FORSCHUNGS museum KOENIG Bonn zoological Bulletin Supplementum Vol. 61 (2014) formerly: Bonner zoologische Monographien A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae) Silvia Fabrizi & Dirk Ahrens An open access journal of organismal zoology, published by Zoologisches Forschungsmuseum Alexander Koenig - Leibniz-lnstitut fur Biodiversitat der Tiere, Bonn BONN ZOOLOGICAL BULLETIN - SUPPLEMENTUM (BzB-S), formerly “Bonner zoologische Monographien”, ss as above, tel. +49 228-9122-425, ;e 1971. It is published by the Zoological Research ;inal zoology papers too long for inclusion in our in rs of this issue: Silvia Fabrizi & D :r. 129, 53113 Bonn, Gen nt of the map of the e> I bzm_61_ahrens_BZB-eml 22.04.14 12:41 Seite 1 A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae) Silvia Fabrizi & Dirk Ahrens Bonn zoological Bulletin - Supplen i61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 2 _ahrens_ Contents Volume 61 Abstract Key words Introduction Material and methods Depositories Systematic part Key to genera of Sri Lanka Periserica Brenske, 1898 Selaserica Brenske, 1 897 Neoserica (s.l.) Brenske, 1894 " Maladera" Maladera lugubris group Maladera (sbg. Cephaloserica) Maladera fistulosa group Maladera madurensis group Maladera setosa group Maladera detersa group Maladera (sbg. Hemiserica) “Serica" Species incertae sedis Zoogeography Acknowledgements References Tables Appendix. Gazetter of localities Index 13 34 39 39 47 58 90 91 91 92 97 97 98 100 101 122 124 Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK April 2014 A monograph of the Sericini of Sri Lanka (Coleoptera: Scarabaeidae) INTRODUCTION sive study of this fauna was conducted by Walker (1859) eration of the island followed later (Moser 1916; Frey 1972, 1974). l61: 1-124 ©ZFMK l61: 1-124 ©ZFMK l61: 1-124 (Figs 2A-C, 16A) spec ” (BMNH). Description. Length: 7.2 mm, length of elytra: 4.3 mm, iiiililiiiiiil 12 serrated line; posterior ventral margin almost straight, weakly widened in apical half, neither serrated ventrally nor dorsally, glabrous. Metatibia moderately wide and short, widest at middle, weakly narrowed apically, ratio width/ length: 1/ 3.6, longitudinally convex dorsally, with two groups of spines, basal one at one quarter, apical one at three quarters of metatibial length; lateral face longi- tudinally convex, smooth, laterally with very sparse and fine punctures; ventral margin with four strong equidis- tant spines; medial face smooth, apex interiorly near tarsal articulation moderately truncated. Tarsomeres impunctate dorsally, with dense, fine setae ventrally; metatarsomeres glabrous ventrally, with a strongly serrated carina, subven- trally with a second, smooth longitudinal carina, first metatarsomere a little shorter than following two tar- someres combined and one third of its length longer than the dorsal tibial spur. Protibia moderately long, tridentate, basal tooth small. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. IF-H. Habitus: Fig. 14C. Variation. Female: Antennal club composed of three an- tennomeres, slightly shorter than remaining antennomeres combined. Remarks. The abdomen and genitalia of the holotype of Periserica nigripennis are lost. However, it is virtually identically in all other external features with the lectotype of Periserica fulvostriata except for the colour of the ely- tra. Although the elytra of P. nigripennis are darker, the pattern of spots is recognizable and virtually identical to that of P. fulvostriata. Selaserica Brenske, 1897 Selaserica Brenske, 1 897: 355 (type species by monotypy: Selaserica nitida (Candeze, 1861)). Key to species of Selaserica (SS): 1 Hypomeron without a transverse carina basiventrally. Metafemur without a serrated line adjacent to anteri- or margin. Antenna with nine antennomeres {Selaserica nitida group) 2 r Hypomeron with a transverse carina basiventrally. Metafemur with a serrated line adjacent to anterior margin. Antenna with ten antennomeres {Selaserica splendiflca group) 13 2 Antennal club in male with 4 antennomeres 3 2’ Antennal club in male with 3 antennomeres 4 3 Antennal club almost as long as remaining anten- nomeres combined Se. nitida (Candeze) 3’ Antennal club distinctly longer than remaining anten- nomeres combined Se. knucklensis sp. n. 4 Frons at least in anterior two thirds shiny 5 4’ Frons extensively dull, shiny only narrowly behind frontoclypeal suture 7 5 Pronotum shiny. Large species, body larger than 11 mm 6 5’ Pronotum dull. Small species, body smaller than 6.5 mm. Frons completely shiny. Elytra glabrous, except the lateral setae Se. kanneliyana sp. n. 6 Dorsal surface dark or dark reddish brown. Elytra dull, intervals deeply impressed. Metatibia wide and strong- ly flattened, dorsal edge sharp. ...Se. lucidicollis sp. n. 6’ Dorsal surface light reddish brovm. Elytra complete- ly shiny, intervals weakly impressed. Metatibia slen- der and moderately flattened, dorsal edge almost lon- gitudinally convex Se. scutellaris Avcosn ($ !) 7 Metatibia wide and strongly flattened, dorsal margin sharp 8 7’ Metatibia slender and moderately flattened, dorsal margin almost longitudinally convex 12 8 Apex of phallobasis (in dorsal view) strongly asym- metrical Se. maculicauda (Arrow) 8’ Apex of phallobasis (in dorsal view) almost symmet- rical 9 9 Antennal club 1.5 times as long as remaining anten- nomeres combined Se. implicata (Brenske) 9’ Antennal club as long as or slightly longer than re- maining antennomeres combined 10 10 Basal lobe of left paramere narrow and long Se. padukkana sp. n. 10’ Basal lobe of left paramere large and lobiform, indis- tinctly defined 11 1 1 Eeft paramere evenly pointed at apex. Right paramere, relative to overall size of aedeagus shorter Se. impexa sp. n. 1 1 ’ Eeft paramere bluntly truncate at apex. Right para- mere, relative to overall size of aedeagus longer Se. convexiuscula sp. n. 12 Antennal club as long as remaining antennomeres combined Se. confirmata (Walker) 12’ Antennal club more than twice as long as remaining antennomeres combined. ..Se. distincticomis (Brenske) 13 Antennal club composed of three antennomeres. Small species, body smaller than 5.5 mm. Frons pos- teriorly narrowly dull 14 13’ Antennal club composed of four antennomeres. Larg- er species, body larger than 6 mm 15 14 Dorsal surface nearly glabrous. Se. wilpattuensis sp. n. 14’ Dorsal surface densely setose. ..Se. meridionalis sp. n. 15 Metatibia laterally with a very short serrated line on- ly. Body larger than 9 mm 16 15’ Metatibia laterally with a serrated line as long as one third of metatibia... 17 16 Dorsal surface in parts dull Se. ser icea Axxosn 16’ Dorsal surface completely shiny Se. nuwarana sp. n. illHlill i im i «i 1 1 1 IIP 1 1 pi I i !i I illl 1 1 nil 1 i i i II 1 1 HI If Hi i | i l!liiPiif!i|liiil!iii!iliifilW lii I 18 i61: 1-124 ©ZFMK ! iiti! ii ifiiiPiflP fiilK 20 22 24 tioii. Length: 9.6 mm, length of elytra: 7.3 mm, l61: 1-124 28 30 l61: 1-124 ©ZFMK I 61: 1-124 ©ZFMK 34 ©ZFMK 1 ’ Body size smaller, total length <5.4 mm. Body surface yellowish brown and shiny N. kalaarensis sp. n. 2 Antennal club in both sexes with four antennomeres.. N. bombycina (Karsch) 2’ Antennal club in male with six antennomeres N. sexfoliata Moser Remarks. Neoserica resulted polyphyletic in a first mo- lecular phylogeny of Sericini (Ahrens & Vogler 2008), however, a revision of the genus concept based on a com- bined molecular and morphological analysis needs a sig- nificantly wider and possibly complete taxon sampling. Therefore, following Ahrens (2004), we consider all species that were so far assigned to Neoserica and that do not belong to the monophyletic Neoserica (sensu stricto) group (Ahrens 2003b) preliminary as Neoserica sensu la- to. Neoserica bombycina (Karsch, 1882) (Figs 5M-0, 141, 17A) Serica bombycina Karsch, 1882: 387. Neoserica bombycina. Brenske 1898: 281; Arrow: 1916: 431. Serica quadriflabellata Brenske, 1896: 154, syn. n. Neoserica quadriflabellata. Brenske 1898: 253. Neoserica sericata Brenske, 1898: 280, syn. n. Type material examined. Syntypes {bombycina)-. 1 $ “61918/ ^/bombycina Karsch* Ceylon. Mus. Colombo” (ZMHB), 1 $ [without labels but the same type of pin - therefore it is to assume that thios specimen is a syntype as well. Consequently, the authors have added a copy of the labels of the other syntype:] “61918/ $/ A. bombyci- na Ceylon, Mus. Colombo” (ZMHB). Syntypes (quadri- flabellata)-. 3 1 $ “Konbir/ 4-jlabellata type Brsk.” (ISNB), 1 $ “Konbir [subsequently labelled by corator]/ 4-jlabellata type Brsk.” (ISNB), 1 6' “Konbir/ A-flabel- lata! coll. Brenske” (ZMHB), 1 6' “Konbir/ Coll. Brenske/4-flabellata” (ZMHB). Syntype {sericataf. 1 $ “Madras/ sericata type Brsk./ Type/ Museum Paris Coll. Generale” (MNHN), 1 $ “Madras” (MNHN). Additional material examined. S. India: 1 ex. ($) “Coll. R.I.Sc.N.B. Inde Barway P. Cardon/ Type/ det. Brenske 1898 Berk Ent. Zeit: 252/ Serica 191. Brsk.” (ISNB), 3 ex. “India: Pondicherry Karalkal, IX. 1988 T.R.S. Nathan” (CMNC), 1 6', 1 9 “Madras” (offensichtlich keine Typen von quadroflabellata, da das Mannchen Brenske unbekan- nt blieb, ZMHB), 1 $ “Mysore S. Indien III.53” (CF), 1 $ “Bengal Mandar/ etwas abweichend [hand written Brsk.]/ coll. Brenske” (ZMHB), 1 ex. “India: Anamalai Hills, Cinchona 700‘ [m] X.1959 / P. S. Nathan Bishop Museum” (BPBM), 1 ex. “S. India: Madras State, Ana- malai Hills, Cinchona 1050 m 1.67/ Collector P. S. Nathan” (BPBM), 2 ex. “South India Pondicherry State, Karikal 1. 1963/ Collector P. S. Nathan” (BPBM), 1 ex. “India (S): Kerala State: Walayar Forests, 213 m, X.1959/ P. S. Nathan Bishop Museum” (BPBM), 1 ex. “Ramnad Hin- doustan” (MNHN), 1 ex. “India Kistna Dt. Avanigadda VII.53 Rimondi” (MSNM), 1 ex. “Cote de Malabar M. Maindron/ Mahe Chass. inidgenes Aout 1901/ Museum Paris Cote de Malabar Mahe M. Maindron” (MNHN), 1 ex. “S. India Coimbatore PS. Nathan” (MNHN), 1 ex. “In- dia Coc[r?]onada/ Simson/Fry Coll. 1905.100.” (BMNH), 1 ex. “W.S. Millard 3rd Nov. 1910 Madras/ Bombay Nat. Hist. Soc. B.M. 1914^11” (BMNH), 4 ex. “Mysore H.K. Slater 1901-182.” (BMNH), 1 ex. “India: ex Mus. Cal- cutta. B.M. 1930-1./ Ind. Mus. Jubbulpore district Cen- tral prov. 1914. Purchased” (BMNH), 2 ex. “India: Oris- sa Bhubaneshwar Feb. 13, 1975, 500’ Mary L. Ripley” (UNSM), 1 ex. “South India Pandioberry State, Karaikal X.1990 leg. Theresa Raj abai Selva Nathan” (CARL). Sri Lanka: 1 S “Sri Lanka Ambalangoda 12.8.1985, Madl” (NHMW), 1 $ “Slid Ceylon Mai 1889 H. Fruhstorfer/ Coll. Brenske” (ZMHB), 1 $ “Ceylon 2de trim 1889 (H. Fruhstorfer)” (ZMHB), 13 ex. “Ceylon, V.1969 50 kmN Colombo leg. P. Schmitz” (CF), 1 S “Inde Anglaise Shi- moga” (MNHN), 1 ex. “Colombo 10.09/ Colombo Cey- lon \9\6. 169! Neoserica bombycina Karsch Determined from Description G. J.A.” (BMNH), 2 ex. “Colombo 9.05/ Ceylon 1910-20.” (BMNH), 1 ex. “Bentota 21.7.1973/ Brancucci” (NHMB), 1 ex. “Ceylan M. Maindron/ Co- lombo 12-20 mai 1901/ Museum Paris Ceylan M. Main- dron” (MNHN), 3 ex. “Colombo Ceylon, India/ 14-VIir44 DE Hardy” (UNSM), 1 ex. “Ceylon, W. Prov. Yakkala, 18 mis NE Colombo, 16-28.11.62 At light Loc. 10/ Lund University Ceylon Expedition Brinck-Andersen- Cederholm” (MZLU), 7 ex. “Sri Lanka: Put. Dist. Wilpettu, National Park, Kali Villu 12-14 June 1975/ Col- lectors: D. H. Messersmith G. L. Williams & P. B. Karunaratne” (USNM), 1 ex. “Ceylon: Ham. Dist. Yala, Palatupana 8-10 March 1972 K. V. Krombein & P. B. Karunaratne” (USNM), 1 ex. “Sri Lanka: Put Dist. Wilpat- tu, Maddikulam 15 June 1975/ Collectors: D. H. Messer- smith G. L. Williams & P. B. Karunaratne” (USNM), 1 ex. “Ceylon: Ham. Dist. Bambowa, 1 . 5mi NW Palatu- pana 23-X-1970, 50’ O. S. Flint, Jr.“ (USNM), 1 ex. “Ceylon: Jaf Dist. Elephant Pass 7-XI-1970, 5’ O. S. Flint, Jr.“ (USNM), 1 ex. “Sri Lanka: Ham. Dist. Palatu- pana Tank 3-A February 1975/ K. V. Krombein P. B. Kara- naratne P. Fernando E. G. Dabrera” (USNM). Burma: 1 S “Victoria Point Tenasserim E.T. Atkinson/ Atkinson Coll. 92-3/ 8.87” (BMNH). Redescription of male syntype of quadriflabellata. Length: 7.0 mm, length of elytra: 4.6 mm, width: 3.9 mm. Body oval, yellowish brown, antenna yellowish, head shiny, dorsal surface dull, nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat- eral margins straight and strongly convergent anteriorly. l61: 1-124 ©ZFMK 38 Silvia] l61: 1-124 ©ZFMK l61: 1-124 ©ZFMK 40 ly serrated longitud] rina beside it, first i Variation. Length: 7.7-8.4 mm, length of elytra: 5.6-6.4 ly shorter than remaining antenneomeres combined; py- [hand written G. Frey]” (CF). Paraty] Bur. det G. Frey, 1972” (CF), 1 6' “Ceylon: Frey, 1972” (CF), 1 “Ceylon: Padukka43km 0 28.IL-2.IIL1971/ J. & M. Sedlacek Bishop Museum” (BPBM), 1 6' “Colombo, Ceylon 1.IL58 leg. G. Frey/ cum Type comparatum/ Autoserica proxima Bur. det G. Frey, 1972” (CF), 1 6', 5 $$ “Ceylon: Kan Dist. Hasalaka, 500’ 22-25-XI-1970 O.S. Flint, Jr.” (USNM), 1 6 ', 2 $$ “Ceylon: Kan. Dist. Laksapana, 1200’ 23-29-IX-1970 O. S. Flint, Jr.“ (USNM), 1 6 ', 1 $ “Cey- lon: Kan. Dist. Kandy 1-15 March 1971 Piyadase & Somapala” (USNM), 1 $ “Sri Lanka: Kan. Dist. Per- adeniya Hantana Hill 29-III-1973, 2000 ft. Baumann & Cross” (USNM), 1 $ “Sri Lanka: Rat. Dist. Kalu Ganga Induruwa Jungle 23-III-1973, 1000 ft. Baumann & Cross” (USNM), 1 $ “Ceylon: Rat. Dist. Uggakaltota 350 ft. Irrigation Bungalow 31 Jan.-8 Feb. 1970 Davis & ma Reservoir 7 Aug. 1975 Peyton & Huang” (USNM), 1 $ “Sri Lanka: N. E. Dist. Hakgala Botanic Garden, 23-25 Feb 1977/ K. V. Krombein P. B. Karunaratne P. Fernan- do D. W. Balasooriya” (USNM), 1 6' “Sri Lanka: N. E. Dist Hakgala Botanic Garden, 23-25 Febl977 blacklight. : Tri. Dist. 1 8-11-X-1977/ K. V. Krombein P. B. Karunaratne P. Fer- nando M. Jayaweera” (USNM), 1 ? “Sri Eanka: Bad. Dist. light” (USNM), 1 $ “Sri Eanka: Pol. Dist. Polounaruwa 6 May 1974 Gans & Prasanna” (USNM), 1 $ “Sri Ean- ka: Mate. Dist. Elkaduwa, Hunas Falls 5-IV-1973, 3000 t” (USNM), 1 $ “Sri I 61: 1-124 l61: 1-124 48 It Loc. 10/ Lund inck-Andersen-C (MZLU), 1 ex. “Ceylon, W. Prov. Yakkala, 18 Colombo, 1.-14.IIL62 At light Loc. 10/ Lund Ui Ceylon (MZLU), 27-VII-1975/Y.-M. D.W. Balasooriya” (USNM), Kandy Tour Inn 31-III-1978 M. D. i.L. Peyton, S. lex. “SriLan (MZLU), 4 ex. “Ceylon, W. 5-13.1.62 Loc. 3 At light / Li (USNM), 1 ex. “Ceylon: Kan. Dist. Nanu Oya, nr Per- adeniya, 1500’ 22-IX-1970 O. S. Flint, Jr.“ (USNM),1 ex. “Ceylon: Kan. Dist. Haslaka, Dam Spillway, 300’ 23-XI-1970 O. S. Flint, Jr.“ (USNM), 1 ex. “Ceylon: Kan. y 1971 F' ■ ), 2 ex. “Ceylon: Kan. E a” (USNM), 3 ex. “SriLan- ht 13-14' ' collected by G.F. Hevel F.E. Dietz IV S. Karunaratne D.W. Balasooriya” (USNM), 1 ex. “Sri Lanka: Col. Dist. Gam- blackl paha Botanical Garden 24 May 1975/ Collectros D.H. maV. Messermilch G.L. (USNM), 4 ex. “Ce: Padaviya 180 ft. 27 Feb.-9 Mar. 1970 Davis & fiiya 1-15 ex. “Cey- Dist. Kandy 1-15 March 1971 Piyadasa & (USNM), 7 ex. “Ceylon: Kan. Dist. Kandy ■ 1971 P. P. Spangler/ Collect at black light” Davies, S. (USNM), 1 ex. “Sri Lanka: Kan. Dist. Kandy, Udawat- 1-3-X-1973 at black light/ K.V. Krombein PB. 2 ex. “Sri Lanka: Kan. (USNM), 2 ex. 15-17 Sept ; 1977/Collected by KV ^ri Lanka: Mate. Dist. Sigiriya, collected by D.M. Davies, S. -unaratne, D. W. Balasooriya” (USNM), 1 ( Mate. Dist. Deniyaya, black light trap 9- lected by D. M. Davies, S. Karunaratne Tiya” (USNM), 2 ex. “Sri Lanka: Kal. D a: Pol. Dist. Polonnamwa 6 May 1974 Gans&Pra- a” (USNM), 2 ex. “Sri Lanka: Rat. Dist. Udawalawa June 1975/ Collecters: D. H. Messersmith G. L. ams & P. B. Karunaratne” (USNM), 1 ex. “Sri Lan- ka: Amp. Dist. Ekgal Am Tank, 100 m 19-23 Febmary 1977/ K. V. Krombein P. B. Karunaratne P. Fernando D. W. Balasooriya” (USNM), 1 ex. “Sri Lanka: Col. Dist. Y. Huang E. (USNM), 1 Sit:- MaaB” (NME), 2 ex. “Ceylon E. Prov. Pottuvil l-12/vii-1983 Ole Mehl leg.” (CDKC). liiiliii 50 i61: 1-124 _BZB-eml 22.04.14 12:41 Seite 51 Sericini of Sri Lanka 5 1 ver 1906-7 E. Bugnion” (MNHN), 9 ex. “S. Ceylon Ha- baraduwa 20.1.^.2.1983 T. Osten leg.” (SMNS), 1 ex. “Ceylon Nuwara Eliya 17.III.78 Renner” (SMNS), 3 ex. “Ceylon N.W.P. Rajakadaluwa 23.VIII.53 F. Reiser” (NHMB), 6 ex. “Sri Eanka (Ceylon) Southern Province Galle, Koggela Lake 14.-28.12.1992 leg. Gerd Kune” (NME), 8 ex. Sri Lanka Vilpattu 8.X.1982 leg. G. Med- vedev” (ZIN), 2 ex. “Beruwela Ceylam 22.IV.75 C.G. Mi- net” (CF), 2 ex. “Beruwela Ceylam 21. IV. 75 C.G. Minet” (CF), 1 ex. “Beruwela Ceylam 23.IV.75 C.G. Minet” (CF), 1 ex. “Ceylon Bentota 21.7.1973/ Brancucci” (NHMB), 3 ex. “Coll. R.I.Sc.N.B. Ceylan/ J.J. Gillet del, vendit.:” (ISNB), 3 ex. “Batticaloa, Ceylon. June 1915. C.F.S. Ba- ker. 1915-338” (BMNH), 1 ex. “Trincomalee, Ceylon. 1915. C.F.S. Baker. 1915-338” (BMNH), 1 ex. “Trinco- malee, Ceylon. C.F.S. Baker. 1914-448” (BMNH), 4 ex. “Nilaveli. Ceylon. C.F.S. Baker. 1914^48” (BMNH), 2 ex. “Trincomalee Ceylon, XI-06/ 1989/ Ceylon, E.E. Green 1910-19” (BMNH), 1 ex. “Batticaloa Ceylon, 5-06/ 1988/ Ceylon, E.E. Green 1910-19” (BMNH), 1 ex. “Ceylon. Trincomali. 16. VI. 07 Oopah Fst. Bainbrigge- Fletcher/ Ceylon T.B. Fletcher. 1909-80” (BMNH), 1 ex. “7.VII.07 Ceylon. Trincomali. / Ceylon T.B. Fletcher. 1909-80” (BMNH), 1 ex. “Colombo, Ceylon/ C.F.S. Ba- ker/April 1915 1915-199.” (BMNH), 4 ex. “Colombo Ceylon, India/ 14-VIIF44 DE Hardy” (UNSM), 14 ex. Sri Lanka Southern Province Galle Distric, Habaraduwa 20.8.^.9.1982 H.J. Bremer leg.” (ZSM), 9 ex. “Ceylon, W. Prov. Yakkala, 18 mis NE Colombo, 14-31.1.62 At light Loc. 10/ Lund University Ceylon Expedition Brinck-Andersen-Cederholm” (MZLU), 6 ex. “Ceylon, W. Prov. Yakkala, 18 mis NE Colombo, 1-14.IIL62 At light Loc. 1 0/ Lund University Ceylon Expedition Brinck- Andersen-Cederholm” (MZLU), 1 ex. “Ceylon, NW. Prov. Chilaw, 31.1.62 At light Loc. 38/ Lund University Cey- lon Expedition Brinck-Andersen-Cederholm” (MZLU), 6 ex. “Ceylon, S. Prov. Hikkaduwa, 11 mis NW Galle, 25.1.62 Loc. 22/ Lund University Ceylon Expedition Brinck-Andersen-Cederholm” (MZLU), 2 ex. “Sri Lan- ka: Put. Dist. 9 mi E Puttalam 19 June 1975 S. L. Wood & J. L. Petty/ Collected in black light trap” (USNM), 10 ex. “Sri Lanka: Man. Dist. Cashew Corp., Ma Villu, 17-21 Feb. 1979, blacklight/ Collected by K. V. Krombein T. Wi- jesinhe S. Siriwaradne T. Gunawaradne” (USNM), 1 ex. “Sri Lanka: Put. Dist. Wilpattu Park, 100ft. Talawila, 9-1 0-IV-73 Baumann & Cross At black lighf ’ (USNM), 2 ex. “Sri Lanka: Man. Dist. Olaithoduvai, lOmi NW of Mannar, 0-50 feet black light, 4-5 November 1976/ Col- lected by G. F. Hevel R. E. Dietz IV S. Karunaratne D. W. Balasooriya” (USNM), 2 ex. “Sri Lanka: Man. Dist. 8mi SE of Mannar, black light, 15 feet 6 November 1976/ Collected by G. F. Hevel R. E. Dietz IV S. Karunaratne D. W. Balasooriya” (USNM), 6 ex. “Sri Lanka: Col. Dist. Hanwella Resthouse 200ft, black light, 2 October 1976/ Collected by G. F. Hevel R. E. Dietz IV S. Karunaratne D. W. Balasooriya” (USNM), 3 ex. “Ceylon: Kan. Dist. Nanu Oya, nr. Peradeniya, 1500’ 22-IX-1970 O. S. Flint, Jr.“ (USNM), 4 ex. “Ceylon: Kan. Dist. Kandy, 1800Ft. Peak View Motel 7-14 Jan. 1970 Davis & Rowe” (USNM), 2 ex. “Ceylon: Kan. Dist. Kandy, 1800 ft. Peak View Motel 15-24 Jan. 1970 Davis & Rowe” (USNM), 1 ex. “Sri Lanka: Kan. Dist. Kandy, 23 November 1974/ Collectors: C. Gans P. Fernando S. Farock” (USNM^ 1 ex. “Sri Lanka: Man. Dist. Kondachchi, Ma Villu 19- IX-1979 In Malaise trap/ T. Wijesinhe L. Jayawick- rema R. Subasinhe” (USNM), 2 ex. “Sri Lanka: Man. Dist. Kondachchi, Ma Villu 1 9-IX-l 979 In Malaise trap/ T. Wi- jesinhe L. Jayawickrema R. Subasinhe” (USNM), 4 ex. “Sri Lanka: Man. Dist. Kondachchi, Ma Villu 20- IX-1979/ T. Wijesinhe L. Jayawickrema R. Subasin- he” (USNM), 1 ex. “Sri Lanka: Man. Dist. Kondachchi, 17-19 Feb 1979Malaise trap/ Collectors: K.V Krombein T. Wijesinhe S. Siriwaradne T. Gunawaradne” (USNM), 1 ex. “Sri Lanka: Man. Dist. Kondachchi, 22-28 January 1978/ PB. Karunaratne T. Wijesinhe M. Jayaweera G. Rat- navira” (USNM), 1 ex. “Ceylon: Kan. Dist. Hantane Hill 10 December 1970 Piyadasa & Sompala” (USNM), 1 ex. “Sri Lanka: Galle Dist. Hiniduma 22 Oct. 1973 M. & R. Robinson” (USNM), 4 ex. “Sri Lanka: Put. Dist. Wilpat- tu National Park, Kali Villu 12-14 June 1975/ Collectors: D. H. Messersmith G. L. Williams P. B. Karunaratne” (USNM), 1 ex. “Sri Lanka: Col. Dist. Beyagama, sea lev- el 28-29 VIII 1973 Ginter Ekis” (USNM), 2 ex. “Sri Lan- ka Bentota 23.5.-15.6. 1990 leg. U. Schmidt/ Coll. A. Ska- le Hof/ Saale” (NME), 2 ex. “Ceylon N.P Mannar 3 1 . 1 .54 F. Reiser/ a. Licht” (NHMB). Redescription. Length: 7.7 mm, length of elytra: 5.5 mm, width: 4.5 mm. Body oblong-oval, light reddish brown, elytra and antenna yellowish brown, labroclypeus shiny, dorsal surface dull, glabrous except for a few small setae on head and lateral margins of pronotum and elytra. Labroclypeus trapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior an- gles, lateral border and ocular canthus producing an in- distinct blunt angle, margins moderately reflexed, anteri- or margin distinctly and narrowly sinuate medially; sur- face slightly carinate along the middle in anterior half, shiny, moderately coarsely and densely punctate, distance between punctures subequal their diameter, partly fusing with each other, behind anterior margin with a few erect setae; frontoclypeal suture feebly incised and weakly curved; smooth area anterior to eye approximately three times as wide as long; ocular canthus moderately long and moderately wide, finely and densely punctate, with two long terminal setae. Frons dull, anterior quarter shiny, with fine, dense punctures, glabrous except for two to three sin- gle setae beside eyes, punctures partly with minute setae. Eyes moderately large, ratio of diameter/ interocular Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK 52 ly- l61: 1-124 ©ZFMK :|||P 58 widest just before apical third, ratio width/ length: 1/ 2.9, Key to species 60 i61: 1-124 ©ZFMK num/ metacoxa: 1/ 2.05. Median apophysis of metacoxa with very dense and evenly short, white, scale-like setae. Ion Expedition 1962 (MZLU). Paratypes: 5 (USNM), 1 $ “Sri La Kandy 29-III-1973, (USNM). ly ly 64 68 iiiiaiiiiiiii isiiWiilSIl ili-iill! 1 61: 1-124 l61: 1-124 ©ZFMK 74 Silvia] ;st just behind middle, st l61: 1-124 ©ZFMK 76 78 (Figs 9P-R, 18D) Loc. 167:1/ Lund L h: 6.2 mm, length of elytra: 4.6 mm, l6I: 1-124 ill iiii i 111 i If filial II 80 I 61: 1-124 ©ZFMK ©ZFMK l61: 1-124 ©ZFMK 86 s, 1 .2 times (6') as long as re- iiiiliiiiiiiii iili 88 90 92 sal margin impunctate, basally and at middle with a few single spines beside the serrated line; lateral face longi- tudinally convex, shiny, with sparse and fine punctures basally and on ventral portion, with short setae in punc- tures; ventral margin coarsely serrate, with three robust and long spines; medial face impunctate and glabrous, apex interiorly near tarsal articulation sharply and deeply truncate. Tarsomeres smooth dorsally, with short, sparse setae ventrally; metatarsomeres with a strongly serrated ridge ventrally, beside the ridge with a fine longitudinal Carina, not carinate laterally; first metatarsomere as long as following two tarsomeres combined and slightly less than twice as long as the dorsal tibial spur. Protibia mod- erately long, bidentate, lateral teeth distinctly displaced apically, slightly but not bluntly widened at middle of the external margin; protarsal claws symmetrical, basal tooth of inner claw simply pointed. Aedeagus: Fig. 13A-C. Habitus: Fig. 15J. Serica lurida Brenske, 1898 (Figs 13D-F, 20B) Serica lurida BxQmkQ, 1898: 261. Serica maculifera Brenske, 1898 (nee Brenske, 1894): 260; Ahrens 1996: 31, syn. n. Serica rubescens 1916: 436, syn. n. Type material examined. Syntypes {lurida)'. 1 S “Cey- lon./ Nietner 860/ lurida Type Brsk./ Typus” (NHMW), 1 S “Ceylon./ Nietner 860/ lurida Type Brsk./ Typus” (NHMW), 1 $ “Ceylon./ Nietner 860/ Type Brsk./ Typus” (NHMW), 1 $ “Ceylon/ lurida m./ coll Brenske” (ZMHB), 1 $ “277/ Haputate Tea estate at 5000 ft./ Mus. Colombo Ceylon/ Serica lurida m./ Museum Leiden Ser- ica lurida Det. Brenske/Cat. No. 1” (RMNH). Syntypes: {maculifera Brsk., 1898 (nee Brsk. 1894); see Ahrens 1996): 1 6 ', 2 $$ “Ceylon Nietner” (ZMHB), 1 $ of these additionally with ^^maculifera Brsk.” (ZMHB), 1 S “Cey- lon Nietner/ 38970” (ZMHB), 1 6' “38970” (ZMHB), 1 6' “Ceylon/ 24943/ Brsk.” (ZMHB), 1 6', 1 ? “24943” (ZMHB), 4 6'6', 3 $$ “Ceylon/ coll. Brenske” (ZMHB), 1 6' of these additionally with "'maculifera Typ.Brsk.” (ZMHB), 1 6 ', 1 $ “Ceylon/ zwaczz/z/era Brsk.” (ZMHB). Syntypes {rubescens)'. 1 S “Bogawantalawa 4.900-5.200 ft. 21.III.^.IV.82/ Ceylon G. Lewis 1910- 320/ Serica rubescens type Arrow/ =S. maculifera Brsk., 1 898 (nee Brsk. 1 894) det. Ahrens 1 995” (BMNH), 4 $ $ “Bogawantalawa 4.900-5.200 ft. 2LIIL^.IV.82/Ceylon G. Lewis 1910-320” (BMNH). Syntype {nano)'. 1 S “Homaloplia nana Ceyl. Dohrn/ nana Brsk. type/ Muse- um Paris ex coll. R. Oberthur” (MNHN). Additional material examined. 8 ex. “Dikaya 3.000^.200 ft. 21. L-7.II.82/ Ceylon G Lewis 1910-320” (BMNH), 1 ex. “Dikaya 3.000-4.200 ft. 21.L-7.II.82/ Ceylon G Lewis 1910-320/ Determined from description G. J. A. Serica maculifera Brsk.” (BMNH), 1 ex. “Hak- gala Ceylon 26-VIII-29/ Ceylon B.M. 1929-370” (BMNH), 3 ex. “Ohiya Ceylon IV-29” (BMNH), 1 ex. “Ohiya Ceylon 6500‘ [feet] 30-IV-28/ Ceylon B.M. 1929-370” (BMNH), 7 ex. “S Sri Lanka 17.-19.III.1994 Polhera - on light 3 km W of Matara Z. Kejval Igt.” (CP), 1 ex. ($)”Belihul-Oya Ceylon 2^^ trin 89 1. Z. Kannegie- ter” (ZMHB), 4 ex. “Ceylon Nietn./ Zool. Inst. Mus. Greifswald Coll. Gerstaecker” (EMAU), 1 ex. ($) ’’Cey- lon, Centr. Prov. Horton Plains, Alt. 7000 ft. 12 mis SSE Nuwara-Eliya 19.111.62 Eoc. 163/ Eund University Cey- lon Expedition 1962 Brinck-Andersson-Cederholm ” (MZEU), 2 ex. “Ceylon N. E. Dist. Kanda-ela Reservoir 5. 6mi SWNuwara Eliya, 6200 ft. 10-21 Feb. 1970 Da- vis & Rowe” (USNM), 1 ex. “Sri Eanka: N. E. Dist. Kan- de-ela 2 June 1975 S. E. Wood & J. E. Petty” (USNM), 1 S “Nuwara-Eliya E. Simon 1892/ Slg. R. Oberthur (Coll. C. Martin) Eing. Nr. 4, 1956” (ZFMK), 1 $ “Cey- lon/ variolosa Ceylan/ Coll. Kraatz/ Serica maculifera Brsk. J. Moser det. 1917.” (DEIC), 1 $ “Ceylon Cande- ze/ Coll. Haag/ Coll. Metzler/ Serica maculifera Brs./ Moser det.” (DEIC), 1 $ “Ceylon/ Coll. Metzler/ Moser det.” (DEIC). Redescription of syntype 1 {lurida). Eength: 5.5 mm, length of elytra: 3.8 mm, width: 3.2 mm. Body moderate- ly oblong, suboval, including antenna and legs reddish brown, head and pronotum with strong green metallic shine, elytra with weak green metallic shine, without dark spots, dorsal surface including entire head dull, nearly glabrous. Eabroclypeus moderately wide, widest at base, lateral margins convexly convergent toward broadly rounded an- terior angles, margins moderately reflexed, anterior mar- gin weakly sinuate medially; surface slightly elevated me- dially and dull, finely and very densely punctate, with a few long, sparsely scattered, erect setae; frontoclypeal su- ture slightly elevate and weakly curved; smooth area an- terior to eye 2.5 times as wide as long, flat; ocular can- thus short but slender (ca. 1/4 of ocular diameter), smooth, without a terminal seta. Frons dull, with fine, dense punc- tures, with minute setae in punctures, with a few setae be- side interior margin of eyes and at middle. Eyes moder- ately large, ratio of diameter/ interocular width: 0.6. An- tenna with ten antennomeres; antennal club in both sex- es with three antennomeres, in male slightly longer than remaining antennomeres combined. Mentum convexly el- evated anteriorly. Pronotum moderately transverse, widest at base, later- al margins evenly convex, moderately convergent anteri- orly, behind anterior angles convex; anterior angles pro- duced and moderately acute, posterior angles almost right-angled, moderately rounded at the tip; anterior mar- gin convexly produced medially, without a fine marginal line, basal margin without marginal line; surface with 96 1 61: 1-124 ©ZFMK Silvia Fabrizi& Dirk Ahrens i61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 101 I bzm_61_ahrens_ Fig. 1. (A-B) Periserica picta Brenske (lectotype), (C-E) P. subsignata (Walker) (syntype of P. picta, ZMHB), (F-H) P. ful- vostriata Brenske (lectotype), (I, J) Periserica interrupta (Walker, 1859) (lectotypeX (K-M) P triflahellata sp. n. (holotype), (N-P) P. densipunctata sp. n. (holotype). (A, C, F, I, K, N) Aedeagus, left side lateral view; (B, E, H, M, P) aedeagus, left side lateral view; (D, G, J, F, O) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 102 102 Silvia Fabrizi & Dirk Ahrens Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK Fig. 3. (A-C) Selaserica distincticornis Brenske (syntype), (D-F) Se. implicata (^renske) (syntype), (G-I) Se. convexiuscula sp. n., (J-L) Se. impexa sp. n., (M-0) Se. padukkana sp. n., (P-R) Se. kanneliyana sp. n. (A, D, G, J, M, P) Aedeagus, left side lateral view; (C, F, I, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N, Q) parameres, dorsal view Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK 12:41 Seite104 Fig. 4. (A-C) Selaserica pusilla Arrow (syntype), (D-F) splendifica (Brenske) (syntype), (G-I) sericea Arrow (holo- type), (J-L) Se. nuwarana sp. n., (M-0) Se. praetexta sp. n., (P-R) Se. sinharajana sp. n. (A, D, G, J, M, P) Aedeagus, left side lateral view; (C, F, I, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N, Q) parameres, dorsal view Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 105 Sericini of Sri Lanka 105 Fig. 5. (A-C) Selaserica wilpattuensis sp. n., (D-F) Se. meridionalis sp. n., (G-I) Neoserica sexfoliata Moser (syntype), (J-L) N. kalaarensis sp. n., (M-0) N. bombycina (Karsch) (syntype Neoserica quadrijlabellata), (P-R) Maladera calcarata (Brenske) (Lectotype). (A, D, G, J, M, P) Aedeagus, left side lateral view; (C, F, 1, L, O, R) aedeagus, left side lateral view; (B, E, H, K, N, Q) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK I bzm_61_ahrens_BZB-eml 22.04.14 12:41 Seite 106 106 Silvia Fabrizi & Dirk Ahrens Fig. 6. (A-C) Maladera breviata (Brenske) (syntype), (D-F) M. breviatella sp. n. (holotype), (G-1) M. rotundata (Walker) (holo- type Autoserica srilanka Frey), (J-L) M. iuga sp. n. (A, D, G, J) Aedeagus, left side lateral view; (C, F, 1, L) aedeagus, left side lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 107 I bzm_61_ahrens_ Sericini of Sri Lanka 107 Fig. 7. (A-C) Maladera mollis (Walker) (Sri Lanka: Marawila), (D-F) M. pseudomollis sp. n. (holotype), (G-I) M carinifrons (Brenske) (syntype), (J-L) M. westermanni (Brenske), (M-0) M. krombeini sp. n. (holotype), (P-R) M. ekisi sp. n. (holotype), (S-U) M cinnaberina (Brenske) (syntype), (V-X) M rufocuprea (Blanchard) (S. India Coimbatore). (A, D, G, J, M, P, S, V) Aedea- gus, left side lateral view; (C, F, I, L, O, R, U, X) aedeagus, left side lateral view; (B, E, H, K, N, Q, T, W) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 108 Dirk Ahrens Fig. 8. (A-C) Maladera fistulosa (Brenske) (lectotype), (D-F) M dubia (Arrow) (holotype), (G-I) M. weligamana (Brenske) (syntypQ Autoserica fistulosa, Ceylon), (J-L) M coxalis (Moser) (syntype). (A, D, G, J) Aedeagus, left side lateral view; (C, F, I, L) aedeagus, left side lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 Seite 109 Sericini of Sri Lanka 109 Sericini of Sri Lanka Fig. 11. (A-C) Maladera kandyensis sp. n. (holotype), (D-F) M hastata sp. n. (holotype), (G-I) M lindulana sp. (J-L) M. tricuspidata sp. n. (holotype), (M-0) M. hortonensis sp. n. (holotype), (P-R) M hadullana sp. n. (holoty n. (holotype), pe), (S-U) M view; (C, F, I, i61: 1-124 _BZB-eml 22.04.14 12:41 Seite112 112 Silvia Fabrizi & Dirk Ahrens Fig. 12. (A-C) Maladera bandarwelana sp. n. (holotype), (D-F) M. setosa (Brenske) (syntype, Konbir), (G-1) M. strata (Brenske) (syntype Autoserica apogonoides Brenske), (J-L) M. brevis (syntype). (A, D, G, J) Aedeagus, left side lateral view; (C, F, 1, L) aedeagus, left side lateral view; (B, E, H, K) parameres, dorsal view. Scale bar: 1 mm. Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 SeitellS I bzm_61_ahrens_ Sericini of Sri Lanka 113 Bonn zoological Bulletin - Supplementum i61: 1-124 ©ZFMK Fig. 14. Habitus. (A) Periserica picta Brenske, (B) R interrupta (Walker), (C) R fulvostriata Brenske, (D) Selaserica lucidicol- lis sp. n., (E) Se. pusilla Arrow, (F) Se. nitida (Candeze), (G) Se. impexa sp. n., (H) Se. meridionalis sp. n., (I) Neoserica bombyci- na (not to scale). Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 SeitellS I bzm_61_ahrens_ Sericini of Sri Lanka 115 Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 SeitellO I bzm_61_ahrens_ 116 Silvia Fabrizi & Dirk Ahrens Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK m_61_ahrens_BZB-eml 22.04.14 12:41 Seite117 Sericini of Sri Lanka group and M. madurensis group (in part D). Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 SeitellS Dirk Ahrens Fig. 18. Distribution of the Sericini species of Sri Lanka (as indicated by the legend): (A-C) Maladera subgenus Cephaloserica and (D) Maladera fistulosa group (part). Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK _BZB-eml 22.04.14 12:41 SeitellQ I bzm_61_ahrens_ Sericini of Sri Lanka 119 Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK 120 Silvia Fabrizi & Dirk Ahrens Fig. 20. Distribution of the Sericini species of Sri Lanka (as indicated by the legend): (A) Maladera setosa group and (B) Seri- ca (sensu lato). (C) Climatic zones (see text), their number of species (D) and their faunistic similarity based on the Jaccard Index (agglomeration method: unweighted pair-group average) in Sericini (E). Bonn zoological Bulletin - Supplementum 61 : 1-124 ©ZFMK Sericini of Sri Lanka 1 61: 1-124 ©ZFMK 1 1 a iiiiiiiiiMiiiiii iiiiiiii I ill iiiiiii ill iiir 3. 1 1 1 iiiii mmmm i m mm m m i rens_BZB-eml 22.04.14 12:41 Seite 124 124 Silvia Fabrizi & Dirk Ahrens anderssoni sp. n. (Maladera) 59, 88, 110, apogonoides Brenske, 1 898; syn carinirostris BTQnskQ, 1896; syn atrata Burmeister, 1855 (nec Reiche 1847); syn atratula Dalla Torre, 1917; syn badullana sp. n. {Maladera) 59, 69, bandarwelana sp. n. {Maladera) 89, bisornata sp. n. {Maladera) 59, 74, bombycina (Karsch, 1882) {Neoserica) ..35, 97, 105, breviata (Brenske, 1898) {Maladera) 39, 41, 44, breviatella sp. n. {Maladera) 39, 43, (Blanchard, USO) {Hemiserica) 91, brincki sp. n. {Maladera) 59, 79, buruensis Brenske, 1899, syn calcarata (Brenske, 1898) {Maladera) ...39, 90, 105, carinifrons (Brenske, 1896) {Maladera) 47, 49, chalybaeaBxQxs&ks, 1898; syn cinerea Brenske, 1898, syn cinnaberina (Brenske, 1 898) {Maladera) confirmata (Walker, 1859) {Selaserica ) ... convexiuscula sp. n. {Selaserica) coxalis (Moser, 1915) {Maladei densipunctata sp. n. {Perisericc distincticornis (Brenske, 1898) {Selaserica) ...13, 25, diyalumana sp. n. {Maladera) 59, 77, dubia (Arrow, 1916) {Maladera) 59, 66, 69, T. ekisi sp. n. {Maladera) A1 , 57, 9^ Brenske, 1898; syn fistulosa (Brenske, 1898) {Maladera) .58, 59, 62, 97, flinti sp. n. {Maladera) 59, 86, fulvostriata Brenske, 1898 {Periserica) ... Al, 54 ....13, 17 ....13, 24 ....59, 60, 62, 64 ica) fusa Brenske, 1898 {Serica) hastata sp. n. {Maladera) heveli sp. n. {Maladera) hortonensis sp. n. {Maladera) . zmmMw/to Brenske, 1898; syn. . impexa sp. n. {Selaserica) implicata (Brenske, 1 898) {Sel interrupta (Walker, 1859) {Pen iuga sp. n. {Maladera) kalaarensis sp. n. {Neoserica) . kandyensis sp. n. {Maladera) .. kanneliyana sp. n. {Selaserica) kitulgalana sp. n. {Serica) knucklensis sp. n. {Selaserica) krombeini sp. n. {Maladera) .... kuruwitana sp. n. {Maladera) . Imdulana sp. n. {Maladera) .... lucidicollis sp. n. {Selaserica) . 92, 97, 113 5, 9 59, 83, 85 59, 78, 80 ....58, 67, 70, 13,23, 103, serica) ....\3, 19, 23, erica) 5, 11, 101, 39, 46, 35, 38, 59, 84, 86, 13,21, 92, 96, 13, 16,21, 47, 56, 58 ..59, 62, 63, 66, 59, 71, 13,20,25, 702 urida Brenske, 1898 {Serica) 92, 94, 96, 97, 113, 120 naculicauda Arrow, 1916 {Serica) 13, 18, 98, 102, 116 naculifera Brenske, 1898 (nec Brenske, 1894); syn 94, 97 navilluensis sp. n. {Maladera) 59, 85, 87, 770, 119 teridionalis sp. n. {Selaserica).... \3, 32, 33, 97, 105, 114, 116 nollis (Walker, 1859) {Maladera) 47, 50, 54, 57, 707, 118 ^ana Brenske, 1898 {Serica) 92, 94, 95, 97, 113, 115, i: 7a(Candeze, \^6\) {Selaserica) ... nilaveliensis sp. n. {Malader padukkana sp. n. {Selasericc pavieana Brenske, 1 899; syn. ...13, 14, 16, 28, 702, 114, 59, 80, 770, 14,27, 104, ...13, 22, 17 , 7 poyagana sp. n. {Maladera) . 9 praetexta sp. n. {Selaserica) . 8 pseudomollis sp. n {Maladera) ;2 pubescens (Arrow, 1916) {Maladei 1 8 pusilla Arrow, 1916 {Selaserica) . . i) 13, 22, 24, 103, 116 91 5, 6, 7, 8, 9, 10, 707, 114, 116 59, 70, 109 14, 29,31,704, 116 47, 52, 707, 118 59, 72, 75, 76, 84, 770, 119 14,27,28, 30, 32, 104, 114, 116 quadriflabellata {BrmskQ, 1896); syn 35, 36, 705 romanoi sp. n. {Maladera) 59, 87, 88, 89, 110, 119 rotundata (Walker, 1859) {Maladera) 39, 44, 47, 106, 117 rubescens Arrow, 1916; syn 94 rufocuprea (Blanchard, 1850) {Maladera) 47, 52, 97, 707, 115, 118 schintlmeisteri sp. n. {Maladera) 59, 64, 66, 109, 119 scutellaris Arrow, 1916 {Selaserica) 1 3, 15 sericata Brenske, 1898; syn. 35 sericea Arrow, 1916 {Selaserica) (Brenske, n96) {Maladera) ... ta Mose 1915 {Neose ’ca) 14, 33, 104, IK 39, 73, 90, 97 772, 115, 12( 35, 36, 705, 111 .49, 50, 5: 14, 30, 704, IK 13, 14, 26, srilanka Frey, 1974; syn straba (Brenske, 1898) {Maladera) stupida Brenske, 1900; syn. . subsignata (Walker, 1859) {Periserica) 5, 6, 8, 9, 7 sumatrensis Bronsko, 1898; syn tricuspidata sp. n. {Maladera) 59, 75, 77, 81, 7 truncatus Mittal, 1976; syn. . uggalkaltotaensis sp. n. {Maladera) 59, 76, 7 vflno7o.sfl Motschulsky, 1863 {Serica) ventralis WdXkor, 1859; syn. .. ;nske, 1900) {Maladera) .59, 60, 61, 108, termanni (Brenske, 1898) {Maladera) 47, 5 attuensis sp. n. {Selaserica) 13, 32, 705, di sp. n. {Maladera) 59, 65, 109, lensis sp. n {Maladera) 59, 81, 110, Bonn zoological Bulletin - Supplementum 61: 1-124 ©ZFMK Bonn zoological Bulletin - Supplementum Vol. 61 (2014) Managing Editor: Thomas Wesener Zoologisches Forschungsmuseum Alexander Koenig - Leibniz-lnstitut fur Biodiversitat der Tiere (ZFMK) Adenaueralllee 160, D-53113 Bonn, Germany ISBN-13: 978-3-925382-65-9 ISSN: 0302-671X Back-Cover illustration: Male genitalia of Maladera tricuspidata sp. n.: newly developed processes of the aedeagus replace functionally the reduced parameres. Cover illustration: Habitus of Selaserica impexa sp. n. in front of the map of the examined material from Sri Lanka. Leibniz-Gemeinschaft Bundesministerium SorsSung Wissenschaft und Forschung Ministerium fiir Innovation, des Landes Nordrhein-Westfalen Bonn zoological Bulletin - Supplementum Volume 61 (2014)