B. variegala, 01. Upper Demerara River.
Genus Monedui^, Latr.
M. pantherina, Handl. A fairly common species in some
parts of the interior, especially where sandy soils exist.
It is quite the largest and fiercest of these insects to
be found in the Colony. Its burrows are stored with
Tabanidae.
M. 'punctata, Lep. An uncommon species in the interior.
M. signata, Latr, The commonest and most widely
distributed Bembex in British Guiana. It may frequently
be seen about mules and cattle in the pastures waitin)^ to
carry off the Tabanidae which are always present about
these animals. Their burrows may be seen iu sandy soil,
and usually occur in colonies of twenty or thirty. A
large species of Bombyliid fly may be seen about these
burrows, occasionally entering them. Its exact relation-
ship to the wasp has never been actually traced, hut it
is quite possibly parasitic. The fly belongs to the genua
Anthrax.
Subfsauly OOBYTINAE.
Genus Gorytes, Latr.
G. brasiliensis, Shuck, Rockstone, I^quebo River.
Subfamily LASSINAE.
Genus Tachytes, Panzer.
T. amelina, Cameron. Rockstone, Essequebo River,
Genus Larra, Fab.
L. rubrkata. Smith, Rockstone, Essequebo River.
u,y,i,A J h, Google
some British Quiana Hym^ioptera. 317
Subfamily TRYPOXYLISAS.
Genus Trypoxylon, Latr.
T. paUidUarse, Sauss. A fairly common species on the
coast lands.
T.fugax, F, This insect has been bred from the disnsed
cells of Eumenes camdiculala, 01. It is of fairly common
occurrence.
T. breificarinalitm, Cameron. A common and widely
distributed species all over the Colony, It usually con-
structs its nest of mud cells beneath a palm leaf. Some-
times as many as twenty-five cylindrical cells are found
cemented together. The largest cells are nearly an inch
in length and about i of an inch in diameter. Small
spiders are stored up by this species.
T. albilarse, F. Bred from a large-sized rounded mud
nest. Potaro River.
T. nitidum, Sm. Issororo, N.W.D.
T. cinereum. Cameron, The habits of this species have
been observed by Mr. C. B. Williams. " An elongate mud
cell was constructed by one of these wasps in the angle
of the woodwork of a door. When making these nests a
' ball ' of mud is collected and applied to the side of the
nest, gradually drawing it downwards as it adheres, A
shrill buzzing sound is emitted the whole time. Thirty
seconds to one minute are occupied in applying one ' ball '
of mud. Spiders are stored."
Family TRIGONALIDAE
Genus Trioonalyh, Westw.
T. melanoleuctt, Westwood. A number of these insects
emerged from the pupae of the Noctuid moth Amyna octo,
Guen., which had been taken from just beneath the surface
of the soil on a sugar estate.
Family CHRYSIDIDAE
SubfamUy CHBT8IDINAE.
Genus Chryris, L.
C. pundalissinHi, Spin. A common species on the coast
lands.
C. distinctissima, Dahtb. Bartica, Essequebo River.
C. parvula, F, East (Joast, Deraerara.
TRAN3. ENT. SOC. LOND. 1917. — PARTS 11, III, IV. (mAY '18) Y
iM-,i,A J h, Google
318 Mr. G. E. Bodkin's Notes on
Family BRACONIDAE.
Subfamily OHBLONISAE.
Genus Foknicia, Brull^.
F. dathrata, Brull6. A larval parasite of the Limacodid
moth, Sibinefusctt, Stoll. This is a verj- abundant moth
on the coast lands and attacks a large variety of economic
plants. The parasite emerges before the lar\-a is half
developed. The percent^e of parasitism is never very
high.
Subfamily BSACONINAE.
Genus Lasiophorus, Hal.
L. forttspinus, Cameron. A fairly common species on
the coast lands.
Genus Iphiaolax, Forater.
/. meAianus, Cameron. This is the well-known parasite
of the several species of Dialraea (Lepidoptera) which
bore into sugar-cane in British Guiana. The species is a
common one, it being an easy matter to capture either
sex in the cane-fields. The life-history of the species has
been observed in the field by Mr. Harold Moore, and the
following is his own description : *— " On the mominp of
the 13th, in a field at Non Pareil, I noticed a female alipht
near the top of a cane-stump, run down the shoot, tapping
it the while with her antennae, till she got to a Diairaea
hole near the base. Into the timnel she began pushing
her ovipositor, when a slight movement on my part, in
an endeavour to obtain a closer view of the proceedings,
disturbed her. Off she flew, but merely to wheel round
and ahght again on the shoot, when she found the hole
the same way, and began inserting her ovipositor. I lay
prostrate on the trash bank, as motionless as possible, and
watche<l. It was 9. "28 a.m. Soon after her ovipositor had
been inserted she gave a slight quick backward jerk with
her abdomen, showing she had evidently felt something.
Back down went her abdomen and again a similar jerk.
This occurred three or four times, after which she kept
quite still for several minutes, her ovipositor being well
in the caterpillar's tunnel. She withdrew her ovipositor
at 9.38, thus occupying about ten minutes in parasitising
the caterpillar, of the presence of which I had no doubt.
• Pablbbed in the " Daily Argosy," Demerara, Maroh 1813.
Ij.y.l.AjL.yCOO'^lL'
some Brituh Guiana Bymenorptera. 319
I cut out the cane-shoot and took it home. I could not
examine it till next morning (13th), when I found it con-
tained a caterpillar, and in the same tunnel with it was
the egg of the parasite. It is about 3 mm. bv about 1 mm.
at the widest part — whitish, translucent, firm or leathery^
narrow, elongated, tapering to a long point posteriorly,
where there is a small curved blackish hook. The hook
at the end is evidently for the purpose of keeping the egg
in place. From the above it seems that the caterpillar
is paralysed bv the parasite at the time of oviposition.
From observations made on previous occasions, however,
it would appear tliat paralysis does not, or at least does
not always, occur immediately, but cornea on gradually.
The firm texture of the egg prevents it from being injured
should the caterpillar happen to wriggle against it. At
3.30 p.m. on the same day (13th) I observed that the egg
had hatched and that the young larva was peacefully
feeding, attached to the side of the ninth segment of the
caterpillar. It had probably hatched during the forenoon.
The egg-period is therefore very short — about one day.
On the morning of the sixteenth the caterpillar was dead,
almost the whole posterior half having been devoured by
the parasite larva. By the morning of the 17th the
parasite larva had completely devoured the caterpillar,
and was spinning a cocoon in order to pupate. The
larval period, too, is very short — about four days. The
sbortnesa of both the egg and larval stages can be very
readily understood, when it is remembered that the larva
requires fresh food, and has only a single caterpillar on
which to feed. It of necessity therefore has to get through
it rapidly, before extreme decomposition is reached, and
do all its growing in that short time. The larva pupated
at the angle between the shoot and the glass jar. It left
the tunnel, probably on account of the latter shrinking
through drying of the shoot. From other observations I
know that the larval period is sometimes certainly a week.
On the morning of the Slst the perfect insect emerged.
Pupal period therefore about fourteen days. Time from
egg-laying to adult is therefore about nineteen days."
The cocoon ia transparent and constructed of closely
woven silk, giving a membranous appearance. In shape
cylindrical, blunted at both ends. Colour whitish. Length
2'1 cm., diameter 4 cm.
/. obscuricarinaliis, Cameron. Issororo, N.W.D.
u,y,l,7rJM,G00glc
320 Mi. G. E. Bodkin'B Notes on
SubfamUy RBOOADINAB.
Genus Meoaruogas, Szep.
M.fusdpaljHs, Cam, Rockstone, Essequebo River.
Subfamily AGATHINAE.
GeiiuB Spilomicrodus, Cam.
S, n{gr{ce/)S, Cam. Bartica, Essequebo River.
Family CHAIX^IDIDAE.
SubfamUy TRICHOOEAXMATINAE.
Genua Trichogramma, Westw,
T. miniilum, Riley. This we! I -known parasite is of
common occurrence in the Colony ae an egg-parasite
of the Oane Borers {Diatraea) and several other species of
Lepidoptera. Its life-histor^' and habits when parasitising
the ova of Dialraea have been fully worked out by myself
and published in the Journal of the Board of Agriculture,
British Guiana, vol. vi, No. 4.
Subfamily OHALCIDINAE.
Genua CHAi-ns, F.
C. pandora, Cwfd. Bred from the pupa case of a species
of Hesperid (sp. indet.] butterfly feeding on cane blades.
An uncommon species.
C. annulaUt, F. A common parasite of Brassulis sopkorae,
L. (Lepidoptera) and also Caligo iUoneus ilioneus, Cramer
(Lepidoptera). It is widely distributed and the commonest
Chalcid in the Colony.
Genus Smicra, Spin.
S./vh'Omaciilfila, Cam. Bredfrom the pupa of a Hesperid
butterfly (sp. indet.). An uncommon apecies.
Family EVANIIDAE.
Subfamily BVANIINAE.
Oeims Evania, F.
E. ert/thrajipis, Cameron. Issororo, N.W.D.
E. append>ga,iler, L. This insect emerged from a large
box containing a quantity of seed rice packed in bt^.
Innumerable Blattid egg-caaea (Periplanfia americana L.l,
as well as adult insects, were in the bnz.
ll,y,l,7rJM,G00glc
Tram. Enl. Sot. Land., igij, Plate XXI.
Attdri, Sleigh &■ Anglo, Ltd.
BRITISH GUIANA HYMENOPTERA.
h,Googlc
some British Guiana Hymenopiera. 321
Family ICHNEUMON ID AE, I«ach.
Subfamily OPHIONINAE.
Genus Eiphosoma, Cresaon,
E. aztecum. Cress.? lasororo, N.W.D.
E. texanum. Cam. Botanic Gardens, G«orgetowii.
Genus Henicospilus, Stephens.
H. major, Morley. Attracted to artificial light. Bock-
stone, Essequebo River,
H. nigricomis, Bmll^. Attracted to artificial light.
Bockstone, Essequebo River.
H. fiavoscutellalus, Brull*. Attracted to artificial light.
Bockstone, Essequebo River.
H. maculiceps, Cam. Bred from the pupa-case of a
Noctuid moth, Amyna octo, Guen. An uncommon species
on the coast lands.
Genus Itoplectis.
/, pUUana, Morley.
Subfamily ICHEUMONINAE.
Genus Joppa, F.
J. atUennat(/r, F. Rockstone, iissequebo River.
J. geminata, Vriich. Courantyne Coast, Berbice.
Explanation op Plates XXI-XXIII.
plate xxi.
FlO. 1. Synoeca surinama, L.
2. S<xliphTonjistulaTf,l>eAAh.
3. Xylocopafimbriala, V.^.
4 , F. ^.
5. Mud neKt of Melipona jruuinae. C'kll.
6. Mud ncHt of Trypoiglon breviatriiuilwm. Cam.
7. Eumenes canalicuhta, Oliv., on its mud cell.
Figs, 1, 2, 3. 4. 6 and 7 arc about the natural size. FJg. 6 is
considerably reduced.
PLATE XXII.
Borings of Xyltxopa fimbriala, F,, in softwood. Note the "wad "
of sawdust wbich divides the cells. Considerably if duccd.
PLATE XXIII.
1 fittu!are, Dahlb. Slightly smaller than
n,g,l,.rJM,GOOglC
{ 322 }
XI, On a CoUeclion of BuUerflies taken in East Africa by
Mt. W. a. Lambom. By H. Eltrinqham, M.A,,
D.Sc. Wiih tiotes on the Pierinae, by Dr. F. A.
DixEY, F,R.S., and descripdon of a new form of
P. dardanuB ?, by Prof. E. B. Poultom, r.R.S.
[Read JuDB eih, 1817.]
The Hope Department at Oxford has recently received a
Collection of Lepidttplera, chiefly Rhopalocera, taken in
1916 by Mr. W. A. Lambom in the northeni central part
of what was German East Africa, and it may be not with-
out interest to give particulars of one of the first conaign-
ments forwarded from one of our newly acquired territories.
The dates and localities with notes as to the type of
country are as follows. The elevations are approximate.
May 3-6 New Moahi 37° 24' E. 3° 21' a Thin woodland,
»-20 Kikuletwa-
Darjama R.,
Noisinak
Bridge.
) Loldiloi
Muruangoin,
Sseiije Drift
Lolkissale
. Plain, 2900 ft.
. Dense evetKi*«n
forest, 4550ft.
Tbombush, 3500
36° 50' I
36°42'i
June 1-0, ID (Jlionii (plain)
„ 2-6 Uflomi (wood-
land)
„ 20 Taranjere River, S
A
July 1 New Moshi 3'
„ 10 Tanga-MoHhi
Railway, Sami
TBANS. ENT. 8O0. LOND.
CSe'S. Wooded borders of
river, dry plains
beyond, 3000 It.
J'aS'S. Thorn-bush, 3550
ft.
E. a'eO'S. Thombush, 4-
6000 ft.
E. 4= 16' S. Dried plain. 4440 ft.
' R 4° 16' S. Green river gorge,
woodland, with
large trees.
E. 4''30'S. Thorn-bush and
woodland, 4500
ft.
S. of, Thom-bnsh and
36° E, 4° S. woodUnd, 3800 ft.
E. 3°24'S. Thin woodland.
2925 ft.
37° 46' E. 4° 8' S. Thom-buah, 2900ft.
1917.— PAKTS II, ni, IV. MAY '18
Dr. H. Eltringham on BuUerJlies taketi in East Africa. 323
July H Tanga-Moehi 37° 56' E. 4° 36' S. Thorn-buah.
Railway, Mabirioni,
("Gonnan Bridge")
„ 23 Handeni About 38° E. 6° S. Woodland and
thom-buBh, 2800ft.
Sept. 6 Bagamoyo, 37 m. N. of Dar-e«- Coco pluitation,
Salam, on coast 100ft.
,, 13 Ngerengere, on Central Railway, Thick thora-buah.
100 m. W. of Dar-ea-Sslani.
„ 30 KondutBchi, 10 m. N. of Dar-ee- Coco plantation,
Salam, on coast fiOft.
Nov. 10-14 Tanga 5° S. on coast. Coco plantation,
50 ft.
At the above dates and localities the wet season prevailed
until about June 16, when the dry began. The rain of the
next change was first encountered on September 6.
Dani^d* formoss, Godm.
A nice series in fine condition.
Same, Mabirioni. 9 S<S, 3 ?9. May 16-July 14.
D. petiverana, Doubl.
Ufiomi (woodland). 1 (J. June 4.
D. chryslppus, Linn.
Forty examples, of which only three are typical chry-
sivpus. One approaches ahippoides, Moore, and three are
of the aMnus, Lanz, form. The rest are all dorippus,
Klug.* The remarkable corresponding predominance of
the daira form of Acraea encedon in this collection is
referred to under that species.
All dorippts unless otherwise stated.
Kikuletwa, Darjama River (thorn-bush)— 1 ? Mar. 19;
Ix>ldi]oi (wooded river-side) — 11 S^, 1 cj ckrysippus, 1 S
• In Seitz" " Macrolepidoptera," Profesaor Aurivilliua aeparatea
tbe dorippiig form as a distiiitt species, with the remark that there
ia not BuJficient evidence of its Bpecific identity with chryaippua I
Any good series, hoicever, thntpg many rtmarkaWe inlennedlales,
and sptcimmn in Ihe Hope Department bred by the late t'l/l. Manders
ahoip in a large number of spedmena the in*labiMy of ihe fvre-u-ing
apical black. There can be no doubt that dotippus is a form of
chrysippua jusl as inaiia is a form of misippua
u,y,i,Ajh,Cooglc
324 Dr. H. Eltringham on a
aJbinus May 20; Lolkissale (thom-busb)— I $ May 26;
Same (thom-bush) 4 (J(J, 3 $5, 1 ^ chtysippus July 10;
1 tJ, 1 ?, 1 (J chrysippus, 1 ^ aUtinus Jaly 1 1 ; Mabirioni
(thoro-bush) 6 cJ(J, 5 $9. 1 cJ chrysipjms (appcoacbing
tdcippoides), I $ aibinus July 14.
Amanrli oehha, Boisd.
Thirty examples, a)] typical.
Bagamoyo. 11 (J(J, 19 ?$. Sept. 6.
A. damocles f . damocUdes, Stgr.
Loldiloi. 1 <?. May 20.
A. nlavlus domlnicanus, Trim.
Mabirioni, Bagamoyo. 3 tj<?- July 14-Sept. 6.
A. alblmaculata, Butl.
New Moshi, Saiija R., Ufiomi (plains). 23 ^^, 5 $?.
May 3-June 6.
[Note. — A. eckeria is rare in Kikuyu, common on Kili-
manjaro (Rogers, Trans. Ent. Soc, p. 511, 1908), and
aUnmacuUua relatively very rare at Entebbe.]
SATYRINAE
Gnaphodes parmeno diversa, Butl.
A single example of the southern and eastern race of
6. parmetto.
tJfiomi (woodland). 1 $. June 3.
Neocoenyra duplex, Butl.
A single example, apparently of this species, though
differing from typical examples in having the fore-wing
eye spot area merely dusted with red brown scales, instead
of being distinctly red brown.
Ufiomi (woodland). 1 cJ. June 4.
Physcaeneura leda, Oerst.
Six specimens somewhat variable in size, but otherwise
normal.
Tanga. 5 <?, 1 ?. Nov. 10-14.
n,g,t,7rJM,GOOglC
CoUedion oj BuUerfiies taken in East Africa. 325
ACRAEINAE.
Acrua quirina, Fab.
Five females of the uauat brownish form. The note
accoDipanying these examples describea the focality aa
dense evergreen forest. This may account for the absence
of male specimens, which probably frequent more opeu
situations.
Anisha. 5 ?$. May 16.
A. neobule, Doubl.
Five examples of this widely distributed species, all of
tjrpical pattern and colour.
Utiomi (plains), Muniangoin, Same. 5 ^^. May 22-
July 10.
A. zetes, Linn.
One male of the acara form so greatly modified in re-
semblance to A. fseudolyda aslrigera, Butl., that the male
armature gives the principal evidence of its distinction
from that species. Zetes and pseudolyda are certainly
closely allied, and the resemblance would appear to be due
to affinity rather than to mimetic association.
Kikuletwa-Darjama River. 1 (J. May 19.
A. Inslgnls, Dist.
A single female approaching the siginna form.
Handeni. 1 $. July 23.
A. chila, Godm.
Of the twelve examples received, ten are of the crystaUina
form of female, the remaining two are males of the ordinary
form. It is remarkable that whereas the female chUo
becomes more transparent as it extends southward, the
female braesia, and to some extent its male also, becomes
more transparent towards the northern limit of its range.
The male cnilo, on the other hand, shows some tendency to
heavier spotting as it extends southward. The localities
where the present examples were taken are further south
than any yet recorded.
Same, Mabirionl. 2 SS, 10 52- July lO-l^-
ll,y,l,7rJM,G00glc
326 Dr. H. Eltringham on a
A. egina. Gram.
Ten Bpecimens, three of which are distioctl}' of the foim
harrisoni, Sharpe, wbibt two other males have slight red
streaks in the fore-wing apical area. The harrisoni form
has previously been reported from Bukuba, L. Kivu
district, by Griinbere, who renamed it kivuensts, and
occasional examples nave been noted from Uganda to
JS^yassa Land. It is merely an intermediate between the
type and the form areca, Mab.
Uflomi. 7 (JJ, 3 ??. June 2-19. (One harrisoni J,
June 2, woodland, the rest plains.)
A. acrita manca, Thur.
A fine series of this interesting form. The characteristic
fore-wing subapical spots show considerable variation in
number and size, and in one or two examples they are
present only on the underside. The fore-wing apical black
IS slightly broader and the black rings on hind-wing margin
are more pronounced than in typical tnanca. The exampes
are thus to some extent intermediate between manca and
manca f. Itdtca. Nearly all the females have a brownish
ground-colour, though in one or two examples there is
in the hind-wing a tendency towards the typical Same
orange of the male. The wet season continued till abont
June 16, BO that these females are fairly consistently of
the wet form.
Ufiomi (plains). 19 <?(?, 21 ?$. June 1-6, 19 (wood-
land). 1 (^, 1 $ . June 5; Ssalanga. 2 $. June 9.
A. ealdarena f . neluska, Oberth.
Two examples of this form occur. They present no
unusual features. The form is rather rare in collections.
Handeni. 1 ^, 1 ?. June 23.
A. pudorella padorella, Auriv.
One dwarf male example.
Ufiomi (plauia). 1 J. June 1.
A. braesia braesla, Godm.
In a long series of this species there is one male example
of the regalis form, and several somewhat int«rmediate
thereto. No specimen shows the peculiarities of the
l„y,l,Ajh,COOl^lC
Coaeciion of Butterflies taken in East Africa. 327
Somaliland foim mentioned in my monograph, though
several of the females are more than usually transparent.
Ufiomi (plains), Same, Tanga Moshi Ry., Ssalanga,
Mabiiioni, Handeni. 34 <J(J, 8 ??. June 5-Ju]y 23.
A. equatorlalis, Neave.
A single male example in tine condition, presenting the
coloration of true equaiorialis combined with the larger
size so usually found in equatorialia anaemia, Eltr. Ako
one other specimen of the anaemia form, imfortunately
without data.
Kikuletwa-Darjama R. 1 ^. May 19.
A. natallca natallca, Boisd.
Five examples of this common species all of typical
appearance.
New Moahi, Sanja R. 4 ^(J, 1 ?. May 6-11.
A. uacieon, Trim.
Two examples of this species were taken, one m^Ie and
one female. Though somewhat faded and worn they are
btereating as showing an intermediate condition between
anacreon bomba f . induna. Trim., and anacreon atiacreotUica,
Gr. Sm, As in the latter, the fore-wing apical black is
very much reduced and the outer half of the wing is
ochreous ; the base of fore-wing and the whole hind-wing
is deep orange as in the induna form. They thus support
my contention that anacretm, bmtAa, and anacr&mtica are
all forms of the same species. The resemblance of the
female example to the specimens of acrita nuxTuxt, with which
it was taken, is very striking.
Ssalanga. 1 J. June 9.
Ufiomi (plains). 1 ?. June 19.
A. encedon, Linn.
Of twenty-one examples there is no specimen of true
encedon. The daira form largely predominates, and
flfcippiMo is absent. It is interesting to note that of forty
examples of D. chrysippus in the same collection, three
Ue typical, one is the (dcippus form, three aUnnus, and all
tfe test are of the doripjms form to which enced(m t.
daira corresponds.
Daira. — ^New Moshi (thin woodUnd) 5 (J(J May 6;
iM:,i,A J 1^, Google
328 Dr. H. Eltringham <m a
Sanja Kiver (plains) 2 $? May 10; Arusba (dense forest)
1 $ May 10, 2 9$ May 16 ; Muruangoin (thom-bush) 1 ^
May 22 ; Ufiomi (plains) 1 $ June 2 ; (woodland) 1 $ June
5 ; Mabirioni (thom-busb) 2 S3 Ju'y 1* ! Handeni (wood-
land and thotn) 1 ? July 23.
Encedon (near infiiscata) — Arusha 1 $ May 16.
Encedon, worn, with rather dull coloration — Ufiomi
(woodland) 1 $ June 4 ; Mabirioni 1 ^ July 14.
Zj/cia New Moshi 1 2 May 6 ; Handeui 1 J July 23.
The above analysis of localities shows that there is Uttle
or no correspondence between the form and the character
of the habitat.
A. sotikensls, Sharpe.
All the specimens are of the form rowena, Bltr., dis-
tinguished from the typical form by having the inner
marginal part of hind-wing patch yellow instead of
red. This form has hitherto only been reported from
Mt. Ruwenzori.
Uflomi. 8 SS, 2 9?. June 1-5 (1 ^, 2 ?$ plains, the
rest woodland).
A. cabira, Hoppf.
Three examples of the typical fonn not calling for
special comment.
Arusha, Sanja R., Mabirioni. 3 ?$. May 10-July 14.
A. acerata. Hew.
Three examples in marking somewhat intermediate
between the type and the vinidia form. One female is
intermediate to the form lenella.
Uflomi (woodland). 1 ^. June 4.
New Moshi. 2 ?$. July 1.
A. terpsichore, Linn.
A long series of this abundant species. All the males
are of the form r&ugett, Guer. The females are not so
variable in form as is usual in this species. Four resemble
the males, the remainder are largely of the form having
dusky fore-wings with more or less whitish subapicol
patch, one or two having a jrreat deal of white on the fore-
wing. Only two ()f the males have any red marks on the
hind-wing underside.
Kikuletwa-Darjama R., Ufiomi, Same, Handeni, New
l„y,l,Ajh,COOl^lC
CoOection of Bvtterfiies taken in Bast Africa . 329
Moshi. 9 S6, 34 ??. (5 SS, 27 ?$. Ufiomi plains, 1 $
woodland.) May 1&-July 23.
A. phanalus, Ward.
Two males of the fonn pharsaimdes, Holl., which seems
generally to replace the typical form in these localities.
New Moshi, Arusha. 2 3^. May 6-16.
A. perenna, Doubl.
One iemale example of the form thesprio, Oberth., in
which the red colour extends over the greater part of.the
fore- wing.
New Moehi. 1 ?. May 6.
A. oreas, Sharpe.
Two examples of which the male is of the alinmacvJata
form, the other, a female, has the tip of the left fore-wing
sienna brown instead of black.
Arusha. 1 ,J, 1 ?. May 13-16.
A. eiebrla, Hew.
A series showing the usual variability. The majority
are of the fonii jacksoni, Sharpe. Two are of the form
pTotea, Trim.; one female is form vumletronis, Butl., and
one female intermediate between motOeironis and nubilata,
Elti.
Aiusba, Ufiomi (woodland). Same, Mabirioni. 9 (J(J,
6 S$. May 14-July U.
A. tyooa, Gcodt.
Two examples of the tona fallax, Bogenh. This ia the
most southern locality I have for this form.
Amsha. 2 $$. May 10-13.
A. johnstoni, Godm.
One male is typical. The other two are of the variety
of confuaa, Rogenh., which haa the hind-wing discal
patch white as well as the fore-wing spots. (See Trana.
Ent. Soc.. p. 342, 1912.)
New Moshi, Mabirioni. 3 (J^. July 1-14.
PUnema aganlce montana, Butl.
Kikuletwa-Dariama R., Ufiomi (woodland), Mabirioni,
Anuha. 1 J, 8 $?. May 16-July 14.
u,y,l,/eJh,G00glc
330 Dr. H. Eltringham an a
P. ifuadrloolor, Bogenh.
Arnsba. 1 $. May 13.
PardopslB pimetaUssiiiu, Boisd.
Same. 1 J, 1 ?. July 10.
With reference to the position of this speciea Professor
Aurivillius ezpresBes the opinion (in Seitz' " Maciolepi-
doptera ") that pending a knowledge of the early stages it
should remain with the Acraeinae, with which it agrees in
certain particulare, including the structure of the fore-
legs. I was at some pains in my monograph of the Genua
Acraea to point out that the species does not agree with
Acraea in this last particular.
NYMPHALINAE
Euxanihe wskefleUi, Ward.
Handeni. 1 (J, 1 ?. July 23.
Charazes etheocles. Or.
New Moshi. 1 ^. May 6.
C. eandiope, Godt.
Ufiomi (woodland). 1 (J, June 4.
C. elthaeron, Feld.
Eikuletwa-Darjama R. 1 (J. May 19.
C. zoolina, Westw.
This dimorphic species is represented by two examples,
one zoolina and the other neanlhes, Hew.
Kikuletwa - Darjama R. $ {zoolina). May 20 (wet
New Moshi. (J [neanthes). July 1 (dry season).
Ufiomi (woodland). 1 <J. June 2,
Euryphura achlys, Hoppf.
Ngereugere. 2 <JcJ. Sept. 13.
n,g,t,7.dh,G00glc
CoUedion of BuOerfiies taken in East Africa. 331
Euryphene senegshnsis ortenUs, Eaisch.
Kondutachi. 4 ^$, 2 $$. Sept. 30.
EphMdra neophron. Hoppf.
Two males of the ordinary form, and one male which
appears to be a rather worn and faded specimen of the
form vwlacea, Batl.
New Moshi. 1 <^ (tnolacea). May 6.
Ngerengere. 2 ^t^ (neophrmi). Sept. 13.
Hamanumlda daedal us. Fab.
Handeni. 1 cJ. July 23.
Neptisagfttha, Stoll.
Ufiomi (plains). 2 ??. Jmie 1.
N. saclava marpessa, Boisd.
The marpessa form is the continental representative of
the Madagascar saclava, and-is very widely distributed.
Ufiomi (woodland). 2 $$. June 5,
Byblla lUthyla, Dnir.
Same. 1 ^, 1 ?. July 10.
B. aohelola, Wallingr.
Wet f. vulgaris, Stgr.
Ufiomi (woodland). 1 ^. June 2.
Eurytela hlarbas tila, R. & J.
Tfae East African race of kiarbas, Dnir.
Ufiomi (woodland). 1 ^. June 2.
E. dryope angulata, Auriv.
The early stages of kiarbas and dryope are, according to
Miss Fountaine, indistinguishable, though their specific
identity seems not yet to have been established.
Ufiomi (woodland). 1 $. June 4.
Hypollmnas mlsippus, Linn.
Ooe female of the typical form.
Same. 1 ?. July 11.
n,g,t,7.dh,G00glc
332 Dr. H. Eltringham on a
Hypollmnas duUa, Pal.
Four examples of the wahJbergi form, showing consider-
able variation in size, the smallest 70 mm. in expanse and
the largest 100 mm. Also four specimens of the mima
form, of which one, a large female, has the hind-wing pale
area white dusted with yellow.
Arusha, Mabirioni. 4 ^^ {uKthlhergi). May 14-JuIy 14.
Ssalanga, Ufiomi (woodland), 2 ^,$, 2 $$ (mima).
June 4-'9.
Pseudaenea lucretia expansa, Butl.
Handeni. 1 ?. July 23.
Salamis parhassus aethiops, Pal.
Two fine examples. The species difFers from anacardU,
L., in having a glossy surface on the underside of both
Ufiomi (woodland). 2 $$. June 4.
^rameis cardui, Linn.
A female of this ubiguitouis species taken at an elevation
of 4500 ft.
Ssalanga. 1 $. June 9.
Catacroptera cloantbe obscurior, Stg.
One female of the dry form of doanthe, though taken
towards the end of the wet season.
Ufiomi (plains). 1 $, Jime 2.
Precis octavia. Gram.
A series of this species of which twelve are of the sesamua
or dry season form, and one naialensis. The first example
of sesamus was taken on June 2, and the wet season con-
tinued till about June 16. The remaining dates extend
to June 19. One seaamus taken June 19 shows, by the
red in the fore-wing cell, an approach towards an inter-
mediate form and to the usual dry fotm of the west coast.
Ufiomi. 6 ^^, 6 5$ (sesamus). June 2-6, 19 (3 ^$,
6 $9 taken June 19, under eaves of native hut, Ufiomi
plains; 3 SS i" woodland, June 2-5).
Ufiomi (woodland). 1 ? {natalensis). June 4.
ih,Cooi^lc
CoUedion of BvUerfiies taken in East Africa. 333
P. UmnorU taveta, Rogenh.
New Moshi, Same. 1 tj, 1 ?. May 6^uiy 11.
P. antllope antilope, Feisth.
The diy season form.
Handeni. 1 S- ^^J 23.
P. terea elglva. Hew.
Ufiomi (wcodland). 1 J. June 6.
LYCAENIDAE.
Teriomlma fraya, S,-& K.
Handeni. 4 cJc? (one doubtful). July 23.
SpalgU lemolea, Dnice.
Tanga. 1 <J. Nov. 10-14.
Dranothauma lalkeostelni. Dew.
Ufiomi (woodland). 1 S- J'"ie 4.
Vliaehola antalus, Hoppf .
Ufiomi (plains). 1 ?■ J^e 2.
Polyommatus boeticus, Linn.
Ufiomi (plains). 1 $. June 2.
Azanus mlrta, Plotz.
New Moshi. 2 SS- May 3.
Azanus slEillAtus, Butl.
New Moshi. 1 cJ. May 3.
PIERINAE.
Terias brigitta. Cram.
NewMoahi. 1 <?. May 3.
Terias r^ularls, Butl.
New Moshi, Tarangere R., Ufiomi (woodland). 9 tjcj-
May3-Juae20.
TRAN9. EHT. SOC. LOKB. 1917.— PARTS II, m, IV. (MAY '18) Z
&34 Dr. H. Eltringham on a
Tencolas enponve. King.
Taiangeie R. 1 S- June 20.
TeneoliB evafore. King.
The form antigone, Boisd.
Same. 1 J. July 11.
Teraeolns baUmede, Klug.
TarangereR. 9 (JfJ, 4 $?. July 10-11.
Teraeolns ehrysonome, Klug.
Loldiloi, Ufiomi (plains), Tarangere R. 3 S3, 9 ??■
May 20-June 20.
Note. — From the last-named locality there were eight
females and only one male.
T«raeolus protomMlia, Klug.
Same. 1 ?. July 10.
" Damaged before capture."
Colias electo, Linn.
Ufiomi (plains). 4 (J<J, one being white. June 2-3,
Eronia teda, Boisd.
Handeni. 1 J. July 23.
Eronia cleodora, Hiibn.
Handeni, Mabirioni. 1 (J, 1 $. July 14-23.
Leueerotiia argia, Fabr.
Ufiomi (woodland), Handeni. 3 (J(J. June 3-July 23.
Leuceronia thatassina, Boisd.
Ufiomi (woodland). 1 ?. June 3.
LeuceronJa buquetil, Boisd.
Handeni, Same. 1 (J, 1 ?. July U -23.
Pinacopteryx vidua, Butl.
Ufiomi (woodland). 1 $, June 5.
iM,Googlc
CoUedion of Buflerfties taken in Bast Africa. 335
PioMoptciyi irifva, Boiad.
New Moshi, Ufiomi. 4 (J(J. May 6-Jime i.
i. ssUrto, Butl.
Ufiomi (woodland). 1 (J. June 4.
Beknois severiiu. Cram.
TarangereR., Ufiomi, Same. 1<?,2?? Jiine4-July 11.
B«lenols mesentiiu. Cram.
Handeni. 1 J. July 23.
Belenois nurK&rftaMS, Sharpe,
Sealanga. 1 ^. Jmie 9.
Hjrlotfarls afalhina. Cram.
Ufiomi, Handeni. 6 ^3, 5 9?. June 2, 19, July 23.
1 (J, 4 $? Ufiomi (plaina) ; 2 (J^, 1 ? (woodland).
Nycbiloiia mednsa f. aleeita. Cram.
Ufiomi (woodland). 4 .JcJ, 4 ??. June 2-6.
PAPILIONINAE.
Papilio Qiraus lyaeus, Doub).
Difiera from true niretis in having a much shorter blue
spot in area 2 of hind-wing.
New Moahi. 9 <S<S- May 3.
P. laonldas leonldas. Fab.
One female example, unfortunately without data.
P. dardanus tibullos, Kirb.
2 f, nov, lamborDl, Poult,
The single female was captured June 3, 1916, at Ufiomi
(woodland). A male was taken in the same locality on
June 6; a second on June 9 at Ssalanga, and a third on
July 14 at Mabirioni.
The female is an extremely interesting form, being very
similar to that described from much further north in Trans.
But, Soc, 1906, p, 290 — ^a Iropkonius form from the Kikuyu
iM-,i,A J h, Google
336 Dr. H. Eltringham on a
Eflcarpment (6500-9000 ft.) near Nairobi, with the pale
markmga not white but letaimng the piimitiTe yellowish
tint of trimeni, and the broad orange marking incompletely
developed, so that it does not quite fill its usual area, the
outer end of the fore-wing patch remaining yellowish. It
was pointed out in the paper referred to, that this specimen
supported the conclusion that Irophonius had arisen direct
from trimeni and not indirectly from it by way of kippocoon.
The existence of another specimen of the same form from a
very different locality aSotds confirmation. The differ-
ences between the two specimens are only such as are found
between different individuals of each of the female forms
of dardanus. Thus, the southern specimen from a lower
altitude ia considerably larger, being just over 90 mm.
in expanse as against just under 80 mm. ; but a small MZe
is characteristic of both males and females of dardamis
polt/tropkus from the high Kiknyu Escarpment. The
southern specimeh is darker and richer in colouring, but
this difference is intensified by its freshness; its hind-wing
orange patch is squarer, with a more pronounced angle
in area 5, and is more encroached upon by the broader
black margin. A vestige of the " tail " involving the
lengthening of vein 4 is seen in the' northern specimen but
not in the southern, just as it is present in some trimeni,
but not in others.
In the fore-wing the band of black ground-colour between
the sub-apical bar and the orange patch on the inner margin
is about twice as wide in the southern specimen, and there
is also far less invasion of the cell by this orange patch.
Furthermore the sub-apical bar and the spot in the cell
are fused in the Kikuyu example, quite distinct in the
southern. The cell spot itself is double in the latter, single
in the former.
In spite of these and other differences both females belong
to a characteristic form for which I propose the name
hmborni. It may be defined as a trimeni form in which
the yellowish ground-colour of the main area of both wings
ia replaced, but incompletely in the fore-wing, by orange.
The specimen from Ufiomi is probably more typical, and
I therefore select it as the type of this female form.
The three males are all of the tibullus form with the black
discal band of the hind-wing heavily marked, although not
so strongly developed as it conmionly is in this sub-species.
The band of the specimens taken June 6 and June 9 shows
l„y,l,Ajh,COOl^lC
Collection of Butler/lies taken in East Africa. 337
in area 5 a marked " costal gap," closed on its outer side
by a narrow black V with apex outwardly directed. In the
male of July 14 a slight indication of the " anal gap " ia
represented by a thinning of the black baod, from without
inwardly, in area 3.
The pattern of these three males together with the
geographical distribution of all four specimens shows that
this laTnbomi female belongs to the subspecies tibullue.
The Kikuyu example of tms form, on the other band,
belongs to pdytrojAus.
E. B. P.
HESPERIIDAE.
Tagiades flesus, Fab.
Ufiomi (woodland). 1 J. June 6.
Cyclopldes, sp. 1
One example not yet ideotiEed. This specimen is
evidently closely allied to Cychpidei Irisignatus, Neave,
from which it differs principally in the total absence of
orange spots in the hind-wing.
Ufiomi (woodland), Jime 6.
n,g,t,7.dh,G00glc
XII. Further notes on recapilulalory attitudes in Lepi-
dopteta. By T. A. Chapman. M.D.
[Read OoUibec 3rd, 1917.]
I HAVE made a few more observations on the methods
followed by some Lepidoptera in passing from the attitude
of drying their winf^s after their expansion to their ordinary
attitude of rest. The subject is interesting from any point
of view, and especially as it points to each si>ecies that has
a special resting attitude adoptingduring this period cerlain
positions that are actually, or in some degree recjill, the
ordinary resting attitude that b not theirs but that of the
group to which they belong, or that is most frequent in
Lepidoptera.
I have only had opportunity to observe three more
species of butterflies. They agree with those reported on
last year, in making certain opening and closing wing
movements, not after the wings are dry, but during the
process of drying ; so far as my few observations go nothing
of this sort occurs in the Heterocera.
In P. rapae this was seen to suggest an effort, which the
limpness of the undried wings prevented being successful,
to spread the wings flatly, as in the resting attitude of so
many Lepidoptera. The other species noticed had similar
alternations of opening and closing the wings, but through
a much smaller angle that would not suggest an eSort to
extend them flatty, but for the fact that they were obvi-
ously of the same character as in P. rapae. Though this
curious habit may have some other meaning, none has
occurred to me but that mentioned in my previous paper.
With regard to the Heterocera observed, the Geometers
follow broadly the same procedure as did those reported in
the previous paper, and suggest that the habit they exhibit
will obtain in all similar (ieometers, that is, similar in having
resting positions other than the typical deltoid one. My
examples come from each of the three largest sub-families
of our British Oeometers, so that the habit is in no way
of classificatory value. This conclusion is confirmed by
finding that precisely the same reminiscence of the typical
resting position occurs in the quite unrelated F.fahataria.
TBAN3. EMT, SOC. LOND, 1917. — PAST.S U, ID, IV. (MAY '18)
l„y,l,Ajh,COOl^lc
Dr. T. A. Chapman on alfUvdes in Lepldoplera. 339
The detailed notes taken minute by minute duriug the
obaen'ations, as I gave them in my firnt notes on this
subject, seem of little value, except as proof that the
observations actually were made, so I do not append them,
but rather describe the general succession of events in each
species, instead of leaving them to be worried out of memo-
randa made hurriedly during the observations, and not
perhaps easily understood, owing to their abbreviated
nature, except by myself.
The species observed were —
P.brassirae. The wings expand to a position the same
as the normal resting position, that in which most Lepi-
doptera dry their wings. Almost as soon as one can say
expansion ia completed, the wings are opened apart to a
position in which the fore and hind wings being together,
the mid-costae are 8 to 12 mm. apart, and the apices 3 to
8 mm., BO that the attitude corre-sponds to that which in
P. rapae I have described as bell-shaped ; but the separa-
tion is so slight in comparison that this attitude would not
in brasskne suggest any resemblance to the outline of a
bell. It is, however, essentially the same, in that the wings
being limp the apices approach each other, more than
the mid-costae, and are in fact 3 to 8 mm. apart instead
of 12 to 20, as they would be if the wings were stiff. They
maintain this attitude for about thirty-five seconds and then
close, so that the costae are in contact to within 4 mm.
of the thorax. They remain so for about forty-five seconds
and are again opened to the open position. This alterna-
tion is repeated about fourteen or fifteen times. The
actual opening and closing takes about ten seconds.
The period during which the wings are open is very similar
to that during which they are closed, viz, about thirty-
five to forty-five seconds. These vary a little in the same
specimen and between different specimens, but only by a
few seconds. I ought to have said that when open there
is the variation of 8 to 12 mm. at mid-costa in different
specimens, not in the same specimen, and that 8 mm. at
Diid-costa corresponds to 3 at apices, 12 mm. to 8 at
apices. At the fifth or sixth opening the hind-wing tends
to open a little before the f()re-wing, separating from them
by a millimetre or two in the process. About the seventh
or eighth opening, the wings are seen to be stiffer than at
first, and the apices are as wide apart as the mid-costae.
About the tenth the apices open wider then the mid-costae,
ih, Google
340 Dr. T. A. Chapman's FuTther NoU^ on
the wings being stiffer and holding themselves straighter.
About this time the bird-wings tend to be a little separate
from the fore-wings when open. Then the fore-winga open
but little and the hind-wings rather more. Gradually by
about fourteenth opening the hind-wings only open, or the
fore-wings hardly pereeptibly; gradually the separation of
the hind-wings diminishes, and somewhere about the six-
teenth to twentieth alternation one may say the process
has finished. When closed, the wings at first close very
close up to the thorax, their limpness causing no resbtance
by the further portion of the wings to this approximation ;
as the wings get stiffer they do not close so far up, only
sometimes for a third or half-way from the apex. Later,
when the final resting attitude is assumed, they close
further up, nearly as far as at the first closing. One or
two specimens opened more than the usual 8 or I'2 min..
one as much as 20 mm. and might fairly be described as
in the bell attitude.
Throughout the process the butterfly at inteivala, with-
out reference to the wings being opened or dosed, makes
a shivering movement, at others rocks to and fro a little —
this more frequently on closing the wings, and more frequent
in later stages— and makes a few fluttering movements of
the wings of an amplitude of about 05 mm. All this lime
the antennae are well separated, much as in the mature
butterfly, but are directed slightly behind a line at right
angles to the line of the body, .which is reached by them
about the time the wings finally close, but do not reach the
final somewhat porrect position tiU an hour or more after ;
their advance to this position is gradual and impereeptible.
The hind-wings have the costae nearly level with those of
the fore-wings when mature, during the opening and shut-
ting movements they are usually a millimetre or two behind.
We have here, though less obvious, just as in P. rapae,
an effort to assume the Lepidopterous resting attitude, t. e.
with the wings flat, but as in P. rapae it occura not after
the wings are Ary but during their drjing. being as it were
pushed backwards in the ontogeny.
Pien's napi. The process in napi is almost intermediate
between those of rapae and brassicae, the Ml attitude
results from the wings being depressed to an angle of about
45^ (90^ between the opposite wings) instead of the 90° of
rapae or the 10^ or 20'^ of brassicae. The details differ a
little, but hardly to a degree worth full description.
ih,Cooi^lc
recapitulatory altitudes in Lepidopfera. 341
Ckrysophanus dispar. var. rulilus emerges about 9.30 a.m.
Winga expand very rapidly in from six to ten minutes,
Ihey expand to butterfly resting attitude, i.e. wings
closed doTsally. There then begins a succession of what
must be called opening and shutting the wings, though
the opening only amounts to a separation of 2 or 3 nun.
When closed, the wings (costae) are in contact for four-
fifths of their length, when open, though the wings are
elsewhere separated, the apices nmy in some cases meet.
They remain closed for about thirty seconds and open for
forty-five seconds, but the time varies a little, for about
sixteen times in twenty minutes and then for several times
more, but so slightly as to be almost doubtful, settling
down to the distal two-thirds of the winga touching at the
end of half an hour. About the middle of the period the
hind-wings open from the fore-wings, so that their tips are
6 01 7 mm. apart during several periods of being open.
Various minor details as to movements of proboscis and
positions of antennae are not perhaps worth reporting.
Smeriiithis populi and ocdlalus only afforded me three
observations ; they have a habit, very trying for this pur-
pose, of emerging hardly before and usually a good time
after midnight. They expand the wings into the drying
attitude, and then gradually and imperceptibly open tbem,
during about half an hour, to the oniinary resting attitude
for the fore-wing; the hind-wing is then only partially
advanced to show its costa, not more, in fact, than is
compelled by its greater width ; they project about 7 mm,,
and it is some time before they advance to show nearly
double this width.
It cannot be said, so far as my meagre materials justify,
that these species show any attempt to exhibit the normal
Lepidopterous resting position,
FtUcaria falcataria follows very closely the habits of the
Georaetere, Epkyras, Acidalias, etc. It emerges about 5 to
5.30 a.m., rarely or never after 6.30. It expands the wings
under an hour, then throws them back into usual drying
attitude with a little sudden jerk, remains so for perhaps
forty minutes, then gradually lowers thera, i. e. with no
sudden jerk to flat attitude, with inner margins against
body, J. e. to normal Geometer resting attitude. Some speci-
mens advance the fore-wings gradually and veiy slowly to
the resting attitude of the species, i. e. exposing much of
the hind-wings; others remain in the triangular position
iM-,i,A J h, Google
342 Dr. T. A. Chapman's Further Notes on
much longer, probably till period of evening flight in moat
cases.
Pkalera bitcepkala. The resting attitude in most Noto-
donts is the normal one with the wings in pent-house
position. In Inicephala this is, one may say, exaggerated,
HO that the wings curl round the body in such a way that
the costae of the fore-wings approach each other beneath
the insect, and the apices are never more than a few milli-
metres apart, and may touch if the object on which the
moth rests permits or favours this.
The wings expand to a position in which their surfaces
form a flat arch over the dorsum, then they are thrown back
in the usual way into butterfly attitude. They remain so,
drying, for half an hour, and are then deflexed to about
the arched position to which they expanded, the surfaces
of the fore-wings being at an angle of about 1 10°, the inner
margins in contact and the apices 25 mm. apart. In about
an hour the angle of the wing surfaces to each other i? 90°,
and the apices are only 16 mm. apart. The wings have
already a little curvature. In another hour the tips are
only 9 mm. apart, and in successive hours they are noted as
6 mm., 4 mm. and 3 mm. apart. Three hours more they
are observed to be touching; this movement, from the
rather flat pent-house position, to that of the wings being
so curled round the body, is gradual, with no actual move-
ment observed at any time, and would appear to depend
on the gradual acquirement by certain wing muscles of
the necessary tonicity.
H. abriiptaria emerges in the afternoon about 4 or 5 p.m.
A specimen kept the wings in ordinary Geometer position
(triangular) from 5.30 to 7.30, though disturbed once about
the middle of this period. Ten minutes later (7.40 p.m.)
it had assumed the ordinary resting attitude of the species
with the fore-wings advanced,
T. consomrria emerges about 4 p.m. An hour later the
wings are expanded and the butterfly attitude taken, and
in another half-hour they are deflexed to the ordinary
triangular Geometer reaiing position, which is maintained
for at least two hours and probably as a rule till the time
of flight, but on earlier disturbance the usual resting posi-
tion witlt adva!iced fore- wings is assumed on resting again.
I did not ascertain after how short a period in the triangular
position tliis would occur.
. In Tephroclifsfis isoijrammnta the wings were maintained
l„y,l,Ajh,COOl^lC
recapUulaiory atlilvdes in Lepidoptera. 343
in the drying (butterfly) attitude only twelve minutes, and
were then placed in the Geometrid (deltoid) position. Being
disturbed some five minutes later, so that it ran two inches,
it settled again in the same attitude. The two costae form
an angle of about 100° ; an hour later it was about 150°,
and in another hour the usual resting position was attained,
with c<ratal angle of 190° to 200°, i. e. with the wings forward
of having the costae in line.
Selenia illuslraria. This species differs 8 good deal from
S. illitnaria in its methixis of proceeding from wing expan-
sion to the normal resting position. Though in a warm
room, specimens would occasionally be more than half
an hour after emergence before any sign of the wings
expanding could be detected. The winp then expanded
rather rapidly, taking, however, usually about or a httle
over thirty-five minutes to expand. When expansion is
completed the wings hang backwards, but only the tips of
the fore-wings touch each other, not indeed always doing
BO. Then gradually the wings approximate to the butter-
fly attitude, that is, closely pressed together dorsally, so
that the costae are coincident for their distal halves.
The moth selects a vertical or, if possible, a slightly
overhanging position in which to expand, so that now they
hang downwards, more or less, whilst in this backward
position. A special feature is that this dorsal position is
gradually attained by quite imperceptible movement, and
this peculiarity continues throughout the whole evolution
from the beginning of expansion to the attainment of the
normal resting position ; there is no sudden itiovement as
is so common, either of throwing the wings back, or assum-
ing the resting position. This closing backwards is attained
in ten or fifteen minutes after expansion is completed ; the
wings are kept ao for about twenty minutes and then are
gradually separated in a minute or two till the costa of
fore-wings are separated to an angle of 10° or 15°; the
separating movement b continued in the same gradual,
haidly noticeable manner, till in about two more minutes
the angle is 60°, in two more 90°, and in two or three more
finally 100° or 110°. Usually the process is rather slower,
and varies a good deal in its rapidity at different stages.
In the position of the moth, gravity no doubt tends to
make the wings hang directly backwards ; nevertheless, the
po-sition of the wings at this stage is with the costae of
the anterior wings at an angle with each other of about
ih, Google
344 Dr. T. A. Chapman's Further Notes on
120°, and of the flat surfaces of the wings of about 140";
the costae not beiii;; so far advanced as to be in a plane
transverse to the line of the insect body, but behind this
in a plane that slopes backwards from it at a consider- -
able angle. This position, which appears to represent the
memory of the normal (triangnlar) resting attitude of Geo-
meters, may last about thirty minutes, and in another
thirty minutes the wings close again doreally to a costal
angle of about 45°. At thus point the costae of the fore-
and hind-wings on either side are nearly coincident.
In the same gradual imperceptible way in which the
other movements occur, the fore-wings advance from the
hind-wings. Measured from costa to costa at the post-
discal line it takes about two hours for the fore-wings to
be 3 mm. in advance of the hind-wings, the fore- and
hind-wings being still in the same plane. Then, still
very gifldually, the fore-wings assume the curious bend in
a line below vein 2 that characterises the ordinaiy
resting attitude of the species, the inner part of the wing
being in the same plane with and close against the hind-
wing, the greater (costal) portion being raised at a con-
siderable angle. At the end of another hour oue may
think this attitude is attained, at the end of two one has
no doubt it is. All the progress and the movements
involved are so gradual and imperceptible that it is difficult
to divide them into stages and say when each is complete.
Itliislraria emerges in the morning about 6.30 to 7.30
a.m.
Ennomos lunaria. The wings are thrown back when
expanded, and some ten minutes later and when appar-
ently dry are opened gradually, so that in about six minutes
they nearly approach a flat position, instead of 180° (flat),
between the two fore-wings they are about 160° or 170°.
In some ten minutes the angle diminishfes to that of norma!
resting, perhaps 80°, but there is still no angulation of fore-
wing, and the hind-wing is only 3 or 4 mm, behind
fore-wing. The complete resting attitude with fore-wit^
advanced and with the longitudinal fold is not attained
till an hour or two later. 1 see that in one specimen I
noted the wings were opened till quite Sat; in another
they did not get quite as far, but I did not note the angle.
The costae of fore-wings made an angle of 90°, but the wing
surfaces were much more.
Fidohia pinUiria cniei^os C to 8 a.m. The wings expand
J,., Cookie
recapUukUory atlitudes in Lepidoplera. 345
to the dor»al (butterfly) position in about twenty minutes;
in about (en minutea more they are gradually separated,
80 that in two or three minutes the wings are separated so
that the apices are some 18 mm. apart, the an^le between
the wings being about 40°. They remain thus for about
six or seven minutes, the wings appearing to be quite dry
and stiif. They then gradually close, at the rate of about
ImiD.(foreachwing)aminute, andin ten or twelve minutes
the butterfly attitude (the normal resting attitude of
piniaria) is reached, to appearance, but they arc not tightly
closed for ten or twenty more minutes. The whole process
is gone through very rapidly (compared with most other
species). After the wings are dry they are separated, but
not widely, so that the reminiscence of the flat deltoid
attitude, though still present, is comparatively brief and
slighf. The process reminds one of that obtaining in the
butterflies, but the important difference is that it takes
place after, not during drying; also, of course, it is one,
not a succession of movements.
iS, illunaria and F.ptniaria both have a normal resting
attitude similar to that of the butterflies, with the wings
raised over the back ; but the reminiscence of the ordinary
Geometrid attitude which they display, in the interval
between the" completion of the drying of the wings and the
assumption of their resting attitude, two attitudes that are
identical and that one would expect to find continuous, is
curiously different in the two species. In both species,
however, the reminiscence is there.
In the Etudes de Lepidopterologie Comparee, Fasc. V (n),
p. 115, Mr. H. Powell records how St/ricklkus mohammed,
Obthr., having expanded its wings in the closed (" butter-
fly") attitude, depresses them to the CJeometrid or tri-
angular position, and gives photographs on PI. Hj, Fasc.
VI and PI. Hb, Fasc. V (n) and of S. proto on PI. 27,
Fasc. VII. He says this position is kept for only a short
period. In complete repose the wings are closed in butter-
fly attitude ; when basking, the depressed wings have the
costae ftt right angles to the body ; when resting briefly
the fore-wings are well raised. This merely summarises a
valuable demonstration that the recapitulatory attitude is
very manifest in Syrichthus. In this habit the " Skipper"
appears to be nearer the Helerocera than to the butterflies.
n,g,t,7.dh,G00glc
( 346 )
XIII. A PreHminary CtUalogtie of Bntisk Cecidomyidae
(Diptera) u>ih special reference to the Gail-midges
oflhe North of England. By Richard S. Bagnall,
F.L.S., and J. W. Heslop Harrison, D.Sc.
(Read H&y2Dd. 19170
For many years tbe Cecidomi/idae, a family of Diptera,
familiar enough to naturalists in a cursory sort of way
(because it embraces the little flies known as " Gall-
midges "), has been neglected by British entomologists, in
spite of its extent and great economic importance.
Of its members a considerable proportion are purely
gall- causers, each producing its particular gall on its
host-plant or plants. These galls are characteristic of the
species and are therefore part of its specific characters, but,
unhke many such distinctions, they can be thoroughly
relied upon as a means of determining the species. They
are not, as many entomologists unacquainted with their
stability think, unworthy of consideration; such an
opinion simply displays ignorance on the part of its
holder.
Any one taking up the study of the group in real
earnest will soon find that many do not fall into the
categorj- of genuine gall-makers. Their modes of life are
exceedingly varied, and thereby render the subject the
more interesting. Some feed as larvae under bark, in
decaying wood, in stems of grasses, sedges, fungi and
mosses; others, again, find their food in epiphytic fungi
such as rusts, smuts and mildews. We find still more
figuring as commensals and inquilines in the galls of
Cynijiiiiae, Gall-midges and other insects, and also of the
Gall-mites (Eriophyidae). whibt others have larvae ecto-
parasitic or predatory- on Aphididae, and Eriophrid and
other mites. Of the remainder, a small number are
Endoparasites, the larvae li\*ing in the abdomens of certain
Hemiptera {Aphkh'dae. PsyUidae and Tingidae).
Some of the Lcxtreminne, Uke those of the genus Miaslor,
reproduce their kind bv paedogenesis. and are of extreme
TRAMS. EKT. SOC. LONO." 1917.— PABTS II, lO, IV. (MAY '18)
ih,Cooi^lc
MeasTS. Bagnall and Harrison's British Cecidomt/idae. 347
importaDce in biology as suitable subjects for studying the
early segregation of germ-ceLs.
There is no excuse for our naturalists to continue this
systematic neglect of the group. Houard (1908-X913)
includes descriptions of all the then-known Cecidomyid
galls of the western section of the Falaearctic area in his
great work on the Zoocecidia. Furthermore, there has
been in recent years a steady Sow of literature as the
result of the researches of continental entomologists such
as Kieffer, Trotter, Tavares and Rubsaamen. The first-
named author's monograph in Wytsman's " Genera
Insectorum " (fasc. 152), which appeared in 1913, will,
with the fine productions of Dr. Felt, in America, form
the groundwork of much further work. Lastly, an
Itahan periodical, " Marcellia," edited by Prof. Trotter,
keeps one up to date with new researches.
It will thus be seen that in the Cectdoniyidae we have a
group not only displaying abundant and varied interest in
its biology, but one provided with a. Uterature unusually
rich for a " neglected order."
During the past three seasons we have devoted much
of our leisure in this fascinating study and with consider-
able success. We have therefore been tempted to publish
the present list as a record of our labours and as a basis
tor future work and workers. We have included references
to (a) original descriptions of genera and species, (6)Kieffer's
1913 Monograph, and {c) to Swanton and Houard as far
as Gall-midges are concerned, quoting only the authors'
Dumbering of the gall. In addition we have appended
an index to the host-plants. Owing to the fact that
Connold's works on galls are very far from complete, and
also because Swanton supersedes them, we have not
deemed it necessary to refer to that author.
Soon we hope to prepare a book on British Zoocecidia,
and later, when we have bred out and studied, as far as
our Umitations will allow, the hfe histories of the insects,
we shall, in all "probability, proceed with a more elaborate
monograph of the Cecidomyida. It is, then, for these
reasons that we characterise this as a " preliminary " hst.
The species noted from the north of England, c. g. from
the counties of Northumberland, Durham, Cumberlind,
Westmoreland, Lancashire, Cheshire and Yorkshire, are
shown in heavy type. Species not yet reported from that
area are shown in italics, whilst those ascribed to older
iM-,i,A J h, Google
318 MesBiB. R. S. Bagnall and J. W. H. HamBon's
authors and requiring confirmation (see italicised names in
Collin, 1904) are enclosed in square brackets. Although
most of Walker's species are capable of being correctly
assigned to Kiefier's genera, as species they are insufficiently
defined and thus unrecognisable by students; they are
therefore neglected here.
By keeping in close touch with each other we have been
able to dupbcate the records of very many of the most
interesting occurrences; in fact, the wealth of records is
a welcome feature. Except where a name is added all
records are our own; outside the Northumberland and
Durham area the first-named author is almost entirely
responsible for the records from Lancashire and Cheshire
(see Bagnall, 1917-1), and the few from Cumberland and
Westmoreland, and from Redcar and Leeds, whibt the
second-named author is similarly responsihle for the
Cleveland area, Yorkshire. We also add Mr. Burkill's
Yorkshire records (Burkill, 1916, 1917).
An asterisk in the main list denotes species not shown
in the hste of Colhn, 1904, and Swanton, 1912.
CONTENTS
L Classified List of British Cecidomyids with
North of England Records . . pp. 348-41 1
II. Check list of named species . pp. 411-417
III. Index to Host-plants . • PP- 417-422
IV. Literature ; references in brief , pp. 4'i2— 426
I. Classified Llst of British Cecidomvidae with
North of Enoland Records.
Subfamily CECIDOMYINAE.
Group Lasiopterariab.
CUnorrhyncha H. Loew, 1850, p. 21.
•1. C. ohrywmtheml H. Loew, 1850, p. 39; Kieffer, 1913-2,
p. 19.
Affecting achenes of Matricaria inodora and AtUhemis
Coinla. See Houard, 5722 and SGO.'J.
Records from all counties excepting Westmoreland.
ih,Cooi^lc
Prdiminary Caialogue of British Ceddomyidae. 349
«2. C. mUfcloUi Wachtl, 1884, p. 161 ; Eiefier, 1913-2,
p. 20.
Ailecting acheaes of Achillea MiOefolium and more rarely
A. Ptarmica. See Houard, 5672 and 5701.
More sparing than C. chrymrUhemi ; records from all
counties excepting Lancashire and Weslmoreland.
*3. C. leDUDtbeml Kieffer {chrysanthemi Scbiner non H.
Loew), 1889-1, p. 285; Kieffer, 1913-2, p. 20.
Affecting achenes of Chrysanthemum Lewanthemiim.
Houard, 5726.
NoRTBUHBERLAND, Whitfield, Durham, Birtley dis-
trict, Fatfield. Also from Scotland.
Trolteria (for ChoriMoneura (praeocc.) Riibs.) Kieffer,
1901-1, p. 561.
*4. T. lalU Riibaaamen, 1912-1, p. 376; Kieffer, 1913-2,
p. 22.
Gall on Gaiium verum, = Cecidomyid sp., Houard, 5290.
Lancashire, several, in a field near Ainsdale. North-
UUBBRLAND, Bamburgh and Warkworth, not rare. Durham,
Blackball Rocks, three examples only. Also from Scotland.
*5. T. svoUuunnl Kieffer, 1890-1, p. 136 (Lasioptera);
Kieffer, 1913-2, p. 22.
Galling seed-pods of broom. Houard, 3411.
Northumberland, near Ovingham; and Durham, Gib-
side; rare, lane between Rowley and Lancbester; Birtley.
•5a. T. nmlMUUerarnin Kieffer, 1913-1, p. 47; 1913-2,
p. 22.
On An^riscus.
Durham, Gunnergate.
Usioptera Meigen, 1818, p. 88.
6. L. alinpennis Meigen, 1804, p. 40 (Cecidmnyia) ; Kieffer,
1913-2, p. 31,
7. L. anindlnU Schiner, 1854, p. 175; Kieffer, 1913-2,
p. 31.
GaUing Phragmites communis. Swanton, 49; Houard,
241.
Northumberland, Warkworth. Durham, Bilhngbam.
trans, ent. soc, lond. 1917. — part* ii, hi, IV. (may '18) aa
ih,Googlc
350 Messrs. R. S. Bagnall and J. W. H. Harrison's
•8. L. eaUnugrostldis Riibsaamen, 1893, p. 164; Kieffer,
1913-2, p. 31.
On Pkalaris arundinacea.
NoRTHUMBEKLAND, Warkwortt. Durham, Birtley and
Gibside.
•9. L. earopUla F. Loew, 1874, p. 149; Kieffer, 1913-2,
p. 31.
Abroad this midge galls many species of Umbelliferae.
Durham, Wolviaton on Antkriscus, and Penshaw Hill
on PimpineUa Saxifraga. Not recorded from the AnthTteciK
in Houard.
10. L. rubi Heeger, 1851, p. 203; Kieffer, 1913-2, p. 32.
Galling stems of Ruhvs. Swanton, 478, 483 and 489;
Houard, 2964, 2976 and 3024.
NoRTHUMBEBLANn, Warkworth, not uncommon. Dur-
ham, on bramble, Ryliope; Tinkler Fell; Birtley. York-
shire, Nunthorpe.
*11. Lirioptsra ap. RUbsaamen; Schlechtendal.
Molinia caerulea, stem. Houard, 251.
Durham, near Lanchester,
Baldratia Kieffer, 1897-3, p. 6; 1913-2, p. 24.
*12. B. salicornlae Kieffer, 1897-3, p. 7; 1913-2, p. 26.
On Saliaymia radtcans. See Houard, 2240,
Durham, Greatham.
Steranlella Kieffer, 1898-2, p. 55; 1913-2, p. 28.
*13. S. brevipalpis Kieffer, 1898-2, p. 53; 1913-2, p. 29.
On Alriphx {OUone) porttihcoides. Houard, 2224.
Durham, Greatham.
Group Oligotrophariae.
[Meoeerata Coquillett, 1900, p. 47.t
•13a. [H.rhodoph«a Coquillett, 1900,p.47; Kieffer, 1913-2,
p. 41.
■f Felt sinks thiansanynonym ot /)(i«yneura (= Perritia).
ih,Cooi^lc
Prdiminary Catalogue of British Ceddomyidae. 351
Northumberland, examples taken in the buds of
cultivated rosea at Ninebanks are possibly referable to
tbis species.
Rhopslomylk RubBaamen, 1892, p. 370.
14. R. mUlerolU (achilleae Inchb, I860) H. Loew, 1850,
p. 37 (Cecidomyia); Kieffer, 1913-2, p. 44.
Gallfi on Achillea Millefolium and more rarely on A.
Plarmica. Swanton, 815, 816, 817, 818 and 823, 824;
Houard, 5673, 5680, 56a5, 5691.
Northumberland and Durham, sparingly but widely
distributed. Lancashire, once only, near Ainsdale.
15. B. UiUMtiMla Karsch, 1879, p. 27 (Oligofropkvs) ;
Kiefter, 1913-2, p. 44.
Somewhat similar galls on Tanacetiim milgare. Swanton,
830, 831, 832-, Houard, 5750, 5752, 5754.
Northumberland, near Chathill; Warkworth. Dur-
ham, several records for the Wear and Team valleys.
Hisopatba Kieffer, 1913-1, p. 48.
•16. M. tloram Kieffer, 1890-3, p. 37 {Ceddomyia);
Kieffer, 1913-2, p. 45.
On Artemisia vulgaris. Houard, 5817.
Durham, between Lambton and Cox Green ; Greatbam
and Port Clarence.
17. M. [oUoram H. Loew, 1850, p. 36 (Cecidtmyia) ;
Kieffer, 1913-2, p. 45.
On Artemisia vulgaris and Abrotanum. Colhn {1904, p.
94} pla«es abrotani of Trail (1886, p. 250} as a synonym of
this species. Swanton, 834 ; Houard, 5824.
Northumberland, Budle Bay. Durham, Billingham;
Birtley.
18. H. ptarmJoae Vallot {forica Winn), 1849, p. 98
(Ceddomyia); Kieffer, 1913-2, p. 46.
On Achillea MiUefolium and A. Ptarmica. Swanton, 813,
814 and 822; Houard. 5676, 5681 and 5706.
Sparingly, all counties excepting Cheshire.
n,g,t,7.dh,G00glc
352 Messrs. R. S. Bagnall and J. W. H. Harrison's
*19. H. sfDgsnosiae H. Loew, 1850, p. 39 {Cecidomyia);
Kieffer, 1913-2, p. 46.
On Anihetnia CUuUt and Matricaria inodora. Houard,
5666 and 5723.
Apparently widely distributed ; records from all counties.
AreeoUiomyia Kieffer, 1913-1, p. 49.
•20. A. valeril Tavares, 1906, p. 299 {Rho^paimnyia);
KieSer, 1913-2, p. 48.
On Juniperus communis. Houard (supplement). 6277
(see H. 135).
Lancashire and Westmoreland, Hampsfelt and Meat-
hop Fell near Grange -over- Sands. The species was origin-
ally described from the Mediterranean on Junipems
oxycedrus, but in 1912 Cotte recorded it from France on
the common Juniper,
OUgotropbus Latreille, 1805, p. 288-1
21. 0. Junlperinus Unn4. 1758, p. 588; Kiefler, 1913-2,
p. 49.
On Juniperus communis. Swanton, 11; Houard, 129.
Durham, near Chester-le-Street. Lancashire and
Westmoreland, Grange-over-Sands district.
*22. 0. pantoli Kieffer, 1898, p. 18; Kieffer, 1913-2. p. 49.
On Juniperus communis, known throughout Europe.
Houard, 126.
Lancashire and West.moreland, Grange-over-Sands
district, locally common.
*23. Oligotrophus sp. Riibssamen and others.
On Juniperus communis. Houard, 125.
Lancashire, Hampsfell near Grange-over-Sands,' rare.
*24. 0. alopecuri Rent.. 1895, p. 3; Kieffer, 1913-2, p. 50.
In seeds of Alopecurus pralensis.
Cheshire, Chester. Miss Ormerod, 1885, p. 32. See
Supplemental Note, Collin, 1904, p. 97. Northumber-
land and Durham, locally common.
t 0. jitniperinus, 0. 'panltli ftnd another Juniper species truly
belong tu tliiH (icnuH as now Ijmiled, but all otlier species ate placed
here temporarily pending further research.
ll,y,l,/eJh,G00glc
Prdiminary Cataloffue of British Ceddwntpdae. 353
25. 0. humrius Breroi, 1847, p. 52 (Cecidomyia) ; Kieffer,
1915-2, p. 50.
Galling leaves of Glechoma hederacea, Swaaton, 706;
Houard, 4809.
Records from all counties.
*26. 0. [agineus Kieffer, 1909, p. 7 (gall) ; 1913-2, p. 50.
Leaves of Fagvs, parenchymatous gall, = Cecidomyid.
Houard, 1156.
Northumberland, rare, Warkworth. Durham, Lamb-
ton ; Gibside. Yorkshire, Leeds.
*27. OUKotropbus sp, Kiibsaamen, Trotter,
A very distinctive gall on beech leaves, previously only
known h^m Russia and Asia Minor. Houard, 1157.
Northumberland, Ninebanks (Rev. J. E. Hull).
•28. 0. harUgi Liebel, 1892, p. 283 (Hormomyia) ; Kieffer,
1913-2, p. 50.
Leaves of TUia, parenchymatous gall; local.
Northumberland, Warkworth and Ovingham, sparingly.
Durham, Gibside, local ; Castle Elden Dene. Cumberland,
near Alston. Lancashire, Grange-over-Sands. York-
shire, Leeds.
*28a. O. loewlinos Kieffer, 1909, p. 4; 1913-2, p. 50.
On Carex arenarius.
Northumberland, Warkworth.
*29. O. rMumnrianus F. Loew,t 1878, p. 387 {Hormomyia} ;
Kieffer, 1913-2, p. 50.
An interesting gall on leaves of Tilia spp.
Yorkshire, Gunnergate, on one tree only. Also re-
corded by Mr. Burkill from W. Gloucestershire.
*30. O. tympanUex Kieffer, 1909, p. 6 (gall); 1913-2, p. 50.
In leaves of Corylus, parenchymatous gall, = Cecido-
myid, Houard, 1061.
Local, though apparently widely distributed ; records
from all counties.
t Referable to the genua Didyomj/ia Biibsaamen, characterised by
Kieffer oa insufficientTy described.
u,y,i,Ajh,Cooglc
354 Meaars. R. S. Bagnall and J. W. H. Harrison's
•31. 0. ulml Kiefler, 1909, p. 31 (gal! and larva); 1913-2,
p. 50.
In leaves of Ulmvs campeslm and V. montana,
parenchymatous gall, = Cecidomyid, Houard, 2046 and
2064.
Also widely distributed; records from all counties
excepting Westmoreland. Very plentiful in 1917.
PhyotidobU Kieffer, 1912-1, p. 220.
32. P. soltnsi Kieffer, 1906, p. 179 (Oligotrophus) ; 1913-2,
p. 51.
Galls on leaves of Viburnum LanJanu. Swanton, 774;
Houard, 5349.
MlkomyU Kieffer, 1912-2, p. 2.
•33. H. ooryU Kieffer, 1901-2, p. 22 {Oli^rophua):
1913-2, p. 52.
On leaves of Corylus. Houard, 1060.
Rare ; records from all counties excepting Westmoreland.
Semudobla Kieffer, 1913-1, p. 55.
•34. S. belulae Winnertz, 1853, p. 234 {Ceddomyia);
Kieffer, 1913-2, p. 52.
Galling fruit of Befwio. Houard, 1067. Note: Swanton
suggests that the species recorded by Bionie and Connold
(= Cecidomyid, Houard, 1068) may be this species. The
following records excepting one are of Houard, 1067,
Northumberland, Crag Lough (H. 1068).' Durham,
several records. Lancashire, Freshfield, Cheshire, Bid-
ston. Yorkshire, Eston.
Itaomyla Kieffer, 1913-1, p. 55.
35. 1. cai^eafl Winnertz, 1853, p. 291 {Cecidomyid) ;
Kieffer, 1913-2, p. 55.
Leaves of Salix Caprea and others.
Not uncommon ; records from all counties.
•36. I. malor Kieffer, 1898-2, p. 22 {Olif/. capreae, vat.
tnaj&r); 1913-2, p. 55.
Rarer than capreae; our records are from SdUx atirila
only.
l„y,l,Ajh,COOl^lC
Preliminary Cataioffae of BrUish Cexndomyidae. 355
Durham, Waldridge, Hart and Easington ; rare.
Lancashire, Ainsdale. Yorkshire, Redcar ; Leeda district.
Janetlella Kiefier, 1898-2, p. 23.
37. J. tomeei Kieffer, 1904, p. 71 {Oligotropkus); 1913-2,
p. 59.
Gall on midrib and nerves of Vlmvs nu»Uana. Swanton,
300; Houard, 2061. A very local species.
Northumberland, Stocksfield and Ovingham (H. S.
Wallace) ; Warkworth. Durham, Winlaton Mill ; dene
near Fencehonses ; Urpeth. Lancashire, Grange-over-
Sands. Yorkshire, Bardsey, near Leeds.
Specimens were taken at Warkworth on U. cantpeslris
and U. suberosa also.
•38. J. thyml Kieffer, 1888, p. 100 (CecicUmyia) ; 1913-2,
p. 59.
On Thymus. Houatd, 4912 and 4917.
Records from all countiea excepting Cheshire and York-
shire.
•39. J. thymlcola Kieffer, 1888, p. 102 {Cmdomyia);
1913-2, p. 59.
Also on Thymus, gall hke preceding, but pilose. Houard,
4921.
Northumberland, Seahouaea, Warkworth, Crag Lough.
Durham, near Sunderland, rare. Lancashiee and West-
moreland, Grange-over- Sands district. Also from Scot-
land.
•40. J. (ubercoli Rubsaamen, 1889, p. 61 .f
GalUng stem of broom {Sarotkamntis). Houard, 34:J4.
Northumberland and Durham, Derwent banks be-
tween Esperehields and Edmiindbyers ; and Durham, in
a lane between Newbiggin and Rowley, not common.
•41. JanetleUa sp. Bagnall and Harrison, 1917-2, p. 207.
On Thymus.
Durham, Stanhope and Penshaw Hill.
1 KiefTcr, 1913-2, unless it is
u,y,l,/eJh,C00glc
356 Messrs. R. S. Bagnall and J. W. H. HarrisoD's
ZyioUa Kieffer, 1913-1, p. 55.
42. Z. carpini F. Loew, 1874, pp. 157 and 322 {Cecidomyia) ;
Kieffer, 1913-2, p. 60.
Galling leaf of Carpinus. Swanton, 211 ; Houaid, 1045-
Craneiobla Kieffer, 1913-1, p. 55.
43. C. comi Giraud, 1863, p. 1301 (Cecidomyia) ; Kieffer,
1913-2, p. 60.
Gall on Cornus sanguinea. Swanton, 672; Houaid,
4543.
Phegobia Kieffer, 1913-2, p. 61.
*44. P. tonulella Bremi, 1847, p. 13 {Cecidomyia) ; Kie&ei,
1913-2, p. 61.
A glabrous gall (like that of the common Harligida
annulipes) on the upper surface of leaf of Fagua, = Houard,
1154.
Northumberland and Durham, several records. Cum-
berland, Alston.
HayetloU Kieffer, 1896-1, p. 89.
^45. H. aveiuw Marchal, 1895, p. 272 (Cecidomyia); KieSer,
1913-2, p. 62.
Gall, stem of Avena Jatva. See Houard, 223 {Avma
saliva).
Yorkshire, Nunthorpe.
*46. M. dac^lldis Kieffer, 1896-2, p. 217; 1913-2, p. 62.
Gall, stem of Daciylis glomeraia. Houard, 258.
Durham, banks of the Wear neat Penshaw.
47. H. destructor Sav, 1817, p. 45 iCecidotnyia) ; Kieffer,
1913-2, p. 62.
On Hordeum imlgare. Houard, 346.
Durham, Penshaw.
48. M. hold Kieffer, 1896-2, p. 333; 1913-2, p. 62.
On Holcxis hnatm, stem, Swanton, 46; Houard, 215.
Northumberland, Seahouses, once only; Warkworth.
Durham, Birtley, Gibside, Edmondsley. Lancashire,
liftthom.
ih,Cooi^lc
Prdiminary Catalogue of British Cecidom^idae. 357
•49. H. Jouinld Kieffer, 1896-2, p. 116; 1913-2, p. 62.
Gall, stein of Poa nemoraUi. Houard, 263.
North CM BERLAND, Warkworth. Dukham, between
Wolviaton and Billingham. Lancashire, Grange-over-
Sands.
*50. BL ventrieoU Riibaaameii, 1899, p. 566 (Oligotrophm);
Kieffer, 1913-2, p. 62.
On MoHnia caerutm. Houard. 249.
Northumberland, between Whitfield and Langley.
Lancashire (Grimshaw). Yorkshire, on all the Cleveland
•51. H. hardel Kieffer, 1909, p. 9 {gall)t; Kieffer, 1913-2,
p. 62.
On Hordeum vulgare, = Cecidorayid, Houard, 347.
Durham, between Vigo and Birtley.
*52. MsyeUota sp. Bagnall and Harrison, 1917-3, p. 228.
On Bromtis erectile.
Durham, Gibside.
Chortomyla Kieffer, 1913-2, p. 63.
53. C. bellwlgl Rubsaanicn, 1912, p. 217 (Poomyia);
Kieffer, 1913-2, p. 63.
On stems, Bmckypodiums sylraiicum, = Cecidoniyid,
Swan ton. 62, and Houard, 297.
Northumberland, Warkworth, local. Durham, Pen-
shaw, Ryhope, Easington, and Horden. locally common.
Lancashjre, Grange- over- Sands, probably not rare. York-
shire, Leeds district.
*o-l. C. moUnlae Rubsaamen, 189">, p. 180 (Oligolropkus);
Kieffer, 1913-2, p. 63.
On Molinia caenileti. Houard, 250.
Durham, Birtley Fell. Che.shire, Bidston Hill.
55. C. poae Bosc, 1817, p. 133 (Ceddomyia); Kieffer,
1913-2, p. 63.
On Poa nemoralis. Swanton, 50; Houard, 264.
t The generic position of thia Bpocies is not yet oerUin.
u,y,l,7rJM,G00glc
358 Meflsrs. R. S. Bagnall and J. W. H. Harrison's
NoHTBUMBERLAND, Warkwoith. DuBHAH, Pciishaw,
Birtley, Gibside. Cumberland, Alston. Lancashire,
Grange -o ver- Sands .
•56. C. ndletflca Rubaaamen, 1895, p. 179 {OligolrophM) ;
Kieffer, 1913-2, p. 63.
On Poa nemoTcdis. Houard, 265.
Northumberland, Ovingham and Langlev Woods
(H. S. Wallace). Durham, Ryhope (H. 8. Wallace);
Fenshaw and Blackhall Rocks. Cumberland, Nentbead
(H. S. Wallace).
Cystiphon Kiefler, 1892, p. 212.
*57. C. hleracU F. Loew, 1874, p. 145 (CecMfoniyifl);
Kieffer, 1913-2, p. 64.
On Hieracium ap., apparently very rare.
Durham, Hylton. Lanc;ashibe, Ainsdale, and West-
moreland, Ravenstondale.
•58. C. (anxaci Kieffer, 1888, p. 98; 1913-2, p. 64.
On Taraxacum officinale, very local and rare. Houard,
6090.
Northumberland, coast near Hamburgh ; Warkworth.
Durham, Blackhall Rocks; Pcnphaw. once only. Cumber-
land, Alston. Lancashire, Grange-over-Sands, once only.
•59. C. leonlodontis Kieffer, 1909, p, 14 (gall) ; 1913-2, p. 64.
On Leemtodon hispidum. Houard, 6053.
Northumberland, Warkworth. Durham, very local,
Fenshaw and Washington.
•60. C. pllosellae Kieffer, 1892, p. 213; 1913-2, p. 64.
On Hieracium Pihsella. Houard, 6207.
Very local ; records from all counties excepting Cheshire.
61. C. sonchi F. Loew, 187.5, p. 18 {Ceddotnyia) ; Kieffer,
1913-2, p. 64.
On Sonckus arven^. Swanton, 880; Houard, 6100.
Northumberland, abundant in N.E. Northumberland;
Ovingham. Lancashire, Ainsdale, one plant only. Also
Stirlinosribe, very common.
ih,Cooi^lc
Preliminary Catalogue of British Cecidomyidae. 359
•62. (^slipbora sp. Bagnall and Harrison, 1916 (iv), p. 248.
On Hf/poehoeris radicata. Apparently a new species.
NoBTHU-MBERLAND, Warkwortb, Durham, Fenshaw and
Edmondsley. Cumberland, Alston.
Hurolabis KiefFer, 189'2, p. 214.
63. H. comigans F. Loew, 1877, p. 11 (CecMfomyta);
Kieffer, 1913-2, p. 66.
(a) On Heracleum Spkondylium. Swanton, 667 ; Houard,
4512.
In all counties; common.
(•6) On H. SphoTidylittm, vat. anffuslifoUum.
Lancashire and Westmobbland, Grange-over-Sands
district.
{*c) On Lamium album. Houard, 4840.
Northumberland, Warkwortb and Ovingham. Lanca-
shire, Lathom. Durham, Biddick; Lambton Park and
Blaydon.
("rf) On Stachys sylvatica.
Durham, Penshaw.
This species is recorded by Houard from six different
Umbels and from three species of Lamium. Swanton (870)
(ref. Connold) records it also from Hieracium boreale, but
this is almost certain to be M. hieracii.
•64. H. hieracii Kieffer, 1888, p. 112.t
Recorded by Houard from Hieracium boreale, H. umbd-
bUum and other species of Hieracium. Apparently the
species (see above) recorded from H. boreale as M. corrugans
by Connold (Swanton, 870).
Durham, Hylton. Lancashire, on Hieracium sp.,
Birkdale.
•65. M. Uppocrepidis Kieffer, 1898-2, p. S9; 1913-2, p. 66.
On Hippocrejns amwsa. Houard, 3683.
Westmoreland, Meathop Fell, near Grange-over-Sands
in October, gall only, too late for larvae.
t We have aa ;et been unable to trace this in Kieffer, 1013-2,
360 Messrs. R. S. Bagnall and J. W. H. Harrison's
•66. M. marteU Kieffer, 1892 (-1), p. 215; 1913-2, p. 66.
On Hypericum kirsutum.
Durham, Middleton-one-Row.
67. M. piloseltae Binnie, 1877, p. 179 (Cecidem^);
Kieffer, 1913-2, p. 66.
On Hieracium Pihsdla. Swanton, 863 ; Houard, 6199.
Records from all counties excepting Cheshire ; local.
*68. H. stellaiiae Liebel, 1889, p. 282 (Cectdomyia];
Kieffer, 1913-2, p. 66.
On SlelUiria graminea. Recorded by Houard from
StdUiria medut.
Durham, Birtley.
Amotdla Kieffer, 1895, p. 7.
*69. A. quereicola Kieffer, 1909, p. 21 (gall and larva);
1913-2, p. 67.
On Quercus. Houard, 1211.
Northumberland, near Whitfield. Durham, neat Fat-
field.
70. A. querais Binnie, 1877, p. 179 (Cecidomtfia) ; Kieffer,
1913-2, p. 67.
On Qiiercus.
•71. Amoldia sp. Kieffer; Lem^e.
On Quercus. Houard, 1212.
Durham, West Cornforth.
Geocrypta Kieffer, 1913-2, p. 68.
•72. C. brauerl Handlirsch, 1884, p. 135 (Cecwfomyw).
A root-Ral! on HyjKricum jmlchrvm; recorded by
Houard (4213) from H. perforatum only. Apparently
very local.
Northumberland, near Staward. Durham, near Lan-
chester.
Rhabdophaca Westwood, 1847, p. 588.
•73. R. alUpennis H. Loew, 1850, p. 35 (Cecidomyia);
Kieifer, 1913-2. p. 70.
On Siilix spp. Houard, S. 32.
n,g,t,7.dt,'G00glc
Prdiminary Caialoffue of British CecidomyHae. 361
Apparently not uncommon; recorda from all counties
excepting Westmoreland. Also recorded by Mr. Burkill
from Derbyshire, and from Scotl^d.
*74. B. oUvUex Kiefler, 1892-2, p. 441 (Cecidmnyia);
1913-2, p. 70.
On Salix app. Houard, S. 14.
Lancashire, Ainsdale, rare.
•76. B. dubiosa Kieffer n. nov. for Cecidmnyia dvbia Kieffer,
1893-2, p. 255; 1913-2, p. 70 (= Cecidomyia
griseicoUis Zett. non Meigen).
On Salix spp. Houard, S. 41.
NoRTUCMBERLAND, Ninebanks, Warkworth. Durham,
Waldridge. Lancashire, near Ainadale.
*76. B. giraudlaUA Kieffer {salidperda Giraud n<m Dufour),
1898-1, p. 159; 1913-2, p. 70.
On Populus alba; also known from P. IremtiUi abroad.
Houard, 475.
Lancashire, Ainsdale and Freshfield, only two examples.
77. R. heterobta H. Loew (saliffna Hardy), 1850, p. 28
{Cecid<mifia); Kieffer, 1913-2, p. 70.
On Salic Bpp. Houard, S. 10; Swanton, 88, 89 and 154.
NoRTHUUBERLAND, Warkwoith, on S. alba. Durham,
Birtley, on 5. repeiu, also affecting the male catkins;
Waldridge, Swalwell ; Tinkler Fell, on S. Caprea. Lanca-
shire, Birkdale and Ainsdale.
•78. R. iteobia Kieffer, 1890-3, p. 201 (Cecidoniyia);
1915-2, p. 70.
On Salix cinerea. Houard, S. 11, Perrisia ile<Aia.
Northumberland, Ninebanks. Durham, near Blancb-
land. Lancashire, Ainsdale. Also from Scotland.
•79. H. kuHhl Kieffer, 1892-2, p. 251 (Cecidomyia);
1913-2, p. 70.
On Salix spp. Houard, S. 34.
Northumberland, Ninebanks, on S. cinerea, and
Warkworth, on S. dnerea and alba var. Durham, Birtley,
u,y,i,Ajh,Cooglc
362 Messrs. R. S. Bagnall and J. W. H. Harrison's
on S. repens, rare. Lancashire, Freehfield, on 5. cinerea.
Tare. Also recorded by Mr. Burkill (on 5. aJha) from
Derbyshire.
80. H. nurginemlorquens Winnertz, 1853, p. 223 (Ceddo-
myia); Kieffer, 1913-2, p. 70.
On Salix spp. Houard, S. 5\,PeTrisia marginemJorquens.
Generally distributed ; records from all counties except-
ing Westmoreland and Cheshire.
*81. B. nervorum Kieffer (nodnli Riibs.), 1895-2 {Dirhe-
hmyia); 1913-2, p. 71.
On Salix spp. Houard, S. 49.
Durham, River Wear near Cox Green, on 5. alba, Gibside
on S. aurita, and Ryhope on S. /ragUis, rare. Lascashibe,
Ainsdale and Freahfiefd, local, on young S. alba. North-
umberland, Oyinghatn and Warkwotth, on S. cinerea,
S. alba and the var. vUellina. Yorkshire, Nunthorpe on
8. viMlina, also see Burkill, 1916, " Entomologist," p. 7,
on S. Caprea. Also recorded by Mr. Burkill from Derby-
shire on 5. aiba and 5. cinerea, and from Surrey on iS.
Cajtrea.
•82. R. pleml Kieffer, 1896-2, p. 218 (Sertieria); 1913-2,
p. 71.
On Salix repens and S. cinerea. rare. Houard, S. 37.
Northumberland, near Warkworth. Durham, Birtiey,
lare.
*82a. B. pseudoeoecns, Riibsaamen 1890-2, p. 307 {Ced-
dotnyia]; Kieffer 1913-2, p. 71.
Flat, broad larva under a scale-like covering on under-
side of leaves of Salix Caprea, gregarious.
Durham. Plentiful on isolated trees in Ryhope Dene
and Hesleden Dene.
*82b. R. pulvlnl Kieffer {salidna Giraud, non Schrank;
khigi? H. Loew mm Meigen) 1891, p. 244 (Ceet-
domyia); 1913-2, p. 71.
On Salix anrita and S. litelUna. Houard, S. 19.
Northumberland, Ovingham.
n,g,t,7.dh,G00glc
Prdiminan/ Cataiogue <^ Brifish Cfcidomyidae 363
83. B. TosuU-f- H. Loew {saiicina Auct.. cinermrum
Hardy), 1800, p. 35; Kieffer, 1913-2, p. 71.
On Salix spp. Honard, S. 8.
Generally attributed ; records from alt counties.
*&1. R. mariella Kieffer, 1901-1, p. 494 (gall); 1913-2,
p. 71.
On Salix spp. Houard, S. 9.
Durham, \Valdridge, on S. aurila and cinerea; Gibside,
on S. repens. Lancashire, near Ainsdale, and Freshfield,
on S. repens. Also recorded by Mr. Burkill from Suirrey
on iS. repens .
85. R, saUeipeNa Lhifour {(erebram H. Loew), 1841, p. 262
[Cecidmnyia); Kieffer, 1913-2, p. 71.
On Salix spp. Houard, S. 36.
Northumberland, Newham. DintHAM, Billingham and
Waldridge. Cumberland, Alston. Lanca.shire, near
Ainsdale. Yorkshire, Eston.
86. R. lalicis Scbrank (degeeri Bremi, galhrvm saU<ns
Hardy), 1803, p. 69 (Tipula) ; Kieffer, 1913-2, p. 71.
On Salix spp. Houard, S. 40.
Records from all counties excepting Westmoreland and
Cheshire.
*87. R. snpema Kieffer, 1897, p. 24.
On Salix. Recorded by Houard, S. 26, from S. Caprea,
aurila and cinerea; we have not succeeded in tracing the
species in Kieffer, 1913-2.
Durham, on a hybrid S. cin«rea x vimifuilis; Billingham.
The larva is distinctive.
88. R. tarminaUs H. Loew, ia50, p. 3-5 (Cecidom\fia)\
Kieffer, 1913-2, p. 72.
On Salix spp. Houard, S. 14, Perrisia terminalis.
Common ; records from all counties.
[89. R. Mmirt«lw,Westwood, 1847, p. .588; Kieffer, 1913-2,
p. 72.
t The Daayaeura aaiicina of Swanton's Catalo^e must be re-
fCarded aa e, synonym, the gall being but't)ic incipient stage of the
" Roee-gall."
ih, Google
364 Messrs. B. S. Bagnall and J. W. H. Hairiaon's
PwrisU Rondani, 1846, p. 371.
•90. P. abieOpwda Henschel, 1880, p. 371 {Cectdomyia) ;
Kieffer, 1913-2, p. 73.
Gall on Picea ftrceisa. Houard, 10().
NoRTHUMBERLAKD, near Hexham. Durham, Gibside
and Eastgate. La^ica.shire, Grange-over-Sands, rare.
YoRKsHntB, Nunthorpe.
*91. P. aeereri^ans Kieffer, 1888-2, p. 266 (Cecidomyia) ;
1913-2, p. 73.
On sycamore. Houard, 3984.
Durham, dene near Fencehouses.
•92. P. acenrispans vat. rtibella Kieffer, 1896-2, p. 37.
On Acer campeslre. Houard, 4025.
Recorded by Mr. Burkill from Monmouthshire.
93. P. aerophiU Winnertz, 1853, p. 233 (C«-tffomy»a) ;
Kieffer, 1913-2, p. 73.
On Fraxinus. Swanton, 681 ; Houard, 4643.
Local ; records from all counties excepting Lancashire
and Cheshire.
94. P. annis Kieffer, 1886, p. 330 {Cecidomifia); 1913-2,
p. 73.
On Viola spp. Swanton. 632, 634, 635 and 637 ; Houard,
4281, 4283. 4284 and 4290.
Local ; records from all counties excepting Cheshire,
•95. P, alni F. Loew, 1877, p. 2 {Cecidotnyui) ; Kieffer,
1913-2, p. 73.
On Ahi'is. Houard, 1127; known throughout Europe.
NoRTHiMBERLAND, Ninebauks, once only.
96. P. alplna F. Loew, 18A>. p. 110 {Cecidomyia) ; Kieffer,
1913-2, p. 73.
On Silene acmtlig. Swanton, 344 ; Houard, 2274.
97. P. anylir., Kieffer, 19<»9. p. 31 (gall and lar\-a) ;
1913-2, p. 73.
On TfiocKiixiH ri/(,v-i(f'(«i, = Cecidomvid ep. Swanton,
676, and Houard, 4071.
ih, Google
pTdiminary Catalogue of British Cecidomyidae. 365
9S. P. apvlnes Kieffer, 1889, p. 208 (Cecidomyia);
1913-2, p. 73.
On Galium Aparine. Swanton. 763 ; Houard, 5303.
Northumberland, Wylam, Belford. Durham, several
localities. Cheshire, Bidston.
*99. P. aueuparla KiefFer.f
On Pynis Auctiparia. Houard, 2907.
Northumberland, Ovingham. Durham, Gibside. Also
from Scotland.
*I00. P. uiUaris Kieffer, 1896-1, p. 87; 1913-2, p. 74.
On Trifdtum sp. On T. medium Houard, 3593.
Lancashire and Westmoreland, once near Grange-
over-Sands. Durham, coast near Horden; Birtley, on
T. froffiferum.
*]01. P. IwlloUa Riibsaamen.
On BaUota nigra, flowers remaining closed.
Northumberland, Denton Burn (H, S. Wallace and
R. S. B.).
•102. P. bACkUna Mik, 1885, p. 140 {6V«fo»nyw); Kieffer,
1913-2, p. 74.
On Inula squarrosa (= /. Conyza). Houard, 5623.
Lancashire, Grange-over-Sands. J. W. H. H. has ob-
served similar galls on Pulicaria dysenlerica at Cowpen
Bewley in Durham.
103. P. bfMSleae Winnertz, 1853, p. 213 [Cecidojntfia) ;
Kieffer, 1913-2, p. 74.
On Brassica spp. Swanton, 415, 420; Houard, 2591
[Dasyneura brassicae).
Durham, Hylton, on *Brassica oleracen (Houard, 2573).
•104. P. brnnellae Kieffer, 1909, p. 3 (gall and larva);
1913-2, p. 74.
On Prunella rtilgaris, = Oecidomyid sp. Houard, 1818.
Northumberland, O^nngton. Yorkshire, Stainton.
t We have been unable to trace tliia in Kieflcr, 1913-2.
TRAKa.EHT, SOC. LOND. 1917.— PARTS II, III, IV. (MAY '18) Bt
u,y,i,Ajh,Cooglc
S66 Messra. R. S. Bagnall and J. W. H. Harrison's
*I05. P. bryonlae BoucM, 1847, p. 144 {Cecidom^);
Kiefter, 1913-2, p. 74.
On Bryonia dioka. Houard, 5476.
Durham, near Lamesley.
''lOG. P. eampanulae Riibsaatnen.
On Campnnula media and C. fersicaefalia.
Durham, Bittley. Cumberland, Alston.
107. P. capitigena Bremi (Eiipkorbiae paitim H. Loew),
1847, p. 50 (Cecidomyin); Kieffer, 1913-2, p. 74.
On Euphorbia Esiiia. Swanton, 597.
108. P. earil»mines Winnertz. 1853, p. 225 (Cecidmttyia);
Kieffer, 1913-2. p, 74.
On Cardamine spp. Swanton, 390 and 393; Houard,
2672 and 2G65.
Durham, in the Derwent Valley.
109. P. eerasUi Binnie, 1877, p. 181 {Cecidomyia); Kieffer,
1913-2, p. 75.
On Ceraslium ridgalum. Swanton, 348; Houard, 2338.
North iMBER LAND, Warkworth, on *C. semidfcandntm
and C. ruigatiim. Durham, Ryhope Dene.
*110. P. elTsU Riibsaanien. 1890. p. 236 {Cecidomyia);
Kieffer. 1913-2, p. 75.
In heads of i'irsium nrrenne. Houard- 5927.
Not uneoninioii ; reconls from all counties.
•111. Perrista sp. Rignall and Harrison. 1916 (iii). p.l99-
In heads of Cir.-'i'iiii arveiixe. like P. cirsii, but larvae
vellowish to orange instead of red ( i form or stage of
P. arsii).
Not uneonminn; nivnls from all counties excepting
Cliesliire.
•112. Perrlsia sp. Ri-juall and Harrison, 1916 (iii). p. 199.
Heads of Vifsi'ni' iiri\n.''e cWt'd ; a (ew petals to one
side only brfiikiiiw dul ; larva solitary, deep vermilion to
NoBTiu MUKRLAND. Nint'Kuiks. Di-RHAM, Pensbaw,
l-attU-ld. and iiiUide.
n,g,t,7.dh,G00glc
Prdimittary Catalogue of British Cecidomyidae. 367
*113. P. Mtnrositanun KieSer, 1888-2. p. 310 Ifiecidmnyia);
1913-2, p. 75.
Heads of Viraium. lanceolatum. Houard, 5911.
NoRTHtfMBERLAND and DuRHAM, numerous records.
LANCA.smRE, Lathom. Yorkshire, ilarton ; Bardjtey near
Leeds.
•114. Perriala sp. Bagnall and Harrieon, 1916 (iii). p. 199.
Heads of Cirsium Uinceolatum. heads remaining closed
or not opening fully; often bent or twisted; florets
apparently normal; larvae wtiite to yellowish.
Records from Northumberland, Durham, Cumberland,
Yorkshire and Lancashire.
*115. P. wrylina KiefTer for cwyli Riibsaamen, 1912,
p. 288; Kieffer, 1913-2, p. 75.
In catkins of Corylus.
Lancashire, Grange-over-Sands, October. Yorkshire,
Guiflbrough, October.
116. P. cralaegl Winnertz, 1853, p. 238 (Ceridomyia);
Kieffer, 1913-2, p. 75.
On Crala^uB monogifna. Swanton, 540 ; Houard, 2942.
Common everywhere ; records from all counties.
*117. P. daphnes Kieffer, 1901-2, p. 18; 1913-2, p. 75.
On Daphne Laureoh. Houard, 4312.
Yorkshire, Gunnergate.
•118. P. engstfeltU Rubsaamen, 1889, p. 375 (Cecidomyia) ;
Kieffer, 1913-2, p. 75.
On Spiraea Vlmaria Houard, 2832 and 2837.
Local but widely distributed ; records from all counties.
Also from Scotland.
•119. Perrisla sp. Bagnall and Harrison, 1916 (iii), p. 200.
On Spiraea Vlmaria, on the underside of leaf, like P.
engstfddi, but always adjacent to midrib or nerve, which
is considerably swollen in such manner as to become a
shelter to the larva.
Extremely local, but plentiful where it occurs. Durham,
Gibside and near Lamesley; Billingham. Cumberland,
Alston.
u,y,i,Ajh,Cooglc
368 Messrs. R. S. Bagnall and J. W. H. HamBon's
120. P. epIIobU F. Loew, 1889, p. 201 (Cccitfomyia) ;
Kieffer, 1913-2^ p. 76.
On Ejnldbium anffuslifoUum. Swanton, 641, 642;
Houard, 4345.
NoETHUMBEBLAND, Seahouses, Spindlestone, Budle Bay,
etc. Durham, Gibside.
*121. P. erleins F. Loew, 1885-2. p. 76 {Cedd^myia);
Kieffer, 1913-2, p. 76.
On Erica dnerea.
Durham, Waldridge. Yorkshire, Great Ayton Moor.
121a. p. flUclns Kieffer, 1889, p. 191 (Cecidomyia);
1913-2, p. 76,
On Pteris nqmlina. Swanton, 3; Houard, 68.
Records from all counties excepting Cumberland. ■
•122. P. florlperda F. Loew, 1888, p. 231 (Cecidomyia);
Kieffer, 1913-2, p. 76.
On Sifene infiala. Houard, 2261.
Durham, Fatfield and Fencehouaee.
*123. P. nosculorum Kieffer, 1890-3, p. 200 {Cecidomyia);
1913-2, p. 76.
Flower of Trifolium pralense. Houard, 3579,
Durham, Penshaw. Lancashire and Westmoreland,
near (■ range-over- Sands.
124. P. fraxinea Kieffer, 1907-1, p. 523; 1913-2, p. 76.
On Fraj-iiuis,^ Dasyneura /raxinea, Swanton, 678;
Houard, 4647.
Apparently widely distributed ; records from all counties.
125. P. rraxlDi Kieffer, 1897-2, p. 301 ; 1913-2, p. 76.
On Fraxiniis. Swanton, 679, 680; Houard, 4644.
Records from all counties excepting Cheshire.
*126. p. Inicluum Riibsaamen, 1895, p. 258 (Dichdmnyid) ;
Kieffer, 1913-2, p. 76.
On Cerasliiim vidgainm. Houard, 2330.
Northumberland, Warkworth, Ovingham. Durhau,
Fatfield, Penshaw, Birtley, Greatham. Cheshire, Bidston.
l„y,l,Ajh,COOl^lC
Prdiminary Catalogue of British Oeddomytdae, 369
Yorkshire, Marton, Cleveland district; Leeds district.
Also taketi (at Warkworth) on Geraslium semidecandrum,
and in Cleveland on C. viscos^im.
127. P. galeobdoUmtis Winnertz, 1853. p, 238 (Cecidomyia) ;
Kieffer, 1913-2, p. 76.
On Lamium Galeobddon. Swanton, 716; Houard, 4847.
128. P. galU H. Loew, 1850, p. 37 (Cecidomyia]'; Kieffer,
1913-2, p. 77.
On Gcdium venim. Swanton, 747, 748; Houard, 5284,
5292.
Northumberland, Seahouses, Bamburgh, Warkworth,
Ovingham. Durham, Blackball Rocks; Hylton; Pensliaw
Hill; near Cornforth. Lancashire, Birkdale, one only.
On Galium vliginosum. Houard, 5268.
Durham, Waldridge Fell.
•129. Perrisla ap. Schlechteiidal, etc.
On Galium verum. Houard, 5289.
Durham, on one patch of bed-straw near Penshaw.
130. P. gaUloola F. Loew, 1880, p. 33; Kieffer, 1913-2,
p. 77.
On Galium spp. Swanton, 7.53; Houard, 5209, also
•5286 and •5275.
Northumberland, Bamburgh, Warkworth. Durham,
coast near Hartlepool, Penshaw Hill and Catcleugh ; rock
neat Sunderland, on G. verum. Cumberland, Alston, on
G. verwrn.
•131. P. genUtamtorquens Kieffer, 1888-2, p. 311 (Cm-
domyia); 1913-2, p. 77.
On Genista Hncloria. Houard, 3371.
Durham, Oibside, rare.
132. P. genistleola F. Loew, 1877, p. 4 {Cecidomyia);
Kieffer, 1913-2, p. 77.
On Genista titiOoria. Swanton, 548, 549; Houard, 3368,
3369, It also occurs on G. anglicn {Swanton, 547 ; Houard,
3349).
Northumberland, Ovingham, Durham, Gibside.
u,y,i,Ajh,Cooglc
370 Messis. R. S. Bagnall and J. W. H. Harrison's
*133. P. KMittaiiu Kieffer, 1909, p. 9 (gall); 1913-2. p. 77.
In flowers and seed-caaea of Genllana campestris and
G. ArtmrdUi. ~ Cecidompd sp. Houard, 4696.
North UMBERLASD, between Alnmouth and AVarkworth.
DoiHAM, Peiishaw Hill and Blackball Rocks. (This species
is recorded in Houard (4696) from British Isles only, but
in Kieffer {1913-2) it is recorded from Central Europe.
Examples from seed-caees cause chloranthia as well ae a
swelling of the ovary, and are possibly referable to a
second species.
• *134. P. Beranil Kieffer, 1907-2, p. 44; 1913-2, p. 77.
We have not cleared up the question of CWidoniyids
affecting Geraniaceae. The records here refer to P. geranii
on Erodium cicularium only (Houard, 3826); but we
append also our notes under the names of various species
of Geranium.
Northumberland, Budle Bay, very common; Wark-
worth. Durham, Cowpen Bewley- one only. LA^■CA.SHIBB,
Freshfield, two only, Yorkshire, Redcar.
Other notes :—
(1) Geranium •prntense.
(fl) Yellow larvae in seeds (Dasyneura geranii).
Northumberland, Budle Bay and Ninebanks.
Durham, Birtley, Eastgate, Wolsingham and Horden.
Cumberland, Alston.
(6) Bright orange-red larvae in seeds with above.
Northumberland and Durham, all above records,
(2) Geranium sylialiaim.
(a) Yellow larvae in seeds (Dasyneura geranii).
(6) Bright orange-red larvae in seeds in company.
Records from Northumberland and Durham.
(3) Geranium sanguimnm.
Yellow larva in seeds,
Durham, Horden and Blackball Rocks. York-
shire, in a moraine garden, Linthorpe.
iM-,i,A J h, Google
Prdiminary Catalogue of British Ceddomyidae. 371
(4) Geranium disseclum.
Yellow larva in seeds.
Durham, Fatfield.
(5) Geranium molle.
Yellow larva in seeds.
YoEKSHraE, Redcar.
(6) Geranium pusilliim.
Yellow larva in seeds.
NoETHUMBERLAND, Warkwoith.
(7) Geranium pusilliim.
One or more transparent, lemon -yellow larvae
amongst seeds and in the flower.
Lancashire, Freshfield and Ainsdale, common.
(8) Erodium cicHlanum.
Cream ish-yellow larvae amongst seeds and in the
flower.
Yorkshire, Redcar. Northumberland, Wark-
worth.
135. P. gUchonuK Kiefter, 1889-2, p. 2C3 {Cecidomyia) ;
1913-2, p. 77.
On Glechoma hederacea. Swanton, 7(17, 708; Houard,
48D7, 4808.
•136.1 P. glycipkylli Rubsaanien, 1912, p. 286; Kieffer,
1913-2, p. 77.
On Astragalus hypoghtlis.
Records from Scotland.
*137. P. holDst«M Kieffcr, 19()9. p. 29 (gall and larva);
1913-2, p. 77.
On ^SteUaria graminea.
Northumberland, between Langley Woods and Whit-
field.
On Stdhria Holoslea.
Northumberland, Stocksfield (H. S. Wallace). Dur-
ham, Gibside, Winlaton Mill,
ll,y,l,7rJM,G00glc
372 Messrs. R. S. Bagnall and J. W. H. HarriBOn's
*138. P. hyKTophlU Mik, 1883, p. 209 {Ceddomyia);
Kieffer, 1913-2, p. 77.
On Galium palustre. Houard, 5278.
Northumberland, Ninebanks. Durham, near Gibside,
Low Fell and Waldridge. Lanca.shire, not uncommon,
Freshfield, Ainsdale aod Lathom. Yorkshire, Nunthorpe ;
Bardsey near Leeds; also recorded by Mr. Burkill. Mr.
Burkill also records this species from Derbyshire, Stafiord-
shire and Surrey,
139. Perrisia sp. Trail, 1878.
On Galium- palvslre. Swanton, 758; Houard, 5277.
140. P. hyperiei Bremi, 1847, p, 53 {Ceddomyia); Kiefier,
1913-2, p. 77,
On Hypericum spp. Swanton and Houard. Recent
records from H. ■perjoraium, H. pulckrum, H. hutnifiisum
and * H. monlanvm.
Records from all counties excepting Cheshire and York-
shire.
*141. Perristasp. Burkill, 1916.
On Hypericum elodes. Terminal leaves thickened at the
base, folded over and crinkled, turning brown at the tips.
Larvae yellow, several in each gall.
Yorkshire, Mr. Burkill's record.
142. P. ignorata Wachtl (medicaginis Bremi. gall; ««>-
brychidis F. Loew mom Bremi), 1884, p. 163
(Ceddomyia); KiefTer, 1913-2, p. 77.
On Medicago salim. Swanton, 561 ; Houard, 3515,
143. P. Inchbaldiana Mik (1 clavdlia Bremi), 1886, p. 317
{Ceddomyia); Kieffer, 1913-2, p. 77.
On Salix alba. Houard, S. 52; Swanton, 98; Houard,
627.
Records from all counties.
144. P. loclusa Frauenfeld, 1862, p. 1175 {Ceddomyia);
Kieffer, 1913-2, p. 78.
On PhragmUes communis. Swanton, 50;* Houard, 245.
Durham, Bilhngham.
n,g,t,7rJM,GOOglC
Prdiminary Caialogue of British Ceddomyidae. 373
*145. P. WeBeri Marchal, 1896, p. 99 ; Kieffer, 1913-2, p. 78.
On Hedara Helix. Houard, 4362.
Northumberland, Warkworth district. Lancasbibe
and Westmoreland, Grange-over-Sanda district, common.
Durham, Oibside and Low Fell. Yorkshire, Middles-
brough and Gunnergate; Leeds. Also from Scotland.
*146. P. klefferlana Riibsaamen, 1891, p. 5 (Cecidomyia);
Kieffer, 1913-2, p; 78.
On EjnUjbium angustifoUum. Houard, 4348.
Durham, Gibside, rare. Yorkshire, see Burkill. Also
recorded by Mr. Burkill from Surrey.
*147. fP- lamii Kieffer.
On Lamium macuUUum, = Cecidomyid sp. Houard,
4836.
Durham, Birtley, in a garden.
•148. ; P. Uriels F. Loew, 1878, p. 393 [Ceddomyia) ;
Kieffer, 1913-2, p. 78.
On Larix decidua. Houard, 86.
Durham, Lanchester, Winlaton Mill ; Gibside. Lasca-
SHiRE, Grange-over-Sanda. Yorkshire, Eaton.
*149. P. Uthyri Kieffer, 1909, p. 13 (gall and larvae);
1913-2, p, 78.
On Lathyrus pralensis, = Perrisia sp. Houard, 3775.
Records from all countiea excepting Cheahire.
*150. P. lathyrlools Riibsaameu, 1890, p. 26 {Cecidomyia) ;
Kieffer, 1913-2, p. 78.
On Lathj/THS pratensis. Houard, 3771.
Records from all counties.
151. Perrisia sp. (lalhyricola Swanton).
On Lathyrus praiensis. Swanton, 595 ; Houard, 3776.
Records from all counties.
t We have been unable to trace this name in Kiefler, 1913-2.
I = KeOne-Ti HeoBchel (gall).
iM-,i,A J h, Google
374 Messrs. R. S. Bagnall and J. W. H. Harrison's
•152. Petriiia sp. Burkill, 1917, p. 83.
On Laikt/nis -praleim*.
Recorded by Mr. Burkill from W. Gloucestershire.
*153. P. Utorm Kieffer, 1909, p. 21 (gall); 1913-2, p. 78.
On Oak, = C«cidom)-id sp. Houard, 1310,
Northumberland, Warkworth; Ovingham. Dckham,
several records. Lancashire, Granfje-over-Sands. York-
shire, Leeds district and near Middlesbro'.
*153a. p. Utho^ermi H. Loew, 1850, p. 36 {Ceddonufta);
Kieffer. 1913-2, p. 78.
On LUhospennum officinale. Houard, 4741,
Northumberland, Ovingham; on an isolated patch of
the host- plant.
154. P. lotharli^as Kieffer, 1888-1, p. 107 (Cecidmnyta);
1913-2, p. 78.
On CerasHum viilgalum. Swanton, 346, 347; Houard,
2331,2334.
Records from all counties excepting Cheshire.
♦155. P. loti Kieffer, 1909, p. 14; 1913-2, p. 78.
On Lofus comicuhlMs, — Cecidoniyid sp. Houard, 3622.
Durham, Gibside, apparently rare.
156, P. loUeoU Rubsaamen, 1889-2, p. 52 {Cecidomyia);
Kieffer, 1913-2, p. 78.
On Loins major and occasionally *L. comiathtia.
Swanton, 584; Houard, 3626, *3010.
Records from all counties excepting Westmoreland,
*157. P. lupuUnaa Kieffer, 1891, p. 258; 1913-2, p. 78.
On Medicngo lupuUna. Houard, 3507.
Durham, near Burnmoor.
•158. PerrUa sp. Kaltenbach and others.
On Medi<vi<fo liipuUna. Houard, 3509.
Durham, Hylton and Penshaw. Recorded by Mr. Burkill
from W. Gloucestershire.
ll,y,l,/eJh,G00t5lc
Prdiminary Catalogue of British Cecidomyidae. 375
•159. P. lyehnidis Hevden, 1861, p. 98 {Ceddmnyia) ;
Kieffer, 1913-2, p. 78.
On Lychnis alba. Houai^, 2292.
Northumberland, coast near Hamburgh. Durham,
Lamesley, Fencehouses. Lancashisb, Ainsdale.
160. P. nuU Kieffer, 1904-2, p. 345; 1913-2, p. 78.
On Pyrus Mains. Swanton, D33; Houard, 2885-
DuRHAM, near Stanley.
•161. P. malplgUi Kieffer, 1909, p. 21 (gall) ; 1913-2, p. 78.
On Quercus. Houard, 1354.
Northumberland, Warkworth, rare. Durham, Gibside,
Laxca.shire, Grange-over-Sande. Yorkshire, Leeds;
Bardfiey.
•162. P. miki Kieffer, 1909, p. 6 (gall); 1913-2, p. 78.
On (a) CerUaurea nigra and (6) C. Scabiosa, = Cecidomyid
sp. Houard, 5981.
Northumberland, near Staward (a). Durham, Fat-
field (a), Billingham (6), Hylton (b), and Gibside (a).
Cumberland, near Nenthead (a). Lancashire and West-
moreland, near Grange-over-Sands (a). Yorkshire,
Mar ton.
•163. Perrtola sp. Bagnall and Harrison, 1916 (iii), p. 199.
Head of (n) Centavrea nigra swollen, breaking round
edges; larvae in large numbers, yellowish to pink. Also
in (6) C. Scabiosa.
Northumberland and Durham, several records in (a)
and (6). Cheshire, Ridston Hill, plentiful (a). Cumber-
land, Nenthead (o).
•164. Perrlsia sp. Bagnall and Harrison, 1917 (v), p, 14.
In old heads of Centaiirea Scabiosa, larvae gregarious,
orange-red to copper-red, small ; not the above species.
Northumberland, Warkworth. Durham, Catcleugh
Rock, near Sunderland ; Ryhope ; Fulwell ; Eaaington.
Also in old beads of C. nigra.
iM-,i,A J h, Google
376 Mesera. R. S. Bagnall and J. W. H. HamBon's
*105. PerrisU ap. Bagiiall and Harrison, 1917 (v), p. 14.
In a garden Hdianthus; gregarious larvae in heads,
golden -yellow-orange to orange -red.
Durham, Fatfield.
106. P. muricatae Meade {caricia H. Loew), 1886, p. 153
(Cectdotnyia); Kieffer, 1913-2, p. 79.
In utricle of Cnrex viilpina, C. contxgua, *C. petuiula,
C. hetrigata, C sylvalica and C palkscms. Swanton, 69,
70; Houard, 361, 362.
Northumberland, Warkworth, on C pendula. Dub-
ham, Gibside, on C. pendula, C. sylvalica and C. paUescens;
Waldridge, on C. laevigata ; Billingham. Lancashire, near
Grange-over-Sands, on C. vulpina.
*167. P. nenrioola Kieffer, 1909, p. 9 (gall); 1913-2, p. 79.
On Hieracirim Pihsella, = Cecidomyid ep. Houard, 6205.
Durham, very local, Birtley Fell, Roker. Yobkshirk,
Huddersaeld district (W. Falconer).
168. P. otwhrychidis (giraudi Prauenf.) Bremi, 1847, p. 53
{Cecidotnyut); Kieffer, 1913-2, p. 79.
On Astragalus danicus. Swanton, 585; Houard, 3646.
Probably P. glyciphylU.
•168a. p. pantoU Kieffer, 1909, p. 21 (gall); 1913-2, p. 79-
On Oak. Houard (Cecidomyid), 1315.
Northumberland, Ovingham, 1 only (H. S, Wallace).
Yorkshire, Leeds, plentiful on one old tree.
169. P. papaverU Winnertz, 1853, p. 229 (Cecid(mpi);
Kiefier, 1913-2, p. 79.
In capsules of Papaier Rkoeas. Swanton, 377 ; Houard,
2479,
Durham, an isolated record, in field near Hylton.
*170. P. perlclymeni Rubsaamen, 1889-2, p. 54 (Cecido-
mijia); Kieffer, 1913-2, p. 79.
On Lonwera Pcrkhjmenum. Houard, 5361.
Northumberland, Warkworth; Ovingham. Durham,
Gibside (Barry Stewart and R. S. B.), Waldridge; Urpeth;
Winlaton; Foncehouses and FasingtOQ. Cueshibb,
Bidaton Hill. Yorkshire, Bardsey, near Leeds.
iM-,i,A J h, Google
Prdimitiary Calahgue of British Cecidomyidae. 377
171. P. penleulae Linn4, 1767, p. 977 (Tipula); KieSer,
1913-2, p. 79.
On Polygonum ampkOnum. Swanton, 321, 325 ; Houard,
2157, 2159.
BoBHAM, Urpeth.
172. P. pUwtrIx ft. Loew, 1850, p.
Kieffer, 1913-2, p. 79.
On Rubm spp. Swanton, 479, 482, 490; Houard, 2966,
2978, 302D.
In some districts very common ; records from all countiea,
•173. P. polygalM Kieffer, 1909, p. 18 (gall); 1913-2, p. 79.
In Polygala vulgaris, larvae in flower. See Houard.
3855.
Durham, Birtley, Horden, Lanchester district (H. S.
Wallace and J. W. H. H.). Cu.mberland, Alston,
•174. P. popDletl Rubsaamen, 1889-2, p. 57 {Cecidomyia) ;
Kieffer, 1913-2, p. 79.
Od Poptdm tr^nuh and. P. IremuUte x alba. Houard, 503.
Durham, Gibside and near Winlaton Mill. Nobthum-
BBRLAND, on the Alnwick road four milea north of Morpeth,
175. P. polerUillae Wachtl, 1885, p. 193 {Cectdomyia);
Kieffer, 1913-2, p. 79.
On PotenlilUi.
•176. P. pratfooU Kieffer, 1892-1, p. 216 {Cendomyia);
1913-2, p. 79,
In flower Lychnis Flos-cuaiH. Houard, 2290.
Durham, Gibside, Billingham.
*177. P. ptoridioola Kieffer, 1901-2, p. 19 (Cecidomyia);
1913-2, p. 80.
On Pleris aquiUna, local. Houard, 69.
Northumberland, Ninebanks. Durham, Fatfield, Birt-
ley.
•178. P. puilulani Riibsaamen, 1889-1, p. 378 {Cecidomyia);
Kieffer, 1913-2, p. 80.
On Spiraea TJlmaria, local. Houard, 2838.
n,g,t,7.dh,G00glc
378 MesBTB. R. S. Bagnall and J. W. H. HamBon's
Eecords from all counties excepting Lancashire and
Westmoreland. Mr. Butkill records it from Derbyshire.
179. P. pyri Bouchfi, 1847, p. 144 (Ceddomtjia) ; Kiefier,
1913-2, p. 80.
On Pyru.* communis. Swanton, 526; Houard, 2864.
Durham, Gibside. Lancashire, Lathom.
•180. f P. plerreana Kieffer.
On Salix cinerea, =^ Perrisia sp. Houard (S. 42), 892.
DuKUAM, Waldridge. Lancashire, near Ainsdale.
181. P. nBimeuli Bremi, 1847, p. 54 (Cecidomyia) ;
Kieffer, 1913-2, p. 80.
X)a Ranunciilus spp. Swanton, 362, 366, 370; Houard,
2423, 2431, 2438.
Records from all counties excepting Cheshire and
Cumberland.
*182. P. raphanlstri Kieffer, 1886, p. 324 {Cecidomifia) ;
1913-2, p. 80.
On Brassica Napus. Houard, 2594 {Dasyneina rapka-
nislri).
Northumberland, Warkworth. Durham, rare and
local, Fatfield, Fencehonsea and Penshaw.
*183. P. rhododeQdroni Kieffer, 1909, p. 95; 1913-2, p. 80.
On Rhododendron ferrugineym. Houard.
YoRK.SHiBE, in a moraine garden at Linthorpe, near
Middlesbrough.
184. P. rosarum Hardy, 1850, p. 186 {Cecidomyia);
Kiefier, 1913-2, p. 80.
On Bosn spp. Swanton, 503, 507, 512; Houard, 3135,
3186.
Records from all counties excepting Westmoreland and
Cheshire.
•185. Perrtda sp.
On Sosu Bpp. like P. rosarum gall, larvae white.
Northumberland and Durham, several records
(= Dirkha rhodophiUt).
-f We hAve been unable to trace this in Kiefler, 1913-2.
l„y,l,Ajh,COOl^lC
Prelxminary CaUdogue of British Ceddomyidae. 379
•186. p. roBtruplans Kieffer, 1909, p. 29 (gall and larva);
1913-2, p, 80.
On Spiraea Vlmaria, = Cecidomyid 8p. Houaid, 2836.
Not common ; records from all counties excepting
Cheshire.
•187. ?P. rubsauneni Kieffer, 1909, p. 6 (gall); 1913-2,
p. 80.
On Carpinus Beiuhts, parenchymatous, = Cecidomyid sp.
Houard, 1041.
Circular clear-cut holes observed in leaves of honibeara
(counties Durham and Lancashire) in October are prob-
ably the results of this insect. Requires confirmation.
•188. P. saUeariM Kieffer, 1888-1, p. 96 (Cecidomyia) ;
1913-2, p. 80.
On Lythrum Salicaria. Houard, 4325, 4326.
Northumberland, not uncommon near Bamburgh.
•189. P. sangulsorbae Kieffer, 1890-1, p. 26 (Ceddomyia) ;
1913-2, p. 80.
On Sanguisorba officinalis. Houard, 3100.
Durham, near Penshaw.
•190. P. scsUosae Kieffer, 1888-1, p. 97 (Ceddmnyia);
1913-2, p. 81.
On Scabiosa C<Aumbaria, Houard, G466, 5470.
Durham, Penshaw Hill, common ; and Catcleugh Rock,
near Sunderland.
•191. P. schlechtendali Kieffer, 1886, p. 328 Ifieddomyia);
1913-2, p. 81.
On Lathyrus macrorhizus. Houard, 3781.
Durham, Birtley ; Fatfield ; Fencehouses ; Gibside, com-
mon; Winlaton (H. S. Wallace), and near Lanchester.
Northumberland, Ovinghara.
•192. P. sebmldU Rubsaamen, 1912, p. 284; Kieffer,
1913-2, p. 81.
Heads of PlatUago lanceohla.
Durham, Greatham. Lancashire, Grange-over-Sands.
ih, Google
380 Mesere. R. S. Bagnali and J. W. H. HarriBon's
193. P. sarotina Winnertz, 1853, p. 316 {Ceddmnyia);
Kieffer, 1913-2, p. 81.
On Hypericum spp. Swanton; Houard.
NoETHUMBERLAND, Dear Staward and Ovingham, on
H. ptdchrum; Warkworth, on H. kumifusum. Durham.
near Lanchester and Winlaton Mill, on H. pulchmm;
Gibside, on H. perforatum and H. pukhrum; Dinsdale and
Castle Eden, on H. kirsutum. Lascashire, Hampsfell,
Grange-over-Sands, on H. pukhrum, very local. Yobk-
SHiRB, Bardsey, near Leeds.
*194. P. sUvleola KieSer, 1909, p. 30 (gall and larva);
1913-2, p. 81.
On *StdUtria graminea.
Northumberland, between Langley Woods and Whit-
field.
On SteUaria Holoslea = Perrtsta sp. Houard, 2311.
Northumberland, Warkworth; Ovingham. Durham,
Gibside and Easington. Yorkshire, Bardsey, near
Leeds.
*195. P. similis F. Loew, 1888, -p. 232 {Ceddomyia);
Kieffer, 1913-2, p. 81.
On Veronuxi spp.
Northumberland, Warkwortb, on V. officinalis and
V. Chamaedri/s. Durham, Gibside, on V. scutellala and
V. officinalis ; Lanchester and Stanhope, on V. officinalis;
Castle Eden and Birtley, on V. Chatmedrys.
196. P. sisymbrii Schranck, 1803, p. 83 {Tipula); Kieffer,
1913-2, p. 81.
On various Crucifers. Swanton; Houard.
Durham, Swalwell, on Sisymbrium officinale.
•197. ? P. sodalis F. Loew, 1877, p. 7 {Cecidomyia);
Kieffer, 1913-2, p. 81.
On Prunus. P. tortrix and this species produce identical
galls, and it will be necessary to breed out specimeua
before this can be definitely included in our Ust. See
records under P. tortrix.
,t,7rJh,G00glc
Prdiminary Cgtfdogue of British Cecidomytdae. 381
198. P. staeliydtc Bremi, 1847, p. 55 {Cecidomyia) ; Kieffer,
1913-2, p. 81.
On Stachys sylvaltca and ^palustris. Swanton, 711-714;
Houard, 4860-62.
Records from all counties excepting Westmoreland.
*199. Perrmo sp. Burkill, 1916.
On Stachys palnstris ; buds galled by white larvae.
Recorded by Mr. Burkill from Derbyshire.
*200. P. strobi Winnertz, 1853, p. 234 {Ceddomyia) ;
Kiefler, 1913-2, p. 81.
In cones of Picea excdsa.
Northumberland, near Corbridge. Mr. Evans records
this species from Perthshire (" Ent. M. Mag.," 1909, p. 17).
•201. P. tetend Rubsaamen, 1892, p. 400 (Ceddomyia);
Kieffer, 1913-2, p. 82,
On leaves of Ribes app.
Durham, Gibside, on R. GrossuUiria; Birtley, on R. nigra.
•202. P. tetrahit Kieffer, 1909, p. 8 (gall and larva) ; 1913-2,
p, 82.
In GaUopsis Telrahil flowers, = Perrisia sp. Houard,
4831.
Lancashire, near Lathom.
•203. P. thomulana Kiefier, 1888-1, p. 95 (Cecidotnyia);
1913-2, p. 82.
On Tilia vulgaris and platyphylloa. Houard, 4124, 4155.
Northumberland, Ovingham, Warkworth. Durham,
several records. Cumberland, Keswick and Alston.
Lancashire, Grange-over-Sands. Yorkshire, Ounner-
gate. Also from Scotland.
•204. Perrisia sp. Bagnall and Harrison, 1917 (v), p. 14.
On TUia, like P. ihmnasiana, but larva milk-white.
NoBTHUMBEELAND, Plesscy. DuRHAM, near Chester-le-
Street; Stanhope.
TRANS. BNT. 8O0. LOND. 1917. — PARTS H, HI, IV. (MAY '18) CO
ih, Google
382 Messrs. R. S. Bagnall and J. W. H. Harrison's
205. P. tlliamvolvens (tiliae Schr., gall), RiibsaameD,
1889-2, p. 55 {Cecidwnyia) ; Kieffer, 1913-2, p. 82.
On TUia ap. Swanton, 618; Houard, 4160.
Durham, Castle Eden Dene.
*206. P. tortrlx F. Loew, 1877, p. 6 (Cectdomyia) ; Kieffer.
1913-2, p. 82.
On (a) Prunus domealica, wild, and (t) P. spinosa.
Houard, 3269, 3274, 3282.
NoBTHUMBFRLAND, near Minsteracrea, {a) H. 3269 and
3274.t YoBKSHiEE, Bardsey near Leeds. Lancashire,
gall (a) H. 3274 only, near Lindale; Blackburn diatrict
(G. W. Nixon), perhaps referable to P. sodalis or both
species. Mr. Burkill, 1916, records this species from P,
spinosa, Devon,
207. P. traehelif Wachtl, 1885, p. 195 {Ceddotngia) ;
Kieffer, 1913-2, p. 82.
On Campanula rolundifdia. Swanton, 796, 797 ; Houard,
5513.
Durham, near Wolsingham.
•208. P. (reili Kieffer, 1909, p. 25 {gall); 1913-2, p. 82.
On Ranunculus acris, = Cecidomyid sp. Houard, 2420,
Northumberland, Ovingham. Durham, near Penshaw.
209. P. (rifolii F. Loew, 1874, p. 143 {Cecidomyia] ;
Kieffer, 1913-2, p. 82.
Onrri/o/iuwspp. Swanton, 568, 569, 572, 577 ; Houard,
3589-91-96-64.
Records from all counties excepting Westmoreland.
*210. Perrisia sp. Burkill, 1916.
On TrifoUnm pralense. leaflets folded into a pod resem-
bling the gall caused by P. trlfoUi, but each gall is occupied
by a white larva.
Durham, ncarHylton. Northumberland, Warkworth.
Recorded by Mr. Burkill from Derbyshire.
t Ptrrieia sodalis may alxo occur in the gail 32T4, but the presence
of 3269 enables one to definitely record P. lortrix.
u,y,l,7rJM,G00glc
Preliminary Catalogue of British Ceddomyidae. 383
*211. Perrisia sp. Burkill, 1916.
On Trifolium repens, leaves thickened and forming a
firm Seshy gall, with aborted buds inside, among which
live several larvae.
Recorded by Mr. Burkill from Surrey.
212. p. tublcola Kiefter, 1889-1, p. 188 {Cectdomyia);
1913-2, p. 82.
On Cylisus scoparius. Swanton, 559 ; Houard, 3423.
Northumberland and Durham, several records. Cum-
berland, Keswick. Lancashire, Freshfield.
*213. P. nUcis Kieffer, 1909, p. 31 (gall); 1913-2, p. 82.
On Vlex, = Cecidomyid sp. Houard, 3396.
Northumberland, between Alnmouth and Warkworth.
Durham, Gibaide, Waldridge (gall only). LANCASHrRE,
Grange-over-Sands (signs). Yorkshire, Bston (old galls).
214. P. ulnuriae Bremi, 1847, p. 52 (Cecidomyia) ; Kieffer,
1913-2, p. 82.
On Spiram JJlmaria. Swanton, 475 ; Houard, 2839.
Common ; records from all counties.
215. Perrisia sp.
On Spiraea Filipendula. Perrisna ulmaria of Swanton,
476 and Houard, 2830. Swanton (1912, p. 64) suggest*
the name P. jiUpenduUte for this gall, but that name has
already been used by Kieffer (1909, p. 29) for another
species affecting the same plant.
•216. P. ulmicola Kieffer, 1909. p. 31 (gall and larva);
1913-2, p. 82.
On Ulmus.
Durham, Bummoor, Gibside, Norton. Northumber-
land, Warkworth, rare. YoRKf^HiRE, Bardsey, near Leeds.
217. P. urtteae Perris, 1840, p. 403 (Ceddomyia); 1913-2,
p. 82.
On Urticadioicn and (more rarely) on U.vrenti. Swantcn,
312, 316 ; Houard, 2095, 2099.
Very common everywhere.
n,g,t,7.dh,G00glc
384 Messrs. R. S. Bagnall and J. W. H. Hamaon's
•218. P. noeinlvonun Kieffer (nom. nov. for nux^nii
Rubs, turn Smith); Riibsaamen, 1895, p. 258
{Dichelomi/ia); Kieffer, 1913-2, p. 82.
On Tacdnium Myrtillua. Houard, 45W.
DcRHAM, rare, Birtley, Waldridge.
219. P. veronicM {chamaedrys Inchb.) Vallot, 1827, p. 93
(Cecidomyia), Kieffer, 1913-2, p. 83.
On Veronica Chainaedrys. Swanton, 728; Houard, 5080.
Recorded also in Houard from the British Isles on *V.
officinale (ullSo) and "F. serpyUifolia {5105).
Yen- common; records from all counties. Taken by
J. W. H. H. at Ninebanks in Northumberland on V.
monttwa, and by R. S. B. at Warkworth on V. officinale.
220. P. vteiM Kieffer, 1888-1. p. 105 (Cecidomyia);
1913-2, p. 83.
On Vicia spp. Swanton, 590, 594 ; Houard, 3731, 3696,
*3723.
General ; records from all counties excepting Westmore-
land.
221. P. violae F. Loew, 1880. p. 34 (Cecidmnyia): Kieffer.
1913-2. p. 83.
On Viola an-ensis. Swanton. 640; Houard, 4293.
Dl'rha.m, Gibside, rare. Cumbeblakd, near Nenthead,
rare.
•222. P. virgae-aureae Liebet. 1889, p. 283 {Cecidomyia);
Kieffer, 1913-2. p. 83.
On SoUdaffo Viryaiirea. Houard, 5560, 5564.
N0RTH1.-MBERLASD, near Staward.
*223. Perrisia sp. Kieffer, 1898.
On Solidago Virgniirea. Houard, 5559.
Records from all counties excepting Cumberland.
*224. P. vitis-ldaeae Kieffer, 1909, p. 31 (gall); 1913-2,
p. 83.
On Vaccinliim VUt.s-idaea. Houard, 4570.
Cumberland, summit of Skiddaw.
iM,Googlc
Preliminary Catalogue of British Cecidomyidae. 385
*225. PertisUsp. Bagnall and Harrieon, 1917 (v), p. U.
On Myosotis arvensis; flower closed and slightly swollen,
containing larva.
Lancashire, Grange-over-Sands. Kieffer has described
two species of Perrieia from the flower of Myosotis paluslris.
HartigtoU Riibsaamen, 1912, p. 161.
226. H. annuUpes Hartig (jyUiger H. Loew), 1844, p. 162
{Cecidomyia); Kiefter, 1913-2, p. 85.
Gall on beech-leaves. Swanton, 292; Houard, 1153.
Records from all counties excepting Westmoreland.
Taxomyla Riibsaamen, 1912, p. 216.
227. T. taxi Inchbald, 1861, p. 76 {Ceddomyia); Kieffer,
1913-2, p. 86.
On Taxus baccata. Swatiton, 14; Houard, 150.
Northumberland, Felton. Durham, Gibaide, Fatfield,
Birtley. Lancashire, Grange-over-feinde, Silverdale.
Yorkshire, Gunnergate.
Group A.HPHONDYLARIAE.
Sehiiomyla KiefEer, 1889-1, p. 183.
228. 5. KaUorum Kieffer, 1889, p. 184; 1913-2, p, 89.
On Galium spp. Swanton, 746, 756; Houard, 5281,
5248.
Records from all counties excepting Westmoreland and
Cheshire.
*228a. S. Ugustrl Riibsaamen 1899-1, p. 598; Kiefter,
1913-2, p. 89.
In flowers of privet. Houard, 4679.
Durham, Hemlington; Hesleden Dene. Also taken at
Bath.
KleOeria Mik, 1895, p. 96.
229. K. plm^ellae F. Loew (umbellataTuin F. Loew)
1874, p. 157; Kieffer, 1913-2, p. 90.
On UmbeUiferae spp. Swanton, 657, 666, 670 ; Houard,
4445, 4499, 4529.
Dubhau, abundant on a plant of * Angelica sylvestris^
iM-,i,A J h, Google
386 Messrs. R. S. Bagnall and J. W. H. Harrison's
Lamesley; once only on PimpineUa saxifraga, Penshaw.
On Daucus carota, Hart (J. Gardner), This species is not
recorded in Houard on Angelica.
Asphondylla H. Loew, 1850, p. 21.
230. A. dorycnii F. Loew, 1880, p. 37; Kieffer, 1913-2,
p. 93.
On Lhrifcnium.
231. A. genistae H. Loew, 1850, p. 38; Kieffer, 1913-2,
p. 93.
On Genista (germanica, Houard, 3345).
*232. A. Inpultnae Kieffer, 1909, p. 14 (gall and nymph);
1913-2, p. 93.
On Medicago lupulina, = Aspfumdylia ap. Houard, 3506.
Northumberland, Warkworth, Durham, Birtley ; near
Penshaw. Cumberland, near Alston.
233. A. mayari Liebel, 1889, p. 266; Kieffer, 1913, p. 93.
On Cylisus scoparius. Swanton, 558; Houard, 3412.
Northumberland and Durham, many records. Lanca-
shire, Freshfield.
*234. ?A. mikli Wachtl, 1880, p. 535; Kieffer, 1913-2,
p. 94.
" Medicago hipulina. Perrisia sp. Seeds swollen and
enlarged, each containing a yellow midge larva."— Burkill,
1916, p. 5.
Yorkshire, Burkill's record.
*235. A. melanopus Kieffer, 1890-1, p. 31; 1913-2, p. 93.
Seed-pods of Lotus corniculatxis, rare. Houard, 3613.
Northumberland, Warkworth. Durham, Birtley,
Tinkler Fell, Penshaw Hill. Lancashire, near Freshfield.
•236. A. ononidls F. Loew, 1873, p. 139; 1913-2, p. 94.
On Ononis repens. Houard, 3501.
Northumberland. Warkworth. Durham, on the coast
etween Horden and Hart.
l„y,|,AJI^,COOl^lC
Preliminanf Catalogue of British Ceddomyidae. 387
*237. A. prozima Kiefter, 1909, p. 30 (larva and gall);
1913-2, p. 94.
On Thymxis Serpyllum, = Cecidomyid sp. Houard, 4914,
Durham, Stanhope. Cumberland. Alston. Lanca-
shire and Westmoreland, Hampsfell and Meathop Fell,
near Grange-over-Sanda.
238. A. urothsmni H. Loew, 1850, p. 38; Kieffer, 1913-2,
p. 94.
On Cylisus scoparitis. Swanton, 556, 557 ; Houard,
3414, 3422.
Records from all counties excepting Westmoreland.
239. A. thyml Kieffer, 1898-2, p. 59; 1913-2, p. 94.
On Thymus Serpyllum. Swanton, 699; Houard, 4913.
Northumberland, Warkworth.
240. A. uUcis VerraU, 1875, p. 225 (gall); Kieffer, 1913-2,
p. 94.
On Vkx europaeus. Swanton, 552 ; Houard, 3395, 3397.
Conunon; records from all counties.
Group Brachyneubahiae.
Phaenotautbla Kieffer, 1912-2, p. 2.
♦241. P. cardui Kieffer, 1904-2, p. 346; 1913-2, p. 111.
In galla of Trypela cardui.
Durham, Penshaw and Edraondsley.
BUUoIa Kieffer, 1896-3, p. 5.
242. H. tagl {Tipuh rubra Hermann) Hartig, 1839, p. 4;
Kieffer, 1913-2, p. 105.
Gall on leaves of Fagus. Swanton, 291 ; Houard, 1151,
Northumberland and Durham, several records. Cum-
berland, Alston. Yorkshire, Marton.
Lasiopteryx Stephens.t 1829, p. 240.
243. L. obfuscaia Meigen, 1818, p. 90; Kieffer, 1913-2,
p. 117.
t Doubtful genus.
n,g,t,7.dh,G00glc
388 Messrs. R. S. Bagnall and J. W. H. Harrison's
Braohyneura Rondani, 1840, p. 18 [type fuaco-grisea
Rondani).
244, B. slygia Walker {nee Meigen).t
Group Cecidomyiariae (Diplosariae).
Hormomyia H. Loew, 1850, pp. 20 and 31.
246. H.fischeri Frauenfeld, 1867, p. 781 ; Kieffer, 1913-2,
p. 138.
On Garex limosa. Swanton, 72.
•246. H. fritenl Kieffer, 1909, p. 5; 1913-2, p. 138.
Durham, Gibside, on Carex binervis; Birtlev Fell, on
C.fiava.
247. ff . yrffn(/i«, Meigen, 1804,p. 39 (Cecid<miyia); Kieffer,
1913-2, p. 138.
*248. H. kneuokeri Kieffer, 1909, p. 5; 1913-2, p. 138.
On Carex sleUulata. — Hormomyia sp, Houard, 368.
Durham, Waldridge.
Dishormomyfa Kieffer. 1912-2, p. 2.
•249. D. oornitex Kieffer, 1898-2. p. 60 (Hortnomyia);
1913-2, p. 139.
On Carex slricta and C. fiava. Houard. 373.
Durham, near Chester-le-Street. Che.shire, BidstOD
Hill.
Trishormomyla Kieffer, 1912-2, p. 2.
•250. T. tuberlflca Riibsaamen, 1899, p. 603 (Hormomyia);
Kieffer. 1913-2, p. 140.
On Carejr spp. Houard, 376.
Cheshire. Bidston Hill, not rare. ? Durham. Gibside,
on C. bineri-is, and coast between Horden and Hart, on
C. glaucn. Northumberland, Warkworth, on C. glauca.
t See Collin, 1904, note (3).
u,y,l,7rJM,G00glc
Prdiminary CaifUoffue of British Ceddomyidae. 389
Cyrtodiplosis Kiefler, 1912-2, p. 1.
*251. C. erassiurva Kieffer, 1901-2. p. 172 (Clitwdiplosis) ;
1913-2, p. 144.
On Slachys sylvatica. Houard, 4859.
Durham, Birtley, Lamesley, LambtoD and Fatfield.
HoDartliropalpiis Rubsaameu, 1892, p. 381.
252. M. buxi Geoffrey, 1764, p. 545 {Scatiypse); Kieffer,
1913-2, p. 150.
Yorkshire, Nunthorpe.
Dlehrona Riibsaamen, 1899, p. 542.
•253. D, gallanim Rubsaamen, 1899, p. 542; Kiefier,
1913-2, p. 151.
On Carex app.
NoRTHUMBBRiAND. Bamburgh, on C. dxstana. Durham,
Waldridge, on C. fiava. Cheshire, Bidston Hill, several,
on C atricUi.
Arthrocnodsx Riibsaamen, 1895-2, p. 189.
254. A.fraxindla Meade, 1888, p. 77 (Diplosis); Kieffer.
1913-3, p. 156.
In galla of the Eriophyid, E. fraxini, on Ash.
Phaenobremla Kieffer, I9I2-2, p. I.
*255. Phaenobremia sp. Bagnall and Harrison, 1917-2, p.
208.
Durham, West Cornforth, larvae feeding on Aphis mali
on apple.
Aphidoletes Kieffer, 1904-2, p. 385.
•256, A. abletls Kieffer, 1896-2, p. 382 (Bremia); 1913-2,
p. 164.
Durham, in galla of Adelges abielis, Gibside.
*257. Aphidoletes sp. Bagnall and Harrison, 1917-3, p. 229.
Durham, in galls of Adeiges strobtlobius, Gibside; larva
entirely red.
iM-,i,A J h, Google
390 Messrs. R. S. Bagnall and J. W. H. Harrison's
Anabramla KieiTer, 1912-2, p. I.
*258. A. DelkToyel Kieffer, 1896-2, p. 384; 1913-2, p. 169.
On Latkyrus pratensis. Clinodiplosis bdlevoyei, Houard,
3774.
Records from all counties,
*259. A. vIclM KieSer (nom. nov.) {Clitutdiphsis longiven-
tris larva, but- not imago), 1909, p. 32; 1913-2,
p. 169.
In flowers of Vida sepium. C. longivetUris, Houard,
3694.
Durham, Fatfield, taken by Mr. W. Hall.
Hadrobremia Kieffer, 1912-2, p. L
*260. H. loaglvenlris Kieffer (imago, not larva ; trifolii
Kieffer larva, I.e. p. 31), 1909,p.34(CIt"norfipfewM);
1913-2, p. 170.
On Trifdium pratense. C. Irifolii Kieffer, Houard
(supplement), 0922.
DoKHAM, Birtley.
Endaphls KieSer, 1896-2, p. 383.
*261. E. perfidus Kieffer, 1896-2, p. 383; 1913-2, p. 172.
Endoparasite of Aphis plaianoides.
Northumberland, Warkworth.
•262. Endaphis sp. Bagnall and Harrison, 1917-2, p. 208.
Endoparasite of an Aphis on Ononis.
Durham, Penshaw Hill.
Thuraula Rubsaamen, 1899-2, p. 38.
•263. Thuraula sp. Bagnall and Harrison, 1917-2, p. 208.
Durham, Waldridge Fell, in leaf-sheaths of Carex
Goodenovii, submerged.
Syndiplosis Rubsaamen, 1910, p. 425.
•264. S. iDDieerearum F. Loew, 1877, p. 17 {Dipiosis);
Kieffer, 1913-2, p. 178.
On Viburnum Opuhts. Cant. Umicereamm, Houard, 5338.
Northumberland, Ovingham. Durham, Fatfield, Gib-
aide, Waldridge Fell.
l„y,l,Ajh,COOl^lL'
PreHminary Catalogue of British Cecidomyidae. 391
265. 5. pe/ioi* Kieffer, 1898-2, p. 37 (ffarmandia) ; 1913-2,
p. 178.
On Popidus Iremula. H. petioli, Swanton, 173, 174;
Houard, 493, 497.
Contarlnia Rondani, 1860, p. 287.
♦266. C. awtosu Kieffer, 1901-2, p. 31 ; 1913-2, p. 179.
In flowers of Rumex Acetosa and AcdoseUa. Houard,
2128, 2136.
Northumberland, Watkworth. Durham, Greenside
and near Lanchester; Beamish. Cheshire, Bidston Hill.
♦267. C. anthobU F. Loew, 1877, p. 16; Kieffer, 1913-2,
p. 179.
In flowers of Crataeffus OxyacarUha. Houard, 2941.
Northumberland and Durham, several records. York-
shire, Gunnergate.
•268. C. barbiehei Kieffer, 1890-1, p. 29; 1913-2, p. 179.
(a) On Lotus comicukUtts. Houard, 3617,
Not rare ; records from all counties.
(b) On Lotus major. Houard, 3627.
Durham, Gibside and Tinkler Fell.
*269. C. totaUcola Kieffer, 1889-1, pp. 155 and 171;
1913-2, p. 179.
On Betula (dba. Houard, 1069.
Northumberland and Durham, several records. Lan-
cashire, Freshfield, Grange-over-Sands. Cheshire, Bid-
ston Hill. Yorkshire, Elston.
270. C. betuUna Kieffer, 1889-1, p. 153; 1913-2, p. 179.
On Belubt alba. Swanton, 192; Houard, 1076.
Northumberland and Durham, several records. Lan-
cashire, sandMUs near Freshfield, plentiful. Yorkshire,
Eiston; Leeds.
271. C. eammnulae Kieffer, 1895-3, p. 9; 1913-2, p. 180.
On Campanula rotuiidiftdia. Swanton, 795; Houard,
KU,
NoBTHUMBBRLANT), Warkworth.
n,g,t,7.dh,GoOglc
392 Messrs. R. S. Bagnall and J. W. H. Harrison's
•272. C. coryU KiefEer, 1909, p. 31 ; 1913-2, p. 180.
On Corylus Avdlnna. = Diplosine s^t.. Honard, 1062-
North ITMBEBLAND and Durham, several records. Lak-
CASHiRE, Grange-over- Sands, rather rare. Westmoreland,
Kirkby-Stephen. Yorkshire, Bardsey, near Leeds.
273. C. craccae Kieffer, 1897-3, p. 15; 1913-2, p. 180.
On Fw-w spp. Swanton, 589, 593 ; Houard, 3721, 3693.
Not uncommon ; records from all counties excepting
Westmoreland.
*274. C. cucuball Kieffer, 1909, p. 28; 1913-2, p. 28.
On Silene inflala. =■ CotUarinia sp., Houard, 2266.
Durham, Birtley, Fatfield and Hytton.
*275. C. gei Kieffer, 1909, p. 9; 191 J-2, p. 180.
On Geum rirah. = Diplostne sp., Houard, 3091.
Cumberland, near Nenthead.
276. C. bellantheml Hardy, 1850, p. 187; Kieffer, 1913-2,
p. 180.
On Helirmthemiiin rulgare. Swanton, 631 ; Houard,
4269.
Northumberland, Cheviot district (Hardy). Durham,
Byhope, Easington, Horden and Blackhall R^cks,
Lanl'a.shire and Westmoreland, Silverdale, Lindale,
and U range -over- San da neighbourhood.
277. C. taeraclel Kiibsaanien, 1889-1, p. 274; Kieffer,
1913-2, p. 180.
On Herachum Sphondylitnn. Swanton, 668; Houard,
451.1.
Nobthu.mberland, Langley W'oods. Durham, Birtley,
; Hylton and neat Washington, rare. CuM-
I, Nenthead.
•278. C. lalhyrl Kieffer. 1909, p. 13; 1913-2, p. 181.
In flowprof lAilhyrxs praleti.iis. = CorUarinia sp., Houard,
3770.
Durham, Penshaw, a few; Lamesley, one only.
iM-,i,A J h, Google
Preliminary Catalogue of British Cecidomyidae. 393
279. C. HnwiM Winnertz, 1853, p. 260; Kiefter, 1913-2,
p. 181.
On Linaria vulgaris. Swantoii, 724 ; Houard, 5028.
Durham, Biddick and Fatfield (B. S. B. and W. Hall),
Birtley. Lancashire, near Freshfield.
280. C. loU De Geer, 1776, p. 420; Kieffer, 1913-2, p. 181.
(a) On Lolus comiculatus. Swant<in,583; Houard,"3614.
Northumberland, Ninebanks. Durham, several records. '
Cumberland, Alston. Lanca.shibe, Birkdale, Ainsdale,
Freshfield, in numbers.
*{b) On Lotus major. Houard, 3625.
Lancashire, Ainsdale, rare. Yorkshire, see Burkill,
1916. Mr. Burkill also records this from Surrey and
Staffordshire on L. major.
•281. ?C. medicaginis Kieffer, 1895^, p. 150; 1913-2,
p. 181.
On Medicago falcata. Swanton, 562.
The species recorded by Connold and Swanton (562),
" Flowers and seed-pod swollen, reddish. Tjar\'ae gre-
garious," would seem to refer to this species and (in the
aeed-pod) Asphondylia mikit. Certainly not to C. loti.
282. C. melanoeera Kieffer, 1909, p. 76; 1913-2, p. 181.
On Genista iincloria. Swanton, 550; Houard, 3372.
Durham, Gibside, plentiful.
•283. C. nastmtii Kieffer, 1888-2, p. 263; 1913-2, p. 181.
See Carpenter, 1911, pp. 68-71, on Brassica Rapa.
Houard, 6701 (supplement).
•284. C. nieolayi Bubsaamen, 189.5-1, p. 183; Kieffer,
1913-2, p. 181.
On Heracleum Sphondiflium. Houard, 4509.
Durham, many records. Cumberland, Alston. Lan-
cashire, Grange and Liiidale. Yorkshire, Redcar.
•285. C. ononidis Kieffer, 1889-3. p. 93; 1913-2, p. 181.
On Ononis repens. Houard, 3.500.
. Northumberland, Warkworth. Durham, between
Horden and Hart; also Kyhope.
l„y,l,Ajh,COOl^lC
394 Messrs. R. S. Bagnall and J. W. H. Harrison's
286^ C. pyri?ora Riley, 1886, p. 283 {Diplosis); Kiefier,
1913-2, p. 181.
On Pyrus communis. Swanton, 525 ; Houard, 2855,
Durham, Wear and Derwent valleys, a few old records.
287. C. quercina Riibsaamen (dryophila Kiefier), 1890,
p. 21 ; Kiefier, 1913-2, p. 181.
On Quercus. Swanton, 278; Houard, 1207.
Durham, Gibaide, Fatfield and West Comforth. York-
shire, Bardaey, near Leeds.
*288. C. rubloola (Kiefier, 1908, gall and larva) Rub-
saamen, 1910, p. 424; Kieffer, 1913-2, p. 181.
In flowers of Rulms caesitis. Houard (supplement), 6775.
Northumberland, Warkworth. Durham, Birtley, Bid-
dick. Lancashire and Westmoreland, near Grange-
over- Sands.
289. C. ruderalis Kieffer, 1890-3, p. 198; 1913-2, p. 181.
On Sisymbrium officinale. Swanton, 403 ; Houard, 2576,
Durham, Birtley. Yorkshire, Redcar.
*290. C. seaUosae Kieffer, 1898-2, p. 60; 1913-2, p. 182.
On Scafnosa Columbaria. Houard, 5463.
Durham, near Birtley ; PenahawHill; TunBtal Hill, near
Sunderland ; Horden and Hart. Cumberland, near Alston.
•291. C. sooparil Rubsaaraen, 1889-2, p. 48; Kieffer,
1913-2, p. 182.
On Cy(isMS scoparius. Houard, 3421 and 3430.
Durham, Horsleyhope and between Newbiggin and
Rowley ; Wolsingham.
•292. C. sorbi Kieffer, 1896-1, p, 99; 1913-2, p. 182.
On Pyrus Aueuparia. Houard. 2909.
Durham, Westgate. Cumberland, near Alston,
Cheshire, Bidston. Yorkshire, Great Ayton Moor,
293. C. steini Karach, 1881, p. 227; Kieffer, 1913-2,
p. 182.
On Lychnis alba and *L. dioica. Swanton, 345 ; Houard,
2291 and '2294.
Records from all counties excepting Westmoreland.
ih,Cooi^lc
Prelimnary Catahffue of British Ceddomyiiae. 395
294. C. tUiarum Kiefler, 1890-2, p. 193; 1913-2, p. 182.
On Tiha platypkyllos and T. vulgaris. Swanton, 615-17,
619; Houard, 4122-23-25-54.
Records from all counties excepting Westmoreland and
Cheshire.
♦295. C. tngopogonis Kieffer, 1909, p. 30; 1913-2, p. 182. '
On Tragopogonpratensis.^CorUariniaRp., Houard, 6077.
DaRHAM, Wear Valley, from Chestet-le-Street to Hylton;
Byhope, Lancashire, Birkdale. Cheshire, Bidston.
Yorkshire, Redcar.
296. C. tralU Kieffer, 1889-2, p. 262; 1913-2, p. 182.
Flowers of Pimjnnella Saxifraga. Swanton, 658 ; Houard,
4446.
Durham, near Penshaw, galls only, rare.
297. C. iTemuhe Kieffer, 1909, p. 78; 1913-2, p. 182.
On Popttlus tremula. =^ Contarinia sp., Swanton, 172;
Houard, 502.
298. C. trUid, Kirby, 1797, p. 246 {Cecidomyia) ; Kieffer,
1913-2, p. 182.
*299. C. Valerianae Riibsaamen, 1890, p. 231 ; Kieffer,
1913-2, p. 182.
Valeriana sambucifiAia. Houard, 5421.
Durham, Gibside.
*300. C. ?ibamorum Kieffer (nom. nov. for C. vibumi
Kieff. non Pelt), 1912-1, p. 230; 1913-2, p. 182.
In flowers of Viburnum Opulua.
Northumberland, Ovingham. Durham, many records.
Cumberland, Keswick.
*301. Contarlnia sp. Riibsaamen; Lagerheim.
On Oalium ventm. Houard, 5288.
DuRHA.M, Penshaw Hill. Northumberland, Hamburgh,
Warkworth. Also from Scotland.
•302. Contarfnia sp. Bagnall and Harrison, 1917-3, p. 229.
On Bupleurum tenuissimum.
Durham, Greatham.
n,g,t,7.dh,G00glc
396 Means. R. S. BagnaD and J. W. H. Harrison's
Stielodiploais Kieffer. 1891, p. 28.
303. S. ewrSm P- Loew, 1878. p. 366; Kieffer. 1913-2,
p. 183.
On Coryliu Af^lana. Swanton, 215; Houard, 1053.
La>'cashibe. Grange-over- Sands, not uncommon. Dur-
ham, wood near Fatfield. Northumberland, Warkworth
and Ovingham. Yorkshire, Bardsey, near Leeds.
*3M. S. hjpoeboeridb Rubsaamen, 1891, p. 52; Kieffer.
1913-2, p. 183.
(a) On Hypochoeris radicata. Houard, 6034.
Records from all counties.
(6) On Crepis biennis. Houard (supplement). 7538.
Yorkshire, Stainton.
305. S. )aeobaea« H. Loew, 1850, p. 29; Kieffer, 1913-2,
p. 183.
On Senecio Jacobaea and *S. erucifolifU. Swanton, 841 ;
Houard, 5865.
Records from all counties excepting Cumberland, bui
only once on 5. emcifolius (Cowpen-Bewley, co. Durham).
*306. S. pUosellae Kieffer, 1896-1, p. 100; 1913-2, p. 183.
On Hieracium Pilosella. Houard. 6197.
Durham, Fatfield, and Tunstal Hill, near Sunderland,
rare.
307. S.sorophul»riae Kieffer, 1896-1, p. 100; 1913-2, p. 183.
On ScTophularia nodosa. Swanton, 726; Houard, 5063.
Northumberland, near Staward. Durham, Oibside,
locally plentiful ; Fencehouaes. Lanca.shtre, Grange-over-
Sands, rare.
♦308. 5. umbellatarum Riibaaamen, 1910, p. 422; Kieffer,
1913-2, p. 183.
Flowers of Pimpinella magna.
Lancashire, near Grange -over* Sands.
Tbecodlplosis Kieffer, 1895-2, p. 194.
309. T. braehynlera Schwagrichen, 1835, p. 162 {Cecido-
myin]; Kieffer, 1913-2, p. 184.
On Pinus sylreslris. Swanton, 21 ; Houard, 76.
iM-,i,A J h, Google
Prdiminaty Catalogue of British Cectdomtfidae. 397
Northumberland, neat Warkworth. DtmHAM, Tinkler
Fell. Lancashire, Oiange-over-Sands.
Hyrleomyia Kieffer, 1900, p. 470.
•310. H. nuditerranes F. Loew, 1885-2, p. 485 (DtpUms) ;
Kieffer, 1913-2, p. 186.
On Erica Telralix.
Northumberland, Blanchland. Durham, Waldridge,
Birtley, Kilihope and Waakerley. CuMBBBLAND, Killhope.
Yorkshire, Eaton and Ayton.
Zeuiddiplosis Kieffer, 1904-2, p. 349.
*311. Z. KUrdl Kieffer, 1896-2, p. 383 {Theeodiplosis
giardiana Kieffer, 1898) ; 1913-2, p. 187.
On (a) Hypericum perforatum and (ft) H. pulcknim. = Z.
giardiana, Houard, 4"210, 4202.
NoRTHUMBEBiwiND, near Staward (fc), rare. Durham,
Gibside (a), rare. Cumberland, near Nenthead (b), rare.
Atrtehosenu Kieffer, 1904-2, p. 347.
312. A. aceris Kieffer, 1904-2, p. 348; 1913-2, p. 189.
On Acer campeslris. Swanton, 606, 607 ; Houard, 4030.
Trlehodiplosis Kieffer, 1912-1, p. 229.
*313. T. oartsts Kieffer, 1898-2, p. 61 (Antichira); 1913-2,
p. 191.
In leaf-sheatha of Carex Goodenovii and C, glnuca.
Durham, Waldridge. Northumberland, Warkworth.
Cumberland, Alston. Cheshire, Bidaton,
Antichirldlum Rubsaamen, 1911, p. 168 (for Antichira
Rubs. 1911, won Eschscholz).
*314. A. strUtiiin Riibsaamen. 1911, p. 122 (Antichira);
Kieffer, 1913-2, p. 192.
In leaf-sheaths of grasses and sedges if large and coarse.
Northumberland, Warkworth, on Varex jtenduh.
Durham, Birtley, on Typha UtfifoUu; Oibside, on Carex
pendula; and near Swalwell, on Phrngmites. Lanca.shire,
near Grange-over-Sanda, on Pkragmiles.
TRANS. ENT. 80C. LOND. 1917. — PARTS II, III, IV. (mAY '18) DD
u,y,i,Ajh,Cooglc
398 Meflsra. R. S. Bagnall and J. W. H. Harrison'B
Lestodiplosis KieSer, 1894, p. 28.
315. i. callida Winnertz, 1853, p. 255 {Diphsis); Kieffer,
1913-2, p. 197.
316. L. cerUralis Winnertz, 1853, p. 277 {Diphsis) ;
Kieffer, 1913-2, p. 197.
Pseudhormomyia Kieffer, 1898-2, p. 57.
317. P. granHex Kieffer, 1898-2, p, 57-8; 1913-2, p. 204.
On Carex aiespitosa, *C. paniculaia, *C. slrttta and
C. flava. Swanton. 71 ; Houard, '374.
Cheshire, Bidston Hill, on C. slricta. Durham, Birtley,
on C.jiava.
Taphodlplosis Kieffer, 1912-2, p. 2.
•318. T. lubterranea Kieffer el Trotter, 1905, p. 65 (Pseud-
hormontyia); Kieffer, 1913-2, p. 205.
On Carex spp. Houard, 366.
Northumberland, Hamburgh, on C. distans. Durham,
near Lanchester, on C. divuUa ; Waldridge, on C. ftava ;
Gibside, on C. binervis. Lancashire, Grange-over-Sands.
Dyodlplosis Riibsaamen, 1912, p. 49.
•319. D. arenariae Rubsaamen, 1899, p. 602 {Hormmnyia).
On Carex arenarm.
North u.mberl AND, Bamburgh, rare.
Loewiola Kieffer, 1896-3, p. 5.
320. L. centaureae F. Loew, 1870, p. 25 (Diplosis) ; Kieffer,
1913-2, p. 207.
(a) On Centaurea nigra, very local. Swanton, 855, 856,
Durham, Fatfield and Ryhope. Lancashire, Lathom.
•(&) On C. Scabiosa. Houard, 599(».
Durham, Ryhope.
Recorded by Mr. Burkill from Surrey, on C. Scabiosa.
Putoniell* Kieffer, 1896, p. 4.
321. P. marsupialls F. Loew, 1889, p. 536; Kieffet,
1913-2, p. 209.
On Primus spinosa. Swanton, 456; Houard, 3295.
Durham, Ryhope Dene, plentiful.
u,y,l,7rJM,G00glc
Preliminary Cataloffue of Brititk Ceeidomyidae. 399
RaidodlplMis Rubsaamen, 1911, p. 393.
•322. H. oquestris B. Wagner, 1871, p. 41; Kieffer, 1913-2,
p. 210.
See Houard, 333.
Lancashire, Grange-over-Sands, on a common grass —
Triticum sp. or ally. Brought forward aa British by the
late F. Enock.
CeeMomyU *Meigen, 1803, p. 261.t
323. C. pini de Geer,* 1776, p. 417 {Tipula); Kieffer,
1913-2, p. 215.
324. G. flava Meigen, 1818, p. 99 {DipUms Theobald) ;
Kieffer, 1913-2, p. 215.
325. a «mia Curtis, 1827, p. 178; Kieffer, 1913-2, p. 220.
HMTOdlplotis Kieffer, 1895-2, p. 194.
326. M. dryoUa F. Loew, 1877, p. 14; Kieffer, 1913-2,
p. 223.
On Quercus. Swanton, 279; Houard, 1306.
NoRTUUUBERLAND and DuRHAM, Several records. Cum-
BERLAm), Keswick. Lancashire, Grange. Yorkshire,
Bardaey near Leeds and Nuntborpe.
327. H. volvens Kieffer, 1904-1, p. 79; 1913-2, p. 223.
On Quercua. Swanton, 280; Houard, 1307.
Northumberland and Durham, several records. Cum-
berland, Keswick. Lancashire, Grange-over-Sands, rare.
Xenodi^ods Felt, 1911, p. 61 {= AUodiphsis Biibs. non
Kieffer).
*328. X.Ueviu8euliRubsaamen,1911,p.85; Kieffer, 1913-2,
p. 228.
On gall of Neuroterus laevinsculus.
Durham, Fatfield. Northumberland, Warkworth.
Yorkshire, Bardsey near Leeds.
t Ajlapeciee in this genus excepting the type pini are insufflciently
deecrib«d, and therefore cannot be relegated to any known genuB.
X Synonyms ftr« lalereHa Zett,, pihaa Btemi, and pini-maritimae
Dup.
u,y,i,Ajh,Cooglc
400 Meesrs. R. S. Bagnall and J. W. H. Harriflon's
PsnIModiplosis Riibsaamen, 1910, p. 287.
•329. P. gaUiperda F. Loew, 1889, p. 202 {Diplom);
Kieffer, 1913-2, p. 229.
On gall of Neurotems lefUicuIaris on Oak.
Northumberland, Warkworth and Ovingham. York-
shire, Nunthorpe.
Rarmandia Kieffer, 1896-3, p. 5.
330. H. tnmulae Winnertz, 1853, p. 263 ; Kieffer, 1913-2,
p. 234.
On Populm tremi/la. Swanton, 175; Honard, 506.
Durham, Low FeU.
•331. H. pustulani Kieffer, 1909, p. 18 (gall), and 1912-1,
p. 230 (larva); 19ia-2, p. 234.
On Pojmliis tremula. = Diplosine sp., Houaid, 513.
Yorkshire, Nunthorpe, very rare.
Dichodlplosls Riibsaamen, 1911, p. 171.
•332. D. lanKeni RubBaamen, 1911, p. 171 ; Kieffer, 1913-3.
p. 235.
On dried plums.
Durham, Gibside.
CUnoiUplosIs Kieffer, 1894-2, p. 121.
•333. C. betonicae Kieffer, 1909, p. 3 ; Kieffer, 1913-2, p- ^^'^'
In flower of Betonica officinalis.
Northumberland, Ninebanks; Warkworth. DuJtP*
Gibside.
334. C. bolulaiia Winnertz, 1853, p. 266 (DipU^^'
Kieffer, 1913-2, p. 237.
Inquiline in galls of Perrisia fraxini. ' ,
Northumberland, near Staward. Durham, PeO»**fL
Hill Cumberland, Alston. Yorkshire, Bardsey **
Leeds and Gu uncinate .
•335. c. rosiperda Rubaaaraen, 1892-2 p 54- Ki«*^^'
1913-2, p. 238,
On Rosa vilhrn (abroad on Rosa centifolia ool/r ^
Houard, R2, 3128).
n,g,t,7.dh,G00glc
Prdiminary Catalogue of British Ceddomyidae. 401
Durham, Billingbam, Latnesley and Gibside. North-
UUBBRLAND, Ovingham. Yorkshire, Bardsey near Leeds.
*336. C. seUechtendftU RUbsaamen, 1911, p. 16; Kieffer,
1913-2, p. 238.
On Convolwlus septum.
Northumberland, Warkworth. Durham, between
Wolviston and Greathara.
Hyeodiploris Rilbsaamen, 1895-1, p. 186.
336a. M. coniophaga Winnertz, 1853, p. 267; Kieffer,
1913-2, p. 242.
*337. Hyeodiplods sp. Bagnall and Harrison, 1917-2, p. 208.
On the aecidia of Ur&myces jund, larva orange-red.
Durham, Billingbam.
•338. Mycodlplosissp. Bagnall and Harrison, 1917-2, p. 208.
On Crepia paludoaa, larvae otange-pink, feeding on
Puannia major.
Durham, Waldridge Fell.
*339. HyeodiplMb sp. Bagnall and Harrison, 1917 (v),
p. 14.
On Cirsium arvense, larvae crimson, feeding on Puccinia
suavoletts.
Durham, Fatfield, Fulwell near Sunderland, Fenshaw.
NORTHUMBBRLAND, WarkwOfth.
*340. Hyeodlplo^ sp. Bagnall and Harrison, 1917-3,
p. 229.
Larvae yellowish-oiange on Puccinia hieracii on H. boreale.
Durham, Winlaton Mill.
Hassalongla KieETer, 1897-3, p. 12.
341. H. rubra Kiefler, 1890-3, p. 199; 1913-2, p. 246.
On Bdula alba. Swanton, 193 ; Houard, 1075.
Northumberland, four miles north of Morpeth ;
l„y,l,Ajh,COOl^lC
402 Messrs. R. S. Bagnall and J. W. H. Haniaon's
Ninebanks. Durham, Gibside, Waldridge, Tinkler Fell,
Urpeth, Castle Eden dene. Cheshire, Bidstoo Hill, local,
Yorkshire, E^ton Moor ; Leeds.
Ametrodlplosis Riibsasmen, 1911, p. 278.
342. A. thalictricola Riibaaamen, 1895-1, p. 257; Kieffer,
1913-2, p. 246.
On Tkaliclrum minus and jlatmm. Clinadtj^osis thalic-
tricola. Swanton, 356, 358; Houard, 2441, 2448; and on
* T. flexuosum, Houard, 2451.
Honodiplosls Riibsaamen, 1910, p. 289.
*343. H. llebell KiefEer (Schizomyia sociabUis Riibs.),
1889 (-1), p. 174; 1913-2, p, 248.
Living in galls of Macrolabis dryobia and volvens.
Northumberland, Ovingham, with M. voh-ena. Dur-
ham, Gibside, Fattield, with M. dryobia.
Atylodjplosis Rubsaamen, 1910, p. 338.
*344. A. rumjcii H. Loew, 1850, p. 190; Kieffer, 1913-2,
p. 257.
In flowers of Rmnex AceloseUa. Houard, 2128.
General and sometimes common ; records from all
counties excepting Westmoreland.
Group PoRRITONDYLAKIAE.
Dirhita H. Loew, 1850, p. 21.
345. D. rhodopUla Hardy, 1850, p. 186 {Cecidomyia);
Kieffer, 1913-2, p. 269.
Northumberland and Durham, several records. See
No. 185.
Porrlcondyla Rondaoi, 1840, p. 14.
346. P. hngipes H. Loew, 1850, p. 38 {Epidoais); Kieffer,
1913-2, p. 272.
Winnartila Rondani, 1860, p. 287.
347. [W. teneUa Walker, 1856, p. 129 {Asynapla); Kieffer,
1913-2, p. 283.
n,g,t,7.dh,G00glc
Prdiminary Catalogue of British Cecidomyidae. 403
Subfamily LBSTREMINAE.
Group Campylomyzariae.
Csmpylomyza Meigen,f 1818, p. 102 (also 1830).
348. [C. aceris Meigen, 1818, p. 102; Kieffer, 1913-2,
p. 296.
349. [C. btcoloT Meigen, 1818, p. 102; Kieffer, 1913-2,
p. 296.
350. [C. flavipes Meigen, 1818, p. 102; Kieffer, 1913-2,
p. 297.
351. C. ghbifera Haliday in Walker, 1856, p. 62; Kieffer,
1913-2, p. 297.
352. [C. hatlerala Zctterstedt, 1852, p. 43.51; Kieffer,
1913-2, p. 297.
Amblyspatha Kieffer, 1913-2, p. 299.
•353. A. ormerodi Kieffer, 1913-la, p. 52; 1913-2, p. 300.
Only known from Scotland.
Group Lestbemiabiae.
Catoclia Haliday, 1833, p. 156.
354. [C. brennernn Zetteratedt, 1851, p. 3770 ILeslremia) ;
Kieffer, 1913-2, p. 307.
355. C. latipes Haliday, 1833, p. 156; Kieffer, 1913-2,
p. 307.
Lettremia Macquart, 18^6, p. 173.
356. L. camea H. Loew, 1844, p. 324 (Cecidomyia) ;
Kieffer, 1913-2, p. 308.
357. [L. cinerea Macquart, 1826, p. 173; Kieffer, 1913-2,
p. 308.
t Due to iiwufficiency of deseriplion, many of the Bpeciea in
KieSer's liat are doubtfully included in tlic genus, doubtless including
eome, if not all, of tbe abuve.
ll,y,l,/eJh,G00glc
VH Memn. R. 8. Bsgnall and J. W. H. HftTriaon's
VjH. L./u»raMeigen, 1830,p.309; Kieffer, 1913-3. p. 3U8.
3ij!). L. leiicnphaea Meigen (Juniperina Fabr. non linn),
1818, p. ;i88 iSciara); Kieffer, 1913-2. p. 308.
BubfamUy HETEROPEZISAE.
(!<>nuH MiAHTOR Meinert, 1864, p. 42 ; EieSer, 1913-2, p. 313.
*3ri<).A HUilor Bp. Bagtiall and Harrison, 1918, p. 61.
UiFKiiAM, Hirtley, increasing paedogenetically.
The f(»lIowin(( cloven CeciAomyid spp. are recorded in
Kwiiiiton'tt Itrititth Catalogue.
;i(SO. ('fruhmyitl ap. Swanton, 1901.
On OphioglonHum vulgtiftim. Swanton, 9.
301. (\iulnmyul sp. Fitch, 1883.
i)\\ J Hni[ierHS communis, ^wanton, 12; Houard, 134'
3(12. (Widomifid ap. Zimmerniann, 19(17.
On i'mUiHPti mtn-n. Swanton, 289; Houard, 1167-
363. Cvoldomyid sp. Trail, 1878.
{)i\ Atitliyllus Viihternria. Swanton, 583; Houard, 360*-
1)||RIIA.M, Hirtley.
361. (Wi<hmuulap. Trail, 1878; Kieffer, 1901.
On Vv-iii .tylmlim, Swanton, 591 ; Houard, 3730.
3li'i. Ceoidomyid sp. Binnie, 1877, and others.
On Aiii/rh'fii .tijhvfilrvi. Swanton. 66i; Houard, ^^ "i .
DrmiAM, Vigo; Hillingham, in countless thou*^" '
l>tiiiiei4loy. YoRKsHiRK, tireat Ayton.
306. r,rii!iw!fiil sp. IJebel, Kieffer and others.
On Vrii'iiiiii strpitlliftJii). Swanton, 737; Houard,
367. (Wi,lo„i>/i,l »p. Trail. 1878; Kieffer. 1901.
On tlolium bomile. Swanton. 744; Houard, 5196.
ih,Cooi^lc
eitfi.
PrfHiminary Catalogue of British Ceddomyidae. 405
368. Ceddrnnyidti^. Trail, 1878; Kieffer, 1897; 1901.
On Galium Aparine. Swanton, 764 ; Houard, 5304.
369. Dlploslssp. Trail, 1878; Kieffer, 1901.
On Valeriana sambucifolia. Swanton, 783; Houard,
542-2.
Durham, Waldridge.
Zlfi. CMldomyld sp. Trail, 1878; KieSer, 19(11.
On Senecio Jacobaea and S. aquatica. Swanton, 842,
845; Houard, 5864, 5a58.
Northumberland and Durham, several records.
■k * * * * *
The following three species are shown in Houard as from
the British Isles, but not in Swanton.
*371. Ceddomt/id ap. Trail, 187^; Kieffer, 1901.
On Rhinanfhm Crista-galli. Houard, 5129.
*372. Ceddomyld 8p. Kieffer, 1901.
On Tkalidrum dunense. Houard, 2456.
Northumberland, Watkworth.
*373. Cecidomyid sp. Kieffer, 1901.
On Rosa sptnosissima. Houard, 3237.
The following species are new or recent records.
•374. Ceeidomyid sp. Bagnall and Harrison, 1917-3, p. 230.
On Gmim urbanum, flowers remaining closed.
Durham, Norton and Fencehouses.
*375. Ceoidomyid sp. Bagnall and Harrison, 1916 (iii),
p. 203.
On Betula alba.
Durham, Waldridge and Gibside,
•376. Ceeidomyid sp. Bagnall and Harrison, 1916 (iii),
p. 203.
Seeds of Trolliits europaeus.
Durham, Billingham.
n,g,t,7.dhvC00glc
406 Messrs. R. S. Bi^alJ and J. W. H. Hftrriaon'B
*377. CecidomyidBp. Burkill, 1916, p. 7.
Salix aurUa and S. aurita cinerea, $ catkins.
Recorded by Mr. Burkili from DerbTshire.
*378. ! CHldomyld sp. Bagnall and Harrison, 1916 (iii),
p. 203.
Pod of Cylisits scoparius; cauaer doubtfully Cecidomyid.
Records from NoRTUUMBERLANDaand Durham.
*379. Ceddomyld sp. Bagnall and Harrison, 1917 (v),
p. 15.
Acorns of Quercus Robur and ceirU.
Records from Durham, Yorkshire and Lancashire.
•380. Csoidomyld Bp.f Bagnall and Harrison, 1916 {iii),
p. '203.
In seeds of Geranium pralense and G. sylmliaim with
Das. geranii, larvae bright orange-red to red.
Records from Northumberland and Durham.
•381 . Ceeldomyid sp. ( ? P. geranii). B^nall and Harrison,
1916 (iv), p. 2ry2.
In seeds of Geranium dissecium; larva yellow.
Durham, Fatfield.
•382. Ceoldomyld sp. ( ? P. geranii) Bagnall and Harrison,
1916 (iv), p. 252.
In seeds of Geranium molle; larva yellow.
Yorkshire, Redcar.
•383. Csoldomyld sp. ( ? P. geranii). Bagnall and Harrison,
1917-3, p. 230.
In seeds of Geranium pmillum.
Northumberland, Warkworth.
•384. Ceiddomyld sp. Bagnall and Harrison, 1916 (iii),
p. -203.
In Sowers and amongst seeds of Geranium pusiUum;
larA'a almost transparent lemon-yellow.
Lancashire, Freshfield and Ainsdale, i
I regaid to these Ger&niam
l„y,l,Ajl^,COOl^lC
Prdiminary CaUdogue of British Cecidomi/idae. 407
*385. CMidomyld sp. Bagnall and Harrison, 1916 (iv),
p. 252.
Erodium cicutarium, in flowers and amongst seeds;
larvae creamy-yellow.
Yorkshire, Redcar.
*386. Ceeldomyld sp. Bagnall and Harrison, 1916 (iv),
p. 252; 1917-2, p. 207.
On Pimpinella Saxifraga.
Durham, Fenshaw HiU.
•387. CMldomyld sp. (Perrisia sp.). Bagnall and Harrison,
1916 (iii), p. 200.
On Vaccinium Myrlillvs. Houard, 4566.
Cumberland, Skiddaw, Durham, Gibside.
*388. Csoidomjrid sp. ( 1 Janetidia). Bagnall and Harrison,
1916 (iv), p. 252.
On Euphrasia officinalis.
HeconiB from all counties ; local.
*3S9. Ceeldomyid sp. B^nall and Harrison, 1916 (iv),
p. 252.
On Atriplex paluta.
Records from Yorkshire and Durham.
•39a Ceddomyid ap. Burkill, 1916, p. 7.
On Slachys ■paluslris.
Recorded by Mr. Burkill from Derbyshire.
•391. Cealdomyid sp. Bagnall and Harrison, 1917-3, p. 230.
On Slachys lanaia.
Durham, in a garden at Fenshaw.
•392. Ceddomyid sp. Bagnall and Harrison, I9I6 (iii),
p. 203.
On Camjxirivla rotuvdtfolia.
Records from Durham and Cumberland.
•393. Ceddomyid ap. Burkill, 1916, p. 8.
On AdiiUea MUUfoUum.
Recorded by Mr. Burkill from Sussex.
n,g,t,7.dh,G00glc
408 Messrs. R. S. Bagnall and J. W. H. Harrison's
•394. Ceeidomyid sp. Bagnall and HamBon, 1917-2, p. 230.
On Achillea Millefolium.
Durham, Hart.
*39[>. CsaldDmyid sp. Bagnall and Hamson, 1916 (iii),
p. 203.
On Taraxacum officinale.
Northumberland, Ninebanks.
*396. Cvcidomyld sp. Bagnall and Harrison, 1916 (iii),
p. 203.
In Hieracium boreale seeds.
Durham, Birtley Fell.
*397. Ceoldomyld ap. Bagnall and Hamson, 1917-3, p. 230.
Laivae feeding on mildew on Oak, yellowish.
Northumberland, Warkworth. Durham, Gibdde.
*398. CeeMomyld sp. Bagnall and Harrison, 1917-3, p. 230.
Larvae feeding on mildew on rose; yellonish.
Durham, Oibside. Known to the late Dr. Hardy
(Hardy, 1850).
•399. Cvcldomyld ap. Bagnall and Harrison, 1917-3, p. 230.
Yellow larvae in galls of Andricus fecundaior,
Northumberland, Warkworth. Durham, Winlaton
Mill.
*4(HI. Ceeidomyid sp. Bagnall and Harrison, 1917-2, p. 230.
I^rva milk-white, in leaf-sheaths of a small Carex.
Durham, Waldridge Fell.
*401. Ceoidomyld sp. Bagnall and Harrison, 1917-2, p. 230.
I>arva white, red at each end; in leaf-sheaths of a small
Carex.
Durham, Waldridge Fell.
•402. CeofdomyEd sp. Bagnall and Harrison, 1907-3, p. 230.
Larvae bright rose-red, under leaf-sheath of Carex fiava.
Durham, Birtlev Fell.
n,g,t,7.dh,G00glc
Preliminan/ Caialoffue of British Ceddomtfidae. 409
*iaZ. CMidomyld sp. Cotte, 1912.
Thomlike gall on Galium verum. Houard (supplement},
7372.
Durham, Penshaw and Seaton Carew, rare. Records
from Scotland (Forth area), where it is plentiful.
*404. Ceeidomyld ep. Bagnall and Harrison, 1917-2, p. 209.
On Tkcdifirwn dunense. See Houard, 2443.
NorthumbesljUxv, Warkworth. Durham, coast near
Hart. Also from Scotland.
■405. Cecidomyid ap. Bagoall and Harrison, 1917-2, p.
209 (o).
*406. CecidomywL sp. Bagnall and Harrison, 1917-2, p.
210 (t).
On Aaragalus kypoghllis.
Records from Scotland.
*407; Cetidomyid sp. Bagnall and Harrison, 1917-3, p. 210.
I^rva yellow, in spikelets of PMeum ■pralense.
Durham, Penshaw.
*408. Cseldonvid sp. Bagnall and Harrison, 1917-2, p. 210.
Minute yellow larva in the spikelet of Poa sp.
Northumberland, Ninebanks.
•409.. CKidomyid sp. Bagnall and Harrison, 1917-2, p. 210.
Larvae in spikelets of Dadylus gUymerala.
Durham, Penshaw and Edmondsley.
*410. C«idomyid sp. Bagnall and Harrison, 1917-3, p. 229.
On privet, flowers remaining closed, leathery, containing
a solitary orange-yellow larva. Not Sclnzomyia liguslri of
Riibsaamen.
NoKTUUHBEBLAND, Warkworth.
*4n. Cecidomyid sp. Bagnal! and Harrison, 1917-3, p. 230.
On dead fungus- attacked seed-cases of Lapsana.
Durham, Gibside.
n,g,t,7.dh,G00glc
410 MeBBTfl. R. S. Bagnall and J. W. H. Harrison's
•412. Ceoidomjid ap. Tavarea, 1907.
On Oak. Houard, 1306a.
NoRTHUMBERLuUJD, Warkwoitt. Durham, Fatfield.
Lamcashihe, Grange-over-Sands.
MIS. Cwldomyld sp. ( ? CorUarinia). Bagnall and HarrUon,
1917-3, p. 229.
On Angelica sylveatris.
DuKHAM, Billingham.
*414. CMldomyld sp. Bagnall and Harrison, 1917-3, p. 229.
Minute rose-red larvae on dead seed-cases of Scropkulana
nodosa.
Northumberland, between Warkworth and Alnmouth.
Durham, Gibside.
•415. Ceoldomyid sp. Bagnall and Harrison, 1917-3, p. 229.
Yellow-orange larvae feeding and pupating under the
epiderm of stems of Heradeum Sph<mdylium growing in
marshy place.
Durham, Gibside.
'*'416. Cecidomyld sp. Bagnall and Harrison, 1917-3, p. 229.
Pale reddish larva feeding externally on what look like
parenchymatous galls on Lonicera.
Northumberland, Warkworth.
•417. Cvcldomyid sp. (f = Lasioptera sp.). Bagnall and
Harrison, 1917-3, p. 228.
A rather large bright salmon-coloured larva feeding in
spikelets of Carex vesicaria.
Durham, Billingham.
•418. Ceotdomjrid sp. Bagnall and Harrison, 1917-3, p. 230.
On Helianthemuvi Chamoedslus, flower remaining closed,
larva solitary, pinkish -yellow. Houard, 4267.
Durham, coast between Horden and Hart.
•419. Ceoidomyid sp. Bagnall and Harrison, 1917-3, p.230.
Larvae semi-transparent, whitish, sometimes with yellow-
ish tinge, feeding gregariously in the inner leaf-sheaths of
Carex gUtuca.
n,g,t,7.dh,G00glc
Prdiminary Catahgue of British Ceddomyidae. 411
NoBTHUMBBBLAND, Dear Warkworth. Durham, coast
between Horden and Hart.
*420. Ceeldoinyld sp. Bagoall and Harrison, 1918, p. 61.
On Achillea Ptmmica.
YoRSSHiRB, Bardaey near Leeda.
*421. Ceeldomyidsp. Houard, 5450.
On KnatUia arvensis, leaf pustules.
NoKTHtiMBKRLAND, OvingtOD. DuRHAM, Ryhope.
*422. CMidomyid sp. Houard, S. 60.
On Salix.
NoRTUUMBERLAMD, Ovingham on S. aurita. Durham,
Easisgton on S. Caprea.
*423. CMldomyid sp. Bagnall and Harrison, 1918, p. 61.
In rotting turnips.
Durham, Birtley.
II. Check List op Named SreciEs.
CECIDOMYIDAE.
Subfamily CECIDOMYINAE. Genus Sletaniella Kieffer.
Group Lasioftbbabuk. 1- S. brevipalpiB Kioffer.
Genus CUnorrhnielu H. ,, . . < „ ,.. < .
Log„ ^ Genus Ladoptera (Meigen).
1. C. chrn^anlhemi H. Loew. '■ f »«""P«»r* M^!««»-
2. C. miOefolii Wachtl. f ■ f ■ »'■""'*""* ^.^'.""^ ■■ .,
3. C. UamTUhemi KieSer. '■ ^- calamagrost^u RubsM-
Gmua Trotlerla Kieffer,
= CAon'^oncura (praeocc.) Riibs.
1. T. gain Rubsoamen.
2. T. mrothamni (Kieffer). Group Ouootkophabiak.
3. T. umbdU/eranim Kieffer.
Genus Baldratia KielTer.
I. B. aalieormtu Kieffer. i. N. rhodophaga Cuquillett.
[Genuf
iM-,i,A J h, Google
412 Messrs. R. S. Bagnall and J. W. H. Harrison's
Genus RboiahttnyU Rabsaa-
1. fl. miOffoiii (H. Loew).
= aehiUeae Inclibald.
2. B. lanaeelicola (Karech).
GeDUB Hlsopatha Kiefier.
i. if. fiorum (KielTer).
2. M. foliorum (H. Loew).
= abrolani Trail.
3. M.plarmicae {VMot).
= fiorica Winnerti.
4. M. gyageneaiae (H. Loew).
Otavm AreeuthaniyU Kiefier.
1. A. valtni CVavarea).
Genue OUgotropliUS Latreille.*
1. 0. jutaperinas (LinnS).
2. 0. pante/i Kieffer.
3. 0. alopecuri Reuter.
4. O. burgariua (Bremi).
5. O. fagijtfua Kieft^T.
6. O. hartigi {UeM).
7. O. Teaaimtrianus (F. Loew).
8. O. lympanifex Kieffer.
9. 0. ulrai Kieffer.
10. O, hewianua Kieffer.
Genua PhyctidObia Kieffer.
I. P. so/nwt {Kieffer).
Genus Hlkomylk Kieffer.
I. if. cori/K (Kieffer).
Genus S«mudobia Kieffer.
I. S. betulae (VVinnertz).
Genua Itoomyla Kieffer.
1. I. capnae (Winnertz).
2. /. ma^r (Kieffer).
Genua Janetiella Kieffer.
1. J. fanwei (Kieffer).
2. J. l/iymi (Kieffer).
3. J. Ihgmkola (Kieffer).
4. J. lubtrculi Rlibaaamen.
Genus ^rgoUa KieS«r.
1. Z. carpini (F. Laev).
Genua Cnuieiobia Kieffer.
1. C. comi (Giraud).
Genus Phegobla Kieffer.
I. P. Umtatttta (Bremi).
Genua MayetloU Kieffer.
1. M. avenat (Marchal).
2. «. rfnrfyitrfw Kieffer.
3. if. ikilraelor (Say).
4. M. hoki Kieffer.
5. M. joaimisi Kieffer.
6. M. venlricola RiibsoaDiai.
7. M. kordei Kieffer.-f
Genua Chorlomyia Kieffer.
\. C. hMwigi (RiibBaamen).
2. C moliniae (Riibsaamen).
3. C. poae (Boac).
4. C. radicifica (Riibsaamen).
Genus CysUphon Kieffer.
1. C.hUracii{F. Loew).
2. C. laraiaci Kieffer.
3. C. Ifonlodmitia Kieffer.
4. C pUoi^iat Kieffer.
5. C. aonchi (f . Loew).
Genus Haorolabls Kieffer.
1. Jf. corragaria (F. Loen).
2. if. hieracii Kieffer.
3. if. hippocrepidia Kieffer.
4. M. jnartdi Kieffer.
5. M. piioaeUae (Binnie).
6. if. 8(eUarta«(Liebel).
Genua Arnoldia Kieffer.
1. A. qnercicoia Kieffer.
2. ^. guercua (Binnie).
Genus G«oerypt« Kieffer.
\. Q. braueri (Handlirsch).
• Tlie first two species truly belong to tlua genus aa now 1
but all tbe otiier H|)Ccies arc placed here temporarily.
f Tbe generic position of this species is not yet certain.
iM-,i,A J h, Google
Preliminary Catahffue of British Cecidomyidae. 413
Genus Rhtbdopluiga We«twood.
1. A. ofin'pfnnig (H. Loew).
2. B. cUtvifex (Kieffer).
3. R. dttbiomt Kieffer,
= dubia Kieffer,
= griaeicoUit Zett. no»
Heigen.
4. It. giraudiana Kieffer,
= aalieiperda Giraud noH
Dufuur.
5. B. hflerMa (H. I>oew),
= ^igiia Hardy.
6. R. iieobia (Kieffer).
7. B. iwwAi (Kieffer).
8. B. margiiifmioTiitiens (Win-
fl. fl. nerrorum (Kieffer),
= nodali Biibfia&men.
10. B. piercEi (Kieffer).
11. B. pteiutoeoecus RubeaaaieB.
12. B. patvini Kieffer,
= gnluina Glraud turn
Schrank.
= T Hugi H. Loew non
Meigen.
13. R. roiaria H. Loew,
= Mlicina Auct,
^ cinemirum Hardy.
14. B. Toaarietla Kieffer.
15. B. galifiperda (Dufour),
= lerfbnirti H. l^oew.
16. B. miticia (.Schmnk),
= digeeri Bremi.
17. A. jiipema Kieffer.
IS. it. terminatia (H. Loew).
GenuH Pairisla Rondani.
1. /*. oiic/iperda (Heiuichel).
2. /*. ac«rcri«paiM (Kiellcr) and
var. rubeUa (Kieffer.)
3. P. aerophUa (WinnertE).
4. P. oy^TiM (Kieffer).
5. P. alni (F. Loew).
6. P. alpina (F. Liiew).
7. P. anglica Kieffer.
8. P. aparinea (Kieffer).
9. P. aiicupariae Kieffer.
10. P. ajMirig Kieffer.
11. P. balhlM (Rubsaamcn).
12. P. beckiana (Mik).
13. P. braaticae (VVinnerIz).
14. P. bnmdUK Kieffer.
15. P. brgtmiat (Bouc)i^).
TRAK8. EKT. SOC. LOND. 1917.-
16. P.campanulae{Rab6iMmBo).
17. P. capiligena {Brtmi).
18. P. eardamintt {Witmertz).
19. /•. cn'a<(H(Binnie).
20. P. rt>ni(Riib8aamen).
21. P. amtpoaitarum {KAefln).
22. P. con/lina Kieffer,
=3 coryli Riibeaaoien.
23. P. erataegi (Winnerti).
24. P. daphnes Kieffer.
25. P. engglfeldi (Rubeaainen).
26. P. Epifo6.i(F. Loew).
27. P. erieina iF. Utev).
28. P. fUkina (Kieffer).
2D. P. fionperda (F. Loew).
30. P. fioiculorum (Kieffer).
31. P. /rajTinai Kieffer.
32. P. fraiini Kieffer.
33. P. /rtiduum (Riilieaamen).
34. P. gaieobdokmlis (Winnertz).
33. P. gain {a. Loew).
36. P. galiieola F. Loew.
37. P. ^enM<anifoi^e>M(Kieffer)
38. P. genidicola (F. Loew).
39. P. gentianat Kieffer.
40. P. gtranii Kieffer.
41. P. fffcfiowuw (Kieffer).
42. T P. gli/cipki/Ui RUbaaamen.
43. P. holosleae Kieffer.
44. P. hffgrophita {m^L).
45. P. Ayperic."(Bremi).
46. P. ignorata (Waclitl),
= medicaginia Bremi,
= onobrychidis F. Loew non
47. P. inchbMiana (Mik),
= ? daugitia BreDli.
48. P. inrJUMi (Frauenfeld).
49. P. tiel/en Marclml.
50. P. hie/feriaHa (Kiibeaamen).
51. P. Utmii Kieffer.
52. P. laricig F. I»ew,
= keUneri Henscliel.
63. P. lalAgn Kieffer.
54. P. kUkyricaUi (Riilmaaiiieu).
55. P. libera Kieffer.
56. P. fo(Aan"n(jiae (Kieffer).
57. P. loti Kieffer.
58. P. /oltco^ (Riilwaamen).
50. P. hipuUme Kieffer.
60. P. Iffch^UU IH^ydm).
61. P. mtJj Kieffer.
62. P. malpighii Kieffer.
63. P. «»jtii Kieffer.
-PAETS II, III, IV. (may '18) EE
u,y,i,Ajh,Cooglc
414 Messrs. B. S. BagnaU and J. W. H. Harrison's
64. P. muricatat (Meade),
65. P. jttrvieola Kieffer.
66. T P. onobrychidit Bremi,
= gimudi Frauenfeld.
67. P. papaveria (Winnertz).
68. P. pmcl^menHRnhBoamm).
69. P. perskariaf (LiimS),
70. P. plieatrix (H. Locw).
71. P. folygaiae Kieffer.
72. P. popufc/t(RubBaainen).
73. P. ■poteidiOae (Wachti).
74. P. praiicola (Kieffer).
75. P. pleridicola (Kieffer).
76. P. pMtulane {Rvibamiaen).
77. P. pyri (Bouch^).
78. P. pierrcana Kieffer,
79. P. ranimeuli {Bremi).
80. P. raphaniefri (Kieffer).
81. P. ThododtndToni Kieffer.
82. P. Toaariiitt (Hardy).
83. P. rosirupiana Kieffer.
84. P. rubaaanuni Kieffer,
85. P. aalicariat (Kieffer).
86. P. Mtiffaisorbae (Kieffer).
87. P. saOiosae (Kieffer).
88. P. achkchleiidali (Kieffer),
8fl. P. achmidli Riibaaamen.
90. P. «erolina (Winnertz).
91. P. ailmaila Kieffer.
92. P. similia (F. Loew).
93. P. msymbni (ticliranck).
94. i P. mdalia (V. Laev.).
95. P. gtadiydis (Bremi).
96. P. glrobi (Winnertz).
97. /*. (e/enstfRiibsaamen).
98. P. lelrakil Kieffer.
99. P. Ihomasiana (KiefieT),
100. P . liliamvolveitB (RUbaaa-
= tiliae iSchranck,
101. P.lorlrirlF. I.oew).
102. P. tracheliHWacM).
103. P. train Kieffer.
104. P. Mfolii (F. Ijoen).
105. /■.(u6(coia (Kieffer).
I0«. P. vlieia Kieffer,
107. P. vlmariae (Bremi).
loa P. ulmicola Kieffer.
109. P. urtUae (PerriM).
110. P.wiwwuwMm Kieffer,
= vaecinii Riibaaamen m
i^mith.
111. P. veronieae (Vallot),
= chamaedryt Inchbald.
112. P. vtctiM (Kieffer),
in. P.violae{F. Loew).
114. P. virgae-aurtae (Uebel).
115. P-vUie-idataeKieBer.
116. P. lilhofpermi H. Loew.
117. P. panf^i Kieffer,
OenuB HartiglolS Riibeaameo.
I, H. anmdipa (Hartig),
= pHifffr H. Loew,
Genus Taxomjris Rubea*men.
1. T. taxi (Inchbald).
Group ASPHONDYLARIAF.
Genus SchiiomyU Kieffer.
1. S. po/ioT-iim Kieffer.
2. S. Iigu4lri Riibsaanien.
Genua KieOeris Hik,
Genus Asphondylla H. Loen-.
1. ^. dorycnii V. Luew.
2. A. genUtae H. Loew.
3. A. lupalinat Kieffer.
4. A. moueri Liebel.
5. A. tniki Wacbtl.
6. A. mtlaaopue Kieffer.
7. A. (Monidie F. Loew.
8. A. proximo Kieffer.
9. A. sarothamni H. Loew.
10. -i.%mt Kieffer.
11. A. u/tcM Verral.
Group Brachynkhkaeub.
Genus PhMnatsuthla Kieffer.
1, P. cardui Kieffer.
Genua HJklola Kieffer,
1. Jf./a^"(Hartig),
= Tipula rubra Hermann.
Genus Brschjmeun Rondani.
1. A slygia Walker (n«;
Meigen).*
• See Collin, 1904, note 3.
u,y,l,/eJt,G00t5lc
Prdiminary Catahgue of British Ceddomyidae. 415
Group CEciDOHnABUB.
Genue Harmomyia H. Loew.
1. H. /M^W Frauenfeld.
2. ff./rirentKieffer.
3. H. grandis (Meigen).
4. H. knmckeri Kieffer.
GenuB Dishormomyia Kie&er.
1. D. comi/tt (Kieffer).
Genna Trisbormomyia Kieffer.
I. T. tvitHfica (RUbBaamen).
Geniu Cyrtodiplosls Kieffer.
1, C. crasstnerva (Kieffer).
Genus HoBarthrop«lpus RQb-
1. M. bazi (Geoffrey).
Genua Dlchrona Rubea«nien.
I. D. gaOaram Rubaaamen.
Genus Arthroenodax RUbsaa-
1. A.fTaxinella (Meade).
Genus Aphldoletes Kieffer.
1. A. abi^is (KiefTer).
Genus Anataemla Kieffer.
1. ^.brffeMjyi (Kieffer).
2. A. viciae KieRei,
=. Clinodiplona longiventrU
Kieffer (larva, not imago).
Genua Hadrobremla Kieffer.
1. H. hngiventrig Kieffer,
= ClinodijiIoeM trifolii Kief-
fer (larva, not imago).
Genus Endaphts Kieffer.
1. E. perfidtit Kieffer.
Genus SyndlplosiS Riibsaamen.
1. S. lonkertarum (F. I»ew).
2, 8. p^ieii (Kieffer).
Genus Contkrinla Bondani.
1. C. acdoaae Kieffer.
2. O. ajitbobia F. Loew.
3. C. barbickei Kieffer.
4. C. bflulicola Kieffer.
5. O. betidina Kieffer.
6. C. eampamilae Kieffer.
7. C. corsit Kieffer.
8. C. craccae Kieffer.
9. C. cucubtdi Kieffer.
10. (7. sei Kieffer,
11. C. hdianthemi (Hardy).
12. C. heraelfi Riibeaamen.
13. C. lalhyn Kieffer.
14. C. linariae (Winnerti).
16. C. hli (De Geer).
16. T C. mfdicaginis Kieffer.
17. C. jnekmocera Kieffer.
18. C. naetUTtii Kieffer.
19. C. nicoiajfi Riibsaamen.
20. C. tnumidit Kieffer.
21. C, pyrivora (Riley).
22. C. qutreina Riibsaamen,
= dryophila Kieffer,
23. C. nOicola (Kieffer) RUbs.
24. C. nuUrrdia Kieffer.
26. C. gcabioeae Kieffer.
26. C. acopuHi Riibsaamen.
27. C. #of6i Kieffer.
25. C. «f«ini Karsoh.
29. C. tiliarum Kieffer.
30. C. tragopogoaie Kieffer.
31. C. (roi(i Kieffer.
32. C. tremulae Kieffer.
33. a Iriiiei (Kirby).
34. G. vaierianat Riibsaamen.
35. C. vihruTUomm Kieffer,
= vibami Kieffer Turn Felt.
Genus SUctodlplO^ Kieffer.
1. S. corylina (F. Loew).
2. 8. hypodiotridis (Rubsaa-
3. S. jaaAaeae (H. Loew).
4. S. piloidlae Kieffer.
5. S. aeropkulanae Kieffer.
6. 8. umbeOalarum RUbsaamen,
Genus Thecidodiplosis Kieffer.
1. T. brackyrUera (SchwSg-
richen).
n,g,t,7.dh,G00glc
416 Messrs. R. S. Bagn&U and J. W. H. Harrison's
GenuB Hyrioomyis Kieff«r.
I. M. medUerranea (F. Luew).
Genua Zeuxidiplosls Kiefler.
1. Z. ffwrrft (Kieffer),
= giarduma Kieffer.
Genua AtrlchoselU^ KiefTer.
1. A.acerisKieSleT.
Genus TrichodiplosiS Kj«fTer.
1. r. carim (Kieffer).
Genua Anticblrldium RUbsaa-
= ^n/tcAi>aRilbsaamenntm
KacliHcholz.
1. A. ttriatiim (Rubeaamen).
Genua Lestodiplosis Kieffer.
1. £. coHiiia (Winnertz).
2. L. ctiiiralie (Winnertz).
Haerodiplosls Kieffer.
1. M. dryobia (F. Loew).
2. Jtf. volvtns Kieffer.
Genus Xenodip[osls Felt,
— Allodiplogig Riibsaamen
turn Kieffer.
1. X. laeviuxtili (Riibeoamen).
Genua Pwallelodlpla^ Riib-
aaamen.
I. P. gatiiperda (F. LoewJ.
Genus Hsrmuidis Kieffer.
1. H. Iremviof. (Winnertz).
2. H. puaulans Kieffer.
(ienu9 ClinodiploslS Kieffer.
1. C. belonieae Kieffer.
2. C. bolularia (Winnertz).
3. C ro«tp<nia (RiibsaaitKn).
4. O. scUechlendali RQbsaamen.
Genua Pseudhofinomyia Kieffer. '^™"" Myeoilpl"!* Rubsaameo.
1. P. granifex Kieffer.
Genus Tftpho4[plosis Kiefler.
I. T. subtermnta (Kieffer tt
Trotier).
Genus Dyodlplosls Rubaaamen.
1. i*. (imiariCTe (Rubsaamen).
Genua Lo«Wio|g Kieffer.
I. L. renlmiTtat (V. Jxtevi).
Genua Putoniell* Kieffer.
1. P. mursupialu (F. Loew).
Genua Haplodlplotis Riibsaa-
I. H. etjiiealris (B. Warmer).
Genua Ceeidomyla Meigen.*
I. C. yin »■ (de Gecrj.-f
niojAaga (Winnertz).
Genus Massalongia Kieffer.
i. M. ruhra (Kieffer).
Genus Amstrodlplosis Riibeaa-
1. A. Ihalw-fnfola (Rubsaamen).
Genua MoDodlpIosiS RDbsoamai.
I. M. lifbdi iKiftSer),
= Seh izomyia «or.iahilis Riiba.
Genua Atylodl^OSlS RUbMameo.
I. A. rumtcis (H. Loew).
Group PORRICONDYLABIAE.
Genua Dlrtliu H. Loew.
1. D. rhodophila (Hardy).
Genua PoiricondylA Rondani.
"■ ^- ^'^"^ ^^'"■"S' 1. P. longipes (M. Loew).
,1 J" '^''T*^'^'"/^''* /«"""' eJicepting the type pini, ape^uflufficientiy
deeenbed, and therefore ^nnot be relegated to any kno>™ genus.
t bynonyma are lalerella Zett.. piloM Bremi, and pini-maritijnae
n,g,t,7.dh,G00glc
Prditninary Catalogue of British Ceddomytdae. 417
SuMamily LESTREMINAE.
Group Cahpylomyzariae.
OenuB Cunpylomyza Meigen
1. C. ghbifera Hftliday.
Genus AmblyspiUu Kieffer.
1. A. ormerodiKi^Ser.
Group Lestbemiariae.
Genus CatOCha Haliday.
1. C. {aftpe« Haliday.
Genus Leslremla Macquart.
1. L. camta (H. Loew).
2. £./u«ca Meigen.
3. L. Uucophafa (Meifien),
= junipeTtJUi Fabriciua no
III, Index to Host-plants.
N.6. — The capiUil initiala of spedfie plant-names are in accordance
with the usual Botanical practice.
In using the following index it is well to note the follow-
ing points: (1) that when a Cecidomyid galls all of the
species of a plant genus, then that fact is notified thus,
" Salix spp." ; (2) that in a few cases an aggregate plant
name is used; (3) that when any Cecidomyid feeds on
rusts, smuts, mildews, aphids and the like, found on any
particular plant, then these are indexed under the name
of ttiat plant; (4) that the numbers refer to the numbering
of the Cecidomyid species in the classified list.
Acer. AnthrUcua.
Anthriacus aylveelria HolTin..
54, 9.
AntkyUU.
Anthyllis Vulneraria L., 363.
Arltmieia.
Artemisia Abrutanum L., 17.
vulgaris L,. 16, 17.
Acer Pseudoplatanus L., 91,
261, on -4pA«» pio/an-
oidta on tliia tree.
„ campestre L., 92, 312.
AdiiUta.
Achillea Hillefolium L., 2, 14,
18, 393, 394.
„ Ptamiica L., 2, 14,
18,420.
AInuB rotundifulia Mill, 96.
AtopecuruB pratensia L,, 24.
Angelica sylveetria L., 229,
366, 413.
Aidhtmia.
AnthemiB Cotula L., 1, 19.
Astragalus
Astmgalus liypiiKloti
136, 168, 406, 406.
Atriplej: (boo also Obione).
Atriplex patula L., 389.
Avena fatua U, 46.
BitOola.
BallotA nigra L., 101.
BelaUi.
Betula alba L., 34, 269.
341, 375.
D. a.
u,y,l,/eJh,G00glc
418 Messrs. R. S. Bagnall and J. W. H. Hairison's
BratAypodium. Centanrea ScsbioHa L., 162,
Braohypodium sylvatioum R. 163, 164, 320.
and 8., 53. CeraHium.
Cerastium Tulgatum L., 109,
126, 154.
„ Tiscosura L., 109.
„ semidecandrum L.,
109, 126.
CItTffsanlhemum.
Qiiyaanthemum Lencanthe-
mumL.,3.
Ciraium.
Ciraiumsp., 241 an inquiline
in galls of Tryptta
„ arvemBe Scop., 110,
)U, 112, 339 on
Puedma suavoltMt,
„ lanceolatum Soop.,
113,114.
Corivolvulue.
Convolvulus aepium L., 336.
ComuB sanguinea L., 43.
Corytus.
Corylus avellana L., 30, 33,
115,272,303.
Cralaegu*.
Crataegus Oiyaoontha L.,
lie. 267.
Crepis biennis L., 304.
„ paludoea Hoench, 338
on Pvtxinia major.
Cytiaus.
Cytisus BCOpariuH Link., 5, 40,
212,233,378.
DaclylU.
Dactylia glomeratA L., 46,
404.
Daphne.
)>aphne Laureola L., 117.
Daiicut.
Daucus CaroUi L., 229.
Doryonium sp., 230.
Epiiobiiim.
Epilobium at^ustifolium L.,
120, 146.
Erica.
Erica cinerea L., 121.
.. Tetraliz L., 310.
Erodium.
Erodiumctcutaiium, 134,385.
BraBsica spp., 103, 182, 283.
Brvmus.
Bromtu erectus Huds., 52.
Bryonia.
Bryonia dioica Jaoq., 105.
Bupleurum.
Bupleurum tenuimimum L.,
302.
BuxuB aemperviveni L., 262.
Carnpanvia.
Campanula media L., 106.
persicifolia L.,106.
rotundifolia L.,
207, 271, 392.
CardamiTu,
CaTdamine spp., 108.
Carex spp., 253, 400, 401.
„ binervis Sm., 246, 250,
„ arenariaL.,280,319.
„ vulpina L., 166.
„ pendulaUuds., 166,314.
„ helodee Link. (= C.
laevigata Sm.), 166.
„ palleacens L., 164.
„ sylvatica Huds., 166.
„ acutiformia Ehr., 166.
„ limosa L., 246.
„ flava Host., 246, 249.
253, 317, 318, 402.
„ echinata Uurr. (= stel-
lulata Good.), 248.
„ elaU All. (= C, stricto
Good.), 249. 250, 253,
317.
„ Goodenovii Gay, 263,
313.
„ paniculata L., 317.
„ divulsa Stokw, 318.
„ distatiR L., 318.
„ giauca, 250, 313,419.
Carpi HUB.
Carpinua Betulua L., 42, 187.
Ca^anea.
Castanea aativa Mill, 362.
CerUaurea.
Centaurea nigra L., 162, 163,
164, 320.
l„y,l,Ajh,COOl^lC
Preliminari/ Catalogue of Briltsk Ceddomyidae. 419
Evpiorbia.
Euphorbia Eoula L., 107.
£uphnuia.
Eupbntsia officinalis U, 388.
Farus sylvatica, 26, 27, 44,
226, 242.
Fraxinus.
FraxinuB excelsior L., 93, 124,
126, 254 in naHot EriophytM
fraxini; 334 inquiiiae in
gall of Per. fraxini.
Oakopsit.
GaleopBiB Tetrahit L„ 202.
Oalium.
Galium epp., 130, 228.
„ verumL..4, 128, 129,
301, 403.
paluBtreL., 138, 139.
„ twreale L., 367.
„ Aparine L., 98, 368.
Oenliana.
Gentiana campestris L., 133.
Amarella, 133.
Genista tinctoria L., 131, 132,
231, 2S2.
Geranium.
Geranium pratense L., 134,
380.
„ sylvaticum L., 134,
380.
„ dJBaectum L., 134,
molle L., 134, 382
pusillum L., 134,
383, 384.
„ saDgiiineum L., 134.
Geum rivale L., 275.
„ urbanum L., 374.
Qlec&imia. See Sepeia.
Hedera Helii L., 145.
Htiianthemum.
Helianthemum Chamaecistus
Mill, 276, 418.
Utlianikus.
HelianthuB sp., 165.
BeracUutn.
Heracleum Sphondylium L.,
63,277,284,415.
„ var. angustifo-
lium, 63
HieTocium.
Hieracium spp., 57. 64.
Pilosella L., 66, 67,
167, 306.
boteale Fr., 396,
340 on Piteciiiia
hieracii.
Hippocrepis.
Hippocrepis comosa L., 65.
HolcuH laaatuB L., 48.
Hordeam.
Hordeum rulgare L., 47,
51.
„ pulclirum L., 72,
140, 103, 311.
„ humifuaum L.,
140, 193,
„ perforatum L., 72,
140, 193, 311.
„ montaDumL.,140,
elodeaL.. 141.
Hypochaerie.
Hypoohaeris radicata L., 62,
304.
Inula squarroaa Bern., 102.
JuniptTna.
Juni{)eni8 communis L., 21,
22, 23, 361.
Kiuinlia.
Knautia arvenais, 421.
Latniuirt.
Lamium album L., 63.
„ maculatum L., 147.
„ GaleoWoIon Crantz,
127.
LMpaana.
Lapsana communia L., 411, on
Larix decidua Mill, 148.
Lathyma.
LatliyruB pratensis L., 149,
150,151,152,258,
278.
„ nmcrorhizus Wim.,
iM-,i,A J h, Google
420 Messrs. R. S. Bagnall and J. W. H. Harrison's
I vulgare h., 228a,
410.
Liimria.
Linaria vulgaris Hoench., 279.
Li t kosptrmv in.
liithospprmum officinale 163 a.
Jjonieera.
l»nicera Periclymenum L,
170, 416.
Lolut
:ulat
156,
Lychnis.
Lychnis dioica L., 293.
„ albn Mill, 15U, 293.
„ FIcw.cuculi L., 176.
Lythnim.
Lvtlinim Salicarift L., 188.
Matricaria.
Matricaria inoduia L., I, 19.
Medicago.
Medicagi' Hativs L., 142.
falcata L., 281.
• „ lupulina L., 157,
158, 232, 234.
Midiuiit.
Molinia cacruleo Muench, II,
50,64.
Myiimlia.
MyoHotis sa)riiioidcH L. (= ar-
venxiM Lam.), 225.
Xf|icia liederacen
lioler
I Hoi
.rd).
25, 135.
Obione.
Ubiiine i>ortulne(>ides Mug.,
Oiionia.
Ononis rcj>enB L., 236 ; 265 on
A^is on Ononis ; 285.
Ophioghisitiim.
OpliiiiBluBsum viilgatum L.,
360.
Faliaver Rlioeas L., 169.
Phragmiieii.
PliraKniitea eommunis Trin.i
7. 144, 314.
Phaturis.
PtiBlariB anindinacea L., 8.
Pioea excdaa Ijam., 90. 200;
256 and 257 botli in
Adelges galls.
Pimpinella.
Pinipinella Saxifraga L., 9.
229. 296, 386.
„ magna, 308.
PinuB aylTeetrie L, 309.
Planlago.
Plantagu lanceolaU L., 192.
Poa.
Poasp., 408.
„ nemoralis L., 49, 35, 56.
Potygala.
Polygala vulgaris L, 173.
Polggonum,
Polvgonum amphibium L.
171.
Pnpulug.
Pupulus alba I^, 76.
„ tremula L., 76, 174,
266. 297,330.331.
„ alba X tremula, 174-
PolfitliUa.
Polentilla ap., 175.
PrunfUa.
Prunella vulgaris L., 104.
PninuBspp.. 197. 206.
„ sjiinosa Toum., 321.
„ (on dry pluma. 332.)
Pifris.
PteriH aquilina L.. 121a, 177.
Pviiraria.
Pull carl a dysenterica S. t-
Qray, T102; 337 on Uro-
mgces junci.
Pyr'us communiH L.. 179, !8B.
Aucuparia L., 99, 292.
„ MaluH L., 160; 255 on
Aphis mali.
Quercus.
Qiiercus Robur L. and seaailis
Ehr., 69. 70, T.
163, 161. 168a,
287. 328, 327. 379.
412; 328, inqoi-
line on A'turoKr*'
latviusculvt; 329,
ll,y,l,/eJh,G00glc
Prdiminary Catalogue of British Cecidomyidae. 421
ScnypKniaria.
^rophotftria nodosa L., 307,
inqiiiline on A'mro.
tentt laUicHiant ;
343. inquiline on
MaCTolabig dryolria
and valrtns; 397,
nokk:
399, inquilii
ffalls of Aadrieus
Quercua cenis, 379.
Ratiuneuhis.
Ranunculus spp., 161.
aciis L., 208.
AAiiHinfAtM.
Rhi nan thus Criata-galli U,
371.
Bhododatdron.
Rhododendron femigineum,
163.
BibrM.
Ribea GroHBularia L., 201.
„ nipnim L., 201.
Rom.
Rona Bpp.. 13o, 184, 185. 335.
34.'}; 398 on mildew.
,. HpinoEissima L., 373.
Rubuaspp., 10, 172.
.. caesiuB L.. 288.
Rumex Acetosa L., 206.
AcetOHPtIa 1^, 266.
344.
Satirnmia.
Salicomia radicana Km., )2.
tiatix.
Sftlixspp., 35, 73. 74. 75, 77.
79.80.81,828.83,84,
87,
„ aurita L., 36,
89. 422 auri
rea. 377.
.. cineiea L., 78, 82, 87.
180.
., rcpens L., 82.
, Caproa L.. 82a, 87. 422,
cinerea x viminalic,
87.
., allia L., 143.
Sanjuiaorba.
Kanguisorba officinalis L., 189.
Scabioaa Columbaria L., 190,
414.
Sfntcio.
Senecio Jacobaea L., 305,
370.
„ erucifoliua L., 305.
„ aquaticus Huda.. 370-
SOtnt.
Silene acaulis L., 96.
„ inflatttSm., 122, 277.
SigymiiTiam.
Sisymbrium officinale iSoop.,
196, 289.
StUdago.
Solida^ro Virgauiea L., 222,
223.
Sonchug.
Sonchus arvenais L., 61.
Spirata.
tjpiraea Ulmana L., 118, 119.
178. 186, 214.
.. FUipendula, 21S.
Slachys.
yiachys sylvalica L., 63. 198,
251.
„ palustria L.. 198, 199,
390.
Belonica Benth., 333.
lanatua. 391.
StrOaiia.
IStellaria graminea L., 68. 137,
HoloBtea L.. 194.
Tanartlam.
Tanacetum vulgare L., Ifl.
TaToiacum.
Taraxacum officinale L., 58,
395.
Tiurua.
Taxus baccata L., 227.
Thalidrum.
Tlialictrum minus (agg.) U,
342, 372.
Thymus.
ThvmuB S«rpvllum L., 38, 39,
41, 237. 239.
TUia.
Tilia Bjip., 28, 20, 203. 204, 205.
Tragiypogon.
Tragopogon pratensis L., 296.
Trifolium.
Trifoliumspp., 100.
„ Iragiierum L., 100.
iM-,i,A J h, Google
422 Messra. R. S. Bagnall and J. W. H. Harriaon'B
Tiifoliiun piutense L.,
210, 260.
repens L., 211.
Triticum ap., 322.
TroUius.
TrolliuB europaeuB L., 376.
Typka.
Typha latifolia L., 314.
Vki.
Ulex euiopaeuB L., 213, 240.
Ulmus spp., 216.
„ campestriB L., 31. 37,
var. BuberoBs, 37.
„ montana Sm., 31, 37.
Urtica. .
Urtica dioiCB L., 217.
„ ureDB L., 217.
Vaccinivta.
Vscciniutn Myrtillus L., 21S,
387.
Vitia-idaea L., 97,
234.
VakriaTta.
Valeriana sambucifolja Hik
(= officinalis auct,), 299,
369.
Veronica.
Veronica ofBoinaiiB L., 195,
219.
„ ChainaediTe L., 196,
219.
„ Berpyllifolia L., 219,
366.
soutellaU L., 196.
„ montana L., 219.
Vifeufnum,
Viburnum Lantuia L., 32.
„ Opulus L., 264,
300.
Vicia.
Vioia Bpp., 220, 273.
„ aepium L., 259.
„ sylvatica L., 364.
Viola.
Vioja app., 84.
„ arvraiBia Murray, 221.
IV. References in Brief.
Bagnall, R. S. 1916. (II) The EntomologUt's Eecord,
V. XXVIII, pp. 158-159.
„ 1917. Lanes, and Cheshire Naturalist,
pp. 253-262 and 282-290.
•Bagnall, R. S., and Harrison, J. W. H. 1916. Jbidm,
(I) pp. 51, (III) pp. 197-203, (IV) pp. 247-252.
Bagnall, R. S„ and Harrison, J. W. H. 1917. Ibidem,
(V), V. XXIX. pp. 206-210.
Bagnall, R. S., and Harrison, J. W. H. 1917-2. IbiJm,
V. XXIX, pp. 12-15.
Bagnall, R. S., and Harrison, J. W. H. 1917-3. Ibidm,
V. XXIX, pp. 228-230,
Bagnall, R. S., and Harrison, J. W. H. 1918. Ibidem,
v.XXX, pp. 61.
BiNNiB. 1877. Proc. Nat. Hist. Soc. Glaegow, v. 3.
ii,y,i,/ejh, Google
■Preliminary Catalogue of British Cecidomyidae. 423
Bosc. 1817. Bull. Soc. Philomat., Paris.
BoucHE. 1847. Stett. Ent. Zeit., v. 8.
Bremi. 1847. Beitr. zu einer Monogr. der Gallmiicken.
BuBKiLL, H. J. 1916. The Entomologist, v. XLIX,
pp. 4-8.
1916-2. Derby- Arohaeol. and Nat. Hist.
Soc. Joum., pp. 29-48.
1917. The Entomologist, v. L, pp.
82-85.
Carpenter, G. H. 1911. Journ. Econ. Biol., v. 6.
CoNNOLD. Plant Galls of Great Britain, 1909.
CoQUiLLETT. 1900. Bull. U.S. Dept. Agric. Washington,
V. 22.
COLUN, J. E. 1904. Ent. Monthly Magazine (2), v. 15,
pp. 93-99.
Curtis. 1827. Brit. Ent.
De Geeb. 1776. M^m. Hist. Ins., v. 6.
DuFOUR. 1841. Ann. Sc. Nat. (2), v. 16. .
Felt, P. 1911. Joum. New York Ent. Soc., v. 19.
Frauenpeld. 1862. Veih. Zool.-bot. Ges. Wien, v. 12,
1867. Ibidem, v. 17.
Geoffrey. 1764. Hist. Ins. Paris, v. 2.
GiRADD. 1863. Verh. Zool.-bot. Ges. Wien, v. 13.
Hauday. 1833. Ent. Mag., v. 1.
in Walker. 1856.
Handlirsch. 1884. Verh. Zool.-bot. Ges. Wien, v. 34.
Hardy. 1850. Ann. Mag. Nat. Hist. (2), v. 6.
Harrison, J. W. H. 1917. Entomologist, v. L, p. 244.
Habtig. 1839. Jahresb. il. Forstschr. d. Foretwiss., v. 1.
,, 1844. In Ratzeburg's forstinseklen, v. 3.
Heegeb. 1851. Sitzber. Akad. wiss. Wien, v. 7.
Hentschbl. 1880. Centralbl. f. d. Ges. Forstwesen, v. 6.
Heyden. 1861. Correspondenzbl. fiir Sammler von In-
sekten, v. 2.
HouARD, C. Les Zooc^idies des Flantes d'Europe. 3
vols. 1908-1913.
Inchbald. 1861. Ent. Weekly Intellig., v. 10.
Karsch. 1879. 7 Jahresber. \Vestf. Prov. Ver.
1881. Beri. Ent. Zeitschr., v. 25.
KlEFFBR. 1886. Zeitschr. f. Naturwiss., v. 59.
1888(-1). Verb. Zool.-bot. Ges. Wien, v. 38.
„ -2. Ent. Nachr. Berhn, v. 14.
1889(-1). Ibidem, v. 15.
„ -^2. Wien Ent. Zeit., v. 8.
n,g,t,7.dh,G00glc
424 Heasn. R. 8. Bagnall and J. W. H. Harrison'^
KrEFPEB. 1899-3. Bull. Soc. Amis. Sc. Nat. Roneo,
V. 34.
1*<9*>{-I). Wien Ent. Zcit.. V. 9.
.. -2. Ent. Nachr. Berlin, v. 16.
.. -3. Verb. Zool.-bot. Ges. Wien, v. 40.
1891. Berl. Ent. Zeitachr., v. 36.
189-i{-I). Wien Ent. Zeit., v. 11.
„ -2. Berl. Ent. Zeitachr., v. 36.
1894(-1}. Bull. Soc. Ent. Franw, V. 63.
„ ., -2. Feuille Jeunes Natural., v. "24.
1895(-1). Wien Ent. Zeit., V. 14.
„ -2.. Bull. Soc. Ent. France, v. 64.
„ .. -3. Feuille Jeunes Natural., v. 26.
„ -4. Miscell. Ent., v. 3.
1896(-1). Wien Ent. Zeit., v. 15.
„ „ -2. Bull. Soc. Ent. France, v. 65.
„ -3. Miscell. Ent., v. 4.
1897(-1). Berl. Ent. Zeitachr., v. 42.
„ -2. Bull. Soc. Ent. France, v. 66.
,, ., -3. Meine Antwort au den Herm Zei-
chenlehrer Rilbsaamen.
]898(-l). Miscell. Ent., v. 6.
„ „ -2, Synopse des Cecidomyes d'Europe et
d'Algerie.
19l)(). Ann. Soc. Ent. France, v. 69.
l'JOl(-I). Ibidetn,v.70.
„ -2. Bull. Soc. Hiat. Nat. Metz, v. 21.
19(I4{-1). Ibidem. V. 2Z.
„ ,, -2. Ann. Soc. Scient. Bruxelles, v. 28.
19()6. Mitt. Philom. Gea. Strassb., 1905,
V. 3.
19»7{-1). Naturw. Zeitschr. f. Land and
Forstw., V. 5.
„ -2. Marcellia, v. 6.
I1HI9. Bull. Soc. Hist. Nat. Metz, v. 26.
1912{-1). Marcellia, v. 11.
,. ,, ~2. Neue Gallniiicken-Gattuugen, Bitech.
1913(1). Bull. Soc. Hiat. Nat. Metz, v. 28.
,, „ -In. Marcellia, v. 12.
,, ,, -2. In \\'ystraan'3 Genera Inaectonim,
fasc. 152.
KiEFFER anil Trotter. 1W).">. Marcellia, v. 3.
KiRBY. 1797. Trans. Linn, Soc. London, Zool., v. 4.
Latreillk. 18(1."). Hiat. Nat. Oust, et Ins., v. 14.
l„y,l,Ajh,COOl^lC
Preliminary Catalogue of British Cecidomyidae. 425
LiEBEL. 1889. Ent. Nachr. Berlin, v. 15.
Linn£. 1758. Syst. Nat., ed. 10.
1767. Ibidem (ed. 12), v. 2.
LoEW, F. 1873. Verb. Zool.-bot. Ges. Wien, v. 23.
1874. llyidem, v. 24.
■ 1875. „ V. 25.
1877. „ V. 27.
1878. „ V. 28.
1880. „ V. 30.
1885(-1). Beri. Ent. Zeitschr., v. 29.
„ -2. Verb. Zool.-bot. Gea. Wien, v. 35.
1888. Ibidem, v. 38.
1889. „ V. 39.
LoEw, H. 1844. Stett. Ent. Zeit.. v. 5.
1850. Dipt. Beitt., v. 4. •
Macquakt. 1826. R«cueil. Soc. Agric. Lille.
Marchal. 1895. Bull. Soc. Ent. France, v. 64.
1896. Ibidem, v. 65.
Meade. 1886. Tbe Entomologist, v. 19.
1888. Ent. Mo. Mag., v. 25.
Meioen. 1803. Dliger's Mag. Ins., v. 2.
""■ Klasaif. Dipt., v. 1.
Syet. Bcschr., v. 1.
Ibidem, v. 6.
. Naturhiat. Tidsskr., Kopenh. (3), v. 3.
ien Ent. Zeit., v. 2.
1885. Verb. Zool.-bot. Ges. Wien, v. 35.
1886. Wien Ent. Zeit., v. 5.
1895. Ibidem, v. 14.
Ormerod. 1885. Eigbtb Rep. Injurious Insects.
Ferris. 1840. Ann. Soc. Ent. France, v. 9.
Reuter. 1895. Acta Soc. Fauna ct Fl. Fennica, v. 11,
No. 8.
Riley. 1886. Annual Rep, Dept. Agric. Washington,
1885.
RoNDANi. 1840. Mem. per Servire alia Ditterol. Itabana.
1846. Nuovi Ann. Sc. Nat. Bologna (2), v. 6.
1860. Atti Soc. Sc. Nat. Milano, v. 2.
1804.
1818.
1830.
Meinert
1864
Mil.
1883. W
BSAAMEN. 1889{-1)
Zeitschr. f. Naturwisa., v. 62.
„ -2.
Berl. Ent. Zeitschr., v. 33.
1890.
Verh. Naturh. vcr. Preus. RheinI
Bonn, V. 47.
1890-2.
Verh. Zool.-bot. Ges. Wien, 40.
1891.
Beri. Ent. Zeitschr., v. 36.
ll,y,l,/eJh,G00glc
426 Messrs. Bagnall and Harrison's British Ceddomytdae.
Rdbsaambn. 1892. Betl. Ent. Zeitschr., v. 37.
1892-2. Verh. Zool.-Bot. Ges. Wien, v. 42.
1893. Ent. Nachr. Berlin, v. 19.
i895(-l). Ibidem, v. 21.
„• -2. Wien Ent. Zeit., v. 14.
1899H)- Biol- Centralbl., v. 19.
„ -2. Wien Ent. Zeit., v. 18.
„ 1910. Zeitschi. Wias. Insektenbiol., v. 6.
1911. Ibidem, v.T.
1912. „ V. 8.
ScHiNER. 1854. Verh. Zool.-bot. Ges. Wien, v. 4.
SCHBANCK. 1803. Fauna Boica, v. 3.
ScHWAGRiCHEN. 1835. In Pfeii'B Krit. Bl. f. Forst.-v.
Jagdwiss., V. 9.
Stephens. 1829. Syst. Cat. Brit. Ins.
gwAKTON, E. W. 1912. British Plant Galls (Catalogue,
pp. 130-261).
„ 1916. The Journal of Botany, pp.
24-29.
Tavabes. 1906. Brot«ria, v. 3.
Trail. 1886. Scottish Nat. (2), v. 1.
Vallot. 1827. Act. Acad. Sc. Dijon.
1849. M6m. Acad. Sc. Dijon.
Verrall. 1875. Ent. Mo. Mag., v. 11.
Wachtl. 1884. Wien Ent. Zeit., v. 3,
1885. Ibidem, v. 4.
Wagner, B. 1871. Stett. Ent. Zeit., v. 33.
Westwood. 1847. The Gardener's Chronicle, No. 36.
WiNNERTZ. 1853. Linnaea Ent., Stettin, v. 8.
Zetterstedt. 1851. Dipt. Scandinav., v. 10.
1852. Ibidem, v. 11.
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
PROCEEDINGS
or THE
ENTOMOLOGICAL SOCIETY
LONDON
1917.
LONDON:
SOLD AT THE SOCIBTY'S ROOMS, 11, CHANDOS STREET
OAVBNDlt>H sqtJARB, W.,
AND BY LONGMANS, GREEN, AND CO.,
P&TBEtNOSTER ROW, K.C. ; AND HEW YORK.
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
PROCEEDINGS
OP TBI
ENTOMOLOGICAL SOCIETY
OF
LONDON
For the Year 1917.
Wednesday, February 7th, 1917.
Dr. C. J. Gahan, M.A., D.So., President, in the Chair,
Nomination of Vice-PreaidenU.
The pRBSiDEKT announced that he had nominated Dr.
T. A. Chafuan, Dr. Q. B. liONOSTAFP and the Honble.
N. Charles Bothschild as Vice-Presidents for the ensuing
Tear.
Death of a former Praident.
The President also announced the death of Mi. C. 0.
Watebhouse, a former President of the Society, and a vote
of condolence with his daughter was passed on the motion
of Mr. Champion, seconded by Mr. Bethune-Baker.
Ekaion of a Feiiow.
Mr. A. W. Rtuek Roberts, M.A., Rothamsted Agri-
cultoial Experiment Station, Harpenden, and The Common,
Windennere, was elected a Fellow of the Society.
New Bye-law read.
The new Bye-law pioposed by the Council was read for
the third time.
FSOC. ENT. BOC. LOND., L 1917 A
iM:,i,A J 1^, Google
Exkibitiotu.
Lepidofteka fboh Salonica. — Mr. A. H. Jones ex-
hibited on behalf of Capt&in E. F. Studd, R.F.A., a Fellow
of the Society, at present serving with the Btitisb Expe-
ditionary Force at Salonica, Tatioua Lepidoptera taken hy
him in 1916, in the neighbourhood of Salonica. He also
furnished notes relating to these captniea and other speciee
observed by him.
PapUio podaliriut, not uncommon on the mountaioB near
Gllvezne on June 1.
Papilio machaon, at Lembet, in April, apparently umilai
to British specimens; a fresh specimen taken on Sept. 17,
near Senikos.
Aporia araiaegi, common. May 16, Lembet.
Pierit brassicae, common (larger than English specimena);
P. rapae and P. napi, common, apparently typical ; P. da^idice,
occasionally met with in June, Akunbar, common from
Aug. 31 until Oct. 8.
Coliaa edma, quite common, still flying plentifully on
Nov. 19 ; one var. kdice seen at Lembet.
Gonepleryx rhamni, several males seen, one on Nov. 18.
Nordmannia (Tbeda) ilieii var. cem. May 30, near
Gilvezne.
Chryaophanut diapar var. mtilui, two i^ taken among
rushes in Langaza Lake, in about two feet of water. They
were the only ones seen, just at this one spot surrounded on
all sides by wat«r.
Rumicia phlaeai, seen occasionally; common in Oct., and
still seen up to Nov. 18.
Aricia medon {Polyommatus aatrarche), Polyommatut icarut,
PJe&etus argus, common, June 2, near Gfivezne.
Vanetsa io, Pyrameia atalanta (first seen April 14, last seen
Nov. 1), common.
Pyrameis cardvi, not uncommon, last seen Sept. 17.
Meliiaea didtpna var, ocoidenUtUs, one 3 June 23, Akunbar ;
M. trivia, two ,Js May 31, near Guvezne.
M. pftoe&e, common.
Brenlkis seiene and Iiioria lathonia, not scarce.
u„,,i,A J 1^, Google
( iii )
M^ntM^ laritsa, Hay 16 to June 3, Lembet ; not tin-
common.
Salynu lemeie commoD on mountains.
Pararge m^gaera, conimon &nd typical.
Bpineph^ jurtina, abundant on mountaina ; E. tiUumue,
not unconunon.
Coenonympha pamphihu, very abundant, still out on
Nov. 19.
Syrichthve malvae and Nitoniadss tages, common.
Augiades sylmnm, several
Proktparce convolvuU, taken in lines, Sept. 19.
Macrogtoisa aleSotarum, occasionally seen.
Setitta fflMomeRd, Lithosia complanula and Procris ttaHoet,
common.
Zygaena JUipetidulae and Z. punctum, locally conimon.
Noctua pleda, common.
Mania maura, in t«nt, Sept. 12.
Amphipyra livida, one specimen, Lembet, June 26.
Plvsia gamma, came to lamp in tent, Nov. 14.
Acontia luctuoaa, Ardia vilUca and Coseinia slriala (gram-
mica), common, locally, May 29, near Guvezne.
Zeuzera pyrina (aescndi), one iS-
AspikUes ochftana, common.
Addalia omata, occasionally seen.
Ematurga aiomaria 3, locally; the yellow coloration of
Wicken specimens.
Co&era pusaria aad PeOonia vAicaria, several; Abraxai
gnttulariata, one specimen only.
Anaitit j^agiata and Cidaria fivctuaria, common.
Camptogramma biUneaia, common.
Commander Walkek said that many years ago he had
taken almost all the species exhibited in the nei^bouihood
of Port Baklar, at the head of the Gulf of Xeros, near the
Boulair Lines. He had found the larvae of M. trivia feeding
on Verbascum in the greatest abundance. The only species
he had not met with there was C. dispar; he remarked
on the very small size of the specimens of this species ; those
of M. larissa were, on the other hand, exceptionally large.
The Pbesident and Hr. W. Q. Sheldon commented on
l„y,l,Ajh,COOl^lC
( i» )
the abnndance of butterflies in Macedonia, the Utter ob-
serving also that North Macedonia and Albania were among
the least known of European localitiw for Lepidoptent.
Local forms op Aobias CLAin>tA. — Mr. G. Tai.bot ex-
hibited on behalf of Mr. J. J. Joicey a series of Agriat dauiia,
Schulz, showing its distribution and local forms. These in-
clude the forms tahlkei, Honr., daudia, Schulz, and amaamica,
Stgr., all from St. Jean de Maioni, French Guiana. A. tiaudia
is the Surinam race, and amazonica an Amazon race, which
are thus shown to occur in French Guiana as aberrations,
the form sahikd representing the Guiana race.
A transitional series may be shown to connect the various
races.
The distribution of daudia follows the coast, riTer-vallejm
and foot-hills. It ranges from Dutch Guiana in the north to
South Brazil in the south. It extends eastwards to Ecuador,
Peru and BoUvia.
Some BsmsH Rhopaloceba. — Dr. £. A. Cockayme ex-
hibited : —
(1) A series of Pararge egeria, bred Nov. and Dec I9I6
and Jan. 1917 from ova laid by several females taken in
August, at Limber, N. Lincolnshire. They showed consider-
able variation. Most of them had very large yellow spots.
One female had a great extension of the yellow colour and
thin scaling over the central portions of all four wings. One
nnderaide aberration was very dark with a deep brown hand
OD the hind-wing.
(2) An aberration of Polyg<m\a c-a/&um taken by Loid
Oanick in Sept. 1916 in Montgomeryshire, the tund-wing^
being nearly black and the fore-wings with costal spots
united into a crescent.
(3) A Qynandromorph of Poh/ommatus tcartM, a dark
female except for stripe and blue scales, with androconia
from the base to the termen of the left fore-wing; taken at
Boyston. Aug. 1916.
(4) A Gynandromorph of P. icarus with a stripe of bright
blue scales on the right fore-wing; no androconia; taken at
Folkestone, June 1915.
(6) A female Agriadeg coridon with one hind-wing marked
l„y,|,AJI^,COOl^lC
( ▼ )
with blue like ab. semUi/ngrapha, th« other hind-wing having
only a thin Bprinkling of blue scales over the same area;
taken at Royston, Aug. 1916.
FuBTBEB Note dealing with the question of the
SPECIFIC IDENTITY OF PeDICULUS CAflTIS AMD PbDICULUS
HUUANUS (VESTiMESTi).— Mr. Baoot redd the following note : —
" At a previous meeting (see p. xiv, 1916), when referring to
this question, I suggest«l that the disparity in the propor-
tions of the sexes in the F. 1 generation resulting from a cross
pairing between a P. capitis ^ and P. humanvs $ lent support
to the view that these insects were distinct species. In a
subsequent conversation the Rev. James Waterston challenged
this view, on the ground that it was necessary to prove that
normal sex- proportion resulted from pairings within the two
races, a point I. had taken for granted, owing to the parity of
the sexes in my stock boxes. 1 have now obtained details
of the sex -proportions of a number of broods of P. kumanvt,
which I propose to lay before you. Faired couples of P.
kumtmtu were taken from a stock box, segregated in small
boxes, and their offspring reared to maturity, with the following
result : —
So. ol F«i«. A A 9 ?
1
1- 2%
43- 98%
44 = 49%
46 = 61%
31 = 66%
17- 36%
m.
67 - 100%
119 - 73%
43- 27%
39 = 66%
20- 34%
116 - 68%
56 = 32%,
Nil.
46 - 100%
64 - 92%
6 = 8%
Na
49 = 100%
10 = 71%
4- 29%
Nil.
39 - 100%
Nil.
48 = 100%
7- 9%
74= 91%
42 = 68%
20= 32%
463 = 46%
676= 66%,
iM-,i,A J h, Google
( vi )
la a numeioas colony of a giegaiions inaect casnal Bex-
production on the part of any given pail will result in a nearly
equal proportion ot S 3 and $ $, and might serve to tninimiae
tlie dangers of too close inbreeding. In this series, however,
aex-production does not appear to be entirely casual, as there
is an evident bias on the $ $ side.
The sez-percentages of the F. 1 generation of four cross
pairings between <J P. cajiitU and ? P. htimanua are as
follows : —
No.1
71 =
= 74%
25 =
= 26%
No. 2
130 =
= 86%
22.
= 14%
No. 3
51.
- 61%
49 =
-49%
No. 4
76 =
328 =
-68%
- 71%
36 =
- 32%
ii^
= 29%
It will be seen that in this series the bias is heavily on
the S aide, which is possibly due to the effects of crossing.
I must admit, however, that in view of the emtio per-
centages from normal pairings of one of the parent races
my argument on tlie basis of abnormal sex- production on the
part of the hybrid insects can carry but little weight.
As experiment dealing with the egg-laying instincts of the
two species affords clearer evidence. The following method
was adopted. Glass-bottomed boxes of 1} inch diameter and
i an inch deep, were lined on the vertical sides with a strip
of flannel, such as is used for Army shirts, human hairs being
placed in central area so as to form a web, but not a felted
mass. The hairs came in contact with the glass bottom, the
flannel-lined sides and the gauze cover through which the
Uce were fed. An equal number of pairs of P. capitit and
P. humanm were placed in each box. At the end of a week
the ? ? of i", capUia had laid 80% of their eggs on the hair
and 20% on the flannel, while the P. humanw $ $ had laid
20% on the hair and 80% on the flannel As the insects had
been taken from congested stock boxes that had been in use
for many months, the P. capitis box containing a felted
mass of hairs and the P. humanut box a rolled flannel strip
l„y,|,AJh,COOl^lC
( vii )
which filled the box, it is poaaible that theit egg-Uying in-
stinots had been dulled daring captivity under these coa-
ditione. The eggs laid were separated into four lots: P.
Aumonuf laid on haii, P. kamanut laid on flannel, P. capitis
laid on hair, and P. capitis laid on flannel. Individuals were
reared from these eggs, and 12 ^ ^ and 12 $ $ of each lot
were placed in boxes containing human hairs and lined with
flannel, similarly to those used in the former experiment.
After five days, during which all four batches were fed and
exposed to the same conditions, the eggs were counted and
their positions recorded as follows : —
P. hvtnanus, bred from eggs laid on hairs : —
12 and 12
produce 359
on hairs 38 = 11%
on gauze cover of box ... 1
on flannel, side next box 37 = 10%
flannel, exposed side . . 283 = 79%
The eggs teere laid on hairs only where these came in contact
with fianttel.
P. hamanus, bred from eggs laid on flannel : —
12 and 12 / on gauze cover of box i= 1%
produce 34i j on flannel, side next box . . 66 = 16%
Q flannel, exposed side . . 285 = 83%
P, capitie, bred from eggs htid on hairs : —
12 and 12 r on gauze cover of box ... 1
produce 274 .| on flannel, exposed aide only . 14 = 5%
\ on hairs 269 = 95%
P. capitis, bred from eggs laid on flannel : —
-- , ._ f on ganze cover of box ... 4 = 1%
ji «/in I on flannel, exposed side only . 2= i%
produce 390 < l ■ . a i o lo/
' 1 on haira close to flannel 2 = t/a
•^' I on hairs 382 = 98%
Ahhoogh this result leaves the question of specific identity
still nndet«nnined, it suggests that there is a clear line of
demarcation as regards the egg-laying instinct, provided the
inaecta an left an element of choice.
u,y,l,/eJh,G00glc
( viii )
SPECIAL MEETING.
The Special Meeting summoned to consider the new Bye-
law proposed by the Council waa then held.
The Secretary read the proposed Bye-law, which runs as
follows :—
" Chap, xxiil. Prohibition in respect of FantU.
" The Society shall not and may sot niake any dividend,
gift, division or bonus in money unto or between any of its
members."
This Bye-law was needed to comply with the Act of Pariia-
ment regulating the Registration of Scientific Societies so that
they may he free from local rates.
On the motion of Mr. Beth one-Baker, seconded by Mr.
Stanley Edwards, it was passed without diacussioB.
Wednesday, March 7th, 19)7.
Dr. 0. J. Gahan, M.A., D.Sc., President, in the Chair.
Death of a MenAer of Coundi.
The death of Mr. A. E. Gibbs, a member of the Council,
and for five years a most valued member of the Busineu
Committee, was announced.
Exhibitions.
South Indian Heuipteba, &c,^Mr. E. A. Butler tx-
hibit«d two species of 8. Indian Hemiptera received from
Mr. T. V. Campbell, M.B., who captured them at Chikka-
ballapuia in the Mysore State; viz. : —
(1) Ureniius echinus, Dist., a species of Tifigididae, which
is a pest feeding on the leaves of the Egg-PUnt {Sdtanvm
melongemi), the larvae living on the under-side of the leaves,
and the adult insects on the upper.
(2) ApoUodolus praefectus. Diet., a species of CapsiJae,
which is found on the same plant, and the larvae of which
prey upon those of the Tingid bug.
n,g,t,7.dh,G00glc
( ix )
AIbo several recently described species of S. Indian Ftdgo-
ridae, together with the S of EurybrwAyt Uymentota, Fabt.,
which has only recently been iec<^niBed. In this sez the
green colour of the t^:niina has almost disappeared, the pale
fasciae are almost obliterated, and the wings are dark with
a white band on the apical area.
Notes ok Miucry in Oriental BUTTERruBS b7 Col.
T. JiBUTH. — Prof. PoDLTON said that he had recently received
the following interesting notes, and had tried to induce the
author to communicate them in person. CoL Jermyn, being
unfortunately prevented from coming, had forwarded with
his manuscript the illuattative specimens now exhibited to
the meeting.
" I have one or two examples of mimicry, which have
atrock me ior some time as rather specially interesting, in a
collection of Indian butterflies made daring the last twenty
years or so. I have long intended to write to you regarding
them, but going on duty again put them out of my mind.
Being at home again, and having just read yoor very interest-
ing address to the Linnean Society, I am now doing as I
intended.
" 1. Two teriet thowing steps in devdopmetU of diaposematic
or redprocai resemblance b&ween Papilio agestor. Gray, and
Danais tytia. Gray. — In the model Danais (Caduga) tytia,
Oray, there is a characteristic black marking nmning from
base of f.-w. along the coata across the end of the cell and
along vein 4 to the ma^;in. This is copied in typical P.
agettor by a stmiUr line, which, however, cute diagonally
acroea the cell at some distance before the end, and is, aa far
aa I have seen, constant. In the specimens of ageslor govindra,
Hoore, however, which I have from Mussoorie, this marking
grades from almost complete absence to a perfect copy as in
Epical agestor. This marking seems to me interesting as
being an attempt to obliterate part of t^ outline of the
large Papilionlne f.-w, cell, thus producing a superficial resem-
blance, in shape aa well as size, to the smallor Danaine cell.
It is also interesting that this mimetic feature is apparently
not yet entirely established, or possibly is undergoing reversion,
in the geographical race govindra.
u,y,l,/eJh,G00glc
( r )
" The specimens eent in iUustiatiioii are five govindra tnm
MnaBoorie (4fi00 ft., Apr. 22, 1907), one govindra from Moiree
(6000 ft., Apr. 29, 1898) and one typical agator from the
Ehuis Hills (1908). The fint six are arranged to show a
gradaal transition in the development of the diagonal mark-
ing across the f.-w. cell and in the filling up basall; of area 5
in the f.-w. with black, in mimicry of area 1 in the model
tyUa. The same series also shows the cutting o5 by a black
lunular line of a row of submarginai spots from the Papilionine
streaks in 1-w. areas 1-4 to match those in tytia.
" A series of the model D. tytia — one from Chakrata (Sept,
1893), four from Tehri Oarhwal, near Hussoorie (May 1-6, 1907)
— ^has been arranged to show the gradaal disappearance of
the black lines separating the disoal and postdiscal spots in
f.-w. areas 2 and 3, thus matching the streaks in oge^or.
The three characteristic spots in areas 2 and 3 of this Danaine
group (conf. meUmea, Cram., agUa, Cram., and mdanotdei,
Moore, sent herewith) and the Fapilionid streak hare thus by
reciprocal variation become a spot and a streak in the inter-
spaces of both species, the PapiUo producing a submarginai
spot out of the end of its streak, the Danait producing a
streak by the fusion of its discal and postdiscal spots.
" 3ome of the imperfections, due to difficulties of structure,
etc., seem almost more interesting than the perfections.
D. tytia, in reducing the black pigment, has lost the streaks
in the interspaces, conunon to both famihes, and fairly pro-
minent in agestor. The difierence in the arrangement of the
median and discoidal nervules seems to be a cause of cross-
purposes. Agetlor is filling f.-w. area 6 with black in imita-
tion of area 4 of tytia, while tytia seems to be clearing area i
in imitation of the same area in agator. On the hind-wing
the difEerence in size of the respective cells seems to he the
cause of a great deal of variation in the proportion of reddish
colouring. Papilios and Fiends seem to have considerable
difSculty in accurately copying the markings about the
S Danaine scent-glands; Nymphalines are much more
successful.
" Calinaga buddha, Moore, appears to be an outlying mem-
ber of the tytia-^igeitor combination. It has no red on the
l„y,l,Ajh,COOl^lC
{ = )
wings, bnt its red thorax is vary conspicaons, and in tho
only valley I have fonnd it (the only plsc«, I tiiitik, when it
has been seen in any nomben) it used to be out in the eariy
spring with agalor, and looked distinctly like it.
" 2. Some pointt in the mimetic retemtlance <4 ^ ^V^
}>&ilifle Parhettinajermyni, S. S. Drwx, to the Pierine Aporia
agathon. Gray, rate phryxe, Boitd. — One specimen of P. jermyni
was taken on Jane 19, 1907, in the Tons valley behind Chakrata,
beyond Hosaoorie, N.W. Himalayas, with a nomber of A.
phryxe, flying about a small detached clump of trees and
shrubs. Another (sent for exhibition) was taken on Jane 25,
1907, farther up the valley, flying by itself in bold circles of
half a mile or so, round the junction of a smaller stnam with
the Tons. It was obviously not A. phryxe, and was secured
on its third circle. One of the specimens of pkryxe shown was
taken with the first Parhestina on the 19th, the other in the
same neighbourhood later. The ParhetUtM forms an interest-
ing contraat with PapUio agettor, because in the Nymphaline
mimic the apparent size of the f.-w. cell has to be increased
and Dot reduced aa in the Swallowtail The bats across the
middle and end of the cell of P. fernmOit, Westw., have
disappeared entirely in jermyni, but there is a dustang of
black scales to indicate the end of a la^er celL It may be
inferred by comparing the sexes of penimiUt that this latter
marking, which has been regained or retained by jermyni, was
tiie first of the three L-w. bars to ondergo reduction. The
black dusting beyond the cell is undoubtedly useful in in-
creasing the leaemblance to the Aporia model, althou^ as
yet not nearly so dark. It b identical in both specimens,
and can be cleariy seeo in Plate XXIX, fig. 1, of Trans. Ent.
Soc., 1911, p. 187, where P. jermytti w described by Mr. H. H.
Druce.
" 3. V»e of proboscis of Parhettina perrimUit, Westa., in
mimiory of Aporia caphusa, Moore. — I have watched at
Hussoorie (4-6000 ft., 1906) one or two individuals of P.
persimilis drinking with a crowd of A. caphusa. The resem-
blance is distinctly enhanced by the yellow proboscis of
pertimilis, which harmonises with the yellow spot at the base
of the hind-wings in eaphuaa. This spot shows up much
ih, Google
( «i )
moie thui vonld be expected, when the wingi ue dosed ow
the back. The proboecis of penmQie is very blight yellow
in life, and when extended, and probably distended, in drink-
ing, is quite conspicuous. In tiie diied specimens the yellow
colour fades to a conuderable extent. There is no yellow on
the underside of persimiiit, of which D. limniace, Cism., is
the model (as ia rendered probable by the leeemblance to tha
pattern around the <3 scent-glands). In P. jemyni, which ia
evidently a direct mimic of phryxe, there b a good deal of
yellow on the h.-w. under surface. It is not 8uggeet«d that
the yellow cotoor of the proboscis has been evolved for the
purpose; all ParheBtinas have it in common with Hf^nii
and the yellow Ditipas. Its use in mimicking Aporia has
possibly preserved it. It does not occur in Euripua.
" i. Syiu^Koematic eutociationt of btue Euploem, Ac., tdbeM
on the same day. — I am sending you one or two sets of mim*iti«
speoiea caught on the same date, as I saw in your separata
you were collecting soch groups. All are common, but yoa
may not have the following five blue Euploeas taken together :
mulciber. Cram, Hvgii, Moore, hofei. Fold., Aomn, Feld.,
and Apfendetu, Buti. Muldber was in swarms, but it was
quite difBcolt to get the others, especially the last two, out
of the throng. I only got one Papdio leleanAvt, Hew. I am
sorry I haven't got Byblia UUhyia, Drury, to complete tix
other little associations of Tddiinia vtoloe, F., and SvUiaUa
nais, Forster, from Central and S. India. I have never {oimd
it common and have only two bad spedmens, both from
Sangor, Central Provinces."
ProL Poulton when exhibiting the specimens sent by Cd.
Jermyn remarked that the example of PapHio ageabjr govindra
from Munee was transitional between the five from Moasoorie
and the typical agator, not only in the development of the
diagonal f.-w. marking, but also in the amount of brown
jugment in the h.-w. and in the size of the specimen. It
would be interesting to compare a long series from these two
localities. The associated characters in which the Mussoorie
specimens differed from the typical agettor rendered it far
more probable that the butterflies from this locality wen
truly ancestral than that they had undergone levereion.
l„y,|,AJI^,COOl^lC
( xiii )
With regard to Calinaga buddha it wu very intoresting
that CoL Jenayn had noted in the field the reaemblance to
the tyHa-agalor combination. It could hardly be doubted
that the Calinagaa were themselves models for certain Chinese
and N. Indian Chalcosiine moths in which the reddish thorax
of the butterfly was represented by a patch of colour — orange-
brown in dried specimens — at the extreme base of the foie-
wingB. Thus Ckdura dejeani, Oberth., and Agahpe davidt,
Oberth., probably mimicked Calinaga davidia, Oberth.;
Cketura eronioides, Moore — Calina^ saka, Moore; and Ch.
banflava, Hoore — CtUinaga gaulama, Moore. Furthermore,
the patterns of these ChalcosiiDe moths were associated with
other species of the same subfamily, which seemed to bear
no direct mimetic relationship with the Calinagas. The
patterns of these anomalous butterflies appeared to belong
to a la^ combination of Lepldoptera, including Vanainae,
NymphaUnae, Pierinae, possibly one or more species of Par-
natsius, and day-flying moths. In addition to this relation-
ship, probably for the most part Miillerian, there was the
teaemblance to the lytia-agesior association, observed by
Col. Jermyn.
With regard to the combination of blue Euploeas, which
had been captured in the Naga Hills, Assam, on June 2, 3,
and 6, 1908, Prof. Poulton said he had long wanted to obtain
examples of the species which could be caught flying tt^ther
at the same time and place; also notes as to their relative
numbers. It was especially interesting that the central
dominant member of the combination exhibited to the meet-
ing should be the male of Buploea (TTepaiehToit) mtdc^xr, of
which the female was quite difierent in appearance, and
mimetic of a prevalent tjrpe of Danaine pattern (" Essays on
Evolution," Oxford, 1908, pp. 334, 335).
AJOIOPHILA SABUL03A, L., WITH ONLY TWO SUBMAKQINAL
CELLS TO BOTH FOBE-wiNQS. — A male A. sabtdosa with two
instead of three submarginal cells in each fore-wing was
exhibited to the meeting by Prof. Poulton. Hr. Charbonnier
had first called attention to this interesting variation in the
specimen, which had been captured by Col. Jermyn at Shap-
wiok Heath, on June 13, 1914. The variation was all the
iM-,i,A J h, Google
( liv )
more interesting becaose the presence of only two enbTnargin*!
cells was, as Prof. Fonltoo had learned from the Bev. V. D.
Morice, characteristic of a subgeons of AmmophUa, wfaidi,
however, did not include any British species.
The Pebsident stated that, at Prof. Ponltoa's request, he
had recently examined the specimen from the Burchell col-
lection (No. 1330), which was ahown that evening, and he
had no besitstioD in saying that it was ^ther a larva or female
of the group Phengodini, a group placed by some authors in
the Telephoridae and by otiiers in the I«mpyridae. The
specimen had been exhibited on a previous occadon by the
late Mr. G. 0. Waterhouse, who was somewhat doubtfully of
opinion that it was an Elat«rid larva (see Proc. Ent. Soc.
1904, p. Ixxxiv). Many years ago an insect supposed to be
the larva of a Pyropkonu was exhibited at a meeting of the
Society by the lat« Mr. Frederick Smith; and from the
description of its luminous properties given (Proc. Ent. Soc.
1869, p. xv) there can be no doubt that that also was the
female or larva of some species of Phengodini. The females
of this group are completely larviform, and have the same
short antennae, simple eyes, and short simple clawed legs that
are met with in tbe larvae. Both larvae and females may be
distinguished from Elaterid larvae by the fact that the tenth
abdominal segment is somewhat conical or tubular in form,
and piojectB beyond the ninth segment so aa to be visible
from above.
New Cbtoniids fbou Bkitish India. — Ur. 0. E. Jausoh
exhibited the four new species of Cetoniidae of the genera
Clerota, Pteudochalcothea, and AnaUma, described in the paper
subsequently read, and made some remarks on their characters.
Teupekatube at which Insect Life is desteotxd. —
Mr. A. Bacot desired to call attention to a very valuable
paper in the Memoirs of the Department of Agriculture in
India (Nov. 1916, Vol. IV, No. 6) dealing with tiie insects
attacking stored wheat in the Punjab, and the metliods of
combating them, by Barnes and Grove, and said : The anthois
show that the germinating powers of wheat submitted to a
tempeiatore of 70° C. for 24 hours were not noticeably im-
paired; as they found that heating to 100° C. generally
Ij.y.l.Ajl^yCOO'^lc
( XV )
destroyed the genninatiiig power, it is to be preanmed that
the conditions of t«Bt were not entirely favourable to the
grain, aa I am told that wheat dried and exposed to heat
under conditions of free transpiration of nioiatnre can with-
stand a temperatoie of 120" C. I am surprised, however,
that the authors do not refer to the beat necessary to destroy
the insect pests. They deal with a number of chemical and
some mechanical devices for destroying the insects, or freeing
the wheat from them. Cannot dry heat be used? So far
as my knowledge goes, the few insects which, have been ex-
perimented on die at temperatures of 45° to 62° C. (= 113"
to IZS-e" Fahr,). Is it Icnown whether the Coleopt«ra destruc-
tive of grain are specially adapted to heat, or is there some
other reason which precludes this apparently simple and
economical remedy b^ng used t
Mr, J. C. F. Fbyek said that he had worked out these tem-
peratures pretty thoroughly, and found that 130°-140° Fahr.
was always fatal to insects; t^eir power of leaistance was,
however, very varied, and moist temperature proved less fatal
than dry.
The Prbstdemt observed that in some French books a
method was described for destroying grain weevils without
interfering with the germinating power of the wheat. Water-
house had experimented succeasfolly on cotton.
Ur. Main pointed out that if the wheat was to be used for
food, its germinating power was of no moment.
Mr. WiLLouQBBY Ellis observed that 104° Fahr. was
always fatal to Staphylinid beetles.
Mr. DuBBANT suggested that heat was most fatal when
applied in shocks ; but Mr. Bacot said that he had not found
this to be the case with mosquito Urvae.
Dr. Chafiuh narrated an incident that occurred to him
many years ago, when, having immersed a number of beetles
in boiling water to kill them, a specimen of Carabus dathralvt
had recovered the next day, and eaten most of the other
insects.
Gift of a Microscope.
The Secretary said that Mr. E. E. Gbeen had ofieied to
the Society a valuable Binocular Microscope, for which objec-
iM:,i,A J 1^, Google
tdvea of 2' to )' were required, and asked whether any Pellow
had spaie objectives which he would preaent.
The following papers were read : —
" On new and little-known Lagriidae from S. America," hj
G. C. CHAMnoN. A.L.S., F.Z.S., F.E.8.
" Additions to the Knowledge of the Cetoniidae of Britiih
India," by 0. E. Janbon, F.E.S.
" The Condition of the Scales in leaden Males of Agrioia
thtiis and other Lycaenids," by E. A. Cockatub, M.A., H.D.,
F.E.S.
" Some Notes on Butterfly Migrations in British Guiana."
by C. B. Williams, M.A., F.E.S.
Commenting on Dr. Cockayne's paper Mr. BaTHCNS-BAKO
observed that the scales in these specimens were curved
triangularly and were very thin instead of fairly sohd. Mr.
Nbwhan said that such specimens needed to be set at once,
if killed with cyanide and relaxed they began to stain in \»»
than six hours, and that this was the case even if kept in ■
dry cyanide bottle.
Prof. PotJLTOM, Rev. G. Whbblbr, Coram. Waiob airf
Mr. Bethu»b-Bakbb commented on Mr. Williams' paper.
Wednesday, March 21st, 1917.
Dr. C. J. Oahah, M.A., D.Sc., President, in the Chair-
Etection of FtHoua.
Messrs. David Huktee, M.A., M.B., The Coppice, Notting-
ham; Nicholas J. Kubnbzov, The Imperi&l Academy o*
Sciences, Petrograd, and Percy A. U. Muscbakp, CkaiU''
house School, Godalmiog, Surrey, were elected Fellows of tW
Society.
ExhibUUma.
A SUPPOSED Hybeuj BcTTERPLY.^Dr. T. A, Chapi***'
exhibited a supposed hybrid between CaUophry* avU •""
l„y,|,AJI^,COOl^lC
( XV" )
CaOophryt rvbi, which for the moet part resembled C. rubi,
but the knobs of the anteiuiae had the red imderaide as in
C. avis, and the androconial brand was rather of the C. avis
form than that of C. rvbi, bat intermediate. The whit« eye
bonders were a Utile leas brilliant than in C. rubi, and the
white line on the nnderude, though distincUy that of C. rvbi,
was slightly nearer die base than usual in that species. The
specimen was taken where the species flew together at Amelie-
les-Bains, in April 1909. The specimen is now in the Museum
at S. Kensington.
A NEW Bbitisb Elatek. — Mr. Donisthokpb exhibited two
specimens of an Eiater, from Ireland, not in the British list.
One was taken hj Mr. F. Bouskell and the other by himself
in Co. Kerry, in June 1902. They had been recorded as
Slater pomonae, Steph., but neither of the captors had ever
been satisfied that they were that species. Mr. Donisthorpe
suggested that they were either E. praeustus, F., or a new
species. In the general collection at the British Museum were
similar specimens mixed in the series of E. •pramaUu, the
latter uther being a very variable species, or an undescribed
species was mixed with it in the National Collection.
He also exhibited a specimen of B. ■pomonae for comparison ;
this particular specimen b^g of interest because it has a
fungus parasitic on it.
Reappka&ance of Sunset Insects at Dawn. — Mr. Collin
said that he had observed that certain Diptera usually to be
seen about sunset were also on the wing about dawn, and
enquired whether the same fact had been observed in other
Orders. Dr. Cbafuan said that he was not aware that the
fact had been observed with regard to Lepidoptera, and
Mr. Green said that when he had been working light for
moths all night, he had never found the early evening moths
return in the morning; Mi. Buxton and Mr. liAiN, however,
both observed that Syria auroraria flies both at dusk and
The " Death-watch " Beetles. — ^The President asked
whether any Fellow could state from his personal knowlea^
that Anobium iomesftcum tape in the manner known as the
" death-watch." Xettcbium tesseUotum and Alropoi divina-
FROC. ENT. 80C. LOND., I. 1917 B
iM-,i,A J h, Google
toria both tap with the mandibles, and this was showii by
Derham to be a sexual call.
Wednesday, April 4th, 1917.
Dr. C. J. Gahan, M.A., D.Sc., President, in the Chair.
Election of Fdimes.
Mr. Thos. W. Kikkpatrick, The Deanery, Ely, and Sm
Charles I^nqham, Bart., Tempo Manor, Co. Fermanagh,
were elected Fellows of the Society.
BxhAUiona.
Forks of Fafilio priamus. — Mr. G. Talbot exhibited on
behalf of Mr. J. J. JoiCEV specimens of PapUvo (Troidxt)
priamus i. coelestis. Roths., from Bossel Island and St. Aignan,
and the allied race urviUeana, Gu4r., from New Ireland and
the Solomons. The specimen shown of coelestit from St.
Aignan was very close to urvilleana. A <? coekstU from
Bossel exhibited on the hind-wing the costal and discal yellow
spots of the type-form, which reappear as an aberration in
most priamus forms.
Ova of Stegomyia fasciata. — Mr. A. Bacot exhibittd
masses of eggs of SUgomyia fatdala, the " yellow-fever
mosquito."
LivixG " Death-watch " Beetle. — The President ex-
hibited a live specimen of XeaUHnum UattUatum, and demon-
strated iU marked power of " ticking " in response to tapfaDK
on the table on which the box stood in which it was contained.
Dr. Chapman, Comm, Walker and Mr. Durrant discussed
the question as to whether Anc^um domeslicKm possessed tlie
same power.
Paper.
The following paper was read, illii8trat«d by the epidi*'
scope: —
" Revision of the Genua Tarwm," by G. T. BbtbO«-
Baker, F.L.S., F.Z.S.
n,g,t,7.dh,G00glc
Wednesday. May 2nd, 1017.
Dr. C. J. Gahas, M.A., D.Sc., Preaident. in the Chair.
Election of a Feiknc.
Mr. Abtbub D1CK8EE, 24, Lyfoid Road, Wandsworth
Common, S.W. 18, waa elected a Fellow of the Society.
Exkibitions.
A Centoxkd proh Madagascar.— Mr. 0. B. Janson
exhibited specimens of Euchroea coeUatis, Burm., a rare and
beautiful species of Celoniidae from Madagascar, and directed
attention to the remarkably brilliant pearly blue coloration
of the underside of the body.
Genitalia of certain Species or Caligo.— Mr. W, J.
Kayb exhibited two cases of Caligo species from the collec-
tion of Mr. J. J. JoiCEY as well as from his own collection,
together with a number of microscopical mountings of the
male genital organs (all prepared at the Hill Museum, where
Mr. Joicey gave every facility) to prove that the forms, or
hitherto supposed species, lUlas from Ecuador and prome-
theus from Colombia, were really forms of memTwn which
more or less typically ranges throughout Central America,
and. that tdamonius, Feld. {— paw, Rob.) and suzanna,
Stich., from Colombia, the former a mountain race, the latter
from low elevations or coastal regions; peieus, Stich., from
Venezuela, cachi from Costa Rica, menu from Chlriqui, semi-
caemiea from S.E. Peru, joata from Upper Amazons, insv-
ttmua from Trinidad, teucer from Guiana, phorkys from Bolivia,
are all really forma of the variable teucer. The point of
difierence in separating these two species lies in the long clasp
having a serrate edge up to the apex in memtum and the tip
tenninating in two short teeth ; while in all the forms of Uueer
the serrations stop before the tip, and at the extremity is
a bunch of long hair instead of a pair of teeth. Telamoniui
at first sight looks very unlike typical teucer from Guiana,
hut specimens from Venezuela are intermediate in colour
and those from Costa Rica are intermediate in size, while
u,y,i,Ajh,Cooglc
{ ^ )
even on the coast of British Guiana specimens occot with
an occasional inward sufiusion of yeUow over the wing. With
memnon the same transitions in blue sufiusion can be found
linking up prometheus through epimetheus to the type form.
Atiaa from Ecuador is only like an enlarged memnon.
The errors made by Herr Fruhstorfer badly need correct-
ing. Mews, Pnih., from Chiriqui, peleus from Venezuela,
telemoniva, Feld. {" pavo. Bob.), and probably jnvonides,
Fruh., from Eastern Colombia are all forms of leucer and
not of menmon. As \o pavonidee I have no specimen to
examine, but suspect it to be a form of leucer.
Then Herr Fruhstorfer makes prometheus, epimetheu$, aticu
and anaximandrtis all forms of another species, while from
the genitalia and the gradual transition of the forms to memnon
there can be no doobt that they are all forms of that species.
As tdamoniut was placed wrongly under memnon it becomes
necessary to find what the form of memnon is that inhabits
Colombia, and the answer without doubt is prometheus, with
its aberration epimetheus. Felder, Reise Novara, p. 455,
distinctly says that prometheits and epimetheus came in the
same collection from Bogota ; yet Fruhstorfer gives different
Bubspecific rank to the two, and even in the same paragraph
says that epimetheits comes from Eastern Colombia and the
Western Cordilleras 2000 m. ! It is, of course, possible that
epimOheus occurs as a race somewhere, but for the elucida-
tion of what species these Gaiigo represent Feldei's information
is illuminating. Anaximand/na has not been examined, as
no specimen ia available.
The undersides respectively of the (eucer forms and the
memntm forms agree pretty well all through, and an inspec-
tion of them alone might have saved a great deal of con-
fusion. Messrs. Godman and Salvia in the Biologia Centrab-
Americana, on PI. xiv, figured what they called Uilamoniut
alongside with memnon, recording both from Panama, so it
should have been clear that letamonttu could not be a sub-
species of memnon. The " felamtmtus " from Costa Rica we
call cachi.
The following statement sets forth the facts as now re-
vealed:—
Ij.y.l.Ajl^yCOO'^lL'
( Mi )
CaUgo metnnon memnon, Feld,
Central America, Mexico to Nicaragua.
Caligo memn<m aUcu, Fnih,
Ecuador.
Caligo memnon prometkeus, Koll., and v. epimetheut.
Colombia.
? Caiigo memnon anaxiTnandrut, Fruh.
W. Colombia.
Caiigo teaoer lewer, Linn.
Goiana.
Caiigo teucer pelevs, Stich.
Venezuela, Caracas, 3000 ft.
Merida, 5000 ft,
Caiigo teucer menes, Fruh.
Chiriqui.
Caiigo teucer cachi, J. and K.
Coata Rica.
Caiigo teucer dbidonus, Fruh.
Amazons, Obidos.
Caiigo teucer joasa, J. and K.
Amazons, Joas Rio.
Caiigo teucer semicaeruUa, J. and E.
8.E. Peru.
Caiigo teucer ecuadora, J. and E.
Ecuador.
Caiigo teucer teUimoniue, Feld.
Colombia, Cauca Val.
Caiigo teucer suzanna, Deyr.
Colombia coast.
Pbotogeaphs of thb Ovipositoes of three SnticiDS
occuBKiNO IN Britain. — The Rev. F. D. Morice exhibited
a set of six photos showing the ovipositor and apex of the
$ abdomen in three species or subspecies of the Siricid genus
PauruTua, viz. jttvencua, ¥., nodUio, F., and cyaneue, F. Of
these the two first are Falaearctic forms, whereas cyaneus
belongs properly to the North American fauna. Yet it seems
to occur not unfrequently in these islands i and the exhibitor
n,g,i,.ejh,G00glc
( »ii )
had ia fact seen mauy more specimens of it taken in Britain
than of juvencus, though it is probably by no means bo com-
mon as noctilio, and it is at least possible that in all cases, as
certainly in some, its occurrence is simply due to the impor-
tation of American timber containing eggs or larvae before it
vas shipped.
The 9 of cyaneus is easily separated both from juvencw
and noctilio by the great length of its ovipositor, of which
about a half projects beyond the dorsal apex of the abdo-
men, so that in this respect it rather resembles our common
black and yellow Sirex, the well-known ffigas, L. It difiers
also from juvencus and agrees with noctilio in having entirely
black antennae, these in both sexes of juvencus being testa-
ceous at the base. The S is diBtinguishable from that of
noctilio by the colour of the abdomen, being testaceous prac-
tically up to the apex, and in this character it agrees with
juvencus (J, nor could Mr. Morice at present suggest any
satisfactory way of separating it from the latter, though the
9 S of the two forms differ so much that he could not believe
them to be mere varieties of one species.
He also mentioned that on Sunday last (April 29th) he had
been surprised by the premature emergence in one of his breed-
ing-cages of a $ Pkymatocera aterrima, which on the same day
laid seventeen eggs in a stem of its proper plant (Solomon's Seal).
He procured this stem by a stroke of good luck from a plant
which had been forced in a greenhouse ; his own plants, growing
in the open air, were as yet hardly out of the ground at all.
He had watched the whole process of oviposition very
carefully through a powerful lens, being desirous of correcting
or conhrming certain details of the account which he gave of
the oviposition of this species in Proc, Ent, Soc. Lond. for
1911, as to some of which Dr. Chapman's later observations
(Trans. Ent. Soc. 1914, pp. 173-184, and Ent. Rec. 1915,
pp. 145-119) were not exactly in accordance with his own.
He had stated that between the formation of each pocket
and the laying of an egg in it the terebra was lifted idtogether
out of the stem and afterwards introduced into it afresh. In
the present case, as in those described by Dr. Chapman, this
had not happened. The apex of the terebra never actually
l„y,l,Ajh,COOl^lC
Proc. Enl. Soc. Land., 1917.
F D Morice. Photo. Hal/-Tane H>ig. Co. Ll,l
Apex ok Female Abdomkn in Sikkx (Faururus).
1. P.juvencus l... dorsal view, , Ai. P.cyam-i.s I-., lateral view,
la do. laleral ,, J, P. iwclilio F.. dorsal .^,[c
2. P.cyaneusV.. dorsal ,. | Ja. do. laieral'..
n,g,t,7.dh,G00glc
( xxiii )
left the slit 1 Either, therefore, the exhibitor had been mis-
taken in his former observations (though they were made so
carefully and repeated so often that he hesitated to believe
this), 01 possibly the phenomena may leally be different in
diSeient cases, as for instance when the stems operated upon
diSer in thickness, and therefore in the curvature of that part
of their surfaces which is affected.
It has sometimes been stated that the attachments of the
two so-called "saws" are such that when one of them ad-
vances the other must necessarily retreat. Frequent dis-
sections of these organs had long made him feel sure that this
was a mistake, and he could now say quite positively that it
is so. On more than one occasion he distinctly saw one of
the two blades advance and retire and again advance, while
the other blade remained absolutely at rest with its apex
touching the apex of the (motionless) supports.
He noticed also, on one occasion, that the cuticle became
separated from the underlying substance of the stem along
a line which the advancing edge of the saw had not yet reached !
This had con6rmed him in the belief he had always held,
that the action of the " tcrebra " ia not simply cutting, but
includes a sort of splitting — in short, that it acts as a " wedge,"
and that in this part of its work the stout thick " supports "
take at least as much share as the delicate and slender " saws."
Dr. Chapman and Measrs. E. E. Green and Betbune-
Baker commented. The President and Mr. C. J. F. Fryer
both observed that they bad received larvae of Siricea from
gromng trees in England.
Further Note on the "Death-watch" Beetle. —The
President remarked that the XestMutrt which he had
exhibited at the previous meeting was still living, and that
he had discovered that it was a $. It had tapped when
touched on the head with a bit of paper, and when this was
continued had extruded its ovipositor.
Many different statements had been made by different
observers as to the part of the head used in tapping; these
were probably all correct, as the part of the head employed
depends on the form of the surface tapped.
Mr. DoNiSTHORPE commented on these observations.
u,y,l,7rJM,G00glc
( xiiv )
Papert.
The following papers were read :—
" New aod Little-known Heterocera from Madagascar,"
by Sir George Eeneick, Bart., F.E.S.
" A preliminary Catalogue of British Cecidomyulae, with
special reference to the Northern Gall-flies," by R. 8. Baqnau,
P.E.S., and J. H. Hakbisok, M.Sc.
Wednesday, June 6th, 1917.
Dr. C. J. Gahan, M.A., D.Sc., President, in the Chair.
Election of FeUom.
Dr. H. G. Breijer, Ph.D., Director of the Transvaal
Museum, Pretoria, Transvaal, 8. Africa, and Dr. Alfkid
£. Cameron, M.A., D.Sc., The Entomological Laboratoiy^
Agaaeiz, British Columbia, were elected Fellows of the Society-
ExkUntums.
Two New British Coccids. — Mr. E. E. Obbbn exhibited
two new and (at present) nodescribed species of British
Cpccidae, both belonging to the genus Lecaniitm and both
occurring on the Birch {Beluia oSm).
One of them is quite a large and conspicuous insect A
single example of this species was taken, at Camberley, hut
year, when a diligent search failed to produce any further
specimens. This year, however, the insect baa turned up in
comparative abundance, upon the stems and branches of
young birch saplings. Its presence is rendered still more
obvious by the attentions of a large ant [Formica rufa). In
life, the dorsum of the insect is more or lees covered with
greyish-white cobwebby secretion which gives it a " mil-
dewed " appearance. This secretion has been removed (by
ether) from one of the examples exhibited, revealing the
actual colour of the insect, which is seen to be bright cas-
taneous, with a narrow black median longitudinal stripe and
six or seven irregularly mottled black transverse bands. An
example of the male puparium is also shown.
l„y,l,Ajh,COOl^lC
(
)
The second epecies is smaller, but very Btrongly convex —
almost globose. Fresh examples exhibit ivory-white trans-
Terse bands on a dark brown background : but this pattern
rapidly fades after tbe death of the insect. The colour pattern
is somewhat similar to that of immature examples of L,
eajrreae; but the insect now under consideration is fully
mature, the scales covering masses of eggs. It is, moreover,
considerably smaller than capreae. This species appears to
be scarce. Repeated search ha^ resulted in the discovery of
four isolated individuals only.
Morpho adonu.
Morjtho eugaiia.
MOBPHO AD0NI3 AND M. EUGENIA DISTISCT SPECIES. —
Hr. W. J. Kate exhibited Morpho adonii, three males and
a fine female from British Guiana, also an behalf of Mr. J. J.
JoiCEY M. adonis males and one $ from French Guiana,
and M. eugenia males and one female also from French Guiana,
together with preparations of the genitalia of both to show
that there was no room for doubt that M. eugenia, Deyr.,
1860, is a distinct species from M. adonis. Cram. He said,
" The harpe of adonis is stouter than that of M. eugenia and
is widened at the centre into a sharp toothed projection which
is absent in eugenia ; the harpe of adonis is also much straighter
than M. eugenia, the latter being more curved. A glance at
the figures will reveal the diSerences at once.
On June 7th of last year (1916) when Mi. G. Talbot ex-
iM-,i,A J h, Google
( xrvi )
hibtted Morpko etigenia under the name of M. marcus, I
Bn^^ted that M. eugenia might possibly be the wet-season
form of M. adonis. In the light of evidence of the genitalia
this theory is untenable, as it is most unlikely that such great
differences in the barpes could be accounted for seasonally.
On the advice of Lord Rothschild I have also prepared the
genitalia of Araschnia Uvana and its summer generation prona
to test if there was any seasonal variation in these oi^ns, but
none was apparent. The settling of the specific distinctness
has automatically removed a great deal of the nomenclatorial
muddle. In British Guiana no male of M. eugenia has ever
been recorded with certainty. But on the Potaro Rivet
M. adonis la fairly common, and the two females that were
sent from there together with t«n males by Mr. Roberts
between the years 1902-1908 unquestionably are paired in-
sects of the same species — adonis. Of the small number of
adonis females that are available for examination it appears
to be clear that the fore-wing apex is rather pointed and the
anal angled projection rather pronounced. On this character
it is pretty certain that Schaller's figure of PapUio marcxis in
Naturforscher, vol. 21, p. 174, pi. iv, ft. 1, 2 (1785) is the ? of
Morpho adonis. Distant recognised this in 1881 in the Trans.
Ent. Soc. for that year, p. 397. The figure which he gives,
PI. XX, really looks as if it might be a female of M. eugenia,
and until a <J eugenia has been taken in British Guiana there
must be a doubt if Distant's figure is a $ of adonis or eugenia.
It certainly looks like a ? eugenia from its very large size and
blunt fore-wing apex.
The synonymy of Morpko adonis is now as follows : —
3. Papilio adonis. Cram., Pap. Ex., i, t. 61, A.B. (1779).
Morpho adonis, Godt., Enc. M^th., ix, p. 439, n. 3 (1823).
?. Papiiio Tnarcus, Schall., Naturf., xxi, p. 174, n. 4,
t. 4, f. 1, 2 (1785).
Zeuxidia tnarcus, Kirby, Syn. Cat. Dium. Lep., p. 116,
8 (1871).
S ?. Morpko adonis, Fruh, in Seitz, vol. v, p. 349 (1913)
(= eugenia, Deyr. (J) err. det.
Morpho adonis. Roths., Nov. Zool. xsiii, p. 317, 191$.
l„y,l,Ajh,COOl^lC
{ xxvii )
Lord Rothschild, with whom I have been in correspondence
for some time over this matter, agrees that he was in eiror
in retaining the name marcus for the pale broader- winged
Urger speciea that has recently been discovered in some
numbers in French Guiana. The geographical races of adoni*
do not appear to be sufficiently well known, but the race named
by Mi. Lathy as major from N. Peru also has a $ with pointed
wings, although the insect is very large. From Iquitos there
appears to be an intermediate race both in size and in the costal
white spot which is smaller than in major, but larger than in
adonis adonis. This form might be known as intermedia.
Morpho eugeriia, Deyr., which until quite recently was so
exceedingly rare and of which we were in so much doubt as
to its specific distinctness, is now established as a perfectly
good species. Apart from the differences of the genitalia
the (J <J are easily separable by the more delicate silvery
blue and the squarer outline of the wings. The $ $ such as
are known are also of squarer build with the apex more blunt
than in adonis, but otherwise the colouring seems to be the
same. It is possible that the dark ground-colour of the
hind-wing extends always well beyond the cell, while in
adonis it is only up to or just beyond the cell. In a photo-
graph of eight $ 2 Morphos in the collection of Mdlle. de
Plorrack of Paris the upper four«ppear to be adonis by the
sharp fore-wing apices and smaller size, and the lower four
eagenia by the blunter fore-wing and larger size. The
synonymy of M. eugenia is as ftJlows :—
J. Morpho eugenia, Deyr., Ann. Soc. Ent. Traver, Ser.
iii, vol. 8, p. 209 (1860). (Oyapock, French Guiana.)
?. Morpho eugettia. Bar., Ann. Soc. Ent. Traver, Ser. iv,
vol. 4, p. 32 (1864). (Portal, Maroni River.)
9. Morpho eagenia, Oberth., Btud. Ent. Jjiv. vi, p. 27,
pi. vi, fig. 1 (1881).
?$. Morpho adonis, Dist., Trans. Ent. Soc, pi. xx, figs. 1,
2, p. 397-399.
Morpho eugenia, Kirby, Syn. Cat. Diur. Lep., p. 122 (187 1 ).
Morpho marcus marciu, Goths., Nov. Zool. xxiii, p. 317
(1916).
u,y,i,Ajh,Cooglc
( xxTJii )
The only race that we know of lot certain outside the type
form is Morpho eugenia vraneis, Bat«e, from the Peruvian
Amazons.
In the Novitatee Zoologicae, vol. xxiii, p. 317 (1916), Lord
Bothachild correctly surmised the specific distinctneaa ol
eugeaia and adonis with the placing of the races of each, but
called eugenia " morcus," thinking the two names applied to
the same insect. Mr. Lathy has obtained from the coUecton
in French Guiana the following interesting notes on eugmia
and adonis. " They are both taken at the same time, though
M. eugenia disappeara at times, while M. adonis appears to
be found all the year round. They are also diSerent in habits,
as M. eugenia is only to be captured at daybreak, and after
about a quarter of an hour it Siea too high to be taken."
I should like here to acknowledge the generous help of
Mr. Joicey, who put the whole of hia specimens at my dis-
posal for examination and dissection notwithstanding the
great rarity of the specimens; also to Lord Rothschild, for
help by correspondence, which has contributed to thresh oat
this difficult problem,
[NOTS ON MOEIPHO BUOENU, DbYR., BT LoBD RoTHSCHILD.
F.E.S.— When I stated, Novit. Zool., vol xxiii, p. 317, that
the Morpho described by Deyiolle in 1860 must stand as
morctM, Schaller, 1786, I gave as my reason that we only
knew a brown and white-banded ? taken in copulation witii
an undoubted ,S of Deyrolle's eugenia. Bates has also stated
that he saw $ adonis flying, and that they were lavender
grey. I now, however, on further study of material not
available at the time I wrote, agree with Mr. Kaye that the
$ $ both of adonis and eu^enta are brown and white-banded.
Seeing also that the adonis $ $ have pointed apices to the
fore-wings, and Schaller's picture shows this clearly, I think
his name is a pure synonym of adonis. Cram., and the rounded
winged silvery blue insect must stand as Morpho eugenia,
Deyr.l
RESEMBI.A24CE, MlUETIC AND NON-tOUETIC. — Mr. 0-
Talbot, on behalf of Mr. J. J. Joicey, exhibited : —
1. A white-banded mimetic group of African Heterocera
from the Cameroons, composed of Mastaga mariiona, ButL,
ih, Google
( «il )
aod Mtusaga sp. ? (Agariilidae), Bypochrosis massagaria,
Karsclt (Geometridae), and OmmoCtxAelx^ grandit, Drutw
{7*netd<M).
2. An example of lesemblance which is not mimeticr seen
in SeorioptU tn/umata, Wair., from Peru, a Geometrid
bearing a strong likeness to a species of LymantrHdae from
Angola.
3. A mimetic group from Dutch New Quinea. — Species of
TeUervo, which are essentially Danaine in character, probably
serving as models for the <; of the Satj^rine, DnuiUopsis
doherlyi, Ob., and with which is associated the black and
white Neplif. Another resemblance appears to exist between
the $ of D. dohertyi and species of Byantis and Taenaru
belonging to the Amaihuaiidae.
4. Two forms of TeUeno from the Island of Misol. This
is further confirmatory evidence of the existence of at least
two species in this puzzling genus. Fruhstorfer and Van
Eecke have declared their opinions that only one species
exists, but Lord Bothschild has noticed the occurrence of
two forms from one locality in the Snow Mountains, Dutch
New Guinea, and Mr. Joicey has received two forms from Biak
Island.
6. PapUio erlaces, with its races, including a new race
from North Peru, and showing the mimetic 9 of P. harmoditu,
Doubl., from the same district.
BlBDS CAPTtnttNO BUTTERFLIES ON THE WINO AT OZFORD. —
Prof. PouLTON said that on the previous day (June 5) Mr.
H. Britten had seen a swallow capture a Lycaenid butterfly.
It was rather a curious coincidence that the observation was
made from a windiTw in the Hope Department, where so
much attention had been devoted to this subject. The
dodging flight of the butterfly made its capture difficult even
for such a bird as the swallow, which only succeeded at the
third attempt. The Lycaenid was bright blue and flying
high opposite the window, in the afternoon. The bird was a
male.
Prof. Foulton said that, after the above not« had been
written, as he was bicycUng to the Museum that morning,
a bird darted from out a garden and struck a Pierine, flying
u,y,l,/eJh,G00glc
( "« )
hekTil; after r&in, close by faim— so close in fact that the
bird waa frightened by the bicycle, and darted back, leaving
the butterfly fluttering in the road. He got off his bicycle
and, looking back, saw the bird return and carry the inaect
into the garden. When he tried to come to closer quarters
the bird flew up into a tree and finally, still carrying the
butterfly, across the road. The butterfly was P. ttapi or
rapae, almost certainly the latter. The bird could not be
obnerved very clearly, but from its size, colour, markinga
(so far as seen), and flight, was evidently a female chaffincJi.
The persistent pursuit in both these observations was worthy
of note.*
Forms or Pafilio polytes rouulcs, Crah., from StNOX-
FORE TSLA^'D AND THE HAIN'LAN'D OPPOSITE. — Prof. PODLTON
exhibited the mimetic polyUt, L., females of two series receotly
sent to him by Dr. R. Hamt«ch of the Rafiles Museum,
Singapore, in extensioD of his eariier consignment, captured
JaQ.-May, 1916, and analysed in our Proceedings for 1916, pp.
btxvi-liiviii. The laiger of the two series, analysed on p.
xxxi, was from the island.
The non-mimetic females, <^ru«, were thus leas abundant
than in the series captured Jan, -May, 1916, when they
approximately equalled (6 to 9) the mimetic females.
The mainland series, only received within the last few
weeks, consisted of 16 specimens from Johore, oppoaite
Singapore Island. Dr. Hanitsch wrote March 26, 1917 : " My
collector was over there last week, and this is the result of
five days' collecting ! As soon as I have the opportunity I
will try to send you more. Johore Bahru, where the speci-
mens were taken, is the capital of Johore, close to the straits
separating Singapore Island from the mainland. ' Bahm '
means ' new.' " The specimens were noted as having been
captured four miles from Johore Bahru on the following
dates :^
" A few (kya after the meeting on June 8 I reoeiTed the following
note from Mr. W. Feather, F.E.S., at Kibweii, B.E. Africa:— "On
September 26, 1916, about 7.30 a.m., 1 ww a gnen BM.e>ter
[probablj Meropt ap.] oUch and eat a Btlatoit mtiaUina i . The
insect wae at rest on a low bunh, and I had gone quite close t4 ~~ ~ ~
it and had disturbed it."— H^ B. P.
l„y,l,Ajh,COOl^lC
( xxxi )
MuTch 22. — 2 mimetic females (jmlylea).
„ 23. — 1 male-like female {cyrua), 4 males.
„ 24.-3 males.
„ 25. — 1 mimetic female (polyla), 3 males.
„ 26.-2 males.
The numbers were quite insufGcient to warrant any con-
clusion, but they suggested the possibility that the ct/rus
females were rarer on the mainland than on the island. Prof.
Poulton very much hoped that Dr. Hanitsch would be able
to send a series sufficiently long to teat this suggestion
thoroughly.
Papilio polyles Twnulus, Cr., at Singapore.
c«p«.r«ini»ie. j "^zT*'" ' *""^°X; *• I '"m^tS:*'
May 12
7 i ;
OoloW
1
Nov. 15
I'll
.. 16
3 : ; 1
„ n
2 ; ; 1
,. 22
1 2 1 11
,. 23
! ^ : >
. " 26
1 2 i 1 '
.. 2T
i 3 i 1 1 !•
Dec. 2
., 4
» !' I ■
.. 6
3 1 1
.. 6
1 3 1 2 1
Tot»U.
1 34 1 5 V
* The form ttieJtiuf with no white in the hind-wing cell.
u,y,l,7rJM,G00glc
Prof. Poulton said that variAtion in the submaigiiial red
spoto and in the white patch and the adjacent led markiiigB
on the hind-wing of the mimetic pt^yUs female made it a
-favoucable example for the study of the heieditAiy trans-
miaaion of very small characters and for testing whether
such transmission was Mendelian. With this object he had
analysed Dr. Hanitsch's material and the result showed
that there were numbers of small characters which might
be selected for breeding experiments.
Considering first the 9 females captured Oct. -Dec., 19IG,
there was great variation in the size of the white patch in
the hind-wing, but it was generally la^ and well developed
in all its parts, viz. the mark in the cell and those in the four
areas from 2 to 6. The white marking In area 2 was always
more oi less replaced by red and that in area 5 very variable
in size. The white marking in the cell was usually loi^, but
very small in the female captured Dec. 4, and absent from
that captured Nov. 27, so that this latter specimen belonged
to the form slichius, HQbn., the first that had appeared among
the specimens sent by Dr. Hanitsch from Singapore. The
female of Dec. 2 combined a large white marking in the cell
with a mere trace in area 5 and a small remnant near the cell
all the rest being replaced by red, in area 2. The white
marks in areas 3 and 4 of this specimen were of moderate
size, the outer end of the former being capped by a red mark-
ing half as long as the white. This red section did not here
appear to replace the white, but to be added to it, for there
was no gradual transition as in area 2, but both white and
red sections ended abruptly and were separated by a trace
of ground-colour, in the form of a thin curved black liiie>
The independent variation of the elements of the white patch
was very evident, especially that in the cell as compared
with those in areas 2-Q. Thus, when the females were arranged
according to the size of the mark in the cell, from ite largest
down to its disappearance in the stichius form, the sizes of
the marks in the areas outside the cell by no means followed
the same arrangement; for they remained of full size even in
the stichius form.
The combination of small submarginal red spots above
l„y,l,Ajh,COOl^lC
( TTTlii )
with large ones below, as noted in some of Dr. Hanitsch'B
Singapore specimens of January and February, 1916 (Proo.
Ent. Soc., 1916, p. Izzvti), was here seen in the ca[ltuieB ot
Nov. 25 and Nov, 27 (the ilickiua form), and to a less marked
extent in those o£ Nov. 17 and Nov. 22. The especially small
Rze of the subma^nal red spot in area 3 (Froc. Ent. 8oc.,
1914, p. zxv) was very evident in these four specimens. It
was altogether wanting from the upper surface of the sltc&tuc
form and represented by a mere trace in the specimen of
Nov. 26.
Comparing these nine polytes females with eight out of
the nine (one had been mislaid) taken JaQ.-May, 1916,
the latter series showed a somewhat higher average develop-
ment of the white markings on the hind-wing. The mark
in the cell was in no case reduced so far as in two of
tiie later captures, and t^ mark in area 2 was nearly re-
placed by red in only one specimen, also characterised by
a smallish white mark in area fi — well developed in the seven
others.
As regards the three polytea females from the mainland, one
of the two taken March 22, 1917, much resembled the island
one of Dec. 2, 1916, although as compared with the latter the
white mark in the cell was much smallet while the spot in
area 6, though very small, was larger and far more distinct.
These two specimens of Dec. 2 and March 22 might indicate
a transition towards the romulus $ form, mimicking P. hector,
L., a transition especially suggested by the increasing red and
diminishing white in area 2, the addition of red to area 3,
and in the tendency of the red to encircle the white patch
by invading the basal border of the mark in Uie cell. This
invasion was especially marked in the mainland female and
in the island one of Nov. 26, in both of which the red scales
were continued across the cell into area 6. The other main-
land female of March 22 showed a small spot in area 6, but
the white markings were otherwise biirly developed, oa they
were in the female of March 26, with a much larger spot in
area 6.
When Dr. Hanitsch's mimetic female polyUs were last
shown to the Society (Proc., 1916, p. btxvi) Mr. J. C. F. Fryer
raoo. BUT. aoo. lond., n-iv. 1917 o
iM-,i,A J h, Google
( zxziv )
&iid Hi. E. B. Gieen lenUTked oq the differeDce between tlie
innei maiginal bordei of theii hind-wing pattern and t^t
of Ceylon specimens. The lattei seemed to them to show a
Bmaller development of the red markings in this region.
Piof. Poulton had since carefully compared the specimenB
and found that the difieience in appeanmce was due to two
out of the three red markings in area Ic of the western females
being fused in the eastern females into a single long basally
placed streak, the outermost marking remaining separate
in both localities. The basal division into 2 separate mark-
ings was sometimes indicated by a notch which occasionally
(lor example in 2 Singapore females taken in January, 1916)
broke througH, producing a pattern like that found in the
great majority of the jxAylea females in Ceylon. Among
these latt«r too the same fusion into a single notched streak
occasionally took place. So tar as could be inferred from the
insufficient material in the Hope Department specimens from
the mainland of India were intermediate in this respect
between those from Ceylon and Singapore. It would be
very interesting to breed from the exceptional females in both
east and west.
Frbdacbous Reditvud buqs and Fossobs, with tbbib
PSBT, FROM TBE S. PaULO DI8TBICT OF SotlTH-ElAST BRAZIL.—
Prof. Poulton exhibited and described a set of predaceous
insects captured 1913-16 by Dr. Giegorlo Bondar in the S. Paulo
district of S.E. Brazil, and presented to the Hope Department
by Ur. Q. A. J. Rothney. In determining the species kind
help had been received from Mr. 0. J. Gahan, Hr. W. h.
Distant, Mr. E. Q. Blair, Mr. R. South, Mr. A. S. Hiret, aod
especially from Dr. G. A. K. Marshall and Mr. Rowland
E. Turner; in interpreting the often indistinctly written
Portuguese on some of the labels, kind help had been given
by Don Fernando de Arteaga. The great majority of the
observations had been made at Piracicaba, about 100 miW
N.W. of S. Paulo, in S.E. Brazil; others at Campinas, about
70 miles N.W. of the same city.
The Reduviid captors and tJieir prey— almost invariably
insects belonging to the specially protected groups — wen
shown in the following table: —
ll,y,l,/eJh,G00glc
triam o» uddtiid bdq.
■rmiM Of »n. tMtum did ntn.
Apiomtnu loMpa, F.
The FoBsor (Sand -
19U.
„ ..
The Honey.bw Jpt»
" 1B13-I9i6.
" ••
The Antbribid beette
«M, Oarm.
"
^£^p.- "•
Tb« C^thuid beetle
<ita,BLuidt.
"
2dt»sp.A.
The Hooey-bee Ajiit
„
.. B.
BroamBp.!.
1913,'bot.2or'ili.T.2
»h;s. • ■
The Homopteron
«nM,Dirt.
" 1913-1916
The FosaoTS and theii prey were recorded in the following
table, the notes being taken from the French and Foitugnese
of the labels on the specimens or from the common labels
written by the collector and pinned beneath two or mote
specimens. " Nid " had been taken to mean burrow, inas-
much as the species belonged to the groups which were well
known to dig in the ground. Dr. B. C. L. Perkins had further-
more kindly examined the specimens and was sure, from their
structure, that all except one were fossorial in habits. The
one for which this evidence was not clear was the Podium,
which on other grounds, viz. its prey, consisting of several
cockroaches, was also sure to be fossorial. When there is
no reference to the bunow it is to be inferred that the prey
was being carried or dragged by its captor. In most cases
it was stated on the label that the Fosaor had been captured
with its prey.
u„,,i,A J.., Google
VMua or rouoB.
BFKns o* rutT. uwiun Ut> MB.
(POMMLlDAa).
A Trap-dooi smder
(C(«..5W«), /Sop.
■p.
Firaoioaba, Deo. 23,1
1914.
Bp. A., muumed in
Brit. Hub., 8 ex-
•mpki.
ZycMo ip., 2 examples.
Jan. "27 and" Deo. 1
1911.
^^^srs.-™^-
Ua^ 26, 1914.
Sia. 27, i914.
A web-building spider
Jm»n«(£jwra)Bp.
d^. 6. 19is.
Podium 8p.ooptured
in it> burrow.
Walk., out of nvenl
BiatUdat of umB
species in the borrow.
Oct. B, 1914.
1
Note: "OissBe iarve
Another label adds
in Portuguese that
the larvae were
green. No speoi-
NodaU butcerUunlyj
in S. Paulo district, |
and betweoi 1913
and 1916.
L., o&ptured in iu
1 9 nr. genua BOero-
•nofiiur, out of
in the burrow.
Kracioaba, H6». 7,
1913.
2 f , 1 immature Lo-
eustids, Conoavhol-
oida sp., probably
»»mIb>L, F., i^
the burrow.
Jul. 13, 1914.
theotherl genus, in
bunow.
Jai'. 26, mo.
ito burrow.
2 Bvbria ip. ? out of
several Loautida* in
the burrow.
No looality, but
certainly 6. Paolo
district. probably
Piiadcab*. as Q.
Bondar was there
Jan. 13 and 27.
Jan. 23, 191i.
ih,Cooi^lc
( zzzrii )
™„.,,.^
SPlCna OF PRET.
U«^^D.T>.
SpAcc Onattu, Sm.
The Aohdian <gTaBs-
hopper) Ottnilia vio-
lacea, Tbimb., 9 .
In oampo, Campinas,
Deo. 7, 1913.
The Acridiftn 8AM0-
G., 9 . The prey ia
buried neat the place
where it was cap-
tured—one Aoridian
for each SjAa Iarv».
In the cemetery,
Compinafl.
Deo. 8, iei3.
aia,J.
Piraeicaba, Jan. 22,
1915.
BjAtx ramptuonu,
Coeta, captured in
is, l9o{ the Locus-
Ota, de G., in the
In the cemetory,
Campinas, Dec. 21,
1913.
C«reen..™tator,8m..
on ono states th»t
it wu oaptored in
its burrow where
it ooUecta nuny
8 of the Enmolpid
with one Crretrta
' 7 in uiother with »
BCoond Ceneria.
Campinas, Deo. 21,
Cm:m.«mpkar,Sm..
Mptnrod in bor-
rows " deep in the
math " (Portu.
guese).
EpUroffiu Bp., tdken
in one of the bur-
Iteo. 21, 1913.
Atlaia up., captured
An immature Penta-
tomid bug.
1916.
Nctagonia dteor<Ua,
Sm.
A 9 Aohetid of genus
Gr^us or OrySodes.
Piraoicaba, April IQIS.
2 i Acbetida as above,
one with each
fVwBor.
"
Tachsia caOota,
Kohl., 2 examples.
" They live in
««uly" pkooe on
one m PortngneBe.
Note: "Chasse des
petite mouohes " on
one Foe»>r. No
■peoimens of prey.
On the football
ground, Oftmpinaa.
'nusnotfl and date
on one specimen,
but probably ap-
plies to both, Deo.
19, 1913.
jTodjtosp.
A i Aoridiu) I gen. et
■P-
Campinas. Deo. 19.
1913.
ll,y,l,/eJh,G00glc
Mtmdttla [Btidia)
adonu, Huidl., 2
eumplee. "Cbawe
HJCToUpidopUres "
label.
The arrangement of No locality. ProbaUj
collection Hhonrd 3 | Piracicaba as Jul
Hetperidat (prob-
ably OorifyAum
' Prittw.
Siajihgliu epieaiU,
Hab., ?, aiajA^iu
»p. rfmnor,Schaus),
associated irilh one
Jf. adonit and 3
otben (probably Sla-
n&uIiMBp. ttcfiramut,
SonanB, r Megittiaa
■p., I SyaUuea sp.
9 ), with the other
— alnuMt oertainly
takoi from the
bairowB.
23, 1914,
odminon label (d.
Spter «(■■
M<nudtila ISlitiia)
gravida, Handl., 2
elamples. " Chasan
Dipt^^e " on com-
moD label.
4 VobtetOa cbaa, F..
prob«bly taken
from burrow of one
or both FosBora.
Firacioaba, Oct. 191t.
Mcnedrda (Stiaia)
fo data, oertaildy
1913-IOie, S. Fknlo
district.
OliT. Probably
from burrow.
- " Chasse larre de
punaises de boia."
Fmtato-
mida, almost cer-
tainly from burrow.
It was of great isterest to have the opportunity of studyiDK
a good seriea of S. American Fosaora and their prey, and to
recognise how constant were the instincta hitherto recorded
mainly from other parts of the world. Thus all the Pom^^iiSM
had taken spiders, all the species of S'phex Orthoptera, the
Cenxria a beetle, tUe Astata a bug, etc. The orthopterooa prey
of the Podium su^^ted the possibility that its true position lay
between Ammophila and Sphex. The most interesting record
was that of Monedula adonis, one of the Bembecides, with
Hesperid prey, not unnaturally classed as " Microlepido-
ptera" by the collector. The specimens were in very bid
condition and so coidd not be determined with certainty, but
there was no doubt about the fact that very difierent speciw
l„y,l,Ajh,COOl^lC
( xixix )
w«ie lepresented. Only three retained their bodies, and of
these Dr. Eltringham had made out two to be females and one
a male. Althongh so different, the general appearance of the
Bpeciee was similar, all being dark brown in colour with no
pattern or one so inconspicuous as to be invisible at a little
distance. It seemed probable that the Fossois had caught
their prey drinking at mnd and had selected females as far
as possible. Dr. Carpenter's observation, recorded on p. zlii,
supported tbeee suggestions.
Mr. G. A. J. Bothney had written Oct. 19, 1916 ;—
" The Foaaor that stores Skippers is new to me but I
have met cases of a first stage in that direction. Vexpa cinda,
F., the common Indian Hornet, captures Skippers on the wing,
strips o& the two wings on one side, then holda the other two and
squeezes the body dry, drops the body and two wings and flies
away. I have recorded the incident in the Entom. Mo. Mag.,
vol. xjii, 1876-7, pp. 254-5, under ' Squirrel versus Hornet.'
I expect FossoTs and Wasps have a taste for the fat juicy
bodies of Skippers, and as they fly by day they fall a natural
prey; but the storing is a distinct advance. The habit of
F. ctncfa is undoubtedly established, not a chance occurrence.
The favourUe Skipper was a thick-set fleshy one — dull in
colour like our tages — ^but with long, pointed wings. There
was a prompt, business-like action about the operation which
told it was an old game or, aa they would say in India,
Shikir."
The only previous observation of the storing of butterflies
by wasps was, so far as Prof. Foulton knew, that recorded
by Belt in " The Naturalist in Nicaragua " (2nd edition,
1888. p. 109) :—
" There is ... a yellow and black banded wasp that
catches them [' Helvxmi4ae,' evidently referring to the trans-
parent-winged Itkomiinae] to store his nest with ; and when-
ever one of theee came about, they would rise fluttering in
the air, where they were safe, as I never saw the wasp attack
them on the wing. It would hawk round the groups of
shrubs, trying to pounce on one unawares; but their natural
dread of this foe made it raUter difficult to do so. When it
did catch one, it would quietly bite ofi its wings, roll it up
ih, Google
( ll )
into a ball, and fly off with it." The deacriptioD snggeBtod
a DijdopteToua tather than a Fossorial wasp.
Id addition to the Bpecimena here tabulated Dr. Bondai'a
collection exhibited to the meeting contained the tnbe of a
trap-door spidei together with the probable owner — a spider
of the genus Idiops without label, but placed in cloee proximity
to the tube. To the latter was attached a Portuguese label
of which the translation was " Destroyed by wasp. Seems
to be a SaHvt with yellow-tipped antennae, of medium size."
The date was Jan. 15, the year being left uncertain. It was
probable that the wasp referred to was the Peptis, taken
with a spider of the genus Idiopt, Dec. 23, 1914, and standing
first in the table on p. xxxvi. This Fossor was of moderate
size and had yellow-tipped antennae. It was also probable
that the spider had been attacked and stored in its own tube
by the Foasor, which was thus saved the trouble of digging
a burrow for itself — an example on a lai^r scale of the tragedy
described by W. H. Hudson in " The Naturalist in La Plata "
(pp. 180-82 of the 1896 edition) :—
" On the grassy pampas, dry bare spots of soil are resorted
to by a class of spiders that either make oi take little holes in
the ground to reside in. . . .
"... Now, in summer, to a dry spot of ground like this,
comes a small wasp, scarcely longer than a blue-bottle fly,
body and wings of a deep shining purplish blue colour, with
only a white mark like a collar on the thorax. ... It visita
and explores every crack and hole in the ground, and, if yon
watch it attentively, you will at length see it, on arriving
at a hole, give a little start backwards. It knows that a
spider lies concealed within. Presently ... it disappeara into
the hole and remains there for some time. Then, just when
you are begiiming to tiunk that the little blue explorer htf
been trapped, out it rushes, flying in terror, apparently, from
the spider who issues close behind in hot pursuit ; but, befors
they are three inches away from the hole, quick as lightnmg
the wasp turns on its follower, and the two become locked
together in a deadly embrace. Looking like one insect, they
epin rapidly round for a few moments, then up springs the
wasp— victorious. The wretched victim is not dead; iU
l„y,l,Ajh,COOl^lC
( xli )
legs move a little, but its soft body is paralysed, and lies
collapsed, flabby, and powerless. . . . When the wasp has
sufficiently rested after the struggle, it deliberately drags the
disable*^ spider back into its own hole, and, having packed it
away at the extremity, lays an egg alongside of it, then,
coming out again, gathers dust and rubbish with which it
fills up and obliterates the hole. . . ."
OBSBBVATlOMa ON FoSSOBS IN SaST AFRICA BY Dft,
0. D. H. Cabpenteb. — Prof. Poultom said that an observa-
tion lecoided in a letter written to liim Jan. 18, 1917, by
Di. Carpenter, threw farther Ught on the storing of Hesperidae
by Bembeddes : —
" Yesterday (on a journey from NdaU, 33° 15' E., 4° 45' 8.,
to my new post Igtdtda, on the Central Railway, aboat 40
miles £. of Tabora) I had such an interesting observation of
a Bembex that I write post haste to tell yon. I was catching
Skippers on mud on the road (and had got several of a beautiful
species, bright golden brown, with the hind-wing below
marked with a number of whitish radiating streaks [Oxypa^ntt
harona, Westw., = ritso, Mab.]. This I had never seen
before; also a lai^ and wonderful white species [Leuco-
cMUmea Mndei, H. H. Druce] which I took at first for a small
Bdenois, among which it was drinking. These will come by
the next box. To return to our muttons) and suddenly saw
an unmistakable Bembex flying round my legs, on which
sat, and bit, numbers of Q. morsUana, Tabanits, and Baema-
topota (indeed, I had been bitten so much that I was quite
jumpy I). I naturally thought the Ben^tex was after these,
though she was not of the species that I have often seen
come round me looking tor fat Olossina or roMniu. She
was large, greenish yellow, with a transverse black band
across each abdominal segment. After buzzing around for
a bit (the hum was the characteristic Bembex hum, which is
of a tone difierent from that of other Hymenoptera, ot AeUidae,
whose bu2z is equally characteristic) she hung in the air a
few inches above a Lycaenid, quietly drinking on the mud,
and pounced on it. I couldn't see exadly what happened,
but the Lycaenid was dropped, and the Bembex moved away
to where a Skipper was also drinking, and hung poised over
u„,,i,A J 1^, Google
( iBi )
that for ft Becond or so — evidently ezamining it cloaely. The
raeult seemed unsatisfactory; she moved away and hung
over another, then, pounce, and she'd got it t Foi a brief
moment she remained in the air, havering juat over the^^und,
holding the Skipper. I struck hut, oh, I mined her ! and
she was o9 like & streak of lightning. I did not see her again.
I waited a little (aa long as I dared, for the afternoon thundet-
stonn was coming up) but she did not come back; bo once
again I have sent you a new Fosaor observatioD, without the
specimen 1 It was interesting that she should catch a Lycaenid
and drop it ; I was so interested watching her that I could not
devote any attention to the discarded piey. I expect the
reason she did not take the jint Skipper was that it was not
fat enough. My observations on the Bembex that preyed
on Olossina on Neadzi Island (see my first Sleeping Sickness
Beport) showed that the prey was the fattest that could be
selected — often a pregnant 2 fly. So the rejected Skipper
may have been a <3 and the chosen a $ ; it was a dark hrown
species with narrow wings, hind-wing beneath with a row of
tiny white dote (I don't know the generic name). Has a
Bmibex ever been known to prey on Skippers or on any other
butterfly? Don't say it was an Asilid, for it was not/ I
could tell an Asilid from a Bembex with my eyes shut by the
buzz alone ! Moreover, I have yet to meet the Asilid that
tekes ite prey sitting. Also the broad abdomen — without
pedicel — ^rendered it unlikely that she was not a Bembex
but some other kind of predatory (perhaps Diplopteroos)
wasp. I should much hke to know if there are any other
notes of this kind."
The following observations on AmmojAtla beninenais. Pal.
de Beauv., or a species very close to it, were recorded in a
letter written by Dr. Carpenter, Jan. 14, 1917, from Ndala,
which appears on the map as a Hisaion Station : —
" On Jan. 3id I disturbed from hei work, on the road, a
fine Sphegid. On the wing her long legs were bunched to-
gether, hanging down very conspicuously. Her prey, a large,
smooth, brown Noctuid larva, lay in a slightly curved posture
at the brink of a vertical burrow. I sat down beside it, and
when the Sphex had returned and, as I did not move, recom-
Ij.y.l.AjL.yCOO'^lL'
( xM )
menced her work, I was able to follow closely. She examined
the larva very carefully from head to tail, and then went
down the hole and brought up an ' armful ' of soil, held by
the fore limbs up against her ' chin.' She walked away with
this, and deposited it about 3 inches away, giving, at the
exact moment when she put it down, a short sharp buzz.
This was repeated once or twice, and then, having placed the
tip of bet abdomen at the orifice of the burrow, she felt her
way down backwards, and before she disappeared seized the
larva just behind the head by bet mandibles and drew it
down after her. But the chamber at the bottom was not
big enough, so with loud buzzing she pushed her way past
the larva and came out again ; then, standing over the hole,
she put her head down and seized the larva aa far in front of
its hinder extremity as she could reach, and dragged it up to
the surface. The next step was to grip, between her fore-
legs, the extreme end of the larva, and to shift her mandibles
so as to get a fresh hold further forward; thus she lifted it
again, and in this way dragged it right out of the hole and
laid it at the brink, the concavity of its posture directed
towards the hole, the two extremities at the very edge. She
then went down again and brought up another armful of earth,
depositing it in the manner previously described. Having
done this several times she again went down backwards,
precisely as before, and dragged her victim down by its ' neck.'
But again the chamber at the bottom of the burrow could not
contain this large larva, and its hinder end projected up into
the burrow. So once more the unskilful wasp had to push
her way out with loud buzzings, and drag the larva out again
in the same way, and further enlarge the chamber. Still it
did not prove big enough, and this all happened four more
times ! On the sixth occasion, the larva not having been
diapoaed quite exactly in the right position at the mouth of
the hole, when she backed down she caught hold of the posterior
and not the anterior end. The larva then moved very feebly
(it had had a lot of pulling about !) and the wasp at once
perceived that something was wrong. She hurried out of the
hole and examined it very carefully. She then adjusted it
to the proper position, went down again, and dragged the
u„,,i,A J.., Google
( div )
larva down by the ' Deck ' as usual. But still it conld not b«
wholly contained in tlie chamber, and had to be dragged oat
once more for further digging : this time, however, a new
difficulty arose ; it waa bo far down that the Sphex, standing
over the hole, coold only giasp the extreme tip of it£ body
(posterior end), so that wheo she drew it up there was nowhere
for the fore limbs to grip ! Thus, every time she tried, the
larva fell down to the bottom of the hole again, and she began
to get very agitated.
" At last, after ten efiorts she managed somehow to get a
grip with the forelegs, and ao pulled the larva out as at Erst
described, enlarged the chamber once more, and pulled the
larva down for the seventh — and last — time; for now it
fitted wholly in the chamber and none of it was visible when
one looked down the burrow. After a pause — quite remark-
ably short — for ovipoeition, the wasp came out and proceeded
to fill up the hole in the usual way, by biting oS small lumpa
of soil, putting them down and ramming them in with her
head, but making very little use of the method of scratching
loose earth backwards. When she had all but finished I
caught her (and send her to you for naming), finishing off the
burrow myself lest hei progeny should suffer I (But I did not
bite off small lumps of soil and ram them down with my
head.)
" Thei« are one or two points worth remarking : —
" 1. Repeated efforts to get a laige larva into a chamber
too small for it. This seems to imply that the wasp digs a
chamber of orthodox size, and finds larvae ajlerwardt. Doea
she use larvae of varying size, or was she inexperienced ?
" 2. The larva must be dr^ged down by the anterior
extremity. One sees no reason why this is necessary.
" 3. On the whole the wasp works quietly. The pro-
longed, high-pitohed, penetrating, buzz which S. marffinatm
makes the whole time she is burrowing, so that one often
hears her at work long before one sees her, is replaced in this
species by a short buzz when the armful of earth is put down.
Also, when she had to push her way out past the larva in the
too small burrow she gave a buzz which one could easily
imagine to express vexation I "
n,g,t,7.dh,G00glc
( »1' )
Mr. E. E. Gbebm observed that in Cejlon a large Reduviid
bug feeds on a etiU Urgei milliped.
Mr. Talbot observed that in his garden he had noticed
that flf-catcbeis paid no attention whatever to butterflies.
Dr. Chafhan also conunented.
Letier from a FdXow irOemed in Q&rmany.
The Secretary read the following interesting letter from
Mr. E. M. Dadd, F.E.8. :—
"Dear Sik,
" The writer, as you are no doubt aware, has been a
Fellow of the Society for many years past. As a resident
in Germany at the outbreak of the war he shared the lot of
his fellow countrymen, and has now been interned at Ruhleben
since Nov. 6, 19U.
" In the meantime those amongst us interested in Natural
Science have formed a Natural History Society, and weekly
meetings are held, papers read, and furthermore a regular
course of Lectures in Natural History, Biology and Botany
satablished under the direction of Dr. Lechmeie. The writer,
as the sole representative of Entomology, has also given several
lectures on various entomological subjects. In the meantime
we have established as well-organised a Biological Laboratory
as could be expected under the circumstances, with microscopes,
and are extending the same as far as possible. It is proposed
this year to form a small section for the study of entomology,
and space has been placed at my disposal for research work.
" I have been able to get in a certain amount of my own
material and a certain amount of literature, but this hardly
suffices. Might I therefore appeal to the Society for assistance
in the way of books or magazines and more especially in
the form of duplicate insects (butterflies in papers), beetles,
hymenoptera, etc. If desired the books could be returned
later on.
* • « • a ■
" Trusting to meet with a favourable response, and with
land regards and best wishes to all friends,
" I remain yours very sincerely,
"Edward M. Dadd."
l„y,l,Ajh,COOl^lC
i
( ^Ti )
Paper.
The following paper was read : —
" On a collection of Lepidoptom made in Eaat Africa by
Mr. W. A. Lambom, F.E.S.," by H. Elthingham, M.A.,
D.Sc., F.E.S.
Wednesday, October 3rd, 1017.
Dr. T. A. Chapmak, M.D., F.Z.8., Vice-Preaident, in the
Chair.
EUclion of a Feliow.
Dr. Gborob Granville Buckley, M.D,, F.S.A., Holly
Bank, Manchester Boad, Bury, Lanes., was elected a Fellow
of the Society.
Dealh of an Bonorary FeUoK.
The death of Dr. Emil Frey-Gessner was annoonced,
and a vote of condolence with his daughter was passed.
ExhQnti&ns.
COLEOFTERA ON CNUSDAL FoOD-pLANTS, ETC. — Ml. DoNIS-
TBOBFB exhibited the following Coleoptera : —
1. Miants camparmtae, L., taken on the Downs at Findon
(Sussex), June 11, 1917. It occurred in some numbers, but
only in a email species of buttercup, of which a specimen was
exhibited. It has only been recorded heretofore in flowers
of Campanula and PkyUuma. A small black Chrysomelid
larva, about the size of the Miama, also occurred in the
buttercups, and they were superficially very like each other.
2. Lycoperdina succincta, L., taken at Barton Mills (Sufiolk),
Sept. 9, 1917, in fungus. This species was first discovered
te be British in October last year by Dr. Nicholson, who
took a series in ripe Lycoperdon gemmatum. Mr. Donisthorpe
was evidently a little early for the beetle, as his specimen
was slightly inunature, and he did not find any more, but ha
took about a dozen larvae in a fresh Lycoperdon genunatvnt,
which he is trying to breed.
3. Casaida faataota, Schall. Taken at Goring Woodi
l„y,|,AJI^,COOl^lC
LABVAE or SAWFIAtS,
1. Cimbtx luUa, L. Coiled imdeT a Sallow-leaf.
2. Abiafateiala, Leach. Coiled under a Snowbeny-leaf.
3. Trichioeampus viminalia. Fall.
(1 and 2 were phol^^aphed from living specimens on their actual
food-ptanta. 3 is from the inflated skin of a larva found on Poplar
in the grounds of the Natural Histary Museum, S.W.)
PLATE C.
LABVAi or SAWnjES.
1. Eriotampa ooata, L. On Alder, covered with a mealy white
secretion.
2. Pteronidta talieis, L. On loDg-leaved Willow. (The colour ol
the body is blue-green in the middle, pale salman colouc at
each end.)
3. Nemalua {Croesut] seplenlriorialie, L. On Birch. The insect
is rising into its "threatening attitude," and has eiserted
its ventral "scent-glands."
4. LygaetMtmalJu crauieotnU, F. On Poplar, surrounded hy its
" stockade " of inspissated secretion formed into Uttle white
6. One of the pillars ( x 10 diameters).
n,g,t,7.dh,G00glc
n,g,t,7.dh,G00glc
Proc. Enl. Soc. Land., 1917.
Half- Torn Eng. Cb. Lid.
Larv^ of Sawfues. '■"■- Ji^v^"t)glC
n,g,t,7.dh,G00glc
( zlvii )
(Sussex), July 28, 1917, on Inula dysenterica (Fleabane).
This U its first record on Fleabane, and all the specimens
were of a bright yellow and black colour when alive, and not,
as is usual, red and black. Though freshly emei^d they were
quite mature, and moreover were kept aUve for a long time
without changing colour in any way. A specimen from Box
Hill of the typical form, taken on Inula aonyza (Ploughman's
Spikenard), May 7, 1909, was also exhibited for comparison.
It was previously only recorded from Senecio jaeobae
(Ragwort).
Fhotooraphs of Sawfly Labvab. — The Rev, P. D.
MoRicB exhibited with the Epidiascope a set of photographs
(mostly taken from living specimens feeding or resting on
their usual food-plants) of the following Sawfly larvae : —
Cimbex luiea, L., coiled up on Sallow, Abia fasdata, L., do.
on Snowberry, Phymatocera aterrima, Kl., on Salomon's Seal,
Rhadinoceraea micane, Kl., on Iris, Empria [Poeoilosoma)
htteola, Kl., on Yellow Loosestrife, Briocampa ovala, L., on
Alder, Nematiu {Croesus) seplentrianalis, L., in "threatening
attitude " on Birch, Pieronidea {Nematus) solids, L., on Birch,
Trichiocampus viminalis. Fall., on Poplar. Also of a cast
skin, showing the Y-shaped spines, of a Periclista larva —
probably mdatwcephala, F. ; and of the sort of " palisade "
— a ring of little columns formed by the rapid drying of some
bubbly, froth-Uke secretion from the mouth — with which the
young larva of Lygaeonematus compressicomis, F., surrounds it-
self while feeding on a leaf of black Poplar. (Plates B and C.)
The Chairman observed that Sawfly larvae in moulting
attach themselves by the tail to a gummy substance pre-
viously deposited by them, because they have no hooks on
the terminal pad of the prolegs such as occur in the Lepido-
pteia; the true legs, which in the latter are generally used
rather as hands than as feet, are in the Sawflies provided with
claws sharp enough to penetrate the leaf.
Paper.
The following paper was read : —
" Further notes on Recapitulatory Attitudes in Lepido-
ptera," by T. A. Chapman, M.D., F.Z.S.
n,g,t,7.dh,G00glc
( zlTili )
Wednesday, October 17th, 1917.
Dr. C. J. Gaean, M.A., D.Sc., Preaident, in tlie Chaii.
Election of FeUowa.
Mr. JoEM Williams Hockin, Castle Street, Launceaton,
Cornwall; Col. Tusenne Jebmyn, HighcliSe, Weston-super-
Mare; Mr. Abthur Wallace PicKARD-CAMBRiDaE, M,A-,
Balllol College, Ozfoid; and the Bev. Prebendary A. P.
WiCKHAM, Kast Brent Vicarage, Uighbhdge, Somerset, were
elected Fellows of the Society.
Exkibiiions.
Hyfer-fabasites on Ap&nteles OLOUERATOS. — Ml.
DoNiSTHOBPE exhibited a. number of small yellov cocoons
vhich were taken on a fence at Putney on Sept. 15tli
last, and which had emerged from the body of a White
Butterfly larva. These cocoons, belonging to a para^te on
the larva of this butterfly, are of course common everywhere
just now where the butterfly has occurred. He stated that
he had found the cocoons just after they had emerged from
the hirva to which they were attached, the latter being still
aUve, and he observed two (or three ^) small Hymenopteroaa
insects hovering about the cocoons. One of these was secnied,
and the larva and cocoons were taken home in a glass-topped
box. On October 8th Hymenopterous insects began to
emerge from the cocoons and were still doing so. Of the 30
specimens exhibited, some of which were alive, 28 belonged
to the insect captured on Sept. 16th, an Ichneumon, and 2
(a S and $) to another species of Hymesoptera, also an
Ichneumon. It would aeem certain that these two species are
hyper-parasites, parasitic on the parasite of the butterfly
larva. The questions arose if the cocoons had not been
disturbed, would they all have produced the hyper- parasites f
and is it necessary for the latter to seek out the cocoons just
after they have emerged from the original host's body ?
A NEW Sub-species or Morpho rhetbnor. — ^Mr. Diceseb
exhibited a probable new sub-species of Morpho rhetawr, and
gave the following description ; —
n,g,t,7.dh,G00glc
( xlii )
" This anb-species, now received foi the fiist time from
Colombia, diSera from the two other Bub-species, cacica from
Pera and etuebea from tlie Amazons, in having a complete
row of twelve white submarginal spots on each side of the
upper surface, six on each wing, instead of from three to five
on each front wing only. It has also a very small white spot
beyond the cell on the front wing, and the small white
crescentfl between each nervore on the margins are more
distinct. On the under side the ground colour is lighter,
maldog all the markings stand out more distinctly. Locality
Bio GuiTiQoiA, April 1917."
An aberrant Wasp. — Dr. CHAFitAN exhibited an aberrant
specimen of a wasp {Vespa germanica), and made the following
observations : —
" Wasps raiely come into my sanctum, but on Oct. 6th
one came and seemed very interested in some preparation I
was working at, so that I easily noticed that there was some-
thing unusual about some abdominal segments. The insect
IB the one exhibited. The second abdominal segment (count-
ing in the usual incorrect manner) is wanting in the middle
dorsal line, and the two sides nearly meet each other, tapering
from a nearly normal width laterally to almost a point where
they are opposed, and where continuity is maintained by
membrane ; the markings on the two sides are not symmetrical.
" The first (basal, really 2nd abdominal) segment is pro-
duced mid-dorsally at its hinder margin, so as to form an
angular projection, largely compensating for the defect of
the following segment.
Sketoh of tenfologioal Bpecimen of Yapa genuauea 1} , x about 4.
" I add a rough outline of the aberrant portion of the
specimen. How did such a defect come about } It might
be the result of an injury at a late larval stage, but it is TOiy
FROC. ENT. BOO. LOND., II-IV. 1917 D
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unlikely for a wasp grub, piot«cted in ita cell in the nest aa
it is, to receive such an isjuiy. It rather more atrocgly
suggests a congenital defect in the closing of the umbilical
opening. It is not easy to say whether the want of symmetry
in the peccant segment supports this view or the contrary.
I will hand the specimen to the British Museum."
A VERT HARE BRITISH BEETLE. — Mr. 0. E. Janson ex-
hibited a fine example of Tapinotw seUalws, Fab., taken by
him on June 9th last near Homing, Norfolk, and recorded in
the October number of the Ent. Mo. Mag. Only two British
specimens were previously known, these having been taken
in the years 1638 and 1846. He also showed some other
Coleopt«ra of interest taken in the same locaUty.
A Bbitish 6?eciubn of Notodonta bicoloku. — Mr. O. E.
Janson also exhibited on behalf of Mr. L. H. Bonaparte-
Wtse, who was present as a visitor, a fine male specimen of
jVotodonfa bicoloria, Schifi., taken by him near Blillamey on
June 7th last, and recorded in the " Irish Naturalist " for
October.
LiviNQ Derhestid Larvae. — Mr. Green exhibited living
larvae of a Dermestid beetle, Tiresias serra, found under dead
bark of an oak tree, in the neighbourhood of Shrewsbury.
The larvae have the abdominal segments clothed with erect
rufous hair. There is also a loose caudal tuft of long greyish
hairs, which is vibrated rapidly when the insect is annoyed
or disturbed.
Mr. Main and Mr. Donistbobfe conunented, the latter
observing that judging from his experience of the larva in
observation ants' nests, the vibration had apparently a
protective value.
Mr. Green also read the following note on the oviposition
of the sawfly Pteronvs ttrtifer.
" The method of oviposition of the sawfly Pterotms serti/er
has been a subject of discussion at several of our meetings
during the present year. Insects raised in captivity, by
Mr. Morice and myself, have refused to throw any light on
the question. 1 have now been successful in obtaining eggs
from a wild female. The parent was captured on the 1st of
this month, and confined in a glass tube with a sprig of pine
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foliage. The contents of the tube were examined, each day,
for the first week, without leeult. The insect remained alive
hot, apparently, uninterested in the question of reproduction.
The tube was put aside and (it must be confessed) completely
foi^otten — ^tuitil this morning, when the insect was seen to
be dead, with its saws choked with a mass of what appears
to be fragments of plant fibre and cell tissue. An examina-
tion of the pine foliage showed some semitranslucent paler
areas along the edges of the needles. A rough dissection
proved that these pale areas each contained a single creamy-
white egg. At the tip of one needle, an egg was partly ex-
posed ; but in every other case the eggs are completely concealed
and their presence indicated solely by the translucent areas.
There is no noticeable disturbance of the tissues, nor is there
any extraneous deposit of fibrous or seoretionary matter —
such as has been observed above the embedded e^s of P.
pini. The egg capsule is very thin and fragile, and is easily
ruptured. It should be noted that the above remarks are
the result of a very hurried and superficial examination,
undertaken just before coming up to attend this meeting."
Wednesday, November 7th, 1917.
Dr. C. J. Gahan, M.A., D.Sc., President, and afterwards
Dr. G. B. LoNGSTAJP, M.A., M.D., Vice-President, in the
Chair.
Election of Fellows.
Prof. T, D. A. CocKEEELL, of the University of Colorado,
U.S.A., Miss D. J. Jackson, Swordale, Evanton, Koss-shire,
and Mr. Jr98E Johnson, 16 and 17 Marston Road, Ste&ord,
were elected Fellows of the Society.
Exhibitions.
Nuptial Flioht of Butterflies. — Dr. F. A. Dixey
exhibited several pairs of Pierines captured by Dr. G. D. H.
Carpenter, at Itigi, 150 miles east of Tabora in what was
German East Africa. In commenting on them, he said —
" Professor Poulton has put into my hands the following
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eztracta from two letter^ writteo by Dr. Carpenter and dated
respectively on August 22 and 31 of the present year.
" ' Talking about pairs in ccjt., 1 have just been re-reading
the " Descent of Han," and when Darwin talks about the $
Pieria and Epinephele supporting the <S, it struck me that my
impression was, out here, that facts would not conform to
this. I fancied that I had never seen the 9 supporting the ^
in Pierines. So 1 have started collecting pairs in cop. and
noting which sex carries the other. Between Aug. 18 and
to-day inclusive I have got pairs of two species of Belenoia
(one each), three of a Teracolus, and four of Pinacopteryx
timana, i^ i^ all supporting $ $. I shall continue to do this,
and wish I had done ao before (one's entomological sins are
OHUally of omission I). Seeing how, broadly speaking, 3 and
$ Belenoia are on a par with our home Pieris, the $ ought to
cany the £. Also F. simajta, in one sense, seeing that the
$ is more blotched with black, though the 3 has black veins
better developed. I am more of the teebng that the 3 Pieris
should be looked on as the " brighter " owing to absence of
black. What is known about say the 8. American mimetic
species, with black and white 3 ^—i. e. which sex carries which t
I remember glancing at some notes by Dr. Diiey on this
point, but as 1 had not devoted any attention to the subject
I did not read them carefully.'
" ' I wrote recently that I was getting interested in the
question of, when butterflies unite in cop., which sex supports
which. What made me pay attention to it was the " Descent
of Man," where Darwin deals with sexual selection among
insects, and telks of the diSerent colours of sexes in butter-
flies (chap. xi. p. 319, in revised second edition of 1899). He
says that in England, with Colias edusa, Pieris, and Epinephele
jurtina " it m ifte ? $ thai support the 3 3, «o thcU the part
wkicA the two sexes play is reversed, as is their rdalive beauty,"
and, later, " the $ $ take the more active part in the final
marriage ceremony, so that we may suppose that they like-
wise do so in the wooing." Well, since 1 last wrote I have
got quite a number of pairs of Pierines t'n cop. : namely,
Belenoit (two species) 7, TeraocAtu i, and Pinacopteryx simana
14, in every case 3 supporting ?. With Teracolus, the 3
Ij.y.l.Ajl^yCOO'^lL'
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being " brighter," this is as it efaould be according to Darwin.
In P. simana the $ is of a duller white, and there is a smatl
extra patch of dark tint on the f-w., in which also the black
veins are leas conspicuous than in the S. I have found in
one case the $ supporting the ij. With Bdetwis, however,
there is a very marked difiereoce between <J and ?, though
it is a question whether the ^ is " brighter " than the <?.
However, the relative difference is of the same type as in
the English whites. Now I see by notes of Dr. Dizey in
Proc. Ent. Soc., March 17, 1916, p. 36, that he got 3 English
Pieris, ^ all supporting $, although as regards Satyxinee his
6 observations all agree with what Darwin says. This seems
cniious. Was Darwin misinjortned 9 I suggest that it would
be a good thing to bring this point before the Ent. Soc. next
spring, asking Fellows to send Pierines caught in cop. to the
Hope Dept., clearly marking which sex supports which. For
here is an opportunity for those who cannot get abroad, to
work at a point of interest to Darwinists. We should in this
way be able to get ample evidence. Of course notes on all
species (but particularly Satyrines and Pierines) would be
valuable.
" ' I may say that from what I have seen of the courtship
of Betenoia, the $ is not the most active partner. After a good
deal of fluttering about one sees a ? settle on the ground
vith wings half or almost fully expanded, and the <? stands
behind or a little above in a state of frantic excitement, and
vigorously scrapes or paws the upper surface of the wings of
the ? with his lore-limbs ( ! first or second pair, I am not sure
of this). I witnessed yesterday the actual union of a pair of
P. simana. The $ settled on the ground between some grass
blades quite quietly, with wings closed; the 3 stood on a
grass stem with wings widely expanded over the $, so that I
could not see her. The union was effected very quickly, and
the S flew off carrying the passive $. Unfortunately, owing
to a gusty wind, I failed to catch this couple.'
" By the kindness of Prof. Foulton I am able to exhibit
the actual specimens sent home by Dr. Carpenter; the species
are as follows: Teraeolm casta, CSerst., 3 pairs; Herpamia
eriphia, Godt., 1 pair; Pinacopteryx simana, Hopff., 6 pairs;
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Beknois gidUsa, Godt., 1 pair; BdenoU mesentitM, Cram., 1
pair. In each of these eleven pairs the S was canyuig the ?.
" Dr. Carpenter refers to a note of mine which appealed
in Proc. Ent. Soc. Loud., March 17, 1915, pp. xlvii, xlviii.
I there recorded six observatioDS on the nuptial flight of
IHerines, including our three common British species, and in
South Africa a Bderwis and a PittacopUryx. Id every one of
these cases, as in Dr. Carpenter's, the male was eupportitig the
female. I have never seen a Colias under these circumstances,
but the combined testimony as to the genera named seems
to be pretty complete. It will be remembered that in an
earlier volume (Proc. Ent, Soc. Lond., Nov. 4, 1914, p. xcviii),
Hr. Wheeler had not«d that, according to his experience, in
Lycaeuids the male always carried the female, but that in
Nymphalines, Satyrines and the common Pierids the female
carried the male. With regard to Lycaeuids and Satyrines
my own observations, as given in the place above cited, are
in full accord with Mr, Wheeler's, but it will be remarked
that in the case of the Pierines Dr. Carpenter's observations
and mine agree with one another, but not with Mr, Wheeler's.
This, I think, gives ground for Dr. Carpenter's suggestion
about asking entomologists to make special observations on
this point, sending their notes and specimens to the Hope
Department.
" It would seem that both among Pierines and Nymphalines
occasional exceptions may occur to what appears to be the
general rule, for while in the only instance that I have noted
of the latter subfamily, Argynnis cydippe {adippe), L., the male
was carrying the female, the detailed testimony of Hr. Wheeler
is entirely in the opposite direction. Dr. Carpenter also
seems to have observed one instance of the female Pino-
copleryx siiiiana carrying the male — the only one where thia
was the case out of many instances of the nuptial flight of
this species and other Pierines observed by him.
[In the first three days of July this year, I observed at
Lyndhurst eight pairs of Dryas paphia in flight. In «x
instances the 3 carried the $, in two the $ carried the o.
Details are given in the " Entomologists' Record," vol. xxix,
p, 166.— G. Whkklkh.]
u,y,l,7rJM,G00glc
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" Dr. Carpenter's series is Id other lespects interesting.
It is to be observed that all the specimens are of the ' dry '
phase, Boine veiy diy. The Terias bngitta are leas dry than
the other species.
" The pairs that I have list«d as Teracolus casta may repre-
sent a new subspecies. On the upper side they strongly
resemble T. xafxtholeuca, described by Miss E. M. Sharpe
from Kavirondo; but the under side is difierent. The dis-
tinction, however, may be seasonal ; the types of zantholeuca
weie taken in January, and Dr. Carpenter's specimens in
June and Augost."
Mr. Kate, Prof. Pouxton and Mr. Bethune-Bakxr com-
mented on Dr. Dizey's exhibit.
Bred Lycaena ARioN.^ilapt. E. B. Purefoy exhibited a
short series of L. arion which had been bred up from the egg.
After the 3rd moult they had been carried into the nests of
Myrmica laevinodis.
Dr. Chapman congratulated Capt. Purefoy on his success
in breeding L. arion, and said that he had larvae of L. aloon,
sent by M. OberthUr, feeding in his ants' nests.
Mr. DoNiSTHORPE corroborated an observation of Dr.
Chapman's, that the ants, on being disturbed, carried ofi the
larvae of Lycaenids, beetles, etc., before removing their own.
FABAStTES AND HYFERPAIUSITEB OF APHIDAB. — Prof.
FouLTON exhibited on the screen enlarged photographs taken
by Mr. Alfred Robinson, of the Oxford University Musenm,
of the parasites and hyperparasites bred from three species
of Aphidae in the Oxford district by Mr. H. Britten of the
Hope Department. The species were as follows : —
HO>r(APnD»B)
puusm (ButmnDAB)
MacTotipham rota*. L.
Aphidi^. sp.
(Chai^idikab)
JfocroniAuni vrtieae,
Sohr. ■
/ffpWrw, sp.
i Ataphtt, ip."
•^ttsxt- "'"■'■
"
1
3. {Ctkipidab)
u,y,i,Ajh,Cooglc
( 1" )
Tbe pAiasites and hjperparaaitee emerged from tfae BwoDen
body of both species of Maoroei^kum, wheieoB in Drepano-
aipkum the Braconid larva emerged and constructed beneath
its host a beautiful disc- or button-shaped cocoon from which
also the hypeipaiasitea, when present, emerged. Into the
roof of this cocoon the ventral euiface of the host was woven.
ThB ChaLCID FARAfilTE OP VaNBSSA UBTICAS, L., WAITING
BESIDE THE LABVA FBSFABI»a FOB FUFATION. GOLDSN PDFAK
NOT CAUSED BY PARAsmsH. — Prof. PoDLTOM exhibited 22
males and 11 female Pteromalut, sp., together with the Vanessid
pupal shell from which they had emei^d. The female parent
had been seen, on Aug. 17 last, at St. Helens, Isle of Wi^t,
patiently waiting beside the butterfly larva, which was rest-
ing motionless on a grey stone wall before it had suspended
itself, and therefore many hours before pupation. On Aug. 20
the pupa was removed, and on Sept. 22 all the exhibited
specimens of Pteromalua had emei^ed and were found alive
and active. The life -cycle thus contrasted sharply with
examples (6 males, 48 females) of Pleromalits puparum, L.,
much larger in average size, bred from Pyranuu atalanla, L.,
in ]90(M)]. In this case the female Chalcid bad been found
seated on the fresh moist pupa, Sept. 7, 1900, at Calverley
station, near Bradford, Torkshire. Ova were laid on this or
the following day, and the offspring emerged at Oxford on
May 22, 1901.
The pupal shell of the V. urlicae was grey, as could still
be seen, and harmonized well with the stone wall, showing
that the power of colour adjustment was not removed, as
some have supposed, by the presence of parasites. The
normal efiect of the green leaves of the nettle was to produce
golden pupae — the nearest approach to green of which this
species was capable. Such golden pupae were sometimes
parasitised, but so were the dark pupae, as exemplified by
the exhibited specimen. In former years Prof. Poulton had,
by the use of the appropriate backgrounds, bred hundreds of
healthy golden pupae which produced normal butterflies.
The opaque cuticle of a parasitised dark pupa showed no
change, but the transparent. cuticle of a parasitised golden
one took on a peculiar brassy appearance, probably, due to
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changes in the stratified thin films of liquid, which, by inter-
feience of light, produced the metallic appearance. Prof.
Poulton believed that a Bimilar effect was to be seen shortly
before normal emergeDce, but, as it lasted for a veiy short
time, it was noticed rarely as compared with the far more
enduring effect of parasitism. It was to be observed, finally,
that the Chalcid parasite laid its eggs in the pupa immediately
after the larval skin had been thrown off and before harden-
ing, and that the pupal colours had been determined mnch
earlier, during the second and third of the prepupational
stages of the larva, viz. " for about 20 hours preceding the last
12 hours " before the skin is thrown off and the pupa revealed
(Phil. Trans. Roy. Soc, vol. 178 (1887), B, pp. 319-98, 438-9).
A STNAPOSBMATIC SERIES OF 272 LtCID BEETLES OP 9
SPECIES TAKEN ON ONE PLANT IN ONE DAY BY G. D. H. CaR-
FBNTEB, IN LATE GERMAN EaST AfRICA. — Prof. POOLTOH
exhibited the series referred to in the following extract from
Captain Carpenter's letter of March 24, 1917, from Itigi (in
34°, 30' E., 5° 46' S., on the Central Railway, about 150 miles
£. of Tabora), where the beetles were captured. The food-
plant, of which a piece was exhibited, had been kindly deter-
mined by Dr. O. Stapf, F.R.S., as the Asclepiad PenlarrhitMrn
insipidum, E. Meyer, common and widely distributed in
tropical and South Africa. Prof. Poulton had added [to
Captain Carpenter's table] the names of the species and the
numbers by which each was represented in the collection.
" I am now sending you a small box of about 270 Lycidae.
On 23. ill. 17 1 came on a flowering creeper which had a number
on it, and started picking off a few of the different species.
While doing so it suddenly struck me how much you would
like a Urge number, for the proportion of species. So I set
to work to collect them all into two empty tins which soon
got full. I couldn't absolutely clear the bush because more
beetles kept arriving on the wing, but, practically speaking,
I got 99 %. It was quite interesting laying them out after
the slaughter and trying to make out how many species there
were, with the aid of 17 pairs found in copuM. I made out
9 species, at least, of two or probably three genera, and the
nnmbers are as follows : —
n,g,t,7.dh,G00glc
( Iviii )
Species 1. [58 (J :B2 5192 plus 6 pairs in eop.—{Lyeui
(Meroiycus) JemoralU, Boui^.]
Species 2. [51 <} : 26 $] 79 plus G paiis. — [Lycus davimeri,
Bourg.]
Species 3. [4 J : I S] 3, no faiie.—[LycM ampliatUB, F.]
Species 4. [20 ,J : 1 S] 13 plus 1 pair and an extra ^
associated together with it. — [Ljfcua trabealus, Gu£r.]
Species 5. [5 (J ; 3 ?] 3 plus 1 pair. — {Lycus sp. near tra-
beatw, Gu^r.]
Species 6. [2 (J ; 1 ?] 2, no pairs. — [Lyoug (Meroiycus) rot-
Irattta, L.]
Species 7. [3 d ; 3 2] 5 plus I pair. — [Lyeus (Meroiycus)
podagricus, Bourg.]
Species 8. [!3 5 : 5 ?] 15 plus 1 pair. — [Lycus marahaUi,
Bourg.]
Species 9. [20 J : 4 ?] 20 plus 1 pair.— [lyous (Lophdyau)
amoenus, Bourg.]
Totals [176 (J : 96 S] 232 plus 17 pairs plus 1 superfluous (!)
male.
" 1 am not at all sure that Species 2 does not include 2
species, but apart from size I could make out no essential
difference. Quite a pretty little exhibit for the Entomological
Society. But, alas, search as I would I could find no Lycoid
mimics : there were only a number of ' Blue-bottles ' and a
large Pompilid which rather frightened me, as she seemed to
resent my presence, and 1 was much afraid of her! There
was one wretched little brown Syntomid moth which (how-
ever ' enthusiastic ') one couldn't pret«nd was Lycoid ! How
different from Bugalla with ttfi lovely Lycoid Fossors and
Longicorn ! I send a bit of the creeper, which you may like
to identify."
The number of specimens was rather larger than that given
by the captor — 272 to 267. In the determination of these
difficult species kind help had been received from the Presi-
dent and also from Mi, H. Britten, who had prepared the
male genitalia of many specimens. The discrepancy between
Captain Carpenter's numbers and those in square brackets
under each of the nine species was in part due to difference
between the totals and in part to the fact that a more exact
Ij.y.l.Ajl^yCOO'^lc
( lil )
compariBon could be made when the material was carefully
mounted. Specimena in the Hope Department identical with
No. 1 had been named by Bourgeois L. (M.) scapularis, Murray,
bat they were certainly dlfFeient from the unique female type
of this species in the British Museum, and the President con-
sidered them to be Bourgeois' jenwralis. No. 6 was similarly
identical with specimens named by Bourgeois L. (Af.) dentines,
Dal., var. flavoscaptdaris, Bourg., but the President conaidered
that denlipes waa a synonym of rostratus, L. Species 4 and 5
were closely similar, b'ut the male genitalia of No. 4 resembled
those of specimens in the British Museum from Abyssinia, the
locality from which Gu£rin described trabeatus, and the
President had therefore con^dered No. 4 to be true trabecUus.
The whole assemblage presented an extraordinarily uniform
appearance, the only marked diSerence, on superficial examina-
tion, being in the degree to which sexual dimorphism was
carried. In species No, 1 the elytra of the males were unex-
panded and female-like, while there was not much difierence
in this respect between the sexes of Nos. 7 and 9. In Nos. 4
and 6 the males were dimorphic, the elytra of some being
female-like and of others highly expanded and of a peculiar
shape, similar in both forms. In all the other species the
males possessed the usual broad elytra, of a very uniform
shape, but differing widely from those of the females. The
relative numbers of the sexes varied in the different species
in an extremely ioteresting way.
Judging from this experience L. (M.) Jemoralis and L. duvi-
vieri were the two dominant species in the locality at the
time when the series was collected.
Method adopted by the African Pohfilid Batozonus
fuligim0su8, kl-uo, in bubyino its spider prey. — the
following note written by Captain Carpenter at Itigi, on
Aug. 22, 1917, was read by Prof. Poulton, who exhibited the
Fossor referred to. The species had been kindly determined
by Mr. Rowland E. Turner as the var. sepvlchralis, Sm., of
the above-named very variable species.
" Here are some notes on a common-looking blue-black
Fompihd with orange antennae found at work on June 12th,
which you will find in the box. She was excavating her
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burrow for the spider which had been caught and paralysed
and \&y close by — a very procryptic species of cmioiudy
irregular shape, and dead grass colour. The Pompilid bur-
rowed ae quickly ae any Fosaor I have seen (except Bemba),
in the ueual way, but, unlike others, FompHidae and SpHegidae,
absolutely quietly, without any of the excited buzzing so
commonly heard. Once an ant approached, and the Pompilid
walked towards it in a threatening manner, with the tip of
the abdomen curved strongly downwards and forwaida.
When the burrow was finished she dragged the spider down,
going down first, backwards, and then stood in the bole and
pulled the loose earth down with her anterior legs. Then
she exhibited a method new to me : other Pompilids that I
have seen ram down the earth firmly with steady pressure
exerted by the tip of the abdomen, emitting the while a
shrill buzz. (It is noteworthy that in the use of the abdomen
they all agree; differing from Sphegids, which employ the
front of the head as a ram.) This one, however, threw her
whole body into such quick, shuddering vibrations that her
outhne became blurred, and one heard a succession o£ quick
raps on the ground reminding one of a pneumatic riveting
machine at work ! The whole process was extremely rapid.
This Pompilid therefore showed two points in her metJiod
which I had not seen before : first, the complete silence in
which she worked ; secondly, the rapid hammering with the
tip of the abdomen."
Spid£r attacking the fierce Ponekihe ant Mboaponbka
FOBTENs, F, — Prof. PouLTON exhibited a spider and its prey
taken at Itigi by Captain Carpenter on Aug. 21, 1917. The
specimens were accompanied by the note : " Spider seen
coming out of a nest of Megaponera bearing one feebly strug-
gling, upside down in its fangs. Caught in a box the spidei
settled down to feed on the ant." The small size of the
spider as compared with ite victim, the redoubtable Termite-
raider, was remarkable.
An East African dbaoonfly-uke Asilid fly op the
GENUS LASiocNBHua. — Prof. PouLTON exhibited the specimen
referred to as follows by Captain Carpenter in a letter of
Aug. 22, 1917: "On June 16 [at Itigi] I caught a curious
l„y,l,Ajh,COOl^lc
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skDder-bodied Asilid of tmusual colouring, which, Testing on
a grass stem, looked rather Uke a very small Agrionine."
The lesembUnce, which waa sufficiently strong in the set
specimen, waa probably closer in the attitude of rest. Di.
G. A. E. Maishall had kindly helped in tracking the species,
which was evidently near L. bigeru, Lw.
Mr. C. 0. Farquhaeson's investigations into the ufb-
HlSTORtES OF S. NlGERIAN LyCAENIDAE. — Prof. POULTON
said that for some years Mr. Farquharson bad corresponded
with him on the above subject, and he hoped, as soon as a
few points had been cleared up, to bring forward a detailed
account of the discoveries. In the meantime the results
were so interesting and important that it was desirable to
m&ke a brief announcement of Mr. Farquharson 'a main
conclusions.
(1) The Liptenine Ltfcaenidae of the genera BewiUonia,
Indoptk, Terabmeura and the genus to which " Epitola "
honoriua, F., belongs, possess hairy Lymantriid-bke larvae
which feed on the delicate filmy lichen encrusting the bark
of certain forest trees infested by Cremaslogaster ante. The
larvae are not molested even when walking in the regular
ant- track.
(2) Lachnocnema larvae not only feed on Jassidae, as shown
by Mr. W. A. Lambom (Trans. Ent. Soc., 1914, pp. 470, 471),
but also on the secretions of immature ant- tended Membracids,
as well as on insect food.
(3) Two species of Argiolaiu feed, as larvae, on the flowers
of a parasitic ant-infested LoratUhus. The Rev. E. St. Aubyn
Rogers has also recently suggested that a parasitic creeper,
probably a Loranthus, is the food-plant of a fine species of
Argiolaus in late German East Africa. Pupae were on several
occasions found attached to the plant.
(4) Pilodeudorvc camerona, Plotz, P. diyUu3, Hew., and
Lycaenesthea musagetes. Holt, (kindly determined by Mr. Q.
T. Bethone-Baker) feed, as larvae, on the ant-infested Sowers
of a species of PUrocarpvs.
(5) Quite recently, on Sept. 10 last, the pupae of Cato-
chrysops ■parsiman, F., were found in large numbers, together
with mature larvae, in the subterranean nest of Camjxmotw
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maculatus, F., var. m^nocnemU, Santschi, at Agege, 16
miles N. of Lagos. The ants' nest had been formed in t
disused part of a termitarium. This discovery confimied
Dr. Guy A, K. MarshaU's conviction expressed in the follow-
ing letter of Oct. 28, 1917 : " I fancy all the larger African
Catochrysvps will prove to be Myrinscophilens. I obtained
eggs of C. patricia. Trim., C. celaeus, Ciam., C. imuAwxi,
Trim., and C, peculiaris, Rogen., in Salisbury, all of them
laid on the buds of a species of Basil [Ocimum) ; but I could
never get the larvae beyond the first stage : they always
died or disappeared. liater Dr. Brauns of Willowmore, Cape
Colony, sent me a specimen of C. ptUrida bred in May, 1899,
from a larva found at BothaviUe, Orange River Colony, in
the nest of Camponotus maadaivs. One of these ante and
an example of the larva may be seen beside the series of
palricia^ the British Museum.
" I believe it will be found that the ants carry off the first
stage larvae of the larger African species of Catockrysops, and
that they pass their whole time in the nests."
An observation which explains why the attacks of
besds on butterflies abe rarely witnessed.— piof
PouLTON read the following extract from a letter received
from Captain G. D. H. Carpenter, and written Aug. 31, 1917.
from Itigi : —
" There is a common black bird here which I should thbk
must be a kind of Shrike. It looks like a Drongo, but has
not the curled tail feathers, and instead of sitting conspicuously
on a tree and catching things in the air, it skulks about (always
in pairs) among thick clumps of bush. It has a harsh ciy,
which is answered by the other one of the pair.
" I noticed from day to day that a pair usually haunts the
same clump of bush, and as they are quite unafraid, I spent
some time a few days ago (on Aug. 23rd) watching to see if
they would catch any of the butterflies which were coming
to the flowers of the bush — Belenois, Teracolus, Pinacopteryx,
and a few Lycaenids. Both birds were in the bush, and I
was watching one when I saw out of the comer of my eye
that the other had come quite to the edge, and suddenly 1
heard a fluttering and the quick snap of a beak, and, looking
l„y,l,Ajl^,COOl^lC
( Mi )
in that direction, saw quite clearly sticking out from one
side of the bird's beak a portion of a Bdeiwis wing, violently
vibrating — to be almost immediately swallowed : however, I
eaw enough to be quite certain. What particularly appealed
to me was the fact that a casual observer (such as I have
hitherto been about birds eating butterflies) would never have
seen this : it had to be looked for. It's exactly as Swynnerton
said in his paper in the Proc. Ent. Soc., and what Trimen
said (' neglect of well-directed and sustained observation ').
I don't in the least wonder that Selous had never seen a bird
eat a butterfly, for he had quite possibly never especially
looked for it.
" Of course I returned to that spot several times again,
but either the birds weren't there, or they were hunting on
the ground, or it was dull and there weren't many butterflies
about, and also the blossoms began to fade, for I didn't see any
more butterflies eaten. But the birds are abundant, so I live
in hope. I feel so much that it's ' up to us ' to produce the
evidence which we know is there and only wants producing ! " •
Dr. G. A. K. Marshall had written concerning the species
of bird : " From his description of the bird and its habits
I can have no doubt that it is the Black Cuckoo Shrike
{Campophaga nigra, Vieill.), a bird I knew well in Mashonalaud,
thongh it was not particularly common there."
A SUGGESTED EXPLANATION OP THE OCCURRENCE OF WET-
SEASON FORUS OP BUTTERFLIES IN THE HIDST OF THE DRY
SEASON. — Prof. PouLTON Bsid that the suggestion contained
in the following extract from a letter from the Rev. K. St.
AuBTN Rogers was of great interest, and offered a probable
explanation of many puzzling irregularities. The letter was
dated Aug. 13, 1917, from late German East Africa. The
precise locality was not given, but Mr. Rogers probably wrot«
from Kongwa on the Central Railway.
abont at the edge of «■ , _
■ettled in the grass, as it iras temporarily overcast. The biid had not
noticed them, vhen one suddenly got up, alarmed at his approach ; and
he darted at it, but only just mi»«d it, and it flew sway. I think it
was a BdtTuju, but couidn't be certain."
u,y,l,/eJh,G00glc
( Mr )
" I have some hopes that some of the thiogs I got in the
vet Beason may be of use to you, and many of the moie
intereBting speciea seem to be confined to the wet season and
the early part of the dry weather.
" 1 venture to think that the whole question of seasonal
forms requires a good deal more investigation. It is not
nearly so simple as might be supposed, being especially com-
plicated by the fact that some plants, e. g. the food-plants of
Belenois and some species of Teramltis, make growth through-
out the dry weather. The food-plant of the common B.
severina, Oram., and B. meseatina. Cram, (the latter greatly
preponderating at present), starts into growth at the b^in-
ning of the dry weather, and thereupon many {Jants are
completely stripped of their leaves by the larvae of the above
speciea and, after a short time, start into growth again, so
that there aie always plants with young leaves on them
during the dry weather at any rate up to date. Now it is
a curious fact that many fresh specimens of B. meserUina, at
the present time after 3 months' absolute drought, are to the
best of my knowledge wet-season forms. I caught a pair
tn cop. last week on purpose to illustrate this.
" I feel sure that Bell's view * (that the seasonal forms are
produced by the state of the food-plant) is correct in cases
of this kind, but then how about PrecU of which dry forms
began to prevail a month before the end of the rains? I
hope that my collection may throw some light on this intenat-
ing question. Just here, in spite of the drought, growth does
not altogether cease, e. g. the millet in the native gardens
sprouts freely after the crop is reaped, and is later used for
grazing. No doubt this is due to the proximity of the range
of high hills to the north, as though all the torrent beds (and
there is nothing else on this side) have been dry for montiis,
yet no doubt a good deal of water must percolate through
underneath. The hills on this side of the range are veiy
steep and stony, and even drier than the plain where we are.
" I wish I were in a position to undertake breeding here,
but it is quite impossible.
" Teraxilus «m, Klug., and T. ceUmene, Luc, I have seen
• Ent. Ho. Hkg., 1M6, p. 121.
u,y,l,7rJM,G00glc
( 1" )
U)ing on the food-plants of B. mesenttna, and T. eupompe.
King., and T. caUidia, Gr.-Sm., on aoothet plant.
. " They are still quite common, aa are many other speciee
of Teracolus, bat they are all old British East African frienda :
still I catch a dry-season form occasionally. T. celimene is
commoner now than in the lains. I have caught several
females of this, as I believe they are not abondant in collec-
tions. They are very like B. teverina ^ on the ving and
mnst be frequently overlooked."
AnOTBEB tNDEPBNDBNT OBaSBVATION OF THE " FAIJBB
HEAD " IN BUTTEBFLiES. — Prof. PorLTON Said that his atten-
tion had be«n directed by Mr. E. A. Elliott to the following
extract from J. Sibree's " Naturalist in Madagascar " (London,
1916, p. 254), referring probably to a Lycaenid butterfly :
" While staying near the forest I was several times struck by
the cnrious formation of the wings of one of the smaller
species of butterfly. The insect in question is of plain incon-
spictious colouring, chieSy shades of brown, and when at rest
sita with the wings erect. The noticeable point is that there
are several strongly-marked and dark-tinted processes from
the hinder part of the wings, which resemble the head, eyes
and antennae of a butterfly, so that when at i«st it is very
difficult to say which is the head and which is the tail of the
insect. The tail markings and points are so much more
strongly emphasized than the actual bead and antennae, that
it is only when the wings slightly open that one is undeceived.
. . . May not the reason of this mimicry of the head by the
tail be of some service in directing the attention of birds and
other enemies to the less vital part of the butterfly's struc-
ture ? It ia evident that the hinder portion of the wings
might be snapped at and broken off, and yet no serious injury
be done to the vital parte of the insect."
This observation added another to the long list recorded
in our Proceedings for 1906, p. lii, to which must be also
added the note by Mr. T. R. Bell in Ent. Mo. Mag., 1906,
p. 128. Prof. Poulton said that there was yet another recent
detailed observation by a Danish naturalist to which he hoped
to direct attention aa soon as it was published.
W. A. I^ahborn's obbkrvations on the mimetic females
?BOC. ENT. eOC. LOND., n-iv. 1917 E
iM:,i,A J 1^, Google
( ll" )
OP Papilio dabdanus, Brown, in late Gebuan East
Africa. — Prof, Poulton said that the following letter was
written by Mr. Lambom from Tanga, on Aug. 10, 1917. It
was of great interest to receive the impressions of so keen an
observer upon the resemblances in life between mimic and
model and between the mimics themselves.
" Ton will have already learnt that I obtained seven
eggs from a hippocoon female of dardanus, captured at Dar-
es-Salaam. The larvae did well and I have the seven pupae,
the imagoB from which may now come out at any time. One
pupa is malformed, but not so badly that the perfect insect
is likely to be entirely valueless. 1 e^cpect the females to be
all hippocoon, for I have as yet seen no other forms along
this coast, and Amauris niavius dotninicatnis. Trim., is more
abundant than I ever saw it on the West Coast, and far more
so than any of the other models. In anticipation of your
probable wish to study the latter with the eastem forms of
hippocoon I have secured a long series (56). These were taken
in association with another Amauris, oMea, Boisd., I think, all
at one spot, mostly at rest on undergrowth beneath a Mango
tree, and within the space of an hour in the early moming.
" Since I last wrote I have seen five more hippocoon females
and have taken three, the last this morning and now in cap-
tivity. I am still not sufficiently familiar with the East
Coast hippocoon and its model to be quite sure what the
species is when on the wing. In this connexion the follow-
ing incident occurred a day or two ago. I saw a hippocoon
feeding at a flower and hovering aa these Fapilios always
then do ; Amau-ris, with the security probably bom of its
protected qualities, always settles to feed. I missed the
Papilio and it flew oS, with me in hot pursuit at midday
over some 200 yaids of burning saudy plain, and it then
escaped over some bushes. But on the other side I saw it
again, as I thought, and pursuing it further took it, and
then discovered it to be a dominicanus. Keturning to tha
bushes I beat out the hippocoon, but again failed to secure
it, for it rose high into the air at once, aa is usual when
alarmed.
" I took recently my second Bypolimnas mambara. Ward,
ll,y,l,/eJh,G00t5lc
( Ixvii )
ft tnale, flitting round a Mango tree in flower, and until I tiad
it in the net, was convinced ttiat I had been watching a
hippoemm.* But my experience of these eastern forms is of
course so limited.
" 18tk August. — The seven imagos are now out, but only
two are females and both hippocoon, one badly malformed.
However, I have put it in the box ready for postage when
an opportunity offers. The other female is a very fine
specimen.
" So far I have not succeeded in obtaining any more eggs,
though I have seen several more hippocoott females, and two
female forms, a cenea, StoU, and a Iropfwniua, Westw., feeding
on flowers high up on a bank and quite out of my reach.
That is the first time I have seen trophoniua alive."
FUBTHBR OBSERVATIONS ON APBICAN HeSPERIDAE OF THE
GENUS SaRANQESA BESTING IN HOLES IN THE GROUND.— Prof.
PoiTLTON said that he had just received from Captain R. S.
Wilson the following observations on the habits of S. elimituUa,
Holl., and S. plisUmicus, Pltitz, or ladius, Mab., in the Nuba
Hountains Province of the Sudan. Among the specimens
sent to Oxford by Capt. Wilson were 2 diminala, taken
Apr. 13, 1917, at Dilling, one of them bearing the note re-
ferred to below, and 14taken the next day in the same locality;
also 1 takcD June 14, 1917, at Talodi. This latt«r, a worn
apecimen, bore the label "flowering plants," probably indi-
cating that the Hesperid was not altogether restricted to holes
and deep shade.
StTi- 14, 1917. TaUtdi.
" He Sarangesa diminata, I was most interested to receive
the proof [Proc. Ent. Soc, 1916, pp. cxxis-cxxxiij you sent.
My note means 'in shade: also in hole in ground.' My
experience of this skipper, as regards its fondness for holes,
ia exactly the same as that recorded in the proof. I used
when at Dilling this year generally to go out collecting for
'See also Trans. Ent. Soc., IBOS, p. 497, and description of Plate
X.JLV1, fig. 3, for the Rev. K. St. Aubyn Rogers' observations on the
living H. utambara. Looking at a spacimea in tbo cabinet it would
be *uppo8ed that the bright orange-brown of both surfaces, especially
the under, is likely to prevent a close mimetic likeness, yet both Mr.
Rogers and Hr. lAmbom observed tJiat the raaomblance ia strong.
iM:,i,A J 1^, Google
( Ixriii )
an hour before BunBet armed with a net and Btick, and got
most of my captures by beating the grasa clumps and bushes
on which the butterflies had settled for the night. On the
first occasion when I saw S. eliminata there, the sun had
just Bet, but had sunk behind a small jebel some few minutes
before, and it was just getting dark. I caught sight of two
or three of the skippers flitting about at the entrance of an
old porcupine burrow (I had often seen butterflies in similar
situations before and recognised them as some sort of skipper).
I then rattled my stick round the sides of the hole, and they
came out in extraordinary numbers and I caught several.
I did this Ave or six times and some came out each time.
If left alone they flitted about for a short time at the entrance
and then went in again. They always settled with wings
fully extended and resting flat against the aides of the burrow.
I often passed this particular burrow afterwards and almost
invariably found some skippers present. The hole was partly
overhung by a bush which did not make it any easier to
capture the skippers. I found them afterwards at Dilling in
mare recently occupied porcupine burrows, and also under
the roots of large trees along the khor, which baa high banks,
where the soil had been washed away and left dark hoUows
amongst the roots, and also in cavities in the khoi sides, but
always on the west aide (the khor here runs N. and S.), I
presume so that the sun should not disturb them in the
afternoon by shining into the hollows, as it would have done
if they had been on the east bank. As regards S. plisloniau,
FiOtz, if this is a black skipper with small whitish transparent
spote on the fore-wing as I think it is,* I found this first at
Dilling in November, 1904, and also at Tira Mandi in the
same month, and in each case recorded that they were always
taken in shade. Those at the latter place I generally took
* Tba examplea lefeTTsd to by Captain Wilson aie ouned 3. flitlo-
nieut in Tnuu. Eat. Soc., 1916, p. 284. Thero are 6 ■pecimeoa at
Oxford in the coHection made bj Him — I from Dillinp, Nov. 15, 1904
(given as 1906 by a clerical error in the paper quoted above); 4 from
Tm. Mandi, Nov. S3-26, 1904. In the collection of ttie Britiab MuBema
flulonicut stands under the genus Bretia, and Captain nelson's speci-
menB are the eame as a series (indudiog an example from the Atban)
named Sarangaa ladim, Hab. The spedee of tbia geonfl will never
be satisfactorily determined until the types have b«en seen and tbs
•tmctuial ehuacters worked out.
Ij.y.l.AjL^yCOO'^lc
( liii )
inside my ' rakuba,' a temporaiy graaB-bnilt shelter which
WBs erected in the shade of some large trees locally called
Tameiza, of the Ficus tycomonts, L., t)rpe. Since then I have
seen both these species resting inside hollow trees snch as
Adamonia digitata (locally called Tebeldi) which grow to a
vast size and are nearly always hoUow,"
Emission of fluid fkom the antennae op Acbaza
QtJiKiNA, F. — Piof. PouLTON exhibited a male example of
A. quirina, captured in Sierra Leone probably in the neigh-
bourhood of Freetown, June 18, 1917, by Ueut. P. A. Buxton,
R.A.M.C., who had made the observations tecorded in the
following letter, written on June 20.
" I am circumnavigating most of the known world on my
way to the Euphrates. I shall post to you a small Acraea
which I took recently. When I pinched it in the net I dis-
tinctly saw drops of yellowish fluid appear, one on the club
of each antenna— drops about the size of the head of a
domestic pin. This seemed interesting, so I took the thing
out of the net, pinched it again, and got the drops again,
but much smaller. Third attempt, no drops. This is the
only time I observed it, though I got lots of small Acraeaa
that day and subsequently, and kept my eyes open, of course.
Is it an old observation ! "
Prof. Poulton said that the fact had been observed by
Dr. G. A. K. Marahall in Planetna aganice. Hew., and Acraea
terptichore, L., f. rottgeli, Gu6r. (sereno, F., buxUmi, Hew.).
See Trans. Ent. Soc, 1902, pp. 413 and 323 respectively.
Labva of Saw-fly with " Palisade. " — The Rev. F. D.
MoRiCE exhibited a photograph of a saw-fly larva with the
epidiascope ami described it as follows : —
" I show a print from a negative already exhibited at a
previous meeting (Oct. 3). It represents a young (living)
larva of the sawfly Lygaeonemalw compressicornu, F., feeding
in the middle of a poplar-leaf, and surrounded with a sort of
' stockade,' or rather circle of Uttle glassy nearly equidistant
' pillars,' which are believed in some way to protect it, but
against what sort of attacks has never been clearly made
out (Plate C, fig. 2). In another photograph I show one
of the pillars separately, at a magnification of 20 diameters
iM:,i,A J 1^, Google
< Ixx )
(Plato C, fig. 3). It. is then seen to be a compact pile of
little transparent bubbles, which must have hardened im-
mediatoly after being ejected by the insect. It is, however,
though hardened, excessively fragile; and it is ditBcult to
believe that it could offer any physical obstruction to the
approach of any imaginable assailant of the feeding larva.
Besides which, the pillars stand far enough one from the
other to allow of a free passage between them. It has been
suggested to me that the so-called ' stockade ' has some
resemblance to various objects (fungoid growths, stalked eggs,
etc., etc.), and that this deceptive mimicry might serve as a
protection to the larva against some particular enemy. At
any rate, for whatever reason, the formation of this stockade
seems to be considered by the larva as an indispensable pre-
liminary to feeding. Miss E. Chawner tolls me that she has
tried the experiment of brushing it off, and that the larva
then seems considerably discomposed, leaves off feeding, and
presently moves away elsewhere, and forms a new circle
before it ventures to recommence its meal. It is curious also
that when the hole eaton in the leaf reaches a certain size,
the larva reaches out and constructs a similar circle of pillars
on the other side of the leaf, and that when it has reached a
certain stage in its own growth it abandons its defensive
ayatom altogether, passes to the edge of the leaf and feeds
there, just like the larvae of other Nematids, without any
visible precautions against disturbance from without. This
remarkable instinct, which apparently is peculiar to this
one species, was first observed and fully described by the
Dutch entomologist v. VoUenhofen (a former Hon. Fellow of
our Society)."
The exhibitor then read extracts from an Knglish transla-
tion (by Mr. May) of v. Vollenhofen'a memoir on the subject
which appeared in vol. xx of the Zoologist (1862). The
insect is there called Netnalus vaUaior, n.sp., its identity with
Fabricius's compresstcomis not having been realized. It is
rather a pity, though it cannot be helped, that the Law of
Priority reduces that which in itself is certainly the more
suggestive and euphonious of these names to the rank of a
mere synonym.
n,g,t,7.dh,G00glc
{ Ixxi )
Tfll COCOOH OF DlCRANUBA VINULA. — Dt. ChAPUAN ei-
hibit«d some cocoons of Dicranura vinida that happened to
show much more plainly than is usually the case the diSeient
and thinner texture where the moth is to emerge. In de-
scribing them he observed that Mr. Latter, Ent. Trans,, 1895,
p. 399, says : " If a cocoon is examined from the inner side
while held towards the light, it will at once be seeo that the
walls are not of unifonn thickness, but that thinner patches
occur here and there. I have examined over a hundred
cocoons in this way and invariably found one of these thinner
areas at the anterior end opposite the head of the pupa."
As viewed from the outside the thin portion of the cocoon
is seldom very evident. Of those exhibited, six, made on
the inner surface of the coik, show these thin places most
obviously as darker patches— darker owing to having less of
the cork material used in them — and these correspond in form
and position to the portion of cocoon that the imago makes
use of for emergence. Two cocoons on the outer surface of
the cork show similar patches, but they are not seen till
looked for. Other cocoons made by the same lot of larvae
are equally difierent, only those on the inner surface of the
cork showing with such marked distinctness the thin places.
There seem to be no thin places at any other part of these
cocoons. As I have never noted these patches before to be
so evident, it seems that this particular cork surface leads to
the structure being so visibly differentiated."
The Generic Naue Tinea.— Mr. Stanley Edwards
asked the following questions : —
" Is the President aware that in the Collections at South
Kensington the generic name Tinea of Linnaeus is employed
for two difierent genera in two diSerent families of the
Lepidoptera ? "
" What steps does he propose to take as to this second
usage in a new and unfamiliar sense of an ancient and well-
known name whose application has been undoubted for
generatioiu ? "
The President, Mr. Bethune-Baker and Mr. Neave
spoke on the subject, and eventually it was resolved, on the
motion of Mr. Bethume- Bakes, seconded by Mr. Dur&aiit,
iM-,i,A J h, Google
{ Uxu )
that Sit Geoboe Hampson should be invited to take part in
the diBcusBion of the question at the December meeting.
Wednesday, November 21st, 1917.
Dr. C. J. Oahan, M.A., D.Sc., President, in the Chair,
domination of Officers.
The following Fellows were nominated as Officers and
Members of Council for the eosuing year : —
President, Dt. C. J. Qahan, M.A., D.Sc. Treasurer, W. G.
Sheldon. Secretaries, Conim, Jaues J. Walkeb, M.A., R.N.,
F.L.8.; Scv. George Wheeler, H.A., F.Z.S. Zt&ronan,
Geoeoe Charles Champion, P.Z.8., A.L.S, Other Members
of Council, A. W. Bacot; E. C. Bedwell; K. G. Blaik;
Dr. T. A. Chapman, M.D., F.Z.8.; W. C. Ceawley, B.A.;
H. WiLLOUOBBY Ellis, F.Z.S.; Dr. H. Eltrdighau, M.A.,
D.Sc, F.Z.S. ; J. C. F. Fryer, M.A. ; A. Huoh Jones ; Rev,
F. D. MoRiCE, M.A.; S. A. Neave, M.A., B.Sc., F.Z.8.;
Herbert E. Paqe.
Eiedion of Fellows.
Mesflre. Frederick Walter Cocks, 26, Crown Street,
Beading, and Willuh Gerald Hakding, St. Hugh's School,
Bickley, Kent, were elected Fellows of the Society.
Exhibitions.
Bbitise Chkysophanus dispab, vab. kdtilus. — Oapt.
PuREFOY exhibited a series of British C. dispar, var. rutUus,
whose ancestors had come from the neighbourhood of Beriin
before the war. They were now firmly established in a marsh
in South Ireland, into which the food-plant, Rumex hj/dro-
lapathum, had also been introduced. The series exhibited
included specimens taken in 1915, 1916 and 1917, and showed
a remarkable increase in size during this short period, the
<; S reaching 60 mm. and one of the $ $ 62 mm. The depth
of colouring on the upper side was also much intensified.
There was still, however, much less blue on the under surface.
Ij.y.l.Ajl^yCOO'^lc
( Izxiii )
and the orange band vae also conaiderabiy less biiUiuit, and
less continuous.
Mr. Kaye enqniied whether the larvae would feed on
other docks, and Gapt. Purefot replied that they would do
BO, bat that other docks were apt to turn brown after flower-
ing, thus depriving the larvae of nourishment. They would
not feed on sorrel.
Mr. Sheldon remarked that the var. rvtilvt must have
alternative food-plantB on the Continent, as he had found it
not only in the Danube Marshes (where R. hydrolajKUhum
grew) in the vicinity of Buda Pesth, but on many of the hills
around this town far out of the range of this plant. In the
neighbourhood of Saiepta also, it was found in the small
cross gullies which had a small stream at the bottom emptying
into the Volga some miles away. In each of these localities,
although several species of dock were found, the usual food-
plant did not grow.
A question having been asked as to whether C. 'dispar had
ever been found out of England, the Bev. 0. Whxslsr re-
minded the Society that it had been discovered last year in
Holland, and exhibited by the Hon. N. C. Rothschild side
by side with English specimens. Some doubt was expressed
later as to whether the Dutch Bpecimens were really indis-
tinguishable from the English.
Pupation op Dmscus iuboinalis. — Mr. Hugh Main
showed a series of lantern slides illustrating the methods he
had successfully adopted for observing the larva of Dytiacut
marginalis " digging itself in " for pupation, also the pupa in
its cell, the disclosure of the imago, and the escape of the latter
from the cell. He said he had exhibited on October 18, 1916,
a clump of earth containing a pupation cell of D. margintdu
which he had found near the margin of a pond in Epping
Forest. Other cells subsequently found agreed in being
dtuat«d along the angle between a horizontal shelf of earth
and the vertical bank rising behind it. It was only necessary
to reproduce such conditions in confinement to get the larva
to form its pupation cell, and it was not difficult so to arrange
matters that the work was done at a convenient time for
observation. A small oblong glasa aquarium was used, on
iM:,i,A J 1^, Google
{ Iniv )
the bottom of which was placed a layer of earth about two
incheB deep, and a bank of earth three or four inches high and
about the same thicknesa was made at one end, the eaith
being damp and tightly pressed. At the other end a small
shallow glass vessel was placed, nearly full of water and con-
taining some water weeds and a larva. The larva was fed on
earthworms and when full fed it easily climbed out of the
water and soon discovered the suitably arranged bank. The
larva could be hindered fiom commencing its work for a few
hours by placing a glass plate in froot of the bank, and when
this was removed the larva at once started operations. Full-
fed larvae were found in July, August, and September. They
are very active on leaving the water, sometimes travelling ten
feet or more from a pond before Ending a suitable rate for
pupation.
In making the cell the larva removes in its jaws a small
mass of earth from the face of the bank, turns round, and
then, opening ite jaws, it pushes out the earth with its anterior
legs, depositing it to one side or behind it, thus gradually
making a cavity in the bank and forming a little curved wall
against the face of the bank. The inner surface of the wall
is made quite smooth by pressure of the flat head, and the
globular cavity which is also smoothed o2 is finally closed in
by the mass of earth removed from the interior. The larva
takes about six to eight hours to dig itself in out of sight,
keeping at work nearly cootinuously.
The pupa was disclosed about a week after the larva had
disappeared from view. It may be easily seen on carefully
removing the exterior wail of the cell or on cutting the earth
away and opening up the cavity from the roof. It is per-
fectly white at first, and only the eyes show any darkening
for a time. It normally rests with its dorsal surface upper-
most, lying across the ceil, supported only by the prothoradc
rim, which bears a number of short stiff bristles, and the two
fringed processes at the tail end. The whole of the body is
thus out of contact with the surrounding earth except the
two small supporting areas at the anterior and posterior
extremities. If disturbed, however, the pnpa gives a wrij^
causing it to fall over on its back, but very soon another
Ij.y.l.AjL.yCOO'^lL'
PLATE D.
Dtjtiacva maryinalis digging itself it
1 to 6. Successive stages of operatione.
All Blightly lesB than J natural size.
PLATE E.
DyliacUi tnargiTtalis. Pupation.
1. Esterior of pupation chamber.
2. Showing opening mode by imago for exit.
3. Pnpa shortly after emergence-
4. Pupa in nonnal resting position, lateral view.
5. „ „ „ anterior view.
6. „ „ „ posterior view.
All slightly less than } natural size.
PLATE F.
Dytiacut tnarginalis.
1. Male, just emerged from pupal akin.
2. Hale, 24 hours later.
3. Female, just emerged from pupal skin.
4. Female, 24 hours later.
6. Artificial pond and bank for the observation of the larva
preparing for pupation, etc.
1 to 4, slightly lees than { natural size; 5, about } natural size.
n,g,t,7.dh,G00glc
n,g,t,7.dhvG00glc
Proc. Ent. Soc. Lottd., 1917.
Hugh Main, Phata. ^^'f' T<^' ^"i- <^'- '■'■'•
Pupation of DvTiSCUS MARGINALIS. : , CoO^lc
n,g,t,7.dh,G00glc
Proc. Eni. Soc. Lond.. 1917.
Hugh Main, Phola. Half-Tom Eng. Co. Ltd.
Pupation of Dytiscus marginalis. ..CoOqIl'
n,g,t,7.dh,G00glc
( Ihv )
wriggle raises it again into its usual resting position. The
enipty laTval skin is pressed flat against the wall of the cell
where the pupa cannot come in contact with it. After the
exit of the imago the larval skin can be removed, spread out
under water and the longitudinal dorsal slit observed through
which the pupa had emei^d.
The pupae observed remained from two to three weeks
before the emergence of the beetle, only the legs and some
of the terminal segments of the abdomen becoming dark.
The thorax and elytra of the imago are soft and quite white
or cream-coloured at first, but in twenty-four hours they have
almost reached their usual coloration. The male is easily
distinguished in the pupal stage by the characteristic appear-
ance of the enlarged joints of the anterioi tarsi. The insects
finally escaped through a round hole made in the exterior
wall towards the top of the dome-shaped interior and above
the part of the wall made by the deposition of the earth from
the interior of the cavity (Plates D, E, F, and 0, fig. 1).
Wednesday, December 6th, 1917.
Dr. C. J. Gahan, M.A., D.Sc., President, in the Chair.
Nomination of Officers aiid Council.
The names of the Fellows nominated by the Council as
Officers and Council for the following year were read for the
second time. No other names had been received.
Election of a Feliov}.
Mr, Charles Ogilvie Farquharson, M.A., B.Sc., Govern-
ment Agricultural Dept., Moor Plantation, Ibadan, S. Nigeria,
was elected a Fellow of the Society.
Nomination of an Honorary Felloie.
The name of M. Paul Marchal (France) was announced
for the first time as having been nominated by the Council
for an Honorary Fellowship, in the place of the late Dr. £.
FREV-GESaNER.
n,g,t,7.dh,G00glc
Exhibitions.
AbEBBATIONS of ClEOBlETBID MoTHS PBOH EAST LOTHIAN.
Miss Balpoue exhibited a strikmg abeirstioD of Lareniia
didynuUa, L., tAken by her at Whittingehame, East Lothian,
on July 24, 1917. This specimen (a J), which was rather
laiga and pale, was Temaikable for having a conapicnoos dark
fuscous marking resembling an airow-head, formed by a
wedge-Uke extension, towards the base, of the fsmiliai " twis-
spots " of' the species. She suggested the name cttnagera
a. ab. for this form. She also exhibited a remarkable aber-
ration of Mdanippe fiwiwUa, L., taken by Mrs. Meldola near
Dunbar, East Lothian, August 12, 1912, and now in Prof.
Meldok's collection in the Hope Department at Oxford.
Sfideb Miuic o? an Amt. — Mr. Donibthobpe exhibited a
large, very ant-Uke, spider together with the ant it mimics,
Potyrhachia schiatacea, Oeis. subsp. rvgidosa, Mayr., which had
been captured and given to him by Mr. Herew&rd Dollman,
F.E.S., at Mwengwa, N.W. Rhodesia, August 13, 1917. Also
a small British beetle, Brachonyx pineti, which he had taken
not uncommonly at Barton Mills, Suffolk, September 9, 1917.
The object of this exhibit was to show the close reaemblance
between the beetle and seeds, and bits of buds from the
young Scots Firs on which the former occurred. Mr. Donia-
thorpe said this might be described as " Camouflage."
Mr. Champion pointed out that it was a very southern
locahty for this Scotch beetle.
Dark Aberration or Hihera pennabu. — Mr, B. E. Greeh
exhibited an unusual variety of Himera pennaria, the front
wings of which were almost entirely covered with dark
plumbeous scales, leaving the veins picked out in bright
fulvous. This form approaches a variety of Smwmot
autumnaria figured by South (" Moths of the British Isles,"
vol. ii, PI. 134, fig. 9).
CoNDmoNs OP Insect Life in Mesopotamia. — ^Ptot
PouLTON said that he had recently received a letter from
Lieut. P. A. Buxton, R.A.M.C, contrasting the conditions
encountered in Mesopotamia with those observed in Africa on
the voyage.
n,g,t,7.dh,G00glc
( Izzvii )
" This is a gieat contrast — really deserUd desert most of it
without even camel thorn — and cultivation, gardens, dates, &c. ;
consequently no insects. Ot course they may buck up in
autumn or spring. The laboratory is efficient : I am doing
flies, and have a great time running eveiywheie at my own
sweet will.
" What strikes one here is the extraordinary comfort of life
compared with what it must have been. You light in at
some little desert post and find the fans running, and the
soda bubbling, and the ice-machine clanking I "
Salt (Chlobide op Sodium) probably sodoht by thb
Hesfeotdae. — Prof. Poulton said that he had received the
following letter from Mr. S. A. Neave, referring to the
observations recorded by Mr. C. 0. Farquharaon :
" Nov. 2&.im.
" As regards the note about the peculiar habit of Bhopalo-
campta foreatan, ou p. Ixxx of the 1916 Proc. Ent. Soc, I
have seen this exactly as described both in this species, in
R. pisislratus, ¥., and in at least one of the common Pamaras,
I think P. fatwUus, HopS. I had not«d in P. Z. S., 1910,
p. 85,* that R. foreatan was much attracted by perspiration,
but had not at that time seen the moisture extruded from
the abdomen. In my case there could have been no question
of ink, the back of my hand oi arm being the site selected,
though the fluid absorbed may have been mixed with the
perspiration. The extruded fluid seems to be usually colour-
less, but in one case, I am not quite sure in which species,
but I think R. pisistratvs, it was of a miUcy appearance."
Mr. Neave had also informed Prof. Poulton that the
skippers would often settle upon the shirt-cufi and there act
as above described. Now besides water the chief constituent
of perapiration, and the only one likely to be of value to the
insect, was salt, while the observed behaviour strongly sug-
gest«d that it was some soluble substance like salt that was
Bought. Bapid evaporation in the sun would leave salt in
concentrated solution or even dry on the skin or in parts of
DDB a dtting OT
a attracted by
n,g,t,7.dh,G00glc
{ Ixzviii )
the clothes in contact with it and exposed to the air, and
the discharge and re-absorption of a watery fluid upon such
surfaces would certainly lead to the ingestion of salt. Mr.
Farquhareon'a observation now probably received its true
interpretation, the arm of an armchair in the tropics being just
the place where salt is likely to be produced by the evapora-
tion of human cutaneous excretions. The dried ink observed
to be moistened by a skipper may have contained sodium
chloride or some other salt which took its place in the insect's
estimation, although it is perhaps more likely that some
gummy or sugary substance was sought.
[While the above note was being brought forward a letter
from Mr. C. 0. Farquhateon, written Nov. 16, was on its way,
containing in the following passage the very same suggestion
as to the object of the habit : —
" By the way a medical man whom I met, who seemed a
fairly observant man, told me that in his district a certain
small Lycaenid was a nuisance (! !} from its habit of settling
on one's hands and arms to quench its thirst on perspiration.
I know of a smalt Hymenopteron [almost certainly the sting-
less bee Melipona] which has the same habit to a most annoy-
ing degree, but why I mention this is that it recalled to me
my theory of the object of Rhojtalocampta forestan when it
settled on the arm of the chair, exuded the drops of clear
liquid and finally drank them up again. The drops might
easily have acquired a certain salinity."
It is to be hoped that specimens of the Lycaenid referred to
may be sent home and the species identified. It is possible
that the insect was a Hesperid.
Furthermore, since the note was read I have found that the
habit is fully described by Mr. J. C. Kershaw in his " Butter-
flies of Hongkong," 1907. The following passage on p. 129
refers to Pamara gnitatus, Bremer and Grey : " It was
exceedingly common at Lo-fu-shan. One individual lit on
my hand and remained for at least a minute, bending the tip
of its abdomen as if about to lay an egg, but it deposited
drops of clear liquid which it eagerly sipped with its proboscis.
It exuded several drops in different spots, and immediately
sucked them dry. This and other Hesperids may occasion-
n,g,i,.rjh,G00glc
( luil )
&II7 be obeerved depositmg liquid on leaves and drinloDg
it up." '
Mr. Kerahaw, who has kindly written, tells me that he has
observed the habit in all the species of Pamara described in
his book. This was at Macao, where the species aie on the
wing and have the habit all the year round, but he has also
seen it in the Malay Islands, Australia and the West Indies.
He regards it as specially characteristic of the genus Pamara.
Uia letter continues : " flo doubt when they do this in a hot
climate on man or other animals, they must imbibe a certain
amount of various salts. They will also emit liquid on any
object whence they can re-imbibe it. Very likely they would
get salts from various rocks and stones and earth when they
void the hquid on these, as I have often (hundreds of times]
seen them do.
" The habit can scarcely arise from want of moisture, since
they merely drink again what they have just voided (-f what-
ever salts, etc., may have been taken up by the Uquid from
the material it has been thrown down on), and moreover they
will do this when only a few inches from a stream or pool.
There may be something in common with the habit of cows,
which will often drink farmyard manure water as black as
ink (and of course full of salts) instead of drinking pure water
from the trough close by."]
Mr. E. £. Green had kindly written on the general subject :
" With regard to curious feeding habits of certain butter-
flies, I remember the late Col. Mandeis, on one of our collect-
ing trips in Ceylon, pointing out to me how the butterflies
were settling on his bare arm and hands — apparently imbibing
the moisture therefrom. Although I was in an equally warm
condition, I was not honoured in the same way. No butter-
flies showed any attraction for my skin. Hsnders told me,
at the time, that they invariably selected him in preference
to others, and he attributed the fact to his having a gouty
tendency. Perhaps you can find out from one of your medical
friends whether the perspiration of a gouty subject contains
any substance that is not present in that of an ordinary
mortal.
" Do you think that the attractive properties of urine (both
u„,,i,Ajh,Googlc J
( tal )
human and bovine) has an; connection with the other facte !
It is moBt noticeable in Ceylon. A fellow collector uaed
systematically to lay a tiap on the sandy margin of a stTeam,
to attract specimens of several species of Cyaniru (Lycaenidae).
And I have seen large numbers of migratory PapUiog (prin-
cipally jason and tered^m) collected round damp spots on the
high road where the cart bullocks bad been micturating. I
I have taken alcibiades under similar conditions; but this is
a very much shyer creature and never associates with tke
common herd.
" I am sorry that I cannot speak with certainty of the
species that were feeding on Manders's arm, but I do not
think that they were Hesferidae. I am almost certain that
L^he drypeiia was one of the visitors, and I think that Pyrameii
cardui was another."
Urea was a constituent of perspirataon, although usually
present in very small quantities, and it might well have been
the attraction in the observation recorded by Hr. Green.
Id the Hesperidae, on the other band, it seemed more probable
that the attraction was salt,
Pafiuo folytbs B0UULU8, Crau., ?ro» the extreme
SOUTH OP India and fbou the mainland opposite Sinoapoee
Isi-AND. — Prof. PouLTON Said that he had recently been given
the opportunity of studying a long series of P. potytet collected
1906-16, by Mr. J. WiUiams Hockin, on the W. slopes of the
Ashamboo Hills, from 6 to 40 miles N.W. of Cape Comorin.
The aeries, which was of great interest in relation to the
forms of the species recorded from Ceylon, was composed as
follows : —
Males-30.
Male-like female (cyrus) — 1.
Aristolochiae-Uke female {polytes) — 12.
Intermediate female — 1.
J7ector-like female (r&mvlus) — 21.
The single intermediate female had the fore-wings of potyles
and the hind-wings of romulus, with reduced red markings
outside the cell and traces of the pale scales of poltftee within
it. The pattern was precisely the same as that of some of
ih,Cooi^lc
( IXHi )
the Bomean females of P. polyUs theseus. Cram., in the Hope
Department.
Of the 12 pdytes females, 4 were the aiichivs form, with
no whit« in the hind-wing cell, 4, with slight indlcatioDB of
white, were intermediate between stichiva and polyta, while
4 were potyteg. The ^1 romuim females varied very greatly
in the development of the red markings in the hind-wing.
Although from the part of the mainland that is very near
to Ceylon, the differences were strongly marked. Thus, in
Ceylon the a/rus female was conuderably commoner than
either mimetic form, while romulvs was generally looked upon
aa only rather commoner than jxAytea. Bat in Mr. Hockin's
series romvlua was nearly twice aa numerous as jiolyta, while
cyrut was apparently much rarer than either. Again the
gtichiua form of polytes, almost unknown in Ceylon, was very
strongly represented in the exhibited series.
The high proportion of rotmdus females was probably to
be accounted for by the relative numbers of the two models;
for H. 3. Ferguson, on p. 446 of his list,* described ariato-
Uxhiae as " common in the low country and the hills," but
hector as " very common " in the same area. Mr. Hockin
agreed, summing up his experiences as follows : " I should
say hector was decidedly more common than arialolochiae,
though the latter would take its place as the second commonest
PapiUo all the year round over the widest area : potyUa would
be third, but several lengths behind."
Prof. Poulton hoped to study the patterns of the Travancore
ariatolochiae, as it was possible that the prevalence of stichiug
and gtichivs-Uke females might be thus explained.
Two of the males, 12 romulus, and all the other female
forma were exhibited to the meeting.
Mr. Hockin had kindly described the character of the inter-
esting locality in which he had collected for so many years : —
" Before saying anything about the range of pclytes, etc.,
I had better give you an account of the different belts of
country. The Ashamboo Hills run parallel with the coast,
with their foot at 16 to 20 miles from it. The hills them-
* " A IM, of the BntterfliM of TraTanoore," in Journ. Bomb. Nat.
Hist. Soc., 1891, p. 132.
PHOC. BUT. SOC. LOND., n.-lV. 1917 F
ihyCOOt^lC
( Izzzii )
selves aaj be about 6 miles wide on tiieir westem faca and
less OD the eastern looking down on llnneTelly, which is very
steep, and I don't think my men ever went to the eastern
face. The top of the ridge is about 6000 ft., of which the
upper half is grass with a few woods scattered about. At
the northern end, however, there is a big block of forest on
a plateau 4600 to 6000 ft., where I fancj PapUio tatmhna
sod buddha, which were unknown to H. S. Ferguson (i&idO,
were found. Bat my men did not often go there, as it was
far away with no inhabitantA but bear, bison, and elephant,
which they feared. The lower slope was tall heavy foieet
(trees 100 to 150 ft. high), with dense bamboo jungle at the
foot. This bamboo jungle (Bamboo mixed with trees such
as Blackwood, Iny and other trees about the height and
habit of EngUeh trees) extends some 6 miles from the foot
of the hill and is succeeded by rounded hills of laterite, 200
to 300 ft. high, with rice-fields in the hollows and grass-fields
OD the higher land. These grass-Gelds have cactus hedges and
hedge-row trees of Palmyra, Jack, Banyan sod Mango and also
leguminous trees grown for leaf manure for the paddy-fields.
Further out in drier country are Tamarind and Neem or
Murgossa trees. On the last belt, next to Tinnevelly and about
7 miles wide, comes a very dry area (similar to Tinnevelly
country) ; where only Palmyra, Tamarind, Neem, and Babul
trees grow with Cactus hedges round grass-fields. These fields
are sometimes planted with peas in the moonsoon, and hone-
grain in October to December in all but the very dry belt.
In the wetter part nearer the hills low scrub jungle grew up
wherever there was no cultivation, and this was felled and
planted with tapioca for two years, then left to grow scrub
again for two or three years. Sometimes hill rice or other
grain was sown for one year on tJie cleared land, or it was
put permanently under Plantain or Banana trees.
" Annual rainfall ISO to 200 inches, over 2000 ft. ; 90 inches
at -foot of hills and 6 miles out, falling to 30 inches in middle
of cultivated tract, and 16 inches in very dry be!t. In Tinne-
velly town, 40 miles E. of Travancore boundary, it is 13
inches. This applies only to the 20 miles at the S. end of
the Ashamboo Hills when I collected.
u,y,i,A"jM,Googlc
( Ixzziii )
" Moat of the PapUioninae and big NymphtUtnae were
found in the lower half of the hills and the bamboo jungle at
their foot. P. hector, arislolochitu and ■polyteii, however, were
plentiful all over the cultivated part with hedgerow trees,
except in the very dry belt. P. entAont'tu and agamemnon
were common in the wetter half of the cultivated belt.
P. diisirmiu is the only one I remember on the higher grass
over 2600 ft. (except perhaps tamlana and bvddha in forest
at that heigKt on the oorthem plat«au). I used constantly
to see P. hector, onale and female, flying in cop., one flying
and carrying the other, whose wings were closed. It is so
difficult to tell Tomvita $ from hector, and polyUs $ from
Ofitltiockiae, when flying that I am afraid I can't add anything
as to their frequency from observation to what the numbers
in the collection will tell you. The same applies to ci/rut $
fuid the males of potytes. As to damage by birds 1 had
better go through all my damaged specimens later and write
you separately about that. The chief preyers on butterflies
were bee-eaters (especially the blue-tailed and chestnut-headed
species) and drongos. We used often to And wings of heOor
under a tree. By the bye, I think I can account for Fer-
guson calling P. momtiM very rare. 1 only once saw it in
September — otherwise only in March to May. The latter is
the unhealthy season when cofFee planters stopped work and
went for a holiday, and no one who was not obliged to
went into the malarious belt at the foot of the hills. Now
we were building a big dam then, which took us four years
to build, and we did most of the work in March to May
because there was no cultivation then and labour was
plentiful."
H. 8. Ferguson {ibid., p. 446) also gave the range of polytes
in nearly the same words as those used for ite models, speak-
ing of it as " common in the low country and up to 2000 ft.
on the hills. The three forms of the female occur."
Prof. Poulton also exhibited a female of the cyrua form,
unfortanately the only example of this sex in a litUe series
sent by Dr. R. Hanltsch in continuation of that described in
our Proceedings for the present year, p. xxx. The specimen
was taken in Johore on July 19, I9I7. Accompanying it
iM-,i,A J h, Google
( IxxziT }
were 3 malee taken oD tlie same day, 1 on July 17, 4 on
June 29, and 3 in May. Dt. Hanitech hoped to aecnie a long
series in August, bo that data would be available foi estimating
the proportions of the two female forms.
Tbb Myhabid Carafhractus cinctus taxen at Oxford
IN 1917. — Prof. PoDLTON read the following communication
by Mr. A. H. Hamm of the Hope Department, and exhibited
examples of the specimens referred to : —
" It ia so long since any record of the captuie of Cara-
phradue dncttis, Hal. {Polynema natant, Lubbock), has been
made that a few notes on its occunence neat Oxfoid may not
be without interest.
" On September 22 last, my friend and colleague Mr. H.
Britten and I weie ' fishing ' for Anopheles larvae and pupae
in small clay-holes in a brick-field in the Kimmeridge Clay on
the west side of Shotover Hill, near Oxfoid. When sorting
over oui captures in the evening we were surprised to find we
had both, quite accidentally, taken C. dnclua. We kept the
specimens alive for a few days in order to observe their mode
of progression in the water. A week later, on September 29,
we again visited the same ponds fully prepared to obtain
more, if possible, of this interesting insect; in this we wete
entirely successful, and between us we took over twenty
individuals of both sexes, all, without exception, obtained
from a very small clay-hole of about two square yards in
area. On October 6 Mr. Britten went alone to the same
Uttle pond and again succeeded in taking s fair number of
both sexes. The next visit was not until October 20, when
I went alone, and found them as before in some numben.
We visited the same pond together, for the last time, on
October 27, but on this occasion, after spending an hour oi
more, we could only find two individuals, one alive and the
other dead.
" On each occasion we took the insects home alive in the
jar containing our other aquatic captures, and then tuined
the contents into a shallow saucer, in order to sort over the
material obtained. It was on one of these occasions that a
number of the Caraphractua were observed to come to the
surface of the water and fly very rapidly from side to side of
Ij.y.l.Ajl^yCOO'^lL'
( IXZXT )
the sancer, just skinuniiig the suriace of the water, in the
manner of a frightened watet-fowl each as the Grebe.
" Their progression under water— comparatively slow, and
effected by means of a series of jerks — was clearly due to
flying rather than swinuning, but they also made use of their
legs, the posterior pair being kicked out behind simultaneously
with the movement of the wings.
" We found that individuals kept in confinement always
preferred to crawl about among the Spirogtfra in prefeienca
to any other water plant. Several were kept alive by Mr,
Britten for about a week.
" One male was observed to make an unsuccessful attempt
to pair with a female on the suriace of the water, by climbing
on to her back. A Corethra larva once seized a male by his
legs and held on for some time until induced to leave go by
means of a camel's-hair brush.
" The small pond or clay-holein which all the examples, except
one, were taken, contained the usual common aquatic insects in
abundance, and the vegetation was also of the usual pond type.
" The occurrence of this species so late in the season is
interesting, as those taken by Lubbock* were obtained in
Angust, and for those recorded by Button f and Enock } the
month is not given."
Mendelun Hebsdipy in relation to selection. — Prof.
PooLTON drew attention to two recent papers by Prof. H. S.
Jennings of Baltimore, D.8.A. : (1) " Observed Changes in
Hereditary Characters in Relation to Evolution " {Jouni. Wash-
ington Acad. Sci., vol. vii. No. 10. May 19, 1917, p. 281), and
(2) " Modifying Factors and Multiple Allelomorphs in Relation
to the Results of Selection " (American Naturalist, vol. li,
May, 1917, p. 301). In (1) the author, referring to the dis-
coveries of the French botanist Jordan and to the work of
Johannsen on " pure lines," expressed the conclusions to
which many naturalists have been led, viz. " that most of
the heritable difierences observed between closely related
organisms — between the members of a given species, for
■ T»iu. Linn. 8do., 1863, vol. zzvi, p. 13S.
t Journ. Quebett Micros. Club, 1S04-T, voL vl, ier. II, p. US.
i Joum. Qnekett Hicroa. Gab, ISM-T, voL vi, tat. U, p. 276.
ih, Google
( Ixxzri )
example — ai« not variaUoits in the senae of alterations; are
not active changes in constitution, but aie pennanent diver-
aities ; they are static, not dynamic. ... All thorough work
has led directly to this resolt : that any species or kind of
organism ia made up of a very great number of diverse stocks,
differing from each other in miiiut« partjculais, but the
diversities inherited from generation to generation." These
facts were particularly evident in the Protozoa, reproducing
by a single parent, as shown in Prof. Jennings's researches on
Paramoecium and Diffiugia, both of which were " found to
consist of a large number of such heritably diverse stocks, each
stock showing within itself many variations that are not
heritable." Selection might indeed operate, but it was the
selection of diversities that were already present, and thus
led to no new steps in evolution. [Just as it was shown on
p. Izxxi that a Bomean mimetic female form of PapUio polytet
was present as a very rare variety in Travancore, so it might
be held that all mimetic 'patt«ma of the species were also
present and could be made predominant by selection without
requiring any new stop in evolution.] " Variations . . . were
not variations at all, in the sense of steps in evolution ; they
were mere instances of the static condition of diversity that
everywhere prevails."
Well might the author conclude " in these days of plots
and spies, the evolutionieta might almost feel that the enemy
had crept into their citadel and was blowing it up from
within." But of course naturalists do not maintain that
changes never take place : " they admit that muUUioiu
occur"; that the permanent germinal constitution or geno-
type " may at rare intervals transform, as a given chemical
compound may transform into another and diverse com-
pound." A favourite theory of evolution nuiy be outlined
thus : " Organisms forming a multitude of diverse strains
with diverse genotypes; the genotype a moaaic of parts that
are recombined in Mendelisn inheritance; selection a men
process of isolating and recombining what already exists;
large changes occurring at rare intervals, through the drop-
ping out of bits of the mosaic, or through their complete
chemical transformation; evolution by saltations."
u„,,i,A J.., Google
( IxxxTii )
The author pointa out the difSculty involved in the very
eziatence of the minute Bti&ius, which must surely have ariaen
at some time or other, and aiiaen " not by saltations, for the
difierencea between the etiaina go down to the very limita
of detectibility." Then there ia the difliculty of "complex
adaptive etnictures, such as the eye."
Therefore many inveatigatoTs could not feel satisfied with
the favouiite theory and have been " looking for something
besides saltations as a basis for evolution; looking for here-
ditary changes that would permit a continuity in transforma-
tion." Among these investigators W. K. Castle has been
searching in the phenomena of biparental inheritance, H. F.
Osbom in the records of palaeontology, and the author in
organisms subject to uniparental reproduction. In this latter
" we meet the problem of inheritance and variation in its
umplest form ; for there is nothing which complicates genetic
problems so enormously as does the continual mixing of
diverse stocks in biparental inheritance. In uniparental re-
production we have but one genotjrpe to deal with; we can
be certfun that no hereditary characters are introduced from
outside that genotype."
Accepting as a foundation the facte already mentioned " as
to the make-up of the species out of a great number of diverse
stocks; as to the usual eSecte of selection being nothing
save the isolation of such pre-existing stocks," the author
undertook " a most extensive and intendve study of heredity,
of variation, and of the eSecte of selection for long periods "
within a single stock of a favourable organism. This he
found in the Rtuzopod Diffiugia corona, which has " numerous
distinctive charactere, all congenitel; all inherited in a high
degree; yet varying from patent to ofispring also; none of
these characters changed by growth or environmental action
during the life of the individual."
" Long-continued work showed that a single strain of this
animal, all derived by fission from a single parent, does
difierentiato gradually, with the passage of generations, into
many hereditarily diverse strains." While these variations
" arose in some few cases by rather large steps, or ' saltations,' "
" the immense majority were minute gradations. Variation is
iM:,i,A J 1^, Google
( Izzzviii )
u continuous as can be detected." Variation occurred la
many diverse cbaiacteis and gave rise to many diverse com-
binations of chaiact«T8. " Any set of chaiacters might v&tj
independently of the rest. The hereditary variations which
arose were of just such a nature as to produce from a single
strain the hereditarily different strains that are found in nature. "
After giving an account of these results obtained from an
organism with uniparental reproduction, the author briefly
refers to Osbom'a palaeontological evidence " for evolution
by minute continuous variations which follow a single definite
trehd," and to Castle's evidence that " in rats he can, by
selection, gradually tucrease or decrease the amount of color
in the coat, passing by continuous stages from one extreme
to the other," the change being " an actual change in the
hereditary characteristics of the stock; and not a mere result
of the recombination of Mendelian factors." From this
point the author proceeds to a critical examination of results
obtained by T. H. Morgan and his associates upon a species
which comes directly within the survey of the Entomological
Society, viz. the fruit-fly Drosophila. The species has normally
a red eye, but in the few years during which these researches
have been conducted it has come to present " seven grada-
tions of color between white and red, each gradation heritable
in the normal Mendelian manner " — " red, blood, cherry,
eosin, buS, tinged, whit«." " Three of these grades have
been discovered in the last five months. It would not require
a bold prophet to predict that as the yeaia pass we shall
come to know more of these gradations, till all detectible
differences of shade have been distinguished, and each shown
to be inherited as a Mendelian unit. Conwdering that the
work on Droaophiia has been going on only about seven or
eight years, this is remarkable progiess toward a demon-
stration that a single unit factor can present as many grades
as can be distinguished. . . ." But this is not all. As regards
the middle member of the series of eye colonrs, eosin, Bridges
has found seven modifying factore, each of which alters ita
intensity and gives rise to a secondary grade of colour. Now
each of these modifying factors are described " specifically as
mutations; as actual changes in the hereditary material."
l„y,|,AJI^,COOl^lC
( Izxziz )
The reeultfl are summarised as follows : " In DrotophUa there
occur mmuto chaagee in the germinal material, such as to
give, so far as our present imperfect knowledge goes, seven
diverse grades of a color which is itself only one grade of
another series of seven known grades. By means of these
graded changes one could obtain, by the mutationist's own
statement, the continuously graded lesulte which selection
actually gives. What more can the selectionist ask ? "
As the author remarks in paper (2), " The work in Men-
delism, and particularly the work on DrosophUa, is supplying
a complete foundation for evolution through the accumulation
by selection of minute gradations. We have got far away
from the old notion that hereditary changes consist only in
the dropping out of complete units, or that they are bound to
occur in lai^ steps. . . . The objections raised by the muta-
tionists to gradual change through selection are breaking down
as a result of the thoroughness of the mutationists* own studies."
The importance of Mendelism as an aid to selection is
insisted on in the same paper. " Hereditary variations, such
as give rise to the multiple allelomorphs and multiple modify-
ing factors, occur in some organisms rather infrequently, as
measured by the time scale of human happenings. If there
were no interchange of factors among individuals and stocks,
it would take a long time to obtain in one individual all the
tax diluteis of the eosin color of the DrosopkUa eye; one
arises in one individual, another in another. But by selective
cross-breeding it is possible to bring together into one stock
all the modifiers that have been produced in diverse stocks.
Mendelism acts as a tremendous accelerator to the efiectiveness
of selection."
Betuming to paper (1), the author finally concludes that
" Evolution according to the typical Darwinian scheme,
through the occurrence of many small variations and their
guidance by natural selection, is perfectly consistent with
what experimental and palaeontological studies show us ; to
me it appears more consistent with the data than does any
other theory."
A VEBY ANCIENT BEETLE. — The Fkesident exhibited a
specimen of a Carabid Beetle, Calovoma Tvgoaum, de Geer,
iM:,i,A J 1^, Google
( » )
found in the stomach of the mumm}' of an Ibis, the date
of which was reckoned to be about 1600 B.C. Foi compariaon
a modern specimen of the same species was shown, no
difierence between them being visible.
Pupation of Geotrupes typhobus. — Mr, Hugh Hain
exhibited a series of lantern slides illuattating the metamor-
phoses of Geotrupea typhoeta {Plates G, H, J).
He said that as he had wished to observe the life-history
of Q. typhoeus he collected a number of the beetles in the
autumn of 1915. They were more easily dug up in their
usual localities before hibernation, as in the spring they were
generally at a much greater distance below the surface of
the ground. After remaining quiescent through December,
they showed signs of activity in Januaiy (1916), and were
separated into pairs, which were placed in Subterraria filled
with fresh sand, together with some food conusting of horse-
or cow-dung at the top. Although rabbits' pellets or those
of the sheep or deer are perhaps the usual food, be had fre-
quently found Uie beetles under the droppings of horses and
cattle. The " Subterrarium " is an observation cage which
he had devised for the purpose of following the life-bistory
of insects whose activities were carried on under the surface
of the earth. It consists essentially of two vertical sheets of
glass fixed half an inch, or more or less as required, apart,
the intervening space being filled with earth or sand, into which
the insects burrow.
Before the end of Januaiy 1916 a number of shafts were
made by the beetles in several of the cages. A quantity of
food was carried down and packed into each shaft, an egg
having previously been deposited in the sand about half an
inch from the first layer of the food. Each food-maaa was
about five inches long, and was followed by a plug of sand
about two inches long. After the first was completed, a
second shaft was sunk from just above the first plug of sand,
and then a third and fourth.
Three such shafts ate shown in Plate G, fig. 2, that on the
right being the first made, then the middle one, and finally the
left one. The beetles were then removed, bo that their further
excavations should not disturb the work already completed.
l„y,|,AJI^,COOl^lC
PLATE G.
1. D!/ti»ms mtrgindlis. Empty larval skin from pupation
chamber; natural aize.
2. Subterrarium with three full-fed larvae of Qtolnijiet typhoeui
in their pupation chambers ; reduced.
3. O. tgphorut, larva, full-fed ; nrarjj' natural nize.
4. 0. li/phoeus, pupa, S, recently disclOBed ; nearly natural size.
PLATE H.
0<atrupea typhoeus.
1. Lorvs, full-fed, in pupation chamber, a
2. Pupa, $, rec^tly diaclosed.
Both about natural size.
1. Pupa, $, mature.
Both about natural si
PLATE J.
Qeotrupta tyyhoeue.
n,g,t,7.dh,G00glc
i
n,g,t,7.dh,G00glc
Prac. Em. Sot. LoitJ., 1917.
I
Hugh Main, FhtU. Ha'l- Tetii Enf.fCo^l
n,g,t,7.dh,G00glc
»f. Eiit. Sot. l.ond., 1917.
Ear. Cc. Uii.
n,g,t,7.dh,G00glc
Prac. Enl. Soc. Land.. 1917.
Hugh Main, Phtlt. Half- Tarn LHg. Co. Ud.
n,g,t,7.dh,G00glc
( »i )
The joimg larvae were first Been about the end of February,
and they ate their way to the top of their food-mass and
then back again to their starting-point, which was only
reached in August 1917, when they were fall-fed. EUtch then
formed an oval pupation chamber in the sand beyond the old
food-mass, and then lay on its back to await pupation. The
larva does not possess the stridulatory apparatus which occurs
on the second and third lege of the other members of the
genus Oeotrupeg. Probably the continental coleopterists are
well advised in placing tyffuxvs in a separate genua (Miwh
taunis) OS account of the morphological peculiarities of the larva.
The larvae pupated towards the end of August, and the
imagines appeared in September. The pupa is quite white at
firet, but the ejres soon darken and then the legs and thorax.
The elytra of the newly disclosed beetle are quit« white, and
pass through various shades of brown to jet black.
I^ck of time and opportunity in the present circumstances
prevented detailed observations being made as desired, but,
having seen the various successive stages as they appeared
and being prepared for them, it should be easy in more
peaceful times to repeat the experiment and to note carefully
the exceedingly interesting operations carried out in full view
in the Subterrarium. M. H. Fabie has written a graphic
account of the life-histories of a number of the Dung Beetles,
but his volaries did not allow him to see actually what was
going on underground, and some of his conclusions are not
confirmed by observations made in the Subterraria.
In answer to a question by the Pbesident, Hr. Main said
that possibly the purpose of the stridulation was to warn
away earthworms, which have a dislike to vibration. *
Larva of Cbakbocampa elfbhor. — Mr. Main also showed
two sbdes of the larva of C, dpmor, one at rest, and one in
its so-called " threatening " attitude after being disturbed.
In reply to a question by Piof . Pottlton he said that the larva
did not adopt that attitude while on the food-plant.
MltOCBY m CERTAIN BuTTSRrLIES OF NeW GuINBA. —
Dr. F. A. DixEY exhibit«d some drawings of butterflies
illustrative of his paper on this subject, of which he gave
the following abstract: —
n,g,t,7.dhvG00glc J
( «ii )
The iesemblanc« between the Pierme Huphina abnormu
and a form of the NTtuphaline genua Mynes waa remarked by
Wallace more than fifty years ago; and in later times an
even closeT likeness has been found to exist between the
Huphina and one or more species of the Fierine genus Ddiat.
The superficial lesemblanca is so great as to have misled good
lepidopterista as to the actual a£Bnitiea of these fonns; and
great confusion has prevailed with regard to their relative
position in a systematic series. The mistakes thus started
have not been confined to the species originally concerned, but
have extended to other forms of a more or less similar aspect.
On the evidence of neuration, scent-scales and genitalia,
there is no doubt that Wallace's aimormis is neither a TiuAyris
nor a Delias. The same is true of Honrath'a euryxantka,
which may possibly be conspecific with abnormis. The afSnit/
of these two Pierine forms, as was first recognised by von
Mitis, is with Moore's genus Huphina, of which they may be
considered to form a subordinate section, characterised by
slight peculiarities of structure. The same combination of
black and scarlet as in H. abnormit is presented by the
underside of the male of Ddiaa irma, Fruhst. ; though here,
there being no white fore-wing area, the resemblance to D.
omylion and B. abnormis applies only to the attitude of the
latter forms during complete rest. There is do scarlet sub-
marginal series in D. irma, nor in D. omytion from Western
New Quinea; but as the range of D. irma does not appear
to extend to the latter region, no particular significance can
be atteched to this fact.
While H. abnormis thus approaches the aspect of one group
of Delias, the closely allied, or, as some tHnk, conspecific
H. euryxantha bears considerable resemblance to another;
viz. the group formed by Delias mysis, Fabr., and its near
allies, particularly the New Guinea subspecies D. lara, Boisd.
With regard to Mynes doryca, Butl., it is observable that the
scarlet mark on the fore-wing corresponds roughly with that
on the hind-wing of H. abnormis, and vice vertd.
The resemblance between these forms extends in many
cases to the upper as well as to the under surface; thongb
here it is naturally less striking, being merely a <
Ij.y.l.Ajl^yCOO'^lc
( xdii )
vendon of a usual lieriue pattern. It U, however, remarkable
that Mynet, a yyinphaliiie, should possess an upper surface
BO strongly Pierine io aspect. A further point of interest is
the fact that Nepheronia jobaea, Boisd. 3, the only member of
its genus to reach New Guinea, shows on its upperside a
UkeneBB to Mynet doryca, which may extend even to the tint
of the pale ground-colour; while its underside, though without
red n[iarkings, is somewhat suggestive of a Ddias.
How far the principle of mimicry enters into these com-
binations may of course be questioned. That it has some
influence can hardly be doubted. More information as to the
bionomics of the forms referred to would be very welcome;
especially with regard to their distribution and habite.
DtpiUtna. X 75.
A NOncUBLX DiPFBBBHCB m THB AnTBNHAB Or CL09ELT
SSLATSD Species. — Mr. Green showed (on the Epidiascope)
an enlarged drawing of single joints of the male antennae of
Sybemia defdiiaria and H. aUTanliaria, and pointed out the
u„,,i,A J 1^, Google
{ xoiv )
Btructiml difFeiences between the antennae of these cloMly
allied species. In defoliaria the pinnae are very short, but
there are two paiia on each joint; whereas aurantiaria (with
the remaining British members of the genus) hae a single
pair of elongated pinnae on each joint. A pieparation of the
two antennae was exhibited under the microscope.
The Rev. F. D. Horice and other Fellowa gave parallel
instances of difierence of antermae in the case of closely ielat«d
iDsects.
Papers.
The following papers wero read : —
" Descriptions of new species of Staphylinidae from Singa-
pore," by Malcolm Caubron, M.D., R.N., F.E.S.
" Goleoptera of the Families Oatomidae, Monotomidae,
Colydiidae and Notiophyaidae from the Seychelles and AMabra
Islands," by the late Antoihe Grodtelle; communicated
by HuQH Scott, M.A., F.L.8., F.E.S.
Discuision.
In accordance with the Resolution passed at the meeting
on Nov. Ist, a discussion took place on Sir George Hampeon'a
method of taking the first species mentioned by the author of
a genus aa the type, without reference to the action of subse-
quent authors in fixing types.
In opening the discussion the President said that he had,
in accordance with the Resolution, invited Sir George Hamp-
son to be present and to state his views, but had received a
letter in reply, which he read, saying that he was unable to
be present, but that a statement of his views was given in
the " Novitetes Zoologicae," and in a paper which he hoped
would be printed in time to send a copy to the meeting. The
President read the first statement referred to, but said that
he had not received the other paper.
Prof. PooLTOM read a further statement from Sir George
Uampson, but took no part in the discussion.
Sir George Hahpson sent the following remarks with
regard to the use of the name Tinea : —
" The position ia this :
n,g,t,7.dh,G00glc
( «w )
" The Clothes-moths were called Tinea by all authors pre-
vious to Lmnffius, but 1758 ttob arbitrarily fixed as the
date of the commencement of Zoological nomenclature, and
Linneua unfortunately described Tinea from the wrong
species ; therefore either the foundation of Zoological nomen-
clature must be abandoned oi the use of Tinea altered.
" I stated the dilemma to Mr. B. Meyrick, whose only
reply was that ' types were made for science, and not science
for types,' i. e. he acknowledged the facts, but thought they
ehoold be ignored."
He also asked, " Why are Scopoli's genera for Butterflies
in 1777 ignored 1 " and gave a list of these genera and of the
names they would displace.
Dr. O. A. E. Marshall pointed out that, is contrast with
the haphazard method of the earlier zoologists of applying to
animals such scientific names as they thought fit, without
regard to the names used by other workers, our modem
system of nomenclature has been built up on the principle
that all valid systematic work done by earlier anthore must
be recognised. With this principle, the so-called " law of
priority," the method adopted by Sir Qeorge Hampson, in
oppositaon to all other entomologists, is in direct conflict ; for
in defining the older genera he has assumed the first species
mentioned by the original author to be the tjrpe, and has com-
pletely ignored the subsequent revisions of such genera by
other systematiste. No adequate reason has been offered for
this arbitrary innovation, except that it will relieve those who
adopt it from the trouble of acquainting themselves with the
work of previous writers. Apart from the hopeless confusion
in zoological nomenclature that would result from the general
adoption of such a system, it is so obviously unjust that it is
not likely to find acceptance among scientific men.
The Rev. F. D. Horice remarked that the Honey-bee had
been accepted by all authors as the typical Apis of Linn^, but
that the first species standing under Apis in the " Systema "
(Ed. X) was not the Honey-bee, but what we now know as
Eucera hngicomie, L. Also, that the first species described
in the same work as a Sphez was an insect from Surinam,
which the author certainly would not have selected as his
iM:,i,A J 1^, Google
( XOTl )
" fcyp*'" 8U"*'* '° Ed. XXI he removed it from Sphex altogether,
and placed it in ApU I Again, the first species described by
Linn^ under Formica wae not what all authors now call a
Formica, but a Camjmnoltu. He felt sure that, whatever
might be the case as to other groupe, no workers on the
Hymenoptera had yet adopted or were ever bkely to adopt
a principle which involved such consequences.
The Pbesident, taking part in the discussion, said he did
not agree with some of the remarks that had been made in
reference to Sir Geoi^ Hampson's motive in adopting the
principle of selecting the first species as the tjrpe of the genua,
for he thought Sir George was firmly persuaded that his was
the only method capable of leading to stabihty in nomencla-
ture, and that the rest of the zoological world would finally
be converted to his way of thinking. He had much sympathy
with the contention that his method would save an immense
amount of time spent in bibliographical leseareh. Had that
method been acted upon from the first, it would, no doubt,
have simplified procedure and placed nomenclature on a
firmer ba«a. But Sir George had come into the world a
century and a half too late. His mission should have com-
menced with Linnieus himself, vrith his pnpil Fabricius, and the
other entomologists of the time, not one of whom entertained
the idea of fixing the firet species as the type of the genus.
As a member of the International Committee on Entomological
Nomenclature, he had recently been asked for an opinion
on the disputed question of the geno-type of Ctmex, Linn.,
and in endeavouring to arrive at one that would be in accord
with the rules of the International Code, he met with difficulties
of various kinds. But this, he thought, was a very excep-
tional case, and the chief difficulties he found in dealing with
it, were the result of the arbitrary action of one or two of the
older authors, who refused to accept the fait accompU, and
proceeded to undo the work already done. Theirs was the
kind of action which Sir George Hampson would hke to see
followed at the present day. It had only led to the contuuon
and the waste of time, which he deplores, and even if we had
no moral obligation to the workers of the past, the great
inconvenienee for zoologists in every branch which would
l„y,|,AJI^,COOl^lC
{ xcvii )
reautt from adopting Iiie principle and appljoog it to the geoeK
of the older authors, ignoring all that baa been done in tJw
meantime and treating the writings of our predeceasore as so
many scrape of paper, condemns it at once aa piepoeteroua
and absurd. He had been looking up aome others of the
genera, besides Tinea, in the 10th Edition of the " Systema
Naturae," to see what changes would follow if Sir Geoi^
Uampeon'e method were to be adopted. Ceramhyx, which
gives its name to a family of Longicoms, and is now applied
to a genus of well-known European species, would be given
inatead to the Harlequin -beetle, which ia a native of tropical
America and belongs to another family. Tbe name I^epttira,
with L. agvatica as the type, would be transferred from the
flower-frequenting group of Longicorna known aa the Lep-
turidae, and given instead to a group of sub-aquatic Fhyto-
phaga. The glow-worm would loae the name of Lampi/ru
nodiluca, to be kiiown henceforth aa Canlharis nodiluca, with
a corresponding change in its family name. It would be
wrong in future to refer to the common house-fly as Mtisca
domesHca, or to place it in the family Muscidae; the latter
muae should be reserved for the Hover-flies, which we now
caU Syrphidae. One had only to think for a moment whit
hundreds of changes of this kind would involve, not merely
Id systematic writings, but in literature of. an economic
or more general character, in order to see how hopeless is the
prospect of getting a majority to adopt the method which
would bring them about. The number of genera whoae
types cannot easily be determined in accordance with the rules
laid down in the International Code of Zoological Nomen-
clature was surely not so great that we must be prepared to
face the revolutionary alternative which Sir George Hampson
so persistently places before us, and to which he adheres in
bifi own writings in spite of the protests of almost every one
of his fellow- workers in entomology. It was to be hoped
that he would be led by the views expressed that evening to
reconsider his position, and that he would fall into line on the
aubject of nomenclature with the syateraatists of his own and
of every other country.
Mr. DufiftANT remarked that the present discussion really
PKOC. BNT. aOC. LOND., V. 1917 O
iM:,i,A J h, Google
( xcviii )
leenlted from the action of the Zoological Society in lefunng
u unjustifiaUe snd vitra vires sd attempt to change the
familiar and wotld-wide application of the generic name Tinea
■ of Linni — to remove this name from the clothes-moths and
traoBfer it to the honey-moths {GaUeriadae), with which tt
had never been specially associated. This would involve not
only the change of generic names but also of the higher groups,
Tineina, Tineidae and GaUeriadae, the only grounds for Ute
change being the numerical sequence of species in the setiee.
The system was founded on a total disregard of all systematic
or critical work done after the original pubbcation of a generic
name, and the result would be to cancel everything that had
been done, and to say that all work was wrong unless the first
species had been select«d as the type. The adoption of Sir
George Hampson's sytem would involve the change of every
familiar generic name, unless by chance the fiist species had
been constituted the type by earlier authors. Nothiog could
be gained by such alterations .except imiformity of treatment
in one series of volumes to which no one else but the author
was committed.
Vears ago Sir George Hampson started a correspjndence
on this subject with the following letter : —
•'SrUiih M uaeum {Xalaral Hialorg).
•'Stpt. \2lh. 18M.
"Dear Sirs,
" As there is a probability of a new British Museum
descriptive Catalogue of all the described species of Heterooera
being shortly commenced it ia thought advisable to consult a
certain number of the best-known authors on the Lepid>pUra
as to the system of nomenclature which is to be adopted.
With this object in view the euclosed list of questions is sent
to you, and if you will be good enough to express your views
on the subject they will be of great assistance in forming the
scheme of the work, and perhaps contribute to attaining a
more uniform system of nomenclature in the Lepidoptera.
" The names selected for consultation are : — Prof. Scudder,
Prof. Femald, Prof. J. B. Smith. Dr. Staudinger, Herr P. C. T.
Snellen, Prof. Aurivillius, Prof. A. R. Grote, Lord Walsingham,
E. Meyrick, Esq,, W. F. Kirby, Esq.. and Sir G. F. Hampson.
l„y,l,Ajh,COOl^lc
( lax )
Other nsmes might easily hava bean added to this list,
^ was thought that eleven would conatttnte a man
■able committee than a larger number would, and those
..ed sufficiently represented the various opinions on the
.DJect and the countries niost interested in the Lepidoptera.
" It is proposed to circulate the papers once for you to
give your opinions, and then again for you to see the answers
given and make any additional remarks that strike you.
" lam, dear Sirs,
" Yours faithfully,
" G, P. Hampson."
Question 7 was : " By what process is the type of a
hetero-typical genus to be ascertained ? " and a special
question under this heading was : " c. What is the type of
Tinea, L. ? "
On Sir Qeorge Hampeon's suggestion that " the first species,
or the first species agreeing with the description, to be con-
sidered the type," the opinions were : —
For: 1, Hampson; 2, Staudinger.
Against: 1, Aurivilhus; 2, Femald; 3, Grote; 4, Kirby:
5, Meyrick; 6, Scudder; T.Smith; 8, Walsingham.
[Snellen stood alone in totaUy rejecting "le syst^me des
types gen4riques.'']
And on the question : " c. What is the type of Tinea, L. ? "
there were in favour of jidlumeUa, L. (an ordinary clothes-
moth) : 1, Walsingham; 2, Meyrick; 3, Kirby; 4, Femald;
5, Smith; 6, Aurivilhus; 7, Grote (8, Scudder and 9, Stau-
dinger assumed to concur).
While Sir George Hampson was unable to say which was
the first species that agreed with the description, after having
wrongly cited as the type geUUeUa, L. (a species which did
not occur in the lOtb edition of linne).
Sir George Hampson has rightly accepted as the type of
Sphinx the eevetUh species Ugustri, which Linne states in the
Fauna Gnecica was " Vulgo Sphinx," and also aa the type
of Bombyx species 18 mori " Vulgo Bombyx " according Ut
linne.
It is therefore evident that the first species included in a
iM-,i,A J h, Google
( » )
genus by linne was not neoessarily tlie type — indeed, in I%il.
Bot. (ed. 1), 197 (1751) and {ed. 2) 201-2 (1763), linn* him-
self wrote : " 8i genns receptum, secundum jus natoiae et
artis, in plura dirimi debet, turn nomen antea commune
manebit vulgatiBsiiuBe et officinal! plantae," showing that his
opinion was that the name should be restricted to the best-
known species — not the fiist on the list.
The whole question was thoroughly thTesbed out in 1896-
1898, and a full report with analyses of raphes to the questions
circulated will be found in Proc. IV, Int. Congress Zool.,
273-312, Cambridge, 1899.
Mr. Durrant concluded by remarking that nothing new was
being brought before us, and he had dealt with the whole
question twenty years ago !
Dr. G. A. K. MARaKALL then proposed the following
Resolution : —
" That in the opinion of this Society the arbitrary method
followed by Sir George Hampson of taking as the Type of a
Genus the first species in the series, disregarding the work
of previous authors, ia contrary to the usage of Entomologists,
and this Society declines to adopt this system."
Mr. G. T, Bbthune-Baker spoke as follows :—
In seconding the motion I would draw your attention to
the fact that the proposition of Sir George Hampson to adopt
the first species as the type has been discarded practically by
all authors of zoology everywhere. Sir George claims Roths-
child and Jordan as adherenta to his method, and in their
great work on the Sphingidae it is true they did adopt that
method, but they have discontinued it and do not adopt it
now. Again, Barnes and McDniinougb, who at first thought
it was a cotivenieiU arrangement, have likewise discarded it,
finding, as they say, the whole of the zoological world against
them. Even Sir George himself makes his own exceptions
(Novit. Zool., xxiv, p. 19; 1917), to wit, Tortrix with viridaua
as the type, the second species; Sphinx with liffustn as the
type, because of the sphinx-like attitude of the larva;
Bombyx with jnori as type. He gives his reasons, it is true,
but similar reasoning ought to apply with equal force to
the genus Tinea, for I demur entirely to his statement
l„y,l,Ajh,COOl^lC
{ ci )
that pdlioneUa when at rest folds it« wings in a t«nt-hke
Sir George's suggestion that most of the authors in " Seitz"
and most of the American authors adopt this principle simply
made me gasp with surprise; whether at the time of the
initiation of this, at the Cambridge Congress, the American
authors adopted the principle I do not remember exactly ; I
doubt it; but to-day there is no question that they do not
Ho so, and looking over my " Seitz ' again I really do not see
any justification for the remark. One ot two do so, but the
bulk of the authors act quite independently of the method,
taking the first species in some cases but not in others.
Further than this, the injustice of the thing is apparent on
the face of it. Why should we at this late date take upon
ourselves to ignore all the revisionary work of earlier students ?
Sir George would scrap the work of all revisers if they did
not adopt his pet theory ; we might quite as well adopt as a
working principle the suggestion that all descriptions should
be ignored if a figure was not published with the description
or shortly after.
Instead of coming rapidly into general use, as Sir George
thinks, it has gone rapidly out of use. It never had general
acquiescence, and to-day in all branches of zoology it is dis-
carded by the great majority of workers, and I can only hope
that Sir George will see that he stands practically alone in
his idea, and that he will soon leave that aoUtaiy position.
Mr. T. Iredale, who was present as a visitor, spoke regarding
Sir G. Hampson's contention that this principle (the first-
species rule) was accepted by a majority of workers in other
classes, and as regards ornithology and conchology denied
that it had any adherents at all. Some years ago the American
Ornithologists' Union debated the subject and inclined to
accept the first-species rule, and advised to that effect. The
result, however, was that immediately the advice was re-
pealed, as it was proved that confusion would ensue, without .
prospect of later stability. It might be further added that
even when the matter was under discussion it was agreed by
all workers that the Linnean genera must be excepted, as it
was a known fact that the first species In Liim^'s syatem was
iM-,i,A J h, Google
{ cii )
generally atypical and that the central one was commonly
the typical species. No exact law as we undeistand laws
was followed by early writere and many ranged their species
alphabetically, and moreover in such cases examples cited
were not displayed for generic chaiacters, but on account
of their beauty, size, or similar object. As regards concbology,
the first-species rule has never been absolutely accepted,
though seventy years ago it was a custom when other methodt
failed of accepting the first species arbilrarily as type. At the
present time certainly no systematic worker in either group
transgresses the Laws of the International Congresses as
regards type selection, nor is there any reason to suspect
any difieience of opinion as to the results to be achieved by
the continuance of such acceptance.
The Sesoiution was then put to the meeting and carried
unanimously.
n,g,t,7.dh,G00glc
( c'" )
THE ANNUAL MEETING.
The Annual Meeting took place on Wednesday, January 16th,
1918, Dr. C. J. Gahan, M.A., D.Sc., Fieeident, being in the
Chair.
Mr. W. G. Sheldon, one of the Auditois, lead the Treaauier's
etatement of Accounts, which was adopted on the motion of
Mr. Stanley Edwabds, seconded by Mr. Frisby.
The Rev. Q. Wheeler, one of the Secretaries, then lead
the following
Report of the Council.
The Society has during the past year felt in many direc-
tions the pressure of the war to a much greater extent than
had previously been the case. Although the attendance at
the meetings in the early part of the year was as large as
could be reasonably expected, yet, owing possibly to air-raids,
there were less than twenty present at either of the October
meetings; there has, however, latterly been a marked im-
provement in attendance, and at the December meeting the
numbers only fell just short of fifty. The standard of interest
of the meetings has, however, been well kept up, although, as
usually occurs, there have been one or two meetings when a
larger number of exhibits would have been welcome.
With regard to numbers we are again somewhat reduced.
While twenty new Fellows have been elected we have lost
eight ordinary Fellows by death, nine by resignation and nine
by removal of names for non-payment of subscription, the
latter including the eight enemy aliens whose names were
still upon the list. We have also lost one Honorary Fellow
by death, viz. Dr. Emil Frey-Gbssneb of Geneva, whose
place has not yet been filled up. The Council has also again
to regret the toss of one of its members in the death of Mr.
A. E. OtBBS, shortly after his re-election. It would be im-
poeeible to pass over the death of Mr. Gibbs without a tribute
to his usefulness both on the Council and more especially on
iM-,i,A J h, Google
( "» )
the Business Committee of which he had been (or five years
a most valued member.
The Society now consists of eleven Honorary and two special
Life Fellows, together with five hundred and eighty-six ordinary
Fellows making a total of five hundred and ninety-nine.
In consequence of the great shortage of labour in the print-
ing trade, as in all others, considerable delay has been expe-
rienced in the production of the Transactions. This fact
combined with the enormous increase in the price of paper
has made it necessary this year to produce Parts II, III and IV
together, both with a view to economy and also to make it
possible for the future to publish Parts I-IV as neariy as
may be within the year for which they are dated. The
volume for 1917 will consist of 405 pages illustrated by one
chromo -lithograph, six 3-coIour plates, fifteen half-tones, one
line block and one sketch-map, twenty-four plates in all.
Sir George Kenrick bears the coat of the 3-colour plates.
The papers are thirteen in number by the following authors : —
Messrs. R. S. Baonall, F.G.S. (in conjunction with Dr.
J. W. H. Harbison, D.Sc.); G. T. Bethune-Bakek. F.L.S.,
F.E.S.; G. E. Bodkin. F.Z.S., F.B.S.; P. A. Buxton, B.A.,
F.E.8., M.R.C.S.; G. C. Champion, A.L,S., F.Z.S.; Dr. T. A.
Chapman. M.D„ F.Z,S„ F.E.S.; Dr. E. A. Cockayne, M.A.,
M.D., F.E.S., et«.; Dr. H. Eltringham, M.A.. D.Sc., F.E.S. ;
Dr. J. W. H. Harrison, D.Sc. (in conjunction with Mr. R. S.
Bagnall); 0. E. Janson, F.E.S.; Sir George H. Kenrick,
Bart., F.E.S.; E. Meyrick, B.A., F.R.S., F.E.S. ; Rowland
B. Turner, F.Z.S., F.E.S. ; and C. B. Williams, M.A., F.E.S.
Of these, eight refer to Lepidoptera, two each to Coleoptera
and Hymenoptera, and one to Diptera. The Proceedings will
consist of about one hundred pages illustrated by nine half-
tone plates and several text-figures ; the cost of two of the
plates is borne by the Rev. F. D. Morice.
At a Special Meeting held on Jan. 7th, the Council drew
up and forwarded to the Prime Minister the following prot«6t
against the proposal of the Government to commandeer the
Natural History Museum ; —
" This Society, founded for the advancement and practical
application of Entomological Science, knowing that this
l„y,l,Ajh,COOl^lC
( CT )
Science, especially at the present moment, plays a most im-
portant part in many queetions, often of extreme urgency,
affecting the health oE the Nation and it« furces at home and
abroad, its food BUpplies, its timber, and the raw material of
its manufactures, views with the gravest concern any action
that would impede work essential to the National welfare.
" Such work includes : —
*' (1) The investigation into the relation between insects
and the spread of such deadly diseases as typhus, plague,
malaria, yellow fever, sleeping sickness, etc., to which our
troops are exposed in various parts of the world.
'* (2) The prevention of the attacks of innumerable insect
pests upon every kind of food crop, whether growing, in
transit, or in storage, by which the available supplies are very
materially reduced, and occasionally even totally destroyed.
" (3) The protection of timber, cotton, and other raw
materials essential to the conduct of the war, from the many
pest« that attack them.
" Towards the solution of these problems the collections at
the Museum have in the past largely contributed, and many
of them are at present nndei investigation.
" The proposed action of His Majesty's Government in
reference to the Natural History Museum, would have a dis-
astrous effect upon work which demands continual reference
to its enormous coUections. It is obvious that to be of any
practical value these must always be readily available, and
moreover their removal would not only be a very lengthy under-
taking, but could not be carried out wit hout irreparable damage.
" The Entomological Society of London feels bound there-
fore to enter the strongest possible protest against such pro-
posed action, the full consequences of which can hardly have
been realized, and in the interests of the Empire urges that
the suggested interference with these important collections
should be abandoned.
" Signed on behalf of the Council,
" G. B. LoKOSTAFF, Vice-President."
The Council desires to express its great regret at the retire-
ment of the Treasurer in consequence of ill-health, and also
iM-,i,A J h, Google
( cvi )
its appreciation of his servicee for the paat fourteen years,
which have been invalu&ble.
The Treasuber reports as follows : —
" The accounts compare favourably with those of last year.
Theie was then an estimated Habihty of £294 against a balance
of £237 14s. 9d., whereas this year the estimated liabilities
are about equal to the balance. There is a falling off of
£44 12s. in the amount receiveil fot Subscriptions compaied
with 1916; but the arrears received (£45 3s,) are in excess
by £18 18s. There are no otliet items which call for comment
with the exception of the ' Subscriptions in arrear.' Hitherto
the return has been made foi those ' considered good.' Under
present conditions it is impossible, with any degree of cer-
tainty, to separat« the ' good ' from the ' bad ' ; I have
therefore returned the total amount of the indebtednesB^ viz.
£232 Is. A considerable portion of this sum is merely in
abeyance during the war, being subscriptions nnpaid mean-
while by Fellows serving at the Front.
" I much regret my inability to continue my Treasureiship,
and I thank the Society for the confidence reposed in me
during the time I have had the honour to act as their Treasurer,
and I take this opportunity of thanking my Colleagues and
the Fellows generally for their unqualified courtesy shown me
during my term of office.
" The Society, I consider, may be congratulated upon
obtaining the services of Mr. W. G. Sheldon as my successor.
" A. Hugh Jonbs."
The LtBRARUN reports as follows : —
" Three hundred and fourteen volumes have been issued
from the library for home reading. Eighteen volumes and a
large quantity of Separata have been presented to the Library.
As was the case last year, very few foreign periodicals are
coming to hand. The Library has been well used for purposes
of reference."
The Report was adopted on the motion of Mr. 0. B. Janson,
seconded by the Rev. F. D. Morice.
No other nonunations in addition to those of the Council
having been received, the President declared the following
l„y,|,AJI^,COOl^lC
( cvii )
Fellows to have been duly elected as Officers and Members of
Council for the ensuing year : —
President. Dr. J. C. Gahan, M.A„ D.Sc. Treasurer, W. G.
Sheldon. Secretaries, Comm. James J. Walker, M.A., B.N.,
F.L.8.; Rev. George Wheeler, M.A.. F.Z.8. Lif^arian,
George Charles Champion, F.Z.S., A.L.S. Other Members of
Council, A. W. Bacot; E. C. Bedwell; K. G. Blaib, B.Sc;
Dr. T. A. Chapman, M.D., F.Z.S.; W. C. Crawley, B.A.;
H. Willoughby Ellis, F.Z.S.; Dr. H. Eltringuam, M.A.,
D.Sc., F.Z.8.; J. C. F. Fryer, M.A.; A. Hugh Jones; Bev.
F. D. MoBiCB, M.A.; S. A. Neave, M,A„ B.Sc., F.Z.S.;
Herbert E. Page.
The President then delivered an Address, aft«r which
Mr. SiHES proposed a Vote of Thanks to him which was
seconded by Dr. Eltbingham; the President in returning
thanks spoke of the neglect of the Biological Sciences on the
part of the Government.
Mr, Collin then proposed a Vote of Thanks to the Officers
which was seconded by Mr. Donisthorpe, both of whom
made special allusion to the regret felt by the Society at the
retirement of Mr. Jones from the Treasurership, after so long
a period of very efficient work in the Society's interests.
The two Secretaries said a few words of thanks, the
Treasurer and the Librarian being absent.
n,g,t,7.dh,G00glc
( cviu J
ENTOMOLOGICAL SOCIETY OP LONDON.
^•Uhm SbMt for tbt T«ar U17.
R1H--BIPTS. I TaIIIBNTS
iting TraiiurtiaDi, c(c 2«
f"' «•« 5(
' mid Office Ei-
BalsDce in h.u,l, Isl Jan .
1917 237 n
Siibscriptioua for 1817 ... 390 12
Arreara « g
S.le.of TmnnctioM " lit » |S I *^""''"'' pf B?"""" for
Int™tonIuvesto«ut.Z '" " "* L'S^' T S'','".*' "'"'■
Con«)l. ... X35 8 0 "«Mni.Meldol.'.gr«nt
Bifininghama I Mpercoot™
5 7 10
Income tn recnvered on
■bove until 5th October,
3C 15 10 , ]»18
1917
d "ill In' fit!
15 7
Intereat on Deposit ... « 7 1
Grant from Mr<. Hd<lola
«percoDtr» 31 ,o „
SubKnplion- 111 AilTODce 15 15 0 (
£957 8 11 !
bKcripticiun
be Front.)'
r !.» Fr!lo«.
£1,351 2j. 2rf, Conaol,
Original coat £1,233 3j.
Present value «t the
pnCB of 64} on the 31st
December, 1917 ... 737 IB in
£238 12;.. 4,1. BimiiDybam
3 pec reula. Oriiiual
coiit£250. Predtnt VKlur
EUTIMATSD I.UBJUTIM.
'■■'St of printing Part« 1 to 5 fti-
mated «t Ibe coat of Part* 1 to 5
forlflie,wj,£3M. E»tiin.ledr«t
of Plates in huid. mj, £18. mtkioK
» total liabiJity of my £liS3
againat the balance of fiSp/n,^
■**^- " '" I f""'"! "Tert. yth January. I9IS^
£1,181
Additional Assets:—
Contents of Library
Valnsd at, aay, £3,8li0.
ThcTtlueofSecuritiBasI
A. HuoH Jones, Trfamrir.
Bth January, 1918.
total depreci
W, G. Sheldon.
H. Willodohbi Ei.lis
S. A. Nbatb.
k; G. elxia.
>f £603 ISj. 8./.
n,g,t,7.dh,G00glc
THE PKESIDENT'S ADDKESS.
Gemtlemen,
Yoir have juet heard the lieport of the Council, and I
think you will agree that the state of affairs which it diBcloses in
reference to the general progress of the Society and its present
financial position is one with which we have no leason to be
dissatisfied. Our meetings have been attended even better
than one could have expected considering the eventful nature
of the times through which we have been passing. The
subjects discussed and the exhibits shown have continued to
be as interesting as ever. We have had no lack of valuable
papers, although you may probably find the volume of Trans-
actions for the past year not quite so bulky as some of those
for preceding years ; but in their decision to Unfit its size the
Council have been influenced not so much by financial con-
siderations as by a desire to return to the old and well-approved
practice of issuing the first four Parts within the year whose
date they bear, That practice seems to be in every way
desirable ; it helps to make the financial situation more easily
understood ; and it has decided advantages for the systematist
by saving him the trouble of giving double dates in his refer-
ences. It was insisted on in the days when for a short period
I was one of the Secretaries, and I remember well the anxious
moments we used to have towards the end of the year for
fear that Part IV would not be ready for issue before the 31st
of December. Knowing some of the difficulties with which
the Secretaries have to contend, I should like to remind
Fellows of the Society that when they contribut« to the
Transactions, exhibit specimens, or speak at the meetings
they can, if they will only take the trouble, make the work
eauer for them.
n,g,t,7.dh,G00glc
< ex )
In addition to the mention of Fellows who have died during
the year, the R«port of the Council relates to oh vtiter matter
which can only be a source of the peatest regret. Needless
to say, I refer to the resignation front office of our worthy
Treasurer. He has torso long a time, and with such great can
and success, looked after the financial affairs of the Society,
tliat it owes him a debt it can never expect to repay.
But if gratitude ta hira for his services can help to wipe
out the debt, he may rest assured that he has it in full
The Fellows of the Society who have died in 1917, fortunately
not a very large number, include Dr. Emile Frey-Gessner, an
Honorary Fellow ; Charles Owen Waterhouse, a former
President; Arthur Emest Gibbs, who was a member of the
Comicil, and six other Fellows.
Dr. Frey-Qessner had a deservedly high reputation as an
Entomologist both here and abroad, and although I had not
the good fortune to know him personally, I mideistand that
he was very popular with his fellow-workers on the Con-
tinent. He was elected an Honorary Member of the Inter-
national Congress of Entomology at the meeting held in
Oxford in 1912, and in the'same year was elected an Honoraiy
Fellow of this Society. Bom in Canton Aargau, he began
collecting Swiss insects of all kinds at an early age, chiefly
for Drs. ImhofI and Schmidt of Basel. Thou^ his own
tastes were at first directed towards the Orthoptera, they were
gradually transferred to the Hymenoptera. Hia monograph
of the Swiss bees (Hymenoptera Helvetica, Apidae), which
was completed only a short time before his death, is considered
by competent judges to be a very thorough one. He had a
general knowledge of exotic insects, hut devoted himself
chiefly to the insects of the Swiss fauna ; and his collections,
which are entirely Swiss, are now mostly in the Bern Museum,
while some are at Geneva.
Charles Owen Waterhouse was the eldest son of George R.
Waterhouse, who was one of the original members of this
Society, and also at one time President; so that his name has
always been cl()aely connected with the Entomological Society
of London and carries us back to the days of ite foundation.
l„y,|,AJI^,COOl^lC
{ cxi )
It is associated also with the names of Charles Darwin and
Bichaid Owen, intimate friends of his father; so that his
brother-Bntoiuologists might feel almost disposed to envy
him the advantages to which he was bom, and which also
he most have derived in his early days from contact with such
distinguished naturalists. But if those advantages are to be
held accountable in any way for the great interest he took in
Entomology and the work he has done in its service, this
Society may claim bo have received some benefits from them.
He had been for forty-eight years a Fellow, and for two years
(1907-8) President of our Society, served at different times on
the Council, rarely missed attending our meetings, and the
mere titles of the papers which he contributed to our Trans-
actions help to fill more than one page in the Catalogue of
our Library. As he was personally famiUai to most of you,
and you have had many opportunities of knowing how wide
and intimate was his knowledge in almost eveiy branch of
Entomology, it will be easy for you to understand how highly
I appreciated him, and how grateful I feel, having been asso-
ciated with him bo long, for the assistance he was always so
ready to give, especially in the days when, almost a novice in
Entomology, I began to work on beetles in the British Museum.
At that time, although I had a good knowledge of biology, the
result of two years' steady work under Prof. Huxley in his
laboratory at South Kensington, I knew comparatively little
about insects. One of the things, however, which I had
studied and thought I knew well was the structure and
homology of the mouth-part« of the cockroach. Quite m-
cently, I have been reading a paper by Prof. E. Bugnion
upon this very subject, and it has reminded me of a difficulty
1 had at one time experienced in reconciling various statements
about the piece known as the 3t^-tnentum. I had been taught
to regard this piece as part of the lower Up and homologous
with the cardinea or basal segments of the first maxillae ; and
this is the view given of it in nearly all of the text-books ; but
the part which in Coleoptera is known by the same name is
merely a continuation of the gula and is so firmly fused with
the head capsule that it can only be regarded as a sternal
part of the head. I went with this difficulty to Waterhouee,
u„,,i,A J.., Google
{ cxii )
and I am inclined to think that hie leadiness to hdp on that
occaBion was the origin of an investigation in comparative
morphology, the result of which he published at his own
expense, in a short paper, with coloured plates, entitled
" The Labium and Sub-mentum in certain Mandibulate
Insects." In this paper he has endeavoured to show, by a
comparison of the parts in various forms, that the sub-me&tum
is, in fact, a part of the head itaeU and need not be taken into
account when-homologizing the parts of the labium with those
of the maxillae. Owing to the manner of its publication, this
paper has, I feat, been almost entirely overlooked, which makes
it all the more desirable to call special attention to it here.
Prof. Bugnion, without in any way referring to it, takes very
much the same view of the sub-mentum as Waterhouse, and
supports it by other statements of fact ; but as the terminology
used by him is not the same, one has to be careful in readmg
his paper to note exactly what he means by sub-mentum.
This name he applies to a part of the labium, using the name
basilaire instead for the part figured and described in most
of out text-books as the sub-mentum. Which particular
part has the right by priority to the name of sub-mttaum,
I cannot, at the moment, definitely say, but I rather think
the name was first made use of in Coleoptera and in the same
sense in which it was applied by Waterhouse.
Arthur Ernest Oibbs was another Pellow of the Society who
took a great interest in its work; and as a member of the
Council and of the Business Committee, as well as in many
other ways, rendered it most valuable service. He was on
active member also of the South London Entomological and
Natural History Society, of which for a time he was Vice-
President. He was a keen student of the Lepidopteia, and in
his frequent travels on the Continent, and with the aid of
collectors in other parte of the world, got together a very
considerable collection. He was very much interested also in
the local museum near his home at St. Albans, which has
benefited largely, I beUeve, through his generosity. Those of
us who have had the pleasure of meeting him frequently and
of knowing his kindly disposition will miss him greatly, as
indeed will this Society as a whole.
n,g,l,.rJM,GOOglC
( cxiii )
' the other Fellows who have died, I wish I wete able
^enonal knowledge to say at least a few fitting words.
irtunately my acquaintance with them was so sli^t
uld be unable to say more than I could gather from
jary Notices which are accessible to all in the
.ological Journals.
At one of our Annual Meetings, the President of the yeai
reminded us that he was not required under any bye-law to
deliver a Presidential Address, but he delivered it all the same,
and an excellent one it was. If you expect me to continue
now and to follow his good esample I fear you will be dis-
appointed. A good Presidential Address, such as yon have
been accustomed to hear at these annual meetings, needs a
good deal of preparation and a lot more besides, even if one
happen to make choice of an interesting subject. The subject,
or rather the mixture of subjects, on which I propose to speak,
is, however, not so much a matter of my own choice, as of
being compelled to it by the necessities of the case. For I
have to admit that so far as preparation ia concerned, yon
may have reason to think that I have been rather neglectful
of my duty to you, and the only excuse I can ofier, which I
trust you will accept, is that, although my intentions were
of the best, I have been quite unable to find the time that was
needful to fulfil them.
When Mr. Bethune -Baker, in his Presidential Address de-
livered three years ago, suggested a subject for investigation
which, he said, would be of the utmost importance to science,
1 had a strong inclination at the time to try and act upon it.
It appealed to me as a Coleoptehst. An investigation of the
male genital organs of beetles would, I thought, be of the
greatest interest, especially if one could go on to compare them
with those of other insects with a view to homologizing the
ports. But an investigation of that kind requires time,
certainly much more than has been at my disposal from that
day to this; and in consequence I have had to abandon the
idea. It is, however, a work that should employ many hands,
notwithstanding that Shaip and Muir in their invaluable
memoir which appeared in our Transactions for 1912 have
PROC. BNT. SOC. L0N1>,, V. 1917 H
u,y,i,Ajh,CoOt5lc
( cxiv )
covend a good part of the ground. That memoir is a veritabU
storehouse of facts on the subject, and enables us to see
almost at a glance the type of structure of the genital tube
and its accessories in almost every family of Coleoptera'; but
it still leaves us without sufficient evidence to judge whether
all the conclusions they seem inclined to draw from them
are right. The organs they describe are, without doubt, of
considerable phylogenetic importance, but whether they have
all the importance which they claim for them is another
question. To me, at any rat«, some of their conclusions seem
open to suspicion, especially those which lead them to hint
that the Coleoptera are not a monophyletic group of insects,
but, starting from some Neuropteroid or other pre-Coleopteran
forms, have reached the Coleopterous stage along difieient
lines of descent. We all admit the great importance which
the structure of the aedeagus has in enabling us to discriminate
between closely allied species in certain genera and in many
whole groups, and we know how difierent it is at times in two
species which in all other respects seem to be very cloEely
related. This applies also to the external accessory part^ —
the parts visible in Coleoptera without dissection — a subject
which did not come within the scope of the memoir by Sharp
and Muir. An investigation that would enable us to see the
whole range of variation and the degree of modification of
the genital organs within certain genera or groups of genera
in the different families would help us to form an idea of their
phylogenetic value, and seems to me to be a great desideratum.
But however neglectful the systematists in Coleoptera may have
been in the past, signs are not wanting to show that they are
now fully alive to the importance of examining these structures,
even if only to help them in their ordinary every-day work.
And to show that their importance had not been overioo^d
in the past, I cannot do better than to quote an interesting
passage from one of H. W. Bates's papers on the Amazonian
Longicoms, a passage which, written for the eyes of the
specialist, may not have received the general attention which
it deserved. The passage is as follows —
" It is a remarkable circumstance, that in many families
of insecto which have accessory sexual parts easy of ezamina-
Ij.y.l.AjL.yCOO'^lL'
( c^ >
tioD, it is found that these differ very considerably in structare
in closely allied speciea. It has been remarked that they
offer some of the beat characters to distinguish species, and
they have been made use of to separate species which scarcely
offered any other distinguishable characters. Mi, Baly has
also discovered that the homy penis concealed In the male
abdomen of Phytophagous Coleoptera differs in form in closely
allied speciea ; and he has shown me a long series of specimens
mouDted for examination under the microscope, belonging
chiefly to the genera Chrysometa and Ewnoljnie, which ofEer
a most instructive study, since by their means some forms
before considered as varieties turn out to be distinct species.
" This class of facts seems to me of great significance, as
throwing light on the segregation of varieties and their passage
into true species. For if we admit that the only sound differ-
ence between allied varieties and allied species is that the
former intermarry and the latter do not, then the abrupt
and great diversities of structure in those oi^ns most directly
involved in the matter must be considered as affording an
explanation why many varieties do not intercross with the
parent stock and therefore remain as independent forms or
species. The difference in the accessory male organs of our
two allied species or local forms of Colobothea is so great that
no one who examines them can believe both to be adapted
to the corresponding organs of the females of each form. At
the same time I have no doubt that, were it not for the great
difference between these organs in our two forms, no entomo-
logist would doubt their being mere local varieties of one and
the same stock. Scores of other local varieties occur in the
same countries, presenting all the successive steps of segre-
gation, from the most partial variation to the full-formed
local race.
" Thus we have only to admit that species disseminate them-
selves over wide areas, and adjust themselves to the diveitdties
of local conditions, or, in other words, segregate local varieties,
to open the way towards an explanation of the way in which
the world has become peopled by its myriads of species. The
inevitable law of Natural Selection which governs the general
process of the adjustment of the local races to new conditions
iM-,i,A J h, Google
( cxvi )
will explain the changes of coDditioos of life in tinne ; and the
laws of variation, diveroified in details as aie the spedee
themselves, vill explain the rest."
Mr. Bethune Baker in his Address had occasion to lefet
to the genera Iduhyunta and PsaUdura, but he could hardly
have suspected how interesting an illustration of the modifi-
cation of parts those genera afford. Ichthywms is a genus id
which I had, myself, previously been very much intoiested
for another reason. It belongs to the family Telephoridae,
and several years ago I had discovered in nearly all the beetles
of this family a series of small apertures, distinct from the
spiraculai openings, along the sides of the first eight dorsal
plates of the abdomen, a pair to each segment. In some
genera they lie well within the lateral margins; in others
they are placed close up to, or quite upon, the postero-lateral
angles, excepting the first pair, which always retain their
position inside the margin of the first t«rgite; and in a few
genera they are situated at the apex of very distinct lateral
processes which point outwards and slightly backwards. In
appearance these apertures are not unlike some forms of
spiracles; each has a circular chitinous rim bordering a small
pit lined with a pale membrane in which is a still smaller
opening fringed with fine hairs, and which evidently is the
external opening of a gland. Segnientallv arranged glandular
apertures of this kind are known in the larvae of some Coleo-
pt«rB, Lepidoptera, and other insecta, and the secretion is
considered to be distasteful and protective. Miss Olga Payne,
with whom I was in correspondence when she was working
on the structure and life-hiatory of one of our species of Tele-
fhonts, has found the glands also in the larva of that genus,
and in the three thoracic as well ae in the first eight abdominal
segments, and has given a description of them in a paper since
published ; but she has expressed a doubt as to whether they
are really functional, since the openings were ver}' small and
no liquid secretion seemed to come from them. So far, how-
ever, as the imago is concerned, there can be no doubt that
they are functional; for in handling some living specimens I
have myself seen drops of a clear liquid of pretty considerable
size issue from the pair of apertures on the eighth segment.
ih, Google
( Gsvii )
and 80 convinced wae I that the secretion serves &b a protection
to these beetles that I mentioned my own observations in
order to confirm the view that they are a distasteful group,
when reading a paper on " Mimicry in Coleopt«ra," since
published in the Proceedings of the South London Entomo-
logical and Natural History Society. Although many other
groups of insects aie at least equally distasteful, this is the
only one, so far as I know, in which a series of segmentally-
arranged glands of the kind described have been met with
in the imago. But to return to Icklht/urus. In this genus,
the apertures of the glands are present as usual, but the last
pair, the largest of all, are placed at the end of those strong
outwardly and backwardly diverging processes of the eighth
tei^it«, which gives it that fish-tail resemblance from which
the name of the genus is derived. It is interesting to
think that while I had been looking upon these processes as
a battery provided to meet the assault of enemies, Mr. Baker
seemed to see in them the arms with which the male is
accustomed to embrace the female. But, strange to say, I
am not certain that we were not both right. The processes
are well developed in all the species of the genus and in both
sexes; but they are more fully developed in the male, in which
sex also they sometimes have a sharply curved point or tooth
at or near the apex, and in such cases the glandular aperture
is to be found not quite at the apes, but ver>' near to it. They
are in most cases fixed pieces continuous with the rest of the
tergite of the eighth segment and only movable with the tergit«
as a whole, and this is the condition in the species referred to
and figured by Mr. Baker. In one species, however, namely
/. apicfiUa,* Mots., the male processes take on the form of a
pair of forceps, like those of an earwig, with the arms curving
inwards and coming in contact behind. Having examined
them with a view to finding out whether they were movable
from side to side, I found this to be the case, to some limited
extent, at least, and that they had all the appearance of being
articulated at the base ; so it seems to me impossible to doubt
that the fixed lateral prolongations of the eighth dorsal plate
met with in the other s[>ecies have here become so far modi-
• = forcipiger, Gostro.
n,g,t,7.dh,G00glc
{ cxriii )
fied as to form a pair of appendages adapted to act as
claspiog organs. They could not have served that purpose
from the beginning, but containiDg, as they still do, the
apertuies of secretory glands, their development at first may
have been to give a mote effective use to those glands, and
later coming to serve also another purpose their further
development may have been in that direction. The genital
opening of the male, I may remark. Ilea juat behind the
stemite of the segment to which the dorsal processes belong,
and is followed by another well-marked stemite which I con-
sider to be that of the ninth segment, though much la^r
fihan the corresponding tergite.
There are, as you all probably know well, two principal
views as to the origin of the wings in insects, and many ad-
vocates of each view. The arguments on both sides are very
well set out in a paper on the subject by G. Crampton which
appeared in the Journal of the New York Entomological
Society in March 1916. The author himself advocates what
is, I think, the now generally accepted view, namely that the
wings have arisen as out-growths or expansions of the dorsal
plates of the nieao- and meta-tfaorax, have gradually developed,
and in time became articulated at the base, finally functioning
as true wings — that their origin and development was, in
fact, very like what we see in the ontogwy of the termites,
bugs and other heiniinetabolous insects of the present day.
To this theory of their origin it has been objected that the
wings must have served some use at every stage of Iheir
development, and that it is very difficult to understand to
what use the notal expansions could have been put which
would have caused them to develop an articulation at the base.
Crampton effectively replies to this objection by saying:
" If an expansion of the integument can acquire an articula-
tion with the body when it develops into a tracheal gill in
the water, why can not a similar expansion acquire an articu-
lation with the tergum when it becomes a wing in the air ?
It is surely no harder to conceive of a rigid outgrowth becom-
ing an articulated appendage in the air, than to conceive of
a similar rigid outgrowth becoming an articulated appendage
in the water ! "
n,g,t,7.dh,G00glc
( cxix )
A Utile further on he gives a fen instances to enforce his
argument; and in reference to one of these, he says that
" On page 242 of hia Cours d'Enlonwhffie, Latraille, 1831,
has described a Coleopteron Aerodnus {sic) lotiffitwmus, whose
prot^orax bears articulated lateral processes {teste Cholod-
kowsky 1886)," and " If these can become articulated to the
prothorax without first passing through a' tracheal gill stage,
why can the wings not do the same ? To demand that the
wingB Imust pass through a tracheal gill stage in wder to
become articulated to the tergum, is asking more than the
facts would warrant, and in the light of the foregoing instances,
this objection to the origin of wings from paranota is not
valid." The reference given to Latreille is, however, wrong;
for in the work cited Latreille does not even mention the
beetle named, though he probably does so elsewhere, as did
most of the old authors when pointmg out the remarkable
character of ite prothoracic processes. For example, we find
it alluded to in Kirby and Spence's Entomology (Vol. Ill,
p. 538) as follows—
" But the prothorax has nwvable as well as fixed appendages ;
of this kind are those spines {umbo'tes) whose base is a spherical
boss moving in an acelabulum of the thoracic shield of the
Capricorn subgenus Macropug, Tbunb. If I might hazard
a conjecture, I should say that these organs were given to
this animal by an all-provident <,)reatoi, to enable it to push
itself forward, when in the heart of some tree it emerges from
the pupa, that it may escape from its con fine niMit."
Lacordaiie, however, who, in his Introduclion d I'Enlomo-
logie (Vol. I, p. 34, 1834), has also described the movable
spines of Aerodnus longimantig, gave a different version of
their origin ; " D'apr^ la loi qui s'oppose k 1 'Introduction
d'une piSce nouvelle, cette epine ne peut ctre qn'une des
pieces du prothorax qui a pris une forme inaolide, et son
analogue doit le retrouvec dans tons les autres Insects ;
aussi d'apr^ ea situation, penEona-nous que ce n'est autre
chose que le trochantin qui a cesse de faire corpa avec la
hanche, eat devenu libre et se portait au dehors, au lieu
de rc8t«r cache i I'interieur, a pris un developpement
inaccoutume. Du rest«, I'usage de ce trochantin, ainsi
l„y,|,AJI^,COOl^lC
( CXI )
mitamotphoB^ est ausei probl^matique que celui dee cocaides
dea M(Uachius."
These inteTeeting explanations of the movable spines of
Acrooinus Umgimanua should serve to remind us how far fiom
infallible aie even the most trustworthy of authorities, and
how careful we should be in our statements to make sure
beforehand of out facta. Many of you are, doubtless, familiar
with the Harlequin -beetle, and know what the truth in regard
to iXx prothoracic spines really is : that they aie not in the
least movable but are absolutely fixed and as firmly consoli-
dated with the sides of the thorax as the corresponding e|HDe«
in many other Longicorn beetles. The idea that they were
movable arose from superficial observation and was due to
the fact that they are surrounded each at the base, but tm
the upper side only, with a rather deep narrow groove; it
began with Linnaeus himself when he described the beetle,
and was never questioned until, just about a century later,
the mistake was discovered and set right by James Thomson
Mr. Crampton evidently bad no knowledge of that fact,
and he is hardly to blame for having accepted as true a state'
ment made on such excellent authority.
But though one of the instances on which he rebed is gone,
there is no reason why he should not find in the forcipate taol
of Ichthyunu another that would answer the purpose of his
argument almost as well. For if I am right in the interpreta-
tion I have given to it, you have there a pair of movable
appendages developed from what were once fixed lateral
processes, comparable with the prothoracic processes of the
Harlequin -bee tie .
In the long forcipate tail appendages of Psalidura, a genus
of Curculionidae, we have another example of perfectly mov-
able structures which have apparently developed from fixed
rigid processes. I say apparently, because, while they ars
free from the eighth tergile, to which they are attached by
membrane on the dorsal side, in one species of a related genus,
they seem to be more or less firmly united to the sternal
plate on the ventral side. But whether developed as prooesses
from the tergum or from the sternum, it is fairly certain that
they have not originated in any other way, and I have little
Ij.y.l.Ajl^yCOO'^lL'
( cxxi )
doubt that further mvestigstioo atnongBt the related forms
will prove this to be the case.
These remarkable Auetralian beetles exhibit a great range
of modification in the external parte placed near the male
genital opening, especially in the stnictuFe'of the seventh
stemit« and in the form and degree of development of the
forceps, the latt«T often being provided on the inner ventral
side with two blades which sometimes, curving downwards
and inwards, come together and overlap so as to form a tdnd
of ring or band, the object of which, so far as one can reason-
ably infer, is to grip the female right round the hinder end
of her body. The characters which these external parts
afford have been made full use of by Dr. Ferguson in his
revision of the genus ; but from what I have seen in the few
forms I have had time to examine, I think the aedeagus and
other internal structures would well repay, in their interest-
ing modifications, any amount of time spent in investigating
them.
Although I have been unable to add more than a very little
to our knowledge of the si^bject, I hope that little will help
to show the great interest which an extended investigatitni
of the whole genital apparatus of the Coleoptera must possess,
and bow completely justified was the opinion expressed by
Hr. Bethune-Baker in the Address to which I have referred.
I think we may congratulate him also on the happy selection
he made when he ventured to illustrate his remarks from
examples amongst the Coleoptera.
As a Coleopterist, I have sometimes wished that I could,
without too much trouble, make a comparison of the parts
in Coleoptera with the conesponding structures in Lepidoptera,
and I have turned more than once to look at some of those
nice photographic figures with which he and some of his
fellow-Lepidopterists illustrate their papers; but I generally
fail to comprehend them. I can quite understand that the
case would be altogether different if 1 were experienced in
the examination of the parts and knew all about their con-
nections and relative positions. I can understand also that
a photograph is more likely to be true to nature than even the
best of drawings; but if it teaches less truth what is the
u,y,l,/eJh,G00glc
( cxzu )
advantage? Prof. Huxley, in hia Anabmy of Invtrtthraie
Animals, says of the male genital armature in the cockroach,
that " It consists of a number of chitinous processes having
the form of plates and hooks, the exact form and disposititn
of which could be made intelligible only byDumerous figuiee."
There is much point in that statement, which api^ies as much,
I think, to many of the Lepidoptera aa to Blatta, and I hope
my friends will bear it in mind.
Goitlemen, there is another subject on which I should like
to ofler a few remarks this evening while it is still fnsh io
your minds, and that is the importance of E!ntomology in
relation to the State. It is a subject in which this Society,
if it is to continue true to the aims of its Founders, is bound
to take a verj- deep interest ; and, aa I need scarcely remind
you, it is one on which there are other views than your own.
There is, for example, the view which evidently bad influenced
the Government in their recent proposal to remove the
collections from the Natural History Museum and to stow
them elsewhere — the .proposal against which thq Society has '
strongly protested, and which, if carried out, would almost
inevitably have had the efFect of interfering greatly with the
scientific work of the entomologists of this country, and of
bringing to a standstill the valuable and important work that
was being done in the interests of the nation by the staffs
of the Museum and of the Imperial Bureau of Entomology.
Although it is satisfactory to the Society to know that that
preposterous proposal has been dropped, the mere fact that
it had been entertained by the Government will give you
some idea of what value and importance they attach to
Entomology. That science has probably formed little or no
part of their education, and they cannot be expected to see
as clearly as you and I how important is the work done by
Entomology, and what great benefit it would be to the State
if it were properly encouraged, assisted, and organised in its
service. There are many and various ways in which Ento-
mology has already proved itself to be of great use to the
State, and the possibilities for the future are also great.
This whole subject deserves your serious consideration, and
I very much regret that I have not come prepared to deal
n,g,t,7rJM,GOOglC
{ cxxiJi )
with it further this evening, but I hope to be able to return
to it on some future occasion.
And now, Gentlemen, I must thank you not only for the
patience with which you have listened to me to-night, but
alao for the great indulgence which you have been so ready
to extend to me at all times during my past year of office.
It is a year that I shall be able to remember always with a
deep sense of gratitude to the Fellowd of the Society.
n,g,t,7.dh,G00glc
GENERAL INDEX.
Tht Arahu fiyvris r^/rr to tht payrt of tkt ' TTaasaction/,' ; Iht Boa,t»
nvineraU tt the yoytt of llit ' Proctedipyt.'
Tlie PreaiJent's Addrcn is uot sepstotel; indexed.
GENERAL SUBJECTS.
Aberrant, wasp, eihibiteil, xlix ; Geometrid moths from Bsst Ijotbiin.
exhibited, liivi ; Hitmni ptnuaria. exhibited, luivi.
Acrata guirina, emission of fluid from the nDtfUDUi of, liii.
Africa, obaervations on Paasora in East, ili ; on a collection of L;pidopler*
made iu East, jlri, 323 ; synapoBematic seriea of Lycid beetle* of
tl speciet taken an one plant in one daj in late German Ea&t, exhibited.
Ivit : in burylDg its spider proy, metbud adopted by the Pompilid ffalv-
coaia fuliyiaeaai, pxliibited. Mi ; dragon fly-like Aiiilid flf of geuiu
LasiKKCaua from East, exhibited, Ix : Dbserratioas on the mimetic
females of Papilio danLinas in late German East, liv; of the genus
tfarangtaii resting in boles iu the ground, further observationa <m
Beiperidat from. Iivii,
Ayriadts thctit and other Lycaenids, condition of scales iu leaden mslea of.
xTi, 165.
Affiioi Claudia, local forms of, exhibited, iv.
America, od new and little-knonn Laijriidni from South, iii, 169 : descrip-
tions of Micio-T^pidoplera from South, 1.
Ammopkiltt .•abulosa, vaiiatiou in, exhibited, liii.
Ant, B{Hder mimic of, exhibited, Ixxri.
Antennae of closely related species, noticeable dilfereuce in, eihibilfd, xclii.
Apanleles ghnieralus, hyper-parasiten on, exhibited, xlviii.
Aphidat, parasites and bypet-pansiles of, tibihited, Iv.
Bato:onas fa/ii/iiiosiii iu burying its spider prey, metboil aclopted by the
African Pompilid, exhibited, lix.
Beetle. Death-watch, xvii, iijii, xiiii ; very ran British, exhibited, 1 ; lining
Dermestid larvae, exhibited, 1 : synaposematic aaries of I,ycid bertles of
9 species taken on one plant in one day iu late OeimaD Bast Africa,
exhibited, Ivii ; very ancient, exhibited, liiiix.
Birds, capturing butterflies on the wing at Oxford, ixix ; arc rarely witDeued,
observation explaiciDg why attacks on butterflies by, Ixii.
Braiil, predaceoiis Beduvitd bugs and Fossora, with their prey, from the
S. Paulo district of Soutb-East, exhibited, xixit.
ihyCOOl^lC
( CJDtV )
Britiib, Rhopsloceri, eihibited, iv; Klaltr, new, eihilnt^d, ivii ; BiiicMs,
plioUigT^hB of uTipoiiitots of three, eihibited, xii ; Cteidomf/idai, ■mllh
special refetcDce to the Nortbcro Oall-flies, preliroinuj utalogae of,
xiiT, 346; Cocvids, two new, ezbibited, iiii; beetle, Tccy rare, exhi'
btied, 1 ; ipeciineii of JVola^iifii bicaleria, eibibited, I ; CkrytopkaHUi
diipar TV. rulilm, exhibited, lixii.
Britisb Gniana, notes on butterSy migtatious in. iTi, IS-li Hymenoptera
(eiolosive of tbe Foraiicidnt), not«9 on some, 397.
British Uuseam, new apeeie* of Hirmeaoptera in the. 53.
BotterHieB, note on mimicrr in Oriental, ii ; in British Guiana, notea on
migrations of, xvi, 154; supposed hybrii^, exhibited, xvi; resemblance in,
mimetic and coD-mimelic, exhibited, iirtii : on the wing captured by
Inrds at Oxford, iiix ; taken in Eafit Africa, on a collection of, ilvi,322;
nuptial flight of, exhibited, H; by birds are rarely witn<j«sed, observation
explaining why attacks on, liii ; in midst of dry season, euggeated ex-
planation of wet-season forms of, liiii; another independent observation
of the " false heul " in, lir.
Bye-law. new, read, i ; special meeting to consider, viii.
Calij/o, genitalia of certain species of, eihilnted, lii.
Camphractua cihcIiu Uken at Oxford in I81T, lixiiv.
Crriilomyidat, with specisl reference to ihe Northern Gall-flies, preliminary
catalogue of British, xiiv, 346.
Ctloiiildat, from British India, new, exhibited, xiv ; additions to knowledge
of, ivi, 102 : from Hadagawar, eihibited, xix.
Ckatrocompa tlptnor, larva of, exhibited, ici.
ChiyaophaHoi dispar var. ratilu), British, eihibited, lixii.
Coccids, two new BritJuh, eihibited, xxiv.
Coleoptera, on unusual food-plants, exhibited, ilvi ; of Ihe Families Oilo-
midat, Monotemidut, Cotydiidat and Xotiophyaidat from the Seychelles
and Aldabra Islands, iciv,
Cotydiidae and Xotiophyaidac from the Seychelles and Aldabra Itlands,
Coleoptera of the Familic* Ostemidat, Movotamidae, iciv.
Council, detth of member of, announced, viii ; nomination of, Ixiv.
Dawn, reappearance of sunset insects at, xtii.
Death-watch beetles, ivii, xiiii ; exhibited, xviii.
Dermeatid Beetle, living larvae of, exhibited, 1.
Diptera, with special reference to the Qall-midges of the North of Boglaod,
preliminary catalogue of British Cecidomyida/, ixiv. 34(1.
Dytiiciit mar^taaJT), pupation of, eihibited, Ixiiii.
East Lothian, aberrations of Geometrid moths from, eihibited, tiivi.
Elalir. new British, exhibited, ivii.
Fellows, election of, i, xvi, xviii, xix, iiiv, ilvj, ilviii, li, lixii, liiv ; interned
in Germany, letter from, ilv.
Food-plants, Coleoptera on unnsual, exhibited, xlvi.
Formieidiif, notes on some British Guiana Uymenopter^i, exclusive ot the.
n,g,t,7.dh,G00glc
( cixvi )
Frey-OmDcr, Dr. Bmil, natiee o( d«*th of, ilri.
Gall-Biea, pnliminarr cutalogne of Brituh t'tridomjuidnr. tritti kprcul
Tcfereoee to tbe Northeni. nir, 346.
G«iutalu of eertaia >pecie« of Caligo, eihibileil, lii.
0«ometrid tnotlu from But LothikD, mbetrktiiHii of, eihibitod, liiri.
Gtolmpet tjiphiitiis, pupatioD of, eibibit«il, ic.
OereuDj, letter from fellow iDterard iD,>lT.
Gibbs, A. B.. Dotite of deatb of, viii.
Hemipldra, Sooth Indian, eibibited, *iij.
Hiiptriiat at the fmiu Saraigna reating in hole* in tbe grmuul, foTther
obaerratioiu do Afrioa. Ixrii ; aalt probably lon^t by. liirii.
Heterocfra from Hadagaacar. new *od little-knowD, iiW. S5.
Himtra ptinaria. dark abemtitin of, eibibited. Ixxvi.
HoDorary Fellow, death of, ilri ; nomination of, Iut.
Hjmenoptecm, in Uw Britiab Hnaenm, oew ipedce of. 53 ; (eielunTe of the
f Vnnir/dcK), Dates on HHiie Britiih Guiana, 291.
lodia, Hemiptera from South, eibibited, Tiii ; new ( rWw'itfac from Britiah,
eibibited, iIt ; addi'tiona to knowledge of Cttoniidtt of BritM, ivi, 102;
Pupilia fiolytet romaUa from eitreme aootb of, eibibited, lixi.
Insect life in Meaopotamiai, conditiooj of, Iiiti.
Loj/riiditt from South America, on new anil liltlc-bnown, iii, 169.
Ltitioctitnat, Bait African diagonBy-like Aiilid fly of genu*, eibibited. Ii.
Lepidopter*, from SilDnica,eihilMted, ii; made in Beat Africa, on a M^lection
of, ilfi, 322 : further not«a on rerapitnlalory attltudea in, iltii. ^38 ; on
the prolocerebnim of Minvpltiyx, 1 1 2.
Ljifatna an'on, bred, eibibited, Iv.
Lucatnidnf, inveBtigationa into tbe lile-hiatorim of »outh Nigerian. l>i.
Hadagaacar, Ctloiuidae from, eibibited, xii; new and little-known Ueterooen
from, ixiv, Sa.
Migapostra fatltni,tf\AtT attacking the Berce Fonerioe ant, eihtbited, li.
Mendelian Heredity in relation to aelection, two paper* on, Ixiir.
UeitopotBmia, conditions of insect life in, liivi.
Hicro-LepidoptEM, descriptioiis of South American, 1.
MicTopttri/x (Lepidopteia), on the protocerebmin of, 1 12.
Hicroscopf, gift of, xi.
Higratiuni of butterBies in British (juiana, notes on, Kvi, IM.
Mimicry, io Orirotal butterflies, uote on. i\ ; reaemblance in bntterfliea,
mimetic and noa-ioimeljc, exhibited, xxTiii ; another independent obaer-
VBtinn of the "false head" in butternies. In; obiervations on tfae
min-.etic females of Papitio Jardnia in late Germ-m Bast AfrMs. Hi ;
of ant by spider, eibibited, liivi ; in certain butterflies af New Oninea,
eibibited. ici.
3IoHotiimidiit, Colgdiidat and yoliepkj/iidiit from Ibe Seychellea and Aldabra
Islands, Coleoptera of tlie Families Ostomidai, leiv.
ilorpha adonis and M. tugtnia distinct species. eibibil«d, ixv; new lob-
apecies of M, rhiltnor, eibibjled, ilviii.
Mymarid Caraphraet«» riuct«» taken at Oiford in 1917. Iiiiii.
New Guinea, mimicry in certain butterflies of, ezbilHted, id.
Nigeria, infestigationi into the lite-histories of Lgeamidat tram Soutb,
ll,y,l,/eJh,G00t5lc
( cxxvii )
^aiiofiyiidiu from the Sejreheltes and Aldabra Ulands, ColeopterB of the
Families OHinuidat, Mmotomidar, Col^diidur aiii), iciT.
yotadmita birataria, British specimen of, eibibited, I.
Nnptikl flight of butterSiea, exhibited, li.
Obitouy, C.O.WalerboQse.i: A.K.Oibbi,viiiiDr.EmilFro;-OesBner,ilTi.
OflScers, nomination of, Uiii, Iiit.
OitBUHtlat. Manatomidat, Catyiliidat and Notiopkyaidae from the Seychelles
and Aldnbra IiUnda, Coleopter& of the Familiea, icir.
OnipDaitoTB of three Siricidi occuiriDg in Britain, photographs at, exhibited,
Oxford in IBtl, the Hymarid CaTaphraelvs rinriat taken at, lixiir.
/"aiiiVioprtaniKf, exhibited, iviii 1 forms of F. polgtti romWiu from Singa-
pore, exhibited, iix ; description of a new form of P. djirdamii, 322 ;
obsermtions on the mimetic females of P. daidanni in late German East
Africa, Ixt ; F. petglti rmnuiiii from extreme south of India and from
mainland opposite Bingapore Island. eibiUted, lixi.
ParaBitea. on ApanteUa ylamrratus, hjper-, exhibited, xlviii; and hyper-para-
sites of Aphidte, exhibited. It ; Chalcid panuite of Vavetta urtitae
naiting beside the laira preparing for pupation, exiiibi ted ; goldeo pupae
EHJt cansed by p&rasitism, Wi,
Fidiattui eapitii and P. hamaaui, further note dealing with question of
speciSc identity of, v,
Fitrinar, on a collection of butterflies talicn in £aBt Africa, with notes on
the, 323,
Prendent, death of former, auoouuced, i.
Pftnmiu atrli/ti; note on oviposition of sawfly, I.
Pupation, oF Dyliafui marginalh, exhibited, Ixiiii ; of Gtolrapei iyphaeas.
exhibited, xc.
TeduTiid bugs and Foubotb. with their prey, from the S. Paulo district of
^utb-East Braiil, predaceous, exhibited, xixiv.
Rbapalocera, British, exhibited, iv.
Salooica, I^pidoptera from, ribibited. ii.
Salt probably sought by the Hnperidae, lixvii.
Sarangeta resting in holes in tbe ground, further obsenatioas on African
Htiptridae of the genlis, Ixvii.
Sawfly, lanae, photographs of, ilvii ; Ptenmui seHifir, note on oviposition
of. I ; with " palisade,^' larva of. exhibited, Ixii.
Scales in leaden males of Agriiidts Ihttii and other Lycaeoids, condition of,
xvi. 165.
Seychellea and Aldabra Islands. Coleoptera of the Families Ostomidut, XUaa-
tomidar, Colydiidat and .Valiophj/iid^e from the, iciv.
Singapore Island and the mainland opposite, forma of Papilio pnlytii romulus
from, exbibiteil, xii. Ixxt ; descriptions of new species of Stapliylimdiu
Siricids occurring in Britain, pbotographs of or jpoaitors of three, exhiliited,
Special meeting to consider new bye-law, viii.
Spider, prey, method adopted by the Africsji Pompilid Batotoint) falifinoaua
in burying its, exhibited, Iix ; attacking the fierce Ponetine ant Mtya-
peHtra/otlm, exUbited, li ; mimic of ant, exhibited, lixvi.
l„y,l,Ajh,COOl^lL'
( cxzviii )
Temperature kt which iniect life dettrojtrd, liv.
f'lHMM «rl>M(, Cbilcid penute of, wutieg beuJe the lam prepariog for
pupttioD, exhibited, Iti.
Vim-PreaideDli, DomioatioD of.i.
Wup, kbemnt, exhibited, ilii.
Walrrhouse, V. O., Dotini of duth of, i.
n,g,t,7.dh,G00glc
SPECIAL INDEX.
abdominalia (Anthicus). 250
(SUtira), 2&6, 267
sbietiperds (I'erruia), 364, 413
abietis(AdelKe»), 369, 420
,, (Apliidolet«9), 389, 415
abnomiis (Huplijna), xvii
sbrotani(Mi«>patlia), SGI, 412
abruptaria (Heiiieropliits), 842 .
abuluta (Phthorim&ea), 44
■caotiiomera (Statira), 177, 26S
ocerata (Acraea), 328
„ r. Unella fAcraea), 32S
,, r. viiiidia(Acra«a), 328
acercriHpaiu (Peirisia), 3tl4, 413
„ vnr. rubella (Ferris
364, 413
aceiia (Atrichosema), 397, 416
„ (Caropjlomyia), 403
acptosae (Contarinia), 391, 415
aclisloia (Bjblia), 331
r. vulgaris (llyblia), 331
Acra«a, 330
Acraeinae, 325
acrita manca (Arraea). 3SS. 327
,. r. lidica (Acraea), 326
scrophila (Perrisia|, 364, 413
acaU(Cliit<>toba), 103
acatipcDDiilU (8f lepta), 99
Adelgea, 420
adipp« (Argynnii), Iiv
adonU (MnnGdiila),xiiviii
„ (Hoii>ho), Kxr, xxvi, ixv
„ adonis (Morpbo), xivii
„ iaUrmedia (Morpho), xxvii
„ race major (Morpho), xxvii
., (Papilio], xivi
„ (Stictia), xxxviii
aegrota (SUtjra), 202, 266
PROC. KNT. SOC. LOND., V. 1917
I oeneomarpiiata (Statira], 226. 266
aeneotincta (Statirol, 212, 225
I aeqiiorcs (Rceurvsria), 39
laerata (Protaetia), 111
I aerobatica (Cacoecia), 7
I oerufrinoaum (Teninosonia), 298
I itcsculi (Zeuzem), iii
'" 1b (Perrisia), 364, 413
iigamemnon (Papilio), Imiii
■ ;e (Plan^rua). Ixix
monUiia [Plunema), 329
A^riatidac
Agatliiuae, f
Bgathon race phrjxe (Aporia), xi, iii
aj^stor (Papilio), ix, x. xi, xii, xiii
, govindra (Papilio), ii, x.iii
_ «(Danai8).x
Agra, 189, 171, 182,200,207
agTacfomiis (Statira), 171, 182
agroides (Statira), 179. IHO, 181, 18«,
266
(Statyra), IBO
alirimancs (Aiio|iliu*),69
„ (PsainniochatM), 69
alliimncula (Argyroploce), 27
alliiniaeulaU (AmanriB), 324
alljipeDaia (Loaioptera), 349, 411
albitarse (Trypoxyfon), 317
albofasciata (HUtint). 257. 25S
alboguttata (Atiatoua), 105, 110
(Protaetia), 103
albolineata (iJtatira). 170, 266
albuniacula (Siugara), 96
albotcrminaliB (Pilodcs), 98
alcibiadea (Papilio), Ixxx
aloon(Lyca*na), Iv
AllodiplosiB. 369, 41S
alni(PeiTiaia), 364,413
alopecuri (OligotropliuB), 352, 412
iM-,i,A J h, Google
klpina (Pcrriam), 364, 413
■lUratua (Tsrucus), 272, 380, !81
altiBona (Plithoriiuseal. 4S
aliunilUu (Ca^ua), 6
AmsllectiB. 1
•malthei (Melipona), 30tl
Amathnaiidae, xxii
AmbljBpatha, 403, 417
Bmcric;aQB (FeriplaneCa). 320
tmethyBtina (PepaU), 313
ametina (Tacliyteii), 31U
AnietrodiptoHia, 402, 410
amicalta (Olyphodes), 100
Ammophtla, xiv. xixvi. ixiviii
BDioena (SUtira), 194, 196, 266
amwnulua (Ani>|itiua), 73
„ (PaammocharFs), 73
>moenus(LopliDircui), Iriii
., (Lycus), Iviji
ampliatus (Lyciia), Iviii
Ampuliciuae, SO
Anabremia. 390, 4tS
anacardii (Sal amis), 332
anacreon (Acrasa), 327
„ anacreontica (Acraea), 327
„ bomba lAcraea), 327
„ „ T- indiina (Acrsea),
327
auali'i (Eiiglosaa), 303
„ (PoTiaUs). 306
ananda (CaaUliil"), 289
„ (Tarucus). 270, 273. 289, 291
■ aatolla, xiv, 102
Andrei
^290
aDKlica (Perrisia\ 364, 413
an^uaUU (Sutira), 202
annulate (Chalcia), 320
(Statira), 252, 253, 266
annulatus (Ueniartes), iixv
annulipes {Harligiola), 3GS, 315, 414
aDUIua(VJrachoU), 333
antennalor (Joppa), 321
anlliiuoide* (SUtira), 261, 262. 260
,, var. staiidingeri (Statira),
261, 267
AnthicUB, 242
■athobia (Contariniu), 391, 415
Aulhopliorinac. 300
anthracina (AntiiolHKKu), 61
Anthrax, 316
Anthrenus, 191
Antichira, 3S7, 416
An tic li iridium, 397. 416
an till arum (Sutira). 230, 267
antilope antiiop« (Precisj, 333
I aiAriaea (Pirriua). 365, 418
[Aphidae, W
, Aphididae, 346
AphidiuB, Iv
I Aphidot?t«8, ^89, 415
I aptiiltra (Aristot«lia), 34
I Aphis, 390, 420
I Apidae, 29S
i Apiomerut, ixxv
1 Apis, icv, icti, 130, 306
I ApoicB, 310
appendiga9t«r(EvaDia), 320
aquatica(Leptura), irvii
aquilina(Phthorimaca), 44
Aranvus, xxxvi
Arwuthoniyia, 3&2, 412
nrcuatipes |3Utira\ 191, 192, 193,
104, 195. 266
arenacea (Poljchrosia), 23
arenanae (D^'odiplosis), S98, 416
argenteolineate (kphiitoaorpba), 57
argia (Leuceronia), 334
Ai^fllans. Ui
Argiopidae, 314
argosema (Geiechial, SO
argua (Plebeiua), ii
arion(Lvcaenal, Iv, 107
aristvlochiae, (Papilio), lixi, lixii,
' Aristotclia, 1, 29
annate (SUtira), 180. 181. 267
amoldi (Elial. 63
„ (Mesa), 63
Amoldia, 360, 412
nrrecta (Capua), 5
arrow) (CleroU), 106, 107
artcmis (Austrosatiua), 76
,, (F^rreolomorpha), 76
Arthrocnodax. 389, 41,'i
Arthroniacrs, 263
artipara (Phlegetonia), 92
arundiuis (Laaioptcra), S49, 411
Asaphes, Iv
' Asilidae, xli
aspasia (CyphoDoiiyi), 78, 80
„ (Mjgnimia), 78
! asperate (Sutira), 230, 233, 240, 260
I Asphondylariae, 385, 414
Aafhondjlia, 386, 414
aaaimilia (Ampulex), 81
AsUte, xxxrii, xiiviii
, aslrarche (Polyomniatus). ij
asvmnietrica (Stetira), 171. 190, 191,
266
^ Aijnapte, 402
I atelanU (Pyramfia), ii, Ivi
I aUlodva (EulU), 9
' Btevus (Anoplius), 89
iM-,i,A J h, Google
(
Dtaviu (Psammocharcs), 6S
atenima (Phjmatocera), ixii
atomaris (Ematurgs), iii
atricspitla (Eucosma], 21
Atrichoaeroa, 397, 416
alricoUis (Statira), 256, 267
atrifascU (Phtborini&ea), 45
AtjlodipIoHia, 402, 418
BUOUpanae (Perrisia), 369, 413
Aagochlora, 299
anloDOIa (AriatuUIia), 30
aurantiaria (Hybemia), iciii, xoiv
anreopicta (EuUlia), 91
aureosericeus (H^mipepsis), 80
anreotinctaltB (Pjrauata), 100
A (CtiD
ia). Ill
anroraria (Hjria),
aurulenlB (XylixKip), 300
auatralaaiaa (HemitiepBia}, 76, 80
(Mygnimia), 80
Auitroaalins, 75, 76
autumnaria (Ennomos), licivi
avenae (Mayetiola). 866, 412
Bvia (Calbptiryi), ivi, xvii
aiillarie (Permia], 36S, 413
aztecum (Eiphoaoma), 321
bsfdadenHta (Aethiesas), 102
Baldratia, 350, 411
balkanica (Tanicus), 272, 275, 276,
277, 278, 279, 281, 287,
278
, (Tan
277,
ballotae (Perruia), 365, 413
barbata (Xjlocopa), 300
barbichei ((jontarinia), 391, 415
bwiflava (Chelura), xiii
baailiciis (Peamniocliaroa), 73
baaimacula (Cryptochilns), 78
baaaianna (Peammocharcs), 72
bat«iai (Statira), 183, 266
hatillifera (Macronota), 108
Batozonna, 65
bauinaani (Cliaraxes), 330
WlfUiiafPerriaia), 365, 413
becga (GorKythion), n
, Iii, 1i
, liv, 1
, I»i
bellBVoyoi (Anabremiaj, 390, 416
,, (ClinodiploaU), SeO
Bembecides, xli
Bembex, ili, xlii, Ix
Betnbicinae, 316
Bembidula, 316
btUKalensIs (Tamcna), 281
beniDenais (ATnmopbila), xiti
bertlioudi (Psamraocbarfs), 70
BertierU, 362
)
BethfUdae, 63
betoiiicae (Clinodiploaia), 400, 41S
betulae (Semudobia), 3S4, 412
betulicola (Contarinia), 391, 416
betnlina (CoDtarinia), 391, 415
bicolor (Camuylomjia), 403
,, (Statira), 25S, 2S9
licoloria (Notodoata), 1
bidentatua ( Dyacinetua), 312
bifiiraaU (Statira), 253, 264, 266
bilmealA (Caniptogramma), iii
bilineolata vai'. niobtana (Nectarina),
309
bodhisattva <Clflrota), 106
boBtiouH (PoljoTiiiiiBtus), 833
Boiiibinae, 304
liombuB, 304
Bombji, xtix, c
botularia (Cliuodiploais), 400, 416
bowkeri (Ljcaena), 294
„ (Tarucus), 873, 294, 296
Bracbyneura. 388, 414
Itrachyneurariae, 387, 414
bracbyntei-a (Tliecodiplusis), 396, 415
Bracon, xxiv
Braconidae, Ir, 31 S
Brauoninaf, 318
braesia (Acraca), 325
braeaia (Acraea], 326
,. f. regalis (Aoraea), 326
brasilianoruin (Xylucopa), 300
braailiensia (Gorytcs), 316
[Statira), 250
bcBssicBa (Daayocura), 865
„ (Perriaia), 365, 413
,, (Pieria), ii, 339, 340
braueri (GcocryiiU), 360, 412
Breoiia, 389
brevicarinatiim (Trypoxylon), 317
breviuervU (Catocba), 403
brevipalpis (Stefaniella), 350, 411
brevi]iilis (SUtira), 263
brigitta (TerUs), Iv, 333
bHsbaneasia {Thytmua), 60
bninellae (Pciriaia), 365, 413
bryaiiti (Statira), 240, 241, 242,
266
bryoniae (Perrisia), 386, 413
buc«>La!a(PhBlera), 342
budda (CIcrota), 107
buddha (Calinaga), i, xiii
.. (Papilio), Ixxxii, Ixxiiii
buquetii (IjCUCL-roniat, 334
buraarius (OligutropbuaV 353. 412
bu\i (Monarthrolffllpus), 389, 415
cabira (Acraca), 328
Google
caaUto (Statin), 1S3, 196
caeruleipenDii (Statira), '211
otenilBotinota (Statira), 211, 212, 218,
217, 218, 288
oalamagTontidia (Lasioptera), 350, 41!
c-albuin (PolyKonia), iv
calduena f. neluaka (Acraea), 328
caliensia (Statim), 21S, 2fi6
faligo, xii, nx
callaxantlm (Bauiana), 91
catlida {Crociduwma), IS
,, (Ustoditilosis), 398, 416
callidia (Terocnfofl), I«v
callimorpha (Eiimenea), 312
callinara (Tanicus), 272, 274, 278,
279
iiigrn (Taniciis), 278
calluiia (Tac)jyt«8), xixvii
Calojihaena, 244
ca)o|>haenoidca (Statira), 243, 244,
24&, 266
ealthella ( Brioocphala], 114
„ (MicropteryiLi, 114
caly|>so [AuKOcliloral, 299
catiierona(Pitodeiidorix), Ui
camiiaimlae (CniituriiiU), 391, 41f>
(Mianut). sivi
,, (Perrisia), 361. 413
cani[«nu1abi (SUtira), 238, 268
CaiiipotiotiiB, xcvi
Cainpjloniyza. 403. 417
Campy lorn jiariae. 403, 417
canaaensis var. aiiiaaiiiiiiiiH (ri)]iat«s),
306
catialiciilaU rKimiKiira). 311, 317
canaliculata (Stalira), 213, 266
nandiope IChaJaxea), 330
caphusa (Aporia). xi
capitigenu (FerriaLa), 368, 413
laipitis (Poiiciilus), v, vi, vii
capnosUctia (Tortrix), 8
ca]>reaf (lt«<iiii^ia}, 354, 412
(U
m). X
r. nnjor (OUjtotrophua}. 354
caracana(StAtira), 219, 266
t^rapliractUB, Uxxiv
cardaniiDi'S (Perrisia), 368, 413
cardui (Phaetiolautliia), Wl. 414
,. (Pvranieia), ii. Ixxx, 332
(Trviictii).
is ( PeiTii
t. 414
(Tiichodipliwis), 397, 416
camea lUstrfiiiiai. 403. 417
carnrotinrla (Boraliu). >''i
caroiiliilatljisioptcra), 3.^0, 411
parpiui iZytJoliia). 356, 412
casDonioides (Statira), 170, 173, 237,
239, 266
oasta (Teracolus), liii, W
CaBUliufl, 270, 272. 273
catiphracta (Argyroploce), 25
cateoata (SUtini), 170, 171, 181, 182,
187, 188, 192. 193, 266
Catocha, 403, 417
CatochrjBOio, Ixij
Caupolicaoa, 300
c^averaoBa iStatira), 205, 266
cayenneiiaia (Bombua), 304
„ (Kugloaaa), 303
(Statira), 2S8, 266
Cecidomyia, 349, 35L, 352, 353, 361,
355, 356, 3G7, SS8, 359, 360, 381,
362, 3S3, 364, 365, 366, 367, 368,
369, 371, 372, 373, 374, 375, S76,
377, 378, 379, 380, 381, 382, 3W3,
384, 385, 388, 395, 396, S99, 402.
403, 416
Cecidoniyiariae, 388. 415
Cecidomyidoe, ixic, 346, 347, 411
Cecidomyitiaa, 348, 411
cetaeuB (UatochrrsolH), IxiJ
celimane (Teraculua), liiv, Ixv
c«utJiuKa« (Locwiola). 398, 41G
centralis (I.estodiploais), 398, 416
Ceiitris, 301
CVrambyx, icvii
Cerapliron, Iv
cerastii (Perrisia), 366, 413
Cerc«riB, xxxvii, xxxvtii, 316
cerviia (Uoliathopaii), 110, 111
CetoDiidac, xiv, xvi, \ix, 103, 103
Oetoniinae, 109
ClialcididBe,lv,S20
Ciialcidinae. 320
Ciialcia, 320
ohalcopt«ta (Sutira), 205, 266
chaniaediTa (PerrUia), 384, 414
championi {Statira). 170
chartariiiB (Chartergus), 309
Chart«rginiia, 309
t'hartijrfcils. 309
oliviograpta (Cacoecia), 7
CbeloniDac, 318
chiio (Acraea), 325
„ i-rystalliDa (Acmm), 325
cliloropt«ra (SUtira), 218. 266
clilorosonia ( Pseiidageiiia), 313
chlorotica (Pepais), 313
ChoristDneura, 349, 41 1
( liortomTia. 357,412
chryaanthemi (UiiDorrhyDcha). S4B.
349.411
ChrFBididae, 317
Cliryaidiiiae. 317
ll,y,l,/eJh,G00glc
( cxxxiii )
chrfsippus (Daoaida), 323, 324, 327
f. albiniu (Danaida), 323,
321, 327
„ r. Kicippoidea (Danaida),
Chrysia, 317
cbryaononic (Teracoliu), 334
chrjBothortii (Poljbia), 308
cJDCta (Veii]ia), xxxix
ciDctua (Caraphnctus), Ixxxiv
ciperea (Lestrnuia), 403
(dnenanim (RhalMlopLa^), 363. 113
cineTeuiD (Try poxy Ion), 317
circamducU (ijlstira), 217, 266
cirsii (Perriaia), 366, 413
dthaeroD (Cliaraiea), 330
clatlinta(Foniicia), 31S
clatliratiu (CarabiiB), xr
(Taruciis), 2B2. 293, 294
dandia (Agriaa), iv
,. f. anmonica (Agriaa), iv
„ r. sahlkei (A^rias), iv
cluuilia (I'errieia), 372, 413
clnvsta (Elu). S2
cUvifei (KhabdopliOKa). 3S1' 113
clavilineatuB (Odyneriia), 313
clu vines (Meli^miB). 304
cleodora(Eroiiia), 331
Clerota, xiv
ClinodiploaU, 389. 390, 100, 4:3
Clinorrliyni'lia, 348, 411
cloanthe (Catacropt«r*), 33'^!
obscurior (CatocropUra), 332
coelestis ( ICilchroea). xix
Coelioxynae, 303
Coelioxys, 303
C«1odera, 109
Coilodera, 109
Colaapoides, xxxvii
Coliaa, liv
ralombica (SUtlra), 2fi7, 267
Colpartiirum, 171, 212, 266
commixtuji (Cryplocl.eilii-.), 78
(Cryptochilus), 78
coni|nrsU (Pseudaf^vnia). 313
complaiiula (Lithoeia), iii
compoiiitarutn (Permia), .1R7, 413
nim|ir«iaicorpiB {I.ygaeonematiia),
ilvii, Ixix, Ixx
coiupresaiventris (Ampulrx), 81
ComjHoaaru, "'
Mlfector (Cj
mrerta (Epifai , ..
conGrmata (Oxrln'hiu), 39
conHicU (Polyhjmno!, 51
co»ru8(u (PUronibniB), 61
conigcni (Ampulex), 82
coniophaga [MycDdtploais), 401, 418
roiiiiexH (Stotira), 242, 296
Conocephaloides, xxivi
coDradti (Dicrogeniuni), 54
consimiiio (Ponipilus), 65
„ ( Psamnioctiares). 65
cunaoua (Gelechia), 50
ritt (Teplirosia), 342
ilata (8t«tim), 17S, 240, 252
:trU(Po]ybia),303
ContarinU. 391, 392, 395, 410, 115
cantigua (KphiituTiiorpha], 54
couvexiuaciila (Sutira), 22S, 266
convolviili(Protoparce), iii
cordatA(Euxl<MSa), 302, S15
Oorethra, ^xxxv
eoiidon (Agriadea), iv
,, ab. saiiu9yiigrepbB{Ag]'iBdes),
corui (Cran«iobia), 356, 412
coruirex (DiHhunuomyia), 368, 415
corona (DiH1uf{is), Ixxxvii
corniRana(MBerolabiB), 359, 412
curyli (CunUrinia), 392. 415
,, (Mikomyia), 351, 412
„ (Perrisia), 367, 413
«.ryliua(Pmri3iB), 367,413
(Stictodiplosia), 396, 415
Cosilidae, 53
cuamugrapha {AristvUlia), 36
coainoptila (CnKidosvma), 17
™atorii;en»ii (Xtatira), 170, 175, .
177. 266
ooNti|i«nnia (iHoduntia), 315
I „ (Sphex), 81B
{ Crabronidac, 80
j CrabroninBL', 83
nracuie (CotifatriDia), 392, 415
CTaiieiobia, 356, 412
I crii)»icornis (RUtira). 17'i, 223
I crawinen-a (Cyrtodiplosis) 389, 41
I oratiKgi (A]>uria). ii
I „ (Pmisia), 367, 413
I crawaliayi (Arapulcx), 80
, CremaJitoKaater. Ixi. 311
mbrici'iKi (Slatira). 2.^9, 230. 206
crimtuB(Poli>it«ii). 307
i-n«riPollis (SUtira), 256
CroddospniB. 1, 16
cruciata (Statira), 254. 263
crueiret»{Antlirafopliora), 102
craatariajPhtlinriinaea). 42
Crvptoubeilua. xxxvi, 78, 80
Crjptufiiilua, 78, 313
u,y,l,7rJM,G00glc
cucubali (ContBTinia), 392. 416
cupratA (Peronea), 14
cupreotiucta (SUtira), 21G
cupreoviridiB(Statira), 207, 208,209,
cuHpidata (Statiro), 184, 1S5, 266
cyaneua (Pouninu), xxi, xxii
cVanipennia (Statira), 2ri7, 25t), 261
Cyclopideg, 83 J
cydippe (ArgyDiiisj, in-
cylindricollia (SUtira), 260, 261, 266
cjmotoina (Argyroploce], 26
Cyuipidar, It, 346
cyntfiia (Ariatotelia), 32
Cyphoiionjx, 78, 80
Cyitudiplosis, 389, 41G
cyniB (Papilio), xxx, x\xi, Ixxx,
Ixxxi, Ixxxiii
Cy8tii*ora, 358, 359, 412
cjtberoea (Ariitotelia). 32
ilKtylidialMayotiola), 356, 412
daedalue (Hamaliumida), 331
dallBtorreaiia (Melipona). 306
datnovleef. darnoclides (Amauris), 324
Danaidae, 323
Danainae, xiii
daphliM (Perri^), 367, 413
daplidice (Pieris), ii
danlaiiua (Papilio). 336
r. ceiiea(i'apilio), Ixvii
,, r. hippocooii (Papilio), Ixvi,
llvii, 336
poIytrophuB (Papilio), 33S,
337
tibuIlUB (Papilio), 336, 336,
337
„ t. trLni«ui(Papilio). 336
,, f. troplioniiia (Papilio),
IxTii, 336, 336
darwiniaiia (Ep1int«nior|ilia), 55
Dasyneura, 350
Daa^proctu«, 84
davidi (Agalop«), xiii
davidii (Calini^ta), xiii
decora (Cliateria), 111
d«coratH(Euglo9Ha), 303
(NotOBorii), xxxvii
a«roIiaria (Hy1>ernia), xdii. xciv
degeeri (Rlinbdophaga), 363, 413
d^eaiii (Clielura), xiii
,, (SUtira). 211, 266
Delias, xcit, iciii
Deltoidae, 96
densata ( Ph thoriiiia«a ), 42
dentata (Cnephasia), 11
denticulata (Statire), 170,179
denticulatum (Podium), 315
dentigera (Stntira), 178, 268
dentipes var. ftavoscapiilam (I.yciu),
tM^ro-
lycus),
ileKtiptaiut (EuiMisiiia), 22
deapeclua (Roliathopflia), 110, 111
destructor (Mayetiola). 356, 412
deviucU (AniallectiB}, 1
dharta (Tanieus), iI91
diardi(UBcronoU), 109
DJatra«a, 318, 320
DichBlomjia, 362. 368. 384
Dicbodiploaia. 400
Dicbrona, 389, 415
Dicrogeniiini, 53
didyma vur. occidfotalis (Melitaea), li
didyniata a)>. cimeigera (lAmitia),
Didyoniyia. 353
Dielis. 313
Ilifflugia, Ixxxvi
di^DA lEacoxnia), 22
diluU (Statira), 210. 211. 266
diiiiidiata (Eu);lo»ia). 301
(Pepsis), 313
„ (Polybia), SOS
Diplosariae, 388
Diplosiuo, 392, 400
DiploaiB, 389, 390, 394, 397, 398, 399,
400, 406
Dirbiza, 402, IIS
diacisa (Beiubidula). 316
Disema, 264, 266
Dialiomiomyia, 388, 415
dispar (CbryBophaniis), iii, Ixxiij
var. nitiluB (Chrysophanus), ii,
Ixxii, Ixxiii, 341
disBimilia (Papilio). Ixxxi ii
diatiRma (Statira), 170. 235. 266
disttncta (ToiTDorrbina). 109
rar |-ilipes (Torynorrhina),
109
distinctiBaima (Cbryaia), 317
diverBJiicB (ICphutomorplu), 55
div'M(DicBifla), 111
dirinatoria (Atropos), xvii
divJBa (Stutiro), 21D, 211, 212, 26S
diylluB (Pilodeudorix), Ixi
l„y,l,Ajh,COOl^lC
doddi (ijiq>1iii9). 86
„ (Peaniniocliares), 6K
doiiertji (Drusillopeis), xxix
dolus TAr. vittatA (PolyoiDmatiis),
lefl, 167, 168
doDiratici (Muxca), xcvii
domeBticum (Anobwin), xvH, nviii
doininuls (Caltiniorpha), 16$
dorsata (Divlis), 313
doryca (Mj-nes), xcii, xciji
dor^cnii (Asphondjlia), 386. 414
droniioides (SUtira), 237, 238, 239,
288
Dromius. 160, 173. SSa, 249
Dnnophita, Ixxxviii, Ixxxix
drvobathra (Gelrahia). 19
do'obia (MacrodiplosiB), 3B9, 418
(Mocrolabin), 402, 421
dryope (Eurylela), 331
aD;culitt*(Eur;t^la), ^M
dryophila (Contarinia), 394, 41*
drypetis (Lethe). Isxx
dubia (Cecidoiuyia), 3S1. 413
,, (HjboiiiDnaa), 332
,, r. minis (Hvpulimnaa), 332
„ f. wahlhen^ (Hvpol<"">B8), 332
diibiou (Rhatidopiiaga), 361, 413
duplex (Xeocoenyra). 324
dureli (litycosia),' 109
davirivri (Lycus), tviii, lix
D^odiplode, 398, 416
eburata (Gelcchia), 50
echeria (Amauns), 324
echinus (Urentius), viii
ednsa (Coliaa), li, In
„ var. helice (Coliaaj, ii
egaensis (Statira), 234, 2S6
egeria (Pararge), tv
tfiflne, (Acrsea), 326
,, areca (Acraea), 326
harriBoiii (Acraea), 326
Eiphasoma, 321
Eiatj-r, xvii
elatii«(Pompiliia), 73
,, (Paammdchares), 73
eleeto (Colias). 334
elegans (Hoterorrhina), 110
(Sutira). 170, 221), 23.^, 236,
237, 244. 266
elefCantula (Statira), 170, 243, 266
Elidiiiae, 60
eliniinata (Sarangesa), Isi'ii, Ixviii
E[ia, 63. 313
ripeitor (Cliaer(icaiii{ia], xci
eluUliB(ryransta), 100
enccdou (Acraea), 327, 32S
„ r. alcippioa (Acravaj, 327
r. daira (Acraea), 323, 327
I encedon f. infiiscata (Acraea). 328
I. lycU (Acraea), 328
I eiiclitica (Fhalonia), 3
, Endaphis, 3S0, 41.'i
ciiKHtieldi (Perrisia), 367, 413
'Epeira, xxxvi
Ephednia, Iv
ephoria (Ariatotelia), 36
Kpbjra, 341
epieaate (Staphylus), ixxviii
Epidiarls, 301
Epidosia, 402
vpilobii (Perrjaia), 368, 413
Epinepkele, lii
Epipaschianae, 97
Epiayrnn, xxxvi. 84. 65, 67
Epi tragus, xxiiii
cpitricha (Phtborimaea), 47, 48
vqiiatarialia (Acraea), 327
,. anaemia (Acrvea), 327
equpatria (Haptodiploais), 399, 416
Eretia. IxriU
ericina(PerriaJa),36S, 413
Eriocephala, 112, 114
Eriucrania, 114
Eriupliyidae, 348
eriphia (KerpaeDia), lili
ei'ia (Teracolua), ixiv
erithoniuB (Papilio), Ixxxiij
eriacea (Papilio). xxix
eronioidea (Clielura), xiji
erubeecens (Parapompilua), xxxvi
erycina (Anstotelia), 31
i'rytbraa[ii9(Evauia). 320
erythronotfl (Anthobosca), 63
erytbropoda(l!:ti8), 62
„ (Mesa), 62
(Pleaia), 82
erjtbrostctbus (Ponipilus). 74
,, (Psa 111 mocli ares), 74
esebria (Acraea), 329
r. jackMini (Acra«a), 329
„ f. nioateironia (Acraea), 329
„ f. Dubilata (Acrsca). 329
„ r. prut«a (Acraea), 329
ctheoclcs (Cliaiaxes), 330
eiibtilc (Callidrjaa), 154, 155, 156, 167,
160, 161, 162, 163. 164
Eiicrra, 300
EiicoBDia, 16
Eucosmidae, 15
eugenia (Morpho), ixv, xxvi, nxvii
nraueU(Morplio),xxvui
Eiialossa. 301,302
I'Mgiossoiiles (MeliUiiiia), SOI
Eumenea, 311,312
Eumeuidinae, 310
n,g,t,7.dh,G00glc
f, 334
Kuri|>us.xu
cDiyderAiStetini), 219. 22% 224, 2Sll
eurjiantba ( Huphina), xc[i
evagore ITencoliiH), »34
r. Biiti)^iin (Teracolns), 331
cronesrens (8tatira), 253
Evaois, 320
Evrniiidar. 320
Evaniinac, 320
evidetm (Eucusms), 19
enocta (Plitfaorimora). 46
ExaeretP. 303
excerjita ( Epliiitomorplia), G5
exigua (Sutira), 25ii
exiniia (Caupolicaua), 300
Exomalopaia, 301
expulsus (Agenioiileui). 74
rxtricatiis (Tanicus), 272, 274, 27B
Taceta (Arfwroploce), 25
rBgi(MikiDla), 887,414
rasiiieiis (OIiKotrophuB), S.^S.lia
falcataria (Falcana), 338, 341
htkensteini (Uranotbanma), 333
bllal (Dirlie), 313
fascials (Anisoptera). xxxvii
■ ','. (Pu^bia),308
(SteKOiiiyia), xviii
faaciatiia (Plebpiiu). 202, 203
(TarucuB), 2lt2, 2C3
fastidiosuaoula var. saiiipniui(Poljl>ia),
faatutwa (Cunida). xlvi
faluelhis (ParnHni), Ixxvii
favosa (Melipoiia), 304
ffcundator (Aiidric;UB),42i
fcinoratis ( Lycus), Iviji, |[x
„ (Mrralycui), Iviii,
ferrugitifa {SUtirn), 24f>
fe»tiva(Kn<!o™), 300
„ (SUtint). 235, 2«7
fif(u1ua(RcHjphron). 3ir
A(P»iTitiia).36it. 413
filicornis (Ktatira), 2t>l, 262, 266
aiii«iidiilae(PvrriHU)..'i83
„ (Zyunena), iii
(InibriaU (Xyloro|n), 299
fiiiclicri (Hormoinyia), 388, 415
tinioulaU(Tortrix), e
ftrtulare ( Seel iph ran), 302, 314
llaTa (Cccidomyia), 399, 416
flnvJcomU (AUotria), Iv
„ (SlAtira). 213, 323
naTipenni3(Me1i|)ona].305
flavipea (Cainpylonijia), 403
HavnfaseiHta (Uacronota), 108
„ (Sotiistocerea), xxxvii
11avopioU(ElU).313
flavoscutellatiiB (KeiiicoapiluB). 321
flaToaiftnatd (Statira), 175, 177, 2tMt
Havovttr«ta (SUtira), 256, 257, 2<iT
flavua (Cyplionoiivx), 7»
flesuslTagiadesJ.SSi
fleiilinennn (Peranra). 14
tlorica (MisoiAtha), 351, 412
llorip«nlB (Perrisia), 368, 413
flonim(Miaoi«tlia|. 351. 412
Hosouloi iim (Perrisia), 38R, 413
i lluctiiaria (Cidaria), iii
fluctuatu {MelBiiipp«1, txxvi
fluTJalis (TiruciiBl. 202, 293
foeUns ( Mp^^poiiera). Ix
folioruni (Minnjifltlia). 351, 41:1
foreslaii ( Rhopalocampta), Ixivii,
IxxHii
Forficaln, 124,134
Formica, xcvi
formosa (Daiiaida), 323
(StBtira). 196. 2«IS
foniiaitB(pBelid«lteriB), 6
Fornicia, 318
TortiBpiiiua (La4io)ihonui), 31S
FosBores, 312
t'oveirollis (SUtim), 236
fragiliconiis (Othryadw). 202
fraxincn(DaBTiieural. 368
„ (Perrisia), 368. 413
fraiinplla (Arthrociiodaxl. 389, 41S
frsxini (EriophyeB), 389, 410
„ (Perriaia). 368, 400, 413, 119
fivya (Tpriomima), 333
fiiteiii (Hormonivm), 388, 415
fronbilii (Poiupifus), 70
„ (PBaiiimocliarm), 66, 70
(Sphe>). 70
rrurtunni (Perrisia), 368, 413
filKax (Tryi»xylon),:tl7
Fiil){oridap, ix
fuliginoBUB rar. iwpillulinilia (BbIaeo-
riilva (Stjitira). 229, 230, 266
rnlrieoHis (Anthiciis), a5«
(.StaUra), 2.M, 237, r.R. 2fiU
„ var. dismidalis (StAlira),
256, 257, 267
riilviUrsiB (Stciiooonis), xxxv
fdvoraBciaUfPotybia). 307
riilromneulata (Smicni), 330
fuBCB (Le^tremia). 404, 417
„ (Sibine). 318
„ (atatira),177
fiwcicomU (Palybia), 308
u,y,i,Ajh,Cooglc
( "
fnscipalpis (MeffAfhogas), 320
fascipennis (Antliicus). 2&6
(Statira). '^56. 267
fuscitATsis (Stetira). 162, 193, 2m
gnltwnea (Plialonia), 2
gsleobdolantia jPerrisU), 36», 413
^lii (Peirigift), 369. 413
„ (Trottari»),349,4ll
g»liicola (PerrisU), 369, ^13
galioram (Schizomyw). 38r>, 414
l^llomm (Dichiona), 389, 4t5
gsUi[>er(lB (ParBlleloJiplosiH), 400, 41
canthin
e, S14
gBuUiiia (CnliDaga).
gui (CoiiUrinia). 392, 41:
tCeUUIU(Tjii»a), xcii
Gelechiadae, 28
geminata <Jo]>[>a), 321
gammaU (Statira). 179.
(SUtj-ra), 189
267
gemnilfer (Stati
g»nuulata(9tatira).170. 1'
188, 189, 190, 196. 266
geuutae{As|>liandyliB). 386, 414
gtnistam torque ne (Perrisia). 3»9,
senisticola (Perriaia), 369, 413
Keutianae (Perrbis), 370, 413
Geocrypta, 380, 412
GemnetriciHe, xiix, 8S
Geotruj>e3, xci
geranii (Dasyneura), 370, 40S
,, (Pcrrisia), 370, 40G, 413
f^nnaiiica (VeB]ia), xlix
gerstackeri (Coliorthnim), 246
giurdi (Zetixtdiplmis), 397. 416
giardiana (ThefodiplosiB), 397
,, (Zeuxifiii'loHis), 397. 41
g{dica ( Belenoia), liv
«2,2t
l3(8Lt1
.. (Zetlius], 312
girandi (Pernsia), 378. 414
giraudiuDa (Rhabdophaga), ItKl. 413
glabrata (SUtira). 170. 179, 208, 209
glahrJRollis (Ftcroiiibrnii), 61
gladiata (Pulyhymnn), !il
glflchomaa (Permia), 371, 413
gloliifcra {Campv]omj7a), 403, 417
ftlubosa (ExomafopsU), 301
IClomeratiia (Apantelca), xSviii
gloriosa (Hypocaljnmia), 90
., 376, 413
Goryt«s, 316
Gorytinac, 316
goyawnsia (SUtira), 254
tvii • )
gntnilU (SUtira), 264, 26G, 266
gramiiiva (Aiigouhlora), 299
grammicft {Coscinia}, iii
,, (Lycaenesthes), 283, 234
,. (TarucTia), 272, 288
grajidia (Horntotnyia), 368, 415
,, (Omiuatothelxis], xxix
granifei ( Pseudhormoiiiyia), 39S, 4H
gratioaa IStatsre). 213, 214. 221, 266
gravida (Monedala), xxxviii
,, (Stictia), xxxviii
gregalis (PhtLorimaea), 43
gri9ealia(l'~ilodpa), 98
griseaU (Perigea). 8S
giiseicollin (Ceciduniyia), 331, 413
grasaulariata (Abraxas), iii
Gryllodes, ixxvii
Otyllus, xixvii
gmanae (HeU|>ona), 30G
guianense (Rhopalosoma), 313
guttatiui (Parnam), lixriii
Hadmbrcmift, 390, 415
Hacniatopoltt, ili
Haemonia, IJb. 266
haeirioDiDidea (»tatira). 265, 266
haitiensis (Statira), 2b5, 266
halimede <T«racolus), 334
lialterata (Cani)-ylouiyza;, 40»
HapaloaariH. 37
Haplodiploais, 399, 41S
Hannatidia, 391. 400. 416
liamiodiua (Paiiiliu), xlix
haroDa (Oiyiialpiis). xli
harrisi (Riiploea), xii
hartigi (Oligotropliua), 353, 412
Hartigiola. 385, 414
hector (Papilio), xxxiii, Ixxi, Ixxxi
heliaiithcnii (ColitarinU), 392, 415
Helioonidoe, xixix
helioiantlia (Amorbia), 12
bellwigi (Chortoniyia), 357 412
Ilemipepsia, 80
Hi!t>Lco»pilus, 321
lieraclni (ConUriuia), 302, 415
Ilfsperidac, xxxviii, xli, txxt, 337
hrtcrabia (Kliulidopbngii), 361, 413
HeteiooBra, xcviii
leUr™-)n>ii[Fl9). 61
(M «a), 61
I rogan nfbU). •!2
Ilet* on all » xxxvi
Heteropez nae 404
H » son A Ix
1 aWfF -jt la), 331
1 ta (F rrtcla), 331
hieraeii (CysUpbora), 358, 412
„ (Macrolabis), 359, 412
iM-,i,A J h, Google
hindei (LfucDi-liitoDea}, xl[
hippocrejiidie (Mocrolabis), 35S, 412
hirtella (Epiooinetis), ]03
hUtrio (SWtyra), ITS, 287
hoici (Hajretiola), 36l1. 412
holostrao (Perrisia), 371, 413
homo^hjla (Tortrix), S
hononus (E[)itfllii), hi
hopei (Euploea). lii
hord«i (Mayetiola), 367, 412
Uorniomyia, 85J, 889, 398, 415
baouniu (Pediculus), t, vi, vti
humeraliB (Stetira), 230
bumiliB (Agenioideiu), "Vs
Huphina, xcii
hniliiui (DielU), 313
HyantJB, xxix
hjtilaeoidea (Bleniun), 92
)iTgrophiU(PerTiBiB), S72. 413
HjmeiiopUra, 2ST, 298, 311
hyperioi (Perrisia), 372, 413
hypochoeHdig(.Stictodiplo«iB),39<^,41.'i
jcsrus (Polyommatus), li, iv, I&7
ichaeumoDcus (Sp1j«x], xxxvi, 315
IchiieumonidHe, 321
IchneuinoiiiliBe, 821
Idiops, xxxvi, xl
itporate (Perrisia), 372, 413
ilici* vtt. cerri (NordmBUnis), ii
„ (Theclft), ii
ilionelu ilioneus (Caligo), 330
ilithyia (Byhlia), xii, 331
illoqucata (Capua), 6
illaniinata (Sparganotliis), 13
illunaria (Selroia), 343, 345
illiMtraria (Seleiiia), 343, 344
imiUtor (CeiverU), xxxvij
imminensfSpilDDoto), 15
impavida (Potosin), 108
„ (Protaetia), 103
impendens (Crovidosema), 17
impolita (Argyroploce), 28
impresaipcnnis (Statira), 171, 206,
207, 210, S66
inchbaldiana (Perrisia), 372, 413
iiicisicDllJB (Stiitira). 248, 206
inclusa (Herrina). 372, 413
incon«l«ns(Statira). 212
inousa (Cuepliaaia), 12
inrerualia (Montezumia), Sll
itifunrnta (8curio]isin). xxjx
infundibuiironnia (Matitpnumia), Sll
ingena (SUtira), 170, 218
inaignu (Acra«a), 325
,, giRinna (Acraea), 325
insularis (Statim), 217, 228, 22S, 268
interrupt* (Melipona), 304
intemiptcpostata (Statira), 180
olVan
»),i
tpliiaulax, 318
Iridopsis, Ixi
' 'iia(S7no«4ra),310
la (Delias), xcii
pgularia (SUlira), 257
isognunmata (Tephroclystis), 342
iteoliia (Prrrisia), 361
., (R)iabdopha|(B), 361, 413
lt«aiu^ia, 354,412
IthomiiDae. xxiix
Itoplectis, 3til
jacobaeae (Stictodiplosia), 396, 415
JaoeUclla, 355, 407,412
jason (Papi)io). Iixx
Jasaidae, xxxv, Ixi
jcrmjiii ( Psrtiffltina), li, xii
juanDisi(Mayetiola), 357, 412
jobaea (S'ephemiiia), iciii
johnstoni (Acraea), 829
,. f. courasa (Acraia), 329
Joppa, 321
juc:nn(la(Pei)8iB), 31S
. unci ( Uroiuyce»), 401. 420
uniperina ( Lestnmia ). 404, 417
uniperinus (Oligotropbus), 352, 412
ariiiri{Polybia).308
iirtioa ( Epmephelp). iii. lii
. uvenons ( Taurunis), xxi, xxii
:arechi (Khabdophat^), 361, 413
kdlueri (Perriaia), 373, 41S
Itipfferi (PcrriaU), 373, 413
Kiclfel'ia, 385, 414
kicfleriaua (Perrisia), 373, 413
kivuensis (Aeraaa), 326
klugi (Cliutcria), 111
„ (RUabdoplis^), 362, 413
klugii {Euplooa)< xii
kneuckeri (Honnoniyia), 3S8, 415
kriatenaeui { Ampulex), 81
kurandeiiaia (Anopliua), 64
„ (ICpisyrou), 64
labiatiia (Mischocjttonia), 310
l8bilia(Poinpilus), 72
„ ( Psammncbar^a), 72, 73
laboriosa (Profopolybia), 308
labl-oss (Centris), 301
Lacliuocuema, Ixi
l^o^La (Gel«c1iia), 48
lacliiis (Sarangew), Ixvii, Ixviii
lacvinodis (Mynuica), Ir
la«VLUsciili (Xeuodiplosis), 399, 416
laeviusciiltia (Nenroterus), 3^9, 420
LaKria. 170
lamii (Peniaia), 373, 413
iM-,i,A J h, Google
Lainpides. 270
luisUi;Ue^hiie), 303
lui^Di (DicbodiploBis), 400
Unipes (Apiomerus), xxxv
„ (Centris), 301
Un (Delias), xcii
lariuiB ( Pemsi&), 373, 413
lariasB (MelaiiDirgis), iii
Lht™. 316
LBrrinae, 31fi
LasiocnemuB, Ix
Laaiophorus. 318
Lasioptcra, 349, 350, 410, 411
LusiopterBritt*, 346,411
LMiopterjx, 387
Laxiotliyris, 4
lateralis (IStatira), 211, 212, 216, 22G,
22fl, 227, 266
UtenilU (Cecidomjia), 399, 416
lateiitUta (Statin), 223, 234, 266
latlionia (Issoris), ii
Uthjri (ConUriuia), 392, 415
,, (Perrisia), 873, 413
lathyricoU (Perrisia). 373. 4 IS
laticoliia (SUtira), 179, 197, 198, 199,
217,219,266
laUl«H (Catocha), 403, 417 *
LecBuiain, xxiv
lechegTiana (Nectarina), 309
leda(Eroiiia), 334
,. (Physcaeaeura), 324 !
leiliu ((>dimon), 155
„ (Umaia), 155
lemeei (Janeticlla), 35ft, 112
lemalca (Spal^), 333
lenta(PlitWimaea}, 41
lenticukris (Neurotus), 400, 421
leonidaa leonidas (Papilio), 335
leontodoutis (Cystipliora), 368, 412
lenpardua (Tanicusj, 292, 293
lepidobirtua (Anopliua), 64
(Episyron), 64
Lepidopteia, xcviii, xcix, 124, 127,
140, 321), 322
iBpricuri(Mouleiumia), 310
Leptura, xrvii
Le>t«diploals, 398. 416
Leatremia, 403, 417
Lastremiariae. 403, 417
Lestreminae, 346, 403, 417
leacanthemi (Clinorrhyucha), 349,411
leucomelaB (SindriH), 98
lencolnesaaa (Eacosma), 23
leucophaea (Leslr«mia), 404, 417
lencopicU (Trachpa), 90
levana (Areachnia),
libera (Perriaia), 374. 413
liebeli (Honwli ptosis), 402, 416
ligncaCrortrix), 9
llKOslri (Schixonijin), 385, 409, 414
,, (Sphinx), itcix, c
lUaciiia (Staiiropus), 85
liinBtuU(LasioLhjTia), 4
limniace ( Danais), xii
ItmDoria tareta (Precis), 333
limpidiiK (Anopliua), 67
,, (EpiaTroli).67
„ (PaanimocbareB), 67, 68
linariav (ConbU'inia), 393, 415
lineata (Melipona), UOS
litliartcfroidca (Borolia). 89
litJioBperrai (Perrisia), 374, 414
litigioea (Gel«;bia), 49
liviaa{Ainphipyra), iii
lobiUirsis (MegBcbile), 303
Locuatidae, xxxvi
loewianus (Oligotiophus), 353, 412
■.oewiola, 398, 416
loiigiceps (Statira), 180. 181, 266
longicollia (Statira), 136, 187, 191,
200, 266
loii)(ioornia (Eucera), icv
longimana (Centris). 301
loHgipes (Pomeondyla), 402, 416
lougiTcntris (CI [nodi ptosis), 390. 415
(Hadrobreniia), 390,415
louicerearum (Contarinia). 390
(Syndiploaia), 390, 415
Loplio^ra, 98
Iwiuax (Pbtborimaea), 45
lotliariagiae (Pcrrisia), 374, 413
luti (Coiitarinia), 393, 415
„ (Perrisia), 374, 413
lotieola (Perriaia). 374, 413
loiiisae (Tarucus), 234
liicretia cxjiansa (Pscudacraea). 332
luctifera (dlyoosia), 109
luctiioaa (Acoutta), iii
lugene (iWumemus), Ixi
lunana (Enuomos), 344
luimlata (Statira). 251, 252, 267
Inpulinac (Aejibondylia), 386, 414
(Perriaia), 374. 418
lurida (Epbutomorpha), 55
lut«a (Cinihex), xlvii
lutosulana (Tortrii), 7
Lycaeaidse, lii, Ixxx, 333
Lycaeninae. 273, 275
lyclniidifl (Perrisia), 375, 413
Lycidae, Ivii
lyuoa (Acraca), S2fl
„ f. rallat(Acrava), 329
Lyman triidae, xx\x
u,y,l,7rJM,G00glc
mschaon (Papilio). n
Hscrodiplosia, ^99, 416
MacralaW, 359, Hi
MscronoU, 109
maculstiu (Camponotiis). Ixii
„ var. molanocncmiB (Csin|ia-
iiot<i»), Ixi
maculiceps (Henicospiliis), 321
niscuUpes (Pompilus), 75
maindroni (Macronota), 109
major (HentcDspilua), 321
„ (Iteomfia), 354, 412
,, (Piimnia), 401, 418
malgaasica (AcoBm^tia). 89
,, (DaaWKihaeto), 87
(Uacalla). 97
,, (Nigramtiia), 94
„ (StauropuB), 35
,, (Sylepto), 99
m&li (A|.hi8), 389, 420
,. (Perrisia), 375. 413
mtiliguuB (Aiistrosaliiis), 7S
,. (Salius). 76
malpigliii (PerrisU). 375, 413
malvae (Syrictithiwl, iii
marcu8(Uorpho), xxvi
„ iiiarpiw(Morplio). ixvii, xKviti
„ (Papilio). xsvi
,, (Zfliixidia), xxvi
niargariU{AppU)i), 162, 163
margnritacca (Belenoia). 335
mnrgiiialia ( lly tisciiB), txxiii
niargiiiatiu (Spliex), xiiv
inargineinton)u?iLH IPRrriaia), 362
,, (Rhnhdophaga), 362,
413
inart^fi (Macrolabis), 3
masliuiia (CatochryBO]is), txii
Maijwga, xxix
massaEitm (HypoolirmiH), xxix
SlassaTonjiUi 401, 416
inaiira (Manin). iii
iiiaxillosiis (Conace|>hatoidra), xxxvi
niaxiii.iini (nivroKcniiim), 53
maypri (Asphnndylia). 386, 414
■ Majetiola, 356, 357, 412
mcitresi (Macronota), 11)9
modialis (Statim), 2^4
mvdiatiUH (Iphianlax), 31S
incdiata(Miitilla), 312
ineditMigiDui (Contarinja). 393, 415
-" ■|i8),372, r-
roednu (Aricia). ii
iiiediisa f. alucsta (Nychrtona), S35
Mi^gaohile, 303
MpKachiliiiae, 303
me^era (Pararge), iii
MegalopU, 299
Megapouera, Ix
MegarliOKaa, 320
Megiatias, xixviii
iiiriancholicus (PampiliiB), 71
,, (Pwiniuiocliarfs) 71,72
ni«lait»>(Daiiais), x
mdanecta (Eulia), 10
melanocaDipU (Phtlioriniaoa), 14
iiiclanocephala ( Pericliata), xirii
(Statira), 229
melaiiocera (Coutarinia), 393, 415
nielauoidea ( Daoaia), x
melanoleuca (Trii^Dalvs). 317
nitjianoptora (Sutira),'2S0, 266
nielaaopiis (Aspliundylia), 386, 4)4
nidcap'is (Statim), 171, 181,187, 266
Meli|iotia, Ixxviii, 904, 305
Mplitonia, 301
mellirera (Apis), 306
mehifica (Apis), XXXV
m^TlonelU (Galleria), 306
iiiemnon (Calijfo), xix. xx
mdnis (Caligo), xx.
,, atlaa (Caligo), xjx, xx, iii
,, epiDi»theus (Ca1i>^), XX, xi
„ mi-muon (Caligo). Hi
,, pronietheua (CaliKu). xi^
iciialcaa (PotyoiiiinatiiB). ir>;. 168
ipBtntina (llclenois), xxx, liv, Ixii
liv, 336
irlla (Setinn), iii
(Cerceris), 316
„ (Trachypua), 316
var. liigi]bnH(/ftliiiii), ni3
Miastor, 346, 404
iiiicanB (SUtira), 195. 209, 2<i6
mirrorepliala (Kiioiii1>»n-|iina|. 109
„ var. Btaiidingcri (Itliam-
borrliina), 109
iiiicTops (Sfntira), 235
Mlcroptcryx, 112, 113, 114, 118, lift,
120, 121, 123, 121, 125, 133, 134,
135, 138, 139, 140, 141, 142. 144,
145, 146, 148,152
niiki(Perriiia), 375,413
mikii (Asphondylia), 386, S93, 414
Mikiola, 387, 414
Mikr.ni3ns, 364, 412
u,y,i,Ajh,Cooglc
( cx!i )
mtllrrolii (Clinorrhjnclia), 349, 411
„ (RlioiialoiDyia), 3M, 4)^
n.inimii{aiycypUiwJ. 110
minor (Sta|ihy](w), xxxviii
Minotaurus, xci
nimiiscula(H¥jioiwrigea), 87
minutum (TnchogTommit), 320
luiras (Amhub), 333
MisohocjttaniB, 310
niisippuslHyjioliniLaH), 331
f. inaria (kypolininas), R23
Uiaopatha, 3S1, 412
mitroUfHeUroirhins), 111
modfsta (CliutPiia). 108
nioliuUe (Cliorlomyia), 357, 412
MonarthTOpslpus, 389, 41ft
Moiled Ilia, BIS
Moiiodi|>li)t<i», 402, 416
MonUziimid, 310
inorbillofia (Stetyrn), 195, 19(1, 2S7
mordax (Plialonia), 4
rnori (Etombyx), xrix, u
raomsella (Acara). 911
manitanH (Gloraina). xli
mucronaU (Statira), 182. 1H4, 26<>
muiri(Psatntnncl]arei), 67
mulnilwr (Kuploea), xii, xiii
{Ti'eiisicbroia), xiii
multifoniiiii(Statii'a], 217
muUiiioUte (Statita), 248, 2fl6
murioatae (PerriBia), a76. 414
nmsaftntrs (LyRa«nvalhps),]xi
mntuta {M^lijiona), 306
Mutilla, 312
Mutlltidar, 54,312
AIyi^odi|>toitis, 401, 416
Myjtnimia, SO
Myrtcumyia, 397, 416
mysU tD«liBij), xcii
liais(EiiUialia), xii
napi(l>iflri8), ii, xxx,.340
nara(L7caena), 273
„ (Tanietia), 269, 272, 273, 274,
275, 276, 279
naBidenx (Odynems), 312
naatartii (Coiilariiiia), SftS, 4IS
natalica naUlica (Acrapa), 327
natans (I'olyncrna). Ixxxiv
tieaiithes (Cliaraxcs), 330
Nectariiia, 309
nef(1ects ( ['otosia), 103
„ (Protaetia), 103
neobulc (Acraca), 325
Kcocersta, 3S0, 411
neogihron (Ejiliafdro), 331
„ f. Tiolacca (Eipliacdra), 331
Neptii, xiix
iicrvicola(PeiTiEia), 376, 414
n«Tvoruni (RliahdopliaKa), 362, 413
iiiavias dominicanus (Amanris), Ixv
324
nicolayi (Conlarinia), 393, 415
nigella (Statira), 177
nigra (Statira), 183, 199
niKi'CBcciia (Pepnia), 313
iiiRricepB (Monteiu
nigroaenea (Statira), 263
iii^roapicalJB (Stutira), 258
iiiRrocaflruleafStatira), 182, 183, 184,
266
nif[ropllnctata|Epicuiita), !ixxv
nigrospitrsa (Statira), 197
iiircua(PHpilio), 336
„ lyaeus(P8i.ilio), 336
iiilidiventriB (Sphex), xnxi-i, xxxviii
iiitidiim {Try poxy Ion), 317
tioctilio (PauiuniB), xxi, xxii
noctiluca (Cantham), xcvii
(Umpyris), xcvii
ncidiili (Rhalido]>lia^), 382, 413
ncimaea (Arwroplmc), 27
Nomiiieia, 53
ncimiila(Papi]io), Ixxxiii
NottidoiitifW, sr.
Notof^nia, xxxvii
imlieciilosa (SUtirj), 2in
niinKi([':liK), 60
„ (M™).flO
,, (Piesitt), fiO
Myiii|>tiBlidae, 325
Nymphalinao, xiii, Ixxxiii. 330
oleosa (Volticetla), xxxviji
olifiiauata ( Laaiopteryx ), 387
i.l)id-'nsi»(Polyl.ia), 308
obliiiiiiraBuiata (Scalniicaiida), SB
obsimm (Statira), 263
obBcuricarinatlin (Iphiaulai), 319
occidcntalia (Polylna), 307
occnans('ninuiiiH^iniiitil1a),312
oc«llatii8 (SnicrintliMJi), 341
oclilca(Aiiiaiins}. Ixvi, 324
ocliroiiria (AsiiilatBs). iii
nchveopicta (Scalmiuaiida), 80
ochreopla^ta (Kntrlia), 91
octavia (Prtcisl, 332
„ r. uatalnnsia (Preoia), 332
„ f. scsaimw (Precis), 332
cKto (Amyna), 317, 321
iirtoliiieata (Statir^), 231, 234, SB6
Odynmia, 312
01 igotrophnriae, 350, 411
u,y,l,7rJM,G00glc
( cxlii
Oiigotroplms, 351, 35*2. 353, 351, 355,
357. 858, 412
olivauetw (CjplionoceplialiiB). Ill
amnhiilopii (Tholcrostota), 40
OQObiychidis (PerriBia), 37^,376, 413,
414
onouidiB (A9['1ion(lylU), 386, 414
{CoiiUriiiia), 3S3, 415
□perta (Eiicoems), 20
Ophioniniie, 321
oreaa (Acraeft), 32»
,, r. alliintaciilata (Acraea), 32tl
oribatis (Aristoteliu), 35
orichalcea (Agoatrata), 111
nrieatalU (Pi-otaetia), lit
omierodi (AmlilyB|iatha), 403, 417
ornata (Acidatia), iij
oniyCion (Del [as), xcii
ortygia (Eucoama), 22
Otbrjftdes, 171
Oxylechia, 39
oiytuoehla (Polychroais). 24
Pachymviica, 313
pacificuB (Foliates). 306
„ (SUzus), 82
pallena (Apoica), 310
pallida (Apoica), 310
„ (Uelipuna), 304
psllidilincatiis (Chartcrfciiius), 308
pslliditarse (Trypoiyloii), 317
pallipea (PatliynienBa), 312
pamphtlus (Ca«naiiyiupha), iii
paudiyanu« (Cryptosatius), 77
pandora (Clialtla), 320
[■anteli (01igotro)>Iiua), 352, 412
., (PriTisia). 376, 414
giulberiua (Moiiiklula), 31B
unurginae, 299
pu]>aTtri» (rerriKia), 876, 414
papbia (Ariatntelia}, 31
,. (Dryos), liv
Pspilio, Ixri, Ixzx, Ixxxt
Papilionidae, 335
Papilioninae, Ixxxiii
papucnais (Episyran). 64
paracusia (titatii-a), 246. 248. 266
parallels (MiitilU),31:i
{Tiijhia). 312
„ (Traumatoiniltilla). 312
ParatlelodiploKia. 400, 416
pantniicalis (UJypliodes), lOU
parhasBUB iU!thio|is (SalamiB), 332
partiiciio ((.iuophodi's). 324
„ diverstt (Guophodea), 324
Pnrnara, lixix
PaniaHaiUB, xiii
parsiiuon (CatoLliryaops), Ixi
parvula (Burolia), h9
., (ClirjsU).317
patrioift (CatochryaoiB). Ixii
|wvo (CaliRoj.xix, XX
pcctinaU (Sticloutera), 91
peculiaris (Catoctirysopa), hii
pediculaCa (Metapolyhia). 308
pellionelU (Tinea), xcix, d
iwiinaria (Hinera), Ixivi
Bjntadora (Gelechia), 51
Bpsinae, 313
PepsiB, XXXV, xxsvi. xl, 313
p*rei]iia(Atraea), 329
r. thesprio (Acraea), 329
periidiosa (Piitfiorimaea), 41
perfiduB (Eudanliia). 390, 415
perforata (Stalira), 262, 263, 268
pcrfossa (AristotelJa), 33
jiericlyTOeni (Perriaia). 376, 414
wriuaueji(Elis).63
Perii'laneta, 127, 141, 147
iwritana (Tortrix), 8
ferpiilclier{Psanimi>chBre9), 74
'erriaia. 350, 364, 366, 367, 369, 372,
373, 374, 376, 376, 378, 380, 381,
382, 3S3, 384, 385, 386, 407, 413
peraicariae (Perriaia), 377, 414
|)ersimilis (iVheatina), si. xij
pereonata (Centris), 301
peruana (Statira), 183. 198. 199, 266
peruTieiiHiB (Teletusa), xxxv
petioli (Hanoaudia). 3SI, 416
„ (Syndiploais), 391,415
petiveraua (Danaida), 323
Eetulaiii)(HapaJoaariB), 37
liaeuobreniiH, 389
Phaeiiolauthia, 387, 414
Plisloniii, I
Phaloniodae, I
Pharniaeis, 4
pharsajus (Acraeu), 3i9
„ r. pbBrsaloidps(Ai-nea), 329
Ptiegol>ia, 356. 412
fhiladelphica(PoiDpilogaster), 313
hiliilithinH«, 316
]ili]aeas(Runiicia), ii
ptioelio(Mv]itaea), ii
plitbi8ica(PolybU),307
Phtliorimai>a, 1, 40
Pliyclidobia, 354. 412
iiicta (Statira), 247, 249
Fieridoe. 164, 33-t
PieriB, li'i. tiii
pierreaua (Perrisis), 378. 414
pierrei (Rhabdopboga), 362. 413
pigea (HnacopWrjx), 335
iM-,i,A J h, Google
( cxliii )
piRger (Hartigioift), 385, 4U
piliveotriB ( Eugloasa). 303
liiloaa (Cecidomvia), 399, 416
(SUtira),'2fl3, 264, 2f
copteiyx, lir, Ixii
]>ineti (iJraclionyic), lixvi
piiii (Cecidoniyift), 399. 41B
„ -maritimae (Cecidonijia), 399,
41S
„ (PUronua), li
[liuiaria (t'idoniii), ^44, 315
jiisistrntiifi ()tho|ialovampta), IxXTJi
pleijiiata (AnaitiH), iii
]>lanta (Lvcarnopsis), 167
platans ( rtoplvctis). 321
platanoiilra (Apliia), 390, 417
„ (DrcpsiiosipliuTn), Iv, Ivi
platyzona [Arg>Toplocc), 28
plebcinna (Crocidnaema), 13, 18. 19
plflcta (Noctlia). iii
Flectroue, 108
plicatrix (I'erHsia), 377, 414
plistonicniB (Eretis), Ixviii
„ (SaranK«8a), Ixvii, Ixriii
plumata (ArialotelLa), 30
piumicomU (Stntira), ISS, 2S7
plato (PsanLmochares). 69
I>OBe (Chortorojia), 357, 412
poda^iiMis (LycusK Iviii
„ ( Memlyuiis), iTiii
H>da1iriu9{Papilio), ii
Podium, XXXV, xxxvi, xxxviii, 315
Polistes, 30fl
Polybia, 307
polyffalae (Peniaia), 377, 414
polytes (Pnpilio), xxx. xxxi. xxxii,
Ixxxiii, Ixxxvi
„ romulua (Pupilio), xxx, xxxi,
xxxiii, Ixxx, Ixxxi, Ixxxiii
„ BtichiuB (rap[li.i), xxxi, xxxii,
., theiKusiPapilio). Ixxxi
pomonae (l^later). xvji
Poiupilidie, xxxvi, sxxviii, Ix
Pompilugaater, 313
Poomyia, 367
populeti (Perriaia), 377, 414
fspoli (Smeriiithua), 341
arricoadyla, 402, 416
Porricondylariae, 402. 41<>
potentillaG (Perridia), 377, 414
praecipua (Caepbaaia). 11
pneiiatua (ElaCer), x ..
praticola (Perriaia). 377, 414
pratti (Acara). 96
,, (Paracaroides), 89
,, (Phlegctonia), 92
„ jPhjtunietra), 94
,, (Scalmicauda), K6
Precis, Ixiv
presuturalia (Statira), 17H, 22.'i. 231,
232, 233, 234, 26S
prismua (Papilio), xviii
,, (Troidea). xviii
,. r. coeliiHtia (Papiliu), xviii
„ ,, (Troid.>8), xviii
„ r. urvilleana (I'apilio), xviii
,. ., (Troides), xviii
(Eiicoema^, 19
H>idi
a. 300
proteoideH (Elaeodca), SO
frokimedia (Teracoliis), 334
rot->polybis, 308
proxiina(ABptioLdylia), 367, 414
pnideiia (Eucostiia), 23
PaaiDincx^tiaridae, xxxvi, 64, 313
PsaTuniocharinac, 313 ^
Pacuda^uia, 313 *
Pseudliormoiuyia, 39S, 41S
Paeiidovlialuutliea, xiv, 108
paaiidocwcus(Uliabdop)iaga), 362, 413
pseudolyoia (Acraea). 325
,, ustrigera (Acraea), 325
Psyllidae, 346
ptannicai: (Miaopatlia), 3r>l, 412
pteridicola (Pornsia), 3/7, 414
Pteronialua, Ivi
PterratichTiB. 208
[itUadelplia (Crocidoaema), 18
Piicciuia, 419
pudurclla pailorella (Acraea), 326
puivitii (Khabdopliaga), 362, 413
punctata (Monedula), 316
puiictatiasima (Chryaia), 817
(Pardopais), 330
pitni:tice|>8 (Statira), 210, 212, 213,
266
punclicollia (Statira), 217, 266
piitictuni (Zygnoiia), iii
pupanim (PtProuialus), ivi
purpurea (Cliloro/ada), 92
purpureipenuis (Statira). 221. 222,
227, 2B«
pnrpureipes (Crjptuoliilua), S13
puanTia (L'a1>i'r»). iii
puspa (Ct'laxtrina). 167
,. (Lycaciopsis), 167
u,y,l,7rJM,G00glc
( cxliv )
puatulana (Petrisia), 377, Hi
„ (Hannandia), 400, ilfl
PutODiella, 398, 41 S
Pyislidse, 96
pyri (Perrieia), 378, 414
pyrius (Zeuzera). iii
fyrivora (CoDtariuia), 304, 415
yro|iliorua, ijv
iiuadratns (Tamcna). 272, 283, 285
quadLicolor(PlaiieTna). 330
<|uadriinaculBU(Statint), 240, 241
tluadriplagiaU (Statira), 25^, 2A4, 266
qiiadrisiKnaU (SUtira), 239, 266
<tu«rcica[a (Amuldia), 360, 412
qnercina {Coutaritiia), 394, 415
quercus (Aruoldia), 360, 412
quiniia (Acraea), Uix, 326
radicata (Ariatocclia), 36
radicitica (Cliortotuyia), 358, 412
ranunculi (Perrisia), 378, 414
rapae (PJeris), ii, xxx, 338. 339, 340
rapliaiiistri (Daayneura), 378
(Perriaia). 378. 414
rava (Cryptoaaliua), 76, 77
,, (PwiidagKiiia), 76
reauniuriaiiu9(<)li(!atn>pliil«),353, 412
Tectaugiilata (Xaiithodesiiia). 96
recuraxciata (Polydestiia), 03
riKtilinealin (Booclnnis), 9ft
recurva(Mi-lipona), 305
reftina (StaCira), 179
rcKuUris (Tpriu), 333
reJecta(Pulybia}, 308
re1i«osa (KucoBiua), 21
Kliabdapliaga, 360, 413
rhaniiii ((io]iopt«rjx), ii
rlieUruor (Muri'lio), xlviii
,, sulMp. caoicB (Murpliu), xlii
,, snU^. casebe8(Morpho), xlix
rhududendi'uiii (Perrisia), 376. 414
KhogaJi
Bliopalocera 322
Kbnpalomyia, 351, 35.J. 412
Kha|ia1uMniia, 313
Kho[M>luioniidiie, 313
rifCilica (I'kruta), lOS
rilarniac (iWut)i>cliBl(-<>t1 it'll). 107, 10(!
n>liUBtii(Mw'iiMriiiyru), 88
rudwayi (Maiib'zuiuia). 310
rorarU (Cmciiluteiua). 16
«m (M «si|,huni). Iv, IvL
■« a(ltU1>l«|>]iaK>). 363. 4)3
isa II (K1abrlo|>lia)Cu), 363, 413
I nwmaniB (Dicrogeiiiniii), 54
I rostratuB ( [^cus), liiii, tix
„ ( burolycna), Itui, liz
' rostnipiana (Perrima), 370, 414
\ rotandioolliB (StaUra), 209, 266
rubi (Callophrya), xvii
„ (Laaioptera), 350, 411
rubicola (CoDtarinia). 394, 4I.''>
nibidata (Perigea), 88
rubra (MaaBslongia), 401, 416
,, (Tijiiila), 387, 414
nibricata (Larra), 816
(Tbvgator), 301
rubrithorax (Statim). 202
rubaaameni (Porriaia), 379. 414
tuderalis (Ctontarinia), 394, 415
rufa (Pormica], xiiv
„ (SUtira), 107, 2)7
ruficollis (AathJcua), 257
,, (SUtira), 257
nitirroua (SUUra), 196, 199, 217, 221,
266
rutipes (rsaniinocbares), 64
nifivciitriB (Helipoiia), 306
I „ rnr. llavulineaU (Mdi-
I i>Dua), 306
nilolnteiia (Kasliibtliynuiis), 60
nigoBisiiiiiiun (EiitomofcnatliUH), 83
rugoauiu (Caltnonia), Ixxxix
rHiiiir.ii« (AtylodiploaiH), 402, 416
luao (Oxytialims). xli
niatica { Epii^haria), 301
sabiiloaa (Amnioliliila), liii
sa.Java (Xqitix), 331
,, ]iiarpi!ssa (Neptis), 331
anka (Caliiia^), xili
aalicanaed'evrUia), 379. 414
salidiis (l>a»ynpiira), 363
„ (KbHl>dapbaga),362, 363, 413
aalioiperda ( Hbabdopbofia). 361. 363.
413
wlicix (nallaniiii), 3S3
„ (Rbslxlophaffi). 36S. 413
aalicomiae (Batdratia), 350. 411
U)Ci>a(RliabdupbaKa). 361, 413
• (1
w |wtd (LI II
in), 378,
Saliii
xl
U9). 400, 4)6
(Ag«itraU), HI
i-ma« (SUUra), 183,266
Han^iiiiicollin (Podilus). 257
saiiKuiaorlw (Perrisia). 379, 414
«.l.|.hiria (Persia). 313
Mai^itbaniiii (Asphundylia), 3S7. 414
(Trottrria), 349, 41)
Hatiiniina(AHHU>tclia), 33
SaltTidar, 324
Hcafiiusap (ConUriiiia), 394, 415
„ (Perrisia), 379. 414
noabrirollia (Statira), 228. 229, 266
ll,y,l,/eJh,G00glc
{ oxiv )
iicapularis (Lf g»b). lii
„ (Merolycus), lix
„ (StAtini), 22G
scariHcata (Pbalonia), 3
Scaups?. 389
ScvliphroD, SI4
scltistacca subsp. rugiilosa (Polyrhu-
chu), Ixivi
Schizomjia. H85, 414
schlechtoiidali (ClinodipluHix), 401,
(PerilBifl), 379, 414
w-limMti (PerrUia), 379, 414
Suiara, 404
scintillanH (SUtira), 203, 266
Sculiadoc, 312
Sculiidae, 60
Sculujicndra, 1 Ifi
Bcoparii (Cnntarinis), 3t>4, 41G
9«,mii.uMSUphytu«l. '"fxviii
HcropliliUriae (SCictudiT'loiua), 398,
415
Aciibrllariii (Nfcturiiiu), 300
(Statire), 2r)3
BegT¥KBU (StaUral, 244, 266
iw^onica (P«iisis). 313
setene (Rrantliis), ii
gelbtiu (Tapinntus), I
Buloiui (Auatonn), 102, 104
svnmlit (SatyruflJ, iii
wmicnprea (Sbitira), IBS, 20S, ses
wmitiictunsis (>Wiinii>cliai¥8), 71
actnituctuoaus (Poni]>iliiH), 70
„ (PaaniiTionlial-es), 70, 71
wnii nigra (Htatira), '24!), 266
MiuiviolacfA (Sutira), 2S8, 266
Sfmndobia, 3E>4, 412
sciifKatp.nais oritntia (Kmyplieiii;), 331
MDt\ (ADoplius;, 68
,, (Psamniocliares), 68
(wrena buxtoni (Acnu'a), Ixix
seria (Bacttra), 24
aencfa (Polybia). 308
„ (.Slatira), 2)3, 220, 221, 266
sevntina (Perrisis), 380, 414
strra (Tireaiaa), I
sertifer (Pteroniw), 1
iwtigertt {E|>hutu[iiur)i]iit), !>7
aeloBA (Cnepliatiia), 10
s«v«riDa (Belenoii), Ixiv, Ixv, SSIi
sil>iri<s(Anipillcx),SI
sichelianus (^thii!i), 312
sifiillatus (AzaiLus), 333
aianata (MonediiU), 316
silvicola (Perrisia). 380, 414
siniana (Phiacupttiryx), lii, liii, liv
BimiliB (Perrifliu), 380, 414
aimiltima (Coeliuxys), 303
Binionis (Statira), 21 1, 212, 266
simpler (Cereeria). xixvii
Bimplicipea (SULira), 194, 266
BiDuacocollis (Heterorrliiua], 110
siaymbrii (Perrisia), 380, 414
amaragdina (ExaereU), 303
„ (Hct«rorr1iina), 110
iinianigdulns (CyplioDotcpbatiis), 100,
111
Smicre. 320
sociubilia (Scbizoniyia), 402, 416
uidaliB (He){alopU), 299
„ (Perriaia), 380, 362,414
sulDisi (Phjctidobia), 354, 412
Bonchi (Cystiphora), 358, 412
aophoroe < BroasoliB). 320
aurbi (Contarinia), 394, 41G
aotikvnaia (Acrava), 328
, , I', rowena ( Acraea), 828
Sphecinae, 314
Spbecodinae, 298
Sphegidao, xixvi, tx
splieDobatlii'a ( I'eroiiea). 13
aphenodera (SUtita), 200, 267
spiienoptera (SUtira), 203, 267
Spbex, xxxvif ■" -I" ''i-
i,315
xlii, iliv, X-
Spliiii
SpLragidupliorua, 2hA
Spiloiiiicrodua, 320
spiniffera (Aiiomoxpnaj, 20
(SUtira), 176, 177, 267
splendena (Euploea), xii
splendicana (SUtira). 209, 267
»i{ua1ida(Ej>icou>etiH), 103
stavtiydiB (Perrisia), 381, 414
sutira, 169, 17o'. 193. 242, 246, 264,
257, 266
Statiro|K<iB,263
St«raiii.:llB, 350,411
sUiubi-li (Sutira), 196
atfliiii (Contarinia), 391, 415
Htellariae (MacrolaHa), S60, 412
atnllatariini ( MacroglDana), iii
steiiocupbala (IStutiia). 200, 267
ateucxlsia (SUtira), 233, 234, 267
Htenoptcra (SUtira), 239, 267
aUnialia (Clinteria), 108
Stictodiploaia, 396, 415
Slizinai.', 82
stiiaU (Coaoinia), iii
strintirrons (/napilotbyimiia). 58
►triatum (Anticniridiimi), 397, 416
atnatiia (Sphrx), iixvii
strobi(reiTi>iia), 331, 414
atrobilobiuH (Adrlges), 389, 420
ih,Cooi^lc
atrongjlioides (Statira), 284, 267
Stron^yiiuni. 2tt4
fltygia [ Uracil jtieutTt), 388, 4U
Bubacida (SpargsTiothU), 13
siiliai^nea (SUtira), iVj, 221
HubBmicBlis (Glyiihodes). 100
Bubcincta (G]jcy[ihiin«), 111
aubfrnesttnta (Stttira), 247, 267
aiibglabral* (Stitir.), 215, '^67
siibindiuta (Corvstlia), 94, 95
8ul>liniiB(Pli*loma), 2
suliinBrgiaata (TBeniaphora), 97
SuUris, xxxvi
snliterrsnea (Tiipli[>diplo8ia), 39S, 416
HiicciJitta (Lycopcrdina), xlvi
aiilfiiaa (SUlira), 211, 2:'6, 227, 267
suriiptuosa (Ststiva), 202. 2a'), 204,
2U5, 206, 207, 267
miniptuoaua (Sphex), xxivii
aiiperna (Rliabdoplinga), 363, 413
Burinama (Svnocca), 309
BiirinanienBia (Euglossa), 302
„ (Houeduln), xxxviii
„ (Stietia), iiiviii
BUtwali. [Statira). 170, 176, 177, 231,
234, 267
Bjbsris (Lycnenn). 284
„ (Tanicua), 272, 274, 283, 28J,
285, 294
syoophanta (Polyhia), 308
sylvan us (Augiodca), iii
Syiidiploata, 390, 415
Byn^^neaiae (Misopatlia), 3ri2, 412
Syii«™, 309
Syrpliidae, icvii
libanidftc, 31«
Tal>aDii8,xli
Taehyris, xk'u
Tachytca, »iiTii, 3Hi
Taenitria, xiix
Taeniaiiliora, 97
ta^L's (Nisoniad«s), iii
TaJHria, 167
tamilana (Paiulio], lixxii, Ixixili
Uuaccticola {Klioralomyia), 351, 412
TaphodiploBiB, 398, 416
taraiaci (CVatiphora), 358, 412
Tai lions, xviii, 269, 270. 273
taxi (Taxomyia). 385,414
Taxomvia, 385, 4U
tckarcfiiis (Papilio), xil
Telptusa, ixxv
Tellervo. xxix
TemnoBoniB, 298
tenebnisa(MacroarinyrB), 88
tfiiellH (Winuortiia), 402
tenuiliniialiB (Bocclioria), 99
Unuiiwa (Statira), 254. 267
tenuis (SlAtire), 233, 267
TeiacoiuB, Iii, Ixii, Ixir, Ixt
Teratoneura, Ixi
teres *lBiva[PrecU). 333
tersliratm (K)ialidophaga), 363, 413
tcradoD (Papilio), Ixxx
tenuinalia (Perriaia), 363
(Kliabdopliiigii), 363, 413
(Statira), 214
terpaichnrc (Acraea). 328
r. rougeti (Acropa). Iiii,
328
terroaa (EmnimimetiBil, 103
,. (Protaetia), 103
t«as(!lUta (Polvdeania), 93
tcsBellutum (XeHtobiuni), xvii, xviii,
testacea (SUtira), 212
t«tensi (Pen-iaia), 381, 414
t«trahit (Penisia), 381, 414
tetraxoa (AnoniQxena), 30
Tettigoniellinae, xxxv
teucer
„ teuc«r (L'aligo). xix, ixi
texanum (Eiphosoma). 321
thala&sina (I^iiceronia). 334
tbatia (Augocliloro), 299
t1ialictricoIa(Ametrodiplosis].402,416
(Clinodiploais). 402
Thauniatomutilla, 312
ThfiMidi ptosis, 306, 415
theoplimatua (Hesperia). 286
(Tanicus), a69, 270. 271 ,
272. 273, 274.27 6, 278,
277,382,286,287,288,
289, 290, 295
thrapia (Papilio), 2B6
(Tamcus), 296
thetia (Agriadm), xvi, 165, 166,107.
168
IhiodeH(Recurvaria), 38
Tha1eroBt«la, 40
tlioinae(HarpactqpuB). 315
„ (Sphex), 315
u,y,i,Ajh,Cooglc
thoin&&isiis (FerrUia), 3S1, 414
Thurenia, 390
Tlijgater, 301
thymi (Aspliondjlia), .187, 414
„ (Jaa«tHta), 35S, 412
thymicola (Jane tie] la). 3:,'^, 412
Thyttnidav, 58
ThjDDUs, 59
thyridoides (Corgatha), 96
tibialis (Statira). 193, 194, 267
liliae(Perrieia), 382, 414
tiliamvoIvenalPerriaia). 382, 414
tiliarain (CoDtariaia), 3e!>, 415
tLiicti|)etinJG(Pqisis), 313
Tin™, liii, Kciv, lev, xcvii, xevi
Tineidae. Kxii, xcviii
Tineina. icviit
TinKida«, 349
Tinvididae, viii
Tiphia, 312 ■
Tiphiiuw, 312
Tipala, 3(13. 399
tithonuB (Epineplielf). iii
tolen9i8(Statira), 229
tomentosa (Eurybraeliya), ix
tornatella (Chegobin), 356, 412
torquata (Glycypliaiia), 111
tortipes (Statira), 171, 101, 287
Tortritidae, 5
Tortrix, e. 7
tartrii(PerriBia), 380. 382, 414
trab«atus(LynuH), Iviii. liK
traclivlii (Pf-rriflia), »83. 414
trachydora (Statira), 227, 267
tragopoKonin (Oontariiiia), 39^1
trail! (C-intarinia). 39.^, 415
,, (Porriaia), 382, 414
tranBferrana{Kui'oama), 22
translucida (SUtira), 211, 214, 2s;
tremutae (Coiitarinia), 395, 415
,, (Hannanilia), 400, 416
trianKulata (Trairlioa),
415
ti'ichinaspia (Plitli
Trichodiploais, 397,
Trichograninia, 320
TrichoKraninmtLnae, 320
TrichopUra, 124, 127, 140
triden» {Stizua). 83
trifolii (ClinofliploKis), 390,
,, ( Hadrobreniia), 390
.), 41
{Pt"
ia), 3!
, 414
rKtci (Codtariuia). 395, 415
riria (Melitaca), ii, iii
Trotteria, 34B, 411
Tr>-poxylinfte, 317
Trvpoxylon, 311, 317
tiiU-culaU (SUtira), 179, 182, 184,
185, 267
tubercnii (Jsnctiulla). 355,412
„ (Perriaia), 355
tuberifica (Trialioniiomyia), 388, 415
tubicola (Perriaia), 383, 414
tympanirex (01if(otrophuB). 353, 412
ty|)hoeu* [Gfot rapes), xo, xci
tytia (CadtipL), ix
„ (Danais), ii, x, xiii
uliois (Aaphoadytia), 387, 414
„ (Perriaia). 383. 414
i]lraHria(Pi>irisia).333
ulniariae( Perriaia), 383, 414
nlmi (DliRotrciphus), 351, 412
iilmicola (Pcrnaia), 333, 414
' " ' I) (Kielferia). 3S5. 414
(Stictodiplosis), 396,
umbelliferarum (TroUeria), 349, 411
unibrosa (Statira), 222. 267
UDirasoiella (lieleciiia), 50
■■■ iata{KpIi ■ ■ ■
la (Phtliu
Lii'ae (Macrosiphuii
„ (Perriaia), 38^1, ii«
„ (Vanessa), Ivi
uaamlHirB (Hypolimnas), Ixvi,
listulatH (Klis), 60
iiiii (Perriaia). 3H4, 414
n(Pen
ia).;
I, 414
Trigonal idae, 31!
Trinoualys, 317
. trilineata (Statira), 232, 233. 267
triBellatJt (Statira), 219, 245, 246, 267
Trisliormomyia, 388. 415
triBtig(Plectrouc), 108
vajji-KJitlata (Statira), 170, 175, 247
248,249,250, 261,267
vaKL-notaU (Slatira), 248, 249, 247
Valerianae (CoDtsrinla), 395, 415
valerii (Arceutliumyia), 352, 412
' valida(Cliiiteria), 111
I vaiidimrnia (Statira), 172, 225
! vallator ( Nematuii), lix
I varia (Mdipoua), 306
variabilis (Statira), 214,221
variatm (Statira). 249
varicgata (RvtiibidiiU), xxxviii, 316
I .. (I)ielia), 313
(Hyptipcrijtea), S7
variicolur ( [Iy|Huilatii'a), 201
vaasei (Paammoc bares), 71, 72
veitchi(Crabro), 84
velutiiia(rulybia), 308
venosns {Tanitiis), 272, 274, 276, 276
veHtrioola (Mayftiola), 357, 412
•nwie (Statira). 25.-i
(C«ciduiiivia), 3911, 416
!
ll,y,l,7rJM,G00glc
( cxlviii )
86. 267
<a(SUtin), 181,
versicolor (Centris), 301
„ (OxjcetoniB), 103
(PoliaUt). 306
„ (Sutira), 220, S21, 267
Vespa,130
Ves|>ida«, 306
Vespinae, 306
Tibiuria (PallouiB), iii
vibiinti (c3<iD tan Dili), 395, *15
viburnoniiD (Contarinia), 395, 415
Ticana (Aiistotulia), 36
viciae (ADabremik), 390, 41B
„ (Pemsia), 364. 414
vidua (PiDaco))teryx),-33l
viiriiitipuilctaU (Statin), 185, 207
267
villica (Arctia), iii
viiuiualiB {IUiabdo|>hagB), 363 '
vintentana (Eucoama), 22
Tiniila (Dicranura), Ixxj
vialacea (Oamili*), ixivii
violie (PerriBia), 384. 414
„ (THicKinia). xii
virens (Paeudochaluotliea), lOS
vir^.aureac (Perneia). 384, 414
vindana (Tortrix), o
Tiridiciiicta (Statira). 216, 241, 242,
267
Tiridifa«ciata (3Utira). 242. 243. 244,
viridiuotxU (SUtira), 241, 243
viridiiiennis (Statira), 170, 171, 187,
188. 189, 190. 192, 1)3,
214. 267
(SutjTa).2U
viriditiacta (SUtira). 198, 169. 367
viridiriltata (Statira), 224, 235, 267
viruUnU (Ephutomorpiia), 64, 55
vitiensis (CjphoooDjx), 78
vitia-ida«ae (Perriaia), 384, 414
TitUta (Statira). 280. 256, !B7
vitliKera(CIeroU), lti6
„ (Ephutomorpha). 66
t'olveiis (MacrodililMia). 399, 416
„ (Hocrolabis). 403, 421
wakefleldi (Kuxantbc). 330
nat«rstradti (Taniriis). 276
n-illiamai (l't«roml>nis), 63
xanthodpra (Sutira). 250. 267
xantholelm (Teracoliis). Iv
xantbotriclia (Rrcurvaria), 3S
Xenodijdoais. 399, 416
Xestobiiim, xxiii
Xybwopa, 299
XyloFopinau, 299
ZospilothjliDua, 69
Zeliu. xixv
?j!te» (Acraea), 325
r. a
a(Aci
ea). 325
Zethus, 312
Zcugopt«ra, 112
Zeiilidililosia, 397, 416
zoolina (Charaxea), 330
ZyifoUa, 356, 412
7Kotoma (Slegaata), 48
iM-,i,A J h, Google
?A
December lo, 1918.
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Part V, 5«.. to Fellows, 3i. 8i.
1914.— Part I, £1 1»., lo Fellowa, 16*. Brf. : Part II, £1 4*., to Fellowa, IS".;
Parts III. IV, £1 2a., to Fellows, 1B». M. ; Psit V, lOj., to Fellows, 7».fl<t
1915.— Part I, 17a. 6(i., to FeUows, 13j. fld.; Part II, lis., tc Fallowf,
8s. Sd.; Parts III. IV, £2 14i., to Fellows, £2 0». M.; Pwt V, it.,
to Fellowa, 3<. 9d.
1B16.— Parti. £1 18»., to Fellows, £\ 8s. S(I,;PartlI, 10s,, to Fellows, 7s. M.:
Parts 111, ]V, £1 2». 6d., to Fellows, 16*. 9d. ; Part V, 6i. Orf., to
Fallows, 4j. 6d.
iei7.~Part 1. 17s. Si, to Fellows, I3l. 9d ; Parti II, III, IV, 17a. Orf., to
Fellows, 12s. Bd.; Part V, 6s. Od., to Fellowa, 4s. ed.
The following may be obtained separately:—
Poseoe's ' Linxgieomia 3falajiana,' formiui; toI. iii, of the Third Series, pablished
price, £2 12«. ; to nou-Fellows. £1 IOj. ; lo Fellows, £1.
BalyV 'Pkjitophaga Maiayana,' forming ji«tt of vol. it, of the Third Series,
published price, ISt. ; lo uon-Fellows, 10s, ; to Fellows, 7s. 6i<.
The 1893 Catalooue of thb Library, with Snppltment to 1900, is pnblUhed
-■. 10s. ; to Fellows 1>. The Supplemaot only, U.id.;\a Fellows, 3s.
THE ENTOMOLOGICAL SOCIETY OF LONDON.
THE FELLOWSHIP AND FEES.
F«Uowg pay tn Adminlon Fm of £2 2i. The Anaual Contributian U £1 It.,
dufl on tlie Brat Jay of Janiwry in each ye*r, and pnjabla in adTance ; or a
Composition Fee of £16 15i. may be paid iu lien thereof, tlie whole payment (or Life
FelloTibip, incladiiii! the Admiuiaii Fee, being £17 17f. Fellowi reeiding per-
manently outside the United Kingdom pay no Admiuion Fee.
All Feee should be paid to the Treasurer, Mr. A. H. Jones, Shmhlanda, Elthnm,
Kent, and not to t)ie Secretaries.
Fellnwe degiiing to pay their Annual Coutribnlion thrangli their bankeia txn
obtain an official farm of iMnker'a order by applying to either the Treasurer or to the
Besiilent TJbrnriKn.
Fellowa vhose ContribatioDB for the carrent year have been paid are entitled
to receive the pahlications of the Society free of charf[e. Further copiea inay l>e
pnrchawd at reduced pricea by applying to the Resident Librarian.
Forms of application for Fellowship and copies of the Bye-lftrs and Litt of
Fellows may be obtained from either of the Secretaries or from the Resident
Lahrarian. >
MEETINGS AND EXHIBITIONS.
Intending exhibitors are required to agnify their names and the nalnre of their
exhibits to the Chairman twfore the beginning of the meeting, in order that lliey
Itiny be calieil upou from tlia cliair. Descriptive notes of all exhibits slionld be
hnnded to tho Secretaries at the snrae meeting for printing in tlie Proceedings. II
the epidisscope is required a week's notice must be given ; exhibits to be satis-
factorily focussed by this inalrumeut must not eicped 7 ins. square.
FelloWM residuntabroBd, or wlio are otherwise unable to attend, are romimlcd that
any specimens, notes, or obscrvatious they may send to lite SEClotaries will be
considered by the Council, with a view to eiliihition or renJing at the meetings
of the Society.
PAPERS AND ILLUSTRATIONS.
Fellows dexiring to communicate ]uiiiers to tho Society must send the full titles of
socli papers either to the Secretaries at the Society's rooms, or to Commander
J. J. Walker, M.A., R.N., Aorsngi, Lonsdale- road, Summcrtown, Oxford, al,
least foui'teen days prior to tbe date of the meeting at which it is proposed that
such papers sball be read.
Anthors proiiosing to illustrate Ibeir papers should communicate with the
Seeretsriea before the drawings are executed. The Council recommend that tbe siie
of the work ou plates should 1» limitad to 81 ina. by 4 ins., and in no case will it be
allowed to exceed 6} ins. by 4} ins.
Attention is called to tlie Instructions to Authors issued with Part I of each
Tolnme, which may also be obtained of the Resident Librarian. Inattcutioa to
these teguUtione may involve on aathoi in considerable expense. Oi.)(.)QIl'
CONTENTS OF PARTS II, III, IV.
IX, A Bevision o[ tbe gmmt Tarve^. Bt O. T. BiTBinil-BlKEB, FX S..
F.Z.S. ■
X. Notes on some Britiih OniaiiB 2fvn<iu>p<(ra (eidmriTe of the F»nitiei<ldt),
By O. B. Bodkin, B.A, Dip. Agrie. (Csatab.), F.Z.B., F.E.a., Ooretn-
meet EooDomic Biologiat, DepBrtment of Sdenoe uid AgncuItiiM.
British Ouiank
XI, On a CollectioD of Butterfliei taken in Eat Africa by Hr. W. A. t«iDbani.
BjH. ELTRiNaHui,H.A.,D.8c. With Dot« on the /'ttn'iuK, by Dr.
F. A. Dixit, F.R.3., and deacriptiou of a niw form of P. dcrJmiu 9 ,
hv Prof. E, B. FouLTOH. F.R.S. .u— — ^
Vsbnlary
Tlu Chair tvill be taken at Bight o'clock.
THE LIBRARY .
18 open to Fellows and their friends every day irom 9 a.m. to
6 p.iii., except Saturdays, when it closes at 2 p.ni. On tlie nights
of meeting it remains open until 10 p.m.
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