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Full text of "World Checklists of Threatened Amphibians and Reptiles"

NATURE 

CONSERVANCY 

COUNCIL 



World Checklist of Threatened 
Amphibians and Reptiles 




Nature Conservancy Council Great Britain 



U)CM-G- klczpLl^ "83 



Digitized by tine Internet Arciiive 

in 2010 with funding from 

UNEP-WCIVIC, Cambridge 



littp://www.arcliive.org/details/worldcliecklistso83groo 



World Checklist of Threatened 
Amphibians and Reptiles 



Bnan Groombndge 



Wildlife Advisory Branch 
Nature Conservancy Council 19/20 Belgrave Square London 3W1X SPY United Kingdom 



Contents 







Page 






Page 


Introdnctory Notes 




7 


Pelomedusidae 


Erymnochelys 

Pelomedusa 

Peltocephalus 


23 
23 
23 


Taxonomic List 








Pelusios 


23 


AMPHIBU 








Podocnemis 


23 


Cryptobranchidae 


Andrias 


11 


Chelidae 


Phiynops 


24 


Ambystomabdae 


Ambystoma 


U 




Platemys 


24 


Salamandndae 


Chioglossa 


11 




Pseudemyduia 


24 


PlethodonQdae 


Batrachoseps 


11 


Alligatoridae 


Alligator 


25 




Eurycea 


12 




Cckiman 


25 




Hydromantes 


12 




Melanosuchiis 


25 




Phaeognathus 


12 




Paleosuchus 


25 




Plethodon 


12 


Crocodylidae 


CitXHxlylus 


26 




Typhlomolge 


12 




Osteolaemus 


27 




TypMotiiton 


12 




Tomistoma 


27 


Proteidae 


Proteus 


12 


Gavialidae 


Gavialis 


27 


Leiopelmatidae 


Leiopelma 


13 


Sphenodontidae 


Sphenodon 


27 


Pipidae 


Xenopus 


13 


Gekkonidae 


Cyrtodactylus 


28 


Discoglossidae 


Discoglossus 


13 




Oedura 


28 


Pelobaddae 


Pelobates 


13 




Phelsuma 


28 


Leptodactyhdae 


Batrachophrymus 


13 


Pygopodidae 


Paradelma 


30 




Eleutherodactylus 


13 


Xantusudae 


Xantusia 


30 


Myobatrachidae 


Phjloiia 


13 


Agamidae 


Hydrosaunis 


30 




Rheobatrachus 


14 




Urotnastyx 


30 


Bufonidae 


Atelopus 


14 


Chamaeleomdae 


Chamaeleo 


31 




Bofo 


14 


Iguamdae 


Amblyrhynchus 


37 




Nectophrynoides 


14 




Anolis 


37 


Hylidae 


Hyla 


15 




Brachylophus 


37 




Pseudacris 


15 




Conolophus 


37 


Sooglossidae 


Nesomantis 


15 




Cyclura 


38 




Sooglossus 


15 






38 


Ranidae 


Conzaua 


15 




Iguana 


38 




Rana 


15 




Phrynosoma 


39 


Hyperoliidae 


Megalixalus 


15 




Sauromalus 


39 








Lacertidae 


Podarcis 


39 


REPTII.TA 








GaUotia 


39 


Kinostemidae 


Kinostemon 


16 


Cordylidae 


Cordylus 


39 




Stemotherus 


16 




Pseudocordylus 


40 


Dermatemydidae 


Denuatemys 


16 


Teudae 


Ameiva 


40 


Emydidae 


Batagur 


16 




Cnemidophorus 


40 




Callagur 


16 




Crocodilurus 


40 




Clemmys 


16 




Dracaena 


40 




Geoclemys 


16 




Tupinambis 


40 




Heosemys 


16 


Scincidae 


Gongylomoiphus 


41 




Kachuga 


16 




Leiolopisma 


41 




Melanochelys 


16 




Macroscincus 


41 




Morenia 


17 


Anguidae 


Gerrhonotus 


41 




Pseudemys 


17 


Aniellidae 


Aniella 


41 




Rhinoclemmys 


17 


Helodermatidae 


Heloderma 


41 




Teiiapene 


17 


Varanidae 


Varanus 


41 


Testudinidae 


Cheisina 


17 


Typhlopidae 


Typhlops 


44 




Geochelone 


17 


Boidae 


Acrantophis 


44 




Gopherus 


19 




Aspidites 


44 




Homopus 


19 




Boa 


45 




Kinixys 


19 




Bolyeiia 


45 




Malacochersus 


19 




Calabaria 


45 




Pscuiunobates 


20 




Candoia 


45 




Pyxis 


20 




Casarea 


43 




Testudo 


20 




Charina 


45 


Chelonudae 


Caretta 


21 . 




Chondropython 


45 




Chelonia 


21 




Corallus 


45 




Eietmochelys 


21 




Epicrates 


46 




Lepidochelys 


21 




Eryx 


47 


Dermochelydidae 


Dermochelys 


22 




Eunectes 


47 


Carettochelyidae 


Carettochelys 


22 




Exiliboa 


48 


Trionychidae 


Lissemys 


22 




Liasis 


48 




Trionyx 


22 




Lichanura 


48 



Acknowledgements 







Page 


Boidae cont. 


Loxocemus 


49 




Python 


49 




Sanzinia 


50 




Trachyboa 


50 




Tropidophis 


50 




Ungaliophis 


51 




Xenoboa 


61 


Colubridae 


Clelia 


52 




Drymarchon 


52 




Elachistodon 


52 




Hydrodynastes 


52 




Masticophis 


52 




Nerodia 


52 




Thamnophis 


52 


Elapidae 


Naja 


53 




Ogmodon 


53 


Viperidae 


Bothrops 


53 




Crotalus 


53 




Vipera 


53 


Additions and amendments 


54 


References 




SS 


Index (to genera, 


synonyms, common. 


62 


and trade names) 







This checklist was compiled and revised in 
October 1982 for the 3rd edition by Dr B 
Groombridge (now at the lUCN Species 
Conservation Monitoring Unit, Cambridge) at the 
Amphibian and Reptile Sechon of the British 
Museum (Natural History), under contract 
NCC/E2/36/04 from the Nature Conservancy 
Council of the United Kingdom, 

The compiler is most grateful to the staff of the 
Amphibian and Reptile Section at the British 
Museum (Natural History), m particular to Miss A G 
C Grandison and Dr E N Arnold, for their valuable 
assistance. 

1st edition published 1979 as a typed document 
with limited circulation 
ISBN 86139 075 X 

2nd edition published 1981 

Designed by Creative Department, Manchester 

Ltd, 

Printed for HMSO by Robendene Ltd. 

ISBN 86139 095 4 

3rd edition published 1983 

Revised design by Margaret Morgan BA 



ISBN 86 139 224 8 

© Nature Conservancy Council 1979, 1981, 1983 



Preface 



This checklist was originally prepared to aid the 
United Kingdom's Scientific and Management 
Authorities for the Convention on International 
Trade in Endangered Species of Wild Fauna and 
Flora (CITES) which the United Kingdom has 
implemented since the 1 January 1976, 

The aim was to provide a summary of basic data 
on each taxon relating to the countries where it 
occurs in the wild, its category in the lUCN Red 
Data Book, its hstmg in CITES and important 
sources of reference. Particular regard has been 
given to listing general names, synonyms. English 
vernacular names and trade names This 
information will help to clarify conditions on the 
issue of permits to trade in amphibians and reptiles, 
especially in those cases where ambiguous or 
erroneous supporting data are supphed It is hoped 
that this publication will be of value to all 
Management and Scientihc Authorities of the Party 
States to CITES and to all those concerned with the 
wider studies of herpetology and related matters. 

This checklist covers only those amphibians and 
reptiles which are themselves threatened in some 
degree, or treated as 'look-alikes' under CITES, at 
the time of compilation. It is clearly desirable for the 
work to be extended to include the remaining taxa 
and therefore complete a full coverage of the 
group 

The present document was completed in 
January 1979 and since then has been subject to 
partial revision to take account of decisions 
affecting the Append ices at the 2nd meeting of the 
Parties to CITES at San Jose, Costa Rica m February 
1979, the 3rd meeting at New Delhi in 1981 and new 
taxa included in Part 1 of the lUCN Amphibia- 
Reptilia Red Data Book (1982), 

The United Kingdom Scientific Authority and the 
Nature Conservancy Council are grateful to Dr 
Brian Groombridge for undertaking this work on 
their behalf, 

AJBRUDGE 

Wildlife Advisory Branch 

Nature Conservancy Council 

Belgrave Square 

London SW IX SPY 

Wmter 1983 



Introductory notes 



The purpose of this work is to provide a 
combined hst of the amphibian and reptile species 
(or subspecies) appearing on Appendices I. II, and 
III. of the Convention on International Trade m 
Endangered Species (CITES), and those included 
in the Red Data Book (RDB), The latter includes data 
sheets from the second edition issued in 1975, 
together with many new or revised sheets dated 
1978 or 1979 (but not generally available until 1980), 
A useful introduction to CITES is provided by 
Inskipp & Wells (1979. see Reference list C), 

This revised edition incorporates changes made 
in the CITES Appendices at the New Delhi meeting 
of the Parties in 1 98 1 , and changes and additions 
among Testudmes, Crocodylia and 
Rhynchocephalia made in Part 1 of the revised 
Amphibia-Reptilia Red Data Book (ref. 1 S2), New 
and fully revised RDB accounts for remaining 
reptiles and for amphibians are currently in 
preparation, the taxa among these groups noted in 
this checklist are thus not fully representative. 

For each taxon. the scientific name is given first, 
with the currently most frequently used synonyms 
in brackets. Common names (English language), 
and trade names (in brackets), appear on the 
following line Very few of the listed taxa have 
regularly or universally used common names. In 
several cases, identical trade names appear to be 
used for different taxa. 

The Red Data Book taxa included are those 
listed in Preamble 8 (i.e , those for which full data 
sheets are provided) as well as those in Preamble 9 
(that are suspected of being threatened, but for 
which full data are currently lacking). 

Geographical Range 

The geographical range of each taxon is given in 
terms of political units, listed in alphabetical order. 
The exceptions are some of those taxa occuring on 
small and/or remote islands with well-known 
names, that are administered by another country; 
for most such taxa the administenng country is 
given in brackets, e.g. Galapagos Islands 
(Ecuador). In certain cases, when a taxon is found m 
a restricted area within a larger political unit, the 
name of that area is given in brackets, eg.. 
Indonesia (West Irian), 

The political names given are those in common 
usage, and may not in all cases be the 
diplomatically correct version. 

When the name of a country is given as within 
the range of a certain species, that species may not 
occur throughout the country, and may even be of 
only peripheral occurence. 

Without surveying the range of each taxon m the 
field. It IS necessary to rely on published records. 
Most taxonomic works give the range of a taxon in 
terms of broad geographical areas, rather than of 
precise political units. While some countries have a 
relatively well-known and well-reported 
herpetofauna, this is not true for others. These 
factors may occasionally have resulted m some of 
the geographical ranges given here being 
incomplete or innaccurate, although every effort 
has been made to counter this. 



Taxonomic sources. 

The most recent available checklist or revision 
has usually been followed for the taxonomic 
treatment of each species or group of species 
herein Certain groups are clearly overdue for 
species-level revision. In the interests of stability, 
the present list follows the taxonomic arrangement 
of the RDB for the forms appearing therein, unless 
there is good reason to do otherwise. The genera 
and species are in alphabetical order within each 
family. The scientific and common names of North 
American species have been modified to accord 
with Standard Common and Current Scientific 
Names for North American Amphibians and 
Reptiles (Collins, Huheey, Knight, & Smith, 1978) 

The following list states the primary taxonomic 
source followed for most of the groups dealt with 
herein, and refers to an authority for each 
modification from that primary source AH works 
cited in this secton appear in part (B) of the 
Reference list. 

Gaudata and Anura: follows Gorham (1974), except: 

(a) Ambystoma tigrinum califomiense here 
treated as a full species. A. califomiense, after 
Collins at aid 978) 

(b) Batrachoseps aridus is added, described by 
Brame(1970) 

(c) Family Atelopodidae no longer recognised, 
Atelopus here assigned to family Bufonidae, after 
McDairmid(1969). 

(d) Bufo nelsoni here reduced to a subspecies, 
B. boreas nelsoni, after Collins et al (1978) 

(e) Five species of Nectophrynoides are added, 
N.cryptus and N.minutus. described by Perret 
(1971, 1972); also N.osgoodi (transferred from Bufo) 
and N.malcomi described by Grandison (1978) and 
N.liberiensis described by Xavier (1 978). 

(f) Sooglossidae here elevated to family rank 
(Nesomantis and Sooglossus listed), after Lynch 
(1973) and Duellman (1975) 

(g) Rana goliath here listed as Conraua goliath, 
after Lamotte & Perret (1 968). 

(h) Rana pipiens lisheri now in synonymy with 
Rana onca, after Pace (1974). 

(i) Hyperoliidae elevated to family rank 
(Megalixalus listed), after Liem (1970). 

0) Myobatrachidae elevated to family rank 
(Philoria and Rheobatrachus listed) after Lynch (in 
Vial) 1973, Heyer & Liem (1976), and others. 

Testudtnes, Crocodylia, Rhynchocephalia: follows 
Wermuth&Mertens (1977), except 

(a) Geoemyda tricarinata here listed as 
Melanochelys tricarirata, after McDowell (1964). 

(b) Pseudemys terrapen malonei is treated as a 
full species, P.malonei, as in Schwartz & Thomas 

(1 975) and Pntchard ( 1 979). The species is assigned 
to the genus Pseudemys rather than Chrysemys, 
following Wermuth & Mertens (1977) and Pntchard 
(1979); there is considerable debate about the use 
of these names, 

(c) Testudo IS broken up into four additional 
genera (Acinixys, Chersina, Geochelone, 
Psammobates) and Testudo is restricted to five 
Palearctic species, following Loveridge & Williams 



0957) and the majority of subsequent authors. 

(d) Podocnemis dumeriliana is listed as 
Peltocephalus dumeriliana (apparently a synonym 
of Peltocephalus tracaxa, unpublished work noted 
in Pritchard ( 1 979) and Rhodm et al) after Rhodin et 
aid 978) 

(e) Podocnemis madagascariensis is treated as 
Erymnochelys madagascariensis, after Williams 
(1954), Smith & James (1958), and Rhodin etal(1978) 

The above modifications within Testudines are 
also all incorporated m Pntchard (1979). (See Bour 
( 1 978) and Obst ( 1 980) for different views on the 
nomenclature of certain of the endemic Malagasy 
tortoises). 

(0 Crocodylus mindorensis here treated as a 
subspecies, C. novaeguinea mindorensis, after 
Wermuth & Fuchs (1978) and several earlier 
authors. 

Squamata; Gekkonidae and Pygopodidae: follows 
Wermuth (1965), except, 

(a) Gymnodactylus serpensinsula here treated 
as Cyrtodactylus serpensinsula, after Underwood 
(1954) and subsequent authors 

(b) Phelsuma agalegae added, described by 
Cheke(1975) 

(c) Phelsuma dubia comorensis elevated to a full 
species P. comorensis, after Mertens (1966a) 

(d) Phelsuma newtoni changed to P. 
edwardnewtoni, following Vinson & Vinson (1969) 

(e) Phelsuma modesta added, described by 
Mertens (1970) 

(0 Rhoptropella ocellata is Created as a species 
of Phelsuma, P. ocellata, after Russel (1977) 

(g) Phelsuma vinsoni changed to P. omata, see 
Mertens (1970) 

Squamata; Agamidae: follows Wermuth ( 1967). 

Squamata; Chamaeleontidae: follows Mertens 
(1966b), except, 

(a) Chamaeleo angeli added, described by 
Brygoo & Domergue ( 1 968). 

(b) Chamaeleo antimena here treated as a full 
species, after Brygoo & Domergue (1968) 

(c) Chamaeleo balteatus here treated as a full 
species, after Brygoo & Domergue (1968). 

(d) Chamaeleo belalandaensis added, 
described by Brygoo & Domergue (1970), 

(e) Chamaeleo capuroni added, described by 
Brygoo, Blanc, & Domergue (1972) 

(f) Chamaeleo eisentiauti added, described by 
Mertens (1968) 

(g) Chamaeleo guentheri here synomised with 
C.pardalis, after Brygoo (1969). 

(h) Chamaeleo labordi treated as a full species, 
after Brygoo & Domergue (1968). 

(i) Chamaeleo peyrieresi added, described by 
Brygoo, Blanc & Domergue (1974). 

(j) Chamaeleo tsaratananensis added, 
described by Brygoo & Domergue (1967/1968). 

(k) Chamaeleo tuzetae added, described by 
Brygoo, Bourgat & Domergue (1972). 

0) Also see Bohme and Klaver for further recent 
changes. 



Squamata; IgueUiidae; several sources, notably: 

(1) Montanucci. (1970), (Crotaphytus wislizenii 
silus should be regarded as a full species C. silus). 

(2) Collins etal, (1978), (Crotaphytus siius 
assigned to Gambelia), 

(3) Carey, (1975), Schwartz & Thomas, (1975), 
Schwartz and Carey, (1977), (Cyclura). 

(4) Peters & Donoso-Barros, (1970), Schwartz & 
Thomas, (1975), aguana), 

(5) Gibbons, (1981), (new Brachylophus sp) 

Squamata; Lacertidae: generic assignment after 

Arnold, (1973) 

Squamata; Cordylidae: after Wermuth, (1968). 

Squamata; Teiidae: several sources, notably: 

(1) Peters & Donoso Barros, (1970) 

(2) Vanzolini & Valencia, (1 965), (Dracaena). 

(3) Presch, (1973), (Tupinambis, only two species 
recognised). 

Squamata; Helodermatidae, Varanidae: follows 
Mertens (1963), except; 

(a) Varanus acanthurus primordius regarded as 
a full species after Cogger (1975), 

(b) Varanus storri added, described by Mertens 
(1966c) 

Squamata; Boidae: several sources, see notes on 
p44 and p46 

Squamata; Colubridae; several sources, notably: 

(1) Peters & Orejas-Mu-anda, (1970), (Clelia 
clelia). 

(2) Cans & Williams, (1954), Fleming and 
Flemmg, (1973), 433 (Elachistodon). 

(3) Dowhng & Gibson, (1970); (Cyclagras gigas 
assigned to Hydrodynastesj. 

(4)Rossman & Eberle, (1977); (Natrix sipedon 
and N. fasciata assigned to Nerodia), 

Squamata; Elapidae, Viperidae: chiefly after 
Klemmer, (1963). 




For each taxon, there are at least two entnes in the 
four columns down the right-hand side of the page 
CITES: 1,11, or III, referring to the CITES Appendix on 

which the taxon is listed as at June 1981 , 
RDB: Red Data Book status categones, 

E = Endangered 

V = Vulnerable 

R = Rare 

0= Out of danger 

Id = Indeterminate (given as "I" in RDB, but 

modified here to avoid confusion with the 

CITES "I" entry). 

K = Insufficiently Known: suspected of being 

threatened, but without full data. 
Exp: Major exploitation of taxon, where known. 

F = Food (may include local and commercial use) 

LA = Live animal trade (primarily for the pet 

trade) 

Pr = Preserved specimens 

SK = Skins, for the leather trade 

TS = Tortoise-shell 

When known, a category of secondary 

importance is given in brackets. 
Ret Entry here isanumberreferringtoanitemin part 

(A) of the list of References providing incidental or 

exhaustive information on the survival status of the 

taxon. 



10 



Class: AMPHIBIA 



CITES RDB Exp Ref 



Order: CAUDATA (Urodela), 
Newts and Salamanders 

FamHy: CRYPTOBRANCHIDAE 

Andrias davidianiis (Megalobatrachus davidianus) 

Chinese Giant Salamander 

China 

Andrias japonicus (Megalobatrachiis japonicus) 

Japanese Giant Salamander 
Japan 

FamHy: AMBYSTOMATIDAE 

Ambystoma califomiense (A. tigrinum calif omiense) 

California Tiger Salamander 
USA (California) 



Id F 43 
LA 



R F 43 
LA 



V - 4, 43 



Ambystoma dumerilii 



II R F 43 



Mexico 

(RDB entry is for A.d. dumerilii, the Lake Patzcuaro Salamander or Achoque, 
from Lake Patzcaiaro, Michoacan; the remaining subspecies A.d. 
queretarensis, Michoacan, is not included) 

Ambystoma lermaensis 

Lake Lerma Salamander 
Mexico 



II R F 43 



Ambystoma macrodactylum croceum 

Santa Cruz Long-toed Salamander 
USA (California) 



E - 1, 4, 14 
43, 67, 75 



Ambystoma mexicaniim 

Axolotl 

Mexico (Lake Xochimilco region) 

Family: SALAMANDRIDAE 

Chioglossa lusitanica 

Gold-stnped Salamander 
Portugal, Spain 

Family: PLETHODONTIDAE 

Batrachoseps aridus 

Desert Slender Salamander 
USA (California) 
Batrachoseps simatus 

Kern Canyon Slender Salamander 
USA (California) 



II R LA 43 



V - 16, 38, 43 



E - 4 43, 75 



R - 4, 43, 75 



11 



CITES RDB Exp Ref 

Batrachosepsstebbinsi - R 4,43,75 

Tehachapi Slender Salamander 

USA (California) 

Euryceanana - R -4,43 

San Marcos Salamander 

USA (Texas) 

Hydromantesbrunus - R - 4,43,75 

Limestone Salamander 
USA (California) 

Hydromantes shastae - R 4 43. 75 

Shasta Salamander 

USA(Califomia) 

Phaeognathus hubrichti - R - 43 

Red Hills Salamander 
USA (Alabama) 

Plethodonlarselli - Id - 4,43 

Larch Mountain Salamander 

USA (Oregon, Washington) 

Plethodonneomexicanus - v - 4,43,73 

Jemez Mountams Salamander 

USA (New Mexico) 

Plethodon netting! (P. richmondi netting!) - K - - 

Cheat Mountam Salamander 

USA (West Virginia) 

Typhlomolgerathbuni(Eiirycearathbuni) - E -4,43,51 

Texas Blind Salamander 

USA (Texas) 

Typhlotriton spelaeus - K - - 

Grotto Salamander 
USA 

Family: PROTEIDAE 

Proteus anguinus - V LA 38,43 

Olm 

Yugoslavia (and introduced at Vicenza, Italy) 



12 



CITES RDB Exp Ref 

Order: ANURA (Salientia) 
Frogs and Toads 

Family: LEIOPELMATIDAE 
(LEIOPELMIDAE; Leiopelma is sometimes 
included m the ASCAPHIDAE) 

Leiopelma axcheyi - R - 11,43,64 

IOC irw 

Archey's Frog, CxDromandel Leiopelma 

New Zealand 

Leiopelma hamiltoni - R - 11,26, 

Hamilton's Frog, Stephen Island Leiopelma j^g j^,^ 

New Zealand 

Leiopelma hochstetteri - R - 1M3 64 

Hochstetter's Frog, North Island Leiopelma 

New Zealand 



106,107 



Family; PIPIDAE 

Xenopusgilli - V - 43 47 

Cape Platanna, Cape Clawed Toad 
South Afnca (Cape Tbwn region) 

FamHy: DISCOGLOSSIDAE 

Discoglossus nigriventer - E - 43 

Israel Painted Frog 
Israel (Lake Huleh) 

FamHy: PELOBATIDAE 

Pelobates fuscns insubricus - E - 38,43 

Italian Spadefoot Toad 

Italy (plains of River Po). Switzerland (Ticmo, probably extinct) 

Family: LEPTODACTYLIDAE 

Batrachophrynusmacrostomus - K - - 

Lake Junin (Giant) Frog 

Peru (Lake Junin) 

Eleutherodactylusjasperi - R - 43 

Golden Coqui Frog 
Puerto Rico 

Family: MYOBATRACHIDAE 

(alternatively regarded as a subfamily of LEPTODACTYLIDAE) 

Philoriafrosti - R - 43 

Baw-baw Frog 

Australia 



13 



CITES RDB Exp Ref 

Rheobatrachus silus - v - 43 

Platypus Frog 
Australia (Queensland) 

Family: BUFONIDAE 

Atelopusvariiiszeteki I - LA 50 

Golden Frog, Zetek's Golden Frog ^^^ 

Panama (Valle de Anton) 

(Atelopus IS sometimes assigned to a separate family ATELOPODIDAE) 

Biifo boreas nelsoni - id - 4,43 

Amargosa Tbad 

USA (California, Nevada) 

Bufoexsul - V LA 4 43,75 

Black Toad, Deep Springs Toad 

USA (California) 

(sometimes treated as a subspecies of B. boreas, B.boreas exsul) 

Bufo houstonensis - E - 4,8,43,79 

Houston Toad 

USA (Texas) 

Bufoperiglenes I E - 43 

Golden Toad, Orange Toad 

Costa Rica 

Bufo retif oimis II v LA 4,43 

Sonoran Green Toad 

Mexico (Sonora), USA (Arizona) 

Bufosuperciliaris I - LA - 

Cameroon Toad 

Cameroon, Equatorial Guinea, Gabon, Ghana, Guinea, Nigeria, 
Zaire, probably also in other countnes of west and west-central 
Africa 

Nectophrynoides cryptiis I - _ _ 

Viviparous African Toad 
Tanzania (Uluguru Mts.) 

Nectophrynoides liberiensis I - _ _ 

Liberia (Mt^ Nimba) 

Nectophrynoides malcomi I - _ _ 

Ethiopia 

Nectophrynoides minutus I - _ _ 

Tanzania (Uluguru Mts.) 

Nectophrynoides occidentalis I v - 43 

Mt. Nimba Viviparous Toad 

Gumea (Mt. Nimba), and adjacent parts of Ivory Coast 



14 



CITES RDB Exp Ref 
Nectophrynoides Osgood! I _ _ _ 

Ethiopia 

Nectophrynoides tomieri I _ _ _ 

Ibnzania (Magrotto and Usambara Mts,) 

Nectophrynoides viviparus I _ _ _ 

Tknzania (Kinga, Rungwe, and Uluguru Mts.) 

Family: HYLIDAE 

Hyla andersoni - R LA 4,43, 

Pine Barrens Tree Frog ^^' 1^3 

USA 

(Flonda populations noted as Endangered m RDB) 

PseudacrisstreckeriiUinoensis - Id - 4,43 

Illinois Chorus Frog 

USA 

Family: SOOGLOSSIDAE 
(sometimes treated as a subfamily of 
RANIDAE) 

Nesomantis thomasseti - id - 43 

Thomasset's Seychelles Frog 

Seychelle Is. (Mahe, Silhouette) 

Sooglossus sechellensis - Id - 43 

Seychelles Frog 

Seychelle Is. (Mahe, Silhouette) 

Sooglossus gardineri - id - 43 

Gardiner's Seychelles Frog 
Seychelle Is. (Mahe, Silhouette) 

Family: RANIDAE 

Conraua goliath (Rana goliath, Gigantorana goliath) - v F 43 

Goliath Frog ^^ 

Cameroon, Equatorial Guinea 

Raziaonca(inc Ranapipiensfisheri) - E -4,43 

Relict Leopard Frog (mc. Vegas Valley Leopard Frog) 

USA (Arizona, Utah, Vegas Valley Leopard Frog restricted to Nevada) 

Family: HYPEROLIIDAE 
(sometimes treated as a subfamily of 
RHACOPHORIDAE) 

Megalixalusseychelleiisis(M.infrarufiis) - K - 43 

Seychelle Islands Tree Frog 

Seychelle Islands (Mahe, PrasUn, La Digue, Silhouette) 

15 



ClassREPTILIA 



CITES RDB Exp Ref 

Order: TESTUDINES, 
Tortoises, Turtles 
Suborder: CRYPTODIRA 

FamHy: KINOSTERNIDAE 

Kinostexnon augustipons _ r - 152 

Narrow-bndged Mud Turtle 

Costa Rica, Nicaragua, Panama 

Kinostexnon creaseri _ r _ 152 

Greaser's Mud Turtle 

Mexico 

Kinostemon dunni _ r _ 152 

Dunn's Mud Turtle 

Columbia 

Stexnotherus minor depressus _ id _ 152 

Flattened Musk Turtle 

USA 

Family: DERMATEMYDIDAE 

Dermatemys mawii n v p 9,152 

Central Amencan River Turtle 
Belize, Guatemala, Mexico 

Family: EMYDIDAE 

Batagurbaska I e f 54,136 

Common Batagur, River Terrapin, Tuntong 

Bangladesh, Burma, India, Indonesia, Kampuchea 

Malaysia, Thailand, Vietnam? 

Callagur borneoensis _ v F 152 

Painted Terrapin 

Indonesia (Kalimantan, Sumatra), Malaysia, Thailand 

Clenunys muhlenbexgii n id (LA) 2,4,25 31 

Bog Turtle , Muhlenberg ' s Turtle 1 1 3 , 1 52 

USA 

Geoclemys hainiltonii (Damonia hamiltonii) I _ la - 

Black Pond Turtle, Spotted Pond Turtle 

Bangladesh, India, Pakistan 

Heosemys silvatica _ id - 1 52 

Kavalai Forest Turtle 

India (Kerala) 

Kachuga tecta tecta I _ _ _ 

Indian Sawback or Tent Turtle, Roofed or Dura Turtle 

Bangladesh, India, Pakistan 

Melanochelys tricarinata (Geoemyda txicarinata, 

Nicoria tricaxinata) I _ _ _ 

Three-keeled Land Tortoise 

Bangladesh, India 

16 



CITES RDB Exp Ref 

Morenia ocellata I _ _ _ 

Burmese Swamp Turtle 

Burma 

Pseudemys alabamensis - id - 1S2 

Alabama Red-bellied Turtle 

USA 

Pseudemys fells - r - 152 

Cat Island Turtle 
Bahamas (Cat Island) 

Pseudemys malonei (Pseudemys terrapen malonei, 

Chrysemys malonei) - r - 152 

Inagua Island Turtle 

Bahamas (Great Inagua Island) 

Pseudemys rubriventris bangs! - r - 152 

Plymouth Red-bellied Turtle 
USA 

Pseudemys scripta callirostris (Pseudemys omata 2 V F 152 

callirostris, Chrysemys omata callirostris) la 

South American Red-lined Turtle 

Colombia, Venezuela 

Rhlnoclemmys areolata - K - 152 

Furrowed Wood Turtle 

Belize, Guatemala, Honduras, Mexico 

Rhlnoclemmys rubida - k - 152 

Mexican Spotted Wood Turtle 

Mexico 

Terrapene coahuila I v LA 9,152 

Aquatic or Water Box Turtle 
Mexico (Coahuila) 

Family: TESTUDINIDAE 

Cheisina angulata (Testudo angulata) ll - LA - 

Angulated or Bowsprit Tortoise 
Namibia, South Afnca 

Geochelone carbonaria (Testudo carbonaria) ll K F 152 

Red-foot Tortoise LA 

Brazil, Colombia, French Guiana, Guyana, 

Lesser Antilles, (several islands, probably introduced), 

Panama, Paraguay, Surinam, Tnnidad, Venezuela 

Geochelone chilensis (inc G.donosobarrosi and II I 

G.petersi; Testudo chilensis) 

Argentine Tortoise, Chaco Tortoise 

Argentina, Paraguay, probably Bolivia 

Geochelone denticulata (Testudo denticulata) II 

Yellow-foot Tortoise, Jaboti, Forest Tortoise, 

Brazilian Giant Tortoise, Hercules Tortoise 

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, 

Peru, Surinam, Trinidad, Venezuela, possible Paraguay 



F 


34,104, 


LA 


152 


F 




LA 





17 



CITES RDB Exp Ref 

Geochelone elegans (Testudo elegans) II - LA - 

Indian Starred or Star Tortoise 

India, Pakistan, Sn Lanka 

Geochelone elephantopus (Testudo elephantopus) I E - 43,114 

Galapagos Giant Tortoise 1^2 

Galapagos Islands (Ecuador) 

Geochelone elongata (Testudo elongata) II - F - 

Yellow Tortoise ^^ 

Bangladesh, Burma, China, India, Kampuchea, Laos, Malaysia, 

Nepal, Thailand, Vietnam 

Geochelone emys (Testudo emys) II K F 54,152 

Burmese Brown Tortoise 

Burma, India, (Assam), Indonesia, Kampuchea, Laos?, 
Malaysia, Thailand, Vietnam? 

Geochelone forstenii (Testudo forstenii) ll R - 152 

Sulawesi (Celebes) Tortoise 

Indonesia (Halmahera, Sulawesi) 

Geochelone gigantea (Testudo gigantea) ll R - 141,152 

Aldabra Giant Tortoise 

Aldabra, Seychelle Islands (introduced) 

Geochelone impressa (Testudo impressa) II K - 48.152 

Impressed Tortoise 

Burma, China, Malaysia (peninsula), Thailand, Vietnam, 

probably also Kampuchea and Laos 

Geochelone paxdalis (Testudo pardalis) II - (?) - 

Leopard Tortoise, Mountain Tortoise 
Angola, Botswana, Ethiopia, Kenya, Lesotho, Malawi, 
Mozambique, Namibia, Somalia, South Africa, Sudan, Swaziland, 
Tanzania, Uganda, Zambia, Zimbabwe 

Geochelone platynota (Testudo platynota) II K - 152 

Burmese Starred Tortoise 
Burma 

Geochelone radiata (Testudo radiata, Asterochelys I V F 46,152 

radiata) LA 

Radiated Tortoise 

Madagascar 

Geochelone sulcata (Testudo siilcata) II - LA - 

Spurred or Grooved Tortoise (F) 

Chad, Ethiopia, Mali, Mauritania, Niger, Nigeria, Senegal, 
Sudan, probably also Cameroon, Central African Republic, 
Djibouti, Upper Volta 

Geochelone travancorica (Testudo travancorica) ll K la 152 

Travancore Tortoise ^ 

India 

Geochelone yniphora (Testudo yniphora, I E F 45,152 

Asterochelys yniphora) LA 

Madagascar Tortoise, Angonoka 

Madagascar 

18 



LA 


4,15,29 




75,144, 




152 


F 
LA 


152 


LA 


101,144, 


F 


152 


LA 


4,32,133 




152 



CITES RDB Exp Ref 

Gopherus agassizii (G. polyphemus agassizii) II V 

Desert Tortoise, Western Gopher Tortoise 

Mexico, USA 

Gopherus berlandieri (G.polyphemus berlandieri) II Id 

Texas Gopher Tortoise, Berlandier's Tortoise 
Mexico, USA (Texas) 

Gopherus flavomarginatus (G.polyphemus I E 

flavomarginatus) 

Bolson Tortoise, Mexican Giant Gopher Tortoise 

Mexico 

Gopherus polyphemus II V 

Florida Gopher Tortoise 

USA 

Homopus areolatus II - LA - 

Parrot-beaked or Areolated Tortoise 

South Afnca 

Homopus boulengeri II - - - 

Boulengers Tortoise 

Namibia, South Afnca 

Homopus femoralis 11 - - - 

Karroo Tortoise 

South Africa 

Homopus signatus II - - - 

Speckled Tortoise 

Namibia, South Afnca 

Kinixys belliana II - F - 

Bell's Hinged Tortoise LA 

Angola, Benin, Botswana, Burundi?, Cameroon, Central African 

Republic, Congo, Djibouti, Ethiopia, Gabon, Gambia, Ghana, 

Gumea, Gumea Bissau, Ivory Coast?, Kenya, Lesotho?, Libena, 

Malawi, Mall, Mozambique, Namibia, Nigena, Rwanda, Senegal, 

Sierra Leone, Somalia, South Afnca, Sudan, Swaziland, Tanzania, 

Togo, Uganda, Zaire, Zambia, Zimbabwe, (introduced on 

Madagascar) 

Kinixys erosa ll - F - 

Schweigger's, Common, or Forest Hinged Tortoise LA 

Angola, Benin, Cameroon, Central Afncan Republic, Congo, 
Equatonal Guinea, Gabon, Gambia, Ghana, Guinea, Ivory 
Coast, Libena, Nigena, Senegal, Sierra Leone, Togo, Uganda, 
Upper Volta, Zaire, (introduced on Madagascar) 

Kinixys homeana 11 - F - 

Home 's Hinged Tortoise LA 

Benin, Cameroon, Congo, Equatonal Guinea, Gabon, Ghana, 
Guinea, Ivory Coast, Liberia, Nigena, Togo, Zaire 

Malacochersus tomieri n K LA 43.152 

Pancake Tortoise 
Kenya, Tanzania 



19 



CITES RDB Exp Ref 

Psammobates geometricus (Testudo geometrica) I V LA 43,47,63 

Geometric Tortoise '52 

South Africa 
Psanunobates oculifer (Testudo oculifera) ll - LA - 

Serrated Tortoise, Toothed Cape Tortoise, Kuhl's Tortoise 

Botswana, Namibia, South Africa 

Psammobates tentorius (Testudo tentoria) II - LA - 

Tent Tortoise 

Namibia, South Afnca 

Pyxis arachnoides ll id LA 43,152 

Spider Tortoise 

Madagascar 

Pyxis planicauda (Testudo planicauda, Acinixys II Id - 152 

planicauda) 

Madagascar Flat-shelled Tortoise 

Madagascar 

Testudo graeca il v la 13,38, 

Spur-thighed, Common, or Greek Tortoise 43,81 . 

Albania, Algeria, Bulgaria, Egypt, Greece, Iran, Iraq, Israel, '31 

Italy (including Sardinia and Sicily, probably introduced), 

Jordan, Lebanon, Libya, Malta, Morocco, Romania, Spain (inc. 

Balearic Islands), Syna, Tunisia, Turkey, USSR, Yugoslavia, 

(introduced m southern France) 

(1 RDB entry is for T.graeca graeca) from Algeria, Libya, Morocco. Spam, 

Tunisia, and introduced in France 

2 Populations in eastern Iran are occasionally treated as a full species, 

T.zarudnyi) 

Testudo hermaimi II V LA 13,38 

Hermann's Tortoise, Greek Tortoise 152 

Albania, Bulgana, France (inc. Corsica), Greece, Italy 
(inc. Sardinia, Sicily), Romania, Spain (Balearic Islands, also 
reported from mainland), Turkey, Yugoslavia 

Testudo horsfieldii (Agrionemys horsfieldii) li - la 13,56 

Horsfield's Tortoise, Afghan Tortoise 

Afghanistan, China, Iran, Pakistan, USSR 

Testudo kleinmanni II Id - 152 

Egyptian Tortoise 

Egypt, Israel, Libya 

Testudo marginata II - LA 13,38 

Margined Tortoise 

Greece, Italy (Sardinia, introduced, possibly mainland also) 



20 



CITES RDB Exp Ref 



Family: CHELONIIDAE 

(Because of the wide and somewhat unpredictable distribution of sea turtles 
away from the nesting beaches, it is impractical to list those countries m whose 
territorial waters individual sea turtles may be expected to occur This applies 
to members of the CHELONIIDAE and DERMOCHELYDIDAE.) 

Caretta caretta 

Loggerhead Turtle 

Nesting recorded on beaches of tropical, subtropical, and 
temperate seas (Atlantic, Indian, Mediterranean, Pacific), 
wandering individuals recorded m temperate and arctic 
waters (to 70°N at Murmansk, USSR, and to 35°S at Rio de la 
Plata, Argentina) 

Chelonia depressa 

Flatback Turtle 

Nests m north and north east Australia, not recorded outside 
adjacent waters 

Chelonia mydas 

Green Turtle 

Nesting recorded on beaches of tropical and subtropical seas 

(Atlantic, Indian, Mediterranean, Pacific), wandering 

individuals recorded in temperate waters (to ca. 52°N in 

English Channel, and to 43°S in Chile). Major breeding and 

feeding areas withm 20°N to 20°S. 

Breeding populations in East Pacific sometimes regarded 

as a full species Chelonia agassizii. 

Eretmochelys imbricata 

Hawksbill Turtle 

Nesting recorded at scattered localities on beaches of tropical 
and subtropical seas, usually between 25°N and 25°S (Atlantic, 
Indian, Pacific, rarely or formerly in Mediterranean), 
wandering individuals occasionally recorded m temperate 
regions, 

Lepidochelys kempii 

Kemp's Ridley, Atlantic Ridley 

Single major nesting site is m Mexico (Rancho Nuevo, on Gulf 
coast of State of Tamaulipas), occurs most frequently in Gulf of 
Mexico, wandering individuals recorded m Atlantic waters (to 
58°N in Scotland), and very occasionally in Mediterranean 

Lepidochelys olivacea 

Olive Ridley, Pacific Ridley 

Nesting recorded on beaches of tropical and subtropical seas 
(Atlantic, Indian, Pacific), most sites between 30°N and 20°S 
occasionally recorded m the Caribbean region although does 
not appear to breed there 



F 


4,6,25,38 




40,41,43, 




54,68,69, 




103.123, 




126,133, 




139,152 


_ 


41,43,130 




152 


F 


4,6,18,25, 


(SK) 


33.37.38, 




40.41,43, 




54,62,65. 




68,69,88, 




99,114, 




117,120, 




124,130, 




133,139, 




146,152 


F 


4,6,22,25 


TS 


30,33,38, 


(SK) 


39,40,41, 




43,54,62, 




68,69,120 




128,130, 




133,139, 




146,152 


F 


120,128, 


(SK) 


130,133, 




139,146, 




4,6,23,25, 




38,41.43, 




69,91,133 




152 


F 


6,40,41, 


(SK) 


43,55,62. 




65,68,90, 




91,121, 




130.139, 




146,152 



CITES RDB Exp Ref 

Family: DERMOCHELYDIDAE 
(DERMOCHELYIDAE) 

(see note under Family CHELONIIDAE. p21) 

Dermochelys coriacea I E F 4,6,25,38, 

Leatherback, Leathery Turtle, Luth 5457 62 

Nesting recorded on beaches of tropical and subtropical seas 68!69 Jo! 

(Atlantic, Indian, Mediterranean, Pacific), most sites between 30°N 87, 1 16, 

and20°S, wandering individuals recorded m temperate waters 121,122, 

(north to Iceland and Norway) 130,132, 

133,146, 
152 

Family: CARETTOCHELYIDAE 

Carettochelys insculpta - K F 152 

New Guinea Plateless Turtle 

Australia (Northern Territory), Indonesia (Inan Jaya), 
Papua New Guinea 



Family: TRIONYCHIDAE 

Lissemys punctata andezsoni (L.p.punctata) 1 - F - 

Indian Flap-shell Turtle 

Bangladesh, Burma (Akyab), India, Nepal, Pakistan 

Trionyxater I - - - 

Cuatro Cienegas or Black Soft-shell Turtle 

Mexico 

Trionyx gangeticus I - F - 

Indian or Ganges Soft-shell Turtle 

Afghanistan, Bangladesh, India, Nepal, Pakistan 

Trionyx hunun I - F - 

Peacock-marked Soft-shell Turtle 

Bangladesh, India 

Trionyx nigricans I R - 152 

Dark Soft-shell Turtle 

Bangladesh 

Trionyx triunguis III _ _ _ 

Nile Soft-shell Turtle (Ghana) 

Angola (inc. Cabmda, and south to Cunene River on Angola- 
Namibia border), Gameroon, Chad, Egypt, Ethiopia, Gabon, 
Gambia, Ghana, Guinea Bissau, Israel, Kenya, Libena, Niger, 
Nigeria, Senegal, Sierra Leone, Somalia, Sudan, Togo, Turkey 
(south), Uganda, Zaire, probably also Benin, Genfral Afncan 
Republic, Congo, Djibouti, Equatorial Guinea, Guinea, 
Ivory Coast, Mali, and Upper Volta 



22 



CITES RDB Exp Ref 

Suborder: PLEURODIRA, 
Side-necked Turtles 

Family: PELOMEDUSIDAE 

Erynuiochelys madagascaiiensis (Podocnemis II? Id - 152 

madagascariensis) 

Madagascar 

(Until recently (see note on p8), a majority of taxonomists have assigned this 
species, and Peltocephalus dumeriliana, to the genus Podocnemis. All species 
of Podocnemis are listed on CITES Appendix II. It is here assumed, pending 
discussion of the matter that the intention was to include these two species also, 
regardless of a change in generic assignment) 

Pelomedusa subrufa III _ _ _ 

Helmeted Turtle (Ghana) 

Angola, Benin, Botswana, Cameroon, Ethiopia, Ghana, Kenya, 
Madagascar, Malawi, Mali, Mauntania, Mozambique, Namibia, 
Nigena, Saudi Arabia (southwest), Senegal, Somalia, South Afnca, 
Sudan, Tanzania, Togo, Uganda, North Yemen, Zaire, Zambia, 
Zimbabwe, probably also Central Afncan Republic, Chad, Congo, 
Djibouti, Equatonal Guinea, Gambia, Guinea, Guinea Bassau, 
Ivory Coast, Libena, Niger, Upper Volta 

Peltocephalus dumeriliana (Peltocephalus tracaxa, IP - F 53,135 

new name; Podocnemis dumeriliana) 

Brazil, Colombia, Ecuador? French Guiana? Peru, Venezuela 
(see note under Erymnochelys madagascariensis, above) 

Pelusios spp. 

Ghana has proposed that all species of Pelusios found in Ghana be 
placed on Appendix III, the following four species, of the eight 
currently recognised by Wermuth & Mertens (1977), may be 
expected to occur m Ghana, The systematics of this genus is 
subject to some dispute 

Pelusios adansonii III _ _ - 

White-breasted Side-necked Turtle, Adanson's Turtle (Ghana) 

Angola, Chad, Congo, Gambia, Mali, Nigena, Senegal, Sudan, 
Zaire, possibly Ghana; Cape Verde Is? 

Pelusios castaneus (formerly mc P.subniger) III _ - _ 

(Ghana) 

Angola, Burundi, Congo, Gambia, Guinea, Liberia, Madagascar, 
Malawi, Rhodesia, Rwanda, Senegal, Seychelle Islands, Sierra 
Leone, Tanzania, Zaire, Zambia, possibly Ghana 

Pelusios gabonensis HI _ _ _ 

Stnpe-backed Side-necked Turtle, Gaboon Turtle (Ghana) 

West Africa (from Liberia east, inc. Ghana, to Gabon) 

Pelusios niger III _ - - 

Black Side-necked Turtle (Ghana) 
West Africa (from Gambia east, possibly inc. Ghana, to Angola) 

Podocnemis erythrocephala (Podocnemis cayennensis) II K F 52,53 

135 152 
Irapuca, Red-headed Amazon Turtle 

Brazil, Venezuela, perhaps Colombia 

23 



CITES RDB Exp Ref 

Podocnemis expansa ll E F 43,53 

Arrau, South American River Turtle ^^^'^ li'^^.n 

lot), 143, 
Bolivia, Brazil, Colombia, Ecuador?, Guyana, Peru, Venezuela, 152 

perhaps French Guiana and Surinam 

Podocnemis lewyana II Id F 53,135 

Magdalena River Turtle ^^^ '^2 

Golombia 

Podocnemis sextuberculata II K F 53,135 

143,152 



(LA) 135,143 
152 



Brazil, Colombia, Peru 

Podocnemis unifilis ll v F 92,119 

Terecay, Tracaja, Yellow-spotted Amazon Turtle 

Bolivia, BrazU, Colombia, Ecuador, French Guiana, Guyana, Peru, 

Surinam, Venezuela 

Podocnemis vogli II - F 53,135 

Colombia, Venezuela 

Family: CHELIDAE 

Phrynops dahli - Id - 152 

Dahl's Toad-headed Turtle 

Colombia (Bolivar) 

Phrynops hogei - id - 152 

Hoge's Sideneck Turtle 

Brazil 

Phrynops rufipes - K - 152 

Red-headed Sideneck Turtle 

Brazil, Colombia 

Platemys pallidipectoris - R L 152 

Chaco Sideneck Turtle 

Argentina, Bolivia?, Paraguay 

Platemys spixii - K - 152 

Sprx's Sideneck Turtle 

Argentina, Brazil, Uruguay 

Pseudemydura tunbrina I E - 43,57 

Short-necked or Western Swamp Turtle ^^' ' ^^' 

130, 13o 
Australia (BuUsbrook) 



24 



CITES RDB Exp Ref 

Order: CROCODYLIA 

For the many and variable trade names of Crocodylia, see King & Brazaitis. 
1971, and Wermuth&Fuchs, 1978. (See Reference list C,p61) 

Family: ALLIGATORIDAE 

Alligator mississippieiisis II - (LA) 4,43.44 

Amencan Alligator (SK) 60,100, 

USA '''''' 

Alligator sinensis I e f 42,43.102, 

Chinese Alligator ^^ ^^2 

China 

Caiman crocodilus sub-spp. 

Caiman crocodilus apaporiensis I Id SK 42,43,152 

(C.sclerops apaporiensis) 

Rio Apapons (Spectacled) Caiman 

Colombia 

Caiman crocodilus (C.sclerops) II Id/V SK 42,43, 

3 to 5 remaining subspecies ^'^^ 152, 1 S3 

Spectacled Caiman 

Argentina, Bolivia, Brazil. Colombia. Costa Rica, Ecuador, 
French Guiana, Guatemala, Guyana, Honduras, Mexico, 
Nicaragua, Panama, Paraguay, Peru, El Salvador. Surinam. 
Tnnidad & Tobago. Uruguay, Venezuela (C.c.yacare, the 
Jacare. from Argentina. Bolivia. Brazil, Paraguay, Uruguay, is 
sometimes treated as a fuU species) 

(In Part 1 of the new Amphibia-Reptilia RDB (ref. 152) the three widely 
recognised subspecies are categorised as follows. C.C. crocodilus 'V; 
C.c. fuscus V and C.c. yacare 'Id') 

Caiman latirostris I E SK 42,43,152 

Broad-nosed Caiman ^ ' 

Argentina, Brazil, Paraguay, Uruguay 

Melanosuchus niger I E SK 42,43,152 

Black Caiman ^^^ 

Bolivia, Brazil. Colombia, Ecuador. French Guiana, Guyana, 
Paraguay, Peru 

Paleosuchus palpebrosus II - (SK) 42,43,152 

Dwarf Caiman. Cuvier's Smooth-fronted Caiman 

Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru. 

Surinam, Venezuela 

Paleosuchus trigonatus n - (SK) 42,43,152 

Schneider's Smooth-fronted Caiman 

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru. 

Surinam, Venezuela 



25 



CITES RDB Exp Ref 

Family: CROCODYLIDAE 

Crocodylus acutus I E SK 4,42,43, 

American Crocodile ^S^^^c 

lo/,14b, 

Belize, Colombia, Costa Rica, Ecuador, Greater Antilles (Cuba, 152 

Domonican Republic, Haiti, Jamaica), Guatemala, Honduras, 
Mexico, Nicaragua, Panama, Peru, El Salvador, USA (Flonda), 
Venezuela 

Crocodylus cataphractus I Id SK 42,43 

African Sharp-nosed or Slender-snouted Crocodile ^' 93, IW 

Angola (Cabinda), Benin, Cameroon, Central African Republic, 
Chad, Congo, Equatorial Guinea, Gabon, Gambia, Ghana, 
Guinea, Guinea Bissau?, Ivory Coast, Liberia, Mali, Mauritania, 
Nigena, Senegal, Sierra Leone, Tanzania (Ujiji), Togo, Upper 
Volta, Zaire, Zambia 

Crocodylus intermedius I E SK 42,43, 

Onnoco Crocodile '^''■'^^ 

Colombia, Venezuela, (and single record from Grenada) 

Crocodylus jotuisom(C.johnstoiii) II V SK 42,43,130, 

Australian Fresh-water Crocodile, Johnson's Crocodile 14 j , lo^ 

Australia 

Crocodylus moreletii I E SK 42,43,60, 

Morelet's Crocodile ^I'^W' 

Belize, Guatemala, Mexico 

Crocodylus niloticus I V SK 42,43 47 

Nile Crocodile ^ ?^'^3,109, 

lo^ 
Angola, Benin, Botswana, Burundi, Cameroon, Central Afncan 
Republic, Chad, Congo, Egypt, Equatorial Guinea, Ethiopia, 
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, 
Kenya, Liberia, Madasgascar, Mali, Malawi, Mozambique, 
Namibia, Niger, Nigena, Rwanda, Senegal, Sierra Leone, 
Somalia, South Africa, Sudan, Swaziland, Tanzania, Togo, Uganda, 
Upper Volta, Zaire, Zambia, Zimbabwe 

Crocodylus novaegumeae sub-spp. I E SK 42,43,152 

C.n.mindoreiisis 

Mmdoro or Philippines Crocodile 

Philippine Islands 

(sometimes treated as a full species, C. mindorensis , as m RDB) 

C.n.novaeguineae ll v SK 42,43 

New Guinea Crocodile 

Indonesia (Aru Islands, West Irian), Papua New Guinea 

Crocodylus palustris I V SK 42,43,44 

Mugger, Marsh Crocodile, Broad-snouted Crocodile 

Bangladesh, India, Iran (Sarbaz River), Nepal, Pakistan, Sn Lanka, 
(Burma?) 



112,152 



85,96,149, 
150,152 



26 



CITES RDB Exp Ref 

Crocodylus porosus I e sk 42,43.44, 



Estuarme or Salt-water Crocodile 



II 85,98,130, 

(Papua New 147,149, 



Australia, Bangladesh, Brunei, Burma, India (inc. Andaman Guinea) 152 

Islands), Indonesia, Kampuchea, Malaysia, Palau Islands, Papua 

New Guinea, Philippine Islands. Solomon Islands, Sn Lanka, 

Thailand, Vanuatu, Vietnam, also records of wandering individuals 

from the Cocos-Keeling Islands and Fiji Islands. Extinct m 

Singapore 

Crocodylus rhombifer I e SK 42,43,60, 

Cuban Crocodile '^5,152 

Cuba (mc. Isla de Pinos) 

Crocodylus siamensis I e sk 42,43,98. 
Siamese Crocodile ^^^ 

Indonesia (Borneo, Java), Kampuchea, Laos, Malaysia 
(peninsula), Thailand, Vietnam 

Osteolaemus tetra^is I Id SK 42.43,93, 

West African Dwarf Crocodile ^ '^^ 

Angola (Cabinda). Benin, Cameroon, Central Afncan Republic, 

Congo, Gabon. Gambia. Ghana. Guinea. Guinea Bissau. 

Ivory Coast, Libena, Nigena. Senegal. Sierra Leone. Togo, 

Uganda, Upper Volta, Zaire 

(the Zaire population is sometimes treated as a full species, O.osbomi) 

Tomistoma schlegelii I E SK 42 43 152 

False Ghanal, False Gavial 

Indonesia (Borneo, Sumatra), Malaysia (peninsula, Sarawak), 
Thailand (possibly extinct) 

FamHy: GAVIALIDAE 

Gavialis gangeticus I E K 3.42,43. 

Ghanal. Gavial TazI'S' 

Bangladesh, Bhutan (probably extmct), India, Nepal, Pakistan 152 



Order: RHYNCHOCEPHALIA 

Family; SPHENODONTIDAE 

Sphenodonpunctatus I R - 27,43, 

Tuatara !?^' 

New Zealand 



152 



27 



CITES RDB Exp Ref 

Order: SQUAMATA, 

Lizards, Snakes, Amphisbaenians 

Suborder: SAURIA, Lizards 

Family: GEKKONIDAE 

Cyrtodactylus serpensinsula 

(Gymnodactylusseipensinsula) II R - 12,43, 

Serpent Island Gecko ^^' ^'^ 

Round Island, Serpent Island, (Mauritius) 

Oedura reticulata - V - 43,130 

Reticulated Velvet Gecko 

Australia 

Phelsuma spp. 

Genus Phelsuma: Day Geckos, certain species may appear m the Live 
Animal trade. The systematics of this genus is subject to some dispute. 

Phelsuma abbotti II _ _ _ 

Abbott's Day Gecko 

Aldabra, Amuante Islands, Assumption, Cosmoledo Island, 

Madagascar Seychelle Islands 

(P.a. longlnsulae and P.a pulchra from the Seychelles are sometimes regarded 
as a separate species, P.longinsulae) 

Phelsuma agalegae II - - 24 

Agalega Island (Mauritius) 

Phelsuma andamanensis ll - - - 

Andaman Day Gecko 

Andaman Islands (India) 

Phelsuma astriata II - - - 

Astove Island, Seychelle Islands 

Phelsuma barbouri II - - - 

Madagascar 

Phelsuma bimaculata II - - _ 

Madagascar 

(sometiraes treated as a subspecies of P.quadiiocellata) 

Phelsuma breviceps II - - - 

Madagascar 

(sometimes treated as a subspecies of P. mutabilis) 

Phelsuma cepediana II - - 77 

Mauritius Greater Day Gecko 

Mauritius, Reunion 

(it has been suggested that P.c borbonica from Reunion should be regarded 
as a fuU species) 



28 



CITES RDB Ebcp Ref 
Phelsuma comorensis II _ - - 

Comoro Islands 

(sometimes treated as a subspecies of P. dubia) 

Phelsuma dubia II - - - 

Madagascar; Tanzania (inc. Zanzibar) 

Phelsuma edwardnewtonii (E newtoni) II E - 43.82 

Rodnguez Day Gecko 

Rodriguez Island (Mauritius) 

(has been reported extinct, see Ref, 82) 

Phelsuma flavigulazis II - - - 

Madagascar 

Phelsuma guentheri II R - 12,43, 

76,77 



78,83 



Round Island Day Gecko 
Round Island (Mauritius) 
Phelsuma g\mnbeaui II - - 78,82 

Mauritius 

Phelsuma guttata II - _ _ 

Madagascar 

Phelsuma laticauda II - - - 

Gold-dust Day Gecko 

Comoro Islands, Farquhar Island (Seychelles)?, Madagascar 

Phelsuma Uneata II _ _ _ 

Side-stnped Day Gecko 

Madagascar 

Phelsuma madagascaxiensis II _ - _ 

Madagascar Day Gecko 

Madagascar, SeycheUe Islands 

(P.m. sundbergi from the Seychelles is frequently regarded as a tuU species) 

Phelsuma modesta II - - - 

Madagascar 

Phelsuma mutabilis II - - - 

Madagascar 

Phelsuma ocellata (Rhoptropella ocellata) II - - 47 

Namibia, South Africa 

Phelsuma omata (P. viitsoni) II - - 12,76 

7783 

Mauritius (mc. Round Island), Reunion 



29 



CITES RDB Exp Ref 
Phelsumaparkeri II _ _ _ 

Pemba Island (Thnzama) 

(sometimes treated as a subspecies of P. abbotti) 

Phelsuma quadriocellata II _ - _ 

Madagascar 

Phelsuma serraticauda II _ _ _ 

Madagascar 

(sometimes treated as a subspecies of P. flavigularis) 

Phelsuma standingi II _ _ _ 

Madagascar 

Phelsuma trilineata II _ _ - 

Madagascar 

Phelsuma v-nigra II _ _ _ 

Comoro Islands 

(sometimes Created as a subspecies of P. abbotti) 

FamHy: PYGOPODIDAE 

Paradelma orientalis II _ _ _ 

Queensland Snake-lizard 
Australia 

Family: XANTUSIIDAE 

Xantusia riversiana (Klauberina riversiana) - R - 43, 

Island Night Lizard 
USA (California) 

Family: AGAMIDAE 

Hydrosauzus pustulatus - v F 43 

Sail-fm Lizard, Soa-Soa Water Lizard, (Crested Lizard) ^^ 

Philippine Islands 
Uromastyx spp. 

Species of Uromastyx are widely termed Spiny-tailed Lizards, Palm Lizards, 
or Mastigures 

Uromastyx acanthinurus II - LA - 

Black Spmy-tailed Lizard ^^^ 

Algeria, Chad, Egypt, Libya, Mali, Mauritania, Morocco, Niger, 
Senegal, Sudan, Tunisia, Western Sahara 

(U.a. geyii from the mountains of soutliem Algena and nortfiem Niger is 
frequently treated as a full species) 



30 



CITES RDB Exp Ref 

Uromastyx aegyptius ll _ _ _ 

Egyptian Spmy-tailed Li2ard 

Egypt, Iraq, Israel, Saudi Arabia, probably Jordan 

Uromastyx asmussi II - SK - 

(Horn Lizard) 

Afghanistan, Iran, Pakistan 

Uromastyx benti II - - - 

South Yemen 

Uromastyx hardwickii II - F 7,72 

Hardwick's Spiny-tailed Lizard 

Afghanistan, India, Pakistan SK LA 

Uromastyx loiicatiis II _ - _ 

Iran, Iraq 

Uromastyx macfadyeni II _ _ _ 



Ethiopia, Somalia 

Uromastyx microlepis II - F 

(Horn Lizard SmaU-grain) 

Bahrain, Iran, Iraq, Kuwait, Oman, Saudi Arabia, South Yemen, 

United Arab Emirates, probably also Qatar and North Yemen 

Uromastyx ocellatus ll 

Egypt, Sudan 

Uromastyx omatus II 

Egypt, Israel, Jordan, Saudi Arabia, Syria 

Uromastyx philbyi n 

Saudi Arabia 

Uromastyx princeps II 

Ethiopia, Somalia 

Uromastyx thomasi 11 

Oman 

Famny: CHAMAELEONIDAE 

Chamaeleo adolfifriderici ll 

Rwanda, Uganda, Zaire 



SK 



31 



CITES RDB Exp Ref 
Chamaeleo af finis II _ - _ 

Ethiopia, Somalia, probably Djibouti 

Chamaeleo africanus II _ _ _ 

African Chameleon 

Cameroon, Chad, Egypt, Ethiopia, Mali, Niger Nigeria, 
Somalia, Sudan, probably Djibouti and Upper Volta 

Chamaeleo anchietae II - - _ 

Angola, Ihnzania, Zaire 

Chamaeleo angeli II - - - 

Madagascar 

Chamaeleo antimena II - _ _ 

Madagascar 

(formerly mc. in C. rhinoceratus) 

Chamaeleo balteatus II _ _ _ 

Two-banded Chameleon 

Madagascar 

(sometimes treated as a subspecies of C.bifidus) 

Chamaeleo belalandaensis II - _ _ 

Madagascar 

Chamaeleo bifidus II - - - 

Madagascar 

Chamaeleo bitaeniatus* II - - - 

Burundi, Ethiopia, Kenya, Rwanda, Somalia, Sudan, Iknzania, 
Uganda, Zaire 

Chamaeleo boettgeri II _ _ _ 

Madagascar 

Chamaeleo brevicomis II - _ _ 

Short-homed Chameleon 

Madagascar 

Chamaeleo calyptratus II _ _ _ 

South Yemen, North Yemen 

Chamaeleo campani II - _ _ 

Madagascar 



*Recently (Bohme and Klaver) three taxa formerly treated as subspecies of C.bitaeniatus have been elevated to 
species rank: C.ellioti, C.hoehnelii and C.kinetensis 

32 



CITES RDB Exp Ref 
Chamaeleo capuroni II - - - 

Madagascar 

Chamaeleo carpenteri n - - - 

Uganda, Zaire 

Chamaeleo cephalolepis II _ _ - 

Comoro Islands 

Chamaeleo chamaeleon II - LA 38,110 

Common or Mediterranean Chameleon 

Algena, Cyprus, Egypt, Gibraltar?, Greece (Crete only), India, 
Israel, Italy (Sicily only), Jordan, Lebanon, Libya, Morocco, 
Oman, Pakistan. Portugal, Saudi Arabia, South Yemen, Spain 
(inc. Canary Islands), Sn Lanka, Syria, Tunisia, Turkey, 
North Yemen 

(populations in India, Pakistan and Sn Lanka are sometimes treated as a full 
species, C.zeylanicus, and Arabian populations as C.arabicus) 

Chamaeleo chapini II _ _ _ 

Zaire 

Chamaeleo czistatus II - _ _ 

Crested Chameleon 

Cameroon, Equatorial Guinea, Gabon, Nigena 

Chamaeleo cucullatus II _ _ _ 

Madagascar 

Chamaeleo deremensis II _ _ _ 

Tanzania (Usambara Mts) 

Chamaeleo dilepis II _ _ - 

Flap-necked Chameleon 

Angola, Botswana, Burundi, Cameroon, Congo, Equatorial 
Guinea, Ethiopia, Gabon, Kenya, Malawi, Mozambique, Namibia, 
Nigena, Rwanda, Somalia, South Afnca, Tanzania, (inc. Pemba 
Island and Zanzibar), Uganda, Zaire, Zambia, Zimbabwe 

Chamaeleo eisentrauti ll _ _ _ 

Cameroon 

Chamaeleo fallax II _ _ _ 

Madagascar 

Chamaeleo fischeri n _ _ _ 

Fischer's Chameleon 

Kenya, Tanzania 

Chamaeleo fuellebomi II _ - - 

Tanzania 

33 



CITES RDB Exp Ref 



Chamaeleo fiircifer ll 

Forked Chameleon 

Madagascar 

Chamaeleo gallus II 

Madagascar 

Chamaeleo gastrotaenia II 

Madagascar 

Chamaeleo globifer II 

Madagascar 

Chamaeleo goetzei II 

Malawi, Tanzama, Zambia 

Chamaeleo gxacilis ll 

Graceful Chameleon 

Angola, Benin, Cameroon, Central African Republic, Chad, 
Congo, Ethiopia, Equatorial Guinea, Gabon, Gambia, Ghana, 
Guinea, Guinea Bissau, Ivory Coast, Kenya, Liberia, Mali, Nigena, 
Senegal, Sierra Leone, Somalia, Sudan, Tanzania, Togo, 
Uganda, Zaire, probably Djibouti 

Chamaeleo guibei II 

Madagascar 

Chamaeleo hoehnelii II 

Kenya, Uganda 

Chamaeleo incomutus ll 

Tanzania 

Chamaeleo jacksonii ll 

Jackson's Three-homed Chameleon 

Kenya, Tanzania 

Chamaeleo johnstoni II 

Burundi, Central African Republic, Rwanda, Uganda, Zaire 
Chamaeleo labordi ll 

Madagascar 

(sometimes treated as a subspecies of C. rhinoceratus) 

Chamaeleo lateralis ll 

Madagascar 

Chamaeleo laterispinis II 

Tanzania 

34 



CITES RDB Exp Ref 
Chamaeleo linotus ll - - - 

Madagascar 

Chamaeleo malthe II - - - 

Madagascar 

Chamaeleo marshalli II - - - 

Short-tailed Dwarf Chameleon 
Mozambique, Zimbabwe 

(alternatively regarded as a member of the Dwarf Chameleon genus 
Rhampholeon) 

Chamaeleo melleri ll - - - 

Meller's Chameleon 

Malawi, Tanzania 

Chamaeleo minor II - - - 

Madagascar 

Chamaeleo mlanjensis II - - - 

Malawi 

Chamaeleo monachus II - - - 

Socotra Chameleon 

Socotra (South Yemen) 

Chamaeleo monoceras II - - - 

Madagascar 

Chamaeleo montium (inc C.feae) II - - - 

Mountain Chameleon 

Cameroon, Equatonal Guinea, Nigena 

Chamaeleo namaquensis ll - - - 

Desert Chameleon 

Angola, Namibia, South Afnca 

Chamaeleo nasutus II - - - 

Madagascar 

Chamaeleo oshaughnessyi II - - - 

Madagascar 

Chamaeleo oustaleti II - - - 

Oustalet's Giant Chameleon 

Madagascar, (and introduced m Ngong Forest 
near Nairobi, Kenya) 

Chamaeleo oweni II - - - 

Owen's Three-homed Chameleon 

Cameroon, Congo, Equatorial Guinea, Gabon, Nigena, Zaire 



35 



Chamaeleo pardalis (mc C.guentheri) 

Panther Chameleon 

Madagascar, (introduced on Reunion) 

Chamaeleo parsonii 

Parson's Giant Chameleon 

Madagascar 

Chamaeleo peyrieresi 



CITES RDB Exp Ref 
II - - 82 



Madagascar 
Chamaeleo pfefferi 

Cameroon 
Chamaeleo polleni 

Comoro Islands 
Chamaeleo pumilis 

Vanegated Dwarf Chameleon 
Mozambique, Namibia, South Afnca 
Chamaeleo quadricomis 
Four-homed Chameleon 
Cameroon, Nigena 
Chamaeleo quilensis 

Angola, Benm, Botswana, Cameroon, Congo, Equatorial Guinea, 
Gabon, Kenya, Malawi, Mozambique, Namibia, Nigena, 
South Afnca, Tanzania (including Pemba and Zanzibar), Togo, 
Uganda, Zaire, Zambia, Zimbabwe 

Chamaeleo rhinoceratus 

Madagascar 
Chamaeleo rudis* 

Burundi, Kenya, Rwanda, Tanzania, Uganda 
Chamaeleo senegalensis 

Senegal Chameleon 

Benin, Burundi, Cameroon, Central Afncan Republic, Ethiopia, 
Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, Kenya, 
Libena, Mali, Nigena, Rwanda, Senegal, Sierra Leone, Sudan, 
Tanzania, Togo, Uganda, Zaire, Zan±)ia 

Chamaeleo spinosus 

Spiny Chameleon 
Tanzania 
Chamaeleo tempeli 



Tanzania 

^Recently (Bohme and Klaver) two taxa formerly treated as subspecies of 
C.rudis have been elevated to species rank C.schoutedeni and C.schubotzi. 



36 



CITES RDB Exp Ref 

Chamaeleo tenuis II - - - 

Slender Chameleon 

Tanzania 

Chamaeleo tigris II - - - 

Seychelle Islands 

Chamaeleo tsaratananensis II - - - 

Madagascar 

Chamaeleo tuzetae 11 - - - 

Madagascar 

Chamaeleo verrucosus II - - - 

Madagascar 

Chamaeleo wemexi II - - - 

Tanzania 

Chamaeleo wiedersheimi II - - - 

Cameroon, Nigena 

Chamaeleo willsii II - - - 

Madagascar 

Chamaeleo xenorhinus II - - - 

Uganda, Zaire 

FamHy: IGUANIDAE 

Amblyrhynchus cristatus II R - 43 

Galapagos Marine Iguana 

Galapagos Islands (Ecuador) 

Anolis roosevelti - E - 43 

Giant Anole, Culebra Giant Ancle 
Isla Culebra (Puerto Rico) 
(possibly extinct) 

Brachylophiis fasciatus I E - 17,43, 

Fiji Banded Iguana ' '^ 

Fiji Islands, Tonga Islands 
Brachylophus vitiensis I - - 153 

Fiji Crested Iguana 

Fiji Islands (Yaduataba Island) 

Conolophus pallidus II R - 43 

Bamngton Land Iguana 

Santa Fe ( = Bamngton) Island in Galapagos group (Ecuador) 



37 



CITES RDB Exp Ref 

Conolophus subcristatus II V - 43.114 

Galapagos Land Iguana 

Galapagos Islands (Ecuador) 

Cyclura carinata I R LA 43,129 

Turks and Caicos Ground Iguana 

Bahama Islands, Turks and Caicos Islands 

Cyclura coUei (C.lophoma) - possibly extinct I _ _ _ 

Jamaica's Ground Iguana 

Jamaica 

Cycluza comuta (mc C.stejnegeri) I v/K - 20,43 

Rhinoceros Iguana 

Dominican Republic, Haiti, Isla Mona, Navassa Id. 

{in RDB C.c.stejnegeri from Isla Mona is in category 'V, while C.c. comuta from 
the rest of the range is in Preamble 9— probably threatened) 

Cyclura cycMura (mc C.baelopha, C.figginsi, plus I R LA 21,43 

Cinomata) ^ 

Bahama Islands 

Cyclura nubila (inc C.caymanensis, C.macleayi) I v - 43 

Cayman Islands, Cuba 

Cyclura pinguis I E - 19,20 

Anegada Ground Iguana ' 

Anegada Id. (British Virgin Islands) 

Cyclura ricordi I K - 20,43 

Hispaniola Ground Iguana, Ricord's Ground Iguana 

Dominican Republic, Haiti 

Cyclura rileyi (mc C.cristata, C.nuchalis) I E/R F 43 

LA 

Bahama Islands 

(In RDB C.r.rileyi is in category E' the other two races are in 'R') 

Gainbeliasilus(Crotaphytussilus, C.wislizeniisilus) - E - 4,43,75 

Blunt-nosed or San Joaquin Leopard Lizard 

USA (California) 

Iguana delicatissima II - F 

Lesser Antilles (inc. Anguilla, Antigua, He Chevreau, 
La D6siderade, Dominica, He Fourchue, Les lies Fregates, 
Guadeloupe, Martinique, Nevis, Les lies des Saints, 
St, Barthelemy, St. Eustatius, St. Martin) 

Iguana iguana ll - F 1 1 1, lis 

Common Iguana, Green Iguana, (South American 'Chameleon') ^^ 

Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French 
Guiana, Guatemala, Guyana, Honduras, Lesser Antilles 
(inc. Grenada, the Grenadines, Guadaloupe, Montserrat, Saba, 
Les lies des Saints, St. Lucia, St. Vincent, Virgin Islands), Mexico, 
Netherlands Antilles, Nicaragua, Panama, Paraguay, Peru, 
El Salvador, Surinam, Tnnidad & Tobago, Venezuela 

38 



CITES RDB Exp Ref 

Phrynosoma coronatum blainvillei II K - 4,43,75 

San Diego Homed Lizard 

Mexico (Baja California), USA (California) 

Sauromalus vazius I - LA - 

San Esteban Island Chuckwalla ^ 

Mexico (San Esteban Island) 

Family: LACERTIDAE 

Podarcis filfolensis filfolensis (Lacerta f.filfolensis) - R - 38,43 

Filfola Lizard 
Filfola Island (Malta) 

Gallotia simonyi (Lacerta simonyi) - E - 43,108 

Hierro Giant Lizard 

Canary Islands (Hierro Island) 

Family: CORDYLIDAE 

Cordylus campbelli II _ _ _ 

Namibia 

Cordylus cataphractus n - - - 

South Africa 

Cordylus coenilopunctatus ll _ _ _ 

South Africa 

Cordylus cordylus II - - - 

Angola, Botswana, Ethiopia, Kenya, Lesotho', Malawi, 
Mozambique, Namibia, South Africa, SwazUand', 
Tanzania, Zaire, Zambia, Zimbabwe 

Cordylus giganteus n - - - 

Lesotho, South Afnca 

Cordylus macropholis II - - - 

South Africa 

Cordylus namaquensis II - - - 

Namibia 

Cordylus peers! II - - - 

South Afnca 

Cordylus polyzonus II - - - 

Botswana, Lesotho', Namibia, South Africa 

Cordylus ukingensis II - - - 

Tanzania ,□ 



CITES RDB Exp Ref 
Cordylus vittifer II - - - 

Angola, Botswana, Lesotho?, Namibia?, South Africa, 
Swaziland 

Cordylus warreni II - - - 

Lesotho? Mozan±iique, South Afnca, Swaziland?, 
Zimbabwe 

Pseudocordylus capensis ll - - - 

South Afnca 

Pseudocordylus langi ll - - - 

Lesotho, South Afnca 

Pseudocordylus microlepidotus II - - - 

Namibia, South Africa 

Pseudocordylus robertsi II - - - 

South Africa 

Pseudocordylus spinosus II - - - 

South Afnca 

Family: TEIIDAE 

Ameiva polops - E - 43,58,59 

St. Croix Ground Lizard 

St. Croix and adjacent cays (US Virgin Islands) 

Cnemidophorus hyperythrus II V - 4,43 

Orange-throated WhiptaU 

Mexico (Baja California), USA (California) 

Crocodilurus lacertinus II - SK - 

Brazil, French Guiana, Guyana, Surinam 

Dracaena guianensis II - LA - 

Caiman Lizard, (Croco-Teju) 

Brazil, French Guiana, Peru, possibly Guyana and other 
countnes bordenng the Amazon basin 

Tupinambis rufescens (mc T.duseni and II - LA - 

T.teguixin sebastiani) SK 

(Argentine Teju, Red Teju) 

Argentina, Bolivia, Brazil, Paraguay, Uruguay 

Tupinambis tegubcin (Argentine or Brazil Teju) II - LA - 

(mc. T.nigropunctatus)Oacamra), SK 

Common Tegu or Banded Tegu 

Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, 
Paraguay, Peru, Surinam, Tnnidad & Tobago, Venezuela, 
probably Ecuador 
40 



CITES RDB Exp Ref 

FamHy: SCINCIDAE 

Gongylomorphus bojerii fontenayi - R - 43,77 

Macabe Forest Skink 

Mauritius 

Leiolopisma telfaizii _ r _ 12,43, 76, 

Round Island Skink 77,78,83 

Round Island (Mauntius) 

Macroscincus coctaei - probably extinct - Id - 38,43,140 

Cape Verde Giant Skmk 
Cape Verde Islands 

FamHy: ANGUIDAE 

Gerrhonotus panamintinus _ K - - 

Panamint Alligator Lizard 
USA (Califorrua) 

Family: ANNIELLIDAE 

Aiuiiella pulchra nigra - E - 4.43 

Black Legless Li2ard 
USA (California) 

FamHy: HELODERMATIDAE 

Heloderma horridum ll id (LA) 43 

Beaded Lizard 

Guatemala, Mexico 

Helodenna suspectum ll v (LA) 43,75 

Gila Monster 

Mexico, USA 

Family: VARANIDAE 

Varanus acanthums II - - - 

Spmy-tailed Pygmy Monitor 

Australia 

Varanus bengalensis (V.monitor, itic V.nebulosus) I - SK - 

Indian Monitor, Common Monitor (India) ^ 

(Agra, Bengal, Indian Small-gram, Penang, Rice, 
Talagoya, - Lizard) 

Afghanistan, Bangladesh, Burma, India, Indonesia, Iran, 
Malaysia, Nepal, Pakistan, Sn Lanka, Thailand, Vietnam, 
probably Kampuchea and Laos 

Varanus brevicauda ll - - - 

Short-tailed Pygmy Monitor 

Australia 

Varanus caudolineatus II - - - 

Streak-tailed Monitor 
Australia 

41 



CITES RDB Exp Ref 

Varanus dumerilii II - SK - 

Dumenl's Monitor, ^ 

(Fish Lizard) 

Burma, Indonesia, Malaysia, Thailand 

Varanus eremius II _ _ _ 

Desert Pygmy Monitor 

Australia 

Varanus exanthematicus II - sK 47 

African Savanna Monitor (Afncan Large-gram Lizard) ^ 

Angola, Benin, Botswana, Burundi, Cameroon, Central Afncan 

Republic, Chad, Djibouti, Ethiopia, Gambia, Ghana, Guinea, 

Gumea Bissau, Ivory Coast, Kenya, Lesotho, Malawi, Mali, ' 

Mauntama, Mozambique, Namibia, Niger, Nigena, Rwanda, 

Senegal, Sierra Leone, Somalia, South Afnca, Sudan, Swaziland, 

Tanzania, Togo, Uganda, Upper Volta, Zaire, Zambia, Zimbabwe 

Varanus flavescens I _ sK - 

Yellow Monitor (Calcutta or Indian Oval-grain Lizard) ^ 

Bangladesh, India, Nepal, Pakistan 

Varanus giganteus II - - - 

Perentie 

Australia 

Varanus gilleni H _ _ _ 

Pygmy Mulga Monitor 

Australia 

Varanus glauerti il _ _ _ 

Long-tailed Rock Monitor 

Australia 

Varanus glebopalma II _ _ _ 

Australia 

Varanus gouldii II - LA - 

Sand Monitor, Gould's Monitor 

Australia, Indonesia (West Inan), Papua New Gumea 

Varanus grayi II _ _ 5 

Gray's Monitor 
Philippme Islands (Luzon) 

Varanus gziseus I v (F) 43,81 

Desert Monitor, Agra Monitor (India) SK 

(Agra, Bengal, - Lizard, Baghdad Small-gram Lizard) 

Afghanistan, Algena, Chad, Egypt, India, Iran, Iraq, Israel, 

Jordan, Kuwait, Lebanon, Libya, Mauntama, Morocco, Oman, 

Pakistan, Qatar, Saudi Arabia, South Yemen, Sudan, Syna, Tunisia, 

Turkey, United Arab Emirates, USSR, Western Sahara, North Yemen 

(the RDB citation applies only to V.g.caspius, the Central Asian or Transcaspian 

Desert (or Grey) Monitor, from Afghanistan, Iran, Pakistan (northern 

Baluchistan), and USSR) 

Varanus indicus il - SK - 

Pacific Monitor, Mangrove Monitor, (Ambon or Flower Lizard) ^ 

Australia, Caroline Islands, Indonesia, Manana Islands, 
^2 Marshall Islands, Papua New Gumea, Solomon Islands 



CITES RDB Exp Ref 

Varanus karlschmidti II _ - _ 

Sepik Monitor 

Papua New Guinea 

Varanus Komodoensis I R LA 43,66 

Komodo Dragon 

Indonesia (Floras, GiliMota, Komodo, OwadiSami, Padar, Rint]a) 

Varanus mertensi (in V.bulliwallah) II - - - 

Merten's Water Monitor 

Australia 

Varanus mitchelli II - - - 

MitcheU's Water Monitor 

Australia 

Varanus niloticus II - SK 47 

Nile Monitor, (African Small-grain Lizard) J^ 

Angola, Benin, Botswana, Cameroon, Central Afncan Republic, 
Chad, Congo, Djibouti, Egypt, Ethiopia, Equatonal Guinea, 
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, 
Kenya, Lesotho, Libena, Malawi, Mali, Mauritania, 
Mozambique, Namibia, Niger, Nigeria, Senegal, Sierra Leone, 
Somalia, South Africa, Sudan, Swaziland, Tanzania, Togo, 
Uganda, Upper Volta, Zaire, Zan±)ia, Zimbabwe 

Varanus prasinus II - LA - 

Emerald Monitor 

Australia (Torres Straits Islands), Indonesia (West Inan), 
Papua New Guinea 

Varanus primordius II - - - 

Ridge-tailed Monitor 

Australia 

(frequently treated as a subspecies of V.acanthurus) 

Varanus rudicollis II - SK - 

Rough-necked Monitor ^^ 

(Tree Lizard) 

Burma, Indonesia, Malaysia, Thailand 

Varanus salvadorii II - LA - 

Papuan Monitor, Tree Crocodile 

Indonesia (West Irian), Papua New Guinea 

Varanus salvator (m V.bivittatus, V.togianus) II - LA - 

Malayan Monitor, Water Monitor 

(very many names, m leather trade accordmg to patternmg and place of 
origin, eg., No-mark, Plain, Rice, Ring, Borneo, Java, Philippines, Sumatra, 
Semi-Java, Singapore, Siam, Ramgodie, Cabragoya - Lizard) 

Bangladesh, Burma, China, India (inc. Andaman Is.), Indonesia, 
Kampuchea, Laos, Malaysia, Philippine Islands, Singapore?, 
Sn Lanka, Thailand, Vietnam 

Varanus semiremex II - - - 

Mangrove Monitor, Rusty Monitor 

Australia 

Varanus spenceri II - - - 

Spencer's Monitor 

Australia .^ 



CITES RDB Exp Ref 

Varanus storri li - - - 

Dwarf Monitor 

Australia 

Varanus timorensis II - LA - 

Timor Tree Monitor, Spotted Tree Monitor 

Australia, Indonesia, Papua New Guinea 

Varanus tristis ll - - - 

Freckled Monitor, Mournful Tree Monitor 

Australia 

Varanus varius II _ _ _ 

Lace Monitor 

Australia 

Suborder: SERPENTES, 
Snakes 

Family: TYPHLOPIDAE 

Typhlops monensis - R - 43 

Mona Blind Snake 
Isla Mona (Puerto Rico) 

Family: BOIDAE 

A few taxa of the family Boidae are listed on CITES Appendix I, the 
remainder are on Appendix II The actual content of this family has been the 
subject of recent discussion among herpetologists. Both McDowell {1 975, see 
Ref list B) and Underwood { 1 976) remove the genera Bolyeiia, Casarea, 
Exiliboa, Trachyboa, Tropidophis and Ungaliophis from the family Boidae. 
McDowell also removes Loxocemus, while Underwood retains Loxocemus 
and adds Xenopeltis. Pending discussion, the contents of the Boidae given 
here closely follows the more conservative list of Stimson (1969), this also 
appears to be the usage of CITES and RDB, Some changes are made at low 
taxonomic levels, following Sheplan & Schwartz (1974), McDowell (197S, 
1979), and Collins et al (1 978), Python oenpelliensis was newly described by 
Cow (1977), P.saxuloides by Miller & Smith (1980). P.bredli by Cow (1981) 
and P.carinatus by Smith (1 98 1 ). 

Acrantophis dumerili I ? LA - 

Dumenl's Boa 
Madagascar 

Acrantophis madagascariensis I ? LA - 

Madagascar Boa 
Madagascar 

Aspidites melancophalus II - - - 

Black-headed Python 

Australia 

Aspidites ramsayi II _ _ _ 

Woma 
Australia 



44 



CITES RDB Exp Ref 

Boa constrictor (Constrictor constrictor) II - LA - 

Boa Constnctor, ^'^ 

(Ampalague, Giboya, Masacuate) 

Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, 

Ecuador, French Guiana, Guatemala, Guyana, Honduras, 

Lesser Antilles (Domiruca, Santa Lucia), Mexico, Nicaragua, 

Panama, Paraguay, Peru, El Salvador, Surinam, Tnnidad & 

Tobago, Venezuela 

Bolyeria multocarinata I E - 12,43 

Round Island Boa ^^'^ 

Round Island (Mauritius) 
Calabaria reinhardtii II - - - 

Calabar Ground Python 

Benin, Cameroon, Congo, Equatorial Guinea, Gabon, Ghana, 
Ivory Coast, Libena, Nigeria, Togo, Zaire, probably 
Central Afncan Republic 

Candoia aspera (Enygrus asper) II - - - 

Papuan Ground Boa 

Indonesia (Moluccas, West Inan), Papua New Guinea 
(inc. Bismarck Archipelago as far east as Bougainville) 

Candoia bibroni (Enygrus bibroni) II - - - 

Pacific Boa 

Fiji Islands (inc. Rotuma), Loyalty Islands, Vanuatu (= New 
Hebndes), (inc. Banks Islands), Western Samoa, also Ta'u Island 
in Amencan Samoa, Solomon Islands (mcl. Santa Cruz) 

Candoia carinata (Enygrus carinata) II - - - 

Solomons Ground Boa 

Indonesia (eastern islands inc. Moluccas, Sulawesi, West Inan), 
Palau Islands, Papua New Guinea (inc. Bismarck Archipelago), 
Solomon Islands, (incl. Santa Cruz) 

Casaxea dussumieri I E - 12,43, 

78 83 
Round Island Keel-scaled Boa 

Round Island (Mauritius) 

Charina bottae II R LA 43,75 

Rubber Boa 

Canada, USA 

(RDB entry is for Charina b. umbratica, the Southern Rubber Boa, 

from 'solated localities in California) 

Chondropython viridis II - LA - 

Green Tree Python 

Australia (Cape York Peninsula), Indonesia (West Inan), 
Papua New Guinea (mainland, not Bismaick Archipelago), 
Solomon Islands? 

Corallus annulatus (Boa annulata) II - - - 

Annulated Tree Boa 

Colombia, Costa Rica, Ecuador, Nicaragua, Panama 

Corallus caninus (Boa canina) ll - LA - 

Emerald Tree Boa 

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, 

Peru, Surinam, Venezuela 

45 



CITES RDB Exp Ref 

Corallus enydris (C.hortulanus, Boa hortulanus, II - - - 

inc. Corallus or Boa cooki) 

Garden Tree Boa 

Bolivia, Brazil, Colombia, Costa Rica, Equador, French Guiana, 
Guyana, Nicaragua, Peru, Surinam, Tnnidad & Tobago, 
Venezuela, Windward Islands (Grenada, Grenadines, 

St. Vincent) 

Epicrates angulifer II Id (SK) 43 

Cuban Tree Boa 

(Maja (de Santa Maria)), Cuba (inc. Isla de Pinos) 

(RDB entry is for E.a.angulifer from tfie Isle of Pines and two small islands off 

the coast of Cuba) 

Epicrates cenchria ll - LA - 

Rainbow Boa ^^*^^ 

(Araramboia) 

Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, 
French Guiana, Guyana, Panama, Paraguay, Peru, Surinam, 
Tnnidad & Tobago, Venezuela 

Epicrates chrysogaster (E.striatus chrysogaster) II - - 10 

Bahama Islands Boa 

Bahama Islands, Caicos Islands, possibly Turks Islands 

Epicrates exsul ll - - - 

Abaco Island Boa 

Bahama Islands 

Epicrates fordii (E.inomatus fordii) ? - - - 

Ford's Boa 

Dominican Republic, Haiti 

(see note under Epicrates inomatus) 

Epicrates gracilis II - - - 

Haitian Vine Boa 

Dominican Republic, Haiti 

Epicrates inomatus I E - 43 

Puerto Rican Boa 

Puerto Rico 

(E.inomatus, as used by CITES and RDB, appears to be in the sense of Stull 
(193S) and Stimson (1969, Das Tierreich 89) In this usage three subspecies are 
recognised, 1 E.i.inomatus (Puerto Rico), 2. E.i.fordii (Dominican Republic 
and Haiti), 3 E.i.granti (St Thomas and Tortola in the Virgin Islands), 
Good evidence has subsequently been proposed (Sheplan & Schwartz, 1974; 
followed by Schwartz & Thomas, 1976) that E.i.fordii is best regarded as a 
distinct species (E.fordii) and that E.i.granti has greater affinities with the 
Epicrates of Isla Mona, the latter two taxa are accordingly treated as a distinct 
species (E.monensis) in the works cited. The present list includes the West 
Indian species of Epicrates as recognised by Sheplan & Schwartz (1 974) and 
Schwartz & Thomas (1976); the proper treatment of E.fordii and E.monensis 
by CITES and RDB thus remains to be determined, it is assumed that the 
current CITES and RDB treatment will continue to apply to Epicrates 
inomatus (of Sheplan & Schwartz) 

Epicrates monensis (E.gracUis monensis plus ^ R - 43 

E.inomatus granti) 

Isla Mona (Puerto Rico), St. Thomas (US Virgin Islands), 

Tortola (Bntish Virgin Islands) 

(see note under Epicrates inomatus (RDB entry is for E.m. monensis 

from the Isla Mona) 

46 



Epicrates striatus 

Fischer's Tree Boa 

Bahama Islands, Dominican Republic, Haiti 

(RDB entry is for E.s.fosteri from the Bahamas) 

Epiczates subflavus 

Jamaica Boa 

Jamaica 

Eryx colubzinus 

Kenya Sand Boa 

Egypt, Ethiopia, Kenya, Niger, Somalia, Sudan, 

Tanzania, North Yemen' 

Eryx conicus (Gongylophis conicus) 

Indian Sand Boa 
(Baby 'Python') 
India, Pakistan, Sn Lanka 
Eryx elegans 

Afghanistan, Iran, USSR (Turkmenistan) 
Eryxjaculus 

Qavelin) Sand Boa, Spotted Sand Boa 
Albania, Algena, Bulgana, Egypt, Greece, Iran, Iraq, Israel, 
Jordan, Lebanon, Morocco, Romania (possibly extinct), 
Saudi Arabia, Syna, Tunisia, Turkey, USSR, Yugoslavia 

Eryx ja'yakari 

Jayakar's Sand Boa 

Bahrain, Kuwait, Oman, Qatar, Saudi Arabia, South Yemen, 

United Arab Emirates, North Yemen 

Eryxjohnii 

Blunt-tailed Sand Boa 
Afghanistan, India, Iran, Pakistan 
Eryx miliaris 

Iran, USSR 
Eryx muelleri 

MiiUer's Sand Boa 

Cameroon, Chad, Ghana, Ivory Coast, Mali, Mauritania, 
Nigena, Senegal, Sudan, Togo, Upper Volta, probably Benin 
and Central Afncan Republic 

Eryx somalicus 

Somalia 

Eryx tatazicus 

Tartary Sand Boa 

Afghanistan, China, Iran, Mongolia, Pakistan, USSR 

Eunectes barbouri 

Brazil 



CITES RDB Exp Ref 

II R LA 43 

I V LA 43 

II - LA - 



11 - LA 148 
SK 



LA 38 



LA 



47 



Eunectes deschauenseei 

Brazil 

Eunectes murinus 

Anaconda 

Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, 
Peru, Surinam, Trinidad, Venezuela 

Eunectes notaeus 

Yellow Anaconda 

Argentina, Bolivia, Brazil, Paraguay, Uruguay 

Exiliboa placata 



CITES 


RDB Exp 


Ref 


II 


- 


- 


II 


- LA 
SK 


- 



SK 



Mexico 

Liasis albertisii (L.fiiscus albertisii) 

D'albertis' Python 

Indonesia (West Irian), Papua New Guinea, Torres Straits Islands 
(Australia) 

Liasis boa (Bothrochilus boa, Nardoana boa) 

Barred Python 

Papua New Guinea (Bismarck Archipelago, mainland 
New Guinea?), Tokelau Islands? 

Liasis childreni 

Children's (Rock) Python 

Australia 

Liasis mackloti (mc L.fuscus) 

Water Python (common name for L.fuscus) 

Australia, Indonesia (Lesser Sunda Islands), Papua New Guinea 

Liasis maximus 



(known only from the type specimen, from New Gumea; a possible synonym of 
L.papuanus (McDowell, 1975 Ref listB) 

Liasis olivaceus 

Olive (Rock) Python 

Australia 

Liasis papuanus (L. olivaceus papuanus) 

Papuan Python 

Indonesia (West Irian), Papua New Guinea 

Liasis perthensis 

West Australian Rock Python 

Australia 

(taxonomic status uncertain, {Cogger, 197S, Ref. listB)), 

Lichanura trivirgata 

Rosy Boa 

Mexico, USA (Arizona, California) 

(L.t.roseofusca from Mexico and California is often regarded as a full species, 
Lichanura roseofusca) 



LA 75 



48 



CITES RDB Exp Ref 
Loxocemus bicolor II - - - 

Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, 
El Salvador 

Python amethistinus (Liasis amethistiniis) ll - - - 

Amethystine (Rock) Python 

Australia (Cape York Peninsula, Torres Straits Is), Indonesia 
(eastern islands. West Inan), Papua New Guinea, Philippines? 

Python anchietae II - - - 

Angolan Python 

Angola, Namibia 

Python boeleni (Liasis boeleni, Liasis taronga) 11 - - - 

Boelen's Python 

Indonesia (West Irian), Papua New Guinea 

Python bredli ll - - - 

Australia (Northern Territory) 

Python carinatus II - - - 

Rough-scaled Carpet Snake 

Australia (Western Australia) 

Python curtus n - SK - 

Short-tailed Python, Blood Python 

Brunei, Indonesia (Bangka, Kalimantan, Sumatra), Malaysia, 
Thailand (m Malay Peninsula) 

Python molurus sub-spp. - - F 43,148 

Indian Python, Burmese Python, Oriental Rock Python 

Qava Tree Python, Molurus Python, Indian Python, Bengal or 
Saigon Python) 



LA 
SK 



P.m.bivittatus ll v - 

Brunei?, Burma, China, Hong Kong, Indonesia, Kampuchea, 
Laos, Malaysia, Thailand, Singapore, Vietnam 

P.m.molurus I V 

Bangladesh, India, Nepal, Pakistan, Sn Lanka 

Python oenpelliensis II - - 

Australia 

Python regius ll - LA 

Ball Python, Royal Python ^^ 

Benin, Cameroon, Chad, Gambia, Ghana, Guinea, Guinea 
Bissau, Ivory Coast, Mali, Nigena, Senegal, Sierra Leone, Sudan, 
Togo, Uganda, Upper Volta, Zaire, probably also Central African 
Republic, Congo, Equatorial Guinea, Gabon, Libena 



49 



CITES RDB Exp Ref 



LA 
SK 



Python reticulatus 11 

Reticulated Python, (Diamond or Java Rock Python) 
Bangladesh, Brunei, Burma, Indonesia (not to West Inan), 
Kampuchea, Laos, Malaysia, Nicobar Islands (India), Philippines, 
Singapore, Thailand, Vietnam 

Python saxuloides n 

Lesser Afncan Rock Python 

Kenya, probably Malawi, South Africa, poss±)ly Ethiopia, 

Somalia, Uganda 

Python sebae H 

African Python, Water Python, Afncan Rock Python 
(West African Python) 

Angola, Benin, Botswana, Burundi, Cameroon, Central African 
Republic, Chad, Congo, Djibouti, Equatonal Guinea, Ethiopia, 
Gabon, Gambia, Ghana, Guinea, Guinea Bissau, Ivory Coast, 
Kenya, Lesotho, Libena, Malawi, Mali, Mauntania, Mozambique, 
Namibia, Nigeria, Rwanda, Senegal, Sierra Leone, Somalia, 
South Afnca, Sudan, Swaziland, Tanzania, Togo, Uganda, 
Upper Volta, Zimbabwe, Zaire, Zambia 

Python spilotus (Morelia spilotes, Morelia argus) II - SK 

P.s.vaiiegata— Carpet Python, P.s.spilotus— Diamond Python 

Australia, Indonesia (West Inan), Papua New Guinea 

Python timorensis II - - 

Timor Python 

Indonesia (Flores, Timor) 

Sanzinia madagascariensis I K LA 

Madagascar Tree Boa 

Madagascar 

Trachyboa boulengeri II - - 

Colombia, Ecuador, Panama 

Trachyboa gularis II - - 

Ecuador 

Tropidophis battersbyi 11 - - 

Ecuador 

Tropidophis canus II - - 

Bahama Islands 

Tropidophis caymanensis II - - 

Caymans Ground Boa 

Cayman Islands 

Tropidophis feicki II - - 

Cuba 



50 



LA 



LA 47 
SK 



Tropidophis greenwayi 

Caicos Islands 
Tropidophis haetianus 

Haitian Ground Boa 

Cuba, Dominican Republic, Haiti, Jamaica 

Tropidophis maculatus 

Cuba 

Tropidophus melanums 

Cuba, Navassa Island 
Tropidophis nigriventris 

Cuba 

Tropidophis pardalis 

Cuba 

Tropidophis paucisquamis 

Brazil, Peru 
Tropidophis pi]sbryi 

Cuba 

Tropidophis semicinctus 

Cuba 

Tropidophis taczanowskyi 

Ecuador, Peru, Brazil? 
Tropidophis wrighti 

Cuba 

Ungaliophis continentalis 

Guatemala, Honduras, Mexico 
Ungaliophis panamensis 

CoIon±)ia, Costa Rica, Nicaragua, Panama 
Xenoboa cropani 

Brazil 



CITES RDB Exp Ref 
II - - - 



51 



CITES RDB Exp Ref 

Family: COLUBRIDAE 

Clelia clelia (Pseudoboa cloelia) II - LA - 

Musurana 

Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, 

French Guiana, Guatemala, Guyana, Honduras, Lesser Antilles 

(inc. Dominica, Grenada, St. Lucia), Nicaragua, Panama, 

Paraguay, Peru, El Salvador, Surinam, Trinidad, Uruguay, 

Venezuela 

(snakes from Mexico and elsewhere m Central America formerly assigned to 

this species are now usually regarded as a full species, Clelia scytalina) 

Drymarchon corais couperi - V LA 43.133 

Eastern Indigo Snake 

USA 

Elachistodon westermaiuii II - - 35 

Indian Egg-eatmg Snake 

Bangladesh, India, Nepal 

Hydrodynastes gigas (Cyclagras gigas) II - LA 

South American Water 'Cobra ' , '^'^ 

(Beach 'Cobra', Surucucu) 

Argentina, Bolivia, Brazil, Paraguay 

Masticophis flagellum ruddocki - R - 4.43 

San Joaquin Coachwhip '° 

USA (California) 

Masticophis lateralis euryxanthus - R - 4,43 

Almeda Striped Racer '° 

USA (California) 

Nerodia fasciata taeniata (Natrix f.taeniata) - R - 4,43 

Atlantic Saltmarsh Snake ' ^'^ 

USA (Florida) 

Nerodia sipedoninsulanun (Natrix s.insulanun) - K - - 

Lake Erie Water Snake 

Lake Ene (Canada/USA) 

Thamnophis couchi gigas (T.elegans gigas) - R - 4,43 

Giant Garter Snake ^^ 

USA (California) 

Thamnophis couchi hammondi II R LA 4.43 

(T. elegans hammondi) 75 

Two-striped Garter Snake 

Mexico (Baja California), USA (California) 

Thamnophis sirtalis tetrataenia - E LA 4,43 

San Francisco Garter Snake "^^ 

USA (California) 



52 



CITES RDB Exp Ref 

Family: ELAPIDAE 

Najaoxiana - E - 43,81 

(frequently treated as a subspecies of the Asiatic Cobra, Naja naja) 
Central Asian or Oxus Cobra 

Afghanistan, India (Kashmir, Punjab), Iran (northeast), Pakistan, 
(north and northwest), USSR (Transcaspian region) 

Ogmodon vitianiis - V - 43 

Fiji Snake 
Fiji (Viti Levu) 

Family: VIPERIDAE 

Bothrops insulaxis - K - - 

Queimada Island Bothrops 

Brazil (Queimada Island) 

Crotalus unicolor - R LA 43 

Aruba Island Rattlesnake 

Netherlands antilles (Aruba Island) 

Crotalus willardi - EA/ LA 4,28,36, 

4? ISl 

Ridge-nosed Rattlesnake ^"''^ 

Mexico, USA (Arizona, New Mexico) 

(in RDB C.w.obscurus, the New Mexico Ridge-nose Rattlesnake, is 

Endangered, the four other subspecies are Vulnerable) 

Vipera ammodytes transcaucasiana - R LA 43,81 

Transcaucasian Long-nosed Viper 

USSR (Georgia), Turkey 

Vipera latifii - E - 43,97 

Latifi's Viper 

Iran (Lar Valley m Elburz Mtns,) 

Vipera lebetina schweizeri - v - 38,43, 

ID] 

Blunt-nosed Viper, Levantine or Lebetine Viper 
(names applied to V.lebetma as a whole) 
Greece (Cyclades only: Kimolos, Milos, Polyagos, Siphnos, 
possibly AntimUos, Kithnos) 

Vipera xanthinaraddei - R LA 43,81 

Armenian Viper 
USSR, Turkey 



53 



Additions and Amendments 



CITES RDB Exp Ref 

Family: KINOSTERNIDAE 

Kinosternon flavescens spooneri - E - 1S2 

Illinois Mud Turtle 
USA 



54 



References 



(A) Sources of information on the survival status of particular species or groups of species. 

The number of each Reference corresponds to entries in the 'Ref column of the taxonoraic list. The utility of the 
References given is rather variable; some works concentrate entirely on survival status and conservation, while 
others may only make a brief comment on these topics in the course of a largely taxonomic treatment Some of the 
works are popular or semi-popular, while others are more rigorous in approach. Emphasis is given to more recent 
works, mcludmg those not cited m the third edition of the Red Data Book; the latter (item 43 in the list) contains 
references to many other works. It is not the intention to provide a complete bibliography of the extensive 
literature treating the biology and conservation of amphibians and reptiles 

1, ANON, 1971, Santa Cruz Long-toed Salamander granted a reprieve Herpetology5(4):14. 

2, ANON, 1974, Pennsylvania protects the Bog Turtle Heipet. Rev 5(2):44. 

3, ANON 1976. Madhya Pradesh Gharial Survey Newsl. Madras Snake Pk. Trust l(2):2-3. 

4, ASHTON, R,E,, Chairman 1973-74 SSAR Liaison Committee to Regional Societies. Endangered and Threatened 
Amphibians and Reptiles in the United States. Society for the study of Amphibians and Reptiles, Herpelological 
Circular No 5 

5, AUFFENBERGW 1979 A monitor lizard in the Philippines. Oryx 15:39-46. 

6, BACON, PR 1975 Reviewonresearch exploitation and management of the stocks of Sea Turtles 
FAO Fisheries Circular No. 334. (Caribbean and western Atlantic regions). 

7 EHATIA, M L & TILAK, R 1976 Some notes on the Spiny-tailed Lizard (Uromastyx hardwickii) Gray 
Newsl. Madras Snake Pk, Trust 1(2):7. 

8, BROWN, L E 1975 The status of the near extinct Houston Toad (Bufo houstonensis) with recommendations for its 
conservation Herpet. Rev 6(2):37-40. 

9, BROWN, W S 1974 Ecology of the Aquatic Box Turtle Terrapene coahuila (Chelonia, Emydidae) m northern 
Mexico Bull. na. St. Mus. (biol. Sci.) 19(1): 1-67 

10, BUDEN, D.W, 1975 Notes on Epicrateschrysogaster(Serpentes Boidae) of the southern Bahamas, with 
description of a new subspecies. Herpetologica31(2):167-177 

11, BULL, PC & WHITTAKER, A] 1975 The amphibians reptiles, birds and mammals In, Kuschel, G, (Ed) 
Biography and Ecology in New Zealand, Junk, The Hague. In series Monographiae biol. 27:231-276. 

12, BULLOCK, D, 1977 Roundlsland-al^leof Destruction Oryx 14(1):5 1-58. 

13, BURTON, J, A, & LAMBERT M. 1977 Tortoises come to Britain. In, Sitwell, N, (Ed.) The World of WUdlife. 
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14, BURY R.B, &RUTH, S,B, 1972, Santa Cruz Long-toed Salamander survival in doubt, Herpet. Rev. 4(l):20-22, 

15, BURY, R.B & MARLOW R,W, 1973, The Desert Tortoise-wiU it survive? Natn. Pk. Conserv. Mag. 47:9-12. 

16, BUSACK, S. D 1976. A review of the biology of the Gold-striped Salamander, Chioglossa lusitanica (Amphibia: 
Salamandndae). Biol. Conserv. 10:309-319. 

17 BUSTARD, H,R 1970 The presentstatusofSeallirtlesandthe Fiji Iguanainthe Fijilslands, Oryxl0:317-322. 

18. BUSTARD, H,R, 1974 The Green Turtle In, Wildl. World Conserv. Yb. 1974:80-87 

19, CAREY M. 1975. Iguanas on the rocks Anim. Kingd. 78(2):15-20. 

20. CAREY M. 1975 The Rock Iguana, Cyclura ptnguis, on Anegada, British Virgin Islands, with notes on Cyclura 
ricordi and Cyclura comuta on Hispaniola BuU, Fla, St, Mus. (biol. Sci.) 19(4):189-234. 

21, CAREY M, 1976, Iguanas of the Exumuas. Wildlife (Lond)18(2):59-61. 

22. CARR, A. & STANCYK, S, 1975. Observations on the ecology and survival outlook of the HawksbiU Turtle, 
Biol, Conserv. 8:161-172. 

23, CARR, A. 1977 Crisis for the Atlantic Ridley Letter m, Mrosovsky, N, (Ed) Marine Turtle Newsl, No. 4:2-3. 
24 CHEKE, AS, 1975, An undescnbed Gecko from Agalega: Phelsuma agalegae sp nov BuU. Mauritius Inst. 

8(l):33-48. 

25. COMMITTEE ON RARE AND ENDANGERED AMPHIBIANS AND REPTILES OF MARYLAND 1973 
Endangered Amphibians and Reptiles of Maryland: a special report. Bull. Md. Herp. Soc. 9(3):42-100. 

26, CROOK, I, & CROOK, G. 1972. New Zealand's rarest frog. Animals (Lond,) 14(5): 188- 190. 

27 CROOK, I 1975 The Tuatara. In, Kuschel, G (Ed,) Biogeography and Ecology m New Zealand, Junk, The Hague, 
In series Monographiae biol. 27:331-352. 

28, DEGENHARDT W G 1973 The Ridge-nosed Rattlesnake: an endangered species. In, Symposium on rare and 
endangered wildlife of the southwestern United States: 104-1 13. New Mexico Dept of Game and Fish, Santa Fe, 
New Mexico, 

29, DESERT TORTOISE PRESERVE COMMITTEE INC. 1975. Desert Tortoise Preserve Chelonia 2(2): 19-20. 

30, DIAMOND, AW, 1976, Breeding biology and conservation of Hawksbill Turtles, Eretmochelys imbricata, 
L. , on Cousin Island, Seychelles. Biol. Conserv. 9(3):199-215. 

3 1 , ERNST C. H & BURY R B. 1977 Clemmys muhlenbergii (Schoepff) Bog Turtle. Catalogue Am, Amphib. Rept. 
204.1-204.2. 

32, FRANZ, R & AUFFENBERG, W. 1974. The Gopher Tortoise in Georgia, Herpet, Rev. 5(3):74-75 (abstr.) 

33, FRAZIER, J 1975. Marine Turtles of the western Indian Ocean, Oryx 13(2):164-175. 

34 FREIBERG, M A 1974, The Argentine land tortoise, Geochelone chilensis, an endangered species. 
Bull. Md. Herp. Soc 10(2):39-41. 

55 



35. GANS, C, & WILLIAMS, E. 1954, Present knowledge of the snake Elachistodon westermarmi Remhardt. 

Breviora No. 36. 
36 HARRIS, H.S, & SIMMONS, R,S, 1975. An endangered species, the New Mexican Ridge-nosed Rattlesnake. 

Bull. Md. Herp. Soc. ll(l);l-7. 

37. HIRTH, H.F. 1971, Synopsisof biological data on the Green Turtle Chelonia mydas (Linnaeus) 1758. 
FAO Fish. Synop. No. 85 (pag. var. ). 

38. HONEGGER, R. E. 1978. Threatened Amphibians and Reptiles in Europe. European Gommittee for the 
Conservation of Nature and Natural Resources Council of Europe, Strasbourg. Nature and Environment 
Series No. 15. 

39. HUGHES, G R. 1973. The survival situation of the HawksbiU Sea-Turtle (Eretmochelys imbricata) in Madagascar. 
Biol.Conserv.5(2):114-118. 

40. HUGHES, G, R, 1974. The Sea Turtles of South-East Africa. Investigational Rep. Oceaitogr. Res. Inst. Durban. 
Part I No. 35: 1-144, Part II No, 36: 1-96. 

41. lUGN. 1971. Marine Turtles. lUCNPubl. New. Ser. Suppl. Pap. No 31 (Proceedings of the 2nd working meeting of 
Marine Turtle specialists). 

42. lUCN. 1973. Crocodiles lUCN Publ. New Ser. Suppl. Pap. No 4 1 (Proceedings of the 2nd working meeting of 
Crocodile specialists). 

43. lUCN, 1979, Red Data Book 3, Amphibians and ReptUes, 3rd Ed. Compiled by R. E. HONEGGER. Merges, 
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44 JONKLASS, R, 1977 Crocodiles m Sri Lanka. Hamadryad (Newsl. Madras Snake Pk, Trust) 2(2):7 

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89. ZWINENBERG, AJ 1975 The westernswamp tortoise (Pseudemyduraumbrina),oneofAustralia'srarest 
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57 



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125 HARDY, G S , & WHITAKER, A. H, 1979. The status of New Zealand's endemic reptiles and thetr conservation. 
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60 



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61 



Index 



Genera, common, and trade names. 



Abaco Island Boa 46 

Abbott's Day Gecko 28 

Achoque 1 1 

(Acinixys) 20 

Acrantophis 44 

Adanson's Turtle 23 

Afghan Tortoise 20 

African Chameleon 32 

African Large-gram Lizard 42 

African Python 50 

African Rock Python 50 

African Savanna Monitor 42 

Afncan Sharp-nosed Crocodile 26 

African Slender-snouted Crocodile 26 

Afncan Small-grain Lizard 43 

Agra Lizard 40. 41 

Agra Monitor 42 

(Agrionemys) 19 

Alabama Red-bellied Turtle 17 

Aldabra Giant Tortoise 18 

Alligator 25 

Almeda Striped Racer 52 

Armargosa Toad 14 

Amblyrhynchiis 37 

Ambon Lizard 42 

Ambystoma 1 1 

Ameiva 40 

American Alligator 25 

American Crocodile 26 

Amethystine (Rock) Python 49 

Ampalague 43 

Anaconda 48 

Andaman Day Gecko 29 

Andrias 11 

Anegada Ground Iguana 38 

Angolan Python 49 

Angonoka 18 

Angulated Tortoise 17 

Amelia 41 

Annulated Tree Boa 45 

Aiiolis37 

Aquatc Box Turtle 17 

Araramboia 45 

Archey'sFrog 13 

Areolated Tortoise 1 9 

Argentine Teju 40 (two entnes) 

Argentine Tortoise 1 7 

Armenian Viper 53 

Arrau 24 

Aruba Island Rattlesnake 53 

Asiatic Cobra 5 1 

Aspidites 44 

(Asterochelys) 18 

Atelopns 14 

Atlantic Ridley 21 

Atlantic Saltmarsh Snake 52 

Australian Fresh-water Crocodile 26 

Axolotl 1 1 

Baby Python 47 

Baghdad Small-gram Lizard 41 

Bahama Islands Boa 46 

Ball Python 49 

Banded Tegu 40 



Barred Python 48 
Barnngfon Land Iguana 37 
Batagnr 16 
Batrachophrynns 1 3 

Batrachoseps 11. 12 

Baw-bawFrog 13 
Beach Cobra 52 
Beaded Lizard 4 1 
Bell's Hinged Tortoise 19 
Bengal Lizard 40, 41 
Bengal Python 49 
Berlandier's Tortoise 18 
Black Caiman 25 
Black-headed Python 44 
Black Legless Lizard 41 
Black Pond Turtle 16 
Black Side-necked Turtle 23 
Black Soft-shell Turtle 22 
Black Spiny-tailed Lizard 30 
Black Toad 14 
Blood Python 49 
Blunt-nosed Leopard Lizard 38 
Blunt-nosed Viper 53 
Blunt-tailed Sand Boa 47 
Boa 44 
(Boa) 45,46 
Boa Constnctor 44 
Boelen's Python 49 
Bog Turtle 16 
Bolson Tortoise 19 
Bolyeria 45 
Borneo Lizard 42 
(Bothrochilns) 48 
Bothiops 53 
Boulenger's Tortoise 19 
Bowsprit Tortoise 17 
Brazil Teju 39 
Broad-nosed Caiman 25 
Broad-snouted Crocodile 26 
Brazilian Giant Tortoise 17 
Brachylophus 37 
Bnfol4 

Burmese Brown Tortoise 1 8 
Burmese Python 49 
Burmese Starred Tortoise 1 8 
Burmese Swamp Turtle 17 

Cabragoya 42 
Caiman 25 

Caiman Lizard 40 

Calaliaiia45 

Calabar Ground Python 45 

Calcutta Lizard 4 1 

California Tiger Salamander 1 1 

Callagnr 16 

Cameroon Toad 14 

Candoia 45 

Cape Clawed Toad 13 

CapePlatanna 13 

Cape Verde Giant Skmk 4 1 

Caretta21 

Carettochelys 22 

Carpet Python 50 

Casarea 45 



Cat Island Turtle 17 

Caymans Ground Boa 50 

Central American River Turtle 16 

Central Asian Cobra 53 

Central Asian Monitor 4 1 

Chaco Sideneck Turtle 24 

Chaco Tortoise 17 

Chamaeleo31 37 

Chariiia45 

Cheat Mountain Salamandar 12 

Chelonia 2 1 

Chersina 1 7 

Childrens (Rock) Python 48 

Chmese Alligator 25 

Chinese Giant Salamander 1 1 

Chondropython 45 

Chioglossa 1 1 

(Chrysemys) 17 

Clelia 51 

Clemmys 1 6 

Cnemidophoms 40 

Common Batagur 16 

Common Chameleon 33 

Common Hinged Tortoise 19 

Common Iguana 38 

Common Monitor 4 1 

Common Tegu 40 

Common Tortoise 19 

Conolophus 37,38 

Conrana 1 5 

(Constrictor) 43 

Corallns 45,46 

Cordylns 39, 40 

Coromandel Leiopelma 13 

Greaser's Mud Turtle 16 

Crested Chameleon 33 

Crested Lizard 30 

Crocodiloms 40 

Crocodylus 26,27 

Croco-Teju 40 

Crotalns 53 

(Crotaphytos) 38 

Cuatro Cienegas Soft-shell Turtle 22 

Cuban Tree Boa 46 

Cuban Crocodile 27 

Culebra Giant Anole 37 

Cuvier's Smooth-fronted Caiman 25 

(Cyclagras) 50 

Cyclnra 38 

Cyrtodactylns 28 

D'Alberti's Python 48 

Dahl's Toad-headed Turtle 24 

(Damonia) 1 6 

Dark Soft-shell Turtle 22 

Day Geckos 28-30 

Deep Spnngs Toad 14 

Dermatemys 1 6 

Dermochelys 22 

Desert Chameleon 35 

Desert Monitor 42 

Desert Pygmy Monitor 42 

Desert Slender Salamander 1 1 

Desert Tortoise 1 9 



62 



Diamond Python 49, 50 
Discoglossns 13 
Dracaena 40 
Drymaichon 52 

Dumeril's Boa 44 
DumerJ's Monitor 42 
Dunn's Mud Turtle 16 
Dura Turtle 16 
Dwarf Caiman 25 
Dwarf Monitor 43 

Eastern Indigo Snake 52 

Egyptian Spmy-tailed Lizard 31 

Egyptian Tortoise 20 

ElachJstodon 50 

Eleutherodactjrlns 13 

Emerald Monitor 43 

Emerald Tree Boa 45 

(Enygros) 45 

Epicrates 46.47 

Eretmochelys 21 

Eiynmochelys 23 

Eryx47 

Estuarine Crocodile 26 

Eunectes 47,48 

Eurycea 1 2 

(Eoiycea) 12 

Exiliboa48 

False Gavial 27 
False Gharial 27 
Fiji Banded Iguana 37 
Fiji Crested Iguana 37 
Fiji Snake 53 
Filfola Lizard 39 
Fischer's Chameleon 33 
Fischer's Tree Boa 47 
Fish Lizard 42 

Flap-necked Chameleon 33 
Flatback Turtle 21 
Flattened Musk Turtle 16 
Florida Gopher Tortoise 19 
Flower Lizard 42 
Ford's Boa 46 
Forest Hinged Tortoise 19 
Forest Tortoise 1 7 
Forked Chameleon 34 
Four-homed Chameleon 36 
Freckled Monitor 44 
Furrowed Wood Turtle 17 

Gaboon Turtle 23 

Galapagos Giant Tortoise 18 

Galapagos Land Iguana 38 

Galapagos Marine Iguana 37 

GaUotia 39 

GambeliaSS 

Ganges Soft-shell Turtle 22 

Garden Tree Boa 46 

Gardiner's Seychelles Frog 15 

Gavialis27 

Gavial 27 

Geoclemys 16 

Geochelonel?, 18 



(Geoemyda) 16 

Geometric Tortoise 20 
Geirhonotiis 4 1 

Ghanal 27 

Giant Anole 37 

Giant Garter Snake 52 

Giboya 43 

(Gigantorana) 15 

Gila Monster 41 

Golden Coqui Frog 13 

Golden Frog 14 

Golden Toad 14 

Gold-dust Day Gecko 29 

Gold-striped Salamander 11 

GohathFroglS 

Gongylomorphus 4 1 

(Gongylophis) 47 

Gophems 19 

Gould's Monitor 42 

Graceful Chameleon 34 

Gray's Monitor 42 

Greek Tortoise 20 (two entries) 

Green Iguana 38 

Green Tree Python 45 

Green Turtle 21 

Grey Monitor 41 

Grooved Tortoise 18 

Grotto Salamander 12 

(Gymnodactylns) 28 

Haitian Ground Boa 51 
Haitian Vine Boa 46 
Hamilton's Frog 13 
Hardwick's Spiny -tailed Lizard 31 
Hawksbill Turtle 21 
Helmeted Turtle 23 
Helodenna41 
Heosemys 16 
Hercules Tortoise 17 
Hermann's Tortoise 20 
Hierro Giant Lizard 39 
Hispaniola Ground Iguana 38 
Hochstetter's Frog 13 
Hoge's Sideneck Turtle 24 
Home's Hinged Tortoise 19 
Homopos 19 
Horn Lizard 31 
Horn Lizard Small-grain 3 1 
Horsfield's Tortoise 20 
Houston Toad 14 
Hydrodynastes 52 
Hydromantes 1 2 
Hydrosaums 30 
HylalS 

Iguana 38 

Illinois Chorus Frog 15 
Illinois Mud Turtle 54 
Impressed Tortoise 18 
Inagua Island Turtle 17 
Indian Egg-eating Snake 52 
Indian Flap-shell Turtle 22 
Indian Monitor 41 
Indian Oval-gram Lizard 42 



Indian Python 49 (two entnes) 
Indian Sand Boa 46 
Indian Sawback Turtle 16 
Indian Small-grain Lizard 40 
Indian Soft-shell Turtle 22 
Indian Star or Starred Tortoise 18 
Indian Tent Turtle 16 
Irapuca 23 

Island Night Lizard 30 
Israel Painted Frog 13 
Italian Spadefoot Toad 13 

Jaboti 17 

Jacar^ 24 

Jacarura 39 

Jackson'sThree-homed Chameleon 34 

Jamaica Boa 47 

Jamaica's Ground Iguana 38 

Japanese Giant Salamander 1 1 

Java Lizard 42 

Java Rock Python 50 

Java Tree Python 49 

Javelin 47 

Jayakar's Sand Boa 47 

Jemez Mountains Salamandar 12 

Johnson's Crocodile 26 

Kachuga 16 

Karoo Tortoise 1 9 

Kavalai Forest Turtle 16 

Kemp's Ridley 21 

Kenya Sand Boa 46 

Kem Canyon Slender Salamander 1 1 

Kiiiixy8l8, 19 

Kinostemon 16,54 

(Klauberina) 3 1 

Komodo Dragon 43 

Kuhl's Tortoise 19 

Lace Monitor 44 

(Laceita) 39 

Lake Erie Water Snake 52 

Lake Junin (Giant) Frog 1 3 

Lake Lerma Salamander 1 1 

Lake Patzcuaro Salamander 1 1 

Larch Mountain Salamander 12 

Latifi's Viper S3 

Leatherback 22 

Leathery Turtle 22 

Lebetine Viper 53 

Leiopelma 13 

LeiolopLsma 4 1 

Leopard Tortoise 18 

Lepidochelys 21 

Lesser Afncan Rock Python 50 

Levantine Viper 53 

Liasis 48 

(Liasis) 48.49 

Lichanura 48 

Limestone Salamander 12 

Lissemys 22 

Loggerhead Turtle 2 1 

Long-tailed Rock Monitor 42 

Loxocemns 49 



63 



Luth 22 



Macabe Forest Skmk 41 
MacToscincns 4 1 

Madagascar Boa 44 

Madagascar Day Gecko 29 

Madagascar Flat-shelled Tortoise 20 

Madagascar Tortoise 18 

Madagascar Tree Boa 50 

Magdalena River Turtle 24 

Maja 45 

Malachocheisus 1 9 

Malayan Monitor 43 

Mangrove Monitor 42, 43 

Margined Tortoise 20 

Marsh Crocodile 26 

Masacuate 43 

Masticophis 52 

Mastigures 30 

Mauritius Greater Day Gecko 28 

Mediterranean Chameleon 33 

Megalixciliis 15 

(Megalsbatrachus) 1 1 

Melanochelys 16 

Melanosuchus 25 

Meller's Chameleon 35 

Merten's Water Monitor 43 

Mexican Giant Gopher Tortoise 19 

Mexican Spotted Wood Turtle 17 

Mindoro Crocodile 26 

Mitchell's Water Monitor 43 

Molurus Python 49 

Mona Blind Snake 44 

(Morelia) 49 

Morelet's Crocodile 26 

Morenia 17 

Mountain Chameleon 35 

Mountain Tortoise 18 

Mount Nimba Viviparous Toad 14 

Mournful Tree Monitor 44 

Mugger 26 

Muhlenberg's Turtle 16 

Mulga Monitor 16 

Muller'sSandBoa47 

Musurana 52 



Naja 53 
(Nardoana) 48 

Narrow-bridged Mud Turtle 16 

(Natrix) 52 

Nectophrynoides 14, 15 

Nerodia 52 

Nesomantis 1 5 

New Guinea Crocodile 26 

New Guinea Plateless Turtle 22 

New Mexico Ridge-nosed 

Rattlesnake 51 

(Nicoria) 16 

Nile Crocodile 26 

Nile Monitor 43 

Nile Soft-shell Turtle 22 

No-mark Lizard 42 

North Island Leiopelma 13 



Oedura 28 
Ogmodon 53 

Ohve Ridley 21 

Olive (Rock) Python 48 

01ml2 

Orange-throated Whiptail 40 

Orange Toad 14 

Oriental Rock Python 49 

Orinoco Crocodile 26 

Osteolaemus 27 

Oustalet's Giant Chameleon 35 

Owen's Three-horned Chameleon 35 

Oxus Cobra 53 

Pacific Boa 45 

Pacific Monitor 42 

Pacific Ridley 21 

Painted Terrapin 16 

Paleosuchus 25 

Palm Lizard 30 

Panamint Alligator Lizard 41 

Pancake Tortoise 19 

Papuan Ground Boa 45 

Panther Chameleon 36 

Papuan Monitor 43 

Papuan Python 48 

Paiadelma 30 

Parrot-beaked Tortoise 19 

Parson's Giant Chameleon 36 

Peacock-marked Softshell Turtle 22 

Pelobatesl3 

Pelomedusa 23 

Peltocephalus 23 

Pelusios 23 

Penang Lizard 40 

Perentie 42 

Phaeognathus 12 

Phelsuma 28 30 

Philippines Crocodile 26 

Philippine Lizard 42 

PhilorialS 

Phrynops 24 

Phrynosoma 39 

Pigmy Mulga Monitor 3 1 

Pine Barrens Tree Frog 15 

Platypus Frog 14 

Plain Lizard 42 

Platemys 24 

Plethodonl2 

Plymouth Red-bellied Turtle 17 

Podaicis 39 

Podocneinis23, 24 

(Podocnemis) 23 

Proteus 12 

Psanunobates 20 

Pseudacxis 1 5 

Pseudeznydura 24 

Pseudemys 16, 17 

(Pseudoboa) 5 1 

Pseudocordylus 40 

Puerto Rican Boa 46 

Pygmy Mulga Monitor 42 

Python 49-50 

Pyxis 20 



Queensland Snake-Lizard 30 
Oueimada Island Bothrops 53 



Radiated Tortoise 18 

Rainbow Boa 46 

Ramgodie 42 

Rana 15 

(Rana) 15 

Red-foot Tortoise 17 

Red-headed Amazon Turtle 23 

Red-headed Sideneck Turtle 24 

Red Hills Salamander 12 

Red Teju 40 

Relict Leopard Frog 15 

Reticulated Python 50 

Reticulated 'Velvet Gecko 28 

(Rhampholeon) 35 

Rheobatrachus 1 4 

Rhinoceros Iguana 38 

Rhinoclemmys 1 7 

(Rhoptiopella) 29 

Rice Lizard 40, 42 

Ricord's Ground Iguana 38 

Ridge-nosed Rattlesnake 53 

Ridge-tailed Monitor 43 

Ring Lizard 42 

Rio Apaporis Caiman 25 

River Terrapin 16 

Rodriguez Day Gecko 30 

Roofed Turtle 16 

Rosy Boa 48 

Rough-necked Monitor 43 

Rough-scaled Carpet Snake 49 

Round Island Boa 45 

Round Island Day Gecko 29 

Round Island Keel-scaled Boa 45 

Round Island Skink 41 

Royal Python 49 

Rubber Boa 45 

Rusty Monitor 43 



Saigon Python 49 

Sail-fin Lizard 30 

Salt-water Crocodile 26 

Sand Boa 47 

Sand Monitor 42 

San Diego Horned Lizard 39 

San Esteban Island Chuckwalla 39 

San Francisco Garter Snake 52 

San Joaquin Leopard Lizard 38 

San Joaquin Coachwhip 52 

San Marcos Salamander 12 

Santa Cruz Long-toed Salamander 1 1 

Saiizinia49 

Sauromalus 39 

Schneider's Smooth-fronted Caiman2S 

Schweigger's Hinged Tortoise 19 

Semi-Java Lizard 42 

Senegal Chameleon 36 

Sepik Monitor 43 

Serpent Island Gecko 28 

Serrated Tortoise 20 



64 



Seychelle Islands Tree Frog 15 
Seychelles Frog 1 5 
Shasta Salamander 12 
Short-homed Chameleon 32 
Short-necked Swamp Turtle 24 
Short-tailed Dwarf Chameleon 35 
Short-tailed Pygmy Monitor 4 1 
Short-tailed Python 49 
Siam Lizard 42 
Siamese Crocodile 27 
Side-stnped Day Gecko 29 
Singapore Lizard 42 
Slender Chameleon 37 
Slender-Snouted Crocodile 26 
Soa-Soa Water Lizard 30 
Socotra Chameleon 35 
Solomons Ground Boa 45 
Sonoran Green Toad 14 
Sooglossns 15 

South Amencan 'Chameleon' 38 
South Amencan Red-lined Turtle 17 
South Amencan River Turtle 24 
South American Water 'Cobra' 52 
Speckled Tortoise 19 
Spectacled Caiman 25 (two entries) 
Spencer's Monitor 43 
Sphenodon 27 
Spider Tortoise 20 
Spiny Chameleon 36 
Spmy-tailed Lizard 30 
Spiny -tailed Pygmy Monitor 41 
Spix's Sideneck Turtle 24 
Spotted Pond Turtle 16 
Spotted Sand Boa 47 
Spotted Tree Monitor 44 
Spurred Tortoise 1 8 
Spur-thighed Tortoise 20 
St CroDt Ground Lizard 40 
Stephen Island Leiopelma 13 
Stemotheros 16 
Steak-tailed Monitor 41 
Stnpe-backed Side-necked Turtle 23 
Sulawesi (Celebes) Tortoise 18 
Sumatra Lizard 42 
Sururucu 52 

Talagoya 40 

Tartary Sand Boa 47 

Tegu39 

Tehachapi Slender Salamander 12 

Tent Tortoise 20 

Teracay 24 

Tenapene 17 

Testndo 20 

(Testndo)17,18,20 

Texas Blind Salamander 12 

Texas Gopher Tortoise 18 

Thaziinophis 52 

Thomasset's Seychelles Frog 15 

Three-keeled Land Tortoise 16 

Timor Python SO 

Timor Tree Monitor 43 

Tomistoina 27 

Toothed Cape Tortoise 20 



Tracai^ 24 
Trachyboa 50 

Transcapian Desert Monitor 41 

Transcaucasian Long-nosed Viper 53 

Travancore Tortoise 18 

Tree 'Crocodile' 43 

Tree Lizard 42 

Trionyx 22 

Tropidophis50. 51 

Tuatara 27 

Tuntongl6 

Turks & Caicos Ground Iguana 38 

Tnpinambis 40 

Two-banded Chameleon 32 

Two-striped Garler Snake 52 

Typhlomolge 12 

Typhlops 44 

Typhlotriton 

Ungaliophis 5 1 
Uroniastyx30, 31 

Vaxanus 41-44 

Variegated Dwarf Chameleon 36 

Vegas Valley Leopard Frog 15 

Vipera51,53 

Viviparous Afncan Toad 14 

Water Box Turtle 17 

Water Monitor 43 

Water Python 48 

West African Dwarf Crocodile 27 

West African Python 50 

West Australian Rock Python 48 

Western Gopher Tortoise 19 

Western Swamp Turtle 24 

White-breasted Side-necked Turtle 23 

Woma 44 

Xantnsia 30 
Xenoboa 5 1 
Xenopus 1 3 

Yellow Anaconda 48 
Yellow-foot Tortoise 1 7 
Yellow Monitor 42 
Yellow-spotted Amazon Turtle 24 
Yellow Tortoise 18 

Zetek's Golden Frog 14 



65 



Lithographed by The Nuffield Press Limited, Cowley. Oxford. Erjgland. 



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