PRESENT TAXONOMIC ATTRIBUTIONS OF THE DINOFLAGELLATE CYSTS AND ACRITARCHS TREATED BY DAVEY, DOWNIE, SARJEANT AND WILLIAMS (1966, 1960) BY LINDA ANN KRUKEWICH AND WILLIAM ANTONY SWITHIN SARJEANT (University of Saskatchewan) Pp 25-54 BULLETIN OF THE BRITISH MUSEUM (NATURAL HISTORY) GEOLOGY Appendix II to Supplement 3 LONDON: 1983 © Trustees of the British Museum (Natural History), 1983 ISSN 0524-644X British Museum (Natural History) Cromwell Road London SW7 5BD PRESENT TAXONOMIC ATTRIBUTIONS OF THE DINOFLAGELLATE CYSTS AND ACRITARCHS TREATED BY DAVEY, DOWNIE, SARJEANT AND WILLIAMS (10966, 1960) By LINDA A. KRUKEWICH and WILLIAM A. S. SARJEANT INTRODUCTION THE major work here reprinted marked the beginning of a new epoch in systematic work on fossil dinoflagellate cysts, in that it embodied not only the first attempt to formulate a separate morphological terminology for these microfossils but also the first extensive application of the ideas of Evitt (1961, 1963), that archaeopyle character and process shape and distribution could be used as prime criteria in taxonomy. In the seventeen years that have elapsed since its publication, there has been a further explosive expansion in our observation and comprehension of fossil dino- flagellate cysts, with a consequent expansion of terminology (see Evitt et al. 1967, William, Sarjeant & Kidson 1978, Sarjeant 1982a). The importance in taxonomy of such additional characters as the shape of process terminations and the direc- tion of their branches, the exact shape of the archaeopyle, the presence and pattern of minute plates (we would now style them “‘paraplates’’) in the mid- ventral region, and the exact relationships of the anterior ventral plates has come to be recognized (see for example Evitt 1967, Lentin & Williams 1976, Sarjeant, 1982b). In contrast, some features - for example, the presence of an ectophragm — are now considered less reliable as means for the separation of taxa. The result has been the reassignment to other genera of many of the species mentioned in the 1966 paper and its first Appendix (1969). The diagnoses of many of these genera and species have been greatly modified, a few taxa have been rejected as junior synonyms and some species, then treated as acritarchs, are now recognized to be dinoflagellate cysts. To simplify understanding of the present taxonomic situation, a checklist is here appended in which the names of all genera and species mentioned in 1966 and in the 1969 Appendix are given in alphabetical order. Any revisions to their diagnoses are noted; and the present names, where different to those cited earlier, are indicated. Where their taxonomic allocation remains a matter for controversy, the alternative ideas are noted. The species placed in 1966 into the acritarch genus Baltisphaertdium are included in proper alphabetical position; if these are still considered to be acritarchs, that fact is likewise noted. Where taxa appeared under different generic names in 1966 and 1969, both names are included in the list. A full reference section, citing all papers referred to in the list, is appended. 28 SECOND APPENDIX TO Two papers by the second author which are currently in press (Sarjeant 1983a,b), and may not be published until after this checklist has appeared, include further revisions to the taxonomy of some of these species mentioned herein. Both are restudies of European type material; the first deals with the Oligocene to Miocene taxa described by Maier (1959) and the second with the Jurassic types of Klement (1960). The imminent revisions are not cited here, to avoid priority problems; but the Maier species concerned are marked with a single asterisk (*), indicating that reference should also be made to Sarjeant (1983a), and the Kle- ment species with a double asterisk (**), suggesting reference to Sarjeant (1983b). CHECKLIST Past Name ACANTHOGONYAULAX Sarjeant, 1966b venusta (Klement, 1960) Sarjeant, 1966b ACHOMOSPHAERA Evitt, 1963 alcicornu. (Eisenack, 1954) Davey & Williams, 1966a grallaeformis (Brosius, 1963) Davey & Williams, 1969 hirundo (Eisenack, 1958) Davey & Williams, 1969 hyperacantha (Deflandre & Cookson, 1955) Davey & Williams, 1969 neptunt (Eisenack, 1958) Davey & Williams, 1966a ramulifera (Deflandre, 1937) Evitt, 1963 vamulifera var. perforata Davey & Williams, 1966a sagena Davey & Williams, 1966a triangulata (Gerlach, 1961) Davey & Williams, 1969 ADNATOSPHAERIDIUM Williams & Downie, 1966c aemulum (Deflandre, 1938) Williams & Downie, 1969 caulleryi (Deflandre, 1938) Williams & Downie, 1969 filamentosum (Cookson & Ejisenack, 1958) Williams & Downie, 1969 multispinosum Williams & Downie, 1966c patulum Williams & Downie, 1966c vittatum Williams & Downie, 1966c Present Name ACANTHAULAX Sarjeant, 1968 [The name Acanthogonyaulax was a jr hom. of a sub- genus of Gonyaulax proposed by Kofoid, 1g11. The subgenus was elevated to generic status by Graham, 1942. ] Acanthaulax venusta (Klement, 1960) Sar- jeant, 1968 Cordosphaeridium? hirundo (Eisenack, 1958) Stover & Evitt, 1978 Spiniferites hyperacanthus (Deflandre & Cook- son, 1955) Cookson & Eisenack, 1974 [Treated by Lentin & Williams, 1973, as a sub- species. ] Rigaudella aemula (Deflandre, 1938) Below, 1982 Rigaudella filamentosa (Cookson & Eisenack, 1958) Below, 1982 Thalassiphora patula (Williams & Downie, 1966c) Stover & Evitt, 1978 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 29 APTEODINIUM Eisenack, 1958* granulatum Eisenack, 1958 maculatum Eisenack & Cookson, 1960 AREOLIGERA Lejeune-Carpentier, 1938; emend. Williams & Downie, 1966c coronata (O. Wetzel, 1933) Lejeune-Carpen- tier, 1938 galea Maier, 1959* lychnea (Maier, 1959) Davey et al., 1969* twistringensis (Maier, 1959) Davey ef al., 1969* BALTISPHAERIDIUM Eisenack, 1958; emend. Downie & Sarjeant, 1963; emend. Eisenack, 1969 armatum (Deflandre, 1937) Downie & Sar- jeant, 1963 asteroideum (Maslov, 1956) Downie & Sar- jeant, 1963 claviculorum (Deflandre, 1938) Downie & Sarjeant, 1963 clavispinulosum Churchill & Sarjeant, 1963 densicomatum (Maier, 1959) Gerlach, 1961* denticulatum (Courtville 7m Deflandre, 1946) Downie & Sarjeant, 1963 ?difforme (Pritchard, 1841) Downie & Sar- jeant, 1963 downier Sarjeant, 1960 echiniplax Churchill & Sarjeant, 1963 Jimbriatum (White, 1842) Sarjeant, 1959 giulsonu (Kufferath, 1950) Downie & Sar- jeant, 1963 horridum (Deflandre, 1937) Downie & Sar- jeant, 1963 huguonioti (Valensi, 1955) Downie & Sar- jeant, 1963 intermedium (O. Wetzel, 1933) Downie & Sarjeant, 1963 longofilum (Maier, 1959) Downie & Sarjeant, 1963* malleoferum (White, 1842) Downie & Sar- jeant, 1963 mariannae (Philippot, 1949) Downie & Sar- jeant, 1963 panniforme Gerlach, 1961 patter (Valensi, 1948) Sarjeant, 1960 paucifurcatum (Cookson & Eisenack, 1961) Downie & Sarjeant, 1965 pectiniforme Gerlach, 1961 Cleistosphaeridium armatum (Deflandre, 1937) Davey, 1969a Regarded as an acritarch. Solisphaeridium claviculorum Deflandre, 1938; emend. Sarjeant, 1968 Regarded as an acritarch. Regarded as an acritarch. Regarded as an acritarch. Acanthaulax? downier (Sarjeant, 1960) Sar- jeant, 1976 Regarded as an acritarch. Regarded as an acritarch. Regarded as an acritarch. Regarded as an acritarch. Cleistosphaertdium huguonioti (Valensi, 1955) Davey, 1969a Hystrichosphaeridium intermedium (O. Wetzel, 1933) Deflandre, 1937 Regarded as an acritarch. Regarded as an acritarch. Achomosphaera mariannae (Philippot, 1949) Stover & Evitt, 1978 Cleistosphaeridium panniforme (Gerlach, 1961) Stover & Evitt, 1978 Lanterna? pattei (Valensi, 1948) Brideaux & Fisher, 1976 Regarded as an acritarch. Areosphaeridium? pectiniforme (Gerlach, 1961) Stover & Evitt, 1978 30 SECOND APPENDIX TO BALTISPHAERIDIUM continued plicatum (Maier, 1959) Downie & Sarjeant, 1963* ?polyceratum Takahashi, 1964 polyozon Brosius, 1963 quaternarium Churchill & Sarjeant, 1963 saturnium (Maier, 1959) Downie & Sarjeant, 1963* seminudum (W. Wetzel, 1952) Downie & Sarjeant, 1963 spiculatum (White, 1842) Downie & Sar- jeant, 1963 stimuliferum (Deflandre, 1938) Sarjeant, 1960 sylheti (Baksi, 1962) Downie & Sarjeant, 1963 telmaticum Churchill & Sarjeant, 1963 tinglewoodense Churchill & Sarjeant, 1963 varispinosum Sarjeant, 1959 whiter (Deflandre & Courteville, 1939) Downie & Sarjeant, 1963 BELODINIUM Cookson & Eisenack, 1960b; emend. Dodekova, 1975 dysculum Cookson & Eisenack, 1960b BROOMEA Cookson & Eisenack, 1958; emend. Lentin & Williams, 1976 Plongicornuta Alberti, 1961 ramosa Cookson & Eisenack, 1958 CALLAIOSPHAERIDIUM Davey & Williams, 1966b; emend. Duxbury, 1980; emend. Below, 1981 asymmetricum (Deflandre & Courtville, 1939) Davey & Williams, 1966b CANNOSPHAEROPSIS O. Wetzel, 1933; emend. Williams & Downie, 1966c; emend. May, 1980; emend. Duxbury, 1980 reticulensis Pastiels, 1948 utinensis O. Wetzel, 1933; emend. Duxbury, 1980 CARPODINIUM Cookson & Eisenack, 1962; emend. Leffingwell & Morgan, 1977 granulatum Cookson & Eisenack, 1962; emend. Leffingwell & Morgan, 1977 Regarded as an acritarch. Cleistosphaeridium? polyozon (Brosius, 1963) Stover & Evitt, 1978 Regarded as an acritarch. Regarded as an acritarch. Regarded as an acritarch. Solisphaeridium stimuliferum (Deflandre, 1938) Staplin, Jansonius & Pocock, 1965 Regarded as an acritarch. Hystrichosphaeridium sylheti Baksi, 1962 Regarded as an acritarch. Regarded as an acritarch. Sentusidinium? varispinosum (Sarjeant, 1959) Sarjeant & Stover, 1978 Cometodinium? white: (Deflandre & Courte- ville, 1939) Stover & Evitt, 1978 Imbatodinium longicornutum (Alberti, 1961) Dorhofer & Davies, 1980 Adnatosphaeridium reticulense (Pastiels, 1948) De Coninck, 1969 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 31 CLEISTOSPHAERIDIUM Davey ¢e al., 1966 ancoriferum (Cookson & Eisenack, 1960a) Davey et al., 1966; emend. Cookson & Eisenack, 1968 ashdodense Rossignol, 1962 danicum (W. Wetzel, 1952) Davey et al., 1969 disjunctum Davey et al., 1966 diversispinosum Davey et al., 1966 echinotdes (Maier, 1959) Davey et al., 1969* ehrenbergi (Deflandre, 1947) Davey et al., 1969 ?flexuosum Davey et al., 1966 heteracanthum (Deflandre & Cookson, 1955) Davey et al., 1966 israelianum (Rossignol, 1962) Davey et al., 1966 leve (Maier, 1959) Davey et al., 1969* lumectum (Sarjeant, 1960a) Davey et al., 1969 machaerophorum (Deflandre & Cookson, 1955) Davey et al., 1966 multifurcatum (Deflandre, 1937) Davey et al., 1969 Poligacanthum (W. Wetzel, 1952) Davey et al., 1969 pectiniforme (Gerlach, 1961) Davey et al., 1969 pilosum (Ehrenberg, 1843) Davey et al., 1966 polypes (Cookson & Eisenack, 1962) Davey, 1g69a polytrichum (Valensi, 1947) Davey et al., 1969 spiralisetum (De Wit, 1943) Davey et al., 1969 tiara (Klumpp, 1953) Davey et al., 1969 tribuliferum (Sarjeant, 1962b) Davey et al., 1969 COMETODINIUM Deflandre & Courteville, 1939 obscurum Deflandre & Courteville, 1939 CORDOSPHAERIDIUM Eisenack, 1963b; emend. Davey, 1969c; emend. Sarjeant, 1981 cantharellum (Brosius, 1963) Gocht 1969 [Wall, 1967 considered this species to be a jr syn. of Lingulodinium machaerophorum} [Reid, 1974, considered this species to be a gr syn. of Lingulodinium machaerophorum| Heterosphaeridium? heteracanthum (Deflandre & Cookson, 1955) Eisenack & Kjellstrom, 1971 } Operculodinium israelianum (Rossignol, 1962) Wall, 1967 Lingulodinium machaerophorum (Deflandre & Cookson, 1955) Wall, 1967 Areosphaeridium? pectiniforme (Gerlach, 1961) Stover & Evitt, 1978 Sentusidinium pilosum (Ehrenberg, 1843) Sar- jeant & Stover, 1978; emend. Erkmen & Sarjeant, 1980 Bacchidinium polypes (Cookson & Eisenack, 1962) Davey, 1979 Holotype lost; exact character and sys- tematic status problematical. Tityrosphaeridium cantharellum (Brosius, 1963) Sarjeant, 1981 32 SECOND APPENDIX TO CORDOSPHAERIDIUM continued ?cracenospinosum Davey & Williams, 1966b diktyoplokus Klumpp, 1953 diktyoplokus subsp. latum Klump, 1953 divergens (Eisenack, 1954) Eisenack, 1963b eotnodes (Eisenack, 1954) Eisenack, 1963b ?erectum (Manum & Cookson, 1964) Davey & Williams, 1969 extlimurum Davey & Williams, 1966b ?fasciatum Davey & Williams, 1966b jibrospinosum Davey & Williams, 1966b floripes (Deflandre & Cookson, 1955) Eisen- ack, 1963b floripes subsp. breviradiatum (Cookson & Eisenack, 1961b) Eisenack, 1963b gracile (Eisenack, 1954) emend. Davey & Williams, 1966b latispinosum Davey & Williams, 1966b microtrriainum (Klump, 1953) Eisenack, 1963b multispinosum Davey & Williams, 1966b CTENIDODINIUM Defiandre, 1938; emend. Sarjeant, 1966b; emend. Sarjeant, 1975 ornatum (Eisenack, 1935) Deflandre, 1938 tenellum Deflandre, 1938 CYCLONEPHELIUM Deflandre & Cookson, 1955; emend. Williams & Downie, 1966c; emend. Ioannides ef al., 1976; emend. Sar- jeant & Stover, 1978; emend. Stover & Evitt, 1978 compactum Deflandre & Cookson, 1955 divaricatum Williams & Downie, 1966c exuberans Deflandre & Cookson, 1955 ordinatum Williams & Downie, 1966c pastielsia Deflandre & Cookson, 1955 Achomosphaera? cracenospinosa (Davey & Williams, 1966b) Sarjeant, 1981 Areosphaeridum diktyoplokus (Klump, 1953) Eaton, 1971 Areosphaeridium diktyoplokus subsp. latum (Klumpp, 1953) Lentin & Williams, 1973 Operculodinium divergens (Eisenack, 1954) Stover & Evitt, 1978 Kleithnriasphaeridium __eoinodes 1958) Davey, 1974 Cleistosphaeridium? erectum (Manum & Cook- son, 1964) Stover & Evitt, 1978 Hystrichosphaerina? exilimura (Davey & Williams, 1966b) Sarjeant, 1981 Kletthriasphaeridium fasciatum (Davey & Williams, 1966b) Davey, 1974 Emmetrocysta? fibrospinosa (Davey & Williams, 1966b) Sarjeant, 1981 Homotryblium floripes (Deflandre & Cookson, 1955) Stover, 1975 Homotrybliium floripes subsp. breviradiatum (Cookson & Eisenack, 1961b) Lentin & Williams, 1977 Tityrosphaeridium gracilis (Eisenack, 1954 emend. Davey & Williams, 1966b) Sar- jeant, 1981 Titryosphaeridium? latisbinosum (Davey & Williams, 1966b) Sarjeant, 1981 [Placed earlier into Achilleodinium by Bujak et al., 1980; this may prove to be a more correct assignment. ] Achomosphaera microtriaina (Klump, 1953) emend. Sarjeant, 1981 Amphorosphaeridum? multispinosum (Davey & Williams, 1966b) Sarjeant, 1981 (Eisenack, Glaphyrocysta divaricata (Williams & Downie, 1966c) Stover & Evitt, 1978 Glaphyrocysta exuberans (Deflandre & Cook- son, 1955) Stover & Evitt, 1978 Glaphyrocysta ordinata (Williams & Downie, 1966c) Stover & Evitt, 1978 Glaphyrocysta pastielsu (Deflandre & Cookson, 1955) Stover & Evitt, 1978 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 33 CYMATIOSPHAERA O. Wetzel, 1933; emend. Deflandre, 1954 membranacea (Philippot, 1949) Davey & Williams, 1969 DEFLANDREA Eisenack, 1938; emend. Williams & Downie, 1966c; emend. Lentin & Williams, 1976 denticulata Alberti, 1959 oebisfeldensis Alberti, 1959 phosphoritica Eisenack, 1938 phosphoritica subsp. australis Cookson & Eisenack, 1961a phosphoritica subsp. phosphoritica Eisenack, 1938 wardenensis Williams & Downie, 1966c DICHADOGONY AULAX Sarjeant, 1966a culmula (Norris, 1965) Loeblich & Loeblich, 1968 pannea (Norris, 1965) Sarjeant, 1969 schizoblatum (Norris, 1965) Sarjeant, 1969 DINGODINIUM Cookson & Eisenack, 1958 Palbertu Sarjeant, 1966c jurassicum Cookson & Eisenack, 1958 DIPHYES Cookson, 1965; emend. Davey & Williams, 1966b colligerum (Deflandre & Cookson, 1955) Cookson, 1965 ?monstruosum (Tasch, 1964) Davey & Williams, 1969 DOIDY X Sarjeant, 1966b anaphrissa Sarjeant, 1966c DUOSPHAERIDIUM Davey & Williams, 1966b nudum (Cookson, 1965) Loeblich & Loe- blich, 1968 EISENACKIA Deflandre & Cookson, 1955; emend. Sarjeant, 1966b; emend. McLean, 1973; emend. Stover & Evitt, 1978 crassitabulata Deflandre & Cookson, 1955; emend. McLean, 1973 Regarded as an acritarch. Ceratiopsis wardenensis (Williams & Downie, 1966c) Bujak et al., 1980 [Considered by Lentin & Williams, 1973 to be ajr syn. of Ctenidodinium| [Transferred to Ctenidodinium by Lentin & Williams, 1973. ] [Transferred to Ctentdodinium by Lentin & Williams, 1973.] [Transferred to Ctentdodinium by Lentin & Williams, 1973.] Corontfera monstruosa (Tasch, 1964) Stover & Evitt, 1978 Shown by Sarjeant & Stover, 1978 to bea jr syn. of APTEA Eisenack, 1958; emend. Davey & Verdier, 1974; emend. Dorhofer & Davies, 1980 Aptea anaphrissa (Sarjeant, 1966c) Stover & Sarjeant, 1978 34 SECOND APPENDIX TO EXOCHOSPHAERIDIUM Davey et al., 1966; emend. Davey, 1969c palmatum (Deflandre & Courtville, 1939) Davey et al., 1969 phragmites Davey et al., 1966 pseudhystrichodinium (Deflandre, 1937) Davey et al., 1969 striolatum (Deflandre, 1937) Davey, 1969a FROMEA Cookson & Eisenack, 1958 amphora Cookson & Eisenack, 1958 GARDODINIUM Alberti, 1961 etsenacku Alberti, 1961 GLY PHANODINIUM Drugg, 1964 facetum Drugg, 1964 GONYAULACYSTA _ Deflandre, 1964; emend. Sarjeant, 1969; emend. Stover & Evitt, 1978; emend. Sarjeant, 1982a Paceras (Eisenack, 1958) Sarjeant, 1966b aculeata (Klement, 1960) Sarjeant, 1969 aichmetes Sarjeant, 1966b amabilis Deflandre, 1939 ambigua (Deflandre, 1939) Sarjeant, 1969 apionis (Cookson & Ejisenack, 1958) Sar- jeant, 1969 aptiana (Deflandre, 1935) Sarjeant, 1966b axicerastes Sarjeant, 1966b cassidata (Eisenack & Cookson, 1960) Sar- jeant, 1966b cladophora (Deflandre, 1938) Sarjeant, 1966b clathrata (Cookson & Eisenack, 1g60b) Sar- jeant, 1966b confusa (Vozzhenikova, 1967) Sarjeant, 1969 ?cornigera (Valensi, 1953) Sarjeant, 1966b crassicornuta (Klement, 1960) Sarjeant, 1966b cretacea (Neale & Sarjeant, 1962) Sarjeant, 1969 crispa (W. Wetzel, 1967) Sarjeant, 1969 dangeardu Sarjeant, 1968 Corontfera striolata (Deflandre, 1937) Stover & Evitt, 1978 [Davey, 1974, considered this species to be a jv syn. of Gardodinium trabeculosum (Gocht, 1959) Alberti, 1961.] Acanthaulax? aceras (Eisenack, 1958) Stover & Evitt, 1978 Acanthaulax aculeata (Klement, 1960) Stover & Evitt, 1978 Millioudodinium aichmetes (Sarjeant, 1966b) Stover & Evitt, 1978 Leptodinium? amabile (Deflandre, 1939) Sar- jeant, 1969 Rhynchodiniopsis ambigua (Deflandre, 1939) Sarjeant, 1982a Millioudodinium apionis (Cookson & Eisen- ack, 1958) Stover & Evitt, 1978 Rhynchodiniopsis aptiana Deflandre, 1935 Rhynchodiniopsis cladophora (Deflandre, 1938) Below, 1981 Leptodinium clathratum (Cookson & Eisenack, 1960b) Sarjeant, 1969 Rhynchodiniopsis? confusa (Vozzhennikova, 1967) Sarjeant, 1982a Hystrichogonyaulax cornigera (Valensi, 1953) Sarjeant, 1969 Rhynchodiniopsis crassicornuta 1960) Below, 1981 (Klement, Hystrichogonyaulax crispa (W. Wetzel, 1967) Sarjeant, 1980b Tubotuberella dangeardi (Sarjeant, 1968) Stover & Evitt, 1978 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 35 GONY AULACYSTA continued diaphanis (Cookson & Eisenack, 1958) Sar- jeant, 1969 dictyophora (Deflandre, 1938) Sarjeant, 1967 edwardsi1 (Cookson & Eisenack, 1958) Clarke & Verdier, 1967 eisenackit (Deflandre, 1938) Dodekova, 1967; emend. Sarjeant, 1982a episoma Sarjeant, 1966b eumorpha (Cookson & Eisenack, 1960b) Sar- jeant, 1966b extlicristata Davey, 1969a fetchamensis Sarjeant, 1966b freakei (Sarjeant, 1963) Sarjeant, 1966a giusepper (Morgenroth, 1966) Sarjeant, 1969 gongylos Sarjeant, 1966b gottisaa Dupin, 1968 granulata (Klement, 1960) Sarjeant, 1969** granuligera (Klement, 1960) Sarjeant, 1969** hadra Sarjeant, 1966b helicoidea (Eisenack & Cookson, 1960) Sar- jeant, 1966b hyaloderma (Deflandre, 1939) Sarjeant, 1967 hyalodermopsis (Cookson & Eisenack, 1958) Sarjeant, 1969 jurassica (Deflandre, 1938) Norris & Sar- jeant, 1965; emend. Sarjeant, 1982a kostromiensis (Vozzhennikova, 1967) Sar- jeant, 1969 longicornis (Downie, 1957) Sarjeant, 1969 Pmamillifera (Deflandre, 1939) Sarjeant, 1969 margaritifera (Cookson & Eisenack, 1960a) Sarjeant, 1966b microceras (Eisenack, 1958) Clarke & Ver- dier, 1967 milliouds (Sarjeant, 1963) Sarjeant, 1966b monacantha (Deflandre, 1935) Sarjeant, 1967 Cribroperidinium diaphanis (Cokson & Eisen- ack, 1958) Stover & Evitt, 1978 Aldorfia dictyophora (Deflandre, 1938) Stover & Evitt, 1978 Cnibroperidinium edwardsu (Cookson & Eisenack, 1958) Davey, 1969a Rhynchodiniopsis episoma (Sarjeant, 1966b) Sarjeant, 1982a Leptodinium eumorphum (Cookson & Eisen- ack, 1960b) Sarjeant, 1969 Cribroperidinium exilicristatum (Davey, 1969a) Stover & Evitt, 1978 Millioudodinium __fetchamense _—_ (Sarjeant, 1966b) Stover & Evitt, 1978 Leptodinium freakez (Sarjeant, 1963) Sarjeant, 1969 Rhynchodiniopsis? giusepper (Morgenroth, 1966) Sarjeant, 1982a Rhynchodiniopsis gongylos (Sarjeant, 1966b) Sarjeant, 1982a Milltoudodinium? gottisi (Dupin, 1968) Stover & Evitt, 1978 Rhynchodiniopsis granulata (Klement, 1960) Sarjeant, 1982a Rhynchodiniopstis granuligera (Klement, 1960) Sarjeant, 1982a Rhynchodiniopsis hadra (Sarjeant, 1966b) Sar- jeant, 1982a Rhynchodiniopsis? hyaloderma (Deflandre, 1939) Sarjeant, 1982a Rhynchodiniopsis hyalodermopsis (Cookson & Eisenack, 1958) Sarjeant, 1982a Nelchinopsis kostromiensis (Vozzhennikova, 1967) Wiggins, 1972 [The transfer of this species to Nelchinopsis was rejected by Duxbury, 1977] Millioudodinium longicornis (Downie, 1957) Sarjeant, 1982a Rhynchodiniopsis? mamillifera (Deflandre, 1939) Sarjeant, 1982a Impagidinium margaritiferum (Cookson & Eisenack, 1960a) Stover & Evitt, 1978 Leptodinium millioudu (Sarjeant, 1963) Sar- jeant, 1969 Milltoudodinium monacanthum (Deflandre, 1935) Stover & Evitt, 1978 36 SECOND APPENDIX TO GON Y AULACYSTA continued muderongensis (Cookson & Eisenack, 1958) Sarjeant, 1966b ?nannotrix (Deflandre, 1939) Sarjeant, 1969 Pnealer (Sarjeant, 1962b) Sarjeant, 1966b nuciformis (Deflandre, 1938) Sarjeant, 1968 obscura (Lejeune-Carpentier, 1946) Sar- jeant, 1969 orthoceras (Eisenack, 1958) Sarjeant, 1966b pachyderma (Deflandre, 1938) Sarjeant, 1969 palla Sarjeant, 1966b parorthoceras Davey, 1968 perforans (Cookson & Eisenack, 1958) Sar- jeant, 1969 ?porosa (Lejeune-Carpentier, 1946) Sar- jeant, 1966b pyra (Drugg, 1967) Sarjeant, 1969 sarjeantit (Vozzhennikova, 1967) Sarjeant, 1969 scarburghensis Sarjeant, 1964 scotti1 (Cookson & Eisenack, 1958) Sarjeant, 1969 serrata (Cookson & Eisenack, 1958) Sar- jeant, 1969 Ptenuiceras (Eisenack, 1958) Sarjeant, 1969 tenuicornuta (Cookson & Eisenack, 1962) Sarjeant, 1969 tenuitabulata (Gerlach, 1961) De Coninck, 1969 ?transparens (Sarjeant, 1959) Sarjeant, 1969 wetzelu (Lejeune-Carpentier, 1939) Sarjeant, 1969; emend. Lejeune-Carpentier & Sar- jeant, 1981 whiter Sarjeant, 19g66b HELIODINIUM Alberti, 1961 patriciae Neale & Sarjeant, 1962 voigtt Alberti, 1961 Cribroperidinium muderongense (Cookson & Eisenack, 1958) Davey, 1969a Rhynchodiniopsis? nannotrix (Deflandre, 1939) Sarjeant, 1982a Hystrichogonyaulax? nealei (Sarjeant, 1962b) Sarjeant, 1969 Millioudodinium nuciformis (Deflandre, 1938) Sarjeant, 1982a Millioudodinium obscurum (Lejeune-Carpen- tier, 1946; emend. Lejeune-Carpentier & Sarjeant, 1981) Sarjeant, 1982a Cribroperidinium orthoceras (Eisenack, 1958) Davey, 1969a Ctenidodinium? pachydermum 1938) Gocht, 1970 Rhynchodiniopsis palla (Sarjeant, 1966b) Sar- jeant, 1982a Millioudodinium parorthoceras (Davey, 1968) Stover & Evitt, 1978 Rhynchodiniopsis perforans (Cookson & Eisen- ack, 1958) Sarjeant, 1982a Leptodinium porosum (Lejeune-Carpentier, 1946) Sarjeant, 1969; emend. Lejeune- Carpentier & Sarjeant, 1981. [Conceivably a fossil motile dinoflagellate! ] Cribroperidinium? pyrum (Drugg, 1967) Stover & Evitt, 1978 Rhynchodimiopsis sarjeantir (Vozzhennikova, 1967) Sarjeant, 1982a (Deflandre, Miullioudodinium? scottii (Cookson & Eisenack, 1958) Stover & Evitt, 1978 Hystrichogonyaulax serrata (Cookson & Eisen- ack, 1958) Stover & Evitt, 1978 Diacanthum tenuiceras (Eisenack, 1958) Stover & Evitt, 1978 Leptodinium? tenuicornutum Cookson & Eisenack, 1962 Millioudodinium tenuitabulatum (Gerlach, 1961) Stover & Evitt, 1978 Rhynchodiniopsis white: (Sarjeant, 1966b) Sar- jeant, 1982a [Considered by Lentin & Williams, 1975 to be a jr syn. of Hystrichodinium Deflandre, 1935. This judgement, and the resultant transfer of its two constituent species to Hystrichodinium, are not accepted by us. } DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) a7) HEMICYSTODINIUM Alberti, 1961 zohary1 (Rossignol, 1962) Wall, 1967 HESLERTONIA Sarjeant, 1966b; emend. Duxbury, 1980 heslertonensis (Neale & Sarjeant, 1962) Sar- jeant, 1966b HOMOTRYBLIUM Davey & Williams, 1966b pallidum Davey & Williams, 1966b tenuispinosum Davey & Williams, 1966b HYSTRICHODINIUM ODeflandre, 1935; emend. Clarke & Verdier, 1967 compactum Alberti, 1961 oligacanthum Deflandre & Cookson, 1955 pulchrum Deflandre, 1935 ramoides Alberti, 1961 HYSTRICHOGONY AULAX Sarjeant, 1969 cornigera (Valensi, 1953) Sarjeant, 1969 nealet (Sarjeant, 1962b) Sarjeant, 1969 HYSTRICHOKOLPOMA Klumpp, 1953; emend. Williams & Downie, 1966a cinctum Klumpp, 1953 clavigerum (Deflandre, 1937) Williams & Downie, 1969 etsenacki Williams & Downie, 1966a eisenacki var. turgidum Williams & Downie, 1966a ferox (Deflandre, 1937) Davey, 1969a poculum Maier, 1959* rigaudae Deflandre & Cookson, 1955 sequanaportus Deflandre & Deflandre- Rigaud, 1958 unispinum Williams & Downie, 1966a Pxupheum (Maier, 1959) Davey & Williams, 1969* HYSTRICHOSPHAERA O. Wetzel, 1933 ex Deflandre, 1937 buccina Davey & Williams, 1966a cingulata (O.Wetzel, 1933) Deflandre, 1954 cingulata var. reticulata Davey & Williams, 1966a cornuta Gerlach, 1961 Jr syn. of Polysphaeridium: see Bujak et al., 1980 Polysphaeridium zoharyt (Rossignol, 1962) Bujak et al., 1980 Florentinia clavigera (Deflandre, 1937) Davey & Verdier, 1973; emend. Davey & Verdier, 1976 [Considered by Damassa, 1979, to be a jr syn. of AHystrichokolpoma cinctum Klumpp, 1953-] Hystrichokolpoma cinctum subsp. turgidum Lentin & Williams, 1973 Florentinia ferox (Deflandre, 1937) Duxbury, 1980 Shown by Sarjeant, 1970 to be a jr syn. of Spiniferites Mantell, 1850 Shown by Lentin & Williams, 1973, to be a jv syn. of Spiniferites pseudofurcatus (Klumpp, 1953) Sarjeant, 1970; emend. Sarjeant, 198T. Pterodinium cingulatum (O. Wetzel, 1933) Below, 1981 Pterodinium cingulatum subsp. reticulatum (Davey & Williams, 1966a) Lentin & Williams, 1981 Spiniferites cornutus (Gerlach, 1961) Sarjeant, 1970 38 SECOND APPENDIX TO HY STRICHOSPHAERA continued cornuta var. laevimura Davey & Williams, 1966a crassimurata Davey & Williams, 1966a crasstpellis Deflandre & Cookson, 1955 leptoderma (Maier, 1959) Davey & Williams, 1969* monolis Davey & Williams, 1966a perforata Davey & Williams, 1966a vamosa (Ehrenberg, 1838) O. Wetzel, 1933 ramosa var. gracilis Davey & Williams, 1966a vamosa var. granosa Davey & Williams, 1966a vamosa var. granomembranacea Davey & Williams, 1966a ramosa var. membranacea (Rossignol, 1964) Davey & Williams, 1966a rvamosa var. multibrevis Davey & Williams, 1966a vamosa var. vamosa (Ehrenberg, 1838) Davey & Williams, 1966a vamosa var. reticulata Davey & Williams, 1966a HYSTRICHOSPHAERIDIUM Deflandre, 1937; emend. Davey & Williams, 1966b Paquitanicum Deunff, 1961 arborispinum Davey & Williams, 1966b Parundum Eisenack & Cookson, 1960 bowerbanku Davey & Williams, 19g66b claviferum (Wilkinson, 1849) Deflandre, 1946 ?clavigerum (Deflandre, 1937) Lejeune- Carpentier, 1940 costatum Davey & Williams, 1966b [Treated as a subspecies by Lentin & Williams, 1973.] Spiniferites crassimuratus (Davey & Williams, 1966a) Sarjeant, 1970 Achomosphaera crasstpellis (Deflandre & Cook- son, 1955) Stover & Evitt, 1978 Spiniferites monolis (Davey & Williams, 1966a) Sarjeant, 1970 Spiniferites perforatus (Davey & Williams, 1966a) Sarjeant, 1970 Spiniferites ramosus (Ehrenberg, 1838) Man- tell, 1857 Spiniferites ramosus var. gracilis (Davey & Williams, 1966a) Corradini, 1973 [Treated by Lentin & Williams, 1981, as a sub- species] Spiniferites ramosus var. granosus (Davey & Williams, 1966a) Corradini, 1973 [Treated by Lentin & Williams, 1981, as a sub- species] Spiniferites ramosus var. granomembranaceus (Davey & Williams, 1966a) Corradini, 1973 [Treated by Lentin & Williams, 1981, as a Subspecies] Spiniferites membranaceus (Rossignol, 1964) Sarjeant, 1970 Spiniferites ramosus var. multibrevis (Davey & Williams, 1966a) Davey & Verdier, 1971 [Treated by Lentin & Williams, 1981, as a subspecies] Spiniferites ramosus var. ramosus (Ehrenberg, 1838) Loeblich & Loeblich, 1966 [Treated by Lentin & Williams, 1981, as a subspe- cies] Spiniferites ramosus var. reticulatus (Davey & Williams, 1966a) Davey & Verdier, 1971 [Treated by Lentin & Williams, 1981, as a subspecies] Litosphaeridium arundum (Eisenack & Cook- son, 1960) Davey, 1979 Florentinia clavigera (Deflandre, 1937) Davey & Verdier, 1973; emend. Davey & Verdier, 1976 Compositosphaeridium costatum (Davey & Williams, 1966b) Dodekova, 1974 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 39 HYSTRICHOSPHAERIDIUM continued deanei Davey & Williams, 1966b Pgliwicense Mack6 1957 hill (Merrill, 1895) Sarjeant, 1964 irvegulare (Merrill, 1895) Sarjeant, 1964 patulum Davey & Williams, 1966b polyplasium Maier, 1959* radiculatum Davey & Williams, 1966b veadet Davey & Williams, 1966b salpingophorum Deflandre, 1935; emend. Davey & Williams, 1966b sheppeyense Davey & Williams, 1966b simplicispinum Davey & Williams, 1966b stellatum Maier, 1959* tridactylites Valensi, 1955 tubtferum (Ehrenberg, 1838) Deflandre, 1937; emend. Davey & Williams, 1966b tubiferum var. brevispinum Davey & Williams, 1966b HYSTRICHOSPHAEROPSIS Deflandre, 1935; emend. Sarjeant, 1966b; emend. Gocht, 1976 borussica Eisenack, 1954 ovum Deflandre, 1935 wetzeli (Deflandre, 1937) Sarjeant, 1966b Florentinia deanei (Davey & Williams, 1966b) Davey & Verdier, 1973 Hystrichosphaeridium speciale (Merrill, 1895) Lentin & Williams, 1981. [At the time of the transfer, this name was a jr hom. of Hystrichosphaeridium irregulare Pocock, 1962; however, the latter species was transferred to Oligosphaeridium by Davey & Williams im Davey et al., 1969. Never- theless, Lentin & Williams (1981) con- sidered that the name H. irregulare (Mer- rill) must still be considered a 77 hom. and accordingly proposed the above substi- tute name. Though Art. 55 of the Inter- national Code of Botanical Nomenclature would require reinstatement of the name irregulare (Merrill) if that species be trans- ferred to any genus other than Oligo- sphaeridium, their procedure appears correct. | Florentinia radiculata (Davey & Williams, 1966b) Davey & Verdier, 1976 Kleithriasphaeridium reader (Davey & Williams, 1966b) Davey & Verdier, 1976 Shown by Stover & Evitt, 1978, to be a 77 syn. of Hystrichosphaeridium pseudorecurva- tum Morgenroth, now Melitasphaeridium pseudorecurvatum (Morgenroth, 1966) Bujak 7m Bujak et al., 1980. Kleithriasphaeridium simplicispinum (Davey & Williams, 1966b) Davey, 1974 Florentinia tridactylites (Valensi, 1955) Dux- bury, 1980 [Treated as a subspecies by Lentin & Williams, 1973. ] Rottnestia borussica (Eisenack, 1954) Cookson & Eisenack, 1961 Spiniferites wetzelu (Deflandre, 1937) Sarjeant, 1970 40 SECOND APPENDIX TO LEPTODINIUM Klement, 1960; emend. Sarjeant, 1982a aceras (Eisenack, 1958) Sarjeant, 1969 aculeatum Wall, 1967 alectrolophum Sarjeant, 1966b amabilis (Deflandre, 1939) Sarjeant, 1969 arcuatum Klement, 1960** clathratum (Cookson & Ejisenack, 1I9g60b) Sarjeant, 1969 crassinervum (Deflandre, 1939) Sarjeant, 1969 ?delicatum (Davey, 1969a) Sarjeant, 1969 eumorphum (Cookson & Eisenack, 1960b) Sarjeant, 1969 freakei (Sarjeant, 1963) Sarjeant, 1969 maculatum Cookson & Eisenack, 1961 margaritiferum (Cookson & Eisenack, 1961) Sarjeant, 1969 membranigerum Gerlach, 1961 millioudi (Sarjeant, 1963) Sarjeant, 1969 mirabile Klement, 1960** mosaicum (Downie, 1957) Sarjeant, 1969 paradoxum Wall, 1967 sphaericum Wall, 1967 strialatum Wall, 1967 striatum (Clarke & Verdier, 1967) Sarjeant, 1969 subtile Klement, 1960 Ptenuicornutum Cookson & Ejisenack, 1962 LITOSPHAERIDIUM Davey & Williams, 1966b; emend. Davey & Verdier, 1973 ?crassipes (Reade, 1839) Davey & Williams, 1969 ?flosculus (Deflandre, 1937) Davey & Williams, 1969 Pinversibuccinum Davey & Williams, 1966b siphoniphorum (Cookson & Eisenack, 1958) Davey & Williams, 1966b Ptruncigerum (Deflandre, 1937) Davey & Williams, 1969 MEIOUROGONY AULAX Sarjeant, 1966b bulloidea (Cookson & Eisenack, 1960b) Sar- jeant, 1969 caytonensis (Sarjeant, 1959) Sarjeant, 1969 Acanthaulax? aceras (Eisenack, 1958) Stover & Evitt, 1978 Impagidinium aculeatum (Wall, 1967) Lentin & Williams, 1981 Impagidinium alectrolophum (Sarjeant, 1966b) Stover & Evitt, 1978 Leptodinium amabile (Deflandre, 1939) Sar- jeant, 1969 Impagidinium maculatum (Cookson & Eisen- ack, 1961) Stover & Evitt, 1978 Impagidinium margaritiferum (Cookson & Eisenack, 1961) Stover & Evitt, 1978 Impagidinium? membranigerum (Gerlach, Ig6r) Stover & Evitt, 1978 Impagidinium paradoxum (Wall, 1967) Stover & Evitt, 1978 Impagidinium sphaericum (Wall, 1967) Lentin & Williams, 1981 Impagidinium strialatum (Wall, 1967) Stover & Evitt, 1978 Impagidinium striatum (Clarke & Verdier, 1967) Stover & Evitt, 1978 Florentinia? flosculus (Deflandre, 1937) Lentin & Williams, 1981 Paucisphaeridium inversibuccinum (Davey & Williams, 1966b) Bujak et al., 1980 Cordosphaeridium truncigerum (Deflandre, 1937) De Coninck, 1975 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 41 MEIOUROGONY AULAX continued ?cristulata (Sarjeant, 1959) Sarjeant, 1969 decapitata (W. Wetzel, 1967) Sarjeant, 1969; emend. Sarjeant, 1980 superornata (W. Wetzel, 1967) Sarjeant, 1969; emend. Sarjeant, 1980 valensi Sarjeant, 1966b MEMBRANILARNACIA Eisenack, 19634; emend. Williams & Downie, 1966c leptoderma (Cookson & Eisenack, 1958) Eisenack, 1963a reticulata Williams & Downie, 1966c MICRODINIUM Cookson & Eisenack, 1960a; emend. Sarjeant, 1966b; emend. Stover & Evitt, 1978 ornatum Cookson & Eisenack, 1960a setosum Sarjeant, 1966b MUDERONGIA Cookson & Eisenack, 1958 mcwhae1 Cookson & Eisenack, 1958 staurota Sarjeant, 1966c NEMATOSPHAEROPSIS Deflandre & Cookson, 1955; emend. Williams & Downie, 1966c balcombiana Deflandre & Cookson, 1955 NETRELYTRON Sarjeant, 1961; emend. Sarjeant, 1966c jurassicum (Alberti, 1961) Sarjeant, 1966c stegastum Sarjeant, 1961; emend. Sarjeant, 1966c trinetron Sarjeant, 1966c ODONTOCHITINA Deflandre, 1935; emend. Davey, 1970 operculata (O. Wetzel, 1933) Deflandre & Cookson, 1955 OLIGOSPHAERIDIUM Davey & Williams, 1966b albertense (Pocock, 1962) Davey & Williams, 1969 anthophorum (Cookson & Ejisenack, 1958) Davey, 1969a [Considered by De Coninck, 1969 and Gocht, 1969 to be a 77 syn. of Membranilarnacia ursulae (Morgenroth, 1966) De Coninck, 1969 now Eatonicysta ursulae (Morgen- roth, 1966) Stover & Evitt, 1978.] (Wiggins, 1975, considered this a 77 syn. of Kalyptea Cookson & Ejisenack, 1960b. This judgement was rejected by Sarjeant, 1976.] Kalyptea jurassica Alberti, 1961. [Since this species was not validly transferred in 1966 (see Lentin & Williams, 1981) it must be retained in Kalyptea.] [The transfer of this species to Kalyptea by Wiggins, 1975, was rejected by Sarjeant, 1976. } [The transfer of this species to Kalyptea by Wiggins, 1975, was rejected by Sarjeant, 1976. } 42 SECOND APPENDIX TO OLIGOSPHAERIDIUM continued ?asterigerum (Gocht, 1959) Davey & Williams, 1969 ?coelenteratum (Tasch, 1964) Davey & Williams, 1969 complex (White, 1842) Davey & Williams, 1969 dictyophorum (Cookson & Eisenack, 1958) Davey & Williams, 1969 ?dispare (Tasch, 1964) Davey & Williams, 1969 ?trregulare (Pocock, 1962) Davey & Williams, 1969 macrotubulum (Neale & Sarjeant, 1962) Davey & Williams, 1966b ?paradoxum (Brosius, 1963) Davey & Williams, 1969 ?perforatum (Gocht, 1959) Williams, 1969 prolixospinosum Davey & Williams, 1966b pulcherrimum (Deflandre & Cookson, 1955) Davey & Williams, 1966b reniforme (Tasch, 1964) Davey, 1969a Davey & reticulatum Davey & Williams, 1966b vasiformum (Neale & Sarjeant, 1962) Davey & Williams, 1966b PARANETRELYTRON Sarjeant, 1966c strongylum Sarjeant, 1966c PAREODINIA Deflandre, 1947; emend. Wig- gins, 1975; emend. Stover & Evitt, 1978 ceratophora Deflandre, 1947; emend. Gocht, 1970 Shown by Stover & Evitt, 1978, to be a 77 syn. of Oligosphaeridium irregulare (Pocock, 1962) Davey & Williams, 1969 Shown by Stover & Evitt, 1978, to be a j7 syn. of Oligosphaeridium irregulare (Pocock, 1962) Davey & Williams, 1969 (Considered by McIntyre & Brideaux, 1980, to bea jr syn. of Olugosphaeridium vastfor- mum (Neale & Sarjeant, 1962) Davey & Williams, 1966b; the latter species is in turn considered a jr syn. of Oligosphaeri- dium asterigerum (Gocht, 1959) Davey & Williams, 1969, by Duxbury, 1977.] Bipolaribucina paradoxa (Brosius, 1963) “Jiabo”’ 1978 Shown by Stover & Evitt, 1978, to be a 77 syn. of Oligosphaeridium irregulare (Po- cock, 1962) Davey & Williams, 1969. [Considered by Duxbury, 1977, to be a jr syn. of Oligosphaeridum asterigerum (Gocht, 1959) Davey & Williams, 1969. This judgment is not accepted by us. } (Considered by Wiggins, 1975, to beajr syn. of Pareodinia Deflandre, 1947. This judge- ment was rejected by Sarjeant, 1976. |] [Considered by Wiggins, 1975, to bea jr syn. of Pareodinia ceratophora Deflandre, 1947. This judgement was rejected by Sarjeant, 1976. | DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 43 PERISSEIASPHAERIDIUM Davey & Williams, 1966b eisenacki (Eisenack, 1958) Davey & Williams, 1969 pannosum Davey & Williams, 1966b PLURIARVALIUM Sarjeant, 1962a osmingtonense Sarjeant, 1962b POLYSPHAERIDIUM Davey & Williams, 1966b; emend. Bujak ef al., 1980 Pasperum (Maier, 1959) Davey & Williams, 1969* Pbelgicum Sarjeant, 1969 ?deflandrer (Valensi, 1947) Davey & Williams, 1969 ?fabium (Tasch, 1964) Davey & Williams, 1969 Pfluctuans (Pastiels, 1948) Davey & Williams 1966b ?follium (Tasch, 1964) Davey & Williams, 1969 ?fucosum (Valensi, 1955) Davey & Williams, 1969 laminaspinosum Davey & Williams, 1966b Pmajor (Lejeune-Carpentier, 1940) Davey & Williams, 1969 Pmarsupium (Tasch, 1964) Davey & Williams, 1969 pastielst Davey & Williams, 1966b ?paulinae (Valensi, 1953) Davey & Williams, 1969 Pperovatum (Tasch, 1964) Davey & Williams, 1969 ?bolypes (Cookson & Eisenack, 1962) Davey & Williams, 1966b pumilum Davey & Williams, 1966b ?rhabdophorum (Valensi, 1955) Davey & Williams, 1969 simplex (White, 1842) Davey & Williams, 1969 (Considered by Davey & Verdier, 1974 and Duxbury, 1980, to be a 77 syn. of Hystrich- osphaerina schindewolfii Alberti, 1961. ] [Considered by Wiggins, 1975 and Dorhofer & Davies, 1980 to be a 77 syn. of Pareodinia Deflandre, 1947. We concur with Stover & Evitt, 1978, who neither accept this judgement nor the resultant transfer of the type species. ] Dapsilidinium asperum (Maier, 1959) Bujak et al., 1980* Dapsitlidinium? belgicum (Sarjeant, 1969) Len- tin & Williams, 1981 Dapsilidinium? deflandre: (Valensi, 1947) Len- tin & Williams, 1981 Shown by Stover & Evitt, 1978, to be a syn- onym of Ktokanstwm unituberculatum (Tasch, 1964) Stover & Evitt, 1978 The taxonomic transfer of this species was not validly made; therefore it must remain in Hystrnichosphaeridium. Shown by Stover & Evitt, 1978, to be a syn- onym of Kziokansium unituberculatum (Tasch, 1964) Stover & Evitt, 1978. Dapsilidinium? fucosum (Valensi, 1955) Lentin & Williams, 1981 Dapsilidinium laminaspinosum (Davey & Williams, 1966b) Lentin & Williams, 1981 Dapsilidinium? major (Lejeune-Carpentier, 1940) Lentin & Williams, 1981 Shown by Stover & Evitt, 1978, to be a syn- onym of Kiokansium unituberculatum (Tasch, 1964) Stover & Evitt, 1978. Dapsilidinium pastielsur (Davey & Williams, 1966b) Bujak e¢ al., 1980 Dapsilidinium? paulinae (Valensi, 1953) Len- tin & Williams, 1981 Shown by Stover & Evitt, 1978, to be a syn- onym of Kiokansitum unituberculatum (Tasch, 1964) Stover & Evitt, 1978. Bacchidinium polypes (Cookson & Eisenack, 1962) Davey, 1979 Dapsilidinium? pumilum (Davey & Williams, 1966b) Lentin & Williams, 1981 Dapsilidinium? rhabdophorum (Valensi, 1955) Lentin & Williams, 1981 Dapsilidinium simplex (White, 1842) Bujak et al., 1980 44 SECOND APPENDIX TO POLYSPHAERIDIUM continued subtile Davey & Williams, 1966b; emend. Bujak et al., 1980 ?tribrachiosum (Tasch, 1964) Davey & Williams, 1969 zoharyi (Rossignol, 1962) Bujak et al., 1980 PROLIXOSPHAERIDIUM Davey et al., 1966 deirense Davey et al., 1966 granulosum (Deflandre, 1937) Davey et al., 1966 mixtispinosum (Klement, 1960) Davey et al., 1969** parvispinum (Deflandre, 1937) Davey et al., 1969 Pxanthiopyxides (O. Wetzel, 1933) Davey et al., 1969 PSALIGONY AULAX Sarjeant, 1966b apatela (Cookson & Eisenack, 1960b) Sar- jeant, 1969 deflandrei Sarjeant, 1966b stmplicia (Cookson & Eisenack, 1961) Sar- jeant, 1969 RAPHIDODINIUM Deflandre, 1936 fucatum Deflandre, 1936 RHAETOGONYAULAX Sarjeant, 1966b; emend. Harland ef al., 1975; emend. Fisher & van Helden, 1979 chalonert (Sargeant, 1963) Sarjeant 1969 rhaetica (Sarjeant, 1963) Loeblich & Loe- blich, 1968; emend. Harland ef al., 1975 RHYNCHODINIOPSIS Deflandre, 1935; emend. Below, 1981; emend. Sarjeant, 1982a aptiana Deflandre, 1935; emend. Sarjeant, 1982a SIRMIODINIUM Alberti, 1961; emend. Warren, 1973 grosst Alberti, 1961; emend. Warren, 1973 SURCULOSPHAERIDIUM Davey et al., 1966 cribrotubiferum (Sarjeant, 1960) Davey et al., 1966 Shown by Stover & Evitt, 1978, to be a syn- onym of Kiokansium unituberculatum (Tasch, 1964) Stover & Evitt, 1978. [Considered by Davey & Verdier, 1974, to be a jr syn. of Prolixosphaeridium parvi- spinum (Deflandre, 1937) Davey et al., 1969. ] Tanyosphaeridium xanthiopyxides (O. Wetzel, 1933) Stover & Evitt, 1978 Tubotuberella apatela (Cookson & Eisenack, 1960b) Ioannides et al., 1976; emend. Sar- jeant, 1982a. [The previous placement of this species as the type for the new genus Glabridinium by Brideaux, 1977, was re- jected by Sarjeant, 1982a. | Impagidinium? simplicium (Cookson & Eisen- ack, 1961) Stover & Evitt, 1978 Rhaetogonyaulax rhaetica subsp. chalonert (Sar- jeant, 1963) Lentin & Williams, 1977 DAVEY, DOWNIE, SARJEANT & WILLIAMS (1966) 45 SURCULOSPHAERIDIUM continued longifurcatum (Firtion, 1952) Davey et al., 1966 vestitum (Deflandre, 1938) Davey et al., 1966 SYSTEMATOPHORA Klement, 1960 areolata Klement, 1960 placacantha (Deflandre & Cookson, 1955) Davey et al., 1966; emend. May, 1980 schindewolfi (Alberti, 1961) Downie & Sar- jeant, 1965 TANYOSPHAERIDIUM Davey & Williams, 1966b elupticum (Cookson, 1965) Davey & Williams, 1969 tsocalamum (Deflandre & Cookson, 1955) Davey & Williams, 1969 regulare Davey & Williams, 1966b variecalamum Davey & Williams, 1966b THALASSIPHORA Eisenack & Gocht, 1960; emend. Gocht, 1968 delicata Williams & Downie, 1966c; emend. Eaton, 1976 pelagica (Eisenack, 1954) Eisenach & Gocht, 1960 WANAEA Cookson & Eisenack, 1958; emend. Fensome, 1981 spectabilis (Deflandre & Cookson, 1955) Cookson & Eisenack, 1958 WETZELIELLA Eisenack, 1938; emend. Williams & Downie, 1966b; emend. Lentin & Williams, 1976 articulata Eisenack, 1938 articulata var. conopia Williams & Downie, 1966b clathrata Eisenack, 1938 coleothrypta Williams & Downie, 1966b condylos Williams & Downie, 1966b homomorpha Deflandre & Cookson, 1955 homomorpha var. quinquelata Williams & Downie, 1966b ovalis Eisenack, 1954 Hystrichosphaerina schindewolfi Alberti, 1961. [The genus Hystrichosphaerina, rejected as ajr syn. of Systematophora by Neale & Sarjeant (1962), was reinstated under an emended definition by Stover & Evitt, 1978. ] Distatodinium ellipticum (Cookson, 1965) Eaton, 1976 [Since the other constituent subgenera have either been raised to generic status or abandoned, the type subgenus Wetzeliella has now fallen into disuse. ] Kisselovia tenuivirgula subsp. conopia (Williams & Downie, 1966b) Costa & Downie, 1979 Kisselovia? clathrata (Eisenack, 1938) Lentin & Williams, 1976 Kisselovia coleothrypta (Williams & Downie, 1966b) Lentin & Williams, 1976 Dracodinium condylos (Williams & Downie, 1966b) Costa & Downie, 1979 Apectodinium homomorphum (Deflandre & Cookson, 1955) Lentin & Williams, 1977; emend. Harland, 1979 Apectodinium quinquelatum (Williams & Downie, 1966b) Costa & Downie, 1979 46 SECOND APPENDIX TO WETZELIELLA continued reticulata Williams & Downie, 1966b similis Eisenack, 1954 solida (Gocht, 1955) Williams & Downie, 1966b symmetrica Weiler, 1956 symmetrica var. lobisca Williams & Downie, 1966b tenuivirgula Williams & Downie, 1966b tenuivirgula var. crassiramosa Williams & Downie, 1966b varielongituda Williams & Downie, 1966b WETZELIELLA Gocht, 1955 draco Gocht, 1955 (RHOMBODINIUM ) Kisselovia reticulata (Williams & Downie, 1966b) Lentin & Williams, 1976 Dracodinium simile (Eisenack, 1954) Costa & Downie, 1979 Dracodinium solidum Gocht, 1955 Wetzeliella meckelfeldensis subsp. Jobisca (Williams & Downie, 1966b) Costa & Downie, 1979 Kisselovia tenuivirgula (Williams & Downie, 1966b) Lentin & Williams, 1976 Kisselovia crassiramosa (Williams & Downie, 1966b) Damassa, 1979 Dracodinium varielongitudum (Williams & Downie, 1966b) Costa & Downie, 1979 Raised to generic status by Gocht, 1955; emend. Bujak, 1979. Rhombodinium draco Gocht, 1955 glabra Cookson, 1956 Rhombodinium glabra (Cookson, 1956) Voz- zhennikova, 1967 XIPHORIDIUM Sarjeant, 1966b alatum (Cookson & Eisenack, 1962) Sar- jeant, 1966b REFERENCES ALBERTI, G. 1959. Zur Kenntnis der Gattung Deflandrea Eisenack (Dinoflag.) in der Kreide und im Alttertiar Nord- und Mitteldeutschlands. Mztt. geol. StInst. Hamb. 28: 93-105, pls 8, 9. — 1961. Zur Kenntnis mesozoischer und alttertiarer Dinoflagellaten und Hystrichosphaerideen von Nord- und Mitteldeutschland sowie einigen anderen Europaischen Gebieten. Palaeontogra- phica, Stuttgart, (A) 116: 1-48, pls 1-12. BaksI, S. K. 1962. 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Dept. of Geological Sciences UNIVERSITY OF SASKATCHEWAN SASKATOON CANADA .% 7 r' ~< 7 a Ba i > we eA PA * ¥ _ J _ j= + 6 ir te =n 4 al or } i 7 ol a ' 4 ‘ =~ a.) ° in a Gael | Gime P= sl we ) min et pe-glls teh abe OSPR .-¥ t. - ”. ad eur d uy HET pat ¢ ae Tay! t y f Soe! ¢ ayy ' +I te i 4% ‘ee ‘. - / \ j = Pa oa ‘ . a ub | ‘ \ 4 ‘ ~ , \ "sy Ul oat in| \ fa Fiery Jie SS —