UNIVERSITY OF KANSAS, LAWRENCE. BULLETIN OF THE DEPARTMENT OF ENTOMOLOGY. ALFALFA, GRASSHOPPERS, BEES: THEIR RELATIONSHIP. BY ‘e¢-1¢ "dd veg (‘soquny Aq op04g) “ALNNOO GCYOdA NI NOILOUS-YALAYVNO IVINAWINAdXH NO VAIVATV JO dOUO CNOOUS oe - —"E % MSF ee oe oe re. ew us & > . . > a ea2" 4 rg sd Sees 5 pet a= UNIVERSITY OF KANSAS, LAWRENCE. ALFALFA. GRASSHOPPERS, BEES: THEIR RELATIONSHIP. A REPORT OF THE FIELD-WORK OF THE DEPARTMENT OF ENTOMOLOGY, SUMMER OF 1898. Contribution from Entomological Laboratory. No. 65. eye ee) ~ EN eR. PRESS OF THE STATE PRINTER: TOPEKA, KANSAS. JANUARY 1899, DEPARTMENT OF ENTOMOLOGY, UNIVERSITY OF KANSAS. F. H. SNOW, Pu. D., PROFESSOR. S. J. HUNTER, A. M., ASSISTANT PROFESSOR. HUGO KAHL, CURATOR AND SysTEMATIC ENTOMOLOGIST. MISS ELLA WEEKS, ARTIST. CONTENTS. PAGE PUNE TONE CE IRISTNG HS, eee. e se GS as, Ge LE ea ee Ae MO aiea\s Meerags DoE Se xi TELCO) mcs Sesser ose BASE a arts PAT, Nas G STREP OE O oy Fhe aay crates ohare iv ayersus Mae eer il PART I.— ALFraLFa AND GRASSHOPPERS. APE NHC AIC aCe Lik OL baetee Peete rota eee te oc canst laos Sper oat Sea, Fea SG o “o yaua Oe ORES at Se ay oe 5 RA TLR PAAR RE TEE ce 0 fs ahs cas nhs aye oe ack 9 cae. c5d dep aie RR RA OE EG 9 ee CASE MO PNGES ID MCATIGAR 8s Ki 5 6-2 Saja w sta« kg hai Pos Be platen Le HT Wye COMM LY ares esto cme Ses rmeenc ccs, ois han ers Gino se os eRe: rae 12 IR OIG! COMTI? pee temtechors Detche Geten b aera RCC MIDE rere aeara MAPS e 6 ea ee 13 [RHIAN (GCONLUTTUN AE AiGenG aren Ie Bie COS ERIE a Crear ona a oie ata contra aawe eae a: 13 id areal liguil: COWiall nA Cosette Cees SE CIA OCI aaa es Not coon ame mo nps 14 Bie AE MICE RTIAGGV AIS SoS chee Fos 6 crane rag eR o Metal «Re Wash ace nia ny ee aut ee 15 VAY nile Grey GOT Un IN tetieet oes ore ee erte oor De eee a oie eC nen ae 16 JG feAR al COLE INT ets Gece IE Hee ie ced eS Tse CCLRC Te oe ona Snir o cet 16 TM GVOTRAE NS CORTICES S aya, Peer ena cee e naee Soest ee R ONES ciel hc MCerite = 16 Rese RUAT EAR CRTER IL GV NS beget oe Shas ost ste giant cin SR eas Date OD LA 16 OMe ed TAN AY AC EMR IER Vata Ale nts ie ick Tie Tape bit a 9.3) anahe dS SARS thet ae nee oe tee 18 NOT GOUNCO UMM V eerie a nay ete Lae erate bie Sle RTD eda MEN ee ee 18 fsmemied of ‘Grasshoppers. © revalents:.-.). 2... 2 6s 26k nk ooo se Racecar 19 bife-Eistory.of, thew itherential Wocust... . 2. 22 2he4cseeaeee tele cec news 20 Tal Tani] OPER ON Layepy A eh ary a rak is aia renee ade A rare eR ree NO 2 20 DISSES UN RTO 0g 0) ie 10 isl 1200 Ra NR a ere rete ce yt ok 20 METH OGTO ONUPOSIUION Gop.) rts oe ee cease APN eters, mae eae ae ee 21 Wiheresthererostane lal Gti. fans) ic see tater oe eee eee 23 Rebea nae aR rete he Bic ins ies ok SAH esta ks RIG U I ao ee 27 Ucar ee eye SB ceeeatO Gee oc ania ow es ee Ba Oe 28 ROOM Mee) 58c Beek AIS, se docs Wi tals Peo ELAR ME Soe ES OL ee aero 28 /AQUITON OS Rh as Aas eS ca ONG Sen oe Reece Aine RE De ene eee et a wire te A CE 28 Bordersiotsthe. teld stripped. an. no sa sete aee sone eee ee 28 Gerson eGR, oye. oOo ois va Ga ae Soe WANES by oe oe ee eee 29 | TENS Oe oe, lt ane ee ee eR See Bek ings ear ae SEEN 29 NUR en ee LE nat b..0 3 bak Sado ae ote Oe o sae ats Wee ee ee 29 GIS O48 a ee a eR RT eA OO eR te, 29 pia rntes Soon ctya Alt ooG as Lok, asin ROIS Gee en ee 30 BPSUSEMBNAKGSS 10 G1 J -o rt. ve8 oe ses Oe eee ose haree ne ee ee 30 MRE RUC 2/5 3, 20° 5.5'3 5 5.08 a 2d WOOS L Se hee Fake eek te eee 31 Mamie MELONS 3s foe ct Pou ees Sek Re ee eine ee ete 31 Reatstninte tM rer vic, stint Sc oe oa ag Bete code wear hee 31 Be tee Spare C hc whe styl Ok i elk EU AO. 51, Ak At A 31 Parasinic: milgsences On-welanoplis. 6.2 oo. wn Sods a tere eae 32 TCU PR MIC CLINOIGUR CN soca ee kiana reece kent the tens 33 WOM F CUI ET VENET CL PECNELE Ds in scorn vi aoe, wep baie in ecareiaietane hatenels eee a 34 1 USUI TEST 07 1151 |S Ry Ook eet RI S aiaae gh cet ebay Sn eae gla 37 Observations upon the Differential Locust made in widely separated LS LANES S elie 8 ant ae grea ais eS DR a eg Ae 2p AR i gle Sh ea 38 vi TABLE OF CONTENTS. Grasshoppers in General— Continued: Anatomy io the Diirerentialwbeocust:«-. sae nase moe ea ae ae reas 39 Hixternal anatomy <2 6.00.5 .306+ ones ok A” A a A a 39 ABTS LOMEL Roto ls Foie kee ines Sane cred Stn ERE UAE Sew Edel e) ie 40 Phe Chorassee ee see e te ee On ee eee io beh ry 41 Ap pendar es Ol thelhmonax. ene: mie ates seein ote oneeees 42 SHE sp CRONNLOU ta 2 for eee er or eae Oss ae ik Pa ee RES he. un I 43 Initernalbanatomiyers sac seo aso store ere acne erate shar oan hore ch hey tee eee 44 PE OSLIVeASHStOMl. «hohe tee sehi nce aie sink Pi dae Duelo ie hs Dian aienceee motes 44 REGINA OCW RY LONE cache se oi Sea = ies a ees Wei eles ok at a 45 NGS PLEO LY SV SUCIA ems Sr lck on aca deinl On CLIN yeah aan ae ee Rr ae 46 Reprogdiuctiversy stems sachs eee hoe tes Re isi eee ee eee 47 INGTVOUSMSYSLCMNS scot ei eee ocd. Ctra ss ed aries Ss eee Ae eR eee 47 Measirestofabrevyention ..2 ssc. ose ae aes ee te ener 62 Technical description of the Differential Locust......................... 63 PART II.—ALFALFA AND BEES. TNO GNC HOM misters tices aseos aie ace RSES ich ve ee EES OL ON Re Reel es ke eee ee 67 Will OLIESIOR DISCOS... = rors a ancrs Serene axes RPNEIR toe Sa Ieee eS eee ee 68 (Germianie ses acct a ai Uher sua) chao’ & seu, te fs Wha shee be dena ay eat LOR Ra OREN aR SUE So 69 Warniolany .co.c.e ves e cbcak onsale oon ae ee eee ee 69 1S) ND eee Re Siete es SRN Manon Preis eae te ard Te ene Rea eS oo ao yal (Gayoink ieee Stee oO ani Cr, Reise shite n G Bing Mona ooo bac geoe 71 Socraleheonomy-ofsthe Hivess: ae sci Bete hc Cert ema ae ae eee eee 72 AME OCOD Ris nite oO he eus sessile oS vine ne aks calor Pe eis eet Renee eee 72 WINES CUNO TTC aie. ko cues) si eae vata ai nveloue eit eeear ls Sieur foes SRO ae ae 73 PPS R WOT OT cc srer secs, ti PAssake ekg Saye ats rake I ae hs SPST nee non a a ie BECHER ROGIICTS : Fics ats She stir sicksrah « pele b asieet Shay Sdtecwiahs al ae guste ys ts Be at be RRS ete eee Oe 74 IBY-YSE| 0} 12216 Cera ee eee ete rr ar enn eee Wat uence en wid on A wa oe 74 IPFOPOLIS Si aic cis. cals crab cssdeabis She seca age ed asada a ewe CE ee ee ee ee 74 Royals elven. oath oy heccor- asa eee eran eee Oe eR Ea ae eee 74 SIU Cea enn ae Seer mbites Seren nee ee E NE Fer ME NS Oe ON Go sane 75 Vitae sere) yao SRG Seosts S eesine PD sa cB NG Wa sv ehe epee eek a ra eS Ot or Po a 75 LE Kole \ OME Orme an et ere BMI EAA Sorina tanner cee coca ones 75 Analyses of various ieee OF homey ncss eas ances ok ene ee eae 79 Gioyns) op OLIN Chin) Un ae Weer ea Neen meen Onn SU betnrn Osea desde 5 sce 79 f blo Veak<) Hh oleae ee er ee See Ne re emer Ie oer er yao otros. a cont sere 80 Method of fertilization of the/alfalfasblossom:.-.. 2-52.51 ee ene 81 Infiuence:of-bees upon the-seed! crop.e.ia. 2 orca. oe eee een as 82 Comparative tables: oo 3 siae Wlecg pees nate deen pene ete Nene ete 84 Adfalia:as‘a.honey plant? i. sc iacs i. Se as aiaeieens eet Bes ele ete Oe EI ene 84 Comparative tables showing alfalfa acreage, stands of bees, pounds of |a10) 012) Aa ene ee etre ee iar Serer mentee 5 ee ia Sina o he oso ch an. 86 Observations «athered from) Wansas Aipiaristsen casei seein erie 88 Sone, ive, Viel sis cia eeic oy acne shee aero Le Ite oo aaa rene sats 98 Record of colony of bees feeding for the year almost exclusively upon alfalfa, 100 TABLE OF CONTENTS. vii tn an INI AGU OGS 5) kw alc sclld tig mak ce a boon wn va ole Sele be a eae epee 102 Witatneons tines ta COLONY OL WOESs:...j. 24 s%ies ccies oe es ele de wie sb hie bela elelene 102 inp mree aie AP OMEN G (seh Te 65 isla Giles oi poe wiacew mean ewidaee aes 103 Stimulative feeding................ gpl Ay -ites ed OR re 408 Pee SATA RS eT eee 105 puiemnecRVvelnanGusiMmOkels yo: uliter nes prea sil 6 Zareryshac artis trees femontare 106 REINER TENIIS,. 2. alice SNE EE Bin HOw! Roe wis Hed Sim Rance: Soave Nee a hee 107 lave secure apa hanl COS a5 526, ices 4) afc) arsiocly Pleven a, he do seccelamio ees «cing Weenie ne 109 The Parker foundation fastener.................. Pie Atte eg ht yi, Beha ng 111 Bnet or Molding Tie SOCLIONS:). .ss.0h 5 9p sw ok kos vind Sales s Ske eta pee, whe 112 MMe sec iON -NOlderMSUPELers sere aeons chien ere ae fala oe Pietersen Siecle clits 113 Sec in OOkes, SHIppiNe CTATOS, CLC. oi 7¢ 250g. 8.2 he aul Oe ete e tles Guere ninete 114 TET Racist Condi) OVE Re voras wee, See ena ree ees ees eC aa Oe ea eae See SR ar 114 ADAMO Said CL OME tees yarn eet cyetaiar aicie tra cape ca cut Tava e e-ons n'a odisece arene ero odva aureeler sca menaes 116 PPG GULP MMOME Vrs sree kt ciaisaitcts eiaiersg Ce Sem ae wd oie © Sele tie tine nee ae tenes te 118 EMM oS UDR Ae Rene tas ee ey Sota n PR ecards cist «, Hlavnlonnere ais eheieasic ee avis ete ssete 119 LE UL PEN ER Octet 1 eo ee nee MES 121 Oneen-cellsrandequecemMearin Gs, oso, fs erase opalel “tSelancvaisis evened Melanoplus differentialis killed. by fungus; ........:- 222-2 aoe eee 38 12. Front view of head of Melanoplus differentialis with clypeus and la- EWM: TeMOVed:.< fies. bibs ate ine ea tage ie ties yee ee 40 13. Front view of head of Melanoplus differentialis with mandibles SPLOAG OU ie fe5.6.Fe cw csis oars wlegn a rete sioteh otthe xed ser) Sees enon oie ee 40 14 Wabium of Melanoplus Quper en cigs ae ceei a eee: nine eee ee 41 15. Inner view of maxilla of Melanoplus differentialis................... 41 16. Clypeus and labrum of Melanoplus differentialis............-025 ses 40 163. Side view of prothorax with leg of Melanoplus differentialis......... 41 17. Side view of thorax of Melanoplus differentialis..................00 41 18. Metathoracic leg of Melanoplus differentialis ..........0 ccc cece eens 42 19. Dorsal aspect of body of female Melanoplus differentialis............ 43 20. Ventral view of thorax of Melanoplus differentialis ................. 44 21. Side view of abdomen of female Melanoplus differentialis........... dt 22. Side view of abdomen of male Melanoplus differentialis ............. 44 23. Dorsal view of caudal appendages of male Melanoplus differentialis, 44 24. Exterior view of auditory organ of Melanoplus differentialis......... 45 25. Digestive, circulatory and nervous systems of Melanoplus differen- HOT SE ore SRSA AONE PITS SACRO SON Om UD n Ma cok SIS ols19 o 45 26. Respiratory system of Melanoplus differentialis.............00..+5: 46 27. Side view, showing spiracles of mesothorax of Melanoplus differen- CUGUAS ceo i fds Bae era ie Ot eT ee OT Car ent cnn 47 28. Reproductive system of female of Melanoplus differentialis......... 47 29% sPlanof construction.of hopper-dozer.... + -erc)aseeie eee eens 57 30. View in front of hopper-dozer, showing quantities of grasshoppers just taken from ‘the pang aye... 5 vars edie 2s elieuelles eee 2) aoe hsp eee epson eet 58) ILLUSTRATIONS. ix PLATES IN PART II. POOLE MINUS A IEUEI OS ce, = ate Sarai tre etets: eiisisiarere-o oars tidhe levers sce Frontispiece. SUraRterePaRE DIGS BE Lis pre eee oe ae eeSeeey Be wale aid a dn Seu aelabr utente 70 DIB OSI IVEgs VS COMM OLW DEO rcs rosso utiee ciciee Seed oie cg ete noi ates aes ONES 76 Shades of color in various kinds of honey........................008: 78 FIGURES IN PART II. Sh GLE UC MPOCE, TD Pe gsc) 21 Nc ae 75 The sting of the worker bee and its appendages....................... 80 Polinamonorn alfalfa: blossom by beG.. 92/6... £3. a6 66550 os Cacland weeres widens 81 Mester oO. alralia with bee Tecate |... ck cae ed Same w an when onstne 83 ARIE MineL Meo atee:s 2 Sasha de oetee iden Lacon didn. 6 LSS Veins Sil Bed eiae eee 90 ERE ECE IDE CLOVES) f 2 6) teers Soka so cepa ed anc cv awe ee a eee 93 Brtiew MeO ON AWERb.ClOVED S520 on Jeph 2GGs oe owe eee etek as ea cser snore ders 96 RRM CERES ITO ER f 65 sc 8. 3h) a Ss Sees sia ett we was bod sot ie es ae a ae ee 107 Hives and appliances for honey production.......... ea ts ican eee 109 Ee ME eC OEPIDEM EO IE TASLOUOT sty oc bios wield, eax c.8 dua, preaieoa Sm aaa ls Sage Selon ae 111 euipee. for naldin® bhe sections 2 2. oS ..s 65. Wie kins ncioares elev eos Fees os 113 See erR MAE NPANGEETE NAOT re S07 Gye ices Se c/n. See's shal ea OR wee Sle are heals Var Oe 114 Becwon ewes, Shining Crate, ChE. 26 6d. ce nc as 0 as eae bs edad vn went 115 TELS Coit OER Stes bn on eee hem ci a ied Re i ee Lone ee Ee aad SRA Gee ese ra 115 Extractor with inside gearing raised and exposed to view .............. 117 Agi etal le SMPTE eS or ae 6 oe Cathey cheks Wieyatn, ie aR. Sal iach TS De AME tre wate 118 GEA ETA Beg Cena Te Ree fone Aaya. Lehane Soaks WEA, ae REE 120 ATED LEWES NOME LIAR reels case Fae wna Grek to ance ea Pike PE woe ae 122 Ren -COlls am GUee TEMPIN OE. .. =: os.g.: eos oes uruk te Maina Medison dew 123 Pranic OUCH pOUDEU AION: .<.)-0 2. 574 wana 2a hae OA oleae es ala te aeented 125 BSCE ee ey iia cr nih dan. ING Deine en. Teas Sle so ios See we SR Seslaresaahe 126 JE IGRUTE SOV ER ETESIO, abet Sis ted e she Gates AOE ie RMN 20 Mon OG nna dead Bi Toon ee 127 Openinehives.ana Handling bees 4... onc 5 chose ns ovate ase aoe 130 RSESE METS MIC UOCIIMIN TS eee) oes ays cach ars acme: NS tea ae wb te aad ie ee 131 MOMS SVIEW-OF HOUSOPIATY .. ss 2 2 a Soca bewath Ueda She eee eae eee 132 Re Blete WHET OLA Bert Ache Sotto Sx d's baci e eae AEG Oe EME es Bees Coles 133 Chat hives.and outdoor: wintering... 2.2.06. Skee. 2 sens actos ae eee 134 SEA SAVE CORA GUO > 37g es ara 2. co lyab ¥en bis lo Act Ue estonia ee ae 135 tmenyar pesd-teady tor Shipment... |). so aco wen suas eee 136 . any es ie tie f 7 Cav re oe daa co Rie Des: cet Ae ee a : “5p Ae >t ee “> ee ; ace > he a, | nt ane ira he « * - ACKOWLEDGMENTS. Referring to Part I, my thanks are due Mr. C. E. McClung, of the Department of Zoology, for the sketch from which figure 4 was made and for the paragraphs upon the egg of Melanoplus differentialis. Plates I, V, VI, VIII and figure 1 are from photographs taken by the author for this publication. Plate VII and figure 30 are produced from photographs by Mr. Earl Cass, under direction. Figures 2 and 3 are from photographs made to be here used by Mr. F. E. Marcy. Figure 11 is from a photograph taken by Mr. E. 8S. Tucker in this laboratory. All the other plates and figures are produced from original drawings made by Miss Ella Weeks, under the author’s direction. In the execution of this work the zeal and fidelity of the artist is worthy of special mention. In will be noted from the above that all the illustrations in Part I, both text and plate, have been prepared expressly for this volume from original subject-matter. Referring to Part II, I am indebted to Prof. W. C. Stevens, of the Department of Botany, for directing the work of the artist, Miss Weeks, in the preparation of figures 3 and 4, and for the use of the photo- graph, taken by himself, from which figure 7 was made. Plate I is made from four photographs. Plate II is produced from original drawings made by Miss Weeks, under direction. Plate III is from a drawing made after Cheshire. Plate 1V—figures 5 and 6 are from photographs made by Mr. F. E. Marcy, under the immediate direction of the author. Figures 1 and 2 show in the explanations where taken from; they were kindly loaned by Chas. Dadant & Son. Figures 8 to 29, inclusive, are from photographs taken under the author’s direction and with the codperation of Mr. A. H. Duff. There should have ap- peared in the explanation after each figure the word “(Original).”’ With the exception of plate III and figures 1 and 2, the illustrations were especially prepared for this publication from original drawings and photographs. (xi) PREFACE. F the products of the season’s labor are damaged by an insect foe, the cause of its appearance can be ascertained and means of pre- vention made prominent factors in subsequent calculations. While the insect tribe contains some foes, it has among its members many insect friends. The grain producer and fruit-grower will do well to cultivate their acquaintance. It is with the desire that intelligent action may be directed against one of our aggressive herbiverous insects, and a more intimate ac- quaintance formed with our melliferous insect friend, that this publi- cation is sent forth. Paradoxical as it may seem, the former insect when properly dealt with proves a blessing in disguise. The latter ealls upon the alfalfa blossom for its nectar to enrich our table, and by its visit increases the seed crop twofold. It is expected that experiments begun one year ago and continued this summer will be carried on during the coming season, and infor- mation upon operation and results will be given at a later date to those interested. In the preparation of this work I am indebted to Chancellor F. H. Syow and Dr.S8. W. WILuiston for valuable suggestions. During the field-work, a report of which this is, an expression of my appreciation is due Supts. G. W. Watson and J. H. Smart, of the firm of Batt & GoppaRp, Kinsley, and Co. Treas. C. H. Hum- PHREY and Hon. Tuomas H. Forp, of Syracuse, for valuable assistance. In behalf of this department, grateful acknowledgment is made to the Atchison, Topeka & Santa Fe, Chicago, Rock Island & Pacific, Union Pacific and Missouri Pacific Railway Companies for material cooperation. A ee Rage ieee SR SS SPS ee F8 ir re ae ON oe eS as. . ; Spee wee ae ee ne fy te ee went eee PART I. ALFALFA AND GRASSHOPPERS. INTRODUCTION. YHE culture of alfalfa has reached such proportions in Kansas that those directly interested are devoting their energies toward making the returns as large as possible. Farmers in some parts of this state are counting their alfalfa fields by the section plats. Whenever such areas are devoted to one particular plant the course of natural events is changed. Nature intends that there shall be an equal distribution of plant life and animal life. Wherever one plant predominates those forms of animal life which thrive best upon it as a food-plant increase in undue proportion. This has been the case with the chinch-bug and wheat, the corn-root worm and corn; each have thriven amidst superabundance of their favorite food-plant. That the relation between alfalfa and insect life might be more fully understood, the writer was called upon to visit Edwards county last fall. He then made note of the conditions, and published, in connection with Professor Snow, a report giving methods of cultiva- tion that would eliminate objectionable insects from alfalfa lands. He was asked to continue these studies at the beginning of this year again, and, with the proffered codperation of those interested, to conduct such investigations and experiments as he saw fit in the alfalfa regions. In acceptance of that request, believing that the best laboratory was the field, the writer went into camp fifteen miles southwest of Kinsley, on the line between Edwards and Ford counties, with tent and full laboratory equipments, prepared to study the entomological conditions of that region with special reference to the alfalfa plant. My first work might be termed a general survey of the territory adjoining the camp. Accordingly, in company with Supt. J. H. Smart, I drove over the alfalfa-producing region of Edwards and Ford counties, and obtained a general knowledge of the situation. I was soon struck with the prevalence of one species of grasshopper, and also the possibilities of bee-keeping in this region, an injurious and a beneficial phase of entomological research. To these two subjects I determined to devote my undivided attention, to the exclusion of all others. While at work here invitations came to visit Finney and Hamilton counties, where grasshoppers were making inroads upon the growing (5) 6 INTRODUCTION. crops. Accordingly, upon the 18th of July I reached Syracuse, in Hamilton county, and visited agricultural regions where grasshoppers were at work. Upon the 20th and 21st I investigated affected fields on the north, east and west of Garden City. I returned to my camp upon the 22d. Being convinced that the damage by the grasshop- pers was due to the prevalence of one species, I concluded to devote my attention to the life-history and habits of this one. Accordingly, observations were largely confined to this one species. The camp was maintained in Edwards county until this form had reached the adult stage, when, having already received reports of flights of grass- hoppers on the western line of the state, and also of the prevalence of grasshoppers in certain localities, I determined to make a survey of all this region, to ascertain the numbers present and causes govern- ing their presence, and also to visit the beekeepers of this region and make such observations upon the actions of their bees toward alfalfa as might seem practicable. Accordingly I left camp for Syracuse, and from Syracuse by team drove north through Hamilton, Greeley and Wallace counties to Sharon Springs; from Sharon Springs by rail to Oakley, where conditions were noted, and from Oakley by slow train to Colby. Conditions were again noted from Colby to Goodland. Reports had reached me from this point of the prevalence of grasshoppers on the tracks. Here, I made special examinations of the cow-catchers of engines entering the roundhouse to ascertain the species of grasshoppers caught along the track. Upon leaving Goodland I was given the op- portunity to occupy a favorable position where flights of grasshop- pers rising in front of the engine could be easily noted. This observation was continued as far as Norton. At Norton, the agri- cultural regions were visited, and the existing state of affairs noted. This survey covered eleven of the twenty-four counties west of the 100th meridian, and two east of this meridian. The pages which follow contain the practical results of the obser- vations made, extending over a period from July 5 to September 4, as well as a fully illustrated summary of the experiments conducted with the codperation of the farmers during the past year. The hearty manner with which all interested have entered into this work, and the many opinions expressed concerning the favorable results, cause me to believe that the work has not been in vain. Since my return numerous inquiries have been received, examples of which are here- with given: Vera, Kan., August 27, 1898. My Dear Sir: I read with much interest the article in the Kansas City Star of August 19, 1898, about your experimenting with alfalfa by disk harrowing, etc. I have quite an acreage of alfalfa that has not done very well this year ow- ing to the wet weather, with perhaps other causes, and the crab-grass and sand- INTRODUCTION. 7 burs have come in badly. I would like the benefit of your experience as to what is best to do; but let me state first, there is a good deal of alfalfa on the field. Now please advise me whether it is best to plow up the field this fall and sow in the spring, or wait until spring and rake off all the dead crab-grass, etc., and disk harrow the fields, sowing where it is needed, or, after the heavy frosts of October, this year, will it be advisable to burn off the crab-grass? If that were done the alfalfa left might winter-kill. Is there any probability that the disk harrow will cut the crown off the alfalfa and kill it; did it kill any in your ex- periments? Would it be advisable to disk harrow the ground in June after the first crop has been cut off? How much of an angle do you set the wheels of the harrow when doing the work? If you will kindly give me the benefit of your ex- perience in the matter, you will greatly oblige me. Very truly yours, ALBERT F. THAYER. Leroy, Kan., October 21, 1898. Dear Srr: I notice in the Capital that youadvise farmers to disk and cross- disk alfalfa fields to rid them of grasshoppers. I have four small fields of alfalfa, sown in 1895, 1896, 1897, and 1898; the grasshoppers are stripping them in spots. I am afraid to use the disk without some instructions. Is there any danger of setting a sharp disk at such an angle as to cut off the alfalfa roots and kill the plant, or is there any danger of disking too deep? Hoping you will be kind enough to give me some light on this subject, I am, Very truly yours, Joun H. Sicssy. P. S.—How old must the plants be before it is safe to use the disk on them ? Sr. JosepH, Mo. Dear Sir: I have over 500 acres in alfalfa [in Norton county, Kansas,] and intend raising that number of acres as soon as I get the ground in order, Some of this alfalfa is over six years old and some was put out as lateas last spring. lb believe that the oldest of my alfalfa yields the most, and I think that it takes at least three years before a full crop can be harvested. The grasshoppers favor al~ falfa fields on account of the tenderness of the plant, which grows so fast during the season. They certainly prefer alfalfa to corn and other cereals, and only leave alfalfa fields when they are destroyed, and then take to the corn-fields or- whatever other cereals adjoin the alfalfa fields. I believe the conditions concern- ing alfalfa appear to be conducive to the multiplication of grasshoppers, as they generally lay their eggs there before leaving. I would like to have you advise me. what headway has been made toward destroying the pest when it is small, and if you know any implement or tool made for that purpose, or have any idea to pre- sent, I will gladly receive it, as I am anxious to save the second and third crops of alfalfa. They do not bother the first crop, as it is generally ready to cut before the grasshoppers do any damage. Kindly let me hear from you at your earliest convenience, and oblige, Yours truly, A. J. Brunswia. That these and many other letters might be clearly answered, and those concerned be informed more fully upon benefits not yet enjoyed from alfalfa, this publication is sent forth. It has been deemed convenient and wise to present the subject in two parts; the first will treat of alfalfa and grasshoppers, the second of alfalfa and bees. It is highly gratifying to state that the conclu- sions of Part I show that the best methods of alfalfa culture are those 8 INTRODUCTION. which are the most conducive to the prevention of grasshoppers, and that the deductions of Part II show that while alfalfa is one of the first of our forage plants, its choicest product is not to be found in the hay-rack, but upon the farmer’s table, carried thither by the busy bee. Dice: webs JANUARY 9, 1899. GRASSHOPPERS IN GENERAL. N the minds of some there exists an opinion that among states Kansas alone has suffered most, is most subject to injury from grasshoppers, that these grasshoppers are migratory, that there is no means of checking them—hence, they are a necessary evil, and the less said about them the better. For the benefit of such, as well as for general information, it may be well to give the situation as it is. Since the Rocky Mountain Lo- cust is the most renowned, we have placed in brief form, as taken from Riley, Bruner, Packard, and others, the data following, showing number of annual visitations the Rocky Mountain Locust has made, and the states which these locusts visited. These insects doubt- less occurred frequently prior to dates given, but no records were retained, since these regions were at that time uninhabited by record- keeping people. The record from 1866 is full; the completeness of the record from 1866 is due, not necessarily to greater prevalence of the locust, but to better facilities for securing such data. I have placed after each state here named the number of annual appearances of the Rocky Mountain Locust within the twenty-eight years between 1851 and 1878. Before 1851 we have no authentic records, and since 1878 this locust has committed little damage in this state. For this reason this period has been chosen. Number of annual visitations of the Rocky Mountain Locust to states named, during the period 1851-1878: Arkansas, 2. Kansas, 14. New Mexico, 3. British N. America, 10. Minnesota, 18. Texas, 15. Colorado, 12. Missouri, 8. Utah, 26. Dakota, 17. Montana, 18. Washington, / 4 Idaho, 5. Nebraska, 13. Oregon, ie Indian Territory, 5. Nevada, 4. Wyoming, 10. Towa, 15. While Kansas has been free from the attacks of these insects since 1878, it has not been so with some of the other states named, notably Minnesota, where, in 1891, the crops of several counties were com- pletely taken by this locust. From a glance at the list, it can be seen that, up to 1878, seven states had a greater number of visitations from this locust than Kansas. Further, it may be well to note that there were only three seasons in Kansas when this grasshopper’s damages (9) 10 GRASSHOPPERS IN GENERAL. were severe or general. To these facts add the statement that other states have been visited since the migratory grasshoppers made their last pilgrimage to Kansas, and it can be readily seen that Kansas by no means stands first nor second, but stands eighth in rank as host for this far-famed locust. This list does not take into account the presence of native grass- hoppers. These have made their appearance at times in destructive numbers in all of the territory named in this table. Not only over this western territory, but the New England states as well have suf- fered from the voracious appetites of resident locusts. Grasshoppers are spoken of historically in the East; they can be made “past tenses” in Kansas. The purpose of this article is not to treat of the migra- tory locusts, but those which flourish and die in the vicinity of their birthplace. Grasshoppers belong to the order of insects known as Orthoptera, or straight-winged insects, such as the cricket, katydid, and praying horse. This order is divided into several families, of which the Ac- rididee or grasshoppers form one. Since the subdivisions or subfami- lies are classed according to habits as well as structure, it might be well to mention them briefly, so that the casual observer may know from the insect in hand whether or not it is a highly injurious form. One group, found more generally in low and marshy land, is char- acterized by a very receding front, forming a sharp angle at vertex of the head. This subfamily, Zryxvalinw, is not abundant enough to cause serious damage. Common along the sides of roads and field is a dusky grasshopper, showing as it flies a dark hind wing with white border. Frequently it pauses in the air, flitting its wings before alighting. This is the Carolina Locust (Dissosteira carolina), or “dust-hopper,” so called from its fondness for roadsides and dusty places. This familiar insect belongs to the subfamily @dipodina, a group characterized by rough prominent ridge on crest of prothorax, the hood-shaped part immedi- ately behind the head. While these are injurious at times, they do not, in this country, possess migratory instinct to any great extent. The next group, the Acridzinw, contains our most injurious forms. The prothorax is usually smooth on top, the breast between front legs usually extended into a tubercle. To this group belong those grass- hoppers that migrate, and those that thrive well on cultivated crops in the locality of birth. GRASSHOPPERS IN GENERAL. it NATIVE GRASSHOPPERS IN KANSAS. The native grasshopper has made himself felt in Kansas, and is lia- ble to do so again unless he becomes the subject of intelligent attack. In a trip for observation, Professor Osborne, in 1891, reported these grasshoppers as causing considerable damage to crops along the Ar- kansas valley, in the western part of this state. Since that time larger areas of cultivated ground have furnished more food and the possi- bility of a greater production of resident locusts. This matter was brought directly to the notice of the writer, by letters received in Au- gust and September, 1897, from Supt. Geo. W. Watson, of the land department of the Alfalfa, Irrigation and Land Company, Kinsley, Kan. These letters were referred to us by Mr. F. D. Coburn, secretary state board of agriculture. From the nature of this correspondence, it was thought advisable to visit that locality, to obtain, in detail, ex- isting conditions. Upon returning, a report was issued, by Professor Snow and myself, an extract from which I herewith give, to show the state of affairs: On the morning of the 30th, Mr. R. E. Edwards, president of the Kinsley Bank, took us to a large tract of alfalfa southwest of Kinsley. There are about 400 acres of alfalfa in this piece, all the property of Mr. Edwards. Here we found a few of the Rocky Mountain Locust,* but many more of what is known as the Differential Locust, or Melanoplus differentialis. The farmers speak of them as ‘‘ those big yellow fellows.’’ Mr. Edwards stated that the grasshop- pers had not been so injurious in that tract as farther west. Yet wherever the alfalfa was young and tender from recent sowing nothing appeared above ground but stubble. Where the plants were older and tougher, the damage was not so noticeable. In the afternoon and the next forenoon we visited a number of alfalfa fields, and found conditions much the same as upon the first day. In the afternoon of the 30th, Hon. A. C. Dyer, county attorney for Edwards county, took us to a locality west of Kinsley where the damage had been great. Here the small Rocky Mountain Locustt was more abundant, with a goodly number of the Two-striped, Melanoplus bivitattux, but, as before, the Differential Lo- cust was by far the more abundant. Adjacent to one piece of alfalfa there had been a piece of corn which they entered after the alfalfa had been cut and com- pletely stripped it, killing it before tassels had appeared. It was not, however, for the purpose of observing the amount of damage done that the visit was made so much as to note existing conditions. These will be discussed farther on under the head of ‘‘Alfalfa and the Grasshoppers.’’ Here we will say that everywhere we were glad to observe that of the Differential Locust (M. differentialis) there were apparently as many dead asalive to be seen. Nat- ural enemies were at work, but too late to be of any assistance in preserving this year’s crop, though certainly of benefit in curtailing the number of eggs which * This was in accordance with a determination made for us. Specimens after- ward sent by request to Mr. W. D. Hunter, Nebraska State University, were cor- rectly determined as the Lesser Migratory Locust, Melanoplus atlanis. + These were afterward determined by Dr. Hough as Sarcophaga cimbicis Town. 13 GRASSHOPPERS IN GENERAL. would hatch to destroy next year’s growth. In many cases the cause for death was apparently the work of a fly which deposits its eggs upon the back of the grasshopper. The larvae —small, whitish, worm-like creatures, or maggots — soon find their way into the interior of the grasshopper and produce the death of the insect. These larve were found in a number of the specimens taken, and later two of them changed in the laboratory to adults. They proved to be flesh-flies (Sarcophaga sp.) Many of the ’hoppers had been deprived of one or both wings —the work of the locust mite. As previously stated, by the request and with the codperation of a number of the leading alfalfa growers of this locality I resumed my investigations upon this subject on the 5th of July. That the situa- tion may be more clearly understood, it is probably better to give by counties the conditions existing in the territory studied : Epwarps County.—In the immediate vicinity of Kinsley no great damage was done. A drive over the greater part of Edwards county, in company with Superintendent Smart, showed the greatest damage to be about fifteen miles southwest of Kinsley. Here large areas of alfalfa are grown. There were 3000 acres in one tract in the immedi- ate vicinity of my camp. The first crop of alfalfa was being cut, and by this time the wingless nymphs were moving into adjacent corn- fields. The numbers were by no means general over the entire quarter- sections of alfalfa, but appeared to be very thick in spots. Lying on the south of one quarter-section which was being cut was about thirty acres of good corn just beginning to tassel. Insects moved into this, and instead of eating the leaves they ascended the stalks and ate the tender young tassels. In some cases where the ear of corn was shoot- ing they devoured that before attacking the leaves. They did not seem to work rapidly, but within two weeks had prevented an ear of corn from forming upon that piece of land. Upon another larger piece of corn directly east of this, they entered at one corner and stripped that corner in the shape of a triangle including about ten acres. In Hd- wards county, I had not observed nor did I hear of damage being done to oats, wheat, or barley. Four quarter-sections of alfalfa in that lo- cality, having a rather thin stand, were kept stripped throughout the season, so that at no time was the forage worth the cutting. In the midst of these was where experiments were conducted last winter, of which extended mention will be made in this report. And here also practical applications of the “hopper-dozer” were conducted, concern- ing which discussions and descriptions will appear later in this report. Mulberry trees and other fruit-trees in the vicinity of the alfalfa fields were, after the first cutting of the alfalfa, entirely stripped of their leaves. Weeds such as the sunflower and lamb’s-quarters were de- nuded, leaving only the bare, white, stripped stalks standing. GRASSHOPPERS IN GENERAL. 13 Forp County.—In Ford county the conditions were much the same asin Edwards county. Damage was local. In one field there would be large numbers of grasshoppers and in other fields near by there would not be an unusual number. An examination of the species doing the damage showed that the one which by its numbers was the cause of a great part of the injury was the yellow grasshopper,* Melanoplus differentialis. There was also present the Two-striped Locust, Welanoplus bivitattus. When I first entered the county this Two-striped Locust was everywhere in the adult stage. While upon the first day of my investigations I did not see more than half a dozen adult individuals of the yellow locust. At different times during my investigations I tried to ascertain the relative numbers of these two species, by catching grasshoppers just as they came, and concluded that there were about twenty-five yellow locusts for every one of the Two-striped Locust. More will be said in another place concerning the different species in cultivated grounds and adjoining pastures. Finney County.— By request, I went to Finney county to make an a investigation of the condition there. In company with Mr. C. 38. Hambleton, of Garden City, I drove first west of Garden City, to the ranch of Mr. John A. Stevens, where on the north side of the road was a fine alfalfa field, bringing forth its second crop, while on the other side of the road was another alfalfa field whose first crop was being stripped by the native grasshoppers. About two miles farther west we came to a large apple, peach and plum orchard on the north side of the road. The apples were on the north boundary of the or- chard, the peaches in the middle, and the plums on the side near the road. The apple orchard had been stripped of leaves, and in some places three or four grasshoppers could be found eating at a single apple. The peaches had been eaten off, leaving the bare pits clinging to the twigs. The young twig growth of peach trees had been eaten in preference to the foliage. A man was there at work building smudges to turn the locusts from the plums, which were almost ready to gather. Upon inquiry, I learned that this farm had been largely seeded to alfalfa, but during the last two years had been pas- tured. The damage done to this place was the greatest that I ob- served in any of the thirteen counties that I visited. I believe the cause is so evident that at this point will say no more than simply to call attention to the fact that the ground upon this plat had not * This was the term commonly applied by the farmers of that region. Another locust bears this common name, and this one is generally spoken of as the Differential Locust. In this report, however, I have frequently used the term yellow locust, and wherever used it has reference to Melanoplus differentialis. I do this because this insect is quite generally known over the region covered by this report as the ‘‘ yellow grasshopper.’’ The words locust and grasshopper are used interchangeably. 14 GRASSHOPPERS IN GENERAL. been disturbed for more than two years past. The previous cultiva- tion had left the alfalfa as a food not only for the cattle, but for the native grasshopper, which flourished undisturbed. Returning by an- other route, and going out east of town, we found the damage nowhere general, but all alfalfa fields showed the effects of grasshoppers in places. Hamiuton County.— Being also requested to visit Hamilton county, I went there upon the 18th of July. Mr. C. M. Humphrey, county treasurer, being aware of my coming, took me to a number of farms in the vicinity of Syracuse. While in his office I met Hon. Thomas H. Ford, who gave me a cordial invitation to accompany him home to his ranch, about five miles east of Syracuse. Having been previously informed that he was one of the largest alfalfa growers in the county, I readily accepted. On our way out we observed grasshoppers glisten- ing, as Mr. Ford termed it, from the tops of the alfalfa stalks grow- ing in fields adjoining the road. I walked through all of these fields, to make sure of the species most prevalent, and in every case found the Differential Locust by far the most numerous. In one corner of a field, comprising less than an acre, I found a fungous disease, Empusa grilli, at work. Here were goodly numbers of locusts clinging in the tops of the alfalfa stalks. I was much interested here, as this was the first time I had observed the disease this season. Upon reaching Mr. Ford’s place, I was told the grasshoppers had been so bad that he saw the only way to save so large a crop of alfalfa was to cut it all down as fast as possible. Accordingly, at that time the whole alfalfa crop—more than 400 acres in all—was either in the windrow, swath, or shock, except a small piece that he had allowed to remain, hoping to secure seed. His young orchard was being rapidly stripped by the grasshoppers which had been driven out of the alfalfa. His neighbor on the east, Mr. Price, was also suffer- ing a great loss of peaches in satisfying the appetites of these creatures. Here was the first time I had had the opportunity to observe the work of the locusts in the irrigated districts. Former reports have stated that these native locusts deposit their eggs along the irrigating ditches, and that along these the damage is the great- est. My own observation, and the testimony of all the farmers con- sulted, did not bear out the above facts. On the contrary, the most luxuriant growth of alfalfa is on the sides of the ditches. Many of the farmers here were of the opinion that these locusts come down from the prairies. I mention this fact here, because in my subse- quent investigations I gave this point special attention. Before leav- ing the subject of Hamilton county, I might state that I returned two weeks later to observe the action of the fungus disease, Hmpusa GRASSHOPPERS IN GENERAL. 1S grilli, in the corner of the field before mentioned, and, while plenty of grasshoppers were still in that corner of the field, 7 could see no per- ceptible increase in the number of dead ones clinging to the branches of the alfalfa. From Syracuse I drove due north through Hamilton county, passing Stowell post-office, two miles and one-half on the east, and continuing north into Greeley county. I was very much interested in noting the species prevalent on these high plains. The Differential Locust, the one to which I gave the closest attention at all times, was found few in numbers among those weeds that thrive only on ground which has previously been broken. Near Stowell I came upon a large peach orchard standing out unprotected on the plain, and found therein some peaches, but not a single yellow grasshopper. An occasional Lubber grasshopper, Brachystola magna, and a number of the smaller species of the genus J/elanoplus, were all that were com- mon on the open prairie away from those weeds which are wont to grow upon cultivated lands. GREELEY County.—I entered Greeley county, six miles north and two and one-half west of Stowell, and found a few grasshoppers be- tween that point and Horace, where I spent the night. Those that I did find were the same as in Hamilton county, and existed under like conditions. North of Horace, in the valley of White Woman creek, there are a number of farms growing corn, Kafir-corn, barley, and sorghum. I visited these and conversed with the owners. From these farmers I learned that grasshoppers were not troubling them, and that they had suffered no serious damage from these insects in recent years. Leaving this valley, I proceeded north in the direction of Sharon Springs. Near the north line of Greeley county, and extending over into Wallace, is a settlement of Swedish farmers, in the vicinity of Stockholm. I found them at this date, 27th of July, in possession of excellent crops of corn and barley, and fine patches of melons and garden vegetables. At no place were the grasshoppers at all numer- ous. Some of these farmers had been residing here for ten years, and they told me that in that time they had never had serious trouble with these insects. My experience with these native grasshoppers has been that they are exceedingly fond of garden vegetables and melons, And from what I have learned of their foraging propensities, I believe that, had they been hatched in any numbers upon the plains adjoining these farmers, they would certainly have found the way to these tooth- some vegetables and tender melon vines. Mr. C. J. Momyer, Hurt, Greeley county, Kansas, wrote me that native grasshoppers were doing slight damage around the edge of his corn and cow-peas. He spoke also of the Lubber grasshopper, Brachystola magna, working upon the vines. He also spoke of the habit, already noted, of the yellow grass- with a heavy roller before sowing the wheat. In substantiation of 16 GRASSHOPPERS IN GENERAL. hopper eating the tassels and silk of the corn before attacking the leaves. Native grasshoppers have troubled him somewhat for five years. Farmers of this vicinity do not, as a rule, plow their ground every year. It was his opinion that the best wheat could be secured only from ground thoroughly cultivated before sowing. Specimens of grasshoppers received from him proved to be the Differential Locust, M. differentialis. WaLLace County.— Conditions continued the same on the road into Sharon Springs. The valley of the Smoky Hill river contains considerable alfalfa. I met a number of farmers of this valley who were in Sharon Springs trading. From them I learned that the grass- hoppers had cut down the alfalfa in patches in the fields, but there was no general or sweeping destruction being caused by them. Loaan County.—From Sharon Springs I went on a slow train over the Union Pacific through Logan county as far as Oakley. From what I could see from the train and ascertain from farmers boarding the train at the smaller stations, I was satisfied that condi- tions were in no way different from the counties previously visited. From Oakley I went north by train to Colby. Tuomas County.—Inquiries made at Colby gave answers much the same as those previously received, and made me believe that a drive out through the country would not add any new facts concern- ing the actions of the grasshopper under consideration, SHERMAN County.— Press reports having been sent out from Good- land concerning the prevalence of grasshoppers along the railroad, their numbers being reported such as to interfere with the action of the drive-wheels upon the rails, | decided to go out and investigate. Upon reaching Goodland, I was taken out in the gardens and yards of the city by Attorney G. L. Calvert, and shown the actions of the lo- custs upon the gardens. From Mr. Calvert I also learned that the damage from grasshoppers in Sherman county had been almost en- tirely confined to the crops of small grains; that the grasshoppers had eaten off the heads of the wheat, oats and barley to a considerable extent, in some fields. In the current issue of the Goodland Lepub- lican, I noticed that a correspondent from one of the outlying post- offices mentioned the fact that grasshoppers were injuring gardens seriously. In conversation with Mr. -William Walker, a hardware merchant of Goodland, and also owner of farm lands managed under his direction, I learned that it was a custom there to sow the wheat upon the ground without giving the land previous cultivation. It was Mr. Walker’s opinion that better yields could be secured if the surface of the ground was thoroughly pulverized and then packed this, he gave the instance of a man who, this year, had sown wheat upon ground so prepared and secured a heavier yield of wheat which GRASSHOPPERS IN GENERAL. 1 tested over sixty pounds to the bushel. Later, I went to the round- house of the Rock Island railroad, at Goodland, and examined a num- ber of the cow-catchers of the engines, as they came in off their runs, in order to ascertain the species of grasshoppers caught along the track. I found the species which were most common there to be the Long-winged Locust, Dissosteira longipennis, the Carolina Locust, Dissosteira carolina, and the yellow locust, Melanoplus differentialis. Through the courtesy of General Foreman Loy and his assistant, I was enabled to meet a number of the engineers and learn from them the state of the case. From Engineer Maclellan I learned that the trouble with the grasshoppers was always at night, and that at no time were they so plentiful but that an engine having forced sand draft could easily pass over them in safety; and further, that they had ex- perienced some difficulty on the run between Goodland and Norton. So, when leaving Goodland, I placed myself in a favorable position to view the grasshoppers as they flew by in front of the engine, and noted that no grasshoppers arose except as the train was passing be- tween fields where the crop was being harvested on the one side and corn or some other crop standing on the other side. The numbers at these places were not great. It was evident, however, that the grass- hoppers, in moving into the new feeding ground in the evening when the ground had become cool, were attracted by the steel rails which retained the heat longer than the ground, and when thus collected caused the trouble to the trains. It seemed conclusive that the inter- ference on the track was not because of the number of grasshoppers, but because those in the vicinity of the track were drawn to the rail by the heat. Some of the newspaper accounts which I read would lead one to believe that they were creeping over the track in a contin- uous mass. Letters received in answer to inquiries, from Messrs. D. A. Long, of Ruleton, and T. W. Simmons, of Goodland, gave the following facts: Mr. Simmons stated that they severely damaged his oats. corn, potatoes, and garden vegetables. He stated that those big yellow grasshoppers mined out the pie-plant; he feared that some of his trees, being stripped so early, are permanently injured. Mr. Sim- mons said that this was the first year that grasshoppers had ever com- mitted any serious depredations, and that the damage this year was quite local in its extent. He also said it is the belief there among many that ground plowed dried out worse than when not plowed, and that it was the custom to drill the wheat in the stubble year after year. Specimens received from Mr. Simmons were as expected —the Differential grasshopper. Mr. Long’s letter is as follows: Dear Sir: In reply to yours of the 18th, in regard to grasshoppers, I would say that they did a great deal of damage here this year in spots, mostly in the fields of wheat, barley, and oats, by cutting off the heads of the grain. Some —2 18 GRASSHOPPERS IN GENERAL, fields suffered one-fourth, some one-third; the >hopper seems to breed right in the field. There was no effort put forth to destroy them. The farmers do not plow their ground; they generally disk or drill in the grain. I think deep plowing would be a success, and in fact it has proven to be the case here this year, as some of the farmers had plowed deep, and then drilled their wheat in deep, and almost doubled the yield, and increased quality as well. The ’hoppers, I have ob- served, appear about the middle of June, and do their work in the month of July. Last year they were killed off by a little red bug or mite so small one could hardly see it with the naked eye. I will send you a few specimens in a few days. They are now working on the corn-fields, on the outer edge only. Yours truly, D. A. Lona, Ruleton, Sherman county, Kansas. Decatur County.— Reports had been sent me of trouble from grasshoppers in Decatur. Observations that I took in the vicinity of Dresden and Jennings showed conditions similar to those already stated. At Dresden I noted a field of forty acres of corn, lying north of a field of oats just harvested, almost entirely stripped, while just. across the road was another field of equal size entirely free from at- tack. Such instances could be observed in every county where I was. Norton Counry.—It was my intention to visit a number of agri- cultural localities of Norton county. I was fortunate, however, in meeting in the office of Judge Case, County Surveyor J. C. Newell, whose work had recently taken him over all parts of the county. Mr. Newell stated that the damage in Norton was almost wholly confined to alfalfa. He said of thirty pieces of alfalfa, ranging in size from ten to eighty acres, he had noticed that the effects of grasshoppers were noticeable more or less in all. He had observed no alfalfa fields that had been entirely stripped. In some the blooms and seed had been taken off, and in others spots here and there in the field had been eaten to the ground. Mr. E. E. Ames, of Norton, also told me that grasshoppers were doing slight damage in 100 acres of alfalfa which he owned. Mr. A. J. Brunswig has large alfalfa interests in this county, and on page 7 I have taken the liberty to quote a letter from him, giving his observations. Light damages were being reported by the press from other locali- ties, but I decided that the territory covered gave me a clear idea of the situation in this state. So, after leaving Norton county, I re- turned to my field laboratory in Edwards county to complete ob- servations upon the life-history of the Differential grasshopper. GRASSHOPPERS IN GENERAL. 19 SPECIES OF GRASSHOPPERS PREVALENT. It has been previously stated that the Differential Locust, Melanop- lus differentialis, was by far the most abundant. Jelanoplus at- lanis, the Lesser Migratory Locust, and Melanoplus bivitattus, the Two-striped Locust, were numerous. The Red-legged Locust, Melan- oplus femur-rubrum, was taken occasionally, generally along the roadside. The Packard Locust, Welanoplus packardi, was not un- common. On September 1, I saw females of the Packard Locust ovipositing in an alfalfa field in Edwards county. The Rocky Mountain Locust, Melanoplus spretus, was also taken. f Fig. 1. (Original.) Melanoplus differentialis ; female. Fie. 2. (Original.) Melanoplus differentialis; male. It is of interest to note the relative numbers of J. spretus and M. atlanis present. With this object in view, I captured, just as they came, a number of these allied species, and found among that number sixty-eight specimens of J/. atlanis and nineteen specimens of MW. spretus. This observation was made in the western part of Edwards county. The Long-winged Locust, Dissosteira longipen- nis, was abundant in one locality, southwestern part of Edwards county; here, in September, the females deposited their eggs in the alfalfa fields. The Carolina Locust, Dissosteira carolina, was numerous along the roads. The determination of these species was made by the writer, and for the accuracy of the work he is responsible. 20 GRASSHOPPERS IN GENERAL. LIFE-HISTORY OF THE DIFFERENTIAL LOCUST. Melanoplus differentialis Thos. This term, in its biologic sense, refers to the cycle of the individual from the embryo until the fulfilment of its mission in the perpetuation of its species and its retirement in favor of its progeny. The life of the locust may be said to begin with the egg, in which the following changes take place, and are noted under the head of — EMBRYOLOGY. By Mr. C. E. McCuuneG, Department of Zoology. Instructor in Embryology. Description of the egg.— The arrangement of the eggs in the ovary and the formation of the capsule have already been described; it is therefore only necessary to add a decription of the individual egg. This, at the time of oviposition, is an ovate cylindrical body, about 5 mm. by 1.25 mm. in size, and of a bright yellow color, gradually be- coming darker with age. The cylindrical form is not perfect, being modified in such a manner that the anterior side of the egg, as it lies in the mother insect, is flattened or even concave, while the opposite side is equally convex (fig. 3). The two ends are very similar in shape, each being bluntly pointed. V1 Fic. 3. Eggs of Melanoplus differentialis. (About five diameters. ) The egg is protected by two coats, the exterior (chorion), a secre- tion of the follicular epithelium; and the interior (vitelline mem- brane), the external, indurated portion of the egg substance. The chorion is a close fitting shell, completely enclosing the ovum. It is usually of a dull yellow color, and is marked over its entire surface by ridges that cross each other in such a manner as to form an irreg- ular hexagonal pattern. This is merely a cast of the lining of the ovarian follicle, registered in its secretion. The chorion is, at first, GRASSHOPPERS IN GENERAL. 21 tough and resistant, but, on exposure to the elements, becomes friable and easily separable. Upon the removal of this external covering, the clear, bright yellow of the underlying egg substance is seen. The structureless vitelline membrane forms an efficient protection for the enclosed egg. It withstands drying, and is broken only by the application of considerable pressure. Im- mediately beneath it is a layer of protoplasmic material, the real liv- ing matter of the egg, and within this a considerable quantity of nutritive material, the yolk. My observations upon these eggs, manner of placing in the ground and the actions of the female during the time of oviposition are here- with given. A female in quest of a suitable position for placing the eggs generally moves slowly about for some time testing the ground over which she passes. During this time the tip of the abdomen is turned downward and, stopping momentarily, the ovipositors (plate Ir) are applied to the ground. J observed one female spending thirty minutes in this way; a crevice in the soil being selected and worked upon for a time, then abandoned for solid ground. Some, however, begin digging and complete the work where the first attempt is made. Small elevated spots on the surface appear to be much chosen. Fre- quently these little hillocks were not noticeable until marked by a locust digging into the crest. From plate 111, figures g, g’, g”, it is evident that there has been a consensus of opinion in these cases con- cerning the suitability of the place for oviposition. Sandy soil, when present, seems preferable. That field observations might be corroborated in the laboratory, I brought home 1380 of the yellow locusts, more than 100 of these being females, for study in the vivarium. In one cage sixty were placed, and blue-grass sod, clover sod, sand and dirt mixed, and pure sand; each class of soil occupied a certain part of the floor, the whole floor being well packed by sprinkling. In this cage, the pure sand was chosen by all except eight, which placed their eggs in the sand and dirt mixed. Refuse from the vegetation given them for food seemed in no way to intefere with their work. I saw three of them hid away in this trash busily engaged in ovipositing. A suitable place chosen, the locust forces a hole in the ground by means of the two pairs of horny-tipped ovipositors at the end of the abdomen. These are opened and closed and the full weight of the body is brought to bear on them. In this way a receptacle is made, often in extremely firm ground, for the eggs. ‘A9Vq ULISOddQ AO WOLLOG AHS ‘NOILATHOSAG YO_ (‘TRUISIQ) “IT WLWId GRASSHOPPERS IN GENERAL. 23 Each egg is preceded by a light-colored mucous fluid. Part of this fluid passes through the walls of the cavity and causes surrounding particles of dirt, sand-and in some cases small clods (see plate 111, fig. ad) to adhere, so that the pods when removed from the ground are protected first by a coat of this sticky substance and an outer layer composed of particles of surrounding earth. This forms a brittle crust which, when pressed, often scales off, as shown by plate 11, ¢c. When the ground is firm, the walls of the pod are generally broken away when the earth is disturbed, thus exposing the naked eggs. This substance before hardening is quite plastic; after hardening it is somewhat fragile. It is insoluble in water. A pod which re- mained in water thirty days appeared to be as firm as when placed there. When the eggs are all deposited the female covers them with asmall amount of this sebaceous fluid. This hardens into a honey- comb structure, as shown is cross-section of top of pod #7 in plate 11. The cross-line near top of pod, at 6’ in plate 111, shows depth of this covering. The whole pod is “finished about one-quarter inch below the surface of the ground, and the ground covered over, leaving no trace of work, as shown on ground surface in plate 11. The arrangement of the eggs is shown in the longitudinal section of the pod in the foreground of plate 11. Number of eggs in pod is about 100, and in the ovary about the same number can be counted. Plate 1 is from nature. The writer had made a section through the vivarium, showing egg pods in position, and, while the artist was sketching this, a locust very accommodatingly came forward, began and completed the work of oviposition. Where the eggs are laid is of greatest interest to the farmer. The alfalfa fields with no intervening weeds between the alfalfa plants furnish excellent open spots for oviposition. Here I saw many species placing their eggs, and was surprised to find the Long-winged Locust, Dissosteira longipennis, among the number. Mr. Wm. Weber, who was with me and rendered valuable assistance during the summer, gave this point careful attention. He was able to cover considerable terri- tory in the southwest portion of Edwards county. This territory in- cluded pastures, alfalfa fields, and cultivated lands. He found the yellow grasshoppers depositing eggs in all these places. They were far more abundant, however, in the alfalfa fields, and remarkably so in the DEscrRIPTION OF Pirate II.— Melanoplus differentialis, female, ovipositing ; egg-pods of same species exposed to view. Grasshopper at right is depositing eggs. Three egg-pods in section of soil show position of pods and depth from the surface. Egg-pod in sand near grasshopper. Egg-pod in foreground, opened on one side to show position of eggs within; this one contained 109 eggs. PLATE III. (Original.) See Borrom or OrrosiTE PAGE. GRASSHOPPERS IN GENERAL. 25 middle ridge of roads passing through the fields. In one place he saw, on a road ridge, in a space about twenty inches wide by three rods long, one grasshopper per every two inches square of ground, oviposit- ing. The first oviposition that I observed occurred September 10, and after making a large number of dissections I feel reasonably certain that few eggs were deposited before that time. Ovipositions were not of general occurrence until September 20, and egg laying was contin- ued until cold weather. Eggs placed in the ground at this late season of the year will not receive heat sufficient to hatch, so that the species pass the rigorous period of winter in the egg-pod. During the first warm month of spring their eggs begin to hatch; then come consumption of food and growth of body. Growth meansexpansion. The insect is surrounded from birth by a rigid skeleton. This, instead of enlarging, yields to another; that is, the outer skin is shed; an under one, at first soft and accommodating, makes allowance for the increase in size. This skeleton soon becomes fixed and the insect’s size soon requires a new armament. This change, commonly called “shedding the skin,” is technically known as molting. Observations upon the number of molts this insect undergoes are not yet completed. The most interesting molt is the last one, the one in which the grasshopper brings out from the wings pads of the skin, which is being cast off, fully developed wings. The best place for watching this transformation was in standing corn, in fields adjacent to alfalfa fields. Plate Iv was made, under direction, from the author’s sketches and a large number of alcoholic specimens taken in every stage of the transformation. The full-grown nymph ceases to eat and, with the head almost in- variably downward and the antennz drooping, fastens the claws firmly into the stalk or blade, remains quiet for a short period, during which it can be handled without being disturbed; a pulsating motion be- gins in the center of the back of thorax; this increases until the whole Descriprion oF Pirate III.—Egeg-pods of Melanoplus differentialis taken from a sandy soil, showing variations in shape. a, pod with top broken off; b, pods made of sand with larger grains of sand or stone adhering; ¢, small portion of outside shell broken off; @, specimens made of sand and dirt with stones or small dead rootlets: d’, specimens composed of sand and dirt, with clod of dirt firmly fixed to the side; e, specimens broken off near the top; /, shows an un- finished rod—the grasshopper was disturbed while depositing eggs, and the pod taken in this unfinished state; g and g’, specimens taken showing two pods firmly fixed to each other, composed of sand; g”, four pods of sand and dirt with small stick and dead rootlets adhering; Fr, cross-section of top of pod, showing honeycomb structure made by the sebaceous fluid when dry. PLATE IY. (Original.) See Borrom or OpposiTE PAGE. GRASSHOPPERS IN GENERAL, 27 thorax moves up and down; soon the skin splits along the back from top of the head to line crossing the base of front wings; the upheav- ing action of the thoracic muscles continues until the body drops to the ground, leaving the nymph skin clinging to the leaf; the antennze lie one on each side of the face and are thus drawn out from under the body; the wings come straight out of the pads as narrow, much- wrinkled portions of cuticle. They are about five-eighths of an inch long when the insect falls to the ground. Inside of an hour, depend- ing upon the weather and time of day, the wings attain their full length, one inch to one and one-quarter inches. The legs are not brought into use in discarding this skin. Frequently the claws of the old skeleton break away from their attachment and the insect falls to the ground. This in no way interferes with the transformation. The insect, when free from the old covering, though its limbs are quite soft and unable to maintain its weight well, crawls to some secluded place where it awaits the hardening of the bones and the expansion of the wings. Before this is fully completed the insect begins eating again. This change occurs more frequently in the morning, though I have observed it at all hours from sunrise until five o’clock in the afternoon. The complete change from beginning of molt until the insects are fully developed occupies about one hour and a half on a bright, warm day. Frequently I observed Sarcophagid and Tachinid flies flitting ner- vously about the newly transformed insect. While I did not observe one light upon the locust, it occurred to me that at this time parasitic diptera might well place their eggs thereon. The first instances of the last molt observed were on July 6, and from the number of adults then apparent I feel safe in saying that none had reached the adult stage prior to July 1. At this time (July 1) adults of the Two-striped Locust, Welanoplus bivitattus, were com- mon, and some of them were pairing. These notes were taken in the midst of nymphs in every state of change, and frequently five or six were in view at one time. While nearly all changed in July, there were some still changing as late as September. DEscrIpTion oF PLATE IV.—Various stages of the last molt of Melanoplus differentialis. 1, nymph just before the breaking of the skin along back of thorax: 2, nymph beginning to come out; 3, mature insect dropping to the ground; 4, cast-off skin, still clinging to the leaf; 5, grasshopper climbing up, spreading wings to dry, and getting ready to eat; 6, fully developed grasshopper on corn-stalk. 28 GRASSHOPPERS IN GENERAL. HABITS. Food.—What this insect will eat depends entirely upon its neces- sities. It is a lover of good food, and knows how to find it when pro- curable. For instance, it will eat the peaches before attacking the tree; it eats the shooting ear and tassel before touching the harden- ing blade. A list of food-plants as personally observed are: Trees — cottonwood, mulberry, cherry, apple, peach, apricot, plum; herbs — alfalfa, Kafir-corn, corn; all cereals; all garden vegetables, including melons; sunflowers, lamb’s-quarters, hogweed, Russian thistle. Han- dles of hay-forks left in the field were frequently roughened by the nibbling of these locusts. Farmers in Edwards and Hamilton coun- ties mentioned the fact that gloves left about the mowing-machines were cut by the insect. Binding-twine has been cut by them so much that a twine prepared by recipe not in accord with their tastes is now much used. In confinement these locusts will feast upon one another. Actions.— Before sunrise nymphs and adults begin to climb to the top of weeds, growing crops, fence-posts, or any object standing above ground, and remain there until about ten o’clock If the ob- ject upon which they rest is edible, they amuse themselves nibbling away. I have frequently seen a portion of an alfalfa field fairly glisten with the bodies of these insects resting on the tops of the plants. About ten o'clock they descend and feed lower down, but ascend about three Pp. M. again. If the day is cloudy and cool the insects are sluggish, and remain the whole day upon the ground; so true is this that we had to abandon work with the ‘“hopper-dozer” upon one cool, cloudy day. If the day be cloudy and sultry, they eat much the same as upon bright, warm days. This habit, I believe, accounts in some instances for the belief among farmers that this locust comes in flights. One day the blades of corn will be ladened down with grasshoppers; the next day not a locust will be in sight as the farmer passes by to his work. They are upon the ground. Several instances of this kind were brought to my notice, where the farmers had reported ‘hoppers having left their corn, or reappeared, when the conditions were caused by this move- ment. Grasshoppers sitting upon the ground, especially when par- tially obscured by weeds, do not appear so numerous or so formidable as when arrayed against a growing crop. Why the borders of the field are stripped.— Damage to alfalfa is most apparent around the edges. This has led to the belief that the insects enter the alfalfa from outside territory. I observed the same conditions around long ricks of alfalfa, along division fences, where the alfalfa joined cane sown broadcast, and around large breaks in the “stand” of alfalfa. GRASSHOPPERS IN GENERAL. 29 It seems evident to me that, while some may enter from surround- ing fields, this stripping of alfalfa at places designated is caused by the insects being.checked in their progress; that is, this grasshopper, when not feeding, moves, walks awhile, rests, walks, jumps, in no evi- dent hurry, each one keeping a somewhat direct course. This action obviously takes it to the borders of the alfalfa field. Here, finding its food-plant extending no farther, it stops; it may turn back, may travel around the field, feeding as it goes. It is evident that this congregat- ing along the borders reduces the vegetation, and alfalfa once cut down furnishes in its young, tender shcots extra attractions, so that the insects remain here and new growth can make no advancement. Around the obstructions above mentioned many more grasshoppers could be seen than in the open parts of the fields. I spent much time in the vicinity of ricks of alfalfa and noted the movements, in many cases following the insects from the open field up to the ricks, and saw them in many cases continue their course until they had reached the rick, then jump off, and finally find their way past the rick. The number seemed to be about the same on every side of the rick. I speak of this matter in detail, for it was the one argument everywhere advanced in favor of the theory of the insect’s entrance from outside territory. And that they do come from outside territory is generally supposed to be the case. Length of Flight.—The flight of this insect is low and heavy, be- ing sustained for only a short distance. It is readily taken in the hop- per-dozer having sheet-iron back three and one-half feet high. I have never seen one carry itself 200 yards at a single flight, though aided by strong winds. Riley and Thomas state that this locust has been seen, though seldom, at considerable heights, apparently migrating. Habitat.—The range of this species, as given by Scudder, embraces the Mississippi valley north of latitude 438°, south to the Gulf, west to the Pacific, south to central Mexico. It has not been found above 6000 feet. Recently it has been taken in Camden county, New Jersey. Its range in Kansas is defined by the cultivated districts. I have seen it upon the streets of our largest cities, and on the tilled soil of the high western plains. NATURAL ENEMIES. VERTEBRATE. All domestic fowls feed upon locusts; chief among these, and highly commendable, is a drove of turkeys. I have seen a cat spend a whole day catching grasshoppers. A whole drove of blackbirds would sometimes light down where these hoppers were plentiful, and hold high carnival. The meadow-lark is a persistent locust catcher, and this lark abounds in the localities visited. The plover picks a locust to pieces: now and then, seemingly preferring 30 GRASSHOPPERS IN GENERAL. smaller forms. I came, however, upon a plover on the high plains in Greeley county, busily dissecting a large western cricket, Anabrus sp. The major part of. the food found in stomachs of quails and prairie-chickens examined was composed of insects belonging to this family. It is safe to say that many birds not generally accredited aid in reducing the number of locusts. Professor Snow first ascertained that the red-headed woodpecker (Jelanerpes erythrocephalus), yellow-billed cuckoo (Coccyzus americanus), cat-bird ( Galeo- scoptes carolinensis), red-eyed vireo (Vireo olivaceus), great- crested flycatcher (Myiarchus crinitus), and crow blackbird ( Qwzis- calus versicolor) feed upon the locust. Prof. Samuel Aughey has found by not less than 630 cases of dissection that ninety species of birds are partial to locusts as food. Many winter birds seek the egg- pods for food. Mr. F. E. L. Beal* states that grasshoppers are favorite food with the yellow-billed cuckoo. Several stomachs examined contained from ten to twenty of these insects, a good meal for so small a bird. Katy- dids and their eggs were found in the stomachs of these birds. The snowy tree-cricket is also used as food. Collectively, this group, Orthoptera, were found in 86 of the 155 stomachs examined, and fur- nish thirty per cent. of the year’s food. Three per cent. of the food in May is composed of these insects, and over forty-three per cent. in July. Mr. 8. D. Judd+ states that the greater part of the insect food of the loggerhead shrike is composed of grasshoppers and crickets, and in summer grasshoppers are given preference as food. The bird at this season impales upon barbed-wire fences and hedges more in- sects than it utilizes, so that an examination of the stomachs would not give a correct estimate of the insects destroyed. I saw a notice in some paper concerning an observation upon snakes, made by Maj. Frank Holsinger, member of the editorial staff of the Western Fruit Grower. I wrote Mr. Holsinger for details. His letter, a part of which follows, is of great interest : “Some years ago, while haying, I had a rake in my hands, when an immense blacksnake ran from the windrow. I placed my rake on it to hold it, but not wishing to kill it. It immediately disgorged an immense amount of grasshoppers, of a kind common in Kansas, a large yellow-legged variety that sometimes becomes destructive to our meadows. I think the discharge was fully one-half pint, and I could discover in it nothing but grasshoppers. I look upon them as our friends — rattlers and copperheads excepted. I believe they should * Bulletin No. 9, U. S. Dept. Agr. Biological Survey, June 15, 1898, p. 12. + Ibid, ‘p. 22. GRASSHOPPERS IN GENERAL. OL be protected, as their food is from insects deleterious to farming in- terests.” From the description given, it is very probable that the yellow grasshoppers referred to belonged to the species now under consider- ation. INVERTEBRATE. The locust finds, however, its most unrelenting enemies within its own class. The accompanying figure 5 shows the immature stage of the locust mite, Zrombidium locustarum, Riley. These were found, in some instances, in such numbers on the under side of the wings of the Differential Locust as to cause the wings to stand out from the body. Lo- custs with one wing or both wings eaten off were found. The wings of 1 many were rendered use- 3'.] less for flight by this : mite. According to Ri- ley, this mite drops from the wings when nearly full grown, passes through the pupal state, and comes forth an eight-legged mite to spend the winter, a part of its food being locusts’ Fie. 5. ag len RS oe hr pel acre locustarum, eggs. In the spring, the females deposit from 300 to 400 eggs, which hatch out as young, six- legged mites, that attach themselves to some host, the one chiefly chosen being the locust. This is one of the most effectual enemies of the locust. It is familiar to all. Newspaper reports concerning locusts, this year, nearly always contained some reference to a “little red bug found under the wings.” Beetles.— Predacious beetles and the larve of beetles are known to prey upon the egg-pods of this species. Flies.— Uast fall, the numbers of this locust seen dead in the fields, destroyed by internal parasites, made me desirous of procuring an es- timate of the interruptions that parasites of this class cause. Accord- ingly, I collected a large number of living forms, retaining and feeding them-in closed boxes. I did the same again this year, collecting 130 for this purpose. The computations upon these observations have appeared elsewhere, and are herewith given. Bye GRASSHOPPERS IN GENERAL. PARASITIC INFLUENCES ON MELANOPLUS.* The relation existing between the host and its parasite is an ever- interesting source of study from a biologic standpoint. Multiplied numbers of the former tend toward greater increase of the latter. When the parasites predominate, the individuals of the host tribe de- crease; should the host disappear, the parasite must follow or adapt itself to new environments. Absence of the parasite grants license to the increase of the host. The prevalence of one is directly dependent upon the other. In order that an estimate of the influence of this condition upon Melanoplus differentialis might be gained, the writer, while conduct- ing the summer field-work of this department during the two seasons past, collected a number of the Differential Locust. Fifty were taken the first week of October, 1897; 130 were collected September 3, 1898. Twelve per cent. of those captured in 1897 had been parasitized by diptera. Of those taken in 1898, 20 per cent. had been attacked by parasitic diptera. When it is taken into consideration that the dates of capture were before the close of the active season of the parasites, and that by reason of capture and confinement, some of the locusts taken were doubtless saved from attack, the estimate can be regarded as conservative. The duration of observation and number of individ- uals considered will not yet allow favorable deductions to be made from the eight per cent. increase recorded this year. In localities where this locust was superabundant in October, 1897, the number of dead forms showing an unmistakable evidence of the work of dipter- ous parasites was nearly equal to those moving about. The number of Melanoplus differentialis that appeared in the same localities this season was equal to, if not greater than, those existing the year pre- vious. This species of locust has been of economic importance an- nually in those regions for some years past. This is, in a measure, due to the peculiarly favorable condition existing there, environments which appear to be highly suitable to the rapid multiplication of this species. The ultimate effect of parasitism upon JMelanoplus differ- entialis with such surroundings is yet to be demonstrated. Observations on this subject will be continued by the department. It is the purpose of this paper to record the data observed and diptera concerned. Dr.8. W. Williston and Dr. Garry de N. Hough have very kindly examined the specimens bred. The descriptions and determi- nations of the Sarcophagide made by Doctor Hough appear below, The description of the Tachinidz which appear to be new will shortly be given by Doctor Williston in a paper on the museum types of Tachinidee. * Kan. Univ. Quar., vol. vit, No. 4, Oct., 1898, series A, pp. 205-210. GRASSHOPPERS IN GENERAL. 33 Concerning the life-history of the diptera described in this article, the following notes have been made. Careful and continued watch- ing for the act and time of oviposition was not fully rewarded. Dur- ing the period of the last molt of Melanoplus differentialis, when frequently a dozen individuals could be seen at one time in various stages of this change, the writer noted numbers of Sarcophagidee flit- ting nervously over and about, alighting near the soft, viscid locust, then taking wing again. While no act of oviposition or darting downward was observed, as is the case with many parasitic Hymenop- tera when placing their eggs, it is the writer's opinion that some at least of the eggs are placed upon the locust at this time. This belief is strengthened by the fact that the insect during the molt is quies- cent, is soft, and lightly coated with a sebaceous fluid, and therefore is an easier prey and a greater attraction for parasitic flies in quest of a host than the active and fully chitinized insect. Fia. 6. (Original.) Different stages in transformation of Sarcophagide. /, 3, 4, white maggot in active stage; 2, maggot beginning to pupate — dirty white in color; 5, pupa case, light brown; 6, 7, fully developed pupa cases of two species. The lot of specimens from which Sarcophaga cimbicis was bred was collected on September 80, 1897; the larva came forth from the host four days later. It emerged October 23, 1897. The material from which Sarcophaga hunteri was bred was taken on September 1, 1898. Three of these dipterous larvae pupated on the 3d, one on the 6th, and the last of the five on the 9th of September. They emerged in the following order: Two on September 6, one on September 8, the remaining two, a male and female, now in Doctor Hough’s collec- tion, hold the labels giving date of emergence, a copy of which I did not retain. There elapsed, however, in each case but a few days be- tween pupation and maturation. Sarcophaga cimbicis Town.; Can. Ent., vol. 24, pp. 126, 127, 1892. This specimen, a female, was determined by Doctor Hough from material in his collection. He wrote that the description by Townsend was not then accessible. Upon looking up the list —3 34 GRASSHOPPERS IN GENERAL. of types in our collection, I found the material, a male and female, upon which Townsend based his description. A careful comparison with literature at hand, made by Doctor Williston, shows the three specimens to be without doubt identical. It is interesting to note, as showing the range of adaptibility of this species, that the types were bred from cocoons of Cimbex americana. Fig. 7. (Original.) Sarcophaga ates a wr Naatert Hough. Male, Here follows Doctor Hough’s description : Sarcophaga ( Tephromyia) hunteri nov. sp. Three males and two females, bred from Melanoplus differentialis by Mr. S. J. Hunter, in whose honor I have named it. Habitat, Kansas. Length, five and one-half to seven millimeters. Color, gray; the male rather brownish, the female whitish. Abdomen without the usual variable spots of a Sarcophaga, but with three black stripes — a median, and on each side a lateral. In the female the lateral stripes are quite faint, and can only be seen well with a favorable incidence of light. Anal segments gray, retracted within the fourth segment in the males. Palpi yellow to yellowish brown. Antenne brown, with the apex of the second joint and the base of the third yellow toa varying extent. Squamule white. Wings grayish hyaline; first. longitudinal vein not spinose; third spinose for two-thirds to three- fourths of the distance to the small cross-vein. Legs black; in the male more or less brownish gray pollinose; in the female whitish gray pollinose. Hind tibiee of male not bearded. Head.—Front of male at narrowest point one-sixth the width of the head. From this point, which is about at the junction of the dorsal and middle thirds, the front widens both dorsad and ventrad: Front of female of uniform width, six-twentieths the width of the head. The exact measurements are: Male, front 0.4 mm., head 2.5. mm.; female, front 0.6 mm., head 2 mm. GRASSHOPPERS IN GENERAL, 3D Antennw.— Third joint more than twice as long as the second. Arista fully as long as the second and third joints together, composed apparently of but two joints, of which the basal is very small and about as long as broad; the terminal tapering as usual (its basal and apical thirds black, its middle third whitish), and feathered for rather more than half its length with rather long, fine hairs. The yellow, or perhaps I should say reddish yellow, color is more extensive on the antenne of the female than of the male. The vibrissal angle is a little above the mouth edge, and slightly but distinctly narrows the clypeus. Dorsad the principal vibrissa, the vibrissal ridge is beset with small bristles its entire length. Ventrad the principal vibrissa are about three smaller vibrisse. The dorso-ventral diameter of the bucca is one-third that of the eye. It is quite evenly beset with small bristles, which are larger toward the edge of the mouth opening, where they form a distinct bordering row. Macrocheete of vertex, front, ete. Male: By far the largest of the vertical bristles is the inner vertical; the outer vertical is scarcely if at all larger than the cilia of the posterior orbit. The greater ocellar are small; the lesser ocellar very small. Of the latter there are several pairs, and they extend over upon the occipital surface of the head, beyond the post-vertical pair, which is small, end very evidently a member of the ocellar group, The occipito-central is present, and is about as large as the post-vertical. There are two or three ascending and about eight decussating transverse frontals. The latter extend down upon the gena about as far as the apex of the second antennal joint. Upon the geno-plate laterad the frontals there are no large bristles, but an irregular row of exceedingly minute hairs, which begins at ora little dorsad of the middle of the geno- vertical plate and extends ventrad on the geno-vertical plate and on the gena nearly or quite to the ventral end of the latter. On the gena this row has a tendency to become double and the last three to five members of the anterior row are much larger than the rest, thus form- ing a rather prominent little group near the lower corner of the eye. The ciliz of the posterior orbit are small, closely set, and well aligned. Parallel to them is a second distinct row of bristles of about the same size. Female: The bristles of the head of the female differ from those of the male as follows: The outer vertical is almost as well developed as the inner vertical. The transverse frontals number but five or six. The row of minute hairs on the geno-vertical plate and gena has a lesser tendency to become doubled on the gena. Two good-sized or- bital bristles are present. 36 GRASSHOPPERS IN GENERAL. Thoraw.—The thorax is striped as is usual in Sarcophagw. The stripes are very distinct in the male, and quite faint in the female. The chetotaxy of the thorax is alike in the two sexes, and is indicated in the accompany-_ ing diagram. The female has a smaller num- ber of minute bristles than the male, and consequently its chetotaxy is more easily made out. In the diagram I have indicated three posthumeral bristles. The two smaller ones are in but one specimen large enough to be distinguished from the other hairs or microchetz. This variation of the posthumerals is common in Sarcophage. Fic. 9. Abdomen.—The macrocheetee of the abdomen are marginal only. Each segment has a complete row. On the first and second segments they are all of insignificant size, except two or three at the lateral border. On the third segment all are of good size and they number twelve to fourteen. On the fourth segment all are of good size and they number fourteen to sixteen. The bristles of the legs are arranged as is usual in Sarcophage. I can make out nothing worthy of special notice here. Wing.— First longitudinal vein not spinose. Third vein not spinose for two-thirds to three-quarters of the distance to the small cross- vein. Elbow of fourth almost exactly rectangular and provided with an apparent appendix, which, however, is not a stump of a vein but a slight fold or wrinkle of the wing. Hind cross-vein sinuous, longer than, but hardly twice as long as, that segment of the fourth vein be- tween it and the elbow. Hind cross-vein and apical cross-vein almost exactly parallel. This species belongs to Brauer’s subgenus Tephromyzia of Sarco- phaga (sens. lat.) In this subgenus the vibrissal angles are distinctly above the mouth edge and, projecting somewhat mesad, distinctly narrow the clypeus. The abdomen does not have the changeable spots, macule spurie, of Sarcophaga, but is either unicolorous or marked with fixed spots or lines. The European species of this group are T. grisea Meig., T. lineata Fall., 7. affinis Fall. and 7. obsoleta Fall. As far as I am aware hunterz is the first Tephromyia to be observed outside of Europe. Through the kindness of Herr Paul Stein, of Genthin, Germany, I have now in my possession specimens of grisea, affinis and obsoleta. From these specimens and the accessible descriptions of /ineata, I am able to construct the fol- lowing table for separating the species of this group. GRASSHOPPERS IN GENERAL. 37 A.— Abdomen unicolorous, squamule yellow, wings strongly yellow at base — grisea Meig. AA.— Abdomen with distinct black markings, squamule not yellow, wings not strongly yellow at base. B. B.— Palpi black. C. C.— Each abdominal segment with a black dorsal line, and on each side with a narrow, oblique, black spot; these spots often united so that the abdomen pre- sents three black stripes. Front of male one-third the width of the head — line- ata Fall. CC.— First abdominal segment blackish: other segments each with a dcrsal black line, and on each side with a large, irregularly shaped black spot. Front of male one-fifth the width of the head; of female, one-third the width of the he d—affinis Fall. ‘BB.— Palpi yellow or brownish yellow. D.— Front of male one-fourth as wide as head; third antennal joint less than one and a half times as long as the second; no intra-alar bristle in front of the suture —obsoleta Fall. DD. Front of male one-sixth, of female less than a third, as wide as the head; third antennal joint more than twice as long as second, with an intra-alar bristle in front of suture —/hunteri n. sp. Dr. S. W. Williston has kindly examined the Zachinida, and will give descriptions in a paper upon Kansas university museum types of Tachinidae in a future number of the Kansas University Quarterly. Among these parasitic Diptera, the female fly deposits the eggs, some- times two, upon the back of the grasshopper, frequently when locust’s wings are spread in flight. The egg soon hatches, and the larva feed until full-grown upon the locust’s vitals. It comes forth as a white maggot; soon its skin turns brown and forms a hard pupa case, from which the flies above mentioned emetge-in: adult form in from ten ©26;20, ((nemat) Brox eineraseens- days to two weeks (see fig. 6). An Asilid fly, Hrax cinerascens, was seen pouncing upon the young grasshoppers. Locust fungus, Empusa grilli Fres. This disease can be readily detected when present, by the general observer, by the numbers of dead locusts clinging to the tops of the alfalfa, weeds, or grass. On the 21st of July I found this fungus, for the first time this year, at work in one corner of a small alfalfa field three miles east of Syracuse, Hamilton county. The grasshoppers were very abundant upon this piece of ground. I returned to this place two weeks later, hoping to attain some valuable data upon the natural spread of this 38 GRASSHOPPERS IN GENERAL. disease. This second visit showed that there had been no noticeable spread of the disease. Dead grass- hoppers, in about the same numbers, in practically the same corner of the field, were to be seen. I did not find the disease working in any other por- tions of this field. I found a few dead grasshoppers that had died from this disease in two spots in two dif- ferent alfalfa fields in Edwards county. In no place, however, did I note anything that could be in any way considered an, epidemic, nor any evidences that would induce favor- able conclusions concerning this dis- ease as a valuable check to the increase of this locust. Observations Made in Widely Sepa- rated Localities. In the vicinity of St. Louis, Mis- souri, the first specimens of this lo- cust were observed to become winged July 19. Eggs were laid Septem- ber 9. As a deviation from the usual egg-laying babies of “the Bore ee Fie. 11. (Original) ‘tela eee ait. eggs are sometimes very numerously foreniaae killed by fungus. On Golden- placed under the bark of logs that have been felled on low lands. The eggs of this species, unlike those of spretus, atlanis, and femur-rubrum, are not quadrilinearly but irregularly arranged. The head ends of the eggs in the pod point mostly outward. One hundred and seventy-five eggs have been counted in a single mass.* In California, they acquired wings from the last week in June to the last week in July, and began laying eggs July 23. A single female occupied seventy-five minutes in depositing an egg mass. The situation chosen for egg laying was invariably the edge of one of the basin-like hollows (for irrigation ?) at the foot of atree. This locust is not easily startled; its ordinary flight is rather heavy, and sus- tained only for a distance of twelve to thirty feet.t * Riley, Summary from First Report U. S. Ent. Com. + Coquillett, Report Dept. of Agr. 1885, pp. 295, 296. GRASSHOPPERS IN GENERAL. 39 This insect has very frequently multiplied in such numbers in limited areas over its range as to do considerable injury to cultivated crops growing upon low, moist ground; and has even been known very frequently to spread over higher and dryer lands adjoining these, its customary haunts. It is one of the few species of locusts that has thus far shown a tendency toward civilization. This it has done readily, since its habits are in unison with the cultivation of the soil. It is only since the settlement of the country where it originally occurred that it has multiplied so as to become sufficiently numerous to become a serious pest. The eggs are laid in cultivated grounds that are more or less com- pact, preferably old roads, deserted fields, the edges of weed patches, and well-grazed pastures adjoining weedy ravines. Egg laying begins about the middle of August and continues into October, varying, of course, according to latitude and climatic conditions. Usually, but not always, only a single cluster of eggs is deposited by each female. Frequently there were two, and in extreme cases perhaps even three, of these clusters deposited by a single female.* ANATOMY. It is believed that a brief general discussion accompanied by fig- ures of the Differential Locust, Welanoplus differentialis, will be of practical value. Such a treatise will enable the farmer to understand more readily the direct action of the external and internal remedies used, ExTEeRNAL ANATOMY. A word, in beginning, concerning skeletons. These are of two kinds: skeletons within the body and surrounded by muscles, and those without the body, having all muscles on the in- terior. The skeletal structure of man comes under the first class; the rigid outer structure of insects under the second class. Every one who has studied human physiology remembers, among the first topics, “Uses of the Skeleton,” and if he were permitted to use another’s lan- guage, instead of his own, he would say, in answer: ‘To protect the delicate organs, to furnish attachment for the muscles, to give form to the body, to furnish levers for the movements of the body.’ And, if this same scholar was to take up comparative anatomy later, he would find that the skeleton of the grasshopper serves identically the same purposes as the human skeleton. The integral parts of the human skeleton we call bones; the separate pieces of the grasshopper skele- ton we term sclerites; that constituent which gives bones their firm- ness we commonly speak of as lime; that which lends rigor to the * Bruner, Report of Ent. to Neb. St. Bd. of Agr., 1896, pp. 120, 121. 40 GRASSHOPPERS IN GENERAL. Fig. 12. (Original.) Front view of head, with clypeus and labrum removed to show mandibles in position. a, antenna; b, ocelli; B, compound eye; (’, mandible; d, maxilla; e, maxillary palpus; f, labium; g, labial palpus. En- larged about five times. sclerite is a horny substance called chitine. An examination of the body wall shows it to be composed of a number of distinct pieces, or sclerites; the lines separating these pieces are known as sutures. Su- tures here, just as in the anatomy of the human skull, are not freely movable joints. The term, “joint,” is reserved for those articulalations where free motion is per- mitted, as is the case at the connections of the parts of the locust’s leg. An examinination of the whole body will readily show three divisions—the head, thorax, and abdomen. The head, apparently one piece, contains the mouth, eyes, and the long horns, known as an- tennze; the thorax is the heavy central part of the body, furnishing attachments for the wings and legs; the abdomen is the slender portion extending backward from the base of hind wings. The head.— The accompanying figures show the head with mouth closed (fig. 12), but upper lip, labrum, removed to reveal comparative size Fig. 15, with mouth open, illustrates the tongue, hypopharynx in the of jaws and head. Fie. 16. (Original.) Cly- peus and labrum, Enlarged about five times. center, mandibles, and part of the maxille on Fic. 13. (Original.) Front view of head, with mandibles spread out. c, mandible; d, maxilla; e, maxillary palpus; /, labium; g, labial palpus; A’, hypopharynx. Enlarged about five times, each side beneath mandibles. With such tools and the power to use them, is it to be wondered that this insect is omnivorous. The compound eyes, one on each side of the head, are the most conspicuous divi- sions of the head. Their surface is made up of a large number of hexagonal plates. Each of these plates forms the surface covy- ering for a simple eye admitting but one ray of light. The compound eye is but a collection of these simple eyes. The simple eyes.—Just above the base of each antenna can be found a small simple eye; between the bases of the horn- like antenne is another simple eye. The many jointed antenne extend from GRASSHOPPERS IN GENERAL. 41 the upper part of the face. upper lip (labrum) are drawn in full in fig- ure 16. The powerful toothed mandibles are Fie. 14. (Original.) Labium. g, gula; S, submentum; M, menl tum; fg, palpiger; Lp, labia- palpus. Ligula consists of the two flaps below mentum. En- larged seven and one-half times. given with figure (fig. 14). The clypeus and shown open and closed ( figs. 12, 13). The movements of the mandibles are confined to the lat- eral action from the median line outward and back again. Just behind them are the smaller, more strongly toothed maxillw, with appendages. These are drawn and described in figure 15. Between the maxille, and rising from the back wall of mouth, is the tongue-like organ known as hypopharynx. The under lip (labium) is shown as it appears from the back of the head. Its subdivisions are The large scle- Fic. 15. (Original.) In- ner view of maxilla. Le, lacinia; gi, galea; p, pal- pus; m, membrane. En- larged about five times. rite composing the front part of the head, in which are situated compound eyes, sim- ple eyes, and antennez, is known as the epicranium. The upper part is called the vertex, the anterior portion the front, and the sides, extending downward from the compound eyes, the cheeks or genw. Fic. 163. (Original.) Side view of prothorax with leg. a, pre- scutum; 6, scutum; c, scutellum ; d, post scutellum ; e, episternum: m, membrane, connecting head with prothorax, containing the jugular sclerites; /, coxa of leg; g, trochanter; h, femur; 7, tibia; j, tarsi; k, pulvillus and two claws. Enlarged three times. The thorax con- sists of three divisions — the prothorax, bear- ing the front pair of legs; the mesothorax, bearing the front wings and the middle pair of legs; the metathorax, bearing the hind wings and last pair of legs. The prothorax is made most conspicu- ous by the pronotum—the large sun- bonnet-shaped piece covering the back —extending out over the back: of meso- thorax and covering the greater portion of the sides of the prothorax as well. The subdivisions of the hood are given beneath Fie. 17. (Original.) Side view of thorax. Enlarged three times. A, Mesothorax. {| B, Metathorax. a, parapteron. b’, episternum. b, episternum. | c’, epimeron. c, epimeron. d', wing. d, wing. 42 GRASSHOPPERS IN GENERAL. figure 164. The sternum of the prothorax is a sclerite having a small tubercle situated between the front legs. On the membrane connecting head and prothorax, on each side underneath the prothorax, are a pair of sclerites looking somewhat like two links in a chain; these are called the jugular sclerites. They are represented in fig- ure 163, as is also the episternum of the prothorax and prothoracic leg. Mesothorax.—The middle part of the thorax, when viewed from above, shows the front wings (tegmina) and mesonotum between the wings (fig. 19). The mesonotum consists of two subdivisions—the scutum occupying the front half and scutellum the back half of this division. The form of each is shown in figure 19. From the side can be seen two sclerites named and referred to in figure 17, also the place of attachment of the middle leg. The ventral portion (mesosternum ) be Fie. 18. (Original.) Metathoracic, or jumping leg. c, coxa; tr, trochanter; /, femur; ti, tibia; ta, tarsi; p, pulvillus and two claws. Enlarged four times. of the mesothorax is a large, flat, nearly rectangular piece, having the corresponding sclerite of the metathorax dovetailed into its caudal margin (fig. 20). Metathorax.— This part resembles both in structure and function the mesothorax. Reference to figures 17, 19, 20, will give names of parts and appearances in structure. Between the mesothorax and metathorax, on the side just above the leg socket, is the largest open- ing in the body for the transmission of air. These openings are called spiracles. See figure 27 for illustration of structure and position. Appendages of the thorax.—These are the legs and wings. The first and second pair of legs are used in walking and grasping; the GRASSHOPPERS IN GENERAL. 43 third pair in walking and jumping. The function of the third pair differs, and yet the parts of each are the same, and these are given, together with their relative forms, in figure 18. The wings differ in structure and texture. The front wings (teg- mina) are large, narrow, and retain the same form whether in flight or at rest. The hind wings, when not in use, are folded like fans and rest under the tegmina. b The membrane of the wings is supported by longitudinal veins and short cross-veins. d Abdomen.—The first abdominal seginent fits accurately into a notch in the metasternum Wi eae, Bert 209 SEEM oa a cae ne aa a == S2= = J a= rat (ODS Fie. 19. (Original.) Dorsal aspect of body (female). A, head; 1, antenna; 2, epicranium; 3, compound eye. #&, prothorax; @, prescutum; 6, scutum; c, scutellum; d, post scutellum, C, mes- onotum; se, scutum; sc/, scutellum; w, tegmina or wing cover. D, metanotum; se, scutum; se/, scutellum; w, wing. LH, abdo- men; /, 2, 3, 4, 5. 6, 7, 8, 9, 10, 11, segments; aud, auditory organ; sp, spiracles; ¢c, cerci; pn, podical plates; v, ovipositor, Enlarged about three and a half times. (figs. 20,21). On the upper half of each side of this first segment is a large opening covered with a delicate membrane (figs. 21, 24). These are the auditory organs. The membrane is the tym- panum. In fig. 24 the auditory organ is enlarged. The margin is the thickened tympanum. The dark structure at the right on tympanum is the cone-shaped prominence which is situated just beneath the tympanum. The dark spot near center is the trian- gular chamber situated just beneath the tympanum. Just in front of each auditory organ is a spiracle (figs. 21,24). The next seven segments contain breathing spores (spiracles) on the 44 GRASSHOPPERS IN GENERAL. sides, at points indicated in fig. 21. The exact use of these will be more fully dis- cussed later. Beginning with the ninth segment, the abdomen is modified, and in the structures which follow are to be found the charac- teristics which distinguish the sexes. In the female the four long, horny-tipped ovi- positors are prominent. How such small instruments can execute so much work, is a matter not easily explained. Their uses were discussed under ‘“Life-History.” ae NY ‘ix Sar aS i PP We i , : hi ag fei A Pies NT gy, dni Ai yn ai ie EMM rere So iy ll in 3 ~~ ps ait On mn, Fic. 25, (Original). Digestive, circulatory and nervous systems. /, 2, 3, 4, 5, 6, 7, 8, 9, 10, 11, 12, segments; a-a, digestive tract; H, hypopharynx; Lb, labium; Lin, labrum; Lp, labial palpus; mp, maxillary palpus; @, esophagus; pp, ovipositors; eg, egg guide; co,colon; r, rectum. The heart is an open tube running along the back; it is so marked, but not easily shown. Enlarged three times. extending from about the tenth segment of ‘the abdomen up into the head. This tube has valves along its sides which admit of entrance of food and do not allow that which has entered to escape until it passes out of the main opening at the end of this organ in the head. This leads us to make a few remarks concerning the blood of insects. 46 GRASSHOPPERS IN GENERAL. The blood of insects differs from that of other animals in having no red corpuscles. It is a thin fluid and, being a mixture of blood and chyle, usually colorless, but sometimes yellowish or reddish. It is carried forward by this tube or heart to the front end, and then flows back, nourishing the organs as it passes, and likewise coming in con- tact with tracheze, which are everywhere present in the body. When in contact with these trachee, action similar to that in the human lung takes place. More will be said upon this phase of the subject under the head of the ‘“ Respiratory system.” It will be seen that the chief function of this heart is to conduct forward newly made blood and unused blood from the back end of the body, pour it out at the front end of the body, and allow it to flow back like a river in its course. This action of the heart can be seen with the naked eye in some caterpillars with light color and delicate skin, when held be- tween the observer and the sun. Fic. 26. (Original.) Respiratory system. sp., spiracles, showing tracheze permeating all parts of the body; 8, airsacs. Enlarged three times. Respiratory system.—This insect, instead of having one portion of the body set apart for the purification of the blood, similar to ani- mals possessing lungs, may be said to have lungs all over its system ; that is, there are tracheze branched and branched until they cover every part of the system and extend to every organ in the system. These tracheze do not depend upon the mouth for their supply of air, but are connected with the body wall direct, the outer portion of this connection being known as spiracles (figs. 21, 27). These spiracles have valves and openings which close and open at intervals, allowing free interchange of air. The tracheze which run from these spiracles are membranous tubes, which do not collapse because they are kept open by continuous rings of cartilage similar, though on a much smaller scale, to the cartilage in the windpipe of those animals bearing lungs. In addition to carrying to all parts of the body, it would seem that this distribution of air within the body tends to make the insect lighter and more capable of flight. In addition to these trachex, GRASSHOPPERS IN GENERAL. 47 Fic. 27. (Original.) Side view, ae Sprracle:/A Ay | / , showing spiracle of mesothorax with TB & spiracle enlarged. ” enlarged : ma ok " — ‘ ess 2 7) ~~ D ae petite however, there are organs es- *. : pecially made to assist in buoying the insect when on ¥ Sa ae the wing. These are com- f \ monly known as air sacs, and ; connect with the spiracles as vA shown in the figure (fig. 26). Reproductive system.— Fig. 25 shows the reproductive organs of the female, when not laden with eggs. It consists of an ovary, egg duct, and receptacle for the fertilizing fluid. Fig. 28 shows the ovary of the female just a few days previous to the time of oviposition. It Fig. 28. (Original.) Reproductive system of female. Large egg sac lying above stomach; oyi- duct leading out above egg guide; 7, rectum; a-a, digestive tract. Enlarged three times. is believed that, in these insects, each egg is fertilized in passing from the ovary out through the egg duct. The number of eggs that the ovary of this insect will contain averages about 100. Nervous system.— The nervous system is shown in Fig. 25. It con- sists of a series of ganglia, or collections of nervous matter, situated under the digestive canal. These ganglia are arranged along the body just next to the digestive tract. They are placed two together, and 48 GRASSHOPPERS IN GENERAL, these pairs, three pairs in the thorax and five pairs in the abdomen, are joined to each other and to the ones of the corresponding side by a cord of nerve tissue known asa commissure. This forms a double chain from the back part of the body up to the head, where a nerve band is formed around the esophagus; on the top of the esophagus are to be found the two largest ganglia in the body of the insect. These two ganglia found above the esophagus might be looked upon as the brain proper. From these there go out branches of nerves to the eyes, to the antenne, to the maxille and mandibles, and other parts of the face. MEASURES OF PREVENTION. It is stated in sacred writ that there may come a time in the life of a man when the grasshopper shall be a burden. While this prophecy referred to the closing days of a man’s life, it is evident that under certain conditions the grasshoppers might become a burden before man will have reached the time when the grinders cease because they are few. To avoid this burden, then, isa matter which will interest every one concerned, and to this subject we will now devote our at- tention, under the head of ‘*Measures of Prevention”; and in this connection we will not discuss remedies in general, but those which we have actually tried in the localities interested, and which have proven most successful. We are glad to say that our experience has shown that in the cases under observation it is not by killing two birds with one stone, but rather three birds, instead of two. have been hit with the same missile. As before stated, the conditions which favor the rapid multiplica- tion of this species are, the soil undisturbed by cultivation, and food- plants which send forth nourishing vegetation early in the spring. These conditions are to be found in Kansas wherever alfalfa is sown and not cultivated thereafter, and where the small grain, such as bar- ley and wheat, are sown without the ground being previously stirred. Wherever we have found these conditions we have found this species of grasshopper in burdensome numbers, together with other kinds of locusts in lesser numbers. In the first place, I should like to call the attention of those inter- ested to the subject of the value of cultivating wheat ground before sowing the seed. In making this suggestion, I am fully aware there is an opinion prevalent in the western part of this state that more wheat can be raised upon ground into which the seed has been placed without previous preparation than upon the same ground after having received some cuitivation. While I am aware that this belief of the farmers is based upon observation covering several years, I believe that observations proving the contrary can be found. GRASSHOPPERS IN GENERAL. 49 In substantiating this statement, I refer to a tract of land near Goodland, Kan., the circumstances of which were related to me by Mr. Wm. Walker and Mr. G. L. Calvert, both of Goodland. This piece of land produced this year thirty-five bushels to the acre, and gave a quality of wheat that tested sixty-three pounds to the bushel. A number of pieces of wheat in the same locality were seeded by sow- ing in the stubble and gave only nominal yields. In this connection, it might be well also to mention the Campbell soil-culture method, which consists largely of thoroughly cultivating the upper layer of the soil and then thoroughly packing the same. Very favorable re- ports have been received concerning the results derived from this sys- tem of tilling the soil. And further, I am glad to make an extract here from a letter received from Hon. B. A. McAllaster, land commis- sioner of the Union Pacific railway. The extensive landed interests of Mr. McAllaster bring him in contact with many different soils, different climates, and different methods of culture. These facts make him speak advisedly when he says: Union Paciric Rattroap Company, LAnD DEPARTMENT, Omana, Nes., August 18, 1898. Dear S1rr—I am particularly impressed with the statement made by you in the latter part of the paper, to the effect that the action of western farmers in not plowing their land except in periods of three to five years apart, is one of the principal causes for the increase of grasshoppers. It isand has been my opinion, very frequently expressed, that the crop failures in the western part of the state, which have been experienced during the past few years, are to a considerable extent due to this failure to properly plow and cultivate the land. It seems to me that a farm should be plowed every year in western Kansas, just the same as it is in other parts of the universe, and if, as alleged by the farmers as an excuse for their methods out there, the plowing of the ground allows the seed to be blown away by the high winds, that could be overcome by rolling the ground after plowing, or by using a subsurface packer. I believe that if I were a west- ern farmer, I would prefer taking my chances of seed being blown away, rather than having the crop eaten up by the grasshoppers. Yours truly, B. A. McAuuaster, Land Commissioner. The reader is also referred to the letters of Mr. C. J. Momyer, on page 15, and Mr. D. A. Long, on page 17, for opinions upon soil cul- ture in western Kansas. ‘ADVd WLISOddOQ 4O WOLLOG AHS ‘NOILdINOSag BOF “A WLVId GRASSHOPPERS AND ALFALFA. 51 The subject of prevention, then, will be considered under two di- visions, methods of cultivation and means of destruction. ALFALFA CULTURE. Since it has been shown in the previous pages of this publication that the conditions essential to the production of large numbers of these native grasshoppers are two in number: an undisturbed soil for the safe retention of the eggs, and an early food-plant for the nourish- ment of the young insect, every one who is familiar with an alfalfa field will readily see that in this field are two conditions which are highly adaptable to the production of these insects. The problem which faced this department when attention was called to the subject two years ago was, how to disturb the solidity of _ the alfalfa field, destroy the eggs of the grasshoppers, and yet main- tain the integrity and life of the plant. This is the problem which was not readily solved, and after a solution was presented it did not find ready acceptance. The mode of procedure was as follows: It was already known that the alfalfa plant has a long tap-root, which extends many feet beneath the surface ; and experiments showed that this root could be chopped and split at the top, and yet the life of the plant maintained with the same degree of vigor. The attention of the farmers had been called to this fact, and they were strongly urged to disturb the soil of their alfalfa fields with the disk harrow. Many were influenced to do this, but some were of the opinion that the fields would be injured by the process. A further discussion of the subject may be better set forth by giving a detailed account of the work carried on upon the land where the experiments of this department were conducted. Early in March, 1898, Mr. J. H. Smart, following instructions, caused disk harrows to be run over 160 acres of alfalfa ground belonging to the firm of Ball & Goddard, for whom he is superintendent. I have asked Superintendent Smart to give a brief statement of his work, manner of conducting the same, and his opinion upon the results. Believing that the same will be of great value to those interested, I have caused it to follow: ALFALFA, IRRIGATION AND LAND Company, CatTLE BREEDERS AND DeraLeRs, Kinstey, Kan. The good effect of cultivation as applied to alfalfa has been very apparent this year on the land of the Alfalfa, Irrigation and Land Company, situated in Edwards and Ford counties, and operated by H. E. Ball, of Topeka, Kan. In DescRIPTion OF PLATE V.—First crop of alfalfa; windrow in immediate fore- ground. This ground was thoroughly disked in March. For details, see Supt. J.H.Smart’s letter, above. Land in Ford county. (Photographed by Hunter on July 23, 1898.) 52 GRASSHOPPERS AND ALFALFA. March, 1898, we started two teams, disking 160 acres of alfalfa in Ford county. The harrow had twenty-inch disks and was set to cut about three and one- half inches deep, as an experiment. We afterward ran a light small-toothed harrow over the ground to level it after the disking. After this harrowing, the ground had the appearance of a wheat field that had been plowed very shallow. Almost immediately the good effect of the cultivation commenced to show, and so continued until the present time, not alone in the increased yield of alfalfa, but by destroying the native grasses which had not been entirely subdued when the seed was sown. Iam of the opinion that the proper time to cultivate is early in the spring, while the ground is mellow, after the winter freezing, and the plant has not started to grow. The cutting up of the ground at this season of the year also exposes all the eggs of the insects to the action of the early frosts. This is very noticeable on this particular field, as there was not one-sixth as many ’hoppers as on an adjacent tract of an equal area, sown in the same season by the same party, and treated in every particular the same, with the exception of the disk- ing given the former in the spring. The yield was one-third more and of a bet- ter quality than the land that received no cultivation. There was no bad effect on the plant to be noticed, such as cutting off the crown of the plant. It seemed to do it good. The yield of the second crop was a decided surprise. We will continue the disking of our alfalfa fields this winter and early spring. I will disk a part of the same quarter-section this spring that we did last spring, and note the effect. In addition to increase of yield and absence of grasshoppers, this field was very noticeable on account of being clear of weeds and free from native grasses. J. H. Smart, Assistant Superintendent. My own observations upon this quarter-section I have endeavored to record, not only by means of words, but by photographs taken at the time, plates of which were made, and appear as plates 1, v, and v1, with detailed descriptions given. The first crop averaged in height twenty- nine inches; stalks of extreme length were found thirty-three inches. This first crop was marked by an evenness in height of growth, and by the remarkable freedom from the presence of native grasses and noxious weeds. The second crop, however, appeared to me to show by far the more striking benefits derived from disking the ground, and, for an idea of it, I should like the reader to return to plate 1, and note the height and luxuriance of the growth. This second crop averaged in height twenty-eight inches; stalks of thirty-two inches were found. I should like to say further, that the photographs were taken from different places in the field, and that one place in the field appeared to be equally as good as the other, with the exception of a low swale, where the ground was poor and non-productive. Con- cerning this second crop, it is not an exaggeration to say that, in thirteen counties traversed, I saw nowhere, wader any conditions, a second crop of alfalfa which excelled the one upon this ground. It is of more striking interest to note that this crop was brought to maturity and harvested with no appreciable loss on account of grass- hoppers, while two quarter-sections near by, of the same kind of soil, gave neither a first nor a second crop on account of the prevalence of GRASSHOPPERS AND ALFALFA, 53 grasshoppers. Here it seems to the writer is an excellent example of the old. expression, ‘“‘killing two birds with one stone.” The destruc- tive influences of grasshoppers are removed, and at the same time and by the same work the yield is made more certain and of much greater quantity. Nothing was so convincing of this effect as a look over a quarter-section thus treated, and that this view might be given the reader, plates numbers I, V and VI are produced. At the close of the season, I asked Superintendent Smart for an estimate of the yield of the alfalfa land under his care. His state- ment I will give verbatim: “In regard to the yield of alfalfa on our lands, I will say that the first crop raised on land that was disked was about one and one-fourth tons per acre, and the second crop, one ton per acre. We have harvested on our lands in Edwards and Ford counties, the past year, about 2500 acres, and the average yield for the first crop was three-fourths of a ton per acre, and the second crop, one-half ton.” Of further interest will be the wide experience of Superintendent Watson upon this subject; I have deemed it advisable to ask him to give his views, and these are as follows: ALFALFA, IRRIGATION AND LAND CoMPANY, Lanp DEPARTMENT. I have been intimately connected with alfalfa culture in western Kansas for five years and have had charge of the seeding of 4000 acres. The advantages and profits arising from this crop are inestimable. No forage-plant is better adapted to the soil of the Arkansas valley than is alfalfa. I have observed, how- ever, in recent years,a prevalence of native grasshoppers. The cause of their increase was not clear to me until my attention was called to their appearance in alfalfa fields, and that in territory away from alfalfa they were not so trouble- some. For instance, I had a half-section of land on the high prairie in cultiva- tion for eight years, and grasshoppers gave no trouble whatever. Four years ago I sowed twenty acres of alfalfa upon this tract. This year the grasshoppers caused damage to alfalfa seed crop and damaged forty acres of wheat joining. I am glad, however, to say that it has become evident that a better degree of culti- vation will not only remove the grasshoppers, but also increase the quality and quantity of alfalfa produced. The alfalfa lands disked, upon our ranches, ac- cording to instructions, have shown remarkable freedom from grasshoppers, and very satisfactory increase in the alfalfa yield. If farmers will codperate in culti- vating their alfalfa fields in the early spring, and use the hopper catcher to capture any *hoppers that may appear, it is very evident, since these are bred and die upon the same farm, that in a year or two there will be no further damage caused by them. Gro. W. Warson, Superintendent. / ‘HOV ALISOddO AAS ‘NOTLAIMOSHq AOA ‘IA ALVId DESTRUCTION OF GRASSHOPPERS, do THE HOPPER-DOZER. See plates vir and vitt, figs. 29 and 30. It has been shown that methods of soil culture which include the turning of the ground annually will destroy the locusts’ eggs placed therein. It is evident, however, that some may escape and others hatch from undisturbed roadsides and pasture lands. To destroy these before the egg-laying time means not only a cessation of the damages caused by these, but also that every female killed causes a reduction of the number of young the following year to the amount of at least 100 individuals. : The simplest and most effectual machine, the one used by the de- _partment in its field-work this year, is commonly known as the hop- per-dozer. The plan of construction, it is believed, can be best set forth by means of a sketch, giving dimensions of material used and showing positions in which each part is placed. Hon. Thos. H. Ford, of Syracuse, kindly sent me the plan of the one made under our direction and successfully used by him this season, and this is here shown. This machine cost Mr. Ford, in complete form ready for use, five dollars. The pans are more readily constructed from two sheets of galvan- ized iron, and more easily handled. These pans should be two feet wide, four inches deep in front and eight inches at back. While it cannot be easily shown in the sketch, yet it must be understood that these pans are laid upon 1x4 boards previously nailed to run- ners. The height of the runners is not given, since that depends upon the height of crop to be protected. It is important that there be no timbers in front of pan, so that the front line of the pan may come in contact with the grain passed over. The insects then fall directly into the fluid. When the machine is ready for use, place two buckets of water and one-half gallon of coal-oil in each pan, and then drive back and forth across the end of the field where the grasshoppers are entering until you have filled the pans, remove insects, replenish with oil and water, and continue until the field is rid of the pest. There will }be many grasshoppers strike the sheet-iron back, drop into the pans and immediately jump out again. Those farmers who watched the experiments this year were at first of the opinion that the locusts that jumped out had jumped away “to live another day.” The writer asked those interested to watch the insects and note the actions DescrRIPTION OF Prare VI.—Photograph of the disked alfalfa field in Ford county, showing the alfalfa yield; in swath in foreground, in windrow in back- ground. Sand-hills in the distance. (Photographed by Hunter, July 23, 1898.) Penrod ‘ fe } é ' ‘ eB For DESCRIPTION, SEE OPPOSITE PAGE. PLATE VII. DESTRUCTION OF GRASSHOPPERS. ar \ AM Galvanized jron N pd iA ZO TT At ER oa — a Fie. 29. (Original.) Plan for hopper-dozer. of a grasshopper that had jumped out. In every case the report was that the insect “became sick and soon died.” In fact, persons going over fields where a day or so before the hopper-dozer had been at work were impressed with the number of dead grasshoppers scattered about on the ground. An examination showed the presence of coal-oil upon the body. This coal-oil and water is an external irritant, and my observations have been that the mixture is more effective than the pure oil alone. The use of the machine may be best shown by examples. In Ford county this season a large tract of alfalfa was cut, and the locusts at once began moving into a large field of Kafir-corn which had been sown broadeast. The hopper-dozer was drawn back and forth across the end of the corn-field nearest the alfalfa land until a portion of the field about twenty rods deep had been gone over. Here it was appar- ent that there were very few grasshoppers; or, in other words, the ad- vance line of the locusts’ march only extended twenty rods into the Description oF PLare VII.— Hopper-dozer at work on the ranch of Hon. Thos. H. Ford, Hamilton county. The photograph is taken in alfalfa field, which is being protected for seed crop. Quantity of grasshoppers just taken from pans can be seen in front of pans. This machine was made after instruc- tions of this department, except the 2x4 fourteen-foot extending forward by the side of each mule. These were added by Mr. Ford, and are of great value in steadying the machine. (Photographed by Cass.) ‘e ms fs} S 6 z & c t ve For DESCRIPTION, SEE OPPOSITE PAGE. PLATE VIII, DESTRUCTION OF GRASSHOPPERS. 59 Fic, 30. View in front of hopper-dozer, showing quantity of grasshoppers just taken from the pans. field. Two days later the same amount of ground was covered, but not as many insects were taken. Grasshoppers no longer entered this corn and the hopper-dozer was no longer used at this point. It has been my experience with this machine that after it has passed over vegetation it does not injure the plants, but in some way renders vegetation distasteful to the grasshoppers, so that they turn their course and seek food elsewhere. I have observed that these native grasshoppers enter a field from one corner or side, and that they are not asa rule scattered over the whole field, but occur in great numbers in patches. This being the case, it is evident that with very little labor with this machine the products of a field can be given fwll opportunity to mature. Mr. Ford, of Hamilton county, used this machine to protect the al- falfa seed crop. He did not stop, however, with guarding this field, but caught them wherever they were to be found. Some weeks after I left, Mr. Ford wrote me: ‘I am catching them whenever I get time, and I am now satisfied it is a solution of the grasshopper problem.” The machine is much more efficient upon bright, warm days, when the insects are upon the vegetation and active, than upon cold, cloudy days, when the young locusts are resting sluggishly upon the ground. DeEscripTION OF PtatTeE VIII.—Hopper-dozer at work in Kafir-corn on north ranch of Ball & Goddard, Edwards county. J. H. Smart, superintendent; Wm. Weber, foreman. Native grasshoppers had entered this corn from newly mown alfalfa field on the left, outside of view in picture. (Photographed by Hunter, July 20, 1898.) 60 GENERAL SUMMARY. In 1891, in Minnesota, more than 200 of these machines were made and used. It was estimated that 5000 bushels of young grass- hoppers were caught with these machines. SPRAYING. There is one crop which at the time of alfalfa harvest is too high for protection by means of the hopper-dozer; that is the growing corn. This can be readily protected by driving along the outside corn rows and spraying the corn thoroughly with Paris green, one pound to sixty gallons of water. The insects entering will feed upon the leaves and die. These rows should be sprayed about once a week until the grasshoppers have disappeared. Having neglected to place in my equipment a spraying pump, I was unable to conduct experi- ments and note results. I give this as a remedy without trial, and be- lieve it will prove a sure destruction to all. locusts entering the corn. The five or ten rows of corn thus sprayed had better be removed be- fore the field is pastured. I see no reason, however, why the ears of corn when husked would be in any way dangerous. SUMMARY. Native grasshoppers require certain conditions for their increased multiplication. The grasshopper prevalent this year is frequently called the yellow locust, better known as the Differential Locust. Large areas of undisturbed soil for deposition and protection of eggs, attended by early spring vegetation for nourishment of newly hatched young, are the essential conditions. | These native grasshoppers, especially the Differential Locust, show decided preference for cultivated ground and roads adjacent to places ‘suitable for oviposition. This Differential Locust, contrary to suppo- sition, does not require soil entirely free from grasses for oviposition, but will sometimes deposit its eggs among the roots of a bunch of buffalo-grass. Wheat drilled in stubble or unplowed ground, alfalfa permitted to grow on the same undisturbed ground from year to year, furnish ideal surroundings for the welfare and productivity of native grasshoppers. Wherever these conditions are found, in any state in the union, native locusts will appear in numbers directly proportionate to the area containing their required conditions. Wheat, oats and barley were not disturbed this season in vicinities where large areas were devoted to alfalfa. Differences of opinion exist concerning the best methods of soil culture for wheat. If grasshoppers damage crop, thorough tilling and packing of the soil previous to sowing seed should be the only method used. Unless this is done, harvesters may be produced on the same ground who will reap the rewards. Crop profits are figured from the granary. GRASSHOPPER LAW. 61 In alfalfa culture, if the grasshopper proves an incentive to proper cultivation, the insect is a blessing in disguise. Disking alfalfa fields in the early spring, after the frost has left the ground and before vegetation has well started, increases the yield of the first crop one-third; matures the second crop earlier, and brings from it an equally increased yield ; destroys the native grasshopper eggs placed therein, and kills the native grasses which frequently threaten to reclaim the field. The young grasshoppers that may hatch from undisturbed ground must be destroyed. They are raised upon the farm where found. The only way to go out of this kind of stock-raising is to kill all the stock. This can be readily done by means of the hopper dozer. The time to use this is as soon as the young insects begin to hop in the spring. Every female grasshopper killed means one hundred grass- hoppers less the next year. Contagious diseases and mortal enemies among the lower animals have thus far failed to keep these native locusts in check. It has been demonstrated that man can do it; upon him rests the responsi- bility. Proper cultivation and vigorous and intelligent use of the hopper-dozer will free any farm of this locust, and the labor expended will be the best investment of the year. The greatest good, the per- manent reduction of this locust, can be brought about by the active, persistent codperation of all concerned. GRASSHOPPER LAW. Below we print an extract from the General Statutes of Kansas re- lating to the destruction of grasshoppers, and found in volume 2 (1897 edition), on page 939. CHAPTER 120, SEss1on LAws OF 1877. Sec. 5. In any senatorial district in the state of Kansas where trouble is anticipated from the ravages of young grasshoppers in the year 1877, and any subsequent year thereafter, it shall be lawful for the counties in said senatorial district to codperate together in the way and manner herein provided for the destruction of the same. Sec. 6. The chairman of the board of county commissioners in the county having the largest number of inhabitants in a senatorial district, where two or more counties form said senatorial district, may notify the chairman of each of the boards of county commissioners of the remaining counties in said district of the time and place when the chairmen of the several boards of commissioners of the respective counties forming said senatorial district shall hold a joint meeting. Sec. 7. At such meeting two of their number shall be chosen to act as chair- man and secretary, and the proceedings of the meeting shall be published in all the newspapers printed in the senatorial district. Sec. 8. Said meeting shall designate the manner of procedure by road over- seers, and what day or days the young grasshoppers should be driven from the cul- tivated Jand onto the unburnt prairie or places of destruction, and shall also 62 GRASSHOPPER BIBLIOGRAPHY. designate on what day or days the grasshoppers shall be destroyed, by burning or otherwise, in said senatorial district, giving at least ten days’ notice of the same by publishing in the newspapers of the said district. Sec. 9. The board of commissioners of each county shall notify the road overseers of said county of the time fixed upon by the joint meeting for the driv- ing and burning, or destroying by other means, of the grasshoppers in the dis- trict; said notice to be given to said road overseers as soon as practicable after the same shall have been determined by the joint meeting. Sec. 10. Said road overseer shall immediately notify the residents of his road district of the time designated and the manner of procedure, in order to carry out the provisions of this act. He shall also specify what tools or implements will be required by each resident in performing the labor required by him; and such no- tice may be enforced the same as in the act authorizing road overseers to warn out the residents to perform road labor; and a refusal shall subject such persons refusing to the same penalties as ere provided by law in such cases. Sec. 11. The road overseer shall direct the manner of performing the labor, and have the supervision of the same, and shall keep a list of the names of those who shall perform labor, and shall certify to the number of days’ work performed by each, and shall place such certified list in possession of the board of county commissioners of his county. Sec. 12. It shall be lawful for two or more senatorial districts to codperate together under the provisions of this act, on a basis of action which they may agree upon. BIBLIOGRAPHY. Melanoplus differentialis. Caloptenus differentialis. Unurr!, MS. (1863). WatsH, Ritey, Amer., Ent., I (1868), p. 16; ibid., I (1869), p. 187. THomas, Proc. Acad. Nat. Sc. Philad., 1871 (1871), p. 149. Gutover, Ill. N. A. Ent., Orth. (1872), pl. vit, fig. 12, pl. 1x, fig. 4, pl. x1, fig. 6. Tomas, Rep. U. S. Geol. Surv> Terr., V (1873). p.. 166, pl., fig. 5; Key Ill. Orth. (1874-75), p. 3. Ritey!, Ann. Rep. Ins. Mo., VII (1875), pp. 124, 173, fig. 33; ibid., VIII (1876), pp. 153, 154. Purnam, Proc. Dav. Acad. Sc., I (1876), p. 266. THomas, Bull. Ill. Mus. Nat. Hist., I (1876), p. 68. Wurrman, Grasshopper (1876), p. 19, fig. Bruner, Can. Ent., [X (1877), p. 144. Brssry, Bienn. Rep. Iowa Agric. Coll., VII (1877), p. 209. Tuomas, Rep. Ent. [ll., VI (1877), pp. 44-45. Rizry, Loc. Plague (1877), pp. 89, 194, 198-201, fig. 34; Amer. Nat., XIT (1878), p. 284; Rep. U.S. Ent. Comm., I (1878), pp. 220, 223, 225-226, 298, 298-299, 301, 327, 447, 459, figs. 32, 110, pl. rv, fig. 1. Tuomas, ibid., I (1878), p. 42; Bull. U. S. Geol. Surv. Terr., IV (1878), p. 500. Ruivey, Bull. U.S. Ent. Comm. III (1880), p. 39; Amer. Ent., IIT (1880), p. 220. Tuomas, Rep. Ent. Ill., [IX (1880), pp. 91, 96, 127-128, fig. 24; Rep. U. S. Ent. Comm., II (1881), pp. 106-107. Linrnrr, Ins. Clover (1881), p. 5. Os- BoRN, Amer. Nat., XVII (1883), pp. 1286-1287. Bruner, Rep. U.S. Ent. Comm., IIT (1883), pp. 54, 60. Forsers, Rep. Ins. Ill., XIV (1884), p. 23. Rizey, Stand. Nat. Hist., IT (1884), pp. 194-195, fig. 271. Ossorn, Bull. Iowa Agric. Coll. Dep. Ent , IIT (1884), p. 83. Bruner, Rep. U. S. Ent., 1884 (1885), p. 399. Rizey, Amer. Nat., XX (1886), pp. 558-559. Coox, Beal’s Grasses N. A., I (1887), p. 373. WEED, Bull. Ohio Agric. Exp. St., Techn. Ser., I (1889), pp. 40-41. Lue- GER, Rep. Agric. Exp. St. Minn. (1889), p. 340, fig. 16. OssBorn, Ins. Life, IV (1891), pp. 50, 51, 55; Rep. Ent. Soc. Ont., XXII (1891), pp. 70-73. Ossorn, Goss, Bull. lowa Exp. St., XIV (1891), p. 175; ibid., XV (1891), p. 267. Riney, Ins. Life, ITV (1891), p. 145; Bull. Div. Ent. U.S. Dep. Agric., X XV (1891), pp. 30-31, fig. 8. OsBorn, ibid., XXVII (1892), pp. 59-60. Ruiney, Ins. Life, IV (1892), pp. 323, 393, 401. GRASSHOPPER BIBLIOGRAPHY. 63 Acridium differentiale. TxHomas, Trans. Ill. St. Agric. Soc., V (1865), p. 450. Cyrtacanthacris differentialis. Waker, Cat. Derm. Salt. Brit. Mus., IV (1870), p. 610. THomas, Proc. Acad. Nat. Se. Philad., 1871 (1871), p. 149. Pezotettix differentialis. Svat, Bih. k. Sv. Vet.-Akad. Handl., V (1878), No. 9, p. 14. WEED, Misc. Ess. Econ. Ent. Ill. (1886), p. 48. Hunt, ibid. (1886), pp. 122-123, 126. Weep, Rep. Ent. Ill., XV (1889), p. 40. Garman, Orth. Ky. (1894), pp. 4, 8. Melanoplus differentialis. Bruner, Bull. Washb. Coll., I (1885), p. 139: -ibid., I (1886), p. 200. Riztey, Rep. U.S. Ent., 1885 (1886), p. 233. CoaquiLuertr, ibid., 1885 (1886), pp. 295, 297. Bruner, Bull. Div. Ent. U. S. Dep. Agric., XIII (1887), p. 33; Rep. Ent. Nebr. Bd. Agric., 1888 (1888), p. 88, fig. 4. Comsrock, Intr. Ent. (1888), pp. 108, 111, fig. 100. Smrrx, Bull. N. J. Exp. St., K. (1890), p. 41. Bruner, Bull. Div. Ent. U. S. Dep. Agric., XXII (1890), p. 104. Brarcx- LEY, Can. Ent., XXIII (1891), p. 99. Bruner, ibid., XXIII (1891), p. 193; Ins. Life, III (1891), p. 229. Wessrer, ibid., IIT (1891), p. 300. Bruner, ibid , IV (1891), p. 22; Rep. Ent. Soc. Ont., XXII (1891), p. 48; Bull. Div. Ent. U. S. Dep. Agric., XXIII (1891), p. 14. Ossorn, ibid., XXIII (1891), p. 59. Bruner, Rep. St. Bd. Agric. Nebr., 1891 (1891), pp. 243, 307, fig. 84. McNerri, Psyche, VI (1891), p. 74. Smrru, Bull. N. J. Exp. St., XC (1892), pp. 4, 31, pl.1. Rivey, Ins. Life, IV (1892), p. 393. Kerttioaa, ibid., V (1892), p.116. Wererp, Can. Ent., XXIV (1892), p. 278. Ossorn, Proc. Iowa Acad. Sc., I, Pt. ii (1892), p.118. Ket- LOGG, Inj. Ins. Kans. (1892), p. 42, figs. 22, 23a. Bruner, Bull. Div. Ent. U.S. Dep. Agric., XX VII (1892), pp. 32-33; ibid., XXVIII (1893), pp. 15-17, fig. 5; ibid., XX X (1893), p. 35. Ossorn, ibid., XXX (1893), p.47. Bruner, Publ. Nebr. Acad. Sc., IIT (1893), p. 27; Rep. Nebr. St. Bd. Agric., 1893 (1893), p. 461, fig. 103. OsBorn, Ins. Life, V (1893), pp. 323-324; Papers Iowa Ins. (1893), p.58. Bruner, _ Ins. Life, VI (1893), p. 34. Ossorn, ibid., VI (1893), pp. 80-81. Bruner, Rep. St. Hort. Soc. Nebr., 1894 (1894), pp. 163, 204, fig. 67; Bull. Div. Ent. U. S. Dep. Agric., XX XIT (1894), p. 12; Nebr. St. Hort. Rep., 1895 (1895), p. 69; Nebr. St. Ag. Rep., 1896, pp. 120, 121, figs. 20,21. ScupprEr, Proc. U.S. Nat. Mus., Vol. XX, pp. 349-354, Pt. xxiii, figs. 3, 4, 1895 (1897). Snow and Hunter, Bull. Dept. Ent. Univ. Kans. Oct. 1897, pp. 9, 10, pl. 11, fig. 4, 1897. Luacrr, 3d Rep. of Ent. of Minn. for 1897 (1898), pp. 204-206, figs. 130, 131. TECHNICAL DESCRIPTION.* The largest of our species of Melanopli, and heavy bodied; excepting the hind Jegs and the lateral lobes of the pronotum, the general color is a nearly uniform brownish testaceous, becoming paler testaceous in specimens from arid regions: in those from Nebraska, Kansas and Colorado it is sometimes a blackish green, while in those from Illinois and Indiana it is often of a dark brownish green. The head has sometimes a pair of dusky, divergent stripes, passing from the pos- terior corners of the fastigium backward across the vertex, and when these are present there are often other but irregular streaks of similar tint on the genz and clouds over parts of the face; the vertex is gently arched, more gently in the female than in the male, with a broad interval between the eyes, the fastigium broadly and not very deeply impressed: frontal costa broad but narrower than the inter- space between the eyes, percurrent, equal except for a slight expansion be- low, broadly and shallowly suleate below (and including) the ocellus, punc- tate: eyes moderately prominent, short, not a great deal longer than broad: antenne fulvo-testaceous, nearly twice as long (male) or fully half as long again (female) as the pronotum. Pronotum subequal, the metazona ex- *S. H. Scudder, in Proc. Nat. Mus. Vol. XX, pp. 350-352. 64 TECHNICAL DESCRIPTION. panding somewhat, the disk of the prozona sometimes (but not always) very feebly tumid, the front margin feebly convex, the hind margin obtusely and roundly angulate, more obtusely in specimens from the Pacific coast than in others, the median carina distinct and sharp on the metazona, less prominent but. distinct on the anterior half of the prozona, still less distinct (occasionally sub- obsolete) between the sulci; prozona subquadrate in both sexes, smooth, divided in the middle, and barely before the middle of the posterior half, by sulci, the principal suleus bent forward in the middle by the posterior emargination of the prozona, the metazona plane finely subruguloso-punctate; lateral lobes nearly ver- tical, separated from the dis] by a well-rounded angle nowhere forming distinct lateral carinze, marked next the upper limit on the proxona by broken blackish patches, frequently reduced to a pair of short, oblique black dashes, one in either longitudinal half of the prozona, each in a clearer field, and also by the black- ening of the sulci in this region. They are sometimes accompanied by slender, oblique, parallel black lines lower down, the hinder the lower: the pleural incis- ures are also heavily marked in black. Prosternal spine rather long, conical as seen from the side, bluntly cylindrical as seen from in front, a very little retrorse. Tegmina at least reaching (female) or distinctly surpassing (male) the hind femora, absolutely free from maculation, the narrowest apical portion about half as broad as the broadest subbasal portion; wings pellucid or (in darkest forms) very feebly infumated, feebly and narrowly opaque along the costal margin, the veins and cross-veins mostly brownish fuscous. Fore and middle femora of male heavily bullate, the hind femora stout and rather short, moderately tumid, generally fulvo-testaceous, sometimes flavo-testaceous beneath, the outer face with alternate, fulvo-testaceous and black, narrow, equal fish-bone markings, the black rarely interrupted in the middle, the upper inner face with small basal and large median and postmedian black patches, the gen- icular are black on both inner and outer sides; hind tibiae yellow or ful- vous (occasionally in California bright coral red), with a postbasal narrow black annulus (in dark specimens more or less infuscated beyond it), the spines black to their very base, ten to eleven, rarely twelve, in number in the outer series. Extremity of male abdomen heavily clavate, the supra-anal plate subclypeate, obtusely angulate at apex, the margins feebly and broadly elevated and the median portion correspondingly elevated and bearing on its summit a moderately shallow, longitudinal sulcus, tolerably broad and subequal on the basal half, narrowing and with falling walls apically; furcula completely absent or indicated only by a thickening of the last dorsal segment at their proper posi- tion; cerci very large and coarse, laminate, boot-shaped, the basal half subequal, punctate and straight, beyond expanding and at the same time feebly bifurcate, the upper fork as long and more than half as broad as the base, feebly invurved, strongly upcurved, apically tapering slightly and well rounded, the lower fork at right angles to it, forming only a rounded, downward and posteriorly projecting lobe, so that the apical margin of the whole is deeply and roundly emarginate below, the whole surpassing a little the supra-anal plate: infracercal plates wholly obscured; subgenital plate short and broad, scarcely so broad apically as long; the apical margin thickened, but hardly otherwise either elevated or pro- longed, entire; upper valve of ovipositor abruptly upturned apically and sharp acuminate, the upper outer carina feebly serrate. Length of body, male 39 mm., female 41 mm.; antennae, male 18 mm., female 16 mm.; tegmina, male 32 mm., female 34.5 mm.; hind femora, male 20 mm., female 23 mm. Some specimens, especially from the north (Illinois, e. g.), are hardly more than half this size. PART II. ALFALFA AND BEES. ‘ASOOVUAS ‘ONIAM ‘H 'f ‘HONGTYOTA ‘XUNOO ‘fF CL “NC ‘SHUINVIdV SVSNVH ANOS *VIXOdNG ‘MATIIN ‘WA “ALIO NAGUVYD ‘NOLIN *a *M INTRODUCTION. 1 fas industry, productivity, adaptability, and utility, the class Insecta produces none superior to the honey-bee. The head secretesa lactic fluid; the tongue collects nectar; the honey stomach begins the trans- formation of nectar into honey; the abdominal plates manufacture wax, and the posterior extremity is the seat of the defensive organ, the gland of which secretes formic acid. Every available bit of space is taxed to produce its required quota of valuable material. In the economy of nature, the bee is depended upon to care for itself and its offspring. Should it be considered a source of profit, intelligent management must be added to secure margins; and it can be safely said the depth of these margins is directly proportionate to the amount of time. thought and attention invested. This super- vision does not consist alone in the manipulation of the hive and its occupants, but includes, as well, provision for requisite pasturage from which ample supplies may be gathered. It has been deemed expedient to place in this connection a treatise upon practical methods in the manipulation of bees. “A Year with the Bees” has been ably prepared by Mr. A. H. Duff, of Larned, Kan., who has devoted his attention almost exclusively to bees and beekeep- ing for the past thirty years. His early experience was gained in Ohio. This enables him to speak advisedly upon methods peculiarly adapted to Kansas. Mr. Duff, however, needs no introduction to the apiarists of Kansas, since he has conducted for years the apiary de- partment of the Kansas Farmer, and is a regular correspondent upon apiculture for a number of other leading journals, During the past summer the writer was placed in an excellent laboratory, the field, for observation and study of alfalfa as a honey plant, and the action of the bees thereon. Many apiaries in the alfalfa region were visited, observations made upon the conduct of bees toward alfalfa grown under varied conditions, and correspondence car- ried on with leading apiarists from all parts of the state. The quali- ties of alfalfa honey have also been chemically compared with the other leading varieties of honey, by Dr. E. C. Franklin, of the de- partment of organic chemistry, a well-known authority upon sugar analysis. Doctor Franklin was assisted by Mr. J. C. Swayze, an ad- vanced student of that department. ( 67) 68 VARIETIES OF BEES. The pages following show that alfalfa, under Kansas conditions, produces a quality and flow of honey surpassed by no other plant. A few speak with great favor of Alsike clover, but the observations upon this clover have in each case been made upon small plats of ground, The plant is by no means generally cultivated in the state. The greatest enthusiasm manifested upon apiculture is found in localities where large areas of alfalfa exist. The wives and daughters of many agriculturists and of some merchants, find pleasure and profit in caring for bees. Many keep bees simply for the table lux- ury derived. The bees gather the nectar from the blossom, and at the same time insure the formation of seed where the blossom was. The results noted show that the seed crop of alfalfa upon which bees worked was 662 per cent. greater than the crops taken from alfalfa dependent upon other insects for pollination. Many flowers cease to secrete nectar after being pollinated. Alfalfa continues to secrete nectar until the blossom begins to wither. The only objection found with alfalfa is the not infrequent removal of the plant for hay during the blooming stage. It is no unusual thing, however, to find within range of the apiary several thousand acres of alfalfa. Here, the second crop is in bloom before the first is all in the swath, and this continual blossoming places the coveted nectar before the bee from the 15th of June until the middle of October. This publication is sent forth with a desire that the resources of the alfalfa plant may receive the attention due them, and that those unacquainted with the honey-bee may become familiar with its good qualities, may appreciate the luxuries of the hive, and find in the bee not only a source of profit, but also a work of pleasure. VARIETIES OF BEES. The common name, bee, is applied to many forms of the order Hymenoptera. It is not within the province of this work, however, to enter into a discussion of the genera of bees within the order, but simply to confine the treatise to the species technically known as Apis mellifica, commonly spoken of as the honey-bee. It is taken for granted that the name honey-bee is sufficiently familiar to require no further definition. Under this head, however, we find a number of kinds of bees; some speak of them as varieties, others as distinct species. It will suffice for our purpose, at present, to consider all as varieties of the one species, Apzs mellifica, and to refer to each variety by the name popularly known. ~The varieties of bees which have been brought to this country, for the honey-bee is originally a foreigner, are the brown or German bee, VARIETIES OF BEES. 69 the Italian, the Egyptian, the Cyprian, the Syrian, the Palestine, the Carniolan, and the Tunisian. Believing that the interest of the bee- keepers in Kansas will be centered around the German bees, the Italians, Carniolans, and Cyprians, I will speak in detail of these only. GERMAN BEES. The German or brown bee was the first bee brought into this country. In the early settlements of North America these bees, after introduction, went in advance of the settlers, notifying the Indians of the encroachments of civilization. Thomas Jefferson, in his history of Virginia, says that the natives called the honey-bee “the white man’s fly”; so the German bee not only established the first honey factories in the United States, but also the first institutions of civilization in many localities. This bee is a native of middle and northern Europe. The species now among us is referred to as the black or brown bee. The charac- teristics of this bee may be summed up as follows: It has a highly irritable temper: by some it is termed cross, in that it not infrequently leaves the hive to attack a passer-by. This disposition varies some- what with different strains. It is not a good neighbor, but frequently quarrels with others in the bee village. When long under manipula- tion it is liable to stampede, if such a term may be used, and the re- sults of this turmoil may be, at least, disagreeable. It is not the best defender against robbers, it is slow to learn a new locality, rather slow in building up in the spring, easily discouraged. It is a good honey gatherer, flies early, ranks high as a comb builder. For those who market comb honey, this bee furnishes artistically white-capped comb. Its irritable disposition frequently overshadows its commend- able qualities and loses it many friends among the apiarists. es CARNIOLAN BEES. The Carniolans may be considered the other extreme in disposi- tion. They are notably gentle. In color they resemble the German bee; in structure, however, they are larger and more robust. The rows of dense silvery white hairs on posterior portion of each of the abdominal segments mark these bees distinctively. This bee seems to obtain the good qualities of the German bee and none of the bad. It is a good comb builder—makes beautifully white-capped comb This bee uses little propolis, winters well, is long-lived. While it isa hard-working bee, a great accumulator of honey, it tends to swarm frequently. This objectionable feature, however, can be controlled by the intelligent apiarist. ‘aHMAOM' NVITVIT ‘aNOUG NVITVIT ; ‘"NaaO’) NVITVIT ‘MAMYOM NVTIOINUVO ‘INOUG NV TIOINYVO ‘ooIMg posielug (‘TBUISO) “II ALVId VARIETIES OF BEES. Tk ITALIAN BEES. The United States department of agriculture in 1860 introduced these bees from Italy, and through the energy of this department they have been widely spread. For this reason they are well and favorably known. The chief distinction in the appearance of this bee is the color. The hairs spoken of previously are present, but are yellow; the first three segments of the abdomen are for the most part tawny yellow. Leather or tan colored is a term sometimes used for this shade upon this bee. The first ring of the abdomen faces toward the front and the band upon this may be overlooked. The second segment is smooth yellow in front ; this is hairless, because it passes beneath the segment in front of it when the body contracts. The central portion of this segment is covered with yellow hairs and the body at. this point is yellow. The back portion of this segment is glossy black and covered with very fine hairs. The markings of the third ring are the same as the second. The fourth and filth segments are black, but covered with yellowish hairs. The sixth segment is black, and bears very few hairs. The queens vary in color. The one shown on plate 11 (page 70) was a beautiful queen, abdomen yellow, with small brown spots upon upper median line of each segment of abdomen. The drones are smaller than the drones of the black bee and darker than the Italian workers. These bees are usually gentle, bear manipulation, and remain firmly upon the comb when this is being handled. They fight hard, long and successfully for their homes; but this strength and this tenacity are sometimes turned in the wrong direction, and these same defend- ers become bold and persistent robbers of others’ gains. They repel the moth with success. These bees are hard workers, at it early and late, excellent honey collectors, but care is required to avoid too great attention on the part of the bees in rearing young with the surplus stores. CYPRIAN BEES. A few colonies of Cyprians are to be found in this state, but not in representative numbers nor in numbers sufficient to draw conclu- sions upon them in thisclimate. Their temper is anything but cheer- ful when thoroughly aroused; their use of their weapon of defense is excelled by none of the varieties previously mentioned. This fault has prevented their general adoption. This variety has the largest honey record per hive for a single season. They winter well and are good defenders. Their comb is better for extracted than for comb honey. They fill the cups so full that cap and honey touch, giving the comb a watery appearance. They build comb well. They are 72 SOCIAL ECONOMY. smaller than the German bee. Orange bands are apparent upon the first three abdominal segments. They are exceeding thrifty, and are said to thrive where others fail entirely. A statement of the qualities of these bees has been given without an attempt to point out the one best adapted for this region. The Italians are justly popular. The black bee is still with us. The Carniolans and Cyprians are comparative strangers within our bor- ders. If an opinion were asked, it would be that the Italians and Car- niolans will merit every attention in this locality. Hybrids are being used to some extent, but a discussion of the various objects sought and qualities attained by this process will hardly apply here. SOCIAL ECONOMY OF THE HIVE. This communistic society contains three divisions, unequal in number but of equal importance in function. The colony is com- posed of a queen, the impregnated female, the drones or male element, and the workers or undeveloped females. THE QUEEN. Before her true function was known she was termed the “king bee” on account of size. This, the most attractive person- age in the hive, is more frequently ruled than ruler. She receives every attention that can be bestowed upon her by her attendants, the workers, and well they may caress her, for around her centers the ex- istence of the hive. The difference between a queen and a worker is caused by the difference in the amount and the character of the food given each in the earlier stages of development. As Cheshire would say, the workers are weaned and the queen nurses during her lifetime. Queens are developed in two ways, each under different circum- stances. In the natural procedure the queen cell is made, the fertil- ized egg placed therein, and the young larva fed, instead of the bee-bread intended for bees, royal jelly, a substance resembling blane-mange, a food given forth from an active gland in the head of the nurse bees. Should the colony be deprived of its queen, the workers hasten the appearance of a new queen by tearing down the partition walls between three surrounding cells, taking the contents away, and leaving one egg to be fed for the throne. The egg hatches in about three days after being laid. About six days are spent in the worm or larval state, then seven in the quiescent or pupal state. In some cases the workers choose a cell containing a larva for the production of an emergency queen. If the workers choose a worm as a princess, the time from formation of cell to emergence of queen will be shorter than the full sixteen days a number of days equal to the age of larva, including egg period. Cheshire has shown that emergency queens are not equal to queens produced in the natu- SOCIAL ECONOMY. te ral way. He further states that it is highly improbable that the bees ean develop a queen from worker larvee which had begun to be fed bee-bread. The large queen cell extending out from the comb, fre- quently hanging down not unlike a peanut in shape, is easily recog- nized. Queen cells are illustrated in Mr. Duff’s article, further on. A few days after maturity the virgin queen leaves the hive on her marriage flight. She is met high in the air by a drone and fertiliza- tion takes place. She returns to her hive and there remains with the colony. The act of pairing takes place but once in the lifetime of a queen. About two days after impregnation the queen begins laying. If a very fertile queen, she will deposit from two to three thousand eggs daily. While the worker exhausts itself and dies in a few weeks, or months at the most, the queen is of greatest service to her colony for two years, and while she will live longer it is not advisable to retain her beyond that time. THE Drone is developed from the unfertilized eggs placed in cells somewhat larger than worker cells. They develop in about twenty- four days, remaining three in the egg, six in the larval, fifteen in the pupal stage. These are reared in larger numbers during the swarm- ing season. This is nature’s provision for the marriage flight of the queen. Were the drones few in number, it would be very probable that many queens would fail to meet a mate in theair. As it is, many drones are in mid-air searching for mates and the fertilization of the queen is assured. The fact that the drones are reared from unfertilized eggs, the un- mixed blood of the queen, and that workers are short-lived, make the introduction of a queen into the hive a matter of much importance. For it will be readily seen that with the introduction of a new queen, new drones of her exact strain soon appear. If the queen has been tested, and found to be purely mated, then the queens reared from the eggs will be pure, and these will now mate with pure-bred drones and their progeny will be a pure strain. The workers of the queen introduced will be pure bred and will shortly replace the native work- ers, who will have lived out their existence. This refers to a single colony. It must be noted in this connec- tion, however, that in an apiary one colony of an inferior strain may contaminate many other colonies in a single season by sending forth drones to breed with the pure queens of the other hives in the apiary. THE WorKER. This is the bee familiar to all of us; the one re- spected for its business air, as well as its powers for defense. The writer has realized the industry and activity of this marvelous little 14 BEE PRODUCTS. honey gatherer when endeavoring to take its photograph while it was at work upon some of its favorite food plants. The number of individuals in a good colony ranges from 20,000 to 80,000, according to the time of the year. The number can never ex- ceed the laying powers of the queen for the season, and rarely, if ever, equals that number. Workers mature in about twenty-one days, spent in the following stages: Three days in the egg, five in the larvee state, and thirteen as a pupa or chrysalis. The division of labor among the workers is discussed in another portion of this work. A young bee is easily known by its pale color and lack of strength. In a few days it grows larger, develops strength and color, and is well covered with hair. The aged worker is known by its tattered wings and bald body. The average life of a bee in the working season is about five weeks. BEE PRODUCTS. The bee is capable of bringing to his hive four things: Bee-bread, propolis or bee-glue, royal jelly, and honey. In addition to these, the bee carries, to be used in case of defense, a gland filled with poison, consisting chiefly of formic acid. The larva has facilities for spin- ning silk. Bee-bread is the pollen gathered from flowers, brought in upon the so-called pollen baskets of the hind legs, and placed, generally, in worker cells, packed down, then covered with honey and the cells sealed. This food is indispensable for the rearing of young brood. Huber demonstrated that young bees could not be reared without this pollen, though this without honey will not support mature bees. Propolis, or bee-glue, is a resinous substance gathered from the buds and limbs of trees. The bees use it to seal over cavities or un- necessary openings in their hives. In the heat of summer this re- mains soft and is used by the bee-moth as a receptacle for eggs. Hives should therefore have as few cavities as possible, and a poor quality of lumber, or boards partly split, should be avoided in the choice of material for hives. Royal jelly is a milk-like substance secreted in a gland within the head of the nurse bees. This is fed to all bees—workers, drones and queens alike. The workers and drones, however, are favored with this highly nitrogenous food during the first days of their larval ex- istence, only until their stomachs become ready to digest the bee- bread. The embryo queens are more favored and this royal jelly is lavished upon them during their development. According to Ches- hire it is the food of queens during their whole lifetime. o~ BEE PRODUCTS. 2) Silk.— The pupal cell is partially lined with silk. This isspun from glands in the head of the larvee. These glands are not used after the bee reaches maturity. Waz is a secretion which in the form of plates appears on the ven- tral surface of the second, third, fourth and fifth segments of the ab- domen. The wax is primarily secreted from the blood-cells as a fluid, and becomes more firm when it is transuded upon the abdominal wax pockets shown in the accompanying figure 1. It was erroneously believed for a long time that wax was pollen digested and then regurgitated. It is what might be termed surplus energy, or a case quite parallel to the secretion of fat in animals. Young bees secrete wax well; old bees produce littlewax. The same may be said regarding fat- producing powers of young and old animals. The production of wax exhausts the strength of the bees, and isa heavy tax upon the stores of honey. The bees intending to produce wax gorge themselves Fic. 1. Secretion With honey, and in about twenty-four hours after- f les. _(En- ; fared Mow Tine ward wax begins to transude and appear upon the hee’) Bienenzei- Wax plates of the abdomen. It requires from ten to sixteen pounds of honey to produce one pound of wax, according to Langstroth. Some authorities state greater weights and some less. When the amount of honey consumed is reckoned, the matter of comb foundation and subject of care of empty comb should receive careful attention. Honey.— The reward most covetedis expressed by this word. The word itself has a pleasing sound. It is the symbol of sweetness. The bee, bearer of this esteemed delicacy, collects nectar from the nec- taries of flowers and sap from the trees—a little here, a little there. These fluids in their watery state are taken into the honey sac (pl. lll), corresponding to the crop of other insects; instead of passing on into the stomach, the sweet fluid is retained here until the hive is reached, A reference to plate 11 at this point will make the subject of the honey sac, stomach mouth and stomach sufficiently clear for the purpose of this discussion. The watery nectar of flowers and sap of trees is reduced to the consistency of honey, either by secretion of the water from the bee’s honey sac, or evaporation caused by heat and currents of air in the hive. These currents of air, forced by the fanning of bees’ wings up through the hive and over the uncapped honey, do much toward ripening and removing unpleasant ( probably volatile) flavors. Formic acid is found in honey; the exact form and amount have not been satisfactorily determined by chemists. Pirate ITI. Digestive system of bee, magnified ten times (after Cheshire). A, Horizontal section of body; Up, labial palpus; mz, maxilla; e, eye; dv, dv, dorsal vessel; v, ventricles of the same; No. 1, No. 2, No. 3, salivary gland systems, J, 2, 3; @, csophagus; pro.t, pro- thorax; mesa.t, mesathorax; meta.t, metathorax; g, g, ganglia of chief nerve chain; n, nerves; As, honey sac; p, petaloid stopper Of honey sac or stomach mouth, c. s, chyle stomach; bé, biliary or malpighian vessels; si, small intestine; 7, lamelle or gland plates of colon; li, large intestine. BEE PRODUCTS. T7 While attending the National Beekeepers’ Association, at Omaha, the writer was greatly impressed by the display of the varieties of honey. This publication being then in course of preparation, Com- missioner Whitcomb and Superintendent Stilson, of the department of bee industries, very generously furnished five samples, from which the accompanying photograph and chemical analyses were made. The varieties sent and donors were: Basswood ( Zilia americana), from the apiary of L. D. Stilson, York, Neb.; knotweed ( Polygonum pennsylvanicum), E. Whitcomb, Friend, Neb.; white clover ( 7rifo- lium repens), L. M. Whitford, Arlington, Neb.; sweet clover (Melli- lotus alba), L. D. Stilson, York, Neb.; alfalfa and melon bloom, G. D. Swink, Rocky Ford, Colo. The alfalfa (M/edicago sativa) honey was sent direct to the department through the kindness of Capt. J. H. Wing, of Syracuse, Kan. The comparative lights and shades of these varieties of honey are shown in plate Iv, and a careful analysis of the same has been made by Dr. E. C. Franklin and Mr. J.C. Swayze. Their results appear on page 79. As may be inferred from the title of this treatise, the subject of alfalfa honey would come under consideration. J can conceive of no better method of procedure than that of comparison, and upon that plan I have here carried on the investigation. Cheshire’s definition of ideal honey is, “An ideal sample would have a delicate but charac- teristic aroma, a rich flavor, leaving a distinct impression on the back of the palate, and would be of a straw or pale amber color. It should possess perfect clearness, and, as distinct from clearness, brightness due to a high refractive index, with density almost amounting to toughness, so that the air beneath the cork should rise very slowly through the mass upon the inversion of the bottle.” Of the six samples submitted, a competent judge placed the alfalfa honey as the one most nearly approaching this standard. Especially was this so with regard to the color and toughness. The six were inverted in the tubes, and the other five were free from the air bubbles in a short time, as compared with the alfalfa sample. The color of pure alfalfa honey is certainly highly desirable. The six test-tubes on plate Iv, while not showing the exact tints, illustrate the comparative degrees of light and shade. They might be said to range from very light straw (6) to dark amber (2). See next page. The tastes and flavors, largely personal elements, are not suscepti- ble to chemical tests or verbal descriptions; to be illustrated and ap- preciated they must be experienced. * sce ele For DESCRIPTION, SEE BOTTOM OF OPPOSITE PAGE, PLATE IV, -] - a BEE PRODUCTS. ANALYSES OF VARIOUS KINDS OF HONEY. By Dr. E. C. FRANKLIN and J. C. Swayze, Department Organic Chemistry, University of Kansas. Polarization. SAMPLE. —— | emp. Sucrose: ee Water. | Ash. Direct. |Indirect. Basswood. ....:..... -9.9 | -14.9 23° 13.77 % | 76.92 %| 11.31 %| .89 % Knotweed.......... —24.2 | -29. 23° |3.62 %| 75.19 % | 11.41%} .05% White clover........ -10.3 | -12.3 23° {1.54 %| 76.34% | 7.98% | 03 % AMINE i Se a igtess (2 ome EE 23° |4.38 %| 75.76 %| 10.00 % | .05 % Sweet clover........ —16. 21. 93° 13.77% | 78.13%| 71.91%) .05% The above samples are all pure, as is shown by the different exam- inations. Pure honey consists of reducing sugars (d-glucose and d-fructose ) and non-reducing sugar ( sucrose.) Pure honey turns the plane of polarized light to the left, generally less than twenty divi- sions. An excess of this might indicate adulteration with invert sugar. This adulteration can be detected with certainty only when large quantities have been added. The sample of knotweed honey may have a slight excess of invert sugar. Dextro rotation may signify either an adulteration of glucose or a normal amount of sucrose. By inverting the sucrose, the presence of glucose is proven, if it is still dextro rotatory. As none of the samples analyzed were dextro rotatory, there was no adulteration with glucose. The sucrose found is in accordance with that of pure honey. It is determined by polarization before and after inverting. It is also de- termined by the Fehling method. The amount of reducing sugar is determined, the sucrose is then inverted, another titration is made, and the sucrose found by difference. We find that this method gives slightly lower results than the polariscope method. The water and ash are normal. From the results, it would seem that a little dust had settled in the basswood honey and had increased the percentage of ash. Comps Buiupine. The architecture of the hive furnishes interesting study for the artisan as well as the apiarist. Contrary to supposition, however, comb is not constructed with geometric calculation. Cellsare made of various sizes and shapes from the wax secreted. The store or honey cells are some what irregular and frequently deeper than DEscRIPTION OF Pirate IV.—Shades in various kinds of honey: /, bass- wood; 2, knotweed; 3, white clover; 4, alfalfa; 5, sweet clover; 6, alfalfa and melon bloom. 80 BEE PRODUCTS. brood cells. The worker brood cells are the most regular, and average twenty-eight to thirty cells to the square inch. The drone cells are larger, and range about eighteen to the square inch. The structure of the queen cells has been described elsewhere. THE Stine. The accompanying figure illustrates the structure of sting and the gland attending it. This weapon is brought into use, — not alone as protection against intruders, such as man, but is much used against the unwelcome visitors of its own tribe. The formic acid and the other toxic element probably present in the fluid injected from the gland into the body of the one attacked, while discomfiting to us, is quite deadly to bees. This poisonous fluid can be secreted from the blood-cells of the bees, but when it is injected into the tissues, by means of another’s sting, death follows When the honey- bee stings the human flesh the sting generally Fic. 2. The sting of the worker bee, and its appenda- ges. (Enlarged, from Girard.) a, sting ; b, poison-sac ; s ; : ec, poison glands; d d, secreting bags. remains, and the bee, if not killed before, dies on account of the wound caused by the tearing away of the organs con- . nected with the sting. “The third sting is composed of two spears of a polished, chestnut- colored, horny substance, which, supported by the sheath, makes a very sharp weapon. In the act of stinging, the spears emerge from the sheath about two-thirds of their length. Between them and on each of them isa small groove through which the liquid coming from the poison-sac is ejected into the wound. “Each spear of the sting has about nine barbs, which are turned back like those of a fish-hook and prevent the sting from being easily withdrawn. When the insect is prepared to sting, one of these spears, having a little longer point than the other, first darts into the flesh, and being fixed by its foremost barb, the other strikes in also, and ALFALFA AND BEES. 81 they alternately penetrate deeper and deeper, till they acquire a firm hold of the flesh with their barbed hooks. “The muscles, though invisible to the eye, are yet strong enough to force the sting to the depth of one-twelfth of an inch through the thick skin of a man’s hand.” * Various remedies are recommended. It is important to scrape away with the finger nail the inserted sting; it should not be taken away by pinching it between the thumb and forefinger, for this act will force all the poison in the gland down into the wound. The place stung should not be rubbed, since this increases circulation and diffuses the poison over larger surface, consequently making the pain greater. This poison will mix freely with water, so that if the affected parts are placed in cold water the pain will be greatly allayed. Ammonia will neutralize the acid, and its use is recommended. While attending the National Beekeepers’ Association, at Omaha, the writer was much interested in the discussion upon the medicinal properties of the sting. It was strongly advocated by some that the sting of the bee, directed toward the seat of rheumatic pains, would give permanent relief. It was the observation of others, based upon experiments, that no permanent value could be attached to the me- dicinal properties of the sting. In each case the speakers spoke from experience, the speaker being the subject of the experimentation. METHOD OF FERTILIZATION OF THE ALFALFA BLOSSOM. The location of the coveted nectar at the base of the flower, the action of the tongue of the bee and the work of the hairs under the head and upon the breast in placing the pollen upon the stigma are shown in figure 5, and also in figure 4 at 64. The flower gives material aid, by causing the stamens and pistils to spring up and strike the insect. A part of Miiller’s observations upon this point are: If in a young flower we eut through the claw of the carina, the column springs upward to some extent, ee ameter) Pollination: “carrying with it the carina and alee. If in another unexploded flower, we carefully cut through one of the digitiform processes of the alee, the parts re- main motionless; but on cutting the processes of the other side, ex- plosion at once follows. The pouched processes of the carina ( fig. 4.63 andc¢3) are thus sufficient to hold the column down without * Langstroth. —6 82 ALFALFA AND BEES. the aid of the processes of the ale (¢ 2), nor are the latter sufficient after section of the carina. Explosion can therefore be effected equally well by separating the anterior pouches, by separating the digitiform processes, or, finally, by depressing the ale and carina. Tf an insect inserts its proboscis in the middle line between the an- terior pouches and the digitiform processes, or if it stands upon the alee and thrusts its head in the middle line under the vexillum, in either case explosion follows. The stigma (¢ 4) projects beyond the anthers, and, therefore, is the first to strike the under surface of the bee’s body or proboscis ; an instant later the anthers come in contact with an area close around the spot that the stigma touched, dusting it with fresh pollen. The first flower that the insect visits is, of course, not cross-fetilized, but as the bee withdraws from the flower, self- fertilization inevitably occurs. Self-fertilization is undoubtedly effi- cient, for Hildebrand has shown that flowers which wither unexploded when insects are excluded produce seed by self-fertilization. The same author finds two imperfections in the mechanism. One is the possibility of the insect securing the nectar without exploding the flower; the other is that the flower continues to secrete honey after it has been fertilized. This certainly is much in its favor as a honey plant, though somewhat against the visitation of other unfertilized flowers. In the case of the alfalfa, however, it will be seen that the stigma of the ovary is higher than the pollen producing anthers, so that the grains of pollen may all drop to the base of the flower and the ovary go unfertilized; such being the case, no seed would be formed. Small forms resembling seed might be found within the ovary at maturity, but these, not being fertilized, would not germinate. From the shape and size of the alfalfa blossom, it is not probable that cross-fertilization could be safely accomplished by means of cur- rents of air. It becomes evident, then, that outside agencies must be called upon, and the plant must provide for these agencies. The agents in this case we find to be insects, and the reward offered by the plant for favors rendered is a sweet drop of nectar; that is, the flower in an enticing way places a tempting sip of nectar in such a position that when the insect has favored the flower with a few grains of pollen un- consciously brought from an adjoining flower and just as uncon- sciously left, the coveted sip may be enjoyed. It is evident, however, that the first flower visited will not be cross-fertilized. INFLUENCE OF BEES UPON THE SEED CROP. It is well known to every stock-breeder that in-and-in breeding will soon cause the strain to deteriorate, and that infusions of new life are ALFALFA AND BEES. 83 Fie. 4. (Original.) @, cluster of alfalfa with bee feeding. b, bee thrusting proboscis into flower: J, vexillum; 2, ale; 3, carina; 4, reproductive organs (gametangia) : 5, calyx. ec, alfalfa bloom with vexillum torn off: 2, ale; 3, carina ; 4, reproductive organs (gametangia) : ae stigma ; 6, anthers; 7, calyx. d: 1, filament; 2, "anther: 3, style; 4, stigma. e, pistil: 7, ovary; 2, style; 8,stigma; 4,ovules. /, cross-section of pistil: i, detached filament; 2, fil ament sheath: 3, ovary; 4,ovule. g, highly magnified pollen grains. required to keep the desired vigor within the breed. As is the case with animals so it is with plants. It was previously supposed that within each flower were the necessary organs and the means for assur- ing the formation of the embryo within the seed. Darwin, however, has clearly shown that many plants, instead of endeavoring to facili- tate self-fertilization (the forming of seed in the ovary from pollen of same flower), are constructed in a manner to hinder or prevent it. Among this number may be classed the plant under consideration. A careful examination of the accompanying figure and explanations will reveal the fact that outside agencies are required to insure fertili- zation, especially cross-fertilization. The process of fertilization in a typical flower is not complex. The stamens have long filaments which bear the pollen-producing anthers high above the ovary; when the pollen is ripe it falls naturally upon the stigma of the ovary, and fer- tilization of the seed is soon accomplished. 84 ALFALFA AND BEES. The writer conducted a number of experiments upon the fertiliza- tion of the alfalfa bloom. The first work consisted in covering a large number of blossoms with fine cheese-cloth. It soon became evident that this would exclude all insects, and the good services of the bee would not be demonstrated, so that this line was discontinued and another taken up. A large number of representative ripened pods were gathered from an alfalfa field less than one-half mile away from a large apiary, and a like number from another field of much the same soil and, practi- cally, under like conditions as the first field, except that the second field was situated twenty-five miles away from a colony of bees. No bees were observed in the field, and the character of the surround- ings, there being no timber or probable living places, was such as to preclude the possibility of wild bees in the vicinity. The pods from each locality were carefully opened and number of seeds in each counted. The results and comparison to be made therefrom are cer- tainly of interest. Seeds taken half mile from bees; Seeds taken twenty-five miles from 87 pods examined. hees; 80 pods examined. No. of pods No. of pods No. of seeds bearing that | Total number] No. of seeds bearing that | Total number in pod. number of of seeds. in pod. number of of seeds. seeds. seeds, 0 0 0 0 2 0 if 1 1 al 8 8 2 5 10 2 18 36 3 8 24 2) 20 60 4 14 56 4 15 60 5 18 90 i) i 35 6 iil 66 6 5 30 7 15 105 7 3 21 8 9 uz 8 i 8 9 2 18 9 0 0 10 4 40 10 1 10 Totalsce- 87 482 Motalsiaeer 80 268 Average number of seeds in a_ pod, 5.58+. Average number of seeds in a pod, 3.35. Seeds Seeds plump; pods numerous in cluster; pods | in at least one-third of the pods were small having several spirals. and shriveled; pods few in a cluster; short, with but few spirals. Per cent. of increase of the first field over the second, 663. ALFALFA AS A HONEY PLANT. My observations upon this subject during the past season have been that it will yield the greatest amount of nectar under circum- stances which tend to give the plant the most vigorous growth ; that is, proper amount of heat and moisture, upon suitable soil. If the plant is upon upland, dry weather will affect the secretion of nectar before it will be affected in the valley, such as the Arkansas valley, ALFALFA AND BEES. 85 where the roots of the plants extend to the water. In September the bees were busy in the alfalfa in the Arkansas valley; while, on the higher ground of one of the counties on the eastern border, I visited a beautiful piece of alfalfa near by an apiary, but no bees were found. They were at that time flying over the alfalfa to the knotweeds be- yond. A strong point in favor of this plant, as shown by Miiller, is that it continues to secrete nectar as long as the blossom flourishes. A greater part of the alfalfa produced in this state is cultivated for forage, and, since for this purpose it is cut while in full bloom, the honey crop is materially less than it would be if alfalfa were allowed to mature. During a dry period bees will fly over alfalfa fields in bloom to a field which has been irrigated a few days previously and has begun to bloom. The relation existing between alfalfa and apiculture can well be obtained by a comparison of the tables following, the data for which Mr. F. D. Coburn, of the state board of agriculture, has kindly fur- nished. The attention of the reader is called to the yield of honey per hive in counties where larger acreages of alfalfa exist, as compared with the yield per hive where the crop is much less. 86 ALFALFA AND BEES. CoMPARATIVE TABLE, showing alfalfa acreage, stands of bees, pounds of honey produced, and the value thereof, in Kansas, for the two years 1897 and 1598. 1897. 1898. Seeiesa Acres | Stands | Pounds Value Acres Stands | Pounds of of of of [o) of of : alfalfa. bees. honey. honey. alfalfa. | bees. honey. The State ..... 171,334 44,345 | 534,925 | $80,238 75 | 231,548 60,941 | 622,703 | Allens eclesces: 9L 526 | 4,295 $644 25 633 416 1,365 Anderson . ... 124 508 3,721 558 15 197 666 3,437 Atchison .:.... 345 546 | 4,516 681 90 171 1,391 12,579 iBanberwcetes 1,453 ADM oats ltacecot eee 1,026 50 Barton soso 1,565 De a Payee! rcepactee irs LOTS an bs cll oe eee anes Bourbon ...... Bit 1,512 | 10,153 1,522 95 98 1,385 7,778 Brownl.ss sees 5U7 1,421 | 23,231 3,484 65 638 2,633 34,317 Butlerwcccsen: 7,755 638 | 6,904 1,025 60 12,029 939 8,295 Ghasecs.sc2 thee 5,753 710| 8,830 1,324 50 7,298 605 1,849 Chautauqua .. 3,481 299 3, 694 553 65 3,998 501 929 Cherokee...... 6 2,046 | 23,058 3,458 70 37 1,476 6,286 Cheyenne ..... GODS ae ies | raee lerenctorstct| etaysray etek cere 652 100 lanles aoe 775 P| Ne ei ee eae eG DLOh |= eer ae Gl nee eee Clay. 207 481} 8,303 | . 1,245 45 381 1,328 7,616 Cloud ane. 4,696 689 | 8,763 1,314 45 5,729 744 10,614 Coffey-sc.<.es5. 757 628 | 5,912 886 80 943 | 736 2,466 Comanche .... Tia llanctckcrere taser nate eects ll Leena acre oe PCM PSA Ieee eyo Cowley ........ 3,904 542 | 2,905 435 75 5,722 1,070 9,702 Crawford ..... 19 1,509 19,234 2,885 10 6 1,922 10,566 Decatur....... 3,307 12 60 9 00 4,278 26 380 Dickinson..... 1,745 397 6,429 964 35 2,000 726 5,897 Doniphan..... 44 927 11,013 1,656 45 126 1,343 12,970 ouglas....... 396 1,258 15,112 2,266 80 604 1,605 20,714 Edwards...... 2,930 14 50 7 50 6,523 15 40 10 ea ata 2,837 467 | 3,649 547 35 3,860 523 3,623 INS ee oe LOOM | Vcore Scot are lias celswas call sine sae eet OLGS | chee Ont cere Ellsworth..... CT apacpeaee Saal or re Same 644 5 80 Finney 11,726 533 13,665 2,049 75 11,795 722 50,535 Horde sce 4,277 5 10 1 50 4,508 39 2,020 Franklin ...... 56 1,432 9,954 1,493 10 175 1,628 13,052 Geary. feces ac 559 | 296 | 3,663 549 45 723 388 2,938 OVE weneen NOI SS a Neer een eee Wg ee DOD | Fe siete oe ace arose Graham SOULE eto rayets arerall cmearote eo a v'| kere aeeteoe TU al ere bated ere teteratnre te Granitiemascsdes ¢) | Atos sada eano.npbel Iboanatnablosssy |ndcoJos Dua Eakonsven. |Maeeob aac Gmayecestencse 1,918 8 40 6 00 2,250 29 867 | GTreBleyic. shee hoe oe ee SOR eer eras Gh ceed sans cael cee enennee Greenwood.... 5,359 596 2,831 424 65 9,518 623 2,281 Hamilton..... 2,422 TOGG| Os Senay een Nes 2,779 3 200 Harper........ 527 AO Reeser Cee al utara tah 1 321 74) kta Rie Hiarvey.sec. oc. 1,535 TL 424 63 60 2,143 123 1,468 la ES} - 4,112 Osages conccss: 965 Osborne....... 3,924 Ottawa...... 2,725 Pawnee ...... 365 EA MUOS oi cin1) 3,142 Pottawatomie, 3,294 Pattee cks ese: 76 Rawlins....... 2,401 Reno .. 3,574 Republic ...... 2,667 RCs atesien s,s 1,830 mbes cas tacts 2 1,927 BOOKS. ..ce0 0s 880 LOCC Uae 45 Fussell se. so. 372 Sale Vocus. s- 5,269 SEOth ys cee acs: 206 Sedgwick ..... 4,389 Seward... 0.5 17 Shawnee ..... 1,115 Sheridan ..... 2,096 Sherman ..... 50 Sil hloeess sae ove 1,026 Stafford 352 PIPATIEOU Mette ec lies ao Seis PEP ENIN pee liaen core Sumner ....... 1,516 THOMAS ie 00 227 WOR Os. 202 55-6 396 Wabaunsee ... 4,295 Wallace....... 570 Washington .. 763 Wichita. J..c2 148 Wilson .... 1,180 Woodson.. 843 Wyandotte 79 1897. 189s. Stands Pounds Value Acres | Stands Pounds Value of of of Gr |. wor of of bees honey. honey. alfalfa. | bees. honey. honey 857 8,589 $1,288 35 152 | 1,006 12,073 $1,810 95 622 8,262 1,239 30 7,015 770) 11,958 1,793 70 456 2,352 352 80 1,637 | 388 742 111 30 440 | 2,075 3il 25 oe 255 1,345 201 75 Powwint save Sees pta-adl pene cae eet LON cap teyae st stiateis stemice acer ralpianatnnteate 1,210 22,905 3,435 75 2,551 2,566 36,512 5,476 80 Til 6,615 992 25 ae } 733 1,023 153 45 ey iee Wate oe eke Danae dineenicaan 178 | tO ete Al [eee ed A 475 10,577 1,586 55 5,965 424 17,439 2,615 85 952 10,987 1,648 05 1,908 1,591 6,406 960 19 60 9 4,076 | 19 356 53 40 390 3,102 465 30 3,308 447 | 3,940 585 00 24 500 75 00 597 21 | 1,620 243 CO 85 790 118 50 4,062 93 | 1,538 275 70 1,235 18,350 2,752 50 4,340 | 1,585 17,560 2,634 00 Be yoratals itsralleeccias ea el Monee Scie ores c UTE ccewae lees «ete Pear eaaense 8 40 6 00 998 10 720 108 00 10 22 3 30 4,385 | 13 64 9 614 9,89) 1,483 50 4,012 1,220 17,696 2,654 40 29 205 30 75 2,871 36 1,160 174 00 1,376 22,118 3,317 70 2,806 1,558 14,723 2,208 45 17 350 54 00 yrs | 18 400 60 AONE EPR pee Ree SR aie 405 esac, oe pees MA Wt eo 614 5,771 865 65 6,228 | 712 5,423 813 45 Woah weer Acie eer Sa 248 |. Fat oct seal eee te oot 468 | 3,510 526 50 6,530 769 6,969 1,045 35 Sho eet Fn enn ete itor 15°). e atee hy fais eke sae lente ees were 872 8, 106 1,215 90 1,740 | 1,192 6,739 1,010 85 AN Ma Ce) ae ee 3,334 | Sc eereonte ee eee Pee rete date ee Spall ocean sae Bete IS) 2eeitust (lecaomes Atadieenie 61 | 708 106 20 1,598 | 90 1,225 183 75 Renta eee ort (see oe 5OL | 2 20 3 00 1 Reet aan eee Wea ray cn meta el rey AMPr sd Cire setae all Iota acte eye 56 128 19 20 2,035 / 150 770 115 50 seine BE selec mate Se stil enacanteoese 372 6 100 15 00 Oras. See] | eee eee a a cehion ORM 2 seh a eee eee mere nee 1,709 13,919 2,087 85 6,439 | 1,196 | 12,770 1,915 50 of Seal 104, |b NRE IY lag ese A 685 | 6 140 21 00 1,389 42,777 6,416 55 1,139 2,312 50,389 7,558 35 uh cebiab reales [hu patin sv eet Nei etna rea bien es 1 {oh Paresh cen a (Soe stele 579 | 2,602 390 30 2,462 | 696 | 2,789 418 35 380 3,720 558 00 1,325 312 1,170 175 50 316 3,045 456 75 136 474 4,538 680 70 88 OBSERVATIONS BY APIARISTS. OBSERVATIONS GATHERED FROM KANSAS APIARISTS. During the field-work this year it was the writer’s privilege to meet a large number of the Kansas apiarists at work among their bees. In conversation, many thoughts concerning their observations upon bees and honey plants were expressed. Some of these, unknown to the speakers, were jotted down and appear here. An extensive corre- spondence with some 400 bee-keepers in this state furnishes much valuable reading for those interested. The substance of some of these letters is given here. The number of personal discussions and opinions which appear here might be increased many times, but in so doing the same ideas would be repeated; want of space also forbids; so that the substance of the views of a few apiarists of Kansas upon the principal honey- producing plants of the state, together with a few other facts concern- ing apiculture, are herewith given. J. F. Hucues, Marquette, McPherson county; thirty stands; Ital- ians. ‘They are good workers, and it takes a very strong wind to keep them in their hives. In central Kansas we must depend almost en- tirely on alfalfa for a honey plant. White clover cannot be cultivated successfully here. Buckwheat can be raised in abundance, but.can we afford to raise it for the honey alone? If season is favorable, Italian bees will gather 100 pounds per stand from alfalfa. Alfalfa will grow here entirely without irrigation.” Mrs. C. E. ANDERSON, Salina, Saline county; eighty colonies; Italians and hybrids. “Alfalfa is our main honey plant, and in a favorable season the honey is equal to the best white honey. Buck- wheat makes a good yield sometimes, but the honey is dark and strong. In 1895 I had 1075 pounds fine comb honey from twelve colonies, and several hundred pounds more not in marketable shape.” Wm. Miuter, Emporia, Lyon county; sixty-three stands. “I re- gard alfalfa superior as a honey plant to buckwheat or white clover, as I have known the latter in Ohio. I believe that a few stands of bees on the lands of every Kansas farmer would be a source of revenue, as well as adding quite a luxury to the living of himself and family, and that with less outlay of labor and money than anything else I know, unless it be the cow and hen. Bee-culture very naturally works in with all small fruits, truck patches, fowls of every descrip- tion, and other kindred industries, as an individual can be working at the one, and yet have his apiary so located as to have an eye over it continually during the swarming season. The bee can be wintered in Kansas without any special preparation. Kansas is destined to be- come a great honey-producing state as the alfalfa gains more and more, as it is bound to do.” OBSERVATIONS BY APIARISTS. 89 Mrs. L. A. Carey, Phillipsburg, Phillips county. “Began bee- keeping in 1895; have now twelve colonies. Five colonies this season produced 300 of pounds surplus honey and gave off two swarms. In order to prevent a second swarm, I changed position of old and new swarms in daytime. This causes all the bees in the field to fly to the young colony, builds it up, and weakens the old swarm, so that it does not have the desire for increase. We have to depend on alfalfa here for the main honey crop. It makes a beautiful white honey, like white clover. Alfalfa produces the most honey during the month of July.” H. M. Hit, Paola, Miami county; twenty stands; Italians. “I do not consider alfalfa a success as a honey plant, as we had a large pasture one-half mile from the bees and received no honey. I ama stock-raiser, and have no time for bees, so they receive but little care. We consider white clover the best of honey plants, and buckwheat good at times.” GEORGE YOXALL, Woodston, Rooks county; twenty stands; all Italians or hybrids. ‘Alfalfa is an excellent honey plant here, yield- ing the best white honey, from the 1st of June until the Ist of Sep- tember, and sometimes as late as the Ist of October. Sweet clover may be superior in yield, in a more continuous flow of honey, as there is no check in mowing as there is in alfalfa, and alfalfa does not give the best results on account of taking off the crop just about the time the bees are commencing to work on it, and if there is no seed crop taken, there is not much benefit conferred.” J. G. Brooks, Pleasanton, Linn county; forty stands. ‘Have spent fifty years with bees. I have had no experience with alfalfa. White clover, buckwheat, corn tassel and tree blossoms are good here. Bees should be cared for the same as other stock. Care will pay.” E. K. Terry, Burlingame, Osage county; thirteen colonies. “Have had no experience with alfalfa as a honey plant. We have members in our town bee-keepers’ association who have quantities of alfalfa on land watered by rains, who report favorably. Mr. Arnold has both al- falfa and alsike, and reports that alsike produces much better-flavored honey and more of it than the alfalfa. I consider the sweet clover one of the very best honey plants. It produces white and good- flavored honey. I would keep bees for the pleasure and honey enough to supply the family. I consider honey one of the greatest table luxuries that it is possible to have. I find that with cellar wintering I can bring out a strong colony in the spring on ten to twelve pounds of honey.” M. B. Gvuarp, Beloit, Mitchell county; eleven colonies; all Ital- ians. “Alfalfa honey is better than white clover, because it is richer. 90 OBSERVATIONS BY APIARISTS. It is better than buckwheat, because it is as thick and rich and a great deal whiter.” Mrs. M. D. Herzet, Kinsley, Edwards county; nine stands. “Our feed for bees here is mostly alfalfa, some wild flowers; no fruit-tree blossoms. The bees find considerable feed on the box-elder trees early in spring. Have a fifty-acre field of alfalfa juining the apiary, with plenty of water within a few rods. The bees have had continuous feed upon alfalfa blossoms from the 10th of May until frost, which this year did not hurt the blossoms until the 16th of October. Do not think white clover or buckwheat could be any improvement on Fie. 5. (Original.) Honey-bee on alfalfa. From photograph. alfalfa for honey, either in quality or quantity. Our best swarm this year produced forty pounds by July 3, and thirty-two more at the close of the season, making seventy-two pounds for the swarm. Five of the other swarms did nearly as well. Our honey this season was pronounced by all who tried it to be as fine, if not the finest, flavored they had ever eaten. I do not know yet as there is: much profit in keeping bees, but it is light and pleasant work. My sixteen-year- OBSERVATIONS BY APIARISTS. 91 old daughter attends the bees almost entirely and enjoys it. She says it is easier than raising poultry, and has more money in it. I know it will pay any farmer’s wife to keep a few stands of bees in order to have plenty of honey to use in the family.” J.C. Batcu, Bronson, Bourbon county; fifteen stands. “I prefer Italians; have found Cyprians cross and almost unmanageable. We have no honey-producing flora in this part of the state. Bees in Kan- sas are not a source of pecuniary income every year; but, taking one year with another, will average with other agricultural pursuits. I have been keeping bees in a small way since 1875, and have never been out of honey for our table but twice, for a few months each time, and I have sold hundreds of dollars’ worth of honey.” D. B. Jones, Mound Ridge, McPherson county. “I have four stands of bees. My experience is that alfalfa is a good honey plant upon lands watered by rain. But I do not think there is any better plant to make fine-flavored honey than white clover. Yet our bees have been doing much better since there has been plenty of alfalfa raised here. I do not think bees have half the care they deserve.” H. H. McGuarn, Mayview, Jewell county; forty stands. “I prefer Italian, but like Carniolans almost as well. I think alfalfa and sweet clover are the best honey-producing plants we have.” P. C. Gress, M. D., Atchison, Atchison county; fifty stands. “I have none but Italians, as I find them far superior in every way to other races. They are better workers, more gentle, less excitable, winter better, remain kindly upon the comb, while other races under like handling run and fall down and scatter, and are more annoying to the apiarist. One of my observations is of special inter- est to fruit-growers. In my estimation the honey-bee is one of the greatest necessities for the proper cross-fertilization of blooms. I have protected limbs of trees by screen cloth during blossoming sea- son, and kept the insects from them, only to find a limb without fruit, where others, without protection, were well fruited. I have received good interest on every dollar invested, and mean to continue the in- dustry, despite the loss and backset which I received last winter, 153 colonies stored for the winter being burned.” Soton STEERE, Asherville, Mitchell county; thirty-five colonies; Italians. ‘“I think that alfalfa is about the only plant here from which bees secure a surplus of honey, and that, of course, with natural moisture. Sweet clover I consider the very best honey plant we have, but there is not enough of it. It is not inclined to spread very fast. I have scattered quite a little of the seed at different times, but re- sults have not been very encouraging. We seem to get the greatest benefit from alfalfa, when it is left for the seed crop. I do not make 92 OBSERVATIONS BY APIARISTS. any special effort to care for the bees, to make money from them. I have sold many colonies in the years gone by. We sell some honey, and have all we want for ourselves. There is no good reason why every family should not have a full supply of honey all the year round, All that is necessary is to get the start in bees; furnish them a home, and protect them. They seem to be willing to work for any one, and board themselves.” RosertT Dovuewas, Long Island, Phillips county. ‘I had six hives, grade Italians, last spring, and by natural swarming I saved thirteen, making nineteen swarms at present. From fourteen hives I have taken 800 pounds of section honey, which is worth at home ten cents per pound. My year’s profit might be reckoned thus: DOr. Cis 6 hives bees, at $5............. $30 00 | 800 pounds of honey, at 10c... $80 00 Suppilesons acta wae es hee 25 00 | 19 hives bees, at $4............ 76 00 a DOD ass esos me ee eae eae 20 00 To balance (profit)............ 81 00 Potala sae ern Sates See $156 00 Potalys ow. g s.r eee $156 00 Ep. HorrMetstEer, Norton, Norton county. ‘“‘My experience shows alfalfa to be a very good honey plant. Bee-keeping is quite profitable if a person tries to make it so. It is some expense to begin, but I have always made expenses and good interest on my money.” R. W. Smitru, Delphos, Ottawa county; twenty-two stands; I[tal- ians. ‘My bees gather their honey from alfalfa watered by rain, and have made forty pounds of honey to the stand. Alfalfa honey is su- perior to white clover or buckwheat honey. I lost several valuable swarms by wintering them in the cellar. Chaff hives are preferable. I use foundation comb for starting. Bees sometimes put comb in crosswise, and the starter prevents this. There is money, good health and honey in the bee business.” W. D. Jones, Neodesha, Wilson county. “I think alfalfa is equal to white clover or buckwheat.” M. A. Burts, Hiawatha, Brown county. ‘“My honey has been mostly secured from white and alsike clover, basswood, buckwheat, and wild flowers. I consider alsike superior to white clover as a honey plant. My experience leads me to the conclusion that bees, with proper care and attention, will pay in this country. It is important to protect bees well in winter and have them strong in spring.” Jacop Swoyer, Winchester, Jefferson county; twenty stands; Italian. “ Buckwheat, white clover and smartweed are the principal honey plants here. I have had no experience with alfalfa. Like white clover best when the season is not too wet. I do not allow my colonies to swarm much, about twice in three years. I prefer one strong colony to a dozen weak ones. In 1896 I had hives which made OBSERVATIONS BY APIARISTS. 93 110 pounds of honey and plenty for their own use. I would not do without my bees, as I think honey and pancakes good enough for anybody.” D. J. Fraser, Peabody, Marion county ; fifty-four stands. ‘I use Italians crossed with the German bee. Alfalfa is best in a medium wet season. Bees seem to care nothing for it if season is very dry or very wet. Like all other stock in Kansas, bees require some intelli- gent care. If properly mismanaged they seem to swarm world with- out end.” Fic. 6. (Original.) Honey-bee on white clover. From photograph. F. H. MIuuer, Great Bend, Barton county. “I had thirteen stands in the spring, and have twenty-eight now. Have taken 1200 pounds of comb honey, all alfalfa. It is a good honey plant. There is but one clover that beats alfalfa, and that is alsike. I have a small patch which blooms eight or ten days earlier than alfalfa, and the bees are thick in it from morning till evening. Alfalfa is a good and delicious honey plant, and produces a fine-flavored honey, better than Colorado honey as there is no other flavor mixed with it. I sell my honey at fifteen cents per pound, while Colorado honey sells for ten and twelve cents. A person cannot expect something for nothing. Take care of your bees and they will pay as good profit as anything I know of. I am in the hardware and agricultural implement business, and have little time for bees myself. Mrs. Miller, however, thinks more of the bees than anything else, and enjoys taking care of them. I think if more wives and daughters would take it up it would be better for them. They would find it a very good and enjoyable business. Ifa bee stings sometimes, it is good for rheumatism, and there seems to be 94 OBSERVATIONS BY APIARISTS. plenty of that in this country. Alfalfa is far ahead of buckwheat or white clover.” R. L. Snoperass, Gordon, Butler county. ‘Have sixty hives and manage, in addition, forty for others. I have kept hybrids and Ital- ians, and prefer the latter. Alfalfa has given me wonderful honey flows. My hives on scales this season and last ran as high as fifteen pounds per day. I have turned my whole attention to the bee busi- ness and have secured for this season’s labor 5000 pounds comb and extracted honey from sixty hives. I secured this season 200 pounds from one hive. Alfalfa, in my experience, is far ahead of buckwheat as a honey plant. I winter my bees both in the cellar and in the open air with success.” W. D. Fuuton, Garden City, Finney county; eighty-five stands. “Alfalfa as a honey plant is second to none for richness of nectar and flavor. It isa very common thing for a single colony to gather 200 pounds of comb honey ina season and sometimes more. I would pre- fer a dry climate for bees, provided there is plenty cf water at com- mand, but usually there is sufficient rainfall to make the business profitable here. I meet with very few difficulties in the care of my bees.” Dr. T. J. Conry, Florence, Marion county; 300 stands. “Alfalfa is our best and surest honey producer, although sweet clover would probably be better if raised in equal quantities.” C. A. D. BENNETT, Garden City, Finney county. ‘Ihave 220 stands of bees. Alfalfa is a splendid honey plant. There is no better in Kansas. We get three distinct flows from it each year. Each bloom gives a flow. The flow on irrigated land is more even than on land watered by rain, but is not so heavy. Alfalfa is superior in every way to white clover, sweet clover, or buckwheat. I do not attempt to build up weak colonics. I believe in the survival of the fittest. I give my bees the best care and attention. I leave plenty of honey in the fall for their winter’s supply. Then if they dwindle I let them go, and pay attention to my other colonies. In this way I rear colo- nies disposd to be strong.” Joun Weir, Carbondale, Osage county. “I have at present fifty- five colonies. I prefer the pure Italian. They are good honey gatherers, hardy, gentle, and beautiful. I have had no experience with alfalfa as a honey plant. There is not much grown in my neighborhood. We have both the yellow and white sweet clover here on waste land. The yellow is of most value, as it blooms six weeks earlier than the white, and just at a time when the bees are much in need of something todo. When the honey flow begins, I place the comb- honey colonies in one brood chamber, and keep them well-shaded, put OBSERVATIONS BY APIARISTS. 95 on forty-eight sections, and prop the hive off the bottom board to give better ventilation. In this way I prevent swarming to a very large degree. I thus secure strong colonies, and in this lies the secret of profits in honey. I have one ton of surplus honey at my elbow taken from forty colonies, spring count—1500 pounds ex- tracted, and 500 pounds comb, or an average of 50 pounds per colony.” Laurens Hawn, Leavenworth, Leavenworth county. “I have had no experience with alfalfa. White clover is the main dependence for a flow of nectar in this vicinity, and as it is very uncertain, the bee business is consequently not very profitable. There seems to be a prevalent though erroneous idea among horticulturists and people in general who raise a little fruit that bees are destructive to the ripe fruit. On the contrary, they are very beneficial to fruit-growers, and this fact should be promulgated among the people. I have had con- siderable experience in this matter for several years, and know posi- tively that without bees our fruit crops would be more often a failure. My apiary is in an orchard between two small vineyards; vines within twenty feet of the hives. I have suspended a bunch of ripe grapes in a hive during a time when the bees were working on decayed grapes in the vineyard, and not a grape was punctured. Of course, when grapes or other fruits are punctured or have rotten spots in them, and there is no nectar in the blooms, they will work on such fruit; hence arise the erroneous ideas concerning their destructive qualities. I have observed that if there is so much rain during the blooming period that bees cannot visit the fruit blossoms, there is always a failure of fruit crops. Many people spray their fruit-trees when they are in bloom, while they should never be sprayed until the bloom is falling. If people spray trees in full blossom they not only kill bees, but run the risk of poisoning the people who eat the honey made at that time. Another point is that in reference to the wholesomeness of honey as food: it can be used in all diseases when sugar and other sweets are prohibited, and if bought of reliable parties is free from all adulterations, which can hardly be said of sugar and other sweets. Bee-keeping in this locality, as a single occu- pation, is not profitable.” J. F. Crocker, Garden City. Finney county. ‘Alfalfa is a splendid honey plant if it has plenty of moisure, either from rain or irrigation. During protracted dry weather it does not secrete nectar, and during excessive wet weather the bloom sloughs off, and we get no honey. If we were situated so we could irrigate the alfalfa fields immediately after each crop of hay was cut, and the fields were not all mowed at the same time, and none mowed until the seed-pods begin to form, we would have a continuous honey flow from June 15 to October 1.” 96 OBSERVAVIONS BY APIARISTS. Fic. 7. (Original.) Honey-bee on sweet clover. From photograph. SAMUEL TEAFORD, Norton, Norton county. ‘Alfalfa clover is the best honey plant for this part of the state. It grows best on bottom lands, and produces most honey in seasons of moderate rainfall. The worst difficulty met with is the man who keeps bees and puts his honey on the market in a dirty, filthy condition, half capped, and no attention paid to grading; who, if he cannot find ready sale, will sell for anything offered him, to the great detriment of the man who is careful to plice on the market only a choice article, nicely cleaned and graded. To the man who wants to keep bees, I would say, do not buy but one or two stands of bees to start with; get pure Italians, in standard hives, with combs built on full sheets of foundation and wired in frame; would use full sheets of foundation, if had to pay one dollar a pound for them. Then get a good text-book on bee-keeping, and pay no attention to the man who tells you that bees do not pay, OBSERVATIONS BY APIARISTS. 97 and that the drones build the combs, and that the wax is gathered from sunflowers. Every farmer who is located within two or three miles of alfalfa or clover fields should get a few stands of bees, and have nice honey for his family, instead of buying the filthy glucose jellies and other stuffs that are for sale in every store.” JoHN W. LaBar, Erie, Neosho county. “TI have not been engaged in bee-culture except as a side issue and to furnish honey for our own table. We now have twenty-five colonies. We have taken 500 or 600 pounds of honey this fall, and the bottoms of hives seem to be full. There seems to be a good deal of white clover in this locality, and it is really the first honey flow we get that we can take off for table use, and is the finest flavor of any we get. This is a business that to be successful must be looked after in every little particular, and I believe it can be made to pay almost any year.” D. P. Nor, Burlington, Coffey county. ‘The difficulty here is the constant winds. Too many bees are lost as they come home laden with honey, and swarms are thus diminished in workers. Alfalfa is a good honey plant. Alsike clover is one of the best. I have visited several bee-keepers and asked them why they do not sow all kinds of clover and plant linden, locust and fruit-trees, raspberries and black- berries. The excuse was, bees do not pay. Neither would any kind of stock pay on a farm if it had to hunt its own living.” S. B. McGrew, Kossuth, Linn county. ‘The study of the honey- bee in its home and ways has always been a fascinating employment for my leisure hours. I have completely failed to secure an alfalfa crop after several trials. Linn county farmers, so far as I have learned, have had no success in growing it. The worst and only diffi- culty that I have met with is the lack of honey-producing plants in sufficient quantities for large numbers of colonies. I have done fairly well with my bees and would like to learn if there are any better plans than I have known.” OxueY Ouston, Cimarron, Gray county. “‘Alfalfa is considered a good honey plant in this section upon irrigated bottom river land.” J. P. Emery, Cimarron, Gray county. “I purchased two stands of Italian bees in spring of 1897. That fall I had increased them to ten strong colonies. I took off about 500 pounds of honey. Alfalfa is the main honey plant in this section wpon irrigated bottom lands.” E. Davison, Garden City, Finney county. ‘Alfalfa is the only honey-producing plant that we have in Finney county, with the ex- ception of fruit bloom in spring.” P. H. Bouuincer, Everest, Brown county: “I have forty stands of bees. Have found Italians to be best. Have never sowed anything but buckwheat.” all 98 OBSERVATIONS BY APIARISTS. D. F. Youne, Long Island, Phillips county: “TI find alfalfa a great honey producer where there is a medium amount of moisture in the ground. I believe alfalfa to be a great honey producer, equal to white clover or buckwheat. Winter in cellar, and think this requires less honey for the winter.” Wo. M. Borrom, Dexter, Cowley county: “Alfalfa is the best honey plant we have for all seasons; not so good as white clover or buck- wheat, but the latter does not do well here. In my opinion, as soon as this southern and western country becomes settled up, and groves and orchards are planted and ponds built, rainfall will increase, and _ as the country grows older the raising of tame grasses will become successful, and with it profitable bee-culture.” Jos—EpH HurrMan, Garden City, Finney county: “Alfalfa under favorable conditions is considered a good honey plant. We have not had sufficient irrigation in this section to test its worth as a honey- producing plant. Alfalfa in this country is a better honey-producing plant than white clover, sweet clover or buckwheat is in the eastern states. Quality of alfalfa honey is the best.” A. L. Dantets, Allendale, Lyon county. ‘‘As to profit derived from bees, have not had enough to pay expenses and labor, but the pleasure of having pure honey of my own has kept me at work.” J. W. Surron, Glasco, Cloud county. “I took 1400 pounds of honey from twenty-six stands. We have a large amount of alfalfa here. I do not think bees make much honey from alfalfa, as I have failed to find many at work on it. I find that my bees make more honey when the corn is in tassel, but the alfalfa is in bloom at the same time. I have seen more bees at work on corn than anything else. Have not given bees special attention, but am beginning to think it will pay to give them the care they require.” Capt. J. H. Wine, Syracuse, Hamilton county; 150 colonies. “Ital- ians and hybrids. I have tried several stands of pure Cyprians and find their disposition quite objectionable. I fiad, however, that bees developed from these Cyprians crossed with Italians or Carniolans, give, all things considered, one of the most satisfactory strains of bees. Alfalfa furnishes our main honey flow. The quality of the honey is second to none.” SOME HIVE YIELDS. Without any direct attempt at procuring the yields of the diftfer- ent apiaries within the state, or securing the highest yields, the re- ports here shown have come to the department giving amounts of honey produced. It is believed that these reports will be read with interest and will give a representative knowledge of what may be ex- pected from bees. SOME BEE STATISTICS. 99 It is regretted that the element of care and attention could not be expressed in figures alongside of the reports shown, for the old adage, “ Keep your shop and your shop will keep you,” is as true in apiculture as in any other line of business. | /NAME. ADDRESS. Year. Reais Yield. daWe sutton’.....5.. Glasco........ 1897 26 1400 pounds. DO. WOUNDS se a= oi Long Island..| 1898 Paine ( 2old | 146 soo eR Ea 7 re SESW sien ias tae 1898 1 1 gallon. (1896 5 50 pounds comb per stand. E. Davison.......... Garden City.. 1 SOT seacass xe 100 pounds comb per stand. | 1898 saciteoies 40 pounds comb per stand. ees MIMOLY s.4.s oes Cimarron.....| 1898 24 500 pounds. H. H. Morten....... Hong Island:.|'.... <6. ll 500 pounds (alfalfa honey). Robert Ferguson....| Galesburg....]........|......... $3 to $7 per hive of twenty sections, John W. LaBar..... BIER) 5 :5.5 gis’siniem | ss ss eas 25 500 to 600 pounds. Jahn Weil...) .) Cloudy, and’ strong: Wind 2. se cverie oe. ccaiecne rs Sale #10 14": | (Clear and: Warm occ 2 ante nc ce Stocco oise eke Re ** 15..| Clear, and strong wind after noon ............ 4162 .)| Raincand wit part Of days. .<+ saseecmus see OS el Cloananahot vcs oe oec cs tacek oe acinieaaeiceioe cs 184)| (Clearand Mobs ese. cases oe emiean one attecniee o19 = I Clegriand not (jer occstasscthe cakaeaemermee aan ©Op IO a VOGAL MNO NOU execs coerce eos cnet neetaaise 04) 1 Cleanmand neti. suoncWosemnae ce ce oe sinters ciemeisicte ‘« 92..| Clear and hot, cloudy after noon.............. $$-~937) | @leamaAnaktoby (ei meee aes acie > rletelnoe tiers ate cretera Sf oe Clear and hot, wind, and cloudy after noon.. Le EDEN NE|| coat Se ciate mfeieos etoren ere crane oe et oie nna ie wie hole wlchatnie asta aniots Ge 96s 4 A@lenr and Hots Gf) caeeensas ccs mceeoadesee aeunvas ‘* 97..| Clear and hot, cloudy and cooler after noon.. ‘* 28..| Clear and BUSIGE Nie i Lo ee eee ee ae S62 199 4@loariand WALI <-aonicairkace enc ae meketeine% ce:cic.cc.se. See ees sean. 96 54 170% EMP GET: ATIC, WATTN” ¢.ctc.c0 siawidscias oaciecclwie at's 00.00 101 36 173 Sealers |G LOudy-ANG GOO). (55 Sinyeiaoo Sia eis aya enema asta hee ore ee ee Nig ao Fm A OGUISE ebiax Sos SAGES ORE ¢ ee Sb ere Paw e BE RASS LOO UR e anid see oleae gers Fea eee lea EEE eae Cee 79 ‘* producing plants of Kansas, principal.........- ee eee eC oe ede cocdenc eres, uk: HO PPCI-GOZET, UEC OLsaiise ase bac ek vicaa he mas issicc awile ae eel ceine secant ne ae Wael oa Gales ee miaee eatene eee 12, 60 ae “¢" less efficient onscloudy Gays. .4\s.~.fccac=cces «ein waaclena species biette viecle nee talons meters i ‘* ability to capture full-fledged ‘ditferential locusts.................0-.2seeeeeee: 29 nf ‘* article on construction, uses, success attending experiments.............. 5d- 60 HMongh, Garry dei Ni woes ice escaves ooh cwicess cache setae dels wale aden tewem eee sekiia aot eit eel ks CeO tame onset plates. 2. ccoease ce hicsncuited Sona Sa eiclel ow eaiana ears one eager nee eee 126, 128 Hiaberi EP rancois 3.5 sosc oes esis sew abs is oo oo ee hoe nuckiele wiealoe whe Rote eae etaiae relate Ries care Ee ee Tee ee 74 Ht SHES He Eis io os ase eis etree n.a neice ne Boa GE Selly Sok CEE 5 NE OEE IT ERC eC aE OEE REE RC RaC en 83. 15 Gy heat oj spush vel Creel « Deen re eee Hr aan ee toy ea ans a eRe OU lien wo eierrmci mace co coet 1 14 PR CTER Whey Dice-se Par et a i 15, 49 EIVUINGVS, EET TTR ES oe Gc ees a ng ne 81, 8&5 150 INDEX. N. National Bee-keepers” Associations . 2%. sect cise sees sie wces|calclecieeeien stil eae same ees woRhees iie ok INGOSIIO' COUMUY cise icios.c cei ccciriniete s sins saiaierereainie clusters ulbinre Siaerasioveeia tee op dhe So laite Walo oa oore sieeiateest tissienter 97 Nervous system: of Melanoplus)differentialis.: :. 2.22. comes ne ce scemnen Se cece cesemicceietdaiistees 47 INGWE ls Sin Cs cee aide aera cies Sisters eto sisie Gal fo. a are ara ccjaialelmaree wha ie) win Diese iat cisteleetiat nie inte muctele saben aterays esi ot eeleters 18 INOG7 ODS Berea ese lacsae iene Scie nl cle oissle SR csheve Stele efsloos aia, bale ehelsiatapeleleretercio mats lata aleve eaaretenerslais erie tels serena 97 IN'OEGON COME Verret uce evita Gaid gsi wdc Sietellons fo Ritmo ale lvasane eter etanieve oct lavaselatevbler clove allerarcte taiatavar ayeselercteum mets 18, 92, 96 oO; BADD UM Be ea era sis > ciara eZ. diuie Glaee © micro wet aie iwivielstels ataial eleva a1a'80uie lnve sieccta Drailclelapate 61 vale iotetatate sere ofay claretoiesteteeenete 10 OMSetevatlons Dy ‘APLATIStS's cisieaats Fa uleieie seo, eoelalee dee ges eehioaine 12, SMP MMCCIE UNS AES MI ANLER TU ERS prey cae. 27 oi ciara cis. al oveiete ay elcte ators esac alesis sraleteeeaiae Leas oules od asred 4ihere 33, 34, Rea PORRSE RENT eh ECU Ee Maeretreerar aye recreates ticys cs Peisis Sa aiat as vie eistchicieieiel we oelcimicia aie niceiale sic.e/sio wins 6 ae Sacra poutesee 32, CSRIVO TYP ECD cS er GR een ie ee eae i ae ee ho ae ae ee ne ca eee ao ee Lae er LE PUREE eye R epee reiare aici otelonsioininie avcia sles olsialel che tay aiciel eisreta torso serach a eWeaite cose cuewet adh oeeaetes un 29, Section-holder super........... Sate caleete acke eee Tarai a asic ainiw ass\ccersiate eam bepasicls eben Aeka aes ae 113, SIM EKO (0G FE 2a oe eB EE OB GORO ISRO GCOS CoMCUO CEE aC CE MAC ICE nats Geet Aen aint ia aes ae ane wy AE Shipping crates for honey................ Sean Sieve Nein See TAaete ale eas URacS SSL neiok a Seow aise mache 114, eS ECMALLIN MEN rer tcrcinin ein crcveraie a eiele wea alolae Rican secre role wat cieleia, oe oreo clisie ai tg eutletaiclaCclacletale sub velo aaednewe teen RSSMERUTSTEDFAPIS MECCA V i ersols aies= cieiisia' em e's « eiayaeis areiia Slavs eiareio eiwieyi/ais clei exste'ee- ao cic’ays 6.3: Sle sie! alate oraheloautiela gine Sie oe Banton PUR RETO RCN KG Uotercei in ain atarayaio sVotnisint sveicisve sonic aiataetera(aclalel siaialaleic mieicle radian’ seiaa'e civ ote si ciels Mon Glorae ae cloacae MPPAVIN AWLP ATIS CTCOM. as ..c.ccsesesosnsinase se esiecuclac Se eine aECIe MATA OMEN TOL COS tains eton reais cfs teas semere oie ol oulaieeae Sena pie Cele oa we wee ne tee Reade es aera OLOMrearceNee Tae ashe eee ae os ho sici a are Parstieratole bciays eres ohardin.s Smae ond stem are eee oe cata Cae RAEI TEINS WN Toll Ortncted take a oran tara c\crat taieiac cle, v ssele syarerarmy Aras Sint aleveau Ate on Mie ena Salhi oka ciate reimiclasaladl SHloah Soa eed aise PR eMNA ROE CS CTMME A TAL levees cerca arciaveie stale aie u'ero' misisiolcVosinen ora eicia sides w'siaid ee lninload hema c oe on OUR ee ae. SSUAL EDN aM erate late vers ihiwcPatala Sie atcyeraveitiarnis, c/aisisvarulcral Aaianaiis Sia’ eioteree ove eee wala svobeialeretetaateseze tart duerclene Mievoa dalsaals Summary, relations existing between alfalfa and grasshoppers, and methods of prevention isl Oss LNGM LASSEN O DOES tekiaais is. oacie eaters daterave visclele samisiclesoBaieecieidete rise oaje’ weeeten acne 00s PSU LENET TION Olea OOS a ites arp nic isssinieler or csete ale a ceraeralolei there a wiaibieeiSnaPanIey oeitere Meee acd SEL ol Toc alos uke 108, us NG WaEGV EH CCC nets. treat costa everereieie ste Sad aaa Mec erate Pare erate esa Loon Tne ee BR WAN ISEIOR SMES CRWPELUV CU or clese al slarcras « s ciaictaialc orars ei oteistom ole la ch clout ee mindiet erahe Seas lone oteidn ede fete bekee pele POMEL AES ee el om Oxstets ehre Tein fais ire cfai vloic ave Sebeteraleies eis aatete Sele ulalaseteeinarate aleveb were hick lett ceresete cette Giecine SNM MB LD aN etna cei ett eens caliente clcta ake smncaceatte Ceatel ci einie rena eee cae aon coe VAMP we TCO iiamrctcccicinssieiaicinaeid rae cn alsaiete wislacnoe eae eas eins suis Go tae ate Sete eee een Jems REN EL EGON AE cicteel vere cm ieee ostvciale ocho nals bcieeia tale win nies Malatadle ooo cine k bee cinema el etind Uolae atl ocinetenea oes Pablo showing influence of bees on) alfalfa Seed s.~..2 :2.6<+ osc. ccee coee caedeanccnenceckunscnces ss sf ehomlcal analysis OL NONSY «kescraceen sees ioncde do enact wtisoa ec catiloa noe is oe alfalfa acreage, stands of bees, pounds of honey..............ceceeeceecee 86, {DEG} ECTS CONES ce SRE MER to or ar Rn Pa A nr SA A ee TO ae a 32, TPE one |, SRIar a) ae SRB BAe annie a ora e tn TASER ce eae ae a ee RR aT UN Technical description of Melanoplus differentialis. ...............cc0ceceseccccccec voceccce 63, BU tesco ENTE yee rete rec masta a eesteys mince alain See ete Ne Ts lorad Hiosalo aa Gis oie date eine w dloveis secs bm gaee Bae nese 34, ae PGEEEL cts mete Ste (cinta Be te laine Sicko eoe Solo a einclo's Wadia mweac one e nak cued tendo dence 36, “s RIBOA vee ntran surements siataletemiaraislateletsietbiayberiaefe-nee oatvce ne Sr Acssieieminaieteaietelcd aera cceeieane 36, be WARIM DOM lige tee Mire hetero ate afaicctad cio sive Pisiviclom asioeioniainats ooe ea Cae eaees Sai asia vars io ciaience ce 36, on LNG cecppeicie. bb a SSO ABO SACLE ICO TE a eT ee ate aE en See RE aR tae 36, as CDSOLG bars seine teig eicie vials cia oe cic dais doieew Seeee he ca Soe Thlsarsadteane ad boiad sebnomee enon 36, RE Hpe vote Al Den tne eet eroet te matech eine cial aie sae, cistorkcy wictals Gare w felcta tea actos oie ue be lve de be, aula av werd thr eoaeeis homas, (GC) 2.5.2 23 Retreat ets ater ote eles sla ievalaicl onerohe is Sicimia eis raesevclaiee a aide oh cca S la aietwete eo ea oman cia caer BES IACWTNE CEC EX ELENG Votes ote ame Terese c(t ee icicle painiol@ cieie Ecce akinaleybiwinfu sad pa adin Ld old recineD eh Wekcd wees Ml LCST 9 Ip Seapine re Metaoteneeyabate otalateaisiaycte et eei ata, fatal cteeisi oe otc < sis hsveiaie sieisi’etsweicvacion @areorctin ears a6 eb wee SLGWDOBENO MCs Hig Wace sce ccccncc ceeweu onesmcene Tao loa Reis csiee oles. LS EH/ ote De tans ecentaeies os 31, 33, Pires LETTE LOC TINEA CUI oe ceisel ers cian ce cb les oni astern sisi becasue siaseu tes eae daieiedcacest ccazacecad MRE UPREELUAIES eat eret tn ctate creas Naiveione oule Giveoe eis ale oa okie Seaie'svendle cede aoe Ge atoiclers aiaisieraiie ceelnarbia’eh ¢ hu Raateuen 92 30 60 103 14 61 91 xi 36 17 105 80 80 81 119 112 98 61 120 108 119 79 17 92 69 152 INDEX. SOT BUDGE EHO so akvt at chai -stue ds neeieis Canin n'ai brsteie ciate me aie ores Gh a SSeS Tate Scie een Sean Dee en eee 112 Two-striped Locust—see Melanoplus bivitattis. MOMMSTAM DROS Scars ceuias ciaisiel seals caree bilge tes Sinelot atin onc st rae ate, ole tle IAP Tr ter ares cto vin ote ah teaten oier ert aerate 69 U. | Minions PaAcitiesmallwayiss swans tes ve.wscciectuwcca-cend oues kee se ee ee rebes Mince aeERe Otome aie 1, 16, 49 United States Department of Agricul tures. in sadectcos macwucciicinees nose ce cece caseuceeeenectere 71 as Blot: NOHO. siiersactorrsietebretehetetelelaresve lve aeiGe caiewnuraieg satersiodiersnamiee oe iis Soe eae 30 We WiITGOLOLIVACCUS cinvre ssc nciseiefe soins ciee's whant cle dewaievlnes gers ©cwciniemeoleelat See alee sevanie tect ecm aeentes 30 Vivarium, Observatlons'on Locusts Wis) 2.55/50 «0-5 sow ssise a cioisreacteceisurecinies sactee a edae cianiee cislaateetee meme W. Wier Mears Wine na ioc sro stinsaycjepartis 2's oie oral je ace 'ararey ciminia fate ayers va ote ecle les vias eisro sian ateiaseralel Sinton ete oats 16, 49 WAT COM COWIE iis oie orntstcoreiclowte a < 4 a aA =)