A monograph of the Sericini of India (Coleoptera: Scarabaeidae) Dirk Ahrens & Silvia Fabrizi Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Contents Volume Abstract 3 Key words 3 Introduction 4 Material and methods 4 Depositories 5 Systematic part 6 Serica MacLeay, 1819 9 Serica (s. stricto) MacLeay, 1819 13 Serica (s. lato) 16 Pachyserica Brenske, 1897 18 Calloserica Brenske, 1894 21 Xenoserica Ahrens, 2005 24 Nepaloserica Frey, 1965 25 Sericania Motschulsky, 1860 31 Gynaecoserica Brenske, 1896 34 Lasioserica Brenske, 1896 41 Amiserica Nomura, 1974 48 Chrysoserica Brenske, 1897 53 Lepidoserica Nikolaev, 1979 54 Meriserica Brenske, 1897 54 Deroserica Moser, 1915 56 Pachyderoserica Moser, 1920 59 Microsericaria Nikolaev, 1979 59 Anomioserica Arrow, 1946 61 Selaserica Brenske, 1897 69 Neoserica Brenske, 1894 72 Neoserica (s. stricto) 72 Neoserica abnormis group 73 Neoserica dichroa group 88 Neoserica lubrica group 97 Neoserica septemlamellata group 100 Neoserica speciosa g ro u p 101 Neoserica uniformis group 101 Neoserica variega fa g ro u p 101 Neoserica incertae sedis 107 Gastroserica Brenske, 1894 122 Bonn zoological Bulletin 65 (1 & 2): 1-355 65 Tetraserica Ahrens, 2004 122 Trioserica Moser, 1922 128 Microserica Brenske, 1894 129 Oxyserica Brenske, 1900 134 Maladera Mulsant & Rey, 1871 136 Maladera assamica group 136 Maladera egregia group 139 Maladera ferruginea group 142 Maladera fistulosa group 147 Maladera granuligera group 149 Maladera indica group 152 Maladera lugubris group 161 Maladera madurensis group 177 Maladera marginella group 179 Maladera modestula group 210 Maladera placida group 211 Maladera quinquidens group 215 Maladera schenklingi group 215 Maladera servitrita group 217 Maladera significabilis group 218 Maladera sikkimensis group 219 Maladera teinzoana group 221 Maladera thomsoni group 222 Maladera (subgenus Cephaloserica) 226 Maladera (subgenus Hemiserica) 250 Maladera (subgenus Omaladera) 257 Maladera incertae sedis 263 Species incertae sedis 271 Zoogeography 273 Acknowledgements 274 References 275 Tables 280 Figures 289 Appendix 1 344 Index 347 ©ZFMK Bonn zoological Bulletin 65 (1 & 2): 1-355 December 2016 A monograph of Sericini of India (Coleoptera: Scarabaeidae) Dirk Ahrens* & Silvia Fabrizi Zoologisches Forschungsm use urn A. Koenig, Centre of Taxonomy and Evolutionary Research, Adenauerallee 160, D-53113 Bonn, Germany; * Corresponding author: E-mail: ahrens.dirk_col@gmx.de Abstract. In this monograph all available type material and specimens of Sericini chafers (Coleoptera: Scarabaeidae) from India and ad¬ jacent countries are revised and results from previous revisionary work are summarised. Here, 127 new species and one new subspecies are described: Amiserica hunliana sp. n., A. lutulenta sp. n., Anomioserica kotagiriensis sp. n., A. maesi sp. n., A. symmetrica sp. n., Deroser- ica championi sp. n., D. koelleri sp. n., Gynaecoserica etalinensis sp. n., Lasioserica verschraegheni sp. n., Maladera alibagensis sp. n., M. allolaterita sp. n., M. amboliensis sp. n., M. anaimalaiensis sp. n., M. andrewesi sp. n., M. antispinosa sp. n., M. ballariensis sp. n., M. balphakramensis sp. n., M. bangaloreensis sp. n., M. bombycinoides sp. n., M. cardamomensis sp. n., M. coimbatoreensis sp. n., M. constans sp. n., M. declarata sp. n., M. decolor sp. n., M. densipilosa sp. n., M. dimidiata sp. n., M. dunhindaensis sp. n., M. faceta sp. n., M. fastuosa sp. n., M. ferekanarana sp. n., M. garoana sp. n., M. geniculata sp. n., M. granigera sp. n., M. hampsoni sp. n., M. hunlien- sis sp. n., M. initialis sp. n., M. jaintiaensis sp. n., M. johannesi sp. n., M. kallarensis sp. n., M. kostali sp. n., M. kumilyensis sp. n., M. lonaviaensis sp. n., M. lorenzi sp. n., M. malabarensis sp. n., M. minops sp. n., M. mussardi sp. n., M. mysoreensis sp. n., M. naduvata- mensis sp. n., M. namborensis sp. n., M. neotridentipes sp. n., M. nigrolucida sp. n., M. padaviyaensis sp. n., M. paraprabangana sp. n., M. pauper sp. n., M. praviforceps sp. n., M. profana sp. n., M. propagator sp. n., M. pseudohongkongica sp. n., M. rudimentalis sp. n., M. sagittula sp. n., M. schoenwitzae sp. n., M. sedida sp. n., M. sempiternella sp. n., M. senfti sp. n., M. seriatoguttata sp. n., M. shimogana sp. n., M. shiva sp. n., M. slateri sp. n., M. songsakensis sp. n., M. subabbreviata sp. n., M. submucronata sp. n., M. sunaiensis sp. n., M. sylhetensis sp. n., M. tempestiva sp. n., M. theresae sp. n., M. tiefermanni sp. n., M. trivandrumensis sp. n., M. trochaloides sp. n., M. tubu- lata sp. n., M. vernacula sp. n., M. versuta sp. n., M. xanthoptera sp. n., Microserica roingensis sp. n., Neoserica agumbeensis sp. n., N. ammattiensis sp. n., N. astuta sp. n., N. aulica sp. n., N. bhalukpongensis sp. n., N. bicolorea sp. n., N. disciplineensis sp. n., N. flagrans sp. n., N. fusiforceps sp. n., N. genieri sp. n., N. gravida sp. n., N. incisa sp. n., N. infamiliaris sp. n., N. kalkadensis sp. n., N. kejvali sp. n., N. keralana sp. n., N. mudigereensis sp. n., N. munnarensis sp. n., N. nilgiriana sp. n., N. parilis sp. n., N. periyarensis sp. n., N. pilis- triata sp. n., N. plagiata sp. n., N. plateosa sp. n., N. plebea sp. n., N. pseudomajor sp. n., N. pushkarensis sp. n., N. rajasthanica sp. n., N. rotundotibialis sp. n., N. sforziae sp. n., N. shillongensis sp. n., N. sparsesquamata sp. n., N. submaculosa sp. n., N. subsetosa sp. n., N. unciforceps sp. n., N. ziyardamensis sp. n., Selaserica karnatakaensis sp. n., Se. vagans sp. n., Serica feresegregata sp. n., Tetraserica bendai sp. n., T. impar sp. n., T. rufimargo sp. n., T. uncinata sp. n., T. univestris sp. n. The study resulted in 21 new synonymies and 28 new combinations: Gynaecoserica Brenske, 1 896 (= Leuroserica Arrow, 1946, syn n ), Maladera (subgenus Cephaloserica Brenske, 1900) (= Coronoserica Brenske, 1902, syn. n.), Anomioserica liliputana (Moser, 1916) comb, n., Anomioserica tarsalis (Frey, 1960) comb, n., Deroserica kidzeri (Frey, 1976) comb, n., Gynaecoserica fulgida (Arrow, 1946) comb, n., G. lateralis (Arrow, 1946) comb, n., G. stemm- leri (Frey, 1975) comb, n., Maladera beata (Brenske, 1902) comb, n., M. bilobata (Arrow, 1945) comb, n., M bombycina (Karsch, 1882) comb, n., M. burmeisteri ssp. alternans (Frey, 1975) stat. n., M. clypeata (Fairmaire, 1887) (= Autoserica spectabilis Brenske, 1898, syn. n.; Autoserica colossica Brenske, 1898, syn. n.), M. fumosa (Brenske, 1898) (= Autosericaperpendicidaris Khan & Ghai, 1980, syn. n.), M. ignava (Brenske, 1894) (= Autoserica distincta Moser, 1915, syn. n.), M. indica (Blanchard, 1850) (= Serica nilgirensis Sharp, 1903, syn. n.; Autoserica singularis Brenske, 1898, syn. n.; Serica laminipes Moser, 1915, syn. n.), M. laterita (Moser, 1915) comb, n., M. lu- teola (Moser, 1918) comb, n., M. mutabdis (Fabricius, 1775) comb. n. (= Melolontha globosa Herbst, 1790, syn. n.; Serica barwayana Brenske, 1896, syn. n.; Autoserica pygmaea Frey, 1972 syn. n.), M. nasuta (Brenske, 1894) comb. n. (= Hemiserica mayarami Khan & Ghai, 1980 syn. n.), M. nasutella (Ahrens, 2004) comb, n., M. trilobata (Khan & Ghai, 1980) comb, n., M. tyrannica (Brenske, 1898) (= Autoserica opalescens Moser, 1915, syn. n.), M. ventriosa (Brenske, 1894) comb. n. (= Autoserica spoliata Brenske, 1898, syn. n.; Au¬ toserica discrepans Moser, 1915, syn. n.), Meriserica kulzeri (Frey, 1976) comb, n., Meriserica setosicollis (Frey, 1976) comb, n., Mi- crosericaria quadrinotata (Moser, 1915) (= Microsericaria arrowi Frey, 1972, syn. n.), Microsericaria stellata (Arrow, 1946) comb, n., Neoserica barberi (Sharp, 1903) comb, n., N. flavoviridis (Brenske, 1896) comb, n., N. quadrilamellata (Brenske, 1896) (= Meriserica chilkensis Arrow, 1923: 260, syn. n.), N. setigera (Brenske, 1894) comb, n., Tetraserica miniatida (Moser, 1915) comb, n., Xenoserica erectosetosa (Ahrens, 1999) comb, n., X. karnaliensis (Ahrens, 1999) comb, n., X. koshiana (Ahrens, 1999) comb, n., X. somathangana (Ahrens, 1999) comb, n., and X. yadongensis (Liu & Ahrens, 2014) comb. n. The following substitution names are given for junior homonymies: Maladera eusericina nom. n. for the primary homonym Autoserica sericina Frey, 1972 [nec Moser, 1916] and Maladera freyi nom. n. for the secondary homonym Maladera opaca (Frey, 1975) [nec Moser, 1924], Hemiserica Brenske, 1894 is downgraded to a subgenus of Maladera Mulsant & Rey, 1871. Lectotypes are designated of the following taxa: Microserica fairmairei Brenske, 1898, Maladera satrapa (Brenske, 1898), M. burmeisteri ssp. burmeisteri (Brenske, 1898), M. granuligera (Blanchard, 1850), M. proxima (Burmeis- ter, 1855), M. significans (Brenske, 1898), M. basalis (Moser, 1915), M. setosiventris (Moser, 1916), M. beata (Brenske, 1902), M. mu- tabilis (Fabricius, 1775), Autoserica colossica Brenske, 1898. The currently known fauna of Sericini of the Indian subcontinent compris¬ es 660 species plus 9 uncertain species group taxa. Most of the species of the Indian subcontinent are highly endemic. Based on the re¬ sults of the species distribution, a ecozone-oriented species checklist is provided and the zoogeography and diversity patterns of Sericini of the Indian subcontinent are explored. Key words. Taxonomy, biogeography, biodiversity patterns, Scarabaeoidea, Melolothinae, chafers beetles, new species, key to species, India. Received: 18.02.2016 Accepted: 07.12.2016 Corresponding editor: R. Peters 4 Dirk Ahrens & Silvia Fabrizi INTRODUCTION The Indian subcontinent bears some of the world’s most diverse regions. It encompasses a wide spectrum of eco- zones, ranging from deserts to high mountains, highlands, tropical and temperate forests, swamplands, plains, as well as grasslands. The region includes several biodiversity hotspots: the Western Ghats and Sri Lanka, the Himalayas and the Tndo-Burmese’ region (Myers et al. 2000), which are characterised by numerous endemic species. The largest part lies within the Indo-Malayan (Orien¬ tal) realm, while the northern parts shape along the Hi¬ malayan range the boundary with the Palearctic region; 2000 to 2500 m are considered to be the altitudinal bound¬ ary between the Indo-Malayan and Palearctic zones (Mani 1974). The climate is variable, although the entire Indian Sub¬ continent is heavily influenced by summer monsoons that cause major seasonal changes in vegetation and habitat. These patterns are reinforced by elevation and exposition to rainfall. Generally, information about the arthropods and beetles of India is quite limited. Significant work having been done only in a few groups of insects, notably select groups of Lepidoptera, Odonata, Hymenoptera, Coleoptera and Heteroptera. Most important concerted attempts to docu¬ ment the biodiversity have been made since the publica¬ tion of the series “The Fauna of British India, Including Ceylon and Burma”, but most of these works date back nearly 100 years or more. While for a number of scarab beetles such volumes exist, in melolonthine chafers (Melolonthinae) there has never been published such a volume and therefore only fragmentary work since the first taxonomic treatments of the Indian fauna exist (Fabricius 1775, Blanchard 1850, Burmeister 1855). Since then, on¬ ly a few authors contributed towards the knowledge in Sericini, with major contributions given by Brenske (1894-1902), Moser (1908-1920), Frey (1962a-1975b). With nearly 4000 described species, Sericini chafers (Coleoptera: Scarabaeidae) represent a very diverse tribe of beetles with a nearly worldwide distribution only ab¬ sent in Australia and circumpolar regions (Ahrens 2006e). Most extant species are found in a monophyletic lineage (“modern Sericini”; Ahrens 2006e; Ahrens and Vogler 2008). It comprises two large palaeotropic subtribes, Seric- ina and Trochalina, which number about 3000 species and 370 species, respectively. The phytophagous Sericini be¬ long to the lineage of Pleurostict chafers (Scarabaeidae), which is thought to have greatly diversified with the rise of angiosperms around 108 million years ago (Mya) (Ahrens et al. 2014b). Compared to the soil dwelling lar¬ val stage, the lifetime of adults is short. The origin of Sericini was proposed to be in Africa, because of the ex¬ clusively African distribution of Ablaberini, the sister group of Sericini (Ahrens 2006e). The divergence of Sericini from Ablaberini dates back to ca 100 Mya (Ahrens et al. 2014b). Our recent taxonomic work on the Indian fauna was started by Ahrens (1995a, b) and was continued with a se¬ ries of papers and monographs on the fauna of the Hi¬ malayas and adjacent regions (Ahrens 1998a-2007c, Ahrens & Fabrizi 2009a-2011, Liu et al. 2014a). Finally, a first part of a comprehensive treatment of the fauna of the Indian subcontinent was given by Fabrizi & Ahrens (2014) who revised the Sericini fauna of Sri Lanka. This monograph represents the second part of this revision sum¬ marising all available information on the Sericini fauna of the Indian subcontinent. MATERIAL AND METHODS The principal terminologies (Fig. 1) and methods used for measurements, specimen dissection and genital prepara¬ tion are described in detail in Ahrens (2004b, 2006e). The examined material is cited with the original label contents given in quotation marks, multiple labels are separated by a “/”, additional comments are given in square brackets. GPS coordinates are given for each newly sampled local¬ ity in Appendix 1, that of previously sampled ones may be found in Ahrens (2004b) or in Fabrizi & Ahrens (2014), or in the web (http://www.serica.gmxhome.de/layout05/ HimGazlNEW.htm). For each species, taxonomy was un¬ derpinned by a comprehensive list of synonymies and ref¬ erences (of faunistic, phylogenetic, and taxonomic treat¬ ments), however, in the latter, citations from pure taxo¬ nomic checklists (e.g., Dalla Torre 1912, Krajcik 2012) were included only when this was crucial for a better un¬ derstanding of the taxonomic background of the species. Male genitalia were glued to a small pointed card and photographed in both lateral and dorsal view with a stere¬ omicroscope Olympus SZX-12 using a Jenoptik ProgRes digital camera (C5 5.0), and at later stage, with a Leica SM-125 and the digital camera DFC 420. Using AU¬ TOMONTAGE software a number of single-focused im¬ ages were combined to obtain an image that was in focus throughout. The resulting images were subsequently dig¬ itally edited to remove the errors of the AUTOMONTAGE reconstruction and to obtain a white background. A part of the figures of male genitalia was done as pencil draw¬ ings (type material revised more than ten years ago) which were subsequently graphically edited to obtain a black and white image similar to a photograph. Distribution maps were generated using QGIS 2.8.2. based on the geographical coordinates. The same software was also used to extract the ecoregion data for each species based on the Terrestrial Ecoregion layer of World Wildlife Fund and Nature Conservancy (2011; Olson et al. 2001). Species richness, sampling density, Shannon diversity (Shannon & Weaver 1949, Spellerberg & Fedor 2003) and Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India Chao diversity estimator (corrected) were calculated in DI- VA-GIS v.5 (Hijmans et al. 2001). The similarity of the species composition of ecoregions and that of the species occurring in different ecoregions was calculated in PAST v.3.1 (Hammer et al. 2001) using the neighbour-joining method based on the Jaccard index (Jaccard 1902). For the calculation of ecoregion-similarity regions with less than three sampled species were excluded, and also a re¬ duced analysis on regions with complete sample investi¬ gation was performed. Currently, most female specimens of Sericini are diffi¬ cult to identify (to genus and species), as diagnostic char¬ acters of genera, as defined currently, are often submit¬ ted to strong sexual dimorphism (e.g., the number of an- tennomeres of club). Additionally, only the shape of aedea- gus often offers sufficient, detailed and stable variation to allow a discrimination of the species, while characters use¬ ful in other scarab groups, such as shape of the head and pronotum or the punctuation and sculpture of the body sur¬ face, are often highly variable in Sericini. For the latter reason, many species keys need to include genital char¬ acters as well, and a reliable identification of species is impossible without examination of male genitalia. Given the enormous diversity of Indian Sericini fauna, this contribution was constrained by some editorial com¬ promises in order to save print space giving priority to species that so far have not yet been revised and to those that are new to science. Taxa that were treated previous¬ ly by the author(s), are referenced, included in the key, but not described in detail nor illustrated. DEPOSITORIES BMNH Natural History Museum, London, UK BPBM Bernice P. Bishop Museum, Honolulu, USA BYU Brigham Young University, Provo, Utah, USA CAKB coll. A. Kleeberg, Berlin, Germany CARL coll. A. Reichenbach, Leipzig, Germany CASH coll. A. Skale, Hof, Germany CAWP coll. A. Weigel, PoBneck, Germany CAZK coll. A. Zubair, Karachi, Paki stan CDEG coll. D. Erber, Giessen-Lahn, Germany CDUL Institute of Systematic Biology, Daugavpils University, Latvia CDKC coll. D. Keith, Chartres, France CF coll. G. Frey (at the NHMB), Switzerland CGST coll. G. Sabatinelli, Trevesin, France CJSB coll. J. Schulze, Berlin, Germany CJWE coll. J. Weipert, Erfurt, Germany CK coll. D. Krai, Prague, Czech Republic CM coll. E. Migliaccio, Roma, Italy CMNC Canadian Museum of Nature, Ottawa, Canada CN coll. M. Nikodym, Prague, Czech Republic CNA coll. A. Napolov, Riga, Latvia CPLM coll. P. Lago, Mississippi University Bonn zoological Bulletin 65 (1 & 2): 1-355 5 CPPB coll. P. Pacholatko, Brno, Czech Republic CRRP coll. R. Rous, Prague, Czech Republic CRSW coll. R. Schuh, Wien, Austria CTIO coll. T. Itoh, Osaka, Japan DEIC Senckenberg Deutsches Entomologisches Institut, Miincheberg, Germany HAHC coll. Henry and Anne Howden, Ottawa, Canada HNHM Hungarian Natural History Museum, Budapest, Hungary IAAS Institute of Agricultural and Animal Sciences of Tribhuvan University, Rampur, Nepal ISNB Institut Royal des Sciences naturelles de Belgique, Bruxelles, Belgium MEMU Mississippi State University, U.S.A. MHNG Museum d’Histoire Naturelle, Geneve, Switzerland MLUH Martin-Luther-Universitat, Wissenschaftsbereich Zoologie, Halle/Saale, Germany MNHN Museum national d’Histoire naturelle, Paris, France MSNG Museo Civico di Storia Naturale G. Doria, Genova, Italy MSNM Museo Civico di Storia Naturale, Milano, Italy MZR Museo di Zoologia, Universita di Roma “La Sapienza”, Italy MZUF Museo Zoologico “La Specola”, Universita di Firenze, Italy NHMB Naturhistorisches Museum, Basel, Switzerland NHMW Naturhistorisches Museum, Wien, Austria NHRS Naturhistoriska Riksmuseet Stockholm, Sweden NME Naturkundemuseum Erfurt, Germany NMPC National Museum Prague (Natural History), Czech Republic NZSI National Zoological Collection, Zoological Survey of India, Calcutta, India SEAN Servicio Entomologico Autonomo, Museo Entomologico, Leon, Nicaragua SMNS Staatliches Museum fur Naturkunde, Stuttgart, Germany SMTD Senckenberg Museum fur Tierkunde, Dresden, Germany UMB Ubersee-Museum Bremen, Germany UMRM University of Missouri, Columbia, W. R. Enns Entomology Museum, U.S.A. UNSM University of Nebraska State Museum, Lincoln, U.S.A. USNM National Museum of Natural History, Washington D.C., U.S.A. ZFMK Zoological Research Museum A. Koenig, Bonn, Germany ZIN Russian Academy of Sciences, Zoological Institute, St. Petersburg, Russia ZMHB Museum fur Naturkunde, Berlin, Germany ZMUC University of Copenhagen, Zoological Museum, Denmark ZSM Zoolog. Staatssammlung, Miinchen, Germany ©ZFMK Dirk Ahrens & Silvia Fabrizi SYSTEMATIC PART Key to genera and species groups 1 Protibia with three teeth.2 1 ’ Protibia with two teeth.4 2 Body unicoloured: yellowish brown or reddish brown. Apical border of elytra with a fine membranous rim. Eyes large (ratio diameter/interocular width: 0.57-0.83). Antennal club in male with three or four antennomeres. Metatarsomeres setose ventrally. Hy- pomeron with or without transverse basoventral Cari¬ na. Selaserica Brenske T Body bicoloured.3 3 Apical border of elytra without membranous rim. Eyes small (ratio diameter/interocular width: 0.42-0.6). An¬ tennal club in male with three to five antennomeres. Hypomeron without transverse basoventral carina. Apex of metatibia concavely sinuate interiorly near tarsal articulation. Periserica Brenske 3’ Apical border of elytra with membranous rim. Hy¬ pomeron with transverse basoventral carina. Apex of metatibia concavely truncate near tarsal articulation. . Trioserica Moser 4 Antennal club in male composed of three anten¬ nomeres.43 4’ Antennal club in male composed of more than three antennomeres.5 5 Hypomeron not carinate.6 5’ Hypomeron carinate.7 6 Dorsal margin of metatibia sharply carinate. Body dark brown to reddish brown, unicolored. . Tetraserica Ahrens 6’ Dorsal margin of metatibia longitudinally convex. Body dark yellow with numerous dark spots. . Neoserica dichroa group 7 Antennal club in female composed of three anten¬ nomeres.8 7’ Antennal club in female composed of more than three antennomeres.25 8 Posterior margin of metafemur serrate ventrally and dorsally.9 8’ Posterior margin of metafemur smooth ventrally. .14 9 Anterior angles of pronotum obsolete.12 9 ’ Anterior angles of pronotum acute and moderately pro¬ duced.10 10 Dorsal body surface yellowish brown with numerous dark spots. Neoserica (s.l.) variegata group 10’Dorsal body surface dark brown with greenish shine .11 11 Antennal club straight. Punctures of body surface red¬ dish brown. . Neoserica (s.l.) inspergata Ahrens & Fabrizi Bonn zoological Bulletin 65 (1 & 2): 1-355 11 ’ Antennal club reflexed. Punctures of body coloured as the rest of the surface. . Neoserica (s.l.) inops Ahrens & Fabrizi 12 Dorsal surface nearly glabrous. . Gastroserica Brenske 12’Dorsal surface densely setose.13 13 Metatibia beside dorsal margin with a serrated longi¬ tudinal line or carina. Neoserica (s.str.) Brenske 13’ Metatibia beside dorsal margin without a serrated lon¬ gitudinal line or carina. Calloserica Brenske 14 Metatibia beside dorsal margin with a serrated longi¬ tudinal line or carina.15 14’ Metatibia beside dorsal margin without a serrated lon¬ gitudinal line or carina.17 15 Metafemur with a continuously serrated line adjacent to the anterior margin of metafemur. . Lasioserica Brenske 15’Metafemur without a continuously serrated line adja¬ cent to the anterior margin of metafemur.16 16 Serrated longitudinal line of metatibia ends about at half of metatibial length. Dorsal surface nearly glabrous. Neoserica (s.l.) probsti Ahrens 16’ Serrated longitudinal line of metatibia ends at apex of metatibia, its apical half directly situated on the dor¬ sal margin. Dorsal surface with dense double pilosity . Neoserica setigera (Brenske), N. subsetosa sp. n. 17 Parameres symmetrical.18 17’Parameres asymmetrical.19 18 Body small (< 6 mm). Phallobase without dorsal api¬ cal process. Oxyserica Brenske 18’ Body large (> 7 mm). Phallobase with long dorsal api¬ cal process. . Neoserica sladeni Ahrens, N. lenangensis Ahrens & Fabrizi 19 Antennal club in males long and reflexed. . Anomalophylla Reitter 19’Antennal club in males short or moderately long and straight.20 20 Elytra bicoloured, yellowish or reddish brown and black. Microserica Brenske 20’Elytra unicoloured.21 21 Apex of metatibia shallowly truncate at interior apex near tarsal articulation.22 21 ’ Apex of metatibia sharply truncate at interior apex near tarsal articulation.24 22 Dorsal surface yellowish brown to reddish brown, strongly and simply shiny. . Neoserica (s.l.) lubrica group 22’Dorsal surface dull or iridescently shiny.23 23 Antennal club with five antennomeres. . Neoserica (s.l.) garlangensis Ahrens 23’Antennal club with four antennomeres. . Neoserica (s.l .)pseudomajor sp. n. 24 Pronotum and elytra always nearly glabrous. . Sericania Motschulsky (see also couplet 36) ©ZFMK Sericini of India 7 24’Pronotum and elytra always distinctly setose. . Gynaecoserica Brenske 25 Labrum without a transverse rim of very dense, short and robust setae.26 25’Labrum short, with a transverse rim of very dense, short and robust setae. Dorsal surface densely setose. . [Neoserica pilosula group] 26 Metatibia slender and long.32 26’Metatibia short and wide.27 27 Body small (< 7.0 mm).28 27’Body large (> 7.5 mm), dorsal surface dark and dull/iridescently shiny.31 28 Dorsal surface light yellow to reddish brown, shiny 30 28’Dorsal surface dark brown, dull.29 29 Antennal club of male composed of six antennomere. . Neoserica multiflabellata Moser, N. keralana sp. n. 29’ Antennal club of male composed of five antennomeres. . Neoserica nathani Frey, N. agumbeensis sp. n., N. periyarensis sp. n., N quinquejlabellata (Brenske) 30 Distal tooth of protibia long and narrower Phallobase with a lateral apical apophysis at right side. . Neoserica quadrilamellata (Brenske) 30’Distal tooth of protibia short and wide. Phallobase without a lateral apical apophysis at right side. . Neoserica speciosa group 31 Labroclypeus strongly trapezoidal, its lateral margins in straight line with ocular canthus; base of labro¬ clypeus with dull toment; margins weakly reflexed... . Neoserica uniformis group 3 L Labroclypeus weakly trapezoidal, its lateral margins produced a blunt angle with ocular canthus; base of labroclypeus shiny; margins strongly reflexed. . Neoserica sexfoliata Moser, N. discipleensis sp. n. 32 Antennal club of males with seven antennomeres . ...33 32’ Antennal club of males with six or less antennomeres. .34 33 Metafemur with a continuously serrated line adjacent to the anterior margin of metafemur. Protibia before basal tooth sometimes bluntly (tooth-like) extended. . Neoserica septemlameUata group 33’Metafemur without a continuously serrated line adja¬ cent to the anterior margin of metafemur. Protibia al¬ ways distinctly bidentate. Nepaloserica Frey 34 Antennal club of males with six antennomeres. Base of labroclypeus dull.35 34’ Antennal club of males with five or four antennomeres. .36 35 Angle between basis of hypomeron and that of prono- tum strongly rounded, angle between surfaces of hy¬ pomeron and pronotum basally blunt. Hypomeron basally strongly produced ventrally and transversely sulcate. Lepidoserica Nikolaev 35’ Angle between basis of hypomeron and that of prono¬ tum sharp, angle between surfaces of hypomeron and pronotum sharp. Hypomeron basally not produced ventrally and not sulcate. Neoserica abnormis group 36 Apex of metatibia shallowly truncate at interior apex near tarsal articulation.40 36’ Apex of metatibia deeply truncate at interior apex near tarsal articulation. ...Sericania Motschulsky (see 14) 36” Apex of metatibia concavely sinuate.37 37 Body size >10 mm. Labroclypeus as long as wide... . Meriserica Brenske 37’ Body size < 8 mm. Labroclypeus wider than long. .. .38 38 Elytra generally bicoloured. In rare cases black, but then protarsal claws asymmetric. . Microsericaria Nikolaev 38’Elytra unicoloured. Protarsal claws always symmet¬ ric.39 39 Margins of labroclypeus moderately reflexed. . Deroserica Moser 39’Margins of labroclypeus very strongly reflexed. . Pachyderoserica Moser 40 Body surface dull. Body larger (8 mm).41 40’Body surface strongly shiny. Body smaller (5.7-6.6 mm).42 41 Body yellow, surface with dense dark spots. . Chrysoserica Brenske 41 ’ Body dark to reddish brown, unicoloured. . Neoserica matura Ahrens, N. rutilans Ahrens & Fabrizi 42 Antennal club composed of six antennomeres. . Neoserica speciosa group 42’Antennal club composed of five antennomeres. . Neosericapushkarensis sp. n., N. rajasthanica sp. n., N. sforziae sp. n. 43 Body extremely small (< 4.5 mm). Body distinctly bi¬ coloured. Anomioserica Arrow 43’Body larger (> 5 mm).44 44 Metafemur with a serrated line adjacent to the anteri¬ or margin.45 44’ Metafemur without serrated line adjacent to the ante¬ rior margin.55 45 Pronotum and elytra glabrous.46 45’Pronotum and elytra more or less densely setose. . Amiserica insperata group 46 Mesosternum between mesocoxae narrow.. . Serica (sensu lato) 46’Mesosternum between mesocoxae wide.47 47 Metatibia at least in basal third with a serrated line be¬ side the dorsal margin.48 47’ Metatibia without serrated line beside the dorsal mar¬ gin.53 48 Posterior longitudinal row of setae on metafemur re¬ duced. Maladera servitrita group 48’ Posterior longitudinal row of setae on metafemur pres¬ ent.49 49 Phallobase without apophysis.50 49’ Phallobase with a dorsal apophysis. Eyes smaller, ra- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Dirk Ahrens & Silvia Fabrizi tio diameter/interocular width ~0.6. . Maladera egregia group 50 Eyes large, ratio diameter/interocular width >0.8. . Maladera teinzoana group 50’Eyes smaller, ratio diameter/interocular width -0.6-0.7.51 51 Ocular canthus narrow and distinctly narrowed to¬ wards apex. M. songsakensis sp. n. 51 ’ Ocular canthus wide, indistinctly narrowed towards apex.52 52 Body larger (> 7.5 mm), dark. Anterior margin of labroclypeus straight. M. hauseri (Moser) 52’Body smaller (< 6.5 mm), reddish brown. Anterior margin of labroclypeus slightly sinuate medially. . M. rosettae (Frey) 53 Phallobase at apex strongly asymmetrical, insertion of left paramere strongly produced distally. . Maladera placida group (part 1) 53’ Phallobase at apex weakly asymmetrical, insertion of left paramere not produced distally.54 54 Parameres symmetrical. Metafemur shiny. . Maladera umbratica (Brenske) 54’Parameres asymmetrical. Metafemur dull. . Maladera lugubris group 55 Apex of metatibia sharply truncate at interior apex near tarsal articulation.56 55’Apex of metatibia bluntly truncate or concavely sin¬ uate at interior apex near tarsal articulation.58 56 Dorsal body surface strongly shiny.57 56’Dorsal body surface dull. Legs short and wide. . Maladera (subgenus Hemiserica ) 57 Dorsal body surface glabrous. Legs narrow and long, metatibia without serrated line beside dorsal margin. . Sericania (S. nepalensis (Frey) group) 57’Dorsal body surface densely finely setose. Legs mod¬ erately wide and long, metatibia with a fine serrated line beside dorsal margin in basal half of metatibia. . . M. senfti sp. n. 58 Dorsal posterior margin of metafemur serrated. ...59 58’Dorsal posterior margin of metafemur smooth.62 59 Anterior angles of pronotum not produced, obsolete. . Pachyserica 59’Anterior angles of pronotum produced, blunt to sharp.60 60 Hypomeron not carinate. Serica (s.str.) 60’Hypomeron carinate.61 61 Parameres symmetric or nearly symmetric. . Xenoserica 61’ Parameres strongly asymmetric. Serica (s.l.) 62 Parameres symmetric.63 62’Parameres asymmetric.65 63 Dorsal surface with dense erect pilosity, between these setae with fine adpressed short setae. Metatibia with a serrated longitudinal line beside the dorsal margin. . Maladera setosa (Brenske) 63’ Dorsal without double pilosity. Metatibia without ser¬ rated longitudinal line beside the dorsal margin. ..64 64 Parameres with distinct basal lobe. Protarsal claws of¬ ten asymmetric. Maladera indica group 64’ Parameres without distinct basal lobe. Protarsal claws symmetric. Maladara assamica group 65 Parameres setose (apically or basally).66 65’Parameres glabrous.67 66 Parameres often reduced in length, very narrow and hook-like. Median phallobase produced distally pro¬ ducing a sclerotised tube. . Maladera margined a group 66’ Parameres not reduced in length, not narrow or hook¬ like. Median phallobase not produced distally but con¬ cavely sinuate. M. trochaloides sp. n. 67 Parameres well-separated from phallobase.68 67’ Parameres basally entirely fused with the apical phal¬ lobase. Maladera fistulosa group 68 Phallobase at apex strongly asymmetrical, insertion of left paramere strongly produced distally. . Maladera placida group (part 2) 68’ Phallobase at apex weakly asymmetrical, insertion of left paramere not produced distally.69 68” Phallobase at apex strongly asymmetrical, insertion of right paramere strongly produced distally, phal¬ lobase before apex with a long lateral apophysis. . M. schoenwitzae sp. n. 69 Basal apodeme of phallobase shortened, corresponding to about 1/5 of phallobasal length. Phallobase tube-like extended.70 69’ Basal apodeme of phallobase not shortened, correspon¬ ding to about half of phallobasal length. Phallobase not tube-like extended.71 70 Phallobase distinctly belly-like convex ventrally, deeply sinuate dorsoapically between parameres. Pro¬ tarsal claws asymmetric. Anterior margin of labro¬ clypeus strongly tridentate. . Maladera quinquidens group 70’Phallobase at maximum moderately evenly convex ventrally, not distinctly sinuate dorsoapically between parameres. Protarsal claws symmetric. . Maladera (subg. Cephaloserica ) 71 Phallobase with large dorsal flattened apophysis. . Maladera schenklingi group 71’ Phallobase without dorsal apophysis.72 72 Parameres with basal appendages.73 72’Parameres without basal appendages.77 73 Basal appendages more or less wide, not filiform. 74 73’ Basal appendages very long and filiform, partly longer than phallobase. Body larger (> 7 mm). Parameres dis¬ tinctly asymmetric.76 74 Body small (4.5-5.5 mm).75 74’Body larger (> 8 mm). M. madurensis group 75 Distal portion of parameres nearly symmetric. Dorsal body surface dull. Maladera sikkimensis group Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 9 75’ Parameres strongly asymmetric. Phallobase at left side with a long lateral apophysis. Dorsal body surface shiny. Maladera punctulata (Frey) 76 Epipleural edge ending behind the anterior third of ely¬ tra. Maladera thomsoni group 76’Epipleural edge ending at the external apical angle of elytra. Maladera modestula group 77 Phallobase at apex lobe-like produced between para¬ meres. Maladera (subg. Omaladera Reitter) 77’Phallobase at apex concavely sinuate between para¬ meres.78 78 Base of labroclypeus dull, otherwise moderately shiny.79 78’Base of labroclypeus shiny.80 79 Phallobase apically at right side with a lateral, more or less pointed apophysis. Parameres well separated. Body size >10 mm. Maladera granuligera group 79’ Phallobase at right side without lateral apophysis. Para¬ meres fused with each other. Body small, < 7 mm. .. . Maladera polunini group 80 Labroclypeus longitudinally elevated medially, mar¬ gins strongly reflexed; its surface coarsely punctate, weakly shiny. Maladera drescheri (Moser) 80’Labroclypeus flat, margins weakly reflexed.81 81 Antennal club short and straight, at maximum little longer than remaining antennomeres combined. Labro¬ clypeus strongly shiny. Maladera ferruginea grovel Maladera significabilis 8 L Antennal club long and reflexed, more than twice as long as remaining antennomeres combined. Labro¬ clypeus moderately shiny. M geniculata sp. n. Serica MacLeay, 1819 Serica MacLeay, 1819: 146 (type species by monotypy: Serica brunnea (Linnaeus, 1758)); Ahrens 2004b: 12, 2005a: 1, 2007a: 450, 2007c: 32. Trichoserica Reitter, 1896: 181 (type species by original designation: Trichoserica fulvopubens Reitter, 1896); syn. by Nomura 1972: 110. Ophthalmoserica Brenske, 1897: 395 (type species by subsequent designation: Serica thibetana Brenske, 1897: 395; Ahrens 1999c); syn. by Nomura 1972: 110. Podoserica Breit, 1912: 202 (type species by monotypy: Podoserica reitteri Breit, 1912: 202); syn. by Ahrens 2005a: 3. Key to Serica species ( 88 ) 1 Hypomeron carinate. Serica (sensu lato) 1 ’ Hypomeron not carinate. Serica (sensu stricto) 2 2 Metastemum anteriorly abruptly and strongly elevat¬ ed. (S. velutina group) 3 2’ Metasternum anteriorly evenly elevated. . (S. brunnea group) 4 3 External lateral margin of protibia smooth. Phallobase dorsally without tubercles. S. velutina Arrow 3’ External lateral margin of protibia serrated. Phallobase dorsally with small tubercles. . S. pilumna Ahrens & Fabrizi 4 Frons shiny. Antenna with 9 or 10 antennomeres. ..5 4’ Frons at least in posterior half dull. Antenna with ten antennomeres.8 5 Antenna with ten antennomeres. Labroclypeus convex medially. External protarsal claw enlarged, its basal lobe small; interior protarsal claw very short, half as long as the external one, its basal tooth reduced. Prono- tum densely punctate. S. olivacea Brenske 5’ Antenna with nine antennomeres. Labroclypeus flat medially. Internal claw at least as long as the exter¬ nal.6 6 Body size 7.6-9.3 mm. Basal lobe of interior protarsal claw large, more than 3/4 of length of apical tooth. Frons sparsely punctate. S. pommeranzi Ahrens 6’ Body size 6.7-7.5 mm. Basal lobe of interior protarsal claw small, less than 3/4 of length of apical tooth. Frons densely punctate.7 7 Left paramere exceeds insertion of right paramere by 1/3 of its length. Right paramere bent dorsally (later¬ al view) . S. tongluana Ahrens 7’ Left paramere exceeds insertion of right paramere on¬ ly by 1/10 of its length. Right paramere always bent ventrally (lateral view) . S. eberti (Frey) 8 Right paramere apically strongly widened, dorsoven- trally flattened; its ventral membrane strongly sclero- tised. Left paramere apically curved externally.9 8’ Right paramere apically not strongly widened, if widened then always very long and dorsoventrally not flattened.17 9 Body size 6.5-7.5 mm.10 9’ Body size > 8.5 mm.11 10 Labroclypeus long, anterior margin deeply semicircu- larly sinuate medially, sharply dentate beside median sinuation. Dorsal surface with a few long erect setae. Right paramere without lateral filiform spine. Apical border of elytra without microtrichomes. . S. bidentata Ahrens 10’Labroclypeus wider and shorter, anterior margin weakly sinuate medially, bluntly dentate beside medi¬ an sinuation. Frons and pronotum with appressed se¬ tae. Right paramere with a lateral filiform spine. Api¬ cal border of elytra with a narrow rim of microtri¬ chomes. S. ramosa Ahrens 11 Parameres almost equal in length. Left paramere al¬ most straight, only at apex weakly curved externally. Dorsal surface with dense and moderately long erect setae. Antenna with nine or ten antennomeres. . S. kingdom Ahrens Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 10 Dirk Ahrens & Silvia Fabrizi 11 ’ Left paramere significantly shorter than the right. 12 12 Labroclypeus shortly transverse, slightly narrowed basally, anterior margin deeply and widely sinuate. Ab¬ dominal sternite five strongly elongate, almost twice as long as abdominal sternite four.13 12’Labroclypeus longer and trapezoidal, widest at base. Abdominal sternites four and five equal in length at the middle.15 13 Labroclypeus finely and densely punctate, with fine transverse wrinkles. Pronotum widest at base, lateral margins in posterior half weakly sinuate. . S. bhaktai Ahrens 13’ Labroclypeus finely and moderately densely punctate. Pronotum widest anterior to base, lateral margins al¬ most evenly convex.14 14 Right paramere strongly widened apically, significant¬ ly wider than the phallobase (dorsal view), apex deeply sinuate. Basal lobe of interior protarsal claw very large, almost as long as the apical tooth. . S. narya Ahrens 14’Right paramere strongly widened apically, but not wider than the phallobase (dorsal view), apical mar¬ gin convex. Basal lobe of interior protarsal claw three quarters as long as the apical tooth. . S. chasilakhae Ahrens 15 Right paramere spoon-like widened. Labroclypeus al¬ most not sinuate medially.16 15’Right paramere flattened dorsoventrally, widened, at apex not wider than at base. Labroclypeus deeply and widely sinuate. S. nepalensis (Frey) 16 Anterior margin of labroclypeus strongly reflexed me¬ dially. Antennal club twice as long as the remaining antennomeres. Disc of pronotum finely appressed se¬ tose, lateral margins straightly convergent from base to the middle. S. filitarsata Ahrens 16’Anterior margin of labroclypeus weakly reflexed me¬ dially. Antennal club 1.5 times as long as the remain¬ ing antennomeres. Disc of pronotum with moderate¬ ly long and erect setae which are bent backwards, lat¬ eral margins posteriorly weakly sinuate. . S. angustatotibialis Ahrens 17 Right paramere in the basal third with a fine lateral spine-like branch.18 17’ Right paramere in the basal third without a fine later¬ al spine-like branch, if present a lateral branch it is ro¬ bust and flattened inserting in the apical half of para¬ mere.28 18 Pronotum and elytra with long erect setae. Metatibia narrow, dorsally weakly carinate. . S. chuttana Ahrens 18’Pronotum and elytra with appressed setae.19 19 Metatibia dorsally longitudinally convex (also in basal third), lateral face smooth, impunctate, without super¬ ficial wrinkles.20 19’ Metatibia dorsally longitudinally carinate (also in basal third), lateral face strongly punctate or with wrinkles. .25 20 Anterior angles of labroclypeus strongly reflexed. ... .21 20’Anterior angles of labroclypeus moderately reflexed. Metatarsomere one 1/3 of its length longer than the dorsal metatibial spur.22 21 Metatarsomere one only slightly longer than the dor¬ sal metatibial spur. Left paramere distinctly shorter than the right one. S. almorae Ahrens 21’ Metatarsomere one 1/3 of its length longer than the dorsal metatibial spur. Left paramere subequal in length to the right one. S. weiperti Ahrens 22 Metatibia slender, ratio maximal with/length: 1/5.5-5.8. Labroclypeus narrow, almost as wide as long. Ocular canthus short (1/4 of ocular diameter). . S. bidigitata Ahrens 22’Metatibia slightly wider, ratio maximal with/length: 1/4.5-5.1. Labroclypeus wider than long. Ocular can¬ thus very short (1/10 of ocular diameter).23 23 Metatibia slender, ratio maximal with/length: 1/5.1... .24 23’Metatibia slightly wider, ratio maximal with/length: 1/4.5. S. ribbei Ahrens 24 Right paramere weakly curved. Dorsomedian incision between parameres deeper. S. jaegeri Ahrens 24’ Right paramere strongly curved. Dorsomedian incision between parameres more shallow. . . S. chautarana Ahrens 25 Antennal club 1.5 times as long as the remaining an¬ tennomeres combined. Body size 8.8-8.9 mm. . S. incognita Ahrens 25’Antennal club at least twice as long as the remaining antennomeres combined.26 26 Body size 7.6-8.5 mm. Labroclypeus superficially punctate and weakly transversely wrinkled. . S. mureensis Ahrens 26’Body size > 9.5 mm.27 27 Labroclypeus coarsely and densely punctate, trans¬ verse wrinkles almost not visible. Left paramere near¬ ly half as long as right one. S. tukucheana Ahrens 27’ Labroclypeus finely and densely punctate, with distinct transverse wrinkles. Left paramere as long as right one. . S. pelelaensis Ahrens & Fabrizi 28 Metatibia longitudinally convex dorsally, not carinate. . S. kumaonensis Ahrens 28’Metatibia strongly carinate dorsally.29 29’Antennal club little longer than the remaining anten¬ nomeres combined.30 29’Antennal club more than 1.5 times as long as the re¬ maining antennomeres combined.31 30 Antennal club 1.3 times as long as remaining anten¬ nomeres combined. Apex of right paramere straight. Left paramere evenly narrowed in apical third. . S. assequens Ahrens & Fabrizi Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 11 30’Antennal club 1.1 times as long as remaining anten- nomeres combined. Apex of right paramere curved in¬ teriorly. Left paramere behind base very narrow. . . S. guidoi Ahrens 31 Labroclypeus almost completely dull. Metatibia rela¬ tively short (ratio maximal width/length: 1/4.3). . S. opaciclypealis Ahrens 3 L Labroclypeus entirely shiny.32 32 Lateral face of metatibia with a few longitudinally im¬ pressed punctures, at maximum only on apex of metat¬ ibia with fine and short longitudinal wrinkles.33 32’ Lateral face of metatibia at least in dorsal portion with distinct longitudinal wrinkles.34 33 Pronotum narrow. Ocular canthus very short (1/6 of ocular diameter). Metatibia very slender (ratio maxi¬ mal with/length: 1/5.5). S. tropdeana Ahrens 33’ Pronotum more or less wide. Ocular canthus long (1/4 of ocular diameter). Metatibia slightly wider (ratio maximal with/length: 1/5). S. nebulosa Ahrens 34 Metatibia on lateral face also in ventral portion with longitudinal wrinkles.35 34’Metatibia on lateral face also in ventral portion with longitudinal punctures only.38 35 Ocular canthus shorter (1/4 of ocular diameter). Metat¬ ibia on lateral face with moderately dense and short longitudinal wrinkles. Metatarsomere one as long as the two following tarsomeres combined.36 35’Ocular canthus extremely long (1/2 of ocular diame¬ ter). Metatibia on medial and lateral face coarsely and densely wrinkled longitudinally. Metatarsomere one little shorter than the following two tarsomeres com¬ bined.37 36 Left paramere not widened in basal half. Right para¬ mere without blunt tooth mesoventrally. . S. thibetana Brenske 36’Left paramere widened in basal half. Right paramere with a blunt tooth mesoventrally. . S. rectidens Ahrens & Fabrizi 37 Phallobase and right paramere longer. Part of the dor- somedian incision at the apex of the phallobase not separated from the rest of phallobase by an incision (lateral view). S. proclivis Ahrens 37’ Phallobase and right paramere shorter. Part of the dor- somedian incision at the apex of the phallobase sepa¬ rated from the rest of phallobase by a small incision (lateral view). S. pigrans Ahrens & Fabrizi 38 Ocular canthus short (1/6 of ocular diameter). Anten¬ nal club almost twice as long as the remaining anten- nomeres combined. Labroclypeus between the trans¬ verse wrinkles superficially punctate only. . S. sherpa (Sabatinelli & Migliaccio) 38’ Ocular canthus longer (1/4 of ocular diameter). Anten¬ nal club at least twice as long as the remaining anten- nomeres combined. Labroclypeus finely and distinct¬ ly punctate.39 39 Protarsal claws symmetric, basal tooth of interior claw sharply truncate. Abdominal sternite five medially el¬ evated. S. lama Ahrens 39’Protarsal claws asymmetric, basal tooth of interior claw lobiform. Abdominal stemite five medially flat. .40 40 Punctation of labroclypeus deeply impressed. Metat¬ ibia slightly wider (ratio maximal width/length: 1/4.7). . S. khajiaris Mittal 40’ Punctation of labroclypeus shallowly impressed. Metatibia slender (ratio max. width/length: 1/5.1). 41 41 Right paramere at middle strongly curved ventrally (lateral view). Left paramere convexly widened exter¬ nally. S. khasiana (Moser) 41’ Right paramere nearly straight (lateral view). Left paramere not widened externally. . S. exhausta Ahrens & Fabrizi Serica velulina group Serica (s.str.) pilumna Ahrens & Fabrizi, 2009 Serica pilumna Ahrens & Fabrizi, 2009b: 253. Material examined. See Ahrens & Fabrizi 2009b (p. 253). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 3A-C, p. 279). Distribution. Endemic to western Arunachal Pradesh (Fig. 40E). Serica (s.str.) velutina Arrow, 1946 Serica velutina Arrow, 1946b: 6; Ahrens 2004b: 25. Material examined. See Ahrens 2004b (p. 25). Aedeagus. See Ahrens 2004b (figs 6-8, p. 381). Distribution. Endemic to northeastern Myanmar and As¬ sam (Mishmi Hills). Serica brunnea group Serica (s.str.) almorae Ahrens, 1999 Serica (s.str.) almorae Ahrens, 1999c: 280; Ahrens 2004b: 20, 2007a: 471. Material examined. See Ahrens 1999c (p. 280). Aedeagus. See Ahrens 1999c (figs 78-89, p. 221). Distribution. Endemic to western Himalaya, only known from the type locality (Pindar valley, Fig. 40C). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 12 Dirk Ahrens & Silvia Fabrizi Serica (s.str.) angustatotibialis Ahrens, 1999 Serica (s.str.) angustatotibialis Ahrens, 1999c: 303, 2004b: 23, 2007a: 471; Ahrens & Fabrizi 2011: 161. Material examined. See Ahrens 1999c (p. 303), 2004b (p. 23); Ahrens & Fabrizi 2011 (p. 161). Aedeagus. See Ahrens 1999c (figs 123-125, p. 226). Distribution. Endemic to Sikkim and the southern face of the Himalaya in southern Tibet between Sikkim and Bhutan (Fig. 40C). Serica (s.str.) assequens Ahrens & Fabrizi, 2009 Serica (s.str.) assequens Ahrens & Fabrizi, 2009b: 249. Material examined. See Ahrens & Fabrizi, 2009b (p. 249). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 2A-C, p. 278). Distribution. Endemic to western Arunachal Pradesh (Fig. 40B). Serica (s.str.) bhaktai Ahrens, 1999 Serica (s.str.) bhaktai Ahrens, 1999c: 294; Ahrens 2004b: 22, 2007a: 471. Material examined. See Ahrens 1999c (p. 294), 2004b (p. 22). Aedeagus. See Ahrens 1999c (figs 108-110, p. 224). Distribution. Endemic to central Nepal (Helambu und Lapchi Kang; Fig. 40C). Serica (s.str.) bidentata Ahrens, 1999 Serica (s.str.) bidentata Ahrens, 1999c: 292; Ahrens 2004b: 22, 2007a: 471. Material examined. See Ahrens 1999c (p. 292). Aedeagus. See Ahrens 1999c (figs 104-106, p. 223). Distribution. Endemic to central Nepal, so far known on¬ ly from the type locality in Helambu Himal (Fig. 40C). Serica (s.str.) bidigitata Ahrens, 2000 Serica (s.str.) bidigitata Ahrens, 2000d: 287; Ahrens 2004b: 19, 2007a: 471. Material examined. See Ahrens 2000d (p. 287), 2004b (p. 19). Aedeagus. See Ahrens 1999c (figs 1-3, p. 288). Distribution. Endemic to western central Nepal, so far known only from the type locality in Dhaulagiri Himal (Fig. 40C). Serica (s.str.) chasilakhae Ahrens, 1999 Serica (s.str.) chasilakhae Ahrens, 1999c: 296; Ahrens 2004b: 22, 2007a: 471. Material examined. See Ahrens 1999c (p. 296). Aedeagus. See Ahrens 1999c (figs 111-113, p. 224). Distribution. Endemic to Bhutan, known only from the type locality (Fig. 40A). Serica (s.str.) chautarana Ahrens, 2005 Serica (s.str.) chautarana Ahrens, 2005a: 122,2007a: 471. Material examined. See Ahrens 2005a (p. 122). Aedeagus. See Ahrens 2005a (fig. 58, p. 156). Distribution. Endemic to central Nepal. Serica (s.str.) chuttana Ahrens, 1999 Serica (s.str.) chuttana Ahrens, 1999c: 278; Ahrens 2004b: 20, 2007a: 471. Material examined. See Ahrens 1999c (p. 278), 2004b (p. 20). Aedeagus. See Ahrens 1999c (figs 84-86, p. 220). Distribution. Endemic to western Nepal (Fig. 40D). Serica (s.str.) eberti (Frey, 1965) (Figs 25B, 40B) Ophthalmoserica eberti Frey, 1965b: 90. Trichoserica eberti : Sabatinelli 1993: 630. Serica (s.str.) eberti. Ahrens 1999c: 310; 2004b: 23, 2005a: 120, 2006a: 409, 2007a: 471; Ahrens & Fabrizi 2009b: 270, 2011: 160; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 278), 2004b (p. 20), 2005a (p. 120), 2006a (p. 409); Ahrens & Fabrizi 2009b: 271, 2011 (p. 160); Shrestha et al. 2012 (p. 382); 8 ex. “India, Uttaranchal Auli, 3000m, 15.-17.VI.2003 N. Nishikawa leg.” (ZFMK). Aedeagus. See Ahrens 1999c (figs 132-139, p. 228). Distribution. From Kumaon Himalaya to eastern central Nepal as well as Bhutan; absent in eastern Nepal and the Sikkim/Darjeeling area (Fig. 40B). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 13 Serica (s.str.) exhausta Ahrens & Fabrizi, 2011 Serica (s.str.) exhausta Ahrens & Fabrizi, 2011: 141. Material examined. See Ahrens & Fabrizi 2011 (p. 140). Aedeagus. See Ahrens & Fabrizi 2011 (fig. 1E-G, p. 140). Distribution. Endemic to Bhutan (Fig. 40C). Serica (s.str.) filitarsata Ahrens, 1999 Serica (s.str.) filitarsata Ahrens, 1999c: 301; Ahrens 2004b: 22, 2007a: 471; Ahrens & Fabrizi 2011: 161. Material examined. See Ahrens 1999c (p. 301); Ahrens & Fabrizi 2011 (p. 161). Aedeagus. See Ahrens 1999c (figs 120-122, p. 226). Distribution. Bhutan, only known from the type locali¬ ty (Fig. 40A). Serica (s.str.) guidoi Ahrens, 1999 Serica (s.str.) guidoi Ahrens, 1999c: 268; Ahrens 2004b: 19, 2005a: 122, 2007a: 471; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 268), 2004b (p. 19), 2005a (p. 122); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999c (figs 69-71, p. 218). Distribution. From central Nepal to Sikkim, with a dis¬ junct occurrence in the Garhwal district (India, Kumaon Himalaya; Fig. 40A). Serica (s.str.) incognita Ahrens, 1999 Serica (s.str.) incognita Ahrens, 1999c: 282; Ahrens 2004b: 20, 2007a: 471. Material examined. See Ahrens 1999c (p. 282), 2004b (p. 20). Aedeagus. See Ahrens 1999c (figs 90-92, p. 221). Distribution. Bhutan to central Nepal, known from Rol- waling- and Helambu Himal as well as the Kali Gandaki valley (Fig. 40B). Serica (s.str.) jaegeri Ahrens, 1999 Serica (s.str.) jaegeri Ahrens, 1999c: 273; Ahrens 2004: 19, 2007a: 471. Material examined. See Ahrens 1999c (p. 273). Aedeagus. See Ahrens 1999c (figs 75-77, p. 221). Distribution. Endemic to central Nepal (Annapurna Hi¬ mal; Fig. 40C). Serica (s.str.) khajiaris Mittal, 1988 Serica khajiaris Mittal, 1988: 157; Ahrens 2004b: 17, 2005: 121, 2006a: 409, 2007a: 471. Serica (s.str.) khaijiaris [sic]: Ahrens 1999c: 262; Ahrens & Fabrizi 2009b: 270, 2011: 161; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 262), 2004b (p. 17), 2005a (p. 121), 2006a (p. 409); Ahrens & Fabrizi 2011 (p. 161); Shrestha et al. 2012 (p. 382); Sabatinelli & Ahrens 2015 (p. 142). Aedeagus. See Ahrens 1999c (figs 60-62, p. 216). Distribution. In the entire Himalaya (Fig. 40A). Serica (s.str.) khasiana (Moser, 1918) (Figs 25A, 40B) Ophthalmoserica khasiana Moser, 1918a: 210. Trichoserica khasiana. Sabatinelli 1993: 630. Serica (s.str.) khasiana. Ahrens 1999c: 265, 2004b: 18, 2005a: 121, 2006a: 409, 2007a: 471; Ahrens & Fabrizi 2011: 161. Material examined. See Ahrens 1999c (p. 265), 2004b (p. 18), 2005a (p. 121), 2006a (p. 409); Ahrens & Fabrizi 2011 (p. 161); 1 ex. “India, Uttaranchal Auli, 3000m, 15.- 17.VI.2003 N. Nishikawaleg” (ZFMK). Aedeagus. See Ahrens 1999c (figs 63-65, p. 217). Distribution. In the western Himalaya, eastward towards central Nepal, no recent material examined from the type locality (Khasia Hills), where its occurrence is therefore highly doubtful as the species is absent from the eastern Himalaya and east Nepal (Fig. 40B). Serica (s.str.) kingdom Ahrens, 1999 Serica (s.str.) kingdom Ahrens, 1999c: 300; Ahrens 2004b: 22, 2007a: 471; Ahrens & Fabrizi 2009b: 270; Ozgiil- Siemund & Ahrens 2015: 171. Material examined. See Ahrens 1999c (p. 300); Ahrens & Fabrizi 2009b (p. 270). Aedeagus. See Ahrens 1999c (figs 117-119, p. 225). Distribution. From the eastern Himalaya (Tibet) to Yun¬ nan (Fig. 40A). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 14 Dirk Ahrens & Silvia Fabrizi Serica (s.str.) kumaonensis Ahrens, 1999 Serica (s.str.) kumaonensis Ahrens, 1999c: 291; Ahrens 2004b: 21, 2007a: 471. Material examined. See Ahrens 1999c (p. 291). Aedeagus. See Ahrens 1999c (figs 101-103, p. 223). Distribution. Endemic to the Kumaon Himalaya (Fig. 40A). Serica (s.str.) lama Ahrens, 1999 Serica (s.str.) lama Ahrens, 1999c: 260; Ahrens 2004b: 17, 2007a: 471. Material examined. See Ahrens 1999c (p. 260), 2004b (p. 17). Aedeagus. See Ahrens 1999c (figs 57-59, p. 216). Distribution. Endemic to Sikkim (Fig. 40C). Serica (s.str.) mureensis Ahrens, 1999 Serica (s.str.) mureensis Ahrens, 1999c: 271; Ahrens 2004b: 19,2007a: 471; Ctegu 1-Siemund & Ahrens 2015: 171. Material examined. See Ahrens 1999c (p. 271), 2004b (p. 19). Aedeagus. See Ahrens 1999c (figs 72-74, p. 218). Distribution. Endemic to east Nepal and the Darjeeling district (Fig. 40C). Serica (s.str.) nary a Ahrens, 1999 Serica (s.str.) narya Ahrens, 1999c: 298; Ahrens 2004b: 22, 2007a: 471. Material examined. See Ahrens 1999c (p. 298), 2004b (p. 22). Aedeagus. See Ahrens 1999c (figs 114-116, p. 225). Distribution. Endemic to the Darjeeling district (Fig. 40C). Serica (s.str.) nebulosa Ahrens, 1999 Serica (s.str.) nebulosa Ahrens, 1999c: 259; Ahrens 2004b: 17, 2007a: 471. Material examined. See Ahrens 1999c (p. 259), 2004b (p. 17). Aedeagus. See Ahrens 1999c (figs 114-116, p. 225). Distribution. Endemic to the Darjeeling district (Fig. 40D). Serica (s.str.) nepalensis (Frey, 1969) Ophthalmoserica nepalensis Frey, 1969a: 519. Trichoserica nepalensis: Sabatinelli 1993: 630. Serica (s.str.) nepalensis: Ahrens 1999c: 286, 2004b: 21, 2005a: 121, 2006a: 410, 2007a: 471; Ahrens & Fabrizi 2009b: 270, 2011: 160; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 286), 2004b (p. 21), 2005a (p. 121), 2006a (p. 410); Ahrens & Fabrizi 2009b (p. 270), 2011 (p. 160); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999c (figs 96, 97, p. 222). Distribution. Endemic to central and east Nepal as well as southern Tibet (Fig. 40D). Serica (s.str.) olivacea Brenske, 1896 Serica olivacea Brenske, 1896: 155; Brenske 1898: 289; Ahrens 1999c: 243, 2004b: 23, 2007a: 471. Trichoserica olivacea'. Sabatinelli 1993: 630. Material examined. See Ahrens 1999c (p. 243). Aedeagus. See Ahrens 1999c (figs 36, 37, p. 212). Distribution. Endemic to Sikkim and the Darjeeling area (Fig. 40B). Serica (s.str.) opaciclypealis Ahrens, 1999 Serica (s.str.) opaciclypealis Ahrens, 1999c: 253; Ahrens 2004b: 17, 2007a: 471. Material examined. See Ahrens 1999c (p. 253). Aedeagus. See Ahrens 1999c (figs 47^19, p. 214). Distribution. Endemic to eastern Nepal and western Sikkim (Fig. 40B). Serica (s.str.) pelelaensis Ahrens & Fabrizi, 2011 Serica (s.str.) pelelaensis Ahrens & Fabrizi, 2011: 140. Material examined. See Ahrens & Fabrizi 2011 (p. 140). Aedeagus. See Ahrens & Fabrizi 2011 (fig. 1A-C, p. 140). Distribution. Endemic to Bhutan (Fig. 40B). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 15 Serica (s.str.) pigrans Ahrens & Fabrizi, 2009 Serica (s.str.) pigrans Ahrens & Fabrizi, 2009b: 251 Material examined. See Ahrens & Fabrizi 2009b (p. 251). Aedeagus. Ahrens & Fabrizi 2009b (fig. 2G-I, p. 278). Distribution. Endemic to western Arunachal Pradesh (Fig. 40A). Serica (s.str.) pommeranzi Ahrens, 1999 Serica (s.str.) pommeranzi Ahrens, 1999c: 305; Ahrens 2004b: 23, 2007a: 471; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 305), 2004b (p. 23); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999c (figs 126-128, p. 227). Distribution. Endemic to western Nepal (Fig. 40C). Serica (s.str.) proclivis Ahrens, 1999 Serica (s.str.) proclivis Ahrens, 1999c: 257; Ahrens 2004b: 17, 2007a: 471. Material examined. See Ahrens 1999c (p. 257). Aedeagus. See Ahrens 1999c (figs 50-52, p. 215). Distribution. Endemic to eastern Nepal, only known from the type locality (Fig. 40B). Serica (s.str.) ramosa Ahrens, 1999 Serica (s.str.) ramosa Ahrens, 1999c: 284; Ahrens 2004b: 21, 2006a: 410, 2007a: 471; Ahrens & Fabrizi 2009b: 270. Material examined. See Ahrens 1999c (p. 284), 2004b (p. 21), 2006a (p. 410); Ahrens & Fabrizi 2009b (p. 270). Aedeagus. See Ahrens 1999c (figs 93-95, p. 222). Distribution. Endemic to western central Nepal (Anna- puma Himal; Fig. 40D). Serica (s.str.) rectidens Ahrens & Fabrizi, 2009 Serica (s.str.) rectidens Ahrens & Fabrizi, 2009: 250; Ahrens & Fabrizi 2011: 160. Material examined. See Ahrens & Fabrizi 2009 (p. 250), 2011 (p. 160). Aedeagus. See Ahrens & Fabrizi 2009 (fig. 2D-F, p. 278). Distribution. Endemic to southern Tibet and Bhutan (Fig. 40D). Serica (s.str.) ribbei Ahrens, 1999 Serica (s.str.) ribbei Ahrens, 1999c: 275; Ahrens 2004b: 20, 2007a: 471. Material examined. See Ahrens 1999c (p. 275). Aedeagus. See Ahrens 1999c (figs 93-95, p. 222). Distribution. Endemic to western central Nepal (Anna¬ purna Himal; Fig. 40B). Serica (s.str.) sherpa (Sabatinelli & Migliaccio, 1982) Trichoserica sherpa Sabatinelli & Migliaccio, 1982: 106; Sabatinelli 1993: 630. Serica (s.str.) sherpa: Ahrens 1999c: 251, 2004b: 16, 2007a: 471. Material examined. See Ahrens 1999c (p. 251), 2004b (p. 16). Aedeagus. See Ahrens 1999c (figs 41^43, p. 213). Distribution. Endemic to southern Tibet, eastern central Nepal and eastern Nepal (Fig. 40B). Serica (s.str.) thibetana Brenske, 1897 Serica ( Ophthalmoserica ) thibetana Brenske, 1897: 395. Trichoserica thibetana: Sabatinelli 1993: 631. Serica (s.str.) thibetana: Ahrens 1999c: 245, 2004b: 15, 2005a: 75, 2006a: 410, 2007a: 471; Ahrens & Fabrizi 2011: 160; Shrestha et al. 2012: 382; Ozgul-Siemund & Ahrens 2015: 171. Serica {Ophthalmoserica) umbrinella Brenske, 1898: 288; Ahrens 1999c: 245. Trichoserica umbrinella: Khan & Ghai 1978: 214; Sabatinelli 1993: 631. Serica umbrina Frey 1965b (non Blanchard 1850): 90. Material examined. See Ahrens 1999c (p. 245), 2004b (p. 15), 2005a (p. 75), 2006a (p. 410); Ahrens & Fabrizi 2009b (p. 270), 2011 (p. 160); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999c (figs 38^10, p. 213). Distribution. The wide range extends over the entire Hi¬ malaya and eastern Tibet (including Yunnan) (Fig. 40D). Serica (s.str.) tongluana Ahrens, 1999 Serica (s.str.) tongluana Ahrens, 1999c: 308; Ahrens 2004b: 24, 2005a: 121, 2007a: 471. Material examined. See Ahrens 1999c (p. 308), 2004b (p. 24), 2005a (p. 121). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 16 Dirk Ahrens & Silvia Fabrizi Aedeagus. See Ahrens 1999c (figs 129-131, p. 227). Distribution. Restricted to eastern Nepal and the Sikkim/Darjeeling area (Fig. 40C). Serica (s.str.) tropdeana Ahrens, 1999 Serica (s.str.) tropdeana Ahrens, 1999: 255; Ahrens 2004b: 17, 2005a: 120, 2007a: 471. Material examined. See Ahrens 1999c (p. 255), 2004b (p. 24), 2005a (p. 120). Aedeagus. See Ahrens 1999c (figs 44^16, p. 214). Distribution. Restricted to southern Tibet, so far known only from the type locality (Fig. 40C). Serica (s.str.) tukucheana Ahrens, 1999 Serica (s.str.) tukucheana Ahrens, 1999c: 276; Ahrens 2004b: 20,2005a: 122,2006a: 410,2007a: 471; Qzgtil- Siemund & Ahrens 2015: 171. Material examined. See Ahrens 1999c (p. 276), 2004b (p. 20), 2005a (p. 122), 2006a (p. 410). Aedeagus. See Ahrens 1999c (figs 81-83, p. 220). Distribution. From western central Nepal to Darjeeling/southern Tibet (Fig. 40A). Serica (s.str.) weiperti Ahrens, 2004 Serica (s.str.) weiperti Ahrens, 2004a: 211; Ahrens 2004b: 20, 2007a: 471. Material examined. See Ahrens 2004a (p. 211), 2004b (p. 20). Aedeagus. See Ahrens 2004a (figs 1-3, p. 212). Distribution. Endemic to western Nepal (Fig. 40A). Serica (sensu lato) Key to Serica (s.l.) species 1 Anterior margin of metafemur without serrated line. .3 1 ’ Anterior margin of metafemur with a serrated line. 2 2 Body larger (length: 9.1-10.0 mm). Labroclypeus wide and trapezoidal, lateral margins strongly convex. In¬ tervals of elytra densely and coarsely punctate. Metat¬ ibia in basal half with a longitudinal serrated line. . S. panchaseana Ahrens T Body larger (length: 7.7-8.0 mm). Labroclypeus mod¬ erately wide, lateral margins weakly convex. Intervals Bonn zoological Bulletin 65 (1 & 2): 1-355 of elytra sparsely punctate. Metatibia without serrat¬ ed line. S. palaea Ahrens 3 Dorsal surface densely setose.4 3 ’ Dorsal surface nearly glabrous or only sparsely setose. .5 4 Phallobase dorsally before apex without a median el¬ evation. Parameres straight. S. velutina Arrow 4’ Phallobase dorsally before apex with a median eleva¬ tion. Parameres curved ventrally. . S. pilumna Ahrens & Fabrizi 5 Main body colour yellowish brown with numerous dark spots. Lateral margin of basal half of pronotum deeply sinuate.6 5 ’ Main body colour dark or reddish brown. Lateral mar¬ gin of basal half of pronotum convex or straight. ...7 6 Sixth antennomeres as long as antennomeres three to five combined. Lateral sinuation of pronotum deeper. Left paramere bifurcate at apex. . S. sticta Ahrens & Fabrizi 6’ Sixth antennomeres slightly shorter than antennomeres three to five combined. Lateral sinuation of pronotum shallower. Left paramere simply pointed at apex. . S. arborea Ahrens 7 Dorsal lobe of left paramere sharply pointed at apex. . S.fashengi Liu & Ahrens, 2014 7’ Dorsal lobe of left paramere rounded at apex.8 8 Dorsal lateral lamina of phallobase interiorly only very weakly widened. Dorsal lobe of left paramere round¬ ed at apex. Dorsal lobe of left paramere bent external¬ ly at middle, only little widened. . S. falcifera Ahrens & Fabrizi 8’ Dorsal lateral lamina of phallobase interiorly with a triangular lamella that extends to the middle of phal¬ lobase. Dorsal lobe of left paramere nearly straight but strongly widened. S. feresegregata sp. n. Serica (s.l.) arborea Ahrens, 1999 (Figs 25C, 40E) Serica (s.str.) arborea Ahrens, 1999c: 210. Serica (s.l.) arborea. Ahrens 2004b: 24, 2007a: 471, 2006a: 409. Material examined. See Ahrens 1999c (p. 210), 2004b (p. 24), 2006a (p. 409). Aedeagus. See Ahrens 1999c (figs 31-33, p. 212). Distribution. Endemic to central Nepal (Fig. 40E). Serica (s.l.) falcifera Ahrens & Fabrizi, 2009 Serica (s.l.) falcifera Ahrens & Fabrizi, 2009b: 254. Material examined. See Ahrens & Fabrizi 2009b (p. 254). ©ZFMK Sericini of India 17 Aedeagus. See Ahrens & Fabrizi 2009b (fig. 3G-I, p. 279). Distribution. Endemic to western Arunachal Pradesh (Fig. 40E). Serica (s.l.) fashengi Liu & Ahrens, 2014 Serica (s.l.) fashengi Liu & Ahrens, 2014: 167. Material examined. See Liu et al. 2014a (p. 167). Aedeagus. See Liu et al. 2014a (fig. 3E-H, p. 166). Distribution. Endemic to eastern Tibet (Fig. 40E). Serica (s.l.) feresegregata sp. n. (Figs 2A-D, 25E, 40E) Type material examined. Holotype: S “NE India Arunachal Pr. Hunli vicinity, 1300±100m 28°19 , 32”N 95°57 , 31 ,, E, L. Dembicky leg., 26.v.-l.vi.2012” (ZFMK). Paratype: 1 S “X-DA3456 - India, Arunachal Pradesh, Roing, 490m, 28°08 , 32 ,, N 95°51\ 2.-5.vi.2012, leg. L. Dembicky” (ZFMK). Description. Length: 8.8 mm, length of elytra: 7.3 mm, width: 5.9 mm. Body oval, dark brown, antenna yellow¬ ish brown, legs reddish brown, dorsal surface dull, labro- clypeus and anterior quarter of frons shiny, head sparse¬ ly setose, dorsal face of pronotum and elytra glabrous. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anteriorly very weakly sinuate medially, margins moder¬ ately reflexed; surface flat and shiny, moderately and densely punctate, with a transverse row of few erect se¬ tae behind anterior margin; frontoclypeal suture indistinct¬ ly incised, not elevated and slightly angled medially; smooth area anterior to eye wide, nearly flat, approximate¬ ly 2.5 times as wide as long; ocular canthus moderately long and broad (1/3 of ocular diameter), finely and sparse¬ ly punctate, with one long terminal seta. Frons dull, part¬ ly narrowly shiny only immediately behind the fronto¬ clypeal suture, with fine, dense punctures on anterior shiny portion, punctation nearly invisible under dull tomenta- tion, with numerous erect setae on posterior half. Eyes moderately large, ratio diameter/interocular width: 0.69. Antenna with ten antennomeres; antennomeres three to seven distinctly wider than long, club with three anten¬ nomeres, three times as long as the remaining anten¬ nomeres combined and strongly reflexed. Mentum elevat¬ ed and slightly flattened anteriorly. Labrum transverse, short, not produced medially, without median sinuation. Pronotum transverse, widest at base, lateral margins convex and slightly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and slightly rounded at tip; anterior margin with a medially widely interrupted marginal line, convexly produced me¬ dially; surface densely and finely punctate, punctures on sides with very minute setae; anterior and lateral border glabrous, around the anterior angles sparsely setose; hy- pomeron basally produced and distinctly carinate. Scutel- lum slender and long, triangular, with fine, dense punc¬ tures, glabrous. Elytra oblong, widest just behind middle, striae mod¬ erately impressed, finely and densely punctate, intervals nearly flat, with fine, dense punctures concentrated along striae, completely glabrous; epipleural edge moderately ro¬ bust, ending at moderately curved external apical angle of elytra, epipleura densely setose, apical border with fine fringe of microtrichomes (visible at ca lOOx magnifica¬ tion). Ventral surface dull, finely and not densely punctate, nearly glabrous, metacoxa with a few short robust setae laterally; abdominal sternites finely and densely punctu¬ ate, with a transverse row of coarse punctures, each bear¬ ing a short seta. Mesosternum between mesocoxae half as wide as the slender mesofemur. Ratio of length of metepis- ternum/metacoxa: 1/1.35. Pygidium strongly convex at apex and dull, finely and densely punctate, without smooth midline, with sparse short setae that are partly erect. Legs moderately slender; femora with two longitudinal rows of setae, finely and sparsely punctate between the rows; metafemur shiny, anterior margin acute, behind an¬ terior edge with a continuously serrated undulating line, posterior margin in apical half ventrally smooth and on¬ ly weakly widened, posterior margin dorsally also smooth, on its entire length with dense short setae. Metat¬ ibia moderately slender and not very long, widest at apex, ratio of width/length: 1/3.6, dorsally moderately carinate, with two groups of spines, basal one just before middle, apical group at three quarters of metatibial length, in basal third with a fine serrated line bearing a few fine single se¬ tae; outside longitudinally convex, finely and moderate¬ ly dense punctate, without wrinkles; ventral edge finely serrated, with four equidistant setae, medial face smooth and impunctate, apex interiorly near tarsal articulation shallowly truncate. Tarsomeres smooth, neither laterally nor dorsally carinate; pro- and mesotarsomeres ventrally with sparse, short setae; metatarsomeres ventrally glabrous, ventrally with a strongly serrated ridge, smooth, first metatarsomere little shorter than the two following tarsomeres combined and one third of its length longer than the upper tibial spur. Protibia moderately long, biden- tate; anterior claws symmetrical, basal tooth of inner claw truncate at apex. Aedeagus: Fig. 2A-D. Habitus: 25E. Female un¬ known. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 18 Dirk Ahrens & Silvia Fabrizi Diagnosis. Serica feresegregata sp. n. is very similar to S. segregata Arrow, 1946 and S. falcifera Ahrens & Fab¬ rizi, 2009 in its external and genital morphology. The new taxon differs from the two previously mentioned species in the slightly shorter antennal club, but significantly on¬ ly in the shape of the aedeagus: the dorsal lateral lamina of the phallobase has interiorly a triangular lamella that extends to the middle of the phallobase, in the former two species this lamina is absent; furthermore, in S. fereseg¬ regata the dorsal lobe of the left paramere is nearly straight but strongly widened, while it is narrower and bent ex¬ ternally at middle in the other two species. Variation. Length: 8.8-9.6 mm, length of elytra: 7.2-7.3 mm, width: 5.0-5.9 mm. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the combined Latin words fere- (nearly) and the species name ‘ segregata ’, with ref¬ erence to the similarity to S. segregata Arrow. Distribution. So far known only from Arunachal Pradesh (Fig. 40E). Serica (s.l.) palaea Ahrens, 2004 Serica (s.l.) palaea Ahrens, 2004b: 26, 2007a: 471. Material examined. See Ahrens 2004b (p. 26). Aedeagus. See Ahrens 2004b (figs 9-11, p. 381). Distribution. Endemic to Sikkim (Fig. 40E). Serica (s.l.) panchaseana Ahrens, 2004 (Figs 25D, 40E) Serica (s.l.) panchaseana Ahrens, 2004b: 28,2006a: 409, 2007a: 471. Material examined. See Ahrens 2004b (p. 28), 2006a (p. 409). Aedeagus. See Ahrens 2004b (figs 12-14, p. 382). Distribution. Endemic to central Nepal (Annapurna mountains) (Fig. 40E). Serica (s.l.) sticta Ahrens & Fabrizi, 2009 Serica sticta Ahrens & Fabrizi, 2009b: 253; Ahrens & Fab¬ rizi 2011: 161. Material examined. See Ahrens & Fabrizi 2009b (p. 253); Ahrens & Fabrizi 2011 (p. 161). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 2P-R, p. 278). Distribution. Endemic to western Arunachal Pradesh and Bhutan (Fig. 40E). P achy serica Brenske, 1897 Pachyserica Brenske, 1897: 355; Ahrens 2004b: 30, 2006b: 487,2007c: 30 (type species by subsequent des¬ ignation: Pachyserica rubrobasalis Brenske, 1897; Yu etal. 1998). Key to Pachyserica species ( ff ) 1 Hypomeron not produced ventrally. Long and erect se¬ tae of elytra scale-like.2 1 ’ Hypomeron strongly produced ventrally.3 2 External margin of protibia smooth. Mesotarsomeres punctate dorsally. P. marmorata (Blanchard) T External margin of protibia distinctly serrate. Meso¬ tarsomeres impunctate dorsally. . P. pellingensis Ahrens 3 Long erect setae on elytra numerous (at least 32 per interval).4 3’ Long erect setae on elytra sparser (at most 20 per in¬ terval).11 4 Metacoxa enlarged (ratio length metepistemum/meta- coxa > 1/1.7). Mesotarsomeres impunctate dorsally. .5 4’ Metacoxa not enlarged (ratio length metepisternum/ metacoxa: 1/1.23-1.6).14 5 Right paramere without basal lobe.6 5 ’ Right paramere with a basal lobe being directed inter¬ nally.7 6 Metatibia moderately long, widened at middle, ratio metatibial width/length: 1/3.3. . P. himalayensis Ahrens 6’ Metatibia long, ratio metatibial width/length < 1/3.5. . P. albosqiiamosa Brenske 7 Ratio metatibial width/length: 1/3.5-3.8.8 7’ Ratio metatibial width/length < 1/3.9.9 8 Pronotum wide. Left paramere long. . P. gracilis Ahrens 8’ Pronotum narrow. Left paramere short. . P. numensis Ahrens 9 Antennal club slightly longer or as long as remaining antennomeres combined. P. olafi Ahrens 9’ Antennal club distinctly longer (1.3 times) than re¬ maining antennomeres combined.10 10 Right paramere only little longer than the apical phal¬ lobase. Left paramere with a blunt lateral tooth at apex. . P. nepalica Ahrens 10’ Right paramere 1.5 times as long as the apical phal- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 19 lobase. Left paramere without blunt lateral tooth at apex. P ambiversa Ahrens 11 Base of labroclypeus shiny, without dull toment. .12 11 ’ Base of labroclypeus with dull toment. Pilosity of metatibia composed also of scales.13 12 Phallobase dorsally before apex without tubercles. ... . P. stabilis Ahrens 12’ Phallobase dorsally before apex with two tubercles. . P. bituberculata Ahrens 13 Phallobase ventroapically medially produced. . P. garoensis Ahrens 13’ Phallobase ventroapically medially concavely sinuate. . P. jendeki Ahrens 14 Apical margin of elytra with a rim of microtrichomes. .15 14’Apical margin of elytra without a rim of microtri¬ chomes. Anterior angles of pronotum strongly convex. (Myanmar) . [P. interruptolineata Ahrens] 15 Anterior margin of labroclypeus weakly sinuate me¬ dially. P. collaris Ahrens 15’Anterior margin of labroclypeus straight, not sinuate medially. P. darjeelingensis Ahrens Pachyserica albosquamosa Brenske, 1898 Pachyserica albosquamosa Brenske, 1898: 356; Ahrens 2004b: 30, 2006: 498; Ahrens & Fabrizi 2009b: 271; Ozgiil-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 30), Ahrens & Fabrizi 2009b (p. 271); 1 ex. “India: West Bengalen Dis- tr. Darjeeling Kurseong: Thingay Bari, 15.VI.1991 800 m NN, leg. N. Dangal” (NME), 1 0 “NE India, Meghalaya, SW of Cherrapunjee, 25°13 , -14”N, 91°40’E, 5.- 24.V.2005, 900m, P. Pacholatko leg.” (CPPB), 1 $ “NE India, Meghalaya, SW of Cherrapunjee, 25°13’-15 , N, 91°47’E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB). Aedeagus. See Ahrens 2004b (figs 15-17, p. 382). Distribution. Some single records from the Garhwal Hi¬ malaya, Nepal, Assam and the Khasi Hills (Fig. 40F). Pachyserica ambiversa Ahrens, 2004 Pachyserica ambiversa Ahrens, 2004b: 43; Ahrens 2006a: 410,2006b: 498; Ozgiil-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 43), 2006a (p. 410). Aedeagus. See Ahrens 2004b (figs 34-36, p. 385). Distribution. Central Nepal (Fig. 41 A). Pachyserica bituberculata Ahrens, 2006 Pachyserica bituberculata Ahrens, 2006b: 525. Material examined. See Ahrens 2006b (p. 525). Aedeagus. See Ahrens 2006b (fig. 12A-C, p. 528). Distribution. Meghalaya area and upper Myanmar (Fig. 40F). Pachyserica collaris Ahrens, 2006 Pachyserica collaris Ahrens, 2006b: 527; Ahrens & Fab¬ rizi 2011: 161. Material examined. See Ahrens 2006b (p. 527); Ahrens & Fabrizi 2011 (p. 161); 1 S “India: 26.VI.95 Cherrapun¬ jee Meghalaya, Werner leg.” (ZFMK). Aedeagus. See Ahrens 2006b (fig. 12D-F, p. 528). Distribution. Assam and the Meghalaya area (Fig. 40F). Pachyserica darjeelingensis Ahrens, 2004 (Fig. 251) Pachyserica darjeelingensis Ahrens, 2004b: 51; Ahrens 2006b: 498. Material examined. See Ahrens 2004b (p. 51). Aedeagus. See Ahrens 2004b (figs 48-51, p. 387). Distribution. Eastern Nepal and the Sikkim/Darjeeling area (Fig. 41A). Pachyserica garoensis Ahrens, 2006 Pachyserica garoensis Ahrens, 2006b: 541. Material examined. See Ahrens 2006b (p. 541). Aedeagus. See Ahrens 2006b (fig. 15D-F, p. 542). Distribution. Meghalaya area and upper Myanmar (Fig. 40F). Pachyserica gracilis Ahrens, 2004 Pachyserica gracilis Ahrens, 2004b: 35; Ahrens 2006b: 498; Shrestha et al. 2012: 382; Ozgiil-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 35); Shrestha et al. 2012 (p. 382); 1 “Coll. R.I.Sc.N.B./ India: Hi¬ malaya Bhim Valley S. Breuning vendit” (ISNB). Aedeagus. See Ahrens 2004b (figs 21-23, p. 383). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 20 Dirk Ahrens & Silvia Fabrizi Distribution. Kumaon and Nepal Himalaya including the Darjeeling area (Fig. 41A). Pachyserica himalayensis Ahrens, 2004 Pachyserica himalayensis Ahrens, 2004b: 37; Ahrens 2006b: 498; Shrestha et al. 2012: 382; Ozgul-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 37); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 2004b (figs 24-26, p. 384). Distribution. Eastern Nepal to Bhutan (Fig. 41 A). Pachyserica jendeki Ahrens, 2004 Pachyserica jendeki Ahrens, 2004b: 45; Ahrens 2006a: 410,2006b: 498; Qzgul-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 45), 2006a (p. 410); 1 S “NE India, Meghalaya, 8km N of Shillong, 25°38‘ N, 91°54‘E, 1200m, L. Dembicky leg., 7.- 9.V.2004” (CPPB), 1 & 1 $ “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-15‘N, 91°47‘E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB), 1 S “NE India, Meghalaya, SW of Shillong, 1600m, 25°34‘N, 91°51‘E, L. Dembicky leg., 14.V.2004” (CPPB), 1 $ “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘N, 91°40‘E, 5.-24.V.2005, 900m, P. Pacholatko leg.” (CPPB). Aedeagus. See Ahrens 2004b (figs 37^10, p. 386). Distribution. India (Darjeeling, Meghalaya, Assam), northern Myanmar, and northern Thailand (Fig. 40F, see also Ahrens 2004). Pachyserica marmorata (Blanchard, 1850) (Fig. 25F) Omaloplia marmorata Blanchard, 1850: 77. Pachyserica marmorata: Brenske 1898: 233; Sabatinelli & Migliaccio 1982: 104; Ahrens 2004b: 48,2006a: 410, 2006b: 498; Ahrens & Fabrizi 2009b: 270; Shrestha et al. 2012: 382; Ozgul-Siemund & Ahrens 2015: 172. Serica marmorata: Barlow 1899: 244. Material examined. See Ahrens 2004b (p. 48), 2006a (p. 410); Ahrens & Fabrizi 2009b (p. 271); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 2004b (figs 44^17, p. 387). Distribution. Kumaon and Nepal Himalaya (Fig. 41 A). Pachyserica nepalica Ahrens, 2004 Pachyserica nepalica Ahrens, 2004b: 39; Ahrens 2006b: 498; Shrestha et al. 2012: 382; Ozgul-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 39); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 2004b (figs 27-29, p. 384). Distribution. Central and eastern Nepal Himalaya (Fig. 41A). Pachyserica numensis Ahrens, 2004 Pachyserica numensis Ahrens, 2004b: 41; Ahrens 2006b: 498. Material examined. See Ahrens 2004b (p. 41). Aedeagus. See Ahrens 2004b (figs 30-33, p. 385). Distribution. Eastern Nepal Himalaya (Fig. 41A). Pachyserica olaji Ahrens, 2004 (Fig. 25H) Pachyserica olafi Ahrens, 2004b: 32; Ahrens 2006a: 410, 2006b: 498; Ahrens & Fabrizi 2011: 161; Ahrens & Fab¬ rizi 2011: 161; Shrestha et al. 2012: 382; Ozgul-Siemu¬ nd & Ahrens 2015: 171. Material examined. See Ahrens 2004b (p. 32), 2006a (p. 410); Ahrens & Fabrizi 2011 (p. 161); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 2004b (figs 18-20, p. 383). Distribution. Central and eastern Nepal Himalaya (Fig. 40F). Pachyserica pellingensis Ahrens, 2004 (Fig. 25G) Pachyserica pellingensis Ahrens, 2004b: 52; Ahrens 2006b: 498; Shrestha et al. 2012: 382. Material examined. See Ahrens 2004b (p. 52); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 2004b (figs 52-54, p. 388). Distribution. Restricted to Sikkim and eastern Nepal (Fig. 41A). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 21 Pachyserica stabilis Ahrens, 2004 Pachyserica stabilis Ahrens, 2004b: 47; Ahrens 2006b: 498. Material examined. See Ahrens 2004b (p. 47). Aedeagus. See Ahrens 2004b (figs 41^13, p. 386). Distribution. Restricted to the Darjeeling area (Fig. 40F). Calios erica Brenske, 1894 Calloserica Brenske, 1894: 10, 56; (type species by mono- typy: Calloserica tigrina Brenske, 1894); Ahrens 1995a: 137; 1999b: 185; 2000a: 815; 2004b: 54; 2005b: 217, 2007c: 12. Key to Calloserica species (88) 1 Anterior margin of labroclypeus weakly sinuate me¬ dially.2 1 ’ Anterior margin of labroclypeus deeply sinuate medi¬ ally.5 2 Dorsal margin of metatibia longitudinally convex, not carinate. C. barabiseana Ahrens 2’ Dorsal margin of metatibia moderately longitudinal¬ ly carinate.3 3 Penultimate abdominal sternite with robust, distinct¬ ly serrate transverse elevation. . C. rupthangensis Ahrens 3’ Penultimate abdominal sternite without elevations. 4 4 Antennal club only slightly longer than remaining an- tennomeres combined. C. raksensis Ahrens 4’ Antennal club 1.5 times as long as remaining anten- nomeres combined. C. gosainkundensis Ahrens 5 Posterior margin of penultimate abdominal sternite with two transverse elevations.6 5’ Posterior margin of penultimate abdominal sternite without elevations.9 6 Elevations on penultimate abdominal sternite sharply pointed, slightly bent anteriorly; last abdominal ster¬ nite produced ventrally and with a transverse carina. Setae on dorsal portion of left paramere lacking. ...7 6’ Elevations on penultimate abdominal sternite blunt, not bent anteriorly; last abdominal sternite simple, not pro¬ duced ventrally and without carina.8 7 Left paramere strongly curved ventrally and evenly narrowed towards apex. C. capillata Ahrens T Left paramere strongly curved straight, and slightly widened towards apex. C. langtangica Ahrens 8 Metatarsomere one shorter than following two tar- someres combined and 1/4 of its length longer than dorsal metatibial spine. Setae on dorsal portion of left paramere present. C. trisuliensis Ahrens 8’ Metatarsomere one as long as two following tar- someres combined and 1/3 of its length longer than dorsal metatibial spine. Setae on dorsal portion of left paramere lacking. C. delectabilis Ahrens 9 Abdominal sternite four strongly and transversely con¬ vex, medially slightly elevated. C. indrai Ahrens 9’ All abdominal stemites simple and without elevations. .10 10 Lateral margins of pronotum behind anterior angles and behind middle less strongly curved.11 10’ Lateral margins of pronotum nearly evenly curved. 17 11 Lateral margins of pronotum behind middle concave. .12 11 ’ Lateral margins of pronotum behind middle straight. .13 12 Long lamellose process sharply pointed, on each side of phallobase ventrally before apex. Parameres long and at apex sharp. C. tigrina Brenske 12’ Sharp process on each side of phallobase at apex. Para¬ meres short and blunt at apex. C. bertiae Ahrens 13 Body < 8 mm. Punctures on labroclypeus moderate¬ ly dense. C. autumnalis Ahrens 13’ Body > 9 mm. Punctures on labroclypeus dense. ..14 14 Eyes small, ratio diameter/interocular width < 0.6. 15 14’ Eyes relatively large, ratio diameter/interocular width >0.7.16 15 Metatarsomere one longer than two following tar- someres combined. C. brendelli Ahrens 15’Metatarsomere one as long as two following tar- someres combined. C. lachungensis Ahrens 16 Metatarsomere one shorter than two following tar- someres combined. C. begnasia Ahrens 16’Metatarsomere one as long as two following tar- someres combined. C. cambeforti Ahrens 17 Eyes small, ratio diameter/interocular width: 0.5-0.61. .18 17’ Eyes large, ratio diameter/interocular width: 0.84. ... . C. hingstoni Ahrens 18 Right paramere dorsally not with a narrow and long process.19 18’Right paramere dorsally with a narrow and long process. Antennal club as long as remaining anten- nomeres combined. C. chiplingensis Ahrens 19 Common basal median process of parameres narrow and long.C. poggii Ahrens 19’ Common basal median process of parameres wide and moderately long.20 20 Left paramere lobe distinctly subequal in size to the right one.C. managensis Ahrens 20’Left paramere lobe distinctly smaller than the right one.C. zhangmuensis Liu & Ahrens Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 22 Dirk Ahrens & Silvia Fabrizi Calloserica autumnalis Ahrens, 1999 Calloserica autumnalis Ahrens, 1999b: 193, 2005: 217. Material examined. See Ahrens 1999b (p. 193). Aedeagus. See Ahrens 1999b (figs 21-23, p. 194). Distribution. Known only from the type locality in cen¬ tral Nepal (Fig. 41C). Calloserica barabiseana Ahrens, 1999 Calloserica barabiseana Ahrens, 1999b: 194,2005b: 217. Material examined. See Ahrens 1999b (p. 194). Aedeagus. See Ahrens 1999b (figs 24-26, p. 196). Distribution. Known only from the type locality in cen¬ tral Nepal (Fig. 41C). Calloserica begnasia Ahrens, 1999 Calloserica begnasia Ahrens, 1999b: 186, 2005b: 217. Material examined. See Ahrens 1999b (p. 186). Aedeagus. See Ahrens 1999b (figs 3-5, p. 187). Distribution. Known only from the type locality in south¬ ern Tibet (Fig. 41C). Calloserica bertiae Ahrens, 2000 Calloserica bertiae Ahrens, 2000a: 816; Ahrens 2005b: 217. Material examined. See Ahrens 2000a (p. 193); Liu et al. 2014a (p. 171). Aedeagus. See Ahrens 2000a (fig. 1, p. 817). Distribution. Known only from the type locality in Sikkim (Fig. 41B). Calloserica brendelli Ahrens, 1999 Calloserica brendelli Ahrens, 1999b: 186; Ahrens 2000a: 821, 2004b: 56, 2005b: 217, 2006a: 410. Material examined. See Ahrens 1999b (p. 186); 2000a (p. 821); 2004b (p. 56), 2006a (p. 410). Aedeagus. See Ahrens 1999b (figs 6-8, p. 187). Distribution. Known only from the type locality in cen¬ tral Nepal (Kathmandu valley) (Fig. 41C). Calloserica cambeforti Ahrens, 2000 Calloserica cambeforti Ahrens, 2000a: 820; Ahrens 2005b: 217. Material examined. See Ahrens 2000a (p. 820). Aedeagus. See Ahrens 2000a (fig. 4, p. 820). Distribution. Known only from the type locality in Sikkim (Fig. 4IB). Calloserica capillata Ahrens, 2005 Calloserica capillata Ahrens, 2005: 225; Ahrens 2005b: 217. Material examined. See Ahrens 2005b (p. 225); Liu et al. 2014a (p. 171). Aedeagus. See Ahrens 2005b (fig. 9A-C, p. 229). Distribution. Known only from the type locality in cen¬ tral Nepal and closeby areas in southern Tibet (Fig. 4IB). Calloserica chiplingensis Ahrens, 1999 Calloserica chiplingensis Ahrens, 1999b: 192; Ahrens 2000a: 821,2005b: 217. Material examined. See Ahrens 1999b (p. 192), 2000a (p. 821). Aedeagus. See Ahrens 1999b (figs 15-17, p. 191). Distribution. Known only from the type locality in cen¬ tral Nepal (Kathmandu valley; Fig. 41C). Calloserica delectabilis Ahrens, 2000 Calloserica delectabilis Ahrens, 2000a: 819; Ahrens 2005b: 217. Material examined. See Ahrens 2000a (p. 819). Aedeagus. See Ahrens 2000a (fig. 3, p. 819). Distribution. Known only from the type locality in cen¬ tral Nepal (Kathmandu valley) (Fig. 4IB). Calloserica gosainkundensis Ahrens, 1999 Calloserica gosainkundensis Ahrens, 1999b: 195; Ahrens 2005b: 217. Material examined. See Ahrens 1999b (p. 195). Aedeagus. See Ahrens 1999b (figs 27-29, p. 196). Distribution. Known only from the type locality in cen¬ tral Nepal (Fig. 41C). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 23 Calloserica hingstoni Ahrens, 1999 Calloserica hingstoni Ahrens, 1999b: 192; Ahrens 2005b: 217. Material examined. See Ahrens 1999b (p. 192). Aedeagus. See Ahrens 1999b (figs 18-20, p. 194). Distribution. Known only from the type locality in south¬ ern Tibet (Fig. 41B). Calloserica indrai Ahrens, 2004 Calloserica indrai Ahrens, 2004b: 57; Ahrens 2005b: 217. Material examined. See Ahrens 2004b (p. 57). Aedeagus. See Ahrens 2004b (figs. 55-57, p. 288). Distribution. Endemic to eastern Nepal, so far known on¬ ly from the type locality (Fig. 4IB). Calloserica lachungensis Ahrens, 2000 Calloserica lachungensis Ahrens, 2000a: 817; Ahrens 2005b: 217. Material examined. See Ahrens 2000a (p. 817). Aedeagus. See Ahrens 2000a (fig. 2, p. 818). Distribution. Known only from the type locality in Sikkim (Fig. 41B). Calloserica langtangica Ahrens, 1999 (Figs 25J, 41C) Calloserica langtangica Ahrens, 1999b: 188; Ahrens 2005b: 217; Qzgul-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 1999b (p. 188). Aedeagus. See Ahrens 1999b (figs 9-11, p. 189). Distribution. Known only from the type locality in cen¬ tral Nepal (Fig. 41C). Calloserica managensis Ahrens, 2005 Calloserica managensis Ahrens, 2005b: 228. Material examined. See Ahrens 2005b (p. 228). Aedeagus. See Ahrens 2005b (fig. 9D-F, p. 229). Distribution. Known only from the type locality in cen¬ tral Nepal. Calloserica poggii Ahrens, 1995 Calloserica poggii Ahrens, 1995a: 137; Ahrens 2005b: 217. Material examined. See Ahrens 1995a (p. 137). Aedeagus. See Ahrens 1995a (figs 4-6, p. 138). Distribution. Known only from the type locality in cen¬ tral Nepal (Annapurna Himal) (Fig. 4IB). Calloserica raksensis Ahrens, 2004 Calloserica raksensis Ahrens, 2004b: 59; Ahrens 2005b: 217. Material examined. See Ahrens 2004b (p. 59). Aedeagus. See Ahrens 2004b (figs. 61-63, p. 389). Distribution. Endemic to eastern Nepal, so far known on¬ ly from the type locality (Fig. 4IB). Calloserica rupthangensis Ahrens, 2004 Calloserica rupthangensis Ahrens, 2004b: 58; Ahrens 2005b: 217. Material examined. See Ahrens 2004b (p. 58). Aedeagus. See Ahrens 2004b (figs. 58-60, p. 389). Distribution. Endemic to eastern central Nepal, so far known only from the type locality (Fig. 4IB). Calloserica tigrina Brenske, 1894 Calloserica tigrina Brenske, 1894: 10, 56; Ahrens 1995a: 137, 1999b: 185, 2000a: 821, 2004b: 55, 2005b: 217, 2006a: 410. Material examined. See Ahrens 1999b (p. 185), 2000a (p. 821), 2004b (p. 55), 2006a (p. 410). Aedeagus. See Ahrens 1999b (figs 1-3, p. 138). Distribution. Endemic to the Darjeeling area (Fig. 4IB). Calloserica trisuliensis Ahrens, 1999 Calloserica trisuliensis Ahrens, 1999b: 190; Ahrens 2000a: 821,2005b: 217. Material examined. See Ahrens 1999b (p. 190), 2000a (p. 821). Aedeagus. See Ahrens 1999b (figs 12-14, p. 191). Distribution. Known only from the type locality in cen¬ tral Nepal (Ganesh Himal) (Fig. 4IB). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 24 Dirk Ahrens & Silvia Fabrizi Calloserica zhangmuensis Liu & Ahrens, 2014 Calloserica zhangmuensis Liu & Ahrens, 2014: 159. Material examined. See Liu et al. 2014a (p. 159). Aedeagus. See Liu et al. 2014a (fig. 1A-C, p. 158). Distribution. Known only from the type locality in cen¬ tral Tibet Himalaya (Fig. 41B). Xenoserica Ahrens, 2005 Xenoserica Ahrens, 2005c: 273; (type species by original designation: Nipponoserica sindhensis Ahrens, 2000); Ahrens 2007c: 40. Remarks. The genus was established by Ahrens (2005c) in a follow-up of the results of a phylogenetic analysis based on morphology. Recently, we were able to include several taxa of Xenoserica into a major molecular phylo¬ genetic analysis including more than 600 Sericini species from all over the World (Eberle et al. 2016). Results from this analysis indicate that X. brachyptera is the sister tax¬ on to the species of the Serica erectosetosa group (Ahrens 2004b). Therefore, we transferred these species now to Xenoserica. Key to Xenoserica species (S$) 1 Metacoxa densely covered with long adpressed setae. Parameres completely symmetric.2 1 ’ Metacoxa glabrous, only laterally with a few robust setae. Parameres asymmetric.5 2 Parameres with a basal process directed basally. Ely¬ tra narrowly chitinous at apex.3 T Parameres without a basal process. Elytra narrowly membraneous at apex. X selaensis Ahrens & Fabrizi 3 Eyes small, ratio ocular diameter/interocular width: 0.5-0.64. Parameres apically narrowed and sharply pointed.4 3’ Eyes large, ratio ocular diameter/interocular width: 0.81. Parameres apically weakly rounded and reflexed outward. X. matthiasi Shrestha et al., X pindarensis (Ahrens) 4 Dorsal surface reddish brown and dull. Antennal club distinctly longer than the remaining antennomeres combined. Wings fully developed. . X. sindhensis (Ahrens) 4’ Dorsal surface blackish and shiny. Antennal club as long as the remaining antennomeres combined. Wings reduced in length. X brachyptera Ahrens 5 Metatarsomeres grooved dorsally. Pronotum and ely¬ tra with appressed scale-like white setae. . X karnaliensis (Ahrens) 5’ Metatarsomeres grooved smooth. Pronotum and ely¬ tra without appressed scale-like white setae.6 6 Anterior angles of pronotum protruding and right-an¬ gled, lateral margin immediately behind anterior an¬ gles weakly sinuate; disc of pronotum throughout with erect long setae. X erectosetosa (Ahrens) 6’ Anterior angles of pronotum blunt, weakly protruding, lateral margin immediately behind anterior angles con¬ vex; disc of pronotum on anterior third only with erect long setae.7 7 Setae on anterior pronotum sparse.8 7’ Setae on anterior pronotum dense. . X yadongensis (Liu & Ahrens) 8 Labroclypeus densely punctate, anterior margin dis¬ tinctly sinuate medially. Intervals of elytra densely punctate. X koshiana (Ahrens) 8’ Labroclypeus moderately densely punctate, anterior margin almost straight medially. Intervals of elytra sparsely punctate. X. somathangana (Ahrens) Xenoserica brachyptera Ahrens, 2005 Xenoserica brachyptera Ahrens, 2005c: 275. Material examined. See Ahrens 2005c (p. 275). Aedeagus. See Ahrens 2005c (fig. 9A-C, p. 276). Distribution. Endemic to western central Nepal (Dhaula¬ giri Himal) (Fig. 4ID). Xenoserica erectosetosa (Ahrens, 1999) comb. n. Serica (s.l.) erectosetosa Ahrens, 1999c: 315; Ahrens 2004b: 24, 2006a: 409. Material examined. See Ahrens 1999c (p. 315); 2004b (p. 25), 2006a (p. 409). Aedeagus. See Ahrens 1999c (figs 140-142, p. 229). Distribution. Endemic to eastern central Nepal (Fig. 41D). Xenoserica karnaliensis (Ahrens, 1999) comb. n. (Fig. 25L) Serica karnaliensis Ahrens, 1999c: 319; Ahrens 2004b: 25; Ahrens & Fabrizi 2011: 161; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999c (p. 319); Ahrens & Fabrizi 2011 (p. 161); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999c (figs 146-148, p. 230). Distribution. Endemic to western Nepal (Fig. 41D). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 25 Xenoserica koshiana (Ahrens, 1999) comb. n. Serica koshiana Ahrens, 1999c: 321; Ahrens 2004b: 25. Material examined. See Ahrens 1999c (p. 321). Aedeagus. See Ahrens 1999c (figs 149-151, p. 230). Distribution. Endemic to eastern Nepal (Fig. 4ID). Xenoserica matthiasi Shrestha, Eberle & Ahrens, 2012 Xenoserica matthiasi Shrestha, Eberle & Ahrens, 2012: 383. Material examined. See Shrestha et al. 2012 (p. 383). Aedeagus. See Shrestha et al. 2012 (figs 1-3, p. 386). Distribution. Endemic to western Nepal (Fig. 4ID). Xenoserica pindarensis (Ahrens, 2000) Nipponoserica pindarensis Ahrens, 2000c: 27; Ahrens 2004b: 61. Xenoserica pindarensis'. Ahrens 2005c: 275. Material examined. See Ahrens 2000c (p. 27). Aedeagus. See Ahrens 2000c (figs 4-6, p. 29). Distribution. Endemic to the Pindar Valley (Uttar Pradesh) (Fig. 4ID). Xenoserica selaensis Ahrens & Fabrizi, 2009 Xenoserica selaensis Ahrens & Fabrizi, 2009b: 255. Material examined. See Ahrens & Fabrizi 2009b (p. 255). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 3M-0, p. 279). Distribution. Endemic to Arunachal Pradesh (Fig. 41D). Xenoserica sindhensis (Ahrens, 2000) (Figs 25K, 4ID) Nipponoserica sindhensis Ahrens, 2000c: 26; Ahrens 2004b: 61. Xenoserica sindhensis. Ahrens 2005c: 275, 2006a: 411; Ahrens & Fabrizi 2009b: 271. Material examined. See Ahrens 2000c (p. 26), 2004b (p. 61), 2006a (p. 411); Ahrens & Fabrizi 2009b (p. 271). Aedeagus. See Ahrens 2000c (figs 1-3, p. 29). Distribution. Endemic to Kashmir (Fig. 4ID). Xenoserica somathangana (Ahrens, 1999) comb. n. Serica somathangana Ahrens, 1999: 317; Ahrens 2004b: 25. Material examined. See Ahrens 1999c (p. 317), 2004b (p. 25). Aedeagus. See Ahrens 1999c (figs 143-145, p. 229). Distribution. Endemic to eastern central Nepal (Fig. 41D). Xenosericayadongensis (Liu & Ahrens, 2014) comb. n. Serica (s.l ) yadongensis Fiu & Ahrens, 2014: 168. Material examined. See Fiu et al. 2014a (p. 168); 1 $ “Yadong, Xizang, 1.VII.2004, leg. Ba Yibin, Shi Aimin” (HBUM). Aedeagus. See Fiu et al. 2014a (fig. 3I-K, p. 166). Distribution. Endemic to Southern Tibet (Fig. 4ID). Remarks. This female specimen from Yadong was in Fiu et al. (2014a) erroneously reported as Serica karnalien- sis Ahrens. Nepaloserica Frey, 1965 Nepaloserica Frey, 1965b: 88 (type species by monotypy: Nepalosericaprocera Frey, 1965); Ahrens & Sabatinel- li 1996: 209; Ahrens 1999b: 198,2004b: 62,2007c: 27, 2012: 305. Key to Nepaloserica species {S3) 1 Fast abdominal sternite with a pair of tubercles. ... 2 1 ’ Fast abdominal sternite without a pair of tubercles or with a transversely elevated carina.18 2 Anterior angles of pronotum sharply pointed.3 2’ Anterior angles of pronotum rounded or blunt.10 3 Tubercles on last abdominal sternite close, separated by less than a metatibial width.4 3’ Tubercles on last abdominal sternite distant, separat¬ ed by more than the metatibial width.6 4 Antennal club shorter (ratio width of base of labro- clypeus/length of club: 0.76-0.8).5 4’ Antennal club longer (ratio width of base of labro- clypeus/length of club: 0.5). . N. vignai Ahrens & Sabatinelli 5 Fabroclypeus wide and distinctly trapezoidal. Anteri¬ or margin of pronotum setose. Setae on frons dense. Eyes moderately large (ratio diameter/interocular dis¬ tance <0.7). N. ganeshi Ahrens Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 26 Dirk Ahrens & Silvia Fabrizi 5 Labroclypeus narrow but trapezoidal. Anterior margin of pronotum glabrous. Setae on frons less dense. Eyes large (ratio diameter/interocular distance < 0.79). . N. pewaensis Ahrens 6 Eyes small (ratio diameter/interocular distance < 0.6). .7 6’ Eyes large (ratio diameter/interocular distance > 0.8). .8 7 Pronotum glabrous. . N. mustangia Ahrens & Sabatinelli 7’ Pronotum with dense erect setae. . N. baglungensis Ahrens 8 Anterior margin of labroclypeus straight, not sinuate medially; surface with sparse punctation. Tubercles on last abdominal sternite large and sharply pointed. . N. fabriziae Ahrens 8 ’ Anterior margin of labroclypeus concavely sinuate me¬ dially; surface with very dense and irregular puncta¬ tion. Tubercles on last abdominal sternite small and blunt.9 9 Lateral margin of labrum separated from that of clypeus by a deep incision. Apex of phallobase asym¬ metrical. Lateral margins of pronotum evenly convex. . N. archolabrata Ahrens & Sabatinelli 9’ Lateral margin of labrum entirely fused with that of clypeus, without incision. Apex of phallobase symmet¬ rical. Lateral margins of pronotum nearly straight be¬ hind anterior angles and in basal half. . . N. schmidti Ahrens & Sabatinelli 10 Disc of pronotum setose. . N, migliaccioi Ahrens & Sabatinelli 10’ Disc of pronotum glabrous.11 11 Metatibia short (ratio width/length -1/3.7). Tubercles on last abdominal sternite distant and sharply point¬ ed. N. brevipes Ahrens & Sabatinelli IE Metatibia longer (ratio width/length -1/4).12 12 Tubercles on last abdominal sternite distinctly point¬ ed, without setae.13 12’ Tubercles on last abdominal sternite blunt, densely covered with robust setae.15 13 Labroclypeus square, anterior angles moderately rounded. Antennal club more than twice as long as an- tennomeres one to three combined. . N. helambuensis Ahrens & Sabatinelli 13’ Labroclypeus rectangular, anterior angles strongly rounded. Antennal club twice as long as antennomeres one to three combined.14 14 Phallobase without tubercles on ventral side. . N. muelleri tuberculata Ahrens, N. telbrungensis Ahrens 14’ Phallobase with two robust tubercles on ventral side. . N. manasluensis Ahrens 15 Tubercles on last abdominal sternite very small. Sides of pronotum weakly convex.16 15’ Tubercles on last abdominal sternite large and blunt¬ ly rounded. . N. procera procera Frey, N. lamjungi Ahrens 16 Body > 11 mm. Parameres narrow and long.17 16’ Body <10 mm. Parameres short and wide. . N. perrecondita Ahrens 17 Lateral anterior margin of pronotum with setae. . N. vilya Ahrens & Sabatinelli 17’ Anterior margin of pronotum without setae. . N. richardsonae Ahrens 18 Last abdominal sternite with a transverse carina; Ca¬ rina may bear 1-3 tubercles.19 18’ Last abdominal sternite simple without any elevation. .21 19 Disc of pronotum with long, moderately dense setae. . N. bruschii Ahrens & Sabatinelli 19’ Disc of pronotum glabrous.20 20 Tubercles on last abdominal sternite situated in its an¬ terior half. Pronotum widest at middle, lateral margins straight or convex in posterior half. Frons with dense long setae. Antennal club as long as antennomeres one to three combined. . N. muelleri muelleri Ahrens & Sabatinelli 20’ Tubercles on last abdominal sternite situated in its pos¬ terior half. Pronotum widest at base. Frons with only a few single setae. Antennal club 2.5 times as long as antennomeres one to three combined. . N. procera rufescens Frey, N. goomensis Ahrens 21 Lateral margins of pronotum straight or convex in pos¬ terior half. .22 21’ Lateral margins of pronotum concavely sinuate in pos¬ terior half. .29 22 Phallobase with a high longitudinal lamina on ventral side.23 22’ Phallobase without a longitudinal lamina on ventral side.26 23 Labroclypeus widest at base, lateral margins strongly curved and convergent anteriorly. . N. baehri Ahrens & Sabatinelli 23’ Labroclypeus square or rectangular, lateral margins on¬ ly weakly convergent anteriorly.24 24 Labroclypeus more rectangular, margins strongly re- flexed. Metatarsi dorsally densely punctate. . N. yeti Ahrens 24’ Labroclypeus more square, margins weakly reflexed. Metatarsi dorsally sparsely punctate.25 25 Anterior angles of labroclypeus strongly convex. . N. longispina Ahrens 25’ Anterior angles of labroclypeus weakly convex. . N. similis Frey 26 Labroclypeus more or less square. Lateral margins of pronotum subparallel in basal half. Body size < 11 mm. .27 26’ Labroclypeus transversely rectangular. Lateral margins of pronotum evenly convex. Body size >11 mm. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 27 Meta-tibia: ratio width/length: 1/4. . N. sankhuwasabhae Ahrens & Sabatinelli 27 Anterior angles of pronotum moderately sharp. Body size <10 mm.28 27’ Anterior angles of pronotum very sharp. Body size > 10 mm. N. rufobrunnea Ahrens 28 Frons on disc glabrous. Metatibia very slender, ratio width/length: 1/5.3. . N. phulcokiensis Ahrens & Sabatinelli 28’ Frons on disc densely setose. Metatibia wider, ratio width/length: 1/3.9. ...N. nielamuensis Liu & Ahrens 29 Body size < 9 mm.30 29’ Body size >10 mm. Labroclypeus never with trans¬ verse wrinkles.31 30 Labroclypeus with transverse wrinkles that link the coarse punctures with each other. Surface of smooth area in front of eyes produce a distinct angle with the surface of the labroclypeus. . N. barbara Ahrens & Sabatinelli 30’ Labroclypeus without transverse wrinkles, only with fine and evenly dense punctures. Surface of smooth area in front of eyes at the same level with the surface of the labroclypeus. . N. thimphui Ahrens & Sabatinelli 31 Lateral margins of pronotum weakly curved in ante¬ rior half, anterior angles sharp and distinctly produced. Labroclypeus widest at base, convexly narrowed an¬ teriorly. Metatibia wider (ratio width/length: 1/3.8). Phallobase with a longitudinal carina on ventral side. . N. induwae Ahrens 31 ’ Lateral margins of pronotum strongly curved in ante¬ rior half, anterior angles bluntly rounded and weakly produced anteriorly. Labroclypeus widest shortly be¬ fore base. Metatibia slender (ratio width/length: 1/4.6-4.7). Phallobase with a pair of teeth on ventral side.32 32 Lateral margins of pronotum straight or convex in basal half. Right paramere without lateral distal tooth . N. hartmanni Ahrens 32’ Lateral margins of pronotum concavely sinuate in basal half. Each paramere with a lateral distal tooth. . N. jumlaica Ahrens Nepaloserica archolabrata Ahrens & Sabatinelli, 1996 Nepaloserica archolabrata Ahrens & Sabatinelli, 1996: 238; Ahrens 1999b: 215, 2004b: 67. Material examined. See Ahrens & Sabatinelli 1996 (p. 238); Ahrens 2004b (p. 67). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 45^17, p. 237). Distribution. Endemic to central Nepal (Kathmandu val¬ ley) (Fig. 42A). Nepaloserica baehri Ahrens & Sabatinelli, 1996 Nepaloserica baehri Ahrens & Sabatinelli, 1996: 214; Ah¬ rens 1999b: 215, 2004b: 65. Material examined. See Ahrens & Sabatinelli 1996 (p. 214); Ahrens 1999b (p. 215), 2004b (p. 65). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 9, 10, p. 213). Distribution. Endemic to eastern central Nepal (Fig. 42 A). Nepaloserica baglungensis Ahrens, 2012 Nepaloserica baglungensis Ahrens, 2012: 311. Material examined. See Ahrens 2012 (p. 311). Aedeagus. See Ahrens 2012 (fig. 5E-H, p. 314). Distribution. Endemic to western central Nepal (Fig. 41F). Nepaloserica barbara Ahrens & Sabatinelli, 1996 Nepaloserica barbara Ahrens & Sabatinelli, 1996: 232; Ahrens 2004b: 73. Material examined. See Ahrens & Sabatinelli 1996 (p. 232); Ahrens 2004b (p. 73). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 38, 39, p. 233). Distribution. Endemic to Sikkim and the Darjeeling area (Fig. 41E). Nepaloserica brevipes Ahrens & Sabatinelli, 1996 Nepaloserica brevipes Ahrens & Sabatinelli, 1996: 231; Ahrens 2004b: 72. Material examined. See Ahrens & Sabatinelli 1996 (p. 231); Ahrens 2004b (p. 72). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 42, 43, p. 233). Distribution. Endemic to the Darjeeling area (Fig. 4IF). Nepaloserica bruschii Ahrens & Sabatinelli, 1996 Nepaloserica bruschii Ahrens & Sabatinelli, 1996: 222; Ahrens 1999b: 200, 2004b: 66. Material examined. See Ahrens & Sabatinelli 1996 (p. 222); Ahrens 1999b (p. 200). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 28 Dirk Ahrens & Silvia Fabrizi Aedeagus. See Ahrens & Sabatinelli 1996 (figs 17-19, p. 217). Distribution. Endemic to central and eastern Nepal (Fig. 41F). Nepaloserica fabriziae Ahrens, 1999 Nepaloserica fabrizii Ahrens, 1999b: 207; Nepaloserica fabriziae '. Ahrens 2004b: 71. Material examined. See Ahrens 1999b (p. 207). Aedeagus. See Ahrens 1999b (figs 46^18, p. 209). Distribution. Endemic to Sikkim (Fig. 4IE). Nepaloserica ganeshi Ahrens, 1999 (Fig. 25N) Nepaloserica ganeshi Ahrens, 1999b: 201; Ahrens 2004b: 66 . Material examined. See Ahrens 1999b (p. 201), 2004b (p. 66). Aedeagus. See Ahrens 1999b (figs 36, 39, p. 203). Distribution. Endemic to central Nepal (Ganesh and Helambu Himal) (Fig. 4IE). Nepaloserica goomensis Ahrens, 1999 (Fig. 25M) Nepaloserica goomensis Ahrens, 1999b: 199; Ahrens 2004b: 64, 2006a: 411, 2012: 309. Material examined. See Ahrens 1999b (p. 199), 2004b (p. 64), 2006a (p. 411). Aedeagus. See Ahrens 1999b (figs 31-33, p. 199). Distribution. Endemic to the Darjeeling area and Sikkim (Fig. 41E). Nepaloserica hartmanni Ahrens, 1999 Nepaloserica hartmanni Ahrens, 1999b: 203; Ahrens 2004b: 72. Material examined. See Ahrens 1999b (p. 203). Aedeagus. See Ahrens 1999b (figs 35, 38, p. 203). Distribution. Endemic to western Nepal (Fig. 41E). Nepaloserica helambuensis Ahrens & Sabatinelli, 1996 Nepaloserica helambuensis Ahrens & Sabatinelli, 1996: 221; Ahrens 1999b: 208, 2004b: 71; Ahrens & Fabrizi 2011: 163. Material examined. See Ahrens & Sabatinelli 1996 (p. 221); Ahrens 1999b (p. 208), 2004b (p. 71); Ahrens & Fabrizi 2011 (p. 163). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 23-25, p. 219). Distribution. Endemic to Helambu und Langtang Himal in central Nepal (Dhaulagiri Himal) (Fig. 42A). Nepaloserica induwae Ahrens, 1999 Nepaloserica induwae Ahrens, 1999b: 211; Ahrens 2004b: 65. Material examined. See Ahrens 1999b (p. 211). Aedeagus. See Ahrens 1999b (figs 54-56, p. 211). Distribution. Endemic to eastern Nepal (Fig. 4IF). Nepaloserica jumlaica Ahrens, 1999 Nepaloserica jumlaica Ahrens, 1999b: 204; Ahrens 2004b: 72; Ahrens & Fabrizi 2011: 163; Shrestha et al. 2012: 382. Material examined. See Ahrens 1999b (p. 204), 2004b (p. 72); Ahrens & Fabrizi 2011 (p. 163); Shrestha et al. 2012 (p. 382). Aedeagus. See Ahrens 1999b (figs 40^12, p. 206). Distribution. Endemic to western Nepal (Fig. 41E). Nepaloserica lamjungi Ahrens, 1999 Nepaloserica lamjungi Ahrens, 1999b: 205; Ahrens 2004b: 71. Material examined. See Ahrens 1999b (p. 205). Aedeagus. See Ahrens 1999b (figs 43^15, p. 206). Distribution. Endemic to western central Nepal (Anna¬ purna Himal) (Fig. 42A). Nepaloserica longispina Ahrens, 1999 Nepaloserica longispina Ahrens, 1999b: 212; Ahrens 2004b: 65. Material examined. See Ahrens 1999b (p. 212). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 29 Aedeagus. See Ahrens 1999b (figs 57-59, p. 214). Distribution. Endemic to eastern central Nepal (Solu Khumbu Himal) (Fig. 4IF). Nepaloserica manasluensis Ahrens, 2004 Nepaloserica manasluensis Ahrens, 2004b: 69. Material examined. See Ahrens 2004b (p. 69). Aedeagus. See Ahrens 2004b (figs 70-72, p. 391). Distribution. Endemic to central Nepal (Manaslu Himal) (Fig. 42A). Nepaloserica migliaccioi Ahrens & Sabatinelli, 1996 Nepaloserica migliaccioi Ahrens & Sabatinelli, 1996: 218; Ahrens 1999b: 207, 2004b: 71, 2006a: 410. Material examined. See Ahrens & Sabatinelli 1996 (p. 218) ; Ahrens 1999b (p. 207), 2004b (p. 71), 2006a (p. 410). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 20-22, p. 219) . Distribution. Endemic to eastern central Nepal (Fig. 4IF). Nepaloserica muelleri muelleri Ahrens & Sabatinelli, 1996 Nepaloserica muelleri Ahrens & Sabatinelli, 1996: 224; Ahrens 1999b: 208. Nepaloserica muelleri muelleri. Ahrens 2004b: 68,2006a: 410. Material examined. See Ahrens & Sabatinelli 1996 (p. 224) ; Ahrens 1999b (p. 208), 2004b (p. 68), 2006a (p. 410). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 29-31, p. 225) . Distribution. Endemic to western central Nepal (Anna- puma Himal) (Fig. 41F). Nepaloserica muelleri luberculala Ahrens, 2004 Nepaloserica muelleri tuberculata Ahrens, 2004b: 68. Material examined. See Ahrens 2004b (p. 68). Aedeagus. See Ahrens 2004b (figs 67-69, p. 390). Distribution. Endemic to western central Nepal (Dhaula¬ giri Himal) (Fig. 4IF). Nepaloserica mustangia Ahrens & Sabatinelli, 1996 Nepaloserica mustangia Ahrens & Sabatinelli, 1996: 226; Ahrens 1999b: 210, 2004b: 72, 2006a: 410; Ahrens & Fabrizi2011: 163; Ctegii 1-Siemund & Ahrens 2015: 171. Material examined. See Ahrens & Sabatinelli 1996 (p. 226); Ahrens 1999b (p. 210), 2004b (p. 72), 2006a (p. 410); Ahrens & Fabrizi 2011 (p. 163). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 26-28, p. 225). Distribution. Endemic to the upper Kali Gandaki valley in western central Nepal (Fig. 4IE). Nepaloserica nielamuensis Liu & Ahrens, 2014 Nepaloserica nielamuensis Fiu & Ahrens, 2014: 163. Material examined. See Fiu et al. 2014a (p. 163). Aedeagus. See Fiu et al. 2014a (fig. 2E-G, p. 162). Distribution. Endemic to southern central Tibet (Fig. 41F). Nepaloserica perrecondita Ahrens, 2004 Nepaloserica perrecondita Ahrens, 2004b: 66. Material examined. See Ahrens 2004b (p. 66). Aedeagus. See Ahrens 2004b (figs 64-66, p. 390). Distribution. Endemic to eastern western central Nepal (Rolwaling Himal) (Fig. 42A). Nepaloserica pewaensis Ahrens, 2012 Nepaloserica pewaensis Ahrens, 2012: 313. Material examined. See Ahrens 2012 (p. 313). Aedeagus. See Ahrens 2012 (fig. 5J-F, p. 314). Distribution. Endemic to western central Nepal (Fig. 41E). Nepaloserica phnlcokiensis Ahrens & Sabatinelli, 1996 Nepaloserica phulcokiensis Ahrens & Sabatinelli, 1996: 236; Ahrens 2004b: 73, 2006a: 410. Material examined. See Ahrens & Sabatinelli 1996 (p. 236) ; Ahrens 2004b: 73, 2006a (p. 410). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 49-51, p. 237) . Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 30 Dirk Ahrens & Silvia Fabrizi Distribution. Endemic to central Nepal (Kathmandu val¬ ley) (Fig. 4IE). Nepaloserica procera procera Frey, 1965 Nepaloserica procera Frey, 1965b: 88; Ahrens & Sabatinelli 1996: 210; Ahrens 2006a: 411. Nepaloserica procera procera'. Ahrens 1999b: 198. Material examined. See Ahrens & Sabatinelli 1996 (p. 210) ; Ahrens 2006a (p. 411). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 1-3, p. 211 ) . Distribution. Endemic to eastern central Nepal (Fig. 4IE). Nepaloserica procera rufescens Frey, 1965 Nepaloserica rufescens Frey, 1965a: 281; Ahrens & Sabatinelli 1996: 210. Nepaloserica procera rufescens : Ahrens 1999b: 198; 2004b: 64, 2006a: 411; Ahrens & Fabrizi 2011: 163. Material examined. See Ahrens & Sabatinelli 1996 (p. 210) ; Ahrens 2006a (p. 411); Ahrens & Fabrizi 2011 (p. 163). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 4-6, p. 211 ) . Distribution. Endemic to western central Nepal (Fig. 42 A). Nepaloserica richardsonae Ahrens, 2012 Nepaloserica richardsonae Ahrens, 2012: 308. Material examined. See Ahrens 2012 (p. 308). Aedeagus. See Ahrens 2012 (fig. 5A-C, p. 314). Distribution. Endemic to western central Nepal (Anna¬ purna Himal) (Fig. 41E). Nepaloserica rufobrunnea Ahrens, 1999 Nepaloserica rufobrunnea Ahrens, 1999b: 201; Ahrens 2004b: 73. Material examined. See Ahrens 1999b (p. 201). Aedeagus. See Ahrens 1999b (figs 34, 37, p. 203). Distribution. Endemic to Bhutan (Fig. 4IE). Nepaloserica sankhuwasabhae Ahrens & Sabatinelli, 1996 Nepaloserica sankhuwasabhae Ahrens & Sabatinelli, 1996: 230; Ahrens 2004b: 72. Material examined. See Ahrens & Sabatinelli 1996 (p. 230); Ahrens 2004b (p. 72). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 35-37, p. 229). Distribution. Endemic to eastern Nepal (Fig. 41E). Nepaloserica schmidti Ahrens & Sabatinelli, 1996 Nepaloserica schmidti Ahrens & Sabatinelli, 1996: 215; Ahrens 1999b: 202, 2004b: 67, 2006a: 411; Ahrens & Fabrizi 2011: 163. Material examined. See Ahrens & Sabatinelli 1996 (p. 215), 2006a (p. 411); Ahrens & Fabrizi 2011 (p. 163). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 14-16, p. 217). Distribution. Endemic to western central Nepal (Anna¬ purna Himal) (Fig. 4IE). Nepaloserica similis Frey, 1969 Nepaloserica similis Frey, 1969a: 520; Ahrens & Sabatinelli 1996: 212; Ahrens 1999b: 211, 2004b: 64. Material examined. See Ahrens & Sabatinelli 1996 (p. 212 ); Ahrens 1999b (p. 211), 2004b (p. 64). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 7, 8, p. 213). Distribution. Endemic to eastern central Nepal (Fig. 4IE). Nepaloserica telbrungensis Ahrens, 1999 Nepaloserica telbrungensis Ahrens, 1999b: 208; Ahrens 2006a: 410. Material examined. See Ahrens 1999b (p. 208), 2006a (p. 410). Aedeagus. See Ahrens 1999b (figs 49-51, p. 209). Distribution. Endemic to western central Nepal (Dhaula¬ giri Himal) (Fig. 42A). Nepaloserica thimphui Ahrens & Sabatinelli, 1996 Nepaloserica thimphui Ahrens & Sabatinelli, 1996: 234; Ahrens 1999b: 200, 2004b: 73. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 31 Material examined. See Ahrens & Sabatinelli 1996 (p. 234). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 41, 44, p. 233). Distribution. Endemic to Bhutan (Fig. 4IE). Nepaloserica vignai Ahrens & Sabatinelli, 1996 Nepaloserica vignai Ahrens & Sabatinelli, 1996: 212; Ahrens 1999b: 213, 2004b: 65. Material examined. See Ahrens & Sabatinelli 1996 (p. 212) ; Ahrens 1999b (p. 213), 2004b (p. 65). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 11-13, p. 213) . Distribution. Endemic to central Nepal (Kathmandu val¬ ley) (Fig. 41F). Nepaloserica vilya Ahrens & Sabatinelli, 1996 Nepaloserica vilya Ahrens & Sabatinelli, 1996: 227; Ahrens 1999b: 202; 2004b: 66. Material examined. See Ahrens & Sabatinelli 1996 (p. 227). Aedeagus. See Ahrens & Sabatinelli 1996 (figs 32-34, p. 229). Distribution. Endemic to central Nepal (Solu Khumbu and Ganesh Himal) (Fig. 4IF). Nepaloserica yeti Ahrens, 1999 Nepaloserica yeti Ahrens, 1999b: 213. Material examined. See Ahrens 1999b (p. 213). Aedeagus. See Ahrens 1999b (figs 60-62, p. 214). Distribution. Endemic to central Nepal (Helambu und Rolwaling Himal) (Fig. 41F). Genus Sericania Motschulsky, 1860 Sericania Motschulsky, 1860: 136 (type species by monotypy: Sericaniafuscolineata Motschulsky, 1860); Ahrens 2007b: 517, 2007c: 36. Mesoserica Matsumura, 1911 (nee Brenske, 1897): 118 (type species by monotypy: Mesoserica sachalinensis Matsumura, 1911). Key to Sericania species of the Indian subcontinent (c? 6 mm.G. yigongensis Liu & Ahrens 7’ Apex of lateral apophysis of phallobase evenly point¬ ed. Body <5.5 mm.G. hani Liu & Ahrens 8 Lateral apophysis of phallobase short, distinctly shorter than half of phallobasal length. .G. compacta Ahrens & Fabrizi 8’ Lateral apophysis of phallobase long, subequal phal¬ lobasal length.G. victori Ahrens & Fabrizi 9 Antennal club composed of three antennomeres. Body length > 6 mm.10 9’ Antennal club composed of four or more anten¬ nomeres.12 10 Anterior angles of pronotum obsolete, disc of prono- tum glabrous. Antennal club 1.5 times as long as the remaining antennomeres combined. .G. singhikensis Ahrens 10’ Anterior angles of pronotum blunt or acute, disc of pronotum setose. Antennal club about as long as the remaining antennomeres combined.11 11 Lateral margins of pronotum evenly convex, in basal half almost subparallel. .G. tawangensis Ahrens and Fabrizi 11 ’ Pronotum basally distinctly narrowed. .G. latesquamosa (Frey) 12 Pronotum basally distinctly narrowed. Antennal club composed of four antennomeres. .G. annuliforceps Ahrens and Fabrizi 12’ Lateral margins of pronotum evenly convex, in basal half almost subparallel.13 13 Antennal club composed of four antennomeres. ...14 13’ Antennal club composed of five antennomeres. ...20 13” Antennal club composed of six antennomeres.24 14 Apical apophysis of phallobase short, about half as long as parameres. Body length < 5 mm.15 14’ Apical apophysis of phallobase long, almost as long as parameres. Body length > 6 mm.16 15 Antenna with nine antennomeres. Antennal club 1.5 times as long as the remaining antennomeres com¬ bined. Parameres in dorsal view narrow, their lateral margins subparallel.G. pellecta (Brenske) 15’ Antenna with ten antennomeres. Antennal club 2.5 times as long as the remaining antennomeres com¬ bined. Parameres in dorsal view broad, their lateral margins convexly widened at middle. .G. tumba Ahrens 16 Parameres short, one third as long as phallobase. .G. bomdilana Ahrens & Fabrizi 16’ Parameres long, at least half as long as phallobase. 17 17 Lateral apophysis of phallobase shorter than the fused parameres that are at apex slightly pointed.18 17’ Lateral apophysis of phallobase slightly longer than the fused parameres that are at apex distinctly round¬ ed.G. aniniensis Ahrens & Fabrizi 18 Punctures on head coarse and moderately dense. Right dorsal margin of paramere not notched at basal third. .G. gogonaica Ahrens 18’ Punctures on head fine, dense. Right dorsal margin of paramere deeply notched at basal third.19 19 Parameres (lateral view) dorsoventrally widened at middle. Lateral apophysis of phallobase basally with¬ out small convexity. .G. variipennis wuermlii (Frey) 19’ Parameres (lateral view) dorsoventrally very slightly widened at middle. Lateral apophysis of phallobase basally with a small convexity. .G. variipennis variipennis (Moser) 20 Lateral apophysis of phallobase in cross-section cir¬ cular. Lateral apophysis of phallobase displaced slightly dorsally being broad at base and evenly nar¬ rowed towards the apex; phallobase apically distinct¬ ly narrowed (dorsal view). .G. ignobilis Ahrens & Fabrizi 20’ Lateral apophysis of phallobase laterally flattened. .. .21 21 Lateral apophysis of phallobase almost as wide as long. .G. lobiceps Ahrens & Fabrizi 2L Lateral apophysis of phallobase at least twice as wide as long (lateral view).22 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 36 Dirk Ahrens & Silvia Fabrizi 22 Lateral apophysis of phallobase sharply pointed at apex. G. perdita Ahrens 22’ Lateral apophysis of phallobase convex at apex. ..23 23 Fused parameres narrowed apically. . G. keithi Ahrens & Fabrizi 23’ Fused parameres distinctly widened apically. . G. cymosa (Brenske) 24 Protarsal claws symmetrical, basal tooth of interior claw sharply pointed. Head entirely shiny. Parameres each with a very long basal extension being almost as long as the paramere itself. .25 24’ Protarsal claws asymmetrical, basal tooth of interior claw bluntly truncate and slightly widened. Basal half of head dull. Basal extension of parameres short, if present.27 25 Lateral apophysis of phallobase absent or very short. Parameres each with a very long basal extension be¬ ing almost as long as the paramere itself. .26 25’ Lateral apophysis of phallobase present, prominent. .G. marginipes (Brenske) 26 Antennal club composed of five antennomeres. Dis¬ tal part of parameres longer. Left paramere with a large lateral subapical tooth.G. etalinensis sp. n. 26’ Antennal club composed of six antennomeres. Left paramere without lateral subapical tooth. .G. lohitensis Ahrens & Fabrizi 27 Ventral apical spine of metatibia sharply pointed at apex.28 27’Ventral apical spine of metatibia slightly truncate at apex.29 28 Parameres and its basal appendages very short, para¬ meres shorter than one sixth of length of phallobase; lateral apophysis of phallobase absent. .G. exilis Ahrens & Fabrizi 28’ Parameres and its basal appendages long, parameres half as long as phallobase; lateral apophysis of phal¬ lobase present.G. dirangensis Ahrens & Fabrizi 29 Parameres on right side strongly widened apically (dorsal view).G. vogleri Ahrens & Fabrizi 29’ Parameres at apex almost as wide as at base (dorsal view).30 30 Ventral apical spine of metatibia distinctly hooked dor- sally. Basal extension of right paramere largely lobi- form.G. arunachalensis Ahrens & Fabrizi 30’ Ventral apical spine of metatibia not hooked. Basal ex¬ tension of right paramere small and sharply pointed. .G. lubosi Ahrens & Fabrizi Gynaecoserica aniniensis Ahrens & Fabrizi, 2011 Gynaecoserica aniniensis Ahrens & Fabrizi, 2011: 152. Material examined. See Ahrens & Fabrizi 2011 (p. 152). Aedeagus. See Ahrens & Fabrizi 2011 (fig. 4I-K, p. 150). Distribution. Endemic to Arunachal Pradesh (Fig. 42F). Gynaecoserica annuliforceps Ahrens & Fabrizi, 2009 Gynaecoserica annuliforceps Ahrens & Fabrizi, 2009b: 256. Material examined. See Ahrens & Fabrizi 2009b (p. 256). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 3P-R, p. 279). Distribution. Endemic to western Arunachal Pradesh (Fig. 42F). Gynaecoserica arunachalensis Ahrens & Fabrizi, 2009 Gynaecoserica arunachalensis Ahrens & Fabrizi, 2009a: 1511. Material examined. See Ahrens & Fabrizi 2009a (p. 1524). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 1G-I, p. 1512). Distribution. Endemic to western Arunachal Pradesh (Fig. 42F). Gynaecoserica bicolorala Ahrens & Fabrizi, 2009 Gynaecoserica bicolorata Ahrens & Fabrizi, 2009a: 1508. Material examined. See Ahrens & Fabrizi 2009a (p. 1528). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 1L-N, p. 1512). Distribution. Endemic to the West Garo Hills (Megha¬ laya state) (Fig. 42E). Gynaecoserica bomdilana Ahrens & Fabrizi, 2009 Gynaecoserica bomdilana Ahrms & Fabrizi, 2009a: 1509. Material examined. See Ahrens & Fabrizi 2009a (p. 1509). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 2A-C, p. 1513). Distribution. Endemic to western Arunachal Pradesh (Fig. 42F). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 37 Gynaecoserica compacta Ahrens & Fabrizi, 2009 Gynaecoserica compacta Ahrens & Fabrizi, 2009a: 1508. Material examined. See Ahrens & Fabrizi 2009a (p. 1533). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 2D-F, p. 1513). Distribution. Endemic to the West Garo Hills (Megha¬ laya state) (Fig. 42D). Gynaecoserica cymosa (Brenske, 1896) Chaetoserica cymosa Brenske, 1896: 154; Brenske 1898: 313. Gynaecoserica cymosa'. Ahrens 1996: 34, 1999b: 183, 2004b: 104; Ahrens & Fabrizi 2009a: 1535; Ozgiil- Siemund & Ahrens 2015: 172. Paragynaecoserica pubescem Khan & Ghai, 1982: 63; Ahrens 1996: 34. Material examined. See Ahrens 1996 (p. 34), 2004b (p. 104); Ahrens & Fabrizi 2009a (p. 1535); 4 ex. “Kurseong 1904 R.P. Verschraeghen” (ISNB), 1 ex. “Inde” (ISNB), 2 ex. “Kurseong Indes” (ISNB), 1 ex. “Indes or.” (ISNB), 1 ex. “Coll. R.I.Sc.N.B./India: Bengal Darjeeling 1904 R.R Verschraeghen ex coll. De Moffarts” (ISNB). Aedeagus. See Ahrens 1996 (figs 78-80, p. 46). Distribution. Darjeeling-Sikkim area (Fig. 42F). Gynaecoserica dirangensis Ahrens & Fabrizi, 2009 Gynaecoserica dirangensis Ahrens & Fabrizi, 2009a: 1511. Material examined. See Ahrens & Fabrizi 2009a (p. 1539). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 2M-0, p. 1513). Distribution. Endemic to western Arunachal Pradesh (Fig. 42F). Gynaecoserica etalinensis sp. n. (Figs 2E-H, 26G, 42D) Type material examined. Holotype S “NE India, Arunachal Pr. Etalin vicinity, 700 m 28°36 , 56”N 95°53 , 21”E, L. Dembicky leg., 12.-25.V.2012” (ZFMK). Paratypes: 1 $ “NE India, Arunachal Pr. Etalin vicinity, 700 m 28°36 , 56”N 95°53 , 21”E, L. Dembicky leg., 12.- 25.V.2012” (ZFMK), 1 S “X-DA3472 - India, Arunachal Pradesh, Hunli, 1300m, 28°19 , 32”N 95°57 , 31”E, 26.v.- l.vi.2012, leg. L. Dembicky” (ZFMK), 1 $ “X-DA3473 - India, Arunachal Pradesh, Hunli, 1300m, 28°19 , 32”N 95°57’31”E, 26.v-l.vi.2012, leg. L. Dembicky” (ZFMK), 1 S “X-DA3474 - India, Arunachal Pradesh, Hunli, 1300m, 28°19’32”N 95°57’31”E, 26.v.-l.vi.2012, leg. L. Dembicky” (ZFMK), 1 S “X-DA3475 - India, Arunachal Pradesh, Hunli, 1300m, 28°19’32”N 95°57 , 31”E, 26.v.- l.vi.2012, leg. L. Dembicky” (ZFMK), 1 “X-DA3476 - India, Arunachal Pradesh, Hunli, 1300m, 28°19 , 32”N 95°57 , 31”E, 26.v-l.vi.2012, leg. L. Dembicky” (ZFMK), 1 S “X-DA3477 - India, Arunachal Pradesh, Hunli, 1300m, 28°19’32”N 95 0 57’31”E, 26.v.-l.vi.2012, leg. L. Dembicky” (ZFMK), 1 $ “834620-Arunachal Pradesh, Etalin vicinity, 800±100m, 28°35’N, 95°52’E L. Dembi¬ cky leg., l-3.vi.2007 Gynaecoserica sp.” (ZFMK). Description. Length: 4.8 mm, length of elytra: 3.4 mm, width: 3.0 mm. Body oblong, dorsal surface light brown, frons and pronotum somewhat greenish, sides of elytra darker, antenna yellowish, except shiny head dorsal sur¬ face dull, sparsely setose. Labroclypeus widest shortly before base, lateral margins moderately convex, anteriorly convergent to strongly rounded anterior angles, lateral border and ocular canthus producing a distinct blunt angle; margins weakly reflexed; anterior margin shallowly sinuate medially; surface near¬ ly flat and shiny, finely and very densely punctate, dis¬ tance between punctures less than their diameter, with a few coarser punctures behind anterior margin each bear¬ ing a long, erect seta; frontoclypeal suture feebly incised and medially moderately curved; smooth area in front of eye little wider than long; ocular canthus short and slen¬ der, finely and densely punctate, without terminal seta. Frons shiny, only posteriorly narrowly dull, with fine, dense punctures, glabrous except some erect setae beside eyes. Eyes large, ratio of diameter/interocular width: 0.73. Antenna yellow, with ten antennomeres; club dark, with five antennomeres, 1.4 times as long as the remaining an¬ tennomeres combined. Mentum weakly elevated and flat¬ tened anteriorly. Pronotum narrow, widest at base, lateral margins straight and subparallel in basal half, in anterior half weak¬ ly curved and convergent anteriorly, anterior angles mod¬ erately produced and sharp, posterior angles blunt; ante¬ rior margin convexly produced medially, with a distinct fine marginal line, basal margin without marginal line; an¬ terior and lateral margins sparsely setose; surface with dense and fine punctures, except on disc with scattered fine white short or moderately long setae in punctures, all setae bent backwards; hypomeron distinctly carinate at base but not ventrally produced. Scutellum narrow and short, triangular, with sparse, fine punctures, on base me¬ dially smooth, punctures with microscopic setae. Elytra moderately long, widest shortly at middle, stri¬ ae weakly impressed, finely and densely punctate; inter- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 38 Dirk Ahrens & Silvia Fabrizi vals weakly convex, with fine and moderately dense punc¬ tures concentrated along striae, punctures with fine white adjacent setae, odd intervals with a few single robust, white, adjacent setae; interior apical angle of elytra with a strong seta; epipleural edge fine ending at strongly curved external apical angle of elytra; epipleura sparsely setose, apical border without short microtrichomes. Ventral surface dull, with fine and moderately dense punctures, sparsely setose; metacoxa only laterally with a few strong adjacent setae. Each abdominal sternite with indistinct transverse row of coarse punctures bearing short setae between fine, dense punctation, penultimate stern¬ ite apically with a shiny smooth sclerotised border, which is one sixth as long as sternite, last sternite medially al¬ most half as long as penultimate one. Mesosternum be¬ tween mesocoxae as wide as mesofemur, with irregular¬ ly scattered very strong setae. Ratio of length of metepis- temum/metacoxa: 1/1.5. Pygidium weakly convex, coarse¬ ly and densely punctate, with fine smooth midline, dull, with numerous long setae. Legs slender and moderately long; femora dull, with two longitudinal rows of setae, finely and sparsely punctate; metafemur shiny, sharply margined anteriorly and with¬ out a submarginal serrate line, posterior margin weakly convex and glabrous, its external part only weakly widened in apical half and not serrate, internally finely ser¬ rate, with short setae. Metatibia slender and moderately long, uniformly widened toward apex, ratio width/length: 1/3.0, dorsally sharply carinate, with two groups of spines, basal one shortly before middle, apical one at three quar¬ ters of metatibial length, basally with a few single, fine setae; external face longitudinally convex, with sparse and moderately coarse punctures, glabrous; ventral margin car¬ inate and serrate, with three strong spines, of which the two distal are widely separated; internal face very finely and sparsely punctate; apex interiorly near tarsal articu¬ lation sharply and deeply truncate. Tarsomeres dorsally glabrous and impunctate, ventrally with sparse, short se¬ tae; metatarsomeres ventrally with a strongly serrate ridge, beside which is a fine longitudinal carina; first metatar- somere as long as following two tarsomeres combined and twice as long as dorsal tibial spur. Protibia moderately long, bidentate, protarsal claws symmetrical. Aedeagus: Fig. 2E-H. Variation. Length: 4.8—4.9 mm, length of elytra: 3.3-3.4 mm, width: 2.8-3.0 mm. Female: Antennal club composed of three antennomeres, slightly shorter than remaining an- tennomeres combined. Diagnosis. Gynaecoserica etalinensis sp. n. is in the gen¬ eral shape of the parameres similar to G. lohitensis Ahrens & Fabrizi, 2009, both sharing the basally strongly pro¬ duced parameres. G. etalinensis may be differentiated from G. lohitensis by the antennal club being composed of only five antennomeres (instead of six as in G. lohiten¬ sis), the slightly longer distal part of the parameres, and the large lateral tooth on the left paramere which is en¬ tirely absent in G. lohitensis. Etymology. The new species is named according to the type locality Etalin (adjective in the nominative singular). Distribution. See map (Fig. 42D). Gynaecoserica exilis Ahrens & Fabrizi, 2009 Gynaecoserica exilis Ahrens & Fabrizi, 2009a: 1511. Material examined. See Ahrens & Fabrizi 2009a (p. 1540). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 2P-R, p. 1513). Distribution. Endemic to western Arunachal Pradesh (Fig. 42E). Gynaecoserica fulgida (Arrow, 1946) comb. n. Leuroserica fulgida Arrow, 1946a: 281; Ahrens 2004b: 150. Material examined. See Ahrens 2004b (p. 150). Aedeagus. See Ahrens 2004b (figs 222-223, p. 414). Distribution. Endemic to the Sikkim-Darjeeling area (Fig. 42F). Gynaecoserica gogonaica Ahrens, 1999 Gynaecoserica gogonaica Ahrens, 1999b: 184. Material examined. See Ahrens 1999b (p. 184). Aedeagus. See Ahrens 1999b (figs 1-2, p. 185). Distribution. So far known only from the type locality in Bhutan (Fig. 42D). Gynaecoserica ignobilis Ahrens & Fabrizi, 2009 Gynaecoserica ignobilis Ahrens & Fabrizi, 2009a: 1510. Material examined. See Ahrens & Fabrizi 2009a (p. 1548). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 3A-C, p. 1514). Distribution. Endemic to Meghalaya (Fig. 42E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 39 Gynaecoserica keithi Ahrens & Fabrizi, 2009 Gynaecoserica keithi Ahrens & Fabrizi, 2009a: 1510. Material examined. See Ahrens & Fabrizi 2009a (p. 1551). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 3G-I, p. 1514). Distribution. Endemic to Meghalaya (Fig. 42D). Gynaecoserica lateralis (Arrow, 1946) comb. n. (Figs 26F, 42D) Leuroserica lateralis Arrow, 1946a: 282; Ahrens 2004b: 151. Material examined. See Ahrens 2004b (p. 151); 17 SS, 20 $$ “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E , 500-950m, L. Dembicky leg., 29.iv.-24.¥2005” (CPPB), 3 ©9 “NE India, Meghalaya, 6.-12.V.2002, 3 km E Tura, 1150 m, 25°30‘N, 90°14 E , M. Tryzna & P. Benda lgt.” (CPPB), 1 S, 2 $$ “NE In¬ dia, Meghalaya SW of Cherrapunjee, 25°13’-15 , N 91°40’E; 500-900 m; L. Dembicky leg., 11.-12.V.2004” (CPPB), 1 ex. “India Arunachal Pradesh 0.3km SSE of Di- rang, 27.iv.-l.v.2008, 1550m, 27° 20 , 32 ,, N, 91° 16’17”E, Podalska, Sipek lgt.” (ZFMK). Aedeagus. See Ahrens 2004b (figs 224, 225, p. 414). Distribution. Endemic to the Meghalaya area (Fig. 42D). Gynaecoserica latesquamosa (Frey, 1975) Lasioserica latesquamosa Frey, 1975b: 230. Lasioserica latesquamata [sic!] Frey: Ahrens 1996: 31. Gynaecoserica latesquamosa : Ahrens 2004b: 110. Material examined. See Ahrens 1996 (p. 31). Aedeagus. See Ahrens 1996 (figs 68-70, p. 45). Distribution. So far known only from the type locality in Bhutan (Fig. 42D). Gynaecoserica lobiceps Ahrens & Fabrizi, 2009 Gynaecoserica lobiceps Ahrens & Fabrizi, 2009a: 1510; Ozgul-Siemund & Ahrens 2015: 172. Material examined. See Ahrens & Fabrizi 2009a (p. 1553). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 3J-L, p. 1514). Distribution. Endemic to Meghalaya (Fig. 42E). Bonn zoological Bulletin 65 (1 & 2): 1-355 Gynaecoserica lohitensis Ahrens & Fabrizi, 2009 Gynaecoserica lohitensis Ahrens & Fabrizi, 2009a: 1510. Material examined. See Ahrens & Fabrizi 2009a (p. 1555). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 3M-0, p. 1514). Distribution. Endemic to the Assam area (Fig. 42D). Gynaecoserica lubosi Ahrens & Fabrizi, 2009 Gynaecoserica lubosi Ahrens & Fabrizi, 2009a: 1511. Material examined. See Ahrens & Fabrizi 2009a (p. 1559). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 4A-C, p. 1515). Distribution. Endemic to Arunachal Pradesh (Fig. 42E). Gynaecoserica marginipes (Brenske, 1898) Serica marginipes Brenske, 1896: 155. Microserica marginipes'. Brenske 1898: 318; Ahrens 2004: 186; Shrestha et al. 2012: 381. Neoserica marginipes'. Sabatinelli 1993: 628. Gynaecoserica marginipes : Ahrens & Fabrizi 2009a: 1564. Material examined. See Ahrens 2004b (p. 186); Shrestha et al. 2012 (p. 381). Aedeagus. See Ahrens 2004b (figs 287-289, p. 425). Distribution. In the hilly and mountainous regions from eastern Nepal to Bhutan (Fig. 42E). Gynaecoserica pellecta Brenske, 1896 Gynaecoserica pellecta Brenske, 1896: 154; Brenske 1898: 235, 314; Ahrens 1996: 34, 2004b: 106, 2006a: 410. Serica minutula Brenske, 1894 (nee Heer, 1862): 11, 50; Brenske 1898: 235. Gynaecoserica minutula: Ahrens 1996: 34, 1999b: 183. Material examined. See Ahrens 2004b (p. 106), 2006a (p. 410); 9 ex. “Darjeeling 1904 R.P Verschraeghen” (ISNB), 56 ex. “Kurseong 1904 R.P. Verschraeghen” (ISNB), 19 ex. “Coll. R.I.Sc.N.B./India: Bengal Darjee¬ ling 1904 R.P. Verschraeghen ex coll. De Moffarts” (ISNB), 1 ex. “Sikkim” (ISNB). Aedeagus. See Ahrens 1996 (figs 75-77, p. 46). Distribution. Eastern central Nepal to Bhutan (Fig. 42E). ©ZFMK 40 Dirk Ahrens & Silvia Fabrizi Gynaecoserica perdita Ahrens, 2004 Gynaecoserica perdita Ahrens, 2004b: 105. Material examined. See Ahrens 2004b (p. 105). Aedeagus. See Ahrens 2004b (figs 159-161, p. 404). Distribution. Endemic to Sikkim (Fig. 42E). Gynaecoserica seinghkuensis Ahrens & Fabrizi, 2009 Gynaecoserica seinghkuensis Ahrens & Fabrizi, 2009: 1577. Material examined. See Ahrens & Fabrizi 2009 (p. 1577). Aedeagus. See Ahrens & Fabrizi 2009 (fig. 5K-M, p. 1516). Distribution. Endemic to northern Myanmar and south¬ eastern Tibet, recorded close to the Indian border (Fig. 42D). Gynaecoserica singhikensis Ahrens, 2004 (Figs 26D, 42D) Gynaecoserica singhikensis Ahrens, 2004b: 109; Ozgiil- Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 109). Aedeagus. See Ahrens 2004b (figs 165-167, p. 405). Distribution. Endemic to eastern Sikkim (Fig. 42D). Gynaecoserica stemmleri (Frey, 1975) comb. n. Microserica stemmleri Frey, 1975b: 225. Leuroserica stemmleri. Ahrens 1995b: 55, 2004b: 151; Ahrens & Fabrizi 2009b: 272. Material examined. See Ahrens 1995b (p. 55), 2004b (p. 151); Ahrens & Fabrizi 2009b (p. 272). Aedeagus. See Ahrens 1995b (figs 55-57, p. 55). Distribution. Endemic to Bhutan and Arunachal Pradesh (Fig. 42D). Gynaecoserica tawangensis Ahrens & Fabrizi, 2009 Gynaecoserica tawangensis Ahrens & Fabrizi, 2009b: 257. Material examined. See Ahrens & Fabrizi 2009b (p. 257); 39 ex. “India: Arunachal Pradesh (11) 0.7km W of Tawang, 1-1.7km N of monastery, 27.-30.iv.2008, 27° 35’37”N, 91° 51 , 27”E, 2950m, Fikacek, Podalska, Sipek lgt.” (ZFMK), 1 ex. “India Arunachal Pradesh 0.3km SSE of Dirang, 27.iv.-l.v.2008, 1550m, 27° 20’32”N, 91° 16’17”JB, Podalska, Sipek lgt.” (ZFMK). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 4A-C, p. 150). Distribution. Endemic to western Arunachal Pradesh (Fig. 42D). Gynaecoserica tumba Ahrens, 2004 (Figs 26E, 42E) Gynaecoserica tumba Ahrens, 2004b: 107. Material examined. See Ahrens 2004b (p. 107). Aedeagus. See Ahrens 2004b (figs 162-164, p. 404). Distribution. Endemic to western Sikkim (Fig. 42E). Gynaecoserica variipennis variipennis (Moser, 1916) (Figs 26C, 42D) Lasioserica variipennis Moser, 1916: 153; Frey 1962a: 134. Gynaecoserica variipennis variipennis'. Ahrens 1996: 35, 1999b: 184,2004b: 107, 2006a: 411; Ahrens & Fabrizi 2009a: 1579; Shrestha et al. 2012: 380; Ozgiil-Siemu- nd & Ahrens 2015: 172. Material examined. See Ahrens 1996 (p. 35), 1999b (p. 184), 2004b (p. 107), 2006a (p. 411); Ahrens & Fabrizi 2009a (p. 1579); Shrestha et al. 2012 (p. 380); 2 ex. “Kur- seong 1904 R.P Verschraeghen” (ISNB), 1 ex. “Bengale Kurseong” (ISNB). Aedeagus. See Ahrens 1996 (figs 81-83, p. 47). Distribution. In the Himalaya, from eastern central Nepal to the Darjeeling-Sikkim area (Fig. 42D). Gynaecoserica variipennis wuermlii (Frey, 1972) Lasioserica wuermlii Frey, 1975b: 231. Gynaecoserica variipennis wuermlii'. Ahrens 1996: 37, 1999b: 184, 2004b: 107. Material examined. See Ahrens 1996 (p. 35), 1999b (p. 184), 2004b (p. 107). Aedeagus. See Ahrens 1996 (figs 84-86, p. 47). Distribution. Endemic to Bhutan (Fig. 42E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 41 Gynaecoserica victori Ahrens & Fabrizi, 2011 Gynaecoserica victori Ahrens & Fabrizi, 2011: 151 Material examined. See Ahrens & Fabrizi 2011 (p. 151). Aedeagus. See Ahrens & Fabrizi 2011 (fig. 4E-G, p. 150). Distribution. Endemic to Bhutan, known so far only from the type locality (Fig. 42D). Gynaecoserica vogleri Ahrens & Fabrizi, 2009 Gynaecoserica vogleri Ahrens & Fabrizi, 2009a: 1511. Material examined. See Ahrens & Fabrizi 2009a (p. 1580). Aedeagus. See Ahrens & Fabrizi 2009a (fig. 5N-P, p. 1516). Distribution. Endemic to western Arunachal Pradesh, known so far only from the type locality (Fig. 42D). Gynaecoserica yigongensis Liu & Ahrens, 2014 Gynaecosericayigongensis Liu & Ahrens, 2014: 160. Material examined. See Liu & Ahrens, 2014 (p. 160). Aedeagus. See Liu & Ahrens, 2014 (fig. 1I-K, p. 158). Distribution. Endemic to southeastern Tibet, known so far only from the type locality (Fig. 42D). Lasioserica Brenske, 1896 Lasioserica Brenske, 1896: 155 (type species by subse¬ quent designation: Serica nobilis Brenske, 1894; Arrow 1946b); Frey 1962: 132; Ahrens 1996: 1, 2000e: 1, 2004b: 111, 2005d: 1, 2006c: 1, 2007c: 17. Orchiserica Miyake & Yamaya, 2001: 38 (type species by subsequent designation: Lasioserica brevipilosa Moser, 1919; Ahrens 2004b: 112); syn. by Ahrens 2004b: 112. Remarks. In the original type species designation of Miyake & Yamaya (2001) the type species was misiden- tified as Lasioserica tuberculiventris Moser, 1915, thus the type species of Orchiserica was corrected by Ahrens (2004b). Key to Lasioserica species (<$$) 1 Protarsal claws asymmetrical. Metatarsomeres longi¬ tudinally sulcate dorsally, and carinate laterally. . L. dekensis Ahrens V Protarsal claws symmetrical. Metatarsomeres dorsal¬ ly smooth or weakly punctate.2 2 Labroclypeus short and wide (ratio length/width: 1/1.8), trapezoidal, with strongly convex anterior an¬ gles. L. breviclypeata Ahrens 2’ Labroclypeus longer and moderately wide (ratio length/width: 1/1.6).3 3 Dorsal surface sparsely setose. Head except for a few setae on labroclypeus and beside eyes glabrous.4 3’ Dorsal surface including head densely setose.23 4 Longitudinal serrated line of metatibia interrupted in apical third.5 4’ Longitudinal serrated line of metatibia not interrupt¬ ed.6 5 Median basal lobe of left paramere extended. Right paramere more strongly curved dorsally (lateral view). Lateral apical apophysis of phallobase ventral - ly weakly sinuate at base, its apical tooth medially strongly produced. Slightly pronounced pair of tuber¬ cles on apical phallobase present. . L. pacholatkoi Ahrens 5’ Median basal lobe of left paramere little extended. Right paramere nearly straight (lateral view). Lateral apical apophysis of phallobase ventrally strongly sin¬ uate at base, its apical tooth medially weakly produced. Slightly pronounced pair of tubercles on apical phal¬ lobase lacking. L. imminuta Ahrens & Fabrizi 6 Labroclypeus in basal third and frons dull. Antennal club as long as remaining antennomeres combined. . . L. assamicola Ahrens 6’ Labroclypeus entirely and at least distal third of frons shiny.7 7 Frons posteriorly dull. Lateral interval of elytra not convex.8 7’ Frons entirely shiny. Lateral interval of elytra convex. Phallobase with long apophysis.22 8 Disc of pronotum with a few short and thick setae. 9 8’ Disc of pronotum nearly glabrous.11 9 Phallobase without dorsal and lateral apophyses. Para- meres very different in length. L. nenya Ahrens 9’ Phallobase with a dorsal distal apophysis.10 10 Phallobase with a lateral distal apophysis on each side, ventrally without teeth. Parameres subequal in length. . L. hamifer Ahrens & Fabrizi 10’ Phallobase without a lateral distal apophysis on each side, ventrally with two paired teeth. Parameres very different in length. L. bispinosa Ahrens & Fabrizi 11 Sides of pronotum behind anterior angles concave. 12 11 ’ Sides of pronotum behind anterior angles convex of straight.13 12 Labroclypeus wider, rectangular. Antennal club 1.5 times as long as remaining antennomeres combined. . L. wittmeri Ahrens 12’ Labroclypeus only little wider than long, lateral mar¬ gins convex and strongly convergent anteriorly. An- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 42 Dirk Ahrens & Silvia Fabrizi tennal club as long as remaining antennomeres com¬ bined. L. nudosa Ahrens 13 Lateral margins of pronotum distinctly sinuate behind middle. L. beibengana Liu & Ahrens 13’ Lateral margins of pronotum distinctly not sinuate be¬ hind middle.14 14 Lateral margins of labroclypeus weakly convex and convergent anteriorly.15 14’ Lateral margins of labroclypeus strongly convex and convergent anteriorly.17 15 Labroclypeus convexly elevated medially. Body elon¬ gate. Frons only posteriorly narrowly dull. Phallobase without a long apical apophysis.16 15’ Labroclypeus not elevated medially. Body oval. Pos¬ terior half of frons dull. Phallobase with long apical apophysis. L. meghalayana Ahrens 16 Metatarsomere one shorter than following two tar- someres combined. Dorsal lobe of right paramere more than half as long as ventral one. Left paramere short, basal piece distinctly pronounced and shorter. . L. sabatinellii Ahrens 16’ Metatarsomere one as long as following two tar- someres combined. Dorsal lobe of right paramere much shorter than ventral one. Left paramere longer, basal piece less distinctly pronounced and shorter. ... . L. verschraegheni sp. n. 17 Phallobase ventrally with transversal elevations. ..18 17’ Phallobase ventrally without transversal elevations. .20 18 Ventral transversal elevations of phallobase large, api- cally concave, in lateral view lobus-like. Right para¬ mere without basal process. L. nepalensis Ahrens 18’ Ventral transversal elevations of phallobase small, not concave, in lateral view like a small tooth. Right para¬ mere with distinct basal process.19 19 Left paramere in dorsal view shorter than wide. Right paramere with a long basal process that is directed api- cally. L. godavariensis Ahrens 19’ Left paramere in dorsal view longer than wide. Right paramere with a short basal process that is directed ex¬ ternally and that may be covered under the left para¬ mere. L. dolakhana Ahrens 20 Left paramere simply convex dorsally, at apex with¬ out reflexed tooth. L. kulbei Ahrens 20’ Left paramere dorsally at apex with a reflexed tooth. .21 21 Right paramere sinuate on mesoventral margin, its basal process (only visible in ventral view) long and narrow, straight, directed distally and curved dorsoven- trally. Reflexed apical tooth of left paramere large. .. . L. modikholae Ahrens 21 ’ Right paramere not sinuate on mesoventral margin, its basal process (visible also in dorsal view) short, direct¬ ed externally. Reflexed apical tooth of left paramere small. L. dolangsae Ahrens 22 Pronotum at base wide, lateral margins convex behind anterior angles. L. nobilis (Brenske) 22’ Pronotum at base narrower, lateral margins straight be¬ hind anterior angles. L. sikkimensis Ahrens 23 Dorsal surface with long and erect pilosity. Phallobase almost always with apical apophysis. ( L. maculata group).24 23’ Dorsal surface with pilosity being bent posteriorly and more or less adpressed.28 24 All antennomeres of club equal in length. Right para¬ mere very long and narrow. . L. silkae Ahrens 24’ First antennomere of club 0.25 times as long as an¬ tennal club.25 25 Dorsal apical apophysis of parameres reduced.26 25’ Dorsal apical apophysis of parameres present, but sometimes short.27 26 Right paramere shortened and widened. Apex of phal¬ lobase at left side with an evenly narrowed process. . L. maculata jiriana Ahrens 26’ Right paramere not or little shortened. Apex of phal¬ lobase at left side with a process that is apically widened. L. maculata bhutanica Ahrens 27 Dorsal apophysis of phallobase small. Apex of phal¬ lobase at left side with an evenly narrowed process. . L. maculata galadrielae Ahrens 27’ Dorsal apophysis of phallobase large and robust. Apex of phallobase at left side without process. . L. maculata maculata (Brenske) 28 Dorsal pilosity fine or thick, but never scale-like. 29 28’ Dorsal pilosity dense and scale-like. . L. turaensis Ahrens 29 Pilosity on disc of pronotum thick and adpressed. Lat¬ eral margins behind anterior angles of pronotum con¬ vex.30 29’ Pilosity on disc of pronotum fine and bent posterior¬ ly, not adpressed.32 30 Phallobase in apical quarter ventrally on each side with an elevated lamella, between lamellae concave. Right paramere wide. L. pilosella Brenske 30’ Phallobase in apical quarter ventrally on each side without lamella.31 31 Right paramere moderately wide and only weakly curved externally. Left paramere small and strongly curved ventrally. L. piloselloida Ahrens 31 ’ Right paramere in apical half narrow and only strong¬ ly curved externally. Left paramere larger and not curved ventrally, slightly sinuate laterally. . L. pseudopilosella Ahrens 32 Penultimate abdominal stemite with a curved trans¬ verse carina. Lateral margins of pronotum behind an¬ terior angles convex. Pilosity on entire surface dense. . L. bumthangana Ahrens 32’ Penultimate abdominal sternite simple. Lateral mar¬ gins of pronotum behind anterior angles straight or concavely sinuate.33 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 43 33 Phallobase apically strongly narrowed, without later¬ al apical apophysis.34 33’ Phallobase apically weakly narrowed, more than half as wide as middle of phallobase (dorsal view), with a lateral apical apophysis.35 34 Parameres more than half as long as phallobase. An¬ tennal club slightly longer than remaining anten- nomeres combined. L. soror Ahrens 34’ Parameres distinctly less than half as long as phal¬ lobase. Antennal club little shorter than remaining an- tennomeres combined, first antennomere in club shortened. L. thoracica Brenske 35 Antennal club little longer than remaining anten- nomeres combined.37 35’Antennal club as long or slightly shorter than remain¬ ing antennomeres combined.36 36 Dorsal mesoapical sinuation of phallobase deep and narrow; apical apophysis of phallobase short and curved ventrally. L. ilamensis Ahrens 36’ Dorsal mesoapical sinuation of phallobase wide and short; apical apophysis of phallobase long and narrow. . L. orlovi Ahrens 37 Phallobase without any distal apophysis. . L. smithi Ahrens 37’ Phallobase with a distal apophysis on one or both sides. .38 38 Right paramere moderately long, apically simply rounded. Phallobase only at right side with short apophysis. L. braeti Brenske 38’ Right paramere short, apically nearly truncate. Phal¬ lobase at right and left side with a moderately long and narrow apophysis. L. chitreana Ahrens Lasioserica assamicola Ahrens, 2004 Lasioserica assamicola Ahrens, 2004b: 124; Ahrens 2006c: 53. Material examined. See Ahrens 2004b (p. 124). Aedeagus. See Ahrens 2004b (figs 185-187, p. 408). Distribution. Endemic to the Assam valley, so far known only from the type locality (Fig. 43A). Lasioserica beibengana Liu & Ahrens, 2014 Lasioserica beibengana Liu & Ahrens, 2014: 161. Material examined. See Liu & Ahrens 2014 (p. 161. Aedeagus. See Liu & Ahrens 2014 (figs 1A-C, p. 162). Distribution. Endemic to southern Tibet, so far known on¬ ly from the type locality (Fig. 43A). Lasioserica bispinosa Ahrens & Fabrizi, 2009 Lasioserica bispinosa Ahrens & Fabrizi, 2009b: 258. Material examined. See Ahrens & Fabrizi 2009b (p. 258). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 4D-F, p. 280). Distribution. Endemic to western Arunachal Pradesh, so far known only from the type locality (Fig. 43B). Lasioserica braeti Brenske, 1896 Lasioserica braeti Brenske, 1896: 155; Ahrens 1996: 8, 2000e: 7, 2004b: 121, 2006c: 53. Lasioserica maculatay ar. braeti : Brenske 1898: 310; Frey 1962a: 132. Material examined. See Ahrens 1996 (p. 8), 2000e (p. 7), 2004b (p. 121); 1 ex. “Kurseong Inde Verschraeghen 1904” (ISNB). Aedeagus. See Ahrens 1996 (figs 20-23, p. 42). Distribution. Endemic to the Darjeeling area (Himalaya) (Fig. 43C). Lasioserica breviclypeata Ahrens, 1999 Lasioserica breviclypeata Ahrens, 1999b: 232; Ahrens 2004b: 125, 2006c: 53. Material examined. See Ahrens 1999b (p. 232). Aedeagus. See Ahrens 1999b (figs 90-92, p. 233). Distribution. Endemic to Bhutan (Fig. 43C). Lasioserica bumthangana Ahrens, 1999 Lasioserica bumthangana Ahrens, 1999b: 225; Ahrens 2004b: 124, 2006c: 53. Material examined. See Ahrens 1999b (p. 225), 2004b (p. 124). Aedeagus. See Ahrens 1999b (figs 80-83, p. 227). Distribution. Endemic to Bhutan (Fig. 43B). Lasioserica chitreana Ahrens, 1999 (Figs 261, 43C) Lasioserica chitreana Ahrens, 1999b: 226; Ahrens 2000e: 7, 2004b: 119, 2006c: 53. Material examined. See Ahrens 1999b (p. 226), 2000e (p. 7), 2004b (p. 119). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 44 Dirk Ahrens & Silvia Fabrizi Aedeagus. See Ahrens 1999b (figs 84-86, p. 227). Distribution. Endemic to eastern Nepal (Fig. 43C). Lasioserica dekensis Ahrens, 1999 (Figs 26L, 43A) Lasioserica dekensis Ahrens, 1999b: 231; Ahrens 2004b: 125, 2006c: 53. Material examined. See Ahrens 1999b (p. 231). Aedeagus. See Ahrens 1999b (figs 87-89, p. 233). Distribution. Endemic to Sikkim and the Darjeeling area (Fig. 43A). Lasioserica dolakhana Ahrens, 2004 Lasioserica dolakhana Ahrens, 2004b: 116; Ahrens 2006c: 53. Material examined. See Ahrens 2004b (p. 116). Aedeagus. See Ahrens 2004b (figs 170-173, p. 406). Distribution. Endemic to central Nepal (Dolakha district) (Fig. 43C). Lasioserica dolangsae Ahrens, 2004 Lasioserica dolangsae Ahrens, 2004b: 117; Ahrens 2006c: 53. Material examined. See Ahrens 2004b (p. 117). Aedeagus. See Ahrens 2004b (figs 174-177, p. 406). Distribution. Endemic to central Nepal, known only from the type locality (Fig. 43C). Lasioserica godavariensis Ahrens, 1999 Lasioserica godavariensis Ahrens, 1999b: 220; Ahrens 2000e: 4, 2004b: 115, 2006c: 53. Material examined. See Ahrens 1999b (p. 220), 2000e (p. 4), 2004b (p. 115). Aedeagus. See Ahrens 1999b (figs 70-72, p. 221). Distribution. Endemic to central Nepal (Kathmandu val¬ ley) (Fig. 43C). Lasioserica hamifer Ahrens & Fabrizi, 2011 Lasioserica hamifer Ahrens & Fabrizi, 2011: 153. Material examined. See Ahrens & Fabrizi 2011 (p. 153). Bonn zoological Bulletin 65 (1 & 2): 1-355 Aedeagus. See Ahrens & Fabrizi 2011 (fig. 5A-C, p. 154). Distribution. Endemic to Bhutan, so far known only from the type locality (Fig. 43A). Lasioserica ilamensis Ahrens, 2000 Lasioserica ilamensis Ahrens, 2000e: 6; Ahrens 2004b: 119, 2006c: 53. Material examined. See Ahrens 2000e (p. 6). Aedeagus. See Ahrens 2000e (figs 7-9, p. 25). Distribution. Endemic to eastern Nepal, so far known on¬ ly from the type locality (Fig. 43D). Lasioserica (s.l.) imminuta Ahrens & Fabrizi, 2011 Lasioserica (s.l.) imminuta Ahrens & Fabrizi, 2011: 155. Material examined. See Ahrens & Fabrizi 2011 (p. 155). Aedeagus. See Ahrens & Fabrizi 2011 (fig. 5E-G, p. 154). Distribution. Endemic to Bhutan, so far known only from the type locality (Fig. 43A). Lasioserica kulbei Ahrens, 1999 Lasioserica kulbei Ahrens, 1999b: 218; Ahrens 2000e: 4, 2004b: 115, 2006a: 411, 2006c: 53. Material examined. See Ahrens 1999b (p. 218), 2000e: 4, 2004b (p. 115), 2006a (p. 411). Aedeagus. See Ahrens 1999b (figs 65,66, 68, 69, p. 219). Distribution. Endemic to central Nepal (Fig. 43B). Lasioserica maculata maculata (Brenske, 1896) (Figs 26J, 43A) Serica maculata Brenske, 1894: 11,49. Lasioserica maculata : Brenske 1898: 310; Frey 1962a: 132, 134; Ahrens 1996: 16. Lasioserica maculata maculata. Ahrens 1999b: 228, 2000e: 8, 2004b: 123, 2006a: 411, 2006c: 53; Ahrens & Fabrizi 2009b: 270; Shrestha et al. 2012: 380; Ozgiil- Siemund & Ahrens 2015: 172. Lasioserica pilosa Moser, 1919: 332; Frey 1962a: 132, 134, syn. by Ahrens 1996: 16. ? Omaloplia umbrina Blanchard, 1850: 77; Ahrens 1999b: 228. Material examined. See Ahrens 1996 (p. 16), 1999b (p. 228), 2000e: 8, 2004b (p. 123), 2006a (p. 411); Ahrens & Fabrizi 2009b (p. 271); Shrestha et al. 2012 (p. 380); 3 ©ZFMK Sericini of India 45 ex. ($) “Coll. R.I.Sc.N.B./ India: Himalaya Bhim Valley S. Breuning vendit” (ISNB). Aedeagus. See Ahrens 1996 (figs 4CM2, p. 44). Distribution. Himalaya, from east of the Sutlej river to central Nepal (Kathmandu valley) (Fig. 43A). Lasioserica maculata bhutanica Ahrens, 1996 Lasioserica bhutanica Ahrens, 1996: 19. Lasioserica maculata bhutanica: Ahrens 1999b: 230, 2004b: 123, 2006c: 53. Material examined. See Ahrens 1996 (p. 19). Aedeagus. See Ahrens 1996 (figs 48^19, p. 44). Distribution. Endemic to Bhutan (Fig. 43A). Lasioserica maculata galadrielae Ahrens, 1996 Lasioserica galadrielae Ahrens, 1996: 18. Lasioserica maculata galadrielae'. Ahrens 1999b: 230, 2000e: 8, 2004b: 123, 2006c: 53; Ahrens & Fabrizi 2009b: 270. Material examined. See Ahrens 1996 (p. 18); Ahrens & Fabrizi 2009b (p. 271). Aedeagus. See Ahrens 1996 (figs 46, 47, p. 44). Distribution. Endemic to eastern Nepal and the Darjee¬ ling area as well as western Sikkim (Fig. 43A). Lasioserica maculata jiriana Ahrens, 1996 Lasioserica jiriana Ahrens, 1996: 18. Lasioserica maculata jiriana. Ahrens 1999b: 229,2000e: 8, 2004b: 123, 2006a: 412, 2006c: 53; Ahrens & Fab¬ rizi 2011: 161; Shrestha et al. 2012: 380. Material examined. See Ahrens 1996 (p. 18), 2000e (p. 8), 2004b (p. 123), 2006a (p. 412); Ahrens & Fabrizi 2011 (p. 161); Shrestha et al. 2012 (p. 380). Aedeagus. See Ahrens 1996 (figs 43^15, p. 44). Distribution. Endemic to eastern central Nepal (Fig. 43A). Lasioserica meghalayana Ahrens, 1999 Lasioserica meghalayana Ahrens, 1999b: 224; Ahrens 2000e: 2, 2004b: 115, 2005d: 777, 2006a: 412, 2006c: 53. Material examined. See Ahrens 1999b (p. 224), 2000e (p. 2), 2004b (p. 115), 2005d: 777, 2006a (p. 412); 3 SS “NE India, Meghalaya, 8km N of Shillong, 1200m, 25°38‘ N, 91°54‘ E, 1200m, L. Dembicky leg., 7.-9.V.2004” (CPPB), 2 4 “Laos, Hua Phan prov., 6. - 18.v.2004b, 20°13‘ N 103°59‘ E, Ban Saluei, Phu Phan Mt. env., 1300 - 2000 m, J. Bezdek leg” (CPPB), 1 “NE India, Meghalaya, SW of Cher- rapunjee, 25°13‘-14‘ N, 91°40‘ E, 500-950m, L. Dem- bicky leg., 29.iv.-24.v.2004” (CPPB). Aedeagus. See Ahrens 2003b (figs 1-3, p. 68). Distribution. Meghalaya and northern Thailand (Fig. 43E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 50 Dirk Ahrens & Silvia Fabrizi Amis erica hunliana sp. n. (Figs 2M-P, 27B, 43E) Type material examined. Holotype: S “NE India Arunachal Pr. Hunli vicinity, 1300±100m 28°19 , 32”N 95°57 , 31”E, L. Dembicky leg., 26.v.-l.vi.2012 ,, (ZFMK). Description. Length: 5.4 mm, length of elytra: 3.9 mm, width: 3.1 mm. Body oblong, dark brown, around punc¬ tures light brown, surface dull, anterior head shiny, anten¬ na yellowish brown, dorsal surface partly with greenish shine, sparsely setose, with minute and white, scale-like setae on elytra and pronotum. Labroclypeus subtrapezoidal, widest at base, lateral mar¬ gins weakly convex and convergent to strongly rounded anterior angles, lateral border and ocular canthus produc¬ ing a distinct obtuse angle, anterior margin concavely sin¬ uate, margins weakly reflexed; surface flat and shiny, coarsely and densely punctate, with a few long erect se¬ tae behind anterior margin; frontoclypeal suture finely im¬ pressed and moderately curved; smooth area anterior to eye 1.5 times as wide as long. Ocular canthus moderate¬ ly long and narrow, finely and sparsely punctate, with a short terminal seta. Frons in posterior third dull, finely and moderately densely punctate, with a few short setae be¬ side eyes and an posterior third. Eyes very large, ratio of diameter/interocular width: 0.92. Antenna with ten anten- nomeres, club with three antennomeres, slightly reflexed externally, 1.7 times as long as remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum widest behind middle, lateral margins mod¬ erately convex and convergent posteriorly, more strong¬ ly curved anteriorly and convergent to acute, distinctly produced anterior angles, posterior angles nearly rectan¬ gular; anterior margin weakly convex, with a distinct mar¬ ginal line; basal margin without marginal line; surface with dense and coarse punctures each bearing a minute seta, along midline less densely punctate; with numerous short and dense, white, scale-like setae; anterior and lateral bor¬ ders sparsely setose; hypomeron carinate, basal margin of hypomeron not produced ventrally. Scutellum subtriangu- lar, apex sharp, with fine and dense punctures, smooth on basal midline, punctures with minute setae. Elytra oblong, widest at middle, striae distinctly im¬ pressed, with fine and dense punctures; intervals moder¬ ately convex, with fine and moderately dense punctures concentrated along striae, with sparse moderately long, white, scale-like setae, otherwise punctures with minute setae only; epipleural edge moderately strong, ending at strongly rounded external apical angle of elytra, epipleu- ra densely setose, apical border narrowly membraneous, with short microtrichomes. Ventral surface dull, with fine and dense punctures, sparsely setose, metacoxa only laterally with a few fine, adpressed setae. Abdominal sternites finely and densely punctate and minutely setose, each sternite with a distinct transverse row of coarse punctures each bearing a short, robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.72. Pygidium convex and dull, with coarse, dense punctures and fine, white setae, midline smooth. Legs moderately slender; femora finely densely punc¬ tate and glabrous, with two longitudinal rows of setae; an¬ terior edge of metafemur acute, with an adjacent contin¬ uously serrated line, posterior margin ventrally weakly widened in apical half but not serrate, dorsally complete¬ ly and finely serrate. Metatibia slender and short, widest at apex, ratio width/length: 1/3.2, weakly carinate dorsal¬ ly, with one group of spines only at 7/8 of metatibial length, basally with a few single spines in punctures, be¬ side dorsal margin with a continuously serrated line con¬ vergent with dorsal margin behind apical group of spines, between serrated line and dorsal densely and fine¬ ly punctate and shortly setose in punctures; lateral face longitudinally convex, with dense and coarse punctures bearing each a minute seta, along middle narrowly smooth; ventral edge serrated, with four fine and long, equidistant spines; medial face impunctate and glabrous; apex inte¬ riorly near tarsal articulation weakly concavely sinuate. Tarsomeres dorsally finely densely punctate and with minute setae, ventrally with short, sparse setae. Metatar- someres ventrally with a strongly serrated ridge, lateral¬ ly not carinate; first metatarsomere slightly longer than fol¬ lowing two tarsomeres combined and a quarter of its length longer then dorsal tibial spur. Protibia short, biden- tate, protarsal claws symmetrical, basal tooth of inner claw simply pointed. Aedeagus: Fig. 2M-P. Habitus: Fig. 27B. Female un¬ known. Diagnosis. The new species is very similar to A. maw- phlangensis Ahrens, but it differs by the parameres being sharply pointed at the apex, while the parameres in A. mawphlangensis are bluntly truncate. Etymology. The new species is named after the type lo¬ cality, Hunli (adjective in the nominative singular). Distribution. See map (Fig. 43E). Amis erica insperata (Brenske, 1898) Lasioserica insperata Brenske, 1898: 357. Amiserica insperata : Ahrens 1996: 29; Ahrens & Fabrizi 2011: 161. Material examined. See Ahrens 1996 (p. 29); Ahrens & Fabrizi 2011 (p. 161). Aedeagus. See Ahrens 1996 (figs 66, 67, p. 45). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 51 Distribution. Khasi Hills and northern Myanmar (Fig. 43E). Amiserica krausei Ahrens, 2004 (Figs 26P, 43E) Amiserica krausei Ahrens, 2004b: 130; Ahrens 2006a: 412; Ahrens & Fabrizi 2011: 162; Shrestha et al. 2012: 380; Ozgiil-Siemund & Ahrens 2015: 172. Material examined. See Ahrens 2004b (p. 130), 2006a (p. 412); Ahrens & Fabrizi 2011 (p. 162); Shrestha et al. 2012 (p. 380). Aedeagus. See Ahrens 2004b (figs 194-196, p. 409). Distribution. Kumaon Himalaya, western and western central Nepal (Fig. 43E). Amiserica longiflabellata Ahrens, 2004 (Figs 27A, 43E) Amiserica longiflabellata Ahrens, 2004b: 134. Material examined. See Ahrens 2004b (p. 134). Aedeagus. See Ahrens 2004b (figs 200-202, p. 410). Distribution. Endemic to Sikkim, so far known only from the type locality (Fig. 43E). Amiserica lutulenta sp. n. (Figs 2Q-S, 27C, 43E) Type material examined. Holotype: 11 mm). . N. ziyardamensis sp. n. 6 Right paramere much shorter than the left one. . N. astuta sp. n. 6’ Both parameres of nearly equal length.7 7 Parameres long, distinctly exceeding more than the half of phallobasal length. N. parilis sp. n. 7’ Parameres shorter, not exceeding the half of phal¬ lobasal length. N. exoleta Ahrens & Fabrizi 8 Phallobase behind middle with distinct pair-wise ven¬ tral processes.9 8’ Phallobase without any distinct pair-wise ventral processes.12 9 Lamina-like processes very small, much shorter than phallobase (at this point) wide. ... N. nilgiriana sp. n. 9’ Lamina-like processes large, subequal or at least half as long as phallobase (at this point) wide.10 10 Lamina-like processes subequal at least half of para¬ mere length.11 10’ Lamina-like processes much shorter than half of para¬ mere length. N. kalkadensis sp. n. 11 Lamina-like processes robust, at apex half as wide (lat¬ eral view) as high. N. mudigereensis sp. n. IE Lamina-like processes finer, at apex 1/6 as wide (lat¬ eral view) as high. N. kejvali sp. n. 12 Both parameres with distinct lateral teeth externally on each side.13 12’ Parameres without lateral teeth externally.14 13 Antennal club with five antennomeres. Parameres strongly curved externally, with one apical tooth and one tiny tooth at middle on each external side of para¬ mere. N. munnarensis sp. n. 13’ Antennal club with six antennomeres. Parameres straight, with one apical tooth and one tiny, more basal tooth (positioned subapically) on each external side of paramere. N. aulica sp. n. 14 Parameres as long as phallobase. ...N. gravida sp. n. 14’ Parameres half as long as phallobase. . N. barberi (Sharp) Neoserica astuta sp. n. (Figs 4V-X, 281, 45C) Type material examined. Holotype: S “NE India, Meghalaya state West Garo Hills, Nokrek Nat. Park 9- 17.V.1996 alt. 1100+150m GPS N25°29.6’, E90°19.5 (WGS 84) E. Jendek & O. Sausa/ IS 40” (CPPB). Paratypes: 3 SS “NE India, Meghalaya state West Garo Hills, Nokrek Nat. Park 9-17. V. 1996 alt. 1100+150m GPS N25°29.6’, E90°19.5 (WGS 84) E. Jendek & O. Sausa/ IS 42/ 98 Sericini Asia spec.” (ZFMK, CPPB). Description. Length: 14.0 mm, length of elytra: 10.0 mm, width: 8.0 mm. Body oblong, dark brown, punctures on elytra reddish brown, antennal club brown, anterior labro¬ clypeus shiny, dorsal surface dull, sparsely setose. Labro¬ clypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and slightly con¬ vergent anteriorly, anterior angles weakly rounded, ante¬ rior margin weakly sinuate medially, margins moderate¬ ly reflexed; surface flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind anterior margin with coarse punctures each bearing a long erect Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 74 Dirk Ahrens & Silvia Fabrizi seta; frontoclypeal suture indistinctly incised, flat and dis¬ tinctly curved medially; smooth area anterior to eye ap¬ proximately 1.5 times as wide as long; ocular canthus moderately long (length = nearly 1/2 of ocular diameter) and slender, impunctate, terminal seta lacking in holotype. Frons dull, with fine and sparse punctures, beside the eyes with two erect setae. Eyes small, ratio diameter/interoc¬ ular width: 0.51. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Labrum distinctly produced medially, with a deep median sinuation and two blunt an¬ terior teeth. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins in basal half nearly straight and moderately convergent, in anterior half evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt, slightly round¬ ed at tip; anterior margin slightly convex, with distinct and complete marginal line; surface densely and finely punc¬ tate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly cari¬ nate, but carina only weakly produced. Scutellum short, triangular, with fine, moderately dense punctures, basal¬ ly at middle smooth, with only minute setae. Elytra oblong, apex slightly truncate, widest at poste¬ rior third, striae weakly impressed, finely and moderate¬ ly densely punctate, even intervals flat, odd ones slight¬ ly convex, with moderately dense, fine punctures being on odd intervals concentrated along striae, intervals with a few fine white adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at blunt external apical angle of elytra, epipleura slightly re¬ flexed, densely setose, apical border narrowly membra¬ neous, with a very fine fringe of microtrichomes (visible at lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.79. Pygidium weakly convex and shiny, coarsely and densely punctate, without smooth midline, with a few short setae beside the apical margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between the rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row near¬ ly completely lacking, posterior margin in apical half ven- trally smooth and slightly widened, posterior margin dor- sally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ra¬ tio of width/length: 1/3.8, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basal¬ ly with a few robust but single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medi¬ al face smooth, apex moderately concavely sinuate inte¬ riorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral ca¬ rina immediately beside it, first metatarsomere distinctly longer than following two tarsomeres combined and lit¬ tle longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4V-X. Habitus: Fig. 281. Female un¬ known. Diagnosis. Neoserica astuta sp. n. is in its external shape and genital morphology rather similar to N. alloputaoana Ahrens et al., 2014. The species differs from N. allop¬ utaoana by the right paramere, consisting of a single lobe only (not two). Etymology. The new species is named with the Latin ad¬ jective ‘ astutus ’ (cunning, artful) (adjective in the nomi¬ native singular). Variation. Length: 13.4-14.0 mm, length of elytra: 10.0-10.6 mm, width: 7.9-8.4 mm. Distribution. See map (Fig. 45C). Neoserica aulica sp. n. (Figs 5A-C, 28J, 45C) Type material examined. Holotype: S “India: Mysore Shimoga dist., Agumbe Ghat, 2000’, V.1990 T.R.S. Nathan” (CMNC). Description. Length: 14.5 mm, length of elytra: 10.0 mm, width: 8.0 mm. Body oblong-oval, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, dis¬ tinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior an¬ gles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface nearly flat and shiny, basis with dull toment, punctation dense, an- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 75 teriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; fronto- clypeal suture indistinctly incised, distinctly curved me¬ dially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside eyes with two single setae. Eyes small, ratio diam¬ eter/interocular width: 0.54. Antenna with ten anten- nomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomeres two to four subequal in length. Mentum elevated and slightly flat¬ tened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest shortly before base, lateral margins evenly convex and strongly convergent anteriorly, anterior angles sharp and distinctly produced, posterior angles blunt; anterior mar¬ gin nearly straight, marginal line widely interrupted me¬ dially; surface densely and finely punctate with minute se¬ tae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, carina weakly pro¬ duced. Scutellum moderately long, triangular, with fine, dense punctures, with only minute setae. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals weakly convex, with unevenly dense, fine punctures con¬ centrated along striae, intervals with a few fine white very short setae, otherwise only with very minute setae in punc¬ tures; epipleural edge fine, ending at strongly rounded ex¬ ternal apical angle of elytra, epipleura sparsely setose, api¬ cal border membranous, with a fine fringe of microtri- chomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.88. Pygidium moderate¬ ly convex and dull, coarsely and densely punctate, with a narrow smooth midline, with a few single setae beside apical margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ pletely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.9, sharply carinate dorsally, with two groups of spines, basal group just before middle, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on basal half; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex mod¬ erately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere slightly longer than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5A-C. Habitus: Fig. 28J. Diagnosis. Neoserica aulica sp. n. differs from N. bar- beri by the slightly larger body and the more robust male genitalia having parameres with external teeth (absent in N. barberi). Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived form Latin adjective, L aulicus\ (princely), with reference of its impressive body size. Distribution. See map (Fig. 45C). Neoserica barberi (Sharp, 1903) comb. n. (Figs 5D-F, 28K, 45C) Melaserica barberi Sharp, 1903: 469. Type material examined. Syntypes: 1 $ “ Melaserica barberi Type D. S. Ootacamund/ Type/ Ootacamund C. A. Barber/ Sharp Coll. 1905-313” (BMNH), 1 $ “Melaserica barberi D. S. Ootacamund Barber 1902/ Sharp Coll. 1905-313” (BMNH), 1 $ “S. India/ Cotype” (BMNH), 1 $ “Cotype/ Malabar/ Melaserica barberi $ D. S.” (BMNH), 1 9 jf Melaserica barberi ^ Type D. S. Ootacamund” (BMNH), 1 S “India Malabar/ 24570/ Fry Coll. 1905-100” (BMNH), 1 $ “Dodabetta Nilgiris 8.000 ft. 1-3-V-16 Ramakrishna Coll./ S-India Coimbatore Mus. 1916-140” (BMNH), 1 S “Nilgiris Hills G. F. Hampson/ Cotype Melaserica barberi S D. S.” (BMNH). Additional material examined. 1 ex. “Nilgiris Hills T. V. Campbell/ G. C. Champion Coll. B.M. 1927-409” (BMNH), 1 ex. “Madras xi. 1907 ex. Tomlin/G. C. Cham¬ pion Coll. B.M. 1927-409” (BMNH), 1 ex. “Ootacamund India ex. Tomlin.//G. C. Champion Coll. B.M. 1927-409” (BMNH), 4 ex. “Nilgiris Hills/ Nilgiris Hills H. L. An- drewes/Andrewes Bequest B.M. 1922-221” (BMNH), 6 ex. “India S. Tamil Nadu, Nilgiris, Odty hills st., 11°25‘N 76°42‘E, 2250 m, 7.-8.V.2000, leg. P. Pacholatko” Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 76 Dirk Ahrens & Silvia Fabrizi (CPPB), 3 ex. “India-Tamil Nadu, Pakyra, Nilgiri Hills, 2250m, 11 0 26,9’N 70°36,9’E, leg. M. Halada, 26.4.2005” (CPPB). Redescription. Length: 10.8 mm, length of elytra: 8.1 mm, width: 6.5 mm. Body oblong-oval, dark brown, an¬ tennal club and ventral surface yellowish brown, anteri¬ or labroclypeus shiny, dorsal surface dull, on head and ely¬ tra sparsely setose. Labroclypeus subtrapezoidal, distinct¬ ly wider than long, widest at base, lateral margins straight but convergent anteriorly, anterior angles moderately rounded, anterior margin distinctly sinuate medially, mar¬ gins moderately reflexed; surface nearly flat and shiny, ba¬ sis with dull toment, punctation dense, anteriorly more sparse, behind the anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indis¬ tinctly incised, distinctly angled medially; smooth area an¬ terior to eye approximately 1.5 times as wide as long; oc¬ ular canthus moderately long (length = 1/3 of ocular di¬ ameter) and slender, very finely punctate, with a fine ter¬ minal seta. Frons dull, with fine and dense punctures, be¬ side eyes and posteriorly at middle with a few long setae. Eyes large, ratio diameter/interocular width: 0.74. Anten¬ na with ten antennomeres, club with five antennomeres, straight, slightly longer than remaining antennomeres combined, antennomere five slightly transversely pro¬ duced; antennomere three slightly longer than anten¬ nomere 2 and 4. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, subtrapezoidal, widest at base, lat¬ eral margins evenly convex and strongly convergent an¬ teriorly, anterior angles nearly rectangular and distinctly produced, posterior angles blunt; anterior margin nearly straight, marginal line widely interrupted medially; sur¬ face densely and finely punctate with minute setae in punc¬ tures; setae of anterior and lateral border sparse; hy- pomeron basally distinctly carinate, carina weakly pro¬ duced. Scutellum moderately long, triangular, with fine, dense punctures that are at middle of base less dense, with only minute setae. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals weakly convex, with dense, fine punctures concentrated along striae, intervals with a few fine very short setae, oth¬ erwise only with very minute setae in punctures; epipleur- al edge fine, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membranous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.5. Pygidium moderate¬ ly convex and dull, finely and densely punctate, with a nar¬ row smooth midline, with a few single setae beside api¬ cal margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ pletely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia narrow and moderately long, widest at apex, ratio of width/length: 1/3.4, sharply carinate dorsal¬ ly, with two groups of spines, basal group just before mid¬ dle, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on later¬ al face present on basal half; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventral¬ ly and a sharp subventral carina immediately beside it, first metatarsomere slightly shorter than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5D-F. Habitus: Fig. 28K. Remarks. Antennal club in female only slightly shorter, composed of four antennomeres, antennomere six sube¬ qual a quarter of club length, antennomere five cylindri¬ cal. Distribution. See map (Fig. 45C). Neoserica exoleta Ahrens & Fabrizi, 2009 Neoserica exoleta Ahrens & Fabrizi, 2009b: 264. Material examined. See Ahrens & Fabrizi, 2009 (p. 264). Aedeagus. See Ahrens & Fabrizi, 2009 (fig. 5J-L). Distribution. The species is so far known only from the type locality (Fig. 45C). Neoserica gravida sp. n. (Figs 5G-I, 28L, 45C) Type material examined. Holotype: $ “Kanara S. India./ Andrewes Bequest. B.M. 1922-221./ Kanara/ 644 Serici- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 77 ni Asia spec” (BMNH). Paratypes: 7 8 $$ “India, Karnataka, 20 km SE Sagar, 600m, 14°06,37’N 75°08,93’E, M. Halada leg., 12.V.2005” (CPPB, ZFMK), 2 SS, 1 ? “India, Karnataka, Sylya, Medikeri, 150m, 12°32’N 75°29’E, M. Halada leg., 24.iv.2005” (CPPB, ZFMK). Description. Length: 14.5 mm, length of elytra: 11.3 mm, width: 9.8 mm. Body oblong-oval, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, dis¬ tinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior an¬ gles moderately rounded, anterior margin weakly sinuate medially, margins moderately reflexed; surface convexly elevated medially, shiny, basis with dull tomentum, punc- tation dense, anteriorly more sparse, behind anterior mar¬ gin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside eyes with three single setae. Eyes small, ratio diameter/interocular width: 0.54. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined; antennomeres two to four subequal in length. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, subtrapezoidal, widest at base, lat¬ eral margins evenly convex and strongly convergent an¬ teriorly, anterior angles sharp and distinctly produced, pos¬ terior angles blunt; anterior margin nearly straight, fine marginal line not interrupted medially; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally dis¬ tinctly carinate, carina weakly produced. Scutellum mod¬ erately long, triangular, with fine, dense punctures, along midline widely impunctate, with only minute setae in punctures. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals flat, with unevenly dense, fine punctures concentrated along striae, punctures on elytra reddish brown, intervals with a few fine white very short setae, otherwise only with very minute setae in punctures; epipleural edge fine, end¬ ing at strongly rounded external apical angle of elytra, epi- pleura sparsely setose, apical border membranous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.57. Pygidium moderate¬ ly convex and dull, coarsely and densely punctate, with¬ out smooth midline, with a few single setae beside apical margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ pletely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group just before middle, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on basal half; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex mod¬ erately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5G-I. Habitus: Fig. 28L. Diagnosis. Neoserica gravida sp. n. differs from N. bar- beri by the slightly larger body and the extremely long parameres being nearly as long as the phallobase. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the Latin adjective ‘ gravidus ’ (full/heavy), with reference to the voluminous body shape. Variation. Length: 14.0-14.5 mm, length of elytra: 10.1-11.3 mm, width: 8.5-9.8 mm. Female: Antennal club composed of four antennomeres, antennomere six slight¬ ly transversely produced, club slightly shorter than the re¬ maining antennomeres combined; eyes as large as in male. Distribution. See map (Fig. 45C). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 78 Dirk Ahrens & Silvia Fabrizi Neoserica infamiliaris sp. n. (Figs 5J-M, 28M, 45C) Type material examined. Holotype: S “Assam W.F. Bad- gley 1906-185” (BMNH). Paratypes: 1 S “Assam W.F. Badgley 1906-185” (ZFMK), 3 SS “NE India, Megha¬ laya state, Jaintia Hills reg., Jowai 6-8.VI.1996 alt. 1350+100 m, GPS N25°27’ E92°12’ (WGS 84) E. Jen- dek & O. Sausa leg./ IS 27/ 647 Sericini Asia spec.” (CPPB, ZFMK). Description. Length: 10.4 mm, length of elytra: 8.2 mm, width: 6.6 mm. Body oval, dark brown, dull, with light greenish shine, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface sparsely setose. Labro- clypeus subtrapezoidal, little wider than long, widest at base, lateral margins strongly convex and convergent an¬ teriorly, anterior angles moderately rounded, anterior mar¬ gin deeply sinuate medially, margins moderately reflexed; surface flat, shiny, basis with dull toment, punctation dense, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinct¬ ly incised, distinctly curved medially; smooth area ante¬ rior to eye as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, without terminal seta. Frons dull, with fine and dense punctures, with sparse long erct setae. Eyes moderately large, ratio diameter/interocular width: 0.57. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined, first joint of club only half as long as club, antennomeres two to four subequal in length. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, subtrapezoidal, widest at base, lat¬ eral margins weakly convex and strongly convergent an¬ teriorly, anterior angles rounded at apex and weakly pro¬ duced, posterior angles blunt; anterior margin straight, marginal line distinct and complete; surface densely and coarsely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally dis¬ tinctly carinate, carina not produced. Scutellum short, tri¬ angular, with coarse, dense punctures, with only minute setae in punctures. Elytra very wide, widest shortly behind middle, striae strongly impressed, finely and densely punctate, intervals convex, with dense, fine punctures concentrated along stri¬ ae, intervals with very sparse, fine, short setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at weakly rounded external apical angle of elytra, epipleura sparsely setose, apical border chitinous, without a fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum with sparse setae; metacoxa glabrous, with a few short setae laterally, posterior margin weakly con¬ vex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a short seta. Mesosternum be¬ tween mesocoxae half as wide as slender mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/1.63. Pygid- ium moderately convex at apex and dull, finely and dense¬ ly punctate, without a smooth midline, with sparse single short setae. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ plete, posterior margin in apical half ventrally smooth and very weakly widened, posterior margin dorsally distinct¬ ly serrated, on its basal portion glabrous. Metatibia slen¬ der and long, widest at apex, ratio of width/length: 1/4.0, sharply carinate dorsally, with two groups of spines, basal group shortly behind first third, apical group at three quar¬ ters of metatibial length, basally with 1-2 shorter and ro¬ bust single setae; lateral face longitudinally convex, along middle slightly concave, finely and sparsely punctate, sub¬ dorsal longitudinal carina on lateral face present up to pos¬ terior third; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tar- someres ventrally with sparse, short setae, laterally and dorsally sharply carinate, impunctate dorsally; metatar- someres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately be¬ side it, first metatarsomere as long as following two tar- someres combined and nearly twice as long as dorsal tib- ial spur. Protibia long, bidentate; anterior claws symmet¬ rical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5J-M. Habitus: Fig. 28M. Female un¬ known. Diagnosis. The new species differs from all other taxa of the Neoserica abnormis group (Ahrens et al. 2014) by the weakly produced anterior angles of the pronotum and the deeply sinuate anterior margin of the labroclypeus. Etymology. The name (adjective in the nominative sin¬ gular) is derived from the combined Latin prefix c in-’( un- / not) and the adjective ‘ familiaris ’ (known/ familiar), with reference to the particular genital morphology. Variation. Length: 10.4-11.2 mm, length of elytra: 8.0-8.2 mm, width: 6.5-6.6 mm. Distribution. See map (Fig. 45C). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 79 Neoserica kalkadensis sp. n. (Figs 5N-P, 28N, 45C) Type material examined. Holotype: S “S India - Tamil Nadu, 7.iv.l999; Kalkad wild life sanctuary; Manjolai 8°15’N 77°27’E; 1000 m, Schintlmeister & Sinaev leg. / 636 Sericini Asia spec ” (CPPB). Paratypes: 4 SS “S In¬ dia - Tamil Nadu, 7.iv.l999; Kalkad wild life sanctuary; Manjolai 8°15’N 77°27’E; 1000 m, Schintlmeister & Sinaev leg” (CPPB, ZFMK), 1 S “S India - Kerala 13.iv.1997 7 km N Munar; 1740 m; Eravikulam nat. p. 10°09’N 77°04’E Schintlmeister & Siniaev leg.” (CPPB). Description. Length: 13.0 mm, length of elytra: 10.6 mm, width: 9.4 mm. Body egg-shaped, dark brown, antennal club yellowish brown, anterior labroclypeus shiny, dor¬ sal surface dull, sparsely setose. Labroclypeus subtrape- zoidal, wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior an¬ gles moderately rounded, anterior margin moderately sin¬ uate medially, margins moderately reflexed; surface weakly convex medially, shiny, basis with dull toment, punctation dense, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal su¬ ture indistinctly incised, distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with one ter¬ minal seta. Frons dull, with fine and dense punctures, be¬ side eyes with a 2-3 single setae. Eyes moderately large, ratio diameter/interocular width: 0.57. Antenna with ten antennomeres, club with five antennomeres, straight, as long as remaining antennomeres combined, antennomere five transversely produced, but not exceeding 1/7 of club length, antennomeres two to four subequal in length. Men- tum elevated and slightly flattened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly con¬ vergent anteriorly, anterior angles sharp and strongly pro¬ duced, posterior angles blunt; anterior margin nearly straight, robust marginal line complete; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, carina weakly produced. Scutellum moderately long, triangular, with fine, dense punctures, on midline narrowly impunctate, with only minute setae in punctures. Elytra very wide, widest shortly before posterior third, striae finely impressed, finely and moderately densely punctate, intervals nearly flat, with dense, fine punctures, intervals with a few fine, very short setae, otherwise on¬ ly with very minute setae in punctures; epipleural edge fine, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membra¬ nous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a short seta. Mesosternum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.54. Pygidium strongly con¬ vex at apex and dull, finely and densely punctate, with a fine smooth midline basally and a few single setae beside apical margin. Legs moderately slender; femora with two longitudinal rows of setae, finely and sparsely punctate between rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row completely lacking, posterior margin in apical half ventrally smooth and very weakly widened, posterior mar¬ gin dorsally distinctly serrated, on its basal portion glabrous. Metatibia wide and short, widest at apex, ratio of width/length: 1/3.1, sharply carinate dorsally, with two groups of spines, basal group shortly at one third, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent up to the posterior third; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventral¬ ly and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5N-P. Habitus: Fig. 28N. Female un¬ known. Diagnosis. Neoserica kalkadensis sp. n. differs from N. nilgiriana by the larger lamina-like processes of phal- lobase being nearly as high as wide (lateral view), while in N. nilgiriana it is not as high as wide. Etymology. The new species is named after its type lo¬ cality, the Kalkad wild life sanctuary (adjective in the nom¬ inative singular). Variation. Length: 13.0-16.4 mm, length of elytra: 10.6-12.5 mm, width: 9.4-10.6 mm. Distribution. See map (Fig. 45C). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 80 Dirk Ahrens & Silvia Fabrizi Neoserica kejvali sp. n. (Figs 5Q-S, 280, 45C) Type material examined. Holotype: S “S India, Kerala, Cardamom hills ca 50 km NW of Pathanamthitta near Pambaiyar riv., alt. 300 m/ 6.-9.V.1994 77°05’E 9°25’N Z. Kejval lgt. / 648 Sericini Asia spec.” (CPPB). Paratypes: 1 $ “S India, Kerala, Cardamom hills ca 50 km NW of Pathanamthitta near Pambaiyar riv., alt. 300 m/ 6.-9.V. 1994 77°05’E 9°25’N Z. Kejval lgt.” (CPPB), 3 SS “S-India, Kerala state, Kallar env., 30 km. NE of Trivandrum, vallay of riv. Kallar, 77°05’E 8°45’N, ca 300- 500 m, 7-13.V.1999 Z. Kejval & M. Tryzna leg” (CPPB, ZFMK). Description. Length: 12.4 mm, length of elytra: 8.9 mm, width: 7.4 mm. Body oval, dark brown, antennal club yel¬ lowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins moderate¬ ly convex and convergent anteriorly, anterior angles mod¬ erately rounded, anterior margin weakly sinuate medial¬ ly, margins moderately reflexed; surface flat, shiny, basis with dull toment, punctation dense, behind anterior mar¬ gin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with one terminal seta. Frons dull, with fine and dense punctures, beside eyes with a 2-3 single setae. Eyes mod¬ erately large, ratio diameter/interocular width: 0.71. An¬ tenna with ten antennomeres, club with six anten- nomeres, straight, 1.5 times as long as remaining anten¬ nomeres combined, antennomere four transversely pro¬ duced, but not exceeding 1/7 of club length, antennomeres two to four subequal in length. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and weakly con¬ vergent in basal half, strongly convergent in anterior half, anterior angles sharp and strongly produced, posterior an¬ gles blunt; anterior margin nearly straight, robust margin¬ al line widely incomplete medially; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally dis¬ tinctly carinate, carina weakly produced. Scutellum short, triangular, with fine, dense punctures, on midline very nar¬ rowly impunctate, with only minute setae in punctures. Elytra very wide, widest shortly before posterior third, striae finely impressed, finely and moderately densely punctate, intervals nearly flat, with dense, fine punctures being slightly concentrated along striae, intervals with a few fine, very short setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at weak¬ ly rounded external apical angle of elytra, epipleura sparse¬ ly setose, apical border membranous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal stemites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a short seta. Mesostemum between mesocoxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.63. Pygidium strongly con¬ vex at apex and dull, finely and densely punctate, with a fine smooth midline basally and a few single setae beside apical margin. Legs moderately slender; femora with two longitudinal rows of setae, finely and sparsely punctate between rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row completely lacking, posterior margin in apical half ventrally smooth and very weakly widened, posterior mar¬ gin dorsally distinctly serrated, on its basal portion glabrous. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.56, sharply carinate dorsally, with two groups of spines, basal group shortly at one third, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent up to the posterior third; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventral¬ ly and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres com¬ bined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5Q-S. Habitus: Fig. 280. Diagnosis. Neoserica kejvali sp. n. differs from N. mudi- gereensis by the long and fine lamina-like processes of phallobase being at its apex 1/6 as wide (lateral view) as high. Etymology. The new species is named after one of its col¬ lectors, Z. Kejval (noun in genitive case). Variation. Length: 12.4-13.6 mm, length of elytra: 8.9-9.7 mm, width: 7.4-8.1 mm. Female: body wider; an¬ tennal club composed of four antennomeres, sixth anten¬ nomere slightly transversely produced, club slightly shorter than the remaining antennomeres combined; eyes Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 81 slightly smaller than in male, ratio diameter/interocular width: 0.63. Distribution. See map (Fig. 45C). Neoserica mudigereensis sp. n. (Figs 5T-V, 28P, 45C) Type material examined. Holotype: S “India Karnata¬ ka Mudigere 27.5.1981 C-R J CR 61/ 660 Sericini Asia spec.” (BMNH). Paratypes: 1 tj, 1 § “India, Karnataka, 20 km SE Sagar, 600m, 14°06,37 , N 75°08,93’E, M. Ha- lada leg., 12.V.2005” (CPPB, ZFMK). Description. Length: 12.0 mm, length of elytra: 8.7 mm, width: 7.9 mm. Body nearly egg-shaped, dark brown, an¬ tennal club yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus sub- trapezoidal, wider than long, widest at base, lateral mar¬ gins moderately convex and convergent anteriorly, ante¬ rior angles moderately rounded, anterior margin moder¬ ately sinuate medially, margins moderately reflexed; sur¬ face convexly elevated medially, shiny, basis with dull to- ment, punctation dense, behind anterior margin with coarse punctures each bearing a long erect seta; fronto- clypeal suture indistinctly incised, distinctly curved me¬ dially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, terminal seta lacking in holotype. Frons dull, with fine and dense punctures, beside eyes with a two single setae. Eyes large, ratio diameter/interocular width: 0.74. Antenna with ten antennomeres, club with six antennomeres, straight, 1.2 times as long as remaining antennomeres combined, an¬ tennomeres two to four subequal in length. Mentum ele¬ vated and slightly flattened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins in basal half straight and subpar¬ allel, at middle convex and in anterior half straight and strongly convergent anteriorly, anterior angles sharp and strongly produced, posterior angles blunt; anterior mar¬ gin nearly straight, fine marginal line complete; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, carina weakly produced. Scutellum moderately long, triangular, with fine, dense punctures, on midline narrowly impunctate, with only minute setae in punctures. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals nearly flat, with dense, fine punctures concentrated along striae, intervals with a few fine white very short setae, oth¬ erwise only with very minute setae in punctures; epipleur- al edge fine, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membranous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.74. Pygidium strongly convex and dull, finely and densely punctate, with a fine smooth midline, with a few single setae beside apical mar¬ gin. Legs moderately slender; femora with two longitudinal rows of setae, finely and sparsely punctate between rows; metafemur dull, anterior margin acute, behind anterior edge without serrated line, setae of anterior longitudinal row completely lacking, posterior margin in apical half ventrally smooth and very weakly widened, posterior mar¬ gin dorsally distinctly serrated, on its basal portion glabrous. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.85, sharply carinate dorsally, with two groups of spines, basal group shortly at one third, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on basal third; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex mod¬ erately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere as long as following two tarsomeres com¬ bined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 5T-V. Habitus: Fig. 28P. Diagnosis. Neoserica mudigereensis sp. n. differs from N. kalkadensis by the larger lamina-like processes of the phal- lobase being subequal to half of the length of the para- mere. Etymology. The new species is named after its type lo¬ cality, Mudigere (adjective in the nominative singular). Variation. Length: 12.0-13.5 mm, length of elytra: 8.7-10.0 mm, width: 7.9-8.1 mm. Female: body wider; antennal club composed of four antennomeres, sixth an- tennomere slightly transversely produced, club slightly shorter than the remaining antennomeres combined; eyes as large as in male. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 82 Dirk Ahrens & Silvia Fabrizi Distribution. See map (Fig. 45C). Neoserica munnarensis sp. n. (Figs 5W-Y, 29A, 45C) Type material examined. Holotype: $ “S India, Kerala; 1250 m; 15 km SWMunnar; 1.-9.V.1997 10,02N 76,58E; Kallar Valley; Dembicky & Pacholatko leg./ IS 67/ 638 Sericini Asia spec” (CPPB). Paratypes: 1 2 99 “S In¬ dia, Kerala; 1250 m; 15 km SW Munnar; 1.-9.V.1997 10,02N 76,58E; Kallar Valley; Dembicky & Pacholatko leg.” (CPPB, ZFMK), 1 S, 2 9$ “Anamalai Hills Cin¬ chona. S. ind. 3500ft. 1959/giganteaBr.” (CF), 1 <$, 2 99 “India, Kerala Trivandrum Dt. Poonmudi Range 3000ft, IV.-V.71 leg. T.R.S. Nathan/ Meriserica oberthuri Br det. G. Frey, 1970” (CF). Description. Length: 13.5 mm, length of elytra: 9.6 mm, width: 8.1 mm. Body oblong-oval, dark brown, antennal club, abdomen including pygidium yellowish brown, an¬ terior labroclypeus shiny, dorsal surface dull, sparsely se¬ tose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent anteriorly, anterior angles moderately rounded, anterior margin weakly sinuate medially, mar¬ gins moderately reflexed; surface convexly elevated me¬ dially, shiny, basis with dull toment, punctation dense, an¬ teriorly more sparse, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal su¬ ture indistinctly incised, distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, with a fine terminal seta. Frons dull, with fine and dense punctures, beside eyes with a single seta. Eyes small, ratio diameter/interocular width: 0.5. Antenna with ten anten- nomeres, club with five antennomeres, straight, as long as remaining antennomeres combined, first joint of club half as long as the club, antennomere five slightly trans¬ verse; antennomeres two to four subequal in length. Men- tum elevated and slightly flattened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly con¬ vergent anteriorly, anterior angles sharp and distinctly pro¬ duced, posterior angles blunt; anterior margin nearly straight, robust marginal line not interrupted medially; sur¬ face densely and finely punctate with minute setae in punc¬ tures; setae of anterior and lateral border sparse; hy- pomeron basally distinctly carinate, carina weakly pro¬ duced. Scutellum moderately long, triangular, with fine, dense punctures, along midline punctures less dense, with only minute setae in punctures. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals weakly convex, with unevenly dense, fine punctures con¬ centrated along striae, punctures on elytra lighter, inter¬ vals with a few fine white very short setae, otherwise on¬ ly with very minute setae in punctures; epipleural edge fine, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membra¬ nous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal stemites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.63. Pygidium strongly convex and dull, finely and densely punctate, with a nar¬ row smooth midline, with a few single setae beside api¬ cal margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ pletely lacking, posterior margin in apical half ventrally smooth and slightly widened, posterior margin dorsally distinctly serrated, on its basal portion with a few short setae. Metatibia moderately slender and short, widest at apex, ratio of width/length: 1/3.1, sharply carinate dorsal¬ ly, with two groups of spines, basal group just before mid¬ dle, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on later¬ al face present on basal half; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres lacking in holotype. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply trun¬ cate at apex. Aedeagus: Fig. 5W-Y. Habitus: Fig. 29A. Diagnosis. Neoserica munnarensis sp. n. differs from N. aulica by the strongly curved parameres that have the more basal external tooth located at the middle instead of subapically as in N. aulica. Etymology. The new species is named according to its oc¬ currence close to Munnar (adjective in the nominative sin¬ gular). Variation. Length: 12.0-14.6 mm, length of elytra: 8.7-10.9 mm, width: 7.9-8.9 mm. Female: body wider; antennal club composed of four antennomeres, sixth an- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 83 tennomere slightly transversely produced, club slightly shorter than the remaining antennomeres combined; eyes as large as in male. Distribution. See map (Fig. 45C). Neoserica nilgiriana sp. n. (Figs 6A-C, 29B, 45C) Type material examined. Holotype: S “Nilgiri Hills. H.L. Andrewes./Andrewes Bequest. B.M. 1922-221./ 685 Sericini Asia spec.” (BMNH). Paratypes: 1 S, 1 $ “Coll. R.I.Sc.N.B./ India: Nilgiri Hills: Naduvatam PS. Nathan” (ISNB, ZFMK). Description. Length: 11.7 mm, length of elytra: 9.2 mm, width: 7.3 mm. Body oblong, dark brown, antennal club and posterior legs yellowish brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins moderately convex and convergent ante¬ riorly, anterior angles moderately rounded, anterior mar¬ gin moderately sinuate medially, margins moderately re- flexed; surface convexly elevated medially, shiny, basis with dull toment, punctation dense, behind anterior mar¬ gin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, terminal seta lacking in holotype. Frons dull, with fine and dense punctures, beside eyes with a single seta. Eyes small, ratio diameter/interocular width: 0.58. Antenna with ten antennomeres, club with six antennomeres, straight, as long as remaining antennomeres combined, first joint of club half as long as the club, antennomeres two to four subequal in length. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly con¬ vergent anteriorly, anterior angles sharp and distinctly pro¬ duced, posterior angles blunt; anterior margin nearly straight, fine marginal line indistinct medially; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally distinctly carinate, carina weakly produced. Scutellum moderately long, triangular, with fine, dense punctures, on midline broadly impunctate, with only minute setae in punctures. Elytra oblong, widest in posterior third, striae finely im¬ pressed, finely and moderately densely punctate, intervals weakly convex, with unevenly dense, fine punctures con¬ centrated along striae, punctures on elytra reddish brown, intervals with a few fine white very short setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membra¬ nous, with a fine fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.63. Pygidium strongly convex and dull, finely and densely punctate, without smooth midline, with a few single setae beside apical mar¬ gin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ pletely lacking, posterior margin in apical half ventrally smooth and very weakly widened, posterior margin dor- sally distinctly serrated, on its basal portion glabrous. Metatibia moderately slender and long, widest at apex, ra¬ tio of width/length: 1/3.4, sharply carinate dorsally, with two groups of spines, basal group shortly behind anteri¬ or third, apical group at three quarters of metatibial length, basally with 1-2 shorter and robust single setae; lateral face longitudinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on later¬ al face present on basal third; ventral edge finely serrat¬ ed, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, dorsally smooth; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventral¬ ly and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6A-C. Habitus: Fig. 29B. Diagnosis. Neoserica nilgiriana sp. n. differs from N. bar- beri by the very small paired laminae on the ventral phal- lobase, these are much shorter than the phallobase is (at this point) wide. Etymology. The new species is named ‘ nilgiriana ’ accord¬ ing to its occurrence in the Nilgiri Hills (adjective in the nominative singular). Variation. Length: 11.7-12.1 mm, length of elytra: 9.2-9.3 mm, width: 7.3-7.4 mm. Female: body slightly wider; antennal club composed of four antennomeres, Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 84 Dirk Ahrens & Silvia Fabrizi sixth antennomere slightly transversely produced, club slightly shorter than the remaining antennomeres com¬ bined; eyes as large as in male. Distribution. See map (Fig. 45C). Neoserica parilis sp. n. (Figs 6D-F, 29C, 45C) Type material examined. Holotype: S “NE India, Meghalaya state, Jaintia Hills reg., Jowai 6-8.VI. 1996 alt. 1350+100 m, GPS N25°27’ E92°12’ (WGS 84) E. Jen- dek & O. Sausa leg./ IS 27/ 647 Sericini Asia spec.” (CPPB). Description. Length: 12.7 mm, length of elytra: 10.9 mm, width: 7.7 mm. Body oblong, dark brown, antennal club brown, anterior labroclypeus shiny, dorsal surface dull, sparsely setose. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and slightly convergent anteriorly, anterior angles weakly rounded, anterior margin nearly straight, margins moderately reflexed; surface flat and shiny, basis with dull toment, punctation dense, anteriorly more sparse, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinctly incised, flat and distinctly curved medially; smooth area anterior to eye ap¬ proximately 1.5 times as wide as long; ocular canthus moderately long (length = nearly 1/2 of ocular diameter) and slender, impunctate, with a fine terminal seta. Frons dull, with fine and sparse punctures, beside the eyes with 2-3 erect setae. Eyes small, ratio diameter/interocular width: 0.5. Antenna with ten antennomeres, club with six antennomeres, straight, only slightly longer than remain¬ ing antennomeres combined, first joint of club subequal to 2/3 of club length. Mentum elevated and slightly flat¬ tened anteriorly. Labrum distinctly produced medially, with a deep median sinuation and two blunt anterior teeth. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins evenly convex and strongly con¬ vergent anteriorly, anterior angles sharp and distinctly pro¬ duced, posterior angles blunt, slightly rounded at tip; an¬ terior margin nearly straight, with distinct and complete marginal line; surface densely and finely punctate with minute setae in punctures; setae of anterior and lateral bor¬ der sparse; hypomeron basally distinctly carinate, but Ca¬ rina only weakly produced. Scutellum short, triangular, apex slightly rounded, with fine, moderately dense punc¬ tures, basally at middle smooth, with only minute setae. Elytra oblong, apex slightly truncate, widest at poste¬ rior third, striae weakly impressed, finely and moderate¬ ly densely punctate, even intervals flat, odd ones slight¬ ly convex, with moderately dense, fine punctures being on odd intervals concentrated along striae, intervals with a few fine white adpressed setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at the blunt external apical angle of elytra, epipleura slight¬ ly reflexed, densely setose, apical border thickened, nar¬ rowly membraneous, with a very fine fringe of microtri- chomes (visible at lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on the disc, glabrous on sides; metacoxa glabrous, with a few short se¬ tae laterally, posterior margin weakly convex; abdominal stemites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.52. Pygidium weakly convex and shiny, coarsely and densely punctate, without smooth midline, with a few semi-erect setae beside the api¬ cal margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between the rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row near¬ ly completely lacking, posterior margin in apical half ven- trally smooth and slightly widened, posterior margin dor- sally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ra¬ tio of width/length: 1/4.2, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basal¬ ly with a few robust but single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on about two third of metatibial length; ventral edge finely serrated, with three robust equidistant setae; medi¬ al face smooth, apex moderately concavely sinuate inte¬ riorly near tarsal articulation. Tarsomeres ventrally with sparse, short setae, laterally not carinate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral ca¬ rina immediately beside it, first metatarsomere distinctly longer than following two tarsomeres combined and 1/4 of its length longer than dorsal tibial spur. Protibia long, bidentate; anterior claws symmetrical, basal tooth of in¬ ner claw sharply truncate at apex. Aedeagus: Fig. 6D-F. Habitus: Fig. 29C. Diagnosis. Neoserica parilis sp. n. is in its external shape and genital morphology highly similar to N. putaoana Ahrens et al., 2014. N. parilis differs from N. putaoana by the much more sinuate parameres, with the left para- mere curved externally at its apex. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 85 Etymology. The new species is named with the Latin ad¬ jective ‘ parilis ’ (similar), with reference to the similarity with N. putaoana (adjective in the nominative singular). Distribution. See map (Fig. 45C). Neoserica pilistriata sp. n. (Figs 6G-I, 29D, 45C) Type material examined. Holotype S “NE India, Meghalaya, SWofCherrapunjee, 25°13‘-15‘N, 91°47‘E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB). Paratypes: 1 S “Megalaya 1976 Wittmer, BaroniU./Up¬ per Shillong 1900 m 13.5./ 601 Sericini: Asia spec.” (NHMB), 1 S “NE India, Meghalaya, SW of Cherrapun- jee, 25°13‘-15‘N, 91°47‘E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (ZFMK). Description. Length: 9.6 mm, length of elytra: 6.8 mm, width: 5.0 mm. Body oblong, dark brown, ventral surface including antennal club yellowish brown, labroclypeus reddish and shiny, dorsal surface dull, with dense, very long pilosity. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and slightly convergent anteriorly, anterior angles strongly rounded, anterior margin weakly sinuate medi¬ ally, margins moderately reflexed; surface flat and shiny, basis without dull toment, punctation moderately dense, anteriorly more sparse, with numerous coarse punctures each bearing a long erect seta; frontoclypeal suture indis¬ tinctly incised, flat and distinctly angled medially; smooth area anterior to eye approximately twice as wide as long; ocular canthus moderately long (length = nearly 1/3 of oc¬ ular diameter) and slender, impunctate, with a long ter¬ minal seta. Frons dull, with fine and dense punctures bear¬ ing each a short adpressed seta, interspersed with moder¬ ately dense, coarser punctures bearing each a long, erect seta. Eyes small, ratio diameter/interocular width: 0.7. An¬ tenna with ten antennomeres, club with five antennomeres, straight, as long as remaining antennomeres combined, an- tennomere five slightly transversely produced. Mentum el¬ evated and slightly flattened anteriorly. Labrum distinct¬ ly produced medially, with a moderately deep median sin- uation and two blunt anterior teeth on each side of it. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins in basal half nearly straight and subparallel, moderately curved at middle, in anterior third margins straight and strongly convergent anteriorly, an¬ terior angles moderately sharp and weakly produced, pos¬ terior angles blunt, distinctly rounded at tip; anterior mar¬ gin strongly convex, with robust, complete marginal line; surface with dense and fine punctures with minute to short setae interspersed with coarser punctures bearing each a long erect seta; setae of anterior and lateral border sparse and long; hypomeron basally distinctly carinate, carina not produced. Scutellum short, triangular, with fine, very dense punctures, basally at middle punctures impunctate, with only minute setae. Elytra oblong, apex slightly truncate, widest at poste¬ rior third, striae weakly impressed, finely and moderate¬ ly densely punctate, intervals slightly convex, with mod¬ erately dense, fine punctures concentrated along striae, in¬ tervals with dense, long, erect setae, otherwise only with very minute setae in punctures; epipleural edge fine, end¬ ing at strongly rounded external apical angle of elytra, epi- pleura densely setose, apical border chitinous, without a fringe of microtrichomes (visible at lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with setae on disc, glabrous on sides; metacoxa glabrous, with a few short setae lat¬ erally, posterior margin weakly convex; abdominal ster- nites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesosternum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Pygidium moderate¬ ly convex and shiny, coarsely and densely punctate, with¬ out smooth midline, with a few short setae beside the api¬ cal margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between the rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row near¬ ly completely lacking, posterior margin in apical half ven- trally smooth and slightly widened, posterior margin dor- sally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ra¬ tio of width/length: 1/4.1, sharply carinate dorsally, with two groups of spines, basal group just before middle, api¬ cal group at three quarters of metatibial length, basally with a few robust but single setae; lateral face longitudi¬ nally convex, very finely, superficially and sparsely punc¬ tate, subdorsal longitudinal carina on lateral face present on about two third of metatibial length; ventral edge fine¬ ly serrated, with three robust setae the posterior two more widely separated; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tar- someres ventrally with sparse, short setae, laterally not car¬ inate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventral¬ ly glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tar- someres combined and 1/4 of its length longer than dor¬ sal tibial spur. Protibia long, bidentate; anterior claws sym¬ metrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6G-I. Habitus: Fig. 29D. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Dirk Ahrens & Silvia Fabrizi Diagnosis. Neosericapilistriata sp. n. is in its genital mor¬ phology rather similar to N. natmatoungensis Ahrens et al., 2014, while both species differ strongly in their ex¬ ternal morphology: the parameres of N. pilistriata are shorter and wider, the surface is more densely setose, and the rim of microtrichomes at the apex of the elytra is ab¬ sent. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the combined Latin words, ‘pilus' (hair), and ‘ striatus ’ (striped), with reference to the longitudinal rows of long setae on the elytra. Variation. Length: 9.6-10.1 mm, length of elytra: 6.8-7.2 mm, width: 5.0-5.6 mm. Distribution. See map (Fig. 45C). Neoserica plagiata sp. n. (Figs 6J-L, 29E, 45C) Type material examined. Holotype: S “NE India, Meghalaya state, West Garo Hills, Nolcrek Nat. Park 9- 17.V.1996 alt. 1100+150 m, GPS N25°29,5’ E90°19,5’ (WGS 84) E. Jendek & O. Sausa leg.” (CPPB). Paratypes: 1 (j “NE India, Meghalaya state, West Garo Hills, Nolcrek Nat. Park 9-17.V.1996 alt. 1100+150 m, GPS N25°29,5’ E90°19,5’ (WGS 84) E. Jendek & O. Sausa leg.” (ZFMK), 3 SS, 2 $ ? “NE India, Meghalaya state, West Garo Hills reg., Tura 5-7.V. 1996 alt. 700+100 m, GPS N25°30,7’ E90°13,9’ (WGS 84) E. Jendek & O. Sausa leg.” (CPPB, ZFMK), 5 SS, 2 $$ “NE India, Megha¬ laya, 1 km E of Tura, 500-600m, 25°30‘N, 90°14‘E,2.- 5.V.2002, M.Tryzna & Benda leg.”(CPPB), 3 SS, 1 ? “NE India; Meghalaya; 1400 m; Nokrek n.p. 3km S Da- ribokgiri 25°27’N 90°19’E; 26.iv. 1999 Dembicky & Pa- cholatko leg.” (CPPB, ZFMK), 1 $ “NE India, Megha¬ laya state West Garo Hills, Nokrek Nat. Park 9-17. V. 1996 alt. 1100+150m GPS N25°29.6\ E90°19.5 (WGS 84) E. Jendek & O. Sausa/ IS 40” (CPPB), 5 §g “NE India, Meghalaya state, Jaintia Hills reg., Jowai 6-8.VI. 1996 alt. 1350+100 m, GPS N25°27’ E92°12’ (WGS 84) E. Jen¬ dek & O. Sausa leg./ IS 27/ 647 Sericini Asia spec.” (CPPB). Description. Length: 11.8 mm, length of elytra: 7.7 mm, width: 6.2 mm. Body oval, dark brown, dull, with light greenish shine, antennal club yellowish brown, anterior labroclypeus shiny, dorsal surface sparsely setose. Labro- clypeus subtrapezoidal, little wider than long, widest at base, lateral margins strongly convex and convergent an¬ teriorly, anterior angles moderately rounded, anterior mar¬ gin deeply sinuate medially, margins moderately reflexed; surface flat, shiny, basis with dull toment, punctation dense, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinct¬ ly incised, distinctly curved medially; smooth area ante¬ rior to eye as wide as long; ocular canthus moderately long (length = 1/3 of ocular diameter) and slender, glabrous, without terminal seta. Frons dull, with fine and dense punctures, with sparse long erct setae beside eyes, other¬ wise nearly glabrous. Eyes moderately large, ratio diam¬ eter/interocular width: 0.53. Antenna with ten anten- nomeres, club with six antennomeres, straight, as long as remaining antennomeres combined, first joint of club on¬ ly half as long as club, antennomeres two to four sube¬ qual in length. Mentum elevated and slightly flattened an¬ teriorly. Pronotum transverse, subtrapezoidal, widest at base, lat¬ eral margins weakly convex and strongly convergent an¬ teriorly, anterior angles rounded at apex and weakly pro¬ duced, posterior angles blunt; anterior margin straight, marginal line distinct and complete; surface densely and coarsely punctate with minute setae in punctures; setae of anterior and lateral border sparse; hypomeron basally dis¬ tinctly carinate, carina not produced. Scutellum short, tri¬ angular, with coarse, dense punctures, along the midline smooth, with only minute setae in punctures. Elytra very wide, widest shortly behind middle, striae strongly impressed, finely and densely punctate, intervals convex, with dense, fine punctures concentrated along stri¬ ae, intervals with very sparse, fine, short setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at weakly rounded external apical angle of elytra, epipleura sparsely setose, apical border chitinous, without a fringe of microtrichomes. Ventral surface dull, coarsely and densely punctate, metasternum with sparse setae; metacoxa glabrous, with a few short setae laterally, posterior margin weakly con¬ vex; abdominal sternites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a short seta. Mesosternum be¬ tween mesocoxae half as wide as slender mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/1.54. Pygid- ium strongly convex at apex, dull, finely and densely punc¬ tate, without a smooth midline, with numerous short se¬ tae, on apical portion also longer setae. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row com¬ plete, posterior margin in apical half ventrally smooth and very weakly widened, posterior margin dorsally distinct¬ ly serrated, on its basal portion glabrous. Metatibia slen¬ der and long, widest at apex, ratio of width/length: 1/3.9, sharply carinate dorsally, with two groups of spines, basal group shortly behind first third, apical group at three quar¬ ters of metatibial length, basally with 1-2 shorter and ro¬ bust single setae; lateral face longitudinally convex, along Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 87 middle slightly concave, finely and sparsely punctate, sub¬ dorsal longitudinal carina on lateral face present up to pos¬ terior third; ventral edge finely serrated, with three robust equidistant setae; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tar- someres ventrally with sparse, short setae, laterally and dorsally sharply carinate, impunctate dorsally; metatar- someres ventrally glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately be¬ side it, first metatarsomere slightly longer than following two tarsomeres combined and nearly twice as long as dor¬ sal tibial spur. Protibia long, bidentate; anterior claws sym¬ metrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6J-L. Habitus: Fig. 29E. Diagnosis. The new species differs from Neoserica in- familaris sp. n. in the presence of only one lamina on the ventral side of the phallobase, as well as in the stouter parameres. Etymology. The new species is named (adjective in the nominative singular) with the Latin adjective ‘ plagiatus ’ (plagiarized) with reference to the highly similar taxon N. infamilaris sp. n. to which N. plagiata seem to be a slight¬ ly modified copy. Variation. Length: 10.5-11.8 mm, length of elytra: 7.2-7.7 mm, width: 6.2-6.6 mm. Female: body as in male; antennal club composed of four antennomeres, 6^ anten- nomere slightly transversely produced, club slightly shorter than the remaining antennomeres combined; eyes as large as in male. Distribution. See map (Fig. 45C). Neoserica ziyardamensis sp. n. (Figs 6M-0, 29F, 45C) Type material examined. Holotype S “Myanmar N (Bur¬ ma) 65 km NE Putao, 1250 m, Zi Yar Dam vill. 18- 21.05.1998 leg. S. Murzin & V. Sinaev/ [ex] Coll. Takeshi Itoh, Osaka (Japan)/ 700 Sericini: Asia spec” (ZFMK). Description. Length: 11.4 mm, length of elytra: 8.2 mm, width: 6.4 mm. Body oblong, dark reddish brown, anten¬ nal club yellowish brown, anterior labroclypeus shiny, dor¬ sal surface dull and with moderately dense to sparse, long pilosity. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex and slightly convergent anteriorly, anterior angles strong¬ ly rounded, anterior margin weakly sinuate medially, mar¬ gins moderately reflexed; surface flat and shiny, basis without dull toment, punctation dense, anteriorly more sparse, behind anterior margin with coarse punctures each bearing a long erect seta; frontoclypeal suture indistinct¬ ly incised, flat and distinctly curved medially; smooth area anterior to eye approximately 1.5 times as wide as long; ocular canthus moderately long (length = nearly 1/2 of oc¬ ular diameter) and slender, impunctate, with a long ter¬ minal seta. Frons dull, with fine and sparse punctures, with dense, long, erect setae. Eyes small, ratio diameter/inte¬ rocular width: 0.51. Antenna with ten antennomeres, club with five antennomeres, straight, as long as remaining an¬ tennomeres combined, first joint of club less than half as long as club. Mentum elevated and slightly flattened an¬ teriorly. Labrum distinctly produced medially, with a shal¬ low median sinuation and two blunt anterior teeth. Pronotum moderately transverse, subtrapezoidal, widest at base, lateral margins in basal half nearly straight and subparallel, moderately curved at middle, in anterior third margins straight and strongly convergent anteriorly, an¬ terior angles sharp and distinctly produced, posterior an¬ gles blunt, distinctly rounded at tip; anterior margin straight, with distinct and complete marginal line; surface densely and finely punctate, interspersed with slightly coarser punctures bearing each a long erect seta, otherwise with minute setae in punctures only; setae of anterior and lateral border sparse; hypomeron basally distinctly cari¬ nate, but carina not produced. Scutellum short, triangu¬ lar, with coarse, very dense punctures, basally at middle punctures less dense, with only minute setae. Elytra oblong, apex slightly truncate, widest shortly be¬ hind middle, striae weakly impressed, finely and moder¬ ately densely punctate, intervals slightly convex, with moderately dense, fine punctures concentrated along stri¬ ae, intervals with sparse long erect setae, otherwise only with very minute setae in punctures; epipleural edge fine, ending at blunt external apical angle of elytra, densely se¬ tose, apical border narrowly membraneous, with a very fine fringe of microtrichomes (visible at lOOx magnifica¬ tion). Ventral surface dull, coarsely and densely punctate, metastemum sparsely covered with setae on disc, glabrous on sides; metacoxa glabrous, with a few short setae lat¬ erally, posterior margin weakly convex; abdominal ster- nites finely and unevenly densely punctuate, nearly glabrous, with a transverse row of coarse punctures, each bearing a robust short seta. Mesostemum between meso- coxae half as wide as slender mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Pygidium moderate¬ ly convex and shiny, coarsely and densely punctate, with¬ out smooth midline, with a few short setae beside the api¬ cal margin. Legs slender; femora with two longitudinal rows of se¬ tae, finely and sparsely punctate between the rows; metafe¬ mur dull, anterior margin acute, behind anterior edge with¬ out serrated line, setae of anterior longitudinal row near¬ ly completely lacking, posterior margin in apical half ven¬ trally smooth and slightly widened, posterior margin dor- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Dirk Ahrens & Silvia Fabrizi sally distinctly serrated, on its basal portion with a few short setae. Metatibia slender and long, widest at apex, ra¬ tio of width/length: 1/4.5, sharply carinate dorsally, with two groups of spines, basal group just before the middle, apical group at three quarters of metatibial length, basal- ly with a few robust but single setae; lateral face longitu¬ dinally convex, very finely, superficially and sparsely punctate, subdorsal longitudinal carina on lateral face pres¬ ent on about two third of metatibial length; ventral edge finely serrated, with three robust setae the posterior two widely separated; medial face smooth, apex moderately concavely sinuate interiorly near tarsal articulation. Tar- someres ventrally with sparse, short setae, laterally not car¬ inate, protarsomeres smooth, meso- and metatarsomeres with a few very fine punctures; metatarsomeres ventral¬ ly glabrous, with a strongly serrated ridge ventrally and a sharp subventral carina immediately beside it, first metatarsomere distinctly longer than following two tar- someres combined and 1/4 of its length longer than dor¬ sal tibial spur. Protibia long, bidentate; anterior claws sym¬ metrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 6M-0. Habitus: Fig. 29F. Diagnosis. Neoserica ziyardamensis sp. n. is in external shape and genital morphology rather similar to N. natma- toungensis Ahrens et al., 2014. The species differs in the left paramere being much shorter and strongly curved dor- sally instead of ventrally (as in N. natmatoungensis ). Etymology. The new species is named after its type lo¬ cality, Zi Yar Dam (= Ziyadam) (adjective in the nomina¬ tive singular). Remarks. This species was omitted by our mistake from the previous revision (Ahrens et al., 2014a), and is here included along with the Indian species, although being from Myanmar. Distribution. Northern Myanmar, however, the type lo¬ cality was not located with GPS data, therefore the species is not presented in our map (Fig. 45C). Neoserica dichroa group Diagnosis. Hypomeron ventrally without carina. Protib¬ ia bidentate. Antennal club in male with three or four an- tennomeres. Pygidium in female shiny, nearly glabrous. Key to species of the Neoserica dichroa group 0<3) 1 Antennal club with three antennomeres in both sexes. .2 1 ’ Antennal club with four antennomeres in males, and three antennomeres in females.5 2 Phallobase at apex without a dorsal process external of left paramere. Left paramere straight. . N. flavoviridis ( Brenske, 1896) T Phallobase at apex with a dorsal process external of left paramere.3 3 Dorsal process of phallobase short and sharply point¬ ed.4 3’ Dorsal process of phallobase long, subequal to left paramere, rounded at apex. N. plateosa sp. n. 4 Left apophysis of phallobase long and straight. . N. genieri sp. n. 4’ Left apophysis of phallobase very short. . N. plebea sp. n. 5 Phallobase with a left lateral distal apophysis of phal¬ lobase.6 5’ Phallobase without a left lateral distal apophysis of phallobase.7 6 Lateral apophysis long and straight. . N. submaculosa sp. n. 6’ Lateral apophysis short and curved ventrally. . N. ammattiensis sp. n. 7 Insertion of left paramere displaced at right side. Para- meres as long as phallobase. . N. rotundotibialis sp. n. 7’ Insertion of left paramere on left side. Parameres less than half as long as phallobase. ...N. dichroa Frey, 1973 Neoserica ammattiensis sp. n. (Figs 6P-R, 29G, 45D) Type material examined. Holotype S “Coll.R.I.Sc.N.B. South India S. Coorg. Ammatti 3100 ft. V-1951 PS. Nathan” (ISNB). Paratypes: 27 S8, 43 $$ “Coll.R.I.Sc.N.B. South India S. Coorg. Ammatti 3100 ft. V-1951 PS. Nathan” (ISNB, ZFMK). Description. Length: 5.1 mm, length of elytra: 3.9 mm, width: 3.3 mm. Body oval, yellowish brown, ventral sur¬ face, frons, disc of pronotum and numerous small spots on elytra dark brown and with greenish shine; dorsal sur¬ face dull and glabrous. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior an¬ gles, anterior margin weakly sinuate medially, margins weakly reflexed; surface weakly convex, moderately shiny, finely and densely punctate, with a few coarse punc¬ tures anteriorly bearing each a short erect seta; fronto- clypeal suture distinctly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a single terminal seta. Frons dull, Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 89 with sparse, fine punctures, with two single erect setae be¬ side each eye. Eyes moderately large; ratio of diameter/in¬ terocular width: 0.57. Antenna with ten antennomeres; club composed of four antennomeres, straight, slightly longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins weakly evenly convex and moderately convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at slightly blunt external apical angle of elytra, epi- pleura densely setose, apical border with a broad fringe of microtrichomes. Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.62. Abdominal stemites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and sparsely punctate, without smooth midline, with a few longer setae along api¬ cal margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.1; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group at middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and very sparsely punctate and glabrous, basally with dense coarse punctures, along middle smooth; ventral edge finely ser¬ rated, with three robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventral¬ ly on distal half, neither laterally nor dorsally carinate, dor- sally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina im¬ mediately beside it, glabrous; first metatarsomere slight¬ ly longer than two following tarsomeres combined, two thirds of its length longer than very short dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6P-R. Habitus: Fig. 29G. Diagnosis. Neoserica ammattiensis sp. n. is in its male genital morphology rather similar to N. flavoviridis but dif¬ fers from the latter by the antennal club being composed of four instead three antennomeres, as well as in the api¬ cal metatibial spines which are in N. ammattiensis much shorter than in N. flavoviridis. The apex of the right (ven¬ tral) paramere is more robust compared to that of N. flavoviridis , the left paramere (dorsal) is shorter and slight¬ ly wider. Etymology. This new species is named after its type lo¬ cality, Ammatti (adjective in the nominative singular). Variation. Length: 5.1-5.6 mm, length of elytra: 3.9^1.5 mm, width: 3.3-3.8 mm. Colour may vary from above de¬ scribed pattern towards the body being entirely dark brown. Female: Antennal club composed of three anten¬ nomeres, slightly shorter than remaining the anten¬ nomeres combined; eyes as large as in male. Distribution. See map (Fig. 45D). Neoserica dichroa Frey, 1973 (Figs 6S-U, 29H, 45D) Neoserica dichroa Frey, 1973: 248. Type material examined. Holotype: S “India, Kerala Tri¬ vandrum Dt. Poonmudi Range 3000 ft., IV.-V.71 leg. T. R. S. Nathan/ Type Neoserica dichroa G. Frey 1972” (CF). Paratype: 1 S “India, Kerala Trivandrum Dt. Poonmudi Range 3000 ft., IV.-V.71 leg. T. R. S. Nathan/ Paratype Neoserica dichroa G. Frey 1972” (CF). Additional material examined. 1 <$ “India, Kerala Trivandrum Dt. Poonmudi Range 3000 ft., IV.-V.71 leg. T. R. S. Nathan/ Paratype Neoserica dichroa G. Frey 1972” (CF), 1 ex. “India Trivandrum distr. Ponmudi rain forest 10.IV.1990 B. Gustafsson” (NHRS). Redescription. Length: 7.3 mm, length of elytra: 4.7 mm, width: 4.3 mm. Body oval, yellowish brown, ventral surface slightly dark¬ er, frons, disc of pronotum and numerous small spots on elytra dark and with greenish shine; dorsal surface dull and glabrous, partly moderately shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 90 Dirk Ahrens & Silvia Fabrizi gin weakly sinuate medially, margins weakly reflexed; sur¬ face weakly convex, moderately shiny, finely and dense¬ ly punctate, with a few coarse punctures anteriorly bear¬ ing each a short erect seta; frontoclypeal suture distinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a sin¬ gle terminal seta. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Eyes moder¬ ately large; ratio of diameter/interocular width: 0.61. An¬ tenna with ten antennomeres; club composed of four an- tennomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum moderately wide, widest at base, lateral mar¬ gins weakly evenly convex and moderately convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at slightly blunt external apical angle of elytra, epi- pleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.75. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and sparsely punctate, without smooth midline, with a few longer setae along api¬ cal margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide,.widest at half of metati- bial length, ratio width/length: 1/3.7; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and very sparse¬ ly punctate, along middle smooth, glabrous; ventral edge finely serrated, with three robust equidistant setae, medi¬ al face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on distal half, neither laterally nor dorsally car¬ inate, dorsally smooth; metatarsomeres with a strongly ser¬ rated ridge ventrally and a parallel subventral smooth Ca¬ rina immediately beside it, glabrous; first metatarsomere slightly longer than two following tarsomeres combined, one third of its length longer than dorsal tibial spur. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6S-U. Habitus: Fig. 29H. Distribution. See map (Fig. 45D). Neoserica flavoviridis (Brenske, 1896) comb. n. (Figs 6V-X, 291, 45D) Serica flavoviridis Brenske, 1896: 154. AutosericaJlavoviridis\ Brenske 1898: 239. Type material examined. Syntypes: 1 “Barway P. Car- don/ Serica flavoviridis typ. Brsk./ Type/ flavoviridis Brsk.” (ISNB), 1 $ “Barway P. Cardon/ Serica flavoviridis typ. Brsk./ Type/ flavoviridis Brsk.” (MSNG), 1 $ “Bar¬ way P. Cardon/ Museum Paris Inde Brenske 1898/ Cotype/ Serica flavoviridis Cotype Brsk.” (MNHN), 1 f “Barway P. Cardon/ coll. Brenske /flavoviridis type Brsk.” (ZMHB), 5 SS, 6 $ “Barway P. Cardon/ coll. Brenske” (ZMHB). Additional material examined. 43 SS, 27 $ $ “Coll. R.I.Sc.N.B. Inde/ Barway P. Cardon” (ISNB), 1 S “Ka- nara” (ZMHB), 1 f “Belgaum” (ZMHB), 3 SS, 1 ? “Ma- he Malabar” (ZMHB), 1 ex. “Matheran 800 m/ India or. Biro 1902” (HNHMB), 1 ex. “Mahe (Inde) Coll. R. Ley/ Comp, au type Musei du Bruxelles det. R. Ley 1923/ Ser¬ ica flavoviridis Brenske det. R. Ley 1923” (MNHN), 5 ex. “Mahe/ Le Moudt vendif ’ (ISNB), 1 ex “Malabaer/ Le Moudt vendit” (ISNB), 3 ex. “Malabar coll. R. Ley” (MNHN), 4 ex. “Coll. R.I.Sc.N.B. Inde/Mandar Beng. P. Cardon 7/91 Det. 92.” (ISNB), 1 ex. “India: Karnataka Balehonnur 21.5.1981/ CR 49/ Brit. Mus. 1984-37” (BMNH), 1 ex. “India: Karnataka Mudigere 22.5.1981 C.R. Coll./ CR 49/ Brit. Mus. 1984-37” (BMNH), 25 ex. “Kanara S. India/ Kanara/ Andrewes Bequest B. M. 1922- 231” (BMNH, CF), 9 ex. “Belgaum Bombay H. E. An¬ drewes/Andrewes Bequest B. M. 1922-221” (BMNH), 4 ex. “Kanara/ India 1900.20” (BMNH), 2 ex. “Kanara, S. India/ Kanara/ Andrewes Bequest B. M. 1922-221” (BMNH), 1 ex. “Belgaum” (BMNH), 2 ex. “Kanara” (BMNH), 1 ex. “India 85.43” (BMNH), 1 ex. “India T. B. Fletcher B. M. 1943-9/ Sidapur Coorg II.V.1917 T. R. N. Coll.”(BMNH), 1 ex. “India T. B. Fletcher B. M. 1943- 9/ Sidapur Coorg 12.IV.1917 T. R. N. Coll.” (BMNH), 1 ex. “India T. B. Fletcher B. M. 1943-9/ Sidapur Coorg 4- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 91 10.IV.1917 T. R. N. Coll.” (BMNH), 1 ex. “India T. B. Fletcher B. M. 1943-9/ Sidapur Coorg III. 1917 T. R. N. Coll.” (BMNH), 1 ex. “India T. B. Fletcher B. M. 1943- 9/ Sidapur Coorg 22.111.1917 T. R. N. Coll.” (BMNH), 1 ex. “India T. B. Fletcher B. M. 1943-9/ Coorg-Mercara 4500 ft. 27. Apr. 13. T. V. R. Coll.” (BMNH), 1 ex. “In¬ dia T. B. Fletcher B. M. 1943-9/Mercara-Coorg 20.V. 1917 T. R. N. Coll.” (BMNH), 2 ex. “Mahe Inde/ Brit. Mus. 1923-572/ compared with type/ Jlavoviridis Brenske det. R. Ley 1923” (BMNH), 1 ex. (?) “SouthMysore H.L. An- drewes/Andrewes Bequest B. M. 1922-221” (BMNH), 1 ex. “Mysore/ on Cashew Nut Plant” (BMNH), 28 ex. “In¬ dia, Mysore v.1973 Coorg distr. Mercara 4000 ft.T. R. S. Nathan leg.” (MHNG, CPPB), 9 ex. “India: Mysore Shi- moga dist., Agumbe Ghat, 2000’, VI. 1990 T.R.S. Nathan” (CMNC), 1 ex. “India Madras Coimbatore 1400 ft. XI.73 T. R. S. Nathan” (MHNG), 1 ex. “Kanara” (CF), 3 ex. “Kanara/ Coll. Kraatz”, 1 ex. “Barway P. Cardon/ C. v. Bennigsen”, 1 ex. “Belgaum” (CF), 2 ex. “Belgaum/ Coll. Kraatz” (DEIC), 7 ex. “Mahe Chass. indigenes Aout 1901/ Museum Paris Cote de Malabar Mahe M. Main- drom 1902/ Cote de Malabar M. Maindrom” (MNHN), 1 ex. “Maissour Shimoga Juin 1897” (MNHN), 1 ex. “Cote de Malabar T. Deschamps 1900” (MNHN), 1 ex. “S-In- dia, Karnataka state, Coorg distr., NE of Virajpet, 75°50‘E 12°13‘N, ca. 500 m, 4.-8.vi.l999, Z. Kejval & M. Tryz- na leg.” (CPPB), 1 ex. “S-India, Kerala state, Ponmudi hill resort, 30 km NE of Trivandrum, 77°06‘E 8°46‘N, ca. 1300-1500 m, 7.-13.v. 1999, Z. Kejval & M. Tryzna leg.” (CPPB), 1 ex. “S. India: Nilgiri Hills: Cherangode, 1050 m, V-50 P. S. Nathan” (BPBM), 1 ex. (?) “South India Kerala St. Quilon Distr., Thenmala V.1993, leg. Theresa Rajabai Selva Nathan” (CARL), 1 ex. ($) “South India Nilgiri Hills, Devala 3200 ft. V.1984, leg. Theresa Rajabai Selva Nathan” (CARL), 66 ex. “S India; Karnataka; W Ghats 18 km E Shiradi; Gundia; 12°47’N 75°43’E; 200- 500 m; P. Pacholatko leg. 16.-21 .v.2002” (CPPB, ZFMK), 6 ex. “India, Maharashta, 15kmN of Mahad, Raigarh Fort env., 250-500m, 18°14’N 73°26’E, 9.vi.2006, Z. Kejval lgt.” (CPPB), 15 ex. “India, Maharashta, 15kmN of Ma¬ had, near river, ca20mm, 18°12’N 73°24’E, 8.vi.2006, Z. Kejval lgt.” (CPPB), 2 ex. “India, Maharashta, ca 30km W of Pune, Mulshi env., 18°29’N 73°30’E, ca700m, 13.- 16.vi.2006, Z. Kejval lgt.” (CPPB), 3 ex. “India-Maha- rashtra state Western Ghat Mts. Raigardh, Mahad env. 7.- 9.VI.2006, O. Safranek leg.” (CPPB), 4 ex. “India, Ker¬ ala, Sabramila 12°32 , N75°29 , EM. Haladaleg., l.v.2005” (CPPB), 211 ex. “S India; Karnataka; W Ghats 18 km E Shiradi; Gunda; 12°47’N 75°43’E; 200-500m; P. Pa¬ cholatko; 16.-21.v.2002” (CPPB), 4 ex. “S India; Karnata¬ ka; W Ghats 20 km W Talguppa; Jog Falls; 14°14’N 74°44’E; 500±200m; P. Pacholatko; 22.-28.V.2002” (CPPB), 3 ex. “India, Maharahstra [sic] state, Alibag env., 45 km S Bombay, 22.-24.vi.2006, O. Safranek leg.” (CPPB). Redescription. Length: 6.2 mm, length of elytra: 4.1 mm, width: 3.5 mm. Body oval, yellowish brown, ventral sur¬ face, frons, disc of pronotum and numerous small spots on elytra dark brown and with greenish shine; dorsal sur¬ face dull and glabrous. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior an¬ gles, anterior margin weakly sinuate medially, margins weakly reflexed; surface weakly convex, moderately shiny, finely and densely punctate, with a few coarse punc¬ tures anteriorly bearing each a short erect seta; fronto- clypeal suture distinctly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with two single erect setae be¬ side each eye. Eyes moderately large; ratio of diameter/in¬ terocular width: 0.55. Antenna with ten antennomeres; club composed of three antennomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins weakly evenly convex and moderately convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, finely and densely punctate, on midline impunctate. Elytra oblong, widest at middle, striae distinctly im¬ pressed, finely and densely punctate, intervals strongly convex, with fine and dense punctures exclusive along stri¬ ae, at centre impunctate, with very minute setae in punc¬ tures, otherwise glabrous; epipleural edge robust, ending at slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.54. Abdominal stemites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium moderately convex and dull, coarsely and sparsely punc¬ tate, without smooth midline, with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 92 Dirk Ahrens & Silvia Fabrizi weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia moderately long and narrow, widest at half of metatibial length, ratio width/length: 1/3.5; dorsal margin distinctly longitudinally convex, with two groups of spines, basal group at middle, apical one at 4/5 of metat¬ ibial length; lateral face longitudinally convex, finely and very sparsely punctate and glabrous, basally with dense coarse punctures, along middle smooth; ventral edge fine¬ ly serrated, with three robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventral- ly on distal half, neither laterally nor dorsally carinate, dor- sally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina im¬ mediately beside it, glabrous; first metatarsomere slight¬ ly longer than two following tarsomeres combined and twice as long as very short dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 6V-X. Habitus: Fig. 291. Distribution. See map (Fig. 45D). Neoserica genieri sp. n. (Figs 7A-C, 29J, 45D) Type material examined. Holotype S “India: Mysore Shimoga dist., Agumbe Ghat, 2000’, VI. 1990 T.R.S. Nathan” (CMNC). Paratypes: 2 $$ “India: Mysore Shi¬ moga dist., Agumbe Ghat, 2000’, VI. 1990 T.R.S. Nathan” (CMNC, ZFMK), 4 SS, 4 $? Mndia, W Karnataka, Udupi distr., E of Bhatkal, Kollur, 26.-29.V.2006, Z. Ke- jval lgt.” (CPPB, ZFMK). Description. Length: 6.1 mm, length of elytra: 4.6 mm, width: 3.9 mm. Body oval, yellowish brown, ventral sur¬ face slightly darker, frons, disc of pronotum, scutellum and numerous small spots on elytra dark and with greenish shine; dorsal surface dull and glabrous, partly moderate¬ ly shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar¬ gin weakly sinuate medially, margins weakly reflexed; sur¬ face weakly convex, moderately shiny, finely and dense¬ ly punctate, with a few coarse punctures anteriorly bear¬ ing each a short erect seta; frontoclypeal suture distinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a sin¬ gle terminal seta. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Eyes moder¬ ately large; ratio of diameter/interocular width: 0.6. An¬ tenna with ten antennomeres; club composed of three an- tennomeres, straight, slightly shorter than remaining an¬ tennomeres combined, 7 th antennomere slightly transverse¬ ly produced. Mentum elevated and slightly flattened an¬ teriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins moderately and evenly convex and convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, coarsely and moderately densely punctate, also on midline. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at slightly blunt external apical angle of elytra, epi- pleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and moderately dense¬ ly punctate, without smooth midline, with a few longer setae at apex. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati¬ bial length, ratio width/length: 1/3.3; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and moderately densely punctate, along middle in apical half smooth, punctures with minute setae; ventral edge finely serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on dis¬ tal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina imme¬ diately beside it, glabrous; first metatarsomere slightly longer than two following tarsomeres combined, twice as Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 93 long as dorsal tibial spur. Protibia short, bidentate; ante¬ rior claws symmetrical, basal tooth of both claws blunt¬ ly truncate at apex. Aedeagus: Fig. 7A-C. Habitus: Fig. 29J. Diagnosis. Neoserica genieri sp. n. differs from the sub¬ sequent species, N. plateosa sp. n., by the dorsal process of phallobase being short and sharply pointed (instead of being long and rounded at apex). Etymology. The new species is dedicated to Franf ois Ge nier (Ottawa) who kindly sent us the material of the Canadian Museum of Nature for study (noun in genitive case). Variation. Length: 6.1-6.9 mm, length of elytra: 4.5^4.6 mm, width: 3.9—4.1 mm. Female: club composed of three antennomeres, straight, slightly shorter than remaining an- tennomeres combined, seventh antennomere not trans¬ versely produced; eyes as large as in male. Distribution. See map (Fig. 45D). Neoserica plateosa sp. n. (Figs 7D-F, 29K, 45D) Type material examined. Holotype S “S India; Karnata¬ ka; W Ghats 18 km E Shiradi; Gundia; 12 0 47’N 75°43’E; 200-500 m; P. Pacholatko leg. 16.-21.v.2002” (CPPB). Paratypes: 8 S3 “S India; Karnataka; W Ghats 18 km E Shiradi; Gundia; 12°47’N 75°43’E; 200-500 m; P. Pa¬ cholatko leg. 16.-21.v.2002” (CPPB). Description. Length: 7.2 mm, length of elytra: 4.5 mm, width: 4.0 mm. Body oval, yellowish brown, frons, ante¬ rior disc, two lateral long spots on pronotum and numer¬ ous small spots on elytra including the penultimate exter¬ nal interval dark and with greenish shine; dorsal surface dull and glabrous, frons moderately shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar¬ gin weakly sinuate medially, margins weakly reflexed; sur¬ face weakly convex, moderately shiny, finely and dense¬ ly punctate, with a few coarse punctures anteriorly bear¬ ing each a short erect seta; frontoclypeal suture distinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, densely finely punctate, with a sin¬ gle terminal seta. Frons dull, with sparse, fine punctures, with one or two single erect setae beside each eye. Eyes moderately large; ratio of diameter/interocular width: 0.57. Antenna with ten antennomeres; club composed of three antennomeres, straight, slightly shorter than remaining an¬ tennomeres combined, antennomere 7 slightly transverse¬ ly produced. Mentum elevated and slightly flattened an¬ teriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins evenly convex and moderately convergent towards sharp and slightly produced anterior angles. Anterior mar¬ gin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; posterior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lateral and later¬ al anterior glabrous. Hypomeron not carinate. Scutellum triangular, finely and densely punctate, on midline narrow¬ ly impunctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at slightly blunt external apical angle of elytra, epi- pleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.62. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and sparsely punctate, without smooth midline, with a few longer setae along api¬ cal margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.6; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and very sparse¬ ly punctate, along middle smooth, glabrous; ventral edge finely serrated, with three robust equidistant setae, medi¬ al face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on distal half, neither laterally nor dorsally car¬ inate, dorsally smooth; metatarsomeres with a strongly ser¬ rated ridge ventrally and a parallel subventral smooth Ca¬ rina immediately beside it, glabrous; first metatarsomere slightly longer than two following tarsomeres combined, nearly twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 94 Dirk Ahrens & Silvia Fabrizi Aedeagus: Fig. 7D-F. Habitus: Fig. 29K. Female un¬ known. Diagnosis. Neoserica plateosa sp. n. differs from N. Jlavoviridis and N. dichroa by the presence of a dorsal process at the apex of its phallobase externally of the left paramere; from N. dichroa it differs also by the antennal club being composed of three antennomeres only. Etymology. The species’ name (adjective in the nomina¬ tive singular) is derived from the Latin word ‘ plateosus ’ (rich with spots), with reference to the dark ornamenta¬ tion of the dorsal surface. Variation. Length: 6.2-7.2 mm, length of elytra: 4.3-4.5 mm, width: 3.9-4.0 mm. Distribution. See map (Fig. 45D). Neoserica plebea sp. n. (Figs 7G-I, 29L, 45D) Type material examined. Holotype: S “S India; Karnata¬ ka; W Ghats 20 km W Talguppa; Jog Falls; 14°14’N 74°44’E; 500±200m; P. Pacholatko; 22.-28.V.2002” (CPPB). Paratypes: 5 SS, 6 ?? “S India; Karnataka; W Ghats 20 km W Talguppa; Jog Falls; 14°14’N 74°44’E; 500±200m; P. Pacholatko; 22.-28.V.2002” (CPPB), 1 S “S India; Karnataka; W Ghats 18 km E Shiradi; Gundia; 12°47’N 75°43’E; 200-500 m; P. Pacholatko leg. 16.- 21.V.2002” (CPPB). Description. Length: 6.6 mm, length of elytra: 4.5 mm, width: 3.9 mm. Body oval, yellowish brown, ventral sur¬ face slightly darker, frons, disc of pronotum except two yellow spots beside middle, scutellum, elytral striae and numerous small spots on elytra dark and with greenish shine; dorsal surface dull and glabrous, partly moderate¬ ly shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar¬ gin weakly sinuate medially, margins weakly reflexed; sur¬ face weakly convex, moderately shiny, finely and dense¬ ly punctate, with a few coarse punctures anteriorly bear¬ ing each a short erect seta; frontoclypeal suture distinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a sin¬ gle terminal seta. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Eyes moder¬ ately large; ratio of diameter/interocular width: 0.54. An¬ tenna with ten antennomeres; club composed of three an¬ tennomeres, straight, slightly shorter than remaining an¬ tennomeres combined, antennomere 7 slightly transverse¬ ly produced. Mentum elevated and slightly flattened an¬ teriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins moderately and evenly convex and convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, coarsely and moderately densely punctate, also on midline. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at moderately rounded external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.88. Abdominal stemites finely and dense¬ ly punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and moderately densely punctate, without smooth midline, with a few longer setae at apex. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.4; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and moderately densely punctate, along middle in apical half smooth, punctures with minute setae; ventral edge finely serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on dis¬ tal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina imme¬ diately beside it, glabrous; first metatarsomere slightly longer than two following tarsomeres combined, nearly twice as long as dorsal tibial spur. Protibia short, biden- tate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 95 Aedeagus: Fig. 7G-I. Habitus: Fig. 29L. Diagnosis. Neosericaplebea sp. n. differs from N. genieri sp. n. in the left apophysis of the phallobase being very short. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the Latin adjective ‘ plebeus ’ (simple/ordinary). Variation. Length: 6.6-7.6 mm, length of elytra: 4.5-5.0 mm, width: 3.9^1.4 mm. Yellow ground colour may be sometimes darker that ornaments are less distinct. Female: club composed of three antennomeres, straight, distinct¬ ly shorter than the remaining antennomeres combined, an- tennomere 7 not transversely produced; eyes as large as in male. Distribution. See map (Fig. 45D). Neoserica rotundotibialis sp. n. (Figs 7J-L, 29M, 45D) Type material examined. Holotype S “S India; Kerala; Thekkady; Periyar Lake; 9,34N 77,10E 900-1000m; 19.- 27.iv.1997 Dembicky & Pacholatko leg./ IS 88/ 624 Sericini Asia spec.” (CPPB). Paratypes: 33 SS, 30 $$ “S India; Kerala; Thekkady; Periyar Lake; 9,34N 77,10E 900-1000m; 19.-27.iv.1997 Dembicky & Pacholatko leg./ IS 88/ 624 Sericini Asia spec.” (CPPB, ZFMK). Description. Length: 5.8 mm, length of elytra: 3.9 mm, width: 3.5 mm. Body oval, dark brown, ventral surface, legs, labroclypeus, sides and two lateral spots of prono- tum, including wide parts of elytra yellowish brown, ely¬ tra with numerous small dark spots, dark parts with green¬ ish shine; dorsal surface dull and glabrous, partly moder¬ ately shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar¬ gin moderately sinuate medially, margins weakly reflexed; surface weakly convex, moderately shiny, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect seta; frontoclypeal suture dis¬ tinctly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a single terminal seta. Frons dull, with sparse, fine punc¬ tures, with 2-3 single erect setae beside each eye. Eyes moderately large; ratio of diameter/interocular width: 0.57. Antenna with ten antennomeres; club composed of four antennomeres, straight, distinctly longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest shortly before base, lateral margins moderately and evenly convex and con¬ vergent towards sharp and slightly produced anterior an¬ gles, slightly narrowed towards base. Anterior margin of pronotum slightly convex, with robust and complete mar¬ ginal line; basal marginal line absent; posterior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lateral and later¬ al anterior margins sparsely setose. Hypomeron not car¬ inate. Scutellum triangular, coarsely and moderately densely punctate, midline widely impunctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at moderately rounded external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex at apex and dull, coarsely and moderate¬ ly densely punctate, without smooth midline, with a few longer setae at apex. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.3; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and moderately densely punctate, along middle in apical half smooth, punctures with minute setae; ventral edge finely serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on dis¬ tal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina imme¬ diately beside it, glabrous; first metatarsomere distinctly longer than the following two tarsomeres combined and distinctly more than twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 96 Dirk Ahrens & Silvia Fabrizi Aedeagus: Fig. 7J-L. Habitus: Fig. 29M. Diagnosis. Neoserica rotundotibialis sp. n. differs from the similar N. dichroa sp. n. by the insertion of the left paramere being displaced at its right side. Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived from the combined Latin words, ‘ rotundus ’ (round) and ‘ tibialis ’ (tibia), a character related with the tibia (with reference to the lon¬ gitudinally strongly convex dorsal margin of the metati¬ bia). Variation. Length: 5.2-5.8 mm, length of elytra: 3.8-3.9 mm, width: 3.1-3.5 mm. Female: club composed of three antennomeres, straight, distinctly shorter than the remain¬ ing antennomeres combined, seventh antennomere not transversely produced; eyes as large as in male. Distribution. See map (Fig. 45D). Neoserica submaculosa sp. n. (Figs 7M-0, 29N, 45D) Type material examined. Holotype <$ “India mer. Ker¬ ala Periyar, Senft lgt. April 1993/ 620 Sericini Asia spec.” (ZFMK). Paratypes: 1 S “India mer. Kerala Periyar, Sen¬ ft lgt. April 1993/ 620 Sericini Asia spec.” (ZFMK), 1 S, 2 $$ “S India Kerala: 1250 m; 15 km SW Munnar; L- 9.V.1997 10,02N 76,5E; Kallar Valley; Dembicky & Pa- cholatko leg./ IS 71” (CPPB, ZFMK). Description. Length: 6.1 mm, length of elytra: 4.2 mm, width: 3.5 mm. Body oval, dark brown, ventral surface slightly lighter, labroclypeus, sides and two lateral spots of pronotum, including wide parts of elytra yellowish brown, elytra with numerous small dark spots, dark parts with greenish shine; dorsal surface dull and glabrous, part¬ ly moderately shiny. Labroclypeus shiny, subtrapezoidal, wider than long, widest at base, lateral margins convex and strongly con¬ vergent to strongly rounded anterior angles, anterior mar¬ gin moderately sinuate medially, margins weakly reflexed; surface weakly convex, moderately shiny, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect seta; frontoclypeal suture dis¬ tinctly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus very short and triangular, sparsely finely punctate, with a single terminal seta. Frons dull, with sparse, fine punc¬ tures, with 2-3 single erect setae beside each eye. Eyes moderately large; ratio of diameter/interocular width: 0.6. Antenna with ten antennomeres; club composed of four antennomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum moderately wide, widest at base, lateral mar¬ gins moderately and evenly convex and convergent to¬ wards sharp and slightly produced anterior angles. Ante¬ rior margin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; pos¬ terior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lat¬ eral and lateral anterior margins sparsely setose. Hy- pomeron not carinate. Scutellum triangular, coarsely and moderately densely punctate, midline impunctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and densely punctate, intervals strong¬ ly convex, with fine and dense punctures exclusive along striae, at centre impunctate, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, end¬ ing at moderately rounded external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.52. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium strongly convex and dull, coarsely and moderately dense¬ ly punctate, without smooth midline, with a few longer setae at apex. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.0; dorsal margin dis¬ tinctly longitudinally convex, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally convex, finely and moderately densely punctate, along middle in apical half smooth, punctures with minute setae; ventral edge finely serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally on dis¬ tal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina imme¬ diately beside it, glabrous; first metatarsomere twice as long as dorsal tibial spur, subsequent tarsomeres lacking in holotype. Protibia short, bidentate; anterior claws sym¬ metrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 7M-0. Habitus: Fig. 29N. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 97 Diagnosis. Neoserica submaculosa sp. n. differs from the similar N. ammattiensis sp. n. by the long and straight left apophysis of the phallobase. Etymology. The name (adjective in the nominative sin¬ gular) of the new sepcies is derived from the combined Latin words, 'sub- (prefix) (almost), and ‘ maculosa ’ (with spots), with reference to the dorsal surface covered with dark spots. Variation. Length: 5.8-6.2 mm, length of elytra: 4.2^1.4 mm, width: 3.5 mm. Female: club composed of three an- tennomeres, straight, distinctly shorter than the remain¬ ing antennomeres combined, seventh antennomere slight¬ ly transversely produced; eyes as large as in male. Distribution. See map (Fig. 45D). Neoserica lubrica group Key to species of the Neoserica lubrica group of the In¬ dian subcontinent (SS) 1 Apex of protibia with a distal twin-tooth.2 1 ’ Apex of protibia simple, with a single tooth.3 2 Left paramere simply pointed. ..N. shillongensis sp. n. T Left paramere bifurcate, with two distal teeth. . N. kaskiensis Ahrens 3 Body stout and bicoloured (yellowish brown and black). N. bicolorea sp. n. 3’ Body more elongate, unicoloured yellowish.4 4 Parameres short, distinctly much less than half of phal- lobasal length. N. pubiforceps Ahrens 4’ Parameres long, subequal or longer than half of phal- lobasal length.5 5 Left paramere not bifurcate at apex but strongly curved. N. incompta Ahrens & Fabrizi 5’ Left paramere bifurcate at apex and nearly straight. . . N. bhalukpongensis sp. n. Neoserica bhalukpongensis sp. n. (Figs 4N-P, 28C, 45E) Type material examined. Holotype: S “NE India, Assam, Bhalukpong, 26.v.-3.vi.2006,27°02’N 92°35’E, 150m, P. Pacholatko leg.” (CPPB). Paratype: 1 $ “NE India, As¬ sam-Arunachal border, Bhalukpong, 150m, 27°00 , 48”N 92°39’08 ,, E, L. Dembicky leg., 1-8.V.2012” (ZFMK). Description. Length: 6.9 mm, length of elytra: 4.5 mm, width: 3.7 mm. Body oval, yellowish brown, dorsal sur¬ face strongly shiny, dorsal surface glabrous. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, mod¬ erately convergent anteriorly, anterior angles weakly rounded, anterior margin shallowly but widely sinuate me¬ dially, margins moderately reflexed; surface convex and shiny, finely and moderately densely punctate, glabrous except a few single seta anteriorly; frontoclypeal suture distinctly incised, flat and weakly curved; smooth area an¬ terior to eye small, weakly convex, approximately 1.5 times as wide as long; ocular canthus very short and broad (1/6 of ocular diameter), finely sparsely punctate, with one terminal seta._Frons with fine, moderately dense punctures, with a few long erect setae beside eyes. Eyes large, ratio diameter/interocular width: 0.75. Antenna with ten anten¬ nomeres, club with five antennomeres and straight, as long as the remaining antennomeres combined, antennomere six subequal to the length of club. Mentum elevated and slightly flattened anteriorly. Labrum not short, transverse¬ ly rectangular, not produced medially, without median sin- uation, with densely setose anterior margin. Pronotum weakly transverse, widest at the base, later¬ al margins moderately evenly convex and more strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin convexly produced medially, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, dense punctures, glabrous. Elytra oval, widest at posterior third, striae finely im¬ pressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures sometimes concentrated along striae, glabrous, with a few very long erect setae on penul¬ timate lateral interval; epipleural edge fine, ending at wide¬ ly rounded external apical angle of elytra, epipleura dense¬ ly setose, apical border with a fine fringe of microtri- chomes (visible at ca lOOx magnification). Ventral surface shiny, finely and densely punctate, metastemum glabrous; metacoxa glabrous, with a few sin¬ gle setae laterally; abdominal sternites finely and dense¬ ly punctuate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.52. Pygidium strongly con¬ vex and shiny, finely and very densely punctate, with a wide smooth midline, with a few long setae along apical margin. Legs short; femora shiny, with two rudimentary longi¬ tudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half and only weakly widened, poste¬ rior margin smooth dorsally. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.5, dorsally sharply carinate, with two groups of spines, basal group Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 98 Dirk Ahrens & Silvia Fabrizi shortly before middle, apical group at three quarters of metatibial length, basally with a few short single setae; lat¬ eral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tar- someres ventrally with sparse, short setae, smooth, nei¬ ther laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth, first metatar- somere distinctly shorter than the two following tar- someres combined and slightly longer than the upper tib- ial spur. Protibia moderately long, bidentate, distal tooth simply pointed at apex; anterior claws symmetrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4N-P. Habitus: Fig. 28C. Diagnosis. This new species is in its shape of the geni¬ talia and its external appearance very similar to N. kask- iensis Ahrens, 2004. N. bhalukpongensis sp. n. differs from the latter by the left paramere being narrowered towards the apex and having the internal distal tooth of the left paramere elongated and sharply pointed at the apex. Etymology. The new species is named with reference to its type locality, Bhalukpong (adjective in the nominative singular). Variation. Length: 6.9-7.2 mm, length of elytra: 4.5-4.9 mm, width: 3.7-4.0 mm. Female: Antennal club composed of three antennomeres, club slightly shorter than the re¬ maining antennomeres combined. Distribution. See map (Fig. 45E). Neoserica bicolorea sp. n. (Figs 4Q-T, 28D, 45E) Type material examined. Holotype: $ “NE India, Meghalaya, SW of Cherrapunjee, 25°-13‘-14‘ N, 91°40‘ E, 5.-24.V.2005, 900m, P. Pacholatko leg.” (CPPB). Paratypes: 64 SS, 63 “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E, 5.-24.V.2005, 900m, P. Pacholatko leg.” (CPPB, ZFMK), 2 2 ?? “India: Meghalaya state E Khasi Hills, 11km SW Cher¬ rapunjee, Laitkynsew, 21-24.iv.2008, 25° 13’N, 91° 39’E, 810m, Fikacek, Podalska, Sipek lgt ” (ZFMK). Description. Length: 6.2 mm, length of elytra: 4.2 mm, width: 3.7 mm. Body oval, yellowish brown, posterior half of elytra black, dorsal surface strongly shiny, dorsal sur¬ face glabrous. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins weakly convex, strongly convergent anteriorly, anterior angles moderately rounded, anterior margin straight medially, margins weakly reflexed; surface convex and shiny, fine¬ ly and densely punctate, glabrous, with a few single se¬ ta; frontoclypeal suture distinctly incised, flat and weak¬ ly curved; smooth area anterior to eye small, weakly con¬ vex, approximately 1.5 times as wide as long; ocular can- thus very short and broad (1/6 of ocular diameter), im- punctate, with one terminal seta. Frons with fine, very sparse punctures, with a few long erect setae beside eyes. Eyes small, ratio diameter/interocular width: 0.41. Anten¬ na with ten antennomeres, club with five antennomeres and straight, as long as the remaining antennomeres com¬ bined, antennomere six subequal to the length of club. Mentum elevated and slightly flattened anteriorly. Labrum transversely rectangular, not produced medially, without median sinuation, with densely setose anterior margin. Pronotum weakly transverse, widest at the base, later¬ al margins subparallel in basal half, moderately evenly convex and strongly convergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin convexly produced medially, with a fine complete marginal line; surface mod¬ erately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly car¬ inate basally, not produced ventrally. Scutellum narrow, triangular, with fine, dense punctures, glabrous. Elytra oval, widest at posterior third, striae finely im¬ pressed, finely and densely punctate, intervals weakly con¬ vex, with sparse, fine punctures concentrated along stri¬ ae, glabrous, only a few single long erect setae on later¬ al intervals; epipleural edge fine, ending at widely round¬ ed external apical angle of elytra, epipleura densely se¬ tose, apical border with a fine fringe of microtrichomes (visible at ca lOOx magnification). Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few sin¬ gle setae laterally; abdominal sternites finely and dense¬ ly punctuate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.63. Pygidium strongly con¬ vex and shiny, finely and densely punctate, without smooth midline, with a few long setae along apical margin. Legs short; femora shiny, with two rudimentary longi¬ tudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half and only weakly widened, poste¬ rior margin smooth dorsally. Metatibia wide and short, widest at apex, ratio of width/length: 1/2.5, dorsally sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metatibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex inte- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 99 riorly near tarsal articulation moderately sinuate. Tar- someres ventrally with sparse, short setae, smooth, nei¬ ther laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth, first metatar- somere little shorter than the two following tarsomeres combined and more than a third of its length longer than the upper tibial spur. Protibia moderately long, bidentate, distal tooth simply pointed at apex; anterior claws sym¬ metrical, basal tooth of inner claw sharply truncate at apex. Aedeagus: Fig. 4Q-T. Habitus: Figs 28D. Diagnosis. The new species is more stout and robust than all other species of the N. lubrica group. In the shape of its genitalia it is most similar to N. pubiforceps Ahrens, 2004. From the latter, N. bicolorea sp. n. differs by the bicoloured elytra, and the shape of the parameres: the sin- utation of the left paramere is situated at the distal mar¬ gins in N. bicolorea , instead of at the lateral margin (as in N. pubiforceps). Etymology. The new species is named (adjective in the nominative singular) with reference to its bicoloured ely¬ tra with the combined latin words, ‘ coloreus ’ (variegat¬ ed) and the prefix L bi- (twice). Variation. Length: 6.2-6.5 mm, length of elytra: 4.2^1.6 mm, width: 37-3.9 mm. Elytra bicoloured, completely light brown, or dark brown. Female: Antennal club com¬ posed of three antennomeres, club slightly shorter than the remaining antennomeres combined. Remarks. Paratypes are labelled as ‘ Neoserica bicolorata sp. n.’ which was the name originally chosen for this species. However, to avoid the name ‘bicolorata’ to be used twice within Sericini (see Gynaecoserica bicolorata Ahrens & Fabrizi, 2009) we decided to use and introduce the name Neoserica bicolorea sp. n. Distribution. See map (Fig. 45E). Neoserica kaskiensis Ahrens, 2004 (Figs 28E, 45E) Neoserica kaskiensis Ahrens, 2004b: 163. Material examined. See Ahrens 2004b (p. 163). Aedeagus: See Ahrens 2004b (figs 249-251, p. 419). Distribution. Central Nepal (Fig. 45E). Neoserica incompta Ahrens & Fabrizi, 2009 Neoserica incompta Ahrens & Fabrizi, 2009b: 265. Material examined. See Ahrens & Fabrizi 2009b (p. 265). Aedeagus: See Ahrens & Fabrizi 2009b (fig. 5M-0, p. 281). Distribution. NE India: Arunachal Pradesh (Fig. 45E). Neoserica pubiforceps Ahrens, 2004 (Figs 28F, 45E) Neoserica pubiforceps Ahrens, 2004b: 165. Material examined. See Ahrens 2004b (p. 165); 1 f “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-15‘N, 91°47‘E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB). Aedeagus: See Ahrens 2004b (figs. 252-254, p. 419). Distribution. Eastern Nepal, Sikkim, Assam and Megha¬ laya (NE India) (Fig. 45E). Neoserica shillongensis sp. n. (Figs 4U-W, 28G, 45E) Type material examined. Holotype: S “Shillong A.G.R. coll. 6-VI-18./ India T.B.Fletcher B.M. 1943-9./6'” (BMNII). Paratypes: 3 $$ “Shillong, Assam F.W.C./G.C. Champion Coll. B.M. 1927-409.” (BMNH, ZFMK). Description. Length: 5.1 mm, length of elytra: 3.8 mm, width: 3.1 mm. Body oval, yellowish brown, dorsal sur¬ face strongly shiny, dorsal surface glabrous. Labroclypeus subtrapezoidal, distinctly wider than long, widest at base, lateral margins straight, convergent anteriorly, anterior angles weakly rounded, anterior mar¬ gin distinctly sinuate medially, margins moderately re- flexed; surface nearly flat, shiny, finely and sparsely punc¬ tate, with a few setae anteriorly; frontoclypeal suture dis¬ tinctly incised, flat and weakly curved; smooth area an¬ terior to eye small, weakly convex, approximately 1.3 times as wide as long; ocular canthus very short and broad (1/6 of ocular diameter), finely sparsely punctate, with one terminal seta. Frons with fine, evenly and moderately dense punctures, with a few long erect setae beside eyes. Eyes very small, ratio diameter/interocular width: 0.39. Antennal club missing in holotype. Mentum elevated and slightly flattened anteriorly. Labrum not short, transverse¬ ly rectangular, weakly produced medially, without medi¬ an sinuation, with densely setose anterior margin. Pronotum weakly transverse, widest at the base, later¬ al margins in basal half nearly straight and weakly con¬ vergent, moderately evenly convex and more strongly con- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 100 Dirk Ahrens & Silvia Fabrizi vergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt and weakly rounded at tip; anterior margin convexly produced medially, with a fine complete marginal line; surface moderately densely and finely punctate, glabrous; lateral and anterior border sparsely setose; hypomeron distinctly carinate basally, not produced ventrally. Scutellum narrow, triangular, with fine, dense punctures, impunctate on basal midline, glabrous. Elytra oval, widest at posterior third, striae finely im¬ pressed, finely and densely punctate, intervals nearly flat, with sparse, fine punctures concentrated along striae, glabrous; epipleural edge fine, ending at widely rounded external apical angle of elytra, epipleura densely setose, apical border with a fine fringe of microtrichomes (visi¬ ble at ca lOOx magnification). Ventral surface shiny, finely and densely punctate, metasternum glabrous; metacoxa glabrous, with a few sin¬ gle setae laterally; abdominal sternites finely and dense¬ ly punctuate, with a transverse row of coarse punctures, each bearing a short robust seta. Mesosternum between mesocoxae as wide as the mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.5. Pygidium strongly con¬ vex and shiny, finely and very densely punctate, without smooth midline, with a few long setae along apical mar¬ gin. Legs short; femora shiny, with two rudimentary longi¬ tudinal rows of setae, superficially and sparsely punctate, glabrous; metafemur with anterior margin acute, without serrated line behind anterior edge, posterior margin smooth ventrally in apical half and only weakly widened, poste¬ rior margin smooth dorsally. Metatibia moderately wide and short, widest at apex, ratio of width/length: 1/2.7, dor- sally sharply carinate, with two groups of spines, basal group at one third, apical group at three quarters of metat- ibial length, basally with a few short single setae; lateral face weakly convex, finely and sparsely punctate, smooth along the middle; ventral edge finely serrated, with three robust nearly equidistant setae; medial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tar- someres ventrally with sparse, short setae, smooth, nei¬ ther laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally, smooth, first metatar- somere slightly shorter than following two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate, distal tooth wide and truncate at apex, slightly sinuate in distal point; anterior claws symmetrical, basal tooth of inner claw sharply trun¬ cate at apex. Aedeagus: Fig. 4U-W. Habitus: Fig. 28G. Diagnosis. The new species is in the shape of the protib¬ ia (apical twin-teeth) similar to N. kaskiensis , but the new species differs in the simple and sharply pointed apex of the left paramere (which is bifurcate in N. kaskiensis). Etymology. This new species is named with reference to its type locality, Shillong (adjective in the nominative sin¬ gular). Variation. Length: 5.1-5.8 mm, length of elytra: 3.8-3.9 mm, width: 3.1-3.4 mm. Female: Antennal club composed of three antennomeres, club slightly shorter than the re¬ maining antennomeres combined. Distribution. See map (Fig. 45E). Neoserica septemlamellata group Remarks. The group which is distributed mainly in south¬ ern China and northern Indochina occurs only with two species in the Himalayas. The group was recently com¬ pletely revised by Ahrens et al. (2014a). Key to species of the Neoserica septemlamellata group of the Indian subcontinent (SS) 1 Left paramere with an additional large external lobe. . N. crenatolineata Ahrens & Fabrizi F Left paramere without an additional large external lobe. N. changrae Ahrens Neoserica changrae Ahrens, 2004 (Figs 28H, 45B) Neoserica changrae Ahrens, 2004b: 167. Material examined. See Ahrens 2004b (p. 167). Aedeagus: See Ahrens 2004b (figs 261-265, p. 421). Distribution. Endemic to Bhutan (Fig. 45B). Neoserica crenatolineata Ahrens & Fabrizi, 2009 Neoserica crenatolineata Ahrens & Fabrizi, 2009b: 262. Material examined. See Ahrens & Fabrizi 2009b (p. 262). Aedeagus: See Ahrens & Fabrizi 2009b (fig. 5A-C, p. 281). Distribution. Endemic to NE India, W-Arunachal Pradesh (Fig. 45B). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 101 Neoserica speciosa group Key to species of the Neoserica speciosa group ($<$) 1 Distal lobe of right paramere strongly convex at its dor¬ sal margin and with numerous small teeth. . N. assamensis (Frey) 1 ’ Distal lobe of right paramere straight at its dorsal mar¬ gin and with a single larger tooth. . N. speciosa Brenske Neoserica assamensis (Frey, 1962) Leucoserica [sic] assamensis Frey, 1962b: 612; Sabatinel- li 1993: 626. Neoserica assamensis'. Ahrens 2004b: 155. Material examined. See Ahrens 2004b (p. 155); 1 S “NE India; Meghalaya, SW of Cherrapunjee, 25°13 , -15’N 91 o 40’E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB), 2 SS “NE India; Meghalaya, 2002 1 km E of Tura, 500-600m, 25°30’N, 90°14’E; 13.-18.V. M. Tryz- na & P. Benda lgt.” (CPPB), 3 “NE India; Megha¬ laya, 2002 3 km E of Tura, 1150m, 25°30’N, 90°14’E; 12.V. M. Tryzna & P. Benda lgt.” (CPPB), 1 ex. “India: Meghalaya state E Khasi Hills, 111cm SW Cherrapunjee, Laitkynsew, 21-24.iv.2008, 25° 13’N, 91° 39’E, 810m, Fikacek, Podalska, Sipek lgt.” (ZFMK). Aedeagus: See Ahrens 2004b (figs 229-231, p. 415). Distribution. NE India: Assam and Meghalaya (Fig. 45D). Neoserica speciosa Brenske, 1898 Neoserica speciosa Brenske, 1898: 353; Ahrens 2004b: 153. Material examined. See Ahrens 2004b (p. 153); 2 SS “NE India; Meghalaya, 2002 1 km E of Tura, 500-600m, 25°30’N, 90°14’E; 13.-18.V. M. Tryzna & P. Benda lgt.” (CPPB). Aedeagus: See Ahrens 2004b (figs 226-228, p. 415). Distribution. NE India: Assam and Meghalaya (Fig. 45D). Neoserica uniformis group Neoserica uniformis Moser, 1920 Neoserica uniformis Moser, 1920: 5; Ahrens 2004b: 161; Ahrens & Fabrizi 2009b: 270. Material examined. Ahrens 2004b (p. 161); Ahrens & Fa¬ brizi 2009b (p. 272); 2 ex. “NE India, Meghalaya, 1 km E of Tura, 500-600m, 25°30‘N, 90°14‘E, 2.-5.V.2002, M.Tryzna & Benda leg.”(CPPB), 1 S “NE India; Meg¬ halaya, 2002 3 km E of Tura, 1150m, 25°30 , N, 90°14’E; 6.-12.V. M. Tryzna & P. Benda lgt.” (CPPB), 1 S “NE In¬ dia Meghalaya, 1999, 9 km NW Jowai, 1400 m 25°30‘N 92°10‘E 12.V. Dembicky & Pacholatko leg.” (CPPB). Aedeagus: See Ahrens, 2004b (figs 246-248, p. 418). Distribution. India: Assam, Nagaland and Meghalaya (Fig. 45B). Neoserica variegata group Key to species of the Neoserica variegata group {$$) 1 Elytra with fine and thick white setae only. . N. variegata Moser 1 ’ Elytra with fine and scale-like white setae.2 2 Dorsal surface of body shiny. N. fusiforceps sp. n. 2’ Dorsal surface of pronotum and elytra dull.3 3 Head shiny. Antennal club distinctly longer than re¬ maining antennomeres combined. .N. flagrans sp. n. 3’ Frons dull. Antennal club as long as or shorter than re¬ maining antennomeres combined.4 4 Pronotum laterally with scale-like setae that are more than five times longer than wide. . N. sparsesquamata sp. n. 4’ Pronotum laterally with scale-like setae that only three times longer than wide.5 5 Right paramere hooked, its apex split into several well separated spines. N. incisa sp. n. 5’ Right paramere hooked, its apex with a single termi¬ nal, not split into several well separated spines. . N. unciforceps sp. n. Neoserica flagrans sp. n. (Figs 7P-R, 290, 45F) Type material examined. Holotype $ “Myanmar N (Bur¬ ma) 25 km E Putao, H-800 m Nan San Bon vill. 06- 09.05.1998 leg. S. Murzin& V. Sinaev/708 Sericini Asia spec.” (ZFMK). Description. Length: 5.5 mm, length of elytra: 3.9 mm, width: 3.3 mm. Body oval, yellowish brown, ventral sur¬ face, frons, two pairs of large spots on disc of pronotum, scutellum, lateral intervals and extended dots on elytra dark; dorsal surface except head dull, nearly glabrous, ex¬ cept scale-like setae on elytra. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to strongly rounded anterior angles, anterior margin distinct- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 102 Dirk Ahrens & Silvia Fabrizi ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, finely punctate, with a single very short terminal seta. Frons shiny, with dense, fine punctures, with a few fine setae.beside eyes. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.65. Antenna with ten anten- nomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral mar¬ gins evenly convex and strongly convergent towards sharp and moderately produced anterior angles, slightly nar¬ rowed basally. Anterior margin of pronotum slightly con¬ vex, with robust and complete marginal line; basal mar¬ ginal line absent; posterior angles blunt, strongly round¬ ed at tip; surface finely and densely punctate, with minute setae on disc, lateral and lateral anterior margins sparse¬ ly setose. Hypomeron carinate, carina not produced ven- trally. Scutellum triangular, coarsely and densely punctate, with minute setae. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals mod¬ erately convex, with fine and dense punctures concentrat¬ ed along striae, with shorter and longer, white scale-like setae; epipleural edge robust, ending at moderately rounded external apical angle of elytra, epipleura dense¬ ly setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.36. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dull, strongly convex, coarsely and moderately densely punctate, with a narrow smooth mid- line and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin serrated dorsally, with a few short setae basally. Metatibia moder¬ ately long and slender, widest at half of metatibial length, ratio width/length: 1/3.6; dorsal margin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally con¬ vex, finely and sparsely punctate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally impunctate; metatar- someres with a strongly serrated ridge ventrally and a par¬ allel subventral smooth carina immediately beside it, glabrous, in addition with a blunt lateral carina; first metatarsomere distinctly longer than following two tar¬ someres combined and distinctly more than twice as long as dorsal tibial spur. Protibia short, bidentate, before basal tooth with a blunt lateral extension; anterior claws asym¬ metrical, basal tooth of interior claw widened and trun¬ cate at apex. Aedeagus: Fig. 7P-R. Habitus: Fig. 290. Diagnosis. Neoserica flagrans sp. n. differs from the rather similar N. variegata Moser by the dull body surface and the large scale-like setae on its elytra. From N. fusiforceps sp. n. it differs by the well-separated parameres. In its gen¬ ital morphology N. flagrans sp. n. is very similar to N. sparsa (Arrow, 1946) from Myanmar, however, the me¬ dian lobe of the left parameres is in the latter species strongly widened at the apex. Etymology. The name of the new species is derived from the Latin word ‘ flagrans ’ (burning) (adjective in the nom¬ inative singular). Distribution. See map (Fig. 45F). Neoserica fusiforceps sp. n. (Figs 7S-U, 29P, 45F) Type material examined. Holotype S “NE India; Assam; 5 km N of Umrongso; 700 m; 25°27’N 92°43’E; 21 .v. 1999 Dembicky & Pacholatko leg./ 614 Sericini Asia spec.” (CPPB). Description. Length: 5.8 mm, length of elytra: 4.3 mm, width: 3.6 mm. Body oval, yellowish brown, ventral sur¬ face, frons, disc of pronotum, scutellum, lateral intervals and extended dots on elytra dark; dorsal surface moder¬ ately shiny and glabrous. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to strongly rounded anterior angles, anterior margin distinct¬ ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, finely punctate, with a single very short terminal seta. Frons shiny, with dense, fine punctures, with a few fine setae beside eyes. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.56. Antenna with ten anten- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 103 nomeres; club composed of four antennomeres, straight, as long as remaining antennomeres combined. Mentum el¬ evated and slightly flattened anteriorly. Pronotum wide, widest shortly before base, lateral mar¬ gins evenly convex and strongly convergent towards sharp and moderately produced anterior angles, slightly nar¬ rowed basally. Anterior margin of pronotum slightly con¬ vex, with robust and complete marginal line; basal mar¬ ginal line absent; posterior angles blunt, strongly round¬ ed at tip; surface finely and densely punctate, with minute setae on disc and short white setae on sides, lateral and lateral anterior margins sparsely setose. Hypomeron car¬ inate, carina not produced ventrally. Scutellum triangular, coarsely and very densely punctate, with minute setae, midline at base widely impunctate. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals mod¬ erately convex, with fine and dense punctures concentrat¬ ed along striae, with short white scale-like setae in punc¬ tures, otherwise glabrous; epipleural edge robust, ending at moderately rounded external apical angle of elytra, epi- pleura densely setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.44. Abdominal stemites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dull, at centre yellow, strongly convex at apex, coarsely and moderately densely punc¬ tate, with a narrow smooth midline and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin serrated dorsally, with a few short setae basally. Metatibia moder¬ ately long and slender, widest at half of metatibial length, ratio width/length: 1/3.3; dorsal margin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally con¬ vex, finely and sparsely punctate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally very finely and sparse¬ ly punctate; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina imme¬ diately beside it, glabrous, in addition with a blunt later¬ al carina; first metatarsomere distinctly longer than fol¬ lowing two tarsomeres combined and distinctly more than twice as long as dorsal tibial spur. Protibia short, biden- tate, before basal tooth with a blunt lateral extension; an¬ terior claws asymmetrical, basal tooth of interior claw shortly lobiform. Aedeagus: Fig. 7S-U. Habitus: Fig. 29P. Diagnosis. Neoserica fusiforceps sp. n. differs from the rather similar N. variegata Moser by the less shiny body surface and the scale-like setae on its elytra, as well as in the shape of the parameres being completely fused with one another in the new species. Etymology. The name of the new species (noun in appo¬ sition) is derived from the combined Latin words ‘ fusils’ (fused) and ‘ forceps’ (forceps), with reference to the para¬ meres being fused with the aedeagus. Distribution. See map (Fig. 45F). Neoserica incisa sp. n. (Figs 7V-X, 30A, 45F) Type material examined. Holotype S “NE India Megha¬ laya state West Garo Hills, Nokrek Nat. Park 9-17. V. 1996 alt. 1100+150 m GPS N25°29.6’ E90 o 19.5 , (WGS 84) E. Jendek & O. Sausa/IS 8/ 606 Sericini Asia spec” (CPPB). Paratypes: 13 SS, 2 ? ? “NE India Meghalaya state West Garo Hills, Nokrek Nat. Park 9-17.V. 1996 alt. 1100+150 m GPS N25°29.6’ E90 o 19.5 , (WGS 84) E. Jendek & O. Sausa” (CPPB, ZFMK), 6 (ft?, 12 “NE India; Assam; 5 km N of Umrongso; 700 m; 25°27’N 92°43’E; 21.V.1999 Dembicky & Pacholatko leg./ 614 Sericini Asia spec.” (CPPB), 1 $ “NE India; Meghalaya; 1400 m; Nokrek n.p. 3km S Daribokgiri 25°27’N 90°19’E; 26.iv. 1999 Dembicky & Pacholatko leg.” (CPPB). Description. Length: 5.8 mm, length of elytra: 4.0 mm, width: 3.5 mm. Body oval, yellowish brown, abdomen in¬ cluding pygidium, frons, disc of pronotum, scutellum, lat¬ eral margin and smaller dots on elytra dark; dorsal sur¬ face except head dull, nearly glabrous, except scale-like setae on elytra. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to strongly rounded anterior angles, anterior margin distinct¬ ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, finely punctate, with a single very short terminal seta. Frons dull in posterior half, with dense, fine punc¬ tures, with a few fine setae beside eyes. Eyes moderate¬ ly large; ratio of diameter/interocular width: 0.58. Anten- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 104 Dirk Ahrens & Silvia Fabrizi na with ten antennomeres; club composed of four anten¬ nomeres, straight, slightly longer than remaining anten¬ nomeres combined. Mentum elevated and slightly flat¬ tened anteriorly. Pronotum wide, widest at base, lateral margins evenly convex and strongly convergent towards sharp and mod¬ erately produced anterior angles. Anterior margin of pronotum slightly convex, with robust and complete mar¬ ginal line; basal marginal line absent; posterior angles strongly rounded; surface finely and densely punctate, with minute setae on disc, beside disc with short, fine, scale-like, white setae, lateral and lateral anterior margins sparsely setose. Hypomeron carinate, carina not produced ventrally. Scutellum triangular, coarsely and densely punc¬ tate, with minute setae, base impunctate medially. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals mod¬ erately convex, with fine and dense punctures concentrat¬ ed along striae, with numerous smaller and larger, white scale-like setae; epipleural edge robust, ending at moder¬ ately rounded external apical angle of elytra, epipleura densely setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.55. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dull, strongly convex, coarsely and moderately densely punctate, with a narrow smooth mid¬ line and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened but distinctly serrate in apical half, pos¬ terior margin serrated dorsally, with a few short setae basally. Metatibia moderately long and slender, widest at half of metatibial length, ratio width/length: 1/3.4; dorsal margin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lat¬ eral face longitudinally convex, finely and sparsely punc¬ tate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally impunctate; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it, glabrous, in addition with a blunt lateral carina; first metatarsomere distinctly longer than following two tarsomeres combined and dis¬ tinctly more than twice as long as dorsal tibial spur. Protib¬ ia short, bidentate, before basal tooth with a blunt lateral extension; anterior claws asymmetrical, basal tooth of in¬ terior claw strongly widened, truncate at apex. Aedeagus: Fig. 7V-X. Habitus: Fig. 30A. Diagnosis. Neoserica incisa sp. n. differs from the simi¬ lar N. sparsesquamata by the broad scales on its prono¬ tum and the right paramere being hooked at the apex, its apex splitting into several well-separated spines. Etymology. The name of the new species (adjective in the nominative singular) is derived from the Latin word c in- cisus’ (incised), with a reference to the morphology of the aedeagus. Variation. Length: 5.1-6.1 mm, length of elytra: 3.6-4.1 mm, width: 3.1-3.5 mm. Dark dots on dorsal surface may vary strongly in density and extension. Female: Antennal club composed of three antennomeres, slightly shorter than the remaining antennomeres combined; eyes as large as in male. Distribution. See map (Fig. 45F). Neoserica sparsesquamata sp. n. (Figs 8A-C, 30B, 45F) Type material examined. Holotype S “Kaziranga 75 m, 7.-9.5.1976/Assam W. Wittmer C. Baroni U. 1976/ 615 Sericini Asia spec.” (NHMB). Description. Length: 6.2 mm, length of elytra: 4.3 mm, width: 3.8 mm. Body oval, yellowish brown, abdomen ex¬ cept pygidium, frons, disc of pronotum, scutellum, later¬ al margins and smaller dots on elytra dark; dorsal surface except head dull, nearly glabrous, except scale-like setae on elytra. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to strongly rounded anterior angles, anterior margin distinct¬ ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, finely punctate, with a single very short terminal seta. Frons dull in posterior half, with dense, fine punc¬ tures, with a few fine setae beside eyes. Eyes moderate¬ ly large; ratio of diameter/interocular width: 0.63. Anten¬ na with ten antennomeres; club composed of four anten¬ nomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 105 Pronotum wide, widest at base, lateral margins evenly convex and strongly convergent towards sharp and mod¬ erately produced anterior angles. Anterior margin of pronotum slightly convex, with robust and complete mar¬ ginal line; basal marginal line absent; posterior angles blunt, strongly rounded at tip; surface finely and densely punctate, with minute setae on disc, beside disc with short, fine, scale-like, white setae, lateral and lateral anterior mar¬ gins sparsely setose. Hypomeron carinate, carina not pro¬ duced ventrally. Scutellum triangular, coarsely and dense¬ ly punctate, with minute setae. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals mod¬ erately convex, with fine and dense punctures concentrat¬ ed along striae, with numerous smaller and larger, white scale-like setae; epipleural edge robust, ending at moder¬ ately rounded external apical angle of elytra, epipleura densely setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.55. Abdominal stemites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dull, strongly convex, coarsely and moderately densely punctate, with a narrow smooth mid¬ line and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened but finely serrate in apical half, posteri¬ or margin serrated dorsally, with a few short setae basal- ly. Metatibia moderately long and slender, widest at half of metatibial length, ratio width/length: 1/3.2; dorsal mar¬ gin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lat¬ eral face longitudinally convex, finely and sparsely punc¬ tate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally impunctate; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it, glabrous, in addition with a blunt lateral carina; first metatarsomere distinctly longer than following two tarsomeres combined and dis¬ tinctly more than twice as long as dorsal tibial spur. Protib¬ ia short, bidentate, before basal tooth with a blunt lateral extension; anterior claws asymmetrical, basal tooth of in¬ terior claw small, lobiform. Aedeagus: Fig. 8A-C. Habitus: Fig. 30B. Diagnosis. Neoserica sparsesquamata sp. n. differs from all other species of the N. variegata group by its dull body surface including the frons, the scale-like setae on its ely¬ tra, and the pronotum having laterally scale-like setae that are more than five times longer than wide. Etymology. The name of the new species (adjective in the nominative singular) is derived from the combined Latin words L sparsus\ (sparse) and ‘ squamatus ’ (bearing scales), with reference to the presence of sparse scales on the elytra. Distribution. See map (Fig. 45F). Neoserica unciforceps sp. n. (Figs 8D-F, 30C, 45F) Type material examined. Holotype S “NE India Megha¬ laya state W Garo Hills, Balphakram Nat. Park 22- 27.V. 1996 alt. 400+150 m GPS N25°1L E90 o 5L (WGS 84) E. Jendek & O. Sausa/ IS 30/ 609 Sericini Asia spec” (CPPB). Paratype. 1 $ “NE India Meghalaya state W Garo Hills, Balphakram Nat. Park 22-27.V.1996 alt. 400+150 m GPS N25°l 1 5 E90°51 ’ (WGS 84) E. Jendek & O. Sausa/ IS 30” (ZFMK). Description. Length: 5.8 mm, length of elytra: 3.9 mm, width: 3.2 mm. Body oval, yellowish brown, metastemum, abdomen including pygidium, frons, disc of pronotum, scutellum, lateral margin and smaller dots on elytra dark; dorsal surface except head dull, nearly glabrous, except scale-like setae on elytra. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to strongly rounded anterior angles, anterior margin distinct¬ ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, finely punctate, with a single very short terminal seta. Frons dull in posterior half, with dense, fine punc¬ tures, with a few fine setae beside eyes. Eyes moderate¬ ly large; ratio of diameter/interocular width: 0.65. Anten¬ na with ten antennomeres; club composed of four anten- nomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum wide, widest at base, lateral margins evenly convex and strongly convergent towards sharp and mod¬ erately produced anterior angles. Anterior margin of pronotum slightly convex, with robust and complete mar¬ ginal line; basal marginal line absent; posterior angles Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 106 Dirk Ahrens & Silvia Fabrizi strongly rounded; surface finely and densely punctate, with minute setae on disc, beside disc with short, fine, scale-like, white setae, lateral and lateral anterior margins sparsely setose. Hypomeron carinate, carina not produced ventrally. Scutellum triangular, coarsely and densely punc¬ tate, with minute setae, base impunctate medially. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals mod¬ erately convex, with fine and dense punctures concentrat¬ ed along striae, with numerous smaller and larger, white scale-like setae; epipleural edge robust, ending at moder¬ ately rounded external apical angle of elytra, epipleura densely setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dull, strongly convex, coarsely and moderately densely punctate, with a narrow smooth mid¬ line and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened but distinctly serrate in apical half, pos¬ terior margin serrated dorsally, with a few short setae basally. Metatibia moderately long and slender, widest at half of metatibial length, ratio width/length: 1/3.6; dorsal margin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lat¬ eral face longitudinally convex, finely and sparsely punc¬ tate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally impunctate; metatarsomeres with a strongly serrated ridge ventrally and a parallel subventral smooth carina immediately beside it, glabrous, in addition with a blunt lateral carina; first metatarsomere distinctly longer than following two tarsomeres combined and dis¬ tinctly more than twice as long as dorsal tibial spur. Protib¬ ia short, bidentate, before basal tooth with a blunt lateral extension; anterior claws asymmetrical, basal tooth of in¬ terior claw small, lobifonn. Aedeagus: Fig. 8D-F. Habitus: Fig. 30C. Diagnosis. Neoserica unciforceps sp. n. differs from the similar N. incisa by the slightly shorter antennal club (male), and the apical hook of the right paramere which is is at the apex not split into several well-separated spines but has a single terminal. Etymology. The name of the new species is derived from the combined Latin words 'uncus' (hook) and ‘ forceps' (forceps), with reference to the morphology of the right paramere (noun in apposition). Variation. Length: 5.2-5.8 mm, length of elytra: 3.6-3.9 mm, width: 3.0-3.2 mm. Female: Antennal club composed of three antennomeres, slightly shorter than the remain¬ ing antennomeres combined; pygidium less convex; pro- tarsal claws symmetric. Distribution. See Map (Fig. 45F). Neoserica variegata Moser, 1915 (Figs 8G-I, 30D, 45F) Neoserica variegata Moser, 1915c: 380. Type material examined. Syntypes: 1 “Naga Hills/ maculosa Brsk./ boops Waterh. afine/ coll. Brenske/ Neoserica variegata Type Mos.” (ZMHB), 1 S “Naga Hills/ maculosa Brsk./ coll. Brenske” (ZMHB), 1 $ “Na¬ ga Hills/ maculosa Brsk./ boops Waterh. afine/ coll. Brenske/ Neoserica variegata Type Mos.” (ZMHB). Additional material examined. 6 ex. “Shillong Assam F. W. C./ G. C. Champion Coll. B. M. 1927-409” (BMNH), 32 ex. “NE India Meghalaya state Jaintia Hills reg. Jowai 6-8.VI.1996 alt. 1350+100 m GPS N25°27‘ E 92°12‘ (WGS 84) E. Jendek & O. Sausa leg.” (CPPB), 6 ex. “Mawphlang 15.5. 1850 m/Meghalaya 1976Wittmer, Baroni U.” (NHMB), 2 ex. “Umtyngar 16.5. Cherrapun- jee/ Meghalaya 1976 Wittmer, Baroni FT.” (NHMB), 2 ex. “India 26.VI.95 Cherrapunjee Meghalaya Werner leg.” (ZFMK), 1 ex. “NE India: Meghalaya East Khasi Hills strada Shillong-Cherrapunji ca. 20 km S Shillong 26.VI.1995/ L. Bartolozzi & K. Werner legit” (MZF), 1 ex. “NE India Meghalaya East Khasi Hills (ca. 1500 m) Mawphlang 25.VI.1995/ L. Bartolozzi & K. Werner le¬ git” (MZF), 1 ex. “Nilghedi Hills” (ZMHB), 2 1 f “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-15‘N, 91°47‘E, 500-900m, L. Dembicky leg., 11.-12.V.2004” (CPPB), 2 $$, 2 §1? “NE India Meghalaya state Jaintia Hills reg., Jowai 6-8.VI.1996 alt. 1350+100 m GPS N25°27’ E92°12’ (WGS 84) E. Jendek & O. Sausa/ IS 29/ 602 Sericini Asia spec.” (CPPB), 3 SS, 2 “NE India Meghalaya, 1999, 9 km NW Jowai, 1400 m 25°30‘N 92°10‘E 12.V. Dembicky & Pacholatko leg.” (CPPB). Redescription. Length: 6.1 mm, length of elytra: 4.1 mm, width: 3.5 mm. Body oval, yellowish brown, ventral sur¬ face, frons, disc of pronotum and small dots on elytra dark; dorsal surface shiny and glabrous. Labroclypeus subrectangular, wider than long, widest at base, lateral margins convex and weakly convergent to Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 107 strongly rounded anterior angles, anterior margin distinct¬ ly sinuate medially, margins moderately reflexed; surface weakly convex, finely and densely punctate, with a few coarse punctures anteriorly bearing each a short erect se¬ ta; frontoclypeal suture finely incised, slightly elevated and weakly curved medially; smooth area anterior to eye 1.5 times as wide as long; ocular canthus moderately long and narrow, sparsely finely punctate, with a single terminal se¬ ta. Frons shiny, with dense, fine punctures, with a few fine setae beside eyes. Eyes moderately large; ratio of diam¬ eter/interocular width: 0.62. Antenna with ten anten- nomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins in basal half straight and moderately convergent, in anterior half moderately convex and strongly convergent towards sharp and moderately produced anterior angles. Anterior mar¬ gin of pronotum slightly convex, with robust and complete marginal line; basal marginal line absent; posterior angles blunt, slightly rounded at tip; surface finely and densely punctate, except minute setae glabrous, lateral and later¬ al anterior margins sparsely setose. Hypomeron carinate, carina not produced ventrally. Scutellum triangular, coarsely and densely punctate, midline at base widely im- punctate. Elytra oblong, widest at posterior third, striae distinct¬ ly impressed, finely and densely punctate, intervals dis¬ tinctly convex, with fine and dense punctures concentrat¬ ed along striae, most intervals completely impunctate me¬ dially, with short white setae in punctures, otherwise glabrous; epipleural edge robust, ending at moderately rounded external apical angle of elytra, epipleura dense¬ ly setose, apical border membraneous, with a broad fringe of microtrichomes. Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.36. Abdominal stemites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium dark and dull, strongly convex at apex, coarsely and moderately densely punctate, with a narrow smooth midline and with moderately dense long setae. Legs slender; femora finely and sparsely punctate; metafemur shiny, anterior margin acute, without submar¬ ginal serrated line; anterior row of seta-bearing punctures complete; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin serrated dorsally, with a few short setae basally. Metatibia moder¬ ately long and slender, widest at half of metatibial length, ratio width/length: 1/3.4; dorsal margin sharply carinate, with two groups of spines, basal group middle, apical one at 4/5 of metatibial length; lateral face longitudinally con¬ vex, finely and sparsely punctate, punctures with minute setae; ventral edge serrate, with two very widely separate robust setae only; medial face smooth, apex interiorly near tarsal articulation sharply truncated. Tarsomeres with fine, very dense setae ventrally, dorsally impunctate; metatar- someres with a strongly serrated ridge ventrally and a par¬ allel subventral smooth carina immediately beside it, glabrous, in addition with a blunt lateral carina; first metatarsomere distinctly longer than following two tar¬ someres combined and distinctly more than twice as long as dorsal tibial spur. Protibia short, bidentate, before basal tooth with a blunt lateral extension; anterior claws asym¬ metrical, basal tooth of interior claw shortly lobiform. Aedeagus: Fig. 8G-I. Habitus: Fig. 30D. Distribution. See map (Fig. 45F). Neoserica incertae sedis Neoserica agumbeensis sp. n. (Figs 8J-L, 30E, 46A) Type material examined. Holotype 8 “India Mysore Shi- moga Dist. Agumbe Ghat 2000 ft. V-74 T.R.S. Nathan/ 621 Sericini Asia spec” (MHNG). Paratypes: 2 88 “S India, Karnataka, Coorg distr. 10 km NE Viraipet 75°46’E, 12°06’N, 500-800 m Z. Kejval & M. Tryzna leg.” (CPPB, ZFMK), 1 8, 2 PP “India: Mysore Shimoga dist., Agumbe Ghat, 2000’, V.1991 T.R.S. Nathan” (CMNC, ZFMK), 1 S, 3f§$ “India: Mysore Shimoga Dist., Agumbe Ghat, 600 m, V.1987 T.R.S. Nathan” (CMNC, ZFMK), 58 88, 11 9$ “S India; Karnataka; WGhats 20 km W Talguppa; Jog Falls; 14°14’N 74°44’E; 500±200m; P. Pacholatko; 22.-28.V.2002” (CPPB), 3 88, 2 $$ “S India; Karnataka; W Ghats 18 km E Shiradi; Gundia; 12°47’N 75°43’E; 200-500 m; P. Pacholatko leg. 16.- 21.V.2002” (CPPB, ZFMK), 1 8 “India, Karnataka, 20 km SE Sagar, 600m, 14°06,37’N 75°08,93’E, M. Halada leg., 12.V.2005” (CPPB), 2 88 “Maissour Shimoga Juin 1897/ Museum Paris ex. Coll. R. Oberthur” (MNHN). Additional material examined. 1 8 [head and thorax lacking] “S India; Karnataka; W Ghats 20 km W Talgup¬ pa; Jog Falls; 14°14’N 74 0 44’E; 500±200m; P. Pa¬ cholatko; 22.-28.V.2002” (CPPB). Description. Length: 6.4 mm, length of elytra: 4.0 mm, width: 4.2 mm. Body oval, black, antenna dark brown, labroclypeus shiny, dorsal surface dull, except some pi- losity on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, later¬ al margins convex and distinctly convergent anteriorly, an¬ terior angles broadly rounded, anterior margin weakly sin¬ uate medially, all margins strongly reflexed; lateral mar- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 108 Dirk Ahrens & Silvia Fabrizi gin and ocular canthus produce an indistinct angle; sur¬ face convexly elevated anteriorly, finely and densely punc¬ tate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area anterior to eye nearly flat, twice as wide as long; ocular canthus short and wide (one third of ocular diameter), coarsely and densely punctate, with a ter¬ minal seta. Frons dull, with fine, dense punctures and a very few erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio diameter/ interocular width: 0.44. Antenna with ten antennomeres; club in male with five antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins evenly moderately convex and convergent an¬ teriorly, anterior angles moderately produced and sharp, posterior angles blunt; anterior margin straight, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, punctures with very minute setae, otherwise glabrous, close to anterior angle with a long seta on each side of disc; lateral and anterior margin densely setose; hypomeron carinate, slightly pro¬ duced ventrally. Scutellum wide, triangular, at apex mod¬ erately pointed, with fine, evenly dense punctures and minute setae. Elytra widest at middle, striae distinctly impressed, fine¬ ly and densely punctate, intervals convex, with fine and dense punctures concentrated along striae, except very minute setae in punctures only a few short, white setae on odd intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra chitinous, only with an ul¬ tra-fine rim of sparse microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metepisternum impunctate posteriorly, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally that are condensed in a short transverse line; abdominal ster- nites finely and densely punctate, the two basal sternites with dense setae, each sternite with a transverse row of very distant (distance equal sternite length) but coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/2.1. Pygidi- um strongly convex and dull, coarsely and densely punc¬ tate, without smooth midline, glabrous except a few sparse longer setae. Legs wide and short; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser behind posterior lon¬ gitudinal row of setae, anterior margin acute, without ser¬ rated line behind anterior edge, anterior row of setae re¬ duced, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at middle, ratio of width/length: 1/2.1, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face nearly flat, coarsely but sparsely punctate, glabrous, smooth along middle; ventral margin finely serrate, with three robust equidistant setae; medial face smooth and glabrous; apex finely serrate, weakly concavely sinuate interiorly near tarsal articula¬ tion. Tarsomeres dorsally smooth and glabrous, neither lat¬ erally nor dorsally carinate, densely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth sub ventral longitudinal carina, glabrous; first metatarsomere slightly shorter than following two tar¬ someres combined and as long as dorsal tibial spur. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8J-L. Habitus: Fig. 30E. Diagnosis. Neoserica agumbeensis sp. n. differs from the, in external characters similar, species N. nathani Frey in the antennal club being composed of five antennomeres (instead of four), as well as in the longer parameres. Etymology. The new species is named according to the type locality, Agumbe Ghat (adjective in the nominative singular). Variation. Length: 5.8-7.8 mm, length of elytra: 3.6-4.5 mm, width: 3.7-4.8 mm. Colour strongly variable, entire¬ ly dark blackish, black with elytra reddish brown, or en¬ tire dorsal surface except black head reddish brown. The antennal club is in some populations (Jog Falls) longer with the fifth antennomere strongly transversely produced and being half as long as the club, which is then 1.2 times as long as remaining antennomeres combined. Female: club with five antennomeres and straight, as long as the remaining antennomeres combined. Distribution. See map (Fig. 46A). Neoserica disciplineensis sp. n. (Figs 24U-W, 391, 46B) Type material examined. Holotype S “India, Tamil Nadu D: Vilupparam, Auroville, Discipline vill. 01. VII.- 30.IX.2014 leg. local collector 12° 0,7’N, 79° 47.97’E” (NME). Paratype: 1 $ “India, Tamil Nadu D: Vilupparam, Auroville, Discipline vill. 01.VII.-30.IX.2014 leg. local collector 12° 0,7’N, 79° 47.97 , E” (ZFMK). Description. Length: 8.1 mm, length of elytra: 5.3 mm, width: 4.9 mm. Body oval, dark brown, antenna yellow- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 109 ish brown, labroclypeus shiny, dorsal surface dull, except dense pilosity on head dorsal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and strongly convergent anterior¬ ly, anterior angles including anterior margin broadly rounded, not sinuate medially, all margins strongly re¬ flexed, lateral margin and ocular canthus produce a very indistinct angle; surface flat, finely and densely punctate, with numerous coarse punctures each bearing an erect se¬ ta; frontoclypeal suture finely incised, not elevated and weakly angled medially; smooth area anterior to eye wide, nearly flat, three times as wide as long; ocular canthus long and wide (one third of ocular diameter), coarsely and densely punctate, with a single terminal seta. Frons dull, with fine, moderately dense punctures and a few erect se¬ tae behind the frontoclypeal suture. Eyes moderately large, ratio diameter/ interocular width: 0.55. Antenna with ten antennomeres; club with five antennomeres and straight, as long as remaining antennomeres combined. Mentum el¬ evated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins in basal half almost straight and slightly convergent anteri¬ orly, weakly convex in anterior half and evenly conver¬ gent anteriorly, anterior angles distinctly produced and acute, posterior angles blunt; anterior margin almost straight, with fine and complete marginal line, base with¬ out marginal line; surface densely and finely punctate, punctures with very minute setae, otherwise glabrous; lat¬ eral and anterior margin densely setose; hypomeron car¬ inate, slightly produced ventrally. Scutellum wide, trian¬ gular, at apex moderately pointed, with fine, evenly dense punctures and minute setae. Elytra widest just behind middle, striae finely impressed, finely and moderately densely punctate, intervals weak¬ ly convex, with fine and evenly dense punctures, except very minute setae in punctures only a few short setae on lateral odd intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra with a fine rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metepisternum impunctate posteriorly, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally; ab¬ dominal sternites finely and densely punctate, the two basal sternites with dense setae, each sternite with a trans¬ verse row of coarse punctures each bearing a robust seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.69. Pygidium moderately convex at apex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few longer setae along apical margin. Legs wide and moderately long; femora with two lon¬ gitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser be¬ hind the posterior longitudinal row of setae, anterior mar¬ gin acute, without serrated line behind anterior edge, pos¬ terior margin smooth and strongly widened at apex ven¬ trally, finely serrate over its entire length dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at middle, ratio of width/length: 1/2.5, sharply carinate dorsally, with two groups of spines, basal group at first third, apical group at three quarters of metat- ibial length, with a few short and robust single spines basally; lateral face weakly longitudinally convex, finely and sparsely punctate, glabrous, smooth along the mid¬ dle; ventral margin finely serrate, with four robust equi¬ distant setae; medial face smooth and glabrous; apex fine¬ ly serrate, weakly concavely sinuate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, robustly densely se¬ tose ventrally; metatarsomeres with a strongly serrated ridge and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tar¬ someres combined and as long as dorsal tibial spur. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 24U-W. Habitus: Fig. 391. Diagnosis. Neoserica disciplineensis sp. n. differs from the very similar N. sexfoliata Moser by the antennal club being composed of only five antennomeres and in the much shorter and wider right and left paramere, as well as in the wider ventral medial sclerotised lobe that is pres¬ ent only in these two species. Etymology. The new species is named according to its type locality, the village Discipline (adjective in the nom¬ inative singular). Variation. Length: 7.8-8.1 mm, length of elytra: 5.1-5.3 mm, width: 4.9 mm. Female: Eyes slightly smaller, ratio diameter/interocular width: 0.53, antennal club with five antennomeres and straight, as long as remaining anten¬ nomeres combined, first joint of club as long as the club. Distribution. See map (Fig. 46B). Neoserica garlangensis Ahrens, 2004 Neoserica garlangensis Ahrens, 2004b: 157. Material examined. See Ahrens 2004b (p. 157). Aedeagus: See Ahrens 2004b (figs 236-238, p. 416). Distribution. Endemic to the central Nepal Himalaya (Fig. 46A). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 110 Dirk Ahrens & Silvia Fabrizi Neoserica inops Ahrens & Fabrizi 2009 Neoserica inops Ahrens & Fabrizi, 2009b: 263. Material examined. See Ahrens & Fabrizi 2009b (p. 263). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 5G-I, p. 281). Distribution. NE India, W-Arunachal Pradesh (Fig. 46B). Neoserica inspergata Ahrens & Fabrizi 2009 Neoserica inspergata Ahrens & Fabrizi, 2009b: 262. Material examined. See Ahrens & Fabrizi 2009b (p. 262). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 5D-F, p. 281). Distribution. NE India, W-Arunachal Pradesh (Fig. 46A). Neoserica keralana sp. n. (Figs 8M-0, 3OF, 46B) Type material examined. Holotype S “India mer. Ker¬ ala Peryiar, Senft leg. April 1993/ 629 Sericini Asia spec.” (ZFMK). Description. Length: 6.0 mm, length of elytra: 4.2 mm, width: 3.4 mm. Body oval, dark brown, antenna yellow¬ ish brown, labroclypeus shiny, dorsal surface dull, except some pilosity on head dorsal surface nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins straight and moderately convergent anteriorly, anterior angles broadly rounded, an¬ terior margin weakly sinuate medially, all margins strong¬ ly reflexed; lateral margin and ocular canthus produce a distinct angle; surface convexly elevated anteriorly, fine¬ ly and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely in¬ cised, weakly angled medially; smooth area anterior to eye nearly flat, twice as wide as long; ocular canthus long and wide (one third of ocular diameter), coarsely and dense¬ ly punctate, without terminal seta. Frons dull, with fine, dense punctures and a very few erect setae beside eyes. Eyes large, ratio diameter/interocular width: 0.81. Anten¬ na with ten antennomeres; club in male with six anten- nomeres and straight, 1.3 times as long as remaining an¬ tennomeres combined. Mentum elevated and slightly flat¬ tened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins in basal half almost straight and slightly con¬ vergent anteriorly, weakly convex in anterior half and evenly convergent anteriorly, anterior angles weakly pro¬ duced and right-angled, posterior angles blunt; anterior margin almost straight, with fine and complete marginal line, base without marginal line; surface densely and fine¬ ly punctate, punctures with very minute setae, otherwise glabrous; lateral and anterior margin densely setose; hy- pomeron carinate, slightly produced ventrally. Scutellum wide, triangular, at apex moderately pointed, with fine, evenly dense punctures and minute setae. Elytra widest in posterior third, striae finely impressed, finely and moderately densely punctate, intervals weak¬ ly convex, with fine and evenly dense punctures, except very minute setae in punctures only a few short setae on lateral odd intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra with a fine rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metepisternum impunctate posteriorly, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally; ab¬ dominal sternites finely and densely punctate, the two basal sternites with dense setae, each stemite with a trans¬ verse row of coarse punctures each bearing a robust seta. Mesostemum between mesocoxae 1.5 times as wide as mesofemur..Ratio of length of metepistemum/metacoxa: 1/1.81. Pygidium strongly convex at apex and dull, coarse¬ ly and densely punctate, without smooth midline, glabrous except a few longer setae along apical margin. Legs wide and moderately long; femora with two lon¬ gitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser be¬ hind the posterior longitudinal row of setae, anterior mar¬ gin acute, without serrated line behind anterior edge, pos¬ terior margin smooth and strongly widened at apex ven¬ trally, finely serrate over its entire length dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at middle, ratio of width/length: 1/2.6, sharply carinate dorsally, with two groups of spines, basal group behind middle first, apical group at 4/5 of metati- bial length, with a few short and robust single spines basal¬ ly, beside dorsal margin with a serrated continuous line ending at first group of spines; lateral face nearly flat, coarsely but sparsely punctate, glabrous, smooth along middle; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, weakly concavely sinuate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, densely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal Ca¬ rina, glabrous; first metatarsomere slightly shorter than fol¬ lowing two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8M-0. Habitus: Fig. 30F. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 111 Diagnosis. N. keralana sp. n. differs from the very simi¬ lar N. multijlabellata Moser by the left paramere being wider and the ventral phallobasal process being shorter. Etymology. The new species is named according to its oc¬ currence in Kerala state (adjective in the nominative sin¬ gular). Distribution. See map (Fig. 46A). Neoserica lenangensis Ahrens & Fabrizi, 2009 Neoserica lenangensis Ahrens & Fabrizi, 2009b: 260. Material examined. See Ahrens & Fabrizi 2009b (p. 260). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 4M-0, p. 280). Distribution. NE India, W-Arunachal Pradesh (Fig. 46A). Neoserica madurana Moser, 1915 (Figs 8P-R, 46B) Neoserica madurana Moser, 1915a: 167. Type material examined. Holotype “India Madura/ Neoserica madurana Type Mos.” (ZMHB). Remarks. The only known syntype preserved in the col¬ lection of Moser (ZMHB) is after (presumable) dermestid damage completely destroyed, only the aedeagus is pre¬ served. There was no additional material available to des¬ ignate a neotype. Therefore, it was not possible to provide a redescription of the species and to include it in the iden¬ tification key. The original description translated from German (Moser 1915a: 167): “Dull, dorsal surface dark reddish brown, ventral sur¬ face light brown. Head sparsely setose, frons with fine punctures. [LabrojClypeus rugosely punctate, slightly con¬ vex at middle, only slightly anteriorly narrowed, anterior margin elevated, indistinctly sinuate medially. Antenna red-yellowish, with ten antennomeres, club in male with four antennomeres, shorter than the remaining anten¬ nomeres combined, 5^ antennomere with spines. Prono- tum with fine, minutely setose and moderately dense punc¬ tures, anterior margin weakly convex, setose as lateral margins. Lateral margins behind middle slightly concave, anteriorly convexly convergent. Sharp anterior angles pro¬ duced, blunt posterior angles weakly rounded at tip. Ely¬ tra with irregularly punctate striae, intervals weakly con¬ vex nearly impunctate. Pygidium densely punctate. Metasternal plate with a finely incised longitudinal line and sparse robust setae. Metacoxa sparsely rugose, with some setae laterally. Abdominal stemite with a transverse row of setae. Metafemur and metatibia little widened. Metafemur dull, with an anterior and a posterior [longi¬ tudinal] row of setae. Metatibia shorted, [lateral] face fine¬ ly sparsely punctate.” Aedeagus: Fig. 8P-R. Distribution. See map (Fig. 46B). Neoserica matura Ahrens, 2004 Neoserica matura Ahrens, 2004b: 160; Ahrens & Fabrizi 2011: 162. Material examined. Ahrens 2004b (p. 160); Ahrens & Fabrizi 2011 (p. 162); 19 ex. “NE India, Meghalaya, 1 km E of Tura, 500 - 600m, 25°30‘N, 90°14‘E, 2.-5.V.2002, M. Tryzna & Benda leg.”(CPPB), 2 $$ “NE India Meghalaya state West Garo Hills, Nokrek Nat. Park 9- 17.V.1996 alt. 1100+150 m GPS N25°29.6’ E90°19.5’ (WGS 84) E. Jendek & O. Sausa” (CPPB). Aedeagus: See Ahrens 2004b (figs 243-245, p. 417). Distribution. Nepal and Meghalaya (India) (Fig. 46A). Neoserica multiflabellata Moser, 1916 (Figs 8S-U, 30G, 46A) Neoserica multiflabellata Moser, 1916: 155. Type material examined. Syntypes: 1 $ “Trichinopoli Ind. or./ Neoserica multiflabellata Type $ Mos.” (ZMHB), 2 SS “Trichinopoli Ind. or.” (ZMHB), 1 S “Pondycher- ry India or.” (ZMHB), 1 S “Pondycherry India or./ Neoserica multiflabellata Type f Moser” (ZMHB), 1 <$, 1 ^ “Madura India or.” (ZMHB), 1 $ “India Madura” (ZMHB). Additional material examined. 4 ex. “Pulney Hills Kodeikanal 6500 ft. IV.53/ S. Indien leg. Nathan” (CF, ZFMK), 1 ex. “Coimbatore Slid Indien 5.54 leg. Nathan” (CF), 1 ex. “Indes Orientales Mts. Kodeicanel J. Castets” (MNHN), 1 ex. “Indes Orient. Trichinopoly R.P.J. Castets” (MNHN), 1 ex. “S-India: 19.VII.96 5 km E Pe- rumalmtal, Tamil Nadu, Palni Hills, Werner/ Lorenz leg.” (ZFMK). Redescription. Length: 5.6 mm, length of elytra: 4.4 mm, width: 3.5 mm. Body oval, reddish brown, antenna yel¬ lowish brown, labroclypeus shiny, dorsal surface dull, ex¬ cept some pilosity on head dorsal surface nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and mod¬ erately convergent anteriorly, anterior angles broadly Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 112 Dirk Ahrens & Silvia Fabrizi rounded, anterior margin weakly sinuate medially, all mar¬ gins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface convexly elevated an¬ teriorly, finely and very densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area anterior to eye nearly flat, 1.5 times as wide as long; oc¬ ular canthus short and wide (one third of ocular diame¬ ter), impunctate, with a fine terminal seta. Frons dull, with fine, dense punctures and a very few erect setae beside eyes. Eyes large, ratio diameter/interocular width: 0.63. Antenna with ten antennomeres; club in male with six an- tennomeres and straight, as long as remaining anten¬ nomeres combined. Mentum elevated and slightly flat¬ tened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins in basal half almost straight and subparallel, weakly convex in anterior half and evenly convergent an¬ teriorly, anterior angles moderately produced and sharp, posterior angles blunt; anterior margin almost straight, with fine and complete marginal line, base without mar¬ ginal line; surface densely and finely punctate, punctures with very minute setae, otherwise glabrous; lateral and an¬ terior margin densely setose; hypomeron carinate, slight¬ ly produced ventrally. Scutellum wide, triangular, at apex moderately pointed, with fine, moderately dense punctures and minute setae. Elytra widest in posterior third, striae finely impressed, finely and moderately densely punctate, intervals weak¬ ly convex, with fine and dense punctures concentrated along striae, except very minute setae in punctures only a few short setae on penultimate lateral intervals; epipleur- al edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra with a fine rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal stemites finely and densely punctate, each sternite with a transverse row of coarse punctures each bearing a robust seta. Mesosternum between meso- coxae 1.5 times as wide as mesofemur..Ratio of length of metepisternum/metacoxa: 1/1.62. Pygidium strongly con¬ vex at apex and dull, finely and moderately densely punc¬ tate, without smooth midline, glabrous except a few longer setae along apical margin. Legs wide and moderately long; femora with two lon¬ gitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser be¬ hind the posterior longitudinal row of setae, anterior mar¬ gin acute, without serrated line behind anterior edge, pos¬ terior margin smooth and strongly widened at apex ven¬ trally, not serrate dorsally, without setae basally. Metati¬ bia wide and flattened, short, widest at middle, ratio of width/length: 1/2.9, sharply carinate dorsally, with two groups of spines, basal group behind middle, apical group at 4/5 of metatibial length, with a few short and robust sin¬ gle spines basally, beside dorsal margin with a serrated continuous line ending at first group of spines; lateral face nearly flat, finely and sparsely punctate, glabrous, smooth along middle; ventral margin finely serrate, with three ro¬ bust equidistant setae; medial face smooth and glabrous; apex finely serrate, weakly concavely sinuate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, densely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal Ca¬ rina, glabrous; first metatarsomere slightly shorter than fol¬ lowing two tarsomeres combined and as long as dorsal tib- ial spur. Protibia short, bidentate; anterior claws symmet¬ rical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8S-U. Habitus: Fig. 30G. Distribution. See map (Fig. 46A). Neoserica nathani Frey, 1972 (Figs 8V-Y, 3OH, 46A) Neoserica nathani Frey, 1972: 190. Autos erica nathani. Patel et al. 1982: 40. Type material examined. Holotype: S “Kerala V.1970 Calicut distr. 3500 ft. Nathan/ Type/ Neoserica nathani det. G. Frey 1970 n. sp.” (CF). Paratypes: 2 SS, 5 “Ker¬ ala V. 1970 Calicut distr. 3500 ft. Nathan/ Paratype Neoser¬ ica nathani det. G. Frey 1970 n. sp” (CF). Additional material examined. 13 ex. “India, Kerala Trivandrum Dt. Poonmudi Range, 3000 ft. IV.-V.71. leg. T. R. S. Nathan” (CF, ZFMK), 2 ex. “Slid Indien lg. Nathan/ Nilgiri Hills Moyar Camp 3000 ft. V.1954/ Neoserica G. Frey det. 1954 5flabellata Br.” (CF), 5 ex. “India Mysore V.1973 Coorg distr., Mercara 4000 ft. T. R. S. Nathan leg.” (MHNG, CPPB), 3 ex. “Sud India Ana- malai Hills, Cinchona V-72 leg. T. R. S. Nathan” (CGST), 1 ex. “Mysore S. Indien/ Byaran Kuppe 800 m 4.53” (CF), 2 ex. “India (S): Kerala State: Calicut Distr.: Chembra Peak Area, 1067 m V.1970/ T. R. S. Nathan Coll. Bishop Museum” (BPBM), 2 ex. “Nilgiri Hills. H.L. Andrewes/ Nilgiri Hills/ Adrewes Bequest. B.M. 1922-221” (BMNH), 1 ex. “Nilgiri Hills. A.K.W. Downing. B.M. 1923-324” (BMNH), 1 ex. ($) “South India Kerala St. Quilon Distr., Thenmala V.1993, leg. Theresa Rajabai Sel¬ va Nathan” (CARL), 1 ex. ($) “South India Mysore St. Shimoga Distr., Agumbe Ghal, 2000 ft. V. 1990, leg. T. Ra- jabai Selva Nathan” (CARL), 3 ex. “South India Nilgiri Hills, Devala 3200 ft. V.1984, leg. Theresa Rajabai Sel¬ va Nathan” (CARL, ZFMK), 2 ex. “India: Kerala Quilon Dist., Thenmala, VI. 1988 T.R.S. Nathan” (CMNC), 4 ex. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 113 “S-India, Kerala state, Ponmudi hill resort, 30 km NE of Trivandrum, 77°06‘E 8°46‘N, ca. 1300-1500 m, 7.- 13.V.1999, Z. Kejval & M. Tryzna leg./ 645 Sericini Asia spec.” (CPPB). Redescription. Length: 6.9 mm, length of elytra: 4.4 mm, width: 4.4 mm. Body oval, black, antenna dark brown, labroclypeus shiny, dorsal surface dull, except some pi- losity on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins convex and distinctly convergent anteriorly, anterior angles broadly rounded, anterior margin straight, all margins strongly reflexed; lateral margin and ocular canthus produce an indistinct angle; surface convexly el¬ evated, very finely and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons dull, with fine, dense punctures and a very few erect setae beside eyes and behind frontoclypeal suture. Eyes small, ratio di¬ ameter/interocular width: 0.73. Antenna with ten anten- nomeres; club in male with five antennomeres and straight, as long as remaining antennomeres combined. Mentum el¬ evated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins evenly moderately convex and convergent an¬ teriorly, anterior angles moderately produced and sharp, posterior angles blunt; anterior margin straight, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, punctures with very minute setae, otherwise glabrous, close to anterior angle with a long seta on each side of disc; lateral and anterior margin densely setose; hypomeron carinate, slightly pro¬ duced ventrally. Scutellum wide, triangular, at apex mod¬ erately pointed, with fine, evenly dense punctures and minute setae. Elytra widest at middle, striae distinctly impressed, fine¬ ly and densely punctate, intervals convex, with fine and dense punctures concentrated along striae, except very minute setae in punctures only a few short, white setae on odd intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra chitinous, only with an ul¬ tra-fine rim of sparse microtrichomes (visible at ca 1 OOx magnification). Ventral surface dull, coarsely and densely punctate, metepisternum impunctate posteriorly, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally that are condensed in a short transverse line; abdominal ster- nites finely and densely punctate, the two basal sternites with dense setae, each sternite with a transverse row of coarse punctures each bearing a robust long seta, these punctures are very distant (distance equal stemite length) on penultimate sternite. Mesostemum between mesocox- ae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few sparse longer se¬ tae. Legs wide and short; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser behind posterior lon¬ gitudinal row of setae, anterior margin acute, without ser¬ rated line behind anterior edge, anterior row of setae re¬ duced, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at middle, ratio of width/length: 1/2.6, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face nearly flat, finely and sparse¬ ly punctate, glabrous, smooth along middle; ventral mar¬ gin finely serrate, with three robust equidistant setae; me¬ dial face smooth and glabrous; apex finely serrate, weak¬ ly concavely sinuate interiorly near tarsal articulation. Tar- someres dorsally smooth and glabrous, neither laterally nor dorsally carinate, densely setose ventrally; metatar- someres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere slightly shorter than following two tar- someres combined and as long as dorsal tibial spur. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 8V-Y. Habitus: Fig. 3OH. Distribution. See map (Fig. 46A). Neoserica periyarensis sp. n. (Figs 9A-C, 301, 46B) Type material examined. Holotype $ “S India; Kerala; Thekkady; Periyar Lake; 9,34N 77,10E 900-1000m; 19.- 27.iv. 1997 Dembicky & Pacholatko leg./ IS 88/ 622 Sericini Asia spec.” (CPPB). Paratypes: 2 SS “S India; Kerala; Thekkady; Periyar Lake; 9,34N 77,10E 900- 1000m; 19.-27.iv.1997 Dembicky & Pacholatko leg./ IS 88/ 624 Sericini Asia spec.” (CPPB, ZFMK). Description. Length: 7.2 mm, length of elytra: 4.6 mm, width: 5.0 mm. Body oval, black, antenna dark brown, labroclypeus shiny, dorsal surface dull, except some pi- losity on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins convex and distinctly convergent anteriorly, anterior angles broadly rounded, anterior margin straight, all margins strongly reflexed; lateral margin and ocular Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 114 Dirk Ahrens & Silvia Fabrizi canthus produce an indistinct angle; surface convexly el¬ evated, finely and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area an¬ terior to eye nearly flat, twice as wide as long; ocular can¬ thus short and wide (one third of ocular diameter), fine¬ ly and sparsely punctate, with a terminal seta. Frons dull, with fine, dense punctures and a very few erect setae be¬ side eyes and behind frontoclypeal suture. Eyes small, ra¬ tio diameter/interocular width: 0.47. Antenna with ten an- tennomeres; club in male with five antennomeres and straight, slightly longer than remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum moderately transverse, widest at base, later¬ al margins evenly moderately convex and convergent an¬ teriorly, anterior angles moderately produced and sharp, posterior angles blunt; anterior margin straight, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, punctures with very minute setae, otherwise glabrous; lateral and anterior mar¬ gin densely setose; hypomeron carinate, slightly produced ventrally. Scutellum wide, triangular, at apex moderately pointed, with fine, dense punctures and minute setae. Elytra widest at middle, striae distinctly impressed, fine¬ ly and densely punctate, intervals convex, with fine and dense punctures concentrated along striae, except very minute setae in punctures only a few short, white setae on odd intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra chitinous, only with an ul¬ tra-fine rim of sparse microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metepisternum impunctate posteriorly, metasternum sparsely covered with fine, short or very minute setae; metacoxa glabrous, with a few single setae laterally that are condensed in a short transverse line; abdominal ster- nites finely and densely punctate, the two basal sternites with dense setae, each sternite with a transverse row of very distant (distance equal sternite length) but coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur..Ra¬ tio of length of metepistemum/metacoxa: 1/2.1. Pygidi- um moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few sparse longer setae. Legs wide and short; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, punctures finer and slightly denser behind posterior lon¬ gitudinal row of setae, anterior margin acute, without ser¬ rated line behind anterior edge, anterior row of setae re¬ duced, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia wide and flattened, short, widest at middle, ratio of width/length: 1/2.2, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face nearly flat, coarsely but sparsely punctate, glabrous, smooth along middle; ventral margin finely serrate, with three robust equidistant setae; medial face smooth and glabrous; apex finely serrate, weakly concavely sinuate interiorly near tarsal articula¬ tion. Tarsomeres dorsally smooth and glabrous, neither lat¬ erally nor dorsally carinate, densely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere as long as following two tarsomeres com¬ bined and as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9A-C. Habitus: Fig. 301. Female un¬ known. Diagnosis. Neoserica periyarensis sp. n. differs from the externally similar N. nathani Frey by the distinctly longer and less curved parameres, the slightly longer antennal club; it differs from N. agumbeensis sp. n. by having the parameres of equal length. Etymology. The new species is named according its type locality, Periyar Lake (adjective in the nominative singu¬ lar). Variation. Length: 6.8-7.2 mm, length of elytra: 4.3-4.6 mm, width: 4.8-5.0 mm. Distribution. See map (Fig. 46B). Neoserica probsti Ahrens, 2004 Neoserica probsti Ahrens, 2004b: 159. Material examined. See Ahrens 2004b (p. 159). Aedeagus: See Ahrens 2004b (figs 239-242, p. 417). Distribution. Eastern Nepal and northern Laos (Fig. 46A). Neoserica pseudomajor sp. n. (Figs 9D-F, 30J, 46B) Type material examined. Holotype: S “NE India; Meghalaya; 1999 3km E of Tura; 500-1150m; 25°30’N 90°14’E; 1.-8.V. J. Rolcik” (CPPB). Paratype: 1 S, 1 f “NE India, Meghalaya, 2002, 3 km E Tura; 1150m; 25°30’N 90°14’E; M. Tryzna & P. Benda” (ZFMK). Description. Length: 8.8 mm, length of elytra: 6.2 mm, width: 5.3 mm. Body oval, dark brown, antenna yellow- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 115 ish brown, ventral surface reddish brown, dorsal surface dull, nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins convex and convergent anteriorly, anterior an¬ gles strongly rounded, anterior margin shallowly sinuate, margins weakly reflexed; lateral margin and ocular can- thus produce a blunt angle; surface nearly flat, finely and very densely punctate, glabrous; frontoclypeal suture in¬ distinctly incised, weakly angled medially; smooth area anterior to eye weakly convex, twice as wide as long; oc¬ ular canthus short and moderately wide (ca. one third of ocular diameter), finely and densely punctate, with a ter¬ minal seta. Frons with fine, dense punctures and a few short setae beside eyes. Eyes moderately large, ratio di¬ ameter/interocular width: 0.52. Antenna with ten anten- nomeres; club with four antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Men- tum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins nearly straight and weakly convergent, in an¬ terior half convex and distinctly convergent anteriorly, an¬ terior angles distinctly produced and sharp, posterior an¬ gles blunt; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; sur¬ face densely and finely punctate, glabrous; lateral and an¬ terior margin sparsely setose; hypomeron carinate, slight¬ ly produced ventrally. Scutellum wide, triangular, with fine, dense punctures, with minute setae in punctures. Elytra widest in posterior third, striae distinctly im¬ pressed, finely and densely punctate, intervals weakly con¬ vex, with fine and moderately dense punctures, with very minute setae in punctures, penultimate lateral interval with a few short sparse setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleu- ra moderately densely setose; apical border of elytra mem¬ braneous, with a fine rim of sparse microtrichomes (vis¬ ible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally. Abdominal sternites finely and densely punctate, sparsely setose, each stemite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Py- gidium convex and dull, coarsely and densely punctate, without smooth midline, glabrous except numerous longer setae beside apical margin. Legs moderately wide and long, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, without ser¬ rated line behind anterior edge, anterior row of setae lack¬ ing, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia moderately long, widest at apex, ratio of width/length: 1/2.78, sharply carinate dorsally, with two groups of spines, basal group at one third, api¬ cal group at three quarters of metatibial length, with a sin¬ gle short spine basally; lateral face longitudinally convex, finely and densely punctate in basal half, glabrous, im- punctate in apical half; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately concavely sin¬ uate interiorly near tarsal articulation. Tarsomeres dorsal¬ ly smooth and glabrous, neither laterally nor dorsally car¬ inate, sparsely setose ventrally; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a smooth sub- ventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, biden- tate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9D-F. Habitus: Fig. 30J. Diagnosis. The new species is very similar to “ Aserica major Arrow, 1946” from Myanmar which formally also belongs to Neoserica sensu lato, but it will be revised in a separate paper. N. pseudomajor differs in the slightly shorter antennal club and the shape of aedeagus: the dis¬ tal portion of the phallobase is narrower and not that broadly pronounced as in A. major ; the distal lateral mem¬ braneous slit is in A. major distinctly longer than in N. pseudomajor. Etymology. The species’ name (adjective in the nomina¬ tive singular) is composed by the Greek prefix ‘ pseudo ’ (false) and the species name ‘ major ’ (greater), with ref¬ erence to its similarity to Aserica major. Variation. Length: 8.8-10.0 mm, length of elytra: 6.2-6.9 mm, width: 5.3-5.8 mm. Female: Eyes slightly smaller than in male, antennal club short, composed by three an¬ tennomeres. Distribution. See map (Fig. 46B). Neoserica pushkarensis sp. n. (Figs 9G-I, 3OK, 46B) Type material examined. Holotype: 8 “India Rajastan Pushkar 7/1995 R. Sauer” (ZFMK). Paratype: 1 8 C-In- dia VIII. 1988 Panna Nat. Park MP, Wemer lgt./ 623 Serici¬ ni Asia spec.” (ZFMK). Description. Length: 6.2 mm, length of elytra: 4.2 mm, width: 3.5 mm. Body oval, reddish brown, antenna yel¬ lowish brown, dorsal surface shiny, except some single se¬ tae on head dorsal surface nearly glabrous. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 116 Dirk Ahrens & Silvia Fabrizi Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins convex and convergent anteriorly, anterior an¬ gles moderately rounded, anterior margin shallowly sin¬ uate, margins strongly reflexed; lateral margin and ocu¬ lar canthus produce a blunt angle; surface nearly flat, fine¬ ly and very densely punctate, with numerous coarse punc¬ tures each bearing an erect seta; frontoclypeal suture fine¬ ly incised, weakly angled medially; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and densely punctate, with a terminal seta. Frons with fine, dense punc¬ tures and a few erect setae beside eyes. Eyes moderately large, ratio diameter/ interocular width: 0.62. Antenna with ten antennomeres; club with five antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins in basal half subparallel, at middle convex and in anterior third straight and distinctly convergent anteri¬ orly, anterior angles distinctly produced and sharp, pos¬ terior angles blunt; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, glabrous; later¬ al and anterior margin sparsely setose; hypomeron cari¬ nate, slightly produced ventrally. Scutellum wide, trian¬ gular, shiny, at apex moderately pointed, with fine, dense punctures, with minute setae in punctures. Elytra widest at middle, striae distinctly impressed, fine¬ ly and densely punctate, intervals weakly convex, with fine and moderately dense punctures concentrated along stri¬ ae, except very minute setae in punctures only a few short, white setae on external intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura moderately densely setose; apical border of ely¬ tra membraneous, with an fine rim of sparse microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally being not condensed in a transverse line. Abdominal sternites finely and densely punctate, sparse¬ ly setose, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/1.56. Pygid- ium weakly convex and shiny, coarsely and sparsely punc¬ tate, without smooth midline, glabrous except a few sparse longer setae along apical margin. Legs moderately wide and long, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without serrated line behind anterior edge, anterior row of setae complete, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia moderately long, widest at apex, ratio of width/length: 1/3.1, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face longitudinally convex, coarsely and densely punctate, glabrous, smooth along middle in apical half; ventral mar¬ gin finely serrate, with five robust equidistant setae; me¬ dial face smooth and glabrous; apex finely serrate, mod¬ erately truncate interiorly near tarsal articulation. Tar- someres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatar- someres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and slight¬ ly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9G-I. Habitus: Fig. 30K. Female un¬ known. Diagnosis. Neosericapushkarensis sp. n. differs from the most similar N. rajasthanica sp. n. by the slightly short¬ er antennal club, as well as by the thicker right paramere (lateral view). Etymology. The new species is named after its type lo¬ cality, Pushkar (adjective in the nominative singular). Variation. Length: 5.6-6.2 mm, length of elytra: 3.9-4.2 mm, width: 3.4-3.5 mm. Distribution. See map (Fig. 46B). Neoserica quadrilamellata (Brenske, 1896) (Figs 9J-L, 30L, 46B) Serica quadrilamellata Brenske, 1896: 154. Neoserica quadrilamellata: Brenske 1898: 253. Meriserica chilkensis Arrow, 1923: 260, syn. n. Type material examined. Holotype ( quadrilamellata ): 1 3 “Madras/ 4-lamellata typ. Brsk.” (ISNB). Syntypes {chilkensis)-. 1 3 “Barkuda Id., Chilka lake Ganjam dist. Madras Presid. 22.VII.22 (at light)/Ind. Mus. Coll. B. M. 1922-249/ 3/ Type/ Meriserica chilkensis type Arrow” (BMNH), 2 $$ “Barkuda Id., Chilka lake Ganjam dist. Madras Presid. 22.VII.22 (at light)/Ind. Mus. Coll. B. M. 1922-249” (BMNH), 1 3 “Barkuda Id., Chilka lake Gan¬ jam dist. Madras Pres. 25.VI.22 Amandale/Ind. Mus. Coll. B. M. 1922-127/ Meriserica chilkensis co-type Arrow” (BMNH), 1 3 “Barkuda Id., Chilka lake Ganjam dist. Madras Pres. 26.VI.22 at light/ 3 ” (BMNH), 1 3 “Ind. Mus. Coll. B. M. 1922-127/Barkuda Id., Chilka lake Gan¬ jam dist. Madras Pres. July 1920 at light” (BMNH), 1 § Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 117 “Barkuda Id., Chilka lake ganjam dist. Madras Pres. 25.VII-4.VIII.22 at light/Ind. Mus. Coll. B. M. 1922-249/ $” (BMNH), 1 S “Barkuda Id., Chilka lake Ganjam dist. Madras Pres. 25.VI.22 at light/ Ind. Mus. Coll. B. M. 1922-127/ S at light/ Meriserica chilkensis Arrow” (CF). Additional material examined. 2 ex. “India New Dehli VII. 1976 M. Helva leg./ IS 59” (CPPB), 4 ex. “India Madras Coimbatore 1400 ft., Nov. 1965 leg. P. S. Nathan” (ZFMK), 6 ex. “Museum Paris Inde Bellary, De Morgan 1896” (MNHN), 3 ex. “Ostinindien/ 16/2/2 Skovgaard” (ZMUC), 1 ex. “Vellore Ostindien J. Skovgaard 1094” (ZMUC), 2 ex. “Coll.R.I.Sc.N.B. Inde/ Inde/ Coll, de Bon- neuil Le Moult vendit” (ISNB), 1 ex. “India: Karnataka Dharwad Mar 1979 NUMR Coll./ CR 29/ Brit. Mus. 1984- 37” (BMNH), 1 ex. “India: Karnataka Dharawady May 1989 C.R. Coll./ CR 29/ Brit. Mus. 1984-37” (BMNH), 1 S “Siid-Indien Prov. Madras Coimbatore 12.X.64 P. Nathan leg.” (ZSM), 1 $ “India, Tamil Nadu, Coimbat¬ ore pr., xi.2002, Tiruppur env. S. Saluk leg.” (CPPB). Doubtful record: 1 S “Darjeeling Atkinson (Ribbe)/ [“Pasinalt”?]” (ZMHB). Redescription. Length: 6.4 mm, length of elytra: 4.4 mm, width: 4.1 mm. Body oval, reddish brown, antenna yel¬ lowish brown, dorsal surface shiny, except some single se¬ tae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins moderately convex and convergent anterior¬ ly, anterior angles moderately rounded, anterior margin distinctly sinuate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface slightly elevated medially, finely and very densely punc¬ tate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocu¬ lar diameter), finely and densely punctate, with a termi¬ nal seta. Frons with fine, dense punctures and a single erect seta beside each eye. Eyes moderately large, ratio diam¬ eter/interocular width: 0.7. Antenna with ten anten- nomeres; club with five antennomeres and straight, as long as remaining antennomeres combined, first joint of club only half as long as club. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins weakly but evenly convex and moderately con¬ vergent anteriorly, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly con¬ vex, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, glabrous; lateral and anterior margin sparsely setose; hy- pomeron carinate, slightly produced ventrally. Scutellum wide, triangular, shiny, at apex moderately pointed, with fine, dense punctures, glabrous. Elytra widest at middle, striae distinctly impressed, fine¬ ly and sparsely punctate, intervals flat, with fine and mod¬ erately dense punctures, on odd intervals concentrated along striae, except very minute setae in punctures only a few short, white setae on external intervals; epipleural edge robust, ending at strongly curved external apical an¬ gle of elytra, epipleura moderately densely setose; apical border of elytra transversely swollen and membraneous, with an broad rim of sparse microtrichomes (visible at ca lOOx magnification). Ventral surface shiny, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally being not condensed in a transverse line. Abdominal sternites finely and irregularly densely punc¬ tate, sparsely setose, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium weakly convex and shiny, coarsely and sparsely punctate, without smooth midline, glabrous ex¬ cept a few sparse longer setae along apical margin. Legs moderately wide and long, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without serrated line behind anterior edge, anterior row of setae complete, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia moderately long, widest at apex, ratio of width/length: 1/2.94; sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, at base glabrous; lateral face flat, coarsely and densely punctate, glabrous, wide¬ ly impunctate on midline; ventral margin finely serrate, with five robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interi¬ orly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparse¬ ly setose ventrally; metatarsomeres with a strongly ser¬ rated ridge ventrally and a smooth subventral longitudi¬ nal carina; first metatarsomere slightly shorter than fol¬ lowing two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9J-L. Habitus: Fig. 30L. Remarks. The male genitalia of the types of both species are virtually identical in shape, consequently, M. chilken¬ sis is to be considered a junior synonym of N. quadril- amellata. The length of the sixth antennomere is variable, in most specimens the antennal club appears to be com¬ posed of five antennomeres with the first joint of club half as long as the club. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 118 Dirk Ahrens & Silvia Fabrizi Distribution. See map (Fig. 46B). Neoserica quinqueflabellata (Brenske, 1896) (Figs 9M-0, 46B) Sericaquinqueflabellata Brenske, 1896: 153. Neoserica quinqueflabellata. Brenske 1898: 254. Type material examined. Syntype: 1 S “Tetara Cardon/ Serica quinqueflabellata Typ. Brsk.” (ISNB). Redescription. Length: 7.5 mm, length of elytra: 5.1 mm, width: 4.5 mm. Body oval, reddish brown, antenna yel¬ lowish brown, dorsal surface dull, except some single se¬ tae on head dorsal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins convex and convergent anteriorly, anteri¬ or angles strongly rounded, anterior margin shallowly sin¬ uate, margins moderately reflexed; lateral margin and oc¬ ular canthus produce an indistinct angle; surface flat, coarsely and very densely punctate, punctures partly fus¬ ing with each other, with a few erect setae; frontoclypeal suture indistinctly incised, weakly angled medially. Frons with fine, moderately dense punctures, with a few erect setae beside eyes. Eyes moderately large, ratio diameter/interocular width: 0.6. Antenna with ten anten- nomeres; club with five antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse, widest at base, lateral margins in basal half straight and subparallel, at middle convex and in anterior half distinctly convergent anteriorly, anterior angles distinctly produced and sharp, lateral margin be¬ fore anterior angles straight, posterior angles blunt; base without marginal line; surface sparsely and finely punc¬ tate, glabrous; lateral and anterior margin sparsely setose; hypomeron carinate, slightly produced ventrally. Scutel- lum wide, triangular, surface dull, at apex moderately pointed, with fine, dense punctures. Elytra strongly convex, widest at middle, striae weak¬ ly impressed, finely and densely punctate, intervals near¬ ly flat, except on second interval fine and dense punctures concentrated along striae, except very minute setae in punctures glabrous; epipleural edge , ending at strongly curved external apical angle of elytra, epipleura moder¬ ately densely setose; apical border of elytra membrane¬ ous, with an fine rim of sparse microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally being not condensed in a transverse line. Abdominal sternites finely and densely punctate, sparse¬ ly setose, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesostemum between mesocoxae as wide as mesofemur. Pygidium moderately convex and dull, coarsely and sparsely punc¬ tate, without smooth midline, glabrous except a few sparse longer setae along apical margin. Legs wide and short, dull; femora with two longitudi¬ nal rows of setae, finely and sparsely punctate. Metafe¬ mur with acute anterior margin, without serrated line be¬ hind anterior edge, anterior row of setae complete. Metat¬ ibia short and wide, widest at apex, ratio of width/length: 1/2.5, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at two thirds of metat- ibial length, with some single small spines and a short ser¬ rated line basally; lateral face weakly longitudinally con¬ vex, finely punctate, glabrous; apex moderately concave- ly sinuate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dor- sally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth sub- ventral longitudinal carina; first metatarsomere as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9M-0. Distribution. See map (Fig. 46B). Neoserica rajasthanica sp. n. (Figs 9P-R, 30M, 46A) Type material examined. Holotype: S “India N: Bharat- pur (Rajasthan) 15/VII/1995 legit Cianfanelli & M. Calcagno (Num Mag. 1764)/ 633 Sericini Asia spec.” (MZF). Paratype: 1 S “Rajasthan Sariska Lug. 83 Leg. Pad. Mai” (ZFMK), 1 £, 1 ■§ “Ajmer Indes angl./ Coll. R.I.Sc.N.B. Inde” (ISNB), 1 S “Inde Agra/Museum Paris 1938 J. Berlioz” (MNHN). Description. Length: 7.0 mm, length of elytra: 4.4 mm, width: 3.7 mm. Body oval, reddish brown, antenna yel¬ lowish brown, dorsal surface shiny, except some single se¬ tae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins moderately convex and convergent anterior¬ ly, anterior angles moderately rounded, anterior margin shallowly sinuate, margins strongly reflexed; lateral mar¬ gin and ocular canthus produce a blunt nearly indistinct angle; surface nearly flat, finely and very densely punc¬ tate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medially; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocu¬ lar diameter), finely and densely punctate, with a termi- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 119 nal seta. Frons with fine, very dense punctures and a few erect setae beside eyes. Eyes moderately large, ratio di¬ ameter/interocular width: 0.6. Antenna with ten anten- nomeres; club with five antennomeres and straight, slight¬ ly longer than remaining antennomeres combined. Men- tum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins in basal half subparallel, slightly concavely sin¬ uate, at middle convex and in anterior third straight and distinctly convergent anteriorly, anterior angles distinct¬ ly produced and sharp, posterior angles blunt; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface densely and fine¬ ly punctate, glabrous; lateral and anterior margin sparse¬ ly setose; hypomeron carinate, slightly produced ventral- ly. Scutellum wide, triangular, surface dull, at apex mod¬ erately pointed, with fine, dense punctures, with minute setae in punctures. Elytra widest at middle, striae distinctly impressed, fine¬ ly and densely punctate, intervals weakly convex, with fine and dense punctures concentrated along striae, except very minute setae in punctures only a few short, white setae on external intervals; epipleural edge robust, ending at strong¬ ly curved external apical angle of elytra, epipleura mod¬ erately densely setose; apical border of elytra membrane¬ ous, with an fine rim of sparse microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally being not condensed in a transverse line. Abdominal sternites finely and densely punctate, sparse¬ ly setose, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/1.5. Pygidi- um moderately convex and shiny, coarsely and densely punctate, without smooth midline, glabrous except a few sparse longer setae along apical margin. Legs moderately wide and long, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without serrated line behind anterior edge, anterior row of setae complete, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia moderately long, widest at apex, ratio of width/length: 1/2.7, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face longitudinally convex, coarsely and densely punctate, glabrous, smooth along middle in apical half; ventral mar¬ gin finely serrate, with five robust equidistant setae; me¬ dial face smooth and glabrous; apex finely serrate, mod¬ erately truncate interiorly near tarsal articulation. Tar- someres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatar- someres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere distinctly shorter than following two tarsomeres combined and slightly shorter than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9P-R. Habitus: Fig. 30M. Diagnosis. Neoserica rajasthanica sp. n. differs from N. quadrilamellata (Brenske) by the lack of a lateral process of the phallobase. Etymology. The new species is named after its occurrence in Rajasthan (adjective in the nominative singular). Variation. Length: 6.5-7.0 mm, length of elytra: 4.1—4.4 mm, width: 3.6-3.7 mm. Female: antennal club as long as remaining antennomeres combined, first joint of club half as long as remaining four joints. Distribution. See map (Fig. 46A). Neoserica rutilans Ahrens & Fabrizi, 2009 Neoserica rutilans Ahrens & Fabrizi, 2009b: 261. Material examined. See Ahrens & Fabrizi 2009b (p. 261). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 4P-R, p. 280). Distribution. NE India, W-Arunachal Pradesh (Fig. 46A). Neoserica setigera (Brenske, 1894) comb. n. (Fig. 9S-U, 46B) Serica setigera Brenske, 1894: 11, 54; Brenske 1898: 235. Type material examined. Syntypes: 1 S “India/ Coll. J. Thomson/ Typus/ setigera type Brsk.” (ISNB), 1 $ “Coll. R. I. Sc. N. B. Inde/ Coll. J. Thomson/ Serica setig¬ era Brsk. det. E. Brenske [not handwritten by Brenske, no original but newly produced locality label of collec¬ tion management” (ISNB). Additional material examined. 1 S “Bombay” (ZFMK), 1 $ “Decan Saunder/ Serica setigera Brsk. Compared with type G.J.A./ pallida Decan Reiche/ so named in Reiches collection C.W” (BMNH), 2 SS “India, Maharahstra [sic] state, Alibag env., 45 km S Bombay, 22.-24.vi.2006, O. Safranek leg.” (CPPB, ZFMK). Redescription. Length: 6.5 mm, elytral length: 4.9 mm, width: 3.7 mm. Body oblong-oval, yellowish brown, dor- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 120 Dirk Ahrens & Silvia Fabrizi sal surface with double pilosity and shiny, without dull to- ment, finely and densely punctate. Labroclypeus weakly trapezoidal, lateral margins pro¬ duce an indistinct angle with the ocular canthus; anterior angles strongly convex, anterior margin shallowly sinu¬ ate; margins weakly reflexed; surface weakly convex me¬ dially, with very dense punctures of variable size. Frons with dense punctures of variable size, with fine yellow, adpressed setae mixed with a few longer erect ones. An¬ tenna yellow, with ten antennomeres, club in male com¬ posed of five antennomeres and straight, distinctly longer than the remaining antennomeres combined. Eyes mod¬ erately large, ratio diameter/interocular width: 0.65. Mentum slightly elevated and convex anteriorly. Pronotum narrow and elongate; lateral margins in basal half subparallel, in anterior third convex and convergent; anterior angles very sharp and distinctly produced, pos¬ terior angles nearly right-angled; basis strongly sinuate, without marginal line; surface with coarse and shallow, dense punctures, distance between punctures smaller than their diameter, with fine, dense, adpressed setae, anterior and lateral margins with long erect setae. Scutellum short, triangular, pilosity and punctation similar to that of prono¬ tum. Elytra elongate, striae weakly impressed, intervals weakly flat and evenly densely punctate; epipleura with dense and robust setae; apical margin of elytra with mem¬ braneous rim. Ventral surface including legs finely densely setose. Mesosternum between mesocoxae as wide as width of mesofemur. Abdominal sternites densely punctate, each sternite with a transversal row of more robust punctures each bearing an erect seta. Pygidium distinctly convex, with double pilosity. Legs moderately wide; metatibia short, ratio length/with: 1/3.2; dorsal margin carinate, with two external groups of spines, basal group at half, apical one at three quarter of metatibial length, basally with two single robust setae; lat¬ eral face longitudinally convex, evenly and densely punc¬ tate, finely setose; ventral margin with robust setae; me¬ dial face completely smooth and glabrous, apex near tarsal articulation weakly truncate. Tarsi dorsally impunctate and glabrous; metatarsomeres ventrally with a strongly serrat¬ ed carina, subventrally with a second, smooth longitudi¬ nal carina, metatarsomere one slightly longer than dorsal tibial spur and distinctly shorter that two following tar- someres combined. Protibia moderately long, bidentate. All claws symmetrical, feebly curved and long, with nor¬ mally developed and simply pointed basal tooth. Aedeagus: Fig. 9S-U. Distribution. See map (Fig. 46B). Neoserica sexfoliata Moser, 1915 Neoserica sexfoliata Moser, 1915a: 166; Fabrizi & Ahrens 2014: 36. Material examined. See Fabrizi & Ahrens 2014 (p. 36); 1 ex. “Sri Lanka = Ceylon Kandy. Summer 1994 leg. G. Strickroth” (CARL). Aedeagus. See Fabrizi & Ahrens 2014 (fig. 5G-I, p. 105). Distribution. Endemic to Southern India and Sri Lanka (Fig. 46A). Neoserica sforziae sp. n. (Figs 9V-Y, 3ON, 46A) Type material examined. Holotype: S “S-India; Tamil Nadu, Tiruchchirappalli distr. fra Perambalur e Turaiyur, alle luci 19.X.1997 legit A. Sforzi & L. Bartolozzi (num Mag. 2091)/ 619 Sericini Asia spec.” (MZF). Description. Length: 5.8 mm, length of elytra: 3.7 mm, width: 3.3 mm. Body oval, reddish brown, antenna yel¬ lowish brown, dorsal surface shiny, except some single se¬ tae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin shallow¬ ly sinuate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface slightly el¬ evated medially, finely and very densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, weakly angled medi¬ ally; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocular diam¬ eter), finely and densely punctate, with a terminal seta. Frons with fine, dense punctures and a few erect setae be¬ side eyes. Eyes moderately large, ratio diameter/interoc- ular width: 0.67. Antenna with ten antennomeres; club with five antennomeres and straight, as long as remain¬ ing antennomeres combined. Mentum elevated and slight¬ ly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins in basal half subparallel, at middle convex and in anterior third straight and distinctly convergent anteri¬ orly, anterior angles distinctly produced and sharp, pos¬ terior angles blunt; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, glabrous; later¬ al and anterior margin sparsely setose; hypomeron cari¬ nate, slightly produced ventrally. Scutellum wide, trian¬ gular, shiny, at apex moderately pointed, with fine, dense punctures, with minute setae in punctures. Elytra widest at middle, striae distinctly impressed, fine¬ ly and sparsely punctate, intervals flat, with fine and mod- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 121 erately dense punctures concentrated along striae, except very minute setae in punctures only a few short, white se¬ tae on external intervals; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleu- ra moderately densely setose; apical border of elytra mem¬ braneous, with an fine rim of sparse microtrichomes (vis¬ ible at ca lOOx magnification). Ventral surface shiny, coarsely and densely punctate, metasternum sparsely covered with fine, short or very minute setae; metacoxa with minute setae in punctures that become laterally longer, with a few single robust and long setae laterally being not condensed in a transverse line. Abdominal stemites finely and irregularly densely punc¬ tate, sparsely setose, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.6. Pygidium weakly convex and shiny, coarsely and sparsely punctate, without smooth midline, glabrous ex¬ cept a few sparse longer setae along apical margin. Legs moderately wide and long, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without serrated line behind anterior edge, anterior row of setae complete, posterior margin smooth and extremely widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia moderately long, widest at apex, ratio of width/length: 1/2.8, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, glabrous basally; lateral face longitudinally convex, coarsely and densely punctate, glabrous; ventral margin finely serrate, with five robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal ar¬ ticulation. Tarsomeres dorsally smooth and glabrous, nei¬ ther laterally nor dorsally carinate, sparsely setose ventral¬ ly; metatarsomeres with a strongly serrated ridge ventral¬ ly and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetri¬ cal, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 9V-Y. Habitus: Fig. 3ON. Diagnosis. Neoserica sforziae sp. n. differs from the most similar N. rajasthanica sp. n. and N. pushkarensis sp. n. by the smaller body, the shiny ventral surface and by the shape of aedeagus. Etymology. The new species is named after one of its col¬ lectors, Alessandra Sforzi (noun in genitive case). Distribution. See map (Fig. 46B). Neoserica sladeni Ahrens, 2004 Neoserica sladeni Ahrens, 2004b: 156. Material examined. See Ahrens 2004b (p. 156). Aedeagus: See Ahrens 2004b (figs 232-235, p. 416). Distribution. Endemic to Mishmi Hills (Assam) (Fig. 46A). Neoserica subsetosa sp. n. (Figs 10A-D, 300, 46A) Type material examined. Holotype: $ “N. Bengal/ 612 Sericini Asia spec.” (BMNH). Paratypes: 1 <$, 1 $ “In¬ dia occ. Maharshtra [sic] st. Bhushi Dam env. 24-28.ix. 4 km S of Lonavala 2005 leg. F. & L. Kantner 500 m” (SMNS, ZFMK). Description. Length: 5.8 mm, elytral length: 4.3 mm, width: 3.3 mm. Body elongate, yellowish brown, dorsal surface with very dense, simple pilosity and shiny, with¬ out dull toment, finely and densely punctate. Labroclypeus trapezoidal, lateral margin weakly convex and moderately convergent anteriorly, anterior angles strongly convex, anterior margin shallowly sinuate, mar¬ gins weakly reflexed, lateral margins produce an indistinct angle with ocular canthus; surface weakly convex medi¬ ally, with fine, dense punctures. Ocular canthus short and wide, finely sparsely punctate, glabrous. Frontoclypeal su¬ ture finely incised and moderately curved. Frons with fine and coarser, dense punctures, with fine yellow, except a few fine setae beside eyes glabrous. Antenna yellow, with ten antennomeres, club in male composed of five anten- nomeres and straight, distinctly longer than remaining an¬ tennomeres combined. Eyes moderately large, ratio diam¬ eter/interocular width: 0.56. Mentum slightly elevated and convex anteriorly. Pronotum narrow and elongate; lateral margins weak¬ ly evenly convex and convergent anteriorly; anterior an¬ gles very sharp and distinctly produced, posterior angles nearly right-angled; base strongly sinuate, without mar¬ ginal line; surface with fine and shallow, dense punctures, distance between punctures smaller than their diameter, with fine, dense, adpressed setae, on sides with a few long erect setae; anterior and lateral margins with long setae. Scutellum short, triangular, pilosity and punctation simi¬ lar to that of pronotum. Elytra elongate, widest in posterior third; striae weak¬ ly impressed, finely punctate, intervals flat, finely, even¬ ly and densely punctate, with fine and dense, short setae being directed posteriorly; epipleura with dense and ro¬ bust setae; apical margin of elytra with fine membrane¬ ous rim. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 122 Dirk Ahrens & Silvia Fabrizi Ventral surface including legs finely densely punctate, with dense adpressed setae. Mesostemum between meso- coxae as wide as width of mesofemur. Metacoxa with fine and dense adpressed setae on entire surface. Abdominal sternites densely punctate and setose, each sternite with a transversal row of more robust punctures each bearing an erect robust seta. Pygidium distinctly convex, finely and densely punctate, with short fine setae and a few longer erect setae. Legs wide; metatibia short, ratio length/with: 1/2.6; dor¬ sal margin carinate, with two external groups of spines, basal group at half, apical one at three quarter of metati- bial length, basally with two single robust setae; lateral face longitudinally convex, finely, evenly and densely punctate, finely setose; ventral margin with robust setae; medial face completely smooth and glabrous, apex near tarsal articulation weakly truncate. Tarsomeres dorsally impunctate and glabrous, sparsely setose ventrally; metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina, first metatarsomere slightly longer than dorsal tibial spur and distinctly shorter that two following tarsomeres com¬ bined. Protibia moderately long, bidentate. All claws sym¬ metrical, feebly curved and long, with normally developed and simply pointed basal tooth. Aedeagus: Fig. 10A-D. Habitus Fig. 300. Diagnosis. This new species is very similar to N. setig- era (Brenske). Neoserica subsetosa sp. n. differs princi¬ pally by the shape of right paramere being (in lateral view) extended at its apex and dorsally strongly curved, instead of being deeply sinuate at its apex like in N. setigera. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the combined Latin prefix ‘ sub- (almost) and ‘setostf (setose). Variation. Length: 5.8-7.8 mm, elytral length: 4.3-5.4 mm, width: 3.3-4.1 mm. Female: Antennal club composed of five antennomeres, as long as remaining antennomeres combined. Distribution. See map (Fig. 46B). Gastroserica Brenske, 1897 Gastroserica Brenske, 1897: 412 (type species by subse¬ quent designation: Serica marginalis Brenske, 1894; Nomura 1973); Ahrens 2000f: 73,2007c: 15; Ahrens & Pacholatko 2003: 1,2007:137; Liuetal. 2011:23,2014b: 1. Remarks. There is so far only one species of this genus known from the Indian subcontinent: Gastroserica patkaiensis Ahrens, 2000. Gastroserica patkaiensis Ahrens, 2000 (Figs 3OP, 46C) Gastroserica patkaiensis Ahrens, 2000f: 108; Liu et al. 2014b: 108. Material examined. See Ahrens 2000f (p. 108); Liu et al. 2014b (p. 108). Aedeagus. See Ahrens 2000f (figs 64-66, p. 111). Distribution. Described from Assam, now also recorded for Yunnan, China (Liu et al. 2014b) (Fig. 46C). Tetraserica Ahrens, 2004 Tetraserica Ahrens, 2004b: 168 (type species by original designation: Neoserica gestroi Brenske, 1898); Ahrens 2007c: 38; Liu etal. 2014c: 83. Remarks. So far 37 species have been formally assigned to Tetraserica (Ahrens 2004b, Ahrens and Fabrizi 2009b, Liu et al. 2014c). All other oriental species (so far grouped with “ Neoserica ”) that potentially belong to Tetraserica species await taxonomic revision. Tetraserica differs from the two closely related genera Microserica Brenske, 1894 and Trioserica Moser, 1922 by the lack of a ventral cari¬ na on the hypomeron. From Microserica Tetraserica al¬ so differs by the lacking sexual dimorphism of the pygid¬ ium, from Trioserica by the bidentate protibia. In contrast to Microserica , species of Tetraserica are active at night and are attracted to light. Distribution. The genus is distributed almost in the en¬ tire Oriental region; we know described species so far as¬ signed to “ Neoserica ” from Philippines, Indochina, Sumatra, Borneo, and China (Ahrens 2004b; Liu et al. 2014c). Except for Meghalaya and the Himalaya, the genus does not occur on the Indian subcontinent south of the Ganges. Key to the species of Tetraserica of the Indian subcon¬ tinent ( 33 ) 1 Punctation of elytra very dense, concentrated exclu¬ sively along striae, intervals strongly convex. Dorsal margin of metatibia longitudinally convex ( Tetraser¬ ica sensu lato).9 1 ’ Punctation of elytra dense, not exclusively concentrat¬ ed along striae; intervals weakly convex to flat. Dor¬ sal margin of metatibia sharply carinate.2 2 Antennomeres of antennal club evenly long, first seg¬ ment at maximum 1/10 of club length, shorter.3 2’ First segment of antennal club distinctly shorter (by at least 1/4 of club length), club sometimes only with Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 123 three antennomeres. T. ferrugata (Blanchard) 3 Metatibia shorter, ratio width/length > 1/3.0.4 3’ Metatibia longer, ratio width/length < 1/3.2.5 4 Dorsal lobe of right paramere with a narrow comb-like basal lobe. T. nmgbongensis Ahrens 4’ Dorsal lobe of right paramere simply pointed, with¬ out a narrow comb-like basal lobe. T. impar sp. n. 5 Articulation of right paramere displaced dorsally. Dor¬ sal lobe of right paramere broad, apically split abrupt¬ ly into many long spikes. T. brahmaputrae Ahrens 5’ Articulation of right paramere not displaced. Dorsal lobe of right paramere narrow, without multiple api¬ cal spikes.6 6 Eyes small, ratio diameter/interocular width <0.5. An¬ tennal club short, as long as remaining antennomeres combined. Sides of pronotum light brown. . T. mfimargo sp. n. 6’ Eyes moderately large, ratio diameter/interocular width >0.6. Antennal club longer, at least 1.2 times as long as remaining antennomeres combined.7 7 Right paramere composed of two distinct lobes. . T. univestris sp. n. 7’ Right paramere composed of one distinct lobe.8 8 Right paramere in lateral view spherical, with numer¬ ous sharp spikes on dorsal margin. Left paramere not displaced dorsally, its dorsal lobe extremely long, with¬ out apical hook. T. bendai sp. n. 8’ Right paramere in lateral view elongate, without spikes on dorsal margin. Left paramere displaced dorsally, its dorsal lobe with a strong apical hook. . T. uncinata sp. n. 9 Parameres nearly equally long.10 9’ Left paramere much longer than right. . T. crenatula Ahrens & Fabrizi 10 Metatibia shorter and wider, ratio length/width: 1/2.96. Left paramere shorter than right. Body short-oval, < 6 mm. T. disoccupata Ahrens 10’ Metatibia narrower, ratio length/width < 1/3.2. Para¬ meres highly similar in length. Body elongate, >7.0 mm.11 11 Left paramere at apex with external tooth. . T. schneideri Ahrens 11 ’ Left paramere at apex without external tooth. . T. hilaris Ahrens & Fabrizi Tetr as erica bendai sp. n. (Figs 10E-G, 31A, 46D) Type material examined. Holotype: 8 “NE India; Meghalaya, 2002 3 km E of Tura, 1150m, 25°30’N, 90°14’E; 12.V. M. Tryzna & P. Benda lgt.” (CPPB). Paratypes: 10 88, 1 $ “NE India; Meghalaya, 2002 3 km E of Tura, 1150m, 25°30’N, 90°14 , E; 6.-12.V. M. Tryz¬ na & P. Benda lgt” (CPPB, ZFMK), 3 88, 1 ? “NE In¬ dia; Meghalaya, 1999 3 km E of Tura, 1150m, 25°30’N, 90°14’E; 4.v. Dembicky & Pacholatko lgt.” (CPPB), 2 88, 1 ? “NE India; Meghalaya, 1999 3 km E of Tura, 500-1150m, 25°30’N, 90°14’E; 1.-8.V. J. Rolcik lgt.” (CPPB), 12 88, 10 ff “NE India; Meghalaya; 1400 m; Nokrek n.p. 3km S Daribokgiri 25°27’N 90°19’E; 26.iv.1999 Dembicky & Pacholatko leg.” (ZFMK), 1 8 “NE India, Meghalaya state West Garo Hills, Nokrek Nat. Park 9-17.V.1996 alt. 1100+150m GPS N25°29.6’, E90°19.5 (WGS 84) E. Jendek & O. Sausa/ IS 40” (CPPB). Description. Length: 9.9 mm, length of elytra: 6.8 mm, width: 5.2 mm. Body oval, dark brown, ventral surface reddish brown, antenna yellowish brown; dorsal surface dull and glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and conver¬ gent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.69. Antenna with ten anten¬ nomeres; club composed of four antennomeres, straight, nearly as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and strongly convex, later¬ al margins evenly convex, more strongly narrowed ante¬ riorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles blunt or strong¬ ly rounded. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum tri¬ angular, finely and densely punctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and moderately densely punctate, in¬ tervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punc¬ tures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.34. Abdominal sternites finely and densely punctuate, with a transverse row of Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 124 Dirk Ahrens & Silvia Fabrizi coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.2; dorsal margin sharply carinate, with two groups of spines, basal group of dor¬ sal spines of metatibia at first third of metatibial length; lateral face finely and sparsely punctate; ventral edge fine¬ ly serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shal¬ low sinuation. Tarsomeres with fine, very dense setae ven¬ trally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrat¬ ed ridge ventrally and glabrous; first metatarsomere slight¬ ly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10E-G. Habitus: Fig. 31 A. Diagnosis. The only so far known species with which this new species shares the spherical shape of the right para- mere is T. ruiliana Liu, Fabrizi, Bai, Yang & Ahrens, 2014. T. bendai differs from T. ruiliana in the flatter right para- mere, and in the shorter ventral lobe of the left paramere. Etymology. The new species is named after one of its col¬ lectors, P. Benda (noun in genitive case). Variation. Length: 9.3-10.0 mm, length of elytra: 67-7.2 mm, width: 5.2-5.9 mm. Female: club composed of three antennomeres, as long as the remaining antennomeres combined. Distribution. See map (Fig. 46D). Tetraserica brahmaputrae Ahrens, 2004 (Figs 3IB, 46D) Tetraserica brahmaputrae Ahrens, 2004b: 172. Material examined. See Ahrens 2004b (p. 172); 7 SS, 4 $ $ “NE India, Meghalaya, 1 tan E of Turn, 500 - 600m, 25°30‘N, 90°14‘E, 2.-5.V.2002, M.Tryzna & P. Benda leg.”(CPPB), 1 cj, 1 ? “NEIndia, Meghalaya, 1 kmEof Tura, 500 - 600m, 25°30‘N, 90°14‘E, 13.-18.V.2002, M.Tryzna & P. Benda leg.” (CPPB), 1 S “NE India, Meghalaya ~8 km N of Shillong, 25°38’N 91°54’E; ~1200m, L. Dembicky leg., 7.-9.V.2004” (CPPB), 2 SS “NE India, Arunachal Pr. 8 km S Jamiri - Sesa vicinity, 27°07’-09’N 92°34’E, 26.v.-4.vi.2005; 350 m, P. Pa- cholatko leg.” (CPPB, ZFMK). Aedeagus. See Ahrens 2004b (figs 272-274, p. 422) Distribution. Bhutan, Assam, Meghalaya, E Nepal (Fig. 46D). Tetraserica crenatula Ahrens & Fabrizi, 2009 (Figs 31C, 46D) Tetraserica crenatula Ahrens & Fabrizi, 2009b: 267 Material examined. See Ahrens & Fabrizi 2009b (p. 267). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 6A-C, p. 282). Distribution. India (Arunachal Pradesh) (Fig. 46D). Tetraserica disoccupata Ahrens, 2004 Tetraserica disoccupata Ahrens, 2004b: 175. Material examined. See Ahrens 2004b (p. 169). Aedeagus. See Ahrens 2004b (figs 278-280, p. 423). Distribution. Endemic to Sikkim (Fig. 46D). Tetraserica ferrugata (Blanchard, 1850) Omaloplia ferrugata Blanchard, 1850: 78 (nec Blanchard, 1850: 82). Serica ferrugata : Brenske 1898: 229. Tetraserica ferrugata : Ahrens 2004b: 169, 2006a: 412. Autoserica alcocki Brenske, 1898: 304; syn. by Ahrens 2004b: 169. Serica alcocki'. Barlow, 1899: 242. Material examined. See Ahrens 2004b (p. 169), 2006a (p. 412). Aedeagus. See Ahrens 2004 (figs 266-268, p. 421). Distribution. India and Nepal (Kumaon-Himalaya and central Nepal) (Fig. 46D). Tetraserica hilaris Ahrens & Fabrizi, 2009 Tetraserica hilaris Ahrens & Fabrizi, 2009b: 266. Material examined. See Ahrens & Fabrizi 2009b (p. 266). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 5P-R, p. 281). Distribution. India (Arunachal Pradesh) (Fig. 46D). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 125 Tetraserica impar sp. n. (Figs 10A-J, 3ID, 46D) Type material examined. Holotype: $ “NE India Assam, 1999, 5 km N of Umrongso, 700 m, 25°27‘N 92°43‘E 17.- 25.V. Dembicky & Pacholatko leg.” (CPPB). Paratypes: 1 s, 2 $$ “NE India Assam, 1999, 5 km N of Umrong¬ so, 700 m, 25°27‘N 92°43‘E 17.-25.V. Dembicky & Pa¬ cholatko leg.” (ZFMK, CPPB), 1 S, 1 $ “NE India, As¬ sam, 2002, Umrongso vill env., 700 m, 25°27‘N 92°43‘E, 3.-8.v. Dembicky & Pacholatko leg.” (CPPB), 1 S, 2 $ $ “NE India Assam, 5 km N of Umrongso, 700 m, 25°27‘N 92°43‘E 21.V. 1999, Dembicky & Pacholatko leg.” (CPPB). Description. Length: 7.0 mm, length of elytra: 4.6 mm, width: 4.2 mm. Body oval, dark brown, labroclypeus, legs and ventral surface reddish brown, antenna yellowish brown; dorsal surface dull and glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and conver¬ gent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.62. Antenna with ten anten- nomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and strongly convex, later¬ al margins evenly convex, more strongly narrowed ante¬ riorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles strongly round¬ ed. Surface finely and densely punctate, except minute se¬ tae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and moderately densely punctate, in¬ tervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punc¬ tures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/ metacoxa: 1/1.71. Abdominal sternites finely and dense¬ ly punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical mar¬ gin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/2.9; dorsal margin sharply carinate, with two groups of spines, basal group of dor¬ sal spines of metatibia at first third of metatibial length; lateral face finely and sparsely punctate; ventral edge fine¬ ly serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shal¬ low sinuation. Tarsomeres with fine, very dense setae ven¬ trally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrat¬ ed ridge ventrally and glabrous; first metatarsomere slight¬ ly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10H-J. Habitus: Fig. 3ID. Diagnosis. This new species is very similar in shape of parameres to Tetraserica miniatula Moser, 1915 comb. n. from Pegu (Myanmar). T. impar differs by the shorter ven¬ tral lobe of the right paramere, and the much shorter dor¬ sal lobe of the left paramere. Variation. Length: 6.7-7.2 mm, length of elytra: 4.6-5.1 mm, width: 4.1—4.4 mm. Female: club composed of three antennomeres, as long as the remaining antennomeres combined. Etymology. The new species is named with the Latin ad¬ jective, ‘ impar’{ unequal), with reference to being differ¬ ent to its relative T. miniatula (adjective in the nomina¬ tive singular). Distribution. See map (Fig. 46D). Tetraserica rufimargo sp. n. (Figs 10K-M, 3IE, 46D) Type material examined. Holotype: S “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E, 500-950m, L. Dembicky leg., 29.iv.-2.v.2005” (CPPB). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 126 Dirk Ahrens & Silvia Fabrizi Paratypes: 2 S3 “NE India, Meghalaya, SW of Cherra- punjee, 25°13‘-14‘N, 91°40‘ E, 5.-24.V.2005, 900 m, P. Pacholatko leg.” (CPPB, ZFMK). Description. Length: 6.9 mm, length of elytra: 4.9 mm, width: 4.5 mm. Body oval, dark brown, ventral surface and lateral margins of pronotum reddish brown, legs and antenna yellowish brown; dorsal surface dull and glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and conver¬ gent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.49. Antenna with ten anten- nomeres; club with four antennomeres, straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and strongly convex, later¬ al margins evenly convex, more strongly narrowed ante¬ riorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles blunt or strong¬ ly rounded. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum tri¬ angular, finely and densely punctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and moderately densely punctate, in¬ tervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punc¬ tures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.41. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.4; dorsal margin sharply carinate, with two groups of spines, basal group of dor¬ sal spines of metatibia at first third of metatibial length; lateral face finely and sparsely punctate; ventral edge fine¬ ly serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shal¬ low sinuation. Tarsomeres with fine, very dense setae ven¬ trally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrat¬ ed ridge ventrally and glabrous; first metatarsomere slight¬ ly longer than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10K-M. Habitus: Fig. 31E. Female un¬ known. Diagnosis. Tetraserica rufimargo sp. n. resembles in ex¬ ternal appearance and genital morphology somewhat T. ferrugata (Blanchard). T. rufimargo sp. n. differs, how¬ ever, by the first segment of the antennal club being sube¬ qual in length with the club, and the shape of the para- meres: the left paramere is apically not sharply pointed, the right paramere possesses instead of the hook-like basal lobe multiple spikes, its ventral lobe is wider and round¬ ed at apex. Etymology. The name (noun in apposition) of the new species is derived from the combined Latin adjectives, ‘ ru- fus' (red) and ‘ margo 5 (margin), with reference to the red¬ dish margin of the pronotum. Variation. Length: 6.9-8.0 mm, length of elytra: 4.9-5.5 mm, width: 4.5-4.9 mm. Distribution. See map (Fig. 46D). Tetraserica rungbongensis Ahrens, 2004 Tetraserica rungbongensis Ahrens, 2004b: 171. Material examined. See Ahrens 2004b (p. 171). Aedeagus. See Ahrens 2004b (figs 269-271, p. 422). Distribution. Endemic to Sikkim (Fig. 46D). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 127 Tetraserica schneideri Ahrens, 2004 Tetraserica schneideri Ahrens, 2004b: 173. Material examined. See Ahrens 2004b (p. 173). Aedeagus. See Ahrens 2004b (figs 275-277, p. 423). Distribution. Endemic to Sikkim (Fig. 46D). Tetraserica uncinata sp. n. (Figs 10N-P, 3IF, 46D) Type material examined. Holotype: <3 “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-15‘N, 91°47‘E, 500-900m, F. Dembicky leg., ll.-12.v.2004b/ 46/04” (CPPB). Paratypes: 1 1 ? “NE India Meghalaya state, Jaintia Hills reg., Jowai 6.-8.VI.1996 alt. 1350 m ± 100 m GPS N25°27‘ E92°12‘, (WGS 84) E. Jendek & O. Sausa/ IS 54/ 423 Sericini Asia spec.” (ZFMK, CPPB). Description. Length: 8.4 mm, length of elytra: 5.8 mm, width: 4.6 mm. Body oval, reddish brown, labroclypeus, legs and ventral surface little lighter, antenna yellowish brown; dorsal surface dull and glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and conver¬ gent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Eyes large; ratio of diameter/inte¬ rocular width: 0.79. Antenna with ten antennomeres; club composed of four antennomeres, straight, 1.4 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and strongly convex, later¬ al margins evenly convex, more strongly narrowed ante¬ riorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles strongly round¬ ed. Surface finely and densely punctate, except minute se¬ tae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and moderately densely punctate, in¬ tervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punc¬ tures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.6. Abdominal stemites fine¬ ly and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, densely punctate, without smooth mid¬ line, almost glabrous, but with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.3; dorsal margin sharply carinate, with two groups of spines, basal group of dor¬ sal spines of metatibia at first third of metatibial length; lateral face finely and sparsely punctate; ventral edge fine¬ ly serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shal¬ low sinuation. Tarsomeres with fine, very dense setae ven¬ trally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrat¬ ed ridge ventrally and glabrous; first metatarsomere slight¬ ly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10N-P. Habitus: Fig. 3IF. Diagnosis. The new species resembles in its morphology of the aedeagus T. miniatula (Moser), however, the dor¬ sal lobe of the right paramere is reduced, and the dorsal lobe of the left paramere is short and strongly hooked at its apex. Variation. Length: 8.4-9.4 mm, length of elytra: 5.8-6.1 mm, width: 4.6-5.4 mm. Female: club composed of three antennomeres, as long as the remaining antennomeres combined. Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived from the Latin adjec¬ tive ‘ uncinatus ’ (hooked), with reference to the shape of the dorsal lobe of the left paramere. Distribution. See map (Fig. 46D). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 128 Dirk Ahrens & Silvia Fabrizi Tetraserica univestris sp. n. (Figs 10Q-S, 31G, 46D) Type material examined. Holotype: S “Assam valley/ Doherty/ Fry Coll. 1905.100./ S n (BMNH). Description. Length: 7.9 mm, length of elytra: 5.3 mm, width: 4.7 mm. Body oval, dark brown, labroclypeus, legs and ventral surface reddish brown, antenna yellowish brown; dorsal surface dull and glabrous. Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and conver¬ gent to strongly rounded anterior angles, anterior margin weakly sinuate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinct¬ ly incised, flat and weakly curved medially; smooth area anterior to eye twice as wide as long; ocular canthus short and triangular, impunctate, with a single terminal seta. Frons dull, with sparse, fine punctures, with single erect setae beside each eye. Eyes moderately large; ratio of di¬ ameter/interocular width: 0.6. Antenna with ten anten- nomeres; club composed of four antennomeres, straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide and strongly convex, later¬ al margins evenly convex, more strongly narrowed ante¬ riorly towards sharp and slightly produced anterior angles. Anterior margin of pronotum slightly convex, with fine complete marginal line. Posterior angles strongly round¬ ed. Surface finely and densely punctate, except minute se¬ tae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate. Elytra oblong, widest just behind middle, striae distinct¬ ly impressed, finely and moderately densely punctate, in¬ tervals distinctly convex, with coarse and dense punctures concentrated along striae, with very minute setae in punc¬ tures; epipleural edge robust, ending at weakly curved and slightly blunt external apical angle of elytra, epipleura densely setose, apical border with a broad fringe of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface weakly shiny, finely and densely punc¬ tate, metasternum sparsely covered with fine, short, or very minute setae, metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.75. Abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, but with a few longer setae along apical margin. Legs moderately wide; femora finely and sparsely punc¬ tate; metafemur dull, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures absent; posterior margin smooth ventrally and only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally, without blunt tooth. Metatibia short and wide, widest at half of metati- bial length, ratio width/length: 1/3.3; dorsal margin sharply carinate, with two groups of spines, basal group of dor¬ sal spines of metatibia at first third of metatibial length; lateral face finely and sparsely punctate; ventral edge fine¬ ly serrated, with four robust equidistant setae, medial face smooth, apex interiorly near tarsal articulation with a shal¬ low sinuation. Tarsomeres with fine, very dense setae ven¬ trally on distal half, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres with a strongly serrat¬ ed ridge ventrally and glabrous; first metatarsomere slight¬ ly shorter than two following tarsomeres combined, one third of its length longer than dorsal tibial spine. Protib¬ ia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10Q-S. Habitus: Fig. 31G. Female un¬ known. Diagnosis. Tetraserica univestris sp. n. is very similar in the shape of the parameres to T. impar; however, the dor¬ sal lobe of the left paramere is reduced and the distal me¬ dian process of the phallobase is exceeding distinctly the ventral lobe of the right paramere. Furthermore, T. uni¬ vestris differs by the longer metatibia. Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived from the Latin adjec¬ tive, ‘ univestris ’ (concordant), with reference to strong similarity among the Tetraserica species’ external mor¬ phology. Distribution. See map (Fig. 46D). Trioserica Moser, 1922 Trioserica Moser, 1922: 111 (type species by subsequent designation: Trioserica lepichaeta Moser, 1922; Ahrens 2002a); Ahrens 2007c: 8. Sinoserica Miyake & Yamaya, 2001: 40; syn. by Ahrens 2007c: 8 (type species by original designation: Sinoser¬ ica maculipennis Miyake & Yamaya, 2001). Remarks. The genus differs from Tetraserica by the body colour being yellowish brown with often dense but dark spots of variable size, the antennal club in male having four or five antennomeres, the prothorax having a basal¬ ly carinate hypomeron, as well as the protibia being tri- dentate. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 129 Distribution. The genus is known so far from the Philip¬ pines (Moser 1922) as well from Taiwan, China and the Himalaya (Ahrens 2002a, 2004b, 2007). Further species described as Neoserica are known to the authors from Bor¬ neo, Indochina and the Malay peninsula. Trios erica tar sat a (Brenske, 1894) (Figs 311, 46D) Serica tarsata Brenske, 1894: 11,41. Autoserica tarsata Brenske, 1898: 226. Trioserica tarsata : Ahrens 2004b: 176. Material examined. See Ahrens 2004b (p. 176); 1 S “NE India Meghalaya state W Garo Hills, Balphakram Nat. Park 22-27.V. 1996 alt. 400+150 m GPS N25°l 1 ’ E90°51 ’ (WGS 84) E. Jendek & O. Sausa” (CPPB), 1 S “NE In¬ dia W Meghalaya Garo Hills; Nokrek N. P. 25,40N 91,04E, 2.-13.VII. 1997 V. Sinaevleg., 1150 m” (CPPB). Aedeagus. See Ahrens 2004b (figs 281-283, p. 424). Distribution. Meghalaya (India) and Bhutan (Fig. 46D). Microserica Brenske, 1894 Microserica Brenske, 1894: 52 (type species by subse¬ quent designation: Serica quadrimaculata Brenske, 1894; Arrow 1946a); Ahrens 1998a: 29,2004b: 178,2007c: 25. Key to Microserica species (SS) 1 Basal portion of metatibia with a more or less contin¬ uously serrated line.2 1 2 3 4 5 Basal portion of metatibia without serrated line.3 2 Dorsomedian sinuation between parameres not or not much deeper than width of phallobase. Antennal club with five antennomeres. Uniformly reddish brown. .. . M. fairmairei Brenske 2’ Dorsomedian sinuation between parameres much deeper than width of phallobase. Antennal club with four antennomeres. Multicoloured, elytra with dark stripes on odd intervals. M. crenatostriata Ahrens 3 Antennal club with three antennomeres. . M. elegans (Frey) 3’ Antennal club with more than three antennomeres. 4 4 Antennal club with six antennomeres.5 4’ Antennal club with four or five antennomeres.7 5 Antennal club 1.5 times as long as remaining anten¬ nomeres combined. Right paramere very small, round¬ ed at apex. Elytra unicoloured. Pygidium in female strongly shiny. M. affinis Arrow 5’ Antennal club twice as long as remaining anten¬ nomeres combined. Right paramere as large as left one. Elytra multi-coloured. Pygidium in female dull.6 6 Metatibia moderately wide, ratio width/length: 1/3.2. Parameres nearly symmetrical. . M. pedongensis Ahrens 6’ Metatibia wide, ratio width/length: 1/2.8. Parameres asymmetrical. M. marginata (Brenske) 7 Antennal club with four antennomeres.8 7’ Antennal club with five antennomeres.15 8 Metatarsomere one as long as dorsal metatibial spine. .9 8’ Metatarsomere one distinctly longer than dorsal metatibial spine.10 8” Metatarsomere one half as long as dorsal metatibial spine.20 9 Body bi-coloured, elytra yellowish brown with dark suture. Body oblong. Lateral margins of pronotum in basal half parallel. M. steelei Ahrens 9’ Body unicoloured. Body short-oval. . M. pruinosa (Hope) 10 Metatibia short and wide, ratio width/length: 1/2.5. . .11 10’ Metatibia narrower, ratio width/length: < 1/3.0. ...13 11 Pronotum yellowish brown, with two dark spots. Ely¬ tra with distinct cross-like signature. . M. cechovskyi Ahrens 11 ’ Pronotum dark, at maximum with light lateral margins. Elytra unicoloured black, brown, or yellowish brown with dark margins.12 12 Apical lobe of right paramere long. . M. truncata (Brenske) 12’ Apical lobe of right paramere short. . M. bhutanensis Frey 13 Antennal club 1.5 times as long as remaining anten¬ nomeres combined. Metatibia narrow, ratio width/length: 1/3.8. M. hispidulaVtQy 13’Antennal club twice as long as remaining anten¬ nomeres combined and slightly reflexed. Metatibia wider, ratio width/length: 1/3.0-3.3.14 14 Phallobase with a distinct dorsal sinuation before apex (lateral view). M. myagdiana Ahrens 14’ Phallobase without dorsal sinuation before apex (lat¬ eral view). M. gandakiensis Ahrens 15 Right paramere very small, rounded at apex. Elytra unicoloured brown or blackish. Pygidium in female strongly shiny, with fine microreticulation. . M. viridicollis Arrow 15’ Right paramere as large as left one.16 16 Metatibia short and wide, ratio width/length: 1/2.5-2.8. .17 16’ Metatibia narrower, ratio width/length: 1/3.2. . M. martensi Ahrens 17 Basal lobe of left paramere as long as paramere itself. .18 17’ Basal lobe of left paramere distinctly shorter than para¬ mere itself. .19 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 130 Dirk Ahrens & Silvia Fabrizi 18 Right paramere rounded at apex, left paramere short. .M schawalleri Ahrens 18’ Right paramere truncate at apex and bluntly toothed, left paramere as long as right one. . M. arunensis Ahrens 19 Left paramere with a lateral somewhat reflexed tooth. . M. interrogator (Arrow) 19’ Left paramere without lateral tooth. . M. schulzei Ahrens 20 Dorsal surface of body strongly shiny. .M lucens Ahrens & Fabrizi 20’ Dorsal surface of body more or less dull.21 21 Phallobase ventrally on the left side only with an el¬ evated carina. M. dohertyi Ahrens & Fabrizi 21 ’ Phallobase ventrally on each side with an elevated ca¬ rina.M roingensis sp. n. Microserica affinis Arrow, 1946 Microserica affinis Arrow, 1946a: 272; Ahrens 2002b: 384, 2004b: 189. Material examined. See Ahrens 2002b: (p. 384). Aedeagus. See Ahrens 2002b (figs 1-3, P. 406). Distribution. Assam, Arunachal Pradesh and northern Myanmar (Fig. 46E). Microserica arunensis Ahrens, 1998 Microserica arunensis Ahrens, 1998a: 45; Ahrens 2004b: 182. Material examined. See Ahrens 1998a (p. 45). Aedeagus. See Ahrens 1998a (figs 35-37, p. 46). Distribution. Eastern Nepal (Fig. 46F). Microserica bhutanensis Frey, 1975 (Figs 31J, 46E) Microserica bhutanensis Frey, 1975b: 227; Ahrens 1998a: 37; 2004b: 182. Material examined. See Ahrens 1998a (p. 37); 27 SS, 5 $ $ “E India, Meghalaya, 2002, 3 km E Tura, 1150 m, 25°30’N, 90°14’E, 6.-12.V., M. Tryzna & P. Benda lgt.” (CPPB). Aedeagus. See Ahrens 1998a (figs 14-16, p. 36). Distribution. Bhutan and Meghalaya (India) (Fig. 46E). Microserica cechovskyi Ahrens, 1999 Microserica cechovskyi Ahrens, 1999a: 243; Ahrens 2004b: 184. Material examined. See Ahrens 1999a (p. 243). Aedeagus. See Ahrens 1999a (figs 1-3, p. 244). Distribution. Eastern Nepal (Fig. 46E). Microserica crenatostriata Ahrens, 2004 Microserica crenatostriata Ahrens, 2004c: 10. Material examined. See Ahrens, 2004c (p. 10); 1 S “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E, 500-950m, L. Dembicky leg., 29.iv.-24.v.2005” (CPPB), 1 1 $ “NE India, Meghalaya, SW of Cherra¬ punjee, 25°13‘-14‘ N, 91°40‘ E, 5.-24.V.2005, 900m, P. Pacholatkoleg”(CPPB),7 SS, 5 ?$ “NEIndia,Megha¬ laya, 1 km E of Tura, 500 - 600m, 25°30‘N, 90°14E, 2.- 5.V.2002, M. Tryzna & P. Benda leg.”(CPPB), 2 “NE India, Meghalaya, SW of Shillong, 1600m, 25°34‘N, 91°5LE, L. Dembicky leg., 14.V.2004” (CPPB), 2 2 99 “India: Meghalaya state E Khasi Hills, 11km SW Cherrapunjee, Laitkynsew, 21-24.iv.2008, 25° 13’N, 91° 39’E, 810m, Fikacek, Podalska, Sipek lgt.” (ZFMK). Aedeagus. See Ahrens 2004c (figs 11-13; p.ll). Distribution. Restricted to Meghalaya (Fig. 46E). Microserica dohertyi Ahrens & Fabrizi, 2009 Microserica dohertyi Ahrens & Fabrizi, 2009b: 268. Material examined. See Ahrens & Fabrizi 2009b (p. 268). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 6G-I, p. 282). Distribution. Endemic to Assam/Patkai Mts. (Fig. 46F). Microserica elegans (Frey, 1975) Autos erica elegans Frey, 1975b: 224. Microserica elegans : Ahrens 1998a: 38, 2004b: 184. Material examined. See Ahrens 1998a (p. 38). Aedeagus. See Ahrens 1998a (figs 20-22, p. 40). Distribution. Endemic to Bhutan (Fig. 46E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 131 Micros erica fairmairei Brenske, 1898 (Figs 10T-W, 3IK) Microserica fairmairei Brenske, 1898: 282. Type material examined. Lectotype (here designated): 5 “Sudl. Ost-Indien Coll. Fairmaire/ Serica fairmairei type Brsk.” (ZMHB). Additional material examined. 1 S, 2 $$ “Gates mer.ales Hindostan” (ZFMK), 3 “S-India, Tamil Nadu, Tiruchchirapalli distr.: fra Perambalur e Turaiyur, alle luci 19.X.1997 legit A. Sforzi & L. Bartolozzi (num Mag. 2091)” (MZF), 1 S “S-India, Kerala, PeryarNat. Re¬ serve: dint. Lake Palace Hotel 25.X.1997 legit A. Sforzi 6 L. Bartolozzi (num Mag. 2091)” (MZF), 1 $ “S-India, Tamil Nadu, boscaglia c/o confine N della Peryar Nat. Re¬ serve 24.X. 1997, alle luci- legit A. Sforzi & L. Bartoloz¬ zi (num Mag. 2091)” (MZF). Redescription. Length: 5.8 mm, length of elytra: 4.0 mm, width: 3.6 mm. Body oval, reddish brown, elytra weakly iridescent, dorsal surface dull and nearly glabrous. Labroclypeus short, nearly semicircular, widest at base, lateral margins convex and strongly convergent to strongly rounded anterior angles; anterior margin distinct¬ ly sinuate medially, margins weakly reflexed; lateral mar¬ gins and ocular canthus produce a blunt angle; surface weakly convex medially, moderately coarsely and dense¬ ly punctate, with a few short erect setae; frontoclypeal weakly impressed and curved medially. Frons dull, with dense, fine punctures, beside eyes and behind fronto¬ clypeal suture with a few single short setae. Smooth area in front of eyes twice as wide as long. Ocular canthus fine¬ ly densely punctate, moderately wide and long, without terminal seta. Eyes large, ratio of diameter/interocular width: 0.77. Antenna with ten antennomeres; club com¬ posed of five antennomeres, straight, as long as remain¬ ing antennomeres combined. Mentum elevated and slight¬ ly flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins nearly straight in basal half and moderately conver¬ gent anteriorly, in anterior half weakly convex and more strongly convergent to sharp and produced anterior angles. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; posterior angles blunt; surface finely and densely punc¬ tate, glabrous, lateral and lateral anterior margins sparse¬ ly setose. Hypomeron carinate, weakly produced ventral- ly. Scutellum triangular, finely and densely punctate. Elytra short, widest at middle, striae distinctly im¬ pressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, otherwise glabrous; epipleural edge robust, ending at slightly rounded exter¬ nal apical angle of elytra, epipleura densely setose, api¬ cal border membraneous, with a fringe of short microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7..Abdominal sternites finely punctate, with a trans¬ verse row of coarse punctures, each bearing a robust se¬ ta. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, with numer¬ ous short setae on apical half. Legs moderately wide; femora finely and sparsely punc¬ tate, with two longitudinal rows of longer setae. Metafe¬ mur dull, anterior margin acute, without submarginal ser¬ rated line; anterior row of seta-bearing punctures present; posterior margin smooth ventrally, only weakly widened in apical half, posterior margin smooth dorsally, with a few short setae basally. Metatibia short and wide, widest at half of metatibial length, ratio width/length: 1/2.6; dorsal mar¬ gin sharply carinate, with two groups of spines, basal group shortly behind middle, apical one at three quarter of metatibial length, in basal half with a fine serrated line beside dorsal margin, in basal third with a single addition¬ al robust seta; lateral face weakly longitudinally convex, finely and densely punctate, with minute setae; ventral edge finely serrate, with four robust equidistant setae; me¬ dial face smooth, apex interiorly near tarsal articulation bluntly truncate. Tarsomeres with fine, sparse setae ven¬ trally, neither laterally nor dorsally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrat¬ ed ridge ventrally and a parallel subventral smooth Cari¬ na immediately beside it; first metatarsomere distinctly shorter than two following tarsomeres combined, slight¬ ly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10T-W. Habitus: Fig. 3IK. Remarks. We were unable to locate the type locality more accurately. Microserica gandakiensis Ahrens, 1998 Microserica gandakiensis Ahrens, 1998a: 39; Ahrens 2004b: 183, 2006a: 412; Ahrens & Fabrizi 2011: 163. Material examined. See Ahrens 1998a (p. 39), 2004b (p. 183), 2006a (p. 412); Ahrens & Fabrizi 2009b (p.272), 2011 (p. 163). Aedeagus. See Ahrens 1998a (figs 23-25, p. 40). Distribution. Western central Nepal (Fig. 46E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 132 Dirk Ahrens & Silvia Fabrizi Microserica hispidula Frey, 1975 Microserica hispidula Frey, 1975b: 226; Ahrens 1998a: 37, 2004b: 183. Material examined. See Ahrens 1998a (p. 37), 2004b (p. 183). Aedeagus. See Ahrens 1998a (figs 17-19, p. 38). Distribution. Bhutan (Fig. 46E). Microserica interrogator (Arrow, 1946) (Figs 31L, 46E) Aserica interrogator Arrow, 1946a: 267. Autoserica interrogator Sabatinelli 1993: 621. Microserica interrogator. Ahrens 1998a: 34; 2004b: 180, 2006a: 412; Ahrens & Fabrizi 2009b: 273. Material examined. See Ahrens 1998a (p. 34), 2004b (p. 180), 2006a (p. 412); Ahrens & Fabrizi 2009b (p. 273). Aedeagus. See Ahrens 1998a (figs 8-10, p. 33). Distribution. Kumaon-Himalaya to eastern central Nepal (Fig. 46E). Microserica lucens Ahrens & Fabrizi, 2009 Microserica lucens Ahrens & Fabrizi, 2009b: 267. Material examined. See Ahrens & Fabrizi 2009b (p. 267). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 6D-F, p. 282). Distribution. Endemic to the eastern Himalaya (Arunachal Prasdesh) (Fig. 46E). Microserica mar gin at a (Brenske, 1896) (Figs 31M, 46F) Serica marginata Brenske, 1896: 154. Microserica marginata: Brenske 1898: 317; Ahrens 1998a: 32, 2004b: 179, 2006a: 412. Aserica prohlematica Arrow, 1946a: 269, syn. by Ahrens 1998a: 32. Autoserica unctiuscula Brenske, 1898: 304, syn. by Ahrens 1998a: 32. Material examined. See Ahrens 1998a (p. 32), 2004b (p. 179), 2006a (p. 412). Aedeagus. See Ahrens 1998a (figs 1-3, p. 31). Distribution. Eastern Nepal and the Darjeeling/Sikkim area (Fig. 46F). Microserica martensi Ahrens, 1998 Microserica martensi Ahrens, 1998a: 43; Ahrens 2004b: 183. Material examined. See Ahrens 1998a (p. 43), 2004b (p. 183). Aedeagus. See Ahrens 1998a (figs 31-33, p. 44). Distribution. Endemic to eastern Nepal (Fig. 46E). Microserica myagdiana Ahrens, 1998 Microserica myagdiana Ahrens, 1998a: 46. Material examined. See Ahrens 1998a (p. 46). Aedeagus. See Ahrens 1998a (figs 40^12, p. 47) Distribution. Western central Nepal (Fig. 46F). Microserica pedongensis Ahrens, 1998 Microserica pedongensis Ahrens, 1998a: 50; Ahrens 2004b: 184, 2006a: 412. Material examined. See Ahrens 1998a (p. 50), 2006a (p. 412). Aedeagus. See Ahrens 1998a (figs 43^15, p. 47). Distribution. Darjeeling/Sikkim area (Fig. 46E). Microserica pruinosa (Hope, 1831) (Figs 3IN, 46F) Serica pruinosa Hope, 1831: 24. Microserica pruinosa. Ahrens 1998a: 39; 2004b: 183, 2006a: 412; Shrestha et al. 2012: 381. Omaloplia subaenea Blanchard, 1850: 79, syn. by Ahrens 1998a: 39. Material examined. See Ahrens 1998a (p. 39), 2004b (p. 183), 2006a (p. 412); Shrestha et al. 2012 (p. 381); 1 ex. “India, Uttarakhand, Govind Ghat [Gobindghat], valley of Alaknada River, N30°62 E079°55,14-15.06.2011, A. An¬ ishchenko & A. Shavrin” (CDUL), 2 ex. “India, Uttarak¬ hand, 2 km N Govind Ghat [Gobindghat], (Josimath- Badrinath pass), valley of Alaknada River, 12-13.06.2011, A. Shavrin & A. Anishchenko” (CDUL). Aedeagus. See Ahrens 1998a (figs 26-30, p. 42). Distribution. Himalaya, from east of the Sutlej river to eastern Nepal (Fig. 46F). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 133 Microserica roingensis sp. n. (Figs 10X-Z, 310, 46E) Type material examined. Holotype: $ “X-DA3458 - In¬ dia, Arunachal Pradesh, Roing, 490m, 28°08’32 ,, N 95°51’E, 2.-5.vi.2012, leg. L. Dembicky” (ZFMK). Description. Length: 5.5 mm, length of elytra: 4.1 mm, width: 3.4 mm. Body oval, yellow, frons dark brown, dor¬ sal surface dull and nearly glabrous. Labroclypeus slightly wider than long, shiny, widest at base, lateral margins strongly convex and convergent to rounded anterior angles; anterior margin weakly sinuate medially, margins weakly reflexed; lateral margins and oc¬ ular canthus produce a blunt angle; surface weakly con¬ vex medially, coarsely and densely punctate, with numer¬ ous long, erect setae; frontoclypeal finely incised and curved medially. Frons dull, with sparse, coarse punctures, beside eyes and behind frontoclypeal suture with a few single short setae, otherwise with minute setae in punc¬ tures. Smooth area in front of eyes twice as wide as long. Ocular canthus impunctate, narrow and moderately long, with a terminal seta. Eyes moderately large, ratio of di¬ ameter/interocular width: 0.68. Antenna with ten anten- nomeres; club composed of four antennomeres, straight, slightly longer than remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum wide, widest at base, lateral margins nearly evenly convex and convergent to blunt and moderately produced anterior angles. Anterior margin of pronotum weakly convex, with fine and complete marginal line; basal marginal line absent; posterior angles blunt; surface finely and densely punctate, glabrous, with minute setae in punctures, lateral and lateral anterior margins sparsely setose. Hypomeron carinate, weakly produced ventrally. Scutellum triangular, finely and densely punctate, on base widely impunctate. Elytra short, widest at middle, striae distinctly im¬ pressed, finely and densely punctate, intervals convex, with fine and sparse punctures concentrated along striae, with very minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at slight¬ ly rounded external apical angle of elytra, epipleura dense¬ ly setose, apical border chitinous, without a fringe of short microtrichomes (visible at ca lOOx magnification). Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.74. Abdominal sternites finely punctate, with a trans¬ verse row of coarse punctures, each bearing a robust se¬ ta. Pygidium strongly convex and dull, with mixed finer and larger punctures, without smooth midline, with numer¬ ous short setae and a few longer in the larger punctures. Legs moderately wide; femora finely and sparsely punc¬ tate, with two longitudinal rows of longer setae. Metafe¬ mur moderately shiny, anterior margin acute, without sub¬ marginal serrated line; anterior row of seta-bearing punc¬ tures present; posterior margin finely serrate ventrally, on¬ ly weakly widened in apical half, posterior margin serrate dorsally, with a few short setae basally. Metatibia moder¬ ately long and wide, widest at middle, ratio width/length: 1/2.85; dorsal margin sharply carinate, with two groups of spines, basal group at middle, apical one at three quar¬ ter of metatibial length, in basal third with a few robust single setae; lateral face longitudinally convex, finely and sparsely punctate, along midline smooth, with minute se¬ tae in punctures; ventral edge finely serrate, with three ro¬ bust equidistant setae; medial face smooth, apex interior¬ ly near tarsal articulation sharply truncate. Tarsomeres with fine, sparse setae ventrally, neither laterally nor dor- sally carinate, dorsally smooth; metatarsomeres glabrous, with a strongly serrated ridge ventrally and a parallel sub- ventral smooth carina immediately beside it; first metatar- somere distinctly as long as following two tarsomeres combined, twice as long as dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 10X-Z. Habitus: Fig. 310. Female un¬ known. Diagnosis. The new species belongs to the Microserica varians group. It differs from the similar species M. do- hertyi Ahrens & Fabrizi, 2009 and M. varians Moser, 1915 by the phallobase that has ventrally on each side an ele¬ vated carina. In the other species of this group the elevat¬ ed carina is present only on the left side. Etymology. The new species in named after its type lo¬ cality, Roing (adjective in the nominative singular). Distribution. India, Arunachal Pradesh (Fig. 46E). Microserica schawalleri Ahrens, 1998 Microserica schawalleri Ahrens, 1998a: 43; Ahrens 2004b: 182. Material examined. See Ahrens 1998a (p. 43), Ahrens 2004b (p. 182). Aedeagus. See Ahrens 1998a (figs 34-36, p. 44). Distribution. Western central Nepal (Fig. 46E). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 134 Dirk Ahrens & Silvia Fabrizi Microserica schulzei Ahrens, 1998 Microserica schulzei Ahrens, 1998a: 32; Ahrens 2004b: 181, 2006a: 412; Ahrens & Fabrizi 2011: 163; Shrest- ha et al. 2012: 381. Material examined. See Ahrens 1998a (p. 32), 2004b (p. 181), 2006a (p. 412); Ahrens & Fabrizi 2011 (p. 163); Shrestha et al. 2012 (p. 381). Aedeagus. See Ahrens 1998a (figs 5-7, p. 33). Distribution. Western central Nepal (Fig. 46F). Microserica steelei Ahrens, 2004 Microserica steelei Ahrens, 2004b: 186. Material examined. See Ahrens 2004b (p. 186). Aedeagus. See Ahrens 2004b (figs 284-286, p. 424). Distribution. SE Tibet and the Mishmi Hills (Fig. 46E). Microserica truncata (Brenske, 1898) Autoserica truncata Brenske, 1898: 308. Serica truncata : Barlow 1899: 246. Microserica truncata. Moser, 1918a: 219; Ahrens 1998a: 35, 2004b: 181; Ahrens & Fabrizi 2009b: 273. Microserica cribriceps Moser, 1915a: 183, syn. by Ahrens 1998a: 35. Material examined. See Ahrens 1998a (p. 35), 2004 (p. 181); Ahrens & Fabrizi 2009b (p. 273). Aedeagus. See Ahrens 1998a (figs 11-13, p. 36). Distribution. Sikkim/Darjeeling area (Fig. 46F). Microserica viridicollis Arrow, 1913 (Figs 3IP, 46E) Microserica viridicollis Arrow, 1913: 192; Ahrens 2002b: 386, 2004b: 188. Material examined. See Ahrens 2002b (p. 386); 27 SS, 25 $ p “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E , 500-950m, L. Dembicky leg., 29.iv.-2.v.2005” (CPPB), 34 ) 1 Apex of parameres convex.2 1 ’ Apex of parameres deeply concave. . M. xanthoptera sp. n. 2 Parameres longer and curved externally. . M. assamica (Moser) T Parameres short and straight. ..M. satrapa (Brenske) Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 137 Maladera assamica (Moser, 1915) Antoserica assamica Mosqy, 1915a: 160. Maladera assamica : Ahrens 2004b: 205; Krajcik 2012: 153. Autoserica ornatipennis Moser, 1915a: 161; syn. by Ahrens 2004b: 205; Maladera ornatipennis : Krajcik 2012: 155. Cephaloserica schereri Frey, 1962b: 613; syn. by Ahrens 2004b: 205; Maladera schererr. Krajcik 2012: 155. Material examined. See Ahrens 2004b (p. 205); 60 ex. “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E, 500-950m, Dembicky leg., 29.iv.-2.v.2005” (CPPB), 11 ex. “NE India, Meghalaya, SW of Cherrapun¬ jee, 25°13‘-14‘ N, 91°40‘ E , 5.-24.V.2005, 900m, P.Pa- cholatko leg.” (CPPB), 8 SS, 4 “NE India, Megha¬ laya, 8km N of Shillong, 1200m, 25°38‘N, 91°54‘E, 1200m, L. Dembicky leg., 7.- 9.V.2004” (CPPB). Aedeagus. See Ahrens 2004b (figs 305-307, p. 428). Distribution. Assam and Meghalaya (India) (Fig. 47B). Maladera satrap a (Brenske, 1898) (Figs 11A-C, 32C, 47B) Autoserica satrapa Brenske, 1898: 341. Maladera satrapa. Krajcik 2012: 155. Type material examined. Lectotype (here designated): S “Khasia Hills 50007 Ex. Museo H. W. Bates 1892/ satrapa type Brslc./ Mus. Paris ex. Coll. R. Oberthiir/ Type” (MNHN). Paralectotypes: 1 S “Naga Hills M. Cal¬ cutta/ satrapa m. (tridens olim./ Coll. Brenske” (ZMHB), 1 $ “Naga Hills M. Calcutta/ satrapa Brsk.” (ZMHB), 1 $ “Khasia Hills 4-50007 Ex. Museo H. W. Bates 1892/ satrapa type Brsk./ Mus. Paris ex. Coll. R. Oberthiir/ Type” (MNHN). Additional material examined. India: 35 ex. “NE India Meghalaya state, Jaintia Hills reg., Jowai 6.-8.VI.1996 alt. 1350 m± 100 m GPS N25°27‘ E92°12‘, (WGS 84) E. Jen- dek & O. Sausa/ IS 23” (CPPB), 39 ex. “NE India Assam, 1999, 5 km N of Umrongso, 700 m, 25°27‘N 92°43‘E 17.- 25.V. Dembicky & Pacholatko leg.” (CPPB), 136 ex. “NE India Meghalaya, 1999,9 km NW Jowai, 1400 m 25°30‘N 92°10‘E 12.V. Dembicky & Pacholatko leg.” (CPPB), 2 ex. “NE India Meghalaya 1999, 3 km E Tura 1150 m, 25°30‘N 90°14‘E, 18.IV. Dembicky & Pacholatko leg.” (CPPB), 1 ex. “India: Meghalaya state E Khasi Hills, 11km SW Cherrapunjee, Laitkynsew, 21-24.iv.2008, 25° 13’N, 91° 39’E, 810m,Fikacek,Podalska, Sipeklgt” (ZFMK), 3 “E India, Meghalaya, 2002, 3 km E Tu¬ ra, 1150 m, 25°30’N, 90°14 , E, 6.-12.V., M. Tryzna & P. Benda lgt.” (CPPB), 1 9 “E India, Meghalaya, 2002, 3 km E Tura, 1150 m, 25°30’N, 90°14’E, 2.-5.v., M. Tryzna & P. Benda lgt.” (CPPB), 1 ex. ($) “Assam Nagas/ Do¬ herty/ Fry Coll. 1905.105” (BMNH), 1 ex. “Shillong A. G. R. coll. 9.IV.18/ India T. B. Fletcher B.M. 1943-9.” (BMNH), 1 ex. ($) “India Or. Manipur/ Doherty/ Fry Coll. 1905.105” (BMNH), 1 ex. “Gauhati APL. 1920, Fletch¬ er Coll. / India T. B. Fletcher B.M. 1943-9.” (BMNH), 2 ex. “Umtyngar 16.5.Cherrapunjee/ Megalaya 1976 Wittmer, Baroni U.” (NHMB), 5 ex. “Assam, Khasi-Hills, Cherrapunji ca. 1300 m, V.1961 leg. G. Scherer/ Cephaloserica mulmeina Br. det. G. Frey 1962” (CF), 4 ex. “Barapani Old Road 1000 m 14.5./ Megalaya 1976 Wittmer, Baroni U ” (NHMB), 7 ex. “Shillong, Assam, In¬ dia 6000 ft. IV-20-28/ L.B. Parker Collector” (USNM), 8 ex. “Cherrapunji India VI-II-29 4300 ft./ L.B. Parker Col¬ lector” (USNM). Myanmar: 2 ex. “NW Myanmar (Bur¬ ma) Somura, Naga Chan; 03.V.2005; leg. Abe Azuma” (ZFMK). Redescription. Length: 5.7-6.4 mm, elytral length: 4.1^43 mm, width: 3.8—4.1 mm. Body short oval; bi¬ coloured, ventral side, head, margins of elytra black, legs, antenna, elytra and partly pronotum yellowish brown, dark parts sometimes with greenish shine; dorsal surface ex¬ cept head, tibiae, and tarsi dull, nearly glabrous. Labroclypeus wide, weakly trapezoidal, widest at base, lateral margins strongly convex and convergent anterior¬ ly, separated by a distinct angle from ocular canthus, slightly incised at transition to labrum; anterior angles moderately convex, slightly elevated; anterior margin slightly sinuate medially, margins weakly reflexed; sur¬ face weakly convex medially, finely densely punctate, with numerous larger punctures each bearing a erect seta. Fron- toclypeal suture finely incised, weakly curved. Smooth area in front of eyes as wide as long, strongly punctate near eyes. Ocular canthus short and wide, externally convex, very finely and densely punctate, with a long terminal se¬ ta. Frons dull, finely and moderately densely punctate, posterior head, beside eyes as well as on posterior quar¬ ter very densely and coarsely punctate, beside eyes with a few long erect setae, otherwise with microscopic setae in punctures only. Eyes small, ratio diameter/interocular width: 0.4. Antenna with nine antennomeres, club with three antennomeres, as long as remaining antennomeres combined and straight. Mentum convexly elevated, ante¬ riorly slightly flattened. Pronotum widest at base, anterior angles moderately sharp and slightly produced, posterior angles strongly con¬ vex, obsolete; anterior margin with fine and complete mar¬ ginal line, weakly convexly produced medially; lateral margins strongly convex and convergent anteriorly, im¬ mediately beside anterior margins convex; surface finely and densely punctate, with microscopic setae in punctures, otherwise glabrous, anterior and lateral margins with sparse but robust setae. Colour black, in basal half more or less widely yellowish, basal and lateral margins always Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 138 Dirk Ahrens & Silvia Fabrizi dark. Scutellum dark or yellowish, narrowly triangular, with fine and moderately dense punctures, with micro¬ scopic setae in punctures. Elytra wide, wides shortly behind middle, external api¬ cal angle widely rounded, yellowish brown, base, sutur¬ al interval, apical margin and external intervals dark, stri¬ ae finely impressed, finely sparsely punctate, intervals very weakly convex or flat, sparse and fine punctures con¬ centrated along striae, odd intervals with a few single short setae, near humerus with a single long seta, otherwise on¬ ly with microscopic setae in punctures. Epipleural edge robust, ending at external apical angle of elytra; apical margin chitinous, without rim of microtrichomes (visible at ca lOOx magnification). Ventral face with coarse and moderately dense punc¬ tures, sparsely setose, with a few robust setae on mesosternum and metastemal disc; otherwise with micro¬ scopic setae. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Metacoxa glabrous except a few robust setae laterally. Abdominal sternites dull, finely densely punctate, with a row of robust punctures each bearing a robust seta. Ratio of length of metepistemum/ metacoxa: 1/1.87. Pygidium dull, in male mostly dark, weakly convex ad moderately dense punctate, apical mar¬ gin with a few single long setae, otherwise only with mi¬ croscopic setae. Legs moderately wide; femora finely and very sparse¬ ly punctate, with two longitudinal rows of setae. Metafe¬ mur moderately shiny, anterior edge acute, without adja¬ cent serrated line; posterior ventral margin almost straight, distinctly widened in apical half, neither ventrally nor dor- sally serrated but smooth, glabrous. Metatibia short and moderately wide, sides subparallel, basally narrowed, ra¬ tio width/length: 1/3.2; dorsal margin weakly carinate, with two groups of spines, basal one at one third, apical one at three quarters of metatibial length; lateral face strongly longitudinally convex, smooth, laterally with sparse and superficial punctures; ventral margin with three nearly equidistant robust setae; medial face smooth and glabrous, interiorly near tarsal articulation shallowly con- cavely sinuate. Tarsi finely and sparsely punctate dorsal- ly, circular in cross section, metatarsomeres ventrally with a strongly serrated carina, subventrally with a second, smooth longitudinal carina, first metatarsomere one as long as following two combined and one quarter of its length longer than the dorsal tibial spur. Protibia moder¬ ately long, bidentate; protarsal claws normal, symmetric. Aedeagus: Fig. 11A-C. Habitus: Fig. 32C. Variation. Female: Antennal club with three anten- nomeres, shorter than remaining antennomeres combined; metatibia wider, ratio width/length: 1/2.78; abdominal ster¬ nites including flatter pygidium as well as the posterior legs being more yellowish brown. Remarks. The species is morphologically very similar to Maladera rufodorsata (Fairmaire, 1888) that is distributed from China (Kiangsi, Yunnan) to Myanmar, including N- Laos and northern Thailand but has been so far not yet recorded from India. Both species can be distinguished on¬ ly in shape of parameres. Distribution. See map (Fig. 47B). Maladera xanthoptera sp. n. (Figs 11D-F, 32D, 47B) Type material examined. Holotype: S “NE India, Meghalaya state, West Garo Hills reg., Tura 5.-7.V. 1996 alt. 700+100 m, GPS N25°30.7’ E90 o 13.9 , (WGS 84) E. Jendek & O. Sausa leg./ IS 18/ 608 Sericini Asia spec.” (CPPB). Paratypes: 1 $ “NE India, Meghalaya state, West Garo Hills reg., Tura 5.-7.V.1996 alt. 700+100 m, GPS N25°30.7’ E90°13.9’ (WGS 84) E. Jendek & O. Sausa leg./ IS 18/608 Sericini Asia spec.” (CPPB), 1 S “NE In¬ dia; Meghalaya; 1999; 3 km E of Tura; 1150 m; 25°30’N 90°14’E; 18.iv. Dembicky & Pacholatko leg.” (ZFMK). Description. Length: 5.8 mm, length of elytra: 3.5 mm, width: 3.5 mm. Body oval, yellowish brown, head, club of antenna, anterior portion and disc of pronotum, mar¬ gins of elytra and ventral surface dark brown to black, dark parts of pronotum and head with greenish shine, dorsal surface dull, except some single setae on head dorsal sur¬ face nearly glabrous. Labroclypeus wide and trapezoidal, widest at base, lat¬ eral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin distinct¬ ly sinuate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface slightly el¬ evated medially, finely and moderately densely punctate, with a few coarse punctures each bearing an erect seta, base of labroclypeus broadly covered with dull toment as the rest of the dorsal surface; frontoclypeal suture finely incised, medially angled; smooth area anterior to eye flat, twice as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons with fine, sparse punctures and a few single erect setae beside eyes. Eyes small, ratio di¬ ameter/interocular width: 0.42. Antenna with nine anten¬ nomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum el¬ evated and slightly flattened anteriorly. Pronotum transverse, widest in posterior third, lateral margins strongly convex, distinctly convergent anterior¬ ly, weakly convergent posteriorly, anterior angles moder¬ ately produced and rectangular, posterior angles blunt, weakly rounded in the tip; anterior margin strongly con¬ vex, with fine and complete marginal line, base without Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 139 marginal line; surface densely and finely punctate, glabrous; lateral and anterior margin finely setose; hy- pomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures. Elytra widest at middle, striae distinctly impressed, fine¬ ly and sparsely punctate, intervals flat, with fine and mod¬ erately dense punctures concentrated along striae, with on¬ ly a few short, setae on external intervals; apex as well sutural interval and four external intervals black; epipleur- al edge robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra not membraneous, without visible rim of microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and moderately densely punctate, metastemum sparsely covered with fine, short or minute setae; metacoxa with a few longer setae later¬ ally. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.93. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few robust setae along apical margin. Legs wide and short, shiny; femora with two longitu¬ dinal rows of setae, finely and sparsely punctate. Metafe¬ mur with anterior margin acute, without serrated line be¬ hind anterior edge, anterior row of setae widely reduced, posterior margin smooth and widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/2.6, sharply carinate dorsally, with two groups of spines, basal group at one third, apical group at two thirds of metatibial length, with a few short robust setae basally; lateral face longitudinally convex, nearly smooth, finely and sparsely punctate on sides, glabrous; ventral margin finely serrate, with four robust equidistant setae; medial face smooth and glabrous; apex finely serrate, moderate¬ ly truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dor¬ sally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth sub- ventral longitudinal carina; first metatarsomere slightly longer than following two tarsomeres combined and sube¬ qual to dorsal tibial spur. Protibia short, bidentate; ante¬ rior claws symmetrical, basal tooth of both claws blunt¬ ly truncate at apex. Aedeagus: Fig. 11D-F. Habitus: Fig. 32D. Diagnosis. Maladera xanthoptera sp. n. differs from the most similar M. satrapa by the shape of the aedeagus: the apex of the parameres is deeply sinuate, in lateral view the parameres are much narrower in the new species. Etymology. The name (noun in apposition) of the new species is derived from the combined latinised Greek words ‘ xanthos ’ (yellow) and ‘ pteron ’ (wing), with refer¬ ence to the yellow elytra. Variation. Length: 5.8-6.1 mm, length of elytra: 3.5—4.1 mm, width: 3.5-4.1 mm. Female: Body slightly larger, head and pronotum reddish brown. Antennal club with three antennomeres and straight, distinctly shorter than the remaining antennomeres combined. Distribution. See map (Fig. 47B). Maladera egregia group Key to species of the Maladera egregia group {1/3.1.6 4 Parameres moderately long, subequal to the slightly angled apophysis of phallobase. M. rolciki Ahrens 4’ Parameres longer, distinctly longer than the apophysis of phallobase being curved dorsally.5 5 Apophysis of phallobase dorsolateral. Fused parameres together moderately narrow (dorsal view). . M. mussooriensis Ahrens 5’ Apophysis of phallobase narrower and entirely dis¬ placed laterally. Fused parameres together distinctly narrower (dorsal view). M. hunliensis sp. n. 6 Parameres dorsoventrally flattened, constricted tip of parameres short. M. impubis Ahrens 6’ Parameres not dorsoventrally flattened, constricted tip of parameres long.7 7 Base of parameres narrow. Right part of phallobasal apophysis ventrolaterally displaced and slightly curved interiorly. M. irididorsis Ahrens 7’ Base of parameres wide. Right part of phallobasal apophysis dorsolaterally, not curved. . M. kazirangae Ahrens Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 140 Dirk Ahrens & Silvia Fabrizi Maladera hmong Ahrens, 2004 Maladera hmong Ahrens, 2004b: 331. Material examined. See Ahrens 2004b (p. 331). Aedeagus. See Ahrens 2004 (figs 530-535, p. 462). Distribution. Mountains of northern Indochina and east¬ ern Nepal. So far not recorded from India (Fig. 47C). Maladera hunliensis sp. n. (Figs 11G-I, 32E, 47C) Type material examined. Holotype S “NE India Arunachal Pr. Hunli vicinity, 1300±100m 28°19 , 32”N 95°57 , 31 ,, E, L. Dembicky leg., 26.v.-l.vi.2012 ,, (ZFMK). Paratype: 1 $ “NE India Arunachal Pr. Hunli vicinity, 1300±100m 28°19 , 32”N 95°57 , 31 ,, E, L. Dembicky leg., 26.v.-l.vi.2012” (ZFMK). Description. Length: 9.1 mm, length of elytra: 6.9 mm, width: 5.8 mm. Body oval, dark reddish brown, shiny, ex¬ cept some single setae on head dorsal surface nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins straight and convergent an¬ teriorly, anterior angles strongly rounded, anterior margin nearly straight, margins moderately reflexed; lateral mar¬ gin and ocular canthus produce a blunt angle; surface slightly elevated medially, coarsely and densely punctate, a few punctures anteriorly bear an erect seta; frontoclypeal suture indistinctly incised, medially angled; smooth area anterior to eye weakly convex, 2.5 times as wide as long; ocular canthus short and wide (one quarter of ocular di¬ ameter), finely and densely punctate, with a terminal se¬ ta. Frons with fine, irregular punctures and two erect se¬ tae beside eyes. Eyes moderately large, ratio diameter/in¬ terocular width: 0.62. Antenna with ten antennomeres; club with three antennomeres and straight, slightly short¬ er than remaining antennomeres combined. Mentum ele¬ vated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins straight and distinctly convergent anteriorly, in ante¬ rior third slightly convex, anterior angles moderately pro¬ duced and rectangular, posterior angles blunt, weakly rounded in the tip; anterior margin weakly convex, with fine and complete marginal line, base without marginal line; surface densely and finely punctate, glabrous; later¬ al margin finely setose, anterior margin glabrous; hy- pomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures, apex smooth. Elytra widest at posterior third, striae distinctly im¬ pressed, finely and sparsely punctate, intervals flat, with fine and evenly dense punctures, with only a few short, nearly entirely glabrous, even minute setae not visible; epi- pleural edge robust, ending at strongly curved external api¬ cal angle of elytra, epipleura densely setose; apical bor¬ der of elytra narrowly membraneous, with a fine rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, metasternum sparsely covered with fine, short or minute setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a fine short seta. Mesosternum between mesocox- ae as wide as mesofemur. Ratio of length of metepister- num/metacoxa: 1/1.6. Pygidium weakly convex and shiny, coarsely and densely punctate, with narrow smooth midline, glabrous except a few robust setae along apical margin. Legs slender and moderately long, shiny; femora with two longitudinal rows of setae, finely and sparsely punc¬ tate. Metafemur with anterior margin acute, with contin¬ uously serrated line behind anterior edge, anterior row of setae complete; posterior margin smooth, widened and finely serrate at apex ventrally, not serrate dorsally, glabrous. Metatibia moderately long, widest at apex, ra¬ tio of width/length: 1/3.2, sharply carinate dorsally, with two groups of spines, basal group at half, apical group at three quarters of metatibial length, with a few short ro¬ bust setae and a short serrated line parallel to the dorsal margin basally; lateral face longitudinally convex, near¬ ly smooth, finely and moderately densely punctate on sides, in apical half punctures sparse, glabrous; ventral margin finely serrate, with the apical of the five robust se¬ tae more distant from the others; medial face smooth and glabrous; apex finely serrate, moderately truncate interi¬ orly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, dense¬ ly setose ventrally; metatarsomeres with a strongly ser¬ rated ridge ventrally and a smooth subventral longitudi¬ nal carina; first metatarsomere distinctly shorter than fol¬ lowing two tarsomeres combined and slightly longer than the dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11G-I. Habitus: Fig. 32E. Female un¬ known. Diagnosis. Maladera hunliensis sp. n. is in its shape of the aedeagus very similar to Maladera mussooriensis Ahrens, 2004, but the lateral apophysis of its phallobase is narrower, being entirely displaced laterally, and the fused parameres are combined distinctly narrower in dor¬ sal view. Etymology. The new species is named with reference to its type locality, Hunli (adjective in the nominative sin¬ gular). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 141 Variation. Length: 7.6-9.1 mm, length of elytra: 6.1-6.9 mm, width: 4.9-5.8 mm. Distribution. See map (Fig. 47C). Maladera impubis Ahrens, 2004 (Figs 32F, 47C) Maladera impubis Ahrens, 2004b: 324. Material examined. See Ahrens 2004b (p. 324); 1 S, 1 9' “NE India Assam, 2002, Umrongso vill. env. 700 m, 25°27’E, 92°43’E, 3.-8.V., M. Tryzna & P. Benda lgt.” (CPPB), 26 6 $ $ “E India, Meghalaya, 2002, 3 km E Tura, 1150 m, 25°30’N, 90°14’E, 6.-12.V., M. Tryzna & P. Benda lgt” (CPPB), 8 SS, 1 9 “E India, Megha¬ laya, 2002, 3 km E Tura, 1150 m, 25°30’N, 90°14 , E, 2.- 5.v., M. Tryzna & P. Benda lgt.” (CPPB), 1 $ “India: Meg¬ halaya state E Khasi Hills, 11km SW Cherrapunjee, Lait- kynsew, 21-24.iv.2008,25° 13’N, 91° 39’E, 810m, Fika- cek, Podalska, Sipek lgt.” (ZFMK). Aedeagus. See Ahrens 2004b (figs 513-515, p. 459). Distribution. Sikkim/Darjeeling Himalaya, Meghalaya (Fig. 47C). Maladera irididorsis Ahrens, 2004 Maladera irididorsis Ahrens, 2004b: 327. Material examined. See Ahrens 2004b (p. 327). Aedeagus. See Ahrens 2004b (figs 519-522, p. 460). Distribution. Known so far from Assam [record of the holotype] only (Fig. 47C). Maladera kazirangae Ahrens, 2004 Maladera kazirangae Ahrens, 2004b: 326. Material examined. See Ahrens 2004b (p. 326). Aedeagus. See Ahrens 2004b (figs 516-518, p. 460). Distribution. Known so far from Assam [record of the holotype] only (Fig. 47C). Maladera mussooriensis Ahrens, 2004 Maladera mussooriensis Ahrens, 2004b: 328 Material examined. See Ahrens 2004b (p. 328). Aedeagus. See Ahrens 2004b (figs 523-526, p. 461). Distribution. So far only known from the type locality (Mussorie - Dehradun District; Uttarakhand state) at the foot of the Indian NW Himalaya (Fig. 47C). Maladera nigrolucida sp. n. (Figs 11J-L, 32G, 47C) Type material examined. Holotype: S “S-India, Kerala state, Kallar env., 30 km. NE of Trivandrum, vallay of riv. Kallar, 77°05’E 8°45’N, ca 300-500 m, 7-13.V.1999 Z. Ke- jval & M. Tryzna leg/ 652 Sericini Asia spec.” (CPPB). Paratypes: 2 SS, 2 99 “S-India, Kerala state, Kallar env., 30 km. NE of Trivandrum, vallay of riv. Kallar, 77°05’E 8°45’N, ca 300-500 m, 7-13.V.1999 Z. Kejval & M. Tryz¬ na leg/ 652 Sericini Asia spec.” (CPPB, ZFMK). Description. Length: 9.2 mm, elytral length: 6.4 mm, width: 5.6 mm. Body oval, uniformly blackish brown, dor¬ sal and ventral face shiny, except lateral setae of elytra and pronotum nearly glabrous. Labroclypeus wide, trapezoidal, widest at base, lateral margins moderately convex and convergent anteriorly, producing an indistinct blunt angle with ocular canthus, not incised towards labrum, anterior angles convex, an¬ terior margin distinctly sinuate medially, margins weak¬ ly reflexed; surface flat, finely and moderately densely punctate. Frontoclypeal suture finely incised, weakly curved. Smooth area in front of eyes twice as wide as long; ocular canthus short and wide, finely and densely punc¬ tate, with a short terminal seta. Frons finely and sparsely punctate, with a few single and short setae beside eyes. Eyes small, ratio diameter/interocular width: 0.54. Anten¬ na with ten antennomeres, club with three antennomeres, as long as remaining antennomeres combined. Mentum convexly elevated, anteriorly slightly flattened. Pronotum widest at base, lateral margins evenly and convexly convergent anteriorly, anterior angles moderate¬ ly sharp, distinctly produced, posterior angles blunt, slight¬ ly rounded at tip; anterior margin with complete margin¬ al line, weakly convexly produced medially; lateral and lateral anterior margin with long and fine setae; surface finely and densely punctate, with microscopic setae in punctures, otherwise glabrous. Scutellum wide, triangu¬ lar, punctures fine and moderately dense, glabrous. Elytra wide, widest shortly behind middle, external api¬ cal angle strongly rounded, striae finely impressed, fine¬ ly and densely punctate, intervals weakly convex, finely and moderately densely punctate, except second one with punctures concentrated along striae and with a few sin¬ gle short setae, otherwise with only microscopic setae in punctures; epipleural edge ending at external apical an¬ gle of elytra; epipleura with long and sparse setae; apical margin with a rim of fine microtrichomes. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 142 Dirk Ahrens & Silvia Fabrizi Ventral surface coarsely and densely punctate, with mi¬ croscopic setae in punctures, with a few longer setae on mesosternum and metasternal plate. Mesosternum be¬ tween mesocoxae 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.78. Metacoxa glabrous, laterally with a few robust setae. Abdominal ster- nites finely and moderately densely punctate, with a trans¬ verse row of coarse punctures each bearing a robust seta, penultimate sternite with a very narrow shiny chitinous rim. Pygidium shiny, weakly convex, coarsely and dense¬ ly punctate, without median impunctate line, with a few fine longer setae apically, otherwise only with microscop¬ ic setae only. Legs moderately wide; femora with two longitudinal rows of setae. Metafemur shiny, superficially punctate, an¬ terior edge acute, with adjacent serrated line, which is un¬ dulated and partly interrupted, anterior row of setae com¬ plete but its setae very short; posterior ventral margin al¬ most straight, weakly widened in apical half, neither ven- trally nor dorsally serrated but smooth, glabrous. Metat¬ ibia wide and short, widest at middle, ratio width/length: 1/2.2, sharply carinate dorsally, with two groups of spines, basal one shortly before middle, apical one at three quar¬ ters of metatibial length, in basal third with 3-4 coarse punctures each bearing a robust seta; lateral face weakly longitudinally convex, impunctate, only ventral part with moderately dense and coarse punctures; ventral margin with four equidistant spines; medial face impunctate and glabrous, apex shallowly sinuate interiorly near tarsal ar¬ ticulation. Tarsomeres impunctate dorsally, circular in cross section, with sparse, fine setae ventrally; metatar- someres ventrally with a strongly serrated carina, subven- trally with a second, smooth longitudinal carina; first metatarsomere distinctly shorter than following two tar¬ someres combined and little longer than dorsal tibial spur. Protibia moderately long, bidentate, teeth moderately large. All claws symmetrical, feebly curved and long, with normally developed basal tooth. Aedeagus: Fig. 11J-L. Habitus: Fig. 32G. Diagnosis. Due to the presence of a presumably motile dorsal lobe of the left paramere Maladera nigrolucida sp. n. resembles somewhat the species of the M. lugubris group. It differs from them by the body being complete¬ ly shiny and having a lamina-like dorsal process at the apex of the phallobase that covers the base of the para- meres. Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived from the combined Latin words ‘ nigris ’ (black) and 7 ucidus’ (shiny), with ref¬ erence to the black and shiny dorsal surface. Variation. Length: 9.2-9.7 mm, elytral length: 6.4-6.7 mm, width: 5.6-5.9 mm. Female: No apparent sexual di¬ morphism, only the antennal club slightly shorter than in male. Distribution. See map (Fig. 47C). Maladera rolciki Ahrens, 2004 Maladera rolciki Ahrens, 2004b: 330. Material examined. See Ahrens 2004b (p. 330); 2 88 “E India, Meghalaya, 2002, 3 km E Tura, 1150 m, 25°30’N, 90°14’E, 6.-12.V., M. Tryzna & P. Benda lgt.” (CPPB). Aedeagus. See Ahrens 2004b (figs 527-529, p. 461). Distribution. Meghalaya (Fig. 47C). Maladera ferruginea group Key to species of the Maladera ferruginea group ( 88 ) 1 Phallobase apically with a lateral apophysis on each side being nearly symmetric.2 1 ’ Phallobase apically without or only with an unilater¬ al apophysis.4 2 Parameres subequal in length.3 T Right paramere half as long as left. . M. namborensis sp. n. 3 Antennal club as long as remaining antennomeres combined. Parameres curved, not cleft before apex. . . M. sempiterna (Brenske) 3’ Antennal club 1.5 times as long as remaining anten¬ nomeres combined. Parameres straight, cleft before apex. M. sempiternella sp. n. 4 Phallobase with long lateral apophysis.5 4’ Phallobase without lateral apophysis.6 5 Lateral apophysis on left side. Antennal club 1.5 times as long as remaining antennomeres combined. . M. festina (Brenske) 5’ Lateral apophysis on right side. Antennal club slight¬ ly longer than remaining antennomeres combined. ... . M. paris Ahrens 6 Metatibia moderately long and wide, ratio width/length: 1/3; with a serrated line in basal two thirds of metatibial length. M. conspicua Ahrens 6’ Metatibia short and wide, ratio width/length: 1/2.5; with a serrated line in basal third of metatibial length. .7 7 Antennal club slightly longer than remaining anten¬ nomeres combined. Dorsoapical sinuation of phal¬ lobase between parameres deep and wide. . M. krali Ahrens 7’ Antennal club 1.5 times as long as remaining anten¬ nomeres combined.8 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 143 8 Dorsoapical sinuation of phallobase between para- meres deep and wide.9 8’ Dorsoapical sinuation of phallobase between para- meres shallow and narrow. .13 9 Right paramere distinctly shorter than the left one. 10 9’ Right paramere distinctly longer than the left one. 11 10 Basal lobe of right paramere short. Left paramere very wide at base, evenly narrowed towards apex. . M. chiruwae Ahrens 10’ Basal lobe of right paramere longer. Left paramere abruptly narrowed in basal third, remainder part long and narrow. . M. spei Ahrens 11 Basolateral extension of left paramere very small. Right paramere nearly twice as long as left one. . M. merkli Ahrens 11 ’ Basolateral extension of left paramere large to very large. Right paramere only 1.2 times of its length longer than left one.12 12 Basolateral extension of left paramere very large, the narrow distal lobe stands dorsally and is curved ven- trally. M. gorkhae Ahrens 12’ Basolateral extension of left paramere large, the nar¬ row distal lobe stands ventrally and is curved dorsal¬ ly. M. wolfgangdierli Ahrens 13 Left paramere with a separated narrow distal lobe in¬ serted ventrally being curved dorsally. Frons glabrous .14 13’ Left paramere without a distinctly separated distal lobe. Frons sparsely setose.15 14 Right paramere distinctly longer than left one. Basal extension of left paramere very large. . M. ferruginea (Kollar Sc Redtenbacher) 14’ Both parameres subequal in length. Basal extension of left paramere small. M. fatigata Ahrens 15 Metatibia shorter (ratio width/length: 1/2.9). Eyes smaller (ratio diameter/interocular distance: 0.68). Left paramere shorter. Phallobase ventroapically lamellous- ly produced. M. schereri (Frey) 15’ Metatibia longer (ratio width/length: 1/3.3). Eyes larg¬ er (ratio diameter/interocular distance: 0.74). Left para¬ mere longer. Phallobase ventroapically not lamellous- ly produced.M duvivieri (Brenske) Maladera chiruwae Ahrens, 2004 Maladera chiruwae Ahrens, 2004b: 300. Material examined. See Ahrens 2004b (p. 300). Aedeagus. See Ahrens 2004b (figs 465-468, p. 452). Distribution. Endemic to eastern Nepal (Fig. 47D). Maladera conspicua Ahrens, 2004 Maladera conspicua Ahrens, 2004b: 293. Material examined. See Ahrens 2004b (p. 293). Aedeagus. See Ahrens 2004b (figs 447-449, p. 450). Distribution. Endemic to Sikkim (Fig. 47D). Maladera duvivieri (Brenske, 1896) Serica duvivieri Brenske, 1896: 152. Maladera duvivieri. Ahrens 2004b: 303; Krajcik 2012: 154. Material examined. See Ahrens 2004b (p. 303). Aedeagus. See Ahrens 2004b (figs 474-476, p. 453). Distribution. Endemic to the lower Darjeeling area (Fig. 47D). Maladera fatigata Ahrens, 2004 Maladera fatigata Ahrens, 2004b: 292. Material examined. See Ahrens 2004b (p. 292). Aedeagus. See Ahrens 2004b (figs 444-446, p. 449). Distribution. Endemic to central Nepal (Fig. 47D). Maladera ferruginea (Kollar & Redtenbacher, 1844) (Figs 32H, 47D) Serica ferruginea Kollar Sc Redtenbacher, 1844: 525; Bar- low 1899: 243. Omaloplia ferruginea Blanchard, 1850: 81; syn. by Ahrens 2004b: 290 Serica punctifrons Dejean, 1837: 182; Gemminger Sc Harold 1869: 1120; Brenske 1898: 230; Dalla Torre 1912: 14 (nomen nudum). Maladera ferruginea. Ahrens 2004b: 290, 2006a: 413; Ahrens Sc Fabrizi 2009b: 274; Krajcik 2012: 154. Material examined. See Ahrens 2004b (p. 290), 2006a (p. 413); Ahrens Sc Fabrizi 2009b (p. 274); Sabatinelli Sc Ahrens 2015 (p. 141). Aedeagus. See Ahrens 2004b (figs 441-443, p. 449). Distribution. Western Himalaya (Pakistan) to western central Nepal (Fig. 47D). Bonn zoological Bulletin 65 (1 Sc 2): 1-355 ©ZFMK 144 Dirk Ahrens & Silvia Fabrizi Maladera festina (Brenske, 1898) Autoserica festina Brenske, 1898: 296. Serica festina. Barlow 1899: 243. Maladera festina. Ahrens 2004b: 305; Krajcik 2012: 154. Material examined. See Ahrens 2004b (p. 305). Aedeagus. See Ahrens 2004b (figs 480-482, p. 454). Distribution. Central Nepal and the Darjeeling district (Fig. 47D). Maladera gorkhae Ahrens, 2004 Maladera gorkhae Ahrens, 2004b: 297. Material examined. See Ahrens 2004b (p. 297). Aedeagus. See Ahrens 2004b (figs 458^160, p. 451). Distribution. Endemic to central Nepal (Fig. 47D). Maladera krali Ahrens, 2004 Maladera krali Ahrens, 2004b: 301. Material examined. See Ahrens 2004b (p. 301). Aedeagus. See Ahrens 2004b (figs 469,470,472,473, p. 453). Distribution. Endemic to Sikkim (without any precise da¬ ta), only known from holotype. Maladera merkli Ahrens, 2004 Maladera merkli Ahrens, 2004b: 296. Material examined. See Ahrens 2004b (p. 296). Aedeagus. See Ahrens 2004b (figs 454^157, p. 451). Distribution. Endemic to the western Himalaya (Fig. 47D). Maladera namborensis sp. n. (Figs 11M-0, 321, 47D) Type material examined. Holotype: S “N.E. India Nam- bor Reserv. Forest Assam, 26°00’N, 94°20’E Garampani 100 m 21.-29.11.97 lg. V. Sinaev & V.S. & M. Murzin/ 604 Sericini Asia spec.” (ZFMK). Paratype: 1 S “NE In¬ dia, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E , 5.-24.V.2005, 900m, P. Pacholatko leg.” (CPPB). Description. Length: 9.2 mm, length of elytra: 6.6 mm, width: 5.8 mm. Body oval, dark reddish brown, dull, labro- clypeus shiny, except some single setae on head dorsal sur¬ face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and con¬ vergent anteriorly, anterior angles strongly rounded, an¬ terior margin weakly sinuate medially, margins weakly re- flexed; lateral margin and ocular canthus produce an in¬ distinct blunt angle; surface flat, finely and densely punc¬ tate, glabrous; frontoclypeal suture distinctly incised, even¬ ly curved; smooth area anterior to eye weakly convex, 2.5 times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely and densely punctate, with a terminal seta. Frons with sparse, fine punctures and a sin¬ gle long seta beside each eye. Eyes large, ratio diameter/in¬ terocular width: 0.71. Antenna with ten antennomeres; club with three antennomeres and straight, as long as re¬ maining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins evenly convex and convergent anteriorly, anterior an¬ gles distinctly produced and sharp, posterior angles blunt, strongly rounded in the tip; anterior margin convex, with the marginal line widely lacking, base without marginal line; surface sparsely and finely punctate, glabrous; lat¬ eral margin finely setose, anterior margin glabrous; hy- pomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures, on median base impunctate. Elytra widest at posterior third, striae distinctly im¬ pressed, finely and sparsely punctate, intervals nearly flat, with fine and dense punctures mainly concentrated along striae, nearly entirely glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membra¬ neous, with a rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each stemite with a loose transverse row of punctures each bear¬ ing a fine seta. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/meta- coxa: 1/1.56. Pygidium weakly convex and dull, coarse¬ ly and densely punctate, without midline, glabrous except a few robust setae near apex. Legs short and wide, shiny; femora with two longitu¬ dinal rows of setae, finely and sparsely punctate. Metafe¬ mur with anterior margin acute, without adjacent serrat¬ ed line, anterior row of setae completely reduced; poste¬ rior margin smooth, widened and smooth at apex ventral¬ ly, not serrate dorsally, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.3, sharply car¬ inate dorsally, with two groups of spines, basal group at anterior third, apical group at three quarters of metatibial Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 145 length, glabrous but with a short serrated line parallel to the dorsal margin basally; lateral face longitudinally con¬ vex, shiny but basal third dull, impunctate and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely ser¬ rate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither later¬ ally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatar- somere nearly as long as following two tarsomeres com¬ bined and as long as dorsal tibial spur. Protibia moderate¬ ly long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11M-0. Habitus: Fig. 321. Female un¬ known. Diagnosis. Maladera namborensis sp. n. is in its external morphology very similar to M. sempiterna. The new species differs from the latter by the distinctly asymmet¬ ric parameres, the right paramere being half as long as the left one. From all other species of the M. ferruginea group M. namborensis can be distinguished by the lateral apophyses at each side of phallobase. Etymology. This new species is named after the Nambor forest reserve in Assam (adjective in the nominative sin¬ gular). Distribution. See map (Fig. 47D). Maladera paris Ahrens, 2004 Maladera paris Ahrens, 2004b: 307. Material examined. See Ahrens 2004b (p. 307). Aedeagus. See Ahrens 2004b (figs 483-485, p. 455). Distribution. Endemic to Sikkim (Fig. 47D). Maladera schereri (Frey, 1975) Autoserica schereri Frey, 1975a: 183. Maladera schereri : Ahrens 2004b: 304; Krajcik 2012: 155. Material examined. See Ahrens 2004b (p. 304). Aedeagus. See Ahrens 2004b (figs 477-479, p. 454). Distribution. Endemic to the lower Darjeeling area (Fig. 47D). Maladera sempiterna (Brenske, 1898) (Fig. 11P-R, 47D) Autoserica sempiterna Brenske, 1898: 328. Maladera sempiterna. Ahrens 2004b: 289; Krajcik 2012: 155. Type material examined. Syntypes: 2 <$<$ “Khasi Hills Mus. Calcutta/ sempiterna Type Brsk./ coll. Brenske” (ZMHB). Additional material examined. India. 1 $ “Naga Hills M. Calcutta/ sempiterna Type Brsk./ coll. Brenske” (ZMHB), 1 S “India: 26.VI.95 Cherrapunjee Meghalaya Werner leg.” (ZFMK), 1 ex. 0) “NE India Meghalaya state Khasi Hills Mawphlang vill. GPS N25°26.7 E 91°45.2‘ 2.-3.VI., 10.VI. 1996alt. 1700mE. Jendek&O. Sausa leg./ IS 93” (CPPB), 1 S “NE India, Megalaya, SW of Cherrapunjee 25°13-14’N 91°40’E, 5.-24.V.2005, 900m, P. Pacholatko leg.” (CPPB), 1 $ “NE India; Mega¬ laya, 2002, 1 km E of Tura, 500-600m, 25°30’N, 90°14 , E; 13.-18.V. M. Tryzna & P. Benda lgt ” (CPPB), 2 ex. “NE India, Meghalaya, 2002, 3 km E Tura, 1150 m, 25°30‘N, 90° 14 E, 6.-12.V., M. Tryzna & P. Benda lgt.” (CPPB), 3 ex. “Khasis July 1894 Nat. Coll./ Coll. C. Felsche, Kauf 20, 1918” (SMTD), 47 ex. “NE India W Meghalaya Garo Hills; Nokrek N. P. 25,40N 91,04E, 2.- 13.VII.1997 V. Sinaev leg., 1150 m” (CPPB), 2 ex. “As¬ sam/ Coll. C. Felsche, Kauf 20, 1918” (SMTD), 1 ex. “Shillong, Assam. F.W.C./ G.C. Champion Coll. B.M. 1927-409.” (BMNH). Thailand. 1 ex. 0) “Thai, Chiang Mai prov., 18°49’N 98°54’E, 1600 m, Doi Pui mt., 2.-6.V. Vit Kuban leg. 1996/Asia Sericini spec. 325” (CPPB). Redescription. Length: 9.8 mm, length of elytra: 6.4 mm, width: 5.6 mm. Body oval, dark reddish brown, dull, labro- clypeus shiny, except some single setae on head dorsal sur¬ face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and con¬ vergent anteriorly, anterior angles strongly rounded, an¬ terior margin weakly sinuate medially, margins weakly re- flexed; lateral margin and ocular canthus produce an in¬ distinct blunt angle; surface nearly flat, finely and dense¬ ly punctate, glabrous; frontoclypeal suture distinctly in¬ cised, medially angled; smooth area anterior to eye weak¬ ly convex, twice as wide as long; ocular canthus short and wide (one quarter of ocular diameter), finely and dense¬ ly punctate, with a terminal seta. Frons with sparse super¬ ficial punctures and a single long seta beside each eye. Eyes moderately large, ratio diameter/interocular width: 0.6. Antenna with ten antennomeres; club with three an- tennomeres and straight, as long as remaining anten¬ nomeres combined. Mentum elevated and slightly flat¬ tened anteriorly. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 146 Dirk Ahrens & Silvia Fabrizi Pronotum moderately wide, widest at base, lateral mar¬ gins evenly convex and convergent anteriorly, anterior an¬ gles distinctly produced and sharp, posterior angles blunt, weakly rounded in the tip; anterior margin convex, with the marginal line widely lacking, base without marginal line; surface sparsely and finely punctate, glabrous; lat¬ eral margin finely setose, anterior margin glabrous; hy- pomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures. Elytra widest at posterior third, striae indistinctly im¬ pressed, finely and sparsely punctate, intervals flat, with fine and evenly dense punctures, nearly entirely glabrous; epipleural edge robust, ending at strongly curved exter¬ nal apical angle of elytra, epipleura sparsely setose; api¬ cal border of elytra membraneous, with a rim of microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each stemite with a loose transverse row of punctures each bear¬ ing a fine seta. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/meta- coxa: 1/1.4. Pygidium weakly convex and dull, coarsely and densely punctate, without midline, glabrous except a few robust setae along apical margin. Legs short and wide, shiny; femora with two longitu¬ dinal rows of setae, finely and sparsely punctate. Metafe¬ mur with anterior margin acute, without adjacent serrat¬ ed line, anterior row of setae completely reduced; poste¬ rior margin smooth, widened and smooth at apex ventral¬ ly, not serrate dorsally, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.5, sharply car¬ inate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short robust setae and a short serrated line parallel to the dorsal margin basally; lateral face longitudinally convex, strongly shiny and impunctate, glabrous; ventral margin finely serrate, with four equidis¬ tant robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal ar¬ ticulation. Tarsomeres dorsally smooth and glabrous, nei¬ ther laterally nor dorsally carinate, moderately setose ven¬ trally; metatarsomeres with a strongly serrated ridge ven¬ trally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tar¬ someres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11P-R. Distribution. Northeastern India and northern Thailand (Fig. 47D). Maladera sempiternella sp. n. (Figs 11S-U, 32J, 47D) Type material examined. Holotype: S “Myanmar N (Burma) 65 km NE Putao, 1250 m, Zi Yar Dam vill. 18- 21.05.1998 leg. S. Murzin & V. Sinaev/ [ex] Coll. Takeshi Itoh, Osaka (Japan)/ 706 Sericini: Asia spec.” (ZFMK). Description. Length: 8.8 mm, length of elytra: 6.4 mm, width: 5.3 mm. Body oval, dark reddish brown, dull, labro- clypeus shiny, except some single setae on head dorsal sur¬ face nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins weakly convex and con¬ vergent anteriorly, anterior angles strongly rounded, an¬ terior margin weakly sinuate medially, margins weakly re¬ flexed; lateral margin and ocular canthus produce an in¬ distinct blunt angle; surface flat, finely and densely punc¬ tate, glabrous; frontoclypeal suture distinctly incised, even¬ ly curved; smooth area anterior to eye weakly convex, 2.5 times as wide as long; ocular canthus short and wide (1/3 of ocular diameter), finely and densely punctate, with a terminal seta. Frons with sparse, fine punctures and a sin¬ gle long seta beside each eye. Eyes large, ratio diameter/in¬ terocular width: 0.78. Antenna with ten antennomeres; club with three antennomeres and straight, 1.5 times as long as remaining antennomeres combined. Mentum el¬ evated and slightly flattened anteriorly. Pronotum moderately wide, widest shortly before base, lateral margins evenly convex and convergent anteriorly, slightly convexly narrowed towards base, anterior angles distinctly produced and sharp, posterior angles blunt, strongly rounded in the tip; anterior margin convex, with the marginal line widely lacking, base without marginal line; surface sparsely and finely punctate, glabrous; lat¬ eral margin finely setose, anterior margin glabrous; hy- pomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, dense punctures. Elytra widest at posterior third, striae distinctly im¬ pressed, finely and sparsely punctate, intervals nearly flat, with fine and evenly dense punctures, nearly entirely glabrous; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra membraneous, with a rim of mi- crotrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each stemite with a loose transverse row of punctures each bear¬ ing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/meta- coxa: 1/1.49. Pygidium weakly convex and dull, coarse¬ ly and densely punctate, without midline, glabrous except a few robust setae along apical margin. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 147 Legs short and wide, shiny; femora with two longitu¬ dinal rows of setae, finely and sparsely punctate. Metafe¬ mur with anterior margin acute, without adjacent serrat¬ ed line, anterior row of setae completely reduced; poste¬ rior margin smooth, widened and smooth at apex ventral - ly, not serrate dorsally, glabrous. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.7, sharply car¬ inate dorsally, with two groups of spines, basal group shortly before middle, apical group at three quarters of metatibial length, with a few short robust setae and a short serrated line parallel to the dorsal margin basally; lateral face longitudinally convex, shiny but basal third dull, im- punctate and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interi¬ orly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, mod¬ erately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitu¬ dinal carina; first metatarsomere as long as following two tarsomeres combined and slightly longer than dorsal tib- ial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 11S-U. Habitus: Fig. 32J. Diagnosis. Maladera sempiternella sp. n. is in the shape of the genitalia and its external morphology very similar to M. sempiterna. The new species differs from M. sem- piterna by the longer antennal club as well as by the straight parameres that are cleft before the apex. Distribution. So far known only from northern Myanmar, in close vicinity to the Indian border (Fig. 47D). Maladera spei Ahrens, 2004 Maladera spei Ahrens, 2004b: 299. Material examined. See Ahrens 2004b (p. 299). Aedeagus. See Ahrens 2004b (figs 461-464, p. 452). Distribution. Endemic to eastern Nepal and the Darjee¬ ling area (Fig. 47D). Maladera wolfgangdierli Ahrens, 2004 Maladera wolfgangdierli Ahrens, 2004b: 295 (nomen no¬ vum). Autoserica dierli Frey, 1972: 188. Material examined. See Ahrens 2004b (p. 295). Aedeagus. See Ahrens 2004b (figs 450-453, p. 450). Distribution. Endemic to central Nepal (Fig. 47D). Maladera fistulosa group Key to species of the Maladera fistulosa group (<$<$) 1 Pronotum and elytra with fine and significantly larg¬ er punctures. Body > 7 mm.2 1 ’ Pronotum and elytra only with fine, dense punctures. Body < 7 mm. M. pubescens (Arrow) 2 Parameres curved and reflexed. M. granigera sp. n. 2’ Parameres more straight and less reflexed. . M. seriatoguttata sp. n. Maladera granigera sp. n. (Figs 11V-X, 32K, 47E) Type material examined. Holotype: S “S-India, Kerala, Cardamom Hills ca 50 km NW of Pathanamthitta near Pambaiyar riv., alt. 300 m/ 6.-9.V. 1994 77°05’N 9°25’E E. Kejval lgt./ IS 90” (CPPB). Description. Length: 8.2 mm, length of elytra: 6.1 mm, width: 5.1 mm. Body oval, dark brown, ventral surface including legs reddish brown, antenna yellow, labro- clypeus moderately shiny, remainder of dorsal surface dull, with moderately dense and erect long setae on head, pronotum and elytra. Labroclypeus narrowly subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent to broadly rounded anterior angles, later¬ al border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin very shal¬ lowly sinuate medially; surface feebly convex and mod¬ erately shiny, finely and densely punctate, distance be¬ tween punctures subequal their diameter, with numerous erect setae; frontoclypeal suture feebly impressed and weakly angled medially; smooth area anterior to eye ap¬ proximately as wide as long; ocular canthus moderately long and narrow, nearly impunctate, with a single short terminal seta. Frons with fine, dense punctures, with dense long erect setae beside eyes and behind frontoclypeal su¬ ture. Eyes small, ratio of diameter/ interocular width: 0.6. Antenna with ten antennomeres; club with three anten- nomeres, slightly shorter than remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins in basal half nearly straight, slightly concave, even¬ ly narrowed towards anterior half, in anterior half mod¬ erately convex and convergent; anterior angles strongly produced and sharp, posterior angles blunt; anterior mar¬ gin weakly produced medially, anterior marginal line widely incomplete medially; surface densely and finely punctate, with numerous long, erect setae on disc, other¬ wise punctures with microscopic setae only; anterior and lateral borders setose, basal margin without marginal line; Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 148 Dirk Ahrens & Silvia Fabrizi hypomeron carinate and slightly produced ventrally. Scutellum wide, triangular, with fine dense punctures each bearing a very minute seta, on basal midline impunctate. Elytra oblong, widest just behind middle, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures, odd intervals with very coarse and slightly elevated punctures each bearing an erect long yellow seta, remainder of punctures with minute setae; epipleural edge robust, ending at broadly rounded external apical angle of elytra, apical declivity with a weak transverse carina; epipleura densely setose, apical border narrowly membranous, apex covered with short microtri- chomes. Ventral surface dull, thorax and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous ex¬ cept for numerous long setae laterally; abdominal stern- ites finely and densely punctate, some punctures with very short or minute setae, each sternite with a distinct trans¬ verse row of coarse punctures each bearing a short seta, penultimate sternite apically with a shiny, smooth chiti- nous border of about a quarter of sternite length. Mesostemum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/2.36. Me¬ dian apophysis of metacoxa nearly glabrous. Pygidium weakly convex, coarsely and densely punctate, midline impunctate, with short setae on apical half, otherwise with minute setae in punctures, along apical margin with a few longer setae. Legs moderately wide; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur dull, anterior edge acute, lacking an adjacent serrated line, densely punctate behind posterior longitu¬ dinal row of setae, posterior ventral margin feebly con¬ cave medially, strongly widened in apical half and very finely serrate apically, posterior dorsal margin not serrat¬ ed, glabrous. Metatibia moderately wide and short, widest behind middle, dorsal and ventral margins subpar¬ allel in posterior two thirds, ratio width/length: 1/2.9, sharply carinate dorsally, with two groups of long spines, basal one at middle, apical one at three quarters of metat- ibial length, beside dorsal margin basally with two single punctures with serrated margins each bearing single spines; lateral face longitudinally convex, with moderate¬ ly dense, fine punctures and minute setae in punctures; ventral margin with five strong equidistant spines; medi¬ al face very sparsely punctate, apex shallowly concave in¬ teriorly near tarsal articulation. Tarsomeres impunctate dorsally, with sparse, short setae ventrally; metatarsomeres ventrally with a strongly serrated, longitudinal ridge and a robust, parallel carina, first metatarsomere a little short¬ er than following two tarsomeres combined and a little longer than the dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with nor¬ mally developed basal tooth. Aedeagus: Fig. 11V-X. Habitus: Fig. 32K. Diagnosis. The species resembles the taxa of the Maladera fistulosa group from Sri Lanka, however, the parameres are still recognisable although being fused with the phallobase. The species differs form all other Indian Maladera species by the elevated coarse punctures at odd intervals and the erect setae on its dorsal face. Etymology. The species name is derived from the Latin word ‘ granigerus ’ (bearing small tubercles) (adjective in the nominative singular). Distribution. See map (Fig. 47E). Maladera pubescens (Arrow, 1916) Autosericapubescens Arrow, 1916: 438. Maladera pubescens'. Krajcik 2012: 154; Fabrizi & Ahrens 2014: 72. Material examined. See Fabrizi & Ahrens 2014 (p. 72). Aedeagus. See Fabrizi & Ahrens 2014 (fig 10A-C, p. 110 ). Distribution. Southern India and Sri Lanka (Fig. 47E). Maladera seriatoguttata sp. n. (Figs 12A-C, 32L, 47E) Type material examined. Holotype: S “India, Karnata¬ ka, 20 km SE Sagar, 600m, 14°06,37’N 75°08,93’E, M. Halada leg., 12.V.2005” (CPPB). Paratypes: 2 <$<$, 6 “India, Karnataka, 20 km SE Sagar, 600m, 14°06,37 , N 75°08,93’E, M. Halada leg., 12.V.2005” (CPPB, ZFMK). Description. Length: 7.7 mm, length of elytra: 5.5 mm, width: 4.5 mm. Body oval, dark brown, ventral surface including legs reddish brown, antenna yellow, labro- clypeus moderately shiny, remainder of dorsal surface dull, with moderately dense and erect long setae on head, pronotum and elytra [partly abraded], Labroclypeus narrowly subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and convergent to broadly rounded anterior angles, later¬ al border and ocular canthus producing an indistinct blunt angle, margins weakly reflexed, anterior margin very shal¬ lowly sinuate medially; surface feebly convex and mod¬ erately shiny, finely and densely punctate, distance be¬ tween punctures subequal their diameter, with numerous erect setae; frontoclypeal suture feebly impressed and weakly angled medially; smooth area anterior to eye ap¬ proximately as wide as long; ocular canthus moderately long and narrow, nearly impunctate, with a single short terminal seta. Frons with fine, dense punctures, with dense long erect setae beside eyes and behind frontoclypeal su- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 149 ture. Eyes small, ratio of diameter/ interocular width: 0.61. Antenna with ten antennomeres; club with three anten- nomeres, as long as remaining antennomeres combined. Mentum elevated and flattened anteriorly. Pronotum moderately wide, widest at base, lateral mar¬ gins in basal half nearly straight, slightly concave, even¬ ly narrowed towards anterior half, in anterior half mod¬ erately convex and convergent; anterior angles strongly produced and sharp, posterior angles blunt; anterior mar¬ gin weakly produced medially, anterior marginal line widely incomplete medially; surface densely and finely punctate, with numerous long, erect setae on disc, other¬ wise punctures with microscopic setae only; anterior and lateral borders setose, basal margin without marginal line; hypomeron carinate and slightly produced ventrally. Scutellum wide, triangular, with fine and dense punctures each bearing a very minute seta, on basal midline impunc- tate. Elytra oblong, widest just behind middle, striae finely impressed, finely and densely punctate, intervals flat, with fine, moderately dense punctures, odd intervals with very coarse and slightly elevated punctures each bearing an erect long yellow seta, remainder of punctures with minute setae; epipleural edge robust, ending at broadly rounded external apical angle of elytra, apical declivity with a weak transverse carina; epipleura densely setose, apical border narrowly membranous, apex covered with short microtri- chomes. Ventral surface dull, thorax and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous ex¬ cept for numerous long setae laterally; abdominal stem- ites finely and densely punctate, some punctures with very short or minute setae, each sternite with a distinct trans¬ verse row of coarse punctures each bearing a short seta, penultimate sternite apically with a shiny, smooth chiti- nous border of about a quarter of sternite length. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/2.18. Me¬ dian apophysis of metacoxa nearly glabrous. Pygidium weakly convex, coarsely and densely punctate, midline impunctate, with short setae on apical half, otherwise with minute setae in punctures, along apical margin with a few longer setae. Legs moderately wide; femora with two longitudinal rows of setae, finely and moderately densely punctate. Metafemur dull, anterior edge acute, lacking an adjacent serrated line, densely punctate behind posterior longitu¬ dinal row of setae, posterior ventral margin feebly con¬ cave medially, strongly widened in apical half and very finely serrate apically, posterior dorsal margin not serrat¬ ed, glabrous. Metatibia moderately wide and short, widest behind middle, dorsal and ventral margins subpar¬ allel in posterior two thirds, ratio width/length: 1/3.4, sharply carinate dorsally, with two groups of long spines, basal one at middle, apical one at three quarters of metat- ibial length, beside dorsal margin basally with two single punctures with serrated margins each bearing single spines; lateral face longitudinally convex, with moderate¬ ly dense, fine punctures and minute setae in punctures; ventral margin with five strong equidistant spines; medi¬ al face very sparsely punctate, apex shallowly concave in¬ teriorly near tarsal articulation. Tarsomeres impunctate dorsally, with sparse, short setae ventrally; metatarsomeres ventrally with a strongly serrated, longitudinal ridge and a robust, parallel carina, first metatarsomere a little short¬ er than following two tarsomeres combined and a little longer than the dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with nor¬ mally developed basal tooth. Aedeagus: Fig. 12A-C. Habitus: Fig. 32L. Diagnosis. Maladera seriatoguttata sp. n. is in the shape of the male genitalia and its external morphology very sim¬ ilar to M. granigera sp. n.. M. seriatoguttata differs from the latter by the more straight and less reflexed parameres (lateral view). Etymology. The species name is derived from the Latin word ‘ seriatus ’ (in rows) and ‘ guttatus ’ (spotted) (adjec¬ tive in the nominative singular). Variation. Length: 7.7-8.9 mm, length of elytra: 5.5-6.1 mm, width: 4.5-5.0 mm. Ground colour of the body varies from reddish brown to dark brown. Female: Antennal club slightly shorter than in male, otherwise no visible sexual dimorphism. Distribution. See map (Fig. 47E). Maladera granuligera group Key to species of the Maladera granuligera group 0(j ) 1 Parameres more than half as long as length of phal- lobase. M. granuligera (Blanchard) L Parameres less than half as long as length of phal- lobase.2 2 Metatibia long and moderately wide, ratio width/length: 1/3.3-3.0.3 2’ Metatibia short and wide, ratio width/length -1/2.6. .4 3 Labroclypeus wide. Parameres longer. Phallobase at right side with a long apophysis. . M. gopaldharae Ahrens 3’ Labroclypeus narrow. Parameres shorter. Phallobase at right side with a short apophysis. . M. weigeli Ahrens 4 Phallobase at right side with an apophysis. Parameres without basal lobes.5 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 150 Dirk Ahrens & Silvia Fabrizi 4’ Phallobase at right side without an apophysis. Each paramere with a small basal lobe. ..M. profana sp. n. 5 Apical dorsal sinuation of phallobase small, phallobase ventrolaterally with small longitudinal elevations. Left paramere in distal half with a sharp longitudinal dor¬ sal carina. M. siniaevi Ahrens 5’ Apical dorsal sinuation of phallobase larger, phallobase ventrolaterally without longitudinal elevations. Left paramere in distal half without longitudinal carina. .. . M. tumida Ahrens Maladera gopaldharae Ahrens, 2004 Maladera gopaldharae Ahrens, 2004b: 286. Material examined. See Ahrens 2004b (p. 286). Aedeagus. See Ahrens 2004b (figs 436^138, p. 448). Distribution. Endemic to Sikkim (Fig. 47F). Maladera granuligera (Blanchard, 1850) (Fig. 12D-F) Omaloplia granuligera Blanchard, 1850: 78. Serica granuligera: Brenske 1898: 225. Autoserica granuligera: Dalla Torre 1912: 23. Maladera granuligera. Krajcik2012: 154. Type material examined. Lectotype (here designated): S “Mus. Paris Indes Orient./ O. granuligera Cat. Mus. In- des orient” (MNHN). Redescription. Length: 8.8 mm, elytral length: 6.8 mm, width: 6.2 mm. Body wide, oval, black, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus wide, trapezoidal, lateral margins slight¬ ly convex and convergent, producing with the ocular can- thus a nearly straight line, anterior angles blunt, weakly rounded, anterior margin very shallowly sinuate medial¬ ly, margins weakly reflexed; surface flat, very densely and finely punctate, distance between punctures equals their diameter, glabrous; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus 4 times as wide as long; ocular canthus short and very wide, finely and very densely punctate ,without terminal seta. Frons with more superficial but dense punctures. Antenna with ten an- tennomeres, club with three antennomeres, as long as re¬ maining antennomeres combined. Eyes small, ratio diam¬ eter/ interocular width: 0.47. Mentum convexly elevated and flattened anteriorly. Pronotum strongly convex, widest shortly before base, lateral margins evenly convex and convergent anteriorly, slightly narrowed towards the strongly rounded posterior angles, anterior angles sharp and distinctly produced; an¬ terior margin with a fine marginal line, weakly convexly produced medially; basal margin without marginal line, also in the posterior angles; surface finely and evenly densely punctate, setae of lateral and anterior margin lack¬ ing, punctures with microscopic setae only. Scutellum wide, triangular, punctation as in pronotum. Elytra strongly convex, widest behind middle, striae finely impressed, finely punctate, intervals flat, very densely and coarsely punctate, distance between punctures equals their diameter; epipleura ending at very strongly rounded external apical angle of elytra; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metastemal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.67. Abdominal sternite finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium moderately convex, coarsely and very densely punctate, lateral and apical margins with very short setae. Legs moderately wide; femora superficially and sparse¬ ly punctate, with two longitudinal rows of setae. Metafe¬ mur shiny, anterior edge acute, without adjacent serrated line, anterior longitudinal row of setae reduced; posteri¬ or ventral margin almost straight, weakly widened in api¬ cal half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia short and wide, flattened, ra¬ tio width/length: 1/2.2, sharply carinate dorsally, with two groups of spines, basal one at nearly half, apical one at three quarters of metatibial length, basally with a few very fine and short setae; lateral face weakly longitudinally con¬ vex, nearly completely smooth, dorsally with a few large but superficial punctures; apex concavely sinuate interi¬ orly near tarsal articulation. Tarsomeres impunctate dor¬ sally, with dense, fine setae ventrally; metatarsomeres ven¬ trally glabrous, with a strongly serrated carina, subven- trally with a second, smooth longitudinal carina, first metatarsomere little longer than following two tar¬ someres combined, dorsal tibial spur lacking. Protibia short, bidentate. Anterior claws lacking in lectotype. Aedeagus: Fig. 12D-F. Remarks. No other specimens than the type have been recorded so far from India. Therefore, its occurrence in India is doubtful. Maladera profana sp. n. (Figs 12G-I, 32M, 47F) Type material examined. Holotype: S “NE India, Meg¬ halaya, 1 km E Tura, 500 - 600m, 25°30‘N, 90°14‘E, 2.- 5.V.2002, M. Tryzna & P. Benda leg.”(CPPB). Paratypes: 11 S3, 7 $ $ “NE India, Meghalaya, 1 km E of Tura, 500 Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 151 - 600m, 25°30‘N, 90°14‘E, 2.-5.V.2002, M. Tryzna & P. Benda leg.”(CPPB, ZFMK). Description. Length: 10.6 mm, elytral length: 7.8 mm, width: 6.7 mm. Body wide, oval, black, dorsal surface dull, labroclypeus, tarsomeres, and tibiae shiny, glabrous. Labroclypeus wide, lateral margins strongly convex and convergent, lateral margins produce with ocular canthus an indistinct angle, anterior angles strongly convex, an¬ terior margin shallowly sinuate medially, margins weak¬ ly reflexed; surface shiny, at base narrowly dull, weakly convex medially, densely and finely punctate, distance be¬ tween punctures partly smaller their diameter, with a few single setae anteriorly; frontoclypeal suture finely incised, bluntly angled medially; ocular canthus moderately wide and long (1/3 of ocular diameter); , finely and densely punctate, with a short terminal seta. Eyes large, ratio di¬ ameter/ interocular width: 0.71. Frons with more super¬ ficial and moderately dense punctures, with a few single setae beside eyes. Antenna with ten antennomeres, club with three antennomeres and straight, 1.5 times as long as remaining antennomeres combined. Mentum convex- ly elevated and flattened anteriorly. Pronotum strongly convex, widest at base, lateral mar¬ gins evenly convex and convergent anteriorly, slightly nar¬ rowed towards strongly rounded posterior angles, anteri¬ or angles sharp and distinctly produced; anterior margin with a fine marginal line, weakly convexly produced me¬ dially; basal margin without marginal line; surface fine¬ ly and evenly densely punctate, setae of lateral and ante¬ rior margin robust but sparse, punctures with microscop¬ ic setae only. Scutellum wide, triangular, punctation as in pronotum. Elytra strongly convex, widest shortly behind middle, striae finely impressed, finely punctate, intervals flat, densely and finely punctate, glabrous; epipleura ending at strongly rounded external apical angle of elytra, sparsely setose; apex of elytra with a fine membraneous rim of short microtrichomes. Ventral face coarsely and densely punctate, nearly glabrous, only metastemal plate and lateral metacoxa with a few robust setae. Mesosternum between mesocoxae wide, 1.5 times as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.91. Abdominal sternite finely and moderately densely punctate, each with a row of coarse punctures bearing each a robust seta. Pygidium moderately convex, coarsely and densely punctate, later¬ al and apical margins with short or long setae. Legs wide and short; femora superficially and sparse¬ ly punctate, with two longitudinal rows of setae. Metafe¬ mur shiny, anterior edge acute, without adjacent serrated line, anterior longitudinal row of setae reduced; posteri¬ or ventral margin almost straight, weakly widened in api¬ cal half, neither ventrally nor dorsally serrated but smooth, glabrous. Metatibia very short and wide, flattened, ratio width/length: 1/2.2, sharply carinate dorsally, with three groups of spines, basal one at one third, median one shortly behind middle, and apical one at three quarters of metatibial length, basally with a few fine and short setae; lateral face weakly longitudinally convex, completely smooth along middle, dorsally and ventrally with a few superficial punctures; medial face impunctate, apex shal¬ lowly and concavely sinuate interiorly near tarsal articu¬ lation. Tarsomeres impunctate dorsally, with dense, fine setae ventrally; metatarsomeres ventrally glabrous, with a strongly serrated carina, subventrally with a second, smooth longitudinal carina, first metatarsomere little short¬ er than following two tarsomeres combined, and as long as dorsal tibial spur. Protibia short, bidentate. Anterior claws symmetrical. Aedeagus: Fig. 12G-I. Habitus: Fig. 32M. Diagnosis. Maladera profana sp. n. differs from all oth¬ er species of the M. granuligera group by the lack of an apical apophysis of the phallobase as well as in the pres¬ ence of a short basal lobe on each paramere. Variation. Length: 10.6-12.2 mm, elytral length: 7.8-8.8 mm, width: 6.7-7.5 mm. Female: Antennal club as long as the remaining antennomeres combined; eyes smaller than in male: ratio diameter/ interocular width: 0.51. Etymology. This new species is named from the Latin ad¬ jective ‘ profanus ’ (non sacred) (adjective in the nomina¬ tive singular). Distribution. See map (Fig. 47F). Maladera siniaevi Ahrens, 2004 (Fig. 32N, 47F) Maladera sinaevi Ahrens 2004b: 284. Maladera siniaevi. Ahrens 2006a: 414; Ahrens & Fabrizi 2009b: 275. Material examined. See Ahrens 2004b (p. 284), 2006a: 414; Ahrens & Fabrizi 2009b (p. 275); 4 ex. “NE India, Meghalaya, 1 km E of Tura, 500-600m, 25°30‘N, 90°14‘E, 13.-18.V.2002, M. Tryzna & P. Benda leg.”(CPPB), 1 ex. “Myanmar (Burma) Putao, 500m, 23.V. 1998” (ZFMK). Aedeagus. See Ahrens 2004b (figs 433-435, p. 447). Distribution. Darjeeling, Sikkim, Bhutan, Meghalaya, As¬ sam, northern Myanmar (Fig. 47F). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 152 Dirk Ahrens & Silvia Fabrizi Maladera tumida Ahrens, 2004 Maladera tumida Ahrens, 2004b: 282, 2006a: 414. Serica pruinosa Frey, 1965b (nec Burmeister, 1855): 89 (fig- 3). Material examined. See Ahrens 2004b (p. 282), 2006a (p. 414); Sabatinelli & Ahrens 2015 (p. 141); 7 ex. “Pak¬ istan Azad Jammu & Kashmir prov., SW from Garhi, 1500m, 5.-15.VII.2003 V. Gurko leg.” (ZFMK), 1 ex. “Pakistan near Islamabad City 600m, 24.VII.2005, V. Gurko leg.” (ZFMK), 1 ex. “N India, Uttarakhand, 10.04.2011 14 km NW New Tehri, h=950 m, N30°28’25.3” E078°20’56.2 Anichtchenko A. leg.” (CDUL), 15 ex. “Pakistan: Pri Sohawa, Islamabad 28.vi.2013 leg. Zubair Ahmed” (CAZK). Aedeagus. See Ahrens 2004b (figs 430^132, p. 447). Distribution. In the Himalaya from Pakistan to central Nepal (Fig. 47F). Maladera weigeli Ahrens, 2004 Maladera weigeli Ahrens, 2004a: 212, 2004b: 287. Material examined. See Ahrens 2004a (p. 212); Shrestha et al. 2012 (p. 381). Aedeagus. See Ahrens 2004a (figs 4-6, p. 213). Distribution. Endemic to western Nepal (Fig. 47F). Maladera indica group Key to species of the Maladera indica groupf ) 1 Metatarsomeres laterally not carinate. Hypomeron car¬ inate. Parameres only distal lobe of parameres entire¬ ly symmetric, basal lobes slightly asymmetric.2 1 ’ Metatarsomeres laterally sharply carinate. Parameres entirely symmetric.8 2 Protarsal claws asymmetrical.3 T Protarsal claws symmetrical. M. decolor sp. n. 3 Basal (i.e., dorsal) lobe of parameres long, nearly as long as paramere itself (i.e. ventral lobe).4 3 ’ Basal (i.e., dorsal) lobe of parameres short, much short¬ er than the paramere itself (i.e. ventral lobe).7 4 Antennal club long, at least 1.5 times as long as re¬ maining antennomeres combined and reflexed. Poste¬ rior dorsal margin of metafemur not convexly widened in basal half..5 4’ Antennal club short, only as long as remaining anten¬ nomeres combined and straight. Posterior dorsal mar¬ gin of metafemur strongly convexly widened in basal half exceeding distinctly the posterior ventral margin. . M. indica (Blanchard) 5 Parameres (larger ventral lobe) sharply pointed at apex. . M. clavata (Frey) 5’ Parameres (larger ventral lobe) rounded at apex.6 6 Length of parameres (lateral view) about one third of length of phallobase. M. magnicornis (Moser) 6’ Length of parameres (lateral view) less than one quar¬ ter of length of phallobase. M. hampsoni sp. n. 7 Protarsal claws symmetrical. Phallobase much longer than phallobasal apodeme, and longitudinally convex ventrally. Parameres small, basal lobe sharply point¬ ed. M. brevistylis Ahrens 7’ Protarsal claws asymmetrical. Phallobase not much longer than phallobasal apodeme, and ventrally lon¬ gitudinally concave. Parameres larger, basal lobe mod¬ erately rounded. M. bhutanensis (Frey) 8 Hypomeron not carinate. . M. consularis Ahrens & Fabrizi 8’ Hypomeron carinate.9 9 Dorsomedian sinuation of phallobase modified, blunt or straight, but not covered by a dorsal process. ...10 9’ Dorsomedian sinuation of phallobase covered by a large dorsal process. M. declarata sp. n. 10 Antennal club strongly reflexed and 2.5 times as long as remaining antennomeres combined. . M. sedula sp. n. 10’ Antennal club nearly straight and 1.5 times as long as remaining antennomeres combined. .M minops sp. n. Maladera bhutanensis (Frey, 1975) Autos erica bhutanensis Frey, 1975b: 224. Lasioserica lata Frey, 1975b: 230; syn. by Ahrens 2004b: 251. Lepidoserica lata: Ahrens 1996: 30. Maladera bhutanensis'. Ahrens 2004b: 251; Krajcik 2012: 153. Material examined. See Ahrens 2004b (p. 251). Aedeagus. See Ahrens 2004b (figs 378-381, p. 439). Distribution. Endemic to Bhutan (Fig. 48A). Maladera brevistylis Ahrens, 2004 Maladera brevistylis Ahrens, 2004b: 249. Material examined. See Ahrens 2004b (p. 249). Aedeagus. See Ahrens 2004b (figs 373-375, p. 439). Distribution. So far with two disjunctive occurrences in Nepal and Uttar Pradesh (Fig. 48A). Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 153 Maladera clavata (Frey, 1972) (Figs 12J-L, 320, 48A) Autoserica clavata Frey, 1972: 191. Maladera clavata: Krajcik 2012: 154. Type material examined. Holotypus (clavata)-. S “Type/ Kerala 1970 Calicut distr. 3500 ft. Nathan/ Autoserica clavata n. sp. det. G. Frey 1970” (CF). Paratypes ( clava¬ ta ): 29 SS, 16 $$ “Kerala 1970 Calicut distr. 3500 ft. Nathan/ Paratype/ Autoserica clavata n. sp. det. G. Frey 1970” (CF, ZMHB). Additional material examined. 5 ex. “Nilgiri Hills. H.L. Andrewes/ Nilgiri Hills/ Adrewes Bequest. B.M. 1922- 221” (BMNH), 1 ex. “Nilgiri Hills. G.F. Hampson” (BMNH), 1 ex. “India: Karnataka Aysikeye 1979 Shivayo- gi Coll./ CR 59/ Brit. Mus. 1984-37” (BMNH), 1 ex. “Byran Kuppe 800 m 4.53/ Mysore S. Indien” (ZFMK), 16 ex. “India, Karnataka, Sulya, Medikeri, 150m, 12°32’N 75°29 , E, M. Halada leg., 24.iv.2005” (CPPB), 12 ex. “S India; Tamil Nadu; Nilgiri Hills 11km SE Kotagiri; llOOilOOm; 11°24’N 76°56’E; Kunchappanai P. Pa- cholatko leg.; 3.-15.V.2002” (CPPB), 1 ex. “S India Ker¬ ala; 1250 m; 15 km SW Munnar; 1.-9.V.1997 10,02N 76,58E; Kallar Valley; Dembicky & Pacholatko leg./ IS 73” (ZFMK), 2 ex. “India, Kerala Trivandrum Dt. Poon- mudi Range 3000 ft., IV.-V.71 leg. T. R. S. Nathan” (CF, ZFMK), 7 ex. “S-India, Tamil Nadu state, Nilgiri hills, 15 km SE of Kotagiri, Kunjappanai env., ca. 900 m, 22.- 30.V.1994 Z. Kejval & M. Tryzna leg.” (CPPB), 18 ex. “S-India, Tamil Nadu, Nilgiri hills, 15 km SE of Kotagiri near Kunjappanai, alt. 900 m/ 13-20.V.1994 Kejval lgt.” (ZFMK), 1 ex. “S India-Kerala; 13.iv.1997 7 km N Mu- nar; 1740 m; Eravikulam nat. p.; 10°09‘N 77°04‘E Schintlmeister & Sinaev leg.” (CPPB), 1 ex. “Shembaga- nur Madura. Inde” (ZMHB), 1 $ “Kerala V. 1970 Calicut distr. 3500 ft. Nathan/ Autoserica clavata Frey” (ZMHB). Redescription. Length: 5.6 mm, length of elytra: 4.6 mm, width: 4.0 mm. Body oblong-oval, yellowish brown, head and pronotum reddish brown, antenna yellow, surface dull, labroclypeus shiny, except some single setae on head dor¬ sal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins nearly straight and convergent anteriorly, anterior angles moderately rounded, anterior margin weak¬ ly sinuate medially, margins distinctly reflexed; lateral margin and ocular canthus produce a distinct angle; sur¬ face flat, finely and densely punctate, with a few larger punctures anteriorly each bearing an erect seta; fronto- clypeal suture distinctly incised, angled medially; smooth area anterior to eye convex, as wide as long; ocular can¬ thus moderately long and narrow (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with dense, fine punctures, with a few single long setae beside each eyes. Eyes moderately large, ratio diameter/ interocular width: 0.69. Antenna with ten antennomeres; club with three antennomeres and distinctly reflexed, 2.5 times as long as remaining antennomeres combined. Men- tum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at middle, lat¬ eral margins evenly convex and convergent anteriorly, slightly convexly narrowed towards base; anterior angles moderately produced and sharp, posterior angles blunt; an¬ terior margin almost straight, with complete fine margin¬ al line, base without marginal line; anterior and lateral margins densely setose; surface densely and finely punc¬ tate, midline narrowly impunctate, with minute setae in punctures, otherwise glabrous; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate on basal midline. Elytra widest shortly behind middle, striae distinctly im¬ pressed, finely and sparsely punctate, intervals slightly convex, with fine and evenly dense punctures concentrat¬ ed along striae, with minute setae in punctures, odd inter¬ vals with a few short setae; epipleural edge robust, end¬ ing at strongly curved external apical angle of elytra, epi- pleura sparsely setose; apical border of elytra very nar¬ rowly membraneous, with a very fine rim of microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal stemites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.63. Pygidium moderately convex and dull, coarsely and densely punctate, without midline, with minute setae in punctures, otherwise glabrous except a few robust se¬ tae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with acute anterior margin, without adjacent serrated line, anterior row of setae reduced to a few sin¬ gle setae; posterior ventral margin weakly widened at apex and serrate at distal third, posterior dorsal margin distinct¬ ly serrate, finely shortly setose, in basal half not widened and not exceeding posterior ventral margin. Metatibia short and wide, widest at middle, ratio of width/length: 1/2.38, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quar¬ ters of metatibial length, with a few robust setae basally subparallel to dorsal margin; lateral face longitudinally convex, shiny, sparsely punctate and with minute setae in punctures; ventral margin not serrate, with four equidis¬ tant long and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interi¬ orly near tarsal articulation. Tarsomeres dorsally impunc¬ tate, glabrous, neither laterally nor dorsally carinate, mod- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 154 Dirk Ahrens & Silvia Fabrizi erately setose ventrally; metatarsomeres ventrally glabrous, with a strongly serrated ventral ridge and a smooth subventral longitudinal carina; first metatarsomere as long as following two tarsomeres combined and slight¬ ly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 12J-L. Habitus: Fig. 320. Remarks. Also the females of M. clavata differ from M. indica in the posterior margin of the metafemur, which is strongly widened in basal half in M. indica which is not the case in M. clavata. The aedeagus of the holotype is damaged. Distribution. See map (Fig. 48A). Maladera consularis Ahrens & Fabrizi, 2009 Maladera consularis Ahrens & Fabrizi, 2009b: 269. Material examined. See Ahrens & Fabrizi 2009b (p. 269). Aedeagus. See Ahrens & Fabrizi 2009b (fig. 6J-L, p. 282). Distribution. So far known only from the type locality in NE India, western Arunachal Pradesh (Fig. 48A). Maladera declarata sp. n. (Figs 12M-0, 32P, 48A) Type material examined. Holotype S “NE India, Meghalaya ~8 km N of Shillong, 25°38’N 91°54’E; ~1200m, L. Dembicky leg., 7.-9.v.2004b/45/04” (CPPB). Paratypes: 2 SS “NE India, Meghalaya SW of Cherra- punjee, 25°13 , -15’N 91°40’E; 500-900 m; L. Dembicky leg., 11.-12.V.2004” (CPPB, ZFMK), 1 S “NE India, Meghalaya state, Jaintia Hills reg., Jowai 6-8.VI. 1996 alt. 1350+100 m, GPS N25°27’ E92°12’ (WGS 84) E. Jen- dek & O. Sausa leg./ IS 32” (CPPB). Description. Length: 7.5 mm, length of elytra: 5.6 mm, width: 4.4 mm. Body oblong-oval, dorsal face dark brown, ventral face dark reddish brown, dull, head moderately shiny, except some single setae on head dorsal surface nearly glabrous. Labroclypeus narrow and subtrapezoidal, widest at base, lateral margins straight and convergent anteriorly, anteri¬ or angles strongly rounded, anterior margin distinctly sin¬ uate medially, margins weakly reflexed; lateral margin and ocular canthus produce a distinct angle; surface flat, fine¬ ly and very densely punctate, with a few larger punctures each bearing an erect seta; frontoclypeal suture indistinct¬ ly incised, evenly curved; smooth area anterior to eye weakly convex, three times as wide as long; ocular can¬ thus moderately long and narrow (1/4 of ocular diameter), finely and densely punctate, terminal seta absent. Frons with dense, fine punctures, with a few long setae beside eyes, on disc and behind frontoclypeal suture._Eyes very large, ratio diameter/ interocular width: 0.91. Antenna with ten antennomeres; club with three antennomeres and straight, 1.2 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum transverse, widest shortly behind middle, lat¬ eral margins evenly convex and strongly convergent an¬ teriorly, slightly convexly narrowed towards base, ante¬ rior angles distinctly produced and sharp, posterior angles strongly rounded; anterior margin convex, with complete fine marginal line, base without marginal line; surface densely and finely punctate, punctures less dense on mid¬ line, with minute setae in punctures; anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, punctures less dense on basal midline. Elytra widest at middle, striae distinctly impressed, fine¬ ly and sparsely punctate, intervals slightly convex, with fine and very dense punctures concentrated along striae, with minute setae in punctures, odd intervals with a very few short setae; epipleural edge robust, ending at strong¬ ly curved external apical angle of elytra, epipleura sparse¬ ly setose; apical border of elytra membraneous, with a rim of microtrichomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.76. Pygidium strongly convex and dull, coarsely and densely punctate, without impunctate midline, glabrous except a few robust setae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with anterior margin acute, without adjacent serrated line, anterior row of setae reduced to a few sin¬ gle setae; posterior margin smooth, weakly widened at apex and smooth ventrally, not serrate dorsally, finely shortly setose. Metatibia moderately long and wide, widest at middle, ratio of width/length: 1/3.0, sharply carinate dorsally, with two groups of spines, basal group at mid¬ dle, apical group at three quarters of metatibial length, with a few robust setae basally subparallel to dorsal margin; lat¬ eral face longitudinally convex, shiny but basal third dull, impunctate and glabrous; ventral margin finely serrate, with four equidistant long and robust setae; medial face smooth and glabrous; apex finely serrate, moderately trun¬ cate interiorly near tarsal articulation. Tarsomeres dorsal- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 155 ly densely and finely punctate, glabrous, neither lateral¬ ly nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatar- somere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderate¬ ly long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 12M-0. Habitus: Fig. 32P. Female un¬ known. Diagnosis. Maladera declarata sp. n. is in the shape of the genitalia and in its external morphology similar to M minops. The new species differs from M. minops by the lacking ventral teeth of the apical phallobase (in lateral view), the presence of a symmetric dorsal phallobasal process that is deeply incised, and in the apically strong¬ ly dilated parameres. Etymology. The name (adjective in the nominative sin¬ gular) of the new species is derived from the Latin adjec¬ tive ‘ declaratus' (declared), with reference to its impres¬ sive differences in the aedeagus from the other species of the group. Variation. Length: 7.5-8.0 mm, length of elytra: 5.6-5.9 mm, width: 4, 4-4 .5 mm. Distribution. See map (Fig. 48A). Maladera decolor sp. n. (Figs 12P-R, 33A, 48A) Type material examined. Holotype: S “NE India; Meghalaya; 19993 kmETura; 1150 m; 25°30’N 90°14’E; 18.iv. Dembicky & Pacholatko leg./ 605 Sericini: Asia spec.” (CPPB). Paratypes: 3 SS, 3 “NE India, Meghalaya, SW of Cherrapunjee, 25°13 £ -14‘ N, 91°40‘ E , 500-950m, Dembicky leg., 29.iv.-2.v.2005” (CPPB), 3 S3, 1 $ “NE India, Meghalaya, SW of Cherrapunjee, 25°13‘-14‘ N, 91°40‘ E, 5.-24.V.2005, 900m, P. Pa¬ cholatko leg.” (CPPB). Description. Length: 9.6 mm, length of elytra: 6.5 mm, width: 6.0 mm. Body oval, yellowish brown, dorsal sur¬ face except labroclypeus dull, some single setae on head dorsal surface nearly glabrous. Labroclypeus wide and trapezoidal, shiny, widest at base, lateral margins convex and convergent anteriorly, an¬ terior angles strongly rounded, anterior margin distinctly sinuate medially, margins strongly reflexed; lateral mar¬ gin and ocular canthus produce an indistinct angle; sur¬ face flat, finely and densely punctate, with numerous coarse punctures anteriorly each bearing an erect seta; frontoclypeal suture finely incised, medially angled; smooth area anterior to eye weakly convex, as wide as long; ocular canthus short and wide (one third of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons dull, with fine, sparse punctures and micro¬ scopic setae in punctures, a few single erect setae beside eyes. Eyes very small, ratio diameter/ interocular width: 0.45. Antenna with ten antennomeres; club with three an- tennomeres, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum transverse and strongly convex, widest short¬ ly before base, lateral margins convex, distinctly conver¬ gent anteriorly, anterior angles moderately produced and rectangular, posterior angles blunt, weakly rounded at tip; anterior margin convex, with fine and complete margin¬ al line, base without marginal line; surface densely and finely punctate, glabrous; lateral and anterior margin fine¬ ly setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, sparse punctures. Elytra widest at middle, striae distinctly impressed, fine¬ ly and sparsely punctate, intervals weakly convex, with fine and sparse punctures, on odd intervals partly concen¬ trated along striae and with a few short, setae; epipleural edge robust, ending at strongly curved external apical an¬ gle of elytra, epipleura densely setose; apical border of ely¬ tra finely membraneous, with a fine rim of microtrichomes (at ca lOOx magnification). Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae 1.5 times as wide as mesofemur. Ra¬ tio of length of metepisternum/metacoxa: 1/1.6. Pygidi- um moderately strongly convex and dull, finely and dense¬ ly punctate, without smooth midline, with a few longer setae on apex. Legs wide and short, shiny; femora with two longitu¬ dinal rows of setae, finely and sparsely punctate. Metafe¬ mur shiny, with anterior margin acute, without serrated line behind anterior edge, anterior row of setae reduced, posterior margin smooth and strongly widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at behind middle, ratio of width/length: 1/2.4, sharply carinate dorsally, with two groups of spines, basal group shortly at middle, apical group at three quarters of metatibial length, with a few short robust setae basally; lateral face longitudinally con¬ vex, sparsely finely punctate on sides and basal half, glabrous; ventral margin not serrate, with four robust equi¬ distant setae; medial face smooth and glabrous; apex fine¬ ly serrate, moderately truncate interiorly near tarsal artic¬ ulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 156 Dirk Ahrens & Silvia Fabrizi metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatar- somere as long as following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, biden- tate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 12P-R. Habitus: Fig. 33A. Diagnosis. Maladera decolor sp. n. differs from the taxa of the M. sikkimensis group by its larger body size and the short antennal club, from somewhat similar taxa of the M. indica group it differs by the smooth and not serrated dor¬ sal posterior margin of the metafemur. Etymology. The species name (adjective in the nomina¬ tive singular) is derived from the Latin adjective ‘ decol¬ or ’ (discoloured) with reference to its variable and poly¬ morphic colour, which is sometimes lost. Variation. Length: 9.6-10.2 mm, length of elytra: 6.5-7.2 mm, width: 6.0-6.1 mm. Colour very variable: entirely reddish brown, or reddish brown with dark brown elytra, in the latter condition there can be larger light spots on elytra too. Female: antennal club with three antennomeres, slightly shorter than the remaining antennomeres com¬ bined. Distribution. See map (Fig. 48A). Maladera hampsoni sp. n. (Figs 24X-Z, 39J) Type material examined. Holotype: S “Nilgiri Hs. G.F. Hampson 94-98” (BMNH). Description. Length: 7.8 mm, length of elytra: 5.1 mm, width: 4.9 mm. Body oblong-oval, dorsal face dark brown, antenna yellowish, ventral face dark reddish brown, dull, labroclypeus shiny, except some single setae on head dor¬ sal surface nearly glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins straight and convergent an¬ teriorly, anterior angles strongly rounded, anterior margin weakly sinuate medially, margins distinctly reflexed; lat¬ eral margin and ocular canthus produce a distinct angle; surface flat, finely and very densely punctate, with a few larger punctures anteriorly each bearing an erect seta; fron- toclypeal suture distinctly incised, angled medially; smooth area anterior to eye convex, twice as wide as long; ocular canthus moderately long and narrow (1/3 of ocu¬ lar diameter), impunctate, terminal seta absent. Frons dull, with dense, fine punctures, with a few single and long se¬ tae beside each eyes. Eyes moderately farge, ratio diam¬ eter/ interocular width: 0.67. Antenna with ten anten¬ nomeres; club with three antennomeres and moderately reflexed, 1.5 times as long as remaining antennomeres combined. Mentum elevated and slightly flattened ante¬ riorly. Pronotum moderately transverse, widest shortly before base, lateral margins evenly convex and strongly conver¬ gent anteriorly, slightly convexly narrowed towards base, anterior angles moderately produced and rectangular, pos¬ terior angles strongly rounded; anterior margin convex, with complete fine marginal line, base without marginal line; surface densely and finely punctate, with minute se¬ tae in punctures; anterior and lateral margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate on basal midline. Elytra widest shortly behind middle, striae distinctly im¬ pressed, finely and sparsely punctate, intervals convex, with fine and evenly dense punctures concentrated along striae, on midline of interval nearly impunctate, with minute setae in punctures, odd intervals with a few short setae; epipleural edge robust, ending at strongly curved external apical angle of elytra, epipleura sparsely setose; apical border of elytra very narrowly membraneous, with a very fine rim of microtrichomes (visible at ca lOOx mag¬ nification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.4. Pygidium moderately convex and dull, coarsely and densely punctate, without midline, with minute setae in punctures, otherwise glabrous except a few robust setae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with acute anterior margin, without adjacent serrated line, anterior row of setae reduced to a few sin¬ gle setae; posterior ventral margin smooth, weakly widened at apex and smooth, posterior dorsal margin not serrate, finely shortly setose._Metatibia short and wide, widest at middle, ratio of width/length: 1/3.0, sharply car¬ inate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, with a few robust setae basally subparallel to dorsal mar¬ gin; lateral face longitudinally convex, shiny but basal third dull, sparsely punctate and with minute setae in punc¬ tures; ventral margin finely serrate, with four equidistant long and robust setae; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderate¬ ly setose ventrally; metatarsomeres glabrous and with a Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 157 strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere as long as follow¬ ing two tarsomeres combined and distinctly longer than dorsal tibial spur. Protibia moderately long, bidentate; an¬ terior claws symmetrical, basal tooth of both claws blunt¬ ly truncate at apex. Aedeagus: Fig. 24X-Z. Habitus: Fig. 39J. Female un¬ known. Diagnosis. This new species differs from the very simi¬ lar M. magnicornis (Moser) by the much shorter para- meres that are at the apex more strongly rounded. Etymology. The new species is named in honour of its col¬ lector, G.F. Hampson (noun in genitive case). Distribution. So far known only from the Nilgiri Hills, without more precise locality information (Fig. 48A). Maladera indica (Blanchard, 1850) (Figs 12S-U, 33B, 48A) Omaloplia indica Blanchard, 1850: 77. Autoserica indica : Brenske 1898: 276; Frey 1972: 209. Serica nilgirensis Sharp, 1903: 469, syn. n. Autoserica singularis Brenske, 1898: 298, syn. n. Serica laminipes Moser, 1915a: 147, syn. n. Maladera indica : Krajcik 2012: 154. Type material examined. Syntypes ( indica)-. <$ “Muse¬ um Paris Pondicherry Perrotet 1840/ Lectotype Autoser¬ ica indica Bl. G. Frey” (MNHN), 1 S “ Omaloplia indi¬ ca Blanch, ex typis/ Brsk. 1/1898 vid./ O. indica Cat. Mus. Pondicherry [yellow label, handwritten Blanchard]” (MNHN), 1 S “Indica Bl. Pondich. c. Dej./ Coll. R.I.Sc.N.B. Inde Pondicherry ex. coll. Dejean/ Collection E. Candeze/ Autoserica indica Blanch, det. Blanchard/ Type” (ISNB). Syntypes ( nilgiriensis ): 1 <$, 1 $ “S', nil- giriensis Types D. S. Ootacamund/ Type/ Ootacamund C. A. Barber/ Sharp Coll. 1905-313” (BMNH), 1 $ “Cotype/ Ootacamund C. A. Barber/ S. India 1903-78/ Serica nil¬ giriensis Ind. typ. D. S.” (BMNH), 2 88, 1 ? “Ootaca¬ mund C. A. Barber/ S. India 1903-78” (BMNH), 2 SS, 1 $ “Ootacamund C. A. Barber/ Sharp Coll. 1905-313” (BMNH), 2 SS, 1 $ “Cotype/ Ootacamund C. A. Bar¬ ber/ Sharp Coll. 1905-313” (BMNH), 2 1 ? “Octa- mund C. A. Barber/ Serica nilgirensis Sharp co-type [Handschr. Arrow]/ Sharp Coll./ 1905-313.” (MSNG). Syntypes (laminipes): 1 S, 1 $ “Bengalen India/ Serica laminipes Type Moser” (ZMHB), 1 S “Bengalen India/ “ (ZMHB). Syntypes (singularis): 3 SS, 1 $ “Indes Ori¬ entals Mts. Kodeicanel J. Castets 1888/ singularis type Brsk./ E. Brenske 1896/ Museum Paris ex. coll. R. Oberthiir” (MNHN). Additional material examined. 1 S “Coll.R.I.Sc.N.B. Inde/Neelgheris/ Coll. J. Thomson/ O. Indica Bl. [hand¬ written Blanch.]” (ISNB), 1 ex. “India Malabar/ Fry Coll. 1905-100.” (BMNH), 37 ex. “Nilgiri Hills. H.L. An- drewes/Nilgiri Hills/Adrewes Bequest. B.M. 1922-221” (BMNH), 3 ex. “Nilgiri Hills. A.K.W. Downing. B.M. 1923-324” (BMNH), 3 ex. “Dodabetta Nilgiris 8000 ft. 1.-3.V.16 Ramakrishna Coll./ S. India Coimbatore Mus. 1916-140.” (BMNH), 1 ex. “Kanara, S. India/Adrewes Bequest. B.M. 1922-221” (BMNH), 2 ex. “S. India: Sham- baganur, Madura. 1921-146” (BMNH), 25 ex. “S India Kerala; 1250 m; 15 km SWMunnar; 1.-9.V.1997 10,02N 76,58E; Kallar Valley; Dembicky & Pacholatko leg./ IS 73” (CPPB), 1 ex. “India (S): Anamalai Hills: Cinchona: 1067 m, IV. 1959/ P. S. Nathan Coll. Bishop Museum” (BPBM), 10 ex. “S. India: Pulney Hills: Kodaikanal 2000 m IV.53” (BPBM), 2 ex. “Nilgiri Hills S. India T. V. Camp¬ bell/ G. C. Champion Coll. B.M. 1927-409” (BMNH), 5 ex. “S. Indien leg. Nathan/ Pulney Hills Kodaikanal 6500 ft. IV.53” (ZMHB, ZFMK), 16 ex. “S India-Kerala; 13.iv.1997 7 km N Munar; 1740 m; Eravikulam nat. p.; 10°09‘N 77°04‘E Schintlmeister & Sinaev leg.” (CPPB), 4 ex. ($) “ praticola Gue r. Nila Ghiri Gue r./ 24898” (ZMHB), 1 ex. “Shembaganur Madura, Sd. Ind.” (ZMHB), 3 ex. “Chambaganour Madura. Inde” (ZMHB, CF), 3 ex. “India Madura” (ZMHB), 1 ex. Madura Ind. or.” (ZMHB), 1 ex. (tf) “Coll. R.I.Sc.N.B. Inde/ Coll, de Bonneuil Le Moult vendit/ Serica praticola Guer. Inde” (ISNB), 1 ex. “Coll. R.I.Sc.N.B. Inde/Inde/Coll, de Bon¬ neuil Le Moult vendit” (ISNB), 8 ex. “Coll. R.I.Sc.N.B. S. India: Madurai Shembaganur 12.IV. 1930 RP Manuel” (ISNB), 2 ex. “Coll. R.I.Sc.N.B. S. India: Madras Shem¬ baganur 1904/05 R.P du Breuil ex. coll, de Moffarts” (ISNB), 1 ex. “Coll. R.I.Sc.N.B. Inde/ Coll. Dohrn/ Col¬ lection E. Candeze” (ISNB), 3 ex. “Coll. RI.Sc.N.B. S. India: Kodaikanal Pulney Hills (6500 ft.) IV-1953 Re c.: PS. Nathan” (ISNB), 1 ex. “India 8514” (BMNH), 1 ex. “Nilgiri Hills G. F. Hampson 94-89.” (BMNH), 1 ex. “Ootacamund India, ex. Tomlin/ G. C. Champion Coll. B.M. 1927-409” (BMNH), 1 ex. “Nilgiri’s, India, ex Tom¬ lin/ G. C. Champion Coll. B.M. 1927-409” (BMNH), 1 ex. “Nilgiri Hills S. India. T.V. Campbell/ G. C. Champi¬ on Coll. B.M. 1927-409” (BMNH), 2 ex. “S-India, Nil¬ giri Hills Naduvatam, 6000 ft. Mai 1958, P.S.N.” (ZMHB), 10 ex. “Naduvatam v-58 S. Nathan” (SEAN), 1 ex ($) “India Devala V-58” (SEAN), 20 ex. “India-Tamil Nadu, Pakyra, Nilgiri Hills, 2250m, 11°26,9’N 70°36,9’E, leg. M. Halada, 26.4.2005” (CPPB), 5 ex. “South India Nilgiri Hills Naduvatam 1958 6000ft. PS. Natan” (US- NM), 3 ex. “Coll. R.I.Sc.N.B./ India: Nilgiri Hills: Nadu- vattam PS. Nathan” (ISNB), 4 ex. “Coll. R.I.Sc.N.B./ S. India: Madras Shembaganur 1904/05 R.P. Du Breuil ex coll, de Moffarts” (ISNB), 22 ex. “Shembaganur Mad. 1904/05 R.P. Du Breuil” (ISNB), 1 ex. “Coll. R.I.Sc.N.B./ S. India: Coimbatore P. Susai-Nathan” (ISNB), 1 ex. “Coll. Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 158 Dirk Ahrens & Silvia Fabrizi R.I.Sc.N.B./ S.India: Trichinopoly 1904/05 R.P. Du Breuil ex coll, de Moffarts” (ISNB), 1 $ “Omaloplia prati- cola Guer. Neelg” (MNHN). Redescription. Length: 7.2 mm, length of elytra: 5.3 mm, width: 4.5 mm. Body oblong-oval, yellowish brown, head and pronotum reddish brown, antenna yellow, surface dull, labroclypeus shiny, except some single setae on head dor¬ sal surface nearly glabrous. Labroclypeus wide and subtrapezoidal, widest at base, lateral margins weakly convex and convergent anterior¬ ly, anterior angles moderately rounded, anterior margin very weakly sinuate medially, margins distinctly reflexed; lateral margin and ocular canthus produce a distinct an¬ gle; surface slightly convex, finely and densely punctate, with a few larger punctures anteriorly each bearing an erect seta; frontoclypeal suture distinctly incised, angled medially; smooth area anterior to eye convex, 1.5 times as wide as long; ocular canthus moderately long and nar¬ row (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with dense, fine punctures, with a few single long setae beside each eyes. Eyes moderate¬ ly large, ratio diameter/ interocular width: 0.62. Antenna with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres combined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest shortly behind middle, lateral margins evenly convex and convergent an¬ teriorly, slightly convexly narrowed towards base; ante¬ rior angles moderately produced and sharp, posterior an¬ gles blunt; anterior margin almost straight, with complete fine marginal line, base without marginal line; anterior and lateral margins densely setose; surface densely and fine¬ ly punctate, midline narrowly impunctate, with minute se¬ tae in punctures, otherwise glabrous; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures, impunctate on basal midline. Elytra widest shortly behind middle, striae distinctly im¬ pressed, finely and sparsely punctate, intervals slightly convex, with fine and evenly dense punctures concentrat¬ ed along striae, with minute setae in punctures, odd inter¬ vals with a few short setae; epipleural edge robust, end¬ ing at strongly curved external apical angle of elytra, epi- pleura sparsely setose; apical border of elytra very nar¬ rowly membraneous, with a very fine rim of microtri- chomes (visible at ca lOOx magnification). Ventral surface dull, coarsely and densely punctate, glabrous, metastemal disc sparsely covered with fine, short setae; metacoxa with a few longer setae laterally. Abdom¬ inal sternites finely and densely punctate, glabrous, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepistemum/metacoxa: 1/1.45. Pygidium moderately convex and dull, coarsely and densely punctate, without midline, with minute setae in punctures, otherwise glabrous except a few robust se¬ tae along apical margin. Legs moderately long and wide, shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur with acute anterior margin, without adjacent serrated line, anterior row of setae reduced to a few sin¬ gle setae; posterior ventral margin weakly widened at apex and serrate at distal third, posterior dorsal margin distinct¬ ly serrate, finely shortly setose, in basal half strongly con¬ vexly widened exceeding by far the ventral posterior mar¬ gin. Metatibia short and wide, widest at middle, ratio of width/length: 1/3.0, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, with a few robust se¬ tae basally subparallel to dorsal margin; lateral face lon¬ gitudinally convex, shiny, sparsely punctate and with minute setae in punctures; ventral margin finely serrate, with four equidistant long and robust setae; medial face smooth and glabrous; apex finely serrate, moderately trun¬ cate interiorly near tarsal articulation. Tarsomeres dorsal¬ ly impunctate, glabrous, neither laterally nor dorsally car¬ inate, moderately setose ventrally; metatarsomeres ven¬ trally glabrous, with a strongly serrated ventral ridge and a smooth subventral longitudinal carina; first metatar- somere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws sym¬ metrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 12S-U. Habitus: Fig. 33B. Remarks. The designation of the lectotype of Maladera indica by Frey is not valid, as it was not published. The tiny lateral tooth of the parameres as well as the length of the interior lobe of the parameres are quite variable in shape, while the external body shape, the shape and punc- tation of the labroclypeus, as well as the metatibia are all identical among the syntypes of the three species. The shape of the aedeagus is quite variable, too, and does not correlate with the length of antennal club as encountered in the type specimens of nilgiriensis , indica, und laminipes. The genitalia of the male syntypes of Autos er¬ ica singular is, probably dissected by Frey, are virtually identical in shape with those of Maladera indica (Blan¬ chard) and d utoserica laminipes Moser, although anten¬ nal clubs of this form (A. singularis) are only as long as the remaining antennomeres. There is no geographic dif¬ ferentiation and many intermediate forms for the length of antennal club, thus we conclude that club lengths rep¬ resents intraspecific variation. The same uncertainty lies in the ratio of the length of the interior lobes of the para¬ meres in relation to the length of the parameres: accord¬ ing to current observations, they seem to be slightly short¬ er in specimens with longer antennal club, however, there is a wide overlap between the forms and thus a differen¬ tiation based on their morphology is impossible. We there- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 159 fore consider these taxa as synonymous. Distribution. See map (Fig. 48A). Maladera magnicornis (Moser, 1920) (Figs 12V-X, 48A) Autoserica magnicornis Moser, 1920: 3. Maladera magnicornis'. Krajcik2012: 155. Type material examined. Syntype: 1 $ “India Manga¬ lore/ Autoserica magnicornis Type Mos.” (ZMHB). Additional material examined. 1 ex. “India: Karnataka 8.5.1981 Mudigere” (BMNH), 1 ex. “Belgaum/Andrewes Bequest B.M. 1922-221/Belgaum Bombay” (BMNH), 1 ex. (10 setae per interval. M. densipilosa sp. n. 6 Lateral phallobasal apodemes strongly displaced ven- trally. Median lobe of apical phallobase robust and strongly produced. Right paramere very small, hidden within the phallobasal apodeme. ..M. mussardi sp. n. 6’ Lateral phallobasal apodemes at one level with medi¬ an lobe. Right paramere small but always well-visi¬ ble.7 7 Anterior margin of labroclypeus convex.8 7’ Anterior margin of labroclypeus straight or concave- ly sinuate. Dorsal surface always dull or with irides¬ cent shine.15 8 Left and right apical phallobasal apodeme subequal in length.9 8’ Right apical phallobasal apodeme produced signifi¬ cantly distally.11 9 Right distal phallobasal apodeme slightly displaced un¬ der median lobe.10 9’ Right distal phallobasal apodeme not displaced under median lobe. Body shiny. M. nigromicans (Frey) 10 Median phallobasal lobe broader (ca 1/2 of width of max. phallobase in dorsal view), right paramere short and more strongly reflexed. M. garoana sp. n. 10’Median phallobasal lobe very narrow ( 10.0 mm. Right paramere straightly di¬ rected forward, at apex abruptly hooked. . M. alibagensis sp. n. 18 Right paramere small, somehow flattened and with a large terminal and sharply pointed hook. Median lobe of phallobase produced. M. constans sp. n. 18’ Right paramere large, not flattened and without a ter¬ minal hook. Median lobe of phallobase very short. .. . M. amboliensis sp. n. 19 Right paramere curved but strongly flattened. . M. holzschuhi Ahrens 19’Right paramere curved but circular in cross section. .20 20 Right apical phallobasal apodeme at maximum half a phallobasal width longer than the left one.21 20’ Right apical phallobasal apodeme one phallobasal width longer than the left one. . M. submucronata sp. n. 21 Body size <8.0 mm. Anterior face of labrum sparse¬ ly setose. M. pauper sp. n. 21’Body size >8.0 mm. Anterior face of labrum very densely setose. M. sagittula sp. n. 22 Left and right apical phallobasal apodeme subequal in length.23 22’Right apical phallobasal apodeme produced signifi¬ cantly distally.25 23 Left paramere dorsoventrally flattened. Right paramere curved and circular in cross section. . M. marginella (Hope) 23’Left paramere circular in cross section.24 24 Right paramere straight and dorsoventrally flattened. . M. naduvatamensis sp. n. 24’Right paramere produced anteriorly but strongly curved and circular in cross section. . M. lonaviaensis sp. n. 25 Right apical phallobasal apodeme produced laterally, widely exceeding the lateral margin of phallobase. .. . M. bengalensis (Brenske) 25’ Right apical phallobasal apodeme not produced later- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK Sericini of India 181 ally, and not exceeding the lateral margin of phal- lobase.26 26 Body size > 9.0 mm. M. coimbatoreensis sp. n. 26’Body size <8.1 mm.27 27 Right apical phallobasal apodeme long and with a strong dorsal convexity. Body colour dark brown. ... . M. poonmudi (Frey) 27’ Right apical phallobasal apodeme without dorsal con¬ vexity.28 28 Right apical phallobasal apodeme not displaced ven- trally. Species from South India. . M. eusericina nom. n. 28’Right apical phallobasal apodeme displaced ventral - ly. Species from NE India, Indochina, and Eastern Hi¬ malaya. M. sericella (Brenske) 29 Median lobe distinctly exceeding parameres. . M. anaimalaiensis sp. n. 29’Median lobe not exceeding parameres.30 30 Distal median lobe of phallobase fused with the more basal portion of the phallobase. Tarsomeres impunc- tate dorsally.31 30’Distal median lobe of phallobase separated by a dis¬ tinct membrane from the more basal portion of the phallobase. Tarsomeres finely punctate dorsally. . M. basalis (Moser) 31 Left paramere basally convexly widened. . M. burmeisteri burmeisteri (Brenske) 3 E Left paramere basally not widened. . M. burmeisteri alternans (Frey) Maladera alibagensis sp. n. (Figs 15G-J, 34C, 48F) Type material examined. Holotype: $ “India, Maharah- stra [sic] state, Alibag env., 45 km S Bombay, 22.- 24.vi.2006, O. Safranek leg.” (CPPB). Description. Length: 10.2 mm, length of elytra: 6.9 mm, width: 5.7 mm. Body oblong-oval, dark brown, antenna yellowish, dull, labroclypeus shiny, except for some sin¬ gle setae on head dorsal surface glabrous. Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins straight and convergent an¬ teriorly, anterior angles strongly rounded, anterior margin straight, margins moderately reflexed; lateral margin and ocular canthus produce a distinct angle; surface weakly convex medially, with coarse and dense punctures inter¬ mixed with a few coarser ones, bearing each an erect se¬ ta; frontoclypeal suture distinctly incised and slightly el¬ evated, angled medially; smooth area anterior to eye flat and narrow, 4 times as wide as long; ocular canthus mod¬ erately long and narrow (1/3 of ocular diameter), finely densely punctate, with a terminal seta. Frons dull, with dense, fine punctures, with a few single setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.85. Anten¬ na with ten antennomeres; club with three antennomeres and straight, as long as remaining antennomeres com¬ bined. Mentum elevated and slightly flattened anteriorly. Pronotum moderately transverse, widest at base, later¬ al margins evenly convex and convergent anteriorly, an¬ terior angles moderately produced and sharp, posterior an¬ gles blunt, slightly rounded at tip; anterior margin weak¬ ly convex, with fine, medially widely interrupted margin¬ al line, base without marginal line; surface moderately densely and finely punctate, with minute setae in punc¬ tures; anterior and lateral margin finely sparsely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, moderately dense punctures. Elytra widest at middle, striae finely impressed, finely and densely punctate, intervals nearly flat, with fine, mod¬ erately dense punctures, with minute setae in punctures, odd intervals with a few single short setae; epipleural edge robust, ending at convexly rounded external apical angle of elytra, epipleura sparsely setose; lateral and apical bor¬ der of elytra membraneous, with a fine rim of microtri- chomes (visible at ca lOOx magnification). Ventral surface moderately shiny, finely and densely punctate, nearly glabrous, metasternal disc sparsely cov¬ ered with fine, short setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each stemite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur, with a semicircular carina bearing robust setae. Ratio of length of metepisternum/metacoxa: 1/1.81. Pygidium weakly convex, finely and densely punctate, with a nar¬ row smooth midline and several long setae at apex. Legs long and moderately wide, dull; femora with two longitudinal rows of setae, finely and sparsely punctate. Anterior margin of metafemur acute, without adjacent ser¬ rated line, anterior row of setae complete; posterior ven¬ tral margin smooth, moderately widened at ventral apex, dorsal posterior edge smooth, neither serrate, glabrous. Metatibia moderately long and wide, widest at middle, ra¬ tio of width/length: 1/3.15, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short single setae and a serrated line subparallel to dor¬ sal margin; lateral face longitudinally convex, finely and moderately densely punctate, with minute setae in punc¬ tures; ventral margin finely serrate, with four equidistant robust setae, between each of these robust setae a short¬ er one; medial face smooth and glabrous; apex finely ser¬ rate, shallowly sinuate interiorly near tarsal articulation. Tarsomeres minutely sparsely punctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventral¬ ly; metatarsomeres with a strongly serrated ridge ventral¬ ly and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tar- Bonn zoological Bulletin 65 (1 & 2): 1-355 ©ZFMK 182 Dirk Ahrens & Silvia Fabrizi someres combined and distinctly longer than dorsal tib- ial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex. Aedeagus: Fig. 15G-J. Habitus: Fig. 34C. Diagnosis. Maladera alibagensis sp. n. differs from the, in external characters similar, M. kanarana (Moser) by the slightly larger body and in the shape of the aedeagus. The right distal phallobasal apodeme is on the same level as the left one in M. alibagensis , while in M. kanarana (Moser) it is produced distally; the right paramere is in its basal two thirds nearly straight in the new species, while in M. kanarana it is strongly curved. Etymology. The new species is named with reference to its type locality, Alibag (adjective in the nominative sin¬ gular). Distribution. See map (Fig. 48F). Maladera amboliensis sp. n. (Figs 15K-N, 34D, 48E) Type material examined. Holotype: $ “India, Maharash¬ tra, ca 20km E of Savantvadi, W of Amboli, 15°57’N 17°53’E [sic!], ca 700m, 21.V.2006, Z. Kejval lgt” (ZFMK). Paratypes: 1 $ “India, Maharashtra, ca 20km E of Savantvadi, W of Amboli, 15°57’N 73°59’E, ca700m, 21.V.2006, Z. Kejval lgt.” (CBBP), 1