i;^ 2!^i^S, YizZ^\. -J^^J^^ BOTANY FOR LADIES; A POPULAR INTRODUCTION Natural ^ps'ttm of ^Slantig, ACCORDING TO TEE CLASSIFICATION OF DE CANDOLLE- LIBRARY HEW YORK BY ^OTANiCAL &ARDBH MRS. LOUDON, Author of " Instructions in Gardening for Ladies," " Year-Book of Natural History," &c. &c. LONDON: JOHN MURRAY, ALBEMARLE STREET. LL LONDON : BRADBURY AND BVAN8, PRINTERS, \AHITEFRIARS PREFACE. When I was a child, I never could learn Botany. There was something in the Linnean system (the only one then taught) excessively repugnant to me ; I never could remember the different classes and orders, and after several attempts the study was given up as one too difficult for me to master. When I married, however, I soon found the necessity of knowing something of Botany, as well as of Gardening. I always acconipanied my husband in his visits to different gardens ; and when we saw beauti- ful flowers, I was continually asking the names, though alks ! these names, when I heard them, conveyed no ideas to my mind, and I was not' any wiser than before. Still the natural wish to know something of what we admire, impelled me to repeat my fruitless questions ; till at last, vexed at my ignorance, and ashamed of not PREFACE. being able to answer the appeals which gar- deners often made to me in doubtful cases, (supposing that Mr. Loudon's wife must know everything about plants,) I determined to learn Botany if possible ; and as my old repugnance remained to the Linnean system, I resolved to study the Natural one. Accordingly I began ; but when I heard that plants were divided into the two great classes, the Vasculares and the Cellulares, and again into the Dicotyledons or Exogens, the Monocotyledons or Endogens, and the Acotyledons or Acrogens, and that the Dicotyledons were redivided into the Dichla- mydese and Monochlamydese, and again into three sub-classes, Thalamiflorse, Calyciflorse, and Corolliflorae, I was in despair, for I thought it quite impossible that I ever could remember all the hard names that seemed to stand on the very threshold of the science, as if to forbid the entrance of any but the initiated. Some time afterwards, as I was walking through the gardens of the Horticultural So- ciety at Chiswick, my attention was attracted by a mass of the beautiful crimson flowers of PREFA.CE. V Malope grandiflora. I had never seen the plant before, and I eagerly asked the name. "It is some Malvaceoiis plant," answered Mr. Loudon, carelessly ; and immediately afterwards he left me to look at some trees which he was about to have drawn for his Arboretum Britannicum. " Some Malvaceous plant," thought I, as I con- tinued looking at the splendid bed before me ; and then I remembered how much the form of these beautiful flowers resembled that of the flowers of the crimson Mallow, the botanical name of which I recollected wasMalva. " I wish I could find out some other Malvaceous plant,'' I thought to myself; and when we soon afterwards walked through the hothouses, I continued to ask if the Chinese Hibiscus, which I saw in flower there, did not belong to Malvaceae. I was answered in the affirmative ; and I was so pleased with my newly-acquired knowledge, that I was not satisfied till I had discovered every Malvaceous plant that was in flower in the garden. I next learned to know the Crucife- rous and Umbelliferous plants ; and thus I acquired a general knowledge of three extensive VI PREFACE. orders with very little trouble to myself. My attention was more fairly aroused, and by learn- ing one order after another, I soon attained a sufficient knowledge of Botany to answer all the purposes for which I wished to learn it, without recurring to the hard words w^hich had so much alarmed me at the outset. One great obstacle to my advancement was the difficulty I had in understanding botanical works. With the ex- ception of Dr. Lindley's Ladies' Botany, they were all sealed books to me ; and even that did not tell half I wanted to know, though it con- tained a great deal I could not understand. It is so difficult for men whose knowledge has grown with their growth, and strengthened with their strength, to imagine the state of profound ignorance in which a beginner is, that even their elementary books are like the old Eton Grammar when it was written in Latin — they require a master to explain them. It is the want that I have felt that has induced me to write the following pages; in which I have endea- voured to meet the wants of those wdiomaybenow in the same difficulties that I was in myself. PREFACE. Vli The course I pursued is also tliat which I shall point out to my readers. I shall first endeavour to explain to them as clearly as I can the botanical characteristics of the orders which contain plants commonly grown in Bri- tish gardens ; and at the end of my work I shall lay before them a slight outline of all the orders scientifically arranged, which they may study or not as they like. Most ladies will, however, probably be satisfied with knowing the orders containing popular plants ; and these, I am con- fident, they will never repent having studied. Indeed, I do not think that I could form a kinder wish for them, than to hope that they may find as much pleasure in the pursuit as I have derived from it myself. Whenever I go into any country I have formerly visited, I feel as though I were endowed with a new sense. Even the very banks by the sides of the roads, which I before thought dull and uninteresting, now appear fraught with beauty. A new charm seems thrown over the face of nature, and a degree of interest is given to even the com- monest weeds. I have often heard that know- PREFACE. ledge is power, and I am quite sure that it contributes greatly to enjoyment. A man knowing nothing of natural history, and of course not caring for anything relating to it, may travel from one extremity of a country to the other, without finding anything to interest, or even amuse him ; but the man of science, and particularly the Botanist, cannot walk a dozen yards along a beaten turnpike-road with- out finding something to excite his attention. A wild plant in a hedge, a tuft of moss on a wall, and even the Lichens which discolour the stones, all present objects of interest, and of admiration for that Almighty Power whose care has provided the flower to shelter the infant germ, and has laid up a stock of nourish- ment in the seed to supply the first wants of the tender plant. It has been often said that the study of nature has a tendency to elevate and ameliorate the mind ; and there is perhaps no branch of Natural History which more fully illustrates the truth of this remark than Botany. COXTEXTS. PART I. PAGE Introduction ......... 1 Miscellaneous Orders — Preliminary Observations . . . 3 CHAPTER I. The Order Ranunculace^ : illustrated by the Garden Raiiunculus; the Butter-cup; the Peony ; the Anemone; the Hepatica ; the Clematis; the Christmas Rose ; the Winter Aconite; Monkshood; the Larkspur; and the Columbine . . . . . The Genus Rammculus, 10— The Genus Ficaria, 16— The Genus Pasonia, 18 — The Genus Anemone, 19 — The Genus Clematis, 23 —The Genus Helleborus, &c. 25— The Genus Aconitum, 27— The Genus Delphinium, 29— The Genus Aquilegia, 31. CHAPTER n. The Order Leguminos.e : illustrated by the Sweet Pea ; the Red Clover ; Acacia armata; the Sensitive Plant; the Baibadoes Flower-fence; the Carob- tree ; the Tamarind ; the Senna; the Gleditschia ; the Logwood ; the Judas-tree ; and the Kentucky Coffee-tree ........ 35 Tribe I. Papilionaceous Flowers, 36—11. Mimosa, 41— III. Csesal- pinese, 44. CHAPTER III. The Order Rosace^e : illustrated by different kinds of Roses; the Potentilla ; the Strawberry ; the Raspberry ; Spiraea ; Kerria or Corchorus japonica ; the Almoud ; the Peach and Nectarine ; CONTENTS. PAGK the Apricot ; the Plum ; tlie Cherry ; the Apple ; the Pear ; The Mountain Ash ; the White Beam Tree ; the Quince ; Pyrus or Cydonia japonica ; the Hawthorn ; the Indian Hawthorn ; the Medlar ; Photinia ; Eriobotrva ; Cotoneaster; Amelanchier; Burnet ; and Alchemilla, or Ladies' Mantle . . .50 Tribel.Roseas, 51— II. Potentilleae or Dryades, 54— III. Spirseeas, 58— IV, Amygdaleas, 60— V. Pomeae, 65— VI. Sanguisorbeae, 73. CHAPTER IV. The Order Onagraceje ; illustrated • by the different kinds of Fuchsia ; Oenothera, or the Evening Tree-Primrose ; Godetia ; Epilobium, or the French Willow-herb ; and Clarkia . . 75 The Genus Fuchsia, 75— The Genus CEnothera, 79— The Genus Clarkia, 83. CHAPTER V. The Order Rubiace^ : illustrated by the Cinchona, or Peruvian Bark ; Luculia gratissima ; Cape Jasmine ; Rondeletia ; Coffee ; Ixora ; Ipecacuanha ; Madder ; Galium ; Woodruff ; and Crucinella stylosa . ...... 85 The Genus Cinchona and its allies, 86— The Genus Gardenia and its allies, 89— The Genus Rondeletia and its allies, 90— The Genus Coffea and its allies, 91— The Genus Galium and its allies, 94. CHAPTER VI. The Order Compositje : illustrated by the Succory ; the Sow- Thistle ; the Dandelion ; the Burdock ; the Daisy ; the Chry- santhemum ; Feverfew ; Pelhtory of Spain ; Wild Chamomile ; True Chamomile ; Yarrow; the Bur-Marigold; Groundsel; Ragwort ; Bird's Tongue ; Purple Jacobaea ; Cineraria ; Sun- flower ; Mutisia ; and Triptilion 98 Tribe I. Cichoracese, 101—11. Cynarocephalae, 103— III. Corymbi- ferae, 104— IV. Labiataefiorae, 107. CHAPTER VII. The Order Ericace^: : illustrated by the Common or Besom Heath ; the Moor Heath ; Cape Heaths ; Ling or Heather ; Androaieda ; Lyonia ; St. Dabseoc's Heath ; Arbutus ; the CO^' TENTS. Bearberrv; Gaultliena; Clethra ; Rhododendron ; Indian or Chinese Azaleas; Yellow Azalea; American Azaleas ; Rho- dora ; Kalmia ; Menziesia : Loiseleuria ; Ledum ; Leiophyl- lura ; the Bilberry ; the AYhortleberry ; the Cranberry ; Pyrola ; and Mouotropa ...... 109 Tribe I. Ericea?, 110— Sub-Tribe I. Ericeae Nonnales, 111—11. An- dromedese, 115— Tribe U. Rhodoreae, 120— III. Vaccinieas, 130 —IV. Pyroleae, 132. CHAPTER VIII. The Order Oleace^, or Jasmineae : illustrated by the Common White Jasmine ; the Yellow Jasmine ; the Privet ; the Phil- lyrea ; the Olive; the Fringe-tree {Chionanthiis virginica)', the Lilac ; the Common Ash ; and the Manna or Flowering Ash 133 Tribe I. Jasmineae, 134—11. Oleineae, 136. CHAPTER IX. The Order SoLANACE^ : illustrated by the Bitter-Sweet ; Garden Nightshade ; Potato ; Egg-Plant ; Tomato ; Capsicum ; Win- ter Cherry ; Cape Gooseberry ; the Deadly Nightshade ; Lycium, or Duke of Argyle's Tea-tree ; Oestrum ; Yestia; Tobacco ; Petunia ; Nierembergia ; Salpiglossis ; Schizanthus ; Henbane ; Datura ; Brugmansia ; Solandi-a ; Yerbascum ; Celsia ; Nolana ; &c. 141 Tribe I. Solanaceas, 142 — II. Nicotianeae, 147 ; HI- Verbascine , 153— IV. Nolaneae. 155. CHAPTER X. The Order Urticace.e : illustrated by the Common Nettle ; the Hop ; the Hemp ; the Pellitory of the Wall ; the Bread-Fruit Tree ; the Jack-tree ; the Cow-tree, or Palo de Yacca ; the Upas or Poison-tree of Java ; the Mulberry ; the Paper Mul- berry ; the Osage Orange, or Madura ; the Common Fig ; Ficus Sycamorus ; the Banyan Tree ; the Indian-Rubber Tree; and Ficus religiosa . . . . . .157 Tribe I. Urticaceae, 158—11. Artocarpae, 163. CONTENTS. """"■" PAGE 38. Hypericinese— the Hyperi- cum Tribe . . .312 39. Guttifera; — the Mangos- teen Tribe . . 313 40. Marcgraaviaees . . lb. 41. Hippocratacese . . . ib. 42. Erythroxylea? — the Red Wood Tribe . . .314 43. 3Ialpigliiace8?— theBarba- does Cherry Tribe . . 314 44. Acerineas the Maple Tribe . . . .315 45. Hippocastanese, or ^scu- laceje— the Horse-chest- nut Tribe . . . 322 46. Rhizoboleae — the Caryocar Tribe . . . .327 47. Sapindacese — the Soap-tree Tribe . . . . ib. 48. Meliacea?— the Bead-tree Tribe . . . ,328 48*.Cedrelea;— the Mahogany Tribe , . . . 329 49. Ampelideas the Vine Tribe . . . . ib. 50. Geraniaceae — the Gera- nium Tribe . . . 332 51. Tropaeolacese — the Nastur- tium Tribe . . .337 51^. Limnautheas . . . ib. 52. Balsamineae— the Balsam Tribe . . . .338 53. Oxalideae — the Wood-sor- rel Tribe . . . 339 54. Zygophylleaj — the Bean- caper Tribe . . . 340 55. Rutaceae— the Rue Tribe . ib. 56. Simarubaceffi . . . 342 57. Ochnaceje . . . ib. 58. Coriareas . . . . ib. § II.— CALYcrFLOR.a;. 59. Celastrinea; . . .343 60. Rhamnaceae . . . 345 61. Bruniaceffi . . . 346 62. Samydese . . . . ib. ()3. Homalineae . . . ib. 64. ChailletiacesB . . . 347 65. Aquilarineae . . . ib. 66. Terebinthaceas — the Tur- pentine Tribe . . . ib. 67. Leguminosae— (See Chap. n. in p. 35) . . . 349 ORDER 68. Rosaceae — (See Chap. iii. in p. 50) . . . . 69. Calycanthaceae 70. Granateae . . . . 71. Memecylea? i 72. Combretacese . . . 73. Vochysieae 74. Rhizophoreae 75. Lophireas 76. Onagrarias — (See Chap. iv. in p. 75) . . . . 77- Halorageae, or Cercodiaceae 78. Ceratophylleas . 79. Lythrarieae, or Salicarise . 80. Tamariscineae — the Ta- marisk Tribe . 81. Melastomaceae . 82. jUangieae. . . . 83. Philadelpheae— the Mock- oi-ange Tribe 84. MyrtacesB — the Myrtle Tribe . . . . 85. Cucurbitaceae— the Gourd Tribe . . . . 86. Passifloreac — the Passion- iiower Tribe . . . 86*. Malesherbiaces 87- Loase^B . . . . 83. Turneriaceae . 89. Portulaceae — the Purslane Tribe . . . . 90. Paronychiese . 91. Crassulaeeae— the House- leek Tribe . . . 92. Ficoideas — the Fig-mari- gold Tribe . 93. Cactacese the Cactus Tribe . , . . 94. Grossularieae — the Goose- berry Tribe . 95. Esealloniaceae 96. Saxifragacese . 97. Cunoniaceae 98. Umbelliferaj — Umbellife- rous Plants, or the Parsley Tribe 99. Araliacea; . ... 99*. Hamamelideae 100. Caprifoliaceae, or the Ho- neysuckle Tribe . . 101. Lorantheae 102. ChlorantheK . . . 350 351 352 353 ib. ib. ib. 354 ib. 355 ib. 356 ib. 357 ib. ib. 360 361 362 ib. 363 364 365 ib. 367 CONTENTS. XV ORDBR PAGE ORDER PAGB 103. Rubiaceae (See Chap. v. 125. Jasmineae (See Chap. viii. p. 85) .... 386 p. 134) 401 104. Opercularie« 387 126. Strychnea? . . . . ib. 105. Valerianeaj— the Valerian 127. Apocj-neae 403 Trihe .... ib. 128. Asclepiadeas . . . ib. 106. Dipsacea?— the Teasel Tribe 389 129. Gentianeae — the Gentian 107. Calycereae 390 Tribe . . . . ib. 108. Compositae (See Chap. vi. 130. Bignoniaceas 404 p. 98) . ib. 131. Cobaeaceae 405 109. Lobeliacese 391 jgg' 1 Pedalineas and Sesameae 110. StylideiE . 393 ib. 111. Goodenovise ib. 134. Polemoniaceas . 406 112. Campanulacea?— the Cam- 135. Hydroleaceas . 407 panula Tribe . 394 136. Convolvulaceas . ib. 113. Gesnerieaj 395 137. Boragineae 409 114. Vaccineas (See Chap. vii. 138. CordiacesB . 410 p. 130) . ib. 139. Hydrophylleae . ib. 115. Ericaceffi (See Chap. vii. 140. Solanaces (See Chap. ix. p. 109) ib. p. 141) . ib. 116. Peneaceae . 396 141. Scrophularinae . 411 142. Labiatse 412 § III.— COROLLIFLORJE. 143. Verbenaceae— the Vervain 117 Epaerideae . 396 Tribe . 414 118. Symplocineae . 397 144. Myoporinas . . 415 119. StjTacinese . 398 145. Acanthaceae ib. 120. Myrsineae . . 399 146. Orobancheae . 416 121. Sapotes . ib. 147. Lentibulariae . ib. 122. Ebenaceaj . . 400 148. Primulaceae — the Prim- 123. Brexieas . ib. rose Tribe . ib. 124. Oleinae (See Chap. viii. p. 149. Globularis . 418 136) . 401 150. Plumbagineae . ib. CHAPTER II. Phanerogamous Plants — Dicotyledone^ — II. Monochlamyde^ 419 151. Plantaginea; . . .419 166. Cytineae 429 152. Nyctagineas . , . 420 167. Euphorbiaceae . 429 153. Amaranthaoea? . . ib. 168. StackhouseJE 431 154. Phytolacea^ . . , . 421 169. Antidesmea? . ib. 155. Chenopodea? . . . ib. 170. Urticeae (See Chap. x. p.l57) ib. 156. Begoniaceffi. . . . 422 171. Ulmaceae. 432 157. Polygoneffi— the Buckwheat 172. Piperaceae . ib. Tribe . . . . 423 173. Juglandaces (See Chap. 158. Laurineae— the Sweet-bay XI. p. 176) . ib. Tribe . . . .424 174. Amentaceas (See Chap. xi. 1.09. Myristicese . . . . 425 p. 174) . .433 160. Proteaceaj . , . ib. 175. Hamamelidese . ib. 161. Thymelae^ . . . . 426 176. Empetreas . ib. 162. Osyridea; . . . .427 177. Conifers; (See Chap. xii. 163. Santalaceae . . . . ib. p. 205) . 434 164. Elaeagneae . . .428 178. Cycadeas (See Chap. xii. 165. Asarine^^,o^A^istolochieaE ib. p. 229) . ib. CONTENTS. CHAPTER III. Phanerogamous Plants — Monocotyledone^ § I.— Petaloide.e 179. Hydrocharideas the Frog's-bit Tribe . 435 180. Alimaceae— the Water- plantain Tribe 436 181. Butomeae— the Flowering- rush Tribe . 437 182. Juncagineas — the Arrow- grass Tribe 438 183. Orchidaceffi ib. 184. ScitaminesB 441 185. Canneae . 442 186. Musaces . ib. 187. Iridaceae . 443 188. Hsemodoraceae 445 189. njT)OxideaB 446 190. Amaryllidaccffi . ib. 191. nemerocallidea; 447 192. Dioscoreas . ib. 19.3. Tamacese . 448 194. Smilaceas . ib. 195. Asphodeleae 449 . 435 PAGE 196. Tulipaceffi . . . . 450 197. ISIelanthacess . . . 451 198. Bromeliaceae . . . ib. 199. Pontederaceae . . . 452 200. Commelinese . . . ib. 201. Palmae— the Palm Tribe . ib. 202. Pandaneas . . . . 453 203. Tj'pliineae — the Bulrush Tribe . . . . ib. 204. Aroideffi — the Arum Tribe 454 205. Fluviales, or Naiades — the Pond-weed Tribe . . 455 206. Junceae— the Rush Tribe . 456 207- Gillesieae . . . ib. 208. Restiaceae — the Pipewort Tribe . . . . ib. § IL — GlumacEjE. 209. Cyperaceae — — the Sedge - Tribe . . . .457 210. Gramineae the Grass Tribe . . . . 458 Cryptogamous Plants CHAPTER IV. 460 Sub-Class I— Foliaoe^. 216. ilusci— the Moss Tribe . 466 211. Filices— the Fern Tribe . 461 217. Hepaticas . . .468 212. Lycopodineffi — the Club- moss Tribe 463 Sub-Class IL— Aphylle.e. 213. Marsileaceas 464 218. Lichenes . . . . ib. 214. Equisetaceae — the Horse- 219. Fungi . . . .470 tail Tribe ib. 220. Alga; 471 215. Characeaj 465 NEV/ YOS BOTANIC, GARDG1> MODERN BOTANY FOR LADIES. PART I. INTRODUCTION. 5^ The following pages are intended to enable my readers to acquire a knowledge of Botany with as little trouble to themselves as possible. ; As, however, Botany is a "wide word," I must \ here premise that I only propose to treat of that '•i part of the science which relates to the classifi- % cation of plants, according to the natural system ^ of Jussieu, as improved by the late Professor *^, De Candolle ; and that the grand object I have ^ in view is to enable my readers to find out the V4 name of a plant when they see it for the first \-i time ; or, if they hear or read the name of a N. plant, to make that name intelligible to them. ^"^othing is more natural than to ask the name of •^ every pretty flower we see ; but unless the in- ' " quirer knows something of botany, the name, if "^^ it be a scientific one, will seem only a collection B r) : — i 2 INTRODUCTION. [part r. of barbarous sounds, and will convey no ideas to the mind. Half the interest of new green- house plants is thus destroyed, as few of them have English names, and strangers will soon cease to make any inquiries respecting them w^hen they find they can obtain no answers that they can understand. Now, a very slight know- ledge of botany will take away this mortifying feeling ; and the name of a new plant, and the ascertaining the order to which it belongs, will recall a variety of recollections that will open up a new source of interest and enjoyment even in such interesting and enjoyable things as flowers — for we never can enjoy thoroughly anything that we do not understand. It now only remains for me to say why I have divided my work into two parts. My reason is my belief that a student will always remember more easily a few strongly marked divisions than a number of smaller ones, the differences between which are only faintly perceptible. In a more advanced state of knowledge, it is delightful to trace the minute shades of difference by which the numerous orders are united, so as to form one great whole ; but these gentle gradations confuse a beginner. On this account I have thought it best to devote the first part of my work to a few of the more important orders, which differ most widely from each other, and FAaxi.] PRELIMINARY OBSERVATIONS. 3 \\ hich I have described at a greater length than my space will allow me to bestow upon the whole; and in the second part of my work, I shall give a short account of the whole natural system, in- troducing the orders described in the first part, in their proper places, so that my readers may see how they are connected with the others. MISCELLANEOUS ORDERS. PRELIMINARY OBSERVATIONS. In this first part I shall endeavour to fami- liarise my readers with botanical details, as all the orders I shall describe contain a great num- ber of genera ; and to begin at the beginning, I must first tell them what is here meant by an order, and what by a genus of plants. A genus then may be compared to a family of children, all the plants in it being known by one common or generic name, in addition to their particular or specific one. Thus, if Rosa alba be spoken of, Rosa is the generic name which is common to all roses, but alba is the specific name which is only applied to the white rose. An order includes many genera, and bears the same afiinity to a nation as a genus does to b2 4 PRELIMINARY OBSERVATIONS. [parti. a family. In many cases the resemblance which the plants in each order bear to each other is sufficiently strong to enable the student to re- cognise them^at first sight ; in the same manner as you may generally know a Frenchman or a German from an Englishman, even before you hear him speak. But unfortunately this general outward resemblance does not always exist, and it is necessary for the student to become ac- quainted with the general construction of flowers before the points of resemblance which have occasioned certain genera to be linked together to form orders, can be understood. It is thus evident that the first step towards a knowledge of systematic botany is to study flowers thoroughl}^ and few objects of study can be more interesting, whether we regard the elegance of their forms or the beauty and brilliancy of their colours. My readers may perhaps, however, be as much surprised as I was, to learn that the beautifully coloured parts of flowers are the least important; and that, as they only serve as a covering to the stamens and pistil, which are designed for the production of seedj they may be, and indeed actually are, wanting in a great many of what ^re considered perfect flowers. In examining a flower, there- fore, it must be remembered that the produc- tion of seed is the object, for which all the curi- PART I.] PRELIMINARY OBSERVATIONS. 5 0U3 contrivances we discover are designed. The germen or ovary {a in fig. 1) is protected by a thick fleshy substance (i), called the receptacle or disk, which serves as a bed or foundation on which the other parts of the flower rest, and which is thence frequently called a thalamus or torus, both words signifying a bed. The ovary itself is hollow, and it is sometimes divided into several cells, each in- closing a number of ovules, which are afterwards to become seeds ; but sometimes there is only one cell, and sometimes only one seed in each cell. The ovary is iuicy and succulent fig-i.-stament ,,.,¥. n ^^^ Pistil. when young, and very dmerent from what it afterwards becomes when the seeds are ripe. Rising from the ovary in most flowers, is a long and slender stalk called the style (c), which supports a kind of head, called the stigma (d). The ovary, the style, and the stigma, constitute what is called the pistil ; but the style is not so essential as the other parts, and indeed it is wanting in many flowers. Sometimes there are many styles, each with a stigma at its summit, forming the pistil; and when this is the case, the ovary will have as many cells as there are stigmas, or each stigma will have a separate ovary to itself. There are generally several stamens in a flower. 6 PRELIMINARY OBSERVATIONS. [part i. each perfect stamen consisting of three parts, — the Filament, the Anther, and the Pollen. The filament (e) is, however, often wanting, and it is only the anther {f), and the powder called the pollen which it contains, that are essential. The anther, when the flower first expands, ap- pears like a little oblong case with a deep groove down the centre, or rather like two oblong cases stuck together. When these cases become ripe, they burst and let out the pollen which was in- closed within them. The pollen is generally very abundant, and it is often seen in the form of yellow dust descending from the catkins of the cedar of Lebanon, or the Scotch fir, or of orange powder, as on the stamens of the orange lily, when it sticks to everything it touches. About the time of the bursting of the anthers, the stigma becomes covered with a glutinous moisture, which absorbs the pollen that falls upon it. The pollen, when absorbed by the stigma, is conveyed down the style to the ovary, where it falls upon and fertilises the ovules or incipient seeds. Nothing can be more beautiful or more ingenious than the mechanism by which this process is effected. It is necessary that the grains of pollen should be separated before they reach the ovary, and they are so in their pas- sage down the style in a manner more fine and delicate than could be done by any exertion of PART I.] PRELIMINARY OBSERVATIONS. mere human skill. We know that we ourselves are " fearfully and wonderfully made," but how few of us are aware that every flower we crush beneath our feet, or gather only to destroy, displays as much of the Divine care and wisdom in its construction, as the frame of the mightiest giant ! I have already mentioned that the most con- spicuous part of the flower is merely a cover- ing to protect the seed-producing organs from injury. In most flowers there are two of these coverings, which form together what is called the perianth ; the inner one, when spoken of separately, being called the corolla, and the outer one the calyx. The corolla is generally of some brilliant colour, and in most cases it is Fig. 2. — Corolla of a Flowkr. Fig. 3.— Calyx of Flower. divided into several leaf -like parts called petals, (see ^ in figs. 2 and 3) ; and the calyx, which is commonly green, is divided into similar por- tions called sepals (see h). Sometimes there is 8 PRELIMINARY OBSERVATIONS. [parti. only one of these coverings, and when this is the case it is called by modern botanists the calyx, though it may be coloured like a corolla; and sometimes the calyx and corolla are of the same colour, and so mixed as hardly to be distin- guished from each other, as in the crocus and the tuhp ; in which case the divisions are called the segments of the perianth. CHAPTER I. THE ORDER RAXUNCULACEX : ILLUSTRATED BY THE RANUNCULUS, THE BUTTERCUP OR CROWFOOT, THE PEONY, THE ANEMONE, THE HEPATICA, THE CLEMATIS, THE CHRISTMAS ROSE, THE WINTER ACONITE, MONKSHOOD, THE LARKSPUR, AND THE COLUMBINE. Such of my readers who may have formed their first ideas of the natural system from some order, the flowers of which bear a strong resemblance to each other, will be surprised at reading the names of the heterogeneous assemblage of plants at the head of this chapter ; for surely no flowers can bear less resemblance to each other than the buttercup and the peony do to the columbine and the larkspur. There are, however, striking points of resemblance which link these flowers together ; the principal of which are the number and disposition of the ovaries, or carpels as they are called in this case, which, though they grow close together, and sometimes even adhere to each other, are yet perfectly distinct ; in the number and position of the stamens, which grow out of the receptacle from beneath the carpels ; and in the leaves and 10 THE GENUS RANUNCULUS. [part i. young stems, when cut or pressed, yielding a thin yellowish juice, which is extremely acrid, and, in most cases, poisonous. The flowers of the plants belonging to Ranunculacese difffer widely in their shapes ; and all the incongruities that are only sparingly met with in other orders, are here gathered together. Some of the flowers have only a coloured calyx, as in the clematis ; in others the calyx and corolla are of the same colour, as in the globe-flower, or so intermingled as to seem all one, as in the columbine ; and in others the calyx forms the most ornamental part of the flower, as in monkshood and the larkspurs. In short, modern botanists seem to have placed this unfortunate order first, as though to terrify students on the very threshold of the science, and to prevent them from daring to advance any farther to penetrate into its mysteries. THE GENUS RANUNCULUS. The word Ranunculus will doubtless conjure up in the minds of my readers those very showy, double, brilliantly- coloured flowers, which flower in spring, and are generally grown in beds like tulips. These flowers form a species of the genus, under the name of Ranunculus asiaticus ; and having been introduced from Asia, they have retained their botanic name from not CHAP. I.] THE GENUS RANUNCULUS. 11 having any English one. The honour of giving a name to the genus does not, however, rest on them, but belongs to a common English weed. Every one who has travelled through England in the months of June and July, must have remarked the almost innumerable buttercups which glitter among the long grass of the mea- dows at that season ; and those who observe closely, will have noticed that these brilliant little flowers are never found in poor soil, or in hilly situations, but in rich valleys where the grass is rank and luxuriant from abundance of moisture. It is this circumstance that has ob- tained for the buttercup the botanical name of Ranunculus, the word being derived from Rana, a frog, a creature that delights in moist places. The buttercup being the type of the genus Ranunculus, and the order Ranunculacese, a close examination of its flowers will show the peculiarities which distinguish both the genus and the order. The characteristics of the order, as far as regards the number and position of the carpels and stamens, are shown in the section of the flower in the lower part o'ijic). 4 ; and those of the genus are, a gre^n calyx of five sepals, and a bright coloured corolla of five petals (see a in fig. 4) ; numerous stamens, the anthers of which are adnate, that is, with the filament growing up the back (see }>) ; and numerous carpels ( Peony retains its calyx till the seeds are ripe, while in all the kinds of Ranunculus the calyx Fig. 6.— Flowbr OF the male Peony, with detached carpel AND STAMEN. drops with the corolla. The carpels of the Peony are also many-seeded, while those of the Ranunculus contain only one seed in each. In the male Peony (P. corallina) there are five petals and five sepals, (see a in Ji^. 6,) with nu- merous stamens, forming a ring round four large woolly carpels in the centre of the flower. The stamens (c) are adnate, like those of the Ranunculus ; and the carpels (b) are each ter- minated by a thick, fleshy, hooked stigma. These CHAP. I.] THE GENUS ANEMONE. 19 carpels open naturally on the side when ripe, to discharge their seeds. The herbaceous Peonies with double flowers, now so common in our gardens, have generally only two carpels, each containing about twenty seeds, arranged in two rows ; and the Chinese tree Peony (P. Moutan) has from five to ten carpels, with only a few seeds in each. This last species is distinguished by the receptacle being drawn out into a thin membrane-like substance, which rises between the carpels like the remains of withered leaves, and partially covers them. THE GENUS ANEMONE. I HAVE already mentioned (p. 10) that some of the genera included in the order Ranuncu- laceee have only a coloured calyx and no corolla; and the Anemone is an example of this peculi- arity of construction. The pasque-flower (Anemone pulsatilla) is divided into six dark purple sepals, which are covered on the outside with long silky hairs. The leaves are so much cut as almost to resemble those of parsley ; and at a short distance below the flowers there are three small floral leaves, or bracts, which grow round the stem, and form what is called an in- volucre. The carpels are small, oblong bodies, pressed close together, and each is furnished with a long, feathery point, called an awn. The 20 THE GENUS ANEMONE. [part i. carpels, though lying so close together, are perfectly distinct, and part readily at the slightest touch; and each contains only one seed. It will be seen from this hasty sketch, that the principal point of resemblance between the . genera Anemone and Ranunculus, in a botanical 1 point of view, lies in the carpels, which are close together, and are yet so distinct as to part at the slightest touch. There is, however, a general resemblance in some of the flowers, from their five sepals, and numerous stamens, that renders it difficult for a beginner to distinguish an Ane- mone from a Ranunculus. In many of the British species, also, the carpels are not awned, but slightly curved, very like those of a buttercup. I remember being once very much puzzled with a beautiful little bright yellow flower, that I found in a wood. At first I thought it was a Ranunculus, but the petals were pointed and not roundish ; and it could not be a Ficaria, because it had only five petals. At last I looked to see what kind of calyx it had, and found none, that is, no green calyx ; and then, ob- serving the involucre of three leaves growing in a whorl round the stem, at some distance below the flower, I knew it was an Anemone ; and on comparing it with the ylates in Sowerby's English Botany, on my return home, I ascer- tained that it was Anemone ranunculoides. CHAP. I.] THE GENUS ANEMONE. 21 My readers will therefore observe that Ane- mones may be always known by their involucre, and by their having only one covering (a showy, coloured calyx) to the flower. The number of sepals in this calyx varies in the different species. The pasque-flower has six; the white wood Anemone generally five ; and the Blue Mountain Anemone from twelve to twenty. The in- volucre also sometimes grows a long way from the flower, as in this last-mentioned species ; and sometimes so close to it, as in the Garland, or Poppy Anemone (A. coronaria)^ as to look almost like a green calyx to the flower. The awns, or feathery tails, are also not found attached to the carpels of all the species ; and this distinction is considered so important, that some botanists make those plants which have awned carpels into a separate genus, which they call Pulsatilla, and of which the pasque-flower is considered the type. This genus, however, has not, I believe, been generally adopted. I have now only a few words to say on florists' Anemones, the tuberous roots of which most of my readers must have seen in the seed-shops. Most of these are varieties of the Garland Anemone, already mentioned as having its in- volucre close to the flower. The sepals of this species are roundish, six in number, and when the flower is in a single state, there are a great 22 THE GENUS ANEMONE. [part i. number of stamens, bearing dark purple anthers in the centre of the flower. When the flower becomes double, the sepals, which retain their form and number, only becoming somewhat more spread out and flattened, are called by- florists the guard-leaves ; and the stamens in the centre are metamorphosed into petals, which generally retain their dark purple colour, or at any rate are much darker than the sepals. The other florists"* Anemones spring from A. stellata^ or hortensis^ and they are distinguished by having pointed sepals, and a white spot at the base of each, so as to form a white circle inside the cup of the flower. -The involucre is a long way from the calyx, and when the flowers become double, the sepals can scarcely be dis- tinguished from the metamorphosed stamens. The hepatica or liverwort, the varieties of which look so pretty in our gardens in spring, was formerly considered to be a species of Ane- mone, and indeed the genus Hepatica appears to rest on very slight grounds. It has, however, been adopted by most modern botanists, and the Anemone Hepatica of Linnseus is now gene- rally called Hepatica triloba. The normal form of the species is the single blue ; and the double blue, the single and double pink, and the single and double white, are all only varieties of this. The hepatica agrees^ in all points with the CHAP. I.] THE GENUS CLEMATIS. 23 Anemone, except in the involucre, which is so very like a green calyx, from the manner in which it enfolds the flower in the bud, as scarcely to be distinguished. I could not, indeed, be persuaded that this calyx-like covering was an involucre, till I turned back the apparent sepals, and found that their glossy surface was within : I also found that there was a very small portion of the stem between them and the flower, a circumstance which always distinguishes an in- volucre from a calyx, the latter forming part of the flower, and being always in some manner attached to the receptacle. THE GENUS CLEMATIS. This genus resembles the Anemone in having only one covering, an ornamental calyx, to its seed-producing organs. It has not, however, any distinct involucre ; though in one species, C. calycina^ there are two bracts, or floral leaves, which bear some resemblance to one. The flowers of the different species vary considerably in form, colour, and the number of the sepals ; C. calydna and C. viticella having four, C.fiorida six, C. vitalba five, &c. All the species agree, however, in the seeds, which are produced singly, each in a separate awned carpel, which does not open, but drops with the seed, and is sown with it. These carpels, which are common 24 THE GENUS CLEMATIS. [part I. to the genera Ranunculus, Anemone, Adonis, and many other kinds of Ranunculacese, are called caryopsides, and seeds thus enclosed are always much longer in coming up than any others. In some species of Clematis the awns of the carpels are smooth ; but in others they Fig. 7. — Carpels of the Clematis Vitalba. are bearded or feathered, as in those of the traveller's joy (C vitalha)^ shown in Jir/. 7. The leaves of the Clematis vary considerably in form and arrangement ; but the stems of the climbing species are furnished with tendrils, or slender twining leafless stems, which some botanists sup- pose to be metamorphosed leaves. The plants composing the genus Atragene have been separated from Clematis ; because they are said to have petals, which the genus Clematis has not. It must not, however, be supposed that the petals of the Atragene bear CHAP. I.] THE GENUS HELLEBORUS. 25 any resemblance to what is generally understood by that word. On the contrary, the showy part of the Atragene is still only a coloured calyx ; while the petals are oblong, leaf-like bodies in the centre of the flower, which look like dilated stamens. In other respects the two genera are scarcely to be distinguished from each other. THE GENUS HELLEBORUS, &C. The Christmas rose (Helleborus niger) bears considerable resemblance in the construction of its flowers to the Atragene, for it has a showy calyx, and narrow oblong petals, encircling the stamens in the centre of the flower. The calyx of the Christmas rose is white, delicately tinged with pink, and the petals are green. The carpels are erect and long, swelling out at the base, and each ends in a curved style with a pointed stigma. The Christmas rose takes its specific name of niger (black) from the root, which is covered with a thick black skin. The common Hellebore takes its name of H. inridis, from its flowers, w^hich are green. The carpels of this plant frequently grow slightly together, and their styles curve inwardly. The British species of Hellebore have no in- volucre, and the Christmas rose has only two bracts or floral leaves, which form a calyx-like covering to the bud ; but the little yellow 26 THE GENUS HELLEBORUS. [part r. garden plant, called the Winter Aconite, which was included by Linnaeus in the genus Helle- borus, has a decided involucre, on which the little yellow, cup-shaped flower reposes, like a fairy bowl upon a leafy plate. The conspicuous part of this flower, like the others, is the calyx, which encloses a number of short tubular petals. This little plant is now separated from Helle- borus, and formed into a distinct genus, under the name of Eranthus hyemalis, from its carpels being each furnished with a very short foot- stalk, by which they are attached to the recep- tacle, instead of growing upon it as in the other genera. The root is tuberous, or rather it forms a kind of underground stem, sending up tufts of leaves and flowers from the different buds. Thus we often see several tufts of the Winter Aconite growing so far from each other as to appear distinct ; but which, in fact, all spring from the same root. The Globe-flower ( Trollius europaus)^ which has a golden yellow, globe- shaped calyx, enclosing a number of small oblong petals, is nearly allied to the Winter Aconite ; and the Fennel-flower, or Devil in a Bush (Nigella damascena), agrees with the common Hellebore in the adhesion of its carpels. CHAP. I.] THE GENUS ACONITUM. 27 THE GENUS ACONITUM. We are so accustomed to see in our gardens the tall showy perennial called monkshood or Fig. 8. — Flower and seed-vessels of the Monkshood. wolfsbane {Aconitum Napellus), that few persons think of examining the flowers in detail. They well deserve, however, to be examined, as they are very curious in their construction. The showy part of the flower is an ornamental calyx of six sepals, but the upper two of these are 28 THE GENUS ACONITUM. [part i. larger than the others, and adhere together so as to form a singular sort of covering, Hke a monk's cowl or hood. (See a mjig. 8.) The stamens are numerous, and they encircle three or five oval carpels, with thread-like styles, and pointed stigmas, as shown at h ; which when ripe burst open at the top (c) to discharge the seed, without separating. Carpels of this kind are called follicles. Under the hood, and en- tirely concealed by it, are the petals (seej^-. 9), which form what may certainly be considered the most remarkable part of the flower, as they are so curiously folded up that they look more like gigantic stamens than petals. The older botanists de- scribed these petals as nectaries. Fig. 9.— Petals of ^^yith crcstcd claws. The leavcs are divided into from three to five principal segments, which are again deeply cut into several others. The stem of the common Monkshood is thickened at the base, or collar, where it joins the root, so as to give it some- what the appearance of celery; and hence ignorant persons have been poisoned by eating it. This knotted appearance of the stem is not common to all the species, and it gives rise to the specific name of Napellus, which signifies a little turnip. CHAP. I.] THE GENUS DELPHINIUM. 29 THE GENUS DELPHINIUM. The plants belonging to the genus Delphinium, that is to say, the Larkspurs, have their flowers constructed in nearly as curious a manner as those of the different kinds of Monkshood ; but they differ in the sepals and petals both forming Fig. 10.— Tub flowers of the Branching Larkspur. conspicuous parts of the flower, though they are generally quite distinct both in form and colour, and may be easily traced through all the dif- ferent forms they assume in the various species. They are, however, perhaps most easily distin- 30 THE GENUS DELPHINIUM. [part I. guished in the branching or autumnal Larkspur {Delphinium consolida). In the flower of this plant the spur [a in Jig. 10) is the upper sepal of the calyx, and it serves as a cover to part of the petals. There are four other portions of the calyx (Z>), which assume the appearance of ordinary sepals. The petals are four in number; and they are united at the lower part, and drawn out into a sort of tail, as shown at c ; while the upper part of two of them stands up like asses' ears {d) in the centre of the flower ; and the others are curiously folded, so as to form a hood over the stamens and carpels, as shown at e. The anthers of the sta- mens resemble those of the Ranunculus ; but the fila- ments are bent, as shown at f. The carpels {g) are up- right, hairy, and terminate in a blunt, fleshy stigma (A). When ripe, they open in the same manner as those of the Fig. 11.-TAP hoot of the Moukshood. The branching Branching Larkspur. Larkspur haS a fusiform Or tap root, as shown in Jig. H, in which a is the cHAi-. I.] THE GENUS AQUILEGTA. 31 collar, or as the Italians call it la noda vitale; and h the fibrous roots, through the points of which the plant takes up its food. The flowers of the other kinds of Larkspur resemble this one in their general appearance, though they differ in the minor details. Those of the Rocket Larkspur (Z). Ajacis) lose their spurs when they become double ; and those of the Bee Larkspurs have their petals nearly black, and instead of standing up like ears, they are so curiously folded as to resemble a bee nestling in the centre of the flower. THE GENUS AQUILEGIA. The common Columbine {Aquilegia vulgaris) differs from all the flowers I have yet described in having the sepals and petals not only of the same colour, but so intermingled as to be scarcely distinguishable from each other. The flower (given on a reduced scale at a 'v^fig- 12) is composed of five horn-shaped petals, which are curved at the upper end, and form a kind of coronet round the stem ; and five oval sepals, which are placed alternately with them; all, generally speaking, being of the same colour. The horn-shaped petal, or nectary as it was called by Linnseus, is attached to the receptacle at the thickened rim (Z>), while the sepal is attached at the point (c) ; d shows the dis- 32 THE GENUS AQUILEGIA. [part position of the stamens ; e a separate stamen, with its adnate anther ; f the inner row of Fig. 12. — Flo^ver and leaf of the Columbine. stamens, which are produced without anthers, and with their filaments growing together, so as to form a thin membranaceous case for the carpels, which are shown exposed at g. The carpels, when ripe, become follicles. The leaf of the Columbine is bi-ternate ; that is, it is cut into three large divisions, each of which is cut into three smaller ones ; so that it is twice- ternate. The petiole or foot-stalk of the leaf sheaths the stem, as shown at A, where the leaf is represented on a reduced scale to suit the flower. CHAP. I.] THE GENUS AQUILEGIA. 33 I would advise such of my readers as are anxious to turn the preceding pages to account, to procure as many of the plants 1 have described as possible, and to compare them with each other, and with any other plants belonging to the order Ranunculacese that they can obtain. Those who have access to a botanic garden will have no difficulty in finding the names of the genera included in the order; and those who have not this advantage, must consult Don's edition of Sweet's Hortus Britannicus, or any other catalogue in which the plants are arranged according to the Natural System. When a number of specimens have been col- lected, the student will be surprised to see how many points of resemblance exist between them. The stems of all, when cut, will yield a watery juice ; which is always acrid, though some of the plants are more poisonous than others. The stamens will be found to be always nu- merous, and always attached to the receptacle below the carpels; and the anthers are generally adnate, that is attached to the filaments from one end to the other (see p. 12). The carpels are in most cases numerous, and either distinct, or adhering in such a manner as to show plainly the line of junction between them ; they are also always one-celled, whether one or many- seeded, and generally either caryopsides (see 34 THE GENUS AQUILEGIA. [part i. p. 24), or follicles (see p. 28). -The leaves are generally divided into three or five lobes, each of which is cut into several smaller divisions; and the petioles or leaf-stalks are very frequently dilated at the base, and sheathing the stem. In most cases, the flowers are of brilliant colours, several of them being cup-shaped, and many with the calyx more ornamental than the corolla. The seeds will generally keep good for several years ; and several of them, particularly those of the kind called caryopsides, when sown, are often a long time before they come up. CHAPTER II. THE OP.DER LEGUMlNOSvE : JLLUSTRATED BY THE SWEET-PEA, THE RED CLOVER, ACACIA ARMATA, THE SENSITIVE PLANT, THE BARBADOES FLOWER-FENCE, THE CAROB-TREE, THE TAMARIND, THE SENNA, THE GLEDITSCHIA, THE LOGWOOD, THE JUDAS- TREE, AND THE KENTUCKY COFFEE-TREE. This order is a very numerous one, contain- ing above three hundred genera, and including several highly important plants, both for food and commerce. As examples of the utility of the Leguminosse for food, I need only mention the pea and bean, and all their numerous allies ; and as examples of their importance in medicine and the arts, I may enumerate senna, liquorice, the tamarind, gum-arabic, and logwood. Among the ornamental plants belonging to this order are, the Laburnum, the Furze or Gorse, the Robinia or False Acacia, the true Acacias, the Sensitive Plant, and the Barbadoes Flower- fence. It will be seen by this enumeration, that the flowers of the Lecruminosse differ from each other nearly as much as those of Ranunculacese ; but when in seed, they are all easily recognised by their seed-vessels, which are always legumes, that is, bearing more or less resemblance to the pod of the common pea. To aid the memory D 2 36 PAPILIONACEOUS FLOWERS. [part I. in retaining the great number of genera included in this order, various methods have been devised of re-dividing it ; and of these I shall adopt the newest, which is also the simplest, by which they are arranged in three tribes, according to their flowers. TRIBE L— PAPILIONACEOUS FLOWERS. The flowers of this tribe are called Papilio- naceous ; because Papilio is the sci- entific name of a genus of butter- ilies, which they were supposed to resemble. The type of this tribe maybe considered the flower of the sweet-pea {La- thjrns odoratus)^ which has a small green calyx, cut into five deep notches, but not divided into regu- OF lar sepals. (See a and h in Jig. 13.) The corolla is in five petals, the largest of Fig. 13. — Flower, pod, and tendril the sweet-pea. CHAP. II.] PAPILIONACEOUS FLOWERS. 37 t wliicli (c) stands erect, and is called the vex- illum or standard; below this are. two smaller petals (d), which are called the algae or wings ; and below these are two petals, joined together so as to form a kind of boat (e), which are called the carina or keel, and which serve as a cradle for the stamens and pistil. There are ten stamens, nine of which have the lower half of their filaments growing together, so as to form a fleshy substance at the base, as shown in Ji.^. 14 at/i and the other ((/) is free. The ovary is oblong, terminating in a filiform style, with a pointed stigma, as shown at q in Jlc/. 13 ; and it is one-celled and many- seeded ; the seeds being what we call the peas. When the petals p,^ u.-stamens fall, the pod still retains the calyx oftheSweet-pea. (b), and the style (g) ; and these remain on till the seeds are ripe, when the pod divides natu- rally into two parts, or valves as they are called, which curl back so as to discharge the seeds. If the pod be examined before it bursts, it will be found that the valves are composed of a fleshy substance, lined with a strong mem- brane or skin, and that they are united by two seams, called the dorsal and ventral sutures. Along the ^'^€t«-tFal -suture (A)l there runs a kind of nerve, called the placenta, to which the peas 38 PAPILIONACEOUS FLOWERS. [part i. are attached, each pea being furnished with a little separate stalk, called a funicle. A cook would be surprised, even in these enlightened times, to be told to take a legume of Pisum sativum^ and after separating the two valves at the dorsal suture, to detach tlie funicles of the seeds from the placenta ; yet these scientific terms would merely describe the operation of shelling the peas. It will be seen by this de- scription that the pod of the pea differs very materially from the seed-vessels of all the other plants I have had 'occasion to describe; and that it thus forms a very distinctive character for the order. The other parts vary in the different genera : the calyx is sometimes tubular, and sometimes inflated ; sometimes it has only four notches, or teeth as they are called, instead of five, and sometimes it has five distinct sepals divided to the base. The parts of the corolla vary also in proportion to each other, the keel in some of the Australian plants is as long as the standard ; as, for example, in Kennedia MaryattcB ; and in others the wings arc so small as to be scarcely visible. The stamens of many of the species are also free, that is, divided to the base ; while in others they resemble those of the sweet-pea, in having nine joined together and one free ; and in others the wliole are joined together at the base. The pods also CHAP. II.] PAPILIONACEOUS FLOWERS. 39 vary very iuuch in size and form ; being some- times nearly round, and only one or two-seeded; and in others long, and containing many seeds, as in the common bean or pea. The seeds themselves are so different that the tribe has been divided, on account of them, into two sections : the one consisting of those plants which, like the common bean, have the seed dividing into two fleshy seed-leaves or cotyledons, when it begins to germinate ; and the other, the seed-leaves of which are thin. The seeds of the papilionaceous plants which have thin coty- ledons are not eatable ; but those with fleshy cotyledons may be safely used as food. The fleshy cotyledons do not always rise above the ground ; but they do so decidedly in the bean and the lupine ; and if either of these seeds be laid in moist soil with the hilum or scar down- w^ards, the seed, as soon as it begins to ger- minate, will divide into two parts (that is, into two cotyledons), which wall rise above the ground, and become green like leaves ; though, from still retaining their roundish form, they are easily distinguished from the true leaves, which rise in the centre. Though my readers will have no difficulty in recognising most of the Leguminosse which have papilionaceous flowers, there are some genera, respecting which they may be interested to learn a few particulars. 40 PAPILIONACEOUS FLOWERS. [part i. Thus, the Chorozema is one of the kinds with 'thin cotyledons, and consequently its seeds are not eatable. The legumes of this genus are roundish, and swelled out, so as to bear but little outward resemblance to a pod. Sophora, Edwardsia, Virgilia, Podolobium, Callistachys, Brachysema, Burtonia, Dillwynia, Eutaxia, Pultensea, Daviesia, and Mirbelia, have all thin cotyledons, and their ten stamens all separate from each other ; but in Hovea, Platylobium, and Bossisea, though the cotyledons are thin, the stamens all grow together at the base. I mention these common greenhouse shrubs, that my readers may have an opportunity of ex- amining their botanical construction, and thus verifying their names. The common furze ( Ulex europceiis)^ the Spanish broom {Spartium jun- ceiim), the Petty whin {Genista Anglica)^ the Laburnum ( Cy^?sw5 Lahurnum)^ and the common broom, all belong to this division, and con- sequently their seeds are not eatable ; those of the Laburnum are indeed poisonous. The distinctions between Spaitium, Genista, and Cytisus, are very slight, lying chiefly in the calyx; and as a proof of this the common broom, which is now called Cytisus scoparius, was formerly supposed to be a Spartium, and afterwards a Genista. The common red clover {Trifolium pratense) CHAP. II.] MIMOSA. 41 has its flowers in such dense heads that it is difficult at first sight to discover that they are Papihonaceous. On examination, however, it will be found that each separate flower has its standard, wings, and keel, though the wings are so large as to hide the keel, and nearly to obscure the standard. The calyx is tubular at the base, but divided above into five long, awl- shaped teeth, that stand widely apart from each other. The legume has only one or tvro seeds, and it is so small as generally to be hidden by the calyx. TRIBE II.— MIMOSA. The second division of Leguminosse com- prises those plants w^hich have heads of flowers either in spikes or balls, like those shown in Jig. 15. This figure repre- sents two heads of flowers oi Acacia ar- mata, a well-known greenhouse shrub, of their natural size ; and Jig. 16 shows a head of similar flowers mag- fig. is.-flowers ast> sprig of • o 1 T J 1 ^ , Acacia armata . nined. In the lat- ■^i;J^ 42 MIMOS.E. [part I. ter, a shows the calyx, which is five-toothed, and h the petals, which are five in number and %a\.. c quite regular in shape; c are the >l'^||l?fj^1|, stamens, which vary from ten to •■^ two hundred in each flower, and which are raised so high above the petals as to give a light and tuft- like appearance to the whole flower. iG. i6.-flo\ver Yi\Q leg^umes are very large in pro- OF Acacia iMAG- ^ ^ j ), which alternate with the petals of the corolla (c). There kkntl'cky are ten stamens, but they are completely enclosed in the tube of the calyx. The pod is very large, the valves becoming hard and bony when dry ; and the seeds are like large beans, the pod being deeply indented between the seeds. The leaves are bi-pinnate, with from four to seven pairs of pinnse ; the lower having only one small leaflet, but the rest bearing from six to eight pairs of leaflets each. This tree mu^t not be confounded with the true Coffee-tree, which belongs to Rubiaceae, and from which it is 19.— Flowers of thi coffke-tree. CHAP. II.] CiESALPINE^. , 49 perfectly distinct in every respect ; and it only takes its American name from its beans having been used as a substitute for coffee. The outer bark of this tree, when it becomes old, splits off in narrow strips and rolls up ; and its timber, like that of the Robinia or False Acacia, having very little sap wood, is thus very strong in quite young trees, though it is of little value when the tree is full-grown. The species contained in the first and second divisions of this order will be easily recognised by botanical students ; and though those of the third division are much more difficult to find out, still there is a kind of family likeness, particu- larly in the leaves, which will enable the eye, with a little practice, to recognise them. The student should visit the hothouses of botanic gardens and nurseries, and should there endea- vour to pick out plants belonging to this order. 50 ROSACEA. [part i. CHAPTER III. THE ORDER ROSACEA, ILLUSTRATED BY DIFFERENT KINDS 0¥ ROSES ; THE POTENTILLA ; THE STRAWBERRY ; THE RASPBERRY ; SPIR.SLA ; KERRIA OR CORCHORUS JAPOMCA; THE ALMOND; THE PEACH AND NECTARINE ; THE APRICOT ; THE PLUM ; THE CHERRY ; THE APPLE ; THE PEAR ; THE MOUNTAIN ASH ; THE WHITE BEAM TREE ; QUINCE ; PYRUS OR CYDONIA JAPO- NICA ; THE HAWTHORN ; THE INDIAN HAWTHORN ; THE MEDLAR ; PHOTINIA ; ERIOBOTRYA ; COTONEASTER ; AMELAN- CHIER ; BURNET ; AND ALCHEMILLA OR LADIES'-MANTLE. All the numerous plants which compose this large order agree more or less with the rose in the construction of their flowers, though they differ widely in the appearance of their fruit. They all agree in having the receptacle dilated, so as toformaliningtothelower part of the calyx, and in the upper part of this lining the stamens and petals are inserted above the ovary ; and the anthers are innate, that is, the filament is inserted only in the lower part. The leaves also have generally large and conspicuous stipules ; and they are frequently compound, that is, composed of several pairs of leaflets, placed exactly opposite to each other ; though the leaves themselves are never opposite to each other, but are placed alternately on the main stem. These CHAP. III.] ROSE^. 51 characters are common to the order; but the plants included in it differ from each other so much in other respects, that it has been found necessary to redivide Rosacese into tribes, of which the following six contain plants com- mon in British gardens. TRIBE I.— ROSE^. The flowers of the wild Rose have the lower part of the calyx tubular and fleshy (from being lined with the dilated receptacle) and the upper part divided into five leafy sepals, which enfold Fig, 20.— Rosa Fosteri. the bud, and remain on after the expansion of the corolla. In Rosa Fosteri^ (see Jig. 20,) and E 2 52 ROSE^. [part I. its near ally the Dog rose(i?. canina), the sepals (a) do not extend far beyond the petals of the bud ; but in some species, as in Bosa cinnamonea and its allies, the sepals are so large and long, that they assume the character of little leaves, The corolla is cup-shaped, and it is composed of five equal petals, each of which is more or less indented in the margin, as shown at h. In the centre of the flower the receptacle forms a kind of disk which completely fills the opening or throat of the calyx ; in most species covering the carpels and their styles and onlyleaving the stigmas free, though in the Ayrshire rose {R. arvensis), and its allies, the styles are united, so as to form a column, which projects considerably above the disk (seejir/. 21). The pitcher - shaped part of the calyxwhen the corolla falls be- comes the hip (Ji[/. 20 c), and serves as a covering or false pericarp to the nu- merous bony carpels or nuts which contain the seed. These nuts are each enveloped in a hairy cover (see Jlf/, 20 d, and Ji^. 21 a,) and each contains only one seed which it does not open naturally to discharge : hence, the seeds of roses when sown Fig. 21.— Ovary of the Ayrshire ROSE WITH A DETACHED SEED. CHAP. HI.] ROSE^. 53 are a long time before they come up. Fig. 22 is the ripe fruit of Rosa cumamonea, cut in two to show the nuts. The leaves are pinnate, consisting of two or more pairs of leaflets, and ending with an odd one. The leaves are furnished with very large stipules (see Jig. 20 e) ; and the stems have numerous prickles fjg. 22.— ripe fruit (/), which differ from thorns in and detached seed being articulated, that is, they may be taken off without tearing the bark of the stem on which they grow, only leaving the scar or mark, shown at g. The leaves of the sweet briar are full of small glands or cells filled with fragrant oil, w^hich may be distinctly seen in the shape of little white dots, when held up to the light ; and this is the reason of their delightful perfume. When the leaf is rubbed between the fingers, the thin skin that covers the cells is broken, and the oil being permitted to escape, the fragrance is increased. There are only two genera in this tribe, viz. Rosa and Loicca^ the latter containing only what was formerly called Rosa herherifolia^ and which has been thought worthy of being made into a separate genus principally on account of its having simple leaves without stipules, and branched prickles. 54 POTENTILLE^, OR [part TRIBE II.— POTENTILLE^ OR DRYADEiE. The plants belonging to this tribe agree more or less in the construction of their flowers with the well-known showy plants called Potentilla, but my readers will probably be surprised to hear that the raspberry and the strawberry are included among them. If, however, they com- pare the flower of the Potentilla with that of the strawberry, they will find them very much alike. In both there is a calyx of ten sepals, and a cup-shaped corolla of five petals ; and in both the stamens form a ring round an ele- vated receptacle, on which are placed numerous carpels. Here, however, the resemblance ceases, for as the seeds of Potentilla ripen, the recep- tacle withers up in proportion to the swelling of the carpels, till it becomes hidden by them; while in the strawberry the receptacle becomes gradually more and more dilated, swelling out and separating the bony carpels still farther and farther from each other, till at last it forms what we call the ripe fruit. I have already had several times occasion to mention the receptacle, which though seldom seen, or at least noticed, by persons who are not botanists, is a most important part of the flower, and one that assumes a greater variety of form than any CHAP. III.] DRYADE^. 55 other. Sometimes, as we have seen in several of the Ranunculaceae and Leguminosae, it is a mere disk or flat substance serving as a founda- tion to hold together the other parts of the flower ; and at other times we have found it drawn out into a thin membrane and divided into a kind of leaves, as it is among the carpels of the tree-peony; but in no plants that I have yet had occasion to describe does it assume such strange forms as in Rosacese. The flower of the strawberry {Fragaria vesca) has a green calyx of ten sepals ; five of which are much smaller than the others, and grow a little behind them, the large and small ones occurring alternately. The corolla is cup-shaped, and in five equal petals ; the stamens are numerous and arrano^ed in a crowded rino^ round the carpels, which are placed on a somewhat raised receptacle. The carpels or nuts resemble those of the rose, but they have no hairy covering, and indeed look hard and shining on the surface of the distended receptacle, or poly- phore as it is called in its metamorphosed state. The carpels when ripe do not open to discharge the seed, and consequently as they are sown with the seeds, the young plants are a long time before they appear. The strawberry has what is called ternate leaves, that is, leaves consisting of three leaflets ; with large membranous stipules. 56 POTENTILLE^, OR [part I. The calyx is persistent, that is, it remains on till the fruit is ripe. The Raspberry {Ruhns IdcBus) differs widely from the strawberry in many particulars, not- withstanding their being included not only in the same natural order, but in the same tribe. The ca-lyx has only five se- pals {a in fi- gure 23) ; and though the co- rolla has five petals (^), they do not form a cup-shaped flower. In the centre are the carpels, the Fig. 23.— Flowers and fruit op the Raspberry. fu„^y> ^f wliipli is shown of the natural size at c, and magnified at d^ the latter showing that each has a separate style and stigma. As the raspberry advances, the petals drop, and the receptacle becomes elevated into what is called a torus, as shown of the natural size at e ; bearing the carpels upon it, which gradually swell out and soften, till each becomes a little pulpy fruit, CHAP. III.] DRYADE^. 57 full of juice, and having the stone or seed in the centre. While this change is taking place, the stamens gradually wither and fall off, and the stigmas disappear, the style shrivelling up to the appearance of a hair ; the pulpy carpels have also become so pressed against each other, as to adhere together, and the whole, with the persistent calyx, now assumes the appearance shown at f. As soon as the carpels become ripe they cease to adhere to the torus, and they may be pulled off and eaten (the torus, or core as it is called, being thrown away) : each carpel will be found to inclose a very hard seed or stone, as shown at^. If the Raspberry, instead of being gathered, be suffered to remain on the stalk, the juicy carpels dry up, and fall with the seed inclosed. The stems of the Raspberry are bien- nial, that is, they do not bear till they are two years old, after which they die ; but the roots are perennial, and they are always sending up fresh suckers, so that the same plants will bear for many years in succession, though not on the same stems. The stems are generally erect, and prickly like the rose ; and the leaves on the bear- ing stems have three leaflets, while those on the barren stems have five ; and in both cases the leaflets are covered with white down on the under side. All the different kinds of Bramble, such as the Dewberry, Blackberry, &c., agree 58 SPIRiEEjE. [part i. with the Raspberry in the construction of their fruit, though they differ in the number of their leaflets, the size and colour of their flowers, and other minor particulars. Several other genera belong to this tribe, among which may be mentioned Geum Aveiis^ or Herh Bennet^ the carpels of which have each a hooked style; Sieversia separated from Geum, because the carpels end in a straight feathery awn ; and Tormentilla^ the flowers of which bear a general resemblance to those of Potentilla, but which have an eight-parted calyx; a corolla of four petals; sixteen stamens, and dry wrinkled carpels on a depressed receptacle. All these genera my readers will find it interesting to procure flowers of, in order to compare them with each other. This and the preceding tribe are considered by some modern botanists to form the order Rosacese ; the other tribes being formed into separate orders. TRIBE III.— SPIRiEE^. The only genera in this tribe which contain well-known plants are Spiraea and Kerria. In Spirsea the calyx is five-cleft (see a in Jig, 24) and lined with the dilated receptacle, forming a shallow tube or rather cup for the reception of the carpels. There are five small roundish CHAP. III.] SPIR^E^. petals (b), and from twenty to fifty stamens (c), which project very far beyond them. In the Fig. 24. — Flower of the Spir.^a. centre are from two to five carpels (d), which are something like those of the raspberry when young, but afterw^ards become of the kind called follicles ; each carpel contains from two to six seeds affixed to its inner suture, and they are dehiscent — that is, they open naturally at the top to discharge the seed (see e). The flowers are set very close together, and from this circum- stance, combined with their small size and pro- jecting stamens, they look like fine filigree work; hence the popular English names given to S. salicifolia or Bride wort, Queen^s needle- work, &c. The flow^ers of this species are in spicate racemes, but others are in corymbs, as in S. bella ; or in panicles, as in S. armfolia. Kerria is a genus containing only one species, the plant which was formerly called Corchorus J-ponica; the calyx is united at the base, but divided in the upper part into five lobes ; three of them obtuse, and the other two tipped with 60 AMYGDALE^. [part i. a little point called a mucro. There are about twenty stamens about the same length as the petals arising from the calyx, and five roundish carpels containing one seed each. The leaves are simple, and the stipules awl-shaped. Till lately only a double-flowered variety was known in Britain; but about 1832, the single-flowered plant was introduced from China. Corchorus, the genus in which this plant was originally placed, is nearly allied to the lime-tree. TRIBE IV.— AMYGDALEiE. This tribe is distinguished by the fruit, which IS what botanists call a drupe, that is, a stone fruit. The principal genera included in this tribe are Amygdalus^ the Almond ; Persica, the Peach and Nectarine ; Anneniaca, the Apricot; Primus^ the Plum: and Cerasus, the Cherry. All these genera contain more or less of prussic acid, which is found to exist principally in the leaves and kernels ; and they all yield gum when wounded. The flowers of the common Almond {Amyg- dalus communis) appear, as is well known, before the leaves, bursting from large scaly buds, which when they open throw off the brown shining bracts in which they had been enwrapped. The calyx is somewhat campanulate, wdth the upper CHAP. III.] AMYGDALE^. 61 part cut into five teeth or lobes, and it is lined by the dilated disk. There are five petals, and about twenty stamens, both inserted in the lining of the calyx. The anthers are innate, and they differ from most of the other plants yet described in being only one-celled. The ovary is also only one- celled, and there are generally two ovules, though the plant rarely ripens more tlian one seed. The leaves are simple, and they have very small stipules. When the petals drop, the ovary appears covered with a thick tough downy pericarp, within which is the hard stone or nut, the kernel or almond of which is the seed. The Peach (Persica vidians) was formerly included in the same genus as the almond ; and in fact there is but little botanical difference. The flow^ers are the same both in construction and appearance ; and the leaves are simple like those of the almond, and, like them, they are conduplicate (that is, folded together at the mid- rib) when young. The only difference indeed is in the fruit ; for, as everybody knows, the stone of the peach has not a dry tough covering, like that of the almond, but a soft and melting one full of juice, and the stone itself is of a harder consistence, and deeply furrowed, instead of being only slightly pitted. The fruit of the peach has thus a fleshy pericarp, the pulp or sarcocarp of 62 AMYGDALE^. [part i. which is eatable, and a furrowed nut or stone, inclosing the seed or kernel, which is '^Tapped up like that of the almond, in a thick loose skin. The Nectarine (P. Icevis) only differs from the peach in the epicarp, or outer covering of the pulpy part, being smooth instead of downy. Of both fruits there are two kinds, one called free-stone, from their parting freely with the stone ; and the other cling-stone, from the stone clinging to the fibres of the pulp. The Apricot (Armeniaca vulgaris) agrees with the preceding genus in its flowers ; but it differs in its fruit, its stone being sharp at one end and blunt at the other, with a furrow on each side, but the rest of the surface smooth. Thus my readers will perceive that the Peach and the Apricot, though so different from each other as to be recognised at a glance, are yet botanically so very closely allied, as to be dis- tinguished only by the stone. The leaves indeed differ in form, but in other respects they are exactly the same. The Sloe (Prunus spinosa) is supposed by some botanists to be the origin of our cultivated plum, though others make it a separate species under the name of Prunus domestica. The flowers in both are solitary (see Jig. 25), and consist of a five-toothed calyx (a) which is united at the base, and in the lining of which the CHAP. III.] AMYGDALEiE. 63 stamens are inserted as shown at h. The ovary has a thick style and capitate stigma (c), Fig. 25. — Flowers and fruit of the Sloe. and the fruit is a drupe {d). In these particulars therefore the plum agrees with the preceding genera ; but it will be found to differ in the skin of the pericarp, which is quite smooth and covered with a fine bloom ; this, indeed, and its stone being pointed at both ends constitute the chief botanical distinctions between the fruit of the plum and that of the apricot, as in other respects they are alike. Both the plum and the apricot have footstalks, and in this differ from the peach and the nectarine, which are without. The leaves of the plum differ from those of the other genera in being convolute, that is, rolled up, in the bud. 64 AMYGDALE^. [part I. The Cherry {Cerasus vulgaris) differs from the plum in the skin of the pericarp being destitute of bloom, and in several flowers springing from each bud, in what botanists call a fascicled umbel (see a in fg._ 26). The pedicels (5) are also much longer; the pe- tals (c) are in- dented in the margin ; the style {d) is more slender ; and the stone (e) is smooth and much more globose. The number of the stamens, and the manner in which they are FxG. 26.- -Flowers ant) stone of the Cherry. inserted in the lining of the calyx, is the same in both genera (seey)'; but the leaves are different, for those of the Cherry are folded down the middle, when young, like those of the peach and almond ; while those of the plum are rolled up. CHAP. III.] POME^. 65 The genus Cerasus is divided into two sections, the first containing those species which have their flowers in bunches, and on long footstalks, as in the common Cherry ; and the second those which have their flowers in racemes on short footstalks, as in the Bird- cherry {Cerasus Padus) ; the Mahaleb, or Bois de Sainte Lucie {Cerasus Mahaleb) ; the common Laurel {Ce- rasus Lauro-Cerasus) ; and the Portugal Laurel {Cerasus lusitanicus) , These plants are so dif- ferent from the common Cherry both in flowers and fruit, as far as can be judged from their general appearance, as scarcely to be recog- nised ; but when closely examined their bo- tanical construction will be found the same. Formerly only two genera were included in this tribe — viz. Amygdalus, which comprised the Peach and Nectarine as well as the Almond ; and Prunus, w'hich included the Apricot and the Cherry. TRIBE v.— POMEtE. The common apple {Pyrus Malus) may be considered the type of this tribe, which com- prehends not only what we are accustomed to call kerneled fruit, but also the Hawthorn, Cotoneaster, and other ornamental shrubs and low trees. The flower of the apple bears con- 66 POME-^. [part I. siderable resemblance to the flowers of the genera already described, but the petals (see a mjig. 27) are oblong, rather than roundish. Fig. 27. — Fruit and part of the Flower of the Apple. The calyx {h) is tubular in the lower part, and the limb is divided into five lobes. The receptacle lines the lower part of the calyx, and forms a disk, filling its throat, in which the stamens and petals are inserted. There are five ovaries, the styles of which are for half their length united, leaving the upper part and the stigmas free ; and the ovaries themselves, now become cells, are enclosed in a cartilaginous endocarp, which forms what we call the core of the Apple, and which adheres firmly to the tubular part of the calyx. There are two ovules in each cell, placed III.] POME^. 67 side by side, but generally only one seed in each becomes perfectly ripe. As the seeds advance, the fleshy tube of the calyx swells out and be- comes what we call the apple ; while the leafy part or lobes of the limb remain on, and form the eye. Fruit of this kind are called pomes. The Pear (Pi/rus communis) differs from the apple in the shape of the fruit (see a in Jir/. 28), Fig. 28.— Fruit axd part of the Flower ov the Pear, which tapers towards the footstalk, instead of being umbilicate, that is, indented at the point of the insertion of the footstalk, as is the case with the Apple. The construction of the flowers in both species is the same, except that the styles are quite free for their whole length in the Pear, and not partially united into a column F 2 68 POME^. [part I. as in the Apple. This distinction, and some others, have been thought, by some botanists, sufficient to constitute the Apple and its allied species into a separate genus under the name of Mains. The leaves of the Pear differ from those of the Apple in being the same colour on both surfaces, whereas those of the Apple are covered with a white down on the under side. Besides the Apple and the Pear, and their respective allies, which form two distinct sections of the genus Pyrus, that genus, being a very extensive one, is divided into several other sections, all the plants contained in which may be arranged under two heads : viz., those that formerly constituted the genus Sorbus ; and those that were once called Aronia. The Mountain Ash {Pyrus aucvparia) may be considered as a fair specimen of most of the trees belonging to the Sorbus division. By the details of the flowers of this species given in Jig. 29, it will be seen that the petals {a) Fig. 29. — Flower and fruit of the Mountain Ash. are very small and concave ; and the calyx (h) CHAP. III.] POME^. 69 is tubular, and five-cleft. There are three styles, as shown at c ; and the stamens (cf), which project far beyond the petals, are inserted in the disk. The fruit (e) is a pome with three seeds (/) enclosed in a cartilaginous membrane, like the core of the apple or pear. The leaves of the Mountain Ash are impari-pinnate, that is, they consist of several pairs of leaflets, ter- minating in an odd one; and the flowers are produced in corymbs. The White Beam-tree {Pyrus Aria), the wild Service (P. torminalis), and several similar trees, belong to this division and have the same kind of fruit as the Moun- tain Ash. The true Service, however, differs in its fruit being generally shaped like a pear, though there is a variety with apple-shaped fruit. One species {P. pinnaiijidd) has the leaves lobed to the midrib, instead of being cut into leaflets ; and this gives the name to the species, leaves of this description being called pinnatifid. The leaves of the genus Pyrus often have their petioles dilated and somewhat stem-clasping at the base ; but they have gene- rally only small stipules. Among the other plants included in the genus Pyrus, may be mentioned the beautiful shrub now called Pyrus arhutifolia, which has been successively included in the genera Crataegus, Aronia, and Mespilus; and P. Chamcemespihis, 70 POME^. [part I. which has been successively called Crataegus, Mespilus, and Sorbus. There are several beau- tiful low shrubs belonging to this division of the genus Pyrus. The genus Cydonia, the Quince, differs from Pyrus in having its seeds arranged in longi- tudinal rows, instead of being placed side by side. In the Chinese Quince there are thirty seeds in each row, arranged lengthways of the fruit. The ovary of this genus consists of five cells, each containing one row of seeds, the seeds being covered with a kind of mucilaginous pulp. The well-known plant, formerly called Pyrus japonica^ has been removed to the genus Cydonia on account of its ovary and the dis- position of its seeds, which are decidedly those of the Quince. It differs, however, from the common Quince in its seeds, which are arranged in two rows in each cell. The common Hawthorn (CratcBgus Oxyacantha) hasgenerally only two styles (see «, Jig. 80) , but the number of styles varies in the many different species included in the genus from one to five. The corolla, ca- FiG. 30.— Flower and fruit of the lyX, and stamcnS Hawthorn. , , are the same as CHAP, iii.l POME^. 71 in the other genera inchided in this tribe, but the petals {b) are rounder and rather more in- dented. The seeds vary from one to five, each being enclosed in a bony covering, or stone, the whole being surrounded by the fleshy part of the calyx, which forms the eatable part of the Haw. In some of the species the haws are so large as to appear like little apples ; but they may be always easily distinguished by the ripe ovary, or case which incloses the seed, being bony ; whereas in all the varieties of Pyrus, the outer part of the ovary is cartilaginous, like the core of the apple. The seeds of the Hawthorn are a long time before they come up, from the hardness of this bony covering, which does not open naturally when ripe. The species com- posing the genus Raphiolepis, the Indian Haw- thorn, have been separated from Cratsegus ; chiefly on account of the covering which en- closes the seeds being of a paper -like texture, instead of bony, and each cell containing two seeds. The hmb of the calyx also falls off" before the fruit is ripe, instead of remaining on to form what is called an eye, as it does in the common Hawthorn. The leaves of the plants belonging to this genus vary in the different species ; but those of the common Hawthorn are wedge- shaped, and cut deeply into three or five lobes. The different species which compose the genus 72 POME^. [part I. Cratsegus were formerly considered to belong to the genus Mespilus. This genus, which is now almost confined to the common Medlar {Mespilus germanica)^ agrees with Crataegus in having each seed enclosed in a bony covering, but it differs in the limb of the calyx being in large leafy segments ; and in the disk being very large and visible even^vhen the fruit is ripe, from the tubular part of the calyx not closing over it. Among the plants formerly included in the genus Mespilus, may be mentioned Photinia serrulafa, and Eriohotrya japonica, both natives of Japan. The first of these was once called Cratcegus glabra^ and it is remarkable for its beautiful glossy leaves, which are of a deep green when old, and beautifully tinged with red when young ; the flowers are white, and they are produced in what botanists call corymbose panicles. There are some other species of the genus Photinia, but only two or three are com- mon in British gardens. Eriohotrya japonica^ the Loquat-tree, was formerly called Mespilus japonica. It is remarkable for its large and handsome leaves, which are woolly on the under side. The flowers, which are small and white, are produced in large panicles, and they are followed by large pendulous bunches of the yellow pear-shaped fruit, which is covered with CHAP. III.] SANGUISORBE^. 73 a woolly substance, and hence the botanic name Eriobotrya, which signifies woolly grapes. The tree will stand out in the open air in England, and it will flower freely in a green- house, but it requires a stove to ripen its fruit. Cotoneaster and Amelanchier were also for- merly included in Mespilus, and they are very closely allied to Photinia and Eriobotrya. The species belonging to Photinia, however, are easily known by their shining leaves, and the petals of their flowers being reflexed, that is, curved back ; and the species of Eriobotrya are distinguished by their woolliness, which spreads over even the flowers and fruit. The Coto- neasters are known by the small petals of their flowers, which curve inwards, and remain a long time without falling. The leaves are also thick, and w^oolly or clothed with rusty hair on the under side ; and the flowers, which are pro- duced in cymes or panicles, with woolly pedi- cels, are followed by bright red havrs, resem- bling those of the hawthorn. Lastly, the genus Amelanchier is known by its long narrow petals, and its ovary having five or ten cells, with five styles united at the base. TRIBE VI SANGUISORBE^E. The plants included in this tribe agree more or less with the common Burnet {Sanguisorba 74 SANGUISORBE^. [part i. officinalis). This plant, which is found in great abundance in rich meadows on calcareous soils, has its flowers produced in a close terminal spike. The flowers have no petals, but the calyx, which is four-cleft, is pink, and there are four glossy brown bracts to each flower ; so that, on the whole, the flowers are rather ornamental, notwithstanding their want of petals. There are only four stamens, and two carpels with slender styles and pointed stigmas. The leaves are pinnate, consisting generally of nine leaflets, and each pair of leaflets is furnished with two stipules. The Alchemilla, or Ladies' Mantle, is nearly allied to the Burnet ; but the flowers are in small corymbs, instead of spikes. The flowers have no petals ; but the limb of the calyx is coloured, and divided into eight un- equal segments. There are generally four stamens and only one style, though sometimes there are two. The ovary contains one or two carpels, each containing a single seed, and these when ripe are enclosed in a capsule, formed by the tubular part of the calyx becoming hardened. The leaves are lobed, plaited, and serrated at the margin ; and those of the Alpine species {A. alpina), which is often found wild on the Scotch mountains, are covered with a beauti- ful silky substance of the most brilliant white- ness. CHAP. IT.] THE GENUS FUCHSIA. CHAPTER IV. THE ORDER ONAGRACE^ : ILLUSTRATED BY THE DIFFERENT KINDS OF FUCHSIA ; CENOTHERA, OR THE EVENING TREE- PRIMROSE ; GODETIA ; EPILOBIUM, OR THE FRENCH WILLOW- HERB ; AND CLARKIA. The type of this order is considered to be the common evening Tree-primrose {CEnothera bien- nis), and it takes its name from Onagra^ the name given by Tournefort to the genus. The Fuchsia seems so unlike the CEnothera, that it appears difficult to any but a botanist to trace the connexion between them ; but, botanically, they agree in the position of the ovary, which in both is so placed as to seem rather to belong to the flower-stalk than to the flower ; and this peculiarity is found in all the genera included in the order. The parts of the flowers are also always either two, four, eight, or twelve ; as, for example, there are four petals and eight stamens in both the Fuchsia and the CEnothera. THE GENUS FUCHSIA. Little more than fifty years ago, the first Fuchsia was introduced into England ; and we are told that small plants of it were sold at 76 THE GENUS FUCHSIA. [parti. a guinea each. Now more than twenty species, and innumerable hybrids and varieties, are in common cultivation, and we find them not onl}^ in greenhouses and windows, but planted in the open air as common border shrubs. The first Fuchsia seen in England was F. coccinea^ intro- duced in 1788 ; and this species is still common in our gardens. It was followed about 1796 by F. lycoides ; and after that no other species was introduced till 1821, since when a full tide of Fuchsias has kept pouring in upon our gardens, from the different parts of Mexico, South America, and New Zealand, to the present time. All the Fuchsias were formerly divided into two sections ; the plants in one of which having the stamens and pistil concealed, and those in the other having the stamens and style exserted, that is, projecting beyond the other parts of the flower. The first division comprises all the small-flowered kinds ; such as F. microphylla^ thymifolia^ cylindracea, and hacillaris, all which have the lobes of the calyx short, and the petals partially concealed. F. parviflora belongs to this division, but it is distinguished by its glau- cous leaves with an entire margin; and F, lycoides is also included in it ; though this last seems to form the connecting link between the two sections, as both its petals and its style and sta- mens are partially exposed. The second division CHAP. IV.] THE GENUS FUCatSiA. 77 comprises all the kinds which have long project- ing stamens. As the general arrangement of the parts of the flower is nearly the same in both divisions, jig. 31, which represents the section of a flower of F. cylindracea, from the Botanical Register^ will give my readers a clear idea of the botanical construction of the Fuchsia. In this figure, a shows two cells of the ovary (which when entire is four-celled, opening when ripe into four valves), with the seeds attached to a central placenta. This ovary is surrounded and protected by the di- lated disk, which also serves as a lining to the tubular part of the calyx, h. The an- thers, in this division, have very short fila- ments, which are inserted in the hning of the calyx, as shown at c ; ^ is the style, which, in fact, consists of four styles united together, and which divides near the apex into four stigmas ; e e are two of the four lobes of the calyx ; and f is one of the four petals. In the second division, of which F. coccinea Fig. 31. — Section of the flowek op- Fuchsia CyiilNDRACEA. 78 THE GENUS FUCHSIA. [part I. may be considered the type, the calyx and the corolla are of different colours. In Jig. 32, which shows a flower of F. discolor, the Port Famine Fuchsia, the calyx {a) is scarlet and the most ornamental part of the flower, while the petals (h) are purple, and wrapped over each other. The ovary (c) is green, and when the petals and calyx fall ofl", it swells into a berry, which becomes of a dark pur- ple when ripe. F. glohosa differs from F. coccinea in the flowers being shorter and more globose, while the limb of the calyx curves inward. In F. macrostemma^ a well- known Fuchsia, the lobes of the limb of the calyx are, on the contrary, recurved, that is, turned backwards. This formation is common, more or less, to several other species. In F. excorticata, the New Zealand Fuchsia, there is a large fleshy knot at the base of the calyx, and strong ribs running up the lobes ; the calyx is green when young, but it afterwards becomes crimson ; and the petals are very small. This species is so different from the others, that it was at first described as a new genus, under the name of Skinnera. The Fig. 32.— Fuchsia discolor. CHAP. IV.] THE GENUS (ENOTHERA. 79 calyx is green at first, but it afterwards becomes crimson, jp. arhorea has pale-purplish flowers, and, like F. hjcoides^ forms a connecting link between the two sections, the stamens being only a little exserted, and the petals hidden. F. radicans, the only Fuchsia yet discovered with a creeping stem, which was introduced in .1841, belongs to this division. These sections include all the Fuchsias known in British gardens previously to 1835 ; but since that period, two kinds have been introduced, which belong to a third division. These are F.fulgens and F. corymhiflora. In these plants the tube of the calyx is about two inches long, and the lobes are very short. The petals are also short, and scarlet or deep-rose colour, though not exactly of the same hue as the calyx. The leaves are large, with the midribs ajid veins red ; and the branches and pedicels are also of a dark reddish purple. THE GENUS GENOTHERA. In the description of the botanical construction of the Fuchsia, my readers may have observed, that the ovary is placed below the calyx, and quite distinct from it. The same construction is still more visible in the CEnothera, as the tube of the calyx is very slender, and often more than two inches long, while the ovary is often vase- THE GENUS (ENOTHERA. [part shaped, and of large size. The cal3'x of CEno- thera biennis^ the common Evening or Tree Primrose, consists of four sepalsgrowing together in the lower part, so as to form a long tube (a v^fy. So), and with tlie upper part or limb gene- rally in two segments (^), which are bent quite Fig. 33. — The Evexin'g Primrose (CEnoUicra biennis). back when the corolla expands, and which may be easily divided with a pin into four. There are four petals in the corolla (c), and they are placed so as to wrap over each other at the base. The calyx is lined with the dilated receptacle, and in this lining are inserted the filaments of the eight stamens (as shown at d) ; the stamens CHAP. IV.] THE GENUS (ENOTHERA. ti having versatile anthers, that is, anthers attached to the filament by the middle, so as to quiver at every breath. The pollen contained in the cells of these anthers feels clammy when touched; and its particles, when magnified, will be found to be triangular, and connected by small threads, a form of construction peculiar to this genus and its allies. The style is long, and the stigma is four- cleft. The ovary (e e) is situated at the base of the calyx, and when ripe, it becomes a four-celled dry capsule, which bursts into four valves, opening at top to discharge the seed. The seeds, when young, are attached to the central placenta, and they are quite free from hair or wool of any kind. The genus CEnothera being a very extensive one, it has been divided by M. Spach, a German botanist residing in Paris, into fourteen new genera ; but only one, or at most two, of these genera have been adopted by other botanists. One of these Godetia, which embraces all the purple-flowered kinds, has been divided from OEnothera, on account of a slight feathery ap- pearance on the seeds ; whereas the seeds of the true yellow-flowered GEnotheras are naked, that is, without the slightest appearance of any feathery substance or wing. The other genus, Boisduvalia Spach, includes only two species, both with pink flowers, which are very seldom 82 THE GENUS EPILOBIUM. [part I. seen in British gardens. The generic mark of distinction consists in four of the stamens in these species being shorter than the other four ; whereas in the true (Enotheras all the eight stamens are of equal length. As M. Spach's other genera have not been adopted by any British botanist, it is not worth while troubling my readers with the distinctions between them. The flowers of the yellow GEnotheras only open in the evening, or in cloudy weather ; but those of the purple kinds, or Godetias, remain open all day. The leaves in both kinds are alternate. THE GENUS EPILOBIUM. This genus is plant often seen in shrubberies, called the French Willow - Herb — (Epilobium an- gustifolium) , and the English weed called Codlings- and - Cream {E. hirsiitum). In this genus, the tubular part of the calyx which incloses the well linown, by the showy Fig. 34.— Epilobivm roseum. ovary, is quadrangular, as CHAP. IV.] THE GENUS CLARKIA. 83 shown at a in Jig. 34, which represents seed- vessels of Epilobium roseum^ a very common weed in the neighbourhood of London. The limb of the calyx is four-cleft, and the corolla has four petals ; and when these fall off, the ovary assumes the appearance shown at a. The quadrangular form is re- tained by the capsule, which, when it ripens, bursts open into the four valves (6), and discharges the seed op'^Epix^Bi^t^:^ which was attached to the central placenta (c) ; each seed being furnished with a little feathery tuft resembling pappus, as shown in^^. 35. The genus Epilobium is divided into two sections ; the plants in one of which have irregular petals, the stamens bent, and the stigma divided into four lobes, as in the French Willow- Herb, and the other showy species ; and the plants in the other section having small flowers with regular petals, erect stamens, and the stigma undivided. THE GENUS CLARKIA. The calyx in this genus is tubular, with the limb in two or four lobes, as in OEnothera. The corolla is, however, very different, the four petals being unguiculate or clawed ; that is, so much narrower in the lower part as to stand widely apart from each other ; they are also three G 2 84 THE GENUS CLARKIA. [part i. lobed. The stamens are very different, onl}^ four of them being perfect, and the anthers of the other four being wasted and destitute of pollen ; and the stigma is divided into four leaf- like lobes, ver}^ different from those of all the other genera included in the order. The cap- sule is cylindrical in shape, and furrowed on the outside ; it is four-celled, and when ripe, it bursts open by four valves. The seeds are quite naked. Among the other genera belonging to this order, I may mention the following : Gaura^ the petals of which are somewhat unguiculate, like those of Clarkia, but not three-lobed as in that genus ; the segments of the limb of the calyx often adhere two together, so as to appear three instead of four ; the ovary is one-celled, and the seeds naked : Lopezia^ which has ap- parently five irregular petals, though, on examination, one will be found to be a meta- morphosed stamen, a four- cleft calyx, two stamens, including the one converted into a petal, and a globular, four-celled capsule : and Circcea, or Enchanter's Nightshade, which has the limb of the calyx apparently in only two segments, and only two petals and two stamens ; the capsule is globular like that of Lopezia, but it is covered with very small hooked bristles, and it is divided into only two cells, each con- taining only one seed. 85 CHAPTER V. THE ORDER RUBIACE^ : ILLUSTRATED BY THE CINCHONA, OR PERUVIAN BARK ; LUCULIA GRATISSIMA ; CAPE JASMINE ; RONDELETIA ; COFFEE ; IXORA ; IPECACUANHA ; MADDER ; GALIUM ; WOODRUFF ; AND CRUCINELLA STYLOSA. This order contains more than two hundred genera ; but by far the greater part of these are composed of tropical plants, many of which are not yet introduced into Britain. Several of the genera, on the other hand, are British weeds ; and this difference in habit, with others in the qualities of the plants, &c., have occasioned some botanists to divide the order into two : one of the new orders being called Cinchonacese, and containing the plants most resembling Cinchona ; and the other Galiacese, containing the plants most nearly allied to Galium or Bedstraw. The characteristics of Rubiacese, in its most extended sense, are that the ovary is surrounded by the calyx, and placed below the rest of the flower ; and that the corolla has a long tube, lined with the dilated receptacle, in which the stamens are inserted. In most of the species, the fila- ments are very short, and the anthers nearly or entirely hidden in the corolla; and in many cases. 86 THE GENUS CINCHONA. [part r. the segments of the calyx remain on the ripe fruit, as they do in the genus Pyrus in Rosacese, where they form what is called the eye in the apple and pear. Tlie qualities of the Cinchona division of the Rubiaceag are generally tonic ; but some of the plants, as for example the Ipecacuanha, are used as emetics, and one (Rcmdia dumetorum) is poisonous. The qualities of the Galium division are not so decidedly marked ; but the roots of some of the plants are used for dyeing. THE GENUS CINCHONA, AND ITS ALLIES. The well-known medicine called Peruvian bark is produced by three species of the genus Cinchona ; the pale bark, which is considered the best, being that of C. lanceolata. The flowers of this species are small, and of a very pale pink. The calyx (see a in fig. '^^) is bell-shaped, and five-toothed ; and the corolla (Z*) is tubular, with the limb divided into five lobes, and silky within, as shown in the magnified section at c. The stamens (d) have very short filaments, which are inserted in the throat of the corolla. The ovary (e), which is deeply furrowed when young, is inclosed in the calyx ; it is two-celled, with a single style, and a two-lobed stigma {f). The capsules retain the lobes of the calyx as a sort of crown (y) ; and they open naturally at the CHAP, v.] THE GENUS CINCHONA. 87 division between the two cells, as shown at A, beginning at the base. The cells (z) each con- FiG. 36.— CiNCHOVA, Peruvian Bark (Cinchona Lanceolata). tain several seeds. C, ohlonpfoUa, which yields the red bark of the shops, has cream-coloured flowers, as large as those of a Jasmine, which they resemble in shape ; and C. cordifolia^ which produces the yellow bark, has flowers like the first species, and heart-shaped leaves. The sin- gular plant called Hillia longiflora^ is nearly allied to Cinchona ; as is also the beautiful and delightfully fragrant Liiculia gratissima. In this last plant the tube of the calyx is very short, and pear-shaped, and the segments of the limb are short, and sharply pointed. The corolla is salver-shaped, with a long tube, and a spreading, five-parted limb. The anthers are nearly sessile, and the short filaments to which they are at- BB THE GENUS CINCHONA. [part i. tached are inserted in the throat of the corolla, only the tips of the anthers being visible. The stigma is divided into two fleshy lobes, and the capsule splits, not like that of Cinchona, but from the apex to the base in the centre of each cell. The seeds are very small, and each has a toothed, membranous wing. The flowers of this beautiful plant are produced in a large head, and at first sight greatly resemble those of a Hydrangea ; but they are easily distinguished by their delightful fragrance. Manettia corcUfoIia, a very pretty stove-twiner often seen in collections, is very nearly allied to Luculia, differing principally in the shape of the flowers, which in Manettia have a long tube and a very small limb. Bouvardia triphylla and the other species of Bouvardia, and Pinckneya pubescens, belong to this division ; and such of my readers as have the living plants to refer to, will find it both interesting and instructive to dissect them and compare the parts of their flowers with the description I have given of Luculia and Cinchona, so as to discover the difference between the different genera ; after- wards reading the generic character of each given in botanical works, that they may see how far they w^ere right. CHAP, v.] THE GENUS GARDENIA. 89 THE GENUS GARDENIA AND ITS ALLIES. The Cape Jasmine {Gardenia radicans) is a well-known greenhouse plant, remarkable for the heavy fragrance of its large white flowers, which (lie off a pale yellow, or buff. The calyx has a ribbed tube, and the limb is parted into long awl-shaped segments. The corolla is salver-shaped, that is, it has a long tube and a spreading limb, the limb being twisted in the bud. There are from five to nine anthers, having very short filaments which are inserted in the throat of the corolla. The stigma is divided into two erect fleshy lobes. The ovary is one- celled, but there are some traces of membranes, which would, if perfect, have divided it into from two to five cells. The seeds are numerous and very small. Gardenia radicans is a dwarf plant, which flowers freely when of very small size, and is easily propagated from the readiness with which its stem throws out roots ; but G.florida is a shrub five or six feet high, and much more difficult to cultivate. In both species the flowers are generally double, and the petals are of a fleshy substance, which gives the corolla a peculiarly wax-like appearance. There are many other species, but the two above-mentioned are the most common in British gardens. Burchellia capensis is gene- 90 THE GENUS RONDELETIA. [part I. rally considered to belong to this division of Rubiacese, though its flowers bear more resem- blance to those of Cinchona ; and the singular plant called MusscBiida puhescens^ the flowers of which are small and yellow, but the bracts are so large and so brilliantly white as to look like flowers ; Posoqueria versicolor, an ornamental plant lately introduced, belong to this division. THE GENUS RONDELETIA AND ITS ALLIES. Rondeleiia odorata, sometimes called M. coccinea, and sometimes R. speciosa, is a very fragrant stove shrub, a native of Cuba. The flowers are produced in corymbs, and their botanical construction is shown in the magnified section Jig. 37. In this a is the ovary inclosed in a hairy calyx ; h shows the limb of the calyx cut into awl-shaped segments ; c shows the manner in which the very short filaments of the anthers are inserted in the throat of the corolla ; d shows the termination of the dilated receptacle which ,, Fig. 37.— Sectiox of the lines the tube oi the corolla ; flower of KoNOiiLETiA. and e the segments of the CHAP, v.] THE GENUS COFFEA. 91 limb. I have given the section of this flower, that my readers may compare it with the sec- tion of the flower of the Cinchona in fig. 36, in p. 87, and may see the general resemblance which connects the two plants in the same order, and the diff*erences which mark them to be of different genera. Fig. 38 is a tuft of flowers of Hon- deletia odorata. Wendlandia is nearly allied to Ronde- letia ; as is the magnifi- cent Portlandia grandiflora^ which somewhat resembles Brugmansia lutea in shape though not in colour, as its fig. 38.— part of the hkad „ - . OF FLOWERS OF RoNDELETIA. flowers are white. THE GENUS COFFEA. AND ITS ALLIES. The Coff-ee-tree {Coffea arahicd) differs from the other Rubiacese in the tube of its calyx being very short and disappearing when the ovary begins to swell ; and in the filaments of the stamens being sufiiciently long to allow the anthers to be seen above the throat of the corolla (see a in Jig. 39). The limb of the corolla ifi) is five-cleft, and the style (c) bifid. Each ovary when its flower falls, becomes dis- tended into a berry {d) or rather drupe, con- taining the nut e, in which are two seeds, flat 92 THE GENUS COFFEA. [part I. on one side, and convex on the other, which are placed with the flat sides together, as shown at f; each seed having a deep longitudinal groove, as shown at g. These seeds are our coffee. Fig. 39 — Coffee. (Cofea Arabica.) The flowers of Ixora coccinea have the same general construction as those of the other plants of the order. The calyx has an ovate tube, and a very small four-toothed limb ; and the corolla is salver-shaped, with a long and very slender tube, and a four-parted spreading limb. There are four anthers inserted in the throat of the tube of the corolla, and just appearing beyond it, and rising a little above them is the point of the style with its two-cleft stigma. The berry CHAP, v.] THE GENUS COFFEA. 98 is two-celled, but it differs from that of the coffee in retaining the lobes of the calyx, which form a sort of crown. There are many kinds of Ixora, all stove shrubs, and all conspicuous for their large heads or rather corymbs of showy flowers. The genus Pavetta has been divided from Ixora, principally because the species com- posing it have the style projecting considerably beyond the corolla, instead of only just appear- ing above it. The drug called Ipecacuanha is the produce of two plants belonging to this order, Cephcelis Ipecacuanha and Rlchardsonia scabra ; though a spurious kind is made from the roots of three species of Viola, all natives of South America, and a still inferior one from the roots of a kind of Euphorbia, a native of Virginia and Carolina. It is important to know this, as the best kinds possess tonic properties as well as emetic ones, while the inferior kinds are only emetics, and they are very injurious if taken frequently. The best brown Ipecacuanha is the powdered root of CephcBlis Ipecacuanha ; a plant with small white flowers collected into a globose head, which is shrouded in an involucre closely re- sembling a common calyx. The true calyx to each separate flower is small and roundish, with a very short five-toothed limb. The corolla is funnel-shaped, with five small bluntish lobes. 94 THE GENUS GALIUM. [part i. The EA, CINERARIA, SUNFLOWER, MUTISIA, AND TRIPTILION. The plants composing the order Compositse have all compound flowers, which differ from other flowers as much as a compound leaf does from a simple one. As the compound leaf is composed of a number of leaflets or pinna? united by a common petiole ; so a compound flower is composed of a number of florets, united by a common receptacle, which is surrounded by a calyx-like involucre, so as to give the whole mass the appearance of a simple flower. Each floret has a calyx, the tubular part of which is rarely sufficiently distinct to be perceptible, but the limb is generally cut into long feathery segments called pappus. The ovary of each floret contains only one seed ; and the fruit, which is called an achenium, retains the pappus when ripe, and falls without opening. There are five stamens, the filaments of which are dis- tinct, but the anthers grow together so as to CHAP. VI.] COMPOSITE. 99 form a kind of cylinder, through which passes the style, ending in a two-lobed stigma (see a in Jig. 41). Most of the corollas are of two kinds : viz. the ligulate, as exem- plified in the floret of the wild Lettuce {Lactuca virosd) shown in Jig. 41 ; and the tubular, as shown in a floret of the Cotton-thistle {Onojwrdium Acantliium) see Jig. Fig. 41.-LIGULATE 42. All the British species of FLORET OF WILD Compositse have their florets either Lettuce. entirely of one of these kinds, or of the two mixed together ; but some foreign genera have florets with two equal lips, cut into three or four lobes, as shown in a floret of Mutisia latifoUa, at e. Jig. 46, p. 108. These flo- rets are called bilabiate. It will be observed that in all these ex- amples, as indeed, in all the flowers belonging to the order, that the pappus {b, in Jgs. 41 and 42), is always on the outside of the corolla, thus plainly indi- floret of the eating its connexion with the cotton-thistle. calyx. The order Compositse is a very large one, above seven thousand species having been H 2 Fig. 42.— Tubular 100 COMPOSITE, [part I. named and described ; and to assist the memory in retaining the names of this great number of plants, various means have been devised for dividing the order into sections and tribes. The principal botanists who have proposed means of arranging this order, are Cassini, Lessing, and lastly the late Professor De Can- doUe, in three voluines of his Prodromus pub- lished in 1840. But as the distinctions between the divisions proposed, lie in the difference found in the stigmas and pappus of the different genera, I have judged them too troublesome for my readers, as I am sure they are for myself, and I have preferred following the plan adopted by Dr. Lindley in his Elements of Botany^ pub- lished in 1841, and dividing the Compositse into four tribes ; viz., the three originally proposed by Jussieu, and a fourth added by Professor De CandoUe, containing the plants with bila- biate florets, which were either not known, or overlooked, by Jussieu. It may perhaps be necessary to add, that this arrangement forms the basis of the new one proposed by De Can- doUe, and that the principal difference consists in the subdivisions. CHAP. VI.] CICHORACE^. 101 TRIBE I.— CICHORACE^. Florets ligulate. Juice milky, narcotic. ' The plants contained in this tribe bear more or less resemblance to the common Succory {Cicliorium Intyhus). This beautiful plant, which is found in great abundance wild in many of the sandy and chalky districts of England, has large bright blue flowers, which when ex- amined will be found to consist of a number of florets, all of the kind called ligulate, that is somewhat like a cornet of paper ; the upper part being broad and flat, and serrated at the edge. The pappus in this genus is very short, and it is scaly rather than feathery. The leaves are bitter, and when broken give out a milky juice; and the fleshy roots when roasted are used to adulterate coffee. The Endive is a variety of this species, or another species of the same genus. The Sowthistle (Sonchus oleraceus) abounds in the same milky juice as the succory, and has the same kind of fleshy root. The flower is composed of a scaly involucre (shown at a in Jig. 43) and a number of ligulate florets (see Z>), which when they fall show the pappus (c), forming a feathery ball. The manner in which the pappus is attached to the seed-vessel is shown at d ; and the receptacle after the florets have been pulled out, but with the involucre still 102 CICHORACE^. [part I. attached to it, at e. A detached floret is shown aty. The DandeUon {Leontodon Taraxacum) Fig. 43.— SottTHisTLE. (Sonckus oleraceus.) differs from the Sowthistle : in its florets, which are flatter and looser ; in its receptacle, which is globular ; and above all, in its pappus, which is what is called stipitate or stalked, that is, the tubular part of its calyx rises to a considerable height above the capsule, before it becomes divided into its feathery segments, as shown in fiq» 44. The leaves of this plant are „,,. , what is called runcinate, that is, the ^pw^. lobes into which they are cut point ^! downwards towards the root instead of upwards from it, and the root is also fleshy. The Lettuce, Salsafy or Goat's- beard. Ox- tongue. Hawk weed, CatV ^ ear. Nipplewort or Swine's Succory, and seedofthe 1 n 1 1 11 Dandelion. many other well-known plants, belong to this tribe. CHAP. Vi.] CYNAROCEPHAL^. 103 TRIBE II=— CYNAROCEPHAL^. Florets tubular. Juice watery, tonic. The plants in this division all bear more or less relation to the common Artichoke (Cynara Scoli/mus). The scales of the involucre are generally fleshy at the base, but terminate out- wardly in a sharp hard point. The florets are tubular, and intermixed with them in the recep- tacle are frequently found the hardened bracts, which in this state are called palese, and which appear to be of a chaffy substance, as exemplified in the choke of the Artichoke, the fleshy recep- tacle being in this plant what we call the Arti- choke bottom. This peculiar formation is shown more in detail in^^. 45, which represents part of the flower of the common Bur or Bur- dock {Arctium Lappa\ so annoying from the strong hold it takes of any part of the dress which it may chance to touch. In Jig. 45 a is the involucre, every scale in which is hooked and turned inwards, so as to hold firmly whatever it may catch ; ^ is a floret showing its tubular shape, and its style proceeding through the Fig. 45. — Part of the flower- head OF THE Burdock. 104 CORYMBIFER^. [part i. united anthers ; c shows the hardened bracts or palese, the other florets having been removed; and d shows a fruit with a palea attached, mag- nified. All the different kinds of thistle belong to this division ; and though many of the kinds have not the hardened bracts, they have all a spiny involucre. The pappus of the thistle is generally attached to a kind of disk, from which it becomes loosened soon after the seed falls, and this thistle, down, as it is cjlUed, being ex- tremely light, is blown about by the winds. All the thistles have fleshy roots, and take firm hold of the soil. The Corn Blue-bottles {Cen- taur ea), the Wild Safli'on (Carthamus tinctorid)^ and many other well-known plants, belong to this division. TRIBE III.— CORYMBIFER^. Florets partly tubular and partly ligulate ; juice watery ; sometimes bitter and tonic, and sometimes acrid. The seeds of some of the species yield oil. The plants included in this tribe all bear more or less resemblance to the common Daisy. In this well-known flower, the white florets are all ligulate, and compose what is called the ray, and the yellow flowers, which are tubular, are called the disk. The involucre is simple and leafy, and the receptacle is conical. The seeds are without pappus. The Chrysanthemum is CHAP. VI.] CORYMBIFER^.' 105 nearly allied to the Daisy, and its seeds also are destitute of pappus ; but it is easily distin- guished by its involucre, which is scaly, and by the flower forming a kind of depressed globe in the bud. The scales of the involucre are strongly marked, from being edged with a thin membrane, and the florets of the ray are much longer in proportion to those of the disk than in the Daisy. The great Ox-eye Daisy, which was formerly called Chrysanthemum leucanthemum^ is now placed in a new genus, and called Leu- canthemum vulgar c; and the Chinese Chrysan- themums have been removed to the genus Fyre- thrum. Both plants, however, will no doubt long continue to be called by their old names. The beautiful yellow-flowered plant often found grow- ing among corn {Chrysanthemum segetum), the three-coloured Chrysanthemum (C tricolor or carinatum)^ and the yellow annual Chrysanthe- mum (C coronarium), with some others, have been left by Professor De CandoUe in their old genus. In the Feverfew (Pyrethrum)^ the re- ceptacle is elevated, and the fruit is crowned with a narrow membrane. The Pellitory of Spain was formerly considered to belong to this genus, and afterwards to the Chamomile, but it is now called Anacyclus Pyrethrum. Matricaria Chamomilla^ the wild Chamomile, has also no pappus; and in this plant the receptacle is 106 CORYMBIFER.^. [part i. almost cylindrical. The true Chamomile {A?i- themis nohilis) greatly resembles the Chrysanthe- mum in its flowers : but they are distinguished by haying a chafiy receptacle, and the fruit ha^dng a membranous margin. The smell of the Chamomile is aromatic, and its qualities highly tonic. The Yarrow {Achillea miUefolium) is another plant destitute of pappus, but with a chaffy receptacle : it is also remarkable for its leayes. which are doubly pinnatifid. It will be seen by the aboye enumeration, that in many plants belonging to this diyision, the pappus is entirely wanting, and in others it wXi. be fotmd to assume a different form to that which it bears in the other tribes. Thus, in the Bur-Marigold {Bidens), the pappus consists of from two to fiye erect awns, which are covered with yery small, bent bristles. The genus Sene- cio has soft, hairy pappus, as may be seen in the common Groundsel (S. vulgaris) ; the leaves of this weed are pinnatifid, and somewhat stem- clasping, and the flowers have no ray florets. In •other species of this division, however, the ray florets are very conspicuous : as, for example, in the common yellow Ragwort (5. Jacola^a)^ in the great fen Ragwort, or Bird's tongue (5. paludosa)^ and in the purple Jacobsea (S-elegajis), Nearly alhed to Senecio, is the genus Cineraria, so much, indeed, that Professor De Candolle, CHAP, vr.] LABIAT-^FLOR-E. 107 in his late arrangement of the Compositce, has included the greater part of the species in Sene- cio. The green-house species, with purple flowers, are among those which have been changed ; but they will probably always retain the appellation of Cineraria, as an arbitrary English name. The Asters, or Michaelmas Daisies, Golden Rod, Elecampane, Leopard's Bane, the Cape Marigold, (now called Dimor- phortheca, instead of being included in the genus Calendula), Coltsfoot, "Wormwood, Southern- wood, Tansy, and many other well-known plants, belong to this dinsion. The Sun-flower {Helianthemum annuus) is an example of one of the plants belonging to this division which has seeds yielding oil. In this plant the pappus is awl-shaped, and deciduous ; and the receptacle, which is broad and some- what convex, is paleaceous. The seeds are large and oblong, and when pressed, yield a consider- able quantity of oil. The ^ladia is another oil plant ; and indeed the seeds of several in this division yield oil. TRIBE IV.— LABIAT.EFLOR.E. Florets bilabiate. The plants belonging to this division are rarely seen in British gardens ; but when they do occur, they are well worth examimng, from 108 LABIAT^FLORtE. [part I. the singularity of their formation. Mutisia latifolia (see fig. 46) has a large, woolly involucre, the |\ A ^ scales of which are {il ) i] I I /] A of two kinds, the VA .,\ t. /' //^' outer ones, (a), being pointed and leaf-like, and the inner ones, (^), having the ap- pearance of scaly bracts. The flo- rets of the ray, (c), are narrow, and spreading in the fully expand- ed flower ; and those of the disk, (f/), are shorter, erect, divided into two lips, which curl back, and the lower one of which is again divided into two segments (as shown at e in the detached floret). The leaves of this plant are very curious ; the mid- rib is lengthened and drawn out into a ten- dril, as shown atyj and the petiole {g) is decur- rent. There are several other genera belonging to this tribe, but none of them are particularly ornamental except Ti^iptilion spinosiim^ which has flowers of the most brilliant blue, that do not lose the intensity of their colour in drying. Fig. 46. — Flower and Leaf of Mutisia Latifoll*.. CHAP. VII.] ERICACE^. 109 CHAPTER VII. THE ORDER ERICACE.E : ILLUSTRATED BY THE COMMON OR BESOM HEATH, THE MOOR HEATH, CAPE HEATHS, LING OR HEATHER, ANDROMEDA, LYONIA, ST.^DAB^OC's HEATH, ARBU- TUS, THE BEARBERRY, GAULTHERIA, CLETHRA, RHODODEN- DRON, INDIAN OR CHINESE AZALEAS, YELLOW AZALEA, AME- EICAN AZALEAS, RHODORAK, ALMIA, MENZIESIA, LOISELEURIA, LEDUM, LEIOPHYLLUM, THE BILBERRY, THE WHORTLE- BERRY, THE CRANBERRY, PYROLA, AND MONOTROPA. The name of Ericaceae, which most people are aware signifies the Heath family, conjures up immediately the image of a number of narrow-leaved plants, with globular, ventricose, or bell-shaped flowers ; and we are apt at first to think that the family is so natural a one, as to require very little explanation. Did the order include only the Heaths, this would be the case, for all the Heaths, different as they are in some particulars, may be recognised at a glance : but as the order includes the Rhododendrons, Azaleas, and Kalmias, besides several other plants which have not so strong a family like- ness to each other as the Heaths, it becomes necessary to say a few words on the botanical resemblances which connect them together. The first, and most striking, of these is the shape of the anthers, each of which appears like two 110 ERICACEAE. [part I. anthers stuck together, and the manner of their opening, which is always by a pore or round hole, in the upper extremity of each cell. The filaments, also, in all the genera, except Vaccinum and Oxycoccus, grow from beneath the seed- vessel, being generally slightly attached to the base of the corolla. There is always a single style with an undivided stigma, though the cap- sule has generally four ceils, each containing several of the seeds, which are small and nume- rous. The calyx is four or five cleft, and the corolla is tubular, with a larger or smaller limb, which is also four or five cleft. The order has been divided into four tribes, which I shall describe in this chapter, though some of these are considered as separate orders by Dr. Lind- ley and other botanists. TRIBE I.— ERICE^. This tribe, which comprehends all the heath- like plants, has been re-divided into two sub - tribes, one con- taining the ge- nera most nearly allied to the heaths, and the other those be- F1G.47.— The Besom Heath (Erica Tdralix). CHAP. Til.] ERICE^ NORMALES. Ill longing to the Andromeda. In both there is a honey-bearing disk under the ovary, and the leaves are generally rolled in at the margin, as shown at a, in Jicj. 47. SUB-TRIBE I.— ERICEiE NORMALES, All the genera in this sub-tribe, twenty-two in number, were formerly included in the genus Erica ; and some botanists still consider all the species to belong to that genus, ^vith the ex- ception of those included in Calluna, while others adopt about half the new genera. In this uncertainty, I shall only describe two of the doubtful genera, partly because the dis- tinctions between them and the true heaths are strongly marked, and partly because the spe- cies they contain are frequently met with in British gardens^ and greenhouses, where they are sometimes labelled with their old names and sometimes with their new ones. In the genus Erica, one of the commonest species is the Besom Heath (Fj. tetralix)^ which is found in great abundance on moorish or boggy ground in every part of Britain. In this plant, the corollas of the flowers appear each to con- sist of a single petal, forming an egg-shaped tube (see h in Jig. 47), contracted at the mouth, but afterwards spreading into a four-cleft limb, through which is seen projecting the style, with 112 ERICACE^. [part I. its flat stigma. The corolla is, however, really in four petals, which, though they adhere to- gether, may be easily separated with a pin. The stamens are concealed by the corolla, but the manner in which they grow is shown at c ; and cf is a single stamen, showing the spurs or awns at the base of the anther, the position of which is one of the characteristics of the genus Erica in its present restricted form ; e is a capsule with the style and stigma attached ; and a is a leaf showing its revolute or curled back margin. The leaves of this species are in whorls, four leaves in each whorl, and they are ciliated, that is, bordered with a fringe of fine hairs. All the true Heaths bear more or less resem- blance to this plant. In some, the corollas are bell-shaped, spreading out at the tip into five teeth, which inclose the stamens, as shown in Jig. 48 ; and in others they are nearly globose swelling out near the calyx, and tapering to a point, beyond which the stigma and anthers pro- ject ; as in the Cape Heath, called Erica hispidaf a flower of which is shown in ^(/, 49. The leaves also Fig. 48— differ exceedingly, in the Bell-shaped number Contained in each Fig.49.-Cape whorl : as in some species heath. . 11 (E. hispida.) there are only three m a wnorl, CHAP. VII.] ERIC E^ NORM ALES. 113 while in others they are five or six. The general features of all the Heaths are, however, the same — viz., there are eight sta- mens, which are generally inclosed in the corolla, though they sometimes project beyond it, as shown in jig, 49, and the anthers of which are two-cleft, and awned or crested at the base, while the filaments are hair-like ; one style, which always projects beyond the corolla, and has a flattened stigma; a four-parted calyx and corolla which is tubular, with a four-parted limb. There are nearly two hundred species of this genus, some of which are natives of Europe, and others of the Cape of Good Hope. The moor Heaths {Gypsocallis) were separated from the genus Erica, by Mr. Salisbury, princi- pally on account of the corolla being campanu- late, or shortly tubular, with a dilated mouth ; and the stamens projecting beyond the corolla. The filaments are also generally flat; the anthers are without awns, and distinctly in two parts ; and the stigma is simple, and scarcely to be distinguished from the style. The com- mon Cornish Heath {G. vagans), and the Mediterranean Heath (G. Mediterranea) ^ are examples of this genus, which appears strongly marked, though, as 1 before mentioned, some botanists do not adopt it. Callista is a genus established by the late 114 ERICE^ NORMALES. [part I. Fig. 50.— Callista bucciniflora. Professor Don, which appears very distinct, though it also has not been generally adopted. It includes all those beautiful Cape Heaths which have a shining, glutinous, ventricose, or cylindrical corolla with a _ spreading limb (see« in^^. 50), and a capitate stigma (b). C. bucciniflora and C. ventricosa, are examples of this genus. The Ling or Heather, which Linnaeus called Erica vulgaris^ is now generally placed by all botanists in a separate genus called Calluna, which was es- tablished by Mr. Salisbury. The calyx of this plant is membranous, and coloured so as to resemble a corolla, and it is furnished with four bracts at the base, which resemble a calyx. The true corolla is bell-shaped, and shorter than the calyx. The stamens are inclosed, and the anthers are of the very singular form shown in Jig. 51. The stigma is capitate, and the flowers are disposed in what is called a racemose spike. The leaves are trigonal ; they are very short, and they are laid over each other like scales in four rows. The Ling is the only species in the genus. Fig. 51.— Stamen of the Ling. CHAP. VII.] ANDROMEDE^. 115 SUB-TRIBE II. ANDROMEDE-^. The plants in this sub-tribe differ decidedly from those of the preceding division, in having ten stamens, while all the genera of heaths have only eight. The calyx is also five-cleft instead of four; and the corolla, which falls before the seeds are ripe, has a five-lobed limb. The sub-tribe is divided into twenty genera, more than half of which are perfectly distinct. The genus Andromeda is distinguished by its globose corolla which has a five-lobed limb ; and its stamens which have their filaments bearded, and their anthers short and two- awned. Tig. 52 shows a stamen of the wild rosemary (Andromeda poll- * m' folia) with its bearded filament («), p « and its two-awned anther with its ^^^ ^^_ pore-like openings {h). The cells stamen of of the capsule open in the middle, ^^'''^o-^r^'^-^ down the back, to discharge the seeds. Professor Don has divided the genus Andromeda into six genera ; some of which contain only one or two species. Thus only Andromeda polifolia and A. rosmarinifolia are left in the genus Andromeda ; Cassandra contains only A. calyculata^ and A. angustifolia ; and Zenobia, only the beautiful 116 ANDROMEDE^. [part I. Andromeda speciosa. In Cassandra the anthers are long and mutic (see a va.Jig. B^), and the leaves {b) are without veins, and white and full of dots on the underside, the edges being curled inwards ; and in Zenobia the corolla Fig. 53 -Leaf is bell-shaped, with the limb, which AND ANTHER jg jjj ^yg lobes, curliug back (see a 'Jig. 54). The stamens have the filaments (h) curiously dilated at the base; Fig. 54.— Flowers, corolla, and stamen of Zenobia. and the point of each cell of the anther is cut into two erect awns (c). The manner in which the stamens are arranged inside the corolla is CHAP, vi..] ANDROMEDEiE. 117 ehovm at (rZ). The cells of the capsule, when ripe, open down the centre, and the seeds which are angular, are attached to a five-lobed placenta. Lyonia is a genus established by the American botanist Nuttall, because the plants it contains have the margins of the valves of their capsules closed by five other narrow external valves. The plants are natives of North America, and their flowers are generally small. Lyonia Mariana may serve as an example of this genus, which is gene- rally adopted by botanists. It would be useless to enter into details of the other genera formed out of Andromeda, as they are not generally adopted ; but, perhaps, it may be worth mentioning, that the well-known Andro- meda floribunda is placed by Professor Don in a new genus which he calls Leucothoe. St. Dabeoc's Heath, or Irish Whorts, a little heath-Hke shrub, common in Ireland, is one of those plants which have puzzled botanists exceedingly. It has been called successively Erica, Andromeda, and Menziesia, Dabcecia ; then Erica Hibemica, next Menziesia polifolia, then Vaccinium Cantabrieum and lastly Daboecia ^yoU- folia. It is probable, however, that it may even yet be doomed to undergo other changes ; as, from the construction of its anthers, which are linear, and arrow-shaped at the base, and 118 ANDROMEDE^. [part which open lengthways, instead of by pores, it does not appear even to belong to the Ericacese. The other genera in this sub -tribe are quite distinct from each other, and contain several well-known plants. The most popular of these genera are Arbutus, Arctostaphylos, Gaultheria, and Clethra. The Strawberry tree (Arbutus Unedo) has little bell-shaped flowers, contracted at the mouth, and with a curling-back limb, which are easily recognised as belonging to the Ericacese. They have ten stamens, the filaments of which are hairy at the base (see a in Ji^. 55) and inserted in the disk; which in this genus is large, and rises up round the ovary (see I). The calyx is permanent, and Fio. 55.— Fruit &c. of Arbutus Unedo. fivC-clcft ; and the flowers are produced in panicles, and each is furnished with a bract. The fruit, which retains the calyx when ripe, is a granular berry, covered with tubercles on the outside ; and it has five cefls (c) containing the seeds. There are nu- merous varieties of this species common in British gardens, besides a very beautiful hybrid CHAP. VII.] ANDROMEDE^. rl.l9 between it and A. Andrachne. The latter species is a native of Greece, and rather more tender than the common kind ; and it is very conspicuous in shrubberies from its red stems and loose bark. The Bearberry (Arctostaphylos Uva- Ursi) was formerly considered to belong to the genus Arbutus, but it differs in the filaments of the stamens being smooth and dilated at the base, and the awns affixed to the middle of the anthers. The berry is without tubercles, and the cells are often only one-seeded. There are two species of Gaultheria common in British gardens : viz. — G. procumhens and G. Shallon : both of which have flowers resem- bling those of the Arbutus and furnished with bracts ; but in the former species the flowers are solitary and produced from the axils of the leaves, and in the latter they are in racemes, of the kind called secund, that is with the flowers growing all on one side. The berries of both kinds are eatable, and those of G. procumhem are called Partridge berries in America, and the leaves Mountain tea. Both species have ten stamens, the anthers of which are two- cleft, each cell being furnished with two horns, as in Zenohia speciosa (see Jig. 54, in page 116). The fruit is five-celled and the seeds are numerous. 120 RHODOREiB. [part I. The genus Clethra differs considerably from the preceding genera, as the limb of the corolla is so large and so deeply cleft, as to make the flower appear to have five petals (see a in Jig, 6Q). There are ten stamens, with broad arrow-shaped anthers (^), and a three-cleft stigma, (c). The cap- sule is dry, with three many-seeded cells. In C. alnifolia^ a native of North America, (of which^?^. 5 6 represents a magnified flower,) the flowers are erect, and produced in a spicate raceme ; but in C. arhorea^ a native of Madeira, the racemes are panicled, and the flowers drooping and somewhat bell- shaped. Both species are very ornamental. Fio. 56 Flower of Custhra Alntfolia. TRIBE II.— RHODORE^. The plants included in this tribe are all con- sidered to bear more or less resemblance to the Rhododendron, though in some of them the family likeness is not very strong ; and the genera I shall describe to illustrate it are CHiP. VII.] RHODORE^. 121 Rhododendron, Azalea, and Rhodora (the last two being by some botanists included in Rhododendron); Kalmia, Menziesia, and Ledum. The species of the genus Rhododendron are easily distinguished by their flower buds, which are disposed in the form of a strobile, or pine- cone, each bud having its accompanying bract, which the flower retains after its expansion, as shown '\xijig. o7 at a, in a flower of R. maximum. Fig. 57.— Back view of a Flower of Rhododendron Maximum, AND Seed-pod. There are five or ten stamens of unequal length, the larger ones curling upwards (as shown at b in Jig, 58), as does the style (c), which has a simple stigma. The flowers have a very small 122 RHODOREiE. [part i. calyx, {d in fig, 57,) and a carapanulate corolla which is deeply five-cleft, the upper segment {e in fig. 58) being somewhat larger than the rest, and spotted in the inside. The capsule is five-celled and five-valved, as shown in^^. 57/^ Fig. 58 — Flower of Rhododendron Ponticu.m. The leaves of nearly all the species are ever- green ; and the flowers are showy, and produced in terminal corymbs. The principal species may be thus distinguished from each other ; R, maximum has drooping leaves, covered with brown or white down on the under surface, and a dense corymb of flowers, the segments of the corollas of which are roundish, and the bracts leafy. In R. ponticum^ on the contrary, the corymbs of flowers are looser, the segments CHAP. VII.] RHODOREiE. 123 more pointed, and the bracts more scale-like ; and the leaves are smooth on both surfaces. The seed-pods also differ: inihose of H, maximum and the other American species, the valves are smooth as shown at/* in ^^. 57 ; and in those of R. ponticum, the valves are somewhat crin- kled as shown in Ji^. 59. This species, and all its hybrids and varieties, are more tender than R. maximum, R. cafawbiense, and all the other American kinds and their offspring. R. cataicbiense has the flower of a darker colour on the outside of the corolla than within, ^^«- ^^-s^^^ and the upper segment is very famtly rhododendron dotted. It hybridises freely with R, ponticum. arhoreumy which R. maximum does not, and the hybrids thus produced are hardier than those raised from R. ponticum, though the latter are by far the most numerous. Most of the species have purple or whitish flowers, but some, such as R. chrysanthemum, and R. anthopogon, have yellow flowers; R, ferrugineum and R. hirsutum, have bright pink or rose-coloured flowers ; and those of R. arhoreum the Nepaul tree Rhododendron, are of a rich scarlet. The commonest small kinds are R. ferrugineum and R. hirsutum, both dwarf shrubs and natives of the north of Europe, 124 RHODORE^. [part i. with funnel-shaped corollasj and leaves dotted on the under surface. They are so much alike as scarcely to be distinguished at first sight, but on examination the leaves of It.ferrugineum will be found to have brown dots, and to be plain on the margin ; while those of R. hir- sutum have white dots and are fringed with fine hairs. Of all the species of the genus, those which differ most widely from the others are the Indian kinds. Of these R. arhoreum has a ten- celled capsule, and the segments of the corolla two-lobed with waved margins. The leaves are long and silvery beneath ; and the capsules, the peduncles, and the calyxes, are all woolly. In R. campanulatum^ a splendid species with very large flowers, the capsule is six-celled, the leaves are somewhat cordate at the base, and the bracts are fringed ; and in R. anthopogon the corolla has a cylindrical tube, woolly inside, and a small but spreading limb, cut into five lobes. There are eight stamens, and the capsule is five- celled. R, Camtschaticum, R. Chamcecistus, and R. dauricum differ from the preceding species in having their corollas rotate, that is, wheel- shaped. The last of these kinds is a fa- vourite greenhouse shrub, from its flowering under shelter in winter. In the open ground CHAP. vii.J RHODOREjE. 125 it flowers in March. The species has rose- coloured flowers which appear before the leaves; and leaves which turn red in autumn before they fall. The roots are knobbed and fibrous ; and the stems are twisted and knobbed in a wild state. There is a variety R. d. atrovirens which has purple flowers, and evergreen leaves, and which is hardier than the species. The genus Azalea may be divided into three kinds, viz., A. indica and its allied species ; A. pontica and its varieties and hybrids ; and the American Azaleas. These divisions are easily distinguished by their flowers. Those of the Indian or Chinese Azaleas have all large showy flowers, on short downy footstalks, and they are produced in small clusters of only two or three flowers each, at the extremity of the shoots. The corollas are bell-shaped and deeply cut, nearly to the base, into broad spreading segments. The stamens are ten in number, shorter than the corolla, and of unequal length. The leaves are evergreen, and they are numerous, thickly set and downy. These Azaleas are all very handsome, but the white Indian Azalea {A. indica alba^ or A, ledifolia) is particularly so, and very fragrant. The species belonging to this division are mostly natives of China, and require either a greenhouse or some slight pro- tection during winter in England. 126 RHODOREiE. [part i. The yellow Azalea (A. pontica or Rhododen- dron Jlavum) differs from A. indica in being quite hardy ; in the flowers being produced in umbels of from eight to twelve, at the ends of the branches, before the leaves ; and in the corollas being funnel-shaped instead of campanulate. The tube of the funnel is, however, shorter than the limb, the segments of which are broad and spreading, the upper three being larger and of a darker yellow than the two below. There are usually five stamens, projecting alittle beyond the corolla, and curving upwards ; the style also curves upwards, and it is crowned by the stigma, which forms a round green head. The calyx is very small, and both it and the corolla feel clammy to the touch. The flowers are fragrant. The leaves are deciduous, and they are ovate, slightly hairy, and terminate in a mucro or stiff point. There are many varieties of this species, and many hybrids between it and the American kinds, all of which are quite hardy in British gardens. The principal American Azaleas are A. nudi- Jlora^ A viscosa, A. iiitida, and A. speciosa, all of which have the corollas of their flowers funnel- shaped. Of these A. nudiflora is easily known by its stamens, which project a long way beyond the corolla, and by the tube of the corolla being longer than the limb. The plant is deciduous ; and CHAP. VII.] RHODOREiE. 127 the flowers, which are produced in large terminal clusters, and which are not clammy, appear before the leaves. The common English name for this plant in some parts of the country is the American Honeysuckle, and the flowers are of various shades of red, pink, white, and purple. A. calendulacea, which some botanists make a variety of this species, has much larger flowers, and the leaves pubescent on both surfaces, whereas, in A, nudiflora the leaves are nearly smooth and green, with only a slight fringe of hairs round the margin. There are numerous varieties of A. calendulacea^ the flowers of which are always either yellow, red, orange, or copper- coloured, and it is supposed to be the parent of the beautiful Ghent Azaleas. A. viscosa has the tube of the corolla equal in length to the limb, and rather short stamens ; the flowers of this species are clammy. A. hispidum, which is generally considered a variety of A. viscosa, is still more clammy, and the tube of the corolla is wider and shorter ; other probable varieties are A. nitida, which has shining leaves, and A. glauca, which has glaucous ones, as in both kinds the flowers are very clammy. A. speciosa has large flowers and leaves tapering at both ends. All the species of Azalea have five stamens, but some of the varieties have ten. Rhodora canadensis is a little American shrub 128 RHODORE^. [part i. with pink flowers, which appear before the leaves, and the corolla of which is bilabiate, the upper lip being the broadest, and cut into two or three teeth, and the lower only once cut. There are ten stamens, and the capsule is five-celled and five-valved. The leaves are deciduous, and slightly pubescent beneath ; and the flowers are pro- duced in small terminal clusters. This plant, as well as all the Azaleas above described, are now included by some botanists in the genus Rhododendron. The genus Kalmia also belongs to this tribe. The flowers of this well known shrub are very curiously constructed. The corolla is salver- shaped, that is, nearly flat, and on the under side of the limb are ten protuberances, producing as many hollows on the upper side, in which lie half-buried the ten stamens. This singular construction gives the corolla that wrinkled appearance which has procured for the plant its American name of Calico flower ; while, from the shape of the leaves, it is also frequently called the Mountain laurel; it is also called Sheep laurel from its being considered poisonous to those animals when they feed on it. There are several species, which differ from each other principally in the shape of their leaves and the size of their flowers. Mcnziesia is a genus containing only three CHAP. VII.] RHODOREiE, 129 Species, of which M. pilosa (fig. 60) may be taken as an example. The flowers are small and bell-shaped, and the anthers {a) are without any awns or bristles ; there are eight stamens, FfG. 60.— Flowers, anthers, and pistil o? Menziesia. and the curious manner in which they are crowded round the style is shown at b. The capsule is four-celled. Loiseleuria, or Azalea procumhens^ is a small plant, having the appearance of thyme, which is the only species left in the genus Azalea by those botanists who include the true Azaleas in the genus Rhododendron. Ledum is the last genus belonging to this tribe that I shall attempt to describe. Ledum palustre, or wild Rosemary, the best-known species, has a corolla in five regular petals, and ten stamens which project beyond it ; but L. latifolium^ the Labrador Tea, has only five stamens, which are 130 VACCINIE^. [part I. not longer than the petals. L. buxifolium, a little thyme-like shrub, is now called Leiophylhim thymifolium. All the species have white flowers. TRIBE III VACCINIE^. The plants comprised in this tribe, which is considered a separate order by many bota- nists, all agree with the genus Vaccinium in having the ovary entirely surrounded by the calyx, which forms a fleshy berry-like fruit when ripe, and in the seeds being scaly. Vac- cinium Myrtillus, the common Bilberry or Blae- berry, is a famihar example of the genus; and Jig. 61 shows the shape of the flowers at a, the manner in which the ovary is enveloped in the calyx at J, and the curious shape of the anthers in the magnified representation of them at c. The berry is five-celled and many-seeded ; and there are eight or ten stamens. Both the anthers and the flower vary in the different species, but the calyx and the manner in which it surrounds the ovary are nearly the same in all, as may be seen m Jig, 62, which represents a specimen of V, tenellum^ the Pennsylvanian Whortle-berry. In Fig. 61. — Common Bilberry ( Vaccinium tenellum). CHAP. VII.] VACCINIE^. 131 this figure a is the flower, h the anther, and c the ovary surrounded by the calyx. Pig. 62. — Amebican Whortlk-berry {Vacciniiim pennsylvanicum'. There are many species, among which may be mentioned the American Bluets {V. angustifo- lium); Deerberries (F. stamineum); Bluetangles ( V. frondosum) ; the Hungarian Whortle-berry ( V. Arctostaphylos) ; and the Cow-berry, or com- mon British Whortle-berry {V. Vitis-IdcBo). The CvanheYry {Oxycoccus palustris) differs from the genus Vaccinium in the shape of its flowers (see^^. 68), and in its anthers being without spurs ; there are eight stamens, the filaments of which are connivent, that is, growing close together. The American Cranberry (0. macro carpus) differs from the European kind, principally in having larger fruit. K 2 Fig. 63.— CflANBERRY. 132 PYROLEjE. [part I. TRIBE IV.— PYROLEiE. This tribe is also considered as a separate order by many botanists ; but the principal dis- tinction is the long arillus or skin which enfolds the seeds and gives them the appearance of being winged. The most remarkable genera are Pyrola^ the Winter Green, of which there are several species common in moist woods in the north of England and Scotland ; and Monotropa^ or Bird's-nest, parasitic plants which grow on the roots of pine and beech-trees, but are by no means common in England. The species of Pyrola are pretty little evergreen plants, with white flowers, the corollas consisting of five distinct petals, and w^hich have ten stamens, the anthers of which are two- celled, each opening by a pore ; the style is single, ending in a capitate stigma cut into five lobes ; and the capsule is five-celled. The yellow Bird's-nest, {Monotropa Hypopitys) has a coloured stem, with drooping flowers, and numerous scales instead of leaves, of which it is destitute. The flowers have a coloured calyx cut into four or five segments, and the corolla is in four or five petals. There is an American species with white flowers. CHAP. VIII.] JASMINEiE. 133 CHAPTER VIII. THE ORDER OLEACE^, OR JASMINES : ILLUSTRATED BY THE COMMON WHITE JASMINE J THE YELLOW JASMINE ; THE PRIVET ; THE PHILLYREA ; THE OLIVE ; THE FRINGE -TREE {Chionanthus Virginica) ; the lilac ; the common ash ; AND THE MANNA OR FLOWERING ASH. This order was established by Jussieu, who dividedit intotwo tribes— Jasminese and Oleinese, which are now very generally considered as distinct orders. I have, however, thought it best to keep them together, as I wish to make as few divisions as possible, to avoid burthening the memory of my readers. All the genera in both tribes agree in their flowers having only two stamens, an ovary with two cells, and two seeds in each cell ; and anthers with two cells, which open with a long slit lengthways. The species of the Ash have no corolla ; but in all the genera where there is one, the fila- ments of the stamens, which are very short, are inserted in it ; and it is generally funnel- shaped — as, for example, the corolla of the Jasmine. Though the ovary is two-celled, and the cells two-seeded, each flower very often only produces one perfect seed. The leaves are generally pinnate. 134 JASMINES. [part I. TRIBE I,— JASMINES. The genus Jasminum is the only one in this tribe which contains plants common in British gardens ; and of all the species contained in it, the common white Jasmine {J. officinale) is perhaps the best known. The flowers are pro- duced in terminal clusters of four or six. The calyx is tubular, with the limb cut into numerous narrow segments ; (see a in Jig. 64 ;) and the Fig. 64. — FtowsR and leaf op the Jasmine. corolla is funnel-shaped, with a spreading limb (h) divided into four or five pointed segments, which are folded over each other, and somewhat twisted in the bud. The two stamens and the style and stigma are enclosed in the corolla; and the fruit is a berry divided into two cells, with one seed in each. There is no albumen CHAP. VIII.] JASMINES. 135 in the seeds. The leaves (c) are impari-pinnate, with the single terminating leaflet larger than the others; and the petioles are articulated. The common yellow Jasmine (J. fruticans) has flowers in terminal clusters of three each, and its leaves are either ternate, that is, with three leaflets, or simple. The branches are angular, and the leaves quite smooth. The Nepaul yellow Jasmine, {J. i^evolutum) has pinnate leaves of five or seven leaflets, which are smooth and shining. The flowers are large and pro- duced in compound corymbs. They are a bright yellow, and very fragrant. The segments of the corolla are obtuse, and the stigma club- shaped. There are above seventy species of Jasmine, more than twenty of which have been introduced into Britain ; but they may be all easily recognised by their flowers, which bear a strong family likeness to each other, and by the petioles of their leaves, which are always articulated or jointed, that is, they will break off the stem without tearing the bark. In other respects the leaves vary exceedingly in this genus, some being simple and others com- pound; and some being opposite, as in the common Jasmine, and others alternate, as in J. revolutum. 136 OLEINEtE. [part I. TRIBE TI.— OLEINE^. This tribe contains numerous genera, among which the most common are the Privet {Ligus- trum)^ Phillyrea, the Olive (Oka), the Fringe- tree {Chionanthus), the Lilac {Syringa), the Ash (Fraximis), and the Flowering or Manna Ash {Ornus). All these genera agree in their general character with Jasminum, except as regards their seeds, which abound in albumen. In the common Privet {Ligustrum vulgare), the flowers, which are produced in terminal compound racemes, have a very short calyx (see a in Jig. 65), w^ith a funnel- shaped corolla, having a w^iJe tube in proportion to the limb (5), which is very short and divided into four segments. The anthers of the stamens and the stigma are seen in the throat of the corolla. The berry is drupe-like, and generally fig. 65.-flow] contains two one-seeded nuts. The of the privet. leaves are simple and opposite. There are many species of Privet, but the handsomest is L. lucidum, the leaves of which are broad and shining, and the panicles of flowers spreading. This tree yields a kind of waxy matter from its leaves and branches when boiled, which is said to be used by the Chinese for candles. CHAP, viii.] OLEINEiE. 137 The Phillyrea is a handsome evergreen shrub, very useful in shrubberies, from its forming a close compact bush of a deep green, which makes a good background to Tree Roses," Almond-trees, Magnolia conspicua, or any other flowering plant that would appear naked if its flowers were not relieved by a back -ground of green. The flowers of the Phillyrea are small and of a greenish white. The fruit is a drupe, containing a two- celled stone or nut, but with seldom more than one perfect seed. The Olive {Oka satlva) has small white flowers, resembling those of the Privet, and a fleshy drupe like a Sloe, with a one or two celled stone or nut. The oil is contained in the fleshy part of the fruit, and the best oil is that which is obtained by crushing the pulp of the fruit without breaking the stone or nut. The Fringe-tree {Chionanthus virginica) differs from the preceding genera in the length of the segments of the limb of its corolla, which is cut into lono: slender shreds like frinoje. In all other respects except that the pulp of the fruit does not contain oil, this genus is closely allied to the Olive. The common Lilac (Syringa vulgaris) has its flowers disposed in a kind of panicled raceme called a thyrsus. The calyx is very small, and obscurely four-toothed (see a in Jig. QQ), and 138 OLEINEiE. [part i. the corolla (b) is funnel-shaped, with a four- parted limb ; the stigma is two-cleft, and both the style and stamens are enclosed in the tube Flower and seed-pods of the Lilac. of the corolla. The fruit is a dry two-celled and two-seeded capsule, which opens with two valves, as shown at c, each valve having a narrow dissepiment down the middle : the shape of the seed is shown at d. The leaves are simple, opposite, and entire ; and the branches are filled with pith, which may easily* be taken out and the branch left hollow like a pipe ; and hence the generic name of Syringa, from Syrinx a pipe. The Ash (Fraxinus excelsior) differs so much from the other genera as to seem scarcely to belong to the same order. The flowers are without any petals, and frequently without any calyx ; and some of them, which are called the female flowers, have no stamens, while others, which are called the males, have no pistil. Some of them, however, have both stamens and pistil. CHAP. VIII.] OLEINE.E. 139 The fruit is what is called a samara or key ; that is, it is furnished with a membrane-like wing so as to resemble a dry leaf. It is two- celled, but very frequently only one-seeded. The shape of the keys, and the manner in which they grow, is shown at a va. Jig. 67; and the leaves, at b. The leaves are opposite and generally pin- nate, with five or six pairs of leaflets ; but there is one species with simple leaves { Fr. simplicifolia ) . The Weeping Ash is only an accidental variety of the common kind. The leaves of the Ash come out late and fall early ; but the tree may easily be recognised when quite bare by the greyness of its bark and its black buds. It will grow in any soil ; but it is injurious to arable land, from its roots spreading widely near the surface. The Manna, or flowering Ash, {Ornus euro- pcBus), differs widely from the common Ash in its flowers, which are white, with a corolla divided into four long narrow segments. The two stamens have long filaments, with a small pistil (c), the stigma of which is notched. The Fig. 67 — American Ash (Fraxinus americana). 140 OLEINEiE. [part i. flowers are produced in great profusion in loose panicles, and they are very ornamental, the samaras and leaves closely resembling those of the common ash. There are several speci es of this genus, which were all formerly included in the genus Fraxinus. The Manna is the sap of the tree, and it is procured by wounding the bark. CHAP. IX.] SOLANACE^. 141 CHAPTER IX. THE ORDER SOLANACE^ : ILLUSTRATED BY THE BITTER-SWEET; GARDEN NIGHTSHADE J POTATO ; EGG-PLANT ; TOMATO ', CAPSI- CUM ; WINTER CHERRY ; CAPE GOOSEBERRY ; THE DEADLY NIGHTSHADE ; LYCIUM, OR DUKE OF ARGYLE's TEA-TREE ; CESTRUM ; VESTIA ; TOBACCO ', PETUNIA ; NIEREMBE^GIA ; SALPIGLOSSIS ; SCHIZANTHUS ; HENBANE ; DATURA ; BRUG- MANSIA ; SOLANDRA ; VERBASCUM ; CELSIA ; NOLANA ; ETC. This large Order is one of those which appear to have been most troublesome to bota- nists, as scarcely any two agree as to the plants to be comprised in it. I have, however, taken it in its most comprehensive sense, as far as popular plants are concerned ; on the same principles as those by which I have been guided throughout ; viz. that it is easier for a beginner to remember a few divisions than a great many ; and that when a student has once learnt what plants are nearly allied to each other, and the general features that con- nect them, it will be comparatively easy to learn the minor distinctions between them. Taking these principles as my guide, I have given the Order Solanaceas as it was formed by Jussieu, adding those plants to it which evidently belong to the several sections, but which have 142 SOLANACEiE. [part i. been discovered since the time of that great naturalist ; and I have divided the Order into four tribes, viz. Solanacese, Nicotianese, Verbas- cinese, and Nolanese. All these plants agree in having the stamens, which are generally five, inserted in the corolla, the calyx and corolla inclosing the ovary, and the calyx remaining on the ripe fruit. TRIBE L— SOLANACE^. The plants included in this tribe are easily recognised by their flowers, which bear a con- siderable resemblance to each other, and by their berry-like fruit, which has always a per- sistent calyx. The corolla is also always folded in the bud ; and the folds, like those of a country woman's clean apron, are often so deeply im- pressed as to be visible in the newly opened flowers. The genera included in this tribe differ widely in their qualities. The genus Solanum is easily recognised by a botanist through all its numerous species by its anthers, which open by two pores like those of the Ericaceae, and which differ in this respect, from the anthers of all the other plants contained in the Order, all of which open by a long slit down each cell. The flowers of all the species of Solanum are of the kind called rotate, or wheel-shaped ; CHAP. IX,] SOLANACE^. 143 Fig. 68. — BrrxER-swEET {Solarium l}ulcainara\. but they are generally cut into five distinct segments: which are sometimes turned back, as in the flower of the Bitter-sweet (S. Dulcamara)^ as shown in^^. 68 a ; and sometimes nearly flat, as in the flower of the common garden Nightshade {S. ni- grum). The berries of the Bitter-sweet (h) are red, and they have a very pretty effect in hedges and wild coppices, where they are produced in great abundance during the latter part of summer and autumn ; and those of the Garden Night- shade are black. Both these plants are poisonous ; but this is by no means the case with all the species of the genus, as the tubers of the potato {S. tuberosum) are, as is well known, whole- some food, and the fruit or apple is not decidedly poisonous ; while the Aubergine, or Egg-plant (