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ALLAN HANCOCK PACIFIC EXPEDITIONS

VOLUME 19 NUMBER 1

QUANTITATIVE SURVEY OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

PART I

PRELIMINARY RESULTS
(Charts 1-2, Plates 1-7)

f

BY

OLGA HARTMAN

~LIBRARL A

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS
LOS ANGELES, CALIFORNIA
1955

OUANTITATIVE SURVEY OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

PART I

PRELIMINARY RESULTS
(Charts 1-2, Plates 1-7)

By Otca HarTMAN

THE UNIVERSITY OF SOUTHERN CALIFORNIA PUBLICATIONS
ALLAN Hancock PAcIFIC EXPEDITIONS

VOLUME 19, NUMBER 1

IssUED JANUARY 31, 1955

Price $4.50

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS

Los ANGELES, CALIFORNIA

TABLE OF CONTENTS

INTRODUCTION
ACKNOWLEDGEMENTS
DESCRIPTION OF THE AREA
CHART OF THE AREA

CRUISES OF THE VELERO IV CONCERNED WITH
QUANTITATIVE SURVEY

LIST OF STATIONS

ANALYSES OF SAMPLES BY DEPTHS
ANALYSES OF BIOLOGICAL MATERIALS
SUMMARY OF RESULTS

APPENDICES

A. Improved Techniques for Ocean Bottom Sampling,
by Floyd E. Durham

B. A List of the Mollusca identified from samples of the Benthos
of San Pedro Basin, California, by Norman T. Mattox

C. A List of Phoxocephalid Amphipoda identified from samples of
the Benthos of San Pedro Basin, California, by J. Laurens Barnard

D. A List of Brachiopoda identified from samples of the Benthos
of San Pedro Basin, California, by Olga Hartman

E. A List of Enteropneusta identified from samples of the
Benthos of San Pedro Basin, California, by Keith Woodwick

F, A List of Polychaetous Annelids identified from samples of
the Benthos of San Pedro Basin, California, by Olga Hartman

PLATES

154

159

164

166

168

187

QUANTITATIVE SURVEY OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

Part I. Preliminary Results

INTRODUCTION

This report outlines the preliminary results of a quantitative study
of the marine animal populations in a limited area of southern California.
The purpose of the project has been to conduct an intensive survey of the
benthic invertebrate metazoan animals of San Pedro Basin, and to at-
tempt a reconstruction of the natural animal associations. Many of the
physical features, including temperature and salinity of the Basin, are
known or are being investigated by the geologists and oceanographers
of southern California. In time, therefore, the biological and physical
data may be correlated and many features of distribution and abundance
explained.

Throughout the San Pedro Basin, measured samples have been taken
at chosen intervals for more than two years (1952 to 1954), using a
bottom sampling device that grabs a sizeable part of the sea bottom
from any depth. The device is dropped as a plummet, presumably without
obliquity ; its measured descent is used to indicate depth to the bottom.
The sample is hauled up and immediately deposited in a large tub that
completely accommodates the contents of the grab, in as undisturbed a
condition as possible. Two half-pint samples are removed and set aside
for future physical studies. From the tub the material is transferred to
a set of screens of selected sizes. Care is taken that the innumerable soft-
bodied animals remain as intact as possible. The screens, with mesh from
coarse to fine, measuring to 24 meshes to the inch, sort out the animals
and other particles; they are washed with seawater and preserved with
a reagent (formalized seawater) to harden protoplasmic tissue. Finally
the labeled samples are taken to the laboratory, where all subsequent
work is done. Tyler Standard screens, with meshes to a fine of 32 to the
inch, are used for further washing of the samples. They are sorted and
analyzed, then transferred to 70% alcohol.

[1]

Z ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

During the first part of the operation, a series of four or five screens
was used on the boat. These screens measured 18 by 24 inches and were
made of stainless steel fitted with a brass mesh that varied from a coarse
one of 12 meshes, through 16, 20, and 24 meshes to the inch. When much
rubble was present in a sample, a coarser screen was placed over the
others to expedite the sorting of the finer materials. The mud was washed
through the screens with the aid of seawater hoses. The process was slow
and physically tiring. As harsh sprays were found to damage soft-bodied
animals, the use of nozzles on hoses was impracticable. Lighter, finger-
controlled sprays were laborious but resulted in better preservation of the
animals.

The methods of washing and clearing of fine debris were considerably
improved when larger screens (figs. 14-16) with greater range in mesh-
size were introduced. These larger screens are now mounted on a shaker
device (see fig. 16 and Description of Equipment, below) which permits
the flow of materials through them without damaging the specimens.
The seawater comes through a system of shower-sprays, and the amount
and force of the sprays can be controlled through valves. Many of the
smaller animals that had become enmeshed around the wires of the
screens, had formerly to be washed or picked off separately; now they
are largely disentangled with the aid of an air hose that causes them to
drop to the retaining sack (fig. 23). The removal of the washed and
cleared animals from the screens was at first performed by raising the
entire screen and washing the contents into a large pan or tub. This
method has been improved so that the contents are removed without
much lifting.

The Velero IV has provided an excellent base from which a grab,
called by its manufacturer the Hayward-orange-peel bucket (known
below as the orange-peel grab), of about two cubic feet capacity (see
figs. 5-8 and Description of Equipment, below), can be suspended. A
second larger one, the Campbell grab (figs. 17-20), named for its de-
signer, Alex Campbell, was introduced in April, 1954, and found more
effective in some kinds of bottoms. For each sample, the living metazoan
animals are sorted out. Many of these have now been identified, counted,
and charted. Since ultimate results can be based only on final counts
of heads or some other unit parts of living specimens, all estimates should
be regarded as minimal. There are, necessarily, losses of unmeasurable
amounts, resulting from the various stages through which each sample
must be processed.

No. 1 HARTMAN: QUANTITATIVE SURVEY 8)

A map of the area (Chart 1) has been marked off by intersecting
lines two minutes of latitude and longitude apart, choosing all even
minutes of each, or about every two miles. It is at once obvious that a
small sample covering approximately two square feet of surface and
with a volume of about two or three cubic feet, is very meager to repre-
sent a seabottom that measures nearly four square miles in surface area.
Furthermore, it was soon found that even though a uniform sampling
device was employed, few of the samples measured as much as two cubic
feet, especially when taken from the shallower waters of the continental
shelf, where measured samples were as small as a tenth of a cubic foot.
Others, from soft oozy muds in great depths, measured well over three
cubic feet. It was found also that the degree of penetration of the grab
varied in different kinds of bottoms; these varied from the finest muds
to compact sandy or shelly gravels to clay. In addition, the incomplete
closure of the jaws of the grab may have caused the loss of contents on
the way to the surface. Some of these difficulties have been met through
improvements in techniques as the sampling continued.

Although the studies are not complete, it seems worthwhile to describe
the methods that have been used, and to indicate preliminary results
which are remarkable for their qualitative and quantitative relations.

4 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

ACKNOWLEDGEMENTS

This program has been supported by Captain Allan Hancock, Direc-
tor of the Allan Hancock Foundation of the University of Southern
California. Captain Hancock has taken an active interest in the project
and accompanied most of the cruises. The field work was performed by
participating members who were largely members of the Foundation
or the University faculty, and zoology students. The technical problems
of constructing and adjusting suitable equipment were solved by the
Engineer, Alex Campbell, and his staff on the Velero IV. The ship was
maneuvered in position by Fred Ziesenhenne so that chosen points were
accurately found; he has also had charge of recording all terrestrial
positions for the charts. During the second half of the survey, Dr. Floyd
E. Durham has had charge of field operations. The photographs of the
submarine scapes were made with the benthograph operated from the
Velero IV. Dr. K. O. Emery and staff of the Geology Department of
the University provided data on temperature and some physical properties
of the San Pedro Basin.

Identifications of animals were made by various authorities. Dr. J.
Laurens Barnard is responsible for the phoxocephalid amphipods; Keith
Woodwick determined the enteropneusts and most of the polydoriid
polychaetes; Dr. Norman T. Mattox was consulted for the mollusks.
Other determinations, except those indicated below, were by the writer.

No. 1 HARTMAN: QUANTITATIVE SURVEY 5

DESCRIPTION OF THE AREA

The area selected for a quantitative investigation is the San Pedro
Basin (Chart 1), the shallowest of a series of about a dozen depressions
along the coast of southern California (see Emery and Shepard, 1945,
“Lithology of the sea floor off southern California.” Bull. Geol. Soc.
America, vol. 56, pp. 431-477, chart). It lies between the mainland of
southern California and Catalina Island, and is continuous northwest-
ward through a narrow channel with the Santa Monica Basin. It is
bounded by a submarine valley, the Redondo Canyon, to the north, and
by the city of South Laguna Beach to the south. Its geographic boun-
daries extend from 33° 16’ to 33° 50’ north latitude, and 117° 46’
to 118° 36’ west longitude. The area comprises about 520 square miles
of sea bottom. Depths range from 4 to 495 fathoms, with the deepest
measured depth about half way between Isthmus, Catalina Island, and
Long Point, on the mainland. There are two poorly marked channels
running approximately parallel to the mainland. There is a northwestern
threshold or sill depth of about 489 fathoms, only slightly above the
deepest part of the basin, and a southeastern one of about 400 fathoms,
east of Avalon, Catalina Island. The oceanward basins, beyond San
Pedro Basin, gradually attain far greater depths, to more than a thousand
fathoms.

There are four submarine mounts within the Basin, one on each side
of the two threshold depths, at the northwestern and southern ends. The
largest one, the Lasuen Seamount (Chart 2), is the most easterly. It
rises abruptly from 385 to 58 fathoms within a mile. There are steep
descents along the rims of Redondo Canyon and along the leeward shores
of Catalina Island. There is a broad shallow submarine shelf, about six
miles wide at most, south of San Pedro Bay, that extends coastwise from
Long Point to Newport Beach. This has an extremely rich and highly
diversified fauna. There is a large sewer outlet (conduit for domestic
pollution) off White’s Point, San Pedro, emptying into the sea nearly
a mile off shore. There are fluvial and terrigenous materials brought in
by rivers, especially the Santa Ana River, northwest of Newport Bay,
and the San Gabriel River, near Long Beach; these carry considerable
loads into the sea during some months of the year. There is a long break-
water at the ocean side of Outer Harbor. There are cable crossings and
possible industrial pollutions from manufacturing plants, all of which
may have some effect on the productivity of benthic populations.

voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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Chart 1.

No. | HARTMAN: QUANTITATIVE SURVEY 7

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fathom contour lines, the northwest and southeast sills. The areas of impov-
erished fauna at the western end are indicated, and the glass sponge bottoms
are shown.

8 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

The latitudinal and longitudinal lines used to mark off this area
result in 267 intersecting points (see Chart 1), from which the samples
originate. The departures from the exact even minutes are due to opera-
tional or economic factors, beyond the control of the experiment. The
precise locations are expressed in the Station List, below.

The numbers, 2107-52 to 2859-54, are a continuation of the series
published earlier in the Allan Hancock Pacific Expeditions (vol. I, no. 3,
1943). Only stations in the San Pedro Basin for which quantitative grabs
have been taken, are here included.

‘Temperatures were taken during the first year of the survey, by
immersing a centigrade thermometer in the mud sample just after it had
reached the deck of the ship. It was found that the readings were unreli-
able, since those from great depths were too high, due perhaps to the
effects of warming on ascent through the upper warmer layers of water.
The samples from shallow depths were frequently very small, making
the process difficult. On the whole it can be stated that the temperatures
of the shallower shelf vary diurnally and seasonally. The deeper parts
of the basin may be nearly homothermal, depending on depth.

Based on readings taken by immersing a thermometer in the mud
sample on the deck of the ship, temperatures from shallow depths were
about as follows:

Station 2124-52, no. 61. June, in 29 fathoms—15.5°C.
Station 2152-52, no. 224d, Sept., in 19 fathoms—14.0°C.
Station 2176-52, no. 164a, October, in 28 fathoms—13.0°C.
Station 2217-52, no. 45b, February, in 12 fathoms—11.5°C.

Temperatures for greater depths, taken with a reversing thermometer
lowered to the water mass just above the surface of the mud, have been
made available by Dr. K. O. Emery, from transects taken from San Pedro
shelf to Catalina Island; these are as follows:

650 feet (108 fms)—8.5°C.
1000 feet (167 fms)—8.0°C.
1200 feet (200 fms) —7.5°C.
1600 feet (267 fms) —6.5°C.
1800 feet (300 fms)—6.0°C.
2100 feet (350 fms)—5.5°C.
2700 to 2850 feet (450 to 480 fms) —5.04°C to 5.07°C.

No. l HARTMAN: QUANTITATIVE SURVEY 9

CRUISES OF THE VELERO IV CONCERNED WITH
QUANTITATIVE SAMPLING OF THE BENTHOS
OF SAN PEDRO BASIN, CALIFORNIA

Thirty-seven cruises, from April, 1952, to June, 1954, have been
made by the Velero IV, concerned partly or wholly with taking quanti-
tative samples in San Pedro Basin and vicinity. These cruises, with dates
and station numbers are:

Cruise 85. iv-20-1952. Stations 2107-52 to 2110-52.
Cruise 88. vi-18/19-1952. Stations 2113-52 to 2122-52.
Cruise 89. vi-25/26-1952. Stations 2124-52 to 2130-52.
Cruise 94. viii-6/7-1952. Stations 2138-52 to 2146-52.
Cruise 97. ix-26/27-1952. Stations 2148-52 to 2153-52.
Cruise 99. x-18-1952. Station 2166-52.

Cruise 100. x-30/31-1952. Stations 2168-52 to 2177-52.
Cruise 104. xii-5-1952. Stations 2189-52 to 2193-52.
Cruise 106. i-17-1953. Station 2202-53.

Cruise 111. ii-27/28-1953. Stations 2217-53 to 2233-53.
Cruise 114. iv-19-1953. Station 2290-53.

Cruise 115. iv-24/25-1953. Stations 2291-53 to 2306-53.
Cruise 117. v-15/16-1953. Stations 2307-53 to 2314-53.
Cruise 118. v-19-1953. Stations 2315-53 to 2317-53.
Cruise 119. vi-24/25-1953. Stations 2321-53 to 2336-53.
Cruise 120. vii-1/2-1953. Stations 2337-53 to 2355-53.
Cruise 121. vii-8/9-1953. Stations 2356-53 to 2374-53.
Cruise 125. vii-28-1953. Stations 2386-53 to 2389-53.
Cruise 126. viii-17-1953. Station 2394-53.

Cruise 129. ix-16/17-1953. Stations 2403-53 to 2414-53.
Cruise 131. ix-29/30-1953. Stations 2417-53 to 2429-53.
Cruise 132. x-10/11-1953. Station 2430-53 to 2445-53.
Cruise 133. x-14/15-1953. Stations 2446-53 to 2454-53.
Cruise 135. x-28-1953. Stations 2470-53 to 2476-53.
Cruise 137. xi-27-1953. Stations 2496-53 to 2500-53.
Cruise 138. xii-10-1953. Stations 2501-53 to 2508-53.
Cruise 143. iii-3-1954. Stations 2606-54 to 2615-54.
Cruise 146. iv-7-1954. Stations 2618-54 to 2620-54.
Cruise 148. iv-20-1954. Stations 2625-54 to 2629-54.
Cruise 149. iv-24/25-1954. Stations 2630-54 to 2646-54.

10 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Cruise 152. v-8/9-1954. Stations 2722-54 to 2740-54.
Cruise 153. v-14/15-1954. Stations 2741-54 to 2757-54.
Cruise 156. v-22/23-1954. Stations 2788-54 to 2802-54.
Cruise 159. vi-17-1954. Stations 2835-54 to 2839-54.
Cruise 160. vi-22/23-1954. Stations 2840-54 to 2850-54.
Cruise 161. vi-25/26-1954. Stations 2851-54 to 2859-54.

The 303 stations, numbered 2107-52 to 2859-54, are described
immediately below, giving the corresponding serial number (1 to 267)
on the chart, date, approximate locality in San Pedro Basin, exact latitude
and longitude, depth in fathoms, volume of grab and kind of gear used,
and general remarks on kind of bottom. Some of the earlier stations
(2107-52 to 2291-53) give temperature of the bottom.

No. l HARTMAN: QUANTITATIVE SURVEY 11

LIST OF STATIONS MADE BY THE VELERO IV, for a
Survey of the Benthos of San Pedro Basin, California, giving corre-
sponding Serial (Sample) number, shown on Chart 1, date, approximate
locality in San Pedro Basin, position, depth in fathoms, volume of sample
and kind of gear used, temperature for some of the earlier stations, and
general remarks.

voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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ALLAN HANCOCK PACIFIC EXPEDITIONS

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JY] Jae M

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voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

24

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25

HARTMAN: QUANTITATIVE SURVEY

NO.

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voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

26

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voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

28

*S[BUIIUB 139q}O

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29

QUANTITATIVE SURVEY

HARTMAN

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NO.

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ALLAN HANCOCK PACIFIC EXPEDITIONS

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voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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HARTMAN : QUANTITATIVE SURVEY

NO.

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VoL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

34

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35

HARTMAN: QUANTITATIVE SURVEY

NO.

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ALLAN HANCOCK PACIFIC EXPEDITIONS

36

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voL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

38

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No. 1 HARTMAN: QUANTITATIVE SURVEY 39

ANALYSES OF SAMPLES BY DEPTHS

The following list groups the numbers of samples taken from various
depths, from shallow (0-25 fms) to deep (476-500 fms). The serial
numbers are the same as those used in the Station List (above) and the
Analyses of samples (below).

51 samples are from 0 to 25 fathoms:
14, 20, 27, 28, 29, 30, 31, 32, 33, 43, 44a, 44b, 45a, 45b, 46a, 46b, 47,
48, 49, 50, 61a, 61b, 62, 63a, 63b, 64, 65, 66a, 66b, 67, 68, 80b, 81a, 81b,
82, 83, 84, 85, 86, near 86, 87, 88, 102, 106, 107, 108, 110, 134, 186a,
186b, 224d.

31 samples are from 26 to 50 fathoms:
7b, 12b, 13, 42b, 80a, 99a, 99b, 100, 101, 103, 105, 123a, 124, 125, 129,
133, 159a, 159b, 164a,* 185, 207, 224a, 224b, 224c, 224e, 238a, 238b,
238c, 250a, 250b, 259.

11 samples are from 51 to 75 fathoms:
3, 4, 5, 7a, 104, 109, 123b, 184a, near 186, 235, 260.

6 samples are from 76 to 100 fathoms:
2,26, 60, 128, 239) 251.

8 samples are from 101 to 125 fathoms:
6a, 19, 79, 122, 130, 149, 208, 261.

8 samples are from 126 to 150 fathoms:
Ge, 42a, 127, 131, 160, 161, 162; 2252.

12 samples are from 151 to 175 fathoms:
6b, 11b, 11c, 126, 132, 148, 158, near 187 (2 lots), 209b, 267a, 267b.

11 samples are from 176 to 200 fathoms:
1, 6d, 41a, 41b, 121a, 174, 184b, 187, 209a, 225b, 236.

7 samples are from 201 to 225 fathoms:
96-147, 150; 151, 152, 1:73; 252.

12 samples are from 226 to 250 fathoms:
12a, 59a, 59b, 121b, 155, 156, 175, 188, 199, 220, 240, 262.

8 samples are from 251 to 275 fathoms:
10b, 135, 163, 178, 179, 180, 226, 237.

18 samples are from 276 to 300 fathoms:
9, 11a, 34, 40, 51, 52, 78, 146, 157, 181, 182, 198, 202, 210, 228, 229,
230b, 249.

*Compare with 164b, from 470 fathoms.

40 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

9 samples are from 301 to 325 fathoms:
10a, 18, 172, 183, 203, 204, 205, 222, 223.

8 samples are from 326 to 350 fathoms:
120, 136, 189, 196, 206, 215, 218, 241a.

9 samples are from 351 to 375 fathoms:
17, 23, 77, 97, 232, 233, 234, 241b, 241c.

14 samples are from 376 to 400 fathoms:
24, 25, 58b, 69, 89, 111, near 171, 195, 214, 230a, 242, 243, 245, 248.

13 samples are from 401 to 425 fathoms:
8, 16, 22, 76, 170, 194, 227a, 227b, 244, 253, 254, 255, 257.

22 samples are from 426 to 450 fathoms:
15,.near 15, 21535; 36,37, 38, 39.53; 57, 58a; 75, 96, near WIZat9:
137, 145, 193, 212, 213, 263, 264.

20 samples are from 451 to 475 fathoms:
54, 55, 56, 70, 71, 73, 74, 90, 95, 112, 115, 118, 141, 144, 164b, 168,
169a, 169b, 192, 211.

20 samples are from 476 to 500 fathoms:
72, 91, 92, 93, 94, 113, 114, 116, 117, 138, 139, 140, 142a, 142b, 143,
165, 166, 167, 190, 191.

5 samples from stations 2846-54 to 2850-54 originate from the deep
basin on the windward side of Catalina Island, in depths of 504 to 715
fathoms.

No. 1 HARTMAN: QUANTITATIVE SURVEY 41

ANALYSES OF SAMPLES, GIVING BIOLOGICAL
COMPONENTS

The serial numbers 1 to 267, shown on Chart 1, correspond to the
samples that have been taken in San Pedro Basin. The following pages
give a summarized analysis of many of the animals, with numbers of
individuals, for some of the samples. Specific determinations are to be
regarded as provisional; they are given to provide an index and approxi-
mation of the sample. Counts of individuals are minimal, especially for
tubicolous forms which have not been entirely examined. The identifica-
tions of species have been made by numerous individuals, indicated under
the proper category, unless specified in one of the Appendices (see below).
The analyses are intended to provide information on abundance and
diversity of species, and associations within a limited area.

With the aid of the specific names and sample numbers, it is possible
to plot the distribution in San Pedro Basin for any particular species,
by placing it on its corresponding number on Chart 1.

1. Station 2790-54. Off Palos Verdes Point, in 180 fms, blue-gray
mud. 2.33 cu. ft.
brissopsid and another urchin
echiuroid
burrowing anemone
crustaceans, including amphipods, isopod, cumaceans, ostracod
phoxocephalid amphipod, Harpinia sp. A—2
polychaetes, including:
A glaophamus sp.—2
A naitides sp.—3
Chone sp.—1
cirratulid—1
Drilonereis sp.—1
Goniada sp.—1
Harmothoé scriptoria—1
another harmothoid—1
Laonice sp.—1
Lumbrineris cruzensis—1
another Lumbrineris sp.—1
Maldane sp.—many
another maldanid—1
Nothria pallida—many, in thick mud-walled tubes

42

Z:

6a.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Onuphis sp.—1 or more
Pectinaria californiensis—many
Prionospio pinnata—several
another Prionospio sp.—2
terebellid—1
Station 2789-54. Off Palos Verdes Point, in 90 fms, sandy blue-
gray mud with sand, several larger stones and much shelly
rubble. 1.7 cu. ft.
echinoderms, including brissopsid and ophiuroids
pelecypod, Tellina sp.—7 or more
echiuroid—1, bright green
polychaetes, including:
Chloeia pinnata—several
Travisia sp.—2 larger
tubes of polyodontid
many smaller annelids, not yet analyzed
Station 2727-54. Off Palos Verdes Point, in 66 fms, green sticky
mud. 1.76 cu. ft.
many ophiuroids and annelids, not yet analyzed.
Station 2726-54. Off Palos Verdes Point, in 70 fms, green sticky
mud with sand. 2.77 cu. ft.
many ophiuroids; urchin; Dentalium rectius; Chloeia pinnata—
many ; diversified smaller annelids
Station 2725-54. Off Palos Verdes Point, in 58 fms, coarse green
shelly mud with rounded stones; many fine white spicules of sponge
resembling rock wool, much rocky shell rubble and fine to coarse
gravel. 1.13 cu. ft.
sea whip—1; echiuroid—1, deep green; mollusks, including scaph-
opods, Chaetoderma, purple tectibranch—1; flatworm—1
polychaetes, including:

A glaophamus sp. Nephtys ferruginea
Brada sp. onuphids
Haploscoloplos sp. Pectinaria sp.
Laonice sp. polyodontid tube
Lumbrineris sp. Sternas pis sp.
maldanids Terebellides sp.

Station 2191-52. Off Redondo Beach pier, in 125 fms, fine sandy

mud. 2.7 cu. ft.

mollusks, including pelecypods and scaphopods; polychaetes, in-
cluding Chloeia, spionids, maldanids, chaetopterids

No. 1 HARTMAN: QUANTITATIVE SURVEY 43

6b. Station 2148-52. Off Redondo Beach pier, in 161 fms, sandy mud.

2.3 Cuedte

nemertean—1 large

T halassema sp.—4 large, in life deep blood red, with white pro-
boscis; length to 140 mm, width about 20 mm, harboring a
pinnotherid crab

echinoid—1

polychaetes, including:

Chloeia pinnata—about 50 nereid—1
A ncistrosyllis sp.—1 or more Pherusa spp.—several
capitellids—many Prionospio spp.—several
Lepidasthenia sp.—1 Travisia sp.—1
Lumbrineris sp.—several many smaller annelids not
Nephtys sp.—several yet identified

6c. Station 2149-52, Off Redondo Beach pier, in 129 fms, sandy mud.
2s Cue tte

echinoid—1 large
crustaceans, including amphipods, ostracods, copepod
phoxocephalid amphipods—Heterophoxus sp. A—1
Phoxocephalus sp. A—1
Paraphoxus sp. A—1
nemertean—1 or more
mollusks, including:
scaphopod—Dentalium rectius—2
pelecypod—Y oldia scissurata—3
echiuroid, including Thalassema sp.—3 large
polychaetes, including:

A naitides sp.—1 maldanid — several, in thick

A ncistrosyllis sp.—1 mud-walled tube

Chloeia pinnata—more than60; Mesochaetopterus sp.—1, in
to 15 mm long, bright orange very long, strawlike tube
red in life Nothria sp.—2

cirratulid—1 Notomastus sp.—many

Cossura n. sp.—several Pectinaria californiensis—

Dorvillea sp.—2 several with many tubes

Glycera sp.—2 Pherusa spp.—several

Goniada sp.—2 polynoid—1

Haploscoloplos elongatus—6 Terebellides sp.—1

Lumbrineris index—6 large other annelids

another Lumbrineris sp.—several

44

ALLAN HANCOCK PACIFIC EXPEDITIONS

6d. Station 2190-52. Off Redondo Beach pier, in 186 fms, fine sandy

mud. 3.02 cu. ft.

echinoid—1

T halassema sp.—1
nemertean—several
holothuroid—several
Chaetoderma—several

polychaetes, inc

luding:

A glaophamus sp.—1 or more

Anaitides sp.—1

Bradasp.—1

?Capitella sp.—1 or more, and
other capitellids

Chloeia pinnata—many

Glycera sp.—1 large

Goniada sp.—1

Harmothoé scriptoria—2, with
internal parasites

Lumbrineris sp.—several
Nephtys sp.—several
Nothria sp.—1
Pectinaria californiensis—12
Pherusa sp.—several
Prionospio spp.—several
sabellid—1
Scalibregma—1|
other annelids not yet

sorted

7a. Station 2192-52. Off Redondo Beach pier, in 61 fms, fine mud and

sand. 1.51 cu. ft.
mollusks, including:
scaphopods—Cadulus fusiformis—1

Dentalium rectius—142

gastropods—A cteon punctocoelata—1

Bittium catalinensis—1
Fusinus arnoldi—2
Turbonilla sp.—l1

V olvulella tenussima—6

pelecypods—Lucinoma annulata—2

polychaetes, inc
annelids

7b. Station 2193-52. Off Redondo Beach pier, in 40 fms, fine sandy

mud. 3.0 cu. ft.

Parvilucina tenuisculpta—14
Solemya panamensis—1|
Tellina bodegensis—1

T hyasira bar barensis—3
Yoldia scissurata—13

luding Pectinaria, a large terebellid, and other

mollusks, including:
scaphopods—Cadulus fusiformis—1

No. | HARTMAN: QUANTITATIVE SURVEY 45

Dentalium rectius—8
gastropods—A cteon punctocoelata—1
Cylichnella diegensis—1
pelecypods—Lucinoma annulata—1|
Nucula carlottensis—1
Pandora filosa—2
Parvilucina tenuisculpta—13
Pitar newcombiana—3
Rochefortia tumida—11
Tellina bodegensis—1
Thyasira barbarensis—1
Yoldia scissurata—19
a commensal crab
polychaetes, including:

Ancistrosy/lis sp—1 or more nephtyid—several
Aricidea sp. and perhaps Ninoé sp.—1

Paraonis sp.—many Nothria sp.—1
capitellids—many, with ?Pareurythoé sp.—1

several species Pectinaria californiensis—
cirratulids—several many
Cossura n. sp.—several Prionospio pinnata—many
Le pidasthenia sp.—1 Prionospio, another species—
Lumbrineris spp.—many several
maldanids—several Spiophanes sp.—several

8. Station 2791-54. Off Palos Verdes Point, in 415 fms, blue-gray
mud. 3.08 cu. ft.
A large sample measuring 3.08 cu. ft. yielded less than a pint of
screenings.
foraminiferans and Phyllochaetopterus tubes only
9. Station 2792-54. Off Palos Verdes Point, in 300 fms, blue-gray
mud. 2.77 cu. ft.
foraminiferans, urchin, echiuroid, gastropod, many annelids, not
yet analyzed
10a. Station 2150-52. Off Redondo Beach pier, in 310 fms, mud. 1.38
Cutt,
urchins—10, pale yellow to pink in life, measuring about 40 mm
across
nemertean—1, bright red ribbonlike in life
scaphopods, possibly Cadulus, many tubes
Chaetoderma—1 or more

46

10b.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

arenaceous foraminiferans, especially Goésella flinti (determined
by Dr. O. Bandy)
many nephtyid and spioniform annelids
Station 2793-54. Off Palos Verdes Point, in 251 fms, blue-gray
mud, much coarse to fine rubble. 0.95 cu. ft.
glass sponge, with long spicules
ophiuroid—1
mollusks, including:
Cadulus sp.—14 or more
Chaetoderma—several
bivalve—1
gastropods—several
crustaceans, including amphipods, isopods
phoxocephalid amphipods:
Harpinia sp. B.—1
Leptophoxus sp. A.—2
burrowing anemone—3
T halassema sp.—2
polyclad—1
many annelids, including:
Ammotry pane sp.—1
ampharetid—2
Hydroides norvegica—2 opercula, possibly from hull of Velero

IV
Lanice sp.—1 large, in tube covered with the foraminiferan,
Goésella flintii

Lumbrineris cruzensis—3

Maldane sp.—more than 50
maldanid—several

Melinna sp.—2

M yriochele sp.—1

Nothria sp.—1

Onuphis, near vexillaria—| or more
Paraonis n. sp.—1

Pectinaria californiensis—1
Petaloproctus sp.—2 or more
Pherusa sp.—2

polynoids—elytra of Lagisca and another polynoid
Prionospio sp.—2

Syllis sp.—1

No. 1 HARTMAN: QUANTITATIVE SURVEY

?T helepus sp.—1, with tube
other annelids
11a. Station 2151-52. Redondo Canyon, in 291 fms, mud. 0.5 cu. ft.
echinoids—2
ophiuroids—many
glass sponge
gastropod, Nitidella sp.— about 30
boring bivalve—many in wood
crustaceans, including:
spider crab—2
shrimp—1
stalked barnacle—1
phoxocephalid amphipod—Heterophoxus sp. A.—1
polychaetes, including:
Acrocirrus ?crassifilis—about 15
Amphicteis s9.—1
A naitides sp.—1
2A sclerocheilus—about 4
capitellid—2
chaetopterid—3 long tubes and
fragments of animals
Cirratulus 2cirratus—1
Cirratulus, another species—1 larger and 3 smaller
?Eumida sp.—1
Euphrosine sp.—1
Evarnella fragilis—1
Glycera tesselata—3
?Hauchiella sp.—1
?Hypoeulalia bilineata—1
Lagisca sp.—1
Laonice sp.—1
Lepidonotus sp.—2
Lumbrineris spp.—9
Nereis sp.—1
Onuphis sp.—1
Pherusa sp.—3
phyllodocid—1
Polydora spp.—about 10, representing 3 species
Protula sp.—several tubes, fully attached to hard surfaces
syllid—3

47

48

11b.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Terebellides sp.—1, pale yellow and orange in life
Station 2361-53. Redondo Canyon, in 167 fms, gray sandy mud.
1.44 cu. ft.

not yet analyzed

lic. Station 2724-54. Redondo Canyon, in 157 fms, fine green sticky

mud with stones (jurassic, metamorphic); much rocky rubble,
gravel and shell fragments. 2.52 cu. ft.
some glass sponge
ophiuroids—few
echiuroids—3, deep green
amphipods—many

phoxocephalid—Heterophoxus sp. A.—7
polychaetes, including:

A naitides sp.—several

Ceratocephala crosslandi americana—|

Drilonereis sp.—1 or more

Glycera sp.—1

Lumbrineris sp.—several

Maldane sp.—many

onuphids—several

polynoid—1 or more

polyodontid tube fragments

spionids and other annelids

. Station 2189-52. Off Redondo Beach pier, in 228 fms, fine sandy

mud. 1.07 cu. ft.

Conspicuous forms include many pelecypods, red Chloeia, ophi-
uroids, urchin, red sand covered Goésella (foraminiferan), and
scaphopods

mollusks, including:
amphineuran, Aplacophora—Chaetoderma sp. B.—13

Limifossor sp.—9

scaphopods—Cadulus tolmiei—33

Dentalium rectius—3
gastropods—Balcis rutila—6

Bittium attenuatum—1
pelecypods—A xionopsis sericatus—12

Cardiomya pectinata—1

Crenella columbiana—1

Macoma incongrua—323

Nuculana conceptionis—42

No. 1 HARTMAN: QUANTITATIVE SURVEY 49

Nuculana spargana—8
Sphenia globula—5
Tellina carpenteri—61
Thyasira barbarensis—1
crustaceans, including:
ostracods—more than 50, with at least 3 species
phoxocephalid amphipod—Heterophexus sp. A—11
cumaceans—5
nemerteans—several
unknown animal—5, reddish purple in life
polychaetes, including:
ampharetid—2 or more, in thick mud-walled tube
another ampharetid—several, in sand-covered tube
A naitides sp.4
?Brada sp.—5
capitellid—1
Chloeia pinnata—more than 90
other cirratulids—several
Cossura n. sp.—2
Glycera americana—1 large
another Glycera sp.—1
Glycinde sp.—1
Goniada sp.—1
Habloscoloplos elongatus—1
maldanids—10 or more
Mar physa sp.—1
Myriochele n. sp.—about 8
nephtyids—several, with 2 or more species
onuphid—1
Pectinaria californiensis—50 or more, including juveniles
Pherusa spp.—several
Polydora sp.—1
polynoid—several
Prionospio ?cirrifera—8
Prionospio pinnata—8
sabellids—2
Scalibregma sp.—1 large and 2 small
Streblosoma sp.—fragment
12b. Station 2360-53. Off Redondo Beach pier, in 49 fms, gray sand,
rock and clay. 1.63 cu. ft.

50

13:

14.

RS

near

15:

16.

Wis

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

not yet analyzed
Station 2359-53. Off Redondo Beach pier, in 31 fms, gray sandy
mud and clay. 0-63 cu. ft.
not yet analyzed
Station 2788-54. South of Redondo Beach, in 9.5 fms, coarse gray
sand and shell. 1.32 cu. ft.
Eight gallons of shell and sandy rubble that did not pass through
the screens, sorted in the laboratory. There were hundreds of mud
filled tests of Dendraster, harboring a sipunculid, each test with
one or more worms. Numerous individuals of a solitary tunicate,
externally covered with bits of shell and sand.
phoxocephalid amphipods—Pontharpinia sp. B.—32
Pontharpinia sp. M.—2

polychaetes, including:

A ricidea jeffreysi—many

Lumbrineris sp.—several, with long head

Mar physa mortenseni—1, large

Pisione, near remota—many
Station 2729-54. Off Point Vicente light, in 445 fms, fine green
sticky mud. 3.4 cu. ft.
foraminiferans, some glass sponge, a large chaetoderm and numer-

ous polychaetes, not yet analyzed.

Station 2728-54. Off Point Vicente light, in 454 fms, fine green
sticky mud, brought to laboratory without screening. Volume not
taken.

foraminiferans and tubes of Phyllochaetopterus only

Station 2474-53. Off Point Vicente light, in 405 fms, green-black
fine mud. 3.9 cu. ft.

foraminiferans and tubes of Phyllochaetopterus only

Station 2362-53. Off Point Vicente light, in 352 fms, fine gray-
green mud. 2.83 cu. ft.

not yet analyzed

. Station 2723-54. Off Point Vicente light, in 325 fms, fine sticky

green mud; most of the sample passed through the screen assembly.

3.4 cu. ft. ;

many foraminiferans

many thick mud tubes formed by at least two polychaetes, including
A mphicteis scaphobranchiata and Maldane sp.

ophiuroid—1

NO.

19.

20.

oA

22.

23:

HARTMAN : QUANTITATIVE SURVEY |

mollusks, including:
Chaetoderma—1 large and several small
Nitidella sp.—several
another gastropod—several
enteropneusts—33 with at least two species, perhaps juveniles
echiuroid—2
sipunculids—several
polychaetes, including (in addition to those named above) :
Aricidea 2jeffreysi—2
capitellid—1
cirratulid—several
Euclymenini—2
?Leocrates sp.—1
Lumbrineris sp.—1
orbiniid, new genus and species—several
Paraonis n. sp.—3
sabellid, possibly unknown—2
sigalionid, possibly Sthenelanella sp.—4
Station 2358-53. Off Point Vicente light, in 125 fms, gray clay.
2.89 cu. ft.
not yet analyzed
Station 2722-54. Off Point Vicente light, in 14 fms, greenish sand
with shell fragments; a very small sample. 0.25 cu. ft.
purple urchins—2
ophiuroids—several
foraminiferans
many smaller annelids, not yet analyzed
Station 2794-54. Off Palos Verdes, in 430 fms, blue-gray mud. 3.4
eu. tt:
foraminiferans, some glass sponge, chaetopterid and serpulid tubes
Phyllochaetopterus sp.
Station 2620-54. Off Point Vicente light, in 418 fms, gray-green
mud. 2.20 cu. ft.
foraminiferans, phyllochaetopterid tubes; little life
Phyllochaetopterus sp.
Station 2475-53. Off Point Vicente light, in 370 fms, fine dark mud.
SiZircuz ft:
much glass sponge
foraminiferans—many
ophiuroids—many large and small

52

24.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

mollusks, including:
Cadulus—5
Chaetoderma—|
gastropod—several
pelecypod—3
Stylatella (sea whip)
anemone—3
amphipod—2
phoxocephalid—Har pinia sp. B.—1
echiuroid—1
polychaetes, including:
ampharetid—many
A ncistrosyllis sp.—2
Laonice sp.—1
Maldane sp.—2 or more
orbiniid—1
Phyllochaetopterus sp.—several tubes
polynoid—1
Protula sp.—1 or more
Station 2476-53. Off Point Vicente light, in 386 fms, fine black
sandy mud, wood fragments. 0.95 cu. ft.
sea whip
enteropneusts—Schizocardium sp.—3
nemertean
anemone—2
mollusks, including:
Cadulus—many
Chaetoderma—1|
gastropods—many
polychaetes, including:
ampharetids—many
capitellid—1
cirratulids—several
?Laonice sp.—1 large, with greatly prolonged branchiae
Lumbrineris sp.—some
maldanid—several
onuphid—juveniles
paraonid—l1, with very long branchiae
Phyllochaetopterus sp.—1 large, in tube
Protula sp.—many, some tubes attached to wood, some clumped

28.

HARTMAN: QUANTITATIVE SURVEY 53

?S piophanes sp.—1
other annelids
Station 2403-53. Off Point Vicente light, in 400 fms, green mud.
3.15-cu: 1!
many foraminiferans
nemertean—a large one
polychaetes, including:
A mphicteis scaphobranchiata—many
other ampharetids
capitellids
terebellids—many
other annelids, not yet analyzed
Station 2357-53. Off Point Vicente light, in 100 fms, fine gray
sandy mud. 1.63 cu. ft.
not yet analyzed
Station 2473-53. Off Point Vicente light, in 16 fms, coarse sandy
mud, shell fragments; very rich in living animals. 0.75 cu. ft.
ophiuroids—many
polychaetes, including:
Chaetopterus sp.—many
cirratulids—many, with several species
Cistenides sp.—several
dexiorbid spirorbids, attached to rocks
Spiophanes sp.—many
Sthenelanella sp.—many
Streblosoma sp.—several
and other annelids
Station 2472-53. Off Point Vicente light, in 11 fms, sandy black
coarse mud, broken shells, considerable debris; diversified fauna.
0.63 cu. ft.
foraminiferans
Glottidia albida—several
nemerteans—several species
nematodes—very many
gastropods and bivalves, including shells of Leda, Yoldia, Lima.
Olivella
anemone—several
Heterocrypta sp. (crab)—several carapaces
amphipods—some
ostracods—many

54 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

polychaetes, including:
Anotomastus gordiodes—several
A phrodita sp.—1
Aricidea spp.—several
Cistenides sp.—1
Dasybranchus sp.—1
Diopatra tridentata—tube fragments
Eumida sp.—1
Exogoninae—several
Halosydna sp.—1
Leocrates n. sp.—1
Lumbrineris spp.—several
Magelona sp.—1
Nephtys sp.—many
nereid—1
Odontosyllis sp.—1
onuphid—1
Pectinaria californiensis—several
Poecilochaetus sp.—several
polynoid—1
Prionospio, near malmgreni—many, the most abundant species
Prionospio pinnata—several
Rhodine sp.—tubes or also living
Scalibregma sp.—1
sigalionids—several
Sthenelanella sp.—some
Streblosoma sp.—1
syllids—some

29. Station 2508-53. In Outer Harbor, in 6 fms, black mud. 2.56

cu. ft.

several small sea whips, some bivalves, nemerteans, anemones, a
few amphipods, a small ophiuroid
polychaetes, including:
ampharetid—1
arabellid—1
capitellid—1
Chaetozone sp.—several
Cossura n. sp.—several
Diopatra sp.—1
Eteone sp.—1

NO.

1

HARTMAN: QUANTITATIVE SURVEY 55

Glycera sp.—1 large
Haploscoloplos sp.—1
Laonice sp.—several
Lumbrineris spp.—many
maldanid—1

Marphysa sp.—10 or more
Nephtys sp.—some

Nereis procera—few
Pectinaria sp.—1

Podarke sp.—1

Prionospio pinnata—several
Scalibregma sp.—1
Spiophanes sp.—1
Terebellides sp.—2

T haryx sp.—hundreds

30. Station 2507-54. In Outer Harbor, off Long Beach breakwater, in

Su

8 fms, black mud ; richly diversified fauna. 3.15 cu. ft.
not yet analyzed
Station 2506-54. In Outer Harbor, in 9 fms, black mud; diversified
fauna. 3.15 cu. ft.
various mollusks
crustaceans, including a ghost shrimp, numerous pinnotherid crabs,
amphipods
ophiuroids—many
enteropneust—Schizocardium sp.—1
polychaetes, including: (specimens not counted )
Ancistrosyllis sp.
Arabella sp.
A mphicteis scaphobranchiata
Brada sp.
Diopatra sp.
disomid
Glycera sp.
Haploscoloplos sp.
Harmothoé sp.
Lumbrineris spp.
maldanid
Mar physa sp.
Megalomma sp.
Nereis sp.

56

a2)

33:

34.

S}5)

36.

37.

38.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Notomastus sp.

Owenia n. sp.

Prionospio pinnata

Scalibregma sp.

Spiophanes sp.

Terebellides sp.

terebellids
Station 2314-53. In Outer Harbor, in 4.5 fms, heavy gray mud.
0.56 cu. ft.

not yet analyzed
Station 2505-53. In Outer Harbor, in 5 fms, black sandy mud; rich
bottom. 0.13 cu. ft.
not yet analyzed
Station 2138-52. Off Point Vicente, in 282 fms, mud and fine
gravel; very small sample. Volume not taken.
brissopsid—2
ophiuroids—many
nemertean—at least |
many annelids, not yet analyzed
Station 2139-52. Off Point Vicente, in 433 fms, mud. 1.89 cu. ft.
not yet analyzed
Station 2619-54. Off Point Vicente light, in 432 fms, gray-green
mud. 3.15 cu. ft.
many foraminiferans
nemertean—1
ophiuroid—1
polychaetes, including:
?Drilonereis—1, white and very slender in life
Lumbrineris sp.—1, dark purple in life
tubes of Phyllochaetopterus, Protula, and onuphid
Station 2363-53. Off Point Vicente light, in 429 fms, fine gray-
green mud. 2.77 cu. ft.
not yet analyzed
Station 2404-53. Off Point Vicente light, in 438 fms, gray sandy
mud. 1.07 cu. ft.
debris contains many bits of wood, fragments of plant stems, fora-
miniferans.
Chaetoderma—1|
scaphopod—1

No. |

39.

40.

4la.

41b.

HARTMAN: QUANTITATIVE SURVEY 57

polychaetes, including:
A mphicteis scaphobranchiata—several
Aricidea sp.—many
cirratulid—4
Lagisca sp.—1
Phyllochaetopterus n. sp.—many with tubes
Polydora sp.—1
serpulid tubes—several long, slender, slightly coiled
terebellid—1
Station 2419-53. Off Point Vicente light, in 437 fms, fine mud.
SLL cust.
foraminiferans
echinoids—2 large
Phyllochaetopterus tubes or specimens
serpulid tubes
a few other animals
Station 2356-53. Off Point Vicente light, in 294 fms, gray-green
mud. 2.57 cu. ft.
not yet analyzed
Station 2220-52. Off Point Fermin light, in 180 fms, fine green
mud. 3.15 cu. ft.
ophiuroids—many
echinoids—several
mollusks, including pelecypods, gastropods, Chaetoderma
smaller crustaceans
many polychaetes, not yet analyzed
Station 2418-53. Off Point Fermin light, in 185 fms, black mud.
2) cu ae
echinoids—3 or more
ophiuroids—some
smaller crustaceans, including amphipods, isopods, cumaceans
mollusks, including Dentalium, pelecypods, gastropods, and Chaeto-
derma
sipunculids
polyclad
nemertean
polychaetes, including:
thick, muddy tubes of ampharetids, onuphids, Pista—many
Ammotry pane sp.—1
A naitides sp.—many

58 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

A ricidea sp. and perhaps Paraonis sp.—several
capitellids
Chloeia sp.
?Chone sp.
cirratulids
Dorvillea sp.—1
Eumida sp.—many
?Eunice sp.—1
Glycera sp.—1
Glycinde sp.—1
Haploscoloplos sp.—1
Harmothoé scriptoria—1
hesionid—1
Laonice sp.—1
Lumbrineris spp.—numerous
maldanids—several
Melinna sp.—1
Nephtys sp.—several
Nothria sp.—1
Onuphis sp.—1
Poweniid
Pectinaria sp.—several
Pherusa sp.—several
Polydora sp.—1
Prionospio pinnata—1
42a. Station 2221-53. Off Point Fermin light, in 147 fms, fine dark
green mud. 2.2 cu. ft.
mollusks
nemerteans—several large
sipunculid
many thick mud-walled tubes of Nothria pallida, terebellid, and
maldanid
polychaetes, including:
A naitides sp.—1 or more
Brada sp.—1
cirratulids
Chloeia pinnata—several
Dorvillea sp.—1
flabelligerids, 2 species—several individuals
Lumbrineris spp.—several

No. | HARTMAN: QUANTITATIVE SURVEY 59

maldanids—several
nephtyids—several
Nothria pallida—many
Pista sp.—some in muddy tubes
orbiniid—1 or more
Pectinaria sp.—several
?Podarke sp.—1
spionids
terebellids
42b. Station 2417-53. Off Point Fermin light, in 34 fms, black mud;
a rich spioniform bottom. 2.14 cu. ft.
sea whip—at least three
Chatoderma—several
polyclad—1
polychetes, including:
Aricidea sp.—1 or more
Chleia pinnata—29 or more
Cossura n. sp.—several
?Eumida sp.—1
Fabricia sp.—several
hesionid—1
Lumbrineris spp.—many
Mar physa sp.—several
nereid—some
orbiniid—several
oweniid—1
Paraonis n. sp.—1 or more
Pectinaria sp.—several small or juveniles
Peisidice aspera—some
Pholoé sp.—several
Polydora spp.—1
Spiophanes sp.—1
other spionids
T haryx, possibly parvus—more than 1530
other small annelids—many
43. Station 2471-53. Off Point Fermin light, in 12 fms, black sandy
mud and shell fragments, very rich in animals, especially smaller
ones. 0.69 cu. ft.
a large living Cardium
many polychetes, especially Chetopterus sp. and spioniform anne-

60 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

lids, not yet analyzed
449. Station 2470-53. Off Point Fermin light, in 10 fms, black sandy
mud, shell fragments, foul odor; very rich in large and small
animals. 1.95 cu. ft.
ophiuroids—many
holothuroids—many
enteropneusts—Schizocardium sp.—8
polychetes, including:
A phrodita sp.—3 or more
Lumbrineris spp.—many
Streblosoma sp.—many
other annelids

44b. Station 2307-53. In Outer Harbor, in 7 fms, black rubbly clay,
friable. 2.14 cu. ft.
ophiuroids
amphipods and other crustaceans
anemones
nematodes
nemerteans—at least two species
sipunculid
polychetes, including:
Amea occidentalis—1
Ampharete 2arctica—3
A mphicteis scaphobranchiata—26
Ancistrosyllis bassi—10
Aricidea sp.—1
Armandia sp.—1 juvenile
Boccardia ?redeki—|
Boccardia sp.—2
Carazzia sp.—1
Chetozone corona—10
Chone sp.—several
Cirratulus ?cirratus—about 10
Cirriformia 2luxuriosa—several
Cossura n. sp.—more than 250
Diopatra tridentata—3 with tubes
Dorvillea articulata—about 10
Eteone californica—1
?Fabricia sp.—1 or more
Glycera americana—2 large and 1 small

NO.

HARTMAN: QUANTITATIVE SURVEY 61

Haploscoloplos elongatus—6

Harmothoé scriptoria—2

Laonice sp.—1

Lumbrineris cruzensis—many

Lumbrineris 2erecta—several

Lumbrineris, other species

Magelona sp.—| or more

Marphysa, resembling conferta—13

Megalomma sp.—3

Melinna sp.—3

Nephtys spp.—about 15

Nereis procera—more than 145, including some epitokes

Notomastus 2hemipodus—about 5

Paraonis n. sp.—more than 92

Pectinaria californiensis—3

Peisidice aspera—32

Pherusa sp.—about 15

Pista sp.—1

Podarke pugettensis—19

Polydora, resembling armata—several

Polydora citrona—9

Polydora, near ligni—several

Prionospio, near malmgreni—8

Prionospio pinnata—3

Spiochetopterus sp.—1 and tubes

Spiophanes missionensis—3

Sthenelanella uniformis—8

Streblosoma crassibranchia—3

Tharyx parvus—hundreds, some ovigerous and with swimming
sete

Travisia sp.—2 juveniles

other annelids, serpulid tubes

45a. Station 2168-52. Off Los Angeles light, in 12 fms, mud with many

large animals. 2.2 cu. ft.
ophiuroids—several

sea whip—1 or more
mollusks, including:

amphineuran, Aplacophora—Chaetoderma sp. A.—1
scaphopods—Cadulus fusiformis—2
Dentalium neohexagonum—1|

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

gastropods—A glaja sp.—1
Crepidula nivea—15
Ferreria belcherii—1
Nassarius per pinquis—2
V olvulella tenussima—3
pelecypods—Macoma yoldiformis—1
Nuculana taphira—2
Pitar newcombiana—1
Solen rosaceus—2
Tagelus californianus—several
Thyasira barbarensis—2
cephalopod—Octopus apollyon—1
crustaceans, including 2 small crabs, a shrimp, 2 cumaceans
phoronids—many in slender stiff tubes externally covered with sand
Glottidia albida—1 small
enteropneusts—Schizocardium sp.—3
nemerteans—about 8
polyclads—4 large, of which 3 come from crevices in the dead conch
leech—1 small
other mollusks include: many pelecypods, gastropods, some living
Crepidula on the dead conch
polychetes, including:
A mphicteis scaphobranchiata—many
Ancistrosyllis rigida—8
A phrodita parva—2
Asychis sp.—several
Brada ?pluribranchiata—7
capitellids—many
Cossura, n. sp.—7
Diopatra tridentata—about 10 with tubes
Drilonereis sp.—many
Glycera sp.—2 juveniles
Goniada sp.—6
Halosydna latior—3, in crevices on dead conch
Harmothoé scriptoria—4
Laonice cirrata—19
Lepidasthenia virens—1
Lumbrineris spp.—many
Magelona, near pacifica—3
maldanids—many

NO.

HARTMAN: QUANTITATIVE SURVEY 63

Mar physa, resembling conferta—18
Melinna sp.—about 5, juveniles

Nereis procera—12, bright green in life
Nothria sp.—many

Owenia sp.—2 large

Pectinaria californiensis—9

Pherusa sp.—about 20

Pholoé sp.—about 6

Phyllodoce sp.—several

Pista sp—many

Podarke sp.—4

Pecilochetus johnsoni—about 12
Prionospio, near malmgreni—8
Prionospio pinnata—about 30
Scalibregma sp.—28

Scoloplos sp.—5 juveniles
Spiochetopterus sp.—several
Spiophanes sp.—about 14
Sthenelanella uniformis—7
Streblosoma crassibranchia—many and tubes
T erebellides sp.—2

T haryx sp.—several

T yposyllis sp.—2, deep orange color in life
other annelids

45b. Station 2217-53. Off Los Angeles light, in 12 fms, shale, chert

rocks, wood fragments, hard packed black mud; many animals.
Oislicus ft:
ophiuroids and asteroids—many
holothuroid—1 large and 3 small
mollusks, including chitons, rock oysters, slipper shells, and others
sipunculids, resembling Golfingia—many, in tortuous burrows in
shale
burrowing anemone—2
nemerteans—3
crustaceans, including a crab, many amphipods, and isopods
Glottidia albida—1
phoronid—1 or more
echiuroid, possibly T halassema—1 small
polychetes, including:
ampharetids—several

64

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

amphinomid—1 small

A naitides sp.—1

A phrodita sp.—6

arabellid—several

Boccardia sp.—6

capitellid—2

Ceratonereis sp.—3

Chatozone sp.—several

Chone sp.—several
cirratulids—many

Cistenides sp.—about 10

dextral spirorbids, on foliose bryozoan
Dorvillea sp.—2

Eteone sp.—1

Eumida sp.—1

Euphrosine—1 juvenile
Exogoninae—1, on bryozoan
flabelligerids—at least 2 species and several individuals
Glycera sp.—1 juvenile

Halosydna sp.—3

Haploscoloplos elongatus—2
Harmothoé scriptoria—several
Hesperalia sp.—1

Leocrates n. sp.—1

Lepidonotus sp.—more than 25
Lumbrineris spp.—many
Megalomma sp.—several

M yxicola sp.—1| juvenile

onuphid—4

Owenia sp.—2

Peisidice sp.—1

Pherusa inflata—many, in U-shaped burrows in clay
Phyllochetopterus, resembling prolifica—about 15
Pista, resembling cristata—several
Pista elongata—1| or more

Podarke sp.—several

Sabellaria sp.—many

Salmacina sp.—clumps of tube masses
Scalibregma sp.—3
spionids—numerous small

No. 1 HARTMAN: QUANTITATIVE SURVEY 65

Sthenelanella sp.—several
Sylline—about 6
terebellids—numerous
T halenessa sp.—several
Tharyx sp.—1
46a. Station 2107-53. Off Los Angeles light, in 13 fms, sand and mud,
much life. 2.0 cu. ft.
sea anemone—1
sea whip—2
Glottidia albida—2
mollusks, including:
chetoderman—1
scaphopods—3
pelecypods—Solen rosaceus—6
Tagelus californianus—6, the longest 8 cm long
Trachycardium quadragenarium—1
sipunculids—8
echiuroid, Listriolobus pelodes—2
phoronids—more than 3
crustaceans, including caprellids, amphipods, ostracod, a juvenile
brachyuran, and a cumacean
nemertean—2
enteropneusts—Schizocardium sp.—2
polychaetes, including:
2A mage sp.—6
A mphicteis sp.—2
Ancistrosyllis bassi—6
Asychis 2lacera—1
Boccardia sp.—2 or more
Brada sp.—1
Capitella sp.—1, male
Chaetozone sp.—18
Cossura n. sp.—20
Diopatra tridentata—more than 3, longest measures about
170 mm
Drilonereis ?nuda—9, longest measures about 100 mm
Exogoninae—1
Glycera ?capitata—1
Haploscoloplos elongatus—5
?Isolda sp.—11

66 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Laonice sp.—1 or more

Lumbrineris index—8

Lumbrineris japonica—\0

Lumbrineris minima—17

Marphysa, resembling conferta—5, longest measures about
80 mm

Nephtys ?caecoides—2

Nereis procera—31, longest measures 38 mm

Notomastus sp.—11

opheliid, anterior fragment

Owenia sp.—2, longest measures 80 mm

paraonids—9

Pectinaria californiensis—5, largest tube measures 61 mm long

Pherusa sp.—several

Phyllochaetopterus sp.—1 or more

phyllodocid—1

Poecilochaetus johnsoni—5 or more

polynoid—2

Scalibregma sp.—12

Spiophanes ?missionensis—12 or more

Sternas pis sp.—2

Sthenelais ?tertiaglabra—3

Sthenelanella uniformis—31

Streblosoma sp.—3 or more

46b. Station 2202-53. Off Los Angeles light, in 11 fms, mud, many
animals. 1.89 cu. ft.
Tagelus californianus—7 large
other mollusks, including scaphopods, gastropods, pelecypods
enteropneust, Schizocardium sp.—3
echinoderms, including an ophiuroid, a holothuroid
nemerteans—many, with several species
echiuroid—1
sipunculid—1
phoronid—l1 or more
smaller crustaceans, including an amphipod, 4 copepods
Glottidia albida—2 large
polychaetes, including:
ampharetids—many
?A mphicteis scaphobranchiata—1
Arabella sp.—2 or more

No. | “HARTMAN: QUANTITATIVE SURVEY 67

Artacaminae, new genus and species—1
Asychis lacera—1 with tube
A xiothella sp.—many
Boccardia sp.—several
Brada sp.—several
capitellids—several
?Ceratonereis sp.—I1 or more
cirratulid—several
Cossura n. sp.—3
Diopatra tridentata—many with tubes
Drilonereis sp.—1
Glycera sp.—1
Goniada sp.—1
Haploscoloplos elongatus—2
Harmothoé scriptoria—3
hesionid—2.or more
Laonice sp.—several
Lepidasthenia sp.—1
Lumbrineris spp.—at least 4 species and about 20 specimens
Magelona sp.—several
Marphysa sp.—1
nephtyid—3
Nereis sp.—several
Nothria sp.—2 or more
Paraonis n. sp.—1
Pherusa ?inflata—1 or more, and other species of the genus—
many
Phyllodoce sp.—3
Pista sp.—several
Poecilochaetus sp.—several with many tubes
Praxillella sp.—many
Prionospio sp.—several
Scalibregma sp.—about 40
sigalionids—at least 2 species and many specimens
Spiophanes sp.—many
Sternas pis sp.—2
other annelids
47. Station 2309-53. Off Long Beach breakwater, in 12 fms, fine
black mud. 1.32 cu. ft.
not yet analyzed

68

48.

49.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2313-53. Off Long Beach breakwater, in 11 fms, fine gray
mud. 0.37 cu. ft.
Richly diversified, abundant fauna, not yet analyzed
phoxocephalid amphipods — Pontharpinia sp. B.—2
Pontharpinia sp. Q.—1
Station 2312-53. Off Long Beach breakwater, in 7.5 fms, brown
sandy mud and gravel, perhaps detritus from Santa Ana River.
Richly diversified fauna. 0.7 cu. ft.
a large ovigerous cancer crab
Lepidopa myops, a spiny sand crab—1
mollusks—various and numerous, especially gastropods, pelecypods
echinoderms—many ophiuroids, some echinoids, a sea star
Glottidia albida—1|
a sea whip
nemerteans, sipunculid, nematodes—many
small crustaceans, including amphipods, isopods
phoxocephalid amphipods—20, including Har pinia sp. C_—2
Pontharpinia sp. B.—7
Ponthar pinia sp. G.—1
Pontharpinia sp. M.—8
polychaetes, including:
Aricidea sp.—1 or more
Chaetozone sp.—1
Chone sp.—1
Eteone sp.—1
Euchone sp.—1
Goniada sp.—1
Harmothoé ?imbricata—1
Lumbrineris spp.—many
Magelona sp.—several
maldanids—many
Mar physa sp.—1
nephtyids—more than | species, many
Nereis ?procera—several
Nothria ?elegans—many tubes
Notomastus sp.—1 or more
Onuphis eremita—1
Ophelia sp.—1
Owenta sp.—1 large
Perinereis sp.—1

NO.

a.

54.

1

HARTMAN: QUANTITATIVE SURVEY 69

Poecilochaetus sp.—1|

Polycirrus sp.—1

Prionospio, near malmgreni—several
Prionospio pinnata—1

Rhynchospio sp.—several
Scalibregma sp.—1

Scoloplos sp.—1

Spiophanes sp.—several

other annelids

Station 2741-54. Off Los Angeles light, in 5 fms, red beach sand
with broken shells; most of sample failed to pass through the
screens. 2.15 cu. ft.
many juveniles of Dendraster and echinoid
Crenella decussata (identified by Dr. T. Soot-Ryen)—many
many polychaetes, including:

Saccocirrus papillocercus—many

juveniles of Anaitides, Dorvillea, Glycera, Owenia, and others

Station 2730-54. Off Point Vicente light, in 297 fms, green rocky
mud. 1.44 cu. ft.
not yet analyzed
Station 2795-54. Off Point Vicente, in 290 fms, much rock and
dark mud; black phosphorite rocks coated black, also basaltic and
other metamorphic rocks not penetrated by animals but some with
attached tubes of annelids, especially maldanids and terebellids.
0.95 cu. ft.
brissopsid, nemertean, small sponge masses on rocks
polychaetes, including:
Lanice sp.—long tube with tesselated tip, externally covered
with arenaceous foraminiferan Goésella flintt
maldanids, in tubes fully attached to rocky surfaces
spionid—few
other annelids not yet analyzed
Station 2432-53. Off Point Vicente light, in 450 fms, silty mud.
3:59 cu. ft
foraminiferans, Phyllochaetopterus and Protula tubes. Few animals.
Station 2420-53. Off Point Vicente light, in 458 fms, fine greenish
mud. 3.15 cu. ft.
foraminiferans, tubes of Phyllochaetopterus and Protula, dead
translucent shells of Pecten. Little life.

70
5D:

56.

Sie

58a.

58b.

59a.

59b.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2405-53. Off Point Vicente light, in 457 fms, sandy mud.
Ze Cutts
foraminiferans, tubes of Phyllochaetopterus and Protula.
Station 2322-53. Off Point Vicente light, in 460 fms, fine grey-
green mud. 2.77 cu. ft.
foraminiferans, small amounts of glass sponge, tubes and animals
of Phyllochaetopterus and Protula, an ampharetid, a nemertean;
little life.
Station 2431-53. Off Point Vicente light, in 431 fms, fine mud.
ZOZicMeits
foraminiferans, tubes of Phyllochaetopterus and Protula, little life.
Station 2110-52. Off Point Fermin light, in 427 fms, oozy mud;
little life. Volume not taken.
foraminiferans, tubes of Phyllochaetopterus, serpulid
Glycera 2branchiopoda—2
ampharetid—2 juveniles.
Station 2321-53. Off Point Fermin light, in 385 fms, fine gray-
green mud. 2.83 cu. ft.
echinoderms, including 2 ophiuroids, 1 holothurian
crustaceans, including amphipods and an isopod
nemerteans, sipunculids, and echiuroids
many polychaetes, not yet analyzed
Station 2218-53. Off Point Fermin light, in 249 fms, fine green
mud. 2.83 cu. ft.
Richly diversified, including:
foraminiferans
ophiuroids, echinoids, asteroids
mollusks, including scaphopods
amphineurans, Aplacophora—C haetoderma sp. A.—1
Chaetoderma sp. C.—1
Limifossor sp.—9
gastropods—WNitidella gouldi—10
a macruran
numerous diversified polychaetes, not yet analyzed
Station 2219-53. Off Point Fermin light, in 235 fms, fine dark
green mud. 2.96 cu. ft.
mollusks, including gastropods, scaphopods, Chaetoderma
echiuroid—l1 large and 1 small
nemertean—1 large ribbon-like
polychaetes, including:

NO.

60.

HARTMAN: QUANTITATIVE SURVEY 71

Asychis sp.—1, in thick mud-walled tube
capitellid—several
Chloeia pinnata—many
Maldane sp.—many
Onuphis sp.—in thick mud-walled tube
orbiniid, new genus and species—1
Pectinaria californiensis—1 or more
Pista ?cristata—1
terebellids and other annelids
Station 2430-53. Off Point Fermin light, in 80 fms, hard packed
mud with many animals. 1.44 cu. ft.
ophiuroids—many
crustaceans, including amphipods and ostracods
phoxocephalid amphipods—25, including:
Harpinia sp. A.—2
Heterophoxus sp. A—2
Phoxocephalus sp. A—6
Pontharpinia sp. E—15
mollusks, including Chaetoderma, many tubes of Dentalium, pele-
cypods
polychaetes, including: (specimens not counted )
ampharetids
Chloeia sp.
goniadid
Harmothoé scriptoria
Lumbrineris sp.
maldanids
Myriochele sp.
Nephtys ferruginea
Pherusa sp.
Pholoé sp.
Prionospio, near malmgrent
Prionospio pinnata
Spiophanes sp.
T haryx sp.

61a. Station 2315-53. Off Los Angeles light, in 20 fms, mud, shell

fragments; richly diversified fauna. 1.07 cu. ft.

not yet analyzed
phoxocephalid amphipods: Metaphoxus sp. A—3
Pontharpinia sp. J.—2

he ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

61b. Station 2618-54. Off Point Fermin light, in 21 fms, muddy sand,
dead shells. 0.44 cu. ft.
echinoderms, including:
ophiuroids—many
asteroid—1 small
holothuroids—a large papillate and a small smocth one
mollusks, including:
Pecten shells—many
tectibranch—l, with large lateral wings
Chaetoderma—several
crustaceans, including a decapod, amphipods, and isopods
phoxocephalid amphipods: Heterophoxus sp. A—1
Pontharpinia sp. E.—3
Ponthar pinia sp. K.—1
nemerteans
burrowing anemone
enteropneust—5
polychaetes, including:
ampharetid
Aricidea sp.
Boccardia sp.
capitellids
cirratulids, several species
Dorvillea sp.
Drilonereis sp.
Eulalia sp.
Eunice sp.
flabelligerids
Glycera sp.
Haploscoloplos sp.
Hyalinoecia sp.
Lumbrineris spp.
maldanids
Nephtys sp.
Pectinaria sp.
Pholoé n. sp.
Phyllochaetopterus ?prolifica
phyllodocids
Polydora sp., boring in dead gastropod shell
Scalibregma sp.

No. l

62.

63a.

HARTMAN: QUANTITATIVE SURVEY 73

sigalionids
Spiophanes sp.
Sternaspis sp.
Sthenelanella sp.
?Streblosoma
other annelids
Station 2446-53. Off Los Angeles breakwater light, in 14 fms,
compact sandy clay. Sample very small, about 0.15 cu. ft.; not yet
analyzed.
Station 2113-52. Off Long Beach light, in 13 fms, red sand, failed
to pass through the screens; much life. 0.95 cu. ft.
echinoderms, including 5 ophiuroids, 5 small echinoids, and a large
brissopsid
crustaceans, including a large sand crab (Lepidopa), amphipods,
cumaceans
phoxocephalid amphipod : Ponthar pinia sp. M—8
burrowing anemone—12
phoronids—more than 15

sea whip—1
nemerteans—2
polyclad—3

mollusks, including 5 scaphopods and others
polychaetes, including:

ampharetid—1 in tube

Aricidea spp.—several

Boccardia sp.—1

Chone sp.—1

?Eumida sp.—1

Lumbrineris spp.—3

Magelona sp.—2

Nephtys caecoides—7

Nothria ?elegans—2

Onuphis eremita—4

O phelia sp.—7

Prionospio spp.—several

Rhynchospio arenincola—55 or more

Scoloplos sp.—1

Spiophanes missionensis—8

syllid—7

T halenessa spinosa—2 large and 1 small

74

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

63b. Station 2445-53. Off Los Angeles breakwater light, in 11 fms,

coarse red sand. 0.81 cu. ft.

Failed to pass through screens; one pail of sand, or about 0.3 cu. ft.
of unscreened and unfixed sand, was sorted in the laboratory and
found to yield richly in polychaetes, notably Dorvillea, Magelona,
Aricidea, Paraonis, and Anaitides species, anemones, and nematodes.
The sand is rich in mollusks, including Caecidae and other gastro-
pods, scaphopods, and pelecypods of various genera, and circular
disk-like colonies of bryozoans. Other animals include:
Dendraster (sand dollar )—several living, to 23 mm across

nudibranch—1
phoxocephalid amphipod : Ponthar pinia sp. M.—1
phoronids—at least 10
anemone (Harenactis ?attenuata, according to the late Dr. W. K.
Fisher )—many hundreds
sipunculids—many
polychaetes, including: (numbers of individuals not counted )
A naitides sp.
Aricidea sp.
A xiothella sp.
Chaetozone sp.
Chone sp.
cirratulids, deep yellow in life
Dorvillea 2gracilis—hundreds
Drilonereis sp.
goniadid
Harmothoé sp., very dark, deeply depressed, resembling a com-
mensal
Leocrates sp., juvenile
Lumbrineris sp.
Magelona sp.
maldanids, the largest polychaetes in the sample
Nephtys sp.
onuphid, in fine red sand-covered tube
Paraonis sp.
Pisione, near remota
Rhynchospfio sp.
Scalibregma sp.
Scoloplos sp.—1
Spiochaetopterus sp.

No. 1 HARTMAN: QUANTITATIVE SURVEY 15

spionids

syllid, long, slender, with pink ova
T halenessa spinosa

other annelids

64. Station 2310-53. Off Long Beach breakwater light, in 15 fms,
sandy mud. 0.37 cu. ft.
several ophiuroids, a small asteroid
mollusks, including pelecypods, gastropods, scaphopods
phoxocephalid amphipods, including Pontharpinia sp. K.—23
Ponthar pinia sp. Q.—10
Pontharpinia sp. V.—11
ostracods
Glottidia albida—\ large
nemerteans ; nematodes; sipunculids; some hydroid branches
others not yet analyzed
65. Station 2291-53. Off Los Angeles breakwater, in 14 fms, fine gray
sandy mud. 0.31 cu. ft.
many ophiuroids, some asteroids
several mollusks
phoxocephalid amphipods—3, including Pontharpinia sp. B.—1
Pontharpinia sp. K.—1
numerous other small crustaceans
Glottidia albida—2 large
nemerteans
polychaetes, including:
A maea occidentalis—3
A mphicteis scaphobranchiata—many
capitellid—1
Chone sp.—5 or more
cirratulid spp.—many
Diopatra tridentata—several
Drilonereis sp.—2
flabelligerids—about 10
Gontada sp.—3
Haploscoloplos elongatus—1
Laonice sp.—1
Lumbrineris spp.—several
Magelona sp.—7
Maldane sp.—1
M elinna sp.—many

76

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

nephtyid—1

nereid—3

Nothria 2geophiliformis—many in tubes
Nothria spp.—several

phyllodocids, several species—many
Pista, possibly cristata—4
?Polycirrus sp.—1 small, ovigerous
Praxillella sp.—1

Prionospio, near malmgreni—several
Prionospio pinnata—1

Rhodine sp.—1 and tubes
Scalibregma sp.—1

Schistocomus sp.—4

Spiophanes sp.—many

Sternaspis sp.—2 large and 9 small
Sthenelais ?tertiaglabra—\
Sthenelanella uniformis—many
syllid—7, in tubes

T halenessa sp.—several

66a. Station 2311-53. Off Long Beach breakwater light, in 12 fms, hard

Dentalium rectius—8

packed sandy mud, rich in living animals. The grabbing device
failed to penetrate; two successive grabs yielded only 0.5 cu. ft.
echinoderms, including ophiuroids, holothurians, asteroids (2 large)
mollusks, including scaphopods, gastropods, pelecypods, chaetoderm
(1 long)

crustaceans, including a cancroid crab, 4 commensal crabs, amphi-

pods, ostracods, phoxocephalid amphipods, 89, including:

Glottidia albida—2
nemerteans—several
nematodes—many
sipunculid—1 or more
leech—1

a few branches of hydroids

Pontharpinia sp. B.—1
Pontharpinia sp. G.—1
Ponthar pinia sp. K.—26
Pontharpinia sp. Q.—26
Pontharpinia sp. R.—12
Ponthar pinia sp. V.—21

HARTMAN: QUANTITATIVE SURVEY

polychaetes including: (counts not made)
A mphictets scaphobranchiata
Anotomastus gordiodes
Arabella sp.
A ricidea spp.
Brada sp.
Chaetozone corona (many)
Chone and perhaps other Fabricinae
cirratulids
Cossura sp. (many)
Diopatra tridentata (many)
Drilonereis spp.
Goniada sp.
Haploscoloplos elongatus
Harmothoé sp.
Laonice sp.
Lepidasthenia sp.
Lumbrineris cruzensis
Lumbrineris minima
Lumbrineris spp.
Magelona sp.
maldanids
Naineris n. sp.
Nephtys spp.
nereids
Nothria spp.
Paraonis sp.
Pherusa sp.
Pholoé sp.
phyllodocids
Pilargis sp.
Pista sp.
Podarke sp.
Poecilochaetus sp.
Prionospio, near malmgreni
Prionospio pinnata
Rhynchospio sp.
Scalibregma sp.
Schistocomus sp.
spionids

78

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Spiophanes sp.
syllid
terebellids

T halenessa sp.
Tharyx sp.

66b. Station 2606-54. Off East Jetty light, Anaheim Bay, in 13 fms,

67.

68.

69.

compact black sandy mud. 0.13 cu. ft.
sea stars; many annelids, not yet analyzed
phoxocephalid amphipods: Pontharpinia sp. B.—7
Ponthar pinia sp. Q.—5
Pontharpinia sp. R.—4
Ponthar pinia sp. V.—4
Station 2504-53. Off Los Angeles light, in 9 fms, black sandy mud.
0.13 cu. ft.
not yet analyzed
Station 2607-54. Off Huntington Beach pier, in 5 fms, compact
black sand. 0.06 cu. ft.
phoxocephalid amphipods, 29, including: Pontharpinia sp. E.—1
Ponthar pinia sp. K.—1
Ponthar pinia sp. S.—10
Pontharpinia sp. T.—14
other animals not yet analyzed
Station 2324-53. Off Point Vicente light, in 400 fms, fine mud
with glass sponge. 3.08 cu ft.
foraminiferans; small calcareous sponge ; 10 small anemones
an enteropneust; compound ascidians attached to needles of glass
sponge
mollusks, including:
amphineurans, Aplacophora — Chaetoderma sp. A—1
Chaetoderma sp. B.—2
neomeniinid sp.—1
gastropods — Leptogyra sp.—6
Nitidella permodesta—25
pelecypods — Kellia sp.—4
crustaceans, including:
pycnogonids—1 1
phoxocephalid amphipods—Heterophoxus sp. A.—3
isopod—36
galatheid crab—1
polychaetes, including:

NO.

70.

7A.

12.

(ES

74.

(Ey

76.

HARTMAN: QUANTITATIVE SURVEY 79

amphinomid—1

Aricidea sp.—3

Autolytus sp.—1 epitoke

capitellid—2

Chone ecaudata—more than 200

cirratulid—2 or more

Harmothoé triannulata—about 125

hesionid—1

Lagisca sp.—many

Lumbrineris sp.—1

Myriochele sp.—1

orbiniid, new genus and species—fragments

Paraonis sp.—1

Phyllochaetopterus sp.—2

Protula sp.—more than 200

sabellid—1 or more

Spionid—1

T yposyllis sp.—several
Station 2433-53. Off Point Vicente light, in 460 fms, fine sandy
mud. 2.07 cu. ft.
foraminiferans, tubes of Phyllochaetopterus and Protula
Station 2421-53. Off Point Vicente light, in 466 fms, fine greenish
mud. 3.08 cu. ft.
foraminiferans, tubes of Phy/lochaetopterus and Protula, dead shells

of Pecten
Station 2323-53. Off Point Vicente light, in 480 fms, fine mud.
Sol 7 cu. sft,
foraminiferans, tubes of Phyllochaetopterus and Protula
Station 2406-53. Off Point Vicente light, in 475 fms, fine greenish
mud. 3.4 cu. ft.
foraminiferans, tubes of Phyllochaetopterus
Station 2740-54. Off Point Vicente light, in 470 fms, fine green
mud. O.31 cw. tt
foraminiferans, tubes of Phyllochaetopterus, dead Pecten shells
Station 2334-53. Off Point Fermin light, in 437 fms, fine mud.
302 :cu, ft:
foraminiferans, tubes of Phyllochaetopterus
Station 2335-53. Off Point Fermin light, in 420 fms, fine mud,
with glass sponge and diversified animals. 2.58 cu. ft.
not yet analyzed

80

77.

78.

7h

80a.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2336-53. Off Point Fermin light, in 355 fms, fine greenish
mud. 2.83 cu. ft.
Diversified fauna, with ophiuroids, echiuroids, foraminiferans, and
polychaetes ; not yet analyzed.
Station 2317-53. Off Point Fermin light, in 280 fms, greenish
mud. 2.83 cu. ft.
not yet analyzed
Station 2316-53. Off Los Angeles breakwater light, in 117 fms,
dark mud. 2.7 cu. ft.
not yet analyzed
Station 2125-52. Off Los Angeles light, in 29 fms, fine mud and
shaley rubble. Volume not taken.
ophiuroids
Glottidia albida
mollusks, including:
gastropod, 4 glaja sp.—1
pelecypods, Tellina sp., and others
crustaceans, including amphipods, isopods, ostracods, cumaceans
phoxocephalid amphipod—Ponthar pinia sp. B.—3
phoronids ;
enteropneust—1
polychaetes, including:
Ammotry pane sp.
Aricidea sp.
Chaetozone sp.
Chloeia pinnata
cirratulids
Fabricinae
Lumbrineris spp.
Magelona sp.
maldanids
Myriochele sp.
Nephtys sp.
onuphid
Paraonis sp.
Pectinaria sp.
Pherusa sp.
phyllodocids
Pista sp.
Prionospio spp.

NO.

HARTMAN: QUANTITATIVE SURVEY 81

sabellids
Scoloplos sp.
sigalionid
Spiophanes sp.
syllid

T erebellides sp.
terebellids
Tharyx sp.

80b. Station 2233-53. Off Point Fermin light, in 23 fms, fine mud.

O31 custe
echinoderms, including ophiuroids, holothuroids, echinoids
anemones ; nemerteans
mollusks, including:
scaphopod, Cadulus tolmiei—1
gastropod, 4 glaja sp.—2
chaetodermans
egg cluster of Loligo sp. (squid)
crustaceans, including amphipods, an isopod, a cumacean
phoxocephalid amphipods, 31, including Metaphoxus sp. A—4
Pontharpinia sp. E—11
Pontharpinia sp. J|.—13
Ponthar pinia sp. L.—2
polychaetes, including:
ampharetids of several genera—6 juveniles
Aricidea sp.—14
Artacaminae, new genus and species—1 large and 5 small
Capitella ovincola—hundreds, in egg cluster of Loligo
Chone sp.—13
cirratulid—7
Drilonereis sp.—2
Halosydna sp.—1
Haploscoloplos elongatus—12
Hyalinoecia juvenalis—1
?Leanira sp.—2
Lumbrineris spp.—11
Magelona sp.—2
maldanids—about 10
?Maldanella robusta—fragments
Nephtys sp.—6
Nereis sp.—1

82

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Nerine foliosa n. subsp.—1
Nothria sp.—1

?Notomastus sp.—3 juveniles
Onuphis sp.—4

Owenia sp.—2

Pectinaria californiensis—1
Pherusa sp.—4

Pholoé sp.—9
phyllodocids—2

Pista, resembling cristata—9
Prionospio, near malmgreni—45
Prionospio pinnata—2
Rhamphobrachium sp.—1
Rhynchospio arenincola—11
Sabella sp.—2

Scalibregma sp.—4 juveniles
Schistocomus sp.—1 or more
sigalionids

Spiophanes sp.—9
Sthenelais sp.—fragment
Sthenelanella uniformis—13
syllid—1

T erebellides sp.—1

other annelids

8la. Station 2124-52. Off Los Angeles light, in 16 fms, shell and fine

81b.

rubble. Volume not taken.
Richly diversified spioniform fauna; not yet analyzed.

Station 2444-53. Off Los Angeles breakwater light, in 18 fms,
black mud and shells. 0.31 cu. ft.
many ophiuroids and 2 small holothurians

Glottidia albida, burrowing anemone, bryozoan colonies (circular

disk-like)

derma

mollusks, including pelecypods, gastropods, scaphopods, a Chaeto-

crustaceans, including a Heterocryptus sp., amphipods, and others

phoxocephalid amphipods, 32, including Metaphoxus sp. A—6

Pontharpinia sp. B.—18
Pontharpinia sp. K.—3
Pontharpinia sp. Q.—4
Pontharpinia sp. S.—1

No. 1 HARTMAN: QUANTITATIVE SURVEY 83

polychaetes, including:
A naitides sp.
Ancistrosyllis bassi
Aricidea sp.
Chaetozone sp.
Chone sp.
Dexiospira sp.
Dorvillea sp.
Eulalia sp.
Eumida sp.
goniadid
Harmothoé scriptoria
Laonice sp.
Lepidasthenia sp.
Lumbrineris spp.
Mazgelona sp.
nereid
Nothria sp.
Pholoé sp.
Praxillella sp.
Prionospio, near malmgrent
Rhynchospio sp.
Scoloplos sp.
Sternaspis sp.
Streblosoma sp.
Tharyx sp.
and others

82. Station 2496-53. Off Los Angeles breakwater light, in 18 fms,
dark sandy mud. 0.5 cu. ft.
echinoderms, including ophiuroids, 2 holothurians, an asteroid
sea whip—2 ; anemones—about 6
bryozoan colonies
Glottidia albida—6
mollusks, including:
amphineuran, Aplacophora — Chaetoderma sp. A.—1
Chaetoderma sp. B.—2
scaphopods — Cadulus fusiformis—5
Dentalium neohexagonum—\
gastropods — 4 glaja sp.—|
Balcis rutila—2

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Eulima caltfornica-—2
O phiodermella incisa—2
pelecypods — 4 xionopsis sericatus—2
Lyonsia californica—l
Nuculana taphira—4
Parvilucina tenuisculpta—2
Rochefortia tumida—3
Solemya volvulus—8
Tellina buttoni—1

crustaceans, including many amphipods, a caprellid, isopods, several

cumaceans, 2 pycnogonid ; phoxocephalid ampipods, 32, including
Metaphoxus sp. A—12
Pontharpinia sp. B.—3
Ponthar pinia sp. Q.—19
Ponthar pinia sp. V.—5

polychaetes, including:

Ampharete sp.—4

Asychis lacera—1

Brada sp.—2

Chone sp.—3

cirratulids—5

Drilonereis sp—1

Euclymenini—3

Eumida sp.—2

goniadid—2

Hyalinoecia sp.—3

Labidognathus sp.—2, endoparasitic in Tharyx sp.

Lumbrineris sp.—more than 3

Magelona sp.—5

Melinna sp.—2

Nephtys sp.—1

nereid—1

Owenia sp.—1

paraonid—2

Pherusa spp.—many

Pholoé sp.—15

Pista sp.—3

Praxillella sp.—several

Prionospio, near malmgreni—8

Prionospio pinnata—2

NO.

83.

84.

85.

near

1

HARTMAN: QUANTITATIVE SURVEY 85

Scalibregma sp.—1
Sternaspis sp.—8
Sthenelanella sp.—18
Streblosoma sp.—2
Terebellides sp.—1

T halenessa spinosa—13
Tharyx sp.—l1 or more
other annelids

Station 2501-53. Off Los Angeles light, in 19 fms, black compact

oily mud. 0.1 cu. ft.

phoxocephalid amphipods, 18, including Metaphoxus sp.A—1
Ponthar pinia sp. B.—2
Ponthar pinia sp. Q.—6
Pontharpinia sp. V.—9

other animals not yet analyzed

Station 2502-53. Off Los Angeles light, in 18 fms, black compact

oily mud. 0.15 cu. ft.

not yet analyzed

Station 2503-53. Off Los Angeles light, in 16 fms, black compact

oily mud. 0.13 cu. ft.

not yet analyzed

Station 2645-54. Off Huntington Beach pier, in 12 fms, greenish-

gray sand. 0.26 cu. ft.

not yet analyzed

Station 2646-54. Off Huntington Beach pier, in 12 fms, sandy
mud. 0.3 cu. ft.
A measured quart, unscreened sample, analyzed in the Jaboratory.
some ophiuroids ; many nematodes; hydroids
crustaceans, including amphipods, isopods
phoxocephalid amphipods, including Pontharpinia sp. K.—11
Pontharpinia sp. Q-—5
Pontharpinia sp. R.—8
polychaetes, including:
arabellid
Aricidea sp.
cirratulid
Diopatra tridentata
Goniada sp.
Lumbrineris spp.

86

87.

88.

89.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Magelona sp.
Nephtys sp.
nereid

Nothria sp.
Prionos pio spp.
sabellid
Scoloplos sp.
Sphaerodorum sp.
spionids

syllid
terebellids
other annelids

Station 2608-54. Off Huntington Beach pier, in 7.5 fms, compact
black sand. 0.06 cu. ft.
not yet analyzed
Station 2609-54. Off Huntington Beach pier, in 4.5 fms, compact
black sand. 0.13 cu. ft.
phoxocephalid amphipods: Ponthar pinia sp. Q.—1
Pontharpinia sp. S.—3
other animals not yet analyzed
Station 2798-54. Off Catalina Island light, in 386 fms, blue-
green-gray mud, glass sponge, fish scales, otoliths, foraminiferans,
and radiolarians. 2.96 cu. ft.
an ophiuroid ; several nemerteans; a ghost shrimp
mollusks, including 2 gastropods, a pelecypod, a scaphopod, Chaeto-
derma
enteropneust, Stereobalanus sp.—1
polychaetes, including:
ampharetids—many
A ricidea sp.—several
capitellid, new genus and species—3
Chloeia sp.—1 juvenile
Chone sp.—1
Glycera branchiopoda—1
?Tsolda or related genus—1
Lumbrineris cruzensis—1
maldanid—1
orbiniid, new genus and species—2
Phyllochaetopterus sp.—tube fragments
Protula sp.—tube fragments

NO.

90.

I

92.

93.

94.

J/5y

o6:

OF:

HARTMAN: QUANTITATIVE SURVEY 87

Spiophanes sp.—l1
Station 2325-53. Off Ship Rock, Catalina Island, in 475 fms, fine
greenish mud. 2.77 cu. ft.
foraminiferans, Phyllochaetopterus and Protula tubes
Station 2799-54. Off Catalina Island light, in 484 fms, blue-green-
gray mud. 3.15 cu. ft.
foraminiferans, tubes of Phyllochaetopterus and Protula sp., shells
of Pecten; also small bits of wood, fish vertebrae, and otoliths
Station 2739-54, Off Catalina Island light, in 478 fms, fine green
mud. 3.15 cu. ft.
foraminiferans, tubes of Phyllochaetopterus, and dead Pecten shells
Station 2407-53. Off Point Vicente light, in 480 fms, fine greenish
mud, a large black oil sludge. 3.4 cu. ft.
foraminiferans, tubes of Phyllochaetopterus, serpulid tubes
Station 2364-53. Off Point Vicente light, in 495 fms, fine mud.
Zi. tt:
foraminiferans, tubes of Phyllochaetopterus
Station 2386-53. Off Point Fermin, in 460 fms, fine mud. 3.3
cus ft:
foraminiferans, tubes of Phyllochaetopterus and Protula
Station 2222-53. Off Point Fermin light, in 440 fms, oozy mud.
3. 15icus ft:
glass sponge; ophiuroids; mollusks, including gastropods, scapho-
pods, pelecypods ; nemerteans
polychaetes, including:
ampharetid—1
cirratulid—1
Melinna sp.—1, in long, thick mud-walled tube about 14 inches
long
Pista sp.—1, in thick mud-walled tube
Protula—several, in white cylindrical tubes
Station 2835-54. Off Point Fermin light, in 370 fms, greenish-
gray sticky mud, otoliths, many foraminiferans. 3.21 cu. ft.
gastropod shell; Chaetoderma—1; enteropneust, possibly Stereo-
balanus—1 ; nemertean—2
polychaetes, including:
A mphicteis scaphobranchiata—1| large and several small
other ampharetids—several
capitellid—1
chaetopterid tubes

88

98.

99a.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Glycera sp.—2
Maldane sp.—2
other maldanids—several
orbiniid—1
Paraonis sp.—1 or more
Prionospio sp.—1
Protula tubes
Station 2306-53. Off Point Fermin light, in 215 fms, clay and
mud. 2.64 cu. ft.
echinoderms, including an ophiuroid, an echinoid, a holothurian
mollusks, including many scaphopods, gastropods, pelecypods,
Chaetoderma
crustaceans, including amphipods, isopods, and copepods
phoxocephalid amphipods, including Harpinia sp. A—6
Heterophoxus sp. A—1
polychaetes, including:
Chloeia pinnata—about 50
Longosoma catalinensis—1 or more
maldanids—many
other annelids
Station 2355-53. Off Los Angeles breakwater light, in 41 fms,
sandy mud. 0.1 cu. ft.
many ophiuroids and annelids; not yet analyzed

99b. Station 2629-54. Off Point Fermin light, in 50 fms, green

sandy mud, passed quickly through screens. 0.18 cu. ft.
crustaceans, including ostracods—hundreds ; amphipods, cumaceans,

and slender linear white isopods—many

phoxocephalid amphipods: Metaphoxus sp. A.—9

Ponthar pinia sp. R.—6

gastropods, pelecypods—many
polychaetes, including:

ampharetids

Artacaminae, new genus

Chloeia sp.

Fabricinae

Myriochele sp.

nephtyids of several species

Pectinaria sp.

Pholoé sp.

Prionospio, near malmgreni

no. 1 HARTMAN: QUANTITATIVE SURVEY 89

syllid
Terebellides sp.
100. Station 2232-53. Off Point Fermin light, in 33 fms, fine mud and
shell. 0.37 cu. ft.

many foraminiferans ; 3 anemones; many ophiuroids
mollusks, including gastropods, pelecypods, scaphopods, and Chaeto-
derma
crustaceans, including amphipods, isopods, and 12 or more cuma-
ceans
phoxocephalid amphipods, 18, including Metaphoxus sp. A—2
Pontharpinia sp. B.—2
Ponthar pinia sp. K.—2
Ponthar pinia sp. L.—4
Pontharpinia sp. P.—4

1 Listriolobus pelodes, nematodes
polychaetes, including:
ampharetids—3 small
Aricidea sp.—17 or more
cirratulids—about 12
Drilonereis sp.—1
Glycera sp.—1
Haploscoloplos elongatus—12
Harmothoé sp.—2
Lumbrineris spp.—about 8
Mazgelona sp.—1
maldanid—1
Myriochele sp.—12 or more
Nephtys sp.—about 10, including 2 large
onuphids—8
Owenia sp.—3 tubes
Pherusa sp.—6 juveniles
Pholoé sp.—about 10
Prionospio spp.—about 20
sabellids—5
Scalibregma sp.—2
sigalionids—several
spionids—several
Spiophanes sp.—1
terebellids—several

90

101.

102.

103.

104.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2394-53. Off Point Fermin, in 29 fms, sandy mud and
broken shells. 0.25 cu. ft.
echinoderms, including 2 urchins, an ophiuroid, holothurians
Glottidia albida
phoxocephalid amphipods, 7, including Pontharpinia sp. B.—1
Ponthar pinia sp. P.—5

a parasitic copepod on a maldanid in sandy tube
polychaetes, including:

Aricidea sp.

capitellid—1 large

Chiloeia sp.—juveniles

Chone sp.

cirratulids

Harmothoé sp.

Lumbrineris sp.

maldanids

Nereis sp.

Onuphis sp.

Peisidice sp.

Prionospio sp.

Scalibregma sp.

Scoloplos sp.

Spiochaetopterus sp.

Spiophanes sp.

Sthenelanella sp.

T haryx sp.

other annelids

Station 2497-53. Off Los Angeles breakwater light, in 24 fms,
sandy mud. 0.16 cu. ft.

ophiuroids; broken shells; many tubicolous annelids; not yet
analyzed

Station 2374-53. Off Los Angeles breakwater, in 27 fms, sandy
mud. 0.62 cu. ft.

diversified fauna; not yet analyzed

Station 2292-53. Off Los Angeles breakwater, in 60 fms, fine mud.

0.81 cu. ft.

echinoderms, including many ophiuroids and one echinoid

mollusks, including pelecypods, gastropods, scaphopods, aplacopho-
ran

NO.

105.

106.

107.

108.

HARTMAN: QUANTITATIVE SURVEY 91

crustaceans, including amphipods, isopods, ostracods, cumaceans,
a copepod, phoxocephalid amphipods, 114, including
Heterophoxus sp. A.—3
Heterophoxus sp.B.—2
Metaphoxus sp. A—1
Pontharpinia sp. E—58
Pontharpinia sp. |.—48
Pontharpinia sp. R—1
nemerteans; an anemone; nematodes, a turbellarian
numerous polychaetes, not yet analyzed
Station 2630-54. Off Huntington Beach pier, in 45 fms, sandy
gray-green mud. 0.86 cu. ft.
many ophiuroids; a large JTravisia and numerous smaller poly-
chaetes ; other animals; not yet analyzed
Station 2611-54. Off Huntington Beach pier, in 19 fms, compact
gray-black sand. 0.06 cu. ft.
phoxocephalid amphipods, including:
Metaphoxus sp. A—1
Pontharpinia sp. B.—3
Pontharpinia sp. K.—2
Pontharpinia sp. P.—5
Pontharpinia sp. R.—2
other animals not yet analyzed
Station 2610-54. Off Huntington Beach pier, in 18 fms, compact
sand. 0.25 cu. ft.
phoxocephalid amphipods, 18, including:
Metaphoxus sp. A.—2
Pontharpinia sp. E—9
Pontharpinia sp. K.—2
Pontharpinia sp. R—4+
Station 2114-52. Off Newport Beach pier, in 17 fms, mud and
Sana. 1¢26.cuy ft:
echinoderms, including ophiuroids, holothurians, asteroids
many small anemones
Glottidia albida—1
phoronid tube
mollusks, including gastropods, pelecypods, scaphopods, 2 nudi-
branchs, Chaetoderma
crustaceans, including amphipods, isopods, and others
nemerteans, sipunculids, nematodes, polyclads

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

polychaetes, including:
Amaea occidentalis—2
Ancistrosyllis sp.—3 juveniles
A phrodita sp.—1 juvenile
Aricidea spp.—about 36
Asychis 2lacera—5
Autolytus sp.—1
Boccardia n. sp.—23
capitellid—5
Chone sp.—about 7
cirratulid—about 11
Cossura n. sp.—)5
Diopatra tridentata—9
Drilonereis ?falcata—5
Ehlersia heterochaeta—20
Eteone sp.—1
flabelligerid, resembling Brada—20 juveniles
Glycera tesselata—\4
goniadids—19
Haploscoloplos elongatus—6
Harmothoé sp.—14 juveniles
Hyalinoecia ?juvenalis—1
Lepidasthenia sp.—2 juveniles
Loandalia fauveli—|
Lumbrineris californiensis—2
Lumbrineris cruzensis—14
Lumbrineris spp.—3, one with very long prostomium
Magelona, near pacifica—10
maldanid—1 large, with 5 or 6 posterior apodous segments
other maldanids—about 6
Marphysa, resembling conferta—1
Melinna and other ampharetids—about 10 juveniles
Nephtys caecoides—7
another Nephtys sp.—about 20
Nereis procera—l1 adult and about 13 juveniles
Nerine foliosa n. subsp.—2
Nothria iridescens—4 large and 9 small
paraonids—about 50
Pectinaria californiensis—S5
Pherusa sp.—about 75 juveniles

No. 1 HARTMAN: QUANTITATIVE SURVEY 93

Pholoé sp.—23

phyllodocids—about 5

Pilargis ?maculata—1

?Pionosyllis sp.—2

Pista 2cristata—4

another Pista sp.—about 20

Podarke sp.—1

Poecilochaetus johnsoni—\|

Praxillella affinis pacifica—5

Prionospio, near malmgreni—more than 60

Prionospio pinnata—31

Rhynchospio arenincola—5

Scalibregma sp.—2

sigalionid—2

Sphaerodorum spp.—2

Spiochaetopterus sp.—2

Spiophanes missionensis—about 20

Sternaspis sp.—18

Sthenelanella uniformis—5 or more

Streblosoma sp.—1

Terebellides sp.—2

T halenessa spinosa—4

Travisia sp.—4

109. Station 2115-52. Off Newport Beach pier, in 61 fms, mud and

sand. 1.51 cu. ft.
ophiuroids and a holothuroid
pelecypods, gastropods, scaphopods
phoxocephalid amphipod : Pontharpinia sp. E.—1
isopod—1
echiuroids, including Listriolobus pelodes and another—about 12
a nemertean, a polyclad, nematodes
polychaetes, including:

?A maea sp.—1

Ammotrypane sp.—2

A naitides sp.—2

capitellid—fragments

Chloeia sp.—2

cirratulid—1

Glycera sp.—1

Glycinde sp.—1

110.

BES

LZ:

near

EZ.

143.

114.

sy

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Haploscoloplos elongatus—2
Lumbrineris sp.—several
Magelona sp.—1
maldanid—several
Nephtys sp.—2
Nereis sp.—1 epitoke
Pectinaria sp.—45, largely juveniles
Pherusa sp.—1
Poecilochaetus johnsoni—1
polynoid—1
Prionospio pinnata—2
Scalibregma sp.—6
spionids—several
spirorbid—1
terebellid—1
other annelids
Station 2745-54. Off Balboa Beach pier, in 8 fms, gray sand and
shell fragments. 0.27 cu. ft.
mollusks and diversified annelids, including Chaetopterus sp.; not
yet analyzed
Station 2326-53. Off Ship Rock, Catalina Island, in 385 fms, fine
greenish mud. 2.52 cu. ft.
not yet analyzed
Station 2732-54. Off west end of Catalina Island, in 464 fms,
green mud. 3.15 cu. ft.
foraminiferans, tubes of Phyllochaetopterus

Station 2731-54. Off west end of Catalina Island, in 450 fms,
green mud, unscreened. Volume not taken.
foraminiferans, tubes of Phyllochaetopterus; no other metazoans

Station 2327-53. Off Ship Rock, Catalina Island, in 490 fms,
fine greenish mud. 3.15 cu. ft.

foraminiferans, tubes of Phyllochaetopterus

Station 2434-53. Off Ship Rock, Catalina Island, in 480 fms,
fine mud. 3.15 cu. ft.

foraminiferans, tubes of Phyllochaetopterus, and serpulid

Station 2166-52. Off Arrow Point, Catalina Island, in 470 fms,
oozy mud. 3.15 cu. ft.

foraminiferans, tubes of Phyllochaetopterus

LW.

118.

lUICP

120.

HARTMAN: QUANTITATIVE SURVEY 95

. Station 2800-54. Off Ship Rock, Catalina Island, in 478 fms,

blue-green-gray mud. 3.27 cu. ft.
foraminiferans, tubes of Phyllochaetopterus, dead shells of Pecten
Station 2333-53. Off Long Point, Catalina Island, in 487 fms,
fine mud. 2.52 cu. ft.
foraminiferans, tubes of Phyllochaetopterus
Station 2454-53. Off Long Point, Catalina Island light, in 460
fms, fine mud. 2.96 cu. it.
foraminiferans, tubes of Phyllochaetopterus
Station 2500-53. Off Point Fermin light, in 450 fms, fine greenish
sticky mud, much glass sponge, diversified fauna. 3.33 cu. ft.
foraminiferans
many ophiuroids, with several species
mollusks, including scaphopods, at least one living Pecten, another
pelecypod, and 2 Chaetoderma
a galatheid crab, Munidopsis depressa Faxon—1 male (identified
by Miss Janet Haig)
a large nemertean
polychaetes, including:
Lagisca sp.—10 or more
Protula sp.—100 or more
Sabellinae, one new genus and species—many
terebellids—several

Station 2628-54. Off Long Point, Catalina Island, in 350 fms,
green sandy mud. 6.1 cu. ft.
many foraminiferans; shark’s tooth; otoliths ; a white holothurian
mollusks, including several scaphopods and a large Chaetoderma
several nemerteans
polychaetes, including:

Amphicteis 2scaphobranchiata—several

capitellid, new genus and species—1 large

flabelligerid—1

Maldane sp.—several

Myriochele sp.—2

orbiniid, new genus and species—1

sabellid—1

121a. Station 2354-53. Off Los Angeles breakwater light, in 200 fms,

muddy clay and rocks. 1.95 cu. ft.
not yet analyzed

96

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

121b. Station 2625-54. Off Point Fermin light, in 230 fms, gray-green

122;

sticky mud. 5.25 cu. ft.
foraminiferans
echinoderms, including 3 echinoids, 2 ophiuroids
a sea whip; a burrowing anemone
many amphipods, including phoxocephalids: Harpinia sp. A—8
Harpinia sp. B.—1
Heterophoxus sp.
B.—1
Paraphoxus sp. A—1
Phoxocephalus sp.A.—1
mollusks, including scaphopods and pelecypods
polychaetes, including:
Ancistrosyllis ?rigida
Aricidea sp.—1| large
Asychis lacera— 1 large, with tube
?Brada sp.—1
Fabricinae—several
Maldane sp.—many
another maldanid—1
Melinna sp.—1
Nephtys sp.—1
Nothria sp.—1 or more
Paraonis sp.—1
Pectinaria sp.—1
polynoid—1
Prionospio pinnata—1 or more
Station 2231-53. Off Point Fermin light, in 115 fms, sandy clay.
1.76 cu. ft.
foraminiferans ; echinoderms, including ophiuroids, holothuroid
mollusks, including:
amphineuran, Aplacophora — Chaetoderma sp. A.—1
Limifossor sp.—3
gastropods — Volvulella californica—1
pelecypods — A xionopsis sericatus—3
Nucula cardara—2
crustaceans, including ostracods and phoxocephalid amphipods:
Heterophoxus sp. A—11
Metaphoxus sp. A—6
Ponthar pinia sp. E.—10

No. 1 HARTMAN: QUANTITATIVE SURVEY 97

polychaetes, including:
A mphicteis scaphobranchiata—2
Aricidea ?pacifica—3
cirratulids—several
Drilonereis sp. — 2
Glycera sp.—1
Harmothoé sp.—3
Lumbrineris sp.—4
maldanids—several
Melinna sp.—2 and tubes
Nephtys sp.—5
Notomastus lobatus—4
Onuphis parva—32
Pherusa sp.—2
Pholoé sp.—3
Pista 2cristata—1
Prionospio pinnata—3
Rhamphobrachium sp.—1
Spiochaetopterus sp.—tubes
spionids—several
Spiophanes sp.—3
Sternaspis sp.—1
Terebellides sp.—1
123a. Station 2126-52. Off Los Angeles light, in 48 fms, rubbly bottom.
Large sample, volume not taken.
coralline clumps; many ophiuroids; pelecypods, 2 chitons, gastropods
T halassema sp.—1
a leech
polychaetes, including:
A glaophamus sp.—1
Ammotry pane sp.—2 or more
ampharetids—several in more than one species
Chaetozone sp.—several
Chloeia sp.—many
Drilonereis sp.—several
Glycera sp.—1
goniadid—1
Lumbrineris spp.—many
Magelona sp.—1 or more
maldanid—several

98

ALLAN HANCOCK PACIFIC EXPEDITIONS

Megalomma sp.—1|

M yriochele sp.—several
Nephtys sp.—several
Oncoscolex sp.—1
Onuphis sp.—many
Owenia sp.—several

paraonids—many, possibly Aricidea and Paraonis spp.

Pectinaria sp.—several
Pherusa spp.—several

Pista spp.—several
polynoid—several

Prionospio spp.—many
sabellid—many

Scalibregma sp.—1

Scoloplos sp.—several
sigalionids—several
sphaerodorid—1
Spiochaetopterus or other chaetopterid—1 or more
spirorbids—several

Sternaspis sp.—1

Sthenelanella sp.—several
syllid—1

Thalenessa spinosa— 2 or more
T haryx sp.—many

voL. 19

123b. Station 2443-53. Off Los Angeles breakwater light, in 60 fms,

pinnules

mollusks, including Te/lina and Chaetoderma
crustaceans, including amphipods, isopods, ostracods, and cumaceans
polychaetes, including:

Chaetozone sp.
Chloeia pinnata
cirratulids

Glycera sp.
Harmothoé scriptoria
Laonice sp.
Lumbrineris spp.

dark mud, sand and shells, much coarse rubble and clumps of
bryozoans. 0.5 cu. ft.
ophiuroids—many

crinoid (Florometra)—2, ovigerous, with bright orange eggs along

NO.

HARTMAN: QUANTITATIVE SURVEY

maldanids

Megalomma sp.
Myriochele sp.

Nephtys sp.
Odontosyllis sp.
Oncoscolex sp.

Onuphis parva
Pectinaria californiensis
Pherusa capulata
Pholoé sp.

Prionospio spp.
sigalionid

sinistral spirorbids—many, on bryozoan colonies
Sternaspis sp.

T halenessa spinosa
Travisia sp.

99

124. Station 2498-53. Off Los Angeles breakwater light, in 50 fms,

sandy mud, considerable pepper-colored sand or fine gravel.

cu. tt.

many ophiuroids; a dark purple holothuroid
numerous pelecypods, some gastropods
ostracods—more than 100

many isopods, tanaids, and others ; cumaceans

phoxocephalid amphipods, including:
Heterophoxus sp. A—1
Pontharpinia sp. B.—2
Ponthar pinia sp. J. —2
Pontharpinia sp. P.—11

polychaetes, including:
?A maea sp.—1
arabellid, parasitic in body cavity of Tharyx fragment
Aricidea sp.—several
Chloeia sp.—more than 50
Harmothoé scriptoria—1| or more
Magelona sp.—2
Paraonis sp.—1
T halenessa sp.—1
Tharyx sp.—1
and others

1.0

100

125.

126.

127,

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2615-54. Off Huntington Beach pier, in 50 fms, gravelly

mud, shell and rubble. 0.63 cu. ft.

parts of a large sea star ; many ostracods, amphipods

polychaetes numerous, including Onuphis sp. in long slender tubes,
and others; not yet analyzed

Station 2614-54. Off Huntington Beach pier, in 155 fms, dark
green mud. 3.02 cu. ft.
an ophiuroid ; 2 larger brissopsids; 6 purple holothurians
mollusks, including:
pelecypods, especially 4 cila sp.
a tectibranch
scaphopods, and others
a large nemertean; a large deep green echiuroid and a smaller one
sipunculids—2
polychaetes, including:
A glaophamus sp.
ampharetids
A naitides sp.
Ancistrosyllis sp.
Brada sp.
Glycera sp.
goniadid
harmothoid
Laonice sp.
Lumbrineris spp.
maldanids
Melinna sp.
?Mesochaetopterus sp.
Nephtys ferruginea
Nothria sp.
Prionospio pinnata
Terebellides sp.
and others
Station 2613-54. Off Huntington Beach pier, in 138 fms, dark
green mud, silt. 1.0 cu. ft.
foraminiferans; otoliths of fishes
a large purple urchin; 2 purple holothurians
mollusks, including a tectibranch, gastropods, pelecypods, Cadulus,
Chaetoderma
ostracods

no. I HARTMAN : QUANTITATIVE SURVEY 101

polychaetes, including:
A glaophamus sp.
Chloeia sp.—abundant
cirratulids
Cossura sp.
?Diopatra sp.
Gontada sp.
?Leocrates sp.
Lumbrineris sp.
Myriochele sp.
Nephtys sp.
Pectinaria sp.
Prionospio pinnata
and others
128. Station 2612-54. Off Huntington Beach pier, in 100 fms, dark
green mud. 2.64 cu. ft.
many ophiuroids; a large purple urchin; 2 holothurians
mollusks: Dentalium rectius (scaphopod) and Acila sp. (pele-
cypod )
amphipods, phoxocephalid :
Heterophoxus sp. A—3
Phoxocephalus sp. A—1
polychaetes, including:
ampharetids
Aricidea sp.
Cossura sp.
Glycera sp.
goniadid
Harmothoé scriptoria
?Leocrates sp.
Lumbrineris sp.
Maldane sp., in thick mud tubes
Melinna sp.
Nephtys sp., numerous
Pectinaria sp.
Pholoé sp., several mature
Prionospio, near malmgreni
Prionospio pinnata
Sternaspis sp.
and others

102

129:

130.

dS.

132;

133;

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2631-54. Off Newport Beach pier, in 50 fms, sandy gray-

green mud. 0.71 cu. ft.

numerous ophiuroids, broken shells, sand-dwelling annelids, espec-
ially Chloeia sp., and others; not yet analyzed

Station 2742-54. Off Newport Beach pier, in 105 fms, sticky green

mud. 5.31 cu. ft.

many large thick muddy tubes, especially of Nothria pallida and
maldanids

other animals not yet identified

Station 2743-54. Off Newport Beach pier, in 150 fms, sticky
green mud. 5.59 cu. ft.
large brissopsids ; many tube worms. Similar to number 130.

Station 2744-54. Off Newport Beach pier, in 155 fms, greenish-
gray mud. 7.1 cu. ft. Much did not pass through the screens; about
5 quarts of debris.
echinoderms, including brissopsid and holothurian
echiuroid, including 2 large Thalassema sp.—deep red with white
proboscis
nemerteans
polychaetes, including:
many onuphids in thick large muddy tubes
Maldane sp., in similar though slenderer tubes
nephtyids
others not yet analyzed
Station 2294-53. Off Newport west jetty light, in 47 fms, fine
mud and clay. 1.76 cu. ft.
foraminiferans
echinoderms, including many ophiuroids and 3 echinoids
coelenterates
mollusks, including gastropod, pelecypods, scaphopods
crustaceans, including many amphipods, ostracods, isopods, cuma-
ceans, a copepod; phoxocephalid amphipods, 126, including
Heterophoxus sp. A—30
Metaphoxus sp. A—5
Paraphoxus sp. A—1
Phoxocephalus sp. A.—6
Ponthar pinia sp. E—66
Ponthar pinia sp. J.—18
nemerteans, sipunculids, a turbellarian

No. 1

HARTMAN : QUANTITATIVE SURVEY

polychaetes, including:

ampharetid—several
Aricidea—several

Capitella sp.—1, and another capitellid
Ceratocephala crosslandi americana—1
Chloeia pinnata—1
cirratulids—many

Cossura sp.—2

Glycera sp.—about 5

Glycinde sp.—3

Harmothoé sp.—1

Lumbrineris spp.—many

Maldane sp.—1

maldanids—some

Megalomma sp.—1

Myriochele sp.—several
nephtyid—many

onuphids—about 6

Panthalis sp.—1

Paraonis sp.—many

Pectinaria californiensis—many
Pholoé sp.—several

Pista sp.—1 large in mud-walled tube
Podarke sp.—1

Praxillella sp.—many
sigalionid—several

Sphaerodorum sp.—2

Spiophanes sp.—several

103

other spionids, including Laonice, Prionospio, or others—many

Sternas pis sp.—3
Travisia sp.—2

134. Station 2746-54. Off Newport Beach light, in 13 fms, fine gray

135;

sand. 0.7 cu. ft.
some shelly debris; sea stars, gastropods, many annelids; not yet

analyzed

T halassema; many annelids; not yet analyzed

Station 2737-54. Off west end, Catalina Island, in 256 fms, fine
green mud. 2.77 cu. ft.

foraminiferans; purple urchin; nemertean; echiuroid, possibly

104

136.

137.

138.

139:

140.

141.

142a.

142b.

143.

144.

145.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2738-54. Off west end, Catalina Island, in 342 fms, fine
green mud. 3.15 cu. ft.
foraminiferans ; many smaller annelids; not yet analyzed
Station 2735-54. Off west end, Catalina Island, in 435 fms, fine
green mud. 3.27 cu. ft.
foraminiferans, tubes of Phyllochaetopterus, a few other annelids
Station 2388-53. Off Ship Rock, Catalina Island, in 482 fms, fine
green mud. 3.1 cu. ft.
foraminiferans, tubes of Phyllochaetopterus
Station 2422-53. Off Ship Rock, Catalina Island light, in 484 fms,
fine greenish mud. 3.15 cu. ft.
foraminiferans, tubes of Phyllochaetopterus and Protula, dead
shells of Pecten
Station 2387-53. Off Ship Rock, Catalina Island, in 482 fms, fine
mud. 2.8 cu. ft.
foraminiferans, tubes of Phyllochaetopterus, serpulid, and dead
Pecten shells
Station 2304-53. Off Ship Rock, Catalina Island, in 470 fms, fine
greenish mud. 2.7 cu. ft.
foraminiferans; tubes of Phyllochaetopterus and serpulid; a pele-
cypod
Station 2146-52. Off Point Fermin, in 490 fms, oozy mud.
3.46 cu. ft.
foraminiferans, tubes of Phyl/ochaetopterus and serpulid
Station 2332-53. Off Long Point, Catalina Island, in 490 fms,
fine mud. 2.68 cu. ft.
foraminiferans, tubes of Phyllochaetopterus
Station 2499-53. Off Long Point, Catalina Island, in 483 fms,
fine mud. 3.27 cu. ft.
foraminiferans; glass sponge; radiolarians; shells of Pecten; an
ostracod ; and otoliths
polychaetes, including:
arabellid—1
Lumbrineris sp.—1
Phyllochaetopterus sp.—3 and many tubes
Protula sp.—1 and many tubes
Station 2627-54. Off Long Point, Catalina Island, in 455 fms,
gray-green sandy mud. 5.75 cu. ft.
foraminiferans, tubes of phyllochaetopterids

Station 2836-54. Off Point Fermin light, in 430 fms, greenish-gray

No. 1 HARTMAN : QUANTITATIVE SURVEY 105

sticky mud, foraminiferans, small polychaetes. 2.88 cu. ft.
sea star—l
mollusks, including Chaetoderma—1 ; Nitidella—1 ; pelecypod—1 ;
Pecten shells
crustaceans, including ghost shrimp—l1 ; isopod—1
Stereobalanus sp.—1
polychaetes, including:
ampharetid—1
A ncistrosyllis sp.—3
Aricidea jeffreysi—3
capitellid—4
hesionid—2
maldanid—1 or more
Protula sp.—tubes and 2 tentacular crowns

146. Station 2230-53. Off Long Point, Catalina Island, in 300 fms,
fine dark green mud. 3.27 cu. ft.
ophiuroids, echinoids; 3 nemerteans; mollusks, including pele-
cypods, gastropods, scaphopods, Chaetoderma
crustaceans, including amphipods, with phoxocephalid amphipod
Har pinia sp. ?>D.—3
1 Listriolobus pelodes
polychaetes, including:
ampharetid—2
Ancistrosyllis sp.—2
Aricidea sp.—8
?Brada sp.—4
capitellid—4
cirratulid—l1 or more
Cossura sp.—1
Euchone sp.—1
Glycera sp.—1
Maldane sp.—7
Melinna sp.—4
Myriochele sp.—2
Pherusa sp.—1
Podarke sp.—1
sabellid—2 larger and 2 smaller
Scoloplos sp.—1
Spiophanes sp.—1
other annelids

106

147.

148.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station 2442-53. Off Los Angeles breakwater light, in 220 fms,
sticky mud. 2.52 cu. ft.
not yet analyzed

Station 2337-53. Off Los Angeles breakwater light, in 170 fms,

fine mud. 1.51 cu. ft.

some glass sponge; many foraminiferans, including Goesella flintit;
otoliths

ophiuroids — many; echiuroids — 2 large; holothurians —3 dark
purple

mollusks, especially Tellina, Cadulus, gastropods, Chaetoderma

crustaceans, especially amphipods and isopods, including the phoxo-
cephalid amphipod Heterophoxus sp. B—5
nemerteans—several small
polychaetes, including:
Aricidea sp.
Brada sp.
capitellid
Chloeia sp.
Eunice sp.—2 large, in chitinized tubes with lateral branches
Glycera sp.
goniadid
Harmothoé scriptoria
Lanice sp.—in tube adorned with Goésella (foraminiferan)
Leocrates sp.
Lumbrineris spp.
Maldane sp.
Nephtys sp.
Nothria sp.
Paraonis sp.
Pectinaria sp.
Pholoé sp.
Pilargis sp.
Prionospio pinnata
Rhodine sp.
Scoloplos sp.
?S piochaetopterus sp.
?S piophanes sp.
Terebellides sp.
Travisia sp.

NO.

149,

i50.

St.

HARTMAN: QUANTITATIVE SURVEY 107

Station 2127-52. Off Los Angeles light, in 125 fms, mud and

sand, some rock. Volume not taken.

Considerable slime, resembling that of a large dead polyodontid ;
poor in living organisms but with some ChJoeia sp. and nephtyid.

Station 2447-53. Off Los Angeles breakwater light, in 208 fms,

compact sticky clay and mud. 2.2 cu. ft.

not yet analyzed

Station 2373-53. Off Los Angeles breakwater light, in 220 fms,

muddy clay. 2.7 cu. ft.

not yet analyzed

. Station 2840-54. Off Los Angeles breakwater light, in 205 fms,

sticky green mud. 2.6 cu. ft.
brissopsids, holothurians; mollusks; echiuroids; many annelids, not
yet analyzed

153 and 154 not yet sampled

15).

156.

Station 2632-54. Off end of Newport Beach pier, in 230 fms,

sandy gray-green mud. 5.45 cu. ft.

2 urchins; echiuroids; sipunculids; chaetopterid tubes and many
other annelids, not yet analyzed

Station 2750-54. Off end of Newport Beach pier, in 238 fms,

greenish mud. 2.77 cu. ft.

brissopsids, holothurians; nemerteans; many annelids, not yet ana-
lyzed

. Station 2749-54. Off Abalone Point, in 277 fms, green mud.

2.63 Cu. ft.

brissopsids ; many larger and smaller annelids, not yet analyzed

. Station 2748-54. Off Abalone Point, in 167 fms, oily sticky green

mud. 2.45 cu. ft.
foraminiferans, an otolith, numerous thick mud-walled tubes
ophiuroids—several
purple holothuroid
brissopsid—2
mollusks, including:

Chaetoderma—2

pelecypods—2 or more

scaphopods—few

gastropods
nemertean—2
polyclad—1
crustaceans, including amphipods, 2 cumaceans, an ostracod

about 10

108 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

polychaetes, including:

goniadid—3

Lumbrineris sp.—1

maldanid—1

nephtyid—numerous small ones, more than one species

Nothria sp.—several, in thick mud-walled tubes lined with
chitinized sheath

Pectinaria californiensis—about 10, with tubes

pilargiid fragment

Pista disjuncta—several, in very thick mud-walled tubes

Prionospio pinnata—1| or more

other spionid—several

159a. Station 2116-52. Off Abalone Point, Laguna Beach, in 26 fms,
mud. 2.45 cu. ft.
ophiuroids—more than 100; holothuroids—more than 25
anemones—about 6
mollusks—many, including 3 Chaetoderma
crustaceans—many amphipods, and other small ones, and 6 stalked
barnacles
phoxocephalid amphipods, including: Heterophoxus sp. A—50
Metaphoxus sp. A—3
Pontharpinia sp. E.—26
Pontharpinia sp. J.—13
echiuroids—8
sipunculid—more than 1
phoronids—many tubes
nemerteans—several small
Glottidia albida—12
enteropneusts, Stereobalanus sp.—19
polychaetes, including:
Ammotry pane sp.—2
A mphicteis sp.—1
Ancistrosyllis sp.—1
Asychis sp.—2 or more
Ceratocephala crosslandi americana—3
Chloeia pinnata—more than 50
cirratulids—many
Cossura n. sp.—many
Diopatra tridentata—\| large
Drilonereis sp.—5

NO.

HARTMAN: QUANTITATIVE SURVEY

Euclymenini—many

Eunice americana—2 in tubes
Glycera sp.—15

goniadid—9

Haploscoloplos elongatus—5
Harmothoé scriptoria—6 (one has a parasitic copepod )
Hyalinoecia juvenalis—4
Lumbrineris californiensis—1
Lumbrineris spp.—many

Magelona, near pacifica—many
Maldane sp.—1

Myriochele sp.—1 or 2

Nephtys sp.—1 large and three small
Nereis sp.—several

Notomastus sp.—about 6
paraonids—9 or more

Pectinaria sp.—many

Pherusa sp.—1| large and 4 small
Pholoé sp.—3

Pista sp.—1 large in tube

Podarke sp.—6

Poecilochaetus sp.—5

Polydora spp.—3 or more
Polyodontes, near panamensis—1 large
Prionospio, near malmgreni—19
Prionospio pinnata—about 15
Rhodine sp.—many, with tubes
Sabellaria sp.—about 3

Scalibregma sp.—1

serpulid tubes

Spiochaetopterus sp.—1

spionid—1 or more

Spiophanes missionensis—many, with tubes
spirorbid—1

Sternaspis sp.—28

Sthenelanella uniformis—10 or more
syllid—1

Terebellides sp.—about 18

Travisia sp.—1

other annelids

109

110

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

159b. Station 2295-53. Off Newport west jetty light, in 26 fms, dark

160.

LGt:

grayish mud. 0.69 cu. ft.
enteropneusts:
Balanoglossus sp.—1
Stereobalanus sp.—11
other animals not yet analyzed
Station 2736-54. Off west end, Catalina Island, in 132 fms, sandy
green mud. 1.57 cu. ft.
not yet analyzed
Station 2389-53. Off Ship Rock, Catalina Island light, in 136 fms,
sandy mud. 2.2 cu. ft.
echinoderms, including many ophiuroids, a very large urchin,
holothurians
mollusks, including pelecypods, Dentalium and Cadulus
burrowing anemone—1
phoxocephalid amphipods, 6, including Heterophoxus sp. A—5
T halassema (echiuroid )—1 large
sipunculid—1
polyclad—1
parasitic copepod—1
polychaetes, including:
ampharetids—several juvenile
A naitides sp.—1
Aricidea sp.—several
Chone sp.—1
cirratulids—many
Cossura sp.—2
Drilonereis sp.—1 or more
Eunice aphroditois—1| large
Glycera sp.—2
goniadid—1
Haploscoloplos elongatus—8 juveniles
Lumbrineris sp.—several
Maldane sp.—several, in tubes
Melinna sp.—many, in tubes
Myriochele sp.—6
Nephtys spp.—few, juveniles
nereid—1
Nothria sp.—1
Paraonis sp.—several

No. l

162.

163.

164a.

HARTMAN: QUANTITATIVE SURVEY 141

Pectinaria californiensis—about 6
Pholoé sp.—many
Pilargis sp.—1
polynoid—1
Prionospio, near malmgreni—several
Prionospio pinnata—several
Sternaspis sp.—1
Terebellides sp.—1
terebellids—several, juveniles
Station 2328-53. Off Ship Rock, Catalina Island, in 150 fms,
fine sandy mud. 1.26 cu. ft.
not yet analyzed
Station 2329-53. Off Ship Rock, Catalina Island, in 260 fms, fine
greenish mud. 2.56 cu. ft.
not yet analyzed
Station 2176-52. Off Salta Verde Point, Catalina Island, in 28
fms, nodular muddy sand. 1.32 cu. ft.
holothurian
anemone—7
mollusks, including 2 Crepidula; a gastropod, Aglaja sp.; a
Chaetoderma; and a nudibranch
crustaceans, including many amphipods
nemerteans—few
sipunculid—1
Glottidia albida—4
branching bryozoans
simple ascidian—3 or more
polychaetes, including:
A glaophamus dicirris—1
A maea occidentalis—1
A mpharete arctica—1
A mphisamytha bioculata—1
A phrodita armifera—1|
Aricidea spp. — several
Artacaminae, new genus and species—2
capitellid—several
Chaetopterus sp.—tubes and fragments
cirratulids—more than 29
Diopatra tridentata—1 or more
Dorvillea sp.—2

1 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Drilonereis spp.—2 or more
Euchone sp.—1 and tube
Eulalia sp.—3

Exogone sp.—l1, ovigerous
Glycera sp.—1

Goniada sp.—3
Haploscoloplos elongatus—9
?H y poeulalia bilineata—3

Lanice sp.—several
Lumbrineris spp.—many
Magelona spp.—2 large
maldanids, several species—many
Nephtys californiensis—1 large
another Nephtys sp.—3
onuphid, in tube
Owenia sp.—about 5
Pherusa sp.—5
Poecilochaetus sp.—6
polydorid—several, with tubes
Prionospio, near malmgreni—18
Prionospio pinnata—3
Pseudopotamilla sp.—1, juvenile
Rhamphobrachium sp.—1 or more
Rhodine sp.—1 or more
Sabella, resembling crassicornis—2
sabellid—2, with tube
Scalibregma sp.—10
serpulid tube fragments
sigalionids—several
Spiochaetopterus sp.—several, with tubes
Spiophanes missionensis—many
Sternaspis sp.—about 10
Sthenelanella uniformis—hundreds
Streblosoma, possibly crassibranchia—many
Terebellides sp.—1
Tharyx, possibly parvus—27
Travisia sp.—6

164b. Station 2303-53. Off Ship Rock, Catalina Island, in 470 fms, in

fine greenish mud. 2.52 cu. ft.

no. l

165.

166.

167.

168.

169a.

169b.

170.

HARTMAN: QUANTITATIVE SURVEY 113

foraminiferans; tubes of Phyllochaetopterus and serpulid; dead
Pecten shells; a living tectibranch
phoxocephalid amphipod, Leptophoxus sp. A.—1
Station 2330-53. Off Ship Rock, Catalina Island, in 490 fms,
fine mud. 3.4 cu. ft.
foraminiferans; tubes of Phyllochaetopterus
Station 2453-53. Off Long Point, Catalina Island, in 480 fms,
very fine mud. 3.02 cu. ft.
foraminiferans; tubes of Phyllochaetopterus
Station 2331-53. Off Long Point, Catalina Island, in 489 fms, fine
mud. 2.7 cu. ft.
foraminiferans; tubes of Phyllochaetopterus
Station 2409-53. Off Long Point, Catalina Island, in 470 fms,
greenish mud. 3.4 cu. ft.
foraminiferans; tubes of Phyllochaetopterus and serpulid; dead
Pecten shells
Station 2305-53. Off Long Point, Catalina Island, in 460 fms,
fine greenish mud. 2.7 cu. ft.
foraminiferans; tubes of Phyllochaetopterus and serpulid; dead
Pecten shells; a small ophiuroid
Station 2626-54. Off Long Point, Catalina Island light, in 460
fms, gray-green mud. 5.75 cu. ft.
foraminiferans; tubes of Phyllochaetopterus
Station 2410-53. Off Long Point, Catalina Island, in 410 fms,
greenish mud. 3.2 cu. ft.
foraminiferans, | large and 6 small ophiuroids
mollusks, including 2 Chaetoderma, scaphopods, a gastropod
crustaceans, including an amphipod, a cumacean
phoxocephalid amphipod, Harpinia sp. B.—1
a nemertean
polychaetes, including:
A ncistrosyllis sp.—1
Aricidea sp.—1 or more
capitellid, new genus and species—1 or more
Chaetozone corona—fragments
other cirratulids—several
Cossura sp.—1 or more
Maldane sp.—2
orbiniid, new genus and species—2
Paraonis sp.—1 or more

114

near

ple

WZ;

ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Station 2411-53. Off Long Point, Catalina Island, in 400 fms,
greenish mud, glass sponge. 3.15 cu. ft.
echinoderms, including 3 brissopsid urchins and 4 ophiuroids
Chaetoderma—3 large and several small
phoxocephalid amphipod, Harpinia sp. B.—1
caprellid—1
enteropneust, Stereobalanus sp.—1
polychaetes, including:

A mphicteis 2scaphobranchiata—6

A ncistrosyllis sp.—2 or more

Aricidea sp.—4

capitellid, new genus and species—about 9

cirratulid—5

Cossura sp.—4

flabelligerid—fragments

Lumbrineris sp.—2

Maldane sp.—1

orbiniid, new genus and species—2

Petaloproctus sp.—3 or more

Protula sp.—several

sabellid, with parasite—1

Spiophanes sp.—3

other annelids
Station 2412-53. Off Long Point, Catalina Island light, in 324
fms, fine mud. 3.2 cu. ft.
4 brissopsid urchins
Chaetoderma—4
pelecypod—2
nemertean—1
polychaetes, including:

A mphicteis sp.—9

A ncistrosyllis sp.—2

Aricidea sp.—12

cirratulid—2

Euchone sp.—1, in mud tube

maldanid—about 3

Myriochele sp.—1

Nephtys sp.—1

orbiniid, new genus and species—1

No. 1 HARTMAN: QUANTITATIVE SURVEY 115

Pista sp.—1
Prionospio sp.—5
serpulid on worn Phyllochaetopterus tube—2
173. Station 2413-53. Off Long Point, Catalina Island light, in 205
fms, muddy clay. 0.5 cu. ft.
many ophiuroids, 2 echinoids
mollusks, including Dentalium, pelecypods, 3 chitons, a Chaeto-
derma
crustaceans, including a ghost shrimp, amphipods, isopods, cuma-
ceans, copepods
phoxocephalid amphipods, 14, including Harpinia sp. F.—1
Heterophoxus sp. A—6
Leptophoxus sp. A—3
Paraphoxus sp. A.—2
sipunculid—2
polyclad—1
polychaetes, including:
Ammotry pane sp.—3
ampharetid—2
A naitides sp.—1
Aricidea sp.—1
capitellid—3 or more
cirratulids—several
flabelligerid, new genus and species—about 20
Glycera sp.—4
Lagisca sp.—fragments
Laonice sp.—fragments
Nephtys sp.—several
Pectinaria sp.—1
Pherusa capulata—1
Polydora spp.—several, from drilled holes in shale
polynoid—1
Prionospio sp.—1 or more
Sphaerodorum sp.—1
?Sphaerosyllis sp.—1
Streblosoma sp.—2
Terebellides sp.—1
174. Station 2414-53. Off Long Point, Catalina Island light, in 177
fms, coarse sand and rocks. 0.4 cu. ft.
several ophiuroids

116 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

mollusks, including:
chitons—7 or more, from crevices in large rocks
Tellina sp.—1
Lima sp. or related genus—2 or more
Dentalium sp.—several
Chaetoderma—1
crustaceans, including a pagurid in dead conch; ostracods; isopods
phoxocephalid amphipods—14, including:
Har pinia sp. F.—2
Heterophoxus sp. A—2
Pontharpinia sp. F.—4
Ponthar pinia sp. J.—3
Pontharpinia sp. N.—2
sipunculids—2
polychaetes, including:
Ammotry pane sp.—3
Anaitides sp.—l1
Aricidea sp.—several
cirratulid—several
Glycera sp.—1
Lumbrineris sp.—3 or more
maldanid—several in tubes fully attached to under side of rocks
polynoids—several
Prionospio spp.—more than 2
sphaerodorid—2
syllid—1
terebellid—several
175. Station 2851-54. Off Long Point, Catalina Island light, in 230
fms, mud. 1.51 cu. ft.
foraminiferans; many diversified smaller animals; not yet analyzed
176 and 177 not yet sampled
178. Station 2841-54. Off Newport west jetty light, in 257 fms, green
mud. 2.93 cu. ft.
passed rapidly through screens, with little debris
a large sea pen; brissopsids ; many smaller animals; not yet analyzed
179. Station 2293-53. Off Newport west jetty light, in 252 fms, oozy
mud. 2.58 cu. ft.
ophiuroids and urchins
a sea pen and an anemone
mollusks, including pelecypods, gastropods, and scaphopods

no. 1

180.

181.

183.

184a.

HARTMAN: QUANTITATIVE SURVEY U7

echiuroids—3
crustaceans, including ostracods, isopods, cumaceans, amphipods
phoxocephalid amphipods, including Harpinia sp. A—2
Paraphoxus sp. A—7
Phoxocephalus sp. A—8
other animals and annelids, not yet analyzed
Station 2757-54. Off Dana Point, in 258 fms, greenish mud. 2.64
cu. ft.
2 large brissopsids; an echiuroid; many small animals, not yet
analyzed
Station 2756-54. Off Dana Point, in 285 fms, greenish mud and
coarse shelly sand. 2.7 cu. ft.
gastropods; polychaetes, including 4 phrodita, sabellid, and many
others, not yet analyzed
Station 2633-54. Off end of Newport Beach pier, in 292 fms, gray-
green clay. 6.03 cu. ft.
foraminiferans; a large brissopsid ; annelids, including ampharetids,
Pectinaria, and others; many small animals, not yet analyzed
Station 2449-53. Off Newport jetty light, in 303 fms, sticky mud.
3.02 cu. ft.
not yet analyzed
Station 2117-52. Off Abalone Point, Laguna Beach, in 54 fms,
mud. 3-65.cu. ft:
many foraminiferans
burrowing anemones—several
echinoderms, including ophiuroids—several hundred ; holothurians,
including 9 Molpadia intermedia
mollusks, including pelecypods, gastropods, scaphopods
crustaceans, especially amphipods, isopods, cumaceans
nemerteans, sipunculids
Glottidia albida—1|
polychaetes, including:
Ammotry pane sp.—1|
ampharetids—5 or more
Aricidea sp.—5 or more
Asychis or Maldane sp.—1
Chloeia sp.—2
cirratulid—1
Glycera sp.—several
Glycinde sp.—1

118

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

other goniadid—several
Lumbrineris spp.—many
Magelona sp.—1
maldanids—many
Megalomma sp.—2
Myriochele sp.—about 15
nephtyid—several

Panthalis pacifica—about 5, in a clump of tubes
Paraonis sp.—3

Pectinaria calif orniensis—15
Pholoé sp.—fragment
Polydora sp.—1

Prionospio spp.—several
Rhodine sp.—1

Spiophanes sp.—1

Sternas pis sp.—about 20
Sthenelais sp.—1
Terebellides sp.—2
terebellids—several

184b. Station 2751-54. Off Dana Point, in 200 fms, greenish-gray mud.

185.

2.95 Cue ite
many thick mud-walled tubes of Nothria pallida and Maldane sp.;
brissopsids ; smaller mollusks, not yet analyzed
Station 2448-53. Off Dana Point, in 30 fms, compact sticky mud.
1.95 cu. ft.
ophiuroids—hundreds
caudate holothuroid, perhaps M olpadia—4
mollusks, including a Chaetoderma and scaphopods
crustaceans, including many amphipods, a pinnotherid and a ju-
venile brachyuran
phoxocephalid amphipods, 34, including Heterophoxus sp. A—7
Ponthar pinia sp. E—8
Ponthar pinia sp. J —8
nemerteans, sipunculid, a polyclad
Glottidia sp.—few
phoronids—in slender sand-covered tube
enteropneusts, Stereobalanus sp.—1
polychaetes, including: (individuals not counted )
Amaea sp.
ampharetids

NO.

HARTMAN: QUANTITATIVE SURVEY

Ancistrosyllis sp.
Aricidea sp.

Brada sp.
Ceratocephala crosslandi americana
Cossura sp.
Glycera sp.
goniadid
Harmothoé sp.
Hyalinoecia sp.
Laonice sp.
Leocrates sp.
Lumbrineris spp.
Magelona sp.
Maldane sp.

other maldanids
Myriochele sp.
nephtyids—several species
Pectinaria sp.
Pholoé sp.

Pista sp.
Poecilochaetus sp.
Prionospio pinnata
Rhodine sp.
sabellid

sigalionid
Sphaerodorum sp.
Spiophanes sp.
Sternas pis sp.

T erebellides sp.
terebellids

119

186a. Station 2142-52. Off Howlands Landing, Catalina Island, in 19

fms, fine sandy mud. 1.9 cu. ft.
Dominant and conspicuous were the large tubes of Chaetopterus

variopedatus, that measured to 5 inches long; and the annulated,
strawlike tubes of Spiochaetopterus, 10 to 12 inches long (see

photograph of bottom, fig. 1)

echinoderms, including many ophiuroids and 7 holothuroids
poriferans—several
anemones—many tube dwelling
sea whips—5

120 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

mollusks, including:

gastropods—C onus californicus—5
Crepidula nivea—3
Kellettia kelletti—1

pelecypods—Botulina denticulata—1
Cardium sp.—1, juvenile
Crenella decussata—1
Gastropteron sp.—1
Lima dehiscens—6
Lyonsia californica—2
Saxicava arctica—|
Semele pulchra—1
Sportella californica—|
V olsella capax—\|

crustaceans, including amphipods, isopods, several shrimps, a crab,
a pycnogonid ; phoxocephalid amphipod, Heterophoxus sp. A.—1
nemerteans—4
sipunculids—12
many bryozoans
Glottidia albida—1
solitary ascidians—7
enteropneusts: Saccoglossus sp.—1 ; ptychoderid—2 ; other—5
polychaetes, including:
A glaophamus dicirris—14
A naitides sp.—1
Chaetopterus variopedatus—11 or more
Chaetozone corona—3
cirratulids—about 6
Cirriformia sp.—several
Eulalia sp.—2
Eumida sp.—1
Lumbrineris sp.—2
Owenia sp—about 60
Paraonis sp.—2
Pherusa sp.—2
Phyllochaetopterus ?prolifica—|
Polycirrus sp.—2
polynoid—2
Psammolyce sp.—3
Pseudopotamilla sp.—1

NO.

HARTMAN: QUANTITATIVE SURVEY 11

Scalibregma sp.—5
?Semiodera sp.—11
Spiochaetopterus sp.—hundreds, with tubes
spionid—1

Sternaspis sp.—1

Sthenelais sp.—several
Sthenelanella uniformis—6
?Streblosoma sp.—fragments
terebellid—1

Trypanosyllis sp.—1

V ermiliopsis sp.—1

186b. Station 2143-52. Off Howlands Landing, Catalina Island, in 25

near

186.

187.

near

187.

near

187.

fms, mud, broken shells and much rubble. 2.0 cu. ft.

mollusks, including gastropods, Conus californicus—3; pelecypods
—Trachycardium quadragenarium—1

many other animals, not yet analyzed

Station 2797-54. Off Ship Rock light, Catalina Island, in 63 fms,

coarse yellow-gray sand with shell and nullipore rubble. 0.12 cu. ft.

ophiuroids; a large urchin; nemerteans; many annelids, including
considerable Ch/oeia sp. and others, not yet analyzed

Station 2302-53. Off Ship Rock, Catalina Island, in 185 fms, fine

greenish mud. 1.57 cu. ft.

echinoderms, including 4 brissopsids, numerous ophiuroids, 3 holo-
thurians

anemone—|

mollusks, including pelecypods, gastropods, scaphopods, Chaeto-
derma

echiuroids—2 ; several nemerteans

crustaceans, including amphipods and isopods; phoxocephalid am-
phipods, Harpinia sp. G.—2; Heterophoxus sp. A—1

numerous annelids, not yet analyzed

Station 2733-54. Off Ship Rock, Catalina Island, in 152 fms, green

sandy mud. 1.13 cu. ft.

a large and a small brissopsid; many ophiuroids; an echiuroid;
many annelids, not yet analyzed

Station 2734-54. Off Ship Rock, Catalina Island, in 154 fms, green

sandy mud. An unscreened, unmeasured quart examined for smaller

122

188.

189:

190.

LO

LOZ.

193:

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

forms that might escape finest screens in field sorting; many smaller
annelids washed out, not yet analyzed
Station 2435-53. Off Ship Rock, Catalina Island, in 244 fms,
clayey mud and sand. 1.95 cu. ft.
not yet analyzed
Station 2301-53. Off Ship Rock, Catalina Island, in 335 fms, oozy
mud. 9.55 Cu. tt.
many foraminiferans
a large echinoid
anemone—2
mollusks, including a pelecypod, 3 scaphopods, 2 Chaetoderma
echiuroid—1
sipunculid—3
nemerteans—several
enteropneust—1
polychaetes, including:
ampharetids—many
Aricidea sp.—6
capitellid—3
Cossura sp.—1
maldanids—several
M yriochele sp.—5
orbiniid, new genus and species—1
Paraonis sp.—3
Protula sp.—1 or more
sabellid—1 or more
Station 2223-53. Off Long Point, Catalina Island, in 480 fms, oozy
mud. 3.1 5\ieus ft:
foraminiferans; tubes of Phyllochaetopterus and serpulid; a small
pelecypod and a Chaetoderma
Station 2408-53. Off Long Point, Catalina Island, in 480 fms,
sandy mud. 3.21 cu. ft.
foraminiferans; tubes of Phyllochaetopterus and serpulid, with liv-
ing specimens
Station 2801-54. Off Long Point, Catalina Island light, in 474
fms, blue-green-gray mud and some glass sponge. 2.64 cu. ft.
foraminiferans; Phyllochaetopterus and a few other animals, not
yet analyzed
Station 2353-53. Off Long Point, Catalina Island, in 430 fms, fine

mud, a rock, some wood fragments, many foraminiferans, minute

NO.

194,

196.

HARTMAN: QUANTITATIVE SURVEY 125

shells, and radiolarians. 3.21 cu. ft.
sea whip—1
pelecypods—2
polychaetes, including:
Amphicteis scaphobranchiata—several
Cossura sp.—l or more
Glycera sp.—several juvenile
Laonice sp.—1
Maldane sp.—1
Pista sp.—1 large, in thick mud-walled tube
tubes of Protula sp.
Spiophanes sp.—1
terebellid fragments

Station 2802-54. Off Long Point, Catalina Island light, in 420

fms, blue-green-gray mud. 2.64 cu. ft.

foraminiferans; glass sponge; a large sea star; scaphopods; gastro-
pod—Nitidella sp. ; large mud-covered tubes of annelids; Protula
sp.; and other animals

Station 2429-53. Off Long Point, Catalina Island light, in 380
fms, fine dark mud. 3.15 cu. ft.
glass sponge and foraminiferans
ophiuroids—about 10
holothuroid—1
sea pen, with broad pannicles—1
mollusks, including a small scaphopod and 2 Chaetoderma
nemerteans—2Z anterior ends
polychaetes, including:
ampharetid—1
A phrodita sp.—1 juvenile
cirratulid—many, some ovigerous, with flesh-pink ova
Maldane sp.—several, including large specimens
Mesochaetopterus sp.tubes
M yriochele sp.—several tubes
Scalibregma sp.—1 large

Station 2428-53. Off Long Point, Catalina Island light, in 350
fms, fine dark mud, with many spherical foraminiferans and a mass
of felt and setae of a large aphroditid. 3.41 cu. ft.

echinoderms, including 3 brissopsids and one ophiuroid
nemertean—1 or more

124

197.
198.

199:

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

polychaetes, including:
ampharetid—about 20, ovigerous
capitellid—2 or more
?Chone sp.—1
Maldane sp.—1
Myriochele sp.—1 or more
orbiniid, new genus and species—2
Pherusa sp.—2
Pilargis n. sp.—1
serpulid, in tube fully attached to tube of Mesochaetopterus
Spiophanes sp.—2
and other species
not yet sampled
Station 2852-54. Off east end, Catalina Island light, in 280 fms,
mude2.92 cu. tt.
foraminiferans; sipunculid; numerous smaller animals, not yet
analyzed
Station 2372-53. Off east end, Catalina Island light, in 230 fms,
sandy mud. 0.81 cu. ft.
not yet analyzed

200 and 201 not yet sampled

202:

203:

204.

205.

206.

Station 2842-54. Off Dana Point, in 288 fms, gray-green mud.

3.) Siew te:

a large dead conch; a large echiuroid; many smaller animals, not
yet analyzed

Station 2755-54. Off Dana Point, in 304 fms, green sticky mud.

SVicutte

foraminiferans; many smaller animals, not yet analyzed

Station 2754-54. Off Dana Point, in 312 fms, greenish-gray sticky

mud. 2.83 cu. ft.

foraminiferans; several long tubes, possibly annelids; a large ovig-
erous nereid; a sabellid; and many smaller animals, not yet
analyzed

Station 2753-54. Off Dana Point, in 322 fms, greenish-gray sandy

mud. 1.26 cu. ft.

foraminiferans; shell fragments; ophiuroids; amphipods; annelids;
not yet analyzed

Station 2752-54. Off Dana Point, in 330 fms, greenish-gray clay.

SOicusat:

foraminiferans, large and small animals, not yet analyzed

no. 1 HARTMAN! QUANTITATIVE SURVEY 125

207. Station 2452-53. Off Ship Rock, Catalina Island light, in 28 fms,
compact sandy clay; many disklike foraminiferans and arenaceous
Goésella flintii, retained in lowest screen. 0.37 cu. ft.
ophiuroids—many
holothurians—several
burrowing anemone
mollusks, including gastropods, pelecypods, Chaetoderma
crustaceans, including amphipods and decapods
nemertean
phoronid
Glottidia albida
solitary ascidian
enteropneust, ptychoderid—1
polychaetes, including: (individuals not counted )

A glaophamus dicirris

A maea occidentalis

ampharetids, several species

Aricidea sp.

capitellid

Chaetopterus sp.

cirratulids, more than one species

Diopatra sp.

Eumida sp.

Glycera sp.

Lanice sp.—in tubes covered with foraminiferan, Goéssela flintii

Lepidasthenia sp.

Lumbrineris spp.

maldanid

Megalomma sp.

Panthalis sp.

phyllodocid, associated with tube of Spiochaetopterus and
ascidian

Pista, resembling cristata

Pista 2elongata

Prionospio pinnata

Rhodine sp.

sabellid, in coarse, sandy chitinized tube

Spiochaetopterus sp.—many tubes and specimens, the dominant
species

Spiophanes sp.

126

208.

209a.

209b.

210.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Sternaspis sp.
2Sthenelais sp.
Sthenelanella sp.
Streblosoma sp.
syllid
Terebellides sp.
Thelepus sp.
Station 2451-53. Off Long Point, Catalina Island light, in 111
fms, compact sandy clay. 1.57 cu. ft.
not yet analyzed
Station 2224-53. Off Long Point, Catalina Island, in 200 fms, fine
dark green mud. 1.76 cu. ft.
echinoderms, including several ophiuroids and 8 echinoids
coelenterates, including one sea anemone and a dead solitary coral
mollusks, including gastropods, pelecypods, scaphopods, Chaeto-
derma
crustaceans, including 6 amphipods and an ostracod
phoxocephalid amphipods, including Harpinia sp. A—2
Heterophoxus sp. B.—1
nemertean—1
nematode—1
polychaetes, including:
A naitides sp.—1
cirratulid—1
Drilonereis sp.—fragment
Glycera sp.—1
Haploscoloplos elongatus—6
Lumbrineris cruzensis—1
Nephtys sp.—2
Nothria sp.—6
Notomastus sp.—1
Pectinaria calif orniensis—3
?Pilargis sp.—2 fragments
Station 2423-53. Off Long Point, Catalina Island light, in 175
fms, mud and shelly sand. 1.13 cu. ft.
many ophiuroids, several urchins; numerous pelecypod shells; many
smaller animals, not yet analyzed
Station 2365-53. Off Long Point, Catalina Island, in 300 fms,
sandy mud. 1.57 cu. ft.
not yet analyzed

211.

au2.

213.

HARTMAN: QUANTITATIVE SURVEY 127

Station 2837-54. Off Long Point light, Catalina Island, in 454
fms, greenish-gray sticky mud. 3.08 cu. ft.
glass sponge ; foraminiferans; an ophiuroid; a pelecypod; a Chaeto-

derma; compound ascidian on tubes of Protula; a sipunculid
polychaetes, including:

A ricidea sp.—3

Paraonis sp.—6

Phyllochaetopterus sp.—6 or more and tubes

polynoid—1

Protula sp.—tubes

Station 2839-54. Off Long Point light, Catalina Island, in 446
fms, greenish-gray sticky mud. 3.46 cu. ft.
foraminiferans; glass sponge; ophiuroids
mollusks, including:

Nitidella sp.—1

a small pelecypod

Chaetoderma—12
enteropneust—Stereobalanus sp.—1
polychaetes, including:

Aricidea jeffreysi—2

Cossura sp.—1

Hydroides sp.—1

Maldane sp.—1

Paraonis sp.—8, including some ovigerous

sabellid—1

Scalibregma sp.—1 or more

Tharyx sp.—2 or more

Station 2229-53. Off Long Point, Catalina Island, in 440 fms,

sandy clay, gravel, and rubbly black groundy clumps to rocks. 0.63

cu. it.

many foraminiferans

echinoderms, including a large ophiuroid and several smaller ones,
and echinoids

mollusks, including pelecypods, gastropods, scaphopods, Chaeto-
derma

branching corals and hydroid

amphipods—phoxocephalid : Harpinia sp. B.—22

a nemertean and a sipunculid

nematodes—4 or more

128 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

polychaetes, including:
Ammotry pane sp.—1
ampharetids—2 or more
A mphicteis scaphobranchiata—1, the largest annelid in the sample
Aricidea sp.—some
chaetopterid fragments
Cossura sp.—3
Drilonereis sp.—2
Glycera branchiopoda—2
Goniada sp.—2
Harmothoé sp.—1
Myriochele sp.—6
Myxicola sp.—1
Naineris sp.—3
Spiophanes sp.—2
Terebellides sp.—1
T haryx sp.—several

214. Station 2838-54. Off Long Point light, Catalina Island, in 394
fms, greenish-gray sticky mud and considerable gravel. 2.7 cu. ft.
many foraminiferans; radiolarians; glass sponge
mollusks, including Nitidella sp. and Chaetoderma
phoxocephalid amphipods; several cumaceans; an isopod with very

large chelae
polychaetes, including:
Aricidea sp.—several
capitellid, new genus and species
Chloeia sp.—1 large
Cossura sp.—3 or more
sabellid—1
Tharyx sp.—many, some with long setae and large ova

215. Station 2441-53. Off Long Point, Catalina Island light, in 340
fms, mud and clay. 2.64 cu. ft.
otoliths, many foraminiferans, some glass sponge
ophiuroids—few
mollusks, including a gastropod shell, 3 scaphopods, 3 Chaetoderma
crustaceans, including 2 amphipods and a caprellid
polychaetes, including: (single or few individuals of each)

ampharetid
Chaetozone sp.
Glycera sp.

No. 1 HARTMAN: QUANTITATIVE SURVEY 129

Maldane sp. or perhaps ?Praxillella sp.
Myriochele sp.
orbiniid, new genus and species—2
216 and 217 not yet sampled
218. Station 2371-53. Off east end, Catalina Island light, in 350 fms,
fine sandy mud. 2.43 cu. ft.
not yet analyzed
219 not yet sampled
220. Station 2843-54. Northern base of Lasuen Seamount, in 230 fms,
gray-green sandy mud and trace of rubble. 1.32 cu. ft.
brissopsids—7 or more; a large nemertean; pelecypods; many anne-
lids and other smaller animals, not yet analyzed
221 not yet sampled
222. Station 2644-54. Off Newport Beach pier, in 310 fms, greenish-
gray clay. 5.74 cu. ft.
foraminiferans; gastropods; smaller animals, not yet analyzed
223. Station 2634-54. Off Dana Point, in 320 fms, gray-green clay.
5.74 cu. ft.
foraminiferans; many larger and smaller animals, not yet analyzed
224a. Station 2120-52. Off Long Point, Catalina Island, in 44 fms, sandy
mud. Two very incompletely filled grabs together totalled 1.07
cu. ft. (This bottom was photographed, see fig. 2.)
echinoderms, including ophiuroids, holothurians, and echinoid
(Lytechinus )
burrowing anemones
mollusks, including pelecypods, gastropods, scaphopods, and Chaeto-
derma
crustaceans, including amphipods, ostracods, isopods, and cumaceans
nemerteans; sipunculid ; Glottidia
enteropneust—1
polychaetes, including:
Ammotrypane sp.—1
Aricidea spp.—many
Chaetozone sp.—several
Chloeia sp.—about 20 juveniles
Chone sp.—1
cirratulids—many
Drilonereis sp.—2
goniadid—1
Haploscoloplos elongatus—6

130 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Harmothoé sp.—fragments
Lumbrineris sp.—2

Magelona sp.—several
maldanids—several

Myriochele sp.—many

Nephtys spp.—many

Nothria sp.—1

Onuphis parva—1|2 or more
Panthalis sp.—2 or more

Pectinaria sp.—1

Pholoé sp.—several

Prionospio, near malmgreni—several
Prionospio pinnata—1

Spio, n. sp.—34 or more

Spiophanes sp. and other spionids—many
Sternaspis sp.—2

Sthenelais sp.—several

Sthenelanella sp.—several

T erebellides sp.—1 or more

T halenessa sp.—many

224b. Station 2144-52. Off Long Point, Catalina Island, in 45 fms, sandy
mud. 1.6 cu. ft. in two grabs
not yet analyzed

224c. Station 2145-52. Off Long Point, Catalina Island, in 44 fms, fine
mud. Volume not taken.
enteropneust, ptychoderid—1 ; other animals not yet analyzed

224d. Station 2152-52. Off Long Point, Catalina Island, in 19 fms, sandy
mud. 0.85 cu. ft.
holothurians—several
mollusks, including:
gastropods—A glaja sp.—1
pelecypods—Parvilucina tenuisculpta—9
Solen rosaceus—3
Sphenia fragilis—1
Tellina buttoni—18
Tellina car penteri—6
T hyasira barbarensis—1

crustaceans, including many amphipods, isopods, cumaceans
polyclad—1

no. l HARTMAN: QUANTITATIVE SURVEY 131

polychaetes, including:
A maea occidentalis—3
Ampharete sp.—5
A naitides sp.—4
A phrodita sp.—1
Aricidea sp.—32 or more
Armandia sp.—1
Artacaminae, new genus and species—3
Chaetozone sp.—12
Cossura sp.—6
Diopatra sp.—3
Euchone sp.—about 25
Fabricia sp.—several
Glycera sp.—1
Goniada sp.—1
Harmothoé sp.—8
Laonice sp.—4 or more
Lumbrineris spp.—43
Megalomma sp.—3
Nephtys sp.—10 or more
Nereis procera—1
Ninoé sp.—1
Odontosyllis sp.—31
Paraonis sp.—50 or more
Pectinaria californiensis—2
Pholoé sp.—1
Poecilochaetus johnsoni—4
Polycirrus sp.—1
Prionospio, near malmgreni—many
Prionospio pinnata—more than 42
sabellid—several, in tubes
Scalibregma sp.—5
Scoloplos sp.—3
Spiochaetopterus sp.—several
Spiophanes missionensis—hundreds
Sternaspis sp.—3 large and 10 small
Sthenelais spp.—several
Sthenelanella sp.—2
Tharyx parvus—more than 16, ovigerous
and other annelids

132 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

224e. Station 2153-52. Off Long Point, Catalina Island, in 45 fms,
broken shells and mud; 2 grabs, both very incomplete, were taken,
to total 1 cu. ft.
holothurians—many ; anemones and sea pen

mollusks, including:

gastropods—A glaja sp.—1
Cavolina tricuspida—1|
eolid—1
Haminoea virescens—2
Puncturella galeata—1

pelecypods—Nezocardium centifilosum—4
Nuculana hamata—1
Parvilucina tenuisculpta—5
Solen rosaceus—3
Sphenia fragilis—6
Tellina buttoni—26
Tellina carpenteri—18

numerous amphipods, including Photis californica, Ampelisca brevi-
similis, lysianassid, oedicerotid ; also phoxocephalids, including
Harpinia sp. D.—2
Heterophoxus sp. A—3
Metaphoxus sp. A—4
Ponthar pinia sp. M.—3
cumaceans, ostracods, and isopods of several species
Glottidia albida and phoronid
polyclad
sipunculid
8 larvae of a wrasse—Oxyjulis californica (identified by Miss Janet
Haig)
polychaetes, including: (individuals not yet counted )
ampharetids—many
A naitides sp.
arabellid, parasitic
?Armandia sp.
Chloeia sp.
Chone sp.
cirratulids, of more than one species
Drilonereis sp.
Euchone sp.

No. |

HARTMAN: QUANTITATIVE SURVEY

Eumida sp.

Eunice sp.

Fabricinae

Glycera sp.

goniadid

Harmothoé sp.
Lepidasthenia sp.
Longosoma catalinensis
Lumbrineris spp.
Magelona spp.
maldanids, of several species
Megalomma sp.
Myriochele sp.
Naineris sp.

nephtyid

nereid

Nothria sp.
Odontosyllis sp.
Onuphis sp.

Owenia sp.

Paraonis sp.
Pectinaria sp.
Peisidice sp.

Pherusa capulata
Pherusa sp.
phyllodocid

Pista elongata

Pista, other species
Polydora sp.
Prionospio spp.
Rhamphobrachium sp.
sabellids

Scalibregma sp.
Scoloplos sp.

serpulid, with glassy tube
sigalionids
Spiochaetopterus sp.
Spiophanes sp.
spirorbids—many
Sternaspis sp.

133

134 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Syllis sp.

terebellids—many

T halenessa sp.

T helepus sp.

trichobranchiid, new genus and species
Vermiliopsis on dead shells of Laqueus

225a. Station 2227-53. Off Long Point, Catalina Island, in 128 fms, fine
dark green mud. 0.81 cu. ft.
ophiuroids
mollusks, including pelecypods, gastropods, scaphopods, Chaeto-
derma
amphipods, ostracods, isopods, cumaceans, one hermit crab
phoxocephalid amphipods Heterophoxus sp. A—2
Ponthar pinia sp. U.—14
polychaetes, including:
Ammotry pane sp.—10
ampharetids—12 juveniles
A mphicteis sp.—1
Aricidea sp.—numerous
capitellid—14
Chone sp.—3
cirratulid—8
Drilonereis sp.—3
Exogoninae—1
Glycera tesselata—4
Goniada sp.—2
Haploscoloplos elongatus—11
Laonice sp.—1
Lumbrineris cruzensis—13
Magelona sp.—8
Maldane sp.—1
maldanids—10
Melinna sp.—1 or more
nereid—1
Nothria ?stigmatis—15
Notomastus sp.—2
Onuphis sp.—5
onuphids in tubes—about 4
Paraonis sp.—several
Pectinaria californiensis—3

No. l

225b.

226.

227a.

227b.

HARTMAN: QUANTITATIVE SURVEY 135

?Phyllodoce ferruginea—1

Pista ?cristata—7 or more
Prionospio ?cirrifera—8
Prionospio pinnata—3
Scalibregma sp.—1

Scionella sp.—1

spiochaetopterid sp.—many tubes
T erebellides sp.—1

T halenessa spinosa—1

Travisia sp.—1

Station 2290-53. Off Long Point, Catalina Island, in 200 fms,

mud. Volume not taken.

The partially analyzed organisms include an ophiuroid, an anemone,
3 scaphopods resembling Cadulus, a deep water gastropod, a
Tellina

polychaetes, including:
capitellid—1
Chloeia sp.—2
Maldane sp.—1
Nephtys sp.—1
Nothria sp.—1
Pectinaria californiensis—1
and others

Station 2426—53. Off Long Point, Catalina Island, in 270 fms,

dark mud. 1.0 cu. ft.

not yet analyzed

Station 2352-53. Off Long Point, Catalina Island, in 420 fms, fine

mud. 2.5 cu. ft.

glass sponge and many foraminiferans

ophiuroids—3

Chaetoderma—1

pelecypod—1

scaphopod—1

sea pen—l1, with digitately arranged branches, red in life

polychaetes, including:

Amphicteis scaphobranchiata—1 large
T helepus sp.—1 large
serpulid tube, attached to tube of Thelepus

Station 2859-54. Off Long Point, Catalina Island, in 425 fms,

greenish-gray sandy mud. 2.58 cu. ft.

136

228.

229.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

foraminiferans; radiolarians; much glass sponge; shells of Pecten

ghost shrimp, Callianassa goniophthalma Rathbun—1 male (?)
(identified by Miss Janet Haig)

mollusks, including a pelecypod and a chiton

a nemertean; ?sipunculid

polychaetes, including tubes of Protula, ?Phyllochaetopterus, am-
pharetid, Aricidea sp., Paraonis sp., ?T haryx sp.

Station 2228-53. Off Long Point, Catalina Island, in 293 fms,

oozy mud. 0.56 cu. ft.

many foraminiferans, some glass sponge

ophiuroids

mollusks, including gastropods, pelecypods, scaphopods, Chaeto-
derma

phoxocephalid amphipods:
Heterophoxus sp. A.—1
Pontharpinia sp. U.—1
polychaetes, including:
A mage sp.—1
A mphicteis sp.—2
A naitides sp.—fragment
Aricidea sp.—several
Cossura sp.—1
Goniada sp.—fragment
Haploscoloplos elongatus—2 juveniles
Laonice sp.—2 or more
Lumbrineris sp.—2
Notomastus lobatus—4 (the largest of the polychaetes)
Streblosoma sp.—1
Terebellides sp.—1
fragments and juveniles of cirratulid, onuphid, and others

Station 2845-54. Off Long Point, Catalina Island, in 277 fms,
gray-green sandy mud. 1.25 cu. ft.
foraminiferans, a brissopsid, many annelids, not yet analyzed

230a. Station 2338-53. Off Long Point, Catalina Island, in 393 fms, fine

mud. 2.58 cu. ft.

glass sponge, many foraminiferans, long slender spines of echinoids
ophiuroid—1 large white

ghost shrimp, resembling Callianassa—1

Pecten shells, delicate in texture—several

NO.

HARTMAN: QUANTITATIVE SURVEY

polychaetes, including:
capitellid—1
Chloeia sp.—1 large, about 40 mm long
cirratulid—several
Glycera branchiopoda—1|
Lumbrineris sp.—posterior fragment
Paraonis sp.—1

137

230b. Station 2427-53. Off Long Point, Catalina Island, in 290 fms, fine

greenish mud. 2.26 cu. ft.
glass sponge, numerous animals, not yet analyzed

231 not yet sampled
232. Station 2370-53. Off east end, Catalina Island light, in 366 fms,

239.

234.

230%

fine mud. 2.83 cu. ft.

foraminiferans and other animals, not yet analyzed

Station 2844-54. Off east end, Catalina Island, at western end of
Lasuen Seamount, in 362 fms, sticky gray-green mud. 3.15 cu. ft.
foraminiferans, large ophiuroids, many small annelids, not yet

analyzed

Station 2299-53. Off east end, Catalina Island, near Lasuen Sea-

mount, in 360 fms, dark oozy mud. 3.08 cu. ft.

glass sponge, dead mollusk shells, foraminiferans

echinoderms, including 1 ophiuroid, 1 echinoid
Chaetoderma—1

pelecypod and gastropod shells

nemerteans, | sipunculid, 1 echiuroid
polychaetes, including:
Ancistrosyllis ?rigida—|
Aricidea sp.—3
capitellid—3
Glycera ?branchiopoda—1|
Maldane sp.—1| or more
oweniid—1
phyllodocid—1
?Scolelepis sp.—1
T halenessa spinosa—1

Station 2298-53. Off east end, Catalina Island light, near Lasuen

Seamount, in 68 fms, shelly sand and a few rocks. 0.37 cu. ft.

echinoderms, including ophiuroids and echinoids
mollusks, including pelecypods and gastropods
an anemone, nemerteans, nematodes, sipunculids

138

236.

231.

238a.

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

crustaceans, including amphipods, isopods, and a copepod
numerous polychaetes, not yet analyzed

Station 2297-53. Off Dana Point light, in 181 fms, large and
small hard rocks and mud; most of the sample failed to pass
through screens. 0.68 cu. ft.

ophiuroids—several
echiuroids—some
sipunculid—more than one
polychaetes, including:
A naitides sp.—several
Boccardia sp.—fragment
capitellid—1
Dodecaceria sp.—1
Eunice sp.—1
Glycera sp.—1
Lepidonotus sp.—1
maldanid—1 or more
Polycirrus sp. —1
syllid—several
and other annelids

Station 2296-53. Off Dana Point light, in 268 fms, dark grayish-
green mud. 1.82 cu. ft.
ophiuroids, annelids, and other animals; not yet analyzed

Station 2121-52. Off Long Point light, Catalina Island, in 32 fms,
black sandy mud, many dead Lagueus and other shells, much
debris. 1.26 cu. ft., in two grabs.
a large white sponge; an urchin
numerous diversified mollusks, including Chaetoderma—2
Laqueus californicus—many shells
T erebratalia transversa—dead shells
crustaceans, including several small crabs, about 8 shrimps, isopods,
amphipods
phoxocephalid amphipods Heterophoxus sp. A—1
Metaphoxus sp. A—3
a small polyclad
polychaetes, including:
A naitides sp.—1
cirratulids—several
disomid—1

No. l

238b.

HARTMAN: QUANTITATIVE SURVEY 139

Glycera sp.—1

Goniada sp.—1

maldanids—numerous

Owenia sp.—2 or more

Pherusa spp.—many

polynoids—several

Prionospio spp.—numerous

Scalibregma sp.—1

Spiochaetopterus sp.—many

spionids

Sternaspis sp.—1

Vermiliopsis sp.—hundreds, attached to dead Laqueus shells
Station. 2638-54. Off Long Point light, Catalina Island, in 40
fms, gray sandy mud. 0.93 cu. ft. (orange-peel grab)
many ophiuroids; a large urchin; many annelids and other animals,

not yet analyzed

238c. Station 2637-54. Off Long Point light, Catalina Island, in 40 fms,

209.

240.

24a.

241b.

241c.

gray sandy mud. 1.14 cu. ft. (Campbell grab)

many ophiuroids; a large caudate holothurian; many annelids and
other animals, not yet analyzed

Station 2639-54. Off Long Point light, Catalina Island, in 82

fms, gray-green sandy mud. 1.14 cu. ft.

many ophiuroids; Chloeia and other annelids; many other animals;
not yet analyzed

Station 2367-53. Off Long Point, Catalina Island, in 230 fms,

gray-green sandy mud. 0.5 cu. ft.

not yet analyzed

Station 2350-53. Off Long Point, Catalina Island, in 350 fms,

rocks, mud, sand. 0.1 cu. ft.

many ophiuroids, annelids, and other animals; not yet analyzed

Station 2640-54. Off Long Point, Catalina Island, in 370 fms,

rock and muddy gravel. 1 quart.

foraminiferans and much rubbly material

2 male galatheid crabs, Munida quadrispina Benedict (identified
by Miss Janet Haig)

numerous annelids, not yet analyzed

Station 2641-54. Off Long Point, Catalina Island, in 373 fms,

green brown sandy mud. 3.01 cu. ft.

a large ophiuroid; 5 brissopsids; Chloeia and other annelids, not
yet analyzed

140 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

242. Station 2368-53. Off east end, Catalina Island, in 385 fms, sandy
mud, rock. 2.26 cu. ft.
foraminiferans and diversified animals; not yet analyzed

243. Station 2369-53. Off east end, Catalina Island, in 390 fms, fine
mud. 3.33 cu. ft.
not yet analyzed

244. Station 2440-53. Off east end, Catalina Island, in 415 fms, clay
and mud. 3.27 cu. ft.
not yet analyzed

245. Station 2339-53. Off east end, Catalina Island, in 394 fms, fine
muds2-2 cul ft:
not yet analyzed

246 and 247 not yet sampled

248. Station 2643-54. Off east end, Catalina Island, in 382 fms, green-
ish-gray rubbly clay. 6.03 cu. ft.
foraminiferans; glass sponge; a large sea star; a large ophiuroid;

Chaetoderma; an amphipod ; an ostracod
polychaetes, including:

Aricidea sp.

cirratulids, including T’haryx sp.

Glycera branchiopoda

Maldane sp.—a large one
Myriochele sp.
tubes of Phyllochaetopterus sp. and Protula sp.
Rhodine sp.

249. Station 2635-54. Off Dana Point, in 278 fms, green clay and
mud, yellow and black marbled clayey mud with small rocks, much
rubble. 2.99 cu. ft.
foraminiferans; ophiuroids; a brissopsid; ?burrowing anemone
mollusks, including Cadulus and Chaetoderma
a shrimp and several amphipods
an echiuroid and a nemertean
polychaetes, including:

ampharetids

Aricidea sp.

Brada sp.

capitellid

Ceratocephala crosslandi americana
Chloeia sp.

Glycera sp.

No. |

250a.

250b.

Zo,

22:

HARTMAN: QUANTITATIVE SURVEY 141

Hydroides sp.

maldanids

nephtyid

orbiniid, new genus and species

tubes of Poecilochaetus sp. or Rhodine sp.

polynoid

spionid

and others
Station 2122-52. Off Jewfish Point, Catalina Island, in 48 fms,
sandy mud, dead brachiopod shells. 0.95 cu. ft. in two grabs.
many ophiuroids; crustaceans, including hundreds of ostracods,
many amphipods, isopods, cumaceans; a pelecypod; tubicolous
anemones
polychaetes, including:

Chaetozone sp.

Chloeia sp.

Lepidasthenia virens

Magelona sp.

maldanid

Myriochele sp.

Nephtys sp.

Prionospio sp.

Sternaspis sp.

Sthenelanella uniformis

Thalenessa spinosa

Tharyx sp.

Thelepus sp.

and others
Station 2436-53. Off Long Point, Catalina Island light, in 44
fms, oily sandy mud and clay. 1.07 cu. ft.
phoxocephalid amphipods, 34, including Harpinia sp. D.—27;

Metaphoxus sp. A—1
other animals not yet analyzed
Station 2347-53. Off east end, Catalina Island light, in 100 fms,
sandy mud. 0.25 cu. ft.
many ophiuroids; a brissopsid ; numerous mollusks; amphipods
a large polyodontid tube, Chloeia sp., and other polychaetes, not

yet analyzed
Station 2344-53. Off east end, Catalina Island light, in 210 fms,
sandy mud, gravelly rocks, rubble. 0.37 cu. ft.

142

253.

254.

299%

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

foraminiferans ; a brissopsid and 2 or more ophiuroids
mollusks, including Te/lina sp. and others
hermit crab in conch; many ostracods
phoxocephalid amphipods—5, including:
Harpinia sp. G.—1
Heterophoxus sp. B.—1
burrowing anemone—3
nemertean—1 or more
polychaetes, including:
Ammotry pane sp.—2
cirratulids
Drilonereis sp.—1
Maldane sp.—several
Onuphis sp.—several
Prionospio pinnata—several
Rhodine sp.—several
and others
Station 2343-53. Off east end, Catalina Island light, in 418 fms,
fine sandy mud. 1.63 cu. ft.
foraminiferans; long spines of urchins; a holothuroid; ophiuroids,
brissopsid ; mollusks, including Jel/ina and others; several am-
phipods, an isopod; polychaetes, including new orbiniid and
others, not yet analyzed
Station 2642-54. Off east end, Catalina Island light, in 422 fms,
greenish-gray mud. 6.17 cu. ft.
foraminiferans; chaetopterid tubes; many smaller animals, not yet
analyzed
Station 2340-53. Off east end, Catalina Island, in 410 fms, fine
mud. 2.26 cu. ft.

not yet analyzed

256 not yet sampled

ZO:

Station 2636-54. Off east end, Catalina Island, in 412 fms, green-
ish-gray clay. 6.03 cu. ft.
richly diversified animals, not yet analyzed

258 not yet sampled

259.

260.

Station 2437-53. Off east end, Catalina Island light, in 48 fms,
fine sandy mud. 1.3 cu. ft.

not yet analyzed

Station 2348-53. Off east end, Catalina Island light, in 75 fms,
sandy mud, rocks, broken shell and rubble. 0.37 cu. ft.

NO.

261.

262.

263.

264.

HARTMAN: QUANTITATIVE SURVEY 143

many ophiuroids; 2 brissopsids; a holothurian
mollusks
1 anemone
4 phoxocephalid amphipods, including Pontharpinia sp. B.—3
polychaetes, including:
A naitides sp.—1
Chloeia sp.—1
Chone sp.—1
Glycera sp.—1
Lumbrineris sp.—1
Magelona sp.—2
Pholoé sp.—1
Prionospio sp.—1
and others
Station 2349-53. Off east end, Catalina Island light, in 110 fms,
sandy mud. 0.1 cu. ft.
ophiuroids, a brissopsid ; 6 phoxocephalid amphipods; many isopods,
some cumaceans; pelecypod mollusks; numerous smaller poly-
chaetes ; not yet analyzed
Station 2342-53. Off east end, Catalina Island light, in 230 fms,
fine sandy mud. 0.31 cu. ft.
not yet analyzed
Station 2341-53. Off east end, Catalina Island light, in 440 fms,
fine sandy mud. 0.1 cu. ft.
not yet analyzed
Station 2439-53. Off east end, Catalina Island light, in 435 fms,
clay and mud. 2.52 cu. ft.
not yet analyzed

265 and 266 not yet sampled
267a. Station 2438-53. Off east end, Catalina Island light, in 153 fms,

hard rocks with surface and crevice dwelling animals. 1 quart.

encrusting bryozoans; an amphipod; polychaetes, including a
sphaerodorid, some maldanids in encrusting tubes on rocks, a
polynoid

267b. Station 2177-52. Off east end, Catalina Island light, in 172 fms,

sandy mud. 0.63 cu. ft.
numerous ophiuroids, some urchins; nemerteans; polychaetes, in-
cluding Chloeia and others, not yet analyzed

144 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

SUMMARY OF RESULTS

The unscreened samples have shown that much of the bottom of San
Pedro Basin is bed rock overlain by silt, ooze, clay or fine detritus in a
layer that exceeds in thickness the depth of the grabbing devices. Most
of the samples have been of this kind. Some have come up with varying
amounts of rock, gravel, or rubbly materials. Such are samples from the
vicinities of submerged mounts, on either side of the northwestern and
southeastern threshholds (see chart 2, and sample 235, above). Large
rocks come from parts of Redondo Canyon (sample 12b), from a canyon
southwest of Newport Bay (12la), and from steep slopes off the eastern
end of Catalina Island (241, 252, and 267). Pleistocene gravel comes
from a canyon wall (at 125) and blackened phosphorite rocks from
sample 52.

It is noteworthy that samples taken with uniform methods, using the
orange-peel grab, have been very irregular in volumetric amounts, rang-
ing from as much as 3.71 cu. ft. in a sample (39), to as little as a quart
(0.033 cu. ft.) in another (267a). These differences are explained in
several ways. For 252, 262, and 263, the grab closed before the bottom
had been penetrated, perhaps through an oversensitivity on the part of the
trigger, or from striking a swimming object, such as a large fish. For
some samples (45b, 241b, and 267a), the jaws were incompletely closed
on ascent, due to the presence of rocks and debris. For others (65, 66a,
and others) the hard packed bottoms of sand, clay or other materials,
could not be penetrated by the device used. Furthermore, it was not
possible to measure the amounts of the sample which might have filtered
out on hauling up the grab, or to retrieve the solid particles, including
animals, which might have escaped through apertures, such as vents for
releasing water pressures. In spite of unmeasurable losses of varying
amounts, the quantities and diversities of the samples are conspicuous.

Most of the bottoms of San Pedro Basin have been found to support
a rich and varied fauna in shallow to great depths, except for a con-
spicuous impoverished area in the western end (see chart 2, and text,
below). The diversity is manifested not only in single samples, where the
numbers of species may run very high, but in samples from adjacent areas,
indicating a patchiness of occurrence which is unpredictable, and at
present unexplainable.

No. 1 HARTMAN: QUANTITATIVE SURVEY 145

Thus, sample 44b yielded more than 50 species of annelids, with more
than 850 individuals, in addition to many other invertebrates (see
Analyses, above). A nearby sample, 45a, contained several large jack-
knife clams, phoronids, ophiuroids, more than 43 species of annelids with
hundreds of individuals, few of which were common to the adjacent
sample, 44b. The Chaetopterus association, sample 43a, is quite different
from that in 44a, even though both come from shallow bottoms near
shore. Adjacent samples, 123a and 123b, about a minute of longitude
apart, differ considerably in the species comprising their populations (see
Analyses, above).

The effects of the breakwater along the outer side of the Los Angeles
Harbor, are well shown by comparison of samples 44b and 44a (see
Analyses, above). Sample 44a, beyond the breakwater, has many speci-
mens of a larger terebellid, Streblosoma sp., enteropneusts, ophiuroids
and holothuroids. Sample 44b, in the Outer Harbor behind the break-
water, contains many specimens of Nereis procera, Tharyx parvus, Cos-
sura sp., Amphictets scaphobranchiata, and other annelids.

Similar habitats, such as sandy bottoms along shore, differ in their
dominant animals, even though ecological conditions appear the same.
For example, sands off Anaheim Bay (sample 50) support an archian-
nelid, Saccocirrus, and a pelecypod, Crenella, in considerable numbers.
Sands off Los Angeles breakwater (sample 63b) have large numbers of
an anemone, Harenactis; those north of Palos Verdes Point (sample 14)
support large numbers of a sipunculid in dead tests of Dendraster, to-
gether with annelids of the genera Pisione and Aricidea.

Masses of biological detritus, such as the thick layers of dead lamp
shells, Laqueus californicus and Terebratalia transversa (from samples
186b, 238a and 224e), and the associations of glass sponge (see text,
below) have been found to contain a great variety of animals.

The number of specimens in some samples runs very high, with some
groups, but seldom single species, of animals dominant. In all soft bot-
toms (rocky bottoms have not been successfully sampled with the grabs),
the marine annelids are most abundant, followed by ophiuroids, pelec-
ypods, holothurians, gastropods, amphipods, scaphopods, burrowing anem-
ones, echiuroids, nemerteans and other invertebrates. Thus, sample
108, off Newport Beach pier, in 17 fms, measuring 1.26 cu. ft., yielded
63 species and more than 500 individuals of annelids, in addition to many
ophiuroids, holothurians, pelecypods, gastropods, amphipods, and other
groups. A sample from Redondo Canyon (6b) in 161 fms, containing
2.8 cu. ft. of mud, had 6 large specimens of an undescribed species of

146 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Thalassema (communication from the late Dr. W. K. Fisher), the
largest measuring 140 mm long, and more than 50 specimens of Chloeia
pinnata, together with many other animals.

Another sample from Redondo Canyon (72) from 40 fms, with 3
cu. ft. of mud, yielded 142 living specimens of Dentalium rectius (see
list of mollusks by Dr. Norman T. Mattox, below), in addition to
hundreds of individuals of other species. Sample 12a, from 229 fms,
yielded 323 living individuals of Macoma incongrua, 42 of Nuculana
conceptionis, 61 of Tellina carpenteri (according to Dr. Mattox), more
than 90 of Chloeia pinnata, and many others (see Analyses, above).
Sample 69 contained more than 200 individuals of a sabellid, Chone
ecaudata, and 125 of a polynoid, Harmothoé triannulata.

The largest count that has been made of a metazoan species comes
from sample 42b, in 34 fms, 2.14 cu. ft. of mud, where a cirratulid
annelid, Tharyx parvus, is represented by more than 1530 individuals.
An interesting sample, 63b, from 11 fms, 0.81 cu. ft., of sandy mud,
contained hundreds of an anemone, Harenactis, in addition to many
other species of animals. Phoxocephalid amphipods have been counted,
numbering to 126 individuals in six species from one sample (133) ; 144
individuals in six species from another (104); and 93 specimens in four
species from a third (159a) (see list in Appendix, by Dr. J. Laurens
Barnard).

Based on analyses of the annelids, it is noteworthy that there are few
species in the benthos of San Pedro Basin which occur also in intertidal
zones of southern California. There are few representatives of nereids,
eunicids, serpulids, and some other large families. There are many genera
and species which are new records for southern California and also the
eastern Pacific (note especially genera and species in Aricidea, Artaca-
minae, capitellids, Cossura, flabelligerids, Hauchiella, Leocrates, Mage-
lona, maldanids, Myriochele, orbiniids, Paraonis, Petaloproctus, Pholoé,
Phyllochaetopterus, Pilargis, Pisione, Saccocirrus, and others). There are
relatively few cosmopolitan species. There are many species which appear
to have prolonged reproductive seasons, or to lack peaks of high repro-
ductivity.

More than 400 species of metazoan invertebrate animals have been
identified from the benthos of San Pedro Basin. The list will be con-
siderably enlarged as the various groups of animals are studied. The
annelids, with more than 283 species, have been examined by the writer.
Twenty-eight species of phoxocephalid amphipods have been identified by
Dr. J. Laurens Barnard; 80 species of mollusks have been examined by

No. 1 HARTMAN: QUANTITATIVE SURVEY 147

Dr. Norman T. Mattox; four species of enteropneusts were studied by
Keith Woodwick (see Appendices, below).

The technical details of the equipment are described by Dr. Floyd E.
Durham (see Appendices, below).

The Impoverished Area of San Pedro Basin

An analysis of the samples shows that there is an area in San Pedro
Basin that is characterized by an impoverished fauna (see chart 2) ; it
is an approximately lozenge-shaped tract measuring about 8 by 15 miles,
with a much smaller area to the north and west. Its outer margins follow
rather closely the 425 fathom contour line. Depths range from about 405
to 495 fms, or the deepest part of the Basin. The area is bounded by a
glass sponge association (see below), especially along the rims where the
current may be most rapid. The few animals existing in the impoverished
association are almost entirely a chaetopterid annelid, Phyllochaeopterus
sp., a serpulid, protulid, and less frequently an ampharetid and a glycerid,
Glycera branchiopoda. There are often delicate, translucent shells of a
Pecten, few of which have been found living; but many have been seen
that were unworn and not long dead.

The impoverished Phyllochaetopterus association in San Pedro Basin
comes from maximum depths in bottoms of very fine oozy mud. The
following lists the sample numbers, depth in fathoms, and volumetric
measurements of the samples obtained.

Volume of Sample,

Sample Number Depth in Fathoms in Cubic Feet
8 415 3.08
near 15 454 (volume not taken)
16 405 3.90
21 430 3.4
OH: 418 2.20
36 432 3:15

39 437 $1

148 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Volume of Sample,

Sample Number Depth in Fathoms in Cubic Feet
53 450 3.59
54 458 3415
bya) 457 2.26
56 460 2
57 431 2552
70 460 2.07
wil 466 3.08
58a 427 (uncertain )
72 480 Sel
73 475 3.40
WS 437 3.02
90 475 Posi fl
93 480 3.40
94 495 2.50
95 460 3.30

3 490 S.5
114 480 S215
5 470 Syils)
117 487 2252.
118 460 2.96
137 435 52:
138 482 3.10
139 484 Sie)
140 482 2.80
141 470 2.70
142a 490 3.46
142b 490 2.68
143 483 Sih
164b 470 252.
165 490 3.40
166 480 3.02
167 489 2.70
168 470 3.40
169a 460 2.70
190 480 S15

191 480 S7h

No. 1 HARTMAN: QUANTITATIVE SURVEY 149

The Glass Sponge Association

Another association that has been identified is characterized by the
presence of glass sponge. It comes only from deep water and is distributed
in its greatest diversity in the vicinities of low submarine mounts, and
alung the margins of the impoverished faunal area. Its geographic extent
is limited by depth, since it gives rise rather rapidly to a shallow water
association in other parts of the Basin. The dominant groups of animals
are listed below, based on samples taken from the stations. The following
gives the sample numbers, depth in fathoms and volumetric measure-

ments of the samples obtained.

Volume of Sample,

Sample Number Depth in Fathoms in Cubic Feet

9 300 2
10b 251 0.95

15 445 3.40

23 370 S71

69 400 3.08

76 420 2.58

89 386 2.96

96 440 S15

119 450 3:33
171 400 Sal
192 474 2.64
194 420 2.64
195 380 3.15
227a 420 2.50
230a 393 2.58
234 360 3.08
241c 373 3.01

248 382 6.03

150 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

List of Animals that occur in Glass Sponge Association

Echinoderms, including ophiuroids, brissopsid, echinoids

Mollusks, especially scaphopods, chaetodermans, gastropods of the genus
Nitidella, pelecypods, including translucent shells of a pecten

ghost shrimp, Callianassa goniophthalma

a broad-pannicled sea pen

amphipods and isopods

echiuroids and nemerteans

an enteropneust, Stereobalanus sp.

polychaetous annelids including:

A mphicteis scaphobranchiata
A ncistrosyllis rigida

A phrodita sp.

capitellid, new gen. and sp.
Chloeia pinnata

Cossura, n. sp.

other cirratulids

Eumida sp.

Glycera branchiopoda
Hydroides sp.

Laonice sp., or 2Spiophanes sp.
Lumbrineris spp.

Maldane sp.
Mesochaetopterus sp.
Myriochele sp.

orbiniid, new genus and species
paraonids

Petaloproctus sp.
Phyllochaetopterus sp.
polynoid, one or more species
Protula sp.

sabellid

Scalibregma sp.

Scolelepis sp.

Spiophanes sp.

T halenessa spinosa

Thelepus sp.

other annelids

NO.

HARTMAN: QUANTITATIVE SURVEY

APPENDICES A TO F

151

sy ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

ix

IMPROVED TECHNIQUES FOR OCEAN BOTTOM
SAMPLING

By Froyp E. DuRHAM

The obtaining of invertebrate specimens from the bottom of the San
Pedro Channel, California, by the Velero IV involves two operations:
(1) bringing to the surface bottom samples, and (2) separating the
specimens from the substrate.

The first operation is effected by a Hayward Standard orange-peel
bucket, with a rated capacity of two cubic feet. The area sampled is 2 3/5
square feet. The bucket is modified by a canvas hood over the open
structure, to prevent the washing out of the bottom sample as the bucket
is hauled up through the water. The hood has twelve four-inch by six-
inch oval vents, each bearing a flap on the outside, hinged at the top to
allow the escape of air and water as the bucket is lowered and to allow
the escape of water as the bucket is raised. Because of the hood, the
bucket will bring to the surface almost double its rated capacity of soft
mud. The bucket is further modified with a lowering pendant which
automatically trips the bucket when it strikes the bottom, and allows its
operation on the single 7/16-inch steel cable wound on a variable-speed
power winch mounted on the working deck.

At a selected spot, the open bucket is swung clear of the stern by the
pneumatically operated A-frame and lowered, with the winch in high
gear, at approximately forty fathoms per minute. A tachometer measures
in feet the cable paid out and indicates the depth from which the sample
is taken. Hauling on the cable closes the four jaws of the tripped bucket
and it is brought to the surface at the same speed as it descended. It is
swung onto the deck by the A-frame and the contents of the bucket are
dropped into a heavy cylindrical tub with a capacity of 4.46 cubic feet.
Particularly when sticky mud is being handled, it is necessary to do some
cleaning of the bucket by hand. Fragments remaining in the bucket are
washed out as the bucket descends for another grab.

No. 1 HARTMAN: QUANTITATIVE SURVEY 153

The tub with the bottom sample is then moved by the four strong
handles, and as many men, over to the screens. The surface of the sample
in the tub is smoothed, if necessary, and the volume is determined by a
calibrated measuring stick. Two small one-half - pint labelled samples
are removed for special studies in sedimentation and foraminifera.

The second operation, that of separating the biological specimens
from the substrate, is done in a mechanical shaker with a set of graded
screens. The process of softening and removing the wastes is accelerated
by a spray of salt water from a system of six sprinkler heads mounted
above the screens. The top screen, in which the bottom sample is placed
by hand-scoops, is of one-half inch diamond mesh, flat, galvanized steel.
The second screen, of brass, is twelve mesh per inch, and the third and
last screen, also of brass, is twenty-four mesh per inch. The latter deter-
mines the minimum size of specimens saved, except for those contained
in the two above-mentioned unsorted samples.

When all the substrate possible has been washed over the side of the
ship, the specimens, with the remaining debris, are collected in jars. The
larger specimens are removed from the screens by forceps, whereas smaller
ones are either gently swept to one corner of the screen with a stream of
water and picked up with a spatula, or the screen is inverted over a
framed, shallow, white, oilcloth tray. In the latter method, the specimens
are dislodged from the screen by tapping it gently and/or applying a
draft of air from the exhaust end of a vacuum cleaner head. The speci-
mens are then easily crowded to the center of the sagging cloth and
scooped up with a large spoon. Preservative, together with a label bear-
ing the station number, is added to each jar of specimens. All apparatus
is hosed clean prior to the collecting of the next bottom sample.

The screens are two feet by four feet and are framed with two inch
by one inch galvanized channel iron with welded corners. Supporting the
screening is one-half inch strap iron on edge every twelve inches. The
sieves are inserted into the carrier from the side, and held by a self-
locking hasp at each end of the carrier. Longitudinal motion within the
carrier is prevented by tightening a winged set screw at the end of each
screen. The carrier is well braced and is mounted on four small, steel
wheels on a short track, making possible a rapid to and fro motion, power
being supplied by a small electric motor directly through an adjustable
eccentric (7/16 inch stroke being satisfactory for the work). The four
screens and the carrier assembly are mounted on a steel stand supporting
the track for the carrier, the mounting for the electric motor, a rack for
specimen trays and tools, and supports for the sprinkler heads.

154 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

B.

A LIST OF THE MOLLUSCA IDENTIFIED
FROM SAMPLES OF THE BENTHOS OF
SAN PEDRO BASIN, CALIFORNIA

By NorMan T. Mattox

Number of Sample Depth in

Name of Species Specimens Number Fathoms
AMPHINEURA 1 45a 12
Aplacophora 1 59a 249
Chaetoderma sp. A 1 69 400
1 82 18
1 122 115
Chaetoderma sp. B 13 12a 228
2 69 400
2 82 18
Chaetoderma sp. C 1 59a 249
Limifossor sp. 9 12a 228
1 59a 249
3 i222 115
Neomeniinid sp. 1 69 400

Polyplacophora
Lepidizona catalinae Willett 4 224e 45
SCAPHOPODA

Cadulus fusiformis Pilsbry 7a 61
and Sharp 7b 40

MmDdOo—
aS
Gn
»
—
bo

No. 1 HARTMAN: QUANTITATIVE SURVEY 155

Number of Sample Depth in

Name of Species Specimens Number Fathoms
Cadulus tolmiei Dall 33 12a 228
1 80b 2S
Dentalium neohexagonum 1 45a 172
Pilsbry and Sharp 1 82 18
Dentalium rectius Carpenter 2 6c 129
142 7a 61
8 7b 40
3 12a 228
8 66a 2

GASTROPODA

Acteon punctocoelata 1 7a 61
(Carpenter) 1 7b 40
A glaja sp. i 45a 12
1 80a 29
Z 80b 23
1 82 18
1 164a 470
1 224d 19
1 224e 45
Balcis rutila (Carpenter) 6 12a 228
2 82 18
Bittium attenuatum Carpenter 1 12a 228
Bittium catalinensis Bartsch 1 7a 61
Cavolina tricuspida (Rivers) 1 224e 45
Conus californicus Hinds 5 186a 19
3 186b 25
Crepidula nivea C. B. Adams i) 45a 12
3 186a 19
Cylichnella diegensis (Dall) 1 7b 40
2 45a 12
1 82 18
Eolid 1 224e 45
Eulima californica (Bartsch) 2 82 18
Ferreria belcherii (Hinds) 1 45a 12
Fusinus arnoldi (Cossmann) 2 7a 61
Gastropteron sp. 1 186a 19
Haminoea virescens (Sowerby) 2 224e 45

156 ALLAN HANCOCK PACIFIC EXPEDITIONS

Number of
Name of Species Specimens
Kellettia kelletti Forbes 1
Leptogyra sp. 6
Nassarius perpinquis (Hinds) 2
Nitidella gouldi (Carpenter) 10
Nitidella permodesta Dall 2

O phiodermella incisa (Carpenter ) 2
Puncturella galeata (Gould) 1
Turbonilla sp. 1
V olvulella californica Dall 1
V olvulella tenussima Willett 6

3

PELECYPODA

A xionopsis sericatus Carpenter

—

Botulina denticulata (Dall)
Cardiomya pectinata (Carpenter)
Cardium sp. (juvenile)

Crenella columbiana Dall
Crenella decussata Montagu
Kellia sp.

Lima dehiscens Conrad
Lucinoma annulata (Reeve)

Lyonsta californica Conrad

W
Do
BR WN KK NO BR RK RB KK WD DY

Macoma incongrua (Martens)
Macoma yoldiformis Carpenter
Nemocardium centifilosum
(Carpenter)
Nucula cardara (Dall)
Nucula carlottensis Dall
Nuculana conceptionis (Dall)
Nuculana hamata (Carpenter )
Nuculana spargana (Dall)
Nuculana taphira (Dall)

aN
bo ke DO

BN CO Fe

Sample
Number

186a

69
45a

voL. 19

Depth in
Fathoms

no. | HARTMAN: QUANTITATIVE SURVEY 157

Number of Sample Depth in

Name of Species Specimens Number Fathoms
Pandora filosa Carpenter 2 7b 40
Parvilucina tenuisculpta 14 7a 61
(Carpenter ) US 7b 40
Z 82 18
9 224d 19
5 224e 45
Pitar newcombiana (Gabb) 3 7b 40
1 45a 12
Rochefortia tumida Carpenter 11 7b 40
3 82 18
Saxicava arctica (Linné) 1 186a 19
Semele pulchra (Sowerby) 1 186a 19
Solemya panamensis Dall 1 7a 61
Solemya volvulus Carpenter 8 82 18
Solen rosaceus Carpenter Z 45a 12
6 46a 13
3 224d 19
3 224e 45
Sphenia fragilis Carpenter 1 224d 19
6 224e 45
Sphenia globula Dall 5 12a 228
Sportella californica Dall 1 186a 19
Tellina bodegensis Hinds 1 7a 61
1 7b 40
Tellina buttoni Dall 1 82 18
18 224d 19
26 224e 45
Tellina carpenteri Dall 61 12a 228
6 224d 19
18 224e 45
Thyasira barbarensis Dall 3 7a 61
1 7b 40
1 12a 228
2 45a 12
1 224d 19
Trachycardium quadragenarium 1 46a 13
(Conrad ) 1 186b 25

158 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Number of Sample Depth in

Name of Species Specimens Number Fathoms
V olsella capax (Conrad) 1 186a 19
Y oldia scissurata Dall 3 6c 129
13 7a 61
19 7b 40

CEPHALOPODA
Octopus apollyon Berry 1 45a 12

No. 1 HARTMAN: QUANTITATIVE SURVEY 159

G.

A LIST OF PHOXOCEPHALID AMPHIPODA
IDENTIFIED FROM SAMPLES OF THE
BENTHOS OF SAN PEDRO BASIN,
CALIFORNIA

By J. Laurens BARNARD

Number of Sample Depth in

Name of Species Specimens Number Fathoms
Harpinia sp. A. 2 1 180
2 60 80
6 98 Z2N5
8 121b 230
Zz 179 252
2 209a 200
Harpinia sp. B. 1 10b 251
1 23 370
ik 121b 230
1 170 410
1 near 171 400
22 213 440

Har pinia sp. C. 2 49 1%
Har pinia sp. D. (2)3 146 300
Z 224e 45
27 250b ++
Harpinia sp. F. 1 73 205
2 174 177,
Harpinia sp. G. 2 187 185
1 252 210

160 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Number of Sample Depth in
Name of Species Specimens Number Fathoms

Heterophoxus sp. A. 1 6c 129
1 lla 291

7 I1c sy

11 12a 228

2 60 80

1 61b ZA

3 69 400

3 104 60

1 98 215

11 122 is)

1 124 50

3 128 100

30 133 47

50 159a 26

5 161 136

6 173 205

2 174 iyi

7 185 30

1 186a 19

1 187 185

3 224e 45

2 225a 128

1 228 293

1 238a 32

1 outside 58

Heterophoxus sp. B. 2 104 60
1 121b 230

5 148 170

1 209a 200

1 252 210

Leptophoxus sp. A. 2 10b 251
1 164b 470

3 173 205

Metaphoxus sp. A. 3 6la 20
+ 80b 23

6 81b 18

12 SOs 18

No. 1 HARTMAN: QUANTITATIVE SURVEY 161

Number of Sample Depth in
Name of Species Specimens Number Fathoms

Metaphoxus sp. A. (cont.) 83 19
99b 50

Paraphoxus sp. A.

Phoxocephalus sp. A.

W
WNHWRNNWOWNKF KE NNN DWDKEP KR DH NIN KP KP KP Re WRWUDD KF KE HY OH
—
1)
—_—
om
i)
WwW
S

Pontharpinia sp. B.

—

162 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Number of Sample Depth in
Name of Species Specimens Number Fathoms
Pontharpinia sp. E. 15 60 80
3 61b 21
1 68 5)
11 80b 23
58 104 60
9 107 18
1 109 61
10 22 115
66 133 47
26 159a 26
8 185 30
Pontharpinia sp. F. 4 174 177
Pontharpinia sp. G. 1 49 1%
1 66a 12
Pontharpinia sp. J. 2 6la 20
13 80b 25
48 104 60
Z 124 50
18 133 47
13 159a 26
3 174 177
8 185 30
Pontharpinia sp. K. 1 61b 21
23 64 5)
1 65 14
26 66a 12
1 68 5
3 81b 18

Pontharpinia sp. L.

Pontharpinia sp. M.

CON RNNNN KS
ioe)
S
or
bo
Ww

No. | HARTMAN: QUANTITATIVE SURVEY

Name of Species

Pontharpinia sp. M. (cont.)

Pontharpinia sp. N.
Ponthar pinia sp. P.

Pontharpinia sp. Q.

Pontharpinia sp. R.

Pontharpinia sp. S.

Pontharpinia sp. T.
Pontharpinia sp. U.

Pontharpinia sp. V.

phoxocephalids

Number of
Specimens

RO — —
We SY ORNKF DAWA NK AUDOANDHDOK KUN Pre DN We

—

——

—

— —
— op

Sample
Number

63b
224e
174

87
100
101
106
124

163

Depth in
Fathoms

164 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

D.

A LIST OF BRACHIOPODA IDENTIFIED
FROM SAMPLES OF THE BENTHOS OF
SAN PEDRO BASIN, CALIFORNIA

By O.tGA HARTMAN

Sample Depth in
Name of Species Number Fathoms

Class INARTICULATA
Order ATREMATA

Glottidia albida (Hinds) 28 11
45a 12
45b 12
46a 13
46b 11
49 LS
64 15
65 14
66a 12
80a 29
81b 18
82 18
101 29
108 17
159a 26
164a 28
184a 54
185 30
186a 19

No. 1 HARTMAN: QUANTITATIVE SURVEY 165

Sample Depth in
Name of Species Number Fathoms
Glottidia albida (cont.) 224a +4
224e 45
Class ARTICULATA
Order TELOTREMATA
Laqueus californicus (Koch) 238a 52

(many dead shells,
mostly unworn)
Terebratalia transversa (Sowerby)
(mostly dead shells, 238a 57
strewn in masses of
Laqueus californicus)

166 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Fr.

A LIST OF ENTEROPNEUSTA IDENTIFIED
FROM SAMPLES OF THE BENTHOS OF
SAN PEDRO BASIN, CALIFORNIA

By Keira Woopwick

Number of Sample Depth in
Name of Species Specimens Number Fathoms
Family PTYCHODERIDAE
Balanoglossus sp. 1 159b 26
ptychoderids 2 186a 19
1 207 28
1 224c a+
Family SPENGELLIDAE
Schizocardium sp. 3 24 386
1 31 9
8 44a 10
3 45a 12
2 46a 13
3 46b 11
Family Harrimaniidae
Saccoglossus sp. 1 186a 19
i, 186b 25
Stereobalanus sp. 1 89 386
1 145 430
19 159a 26
11 159b 26
1 near 171 400
1 185 30
1 22 446

No. | HARTMAN: QUANTITATIVE SURVEY 167

Sample Depth in

Name of Species Number Fathoms
unidentified enteropneusts 32 18 325
5 61b 21
1 80a 29
5 186a 19
i 189 335
1 224a 44

168 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

F.

AOLIST OF POLYCHAETOUS ANNELIDS
IDENTIFIED FROM SAMPLES OF THE BENTHOS
OF SAN PEDRO BASIN, CALIFORNIA

By O.tcA HARTMAN

Name of Species Sample Number Name of Family

Acrocirrus ?crassifilis lla Cirratulidae
Moore, 1923

Aglaophamus dicirris 164a, 186a, 207 Nephtyidae
Hartman, 1950

A glaophamus spp. 15, (0d, 123a) 126) 127 Nephtyidae

A maea occidentalis 44b, 65, 108, 164a, 207, Terebellidae
Hartman, 1944 224d

?A maea sp. 109, 124, 185 Terebellidae

A mage sp. 46a, 228 Ampharetidae

Ammotry pane sp. 10b, 41b, 80a, 109, Opheliidae

123a, 159a, 173, 174,
184a, 213, 224a, 225a, 252

Ampharete 2arctica 44b, 164a Ampharetidae
Malmgren, 1866

Ampharete sp. 82, 224d Ampharetidae

ampharetids LOD, 124,23. 24.- 25229.

41b, 45b, 46b, 56, 58a, 60,
61b, 63a, 80b, 89, 96, 97,
99b, 100, 123a, 126, 128,
133, 145, 146, 161, 173,
182, 184a, 185, 189, 195,
196,207, 213, 215, 224e,
225a, 227b, 249

No. | HARTMAN: QUANTITATIVE SURVEY

Name of Species

A mphicteis
scaphobranchiata
Moore, 1906

Amphicteis spp.

amphinomid

Amphisamytha bioculata

(Moore) 1906
A naitides spp.

Ancistrosyllis bassi
Hartman, 1945
Ancistrosyllis rigida
Fauvel, 1919
Ancistrosyllis spp.

Anotomastus gordiodes
(Moore) 1909

A phrodita armifera
Moore, 1910

A phrodita parva
Moore, 1905

A phrodita spp.

Arabella sp.

arabellids

arabellid, parasitic

Aricidea jeffreysi
(McIntosh) 1879

Aricidea ?pacifica
Hartman, 1944

Sample Number

18, 25, 31, 38, 44b,
45a, 46b, 65, 66a, 97,
?120, 122, ?near 171,
193,213, 2274

lla, 46a, 159a, 172,
225a, 228

45b, 69

164a

1. 6c 6d, Pla ii;
12a, 41b, 42a, 45b, 50,
63b, 81b, 109, 126,
161, 173, 174, 186a,
209a, 224d, 224e, 228,
236, 238a, 260

44b, 46a, 81b

45a, ?121b, ?234

6b, 6c; 7b, 23, 31,
108, 126, 145, 146,
159a, 170, near 171,
172, 185

28, 66a

164a
45a

28, 44a, 45b, 108,
181, 195, 224d

31, 46b, 66a

29, 45b, near 86, 143
124, 224e

14, 18, 145, 212

122

169

Name of Family
Ampharetidae

Ampharetidae

Amphinomidae
Ampharetidae

Phyllodocidae

Pilargiidae
Pilargiidae

Pilargiidae

Capitellidae

Aphroditidae
Aphroditidae
Aphroditidae

Arabellidae

Paraonidae

Paraonidae

170 ALLAN HANCOCK PACIFIC EXPEDITIONS

Name of Species

A ricidea spp.

Armandia sp.

Artacaminae, new genus
and species

Asclerocheilus sp.

Asychis lacera

(Moore) 1923
Asychis sp.
Autolytus sp.

A xiothella spp.
Boccardia ?redeki

(Horst) 1920

Boccardia spp.

Brada ?pluribranchiata
(Moore) 1923
Brada spp.

Capitella ovincola
Hartman, 1947
Capitella sp.
capitellid, new genus
and species
capitellids

Sample Number

7b, 28, 38, 41b, 42b,
44b, 49, 61b, 63a,

63b, 66a, 69, 80a,

80b, 81b, near 86, 89,
100, 101, 108, 121b,
123a, 124, 128, 133,
146, 148, 161, 164a

170, near 171, 172,

173, 174, 184a, 185, 189,
207, 211, 213, 214, 224a,
224d, 225a, 227b, 228,
234, 248, 249

44b, 224d, ?224e

46b, 80b, 99b, 164a, 224d

lla
?46a, 46b, 82, 2108, 121b

45a, 59b, 159a, 184a,
69, 108

46b, 63b

44b

44b, 45b, 46a, 46b,
61b, 63a, 108, 236
45a

5, 6d, 12a, 31, 42a,
46a, 46b, 66a, 82,
?121b, 126, 2146, 148,
185, 249

80b

6d, 46a, 133

89, 120, 170, near 171,
214

6b:.6d,, 7b, tay 12a. 18:
24, 25, 29, 41b, 45a, 45b,

voL. 19

Name of Family

Paraonidae

Opheliidae
Terebellidae

Scalibregmidae
Maldanidae

Maldanidae
Syllidae
Maldanidae
Spionidae

Spionidae
Flabelligeridae

Flabelligeridae

Capitellidae

Capitellidae
Capitellidae

No. 1 HARTMAN: QUANTITATIVE SURVEY 171

Name of Species Sample Number Name of Family
capitellids (cont. ) 46b, 59b, 61b, 65, 69, 97,
101, 108, 109, 145, 146,
148, 164a, 173, 189, 196,
207, 225a, 225b, 230a, 234,

236, 249
Carazzia sp. 44b Spionidae
Ceratocephala crosslandi 1c, 133, 159a, 185 Nereidae
americana 249
Hartman, 1952
Ceratonereis spp. 45b, 46b, Nereidae
chaetopterids Gay Wa: 975 213 Chaetopteridae
Chaetopterus variopedatus 186a Chaetopteridae
(Renier) 1804
Chaetopterus sp. 27, 43, 110, 164a, 207 Chaetopteridae
Chaetozone corona 44b, 66a, 170, 186a Cirratulidae
Berkeley and Berkeley,
1941
Chaetozone spp. 29, 45b, 46a, 49, 63b, Cirratulidae
80a, 81b, 123a, 123b,
215, 224a, 224d, 250a
Chloeia pinnata 2, 4, 6b, 6c, 6d, 12a, Amphinomidae
Moore, 1911 42a, 42b, 59b, 80a,
98, 123b, 133, 159a
Chloeia spp. 6a, 41b, 60, 89, 99b Amphinomidae
101, 109, 123a, 124,
127, 129, 148, 149,
184a, 186, 214, 224a, 224e,
225b, 230a, 239, 241,
249, 250a, 251, 260, 267b
Chone ecaudata 69 Sabellidae
(Moore) 1923
Chone spp. 1, 41b, 44b, 45b, 49, Sabellidae

63a, 63b, 65, 66a,
80b, 81b, 82, 89, 101,
108, 161, ?196, 224a,
224e, 225a, 260
Cirratulus ?cirratus lla, 44b Cirratulidae
(Miller) 1776

172 ALLAN HANCOCK PACIFIC EXPEDITIONS

Name of Species

Cirratulus sp.

cirratulids

Cirriformia ?luxuriosa
(Moore) 1904

Cirriformia sp.

Cistenides sp.

Cossura n. sp.

Dasybranchus sp.

Dexiospira sp.

Diopatra tridentata
Hartman, 1944

Diopatra sp.
disomid
Dodecaceria sp.
Dorvillea articulata
(Hartman) 1938
Dorvillea 2gracilis
(Hartman) 1938
Dorvillea spp.

Drilonereis falcata

Moore, 1911

Sample Number

lla

Li6e, 7b 12a: 18: 24
27, 38, 41b, 42a, 45b,
46b, 61b, 63b, 65, 66a,

69, 80a, 80b, 82, near 86,

96, 100, 101, 108, 109,

122, 123b, 127,133; 146,

159a, 161, 164a, 170,
near 171, 172, 173, 174,

184a, 186a, 195, 207, 209a,
224a, 224e, 225a, 230a, 238a,

248, 252
44b

186a

27, 28, 45b

6c, 7b, 12a, 29,

42b, 44b, 45a, 46a, 46b,
66a; 108, 12751285 133,
146, 159a, 161, 170,
near 171, 185, 189, 193,
212, 213, 214, 224d, 228
28

81b

28, 44b, 45a, 46a, 46b
65, 66a, near 86,

108, 159a, 164a

29, 31, 127, 207, 224d
31)238a

236

44b

63b
6c, 41b, 42a, 45b,

50, 61b, 63b, 81b, 164a
?108

voL. 19

Name of Family

Cirratulidae

Cirratulidae

Cirratulidae

Cirratulidae
Pectinariidae

Cirratulidae

Capitellidae

Serpulidae
Onuphidae

Onuphidae
Disomidae
Cirratulidae
Dorvilleidae
Dorvilleidae

Dorvilleidae

Arabellidae

No. l

Name of Species

Drilonereis nuda
Moore, 1909

Drilonereis spp.

Ehlersia heterochaeta

Moore, 1909

Eteone californica

Hartman, 1936

Eteone spp.
Euchone sp.

Euclymenini
Eulalia spp.
Eumida spp.

Eunice americana
Hartman, 1944
Eunice aphroditois
(Pallas) 1788
Eunice spp.
Euphrosine sp.
Evarnella fragilis
(Moore) 1911
Exogone sp.
Exogoninae
Fabricia sp.
Fabricinae
flabelligerid, new genus
and species

flabelligerids

Glycera americana

Leidy, 1855

HARTMAN: QUANTITATIVE SURVEY

Sample Number
?46a

1, 11c, 36, 45a,

46b, 61b, 63b, 65,
66a, 80b, 82, 100,
12251234, 159a9 161:
164a, 209a, 213, 224a,
224e, 225a, 252

108
44b

29, 45b, 49, 108

49, 146, 164a, 172,
224d, 224e

18, 82, 159a

61b, 81b, 164a, 186a,
lla, 28, 41b, 42b, 45b,
63a, 81b, 82, 186a, 207,
224e

159a

161

41b, 61b, 148, 224e, 236
lla, 45b
lla

164a

28, 45b, 46a, 225a
42b, 44b, 224d

80a, 99b, 121b, 224e
iS

42a, 45b, 61b, 65, 108,
120, near 171
12a, 44b

WE

Name of Family

Arabellidae

Arabellidae

Syllidae
Phyllodocidae

Phyllodocidae
Sabellidae

Maldanidae
Phyllodocidae
Phyllodocidae

Eunicidae
Eunicidae

Eunicidae
Euphrosinidae
Polynoidae

Syllidae
Syllidae
Sabellidae
Sabellidae
Flabelligeridae

Glyceridae

174

Name of Species

Glycera branchiopoda
Moore, 1911

Glycera ?capitata
Oersted, 1843

Glycera tesselata

Grube, 1863
Glycera spp.

Glycinde sp.
Goniada spp.

goniadids

Halosydna latior
Chamberlin, 1919

Halosydna spp.

Hapbloscoloplos elongatus
(Johnson) 1901

Hapbloscoloplos sp.
Harmothoé ?imbricata
(Linnaeus) 1767
Harmothoé scriptoria

Moore, 1910

Harmothoé triannulata
Moore, 1910
Harmothoé spp.

ALLAN HANCOCK PACIFIC EXPEDITIONS

Sample Number

?58a, 89, 213, 230a,
2234, 248
46a

11a, 108, 225a

6c, 6d, 11c, 12a, 29, 31, 41b,
45a, 45b, 46b, 50, 61b, 97,
100, 109, 122, 123a, 123b,
126, 128, 133, 146, 148, 159a,
161,164a,173,174,184a,185,
193, 207, 209a, 215, 224d,
224e, 236, 238a, 249, 260
12a, 41b, 109, 133, 184a

1, 6c, 6d, 12a, 45a, 46b, 49,
65, 66a, near 86, 127, 164a,
213, 224d, 225a, 228, 238a
60, 63b, 81b, 82, 108, 123a,
126, 128, 148, 158, 159a,
161, 184a, 185, 224a, 224e
45a

28, 45b, 80b

6c, 12a, 44b, 45a, 45b,

46a, 46b, 65, 66a, 80b, 100,
108, 109, 159a, 161, 164a,
209a, 224a, 225a, 228

5, 29, 31, 41b, 61b

49

1, 6d, 41b, 44b, 45a, 45b,
46b, 60, 81b, 123b, 124,
128, 148, 159a

69

31, 63b, 66a, 100, 101,
1081220133. 185,:213;
224a, 224d, 224e

voL. 19

Name of Family

Glyceridae
Glyceridae
Glyceridae

Glyceridae

Goniadidae
Goniadidae

Polynoidae

Polynoidae
Orbiniidae

Orbiniidae
Polynoidae

Polynoidae

Polynoidae

Polynoidae

No. | HARTMAN : QUANTITATIVE SURVEY VS)

Name of Species Sample Number Name of Family
harmothoid 1, 126
?Hauchiella sp. tla Terebellidae
hesionids 41b, 42b, 46b, 69, 145
Hesperalia sp. 45b Syllidae
Hyalinoecia juvenalis 80b, 2108, 159a Onuphidae
Moore, 1911
Hyalinoecia sp. 61b, 82, 185 Onuphidae
Hydroides norvegica (usually contaminant from Serpulidae
Gunnerus, 1768 hull of Velero IV)
Hydroides sp. 212, 249 Serpulidae
?Hypoeulalia bilineata lla, 164a Phyllodocidae
(Johnston) 1840
?Tsolda sp. 46a, 89 Ampharetidae
Labidognathus sp. 82 Arabellidae
Lagisca sp. 10b, 11a, 38, 69,119,173 Polynoidae
Lanice sp. 10b, 52, 148, 164a, 207 Terebellidae
Laonice cirrata 45a Spionidae
(Sars) 1851
Laonice spp. 1953, bla. 23. 24 29: Spionidae

41b, 44b, 46a, 46b, 65, 66a,
Sib} 123b, 126, 133,173,
185, 193, 224d, 225a, 228

?Leanira sp. 80b Sigalionidae

Leocrates n. sp. 18, 28, 45b, 63b, 2127, Hesionidae
?128, 148, 185

Lepidasthenia virens 45a, 250a Polynoidae

(Blanchard) 1849

Lepidasthenia spp. 6b, 7b, 46b, 66a, 81b, Polynoidae
108, 207, 224e

Lepidonotus spp. 11a, 45b, 236 Polynoidae

Loandalia fauveli 108 Pilargiidae

Berkeley and
Berkeley, 1941

Longosoma catalinensis 98, 224e Longosomidae
Hartman, 1944
Lumbrineris californiensis 108, 159a Lumbrineridae

Hartman, 1944
Lumbrineris cruzensis 1, 10b, 44b, 66a, 89, 108, Lumbrineridae
Hartman, 1944 209a, 225a

176

Name of Species

Lumbrineris 2erecta
(Moore) 1904

Lumbrineris index

Moore, 1911

Lumbrineris japonica
(Marenzeller) 1879

Lumbrineris minima

Hartman, 1944

Lumbrineris spp.

Magelona, near pacifica

Monro, 1933
Magelona spp.

Maldane spp.

?Maldanella robusta
Moore, 1906

maldanids

ALLAN HANCOCK PACIFIC EXPEDITIONS

Sample Number
44b

6c, 46a
46a
46a, 66a

1, 5, 6b, 6c, 6d, 7b, 11a,
1c, 14, 18, 24, 28, 29, 31,
36, 41b, 42a, 42b, 44a, 44b,
45a, 45b, 46b, 49, 60, 61b,
63a, 63b, 65, 66a, 69, 80a,
80b, 81b, 82, near 86, 100,
101, 108, 109, 122, 123a,
IZ3 D126) 12701285133;
143, 148, 158, 159a, 161,
164a, near 171, 174, 184a,
185, 186a, 207, 224a,
224d, 224e, 228, 230a, 260
45a, 108, 159a

28, 44b, 46b, 49, 63a, 63b,
65, 66a, 80, 80a, 80b, 81b,
82, near 86, 100, 109, 123a,
124, 164a, 184a, 185, 224a,
224e, 225a, 250a, 260

1. Ob Ierls! 23. 59b: 65,
97 12012 Wb Wl 28, 1324 33)
146, 148, 159a, 161, 170,
near 171, 184a, 184b, 185,
[9SP195, 196212215,
225a, 225b, 234, 248, 252
80b

1, 5, Ga, 6c; 7b; 10b;, 12a, 24,
29, 31, 41b, 42a, 45a, 49, 52,
60, 61b, 63b, 66a, 80a, 80b,

voL. 19

Name of Family

Lumbrineridae
Lumbrineridae
Lumbrineridae
Lumbrineridae

Lumbrineridae

Magelonidae

Magelonidae

Maldanidae

Maldanidae

No. |

Name of Species

maldanids (cont.)

Mar physa mortenseni
Monro, 1928

Mar physa, resembling
conferta Moore, 1911

Mar physa spp.

Megalomma sp.

Melinna spp.

Mesochaetopterus sp.
Myriochele sp.

Myxicola sp.

Naineris sp.

Nephtys 2caecoides
Hartman, 1938

Nephtys californiensis
Hartman, 1938

Nephtys ferruginea
Hartman, 1940

Nephtys spp.

HARTMAN: QUANTITATIVE SURVEY

Sample Number
89, 97, 98, 100, 101, 108,

109; 120, 122, 1234, 123,

126130335145, 198;

164a, 172, 174, 184a, 185,
189, 207, 224a, 224e, 225a,
236, 238a, 249, 250a, 267a

14

12a, 29, 31, 42b, 44b,
45a, 46a, 108

46b, 49

31, 44b, 45b, 123a, 123b,

133, 184a, 207, 224d, 224e

10b, 41b, 44b, 45a, 65,
$Z, 96, 108, 12 1b, 122,
126, 128, 146, 161, 225a
6c, 2126; 195

10b, 12a, 60, 69, 80a,

99b; 1005120) 123.4, 23h;

127, 1335146;,159a; 161,
172, 184a, 185, 189, 195,

1962135219. 2242,224e,

248, 250a
45b, 213

66a, 213, 224e
46a, 63a, 108

164a
5, 60, 126

6b, 6d, 28, 29, 41b, 42a,
44b, 61b, 63b, 66a, 80a,

80b, 82, near 86, 100, 108,
109, 121b, 122, 1234, 123b,

127, 128, 148, 159a, 161,

164a, 172, 173, 209a, 224a,

224d, 225b, 250a

La

Name of Family

Eunicidae
Eunicidae

Eunicidae

Sabellidae

Ampharetidae

Chaetopteridae
Oweniidae

Sabellidae
Orbiniidae
Nephtyidae
Nephtyidae
Nephtyidae

Nephtyidae

178

Name of Species

nephtyids

Nereis procera
Ehlers, 1868
Nereis spp.

nereids

Nerine foliosa

(Audouin- M. Edwards)

1834, new subspecies
Ninoé sp.
Nothria ?elegans
(Johnson) 1901
Nothria ?geophiliformis
(Moore) 1903
Nothria iridescens
(Jchnson) 1901
Nothria pallida
Moore, 1911
Nothria 2stigmatis
(Treadwell) 1922
Nothria spp.

Notomastus 2hemipodus
Hartman, 1947
Notomastus lobatus

Hartman, 1947
Notomastus spp.

Odontosyllis sp.

Oncoscolex sp.

Onuphis eremita
Audouin and

M. Edwards, 1834

ALLAN HANCOCK PACIFIC EXPEDITIONS

Sample Number

7b, 12a, 46b, 49, 65, 99b,
132, 133, 149, 158, 184a,
185, 224e, 249

29, 44b, 45a, 46a, 49, 108,
224d

11a, 31, 46b, 80b, 101,
109, 159a

6b, 28, 42b, 65, 66a, 81b,
82, near 86, 161, 181, 204,
224e, 225a

80b, 108

7b, 224d
49, 63a
65

108

1, 42a, 129, 130, 184b, 185a

225a

6c, 6d, 7b, 10b, 41b, 45a,
46b, 65, 66a, 80b, 81b, near
86, 121b, 126, 148, 158,

161, 209a, 224a, 224e, 225b

44b
122, 228

6c, 31, 46a, 49, 80b, 159a,
209a, 225a

28, 123b, 224d, 224e
123a, 123b

49, 63a

voL. 19

Name of Family

Nereidae

Nereidae

Spionidae
Lumbrineridae
Onuphidae
Onuphidae
Onuphidae
Onuphidae
Onuphidae

Onuphidae

Capitellidae
Capitellidae
Capitellidae
Syllidae

Scalibregmidae
Onuphidae

No. l HARTMAN: QUANTITATIVE SURVEY 179

Name of Species Sample Number Name of Family
Onuphis parva 122, 123b, 224a Onuphidae
Moore, 1911
Onuphis, near vexillaria 10b Onuphidae
Moore, 1911
Onuphis spp. 1, lla, 41b, 59b, 80b, 101, © Onuphidae
123a,125, 2246) 225a, 252
onuphids 5, llc, 12a, 24, 28, 36, 41b,
45b, 63b, 80a, 100, 132, 133,
164a, 225a
Ophelia sp. 49, 63a Opheliidae
opheliid 46a
orbiniid, new genus and 18, 59b, 69, 89,120,170, | Orbintidae
species near 171, 172, 189, 196,
215, 249, 253
orbiniids 23, 42a, 42b, 97
Owenia sp. 31, 45a, 45b, 46a, 49, 50, Oweniidae

80b, 82, 100, 123a, 164a,
186a, 224e, 238a

owenlids 41b, 42b, 234

Panthalis pacifica 184a Polyodontidae
Treadwell, 1914

Panthalis spp. 133, 207, 224a Polyodontidae

Paraonis n. sp. 7b, 10b, 18, 42b, 44b, 46b, Paraonidae

63b, 66a, 69, 80a, 97, 121b,
123a, 124, 133, 148, 161,
170, 184a, 186a, 189, 211,
212, 224d, 224e, 225a, 227b.

230a
paraonoids 24, 46a, 82, 108, 123a, 159a
?Pareurythoé sp. 7b Amphinomidae
Pectinaria californiensis 1, 6c, 6d, 7b, 10b, 12a, 28, —Pectinariidae
Hartman, 1941 44b, 45a, 46a, 59b, 80b, 108,

123b, 133, 158, 161, 184a,
209a, 224d, 225a, 225b
Pectinaria spp. 5, 7a, 29, 41b, 42a, 42b, Pectinariidae
61b, 80a, 99b, 109, 121b,
123a, 127, 128, 148, 159a,
173, 182, 185, 224a, 224e

180 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19
Name of Species Sample Number Name of Family
Peisidice aspera 42b, 44b, 45b Polyodontidae
Johnson, 1897
Peisidice sp. 101, 224e Polyodontidae
Perinereis sp. 49 Nereidae
Petaloproctus sp. 10b, near 171 Maldanidae
Pherusa capulata 123b, 173, 224e Flabelligeridae
(Moore) 1909
Pherusa inflata 45b, 46b Flabelligeridae
(Treadwell) 1914
Pherusa spp. 6b, 6c, 6d, 10b, 11a, 12a, Flabelligeridae
41b, 44b, 45a, 46a, 60,
66a, 80a, 80b, 82, 100,
108, 109, 122, 123a, 146,
159a, 164a, 186a, 196, 224e
238a
Pholoé sp. 42b, 45a, 60, 61b, 66a, Sigalionidae
80b, 81b, 82, 99b, 100,
108, 122, 123b, 128, 133,
148, 159a, 161, 184a, 185,
224a, 224d, 260
Phyllochaetopterus 45b, 61b, 186a Chaetopteridae
?prolifica Potts, 1914
Phyllochaetopterusn.sp. 8, near 15, 16, 21, 22,23, | Chaetopteridae
24, 36, 38, 39, 53, 54, 55,
50) OF 00a, 70/74, feuds:
74, 75, 91, 92, 93, 94, 95,
112, near 112, 113, 114,
RIS 1G) WIAs 137,
138, 139, 140, 141, 142a,
142b, 143, 144, 164b, 165,
166, 167, 168, 169a, 169b,
190, 191, 192, 211, 2?227b,
248
Phyllochaetopterus sp. 46a, 69, 89, 90 Chaetopteridae
?Phyllodoce ferruginea 225a Phyllodocidae
Moore, 1909
Phyllodoce sp. Ila, 45a, 46b Phyllodocidae
phyllodocids 46a, 61b, 65, 66a, 80a, 80b,

108, 207, 224e, 234

No. 1

Name of Species
Pilargis ?maculata

Hartman, 1947
Pilargis spp.

?Pionosyllis sp.
Pisione, near remota

(Southern) 1914
Pista 2cristata

(Miller) 1776
Pista disjuncta

Moore, 1923
Pista elongata

Moore, 1909
Pista spp.

Podarke pugettensis
Johnson, 1901
Podarke spp.

Poecilochaetus johnsoni
Hartman, 1939
Poecilochaetus sp.

Polycirrus sp.
Polydora, near armata
Langerhans, 1880

Polydora citrona
Hartman, 1941
Polydora, near ligni
Webster, 1879

Polydora spp.

polydorid
polynoids

HARTMAN: QUANTITATIVE SURVEY

Sample Number
108

66a, 148, 158, 161, 196,
2209a

108

14, 63b

45b, 59b, 65, 80b, 108,
122, -207,.2252

158

45b, 2207, 224e

41b, 42a, 44b, 45a, 46b,
66a, 80a, 82, 96, 108, 123a,
133, 159a, 172, 185, 193,
224e

44b

29, 42a, 45a, 45b, 66a,
108, 133, 146, 159a
45a, 46a, 108, 109, 224d

28, 46b, 49, 66a, 159a,
164a, 185, 249

49, 265, 186a, 224d, 236
44b

44b
44b

11a, 12a, 38, 41b, 42b, 61b,
159a, 173, 184a, 224e
164a

6c; 10b, tlic, 122,23, 28,
46a, 109, 121b, 123a, 161,
173, 174, 186a, 211, 238a,

181

Name of Family

Pilargiidae
Pilargiidae

Syllidae
Pisionidae

‘Terebellidae
Terebellidae

Terebellidae

Terebellidae

Hesionidae
Hesionidae
Disomidae
Disomidae

Terebellidae

Spionidae
Spionidae
Spionidae

Spionidae

182 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19
Name of Species Sample Number Name of Family
polynoids (cont. ) 249, 267a
Polyodontes, near 159a Polyodontidae
panamensis
(Chamberlin) 1919
polyodontids Zeolite
Praxillella affinis pacifica 108 Maldanidae
Berkeley, 1929
Praxillella sp. 46b, 65, 81b, 82, 133,215 Maldanidae
Prionospio 2cirrifera 12a, 225a Spionidae
Wirén, 1883
Prionospio, near 28, 44b, 45a, 49, 60, 65, Spionidae
malmgreni 66a, 80b, 81b, 82, 99b,
Claparéde, 1870 108, 128, 159a, 161, 164a,
224a, 224d
Prionospio pinnata 1, 7b, 12a, 29, 31, 41b, Spionidae
Ehlers, 1901 44b, 45a, 49, 60, 65, 66a,
80b, 82, 108, 109, 121b,
N22, 1260, 12728. 148;
158, 159a, 161, 164a, 185,
207, 224a, 224d, 225a, 252
Prionospio spp. 1, 6b, 6d, 7b, 10b, 46b, 63a, Spionidae
80a, near 86, 97, 100, 101,
123 a 123 b6133, 172073)
174, 184a, 224e, 238a, 250a,
260
Protula sp. lla, 23, 24, 36, 53,54,55, Serpulidae
56; 9/5109) 207k, 12089:
90, 91, 95, 96, 97, 119,
139, 143, 145, near 171, 189,
193, 194, 211, 227b, 248
Psammolyce sp. 186a Sigalionidae
Pseudopotamilla sp. 164a, 186a Sabellidae
Rhamphobrachium sp. 80b, 122, 164a, 224e Onuphidae
Rhodine sp. 28, 65, 148, 159a, 164a Maldanidae
184a, 185, 207, 248, 249,
252
Rhynchospio arenincola 63a, 80b, 108 Spionidae

Hartman, 1936

NO. 1 HARTMAN: QUANTITATIVE SURVEY 183

Name of Species Sample Number Name of Family
Rhynchospio sp. 49, 63b, 66a, 81b Spionidae
Sabella, resembling 164a Sabellidae

crassicornis Sars, 1851
Sabella sp. 80b Sabellidae
Sabellinae, new genus 119 Sabellidae

and species
sabellids 6d, 12a, 18, 69, 80a, near

86, 100, 120, 123a, 146,
164a, near 171, 181, 185,
189, 204, 207, 212, 214,

224d, 224e
Sabellaria sp. 45b, 159a Sabellariidae
Saccocirrus papillocercus 50 Saccocirridae
Bobretzky, 1871
Salmacina sp. 45b Serpulidae
Scalibregma sp. 6d, 12a, 28, 29, 31, 45a, Scalibregmidae
45b, 46a, 46b, 49, 61b, 63b,
65, 66a, 80b, 82, 100, 101,
108, 109, 123a, 159a, 164a,
186a, 195, 212, 224d, 224e,
225a, 238a
Schistocomus sp. 65, 66a, 80b Ampharetidae
Scionella sp. 225a Terebellidae
?Scolelepis sp. 234 Spionidae
Scoloplos spp. 45a, 49, 63a, 63b, 80a, Orbiniidae
81b, near 86, 101, 123a,
146, 148, 224d, 224e
?Semiodera sp. 186a Flabelligeridae
serpulids 38, 39, 159a, 164a, 172, 196
224e, 227a
sigalionids 18, 28, 46b, 61b, 80a, 80b,
100; 108; 123a, 123b, 133;
164a, 185, 224e
Sphaerodorum spp. near 86, 108, 133, 173, 185 Sphaerodoridae
sphaerodorids 123a, 174, 267a
Sphaerosyllis sp. ?173 Syllidae
Spio n. sp. 224a Spionidae

Spiochaetopterus sp. 44b, 45a, 63b, 101, 108, Chaetopteridae

184 ALLAN HANCOCK PACIFIC EXPEDITIONS

Name of Species

Spiochaetopterus sp.
(cont. )

spionids

Spiophanes missionensis

Hartman, 1941
Spiophanes sp.

spirorbids

Sternas pis sp.

Sthenelais ?tertiaglabra
Moore, 1910
Sthenelais spp.

Sthenelanella uniformis

Moore, 1910

Sthenelanella sp.

Streblosoma crassibranchia

Treadwell, 1914

Streblosoma sp.

Syllinae
Syllis sp.

Sample Number

122, 123a, 148, 159a, 164a,
186a, 207, 224d, 224e, 225a,

238a

6a, 11c, 42a, 42b, 45b, 52,
63b, 66a, 69, near 86, 100,
1095 1225133, 158,159:

186a, 238a, 249

44b, 46a, 63a, 108, 159a,

164a, 224d

7b, 24, 27, 29, 31, 42b, 45a,
46b, 49, 60, 61b, 65, 66a,
80a, 80b, 89, 100, 101, 122,
133, 146, 2148, near 171,
184a, 185, 193, 196, 207,

213, 224a, 224e

27, 45b, 109, 123a, 123b,

159a, 224e

5, 46a, 46b, 61b, 65, 81b,
82, 108 127 W234. 1235p,
128))133, 1594, 161, 1644,
184a, 185, 186a, 207, 224a,

224d, 224e, 238a, 250a
46a, 65

80b, 184a, 186a, ?207, 224a,

224d

44b, 45a, 46a, 65, 80b,
108, 159a, 164a, 186a,
250a

18, 27, 28, 45b, 61b, 82,

101, 123a, 207, 224a, 224d

44b, 45a, ?164a

12a, 27, 28, 44a, 46a,
?61b, 81b, 82, 108, 173,
?186a, 207, 228

45b

10b, 224e

VoL. 19

Name of Family

Spionidae

Spionidae

Serpulidae

Sternaspidae

Sigalionidae
Sigalionidae

Sigalionidae

Sigalionidae

Terebellidae

Terebellidae

Syllidae
Syllidae

No. 1

Name of Species

syllids

Terebellides sp.

terebellids

Thalenessa spinosa
(Hartman) 1939

T halenessa sp.

Tharyx parvus
Berkeley, 1929
Tharyx spp.

T helepus sp.

Travisia sp.

trichobranchiid
Try panosyllis sp.
Ty posyllis sp.

V ermiliopsis sp.

HARTMAN: QUANTITATIVE SURVEY

Sample Number

lla, 28, 63a, 63b, 65, 66a,
80a, 80b, near 86, 99b, 123a,
159a, 174, 207, 236

5, 6c, 11a, 29, 31, 45a,
80a, 80b, 82, 99b, 108, 122,
126, 148, 159a, 161, 164a,
173, 184a, 185, 207, 213,
224a, 225a, 228

1, 25, 31, 38, 42a, 45b, 52,
59b, 66a, 80a, near 86, 100,
109, 119, 161, 174, 184a,
185, 186a, 193, 224e

63a, 63b, 82, 108, 123a,
123b, 225a, 234, 250a

45b, 65, 66a, 124, 224a,
224e

42b, 44b, ?164a, 224d

29, 45a, 45b, 60, 66a, 80a,
81b, 82, 101, 123a, 124,
212, 213, 214, ?227b, 248,
250a

10b, 207, 224e, 227a, 250a
2, 6b, 44b, 105, 108, 123b,
133, 148, 159a, 164a, 225a
224e

186a

45a, 69

186a, 224e, 238a

185

Name of Family

Tricho-

branchidae

Sigalionidae
Sigalionidae
Cirratulidae

Cirratulidae

‘Terebellidae
Opheliidae

Syllidae
Syllidae
Serpulidae

PLATES

PL. 1A

ALLAN HANCOCK PACIFIC EXPEDITION VOL. 19

1 1

Emerald Cove, Catalina Island, California. Nov. 17, 1951. In
17 fms. 33° 28’ 05” north latitude, 118° 31’ 17” west longitude.
The rubbly appearance is caused by masses of Chaetopterus
tubes, tubicolous anemones, and other tube-dwelling animals.
A two sq. ft. sample yielded at least 60 species of metazoan
invertebrates, of which 28 are annelids, 3 are enteropneusts,
and the others are representatives of different phyla or classes
of animals. The bottom is highly productive. The base line
represents about five feet.

1 Photographs 1 to 4 were made with the benthograph operating from the Velero IV,
by William Fortin, staff photographer of the Allan Hancock Foundation.

No. |

bo

HARTMAN : BOTTOM SURVEY

Off Jew Fish Point, 200 yards NNE of Abalone Point and
southeast of Avalon Bay, at the east end of Catalina Island,
California. Sept. 22, 1951. In about 48 fms. 33 20’ 47” north
latitude, 118° 18’ 54” west longitude.

Surface features show tests of Lytechinus, sea whips, Chloeta.
Hummocks are those of large maldanids, burrowing anemones,
onuphids, and sigalionids. Depressions are made by ophiuroids,
other annelids, etc. The bottom is richly populated. Size
proportions can be estimated from the tests of Lytechinus, which
measure 30 to +0 mm across.

be Wi:

ALLAN HANCOCK PACIFIC EXPEDITION VOL. 19

we

In a deep part of San Pedro Basin, 11 miles northeast of
Avalon, Catalina Island, California. Aug. 11, 1951. In 350
to 400 fms. 33° 28’ 00” north latitude, 118° 11° 38” west
longitude.

Surface features show tests of d/locentrotus (measuring 30 to
80 mm across), a crinoid at right center, a sea star at lower
right. A two sq. ft. sample from the bottom yielded glass
sponge, many foraminiferans, 20 or more species of annelids,
many ophiuroids, and a high percentage of new or little known
animals.

No. | HARTMAN : BOTTOM SURVEY PL. 2B

4+. Off Jew Fish Point, southeast of Avalon Bay, Catalina Island,
California. Sept. 22, 1951. In about 30 fms.
Surface features show many tests of Lytechinus (measuring
30 to 40 mm across), a sea whip, tangled tubes of Phyllochae-
topterus at lower left; the bottom is inhabited by many species
of annelids, ophiuroids, and other invertebrate animals. The
large hummocks are those of a large maldanid, the smaller are
onuphid.

PAL YAN IPS, 3}

5. The Hayward orange-peel-grab, modified with canvas sleeve,
hooked up for descent to the bottom, shown on the rear working
deck of the Velero IV’.

6. Hoisted through the A-frame with the aid of air compression,
showing position of the claws at greatest extension.

7. Turned to show arrangement of canvas flaps which permit
escape of air or water, but prevent escape of solid particles.

8. About to enter the water over the point selected for investiga-
tion, showing the hook-up mechanism with the end of the cable.

2 The following photographs were made by Roy V. George, staff photographer
of the Allan Hancock Foundation, except No. 15, which was made by Floyd E.

Durham.

~~
-

eb, OY ALLAN HANCOCK FACIFIC EXPEDITION VOL. 19

Fig.

Cn

Fig. 6

Fig. 7 Fig. 8

10.

Mate

12.

PLATE 4

The A-frame at its greatest extension, with the orange-peel-
grab still descending; the cable taut, shown diagonally across
the picture.

The grab loaded with a sample from the bottom, just after
breaking the surface of the water; the jaws are well closed
to prevent the loss of contents.

The sample is hoisted through the A-frame and about to be
hooked to the chain and released from its cable, before it is
brought down.

It is placed over its large retaining tub on the rear platform
of the ’elero IV, and opened, to empty its contents in the tub.

PL. +A ALLAN HANCOCK FACIFIC EXPEDITION VoL. 19

18%

14.

15.

16.

ALANIS, 5}

The tub with its load of mud is pulled out from under the grab
and its contents are about to be measured with a graduated
brass rule. Two half-pint samples are set aside for geological
study.

The tub is moved to a screen assembly, consisting of 4 assorted
screens (ranging in size from coarse to fine mesh) on a table
of convenient height. The finest or lowest screen has a mesh
that measures 24 meshes to the inch.

The screen assembly is equipped with an overhead shower
spray which washes the sample; the particles are thus sorted
to size; then they are transferred to jars and fixed with for-
malized seawater.

The screen assembly is provided with a shaker device, located
under the bottom screen and operated with a small motor
housed at its far end; the sorting trays are on a raised platform,
shown to the left of the picture.

PL. JA ALLAN HANCOCK PACIFIC EXPEDITION VOL. 19

We

PLATE 6

The Campbell grab, named for its designer, Alex Campbell,
has a capacity of about 4.46 cu. ft., and weighs about 900
pounds; it was devised to take samples from hard-packed
bottoms; it is shown open, suspended in the A-frame on the
working deck of the Velero IV. Area covered, 6 sq. ft.

It is attached to the cable and hoisted overboard, with its
trigger mechanism set to shut on striking bottom; the vents
to permit escape of air or water pressure, are visible at the
upper end of the shell; they are covered by flaps to prevent
escape of mud.

The grab returns from the bottom with a load of mud; its
jaws close tightly so that even water is retained.

It is hoisted over its large retaining tub and hooked to its
chains, which are mechanically pulled to open the grab.

PL. OA ALLAN HANCOCK PACIFIC EXPEDITION vot. 19

Fig. 19 Fig. 20

PEATE 7

The Campbeil grab has emptied its contents into a rectangular
tub which easily accommodates its load; the tub is mounted
on a castored base to permit moving across the deck of the
Velero IV.

The mud in the tub is measured and two half-pint samples are
removed for future geological study; the grab is cleaned and
readied for another sample.

The screen assembly seen from the top, showing the six shower-
heads and the sorting trays; a canvas bag is in place over
the screen base, to catch smaller bits that are then washed
into jars for preservation.

The large Dragger winch on the working deck of the Velero
Il’, showing the drum within its housing and the extending
cable seen across the lower left of the picture.

PL. 7A ALLAN HANCOCK PACIFIC EXPEDITION VoL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

VOLUME 19 NUMBER 2

QUANTITATIVE SURVEY OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

PART If

FINAL RESULTS AND CONCLUSIONS
(Maps 1-2, Plates 1-13)

BY

OLGA HARTMAN

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS
LOS ANGELES, CALIFORNIA
1966

ALLAN HANCOCK PACIFIC EXPEDITIONS

VOLUME 19

OUANTITATIVE SURVEY OF THE

BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

BY

OLGA HARTMAN

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS
LOS ANGELES, CALIFORNIA
1955-1966

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ALAIN “EEANCOCK. FACIFPIC EXPEDITIONS

VOLUME 19 NUMBER 2

QUANTITATIVE SURVEY “OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN ‘CALIFORNIA

PRI EE

FINAL RESULTS AND CONCLUSIONS
(Maps 1-2, Plates 1-13)

BY

OLGA HARTMAN

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS
LOS ANGELES, CALIFORNIA
1966

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‘A SY AE

QUANTITATIVE SURVEY OF (THE
BENTHOS OF SAN "PEDRO BASIN,
SOUTHERN CALIFORNIA

PARIS Jil

FINAL RESULTS AND CONCLUSIONS
by
OLGA HARTMAN

THE UNIVERSITY OF SOUTHERN CALIFORNIA PUBLICATIONS
ALLAN HANCOCK PACIFIC EXPEDITIONS
VOLUME 19, NUMBER 2

IssUED: DECEMBER 5, 1966

PRICE: $7.50

THE UNIVERSITY OF SOUTHERN CALIFORNIA PRESS
Los ANGELES, CALIFORNIA

TABLE OF CONTENTS

Pages
PNR ODE THOM oS a csc lt ec det ght Oe eon
RCO Ss en ee, Pe Se cat, oe hy el Te melt
SIVA Y “OR RESUMES S25) sees 8 Ree of ot GO
Re NOVILEDGEMENDS =.) 2a 2 Soest 2 6 os. ce MO
DESCRIPTIONS OF NEW AND LITTLE KNOWN
LOLVCHAELTOUS ANNEEIDS. 34 < 2 & & =» « 192
ANALYSES, BY AREAS el acts “Pees ence yet were
MAPCOE. DHE SAN PEDRO REGION... = < 4 2.’ 207

MAP OF THE SANTA CATALINA ISLAND AREA . 208

LOS ANGELES-LONG BEACH OUTER HARBOR,

mUMBChICUTE, te key Vast te ed ao Gee eres hey? a ae os ae Vo A ee
PALOS VERDES SHELF AND SLOPE, with chart . . 219
SANA PEDRO SEBER, with charts 2 = = . < 31.) Zor
SAN PEDRO SEOPEwith*chart 2 3. 2 . «< = 2. «289
NEWPORT SHEER “with chart: . . . 2 « = «2 dil
SUBMERGED SEA MOUNTS, with chart . © = = 4 326

SANTA CATALINA ISLAND, LEEWARD SIDE,
ECAR Me. ccd sc SUA TR fo cil tate Seen iy uate FOS

SANTA CATALINA ISLAND, WINDWARD SIDE,
CULE Roel 0 17 i a ne SU Rene ew mr os. Aa eee 17/4

PHOTOGRAPHIC RECORDS, with chart ££ « . . « 389

TABLE OF CONTENTS

Pages
ALPHABETICAL LISTS OF SPECIES NAMED FROM
THESE AREAS: o> S281 4 25) 06h) Ee odo, oe eee
POLY CHAE TES... 3% Goce 2.98 em eee
ECHINODERMS <6 sis. av Sectow at eaeegiagee nll lon Sem
MOLLUSKS. oo 3 es. Si ee ta se = eres ee et
BRYOZOANS FROM FARNSWORTH BANK,
by? Walliam Banta 25%. a. «se 2 =<. 6) een ere
SERIAL NUMBERS 11 to 267, WITH STATION
INUMIBERS i. 0 os se a\bsyaeg = pecan ueek oo eee eee
ELFERATURE CLIVED,) cc see eres ce ren he ape eee
PICA ES ace fey trek 8S) ered ay yeh ass tot ee) cee & eed oe

DNIDE Xeigs aes Teg she ee ce ee

QUANTITATIVE SURVEY OF THE
BENTHOS OF SAN PEDRO BASIN,
SOUTHERN CALIFORNIA

PARE If

FINAL RESULTS AND CONCLUSIONS

INTRODUCTION

This is a continuation and conclusion of the study of the San Pedro
area for which preliminary results have been published (Hartman, 1955).
This area (Map 1) lies between the mainland of southern California
and Santa Catalina Island (Map 2), in 33° and 34° north Latitude,
and 117° to 119° west Longitude. The submarine lands comprise more
than 1060 square miles, of which the shelflands make up about a fourth,
the slopes and canyons together about half, and the San Pedro Basin
nearly a fourth. The terrain is rugged, consisting of shelf, slope, canyon,
basin and sea mount features; depths range from a few to 495 fathoms.
This large, partly enclosed, oceanic area receives not only the discharge
of the Los Angeles, San Gabriel and Santa Ana Rivers which drain the
slopes of the lofty San Gabriel and San Bernardino Mountains, but also
the liquid efHuent wastes of populous Los Angeles County. As a result,
much terrigenous material is carried downstream to the sea, where long-
shore currents, chiefly with a southeasterly drift, pick up the loads, to
deposit the coarser gravels along the beaches and shallow shelf; the
sands, silts and muds are carried successively farther out, or may be
deposited in the canyons and San Pedro Basin. All but the finest particles
may have been filtered out before the nutrient-rich waters reach the
outer shelf and slope. As a result, the long-shore shelves and slopes are
heavily populated with a richly diversified benthic fauna, which attains
its maximum development along the Palos Verdes and the San Pedro
shelves. These benthic populations chiefly represent a few groups, includ-

187

188 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

ing infaunal polychaetes, nemerteans, enteropneusts, and some others.
‘They maintain a dynamic stability, showing no seasonal differences, and
have continued to flourish through several years.

This broadly oval submarine area was marked off as a grid, at about
two-mile intervals, resulting in 267 intersecting points (Hartman, 1955,
pp. 6, 7). Grab samples were taken from these pre-selected sites, and
analyzed for their specific components. Ihe samples varied greatly in
volumetric sizes, ranging from 0.06 to nearly 3 cubic feet. The surface
areas covered by the sampler probably varied far less, because the initial
bite-size was more nearly equal for all samples. The sampler was effec-
tive in taking surface scratchings, where the largest numbers of animals
are known to occur. In assessing unit quantitative values, therefore, not
only differences in grab sizes, but kinds of substrata are to be considered.
It is further noteworthy that all final values must be considered minimal,
for only those organisms which finally reached the microscope and the
specialist could be fully named and recorded. In this study, therefore, it
would be meaningless to re-assess these areal quantities to units of a
square meter, for it has been observed repeatedly that the aggregates of
animals in the San Pedro area occur in small patches, with surface units
perhaps much less than a square meter. Even more importantly, it has
been shown that the faunal units are patchy, differing greatly from one
sample to the next.

Except for differences in contour, the physical features are fairly
uniform. Salinity is nearly constant throughout the year, ranging from
33.5% near the surface (physical data are from Emery, 1960, p. 108)
to 34.29% in 100 fms. There is some indication that lowered salinities
occur in scattered spots, possibly near aquifers, as indicated by the pres-
ence of peak numbers of the polychaete Capitella capitata subspp. Such
spots have been found in the San Pedro shelf as well as in other offshore
places (Hartman, 196la). Temperatures in the sediments vary little,
from 12.5°C near the surface, to 8.5°C in 100 fms, and 5.06°C in 495
fms, in the San Pedro basin. Dissolved oxygen varies from 5.5 ml/I at
50 fms, 1.5 ml/I at 100 fms, to only 0.2 ml/I in the San Pedro basin,
where there is almost no life. Sediments are chiefly sands, with coarser
fractions near beaches and increasingly finer sands at greater distances
from shore; sands change to silty sands, sandy silts and then to clays and
oozes along deep slopes, in canyon and basin depths, and axes of canyons,
and their lower ends may have coarse gravels (Emery and Hiulsemann,
1963, p. 62). Some rocky outcrops, such as the sea mounts, are conspicu-
ous in a few places. Red sands, near Alamitos Bay, in 3 to 7.5 fms, and
near the center of San Pedro shelf, are sparse.

No. 2 HARTMAN : QUANTITATIVE SURVEY 189
METHODS

This planned procedure was more or less consistently followed:
(1) The samples were collected, usually with the orange-peel grab, less
often with the larger Campbell grab, and processed aboard ship, using a
screen with finest mesh slightly less than a millimeter. (2) “Che washed
samples were fixed in formalin-seawater, and then transferred to the
laboratory, where more complete sorting and washing were done. Larger
macroscopic animals were removed, some of the larger ones weighed, and
assessments made of the largest individuals in the sample, the most con-
spicuous species, and the general characteristics of the faunal population.
(3) As many as possible of the invertebrate animals were identified to the
specific level, and counts made of individual species. Smaller animals,
including nematodes, small crustaceans (amphipods, isopods, cumaceans,
tanaids, ostracods, and others) as well as juvenile stages of other animals,
were probably taken only very incompletely because of the coarse tech-

niques employed. It is therefore all the more noteworthy that large num-
bers of these and other animals were retained in the individual samples.

(4) Many species from southern California were newly described,
based on specimens found in the materials. Separate reports have been
issued for some major groups. Amphipods have been studied by Dr. J.
Laurens Barnard (1957-1963) and Barnard and Given (1960), cuma-
ceans by Mr. Robert Given (1961, 1964) and Barnard and Given
(1961) and some isopods by Dr. Robert Menzies (1959). Echinoderms
were identified by Mr. Fred Ziesenhenne (1951) and Barnard and
Ziesenhenne (1961) and polychaetes by Hartman (1955-1963). Mol-
lusks were studied by Mr. Gilbert F. Jones (1963, 1965), the late
Dr. Norman Mattox (1955, 1958) and Dr. Mathilde Schwabl (1961,
1963). There still remain many groups of animals requiring study, the
most important of which are nemerteans, sipunculids, enteropneusts,
ostracods, tanaids, ceriantharian and other coelenterates, and nematodes.

(5) The distributions of individual species, with their relative abun-
dance, were named for each of the selected areas, and plotted by increasing
depth (see charts following the analyses for each area). These show not
only the recurrence of each species in single areas but throughout the
region, and associated ecological groupings, in a frame of reference.

190 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

SUMMARY OF RESULTS

A sampler lowered almost anywhere off the coast of southern Califor-
nia recovers an amazing array of kinds of animals. These kinds and
numbers vary from one sample to the next and attain large numbers in
shallow bottoms. This diversity extends not only to most samples and in
all kinds of sediments, but throughout the year, with little indication of
season. It is difficult to relocate the exact spot from which a sample was
taken, except in shallowest bottoms or in few places where one or few
species are known to occur in massed numbers. Such are some known
Dendraster and Capitella bottoms, where exceptional aggregates occur.
Usually the numbers of species in a sample from shallow bottoms range
to a hundred or more, with individuals numbering to near five thousand.
These species occur as aggregates, where the presence of one generally
indicates that of other kinds; they seem to maintain their restricted
locations over periods of years, as proven by subsequent probes in suc-
cessive seasons and years.

The kinds of species and their respective numbers, the stages of devel-
opment, and the state of maturity, as well as the associations with other
groups in the sample, are repetitive characteristics and should therefore
be predictable.

Samples with large numbers of specimens are generally characterized
by having one or a few species with exceptionally high numbers of indi-
viduals ; these specific units may change from one sample to the next, and
their occurrences are not as predictable as are the aggregated species with
which they occur. Their kinds and distributions are indicated by an aster-
isk in the charts, below. Exception must be made for the Amphiodia
urtica-Pectinaria californiensis association, in which individual numbers
continue high along most of the edge of the San Pedro shelf and in other
areas remote from shore, especially where sediments are somewhat impov-
erished. Isolated peak numbers are indicated for many of the more than
600 species named below. Peak numbers may be partly explained by the
successful establishment of spatfalls of larvae, but they do not explain the
continuance of the colony through successive generations.

Specific diversity in southern California is so characteristic that large-
scale communities of organisms cannot be identified. Exception may be
made for a Listriolobus pelodes colony on the Santa Barbara shelf (Bar-
nard and Hartman, 1959) where a continuous shelf depth is occupied by
an identifiable community of organisms.

Numbers of species and specimens are highest in shelf depths, and
lowest in outer slopes and the oxygen-impoverished, long-shore basin

No. 2 HARTMAN : QUANTITATIVE SURVEY 191

(Hartman and Barnard, 1958, p. 29). Numbers of species and specimens
have ranged to 88 species and about 4500 specimens per sample.

Biomasses are highest in long-shore areas, especially the Palos Verdes
and San Pedro shelves, where the sediments are enriched by the runoff of
the mainland. Single samples have yielded a gross moist weight of 768
grams per sample, and similar high biomasses have been found on the
San Pedro shelf and other places along the shelflands of southern Cali-
fornia (see California, 1965a, 1965b, pp. 104-319).

Most of the species named below may be expected to produce individ-
uals with a short life span, not exceeding a year. Cursory examinations
have shown that ova are usually numerous in mature individuals, insuring
the wide dispersal of larval stages, well beyond the limits in which the
species normally settles and matures. Nearly all can be expected to occur
in competition with others, where space, food, protection from predators
are factors determining survival and further reproduction. The fact
remains that most of the benthic stages of these species continue to exist
only in selected places. It must be assumed that these areas are previously
biologically conditioned by the predecessors of the same species.

This horizontal distributional pattern is closely linked to that of ver-
tical zonation, for it can be seen (see charts) that replacement of closely
related species, by depth, is a constant factor of these benthic animals.
The result is a stepping-stone effect, where greater depths result in a
dropping out of many species, and additions of others. This effect is illus-
trated not only for the polychaetes, but for echinoderms and mollusks,
and may extend to other well represented groups. It has also been demon-
strated for the faunas in the submarine canyons of southern California

(Hartman, 1963a).
ACKNOWLEDGEMENTS

Many scientists from the Allan Hancock Foundation of the Uni-
versity of Southern California participated in various aspects of this
study. The collections of samples were taken aboard the Research Vessel
VELERO IV, and processed aboard ship by members of the Hancock
staff. The tedious tasks of sorting and identifying the thousands of speci-
mens were performed by numerous specialists and assistants; to them
much gratitude is due for their patience and care. These assistants were
supported by the Allan Hancock Foundation, the California State Water
Pollution Control Board, and the National Science Foundation; the
administrations of these organizations are gratefully recognized for their
cooperation. The identifications of invertebrate specimens were under-
taken by various specialists (see Bibliography at end). The echinoderms

192 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

were identified by Mr. Fred Ziesenhenne; the mollusks were studied by
Mr. Gilbert F. Jones and the late Dr. Norman Mattox. Crustaceans
were identified or described by various specialists; Dr. J. Laurens Bar-
nard studied the amphipods, Mr. Robert Given the cumaceans, Dr. Rob-
ert Menzies some of the isopods, and Mr. William Banta the bryozoans
from the Farnsworth Bank. Professor K. O. Emery provided copies of
photographs from submarine areas off Santa Catalina Island. The
National Science Foundation, through grant no. B5-1780, supported
Mrs. Hannelore Paxton, who assisted in the organization and prepara-
tion of the manuscript, and Mr. Carl Petterson, illustrator, who prepared
the illustrations for plates 1 to 6, and arranged the photographs for plates
8 to 14. Mr. Anker Peterson prepared the figures on plate 7. Mrs.
Dorothy Halmos, Editor of the Hancock Publications, gave invaluable
aid in the final preparation of the manuscript. To all of these people and
institutions, | am most deeply indebted.

DESCRIPTIONS OF NEW AND LITTLE KNOWN
POLYCHAETOUS ANNELIDS FROM THE
SAN PEDRO REGION

‘The benthic samples taken from the San Pedro region were unusually
productive of polychaetous annelids, and resulted in the recovery of species
heretofore unknown. Some are here described.

Family POLY NOIDAE
Genus Gattyana McIntosh, 1900

Gattyana brunnea, new species
(Plate 1, Figs. 1-3)

Collection: Sta. 2154 (TYPE).

A small, linear, depressed species, it measures 15 mm long by 2 mm
wide with parapodia, and has 30 segments. Elytra number 15 pairs and
completely cover the dorsum. ‘The prostomium has two pairs of eyes; the
anterior ones are in front of midlength and at sides, and the posterior pair
nearer together, and at the posterior margin of the prostomium. FElytra
are subcircular (Fig. 1) in anteriormost segments, where they are large,
dusky brown and have a chalky white spot over the elytral scar; their
lateral margins are smooth in the first pairs, then have a sparse marginal
fringe at outer lateral edges, with the fringe in a single row; they are
never fimbriated as in other species of the genus. The upper surface of
elytra appears smooth but is sparsely covered with low, inconspicuous
surface nodules.

No. 2 HARTMAN : QUANTITATIVE SURVEY 193

Notosetae occur in small, close fascicles and do not extend to the
distal ends of neuropodial lobes. Notosetae are hairlike ; under high mag-
nification they show deep lateral serrations (Fig. 2). Neurosetae (Fig. 3)
are much coarser; each is distally entire, the tip long, smooth, slightly
falcate, and with four to seven rows of serrations subproximally.

Gattyana brunnea approaches G. ciliata Moore (1902, p. 263) from
which it differs in lacking heavy elytral fringe, and in the details of para-
podial setae.

Gattyana brunnea was taken only at Sixmile bank, in 300 fms, in
rocky bottom.

Genus Halosydna Kinberg, 1855
Halosydna latior Chamberlin, 1919

Flalosydna latior Chamberlin, 1919:1-2; Rioja, 1941 :680.

Collections: Many specimens come from shallow depths off southern
California, usually in a large gastropod shell occupied by a pagurid crab,
especially in shells of Polinices and Strombus occupied by Holopagurus
pilosus Holmes. ‘The commensal polynoid is usually paired, with the indi-
viduals near the anterior end of the crab. The body is greatly depressed,
unusually broad for a Halosydna; its length is 40 to 50 mm, its width
14 to 20 mm, and segments number 37. The 18 pairs of elytra are broadly
imbricated and completely cover the body. The prostomium has two pairs
of moderately large eyes, the anterior pair just in front of mid-length, the
posterior pair nearer together and near the posterior margin. Parapodia
are distinctly biramous with smaller notopodia and larger neuropodia.
Neurosetae are thick, distally falcate, the tip entire. Elytra are character-
istic; they have broad, smooth surfaces and enlarged, very flat, pale pus-
tules in an area behind the elytral scar.

Halosydna latior is recorded from southern California and western
Mexico, in shelf depths; its association with pagurid crabs is not previ-
ously mentioned.

Genus Harmothoe Kinberg, 1855
Harmothoe priops Hartman, 1961
(Plate 1, Figs. 4-6)
Harmothoe priops Hartman, 1961b:50-51; Hartman, 1963b:3.
Collections: Sta. 2314; Sta. 4886.

The body is about 7 mm long, 0.7 mm wide, and segments number
39. The dorsum is crossed by broad, reddish brown pigment bars which

194 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

are most intense between dorsal parapodial bases. The prostomium (Fig.
4) has distinct peaks and a deep median groove at the insertion of the
median antenna. The anterior prostomial eyes are under the frontal
peaks, and the posterior ones on the posterior third of the lobe; in addi-
tion, 10 to 20 irregularly dispersed small black spots are on the posterior
third of the lobe. Notosetae (Fig. 5) are of one kind; each is long,
slender, distally whiplike, and serrated along the cutting edge. Neuro-
setae are of two kinds; supra-acicular ones are similar to the notosetae;
subacicular setae are thicker and distally bifid (Fig. 6).

Harmothoe priops is known from southern California, in shelf, slope
and canyon depths, in silty mud bottoms.

Family HESIONIDAE
Genus Hesionura Hartmann-Schroeder, 1958

Hesionura coineaui difficilis (Banse, 1963)

Eteonides coineaui difficilis Banse, 1963:197-200, fig. 1; Hartmann-
Schroeder, 1963 :223-225, figs. 33-35.

Collection: Sta. 3595 (15*).

The body is small, slender, measures 3.5 to 5.6 mm long; its color is
translucent yellow, with a dark brown pharyngeal region. The everted
proboscis is dark brown, completely covered with coarse papillae in dis-
persed arrangement. The long trapezoidal prostomium has two pairs of
long, cirriform antennae inserted at the frontal margin, and two or three
small black eyespots near its posterior end. The first segment is a simple
ring with a pair of long, tentacular cirri; the second segment has a pair of
similar long, dorsal cirri and a pair of normal ventral cirri; the third
segment is the first setigerous and has embedded acicula. he formula

1 0
may be expressed thus: 1 + — + Sa—. Parapodia are uniramous,

N N

with smaller dorsal and larger ventral cirri. Setae are in small, fan-
shaped fascicles, number about five or six in a bundle; all are composite,
the appendage bladelike and ridged along its length.

Previously known only from San Juan Archipelago, Washington, in
20 and 100 m, from coarse, clean sand with broken shells, it is here

reported from Farnsworth Bank, outer side of Santa Catalina Island,
in 16 fms.

NO. 2 HARTMAN : QUANTITATIVE SURVEY 195

Family SY_LLIDAE
Genus Eusyllis Malmgren, 1867

Eusyllis transecta, new species
(Plate 2)
Collection: Sta. 2006 (12, TYPE).

The body is short, thick and fragile; it measures 6.1 mm long by
0.8 mm wide between segments 10-20, or at its widest part; it has 46
segments. The dorsum (preserved) is crossed by black bars, at segmental
grooves. Ihe prostomium (Fig. 1) is subquadrate, wider than long, has
four small eyes in trapezoidal arrangement, behind the insertion of the
paired antennae; the anterior, slightly larger pair are wider apart than
the posterior pair. Ihe median antenna is much the longest, inserted
behind the paired antennae; its length is about 2.5 times that of the
paired ones, which are about as long as the second dorsal cirri. Antennae,
dorsal and tentacular cirri appear smooth, but show weak annulations
under magnification.

Palpi are short, broad; the two are fused at the base; each is directed
forward for a length equal to that of the prostomium. The retracted
proboscis and proventriculus extend through segments | to 5 and 6 to 11
respectively. The dissected pharynx is cylindrical ; it terminates anteriorly
in a serrated margin (Fig. 2); its dorsal arc is slightly crenulated and
the ventral arc has 18 pointed teeth which increase in length midventrally.
A large, middorsal tooth (Fig. 3) is embedded in the pharyngeal tissue
some distance beyond the anterior edge (Fig. 3, seen in lateral view).

The first segment, or peristomium, slightly overlaps the prostomium
medially; its tentacular cirri are long, with the dorsal pair the longest,
exceeded only in length by the median prostomial antenna. The second
segment is the first parapodial one; its ventral cirri are cirriform, and
farther back they are short, triangular. All parapodia (Fig. 4) are lateral,
have long, pseudoarticulate dorsal cirri which diminish in length posteri-
orly, or alternate long and short in anterior segments. Ventral cirri, from
the second, are short, triangular, exceeded in length by the acicular lobes.
Anterior and median parapodia have setae of two kinds; the superior-
most, supra-acicular one is long, slender, distally pointed and laterally
slightly dentate (Fig. 5). Subacicular setae number 9 to 12 in a fascicle;
each is composite, distally falcate; anterior ones (Fig. 6) have a slightly
longer appendage than posterior ones, and are dentate along the cutting
edge; posterior setae have a shaft distally dentate and the appendage is
smooth (Fig. 7). Acicula occur singly in a ramus; each is straight and

196 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

distally slightly club-shaped (Fig. 8). Posterior segments have simple,
slender, superiormost setae, occurring singly in a fascicle ; each has widely
spaced denticles along the cutting edge. The body tapers posteriorly and
terminates in a pair of long, cirriform, pygidial appendages.

Eusyllis transecta differs from other species of the genus in its pharyn-
geal formula; it has falcigerous setae which are distally entire. It has been
recovered only from the San Pedro shelf, in 14 fms, where the trawl took
algae, sand, rock and ascidians.

Genus Sphaerosyllis Claparede, 1863

Sphaerosyllis californiensis, new species
(Plate 3)

Collections: Sta. 5028 (66, TYPE); Sta. 4806 (57); Sta. 5102
(100°).

‘The body is small, depressed, linear and trim in appearance. Length
is about 3.0 mm, width 0.5 mm, and segments number 32 to 34. The
dorsal surface is papillated and convex; the ventral surface is nearly
smooth and flat. The quadrate prostomium (Fig. 1) is wider than long,
has a pair of broad palpi completely fused, and a slight, midfrontal emar-
gination. he two pairs of eyes are in quadrate arrangement, with the
two on a side nearly touching; the anterior pair are slightly the larger.
Prostomial antennae resemble one another; all are short, clavate, their
length less than that of the prostomium; the median one is inserted
behind the middle of the lobe, and the paired ones are in front, within the
anterior eyes.

‘The proventriculus, seen through the body wall (Fig. 1), is short,
extends through setigerous segments 3 to 5, and is crossed by about 13 or
14 muscular bands. The anterior end of the pharynx terminates in 10
soft, widely spaced, small papillae ; a conspicuous, blunt yellow, middorsal
tooth is embedded in the pharyngeal wall.

‘The first segment is reduced to a lower lip or peristomium and a pair
of bases for the tentacular cirri, best seen in ventral view; they project
laterally, and in dorsal view are seen in line with the prostomial eyes.
‘These cirri, and all other dorsal cirri are clavate, with a thick, bulbous
base and a terminal digitate lobe exceeding in length that of the base.
‘The second parapodia lack dorsal cirri, but all others have well developed
ones.

Parapodia (Fig. 3) are similar throughout, but diminish in size poste-
riorly. Dorsal and ventral cirri are clavate, and taper distally. A superior-
most, nearly straight acicular spinelike seta is present from the first, and

NO. 2 HARTMAN : QUANTITATIVE SURVEY 197

continues in posterior segments. This is followed inferiorly by three or
four composite falcigers in which the appendage is longest (Fig. 4) in
anteriormost segments, and diminishes in proportionate length farther
back (Fig. 5) ; the distal end is falcate; the cutting edge has a few long,
slender teeth in a single row. Each of the last few segments has, in addi-
tion, an inferiormost simple, slightly curved, spinelike seta (Fig. 6)
directed ventrolaterally. Acicula occur singly in a ramus; each is a slender
rod, distally club-shaped (Fig. 7).

The posterior end of the body terminates in a pair of short, laterally
directed, clavate processes (Fig. 2) resembling the dorsal cirri.

Sphaerosyllis californiensis is characterized as follows: the epithelium
is papillated ; the first segment is reduced to a lower lip and a pair of ten-
tacular bases, lying in line with the prostomial eyes; parapodia have
simple setae in superiormost and inferiormost positions, and composite
falcigers; acicula occur singly. It is unique for having the paired pros-
tomial antennae inserted far forward, and the median one near the pos-
terior end of the lobe. Composite falcigers have a few long teeth limited
to the basal half of the appendage.

Sphaerosyllis californiensis occurs in San Pedro shelf, in silt and

mixed sediments.

Family QUESTIDAE, new family
Genus Questa, new genus
Genotype: Questa caudicirra, new species

The body is long, linear and consists of many segments. Ihe prosto-
mium is a simple, triangular lobe without appendages. The first segment
or peristomium is a smooth ring which is somewhat biannulated ; all other
segments have lateral, biramous parapodia in which notopodia and neuro-
podia are ventrolateral in median and posterior segments. Setae are sim-
ple, of two kinds, including slender, distally tapering ones and shorter,
thicker, distally bifid hooks without a hood. he pygidium terminates in
an anal pore bounded by paired cirri. The alimentary tract is simple,
linear, with a pharyngeal region in the first four or five segments without
jaws or other hard parts. A single species is known.

Questa caudicirra, new species
(Plate 4)
Collections: Sta. 3595° (20%, TYPE) 3 Sta. 2298 .(1).

The body is long, linear, measures 9 to 10 mm long by 0.275 mm
wide and consists of 50 to 58 segments. All segments are smooth, plain,

198 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

with inconspicuous biramous parapodia (Fig. 1) from which the setae
project in sparse fascicles; the last 9 to 15 segments have long, dorsally
inserted branchiae. The posterior anal pore is surrounded by a pair of
dorsolaterally and similar, slightly longer ventrolaterally inserted cirri-
form processes (Fig. 2). The prostomium (Fig. 3) is a simple, depressed,
bluntly triangular lobe without eyes. The first ring is smooth, or some-
what biannulate; all other segments are simple, uniannulate, with bira-
mous parapodia. The first three segments have only slender, distally
pointed setae in notopodia and neuropodia. The fourth segment has simi-
lar setae accompanied by thicker, distally falcate hooks with bifid tip;
these hooks usually number one in a fascicle (Fig. +), but an occasional
one has two or three hooks, all of one kind. Numbers of setae and hooks
in fascicles remain about the same except in posteriormost, branchial seg-
ments where setae are less conspicuous.

Each long seta (Fig. 5) is serrated along the cutting edge and its tip
is pointed. The hook consists of a straight, embedded shaft, a slightly
curved free region, and a broader, laterally directed fang, nearly at right
angles to the shaft (Fig. 6) ; the accessory tooth is small and distal. The
alimentary tract of some specimens contains white calcareous ooze, such
as the sediments in the sample from which they were removed.

These small, linear worms may be allied to the PARAONIDAE,
with which they agree in their small size, the presence of two kinds of
setae, and the presence of a well developed, simple prostomium. They
were at first thought to have affinities with oligochaetes, but this appears
unlikely in view of the characteristics named.

Questa caudicirra was taken only on Farnsworth bank, Santa Cata-
lina Island in 16 fms, and on Lasuen sea mount, in 68 fms.

Family CIRRATULIDAE
Genus Cirratulus Lamarck, 1801
Cirratulus, unknown species
Collection: Sta. 2737 (2).

These individuals differ from known species of the genus, and may
represent an undescribed species. Length of the body is more than 30 mm,
width 3 to 4.5 mm, and segments number more than 67 (the tail is
lacking). The body is inflated through setigers 10 to 25, then depressed
cylindrical. Segments are uniannulate, wider than long, smooth, their
width up to 25 times that of their length in front, but only 1.5 times as
wide as long in middle and posterior segments. Setae are long, silky,

NO. 2 HARTMAN : QUANTITATIVE SURVEY 199

laterally directed, their extended length 3.5 to 5 times as long as the body
is wide. The prostomium is broad, thick, nearly semicircular; eyes have
not been identified. The buccal region is triannulate, with each ring
smooth, and longer than the prostomium or any body segment.

The first pair of branchiae are on the posterior margin of the third
buccal segment. Branchiae are inserted immediately above the notopodial
ridge, in median and posterior segments. Most branchiae have been lost
from the specimen, so that it looks rather like a spindle-shaped oligo-
chaete. All setae in anteriormost segments are slender and capillary.
Thick, short, acicular spines are first present in neuropodia 32, or behind
the anterior, inflated region; they number only one or two in a fascicle,
and are located at the lower end of the fascicle; they are accompanied
by slenderer, longer capillary setae. Similar but slenderer acicular spines
also occur with long, slender, capillary setae. These spines are translucent
yellow, nearly straight, taper to blunt points, and never number more
than two or three in a fascicle, where they are accompanied by capillary
setae.

This Cirratulus sp. has been taken only from West End, Santa Cata-
lina Island, in 256 fms.

Family SCALIBREGMIDAE
Genus Sclerocheilus Grube, 1863

Sclerocheilus acirratus, new species
(Plate 5, Figs. 1-5)
Collections: Sta. 1370 (4, TYPE) ; Sta. 1378 (2), White Cove, in
Eisenia holdfasts (2).

Small individuals, yellow-orange in life, fading to colorless in alcohol,
measure about 8.6 mm long by 1.4 mm wide near the middle of the body ;
they consist of 31 to 50 segments. The surface epithelium is rugose, with
transverse annulations, or papillate, coarsest on the dorsum and less so
ventrally. Median and posterior segments are quadri-annulated, with the
rings equally long and most distinct in middle segments. Branchiae are
absent. The prostomium is broadly T-shaped, with the lateral extensions
continuous with the prostomium (Fig. 1). Two pairs of transverse red-
dish eye patches are conspicuous on its posterior half, with the two patches
on a side near together, and the anterior ones the larger. The first seg-
ment is a short, smooth ring. The second, or first setigerous, segment is
about twice as long as the first one; it has biramous parapodia with noto-
podia dorsolateral, and neuropodia lateral.

200 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Notosetae of the first setiger are of two kinds; an anterior transverse
row of five or six, coarse, spinelike setae (Fig. 2) is located in front of a
posterior row of eight or nine slenderer, capillary ones (Fig. 3). Uhe
corresponding neurosetae are slender, capillary only, as in all successive
parapodia. Median and posterior parapodia have, in addition, shorter,
furcate setae in which the tines are spreading (Fig. +) and the inner
margins are serrated (Fig. 5).

Parapodia are short and truncate throughout; the setae emerge in
front of short, postsetal lobes. Cirri are altogether absent, hence the spe-
cific name. The pygidium is a smooth ring without cirri.

Sclerocheilus acirratus differs from other species of the genus in lack-
ing cirri. It may approach S. minutus Grube, from the Mediterranean
Sea (Fauvel, 1927, p. 125), in having a single anterior segment with
acicular setae. It differs from S. minutus in that these acicular spines are
distally prolonged to slender tips, and the prostomium has four, instead of
two eyes. [he frontal extensions of the prostomium are continuous with
the lobe, not set off by constrictions.

S. acirratus has been taken only in White Cove, Santa Catalina
Island, in shallow depths, from holdfasts of Eisenia arborea (kelp). Fhe
species was first identified from samples taken by the VELERO III
(See Fraser, 1943, for Station data), in holdfasts of this kelp. The same
locality was investigated by Mr. Robert Given, where the same species
was again located. It has been found nowhere else.

Genus Asclerocheilus Ashworth, 1901

Asclerocheilus californicus Hartman, 1963
(Plate 5, Figs. 6-9)
Asclerocheilus californicus Hartman, 1963b:56-57.

‘The body is long, arenicoliform, measures +3 mm long by 6 mm wide,
and consists of more than 70 segments. The prostomium is triangular,
broadest behind, and has a pair of thick, short, frontal processes resem-
bling a pair of short antennae (Fig. 6); eyes are absent. ‘Ihe posterior
end terminates in a pygidium surrounded by five short, tapering cirri;
one is midventral and the others are lateral; the anal aperture is cen-
tered. Dorsal and ventral cirri are absent. Posterior parapodia are char-
acterized by the great prolongation of superior and inferior lobes (Fig.
7) resembling dorsal and ventral cirri; they are first present from neuro-
podium six and notopodium seven, and continue long farther back. The

first two setigerous segments have thick, acicular curved spines (Fig. 8)

NO. 2 HARTMAN : QUANTITATIVE SURVEY 201

in an anterior series, accompanied by longer, slenderer ones in a posterior
row. Farther back, notopodia have only slender setae, and from about
segment 20 they are accompanied by fewer, shorter furcate spines (Fig.
9). Neurosetae are similar to notosetae but shorter.

Asclerocheilus californicus is characterized by its greatly prolonged
parapodial lobes in posterior parapodia. It is known from slope and can-
yon depths of southern California, in mud.

Asclerocheilus, unknown sp.
Collection: Sta. 3616 (2).

The larger specimen, in two pieces, measures 14+ mm long by 1.6 mm
wide in the thoracic or widest region. Segments number at least 62. The
body is broadest and conspicuously rugose in the anterior third, and it
tapers posteriorly, where the epithelium is less roughened. The pros-
tomium is I-shaped, widest in front and laterally prolonged as a pair of
horns which are continuous with the prostomium; a pair of conspicuous
ocular patches occupies much of its first half, nearly meeting medially.
The prostomium is posteriorly slightly overhung by the first short seg-
ment, a plain, incomplete ring surrounding the lobe only middorsally and
laterally; the second segment is the first complete ring and its ventral
part forms the lower lip.

The first two setigerous segments have thick, curved acicular spines, in
an anterior series, accompanied by a posterior series of capillary setae,
present in both notopodia and neuropodia. From the third segment the
setae are smooth, long, capillary, and accompanied by conspicuous series
of furcate setae, in both notopodia and neuropodia. Each furca has un-
equally long tines, with long, slender spinelike teeth along the inner cut-
ting edges. Dorsal and ventral cirri, as well as branchiae, are totally lack-
ing. Parapodia are clearly biramous, with notopodia and neuropodia rep-
resented by low, broad lobes, from which the setae project; the superior
bases of setal fascicles are bounded by pale, glandular ridges. The pygid-
lum terminates in 2 longer and 2 shorter, cirriform processes, directed
laterally.

‘These specimens differ from known species of the genus in having
ocular areas on the prostomium (see Uschakov, 1965 :292).

‘The species has been taken only off East End, Santa Catalina Island,

in 88 fms, in glauconitic sand.

202 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Family SABELLIDAE
Genus Euchone Malmgren, 1866

Euchone arenae, new species
(Plate 6, Figs. 1-6)
Collections: Sta. 2788 (23, TYPE) ; Sta. 2417 (2).

The body is short, thick, tumid and tapers posteriorly. It measures
3.8 mm long without, and 6.2 mm long with, the tentacular crown;
width is 0.7 mm. Segments include 8 thoracic and 13 to 15 abdominal
setigers ; each is biannulate. A narrow, glistening white band, behind the
second setigerous fascicle, surrounds the body. The tentacular crown is
well developed ; it consists of five to seven pairs of radioles, and two pairs
of long slender filaments at the ventral ends of the radiolar base (of
which only one is shown in Fig. 1). Peristomial eyes are not visible, but
may be faded. A palmate membrane comprises less than a fourth of the
radiolar length; it is inconspicuous. A pair of reddish eyes is visible in
the pygidium, in some individuals.

The thoracic collar is continuous all around and of uniform length; it
is deeply cleft middorsally, where it joins the thoracic segment; a slight
midventral cleft is distinguishable.

The collar segment resembles those farther back but lacks neuro-
setae. Its notosetae are slender, limbate and number four or five in a
fascicle. More posterior thoracic notopodia have setae of two kinds;
three or four in superior position are long, limbate (Fig. 2), and four
to six are short, subspatulate (Fig. 3). The corresponding neuropodia
have long-handled uncini, in which the distal end is a large fang (Fig.
4+) surmounted by three smaller teeth in a row.

Abdominal parapodia have thick, short uncini in notopodia; they
number six to eight in a row; each has a thick, broad base, a large curved
fang surmounted by three or four rows of small denticles, in transverse
rows (Fig. 5). The corresponding neuropodia have capillary setae, num-
bering three to five in a fascicle.

‘The posterior end terminates in an anal groove (Fig. 6) in which the
last six setigerous segments are involved; this may be broadly open, as
shown in the figure, or somewhat curled inward. The groove is followed
by a small triangular pygidium with dorsal anal pore.

Euchone arenae differs from known species of the genus in having
six setigerous segments in the anal groove. Thoracic notosetae are essen-
tially of one kind, with the superior longer and less curved but otherwise
similar to the inferior shorter ones.

NO. 2 HARTMAN : QUANTITATIVE SURVEY 203

Euchone arenae has been taken near Redondo Beach, in 9.5 fms, off
Point Fermin, in 34 fms, and off Point Loma light, in 23 fms, in sedi-
ments of reddish brown sand.

Euchone incolor Hartman, 1965
Euchone incolor Hartman, 1965 :231-232, pl. 51.
Collections: Sta. 2298 (15) ; Sta. 3585 (3).

Total length, excluding the tentacular crown, is 2.1 to 2.2 mm;
width is 0.3 mm; segments include 8 thoracic and 8 abdominal setigers.
The body is slender and linear. The flaring tentacular crown consists of
four pairs of radioles, each with paired rows of pinnules; they extend
distally for a length nearly one-fourth that of the body. A pair of slender
cirriform radioles is present at the ventralmost end of the crown. The
thoracic collar is large, voluminous and covers most of the peristomium ;
it has a straight, continuous margin, and is incised middorsally. Eyes have
not been distinguished. The anal groove extends through the last three
segments. Collar setae are slender, limbate and number about five in a
fascicle. Other thoracic notosetae are slender and limbate; neurosetae are
long-handled uncini with a large fang, surmounted by three smaller teeth.

Previously known from the western Atlantic Ocean, in shelf to
abyssal depths (Hartman, 1965, p. 232), this is here recorded from the
Lasuen sea mount, in 68 fms, and off Farnsworth bank, in 23 fms.

Euchone limicola Reish, 1959, redescribed
(Plate 6, Figs. 7-10)
Euchone limicola Reish, 1959 :717-719, figs. 1-5.

Collection: Type collection from Alamitos Bay marina, and Long
Beach outer harbor.

Length of the body is 8 mm of which the crown measures 2 mm; seg-
ments include 8 thoracic and 18 abdominal, of which the last 10 setigers
are involved in the anal groove (Fig. 10). The tentacular crown consists
of seven pairs of radioles, each with many filaments in pairs; the palmate
membrane is very low and nearly lacking. Ventral scutes or glandular
areas continue through seven thoracic setigers; the last one is least devel-
oped, and all are transversely divided, in line with the biannulation of the
thoracic segments. Thoracic notosetae are of two kinds; the superior ones
are longest and slenderly limbate; the inferior ones are abruptly shorter,
and broadly limbate (Fig. 7). Abdominal uncini are avicular, with short,
thick base, large, curved fang surmounted by many teeth in four to six
transverse rows (Figs. 8, 9), each row with four to seven teeth.

204 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Euchone limicola is an estuarine form, limited to harbor areas, in
sandy mud.

Family SERPULIDAE
Genus Spirobranchus Blainville, 1818

Spirobranchus spinosus Moore, 1923
(Plate 7)

Spirabranchus spinosus Moore, 1923 :248-250, pl. 18, fig. 47.
Collections: Isthmus Cove, Santa Catalina Island.

This brilliantly multi-colored serpulid is one of the conspicuous en-
crusting forms along the leeward side of Santa Catalina Island, and along
outer, rocky insular coves off southern California. Its hard, stony, white
tubes encrust rocky walls at and below intertidal levels. When the ani-
mals are extended, they present a variety of red and white banded, or
black, blue and brown banded crowns. A slight disturbance at the surface
will cause the entire colony to withdraw, revealing the purple-white
tubes; these are longitudinally ridged and often diffused with red and
blue pigments along the ridges.

The only known account of this species was made from a single
imperfect specimen, 23 mm long, taken off Santa Barbara Island, in 29
fms, in fine yellow sand and coralline rock (possibly only from the coral-
line rock). Because of its abundance at Santa Catalina Island, the species
is here more fully described. The body of adult specimens measures 20 to
25 mm long, the radioles being 6.2 mm long. Width in the thorax is 2.5
mm and in the collar region + to 5 mm. The tentacular crown has paired,
inrolled halves; each consists of +45 to 50 radioles, coiled in about three
spirals, and the lateral pinnae are paired with the longest outermost and
the shortest innermost and most distal. The radioles are basally united by
a palmate membrane for about two-fifths of the total length. The oper-
cular stalk is unpaired (Fig. 1), formed of the dorsalmost left radiole ;
it terminates in a broad, circular, calcareous disk which extends distally
to near the ends of the radioles; the disk has a pair of smaller inner
branched spines (Fig. 2), and a pair of larger outer branched spines.
The pedicle is cylindrical for half its basal length, then has a pair of
broadly expanded lateral wings which extend distally to near the base of
the distal disk. The operculum is sometimes partly overgrown with other
organisms, especially bryozoans, spirorbids, sponge masses, etc.

The collar membrane is large, consists of a pair of long, broad, dor-
solateral lobes, a pair of much smaller lateral lobes, and a pair of large

Gn

NO. 2 HARTMAN : QUANTITATIVE SURVEY 20

ventral lobes resembling the dorsal lobes. In damaged, or regenerated
specimens these lobes may be otherwise. Ihe thoracic membrane is re-
duced. The first, or collar, fascicle emerges from the dorsalmost part of
the collar membrane; its setae are in thick, projecting tufts; each seta is
simple and bayonet-shaped. Segments two to five have smaller notoseta
fascicles and neurosetae in transverse, linear ridges; the uncini form a
single row of many flat platelets; each uncinus (Fig. 3) consists of a thin
rectangular base with a series of twelve teeth along the cutting edge.

‘The thorax is separated from the abdomen by a long, smooth, apodous
region. Abdominal segments are very short, appear crowded, have short
uncinial ridges and inconspicuous setal tufts. Abdominal uncini are
smaller than thoracic uncini, and each has about seven marginal teeth.
‘Tubes are fully attached to the substratum; they measure about 7 mm
across, with the lumen only about 2.5 mm across. Mature females are
bright orange red, males pale; the opercular stalk is deep maroon to
brown, splashed with white.

Distribution: Long-shore islands of southern California, on rocky
surfaces in low intertidal or littoral zones.

ANALYSES, BY AREAS

The San Pedro region is divided into areas which are believed to have
unique biological characteristics. Nearly a third of the region is taken
over by the San Pedro Basin (Hartman and Barnard, 1958, 1960). Sub-
marine canyons include the San Pedro sea valley, the Newport canyon
(Hartman, 1963), and the San Gabriel sea valley (see Map 1). The

following areas are here further identified :

LH refers to the Los Angeles-Long Beach outer harbor, a shallow
enclosure bounded on the east by the mainland, and on the west by an
artificial breakwater.

PV refers to the Palos Verdes shelf and slope, bordered on the east
by the Redondo canyon, on the west by the sill of the San Pedro Basin,
and on the south by the San Pedro sea valley. It is the site of an important
outfall line for Los Angeles County.

SP refers to the San Pedro shelf and slope; it is bordered on the east
by the Los Angeles-Long Beach breakwaters, on the south by the New-
port canyon, on the north by the Palos Verdes shelf and slope; on the
west it surrounds the Lasuen sea mount and Sixmile bank, then continues
seaward to the Catalina rise.

206 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

NE refers to the Newport shelf, bounded on the east by Newport
Bay, on the north by Newport canyon, and on the west by the San Pedro
slope ; it extends oceanward as part of the San Pedro slope.

LS refers to the Lasuen seamount, located beyond the outer end of
Newport canyon, in depths of 58 to 385 fms, and to Sixmile bank, ap-
proximately six miles northeast of Avalon, Santa Catalina Island.

CL refers to the leeward side of Santa Catalina Island. A narrow,
long shore shelf borders the island, from East End at the southeast, to
West End at the northwest ; oceanward the shelf gives rise to the slope
which merges with the San Pedro basin (Hartman and Barnard, 1958,
1960).

CW refers to the windward side of Santa Catalina Island. The nar-
row, long shore shelf extends westward as a slope, offshore rocks and
banks; it is dissected by the Catalina canyon, and in greater depths it
merges with the Catalina basin.

Each of these areas has characteristic associations of organisms, for
which the details are explained below.

An inshore area, in 2-5 fm depths, can be recognized as having some
species characteristic to it. Analyses for this area have been previously

published (California, 1965b, pp. 302-318).

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NO. HARTMAN : QUANTITATIVE SURVEY 209

LOS ANGELES-LONG BEACH OUTER HARBOR
The Los Angeles-Long Beach outer harbor (LH) is a shallow, four

to nine fathom deep, artificially constructed basin, located between Ter-
minal Island and the San Pedro shelf. It is separated from the open sea
by a breakwater with openings which allow ships and seawater to move
freely in and out of the basin. Sediments in the bottom are mainly silty
to sticky muds and rubbly shales, except along the shipping lanes, where
hard sandy bottoms prevail. Benthic animal populations, taken with the
grab, are predominantly annelids of many kinds and in moderate to high
numbers; they are accompanied by much smaller numbers of mollusks,
crustaceans, and sparse numbers of echinoderms. They are the same kinds
as those occurring more abundantly on the San Pedro shelf where the
quality of the sediments and proximity to food supplies are similar.

The outer harbor receives much organic and waste materials from
the drainage of the Los Angeles and San Gabriel rivers. The effects of
pollution on bottom faunas in Alamitos Bay, an arm of the San Gabriel
River, have been studied by Reish (1955); the kinds and numbers of
animals are much like those in the outer harbor or the shallow, sandy
shelf (California, 1959). The fauna is fairly uniform throughout, de-
pending on whether sediments are chiefly mud, clay or sand.

Seven samples were analyzed:

1.1 Sta. 6107 (near no. 29), in 4 fms. OPG took 2.96 cuft of smooth,
black silty mud. Wet weights measured 28 grams, of which polychaetes
were 14.7, mollusks 5.4, enteropneusts 4.2 and pinnotherid crabs 3.7
grams. In addition to those listed on the chart, a cumacean (1), a pinno-
therid crab (17), and an enteropneust (6) were present. Largest species
were Glycera americana, an enteropneust, and MJarphysa disjuncta. Most
abundant were Ancistrosyllis tentaculata (169), Haploscoloplos elon-

gatus (50), and Cossura candida (35).
2. Sta. 2314 (no. 32), in 4.5 fms. OPG took 0.56 cuft of mud, silt

and many animals. A sea whip (1), ?/arenactis (24), two kinds of
nemerteans (10), amphipods (1), isopod (1), cumaceans (5), ostracods
(9), and a pinnixid (7), accompanied the species named on the chart.
Largest individuals were Asychis sp. and Praxillella affinis pacifica; most
abundant were Haploscoloplos elongatus (82), Lumbrineris spp. (61),
Tharyx tesselata (52), and Nereis procera (41).

1JIn all charts, below, the first numbers (i.e. 1-7) refer to those used in the
chart-headings. Station numbers (1370 to 7726) are given in the LIST OF

STATIONS. The serial numbers, 11 to 267, refer to locations on the grid, vol. 19,
pt. 1, p. 6).

210 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 46 species, 512 specimens
echinoderms 3 $5)
mollusks Y) 24
crustaceans 6+ 32
others 5 44

Total: 69+ species, 647 specimens

3. Sta. 5808 (near no. 30), in 5.8 fms. OPG took 2.52 cuft of dark
olive-green silt with many animals. Wet weights totalled 87.6 grams, of
which polychaetes comprised 64.1, mollusks 17.5, nemerteans 4.5, and
anemones 1.5 grams. A ceriantharian with dark purple stalk and red
crown (1), a small anemone (3), Cerebratulus rubra (1), a small
nemertean (3), a phoronid (2), and an ostracod (3) accompanied the
species named on the chart. Largest individuals were Solen sicarius and
Cerebratulus rubra. Most conspicuous were Diopatra tridentata and
Compsomyax subdiaphana, and most abundant were Marphysa disjuncta
(57), Tharyx tesselata (55), Pectinaria californiensis (34), and Chaeto-
zone corona (32).

Numbers of species and specimens totalled :

polychaetes 33 species, 328 specimens
echinoderms 1 3)
mollusks 25 82
crustaceans 1 2,
others 5 10

Total: 65 species, 425 specimens

4. Sta. 2508 (no. 29), in 6 fms. OPG took 2.56 cuft of black mud
and clay with many animals. A sea-whip (2), ?Harenactis (1), a nemer-
tean (3), an amphipod (5), a pinnotherid (1), were present in addition
to those listed on the chart. Largest individuals were Marphysa dis-

juncta and Glycera americana. Most abundant species were Tharyx tes-
selata (301), Cossura candida (24) and Lumbrineris ?zonata (20).

NO. 2 HARTMAN : QUANTITATIVE SURVEY 211

Numbers of species and specimens totalled :

polychaetes 28 species, 465 specimens
echinoderms ] 1
mollusks 1 2
crustaceans 2 6
others 3 6

Total: 35 species, 480 specimens

5. Sta. 4718 (near no. 33), in 6 fms. OPG took 0.5 cuft of coarse
greenish gray sand with much flocculent debris, white branching bryo-
zoans and many animals. In addition to species named in the chart, there
were a ceriantharian (1), ?Harenactis (2), a sea-whip (1), a nemer-
tean (4), a sipunculid (2), Glottidia albida (1), + kinds of amphipods
(8), 2 isopods (4), 4 cumaceans (5), an ostracod (1), a cancroid crab
(1), a pycnogonid (5), and an enteropneust (1). Largest species were
Pista disjuncta and a sea-whip; most abundant were Tharyx tesselata
(104) and Mediomastus californiensis (16).

Numbers of species and specimens totalled :

polychaetes 57 species, 354 specimens
echinoderms S 5
mollusks 17. 46
crustaceans 11 15
others 8 12

Total: 96 species, 432 specimens

6. Sta. 2307 (no. 29), in 7 fms. OPG took 2.14,cuft of shale, black,
friable, rubbly clay and many animals. In addition to species named in the
chart, there were ?Harenactis (2), 2 nemerteans (6), a sipunculid (1),
an amphipod (5), a pinnixid crab (1). There were no large individuals.
Most abundant were Cossura candida (250*), Nereis procera (145),
Tharyx tesselata (more than 100), and Paraonis gracilis (92).

Numbers of species and specimens totalled :

polychaetes +9 species, 846 specimens
echinoderms 1 +

mollusks 9 4]
crustaceans Y 6

others 5 10

Total: 66 species, 905*specimens

212 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

7. Sta. 2507 (no. 30), in 8 fms. OPG was filled with black mud,
shelly debris and many animals. A large ceriantharian, a nemertean (2),
an amphipod (5), a cumacean (2), an ostracod (28), a commensal crab
(5), Callianassa (1), a pycnogonid (1) and an enteropneust (1) were
present, in addition to those listed on the chart. Largest individuals were
Cerebratulus rubra and Glycera americana. Most numerous species were
Marphysa disjuncta, Pectinaria californiensis and Cossura candida, each
with 10 individuals.

Numbers of species and specimens totalled :

polychaetes 25 species, 95 specimens
echinoderms 1 6
mollusks 11 20
crustaceans 5 4]
others 5 5

Total: 47 species, 174 specimens

HARTMAN : QUANTITATIVE SURVEY

Polychaeta in the Los Angeles-Long Beach Outer Harbor,

showing order of occurrence in 4 to 8 fms.
+ indicates presence, - absence.
Species represented by more than 10 specimens are indicated by

Ancistrosyllis tentaculata

Chaetozone corona : 5
Chone mollis . ¢ 2
Cossura candida . c c
Dorvillea articulata . 5
Drilonereis sp. . : :
Eteone ?alba . 5 : C

Glycera americana : ,

Gyptis a. glabra . : :
Haploscoloplos elongatus .

Harmothoe lunulata
Loandalia fauveli 5 é
Lumbrineris cruzensis c
Lumbrineris pallida . c
Magelona sacculata 5

Marphysa disjuncta : :

Mediomastus californiensis

Notomastus tenuis - :
Paraonis gracilis : :
Pectinaria californiensis
Pherusa neopapillata . A
Pilargis hamatus . : :
Polydora sp. . ° ° °

Prionospio cirrifera . =
Telepsavus costarum .
Tharyx tesselata . A :
Amphicteis scaphobranchiata
Anaitides sp., checkered

Anotomastus gordiodes

ab

a
*

2

3

+

m1

+

5

+

213

214 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Polychaeta in Los Angeles-Long Beach Outer Harbor (continued)

1 2 3 4 5 6

Aricidea lopezi . A : 6 A ‘ Hy : + = 2 + A
Aricidea, other sp. . 5 : : 5 : : + - - - =
Asychis sp. . : : : S c 6 5 + = = + =
Brada pilosa . : : 5 : : : 5 : x - - - =
Cirrophorus furcatus . : A é . ‘ 3 + = = + =
Glycera ?capitata 6 , 0 ; : 5 + ~ - = =
Goniada brunnea . c 5 c : 0 5 é + - - + =
Goniada littorea . é ; A : 6 : + - - = =
Laonice cirrata . ° : : : 3 : 5 * + + + +
Lumbrineris californiensis : ° ‘; F 3 ae es = + +
Lumbrineris limicola . 6 5 Ps 5 5 5 * % - = +

Pd

Lumbrineris minima - 3 3 - 5 é : - - = =

ba

Nephtys cornuta . C ° 5 5 . - - - -

Nereis procera : ‘ . ‘ ; ; z : * + + + *

Nothria iridescens 0 : 5S 6 5 $ a + - - + =
Pholoe glabra 0 . : C : . 5 : + - + - -
Pista cf. cristata : : 6 6 ; 5 : + - - - +
Polydora ?limicola ; : é A 5 6 : + - - - -

Praxillella a. pacifica . 4 . : . 5 + ~ + - -

Prionospio malmgreni . : 5 3 A 5 3 + = = + +
Prionospio pinnata 5 5 3 3 5 5 ; * + + + +
Scalibregma inflatum . 5 5 A ; : C + - + + -
Sphaerosyllis sp. : 3 5 0 3 : . + - - - -
Spiophanes fimbriata . - c ® : 6 5 + - + - -
Spiophanes missionensis . 5 6 5 5 6 + + = + +
Sternaspis fossor 5 5 2 5 : ; C + - ~ + -

Sthenelanella uniformis . 5 5 : 5 & + + - + +
Streblosoma sp. . C a A : : 5 ° + - - - -
Terebellides stroemii ; : . ; 5 : + - + - ~

Ampharete sp. 5 . ° c ° ° : . + = = =

Boccardia basSilaria . : - é 3 b : : + - - -

No. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta in Los Angeles-Long Beach Outer

Diopatra tridentata . ‘ .

Harmothoe priops . 5 ; :

Marphysa disjuncta : 7
Ninoe gemmea . C - S é
onuphid . : 2 6 3 2

Paraonis gracilis : 3 :
phyllodocid . ° : é
Pista disjuncta . : $ &
Poecilochaetus johnsoni . ;
Streblosoma crassibranchia

Tharyx monilaris . : 5 ;

Ampharete labrops 5 i 5
Diopatra ornata . : n 5
Drilonereis ?longa “
Lumbrineris ?zonata . : 5
Nephtys ferruginea 5 9
Polycirrus sp. 4 > 0 a
Amaeana occidentalis . : :

Anaitides, sp. B [trilineate]

Arabella semimaculata 5 5
Aricidea neosuecica . 5
Boccardia sp. Fe : ‘ :
Chone gracilis : : : 3
Cirriformia spirabrancha . :
Eteone californica : 5
Eteone ?dilatae . A :
Eumida bifoliata . * c 5
Maldane sarsi ; : 5

Melinna denticulata . y

Nephtys caecoides : 5

Owenia f. collaris : : :

i

Harbor (continued)

2

3
+

+

*

4

5

216 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta in Los Angeles-Long Beach Outer

Pherusa capulata . : 5 5
Pilargis maculata 5 d .
Prionospio pygmaeus . 5 6
Schistocomus hiltoni . . :

Ancistrosyllis sp. : ° S

Armandia bioculata 5 5 5
Boccardia nr. redeki . 5 5
Carazzia sp. . 6 6 6 5
Cirratulus cirratus . : :
Cirriformia luxuriosa : 5

Diopatra tridentata . ; ;

Dorvillea articulata . 6 6
?Fabricia sp. 0 6 C

Megalomma sp. ° é . °
Melinna sp. . C 5 C :
Nephtys sp. . 0 6 5 9

Ophiodromus pugettensis 5 ‘
Paraonis gracilis A ; :

Peisidice aspera . : 6 S
Polydora nr. armata . : C
Polydora citrona . : : 6
Polydora ligni ; ; 6
Travisia sp. . é : 5 ‘

Drilonereis cf. nuda. , E

Magelona sp. . c . 5 °
Pilargis berkeleyi S 5

Echinodermata in the Los
showing order of

Dendraster excentricus ; ;

Amphiodia urtica . ; A ‘

Pentamera pseudopopulifera 2

it

VOL. 19

Harbor (continued)

2

3

4

5

“
*

Angeles-Long Beach Outer Harbor,

occurrence in 4 to 8 fms.

NO. 2 HARTMAN : QUANTITATIVE SURVEY

217

Echinodermata in Los Angeles-Long Beach Outer Harbor (continued)

ophiuroid * : ° . . ° 5 . : ° : + -
Amphiodia occidentalis = - : 5 : C 5 S A +
Pachythyone rubra . . . . . . . . ° ° +

Amphiodia digitata s : 5 ; C : : : : : P

Mollusca in the Los Angeles-Long Beach Outer Harbor,
showing order of occurrence in 4 to 8 fms.

Asthenothaerus villosior . : : 0 : + - - - -
Chione undatella . : S 5 : 5 . + - - - -
Compsomyax subdiaphana : C . 8 c t + + - -
Cylichna attonsa . ; & : . 6 6 + - - - -
Lucinisca nuttalli 5 5 : C . + - + - +
Macoma yoldiformis 4 : 0 5 6 é + + + - +
Nassarius cooperi : : 5 ; : 5 + - tc - -
Nassarius perpinguis . ‘ . ‘ 6 : + = a = +
Nuculana taphria . . 3 5 co 6 5 + = = = =
Olivella baetica . 6 5 5 c ; + - + - -
Rochefortia sp. . 6 é : 2 5 + - ar - -
Thyasira sp. . : : 5 a 0 : C + - + - -
Turbonilla sp. s 5 5 5 : ; ° + + - +
Aglaja sp. é 5 . é < ‘ a + - = =
Balcis rutila s : 5 5 : 9 5 + - ~ -
Cadulus fusiformis 2 5 3 Cc ‘ - ; + = = =
Cylichna diegensis 5 5 $ . 3 ; . + - - -
Lyonsia californica . ‘ 5 A , A it - ~ +
Tellina idae . ‘ ; 5 . . 3 ‘ . + - - =
Aceton punctocoelata . c A C ° . ° ° a - *
Adontorhina cyclia 2 5 : 5 S : . + - -
Cadulus sp. . - 5 : : 5 E : ° ° + - +
Chione undatella . : ; ‘ : . 3 é ‘ * - -
Crepidula sp. 3 ‘ : . A ‘ : . ‘ + = =

Dentalium sp. - : 5 5 - C . ° ° + - -

218 ALLAN HANCOCK PACIFIC EXPEDITIONS

VOL. 19

Mollusca in Los Angeles-Long Beach Outer Harbor (continued)

Macoma sp.

Mangelia sp. . 5
Odostomia sp.
Periploma discus .
pholad C F .
?Poromya sp. . 5
Solamen columbianum

Solen sicarius 3

Tellina carpenteri

Vitrinella sp. ;
clam. C O 0

Macoma indentata .
Chlamys sp. . 6
CxGublayay Be. 6
Macoma nasuta 5
Mactra ?californica
Mangelia barbarensis

Modiolus sp. . 5

Ophiodermella incisa
Crepidula aculeata

Lucinoma annulata
Protothaca tenerrima

Solen rosaceus A

Tagelus californicus .

Cadulus fusiformis

Modiolus neglectus

Parvilucina tenuisculpta

Tellina buttoni

Thyasira barbarensis

Volvulella tenuissima

.

al

2

3

i

m

5

NO. 2 HARTMAN : QUANTITATIVE SURVEY 219

PALOS VERDES SHELF AND SLOPE
The Palos Verdes shelf (PV), in depths of 7 to 10 fms, supports

conspicuous Chaetopterus associations consisting chiefly of this genus with
other kinds of polychaetes. In its shallowest part, where sediments are
somewhat sandy, the parchment-worm is associated with enteropneusts
harboring pinnotherid crabs, the large swarming nereid, Neanthes
brandti, juvenile sand-dollars Dendraster excentricus, a tubicolous an-
nelid Eupolymnia crescentis with a commensal polynoid FHalosydna
brevisetosa, the proboscis worms, G/ycera americana and G. robusta, and
small spioniform worms, especially species of Polydora and Prionospio.
At about 10 fms, the parchment-worm is associated with the case-building
clam Lima dehiscens, and the worms Myxicola infundibulum and Gol-
fingia ?hespera; in bottoms where rocks and gravel occur, it is present
with gorgonian coelenterates, T'elepsavus costarum and Phyllochaetop-
terus prolifica.

In depths of 11 to 20 fms, Chaetopterus is associated with decreasing
numbers of Lima dehiscens and, where gravel or rocks occur, with the
spiny ophiuroid, Ophiothrix spiculata, the elbow crab, Heterocrypta
occidentalis, the cone-snail, Conus californicus, other mollusks, including
Nassarius, Megasurcula, Polinices, together with large nemerteans,
Cerebratulus rubra, the large cageworm, Pherusa capulata, as well as
many other kinds of polychaetes, anemones of several kinds, and sipun-
culids. Red-brown sands support Sipunculus nudus, Lytechinus anamesus,
other ophiuroids, and many kinds of shelf-polychaetes. In depths of 14
to 20 fms, a sea-star, Astropecten californicus, ophiuroids, such as 4 mphi-
oplus hexacanthus, Amphipholis squamata and Amphiodia urtica, are
prominent, together with increasing numbers of small crustaceans and
polychaetes.

In 21 to 30 fms, the sediments of the Palos Verdes shelf are black
muds with odor of hydrogen sulfide, flocculent black debris and decaying
vegetation. Animals are chiefly polychaetes, Dorvillea articulata, Ampha-
rete arctica, Lumbrineris pallida, other Lumbrineris species, O phiod-
romus pugettensis, Boccardia basilaria, Tharyx species, Diopatra ornata,
and other smaller polychaetes, associated with a clam, So/emya sp., sipun-
culids, and Amphiodia urtica. At this depth Chaetopterus and crusta-
ceans are nearly absent.

In 31 to 40 fms the animals in the sediments are mainly polychaetes,
especially cirratulids, spionids, some Pectinaria californiensis, and nemer-
teans, such as the large Cerebratulus; surface forms include Chloeia
pinnata, O phiothrix spiculata, and the large sea-whip, Sty/atula. Crusta-

220 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

cea are absent except for the barnacle, Scalpellum, attached to stiff tubes
of polychaetes.

In deeper bottoms, sediments are silt and mud; polychaetes continue
the most prominent, and occur with ophiuroids, echiuroids, sipunculids,
small solenogasters and coelenterates. The most conspicuous polychaetes
are Chloeia pinnata, Lumbrineris pallida, Spiophanes and Tharyx species,
together with onuphids and maldanids. Brissopsids are frequent, as well
as large echiuroids, Arynchite, and nemerteans, Cerebratulus. Charac-
teristic animals in 195 fms are the large mud-tube polychaetes, Pista
disjuncta and Maldane sarsi, and a tubicolous ceriantharian.

Specific diversity ranges from 25 to 129 in a sample, and numerical
count from 100 to 2448. Biomasses are moderately high, ranging to
840.0 grams for a sample measuring 0.16 cuft.

The following stations, by depth, were useful in indicating the diver-
sity and kinds of species in the Palos Verdes area; they are numbered 1
to 27 (see charts), with station (4801-57 to 4832-57) and serial number
(11 to 60) (see Map, vol. 19, p. 6).

1. Sta. 4801 (no. 28). 0.3 mi offshore from Portuguese Point, in 7
fms. OPG took 1.0 cuft of organic debris, shelly silty mud and red algae.
In addition to species shown on the chart, the sample contained a small
anemone (1), nemerteans (10), four kinds of amphipods (20 speci-
mens), a Caprella (11), a tanaid (1), and a cumacean (2).

Numbers of species and specimens were estimated at:

polychaetes 34* species, 381 specimens
echinoderm 1 84
mollusks 6 1]
crustaceans 7 34
others 2 11

Total: 50+species, 521 specimens

‘The most conspicuous was Chaetopterus, followed by Diopatra and ju-
veniles of Dendraster. The most abundant were Tharyx marioni (91),
Dendraster sp. (84), Ophiodromus pugettensis (56), with 20 or more
of each of the following: Chaetopterus variopedatus, Eupolymnia crescen-
tis and Halosydna brevisetosa.

2. Sta. 4830 (no. 28). 2.5 mi from Pt. Vicente light, in 7 fms. OPG
took 0.95 cuft of fine olive-gray sand. The screenings contained large
numbers of Dendraster sp. and other animals. In addition to those shown

NO. 2 HARTMAN : QUANTITATIVE SURVEY 224

on the chart, crustaceans were represented by many amphipods and caprel-
lids, and a crab, Heterocrypta occidentalis. Largest species were Polin-
ices and Neanthes brandti; most abundant were Polydora ?caulleryi

(312+) and Diopatra ornata (37).

3. Sta. 4806 (no. 28). 1.25 mi from Pt. Vicente light, in 7.5 fms.
OPG took 1.8 cuft of debris with many algae, large tubes of Chaetop-
terus variopedatus and Lima dehiscens in cases. In addition to those
listed on the chart, coelenterates were represented by a small anemone
(2) and a ceriantharian (1); polyclads by two species, nemerteans by
three species, oligochaetes by one, and Glottidia albida (1); bryozoans
were also present. Amphipods were estimated at 11 species and 128 speci-
mens, and decapods at two small individuals; one enteropneust was pres-
ent. Ihe largest species was Chaetopterus variopedatus and the most
abundant Dorvillea articulata (754 individuals), followed by Chaetop-
terus with about 100, Lima dehiscens with 90, Diopatra ornata and
Ophiodromus pugettensis with 54 each, Flabelligera commensalis with
46, and Lumbrineris cruzensis with 53.

Numbers of species and specimens totalled :

polychaetes 51 species, 1509 specimens
echinoderms 3 7
mollusks My, Zaid
crustaceans 14 130
others 12 94

Total: 102 species, 2017 specimens

Sub. 3. Sta. 4833 (no. 27). 0.6 mi from Pt. Vicente light, in 8.5
fms. OPG took 0.95 cuft of coarse gray sand, which contained chiefly
Dendraster excentricus and Chaetopterus variopedatus, also a few small
pelecypods.

4. Sta. 4807 (no. 28). Off Pt. Vicente light, in 9 fms. OPG took
2.4 cuft of massed Chaetopterus tubes with gravelly sand. The sample
contained nemerteans of 3 kinds and many specimens, a sipunculid, crus-
taceans with | amphipod, 10 caprellids, and a large brown enteropneust
(2), in addition to the animals listed on the chart. ‘The largest and most
conspicuous individuals were Chaetopterus variopedatus (ca. 100) and
Lima dehiscens (about 40). Other numerous species were Dorvillea artic-
ulata, Lumbrineris pallida, Ophiodromus pugettensis, Telepsavus cos-
tarum, Tharyx multifilis, and nemerteans of at least 3 kinds.

Wa? ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 22 species, 240* specimens
echinoderms Z +
mollusks 6 61
others 3 133
Total: 33 species, 318+ specimens

5. Sta. 2788 (no. 14). South of Redondo Beach, in 9.5 fms. OPG
took 1.32 cuft of coarse gray sand and shell, with dead shells of Olivella,
Pecten, Cadulus, Nassarius, Dendraster occupied by sipunculids, and
solitary tunicates. The sample contained many worm-like animals, with
the most numerous being Golfingia (sipunculid), Pisione remota (50),
cirratulids and hesionids. Crustaceans included several kinds of amphi-
pods with 34 specimens and a caprellid, and an anemone (1). Largest
individual was Marphysa mortenseni.

Sub 5. Sta. 2470 (no. 44). 1.4 mi E of Pt. Fermin light, in 10 fms.
OPG took 1.95 cuft of black sandy mud with foul odor. Most abundant
were many kinds of polychaetes, like those in adjacent stations, with un-
usual abundance of Lumbrineris spp., Streblosoma crassibranchia, and
an enteropneust, possibly Spengelia sp.

6. Sta. 4805 (no. 28). 1.8 mi from Pt. Vicente light, in 10 fms.
OPG took 1.8 cuft of Chaetopterus-Diopatra ornata association in sand
with red algae. A few small ?Harenactis (anemone), a large speckled
gray nemertean, several smaller ones, and two kinds of sipunculids, ac-
companied by large numbers of polychaetes and Lima dehiscens, com-
prised most of the sample. Crustaceans were present with a few amphi-
pods, a Caprella, and Heterocrypta occidentalis (3). he largest and
most conspicuous animals were Chaetopterus variopedatus, Lima dehis-
cens, Chone sp., Flabelligera commensalis, Spiophanes missionensis and
Tharyx multifiilis. Echinoderms were absent.

7. Sta. 4856 (near no. 28). 2.9 mi from Pt. Vicente light, in 10
fms. OPG took 0.81 cuft of fine olive-green sand and rocks with small
white attached sponges. In addition to those on the chart, the lot con-
tained a small white anemone (2), a large Cerebratulus rubra and a
small nemertean (10), Glottidia albida (1), Terebratalia sp. (5), ten
kinds of amphipods (77), two cumaceans, an isopod (15), an ostracod,
an elbow crab (2) and a spider crab, a pycnogonid (6), and a large dark
brown enteropneust (1). The largest species were Burchia redondoensis
and Pista disjuncta, the most abundant Paraonis gracilis (50+), Priono-
spio malmgreni (45), and Ophiothrix spiculata (29).

NO. 2 HARTMAN : QUANTITATIVE SURVEY 223

Numbers of species and specimens totalled :

polychaetes 54 species, 350 specimens
echinoderms - 47
mollusks 17 34
crustaceans 20 99
others 10 40

Total: 105 species, 570 specimens

8. Sta. 3050 (near no. 28). 2.1 mi west of Pt. Fermin light, in 10-11
fms, dredged, in rocky and mixed bottom. Gorgonian corals (Leptogor-
gia hebes) were most conspicuous; other species are listed on the chart.

9. Sta. 2472 (no. 28). 2.25 mi E of Pt. Vicente light, in 11 fms.
OPG took 0.63 cuft of coarse black sandy mud with broken shells and
many animals. In addition to those shown on the chart, the sample con-
tained a few small anemones, several kinds of nemerteans, Glottidia
albida (1), and few crustaceans (amphipods, caprellids, ostracods, and
Heterocrypta occidentalis). Echinoderms were absent.

10. Sta. 5102 (near no. 28). 3.35 mi from Pt. Vicente light, in 11
fms. OPG took 0.25 cuft of coarse black sand, with tubes of Chaetop-
terus. A ceriantharian (2), a polyclad (6), at least two kinds of nemer-
teans (21), a sipunculid (4), an oligochaete (30+), caprellids (100*),
an amphipod (1), and Heterocrypta occidentalis (2), were present in
addition to those listed in the chart. The largest species was Polinices ;
the most conspicuous Chaetopterus variopedatus (65); the most abun-
dant Dorvillea articulata (550*), Sphaerosyllis californiensis (100*),
Paraonis gracilis (85*), and Lima dehiscens (41). Echinoderms were
nearly absent.

Numbers of species and specimens totalled :

polychaetes 60* species, 4200+ specimens
echinoderms 1 Z
mollusks nS 79
crustaceans oy 103*
others 8 64+

Total: 87+ species, 4448+ specimens

Sub 10. Sta. 5101 (near no. 28). 1.85 mi from Abalone Cove, in 11
fms. OPG took 0.88 cuft of coarse gray-black sand with many animals.
Screenings consisted of large clumps of Chaetopterus variopedatus with
Lima dehiscens, a tubicolous ceriantharian, nemerteans of several kinds,

224 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

sipunculids, many kinds of polychaetes, resembling those in adjacent
samples, Heterocrypta occidentalis, and many other smaller kinds.

11. Sta. 4802 (near no. 27). Near Marineland, in 12 fms. OPG
took 2.2 cuft of mixed gravel and sand, tubicolous polychaetes and
massed cases of Lima. In addition to those listed on the chart, there were
two small anemones, a polyclad (3), two kinds of nemerteans, a sipun-
culid, three kinds of amphipods (7), two caprellids (8), a tanaid (1),
Heterocrypta sp. (3). Most conspicuous species were Chaetopterus vario-
pedatus and tubes of Polyodontes; most abundant were Ampharete
(100+), and Flabelligera commensalis, Lumbrineris pallida, O phiod-

romus pugettensis and Lima dehiscens, each with more than 10.

Numbers of species and specimens totalled :

polychaetes 31*species, 300*specimens
echinoderms 1 |
mollusks 14 103
crustaceans 7 19
others 4 10

Total: 57* species, 433+ specimens

12. Sta. 5030 (near no. 43). 2.7 mi from Pt. Fermin, in 12 fms.
OPG took 1.13 cuft of gray and black sand with tubes of Chaetopterus.
The sample contained a ceriantharian, small nemerteans (5), a sipun-
culid (5), an oligochaete (10*), five kinds of amphipods (20), a caprel-
lid (3), two kinds of isopods (8), Heterocrypta (1), a shrimp (4), a
pycnogonid (1), an enteropneust (1). Largest and most conspicuous
species was Chaetopterus ; most abundant were Exogone uniformis (78),
Dorvillea articulata (70), Melinna denticulata (35), and Conus cali-
fornicus (33).

Numbers of species and specimens were estimated at:

polychaetes 47* species, 468*specimens
echinoderms 3 20
mollusks 24 128
crustaceans 10 33
others 7. 24

Total: 91+ species, 673*specimens

13. Sta. 5541 (no. 28). 8.8 mi from Pt. Fermin light, in 12 fms.
OPG took 1.63 cuft of coarse black sand with odor of hydrogen sulfide.

bo
ie)
Ga

NO. 2 HARTMAN : QUANTITATIVE SURVEY

‘The wet weights of larger animals were:
ceriantharian (1) 7.0 grams
Cerebratulus (2) Shs
?Golfingia hespera (245) 106.0

polychaetes, without tubes 617.6

echinoderms 0.5
decapods 32.0
Eotal: 768.8 grams

In addition to those listed in the charts, the sample contained: a large
ceriantharian, a small white anemone, Cerebratulus (2), small nemer-
teans (10*), two kinds of sipunculids (256), five kinds of amphipods
(42), an isopod (1), a tanaid (1), Cancer sp. (3), Heterocrypta occt-
dentalis (14), Scalpellum (1), a shrimp (3). Largest species were Pher-
usa capulata, Myxicola infundibulum, Cerebratulus, ceriantharian and
Heterocrypta; most abundant were Chaetopterus (200), Phyllochaetop-
terus prolifica (100*) and Telepsavus costarum (100*).

Numbers of species and specimens totalled:

polychaetes 28 species, ca. 686 specimens
echinoderms is 3
mollusks Wee 156
crustaceans ca. 12 65
others 6 270

Total: ca. 60 species, ca. 1180 specimens

14. Sta. 5028 (near no. 43). 0.9 mi from Pt. Fermin light, in 12
fms. OPG took 0.1 cuft of coarse, gray-black sand, with many tubes of
Chaetopterus. Coelenterates were represented by a small Harenactis and
another anemone (3); two kinds of polyclads (3), a nemertean (5),
two kinds of sipunculids (83), an oligochaete (10), an amphipod (10),
a caprellid (6), and Glottidia albida (3). The largest species was Chae-
topterus, the most abundant Pherusa neopapillata (197), Dorvillea artic-
ulata (130), Golfingia (79), Ampharete labrops (50), and Sphaerosyl-
lis californiensis (66). Echinoderms, mollusks and crustaceans were

nearly absent.

226 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 54 species, 797 specimens
echinoderms l 1
mollusks 7 8
crustaceans 2 16
others 10 114

Total: 74 species, 936 specimens

15. Sta. 3051 (no. 42). 2.4 mi off Pt. Fermin light, in 11-12.5 fms.
Biol. dredge with tangles, took large quantities of Chaetopterus with asso-
ciated animals. In addition to those on the chart, crustaceans were rep-
resented by a few Heterocrypta, amphipods, isopods. ‘The most conspicu-
ous animals were Chaetopterus, Diopatra ornata and Ophiothrix spic-

ulata.

Sub 15. Sta. 2473 (no. 27). 0.75 mi SE of Pt. Vicente light, in 16
fms. OPG took 0.75 cuft of coarse sandy mud, shell fragments, and
Chaetopterus with O phiothrix (see Hartman, 1955: 53).

Sub 15. Sta. 2962-54 (near no. 42). 1.5 mi W of Pt. Fermin light,
in 20 fms. Biological dredge took black mud with Chaetopterus, large
Stylatula, Listriolobus pelodes, sipunculids, Modiolus and a few other
pelecypods, also many polychaetes. Echinoderms and crustaceans were
nearly absent.

16. Sta. 3049 (no. 42). 2.1 mi from Pt. Fermin light, in 23 fms.
OPG took a small volume of black mud with dead shells of Lima, Modi-
olus, Solen, Polinices, Olivella, Pecten, scaphopod and caecid shells. The
sample contained Stylatula (4, very long), a ceriantharian (2), poly-
clads (10), few amphipods and cumaceans, and others listed on the chart.
The largest individuals were Stylatula and ceriantharian. Most abun-
dant were Capitella capitata subsp. (985), Dorvillea articulata (700*),
and A mpharete arctica (200+).

17. Sta. 4803 (no. 27). 1.6 mi from Pt. Vicente light, in 24 fms.
OPG took 2.2 cuft of black detrital flocculent debris with foul odor. The
sample contained large Sty/atula (6), measuring to 50 cm long, a large
Cerebratulus rubra, and smaller nemerteans, a caprellid (3), and other
species shown on the chart. Most abundant species were Dorvillea articu-
lata (100+), Lumbrineris pallida (20*), Phyllochaetopterus limicolus,
and Tharyx spp. (70+).

18. Sta. 5096 (near no. 28). 2.5 mi from Pt. Fermin light, in 25
fms. OPG took 1.57 cuft black silt with many small animals. A sea

NO. 2 HARTMAN : QUANTITATIVE SURVEY 227.

whip (1), a polyclad (3), two kinds of nemerteans (15), a sipunculid
(1), were present in addition to those listed on the chart. The largest
individual was the sea whip, the most abundant species Dorvillea articu-
lata (437), Capitella capitata subsp. (190*), and Telepsavus costarum
(64). The rare cirratulid, Raricirrus maculata, was well represented

(eae

19. Sta. 2417 (no. 42). Off Pt. Fermin light, in 34 fms. OPG took
2.14 cuft of black mud. A slender sea whip (3), a polyclad (1), and
nemerteans (2+), accompanied the species listed on the chart. The largest
animal was Molpadia intermedia, the most abundant species T’haryx
spp. (1620*) and Spiophanes (100*). Crustaceans and echinoderms were
nearly absent.

20. Sta. 5027 (near no. 42). 1.9 mi from Pt. Fermin light, in 38
fms. OPG took 1.82 cuft of green-black silty sand. A large Stylatula
(1), ?Harenactis (1), Cerebratulus rubra (1), and two other smaller
nemerteans (30), a phoronid (4), a hirudinean (1), an amphipod (2),
a caprellid (1), Scalpellum (12), were taken in addition to those listed
on the chart. Largest individuals were Cerebratulus rubra and Stylatula
spp. Most abundant species were Tharyx multifilis (2634) and Spio-
phanes spp. (214).

Numbers of species and specimens totalled :

polychaetes 63 species, 3357 specimens
mollusks 12 30
crustaceans 5 34
others 8 43

otal: 88 species, 3464 specimens

21. Sta. 4855 (no. 12). 3.8 mi from Pt. Vicente light, in 39 fms.
OPG took 1.26 cuft of fine gray sand with detritus. Coelenterates were
represented by Monobrachium parasitum (many, on a bivalve), ?Haren-
actis (1); a polyclad (2), two kinds of nemerteans (13), a sipunculid
(4), nine kinds of amphipods (52), three kinds of cumaceans (13), three
kinds of isopods (45), tanaids (7), two kinds of ostracods (12), a
shrimp (1), were also present. The largest animals were 4 phrodita
refulgida and Amphiura arcystata. Most conspicuous and abundant spe-
cies were Chloeia pinnata (126) and Spiophanes missionensis (56).

228 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 76 species, 675 specimens
echinoderms ii 24
mollusks Z| 55)
crustaceans 20 131
others 5 20

otal: 129 species, 905 specimens

22. Sta. 5502 (no. 42). 2 mi from Pt. Fermin light, in 49 fms. OPG
took 2.39 cuft of green silty sand. Nemerteans included a large Cerebra-
tulus (1), a small, red-banded (4), and a longitudinally striped one.
A sipunculid (1), and Branchiostoma (1) were also present, in addition
to those listed on the chart. Largest animal was Pherusa capulata, and
most conspicuous was Chloeia pinnata (320) ; Tharyx tesselata was the
most abundant (1700+). Echinoderms and crustaceans were sparse or
absent.

23. Sta. 5029 (no. 41). 2.95 mi from Pt. Fermin light, in 50 fms.
OPG took 1.76 cuft of fine greenish black silty sand with flocculent
debris. In addition to those listed on the chart, the sample contained Cere-
bratulus (2 large, weighing 11 grams), and another small kind (many),
also two kinds of sipunculids (2). The largest individual was Cerebra-
tulus, the most abundant species Capitella capitata subsp. (300), Dor-
villea articulata (110), and Spiophanes missionensis (75+). Echinoderms
were absent, and crustaceans were represented only by large Scalpellum
(5). Mollusks were few and small. Total numbers were estimated at 38
species and 803 specimens.

24. Sta. 2430 (no. 60). 2.3 mi off Pt. Fermin, in 80 fms. OPG took
1.44 cuft of hard-packed mud with many animals. In addition to those
listed on the chart, there was a nemertean (2 small), four kinds of
amphipods (25), an anthurid isopod (1), a tanaid (1), and an octracod
(1). There was no conspicuously large individual, and none were out-
standingly abundant. Total numbers were estimated at 41* species, and
185* specimens.

25. Sta. 4854 (near no. 11). 5.25 mi from Pt. Vicente, in 98 fms.
OPG took 1.57 cuft of dark gray silty sand, gravel and rocks. The grab
was lowered twice. Nine colonies of Alonobrachium were recovered on
Pseudopythina sp.; others present were a nemertean (2), an echiuroid,
Arynchite (2, measuring 62 mm long), three kinds of amphipods (4),
an isopod (1), a cumacean (5), and ostracods (4). The largest indi-

NO. 2 HARTMAN : QUANTITATIVE SURVEY 229
viduals were brissopsids, Arynchite and Travisia pupa. Vhe most con-
spicuous species was Amphiodia digitata and the most abundant were
Tharyx tesselata (100*), and onuphids (65*). ‘Total numbers were esti-
mated at 76 species and more than 600 specimens.

26. Sta. 4831 (near no. 41). 3.6 mi from Pt. Vicente light, in 195
fms. OPG took 2.83 cuft of olive-green silt. A ceriantharian (1), Cere-
bratulus sp. (+), and a small nemertean (2*) were taken, in addition to
the species listed below. Largest individual was Eunice americana, and
most abundant species was Pectinaria californiensis (25).

Numbers of species and specimens totalled :

polychaetes 19 species, 85 specimens
echinoderms 2 s
mollusks 1 4
others 5) 8
‘Total: 25 species, 100 specimens

27. Sta. 4832 (near no. 19). 2.2 mi from Pt. Vicente light, in 195
fms. OPG took 1.32 cuft olive-green sandy and sticky silt. No large ani-
mal was present. In addition to those listed below, there were a cerian-
tharian (1), a white polyclad (1), and an echiuroid (1). Most abundant
were Chioeia pinnata (96), and Pista disjuncta (5+).

Numbers of species and specimens totalled :

polychaetes 15 species, 254 specimens
echinoderms 6 48
mollusks 10 67
others 5} 3)

Total: 34 species, 372 specimens

VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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NO. 2 HARTMAN : QUANTITATIVE SURVEY pay |

SAN PEDRO SHELF

The San Pedro (SP) shelf, in depths to 50 fms, supports a diversi-
fied and complex fauna, changing from one location to the next, accord-
ing to kinds of sediments, location, and other physical factors. Some shal-
lowest depths, off Seal Beach, characterized as reddish brown beach-sand,
have a unique association of animals; similar deeper sands, in 17 fms,
have different kinds and numbers of animals. Vertical zonation is simi-
larly demonstrable in gray to black silts, in hard-packed sands, and in
rocky or mixed sediments—all of which are represented in the San Pedro
area. Changes of specific units with depths, kinds of sediments and other
factors, are indicated on the charts of species; they show not only eco-
logical groups of species, but vertical zonation, and relative abundance
of the more numerous kinds. Proportionately few species occur in all
depths, from shallowest to greatest; most are limited to one or few sta-
tions; some other species show replication by association with other bio-
logical units.

The charts demonstrate further the repeated occurrence of species
within genera; in polychaete genera it is shown for species of Lumbri-
neris, Pherusa, Prionospio, Glycera, Spiophanes, and others. ‘hese oc-
currences may be more than incidental, for it can be assumed that the
“rain of larvae” is present over all; their settlement and growth may be
restricted by factors concerned with peculiarities of larval or planktonic
adaptability.

Another process of some interest is the frequent recurrence of non-
reproductive individuals, especially in outer, slope areas ; here shelf-species
are present but do not attain maturity; they show the same patterns of
distribution and growth as do those below sill depths of offshore basins.

The changes of species with depth, kinds of sediments, faunal com-
position, and ecological associations are shown for the polychaetes, echi-
noderms and mollusks, in the charts below. he amphipods, which were
equally numerous in some samples, and other crustaceans are omitted from
the tabulations because they have been incompletely named, although
more than 200 species have been named or identified in reports by Dr. J.
L. Barnard (1959-1963) and many identified collections are consultable
in the collection of the Allan Hancock Foundation.

A comparison of numbers of species and specimens, in depths of 6.5
to +48 fms, shows no decline or rise with depth of bottom, except in rocky
areas, where numbers appear lower; this is due, in part, to the ineffective
sampling methods, for when samples are hand-picked, as by diving, the
numbers rise remarkably. The values of specific diversity, on the San
Pedro shelf, by increasing depth, are illustrated :

252 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19
Size of
Depth sample No. of No. of

Sta. no. in fms. in cuft species specimens
4885 6.5 Isreis 85 800
5740 8 0.06 66 529
2504 9 0.13 41 LS
3047 10 — 63 170
5753 10 0.13 64 320
4886 10.5 1.44 94 1097
6102 125 0.63 o2 574
2107 13 2.0 68 316
4719 13 0.44 76 266
6100 11 Se) 0.78 80 433
5/52 14.5 0.06 Ty 494
6104 14.5 On 1S) 945
5741 15 0.31 101 899
5844 15 0.06 48 239
5754 16 0.88 72 716
5751 16.5 0.31 65 666
5742 17 0.44 63 358
5743 19 0.44 127, 1412
5820 24.5 0.18 102 734
5750 Se) 0.13 67 376
5087 27 0.56 98 522
5003 30 0.18 78 By
5086 38 0.37 79 1402
5748 38 0.25 108 1347
2355 4] 0.10 32% 240
5749 42 0.13 87 924
2630 45 0.86 42 689
5745 46 0.44 82+ 1401
2126 48 — 64+ So}

The following samples (1 to 35) from the San Pedro shelf, come from
depths of 5 to 48 fms. Polychaetes, echinoderms and mollusks are listed
on the chart (immediately following) ; others are summarized in the

NO. 2 HARTMAN : QUANTITATIVE SURVEY 255

Analyses. Some of the crustaceans are more completely named in the

State report (California, 1965, App. pp. 104-301).

1. Sta. 2741 (no. 50). Off Seal Beach, in 5 fms. Campbell grab
took 2.15 cuft of red-brown beach sand with broken shells. The fauna
was a limited one, comprising chiefly three species of polychaetes (see
chart), two echinoderms, and two mollusks, each in considerable

numbers.

2. Sta. 4885 (no. 49). 2 mi from east end of Long Beach breakwater,
in 6.5 fms. OPG took 1.89 cuft of red sand over grayish black silt. In
addition to the species listed on the chart, there were a small anemone,
a polyclad (3), nemerteans (21), a phoronid (1), Glottidia albida
(1), 11 kinds of amphipods (26), two cumaceans (5), three ostracods
(6), a pinnixid crab (3), a shrimp (1), a pycnogonid (7), Branchio-
stoma sp. (22), and an enteropneust (3). Largest species was Solen
sicarius, and most conspicuous Branchiostoma. Most numerous species
were Mediomastus californiensis (125), Tharyx tesselata (113), Prio-
nospio pinnata (61), and Amphipholis squamata (52).

Numbers of species and specimens totalled :

polychaetes 45 species, 619 specimens
echinoderms + 64
mollusks v 9
crustaceans 20 44
others 9 64

Dota: 85 species, 800 specimens

3. Sta. 5740 (no. 68). 9.75 mi from Los Angeles harbor light, in 8
fms. OPG took 0.06 cuft of fine gray sand. In addition to those named in
the chart, the sparse sample yielded a small coelenterate, two nemerteans
(9), bryozoans, eight amphipods (33), two cumaceans (9), a Callia-
nassa sp. (4), and a crab (2). Largest species were Terebra pedroana
and O phiodermella incisa; most abundant were Chaetozone corona (65),
Tharyx tesselata (51), Prionospio malmgreni (46) and P. pinnata
(42), and Olivella baetica (31).

Numbers of species and specimens totalled :

polychaetes 37 species, 382 specimens
echinoderms 2 8

mollusks 13 67
crustaceans 13 50

others 3 Dili

‘Total: 68 species, 529* specimens

254 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

4. Sta. 2504 (no. 67). Off Los Angeles light, in 9 fms. OPG took
0.13 cuft of black sandy mud and gray sand and broken shells. In addi-
tion to those named on the chart, there were nemerteans (2), amphipods
(15), a tanaid (1), cumaceans (3) and ostracods (6). All were small,
none was abundant, with Tellina buttoni (17) the best represented.

Numbers of species and specimens totalled :

polychaetes 28 species, 61 specimens
echinoderms 1 2
mollusks 5 Fl
crustaceans 4+ 25
others 3 6

Total: 41+species, 115 specimens

5. Sta. 3047 (near no. 45). 1.1 mi east of Los Angeles light, in 10
fms. OPG took a small, mud sample with many animals. Small anemones
and a large enteropneust accompanied the specimens listed on the chart.
None was abundant.

Numbers of species and specimens totalled :

polychaetes ca. 50 species, 100*specimens

echinoderms 2 5
mollusks 3 5
Total: ca. 55 species, 110* specimens

6. Sta. 5753 (near no. 45). 2.45 mi from Los Angeles harbor light,
in 10 fms. OPG took 0.13 cuft of fine gray sand. In addition to those
listed on the chart, there were two nemerteans (14), five amphipods
(23), five cumaceans (14), an isopod (1) and a tanaid (3). The largest
species was T'erebra pedroana, the most abundant Prionospio malmgreni
(36), Mediomastus californiensis (35) and Tellina carpenteri (18).

Echinoderms were absent.

Numbers of species and specimens totalled :

polychaetes 34 species, 217 specimens
mollusks 2 44
crustaceans 14 4]
others + 18

Total® 64 species, 320 specimens

7. Sta. 3048 (near no. 45). 1.25 mi east of Los Angeles light, in
10.5 fms. OPG took a small mud sample with animals. Coelenterates,
with a ceriantharian, a broad-pannicled sea-pen and branching hydroids,
several nemerteans, a phoronid, a Glottidia albida, and a few amphipods,

No. 2 HARTMAN : QUANTITATIVE SURVEY 253

caprellids, an ostracod, a spider crab and an enteropneust, ?Schizocar-
dium (2) were present, in addition to those listed on the chart. The
largest individuals were the sea-pen and the enteropneust. Most abun-
dant species were Nothria iridescens and Phyllochaetopterus prolifica.
Numbers of species is estimated at 58*, and specimens at more than 200.

8. Sta. 4886 (near no. +8). 0.85 mi east of Long Beach, near break-
water, in 10.5 fms. OPG took 1.44 cuft of gray to black, medium to
coarse sand. Coelenterates were represented by ?Harenactis (23) and a
sea-whip; one nemertean (32), 11 amphipods (35), two isopods (2),
three cumaceans (26), three ostracods (4), Callianassa sp. (1), two pin-
notherid crabs (19), two pycnogonids (8), and an enteropneust (7).
The largest animal was 4 mphiodia occidentalis, and the most conspicu-
ous were T’haryx tesselata (300*), Scalibregma inflatum (61) ,, Medio-
mastus californiensis (45), Amphipholis squamata (41), and Brada
pilosa (34).

Numbers of species and specimens totalled:

polychaetes 49 species, 836 specimens
echinoderms 3 69
mollusks 9 16
crustaceans 23 92
others 11 84+
Totals: 95 species, 1097+ specimens

9. Sta. 3053 (near no. 47). 3.5 mi off Los Angeles light, in 11-12
fms. Biological dredge, in rocky and rubbly bottom, took a diversified
fauna, with a large pagurid, echinoderms, fishes, and many other ani-
mals. Coelenterates were present with gorgonian corals, Euplexaura
marki, coarse brown Eunicea, Ptilosarcus quadrangularis (broad-pan-
nicled sea-pen) and Corynactis californica. Bryozoans were encrusted on
rocks, and crustaceans included many amphipods, caprellids, and barnacle
clumps. The largest individuals were Holopagurus pilosus, and Astropec-
ten sp. Number of species was estimated at more than 57.

10. Sta. 2311 (no. 66). Off Long Beach breakwater light, in 12 fms.
OPG took 0.5 cuft, in 2 drops. In addition to the species named (Hart-
man, 1955, p. 76) additional species are indicated in the chart. Total
number of species is well over 100.

11. Sta. 6102 (near no. 47). Off Long Beach breakwater, in 12.5
fms. OPG took 0.63 cuft of dark gray fine sand. A polyclad (1), a
small nemertean (28), six amphipods (18), four cumaceans (8), two
ostracods (10), a mysid (1), and a pinnixid (1), accompanied the spe-

256 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

cies named on the chart. All species were moderately small; most abun-
dant were Prionospio pinnata (47), Chaetozone corona (33), Tellina
carpenteri (32).

Numbers of species and specimens totalled :

polychaetes 50 species, 373 specimens
echinoderms 3 15)
mollusks 17 4]
others p) 4]
Total: 75 species, +70 specimens

12. Sta. 2107 (no. 46). 2.2 mi off Los Angeles light, in 13 fms.
OPG took 2.0 cuft of sand and mud with many animals. Present were
a sea-whip (2), a small anemone, a sipunculid (8), an echiuroid, Listrio-
lobus pelodes (2), a phoronid (3), Glottidia albida (2), an amphipod
(1), a caprellid (3), a cumacean (1), an ostracod (1), a brachyuran
crab (1), and an enteropneust (1). Largest animal was Tagelus califor-
nicus (6) measuring to 80 mm long, and most abundant species were
Nereis procera and Sthenelanella uniformis, each with 31, and Cossura
candida with 20.

Numbers of species and specimens totalled :

polychaetes 38 species, 256* specimens
echinoderms 2 15
mollusks 5 1
crustaceans 5 ii
others § 21

Total: 58 species, 316* specimens

13. Sta. +719 (near no. 48). East end of Long Beach breakwater,
in 13 fms. OPG took 0.44 cuft of gray and green sand, with many ani-
mals. ?/larenactis sp. (4) and a nemertean (3), Glottidia albida (2),
seven amphipods (19), two isopods (2), a tanaid, two ostracods (8),
and a cumacean (1) accompanied the species named on the chart. Largest
species were Nephtys caecoides and Nothria elegans, and most abundant

was Scalibregma inflatum (26).

Numbers of species and specimens totalled:

polychaetes 37 species, 175 specimens
echinoderms 5 9
mollusks 14 38
others 3) 9

Total: 59 species, 231 specimens

NO. 2 HARTMAN : QUANTITATIVE SURVEY 25

14. Sta. 6100 (near no. 46). 5.65 mi from Pt. Fermin light, in 13.5
fms. OPG took 0.78 cuft of dark gray medium to fine sand. Small nemer-
teans (14), 11 amphipods (85), a caprellid (1), four isopods (6), a
tanaid (2), three cumaceans (26), five ostracods (56), Heterocrypta
occidentalis (2), a pinnixid (1), and an enteropneust (1), accompanied
those named on the chart. The largest individual was Lovenia cordi-
formis. No species was outstandingly abundant.

Numbers of species and specimens totalled:

polychaetes 37 species, 180 specimens
echinoderms + 18)
mollusks 7 35
crustaceans 29 180
others 3 15

Total: 80 species, 433 specimens

15. Sta. 2006 (near no. 47). 2.5 mi south of Long Beach, in 14
fms. The trawl took ascidian clumps, chiefly Stye/a and Amaroecium
colonies. A polyclad and a pycnogonid accompanied the polychaetes
named on the chart.

16. Sta. 5752 (near no. 64). 5 mi from Los Angeles light, in 14.5
fms. OPG took 0.06 cuft of fine gray sand with many small animals,
chiefly polychaetes, together with a ceriantharian, a small nemertean
(7), Glottidia albida (5), eight amphipods (21), two isopods (5), tana-
ids (5), three cumaceans (5), two crabs (3). Largest species were Glot-
tidia albida, a ceriantharian and Amphipholis squamata; most numerous
were Prionospio pinnata (64), Mediomastus californiensis (56), Prio-
nospio malmgreni (39), and Tellina buttoni (31).

Numbers of species and specimens totalled :

polychaetes 37 species, 322 specimens
echinoderms 3 8
mollusks 16 100
crustaceans 16 50
others i 14

Total: 76 species, +94 specimens

17. Sta. 6104 (near no. 65). Nearly 10 mi from Pt. Fermin light,
in 14.5 fms. OPG took 0.37 cuft of dark gray silty fine sand. A sea-whip,

258 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

a polyclad (1), a nemertean (4), a sipunculid (2), 11 amphipods (35),
a caprellid (1), an isopod (9), five cumaceans (6), an epinebalian (3),
and a kelp crab (2), accompanied the species named on the chart. Larg-
est species were Astropecten californicus and Glycera robusta; most
abundant were 4 mphiodia urtica (56), Aricidea lopezi (56), and Chone
sp. (48).

Numbers of species and specimens totalled:

polychaetes 59 species, 567 specimens
echinoderms 6 121
mollusks 24 15
crustaceans 20 56
others oF 66

Total: 115*species, 945 specimens

18. Sta. 5741 (no. 67). 9.25 mi from Los Angeles light, in 15 fms.
OPG took 0.31 cuft of fine gray sand with many small animals. Weight
of animals was estimated at 19.2 grams, of which polychaetes comprised
9.3, ophiuroids 7.1, and mollusks 2.5 grams. In addition to those listed
on the chart, there were present a small nemertean (9), a phoronid (1),
Glottidia albida (131 juveniles), nine amphipods (43), a tanaid (7),
two ostracods (29), three cumaceans (5), Callianassa sp. (1), and a pin-
nixid (5). The largest individual was Astropecten californicus, the most
conspicuous species Amphioplus hexacanthus (88), and the most abun-
dant Glottidia albida (131) and Chaetozone corona (88*).

Numbers of species and specimens totalled :

polychaetes 49 species, 464 specimens
echinoderms + 99
mollusks 24 102
crustaceans 21 93
others 3 141

Total: 101 species, 899 specimens

19. Sta. 5844 (no. 64). 2.6 mi from Los Angeles harbor light, in 15
fms. OPG took 0.06 cuft of coarse red sand and a few rocks. In addi-
tion to those shown on the chart, this very small sample took two anemo-
nes (4), a nemertean (4), a sipunculid (2), an oligochaete (12), five
amphipods (29), an isopod (3), and a cumacean (1).

NO. 2 HARTMAN : QUANTITATIVE SURVEY 259

Numbers of species and specimens totalled :

polychaetes 28 species, 159 specimens
echinoderms L i
mollusks - i
crustaceans 8 36
others ) 22

Total: 47 species, 235 specimens

20. Sta. 5754 (near no. 62). 1.65 mi from Los Angeles light, in 16
fms. OPG took 0.88 cuft of coarse dark gray to black sand, with woody
debris and many animals. Wet weights totalled 43.4 grams, of which
polychaetes were 13, echinoids 10.4, mollusks 9.2, holothuroids 4.6, anem-
ones 3.0, nemerteans 2.9 grams, and others less than 1.0 gram. In addi-
tion to those listed on the chart, the lot contained a ceriantharian (1),
a sand-covered anemone (12), ?//arenactis (1), three nemerteans (35),
Glottidia albida (4), seven amphipods (12), two isopods (4), two ostra-
cods (8), three cumaceans (23), Callianassa (1) and Heterocrypta (1).
Largest individuals were Lytechinus anamesus (12), and Pentamera
pseudopopulifera (2) ; most conspicuous or numerous species were Pher-
usa capulata (10), Tharyx tesselata (275), Lysippe annectens (15) and
Prionospio pinnata (20).

Numbers of species and specimens totalled :

polychaetes 42 species, 585 specimens
echinoderms 2 14
mollusks 5 14
crustaceans 16 49
others | 54

‘otal: 72 species, 716 specimens

21. Sta. 5751 (near no. 63). 4.8 mi from Los Angeles light, in 16.5
fms. OPG took 0.31 cuft of gray sand with animals. In addition to those
listed on the chart, there were a ceriantharian (1), a polyclad (1), two
nemerteans (4), Glottidia albida (12), four amphipods (26), a tanaid
(4), cumaceans (4), ostracods (21), and Heterocrypta occidentalis (1).
Largest individual was Astropecten californicus (2), and most conspicu-
ous Ophiothrix spiculata (144) ; high counts occurred also in Aricidea
lopezi (96), Prionospio malmgreni (62), Onuphis nebulosa (52) and
Melinna denticulata (45).

260 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 42 species, +16 specimens
echinoderms 5) 166
mollusks + 9
crustaceans 8 56
others 6 19

Total: 65 species, 666 specimens

22. Sta. 3052 (near no. 63). 4.4 mi off Los Angeles light, in 16.5 to
17 fms. The biological dredge took rocks overgrown with organisms, es-
pecially sponges, Corynactis (in crevices on rocks), branching hydroids,
encrusting and branching bryozoans, and tubicolous amphipods. Echi-
noderms and mollusks were most conspicuous; smaller animals had per-
haps been lost in recovery of the sample.

23. Sta. 5742 (near no. 86). 11.7 mi from Los Angeles light, in 17
fms. OPG took 0.44 cuft of reddish brown sand with many animals. In
addition to those listed on the chart, the sample contained a rhabdocoele
(2), a sipunculid (1), Glottidia albida (5), a phoronid (6), seven am-
phipods (46), an isopod (5), an ostracod (5), and two cumaceans (5).
Largest individuals were Lovenia cordiformis, Nephtys californiensis
and Onuphis eremita. Most abundant were Lumbrineris cruzensis (33),
Spiophanes bombyx (27), and Dendraster excentricus and Lumbrineris
cf. acuta, each with 19 individuals.

Numbers of species and specimens totalled :

polychaetes 32 species, 216 specimens
echinoderms fi 28
mollusks 8 32
crustaceans 12 68
others + 14

otal: 63 species, 358 specimens

24. Sta. 5743 (near no. 83). 7.2 mi from Los Angeles light, in 19
fms. OPG took 0.44 cuft of fine green sand with many small animals.
Gross weight was only 17.4 grams, of which polychaetes were 10.1 and
ophiuroids 5.5 grams. A polyclad (1), a nemertean (25), Glottidia
albida (151), 13 kinds of amphipods (172), three isopods (12), two
tanaids (21), five ostracods (151), five cumaceans (23), Heterocrypta
(1), another crab (1), and Scalpellum (6) accompanied the species
named on the chart. Largest animals were Polinices lewis, Lumbrineris
californiensis and Nephtys californiensis. Most abundant or conspicuous
were Amphioplus hexacanthus (39), Onuphis nebulosa (15), Prionospio

NO. 2 HARTMAN : QUANTITATIVE SURVEY 261

pinnata (168), and Glottidia albida (151). Others attaining peak num-
bers were A xiothella rubrocincta (32), Aricidea lopezi (52), Mediomas-
tus californiensis (47), and Sthenelanella uniformis (38).

Numbers of species and specimens totalled :

polychaetes 60 species, 658 specimens
echinoderms if [PP
mollusks 25 110
crustaceans 32 395
others 3 er)

Total: 127 species, 1412 specimens

25. Sta. 5820 (near no. 101). 7.2 mi from Los Angeles light, in 24.5
fms. OPG took 0.18 cuft of fine olive-green sand with many animals.
A ceriantharian (29), ?Harenactis (2), a nemertean (4), Glottidia
albida (42), a phoronid (6), 13 amphipods (86), a caprellid (1), two
isopods (2), tanaids (22), cumaceans (54) and Heterocrypta (1) ac-
companied the species named on the chart. Largest animal was O phiura
lutkeni; most abundant species were Spiophanes missionensis (47), Arici-
dea lopezi (42), and Prionospio pinnata (28).

Numbers of species and specimens totalled :

polychaetes 44 species, 360 specimens
echinoderms 5 Se)
mollusks 21 56
crustaceans 26 194
others 6 89

Total: 102 species, 734 specimens

26. Sta. 5750 (near no. 100). 5.5 mi from Los Angeles light, in 25.5
fms. OPG took 0.13 cuft of fine green sand, with many animals. A ceri-
antharian (3), Glottidia albida (12), six amphipods (42), caprellid
(1), two isopods (4), two tanaids (39), and four cumaceans (11) ac-
companied the species named in the chart. Largest individual was Nephtys
caecoides ; most conspicuous were Amphiodia urtica (17), Amphioplus
hexacanthus (14), and Chloeia pinnata (3).

Numbers of species and specimens totalled :

polychaetes 30 species, 192 specimens
echinoderms + 35
mollusks 15 40
crustaceans 15 95
others 3 16

Total: 67 species, 376 specimens

262 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

27. Sta. 5087 (near no. 108). 3.6 mi from Newport pier, in 27 fms.
OPG took 0.56 cuft of dark green sandy silt with many animals. Wet
weight totalled 65.3 grams, with ophiuroids 40, polychaetes 10, crusta-
ceans 7, gorgonian 4.6, and others less than 1.0 gram. A gorgonian, a
ceriantharian, a nemertean (5), a sipunculid, small Listriolobus pelodes
(3), a papillated leech (1), Glottidia albida (1), 16 amphipods (69),
a caprellid (4), a tanaid (2), a cumacean (2), an ostracod (2), an
epinebalian (4), Scalpellum (14), Heterocrypta occidentalis (1), a
shrimp (2), a pycnogonid (1), an enteropneust (1) and Pyrosoma (1)
were present, in addition to those listed on the chart. Largest species was
O phiura lutkenit (37), and most abundant Amphiodia urtica (119).

Numbers of species and specimens totalled :

polychaetes 35 species, 162 specimens
echinoderms ih 194
mollusks ZA 5)5)
crustaceans 25 97
others 10 16

Total: 98 species, 522 specimens

28. Sta. 5003 (near no. 108). Off Newport Bay, in 30 fms. OPG
took 0.18 cuft of fine grayish green sand with many animals. Present
were a loose-pannicled sea-whip (3), ?Harenactis (1), a nemertean (8),
12 amphipods (42), a caprellid (2), an isopod (5), a tanaid (8), three
cumaceans (5), two ostracods (176). Largest individuals were a sea-
whip and Nephtys californiensis. Most numerous animals were ostracods

(176) and Prionospio malmgreni (36).

Numbers of species and specimens totalled :

polychaetes 33 species, 206 specimens
echinoderms 3 35
mollusks 18 35
crustaceans 21 239
others 3 12

Total: 78 species, 527 specimens

29. Sta. 5086 (no. 129). 4.7 mi from Newport pier, in 38 fms.
OPG took 0.37 cuft of dark green silt and sand, with many animals.
Wet weight was 12.8 grams, of which polychaetes were 5, sea-whips 4,
ophiuroids 2.8, and others less than 1 gram. A sea-whip (5), a small
anemone, a nemertean (2), 16 amphipods (64), an isopod (6), two
tanaids (16), five cumaceans (23), five ostracods (880), and an epine-
balian (2) were present, in addition to those named on the chart. The
largest individuals were a sea-whip and Nephtys caecoides; most con-

NO. 2 HARTMAN : QUANTITATIVE SURVEY 263

spicuous species were 4 mphiodia urtica (56), Chloeia pinnata (103), fol-
lowed by Pectinaria californiensis (19) and Prionospio malmgreni (43).

Numbers of species and specimens totalled :

polychaetes 26 species, 283 specimens
echinoderms 2 60
mollusks 17 5/
crustaceans 30 993
others 7 9

‘Total: 79 species, 1402 specimens

30. Sta. 5748 (near no. 101). 8.2 mi from Los Angeles light, in 38
fms. OPG took 0.25 cuft green sand and shell, with many animals. They
included a sea-whip (2), a polyclad (2), Glottidia albida (2), 22 am-
phipods (123), three isopods (18), tanaids (48), four cumaceans (37),
four ostracods (425), Scalpellum (2), a shrimp, a pycnogonid and a
colonial ascidian. Largest species were Polinices lewisti and Scalpellum,
and most abundant were Prionospio malmgreni (106), Amygdalum pal-
lidulum (90), Chloeta pinnata (47), Axinopsida serricata (40), Chaeto-
zone setosa (35), Aricidea lopezi (32), Lumbrineris cruzensis (31), and
Spiophanes missionensis (30).

Numbers of species and specimens totalled :

polychaetes 44 species, 486 specimens
echinoderms = 41
mollusks 17 157
crustaceans 38 655
others 5 8*

Total: 108 species, 1347+ specimens

31. Sta. 2355 (no. 99). 6.3 mi off Los Angeles light, in 41 fms.
OPG took 0.10 cuft of green sand with many animals. In addition to
those listed on the chart, Monobrachium (a colony), amphipods (19),
caprellids (2+), an isopod (1), a tanaid (6), and two ostracods (55)
were present. Largest individual was a maldanid, and most abundant was
A mphiodia urtica (52).

Numbers of species and specimens totalled :

polychaetes 23 species, 64 specimens
echinoderms 2 60
mollusks 5 8
crustaceans Ge 107
other | 1

Total: 37* species, 240 specimens

264 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

32. Sta. 5749 (near no. 99). 6.4 mi from Los Angeles light, in 42
fms. OPG took 0.13 cuft of fine green sand with many animals. Wet
weight of animals was estimated at 9.5 grams, with ophiuroids 5.3, poly-
chaetes 3.4 grams, and all others 0.8 grams. In addition to those listed
on the chart, 13 amphipods (47), two isopods (6), tanaids (16), two
ostracods (23), a mysid (1) and Monobrachium colonies (61) on Axi-
nopsida and Erycina, were present. The largest species were Nephtys
californiensis and Pentamera pseudopopulifera; most abundant were
Prionospio malmgreni (120), Axinopsida serricata (66), Chloeia pin-
nata (60), Amphiodia urtica (57), Aricidea lopezi (51) and Chaetozone
setosa (42).

Numbers of species and specimens totalled :

polychaetes 40 species, 536 specimens
echinoderms . 67
mollusks 19 117
crustaceans 23 136
others 1 68

Total: 87 species, 924 specimens

33. Sta. 2630 (no. 105). 4.4 mi SW of Huntington Beach pier, in
45 fms. Campbell grab took 0.86 cuft of sandy grayish green mud with
many small animals. The following were present in addition to those
named on the chart: Monobrachium parasitum (1), a polyclad (2), a
nemertean (2), amphipods of several kinds (36), two cumaceans (3),
three ostracods (9), a pinnotherid crab and a sand crab. Largest indi-
vidual was T’ravisia ?pupa and most abundant A mphiodia urtica (390).

Numbers of species and specimens totalled :

polychaetes 26 species, 140 specimens
echinoderms 4 458
mollusks 9 27
crustaceans a 50
others 3 14

Total: 51* species, 689 specimens

34. Sta. 5745 (near no. 103). 9.5 mi from Los Angeles light, in +6
fms. OPG took 0.44 cuft of fine green sand. Wet weights were esti-
mated at 15.3 grams, of which ophiuroids comprised 12.3 and polychaetes
2.2 grams, with all others 0.8 grams. The following species were present:
?Harenactis (1), a polyclad (1), anemertean (12), 11 amphipods (72),
three isopods (11), several tanaids (46), three cumaceans (48), and five
ostracods (571), in addition to those named in the chart. Largest indi-

No. 2 HARTMAN : QUANTITATIVE SURVEY 265

viduals were Panthalis pacifica and Megalomma splendida. Most con-
spicuous and abundant species was 4 mphiodia urtica (98), followed by
Lumbrineris cruzensis (98), Prionospio malmgreni (70), Chaetozone

setosa (65), Aricidea lopezi (37), and Scoloplos armiger (34).

Numbers of species and specimens totalled :

polychaetes 36 species, +446 specimens
echinoderms 6 127
mollusks 14 60
crustaceans 23; 754
others 3 14

Total: 82+ species, 1401 specimens

35. Sta. 2126 (no. 123). 9.2 mi from Los Angeles light, in 48 fms.
OPG took a moderately large sample of coralline clumps, and many
kinds of animals. In addition to those named in the chart, there were
present a stalked hydroid, a nemertean, ?4rynchite sp. (1), a leech,
branching bryozoans, amphipods (12), two isopods (2), tanaids (9), a
cumacean, three ostracods (57) and Scalpellum (2+). The largest species
was ?Arynchite sp., the most abundant Chloeia pinnata (33). Total
numbers were estimated at more than 6+ species and 239 specimens.

VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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ALLAN HANCOCK PACIFIC EXPEDITIONS

276

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ALLAN HANCOCK PACIFIC EXPEDITIONS

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ALLAN HANCOCK PACIFIC EXPEDITIONS

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

288

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NO. 2 HARTMAN : QUANTITATIVE SURVEY 289

SAN PEDRO SLOPE

The San Pedro (SP) slope, in depths below 60 fms, is an oceanward
down-sloping extension of the San Pedro shelf. It is dissected by the San
Gabriel sea valley; near its outer limits it surrounds the Lasuen sea
mount and Sixmile bank. One of its greatest depths, +40 fms, is attained
north of Sixmile bank (no. 213 on Map, Pt. 1). Its terrain is overall
irregular, compared with that of adjacent areas. Surface sediments are
mainly silt, mud and clay, with rocky rubble and gravel in the vicinities
of the sea mounts.

The fauna in the soft bottoms is diversified and patchy. Biomass
values at depths of 90 to 100 fms approach 100 grams to a sample; these
values diminish to a fraction of a gram at greatest depth, showing a more
or less steady decline with depth. Rocky outcrops occur in 170 and 180
fms near no. 174 (see Map, Pt. 1), and in 240 fms at no. 197; these are
also the sites of siliceous sponge and conspicuous representatives of echi-
noderms (Sta. 2987, 2416). The values of specific diversity, by increasing
depth, are illustrated :

Size of
Depth sample No. of No. of
Sta. no. in fms. in cuft species specimens
2292 60 0.81 87* 5767
5746 68 113 O72 822
5747 90 0.88 98 1546
4778 100 2.96 40 iaped
2423 IVE: [eis 42+ 107*
2884 190 1.93 23 123
2306 ZS 2.64 29° 158+
2342 230 0.31 29 146
2372 230 0.81 36* 203
2898 240 Het 3 26 13
2894 253 0.81 17 3,
2893 265 2.58 Zoe a
2886 270 2.60 20; 49
2635 278 Z.99 40* iB)
2644 310 5.74 21 37
2901 S12 3.02 pH | 40
2900 345 3.08 23 62
2844 362 S215 7 67
2370 366 2:83 17 47
2368 385 2.26 17 37
2369 390 3,33 De 52
2440 415 3.27 19 We,
2802 420 2.64 26 48
2836 430 2.88 15 $4

2229 440 0.63 20 48

290 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

The following samples, numbered | to 31 on the charts, come from
the San Pedro slope, in depths of 60 to +46 fms. Polychaeta, echinoderms
and mollusks are listed on the charts by increasing depths; others are
summarized in the following analyses.

1. Sta. 2292 (no. 104). Upper end of San Gabriel sea valley, in 60
fms. OPG took 0.81 cuft of fine grayish green sand, with many animals.
In addition to those named on the chart, the following were present:
Harenactis (1), Monobrachium (2 colonies), a nemertean (2), a tur-
bellarian (1), four amphipods (113), an isopod (2*), ostracods (many)
and a mysid (1). The largest species was Lumbrineris bicirrata; the most
abundant were Amphiodia urtica (177), Adontorhina cyclia (78), Ali-
gena sp. (74), Amphipholis squamata (53), and Chloeia pinnata (47).

Numbers of species and specimens totalled :

polychaetes 53 species, 350 specimens
echinoderms 11 310
mollusks 15 19]
crustaceans 6* b22F
others 2 5)

otal: 87+ species, 976* specimens

2. Sta. 5746 (near no. 129). SW of Newport, in 68 fms. OPG took
1.13 cuft of light green, foraminiferan sand with many animals. In addi-
tion to those named on the chart, the sample contained a nemertean (3),
a sipunculid (1), 16 amphipods (86), three isopods (7), two or more
tanaids (20) six ostracods (236) and a juvenile crab (1). The largest
individual was Antiplanes ?perversa, and the most abundant species were
Onuphis nebulosa (135), Tharyx tesselata (36), and Magelona pacifica
(20).

Numbers of species and specimens totalled :

polychaetes 48 species, 429 specimens
echinoderms 2 3

r10llusks 6 13
crustaceans 33 369
others 3 8

Total: 92 species, 822 specimens

3. Sta. 5747 (near no. 123). 9.2 mi south of Los Angeles light, in
90 fms. OPG took 0.88 cuft of very fine green sand. Wet weight of the
larger animals was 96.7 grams, of which echinoids were 62.7, ophiuroids
17.0, polychaetes 16.6, and all others less than 0.5 grams. In addition to

NO. 2 HARTMAN : QUANTITATIVE SURVEY 291

those named on the chart, the sample contained Mlonobrachium parasitum
(11 colonies on Pseudopythina chacei), a nemertean (2), a polyclad (1),
a sipunculid (4), ten amphipods (137), three isopods (11), tanaids (8),
three ostracods (about 400), and four cumaceans (32). Largest species
were Nephtys glabra and brissopsid urchins ; most abundant were Chloeia
pinnata (102) and Amphiodia urtica (100). Peak numbers were at-
tained also by Tharyx monilaris (70), Paraonis gracilis (58), Pista
moorei (36), and Adontorhina cyclia (35).

Numbers of species and specimens totalled:

polychaetes +6 species, 667 specimens
echinoderms 7 138
mollusks 18 130
crustaceans 23 593
others 4 18

gral 98 species, 1546 specimens

4. Sta. 4778 (no. 184). S of Laguna Beach, in 100 fms. OPG took
2.96 cuft of green mud. Animals included 8 amphipods (20), an isopod
(3), a cumacean (3) and an ostracod (1), in addition to those named
on the chart. Largest individuals were MJolpadia intermedia, Pista dis-
juncta and Travisia pupa. Most abundant or conspicuous were Chloeia
pinnata (56), Amphiodia urtica (44), and Pectinaria californiensis (37).

Numbers of species and specimens totalled :

polychaetes 21 species, 137 specimens
echinoderms 6 5D
mollusks Zs 8
crustaceans 11 2h

Total: 40 species, 227 specimens

Sub. 4. Sta. 2177 and 2438 (no. 267). Outermost end of the San
Pedro area, in 172 and 153 fms. OPG took sandy mud (2177) and rock
(2438) with attached tubes of unnamed polychaetes, representatives of
a polynoid, a sphaerodorid and a maldanid. ‘he sample from sandy mud
contained many polychaetes with Chloeia pinnata the most conspicuous ;
three kinds of echinoderms, Amphiodia urtica (18), Amphiura arcystata
(1), and Brissopsis pacifica (5), and five mollusks: Cadulus tolmiei (8),
Cardita ventricosa (7), Cyrilla minuta (41), Solamen columbianum

(2) and Tellina carpenteri (16).
Sub. 4. Sta. 2987 (no. 174). 13.7 mi SE of Los Angeles light, in

177-189 fms. The biological dredge took rocks and echinoderms with
siliceous sponge and a lamprey eel.

292 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

5. Sta. 2416 (no. 174). 13.5 mi SE of Los Angeles light, in 180
fms. The biological dredge took large boulders and many echinoderms
(see chart). Most abundant were Ophiopholis bakeri (42) and Ophia-
cantha diplasia (22).

6. Sta. 2884 (no. 154). 5.6 mi SW of Newport pier, in 190 fms.
OPG took 1.93 cuft of gray-green mud. Present were a ceriantharian
(1), an anemone with warty epithelium (2), ?drynchite (1), small
crustaceans (not examined), and those named on the chart. Largest
species was Pista disjuncta, and most abundant Chiloeia pinnata (70).

Numbers of species and specimens totalled :

polychaetes 12 species, 103 specimens
echinoderms - 11
mollusks + 5
others 3 -
Total: 23 species, 123 specimens

7. Sta. 2306 (no. 98). 6.2 mi from Pt. Fermin light, in 215 fms.
OPG took 2.64 cuft of mud and gray clay with large animals. In addi-
tion to those named on the chart, there were a few small crustaceans.
Largest species was Asychis disparidentata, and most abundant Chloeia
pinnata (51). Total numbers were estimated at more than 38 species,
and 175 specimens.

8. Sta. 2342 (no. 262). 5.9 mi E of East End, SCI, in 230 fms.
OPG took 0.31 cuft of sandy mud with sticky oil globules. The sample
contained siliceous sponge, a nemertean (1), a tongue of a large echi-
uroid (1), amphipods (+), an anthurid isopod (1), ostracods (32), in
addition to those named on the chart. Largest species was Nephtys
assignis, and most abundant Chloeia pinnata (46).

Numbers of species and specimens totalled :

polychaetes 16 species, 73 specimens
echinoderms 2 9
mollusks 5 24+
crustaceans is ey
others 3 3

9. Sta. 2372 (no. 199). 16 mi NE of East End, in 230 fms. OPG
took 0.81 cuft of gray sandy mud, with many foraminiferans, small tes-
selated sponge balls, a nemertean (3), a polyclad (1), amphipods (14),
an anthurid isopod (6), two ostracods (27), a small crab (1), in addi-
tion to those named on the chart. Largest species was Nephtys punctata,

NO. 2 HARTMAN : QUANTITATIVE SURVEY 293

and most abundant were Praxillella affinis pacifica (26), and Tellina car-
penteri (24).

Numbers of species and specimens totalled:

polychaetes 23 species, 102 specimens
echinoderms 1 +
mollusks 5 42
crustaceans = 48
others 3 y|

Total: 37* species, 203 specimens

10. Sta. 2898 (no. 197). 12.7 mi SW of East End, in 240 fms. OPG
took 1.13 cuft of blue-green mud with rocky rubble and shale. In addi-
tion to those named on the chart the sample contained a smooth anemone
(1), and a sand-covered one (1), a nemertean (1), an amphipod (2),
a caprellid (7), a gnathid isopod (1). Largest individual was Thelepus
setosus ; none was outstandingly abundant.

Numbers of species and specimens totalled :

polychaetes 14 species, 35 specimens
echinoderms 1 8
mollusks 5 17
crustaceans 3 10
others 3 3

Total: 26 species, 73 specimens

11. Sta. 2894 (no. 219). 11.5 mi SW of Newport pier, in 253 fms.
OPG took 0.81 cuft of gray-green, sandy mud with few animals. In addi-
tion to those named on the chart, there were a nemertean (3), an amphi-
pod (3) and a cumacean (1). Largest animal was a nemertean, and
most conspicuous Onuphis nebulosa (6).

Numbers of species and specimens totalled :

polychaetes 11 species, 19 specimens
mollusks 3 11
crustaceans 2 +
other 1 3

Total: 17 species, 37 specimens

12. Sta. 2895 (no. 200). 9.9 SW of Newport pier, in 265 fms.
OPG took 2.58 cuft of grayish green mud with arenaceous foramini-
ferans, small tesselated sponge balls, a large nemertean (2), an echiuroid
(1), amphipods (5), two isopods (2), in addition to those named on the
chart. Largest individuals were an echiuroid and a nemertean; none was
outstandingly abundant.

294 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled:

polychaetes 13 species, 31 specimens
echinoderms 2 2
mollusks 5 12
crustaceans af 7

Total: 23* species, 52 specimens

13. Sta. 2886 (no. 201). 9.2 mi SW of Newport pier, in 270 fms.
OPG took 2.6 cuft of grayish green mud. A large nemertean (2), an
echiuroid (1), amphipods (12), a cumacean (1), an ostracod (1), and
a small enteropneust, were present in addition to those named on the
chart. Largest animals were Brissopsis pacifica and a nemertean. None
was abundant.

Numbers of species and specimens totalled :

polychaetes 13 species, 18 specimens
echinoderm 1 1
mollusks wh 12
crustaceans 5 14
others 3 +

Total: 29* species, +9 specimens

14. Sta. 2635 (no. 249). S of Lasuen sea mount, in 278 fms. OPG
took 2.99 cuft of greenish yellow and black, marbled clay, with small
rocks and considerable rubble. Screenings contained small tesselated
sponge balls, two anemones (5), a nemertean (2), an echiuroid (2),
amphipods (6), a shrimp (1), in addition to those named on the chart.
Largest species was Notomastus magnus; none was notably abundant.

Numbers of species and specimens totalled :

polychaetes 26 species, 38 specimens
echinoderms 6 13
mollusks 2 8
crustaceans Os il
others + 9

Motal: 40* species, 75 specimens

15. Sta. 2644 (no. 222). NE of Lasuen sea mount, in 310 fms.
Campbell grab took 5.74 cuft of greenish gray clay with foraminiferans.
In addition to those named on the chart, there were a large nemertean
(1), 2?drynchite (1), and an amphipod (6). Largest individuals were a
nemertean and an echiuroid ; none was abundant.

NO. 2 HARTMAN : QUANTITATIVE SURVEY 295

Numbers of species and specimens totalled :

polychaetes 12 species, 19 specimens
echinoderms 2 S)
mollusks 4 6
crustaceans 1 6
others 2 3)

16. Sta. 2901 (no. 217). Lower end of San Gabriel sea valley, in
312 fms. OPG took 3.02 cuft of grayish green mud with many arena-
ceous and calcareous foraminiferans. Ihe sample contained a large ceri-
antharian, Cerebratulus (1), two amphipods (3), a caprellid (1), a
cumacean (1), and an enteropneust (2). Largest animals were Brissopsis
pacifica, a ceriantharian, and a nemertean; none was abundant.

Numbers of species and specimens totalled :

polychaetes 14 species, 22 specimens
echinoderms 5 i
mollusks 6 8*
crustaceans + 5
others 3 5

Total: 30 species, +7* specimens

17. Sta. 2900 (no. 231). E of Sixmile bank, in 345 fms. Grab took
3.08 cuft of mud. The sample contained those listed in the chart, together
with an anemone with wrinkled epithelium (3), a large red ceriantharian
(1), a red nemertean (1), a sipunculid (3), two amphipods (3), a
caprellid (1) and a cumacean (1). Largest animal was a ceriantharian,
and none was abundant.

Numbers of species and specimens totalled :

polychaetes 11 species, 33 specimens
echinoderms 5 9
mollusks 2 7
crustaceans 3 5
others 7 8

Total: 25 species, 62 specimens

ISe Sta. 2350 (no. 241)2 69/5 mi SE of Long Pt. SCI; in’ 350. ims:
OPG took 0.1 cuft of rocks, mud, sand, rubble and siliceous sponge. All
animals were small, with the largest Chaetozone and Lepidonotus. The
most abundant was O phiacantha sp. (10). Numbers of species and speci-
mens included polychaetes with 9 species and 11 specimens, and _ echi-
noderms with 3 species and 18 specimens.

296 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

19. Sta. 2844 (no. 233). SW of Lasuen sea mount, in 362 fms.
OPG took 3.15 cuft of sticky grayish green mud with many calcareous
foraminiferans, long sponge spicules and ophiuroid fragments. In addi-
tion to the species named in the chart, there were an anemone (1), a
nemertean (4), and an amphipod (1). Largest species was A mphiura

arcystata, and most abundant Aricidea uschakowi (16) and Tharyx spp.
(23ne

Numbers of species and specimens totalled :

polychaetes 11 species, 55 specimens
echinoderms 2 3
mollusk 1 3
crustacean Il 1
others 2 2

20. Sta. 2370 (no. 232). Lower end of San Gabriel sea valley, in
366 fms. OPG took 2.83 cuft of grayish green mud with foraminiferans,
small amount of siliceous sponge, and many small animals. In addition
to those named on the chart, an amphipod (3) and a cumacean (1)
were present; there was no large individual; most abundant was
Tharyx sp.

Numbers of species and specimens totalled :

polychaetes 11 species, 26 specimens
echinoderm 1 3
mollusks 3 14
crustaceans 2 +

Total: 17 species, +7 specimens

21. Sta. 2640 (no. 241). S of Sixmile bank, in 370 fms. OPG took
about a liter of brown-black, friable rock, clayey gravel, animals and
siliceous sponge spicules. In addition to the species named on the chart,
there were present an isopod (3), Alunida quadrispina (galatheid crab)
(1), and a colonial tunicate. Largest animal was the crab, and none was
abundant. Total number of species 15, and specimens 27.

22. Sta. 2368 (no. 242). S of Sixmile bank, in 385 fms. OPG took
2.26 cuft of sandy mud, rocks, siliceous sponge and foraminiferans. A
dark, warty anemone was present, an amphipod (3), and a tanaid (2),
in addition to those named on the chart. Largest individual was Chloeia
pinnata, and most abundant were Lumbriclymene lineus (9) and Tharyx

Sp.(9))e

No. 2 HARTMAN : QUANTITATIVE SURVEY 297

Numbers of species and specimens totalled :

polychaetes 11 species, 28 specimens
mollusks 2, Z
crustaceans 2 5)
others 2 2

Total : 17 species, 37 specimens

23. Sta. 2369 (no. 243). 3 mi SE of Sixmile bank, in 390 fms. OPG
took grayish green mud with siliceous sponge, calcareous foraminiferans,
and many small animals. In addition to those named on the chart there
were two amphipods (8), and a pycnogonid (1). None was large and
none abundant.

Numbers of species and specimens totalled :

polychaetes 11 species, 26 specimens
echinoderms 4 11
mollusks + 6
crustaceans 2 8
other 1 |

Total’: 22 species, 52 specimens

24. Sta. 2440 (no. 244). Lowest end of San Gabriel sea valley, in
415 fms. Grab took 3.27 cuft of greenish gray clay and mud, with many
foraminiferans, minute mud balls, tube fragments of Phyllochaetopterus
limicolus and a few animals. An anemone measuring 10 by 28 mm, a
nemertean (1), four amphipods (5), an isopod (1), were present in ad-
dition to those named in the chart. The largest individual was an
anemone.

Numbers of species and specimens totalled :

polychaetes 7 species, 34 specimens
echinoderm 1 12
mollusks 4 18
crustaceans 5 6
others 2 2

Total: 19 species, 72 specimens

25. Sta. 2352 (no. 227). W of Sixmile bank, in 420 fms. OPG took
2.5 cuft of grayish green mud with considerable siliceous sponge and
orbicular foraminiferans. In addition to those named on the chart, the
lot included a small sea pen. The largest individual was Thelepus setosus,
in a tube 170 mm long by 12 mm across; it was externally adorned with
attached foraminiferans and tubes of Protis.

298 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled:

polychaetes 3 species, 17 specimens
echinoderms S) +t
mollusks 3) 3
other 1 1
Total: 10 species, 25 specimens

26. Sta. 2802 (no. 194). N of Sixmile bank, in 420 fms. Grab took
2.65 cuft of bluish greenish gray mud with siliceous sponge and foramini-
ferans. In addition to those named on the chart, there was a large and
a small anemone, a nemertean (1), and a cumacean (1). Largest species
were Melinnexis pacifica and Myxoderma platyacantha.

Numbers of species and specimens totalled:

polychaetes 13 species, 38 specimens
echinoderms 2 3
mollusks 2 3)
crustacean | 1
others 2 3

Total: 20 species, +8 specimens

27. Sta. 2859 (no. 227). W of Sixmile bank, in 425 fms. OPG took
2.58 cuft of greenish gray sand and mud with many foraminiferans and
much siliceous sponge. The lot contained a sea pen (2), a nemertean
(1), a sipunculid (1), an ostracod (1) and Callianassa goniophthalma
(1). The large individual was the ghost shrimp.

Numbers of species and specimens totalled :

polychaetes 10 species, 12 specimens
mollusks 3 6
crustaceans 2 Wh
others 3 5)

Total: 18 species, 23 specimens

28. Sta. 2836 (no. 145). Eastern sill of San Pedro basin, in 430
fms. OPG took 2.88 cuft of greenish gray sticky mud, with many fora-
miniferans, tubes of Phyllochaetopterus and diversified animals. In addi-
tion to those named in the chart, a ghost shrimp (1), an isopod (1), and
a small enteropneust (1) were present.

Numbers of species and specimens totalled :

polychaetes 8 species, 2+ specimens
echinoderm 1 1
mollusks 3 3
crustaceans 2 2
other 1 1

Total: 15 species, 31 specimens

NO. 2 HARTMAN : QUANTITATIVE SURVEY 299

29. Sta. 2229 (no. 213). N of Sixmile bank, in 440 fms. OPG took
0.63 cuft of sandy clay, gravel, rubbly rocks, with siliceous sponge, fora-
miniferans, and diversified animals. A gorgonian and sea pen were pres-
ent, in addition to those named in the chart.

Numbers of species and specimens totalled :

polychaetes 17 species, 44 specimens
echinoderm | 2
others Z 2

‘Total: 20 species, +8 specimens

Sub 29. Sta. 2341 (no. 263). 7.6 mi E of East End, SCI, in 440
fms. OPG took 0.1 cuft of fine sandy mud with almost no life; only a
small ophiuroid and fragments of a maldanid and Tharyx were present.

30. Sta. 2839 (no. 212). NW of Sixmile bank, in +46 fms. Camp-
bell grab took 3.4 cuft of greenish gray sticky mud with foraminiferans,
and siliceous sponge. In addition to those named on the chart, a nemer-
tean (1) and an enteropneust (1) were present. Solenogasters, with 3
species and 12 specimens, were unusual.

Numbers of species and specimens totalled :

polychaetes 7 species, 17 specimens
echinoderms 2 2
mollusks 5 14+
others 2 my
Wotali: 16 species, 35 specimens

31. Sta. 2837 (no. 211). NW of Sixmile bank, in 454 fms. Campbell
grab took 3.08 cuft of greenish gray sticky mud with foraminiferans,
tubes of Phyllochaetopterus limicolus, Protis pacifica and siliceous sponge.
In addition to those named on the chart, a sipunculid (1) and a com-
pound ascidian (2) were present.

Numbers of species and specimens totalled :

polychaetes 5 species, 18 specimens
echinoderm | ]
mollusks 3 4
others 2 3

Motali: 11 species, 26 specimens

VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

310

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

312

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

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NO. 2

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316 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

NEWPORT SHELF

The narrow Newport Shelf (NE) has its shoreward end between
Newport Bay and Laguna Beach; it is bounded on the west by Newport
canyon (Hartman, 1963, 27) and merges oceanward and in greater depth
with the San Pedro slope (see Map 1). Sediments of the bottom are sand
and silt mixed with varying amounts of shelly and other biological debris.
Faunal diversity changes with location, depth and size of sample, as
shown by the following analysis:

Size of

Depth sample No. of No. of
Sta. no. in fms. in cuft species specimens
2745 8 0.27 47 190
5877 9 0.69 59 306
4777 10 0.88 70 437
4871 Oy egy 101 701
5354 31 4 82 1250
5092 50 hes 78 1134
4872 Sy 3.30 61 540

Most of the benthic animals of this shelf are infaunal and thus not
visible in surface views except as pores, small mounds or extended parts
of animals. Eight samples are analyzed, with three of the major groups
of species named on the following chart. Greater depths than 52 fms are
referred to the San Pedro slope area.

1. Sta. 2745 (no. 110). 0.1 mi from end of Balboa pier, in 8 fms.
OPG took 0.27 cuft of gray sand and shelly fragments and numerous
branches of Cellaria (bryozoans). In addition to those named on the
chart, Harenactis sp. (28), a nemertean (3), amphipods (45), a caprel-
lid (1), cumaceans (14), ostracods (5) and a small crab (6), were pres-
ent. The largest species were Diopatra ornata and Chaetopterus variope-

datus. None was outstandingly abundant.

Numbers of species and specimens totalled:

polychaetes 34 species, 77 specimens
echinoderms 3 3

mollusks 3 7
crustaceans Sy (Bl

others 4 a2

Total: 49* species, 190* specimens

NO. 2 HARTMAN : QUANTITATIVE SURVEY 317

2. Sta. 4877 (no. 134). About half way between Newport Bay and
Laguna Beach, in 9 fms. OPG took 0.69 cuft of medium dark gray sand.
Wet weight of macroscopic animals was 19.9 grams, of which polychaetes
comprised 10.7, ophiuroids 4, Astropecten (2) 1.7, nemerteans 1.4, and
anemones 1.3 grams. In addition to the species named on the chart, a
white anemone with flat base (8), small anemone (8), Cerebratulus sp.
(2), a polyclad (2), a sipunculid (15), about ten species of amphipods
(107), an isopod (1), cumaceans with six species (18), ostracods with
four (18), ashrimp (1), and Branchiostoma sp. (3), comprised the lot.
The largest individual was Nothria iridescens, the most conspicuous
Owenia f. collaris and Onuphis eremita, and the most abundant Lumbri-
neris cruzensis (25) and Glycera oxycephala (21).

Numbers of species and specimens totalled :

polychaetes 25 species, 101 specimens
echinoderms + 33
mollusk 1 |
crustaceans 25 133
others 6 38

Total: 59 species, 306 specimens

3. Sta. 4777 (near no. 185). S of Laguna Beach, in 10 fms. OPG
took 0.88 cuft of fine gray to black sand, flocculent debris having an odor
of hydrogen sulfide, and many animals. Total wet weight of animals was
21.2 grams, of which polychaetes comprised 12.4, mollusks 4.6, an 4 stro-
pecten 2.0, and crustaceans 1.0 grams. In addition to those listed on the
chart, the sample had two kinds of nemerteans (18), Glottidia albida
(2), 18 species of amphipods (284), a tanaid (23), two kinds of cuma-
ceans (6), an epinebalian (1), an ostracod (2), Callianassa (3), a pag-
urid (1) and a crab (4). Largest individuals were Astropecten and
Lytechinus ; and most conspicuous was Diopatra ornata.

Numbers of species and specimens totalled :

polychaetes 26 species, 75 specimens
echinoderms ) i
mollusks 8 1]
crustaceans 30 324
others 3 20

Total: 70 species, +37 specimens

4. Sta. 4871 (near no. 185). 2.7 mi from Dana Pt., in 27 fms.
OPG took 1.32 cuft of green silt. Wet weights of animals totalled 54.5
grams, of which polychaetes comprised 22.2, ophiuroids 19.6, tunicates

318 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

5.7, mollusks 2.8, and decapods 1.8 grams. In addition to the species
named on the chart, the lot comprised a small sea-whip, a hydroid stalk,
several small nemerteans, many slender tubes of phoronids, 16 kinds of
amphipods (115), an isopod, a tanaid, two kinds of cumaceans (3), three
kinds of ostracods (40), Callianassa sp. (4), Pinnixa sp. (4), Randallia
sp. (1), Scalpellum (1), a pycnogonid (1), an enteropneust (5) and a
tunicate (2). There was no conspicuously large specimen ; most abundant
were Amphiodia urtica (240) and Pectinaria californiensis (40).

Numbers of species and specimens totalled :

polychaetes 35 species, 175 specimens
echinoderms 8 267
mollusks 21 56
crustaceans BY 173
others 5 SUF

Total: 101 species, 701+ specimens

5. Sta. 2747 (near no. 134). Between Newport Bay and Laguna
Beach, in 30 fms. Biological dredge took a small sample in kelp and sandy
bottom. The most conspicuous species were Pectinaria californiensis and
Platynereis bicanaliculata.

6. Sta. 5354 (near no. 185). 7.5 mi from Newport jetty light, in 31
fms. OPG took 1.51 cuft of green sandy silt with many animals. Wet
weights totalled 88.9 grams, of which ophiuroids comprised 57.5, poly-
chaetes 17.5, holothurians 10.8, nemerteans 1.2, and others less than a
gram. In addition to the species named on the chart, there were present
a cenantharian (3), a broad white polyclad (1) and another with encir-
cling black eyes (1), nemerteans (12), a phoronid (10), 11 kinds of
amphipods (106), two cumaceans (2), four ostracods (99), two pinnixid
crabs (5), and an enteropneust (5). Largest individuals were Molpadia
intermedia and Travisia pupa. Most abundant were Amphiodia urtica
(595), Pholoe glabra (112), Lumbrineris cruzensis (38) and Pectinaria
californiensis (30).

Numbers of species and specimens totalled :

polychaetes 41 species, 358 specimens
echinoderms 5 618
mollusks 10 29
crustaceans 19 DANS
others 7 32

Total: 82 species, 1250 specimens

NO. 2 HARTMAN : QUANTITATIVE SURVEY 319

7. Sta. 5092 (near no. 185). 1.4 mi from Newport jetty light, in
52 fms. OPG took 3.3 cuft of sand and shell fragments. Wet weight was
131.4 grams, of which ophiuroids comprised 69.2, Molpadia 32.5, poly-
chaetes 25.2, echiuroids 2.6, mollusks 1, and others less than 1 gram. In
addition to the species listed on the chart, the lot contained a small white
anemone (1), ?Harenactis sp. (1), Monobrachium parasitum colonies
(6), a polyclad (1), nemerteans (5), sipunculid (1), Arynchite (1),
11 kinds of amphipods (151), two kinds of isopods (3), ostracods (4),
two cumaceans (4), and small colonies of Pyrosoma (5). Largest indi-
viduals were Molpadia intermedia and Travisia pupa, and most abun-
dant was Amphiodia urtica (648), followed by Amphipholis squamata
(72) and Pectinaria californiensis (36).

Numbers of species and specimens totalled :

polychaetes 34 species, 161 specimens
echinoderms ih 747
mollusks 12 42
crustaceans 17 162
others 8 20

Total: 78 species, 1132 specimens

8. Sta. 4878 (near no. 185). 1.4 mi from Newport jetty light, in 52
fms. OPG took 3.3 cuft of sand and shell fragments. Wet weights
totalled 58.4 grams, of which polychaetes were 36.5, ophiuroids 15.6,
mollusks 2.8, nemerteans 1.1, and others each less than 1 gram. In addi-
tion to the species named on the chart, the sample contained a small sea-
whip, ?Harenactis (3), Listriolobus pelodes (9), eight kinds of amphi-
pods (50), an isopod (2), an ostracod (2), a pinnotherid crab (2), a
small crab (1), and an enteropneust (3). Largest individual was Pista
disjuncta, and most abundant species were 4 mphiodia urtica (252), Pec-
tinaria californiensis (43), and Pista disjuncta (20).

Numbers of species and specimens totalled :

polychaetes 31 species, 168 specimens
echinoderms 4 P|
mollusks 9 liv
crustaceans 12 57
others 5} ZA

Total: 61 species, 540 specimens

320 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta in the Newport shelf,

showing order of occurrence in 8 to 52

+ indicates presence, - absence.

Species represented by more than 10 specimens are

Amaeana occidentalis . A
Anaitides sp. 5 5
Aricidea lopezi . A :
Aricidea neosuecica

Chaetopterus variopedatus

Chaetozone corona ‘

Diopatra ornata

Glycera convoluta 0 é
Goniada littorea . C

Gyptis a. glabra . :

Haploscoloplos elongatus .

Harmothoe lunulata 6 :
Harmothoe priops . . 5
Heterospio catalinensis .

Laonice cirrata

Lumbrineris spp. . 9
Magelona sacculata “ 0

Mediomastus californiensis

Nephtys spp. . 3 C 2
Nereis procera 5 5 o

Nothria iridescens fe

Odontosyllis phosphorea .

Owenia f. collaris :

Pareulepis fimbriata . 0

Phyllochaetopterus limicolus

Prionospio malmgreni
Sthenelais tertiaglabra .
Enigiglgts ue ee. kt

Telepsavus costarum

1

+

2

3

fms.

indicated by #*.

y

+

5

6

VOL. 19

No. 2 HARTMAN:

QUANTITATIVE SURVEY

Polychaeta in Newport shelf (continued)

Thalenessa spinosa c :
Tharyx spp. . : . 5
Exogone uniformis c :

Glycera oxycephala ¢ c
Glycinde armigera 5

Goniada brunnea . 3 :
Loimia medusa ‘ - :
Lumbrineris nr. acuta é

Lumbrineris californiensis

Lumbrineris cruzensis
Nephtys caecoides . c
Onuphis eremita . c

Pholoe glabra : : c

Pista disjuncta . : C
Prionospio pinnata : 6
Prionospio, another sp. .
Rhynchospio sp. . . *
Spiophanes missionensis .

Amphicteis scaphobranchiata

Axiothella rubrocincta F
Brada pilosa . Ce :
Glycera americana c :
Harmothoe sp. 5 ° 5

Pherusa neopapillata . -

Pista cf. cristata E :

Platynereis bicanaliculata

Poecilochaetus johnsoni .

Pseudopotamilla occelata .

Schistocomus hiltoni . c
Tharyx multifilis :

Ancistrosyllis tentaculata

al

. : +

2

+

+

3

+

4

32

322 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta in Newport shelf (continued)

Anotomastus gordiodes :

Aricidea sp. . C 6
Artacamella hancocki . ‘

Brada glabra . : 5 C

Ceratocephala c. americana

Cossura candida . . $
Glycera capitata . : x
Loandalia fauveli ‘ 2

Magelona pacifica 6 B

Myriowenia californiensis
Naineris uncinata C 2
Paraonis gracilis 7

Pectinaria californiensis
Pilargis berkeleyi C 3
Polydora sp. . < : :

Praxillella a. pacifica .

Rhodine bitorquata é 6
Sternaspis fossor ; ‘

Sthenelanella uniformis .
Terebellides stroemii

ampharetid C C

Asabellides ?lineata or A. sp.

Hyalinoecia juvenalis 5
Isocirrus longiceps
Notomastus magnus c
Pherusa capulata . C :
Travisia sp. . : 3 :
Anaitides, checkered . :
Ancistrosyllis ef. rigida

Aricidea uschakowi ;

Chloeia pinnata .

il

2

3

4

+

VOL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta in Newport shelf (continued)

Drilonereis sp. . :

Eunice americana .

Hesperonoe sp. . .
Nephtys ferruginea :

Notomastus tenuis
Praxillella gracilis .
Spio punctata : c
Travisia pupa “
Brada pluribranchiata
Chone gracilis : :

Leanira sp. . c :
Lumbrineris bifilaris
Megalomma sp. :
Myriochele gracilis

Onuphis parva . :
Lepidasthenia sp. 6
Lumbrineris bicirrata
Ninoe gemmea . : 5
Panthalis pacifica 9

Travisia brevis . :

iL

2

3 4

Echinodermata in the Newport shelf,

showing order of occurrence in 8 to

Amphiodia psara . :

Amphiodia urptLCa s F

Astropecten californicus .

Amphipholis squamata .
Dendraster excentricus
Amphiodia digitata C
Astropecten sp. .

Lytechinus anamesus .

Amphiura arcystata C

we
“

52 fms.
ale

= +

= *

+ =

+ =

+ te

; +

324 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL.

Echinodermata in Newport shelf

Ophiothrix spiculata .

Ophiura lutkeni .

Pentamera pseudopopulifera

Amphioplus strongyloplax .

Leptosynapta albican

Molpadia intermedia

Ss

Amphiacantha amphacantha .

holothurian .

Epitonium tinctum
Rochefortia tumida
Turbonilla sp. 5
Olivella baetica .

Crepidula excavata

?Lacuna sp.

iL

(continued)

2 3 4 5 6 U

Mollusca in the Newport shelf,
showing order of occurrence in 8 to 52 fms.

Nassarius perpinguis .

Nitidella carinata
Olivella ?pedroana
Polinices lewisii
Solen ?sicarius

Tellina modesta

Acteocina intermedia .

Axinopsida serricata

Bittium sp. . :

Caduilus) spr = :

Compsomyax subdiaphana

Crenella columbiana
Cylichna diegensis
Lyonsia californica

Macoma yoldiformis

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‘ + 2 2 Pe
+ = = =

+ = = 2

+ 2 = 2

19

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Mollusca in Newport shelf (continued)

il 72 3
Mangelia sp. . A 5 2 3 = 5 5 5 5
Megasurcula ?carpenteriana 3 ° ° C , .
Modiolus sp. . 5 : : 5 : 6 : 5 :
Nuculana taphria . : = : 5 = : : :
Pandora sp. . “ 2 : : : . ° . A
?Rochefortia sp. . ; 5 4 : 0 5 é
Saxicavella pacifica . ; 3 c 5 : ‘ é
solenogasters C 6 . i 5 : 5 Q
Thyasira sp. . : 5 ° 5 5 : C 3 C
Volvulella tenuissima 5 : : ; : 6 -
Cylichna attonsa . 5 : : = 6 : 3 4
Epitonium sp. c : ; 5 3 a 5 é
Nucula tenuis 5 : ; C ; : ; 5 S
Pandora bilirata . a 4 C 6 6 : 5 5
Rochefortia aleutica . cS a : é : 5 .
Adontorhina cyclia 5 : 3 5 . 6 5 A
?Macoma sp... : : 7 0 : é : : :
Pseudopythina chacei . 5 : 5 5 : : 3
?Turbonilla sp. . - : 4 : : - 6 ;
Balcis sp. 5 5 6 : 6 : : : : 5
Cardita ventricosa : 5 5 : : 3 3
Cuspidaria sp. - C és 6 - 5 3 5

Saxicavella pacifica . ; 5 é : : 6

Gn

326 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

SUBMERGED SEA MOUNTS IN THE
SAN PEDRO CHANNEL

‘Two sea mounts (LS) in the San Pedro Channel have been biologi-
cally explored. The larger Lasuen sea mount is on the eastern and the
smaller Sixmile bank on the western side of the channel (see Map 1).

‘The Lasuen sea mount is located 13 nautical miles west of Newport
and beyond the outer end of Newport Canyon; its base is in depths of
about 250 fms and it rises to 40 to 68 fms at its top. It measures about
3.25 by 5.33 miles and covers more than 16 square miles. Six samples
were recovered and analyzed. The sediments at the top are rocky, calca-
reous, with much debris; downward the sediments contain much shelly
debris, mucoid stringy masses, flocculent debris; this gives way farther
down to gravelly debris and then sandy green mud. Siliceous sponge
occurs at its deepest perimeter. Ihe faunal components are richly diversi-
fied ; the most abundant and largest are echinoderms, coelenterates, mol-
lusks, polychaetes and crustaceans.

1. Sta. 2298 (no. 235). SW end of sea mount, in 68 fms. OPG took
0.37 cuft of shelly sand and a few rocks. In addition to the species listed
on the chart, there were single individuals of anemone, nemertean, sipun-
culid (2), oligochaete (1), and crustaceans (amphipods and isopods) in
sparse numbers. Largest individual was Lanice conchilega, and most
abundant species were Spio punctata (148), and Chloeia pinnata (16).

Numbers of species and specimens totalled :

polychaetes 33 species, 254 specimens
echinoderms + 2
mollusks 3 3
crustaceans fie os
others + 14

Total: 46* species, 285* specimens

2. Sta. 2297 (no. 236). Southeast end of Lasuen sea mount, in 181

fms. OPG took 0.68 cuft of large and small rocks, mud, gravelly debris,
with many small animals and a large brown, coarse sponge measuring
3 by 5 cm. In addition to those named on the chart, there were also a
sipunculid (2), an amphipod (2), a caprellid (1), and a tanaid (3).
None was abundant, and most were under-developed. ‘otal number of
species was estimated at less than 30, and specimens numbered about 50.

No. 2 HARTMAN : QUANTITATIVE SURVEY S27

3. Sta. 2843 (no. 220). Northern end of Lasuen sea mount, in 230
fms. OPG took 1.32 cuft of grayish green sandy mud with little rubble
and numerous animals. The lot included a small sea-whip, a nemertean
(2 large), amphipods (few), an isopod (1), a cumacean (1), ostracods
(3). Largest individual was Brissopsis pacifica and most abundant were
Nuculana conceptionis (16), and Lysippe annectens (15).

Numbers of species and specimens totalled :

polychaetes 20 species, 46 specimens
echinoderm 1 5
mollusks 8* 41
crustaceans jf le
others 2 3

Total: 36* species, 102+ specimens

4. Sta. 2887 (no. 221). Northeast end of Lasuen sea mount, in 284
fms. OPG took 2.89 cuft of gray-green mud with many foraminiferans
and small animals of few kinds. In addition to those listed on the chart,
an amphipod (3) and a cumacean (2) were present. Largest was a bris-
sopsid, and most abundant Nitidella permodesta (13). Most of the spe-
cies are those characteristic of slope depths,

5. Sta. 2299 (no. 234). Southwest end of Lasuen sea mount, in 360
fms. OPG took a full sample of dark gray oozy mud with siliceous sponge
and many kinds of animals. In addition to those listed, there were present
a nemertean (1), and a sipunculid (1). Largest individuals were Thal-
anessa spinosa and Maldane sarsi. None was conspicuously abundant.
Number of species totalled about 16, and specimens less than 20.

6. Sta. 2154-52 (near 228). Lasuen sea mount, in 300 fms. Biologi-
cal dredge, with tangles, pulling upslope on a rocky incline, recovered
coarse brown sponge, an alcyonarian, bryozoans encrusting rocks, a gala-
theid crab, and those species listed on the chart. Most, if not all, are
unique to this area, and may have their affinities with animals off the
deeper areas of Santa Catalina Island.

Sixmile bank is a broadly elongate seamount, 6.5 nautical miles north-
east of Avalon, Santa Catalina Island; it rises from a depth of 350 fms
to a flat top in 228 fms. It measures about 3.25 by 4.5 mi and covers an
area of about 15 square miles. It is located in the serial numbers 213, 214
and 228, 229. ‘Three samples were useful to indicate the kind of fauna.

Sta. 2204 (near no. 228), in 230 fms, was a dredged one and took
rocks with attached ophiuroid, Ophiacantha phragma, asteroids, and

328 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Psolus, also egg-capsules of a hag-fish, a chiton, a munnid crab, horny
brown sponges and associated bryozoans and polychaetes. The polychaetes
appear to be largely unknown species in the genera Euchone, Euphrosine,
Exogone, Glycera, Lagisca, Lumbrineris, Odontosyllis and Thelepus.

Sta. 2845 (no. 229), in 227 fms. OPG took 1.25 cuft of grayish
green sandy mud with small mud-balls, dark gravel, and wormlike ani-
mals. Largest individuals were a nemertean, Cerabratulus, a capitellid,
and a brissopsid, Brissopsis pacifica. Polychaetes included species in 4 m-
motrypane, ampharetids, capitellid, cirratulid, Cirrophorus furcatus,
Haploscoloplos elongatus, Laonice sp., Onuphis sp., Streblosoma sp. and
Terebellides stroemti. Mollusks were represented by Cadulus tolmiei and
solenogasters, and crustaceans by amphipods and an apseudid isopod.

Sta. 2228 (no. 228) in 293 fms. OPG took oozy mud with siliceous
sponge, orbicular foraminiferans, and metazoan animals including ophi-
uroids, gastropods, pelecypods, scaphopods, solenogasters, together with
more numerous small polychaetes and crustaceans (see Hartman, 1955:

136).

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta from Lasuen sea mount (1-5) and Sixmile bank (6)

showing order of occurrence in 68 to 360 fms.

+ indicates presence, - absence.
Species represented by more than 10 specimens are indicated by

Anaitides ?madeirensis : 5
Anobothrus gracilis . c : 2
Capitella capitata subsp. 3
Chaetozone armata :

Chloeia pinnata .

Euchone incolor . c 0 < .
Exogone uniformis 5 6 5
Glycera americana : : : C
Glycera cf. capitata . ; .
hesionid, unknown :

Lanice conchilega

Laonice cirrata

Lumbrineris latreilli 5 4 5
Lumbrineris spp. .

Lysippe annectens

Magelona ?californica

Magelona pacifica s : :
Nephtys, unknown sp.

Notomastus lineatus .

Notomastus magnus : : : :
Onuphis sp. . a : :

Peisidice aspera

phyllodocid . : ‘ : 3
Polycirrus sp. - C : : °
Polydora cf. websteri I é
Prionospio pinnata :
Protodorvillea gracilis . c 6

Questa caudicirra n. gen., n. SD.

Sphaerosyllis sp. - 5 :

1

+

2

+

3

4

o29

330 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Polychaeta from Lasuen sea mount and Sixmile bank (continued)

al 2 3 4 5 6

Spio punctata : c 5 Q 5 5 : : * - - = + =
Thalenessa spinosa : c 0 5 : 4 : + - - = - +
Tharyx tesselata . 3 ° : : ° 5 ° + - - - - -
?Typesyllis sp. .- 5 0 é 5 : : + - - - - =
Aricidea spp. 5 é 6 5 P : : é + = S ps =
Boccardia sp. : : C ; : 5 : 3 - + = = = E
Brada glabra . a 5 6 a . C : . 5 + + a = =
capitellid 5 é : : c : 5 5 : + = = = =
Cossura candida . : : ¢ : . : : : + - - = -
Dodecaceria concharum 5 4 9 : .: é 2 + = = E. =
Eunice sp. : : 5 : a c : : 7 + = = o =
GlyiceraNsp. 0 C - : : : : c ° a - - - -

Haploscoloplos elongatus . : . - : : : + + = = -

Lepidonotus caelorus . 6 : : 5 : : nF - - - -
maldanid . 5 A : 0 0 : 6 : : : + - - - -

Phyllochaetopterus limicolus . ; b 5 . < + + - - -

syllid : : : : 5 : 5 é : 3 + - - - -
Terebellides stroemii : 5 ; ‘ 5 ‘ : + - - - -
unknown annelid . 5 c : 5 : : é 5 + - - - -
Ammotrypane aulogaster ¢ o 5 C 6 6 5 o + - - -
Anaitides sp. 2 2 C é c : c : 5 : + - - -
Cirrophorus aciculatus 3 5 5 5 3 : 3 : + - -
Cirrophorus furcatus . 5 : = 6 : : f 5 + - - -
Drilonereis ?longa c : 5 6 : A 5 C + - - -
“Goniada brunnea . 5 é : E ‘ : : 3 5 + - - -
Gyptis a. glabra . : ° : : : : . : : + - - ~
Myriochele sp. .. : 6 : 5 : c c c : x - + -
Nephtys ?ferruginea . c C 5 : : 5 ¢ c 7 - - -
Paraonis gracilis : 3 5 é P ° A : + - + -
Potamethus sp. c : : : q : 2 . . : + = = =

Praxillelilal a. pacifiica . . S 4 F : : - + - - -

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta from Lasuen sea

Prionospio sp.

Tharyx sp. ° : 5
Aglaophamus sp.

Ammotrypane sp. . :
Aricidea lopezi . :

Caulleriella sp.
goniadid . 5 A :

Ancistrosyllis sp. 6

Eumida sp. . é c

Glycera branchiopoda .

Maldane sarsi -
Notomastus sp. cC

Gattyana brunnea n. sp.
Glycera tesselata :
Lagiseca nr. multisetosa
Lepidonotus caelorus .
Macellicephala remigata
?Nicomache, unknown .
serpulid fragment

terebellid, unknown .

mount

331

and Sixmile bank (continued)

al 2 3 4

: ‘ ; : : + =
+

3 é : +
: : . : +
+

: 4 4 +
5 : +

Echinodermata from Lasuen sea mount (1-5) and Sixmile bank
showing order of occurrence in 68 to 360 fms.

Amphiacantha amphacantha
Amphiodia urtica . :
Amphipholis squamata .
Leptosynapta albicans
Spatangus californicus
Amphioplus strongyloplax
Amphipholis pugetana
Ophiothrix spiculata .

ee
g a ae. ee

Se hie ed

: : We fas FS Se
a ee a

: f= =

+ - =

: oe

(6)

S52 ALLAN HANCOCK PACIFIC EXPEDITIONS

Echinodermata from Lasuen sea

Brissopsis pacifica . 6

brissopsids . C o 5
ophiuroid
echinoid . 5 ; C 2

Allocentrotus fragilis :

Amphiura seminuda :
Astrophiura marionae . :

Luidiastercalifornicus

Ophiacantha bairdi 6
Ophiacantha diplasia . g

Ophiocynodus corynetes C
Ophiopholis bakeri 3

Mollusca from Lasuen
showing order

Glycimeris subobsoleta C

Parvilucina tenuisculpta .
Tellina carpenteri .

Cadulus tolmiei . 5 6
Lepidopleurus nexus . 6
Limifossor sp. : : :
Saxicava arctica . :
Cardita sp. . é

Nuculana conceptionis ;

Nuculana hamata . ° .
Solamen columbianum .
solenogasters . . °
Dacrydium sp. 6 3

Nitidella permodesta .
Saxicavella pacifica .

sea mount (1-5) and
of occurrence in 68

al

2

3

+

4

VOL. 19

mount and Sixmile bank (continued)

5

Sixmile bank (6)
to: 360) fms

+

+

NO. 2 HARTMAN : QUANTITATIVE SURVEY 333

SANTA CATALINA ISLAND, LEEWARD SIDE
The leeward side of Santa Catalina Island (CL) has been sampled

on many occasions. Quantitative grab samples noted below were taken
along transects extending from East End, Avalon, Willow Cove, White
Cove-Long Point, Isthmus Cove, Howlands Landing, and West End,
leeward side. The details of kinds and abundance of animals are indi-
cated on the separate charts (below).

East End
1. Sta. 3611 (no. 259), in 30 fms. OPG took 0.56 cuft of green mud,

sand, strands of algae (Egregia) measuring to 11 feet long, and many
animals, especially amphipods, red-striped spirontocarid shrimps, poly-
chaetes, echinoderms and coelenterates. Polyclads, nemerteans and sipun-
culids were occasional. Encrusting bryozoans on kelps were varied. ‘he
most conspicuous animals were O phiothrix spiculata, Chaetopterus vario-
pedatus and Platynereis bicanaliculata; most abundant were amphipods
and Capitella capitata subspp. Total number of species was more than 50.

2. Sta. 5095 (near no. 259), in 40 fms. OPG took 0.81 cuft of fine
greenish brown sand. Largest individual was a red nemertean, Lepidas-
thenia ?virens, and most conspicuous was Sternaspis fossor. Amphiodia
urtica was most abundant, with 186 specimens. Other animals than those
shown on the chart included an anemone, three each of ceriantharians
and enteropneusts, and numerous small Pyrosoma.

The number of species and specimens totalled :

polychaetes 17* species, 116* specimens
echinoderms 6 226
mollusks 6 13
crustaceans 17 45
others 5) 9*

Total: 51* species, +09* specimens

3. Sta. 2122 (no. 259), in 48 fms. OPG took 0.95 cuft of sandy
mud with dead Lagueus and other brachiopod shells, and living animals.
The largest were a ceriantharian anemone, and a nemertean. Crustaceans
were numerous, with ostracods, amphipods, cumaceans, isopods.

4. Sta. 3613 (near no. 259), in 50 fms. OPG took 0.31 cuft of
glauconitic sand and white shelly debris, with many animals. Kelps were
absent and there was no large or conspicuous animal. Most abundant was
Nothria elegans, with 145* specimens, followed by Chaetozone setosa
with 68, and Amphiodia urtica with 50. The polychaetes were followed

334 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

in diversity and abundance by small crustaceans, especially amphipods,
ostracods and cumaceans.

5. Sta. 4045 (near no. 259), in 57 fms. OPG took about 2 liters of
sandy debris with stony hydrocoral skeletons, broken shells, a large bris-
sopsid and many animals. The largest individual was Spatangus califor-
nicus, which measured 46 mm in diameter. Most conspicuous were 4 m-
phiodia digitata, with 43 individuals, and a large Thelepus setosus. Shells
of Bittium (snail) had hydranth colonies. Crustaceans were small, in-
cluded a pagurid, cumaceans, amphipods, isopods and ostracods.

6. Sta. 3615 (near no. 259), in 60 fms. OPG took 0.63 cuft of green
shelly sand, with rock, white coralline rubble, and much diversified life.
In addition to those animals listed in the chart, the sample contained a
small amount of sponge, a solitary coral, sand covered anemones, a smooth
brown-spotted anemone, a nemertean, and a sipunculid. Crustaceans in-
cluded many amphipods, at least four kinds of isopods, 50 or more ostra-
cods, crabs with eight pagurids, ten dromiids, a spider, and another crab.
The most conspicuous animals were echinoderms: Brissopsis pacifica
(4), Lovenia californicus (2), and Amphipholis squamata (21). The
most abundant and diversified were polychaetes.

7. Sta. 2348 (near no. 260), in 75 fms. OPG took gray-green sand,
rock, coralline and shelly rubble, with few living animals. The most
abundant animal was Nothria stigmatis, with more than 100 specimens.
Most conspicuous were two brissopsids, small ophiuroids, and a holo-
thurian.

8. Sta. 3616 (near no. 260), in 88 fms. OPG took 0.18 cuft of glau-
conitic white shelly sand and rubble with trace of siliceous sponge. The
screenings retained no large animal but numerous small ones, with poly-
chaetes the most abundant and diversified. “he most conspicuous was
Chloeia pinnata (25). Amphipods were numerous and diversified, fol-
lowed by ostracods, isopods, small crabs, and a pagurid, with six indi-
viduals, in dead shells of Nassarius.

9. Sta. 3617 (near no. 260), in 93 fms. OPG took white coralline
sand and silt, the sand compacted but friable. The largest animals were
single individuals of Brissopsis pacifica and Brisaster townsendi; most
abundant were species of Tharyx, Nothria elegans, and amphipods.

Avalon Harbor
1. Sta. 3601 (near no. 239), in 18 fms. OPG took 0.69 cuft of gray

shelly sand with flocculent debris, many animals, with large tubes of
Chaetopterus, Telepsavus and Phyllochaetopterus. Coelenterates included

NO. 2 HARTMAN : QUANTITATIVE SURVEY 335

ten sea pens, more than a hundred sand covered anemones and about as
many smooth anemones. Among the polychaetes the best represented were
Owenia, Lumbrineris, spionids and Telepsavus. ‘The most conspicuous
animals were Ophiothrix spiculata and Chaetopterus variopedatus. Total
numbers of species exceeded 79. Crustaceans were represented by many
amphipods and tanaids; also some isopods, ostracods, cumaceans, a few
small shrimps and crabs, and a pinnotherid crab. Included also were a
pycnogonid (4) and an enteropneust (3). This fauna is similar to, but
more diversified than, that of the mainland shelf, in comparable depths.
A photograph (Plate 8) of the bottom in Emerald Bay in 17 fms (near
no. 207) dominated by Chaetopterus, depicts a similar area.

2. Sta. 3045 (no. 250), in 42-43 fms. Biological dredge took mud,
echinoids, ophiuroids, a holothuroid, a small squid, numerous polychaetes,
and many kinds of small crustaceans, especially amphipods and isopods.
The dredge was best suited to recover echinoderms, and least effective
for infaunal animals.

3. Sta. 2436 (no. 250), in 44 fms. OPG took 1.07 cuft of oily sandy
mud and clay. The largest animal was a large red-ribbon nemertean;
other animals included a small ceriantharian, amphipods, and many poly-
chaetes. The species are those characteristic of the shelf fauna.

4. Sta. 3603 (near no. 238), in 47 fms. OPG took 0.37 cuft of green
shelly sand. Conspicuous were brissopsids, ophiuroids and many other
animals. Animals other than those shown on the chart include coelen-
terates with a small sea pen, smooth and sand-covered burrowing anemo-
nes; a small Glottidia albida, two small sipunculids; crustaceans with 84
amphipods, 36 cumaceans, 5 anthurid isopods, 2 gnathid isopods, about
90 ostracods, 10 tanaids, and 6 pagurids in small gastropod shells. The
largest specimen was Brissopsis pacifica, and the most abundant Nothria
stigmatis with 215 specimens, followed by Amphiodia urtica with 126.
‘Total number of species is more than 80, and specimens more than 850.

5. Sta. 2639 (no. 239), in 82 fms. OPG took 1.14 cuft of gray-green
sandy mud, with white shelly sand and fine gravel. Most conspicuous
were Ophiothrix spiculata, with about 25, and Chloeia pinnata with 13
specimens. Brown ostracods were numerous, and amphipods less abun-
dant. The fauna is limited in kinds of animals and characteristic of that
at the outer edge of the shelf.

6. Sta. 2347 (no. 251), in 100 fms. OPG took 0.25 cuft of gray-
green sandy mud with broken shells, echinoid spines and many animals.
The largest was Brisaster townsendi, the most conspicuous Chloeia pin-
nata, with 40 specimens, and crustaceans were represented by many
amphipods and ostracods.

336 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

7. Sta. 3605 (near no. 238). OPG took 0.25 cuft of green mud and
sand, in 100 fms. All animals were small and inconspicuous. The most
abundant were brown ostracods, with more than 100, and Amphipholis
squamata, with 15 specimens.

8. Sta. 2367 (no. 240), in 230 fms. OPG took 0.5 cuft of gray-green
sandy mud and gravel, with dead Cadulus shells. The largest species was
Travisia pupa, the most abundant and conspicuous Chloeia pinnata, with
Sil
(2), and many ostracods.

Sta. 2344 (no. 252), in 210 fms. OPG took 0.37 cuft sandy mud,
gravel, rocks, with animals. ‘he sediments contained many Rhabdamina

and disklike foraminiferans. Ihe most conspicuous individual was a bris-
sopsid. Other animals are those characteristic of the slope.

specimens. Crustaceans were represented by amphipods (7), tanaids

Willow Cove

1. Sta. 2121 (no. 238), in 32 fms. OPG dropped twice, recovered
1.26 cuft of sand, mud, dead Laqueus shells, a large white sponge, large
asteroid, solitary corals, a surface echinoid, mollusks (not identified ),
numerous crustaceans (not identified), with small shrimps, crabs, many
amphipods and some isopods. Ihe polychaetes were represented by more
than 39 species and many specimens. The species are those characteristic
of the shelf fauna.

2. Sta. 2853 (no. 238), in 35-38 fms. Sigsbee trawl, in rocky bottom,
took rocks, sand, silt, siliceous sponge, dead brachiopod (Laqueus and
Terebratalia) shells. Coelenterates included two or more sea pens, two
small colonies of hydrocorals, a few solitary corals, several nemerteans, a
sipunculid, several polyclads, small crustaceans, especially tanaids (38),
amphipods (7), a few isopods, stalked barnacle, Scalpellum attached to
tubes of Phyllochaetopterus, and many different kinds of polychaetes.
The most abundant, 4 naitides madeirensis, had 30 specimens.

3. Sta. 2637 (no. 238), in 40 fms. Campbell grab took 1.14 cuft of
gray sandy mud with white shelly debris. The largest species was A1ol-
padia intermedia, and the most abundant 4 mphiodia digitata (93). Also
present were a ceriantharian (1), Glottidia albida (1), crustaceans rep-
resented by many amphipods and an epinebalian (2).

4. Sta. 2638 (no. 238), in 40 fms. OPG took 0.93 cuft of gray sand
and mud. The largest species were Lytechinus anamesus (urchin) and a
red nemertean. Most abundant were Amphiodia urtica (109) and cir-

ratulid polychaetes.

NO. 2 HARTMAN : QUANTITATIVE SURVEY 337

5. Sta. 2426 (no. 226), in 270 fms. OPG took 1.0 cuft of dark
green mud, with siliceous sponge, tesselated small sponge, foraminiferans,
and many animals. Largest species was Thelepus setosus with commensal,
Lepidasthenia virens ; none was outstandingly abundant. The sample con-
tained also a small, smooth anemone, crustaceans with eight amphipods,
a caprellid, five isopods, a tanaid, three cumaceans, an ostracod, and a
galatheid crab. ‘Total number of species exceeded +5, and specimens more
than 100. Species are those usually found on slopes, characterized by
siliceous sponge, the ophiuroid, Sc/erasterias heteropaes, the galatheid
crab, and polychaetes, Thelepus setosus, Onuphis vexillaria, Potamethus

mucronatus, and others.

White Cove-Long Point

‘Two samples from White Cove, taken from shore and by shallow
diving, numbered 1370 and 1378 in the Velero series (Fraser, 1943, pp.
358, 359) were useful in disclosing the presence of an unknown scalibreg-
mid, found only in the holdfasts of Eisenia arborea; the occurrence of
this species was verified by recent samples from the same locality, taken
by Mr. Robert Given. The species is described as Sclerocheilus acirratus

(see above).

1. Sta. 2152 (no. 224). 0.8 mi S of Long Pt., in 19 fms. OPG took
0.85 cuft of sandy mud with many small animals. Only the polychaetes
have been analyzed (see chart) ; these number more than 37 species and
362* specimens. The most abundant were Paraonis gracilis with 50,
Prionospio malmgreni with 42, Lumbrineris spp. with 43, Aricidea spp.
with 32, and Odontosyllis phosphorea with 31 specimens.

2. Sta. 2450 (near no. 224). Near White Cove, in 30-35 fms. Sigs-
bee trawl took a large sample from a rocky bottom with sand. Largest
individual was a sponge, Geodia mesotriaena Lendenfeld, measuring 90
mm across; also present were coelenterates, Corynactis, Stylatula and
solitary corals; bryozoans of several kinds; a polyclad, and dead Laqueus
shells. Crustaceans included amphipods, isopods, ostracods, a few epine-
balians, Scalpellum, Fupagurus in dead Bursa shells, and two large spider
crabs; also an ascidian with tough tunic. The fauna is that characteristic
of a rocky, shelly bottom. Dendropoma lituella (vermetid) was repre-
sented by numerous colonies, and Saxicava arctica burrowed in sponge
and rocks. Octopus apollyon was represented by a large male and female;
Bursa californica, dead shell, contained a large hermit crab with com-
mensal polynoid, Halosydna latior. Vhe fauna is large, diversified, asso-
ciated with a rocky, shelly bottom.

338 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

3. Sta. 2128 (no. 224). 0.3 mi off Long Pt., in 42-67 fms. Biological
dredge took sand, dead Laqueus with Vermiliopsis, alcyonarian and soli-
tary corals; echinoderms of diversified kinds, including asteroids, ophi-
uroids, echinoids, holothurians, a crinoid. Mollusks included an octopus,
key hole limpets, and various pelecypods and gastropods. Bryozoans were
encrusting and stalked. Crustaceans were diversified, with amphipods,
isopods, tanaids, cumaceans, ostracods, stalked barnacle, commensal
crabs, spider crabs, and cancroids. Polychaetes were diversified, with at
least 80 species. No one species was conspicuously abundant.

4. Sta. 2855 (no. 238). Off White Cove, in 36 fms. OPG took an
unmeasured amount of black sandy mud with reddish brown waxy
lumps. No species was outstandingly large; most abundant was Chloeia
pinnata with 70 specimens; Glottidia albida was present with 14 small
individuals. Coelenterates were represented by ceriantharians (4 small),
and a small anemone. Several small nemerteans and nematodes; crusta-
ceans with amphipods (12), isopods (2), and ostracods were also
present.

5. Sta. 2144 (no. 224+). Off Long Pt., in 45 fms. OPG took 1.6 cuft
of fine black sand in two grabs. Coelenterates were represented by small
specimens of ceriantharian (6), anemones (2), nemerteans (2), and a
long, slender sipunculid (1); crustaceans by amphipods (25), brown
ostracods (31), cumaceans (3), isopods (3), a tanaid (1), and a small
crab (2). The most abundant species were Aricidea spp. (112), Spio
punctata (33), Chaetozone setosa (27). The fauna is that characteristic
of the outer shelf.

6: Sta. 5/27 (no. 225). S of Long Pt., in 57 tms. OPG took 0237
cuft of glauconitic white shelly sand, with broken Laqueus shells, and
diversified small animals. Crustaceans included a brown ostracod (more
than a hundred), numerous ostracods and amphipods, and sparse num-
bers of cumaceans and isopods. The most conspicuous animals were
O phiothrix spiculata (more than 10) and Nothria iridescens (25) ; the
most numerous were 4ricidea spp. (more than 84).

7. Sta. 2365 (no. 210). Off Long Pt., in 300 fms. OPG took 1.57
cuft of gray-green sand and mud, with dead shells of Limopsis, Cadulus,
small moon snail, turret top shells. Largest specimen was Brisaster town-
sendi, and most conspicuous Chloeia pinnata (6). The sample contained
an echiuroid fragment, two amphipods, and a polyclad. Characteristic
species of this depth are Maldane sarsi, Limifossor diegensis, and Brisas-

ter townsend.

bo

NO. HARTMAN : QUANTITATIVE SURVEY 339

8. Sta. 2393 (near no. 211), in 250-270 fms. Trawl took kelp hold-
fasts, rubbly sand and rocky debris. he animals taken are those charac-
teristic of a rocky bottom. Echinoderms (not identified) included echi-
noids and ophiuroids. Crustaceans were represented by a few amphipods,
an isopod and a crab.

Isthmus Cove

1. Isthmus Cove, in diving depths along the rocky north wall, was
sampled Nov.-Dec. 1965 (coll. Mr. Robert Given) ; part of an encrust-
ing colony of the serpulid, Spirobranchus spinosus, yielded an association
of animals (see chart) ; the long-spined echinoid, Centrostephanus coro-
natus, occurs in rocky pockets along the wall.

2. Intertidal holdfasts of kelps, Eisenia arborea, and larger kelps
(coll. Mr. Robert Given) yielded characteristic animals (see chart)
with grapsid and pagurid crabs.

3. Sta. 2452 (no. 207) near Isthmus, in 28 fms. OPG took 0.37 cuft
of compact sandy clay with many animals and much calcareous debris.
In addition to the species named in the chart, a burrowing anemone (2),
two nemerteans (2), Phoronopsis californica, red in life (1), Glottidia
albida (1), many amphipods and other small crustaceans, several ascidi-
ans and an enteropneust (1) were present. The largest species was
Lanice conchilega; none was conspicuously abundant. Total number of
species was estimated at more than 60, with polychaetes about 30, echi-
noderms at least 2, mollusks 15, crustaceans many, and others, 8 or more.

A concentration of Laqueus californicus is believed to exist along the
rocky exposures between Empire Landing and Ship Rock, in 30 to 70
fms, indicated by concentrations of dead shells in these areas. Several
probes (4a-4+f) were attempted to procure living lamp-shells; the fol-
lowing stations indicated their presence, with associated organisms:

3. Sta. 2955 (near no. 186). SW of Ship Rock, in 34 fms. The
dredge took sand, rock, shells of Laqueus, Terebratalia, Lytechinus and
other echinoderms, together with Conus californicus; the echinoderms

included :
A mphiodia urtica O phiopsila californica
Amphipholis pugetana O phiopteris papillosa
Luidia asthenosoma O phiothrix spiculata
Luidia foliolata Sclerasterias heteropaes

Ophiopholis bakeri

340 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

4a. Sta. 2960 (near no. 186). SW of Ship Rock, in 34-36 fms.
Dredge took shelly sand with Laqueus, Terebratalia and other animals,
with the following echinoderms:

Amphipholis pugetana O phiopteris papillosa
Astropecten californicus O phiothrix spiculata
Henricia sp. O phiura lutkeni
Mediaster aequalis Sclerasterias heteropaes

O phiopholis bakeri

+b. Sta. 2959 (near no. 186). NW off Ship Rock, in 42 fms. Dredge

took-sand with Lytechinus, small pelecypods and other echinoderms,

including:
A mphiodia digitata Luidtia ludwigt
Amphioplus hexacanthus Lytechinus anamesus
Amphiura arcystata O phiopholis bakeri
Florometra perplexa O phiura lutkeni

4c. Sta. 3310 (near no. 186). NW off Ship Rock, in 47-52 fms.
Dredge took sand with a sea pen, few brachiopods and mollusks, several
crabs, and the following echinoderms :

Allocentrotus fragilis O phiura lutkenti
A stropecten californicus

4d. Sta. 2952 (near no. 208). SE off Ship Rock, in 63 fms. OPG

took sand with some living Laqueus, Florometra perplexa and gastropods.

te. Sta. 2957 (near no. 186). NW off Ship Rock, in 64 fms. OPG
took Laqueus with Allocentrotus fragilis, Spatangus californicus and

Florometra perplexa.

4f. Sta. 2953 (near no. 208). SE off Ship Rock, in 52-65 fms.
Dredge took shelly sand, Laqueus, heart-urchins, gastropods, with other
animals,

5. Sta. 2451 (no. 208). Between Isthmus and Long Pt., in 111 fms.
OPG took 1.57 cuft of compact sandy clay with many animals. In addi-
tion to those named in the chart, the sample contained amphipods (20),
ostracods (8), an anthurid isopod (3), a tanaid (1), a pagurid in
Amphissa shell (1). The largest was Molpadia, and the most abundant
were Alaldane sarsi (20), and Tellina carpenteri (16).

NO. 2 HARTMAN : QUANTITATIVE SURVEY 341

Numbers of species and specimens totalled :

polychaetes 21 species, 60 specimens
echinoderms + 20
mollusks 8 39
crustaceans De 33

6. Sta. 2733 (near no. 187). NE of Ship Rock, in 152 fms. OPG
took 1.13 cuft of sandy mud with many animals. In addition to those
named in the chart, the sample included a ceriantharian (2), an echiu-
roid (1), a nemertean (1), amphipods (several), and a gnathid isopod
(1). The largest animal was the echiuroid, the most abundant 4 mphi-
odia digitata (48) and solenogasters (23).

Numbers of species and specimens estimated :

polychaetes 22 species, 40*specimens
echinoderms j| ie
mollusks 6* Vas
crustaceans Ob, Mee

Total: 37*species, 144+ specimens

7. Sta. 2302 (no. 187). Between Isthmus and Ship Rock, in 185
fms. OPG took 1.57 cuft of fine greenish mud with many animals. In
addition to those named in the chart, the sample contained a ceriantha-
rian (2), nemerteans (2+), an echiuroid (2), numerous amphipods and
a few isopods. The largest individual was a brissopsid echinoid, and the
most abundant were ophiuroids. Total numbers were estimated at more
than 29 species and 69 specimens.

8. Sta. 2301 (no. 189), in 335 fms. Campbell grab took 3.33 cuft of
fine grayish green mud with foraminiferans. In addition to those named
in the chart, there were an anemone (2), an echiuroid (1), a sipunculid
(3), nemerteans (2+) and an enteropneust (1). The largest animal was
an echinoid, and none was outstandingly abundant. Numbers were esti-
mated at 21 species and about 60 specimens.

Howlands Landing
1. Sta. 2142 (no. 186), in 19 fms. OPG took 1.9 cuft of sand and

mud with many animals. In addition to those named in the chart, there
were several kinds of sponges, some large and spherical, a ceriantharian
(1), small sea whips (3), a burrowing anemone covered with coarse
sand (4), four kinds of nemerteans (7), sipunculids of more than one

342 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

kind (12), a reddish purple echiuroid (1), Glottidia albida (1), Phoro-
nopsis californica (7), bryozoans of several kinds, an amphipod (1),
isopods (2+), two shrimps (5), a crab (1), two enteropneusts (6) and
an ascidian (7). The largest animal was a spherical sponge, the most
conspicuous Chaetopterus variopedatus, and the most abundant Owenia
f. collaris (60). Total numbers were estimated at more than 60 species,
and more than 200 specimens.

2. Sta. 2961 (near no. 186). SW of Ship Rock, in 36 fms. OPG
took shelly sand and many animals. In addition to those named in the
chart, the sample contained two sponges (2), four coelenterates (20), a
nemertean (4), two sipunculids (7), a leech (1), a bryozoan colony
(1), a phoronid (6), Laqueus californicus (3), Terebratalia occidentalis
(2), several kinds of amphipods (18), ostracods (31), isopods (5), a
cumacean (1), Mitella polymerus (22), a decapod (2), and an enterop-
neust (2).

Numbers of species and specimens totalled :

polychaetes 56 species, 102 specimens
echinoderms 7 69
mollusks 23 105
crustaceans 8+ 81
others 7; 52
Total: 111*species, about +09 specimens

3. Sta. 3569 (near no. 162). N of Howlands Landing, in 100 fms.
OPG took 0.69 cuft of green, sandy mud with many animals. The lot
contained a large sea-whip, Cerebratulus rubra, many polychaetes char-
acteristic of muds in slope depths, small crustaceans, and the following
echinoderms:

ie)

bo
CSP hort

A mphiacantha amphacantha
Amphiodia digitata

—"

Amphioplus strongyloplax
Amphipholis squamata
Amphiura arcystata

nN

Brissopsis pacifica
Molpadia sp. 1, measuring 75 mm long

4. Sta. 2738 (no. 136). Off Howlands Landing, in 342 fms. Camp-
bell grab took 3.15 cuft of green sticky mud with oily odor and waxy
nodules. In addition to those named on the chart, there was a trace of
siliceous sponge, a sipunculid (2), a nemertean (3), and an enteropneust
(2). Echinoderms were represented by a small ophiuroid.

No. 2 HARTMAN : QUANTITATIVE SURVEY 343

Numbers of species and specimens totalled :

polychaetes 10 species, 31 specimens
ophiuroid 1 1
mollusks 2 6
others 3 6
otal: 16 species, +4 specimens
West End

1. Sta. 3570 (no. 162), in 48 fms. OPG took 0.56 cuft of sandy
mud with shelly debris and considerable life ; dead remains included those
of Laqueus, Terebratalia, solitary corals, Protula tubes and coralline
algae. The most conspicuous living animals are listed in the chart.

2. Sta. 5148 (near no. 162), in 98 fms. OPG took 0.31 cuft of glau-
conitic sand with foraminiferans and large oil globules. Ophiuroids were
most conspicuous. In addition to the species named in the chart, there
were six amphipods (19), two cumaceans (3) and three ostracods (37).
The largest animals were ophiuroids, and the most abundant Prionospio
malmegreni (35), Amphiodia digitata (29), Nothria stigmatis (30), and
Amphiacantha amphacantha (24). Total numbers were estimated at 45
species and 241 specimens.

Seta. 2/56 (no. 160) un 132 ims. \OPG took 1.5/7 cuit of sandy
green mud with reddish brown waxy lumps and many dead shells of
Cardita, Cuspidaria, Cadulus, Nassarius and Thyasira. In addition to the
species named in the chart, there were a few amphipods and isopods. The
largest animal was Brissopsis pacifica, and the most abundant were
Amphiodia digitata (64) and Amphioplus strongyloplax (22). Total
numbers were estimated at 32 species and 140 specimens.

4. Sta. 2389 (no. 161), in 136 fms. OPG took 2.2 cuft of gray sand
and mud with many animals. In addition to those listed in the chart,
there were a burrowing anemone (1), a polyclad (1), a sipunculid (1),
a large echiuroid (1), and a few small crustaceans. Largest species were
Brissopsis pacifica and the echiuroid, and the most abundant were the
ophiuroids. Total numbers were estimated at more than +5 species and
175 specimens.

Sub 4. Sta. 5146 (near no. 160), in 140 fms. Grab took a large
basaltic rock, 14” by 7.5’, with attached large sponge (Heathia), soli-
tary coral, alcyonarians (2), tunicates, and bored by pholad mollusks ;
ophiuroids occupied the crevices, and serpulid polychaetes were attached

344 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

on the surfaces. The ophiuroids and mollusks are named in the chart.
Most abundant species was O phipholis bakeri (30).

5. Sta. 2737 (no. 135), in 256 fms. OPG took 2.77 cuft of sticky
green mud with odor of oil, foraminiferans, and numerous animals.
Glassy sponge spicules characterized the mud. In addition to the species
named in the chart, the sample contained a large nemertean (2), an
echiuroid (2), and an amphipod (1). Largest individuals were the
echiuroid and nemertean. Total numbers were estimated at 29 species
and 72 specimens.

6. Sta. 2798 (no. 89), in 386 fms. OPG took 2.96 cuft of bluish
green-gray mud, with siliceous sponge spicules, radiolarian shells, fora-
miniferans, otoliths, and deep-water animals. In addition to those named
in the chart, there were a nemertean (2), a ghost shrimp (1), and an
enteropneust (1). Total numbers were estimated at 20 species and about
35 specimens.

345

HARTMAN : QUANTITATIVE SURVEY

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ALLAN HANCOCK PACIFIC EXPEDITIONS

354

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

356

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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

358

ay eg a ES AE pbs 2 ee eR Sf ye ERE et AS eee eee i nodoroad
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VOL. 19

ALLAN HANCOCK PACIFIC EXPEDITIONS

360

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HARTMAN : QUANTITATIVE SURVEY 361

NO. 2

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os * © * + + syouraqrpnu

362 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta from Santa Catalina Island,
showing distributions by profiles,

VOL. 19

from Isthmus Cove, Howlands Landing, and West End.
+ indicates presence, - absence.
Species represented by more than 10 specimens are indicated by *.

Dodecaceria concharum

Pherusa inflata. .

Polycirrus californicus

Poliydora (Spe sts

Spirobranchus spinosus

Typosyllis sp. .. .

Vermiliopsis infundibulum

BUMS Epo 6 o oo oc
Exogone uniformis . .
Fabricia pacifica. .
Harmothoe hirsuta. .
Lepidonotus caelorus

Lumbrineris latreilli

Nereis mediator...

Phragmatopoma californica

phyllodocid, unknown

Polycimrus Sp. . « .

Polydora ligni . .

.

Polyophthalmus pictus .

Salmacina dysteri .

Selerocheilus acirrata

Sphaerosyllis sp. . .
Splonwdsiy.

SpLrorbildS mae wesiemaco
Typosyllis aciculata
Aglaophamus dicirris

Amaeana occidentalis

amphanetudse 0k sie fe

Isthmus Cove

2

sie UN ey Wee 7/4
as e- = + — =
foo obese
fo See =
os Se Se

Howl.

i

2

Land.

3

4

West End
aed aah By fe)

NO. 2

HARTMAN : QUANTITATIVE

Polychaeta from Santa Catalina Isl

AmLcildea Spp. . . . «

Capmterird= 3... «

Chaetopterus variopedatus

cirratulid ..

Diopatra ornata .

Eumida, trilineate .

Glycera spp. .

Lanice conchilega. .
Lepidasthenia spp. .
Lumbrineris spp. .
Malidanvasi <9. < «
Megalomma sp. ... .
Panthalis pacifica .
Pista cf. cristata
Pista elongata ..
Prionospio pinnata .
Rhodine bitorguata
Spiophanes sp. ...
Sternaspis fossor. .

Sthenelanella uniformis

Streblosoma crassibranchia

SVAEUdSieere 6 se

Telepsavus costarum .

Terebellides stroemii

Thalenessa spinosa

Thelepus setosus . .

Aedicira ramosa .

Amphicteis scaphobranchiata

.

.

Anaitides medipapillata .

Brada pilosa ..

Chloeia pinnata...

1

2.

3
+

+

Isthmus Cove

Eo 6 71.8

- -+t+ - +

SURVEY

Howl.

a

2
+

+

Land.

3

and (continued)

4

+

363

364 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta from Santa Catalina Island (continued)

Gossumal candidate n-ne
Drilonereis nuda ....

Glycera Caplitaita |. = -

Goniada brunnea .... -
Haploscoloplos panamensis
Harmothoe spp. ..--. . -
Laonice cirrata .... .
Lumbrineris cruzensis .

Mailidanel Savsay 2 jo) =) -
Nerevsmspe. 3 «+ 5
Notomastus Sp. .-.-. + -
Pholoe glabra ...--s -
Praxillella a. pacifica .
Sphaerodoridium minutum .
Tharyx monilaris ... .

Anaitides spp. .... -

Bradamedabmay rye eel

Haploscoloplos elongatus

Nohstentet Sg o 6 6 8 Gd oc

Nephtys cornuta. . -

Nephitys: Spps = «= « . « =

Onuphis parva ....-. -

Pectinaria californiensis
Phyllochaetopterus sp. .
jELdkeveyeniel Go 6 Oo o 9 6 O
jpolvanontl “Glog mG o o oO 6
Tharyx tesselata ... -
ihe Walsall lg lo a om oO ©
Aglaophamus sp. . .
Ammotrypane aulogaster
Ancistrosyllis breviceps

Isthmus Cove

2

3

m1

5
+

+

6

+

7

+

Howl.

1

22

Land.

3

4

VOL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta from

Aricidea lopezi...
Amphicteis sp. ...
Brada pluribranchiata .
Chonegsp. 7 = = 2 - « -
Harmothoe scriptoria .
Laonice foliata...
Melinna denticulata .
Nothria iridescens ..

Paraonis g. oculata. .

Santa Catalina Island (continued)

Isthmus Cove

L234

Phyllochaetopterus limicolus ... .

Prionospio cirrifera .
Spiophanes fimbriata .
Travisia pupa... .

Anobothrus gracilis . .
Caukifmagcaliida cn. 5 -
?Leiochrides sp. ...
Lysippe annectens .. .
Myriochele gracilis . .

Potamethus mucronatus .

Protula superba... .
Chaetozone setosa...

Cimmitonmia Sp. . . « -
Eulalia myriacyclum .

MIM aS pPlern e) Glen) clay te
Notomastus latericeus .

Owenia f. collaris ..

Paraonis gracilis ...
Pherusa capulata . .
Pherusa neopapillata

Phyllochaetopterus prolifica....

Prionospio malmgreni

5

6

7

+

+

Howl.

al

2

365

366 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta from Santa Catalina Island (continued)

PSiammMolvice ssp ct itn eens

Pseudopotamilla sp. . .

Scalibregma inflatum ..

Sthenelais tertiaglabra .

tenebedsltdss -) (53 came es 0.

Mees Sehia oo oO oF 6 5 4
Inge iloenalilstss Silo o 6 oo
Vermminlilopsasm sip) us) « i
Dorvillea articulata ..
EW alatayesippe 0 sess cee ye

Gyptiswdi glabra 2 . -

Nereis ?procera ....

Redsiidic easpenciee iis) \-
Rasitawdaisauncitaus ean).

RUaCOSsiteeUsSmsipiem ts)
Rhamphobrachium longisetosum
SeNoiglael AS GG cee eo Gmc

Suis cullonaicd sissy ene money jen
WELT BiG 6 Go oo a 6
Halen ile 5 6 of oe 6

Vermiliopsis biformis .
ANAS EM ZaNOPSH Ns) in le
Aricidea neosuecica .
Chaetozone corona. .
Naineris uncinata ...
Nothria stigmatis . .
?Mesochaetopterus sp.
Drilonereds! spi ais |s
Eunice aphroditois ..
Ancistrosyllis tentaculata

Cirratulis, unknown sp.

1

Isthmus Cove

2

3

4

5

6 7 8

Howl.

1
+

+

2

Land.

3

u

VOL. 19

West End
12 oe ee See

sc o- — — =

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Polychaeta from Santa Catalina Island (continued)

Cirrophorus aciculatus .
Leiochrides hemipodus . .

Pherusa, unknown sp. ..
Aricidea uschakowi ...
capitellid, unknown...

Glycera branchiopoda . .

US OLGANS Peis) eo) es

Isthmus Cove
Ne?!

3) 495" 6) 7) 8

Howl.

1

2

Land.

3

Echinodermata from Santa Catalina Island
showing distributions by profiles,
from Isthmus Cove, Howlands Landing, and West End.

Centrostephanus coronatus

Lytechinus anamesus .. .
Ophiothrix spiculata ..

noLothurords = « «. «. « «
Allocentrotus fragilis .
Amphiodia digitata ...
Amphiodia urtica ....
Amphioplus hexacanthus .
Amphipholis pugetana ..
Amphiura arcystata ...

Astropecten californicus
Florometra perplexa.. .
Henmiiciiayispe <6 « « «
Luidia asthenosoma ...
Luvidita foldiolata . . . «
Luidia ludwigi .... .
Mediaster aequalis ...
Ophiopholis bakeri ...
Ophiopsila californica

Ophiopteris papillosa .
Ophiura lutkeni .....

+

+

+

+

m1

367

Le 72) 3) Sao

a

368 ALLAN HANCOCK PACIFIC EXPEDITIONS

Echinodermata from Santa Catalina Island (continued)

Isthmus Cove Howl. Land.

As & Hb 7 8 2 See ale

Sclerasterias heteropaes oe feet == = = fo =
Spatangus californicus . she ss os =
Amphipholis squamata . . a, oe ee sage =
Molpadia intermedia .. . eee So PS = = sage =
Amphioplus strongyloplax Se Re stgiet he so tS +
Brisaster townsendi .. . eee see et ---- =
Brissopsis pacifica... fe fees eee e tec sabe =
Opiates! 5 6 6 6 6:0 6 SO seat oe Choo = ob =
Amphiacantha amphacantha A ae ie a, ae oo AC Bee th *
Leptosynapta albicans . . Eis, coratromee Mrrerar sce ee et -
Parastichopus californicus Baie Oh be tr SOE? VC == -
Pentamera pseudopopulifera So 6 0 6 Oo 0 5 3 t=. = -
Woiy=iGlieh SG 6 G oF oO oO Co 5 Oo Oo oO oO 0 SC «1%? i= -
Ophionereis eurybrachiplax oo 8 Oo Oo Oo 8 ene -
Ophiacanthia diplasia. . 6 Oo Of 6 Oo oO 6 4 oO oS :

Mollusca from Santa Catalina Island,
showing distributions by profiles,

from Isthmus Cove, Howlands Landing, and West End.

Acteon punctocoelata .. Geo See So ---- =
Amygdalum pallidulum .. ee RSS = sO Ss = =
Axinopsida serricata .. oo foe SS = ---- =
Cylichna diegensis ... a are pen s
Elaeocyma halocydne.. . oe Vou abe ee cs ---- =
Euiima Ccalatornaca i 2% joa es) Sy ye = Ss ae =
Lyonsia californica... oe) fa) === aSes -
Nemocardium centrofilosum ee fee ses Site =
Nucullay Damien icy Ae Qe tees ee 25 Sie =
Nuculana taphria .... == = = S fees = =
Parvilucina tenuisculpta Cs Eee =
SaxalCaVide aMGib Cay ye tel tien Seek =) ===) = =i Ss =

Sinum Scopulliosum <1. 2 . Bitte See ea eS ep ae ao

VOL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Mollusca from Santa Catalina Island (continued)

Solamen columbianum .

Tellina carpenteri .
Cardiomya pectinata .
Cardita ventricosa .
Cuspidaria apodema .
solenogasters ...
Thyasira barbarensis
Cadulus tolmiei ...
Cuspidaria sp. ..
Lima dehiscens .. .
gastropods ....
pelecypods ...
Acteocina intermedia

Aglaja sp. ....-.

Balcis catalinensis .

Clinocardium nuttalli

Conus californicus .

Cyrilla munita

Kellia suborbicularis

Lima subauriculata .

Micranellum crebricinctum

or M. sp.
Nuculana hamata 5

Pandora bilirata ..
Pseudochama exogyra .
Sphenia fragilis ..
Turbonilla sp. ...

Verticordia ornata .

Volvulella tenuissima .

Dentalzum Sp... « -

Epitonium tinctum or E.

Tellina modesta .

sp.

Isthmus Cove

ib ee) TS

. wos SS SS
Se haan seh
ep chy Cami gum es
ao 7
ee oe te
io oe age
. 6 Oo hee
. eee e Gt
=) aeryee, Os
+.

7

Land.

3

m

369

West End
a ederel TS 5)
SSS ee se
SS SS ae
See Soe
S57 SoS co Ss
Si ee SS
ay
poe ae
eo

370 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Mollusea from Santa Catalina Island (continued)

Isthmus Cove Howl. Land. West End

2s: YS 6s 7s 2. Si 4 23 GS
Saxicava ef. rugosa... Big "SOF GO Si Som.c yo 6. 6 Ba on. ee =
Anphusisa Drecollom 7) = =< yao Bo ao 4 =) fel etot eke oy ee) Wares t:
Dacrydium pacificum. . . Shoo, coo 62 5 Se he ce iceso ote
Protochaetoderma californica ........ yey ome SG Go 6

Nitidella permodesta .. Sates. peek ecu ete a. iS wen es away racers

No. 2 HARTMAN : QUANTITATIVE SURVEY 371

SANTA CATALINA ISLAND, WINDWARD SIDE

The grab was lowered along transect lines extending oceanward,
from the outer side of West End, Ribbon Rock (see Map 2), Catalina
harbor, Farnsworth bank, China Point to Salta Verde point, and Pali-
sades. Analyses disclosed changes in faunal composition by place and
depth. Many species were found which are unknown on the leeward
side of the island. The finding of a conspicuous bryozoan fauna on
Farnsworth bank, and a large population of the lingulate brachiopod at
Palisades were noteworthy.

West End

1. Sta. 3577, in 15 fms. A snapper sampler took black debris with
large tubes of Chaetopterus variopedatus and many associated animals,

resembling those at White Cove, SCI, in comparable depths.

2. Sta. 3576, in 36 fms. OPG took 0.37 cuft of shelly sand with
broken shells, chiefly A/ytilus and other shelf-mollusks, also bryozoan
clusters, fragmented tubes of Telepsavus costarum and Phyllochaetop-
terus prolifica. In addition to the species named in the chart, there were
a small ceriantharian (5), an anemone covered with coarse, black sand
(12), a long ribbonlike polyclad (1), a nemertean (3), a slender sipun-
culid (12), a phoronid in coarse, sand-covered tube (12), amphipods of
several kinds (many), an anthurid isopod (several), a tanaid (7), an
ostracod (7), a cumacean (2), a dromiid crab (2), and Glottidia albida
(5). The largest species was Glycera americana; none was conspicuously
abundant. Total numbers were estimated at more than 80 species and
575 specimens.

3. Sta. 3575, in 44 fms. OPG took 0.25 cuft of green shelly sand
with rock chips. The sample contained many small to very small animals,
like those on the leeward side of the island. Rocks had attached V’ermili-
opsis and tubes of Pista. Polychaetes, echinoderms, mollusks and small

crustaceans were diversified and represented by immature individuals.

4. Sta. 3573, in 88 fms. OPG took 0.31 cuft of glauconitic white
sand, with echinoid ( ?Lytechinus) spines, foraminiferans and many
small animals, like those of the leeward side of the island. None was
large; most abundant species were Nothria stigmatis (140), Chloeia pin-
nata (52) and Amphiodia digitata (45).

Ry) ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens were estimated at:

polychaetes 35 species, 302 specimens
echinoderms + Sy
mollusks 2 few
crustaceans 5 24
others 1 1

Total: 47+ species, 378+ specimens

5. Sta. 3572, in 103 fms. OPG took 0.25 cuft of glauconitic white
sand with shelly rubble, siliceous sponge spicules and many small ani-
mals; none was conspicuous or large. Most numerous and abundant were
polychaetes, chiefly onuphids, cirratulids and spionids. Most abundant
were Nothria stigmatis (56), Amphiodia urtica (43), Chloeia pinnata
(23) and Haploscoloplos elongatus (10). Polychaetes numbered 26
species; none was unique to the area. Echinoderms were represented by
two ophiuroids (49), and mollusks by four or more species, each with
few specimens. Small crustaceans were represented by amphipods, iso-
pods, tanaids and cumaceans.

Ribbon Rock

1. Sta. 3580, in 40 fms. OPG took 1.26 cuft of grayish green, sandy
mud. Most conspicuous were a large Luidia foliolata, purple holothu-
rians, Ophiothrix spiculata and other ophiuroids. The sample contained
also a small sea pen, an anemone, polychaetes with about 35 species, mol-
lusks with Cadulus (28); and small pelecypods and gastropods; small
crustaceans were represented by many amphipods, cumaceans (26), an
isopod (1), a small cancroid crab (2), and a pinnotherid crab (2).

2. Sta. 3579, in 48 fms. OPG took 1.76 cuft of grayish green, sandy
mud with shelly rubble and many animals. Largest was Cerebratulus
rubra, and most conspicuous were Amphipholis squamata (82), Amphi-
oplus strongyloplax (7), Amphiacantha amphacantha (4), Leptosynapta
albicans (16) and Pentamera pseudopopulifera (3). In addition to the
species named in the chart, crustaceans were present as amphipods, 1so-
pods, cumaceans and tanaids, each with few individuals; other animals
were small anemones (2), and a large Cerebratulus rubra.

Numbers of species and specimens totalled :

polychaetes 42 species, ca. 400 specimens
echinoderms 5) 12
mollusks 8 18
crustaceans at 10+
others 4 DA

Total: 64+ species, 561+ specimens

NO. 2 HARTMAN !: QUANTITATIVE SURVEY 33

Catalina harbor

Catalina harbor was sampled in 4 to 100 fms. Shallower depths, in
0.5 to 10 fms, had been previously reported (Reish, 1964) when 43
species of polychaetes were named, identical with those of the mainland
shelf, in comparable depths.

1. Sta. 3586, in + fms. A snapper took about a liter of gray mud with
many small animals and a large nemertean, Cerebratulus lineolatus.
Most abundant were Lumbrineris limicola and L. minima (together
260), Prionospio malmgreni (177), Paraonis gracilis (40), Haploscolop-
los elongatus (35), and Tharyx monilaris (23+). Small crustaceans
were present as amphipods, tanaids and cumaceans. Echinoderms were

absent.

Numbers of species and specimens totalled:
polychaetes 29 species, 600* specimens
mollusks 55 10
crustaceans oe 50+
others + i

Total: 43*species, 677* specimens

2. Sta. 3585, in 23 fms. OPG took 1.89 cuft of gray mud with much
shelly and mucoid debris and many small animals. In addition to those
named in the chart, there were a sand-covered and a smooth anemone
(2), a nemertean in mucoid tube (4), Golfingia sp. (7), an echiuroid
(1), Dendrostoma sp. (2), a phoronid (1) and Glottidia albida (14).
The most conspicuous were ophiuroids. Most numerous were Tharyx
multifilis (56), Amphiodia urtica (38), Nephtys ferruginea (24),
O phiopholis longispina (21) and Amphipholis pugetana (20). Crusta-
ceans were present as amphipods (many) isopods, cumaceans, tanaids.

Numbers of species and specimens totalled :

polychaetes 52 species, 289 specimens
echinoderms 11 106
mollusks i 10
crustaceans 10+ many
others 7 39

Total: 87* species, +44* specimens

3. Sta. 3584, in 48 fms. OPG took 2.0 cuft of gray mud with shell
fragments, mucoid debris, and many animals. In addition to those listed
in the chart, there was a loose-pannicled sea pen (1), a nemertean (2),
amphipods (7), anthurid and gnathid isopods (2), a cumacean (1) and
an enteropneust (2). Largest species were the sea pen and Pista dis-
juncta, and most conspicuous was 4A mphiura arcystata. Most abundant
were 4 mphiodia urtica (97) and Amphiura arcystata (14).

374 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Numbers of species and specimens totalled :

polychaetes 34 species, 77 specimens
echinoderms 7 110
mollusks 5 5
crustaceans 4+ re
others 3 11

Total: 53* species, 214 specimens

4. Sta. 3581, in 100 fms. OPG took 1.76 cuft of grayish green, sandy
mud with shell fragments, mucoid debris, and many animals. In addition
to those named in the chart, there was a nemertean (1), a sipunculid
(1), amphipods (12), an anthurid isopod (1), and an ostracod (1).
The largest species was Nephtys glabra (weight 53 grams), and the
most numerous was 4 mphioplus strongyloplax (25).

Numbers of species and specimens totalled :

polychaetes 38 species, 104 specimens
mollusks 6 6
echinoderms 6 33
crustaceans Sh 14
others Z 2

Total: 55+ species, 159 specimens

Farnsworth Bank
1. Sta. 3594, in 16 fms. OPG took a fraction of a rocky shoal, with

hydrocorals, bryozoans, red algae and many associated animals. Note-
worthy were ophiuroids, snapping shrimps, many amphipods associated
with algae, many branches of salmon-pink, fenestrated bryozoans, Phi-
dolophora pacifica, purple hydrocoral, Allopora californica with com-
mensal gastropod, Pedicularia californica, and many other animals. In
addition to those named in the chart, there were present also a nemer-
tean (1), a sipunculid (1), a polyclad (1), a caprellid (1), isopods
(10+), tanaids (2), an epinebalian (3), small crabs (2+), and a pycno-
gonid (3). Some of the bryozoans are named elsewhere (see list of bryo-
zoans, identified by Mr. William Banta). The total numbers of species
and specimens are not estimated, but are expected to be much higher than
that in adjacent soft-bottoms, because of the larger numbers of epifaunal
animals.

2. Sta. 3595, in 16 fms. OPG took 0.47 cuft of coralline nodules,
with green Ulva, many bryozoan clusters, broken shells of large Hin-
nites, Astarte, Astrea, slipper limpets, many large fragments of dead
hydrocorals, and dead shells penetrated by boring sponge. Animals were

NO. 2 HARTMAN : QUANTITATIVE SURVEY 375

very numerous and diversified. In addition to those listed in the chart,
there were a ceriantharian (1), a polyclad (3), small nemerteans (2),
sipunculids penetrating coral stems (6*), an oligochaete (6), many kinds
of bryozoans, differing from those in the preceding sample, amphipods
(many), isopods, especially anthurids (many), tanaids, pagurid crabs,
shrimps, a pycnogonid (3), and a small enteropneust (1). The most con-
spicuous species were Ophiothrix spiculata (30), Amphipholis squamata
(38), and a large Asterina miniata.

Numbers of species and specimens, excluding bryozoans (see list of
species elsewhere ), totalled:

polychaetes 62 species, 459 specimens
echinoderms 8 105
mollusks Is 20*
crustaceans Si 50*
others (excl. 7 17
bryozoans )
Total: 89* species, 651+specimens

(excluding bryozoans)

Sta. 3597, in 35 fms. OPG took 1.44 cuft of fine gray sandy mud
with many small animals. The most conspicuous and abundant species
was Amphiodia urtica (177), and other species were like those of the
mainland shelf in comparable depths and sediments.

Sta. 3593, in 52 fms. OPG took 0.5 cuft of glauconitic sand and
much biological rubble. Animals were like those of the shelf, but in
sparser numbers. Characteristic echinoderms were 4 mphiodia occidentalis
and Amphipholis squamata, and polychaetes were Pherusa capulata and
Chloeia pinnata, with many other smaller kinds.

Sta. 3592, in 57 fms. OPG took 0.44 cuft of shelly, glauconitic sand,
calcareous algae and gravelly rubble. There were no large, conspicuous
animals; all were small, diversified, best represented by polychaetes and
crustaceans, like those of the mainland shelf in comparable depths.

Sta. 3591, in 88 fms. OPG took 0.14 cuft of glauconitic sand with
many kinds of small animals, most in sparse numbers, and Rhabdamina
foraminiferans. Characteristic polychaetes were Nothria stigmatis, Pista
cf. cristata, small maldanids, cirratulids, and other diversified kinds.
Echinoderms were represented by Amphipholis squamata, Amphiodia
urtica and Astropecten californica, and mollusks by Amygdalum pallidu-
lum, Cadulus, Tellina, and others. Crustaceans were present as many
amphipods, a few isopods, tanaids, many ostracods, some cumaceans and
a small shrimp.

376 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Between China Point and Salta Verde Point
1. Sta. 3600, in 25 fms. OPG took 0.25 cuft of sandy mud with

many animals. In addition to those named in the chart, there was a
ceriantharian (1), small anemones (2), a hydroid branch (2), a nemer-
tean (2), a branching bryozoan, Glottidia albida (13), amphipods (20),
a gnathid isopod (1), a cumacean (3), a small crab (1), a pinnotherid
crab (1), a pycnogonid (2), and a sand-covered ascidian (2). Most con-
spicuous and abundant species were Glottidia albida (13), tubes of
Streblosoma crassibranchia (many) and Sthenelanella uniformis (10*).

Numbers of species and specimens totalled :

polychaetes 41 species, 146 specimens
echinoderms 6 19
mollusks 10 32
crustaceans a 26
others f| 24

Total: 69* species, 247 specimens

2. Sta. 2176, in 28 fms. OPG took 1.32 cuft of nodular muddy sand
and mud. Branching bryozoans and simple ascidians resembling small
cindery balls, characterized the screenings. In addition to the species
named in the chart, there were an anemone (7), a nemertean (2*), a
sipunculid (1), Glottidia albida (4), an ascidian (3), and small crusta-
ceans, especially amphipods.

Numbers of species and specimens totalled :

polychaetes 54 species, 300* specimens
echinoderms I Das
mollusks 4+ 8*
crustaceans 10* many
others 6 (ae

Total: 75* species, 322+ specimens

Sta. 2175, in 58 fms. OPG took a small sample of mud and shell
fragments, with many diversified animals. The polychaetes which were
best represented included some not recovered in other samples, such as
A phrodita refulgida, Magelona with serrated prostomial margin, uniden-
tified cirratulids, maldanid with parasitic copepod, ampharetids of sev-
eral kinds, Megalomma sp., and spirorbids. The fauna may have affinities
with that of the outer shelf.

Sta. 2174, in 161 fms. OPG took 0.95 cuft of mud with shell frag-
ments. Ihe largest animal was a large seastar, [ippasteria sp., accom-

NO. 2 HARTMAN : QUANTITATIVE SURVEY 377

panied by sea urchins (5), many small mollusks, numerous small crus-
taceans, and many diversified polychaetes, some not recovered elsewhere.

Sta. 2173, in 252 fms. OPG took 1.57 cuft of mud, with a nemer-
tean, brissopsid urchins (2), a few ophiuroids, a small crustacean, and
polychaetes including some not found elsewhere; they include 4 naitides
sp.; Glycinde polygnatha; Lepidasthenia virens; Lumbrineris index ;
Oncoscolex pacificus ; Prionospio sp.; Scalibregma inflatum ; Thelepus sp.

Sta. 2172, in 517 fms. OPG took 2.52 cuft of fine green mud with
glass sponge, disklike foraminiferans, and animals differing from those in
shallower depths. The largest individuals were a philinid mollusk (1),
and a tube of MWaldane, with lateral branches, measuring 30 mm long
by 5 mm wide. Other animals were a pelecypod (3), an amphipod (1),
an ovigerous isopod (1), and polychaetes: A4ricidea sp., Chaetozone sp.,
Drilonereis sp., Laonice sp., Lumbrineris index (1), Maldane sp. with
branched tube (4), Ninoe sp. and Tharyx ?multifilis (3).

Sta. 2170, in 587 fms. OPG took 2.9 cuft of fine mud with glass
sponge spicules, many foraminiferans, and few animals. They were poly-
chaetes with Aricidea uschakowi (several), Onuphis sp. (1), Lumbri-
neris 2?index (1), a harmothoid (1), a terebellid (1) ; a small pelecypod
and amphipods (few).

Palisades

1. Sta. 3610, in 20.5 fms. OPG took 0.5 cuft of gray muddy sand
with black algal detritus, little sand and gravel, and many animals. Most
abundant and conspicuous was Glottidia albida (50 large, measuring to
27 mm long, and + small, only 9 mm long). In addition to the species
named in the chart, there were a small ceriantharian (3), a large Cere-
bratulus rubra (1), ectoproct bryozoans, crustaceans with an amphipod
(2), a cumacean (1), a spider crab (1 jv), a box crab (1 jv), and
a pinnotherid crab (1).

Numbers of species and specimens totalled :

polychaetes 28 species, 102 specimens
echinoderms 4 18
mollusks oy 125
crustaceans 5 6
others + 59

Total: 46* species, 197* specimens

2. Sta. 3609, in 34.5 fms. OPG took 1.07 cuft of gray mud and fine
sand with shell bits and conspicuous ophiuroids. The largest animal was

378 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Cerebratulus rubra, and the most abundant Amphiodia urtica (144).
In addition to those named in the chart, there were nemerteans (2),
Glottidia albida (3), amphipods (many), an anthurid isopod (1).

Numbers of species and specimens totalled :

polychaetes 35 species, 158 specimens
echinoderms 4 171

mollusks 2 ep
crustaceans a =

others 2 5)

Total: 43* species, 336* specimens

bo

NO.

Polychaeta from Santa Catalina Island windward side

HARTMAN :

QUANTITATIVE SURVEY

showing distributions by profiles,

from West End,

+ indicates presence, - absence.
Species represented by more than 10 specimens are indicated by *.

Chaetopterus variopedatus

Ampharete sp. ..

ampharetids ...

Anaitides madeirensis

Anobothrus gracilis
Aricidea lopezi
Aricidea spp. ...
Armandia bioculata .
Artacamella hancocki

Chaetozone armata

Chloeia pinnata ..
Chone gracilis or C.
Cirrophorus furcatus

Cossura candida .

Drilonereis spp. .

Eteone sp. Ss. « « «
BUCHONE Sp. . = » -«
Eulalia myriacyclum
Eurysyllis americana
Exogone uniformis .
Glycera americana .
Glyceralispp. << =. «= «+
Goniada brunnea .

Gyptis a. glabra. .

Harmothoe lunulata .

Isocirrus longiceps

Laonice cirrata ..

ab

+

Lepidasthenia ?longicirrata.

2

West

3

E
4

R.

1

R.

2

Gabe it

1

2

3

4

EisiDs

1

2

Ribbon Rock, Catalina Harbor, Farnsworth Bank,
China Point- Salta Verde, and Palisades.

C+S

1

2

+

Sy/i)

380

Polychaeta from Santa Catalina Island

Loandalltal fauvedil “i921

Loimia montagui....
Lumbrineris spp. ....

Lysippe annectens .. .
Magelona ?pacifica...

Melinna denticulata ..
Myriochele gracilis . .
Nephity.S@S ppm e-nsiel)
Nereis procera... . .-
Newmimnaidesm sip. e616)

Nothria iridescens ...
Notomastus latericeus
Owenia f. collaris ...
Paraonis gracilis ..
Peisidvce aspera. . = «
Photoesgillabray \ . 5 = -
Phyllochaetopterus
limicolus

IPslisieel Sano a ¢ 4 0 ©

Praxillella a. pacifica

Prionospio malmgreni .
Prionospio pinnata.. .
Prionospio pygmaeus
Protodorvillea gracilis
Scalibregma inflatum. .
Scoloplos sp. ....
Spio punctatal 5 3) 27
Sphaerodorum sp. ... .
Sphaerosyllis sp. .. .;

Sthenelanella uniformis

Streblosoma crassi-
branchia
SVALUICISI TG SG G8 ao 5 56

1

West E.

2
+

+

%

3

4

Re
aL

windward side (continued)

Re
2

Cat.

i

2

ALLAN HANCOCK PACIFIC EXPEDITIONS

H.

4

C
1

VOL. 19

ES
2

bi

Pal.

1

2

NO. 2

Polychaeta from Santa Catalina Island

Telepsavus costarum

Terebellides stroemii

Tharyx monilaris . .

Tharyx tesselata .

Vermiliopsis sp. . . .

Aglaophamus sp. .

Ammotrypane aulogaster

Capiteliads <. . .

Cirratulids ...

Haploscoloplos elongatus

Harmothoe spp. . .
Nereis spp. ...
OMUp Madi ae 2 ae en
phyzkliodocid = 2 5
Sabediliade. i.

terebellids ...

Amage anops S6'h 3

Amphicteis scapho-
branchiata

Maldane sarsi ..

mMaldanids ....

Nerinides maculata

Nothria conchylega
Nothria stigmatis
Onuphis parva ..
Onuphis ?vexillaria

Pectinaria californiensis
Pherusa neopapillata
Pista cf. cristata .

Thalenessa spinosa .

Travisia spp. ..

Amaeana occidentalis

il

West

2

+

+

+

bd

3

ve

m

+

Re
al

HARTMAN : QUANTITATIVE SURVEY

381

windward side (continued)

R.
2

*
n

Cat.

a

2

+

+

3

+

H.

m1

+

eB

i

2

+

C+S

a

2

Pals

a

2
+

+

382 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Polychaeta from Santa Catalina Island windward side (continued)

West E. Iai (Chea hs leoletg = (Che; Teel
LS iL 2 2 Ay i, @ 12 al «2

FAR VICE NES GIS o 6 o 6 t sMicmis’ fat -= i =< a Se
MMA! Gb 6 iG 6 6 6 6 aly ome, ware ——— Sp os Ses aes a
Irae NZASabel “EES 5 5 GO Cer a at: —— 35. 55 oy ee ae

Aen des wali Spo 6 64a 6 Oc + - ---- So == es

AncEOnOemS pry <n is! <i ss Oo & wy 6 < + - ---- -- == Ss
Diopatra ?splendidissima Scat oat cles + - ---- a =s ws
Pherusa Gapulaitay ssn Sonor ete Me + - sp Sse {PS oe 7
Poecilochaetus johnsoni Besa + - eee ——— + + <=

Rhodine bitorquata... Se hs saone + + ee == Seca - +

ie

*
+
+

Spiophanes spp. .... od oop + + Sa es =
Sitemnaspas, Ossor! |.) - eee oo + + Pa oe ee + % + &

Brada pluribranchiata . Ai bi, ko or --++# = = S= = 5
Prionospio cirrifera . . oO ho 5 --++# - + ee so
Sthenelais tertiaglabra eS. se IB at --+-— Ss = tem Ss
(NoEeGlAS Hh Go po 6 BiG. hg €e 3 2 + - - - + - -- ==
Apistobranchus ornatus . do Oo oO ae + - - - -- = = = =
Armandia bioculata ... 5 0. etd 5 oo 4+ - = = So = = a
Dorvillea articulata . . Mab he (\ ceteire ae tp = = == = =
Lumbrineris limicola. . SWiswisr tee aac Boo = Sc Se =e
Lumbrineris minima... Ah aesy youu ae *—- — - == =< ss
Mediomastus californiensis ..... 56 ++ - - - - -- =o

Aglaophamus dicirris . . Dd soem A er SP oe = + + fae

Aricidea uschakowi ... sect cte Tas 5 ob -+-- +i a ==
Capaitelila:icapiitatas SuUbSpe) .) ce care - += - + - - - --
Cauilteniedilay sila yeu tien Ot NOES. Same Co = - - - - --
Euchone ancolion .) .) 4 - et “alee oc Ae 5 SS 5 = a =
Eumida, bilineate ... SG tree ae ais ap te - - -- - =
NESTONndGN sic sa ee ac sees Peicehicets® vc ae ee tan - - -- - -
HEsperonoel(Sipieeen ieee ius SG 40 °o Bo rd settee -- - - - -
Langerhansia heterochaeta ..... SC - t+ - - -- - - - -
Lumbrineris palalidaigy sar. dp cch oe teees ATES . ® = - - - i = =

NO. 2

Polychaeta from Santa Catalina Island

Nephtys ferruginea .
Nerinides pigmentata .

Notomastus tenuis

Odontosyllis phosphorea

Polyeirrus Sp... =. « «

Rollydora Sp. . - -

Rhynchopsio sp.

Tharyx ?multifilis or
T. sp.
Ceratocephale c. americana

Dorvillea moniloceras

Glycera capitata
Pista disjuncta .

.

Praxidvella gracilis .

Praxillella, collared

BURQCeE SPs) sis. «
Laonice Sp. «. . «
Lumbrineris

Lumbrineris index

Nephtys cornuta .
Nephtys glabra. .

Spiophanes fimbriata

AULOLYEUS SP. = «

Halosydna sp. ..

Harmothoe

Laeospira

Platynereis bicanaliculata

Arabella sp. ...

Armandia sp... .

Aricidea neosuecica

Caulleriella alata

Dodecaceria sp. .

extenuata

?borealis

cruzensis

.

1

West E.

2

3

m

R.
il

windward side

R.
2

Gate

2
+

rm

HARTMAN : QUANTITATIVE SURVEY

lle

(continued)

EBs

i

2

C+S

i

2

383

Pals

il

+

2

+

384

Polychaeta from Santa Catalina Island windward side (continued)

Eurythoe complanata ..

EXOZONE Spey 6) =). s
(CSmveryAs Spe 4 6 o 09 4

Harmothoe hirsuta ...
HOSTOMS Bo G6 6 o o 6 oC
Hesionura c. difficilis
Lepidonotus! spi. = = = «
Lumbrineris acuta ..
Lumbrineris japonica. .
Lumbrineris latreilli .
LyisudaCemnuineiticar wae -)
Weyetlomel Gig 6 9 5 6 6 C
maldanid, collared...
Nereis ?neonigripes . .
Neninelspree <<
Ophiodromus pugettensis
Ore? Biyo. o 6 4a oo a
Pareurythoe californica

Pherusa papillata .

Pisione remota .....
RELONOSPVONSP ces iene =
Psammolyce spinosa...
Questa caudicirra ..

Salmacina dysteri ..

Scoloplos acmeceps . .

SOrpwlsiels 5 Gin 0 0 Go
Spiophanes bombyx .. .

Sprironrbidse wm ves
Vermiliopsis biformis .
Asychais) Vacenay a). a.

Cirriformia spirabrancha

ALLAN HANCOCK PACIFIC

R.R.

a

EXPEDITIONS

2

Cat.

1

2

3

H.

4

PeBe

i,

2
+
+

+

td

*

*

VOL. 19

C+S

1

2

Pal.

1

2

NO. 2

Polychaeta from Santa Catalina Island windward side

Gonadal sp. 9 =. . . = =

Sthenelais verruculosa

Ampharete arctica ..

Amphisamytha bioculata

Aphrodita armifera .
Diopatra tridentata .
Dorvillea articulata .
Eulalia bilineata .
OWE) Sieg woo om

GANUCemSDs ie) 3 0 se

Megalomma sp. ... .

Nephtys californiensis

Phenusa Sp. . « « « «

Pseudopotamilla sp. .
Rhamphobrachium longi-

setosum
Sabella ?crassicornis

Sabella, another sp. .

Saleulsteiquysl 5 4 Go GG

Chaetozone sp. ....
Notomastus ?magnus . .

Potamilla, oculate..

Myriowenia californiensis

2

West

3

E
4

R.R.

il

2

Cat.

1

2

HARTMAN : QUANTITATIVE SURVEY

H.
3 4

385

(continued)

ial

-B
2

c+S

a
+

+

2

Rae

1

2

386 ALLAN HANCOCK PACIFIC EXPEDITIONS

Echinodermata from Santa Catalina Island windward side,
showing distributions by profiles,

from West End, Ribbon Rock, Catalina Harbor, Farnsworth Bank,

China Point-Salta Verde,

Amphiacantha amphacantha
Amphipholis squamata. .
Leptosynapta albicans .

Lytechinus anamesus . .
Gyojorwnaowucls Go 4 a 6

Amphiodia digitata...
Amphioplus hexacanthus .
Brissopsis pacifica ..
Amphiodia urtica... .

DSieeoMGs 6 6 5 6 6 Go
holothunrords) <=

Lusdvassolwvollatal.) sn.

Ophiothrix spiculata. .
Amphioplus strongyloplax

Pentamera pseudo-
populifera

Amphipholis pugetana. .
Amphiura arcystata.. .

Astropecten californicus

Lovenia cordiformis .

Ophiacantha diplasia. .
Ophiocynodus corynetes .
Ophiopholis longispina .
Leptosynapta sp... .
Molpadia intermedia
Ophiomusium jolliensis .
Op elen Sor ge wo A oo x
Ophiopteris papillosa

Asterina miniata =. 2 .

Ophiopsila californica

al

West

2

+

+

3

15;
m1

+

R.

al

and

R.
2

+

&
n

Palisades.

Gate iH. aldig (ches
2

ala) 3}, nh 73
=) xe =- = + =
ee *& % ee
ee ita ort - - - =
i ns ———
e & 2&6 a ———
=. aS = =
= Son k ——
oo Sa a =e
Sco mS a a
5 @o < + # a
Ae Get a =o
pS == wos
a) tee = i =)
A -f.e 4 a Se
ety ee = 5S a
AAC es SS ae
a - - - =
si ot ote == ==

. + = S 5

hi oleae nas + = =--
: c + - -
— + =

+ - =

VOL. 19

ae

\ \ Kp

be

oo ihe ainsie
<_<

NO. 2

HARTMAN : QUANTITATIVE SURVEY

387

Echinodermata from Santa Catalina Island windward side (continued)

Ophiura iutkent « «

purple holothuroid. .

Mollusca from Santa Catalina Island windward side,

West E.
2 more eS

R.R.

1

2

Cater. [Ps

23 4 HL

showing distributions by profiles,
from West End, Ribbon Rock, Catalina
China Point-Salta Verde,

Sastropods = =<. . = « «
pelecypods .... .
solenogaster .... .

Pecten Sp. .*. « «© « «

Tellina carpenteri. .
Cuspidaria sp. ....
Nassarius sp. ... =.
Antiplanes sp. ... .
Cadudussispp. # = - «= -

Acila castrensis ...

Amygdalum pallidulum .
Epitonium sp. ... .-

Lima subauriculata. .
Lucinoma annulata ..

Nemocardium sp. ...

Compsomyax subdiaphana .

ihyasima Sp. = % = « «
Tellina idae .....
Cardita ventricosa .

Calliostoma sp. ...

Conus californicus . .

Hyalina sp. (olive-shell)

Lima dehiscens ....

Pedicularia californica

HeaniastceS eS Diets hs. ©) sure

+

+

+

we
”

Harbor,

and Palisades.
= A eas ay *
= ee =
= ee cae =
= Sy Z
= ee a a
+ are =
= ete =
+ Soe =
+ yy te
+ = fs =
+ ws =
De ee
+ ae =
+ - + =
+ - + =
: ee
‘5 + =
. 2. e ;
aitelane =
° Suetae a2
ne OSES So
+

C+S
2

+ -

x +

Farnsworth Bank,

Pal.

1

2

*

388 ALLAN HANCOCK PACIFIC EXPEDITIONS VOR. 19

Mollusca from Santa Catalina Island windward side (continued)

West E. Rains (Gaue5 Inls oly (ChPS} eh

ae Se i 7 2 gu iy il, 2 aL
GHEY Sdn o 6 oO Go OC ao Olle “US 5 Oo ac 5 - - -
Saxicava arctica... . do Ga o 5 0 oie Coe 5 - - -
Weuanieneael 5 56 5 Gg 6 Oo a o SG. 6 o « ae 6 om 6 ome -- -
Npileiyel YG 6 6 OD oO oO f a bo oC 9 6 55 0 6 co + - ~

Nuculana taphria.... Bo 5 oo 5 6 weal: 5 0 * = -
I GEUONAeINVAN 5 G G Oo 50 6 6 Alec oy lomo ah ie Aue Geto a % ar -
SubilauaWwicy JUebGCEy G iG 5 4 Sued seu cice ae Seas ree te es + = -
(EUICTS C—O Dees nel olel nie) iets Ol Go «ce 26 ns 5 Gide ar <6 & £0 + - -

Caspicwile Bog of6 0 6 @ S65 560 ©6 8665 oof oo &

SOLGN SDd| c SB olo-o G@ a 6 66 oo S & 5» 646 od cso &

NO. 2 HARTMAN : QUANTITATIVE SURVEY 389

PHOTOGRAPHIC RECORDS ALONG THE LEEWARD
SIDE OF SANTA CATALINA ISLAND

The sea bottoms on the leeward side of Santa Catalina Island were
sampled with a Campbell grab equipped with camera. I am indebted to
Professor K. O. Emery, Woods Hole Oceanographic Institution, for the
copies of the photographs. The methods for taking the samples were like
those described in the canyon report (Hartman, 1963, p. 102). The
Campbell grab, with coverage of about a fourth of a square meter, was
equipped with camera and lights, and lowered so that the size of the
photograph and the surface taken by the grab were about equal. There
was reasonable agreement with what the pictures revealed, and the
numbers and kinds of larger surface organisms taken in the grab.

The locations of photographs and their analyses are listed in the
chart and summaries, below.

1. Sta. 7707 (N of no. 250), off Abalone Pt., in 22.5 fms. The
photograph (PI. 8, top) shows numerous slender, projecting tubes of
Telepsavus costarum, and larger, recumbent tubes of Chaetopterus vario-
pedatus (the largest tubes measured 225 by 18 mm) ; three echinoids and
two cone-snails are seen on the surface. The largest animals in the sample
are shown (PI. 8, bottom). Analyses revealed the presence of 21 species
and 155 specimens; in addition to those listed in the chart (see below),
there were bryozoan clusters, an amphipod (1), a commensal crab (1)
and an ascidian. Largest were Chaetopterus and Lytechinus.

2. Sta. 7719 (N of no. 250). Shelf off Abalone Pt., in 23.5 fms. The
photograph (Pl. 9, top) shows a silty surface interrupted by two tubes of
Chactopterus variopedatus, distal ends of slenderer tubes of Te/epsavus
costarum, and Lytechinus anamesus. Other animals in the sample resem-
ble those from Sta. 7707. The photographs appear different because the
surface is more completely silt-covered in the second than in the first.

3. Sta. 7720 (N of no. 253). Shelf off Abalone Pt., in 23.5 fms.
The photograph (PI. 9, bottom) shows a bottom similar to that in the
preceding one, except for the presence of a seastar, Mediaster aequalis.
Chaetopterus and Telepsavus were the most prominent organisms.

4. Sta. 7723 (near no. 250). Shelf off Jewfish Pt., in 39.5 fms. The
photograph (PI. 10, top) shows a level silty bottom, with rocks project-
ing at lower left, overgrown by solitary corals, and a smaller clump of
the same in the upper left-hand corner; an ophiuroid arm is seen in upper
right, and the surface shows pittings of smaller organisms embedded in

390 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

the sediments. he grab failed to penetrate the bottom, perhaps because
the jaws struck hard rock before they closed.

5. Sta. 7725 (near no. 250). Shelf off Jewfish Pt., in 40.5 fms. The
photograph (PI. 10, bottom) shows a silty surface with a few long,
ophiuroid arms, many small pits, mounds and low ridges. The sample
contained an echinoid and numerous polychaete tubes, together with a
ceriantharian (8), a sipunculid burrowing in the foraminiferan, Rhab-
damina (2), a chaetognath (15), amphipods (65), a gnathid isopod
(1), an anthurid isopod (1), tanaids (12), cumaceans (22), ostracods
(41), a shrimp (3), a pycnogonid (2), in addition to those named in
the chart. The most abundant species were 4ricidea neosuecica (42) and

Aricidea lopezi (25).

6. Sta. 7726 (near no. 251). Shelf off Jewfish Pt., in 40 fms. The
photograph (PI. 11, top) shows silty sediments overlain by dead shells of
Laqueus californicus. Analyses of the sample revealed the presence of
more than 100 species and 300 specimens. In addition to those named in
the chart, the sample contained a ceriantharian (3), a solitary coral (2),
a hydroid stalk (1), a nemertean (1), a polyclad (1), a sipunculid (1),
a phoronid (1), encrusting bryozoans, amphipods of numerous kinds
(60*), a caprellid (1), isopods (5), tanaids (14), cumaceans (3), three
kinds of ostracods (13), a shrimp (5), an oxyrhynch crab (3), Scalpel-
lum (1) attached to Phyllochaetopterus tube. The largest animals were
Euclymene sp. (Pl. 11, bottom) and dead shells of Laqueus. The most
abundant species was Peisidice aspera (17).

7. Sta. 7712 (near no. 260). Slope SE end of Santa Catalina Island,
in 133 fms. The photograph (Pl. 12, top) shows a coarse black sandy
bottom, its surface marked with a broad, shallow, depressed trail in the
lower half, a patch of ophiuroid arms in center right, and the surface
pitted with pores, mounds and other irregularities. The largest animals
were Brissopsis pacifica, (Pl. 12, bottom), Eunice americana, Nothria
iridescens, Maldane sarsi and tubes of Phyllochaetopterus limicolus. In
addition to those named in the chart, there were a ceriantharian (1), a
nemertean (1), a sipunculid (2), amphipods of several kinds (12), an
anthurid isopod (1), a tanaid (3), a cumacean (1), and three kinds of
ostracods (43). None was conspicuously abundant.

8. Sta. 7727 (near no. 251). Shelf off Jewfish Pt., in 44 fms. The
grab failed to take a sample. The photograph (PI. 13, top) reveals a
silty sand surface riddled by pores and depressions, suggesting the pres-
ence of many animals.

No. 2 HARTMAN : QUANTITATIVE SURVEY 391

9. Sta. 7713 (near no. 267). Basin slope, SE of Santa Catalina
Island, in 286 fms. The sediments were black sand and gravel with silt.
The photograph (PI. 13, bottom) showed no large surface animals, but
many small to large pores, mounds, trails and white shelly fragments.
The analyses disclosed the presence of 32 species and 139 specimens. In
addition to those named in the chart, there were glassy sponge spicules,
a sea-whip with bulbous base (1), a nemertean (1), an echiuroid with
anterior end lightly papillated (1), amphipods (3), a caprellid (1), a
cumacean (1), and an ostracod (1). Capsules of unknown origin, meas-
uring 4+ by 1.2 mm, surrounded by a chitinized layer, were filled with
uniformly minute, spherical granules. ‘Ihe most abundant species were
Paraonis gracilis oculata (62), and Crystallophrisson sp. (15), and the
largest individuals were Scalibregma inflatum and an echiuroid.

392 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Polychaeta on photographic records
from the leeward side of Santa Catalina Island,
showing order of occurrence in 22.5 to 268 fms.
+ indicates presence, - absence.
Species represented by more than 10 specimens are indicated by *.

al 2 3 4 5 6 7 8

Aglaophamus dicirris . C 5 4 + - - - = = = =
Anaitides, checkered . 3 4 5 + = = = = = = =

Aphrodita sp. 5 5 oC : . + = = = = = = =

Chaetopterus variopedatus 3 ‘ % + + = - = = =
Langerhansia heterochaeta c ; + - - - = = - =
Lepidasthenia sp. : 5 2 - + = = = = ~ = as
Owenia f. collaris . : O + + + = - = = =
Telepsavus costarum . : ° : * - - = + + = +
terebellid g fi : 6 4 A + = - = = + = =
Amphicteis scaphobranchiata . : ; 6 : : + - - -
Aricidea lopezi . a 4 . é : 5 , 5 * + * *
Aricidea neosuecica . a : : ; : . 6 % + * -
Aricidea uschakowi ‘ < 3 - A ‘ 6 ; * = * +
Chaetozone setosa . 2 : C : f > : + + - -
Chone sp. > 5 ° 5 . . 5 2 . ° + + + -
Eteone ?californiensis A 5 A . : é + - - -
goniadid . 3 5 : : 6 : q ; 6 : + + = =
Harmothoe lunulata 6 C f : 6 : 5 + + - -
Lumbrineris bicirrata : : C 5 : : : + - we -
Myriowenia californiensis 9 6 . 5 6 5 + - - =
Nephtys cornuta . : A 4 6 : 5 : x - - -
Nephtys ferruginea . : 55 C A c ; O + - + -
Nerinides pigmentata . : : 5 5 5 : : + a + -
Nothria iridescens - : A 5 : . : 3 + - - +
Onuphis parva 2 S 5 5 : 5 é 5 5 + - - +
Paraonis gracilis A 6 5 S 6 8 : : + + + +
Pectinaria californiensis 3 A 5 . A : + - + +
Pherusa neopapillata . : ; C 6 & : < + - - -

Pholoe glabra ' : . ; : C ; : 6 + + + c

NO. 2

HARTMAN : QUANTITATIVE SURVEY

Polychaeta on photographic records (continued)

Polydora sp. . ° :
Rhodine bitorquata :
Spio punctata : :
Sthenelais tertiaglabra

Sthenelanella uniformis
Terebellides stroemii
Thalenessa spinosa :
Tharyx monilaris . -
Aglaophamus sp. . °

Amaeana occidentalis .

Ammotrypane sp. . 5
Anaitides sp. :
Autolytus sp. ° é
Euclymene sp. °
Eulalia myriacyclum .

Euphrosine sp. G
Exogone uniformis :

Flabelligera infundibularis

Glycera americana :
Glycera cf. capitata .
Lanice sp. : ;
Laonice foliata . .
Lepidonotus caelorus .
Lumbrineris cruzensis

Lumbrineris, other sp.

Magelona pacifica :

Mediomastus californiensis

Notoproctus cf. pacificus

Odontosyllis phosphorea
Ophiodromus pugettensis

Peisidice aspera .

1

2

3

4

5

-

6

Ge

393

394 ALLAN HANCOCK PACIFIC EXPEDITIONS

Polychaeta on photographie records (continued)

Pherusa capulata . 5 5

Phyllochaetopterus prolifica
Platynereis bicanaliculata

Polycirrus sp. . : :
Praxillella sp. . 5 C
Prionospio malmgreni . :

Prionospio pinnata : ;
Pseudopotamilla occelata .
Scalibregma inflatum .
Tharyx tesselata .
Vermiliopsis biformis
other serpulid 6 5
Myriochele gracilis . :
Prionospio pygmaeus . 6
Spiophanes sp. : 5
Sternaspis fossor : :
Thanysc (SPP 5 = C
Ampharete sp. : $ .
Cirrophorus aciculatus é

Cossura candida . 4 s
Eunice americana . 6 5
Goniada brunnea . B 5
Maldane sarsi a : ‘
Notomastus sp. 5 ° 4
Pherusa sp. . 3 : 5

Phyllochaetopterus limicolus
Prionospio cirrifera . C
Spiophanes fimbriata . 5

Ancistrosyllis tentaculata
Anobothrus gracilis .

Brada glabra . A 0

2

3

4

5

6

+

VOL. 19

HARTMAN : QUANTITATIVE SURVEY

Polychaeta on photographic records (continued)

Brada pluribranchiata
Califia calida
Glycera branchiopoda .
Gyptis a. glabra . .
Harmothoe sp. 4
Nephtys sp. . : A
Notomastus magnus
Paraonis g. oculata .

Praxillella a. pacifica

Echinodermata on photographic records

1

2

3

mn

5

6

from the leeward side of Santa Catalina Island,
showing order of occurrence in 22.5 to 268 fms.

Lytechinus anamesus .

Ophiothrix spiculata .
Mediaster aequalis -

ophiuroids : 5 2
Amphiodia urtica .
holothurian . - 7
seastar . . : .

Leptosynapta albicans
Brissopsis pacifica .
Amphipholis squamata .

+

+

+

+

+

*

395

%

396

ALLAN HANCOCK PACIFIC EXPEDITIONS

Mollusca on photographic records
from the leeward side of Santa Catalina Island,
showing order of occurrence in 22.5 to 268 fms.

Compsomyax subdiaphana 3

Conus calif

Pecten sp.

ornicus A 5

Amygdalum pellucidum . :

Cadulus tolmiei . 9 C

gastropods

Nemocardium sp. . c

pelecypods
Baleis sp.

chiton ;

Cyclopecten sp. . g ‘

Nassarius s
seaslug .

Tellina sp.

Axinopsida

p- : C 5

serricata . S

Crystallophrisson sp. 5
Dacrydium pacifica 6 5

turret -top

Cadulus sp.

Protochaetoderma sp. . 5

scaphopod

uf

+

2

3

4

5

6

VOL.

19

NO: 2 HARTMAN : QUANTITATIVE SURVEY 397

ALPHABETICAL LISTS OF SPECIES NAMED FROM
THESE AREAS

POLYCHAETES

A edicira ramosa (Annenkova) ; SP, CL

A glaophamus dicirris Hartman; CL, CW

A glaophamus erectans Hartman; SP

A glaophamus spp.; SP, LS, CL

A maeana occidentalis (Hartman) ; PV, SP, NE, CL, CW
A mage anops ( Johnson) ; PV, SP, CL, CW

Ammotry pane aulogaster Rathke; SP, LS, CL, CW
Ammotrypane sp.; SP, LS, CL

A mpharete arctica Malmgren; PV, SP, CL, CW
Ampharete labrops Hartman; PV

Ampharete spp.; PV, SP, CL

Amphicteis mucronata Moore; SP, CL

A mphicteis scaphobranchiata Moore; PV, SP, NE, LH, CL, CW
A mphicteis spp. ; SP, CL

Amphiduros pacificus Hartman; PV

A mphisamytha bioculata (Moore) ; CL, CW

A naitides madeirensis (Langerhans) ; PV, SP, LS, CL, CW
A naitides medipapillata (Moore) ; PV, SP, CL

A naitides, checkered ; SP, NE, LH, CL

Anaitides spp.; PV, SP, NE, LS, CL, CW

Ancistrosyllis cf. bassi Hartman; SP

Ancistrosyllis breviceps Hartman; CL

Ancistrosyllis cf. rigida Fauvel; SP, NE

Ancistrosyllis tentaculata Treadwell; LH, PV, SP, NE, CL, CW
Ancistrosyllis spp.; LH, SP, LS

Anobothrus gracilis (Malmgren) ; SP, LS, CL, CW
Anotomastus gordiodes (Moore) ; LH, PV, SP, NE
Antinoella sp.; PV, SP

A phrodita armifera Moore; SP, CW

A phrodita parva Moore; CL

A phrodita refulgida Moore; PV

A phrodita spp.; PV, SP, CL

398 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

A pistobranchus ornatus Hartman; SP, CW

A pomatus geniculata (Moore and Bush) ; CL

Arabella semimaculata (Moore) ; SP

Arabella sp.; SP, CW

Arctonoe sp.; CL, CW

Aricidea lopezi Berkeley and Berkeley; LH, PV, SP, NE, LS, CL, CW
Aricidea neosuecica Hartman; PV, SP, NE, CL, CW
Aricidea uschakowi Zachs ; PV, SP, NE, CL, CW
Aricidea spp; HYP V. SP. Ls, Cl, Cw

Armandia bioculata Hartman; LH, PV, SP, CL, CW
Artacamella hancocki Hartman; PV, SP, NE, CL, CW
Asabellides lineata ( Berkeley and Berkeley) ; NE
Asabellides sp.; PV, NE

Asclerocheilus californicus Hartman; SP

Asclerocheilus sp.; CL

Asychis disparidentata (Moore) ; PV, SP

Asychis lacera (Moore) ; SP, CW

Asychis spp.; LH, PV, SP

Autolytus sp.; PV, SP, CW

A xiothella rubrocincta (Johnson) ; PV, SP, NE

A xiothella spp.; SP, NE, CL

Boccardia basilaria Hartman; PV

Boccardia nr. redeki (Horst) ; LH

Boccardia uncata Berkeley ; PV

Boccardia spp.; PV, SP, LS

Brada glabra Hartman; SP, NE, LS, CL

Brada pilosa Moore; LH, PV, SP, NE, CL

Brada pluribranchiata (Moore) ; PV, SP, NE, CL, CW
Brada sp.; SP

Califia calida Hartman; SP, CL

Capitata ambiseta Hartman; PV

Capitella capitata ( Fabricius), subspp.; PV, SP, LS, CL, CW
Capitellides sp.; PV

Capitomastus sp.; PV, SP

Carazzia sp.; LH

Caulleriella alata (Southern) ; PV, CW

NO. 2 HARTMAN : QUANTITATIVE SURVEY 399

Caulleriella bioculata (Keferstein) ; PV, SP

Caulleriella spp. ; LS, CL, CW

Ceratocephala c. americana Hartman; PV, NE, SP, CW
Ceratocephala 1. pacifica Hartman; SP

Ceratonereis sp.; SP

Chaetopterus variopedatus (Renier) ; PV, SP, NE, CL, CW
Chaetozone armata Hartman; LS, CL, CW

Chaetozone corona Berkeley and Berkeley ; LH, PV, SP, NE, CL
Chaetozone gracilis (Moore) ; PV, SP

Chaetozone hamata (Hartman) ; SP

Chaetozone multioculata Hartman; CL

Chaetozone setosa Malmgren; PV, SP, CL

Chaetozone, oculate ; PV

Chaetozone spp.; SP, CL, CW

Chloeia pinnata Moore; PV, SP, NE, LS, CL, CW

Chone ecaudata (Moore) ; LH, PV

Chone gracilis Moore; PV, SP, NE, CW

Chone minuta Hartman; SP

Chone mollis (Bush) ; PV, SP

Chone spp.; PV, SP, NE, CL, CW

Chrysopetalum occidentale Johnson; PV

Cirratulus cirratus (Miller) ; LH, PV, SP

Cirratulus sp.; CL

Cirriformia luxuriosa (Moore) ; LH, SP

Cirriformia spirabrancha (Moore) ; PV, SP, CW
Cirriformia sp.; CL

Cirrophorus aciculatus Hartman; PV, SP, LS, CL
Cirrophorus furcatus Hartman; LH, PV, SP, LS, CL, CW
Cistenides brevicoma ( Johnson) ; PV, SP

Cossura candida Hartman; LH, PV, SP, NE, LS, CL, CW
Dasybranchus glabrus Moore; PV

Dasybranchus sp.; PV

Dexiospira sp.; CL

Diopatra ornata Moore; PV, SP, NE, CL

Diopatra ?splendidissima Kinberg ; CW

Diopatra tridentata Hartman; LH, PV, SP, CL, CW

400 ALLAN HANCOCK PACIFIC EXPEDITIONS

Diopatra sp.; CL

Distylidia rugosa (Moore) ; PV
Dodecaceria concharum Oersted ; LS, CL
Dodecaceria spp. ; PV, CL, CW

Dorvillea articulata (Hartman) ; LH, PV, SP, CL, CW
Dorvillea moniloceras (Moore) ; CW
Drilonereis longa Webster ; PV, SP, LS
Drilonereis nuda Moore; LH, PV, SP
Drilonereis spp.; LH, PV, SP, NE, CL, CW
Eteone ?alba Webster ; LH

Eteone californica Hartman; PV, SP

Eteone dilatae Hartman; SP

Eteone spp.; CL, CW

Euchone incolor Hartman; CW

Euchone spp.; PV, SP, LS, CL, CW
Euclymene reticulata Moore; PV, SP
Euclymene sp.; PV, SP, CL, CW
euclymenid ; SP

Eulalia myriacyclum Schmarda; PV, SP, CL
Eulalia quadrioculata Moore; PV, SP
Eulalia, cross-barred ; PV

Eulalia spp. ; SP, CL

Eumida sanguinea (Oersted) ; PV, SP
Eumida tubiformis Moore; CL

Eumida, bioculate; PV, SP

Eumida, trilineate; PV, SP, CL

Eumida spp: PV «Sb. nes, Cl

Eunice americana Hartman; PV, SP, NE, CL
Eunice antennata (Savigny), PV

Eunice aphroditois (Pallas) ; CL

Eunice sp.; LS, Cre

?Eunoe sp.; CL

Euphrosine sp.; SP, CL

Eupolymnia crescentis Chamberlin; PV
Eupolymnia heterobranchia ( Johnson) ; CL

Eupolymnia spp.; SP, CL

VOL. 19

No. 2 HARTMAN : QUANTITATIVE SURVEY

Eusyllis transecta n. sp.; SP

Euzonus (Thoracophelia) sp.; PV

Exogone uniformis Hartman; PV, SP, NE, LS, CL
Exogone spp.; PV, CL

Exogonella brunnea Hartman; PV, CL
EXOGONINAE; PV, SP

Fabricia pacifica Berkeley and Berkeley ; CL
Fabricia spp.; LH, PV, SP, CL

Flabelligera commensalis Moore; PV, SP, CL
Flabelligera infundibularis Johnson ; CL

Gattyana brunnean. sp.; LS

Genetyllis castanea (Marenzeller) ; SP

Glycera americana Leidy ; LH, PV, SP, NE, LS, CL
Glycera branchiopoda Moore; SP, LS, CL

Glycera ?capitata Oersted ; LH, PV, SP, NE, LS, CL
Glycera convoluta Keferstein; NE

Glycera oxycephala Ehlers ; SP, NE

Glycera robusta Ehlers; SP

Glycera tenuis Hartman; SP

Glycera tesselata Grube; LS, CL

Glycera spp.; PV, SP, LS, CL

Glycinde armigera Moore; SP, NE

Glycinde wireni Arwidsson; SP

Goniada brunnea Treadwell; LH, PV, SP, NE, LS, CL

Goniada littorea Hartman; LH, PV, SP, NE
Goniada sp.; SP

Gy ptis arenicola glabra (Hartman) ; LH, PV, SP, NE, LS, CL

Gy ptis brunnea (Hartman) ; PV
Gypftis sp.; SP

Halosydna brevisetosa Kinberg ; PV, SP
Halosydna latior Chamberlin ; SP, CL
Halosydna spp.; PV, CL

Haploscoloplos elongatus (Johnson) ; LH, PV, SP, NE, LS, CL

Haploscoloplos panamensis Monro; CL
Haplosyllis sp.; PV
Harmothoe ?fragilis Moore; CL

401

402 ALLAN HANCOCK PACIFIC EXPEDITIONS

Harmothoe hirsuta Johnson; CL

Harmothoe ct. imbricata (Linnaeus) ; SP
Harmothoe lunulata (delle Chiaje) ; LH, PV, SP, NE, CL
Harmothoe priops Hartman; SP, NE

Harmothoe scriptoria Moore; PV, SP, CL
Harmothoe spp.; PV, SP, NE, CL

harmothoid ; SP

Hesperonoe laevis Hartman; PV, SP

Hesperonoe sp.; SP, NE

Heteromastus filiformis (Claparede) ; PV
Heteropale bellis ( Johnson) ; SP

Heterospio catalinensis (Hartman) ; PV, SP, NE, CL
Hyalinoecia juvenalis Moore; NE

?Tlyphagus sp.; SP

Tsocirrus longiceps (Moore) ; SP, NE

Tsocirrus sp.; CL

?Isoldasp.; SP, CL

Lagisca nr. multisetosa Moore; LS

Lagisca sp.; SP

Langerhansia heterochaeta (Moore) ; PV, SP, CL
Lanice conchilega (Pallas) ; PV, SP, LS, CL
Lanice sp.; CL

Laontce cirrata (Sars) ; LH, PV, SP, NE; LS, Ck
Laonice foliata (Moore) ; CL

Laonice sp.; SP

Leaena caeca Hartman; SP

Leanira sp.; NE

Leiochrides hemipodus Hartman; CL

Leiochrides sp.; CL

Lepidasthenia interrupta (Marenzeller) ; SP
Lepidasthenia longicirrata Berkeley ; CL
Lepidasthenia virens (Blanchard) ; PV, SP, CL
Lepidasthenia spp.; PV, NE, CL

Lepidonotus caelorus Moore; PV, SP, LS, CL
Loandalia fauveli Berkeley and Berkeley; LH, PV, SP, NE
Loimia medusa (Savigny) ; PV, NE

voL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY 403

Lumbriclymene lineus Hartman; SP

Lumbriclymene sp.; SP

Lumbrineris nr. acuta Verrill; PV, SP, NE, CL
Lumbrineris cf. bassi Hartman; PV

Lumbrineris bicirrata Treadwell; PV, SP, NE, CL
Lumbrineris bifilaris (Ehlers) ; NE

Lumbrineris californiensis Hartman; LH, PV, SP, NE, CL
Lumbrineris cruzensis Hartman; LH, PV, SP, NE, CL
Lumbrineris nr. impatiens (Claparéde) ; CL
Lumbrineris index Moore; SP

Lumbrineris inflata Moore ; PV

Lumbrineris japonica (Marenzeller) ; SP

Lumbrineris latreilli Audouin and Edwards; PV, SP, LS, CL
Lumbrineris limicola Hartman; LH, PV, SP
Lumbrineris pallida Hartman; LH, PV, SP
Lumbrineris nr. sarsi (Kinberg) ; SP

Lumbrineris ?tetraura (Schmarda) ; PV, SP
Lumbrineris spp.; PV, SP, NE, LS, CL

Lysippe annectens Moore; PV, SP, LS, CL
Macellicephala remigata (Moore) ; LS

Magelona ?californica Hartman; LS

Magelona pacifica Monro; PV, SP, NE, LS, CL
Magelona pitelkai Hartman; SP

Magelona sacculata Hartman; LH, SP, NE

Magelona spp.; LH, PV, SP, CL

Maldane sarsi Malmgren; PV, SP, LS, CL

Maldane sp.; SP

Maldanella robusta Moore; SP

Mar physa disjuncta Hartman; LH, PV, SP

Marphysa mortenseni Monro; PV

Mar physa sanguinea (Montagu) ; CL

Mediomastus californiensis Hartman; LH, PV, SP, NE, CL
Megalomma splendida (Moore) ; PV, SP, CL
Megalomma, bioculate ; SP

Megalomma spp.; LH, PV, NE, CL

Melinna denticulata Moore; PV, SP, CL

404 ALLAN HANCOCK PACIFIC EXPEDITIONS

Melinna spp.; LH, PV, SP

Melinnexis moorei Hartman; SP

?Mesochaetopterus sp.; CL

Myriochele gracilis Hartman; PV, SP, NE, LS, CL, CW
myriocheliid ; SP

Myriowenia californiensis Hartman; PV, SP, NE, CL, CW
Myxicola infundibulum (Renier) ; PV, SP

Naineris nr. quadricuspida (Fabricius) ; SP

Naineris uncinata Hartman; PV, SP, NE, CL

Neanthes brandti (Malmgren) ; PV

Nephtys assignis Hartman ; SP

Nephtys caecoides Hartman; SP, NE

Nephtys californiensis Hartman; SP, CW

Nephtys cornuta Berkeley and Berkeley ; LH, SP, CL, CW
Nephtys ferruginea Hartman; PV, SP, NE, LS, CL, CW
Nephtys glabra Hartman; SP, CW

Nephtys punctata Hartman; SP

Nephtys spp.; LH, PV, SP, NE, LS, CL, CW
Nereiphylla sp.; SP

Nereis latescens Chamberlin ; SP

Nereis mediator Chamberlin ; CL

Nereis pelagica neonigripes Hartman; PV, CW

Nereis procera Ehlers; LH, PV, SP, NE, CL, CW
Nereis zonata Malmgren; SP

Nereis spp.; PV, CL, CW

Nerine sp.; CW

nerinid ; PV

Nerinides ?acuta (Treadwell) ; CL

Nerinides maculata Hartman; SP, CL, CW

Nerinides pigmentata (Reish) ; SP, CL, CW

Nerinides sp. ; CW

Nicomache sp.; SP, LS, CL

Ninoe gemmea Moore; PV, NE, SP

Ninoe sp.; CL

Nothria conchylega (Sars) ; CL, CW

Nothria elegans (Johnson) ; PV, SP, CL

voL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY 405

Nothria geophiliformis (Moore) ; SP

Nothria hiatidentata Moore; CL

Nothria iridescens (Johnson) ; LH, PV, SP, NE, CL, CW
Nothria pallida Moore; PV, SP

Nothria stigmatis (Treadwell) ; SP, CL, CW

Nothria spp. ; PV, SP, CL

Notocirrus sp.; PV

Notomastus latericeus Sars ; CL, CW

Notomastus lineatus Claparede; PV, SP, LS, CL
Notomastus magnus Hartman; PV, SP, NE, LS, CL, CW
Notomastus tenuis Moore; LH, PV, SP, CW

Notomastus spp.; PV, SP, LS, CL

Notoproctus pacificus (Moore) ; CL

Odontosyllis phosphorea Moore; PV, NE, CL, CW
Odontosyllis sp.; PV, SP, CL

Oncoscolex pacificus (Moore) ; SP, CL

Onuphis eremita Audouin and Edwards; SP, NE

Onuphis nebulosa Moore; SP, CL

Onuphis parva Moore; PV, SP, NE, CL, CW

Onuphis vexillaria Moore; CL, CW

O phelia limacina (Rathke) ; SP

Orseis sp.; CW

Owenia fusiformis collaris Hartman; PV, SP, NE, CL, CW
Panthalis pacifica Vreadwell; PV, SP, NE, CL

Panthalis sp.; SP

Paradexiospira vitrea (Fabricius) ; SP

Paranitis polynoides (Moore) ; SP

Paraonis gracilis (Tauber) ; LH, PV, SP, NE, LS, CL, CW
Paraonis gracilis oculata Hartman ; SP, CL

Paraonis sp.; abranchiate ; CL

Pareule pis fimbriata (Vreadwell) ; NE

Pareurythoe californica ( Johnson) ; PV, SP, CL, CW
Pectinaria californiensis Hartman; LH, PV, SP, NE, CL, CW
Peisidice aspera Johnson; LH, PV, SP, LS, CL, CW
Pherusa capulata (Moore) ; PV, SP, NE, CL, CW
Pherusa inflata (Vreadwell) ; PV, SP, CL

406 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Pherusa neopapillata Hartman; LH, PV, SP, NE, CL, CW
Pherusa papillata ( Johnson) ; CL, CW

Pherusa spp.; PV, SP, CL, CW

Pholoe glabra Hartman; LH, PV, SP, NE, CL, CW
Phragmatopoma californica (Fewkes) ; CL
Phyllochaetopterus limicolus Hartman; PV, SP, NE, LS, CL, CW
Phyllochaetopterus prolifica Potts; PV, SP, CL
Phyllochaetopterus sp.; CL

Phyllodoce ferruginea Moore; SP

Phyllodoce cf. papillosa Uschakov and Wu; SP
Phyllodoce sp.; SP

Phylo felix Kinberg ; CL

Pilargis berkeleyi Monro; LH, PV, SP, NE

Pilargis hamatus Hartman; LH, PV, SP

Pilargis maculata Hartman; SP

Pisione remota (Southern) ; PV, CW

Pista alata Moore; PV

Pista cf. cristata (Miller) ; LH, PV, SP, NE, CL, CW
Pista disjuncta Moore; PV, SP, NE, CW

Pista elongata Moore; PV, CL

Pista moorei Berkeley and Berkeley ; SP

Pista spp. ; LH, PV, SP, NE, CL, CW

?Placostegus sp.; CL

Platynereis bicanaliculata (Baird) ; PV, SP, NE, CL, CW
Poecilochaetus johnsoni Hartman; PV, SP, NE, CL, CW
Polycirrus californicus Moore; CL

Polycirrus spp.; PV, SP, LS, CL, CW

Polydora nr. armata Langerhans ; LH, CL

Polydora nr. caulleryi Mesnil; SP

Polydora citrona Hartman; LH

Polydora ligni Webster ; LH, CL

Polydora limicola Annenkova; LH, PV

Polydora socialis (Schmarda) ; PV

Polydora ?spongicola Berkeley and Berkeley ; SP
Polydora cf. websteri Hartman; LS

Polydora spp.; LH, PV, SP, NE, CL, CW

NO. 2 HARTMAN : QUANTITATIVE SURVEY 407

Polyophthalmus pictus (Dujardin) ; CL

Potamethus mucronatus (Moore) ; SP, CL

Potamethus sp.; LS

Potamilla sp.; PV, CW

Praxillella affinis pacifica Berkeley ; LH, PV, SP, NE, LS, CL, CW
Praxillella gracilis (Sars) ; PV, SP, NE, CW
Praxillella, collared ; CW

Praxillella spp.; SP, CL

Praxillura maculata Moore; CL

Prionospio cirrifera Wirén; LH, PV, SP, CL, CW
Prionospio malmgreni Claparéde; PV, SP, NE, CL, CW
Prionospio pinnata Ehlers; PV, SP, NE, LS, CL, CW
Prionospio pygmaeus Hartman; SP, CL

Prionospio spp.; PV, SP, NE, LS, CL

Protis pacifica Moore; SP, CL

Protodorvillea gracilis (Hartman) ; PV, SP, LS, CL, CW
Protula superba Moore; CL

Psammolyce spinosa Hartman ; CW

Psammolyce sp. ; CL

Pseudopotamilla occelata Moore; PV, NE, CL
Pseudopotamilla, oculate ; PV

Pseudopotamilla spp.; PV, SP, CL, CW

Questa caudicirra n. gen., n. sp. ; LS, CW

Raricirrus maculata Hartman; PV

Rhamphobrachium longisetosum Berkeley and Berkeley ; SP, CL, CW
Rhodine bitorquata Moore; SP, NE, CL, CW
Rhynchospio spp.; NE, CL, CW

Sabella crassicornis Sars; SP, CW

Sabella sp.; PV, SP, CW

Sabellaria cementarium Moore; PV, SP, CL

Saccocirrus papillocercus Bobretzky ; SP

Salmacina dysteri Huxley ; CL, CW

Scalibregma inflatum Rathke ; PV, SP, CL, CW
Schistocomus hiltoni Chamberlin; PV, SP, NE
Sclerocheilus acirrata n. sp.; CL

Scoloplos acmeceps Chamberlin ; SP, CW

408 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Scoloplos armiger (Miiller) ; SP, CL

Scoloplos (Leodamas) sp.; CL

Scoloplos sp.; SP, CL, CW

Sphaerodoridium ?biserialis (Berkeley and Berkeley) ; SP
Sphaerodoridium minutum (Webster and Benedict) ; PV, SP, CL
Sphaerodoridium sp.; SP

Sphaerodorum papillifer Moore; SP, CL

Sphaerodorum sp.; PV, SP, CW

Sphaerosyllis californiensis n. sp.; PV

Sphaerosyllis sp.; PV, SP, LS, CL, CW

Spio punctata Hartman; PV, SP, NE, LS, CL, CW
Spiophanes anoculata Hartman; SP

Spiophanes bombyx (Claparede) ; PV, SP, CW
Spiophanes fimbriata Moore; PV, SP, CL, CW
Spiophanes missionensis Hartman; PV, SP, NE, CL
Spiophanes spp.; PV, SP, CL, CW

Spirobranchus spinosus Moore; CL

spirorbids ; PV, CL, CW

Sternaspis fossor Stimpson; PV, SP, NE, CL, CW
Sthenelais fusca Johnson; CL

Sthenelais tertiaglabra Moore; PV, SP, NE, CL, CW
Sthenelais verruculosa Johnson ; CL, CW

Sthenelais sp.; SP, CL

Sthenelanella uniformis Moore; PV, SP, NE, CL, CW
Streblosoma crassibranchia Treadwell; PV, SP, CL, CW
Streblosoma sp.; SP, CL

Syllis gracilis Grube ; PV

Syllis sp:; SP, CL

Telepsavus costarum Claparéde; LH, PV, SP, NE, CL, CW
Terebellides stroemti Sars; PV, SP, NE, LS, CL, CW
Thalenessa spinosa (Hartman) ; PV, SP, NE, LS, CL, CW
Tharyx marioni (Saint-Joseph) ; PV, CW

Tharyx monilaris Hartman; PV, SP, CL

T haryx multifilis Moore; PV, SP, NE, CW

T haryx tesselata Hartman; LH, PV, SP, LS, CL, CW
Tharyx spp.; PV, SP, NE, LS, CL, CW

NO. 2 HARTMAN : QUANTITATIVE SURVEY

T helepus setosus (Quatrefages) ; SP, CL
Thelepus sp.; CL

Thormora johnstoni (Kinberg) ; PV
Timarete sp. ; CL

Travisia brevis Moore; SP, NE

Travisia gigas Hartman; PV, SP, CW
Travisia pupa Moore; PV, SP, NE, CL
Travisia spp.; LH, CL, CW
Trypanosyllis gemmipara Johnson; PV

Trypanosyllis sp. ; CL

Typosyllis aciculata Treadwell ; CL
Typosyllis spp.; PV, SP, LS, CL

Vermiliopsis biformis Hartman; SP, CL, CW
Vermiliopsis cornuta Rioja; CL

Vermiliopsis infundibulum Philippi; CL
Vermiliopsis sp.; SP, CL, CW

O

oO

ECHINODERMS

(E=echinoid, Oophiuroid, H=holothuroid, A—asteroid, C=crinoid)

Allocentrotus fragilis (Jackson) ; SP, LS, CL

A mphiacantha amphacantha (McClendon) ; PV, SP, NE, LS,
Ch, CW

A mphiodia (A mphispina) digitata Nielsen; LH, PV, SP, NE,
Ch.-CWw.

Amphiodia (Amphispina) urtica (Lyman) ; PV, SP, NE, LS,
CEeCw.

Amphiodia occidentalis Lyman; LH, SP, CL

Amphiodia psara H. L. Clark; SP, NE

A mphiodia, rugose ; SP

A mphioplus hexacanthus H. L. Clark; SP, CL, CW

Amphioplus strongyloplax (H.L. Clark) ; PV, SP, NE, LS,
CE-CW.

Amphipholis pugetana (Lyman) ; PV, SP, LS, CL, CW

409

Amphipholis squamata (delle Chiaje) ; PV, SP, NE, LS, CL, CW

Amphiura arcystata H. L. Clark; PV, SP, NE, CL, CW
Amphiura seminuda Liitken and Mortensen; SP, LS
Asterina miniata (Brandt) ; PV, CW

410 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Astropecten californicus Fisher ; PV, SP, NE, CL, CW
Astropecten sp.; NE, CL

Astrophiura marionae Ziesenhenne ; LS
Benthopecten acanthonotus Fisher ; SP
Brisaster townsendi (A. Agassiz) ; PV, SP, CL
Brissopsis pacifica (A. Agassiz) ; PV, SP, LS, CL, CW
?Caudina sp.; CL

Centrostephanus coronatus (Verrill) ; CL
Cryptopeltaster lepidonotus Fisher ; SP j
Cucumaria sp. ; SP, CL

Dendraster excentricus (Eschscholtz) ; LH, PV, SP, NE
Dendraster laevis H. L. Clark; SP

Dendraster sp.; SP_

Florometra perplexa (H. L. Clark) ; SP, CL

Gonimaretia laevis H. L. Clark; SP

Gorgonocephalus caryi (Lyman) ; SP

Henricia sp.; SP, CL, CW

Leptosynapta albicans (Selenka) ; PV, SP, NE, LS, CL, CW
Leptosynapta sp.; SP, CW

Leptychaster (Parastropecten) pacificus Fisher ; SP

Lovenia cordiformis A. Agassiz ; SP, CL, CW

Luidia asthenosoma Fisher ; CL

Luidia foliolata Grube ; SP, CL, CW

Euidia ludwigi Fisher ; PV, CL

Luidiaster californicus Ziesenhenne; SP, LS

Lytechinus anamesus H. L. Clark; PV, SP, NE, CL, CW
Mediaster aequalis (Stimpson) ; CL

Molpadia intermedia (Ludwig) ; PV, SP, NE, CL, CW
Molpadia sp.; CL

Myxoderma platyacantha H. L. Clark; SP

Odontaster crassus Fisher ; SP

?0 phiacantha bairdi Lyman; LS

O phiacantha diplasia H. L. Clark; SP, LS, CL, CW

O phiacantha paucispina Liitken and Mortensen ; SP

ee ae

O phiacantha phragma Ziesenhenne ; SP
O phiacantha rhachophora H. L. Clark; SP

OOO OsS> EE Se eb fe eee hea te eee ae Soe eS ees

NO. 2 HARTMAN : QUANTITATIVE SURVEY 411

OQ Ophiacantha sp.; SP

O Ophiocynodus corynetes H. L. Clark; SP, LS, CL, CW
O Ophioderma sp.; SP

O Ophiomusium jolliensis McClendon ; SP, CW

OQ Ophionereis eurybrachyplax H. L. Clark; CL

O Ophiopholis bakeri McClendon; SP, LS, CL

O Ophiopholis longispina H. L. Clark ; CW

O Ophiopsila californica A. H. Clark; CL, CW

OQ Ophiopteris papillosa (Lyman) ; CL, CW

O Ophiothrix spiculata Le Conte; PV, SP, NE, LS, CL, CW

O Ophiura kofoidi McClendon; SP

O Ophiura leptoctenia H. L. Clark; SP, CL

O Ophiura lutkeni (Lyman) ; PV, SP, NE, CL, CW

O Ophiuroconis bispinosa Ziesenhenne ; PV, SP

H Pachythyone rubra (H. L. Clark) ; LH, SP, CL

H_ Parastichopus californicus (Stimpson) ; CL

H_ Pentamera pseudopopulifera Deichmann; LH, PV, SP, NE,
CL, CW

A Pisaster capitatus (Stimpson) ; PV

H_ Psolus sp.; SP

A Rathbunaster californicus Fisher ; SP

A Sclerasterias heteropaes Fisher ; PV, SP, CL

E Spatangus californicus H. L. Clark; LS, CL

H_ Stichopus sp.; CL

E Strongylocentrotus franciscanus (A. Agassiz) ; SP

E Strongylocentrotus purpuratus (Stimpson) ; PV, SP

E Strongylocentrotus sp.; SP

A Stylasterias forreri (de Loriol) ; SP

MOLLUSKS

(P==pelecypod, G—gastropod, S=scaphopod, SO==solenogaster, A—amphineuran,
C=cephalopod, V=vermetid. Including those named in the brachiopod associ-
ation from Mattox, 1955:81).

P Acila castrensis (Hinds) ; SP, CL, CW

G_ Acteocina culcitella (Gould) ; SP

G_ Acteocina culcitella intermedia Willett ; SP, NE, CL
G Acteon punctocoelata (Carpenter) ; LH, PV, SP, CL

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

aS
aaR
Do

Admete californica Dall; SP

Admete sp.; SP

A dontorhina cyclia Berry; LH, PV, SP, NE, CL
A equipecten circularis aequiculcatus (Carpenter) ; SP
A glaja purpurea Bergh; PV

A glaja sp.7 UA, PV, SE, Cl Cw

Aligena sp.; SP, CL

Amphissa bicolor Dall; SP, CL

A mphissa cf. reticulata Dall; PV

Amphissa undata Carpenter; PV, SP

Amphissa sp.; PV

Amygdalum pallidulum (Dall) ; PV, SP, CL, CW
Anisodoris nobilis (MacFarland) ; CL
Antiplanes ?perversa (Gabb) ; SP, CL
Antiplanes sp. ; CW

Armina californica (Bergh) ; CL

Asthenothaerus villosior Carpenter ; LH, PV
Axinopsida serricata (Carpenter) ; PV, SP, NE, CL
Balcis catalinensis (Bartsch) ; CL

Balcis ?compacta (Carpenter) ; SP

Balcis rutila (Carpenter) ; LH, SP, CL

Balcis sp.; PV, SP, NE

Bittium catalinensis Bartsch; SP, CL

Bittium ?larum Bartsch; SP

Bittium subplanatum Bartsch; SP

Boreotrophon triangulatus (Carpenter) ; CL
Borsonella dalli Arnold ; SP

Botulina denticulata (Dall) ; CL

Burchia redondoensis (Burch) ; PV

Bursa californica (Hinds) ; SP, CL

Cadulus fusiformis Pilsbry and Sharp; LH, CL
Cadulus tolmiei Dall; SP, LS, CL

Cadulus spp.; LH, PV, SP, NE, CL, CW
Calliostoma tricolor Gabb; PV, CL

Calliostoma sp.; PV, CW

Cancellaria cooperi (Gabb) ; CL

GO) Oo om min @ so GGG GG Gr Gir Gr @r res GaGa Gr Gr =o anGe Gn ann ec Gul Gn sien an ian

NO. 2 HARTMAN: QUANTITATIVE SURVEY

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Cancellaria crawfordiana Dall; CL
Cancellaria sp.; PV

Capulus californicus Dall; SP

Cardiomya pectinata (Carpenter) ; SP, CL
Cardiomya sp.; PV

Carolina tridentata Forskal; CL

Cardita ventricosa Gould ; PV, SP, NE, CL, CW
Cardita spp.; SP, LS, CL

Chione undatella Sowerby ; LH

?Chione sp.; PV

Chlamys hastatus Sowerby ; CL

Chlamys hericius (Gould) ; CL

Chlamys latiauratus Conrad ; SP

Chlamys sp.; LH, PV, SP, CL
Clinocardium nuttalli (Conrad) ; CL

Compsomyax subdiaphana (Carpenter) ; LH, PV, SP, NE,

CL, CW
Conus californicus Hinds; PV, SP, CL, CW
Crenella columbiana Dall; NE
Crenella decussata Montagu; SP, CL
Crenella divaricata d’Orbigny ; SP
Crenella sp.; SP
Crepidula aculeata (Gmelin) ; LH
Crepidula excavata Broderip ; NE
Crepidula nivea Adams; PV, CL
Crepidula norissiarum Williamson ; SP
Crepidula onyx Sowerby ; PV
Crepidula spp.; LH, PV, SP, CW
Crepipatella lingulata (Gould) ; PV
Crystallophrisson hancocki Schwabl; SP
Crystallophrisson hartmani Schwabl ; SP
Crystallophrisson marinellii Schwabl ; SP
Crystallophrisson rectum Schwabl; SP
Crystallophrisson spp.; PV, SP
Cuspidaria apodema Dall; SP, CL
Cuspidaria sp.; NE, CL, CW

413

414 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

P Cyathodonta pedroana Dall; PV

P Cyclopecten vancouverensis (Whiteaves) ; PV, CL
Cylichnella attonsa (Carpenter) ; LH, PV, SP, NE
Cylichnella diegensis (Dall) ; LH, PV, SP, NE, CL
Cyrilla minuta Dall; SP, CL

Dacrydium pacificum Dall; SP, CL

Dacrydium sp.; LS

Dendropoma lituella (Morch) ; CL

Dentalium neohexagonum Pilsbury and Sharp; SP
Dentalium rectius Carpenter; PV, SP, CL
Dentalium sp.; LH, CL

Dermatomya tenuiconcha (Dall) ; CL

Diadora aspera Eschscholtz ; PV

Donax gouldi Dall; SP

Elaeocyma empyrosia (Dall) ; SP, CL

Elaeocyma halocydne (Dall) ; CL

Ensis myrae Berry ; SP

a
yd

Epitonium bellastriatum Carpenter ; SP
Epitonium tinctum (Carpenter) ; NE, CL
Epitonium spp.; PV, SP, NE, CL, CW
Erycina chacei Dall; SP

Eulima californica Bartsch; SP, CL
Eulima rutila (Carpenter) ; SP

Eulima sp.; CL

G_ ?Fusinus sp.; PV

P Gari edentula (Gabb) ; SP

G_ Gastropteron sp.; CL

G_ Glans carpenteri (Lamy) ; PV

P Glycimeris subobsoleta Carpenter ; LS, CL
x Faminoea virescens (Sowerby) ; CL

G Hemitoma bella (Gabb) ; CL

P Hinnites sp.; CW

G_ Hyalima sp.; CW

G Kellettia kelleti Forbes; CL

P Kellia laperousti Deshayes ; PV

P Kellia suborbicularis (Montagu) ; CL

@ GQ rag Gy) Gc: Gr Gy tc) Glee cacao ay ee

Q

NO. 2 HARTMAN : QUANTITATIVE SURVEY 415

e Kellia'sp:; UH, PV,SP,CW

G_ ?Lacuna sp.; NE

P Ledasp:; PV

A Lepidopleurus nexus Carpenter; LS

A Lepidozona catalinae Willett ; CL

P_ Lima dehiscens Conrad ; PV, SP, CL, CW
P Lima subauriculata Montagu; PV, CL, CW
SO Limifossor fratula Health; SP

ie)

O Limifossor sp.; LS

Limopsis diegensis Dall; CL

Linga lichthofeni (Gabb) ; SP

Linga sp.; SP

Lithophaga plumula kelseyi Hertlein and Strong; PV
Lithophaga sp. ; CL

Lucinisca nuttalli (Conrad) ; LH, PV, SP
Lucinoma annulata Reeve; LH, PV, SP, CW
Lyonsia californica Conrad; LH, PV, NE, CL
Lyonsia sp.; SP

Macoma indentata Carpenter ; LH

Macoma nasuta Conrad ; LH, SP

Macoma yoldiformis Carpenter; LH, PV, SP, NE
Macoma sp.; LH, PV, SP, NE

Mactra ?californica Conrad ; LH

Mangelia arteaga Dall and Bartsch; SP
Mangelia barbarensis Oldroyd ; LH, SP
Mangelia spp.; LH, PV, SP, NE

?Margarites sp.; SP

Megasurcula carpenteriana (Gabb) ; NE, CL
Melanella sp.; CL

Micranellum crebricinctum (Carpenter) ; CL
G Micranellum sp.; SP

G Miitrellasp.; SP

P Modiolus capax (Conrad) ; CL

P Modiolus modiolus Linnaeus ; PV, SP

P Modiolus neglectus Soot-Ryen; LH, SP

P Modiolus sacculifer (Berry) ; PV, CL

ag iqe} ge} !as) ae) las) Iq) Ine} toe} Ine} Ine} tae) Ie] Ise,

Sa’
he 4

OO) G) G2 Gi 1G

+16

GED re rs es re rs ret = GaG) GaGa Gy eG lean c he ga) Gal Ga Ga Ga Gy) bl

an
1 wd

~

4

~

4

~

~

of Mopofl Doyafh Var

ALLAN HANCOCK PACIFIC EXPEDITIONS

Mytilus sp.; PV

Nassarius cooperi (Forbes) ; LH

Nassarius insculptus (Carpenter) ; CL

Nassarius mendicus (Gould) ; PV

Nassarius perpinguis (Hinds) ; LH, PV, SP, NE
Nassarius sp.; PV, CL, CW

?Natica clausa Broderip and Sowerby ; SP
Nemocardium centrifilosum (Carpenter) ; SP, CL
Nemocardium sp.; CL, CW

Neosimnia acicularis (Lamarck) ; PV

Neosimnia loebbeckeana (Weinkauft) ; CL
Neosimnia sp.; CL

Nitidella carinata (Hinds) ; PV, NE

Nitidella ?gouldi Carpenter ; SP

Nitidella permodesta (Dall) ; SP, LS, CL
Nitidella ?tuberosa (Carpenter) ; PV

?Nitidella sp.; PV

Nucula carlottensis Dall; PV, SP, CL

Nucula linki Dall; CL

Nucula tenuis Montagu; SP, NE

Nucula sp.; SP, CL

Nuculana conceptionis (Dall) ; SP, LS
Nuculana hamata (Carpenter) ; PV, SP, LS, CL
Nuculana minuta (Fabricius) ; SP

Nuculana taphria (Dall) ; LH, SP, NE, CL, CW
Octopus apollyon Berry ; CL

Odostomia herilda Dall and Bartsch; SP
Odostomia sp.; LH, PV, SP

Olivella baetica Carpenter; LH, PV, SP, NE
Olivella ?pedroana (Conrad) ; NE

O phiodermella incisa Carpenter ; LH, PV, SP
Pandora bilirata Conrad; PV, SP, NE, CL
Pandora sp.; NE

Parvilucina tenuisculpta (Carpenter) ; LH, PV, SP, LS, CL

Pecten (Pecten) diegensis Dall; CL
Pecten spp.; PV, SP, CW

VOL.

NO.

P

Cotto wewe we We ODODADAAaAAAs

2 HARTMAN : QUANTITATIVE SURVEY

Pedicularia californica Newcomb ; CW
Periploma discus Stearns ; LH, SP
Periploma planiuscula Sowerby ; PV, SP
Periploma sp.; SP

Philine sp.; CL

?Plagioctenium circularis (Sowerby) ; PV, SP
Pleurobranchaea sp.; CL

Pododesmus macroschismus Deshayes ; PV
Polinices cf. draconis Dall; PV

Polinices lewisti Gould ; SP, NE
Polinices sp.; PV, SP

Poromya sp.; LH, SP

Prochaetoderma californica Schwabl; SP, CL
Protothaca staminea (Conrad) ; PV
Protothaca tenerrima Carpenter ; LH
Psephidia lordi Baird ; SP

Psephidia sp. ; SP

Pseudamusium ?incongruus (Dall) ; SP
Pseudochama exogyra (Conrad) ; CL
Pseudopythina chacei (Dall) ;SP, NE
Pseudopythina sp.; PV

Pterynotus carpenteri Dall; CL
Pterynotus petri (Dall) ; CL
Pterynotus sp.; CL

Puncturella cucullata (Gould) ; CL
Puncturella galeata (Gould) ; CL
Pusula californica (Gray) ; CL

Retusa harpa Dall; SP

Rochefortia aleutica Dall; PV, SP, NE
Rochefortia cf. golischi Dall; SP
Rochefortia tumida Carpenter ; NE
Rochefortia spp.; LH, PV, SP, NE

Saxicava arctica (Linnaeus) ; PV, SP, LS, CL, CW

Saxicava cf. rugosa (Linnaeus) ; CL
Saxicavella pacifica Dall; PV, SP, NE, LS
Semele pulchra (Sowerby) ; CL

417

aS
—
ioe)

QQOQAQ tater tt Hee er OO Se ee Oo oO as

ALLAN HANCOCK PACIFIC EXPEDITIONS

Siliqua lucida Conrad ; SP, CW

Sinum debile Gould ; PV

Sinum scopulosum Conrad ; PV, CL
Solamen columbianum (Dall) ; LH, PV, SP, LS, CL
Solariella peramabilis Carpenter ; CL
Solariella sp.; SP

Solemya panamensis Dall; SP

Solemya sp.; PV

Solen rosaceus Carpenter ; LH, SP, CL
Solen sicarius Gould ; LH, SP, NE

Solen spp.; PV, SP, CW

Sphenia fragilis Carpenter ; CL

?Sphenia sp.; PV

Spisula planulata Conrad ; SP

Spisula sp.; SP

Sportella californica Dall; CL
Strombiformis californicus Bartsch ; SP
Sulcuretusa taphria (Dall) ; SP

Tagelus californicus Conrad ; LH, SP

?T apes sp.; PV

Tellina buttoni Dall; LH, SP, CL

Tellina carpenteri Dall; LH, PV, SP, LS, CL, CW
Tellina idae Dall; LH, SP, CW

Tellina modesta Carpenter ; PV, NE, CL
Tellina spp.; PV, SP, CL

Thracia curta Conrad ; PV

Thracia trapezoides Conrad ; SP

Thracia sp.; PV

T hyasira barbarensis Dall; LH, CL
Thyasira ?gouldi (Philippi) ; SP

T hyasira trisinuata Orbigny ; SP, PV, CL

T hyasira spp.; LH, PV, SP, NE, CL, CW
Trachycardium quadragenarium (Conrad) ; PV, SP, CL
Triophora sp.; CL

Tritoniopsis aurantia Mattox ; CL
Turbonilla antimunda Dall and Bartsch; SP

voL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY 419

G Turbonilla cf. attrita Dall and Bartsch; SP
G Turbonilla santarosana Dall and Bartsch; SP
G Turbonillaspp.; LH, PV, SP, NE, CL

G_ Turritella cooperi Carpenter ; CL
V_vermetid ; PV, SP, CW

P Verticordia ornata (d’Orbigny) ; CL

G Vitrinellasp.; LH, PV

G_ Volvulella cylindrica (Carpenter) ; SP

G Volvulella tenuissima Willett ; LH, PV, SP, NE, CL
G Volvulellasp.; PV, SP
G Zonariasp.; PV

SYS DEMATIC LISh OF BRYOZOA IDENTIFIED FROM
FARNSWORTH BANK, SANTA CATALINA ISLAND,
IN SAMPLES OF VELERO IV, STATIONS
NUMBERED 3594, 3595 AND 10334

by William Banta

The following bryozoan species have been identified in three sam-
ples taken by the VELERO IV, from Farnsworth Bank, near Santa
Catalina Island. Specific accounts of all species may be consulted in the
monograph by Osburn (1950-1953, Hancock Pac. Exped., vol. 14).
The relative abundance of species is indicated by symbols, as follows:
P = present; U = uncommon; O = occasional; C = common, and

A = abundant ; — = not recovered.
Station Numbers of
Name of Species Velero TV

Family HINKSINIDAE 3594 3595 10334
Antropora tincta ( Hastings 1930) — P C
Hinksina velata (Hinks 1881) ly — P
Cauloramphus spiniferum (Johnston 1832) O — Pe
Cauloramphus brunnea (Canu and Bassler

1930) — — C

Family ALDERINIDAE
Alderina smitti Osburn 1950 = = P
Mollia patellaria (Moll 1803) — — C
Callopora circumclathrata (Hinks 1881) P iP —
Callopora corniculifera (Hinks 1884) — — P
Callopora 2inconspicua (O’donoghue 1923) — — ee
Copidozoum tenuirostre (Hinks 1880) lf — U

420 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

Station Numbers of
Name of Species Velero IV

Family CHAPERIELLIDAE 3594 3595 10334
(= CHAPPERTIDAE)
Chaperiella (= Chapperia) californica

(Osburn 1950) — P Cc
Chaperiella (= Chapperia) patula (Hinks 1881) — — Pp
Family MICROPORIDAE
Micropora coriacea (Esper 1791) P. ¢ C
Family CELLARIIDAE
Cellaria mandibulata Hinks 1882 P = C
?Cellaria diffusa? Robertson 1905 — — \
Family SCRUPOCELLARIIDAE
A mastigia rudis ( Busk 1852) — — P

Caberia boryi (Audouin 1826)
Scrupocellaria varians (Hinks 1882)

Family BICELLARIELLIDAE
Dendrobeania longispinosa (Robertson 1905)

Family CRIBRILINIDAE
Reginella mucronata (Canu and Bassler 1923 ) =
Lyrula hippocrepis (Hinks 1882) P
Colletosia radiata (Moll 1803) P
Colletosia sp. —
Figularta hilli Osburn 1950 —

ala
|
|

|
|
ry

Ps ao) ao@
ae)

Division AACOPHORA

Family HIPPOTHOIDAE
Hippothoa distans (MacGillivary 1896) = P —
Celleporella (= Hippothoa) hyalina

(Linnaeus 1758) B P
Try postega claviculata (Hinks 1884) — P

Family SCHIZOPORELLIDAE
?Schizoporella linearis var. inarmata Hinks 1884 — —
Emballotheca obscura Osburn 1952 — —-
Schizomavella auriculata ( Hassall 1842)
Hippodiplosia insculpta (Hinks 1882)

Family HIPPOPORINIDAE
Hippoporina porcellina ( Busk 1860) le iP Ss
Hippoporella nr. gorgonensis Hastings 1930 -- — P

e) |

EJ 1@

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Name of Species

Stephanosula vitrea Osburn 1952
Hippomonavella longirostrata (Hinks 1883)
hippoporinid, unidentified

Family MICROPORELLIDAE
Microporella ciliata ( Pallas 1776)
Microporella vibraculifera (Hinks 1884)
Microporella cribosa Osburn 1952
Microporella californica (Busk 1856)
Fenestrulina malusi (Audouin 1826)

Family SMITTINIDAE
Porella porifera (Hinks 1884)
Porella patens Osburn 1952
Rhamphostomella curuirostrata (O’donoghue
1923;)
Smittina landsborovi ( Johnston 1847)
Parasmittina trispinosa ( Johnston 1838)
Escharella (= Mucronella) major
(Hinks 1884)
Family RETEPORIDAE
Phidolopora labiata Gabb and Horn 1862
[= P. pacifica (Robertson 1908) ]
Rhynchozoon rostratum (Busk 1856)
[incl. R. tumulosum |
Rhynchozoon grandicella Canu and Bassler 1923
Family CREPIDACANTHIDAE
Crepidacantha poissoni ( Audouin 1826)
Family PHY LACTELLIDAE
Lagenopora socialis (Hinks 1877)
Lagenopora spinulosa (Hinks 1883)
Lagenopora punctulata (Gabb and Horn 1862)
Family CELLEPORARIIDAE
(=CELLEPORIDAE)
Celleporaria (= Holoporella) brunnea
(Hinks 1884)
Celleporina (= Costazia) robertsonae
(Canu & Bassler 1923)
Celleporina (= Costazia) procumbens

(Osburn 1952)

421

Station Numbers of

Velero IV

3594 3595
— P
— P
P me
P 7
P P
O Cc
= Pp
iP le
A —_
iz P
Pp ==
P =
P ae
(Gs P

10334
P
P
P.

Se

7 © |

@

422 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Station Numbers of
Name of Species Velero IV

3594 13595 M0334
Family ONCOUSOECIIDAE

Proboscina ?sigmata Osburn 1953 — — Pp

Proboscina, two other species — — ie
Family DIASTOPORIDAE

Diaperoecia californica (d’ Orbigny 1852) (E — —
Family TUBULIPORIDAE

Tubulipora tuba (Gabb and Horn 1862) E P U

Tubulipora pacifica Robertson 1910 |e — —

Platonea ?expansa Osburn 1953 — — P
Family CRISIIDAE

Crisidia corunata (Linnaeus 1758) — — O

Crisia occidentalis ‘Vrask 1857 — — je
Family PENETRANTIIDAE

?Penetrantia concharum Silen 1946 — RP P

Serial numbers 11 to 267 with Velero Station numbers from each
station for which analyses are given

(See also vol. 19(1) :6 and 12-38, for map and additional data).

Serial area
number code Velero Station no.
1] PV 4854
12 PV 4855
14 PV 2788
19 EV 4832
27 Py. 2473 ; 4802 ; 4803 ; 4833
28 PV, 2472 ; 3050; 4801 ; 4805 ; 4806 ; 4807 ;
4830 ; 4856 ; 5096; 5101; 5102; 5541
29 jel 2307 ; 2508 ; 6107
30 LH 2507 ; 5808
32 eH 2314
33 iDjat +718
41 PV +831; 5029
42 PV 2417 ; 2962 ; 3049 ; 3051 ; 5027 ; 5502

43 PV 5028 ; 5030

NO. 2 HARTMAN : QUANTITATIVE SURVEY

Serial area
number code Velero Station no.
+4 PV 2470
45 SE 3047 ; 3048 ; 5753
46 SP 2107; 6100
47 bie 2006 ; 3053 ; 6102
48 SP 4719; 4886
49 SP 4885
50 SP 2741
60 PV 2430
62 SP 5754
63 SP 30525751
64 SP 5752 ; 5844
65 SP 6104
66 SP 2314
67 SP 2504; 5741
68 SP 5740
83 SP 5743
86 SP 5742
89 (SF 2798
98 SP 2306
99 SE 2355 ; 5749
100 SP 5750
101 SP 5748 ; 5820
103 SP 5745
104 Se 2292
105 Sie 2630
108 SP 5003 ; 5087
110 NE 2745
123 ile ANA6G S71;
129 SP 5086 ; 5746
134 NE 2747 ; 4877
135 CL 2131
136 CE 2738
145 SP 2836
154 SP 2884
160 CL 2736; 5146
161 Cl 2389
162 CL 3569; 3570; 5148
174 SP 2416; 2987
184 SE 4778

185 NE 4777 ; 4871 ; 4878 ; 5092 ; 5354

424 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

Serial area

number code Velero Station no.

186 CL 2142; 2955 ; 2957 ; 2959 ; 2960’; 2961;
3310

187 Cr 2302 ; 2733

189 Cr 2301

194 SP. 2802

197 SP 2898

199 SP 2372

200 SP 2895

201 SP 2886

207 GIL, 2452

208 CE 2451; 2952; 2953

210 CL 2365

233\)| SP 2393128371,

PAIS SP 2O29

Ziyi SP 2901

219 SP 2894

220 Es 2843

DA) ES 2887

222 SP 2644

224 CL 1370; 1371; 2128 ; 2144; 2152; 2450

225 CE 5727

226 CL 2426

227 SP 235232859

228 6M 2154; 2204; 2228

229 6M 2845

23il SIE 2900

232 SIP 2370

233 Siz 2844

234 1S 2299

235) ES 2298

236 Is) 2297

238 Cr 212152637 ; 2638; 2853'; 2855 = 3603-
3605

239 CE 2639 ; 3601

240 Ci 2367

241 SE 2350 ; 2640

242 SP 2368

243 SP 2369

244 poy es 2440

249 SP 2635

NO. 2

Serial
number

250
251
Zoe
253
259
260
262
263
267

HARTMAN : QUANTITATIVE SURVEY 425

area
code

CL
CL
CL
CE
CL
CL
SP
SP
SP

Velero Station no.

24302 S049 31/073 171957 123% Lie)
2347; 7709 ; 7726

2344

7720

2122; 3614 = 3613 ; 3615; 4045 ; 5095
2348 ; 3616; 3617; 7712

2342

2341

277 2 24382 7713

426 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

LITERATURE CrrEeD

BANSE, KARL

1963. Polychaetous annelids from Puget Sound and the San Juan Archi-
pelago, Washington. Biol. Soc. Wash., Proc., 76:197-208, figs. 1-3.

BARNARD, J. L.

1957. A new genus of phoxocephalid Amphipoda (Crustacea) from Africa,
India, and California. Ann. and Mag. Nat. Hist., (12) 10:432-438,
4 figs.

1958. Amphipod crustaceans as fouling organisms in Los Angeles-Long Beach
harbors, with reference to the influence of seawater turbidity. Calif.
Fish and Game, 44:161-170, 2 figs.

1959a. Liljeborgiid amphipods of southern California coastal bottoms, with a
revision of the family. Pacific Naturalist, 1 (4) :12-28, 12 figs.

1959b. The common pardaliscid Amphipoda of southern California, with a
revision of the family. Pacific Naturalist, 1 (12) :36-43, 4 figs.

1960. New Bathyal and Sublittoral Ampeliscid Amphipods from California,
with an illustrated key to Ampelisca. Pacific Naturalist, 1 (16) :1-36,
11 figs.

1961. Relationship of Californian Amphipod Faunas in Newport Bay and in
the Open Sea. Pacific Naturalist, 2 (4) :166-186, 2 figs.

1962a. Benthic marine Amphipoda of southern California: Families Aoridae,
Photidae, Ischyroceridae, Corophiidae, Podoceridae. Pacific Naturalist,
3 3-163, 78 figs.

1962b. Benthic marine Amphipoda of southern California: Family Oedicero-
tidae. Pacific Naturalist, 3 :349-371, 10 figs.

1963. Relationship of benthic Amphipoda to invertebrate communities of in-
shore sublittoral sands of southern California. Pacific Naturalist, 3:
437-467, 7 figs.

BARNARD, J. L. and R. R. GIvEN

1960. Common Pleustid Amphipods of southern California, with a projected
revision of the family. Pacific Naturalist, 1 (17) :37-48, 6 figs.

1961. Morphology and Ecology of Some Sublittoral Cumacean Crustacea of
Southern California. Pacific Naturalist, 2:153-165, 4 figs.

BARNARD, J. L. and OLGA HARTMAN

1959. The sea bottom off Santa Barbara, California: Biomass and commu-
nity structure. Pacific Naturalist, 1 (6) :1-16, 7 figs.

BARNARD, J. L. and F. C. ZIESENHENNE

1961. Ophiuroid Communities of Southern California Coastal Bottoms. Pa-
cific Naturalist, 2 (2) :131-152, 8 figs.

CALIFORNIA STATE WATER QUALITY CONTROL BOARD

1959. Oceanographic survey of the continental shelf area of southern Cali-
fornia. Its Publ., 20 :1-560.

1965a. An oceanographic and biological survey of the southern California
mainland shelf. Its Publ., 27 :1-232.

1965b. An oceanographic and biological survey of the southern California
mainland shelf. Appendix-Data. Its Publ., 27 (App.) : 1-445.

CHAMBERLIN, R. V.

1919. New Polychaetous Annelids From Laguna Beach, California. Journ.
Entom. Zool., 11:1-23.

No. 2 HARTMAN : QUANTITATIVE SURVEY 427

Emery, K. O.
1960. The Sea Off Southern California. Wiley, New York, 366 pp., figs.
1-248.

Emery, K. O. and Jossr HULSEMANN

1963. Submarine canyons of southern California. Part I. Topography, water,
and sediments. Allan Hancock Pac. Exped., 27 (1) :1-80, figs. 1-22.

FAUVEL, PIERRE
1927. Polychétes sédentaires. Faune de France, 16 :1-494.

FRASER, C. M.

1943. General account of the scientific work of the Velero III in the eastern
Pacific, 1931-41. Allan Hancock Pac. Exped., 1:1-431, pls. 1-128, charts
1-115.

GIvEN, R. R.
1961. The cumacean fauna of the southern California continental shelf. No.
1, Family Leuconidae. So. Calif. Acad. Sc., Bull., 60:130-146, pls. 34-37.

1964. The cumacean fauna of the southern California continental shelf. No.
2. The new family Mesolampropidae. Crustaceana, 7 :284-292, figs. 1, 2.

HARTMAN, OLGA

1955. Quantitative survey of the benthos of San Pedro Basin, southern Cali-
fornia. Part I. Preliminary Results. Allan Hancock Pac. Exped., 19
(1) :1-185, pls. 1-7.

1961a. A New Monstrillid Copepod parasitic in Capitellid Polychaetes in
Southern California. Zool. Anz., 167 :325-334, figs. 1-5.

1961b. Polychaetous annelids from California. Allan Hancock Pac. Exped.,
25 :1-226, pls. 1-34.

1963a. Submarine canyons of southern California. Part II. Biology. Allan
Hancock Pac. Exped., 27 (2) :1-424, figs. 1-27.

1965. Deep-water benthic polychaetous annelids off New England to Ber-
muda and other North Atlantic areas. Allan Hancock Found. Publ.,
Occ. Pap., 28 :1-378, pls. 1-52.

HARTMAN, O1cGa and J. L. BARNARD
1958. The benthic fauna of the deep basins off southern California. Pt. I.
Allan Hancock Pac. Exped., 22 (1) :1-67, pls. 1-2.

1960. The benthic fauna of the deep basins off southern California. Pt. II.
Allan Hancock Pac. Exped., 22 (2) :69-297, pls. 1-19.

HARTMANN-SCHRODER, GESA
1963. Revision der Gattung Mystides Théel (Phyllodocidae; Polychaeta Er-
rantia). Mit Bemerkungen zur Systematik der Gattungen Eteonides
Hartmann-Schroéder und Protomystides Czerniavsky und mit Beschrei-
bungen zweier neuer Arten aus dem Mittelmeer und einer neuen Art
aus Chile. Zool. Anz., 171:204-243, figs. 1-61.

JONES, G. F.
1963. The distribution and abundance of subtidal benthic Mollusca on the
mainland shelf of southern California. Malacologia, 2:43-68, 5 figs.

1965. The distribution and abundance of the pelecypod Adontorhina cyclia
on the mainland shelf of southern California. Int. Rev. Hydrobiol.,
50:127-138, 5 figs.

KEEN, A. Myra
1961. A proposed reclassification of the gastropod family Vermetidae. Brit.
Mus. (Nat. Hist.), Bull., Zool., 7:181-213, pls. 54-55.

428 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

Mattox, N. T.

1955a. Studies on the Opisthobranchiata: I. A new species of the genus Tri-
toniopsis from southern California. So. Calif. Acad. Sci., Bull., 54
(1) :8-13, pls. 4-5.

1955b. Observations on the brachiopod communities near Santa Catalina
Island. Jz Essays in the Natural Sciences in Honor of Captain Allan
Hancock. Los Angeles, Univ. of South. Calif. Press, pp. 73-86, 5 figs.

1958. Studies on the Opisthobranchiata: II. A new Tectibranch of the Genus
Philine. So. Calif. Acad. Sci., Bull., 57:98-104, pls. 33, 34.

Menzies, R. J. and J. L. BARNARD

1959. Marine Isopoda on coastal shelf bottoms of southern California: Syste-
matics and ecology. Pacific Naturalist, 1 (11) :1-35, figs. 1-28.

Moore, J. P.

1902. Descriptions of some new Polynoidae, with a list of other Polychaeta
from north Greenland waters. Acad. Nat. Sci. Philadelphia, Proc.,
54:258-278, 2 pls.

1923. The Polychaetous Annelids dredged by the U.S.S. “Albatross” off the
Coast of Southern California in 1904. IV. Spionidae to Sabellariidae.
Acad. Nat. Sci. Philadelphia, Proc., 75:179-259, 2 pls.

REIsH, D. J.
1955. The relation of polychaetous annelids to harbor pollution. U. S. Public
Health Sery., Public Health Rept., 70:1168-1174.

1960. A new species of Sabellidae (Annelida, Polychaeta) from southern
California, Ann. Mag. Nat. Hist., (13) 2:717-719, figs. 1-5.

1964. A quantitative study of the benthic polychaetous annelids of Catalina
Harbor, Santa Catalina Island, California. So. Calif. Acad. Sci., Bull.,
63 :86-92, 1 fig.

Rioja, ENRIQUE
1941. Estudios Anelidologicos. III. Datos para el conocimiento de la fauna

de poliquetos de las costas del Pacifico de Mexico. Mexico, Uniy. Nac.,
Inst. Biol. Anal., 12:669-746.

SCHWABL, MATHILDE
1961. Crystallophrisson (==Chaetoderma) hartmani, nov. spec., eine neue
Aplacophore aus dem Ostpazifik. Zool. Anz., 166:258-277, figs. 1-10.
1963. Solenogaster mollusks from Southern California. Pacific Science, 17:
261-281, 28 figs.

UsHAKOov, P. V.
1955. Mnogoshchetinkovye chervi dal’nevostochnykh morei SSSR (Poly-
chaeta). Akad. Nauk SSSR, Zool. Inst., Opred. po faune SSSR, 56:1-
445, 164 figs.
1965. Polychaeta of the Far Eastern Seas of the U.S.S.R., Jerusalem, Israel
Program for Scientific Translations, 419 pp. [English translation of
above title].

ZIESENHENNE, F. C.
1951. A new brittle star of the genus 4sfrophiura from southern California.
So. Calif. Acad. Sci., Bull. 50 (1) :25-33, figs. 1, 2.

PLATES

430) ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

LANES, il

Gattyana brunnea, new species (Sta. 2154).
1. Elytrum from seventh segment, in dorsal view, x 30.
2. Distal end of a notoseta from eleventh parapodium, in lateral
view, x 310.
3. Distal end of a neuroseta from eleventh parapodium, in lat-
eral view, x 310.

Harmothoe priops Hartman (Sta. 5402).
4. Prostomium and first few segments, in dorsal view, x 94.
5. Distal end of notoseta from a median parapodium, x 140.
6. Distal end of neuroseta from a median parapodium, x 140.

431

HARTMAN : QUANTITATIVE SURVEY

NO. 2

Qtiitlcercecs

psy
LLALALLL A,

CALLA

ed

432 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

RWASE 2

Eusyllis transecta, new species (Sta. 2006)

1. Anterior end in dorsal view, x 92.

2. Pharyngeal tooth, in lateral view, x 245.

3. Distal end of pharynx, x 490.

4. A median parapodium in anterior view, x 145.

5. Superiormost seta from an anterior parapodium, x 1042.
6

Short falcigerous seta, from anterior segment, in lateral view,
x 1042.

Short falcigerous seta, from posterior segment, in lateral view,
x 1042.
8. Aciculum with bulbous tip, x 1042.

“I

HARTMAN : QUANTITATIVE SURVEY 433

i)

NO.

434 ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

JILNIPIS, 3

Sphaerosyllis californiensis, new species (Sta. 5028)

1. Anterior end, in dorsal view, x 125.
Posterior end, in dorsal view, x 125.
A median parapodium, in anterior view, x 270.
Falcigerous seta with dentate cutting edge, x 1450.
Falcigerous seta with smooth cutting edge, x 1450.
Distal end of an inferior simple seta, x 1450.
Distal end of an aciculum, x 1450.

No. 2 HARTMAN : QUANTITATIVE SURVEY 435

436 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

PLATE 4

Questa caudicirra, new genus and species (Sta. 3595)

1. Entire body, in dorsal view, x 16.

2. Posterior end in three-fourths dorsal view, x 30.

3. Anterior end, through five setigerous segments, in dorsal view,
x 32:
A median parapodium, in posterior view, x 120.
Seta from a median parapodium, x 310.
Bidentate hook from a median parapodium, x 310.

No. 2 HARTMAN : QUANTITATIVE SURVEY 437

4

)

ALLAN HANCOCK PACIFIC EXPEDITIONS VoL. 19

IAN IDI, 5}

Sclerocheilus acirratus, new species (Sta. 1370)

1. Anterior end, in dorsal view, x 74.

2. Acicular hook from first parapodium, x 490.

3. Slenderer seta from first parapodium, x 490.

4. Fureate seta from a median parapodium, x 490.
Ss

Tip of furcate seta, showing teeth along tines, x 980.

Asclerocheilus californicus Hartman (Sta. 3037)
6. Prostomium and first few segments, in dorsal view, x 29.
7. A posterior parapodium, showing prolonged lobes, x 29.
8. Acicular hook from first segment, in lateral view, x 260.
9. Tip of furcate seta, showing teeth along tines, x 980.

=
)

HARTMAN : QUANTITATIVE SURVEY

2

NO.

>

ccm

TTT, MQziziiii_

ea
SSS

440

ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

PLATE 6

Euchone arenae, new species (Sta. 2788)

1. Ventral end of radiolar base, showing one of two simple radi-
oles, x 48.

Thoracic superior seta, in lateral view, x 588.
Thoracic inferior seta, in lateral view, x 588.

Ze

3.

4. Thoracic uncinus, in lateral view, x 588.

5. Abdominal uncinus, in lateral view, x 1440.
6.

Posterior anal groove, in ventral view, x 40.

Euchone limicola Reish (Alamitos Bay)
7. Thoracic seta, in lateral view, x 336.
8. Abdominal uncinus, in frontal view, x 336.
9. Abdominal uncinus, in three-fourths view, x 780.
10. Posterior anal groove, in ventral view, x 46.

NO. 2 HARTMAN : QUANTITATIVE SURVEY 441

i

<J

442 ALLAN HANCOCK PACIFIC EXPEDITIONS voL. 19

ILI, 7/

Spirobranchus spinosus Moore (Isthmus Cove)
1. Entire animal, in left lateral view, x 6.3.
2. Operculum, in dorsal view, x 12.6.
3. Thoracic uncinus, in lateral view, x 875.

443

QUANTITATIVE SURVEY

.
.

HARTMAN

the

WA (7
pine

444 ALLAN HANCOCK PACIFIC EXPEDITIONS VOL. 19

PIAS E 8

Top. Seabottom at Sta. 7707, off Abalone Point, in 22.5 fms, showing
projecting tubes of chaetopterids in a silty sediment.

Bottom. Largest individuals from the same sample, showing, from left
to right, two tubes of Chaetopterus variopedatus, one of Tel-
epsavus costarum, three Lytechinus anamesus, and two Conus
californicus (the scale at bottom, refers to these specimens).

NO. 2 HARTMAN : QUANTITATIVE SURVEY 445

0 " 1z rr] 4 iS 6 ‘7 8 ’
Wii ie
mils ae

446

Top.

Bottom.

ALLAN HANCOCK PACIFIC EXPEDITIONS

PAWNIDIS, &)

Seabottom at Sta. 7719, shelf off Abalone Point, in 23.5 fms,
showing a silty surface with projecting tubes of Chaetopterus
variopedatus, slenderer Telepsavus costarum, and Lytechinus
anamesus.

Seabottom at Sta. 7720, shelf off Abalone Point, in 23.5 fms,
showing a silty bottom, similar to that at Sta. 7719, but with a
seastar, Mediaster aequallis.

voL. 19

NO. 2 HARTMAN ! QUANTITATIVE SURVEY 447

448

Top.

Bottom.

ALLAN HANCOCK PACIFIC EXPEDITIONS

ALINE; ill

Seabottom at Sta. 7723, shelf off Jewfish Point, in 39.5 fms,
showing a level silty bottom, with projecting rocks at lower
left, solitary corals at upper left, and a dead shell fragment
near center. The sampler failed to take a sample.

Seabottom at Sta. 7725, shelf off Jewfish Point, in 40.5 fms,
showing a silty surface with many small pits, mounds and
projecting ophiuroid arms.

voL. 19

NO.

bo

HARTMAN : QUANTITATIVE SURVEY

449

450

Top.

Bottom.

ALLAN HANCOCK PACIFIC EXPEDITIONS

IG ANIEIS, al

Seabottom at Sta. 7726, shelf off Jewfish Point, in 40 fms,
showing a silty surface overlain with dead shells of the
brachiopod, Laqueus californicus.

Largest specimens from Sta. 7726, showing anterior end of
Euclymene sp. with tube, and four dead valves of Laqueus
californicus. The scale at bottom refers to these individuals.

voL. 19

NO. 2 HARTMAN : QUANTITATIVE SURVEY 451

TTT Ty MARUIASUI LUTTE UL

16 17 18 19 20 21 22 23 24 25 26 27 2

4s
Cn
bo

Top.

Bottom.

ALLAN HANCOCK PACIFIC EXPEDITIONS

AWAITS, WZ

Seabottom at Sta. 7712, basin slope southeast end of Santa
Catalina Island, in 133 fms, coarse black sandy sediments,
showing patch of ophiuroid arms at far right, and crescentic
depressions formed by brissopsid urchins; the large trail across
lower left has not been identified.

The larger organisms from Sta. 7712, including Brissopsts
pacifica, and tubes of polychaetes; the scale at bottom refers
to these organisms.

voL. 19

iS

16

14M

17

no
rx)

{
\
es

pio

HE ii

2? 28

454

Top.

Bottom.

ALLAN HANCOCK PACIFIC EXPEDITIONS

JILANIEIS, i}

Sta. 7727, seabottom on the shelf off Jewfish Point, in 44 fms,
at which no sample was taken. The photograph shows the sur-
face riddled by pores and depressions, indicating the presence
of many animals in the sediments.

Sta. 7713, seabottom on basin slope, southeast of Santa Catalina
Island, in 286 fms. Surface sediments are black sand and
gravel with silt, and show pores, mounds and shallow trails
with white shelly fragments.

VoL. 19

HARTMAN : QU

ANTITATIVE SURVEY

INDEX OF SCIENTIFIC NAMES IN
SYSTEMATICS SECTION

(Numbers in bold face refer to illustrations)

acirratus, Sclerocheilus, 199, 439
arenae, Euchone, 202, 441
Asclerocheilus, 200

californicus, 200, 439

spe Ol
brunnea, Gattyana, 192, 431
californicus, Asclerocheilus, 200, 439
californiensis, Sphaerosyllis, 196, 435
caudicirra, Questa, 197, 437
ciliata, Gattyana, 193
Cirratulidae, 198
Cirratulus, 198

sp., 198
coineaui difficilis, Eteonides, 194

Hesionura, 194

difficilis, Eteonides coineaui, 194

Hesionura coineaui, 194
Eteonides coineaui difficilis, 194
Euchone, 202

arenae, 202, 441

incolor, 203

limicola, 203, 441
Eusyllis, 195

transecta, 195, 433
Gattyana, 192

brunnea, 192, 431

ciliata, 193
Halosydna, 193

latior, 193
Harmothoe, 193

priops, 193, 431

Hesionidae, 194

Hesionura, 194
coineaui difficilis, 194

incolor, Euchone, 203
latior, Halosydna, 193
limicola, Euchone, 203, 441
minutus, Sclerocheilus, 200
Paraonidae, 198
Polynoidae, 192

priops, Harmothoe, 193, 431

Questa, 197
caudicirra, 197, 437

Questidae, 197
Sabellidae, 202
Scalibregmidae, 199

Sclerocheilus, 199
acirratus, 199, 439
minutus, 200

Serpulidae, 204

Sphaerosyllis, 196
californiensis, 196, 435

spinosus, Spirabranchus, 204
Spirobranchus, 204, 443

Spirabranchus spinosus, 204

Spirobranchus, 204
spinosus, 204, 443

Syllidae, 195
transecta, Eusyllis, 195, 433

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