NATURE
CONSERVANCY
COUNCIL
World Checklist of ‘Threatened.
Amphibians and Reptles
Nature Conservancy Council Great Britain
Digitized by the Internet Archive
in 2010 with funding from
UNEP-WCMC, Cambridge
http://www.archive.org/details/worldchecklistso83groo
World Checkhist of ‘Threatened
Amphibians and Reptiles
Bnan Groombridge
Wildlife Advisory Branch
Nature Conservancy Council 19/20 Belgrave Square London SW1X 8PY United Kingdom
om ’
=
Contents
Introductory Notes
Taxonomic List
AMPHIBIA
Cryptobranchidae
Ambystomatidae
Salamandndae
Plethodontidae
Proteidae
Leiopelmatidae
Pipidae
Discoglossidae
Pelobatidae
Leptodactylidae
Myobatrachidae
Bufonidae
Hylidae
Sooglossidae
Ranidae
Hyperolidae
REPTILIA
Kinosternidae
Dermatemydidae
Emydidae
Testudinidae
Cheloniudae
Dermochelydidae
Carettochelyidae
Tnonychidae
Andrias
Ambystoma
Chioglossa
Batrachoseps
Eurycea
Hydromantes
Phaeognathus
Plethodon
Typhlomolge
Typhlotriton
Proteus
Leiopelma
Xenopus
Discoglossus
Pelobates
Batrachophrynus
Eleutherodactylus
Philoria
Rheobatrachus
Atelopus
Bufo
Nectophrynoides
Hyla
Pseudacris
Nesomantis
Sooglossus
Conraua
Rana
Megalixalus
Kinosternon
Sternotherus
Dermatemys
Batagur
Callagur
Clemmys
Geoclemys
Heosemys
Kachuga
Melanochelys
Morenia
Pseudemys
Rhinoclemmys
Dermochelys
Carettochelys
Lissemys
Trionyx
Page
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Pelomedusidae
Chelidae
Alhgatondae
Crocodylidae
Gavialidae
Sphenodontidae
Gekkonidae
Pygopodidae
Xantusudae
Agamidae
Chamaeleonidae
Iguanidae
Lacertidae
Cordylidae
Tedae
Scincidae
Anguidae
Aniellidae
Helodermatidae
Varanidae
Typhlopidae
Boidae
Erymnochelys
Pelomedusa
Peltocephalus
Pelusios
Podocnemis
Phrynops
Platemys
Pseudemydura
Gavialis
Sphenodon
Cyrtodactylus
Oedura
Phelsuma
Paradelma
Xantusia
Hydrosaurus
Uromastyx
Chamaeleo
Amblyrhynchus
Anolis
Brachylophus
Conolophus
Cyclura
Cordylus
Pseudocordylus
Ameiva
Cnemidophorus
Crocodilurus
Dracaena
Tupinambis
Gongylomorphus
Leiolopisma
Macroscincus
Gerrhonotus
Aniella
Heloderma
Varanus
Typhlops
Acrantophis
Aspidites
Boa
Corallus
Boidae cont.
Colubridae
Elapidae
Viperidae
Loxocemus
Python
Sanzinia
Trachyboa
Tropidophis
Ungaliophis
Xenoboa
Clelia
Drymarchon
Elachistodon
Hydrodynastes
Masticophis
Nerodia
Thamnophis
Naja
Ogmodon
Bothrops
Crotalus
Vipera
Additions and amendments
References
Index (to genera, synonyms, common,
and trade names)
Acknowledgements
This checklist was compiled and revised in
October 1982 for the 3rd edition by Dr B
Groombridge (now at the IUCN Species
Conservation Monitoring Unit, Cambndge) at the
Amphibian and Reptile Section of the British
Museum (Natural History), under contract
NCC/E2/36/04 from the Nature Conservancy
Council of the United Kingdom.
The compiler is most grateful to the staff of the
Amphibian and Reptile Section at the British
Museum (Natural History), in particular to Miss A G
C Grandison and Dr E N Arnold, for their valuable
assistance.
lst edition published 1979 as a typed document
with limited circulation
ISBN 0 86139 075 X
and edition published 1981
Designed by Creative Department, Manchester
Ltd.
Printed for HMSO by Robendene Ltd.
ISBN 0 86139 095 4
3rd edition published 1983
Revised design by Margaret Morgan BA
ISBN 0 86139 224 8
© Nature Conservancy Council 1979, 1981, 1983
Preface
This checklist was originally prepared to aid the
United Kingdom's Scientific and Management
Authorities for the Convention on International
Trade in Endangered Species of Wild Fauna and
Flora (CITES) which the United Kingdom has
implemented since the 1 January 1976.
The aim was to provide a summary of basic data
on each taxon relating to the countries where it
occurs in the wild, its category in the IUCN Red
Data Book, its listing in CITES and important
sources of reference. Particular regard has been
given to listing general names, synonyms, English
vernacular names and trade names. This
information will help to clarify conditions on the
issue of permits to trade in amphibians and reptiles,
especially in those cases where ambiguous or
erroneous supporting data are supplied. It is hoped
that this publication will be of value to all
Management and Scientific Authontties of the Party
States to CITES and to all those concerned with the
wider studies of herpetology and related matters.
This checklist covers only those amphibians and
reptiles which are themselves threatened in some
degree, or treated as ‘look-alikes’ under CITES, at
the time of compilation. It is clearly desirable for the
work to be extended to include the remaining taxa
and therefore complete a full coverage of the
group
The present document was completed in
January 1979 and since then has been subject to
partial revision to take account of decisions
affecting the Appendices at the 2nd meeting of the
Parties to CITES at San Jose, Costa Rica in February
1979, the 3rd meeting at New Delhi in 1981 and new
taxa included in Part 1 of the IUCN Amphibia-
Reptilia Red Data Book (1982).
The United Kingdom Scientific Authority and the
Nature Conservancy Council are grateful to Dr
Brian Groombridge for undertaking this work on
their behalf.
AJBRUDGE
Wildlife Advisory Branch
Nature Conservancy Council
Belgrave Square
London SW1X 8PY
Winter 1983
Introductory notes
The purpose of this work is to provide a
combined list of the amphibian and reptile species
(or subspecies) appearing on Appendices I, II, and
III, of the Convention on International Trade in
Endangered Species (CITES), and those included
in the Red Data Book (RDB). The latter includes data
sheets from the second edition issued in 1975,
together with many new or revised sheets dated
1978 or 1979 (but not generally available until 1980)
A useful introduction to CITES is provided by
Inskipp & Wells (1979, see Reference list C).
This revised edition incorporates changes made
in the CITES Appendices at the New Delhi meeting
of the Parties in 1981, and changes and additions
among Testudines, Crocodylia and
Rhynchocephalia made in Part | of the revised
Amphibia-Reptilia Red Data Book (ref. 152). New
and fully revised RDB accounts for remaining
reptiles and for amphibians are currently in
preparation, the taxa among these groups noted in
this checklist are thus not fully representative.
For each taxon, the scientific name 1s given first,
with the currently most frequently used synonyms
in brackets, Common names (English language),
and trade names (in brackets), appear on the
following line. Very few of the listed taxa have
regularly or universally used common names. In
several cases, identical trade names appear to be
used for different taxa.
The Red Data Book taxa included are those
listed in Preamble 8 (i.e., those for which full data
sheets are provided) as well as those in Preamble 9
(that are suspected of being threatened, but for
which full data are currently lacking).
Geographical Range
The geographical range of each taxon is given in
terms of political units, listed in alphabetical order.
The exceptions are some of those taxa occuring on
small and/or remote islands with well-known
names, that are administered by another country;
for most such taxa the administenng country is
given in brackets, e.g. Galapagos Islands
(Ecuador). In certain cases, when a taxon is found in
a restricted area within a larger political unit, the
name of that area is given in brackets, eg.,
Indonesia (West Inan).
The political names given are those in common
usage, and may notin all cases be the
diplomatically correct version.
When the name ofa country is given as within
the range of a certain species, that species may not
occur throughout the country, and may even be of
only penpheral occurence.
Without surveying the range of each taxon in the
field, itis necessary to rely on published records.
Most taxonomic works give the range of a taxon in
terms of broad geographical areas, rather than of
precise political units. While some countries have a
relatively well-known and well-reported
herpetofauna, this is not tue for others. These
factors may occasionally have resulted in some of
the geographical ranges given here being
incomplete or innaccurate, although every effort
has been made to counter this.
8
Taxonomic sources.
The most recent available checklist or revision
has usually been followed for the taxonomic
treatment of each species or group of species
herein, Certain groups are clearly overdue for
species-level revision. In the interests of stability,
the present list follows the taxonomic arrangement
of the RDB for the forms appearing therein, unless
there is good reason to do otherwise. The genera
and species are in alphabetical order within each
family. The scientific and common names of North
American species have been modified to accord
with Standard Common and Current Scientific
Names for North American Amphibians and
Reptiles (Collins, Huheey, Knight, & Smith, 1978)
The following list states the primary taxonomic
source followed for most of the groups dealt with
herein, and refers to an authority for each
modification from that primary source. All works
cited in this section appear in part (B) of the
Reference list.
Caudata and Anura: follows Gorham (1974), except:
(a) Ambystoma tigrinum californiense here
treated as a full species, A. californiense, after
Collins et al (1978)
(b) Batrachoseps aridus is added, described by
Brame (1970).
(c) Family Atelopodidae no longer recognised,
Atelopus here assigned to family Bufonidae, after
McDairmid (1969).
(d) Bufo nelsoni here reduced to a subspecies,
B. boreas nelsoni, after Collins et al (1978).
(e) Five species of Nectophrynoides are added,
N.cryptus and N.minutus, described by Perret
(1971, 1972); also N.osgoodi (transferred from Bufo)
and N.malcomi described by Grandison (1978) and
N.liberiensis described by Xavier (1978).
(f) Sooglossidae here elevated to family rank
(esomantis and Sooglossus listed), after Lynch
(1973) and Duellman (1975).
(g) Rana goliath here listed as Conraua goliath,
after Lamotte & Perret (1968).
(h) Rana pipiens fisheri now in synonymy with
Rana onca, after Pace (1974).
() Hyperoliidae elevated to family rank
(Megalixalus listed), after Liem (1970).
G) Myobatrachidae elevated to family rank
(hiloria and Rheobatrachus listed) after Lynch (in
Vial) 1973, Heyer & Liem (1976), and others.
Testudines, Crocodylia, Rhynchocephalia: follows
Wermuth & Mertens (1977), except
(a) Geoemyda tricarinata here listed as
Melanochelys tricarinata, after McDowell (1964).
(b) Pseudemys terrapen malonei is treated asa
full species, P.malonei, as in Schwartz & Thomas
(1975) and Pnitchard (1979). The species is assigned
to the genus Pseudemys rather than Chrysemys,
following Wermuth & Mertens (1977) and Pritchard
(1979); there is considerable debate about the use
of these names.
(c) Testudo is broken up into four additional
genera (Acinixys, Chersina, Geochelone,
Psammobates) and Testudo is restricted to five
Palearctic species, following Loveridge & Williams
(1957) and the majority of subsequent authors
(d) Podocnemis dumeriliana is listed as
Peltocephalus dumeniliana (apparently a synonym
of Peltocephalus tracaxa, unpublished work noted
in Pntchard (1979) and Rhodin et al) after Rhodin et
al (1978)
(e) Podocnemis madagascariensis Is treated as
Erymnochelys madagascariensis, after Williams
(1954), Smith & James (1958), and Rhodin etal (1978)
The above modifications within Testudines are
also all incorporated in Pritchard (1979). (See Bour
(1978) and Obst (1980) for different views on the
nomenclature of certain of the endemic Malagasy
tortoises)
(f) Crocodylus mindorensis here treated asa
subspecies, C. novaeguinea mindorensis, after
Wermuth & Fuchs (1978) and several earlier
authors
Squamata; Gekkonidae and Pygopodidae: follows
Wermuth (1965), except;
(a) Gymnodactylus serpensinsula here treated
as Cyrtodactylus serpensinsula, after Underwood
(1954) and subsequent authors
(b) Phelsuma agalegae added, described by
Cheke (1975)
(c) Phelsuma dubia comorensis elevated to a full
species. P. comorensis, after Mertens (1966a),
(d) Phelsuma newtoni changed to P.
edwardnewtoni, following Vinson & Vinson (1969)
(e) Phelsuma modesta added, described by
Mertens (1970).
(f) Rhoptropella ocellata is treated as a species
of Phelsuma, P. ocellata, after Russel (1977).
(g) Phelsuma vinsoni changed to P. ornata, see
Mertens (1970)
Squamata; Agamidae: follows Wermuth (1967)
Squamata; Chamaeleontidae: follows Mertens
(1966b), except;
(a) Chamaeleo angeli added, described by
Brygoo & Domergue (1968).
(b) Chamaeleo antimena here treated as a full
species, after Brygoo & Domergue (1968).
(c) Chamaeleo balteatus here treated as a full
species, after Brygoo & Domergue (1968).
(d) Chamaeleo belalandaensis added,
described by Brygoo & Domergue (1970).
(e) Chamaeleo capuroni added, described by
Brygoo, Blanc, & Domergue (1972).
(f) Chamaeleo eisentrauti added, described by
Mertens (1968).
(g) Chamaeleo guentheni here synomised with
C.pardalis, after Brygoo (1969).
(h) Chamaeleo labordi treated as a full species,
after Brygoo & Domergue (1968).
(i) Chamaeleo peyrieresi added, described by
Brygoo, Blanc & Domergue (1974).
() Chamaeleo tsaratananensis added,
described by Brygoo & Domergue (1967/1968).
(k) Chamaeleo tuzetae added, described by
Brygoo, Bourgat & Domergue (1972).
() Also see BOhme and Klaver for further recent
changes.
Squamata; Iguanidae: several sources, notably;
(1) Montanucci, (1970), (Crotaphytus wislizenii
silus should be regarded as a full species. C. silus).
(2) Collins et al, (1978), (Crotaphytus silus
assigned to Gambelia).
(3) Carey, (1975), Schwartz & Thomas, (1975),
Schwartz and Carey, (1977), (Cyclura).
(4) Peters & Donoso-Barros, (1970), Schwartz &
Thomas, (1975), (iguana).
(5) Gibbons, (1981), (mew Brachylophus sp.)
Squamata; Lacertidae: generic assignment after
Arnold, (1973).
Squamata; Cordylidae: after Wermuth, (1968).
Squamata; Teiidae: several sources, notably;
(1) Peters & Donoso Barros, (1970)
(2) Vanzolini & Valencia, (1965), (Dracaena).
(3) Presch, (1973), (Tupinambis, only two species
recognised)
Squamata; Helodermatidae, Varanidae: follows
Mertens (1963), except;
(a) Varanus acanthums primordius regarded as
a full species after Cogger (1975).
(b) Varanus storm added, described by Mertens
(1966c).
Squamata; Boidae: several sources, see notes on
p44 and p46
Squamata; Colubridae: several sources, notably;
(1) Peters & Orejas-Miranda, (1970), (Clelia
clelia).
(2) Gans & Williams, (1954), Fleming and
Fleming, (1973), 433 (Elachistodon).
(3) Dowling & Gibson, (1970); (Cyclagras gigas
assigned to Hydrodynastes).
(4) Rossman & Eberle, (1977); (Natrix sipedon
and N. fasciata assigned to Nerodia).
Squamata; Elapidae, Viperidae: chiefly after
Klemmer, (1963).
Explanatory Key
For each taxon, there are at least two entries in the
four columns down the right-hand side of the page.
CITES: LII, or III, referring to the CITES Appendix on
which the taxon is listed as at June 1981.
RDB: Red Data Book status categones,
E = Endangered
V = Vulnerable
R= Rare
O = Outof danger
Id = Indeterminate Given as ‘Tl’ in RDB, but
modified here to avoid confusion with the
CITES ‘T’ entry).
K = Insufficiently Known; suspected of being
threatened, but without full data.
Exp: Major exploitation of taxon, where known.
F = Food (may include localand commercial use)
LA = Live animal trade (primarily for the pet
trade)
Pr = Preserved specimens
SK = Skins, for the leather trade
TS = Tortoise-shell
When known, a category of secondary
importance is given in brackets.
Ref. Entry hereisanumberreferringtoanitem inpart
(A) ofthe list of References providing incidental or
exhaustive information on the survival status of the
taxon.
10
Class: AMPHIBIA
Order: CAUDATA (Urodela),
Newts and Salamanders
Family: CRYPTOBRANCHIDAE
Andrias davidianus (Megalobatrachus davidianus)
Chinese Giant Salamander
China
Andrias japonicus (Megalobatrachus japonicus)
Japanese Giant Salamander
Japan
Family: AMBYSTOMATIDAE
Ambystoma californiense (A. tigrinum californiense)
California Tiger Salamander
USA (California)
Ambystoma dumerilii
Mexico
(RDB entry is for A.d. dumerilii, the Lake Patzcuaro Salamander or Achoque
from Lake Patzcuaro, Michoacan; the remaining subspecies A.d.
queretarensis, Michoacan, is not included)
Ambystoma lermaensis
Lake Lerma Salamander
Mexico
Ambystoma macrodactylum croceum
Santa Cruz Long-toed Salamander
USA (California)
Ambystoma mexicanum
Axolotl
Mexico (Lake Xochimilco region)
Family: SALAMANDRIDAE
Chioglossa lusitanica
Gold-stnped Salamander
Portugal, Spain
Family: PLETHODONTIDAE
Batrachoseps aridus
Desert Slender Salamander
USA (California)
Batrachoseps simatus
Kem Canyon Slender Salamander
USA (California)
CITES RDB
Exp Ref
- 41,414
43, 67, 75
LA 43
-— 16,38, 43
— 4 43,75
— 443,75
CITES
Batrachoseps stebbinsi =
Tehachapi Slender Salamander
USA (California)
Eurycea nana ~
San Marcos Salamander
USA (Texas)
Hydromantes brunus -
Limestone Salamander
USA (Califommia)
Hydromantes shastae =
Shasta Salamander
USA (California)
Phaeognathus hubrichti =
Red Hills Salamander
USA (Alabama)
Plethodon larselli =
Larch Mountain Salamander
USA (Oregon, Washington)
Plethodon neomexicanus =
Jemez Mountains Salamander
USA (New Mexico)
Plethodon nettingi (P. richmondi nettingi) =
Cheat Mountain Salamander
USA (West Virginia)
Typhlomolge rathbuni (Eurycea rathbuni) =
Texas Blind Salamander
USA (Texas)
Typhlotriton spelaeus -
Grotto Salamander
USA
Family: PROTEIDAE
Proteus anguinus -
Olm
Yugoslavia (and introduced at Vicenza, Italy)
RDB Exp Ref
R 4 43, 75
R - 443
R — 443,75
R 4 43,75
R - 43
Id -— 448
Vi > 4743/78
Ke £
E - 44351
K = =
VLA 38,43
CITES RDB Exp Ref
Order: ANURA (Salientia)
Frogs and Toads
Family: LEIOPELMATIDAE
(LEIOPELMIDAE; Leiopelma is sometimes
included in the ASCAPHIDAE)
Leiopelma archeyi = Re 43, 64
Archey’s Frog, Coromandel Leiopelma 06,107
New Zealand
Leiopelma hamiltoni = R - 26,
Hamilton's Frog, Stephen Island Leiopelma ee
New Zealand
Leiopelma hochstetteri = R= 43, 64
Hochstetter'’s Frog, North Island Leiopelma 106,107
New Zealand
Family: PIPIDAE
Xenopus gilli = Vi sonar
Cape Platanna, Cape Clawed Toad
South Afnca (Cape Town region)
Family: DISCOGLOSSIDAE
Discoglossus nigriventer = Em 4s
Israel Painted Frog
Israel (Lake Huleh)
Family: PELOBATIDAE
Pelobates fuscus insubricus = Ey =) oss
Italian Spadefoot Toad
Italy (plains of River Po). Switzerland (Ticino, probably extinct)
Family: LEPTODACTYLIDAE
Batrachophrynus macrostomus - Kel
Lake Junin (Giant) Frog
Peru (Lake Junin)
Eleutherodactylus jasperi = R = &
Golden Coqui Frog
Puerto Rico
Family: MYOBATRACHIDAE
(altematively regarded as a subfamily of LEPTODACTYLIDAE)
Philoria frosti = R - 43
Baw-baw Frog
Australia
CITES RDB Exp Ref
Rheobatrachus silus = Wi 2g
Platypus Frog
Australia (Queensland)
Family: BUPFONIDAE
Atelopus varius zeteki I — LA 50
Golden Frog, Zetek's Golden Frog (Pr)
Panama (Valle de Anton)
(Atelopus is sometimes assigned to a separate family, ATELOPODIDAE
Bufo boreas nelsoni - el = whee
Amargosa Toad
USA (Califommia, Nevada)
Bufo exsul - V LA 4,43,75
Black Toad, Deep Springs Toad
USA (California)
(sometimes treated as a subspecies of B. boreas, B.boreas exsul)
Bufo houstonensis = Ee Aner4gh79
Houston Toad
USA (Texas)
Bufo periglenes I 5) oe 28
Golden Toad, Orange Toad
Costa Rica
Bufo retiformis Il VLA 4,43
Sonoran Green Toad
Mexico (Sonora), USA (Arizona)
Bufo superciliaris I Se id= az
Cameroon Toad
Cameroon, Equatorial Guinea, Gabon, Ghana, Guinea, Nigeria,
Zaire, probably also in other countries of west and west-central
Nea
Africa
Nectophrynoides cryptus I a ks ey
Viviparous African Toad
Tanzania (Uluguru Mts.)
Nectophrynoides liberiensis I ree =
Liberia (Mt. Nimba)
Nectophrynoides malcomi I a Se La
Ethiopia
Nectophrynoides minutus I eo en ee
Tanzania (Uluguru Mts.)
Nectophrynoides occidentalis I Vo = &
Mt. Nimba Viviparous Toad
Guinea (Mt. Nimba), and adjacent parts of Ivory Coast
CITES RDB Exp Ref
Nectophrynoides osgoodi
Ethiopia
Nectophrynoides tomieri
Tanzania (Magrotto and Usambara Mts.)
Nectophrynoides viviparus
Tanzania (Kinga, Rungwe, and Uluguru Mts.)
Family: HYLIDAE
Hyla andersoni
Pine Barrens Tree Frog
USA
(Florida populations noted as Endangered in RDB)
Pseudacris streckeri illinoensis
Ilhnois Chorus Frog
USA
Family: SOOGLOSSIDAE
(sometimes treated as a subfamily of
RANIDAE)
Nesomantis thomasseti
Thomasset's Seychelles Frog
Seychelle Is. (Mahe, Silhouette)
Sooglossus sechellensis
Seychelles Frog
Seychelle Is. (Mahe, Silhouette)
Sooglossus gardineri
Gardiners Seychelles Frog
Seychelle Is. (Mahe, Silhouette)
Family: RANIDAE
Conraua goliath (Rana goliath, Gigantorana goliath)
Goliath Frog
Cameroon, Equatonal Guinea
Rana onca (inc. Rana pipiens fisheri)
Relict Leopard Frog (inc. Vegas Valley Leopard Frog)
USA (Anzona, Utah, Vegas Valley Leopard Frog restricted to Nevada)
Family: HYPEROLIDAE
(sometimes treated as a subfamily of
RHACOPHORIDAE)
Megalixalus seychellensis (M. infrarufus)
Seychelle Islands Tree Frog
Seychelle Islands (Mahe, Praslin, La Digue, Silhouette)
I
4,43
43
43
43
4,43
43
Class: REPTILIA
Order: TESTUDINES,
Tortoises, Turtles
Suborder: CRYPTODIRA
Family: KINOSTERNIDAE
Kinosternon augustipons
Narrow-bndged Mud Turtle
Costa Rica, Nicaragua, Panama
Kinosternon creaseri
Creaser's Mud Turtle
Mexico
Kinosternon dunni
Dunn's Mud Turtle
Columbia
Sternotherus minor depressus
Flattened Musk Turtle
USA
Family: DERMATEMYDIDAE
Dermatemys mawii
Central Amenican River Turtle
Belize, Guatemala, Mexico
Family: EMYDIDAE
Batagur baska
Common Batagur, River Terrapin, Tuntong
Bangladesh, Burma, India, Indonesia, Kampuchea
Malaysia, Thailand, Vietnam?
Callagur borneoensis
Painted Terrapin
Indonesia (Kalimantan, Sumatra), Malaysia, Thailand
Clemmys muhlenbergii
Bog Turtle, Muhlenberg's Turtle
USA
Geoclemys hamiltonii (Damonia hamiltonii)
Black Pond Turtle, Spotted Pond Turtle
Bangladesh, India, Pakistan
Heosemys silvatica
Kavalai Forest Turtle
India (Kerala)
Kachuga tecta tecta
Indian Sawback or Tent Turtle, Roofed or Dura Turtle
Bangladesh, India, Pakistan
Melanochelys tricarinata (Geoemyda tricarinata,
Nicoria tricarinata)
Three-keeled Land Tortoise
Bangladesh, India
16
CITES
RDB Exp Ref
R ~ 52
R - 52
R - 52
Id - 152
V le 9, 152
FE IF 54,136
Vv le 152
leis (iA) 274'25;31,
113,152
= LA =
Id - 152
Morenia ocellata
Burmese Swamp Turtle
Burma
Pseudemys alabamensis
Alabama Red-bellied Turtle
USA
Pseudemys felis
Cat Island Turtle
Bahamas (Cat Island)
Pseudemys malonei (Pseudemys terrapen malonei,
Chrysemys malonei)
Inagua Island Turtle
Bahamas (Great Inagua Island)
Pseudemys rubriventris bangsi
Plymouth Red-bellied Turtle
USA
Pseudemys scripta callirostris (Pseudemys ornata
callirostris, Chrysemys ornata callirostris)
South Amenican Red-lined Turtle
Colombia, Venezuela
Rhinoclemmys areolata
Furrowed Wood Turtle
Belize, Guatemala, Honduras, Mexico
Rhinoclemmys rubida
Mexican Spotted Wood Turtle
Mexico
Terrapene coahuila
Aquatic or Water Box Turtle
Mexico (Coahuila)
Family: TESTUDINIDAE
Chersina angulata (Testudo angulata)
Angulated or Bowspnit Tortoise
Namibia, South Africa
Geochelone carbonaria (Testudo carbonaria)
Red-foot Tortoise
Brazil, Colombia, French Guiana, Guyana,
Lesser Antilles, (several islands, probably introduced),
Panama, Paraguay, Surinam, Trinidad, Venezuela
Geochelone chilensis (inc. G.donosobarrosi and
G.petersi; Testudo chilensis)
Argentine Tortoise, Chaco Tortoise
Argentina, Paraguay, probably Bolivia
Geochelone denticulata (Testudo denticulata)
Yellow-foot Tortoise, Jaboti, Forest Tortoise,
Brazilian Giant Tortoise, Hercules Tortoise
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana,
Peru, Surinam, Trinidad, Venezuela, possible Paraguay
CITES
ine)
I
II
RDB Exp Ref
Id — 152
R = 152
R ~ 152
R ~ 152
V Fe 152
LA
K - 152
K - 152
V LA 9,152
ms ie =
K E 152
LA
K IF 34,104
LA 152
= B i=
LA
17
Geochelone elegans (Testudo elegans)
Indian Starred or Star Tortoise
India, Pakistan, Sn Lanka
Geochelone elephantopus (Testudo elephantopus)
Galapagos Giant Tortoise
Galapagos Islands (Ecuador)
Geochelone elongata (Testudo elongata)
Yellow Tortoise
Bangladesh, Burma, China, India, Kampuchea, Laos, Malaysia,
Nepal, Thailand, Vietnam
Geochelone emys (Testudo emys)
Burmese Brown Tortoise
Burma, India, (Assam), Indonesia, Kampuchea, Laos?,
Malaysia, Thailand, Vietnam?
Geochelone forstenii (Testudo forstenii)
Sulawesi (Celebes) Tortoise
Indonesia (Halmahera, Sulawesi)
Geochelone gigantea (Testudo gigantea)
Aldabra Giant Tortoise
Aldabra, Seychelle Islands (introduced)
Geochelone impressa (Testudo impressa)
Impressed Tortoise
Burma, China, Malaysia (peninsula), Thailand, Vietnam,
probably also Kampuchea and Laos
Geochelone pardalis (Testudo pardalis)
Leopard Tortoise, Mountain Tortoise
Angola, Botswana, Ethiopia, Kenya, Lesotho, Malawi,
Mozambique, Namibia, Somalia, South Africa, Sudan, Swaziland,
Tanzania, Uganda, Zambia, Zimbabwe
Geochelone platynota (Testudo platynota)
Burmese Starred Tortoise
Burma
Geochelone radiata (Testudo radiata, Asterochelys
radiata)
Radiated Tortoise
Madagascar
Geochelone sulcata (Testudo sulcata)
Spurred or Grooved Tortoise
Chad, Ethiopia, Mali, Mauntania, Niger, Nigena, Senegal,
Sudan, probably also Cameroon, Central Afncan Republic,
Djibouti, Upper Volta
Geochelone travancorica (Testudo travancorica)
Travancore Tortoise
India
Geochelone yniphora (Testudo yniphora,
Asterochelys yniphora)
Madagascar Tortoise, Angonoka
Madagascar
18
CITES
II
Il
II
Il
I
RDB Exp
LA
)
LA
LA
Ref
54,152
48,152
46,152
152
45,152
Gopherus agassizii (G. polyphemus agassizii)
Desert Tortoise, Western Gopher Tortoise
Mexico, USA
Gopherus berlandieri (G.polyphemus berlandieri)
Texas Gopher Tortoise, Berlandier’s Tortoise
Mexico, USA (Texas)
Gopherus flavomarginatus (G.polyphemus
flavomarginatus)
Bolson Tortoise, Mexican Giant Gopher Tortoise
Mexico
Gopherus polyphemus
Flonda Gopher Tortoise
USA
Homopus areolatus
Parrot-beaked or Areolated Tortoise
South Afnca
Homopus boulengeri
Boulengers Tortoise
Namibia, South Afnca
Homopus femoralis
Karroo Tortoise
South Afnca
Homopus signatus
Speckled Tortoise
Namibia, South Afnca
Kinixys belliana
Bell's Hinged Tortoise
Angola, Benin, Botswana, Burundi?, Cameroon, Central Afncan
Republic, Congo, Djibouti, Ethiopia, Gabon, Gambia, Ghana,
Guinea, Guinea Bissau, Ivory Coast?, Kenya, Lesotho?, Libena,
Malawi, Mali, Mozambique, Namibia, Nigena, Rwanda, Senegal,
Sierra Leone, Somalia, South Africa, Sudan, Swaziland, Tanzania,
Togo, Uganda, Zaire, Zambia, Zimbabwe, (introduced on
Madagascar)
Kinixys erosa
Schweigger's, Common, or Forest Hinged Tortoise
Angola, Benin, Cameroon, Central Afncan Republic, Congo,
Equatonal Guinea, Gabon, Gambia, Ghana, Guinea, Ivory
Coast, Liberia, Nigeria, Senegal, Sierra Leone, Togo, Uganda,
Upper Volta, Zaire, (introduced on Madagascar)
Kinixys homeana
Home's Hinged Tortoise
Benin, Cameroon, Congo, Equatorial Guinea, Gabon, Ghana,
Guinea, Ivory Coast, Libena, Nigeria, Togo, Zaire
Malacochersus tornieri
Pancake Tortoise
Kenya, Tanzania
CITES
II
RDB Exp
V
Id
LA
LA
LA
LA
LA
LA
LA
Ref
4,15,29
75,144,
52
52
101,144,
152
432,133,
152
43,152
19
Psammobates geometricus (Testudo geometrica)
Geometric Tortoise
South Afnca
Psammobates oculifer (Testudo oculifera)
Serrated Tortoise, Toothed Cape Tortoise, Kuhl's Tortoise
Botswana, Namibia, South Afnca
Psammohbates tentorius (Testudo tentoria)
Tent Tortoise
Namibia, South Afnca
Pyxis arachnoides
Spider Tortoise
Madagascar
Pyxis planicauda (Testudo planicauda, Acinixys
planicauda)
Madagascar Flat-shelled Tortoise
Madagascar
Testudo graeca
Spur-thighed, Common, or Greek Tortoise
Albania, Algena, Bulgana, Egypt, Greece, Iran, Iraq, Israel,
Italy (including Sardinia and Sicily, probably introduced),
Jordan, Lebanon, Libya, Malta, Morocco, Romania, Spain (inc.
Balearic Islands), Syna, Tunisia, Turkey, USSR, Yugoslavia,
(introduced in southem France)
(1. RDB entry is for T.graeca graeca) from Algena, Libya, Morocco, Spain,
Tunisia, and introduced in France
2. Populations in eastern Iran are occasionally treated as a full species,
T.zarudnyi)
Testudo hermanni
Hermann's Tortoise, Greek Tortoise
Albania, Bulgana, France (inc. Corsica), Greece, Italy
(inc. Sardinia, Sicily), Romania, Spain (Balearic Islands, also
reported from mainland), Turkey, Yugoslavia
Testudo horsfieldii (Agrionemys horsfieldii)
Horsfield’s Tortoise, Afghan Tortoise
Afghanistan, China, Iran, Pakistan, USSR
Testudo kleinmanni
Egyptian Tortoise
Egypt, Israel, Libya
Testudo marginata
Margined Tortoise
Greece, Italy (Sardinia, introduced, possibly mainland also)
20
CITES
II
II
I
RDB. Exp
VA:
= Ibis
= Iba
Id LA
Idi =
Ven ba
VLA
= by
igh =
= Ive
Ref
43,47,63
152
43,152
13,38
152
13,56
152
13,38
CITES RDB Exp
Family: CHELONIIDAE
(Because of the wide and somewhat unpredictable distribution of sea turtles
away from the nesting beaches, it is impractical to list those countries in whose
territorial waters individual sea turtles may be expected to occur. This applies
to members of the CHELONIIDAE and DERMOCHELYDIDAE.)
Caretta caretta
Loggerhead Turtle
Nesting recorded on beaches of tropical, subtropical, and
temperate seas (Atlantic, Indian, Mediterranean, Pacific),
wandering individuals recorded in temperate and arctic
waters (to 70°N at Murmansk, USSR, and to 35°S at Rio de la
Plata, Argentina)
Chelonia depressa
Flatback Turtle
Nests in north and north east Australia, not recorded outside
adjacent waters
Chelonia mydas
Green Turtle
Nesting recorded on beaches of tropical and subtropical seas
(Atlantic, Indian, Mediterranean, Pacific), wandering
individuals recorded in temperate waters (to ca. 52°N in
English Channel, and to 43°S in Chile). Major breeding and
feeding areas within 20°N to 20°S.
Breeding populations in East Pacific sometimes regarded
as a full species Chelonia agassizii.
Eretmochelys imbricata
Hawksbill Turtle
Nesting recorded at scattered localities on beaches of tropical
and subtropical seas, usually between 25°N and 25°S (Atlantic,
Indian, Pacific, rarely or formerly in Mediterranean),
wandering individuals occasionally recorded in temperate
regions,
Lepidochelys kempii
Kemp's Ridley, Atlantic Ridley
Single major nesting site is in Mexico (Rancho Nuevo, on Gulf
coast of State of Tamaulipas), occurs most frequently in Gulf of
Mexico, wandering individuals recorded in Atlantic waters (to
58°N in Scotland), and very occasionally in Mediterranean
Lepidochelys olivacea
Olive Ridley, Pacific Ridley
Nesting recorded on beaches of tropical and subtropical seas
(Atlantic, Indian, Pacific), most sites between 30°N and 20°S
occasionally recorded in the Caribbean region although does
not appear to breed there
(SK)
(SK)
(SK)
(SK)
Ref
4,6,25,38
40,41,43,
54,68,69,
103,123,
126,133,
139,152
41,43, 130,
152
4,6,18,25,
33,37,38,
40, 41,43,
54,62,65,
68,69,88,
99,114,
117,120,
124,130,
133,139)
146,152
4,6,22,25
30,33,38,
39,40,41,
43,54,62,
68,69,120,
128,130,
133,139,
146,152
120,128,
130,133,
139,146,
4,6,23,25,
38,41,43,
69,91,133,
152
6,40,41,
43,55,62,
65,68,90,
91,121,
130,139,
146,152
al
CITES RDB Exp
Family: DERMOCHELYDIDAE
(DERMOCHELYIDAE)
(see note under Family: CHELONIIDAE, p21)
Dermochelys coriacea I ig oP
Leatherback, Leathery Turtle, Luth
Nesting recorded on beaches of tropical and subtropical seas
(Atlantic, Indian, Mediterranean, Pacific), most sites between 30°N
and 20°S, wandenng individuals recorded in temperate waters
(north to Iceland and Norway)
Family: CARETTOCHELYIDAE
Carettochelys insculpta = KF
New Guinea Plateless Turtle
Australia (Northem Termitory), Indonesia (Inan Jaya),
Papua New Guinea
Family: TRIONYCHIDAE
Lissemys punctata andersoni (L.p.punctata) I =
Indian Flap-shell Turtle
Bangladesh, Burma (Akyab), India, Nepal, Pakistan
Trionyx ater I = =
Cuatro Cienegas or Black Soft-shell Turtle
Mexico
Trionyx gangeticus I =SE
Indian or Ganges Soft-shell Turtle
Afghanistan, Bangladesh, India, Nepal, Pakistan
Trionyx hurum I = iF
Peacock-marked Soft-shell Turtle
Bangladesh, India
Trionyx nigricans I i oe
Dark Soft-shell Turtle
Bangladesh
Trionyx triunguis Ill - =
Nile Soft-shell Turtle (Ghana)
Angola (inc. Cabinda, and south to Cunene River on Angola-
Namibia border), Cameroon, Chad, Egypt, Ethiopia, Gabon,
Gambia, Ghana, Guinea Bissau, Israel, Kenya, Libena, Niger,
Nigena, Senegal, Sierra Leone, Somalia, Sudan, Togo, Turkey
(south), Uganda, Zaire, probably also Benin, Central Afncan
Republic, Congo, Djibouti, Equatonal Guinea, Guinea,
Ivory Coast, Mali, and Upper Volta
22
Ref
4,6,25,38,
40,41,43,
54,57,62,
68,69, 70,
87,116,
121,122,
130,132,
133,146,
152
152
152
CITES RDB Exp
Suborder: PLEURODIRA,
Side-necked Turtles
Family: PELOMEDUSIDAE
Erymnochelys madagascariensis (Podocnemis II? il =
madagascariensis)
Madagascar
(Until recently (see note on p8), a majority of taxonomists have assigned this
species, and Peltocephalus dumeniliana, to the genus Podocnemis. All species
of Podocnemis are listed on CITES Appendix II. It is here assumed, pending
discussion of the matter that the intention was to include these two species also,
regardless of a change in generic assignment)
Pelomedusa subrufa Ill = =
Helmeted Turtle (Ghana)
Angola, Benin, Botswana, Cameroon, Ethiopia, Ghana, Kenya,
Madagascar, Malawi, Mali, Mauntania, Mozambique, Namibia,
Nigena, Saudi Arabia (southwest), Senegal, Somalia, South Afnca,
Sudan, Tanzania, Togo, Uganda, North Yemen, Zaire, Zambia,
Zimbabwe, probably also Central Afncan Republic, Chad, Congo,
Djibouti, Equatorial Guinea, Gambia, Guinea, Guinea Bassau,
Ivory Coast, Libena, Niger, Upper Volta
Peltocephalus dumeriliana (Peltocephalus tracaxa, II? = aes
new name; Podocnemis dumeriliana)
Brazil, Colombia, Ecuador? French Guiana? Peru, Venezuela
(see note under Erymnochelys madagascanensis, above)
Pelusios spp.
Ghana has proposed that all species of Pelusios found in Ghana be
placed on Appendix III, the following four species, of the eight
currently recognised by Wermuth & Mertens (1977), may be
expected to occur in Ghana. The systematics of this genus is
subject to some dispute
Pelusios adansonii I - =
White-breasted Side-necked Turtle, Adanson's Turtle (Ghana)
Angola, Chad, Congo, Gambia, Mali, Nigena, Senegal, Sudan,
Zaire, possibly Ghana; Cape Verde Is?
Pelusios castaneus (formerly inc. P.subniger) Il = =
Angola, Burundi, Congo, Gambia, Guinea, Liberia, Madagascar,
Malawi, Rhodesia, Rwanda, Senegal, Seychelle Islands, Sierra
Leone, Tanzania, Zaire, Zambia, possibly Ghana
Pelusios gabonensis Il = =
Stnpe-backed Side-necked Turtle, Gaboon Turtle (Ghana)
West Africa (from Liberia east, inc. Ghana, to Gabon)
Pelusios niger I = =
Black Side-necked Turtle (Ghana)
West Africa (from Gambia east, possibly inc. Ghana, to Angola)
Podocnemis erythrocephala (Podocnemis cayennensis) II Kael
Irapuca, Red-headed Amazon Turtle
Brazil, Venezuela, perhaps Colombia
Ref
53,135
23
Podocnemis expansa
Arrau, South Amenican River Turtle
Bolivia, Brazil, Colombia, Ecuador?, Guyana, Peru, Venezuela,
perhaps French Guiana and Sunnam
Podocnemis lewyana
Magdalena River Turtle
Colombia
Podocnemis sextuberculata
Brazil, Colombia, Peru
Podocnemis unifilis
Terecay, Tracaja, Yellow-spotted Amazon Turtle
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru,
Surinam, Venezuela
Podocnemis vogli
Colombia, Venezuela
Family: CHELIDAE
Phrynops dahli
Dahl's Toad-headed Turtle
Colombia (Bolivar)
Phrynops hogei
Hoge's Sideneck Turtle
Brazil
Phrynops rufipes
Red-headed Sideneck Turtle
Brazil, Colombia
Platemys pallidipectoris
Chaco Sideneck Turtle
Argentina, Bolivia?, Paraguay
Platemys spixii
Spix's Sideneck Turtle
Argentina, Brazil, Uruguay
Pseudemydura umbrina
Shortnecked or Western Swamp Turtle
Australia (Bullsbrook)
24
CITES RDB Exp
II
II
II
E
Id
P
LA)
(LA)
(LA)
Ref
43,53
14,92
135,143,
152
53,135
152
53,135
143,152
92,119
135,143
152
53,135
152
152
152
43,57
89,105,
130,138
CITES RDB Exp Ref
Order: CROCODYLIA
For the many and variable trade names of Crocodylia, see King & Brazaitis,
1971, and Wermuth & Fuchs, 1978, (See Reference list C.p61)
Family: ALLIGATORIDAE
Alligator mississippiensis
Amencan Alhgator
USA
Alligator sinensis
Chinese Alligator
China
Caiman crocodilus sub-spp.
Caiman crocodilus apaporiensis
(C.sclerops apaporiensis)
Rio Apaponis (Spectacled) Caiman
Colombia
Caiman crocodilus (C.sclerops)
3 to 5 remaining subspecies
Spectacled Caiman
Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador,
French Guiana, Guatemala, Guyana, Honduras, Mexico,
Nicaragua, Panama, Paraguay, Peru, El Salvador, Surinam,
Tnnidad & Tobago, Uniguay, Venezuela (C.c.yacare, the
Jacaré, from Argentina, Bolivia, Brazil, Paraguay, Uruguay, is
sometimes treated as a full species)
(In Part 1 of the new Amphibia-Reptilia RDB (ref. 152) the three widely
recognised subspecies are categorised as follows. C.C. crocodilus ‘V’;
C.c. fuscus 'V' and C.c. yacare 'Id’.)
Caiman latirostris
Broad-nosed Caiman
Argentina, Brazil, Paraguay, Uruguay
Melanosuchus niger
Black Caiman
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana,
Paraguay, Peru
Paleosuchus palpebrosus
Dwarf Caiman, Cuvier's Smooth-fronted Caiman
Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru.
Surinam, Venezuela
Paleosuchus trigonatus
Schneider's Smooth-fronted Caiman
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Per,
Sunnam, Venezuela
II
— (LA) 4,43,44
(SK) 60,100,
133,152
E F 42,43, 102,
SK 152
Id SK 42,43,152
Id/V SK 42,43,
(LA) 152,153
E SK 42,43,152
(LA)
E SK 42,43,152
(LA)
— (SK) 42,43,152
— (SK) 42,43,152
25
Family: CROCODYLIDAE
Crocodylus acutus
American Crocodile
Belize, Colombia, Costa Rica, Ecuador, Greater Antilles (Cuba,
Domonican Republic, Haiti, Jamaica), Guatemala, Honduras,
Mexico, Nicaragua, Panama, Peru, El Salvador, USA (Florida),
Venezuela
Crocodylus cataphractus
African Sharp-nosed or Slender-snouted Crocodile
Angola (Cabinda), Benin, Cameroon, Central African Republic,
Chad, Congo, Equatorial Guinea, Gabon, Gambia, Ghana,
Guinea, Guinea Bissau?, Ivory Coast, Liberia, Mali, Mauntania,
Nigena, Senegal, Sierra Leone, Tanzania (Uji), Togo, Upper
Volta, Zaire, Zambia
Crocodylus intermedius
Onnoco Crocodile
Colombia, Venezuela, (and single record from Grenada)
Crocodylus johnsoni (C.johnstoni)
Australian Fresh-water Crocodile, Johnson's Crocodile
Australia
Crocodylus moreletii
Morelet's Crocodile
Belize, Guatemala, Mexico
Crocodylus niloticus
Nile Crocodile
Angola, Benin, Botswana, Burundi, Cameroon, Central Afncan
Republic, Chad, Congo, Egypt, Equatorial Guinea, Ethiopia,
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast,
Kenya, Liberia, Madasgascar, Mali, Malawi, Mozambique,
Namibia, Niger, Nigena, Rwanda, Senegal, Sierra Leone,
CITES RDB-Exp
I
Somalia, South Afncea, Sudan, Swaziland, Tanzania, Togo, Uganda,
Upper Volta, Zaire, Zambia, Zimbabwe
Crocodylus novaeguineae sub-spp.
C.n.mindorensis
Mindoro or Philippines Crocodile
Philippine Islands
(sometimes treated as a full species, C. mindorensis, as in RDB)
C.n.novaeguineae
New Guinea Crocodile
Indonesia (Aru Islands, West Inan), Papua New Guinea
Crocodylus palustris
Mugger, Marsh Crocodile, Broad-snouted Crocodile
Bangladesh, India, Iran (Sarbaz River), Nepal, Pakistan, Sn Lanka,
(Burma?)
26
II
SK
SK
)
SK
SK
SK
SK
(F)
SK
SK
SK
Ref
4,42,43,
60,133,
137,145,
152
42,43
93,152
42,43,
134,152
42,43, 130,
147,152
42,43,60,
61,80,94,
127,152
42,43,47,
86,93, 109,
152
42,43, 152
42,43,
112,152
42,43,44,
85,96, 149,
150,152
CITES RDB Exp
Crocodylus porosus I ig” = Sx
Estuanine or Salt-water Crocodile ui
(Papua New
Australia, Bangladesh, Brunei, Burma, India (inc. Andaman Guinea)
Islands), Indonesia, Kampuchea, Malaysia, Palau Islands, Papua
New Guinea, Philippine Islands, Solomon Islands, Sn Lanka,
Thailand, Vanuatu, Vietnam, also records of wandering individuals
from the Cocos-Keeling Islands and Fiji Islands. Extinct in
Singapore
Crocodylus rhombifer I is. SK
Cuban Crocodile
Cuba (inc. Isla de Pinos)
Crocodylus siamensis I Is Six
Siamese Crocodile
Indonesia (Borneo, Java), Kampuchea, Laos, Malaysia
(peninsula), Thailand, Vietnam
Osteolaemus tetraspis I Id SK
West Afncan Dwarf Crocodile ©)
Angola (Cabinda), Benin, Cameroon, Central Afncan Republic,
Congo, Gabon, Gambia, Ghana, Guinea, Guinea Bissau,
Ivory Coast, Libena, Nigena, Senegal, Sierra Leone, Togo,
Uganda, Upper Volta, Zaire
(the Zaire population is sometimes treated as a full species, O.osbomni)
Tomistoma schlegelii I ig 9K
False Ghanal, False Gavial
Indonesia (Borneo, Sumatra), Malaysia (peninsula, Sarawak),
Thailand (possibly extinct)
Family: GAVIALIDAE
Gavialis gangeticus I Evi y ik
Ghanal, Gavial
Bangladesh, Bhutan (probably extinct), India, Nepal, Pakistan
Order: RHYNCHOCEPHALIA
Family: SPHENODONTIDAE
Sphenodon punctatus I R -
Tuatara
New Zealand
Ref
42,43, 44,
85,98, 130,
147,149,
152
42,43,60,
145,152
42,43,98,
152
42,43,93,
152
42,43, 152
3,42,43,
65,71,84,
142,149,
152
Me
CITES RDB Exp Ref
Order: SQUAMATA,
Lizards, Snakes, Amphisbaenians
Suborder: SAURIA, Lizards
Family: GEKKONIDAE
Cyrtodactylus serpensinsula
(Gymnodactylus serpensinsula) II Ree eela4e:
Serpent Island Gecko a 7,
Round Island, Serpent Island, (Mauritius)
Oedura reticulata = We = ZSHIE0)
Reticulated Velvet Gecko
Australia
Phelsuma spp.
Genus Phelsuma: Day Geckos, certain species may appear in the Live
Animal trade. The systematics of this genus 1s subject to some dispute.
Phelsuma abbotti I tbe ST Fie
Abbotts Day Gecko
Aldabra, Amirante Islands, Assumption, Cosmoledo Island,
Madagascar, Seychelle Islands
(P.a. longinsulae and P.a pulchra from the Seychelles are sometimes regarded
as a separate species, P.longinsulae)
Phelsuma agalegae II - — 24
Agalega Island (Mauritius)
Phelsuma andamanensis Il yo Zee
Andaman Day Gecko
Andaman Islands (India)
Phelsuma astriata Il geare STR
Astove Island, Seychelle Islands
Phelsuma barbouri Il Se ae
Madagascar
Phelsuma bimaculata II Fey eat aR
Madagascar
(sometimes treated as a subspecies of P.quadriocellata)
Phelsuma breviceps Il ee LS
Madagascar
(sometimes treated as a subspecies of P. mutabilis)
Phelsuma cepediana I = = Th
Mauntius Greater Day Gecko
Mauntius, Reunion
(ithas been suggested that P.c. borbonica from Reunion should be regarded
as a full species)
28
Phelsuma comorensis
Comoro Islands
(sometimes treated as a subspecies of P. dubia)
Phelsuma dubia
Madagascar, Tanzania (inc. Zanzibar)
Phelsuma edwardnewtonii (P newtoni)
Rodriguez Day Gecko
Rodnguez Island (Mauntius)
(has been reported extinct, see Ref. 82)
Phelsuma flavigularis
Madagascar
Phelsuma guentheri
Round Island Day Gecko
Round Island (Maunitius)
Phelsuma guimbeaui
Mauritius
Phelsuma guttata
Madagascar
Phelsuma laticauda
Gold-dust Day Gecko
Comoro Islands, Farquhar Island (Seychelles)?, Madagascar
Phelsuma lineata
Side-smped Day Gecko
Madagascar
Phelsuma madagascariensis
Madagascar Day Gecko
Madagascar, Seychelle Islands
(P.m. sundbergi from the Seychelles is frequently regarded asa full species)
Phelsuma modesta
Madagascar
Phelsuma mutabilis
Madagascar
Phelsuma ocellata (Rhoptropella ocellata)
Namibia, South Afnca
Phelsuma omnata (P. vinsoni)
Mauritius (inc. Round Island), Reunion
CITES RDB Exp Ref
II
43,82
47
12, 76
17, 83
2g
CITES
Phelsuma parkeri II
Pemba Island (Tanzania)
(sometimes treated as a subspecies of P. abbotti)
Phelsuma quadriocellata II
Madagascar
Phelsuma serraticauda Il
Madagascar
(sometimes treated as a subspecies of P. flavigularis)
Phelsuma standingi II
Madagascar
Phelsuma trilineata II
Madagascar
Phelsuma v-nigra II
Comoro Islands
(sometimes treated as a subspecies of P. abbotti)
Family: PYGOPODIDAE
Paradelma orientalis II
Queensland Snake-lizard
Australia
Family: XANTUSIDAE
Xantusia riversiana (Klauberina riversiana) =
Island Night Lizard
USA (Califormia)
Family: AGAMIDAE
Hydrosaurus pustulatus =
Sail-fin Lizard, Soa-Soa Water Lizard, (Crested Lizard)
Phihppine Islands
Uromastyx spp.
Species of Uromastyx are widely termed Spiny-tailed Lizards, Palm Lizards,
or Mastigures
Uromastyx acanthinurus II
Black Spiny-tailed Lizard
Algena, Chad, Egypt, Libya, Mah, Mauntania, Morocco, Niger,
Senegal, Sudan, Tunisia, Wester Sahara
(U.a. geyni from the mountains of southem Algena and northern Niger is
frequently treated as a full species)
30
RDB Exp
R =
Wrote
SK
—- LA
(SK)
Ref
43,
Uromastyx aegyptius
Egyptian Spiny-tailed Lizard
Egypt, Iraq, Israel, Saudi Arabia, probably Jordan
Uromastyx asmussi
(Horn Lizard)
Afghanistan, Iran, Pakistan
Uromastyx benti
South Yemen
Uromastyx hardwickii
Hardwick's Spiny-tailed Lizard
Afghanistan, India, Pakistan
Uromastyx loricatus
Iran, Iraq
Uromastyx macfadyeni
Ethiopia, Somalia
Uromastyx microlepis
(Horn Lizard Small-grain)
Bahrain, Iran, Iraq, Kuwait, Oman, Saudi Arabia, South Yemen,
United Arab Emirates, probably also Qatar and North Yemen
Uromastyx ocellatus
Egypt, Sudan
Uromastyx ornatus
Egypt, Israel, Jordan, Saudi Arabia, Syna
Uromastyx philbyi
Saudi Arabia
Uromastyx princeps
Ethiopia, Somalia
Uromastyx thomasi
Oman
Family: CHAMAELEONIDAE
Chamaeleo adolfifriderici
Rwanda, Uganda, Zaire
CIT!
ES RDB
Exp
SK
SK
Ref
31
Chamaeleo affinis
Ethiopia, Somalia, probably Djibouti
Chamaeleo africanus
Afncan Chameleon
Cameroon, Chad, Egypt, Ethiopia, Mali, Niger, Nigenia,
Somalia, Sudan, probably Djibouti and Upper Volta
Chamaeleo anchietae
Angola, Tanzania, Zaire
Chamaeleo angeli
Madagascar
Chamaeleo antimena
Madagascar
(formerly inc. in C. rhinoceratus)
Chamaeleo balteatus
Two-banded Chameleon
Madagascar
(sometimes treated as a subspecies of C.bifidus)
Chamaeleo belalandaensis
Madagascar
Chamaeleo bifidus
Madagascar
Chamaeleo bitaeniatus*
Burundi, Ethiopia, Kenya, Rwanda, Somalia, Sudan, Tanzania,
Uganda, Zaire
Chamaeleo boettgeri
Madagascar
Chamaeleo brevicornis
Short-homed Chameleon
Madagascar
Chamaeleo calyptratus
South Yemen, North Yemen
Chamaeleo campani
Madagascar
CITES RDB
I -
Exp
Ref
*Recently (Bohme and Klaver) three taxa formerly treated as subspecies of C.bitaeniatus have been elevated to
species rank: C.ellioti, C.hoehnelii and C.kinetensis
32
Chamaeleo capuroni
Madagascar
Chamaeleo carpenteri
Uganda, Zaire
Chamaeleo cephalolepis
Comoro Islands
Chamaeleo chamaeleon
Common or Mediterranean Chameleon
Algena, Cyprus, Egypt, Gibraltar?, Greece (Crete only), India,
Israel, Italy (Sicily only), Jordan, Lebanon, Libya, Morocco,
Oman, Pakistan, Portugal, Saudi Arabia, South Yemen, Spain
(inc. Canary Islands), Sn Lanka, Syna, Tunisia, Turkey,
North Yemen
(populations in India, Pakistan and Sn Lanka are sometimes treated as a full
species, C.zeylanicus, and Arabian populations as C.arabicus)
Chamaeleo chapini
Zaire
Chamaeleo cristatus
Crested Chameleon
Cameroon, Equatonal Guinea, Gabon, Nigeria
Chamaeleo cucullatus
Madagascar
Chamaeleo deremensis
Tanzania (Usambara Mts)
Chamaeleo dilepis
Flap-necked Chameleon
Angola, Botswana, Burundi, Cameroon, Congo, Equatonial
Guinea, Ethiopia, Gabon, Kenya, Malawi, Mozambique, Namibia,
Nigena, Rwanda, Somalia, South Afnea, Tanzania, (inc. Pemba
Island and Zanzibar), Uganda, Zaire, Zambia, Zimbabwe
Chamaeleo eisentrauti
Cameroon
Chamaeleo fallax
Madagascar
Chamaeleo fischeri
Fischer's Chameleon
Kenya, Tanzania :
Chamaeleo fuelleborni
Tanzania
II
II
II
II
II
II
II
II
CITES RDB Exp
LA
Ref
38,110
33
Chamaeleo furcifer
Forked Chameleon
Madagascar
Chamaeleo gallus
Madagascar
Chamaeleo gastrotaenia
Madagascar
Chamaeleo globifer
Madagascar
Chamaeleo goetzei
Malawi, Tanzania, Zambia
Ghamaeleo gracilis
Graceful Chameleon
Angola, Benin, Cameroon, Central Afncan Republic, Chad,
Congo, Ethiopia, Equatonal Guinea, Gabon, Gambia, Ghana,
Guinea, Guinea Bissau, Ivory Coast, Kenya, Liberia, Mali, Nigena,
Senegal, Sierra Leone, Somalia, Sudan, Tanzania, Togo,
Uganda, Zaire, probably Djibouti
Chamaeleo guibei
Madagascar
Chamaeleo hoehnelii
Kenya, Uganda
Chamaeleo incornutus
Tanzania
Chamaeleo jacksonii
Jackson's Three-horned Chameleon
Kenya, Tanzania
Chamaeleo johnstoni
Bumnundi, Central Afncan Republic, Rwanda, Uganda, Zaire
Chamaeleo labordi
Madagascar
(sometimes treated as a subspecies of C. rhinoceratus)
Chamaeleo lateralis
Madagascar
Chamaeleo laterispinis
Tanzania
34
CITES
Il
I
II
II
RDB
Exp
Ref
Chamaeleo linotus
Madagascar
Chamaeleo malthe
Madagascar
Chamaeleo marshalli
Shorttailed Dwarf Chameleon
Mozambique, Zimbabwe
(alternatively regarded as a member of the Dwarf Chameleon genus
Rhampholeon)
Chamaeleo melleri
Meller's Chameleon
Malawi, Tanzania
Chamaeleo minor
Madagascar
Chamaeleo mlanjensis
Malawi
Chamaeleo monachus
Socotra Chameleon
Socotra (South Yemen)
Chamaeleo monoceras
Madagascar
Chamaeleo montium (inc. C.feae)
Mountain Chameleon
Cameroon, Equatonal Guinea, Nigena
Chamaeleo namaquensis
Desert Chameleon
Angola, Namibia, South Afnca
Chamaeleo nasutus
Madagascar
Chamaeleo oshaughnessyi
Madagascar
Chamaeleo oustaleti
Oustalet’s Giant Chameleon
Madagascar, (and introduced in Ngong Forest
near Nairobi, Kenya)
Chamaeleo oweni
Owen's Three-homed Chameleon
Cameroon, Congo, Equatonal Guinea, Gabon, Nigena, Zaire
CITES
RDB
Exp
Ref
35
CITES
Chamaeleo pardalis (inc. C.guentheri) II
Panther Chameleon
Madagascar, (introduced on Reunion)
Chamaeleo parsonii II
Parson's Giant Chameleon
Madagascar
Chamaeleo peyrieresi Il
Madagascar
Chamaeleo pfefferi II
Cameroon
Chamaeieo polleni Il
Comoro Islands
Chamaeleo pumilis II
Variegated Dwarf Chameleon
Mozambique, Namibia, South Africa
Chamaeleo quadricornis II
Four-horned Chameleon
Cameroon, Nigena
Chamaeleo quilensis II
Angola, Benin, Botswana, Cameroon, Congo, Equatonal Guinea,
Gabon, Kenya, Malawi, Mozambique, Namibia, Nigena,
South Africa, Tanzania (including Pemba and Zanzibar), Togo,
Uganda, Zaire, Zambia, Zimbabwe
Chamaeleo rhinoceratus II
Madagascar
Chamaeleo rudis* Il
Burundi, Kenya, Rwanda, Tanzania, Uganda
Chamaeleo senegalensis II
Senegal Chameleon
Benin, Burundi, Cameroon, Central African Republic, Ethiopia,
Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, Kenya,
Libena, Mali, Nigeria, Rwanda, Senegal, Sierra Leone, Sudan,
Tanzania, Togo, Uganda, Zaire, Zambia
Chamaeleo spinosus II
Spiny Chameleon
Tanzania
Chamaeleo tempeli II
Tanzania
*Recently (Bohme and Klaver) two taxa formerly treated as subspecies of
C.mdis have been elevated to species rank: C.schoutedeni and C.schubotzi.
36
RDB
Exp
Ref
CITES RDB Exp
Chamaeleo tenuis II =
Slender Chameleon
Tanzania
Chamaeleo tigris II -
Seychelle Islands
Chamaeleo tsaratananensis II =
Madagascar
Chamaeleo tuzetae II -
Madagascar
Chamaeleo verrucosus II =
Madagascar
Chamaeleo werneri II =
Tanzania
Chamaeleo wiedersheimi II =
Cameroon, Nigena
Chamaeleo willsii II =
Madagascar
Chamaeleo xenorhinus II =
Uganda, Zaire
Family: IGUANIDAE
Amblyrhynchus cristatus II R
Galapagos Marine Iguana
Galapagos Islands (Ecuador)
Anolis roosevelti - E
Giant Anole, Culebra Giant Ancle
Isla Culebra (Puerto Rico)
(possibly extinct)
Brachylophus fasciatus I E
Fiji Banded Iguana
Fiji Islands, Tonga Islands
Brachylophus vitiensis I =
Fiji Crested Iguana
Fyji Islands (Yaduataba Island)
Conolophus pallidus . II R
Barnington Land Iguana
Santa Fé (=Barnington) Island in Galapagos group (Ecuador)
CITES RDB Exp Ref
Conolophus subcristatus II View = 43,114
Galapagos Land Iguana
Galapagos Islands (Ecuador)
Cyclura carinata I Rie LAS 434129
Turks and Caicos Ground Iguana
Bahama Islands, Turks and Caicos Islands
Cyclura collei (C.lophoma) - possibly extinct I = = =
Jamaica's Ground Iguana
Jamaica
Cyclura cornuta (inc. C.stejnegeri) I V/K - 20,43
Rhinoceros Iguana
Dominican Republic, Haiti, Isla Mona, Navassa Id.
(in RDB C.c.stejnegeri from Isla Mona is in category ‘V', while C.c.cornuta from
the rest of the range is in Preamble 9—probably threatened)
Cyclura cychlura (inc. C.baelopha, C.figginsi, plus I Re anata 21043
C.inornata) F
Bahama Islands
Cyclura nubila (inc. C.caymanensis, C.macleayi) I WS 43
Cayman Islands, Cuba
Cyclura pinguis I Een 19,20
Anegada Ground Iguana 21,43
Anegada Id. (Bntish Virgin Islands)
Cyclura ricordi I K - 20,43
Hispaniola Ground Iguana, Ricord's Ground Iguana
Dominican Republic, Haiti
Cyclura rileyi (inc. C.cristata, C.nuchalis) I E/R F 43
Bahama Islands
(In RDB C.xz.rileyi is in category 'E’ the other two races are in 'R’)
Gambelia silus (Crotaphytus silus, C.wislizenii silus) - Ets 443,75
Blunt-nosed or San Joaquin Leopard Lizard
USA (California)
Iguana delicatissima II SP NE =
Lesser Antilles G@nc. Anguilla, Antigua, Ile Chevreau,
La Désiderade, Dominica, Ile Fourchue, Les Iles Frégates,
Guadeloupe, Martinique, Nevis, Les Iles des Saints,
St. Barthélemy, St. Eustatius, St. Martin)
Iguana iguana II - F 111,118
Common Iguana, Green Iguana, (South Amencan ‘'Chameleon’) LA
Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French ae
Guiana, Guatemala, Guyana, Honduras, Lesser Antilles
(inc. Grenada, the Grenadines, Guadaloupe, Montserrat, Saba,
Les Iles des Saints, St. Lucia, St. Vincent, Virgin Islands), Mexico,
Netherlands Antilles, Nicaragua, Panama, Paraguay, Perm,
El Salvador, Sunnam, Tnnidad & Tobago, Venezuela
38
CITES RDB Exp’ Ref
Phrynosoma coronatum blainvillei II Keo 4,43,75
San Diego Homed Lizard
Mexico (Baja California), USA (Califomia)
Sauromalus varius I Se In =
San Esteban Island Chuckwalla P
Mexico (San Esteban Island)
Family: LACERTIDAE
Podarcis filfolensis filfolensis (Lacerta f.filfolensis) = R - 38,43
Filfola Lizard
Filfola Island (Malta)
Gallotia simonyi (Lacerta simonyi) ~ I} = 43,108
Hierro Giant Lizard
Canary Islands (Hierro Island)
Family: CORDYLIDAE
Cordylus campbelli II AS =
Namibia
Cordylus cataphractus II = = =
South Afnca
Cordylus coerulopunctatus Il mes e
South Afnca
Cordylus cordylus Il <a gece x
Angola, Botswana, Ethiopia, Kenya, Lesotho?, Malawi,
Mozambique, Namibia, South Africa, Swaziland?,
Tanzania, Zaire, Zambia, Zimbabwe
Cordylus giganteus Il = apt ks a
Lesotho, South Afnca
Cordylus macropholis I Sah e: a
South Afnca
Cordylus namaquensis Il =. =
Namibia
Cordylus peersi Il = = =
South Afnca
Cordylus polyzonus Il aot qh if
Botswana, Lesotho?, Namibia, South Africa
Cordylus ukingensis II = = -
Tanzania 39
CITES RDB Exp Ref
Cordylus vittifer ll ie = =
Angola, Botswana, Lesotho?, Namibia?, South Africa,
Swaziland
Cordylus warreni II = ae -
Lesotho? Mozambique, South Afncea, Swaziland’,
Zimbabwe
Pseudocordylus capensis Il eee =.
South Afnca
Pseudocordylus langi II ya ar = te
Lesotho, South Africa
Pseudocordylus microlepidotus I te ves a
Namibia, South Afnca
Pseudocordylus robertsi Il Ly ee 3:
South Africa
Pseudocordylus spinosus Il i A aie =
South Africa
Family: TEIIDAE
Ameiva polops = E - 43,58,59
St. Croix Ground Lizard
St. Croix and adjacent cays (US Virgin Islands)
Cnemidophorus hyperythrus II Vv 4,43
Orange-throated Whiptail
Mexico (Baja California), USA (California)
Crocodilurus lacertinus II =" Sx ~
Brazil, French Guiana, Guyana, Sunnam
Dracaena guianensis II SHAN =
Caiman Lizard, (Croco-Teju)
Brazil, French Guiana, Peru, possibly Guyana and other
countnes bordering the Amazon basin
Tupinambis rufescens (inc. T.duseni and II SW =
T.teguixin sebastiani) SK
(Argentine Teju, Red Teju)
Argentina, Bolivia, Brazil, Paraguay, Uruguay
Tupinambis teguixin (Argentine or Brazil Teju) II - LA -
(inc. T.nigropunctatus)(Jacarura), SK
Common Tegu or Banded Tegu
Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana,
Paraguay, Peru, Surinam, Tnnidad & Tobago, Venezuela,
probably Ecuador
40
Family: SCINCIDAE
Gongylomorphus bojerii fontenayi
Macabé Forest Skink
Mauritius
Leiolopisma telfairii
Round Island Skink
Round Island (Mauritius)
Macroscincus coctaei — probably extinct
Cape Verde Giant Skink
Cape Verde Islands
Family: ANGUIDAE
Gerrhonotus panamintinus
Panamint Alligator Lizard
USA (California)
Family: ANNIELLIDAE
Anniella pulchra nigra
Black Legless Lizard
USA (California)
Family: HELODERMATIDAE
Heloderma horridum
Beaded Lizard
Guatemala, Mexico
Heloderma suspectum
Gila Monster
Mexico, USA
Family: VARANIDAE
Varanus acanthurus
Spiny-tailled Pygmy Monitor
Australia
Varanus bengalensis (V.monitor, inc. V.nebulosus)
Indian Monitor, Common Monitor (India)
(Agra, Bengal, Indian Small-grain, Penang, Rice,
Talagoya, - Lizard)
Afghanistan, Bangladesh, Burma, India, Indonesia, Iran,
Malaysia, Nepal, Pakistan, Sn Lanka, Thailand, Vietnam,
probably Kampuchea and Laos
Varanus brevicauda
Short-tailled Pygmy Monitor
Australia
Varanus caudolineatus
Streak-tailed Monitor
Australia
CITES
RDB Exp
R =
R 4
Id -
K be
is =
Id (LA)
V (LA)
- SK
(F)
Ref
43,77
12,43, 76,
77,78,83
38,43, 140
4,43
43,75
4]
CITES RDB. Exp
Varanus dumerilii II =
Dumeénl's Monitor,
(Fish Lizard)
Burma, Indonesia, Malaysia, Thailand
Varanus eremius II =
Desert Pygmy Monitor
Australia
Varanus exanthematicus II =
Afncan Savanna Monitor (African Large-grain Lizard)
Angola, Benin, Botswana, Burundi, Cameroon, Central African
Republic, Chad, Djibouti, Ethiopia, Gambia, Ghana, Guinea,
Guinea Bissau, Ivory Coast, Kenya, Lesotho, Malawi, Mali,
Maunitania, Mozambique, Namibia, Niger, Nigena, Rwanda,
Senegal, Sierra Leone, Somalia, South Africa, Sudan, Swaziland,
Tanzania, Togo, Uganda, Upper Volta, Zaire, Zambia, Zimbabwe
Varanus flavescens I =
Yellow Monitor (Calcutta or Indian Oval-grain Lizard)
Bangladesh, India, Nepal, Pakistan
Varanus giganteus II =
Perentie
Australia
Varanus gilleni II -
Pygmy Mulga Monitor
Australia
Varanus glauerti II -
Long-tailed Rock Monitor
Australia
Varanus glebopalma Il —
Australia
Varanus gouldii II =
Sand Monitor, Gould's Monitor
Australia, Indonesia (West Irian), Papua New Guinea
Varanus grayi II =
Gray's Monitor
Philippine Islands (Luzon)
Varanus griseus I Vv
Desert Monitor, Agra Monitor (India)
(Agra, Bengal, - Lizard, Baghdad Small-grain Lizard)
Afghanistan, Algeria, Chad, Egypt, India, Iran, Iraq, Israel,
Jordan, Kuwait, Lebanon, Libya, Mauritania, Morocco, Oman,
Pakistan, Qatar, Saudi Arabia, South Yemen, Sudan, Syria, Tunisia,
Turkey, United Arab Emirates, USSR, Western Sahara, North Yemen
(the RDB citation applies only to V.g.caspius, the Central Asian or Transcaspian
Desert (or Grey) Monitor, from Afghanistan, Iran, Pakistan (northern
Baluchistan), and USSR)
Varanus indicus Il =
Pacific Monitor, Mangrove Monitor, (Ambon or Flower Lizard)
Australia, Caroline Islands, Indonesia, Mariana Islands,
49 Marshall Islands, Papua New Guinea, Solomon Islands
SK
LA
LA
)
LA
SK
LA
Ref
47
43,81
Varanus karlschmidti
Sepik Monitor
Papua New Guinea
Varanus Komodoensis
Komodo Dragon
Indonesia (Flores, Gili Mota, Komodo, Owadi Sami, Padar, Rintja)
Varanus mertensi (in. V.bulliwallah)
Merten's Water Monitor
Australia
Varanus mitchelli
Mitchell's Water Monitor
Australia
Varanus niloticus
Nile Monitor, (Afncan Small-grain Lizard)
Angola, Benin, Botswana, Cameroon, Central Afncan Republic,
Chad, Congo, Djibouti, Egypt, Ethiopia, Equatorial Guinea,
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast,
Kenya, Lesotho, Libena, Malawi, Mali, Mauntania,
Mozambique, Namibia, Niger, Nigena, Senegal, Sierra Leone,
Somalia, South Afnca, Sudan, Swaziland, Tanzania, Togo,
Uganda, Upper Volta, Zaire, Zambia, Zimbabwe
Varanus prasinus
Emerald Monitor
Australia (Torres Straits Islands), Indonesia (West Inan),
Papua New Guinea
Varanus primordius
Ridge-tailed Monitor
Australia
(frequently treated as a subspecies of V.acanthurus)
Varanus rudicollis
Rough-necked Monitor
(Tree Lizard)
Burma, Indonesia, Malaysia, Thailand
Varanus salvadorii
Papuan Monitor, Tree Crocodile
Indonesia (West Iran), Papua New Guinea
Varanus salvator (in. V.bivittatus, V.togianus)
Malayan Monitor, Water Monitor
(very many names, in leather trade according to patterning and place of
origin, eg., No-mark, Plain, Rice, Ring, Borneo, Java, Philippines, Sumatra,
Semi-Java, Singapore, Siam, Ramgodie, Cabragoya - Lizard)
Bangladesh, Burma, China, India (inc. Andaman Is.), Indonesia,
Kampuchea, Laos, Malaysia, Philippine Islands, Singapore’,
Sn Lanka, Thailand, Vietnam
Varanus semiremex
Mangrove Monitor, Rusty Monitor
Australia
Varanus spenceri
Spencer's Monitor
Australia
CITES
II
II
II
I
I
II
II
I
RDB Exp
LA
)
LA
SK
Ref
43,66
47
43
CITES RDB Exp
Varanus storri II Se
Dwarf Monitor
Australia
Varanus timorensis II =v
Timor Tree Monitor, Spotted Tree Monitor
Australia, Indonesia, Papua New Guinea
Varanus tristis II =
Freckled Monitor, Mournful Tree Monitor
Australia
Varanus varius I =
Lace Monitor
Australia
Suborder: SERPENTES,
Snakes
Family: TYPHLOPIDAE
Typhlops monensis ST Wwe =
Mona Blind Snake
Isla Mona (Puerto Rico)
Family: BOIDAE
A few taxa of the family Boidae are listed on CITES Appendix I, the
remainder are on Appendix II. The actual content of this family has been the
subject of recent discussion among herpetologists. Both McDowell (1975, see
Ref list B) and Underwood (1976) remove the genera Bolyeria, Casarea,
Exiliboa, Trachyboa, Tropidophis and Ungaliophis from the family Boidae.
McDowell also removes Loxocemus, while Underwood retains Loxocemus
and adds Xenopeltis. Pending discussion, the contents of the Boidae given
here closely follows the more conservative list of Stimson (1969), this also
appears to be the usage of CITES and RDB. Some changes are made at low
taxonomic levels, following Sheplan & Schwartz (1974), McDowell (1975,
1979), and Collins et al (1978). Python oenpelliensis was newly described by
Gow (1977), P.saxuloides by Miller & Smith (1980), P.bredli by Gow (1981)
and P.carinatus by Smith (1981).
Acrantophis dumerili I Can Al
Duménl's Boa
Madagascar
Acrantophis madagascariensis I ? LA
Madagascar Boa
Madagascar
Aspidites melancophalus II oe
Black-headed Python
Australia
Aspidites ramsayi I = =
Woma
Australia
44
Ref
43
Boa constrictor (Constrictor constrictor)
Boa Constrictor,
(Ampalague, Giboya, Masacuate)
Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica,
Ecuador, French Guiana, Guatemala, Guyana, Honduras,
Lesser Antilles (Dominica, Santa Lucia), Mexico, Nicaragua,
Panama, Paraguay, Peru, El Salvador, Surinam, Tnnidad &
Tobago, Venezuela
Bolyeria multocarinata
Round Island Boa
Round Island (Mauntius)
Calabaria reinhardtii
Calabar Ground Python
Benin, Cameroon, Congo, Equatorial Guinea, Gabon, Ghana,
Ivory Coast, Libena, Nigena, Togo, Zaire, probably
Central Afncan Republic
Candoia aspera (Enygrus asper)
Papuan Ground Boa
Indonesia (Moluccas, West Inan), Papua New Guinea
(inc. Bismarck Archipelago as far east as Bougainville)
Candoia bibroni (Enygrus bibroni)
Pacific Boa
Fyji Islands (inc, Rotuma), Loyalty Islands, Vanuatu (= New
Hebrides), (inc. Banks Islands), Western Samoa, also Ta'u Island
in Amencan Samoa, Solomon Islands (incl. Santa Cruz)
Candoia carinata (Enygrus carinata)
Solomons Ground Boa
Indonesia (easter islands inc. Moluccas, Sulawesi, West Inan),
Palau Islands, Papua New Guinea (inc. Bismarck Archipelago),
Solomon Islands, (incl. Santa Cruz)
Casarea dussumieri
Round Island Keel-scaled Boa
Round Island (Mauritius)
Charina bottae
Rubber Boa
Canada, USA
(RDB entry is for Charina b. umbratica, the Southern Rubber Boa,
from isolated localities in California)
Chondropython viridis
Green Tree Python
Australia (Cape York Peninsula), Indonesia (West Inan),
Papua New Guinea (mainland, not Bismarck Archipelago),
Solomon Islands?
Corallus annulatus (Boa annulata)
Annulated Tree Boa
Colombia, Costa Rica, Ecuador, Nicaragua, Panama
Corallus caninus (Boa canina)
Emerald Tree Boa
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana,
Peru, Surinam, Venezuela
CITES
Il
II
II
II
II
II
I
RDB Exp
- LA
SK
E =
E =
R LA
- LA
- LA
Ref
12,43,
78,83
43,75
45
Corallus enydris (C.hortulanus, Boa hortulanus,
inc, Corallus or Boa cooki)
Garden Tree Boa
Bolivia, Brazil, Colombia, Costa Rica, Equador, French Guiana,
Guyana, Nicaragua, Peru, Surinam, Tnnidad & Tobago,
Venezuela, Windward Islands (Grenada, Grenadines,
St. Vincent)
Epicrates angulifer
Cuban Tree Boa
(Maja (de Santa Mania)), Cuba (inc. Isla de Pinos)
(RDB entry is for E.a.angulifer from the Isle of Pines and two small islands off
the coast of Cuba)
Epicrates cenchria
Rainbow Boa
(Araramboia)
Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador,
French Guiana, Guyana, Panama, Paraguay, Peru, Sunnam,
Tnnidad & Tobago, Venezuela
Epicrates chrysogaster (E.striatus chrysogaster)
Bahama Islands Boa
Bahama Islands, Caicos Islands, possibly Turks Islands
Epicrates exsul
Abaco Island Boa
Bahama Islands
Epicrates fordii (E.inornatus fordii)
Ford's Boa
Dominican Republic, Haiti
(see note under Epicrates inornatus)
Epicrates gracilis
Haitian Vine Boa
Dominican Republic, Haiti
Epicrates inornatus
Puerto Rican Boa
Puerto Rico
(E.inornatus, as used by CITES and RDB, appears to be in the sense of Stull
(1935) and Stimson (1969, Das Tierreich 89). In this usage three subspecies are
recognised, |. E.i.inornatus (Puerto Rico), 2. E.i.fordii (Dominican Republic
and Haiti), 3. E.i.granti (St. Thomas and Tortola in the Virgin Islands)
Good evidence has subsequently been proposed (Sheplan & Schwartz, 1974;
followed by Schwartz & Thomas, 1976) that E.i.fordii is best regarded asa
distinct species (E.fordii) and that E.i.granti has greater affinities with the
Epicrates of Isla Mona, the latter two taxa are accordingly treated as a distinct
species (E.monensis) in the works cited, The present list includes the West
Indian species of Epicrates as recognised by Sheplan & Schwartz (1974) and
Schwartz & Thomas (1976); the proper treatment of E.fordii and E.monensis
by CITES and RDB thus remains to be determined. it is assumed that the
current CITES and RDB treatment will continue to apply to Epicrates
inornatus (of Sheplan & Schwartz)
Epicrates monensis (E.gracilis monensis plus
E.inornatus granti)
Isla Mona (Puerto Rico), St. Thomas (US Virgin Islands),
Tortola (British Virgin Islands)
(see note under Epicrates inornatus (RDB entry is for E.m.monensis
from the Isla Mona)
46
CITES
II
II
I
RDB Exp
Ide (SK)
- LA
(SK)
EB =
R =
Ref
43
43
43
Epicrates striatus
Fischer's Tree Boa
Bahama Islands, Dominican Republic, Haiti
(RDB entry is for E.s.fosteri from the Bahamas)
Epicrates subflavus
Jamaica Boa
Jamaica
Eryx colubrinus
Kenya Sand Boa
Egypt, Ethiopia, Kenya, Niger, Somalia, Sudan,
Tanzania, North Yemen?
Eryx conicus (Gongylophis conicus)
Indian Sand Boa
(Baby ‘Python’)
India, Pakistan, Sn Lanka
Eryx elegans
Afghanistan, Iran, USSR (Turkmenistan)
Eryx jaculus
(Javelin) Sand Boa, Spotied Sand Boa
Albania, Algeria, Bulgania, Egypt, Greece, Iran, Iraq, Israel,
Jordan, Lebanon, Morocco, Romania (possibly extinct),
Saudi Arabia, Syma, Tunisia, Turkey, USSR, Yugoslavia
Eryx jayakari
Jayakar’'s Sand Boa
Bahrain, Kuwait, Oman, Qatar, Saudi Arabia, South Yemen,
United Arab Emirates, North Yemen
Eryx johnii
Blunttailed Sand Boa
Afghanistan, India, Iran, Pakistan
Eryx miliaris
Iran, USSR
Eryx muelleri
Muller's Sand Boa
Cameroon, Chad, Ghana, Ivory Coast, Mali, Mauntania,
Nigeria, Senegal, Sudan, Togo, Upper Volta, probably Benin
and Central Afnican Republic
Eryx somalicus
Somalia
Eryx tataricus
Tartary Sand Boa
Afghanistan, China, Iran, Mongolia, Pakistan, USSR
Eunectes barbouri
Brazil
CITES
RDB Exp
R
LA
LA
LA
SK
LA
LA
Ref
43
43
148
38
47
Eunectes deschauenseei
Brazil
Eunectes murinus
Anaconda
Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana,
Peru, Surinam, Trinidad, Venezuela
Eunectes notaeus
Yellow Anaconda
Argentina, Bolivia, Brazil, Paraguay, Uruguay
Exiliboa placata
Mexico
Liasis albertisii (L.fuscus albertisii)
D’albertis' Python
Indonesia (West Inan), Papua New Guinea, Torres Straits Islands
(Australia)
Liasis boa (Bothrochilus boa, Nardoana boa)
Barred Python
Papua New Guinea (Bismarck Archipelago, mainland
New Guinea’), Tokelau Islands?
Liasis childreni
Children's (Rock) Python
Australia
Liasis mackloti (inc. L.fuscus)
Water Python (common name for L.fuscus)
Australia, Indonesia (Lesser Sunda Islands), Papua New Guinea
Liasis maximus
(known only from the type specimen, from New Guinea; a possible synonym of
L.papuanus (McDowell, 1975 Ref, list B)
Liasis olivaceus
Olive (Rock) Python
Australia
Liasis papuanus (L.olivaceus papuanus)
Papuan Python
Indonesia (West Irian), Paonua New Guinea
Liasis perthensis
West Australian Rock Python
Australia
(taxonomic status uncertain, (Cogger, 1975, Ref. list B))
Lichanura trivirgata
Rosy Boa
Mexico, USA (Anzona, California)
(L.t.roseofusca from Mexico and California is often regarded asa full species,
Lichanura roseofusca)
48
CITES
I
I
I
Il
II
II
RDB
SK
SK
LA
75
Loxocemus bicolor
Costa Rica, Guatemala, Honduras, Mexico, Nicaragua,
El Salvador
Python amethistinus (Liasis amethistinus)
Amethystine (Rock) Python
Australia (Cape York Peninsula, Torres Straits Is.), Indonesia
(easter islands, West Inan), Papua New Guinea, Philippines?
Python anchietae
Angolan Python
Angola, Namibia
Python boeleni (Liasis boeleni, Liasis taronga)
Boelen’s Python
Indonesia (West Inan), Papua New Guinea
Python bredli
Australia (Northern Ternitory)
Python carinatus
Rough-scaled Carpet Snake
Australia (Westem Australia)
Python curtus
Short-tailed Python, Blood Python
Brunei, Indonesia (Bangka, Kalimantan, Sumatra), Malaysia,
Thailand (in Malay Peninsula)
Python molurus sub-spp.
Indian Python, Burmese Python, Onental Rock Python
Java Tree Python, Molunis Python, Indian Python, Bengal or
Saigon Python)
P.m.bivittatus
Brunei?, Burma, China, Hong Kong, Indonesia, Kampuchea,
Laos, Malaysia, Thailand, Singapore, Vietnam
P.m.molurus
Bangladesh, India, Nepal, Pakistan, Sn Lanka
Python oenpelliensis
Australia
Python regius
Ball Python, Royal Python
Benin, Cameroon, Chad, Gambia, Ghana, Guinea, Guinea
Bissau, Ivory Coast, Mali, Nigeria, Senegal, Sierra Leone, Sudan,
Togo, Uganda, Upper Volta, Zaire, probably also Central Afncan
Republic, Congo, Equatorial Guinea, Gabon, Libena
CITES
II
RDB Exp
SK
Ref
49
CITES RDB Exp Ref
Python reticulatus II = A
Reticulated Python, (Diamond or Java Rock Python) SK
Bangladesh, Brunei, Burma, Indonesia (not to West Inan),
Kampuchea, Laos, Malaysia, Nicobar Islands (India), Philippines,
Singapore, Thailand, Vietnam
Python saxuloides Il = — Ibs -
Lesser African Rock Python
Kenya, probably Malawi, South Afnca, possibly Ethiopia,
Somalia, Uganda
Python sebae II = hiv et
African Python, Water Python, African Rock Python SK
(West African Python)
Angola, Benin, Botswana, Burundi, Cameroon, Central Afncan
Republic, Chad, Congo, Djibouti, Equatorial Guinea, Ethiopia,
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast,
Kenya, Lesotho, Liberia, Malawi, Mali, Mauritania, Mozambique,
Namibia, Nigeria, Rwanda, Senegal, Sierra Leone, Somalia,
South Africa, Sudan, Swaziland, Tanzania, Togo, Uganda,
Upper Volta, Zimbabwe, Zaire, Zambia
Python spilotus (Morelia spilotes, Morelia argus) II = ik -
P.s.variegata—Carpet Python, P.s.spilotus—Diamond Python
Australia, Indonesia (West Irian), Papua New Guinea
Python timorensis II == =
Timor Python
Indonesia (Flores, Timor)
Sanzinia madagascariensis I K LA -
Madagascar Tree Boa
Madagascar
Trachyboa boulengeri Il =a le =
Colombia, Ecuador, Panama
Trachyboa gularis II ate ie
Ecuador
Tropidophis battersbyi Il a ams 2
Ecuador
Tropidophis canus Il = 3
Bahama Islands
Tropidophis caymanensis Il = =
Caymans Ground Boa
Cayman Islands
Tropidophis feicki Il ee S
Cuba
50
CITES RDB
Tropidophis greenwayi II =
Caicos Islands
Tropidophis haetianus II =
Haitian Ground Boa
Cuba, Dominican Republic, Haiti, Jamaica
Tropidophis maculatus II -
Cuba
Tropidophus melanurus II ~
Cuba, Navassa Island
Tropidophis nigriventris II -
Cuba
Tropidophis pardalis II -
Cuba
Tropidophis paucisquamis II -
Brazil, Peru
Tropidophis pilsbryi II -
Cuba
Tropidophis semicinctus II -
Cuba
Tropidophis taczanowskyi Il =
Ecuador, Peru, Brazil?
Tropidophis wrighti II -
Cuba
Ungaliophis continentalis II -
Guatemala, Honduras, Mexico
Ungaliophis panamensis Il -
Colombia, Costa Rica, Nicaragua, Panama
Xenoboa cropani II -
Brazil
CITES RDB Exp Ref
Family: COLUBRIDAE
Clelia clelia (Pseudoboa cloelia) II = aA =
Musurana
Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador,
French Guiana, Guatemala, Guyana, Honduras, Lesser Antilles
(inc. Dominica, Grenada, St. Lucia), Nicaragua, Panama,
Paraguay, Peru, El Salvador, Surinam, Trinidad, Uruguay,
Venezuela
(snakes from Mexico and elsewhere in Central America formerly assigned to
this species are now usually regarded as a full species, Clelia scytalina)
Drymarchon corais couperi = Vie Ave 435133
Eastern Indigo Snake
USA
Elachistodon westermanni II = = 35
Indian Egg-eating Snake
Bangladesh, India, Nepal
Hydrodynastes gigas (Cyclagras gigas) II = lst -
South Amencan Water ‘Cobra’, SK
(Beach ‘Cobra’, Surucucu)
Argentina, Bolivia, Brazil, Paraguay
Masticophis flagellum ruddocki = R = 4,43
San Joaquin Coachwhip
USA (California)
Masticophis lateralis euryxanthus - R - 4,43
Almeda Striped Racer
USA (California)
Nerodia fasciata taeniata (Natrix f.taeniata) - R = 4,43
Atlantic Saltmarsh Snake
USA (Florida)
Nerodia sipedon insularum (Natrix s.insularum) = Ke =
Lake Ene Water Snake
Lake Ene (Canada/USA)
Thamnophis couchi gigas (T.elegans gigas) = R=) = 4,43
Giant Garter Snake as
USA (California)
Thamnophis couchi hammondi II Re LAY 4:48
(T. elegans hammondi) 75
Two-striped Garter Snake
Mexico (Baja California), USA (California)
Thamnophis sirtalis tetrataenia = E LA 443
San Francisco Garter Snake 1
USA (California)
52
CITES RDB Exp
Family: ELAPIDAE
Naja oxiana - En)
(frequently treated as a subspecies of the Asiatic Cobra, Naja naja)
Central Asian or Oxus Cobra
Afghanistan, India Kashmir, Punjab), Iran (northeast), Pakistan,
(north and northwest), USSR (Transcaspian region)
Ogmodon vitianus = Wo =
Fy Snake
Fuji (Viti Levu)
Family: VIPERIDAE
Bothrops insularis = K =
Queimada Island Bothrops
Brazil (Queimada Island)
Crotalus unicolor = ILA
Aruba Island Rattlesnake
Netherlands antilles (Amba Island)
Crotalus willardi - E/V LA
Ridge-nosed Rattlesnake
Mexico, USA (Anzona, New Mexico)
(in RDB C.w.obscumus, the New Mexico Ridge-nose Rattlesnake, is
Endangered, the four other subspecies are Vulnerable)
Vipera ammodytes transcaucasiana = R LA
Transcaucasian Long-nosed Viper
USSR (Georgia), Turkey
Vipera latifii = Ee
Latifi's Viper
Iran (Lar Valley in Elburz Mtns.)
Vipera lebetina schweizeri = Wo =
Blunt-nosed Viper, Levantine or Lebetine Viper
(names applied to V.lebetina as a whole)
Greece (Cyclades only: Kimolos, Milos, Polyagos, Siphnos,
possibly Antimilos, Kithnos)
Vipera xanthina raddei = R LA
Armenian Viper
USSR, Turkey
Ref
43,81
43
43
428,36,
43,151
43,81
43,97
43,81
53
Additions and Amendments
CITES RDB Exp Ref
Family: KINOSTERNIDAE
Kinosternon flavescens spooneri - i -= — 1g
Illinois Mud Turtle
USA
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(A) Sources of information on the survival status of particular species or groups of species.
fe OD YH
The number of each Reference corresponds to entries in the ‘Ref column of the taxonomic list. The utility of the
References given is rather variable; some works concentrate entirely on survival status and conservation, while
others may only make a brief commenton these topics in the course ofalargely taxonomic treatment. Some of the
worksare popular orsemi-popular, while others are more rigorous inapproach. Emphasisisgiven tomore recent
works, including those not cited in the third edition of the Red Data Book; the latter (item 43 in the list) contains
references to many other works. It is not the intention to provide a complete bibliography of the extensive
literature treating the biology and conservation of amphibians and reptiles.
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DODD, C.K. 1979. A bibliography of endangered and threatened amphibians and reptiles in the US. and its
territories (conservation, distnibution, natural history, status). Smithsonian Herpetological Information Service,
No. 46:1-35. (Washington, D.C.)
FUCHS, K. 1974. Die Krokodilhaut. Darmstadt: Eduard Roether Verlag
HOOGMOED, M.S. 1977. De handel in reptielehuiden in Singapore. Lacerta 35(5):66-74. (inc. English summary),
INSKIPP, T., & WELLS, S. 1979. International trade in Wildlife. Earthscan (10 Percy St., London W1P ODR).
KING, F.W., & BRAZAITIS, P. 1971. Species identification of commercial crocodile skins Zoologica (NY)
56(2): 15-70.
MACK, D., DUPLAIX, N., & WELLS, S. 1979. International Trade in Sea Turtle Products (The Sea Turtle: an animal
of divisible parts). Traffic (International) & Traffic (USA). Revised edition, 1980
MROSOVSKY, N. (Ed.) Marine Turtle Newsletter. (Depts. of Zoology & Psychology, Univ. of Toronto, Ontario,
Canada MSS 1 Al).
WERMUTH, H., & FUCHS, K. 1978. Bestimmen von Krokodilen und ihrer Haute. Stuttgart: Gustav Fischer.
WOOD, D.A. 1977. A bibliography of the World's rare, endangered, and recently extinct wildlife and plants.
Environmental Ser. No.3. Environmental Inst., Oklahoma State Univ
WORLD CONFERENCE ON SEA TURTLE CONSERVATION. Nov. 1979, Washington D.C.—a symposium volume
on the Conference is anticipated in 1981. Among reports on the Conference is: CHERFAS, J. 1979. The song of the
turtle. New Scientist 84 (1185) 880-882:
61
Index
Genera, common, and trade names.
Abaco Island Boa 46
Abbott's Day Gecko 28
Achoque 11
(Acinixys) 20
Acrantophis 44
Adanson's Turtle 23
Afghan Tortoise 20
African Chameleon 32
African Large-grain Lizard 42
African Python 50.
African Rock Python 50
African Savanna Monitor 42
African Sharp-nosed Crocodile 26
African Slender-snouted Crocodile 26
African Small-grain Lizard 43
Agra Lizard 40, 41
Agra Monitor 42
(Agrionemys) |9
Alabama Red-bellied Turtle 17
Aldabra Giant Tortoise 18
Alligator 25
Almeda Striped Racer 52
Armargosa Toad 14
Amblyrhynchus 37
Ambon Lizard 42
Ambystoma ||
Ameiva 40
Amencan Alligator 25
American Crocodile 26
Amethystine (Rock) Python 49
Ampalague 43
Anaconda 48
Andaman Day Gecko 29
Andrias | |
Anegada Ground Iguana 38
Angolan Python 49
Angonoka 18
Angulated Tortoise 17
Aniella 4]
Annulated Tree Boa 45
Anolis 37
Aquatic Box Turtle 17
Araramboia 45
Archey's Frog 13
Areolated Tortoise 19
Argentine Teju 40 (two entnes)
Argentine Tortoise 17
Armenian Viper 53
Arrau 24
Aruba Island Rattlesnake 53
Asiatic Cobra 51
Aspidites 44
(Asterochelys) |8
Atelopus | 4
Atlantic Ridley 21
Atlantic Saltmarsh Snake 52
Australian Fresh-water Crocodile 26
Axolotl 11
Baby Python 47
Baghdad Small-grain Lizard 41
Bahama Islands Boa 46
Ball Python 49
Banded Tegu 40
62
Barred Python 48
Barrington Land Iguana 37
Batagur |6
Batrachophrynus |3
Batrachoseps | 1, 12
Baw-baw Frog 13
Beach Cobra 52
Beaded Lizard 41
Bell's Hinged Tortoise 19
Bengal Lizard 40, 41
Bengal Python 49
Berlandier's Tortoise 18
Black Caiman 25
Black-headed Python 44
Black Legless Lizard 41
Black Pond Turtle 16
Black Side-necked Turtle 23
Black Soft-shell Turtle 22
Black Spiny-tailed Lizard 30
Black Toad 14
Blood Python 49
Blunt-nosed Leopard Lizard 38
Blunt-nosed Viper 53
Blunttailed Sand Boa 47
Boa 44
(Boa) 45,46
Boa Constnictor 44
Boelen's Python 49
Bog Turtle 16
Bolson Tortoise 19
Bolyeria 45
Borneo Lizard 42
@othrochilus) 48
Bothrops 53
Boulenger's Tortoise 19
Bowsprit Tortoise 17
Brazil Teju 39
Broad-nosed Caiman 25
Broad-snouted Crocodile 26
Brazilian Giant Tortoise 17
Brachylophus 37
Bufo 14
Burmese Brown Tortoise 18
Burmese Python 49
Burmese Starred Tortoise 18
Burmese Swamp Turtle 17
Cabragoya 42
Caiman 25
Caiman Lizard 40
Calabaria 45
Calabar Ground Python 45
Calcutta Lizard 41
California Tiger Salamander 11
Callagur |6
Cameroon Toad 14
Candoia 45
Cape Clawed Toad 13
Cape Platanna 13
Cape Verde Giant Skink 41
Caretta 21
Carettochelys 22
Carpet Python 50
Casarea 45
Cat Island Turtle 17
Caymans Ground Boa 50
Central American River Turtle 16
Central Asian Cobra 53
Central Asian Monitor 41
Chaco Sideneck Turtle 24
Chaco Tortoise 17
Chamaeleo 31-37
Charina 45
Cheat Mountain Salamandar 12
Chelonia 21
Chersina 17
Childrens (Rock) Python 48
Chinese Alligator 25
Chinese Giant Salamander 11
Chondropython 45
Chioglossa | |
(Chrysemys) 17
Clelia 51]
Clemmys |6
Cnemidophorus 40
Common Batagur 16
Common Chameleon 33
Common Hinged Tortoise 19
Common Iguana 38
Common Monitor 41
Common Tegu 40
Common Tortoise 19
Conolophus 37,38
Conraua 15
(Constrictor) 43
Corallus 45,46
Cordylus 39, 40
Coromandel Leiopelma 13
Creaser's Mud Turtle 16
Crested Chameleon 33
Crested Lizard 30
Crocodilurus 40
Crocodylus 26,27
Croco-Teju 40
Crotalus 53
(Crotaphytus) 38
Cuatro Cienegas Soft-shell Turtle 22
Cuban Tree Boa 46
Cuban Crocodile 27
Culebra Giant Anole 37
Cuvier's Smooth-fronted Caiman 25
(Cyclagras) 50
Cyclura 38
Cyrtodactylus 28
D'Albert's Python 48
Dahl's Toad-headed Turtle 24
(Damonia) 16
Dark Soft-shell Turtle 22
Day Geckos 28-30
Deep Spmings Toad 14
Dermatemys 16
Dermochelys 22
Desert Chameleon 35
Desert Monitor 42
Desert Pygmy Monitor 42
Desert Slender Salamander 11
Desert Tortoise 19
Diamond Python 49, 50
Discoglossus |3
Dracaena 40
Drymarchon 52
Dumeril's Boa 44
Dumeril's Monitor 42
Dunn's Mud Turtle 16
Dura Turtle 16
Dwarf Caiman 25
Dwarf Monitor 43
Eastern Indigo Snake 52
Egyptian Spiny-tailed Lizard 31
Egyptian Tortoise 20
Elachistodon 50
Eleutherodactylus | 3
Emerald Monitor 43
Emerald Tree Boa 45
(Enygrus) 45
Epicrates 46,47
Eretmochelys 21
Erymnochelys 23
Eryx 47
Estuarine Crocodile 26
Eunectes 47,48
Eurycea |2
(Eurycea) |2
Exiliboa 48
False Gavial 27
False Gharial 27
Fiji Banded Iguana 37
Fyi Crested Iguana 37
Fyi Snake 53
Filfola Lizard 39
Fischer's Chameleon 33
Fischer's Tree Boa 47
Fish Lizard 42
Flap-necked Chameleon 33
Flatback Turtle 21
Flattened Musk Turtle 16
Florida Gopher Tortoise 19
Flower Lizard 42
Ford's Boa 46
Forest Hinged Tortoise 19
Forest Tortoise 17
Forked Chameleon 34
Four-horned Chameleon 36
Freckled Monitor 44
Furrowed Wood Turtle 17
Gaboon Turtle 23
Galapagos Giant Tortoise 18
Galapagos Land Iguana 38
Galapagos Marine Iguana 37
Gallotia 39
Gambelia 38
Ganges Soft-shell Turtle 22
Garden Tree Boa 46
Gardiner's Seychelles Frog 15
Gavialis 27
Gavial 27
Geoclemys 16
Geochelone 17, 18
(Geoemyda) 16
Geometric Tortoise 20
Gerrhonotus 41
Ghanial 27
Giant Anole 37
Giant Garter Snake 52
Giboya 43
(Gigantorana) 15
Gila Monster 41
Golden Coqui Frog 13
Golden Frog 14
Golden Toad 14
Gold-dust Day Gecko 29
Gold-striped Salamander 11
Goliath Frog 15
Gongylomorphus 41
(Gongylophis) 47
Gopherus 19
Gould's Monitor 42
Graceful Chameleon 34
Gray's Monitor 42
Greek Tortoise 20 (two entries)
Green Iguana 38
Green Tree Python 45
Green Turtle 21
Grey Monitor 41
Grooved Tortoise 18
Grotto Salamander 12
(Gymnodactylus) 28
Haitian Ground Boa 51
Haitian Vine Boa 46
Hamilton's Frog 13
Hardwick's Spiny-tailed Lizard 31
Hawksbill Turtle 21
Helmeted Turtle 23
Heloderma 41
Heosemys |6
Hercules Tortoise 17
Hermann's Tortoise 20
Hierro Giant Lizard 39
Hispaniola Ground Iguana 38
Hochstetter's Frog 13
Hoge's Sideneck Turtle 24
Home's Hinged Tortoise 19
Homopus |9
Horn Lizard 31
Horn Lizard Small-grain 31
Horsfield's Tortoise 20
Houston Toad 14
Hydrodynastes 52
Hydromantes | 2
Hydrosaurus 30
Hyla 15
Iguana 38
Illinois Chorus Frog 15
Illinois Mud Turtle 54
Impressed Tortoise 18
Inagua Island Turtle 17
Indian Egg-eating Snake 52
Indian Flap-shell Turtle 22
Indian Monitor 41
Indian Oval-grain Lizard 42
Indian Python 49 (two entries)
Indian Sand Boa 46
Indian Sawback Turtle 16
Indian Small-grain Lizard 40
Indian Soft-shell Turtle 22
Indian Star or Starred Tortoise 18
Indian Tent Turtle 16
Irapuca 23
Island Night Lizard 30
Israel Painted Frog 13
Italian Spadefoot Toad 13
Jaboti 17
Jacaré 24
Jacarura 39
Jackson's Three-hommed Chameleon 34
Jamaica Boa 47
Jamaica's Ground Iguana 38
Japanese Giant Salamander ] 1
Java Lizard 42
Java Rock Python 50
Java Tree Python 49
Javelin 47
Jayakar's Sand Boa 47
Jemez Mountains Salamandar 12
Johnson's Crocodile 26
Kachuga |6
Karoo Tortoise 19
Kavalai Forest Turtle 16
Kemp's Ridley 21
Kenya Sand Boa 46
Kem Canyon Slender Salamander 11
Kinixys 18, 19
Kinosternon 16,54
(Klauberina) 31
Komodo Dragon 43
Kuhl's Tortoise 19
Lace Monitor 44
(Lacerta) 39
Lake Ene Water Snake 52
Lake Junin (Giant) Frog 13
Lake Lerma Salamander 11
Lake Patzcuaro Salamander 11
Larch Mountain Salamander 12
Latifi's Viper 53
Leatherback 22
Leathery Turtle 22
Lebetine Viper 53
Leiopelma 13
Leiolopisma 4]
Leopard Tortoise 18
Lepidochelys 21
Lesser African Rock Python 50
Levantine Viper 53
Liasis 48
(Liasis) 48,49
Lichanura 48
Limestone Salamander 12
Lissemys 22
Loggerhead Turtle 21
Long-tailed Rock Monitor 42
Loxocemus 49
63
Luth 22
Macabe Forest Skink 41
Macroscincus 4]
Madagascar Boa 44
Madagascar Day Gecko 29
Madagascar Flat-shelled Tortoise 20
Madagascar Tortoise 18
Madagascar Tree Boa 50
Magdalena River Turtle 24
Maja 45
Malachochersus 19
Malayan Monitor 43
Mangrove Monitor 42, 43
Margined Tortoise 20
Marsh Crocodile 26
Masacuate 43
Masticophis 52
Mastigures 30
Mauritius Greater Day Gecko 28
Mediterranean Chameleon 33
Megalixalus |5
(Megalsbatrachus) | |
Melanochelys 16
Melanosuchus 25
Meller's Chameleon 35
Merten's Water Monitor 43
Mexican Giant Gopher Tortoise 19
Mexican Spotted Wood Turtle 17
Mindoro Crocodile 26
Mitchell's Water Monitor 43
Molurus Python 49
Mona Blind Snake 44
(Morelia) 49
Morelet's Crocodile 26
Morenia 17
Mountain Chameleon 35
Mountain Tortoise 18
Mount Nimba Viviparous Toad 14
Mournful Tree Monitor 44
Mugger 26
Muhlenberg's Turtle 16
Mulga Monitor 16
Miiller's Sand Boa 47
Musurana 52
Naja 53
(Nardoana) 48
Narrow-bridged Mud Turtle 16
(Natrix) 52
Nectophrynoides | 4, 15
Nerodia 52
Nesomantis 15
New Guinea Crocodile 26
New Guinea Plateless Turtle 22
New Mexico Ridge-nosed
Rattlesnake 51
(Nicoria) 16
Nile Crocodile 26
Nile Monitor 43
Nile Soft-shell Turtle 22
No-mark Lizard 42
North Island Leiopelma 13
64
Oedura 28
Ogmodon 53
Olive Ridley 21
Olive (Rock) Python 48
Olm 12
Orange-throated Whiptail 40
Orange Toad 14
Oniental Rock Python 49
Oninoco Crocodile 26
Osteolaemus 27
Oustalet's Giant Chameleon 35
Owen's Three-horned Chameleon 35
Oxus Cobra 53
Pacific Boa 45
Pacific Monitor 42
Pacific Ridley 21
Painted Terrapin 16
Paleosuchus 25
Palm Lizard 30
Panamint Alligator Lizard 41
Pancake Tortoise 19
Papuan Ground Boa 45
Panther Chameleon 36
Papuan Monitor 43
Papuan Python 48
Paradelma 30
Parrot-beaked Tortoise 19
Parson's Giant Chameleon 36
Peacock-marked Softshell Turtle 22
Pelobates |3
Pelomedusa 23
Peltocephalus 23
Pelusios 23
Penang Lizard 40
Perentie 42
Phaeognathus | 2
Phelsuma 28-30
Philippines Crocodile 26
Phihppine Lizard 42
Philoria |3
Phrynops 24
Phrynosoma 39
Pigmy Mulga Monitor 31
Pine Barrens Tree Frog 15
Platypus Frog 14
Plain Lizard 42
Platemys 24
Plethodon |2
Plymouth Red-bellied Turtle 17
Podarcis 39
Podocnemis 23, 24
(Podocnemis) 23
Proteus |2
Psammobates 20
Pseudacris 15
Pseudemydura 24
Pseudemys |6, 17
(Pseudoboa) 51
Pseudocordylus 40
Puerto Rican Boa 46
Pygmy Mulga Monitor 42
Python 49-50
Pyxis 20
Queensland Snake-Lizard 30
Queimada Island Bothrops 53
Radiated Tortoise 18
Rainbow Boa 46
Ramgodie 42
Rana 15
(Rana) 15
Red-foot Tortoise 17
Red-headed Amazon Turtle 23
Red-headed Sideneck Turtle 24
Red Hills Salamander 12
Red Teju 40
Relict Leopard Frog 15
Reticulated Python 50
Reticulated Velvet Gecko 28
(Rhampholeon) 35
Rheobatrachus | 4
Rhinoceros Iguana 38
Rhinoclemmys |7
(Rhoptropella) 29
Rice Lizard 40, 42
Ricord's Ground Iguana 38
Ridge-nosed Rattlesnake 53
Ridge-tailed Monitor 43
Ring Lizard 42
Rio Apaporis Caiman 25
River Terrapin 16
Rodriguez Day Gecko 30
Roofed Turtle 16
Rosy Boa 48
Rough-necked Monitor 43
Rough-scaled Carpet Snake 49
Round Island Boa 45
Round Island Day Gecko 29
Round Island Keel-scaled Boa 45
Round Island Skink 41
Royal Python 49
Rubber Boa 45
Rusty Monitor 43
Saigon Python 49
Sail-fin Lizard 30
Salt-water Crocodile 26
Sand Boa 47
Sand Monitor 42
San Diego Horned Lizard 39
San Esteban Island Chuckwalla 39
San Francisco Garter Snake 52
San Joaquin Leopard Lizard 38
San Joaquin Coachwhip 52
San Marcos Salamander 12
Santa Cruz Long-toed Salamander 11
Sanzinia 49
Sauromalus 39
Schneider's Smooth-fronted Caiman 25
Schweigger's Hinged Tortoise 19
Semi-Java Lizard 42
Senegal Chameleon 36
Sepik Monitor 43
Serpent Island Gecko 28
Serrated Tortoise 20
Seychelle Islands Tree Frog 15
Seychelles Frog 15
Shasta Salamander 12
Short-horned Chameleon 32
Short-necked Swamp Turtle 24
Short-tailed Dwarf Chameleon 35
Short-+tailed Pygmy Monitor 41
Shorttailed Python 49
Siam Lizard 42
Siamese Crocodile 27
Side-stnped Day Gecko 29
Singapore Lizard 42
Slender Chameleon 37
Slender-Snouted Crocodile 26
Soa-Soa Water Lizard 30
Socotra Chameleon 35
Solomons Ground Boa 45
Sonoran Green Toad 14
Sooglossus 15
South American ‘Chameleon’ 38
South American Red-lined Turtle 17
South American River Turtle 24
South American Water ‘Cobra’ 52
Speckled Tortoise 19
Spectacled Caiman 25 (two entries)
Spencer's Monitor 43
Sphenodon 27
Spider Tortoise 20
Spiny Chameleon 36
Spiny-tailed Lizard 30
Spiny-tailed Pygmy Monitor 41
Spix's Sideneck Turtle 24
Spotted Pond Turtle 16
Spotted Sand Boa 47
Spotted Tree Monitor 44
Spurred Tortoise 18
Spur-thighed Tortoise 20
St. Croix Ground Lizard 40
Stephen Island Leiopelma 13
Sternotherus |6
Steak-tailed Monitor 41
Stnpe-backed Side-necked Turtle 23
Sulawesi (Celebes) Tortoise 18
Sumatra Lizard 42
Sururucu 52
Talagoya 40
Tartary Sand Boa 47
Tegu 39
Tehachapi Slender Salamander 12
Tent Tortoise 20
Teracay 24
Terrapene |7
Testudo 20
(Testudo) 17,18, 20
Texas Blind Salamander 12
Texas Gopher Tortoise 18
Thamnophis 52
Thomasset's Seychelles Frog 15
Three-keeled Land Tortoise 16
Timor Python 50
Timor Tree Monitor 43
Tomistoma 27
Toothed Cape Tortoise 20
Tracaja 24
Trachyboa 50
Transcapian Desert Monitor 41
Transcaucasian Long-nosed Viper 53
Travancore Tortoise 18
Tree ‘Crocodile’ 43
Tree Lizard 42
Trionyx 22
Tropidophis 50, 51
Tuatara 27
Tuntong 16
Turks & Caicos Ground Iguana 38
Tupinambis 40
Two-banded Chameleon 32
Two-stnped Garter Snake 52
Typhlomolge | 2
Typhlops 44
Typhlotriton
Ungaliophis 5!
Uromastyx 30, 31
Varanus 41-44
Vanegated Dwarf Chameleon 36
Vegas Valley Leopard Frog 15
Vipera 51,53
Viviparous African Toad 14
Water Box Turtle 17
Water Monitor 43
Water Python 48
West African Dwarf Crocodile 27
West African Python 50
West Australian Rock Python 48
Western Gopher Tortoise 19
Western Swamp Turtle 24
White-breasted Side-necked Turtle 23
Woma 44
Xantusia 30
Xenoboa 5)
Xenopus |3
Yellow Anaconda 48
Yellow-foot Tortoise 17
Yellow Monitor 42
Yellow-spotted Amazon Turtle 24
Yellow Tortoise 18
Zetek's Golden Frog 14
65
Lithographed by The Nuffield Press Limited, Cowley, Oxford, England.
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