fate mT VS atte oe a ne Taha te eagae ye Lee! eR, vide Stet et RYE aan rainy Ort LOEW bere Min tee 7 yey Se Oh ete Ve ApA te hae ESM cyst he fetes RUD og an N BAGY A he Srben la % , seats eed wee : Th tg tee ih Awta ty We eae Sedge ATTN TOR nhs Sse Le phen word ge wee typ eek ny Sed Coe Ves ia easy acai don vir ye eis ayes ’ Yop owe HARVARD UNIVERSITY e Library of the Museum of Comparative Zoology - ; ml a J | 5 i i 3 ary 1% baa f Hi Bulletin of the Museum of Comparative Zodlogy ‘AT HARVARD COLLEGE. Vou. LVII. No. 1. THE HENICOPIDAE OF AMERICA NORTH OF MEXICO. By Rates V. CHAMBERLIN. Wits Five Puares. CAMBRIDGE, MASS., U. S. A.: PRINTED FOR THE MUSEUM. DrEcEMBER, 1912. Pa atl: ; ' PA ey be hoitieonnen 3 iff a: a ae Reis, ie ae ns eh a CS | ie ee ad i" a ‘posh A ce nh ink Ware No. 1.— The Henicopidae of America north of Mexico. By Raupa V. CHAMBERLIN. THE genus Henicops was proposed by Newport in 1844 for two species, H. maculatus, from Van Diemen Land, and H. emarginatus, from New Zealand. Twenty-four years later Meinert described from Europe a species obviously related to these, placing it under a new genus as Lamyctes fulvicornis; but by common consent subsequent workers have, until recently, dropped Lamyctes as a synonym of Henicops. However, in 1901, Pocock restudied Newport’s species and pointed out that maculatus, which he specified as the type of Henicops, differed from the European species in having the tarsi of the fourteenth and fifteenth pairs of legs, six segmented and those of the anterior pairs trisegmented, the last three pairs of legs in fulvi- cornis having the tarsi only bisegmented and the anterior legs having the tarsi wholly undivided. In accordance with these important differences Pocock again recognized Lamyctes as a distinct genus, at the same time describing two new genera evidently closely allied, Haasiella and Paralamyctes.! Other genera having more or less close affinities with Henicops and Lamyctes have also been described by Silvestri ? and by the present writer. Silvestri and Pocock place these genera, which, it may be noted, occur in the main in the Southern Hemisphere, in a family Henicopidae in contrast particularly with the Lithobiidae, including Lithobius and allies which abound for the most part in the Northern Hemisphere; but Verhoeff in his recent revision of the Lithobiomorpha in Die klassen und ordnungen des thierreichs (1907, 5, p. 245f), does not recognize the family, denying the sufficiency of the basis for it. Careful study of the genera of the Lithobiomorpha, however, makes it very obvious that these (omitting those of the Cermatobiidae) fall into two compact and clearly sepa- rated groups and that, as one of these, the Henicopidae must be given recognition as a distinct and well-defined family. 1R. I. Pocock, Some new genera and species of lithobiomorphous chilopods. Ann. mag. nat. hist., 1901, ser. 7, 7, p. 448-451. 2 F. Silvestri, Descrizioni di aleuni generi especie di Henicopidae. Boll. Lab. zool R. scuola sup. agric. Portici, 1910, 4, p. 38-50. 4 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. The following points in which the Lithobiidae differ from the Heni- copidae may be mentioned as the most important thus far noted :— 1. The labrum is tridentate instead of unidentate. 2. The inner branch of the first maxillae is armed with ramose or plumose hairs. 3. The outer branch of the first maxillae is only biarticulate. 4. When present, the eyes consist of from several to many ocelli. 5. Sternite of prehensorial segment absent, never constituting a distinct transverse sclerite or collar. 6. On none of the legs is a true tibial process or spur ever present. 7. Articular spines are present on at least part of the legs cephalad of the fourteenth pair. 8. Anal pores never present in adults. 9. Gonopods of male composed of one or, at most, of two articles and never terminating in a pointed process or rudimentary claw. In regard to point 7 it may be mentioned that in Watobius and allies (subfamily Watobiinae mihi), which differ from the typical Lithobiidae in other points also, spines are present only on the tibiae in the place occupied by the tibial spur in the Henicopidae; while in the Henicopidae spines may very rarely occur on trochanter or on tro- chanter and prefemur of the fourteenth and fifteenth pairs of legs. The following definition of the Henicopidae is based primarily upon a study of the North American representatives. HENICOPIDAE (char. emend.). Labrum mesally deeply incised; unidentate; margins of lateral divisions bearing either plumose or else simple or nearly simple hairs. Inner branch of first maxillae armed with simple hairs exclusively, or in part with hairs simply and shortly laciniate at tips. Outer branch triarticulate; clothed distally with plumose hairs. Palpus of second maxillae triarticulate; claw four or five partite; distal article bearing on ventral surface toward distal end few to many plumose (ramose) hairs. Eyes either entirely absent or consisting of a single large ocellus on each side. Organ of Témésvary large; situated on ventral border of head, typically caudad of position of ocellus. Sternite of prehensorial segment well developed, extending trans- CHAMBERLIN: HENICOPIDAE OF AMERICA NORTH OF MEXICO. 5 versely as a narrow sclerite or “collar”? caudad of the prosternum proper. . Tibia of all legs, or of all excepting the last one to three pairs, bear- ing at the distal end on the cephalic or cephalodorsal side an acute or spinous tipped process or spur. ‘True articular spines either wholly absent (as in all N. Amer. species) or present only on last two pairs of legs. Spiracles present on 3rd, 5th, 8th, 10th, 12th, and 14th segments, as in the Lithobiidae, or, in addition also on the Ist; or rarely present only on the 3rd and 10th segments (Catanopsobius). Coxal pores present on the last two to five pairs of coxae. Anal pores always present in adults and consisting of a single well- developed pair opening into a common median passage. Gonopods of female long, triarticulate, and ending in a stout, strongly chitinized claw which is always entire; basal spines, so far as known, 2 + 2 or 2+3. 2nd article glabrous dorsally. Gonopods of male likewise long and triarticulate; ending in a straight, weakly chitinized process usually terminating in a bristle. Three genera of this family are at present known to occur in America north of Mexico. They may be separated by means of the following key. aa. SKE ee ana PLATE 3. BUETHOBIUS OABITUS Chamberlin. Prosternum and prehensors. First maxillae. Mandible. Branched aciculus from tip of outer branch of first maxilla. Ramose or plumose hair from outer branch of first maxilla. Labrum. Head, dorsal aspect. Anal leg. Fifth leg, cephalic aspect. Sternite of genital segment and gonopods of female. Bull. Mus. Comp. Zodl. Henicopidae Plate 3 OL OLE LEP AE s~ Jy cine a “wen : rAVS- a7, ee ee? Le haa ath pit e, 5 ity n ) iy? s ” oa an ‘ ; "i . rs ‘ OF Ae vn oh Nv ates ue) : as ay = i? i! hivyaas*) » ‘ ne r ‘sures | ; , ah at | iy t i t 5 » - uy hes [2° . one + ‘ 9 . an) ae | Pa 7 4 4 pvt fal heii 1 i é / in f j . \ ‘ Chamberlin: — Henicopidae of America north of Mexico. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. — iis J SAN 2 PLATE 4. BUETHOBIUS OABITUS Chamberlin. Head, ventral aspect, showing organ of Témdésvary. Distal joints of palpus of second maxilla. BUETHOBIUS CONIUGANS Chamberlin. Ventral view of posterior region of body of male, showing gonopods, etc. ZYGETHOBIUS SOKARIENUS Chamberlin. Claw of female gonopods, dorsal aspect. Head, dorsal aspect. Prosternum and prehensors. Anal leg. Leg of fifth pair, cephalic aspect. ZYGETHOBIUS PONTIS Chamberlin. Anal leg. Henicopidae Plate 4 Bull. Mus. Comp. Zool. PLATE 5. Chamberlin: — Henicopidae of America north of Mexico. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. PLATE 5. ZXGETHOBIUS SOKARIENUS Chamberlin. Distal article of palpus of second maxilla. Labrum. First maxilla. Laciniate bristles from inner branch of first maxilla more highly magnified. Mandible. Sternite of genital segment and gonopods of female. Antenna. Ventral view of caudal region of body of male showing gonopods, etc. ZXYGETHOBIUS PONTIS Chamberlin. Antenna. Bull. Mus. Comp. Zodl. Henicopidae Plate 5 Bulletin of the Museum of Comparative Zodlogy AT HARVARD COLLEGE. Vou VIE. No: 2. THE LITHOBIID GENERA NAMPABIUS, GARIBIUS, TIDA- BIUS, AND SIGIBIUS. By Raupo V. CHAMBERLIN. Wits Five Puates. CAMBRIDGE, MASS., U.S. A., PRINTED FOR THE MUSEUM. Novemser, 1913. No. 2.— The Lithobiid Genera Nampabius, Garibius, Tidabius, and Sigibius. By Raupa V. CHAMBERLIN. - Tue four new genera dealt with in the present paper include the smallest of the known North American species of Lithobiidae. None of them exceeds ten mm. in length, most of them averaging materially less than this. They occur principally among the layers of damp fallen leaves, under the bark of logs, and in the looser loam of open woods. All seem to have the death-feigning instinct more or less pronounced, when suddenly jarred or otherwise frightened often flex- ing the body laterally into a circle and lying rigid for some time, when they may be handled with forceps or otherwise without their relaxing. Because of their small size and the effectiveness with which they conceal themselves, specimens of these species are not often secured by the general collector; but one looking for them especially and understanding their habits may nearly always secure one or more species in appropriate habitats throughout the United States excepting the Southwest where they are replaced by Oabius (pylorus Chamb., tiganus Chamb., etc.), Hesperobius (medius Chamb., obesus Stuxb., and allies), and related genera to be considered in a suc- ceeding Bulletin. They may sometimes be secured in considerable numbers by sifting leaves. In addition to the similarity in many features of habit, the species of these four genera likewise have much in common in general appearance and structure. The genera are, however, readily separated upon critical examination. The discrimination of the species has presented more difficulty. Heretofore the species have remained practically unknown, not only because of the paucity of material in collections, but especially because the characters studied and given in descriptions are, in the main, not truly specific, being either generic in extent or common to a number of species. More detailed and careful study and comparison of the material in extensive collections made by the author personally in various parts of the country has revealed the considerable number of species here described. The four genera may readily be separated by means of the following key. 40 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Key to the Genera. a. Articles of antennae from 25 to 35, the number being mostly near 32. Posterior legs of co without special processes or lobes. (Posterior coxae wholly unarmed). b. Dorsal spines of anal and penult legs 0, 0, 1, 0, 0, ventral Ope de WO: Sigibius, gen. nov. / | bb. Dorsal spines of anal legs 0, 0, 2, 0, 0, ventral 0, 1, 3, 1, 0, or’ 0, 1,3, 2,0; dorsal spines of penult legs 0, 0, 3, 1, 0, or 0, 0, 3, 1, tevemtraliO. Leics. Or On deel. tonal, Tidabius, gen. nov. aa. Articles of antennae normally 20. Either anal or penult legs of o&' with a special lobe. (Coxae armed or unarmed). b. Posterior coxae wholly unarmed; median incision of proster- num semicircular; penult legs of o always bearing at distal end of tibia on dorsal side a small but conspicuous lobe, the anal legs without lobes or crests; posterior angles of 11th and 13th or at least of 13th more or less produced. Nampabius, gen. nov. bb. Last 3 or 4 pairs of coxae dorsally armed; median incision of prosternum V-shaped; penult legs of co’ without any such lobe, but anal legs bearing at distal end of tibia a small crest; posterior angles of none of the dorsal plates at all produced. Garibius, gen. nov. © NAMPABIUS, gen. nov. Body conspicuously attenuated cephalad, with the first dorsal plate much narrower than the head and also narrower than the third plate. Mostly robust, varying from 6 to 8.5 times as long as the width of the 10th plate. Antennae short; composed of 20 articles. Ocelli in 2, or, less commonly, in 3, longitudinal series; the,single ocellus mostly a little smaller than the most caudal one of the dorsal series. { Prosternal teeth 2+2. Median incision semicircular in outline, never acute at bottom. (Plate 1, fig. 2). Eleventh and thirteenth dorsal plates with the caudal angles more or less produced or sometimes that of thirteenth alone distinctly extended; angles of other plates straight or excised. Claw of the female gonopods tripartite, with the lobes subequal or CHAMBERLIN: LITHOBIID GENERA. 41 the median longest, all usually more or less rounded distally. Basal spines 2+2, long and usually slender, the sides, as seen in ventral view, converging to near the apex or beginning of the acuminate distal portion which is very short. (Plate 2, fig. 6). Coxal pores very small; circular, 2, 2, 2, 2 to 2, 3, 3, 3 in number. Ventral spines of the anal legs 0, 1, 1, 0, 0 or 0, 1, 1, 1, 0; dorsal 0, 0, 0, 0, 0 or 0,0, 2, 0,0; claws 2 or, rarely, but 1. Ventral spines of penult legs 0, 1, 1, 1, 0 or 0, 1, 2, 1, 0; dorsal spines 0, 0, 0, 0, 0 or 0, 0, 2, 0, 0; claws 2 or, rarely, but 1. Ventral spines of thirteenth lezsi0;0;;2, 1, Isto 0/0;0) 1, i; dorsaltirom 0; 0;'0,:0; 0'to 00, 2, 1, 1. Ventral spines of twelfth legs from 0, 0, 0, 0, 1 to 0, 0, 0, 2,1; dorsal from 0, 0, 0, 0, 0 to 0, 0, 2, 1, 2. Ventral spines of eleventh legs OO 02010) co .0,.02)0)) 2) i Sdorsall from 0309 0)'0;)t\to OF0, 0, 2, 2. First two to first four pairs of legs with no ventral spines on any joint; and the third joint of the first twelve or thirteen pairs of legs ven- trally unarmed. The posterior coxae are wholly unarmed. Anal legs in the male short; moderately and uniformly inflated; with- out special lobes or furrows. Penult legs with the fifth article produced at its distal end on the dorsal or dorsomesal side into a small sub- eylindric or distally expanded lobe bearing a number of sensory hairs. Typr.— N. virginiensis, sp. nov. DistRIBUTION.— Eastern and Southeastern States, in the latter section occurring chiefly in or near the mountainous parts. The species of this genus are the smallest members of the family, all known forms being under 7.5 mm. in length. In proportions they are mostly relatively robust and conspicuously narrowed cephalad. They vary considerably in color, some being testaceous, some ferrugi- nous, and others chestnut. While the head may be paler than the dorsum, it is usually darker, frequently being a shining reddish black. As in Tidabius, the posterior legs, especially distally, are nearly always more brightly pigmented, being commonly bright yellow or orange. These forms occur commonly in and beneath layers of fallen leaves and loose piles of sticks. The following key will aid in the identification of the known species. Key to species of Nampabius. a. Neither the anal nor the penult legs with any dorsal spines. Ventral spines of anal legs 0, 1, 1, 0, 0. N. fungiferopes (Chamberlin). aa. Anal and penult legs with the third joint armed with 2 dorsal spines. 42 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. b. Dorsal spines of twelfth legs 0, 0, 2, 1, 2; (ventral spines of ninth to twelfth legs 0, 0, 0, 2, 1). N. virginiensis, sp. nov. bb. Dorsal spines of twelfth legs not 0, 0, 2, 1, 2. ec. Third joint of twelfth legs unarmed dorsally (spines 0, 0, 0, 1, 2 or fewer). d. Dorsal spines of twelfth legs 0, 0, 0, 0 (1), 1; of the eleventh 0, 0, 0, 1, 1. N. parienus, sp. nov. dd. Dorsal spines of twelfth legs 0, 0, 0, 1, 2; of the eleventh OOO 2 ori0s' 0; O22. e. Dorsal spines of tenth and eleventh legs 0, 0, 0, 1, 2; ventral spines of thirteenth legs 0, 0, 1, 1, 1. N. embius, sp. nov. ee. Dorsal spines of eleventh legs 0, 0, 0, 2, 2; ventral spines of thirteenth legs 0, 0, 2, 1, 1. f. Ventral spines of twelfth legs 0, 0, 1, 2,1; of eighth to eleventh 0, 0, 0, 2, 1; ocelli cir. 6 in two series. N. inimicus, sp. nov. ff. Ventral spines of twelfth legs 0, 0, 0, 1, 1; of eighth to eleventh also 0, 0, 0, 1, 1; ocelli 10 or more in three series. N. georgianus, sp. nov. ce. Third joint of twelfth legs armed with one or two dorsal spines. d. Dorsal spines of twelfth legs 0, 0, 1, 1, 2 or 0, 0, 2, 1, 1; of tenth and eleventh 0, 0, 0, 2, 2. N. carolinensis, sp. nov. dd. Dorsal spines of twelfth legs 0, 0, 2, 1, 0 or 0, 0, 2, 1, 1; of tenth and eleventh 0, 0, 0, 0, 1 or 0, 0, 0, 1, 1. e. Dorsal spines of twelfth and thirteenth legs 0, 0, 2, 1, 0; of eleventh 0, 0, 0, 1, 1. N. tennesseensis, sp. nov. ee. Dorsal spines of twelfth and thirteenth legs 0, 0, 2, 1, 1; of eleventh 0, 0, 0, 0, 1. f. Near 8.5 times longer than width of tenth plate. N. longiceps, sp. nov. ff. Not more than 7 times longer than width of tenth plate. N. lulae, sp. nov. Notr.— The key does not include NV. lundiz (Meinert) the insuffi- cient description of which must leave its identity in doubt until the types can be restudied. See further under the account of this species. CHAMBERLIN: LITHOBIID GENERA. 43 NAMPABIUS FUNGIFEROPES (Chamberlin). Lithobius fungiferopes Chamberlin, Proc. Acad. nat. sci. Phil., 1904, 56, p. 652. DraGnosts.— Dorsum testaceous to dark, more or less reddish brown, in life more or less suffused with purplish. Head darker, almost black, in front of suture paler and with a tendency toward ferruginous. Body about 62 times longer than width of tenth plate. Head considerably wider than long. Ocelli mostly 4 to 6 in two series. Prosternum about 1.54—1.6 times wider than long; distance between chitinous spots about 2.3-2.4 times greater than width at median incision and 3.8 or 3.9 times the dental line. First dorsal plate about 1.9 times wider than long. Coxal pores 2, 2, 2, 2. Claw of female gonopods short and wide with the median lobe considerably largest, the lateral equal. Spines of moderate width, the sides converging distad to the short, abruptly subacute apex. Twelfth to anal legs inclusive with no dorsal spines. Ventral spines of anal 0, 1, 1, 0,0; of penult, 0, 1,1, 1,0. Ventral spines of ninth to thirteenth legs (also of fifth to ninth) inclusive 0, 0, 0, 0, 1. Dorsal spines of tenth and eleventh legs (also of second to tenth) 0, 0, 0, 0, 1. Process of the penult legs in the male conspicuously expanding distad; bearing the usual hairs. Tarsal joints abruptly and consid- erably more slender than the tibia. (Plate 1, fig. 6-7). Description.— Above testaceous to dark, more or less reddish, brown, in life especially commonly conspicuously suffused with viola- ceous or purplish. Head deeper in color, almost black, but paler in front of suture where it may show a tendency toward ferruginous. Antennae rufous distally, proximally brown or purplish brown. Prosternum dark brown with the prehensorial feet lighter. Venter light brown or testaceous, in life suffused with purplish. Legs pale brown, in life of a distinctly purplish or violaceous tinge proximally, being yellow distally; posterior pairs more strongly pigmented as usual, bright yellow except proximally. Body strongly attenuated cephalad of tenth plate as usual. Pro- portionately robust, being mostly 6.75+ times longer than width of tenth plate. The widths of head and first, third, eighth, tenth, and twelfth plates to each other as 37: 31: 33: 42: 42: 39. 44 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Head wider than long (mostly from 37:34 to 37:36); widely rounded anteriorly. Hairs long and sparse. Antennae short; composed of the normal twenty articles, all of which are short. Hairs of moderate length, not very dense, more sparse proximally. Eyes composed mostly of from 4 to 6 ocelli which are arranged in two series: — thus, 1+2,1 to1+3,2. The single ocellus smaller than most of the others. All ocelli well pigmented. Prosternum 1.54-1.6 times wider than long. Distance between chitinous spots near 2.38 times the width at level of bottom of mesal incision; about 3.87 times the dental line, the anterior portion being more strongly narrowed than in the related species. Median incision shallow, not large, of the usual semicircular form. Teeth small and acute, equal or nearly so, the interval between each pair narrow and well rounded at bottom. Sides slanting directly from close to base of each ectal tooth, nearly straight, being slightly concave near middle of length. First dorsal plate widest at anterior end, the sides at first but little and then more strongly converging caudad and rounding in about the caudal corners; nearly 1.9 times wider than long. Posterior corners of the eleventh and more distinctly of the thirteenth plates weakly produced, those of the ninth straight. Coxal pores small and circular; mostly 2, 2, 2, 2. : Fp OnOn OA0n 0. . . Spines of first legs 5-9-5-9'9; of second to fourth inclusive 0. 0, 0, 0, 1. . a SUMO OAR ale Sa of fifth to eleventh inclusive cHoaawaree of twelfth and . 0, 0, 0, 0, 0. 0, 0, 0, 0,0 : thirteenth, Saaremaa of the penult coat arene the claws 2; of the anal 0: 0, 0.0 claws 2, the accessory claw easily lost from preserved 0; 1,-1, On0 specimens in both anal and penult pairs. In the male the anal and penult legs are short and but moderately inflated; longitudinally suleate along ectal or anterior side, more distinctly so on fourth and fifth articles. In the penult legs the tarsal joints are abruptly and considerably more slender than the tibia; tibia with the distal mesodorsal corner obliquely complanate and there bearing the usual small process, this being in this species conspicu- ously widened at the distal end (see Plate 1, fig. 6 and 7) and bearing on its distal surface the usual sensory hairs. Claw of female gonopods short and relatively wide, scarcely curved; tripartite, the middle lobe considerably larger than the two lateral which are about equal, all lobes distally narrowly rounded. Spines CHAMBERLIN: LITHOBIID GENERA. 45 moderately wide, with the sides converging from base distad, toward ends more abruptly bending in to the acute or subacute apex. Mesal side of first article of gonopods nearly straight and parallel with its fellow. Length from 5 to nearly 7 mm. Type Locatiry.— New York (Ithaca!). Locatitres. —New York (Ithaca!); Vermont (Jamaica! Grout’s Mill!). NAMPABIUS VIRGINIENSIS, Sp. NOV. D1acenosis.— Dorsum testaceous to dark reddish or ferruginous brown. Head more or less ferruginous, darker caudad of suture. Body robust, being mostly but 6.25 or 6.3 times longer than width of the tenth plate. Head equal in length and breadth; cordate. Ocelli mostly 6 or 7 in two series. Prosternum about 1.63 times wider than long, the anterior portion being short. Distance between chitinous spots only 1.7 times greater than width at median incision and but 2.9 times greater than the dental line. First dorsal plate nearly 1.9 times wider than long. Coxal pores 2, 2, 2, 2-2, 3,'3, 3. Claw of female gonopods with the outer latest tooth somewhat smaller than the inner. Spines long, their sides moderately converg- ing to near apex where curving in more abruptly to the acute apex. Dorsal spines of anal and of penult legs 0, 0, 2, 0, 0; ventral spines of these legs 0, 1, 1, 1,0. Dorsal spines of thirteenth legs 0, 0, 2, 1, 0; ventral 0,0, 2,1,1. Dorsal spines of twelfth legs 0, 0, 2, 1,2. Dorsal spines of tenth and eleventh legs 0, 0, 0, 2,2. Ventral spines of ninth to twelfth legs inclusive 0, 0, 0, 2, 1. Process of penult legs of male subcylindric; expanded somewhat distally, flat-topped and on the distal surface bearing a subdense patch of moderately long sensory hairs. (Plate 1, fig. 4-5). Description.— Dorsally testaceous to dark reddish or ferruginous brown, more reddish anteriorly and posteriorly. Head more or less ferruginous, darker, dusky, caudad of the suture. Antennae like the head; pale at tips. Prosternum brown, with the prehensorial feet paler. Venter pale yellowish to light reddish brown, the caudal plates darker. Legs pale yellow to testaceous, with the posterior pairs, excepting at base, bright yellow or somewhat orange. 46 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body attenuated as usual. Typically near 6.25 or 6.3 times longer than width of the tenth plate. A female 5 mm. long has the widths of head and first, third, eighth, tenth, and twelfth plates to each other as 37:33:35: 42: 42: 42. Head nearly equal in length and breadth. Cordate; widest a little in front of marginal breaks from where it narrows continuously forwards. : Antennae very short; moderately attenuated. Composed of the usual twenty articles which are short and very short; decreasing distad from the second. Eyes composed mostly of 6 or 7 ocelli arranged in 2 series: — thus 1+3, 2 or 1+4, 2. Dark. The most caudal ocellus of upper row considerably larger than the single one, the first of lower row less markedly larger. Prosternum about 1.63 times wider than long. Distance between chitinous spots only 1.7 times greater than width at level of bottom of mesal incision which is nearly in line with caudal ends of lateral slopes; 2.9 times longer than the dental line which is also thus longer than usual. Median incision large and conspicuous, of the usual form. Teeth subequal, small and acute, the interval between each pair semicircular; line of apices recurved. Sides straight or a little in- curved, short. First dorsal plate widest at anterior end from where the sides con- verge caudad considerably; nearly 1.9 times wider than long. Thir- teenth dorsal plate with posterior corners moderately produced, those of the eleventh less so as usual; caudal margin of ninth plate straight. Coxalsporesismialll: 25212512) tol23n3; 3: Spines of the first legs 0-0-0. 0-0, 0,0,0,1. of the second 0, 0,0, 0, 1 Oooo of the third to seventh inclusive, ae of the eighth o o ocr of the ninth, 0 One ea of the tenth and eleventh, O-o-o-g-43 of the thirteenth 0-02-49; of the penult eee aE of the anal Oo . - i: . In the male the anal legs are short, moderately and uniformly inflated, and without special modifications. Penult legs similar to anal but the fifth article obliquely complanate at distal end on meso- dorsal side and there bearing the usual small lobe which is subcylindrie and somewhat expanded distad, especially on side toward distal end of leg; sensory hairs borne on distal surface of lobe rather long and subdense. (Plate 1, fig. 4-5). CHAMBERLIN: LITHOBIID GENERA. 47 Claw of female gonopods of moderate size; considerably curved; tripartite; the inner and median teeth not much different in size, the outer one smaller. Spines rather long, the sides as usual converging from base distad, toward end more strongly curving in to meet at the acute apex. Inner side of first article nearly straight, a little diverging from its fellow proximad. Length near 5 mm. Type Locatity.— Virginia (Natural Bridge! August, 1910). Locauities.— Virginia (Natural Bridge!); Tennessee (Unaka Springs!). NAMPABIUS PARIENUS, sp. Nov. Diacenosis.— Dorsal spines of anal, penult, and thirteenth legs 0,0,2,0,0. Ventral spines of anal leg 0,1, 1,1, 0, of penult 0, 1, 1, 1, 0- 0, 1, 2, 1, 0; of ninth to thirteenth, 0, 0, 0, 1, 1. Dorsal spines of tenth and eleventh pairs of legs (as also of the second or third to ninth) O05 0; 1. 31. Ocelli 5 or 6 in two series. Prosternum 1.57 or 1.6 times wider than long. Distance between chitinous spots about 3.6 times greater than the distal line; 2.25 times longer than the width at median incision. Coxal pores. 2;,2, 2, 2-2,.3,:3, 2. Process of tibia of penult legs in male rather long and slender, in proximal aspect with lateral lines parallel; in lateral aspect appearing thicker and projecting over base on side at distal end of article; sen- sory hairs long and slender. Body 7.7 or 7.75 times longer than width of tenth plate. Head broadly subcordate; distinctly wider than long. First dorsal plate (in type) between 1.75 and 1.8 times wider than long. Dorsum yellow with the caudal plates tinged with orange. Head orange, with the frontal region paler. DescripTion.— The body above is yellow with the caudal plates tinged with orange. Head orange, the frontal region paler, more yel- lowish. Antennae orange, yellow at tips. Prosternum orange brown. Venter pale, the caudal plates more strongly pigmented as usual. Legs pale excepting the posterior pairs which are bright orange. Body more slender than in most related species, in the type being 7.73 times longer than the width of the tenth dorsal plate. Width of head and first, third, eighth, tenth, and twelfth plates to each other about as 36: 32:35: 43: 44: 41. 48 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Head broadly subcordate, considerably bulging caudad of eyes. Clearly wider than long (12:11). Hairs short and sparse. Antennae short. Articles short and very short, decreasing distad of the second as usual. Hairs short and not dense. Eyes typically composed of five or six ocelli in two series: — thus, 1+2,2 or 1+8,2. Single ocellus smaller than the first one of upper row and the others of the usual relative sizes. Prosternum in type 1.57 times wider than long. Distance between chitinous spots 3.6 times the dental line; 2.25 times width at level of bottom of median incision. Median incision of the usual semicircular form, rather small, the distance across it between the two mesal teeth not greater than that between the teeth of each pair. Teeth moderate; line of apices distinctly and considerably recurved. Spine at base of ectal tooth; slender, somewhat curved mesad distally, not exceeding the tooth in length. First dorsal plate widest near anterior end, the sides from there at first slightly and then more strongly converging caudad; plate in type 1.77-++ times wider than long. Posterior corners of the eleventh and thirteenth dorsal plates a little produced, those of the ninth straight or also showing a very slight tendency toward production, rounded; other minor plates with corners oblique or rounded. Coxal pores very small; 2, 2, 2, 2-2, 3, 3, 2. Spines of first legs, Ooo: of second, 2: 50° é an 1: of third, O--0- 0-0} of the fourth to seventh, ear p of the ninth to eleventh, e = Ooort i, 75 of the twelfth, TCE 1 the dorsal spine of the atth nrucle being on the cephalic side; of the thirteenth, ear wee of the penult, an tee: Tan claws 2; of the anal, . . 1: os claws 2. In the male the anal and penult legs are short and uniformly and but little inflated; both are laterally longitudinally suleate. The process at the distal end of the tibia of the penult legs is rather long and slender; when viewed from proximal aspect of leg its sides appear nearly parallel; in lateral aspect the process is somewhat thicker and projects distad above its base; distal surface rising obliquely dorso- distad from its proximal end to its distal; sensory hairs long and slender. (Plate 8, fig. 1-2). Length (type) 6.5 mm.; width of tenth plate .84 mm.; length of antennae 2 mm. Locauiry.— North Carolina (Hot Springs’). CHAMBERLIN: LITHOBIID GENERA. 49 NAMPABIUS EMBIUS, sp. Nov. D1iacnosis.— Dorsal spines of the anal and penult legs 0, 0, 2, 0, 0. Dorsal spines of the thirteenth legs 0, 0, 2, 1, 0; ventral 0, 0, 1, 1, 1. Dorsal spines of the tenth to twelfth legs 0,0, 0, 1,2; ventral 0, 0, 0, 2,1. Dorsal spines of the sixth to ninth legs also 0, 0, 0, 1, 2. Ocelli 6 to 8 in two series. Prosternum about 1.56 times wider than long. Distance between chitinous spots 2.14 times width at median incision; 33% times the dental line. Spines of the gonopods rather long; sides parallel or but slightly converging from base to the very short acuminate tip, the sides of which are straight and meet at an almost obtuse angle. Body more slender than in most related species being (in type) about 84 times longer than the width of the tenth dorsal plate. Head clearly wider than long. First dorsal plate near 1.75 times wider than long. Dorsum brown of dilute chestnut caste. Head dark brownish red. DescripTion.— Brown of dilute chestnut caste. Head dark brownish red. Antennae brown, pale distally. Prosternum dark, more brownish than the head with the prehensorial feet paler. Venter pale brown, the caudal plates darker. Legs colored nearly like the corresponding plates of venter; posterior pairs light distad. Body comparatively slender being in the type 8% times longer than the width of the tenth plate. Widths of head and of the first, third, eighth, tenth, and twelfth plates to each other about as 39: 35: 38: 48: 49: 47.5. Head wider than long in ratio 13: 12; widest behind eyes from where the convergence of the sides to the caudal corners is marked. Antennae short as usual. Moderately attenuated. The twenty articles short and very short, decreasing regularly distad. Hairs of moderate length, short, not dense. (Plate 2, fig. 7). Eyes composed of 6 to 8 ocelli in two series: — thus, 1+3, 3 — 1+ 4,3—1+8, 2. Ocelli of moderate size, black. Prosternum 1.56 times wider than long. Distance between chitin- ous spots 2.14 times width at level of bottom of mesal incision; 34 times the dental line. Mesal incision wide and shallow, of the usual form at bottom. Teeth small, distally narrowly rounded; line of apices rather strongly recurved. Spine unusually long, bristle like. First dorsal plate widest a little caudad of anterior end from where 50 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. the sides converge considerably; 1.75 times wider than long. Poste- rior corners of the thirteenth plate a little produced; those of the eleventh nearly straight and those of ninth rounded cephalad. Coxal pores small and circular as usual, 2, 3, 3, 3. . . 04.0/10)0, 0 10,0" 0) Olt Spines of first pair of legs, 5~9'9-9-9 © o-o: b- 0,0) Of second and third, aS TEE of fourth and fifth, a e é 1; of sixth, EURO eet or 0, ONG ; 0, 0, 0, 1, 2. . 0, 0, 0, 1, 2. Tae of seventh, SS: of tenth to twelfth, aoa ate of . OOH, 0% 0, 0, 2, 0, 0 Da thirteenth, ae, of the penult, poe claws 2; of the anal 0;.0),2,,0350: Claw of female gonopods of moderate size; distinctly tripartite. Spines rather long, with sides parallel or but slightly converging from base to acuminate apical portion which, as usual, is relatively very short with sides strictly straight and running to an almost obtuse angle. Inner side of first article of gonopod considerably diverging proximad from its fellow. Sternite mesally straight, with no distinct process. Length (type) 7.5 mm., width of tenth plate .91 mm.; length of antenna, 2 mm. Loca.iry.— South Carolina (Taylors!). NAMPABIUS INIMICUS, Sp. nov. DraGnosis.— Dorsal spines of penult legs 0, 0, 2, 0, 0; ventral 0, 1, 2, 1, 0. Dorsal spines of thirteenth legs, 0, 0, 2, 1, 0; ventral, 0, 0, 2, 1, 1. Dorsal spines of twelfth legs, 0, 0, 0, 1, 2; ventral, 0, 0, 1, 2, 1. Dorsal spines of tenth and eleventh legs, 0, 0, 0, 2, 2. Ventral spines of tenth and eleventh (as also of eighth and ninth) legs, O70 70, 250% Ocelli (in type) 6 in two series. Prosternum about 1.57 times wider than long. Distance between chitinous spots 2.1 times the width at median incision; 3.1 times as great as the dental line. Coxal pores 2, 2, 2, 2—2;'3,'3} 2: Body about 7.8 times longer than width of tenth dorsal plate. Head subrotund, considerably wider than long. First dorsal plate 1.7 times wider than long. Process of penult legs in male small, cylindric proximally but ex- panded at distal end on side toward distal end of jot; sensory hairs short and spinescent. (Plate 8, fig. 5-6). CHAMBERLIN: LITHOBIID GENERA. Bik Dorsum light testaceous, the caudal plates of ferruginous tinge. Head dark orange. DescripTion.— Dorsum light testaceous, the caudal plates of ferruginous or orange tinge. Head dark orange. Antennae reddish brown, light at tips. Prosternum brown. Venter very pale, excepting the caudal plates which are orange. Legs pale like the venter, but the posterior pairs more brightly pigmented as usual. Body more slender than typical in the genus the length of type specimen being 7.77-+ times greater than the width of the tenth plate. Width of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other as 39: 34: 37: 46:47:43 (7). Head subrotund; considerably wider than long (39:35); the sides convexly bulging caudad of eyes; widely rounded cephalad. Antennae moderately short. Joints short, decreasing markedly in size from the second distad. Sides of articles straight, moderately flaring distad. Hairs moderately short, not very dense. Eyes in type composed of 6 ocelli in two series:— thus, 1+8, 2. Single ocellus of about same size as the first one of upper series or a very little smaller; others decreasing in size cephalad. Prosternum 1.57-+ times wider than long. Distance between chitinous spots 2.1 times width at level of bottom of median incision; 3.1 times the dental line. Lateral edges nearly straight or very slightly convex cephalad and slightly bending ectad at posterior end. Median incision wide, semicircular as usual; the interval between the two mesal teeth decidedly greater than that between the teeth of each lateral pair. Teeth acute; line of apices distinctly recurved. First dorsal plate 1.7 times ees than long; widest near middle, the sides converging cephalad and, more conspicuously, caudad. Posterior angles of eleventh and thirteenth dorsal plates a little pro- duced as usual, those of the other minor plates oblique or simply rounded. Coxal pores very small and circular; 2, 2,2, 2-2, 3, 3, 2 : 0, 0, 0, 0, 1. Glonossialh 0710; 0,081: Spines of first legs, MONON? of the second, See aR OT ventral spines of fourth and fifth, 0, 0, 0, 0, 1 and of the sixth the same or else 0, 0, 0, 1, 1; spines of seventh legs, 21-2; of eighth and 2NOMO TOs Lane . 0, 0, 0, 1, 2 OnOnGn2 2. 2 ninth, wae of tenth and eleventh, Gnosueoaa of twelfth, 00;/0; 152° 0, 0, 2, 1, 0. 0, 0, 2, 0, 0. equate of thirteenth, ar ane of the penult, aoa ae: anal lost. Penult legs of male short and uniformly inflated. Process at distal. 52 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. end of tibia small, cylindric at base and expanded at free end on side toward distal end of joint; process extending dorsodistad or also a little caudad; sensory hairs on distal surface of process short and spinescent. (Plate 3, fig. 5-6). Length of type 7 mm.; width of tenth plate .9 mm.; length of antenna, 2.3 mm. Locatitry.— Tennessee (Russellville!). NAMPABIUS GEORGIANUS, sp. nov. Draanosis.— Dorsal spines of anal and penult legs 0, 0, 2, 0, 0; ventral 0, 1, 1, 1, 0. Dorsal spines of thirteenth legs 0, 0, 2, 1, 0; ventral 0, 0, 2, 1,1. Dorsal spines of twelfth legs, 0, 0, 1, 2; ventral, 0,0, 0, 1,1. Dorsal spines of eleventh legs, 0, 0, 0, 2,2; of the tenth, 0, 0,0, 1,2. Ventral spines of tenth and eleventh legs (as also of the sixth to ninth) 0, 0, 0, 1, 1. Ocelli 10-12 in three series. Prosternum 1.5 times wider than long; distance between chitinous spots only 3.1 times as great as dental line; about 23 times as great as width at median incision. Body 6.6 to nearly 7 times longer than width of tenth dorsal plate. Head much wider than long. First dorsal plate twice as wide as long or nearly so. Process of tibia of penult leg of male small and subcylindrie, (Plate 2; fig. 2). Dorsum testaceous. Head light ferruginous; frontal region palest. DescripTion.— Dorsum testaceous. Head somewhat light fer- ruginous with caudal portion covered with a darker brown area, leaving the frontal region palest. Antennae somewhat pale ferrugin- ous brown, lighter at tips. Prosternum brown of more or less reddish caste, the prehensorial feet paler. Venter pale yellow, with the caudal plates darker. Legs like adjoining part of venter with caudal pairs somewhat brighter, distally. Body strongly narrowed cephalad, moderately robust, being from 6.6 to 6.9 times longer than width of the tenth plate, being thus relatively more slender than in N. virginiensis. Widths of head and of first, third, eighth, tenth, and twelfth plates as 37: 34: 38: 46: 48: 43. Head much wider than long (average ratio about 33.5: 29.5), being wider by from $th to 7oth. Broadest a little back of eyes where it bulges considerably on each side. Broad anteriorly, not subcordate CHAMBERLIN: LITHOBIID GENERA. 53 as in virginiensis. Hairs moderately short, very sparse. (Plate 8, fig. 8). Antennae short and but moderately attenuated. Articles, as usual, 20, which are short though rather longer than in virginiensis and also less closely telescoped; decreasing distad from the second. Hairs moderately short, not especially dense. Eyes composed of ten to twelve ocelli arranged in three series: — thus 1+4, 4, 3-1+4, 3, 3. Ocelli black; closely but somewhat irregularly arranged. Prosternum 1.5 times wider than long. Distance between chitinous spots 3.1 times as great as the length of dental line; about 23 times the width at level of bottom of median incision which is clearly farther cephalad of caudal ends of lateral slopes than in vwirginiensis. Sides or slopes of anterior portion straight, slanting directly ecto- caudad from the teeth. Teeth small and acute. Median incision conspicuous, of the usual semicircular form. First dorsal plate widest at anterior end, the sides at first moderately converging and then strongly rounding in about the caudal corners; about twice as wide as long. Ninth plate with caudal margin bent forward a little at ends. Eleventh plate with caudal margin straight all the way across or very nearly so. Thirteenth plate with caudal corners a little produced. Coxal pores small; >, Deoaaee Spines of first legs, % a ae of second, ao oOo i 1; of third, O508001, 20, OnOnOn ls 1. 0),:0; Ot: ae of fourth and fifth, 5-5 ee of sixth, SSS of On 0) Ona, 2: : 0, 0, 0, 2, 2. eighth to tenth inclusive, carci of eleventh, SES of O00 12 0205.2) 1, @. twelfth, ahs of thirteenth, 3 Sees of penult and anal, 0, 0, 2, 0, 0 mete each with two claws. In the male the anal legs are short; uniformly and but little crassate. (Plate 2, fig 3). Penult legs similar to anal but bearing the charac- teristic lobe at distal end of the fifth joint, or tibia; lobe small and subcyclindric, proportionately longer than in virginiensis (Plate 2 fig. 1-2); extending in a mesodorsal and somewhat distal direction, bearing on the free surface a limited number of rather short sensory hairs. Length 5 to 6.25 mm. Locauity.— Georgia (Tallulah Falls!). 54 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. NAMPABIUS CAROLINENSIS, Sp. Nov. Draqnosis.— Dorsal spines of the anal and penult legs, 0, 0, 2, 0, 0; ventral of penult, 0, 1, 2, 1, 0, of anal 0, 1, 1, 1, 0. Dorsal spines of thirteenth legs, 0, 0, 2, 1,0; ventral, 0, 0, 2, 1,1. Dorsal spines twelfth legs, 0, 0, 1, 1, 2. Dorsal spines of tenth and eleventh legs, 0, 0,0, 2,2. Ventral spines of ninth to twelfth (as also of seventh and eighth) legs, 0, 0, 0, 1, 1. Ocelli near 6 in number, in two series. Prosternum 1.6 times wider than long. Distance between chitinous spots only 2.1 times width at median incision; 3.5 times as great as dental line. Body near eight times longer than width of tenth plate. Head conspicuously wider than long. First dorsal plate near 1.7+ times wider than long. Process of tibia of penult leg in male relatively slender and moder- ately long, broadening at base but not distally where it is rounded. Hairs at distal end straight and divergent. (Plate 2, figs. 4-5). Dorsum yellowish, uniform. Head reddish orange. DescriptTion.— Dorsum yellowish, uniform. Head reddish orange. Antennae brown, pale distally. Prosternum somewhat paler than head, the prehensorial feet still lighter. Venter light yellow, the caudal plates orange. Legs nearly like corresponding plates of venter. Body moderate, being 7.9 or 8 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other as 36: 31: 34: 42: 43: 41. Head conspicuously wider than long (9: 8), widely rounded in front of eyes; sides well rounded caudad of eyes. Hairs of moderate length and sparse, as usual. Antennae short and moderately attenuated. Articles beyond the second short and very short, decreasing distad. Clothed as usual. Eyes composed of about 6 ocelli arranged in two series: — thus, 1+3, 2. Ocelli distinct, regular in form and arrangement. Prosternum 1.6 times wider than long. Distance between chitinous spots 2.1 times width at median incision; 3.5 times the dental line. Median incision wide and shallow, strictly semicircular in form. Teeth small, the outer of each pair rather larger than the inner, the interval between the two well rounded; line of apices strongly re- curved. Spine contiguous with base of outer tooth, fine. Lateral CHAMBERLIN: LITHOBIID GENERA. 55 margin of anterior portion beginning at spine and sloping ectocaudad; a little convex, bent ectad near caudal end for a short distance. First dorsal plate 1.72 times wider than long; sides convex, widest a little in front of middle of length. Posterior angles of thirteenth plate a little produced; those of the eleventh slightly moreso. Caudal margin of other minor plates more or less bent forwards at ends; or corners of the ninth obliquely excised. Coxal pores small and circular; 2, 2, 2, 2. Spines of first to third legs inclusiv eo 0.0, 0.1, of fourth and fifth, 0:00) Fn 0,080) 1.2) ; atk GLO, Oty 2. fnEoatee? of sixth, 0,0,0,0,4) Soa) of seventh and eighth, 9-9-977'4; SOs, Ct aoe Fee : of tenth and eleventh, ° Sooo: = ; of twelfth, GEusOsE Te of thirteenth Oi0t ir bso: O205:2,105 0, 0, 2, 0, 0 Shea of penult, eee OT : Melek of the anal, eat ass 0? armed * with two claws. In the male the anal legs are short and moderately, uniformly in- flated. The penult legs are similar excepting for the tibial lobe; lobe a relatively slender and moderately long cylindric process which widens at base but distally is rounded and not at all expanded; distal hairs straight and divergent. Length of type 6.5 mm. Locauity.— South Carolina (Landrum! August, 1910). NAMPABIUS TENNESSEENSIS, Sp. Nov. Diaenosis.— Dorsal spines of anal and penult legs 0, 0, 2, 1, 0; ventral, 0,1,1,1,0. Dorsal spines of twelfth legs, 0, 0,2,1,0. Dorsal spines of tenth and eleventh legs (as also of the first to ninth), 0, 0, 0, 1, 1. Ventral spines of ninth to twelfth legs (as also of the fifth or sixth to eighth), 0, 0, 0, 1, 1. Ocelli 8 or 9 in two or three series. Prosternum 1.54 times wider than long. Distance between chitin- ous spots 24 times width at median incision; 3.5-3.8 times the dental line. Body eight times longer than width of tenth plate. Head wider than long. Process of tibia of penult legs of male flattened in a distoectal proximomesal direction; considerably longer along base than the height above same; expanded at distal end. (Plate 3, fig. 3-4). Dorsum pale chestnut. Head darker chestnut, shining. Description.— Dorsum pale chestnut. Head a deeper, shining 56 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. chestnut. Antennae chestnut, excepting the tips which are lighter. Prosternum rather more brownish than the head, with the prehenso- rial feet lighter and at tips somewhat light ferruginous. Venter pale; the anterior plates suffused with purplish; the caudal plates burnt brown. Legs pale, excepting the posterior pairs which are brown or brownish orange. Body moderate, in the male type being near eight times longer than wide. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 36: 30: 32: 41: 42: 40. Head wider than long in ratio 12:11; not varying much in width between eyes and level of marginal breaks; broadly rounded cephalad. Hairs short and sparse. Antennae short as in related species. Moderately attenuated. Articles short, decreasing distad. Hairs on proximal articles rather sparse, becoming denser distad. Eyes composed of eight or nine ocelli arranged in two or three series: — thus, 1+4, 3; 1+3, 3,2; 1+4, 3, 1. : Prosternum 1.54 times wider than long. Distance between chitin- ous spots 23 times the width at level of bottom of median incision; 3.8-3.5 times the dental line. Median incision wide and shallow, sides strongly converging, rounded at bottom. Teeth moderately small, acute; line of bases forming a strongly reentrant angle and the line of apices accordingly recurved. Sides of anterior portion sloping immediately from the outer teeth, straight, slanting more strongly ectad than usual. First dorsal plate widest near the middle, from where the sides are subparallel cephalad and rather strongly converging caudad. Caudal angles of eleventh and thirteenth dorsal plates a little produced in the typical manner; those of the ninth straight or a little oblique. Coxal pores small and circular, the proximal ones in each series somewhat smaller than the distal; 2, 2, 2, 2 or 2, 3, 3, 2. Spines of first legs,f0-0.0.1.1. of the Loa On070 stl sopethie 0, 0, 0, 0, 0” SOBER De 020)10)1 1, 0,:0,:0, 1, 1 OsOfOR NIC third and fourth; ioacacirat of the fifth, ae 0 ; or iD Sie ae 010, Oats ‘ 0,25 a of the sixth to eleventh, paaonat of the twelfth, ° erst of the thirteenth, 90 2» 1,0. of the penult, 2% 2-9 © claws 2: of the anal, 0,01, ta? Pp MORE EIR SIAKO ar pace 0, 0, 2, 0,0 2. ges aaieaiean 5» Claws 2 4 | Anal legs of the male short; uniformly and but little thickened. Penult legs as usual; lobe or process extending dorsocaudo-distad, flattened in a distoectal and proximo-mesal direction, considerably CHAMBERLIN: LITHOBIID GENERA. 57 longer along line of attachment than its height above surface of joint, at distal end expanded over proximal portion, hairs at distal end of process as usual. (Plate 3, fig. 3-4). Gonopods of male broad but very short, rounded; bearing two bristles. Length of male type, 6.25 mm.; width of tenth plate, .78 mm.; length of antennae 1.75 mm.; of anal legs, 2 mm. Locatiry.— Tennessee (Russellville!). NAMPABIUS LONGICEPS, sp. nov. DiaGnosis.— Dorsal spines of anal and penult legs 0, 0, 2, 0, 0; ventral, 0, 1, 1, 1, 0. Dorsal spines of thirteenth legs, 0, 0, 2, 1, 0; ventral 0, 0, 1, 1, 1. Dorsal spines of twelfth legs 0, 0, 2, 1, 1- 0,0, 1, 1,1. Dorsal spines of eleventh legs, 0, 0, 0, 0, 1; of tenth (as also of the second to ninth), 0, 0,0, 1, 1. Ventral spines of ninth to twelfth pairs of legs (as also of the fourth or fifth to eighth), 0, 0, 0, 1, 1. Ocelli 7 or 8 in two series. Prosternum about 1.48 times wider than long. Distance between chitinous spots 2.15 times the width at median incision; 3.5 times the dental line. Median lobe of claw of female gonopods but little longer than the lateral. Spines wide with the sides parallel from base to the very short, acuminate distal portion. Body slender, being 8.5 times longer than the width of the tenth plate. Head only slightly wider than long, less rotund than in N. lulae. First dorsal plate 1.8++ times wider than long. Dorsum testaceous with anterior and posterior plates darker. Head reddish or chestnut. DEscrIPTION.— Dorsum testaceous, with the anterior and the posterior plates darker. Head reddish or chestnut. Antennae brown, pale at tips. Prosternum brown. Venter very pale, with the caudal plates darker and the anterior ones strongly suffused with violaceous. Legs pale like the venter, the posterior pairs darker brown. Body comparatively slender being about 8.5 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 36: 30: 33: 41: 41: 39. Head only slightly wider than long (36:35); less rotund than in lulae. Hairs moderate in length, few. 58 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae very short, reaching only the fourth or fifth segment. Articles short and very short, decreasing distad as usual. Hairs of moderate length, rather sparse proximally, becoming more dense distad. Ocelli 7 or 8; arranged in two series: — thus, 1+3, 3; 1+4, 3. Prosternum 1.48— times wider than long. Distance between chitinous spots 2.15 times the width at level of median incision; 3.5 times as great as the dental line. Teeth subequal, acute, with sides. nearly straight but with interval on each side narrowly rounded at the bottom; line of apices recurved. Median incision well rounded at bottom as usual; but the sides more nearly straight and less converg- ing. Sides of anterior portion but slightly convex, sloping ecto- caudad immediately from base of spine. First dorsal plate widest near anterior end; the sides moderately converging caudad; 1.8 times wider than long. Thirteenth plate with caudal angles a little produced; but the eleventh and ninth with posterior margin straight all the way across or nearly so, or the eleventh showing a tendency for the et to be weakly produced. Coxal pores very small; 2, 3, 3, 3. Spines of the first legs, °: aah 3; of the second, i a the third, BO of the fourth, . - 5 a: Tore os “ a +; of the fifth to tenth oso t of the eleventh, je OF of the twelfth, 0, OTE OS OMIT 0, 0,2, 1,0 puro mimi anor Ota of the thirteenth, 5-5-7°4°7; of the penult oy2o2 0 . * and anal, 2 Tae with two claws. Claw of the gonopods of female short, moderately curved, tripartite; lobes short and acute, the median being but little longer than the lateral. Basal spines wide with sides parallel from base to the acumi- nate distal portion which is relatively very short with its straight sides converging to an almost obtuse angle, its outer side being longer than itsinner. Inner side of first article of gonopods a little convex distally and slightly concave wees subparallel with its fellow. Length of type 6.5 mm.; of antenna, 1.9 mm.; of anal leg, 1.8 mm.; width of tenth dorsal re .76 mm. Locauiry.— North Carolina (Asheville!). NAMPABIUS LULAE, Sp. nov. DraGnosts.— Dorsal spines of anal and penult legs, 0, 0, 2, 0, 0; ventral, 0, 1, 1, 1, 0. Dorsal spines of thirteenth legs, 0, 0, 2, 1, 1 ventral, 0, 0,0, 1, 1-0, 0, 1, 1, 1. Dorsal spines of twelfth legs, CHAMBERLIN: LITHOBIID GENERA. 59 0, 0, 2, 1, 1; of the eleventh, 0, 0,0, 0, 1; of the tenth (as also of sec- ond to ninth), 0, 0, 0, 1, 1. Ventral spines of ninth to twelfth pairs of legs (as also of the sixth to eighth), 0, 0, 0, 1, 1. Ocelli 7 to 11 in 2 or 8 series. Prosternum 1.55+ times wider than long. Distance between chitinous spots 2.1 times the width at median incision; 3.5 times the dental line. Body about seven times longer than the width of tenth plate. Head subrotund; wider than long: First dorsal plate 1.75 times wider than long. Dorsum rather dark brown, in some of reddish caste. Head deep reddish brown or almost black. Description.— Dorsum rather dark brown, which may be some- what reddish in caste. Head deep reddish brown or almost black. Antennae dark brown proximally, pale at tips. Prosternum brown, the prehensorial feet considerably lighter in shade. Venter dilute greyish brown to light brown, the caudal plates darker brown as usual and the anterior segments commonly distinctly suffused with viola- ceous. Legs like corresponding portion of venter, the posterior pairs being thus darker, brown, paler distally. Body seven times, a little more or less, longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 37:31: 34: 42: 44: 42. Head subrotund, wider than long in ratio 35.5: 33.5. Hairs moder- ately short, sparse. Antennae short, only moderately attenuated. Articles short, decreasing distad as usual. Hairs of moderate length, not especially dense. Eyes each composed of from 7 to 12 ocelli in two or three series: — thus, 1+4, 4,2; 1+4, 4,1; 1+3,3; 14+4,3. Single ocellus relatively large. Prosternum 1.55+ times wider than long. Distance between chitinous spots 2.1 times width at level of bottom of median incision; 3.5 times the dental line. Teeth small and acute; line of apices clearly recurved; interval between teeth of each pair angular. Median inci- sion relatively wide, semicircular, being widely curved at bottom and with sides considerably converging. Spine moderately stout at base, long and acutely, finely acuminate as usual; parallel with ectal edge of outer tooth. Lateral slopes beginning at spine and extending directly ectocaudad. First dorsal plate 1.75 times wider than long; sides from middle to anterior end nearly straight and parallel; caudad of middle strongly 60 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. convexly rounding in mesad about the caudal corners. Ninth plate with its caudal margin straight or a little bent forwards at ends. The eleventh and thirteenth plates with posterior corners only slightly produced caudad. Coxal pores small; on distal portion of coxa; in number 2, 2, 2, 2, mostly. Spines of first legs, 0 0-0-0:1. of second, 0:0 1:1. of third to Bt ie 0, 0, 0, 0, 0? ? 0, 0, 0, 0, U? tenth, Se the dorsal spines on tenth legs being on anterior . ent te 9 ° e side of joint; of twelfth, eet ene the dorsal spine of fourth article being posterior and that of fifth joint anterior in position; of thir- One Datel e002) tit obit . teenth, 9-0:0°1,1~ 00,1. 1,1 the two single dorsal spines situated on posterior side; of penult and anal legs, Oa each with two claws. Anal and penult legs of male short and moderately, uniformly in- flated; fourth joint longitudinally sulcate above, the third and fifth less distinctly so. Process of penult legs present in usual position. Gonopods of male each extending somewhat ectocaudad, the mesal side being very long and convex and the ectal relatively very short, bearing two (or one) bristles. Length from 5 to6.5 mm. A male 6 mm. long has antenna 2 mm. long; anal leg 1.8 or 1.9 mm. long; and the tenth plate .87 mm. wide. Type Locatiry.— Georgia (Lula). Known Locatitres.— Georgia (Lula!); Tennessee (Johnson City! Russellville!). NAMPABIUS LUNDII (MEINERT). Lithobius lundwi Meinert, Myriap. Mus. Haumensis, 1886, 3, p. 12. Description.— The original description, somewhat rearranged, is as follows: Brunneus, capite cum antennis obscuriore, subtus pallidiore, pedibus flavis. Sat gracilis, sublaevis. Capite suborbiculare, fere aeque longo ac lato. Antennae breviusculae, 19-articulatae, articulis longiusculis. Oculi ocellis 6-7, in series 3 digestis. Dentes prosternales bini. Laminae dorsales omnes angulis rectis. CHAMBERLIN: LITHOBIID GENERA. 61 Pori coxales 2, 3, 3, 2, rotundi, parvi vel perparvi. Pedes corporis primi paris calearibus 0, 0, 1; pedes anales calcari- bus 1, 1, 1, 0 armati, unguibus binis; pedes penultimi paris unguibus binis armati. Pedum analium coxae inermes. Pedum penultimorum tibiae in angulo interno appendice parvo nodiformi instructae. Pedes postici breviusculi, crassiusculi. Long. (maris) 6.5 mm. Locatity.— Near New York City (L. Lund): the type is in the Museum at Copenhagen. GARIBIUS, gen. nov. Body considerably narrowed from the tenth segment cephalad to the first, the dorsal plate of which is much narrower than the head a little narrower than that of the third segment. Moderate to slender, the length varying from 7.5 to 9.5 times the width of the tenth plate. Varying from 5 to 9.5 mm. in length. Antennae short; uniformly composed of twenty articles. Ocelli in three or, less commonly, in two series; 6 to 14 in number. Single ocellus typically not larger than the others. Prosternal teeth 2+2; line of apices straight or but little recurved. Median incision v-shaped, strictly angular at bottom. Posterior angles of none of the dorsal plates produced. Claw of the female gonopods in all known species tripartite. Basal spines 2+2, slender. Mesal side of first article nearly straight. Coxal pores small and circular; uniseriate; from 2, 3, 3, 2 to 3, 4, 4, 3. Dorsal spines of the anal legs 1, 0, 2, 0, 0 or rarely 1, 0, 2, 1,0; ven- tral, 0, 1, 3, 1,0; claw 1. Dorsal spines of penult legs 1, 0, 2, 1,0 or 1,0, 2, 1,1; ventral, 0, 1,3, 2,1; claws3. Dorsal spines of thirteenth legs, 1,0, 2,1, 1; ventral, 0, 1, 3, 3, 2 to0, 0, 2, 3,2. Dorsal spines of twelfth legs, 1, 0, 2, 1, 1 or 0, 0, 0, 2, 2 to1, 0, 2,1,2. Dorsal spines of eleventh legs 0, 0, 2, 1, 1 or 0, 0, 0, 2, 2 to 1, 0, 2, 1, 2; ventral, 0, 0, 0, 2,1 to0,0,2,3,2. Dorsal spines of first legs 0, 0, 1, 1, 1 to 0, 0, 2, 2, 1; ventral, 0, 0, 0, 2, 1 to 0, 0, 1, 2, 1. Last three or four pairs of coxae thus dorsally armed; while rarely none of the coxae are laterally armed, in most those of the last pair are, and those of the last two pair may be. Anal legs of male moderately thickened; tibia always bearing at distal end on dorsomesal side a low crest bearing sensory setae 62 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. (Plate 4, fig. 6, 7,9) muchas in the genus Nadabius (collium Chamb., towensis Mein., and allies). The tibia of the penult legs may or may not show a more weakly developed similar crest. Typr.— G. monticolens, sp. nov. DiIsTRIBUTION.— Southeastern States and northward to Pennsyl- vania and New Jersey. These forms seem particularly to prefer to live under the bark of fallen trees. Nore.— In my paper on the Lithobiomorpha of the Southeastern States (Ann. Ent. soc. America, 1911, 4, p. 32-50) several of the species now described were placed together under the name Lithobius brannert Bollman, the several forms being considered tentatively as varieties. The following key is given as an aid in the identification of the species. Key to species of Garibius. a. Dorsal spines of anal legs 1, 0, 2, 1 aa. Dorsal spines of anal legs 1, 0, 2, 0 b. First seven or more pairs of legs with the third joint ventrally unarmed. c. Third joint of all legs dorsally armed; none of the coxae laterally armed; dorsal spines of penult legs 1, 0, 2, 1, 0; of the twelfth, 0, 0, 2,1, 1 o0r1,0,2,1,1.° G. georgiae, sp. nov. ce. ‘Third joint of first ten or more pairs of legs dorsally unarmed; last pair of coxae laterally armed; dorsal spines of penult legs 1, 0, 2, 1, 1; of the twelfth 0, 0, 2, 2, 2. G. alabamae, sp. nov. bb. All legs, or all excepting the first pair, with the third joint armed both dorsally and ventrally. c. Last two pairs of coxae laterally armed. d. Ventral spines of second to sixth legs 1, 3, 1; of Ist, 1, 2, 1, with the dorsal 2, 1, 1; head equal in length and breadth. G. pagoketes, sp. nov. dd. Ventral spines of second to seventh legs 1, 2, 1; of first 0, 2, 1, with the dorsal 1, 1, 1; head distinctly wider than long. G. mississipplensis, sp. Nov. cc. Either none, or only the last pair, of coxae laterally armed. d. None of the coxae laterally armed; dorsal spines of the eleventh legs 2, 1, 2, the ventral 1, 3, 2; ventral spines of the first legs 1, 3, 2. G. dendrophilus, sp. nov. 50} G. catawbae, sp. nov. 50. CHAMBERLIN: LITHOBIID GENERA. 63 dd. Last pair of coxae laterally armed; dorsal spines of eleventh legs 2, 1, 1, ventral 2, 3, 2 or 3, 3, 2; ventral spines of first legs, 1, 2, 1 or 1, 3, 1. e. Ventral spines of eleventh and twelfth legs 2, 3, 2; of second to sixth, 1, 3, 2. G. opicolens, sp. nov. ee. Ventral spines of eleventh and twelfth legs 3, 3, 2; of second to sixth 1, 3, 1. . G. monticolens, sp. nov. GARIBIUS CATAWBAE, Sp. Nov. D1aGnosis.— Dorsal spines of anal and of penult legs, 1, 0, 2, 1, 0. Ventral spines of thirteenth legs, 1, 3, 3, 2 or 1, 2,3, 2. Dorsal spines of twelfth legs, 0, 0, 2, 1, 1; ventral, 0, 0, 2, 3, 2‘ Dorsal spines of eleventh legs, 0, 0, 2, 2,2; ventral, 0, 0,2, 3,2. Dorsal spines of first legs, 0, 0, 2, 1,1; ventral, 0,0, 1, 2,1. Third joint of all legs dorsally and also ventrally armed. Last pair of coxae laterally armed. Antennae as usual. Ocelli about 14 in three series. Prosternum 1.65 times wider than long. Distance between chitin- ous spots 2.7 times the width at median incision; 3.78 times the dental line. Median incision strictly V-shaped. Coxal pores 3, 4, 4, 3. Body about 8.4 times longer than width of tenth plate. Length (of type) 8 mm. First dorsal plate 1.73 times wider than long. Anal legs in the male with the usual crest. (Plate 4, fig. 8-9). Penult legs with the tibia also bearing a sharp longitudinal ridge along dorsocaudal side the form of which at its distal end is some- what suggestive of the crest of the anal leg. Dorsum light chestnut; head scarcely darker. Description.— Light chestnut above. Head scarcely darker, Antennae in color like the head, pale at tips. Prosternum light brown, with the prehensorial feet paler. Venter pale, the caudal plates darker, burnt brown. Legs pale, with tarsi most pigmented; posterior pairs darker, brown or pale chestnut, lighter distally. Body of usual form. Length about 8.4 times width of the tenth dorsal plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 45: 40: 42: 49: 50: 44. Head of equal width and length or very nearly so. Nearly of uni- 64 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. form width between eyes and marginal breaks; narrowed rather more than usual cephalad. Antennae short. Composed of the usual twenty articles which are short, articles more loosely united distally. Eyes composed of about 14 ocelli arranged in three series: — thus, 1+5, 4, 4. Prosternum 1.65 times wider than long. Distance between chitin- ous spots 2.7 times width at level of bottom of median incision; 3.78 times the length of dental line. Mesal incision strictly V-shaped, the sides being straight and meeting in an acute angle at bottom. Teeth acute, with their sides straight; the interval between each pair subacute at bottom or but narrowly rounded; inner tooth distinetly larger than the outer; line of apices straight or very nearly so. Lateral slope beginning at outer tooth; convex for entire length, most strongly so near its middle. First dorsal plate with sides convex anteriorly; widest a little back of anterior end from where to the rounded caudal corners the sides are nearly straight and converge conspicuously; about 1.73 times wider than long. Posterior corners of ninth, eleventh, and_ thir- teenth dorsal plates simply rounded. Coxal pores small, regularly decreasing proximad on each coxa; in number, 3, 4, 4, 3. Spines of first legs, ule 204; of the second to oe 0, ot; 2 O, 0, ’ 2, ’ 3, z 0, 0, O, of eighth, 220: 2: 272 op 0:0 2.2;2- of minth, ONO; 2 i 2582) 2 Op Oy ee why ) OFOS UR sy al SS : MOV) Bak 0,0 25rlenle of ke and eleventh, ? nossa of Haas Gaeoaoaes tn ba 1 AO522) “4 10 4 ae One cies of penult, 4 = + 3 at claws 3; ) 9; Claw single. Coxae of anal legs laterally of seventh OO 2 ena ee ONOMINS 2? of thirteenth, 4 = + 3 1. (0) '2, Odes of the anal, aay 1 armed. Anal legs of male moderately crassate, with the tarsi more or less abruptly more slender than the tibia, tibia at distal end with dorso- mesal corner flattened and there bearing the typical low ridge or crest. Dorsomesal edge of crest forming almost a straight line, at distal end convex; bearing a number of moderately long hairs. (Plate 4, fig. 8-9). Joints longitudinally suleate above and laterally. Penult legs. with tibia bearing a sharp longitudinal ridge along dorsocaudal side, this at distal end in appearance suggestive of the crest of anal legs. Length of type (o’) 8 mm.; length of antennae, 2.5 mm.; of anal legs also near 2.5 mm.; width of tenth plate, .95 mm. Locatiry.— North Carolina (Catawba!). CHAMBERLIN: LITHOBIID GENERA. 65 GARIBIUS MONTICOLENS, sp. nov. DraGnosis.— Dorsal spines of anal legs 1, 0, 2, 0,0. Dorsal spines of penult legs 1, 0,2, 1,0. Ventral spines of thirteenth legs, 0, 1, 3, 3, 2. Dorsal spines of twelfth legs, 1, 0, 2, 1, 1; ventral, 0, 0,3, 3,2. Dorsal spines of eleventh legs, 0, 0, 2, 1, 1; ventral, 0, 0, 3, 3, 2. Dorsal spines of first legs, 0, 0, 2, 2, 1; ventral, 0, 0, 1, 2, 1. All legs with third joint armed both dorsally and ventrally. Last pair of coxae laterally armed. Antennae of usual form. (Plate 4, fig. 3). Ocelli about 14 in three series. (Plate 4, fig. 4). Prosternum 1.53 times wider than long. Distance between chitin- ous spots 2.38 or 2.4 times width at median incision; 3.58 times length of dental line. (Plate 4, fig. 1-2). Anal leg of male moderately inflated; tarsi abruptly more slender; tibia bearing a crest in the usual position, this clothed with a subdense series of long hairs. (Plate 4, fig. 6-7). Penult legs without lobes. Body slender; 9.5 times as long as width of tenth plate. Head cordate; equal in length and width or nearly so. First dorsal plate 2.5 to 2.6 times wider than long. Dorsum light brown. Head dilute ferruginous brown or somewhat orange with frontal region paler. Description.— Above, light brown. Head dilute ferruginous brown or somewhat orange, with the frontal region paler. Antennae light brown, paler distad as usual. Prosternum lighter than head. Venter pale testaceous; the caudal plates more deeply pigmented, more or less orange. Legs pale testaceous with the tarsi more pig- mented, the posterior pairs darker, brownish. Body slender, being nearly 9.5 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 54: 46: 47:54: 56: 55. Head cordate. Equal in length and width or very nearly so. Nearly constant in width between level of eyes and that of lateral breaks or a little caudad of then. Antennae short. Composed of the usual twenty articles, these being loosely joined. (Plate 4, fig. 3). Eyes consisting of about 14 ocelli arranged in three series: — thus, 1-+5, 5,3. Single ocellus oval. Ocelli of upper series, especially the more caudal ones, conspicuously larger than the other ones, the ocelli decreasing in size cephalad and ventrad. (Plate 4, fig. 4). 66 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Prosternum 1.53 times wider than long. Distance between chitin- ous spots 2.38—2.4 times width at level of bottom of median incision; 3.58 times length of dental line. Mesal incision v-shaped, with sides nearly straight. Teeth acute, the mesal one larger than the ectal; line of apices straight or slightly recurved. Spine considerably stouter at base than the hairs, slender distally, of moderate length. Lateral slopes convex cephalad, concave and bending ectad caudally. (Plate 4, fig. 1-2). First dorsal plate widest a little caudad of the anterior end from where the sides are nearly straight and moderately converging to the rounded posterior corners; 2.5 to 2.6 times wider than long. Posterior angles of ninth, eleventh, and thirteenth dorsal plates simply rounded, with no tendency toward production. Coxal pores small and circular; on each coxa decreasing regularly proximad. Spines of first legs, o o: . * i; of second to fifth, t a ae of sixth, ro Ot i or 3 z ae of seventh, OOo of eighth, 0.0.2.2, . or SOS 353 of ninth and tenth, OOo so of the, eleventh, eet of the twelfth, eet sg 5) Of ‘the’ thirteenth; ee 1. of the penult, }: wats . 3 », with three claws, of which the anterior is often very eth sometimes spine of coxa is doubled, making the dorsal formula thus 2, 0, 2, 1,0; of the anal, oe the claw single, long. Anal legs of male moderate in length, moderately inflated distad to the tarsus which is abruptly more slender; tibia obliquely complanate at distal end on mesal side toward dorsal surface and there bearing a low subtriangular crest, the elevated end of which is caudad, its mesal, gently convex edge gradually resuming the level of the joint proximad, the crest bearing a subdense series of moderately long hairs which tend to be somewhat curved caudad at their distal ends (Plate 4, fig. 6-7). Penult legs more slightly thickened, without special lobes. Third, fourth, and fifth joints of both anal and penult legs longitudi- nally suleate dorsally. Length of male type 9.5 mm.; width of tenth plate 1.07 mm.; length of antenna, 3.5 mm.; of anal leg also 3.5 mm. Type Locatiry.— Tennessee (Unaka Springs!). Loca.ities.— Tennessee (Unaka Springs!); North Carolina (Ash- ville! Brown’s Summit!). CHAMBERLIN: LITHOBIID GENERA. 67 GARIBIUS OPICOLENS, sp. nov. D1aGnosis.— Dorsal spines of anal legs 1, 0, 2, 0, 0; of penult 1, 0, 2, 1,0. Ventral spines of thirteenth legs, 0, 1, 3, 3, 2. Dorsal spines of twelfth legs, 1, 0,2, 1,1, ventral, 0,0, 2,3, 2. Dorsal spines of eleventh legs, 1, 0, 2, 1, 1; ventral, 0, 0, 2,3, 2. Dorsal spines of first legs, 0, 0, 2, 2,1; ventral, 0,0, 1,2,1,o0r0,0,1,3,1. Third joint of all legs armed both dorsally and ventrally. Last pair of coxae laterally armed. Brown to light chestnut; head subferruginous, dusky caudad_ of suture. Body moderate, being about 8.25 times longer than width of tenth plate. Head broadly subcordate, nearly equal in length and breadth. Eyes composed of from thirteen to fifteen ocelli in three series. Prosternum 1.54 times wider than long, as in G. monticolens. Penult legs without lobes; anal legs moderately inflated with the tarsi abruptly more slender, and the tibia bearing the usual crest which is nearly as in monticolens. Gonopods of female with claw short and wide, tripartite, outer- most lobe smallest; outer spine considerably longer than the inner on each side, sides of spines subparallel to the short and abruptly acuminate distal portion. Descrietion.— Dorsum brown to light chestnut, with nearly always a blackish median longitudinal stripe for all or most of length while two or more of the most anterior plates may be margined, caudally with blackish; fifteenth plate abruptly lighter, yellowish or light orange. -Head subferruginous, dusky caudad of suture with a blackish median spot or stripe on caudal portion. Antennae dark proximally, orange or rufous distad. Prosternum orange or light ferruginous. Venter brown, the last plate orange like the correspond- ing dorsal one, with the immediately preceding one or two also often lighter. The posterior pairs of legs more densely pigmented as usual, the distal joints being light orange. Body of moderate build, being about 8.23 to 8.26 times longer than width of tenth dorsal plate. Head broadly subcordate. Equal in length and breadth or very slightly longer (49:48). Widest just caudad of the marginal breaks from where the sides converge slightly cephalad to the eyes. Antennae short. The twenty articles short, of uniform proportions, 68 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. moderately compactly joined, being less loose than in types of monti- colens; ultimate article long and pointed. Hairs rather long, straight. Eyes composed mostly of from thirteen to fifteen ocelli arranged in three series: — thus, 1-++5, 4, 3; 1+5, 4, 4; 14+5, 5, 4. The single ocellus is typically broad and may be materially the largest; of the others those of the dorsal series are materially the largest, these de- creasing in size cephalad. Prosternum cir. 1.54 times wider than long. Distance between chitinous spots 2.4 times width at level of median incision; 3.78+- times the dental line. Outer tooth of each pair smaller than the inner one, both subacute; line of apices very slightly recurved. _Inter- val between teeth of each pair semicircular. Sides of median incision straight, the bottom well rounded. __Ectal spine materially stouter at base than neighboring bristles, rapidly acuminate to about middle of length, distad of where it is bristle-like; straight and moderately long. Coxal pores circular and small; nearly uniform in size or the most proximal on coxa reduced. Common formulae are these: 3, 4, 4, 3; 4,4,4,3; 4,5,4,3; 4,5, 4,4; 4, 5, 5, 4. 0,0, 2, 2, 1 op 0. 0: 2; 7 = 25 Al 4 — spimes*of “arst legs, o"6"4-o: 4 ©) 00,1, 3, of the second to sixth O70, 2204 pairs, 0-5-4" s'9; of the seventh, eee of the eighth to tenth, »2)2. of the eleventh and twelfth, oe 2 3° a3 of the thirteenth, € +; of the penult, eee with three claws, of which the accessory ones are small but distinct; of the anal, 1-9 2-9 claw > OWS Oo? single, long. The median ventral spines of legs especially long. In one specimen the inner ventral spine of prefemur of both penult legs is bifid for about half of length. Coxae of anal legs laterally armed. Anal legs of male of moderate length, considerably longer than the penult; moderately inflated, with the tibia a little clavately widening distad and the tarsal joints abruptly more slender; tibia at distal end with dorsomesal surface obliquely complanate and there bearing the usual crest, the free edge of which toward caudal end is convex and somewhat lowering distad but materially less so than the general sur- face of joint and sides of crest; crest bearing a series of hairs which are straight or with some of them a little bent caudad at ends, these hairs more slender than bristles of other parts of joint; the crest very simi- lar to that of monticolens, see Plate 4, fig. 6-7. Penult leg of male also moderately inflated, without any crest or special lobe. Prefemur, femur and, less distinctly, the tibia of anal and penult legs longi- tudinally dorsally suleate. CHAMBERLIN: LITHOBIID GENERA. 69 The single article of the gonopods of male small but well exposed. Distally obliquely subtruncate. Bearing on distal surface mostly two long bristles. Claw of female gonopods short and relatively wide; tripartite, the divisions acute and rather longer than usual, the middle lobe but slightly longer than the inner one, the outer one considerably shorter and more slender. Basal spines with outer one of each pair con- spicuously longer than the inner; sides of spines subparallel or slightly diverging from base to the acuminate distal portion which is very short, occupying about one fourth of the total length, its sides strongly converging, its apex narrowly rounded; a denticle usually manifest on margin at beginning of acuminate portion on each side. Length 8 toll mm. A specimen 10.5 mm. in length has antennae 2.6 mm. long and anal legs 3.5+ mm. long. Immaturus.— Body pale yellow or dilute brown tinged with violaceous. Head yellow or pale orange. Anal legs whitish to yel- low. Antennae pale yellow. Antennae very short. Articles twenty, these distad of the fourth or fifth being very short, proportionately more so than in the adult, closely compacted. Hairs long, not dense. Eyes composed of six to eight ocelli arranged in two series: — thus, 1+3, 2; 1438, 3; 144, 3. Ocelli pale. Single ocellus somewhat largest; caudal two of dorsal row large, subequal the most anterior and those of the lower row reduced. Prosternum practically as in the adult as regards the form and rela- tions of teeth and ectal spines. Teeth pale. Coxal pores small, with the most proximal one on each coxa only very slightly smaller than the distal one; 2, 2, 2, 2. Spines of first legs, CEE ore 0. 0, (0, 9.0.0, 1,1; oy 0.0, 1,2, 0, 0, 0, 1 oat G) ovo; nt spp of the 0.0, 1, 2.1 sixth, op os0Fa-1 > + <>: of the eighth, () lar | second to fifth pair, . ie} ROVE On 04 Tea 21: 0, 0, 1, mle tOly oo of the sev a: 00: to ONOP 2a B09 1st SS eee ot “iite fee 2 2, of the tenth, Ouest to 0. Ona: of the twelfth, 2 1 ee f WONZ Ie te: 07.0; 2 B.0 05.0; See of the thirteenth, oa Eee O44 es of the deals coxae armed laterally as in the adult. Claw of female gonopods very small and weak, consisting of a single pointed process or with the lateral divisions indicated as slender ojo oO ee Led ce Ll ae io - w bo ; of the penult, 70 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. spinous points at base of the median process. Either two or three articles present. Basal spines 1+1; slender and acutely acuminate from base distad. Length 4.75 to 5.75 mm. Agenitalis I.— Body very pale, suffused with violaceous. Head, prehensorial feet and prosternum light orange. Antennae dilute yellow. Legs almost without pigment excepting the caudal pairs which are yellow. Antennae composed of seventeen articles. Distally submoniliform. Bristles long, subsparse. Ocelli four in two series: — thus, 1+2,1. The single ocellus smaller than the caudal.one of the upper series; the two ocelli of upper series well separated. Prosternal teeth and spines of nearly same form and relations as in the adult. Coxalsperes, lly, te Anal glands still evident. Length cir. 4 mm. Type Locatity.— New Jersey (Masonville!). Locauities.— New Jersey (Masonville! Camden! Westfield!) ; Pennsylvania (Upsal! Philadelphia!). In New Jersey this species is found occurring commonly under the bark of fallen trees in pine woods. This form is evidently very close to monticolens with which the first inclination was to unite it; but the examination of a considerable number of specimens from the localities above listed showed no varia- tion in a number of differences presented. The most definite and readily usable difference is in the spining of the eleventh and twelfth pairs of legs, the formula for the ventral surface of which in monticolens is 0, 0, 3, 3, 2 whereas in all specimens of opicolens examined it is uni- formly 0, 0, 2, 3, 2. A larger number of the posterior coxae are dor- sally armed, but this in other species is subject to considerably varia- tion so that it may not prove of distinctive value in the present case. Garibius opicolens is more robust and also presents well marked differ- ences in color and color-pattern from that shown by the individuals of monticolens thus far observed. GARIBIUS DENDROPHILUS, sp. NOV. D1aagnosis.— Dorsal spines of anal legs, 1, 0, 2, 0, 0; of the penult, 1,0,2,1,0. Ventral spines of the thirteenth legs, 0, 0,3, 3,2. Dorsal CHAMBERLIN: LITHOBIID GENERA. TA spines of the twelfth legs, 1, 0, 2, 1, 1; ventral, 0, 0, spines of the eleventh legs, 0, 0, 2, 1, 2; ventral, 0, 0, spines of the first legs, 0, 0, 2, 2, 1; ventral 0, 0, 1, : coxae laterally armed. Ocelli mostly near twelve, in three series. Prosternum cir. 1.46 times wider than long. Penult legs of male without lobes and the crest of tibia of anal legs almost obsolete. Head subcordate; wider than long. Robust, in type not fully seven times longer than width of tenth plate. Dorsum reddish brown, with caudal margins of plates darkened; head light ferruginous, darker along caudal portion. DescriptTion.— Dorsum reddish brown, somewhat darker along caudal borders of plates; last two plates brighter, more reddish in color. Head light ferruginous, darker across caudal portion. Pro- sternum yellow ferruginous, the prehensors being yellow. Antennae yellow prox imally and more or less violaceous brown elsewhere except- ing at very tip which is again lighter. Venter light brown of viola- ceous tinge; the anterior plates somewhat darker, more reddish, and the posterior ones decidedly darker, subferruginous. Legs pale; the anal and penult pairs bright yellow excepting at bases. Body considerably more robust than in G. monticolens and opicolens, in the type being not fully seven times as long as width of tenth dorsal plate. Head subcordate. Widest at or a little in front of marginal breaks between which and the eyes the sides are convex; sides converging from breaks caudad about the well rounded corners to the caudal margin which is straight or slightly incurved. Anterior margin between eyes substraight, crenately indented a little each side of median line. Hairs of moderate length; sparse. Wider than long about in ratio 58:55. Antennae rather short; 2.75— times longer than the head. Com- posed of the usual twenty articles which decrease uniformly in diameter and length distad; rather loosely joined; ultimate article relatively shorter than in monticolens and opicolens. Hairs long and straight, of moderate denseness. Eyes consisting mostly of about twelve ocelli arranged in three series: — thus, 1+5, 4,2; 1+5,3,2. Single ocellus largest. Prosternum cir. 1.46 times wider than long. Distance between chitinous spots nearly three times (in type 2.9—) as great as width at 2 aec2) Dorsal 1, 3; 2. Dorsal 2 Yone of the Zs b ie BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. level of bottom of median incision; 3.84 times the length of the dental line. Mesal incision v-shaped, subacute at bottom but with the converging sides a little concave. Laine of bases of teeth distinctly recurved; but the inner tooth on each side larger than the outer so that the line of apices is nearly straight. ‘Teeth acute, the interval between those of each pair angularly rounded at bottom. Ectal spine stouter at base than the bristles, rapidly acuminate, short. Coxal pores moderately small; in type, 2, 3, 3, 2. The single article of the gonopod of male broad but short and only slightly exposed. Broadly truncate. A moderately long median apical bristle and at least one more ectal in position. Spines of first to eighth pairs of legs, 2-°-?-2-1; of ninth and tenth OROMIS B22 > F pairs, CE of the Batt 80,1, 8.29 of the twelfth, 1, 0, 2, 1, 1. 1 FON Oe 0,072,379) Of the thirteenth, o, 7 - : +; of the penult, cpa of the anal, a + 8 O: aa Length cir. 7.5 mm. Length of antennae 2.7 mm.; of anal legs, 3+ mm. Locatity.— Pennsylvania (Philadelphia!). One male thus far known. Like the preceding species, the type of this one was found under the bark of a fallen log. This species appears to be related somewhat closely to the two pre- ceding ones; but it is very readily distinguishable through lacking a lateral spine on the anal coxae; the ventral spines of the anal leg being 0, 1, 3, 2, 0 instead of the more usual 0, 1, 3, 1, 0; the absence of spine from trochanter of thirteenth legs; the dorsal spines of the eleventh legs being 0, 0, 2, 1, 2 instead of 0, 0, 2, 1, 1 or 1, 0, 2, 1, 1, and the ventral 0, 0, 1, 3, 2 instead of 0, 0, 2, 3, 2 or 0, 0, 3, 3, 2; the ventral spines of the first legs being 0, 0, 1, 3, 2 instead of 0, 0, 1, 2, 1 or 0, 0, 1, 3, 1; through the almost obsolete crest of tibia of anal legs; and finally through apparently greater robustness. claw one. None of the coxae laterally armed. GARIBIUS PAGOKETES, sp. nov. D1acnosis.— Dorsal spines of anal legs 1, 0, 2, 0,0. Dorsal spines of penult legs, 1, 0, 2, 1,0. Ventral spines of thirteenth legs, 0, 1, 3, 2,2. Dorsal spines of twelfth legs 0, 0, 2, 1, 1; ventral 0, 0, 2, 3, 2. Dorsal spines of eleventh legs 0, 0, 2, 1,2; ventral, 0,0,2,3,2. Dorsal spines of first legs 0, 0, 2, 1, 1; ventral, 0,0, 1, 2,1. All legs with the CHAMBERLIN: LITHOBIID GENERA. 73 third joint armed both dorsally and ventrally. Last two pairs of coxae laterally armed. Antennae of usual structure and relations. Ocelli 10 in two or three series. Prosternum 1.65— times wider than long. Length (type) 8 mm. Body eight times longer than width of tenth plate. Head equal in length and breadth. First dorsal plate 1.65 times wider than long. Dorsum brown, darker caudad. Head light in front of suture. Antennae rufous yellow distad. DescripTion.— Dorsum brown, the caudal plates becoming some- what dilute chestnut. Head pale in front of the suture. Antennae brownish, distally lighter, yellow of somewhat rufous tinge. Legs brownish yellow, the anterior pairs lightest, caudal pair more yellow distally. Venter light brown, darker, more reddish, caudad. Pro- sternum testaceous. Head subcordate; equal in length and breadth; of nearly uniform width between eyes and lateral breaks or a little widest at the latter; widely rounded from the breaks caudad about the corners. Antennae short. Articles distad of the third short and very short, the distal ones proportionately somewhat longer than the more proximal. Eyes in type consisting of ten ocelli arranged in two or three series: — thus, 1+5, 4 to 1+4, 3, 2. Single ocellus large. Prosternum about 1.65— times wider than long. Distance between chitinous spots 3.75 times width at median incision, 4 times the dental lines. First dorsal plate 1.65 times wider than long. Posterior angles of eleventh and thirteenth dorsal plates obliquely excised. Coxal pores small, circular, decreasing proximad; in number, 3, 3, 3, 3. Spines of first legs, (» oe 2 5 “4; of second to sixth, EE of . 0,0, 2, 2,2. 0, 0, 2. seventh to ninth, OjUuLaaS: of tenth, RENEE of eleventh, 0, 0, 2, 1, 2. s 0, 0, sa ie TaGaoeaes OF a twelfth, OSS 5; of the thirteenth, > = aL of sel O 1082710710 r the penult, § ae eT claws 3; of the last, 1 Resa claw single. Claw of Pb gonopods proportionately wide, triparite, the divi- sions not differing much in length, acute. Inner spine of each pair markedly shorter than the outer (maturus). Spines not acuminate 74 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. from base, but somewhat constricted near middle of length and of greatest width immediately below the flattened acuminate opical portion, which is short. In the type (Q ) the widths of head, and first, third, eighth, tenth, and twelfth dorsal plates are as 50: 46: 48: 56: 57: 56. Length of type 8 mm.; width of tenth plate 1 mm.; length of antennae 2.4 mm.; of anal legs about 3 mm. Type Locaiiry.— Massachusetts (Blue Hills!). The closest allies of this species would seem to be G. monticolens, opicolens, and dendrophilus, from which it is to be separated by the characters assigned in the key and the diagnosis. The male is not known. GARIBIUS MISSISSIPPIENSIS, sp. nov. DtiaGnosis.— Dorsal spines of anal legs 1, 0, 2, 0,0. Dorsal spines of penult legs 1, 0, 2, 1,0. Ventral spines of thirteenth legs 1, 3, 3, 2. Dorsal spines of twelfth legs 1, 0, 2, 1, 1; ventral, 0, 0,2,3,2. Dorsal spines of eleventh legs 1, 0, 2, 1, 2; ventral, 0, 0, 2, 3, 2. Dorsal spines of first legs, 0,0, 1, 1,1; ventral, 0,0, 0, 2,1. Alllegs with the third joint dorsally armed, and all excepting the first with this joint also ventrally armed. Last two pairs of coxae laterally armed. Ocelli about 12, in three series. Antennae as usual. Prosternum 1% times wider than long. Distance between chitinous spots 2.5 times width at median incision; 43 times the dental line. Incision strictly v-shaped. Body 7.5 times longer than width of tenth plate. Length (type), 6.75 mm. Head cordate; clearly wider than long. First dorsal plate 1.8++ times wider than long. Fifth article of anal leg of male raised into an edge along dorso- mesal side with the ridge at caudal end elevated into a crest bearing a number of hairs of the usual character. Penult legs without lobes. Pale testaceous with the posterior plates orange. Head orange or hight ferruginous. DeEScRIPTION.— Dorsum pale testaceous with the posterior plates orange. Head orange or light ferruginous. Antennae yellow. Pro- sternum orange of paler shade than head. Venter pale with cau- dal plates more or less orange. Legs pale excepting the posterior pairs which are bright orange. CHAMBERLIN: LITHOBIID GENERA. (5) Body more robust relatively than in related species; 7.5 times longer than width of tenth dorsal plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 43:37: 38: 46: 47: 46. Head cordate. Widest a little back of eyes from where the sides gently converge caudad; rather strongly narrowed cephalad. Wider than long in about ratio 43:40. Hairs rather short, sparse. Antennae short, reaching the sixth segment or nearly so. Composed of the usual twenty articles which decrease in size distad from the second, and, excepting those at the two ends, are very short. Hairs short and not very dense. Eyes composed mostly of about 12 ocelli which are arranged in three series: — thus, 1+5, 4, 2; 1+4, 4,3. Single ocellus small, not larger than others. Seriate ocelli not contiguous, the caudal ones not larger than the anterior; all small. Prosternum 12 times wider than long. Distance between chitinous spots 2.5 times width at level of bottom of median incision; 43 times the dental line. Median incision strictly v-shaped, being angular at bottom with sides straight or but very slightly concave. Teeth small and acute; the inner slightly larger than the outer; line of apices weakly recurved or straight. Spine inserted a little ectad of outer tooth as usual; slender, straight. First dorsal plate widest at or near anterior end, the sides at first gently and then more strongly converging caudad; 1.8+ times wider than long. Caudal corners of ninth, eleventh, and thirteenth dorsal plates straight or a little rounded, none of these either at all produced or excised. Coxal pores very small: 2, 3, 3, 2-3, 3, 3, 3. Spines of first legs, ate 7. 5 ; of the second to seventh inclusive, ae of the eighth, oe e a = 2 the ninth, eT of the tenth, 9-0-3235; of the elev enth, 5-3 = 3 z ee of the twelfth, + = ae of the thirteenth, oe Ss) of the penult, sey with three claws; of the anal, (92-9, claw single. Last two pairs of coxae laterally armed. Anal legs of male short and moderately inflated; fifth article nar- rowed to an edge along dorso-mesal line with at caudal end of this ridge a low, dorsally gently convex, crest bearing a number of hairs of the usual character; longitudinally sulcate laterally. Penult legs less inflated; without lobes. 76 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Length of type 6.75 mm.; width of tenth plate, .9 mm.; length of antenna, 2.25 mm.; of anal leg, 2.5 mm. Loca.ity.— Mississippi (Brookhaven!). GARIBIUS GEORGIAE, sp. nov. Diacnosts.— Dorsal spines of anal legs, 1, 0, 2, 0,0. Dorsal spines of penult legs, 1, 0,2, 1,0. Ventral spines of thirteenth legs, 0, 0, 2, 3, 2. Dorsal spines of twelfth legs, 0, 0, 2, 1, 1-1, 0, 2, 1, 1; ventral, 0, 0, 2, 3, 2. Dorsal spines of eleventh legs, 0, 0, 2, 1, 1; ventral, 0, 0, 2, 2,2. Dorsal spines of first legs, 0, 0, 1, 1, 1; ventral, 0, 0, 0, 2,1. Third joint of all legs dorsally armed; but about first eight or nine pairs with this joint unarmed ventrally. None of the coxae laterally armed. Antennae as usual. Ocelli about 9 to 11 in three series. Prosternum 1.9 times wider than long. Distance between chitin- ous spots 2.44 times width at median incision; 3.44 times the dental line. Coxal pores 2/'3;.3, 2—2, 6, os Claw of gonopods of the female small; tripartite; lobes acute with the median considerably longest. Outer spine longer than the inner; moderately long and slender, attenuated from the base distad, the sides more strongly rounding in toward the apex. Anal leg of male modified as usual, the fifth article bearing above a low ridge elevated into a low crest at caudal end. Length 5.5 to 7.5 mm. Body 9 times as long as width of tenth plate. Head equal in length and breadth. First dorsal plate 1.84 times wider than long. Dorsum yellow, with caudal plates verging to light orange. Head light orange. Description.— Yellow above, with the caudal plates tending toward light orange. Head light orange. Antennae yellow. Pro- sternum paler than head, being more of a lemon yellow. Venter very pale excepting the caudal plates which are like the prosternum. Legs pale, almost whitish, excepting the caudal pairs which are orange brown. Rather slender, being nine times as long as the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 40: 35: 36: 42: 43: 40. CHAMBERLIN: LITHOBIID GENERA. 77 Head equal in length and breadth or in the male it may be a little longer. Widest a little caudad of eyes from where the sides moder- ately converge caudad; rather strongly narrowed cephalad. Hairs short and sparse. Antennae short; moderately attenuated; composed of twenty articles, most of which are short and very short, the ultimate relatively long. Hairs of moderate length, not very dense. Eyes each composed of from 9 to 11 ocelli arranged in three series: — thus, 1+4, 4, 2; 144, 3,3; 1+4, 3,1; 144, 3,2. Single ocellus of same size as or a little smaller than those of the upper series, the seriate ocelli decreasing in size cephalad and ventrad. All ocelli pale. Prosternum 1.9+ times wider than long. Distance between chitinous spots 2.44 times the width at level of bottom of median in- cision; 3.44+ times the length of the dental line. Median incision rather narrowly v-shaped, the sides straight. Teeth nearly equal or with inner on each side somewhat larger making the line of apices straight or nearly so; acute, with sides strictly straight and the inter- val between the two of each pair angular, not at all rounded. Spine fine, but slightly stouter than the hairs; straight; inserted a little ectad of base of outer tooth. Lateral edges of anterior portion gently convex cephalad and bending ectad caudally as usual. First dorsal plate with sides almost parallel from caudal third cephalad; 1.84 times wider than long. Posterior corners of the ninth, eleventh, and thirteenth plates rounded or rather obliquely excised. Coxal pores very small; 2, 3, 3, 2-2, 3, 3, 3. Spines of first legs, SOO : I; of the second and third py a! /Os2: 0, on 0, 0, 2, 2,1. Fie os "ove OFONNF aaa of the seventh, 2; of the : of the 1 ods Ov 2 of the fourth to sixth, 0,0, 0,2 h GaOmenle = O10) 25251. 0,0,0, 2.1% 0:0 of the eighth more OwOy Si 2h2s wie ols , ele Flt selene Pe, Onset as oS ee ete. aes 2 rif Ono WO ware O. thinteenth, 00,3, 3-2) of the penult, 9-7 -3°9" LOZ OF0 34-319 the claw single. None of the coxae laterally armed. Anal legs of male short and moderately inflated; longitudinally suleate laterally but not dorsally; fifth joint complanate above, extended in a low, rather sharply edged, longitudinal ridge along dorso- mesal side which ends at caudal end in the usual crest, this being low with its free edge a little convex and bearing a series of hairs which are claws 3; of the anal, 78 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. curved at distal ends. Penult legs without any special modifications; moderately and uniformly inflated. Claw of female gonopods small; considerably curved; tripartite; lobes acute with the median considerably longest. Outer of basal spines larger than the inner; moderately long and slender, a little attenuated from base distad, the sides toward end more strongly rounding in to the apex. Inner side of first article straight, a little diverging from its mate at anterior end. Length of types from 5.5 to7.5 mm. A specimen 7.3 mm. long has tenth plate .82 mm. wide, antennae 2.4 mm. long, and anal leg about 2.2 mm. long. Locatiry.— Georgia (Atlanta!). GARIBIUS ALABAMAE, sp. nov. DiacGnosis.— Dorsal spines of anal legs, 1, 0, 2,0, 0. Dorsal spines of penult legs 1, 0, 2, 1, 1. Ventral spines of thirteenth legs, 0, 0, 2, 3,2. Dorsal spines of twelfth legs, 0, 0, 2, 2, 2; ventral, 0, 0, 1, 2, 1. Dorsal spines of eleventh legs, 0, 0, 0, 2, 2; ventral, 0, 0, 0, 2, 1. Dorsal spines of first legs 0, 0, 0, 1, 1; ventral, 0, 0,0, 1, 1. Third joint of first eleven pairs of legs or thereabouts unarmed either dor- sally or ventrally. Last pair of coxae laterally armed. Antennae as usual. Ocelli about 6 in two series. Prosternum 1.53 times wider than long. Distance between chitin- ous spots 2.4+ times width at median incision; 3.42 times dental line. Crest of anal legs of male broader and more rounded than in related species; bearing numerous hairs of the usual type. Length of type 5 mm. Body 7.6-+ times longer than the width of the tenth plate. Head considerably longer than wide. Dorsum anteriorly greyish yellow, becoming clearer yellow and light orange caudad. Head light orange. Description.— Above the body is greyish yellow, becoming clearer yellow and light orange caudad. Head light orange. Anten- nae yellowish. Prosternum pale orange or yellow. Venter dilute yellow, deeper yellow caudad. Legs very pale excepting the caudal ones which are bright yellow. Length about 7.6+ times the width of tenth dorsal plate. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to CHAMBERLIN: LITHOBIID GENERA. 79 each other as 31: 31+: 31: 34: 34: 33, the body being thus not strongly attenuated. Head elongate, being considerably longer than wide (34:31). Hairs of moderate length, sparse. Antennae short; moderately attenuated. Articles mostly very short, decreasing distad as usual; the ultimate moderately long. Hairs as usual in the genus. Eyes composed of about 6 ocelli in two series: — thus, 1+3,2. Single ocellus about equal in size to, or a little smaller than, the first one of the upper series. Seriate ocelli decreasing cephalad and ventrad; all pale. Prosternum 1.53 times wider than long. Distance between chitin- ous spots 2.4+ times width at level of bottom of mesal incision; 3.42 times as great as the dental line. Teeth small and acute; inter- val between each pair rounded at bottom; inner tooth somewhat larger than the outer and the line of apices straight. Mesal incision rather wide and shallow, sides concave but meeting at an acute angle at bottom. First dorsal plate widest a little back of the anterior end, the sides from there caudad at first slightly and then rather strongly converging. Posterior corners of ninth, eleventh, and thirteenth dorsal plates all straight on caudal side or a little rounded. O2000. Nalhy: 0, 10,0; 1@)1 2). Spines of first legs, Sareea nate of the second, 5°59: a a23-of the third to ce oo 0. 0, 0, 2,2. of the twelfth, 20-222: of 0, 0, 0, 2, 1 O Ov Is 20a the thirteenth, 4s 2 5'; of the penult, ip claws 2 (3); of Owns ‘i o 70,1, 3; 1,07 Gonopods of male well exposed, convexly rounded, wart-like; bearing a single moderately long bristle. Anal legs of male short, considerably inflated; fifth joint bearing at its distal end a low eminence which is relatively broader and more rounded than in related species, this lobe bearing over its surface rather numerous hairs of the usual character. Penult legs also in- flated, but without any special lobes. Length of type 5 mm.; width of tenth plate .65 mm.; length of antenna 1.6 mm.; anal leg of same length as antenna or nearly so. Locauity.— Alabama (Morgan!). A single male was secured. the anal eed single. Last pair of coxae laterally armed. 80 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. GARIBIUS BRANNERI (Bollman). Inthobius brannert Bollman, Ann. N. Y. acad. sci., 1888, 4, p. 107. Descriprion.— The original description, rearranged, is as follows:— Light chestnut-brown or orange; head and antennae scarcely darker; feet orange. Slender, smooth, very sparsely pilose. Head rounded triangular, narrowest before. Antennae short; joints 20, short. Ocelli 6-8, arranged in four or five rows. Prosternal teeth 4, small. [Posterior angles of none of the dorsal plates produced]. Coxal pores 2, 3, 3, 2-3, 4, 4, 3, small and round. First pair of feet armed with 0, 2, 1 spines; penultimate with 1, 3, Ll, 2,90; last wath 1o,3, 50: In the male the fifth joint of the anal leg is produced into a short pilose lobe, and is depressed. Claw of the female genitalia short, wide, bi- or tri-partite; spines slender, subequal, outer strongly toothed. Length of body 5-10 mm. Type Locariry.— Tennessee (Knoxville!). Locaitres.— Tennessee (Knoxville! Beaver Creek; Mossy Creek). TIDABIUS, gen. nov. Antennae short; composed of from 25 to 35 articles, but of mostly near 32. Eyes composed mostly of from 9 to 14 ocelli which are arranged usually in three series; single ocellus largest. Prosternum from 1.55 to 1.65 times wider than long, never less. Teeth 2+2; line of apices somewhat recurved. Spine bristle-like. Median incision v-shaped, being acute at bottom, or very rarely more or less rounded. Body always considerably attenuated from the tenth plate cephalad with the first plate always and mostly considerably narrower than the third. Posterior angles of none of the dorsal plates produced; or, often, with thirteenth or with eleventh and thirteenth showing a very weak production of these angles. CHAMBERLIN: LITHOBIID GENERA. 81 Coxal pores small and circular, uniseriate. Gonopods of female tripartite with lobes short, the lateral ones equal and with their apices usually in nearly same transverse line. Basal spines 2+2 or, rarely, 3+3; proximal portion of spine relatively broad with sides parallel or diverging to where the acutely acuminate terminal portion begins more or less abruptly (Plate 4, fig. 5). Posterior coxae always wholly unarmed. Anal legs with the dorsal spines uniformly 0, 0, 2, 0, 0; ventral spines, 0, 1, 3, 1, 0 or, in some, 0, 1, 3, 2, 0; claws always 2 or 3. Penult legs with dorsal spines OOo I Onor, rarchy,010,03,9l el ventralispines, 01,13, 3,1, to 0, 1, 2, 3, 1; claws 2 or 3. Third joint of first 7 to 9 pairs of legs mostly unarmed, rarely armed. Tibia of none of the legs with more than one ventral spine. All tarsi biarticulate. Posterior legs in both sexes short and moderately inflated, longi- tudinally more or less suleate dorsally; in male presenting no special modifications, or with legs but little more crassate. Length between 5.5 and 10 mm. Typr.— 7. tivius Chamberlin. Distrisution.— The known species are distributed across the continent through the Northern United States from Washington and Oregon to the Atlantic and from there into the Southeastern States. These small and wide-spread lithobiids never exceed 10 mm. in length. In most of the species the head varies from yellowish to ferruginous and is distinctly lighter than the dorsum. The venter commonly is suffused anteriorly with violaceous. The posterior legs are typically more brightly colored distad. These forms occur com- monly under leaves and sticks of willows and other trees and shrubs along streams, under bark of logs, among stones at the water’s edge and in other similar places. Where found they usually occur in considerable numbers. The species to one examining them at first commonly seem very close in general structure and appearance and are to be discriminated only upon careful study and comparison. The following key will aid in separating average individuals. Key to species of Tidabius. a. Ventral spines of anal legs 0, 1, 3, 2,0 or 0, 1, 3, 3, 0. b. Third article of all legs, or of all excepting the first two pairs, ventrally armed. T. poaphilus, sp. nov. bb. Third article of at least the first five pairs of legs unarmed ventrally. 82 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. c. Dorsal spines of penult legs 0, 0, 3, 1, 1. T. anderis, sp. nov. ce. Dorsal spines of penult legs 0, 0, 3, 1, 0. d. Anal leg with 3 distinct claws; basal spines of 2 gonopods 3+3. T. bonvillensis Chamberlin. dd. Anal leg with but 2 distinct claws; basal spines of 9 gonopods 2+2. e. Body 8 times, or but little more, as long as width of tenth plate. T’. suitus, Chamberlin. ee. Body 9 or more times as long as width of tenth plate. 1’. nasintus, sp. nov. aa. Ventral spines of anal legs 0, 1, 3, 1, 0. b. Length of body of adults less than 8 mm.; mostly from 7.25 to 8.25 times longer than width of tenth plate. T. tiwius Chamberlin. bb. Length of body of adults 8 mm. or more; 8.5 or more times longer than width of tenth plate. c. Ventral spines of eleventh legs 0, 0, 1,3, 1; of the thirteenth, OF 2,55, lore; Oneas, 1: Body 9 to 9.5 times longer than width 10th plate. T. opiphilus, sp. nov. ce. Ventral spines of eleventh legs 0, 0, 2,3, 1; of the thirteenth, Op lyeares 1 d. Dorsum brown; head brown of ferruginous caste; body about 8.5 times longer than width of 10th plate. T. pallidus alabamensis, subsp. nov. dd. Dorsum yellow; head light orange; body about 9 times longer than width of 10th plate. 7. pallidus, sp. nov. TIDABIUS NASINTUS, sp. nov. D1aGnosis.— Ventral spines of anal legs 0, 1,3, 2,0. Dorsal spines of penult legs, 0, 0,3, 1,0. Third joint of eleventh to fourteenth legs with three dorsal spines. Ventral spines of tenth and eleventh legs 0,0, 1,2,1. Ventral spines of first legs, 0, 0, 0, 1, 1. Articles of antennae (in type) 30. Ocelli 11 in three series. Distance between chitinous spots of prosternum 2.58 times width at median incision; 3.75—4 times the dental line. Head slightly wider than long. Coxal pores 2, 3, 3, 3. CHAMBERLIN: LITHOBIID GENERA. 83 Slender, about 9.3 times longer than the width of tenth plate. First dorsal plate only 13 times wider than long. Outer of basal spines on each side longer than the inner with which it stands at an angle; spines broad at base with sides slightly diverg- ing to middle where acuminate portion begins. Dorsum light brown, the caudal plates darker. Head darker brown or dilute chestnut with frontal region paler. Description.— Above light brown with the caudal plates darker. Head darker brown or dilute chestnut, the frontal region paler. Antennae brown proximally, light at tips. Prosternum brown; prehensorial feet yellowish. Venter pale with the caudal plates darker, brownish. Legs like the corresponding plates of venter, the posterior pairs there being darker, brownish, but brighter distad as usual. Body slender, being in the type 9.3 times longer than the width of the tenth plate. The widths of head and of the first, third, eighth, tenth, and twelfth plates to each other approximately as 41:35:38: 43:45: 43. Head subcordate. Slightly wider than long (ratio about 41: 40). Not much varying in width between caudal edge of eyes and the marginal breaks. Antennae short. Composed of thirty articles which distad of the second or third become very short and closely arranged. Hairs short extending in part distoectad and in part proximoectad as usual. Eye, in the type, composed of eleven ocelli arranged in three series: — thus, 1+4,3,3. Single ocellus largest. Ocelli of upper series larger and paler than others which decrease in size ventrad. Prosternum, in type, 1.58— times wider than long. Distance between chitinous spots 2.58 times width at level of bottom of median incision; 3.87 times the dental line. Median incision rather small, v-shaped. Teeth small, acute, with sides nearly straight; inner tooth a little larger than the outer but the line of apices clearly re- curved. Distance from tip to tip of mesal teeth about equal to or a little less than that from tip to tip of the two of each lateral pairs. Lateral margin of anterior portion convex anteriorly and concave posteriorly, where it bends ectad at end as usual. First dorsal plate 12 times wider than long; widest a little back of the anterior end where the sides are convex, the sides then moderately converging caudad to the rounded corners. Caudal margin of the ninth plate straight or nearly so; angles of eleventh and thirteenth very slightly produced, the process rounded; posterior corners of other minor plates more or less obliquely excised. 84 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Com pores circular; small, decreasing in size proximad on each coxa; 2,3; 3,0. Spines of first legs, OOO of the second, See iar eat of the third and fourth, OOS: 5 i; of the fifth, Eee of the seventh and eighth, ? aa 5 2; of the ninth, 3: ih 4 oO °: OF ? 5 25 of the tenth, 0.0.2.2, of the eleventh, 5-0 os 3: 5 4: of the twelfth, ae ee of the thirteenth, eee of the fourteenth, 0° Say : °; 0, 0, 2, 0, O of the anal, 0-13-39? With two claws. None of the coxae at all armed. Anal and penult legs of the female short, weakly inflated; especially the fourth joint longitudinally suleate above. Claw of female gonopods short and relatively wide; tripartite; the lobes short but distinct, the median longer than the lateral which are equal in size and at the same level. Outer of the two basal spines of each pair with which its wider surface stands at an angle as usual; spines broad at base with sides slightly diverging to the middle and then converging to a very acute apex, the sides of the acuminate por- tion being somewhat incurved. Inner side of first article of gonopods slightly curving ectad at proximal end and then a little diverging from its fellow. Length of type 8 mm.; width of tenth plate, .86 mm.; length of antennae, 2.3 mm.; of anal leg, 2.1 mm. Locatity.— Mississippi (Jackson!). Tipasius suitus (Chamberlin). Lithobius cantabrigensis var. suitus Chamberlin, Ann. Ent. soe. America, 1911, 4, p. 41. Diacnosis.— Ventral spines of anal legs, 0, 1, 3, 2, 0. Dorsal spines of penult legs 0, 0, 3, 1, 0 or 0, 0, 2, 1,0. Third joint of tenth to fourteenth legs inclusive armed with three dorsal spines. Ventral spines of tenth and eleventh legs, 0, 0, 1, 2,1. Ventral spines of first legs, 0, 0,0, 1, 1 or 0, 0,0, 2,1. None of the coxae armed. Articles of antennae 29-32. Ocelli 8-11 in 2 or 3 series. Distance between chitinous spots of prosternum 2-2.25 times width at median incision; 3.75-4 times as great as the dental line. Median incision rather large, widely rounded at bottom. CHAMBERLIN: LITHOBIID GENERA. 85 Coxal pores 2, 3, 3, 2-3, 4, 4, 3. Head subcordate, wider than long. Body moderate; about eight times longer than the width of the tenth plate. Median lobe of claw of female gonopods but little longer than the lateral. Basal spines very broad proximally, sides parallel or slightly diverging to about the middle where the acuminate portion begins. Dorsum brown, with posterior segments darker and more reddish as are also sometimes the first ones. Head brown of dilute ferrugin- ous tinge, especially anteriorly. DescrieTion.— Brown above, with the posterior segments con- siderably darker and more reddish; some of the anterior plates may also be darker, especially along the caudal borders, these also often showing a violaceous tinge. Head brown of dilute ferruginous tinge, especially anteriorly. Antennae brown of purplish tinge, distally lighter, yellow or yellowish of dilute ferruginous tinge. Prosternum brown, the prehensorial feet lighter. Venter very pale, the anterior plates showing a distinct violaceous or purplish tinge and the caudal plate much deeper in color and more or less of reddish tinge. Legs like the corresponding plates of venter; the posterior pairs darker brown with the distal joints light. Body moderate, being on an average 8+ times longer than wide. Widths of head and of first, third, eighth, tenth, and twelfth plates to exci ornertas 29.9): 22: 22:20: 27 320. Head subcordate; wider than long in about ratio 25.5: 24. Hairs short, sparse. Form and markings as usual. Antennae short, reaching the fifth segment composed of from 29 to 32, mostly very short, articles. Hairs moderate, straight, in part extending distoectad and in part almost directly transversely or somewhat caudoectad. Eyes each composed mostly of about 8 or 9 to 11 ocelli in two or three series: — thus, 1+4, 4; 1+3, 3, 1; 1+4, 4,2. Single ocellus considerably the largest. Seriate ocelli decreasing in size cephalad and ventrad. Prosternum 1.58 times wider than long. Distance between chi- tinous spots 2 to 2.25 times greater than width at level of bottom of mesal incision; 3.75 to 4 times as great as the dental line. Teeth small; subacute; the interval between each pair concave; line of apices clearly recurved. Mesal incision rather large, sides but little converging caudad, the bottom being rather widely rounded. Spine slender and acuminate; inserted a little ectad of outer tooth and 86 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. extending somewhat mesad of directly cephalad. Lateral margin ectad of spine, convex, more caudad becoming concave and bending ectad. Coxal pores small and circular; 2, 3, 3, 2 to 3, 4, 4, 3. . 0.0 silts / ofaeti Na. Spines of first legs, 5-5: o 7 + or 5 o c 5 i; of the second to fifth inclusive, a a 1; of the sixth and seventh, ? SOD of the eighth and ninth, 0-0-2 3-?; of the tenth and eleventh, §-9-4'5-73 of the twelfth, Be SB a the thirteenth, ee of the penult, % - * a a r fy °. ay == +, claws 2 2 (or with obsolete third one); 0, O of the anal, 2 os claws 2 (in regenerated legs may be but 1). None of the coxae at all armed. Anal legs of male short and a little inflated; the third, fourth and fifth, but most distinctly the fourth, joints longitudinally suleate on dorsal side. Penult legs similar but a little more slender. Claw of the female gonopods short and relatively wide; tripartite; the median lobe but little longer than the two lateral, the distal margin of the latter almost at right angles to the longitudinal axis of claw. Basal spines very broad at base as usual, the sides parallel or a little diverging to about middle of length, then converging to an acute point. Inner side of first article nearly straight, being a little convex distally and slightly concave proximally, nearly parallel to that of its opposite gonopod. Length 7-9 mm. A female 8 mm. long has the tenth plate .98 mm. wide; antenna 4 mm. long; and the anal leg 3.8-4 mm. long. Praematurus.— Coloration similar to that of adult but paler throughout; light brown of a pale violaceous caste, with the four- teenth plate darker and the fifteenth lighter, the latter being yellowish as in the adult. Head light orange. Antennae brown, light at tips. Prosternum yellowish. Venter pale, the first plates more strongly tinged with violaceous, and the last one more strongly pigmented, yellowish or light orange. Legs pale, suffused with very dilute violaceous, the last pairs light yellow excepting proximally. Antennae composed of twenty-five articles of which those beyond the third or fourth are very short and compactly arranged and those near the middle of length the shortest. Hairs straight, of moderate length, not dense. Eyes composed of seven or eight ocelli arranged in two series: — thus, 1+4, 3; 1+4,2. Ocelli small and pale, the single one largest and the most anterior ones smallest. CHAMBERLIN: LITHOBIID GENERA. 87 Prosternum is in adult or nearly so. Coxal pores very small; 2, 3, 3, 3. Spines of first legs, 5 o, a0 a 7 -+; of the second, a: ao “ x } to? : a +: 5 of the third to sixth, Ce +; of the sev re a: rs 3 1; of the eighth, 2-05-34 oF 675: +24; of the ninth, §-$45:+75 of the tenth and eleventh, ° a oO ‘ - * or . a ao ; of the twelfth, OOo oot or $54 B z =a i of the thirteenth, > Bo 3 z t; of the penult, S . 5 o; of the anal, 5 = aa = 2. Claw of feinal gonopods short and relatively wide, with the three divisions acute and the outermost somewhat smallest. Spines 2+2; the outer one of each pair of nearly adult form, being broad or some- what clavate proximally and acuminate from near the middle of length, the apex very narrowly rounded, the inner spine about half the size of the outer one, acuminate from very base. Length cir. 5.25 mm. Length of antennae 2 mm.; of anal legs, cir. 2.2 mm. (Haddonfield, N. J., Oct. 27, 1912). Immaturus.— Still paler than the prematurus; of a violaceous tinge throughout; the last dorsal slate yellow. Head and prosternum yellow. Anal and penult legs with tibiae and tarsi yellow, the more proximal joints having the pale violaceous tinge of the rest of the legs. Antennae very short. Composed of nineteen articles which distad of the third are very short. Hairs about as in the prematurus. Eyes composed of about six ocelli arranged in two series: — thus, 1+3,2. The most caudal ocellus of the dorsal series largest, the single one next, with the anterior one and those of the lower series much smaller. Prosternum essentially as in older stages. Coxal pores very small and pale; 2, 2, 2, 2. Spines of first to third pairs of legs, 2-°: oul nea: 00-00-17 Of the fourth to sixth pairs, §-0-0-4-1; of the seventh to ninth pairs, $9944) of the tenth and eleventh pairs, a 7 4 = *; of the twelfth, oe L x 1; of the thirteenth, e a 5 + or ee of the penult, asl : of the anal, STO Gonopods of female very short and pale; the three articles present, but the division between the second and third not yet very distinct. Claw indicated as a mere spinous point, the article beneath it distally 88 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. well rounded and bearing also a single bristle. Second article like- wise bearing a single bristle. Proximal article bearing three bristles in a transverse row. Spines 1+1, small and pale; acutely acuminate from the very base. Length cir. 4.2 mm. (Haddonfield, N. J., Oct. 27, 1912). Type Locatiry.— North Carolina (Hot Springs). Locaities.— North Carolina (Hot Springs!); Alabama (Birming- ham!); New Jersey (Haddonfield!). TIDABIUS POAPHILUS, Sp. nov. DiaGnosts.— Ventral spines of anal legs 0, 1, 3, 2,0. Dorsal spines of penult legs, 0, 0, 3, 1, 0. Third joint of tenth to fourteenth legs bearing 3 dorsal spines. Ventral spines of tenth and eleventh legs, 0,0, 2,3, 1. Ventral spines of the first legs, 0, 0, 1, 1, 1, or, occasion- ally, 0,0, 0,1, 1. None of the coxae armed. Articles of antennae 32. Ocelli 9-12 in three series. Distance between chitinous spots about 2.25 times the width at median incision; 4 times the dental line. Coxal pores 3, 4, 4, 3. Head wider than long. Body moderately robust, 7.5-7.75 times longer than the width of the tenth plate. Claw of female gonopods as typical in the genus. Spines broad proximally, the sides of this portion parallel or a little diverging; acuminate portion more than half the total length. Dorsum brown or greyish brown, suffused with dark purplish; the first and last several plates darker. Head brown to orange-brown, with the frontal region lightest. Description.— Above brown or greyish brown, commonly suffused with dark purple; the first and the last several plates darkest. Head brown or orange-brown, the frontal region lightest. Antennae light brown, lighter at tips, often tinged with purplish. Prosternum and prehensors brown. Venter greyish or dilute greyish brown, the anterior plates typically tinged with purple and the caudal plates darker, brown. Legs like the corresponding plates of venter, the first commonly with a purplish tinge and the caudal pairs brownish proxi- mally, brighter, yellowish, distally. CHAMBERLIN: LITHOBIID GENERA. 89 Body moderate or rather robust; the type 7.65 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 23: 21: 23: 26: 27: 26. Head wider than long in ratio 50:47.5. Caudal margin slightly and widely excurved. Sides between eyes and marginal interruptions very little excurved, subparallel. Hairs long, sparse. Antennae of the usual length. Composed of 32 articles which beyond the second are very short and closely united, submoniliform. Hairs of moderate length and short; not very dense, sparser proximad; extending in various directions as usual. Eyes each composed of 9-11 ocelli arranged in three, or occasionally, in but two, series: —e. g., 1+4, 3,3; 1+1,4,3; 1+5,3. The lowest series almost on ventral side of head. Single ocellus largest. Of the seriate ocelli the first or most caudal of the uppermost series is largest, the first of the second series also large; others decreasing cephalad and ventrad as usual. Prosternum 1.64 times wider than long. Distance between chitin- ous spots 2.25 times width at level of bottom of median incision; 4 times as great as the length of the dental line. Median incision mod- erately large, sides gently converging caudad; rather widely rounded at bottom. Teeth small, acute, equal; line of apices recurved. Tubercle of spine contiguous with base of outer tooth as usual; spine rather stout at base, running to a fine point with sides straight through- out length or nearly so, extending directly cephalad or nearly so, as in 7’. anderis. Lateral margins of anterior portions sloping directly caudoectad immediately from base of spine; the margins strongly incurved with caudal portion almost transverse. (Plate 5, fig. 3). First dorsal plate with sides diverging cephalad to about the middle and then parallel or nearly so. 1.8-+ times wider than long. Caudal angles of thirteenth plate a little produced as usual; angles of eleventh rectangular or showing a slight tendency to production; other minor plates with caudal margins bending a little forward at ends about the rounded corners. All plates finely but distinctly roughened with tubercular and rugose elevations. Coxal pores small, circular; 3, 4, 4, 3; on each coxa decreasing in size proximad with the most proximal often very small. Spines of first legs, 9-0.1:1:1 oy 9.0.1,1,1 of the second, Ono QnONO; ds 1? 0, 0, 2, 2, 1 OMOMZ arm . < < Os Opes 2p al Gaon oan anes of the third to fifth inclusive, Wa ae ed OROR2 22s - 7 0, 0, 2, 2, 2 of the sixth, ie a of the seventh and eighth, Or 0nis oy? OF on 90 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. one side this and on the other, 0-0-*-2-2; of the ninth, 0-92-22, or 0-02.24; of the tenth and eleventh, pos: 2:2; of the twelfth, OOS) bade . OvONS. deel» 0,053, 1.0 joe of the thirteenth, Fee: of the penult, pee 0, 0, 2, 0, O tarwnncoaae claws 2. None of the coxae at all claws 3; of the anal armed. Posterior legs but little, uniformly inflated. Third, fourth, and fifth articles longitudinally furrowed on dorsal side as usual. Not differing in male and female. Gonopods of the male small and rounded; bearing 1 or 2 bristles. Claw of female gonopods short and but slightly curved; distinctly tripartite in the usual way; median lobe longest, obtusely rounded, less acuminate than in anderis; lateral lobes with distal margins nearly at right angles to axis of claw. Basal spines broad at base as in related species; the sides proximally parallel or a little diverging distad; acuminate portion long, more than half the total length. Mesal margin of first joint nearly straight, a little diverging from its fellow cephalad. Sternite mesally a little incurved as in related spe- cies and without a distinctly chitinized process. The & type is 7.5 mm. long; width of tenth dorsal plate .98 mm.; length of antenna 2.5 mm.; of anal leg, 3 mm. Type Locatiry.— Nebraska (Fremont). Locauities.— Nebraska (Fremont! Omaha!). TIDABIUS ANDERIS, sp. Nov. Dracnosis.— Ventral spines of anal legs 0, 1, 3, 2,0. Dorsal spines of penult legs, 0, 0, 3, 1, 1. Third joint of twelfth to fourteenth pairs of legs inclusive with three dorsal spines. Ventral spines of tenth and eleventh legs, 0, 0, 1, 2, 1. Ventral spines of first legs 0, 0, 0, 1, 1. None of the coxae armed. Articles of antennae (in type) 31. Ocelli 8 in two series. Distance between chitinous spots 4 times the dental line; 2.46 times width at median incision. Coxal pores 3, 4, 4, 3. Claw of female gonopods as typical in the genus. Spines broad at base as in 7’. tivius but the acuminate apical portion embracing con- siderably more than half the total length. Head longer than wide. CHAMBERLIN: LITHOBIID GENERA. 91 Body robust, being only about 7.4 times longer than width of tenth plate. Dorsum testaceous, usually of dilute olivaceous or, sometimes, of somewhat purplish tinge; first and last plates darker as usual. Head yellowish orange to light ferruginous. DescripTion.— Above testaceous, in some of dilute olivaceous tinge, in others more or less purplish; the first plate and the most caudal ones often darker and some of the plates with their caudal borders crossed by a transverse dark stripe. Head yellowish orange to light ferruginous with a dark area on caudal portion along margin and extending forward along median line part way to the suture. Antennae brownish proximally, becoming yellow distad. Prosternum and prehensorial feet yellow to brown, the prosternum usually dusky over a yellow background. Venter pale, dilute testaceous or yellow- ish, the caudal plates darker brown. Legs like the contiguous plates of venter excepting that the posterior pairs beyond the proximal portion are bright yellow. Body robust; in Q type, 7.4 times longer than width of tenth plate. Widths of head and of the first, third, eighth, tenth, and twelfth plates to each other as 25: 23: 24: 28: 26.5. Head longer than wide in ratio 50: 47; a little more elongate than in 7’. tivius with the anterior portion somewhat narrower; not much varying in width between the marginal breaks and eyes; caudal margin straight. Antennae short or very short. Composed of 31 articles which are very short, especially distad, and are closely aggregated. Bristles moderate in length, subdense on distal articles, becoming considerably sparser proximad. Eyes in type each composed of about 8 ocelli arranged in two series: — thus, 1+4, 3. Ocelli large and distinct; single ocellus somewhat obliquely elliptic. Prosternum 1.64 times wider than long. Distance between chitin- ous spots 2.46 times width at level of bottom of median incision; 4 times as great as length of the dental line. Teeth small and acute, pale; line of apices recurved. Tubercle of spine contiguous with base of outer tooth; spine manifestly stouter than the hairs, acutely acuminate as usual, extending almost directly cephalad and thus not parallel with outer edge of tooth as is more typically the case in tivius. Median incision rather narrow, the sides nearly parallel, but little converging caudad excepting toward bottom, the latter some- what rounded. 92 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. First plate moderately narrowed caudad as usual, 1.9 times wider than long. Minor plates as in tivius or nearly so; in the anterior ones the caudal margin moderately bending forward to corners at ends; in the eleventh the posterior corners are nearly rectangular; in the thirteenth they are a little extended caudad. Plates all dis- tinctly finely roughened. Coxal pores small, circular, the most proximal one on some coxae minute; 3, 4, 4, 3. COMO Ee a 050,008, 1° ONOR2 2.1” ; . : Wot Ou Ove2 nana 0-070-2-1; Of the seventh to ninth inclusive, 9-9-9-5°7; of the tenth and eleventh, 0-?-*3-?; of the twelfth and thirteenth, pS at |, 0.0, 2, 0, 0 OLO Nei 9 . i aaa Nae aa claws 2. No spines detected on any coxae. Anal and penult legs in both sexes short; a little thickened. Claw of female gonopods short; but little curved; median lobe longest, distally more or less narrowly rounded; lateral teeth with their distal edges in line and at right angles to the axis of the claw or nearly so. Spines broad at base as in tivius, but the acuminate por- tion longer, embracing considerably more than half the total length, apically acutely rounded with edge often showing a few minute denticulations. Vental edge of mesal side of first joint well chitinized; nearly straight excepting proximally where it bends a little ectad. The female type is 7.4 mm. long; width of tenth plate 1 mm.; length of antenna 2.25 mm. Locautity.— Washington (Pullman! W.M. Mann, 1908). Spines of first legs, of the second to sixth inclusive, of the penul of the ana TIDABIUS PALLIDUS, sp. nov. D1aGnosis.— Ventral spines of anal legs 0, 1,3, 1,0. Dorsal spines of penult legs 0, 0, 3, 1, 0. Third joint of tenth to fourteenth legs with three dorsal spines. Ventral spines of tenth and eleventh legs, 0, 0, 2, 3, 1. Ventral spines of first legs, 0, 0, 0, 1, 1. None of the coxae armed. Articles of antennae, in type, 27. Ocelli 9 or 10 in three series. Distance between chitinous spots of prosternum 3.5 times the dental. line; only 2.1 times width at median incision. Head cordate, very little wider than long. CHAMBERLIN: LITHOBIID GENERA. 93 Coxal pores 3, 4, 4, 3. Body moderately slender, about nine times longer than the width of the tenth plate. Spines of Q gonopods large and broad, the sides parallel to near middle from where the acutely acuminate portion begins. Dorsum yellow. Head light orange. DescripTion.— Yellow above with the caudal plates light orange. Head light orange. Antennae pale. Prosternum yellow of pale orange tinge. Venter very pale with caudal plates orange. Legs pale, the posterior pairs more densely pigmented, brighter in color. Body slender, being 9 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 45: 40: 43: 51: 52: 48. Head cordate; slightly wider than long (ratio about 45.5: 44). Hair short, sparse. Antennae short and moderately attenuated as usual. In type composed of 27 articles which beyond the second are very short, shorter ones occurring in pairs at intervals among somewhat longer ones. Hairs as usual. Eyes in type each composed of 9 or 10 ocelli arranged in three series: — thus, 1+4, 3, 1; 1+4, 3, 2. Single ocellus largest; first ocellus of top row also large and the others of this series also larger than the lower ones which decrease in size ventrad. Prosternum 1.58 times wider than long. Distance between chi- tinous spots 2.1 times the width at level of bottom of median incision; 3.5 times the dental line. Median incision wide; sides distally concave, meeting at bottom in a wide angle; distance from tip to tip of mesal teeth clearly greater than between tips of teeth of each pair. Teeth small and acute; line of apices slightly recurved. Anterior lateral margin at first strictly transverse ectad of each outer tooth for a short distance, this portion bearing the spine, the caudally sloping portion at first conspicuously convex, becoming caudally concave and at caudal end bending directly ectad. Spine conspicuously stout at base, slender and fine pointed distally. First dorsal plate 1.78 times wider than long, shaped as in 7. nasin- tus. Caudal corners of thirteenth plate a little produced as usual; those of the eleventh caudally straight or somewhat excised; those of the ninth mostly a little excised or with caudal edge bent forward. Coxal pores of the usual size and form; 38, 4, 4, 3. Spines of first legs, Oe of the second to seventh inclusive, POs 2) T. : CNC HE AP I I Fatmuacane of the ninth, aaa of the tenth and eleventh, 94 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 0,0, 3,2, 2. 0, 3.1, - OnOesNIy a, ae ae of the twelfth, 2 CREE of the thirteenth, Soe of the penult, eee claws 2; of the anal, 9-0 2.0.0 Claws 2 1-0, aaa c0? None of the coxae armed. Posterior legs as usual. Claw of female gonopods relatively short and wide as usual; lobes short but distinct, the median longest, the two lateral subequal and at same level with their caudal edges almost transverse. Spines’ large, broad, with sides from base to about middle parallel, spines distad from middle acutely acuminate. Inner side of first article diverging moderately from its fellow proximally. The female type is 8.7 mm. long; width of tenth plate .97 mm.; length of antenna 3 mm.; of anal leg, 2.7 mm. Locauity.— Mississippi (Jackson!). TIDABIUS PALLIDUS ALABAMENSIS, subsp. nov. Diacenosis.— Light brown, with the first and last plates darker. Head brown of dilute ferruginous tinge. Somewhat more robust than type of 7. pallidus, being but 8.5 times longer than width of the tenth plate. Spining of legs as in pallidus or essentially so. Coxal pores 3, 4, 4, 4. Head as in pallidus. Gonopods and spines nearly as in pallidus. Distance between chitinous spots 3.1 times the dental line. Description.— Dorsum light brown, the first two or three plates and, more especially, those of the caudal third distinctly darker. Head brown of pale ferruginous tinge. Antennae brown proximally, pale distally. Prosternum and prehensorial feet brown. Venter pale, the anterior plates suffused with violaceous, the posterior plates darker, brownish. Legs like adjoining plates of venter; the posterior pairs brown proximally and bright yellow or yellow of dilute orange tinge distally. Body 8.5 times longer than width of the tenth plate. Widths of head and of the first, third, eighth, tenth, and twelfth plates to each other as 42: 40: 42: 47: 50: 48. Head a little wider than long (near 42:41); widest immediately back of eyes, with the sides conspicuously converging from the breaks caudad. CHAMBERLIN: LITHOBIID GENERA. 95 Antennae in type broken off at tips so precise number of articles is not determinable. Eyes each composed of 12 ocelli in three series: — thus, 1+4, 4, 3; ocelli of the usual relative sizes and arrangement. Prosternum 1.55-+ times wider than long. Distance between chitinous spots about 2 times width at level of bottom of median in- cision; 3.1 times the dental line. Median incision wide, well rounded at bottom; distance from tip to tip of mesal teeth clearly greater than distance between tips of tenth of each pair. Teeth small and acute; line of apices almost straight, being only very slightly recurved. Sides of anterior portion convex anteriorly and bending ectad caudally as usual. Spine of usual form. First dorsal plate of the usual shape, slightly shorter relatively than in type of pallidus. Thirteenth plate with posterior angles weakly produced; other minor plates with caudal corners more or less ob- liquely excised. Coxal pores of usual size and shape; 3, 4, 4, 4. Spines of first legs, 5 ue : 2,1,1. of the second to fifth, 9% 2-2-1. Bp 8 rOedeat OG Oy aa of the sixth to eighth, “o. 0 2: aa > of the) ninth, & OO of the tenth, 220sa202) oy O0:8.2.2. of the eleventh, 0.0, 22). of) the O,On2 2n1 On0nt, 2) 1 Oe wae twelfth, SOS ED of the thirteenth, aE of the penult, G40R3i 1, , 0, 2, 0, 0 ra i Osirees: -°, claws 2; of the anal, 5 claws 2,long. None of Serr O the coxae armed. Claw of female gonopods short and relatively wide; lobes short, with the median only a little longest, the two lateral equal and at the same level. Spines broad proximally, the sides diverging from base to a little below middle, the spine then acutely long acuminate. Inner side of first article subparallel with that of its mate excepting proximally, where it moderately diverges from it. Length of female type 8 mm.; width of tenth plate .94— mm. Locauity.— Alabama (Selma!). While manifestly very close to pallidus, the type of alabamensis is strikingly different in color, being much more in appearance like nasintus. Otherwise it differs a little also in robustness, being a little stouter. The prosterna are very similar. As only the type of each form has been examined, the precise relationship of the two cannot at present be definitely determined; but it seems best to keep them apart tentatively and to range one as subspecies to the other. 96 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. TipaBius Trvrus (Chamberlin). Lithobius tivius Chamberlin, Ann. Ent. soc. America, 1909, 2, p. 188. DraGnosis.— Ventral spines of anal legs 0, 1,3, 1,0. Dorsal spines of penult legs, 0, 0, 3, 1, 0. Third joint of tenth to fourteenth legs armed dorsally with three spines. Ventral spines of eleventh legs, 0,0, 2, 2,1; ventral spines of first legs, 0,0, 0,0, 1. None of the coxae armed. Articles of antennae 25 to 31. Ocelli mostly 9 to 12 in three series. Distance between chitinous spots 3.8 to 4.25 times the dental line; 2.3 times width at median incision. Head equal in length and breadth or very nearly so. Body robust, from 7.25 to 8.25, or thereabouts, times longer than the width of the tenth plate. Coxal pores 2, 3, 4, 3-3, 4, 4, 3. Claw of gonopods as usual. Spines broad with sides diverging to about the middle where the acuminate distal portion begins. (Plate 4, fig. 5). Dorsum pale brown or of pearl-grey caste, with the caudal plates more or less of a ferruginous tinge. Head light ferruginous. Description.— Above pale brown faintly suffused with violaceous or sometimes of pearl-grey caste; the caudal plates somewhat ferru- ginous. Head light ferruginous, commonly darkened over median region of caudal portion. Antennae brown to somewhat ferruginous proximally, lighter distad. Prosternum paler than head, the prehen- sorial feet lighter. Venter very pale, suffused with violaceous; the caudal plates darker, more or less of ferruginous tinge. Legs also pale, the caudal pairs darker, brown, proximally, and brighter, yel- lowish, distad. Body robust, being about 7.25 to 8.25 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 40: 34: 37: 40: 48: 42. Head subcordate; equal in length and breadth or very nearly so. Hairs moderately long with shorter ones intermixed, sparse. Antennae short and moderately attenuated. Composed of 25 to 31 articles which distad of the second are very short. Bristles almost transverse not especially dense. (Plate 5, fig. 6). Eyes each consisting of 10 or 11 ocelli in the types: —e. g., 1+4, 4, 2; 1+4, 3,2. Single ocellus largest; others regular in form and arrange- ment. (Plate 5, fig 7). CHAMBERLIN: LITHOBIID GENERA. 97 Prosternum 1.58 times wider than long. Distance between chitin- ous spots 2.3 times width at level of bottom of median incision; 3.8 to 4.25 times the dental line. Median incision v-shaped, with sides rather strongly converging. Teeth small and acute; interval between the two on each side semicircular; line of apices clearly recurved. An- terior lateral margins rather strongly convex anteriorly, concave caudally. (Plate 5, fig. 2-3). First dorsal plate 1.8 times wider than long; widest a little behind the anterior end where the sides are convex; sides then nearly straight and moderately converging to the rounded caudal corners. Posterior angles of the elventh and thirteenth plates a little extended caudad, rounded, those of ninth straight or obliquely excised. Coxal pores small and circular as usual; 2, 3, 4, 3 — 3, 4, 4, 3. Spines of first legs, ?-$3:9-4 °f o-oo: t; of the second, 9-05-24 Or O-5¢G! i of the third to fifth, Ceres of the sixth, ¢-p-4-3+4 or 0-0:5-5-7; of the seventh to ninth, pshetate2 of “the tenth, °: 7 3: 5 2; of the eleventh, B 33 3 2 or Oops of the twelfth, 6.0. 3.1.1; of the thirteenth, $0334 to 9-7-3+3°45 of the penult, o-9-3-5-9; of the anal, §:9:3° 9" Claws 2. None of the coxae armed. Anal and penult legs (2) short; uniformly a little inflated. (Plate 5, fig. 4-5). Claw of gonopods of female moderate; relatively wide; but weakly curved; the median lobe longest and the lateral ones equal and at the same level as usual. Basal spines broad proximally; sides diverging a little from base to about middle where the acutely acuminate distal portion begins, the sides of the latter straight. Mesal side of first article moderately diverging in proximal half from its fellow. (Plate 4, fig. 5). Length 5.5-7 mm. A specimen 5.8 mm. in length has tenth plate .8-++ mm. wide; and antenna 2 mm. long. Pseudomaturus.— Freshly matured individuals seem to agree in nearly all features with the older ones; but the sixth legs more fre- quently bear but a single dorsal tibial spine instead of two and the eleventh legs more frequently bear a single ventral spine on the pre- femur instead of two. Thus the spining, as might be expected, more frequently remains in the condition prevalent in the praematurus stage. Praematurus.— Dilute brown, suffused more or less with violaceous, this being evident more especially on the anterior and posterior seg- 98 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ments both above and below. Head light ferruginous. Posterior legs as usual more pigmented than the others. Antennae composed of twenty-six or twenty-seven articles of nearly adult proportions. Ocelli six, in two series: — thus, 1+3, 2. The single ocellus and the first of the upper row of the same size or nearly so and much larger than the others, the most anterior smallest. Prosternum as in the adult. Coxal pores small and pale; 2, 3, 3, 2. : Ov0;(0,1, 1. 0, 0, 0, 1, 1 Spines of the first legs, 0 OF 070s? of the second pair, Gee OE 0, 0, 0, 2, 1. : osiys OMONOMOET | 0,10; Dia Gosac of the third to sixth, OnOmOrsiesTe of the seventh, cOMORGe ast of the eighth and ninth, Oot of the tenth and eleventh, OL0 Bene. oo ala, Oak ayes of the twelfth, 5 ew of the thirteenth, Se MO e2 Ibe 0, 0, 2, 0, 0 of the penult, 2 ae of Re aap Ss Claw of the female gonopods small and pale, the three lobes distinct but appearing more slender and acute than in the adult. Proximal article with about six bristles on its ventral surface. Spines 2+2; the inner one of each pair shorter and much more slender than the outer and acutely acuminate from the very base; in the outer spine also the demarcation between basal and acuminate portions less distinct than in the adult. (Canton, Miss.). Agenitalis I.— Very dilute yellowish, with the head as usual more densely pigmented. Antennae and legs pale; the last pairs of legs with more pigment, very pale yellowish. Antennae composed of from twenty to twenty-three articles. Sub- moniliform. The last article conspicuously long. Hairs straight; of moderate length; not dense. Ocelli four in one or two series: — thus, 1+3, or 1+2,1. Small and pale. Teeth of prosternum pale, their form and relations, as those of the ectal spines, as in the adult or nearly so. Coxal pores small and weak edged; 1, 1, 1, 1 or 1, 2, 2, 1, the more proximal one when two are present on a coxa being considerably smallest. Anal glands still evident but not very distinct. Spines of first and second pairs of legs, 0.0. OOF; of the third to O00, 0,2 . 0,10:.0,.1, t cs casenatoda ninth, ONO T Unie of the tenth to thirteenth, GROMOMIENE of the penult, ? as ic Bs of the anal, oOo" CHAMBERLIN: LITHOBIID GENERA. 99 (Sterling, Ill., July). Tyre Locauity.— Utah (Provo!). Loca.ities.— Utah (Provo! Salt Lake City!); Colorado (Colo- rado Springs! Ft. Collins!); Illinois (Sterling! Dwight! Peoria!); Kentucky (Lexington!); Tennessee (Johnson City!); Virginia (va- riety, Lynchburg! Balcony Falls!); New Jersey (Haddonfield!) ; Pennsylvania (Philadelphia! Ardmore!); Wisconsin (Janesville!) ; Massachusetts (Hough’s Neck!); Alabama (Birmingham!); Missis- sippi (Brookhaven! Canton!). TIDABIUS OPIPHILUS, sp. NOV. DraGnosis.— Ventral spines of anal legs 0, 1,3, 1,0. Dorsal spines of penult legs 0, 0, 3, 1, 0. Third joint of tenth to fourteenth pairs of legs with three dorsal spines. Ventral spines of eleventh legs, 0,0, 1,3, 1. Ventral spines of first legs, 0, 0, 0, 1, 1. Articles of antennae 27 to 32. Ocelli 9 to 12 in three or, rarely, in four longitudinal series. Head subcordate; wider than long. Distance between chitinous spots 3.7 times as great as the dental line; 2.3 times the width at median incision. Body 9 to 9.5 times longer than the width of the tenth plate. Claw of the female gonopods much as usual. Basal spines broad proximally with sides subparallel; acuminate portion about two thirds the total length. (Plate 5, fig. 9). Brown; first and last plates darker. Head dark brown of a dilute ferruginous tinge. Descrietion.— Dorsum brown; first and caudal plates darker; the median region pale. Head darker, brown of a dilute ferruginous tinge, lightest in front of suture. Antennae brown proximally, lighter, yellowish or pale ferruginous, distally. Prosternum brown, the prehensorial feet paler. Venter very pale, tinged with violaceous anteriorly and the caudal plates darker, burnt brown. Legs pale like the venter; the caudal pairs darker, brown, commonly brighter distally. Slender, moderately attenuated cephalad; 9 to 9.5 times longer than width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 48: 44: 45: 51: 52: 49. Head subcordate, being considerably narrowed cephalad; wider than long in about ratio 49:47. Hairs of moderate length, sparse. 100 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae short. Composed of from 27 to 32 articles which are short and very short. Hairs rather short, subdense. Eyes each composed of from 9 to 12 ocelli in three or four series: — CG e4, ay wila-4, 19) oni aasna.. Prosternum 1.56 times wider than long. Distance between chi- tinous spots 2.31 times width at level of bottom of median incision; 3.7 times the dental line. Median incision v-shaped; sides Subparaltel distad, concave and converging caudad. Teeth small and acute, _ the interval between the two of each pair well rounded at bottom, not at all angular. Spine a little ectad of tooth, straight and slender; extending directly cephalad. Anterior lateral margins convex distad then Bacomune concave and bending out almost directly ectad caudally. First dorsal plate widest a little back of anterior end where the sides are rounded, becoming there nearly straight and gently converging te the rounded caudal corners; 1.9 times wider than long. Poste- rior angles of eleventh and timecnth plates very weakly produced caudad; those of the ninth straight to more or less obliquely excised. Comal pores 3, 4, 4, 3-3, 3, 3, 3; often decreasing markedly in size proximal on each coxa, the most aacsimel then being unas Spines of first legs, 0° 11:1. of the second to fifth, 90 2-2-1, 0, O10iel? > 0,0, 0,2, 1 of the sixth, i‘ acs one 3: - of the seventh, 0 °: = ae or . o . 5 2; of the Rite o- 7 e 5 -2; of the ninth, 2 a 7 sa OS ae - 4 (noted on one side only, the other leg plese the first formula); of the one OO ea of the eleventh, oO? of the twelfth, 7 oe 3 ; of the thirteenth, o es et or, SET of the penult, oa oO; We a claws 2; of the anal, aa °, claws 2. None of the coxae armed. Anal legs short; uniformly but only a little thickened; longitudi- nally sulcate above, especially the fifth article. Penult legs similar, but a little more slender. Claw of female gonopods very short; bent; median lobe longest and the lateral subequal as usual. Sides rather broad with sides of proximal division subparallel; acuminate portion rather long, being mostly about two thirds of the total length, the apex acute or narrowly acutely rounded. (Plate 5, fig. 9). Mesal side of first article more or less convex, diverging from its fellow proximately. Length from 8 to 10 mm. A female 9 mm. long has tenth plate 1 mm. wide; antenna 2.8-3 mm. long; anal leg 3 mm. long. Loca.ity.— Wisconsin (Beloit!). CHAMBERLIN: LITHOBIID GENERA. 101 TIDABIUS BONVILLENSIS (Chamberlin). Iithobius bonvillensis Chamberlin, Ann. Ent. soc. America, 1909, 2, p. 189. DescriptTion.— The original description of the types is as follows: — Dorsum from light to blackish brown, the posterior border of most of the major plates usually lighter; head paler, brown or light brown, sometimes with a darker median area caudad of the frontal suture; antennae brown to blackish, yellowish or rufous distally; praesternum and feet concolorous with head; venter light brown to yellowish, the first and the last plates darkest; legs yellowish, the last few pairs commonly darker. Head a little wider than long (3: 2.9). Antennae short; the first two articles long, others short, closely crowded; articles 29 in number. Eyes each composed of 8-9 ocelli arranged in two series (1+4, 3; 1+5, 3); ocelli of moderate size. Prosternal teeth small, slightly darkened. Angles of none of the dorsal plates produced. Coxal pores small, round, the distal ones of each series largest as usual; 3, 4, 4, 4. Ventral spines of anal legs 0, 1, 3, 2, 0, claws 3. Ventral spines of penult legs, 0, 1,3, 3,1, claws3. Ventral spines of first legs 0, 0,0, 0, 1. None of the coxae at all armed. Gonopods of female with the claw tripartite as usual, the lobes short and rather blunt. Basal spines 3+3; decreasing in size from the most ectal to the most mesal on each side; each spine acuminate distad of middle as usual. Locatity.— Utah (Lake Point!). This species was found in May under stones along the foot-hills. SIGIBIUS, gen. nov. Dorsal spines of anal legs 0, 0, 1, 0,0; ventral, 0, 1,1, 1,0; claws 2. Dorsal spines of penult legs, 0, 0, 1, 0,0; ventral, 0, 1, 1, 1, 0, claws 2. Dorsal spines of thirteenth legs, 6, 0, 1, 0, 0; ventral, 0, 0, 1, 1, 1. Dorsal spines of twelfth legs, 0, 0, 0,0, 0; ventral, 0,0,0,1,1. Dorsal spines of eleventh legs, 0, 0, 0, 0, 1; ventral, 0, 0, 0, 1, 1. Dorsal spines of first legs, 0, 0, 0, 0, 1; ventral, 0, 0, 0, 0,1. None of the coxae at all armed. 102 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae short; articles (in type) 25. Eyes composed of few ocelli (6 or less in type) which are arranged in either one or two series; single ocellus decidedly largest as in Tidabius. Prosternal teeth 2+2; line of apices rather strongly recurved. Coxal pores circular; uniseriate. Posterior legs of male short; uniformly a little thickened; without any special lobes or processes. Claw of female gonopods in type nearly bipartite, the ectal lobe commonly obsolescent, then contracting with all known species of Tidabius. Basal spines moderately slender, acutely and evenly acuminate from very base to apex. Posterior angles of none of the dorsal plates at all produced. Small in size (max. length of type species cir. 7.5 mm.). Typr.— S. puritanus, sp. nov. DistrrpuTIoN.— New England States. Evidently this genus is closest to the preceding one, Tidabius. From this genus, however, it is clearly distinct in the spining of the legs; e.g. the dorsal spines of the anal legs being 0, 0, 1, 0, 0 whereas in all species of Tidabius the corresponding formula is 0, 0, 2, 0, 0; also the ventral spines of these legs are 0, 1, 1, 1, 0, whereas in the other genus the formula is 0, 1, 3, 1, 0 or, more rarely, 0, 1, 3, 2,0; and the contrast in spining of the penult legs is still greater and equally con- stant. The prosternum is similar to that of Tidabius though the line of apices of teeth is more strongly recurved than in any species of the latter genus. The basal spines of the female gonopods are conspicu- ously different in shape, being more slender and not showing any division into broader proximal portion, the sides of which are divergent or parallel, and an acuminate distal portion, so characteristic in all species of Tidabius, but attenuated uniformly from base to apex; ete. The ocelli are fewer in number and show a pronounced tendency to form but a single series. Tidabius and Sigibius may be at once separated from the other two genera dealt with in the present paper by the larger number of antennal articles and by their not having the posterior legs of males characterized by any special processes or lobes. SIGIBIUS PURITANUS, sp. nov. DraGnosts.— Dorsal spines of third to eleventh pairs of legs, 0, 0, 0, 0, 1; ventral 0, 0,0, 1, 1. Dorsal spines of first and second pairs, 0, 0, 0,0, 1; ventral, 0, 0, 0, 0, 1. 1 ’ CHAMBERLIN: LITHOBIID GENERA. 103 Ocelli 3 to 6 in one or two series. Prosternum near 1.57 times wider than long, Distance between chitinous spots 23 times the width at median incision; 3.86 times the dental line. Coxal pores 2, 3, 3, 3-3, 4, 4, 3. Claw of female gonopods wide bipartite or with obsolescent third tooth; the two main lobes short and rounded. Body on average about 7.78 times longer than width of tenth plate. Head a little longer than wide (42: 40 to 41: 40). First dorsal plate about 1.7 times wider than long. Brown to testaceous, often of dilute olivaceous caste; caudal dorsal plates‘:and commonly Ist one darker, reddish brown. Head chestnut, light ferruginous in front of suture. DescrreTion.— The dorsum is brown to testaceous, often of dilute olivaceous caste; caudal plates, and usually also the first one darker, reddish brown or chestnut, and one or more of the plates immediately caudad of the first one often with their posterior borders also of the darker color or the first plate also similarly darker only along its caudal border. Head chestnut in appearance, of a lighter ferruginous color in front of suture, behind which there is present a median black spot extending toward the caudal margin. Antennae mostly dark brown proximally, becoming lighter distad, the distal portion being mostly yellowish or sometimes of a rufous tinge. Prosternum dusky brown, the prehensorial feet lighter, rufous distally. Venter pale; the caudal plates darker, ferruginous to burnt brown; the anterior segments or entire venter suffused with purplish. Legs mostly pale, but the caudal pairs bright yellow or light orange excepting proxi- mally, where they are brown. Body strongly attenuated cephalad, about 7.8 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 39: 38: 42: 50: 50.5: 50 in a female type; in male as 39: 37.5: 39: 46: 47: 46, the female being thus pro- portionately the broader caudad. Head subcordate, mostly a little longer than wide (42: 40 to 41: 40). Antennae very short; articles mostly 25, very short; hairs moder- ately long, not very dense. Eyes each consisting of from 3 to 6 ocelli arranged in one or two series: —e. g., 1+2; 1+2,1; 1+3, 2. Single ocellus largest. Prosternum about 1.57 times wider than long. Distance between chitinous spots 23 times width at level of bottom of median incision; 3.86 times length of dental line. Median incision moderately wide, 104 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. sides straight and strongly converging. Teeth small and acute; the line of apices strongly recurving, the ectal tooth on each side being borne considerably farther forward than the mesal one. Spine in- serted at base of outer tooth, scarcely stouter at base than the hairs, becoming slender and finely pointed distad. First dorsal plate 1.7 times wider than long. Minor plates all with caudal corners obliquely somewhat excised; none with them at all produced. Coxal pores 2, 3, 3, 3-38, 4, 4, 3; increasing in size distad on each coxa. Spines of first and second pairs of legs, aE of third to eleventh, co ae of twelfth, teint of thirteenth, ?: ON0515.0, 08 Oe , 0, 1, 0, ¢ » Oi1o Claws 2; of the anal also, 4 5! of the penult claws 2. Posterior legs in male, short, very moderately and uniformly en- larged. Claw of female gonopods broad; the ectal lobe commonly almost or wholly obliterated, the other two lobes distinct, with the median usually considerably the larger, or sometimes the two lobes are nearly equal and bluntly rounded. Basal spines differing from those of species of Tidabius in being more slender and in being attenuated evenly from the very base to the acutely rounded apex, their being no distal acuminate portion sharply delimited from a basal. Inner edge of first article straight. Tyrese Locauity.— Massachusetts (Cambridge!). Locauitres.— Massachusetts (Cambridge! Arlington Heights! Forest Hills! Waverley! Mattapan! Blue Hills! Wellesley! Hough’s Neck!); Maine (Orono! Bangor! Hampden! Old Town!). Vari us ah >» " - ahha : ts Be ee hae | obi ane V “ety fy Tos Sled NATAL yet pe id day 1 U0] AH va “ey 4 ceVith relia Fei 1 oo mt 4 teapteal th 5 Pe st aR (co Sara CHAMBERLIN.— Lithobiid Genera. Fig. Fig. Fig. Fig. Fig. Fig. Fig. PLATE 1. NAMPABIUS VIRGINIENSIS Chamberlin. Prosternum. Anterior border of prosternum more highly enlarged, showing teeth, ectal spines and form of mesal incision characteristic of the genus. Penult leg of o& showing the characteristic process at distal end of tibia. Process of penult leg more highly enlarged. NAMPABIUS EMBIUS Chamberlin. Left eye, showing characteristic form and arrangement of ocelli (single ocellus at the right of fig.). NAMPABIUS FUNGIFEROPES Chamberlin. Penult leg. Process of penult leg more highly enlarged. Bull. Mus. Comp. Zool. Lithobiid Genera Plate 1 | iS thet sare YA bak | Cann Lage Seek Ase ‘ Ps ‘q ee ae rey ieee ee Led BEA My ; ¢ oe a fu ie i =~ 1 j . ‘ ch " f eu i - 1 ‘ay nr any 7 it i ; \ 4, i i * hee \ Pans e ‘ ‘ tet j CHAMBERLIN.— Lithobiid Genera. Fig. Fig. Fig. Fig. Fig. Fig. Fig. | OO PLATE 2. NAMPABIUS GEORGIANUS Chamberlin. Penult leg, caudal (mesal) aspect. Process of penult leg more highly enlarged. Anal leg ectal aspect. NAMPABIUS CAROLINENSIS Chamberlin. Penult leg, caudal aspect. Process of penult leg more highly enlarged. NAMPABIUS VIRGINIENSIS Chamberlin. (See also Plate 1). Left gonopod showing typical form of claw, basal spines, and proxi- mal article. NAMPABIUS EMBIUS Chamberlin. Antenna. Bull. Mus. Comp. Zodl. Lithobiid Genera Plate 2 CHAMBERLIN.— Lithobiid Genera. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. = PLATE 8. NAMPABIUS PARIENUS Chamberlin. Penult leg, caudal aspect. Process of penult leg more highly enlarged. NAMPABIUS TENNESSEENSIS Chamberlin. Penult leg. Process of penult leg more highly enlarged. NAMPABIUS INIMICUS Chamberlin. Penult leg, caudal aspect. Process of penult leg more highly enlarged. NAMPABIUS VIRGINIENSIS Chamberlin. (See Plates 1, 2). Head in outline, showing the more usual form in the genus. NAMPABIUS GEORGIANUS Chamberlin. (See also Plate 2). Head in outline. Bull. Mus. Comp. Zo0l. Lithobiid Genera Plate 3 : a entice ee he 2 ee TLL a - ay wks ee A \ Pt as 7 i iy ernk £) tibetan Yenyounn i eiet? A yi mt ey > é ans x ; BO ay i H o4@ Pah : ; "5, : r iy : 7 ~ r 1 ‘ee. i denad: webs it7 rao i poses): , + Gus pers heres } ta nn ison att nil i, swaqae tee rere ’ Bee vy , an elnetn po alt etl jn: ka orisiy Pesan dewey | in jl omer iy Lades oo Tui No terns bagel 2 gil: » JM . : sy art 18 is ey : | | wiodaanst? 5 anwatKD anitiaas) ~s silty « ‘ iid io e Taunt wile wal} Yo ‘tel uit wide fy vrictors Hitaadk = CHAMBERLIN.— Lithobiid Genera. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. Fig. PLATE 4. GARIBIUS MONTICOLENS Chamberlin. Prosternum. Anterior border of prosternum more highly enlarged, showing teeth, spines and form of median incision characteristic in the genus. Antennae. Left eye showing typical form and arrangement of ocelli (single ocellus at left). Anal leg, mesal aspect, showing position and usual form of the tibial crest characteristic of males of the genus. Tibial crest of anal leg, ectal view, more highly enlarged. GARIBIUS CATAWBAE Chamberlin. .Anal leg, mesal view. Tibial crest of anal leg, ectal view, more highly enlarged. TIpABIUS TIVIUS Chamberlin. Right gonopod, showing form of claw and of basal spines character istic of the genus. Bull. Mus. Comp. Zod0l. Lithobiid Genera Plate 4 io . “elo ee ph vce a a5 , ere ae ie wh: i P a A | r Tat ae. Wi a) tor Lar ‘oe el, i ei Fe ‘ the | Bary: fs hired ¥) « | (hee pas | y wh ! i i y P| x di i ayaa i aha yd e) w CHAMBERLIN.— Lithobiid Genera. PLATE 5. GARIBIUS MONTICOLENS Chamberlin. (See also Plate 4). Fig. 1. Penult leg, caudal aspect. TipABIUS TIvIuS Chamberlin. (See also Plate 4). Fig. 2. Prosternum. Fig. 3. Anterior border of prosternum showing teeth spines and mesal incision, the latter of the form prevailingly but not exclusively found in the genus. Fig. 4. Anal leg of 9, mesal aspect, showing the usual form for both sexes in the genus. Fig. 5. Penult leg mesal (caudal) aspect. Fig 6. Antenna, representing the form typical in the genus. Fig. 7. Right eye, showing characteristic form and arrangement of ocelli, (single ocellus at left of figure). TIDABIUS OPIPHILUS Chamberlin. Fig. 9. Basal spines of left gonopod of 9. Lithobiid Genera Plate 5 Mus. Comp. Zool. Bull, _ ctl 2s wae M., C..Z:,. No. 369, santan Cruz Island; M. C. Z., No. 370, 376, Pacific Grove; M. C. Z., No. 371, Felton Big Trees; M. C. Z., No. 372, Mill Valley; M. C. Z., No. 379, Berkeley. R. V. Chamberlin. Also taken in California at Capitola and Santa Barbara. This species is subject to considerable variation in slenderness. The typical specimens from Pacific Grove vary from 7.5 to 8 times longer than the width of the tenth plate; while those from Sausalito and Mill Valley seem on the average to be considerably more slender, from 9 to 9.5 times longer than width of that plate. The head, too, varies more than usual in its relative length and breadth. The spining, e. g. of the twelfth legs, likewise varies, as indicated above. It does not seem possible, however, clearly to establish any well- defined local varieties as intergrades are everywhere to be found. As 0 O° 144 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. a species it is very distinct and easily separated from the others of the genus. The rather limited extent of variation in the form of the characteristic basal spines of the female gonopods may be judged from Plate 3, fig. 1-3, drawn from specimens from Brookdale, Santa Barbara, and Sausalito respectively. OaBius SASTIANUS (Chamberlin). Lithobius sastianus Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 157. Pomona college journ. ent., 1910, 2, p. 371, 374. Driagnosis.— Moderate; first plate decidedly narrower than third, and the head than the tenth. Brown of somewhat reddish tinge, the posterior legs also brown. Ocelli near ten in three series. Prosternum 12 times wider than long; distance between chitinous spots 3.5 + times the dental line; sinus v-shaped. Median lobe of genital forceps longest; basal spines long and slender, outer longer. Coxal pores very small, 2, 2, 3, 3 to 3, 4, 4, 4. Claw of anal leg single. Third joint of all legs armed dorsally and ventrally. Spines of anal legs, ventral: 0, 4,°3;'3; Lor’ 0,°1,'3, 2, 1; dorsal; 150) 3391-0" of penult; ventral, 0, 1, 3, 3, 2; of twelfth, dorsal, 0, 0, 3, 2, 2; of first ventral, 0, 0, 1, 2,1 to 0, 0,1, 3,2. Length 8 to 10 mm. Description.— Dorsum and head with antennae brown of some- what reddish tinge; antennae pale distad as usual. Venter paler brown. Most legs yellowish, the posterior pairs brown. - Body moderate, in the types being from 7 (2) to 8.5 (co) times longer than width of the tenth plate. Moderately narrowed cepha- lad, with first plate of same width as tenth and narrower than head. Ratio of widths of head and of first, third, eighth, tenth, and twelfth plates near 49 : 45 :45 :53 : 53 : 52. Head equal in length and breadth or very nearly so. Widest near marginal breaks, between which and the eyes the sides are nearly straight. Bristles few, long. Antennae short, reaching the fifth segment. Composed of the usual twenty short and very short articles which decrease in length from the third distad. Each eye composed of about ten ocelli which are arranged in three series: e. g., 1 +4, 3, 2. Ocelli decreasing in size from the dorsal row ventrad. Prosternum 1% times wider than long. Distance between chiti- nous spots 2 times the width at level of bottom of sinus; 3.55-++ CHAMBERLIN: LITHOBIID GENERA. 145 times the dental line. Teeth small, the interval between those of each pair relatively wide, semicircular. Mesal incision v-shaped. Spine distally bristle-like as usual. First dorsal plate nearly 1.8 times wider than long; conspicuously narrowing from anterior end caudad. Posterior angles of the ninth, eleventh, and thirteenth dorsal plates straight or with a tendency to very slight extension, not at all excised. Coxal pores very small: ae Dee ae DOV ON ot oF 4, Ay Bs Spines of first legs, 9 ae 735; of second, feq34 to ¢ : 2,22 0,22 2; t » 2, 0, 0, 3, 2, 0, 0, 2 of third to fifth, % 0. 0, Bal a 0; 0752: 3: 0, 0, ¢ 9s TBR ° 29 ee (Si. io) S fa) po 1: L z of sixth to ninth, 335; of the tenth and eleventh, §--33°3; of the twelfth, ° 0):0;.3;.2;;2 o DON el omiee 1, 0, ; Or 90-33-23 of ihe thirteenth, §-73:3:5; of the penult, 977°3'3'3; il the anal, §°3°371 with one claw. Last two pairs of coxae laterally armed. Anal and penult legs in both sexes moderately uniformly inflated. Claw of female gonopods of moderate length, well curved; dis- tinetly tripartite with the middle lobe somewhat longest as usual. Basal spines moderately long and slender; acuminate from base to apex, with the outer longer than the inner. Length of types from 8 to 8.5 mm. A male 8.5 mm. long has antennae 2.8 mm. and anal leg 2.8 mm. long with width of tenth plate 1 mm. A female 8 mm. long has tenth plate 1.13 mm. wide. Typr.— M. C. Z., No. 336, California: Shasta Springs. R. V. Chamberlin. A divergent species known only from the types, a single pair. OABIUS TABIPHILUS, sp. nov. Plate 1, fig. 9; Plate 2, fig. 8. DiaGnosis.— Body moderate, first plate much narrower than third and head a little narrower than tenth. Dorsum dilute orange-yellow, head the same or a little deeper. Ocelli seven to eleven in two or three series. Prosternum 1.5 to 1.6 times wider than long; distance between chitinous spots 3.5 to 3.8 times the dental line; sinus nar- rowly v-shaped. Lobes of female genital forceps short and blunt, median but little longest; basal spines small. Coxal pores 3, 3, 3, 3- 3, 4, 4, 3. Claw of anal leg single. Third joint of all legs armed dorsally and ventrally. Spines of anal legs, ventral, 0, 1, 3, 2, 0, dorsal, 1, 0, 3, 0, 0; of penult, ventral, 0, 1, 3, 3, 1; of twelfth, dorsal, 146 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 150, 331, 20rd, 023,01; 45, of first legs yentral)j0; 0,153) ed seneth 7-10 mm. ; DescriptTion.— Dorsum dilute orange-yellow to light brownish yellow. Head like dorsum or with orange a little deeper; the frontal region pale. Antennae dilute orange-yellow, paler distad. Pro- sternum like head but more dilute. Venter pale yellowish, excepting the posterior plates which are light orange. Legs very pale or whitish yellow excepting the posterior pairs which are more densely pig- mented, commonly bright lemon-yellow. Body proportionately wide the length being from 7.25 to 8 times greater than width of the tenth dorsal plate. Body only moderately narrowed cephalad, the widths of head and of first, third, eighth, tenth, and twelfth dorsal plates being to each other about as 28 26: 27.5 232 : 32:31. Head usually slightly longer than wide (28:27). Of the usual form. Antennae very short, reaching the fifth segment; moderately attenuated distad but tip not very slender. The twenty articles mostly very short and closely compacted, decreasing distad from the second or third. Ocelli of each eye (Plate 1, fig. 9) from seven to eleven in number, arranged in two or three series: e. g., 1 + 3,3; 1+ 4, 4; 1+4, 3; 1+ 2,3, 2; 1+ 4, 3, 2; 1+ 4,4, 2. Single ocellus of about same size as most caudal one of upper series, or sometimes smaller and at others somewhat larger. Ocelli decreasing in size cephalad and ventrad, those of third series, when present, commonly very small. Prosternum 1.5 to 1.6 times wider than long. Distance between chitinous spots 2.5 to 2.8 times width at bottom of sinus, 3.5 to 3.6 times as great as length of dental line. Teeth small, dark tipped. Sinus narrowly v-shaped, but sides a little concave. Spine nearly as in related species, stout at base and bristle-like distally; straight or nearly so. First dorsal plate 1.73 to 1.75 times wider than long; caudal corners well rounded; sides moderately diverging cephalad. Eleventh and thirteenth plates with caudal margins straight or corners showing even a slight tendency toward extension, not at all excised. Coxal pores small, circular, the most proximal often much smallest: 345 4 Seid; By 3,3: ° q 0,0; 23:2; 1 0, 0, 2, 2, 2 Spines of first legs, g-o:i'31; of the second to seventh, 9-9:7°3°33 : 0, 0, 2, 2, 2 0, 0, 3, 2, 2 of the eighth to tenth, oo-¢3:5; of the eleventh, 9-o-333; of the CHAMBERLIN: LITHOBIID GENERA. 147 twelfth, $333 Or (saa; of the thirteenth, §3:3:3:3; of the penult, ores; of the anal, 9-733; laterally armed. Anal and penult pairs of legs in both sexes short and rather slender, being but moderately uniformly thickened as in OQ. tiganus, ete., scarcely if at all more so in male than in female. Gonopods of male considerably extended; pale; slender. In ven- tral view commonly appearing rounded apically but sometimes sub- truncate. Bearing mostly two bristles. Claw of female gonopods moderately bent and concave; tripartite with the teeth blunt and short, the median one being a little longest. Spines (Plate 2, fig. 8) small; broadest at base, the sides incurving and moderately converging to the acuminate distal division; apex rounded. Process of sternite distally truncate or slightly convex. Length from 7 to 10 mm. A male 8.5 mm. long has antennae and anal legs both 3 mm. long, and the tenth plate 1.17 mm. wide. Pseudomaturus.— Coloration as in maturus. Antennae very short. Articles distad of second very short and closely compacted. Ultimate article in specimen described longer than the two preceding together and nearly equalling the three pre- ceding. Ocelli 1 + 4,4. The first ocellus of dorsal series largest, the single one small. Ocelli somewhat irregular in form, size, and position. Prosternum 1.4 times wider than long. Distance between chitin- ous spots 2.4 times width at level of bottom of sinus; 3.8 times the length of dental line. Line of apices of teeth a little recurved. Coxal pores 3, 3, 3, 2 — 3, 3, 4, 2. Spines of first legs, (+0754; of the second, (7734; of the eighth, cae Ar zo 5; of others as given for maturus. ; claw one. Last two pairs of coxae Length of specimen described (a o) 6.5-+ mm. Praematurus (Late).— Color as in maturus or somewhat paler throughout. Antennae as in pseudomaturus. Ocelli 1+ 3,2; 1+ 4, 3. First ocellus of dorsal series the largest. Prosternum nearly as in maturus; teeth smaller and more acute. 1.47 times wider than long. Distance between chitinous spots 23 times width at level of pe rem of sinus; 3.5 times dental line. Coxal pores very small: 2, 3, 3, 2. Spines of first legs, [-:t-34; of the secon nd, foto (left) or pos 0, 0,0,2 (right); of the third, }-j3:34 (left) or S43 vp Gan. of the fourth 148 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. to eighth, [t34; of the ninth and tenth, O33; Of the eleventh, oes; of the twelfth, (0234 oe zi of the thirteenth, §1:3:33; of the DONO nsel 1, 0, 3, 0, 0 penult, o-333; of the anal, o73-3. Last two pairs of coxae laterally armed. Claw of female gonopods short and pale; the lateral teeth consid- erably the smaller. Inner spine about two thirds as long as the outer; both acuminate from base. ee (2) about 6.5 mm. Immaturus.— Of a pale, slightly yellowish, tinge, the head and most caudal segments being of a denser yellow. Anal legs nearly white, darker proximad. Antennae pale yellow. Antennae with articles rather more loosely joined than in the later stages, submoniliform. Ocelli 1 + 3, 2. The single ocellus and the first two of the dorsal series largest and about equal in size; the others distinctly smaller; pale and well separated. Prosternal teeth pale, in relative size and form as in praematurus or nearly so, 1.5 times wider than long. Distance between chitinous spots only 2.1 times width at bottom of sinus; 3 times length of dental line. Coxal pores very small and pale margined; 2, 3, 3, 2. Spines of first legs, ma 0 est of the second and third, (34; of the fourth to seventh, 5 0, otis a of the oon ¢ i TES of the ninth, * 0-0; nee : or §0121; of the tenth and eleventh, 9.3 a of the twelfth, (22333 = 1 0, of the thirteenth, j33'4; of the penult, §1:3:3:4; of the anal, j Last two pairs of coxae laterally armed. Female gonopods short and pale but distinetly triarticulate. Ulti- mate article with claw represented by a slight pale and acute spinule or tooth; without hairs. Second article apparently with a single short ventral bristle. First article with two ventral bristles. Spines 1+ 1, small and pale, acutely conical. Length ad 6 mm. Agenitalis II.— Pale, the yellowish tinge very dilute, clearer in head and prosternum and on caudal segments. Anal legs white with a tinge of yellow. Articles of antennae distad of the second very short; the ultimate article proportionately longer than in older stages, being equal in length to the four preceding taken together, or very nearly so. Ocelli 1 + 3. The two caudal ocelli of the series proper the largest, Ait 0 70,2, 0,3, 1.3,20° CHAMBERLIN: LITHOBIID GENERA. 149 irregular; the single one a little smaller and the most anterior one much smaller; middle ocellus of series a little lower than the other two. Organ of Toémésvary in outline larger than anyone of the ocelli. Prosternum essentially as in later stages (specimen measured had prosternum 1.55 times wider than long; distance between chitinous spots 2.4 times width at sinus and 3.7 times the dental line). Coxal pores, 1, 2, 2, 2. Spines of first legs, oat; of the second to sixth, Ppt; of the seventh to ninth, 9°5 cone i; of the tenth, 5° 44 ih i i; of the eleventh, } 0: z h i; ae of the twelfth, 0 eed 2 41 1; of the thirteenth, ¢ {7743 of the penult, }33°4; 2 of the anal, 0.0.2 0.0 None of coxae armed. Gonopods of male appearing as slight, rounded, and wholly gla- brous protuberances. Gonopods of female appearing as much longer processes which are slender and conically pointed; biarticulate; glabrous; with no trace of claw or of basal spines. In the male studied there is no longer any trace of anal glands; but in a female, the glands are traceable in an apparently degenerate condition. Length of a female 5.38 mm.; of a male, 5.7 mm. Typrs.— M. C. Z., No. 411, California: Santa Barbara. R. V. Chamberlin. Paratypres.— M. C. Z., No. 328-333, 407-410, California: Santa Barbara. R. V. Chamberlin. Many specimens of this divergent and very distinct species were collected by the author in June, 1909, and in March, 1913. It is most easily detected through the uniform absence of any dorsal spine on the fourth joint of the anal legs. Oaxsius (NYCTOBIUS) DECIPIENS, sp. nov. Plate 13; fig.0 5: Diacnosis.— Slender; first plate decidedly narrower than third; head wider than long, narrower than tenth plate. Testaceous to hght chestnut; head brown to chestnut. Ocelli eight to ten in three series, the single ocellus distinctly largest. Prosternum less than 1.5 times wider than long; distance between chitinous spots 3.4— times dental line; sinus wide and shallow. Lobes of genital forceps short, median little longest; basal spines short and broad. Coxal pores 150 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 1, 2, 2, 2 to 3, 4, 4, 3, small. Anal leg with two claws. Third joint of all legs armed dorsally and ventrally. Spines of anal legs, ventral, 0; 13, 240; dorsal, 12'0, 3; al; 0; for penuligventrals 0; 3:3, bor thirteenth, ventral; 0, 1, 2, 3, 2 or, rarely 0,1, 3, 3, 2; of twelith, dorsal, 1, 0, 3, 1, 1; of first, ventral, 0,0,1,2,1. Length 8.5 to 10mm. DescripTion.— Dorsum testaceous to light chestnut, with the first and the last plates darker than the median. Head brown to chestnut. Antennae nearly same color as head; paler at tips. Prosternum similar to head but somewhat paler. Venter pale testaceous to dilute brown, the caudal plates darker. Legs nearly same in color as the contiguous plates of venter, the posterior pairs being thus darker, but these paler along mesal and ventral surfaces, especially distad. Body in types from 8.5 to 9 times longer than width of the tenth plate. Rather conspicuously narrowed cephalad, with the first plate much narrower than the third. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other about as 27 :25 : 28 : 32 :33 : 30. Head wider than long (27 : 26). Subrotund; the sides well rounded posteriorly; caudal margin mesally straight. On the caudal portion are two rather distinctly impressed furrows which diverge forwards. Antennae short, reaching the fifth segment. Articles short, de- creasing in size distad, the last ones being decidedly shorter than the proximal ones. Hairs moderately long. Each eye consisting of from eight to ten ocelli arranged in two or three series: ¢. g.. 1+ 4, 4; 1+ 4, 3; 1+ 1, 4, 3; 1+ 3, 3, 1; 1+ 2,4, 1. All ocelli rather large; the single one distinctly largest. Prosternum 1.47 times wider than long. Distance between chiti- nous spots 2.2 times the width at level of bottom of sinus; 3.36 times the dental line. Teeth small and acute with sides more or less in- curved; line of apices recurved. Sinus wide and rather shallow; sides and bottom rounded. First dorsal plate 13 times wider than long; sides widely rounded behind, diverging moderately cephalad. Dorsum smooth and shining or but finely and obscurely roughened caudad. Coxal pores small, oe ee proximad as usual. In number and arrangement, é. g., 1, 2;2,2; 2,2, 2,2; 2,3,3,2; 3, 4, 4, 3. 1, 1(2), 1 e pul ies Spines of first legs, os : of the second to seventh, 9-94°353 0, a outa of the eighth and ninth, 9 OF f-o7-33; Of the tenth, ess OF i a 0, 0. 3 2 ORORS et LVOnsi ar hat " DET oe ee ot the ere LOR2R2. 6 . ot the twelfth, 9° 0, Od 3h, 2 or 0, 0, 2, 3,29 . 1,0; 3)1)1. of the thirteenth 3.5 or occasionally (425-33; , i ae 2 3,2 Y 0,123.2 2 2, .2, 2, of the penult, CHAMBERLIN: LITHOBIID GENERA. 151 TOSS adele 1, 0, 3, 1,0 (in SESE IID of the anal, 0, 1, 3, 2,0 ; claws two. Last two pairs of coxae laterally armed. Anal and penult legs very short; slender; fourth and fifth articles weakly longitudinally furrowed dorsally. Claw of female gonopods short and relatively wide; tripartite, with the lobes short, the median being not much longest. Spines wide and short, the sides parallel from base to the short acuminate distal portion; the latter subacute but with apex narrowly rounded (Plate 3, fig. 5). Inner side of first article convex. ' Length of types from 8.5 to 10 mm. A female 8.5 mm. long has antennae 2.8 mm. long and tenth dorsal plate 1 mm. wide. Typre.— M. C. Z., No. 405, California: Pacific Grove. R. V. Chamberlin. PaRATyPEs.— M. C. Z., No. 334, California: Pacific Grove. R. V. Chamberlin. Oasius (NycToBIus) INEPTUS, sp. nov. Plate 2, fig. 3, 4; Plate 3, fig. 4. Diacnosis.— Moderate; first plate narrower than third and head a little narrower than tenth. Dark brown, chestnut at ends, head and first tergite deeper chestnut. Ocelli six or seven in two series, the single ocellus clearly largest. Prosternum near 1.65 times wider than long; distance between chitinous spots 3.25+ times the dental line. Lobes of genital forceps short and rounded; basal spines short and broad, close together, outer spine but little longer than inner, not more robust. Coxal pores small, 1, 2, 2, 2—2, 2, 2, 2. Anal leg with two claws. Third joint of all legs armed dorsally and ventrally. Spines of anal legs, ventral, 0, 1, 3, 2, 0, dorsal, 1, 0, 3, 1, 0; of penult, ventral, 0, 1, 3, 3, 1; of thirteenth, ventral, 0, 1, 2, 3, 2; of twelfth, dorsal, 0, 0, 3, 1, 1; of first, ventral, 0, 0,1, 2,1. Length 7-7.3 mm. Description.— Dark brown or chestnut, deeper chestnut anteriorly and posteriorly. Head and first dorsal plate deeper in color, appear- ing almost black caudad of the suture in front of which it is rather pale. Antennae chestnut, paler distally. Prosternum dark chestnut. Venter brown, the anterior and posterior plates darker and the latter especially more reddish. Legs brown. Body 7.5 to 8 times longer than width of tenth plate; conspicuously narrowed cephalad to the first plate, with the head rather large. Widths of head and of first, third, eighth, tenth, and twelfth plates to éach other nearly as 25 : 23 : 25 : 27 : 27 : 27. 152 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Head subrotund as in O. decipiens, the sides being conspicuously convex. Clearly wider than long, the ratio being near 25 : 23. Antennae short as usual. Articles nineteen or twenty; decreasing in size distad as in other species. In types, the ocelli are six or seven in number and are arranged in two series: ¢. g.. 1+ 3, 2; 1+ 3, 3. Single ocellus clearly largest though proportionately smaller than in O. decipiens. Other ocelli of moderate size. Teeth of prosternum moderate; acute, with sides straight and intervals angular; inner tooth larger than the outer and the line of apices straight or nearly so. Prosternum 1.65 times wider than long. Distance between chitinous spots 2.1 times width at level of bottom of sinus; 3.26 times the dental line. First dorsal plate with the sides between the rounded anterior and posterior portions nearly straight, moderately diverging cephalad; 1.9 times wider than long. Plates strongly arched, smooth and shining. ‘ Coxal pores 1, 2, 2, 2 — 2, 2, 2, 2, small. Spines of first legs, (0q34; of the second to seventh or ieee 0, 0,1, 2.2, 0, 0, 3) 252 0, 0, 3, 2, 2 Trai; Of the eighth and ninth, ¢-o:737; of the tenth, (pis3 OF 0, 8, 2, 2 0,0, 3, 2,2 0, 0, 3,1,1 0, 0, 2,2, 25 of the eleventh, { 0, ), 0, 2,2, 23 of the twelfth, 0, 0; 2, 3, 25 of the 1, 0, 3,1,1 1 DUB VA ye 1, 0, 3,1,0, thirteenth, (3°33; of the penult, g73:34; of the anal, 73203 claws 2. Last two pairs of coxae laterally armed. Spining thus as in O. decipiens. Anal and penult legs short; a little and uniformly thickened. Claw of female gonopods (Plate 2, fig. 3) relatively wide and short, moderately curved; tripartite with the lobes small and distally obtusely rounded, the median longest though not greatly so. Basal spines (Plate 2, fig. 4; Plate 3, fig. 4) of each side close together on a distinct low lobe or tubercle; broad, with the sides parallel or a little diverging from base to the acuminate apical portion which is sub- acute. Sternal process distally subacutely rounded. Length of types 7 to 7.3mm. A female 7.3 mm. long has antennae 2.6 mm. long and anal leg 2.8 mm. long, and the tenth dorsal plate .98 mm. wide. Typr.— M.C. Z., No. 335, Oregon; Portland, August, 1902. R. V. Chamberlin. Paratype.— M. C. Z., No. 406, Oregon: Portland, August, 1902. R. V. Chamberlin. The types consist of two specimens. CHAMBERLIN: LITHOBIID GENERA. 153. This species is very close in general structure to O. decipiens. These two species, aside from differing in size, in ocelli, in proportions of prosternum, and of first dorsal plate, etc., are at once to be separated by the differences in the basal spines of the female gonopods (Plate 3, fig. 4, 5). KIBERBIUS, gen. nov. Head with the usual marginal interruptions. Antennae short; composed normally of twenty articles. Eyes composed of from one to three series of ocelli. Single ocellus but little enlarged. Prosternal teeth 2+ 2; small; the dental line short with line of apices straight or a little recurved. Median sinus wide v-shaped. Posterior angles of none of the dorsal plates produced. Coxal pores small and circular, uniseriate. Claw of female gonopods tripartite with the median not greatly exceeding the lateral. Basal spines long and slender; 2 + 2. Gono-_ pods of male uniarticulate. Tarsi of anterior legs with division typically indicated by difference in diameter, clear transverse line and partial suture (Plate 4, fig. 6), but indication of division may be lacking (in K. nannus only). Anal legs in both sexes conspicuously longitudinally furrowed along the mesal surface, the furrow broad and deep and the leg having a consequent thin or flattened appearance (Plate 5, fig. 1). None of the coxae laterally armed. None of the legs cephalad of the twelfth having the third joint with three dorsal spines. Legs between the first or second and the twelfth pairs all with the fifth joint bearing two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral 0, 1, 3, 2, 0 or 0, 1, 3, 1,0 Gn K. nannus only); claw single. Dorsal spines of penult legs, 1, 0, 3, 1, 1 or 1, 0, 2, 1, 1; ventral 0, 1, 3, 3, 1 to 0, 1, 2, 3, 0 (K. nannus); claws 3, the anterior one distinct, long, straight and spine-like. Dorsal spines of thirteenth legs, 1, 0, 3, 1, 1 ore! 0, 2, i. i ventral, 0,0, 2;.3;2 or 0; 0):2,3;1.. Dorsal spines of twelfth legs, 0,0, 2, 1,1; ventral, 0, 0, 2,3, 2 or0,0,1,3,2. Dorsal spines of eleventh, 0, 0, 2, 2, 2 or 0,0, 0, 2,2; ventral, 0, 0, 2, 3, 2 .to 0,.0, 0, 3, 2. Dorsal spines of first legs, 0, 0, 1, 1, 1 or 0, 0, 1, 2, 1; ventral, 020; 1, 1,3 or 0/051, 2, \1. Length under 9 mm. Typr.— K. ogmopus, sp. nov. 154 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. This genus, at present represented by three Californian species, 1s most readily distinguished by the presence in males and females alike of the conspicuous wide and deep longitudinal furrow along mesal surface of anal legs or by a pronounced corresponding flattening of the same. The genus is also separable through the absence of lateral spines from the posterior coxae. Key to Species of Kiberbius. a. Ventral spines of anal leg, 0, 1, 3, 1, 0; third joint of all or most legs between second and twelfth unarmed dorsally; length UNGER 7 MAIN sept cS ET Ae RISE, a K. nannus, sp. nov. aa. Ventral spines of anal legs, 0, 1, 3, 2, 0; third joint of all legs dorsally armed; length 8 mm. or more. b. Head longer than wide, of nearly same width as tenth plate; first and third plates of same width or very nearly so. K. remex (Chamberlin) bb. Head equal in length and breadth; head distinctly narrower than the tenth plate and the first plate than the third. K. ogmopus, sp. nov. KIBERBIUS OGMOPUS, sp. Nov. Plate 4, fig. 5, 6; Plate 5, fig. 1-3. DraGnosis.— Body moderately stout; first dorsal plate 1.6 times wider than long, narrower than the third; head equal in length and breadth, narrower than tenth plate. Pale brown, first and last plates darker; head dark brown.’ Ocelli about ten in three series. Pro- sternum 1.5 -+ times wider tham long; distance between chitinous spots 4.25 — times dental line. Lobes of genital forceps acute; basal spines long and slender, narrowing toward middle. Coxal pores 2, 2, 2, 2. Third joint of all legs dorsally armed. Spines of anal legs, ventral, 0, 1, 3, 2, 0; of penult legs, ventral, 0, 1, 3, 3, 1, dorsal, 1, 0, 3, 1, 1; of twelfth, ventral, 0, 0, 2, 3, 2; of first, ventral, 0, 0, 1, 2, 1, dorsal, 0, 0,1, 1,1. Length 8 to 9 mm. Description.— Dorsum pale brown; the anterior plates darker and suffused with purplish; the posterior plates also darker. An- tennae light purplish brown, paler at tips. Prosternum pale brownish. Venter very pale mesally, the anterior portion strongly suffused with 155 LITHOBIID GENERA. CHAMBERLIN: Fic. 2.— Distribution of Kiberbius. 156 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. purplish, the caudal ones distinctly darker. Anal and penult legs more or less suffused with purplish proximad of the tarsi. Body typically from 7.5 to 8 times'longer than width of tenth plate; conspicuously narrowed cephalad, the widths of head and of first, third, eighth, tenth, and twelfth plates in female type to each other as 47 : 42 249 :55 : 55 : 52, the first plate thus conspicuously narrower than the third. Head subcordate; caudal margin straight. Equal in length and breadth or a very little wider than long (Plate 4, fig. 5). Antennae moderately short. Articles of medium length; sub- cylindric; not closely compacted; those immediately preceding the ultimate short. Ultimate longer than the two preceding together. Each eye consisting mostly of about ten ocelli arranged in three series: ¢. g., 1+ 4, 3, 2. Ocelli of purplish cast. Single ocellus paler and a little largest. Those of most ventral row considerably smaller than the others. Organ of Témésvary in outline about size of the smaller ocelli. Prosternum (Plate 4, fig. 6) 1.53 times wider than long. Distance between chitinous spots 2.57 times width at level of bottom of sinus; 4.23 times the dental line. Teeth acute with sides straight; line of apices only slightly recurved. Sides sloping directly from ectal tooth. Spine of usual type. First dorsal plate only moderately narrowed caudad; caudal margin slightly mesally incurved; 1.6 times wider than long. Poste- rior corners of thirteenth dorsal plate straight; of the ninth and eleventh rounded a shortly obliquely truncate. Coxal pores 2, 2, 2, 2. Spines of first legs, O21; of the second, COT OF gers; of the third to fifth, jt i343; of the sixth and seventh, OEE of the eighth, ? Coe OF vox; of the ninth to eleventh, eer of the twelfth, (335; of the thirteenth, oeas; of the penult, 1,0,3, 1,1 3, 1, 0 0,1,3,3,13 of the anal, 77:33. Anal leg short. The tibia and the tarsal joints characteristically very deeply longitudinally furrowed or channelled along ventral surface, the femur less deeply so (Plate 5, fig. 1). The penult leg has the tibia and tarsi similarly but much less conspicuously fur- rowed. Claw of female gonopods short and proportionately wide; tri- partite, the teeth and the incisions between them acute. Basal CHAMBERLIN: LITHOBIID GENERA. 17 spines relatively long and slender, the outer one in ventral aspect appearing narrowest near middle of length, the distal division short and acute; inner spine presenting edge ventrad and appearing in outline acutely acuminate from the very base (Plate 5, fig. 3). Length mostly 8 to 9 mm. Typr.— M. C. Z., No. 339, California: Eaton’s Canyon, near Altadena. R. V. Chamberlin. ParatyPEs.— M. C. Z., No. 419, California: Eaton’s Canyon, near Altadena. R. V. Chamberlin. The specific name of this species refers to the characteristic chan- nelling of the anal legs. KIBERBIUS NANNUS, sp. nov. Plate 4, fig. 7; Plate 5, fig. 4. Diacnosis.— Slender; first plate 1.6— times wider than long, much narrower than third; head longer than wide, narrower than tenth plate. Yellow, the head brightest. Ocelli four or five in one or two series. Prosternum 1.5— times wider than long; distance between chitinous spots four times the dental line; sinus v-shaped. Median lobe of genital forceps considerably longest; basal spines long and slender, inner nearly as long as outer. Coxal pores 1, 1, 1, 1. Third joint of all or most legs between second and twelfth unarmed dorsally or ventrally. Spines of anal legs, ventral, 0, 1, 3, 1, 0; of penult, ventral, 0, 1, 2, 3, 0, dorsal, 1, 0, 2, 1, 1; of twelfth, ventral, 0,0, 1, 3, 2; of first, ventral, 0, 0, 1,1, 1, dorsal, 0, 0, 1, 1, 1 or 0, 0, 1, 2,1. Length 6+ mm. Description.— Yellow. The head and posterior segments a brighter yellow. Antennae and posterior legs yellow. Body between 8.5 and 9 times longer than width of tenth plate. Conspicuously narrowed cephalad. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 35 : 33 : 37: 43 :44 : 42, the first plate thus being considerably narrower than the third. Head (Plate 4, fig. 7) strongly narrowed cephalad of the eyes. Caudal margin straight. Considerably longer than wide (39 : 35). Antennae short. Articles short and decreasing cephalad in the usual way. Ultimate article distinctly longer than the two preceding together. 158 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli in type three or four in one or two series: e.g., 1-+3; 1+3.1. Single ocellus not specially enlarged. Prosternum 1.48 times wider than long. Distance between chiti- nous spots 2.9 times width at level of bottom of median sinus; 4:times the dental line. Prosternal teeth small, the mesal one of each pair in the type somewhat larger than the outer and the line of apices thus straight or very slightly procurved. Sinus v-shaped with the sides a little convex. First dorsal plate gradually narrowed caudad of the middle with posterior corners well rounded; caudal margin slightly incurved mesally. Plate as a whole 1.57 times wider than long. Coxal pores 1, 1,1, 1: : 0,0, 1, Ose: 2, Spines_ of ist legs). coi Of toi; oO: the second, $4143 2 1 , 0, 2, 2 0, 0, 0, 2, 0, 0, 0, 2, 2 of the third, 1, Ti OF poi; of the fourth, o-o-o-77; of the fifth WOO} iste, 0, 0, 0, 2, to ninth, 9-p°9-3-3; of the tenth and eleventh, 9-7; O33; of the twelfth, 0,0,2,1,1 . 1, 0,2,1,1 TOL32; of the thirteenth, ¢o-¢34; of the Jems reso; of the j, 20.3.1.0 anal, 0,1, 3,1,0° Last three joints of the anal legs conspicuously longitudinally channelled along mesal side as in K. ogmopus but not quite so strongly so. . Claw of female gonopods scarcely curved, being nearly flat; tri- partite, with the median lobe decidedly longest, the lateral ones at same level considerably proximad of distal one. Basal spines espe- cially long and slender, the inner as long as or nearly as long as the outer; in ventral view the spines are narrowed gradually from base but the sides incurve midway between base and apical portion (Plate 5, fig. 4). Length of female type, 6.2 mm.; width of tenth plate .7 mm. Typr.— M. C. Z., No. 340, California: Eaton’s Canyon near Altadena, April, 1913. R. V. Chamberlin. At first, it seemed likely that this form was to be identified with an immature stage of the preceding species because of certain apparently immature features, such as, to some extent, the spining of the legs. However, the gonopods are large and fully developed with no sugges- tion whatsoever of immaturity. Upon clearing and mounting the specimen showed well-developed eggs within the body. CHAMBERLIN: LITHOBIID GENERA. 159 KIBERBIUS REMEX (Chamberlin). Lithobius remex Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 156; Pomona college journ. ent., 1910, 2, p. 370, 374. DraGnosts.— Body moderate; head longer than wide of nearly same width as tenth plate; first dorsal plate 1.8 times wider than long. Brown. Ocelli about six in two series. Prosternum 1.6 times wider than long. Coxal pores 3, 4, 4, 3. Third joint of all legs dorsally armed. Spines of anal legs, ventral, 0, 1, 3, 2, 0; of the first, ventral, 0; 0, 1, 2; 1.. Length 8 mm. DescripTion.— Head and dorsum brown. Legs pale brown. Body slightly less than 8 times longer than width of tenth plate. Body nearly parallel sided, the head and the tenth plate of same width. Widths of head and of first, third, eighth, and tenth plates to each other as 28 : 26 : 26 : 27 : 28, the third and first plates being of same width and the head much wider than the first. Head cordate, considerably more strongly narrowed cephalad of eyes than usual. Widest near marginal interruptions. Longer than wide in ratio 15 : 14, very nearly. Hairs long, few. Antennae short. Consisting of the usual twenty articles which, excepting the first two and the ultimate, are short. Ultimate nearly equalling the three preceding taken together. Eye consisting of six ocelli arranged in two longitudinal series: é. g., 1+ 3, 2; 1+ 2, 3. Deeply pigmented. Single ocellus but little enlarged. Those of upper series larger than those of lower. Prosternum 1.6 times wider than long. Teeth of moderate size with sides somewhat concave and apices narrowly rounded. Spine as usual. First dorsal plate with sides convex toward ends but between these nearly straight, considerably diverging cephalad. Posterior minor plates with their corners obliquely truncate, the line of truncation making only a slight angle with the posterior margin. Hairs short and very sparse. Coxal pores of moderate size: 3, 4, 4, 3. Ventral spines of first legs, 0, 0, 1, 2, 1; of the sixth and seventh, 0,0; 3: 25 of themmth, 0; 0,2; 3, 2:, of the anal, 0,.1,.3, 2, 0. Anal legs distally conspicuously flattened with tarsus somewhat clavately widening distad. Proximal joints laterally longitudinally suleate and at some time complanate. 160 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Length 8 mm.; width of tenth plate 1-+ mm.; length of antenna and of anal leg, 2.9 mm. Tyre Locauity.— California: Shasta Springs, 1902. R. V. Chamberlin. Known only from the type specimens. PAoBIUS, gen. nov. Lateral margins of head continuous, without any abrupt interrup- tions (Plate 4, fig. 3). Antennae short, consisting normally of twenty articles. Eyes composed of ocelli arranged in two or three series. Single ocellus differentiated, equal to or larger than largest of others. Prosternal teeth 2 + 2, with line of apices straight or nearly so, and the dental line one fourth or less the distance between the chiti- nous spots. Sinus proportionately large, v-shaped. Spine stout at base but distally attenuated and bristle-like (Plate 3, fig. 3). Posterior angles of none of the dorsal plates produced. Coxal pores small and circular, uniseriate. Claw of female gonopods tripartite; the lateral lobes equal and at same level, smaller than the median though not much so; or bipartite and with lobes equal. Basal spines 2 + 2, long and narrowest near middle (Plate 4, figs. 2, 4). Tarsi of anterior legs with division indicated by a pale transverse line. Anal and penult legs without special lobes or other modifications in either sex. Last one or two pairs of coxae laterally armed or rarely, none laterally armed (P. vagrans). Third joint of all but the first pair of legs with three dorsal spines or rarely that of first 12 or 13 pairs with but 2 (P. vagrans). First seven to eleven pairs of legs with but a single dorsal spine on the fifth joint. Dorsal spines of anal legs 1,0,3,0,0 or 1,0,3,1,0; ventral, 0,1, 3, 2,0; claw single. Dorsal spines of penult legs 1, 0,3, 1,0; ventral, 0,1, 3,3, 1; claws 3, the anterior accessory being long, straight and spine-like. Dorsal spines of thirteenth legs 1, 0, 3, 1,0 or 1, 0, 3, 1, 1; ventral, 0, 1, 3, 3, 2 to 0, 1, 2, 3, 2 or 0, 0, 2, 3, 2 and 0, 1, 3,3, 1. Dorsal spines of twelfth legs 41,.0,.3::1,:0 to0n0}S; 451 anid 052,115 ventral). 0;.052. 3552: Dorsal spines of eleventh legs 0, 0, 3, 1, 1 to 0, 0, 2, 1, 2 and 0, 0, 3, 2, 2; ventral, 0, 0, 2, 3, 1 or 0, 0, 2,3, 2. Dorsal spines of first legs 0, 0, 1, L, lito 0,0; 2,.1,.1 and 2; 2, 1; ventralPOs0,1 1k to OF O21 3 and Dalai CHAMBERLIN: LITHOBIID GENERA. 161 { ‘Fic. 3.— Distribution of Paobius. 162 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Length 9 mm. and less. Typr.— P. boreus, sp. nov. The five species at present known are small forms from 7 to 9 mm. in length, these being known at present as occurring in British Colum- bia, on Forrester Island off the southern Alaskan Coast, and in the mountains of Vermont. It is evidently a boreal genus and will probably be found at various points in Canada. In general appear- ance the species resemble the more typical members of Oabius. They are at once distinguished from species of other genera by the character- istic structure of the lateral margin of the head which, as in Bothro- polys, is not interrupted-or broken between the caudal corners and the eyes or only very vaguely so. The partial division of the anterior tarsi indicated by pale transverse line or by partial suture on ventral side is a correlated character of importance. The three distinct claws of the penult legs, of which the anterior one is long, is an important characteristic in comparing with Oabius. The presence of only a single dotsal spime on the fifth joint of the first nine to eleven pairs of legs and the absence of any spine at all from this joint in the penult legs, are unusual characters also differentiating these species. The presence of three dorsal spines on the third joint of all but the first pair of legs in all but the Vermont species (P. vagrans) may also be noted. The form of the basal spines of the female gonopods is generally characteristic, though in this regard the Vermont species is again divergent. Key to the Species of Paobius. a. Dorsal spines of anal legs 1, 0, 3, 0, 0 or 1, 0, 2, 0, 0. b. None of the posterior coxae laterally armed; ventral spines of thirteenth legs 0,1, 3,3,1; third joint of first ten or more pairs of legs with but two dorsal spines. P. vagrans, sp. nov. bb. Last two pairs of coxae laterally armed. Ventral spines of thirteenth legs 0, 1,2,3,2 or 0,0,2,3,2; third joint of all legs with three dorsal spines........ P. boreus, sp. nov. aa. Dorsal spines of anal legs 1, 0, 3, 1, 0. b. Fifth joint of sixth to ninth pairs of legs with but one ventral spine; head equal in length and breadth. P. columbiensis, sp. nov. bb. Fifth joint of sixth to thirteenth pairs of legs with two ventral spines; head wider than long........ P. orophilus, sp. nov. CHAMBERLIN: LITHOBIID GENERA. 163 PAOBIUS BOREUS, sp. nov. Plate 3, fig. 7-9; Plate 4, fig. 1-3. DraGnosis.— Slender; first dorsal plate 1.6 times wider than long; head much wider than long. Brown to chestnut, head lighter. Ocelli twelve to fourteen in three series. Prosternum with distance between chitinous spots 4.25 times dental line; sinus wide, u-shaped. Lobes of genital forceps acute; basal spines narrowed near middle, clavate distad. Coxal pores 2, 3, 3, 2. Last two pairs of coxae laterally armed. Third joint of all legs with three dorsal spines. Spines of anal legs, dorsal, 1, 0, 3,0, 0; of penult, dorsal, 1, 0, 3, 1, 0; of thirteenth, ventral, 0, 1, 2, 3,2 or 0, 0, 2, 3, 2, dorsal, 1, 0, 3, 1, 0; of twelfth, dorsal, 1, 0, 3, 1, 0; of first, ventral, 0, 0, 1, 3, 1, dorsal, 0,0, 2,1,1. Length 8 to 9 mm. Description.— Dorsum brown to chestnut. Head lighter, of ferruginous tinge; commonly with a dark spot in front of the caudal border. Antennae dark, paler, rufous or yellowish, at tips or in same over distal half or more of length. Posterior legs usually but slightly, if at all, more brightly colored than others. Venter much paler than the dorsum, clear brownish grey; most of the caudal plates chestnut. Prosternum and prehensors dilute chestnut. Body rather slender, being from 8.5 to 9 times longer than wide. Conspicuously narrowed cephalad, with first plate narrower than third and much narrower than head. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other about as 58 : 51 : 54 :65 : 67 : 65. Head (Plate 4, fig. 3) wider than long in about ratio 58 : 53. Antennae short. Articles distad of the second decreasing regularly, short and very short, compact. Ultimate article distinctly longer than the two preceding taken together. Eyes each consisting of mostly twelve to fourteen ocelli (Plate 3, fig. 7) arranged in three series: e. g., 1 + 5, 4, 2, which is the com- monest number and arrangement; 1 + 6, 4, 3; 1+.5, 4, 3; 1+3, 4, 4. Single ocellus distinctly largest. Other ocelli dark, regular. Ocelli of most ventral row reduced, those of others decreasing cepha- lad. Organ of Témésvary small, ordinarily not detectable in lateral view of head. Prosternum in type 1.57 times wider than long. Distance between chitinous spots 13 times width at level of bottom of median sinus; 164 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 4.25 times length of dental line. Teeth acute, the inner one of each pair a little larger than the outer so that the line of apices is straight or very nearly so. Median sinus wide, somewhat u-shaped but subacute at bottom (Plate 3, fig. 8). First dorsal plate widest cephalad, moderately narrowing caudad; 1.6 times wider than long. Posterior corners of ninth, eleventh, and thirteenth dorsal plates more or less oblique. Coxal pores, 2, 3, 3, 2. Spines of first legs, ma Os a of the second, oasT OF ty :. a3 of the third to seventh, }+ o: 134; of the eighth, } ogy OF eeeed, the dorsal spine of fifth joint on anterior side of joint; of the ninth to eleventh, (3:3, With dorsal spine of fifth joint also on anterior . 1, 0, 3, 1, 0 : ONS. 0, 3, 1, 0 side; of the Subs o023-2; of the eee 0: i 3 23 or EEE of the penult, 7733, of the anal, ¢7so5. Last two pairs of coxae laterally armed. Anal legs a little and uniformly thickened to the tarsi which are slender; but slightly if at all more thickened in male than in female; dorsally somewhat complanate and weakly longitudinally furrowed. Gonopods of male small, wart-like, each bearing two moderately long bristles. Claw of female gonopods (Plate 4, fig. 1) short, of uniform width from base to teeth; tripartite, teeth acute, the median longer than the lateral which are subequal. Basal spines with the inner one of each pair shorter than the outer; constricted near middle of length and then, especially in the outer one, widening clavately rather conspicu- ously and characteristically to the short acuminate portion (Plate 4 fig, 2) Length 8 to 9 mm. Immaturus.— Light brown. Head and prehensors somewhat orange. Anal legs brighter at distal ends. Antennae with twenty articles present. First three or four articles conspicuously longest; the others very short; ultimate article pro- portionately longer than in adult. Ocelli 1+ 3, 2. The single ocellus and adjacent seriate one of same size or nearly so. Ocelli pale and distinct, anterior ones smallest as usual. Prosternum nearly as in adult. Line of apices of teeth nearly straight. Sinus v-shaped. Coxal pores very small: 2, 2, 2, 2. 1. 0,0, 2, 0, 0 CHAMBERLIN: LITHOBIID GENERA. 165 0, 0, 1, 1, 1 . 0, 0, 2 Spines of first legs, oocaii; of the second to sixth, (744; of 0, 2,1, 1 the seventh to ninth, }:3'4, the dorsal spine of fifth joint anterior 0, 0, 3, 1,1 in position; of the tenth, 0.0.2.8.0 dorsal spine of tibia anterior in 0, 3,1, 0 p 0¥3,. q position; of ie eleventh, CesT of the twelfth, ee of the bse oO Oe O 1, 0, 3, 1,0 thirteenth, (733°) or fo334; of the penult, 6133-1; Of the anal, 1, 0, 3, 0,0 0, 1,3, 2,0° Gonopods of female with the three articles distinct, the ultimate bearing a claw which is small, slender and acute and spine-like. Third article without bristles; the second one with a single ventral bristle; and the first one with two. Length 5.5 to 6.5 mm. Agenitalis II.— Dilute brown. Head orange. Antennae pale brown, orange at tips. All legs with tarsi somewhat orange. Articles of antennae nineteen. Articles distad of the third, excepting the ultimate, very short; the twelfth and thirteenth in specimen described shorter than the adjacent ones, having been apparently formed by recent subdivision of one article. Ultimate article about as long as the three preceding ones taken together. Ocelli 1 + 3, 2. First ocellus of upper series largest, the anterior one of this series very small. Ocelli all pale. Prosternum as in older stages. Coxal pores very small: 2, 2, 2, 2. c q 0, 0, 0, 0, 1, q 0, 0, 0, 0, 1 S Spines of first legs, o-o:0:0-1; of the second, ppg; of the third MOE elenT 0, 0, 2, 1, 0, 1 0, 0, 2, 1, 0 to ninth, 0, 0,0, 1, a et ane tenth, aie is Be of the elev enth, 0,0, 1,1, 13 of the twelfth, [a3 43 of the thirteenth, }}-?14; of the penult, Ti3i1; of the ana al, 9 35. None of the posterior coxae as yet laterally armed. Gonopods of female biarticulate. No claw indicated but one ready to have appeared with the next moult showing through integu- ment in cleared specimen. No bristles present but a spine is indi- cated on proximal article as a small acute chitinous point. Length 5 mm. Pullus IV (Larva quarta).— Pale, for the most part unpigmented or nearly so. Head and antennae yellowish, also the caudal end of the body and slightly so the tarsi of some of the legs. Antennae composed of seventeen articles of which those between the third and the ultimate are very short and closely compacted. Ocelli 1 + 1; the anterior one the larger; both somewhat angular. Prosternum nearly as in older stages. Sinus v-shaped with sides a little excurved. 166 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. One coxal pore present on coxa of each twelfth leg. Twelve pairs of developed legs present, with the remaining three pairs appearing as slender buds. Spines of first to third pair of legs, jjtu1; of fourth to ninth, oot; of tenth to twelfth, OT Anal glands distinct and well developed. Length 2.8 mm. Pullus III (Larva tertia).— Very pale and almost without pigment excepting the head, prehensors, and antennae which are of yellow or orange cast. Antennae composed of fourteen articles. Ocelli 1 + 1 of which the anterior is the larger. Prosternal teeth small, acute; line of apices slightly recurved. Sinus strictly v-shaped. Pairs of developed legs ten, behind which are two pairs of slender buds which distad of the coxa are unsegmented. Spines of first legs, 0,0, 0, 0, 1 0,0, 0, 0,1 . . 0-00-03 Of the second to seventh, ¢o-o-113 of the eighth and ninth, 0, 0, 0, 0, 0 0,0, 0,.0, 0 iO OLA; Wok the tenthig-p--at0- Anal glands well developed. Length 2.5 mm. Typr.— M. C. Z., No. 345, Alaska: Forrester Island. Harold and R. W. Heath. ParatypPEes.— M. C. Z., No. 346-349, 418, Alaska: Forrester Island. Harold and R. W. Heath. This species is readily recognized through the absence of a dorsal spine on the fourth joint of the anal legs. The only other known western lithobiid having this characteristic spining in the anal legs is Oabius tabiphilus, from Santa Barbara. PAOBIUS COLUMBIENSIS, sp. Nov. Plate 4, fig. 4. Diacnosis.— First dorsal plate 1.5 times wider than long; head equal in length and breadth. Body and head above light yellowish | brown. Ocelli ten to twelve in two or three series. Prosternum with distance between chitinous spots four times the dental line; sinus widely v-shaped. Two lateral lobes of genital forceps equal, a little smaller than the median; basal spines long and slender, nar- CHAMBERLIN: LITHOBIID GENERA. 167 rowest near middle. Coxal pores 2, 3, 3, 3. Last pair of coxae laterally armed. Fifth joint of sixth to ninth pairs of legs with but one ventral spine. Spines of anal legs, dorsal, 1, 0, 3, 1, 0; of penult, dorsal, 1, 0, 3, 1, 0, ventral, 0, 1, 3, 3, 1; of thirteenth, ventral, Oia o.e2; OFskOROl 2253, 2eldorsal, 0,-3.5 11s cof twelfth; dorsal, O31, &otnrst ventral, OF031,1, 1 or 0; 0, 15:251,-dorsal: 0) O45 1, 1. Length 7-8 mm. DescriptTion.— Dorsum light yellowish brown or orange-brown, with head not darker in types. Antennae yellowish brown. Poste- rior legs bright yellow. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 52 : 44:46 :59:61:58. Head precisely equal in length and breadth. Antennae of moderate shortness. Articles short, decreasing regu- larly distad in the usual way. Ultimate article considerably longer than the two preceding taken together but shorter than the. three preceding. Ocelli ten to twelve in two or three series: e. g., 1 + 4, 4,3; 1+ 5, 4, 1; 1+.5, 5. Single ocellus but a little larger than the most caudal ones of series. Other seriate ocelli mostly very small. Deeply pigmented. Prosternum 1.6 times wider than long. Distance between chiti- nous spots 2.5 times width at sinus and 4 times length of dental line. Teeth small and rather close, the interval between those of each pair semicircular. Line of apices very nearly straight. Spine close to base of outer tooth, bending mesocephalad over it, of usual character. Sinus widely v-shaped, the interval between the two mesal teeth being greater than that between teeth of each pair. First dorsal plate moderately narrowed caudad of middle in front of which the sides are subparallel. 1.5 times wider than long. Poste- rior angles of ninth, eleventh, and thirteenth dorsal plates straight. Coxal pores 2, 3, 3, 3. a of first legs, ¢ Or poi; Of the second to eighth, 0, 0, 3 0, 0 3, 2 0, 0,.3, 2, 2 Oot; of the he vured; of the tenth and eleventh, > 02,333 of the twelfth, (7333; of the thirteenth, ees OF Peas; Of the penult, ¢73:3:4; of the anal, 7335. Last pair of coxae laterally armed. Claw of female gonopods tripartite, the two lateral lobes equal and at same level, a little smaller than the median, all rather blunt. Spines 2 + 2 or 2+ 3; long and slender, broadest at base and nar- Una hs bal OF ON Oso 168 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. rowing to near middle above which again widening to beginning of the acuminate tip which is of moderate length; inner spine curving somewhat mesad (Plate 4, fig. 4). Length 7 to 8 mm. Typr.— M. C. Z., No. 350, British Columbia: Kaslo. PaRATYyPEs.— M. C. Z., No. 417, British Columbia: Kaslo. PAOBIUS OROPHILUS, sp. nov. DiaGnosis.— First dorsal plate 1.7— times wider than long; head wider than long. Brown; head much darker, nearly black. Ocelli about thirteen in three series. Prosternum with distance between chitinous spots four times the dental line; sinus v-shaped. Genital forceps bipartite or tripartite with outer tooth small; basal spines long, inner as long as or a little longer than outer. Coxal pores 2, 3, 3, 3. Last pair of coxae laterally armed. Fifth joint of sixth to thirteenth pairs of legs with two ventral spines. Spines of anal legs, dorsal, 1, 0, 3, 1, 0; of penult, dorsal, 1, 0, 3, 1, 0; of thirteenth, ventral, 0, 0, 3, 3, 2, dorsal, 1, 0, 3, 1,1; of twelfth, dorsal, 0, 0, 3, 1, 1; of first, ventral, 0, 0, 1,2, 1, dorsal, 0, 0, 3, 2,1. Length 9 mm. Description.— The dorsum in type is uniform brown. Head much darker, blackish with no tinge of red, uniform. Antennae like head excepting the tips which are light, yellowish. Venter, proster- num, and prehensors light brown or testaceous. Legs like venter, the caudal pairs a little darker, uniform. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 52 : 42 : 47:56:57 :55. Head wider than long in ratio 52 : 50 or 52 : 49. Antennae short, rather stout and only moderately attenuated distad. Articles short, each a little clavate. Ultimate article a little longer than the two preceding taken together, as usual. Ocelli in type thirteen in three series: thus, 1 + 5, 4, 3. Single ocellus distinctly largest, circular. Other ocelli large, distinct, and regular. Ocelli deeply pigmented excepting the single ocellus and the most caudal one of the uppermost series. Prosternum in type 1.53 times wider than long. Each side of anterior portion conspicuously inbent between spine and caudal or end or that adjacent to prehensor. Distance between chitinous spots 2.85 times width at level of bottom of sinus; 4 times the dental line. CHAMBERLIN: LITHOBIID GENERA. 169 Teeth small, acute, the inner of each pair the larger and the line of apices thus very nearly straight. Sinus strictly v-shaped. Spine as usual. First dorsal plate moderately narrowed caudad; 1.68 times wider than long. Caudal margins of ninth, eleventh, and thirteenth dorsal plates rounding forward at lateral ends, but not truly obliquely truncate. Coxal pores very uals z Bayt Spines of first legs, §-0:73:7; of the second, coed; of the third, Oete4; of the fourth and fifth, na a3 of the sixth to eighth, 0 0,3,.2,1 2,3, 29 Qo132; Of the ninth to eleventh, [:7:3:3:3, the dorsal spine of tibia being on the anterior side instead of on the posterior as more usual; of the twelfth, ¢:y332; of the thirteenth, j-}:33:3; of the penult, 3, 1,0 nee . 3,1, 0 Osa claws three, all distinct; of the anal, On i320- Only anal coxae laterally armed. Claw of female gonopods long and strongly curved, not excavated, thin and rather slender; bipartite, with the lobes equal, acute or tripartite, a smaller outer claw being thus present. Spines long and slender, equal in length or the inner a little the longer; in ventral view the inner spine is acuminate from base and is distally acute, but the outer is narrowed about one third the distance from the distal end toward which it is a little expanded and which is truncate or at middle a little incised. Length of type 9 mm. Typr.— M. C. Z., No. 351, British Columbia: Kaslo. ParatTyPes.— M. C. Z., No. 416, British Columbia: Kaslo. This species differs from P. columbiensis, occurring in the same locality, in larger size, coloration, spining of part of legs, different proportions of head, form of prosternum and especially in having the claw of the female gonopods more slender and curved and_ only bipartite, with the basal spines of a distinctly different form. PAOBIUS VAGRANS, sp. nov. DtaGnosis.— First dorsal plate 1.65-+ times wider than long; head wider than long. Dorsum yellowish brown with head chestnut or cherry-red. Ocelli six to nine in two or three series. Prosternum with distance between chitinous spots 4— times the dental line; 170 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. sinus large, v-shaped. Claw of gonopods tripartite; outer basal spine broad, much larger than the inner. Coxal pores 2, 3, 3, 2 to 3, 4, 4, 3. None of the coxae laterally armed. Third joint of first ten or more pairs of legs with but two dorsal spines. First seven pairs of legs with but a single dorsal tibial spine. Spines of anal legs, dorsal, 1, 0, 3, 0, 0 or 1, 0, 2, 0, 0; of penult, dorsal, 1, 0, 3, 1, 0 or 1,0, 2, 1,0; of thirteenth legs, ventral, 0, 1, 3, 3, 1, dorsal, 1, 0, 2, 1, 1; of twelfth, dorsal, 0, 0, 3, 2, 2 or 0, 0, 3, 1, 2; of first, ventral, 0, 0,1, 1,1 or 0, 0, 2, 1, 1; dorsal, 0,0, 2,2, 1. Length, 7.5-9 mm. Description.— Dorsum yellowish brown, with some of the plates occasionally showing a greenish tinge. Head from somewhat ferru- ginous to chestnut and cherry-red in individuals in full color, the first dorsal plate also more or less darkened like the head. Antennae colored like the head proximally, yellowish or orange distally. Venter a little paler than dorsum, sometimes in part showing a greenish tinge. Prosternum and prehensors a little paler than the head. Legs pale brown or brownish yellow, the caudal pairs more densely pigmented, bright orange-yellow. Body considerably narrowed cephalad from the tenth plate, with the first plate much narrower than the head but only slightly or not at all narrower than the third. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 48 : 43 : 43+: 52: 00 : 04. Head wider than long in about ratio 48:45. Caudal margin widely incurved. Lateral breaks may be more or less vaguely indi- cated. Antennae very short, the articles decreasing regularly from second to the penult, short and very short. Ultimate article distally acumi- nate, longer than the two preceding taken together. Ocelli six to nine in two or three series: e. g., 1 + 3, 2; 1+ 3, 3; 1+ 3, 2,1; 1+ 4, 3,1. Single ocellus distinctly largest. Prosternum with teeth equal, the line of apices a little recurved. Incision of good size, of the usual v-shape. Spine stouter at base than the bristle, slenderly acuminate and bristle-like distad, inserted in usual position. In type 1.54 times wider than long. Distance between chitinous spots 2.6 times width at level of bottom of incision; 3.9 times length of the dental line. First dorsal plate with sides convex anteriorly and caudally, sub- straight between, considerably converging caudad; 1.65 times wider than long. Caudal margin of ninth, eleventh, and thirteenth dorsal plates straight, the corners not excised or but indistinctly so. CHAMBERLIN: LITHOBIID GENERA. Lil Coxal pores small: 2,3 Dp tovoonaw ayaa 1. 44: 3. ovo aiora, Spines of first legs, = i" H iq OF fost of the second, oot or Soeei; of the third, (RTS, ooesT OF Goeet; of the fourth, Coes OF ooosi; Of the th ©0221, of the sixth, 22221 of 20221, of the seventh, S222, of the eighth, LAE on RAER2, of the ninth, (0:33:33 of ee tenth, ° oss; Of the eleventh, $2242: of the twelfth, (°f°33:2; of the thirteenth, & r 334; of the penult, E337 OF gtsa4, Claws three; of the anal, Ese OF eee, the coxal spine sometimes doubled, claw one. None of the coxae laterally armed. Claw of female gonopods short and relatively wide, well curved, tripartite, the median lobe longest, all acute. Outer basal spine of each pair longer and much wider than the inner with its sides proximad of the apical portion straight and parallel; apical portion moderate in length and acuteness, weakly denticulate on outer edge or also on mesal. Inner spine in ventral view acuminate from near base, denticulate along ectal edge. Length from 7.5 to 9 mm. Type.— M. C. Z., No. 352, Vermont: St. Johnsbury. R. V. Chamberlin. ParatypEs.— M. C. Z., No. 412, Vermont: St. Johnsbury; M. C. Z., No. 414, Lake Carmi. R. V. Chamberlin. This species is aberrant, especially in the character of basal spines of the female gonopods and in lacking lateral coxal spines. AREBIUS, gen. nov. Head with the usual marginal interruptions. - Antennae short; composed of twenty articles. Eyes composed of moderate or large ocelli arranged in two or three or occasionally in four longitudinal series. Single ocellus clearly differentiated and conspicuously larger than the others. Prosternal teeth 2 + 2; line of apices recurved; sinus v-shaped; spines slender and distally bristle-like. Posterior angles of ninth, dorsal plate straight or obliquely some- what excised; of the eleventh and thirteenth from somewhat obliquely truncate to straight or very weakly extended caudad. Coxal pores small and circular; uniseriate. 1¥2 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Claw of female gonopods either large and strictly entire (Hespero- bius sens. str.); or smaller and bipartite or tripartite with the main lobe or lobes greatly exceeding the lateral one or ones (Pagobius). Basal spines 2 + 2, long and mostly moderately slender. Tarsi of anterior legs slender with division indicated by an abrupt narrowing evident especially on the ventral side and by a paler, less chitinous transverse line or narrow band, and sometimes in addition by a mostly partial suture. Posterior legs comparatively slender in both sexes; without special modification in the male. Last one or two pairs of coxae laterally armed. From six to eleven last pairs of legs with the third joint bearing three dorsal spines. Either all legs from the first to the eleventh, twelfth, or thirteenth, or all excepting the first one, two, three, or four pairs bearing two dorsal spines on the fifth joint. Usu- ally the first five to eight pairs of legs, but rarely only the first two pairs, with but a single ventral spine on the third joint. Dorsal spines of anal legs, 1, 0, 3, 1,0; ventral 0, 1, 3, 2,0 (normal in Pago- bius) to 0, 1, 3, 2,1 or 0, 1,3, 3,1 (Arebius sens. str.); claws either one or two. Dorsal spines of penult legs 1, 0,3, 1,1; ventral, 0, 1,3, 3,2; claws three, the anterior commonly short but rather stout and somewhat truly claw-like. Dorsal spines of thirteenth legs Ors, Ato dO, 3) 262 ventral, 0.13, a2, 08-040; 3. 32. 5 Dorsal spines of twelfth legs from 0, 0, 3, 1, 1 and 1, 0, 3, 1, 1 to 0, 0, 3, 1, 2 and 1; 0,3, 2,2; ventral! 0, 0,3, 3,2 or rarely 0,0,2,3,2: Dorsal spines of eleventh legs 0,0, 3, 2,2; ventral, 0,0,2,3,2. Dorsal spines of first legs 0, 0, 2, 1, 1 or less commonly, 0, 0, 2, 2, 1; ventral, OO 2 or 0s OF Si le Length’ 12 mm. and less. Typre.— A. medius, sp. nov. The species of this genus at present known occur in California with the exception of A. oregonensis, the types of which were taken at Portland, Oregon. They are forms of small or medium size, from brown to chestnut in color, in which the body is rather conspicuously narrowed cephalad from the tenth plate, with the head proportionately rather small. The species fall into two natural groups or subgenera; Arebius sens. str., embracing species in which the claw of the female gonopods is large and strictly entire and the ventral spines of the anal legs normally 0,1,3,2,1 or occasionally 0,1,3,3,1; and Pagobius, subgen. nov., with P. diplonyx, sp. nov. as the type, embracing species in which the claw of the female gonopods 1s smaller and either bipartite CHAMBERLIN: LITHOBIID GENERA. We or tripartite, one or two small teeth appearing on sides of the main claw, and the ventral spines of the anal legs normally 0, 1, 3, 2, 0. Sometimes immature specimens of species of Arebius sens. str. (e. g. A. obesus) have minute lateral teeth near the base, but these invari- ably disappear by the time the maturus stage is reached. 'The small- ness of the lateral tooth or teeth in the claw of the gonopod of species of Pagobius and the proximal position of the same clearly indicate a transition to the condition of integrity normal in Arebius proper. Key to Subgenera and Species of Arebius. a. Claw of @ gonopods strictly entire; ventral spines of anal legs normally 0, 1, 3, 2, 1 or 0, 1, 3, 3, 1..Arebius sens. str. b. Only the last pair of coxae laterally armed. A. oregonensis, sp. Nov. bb. Last two pairs of coxae laterally armed. c. Anal leg with the claw single.......... A. obesus (Stuxberg) cc. Anal legs with two claws. d. Ventral spines of anal legs 1, 3, 3, 1; only the first two pairs of legs with third joint bearing but a single ventral spine, the others with 2 or 3....A. elysianus, sp. nov. dd. Ventral spines of anal legs 1, 3, 2, 1; first 7 pairs of legs with the third joint bearing a single ventral spine. A. medius, sp. nov. aa. Claw of 2 gonopods bipartite or tripartite; ventral spines of anal legs normally. 1,3,2,0........... Pagobius, subgen. nov. bur Claw. ‘of fanalleasimgle i. 2 hae FO. <2 A. dolius, sp. nov. bb. Anal leg with two claws. c. Claw of 2 gonopods bipartite; basal spines slender, narrowed from base distad; head wider than long; first dorsal plate cir. 1.8 times wider than long....A. kochii (Stuxberg) cc. Claw of Q gonopods tripartite; basal spines stout, only slightly or not at all narrowed from base to the acuminate apical portion; head equal in length and breadth; first dorsal plate cir. 1.6 times wider than long. A. diplonyx, sp. nov. BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 174 Fig. 4.— Distribution of Arebius. CHAMBERLIN: LITHOBIID GENERA. 175 AREBIUS OBESUS (Stuxberg). Plate 7, fiz. 3, 4; Plate 8, fig. 1. Lithobius obesus, Stuxberg, Ofvers. K. vet.-akad. Férhandl., 1875, 32, no. 2, p. 67; Ann. mag. nat. hist., 1875, ser. 4, 15, p. 189. Chamberlin (in part), Pomona college journ. ent., 1910, 2, p. 371, 373. Can. ent., 1911, 43, p. 380. Lithobius (Archilithobius) obesus Stuxberg, Ofvers. K. vet.-akad. Foérhandl., 1875, 32, no. 3, p. 18, 31; Proc. Cal. acad. sci., 1877, 7, p. 138. Now Lithobius obesus Bollman, Proc. U.S. N. M., 1889, 11, p. 347. Dracnosis.— Slender; first dorsal plate 1.5 + times wider than long; head as long as wide. Brown; head, and sometimes posterior plates, reddish. Ocelli eight to fourteen in three series. Prosternum 1.6 + times wider than long; distance between chitinous spots four times the dental line; sinus v-shaped. Claw of gonopods large, entire; basal spines moderate, acuminate from base. Coxal pores 2, 3, 3, 2 to 4, 4, 4, 4. Claw of anal leg single. Last two pairs of coxae laterally armed. First eight pairs of legs with third joint bearing but a single ventral spine. Spines of anal legs, ventral, 0. tea; 25 1 cot thirteenth; dorsal, 1, 0, 3, 1,. 1, ventral, 0, 1; 3,.3, 2: of twelfth, dorsal, 1, 0, 3, 1, 1, ventral, 0, 0, 2, 3, 2; of first, dorsal, 0, 0, 2, 2,1. Length, 9 to 12 mm. DescrietTion.— Dorsum brown, the caudal plates sometimes with tendency toward reddish. Head of a similar brown color or in some more reddish, the frontal region paler. Antennae brown, but little paler at tips. Prosternum somewhat paler than head with the pre- hensors a little lighter. Venter testaceous or yellowish with the caudal plates darker. Legs pale excepting the posterior pairs which are darker in correspondence with the ventral plates. Body rather slender, usually from 8.5 to 9.25 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 30 : 28 : 29 : 33:34 : 33. Head as long as wide or slightly longer; caudal margin mesally slightly incurved. Obscurely punctate. Antennae short, reaching the fifth body-segment. Articles short, sides nearly straight, each article a little clavately widening distad. Each eye consisting of from eight to fourteen ocelli arranged in three series: ¢. g., 1 + 3, 3,1; 1+ 4, 3,1; 1+ 4, 3,2; 1+ 4, 4, 2; 1+ 5, 4,3; 1+ 5, 5,3. Single ocellus much larger than any of the 176 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. seriate ocelli, the most caudal ocellus of each of the two upper rows also commonly large, others decreasing in size cephalad and ventrad. Prosternum about 1.62 times wider than long. Distance between chitinous spots near 2.36 times width at level of bottom of sinus; 4 times the length of the dental line. Teeth small, acute, dark; line of apices a little recurved. Sinus v-shaped. Spines slender and distally bristle-like. First dorsal plate 1.55-+ times wider than long. Sides caudad more than usually strongly rounded and converging about caudal corners, cephalad only gently convex and diverging. Caudal corners of minor plates and of the seventh obliquely excised. Plates smooth and shining. Coxal-pores very small, circular: 2, 3, 3, 2; 3, 3, 3, 3; 3, 4, 4, 3; 4, 4, 4, 4. : ONO Zoid 0,0; '2..2,.0 0, 2, 2, 2 Spines of first legs, g-o-as4; of the second, g-oPo4q OF gets o 0, 0, 2, 2,.2 25 , 0, 2, 2, 2 of the third ae fourth, ¢oioa;. of the fifth, ¢oaS4 or peaes 2, 2 0, 0, 2, 2, 2 0, 0, 3, 2, 2 0, 0, 2, 2, 2 of the sixth, (¢¢34 or foe tie LO Goss; Of the poeane 0-0: 122 0, 0, 3, 2, 2 0,0, 3,22 0,0, 3, 2 Or g-o122; Of the eighth, ¢o-7°5'5; of the ninth to eleventh, 9-335; of the twelfth, (33:5; of the sper tae cres2; of the penult, anit 0, 3,1 ‘ 13-373, Claws three; of the anal, j tae Claw one. Last two pairs of coxae laterally armed. Claw of female gonopods (Plate 7, fig. 4) moderately large and rather strongly curved; entire, with no trace of lateral teeth, acute. Basal spines (Plate 7, fig. 3) 2+ 2 or 2+ 3; of moderate length, slender, acuminate from near base; the plane of dorsal excision long. Mesal side of first joint (Plate 8, fig. 1) diverging considerably from its fellow cephalad. Process of sternite truncate. Length mostly 9 to 12 mm. A male 11 mm. long has antennae 3.5 mm. in length and the tenth dorsal plate 1.24 mm. wide. Immaturus (o).— Head slightly longer than wide as in maturus. Antennae with the full number of articles present but those of distal half, especially, relatively shorter than in adult. Ocelli of same form and arrangement as in maturus, the number being also nearly the same. Prosternum essentially as in adult but distance between chitinous spots may be as much as 3 times width at level of bottom of sinus. Coxal pores, 2, 2, 2, 2. 0,0, 1, 2,1 Spines of first legs, g-p:p-34; of the second, i; of the third 0, 0, 2, 2,1, and fourth, (}434; of the fifth to eighth, 73 22, of the ninth, CHAMBERLIN: LITHOBIID GENERA. ura? ON S022 0; 0, 3, .2,.2 3, 2, 2 y foe; of the tenth, f+ iss; Of the eleventh, (233; of the 0,0, 3, 1, 1 : 1, 0, 3, 1, 1 pt dit. twelfth, 9-95-33; of the thirteenth, jp-33-5; of the penult, 973353; of the anal, 2.0.3.1. Last two pairs of coxae later alhy armed. Specimen described from Stanford. Pullus IV (Larva quarta).— Very pale, yellowish brown, apparently violaceous in life. Head yellow of a dilute orange or ferruginous tinge, paler in front of suture. Antennae pale brown, lighter at tips. Prosternum nearly like head. Venter pale. Legs white or nearly so proximally, the tarsi with some yellow pigment. Head longer than wide (29 : 28). Antennae composed of seventeen articles, of which those between the third and the ultimate are very short, and more or less moniliform. Ocelli 1 + 2, 1 single ocellus already largest. Prosternal teeth small and pale. Prosternum 1.53 times wider than long. Distance between chitinous spots 2.56 times width at level of bottom of sinus; 4.1 times length of dental line. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other as 28 : 23 : 25 : 27 :25 ; 22. First dorsal plate conspicuously narrowed caudad; 1.6 times wider than long. Coxal pore one on each twelfth coxa. . 0,0, 0,0 0, 0, 0, 0, 0 Spines of first oe ooo; Of the second to ninth, TOOL; Of the tenth, Soa to ori; of the eleventh and twelfth, 2¢9%?. The spine on femur decreases in length cephalad. Length of specimen described 4 mm., with antennae 1.5 mm. long. Type Locauiry.— California: Sausalito. Also taken in California at Stanford, (M. C. Z., No. 354, 445, 447); Santa Barbara, (M. C. Z., No. 449); Fresno, (M. C. Z., No. 448). R. V. Chamberlin. AREBIUS MEDIUS, sp. nov. Plate 7, fig. 1. Dtacnosis.— First dorsal plate 1.45 times wider than long; head as long as wide. Dark brown to chestnut with head blackish. Ocelli eight or ten in two or three series. Prosternum 1.5 times wider than long; distance between chitinous spots 32 times the dental line; sinus v-shaped. Claw of female gonopods large, acute, entire; basal spines long and stout (Plate 7, fig. 1). Coxal pores 2, 2, 2,2 to3,4,4,3. Anal 178 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. leg with two claws. Last two pairs of coxae laterally armed. First seven pairs of legs with third joint bearing but a single spine. Spines of anal legs, ventral, 0, 1, 3, 2, 1; of thirteenth, ventral, 0, 1, 3, 3, 2, dorsal, 1, 0, 3, 1, 1; of twelfth legs, ventral, 0, 0, 2, 3, 2, dorsal, 0, 0, 3, 1, 2; of first legs, dorsal, 0, 0, 2, 1,1. Length 8-11 mm. Description.— Dark brown to chestnut, shining. Head darker, blackish. Antennae chestnut to blackish, light distad: Prosternum and prehensors dusky brown, the prehensors lighter, especially distad. Venter lighter, brown to light chestnut, the caudal plates sometimes darker. Legs brown, the caudal pairs darkest, sometimes irregularly dusky; tarsi of all legs paler, testaceous. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 60 : 55 : 60 : 69 : 69 : 68. Head subrotund, broadly rounded anteriorly; caudal margin mesally a little incurved. ‘Widest between eyes and marginal breaks. Equal in length and breadth or slightly wider. Antennae and its articles of intermediate length. Articles .cylindri- cal, those of distal portion becoming slender. Ultimate article a little clavately widening distad. Ocelli in specimens examined mostly eight to ten in two or three series: e. g., 1+ 4, 3; 1+ 4, 8, 2, which is the much commonest number and arrangement. Single ocellus much the largest. Most caudal one the largest of seriate ocelli. Organ of Témésvary in outline a little smaller than the ocelli of lowermost series, situated ventrad of anterior end of eye patch. Teeth of prosternum of moderate size, acute, uniform; the line of apices a little recurved. Sinus v-shaped, sides a little concave, of intermediate size; distance between two mesal teeth less than that between teeth of each pair. Spine slenderly acuminate, curving mesocephalad over ectal tooth. 1.5 times wider than long. Distance between chitinous spots 2.2 times width at level of bottom of sinus; 3% times length of dental line. First dorsal plate 1.45 times wider than long; sides convex, con- siderably converging caudad. Posterior corners of ninth and eleventh dorsal plates obliquely excised or truncate, the line of truncation short, especially on Hirtea plate. Coxal pores:))25 2:2, 25) 2, /dtio5 leno, 45 4oe Spines of first legs, p'p'j'24; of the second, §-j:73:4; of the third to Us ores; of the eighth, ¢¢¢33; of the ninth to eleventh, (7333; of the twelfth, (0343; of the thirteenth and CHAMBERLIN: LITHOBIID GENERA. 179 TONS eI i u 0 penult, 9-7-3373; of the anal, 97 334, claws two. Last two pairs of coxae laterally armed. Claw of female gonopods large and strongly curved, entire; acutely pointed. Mesal side of first article straight, well chitinized and keel- like. Basal spines long and stout, the outer a little longer than the inner; both gradually acuminate from base distad, the tip very nar- rowly rounded (Plate 7, fig. 1). Length 8 to 11 mm. Typre.— M. C. Z., No. 356, California: Brookdale, March, 1913. R. V. Chamberlin. Paratypres.— M. C. Z., No. 438, California: Brookdale, March, 1913. R. V. Chamberlin. AREBIUS ELYSIANUS, sp. Nov. Plate 7, fig. 2. Diacenosis.— First dorsal plate 13 times wider than long; head slightly wider than long. Testaceous, with head darker reddish brown. Ocelli eight to twelve in three series. Prosternum 1.5 + times wider than long; distance between chitinous spots 3.6— times dental line. Claw of gonopods long, entire; spines moderate in length. Coxal pores 2, 2, 2, 2 to 3, 3, 3, 3. Anal leg with two claws. Last two pairs of coxae laterally armed. Third joint of all legs excepting the first two pairs bearing two or three spines. Spines of anal legs, ventral, 0, 1, 3, 3, 1; of thirteenth legs, ventral, 0, 1, 3, 3, 2, dorsal, 1, 0, 8, 2, 2; of twelfth legs, dorsal, 1, 0, 3, 2, 2, ventral, 0, 0, 3, 2, 2; of first legs, dorsal, 0, 0, 2, 2, 1. Length 8-12 mm. Descrietion.— Dorsum light yellowish or testaceous brown. Head dark brown of reddish cast, always distinctly and considerably darker than the dorsum; frontal region paler. Antennae like head, reddish brown, with tips in some degree paler. Prosternum some- what lighter than head. Venter yellow or pale testaceous, caudal plates a little darker, usually somewhat reddish. Legs pale, the posterior pairs more deeply pigmented, mostly yellow or of a some- what orange tinge. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 32 : 30:31 : 37 : 37 :35. Head slightly wider than long (32 :31). Caudal margin straight, 180 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. / otherwise head is suborbicular or sides somewhat flattened caudad of eyes. Antennae short; composed of the usual twenty articles which are short; articles subcylindric or a little widening distad. Ocelli eight to twelve in three series: e. g., 1 + 3, 3,2; 1+ 4, 3, 2; 1+ 4, 4, 3. Single ocellus largest. Of the seriate ocelli the caudal one of first row and of the second are largest, those of lowermost row smallest. Prosternum in type 1.5+ times wider than long. Distance between chitious spots 2.27 times width at level of bottom of sinus; 3.57 times the dental line. ‘Teeth moderately small and acute with sides a little concave. Sinus rather shallow in proportion to its width; v-shaped but with sides a little concave. Spine moderately stout at base, becoming bristle-like distally. First dorsal plate 13 times as wide as long; widest anteriorly, the sides rather conspicuously converging caudad. Minor plates mostly with the caudal margin straight entirely across width or with the caudal corners a little rounded or occasionally one corner excised. Coxal pores small and circular, nosey well separated, few in number: ¢..9:,2, 2,2, 2; .2,3,0, 2; 2, 23,2; 2,3, 3,09; 8), 35a * 0, if Zia, 0; '2;.2,.2 * Spines of first legs, (:y-q3:4; of the second, ¢°:73:4; of the third, to 2, 2 s 0, 0, a 2,2 fifth, (-os:33; of the sixth to eleventh, (o-s33; of te twelfth, 2,2 0,3, 2 1,.0,3,1 roses; of the thirteenth, ¢7:3:3; of the penult, 9y3+3> 5; of the 0. . anal, (3-3-1 Claws two. Last two pairs of coxae laterally armed. Gonopods of male slender; well exposed; somewhat obliquely sub- conically rounded distally and bearing apparently but a single bristle. Claw of female gonopods rather long and narrowly acutely pointed, moderately curved and entire with no indication of lateral teeth. Basal spines moderate in stoutness and length, acuminate regularly from base to apex; inner spine somewhat smaller than the outer (Plate 7, fig. 2). Inner side of first article weakly convex, only a little diverging cephalad from margin of its fellow. Length 8 to 12mm. A male 10 mm. long has antennae 3 mm. long and anal leg of same length or very nearly so. M. C. Z., No. 444, California: Los Angeles, Elysian Park, June, 1909. R.V. Chamberlin. Paratyprs.— M. €. Z., No. 355, 442, California: Pasadena; M. C. Z., No. 441, San Ysidro. W. M. Wheeler; M. C. Z., No. 443, Claremont. R. V. Chamberlin. CHAMBERLIN: LITHOBIID GENERA. 181 AREBIUS OREGONENSIS, sp. Nov. Plate 7, fig. 6. Diacnosis.— First dorsal plate 1.7 times wider than long; head as long as wide. Brown, head dark brown to chestnut. Ocelli ten to fourteen in four series. Prosternum with distance between chitin- ous spots only 33 times the dental line. Claw of female gonopods entire; basal spines long and slender, of uniform width, the outer but little longer. Coxal pores 4, 4, 4, 4. Only last pair of coxae laterally armed. Third joint of first five pairs of legs with a single ventral spine. Spines of thirteenth legs, ventral, 0, 1, 3, 3, 2, dorsal, 0, 0, 3, 1,1; of twelfth ventral, 0, 0, 2,3, 2. or 0, 0, 2, 3, 3, dorsal, 0, 0, 3,1, 2. Length 10-12 mm. DerscripTION.— Brown. Head darker, dark brown or chestnut. Antennae like head, uniform or paler distad. Prosternum and pre- hensors paler than the head. Venter brown with the caudal plates a little darker. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 75 : 69 : 72 : 88 : 90 : 84. Head as wide as long or slightly longer (75 : 77). Rather strongly narrowed caudad of marginal breaks; caudal margin mesally a little incurved. Antennae of moderate shortness. The articles of corresponding moderate length; usually not decreasing distad in usual way or only but few preceding the ultimate thus shortened, but sometimes those distad of third or fourth decrease in the ordinary way. Ocelli ten to fourteen in four series: e. g., 1+ 2, 4, 3, 1; 1+ 4, 4, 3,1; 1+ 4,4, 4,1. Single ocellus much exceeding the others in size. Seriate ocelli comparatively rather large, regular. Organ of Toémoésvary smaller in outline than the smaller ocelli, below the anterior end of which it is placed. Teeth of prosternum very small and acute, equal; line of apices recurved. Sinus large, v-shaped, with sides a little concave. Spine not close to outer tooth, not much stouter at base than the bristles, long and slenderly acuminate, nearly straight. Sides of anterior portion rounding out a considerable distance from the outer tooth in a direction but little caudad of directly ectad before extending caudad at the usual angle (Plate 7, fig. 6). Distance between chitinous spots only twice or slightly less times width at level of bottom of sinus; 33 times the dental line. 182 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. First dorsal plate 1.7 times wider than long; caudal corners widely rounded; sides but little diverging cephalad. Caudal margins of ninth and eleventh plates bent forwards at ends, that of the ninth the more strongly so. Thirteenth plate with the caudal margin curving caudad at ends, the corners being slightly extended. Coxal pores in type 4, 4, 4, 4. Ventral spines of first and second legs 0, 0, 1, 3,1. Spines of second to fourth legs, (y'q'3'3; of ihe a oi33; of the sixth to eleventh, OES of the twelfth, §:3:3:3:5 or on one side, oeeg3; of the thir- teenth, hate ; 33 5; Others lost in types. Only last pair of coxae in types laterally armed. Claw of female gonopods large, well curved, concave, entire. Inner side of first article straight. Basal spines long and slender, with the outer but little the longer; outer spine of nearly uniform width from base to the short acutely pointed apical portion. Process of sternite acutely pointed. Length 10 to 12 mm. Typr.— M. C. Z., No. 353, Oregon: Portland, 3 August, 1902. R. V. Chamberlin. ParatypEs.— M. C. Z., No. 439, Oregon: Portland, 3 August, 1902. R. V. Chamberlin. AREBIUS (PaGosius) KocHi (Stuxberg). Plate 5, fig. 6; Plate 6, fig. 1-3. Lithobius kochii Stuxberg, Ofvers. K. vet.-akad. Férhandl., 1875, 32, no. 2. p. 68; Ann. mag. nat. hist., 1875, ser. 4, 15, p. 190. Chamberlin (in part), Ann. Ent. soc. Amer., 1909, 2, p. 187; Pomona college journ. ent., 1910, 2, p. 371, 374; Can. ent., 1911, 43, p. 380. Lithobius (Archilithobius) kochii Stuxberg, Ofvers. K. vet.-akad. Forhandl., 1875, 32, no. 3, p. 18, 30; Proc. Cal. acad. sci., 1877, 7, p. 138. Non Lithobius kochit Bollman, Proc. U. 8S. N. M., 1887, 10, p. 254, 255; 1889, 11, p. 348. * Diacnosis.— First dorsal plate 1.8 times wider than long; head a little wider than long. Testaceous to chestnut, frontal region of head paler. Ocelli about ten in three series. Prosternum 1.6 + times wider than long; distance between chitinous spots 3.8 times dental line. Claw of female gonopods long, bipartite; basal spines attenu- ated from base, outer a little longer. Coxal pores 2, 2, 2, 2 to 3, 3, 3, 3, CHAMBERLIN: LITHOBIID GENERA. 183 small. Anal leg with two claws. Last two pairs of coxae laterally ‘armed. Third joint of first seven pairs of leg with a single ventral spine. Spines of anal legs, ventral, 0, 1, 3, 2, 0 or 0, 1, 3, 2, 1; of thirteenth, ventral, 0, 1, 3,3, 2; of twelfth, dorsal, 0, 0,3, 1,2. Length 7.5 to 11 mm. DeEscrIPTION.— Dorsum testaceous to dark brown, often varying toward chestnut; caudal margin of plates sometimes darkened. Head dilute to dark chestnut with the frontal region paler. Venter pale testaceous or very dilute brown, commonly showing anteriorly a somewhat violaceous tinge, while the caudal plates are darker as usual, burnt brown. Legs nearly like corresponding plates of venter; last pair brown above, paler distally and ventrally. Body strongly narrowed cephalad with head rather small and first plate narrower than third, the widths of head and of first, third, eighth, tenth, and twelfth plates being to each other nearly as 28 : 26 : 28 :34 :34:33. Body typically about eight. times wider than long. Head suborbicular, the sides being considerably convex; caudal margin mesally nearly straight. A little wider than long (29 : 28). Antennae very short, reaching only to the fifth segment of body. The twenty articles short and cylindric, rather uniform in proportions. Ocelli most frequently about ten in three series: e. g., 1 + 4, 3, 2. Decreasing in size cephalad and ventrad. Single ocellus typically much larger than any other, well differentiated (Plate 6, fig. 1). Prosternum (Plate 5, fig. 6) 1.63 times wider than long. Distance between chitinous spots 2% times the width at level of bottom of sinus; 3.8 times the dental line. Median sinus v-shaped with the sides straight; shallow, scarcely or not at all extending below level of chitinous dental plate. Teeth small with sides nearly straight, the inner a little larger than the outer and the line of apices but slightly recurved. Spine stout at base, narrowing to bristle-like form distally as usual. First dorsal plate 1.8 times wider than long; sides moderately con- vex and converging caudad, widest a little back of the anterior end. Minor plates mostly simply rounded at caudal corners, not obliquely truncate or excised. Coxzal pores small ):252)/2, 23) 2,335.23) 2.08, 35.33 3513; 3, 3. 0 0 —) ’ 0,0, 2, 2.1 00 3; Of the second, Spines of first legs, 0, 0,2,2,1 0,0, 2, 2,1 0,.0, 2, 2, 2, of the third, ONO SaN2 el OND , 1.3, 1 or 0, 0, 1, 3, 19 of the fourth, ° 0, oy Dali or 0,0, 2, 2, 2, of tl fifth |e ee of h ht} 0, 2, 2, 2 0.0.1.3.13 ne hith to seventh, 9°9°7°3:13 the eighth, ones 5 OF 184 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 0, 0, 3. BZ 2, 2A 4 2 0023-5; Of the ninth to eleventh, ° oases; of the twelfth, (93333; of ite thirteenth, ¢-7343; of the penult, 9-333; of the anal, iss with two claws. Coxae of last two pairs of legs laterally armed but the spines, especially of the penult pair, easily lost off and coxae SO appearing unarmed. Posterior legs short and moderately slender in female, being a little, uniformly, thickened. In male as in female without special lobes or furrows. Gonopods of male short and rather wide; distal edge straight, bearing two bristles. Claw of gonopods of female (Plate 6, fig. 3) rather long and strongly curved; bipartite, the inner or dorsal lateral lobe being wholly ob- literated, the ventral lateral lobe much smaller than the main one, both acute (Plate 6, fig. 2). Basal spines attenuated more or less uniformly from base to apex, the outer a little the larger. Length from 7.5 to 11 mm. A male 10 mm. long has antennae 3.6 mm. and anal leg 3.25 mm. long, with the tenth dorsal plate 1.24 mm. wide. Tyree Locauity.— California: Sausalito. Torotyrrs.— M. C. Z., No. 344, 437, California: Sausalito. R. V. Chamberlin. Also taken in California at Monterey and Pacific Grove, R. V. Chamberlin, and at Ukiah, Oregon. AREBIUS (PAGOBIUS) DIPLONYX, sp. nov. Plate 5, fig. 5: Plate, 6, fig. 73. 8: Diacgnosis.— Dorsal plate 1.54 to 1.58 times wider than long; head equal in length and breadth. Brown, first one and last several plates darker, head chestnut. Ocelli eight to ten in three series. Prosternum 1.5 to 1.58 times wider than long; distance between chitinous spots 3.53 to 3.8 times the dental line; sinus v-shaped, opening wide. Claw of female gonopods tripartite with median lobe much the longest; spines subcylindric. Coxal pores 2, 2, 2, 2 to 2, 3, 3, 3. Anal leg with two claws. Last two pairs of coxae laterally armed. Third joint of first seven pairs of legs with single ventral spine. Spines of anal legs, ventral, 0, 1, 3, 2, 0 or 0, 1, 3, 2, 1; of thirteenth, ventral, 0, 1, 3, 3, 2 or 0, 1, 2, 3, 2; of twelfth, dorsal, 0; 0, 3, 2; 2 or 0, 0, 3,1, 2. Length 8.5 to 10 mm. CHAMBERLIN: LITHOBIID GENERA. 185 DEscrIPTION.— Dorsum brown, with the first one and the last several plates commonly darker, more or less chestnut. Head chest- nut, nearly uniform or but slightly paler in front of the suture. An- tennae usually dark brown or chestnut like the head in individuals in full color, with the ultimate article alone or sometimes with more of the distal portion paler; in some the antennae are lighter throughout. Venter pale brown to almost colorless, sometimes suffused with purplish and at others having a greenish tinge, the posterior plates a denser, clearer brown. Prosternum testaceous and brown to chestnut, the prehensors paler, especially distally. Legs pale, with the tarsi more pigmented; the posterior pairs brownish, brighter testaceous or yellowish distally. Body mostly from 7.5 to 8.5 times longer than width of tenth plate; but one female 9 mm. long is but 6.4 times longer than the width of this plate. Body conspicuously narrowed cephalad with first plate clearly narrower than the third. Widths of head and of first, third, eighth, tenth, and twelfth plates in a female measured are to each other as 60 : 54 : 60:70 : 72 : 68, and in a male as 55 : 50 : 54 : 65: 64 : 60. Head precisely equal in length and breadth. Widest just back of eyes; sides considerably converging from the marginal breaks caudad. Antennae short. Composed of from twenty to twenty-three articles which are of moderate length or short and subcylindric in form, not closely compacted. When the number of articles is above the usual twenty, this is due to subdivision of some of those immedi- ately preceding the ultimate which are, as a result, very short. Ulti- mate article normally but slightly longer than the two preceding taken together. Each eye consisting usually of from eight to ten ocelli arranged in three series: e. g., 1 + 4, 3, 2, the most frequent arrangement and number; 1+ 4, 3,1; 1+.3, 3, 1. Single ocellus distinctly largest, it and the first one of the most dorsal series commonly of a distinct color cast. Prosternum 1.5 to 1.58 times wider than long. Distance between chitinous spots 2.4 to 2.65 times width at level of bottom of sinus; 3.53 to 3.8 times the dental line. Sinus v-shaped with the opening wide. Teeth small, uniform, well separated with the interval widely semicircular; line of apices nearly straight. Spine long, stout at base and slender distally as usual. First dorsal plate 1.54 to 1.58 times wider than long. Caudal margins of ninth, eleventh, and thirteenth dorsal plates straight, with 186 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. the corners rectangular or narrowly rounded, not at all obliquely truncate. Coxaltpores 2572.92) 2 to 2333 . 0,0,2, 1.1 0,0, 21,1 0,0,2,2.1 Spines of first legs, gg-o4 OF o-aai31; Of the second, 5-94 131) s 0,0;'2, 2 1 ONO ee Neale of the third, 99:23 OF o-o131; Of the fourth and fifth, oo7: Opies Ow, )2 0, 0, 1, 2 0 ONO? . 0, 0, 3, 2, 0, inet 5 ot the sixth, 0, 0, 1, 2, + Ea en Ea the ‘Gghih, COL 3 1 OF o-0-331; of 3 of 2 0, 0, 3, 1,2 we 1, 0,3, 1,4 of the twelfth, ¢-o3:35 or joes; of the thirteenth, ees OF 9 i 34; of the seventh, 3, 1, t he ninth to eleventh, ® 5, the spines of the trochanter small or minute; of the penult, 3 10; 0, 3, 1, 0 0: 8.11, of the anal, 7-733 or ¢ L321, Claws two with the acces- sory one moderately large and distinct. Last two pairs of coxae laterally armed. Gonopod of male very slightly protruding, bearing apparently but one bristle. Claw of female gonopods strongly curved and concave mesally; tripartite, with the median lobe much largest and the outer tooth often inconspicuous (Plate 6, fig. 8). Spines subeylindric, but slightly narrowed from the base to the acuminate distal portion which is moderate (Plate 6, fig. 7). Length from 8.5 to 10 mm. Typr.— M. C. Z., No. 342, California: Santa Barbara. R. V. Chamberlin. ParatTyPEes.— M. C. Z., No. 341, 434-436, California: Santa Barbara. R. V. Chamberlin. A considerable number of specimens of this species were collected by the author in March, 1913. AREBIUS (PAGOBIUS) DOLIUS, sp. nov. Plate 6, fig. 4-6. Diacnosis.— Body strongly narrowed cephalad. First dorsal plate 1.55 to 1.6 times wider than long; head longer than wide. Dorsum brown, darker at ends, head chestnut. Ocelli seven to nine, mostly in two series. Prosternum about 1.5 times wider than long; distance between chitinous spots 3.8 times dental line. Claw of female gonopods bipartite; basal spines narrowed from base, outer one considerably larger. Coxal pores 3, 3, 3, 3 to 3, 4, 4, 3. Anal leg with claw single. Last two pairs of coxae laterally armed. Third CHAMBERLIN: LITHOBIID GENERA. 187 joint of first eight pairs of legs with single ventral spine. Spines of anal legs, ventral, 0, 1, 3, 2, 1; of thirteenth legs, ventral, 0, 0, 3, 3, 2; of twelfth, dorsal, 1, 0, 3, 1,1. Length 9-11 mm. DescripTion.— Brown, with the first and one or more of caudal plates sometimes darker, more or less of chestnut tinge. Head chest- nut, often dusky, paler in front of suture. Antennae brown to chest- nut, lighter at tips. Prosternum chestnut like the head, with the prehensors somewhat lighter. Venter much lighter than dorsum, dilute brown of frequently greyish cast; caudal plates darker as usual. Posterior legs darkest, brownish, lighter at tips but not clear yellow. Body strongly narrowed cephalad with the head comparatively small, the widths of head and of first, third, eighth, tenth, and twelfth plates being to each other, in a male, as 54 : 52 :57 : 65 : 68 : 68 : 65 and in a female as 52 : 51 : 57 : 67 : 69 : 69. Head longer than wide, typically in about ratio 54 : 52. Antennae short; the twenty articles short, decreasing gradually distad from the second. The ultimate article normally distinctly longer than the two preceding ones together. Each eye commonly composed of from seven to nine ocelli in two series: e. g.. 1+ 4,3; 1+ 5, 3; 1+ 4, 2. Single ocellus (Plate 6, fig. 5) always distinctly much larger than any other, the first of the dorsal series being next in size and the others decreasing cephalad. Prosternum 1.5— times wider than long. Distance between chiti- nous spots 2.8 times width at bottom of median sinus and 3.8 times the dental line. Teeth rather small and acute; the line of apices distinetly recurved. Sinus strictly v-shaped. Spine much stouter than the bristles and acutely acuminate but ordinarily not strictly bristle-like at tip. First dorsal plate with sides moderately converging caudad to the rounded corners; 1.55 or 1.6 times wider than long. Minor plates with caudal corners simply rounded, not obliquely truly truncate. @oxalpores: small: 3/3, 3,07) 3,0;4,a; 3,4, 4, 3. Spines of first les O21; of the second to fifth, jos; of the » 0,0 jp Dy Qn 50,3, 2. 2 sixth to eighth, (3:33; of the ninth to eleventh, }}335; of the a (ip og AUR Eat : sy oh abe! twelfth, (335 or gosgs; of the thirteenth, ff335; of the 1, 0, 3,1, : penult, 5 aa 33:3; of the anal, ¢1:3:34, with the claw single. Last two pairs of coxae laterally armed. Claw of female gonopods bipartite, with the inner or more dorsal lobe much smaller than and much below apex of the main one, the outer lobe being wholly obliterated (Plate 6, fig. 6). Basal spines 188 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. distinctly narrowing from the base distad, the apical portion more abruptly acute; sometimes the sides are somewhat incurved toward middle of length; outer spine stouter and considerably longer than the inner (Plate 6, fig. 4). Length from 9 to 11 mm. Typr.— M. C. Z., No. 343, California; Friant, March, 1913. R. V. and §. C. Chamberlin. ParatyPes.— M. C. Z., No. 431, California: Santa Barbara, R. V. Chamberlin; M. C. Z., No. 432, 433, 450, Friant, March, 1913, R. V. and S. C. Chamberlin. ; . NOTHEMBIUS, gen. nov. Head with lateral marginal interruptions. Antennae short; composed normally of twenty-two articles, or in some individuals of but twenty-one. Eyes composed of ocelli in two or three seriés; single ocellus differ- entiated and enlarged. Prosternal teeth 2+ 2 or 3+ 3; line of apices, more or less re- curved; sinus v-shaped; spine slender, distally bristle-like. Posterior corners of ninth dorsal plate rounded; of eleventh straight or slightly rounded; and of thirteenth produced. Claw of female gonopods large; entire. Basal spines 2 + 2 or 2+ 3; stout, attenuated distad, with the outer of each pair con- siderably exceeding the inner. Tarsi of the anterior legs with a division more or less clearly indi- cated, the suture being mostly complete or nearly so. Anal and penult legs of male moderately thickened, the fifth joint of the former bearing on its ventral surface at distal end a more or less conspicuous pilose lobe. Last. one or two pairs of coxae laterally armed. Either all legs or all excepting the first one to first five pairs with the third joint bearing three dorsal spines and either two or three ventral ones; fifth joint of legs from first to twelfth, or all of these excepting the first one or two pairs with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral in female, 1, 3, 2, 1 and ina male either 1, 3, 2, 0 or 1, 3, 2, 1; claw one. Dorsal spines of penult legs 1, 0, 3, 1, 1; ventral, 0, 1, 3, 3, 2 or in individual exceptions 0, 1, 3, 3, 1; claws three. Dorsal spines of thirteenth legs 1, 0, 3, 1, 1 or 1, 0, 3, 1, 2; ventral, 0, 1, 3, 3, 2 or in exceptional individuals, 0, 0, 3, 3, 2. Dorsal spines of twelfth legs 1, 0, 3, 1, 2 or 0, 0, 3, 1, 2 to 189 GENERA. XN CHAMBERLIN: LITHOBIID 5.— Distribution of Nothembius. Fic. 190 BULLETIN MUSEUM OF COMPARATIVE ZOOLOGY. 0, 0, 3, 1, 2 or 0, 0, 3, 2, 2; ventral, 0, 0, 3, 3, 2. Dorsal spines of elev- enth) legs,0;0) 302,023 ventral.J0,/0.53,,3,/2\o0r-0.092,.3,.2. “Dorsal spines of first legs mostly 3, 2, 1 or 2, 2, 1, in individual exceptions sometimes only 2, 1,1; ventral nearly always 0, 0, 2, 3, 2, occasionally 030;.2.53, or 08022. 2) le Length between 8.5 and 13.5 mm. Typr.— JN. insulae, sp. nov. Three species of this interesting genus occur in California. They may be separated as follows. Key to Species of Nothembius. a. Prosternal teeth 3+ 3; ventral spines of anal legs in both @ DIGITS 4 Opaal” oyu 2a; meincccle.. oes N. aberrans, sp. nov. aa. Prosternal teeth 2 + 2; ventral spines of anal legs in o, 0, 1, 3, DAO wand Ov, Misa 22 Le b. Dorsal spines of third joint of first four to six pairs of legs only two in number; body typically 8.5 or more times longer than width of the tenth plate... .N. insulae, sp. nov. bb. Dorsal spines of third joint of all legs three in number or of the first alone rarely only two; body mostly only between seven and eight times longer than width of tenth plate. N. nampus, sp. nov. NOTHEMBIUS INSULAE, Sp. nov. Pilate 8, fig. 6; Plate 9, fig. 1-3. Diacnosis.— Slender; first dorsal plate 1.7 times wider than long; head wider than long. Dorsum brown to chestnut, head and first dorsal plate darker. Ocelli six to eight in two or three series. Pro- sternum 1.6 times wider than long; distance between chitinous spots 3.6 times dental line; teeth, 2-+ 2. Basal spines of gonopods acumi- nate from base. Coxal pores 3, 3, 3, 3 to 3, 4, 4, 3. Third joint of first four or five pairs of legs with but two dorsal spines. Last one or two pairs of coxae laterally armed. Spines of anal legs, ventral, in female 0, 1, 3, 2, 1, in male 0, 1, 3, 2, 0; of thirteenth, dorsal, 1, 073505 1; of twelith,-dorsal;, 1,0: 3,45, 2,07 1,:0,3..1,) Ja Loberor anal leg of male as in Plate 9, fig. 1. -Length 10 to 13.5 mm. DescripTion.— Dorsum brown to chestnut, typically the latter. Head and usually with it the first dorsal plate a little darker than the CHAMBERLIN: LITHOBIID GENERA. 191 dorsum of body with the region in front of suture usually paler and in some a black spot just back of the suture. Antennae chestnut, rufous distally. Prosternum dusky brown; prehensors testaceous. Venter brown, the caudal plates darker and more reddish as usual. Legs brown like the venter, the tarsi lighter, testaceous; posterior pairs more chestnut, the tarsi, or tibiae and tarsi, commonly lighter, testaceous. Body usually 8.5 or more times longer than width of tenth plate; Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 67 : 63 : 71 : 82 : 84: 78. Head suborbicular; wider than long in about ratio 67 : 63. Antennae short and strongly attenuated. Articles short, uniform, very regularly decreasing distad; twenty-two in number. Ocelli six to eight in two or three series: e. g., 1 + 3, 2; 1+ 4, 3; 1+ 3, 2, 1; 1+ 4, 2, 1. Single ocellus greatly exceeding in size the others, which are mostly small; first ocellus of uppermost row clearly the largest of the seriate ocelli, often paler than the others which decrease cephalad. Prosternum (Plate 9, fig. 3) 1.6 times wider than long. Distance between chitinous spots 2.26 times width at level of bottom of sinus; 3.6 times length of dental line. Sides of anterior portion round- ing out well from ectal teeth. Line of apices of teeth nearly straight. Sinus large, v-shaped; distance between mesal teeth somewhat greater than that between teeth of each pair. First dorsal plate near 1.7 times wider than long. Coxalipores.3,3,.0; 4; 3,3,4,3; 334, 4, 3. Spines of first legs, $-o:3:3'3 OF posae; of the second to fourth, tosses; of the fourth to fifth, P¢¢33 or EEE; of the sixth to eleventh, (03:33; of the twelfth, oss OF Geese Where the latter the anterior dorsal spine of tibia very small; of the Gattespth 1, 0,3,1,1. 1, 0, 3, 1,0 1, 0,3,1 and penult, 9-7°3:3:2; of the anal, Os 4 oes OF ae (ot); claw one. Either last pair or last two pairs of coxae laterally armed; when the last two the spine on penult coxa is very small. Anal legs of male with the fifth joint thickened and at distal end ventromesally excised and on plane of excision bearing an oblong lobe, the longitudinal axis of which is a little oblique to that of joint; the lobe expands distally above base and is flat or somewhat concave topped (Plate 9, fig. 1). Claw of female gonopods large, well curved, moderately excavated. Spines acuminate from base gradually to distal division and then more 192 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. abruptly acutely pointed. Outer spine always considerably longer than the inner (Plate 8, fig. 6). Mesal edge of first article diverging from the median line, proximally somewhat concave (Plate 9, fig. 2). Length 10 to 13.5 mm. Praematurus.— Color of dorsum decidedly green. Head and first plate light brown. Antennae light brownish, lighter distad. Pro- sternum like head. Venter green. Legs pale green, lighter, freer from green, distad; posterior pairs brighter, yellowish. Articles of antennae already twenty-one or twenty-two. Ocelli five in two series: 1-+ 3, 2. Single ocellus largest, pale; first of upper series next in size, also pale. General form and proportion of prosternum as in adult. Sinus between v- and u-shaped, large. Line of apices of teeth slightly re- curved. } Coxal pores 2, 2, 2, 2. 0, 0.1.2.1, Spines of first legs, goa-24; of the second, ¢'p'3-3-4; of the third, 0, 0, 2, 2, 2 0, 0, 2,2,2 002313 Of the fourth to sixth, 9-p-33-3; of the seventh to eleventh, ONO Bene i Oyeha 1, 0, 3,1,1, 0, 0, ogo; Of the twelfth, p34 m z el Ae thirteenth and penult, (-7°3°3°3; of the anal, ¢-7232 (2), org ally armed. Anal legs in male slender, the fifth joint not much thickened but showing at the distal end the lobe which may have nearly the char- acteristic form excepting for smaller size or which may appear in the form of merely a rounded, bulging eminence or knob not yet showing the characteristic shape. Claw of female gonopods moderately curved, rather stout, strictly entire. Basal spines two plus two, but the inner spine of each pair very much smaller, both spines acutely conical. Second and third articles each with three ventral bristles, the first with three or four. Length cir. 8.25-8.5 mm. aa Prevailingly green. Head and first plate same as dorsum or brownish with frontal paler. Antennae paler distad of middle. In green individuals the legs are somewhat whitish distad. Antennae with full number of articles, twenty-two. Ocelli 1-+ 2, 2. Well separated; slightly pigmented. Single ocellus and the caudal one of each series not much differing in size. Prosternum with same form and proportions as in adult. Coxal pores very small, 2, 2, 2, 2. . 0,0, 1, 1, 0,0, 2,21 Spines of first legs, i{:z74; of the second, p:joi1; of the third 0, 2, 2, 2 to fifth, 7744; of the sixth and seventh, [}¢ 734; of the eighth *(%). Last pair of coxae later- CHAMBERLIN: LITHOBIID GENERA. 193 , 0, 2, 2, 2 0,3, 2,2 0,3, 2 and ninth, §97'3:4; of the tenth, os OF tt iss; of the eleventh, 0,0, 3,1, 0, 0, 3, 1, 1 0, 0, 3, 1,1 ONO SS ed Cosi; Of the twelfth, Pp233; of the thirteenth, Oars tO OSes 0, 0, 3, 1,1, 1, 0, 3, 1,0 1, 0, 3,1, 0 of the penult, gis; of the anal, girso7 (2) or gigs (co). No lateral spines detected on any of the coxae. Length cir. 7.5 mm. Agenitalis I.— Color very dilute brown, with head and first dorsal plate a little darker. Legs pale, whitish. Antennae composed of twenty articles which are short and closely compacted, the ultimate being longer than the three preceding ones taken together. Ocelli 1+ 2, Lor 1+ 2,2. Single ocellus and caudal one of upper series equal or single one rather smaller. Prosternum nearly as in older stages. Coxal pores very small, 2, 1, 1, 1. . 0, 0, 0, 1, 1 0, 0, 1 Spines of first legs, f-p-{4444 or (att 1 the ventral spine of third joint being minute; of the second, {7 0 74 or gga; of the fourth, Sore; of the fifth to ninth, Cored; of the tenth, eos to Coi33; of the eleventh, Coase OF oeess; of the twelfth, 1 oe or goq33; of the thirteenth and_penult, Soria; of the anal, 0, 0, 0, 0, 0 0-0-11,0: Posterior coxae with neither bristles nor spines. Female gonopods elongate; conical; biarticulate; with no trace of claw or spines. Anal glands still present in fully developed form. Length cir. 6.4 mm. Pullus IV (Larva quarta).— Very pale, dilute yellowish. Head more pigmented, somewhat orange. Antennae composed of seventeen articles. Ocelli 1+ 2, 1. Caudal ocellus of upper series and that of the lower larger than the single ocellus; anterior one small. All distinct and well separated. Prosternum with line of apices of teeth recurved. Sinus propor- tionately somewhat smaller than in the adult. Each twelfth coxa with a single pore. Twelve developed pairs of legs present, the other three pairs ap- pearing as slender pressed buds. Spines of first legs, [0-34 or Cora; of the second, p-e114; of the third, (:¢:¢74; of the fourth to eighth, cera; of the ninth, teti41; of the tenth to twelfth, niet 194 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Anal glands well developed. Length circa 6 mm. Type.— M. C. Z., No. 429, California: Santa Cruz Island, La Playa Canyon. R. V. Chamberlin. Paratypes.— M. C. Z., No. 359-364, 427, 428, California: Santa Cruz Island, La Playa Canyon. R. V. Chamberlin. This species is very close to NV. nampus but the differences appear clearly and definitely upon close study. It is a more slender species with a proportionately smaller head, being 8.5 or more times longer than width of the tenth plate whereas the ratio in NV. nampus is usually between seven and eight or but slightly more, to one. The pro- sternum is proportionately narrower. It is a distinctly more reddish species, being normally chestnut in the adult. The spines of the female gonopods are somewhat more slender and are more uniformly and acutely acuminate. Finally, the first four or five pairs of legs have the third joint with but two dorsal spines as against three in NV. nampus on all or on all but the first pair. There appears to be a difference in habitat, N. znsulae having been found only in the cooler, moister portions of La Playa Canyon on Santa Cruz Island, where N. nampus occurs on the hills and in the more open places as it does on the mainland. NoOTHEMBIUS NAMPUS, sp. nov. Plate 8, fig. 4; Plate 9, fig: 6, -7. Draanosts.— Less slender; first dorsal plate near 1.64 times wider than long; head wider than long. Testaceous to brown, head and first dorsal plate light chestnut. Ocelli seven to nine in two or three series. Prosternum 1.66 to 1.68 times wider than long; distance between chitinous spots 3.75 times dental line; sinus shallow and wide; teeth 2+ 2. Basal spines of gonopods less slender (Plate 8, fig. 4). Coxal pores 3, 4, 4, 3. Third joint of all legs, or of all but first pair, with three dorsal spines. Last one or two pairs of coxae laterally armed. Spines of anal legs, ventral, in female 9, 1, 3, 2, 1, in male 0, 1, 3, 2, 0; of thirteenth, dorsal, 1, 0, 3, 1, 1; of twelfth, dorsal, 0, 0, 3, 1, 2. Lobe of anal leg of male as in N. insulae. Length 7 to 12 mm. DescrIPpTION.— Dorsum testaceous to brown, the plates caudad of the middle in most showing a distinct median longitudinal dark stripe which may also sometimes be present on the anterior ones. CHAMBERLIN: LITHOBIID GENERA. 195 Head and usually the first dorsal plate pale chestnut, frontal region paler and on caudal portion a darker median longitudinal stripe. Prosternum mostly light brown, the prehensors paler. Venter yellow to testaceous, with the caudal plates darker, burnt brown. Legs yellowish or testaceous, the posterior pairs more densely pigmented. Body from 7 to 83 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other as 29 : 28 : 29 : 31 : 32 : 30. Head wider than long. Caudal margin mesally a little incurved. Sides between eyes and lateral breaks characteristically nearly straight and subparallel. Antennae very short, reaching or not fully attaining the fifth body segment. Articles normally twenty-two, sometimes only twenty-one; short and mostly rather slender; distally shorter ones may occur at intervals between longer ones. Ocelli seven to nine in two or three series: e. g., 1 + 4, 3, a very common arrangement; 1+ 3, 3; 1+ 4, 2,1; 1+ 4, 3, 1. Single ocellus largest, the others, excepting most caudal, of small or very small size. Prosternal teeth 2 + 2 2; inner one of each pair larger than the outer so that line of apices is commonly almost straight. Sinus shallow and wide, with sides concave. Spine slender and straight. 1.66 to 1.68 times wider than long. Distance between chitinous spots 2.64 times width at level of bottom of sinus; 3.75 times the dental line. Distance between apices of mesal teeth equal to that between apices of teeth of each pair (Plate 9, fig. 6). First dorsal plate about 1.64 times wider than long; sides back of about anterior third of length considerably converging and but little convex; caudal corners well rounded. Caudal margin of ninth plate straight; the eleventh and especially the thirteenth plate with the caudal corners obliquely excised, the line of excision straight and making but a slight angle with the median portion of margin. Coxal pores small, decreasing in size proximad on each coxa in the usual way: 2, 3, 3, 2; 3, 4, 4, 3, the most usual number and arrange- ment. i: of the second to eleventh, . 9 Spines of first legs, (3333 or 0, 0, 3, cease; of the twelfth, eT ce ae thirteenth and penult, Gress; of the anal, p¢esy (9) or E330 (o7). Last two pairs of coxae, or rarely but the last pair, ae ee Anal legs in male short, of uniform thickness. The tibia at its 196 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. distal end with mesoventral side obliquely excised and bearing at this place a characteristic oblong lobe set somewhat obliquely to the longitudinal axis of the jomt, somewhat expanded above base and distally flattened as in NV. insulae; bristles of lobe rather few, straight. Gonopods of male pale; small, rounded and wart-like; bearing one or two bristles. Claw of female gonopods short and unusually thick and stout; rather strongly curved; strictly entire; distally narrowly acutely rounded. Spines 2 + 2 or 2 + 3, the outer considerably larger than the inner; in ventral view the outer is attenuated from near base, the inner being proportionately broader with the sides parallel from base to an acuminate apical portion (Plate 8, fig. 4; Plate 9, fig. 7). Inner side of first article considerably diverging proximad from that of the opposite one, proximally concave. Length 7 to 12 mm., the female types varying from 9 to 12 mm. and the male types from 7 to 10 mm. in length. A male 9.75 mm. long has antennae 3.5 mm. and anal legs 3.4 mm. long and the tenth plate 1.17 mm. wide. Typre.— M.C. Z., No. 426, California: Claremont. R.V.Chamber- lin. ParatyPes.— M. C. Z., No. 420, 424, California: Claremont; M. C. Z., No. 358, 425, Eaton’s Canyon; M. C. Z., No. 421-428, Santa Cruz Island. R. V. Chamberlin. NoTHEMBIUS ABERRANS, sp. Nov. Plate 8, fig. 5; Plate 9, fig. 4, 5. Dracnosts.— First dorsal plate near 1.86 times wider than long; head wider than long. Brown, head a little darker and of chestnut cast. Ocelli nine to eleven in three series. Prosternum 1.64 times wider than long; distance between chitinous spots 4 times dental line; teeth 3 + 3. Outer spine longer and stouter than inner. Coxal pores 3, 3, 3, 3 to 4, 4, 4, 3. Third joint of first four or five pairs of legs bearing only two dorsal spines and of first nine only two ventral ones. Last two pairs of coxae laterally armed. Spines of anal legs, ventral, in both sexes 0, 1, 3, 2, 1; of thirteenth legs, dorsal, 1, 0, 3, 1, 2 or, rarely, 1, 0, 3, 1, 1; of twelfth, dorsal, 0, 0, 3, 2, 2 or 0, 0, 3, 1, 2. Lobe of anal leg in male as in Plate 9, fig. 5. Length 8. 5 to 10 mm. Description.— Dorsum typically brown. Head a little darker CHAMBERLIN: LITHOBIID GENERA. 197 and of chestnut cast. Antennae dark brown, light at tips. Venter brown. Prosternum darker than venter, of chestnut tinge. Legs pale brown, posterior pairs darker, all legs lighter distally. In some specimens the body is paler throughout and of a greenish tinge, as is often the case in the related species. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 58 : 56:60:71 : 75 : 73. Head wider than long in about ratio 58:53; widest at level of breaks, widely rounded and but moderately narrowed anteriorly. Antennae short and well attenuated. Articles twenty-two, short and very short, decreasing distad in the usual way. Ocelli nine to eleven in three series: e. g., 1 + 4, 3, 2, the commonest arrangement, 1 + 4, 3, 1; 1+ 4, 3, 3. Single ocellus largest; the most caudal one of first and of second series also large, the others decreasing anteriorly. Prosternum in type 1.64 times wider than long. Distance between chitinous spots 2.35 times width at level of bottom of sinus; four times the dental line. Teeth 3 + 3, all moderately acute and with sides straight, the most mesal on each side smaller than the other two which are subequal, and the interval separating it from the median one of the three shallower and much narrower than that between the outer two. Line of apices recurved. Sinus narrow, narrowly rounded at bottom. Spine as usual (Plate 9, fig. 4). First dorsal plate in type about 1.86 times wider than long; widen- ing moderately cephalad. Posterior angles of ninth plate rounded, of the eleventh straight, and of thirteenth distinctly, although but moderately, produced. (Gocal-porés 3. on On oe Arc on ae Aa 44, 4. 1 0, 2,21 1 0,2,2,1 ’ 2, 2, 0,052, 2a Spines of first legs, or goss (on one side) to g--3333 0, 0, 2, 2, 2 ‘0, 0, 2,22 ee of the secant 0023-1 OF ooa32; Of the third and fourth, 9-9-33°3; 0, .0;.2, 2, 2 , 0, 3, 2, 2 1 By ¢ 2,2 of the fifth, (¢335 Or gases; Of the sixth to ninth, ¢p335; of 3, 2, 2 Ay? Os dba the tenth and ents fates 5 of the twelfth, (ess OF oro ss3 f h } I 0,0, 3,1,2 0,0, 3,1, 2 id l LESS Hale of the thirteenth, (3°33, 003-32 (on one side), or rarely Salen uae of the pam ress or occasionally, on one side, $1337; of the anal, (73:21, alike in both sexes; claw one. Last two pairs of coxae laterally armed. Anal and penult legs considerably thickened in both sexes, especially the third, fourth, and fifth joints, the tibiae and tarsi remaining in comparison abruptly more slender. In the male the fifth joint of the 198 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. anal leg is less markedly excised at distal end than in the other two species. The lobe borne is comparatively thin, is higher caudally than anteriorly and has the distal or free surface convex; it bears numerous long, distally curved, hairs; the lobe is set somewhat obliquely to the long axis of the joint and is proportionately lower and smaller than in the other species (Plate 9, fig. 5). The claw of the female gonopods is long and rather slender, acutely pointed, strongly curved and moderately excavated. The outer spine is distinctly and considerably stouter and longer than the inner; ‘spines narrowed from base to apical portion which in outer spine is acute and in inner is almost obtuse (Plate 8, fig. 5). Inner edge of first article straight but distinctly diverging from the median line cephalad. Length 8.5 to 10 mm. Type.— M. C. Z., No. 357, California: Eaton’s Canyon, near Altadena. R. V. Chamberlin. ParatyPEs.— M. C. Z., No. 365, 480, California: Eaton’s Canyon, near Altadena. R. V. Chamberlin. This species is readily separated from the two other species in having the prosternal teeth 3 -+ 3 instead of 2+ 2. The male differs from those species not only distinctly in the character of the lobe borne on the anal leg but also in having the ventral spines of that leg 0, 1, 3, 2, 1 as in the female. In the spining of the legs the species is nearer to N. insulae than to N. nampus. ‘Typically it differs from both species in having the dorsal spines of the thirteenth legs 1, 0, 3, 1, 2 instead of 1, 0, 3, 1, 1. The posterior legs are proportionately stouter. The single ocellus is less enlarged. TIGOBIUS, gen. nov. Head with lateral marginal breaks. Antennae very short; articles twenty. Eyes composed of seriate ocelli; single ocellus not enlarged, smaller than one or more of the other ocelli. Prosternal teeth 2 + 2, the line of apices recurved. Spine slender. Incision rather large, v-shaped. Posterior angles of none of the dorsal plates produced. Coxal pores small and circular, uniseriate. Claw of female gonopods short, tripartite, with median tooth larg- est. Basal spines 2 + 2 acuminate from base. 199 CHAMBERLIN: LITHOBIID GENERA. Fg a i ies © t—t-- i ave : t 38 Fic. 6.— Distribution of Tigobius. 200 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Tarsi of anterior legs strictly entire, with no trace of division. Anal and penult legs in male moderately and uniformly crassate. Anal leg with sixth joint bearing at distal end on dorsal side a small but conspicuous process or lobe. Anal coxae laterally armed. Third joint of none of the legs cephalad of the ninth or tenth pair dorsally armed. Fifth jomt of legs from second to near eleventh with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1,0; ventral, 0, 1, 3, 2, 0, the claw single. Dorsal spines of penult legs 1, 0, 3, 1, 1; ven- tral, 0, 1,3, 3,1; clawstwo. Dorsal spines of thirteenth legs 0, 0, 2, 1, 1; ventral, 0, 0, 2, 3, 2 or 0, 1, 2, 3,2. Dorsal spines of twelfth legs 0, 0, 2, 1, 1; ventral, 0, 0, 2, 3, 2. Dorsal spines of eleventh legs 0, 0, 0, 2, 2 or 0, 0, 1, 2, 2; ventral, 0, 0, 1, 3, 2. Dorsal spines of first legs 0; 0, O;.1;4sor 0; 00s 10"t- ventral, O70) 1290. Length 7 mm. and under. Typr.— T. paralus, sp. nov. The characteristic lobe on the sixth joint of the anal legs much resembles that found on the fifth joint of the penult legs in the males of Nampabius. The genus is at present known from a single species. TIGOBIUS PARALUS, sp. nov. Plate 10, fig. 1-4. 4 Dorsum brown, lightest at middle. Caudal portion of head simi- lar to dorsum; but anteriorly it is lighter, yellowish or subferruginous. Antennae light yellowish brown. Legs dilute brown, posterior pairs lighter, bright yellow. Venter brown, with caudal plates darker, and anterior ones suffused with violaceous. Prosternum and prehen- sors somewhat testaceous. é Body attenuated considerably cephalad from tenth plate, the first plate being much narrower than the head and than the third plate. Width of head and of first, third, eighth, tenth and twelfth plates to each other as 37 : 34.5 : 37 :42 :43 : 41. Head varying in proportions from nearly equal in length and breadth (2 paratype) to considerably longer than wide (41:37 in o& paratype). Sides substraight and but slightly converging caudad between eyes and caudal corners. Lateral marginal breaks slight. Antennae very short. Articles between third and ultimate very short. Ultimate article longer than the two preceding taken together. Ocelli 1 + 4 and 1+ 1, 3. Single ocellus somewhat smaller than CHAMBERLIN: LITHOBIID GENERA. 201 the first of series (Plate 10, fig. 2). Ocelli pale. Organ of Témésvary circular in outline, larger than the adjacent ocelli. Prosternum (Plate 10, fig. 3) strongly narrowed anteriorly, with sides of anterior portion incurving. Teeth low, angularly pointed, with line of apices distinctly recurved. Incision proportionately rather large, v-shaped, with sides a little concave. Spine slenderly acuminate, bending a little mesad. In type 1.42-+ times wider than long. Distance between chitinous spots 2.23 times width at level of bottom of incision; 3.625 times as great as length of dental line. First dorsal plate widest at or a little in front of the middle, widely rounded caudad from a little back of middle to and about the caudal _ corners; 1.7 times wider than long. Sixth, seventh, and ninth plates with the caudal corners excised, the line of excision or trunca- tion being long; eleventh plate with caudal corners only shortly excised; fourth and thirteenth with caudal margins straight. Coxal pores‘ very small: 2, 2, 2, 2. ws a ee OnOr I hae Spines of first legs, ¢o-¢34 OF Gudea; of the second, COST OF orei; of the third, 3 ys 0,1, 0,1 of the fifth to eighth, v1 0, 2 0, , 0, 0, 0, 0, 2 0, 0, 2 ¢ =i; of «the ee Oe «OF ob 013 0, 0, 2, “i; of the ninth, j734; of the tenth, "0,0,0,2,2 0,0,1,: ;) Of ‘the eleventh, ggaa-5) dot , 2, 2, 2 0, 0, 2, 1,1 0,.0, 2, 1, co0ra2; Of the twelfth, gos35; of the thirteenth, 99-335 or . 0,0,2,1,1 1,.0,3, (on one ade), Toes2; of the penult, 77334, claws two; of the anal, (-7'3'2'» Claw single. Anal coxae alone laterally armed. Anal and penult legs of male moderately and uniformly crassate. Penult not otherwise modified but sixth joint of anal leg bearing on dorsal surface at distal end a small but conspicuous subcylindric process very similar to that occurring on tibia of penult legs in species of Nampabius (Plate 10, fig. 1). Gonopods of male well exposed, sides subparallel and distal edge substraight; each bearing a single bristle. Claw of female gonopods moderately short, but little curved, tripartite, with median tooth considerably largest and the lateral ones subequal. Spines acutely acuminate from base to apex; the outer longer than the inner (Plate 10, fig. 4). Length from 5 to 7 mm. TypE.— M. C. Z., No. 366, California: Pacific Grove. R. V. Chamberlin. Paratypes.— M. C. Z., No. 367, California: Pacific Grove; M. C. Z., No. 368, 452, Santa Barbara; M.C. Z., No. 451, 453, Santa Cruz Island. R. V. Chamberlin. 0, if 0, 0 0, 0, 0,22 0,0,1,2,2 0, 0, 0, 2, 2 See Wdiben WE iiaeehal 2 | pone ta ea pare nnn} ab alt Pee fie : Rubee ahha die Ha. G1, thoi seks cae i A | Ney ius rate yi i ha ; Bs a he ; iu | ut | 7G ¥ ih mn a i | he ak ra Wy Padi hy | Ve nm vue ae) aes a ety a el sit ae Me AL o y hy , i vate ar ‘ i , Cint n he Va se dt rae e ik aoe, as ae as Phot a Phy X Pias ak Hy ball ae ' teat Mad ma aed Mid ee Tei ahs oan : he CNA WEN ie thas ntinte’ ee bp) ie UE ah re BS Aah * OK th dy. ‘a weed, Mi wisks oa abi) if ve. rit at in 0 [ey ri oy im oe : jal Aen WO) ed | » TA 2 ‘ 7 i aaa TNR) rales th Ti se i) ‘bbe. foriiat, meray in) sl), Bau mit pea cmatladuedt awk agsrheg Fn face Suet " its ea! | Oi Lain pean’, My Ny vate | Coe, ui ae hia Bt noe red) ea ead i ‘ ie aan? ip shige Mgae ’ oped he TAlty AS eet # ’ ebmneteadoyncl eBhiatseas Ate shbadh Res wee i Koad Allaah tye erro Bart RKO aie oy ae gi Ceelahaeery it " be aay) saredlngi oe if w poet vy s Ae ie ony eapwal he Layap tolf oy aq, * panty Livery Ve : i iv aly sh Pate 4 apy: seas ae f i : Youd o . hier Peg ; Pe ee hy 6 ty Pious ies eh rath fos semiesh in? i i 4 Oa Myint? a) Wa pO z a ah mr Prat Reh vag Rent Kahiin hi fants: TES | ini bout fi] ne ite rer ri eva BL ff a a ent Ne we te a Rit " 7 i hei if neu, ry Rear nieh. ant het} ayitli. lng Ki at ae : uf 4 Gh i Keay ne We a pays 7 Nin Hick WS if REALE i ha vas TAP Br \ ua i Sema Leahy ta stdin) dass t ow ee) Tay ‘at pI ) Mei aye: p TAD | iy sy “git Sees en Hate as ; ai wey ; et chad %.) ida Ball inertia ; at Ria a na ca \ ’ ii f : PEATE I, CHAMBERLIN.— Lithobiid Genera. PLATE 1. Oabius patonius (Chamberlin). (Mill Valley). Fig. 1. Last three legs of right side, dorsal view. Fig. 2. Anterior view of distal portion of the seventh left leg, showing the undivided tarsus. Fig. 3. Anterior portion of prosternum showing teeth, spines, sinus, etc. Fig. 4. Right eye, showing arrangement and form of ocelli. Organ of Témésvary appears in outline at the right and below. Oabius patonius flavus Chamberlin. (Santa Barbara). Fig. 5. Right eye and organ of Témésvary (same scale as for fig. 4). Oabius pylorus Chamberlin. (Capitola). Fig. 6. Right eye and organ of Témésvary (same scale as for fig. 4). Oabius tiganus (Chamberlin). (Pacific Grove). Fig. 7. Right eye and organ of Témésvary (same scale as for fig. 4). Oabius fratris Chamberlin. (Friant). Fig. 8. Right eye and organ of Témésvary (same scale as for fig. 4). Oabius tabiphilus Chamberlin. (Santa Barbara). Fig. 9. Right eye and organ of Témésvary (same scale as for fig. 4), Oabius patonius micrus Chamberlin. (Pacific Grove). Fig. 10. Anterior portion of prosternum. Bull. Mus. Comp. Zodl. Oabius Plate 1. PLATE 2. CHAMBERLIN.— Lithobiid Genera. PLATE 2. Oabius pylorus Chamberlin. (Capitola). Fig. 1. Claw of right gonopod of female, ventral view. Fig. 2. Basal spines of right gonopod of female, more highly magnified. Oabius (Nyctobius) ineptus Chamberlin. (Portland). Fig. 3. Claw of left gonopod of female (same magnification as for fig. 1). Fig. 4. Basal spines of left gonopod of female (same scale as for fig, 2). Oabius tiganus (Chamberlin). (Pacific Grove). Fig. 5. Basal spines of left gonopod of female, showing also straight mesal edge of proximal article (same scale as for fig. 2). Oabius patonius micrus Chamberlin. (Pacific Grove). Fig. 6. Basal spines of left gonopod of female (same scale as for fig. 2). Oabius patonius flavus Chamberlin. (Santa Barbara). Fig. 7. Basal spines of left gonopod of female (same scale as for fig. 2). Oabius tabiphilus Chamberlin. (Santa Barbara). Fig. 8. Basal spines of left gonopod of female (same scale as for fig. 2). Oabius uleorus Chamberlin. (Forrester Island). Fig. 9. Basal spines of left gonopod of female (same scale as for fig. 2). Oabius fratris Chamberlin. (Friant). Fig. 10. Basal spines of left gonopod of female (same scale as for fig. 2). Oabius Plate 2. Bull. Mus. Comp. Zo6l. piste ea rear BANS iad - as att) i Ls : : oan le | i a nu i " ‘ AT : 7 i et | fy sa aR i jan te A heat a i a reer, wou u ‘ - ; ie ye oe oa oA i a ny) 71% é fi i ur if x a 0 cr | PLATE 3. ‘CHAMBERLIN.— Lithobiid Genera PLATE 3. Oabius dissimulans Chamberlin. Fig. 1. Basal spines of left gonopod of female from Brookdale (same scale as for Plate 2, fig. 2). Fig. 2. Basal spines of left gonopod of female from Santa Barbara. Fig. 8. Basal spines of left gonopod of female from Sausalito. Oabius (Nyctobius) ineptus Chamberlin. (Portland). Fig. 4. Basal spines of left gonopod of female. Oabius (Nyctobius) decipiens Chamberlin. (Pacific Grove). Fig. 5. Basal spines of left gonopod of female. Paobius boreus Chamberlin. (Forrester Island). Fig. 7. Ocelli of left eye. Organ of Témésvary not represented (same scale as for Plate 1, fig. 4). Fig. 8. Anterior portion of prosternum. Fig. 9. Second leg of right side, anterior view, showing line separating the two parts of tarsus. Bull. Mus. Comp. Zodl. Oabius Plate 3. Bl ts An. Wie ne sy a ell iy on % " ij “hs : \ 1 LY na) : y ity # ; i A i ie 3 e( rn _ o } a J) - - : ' _ ' 7 é Gr i i | : La) ' ‘ { \ Pn uceeringt a Neen ike yi ui rh yin e - coin “4 a] yi 3 ¥y , oo if Hiss eae OG aw - van Ne a _- 5" < ' F 7 t i iy PLATE 4. (CHAMBERLIN.— Lithobiid Genera. PLATE 4. Paobius boreus Chamberlin. (Forrester Island). Fig. 1. Claw of left gonopod of female, ventral view. Fig.2. Basal spines of right gonopod of female more highly magnified (same scale as for Plate 3, fig. 1). Fig. 3. Head in outline, showing character of margin. Paobius columbiensis Chamberlin. (Kaslo). Fig. 4. Basal spines of right gonopod of female (same scale as for fig. 2). Kiberbius ogmopus Chamberlin. (Eaton’s Canyon). Fig. 5. Head in outline. Fig. 6. Anterior portion of prosternum. Kiberbius nannus Chamberlin. (Eaton’s Canyon). Fig. 7. Head in outline (same scale as for fig. 5). Bull. Mus. Comp. Zodl. Oabius Plate 4. CHAMBERLIN.— Lithobiid Genera. PLATE 65. Kiberbius ogmopus Chamberlin. (Eaton’s Canyon). Fig. 1. Right anal leg of female, ventral view, showing the characteristic furrowing. Fig. 2. Seventh left leg, anterior view, showing demarcation of two parts of tarsus. Fig. 3. Basal spines of left gonopod of female (same scale as for Plate 3, fig. 1). Kiberbius nannus Chamberlin. (Eaton’s Canyon). Fig. 4. Basal spines of left gonopod of female (same scale as for fig. 3). Arebius (Pagobius) diplonyx Chamberlin. (Santa Barbara). Fig. 5. Distal portion of fourth leg on left side, showing structure of tarsus. Arebius (Pagobius) kochii (Stuxberg). (Sausalito). Fig. 6. Anterior portion of prosternum. Oabius Plate 5. Bull. Mus. Comp. Zool. oT ae ihn CHAMBERLIN.— Lithobiid Genera. PLATE 6. Arebius (Pagobius) kochii (Stuxberg). (Sausalito). Fig. 1. Ocelli of left eye. Organ of Témésvary not shown (same scale as for Plate 1, fig. 4). Fig. 2. Basal spines of gonopod of female (same scale as for Plate 2, fig. 2). Fig. 3. Claw of left gonopod of female, ventral view (same scale as for Plate 2, fig. 1). Arebius (Pagobius) dolius Chamberlin. (Friant). Fig. 4. Basal spines of left gonopod of female. Fig. 5. Ocelli of right eye, with organ of Témésvary in outline. Fig. 6. Claw of right gonopod of female, ventral view (same scale as for fig. 3). Arebius (Pagobius) diplonyz Chamberlin. (Santa Barbara). Fig. 7. Basal spines of left gonopod of female (same scale as for fig. 2). Fig. 8. Claw of right gonopod of female (same scale as for fig. 3). Oabius Plate 6. i: Zoo Bull. Mus. Comp PLATE 7. CHAMBERLIN.— Lithobiid Genera. PLATE 7. Arebius medius Chamberlin. (Brookdale). Fig. 1. Basal spines of right gonopod of female, ventral view (same seale as for Plate 2, fig. 2). Arebius elysianus Chamberlin. (Pasadena). Fig. 2. Basal spines of left gonopod of female. Arebius obesus (Stuxberg). (Stanford). Fig. 3. Basal spines of left gonopod of female. Fig. 4. Claw of right gonopod of female. Arebius oregonensis Chamberlin. (Portland). Fig. 6. Anterior portion of prosternum (bristles not represented). Bull. Mus. Comp. Zo0l. Oabius Plate 7. PLATE 8. CHAMBERLIN.— Lithobiid Genera. PLATE 8. Arebius obesus (Stuxberg). (Stanford). Fig. 1. Portion of right gonopod of female in outline, ventral view, showing form of the first article. Nothembius nampus Chamberlin. Fig. 4. Basal spines of left gonopod of female. Nothembius aberrans Chamberlin. (Eaton’s Canyon). Fig. 5. Basal spines of left gonopod of female. Nothembius insulae Chamberlin. (Santa Cruz Island). Fig.6. Basal spines of left gonopod of female. Bull. Mus. Comp. Zool. Oabius Plate PLATE 9. CHAMBERLIN.— Lithobiid Genera. PLATE 9. Nothembius insulae Chamberlin. (Santa Cruz Island). Fig. 1. Portion of right anal leg of male, ectoventral view, showing form of lobe at distal end of tibia. Fig. 2. Outline of part of right gonopod of female, ventral view, showing form of first article. Fig. 3. Anterior portion of prosternum. Nothembius aberrans Chamberlin. (Eaton’s Canyon). Fig. 4. Anterior portion of prosternum. Fig. 5. Part of left anal leg of male, mesal view, showing form of lobe at distal end of tibia. Nothembius nampus Chamberlin. (Eaton’s Canyon). Fig. 6. Anterior portion of prosternum. Fig. 7. Outline of portion of right gonopod of female, ventral view, showing form of first article. Bull. Mus. Comp. Zodl. Oabius Plate 9. ys tite wad iit CHAMBERLIN.— Lithobiid Genera. PLATE 10. Tigobius paralus Chamberlin. (Pacific Grove, Cal.). Fig. 1. Distal portion of right anal leg in mesal aspect showing the char- acteristic lobe on the first tarsal joint. Fig. 2. Ocelli of left eye. Fig. 3. Anterior portion of prosternum (the hairs not represented). Fig. 4. Basal spines of left gonopod of 9 in ventral view. Pokabius species. (British Columbia). Fig. 5. Ocelli of right eye. Fig. 6. Basal spines of left gonopod of 9, ventral view. Bull. Mus. Comp. Zodl. Oabius Plate to. Bulletin of the Museum of Comparative Zodlogy AT HARVARD COLLEGE. Vor. -bVEL No.5. THE GOSIBIIDAE OF AMERICA NORTH OF MEXICO. By Raupx V. CHAMBERLIN. With Six PLartes. CAMBRIDGE, MASS., U.S. A. PRINTED FOR THE MUSEUM. DECEMBER, 1917. aeons Janet i saan aN No. 5.— The Gosibiidae of America North of Mexico. By Rapa V. CHAMBERLIN. THE genera included in the family Gosibiidae occur in Mexico and across the southern portion of the United States. Nearly all of the lithobioid chilopods thus far reported from Mexico, in fact, appear to belong here. When Mexico shall have been well explored, there is little doubt that the family will be found to be a large one. From the southwest it has spread in the United States northward to north central California and to Colorado and northeastward to Tennessee: and North Carolina. LirHosiomiwEA Chamberlin. Bull. M. C. Z., 1915, 59, p. 531. Erected in contrast with the new superfamily Henicopoidea to include the Lithobiidae proper, Watobiidae, Ethopolidae (Ethopolys, Bothropolys and allies), and Gosibiidae. GOSIBIIDAE Chamberlin. Bull. M. C. Z., 1915, 59, p. 531. The members of this family have the claw of the female gonopods always large and strictly entire, with never any trace of teeth or partite condition, any element giving such trend or expression being apparently wholly absent. First article of gonopods broad, pro- truding well mesad, with mesal edge usually strongly chitinized and either conspicuously and characteristically excavated at base or there curving more evenly but distinctly ectad; the excavation commonly the deeper continuation of a transverse furrow or constriction extend- ing across the proximal end of the article. Basal spines large and stout, the article beneath them compressed more or less to a thin 206 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. edge which is typically well chitinized. Spines of each gonopod nearly always in a plane, and subparallel with or making but a small angle with the principal plane of the gonopod (7. e. the one including the axis and the greatest diameter of the first article), and thus making a distinct angle with that of the spines of the other gonopod (7. e., when seen in caudal view the two rows of spines usually meet at a distintt angle instead of lying in the same horizontal plane as in the Lithobiidae). In the males either the penult or the anal or, much more commonly in ours, both the penult and anal legs showing conspicuous special lobes or modifications, the penult being more constantly and usually more conspicuously modified than the anal. Excepting in the subgenus Guambius, the dorsal spines of the penult legsvare Uy 051332, .2)0r 1,/0;7.3): 25-15 those of the anal legs also, with rare exceptions, being the same. In mouthparts, eyes, general structure of prehensors, and _ pre- hensorial segment, and in most other general structural features, agreeing with the Lithobiidae proper. Key to Genera. a. Coxae of last five pairs of legs bearing pores. Pseudolithobius Stuxberg. aa. Coxae of only the last four pairs of legs bearing pores. b. Prosternal spines very stout, exceeding the teeth in size; articles of antennae normally twenty. " Arenobius Chamberlin. ® bb. Prosternal spines small and mostly bristle-like, at least distally, always more slender than the teeth; articles of antennae twenty-one to thirty-five, mostly above twenty-four. c. Penult legs of @ having fifth article more or less obliquely depressed or excavated above at distal end and there bear- ing at very end a conspicuous lobe or crest. (Claws of anal legs two or three; species 10-15.5 mm. long). Guambius Chamberlin. ce. Penult legs of &@ with fifth article thickening distad, but never excavated above nor bearing at end any such crest. (Claws of anal leg one or two; species 15-30 mm. long). Gosibius Chamberlin. CHAMBERLIN: THE GOSIBIIDAE. 207 Gosrpius Chamberlin. Can. ent., 1912, 43, p. 204; Ann. Ent. soc. Amer., 1912, 5, p. 146. Lateral marginal breaks of head distinct. Antennae from short to long, mostly moderately long; articles twenty-one to thirty-five, nearly always above twenty-four. Eyes composed of ocelli in several series. Single ocellus distinctly largest. Prosternal teeth 2 + 2 or, less commonly, 3+ 3. Line of apices recurved. Sinus large; between V- and U-shaped. Ectal spines much more slender than the teeth, usually, but not always, distally bristle-like. Body moderately narrowed cephalad, distinctly less so than, e. g., in Arenobius. First dorsal plate nearly always as wide as or wider than the third. Posterior angles of none, of ninth, eleventh, and thirteenth, or of the sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced. Coxal pores circular, uniseriate; present on last four pairs of coxae. In the male the penult legs have the fifth joint distinctly elevated distally and thus there manifestly thicker than proximally, with or without a nodular protuberance on the elevated portion but this never at the very distal end and never cristate. While rarely essen- tially unmodified, the anal legs usually have the fourth joint distinctly longitudinally furrowed above and widened, and in some the fifth also more or less widened and bowed ventrad, the joint along side of furrows more densely pilose. Dorsal spines of anal legs mostly 1,0, 3,2; 1 or 1, 0).3, 2, 2; more rarely 1, 0;3, 2;,0 or even 1, 0, 3, 1, 0; ventral spines 0, 1, 3, 3 (2), 1 or 0, 1,3, 3, 2; claws one or two. Dorsal spines of penult legs 1, 0, 3, 2, 2, rarely 0, 0, 3, 2, 2; ventral 0, 1, 3, 3, 2 or 0, 1,3,3,3 or very rarely only 0,1,3,3,1. Dorsal spines of fourth to thirteenth legs, and usually of second and third as well, 0, 0, 3, 2,2; of first legs either likewise 0, 0, 3, 2, 2 or 0, 0, 2, 2, 1 to 0, 0, 2, 1,1. Some of the posterior coxae dorsally armed; and either none or from last one to four pairs laterally armed. Gonopods of male uniarticulate. Claw of. female gonopods large and entire as usual. Basal spines 2+2 or 3+ 3. First article large, protruding mesad over base, with the inner side strongly chitinized and usually appearing con- spicuously excavated proximally, commonly distinctly constricted or furrowed across base. 208 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Species from 15 to 30 mm. in length. Typr.— G. paucidens (Wood). The species of this genus so far as known from our region are con- fined to the extreme southwestern United States; 7. e. to Arizona and California (Figure 1). The genus is close to Guambius; but it differs, e. g., conspicuously and constantly in the character of the modification in the penult legs, as also in most cases equally clearly in that of the anal legs of the male. Key to Subgenera and Species of Gosrbius. a. Posterior angles of none of the dorsal plates produced. Abatobius, subgen. nov. b. Claw of anal legs single; fifth joint of first thirteen pairs of legs with three ventral spines; dorsal spines of anal legs DOORS desu ho pee retrecy aNte een BN Ce G. arizonensis, Sp. nov. bb. Anal legs with two claws; fifth joint of none of the legs with three ventral spines; dorsal spines of anal legs 1, 0, 3, 2, 0. G. montereus, sp. nov. aa. Posterior angles of ninth, eleventh, and thirteen dorsal plates BEOUUCEHRY: oie ra Ree ee ae Gosibius, sens. str. b. Anal legs with two claws; basal spines of 9 gonopods 2 + 2. c. Ventral spines of anal legs 0, 1, 3, 3, 1. G. monicus Chamberlin. cc. Ventral spines of anal legs 0, 1, 3,3, 2; tibia of first and second legs with 2 dorsal spines. G. claremontus, sp. nov. bb. Anal legs with but one claw (Basal spines of Q gonopods normally 3+ 3, rarely 2 + 2). c. Ventral spines of anal legs 0, 1, 3, 3, 2; articles of antennae twenty-eight to thirty-five........ G. paucidens (Wood). ce. Ventral spines of anal legs 0,1, 3,2,1 or 0,1, 3,3, 1; articles of antennae twenty-one to twenty-three. G. brevicornis, sp. nov. aaa. Posterior angles of sixth, seventh, ninth, eleventh, and thir- teenth dorsal plates produced...... Timiobius, subgen. nov. Anal legs with two claws.............. G. intermedius, sp. nov. 209 THE GOSIBIIDAE. CHAMBERLIN: Distribution of Gosibius. ie Fig. 210 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Gosipius (GOSIBIUS) PAUCIDENS (Wood). Plate 1, fig. 1-4. Lithobius paucidens Wood, Journ. Acad. nat. sci. Phil., n. s., 1862, 5, p. 14; Trans. Amer. phil. soc., n. s., 1865, 13, p. 151. Lithobius paucidens Stuxberg, Ofver. K. vet.-akad. Férh., 1875, p. 12; Proce. Calacad. sc, US, . po lav. Lithobius paucidens Chamberlin, Ann. Ent. soc. Amer., 1909, 2, p. 191; Po- mona college -journ. ent., 1910, 2, p. 372. Gosibius paucidens Chamberlin, Can. ent., 1912, 43, p. 204. Draenosis.— Dark brown with a darker median dorsal stripe; head and first and last tergites orange or ferruginous. Antennae long; articles twenty-eight to thirty-five. Ocelli fourteen to twenty in’ four or five series. Prosternal teeth 2 + 2. First dorsal plate wider than third. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. Basal spines of female gonopods 3 + 3, or rarely 2-+ 2. Dorsal spines of from last seven to last ten pairs of legs 1,0, 3, 2,2; of the others, 0,0, 3,2,2. Anal legs with claw single; ventral spines 0, 1,3,3,2; dorsal 1,0,3,2,2. Ventral spines of twelfth and thirteenth legs 0, 1,3,3,3; of seventh to eleventh 0,0, 3, 3,3; of others 0,0, 2,3,3. Trochanter of last four pairs of legs armed. Anal legs of male with fourth joint widened, com- planate, and longitudinally furrowed above (Plate 1, fig. 3). Penult legs of male with fifth joint elevated distad (Plate 1, fig. 4). Length 18-30 mm. DerscripTion.— Dorsum testaceous to dark brown and subferrugi- nous, usually with a dark median longitudinal stripe which is often black and rather wide, sometimes embracing most of width of dorsum, the dark color becoming more dilute ectad. Head orange or ferru- ginous, with the first and the last dorsal plate usually of similar color but this paler or more dilute, though sometimes these plates scarcely differ from the rest of the dorsum; the dark median stripe of dorsum often extending upon head. Antennae like head or nearly so. Venter yellow to testaceous and almost clear brown, the last 3 or 4 plates darker, commonly of orange or ferruginous cast. Prosternum and prehensors orange to ferruginous. Legs yellow to testaceous; the caudal pairs a little darker than others, uniform. Body mostly near 7.75 times longer than width of tenth plate. Only moderately narrowed cephalad to first plate which is wider CHAMBERLIN: THE GOSIBIIDAE. 211 than the third but much narrower than the head. Width of head and of first, third, eighth, and tenth plates to each other (in 9) as 69 : 63 : 62 : 69 :-72. Head broadly subcordate; the sides caudal of marginal breaks moderately converging, the caudal corners well rounded and the caudal margin mesally incurved. Considerably wider than long (7:6 or nearly so). Appearing smooth and shining, being only obscurely roughened and with the punctae fine. A rather deep sulcus, this usu- ally angular and often pit-like at middle; sometimes a median longi- tudinal sulcus leading forward from it. Antennae long, reaching mostly somewhere between beginning of eighth and end of ninth segment, varying considerably. Articles twenty-eight to thirty, all long and cylindric, usually more or less uniformly and gradually decreasing distad, occasionally those beyond the first 12-16 rather abruptly shorter. Hairs short. Ocelli from fourteen to twenty in number, nearly always in four series but occasionally in five. The most frequent arrangement is 1+ 4,4,4,3. Examples of other arrangements are: 1 + 4, 4, 3, 2; 1+4,4,3,3; 1+ 4,5,5,4; 1+ 4,4,4,3,1; 1+4,4,4,3,2. The single ocellus not contiguous with the others, much largest. Of the seriate ocelli those of most dorsal row are largest, often much differ- ing among themselves, those of second row but little smaller or not at all so, those of lowest row considerably smallest as usual. Prosternum 1.6 times wider than long. Anterior margin widely rounded. Teeth 2 + 2; small, well separated, but dental line as a whole relatively short; mesal tooth larger than the ectal, but its apex farther caudad and thus line of apices is recurved. Spine well ectad of outer tooth, long and slender but stouter than the bristles and also paler than the latter; springing from a rounded nodule. Distance between chitinous spots 4.66 times dental line; 1.75 times width at level of bottom of sinus. Surface of first dorsal plate nearly like that of head, appearing smooth and shining, or only obscurely uneven. Other tergites more distinctly roughened, especially the more caudal ones; these usually showing the short, submarginal, transverse sulcus on each side at about beginning of caudal third of length and a second similar, but less deeply impressed, one at middle of length; usually more or less evident also is a longitudinal furrow extending somewhat mesad of caudad from anterior margin between median line and lateral margin and bending mesad near beginning of middle third of length to join, or nearly join, the corresponding one of the other side, sometimes a branch continuing farther caudad also evident. 212 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ventral plates appearing mostly smooth and shining; with the usual three longitudinal furrows, the lateral ones, however, absent or obscure on the last one or two. Coxal pores circular or transversely a little elongate and broadly elliptic. A common number and arrangement is 5,4,4,4. The twelfth coxa has practically always one more pore than the others which have typically uniformly the same number. Other numbers ares +4, 3).3No “6/15; Oioy) ‘and 77; Gh6p6: 5 0; 0,.3,'2, 2), Q 0, 0, 3, 2, 2 Spines of first to fifth legs, oss3; of sixth, the same or, o-o-3333 2 0; 0, 3..212 POPS i252 = seventh and eighth, 9:0:3°3°3 op 003-33; Of ninth to eleventh, 1.0.3.53; of twelfth to penult, (3:33, all with 3 claws; of the anal, 1.0.8 33, claw 1. Last 7 to 9 pairs a coxae laterally armed. "Tn the female the posterior legs are short and slender; the prefemur and femur of anal legs longitudinally furrowed above, the latter the more distinctly so; the corresponding articles in penult legs more indistinctly furrowed. In the male the posterior legs are also slender; but the femur in the anal pair is typically considerably crassate and wider than the other articles, dorsally complanate and longitudinally deeply furrowed, with numerous hairs along and each side of furrow (Plate 1, fig. 3). The penult legs in the male have the prefemur and especially the femur longitudinally dorsally furrowed, while the tibia is clavately swollen, being wider and especially conspicuously thicker dorsoventrally at the distal end (Plate 1, fig. 4). There is considerable variation in the conspicuousness of these modifications, which in some specimens are weak. Gonopods of male distinctly exposed. The single article in ventral view appears truncate distally, in lateral view seem to be narrowed to an edge. Claw of female gonopods as usual; long and acute, strongly curved. Basal spines 3 + 3, rarely only 2 + 2; margin along which attached thin, strongly chitinous, blackish. lee edge strongly chitinized. First article conspicuously excavated within proximally; transversely furrowed or constricted across base! (Plate 1, fig. 2).- Length 18-30 mm. A female 20 mm. long has antennae 10 mm. long, anal leg 7.5 mm. long; and tenth dorsal plate 2.6 mm. wide. Immaturus (o).—Dorsum dilute testaceous to light brown. Head brown or paler, yellowish, the pigment denser than in dorsum of body. 1 Several females from Claremont have a stout spine at distal end of second article of gonopod, on left side in all. I have never noted myself or seen record of such occurrence of a spine on this article in any other species of the Lithobioidea. . CHAMBERLIN: THE GOSIBIIDAE. 213 Antennae brown to yellow. Prosternum and prehensors like head. Venter pale yellow to dilute brown, in most strongly suffused with green. Legs pale yellow to dilute brown, nearly uniform. Antennae varying considerably in length but mostly approaching the proportion of those of adult. Articles twenty-seven to thirty. Oceliit: 34.04, 3-6 1-4 4,.3, 2; 14-3,,4,.38, 13-144, 4,73, 1; 1+ 4, 3, 3, 2; 1+4, 4, 3, 2; proportioned nearly as in maturus. Prosternum essentially as in matwrus. Coxal pores mostly 3, 2, 2, 2. Spines of first legs, oe; of second to fifth as in adult; of 0,0, 3,22 0, 0, 3, 2,2 2 OSES ee et ias sixth and seventh, j-p-3-3-3; of eighth, o-9'3°373 OF o-0,3-3-3 Of ninth, 1, 0, 3, 2,2 1, 0, 3, 2,2 Ned 5 eoes3 OF posa3; of tenth to anal as in maturus. Last four pairs of coxae laterally armed. Anal legs in male slender, the fourth article but little enlarged, and other modifications usually more weakly indicated but their expres- sion somewhat variable. Length 14.5-17 mm. Agenitalis II. Greenish. Head dilute olivaceous with a tendency toward dull dilute orange in the frontal region; a dark spot caudad of the suture. Antennae brown, paler distad. Legs all pale, uniform. Antennae short, reaching fifth or sixth segment. Articles twenty- six to twenty-eight, short ones intermingled with longer ones. Ocelli 1 + 4, 4,3; 1+ 4,3,2; 1+ 4,4,3,1; 1+ 4,3, 2,1 Prosternal teeth 2 + 2. Sinus and spines as in adult or nearly so. Sides of anterior portion a little less strongly convexly rounded. Coxal pores 2, 1, 1, 1. Spines of first legs, (03:34; of the second, (0337; of the third and fourth, (33:3; of the fifth to math, oss; of the LeaU b] 0,0, 3.2.2 0, 0, 3, 2, 2. 322, 1,0, 3, 2 2 oss OF, sss; Of the eleventh, g-yss3; of the twelfth, jy3373; 0, 2, 2, 2 of the thirteenth and penult, °. a + 3; of the anal, coxae armed laterally. Anal glands degenerate. Length of specimens described 11.5-14.5 mm. (Santa Barbara; March, 1910). Female gonopods straight, pale. Either two or all three articles present; glabrous and with no indications of spines. Agenitalis I.— Greenish, with caudal borders of most plates more brownish. Head and antennae yellowish brown, the greenish tinge being obscure. Legs pale, uniform. Antennae wanting. 214 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli 1 + 4,3, 2. All pale, the anterior and ventral ones espe- cially so. Single ocellus largest. Prosternal teeth 2+ 2. Sinus apparently same as in adult or relatively a little narrower. Sides of anterior portion less convex and less extended laterad than in maturus. Coxal pores 2, 1, 1, 1. te Spines of first legs, (:g:i3:4; of the second, 9:33:34; of the third, oss; of the fourth, go33:3; of the fifth to eighth, j-7233; of the 0,2, 3, 0, 0, 3, 2, 2, 0,.0,.3,.2, 2), ninth, 9; O28 38 of the tenth, o:o:3:373; of the eleventh, 9:9°3:3°5 or on 1,.0,3,2,2 one side §-p-9,33; of the twelfth, §-3:3:"2; of the thirteenth and penult, 0,0, 0, 0, 0, 0, 0, 0, 0, 0 OO0111; Of the anal, o-p-7-47-9- Twelfth coxae laterally armed. Anal glands present. Length of specimen described, 9.2 mm. (Santa Barbara, March, 1910). Pullus IV (Larva quarta). A strong greenish tinge marking the other pigment, but in one specimen the more usual purplish tinge prevailed. Head lighter, in front of suture dilute dull orange, a dark spot just back of the suture. Antennae brown, lighter distad. Legs pale, uniform. Antennae already with twenty-six to twenty-eight articles; shorter and longer ones occurring variously. Length variable. Ocelli 1 + 1, 3,.3,2; 1+ 1,4,4,2; 1+4,4,2; 1+4,3,2. The single ocellus of top row in first two cases at anterior end of patch. Single ocellus largest. ' Prosternal teeth 2 + 2. Sinus V-shaped but narrowly rounded at bottom. Sides of anterior portion less convex than in adult and less extended laterad of teeth. Twelfth coxae each with single pore. Last three pairs of legs present as pale slender buds in which the division into articles is indicated in surface view by pale transverse lines. Spine: of first legs, (0-774 OT, gota; Of the second, 9:p-7°3:4; of the third, ¢-o73'3 OT, p-o-gs4; of the fury OLE of the fifth, (9:73: of the sixth the same as fifth or, (733; of the seventh, jo'7'3'5 or, 2, 2 0, 3, 2,2, eat of the eighth the same as latter or, §°o°73:3; of the ninth, 0,0, 3, 2,2 0,0,3,22. 0,0, 3, 2,2 0, 0, 3, 2, 2 0, 0, 1, 2,2 or, on 35 ot the tenth, 0; 0nds saz to 0, 0, 2, 3, 35 of the eleventh EE 0.0,0,0,0 4. 0,0,222 and twelfth, 9-¢atia to o07i°17- 0, 1, Anal aan large and conspicuous. Length 8.25-9.25 mm. (Santa Barbara, March, 1910). or CHAMBERLIN: THE GOSIBIIDAE. PAI Tyee Locauiry.— California: Fort Tejon. Also taken in California at San Bernardino, (M. C. Z., No. 557, 560); Los Angeles, (M. C. Z., No. 559, 569); Laurel Canyon, (M. C. Z., No. 567); Santa Monica, (M. C. Z., No. 573); Claremont, (M. C. Z., No. 554, 556, 558, 561, 563, 570, 576, 577); Santa Barbara, (M. C. Z., No. 562-566, 575). A very common species in southern California. Gostpius (GOSIBIUS) BREVICORNIS, sp. nov. Platev2, fig 1-4. Dracnosis.— Light to dark brown. Antennae short, with articles twenty-one to twenty-three. Ocelli nine to fourteen in three or four series. Prosternal teeth 3+ 3. First dorsal plate of same width as the third. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. Basal spines of female gonopods 3 + 3. Anal leg with claw single; dorsal spines 1, 0,3, 2,1, ventral 0, 1,3, 2,1 or 0,1,3,3,1. Dorsal spines of ninth to penult legs 1, 0, 3, 2, 2; ventral spines of penult 0, 1,3, 3,1 (c*) or 0,1,3,3,2 (2). Ventral spines of tenth to thirteenth legs 1, 0, 3, 3, 3; of second to eighth 0, 0, 3, 3, 3. Dorsal spines of third to eighth legs 0,0, 3, 2,2. None of coxae laterally armed. Last six pairs of trochanters armed. Anal legs of male with fourth and fifth articles strongly widened, the fifth bowed ventrad and both articles dorsally furrowed. (Plate 2, fig. 2). Fifth joint of penult legs of male elevated distad, bearing a rounded eminence at beginning of elevated region (Plate 2, fig. 4). Length 16 to 30 mm. DeEscrIPTION.— Dorsum light or testaceous brown to deep brown, the head either the same or, in lighter specimens with a tendency toward very dilute orange while in darkest specimens it and the first plate are darker, chestnut. Antennae like the head, 7. e., chestnut in specimens in full color either uniform or, in some, paler distad. Prosternum and prehensors like head. Venter from light yellowish brown to dark brown, the last plates darker as usual. Legs dilute yellow to pale brown, the posterior pairs in females and lighter males not darker than the others, but in darker ones of weak chestnut color. In males body nearly parallel-sided from the tenth segment cepha- lad, strongly narrowed caudad of twelfth plate, first and third tergites of same width. In females first plate is narrower than the third, and 216 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. body narrower caudad from eighth plate which is of same width as seventh. In females widths of head and of first, third, seventh, eighth, tenth, and twelfth plates to each other as 46: 45: 47: 50: 50: 47: 43. In males, omitting the seventh plate, the relations are 66: 67: 67:73: 73: 70. Body rather slender, typically 9-9.3 times longer than width of eighth plate. Head suborbicular, the caudal margin being typically convex, often considerably so, though sometimes varying to nearly straight. Margi- nal breaks weak, oblique. Head smooth excepting for several pairs of impressed dots between suture and caudal margin and two impres- sions in median line in front of suture. A little wider than long - (66:65 to 46: 45). Antennae short, reaching only to the fourth or fifth segment. Strongly narrowed distad. Articles twenty-one to twenty-three; mostly proportionately long, but the seventh and eighth, and the fourteenth and fifteenth (or sometimes thirteenth and fourteenth) articles typically abruptly much shorter than adjacent ones. Ocelli nine to fourteen in three or four series: e. g., 1 + 4, 4, 3; PAF 3,05 230) 42,55) 3; 33) e844, 425i 40463, 25 Byve-pateh very small in proportion to size of head, more toward ventral side of head than usual. Ocelli pale and inconspicuous. Single ocellus much largest, well separated. Other ocelli decreasing conspicuously cephalad and ventrad. Prosternal teeth 3 + 38; moderately large, the median on each side largest, so that if most mesal one on each side were omitted the line of apices would be procurved. Sinus between V- and U-shaped. Spines much stouter than bristles, truly spiniform, inserted close to edge of ectal tooth. Sides of anterior portion straight, slanting directly back from spine. Prosternum 1.4 to 1.5 times wider than long. Distance between chitinous spots 3.25-3.27 times dental line; 2.5-2.66 times the width at level of bottom of sinus (Plate 2, fig. 1). First dorsal plate 1.63-1.66 times wider than long. Especially the more caudal dorsal plates conspicuously depressed transversely in front of the caudal border and within and parallel with each lateral border, especially of caudal portion, of plate and the edges outside the depression more or less bent upwards. Plates not roughened: Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced, the processes rather wide and distally narrowly rounded, not acute. Coxal pores small, circular: 4, 4, 4, 4; 4, 3, 3,3; 4,3, 4,4; 4,3, 3,4; etc. ' CHAMBERLIN: THE GOSIBIIDAE. ONE Spines of first legs, 003,83 (M) or, {3-3-3 (observed only in 9); of the second to eighth, (3:33; of the ninth, cosas; of the tenth to thirteenth, mona of the penult, P3235 (7) or, gr 333 33:5 (mostly Q); of the anal, ¢7#434 or, i337 the claw single. None of the coxae laterally armed. Posterior legs of females short and very slender, not at all thickened. In the anal legs of the male the fourth and fifth joints are conspicu- ously expanded laterally but are not thickened dorsoventrally and thus appear flattened; the fifth joint as a whole markedly bowed ventrad and longitudinally furrowed along the dorsal surface, the fourth joint also furrowed but not so deeply so; fourth and, especially, the fifth joints with numerous straight hairs, the other joints being more sparsely clothed; sixth joint abruptly much more slender than the fifth (Plate 2, fig. 2, 3). Penult legs of male with fourth and fifth joints somewhat enlarged, the fifth joint decidedly more elevated distad and bearing upon dorsal surface toward mesal side of anterior end of elevated portion a low rounded eminence (Plate 2 fig. 4). Gonopods of male distinctly exposed; flattened; distally sub- truncate; bearing many hairs. Claw of female gonopods long, moderately curved, excavated and thin, acute. Spines 3 + 3, conical from base, acute; ectal one much longer than most mesal. Inner side of first article well chitinized, bent ectad at base but not truly excavated. Males clearly exceeding the females in average size, ranging from 21 to 30 mm. (mostly 25-30 mm.) in length as against 16-21 mm. (mostly 18-20 mm.) for the females. In a male 30 mm. long the eighth and tenth plates are 3.2 mm. wide; and in a female 20 mm. long the width of eighth plate is 2.2 mm. Praematurus.— Light brown, head somewhat darker and of dilute chestnut tinge. Antennae like head as are also prosternum and prehensors or those a little lighter. Venter pale, often but not al- ways of a strong greenish tinge. Legs pale, uniform. Antennae as in adult. Ocelli 1 + 3,3,1; 1+3,3,2. Organ of Témésvary in outline smaller than largest seriate ocelli and larger than the smallest ones. Prosternum as in maturus, on the teeth adjacent to sinus in some relatively smaller. Spines a little more slender. Coxal pores 4, 3, 3, 3; 3, 3, 3, 3. Spines 3-+ 3 or sometimes 3+ 2 and more rarely only 2 + 2, 218 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. but the most mesal spine on each side much smaller, the median one of each side intermediate; all more slender and acute than in maturus. Spines of first legs, ('3:3:3; of second to fifth, 0 eeRt of sixth and seventh the same or, ETT of the other legs same as in adult. Claw of female gonopods shorter than in adult and only slightly curved distad or in some essentially straight; acute, pale. Length 15-17 mm. Immaturus.— Dorsum pale Preeti brown. Head commonly a little darker and a clearer brown. Antennae light brown to dilute chestnut. Venter more distinctly greenish than the dorsum. All legs pale. Body apparently more decidedly narrowed caudad from the middle than in maturus. Antennae composed of nineteen to twenty articles; articles five, six, and seven, and twelve and thirteen shorter than adjacent ones. Ultimate article proportionately long, equalling or exceeding in length the three preceding ones taken together. Ocelli 1+ 3,2,1; 1+2,2; 1-+2,1. Single ocellus clearly largest. Prosternal teeth 3 + 3 but the most mesal on each side minute or indicated by a mere point. Spines distally more slender and bristle- like than in the adult. Coxal pores, 3, 2, > a ee EWI ep 2, 2.1 ), 0, 0,.3,.2 Spines of first legs, ¢ (2-3-3; of the second 4 tees4 or dorsal spines 0, 0,3, 2,2; of the third to seventh, jt a4 3-3; of the others as in maturus excepting for lack of dorsal spine on coxae of ninth to thirteenth and sometimes fourteenth legs. The dorsal spines of coxae of anal and penult legs very small and slender. Claw of female gonopods short, pale, acute, not at all curved. Spines 1 + 1; appearing as minute, acute points. Length 11-14 mm. Pullus IV.— Pale brown. Head and antennae a little darker, of chestnut tinge. Venter lighter than dorsum. Legs all pale. Antennae composed of sixteen articles, of which the fifth and sixth and the tenth and eleventh are distinctly shorter than the adjacent ones. Ocelli 1 + 2,1. All pale. Single one largest. Prosternal teeth 2-++ 2. Sinus subsemicircular. Spine short, acute, more slender than in older stages. A single pore on each twelfth coxa. CHAMBERLIN: THE GOSIBIIDAE. 219 . 0,0,1,1, 1 0,0,1,2,1 Spines of first legs, JULI; of the second, 0 Sis SOL nai os of pe third to fifth, 0734; of the seventh, } CoOL; of the eighth, oois3 or ventral spines may be 0, 0, 2, 3,3; of the ninth, oo gS OF ventral spines 0, 0, 2,3, 3; of the tenth, ee " 353 of the eleventh and twelfth, Ppa4:4- Anal glands very large and distinct. Length circa 7.75 mm. Typr.— M. C. Z., No. 552, California: Friant. Paratypes.— M. C. Z., No. 545-548, 550-551, 553, California: Friant; M.C. Z., No. 549, Fresno. R. V. andS.C. Chamberlin. Numerous specimens were taken at Friant under stones in the open near the mouth of the canyon. The conspicuous difference in size between the sexes suggests that occurring in Pseudolithobius megaloporus. GosiBius (Gosiprus) MonIcUS Chamberlin. Ann. Ent. soc. Amer., 1912, 5, p. 146. Diacnosis.— Brown with a darker middorsal stripe; head light ferruginous. Articles of antennae above twenty-nine in number. Ocelli fifteen to seventeen in four series. Prosternal teeth 2 + 2. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced. Basal spines of female gonopods 2 + 2. Dorsal spines of anal legs 107 3.2,2 (or 12). ventral. 0) 1, 3,3, 1; aN two. Dorsal spines of tenth or eleventh to penult legs 1, 0, 3, 2, 2; of third to ninth or tenth, 0,0, 3, 2,2. Ventral spines of fifth to ninth legs 0, 0, 2, 3, 3; of tenth and eleventh 0, 0, 3, 3,3; of twelfth and thirteenth 0, 1, 3, 3,3; of penult 0, 1,3, 3,2; of first to fourth 0,0, 2,3,2. Length 14-15 mm. DescriPtion.— Dorsum brown with a darker median longitudinal stripe which shows a marked tendency to spread laterad at the caudal end of each plate and may reach the lateral margins along the caudal border. Head subferruginous, the median longitudinal stripe of dorsum of body continuing upon it as far as a little caudad of frontal suture where it ends abruptly at a pale transverse band along the suture. Antennae similar to head. Prosternum and _prehensors pale ferruginous. Venter yellow or testaceous, the caudal segments darker, somewhat ferruginous. Legs yellow or testaceous like venter, nearly uniform or the caudal pairs slightly darker dorsally. 220) BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body rather robust, in type being only seven times longer than width of tenth plate. Moderately narrowed cephalad; strongly and uniformly narrowed caudad. Eighth and tenth plates of same width or the eighth a little wider, these clearly wider than the head. First plate slightly narrower than the third in female. Widths of head and of first, third, eighth, and tenth plates to each other as 53:47: 48: 57: 57. Head subcordate. Wider than long in ratio 47:45. Caudal margin mesally gently incurved, the sides conspicuously converging from marginal breaks caudad and rounding about corners. A median longitudinal sulcus extends cephalad from the suture to a transverse depression between the antennae. A short, deep transverse sulcus a little in front of caudal marginal elevation, the same more weakly indicated farther laterad on each side. Head smooth and shining, not punctate or rugose. In types the antennae have their ends broken off so that precise number of articles cannot be determined; but at least twenty-nine are present. The articles present are of nearly same form and pro- portions as in G. paucidens. Ocelli fifteen to seventeen in four series: e¢. g., | + 4,4, 4,2; 1 +5, 4,4,2; 1+4,4,4,4; 1+4,4,3,4. Much as inG. paucidens. Prosternum 1.6 times wider than long. Distance between chiti- nous spots 2.1 times width at sinus; 3.9 times dental line. Teeth 2+ 2, stout, conical, relatively larger than in paucidens. Spine long, distally drawn out in bristle-like form, its tubercle contiguous with base of outer tooth, attached on ventral side of edge. First dorsal plate smooth and shining like the head or only obscurely roughened. Other plates more or less roughened, the more caudal ones most strongly so. Major plates with the usual short transverse submarginal sulci near caudal third, the seventh having an additional similar one on each side at about middle of length. A longitudinal sulcus on each side midway between median line and lateral margin, the suleus at about beginning of the middle third sending a more clearly impressed branch directly mesad where it often unites with corresponding one of other side; often also a shorter sulcus running from near anterior margin obliquely caudoeetad toward point of origin of this transverse branch. Ventral plates appearing smooth and shining. The usual three longitudinal impressions indicated with varying degrees of distinct- ness. On several of the caudal sternites the median mark may end caudad in a deeper pit-like and somewhat transverse depression a little in front of the caudal margin. CHAMBERLIN: THE GOSIBIIDAE. 22h: Coxal pores circular and small: 4, 3, 3, 3 in types. . 0,0, 2,21 00.321, —¢ . Spines of first legs, ¢-9:s-3-9; of the second, 9:973:3-3; of the third 3, 2,2 . 0,0, 32,2 and fourth, (333; of the fifth to ninth, jqogr3; of the tenth and aed : ; eleventh, §):3-33; or coxal spine may be absent from tenth; of the . 1, 0, 3, 2 2 1,0,3,2.2, ¢ twelfth and thirteenth, j3:3:3; of the penult, 97-335; of the anal, By Bee ¢ a . 0: ere claws 2. Last four pairs of coxae laterally armed. Dorsal spine of eleventh coxae small. Anal legs of female short and moderately slender; the dorsal surface of femur and tibia distinctly longitudinally furrowed, the same articles of penult legs, especially the femur, similarly but less strongly fur- rowed. Male unknown. Gonopods of female long. Claw large, of usual form. First article strongly constricted at base leaving a conspicuous lobe distad pro- jecting mesad and touching corresponding lobe of the other gonopod, the anterior edge of these lobes at right angles to longitudinal axis of body or nearly so; mesal side strongly chitinized as in G. paucidens. Spines 2 + 2; stout; inner smaller than the outer one on each side. Process of sternite first clavately widening, presenting a point on each side, and then narrowing distad and becoming very slender and acute. Length of types (9 2) 14-15 mm. The specimen 14 mm. long has anal leg 5.8 mm. long and tenth plate 2 mm. wide. Type.— M. C. Z. No. 578, California: Santa, Monica, June, 1909. This species is manifestly close in many features to G. paucidens. It differs clearly, however, in much smaller size, in having the anal legs armed with two distinct claws, in ventral spines of anal legs, in dorsal spines of anterior pairs, in having the basal spines of female gonopods 2 + 2, ete. Gosrpius (TmM1oBIUS) INTERMEDIUS, sp. nov. Plate 3, fig. 2,3. DraGnosts.— Brown; head dilute chestnut. Marginal breaks of head small and inconspicuous. Antennae moderate; articles twenty-eight. Ocelli seventeen in four series. * Prosternal teeth 3 + 3 Posterior angles of sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced. Claws of anal legs two; dorsal spines 1,0, 3, 2,0; ventral 0, 1,3,3,1. Dorsal spines of ninth or tenth apap BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. to penult legs 1,0,3,2,2; of first or second to eighth or ninth 0, 0,3, 2,2. Ventral spines of thirteenth and penult legs 0, 1, 3, 3, 3; of fourth to twelfth, 0, 0, 3, 3, 3; of first to third, 0, 0,2, 3,3. Last four pairs of coxae laterally armed. Anal legs in male slender, unmodi- fied. Penult legs with a low rounded eminence on dorsal surface at distal end of tibia. Length 19 mm. DerscripTion.— Brown, the first plate darker. Head darker than dorsum, in type dilute chestnut. Antennae light or yellowish brown. Legs pale, the caudal pairs more densely pigmented, yellowish. Prosternum and prehensors dilute chestnut. Venter pale testaceous, the caudal plates darker, burnt brown. Body very conspicuously narrowed both cephalad and caudad from the eighth segment. First and third tergites equal in width. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 50: 46: 46: 66: 60: 51. Head broadly subcordate; wider than long in ratio 25: 23. Lateral margined breaks very small and inconspicuous. A sharply impressed transverse sulcus in front of caudal border, slightly convex; antero- laterad from each end of this a short, oblique impressed line. The usual broad and shallow longitudinal depressions in front of the transverse sulcus, these nearly parallel and extending to the anterior end of head. Antennae of moderate length. Articles in type twenty-eight; of these the first four or five are rather long, the others being moderate with a longer one occurring at intervals. Ultimate article shorter than the two preceding taken together. Ocelli seventeen in four series: 1 + 4, 4,4, 4. Single ocellus much largest. Of the seriate ocelli the most ventral are much the smallest; the dorsal ones conspicuously largest, somewhat irregular in form. Prosternum 1.6 times wider than long. Distance between chitinous spots 2.44 times width at level of bottom of sinus; 4 times the dental line. Teeth 3+ 3; small; the most mesal on each side smallest and lowest; line of apices of teeth, omitting innermost on each side, nearly straight. Sinus small, rounded at bottom. Spines small and hair-like. Sides of anterior portion of prosternum nearly evenly convexly rounded from spine to prehensor. First dorsal plate moderately narrowing caudad; 1.6— times wider than long. Major dorsal plates with the usual broad longitudinal depressions or furrows which diverge a little caudad; commonly each gives off anteriorly a branch which runs nearly straight cephalad to anterior margin. Plates roughened. Clothed with short, acute hairs CHAMBERLIN: THE GOSIBIIDAE. 228 which are more numerous toward lateral margins and which arise mostly from small tuberculiform elevations. The seventh to tenth plates especially with the lateral margins distinctly denticulate or serrulate, the teeth on seventh to ninth longest and most acute; on the plates each way from these the irregularities become more crenu- late and weak and there disappearing. Marginal hairs spring from the incisions caudad of the teeth (Plate 3, fig. 2). The fourteenth plate is narrowest a little cephalad of caudal end, the sides from there then again weakly diverging caudad; caudal margin mesally incurved (Plate 3, fig. 3). Posterior angles of sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced, but those of the sixth rather weakly so; processes broad, becoming more acute on the caudal plates. Coxal pores circular, of moderate size: 4, 4, 4, 4. , 0, 2, 3, 2 0, 0, 3, 22, ° 0, 0, 3, 2, Spines of first legs, 9: 9:3:3°3 OF 0° ae a ; of second and third, 9°9°5 of fourth to eighth or ninth, 0:33:33 1, 0, 3, 2, 2, 2. 3) 0-0-3-3-3, Of thirteenth and penult, 9:7 0 a : 0,2,3,3 2, + oh ninth or tenth to twelft Tere, claws 3; of anal, p¢s34, claws 2. Last four pairs of coxae Peat armed. Anal legs in male slender, without special modifications. Penult legs also slender but with the tibia bearing a low rounded lobe or elevation on dorsal surface at distal end. Length of type (o7) near 19 mm.; width of eighth plate 3 mm. Type.— M. C. Z., No. 579, California: Santa Barbara, March, 1913. Paratypes.— M. C. Z., No. 580, California: Santa Barbara, March, 1913. One mature and two not fully mature males. The serrulation of part of the dorsal plates and the unusual form of the fourteenth plate in this form are features of especial interest. GosiBius (ABATOBIUS) ARIZONENSIS, sp. nov. Plate 27 fig, 5:\ Plate! 3), ‘he.. 1. D1aGnosts.— Testaceous to brown, head not differing. Antennae with articles twenty-five to pene ene: Ocelli seventeen to nine- teen in four series. Prosternal teeth 2+ 2. Posterior angles of none of dorsal plates produced. Basal spines of female gonopods 2+ 2. Anal leg with claw single; dorsal spines 1, 0, 3, 1,0; ventral 0, 1, 3,2, 1. Dorsal spines of twelfth to penult legs 1, 0, 3, 2,2; of 224 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. all others 0, 0,3, 2,2. Ventral spines of penult legs 0, 1, 3, 3, 2; of thirteenth 0, 1, 3, 3,3; of eighth to twelfth, 0, 0,3, 3,3. Last four pairs of coxae laterally armed. Anal legs in male type missing. Penult with tibia enlarged distad as usual. Length 18-22 mm. DESCRIPTION.— Dorsum testaceous to brown, the head the same, but fifteenth plate abruptly paler; the caudal borders of plates and head may be darker as they may also be in a longitudinal median stripe. Antennae yellow to yellowish brown. Prosternum and pre- hensors like head or a little paler. Venter yellow or brownish yellow, the caudal plates more densely pigmented. Legs similar in color to corresponding plates of venter, the posterior pairs somewhat orange. Body moderately narrowed cephalad from the tenth segment. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other (in co’ type) as 52: 52: 51: 58: 57: 55. Head wider than long in ratio 52:47. Smooth. Hairs sparse. Antennae moderately long, reaching upon the seventh segment. Articles twenty-five to twenty-eight; mostly long, those composing a varying proportion of distal portion short. Ocelli seventeen to nineteen in four series: e.g., 1 +5, 4, 4, 3; 1+ 4, 5,5,3; 1+5,4,5,4; 1+ 5,5,4,3. Single ocellus largest. Seriate ocelli conspicuously decreasing in size cephalad and ventrad, varying considerably. Prosternal teeth 2 + 2, small, well separated. Inner one on each side larger than the outer; line of apices a little procurved. Sinus between V-shaped and semicircular, the sides being concave and the bottom more or less rounded (Plate 3, fig. 1). Prosternum 1.6 times wider than long. Distance between chitinous spots 3 times width at level of bottom of median sinus; 4.4 times the dental line. Spines straight, slender, and acute, inserted a little ectad of tooth. Margin extending laterad almost horizontally a little distance from spine and then bending back caudoectad to prehensor. First dorsal plate equalling head in width; 1.7 times wider than long. Posterior angles of none of the dorsal plates produced; posterior corners of ninth shortly obliquely excised, those of eleventh and thirteenth straight or excised on one side. Major plates mostly showing a median longitudinal furrow, and one diverging caudad from it on each side; transversely depressed in front of caudal border; the short, transverse, submarginal sulcus on each side near beginning of caudal fourth of length. Coxal pores very small, circular; porigerous surface depressed: 4,5,4,4; 4,5,5,5; 4,5,5,4; 5,5,5,5; 4, 5, 6, 5. t CHAMBERLIN: THE GOSIBIIDAE. 225 Spines_ of bist to seventh pairs of legs, pg333; of elehth to 0, 3,22 0, 3, 2, 2 eleventh pase ses; of twelfth, eos ag; Of thirteenth, (73:33; of 0, 3, 2 BONS ROG 4 penult, (33:35; of anal, oy3:24, with one claw. Last four pairs of coxae laterally armed. Penult legs of male slender; the tibia enlarging distad as usual. (Anal legs missing in o of types). Claw of female gonopods long and acute, flattened and moderately curved. Inner side of first article proximally bending ectad, well chitinized; joint constricted or broadly channelled across base (Plate 2 fig. 5). Length 18-22 mm. No. 581, Arizona: Fort Williams, January, 1911. W. M. Wheeler. Paratypes.— M. C. Z. No. 612, Arizona: Fort Williams, W. M. Wheeler; M. C. Z. No. 611, Grand Canyon, W. M: Wheeler; M. C. Z. No. 610, Huachucha Mts., W. M. Wheeler; M. C. Z. No. 608, 615, Arizona, H. K. Morrison. Goststus (ABATOBIUS) MONTEREUS, sp. Nov. Plate 1, fig. 5. DraGnosts.— Dark brown to chestnut, head of nearly same shade. Antennae short, articles twenty-three to twenty-seven. Ocelli fifteen or sixteen in four or five series. Prosternal teeth 2 + 2. Posterior angles of none of the dorsal plates produced. Basal spines of female gonopods 2+ 2. Claws of anal leg two; dorsal spines 1, 0, 3, 2, 0; ventral spines 0, 1,3,3,1. Dorsal spines of penult and thirteenth legs 0, 0, 3, 2,2 or 1, 0,3, 2,2; ventral, 0, 1,3, 3,2. Ventral spines of twelfth legs 0, 0,3, 3,2; of fourth to eleventh 0, 0, 2, 3,2; of first, 0,0, 1,3,1. Last one or two pairs of coxae laterally armed. Length of female 15-17 mm. DescrieTion.— Dorsum dark brown to chestnut, darker along a median dorsal line. Head of nearly same shade as dorsal plates or, in paler specimens, darker. Antennae chestnut, uniform. Leg light brown excepting the posterior pairs which, excepting proximally, are dusky, appearing blackish brown or dark chestnut. Prosternum and prehensors chestnut, sometimes more of a dark brown. Venter brown to testaceous. 226 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body typically strongly narrowed cephalad from the eighth segment, with the first segment distinctly narrowest. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 36: 32: 34: 44: 45: 44. Head suborbicular, widest at level of lateral breaks; wider than long in ratio 18:17. A semicircular depression in front of the caudal border, its ends continuing cephalad as more obscure and broader diverging furrows or depressions. Punctae fine and sparse. Antennae short, strongly attenuated distad. Articles, so far as noted, from twenty-three to twenty-seven; decreasing distad from the second, and most moderately short in antennae with the lesser number of articles to very short and crowded in those with the larger number. The ultimate article typically longer than the two preced- ing ones taken together. Ocelli fifteen to sixteen in four or five series: 1 + 4, 4,3}3; 1+ 1, 4,4, 3, 2, the ocellus of first series at anterior end of patch; 1 + 4, 3, 3, 3, 2. Single ocellus very much the largest. Ocelli of top series larger than others, the ocelli of others decreasing cephalad and ven- trad as usual. Prosternum 1.54 times wider than long. Distance between chiti- nous spots 2.4 times width at level of bottom of sinus; 3.7 times the dental line. Median sinus very shallow; wide and semicircular. Teeth 2 + 2; equal in size or inner one of each pair a little the larger, moderate, acute; line of apices recurved. Spines fine and hair-like, inserted caudad of margin close to ectal tooth. Margin rounding _ out ectad from teeth on each side as usual and then bending back straight ectocaudad. First dorsal plate widest cephalad, its sides moderately converging caudad; 1.68 times wider than long. Other major dorsal plates showing more or less clearly a longitudinal furrow each side of the middle line from which it diverges a little caudad. Under lens these plates, excepting the first one or two and the fifteenth, those of the posterior region especially, are seen to be roughened, tuberculo- rugose, most strongly so on caudal and lateral portions of plate. Hairs short. Posterior corners of none of the dorsal plates produced; those of ninth, eleventh, and thirteenth rounded or obliquely excised. Coxal pores small: 3, 3, 3,3; 3,3,4,3; 3,4, 4, 3. eae of first legs, $:j-3:7 or dorsal spines 0,0, 2,1,1; of the second, (-g-y-33 to Goose; of the third, OO to t¢e33; of the fourth to eleventh, (0-333; of the twelfth, (03:33; et the thir- teenth and penult, (13-35 or, (73:33; of the anal, (3:34, claws two. CHAMBERLIN: THE GOSIBIIDAE. P| Last one or two pairs of coxae laterally armed, the spine on penult, when present, small. Claw of female gonopods long and acute, excavated and strongly curved. Basal spines 2 +2; long and moderately stout, conically | acuminate from very base and uniformly so or a little more abruptly narrowing toward tip. First article conspicuously excavated on mesal side at proximal end. Process of sternite distally, truncate or convex. Length 15-17 mm. (29. The co unknown). Type.— M. C. Z. No. 582, California; Pacific Grove, March, 1913. PSEUDOLITHOBIUS Stuxberg. Ofvers. K. vet.-akad. Férhandl., 1875, no. 3, p. 8; Proc. Cal. acad. sci., 1877, 7, p. 137. Latzel, Myr. Ost.-Ung. Mon., 1880, th. 1, p. 35. Boll- man, Bull. 46, U.S. N. M., 1893, p. 164. Verhoeff, Die Chilop., 1907, p. 240. Chamberlin, Pomona college journ. ent., 1910, 2, p. 369; Can. ent., 1911, 42, p. 377.. Head with lateral margins essentially continuous, no true breaks occurring. Antennae short; articles twenty to twenty-two. Eyes composed of seriate ocelli. Prosternal teeth 3 + 3 or4 +4. Spine near ectal angle of anterior margin, stout and subdentiform. Pores present on last five pairs of coxae; circular and uniseriate. Penult legs in male with the fifth article enlarging distad as in Gosibius (Plate 3, fig. 5). Anal leg with fifth article conspicuously longitudinally furrowed dorsally and bowed ventrad as in some species of Gosibius (Plate 4, fig. 1). Dorsal spines of anal legs 1, 0, 3, 2,2; ventral, 0,1, 3,3, 1; (claws in type species 2). Dorsal spines of penult legs 1, 0, 3, 2,2; ventral 0, 1, 3, 3, 2 or 0, 1, 3, 3,3. (Third and fifth joints of all legs cephalad of fourteenth with three ventral spines in type species). Gonopods of male short; uniarticulate. Claw of female gonopods large and entire. Basal‘ spines stout; (in type species 3 + 3 or 3+ 4). First article broad, inner side well chitinized, constricted at base as usual in the family. Type.— P. megaloporus Stuxberg. This appears to be the only species of the Lithobioidea to bear pores on ail the last five pairs of coxae; but in the Henicopidae, Zygethobius agrees in this distribution of the pores. 28 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Fig. 2. Distribution of Pseudolithobius. bo bo We) CHAMBERLIN: THE GOSIBIIDAE. PSEUDOLITHOBIUS MEGALOPORUS (Stuxberg). Plate 3, fic.'4, 5; Plate 4) fig. 1, 2: Lithobius megaloporus Stuxberg, Ofvers. K. vet-akad. Férhandl., 1875, no. 2, p. 69; Ann. mag. nat. hist., 1875, ser. 4, 15, p. 190. Lithobius (Pseudolithobius) megaloporus Stuxberg, Ofvers. K. vet.-akad. Forhandl., 1875, no. 3, p. 14; Proc. Cal. acad. sci., 1877, 7, p. 137. Pseudolithobius megaloporus Chamberlin, Pomona college journ. ent., 1911, 3, p. 470; Can. ent., 1911, 42, p. 381. D1acnosis.— Dorsum brown, first tergite and caudal borders of others often darker; head darker and more reddish. Antennae short, articles twenty to twenty-two. Ocelli only five to seven in two or three series. Prosternal teeth 3-+3 or 4+ 4. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. Basal spines of female gonopods 3 + 3 or 4+4. Dorsal spines of tenth to anal pairs of legs 1,0,3,2,2. Ventral spines of anal legs 0, 1, 3, 3, 1; claws two but the accessory one minute. Ventral spines of tenth to penult legs 0, 1,3, 3,3 or those of penult sometimes Ot Bt3) ONCE the others 0, 0, 3, 3, 3 with dorsal spines 0, 0, 3, 2,2. Last two pairs of coxae laterally armed. Length of female up to 31 mm.; of male up to 41 mm. Description.— Dorsum brown, often reddish, the first tergite commonly darker and more reddish and the other tergites also fre- quently darkened along the caudal borders. Head and prosternum with prehensors usually but not always darker and more reddish than the dorsum. Antennae brown to reddish brown, uniform. Venter lighter brown, mostly devoid of any red tinge excepting on caudal segments, sometimes with olivaceous cast. Legs light brown, mostly uniform, the caudal pairs in some a little darker than others. Body slightly widest at about fifth segment from where it is nearly of the same width caudad to the tenth segment; only little narrowed cephalad of fifth segment, the sides thus being nearly parallel over most of length. Males proportionately more slender than females, being commonly 10 times longer than width of tenth plate, as against 8.5 times in the female. Head nearly equal in length and breadth but commonly a little wider (20:19). Widest a little in front of the well-rounded caudal corners from where nearly of same width forward to middle. Strongly margined caudally and along the sides. Commonly elevated in front of suture and complanate behind it; a short median longitudinal 230 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. furrow bisecting the elevated portion. Finely punctate and weakly irregularly rugose. Antennae short. Articles twenty to twenty-two, of which the arrangement is usually about as follows: — the first four long, the seventh and eighth short with the fifth and sixth intermediate; the ninth, tenth, and eleventh long, the twelfth and thirteenth inter- mediate and the fourteenth and fifteenth very short; the next three long, the nineteenth intermediate, the twentieth and twenty-first short. That is, the antennae thus show several successive divisions beginning with long articles and ending in short ones which are mostly in groups of two, the seventh and eighth and fourteenth and fifteenth especially being normally very short. Eye area small and elongate in form. Consisting mostly of five to seven, but sometimes up to nine ocelli arranged in two series, occasionally im three series: ¢. g.. 1+3,2; 1+3,3; 1+ 4,4; 1+ 3, 3,2. Organ of Toémésvary in outline small. Anterior portion of prosternum trapeziform, the lateral margins being substraight. Teeth 3+ 3 or 4+ 4, the innermost tooth on each side is typically on side of smus; outermost tooth on each side a little more widely removed. Spine stout and conical, inserted a little proximad of anterior margin on ventral side. Prosternum 1.66 times wider than long. Chitinous lines strongly marked cephalad, but fading out caudad. Dorsal plates depressed longitudinally immediately mesad_ of turned up lateral edges, median portion not strongly convex, caudal portion depressed. A weak longitudinal furrow each side of middle diverging from its mate caudad; from its caudal end or near it a second furrow diverges from it cephalad. Surface moderately uneven or rugose. The major plates unusually long, longer relatively in the male than in the female as shown by the following measurements :— Male Female Tergite No. Length Width Length Width 1 2.6mm 4mm. 2mm 3.1mm 3 4 4.1 3 3+ 5 4.9 4.2 Sel! 352 7 4 4.1+ 22 3367) 8 4.1+ 4.1+ 3 3.2 10 5) 4.1 Ben See 12 5 4 3+ 3+ (The total length of male measured 41 mm.; of female 27 mm.). CHAMBERLIN: THE GOSIBIIDAE. Tal The ventral plates are also long and similarly relatively longer in the male than in the female. Measurements from the same two individuals are as follows: — Male Female Sternite No. Length Width Length Width il 1.1mm 1mm. .6mm. 1mm. 2 2 1.6 ileal 1.8 3 2.4 eal 1.4 2 4 2.6 2a. 1S eh 2 5 2.8 2h Noe 2.1 6 3— 29 1.9 2.2 uf 3.1— 2.3 2.1 2.8 8 aia IL PAE Yh 2.1 PPA 9 aye 2.9 2 2.3+ 10 3.2 3 2.1 220 11 one 2.8 1.6 hall Wea 3 2.8 1.3 esi 13 2.9 3+ 1.2 Pond 14 1.5 2.1 iil 2 From this table it will be seen that, excepting the last few, the sternites in the male are all longer than wide whereas in the female they are wider than long. The ventral plates are bowed ventrad, from end to end; the anterior end of outer more anterior plates more strongly bent dorsad and making a distinct angle, which in some is nearly right, with adjoining portion of plate. Depressed within each lateral edge. Median and caudal plates with a median and two . lateral longitudinal sulci which do not extend over anterior portion of plate; especially the median sulcus disappearing in more cephalic plates. Plates irregularly roughened and densely punctate. Coxal pores moderately large, circular or a little transversely elon- gate or elliptical; their edges elevate and strongly chitinized; poriger- ous surface depressed, the bordering rims well chitinized: 4, 4, 4, 4, 4; Dosa o. De uoyon(O)e oso). ties; of the tenth to thirteenth, Shines of first to ninth legs, or : 33; of the pen fis33 or with ventral spines 0, 1, 3, 3, 2, 0, 3,22 claws two; of anal, §°3:3:7, claws two but the accessory one very small. Last two pairs of coxae laterally armed. Anal and penult legs in female slender, their prefemora, as also in the male, strongly laterally compressed, the immediately preceding ones also showing this compression, broader dorsally than ventrally, ‘ Fae, BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. a section being wedge-shaped. In the male the tibia of anal legs is widened and is conspicuously bowed ventrad, furrowed longitudi- nally above (Plate 4, fig. 1). In the penult legs of male the tibia is elevated over distal half of length (Plate 3, fig. 5). Gonopods of male short and broad, rounded; bristles long. Claw of female gonopods very long, strongly curved; blackish. Basal spines 3 + 3 or 3 + 4; conically acuminate from base or near it, decreasing in size from outermost to innermost. Proximal article broad, constricted at base (Plate 8, fig. 4). Length of males up to 41 mm.; of females up to 31 mm. A male 30 mm. long has antennae 11 mm. and anal legs (exclusive of coxae) 12.5 mm. long. Agenitalis.— Stuxberg’s original description was manifestly based upon very young specimens, these being of the agenitalis stage as indicated by several points in his description. The original descrip- tion, rearranged, is as follows:— Lamina cephalica obcordata, hirsuta, margine postico subrecto, elevato, parte antica sulco profundiore a postico sejuncta. Antennae perbreves, articulis 19-20 compositae, ex quibus ultimus longissimus, tres antecedenta junctos longitudine aequansy 20.0. ; 8°, 9°, 10°, 14°, 15°, 16° mediocribus, ceteri latitudine duplo majore quam longitudine minime. Oculi ocellis 7 magnis in 2 series (1 + 3, 3) digestis. Coxae pedum maxillarium secundi paris dentibus 2 + 2-3 +3 crassis acumine nigris armatae, sinu mediano lato, hand profundo. Seuta dorsualia 9%™, 11¥™, 13¥™ angulis productis, 2%, 44, 62™, 7"™ margine portico recto, 1°, 98” convexo, EE yD ees cal (Di 14"™ sinuato. Scuta ventralia omnia, praesertim marginibus, setis longis, magnis vestito, medii corporis sulco profundiore longitudinali mediano laterali minore et breviore, ex angulo postico laterali excurrente. Pori coxales 2, 2, 1, 1, 1 maximi, rotundi. Pedes primi paris calcaribus 1,1,1. Pedum analium articulus primus calearibus nullis, setis 2 longioribus, altera ventrali, altera laterali armatus. Pedes anales perbreves, longitudinem antennarum non assequentes, incrassati, ungue singulo, calearibus 0, 1, 1, 0 armati. Color dorsi testaceo-brunneus, laminis ventralibus pedibusque pallidioribus. Longitudino corporis 12 mm.; antennarum 4 mm.; pedum analium 2.5 mm. CHAMBERLIN: THE GOSIBIIDAE. 233 Type Locauity.— California: near San Francisco (G. Eisen). Also taken in California at Sausalito, (M. C. Z., No. 586, 587); Oro- ville, (M. C. Z. No. 584, 585). The type locality, “near San Francisco”, was probably Sausalito, from where numerous specimens have been examined. I found the species to be very common about Oroville. In April it occurred under stones and other objects lying in open treeless areas. They seemed slow to take alarm, often remaining quite unconcerned after stones had been removed from over them and they themselves touched or jarred, and appearing generally more sluggish than most related forms. The conspicuous difference in size and form of the sexes in this species is noteworthy. GuampBius Chamberlin. 7* Ann. Ent. soc. Amer., 1912, 5, p. 144. Head with lateral marginal breaks. Antennae short or moderate; articles from twenty-four to thirty- five. Eyes composed of seriate ocelli; single ocellus enlarged. Prosternal teeth 2 + 2; sinus of good size, mostly between V- and U-shaped. Spines always more slender than the teeth, varying from truly spiniform to the more slender, distally bristle-like form. Either posterior angles of ninth, eleventh, and thirteenth dorsal plates produced (Sibibius) or those of eleventh and thirteenth, at least of latter, more weakly produced (Guambius sens. str.). Anal legs in the male with the fourth article more or less conspicu- ously crassate and longitudinally furrowed above; the fifth article commonly much less crassate, somewhat elevated along dorsal side but not conspicuously modified. Fifth article of penult legs obliquely planate or excised at distal end above and there bearing a conspicuous lobe or crest of characteristic uy (Plate 4, fig. 5; Plate 5, fig. 3, 5, 6; Plate 6, fig. 1). Dorsal spines of anal legs 1, 0,3, 2,1 or 1, 0,3, 2,2 (Sibibius) to 0, 0, 3, 1,0 or 1,0, 3, 1,0 (Guambius sens. str.); ventral 0, 1, 3, 3, 1 to 0,1,3,2,0; claws two or three. Dorsal spines of penult legs iGaar 2 2-ors0sOrar 2 2 49) and 7,0, 3, 2,2 or 0,0, 372) 1(e) (Sibibius) to 1, 0, 3, 1, 1 or 0, 0, 3, 1, 1 (Guambius sens. str.); ventral, 0, 1, 3,3, 2 or 0, 1,3, 3,1; claws three. Dorsal spines of thirteenth 234 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Fig. 3. Distribution of Guambius. CHAMBERLIN: THE GOSIBIIDAE. 230 legs 1, 0,3, 1,1 or 0,0, 3, 1,1 to 0,0, 3, 2,2; of twelfth 0, 0, 3, 2,2 to 0,0,3,1,2. Last one or two pairs or none of posterior coxae laterally armed. Gonopods of male uniarticulate. Claw of female gonopods large and strictly entire. Basal spines stout, 2-+2. Mesal side of first article well chitinized; article con- stricted or furrowed about base, commonly appearing excavated on mesal side at proximal end as usual in the family. Length 9 to 15.5 mm. Type.— G. euthus (Chamberlin). Sibibius is removed from Arenobius, under which it was originally placed as a subgenus, chiefly because of the pronounced differences in the prosternum and in the modifications of the posterior legs. It 1s united tentatively with Guambius because of close agreement in the secondary sexual characters of the males, although the differences in the spining of the legs and in the angulation of the dorsal plates with other lesser ones make it seem likely that, with more thorough acquaintance with the chilopod fauna of the southern United States and Mexico, it will be found advisable or necessary to treat it as a separate genus. The distribution of the known species is indicated on (Fig. 3). Key to Subgenera and Species of Guambius. a. Posterior angles of only the eleventh and thirteenth dorsal plates produced, or production of these also but slight; dorsal spines of anal legs 0,0, 3, 1,0; of penult, 0, 0, 3, 1, 1. Guambius Chamberlin sens. str. b. Ventral spines of penult legs 0, 1, 3, 3, 2. G. euthus (Chamberlin). bb. Ventral spines of penult legs 0, 1, 3,3, 1° or 0,1,3,2,1. ~ c. Ventral spines of first legs, 0, 0, 0, 0, 1. G. pinguis (Bollman). ce. Ventral spines of first legs 0,0, 1, 1,1 or 0, 0, 1, 2, 1. d. 'Trochanter of thirteenth legs armed; ventral spines of anal legs 0, 1,3, 2,1 or 0,-1, 3, 3, 1. G. mississippiensis (Chamberlin). dd. 'Trochanter of thirteenth legs unarmed; ventral spines of anal! legs: 0; 1,/3, 2,0... 2G: curtior) sp. nov. aa. Posterior angles of ninth, eleventh, and thirteenth dorsal plates 236 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. produced; dorsal spines of anal legs 1 (10), 0,3, 2,1; of the penult, 1 (10), 3, 2,2 or 1 (10), 3, 2, 1.. .Sabibtws Chamberlin. b. Articles of antennae 30-35; ventral spines of thirteenth legs normally 0, 0, 3, 3, 2; claws of anal legs two G. coloradanus (Chamberlin). bb. Articles of antennae mostly 24-26; ventral spines of thir- teenth legs normally 0, 1, 2, 3,2; claws of anal legs three. G. oedipes (Bollman). GuAMBIUS (GUAMBIUS) CURTIOR, sp. nov. Plate 5, fig. 6; Plate 6, fig. 1, 2. DraGnosis.— Dorsum testaceous, caudal plates darker; head deep cherry-red. Antennae very short; articles twenty-five. Ocelli nine or ten in three series. Body about 6.3 times longer than width of tenth plate. First tergite two times as wide as long. Anal leg of male crassate, the fourth joiht particularly so. Penult legs more slender, the fourth joint most inflated; crest of fifth article subtri- angular in lateral view, highest distad (Plate 5, fig. 6). Anal legs with two claws; dorsal spines 0,0,3,1,0; ventral, 0, 1, 3, 2,0. Dorsal spines of penult legs 0, 0, 3, 1, 0; nen OM tao. Ios claws two. Dorsal spines of thirteenth legs 0, 0, 3, 2, 2; ventral, 0, 0, 3, 3, 2. Ventral spines of twelfth legs 0, 0, 2, 3, 2. Dorsal spines of second legs 0, 0, 1, 2,1; ventral same or 0,0, 1, 1,1. None of coxae armed. Length 8.5 mm. (male type). DeEscriPTion.— Dorsum testaceous, posterior plates and caudal margins of same anterior ones darker, more reddish. Head deep cherry-red. Antennae similar to head proximally, lighter distally. Prosternum dark but less reddish than the head. Venter pale testa- ceous, the anterior and especially the posterior plates darker. Legs pale testaceous, the posterior pairs more densely pigmented, more or less orange. Body wide, relative to length, being in the type 6.3 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 30: 27: 30: 37: 37: 33. Head as wide as long or only slightly wider. Of uniform width between eyes and marginal breaks. Hairs short and sparse. Antennae very short; articles twenty-five short and decreasing distad as usual, closely united. Hairs short. Ocelli nine or ten in three series: 1 + 4, 3,2; 1+ 3,3, 2. CHAMBERLIN: THE GOSIBIIDAE. 237 Prosternum 1.6 times wider than long. Distance between chitinous spots 2 times width at level of bottom of mesal incision; 3.1 times the dental line. Median sinus rather small, widely V-shaped with sides somewhat concave. Teeth acute, subequal; line of apices recurved. Tubercle distinct, contiguous with base of ectal tooth; spine slender, bristle-like. First dorsal plate 2 times wider than long; sides nearly straight, considerably diverging cephalad, rounded only at ends. Eleventh and thirteenth plates with the caudal corners distinctly produced; ninth plate an the corners aaa ora nts ae excised. second, 0.0,3,2.1 of he. fount and ith, none at ne the sixth and SANs eae; of the eighth, (232; of the ninth, -°222: of the tenth, oo ja3°9; Of the eleventh and twelfth, (2-333; of the thirteenth, 0, 0, 3, 2, 2 0, 0, 3, 1, 0 O33; of the penult, (4'237; claws two; of the anal, i320; Claws two. None of the coxae at all armed. Anal legs of male with the fourth joint in particular strongly in- flated and conspicuously longitudinally furrowed dorsally; fifth joint uniformly rounding up dorsally as usual; tarsi slender. Penult legs more slender than the anal, the fourth joint not relatively more inflated than the adjoining ones; fifth article obliquely dorsocaudally complanate or excised at distal end and their bearing a longitudinally placed, laterally compressed low crest, the dorsal edge of which begins proximally at surface of joint and rises to distal end, in lateral view being thus subtriangular (Plate 5, fig. 6). Gonopod of male bearing about four bristles. Length of type (co) 8.56 mm.; antennae 2.5 mm.; width of tenth plate 1.35 mm. Type.— M. C. Z., No. 605, Mississippi: Gulfport. GuaAMBIUS (GUAMBIUS) MISSISSIPPIENSIS Chamberlin. Plate 5, fig. 4,5: Ann. Ent. soc. Amer., 1912, 5, p. 144. Lithobius oedipes Chamberlin (non Bollman), Ann. Ent. soe. Amer., 1911, 4, p. 43. DiaGnosis.— Testaceous to clear brown; head cherry-red to al- most black. Antennae short; articles twenty-fivé or twenty-six. Ocelli nine to fifteen in three or four series. Body 7.25 to 7.85 times 238 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. longer than width of tenth plate. First dorsal plate near 1.8 times wider than long. Posterior angles of eleventh and thirteenth dorsal plates more or less produced. Anal legs of male strongly crassate, the prefemur elevated and deeply longitudinally furrowed. Penult ‘legs similarly crassate; tibia with lobe (Plate 5, fig. 5). Claws of anal legs two; dorsal spines 0, 0, 3, 1, 0; ventral, 0, 1, 3, 3, 1-0, 1, 3, 2, 1. Dorsal spines of penult legs 0, 0,3, 1,1; ventral 0, 1, 3, 3, 1- 0, 1, 3, 2, 1; claws two. Dorsal spines of thirteenth legs, 0,0, 3, 2, 2; ventral 0,1, 2,3, 2. Ventral spines of twelfth legs, 0, 0, 2, 3, 2. Dorsal spines of second legs 0, 0,1, 2,1; ventral the same. Dorsal spines of first legs 0,0, 1, 1,1; ventral, 0,0, 1,2,1. None of coxae armed. Length 10-14 mm. Description.— Dorsum dilute chestnut with the caudal plates and a median longitudinal line somewhat darker. Head and an- tennae dark chestnut, the latter paler distad. Venter testaceous, darker brown caudad. Prosternum and prehensorial feet nearly the same as head. Legs testaceous, the caudal pairs a little darker. Body strongly narrowed from eighth plate cephalad to the first which is clearly narrower than the third; eighth and tenth plates nearly equal in. width, much wider than the head, the latter being typically even a little narrower than third plate, widths of head and of first, third, eighth, tenth, and twelfth plates as 31: 29: 32: 41: 42: 39. Head equal in length and width or only slightly wider (36:35 or 34.5). Widest at marginal interruptions between which and the eyes the diameter is nearly uniform, the sides caudad of this convex and strongly converging; median portion of caudal margin straight. Rather strongly transversely furrowed or depressed immediately in front of caudal marginal thickening, the furrow extending entirely across the head and continuous with the caudal end of a wide longi- tudinal furrow on each side. A short. transverse sulcus a little cephalad of and parallel with median part of frontal suture, a me- dian longitudinal sulcus extending forward from this transverse one and showing or passing through a deeper pit-like depression at middle of length. Smooth and shining, or only very obscurely uneven. _ Antennae very short, reaching only to the fifth or sixth body-seg- ment; strongly attenuated. Composed of twenty-five or twenty-six articles of which the second is longest, those immediately following not abruptly shorter but only very gradually decreasing in size distad. Ocelli ten to fourteen in number arranged in three or four series: e. g., L On On Osi te Akt Ss ol a Oidy: 25, lias Oy ay Oaylie lo ae eoeale The single ocellus much the largest, subvertically elliptic or oval. CHAMBERLIN: THE GOSIBIIDAE. 239 Prosternum 1.5-1.55 times wider than long. Distance between chitinous spots near 2.23 times width at level of bottom of sinus; 3.35 times dental line. Teeth subequal, acute, with line of apices manifestly recurved. Median sinus wide and deep with its sides concave. Tubercle of spine well developed, contiguous with base of ectal tooth; spine slender and bristle-like, not much longer than the teeth. First dorsal plate with sides moderately diverging cephalad; 1.8 times wider than long. Plates under lens appearing finely and in part only obscurely roughened, the roughening as usual more distinct caudad. Major plates excepting the seventh, showing distinctly the short transverse submarginal sulcus on each side, the seventh plate showing a similar sulcus near each caudal corner and one near each anterior corner as well. Most of the major plates marked with two distinct longitudinal furrows which diverge more or less from near thé caudal margin cephalad, these furrows in some distinct only in front of the submarginal transverse marks. Posterior angles of eleventh and thirteenth plates a little produced, those of the others rounded or obliquely excised. Most ventral plates with a rather deep longitudinal furrow toward each side but mesad of a weaker submarginal furrow. Thé usual median longitudinal furrow with also the transverse depression cephalad of caudal margin more or less evident. Coxal pores small or moderate: 3,4,4,3; 4,5,5,5; 5,5, 5, 5; Db DONA 319,09570; oO: Spines of first legs, (tas; of the second, Corea; of the third 0, 0, 1, 2, 2 0, 0, 1, 2; 2 . same as second to, (jaa; of the fourth, jo4:g7 or ventral spines 0, 0,2, 2,1; of fifth same as fourth or ventral spines may also be Sc, é C 0, 0, 2, 2, 2 9 « 2 0, 0, 2,.2,2; of the sixth, g-o-s24 or ventral. spines 0, 0, 2, 2,2; of a aan ‘ the seventh, dp2a3 or dorsal spines 0, 0, 2,2, 2; of the eighth and 0, 0,2, 2, 2 RZ 252 ° ninth, ¢-p-s-2-93 of the tenth and eleventh, 333 or ventral spines 2 « 0, 0, 3, 2.2 ¢ ‘ . 0, 0, 2, 3,2; of the twelfth, (-p-s-35 or ventral spines 0, 0, 2, 3, 2; ot 0, 0, 3,.2, 2 é the thirteenth, (ag3'2; of the penult, usgq or ventral spines ¢ ¢ 0, 0, 3, 1, 0 =n ic Orla o2s4, claws: 25, ol. the anal, 7--.-4.,,er.ventrad 0; 1,3, 2,.1, claws two. None of the coxae armed either above. or laterally. Gonopods of male wide, truncate distad, flattened in an ectoventral dorsomesal direction; bearing a few short bristles in a row along distal edge of ventral side. Claw of female gonopods long and moderately stout, considerably 240 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. curved, subacute but apex narrowly rounded. Spines 2 + 2, the outer clearly longer than the inner, acuminate from base. First article with inner side sharply defined and strongly chitinized edge; considerably curving ectad proximally. Median process of sternite long and acutely acuminate, strongly chitinized. Length 10 to 14 mm. A male (type) 12.5 mm. long has antennae 5 mm. long and width of tenth plate 1.7 mm. The widths of head and of first, third, eighth, and tenth plates are to each other as 18: 17: 19: 24: 24. Typre.— M. C. Z., No. 604, Mississippi: Byram, July, 1910. ParatyPes.— M. C. Z., No. 603, Mississippi: Byram, July, 1910. GuaAmBius (GUAMBIUS) EUTHUS (Chamberlin). Lithobius euthus Chamberlin, Proc. Acad. nat. sci. Phil., 1904, p. 652. Diagnosis.— Dorsum brown, first and last plates more reddish; head darker reddish brown. Antennae short; articles twenty-five to twenty-eight. Ocelli about eight in three series. First dorsal plate 1.72 times wider than long. Posterior angles of eleventh and thirteenth dorsal plates produced weakly. Dorsal spines of anal legs, 0, 0, 3, 1,0; ventral, 0, 1, 3, 3, 0 or 0,1, 3,2,0; claws 2, the accessory onelong. Dorsal spines of penult legs, 0,0, 3, 1,1; ventral, 0, 1,3, 3, 2; claws three. Dorsal spines of thirteenth, 0, 0, 3, 2,2; ventral, 0, 1, 3, 3, 2. Ventral spines of twelfth legs, 0, 0, 2, 3, 2. Dorsal spines of second legs, 0, 0, 2, 2, 1; ventral, 0, 0, 2, 3, 1. Dorsal spines of first legs ORO; 2)4 0 or 050) 23>... 14 ventral, 0,0) 15 Ae. or. 0 Osi 2s I: None of coxae armed. Length 12-15 mm. DEscRIPTION.— Dorsum brown, the first and the last tergites darker and more reddish. Head darker reddish brown or chestnut. Antennae brown or like head, paler distad. Prosternum and prehensors like head. Venter brown. Legs light brown. Body distinctly narrowed cephalad to the first plate which is clearly narrower than the head and than the third plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 76: 69: 76: 82: 80: 77. Head wider than long (43: 38.) Caudal margin straight or nearly so. Narrowed from breaks caudad; moderately widely rounded cephalad. Antennae short; articles twenty-five to twenty-eight, moderate. CHAMBERLIN: THE GOSIBIIDAE. 241 Ocelli near eight in three series: e¢. g., 1 + 3,3, 2, which is the commonest arrangement. Prosternal teeth 2 + 2; the mesal one of each pair distinctly larger than the outer and the line of apices mostly a little procurved. Sinus typically between semicircular and U-shaped. Spine in usual posi- tion, stouter than the bristles, acute. Prosternum 1.45-1.5 times wider than long. Distance between chitinous spots 2-++ times wider than width at level of bottom of sinus; 3 times the dental line. First dorsal plate 1.72 times wider than long. Posterior angles of eleventh and thirteenth dorsal plates weakly produced but the pro- cesses varying considerably in distinctness and size. Coxal pores small; mostly 4, 4,4,4; also 4,5, 5,4 and 5, 5, 5, 4, etc. Spines of first legs, P74 to -oy'34; of the second, f-oye4; of the ies pogo; Of the fourth i twelfth, ¢-?335; of the Wide irae, 0, 0, 3, 2 or ROBE RAL peo; of the penult, j{3°3°9, claws three, of the anal, ¢-q-3°3-5 OF ventr al 0, 1, 3, 2,0, claws two, the accessory being nearly as long as the principal one. None of the coxae at all armed. Claw of female gonopods of usual type; long, acutely pointed, moderately curved, mesally hollowed out and rather thin. Spines 2 + 2, long and stout, acutely and evenly acuminate from very base; the inner spine shorter than the outer and also attached farther proximad so that the actual difference is exaggerated in appearance. First article with the mesal side strongly chitinized as usual, proxi- mally bent ectad or appearing excavated. Length 12-15 mm. Typre.— M. C. Z., No. 606, Texas: Austin, J. H. Comstock. The types are all females. Guameius (GuAMBIUS) PINGUIS (Bollman). Lithobius pinguis Bollman, Ent. Amer., 1888, 4, p. 7; Bull. 46, U.S. N. M., 1893, p. 79. The original description, rearranged, is as follows: — Dark chestnut-brown. Head and antennae dark. Legs paler. Slender; not smooth; sparsely pilose. Head wider than long (3.5:3). . [The ratio given, possibly includ- ing prehensors, is doubtless too great]. Polished, not pilose. Antennae short. Articles twenty-two to twenty-four, short. 242 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli four to six arranged in two or three series. Prosternal teeth 2 + 2. Posterior angles of all the dorsal plates straight. Coxal pores 3, 3, 3, 2 to 4, 4, 4, 4, round. . Spines [ventral] of the first pair of legs, 0,0, 1; of the penultimate pair 1, 3, 2, 1 fo 1,3, 3, 1; of the anal pair, 1, 3, 2, 0. Posterior legs short. 7 Claw of female genitalia [gonopods] entire; stout and much curved. Spines strong, subequal. Length of body 9-10 mm. Type Locauiry.— Arkansas: Little Rock (Bollman). Based upon three female specimens. A much rubbed specimen (M. C. Z., No. 602), referable with some doubt to this species, was secured by the writer at Hudsonville, Miss. GUAMBIUS (SIBIBIUS) COLORADANUS (Chamberlin). Plate 4, fig. 3-5; Plate 5, fig. 1. Arenobius coloradanus Chamberlin, Ann. Ent. Soc. Amer., 1912, 5, p. 141. DraGcnosis.— Testaceous to dark brown and dull chestnut; head of same color. Antennae short, articles thirty to thirty-five. Ocelli nine to twelve in three series. Anal legs of male rather slender but tibia more crassate. Penult legs with the usual crest (Plate 4, fig. 5). Dorsal spines of anal legs 1, 0, 3, 2, 1; claws two. Dorsal spines of penult legs 1(0), 0, 3, 2, 2 (2) or 1(0), 0, 3, 2, 1 (o7). Ventral spines of thirteenth legs 0, 0,3, 3,2. Ventral spines of twelfth legs 0, 0, 3, 3, 2; of eleventh, 0, 0, 2, 3, 2. Dorsal spines of first legs 0, 0, 2, 1, 1; ven- tral, 0, 0, 1, 2, 1 or 0, 0, 1, 3, 1. Last pairs of coxae laterally armed. Length 12 to 15 mm. DescripTION.— Dorsum from testaceous to dark brown and dull chestnut with the posterior plates and the first one usually not at all darkened. Head mostly the same color as dorsum or very nearly so, or somewhat more reddish. Antennae concolorous with head, pale distad. Venter from yellow or testaceous to dark brown, the caudal plates usually a little darker. Prosternum and prehensorial feet of same color as head or nearly so. Legs of same color as adjacent portion of venter. Head widest at level of marginal interruptions; clearly wider than long (11:10); caudal margin straight. Depressed or furrowed CHAMBERLIN: THE GOSIBIIDAE. 243 parallel with frontal suture a little cephalad of the latter, from this furrow a median longitudinal furrow extending toward anterior margin. Two, usually clearly impressed, longitudinal furrows on caudal region of head, one a little each side of the middle and the two united in front of the posterior margin by a transverse furrow. Smooth and shining. Antennae short or very short, reaching the sixth or seventh seg- ment; articles thirty to thirty-five, the second very long with those more distad much shorter, cylindric, subuniform or, more usually, with longer articles occurring singly at intervals among the shorter ones, in general decreasing in size distad. Hairs of medium length. Eyes composed mostly of from nine to twelve ocelli arranged in three series: e. g.. 1+ 3, 3, 2,1+4, 3, 3,1+4, 4,3. The single ocellus much largest, subvertically elliptic or oval. Dorsal plates all very finely or obscurely roughened, appearing smooth and shining to the naked eye; usually no furrows distinctly developed excepting the usual depression or furrow immediately within the caudal and lateral margins and on some the short transverse mark on each side near margin at about one third length of plate from caudal margin. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced; posterior angles of other minor plates rounded or obliquely excised. Ventral plates all punctate and finely roughened; impressed with three longitudinal furrows of which the median is most distinct, the latter on some of the more posterior plates ending caudad in a deeper pit or depression about one third the length of plate from caudal edge; mostly with a rather wide transverse depression or furrow caudad of middle of plate. Sometimes an additional longitudinal furrow show- ing on each side between the median and the lateral, the caudal end often curving mesad to unite with its mate at middle; this furrow often more or less united with the lateral one. The last several plates sometimes with furrows very obscure or practically absent. Prosternum about 1.6 times wider than long. Teeth 2 + 2, small acute. Median incision wide and moderately deep, its sides concave. Spine on each side well ectad of outer tooth, much more slender than teeth but stouter at base than neighboring hairs, bristle-like distad. Coxal pores circular, well separated; in number arranged as fol- lows: — 3, 4, 4, 3; 3, 4, 4, 4; 4, 5, 5, 5. Coxae of anal legs armed laterally and corsally other coxae seem- ingly unarmed. Spines of first legs, (0:21 OT) 0: ois; of the second 244 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. and san oes; of the fourth, :}:5+4 or, occasionally, Coss; of the fifth, joo oy of the sixth to the eleventh, (20:33:33; of the twelfth 0,0, 3, 2 2, . and thirteenth, j-j-3:3:3; of the penult, (13:33, in the female, or, O1-3-3-2 in the adult male, each having claw armed with two accessory ns Pare | claws; of anal, 9-7°3°3:4, the dorsal spine of tibia and also outer one of femur readily lost so that spining may appear thus, *)2,°;°32°, two accessory claws present as in the penult pair. In the female the anal legs are short and very slender, decreasing gradually in diameter from the femur distad, the tibia being intermediate in diameter between femur and first tarsal article as the latter is between tibia and second tarsal article; tibia rather weakly longitudinally furrowed along dorsal surface. Penult legs similar to anal except for smaller size. In the male the anal legs are also rather slender but the tibia is crassate, being fully as thick as the femur and being widest at distal end and abruptly thicker than the first tarsal joint, its dorsal surface shallowly depressed, a longitudinal dorsal furrow also present on femur and prefemur, that of the latter less distinct. Penult legs of male also slender, with joints dorsally longitudinally furrowed; the tibia obliquely excised at dorsocaudal corner of distal end and bearing at this place a small, flattened lobe or crest which is trans- verse to the axis of the joint, this lobe bearing a few short hairs but nothing like the brush in Arenobius manegitus. Gonopods of male relatively wide, flattened, truncate distad; bear- ing mostly 4-6 bristles in a transverse row along distal edge of ven- tral side. Gonopods of female with claw entire, long and stout, acutely pointed, considerably curved, darkened distad. Basal spines 2 + 2, subequal or with the inner in some individuals considerably shorter, stout; in ventral view acutely conical in outline. Articles, especially the second and third, glabrous or nearly so on ventral and mesal sur- faces, but clothed with a moderate number of bristles on ectal and dorsal surfaces. Body rather slender being about 7.5 times as long as width of tenth plate; conspicuously attenuated cephalad from eighth plate, with the first plate much narrower than the third. Width of head, first, third, eighth, and tenth dorsal plates to each other as 40, 33, 37, 47 and 47, the eighth and tenth plates being equal and manifestly considerably wider than the head. Length from 12 to 15 mm. A male 13 mm. has antennae 5 mm. CHAMBERLIN: THE GOSIBIIDAE. 245 long and anal legs exclusive of coxa, circa 4.8 or 5 mm. long, with the tenth plate about 1.7 mm. wide. Types.— M. C. Z., No. 599, Colorado: Manitou, August, 1910. Also taken at Durango, Colorado, C. F. Baker, 1894, (M. C. Z., No. 600). GuamBiIus (SIBIBIUS) OEDIPES (Bollman). Plate 5,. fig. 2; 3. Lithobius oedipes Bollman, Ent. Amer., 1888, 4, p. 8. DiaGcnosis.— Brown to somewhat chestnut, the posterior plates sometimes with a median longitudinal dark stripe. Head darker than dorsum, chestnut. Antennae short; articles twenty-four to twenty-six. _Ocelli nine to eleven in three series. Body but little more than six times longer than width of tenth plate. In male fourth article of anal legs very strongly crassate, with a conspicuous bunch of long hairs at distal end (Plate 5, fig. 2). Penult legs much more slender; fifth article with excision and crest (Plate 5, fig. 3). Dorsal spine of anal legs 0, 0, 3, 2, 1, claws three. Dorsal spines of penult legs 0, 0, 3, 2, 1 (o*) or 0, 0, 3, 2,2 (2). Ventral spines of thirteenth legs 0,1,2,3,2. Ventral spines of twelfth legs 0, 1, 2, 3, 2 or 0, 0, 2, 3, 2; of the eleventh 0, 0, 2, 3, 2. Dorsal spines of first legs 0, 0, 2, 1, 1; ventral, 0, 0, 2, 1, 1. None of the coxae armed. Length 12.5 to 15.5 mm. Descrietion.— Dorsum brown to somewhat chestnut, the latter color especially affecting the first and the last few plates; the posterior plates may have a median longitudinal dark mark which may be either broad or narrow or with one or more longitudinal dark lines each side of the middle. Head darker than the dorsum, chestnut. Antennae chestnut, paler distad. Prosternum deep brown, the prehensors much paler. Venter testaceous to rather dark brown, the posteriorly darker and sometimes decidedly reddish from the middle of the body caudad, the darker area usually showing a pale median area. Legs brownish yellow to light brown, the posterior pairs darker proximally, but distally, especially distad of fourth article, light, bright yellow or orange. Body proportionately ‘broad, being only about 6.1 times longer than the width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 47: 43: 50: 61: 64: 58. 246 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Head considerably wider than long (47:42). Of usual shape. Surface punctate, only obscurely uneven. Antennae short, composed of twenty-four (2) or twenty-six (c7) articles which are short. Ocelli nine to eleven, nearly uniformly in three longitudinal series or partially am tour: ie-j=23,105. a3, oy 2s oe aeons ek ale 3,3, 1. The single ocellus ovate. The first ocellus of top row usually conspicuously larger than other seriate ocelli. Prosternum wider than long in about ratio 45:26, being thus about 1.7-+ times wider than long. Distance between chitinous spots 2.26 times as great as width at level of mesal incision, 3.4 times as great as length of the dental line. Teeth 2 + 2, equal, moderately large, acute, and rather pale. Tubercle of spines well developed, situated immediately ectad of outer tooth; the spine rather stout but much more slender than the teeth. All dorsal plates excepting the first finely roughened, the first being smooth and shining like the head. Posterior angles of ninth, eleventh and thirteenth acutely produced. Median ventral furrow extending across entire plate in each case, broad and rather deep. Lateral ventral furrows less distinct. All ordinarily much weaker on posterior plates. Coxal pores circular, moderate: e. g., 5, 5, 5, 4—6, 5; 5,4; 6(5) 5, 5y D2 oy Dy ls oe lO ona) OFOR 2h el 12 Dye q © Se et isl legs, joa21; of the second, 00,2.3,23 of the third and - ONOMSst2r eo 0; 03:32, 2 fourth, o: a 0 335; of the fifth to eleventh, 9-9-5-3-5; a the twelfth, 9473335 0, 0, a, Ons Bie, Or, eas; of the thirteenth, (2343; of the fourteenth or penult, 0, 05.3, 2. 1 Do Bik, 3 ae . QuingsesaOe Oly g= 3 (9 ), the claws three, both accessory claws very Slips ON0N3525 102 distinct; of the al ‘0:1-3-3,1-> claws three, the outer accessory very small and slender. No coxal spines could be detected in the types. In the male the third article of the anal legs is clavately thickened and ridged along mesal or dorsomesal surface. Fourth article very strongly crassate; marked on dorsomesal surface with a broad deep longitudinal furrow which bends somewhat obliquely more ventrad from proximal to distal end; bordered along entire dorsal or ectal side by a very conspicuously elevated thin ridge, which near the beginning of its distal fourth, bears a brush of long hairs; this brush projects mesad; scattered similar hairs also occur on the edge more proximad; the furrow is also bordered on the mesal and more ventral side with a lower thin ridge which bears sparsely along its edge long hairs. Fifth article or tibia abruptly much more slender than the fourth (Plate 5, fig. 2). CHAMBERLIN: THE GOSIBIIDAE. . 247 Penult legs in the male with third and fourth articles much more slender than in the anal, the fifth not greatly more slender than the fourth. Fourth article but little enlarged, dorsally weakly longi- tudinally furrowed. Fifth article or tibia obliquely excised at distal end on mesal side and bearing toward dorsal edge of excised surface a thin keel-like triangular lobe or crest which is long in comparison with height, the highest end being distad, and projects mesad or dorsomesad. Crest not pilose. Gonopods of male of usual shape, flattened and parallel-sided, bristles long. Proximal articles of female gonopods strongly chitinous and con- spicuously excavated mesally at base. Claw long and_ strongly curved, acutely pointed, entire. Spines 2+ 2, large and broad, flattened distad, the outer of about equal breadth to beginning of distal fourth where it narrows to a rounded point, the outer side of tip longer and more oblique than the inner. Inner spines set nearly at right angles to the outer, a little smaller. In a male 15 mm. long the antennae are 5.5 mm. long and the anal legs 6 mm. The tenth plate is 2.34 mm. wide. A second male is 12.5 mm. long; and a female 15.5 mm. long. Locauiry.— Arkansas: Little Rock. The description above is based upon the type specimens. Bollman gives the length of the male as 15.4 mm. and that of the female as 20 mm., but this is an error. ARENOBIUS Chamberlin. Can. ent., 1912, 43, :p. 177. Head with marginal breaks present but small or incomplete. Antennae short; articles normally twenty. Eyes consisting of seriate ocelli; single ocellus enlarged. Prosternal teeth 2 + 2, small. Spines very stout and_ tooth-like. No true median sinus present. ; Posterior angles of ca eleventh, and thirteenth dorsal plates, produced. Pores on only last ae pairs of coxae; uniseriate. Anal legs of male crassate; fourth and fifth articles dorsally, ectally and mesally longitudinally furrowed, the dorsal furrows of fifth article most conspicuous and in type limited by marginal pilosé 248 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ridges. Penult legs more strongly crassate; mesal furrow of tibia deep, conspicuously deepened and widened distally and in type embracing a lobe bearing a conspicuous brush of long hairs. Dorsal spines of anal legs in both male and female normally 1, 0, 3, 2, 1, occa- sionally only 1, 0, 3, 2, 0; ventral 0, 1, 3, 3, 1; claws two. Dorsal spines of penult legs in female 1, 0, 3, 2, 2, in male 1, 0, 3, 2, 1; ven- tral 0, 1, 3, 3, 2; claws three. Dorsal spines of eleventh to thirteenth legs 0, 0, 3, 2, 2. (Some of coxae armed laterally as well as dorsally). Gonopods of male uniarticulate. Gonopods of female with first article mesally strongly chitinized and excavated at base which is constricted, in the way usual in the family. Claw large. Spines 2 + 2. TypPE.— A. manegitus (Chamberlin). The only species of this genus as here restricted known to occur north of Mexico is the type form. The species is peculiar especially in the character of the prosternum and the nature of the modification of the penult legs in the male. ARENOBIUS MANEGITUS (Chamberlin). Plate 6, fig. 3-6. Lithobius manegitus Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 43, pl. 4, fig. 4. Arenobius manegitus Chamberlin, Can. ent., 1912, 48, p. 178. Dracnosis.— Dorsum testaceous to chestnut; head and usually first plate conspicuously darker, dark brown to deep mahogany. Antennae short; articles twenty to twenty-three. Ocelli ten to fifteen in three or four series. Body 6.65 to 6.75 times longer than width of tenth plate. Dorsal spines of anal legs 1, 0, 3, 2, 1 (2 o%); claws two. Dorsal spines penult legs 1, 0, 3, 2, 2 (2) or 1, 0, 3, 2, 1 (). Ventral spines of thirteenth legs 0, 1, 3, 3, 2; of twelfth 0, 0, 3, 3, 2 or 0, 0, 2, 3, 2; of eleventh 0, 0, 2, 3, 2. Dorsal spines of first legs 0,0; 1:1, 1.or0, 0, 2; 1,1; ventral, 0, 0, 2; 1, 1,07 0,0; 1) 25 1; Last two pairs of coxae armed laterally. Anal legs of male crassate; fourth and fifth articles furrowed ectally, mesally and dorsally; ridges branching dorsal furrow of fifth densely pilose (Plate 6, fig. 5). Penult legs more strongly pilose; tibia modified (Plate 6, fig. 6). Length 15-17 mm. CHAMBERLIN: THE GOSIBIIDAE. Fig. 4. Distribution of Arenobius. 249 250 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Description.— Dorsum testaceous to chestnut, nearly uniform, excepting for the first plate which is of same color as or but little paler than the head. Head conspicuously darker, varying from dark brown to deep mahogany, sometimes paler along the caudal border. Antennae of same color as head, becoming paler and more or less rufous distad. Venter from testaceous to rather dark brown, the caudal segments darker, often dull chestnut. Legs nearly of same color as adjacent portion of venter, the posterior pairs thus varying from dark brown to chestnut but always paler distally. Prosternum of almost same color as head, the prehensors somewhat paler and commonly of a more reddish tinge distad. Body proportionately robust being only 6.66 to 6.75 times longer than width of tenth plate. Strongly narrowed cephalad from eighth segment to the first. Eighth and tenth plates equal in width and the first narrower than the third and equal in width to the head. Widths of head and of first, third, eighth, and tenth plates to each other as 95: 53: 55: 66: 66. Marginal lateral breaks of head weak and but partially extending through margining. Sides of head caudad of breaks moderately convex and converging considerably to the rounded caudal corners; caudal margin a little incurved mesally. Head very nearly exactly equal in length and breadth; widest at level of breaks. Smooth or obscurely uneven, finely punctate, shining. Hairs sparse, moderately long. The most conspicuous mark is.a sharply impressed median longitudinal suleus running from anterior margin caudad to end abruptly a little in front of suture where it meets a weaker transverse furrow extending part way to each lateral margin. Antennae short, reaching fifth to seventh segment. Composed normally of twenty articles of moderate length which decrease regu- larly and very gradually in size from the second distad to the penult. Articles sometimes as many as twenty-three. Ocelli ten to fifteen in number, arranged in three or more frequently four series: ¢. g., 1 +- 3, 4,3; 1+ 4, 3,2; 1+ 4,4,3; 1+ 4,3, 2, 1; V4 4 4 Ta, Bo ee og ei a oo Single ocellus large, somewhat cuneate with dorsal margin convex and apex ventrad, often indistinctly limited. Most caudal ocellus of the top series larger than the others sometimes more dorsal in position and often elongate as may also be some of the others. The first ocellus of second row also large. All ocelli deeply pigmented. Organ of Toémésvary cephalad of anterior end of second row, smaller in outline than the small ocelli. CHAMBERLIN: THE GOSIBIIDAE. 251 Prosternum 1.7-+ times wider than long. ‘Teeth 2 + 2, very small, rounded, attached a little caudad of edge of prosternum on dorsal side. Spine a little mesad of each ectal angle and attached a little caudad of edge on ventral side; very stout, acutely conical, greatly exceeding in length and breadth the teeth. Anterior margin on each side straight, the two sides meeting in a very obtuse reentrant angle. Chitinous lines very distinct anteriorly but obscure below. (Plate 6, fr 3). Surface of first dorsal plate similar to that of head; smooth or but obscurely uneven and shining. Major plates depressed within lateral and caudal marginal rims, the depression on caudal plates more conspicuous and more furrow-like. Plates smooth and polished, no distinct furrows usually present. On the ventral plates the most distinct mark is a median longi- tudinal sulcus which is longest and most distinct in the median and caudal regions, on the more caudal plates ending abruptly a little in front of the caudal margin. Lateral longitudinal furrows obscure, more often evident in anterior region of body, the plates mostly not distinctly depressed within the lateral borders. Obscurely roughened. Coxal pores small, circular: mostly 5, 5, 5, 5, also 5, 4, 4, 4 and 5, 5, 5,4. In one specimen 5, 5, 4, 3 on right side while 4, 5, 5, 5 on the left. Another with 5, 5, 5, 4 on left side and 5, 5, 5, 3 on right. 0,0,.2. 11 GBs oak oe: ONO Mal Spines of first legs mostly, ¢-o-3-34, but also, pogcsa and, pors4 ; of the wa ooo (observed in @) or, issy; of the third, the same or, 2a; of the fourth and fifth, po 335 or, 0 oes; of the sixth to eighth, (:9°5°3-5) o-0-3-a-m OT, oy ego; of the ninth, (233 or ventral epines, p; 0, 2; 3,2; of tenth = eleventh, 1a: fee 22, of the twelfth, ain RP ra 2 (in 2 and o”) or, B08 23-3 > (o& chiefly); of the thir- mo 22 2 5 fas 1.0, 35:25 1 TeeMtihy ig 47s 4-3; is the penult, ¢7:3:35 (Q) or, (1333, Claws three; of the ana all = 1.0.3, 74 normally, but occasionally the dorsal spines vary imcels to 1, 0, 3, 2,0 or even 1, 0, 3, 1, 0, claws two, the accessory distinct. Last two pairs of coxae laterally armed. In the males the anal legs are conspicuously thickened; the fourth and fifth joints being most strongly crassate, these distinctly furrowed longitudinally along dorsal, mesal, and ectal surfaces; the dorsal furrow of fifth joint especially deep and conspicuous, not entirely reaching distal end, distinct ridges bounding the furrow, these densely pilose, the hairs being longest where the ridges unite at distal end and there often forming a conspicuous bunch (Plate 6, fig. 5). The 252 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. fourth joint or prefemur typically with two longitudinal dorsal fur- rows, one toward mesal surface and one toward the ectal, the dorsal surface swollen and elevated between them. Penult legs of male more strongly swollen than the anal, the femur and tibia being espe- cially strongly crassate, the tarsal joints abruptly more slender. A longitudinal furrow on ectal side of femur and tibia. Mesal surface of these articles also longitudinally furrowed, the furrow of the tibia conspicuously deepened at its distal end were it embraces a peculiar process bearing a brush of long hairs which projects mesad (Plate 6, fig. 6). Two dorsal longitudinal furrows on the femur, one toward mesal and one toward ectal surface with dorsal surface swollen and elevated between them, the prefemur similarly but less conspicuously modified. Gonopods of male short but very broad with distal edge long and straight and well chitinized. Claw of female gonopods long and very strongly curved, gradually narrowed to an acute point. Basal spines 2 + 2, subequal or the outer a little longer; robust, cylindric proximally, the distal portion subconic in ventral outline, the apex frequently notched (Plate 6, fig. 4). Length 15-17 mm. A male 15.5 mm. long has antennae 7 mm. long and anal leg nearly 5.8 mm. long with tenth plate 2.35 mm. wide. Praematurus.— Dorsum testaceous or light brown, with sometimes a paler, yellow, median longitudinal line and also a less distinct line on each side. Head and first dorsal plate darker brown. Antennae like head but pale and somewhat rufous distad. Venter paler, more yellow, darker caudad. Legs similar ventrally, darker dorsally; the posterior pairs darker, light distally as in adults. Antennae as in maturus. Ocelli 1 + 3, 4, 3. Single ocellus and first one of top row may be paler. Prosternum as in adult excepting that hairs are fewer. Coxal pores 4, 4, 4, 4. Spines of first legs, (aa; of the sixth, 93:55; of seventh the 3, 2, 2 0. . . . . same OF, 2-3-3; Others as in maturus (the lesser spining in each “> case, but ventral spines of twelfth 0, 0, 3, 3, 2). Claw of female gonopods shorter and paler than in adult. Bristles of articles fewer. Basal spines 2 + 2 but the inner one of each pair only about half the length of the outer one and much more slender, acute. CHAMBERLIN: THE GOSIBIIDAE. 253 A female 13.5 mm. long has antennae 5 mm. long; anal leg near 4.5 mm. long; and eighth plate 1.75 mm. wide. (Unaka Springs, Tenn.). Immaturus (Late)— Color nearly as in praematurus. Dorsum light brown or testaceous, the head darker brown or light chestnut. Antennae darkest over middle portion of length, palest distad. Venter and legs testaceous or yellowish. Posterior legs not much darkened, often pale throughout. Articles of antennae twenty to twenty-three; those distad of the first four or five sometimes very short but with longer ones intermin- gled at intervals; in other cases of moderate length, decreasing gradually distad. Ocelli 1 + 4, 2, 1, the anterior ocellus of top row, as usual, not directly in line. Also 1 + 4, 3, 3 with first and last of ventral row very small; and 1 +3, 3, 2. Prosternum as in older stages or very nearly so. Coxal pores 3, 3, 3, 3 or 3, 3, 3, 2 (observed on one side only); most distal one of each series usually conspicuously larger than the others. Spines_ of peek legs, touo3; of the third, ooos; of the sixth , 2, 2, : 3.2, and seventh, -j:3-3°33 of the eighth and ninth, ooa os (ventral once noted also as 0, 0, 1, 2, 2); of others as in adult excepting penult of male of which formula is, (-4°3-35 as in the female. In the male the anal legs are a little more enlarged than in the female. Penult, if different, rather more slender than the anal with the char- acteristic furrows present, but as in the female. Characteristic pilose ridges bounding furrow on dorsal surface of fifth joint of anal legs in adult not present nor is the peculiar excavation and setigerous lobe of tibia of penult legs, this joint also at this stage bearing two dorsal spines at distal end. Claw of female gonopods very short and subtriangular, moderately acute distad, pale yellowish throughout. Basal spines 1 + 1, small and acute, pale. Bristles few. A female 11 mm. long has antennae 4.25 mm. long and anal leg 4 mm. long with tenth and eighth plates 1.6 mm. wide. (Unaka Springs, Tenn.). Immaturus (Early).— Pale testaceous. Head brown with a paler band along the suture. Antennae brown, pale distad. Pro- sternum light brown, prehensors testaceous. Venter dilute yellow of a violaceous tinge, darker caudad. Legs all pale with the tarsi brighter; the caudal pairs not dark, scarcely differing from the others. 254 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae very short, already consisting of twenty articles of which those beyond the third are short and decrease to the ultimate which is long. Ocelli 1 + 3, 2,1. The first of top row about the same size as the single ocellus which is vertically elliptic or oval as in older stages. Prosternum nearly same as in preceding stages. Coxal pores 3, 2, 2, 2. ne 0, 0, 0,1, 1 Tai ee Spines of first legs, p-oq4; of the second, cat i? Olathe third, 0-01-31 or dorsal spines 0, 0, 2, 2, 1; of the fourth p 0.222. of the fifth led to seventh either same as fourth or ventral spines 0, 0, 1, 2,2; of the 3, eighth to eleventh, (73:3; of the twelfth, foes; of the thir- 0, 0, 3 1..0,.3, 2, 1 10,321 1,10;.35-1,.0 teenth, (354; of the penult, ¢73:3:4; of the anal, 234 to G23 sy Anal and penult legs of male already distinctly swollen but more or less uniformly so and the anal clearly more so than the penult. The lateral furrows present but none of the characteristic developments of the adult indicated. Length (o7) 9.5 mm.; anal legs 3.5 mm.; antennae, also 3.5 mm.; width of eighth plate near 1.3 mm. (Unaka Springs, Tenn.). Agenitalis I.— Pale testaceous, of violaceous tinge in life. Head more brownish, paler along the suture. Antennae yellow distad. Legs all pale, the caudal pairs mostly lighter than the others and but little darker proximad. The twenty articles of antennae already present. In the portion distad of a few longer proximal articles, very short ones occur at intervals among longer ones. Ultimate article long. Ocelli 1 + 3, 3, the first ocellus of each of the two series large and: equal to the single ocellus or nearly so. Prosternum as in older stages, but spine relatively even stouter, it- and teeth pale. Posterior angles of ninth, eleventh, and thirteenth dorsal plates already produced. Coxal pores 2, 15, 1,1 Anal glands still strongly developed with no trace of degeneration. Spines of first legs, (-a-i44; of the second, jtoa4; of the third, hate cid i; of the fourth and fifth, 23:1; or dorsal spines of the latter may be 0,0, 1, 2,2; of the sixth to ninth, 04°33; of the tenth and eleventh, (0:73:43; of the twelfth, }}j24; of the thirteenth and penult, (or34; of the anal, (jt. Trochanters as yet all gla- brous and unspined. CHAMBERLIN: THE GOSIBIIDAE. 255 Anal and penult legs in the male already much more crassate as well as longer than the preceding pairs but otherwise not showing adult peculiarities. Length close to 7 mm.; antennae 3 mm.; anal leg (exclusive of coxa) near 2 mm.; width of eighth plate .9+ mm. (Unaka Springs, Tenn.). Pullus IV (Larva quarta).— Pale, suffused with violaceous through- out; the dorsum cephalad testaceous, caudad pale yellow. Head light brown with a rather broad paler band along the suture. An- tennae paler, yellowish distad. Venter yellow caudad, darker cephalad. Of the developed legs the posterior pairs are lightest. Antennae composed of twenty articles. Very short articles oc- curring at intervals in groups between longer ones, the first three or four longer than the others as in later stages. Hairs more sparse. Ocelli 1 + 2, 1. Single ocellus largest, the first of the upper row much larger than other seriate ocelli. Prosternum as in older stages excepting for fewer bristles; teeth and spines paler. A single pore on each coxa of twelfth pair of legs. Anal glands conspicuous. The last three pairs of legs appearing as slender buds, closely ap- pressed, in which the portion distad of the coxa is unsegmented. . : 0, 0 : q Spines of first legs either none at all or, ooo; Of the second, a. 4 o 43; of the third, oat iateel i (right) or dorsal 0,0, 0,0, 1 (left); of the fourth to seventh, fU:p44; of the eighth, CooL of the ninth, EEE of the tenth, 0.0.1 right) or dorsal 0, 0, 1, 1, 2 (left); of the eleventh and twelfth, }-o-41- Length near 5.8 mm. (Unaka Springs, Tenn.). Type.— M. C. Z., No. 596, North Carolina: Hot Springs. Paratypes.— North Carolina: Hot Springs, M. C. Z., No. 588; Catawba, M. C. Z., No. 597; Saluda, M. Z. C., No. 594; Linville Falls, M. C. Z., No. 591. Tennessee: Johnson City, M. C. Z., No. 590; Unaka Springs, M. C. Z., No. 593; Altapass, M. CZ No:; 595. LED et pany ‘ ve Nas rt ; ba i me, fre PLATE 1. CHAMBERLIN.— The Gosibiidae. Fig. Fig. Fig. Fig. Fig. Nw — PLATE 1. Gosibius paucidens (Wood). Anterior portion of prosternum. Left gonopod of 2, ventral view. (Claremont, Cal.). Third and fourth and part of fifth article of anal leg of o, dorsal view. (Laurel Canyon, Cal.). Fifth and part of third and fourth articles of right penult leg of 7, mesal view. (Laurel Canyon, Cal.). Gosibius montereus Chamberlin. (Type, Pacific Grove, Cal.). Proximal portion of gonopod of 92, ventral view. BULL. MUS. COMP. ZOOL. GOSIBIIDAE. PLATE 1 CHAMBERLIN.— The Gosibiidae. PLATE 2. Gosibius brevicornis Chamberlin. (Type, Friant, Cal.). Fig. 1. Anterior portion of prosternum. Fig. 2. Fourth and fifth articles of right anal leg of <, dorsal view. Fig. 3. Mesal view of fifth article of left anal leg of &. Fig. 4. Mesal view of fifth article of left penult leg of @. Gosibius arizonensis Chamberlin. (Type, Fort Williams, Arizona). Fig. 5. Proximal portion of left gonopod of 9. BULL. MUS. COMP. ZOOL. GOSIBIIDAE. PLATE 2 Lt A) | ¥ H \ ey ASS say | VAAN SSeS * \ iff ti/ ef lie aR | \I {Ae + La Si NUNNAUE Sc ye eee PLATE 3. CHAMBERLIN.— The Gosibiidae. PLATE 3. Gosibius arizonensis Chamberlin. (Ft. Williams, Arizona). Fig. 1. Aniterior portion of prosternum. Gosibius intermedius Chamberlin. (Santa Barbara, Cal.). Fig. 2. Lateral portions of sixth, seventh, and part of eighth dorsal plates in outline. Fig. 3. Fourteenth dorsal plate in outline. Pseudolithobius megaloporus Stuxberg. (Oroville, Cal.). Fig. 4. Left gonopod of 9, ventral view. Fig. 5. Fifth article and part of two adjoining ones of penult leg of ¢. GOSIBIIDAE. PLATE 3 BULL. MUS. COMP. ZOOL. PLATE 4. CHAMBERLIN.— The Gosibiidae. Fig. Fig. Fig. Fig. Fig. _ Pence PLATE 4. Pseudolithobius megaloporus Stuxberg. Portion of right anal leg of <, dorsal view. Anterior portion of prosternum. Guambius coloradanus Chamberlin. (Durango, Col.). Left gonopod of 9, ventral view. Dorsal view of fifth and part of adjoining articles of anal leg of o. Fifth and part of adjoining articles of penult leg of <, lateral view. GOSIBIIDAE. PLATE 4 BULL. MUS. COMP. ZOOL. ts ~_ al eG : ~— ‘ PEUAEE Bey) el * CHAMBERLIN.— The Gosibiidae. Fig. Fig. Fig. Fig. Fig. Fig. PLATE 5. ‘ Guambius coloradanus Chamberlin. Anterior portion of prosternum. Guambius oedipes (Bollman). (Type, Little Rock, Ark.). Second to fourth and part of fifth articles of right* anal leg of , mesal view. Fourth and fifth and part of adjacent joints of left penult leg of &, mesal view. Guambius mississippiensis Chamberlin. (Type, Byram, Miss.). Left gonopod of @, ventral view. Fifth and part of adjoining articles of left penult leg of co’, mesal view. Guambius curtior Chamberlin. (Type, Gulfport, Miss.). Fifth and part of adjoining articles of left penult leg of <7, mesa! view. BULL. MUS. COMP. ZOOL. GOSIBIIDAE. PLATE 5 ape “ CHAMBERLIN.— The Gosibiidae. Fig. Fig. Fig. Fig. Fig. Fig. SECS ce PLATE 6. Guambius curtior Chamberlin. Outline of adjoining portions of fifth and sixth articles of penult leg of <, dorsal view. Third and fourth and part of fifth joints of anal leg, dorsal view. Arenobius manegitus. (Chamberlin) (North Carolina). Anterior portion of prosternum. Right gonopod of 92, ventral view. Fifth article and part of adjoining ones of anal leg of <”, dorsal view. Ventral view of fifth and part of adjoining articles of right penult leg. PLATE 6 GOSIBIIDAE. BULL. MUS. COMP. ZOOL. al a ad z ay : ne ‘ a ¢ 1 Segoe oe ae Sie, Baaveny ; od a « *at sy 1 4 7 ‘ « > . payee . \ : . . ‘ a r f ‘ i *% om 5 q = i = 1 j i i ' ; ~ = Bulletin of the Museum of Comparative Zodlogy AT HARVARD COLLEGE. Vou, LVI No. 6: FURTHER STUDIES ON NORTH AMERICAN LITHOBIIDAE. By RaupH V. CHAMBERLIN. Wita Twecve Puates. CAMBRIDGE, MASS., U.S. A.: PRINTED OR THe MUSEU M. JUNE, 1922. tg Ui in? ¥ 7 y vo. i No. 6.— Further Studies on North American Lithobiidae. By Raupo V. CHAMBERLIN. THE present paper, written in 1914, consists of revisions of new or recently described genera of Lithobiidae as represented in America, north of Mexico. All of the material upon which this paper is based was collected by the author, unless specifically stated to the contrary. The key will aid in the separation of the genera. Key to the Genera. GEmINGOcelipresentts sown si Maes dare tees eel Typhlobius, gen. nov. aa. Ocelli present. b. Prosternal teeth 5-++5 or more. Posterior angles of none of the dorsal plates produced; articles of antennae more than 25............. Soztbius Chamberlin. bb. Prosternal teeth normally 2+2, or occasionally up to 4+4. c. Fourth joint in anal leg of male with a lobe at proximal end bearing a dense brush of very long hairs. Posterior angles of none of dorsal plates produced; articles of anteniae/ ra pou 24.5e)xa ee os Pampibius, gen. nov. cc. Fourth joint of anal legs of male not thus modified. d. Articles of antennae 26 or more. (Posterior angles of ninth, eleventh, and thirteenth dorsal plates, or of seventh, or of sixth and seventh in addition, produced). e. Line of apices of prosternal teeth more or less procurved, the inner tooth of each pair being the larger; basal spines of gonopods of female slender. Paitobius Chamberlin. ee. Line of apices of prosternal teeth recurved; basal spines of gonopods of female short and very broad. Taiyubius Chamberlin. dd. Articles of antennae normally 20, rarely as many as 24 (but when so none of dorsal plates with posterior angles produced). e. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced; neither anal nor penult legs of male with any special process or lobe; anal legs with two or three claws...Sontbiws Chamberlin. 260 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ee. Posterior angles of none of dorsal plates produced or, rarely, those of eleventh and thirteenth, or of ninth, eleventh, and thirteenth weakly produced, but, when so, claw of anal leg single; with few exceptions, either anal or penult legs in male with special lobes or ridges, f. Fifth joint of anal leg of male always bearing on dorsal surface at distal end a conspicuous crest. Nadabius Chamberlin. ff. Fifth joint of anal legs of male with no such crest. g. Penult legs of male bearing a conspicuous lobe at distal end of fifth article, but anal legs without Special processi epee ee Simobius, gen. nov. gg. Penult legs of male bearing no such lobe, or if so the anal legs also strongly modified; the third and fourth joints of the latter nearly always longitudinally ridged and produced into lobes, rarely not modified. ..... Pokabius Chamberlin. Sozts1us Chamberlin. Ann. Ent. soc. Amer., 1912, 5, p. 152. Head with lateral marginal interruptions distinct. Antennae moderate in length or short; composed of from twenty- five to thirty-five articles. Eye composed of seriate ocelli; a single ocellus usually moderately enlarged, but sometimes subequal to caudal ocellus of upper series. Prosternal teeth 5+5 to 7+7; spines slender, distally bristle-like. Posterior corners of ninth, eleventh, and thirteenth dorsal plates straight or somewhat excised, or those of thirteenth, or of eleventh and thirteenth, weakly produced. Coxal pores circular uniseriate. Gonopods of male uniarticulate. Claw of genital forceps of female large; either entire or with one or two relatively small lateral teeth situated well proximad. Spines 2+2; moderately long and slender, mostly more or less acuminate from base. First article of forceps not excavated proximally. Tarsi of all legs divided. In the anal legs of the male the fourth joint is more or less thickened, especially distally and is dorsally longitudinally furrowed and typically concave or saddle-shaped; at distal end more or less elevated dorsad (Plater tie 13 °6): CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 261 Last one or two pairs of coxae laterally armed. Third joint of all legs caudad of first to sixth pairs with three dorsal spines, the others with two. Fifth joint of all legs between third and eleventh pairs with two dorsal spines. Dorsal spines of anal legs 1, 0,3, 1,0; ventral, 1,3, 3,2; the claw single. Dorsal spines of penult legs, 1, 0, 3, 1, 1; ventral, 0, 1,3, 3,2; claw single. Dorsal spines of twelfth legs, 1003 leor O.0).3,.0,.05 ventral, 00,33, 2. “Dorsal, spines. of first legs mostly 2, 1, 1, oceasionally 1, 1,1 or 2, 2,1; ventral 2, 2, 1 tO, ia, 2: Length 8-14 mm. Tyre.— S. tuobukus (Chamberlin). This interesting genus, at present known from three species, has a rather restricted distribution (Fig. 1). It is very common in North Carolina, Tennessee, and Virginia, especially, it would seem, in the mountainous sections. This region appears to be the center of dis- tribution. It is known from as far north as Philadelphia and as far west as Arkansas. Key to Species of Sozibius. a. Dorsal spines of second legs 0,0, 3, 2,1; ventral-spines of first legs in adults nearly always 0, 0, 2, 3,2; claw of 2 gonopods SUTIN NE ONOIR SR Us a Wien 5 See A eer S. providens (Bollman). aa. Third joint of first three to six pairs of legs with but 2 dorsal spines, the formula of the second pair being 0, 0, 2, 2, 1; ven- tral spines of first legs 0, 0, 2, 2,1; claw of 9 gonopods either bipartite or tripartite. b. Last pair of coxae alone laterally armed; first five or six pairs of legs with third joint bearing but 2 dorsal spines. S. pennsylvanicus, sp. nov. bb. Last two pairs of coxae laterally armed; only the first three or four pairs of legs with but 2 dorsal spines on third joint. S. tuobukus (Chamberlin). Sozisius TuoBUKUS (Chamberlin). Rlatesis fie 6; Plate 2; fig, eZ: Lithobius tuobukus Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 36. Sozibius tuobukus Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 152. Description.— Dorsum light brown or testaceous, often of a more or less orange cast, the posterior plates commonly in some degree 262 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ' Fie. 1.— Distribution of Sozibius. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 263 darker. Head either the same in color as the dorsum or, more typi- eally, a darker brown with the frontal region paler, somewhat orange or ferruginous; the first dorsal plate commonly agreeing in color with the posterior portion of head. Antennae light yellowish brown proxi- mally, yellow distad. Venter pale brown or brownish yellow with the caudal plates darker. Prosternum and prehensors somewhat orange- brown to very dilute chestnut. Legs pale brownish yellow, the caudal pairs bright yellow, especially distad. Body moderately narrowed cephalad to the first dorsal plate, the widths of head and of the first, third, eighth, tenth, and twelfth plates being to each other as 64 : 58 : 60 : 69 : 72 : 67. Head from equal in length and breadth to wider than long in ratio 20 : 19 or less, the head in the & appearing to be wider on the average than in the 9. Widest just caudad of eyes, rather conspicuously narrowed cephalad; caudal margin mesally slightly incurved. Mar- ginal interruptions very slight, sometimes essentially obsolete. Antennae commonly reaching to the seventh segment but some- times considerably shorter and not surpassing the fifth. Articles twenty-five to thirty of intermediate length; the ultimate usually distinctly shorter than the two preceding taken together, distally pointed. Ocelli from nine to eighteen, usually in four series, sometimes in three:¢:9, bo- byd, 2; L-- 4,5, 4; 14+ '3,3,2;1 + 4,5, 4,2; 1.5, 4,4,3; 1+ 5, 4, 3,2; 1+ 2, 5, 5,3; 1+ 6,5, 4, 1. Single ocellus usually only a little, or sometimes not at all, larger than the most caudal ocellus of the dorsal series, the corresponding one of the second series also often large; mostly vertically subelliptic, but sometimes circular and sometimes angular. The seriate ocelli vary, often irregu- larly, in an unusual degree in size, form, and arrangement, with the series frequently very irregular. Prosternum 1.6-1.7, mostly near 13, times wider than long. Dis- tance between the chitinous spots 1.87 + 2 times the width at level of bottom of median incision; 2.5 to 2.7 times the dental line. Teeth 5+ 5 or 6+ 6, rather small, uniform, the line of apices forming a distinctly reentrant angle. Spine slenderly acuminate, bristle-like, and straight or nearly so. Incision distinctly u-shaped or with the bottom sometimes somewhat angular. First dorsal plate with the sides weakly convex, moderately con- verging caudad to the widely rounded caudal corners; caudal margin mesally incurved; plate 1.6-++ times wider than long. Caudal margin of the ninth plate straight all the way across. Eleventh plate with 264 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. the posterior corners showing tendency toward weak production, while the corners of the thirteenth plate are more distinctly, but still only slightly, produced. Coxal pores circular: 3,4,4,3; 3,4,5,4; 4,5, 5,4; 4, 6,(6, 4; 5; 6,105.5 : (1h, a al OOM sel al 0, 2,1, 1 , Spines os he bile 8S, o-o-sa7 OF 00,3. 213 of the second, { 050, 2 a and 2,2 ON22s ® 25.2, 0-0,2,3,1 to 0, 0,.2,8,23 of the third, 9:o:3'3'2 or, occasionally, { 0-02-31; Of 0, 0, 2, 2, 2 Peete 0, 6, 3, 2, 2 the fourth, §:03°3°3 or 00,232; Of the fourth to the alcontt 1033,3; 1), 0,3 ee (0) ONS eel. iL, @, Shik It of the twelfth, —,0,3-3,33 Of the thirteenth, cians O31 op 0-0-373723 0 Ona i the penult, } 0 » OF sometimes, on one side, j e334 Ly the claw single; A me “a aS of the anal, ¢°1 : $9 OF occasionally, ¢ te4'q, or the spine of trochanter rarely missing. Last two pairs of coxae normally armed, rarely only the anal. Anal legs of the male with the third article thickened into a rounded longitudinal ridge which is subdensely clothed with long bristles. Fourth article elevated into a rounded prominence at proximal end, this being clothed like the ridge of the preceding joint; caudad of this elevation the joint is longitudinally excavated or furrowed, with a ridge-like elevation on each side, the mesodorsal one of which increases in height, and in the number of hairs borne, proximad, while the ecto- dorsal one increases in similar manner distad and terminates in an elevation at distal end of joint which is often conspicuous and which bears the dorsal spine, the latter frequently projecting directly caudad (Plate 1, fig. 6). Penult legs more slender but they may be similarly though more weakly modified. Gonopods of male small but distinctly exposed; oblique. Claw of female gonopods large, long, well curved, and very acutely pointed; bipartite, the main lobe bearing the second near the middle of its length on the outer or ventral Bape as a very small denticle, or tridentate, a denticle appearing at nearly the same level on the outer edge (Plate 2, fig. 1). Basal spines 2+ 2; the outer one of each pair only a little or not at all longer than the inner; in ventral view moderately narrowing from base to near middle and then of uniform width to the beginning of the acuminate division which is long and acute; outer edge often denticulate (Plate 2, fig. 2). Length 9 to 13 mm. Early pseudomaturus.— Body yellow or brownish yellow. Head and prosternum darker, orange. Legs yellow, the posterior pairs brightest. Antennae consisting of twenty-six articles which distad of the third are very short and closely united. Ultimate article much longer than the two preceding taken together. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 265 Ocelli 1 + 3,2. Single ocellus and first two of the upper series subequal in size; the two ocelli of the ventral series much smaller. Single ocellus not distinctly separated or otherwise differentiated. Prosternum of same form and structure asin maturus. Teeth5 + 5, on each side decreasing mesad and the most mesal one considerably reduced. Coxal pores very small: 2, 3, 3, 3. Claw of female gonopods proportionately shorter than in the maturus, with the lateral tooth or teeth occasionally only vaguely indicated. Spines 2 2-+ 2, more attenuated distad than in maturus and the inner spine proportionately shorter. Spines of first legs, ROPES of the second and third, (334; of the fourth, ? 0: ae 3 i, OF on 0: 023,35 of the fifth, a8 he 02 23 > 5; of the sixth to eighth, do232; of the remaining pairs as in the maturus. Last two pairs of coxae similarly laterally armed. Length near 7.5 mm. Specimens described, collected at Johnson City, Tenn., in August, 1910, are nearly between usual praematurus and pscudomaturus stages. Immaturus.— Coloration as in preceding form. Articles of antennae twenty-four. (Regenerating antenna on one side has but fifteen). Ocelli 1 + 2,2; 1+ 2,3. First ocellus of upper series larger than the single ocellus. All contiguous. Prosternum with teeth 5 + 5, the most: mesal on each side being very small. Incision, spines, and general structure as in the older stages. Coxal pores'very smalli:/2;'353;'23;)2;3)-3)3! In the gonopods of the female the three articles are present. Third article with claw indicated as a small, pale, acute point. Second article with one bristle, the first with three. Spines 1 + 1, minute, acute, pale. Spines of first legs, #774; of the second, 0,0,1,1,1 OF 9 eI; of the hird, Taeteiel ?: “ : t; of the fourth, to seventh, COs of the eighth to tenth, rahe 2. of the twelfth, ¢-:3:34; of the thirteenth, } Neer Es (right side) or 1022.1 (left side); penult and anal legs missing from specimen described. Last three pairs of coxae dorsally armed, but no lateral spines detected on any. Length near 6.7 mm. (Johnson City, Tenn., August, 1910). Agenitalis I.— Pale, very dilute yellowish, throughout. Antennae very short; articles twenty. Ocelli 1 + 2,1, also in one case 1 + 3, 2 (South Carolina). First ONONO Sa EL t 0, 0 266 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ocellus of upper series largest, the single one being next in size, while the other two are much smaller. Prosternal teeth 4 + 4. Coxal pores very small: 1, 1, 1, 1. Anal glands fully dev eloped and very distinct. Spines of the first legs, (0-0-4; of the second, (+t 4; of the third, Tq OF por; of the fourth to seventh, Mapa a of the ninth, 0.hi the two dorsal spines being on the anterior side; of the tenth, (441, the dorsal spine of fourth joint being caudal and that of fifth joint anterior in position; of the eleventh to thirteenth, 9-9-9443 others wanting. None of the coxae armed dorsally or laterally. Length near 5mm. (Johnson City, Tenn., August, 1910). Tyre Locauiry.— North Carolina: Hot Springs. Also taken in North Carolina: Linnville Falls, Brown’s Summit. South Carolina: Greenville. Tennessee: Johnson City, Unaka Springs. Virginia: Chatham, Lynchburg, Natural Bridge. West Virginia: White Sulphur. Kentucky: Lexington. The specimens recorded by Bollman from Washington, D. C., as L. providens probably belong to the present species. SozZIBIUS PENNSYLVANICUS, sp. nov. Plate 1, fig. 1-5. Derscription.— From light brown to chestnut, the normal color in specimens in full color, the species being darker than the other two known. Head and, usually, the first dorsal plate darker, the head having a dusky or black area or spot caudad of the suture in front of which it is light ferruginous or orange. Antennae brown proximally, yellowish distally. Venter but little paler than the dorsum, darkest caudally as usual. Prosternum and prehensors typically dark brown to dilute chestnut, the prehensors paler distad. Legs brownish yellow or brown, the posterior pairs more brightly colored, commonly bright yellow, especially distally and ventrally. Body strongly narrowed cephalad, the widths of head and of first, eighth, tenth, and twelfth plates being to each other as 56 : 52:56: 64 : 66 : 60. Head equal in length and breadth strongly narrowed cephalad. Marginal breaks very small as in related species. Antennae short, reaching fifth to seventh segments. Articles nearly always twenty-seven or twenty-eight, mostly the latter; short and very short and usually closely compacted. Ultimate article of moderate length. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE, 267 Ocelli nine to fifteen, usually in three but also often in four series: Agel >,4 noel 0.6) 43a 4543; el 33372; 1 4-4, 3,3, 1; 1+.5,4,3,2. Single ocellus somewhat largest or equal to or smaller than first of upper series and scarcely differentiated. Ocelli varying much in relative and absolute size and in arrangement. Organs of Témésvary in outline usually clearly larger than any ocellus (Plate eien 203): Prosternum with median incision small, u-shaped. Teeth mostly 5 + 5, apically rounded, on each side somewhat decreasing mesad. Spine slender, only a little curved, inserted close to the ectal tooth as usual. 1.73-1.76+ times wider than long. Distance between chitin- ous spots 2+ times width at level of bottom of incision; 2.75 times the dental line. First dorsal plate 1.75-+ or more times wider than long; widest cephalad but only weakly narrowed caudad to the rounded corners. Posterior corners of the ninth, eleventh, and thirteenth dorsal plates with caudal side straight or slightly rounded, in some a little obliquely excised on one side, not at all produced or rarely showing a very vague tendency yee production. Coxal pores: 2,3, 3,2; 3,3, 3, 3; BUSH otEZ oe atone oo ae as ’ 25 2, 1,1 0, 0, oe : Spines of first tee ee of the second, nos; Of the third, 0,0, 2,2 22,2 0,0.23% . oo 33; of the fourth to sixth, ° Ss OF Ges, or of sixth some- times 23:3; of the seventh to elev enth, ieee of the twelfth, aE yoh ale ib Oh ehalya! Shas bal 0-0-3373} of the thirteenth, ¢-7°33:3; of the penult, (73:33; of the anal, —— Only the last pair of coxae laterally armed. Fourth joint of the anal legs in the male moderately thickened; the dorsal surface bowed in concavely between the ends and longi- tudinally shallowly furrowed;, elevated at distal end; hairs straight, sparse, much fewer than in providens and tuobukus (Plate 1, fig. 1). Claw of female gonopods large; only moderately curved; subacute; always with a distinct small lobe or tooth on the outer or ventral edge a little distad of middle of length and there may also be a smaller denticle on the dorsal edge farther distad (Plate 1, fig. 4). Spines 2+ 2: outer one of each pair a little longer than the mesal one; in ventral view both spines appear distinctly and uniformly acuminate from base to apex with the sides appearing smooth or the ectal edge of inner spine showing denticulations, but these commonly bent dorsad and not always evident in ventral view (Plate 1, fig. 5). Length 8 to 10 mm., this species averaging considerably less in size than either of the other two species. Praematurus.— (Male.) Yellow throughout. Head caudad of su- 268 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ture darker, dusky yellow; frontal region dilute orange. Prosternum dusky yellow, prehensors distally a brighter yellow. Legs yellow. Antennae composed of twenty articles which are short and closely united. Ocelli 1+ 3, 38; 1+ 3, 2. Single ocellus smaller than the first one of the upper series which is much larger than the other seriate ocelli. Prosternum with incision, spines, and general structure as in maturus stage. Teeth 5+ 5 Coxal pores very small: 2, 3, 3, 2. spines of first legs, D cura; of the second, Ss Tt: fy Oryaaii; of the third, 9, 0. a Tig to § 0 vB $4; of the fourth, (9-5 eal of The fifth to eleventh Paeee Ole HIG twelfth, toss; of the thirteenth, meer of the penult, (13:34; of the anal, eee Anal leg of male with the fourth saint but little modified, the dorsal surface being slightly bowed and a little elevated distad. Length 7 mm. Pullus IT (Larva secunda).— Body and legs essentially without pigment; the head, prosternum, and antennae with a very dilute yellowish tinge. Ocelli 1 + 2,1. First ocellus of upper series largest, the single one next, the other two being much smaller. Prosternum with median incision relatively wide, semicircular. Teeth 3 + 3. Eight pairs of developed legs are present, the slender appressed buds of two additional pairs appearing caudad of these. The sixth, seventh, and eighth pairs of legs have a-division indicated in their tarsi while the tarsi of the more anterior pairs appear entire. Spines of first to seventh pairs of legs, ff-0-4443 of the eighth, 9:9-73:7- Ventral spine of tibia relatively long and stout, much exceeding the others in size. Ventral spine of third joint, when present, small. Dorsal spine of tibia slender and almost bristle-like. Hairs relatively long. Length near 3.5mm. Type.— M. C. Z., No. 94, Pennsylvania: Upsal, October 20, 1912. Many specimens. SOZIBIUS PROVIDENS (Bollman). Plate 2, fig. 3-6. Lithobius providens Bollman, Amer. nat., 1887, 21, p. 81. Proc. U.S. N. M., 1887, 10, p. 258. Bull. 46, U. S. N. M., 1893, p. 19, 24, 79; 82, 85, 92, 100, 110, 133. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 269 Derscriprion.— Dorsum light brown to yellowish. Head darker, light chestnut, with the frontal region paler, often ferruginous. An- tennae brown or brownish yellow proximally, paler distally. Venter brown to yellow of paler cast than dorsum; the posterior plates darker as usual. Prosternum and prehensors light chestnut of nearly same shade as head. Legs like corresponding plates of venter excepting that the caudal pairs may be lighter, sometimes being bright yellow, especially ventrally and distally. Body robust, 7 to 7.5 times longer than width of tenth plate. Body considerably narrowing cephalad to the first dorsal plate, which is decidedly narrower than the third, the widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 65 : 58 : 62: fordo 270: Head precisely equal in length and breadth, or very nearly so. Widest just caudad of eyes, from these narrowing gradually and but little to the rounded caudal corners. Posterior margin mesally incurved. Lateral marginal breaks small but usually distinct. Antennae usually reaching the sixth segment but sometimes longer and attaining the eighth. Slender distally. Articles twenty-eight to thirty-five; mostly short; rather loosely joined; ultimate long and slender, usually a little longer than the two preceding ones taken together. Ocelli nine to fifteen in four series or, less commonly, in three: e. g., Sar anay Li oon ep ks D353) 3, 25°) 93,44, 35 LF 4, 454,:2- 1+ 4, 3, 3, 1, and 1 + 4, 3, 3, 2, the two last seeming to be the com- monest arrangements. Single ocellus commonly largest but some- times equal to or smaller than the first of the most dorsal series. Seriate ocelli varying considerably in relative and absolute size in different specimens and also in form, being sometimes angular or elongate rather than circular. Prosternum with teeth acute, uniform, moderate; decreasing some- what in size from most ectal to most mesal on each side; line of apices on each side a little convex, as a whole forming a reentrant angle; teeth in number 5 + 5to7-+ 7. Median incision narrowly u-shaped, often narrower at mouth than at bottom. Spine straight, slenderly acuminate and bristle-like, inserted near ectal tooth. 1% to 1.7 times wider than long. Distance between chitinous spots 1.85-1.88 times width at level of bottom of mesal incision; 2.75-288-+ times the dental line (Plate 2, fig. 3). First dorsal plate 1.6+ times wider than long. Considerably narrowed caudad, with the sides convex. Ninth plate with the 270 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. caudal margin straight, or one corner may be obliquely excised. Caudal corners of eleventh and thirteenth plates slightly produced, those of the thirteenth the more distinctly so. Coxal pores: 3, 4, 4,3; 4, 4,4, 3; 4,5, 4,3; 4,4,5,4; 4,5, Hae 4, 5, 5, 3; 5, 6, 6, 4; ete. , 2, 2,1 0,0,2,2,1 = : Ses of first hes usually, $-933°35 0,232 Sometimes, 9°93" ei in specimens toward pseudomaturus stage; of the second and third, ees: of ONONSH 252 0,8, 2D }, 0, 3, 2, the fourth to tenth, 99:33:23; of the eleventh, 9:93:33 or Osa; of the twelfth, eS of the thirteenth, aa claws three; of the 4. t 3. 1 of the anal, j= ae 5. Last two pairs of coxae, laterally penult, 973533 2 armed. As in tuobukus, the fourth joint of the anal legs in the male is typi- cally conspicuously thickened, more strongly so distad than proximad, and is elevated dorsad at the distal end; cephalad of this rounded dorsal elevation the joint is longitudinally depressed and furrowed; the bulge or ridge along mesodorsal surface mesad of the furrow clothed subdensely, especially toward proximal end, with very long bristles. Usually the fifth joint is slender but is somewhat clavately and usually irregularly enlarged distad and may in fact project ventrad in a con- spicuous lobe at the distal end. Tibial and tarsal joints longitudinally sulcate along mesal surface. Penult legs slender, with the last three or four joints sulcate longitudinally on caudal (mesal) side. Claw of female gonopods long, not much curved, scarcely excavated on inner side; comparatively narrow and acutely pointed; strictly entire, there being no trace of lateral teeth. Basal spines 2 + 2; long and rather slender; acuminate from base to tip which may be very narrowly rounded; in ventral view the edges are either smooth or one or more denticles may appear on the ectal edge of inner spine, or occasionally on outer one, or the edges sometimes finely undulate; if the spines are viewed from side, how- ever, denticles directed dorsad from the edges, which are bent or partly rolled, may be seen. Extremes of variation in form of spines are shown (Plate 2, fig. 5, 6). Length 11 to 14mm. A male 14 mm. long has antennae 7+ mm. long; anal legs cir. 4.25 mm. long; and tenth plate 2 mm. wide. Immaturus.— Light yellow. Head a little darker, of dilute orange tinge. Prosternum and prehensors like head. Venter yellow. Legs light yellow, the posterior pairs brightest. Antennae composed of twenty-five to twenty-seven articles which are mostly short, sometimes with two shorter ones appearing together at intervals and being, apparently, the result of more recent subdivision CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. Lait i of a preceding single article. Ultimate article long, conically acumi- nate, distally rounded. Ocelli 1 + 2, 3. The first ocellus of upper series much larger than the other seriate ocelli and as large as or, more usually, considerably larger than the single ocellus. Organ of Témdésvary relatively large as in the maturus. Prosternum in general proportions and form as in the maturus. Median incision u-shaped, with sides more widely divergent than in adult. Teeth = ae 4or5+ 5. Coxallipores:: 2,3, 35:25 13,05 5,.0- The three heticles of gonopod of female are present, the claw on the distal one being also represented as a small, acute, chitinous point. Basal spines 1 + 1, these appearing merely as acute chitinous points. Anal glands absent or obscurely traceable as more or less degenerated remnants. ; 0,0,0,1,1 0,0,1, 1,1 Spines of first legs, 9-0-7743 of the second, goii1; of the third 001,21 00,221 0.1.2.2 0,0,.2,217," 00,222 0,0, 1,1,1 OT 0,0,1,1, 1) of the ourth, 9 0,0,1,1,1 Ore 0,0, 4.1 i oo221; Of the 0.0.1, 2,24 0,0, 2 2,2 222 "0.2, 2, fifth, j-p-7-74 to poo4; of the sixth, popy4 OF 9.0, 2.313 of the seventh 0,0,2,2%2 ” 0,0,2,22 0,222 00,222 — 0,0,2,22 and eighth, > 0, 0,154,208 0, 0, 2 2, 2», of the ee 0, 0, ‘1, 1, 2» 0,0, 1,232 or 0, 0,3 23 of the tenth, 9-9-1735 to o-0-a23; of the eleventh, 9-o-7773:5 or 0-0-2373; ORDA ee ONONDA Te 1 B0R41, OO2L1 of the eno. 0,0,3, 2,2 or 0-0-3-3-2; Of the thirteenth, ¢-3:-34 to o-7°3°3733 penult, 944734 Or o-:1:2°0, the coxal spine minute. Either none of the coxae laterally armed or the last pair with a very small lateral spine. Length 6.5-7.5 mm. (Russellville, Tenn., August, 1910). Agenitalis II.— Light yellow throughout. The head, prosternum, and prehensors a little darker, of an orange cast. Antennae in specimens studied with twenty-two or twenty-three articles which are short or very short, the shortest ones occurring in pairs at intervals as usual. Ultimate article relatively long, clearly exceeding the two preceding together. Ocelli 1+ 2, 3; 1+ 2,2; 1+ 2,1. The most caudal eye of dorsal series much larger than any other, with the single ocellus next in size. All ocelli pale; well separated. Incision of prosternum with sides divergent as in preceding stage. Teeth 4 + 4. Coxalipores: (192525 2:202,.2),2-) very small Gonopods of female apparent as short, biarticulate buds. Anal glands absent or degenerate. 0, 0, 0, 0, 0 0, 0, 0, 0, 1 0, 0, 0, 0, 1 Spines of first legs, 9-9-0714 OF 00-071. Eau of tbesserond, Joi; of the thire d, i tit4; of the fourth to sixth, § 10,04, 1,1 OF 0,0,1,1,13 of the seventh, OO Bd. of the eighth and ninth, we of the tenth, ® oes; of the 272 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ORO aer0 : 0, 0, 0 0, 0, 1, 0, eleventh and twelfth, ooaa; of the thirteenth, 4 nie e I OF 00-1 a ae 0, 0, 1, 0, 0 0, 1, 0 of the penult, oo-77; of the anal, ? TOLL 1, the trochanter bearing a bristle distally. None of the coxae armed. Length 5-6 mm. (Russellville, Tenn., August, 1910). Tyre Locauiry.— Indiana: Bloomington. Also taken in Indiana at La Fayette, Richmond, Brookville, Salem, New Providence, Wyandotte. Tennessee: Russellville, Knoxville, Beaver Creek, Mossy Creek. Arkansas: Little Rock. TYPHLOBIUS, gen. nov. Head with lateral marginal interruptions only vaguely indicated by a slight obliquity at the point where usually developed. Antennae short to moderate; composed of twenty to twenty-eight articles. No ocelli present. Prosternal teeth 2+ 2; line of apices recurved. Spines slender and bristle-like. None of the dorsal plates with caudal angles produced. Coxal pores circular; uniseriate. Gonopods of male uniarticulate. Claw of female gonopods long and slender; bipartite (subentire) or tripartite, the lateral tooth small and situated well proximad. Spines 2 + 2; long and slender, attenuated from base distad. Tarsi of anterior legs slender; distinctly divided and the distal segment more slender. Only the last pair of coxae laterally armed. Third joint of first eight to eleven pairs of legs with but two dorsal spines, or the most anterior one to three with but one. Fifth joint of all legs between third and thirteenth with two dorsal spines. Dorsal spines of anal legs 1,0, 3, 1,0; ventral, 0, 1, 3, 2,0 or 0, 1, 3, 2, 1; claw single, long. Dorsal spines of penult legs, 1, 0, 3, 1, 1; ventral, 0, 1, 3, 2, 1; one accessory claw which is very small or obsolete. Dorsal spines of thirteenth legs, 1, 0, 3, 1, 1; le OF Sr 3eZ.0r OOnoron2 Dorsal spines of twelfth legs 0, 0, 3, 1, a wenieal 0, 0, 3, 3, 2. Dorsal spines of firstlecs 0; 0, 1, 1, 1 or0, 0} 23251 ventrall0,05 2525 onOnO2noa lk Length 8 to 11 mm. fiver 1. /kebus\ sp. nove The two species for which this genus is established are small, much alike in general appearance, and characteristically slender, being 9.5 or 10 times longer than the width of the tenth plate. They are light CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 273 Fic. 2.— Distribution of Typhlobius. 274 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. yellow in color with the head pale orange. Their habits are not known; but the absence of eyes and the weak pigmentation would indicate that they are probably more strongly lucifugous than usual. Key to Species of Typhlobius. a. Articles of antennae twenty; trochanter of thirteenth legs with a spine; ventral spines of anal legs 0, 1, 3, 2, 1. . 7. kebus, sp. nov. aa. Articles of antennae twenty-eight; trochanter of thirteenth legs unarmed; ventral spines of anal legs 0, 1, 3, 2, 0. T. coecus (Bollman). TypHuosius coEecus (Bollman). Plate 8, fig. 7; Plate 4, fig. 1, 2. Lithobius coecus Bollman, Ann. N. Y. acad. sci., 1888, 4, p. 111. Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 36. DescripTion.— Dorsum yellow of dilute orange cast. Head more distinctly orange. Antennae light distally, proximally approaching the head in color. Prosternum like head but a shade lighter. Venter a little paler than dorsum, the caudal plates more distinctly orange than the others. Legs yellow, the caudal pairs most densely pig- mented. Body slender, usually ten times longer than width of tenth plate ora little less. Head as wide as, or wider than, the tenth plate and the third plate narrower than the first. Widths of head and of first, third, eighth, and tenth plates in a female measured as 25 : 22:21: 92 94-2 23- im a male as 28.0726 824 220 220 a2. Head cordate, conspicuously narrowed from caudal edge of eyes cephalad. Equal in length and breadth. Antennae of moderate length; strongly attenuated distad. Articles mostly twenty-eight which are short and often submoniliform. Bristles long and subdense. Prosternum 1.6 times wider than long. Distance between chitinous spots 2.25 times width at level of bottom of mesal incision; four times length of the dental line. Median incision v-shaped, its sides from moderately concave to nearly straight. Teeth moderate in size, the mesal one of each pair a little larger than the outer, acute; line of apices a little recurved. Spine inserted at base of outer tooth, small and slender. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. DTS First dorsal plate about 1.65 times wider than long. Caudal margin of ninth, eleventh, and thirteenth plates a little curved for- wards at ends, the corners being rounded. Coxal pores small, the most proximal ones often much reduced: BCS CHESS he pee Spines of first legs, (334 334; of the second, Soest; of the third, oad; of the fourth and fifth 0231; Of the sixth to eighth, ?: ee 2. of the ninth to eleventh, (4 3 ego; of ith 1e twelfth, ae of the thir- teenth, DET of the penult, or3o7; of the anal, e3 > Last pair of coxae laterally armed. Posterior legs of male short. Penult laterally somewhat com- pressed, distinctly longitudinally suleate on the caudal, or mesal, side; fourth joint raised along dorsal surface into a ridge which is elevated at distal end. Claw of female gonopods long and acute, with a single small acute tooth near base on outer or ventral edge. Spines 2+ 2; long, the outer larger than the inner; slender, attenuated from base to tip. Length 8.5 to 11 mm. Type Locatiry.— Tennessee: Beaver Creek. Also taken in North Carolina at Saluda. In Bollman’s description of the type the following differences may be noted: ventral spines of anal legs 0, 1, 3, 1, 0 and of the penult 0, 1, 3, 2, 0, a spine probably having been overlooked on fourth joint of the anal and on fifth of the penult. TYPHLOBIUS KEBUS, sp. nov. Plate 3, fig. 5, 6 DescrieTion.— Dorsum very dilute yellow, the pigment a little denser on the first one and the several most caudal plates. Head light orange-yellow. Antennae yellowish proximally, whitish dis- tally. Prosternum like head but a little paler. Venter similar to dorsum, the caudal plates, as usual, more densely pigmented. Legs with faint yellow tinge. Body very nearly 9.5 times longer than width of tenth plate, being very slender and appearing almost parallel-sided, with the head wider than any dorsal plate and the first plate wider than the third. Widths of head and of first, third, eighth, tenth, and twelfth plates to each othen asta0ii2/ = 26): 26) 327-226: Head nearly equal in length and breadth, being slightly wider. 276 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Widest a little in front of marginal breaks and strongly narrowed cephalad. Hairs long, very sparse. Antennae short, attaining the fifth segment. Distal portion un- usually slender. Articles twenty, moderately short, the distal ones slender and almost strictly cylindric. Ultimate article very long. Organ of Témésvary in usual position; not enlarged. Prosternum with median incision very narrow and relatively deep, its sides straight. Teeth moderately large and well chitinized; inner tooth of each pair distinctly larger than the outer; line of apices recurved; ectal side of outer tooth nearly parallel with longitudinal axis while the inner edge is very oblique, the reverse being true of the inner tooth (Plate 3, fig. 5). Spine small and_bristle-like, straight. Prosternum 1.52 times wider than long. Distance between chitinous spots 2.4 times width at level of bottom of mesal incision; 3.8 times as great as length of dental line. First dorsal plate 1.8 times wider than long; sides moderately convex and diverging cephalad. Minor plates with caudal corners subrectangular or simply rounded. Plates very sparsely clothed with short hairs, and along margins, with some longer hairs. Coxal pores very small: 3, 4,4, 3. Spines of first legs, ia arse of the second, 0: 72 % OF Hoo the third to eleventh, 9:0:3°3:3; of the twelfth, oO33-3; of the tee Ot33-2; Of the penult, 9:73:34, one accessory Sa present, this being 0. POOH en eee 1, 0, 3, 1, 0 very small and the main claw long; of the anal, 9-7°3°3'4, claw long. Last pair of coxae laterally armed. Claw of female gonopods of moderate size, only slightly curved; distally showing two blunt teeth or lobes separated by only a slight reentrant angle and the outer larger than the inner; farther toward base is a small tooth on the ventral or outer edge. Basal spines 2+ 2. Length 9.25 mm. Typr.— M. C. Z., No. 151. California: Santa Barbara. PAMPIBIUS, gen. nov. Head with lateral marginal breaks well marked. Antennae short; in type species composed normally of twenty-four articles. Ocelli present; seriate; single ocellus not largest. Prosternal teeth 2 + 2; spines slender and distally bristle-like. Posterior angles of none of the dorsal plates at all produced. NORTH AMERICAN LITHOBIIDAE. . CHAMBERLIN Fic. 3.— Distribution of Pampibius. 278 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Coxal pores circular; uniseriate. Gonopods of male uniarticulate. Claw of female gonopods tripartite. Basal spines 2 + 2, long and slender. First article not excavated proximally. Tarsi of anterior legs undivided. In the male the fourth joint is conspicuously crassate, and bears a lobe at proximal end from which projects a brush of very long hairs. Last two pairs of coxae laterally armed. Third joint of first ten pairs of legs with but one dorsal spine, that of only last two pairs with three, that of about first seven pairs unarmed ventrally. Fifth joint of first eleven pairs of legs, or of all these excepting first one or two, with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral 0, 1, 3, 2, 0; claw single. Dorsal spines of penult legs 1; 0,3, 1, 1; ventral, 0; 1,3, 2; 1 or 0, 1, 3,250; claw single: “Dorsal spines of twelfth legs, 0,0, 2, 1,1; ventral 0,0, 2, 3,2. Dorsal spines of first Weesi0) 0/2 2hors0nOnie tle) ventral, 0; ONO leks Length of type species 6.5-7 mm. Type.— P. paitius (Chamberlin). Only one species, found in North Carolina and Tennessee, is at present known. ' Pampisius paitius (Chamberlin). Plate 3, fig. 1-4. Lithobius paitius Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 37, pl. 3, fig. 6. Description.— Dorsum pale brown. Head and posterior seg- ments dark orange. Antennae dull yellow. Venter grey or greyish yellow. Legs pale, greyish, excepting the posterior pairs which are bright yellow, with the brush of hairs on the anal legs of male reddish proximally and yellow distally. Body a little narrowed cephalad, with the first plate narrower than the third. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 50 : 45 : 47 : 53 : 52 : 50. Head subcordate; caudal margin mesally distinetly incurved; moderately narrowed and rounded cephalad. Marginal breaks proportionately large and distinct. Wider than long in ratio 45 : 42. Antennae short. Composed normally of twenty-four articles which are short between the second and the ultimate, and decrease distad; often submoniliform; ultimate article rather slender, moder- ately acuminate, clearly longer than the two preceding together. Ocelli six to nine in two, or sometimes in three, series: e.g., 1 + 4, 1; CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 279 1+ 4,2; 1+ 4, 3,1. Single ocellus smaller than any of upper series, or equal to most anterior. Ocelli of lower series very small (Plate 3, fig. 2). Prosternum with median incision very narrow and sides nearly parallelin same. Teeth of each pair widely separated. Spine acutely acuminate, attached caudad of outer edge of outer tooth (Plate 3, fig. 1); 1.7 times wider than long. Distance between chitinous spots 2.25-2.32 times width at level of bottom of mesal incision; near 4.25 times the dental line. First dorsal plate 1.8-++ times wider than long; sides very convex; strongly narrowed caudad, with posterior corners strongly rounded. Caudal corners of ninth and eleventh dorsal plates rounded or shortly excised; those of thirteenth with caudal edge straight. Coxaliporessmalls aes asas la; 3,09 lon 4;07 42) 5,.0,2) 2nonore; Zar atoy 24 Anos cue, cree of first legs, 0, - 6 : or i: i i 1; of the second, Oe. é oat of the Pat or a 00 2 1; of the fourth to sixth, Ta of the seventh, Le of the eighth to tenth, Deis ee of the eleventh, of the twelfth, ooas; of the thirteenth, res; Of the penult, r. . 321 or ¢: r - ah dy claw single; of the anal, 7 OLS or, af al claw single. Last two pairs of coxae laterally armed. In the anal legs of the male the third joint is clavately thickened and the fourth joint, in particular, is rather strongly crassate; the latter joint has at its proximal end on the dorsomesal side a lobe on the posterior surface of which is born a dense brush of very long hairs which extend caudad to or beyond the distal end of the joint (Plate 3 fig. 3). Claw of female gonopods short; parallel-sided; well curved; tri- partite, with the lateral lobes equal and not much shorter than the median. Spines relatively long and slender, the sides a little incurv- ing at middle, apical acuminate portion short with denticle on each side at base; the two of each pair of nearly same length (Plate 3, fig. 4). Length 6.5-7 mm. Type Locauiry.— Tennessee: Unaka Springs. Also taken in North Carolina at Catawba. SO Se a) S =) Lay i) w io —_ Patrostus Chamberlin. Can. ent., 1912, 44, p. 175. Head with distinct marginal lateral interruptions. Antennae short but often reaching seventh or eighth segment; composed of from twenty-six to thirty-four articles. 280 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli from seven to twenty in from two to four series; single ocellus enlarged. Prosternal teeth 2 + 2; line of apices typically more or less pro- curved, the inner tooth of each pair being borne farther cephalad than the outer (Plate 4, fig. 3, 5). Spines distally bristle-like, median incision v-shaped. Of the dorsal plates the posterior angles of ninth, eleventh, and thirteenth; of seventh, ninth, eleventh, and thirteenth; of sixth, seventh, ninth, eleventh and thirteenth, or sometimes of only the eleventh and thirteenth, are produced. Coxal pores circular; uniseriate. Gonopods of male uniarticulate. Claws of female gonopods tripartite or rarely only bipartite; basal spines 2 + 2, slender, acuminate from base. First article not exca- vated at base. Tarsi of all legs divided. Posterior legs of male not specially modified. Third to fifth article of anal legs more or less distinctly longitudinally furrowed above, a little more crassate in male. Last one or two pairs of coxae laterally armed. Third joint of first six to eleven pairs of legs with two dorsal spines, or occasionally the first one to three pairs, and in one species the first eleven pairs, with but one dorsal spine or first one or two with none; others with three. Fifth joint of all legs between the second or third and twelfth or thirteenth pairs with two dorsal spines. Dorsal spines of anal legs dh'07 3,1; Os ventralOM1e3; 2. lor0,143.2,.0) rarely,0-mloioe a,c: claws two or sometimes but one. Dorsal spines of penult legs 1, 0, 3, ie less commonly, 07.3321,,0, 15,052. loi), and even dlsOia.s2eele ventral 0, 1, 3, 3, 1 or 0, 1, 3, 3, 2; claws two or three. Dorsal spines of twelfth legs 1,0; 3,2, 2 or 0;,0, 3,2, 2, to 0;/0;.3,, 1, 2.and 0; 0F 2; 1, 2: Dorsal spines of first legs 0, 0,0, 1, 1 to 0, 0, 2, 2,1; ventral 0, 0, 0, 0, 0 to 0, 0, 1, 2, 1, the usual formula. Length of species 6.5 to 12 mm. Typr.— P. carolinae (Chamberlin). The species of this genus, even in the case of the adults, in most cases show a purplish pigment which, at least in alcoholic specimens, tinges the muscles distinctly and modifies the color of the entire body, but is most evident in the anterior ventral plates, antennae and head, and the caudal pairs of legs. The anal legs are always dark proxi- mally with the tarsi, or the tibiae and tarsi, conspicuously paler and usually yellow. The head and dorsum are smooth and shining, never rugose. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 281 Fic. 4.— Distribution of Paitobius. 282 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. The species occur chiefly in the southeastern states, though one is known from Pennsylvania and another from New York City (Figure 4). The species embraced under the fourth division (aaaa) in the fol- lowing key may be regarded as constituting a separate subgenus, Tunabius (P. zinus Chamberlin, TYPE) differing from Paitobius sens. str., including all the remaining species, in having either none or only ns eleventh and thirteenth of the dorsal plates produced and in having the line of apices of prosternal teeth typically straight or slightly recurved. Key to Species of Paitobius. a. Posterior angles of sixth, seventh, ninth, eleventh, and_thir- teenth dorsal plates produced. Ventral spines of anal legs 1, 3, 2, 1; of penult, 1, 3, 3, 2; dorsal spines of thirteenth legs 1, 0, 3, 2, 2... P. arvenus (Chamberlin). aa. Posterior angles of seventh, ninth, eleventh, and thirteenth dorsal plates produced. b. Ventral spines of anal legs 1,3, 3,1; of the penult 1, 3, 3, 1. P. juventus (Bollman). bb. Ventral spines of anal legs 1, 3, 2,0; of the penult, 1, 3, 2, 1 P. carolinae (Chamberlin). aaa. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. b. Anal leg with the claw single. Ventral spines of anal legs 1,3, 2,1; of the penult, 1, 3, 3, 2; third joint of all legs with two or three dorsal spines. P. naiwatus (Chamberlin). bb. Anal leg with two claws. c. Ventral spines of anal legs 1, 3, 2, 1 and of penult 1, 3, 3, 2. d. Dorsal spines of thirteenth legs 1, 0, 3, 2, 2; dorsal spines of first legs 2, 2,1; third joint of first seven pairs of legs with two dorsal spines... .P. tabrus (Chamberlin). dd. Dorsal spines of thirteenth legs 0, 0, 3, 1, 1 or 1, 0, 3, 1, 1; dorsal spines of first legs 2,1,1; third joint of first eleven or twelve pairs of legs with two dorsal spines. P. adelus, sp. nov. cc. Ventral spines of anal legs 1,3, 2,0; of the penult also ft a2 OR ge Sek siren: Bee eee. a P. simitus (Chamberlin). aaaa. Posterior angles of only eleventh and thirteenth or of none of the dorsal plates produced.......... Tunabius, subgen. nov. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 283 b. Anal leg with the claw single. c. Ventral spines of penult legs 1, 3, 3, 1; third joint of first eleven pairs of legs with 2 dorsal spines. P. watsuitus (Chamberlin). cc. Ventral spines of penult legs 1,3, 2,0; third joint of first eleven pairs of legs with only one dorsal spine. P. atlantae, sp. nov. bb. Anal leg with two claws. (Ocelli ten or more in three series). c. Dorsal spines of penult and of thirteenth legs 1, 0, 2, 1, 1; ventral spines of penult legs 1,3, 2,1; dorsal spines of tweltth, degsetOW0i2, 2a a. P. exceptus, sp. nov. ce. Dorsal spines of penult and of thirteenth legs 1, 0, 3, 1, 1; ventral spines of penult legs, 1, 3, 3,1; dorsal spines of twelfth legs 0;0,3,2,2 or 0,0, 2, 2, 2. P. zinus (Chamberlin). The key does not include P. exiguus (Meinert). (Cf. p. 302). ParroBius ARIENUS (Chamberlin). Lithobius arienus Chamberlin, Ann. Ent. soc: Amer., 1911, 4, p. 47. Descrietion.— Dorsum brown, darkened caudad and sometimes, though less, cephalad. Head from brown and but little darker than the dorsum, with a transverse band along the frontal suture palest, to a darker, dusky or dull purplish, brown. Antennae brown suffused as usual, with purplish, distinctly darkened distad to the ultimate article, which is alone paler. ‘Venter pale testaceous to brown, becom- ing, as usual, darker caudad, and the last several segments much darker, of a somewhat burnt brown tinge; anterior segments showing a weak violaceous or purplish tinge. Prosternum testaceous, not at all, or but little, darker than anterior sternites, or appearing in some dusky from a dull purplish pigment; prehensors paler ectad and distad, excepting the claw. Legs testaceous, with the tarsi more pigmented, or they may be suffused with the purplish pigment; anal legs much darker, deep purplish brown, but with the tarsi lighter. Body robust, being seven times, a little more or less (co), as long as width of the tenth dorsal plate; conspicuously narrowed from the eighth segment cephalad to the first. Eighth and tenth plates of equal width, much wider than the head; first plate clearly narrower than the third. Widths of head and of first, third, eighth, and tenth plates to each other as 41 : 39 : 43 : 46 : 50 : 50. 284 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Head wider than long in about ratio 41 : 38. Subcordate, widest at marginal breaks. Caudal margin a little incurved mesally. A transverse furrow a little in front of and parallel with caudal marginal rim, its ends curving forwards to lateral breaks. A broad furrow on each side parallel with lateral margin as far forward as eye, and also a furrow each side of middle extending cephalad from transverse furrow. Smooth and shining, punctae few and indistinct. Antennae reaching the seventh segment. Articles thirty-two to thirty-four, of moderate length; gradually decreasing in size from the second distad to the ultimate. Ocelli fourteen to twenty in four or five series: ¢. g., 1 + 4, 3, 3, 3, 1; 1+ 4, 4, 3, 3, 2; 14+ .5, 5, 5, 3, 1. Single ocellus much the largest, subvertically oval, with the narrower end ventrad. Seriate ocelli decreasing in size ventrad as usual. Prosternum 1.5+ times wider than long. Teeth proper small and dark, but elevated on the usual obtusely angular protrusions of the anterior margin; line of apices distinctly procurved. Sixth, seventh, ninth, eleventh, and thirteenth dorsal plates with posterior angles strongly produced, the processes of the sixth being more acute than those of the seventh. Major plates depressed along lateral and caudal borders. On elevated portion of each plate in front of caudal depressed border a transverse sulcus continuous at each end with a sublongitudinal sulcus running a little ectad of cepha- lad; and mesad of the latter are two other sulci the inner of which diverges from its mate of the opposite side caudad. Sternites with a median longitudinal furrow most distinct on ante- rior portion of plate and a submarginal furrow as usual on each side. Two transverse furrows, one near middle and one a little cephalad of caudal margin. Coxal pores small and circular: 3, 4,4, 3; 4,5, 5,3; 4, 5,5, 4 0,0,2.1,1 0,.0,.2,2.1 Spines of first legs, uot; of the second to fourth, 9°o-t-3'2; of the ONON 22a 0, 0, 3 21 tN Ds Game (or, in one specimen on one side only, 0-0-1322); of the pee 0. 0, 2 2,2 0,0, 3. 2.2 0.3.22 020,322, f sixth? iqaoses MOLS ood opel he ISeV enth, § COL22 OF o0223; O ° 0, 0, 3, 2,2 "023,22 0,0, 3.2.2 the eighth, 9-o-7-3'3 or °: 023-9; of the ninth and tenth, 9:9°2°3°3; 0, 0, 8, 2,2 1,0,3,2,2 1,0,3,22 . elev enth, 9, 0-232; of the twelft h, (o-33-3 OF 013-333 Of the thirteenth, TO oscese 3, 1, 1 T3373; of the penult, ; T3323) Claws two; of the anal, } 0: + = : 1, claws two. Last two pairs of coxae late rally armed. Male with anal and penult legs, as usual, moderately crassate. Fourth joint of anal legs thickened and dorsally complanate, with a conspicuous median longitudinal dorsal furrow, the third and fifth articles also similarly but more weakly furrowed. Corresponding articles of penult legs similarly but less strongly modified. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 285 Gonopods of male showing a pale, conical, and apparently glabrous process extending caudoectad a little beyond edge of sternite, or sometimes wholly covered by the latter. Length of types 10.5 to 11.5 mm. A specimen 10.5 mm. long has antennae 5 mm. long; anal legs (exclusive of coxae) 4.5 mm. long; tenth plate 1.6 mm. wide. Tyre Locauiry.— North Carolina: Hot Springs, August 6, 1910. Also taken in South Carolina at Taylors, August 3, 1910. The female of this species is as yet unknown. Pairopius JUVENTUS (Bollman). Lithobius juventus Bollman, Proc. U. S. N. M., 1887, 10, p. 263, 1888, 11, p. 342, 1889, 11, p. 410. Chamberlin, Can. ent., 1911, 48, p. 99, 101. Paitobius juventus Chamberlin, Can. ent., 1912, 44, p. 175. Descrietion.— Brown. Head darker. Antennae brown proxi- mally, fulvous at tips. Venter yellowish to brown. Legs like adjoin- ing portion of venter; posterior legs apparently paler distally. Pre- hensors paler than prosternum. Body about 7.25 times longer than width of tenth plate. Width of head and of first, third, eighth, tenth, and twelfth plates to each other as\ 50)247 249 :.59-. 61 : 56. Head longer than wide in about ratio 52 or 53:50. Sides strongly rounded from breaks caudad about corners. Posterior portion of head with two longitudinal sulci diverging cephalad. Antennae short. Articles, in type, thirty-one to thirty-two; short. Ocelli 1 + 53; 1+ 4, 3, 3. Single ocellus moderate in size, the most caudal of uppermost series large. Prosternum 1.6 times wider than long. Distance between chitinous spots 2 times width at level of bottom of median incision; 3-3.25 times the dental line. Line of apices conspicuously procurved as usual. Posterior angles of seventh, ninth, eleventh, and thirteenth dorsal plates strongly produced, the processes of the last three being espe- cially acute. Posterior margin of sixth plate straight. Coxal pores small, circular; in types: 3, 4,4, 3 and 4, 4, 4, 3. Spines of first legs, 0.0, 21.1, of the second and third, 0 34; of the fourth to _ ee 2. of the eleventh and twelfth, j-p-333; of the 1LA thirteenth, 5°. i. of the penult, ¢-4°3:3:1, claws two; of the anal, AiO E1 0} 40, 8.4 . . ae CL337 OF o1-3°37p Claws two. In types, which are in poor condition, no lateral spines are detectable on posterior coxae but they may have been lost. 1, 0,0 LO.skat 23,2, 0 286 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Claw of female gonopods of moderate length, rather wide, tripartite, the lateral lobes small and acute, the outer one being a little more proximal in position, and the median one longest and much broadest, acute. Basal spines slender, long and acutely pointed, subequal. Gonopods of male small and wart-like, almost completely concealed. Length 6.5-9 mm. A female 8 mm. long has antennae 3 mm. long, anal legs 3.5— mm. long; and tenth plate 1.1-+ mm. wide. Tyre Locauiry.— Indiana: Bloomington (Bollman coll.). Also taken in Tennessee at Mossy Creek (C. B. Branner coll.). The description above is based upon two of the four type specimens from Bloomington. PAITOBIUS CAROLINAE (Chamberlin). Plate 4, fig. 5, 6. Lithobius carolinae Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 47. Paitobius carolinae Chamberlin, Can. ent., 1912, 44, p. 175. DeEscription.— Dorsum from pale yellowish brown to dark brown and light chestnut, the posterior and anterior plates, especially the first one, darker and usually tinged with red or purplish red. Head dull purplish red to reddish chestnut, paler in a characteristic band along the frontal suture. Antennae dull purplish red proximally but pale distad. Prosternum dull brown of reddish tinge; prehensors paler ectad, testaceous, or with slight tinge of ferruginous. Venter from dilute brown or brownish grey to darker burnt brown, the more caudal plates darker and having the characteristic reddish tinge, while the anterior plates are often suffused with purplish. Legs nearly like corresponding portions of venter, becoming darker caudad, but the anal and penult pairs yellowish distad. Body conspicuously narrowed from the eighth segment cephalad as usual. Tenth and eighth plates nearly equal in width, clearly wider than the head. 7.25 to 7.5 times longer than width of tenth plate. Widths of head and of first, third, eighth, and tenth plates to each other as 32 : 29 : 30 : 37.5 : 37. Head suborbicular; almost precisely equal in length and breadth. Caudal margin straight. Somewhat depressed transversely in front of caudal marginal thickening and longitudinally a little mesad of each lateral margin and also a little each side of the median line, the depres- sions usually wide and shallow. Head smooth and shining, the few punctae present being very fine. Hairs sparse, in part long. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 287 Antennae short, reaching to the sixth or the beginning of the seventh segment. Composed mostly of from thirty to thirty-five articles which distad of the third are short and very short, the shortest ones often alternating singly or in groups with longer ones. Ocelli from eleven to sixteen in three or, less commonly, in four series: ¢. g., 1 +5, 4,3; 1+ 4, 3,3; 1+ 5, 4,3, 3. Single ocellus moderate in size, a little obliquely elliptic, commonly not much larger than the most caudal one of the uppermost series which, as usual, is elongate longitudinally; other ocelli decreasing in size cephalad and ventrad. Prosternum 1.5 times wider than long or slightly wider. Teeth proper small but borne forwards as usual, with the line of apices dis- tinctly procurved. Spines in usual position, bristle-like distally. Posterior angles of seventh, ninth, eleventh, and thirteenth dorsal plates produced, the processes of the last three being acute, those of the seventh more obtuse. Posterior angles of sixth plate typically rounded or somewhat excised. All major plates furrowed transversely a little in front of caudal margin and longitudinally within each lateral margin. Elevated portion of each plate with a longitudinal furrow a little each side of median line and a second one farther ectad. Otherwise the plates are smooth and shining, not at all rugose. Hairs very short and sparse. Ventral plates longitudinally depressed or furrowed between median line and each lateral margin, the furrows most distinct caudad. Ante- rior plates transversely depressed between bases of legs and, on more caudal plates, a second transverse furrow appearing as usual, in front of the caudal margin. Coxal pores circular, the most proximal on each coxa often very small? io2454 35°45, 5) 4s 4 543: Tarsi of all legs biarticulate, but those of anterior pairs with division often indistinct. Anal and penult legs short, considerably inflated in both sexes, but somewhat more so in the male. In both the femur is widened and is more or less flattened above and longitudinally fur- rowed, the prefemur and tibia likewise distinctly longitudinally fur- rowed. Penult leg similarly furrowed, but less crassate. Thirteenth legs with third, fourth, and fifth joints longitudinally furrowed above toward each side, the two furrows leaving a low ridge-like elevation between them. The other legs similarly furrowed but less and less distinctly so in going cephalad, the legs being more or less distinctly laterally compressed and somewhat furrowed longitudinally along sides of joints. 288 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 4 Pl nes of first legs, 0: 3 i: a4 a ry 1b rarely On 2 Ser ou the second, 00121; of the third to seventh, 9:9-7°3'43 of the eighth, °° 00-121 OF 0-0-1313 of the ninth, $3434 to 0.0.3.2.2, of the tenth and eleventh, caters of the twelfth, ? iki or, rarely, on one side, (4 a 35; of the UME ReL nerer of ae penult, ¢ 1 ae 4, claws two; of the anal, « ET) (or ina 1, 0, 3, 0, 0 partly regenerated leg observed also as 0-1. 3-070)5 lacs a6 Last pair of coxae laterally armed. Claw of female gonopods short and relatively rather broad, thin, and well curved; tripartite, the outermost, or most ventral, lobe smallest and sometimes almost obliterated, the median considerably longest, all acute. Basal spines moderately long and slender, the inner usu- ally somewhat more slender than the outer; both gradually narrowing distad from base to more acuminate apical portion, with edges of latter portion sometimes minutely denticulate. : Length 8.5 to10 mm. A female 9.5 mm. long has antennae 3.8 mm. long; anal leg 3.2 mm. long; and tenth plate 1.38 mm. wide. Praematurus.— Testaceous or pale brown, with anterior and _ pos- terior plates reddish. Head brownish to dull purplish red, with the transverse pale band along the frontal suture as in the adult. An- tennae from light to dark dull purplish red, pale distad. Prosternum and prehensorial feet as in maturus. Venter pale testaceous, the anterior plates suffused with light purple, the caudal plates darkened as usual. Caudal pairs of legs dark proximally and light, yellowish, distad. Antennae as in adult or nearly so, the number of articles complete, short, and very shone Ocelli 1+ 4, 3, 2; 1+ 4, 3. Single ocellus frequently smaller than most caudal one of upper series, which is large. Ocelli of most ventral row small. Prosternum as in adult. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced acutely, those of the seventh somewhat more weakly pro- duced than in the adult. Coxal pores: 3,3,4,3; 3,4,4,3; small, the most proximal especially so. Claw of female gonopods shorter and less curved than in adult, pale and thin; the three lobes distinct but the outermost minute and the median much the longest, all acute. Spines nearly conical, uni- formly narrowing from base to apex, the inner spine on each side only about one fifth shorter than the outer, the two spines in this form appearing to develop more nearly simultaneously than is usual in most of the Lithobiidae. Bristles fewer than in maturus. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 289 1 ONO ala d 0, 0, mee hal 1 tO 0,0, 0.0.05 of the second to fourth, 9-9-7734 ih 1 6 al = , 0, al Spines of first legs, 5 0,0, 0, 0,0 2,2,1 By Ay-Ar? a4 : OF 0.0/9, 2,15 of the fifth, ¢¢to4 “34; of the sixth and sev enth, 0, 2 22 0, Oe ee OR ON 222 : 0, 0, 2, 2, 2 031; Of the eighth, ¢ 0.0.21 OF 0.0,0,3,23 of the ninth, ¢7 0,2.1 OF 0, , 0, 2, 2, 2 0, 0,2, 2; 2 0, 0, 23, 2 0 of the tenth, ? 001,21 Or 9-01-35; Of the elev enth, T0221 OF , Ae hos Teal : of the twelfth, ° oso; of the thirteenth, -74330; of the Uae eal saTe 0; 3..1,.0 eee a ak 39-1; Of the anal, 97°33. Length 7 to 7.5mm. A specimen 7 mm. long has antenne 3.25 mm. long; and anal legs 2.75 mm. long. Tyre Loca.tiry.— North Carolina: Asheville. Found also in North Carolina at Hot Springs. South Carolina: Landrum. ‘Tennessee: Russellville. The statement (Ann. Ent. soc. Amer., 1911, 4, p. 47) that the angles of the sixth plate are produced was due to several specimens of arvenus (from Taylors, S. C.) having been included in the description. Parropius NAtwaTus (Chamberlin). Plate 4, fig. 3, 4. Lithobius naiwatus Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 42. Paitobius naiwatus Chamberlin, Can. ent., 1912, 44, p. 175. Descrietion.— Dorsum from light brown to dark brown and chestnut, the caudal segments darkest, often dark brownish purple. Head from dark brown, in the paler specimens, to deep purplish brown or purplish chestnut in the darker ones; mostly uniform, but the frontal region sometimes a little lighter. Antennae concolorous with head proximally, becoming pale distad, the end portion being from yellow to rufous. Venter yellowish brown to brown, the sternites normally becoming darker from the anterior ones caudad, the last four or five plates being especially dark, often dull reddish brown. Prosternum dark brown, usually suffused with purplish; prehensors paler, light ferruginous or rufous distad. Legs yellow to brown, the last two pairs dark purplish brown to purplish black excepting the tarsi, or tibiae and tarsi, which are light. Body robust, being relatively very wide; conspicuously narrowed cephalad; typically only about six times longer than width of the tenth plate. Widths of head and of first, third, eighth, and tenth plates to each other as 87 : 80 : 77 :95 : 100. Head widest between eyes and marginal breaks as usual. Caudal margin nearly straight. Smooth and shining, not at all roughened; a few minute and scattered punctae. Equal in length and breadth, or slightly wider. 290 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae of moderate length, normally reaching to the eighth segment, or sometimes to the ninth. Articles beyond the second of moderate length and either uniform throughout or those of the distal two thirds may be considerably shorter than the others. Articles from thirty-two to thirty-six. Bristles long, not very dense. Ocelli ten to sixteen in three or four poe 5s gs) Jet A oa: 1+ 4, 4,4; 1+ 4,4,3,1; 1+ 4, 3, 3,2; 1+ 4, 4, 4, 3. Single ocellus much the largest, subvertically elliptic, often paler than the others. Series regular, the ocelli of the most dorsal one often sepa- rated by a small space from each other, especially so those toward the caudal end. Organ of T6mésvary in outline of about same size as an average sized ocellus, a little cephaloventrad of eye-patch. Prosternal teeth 2 + 2 or rarely 1 + 2; those of each pair separated by a rather wide space; margin extended beneath each tooth; line of apices procurved. 1.6 times wider than long. All major dorsal plates with a lateral longitudinal furrow parallel with, and but little removed from, lateral margin, and all but the first with a sublongitudinal furrow each side of the middle running from anterior end obliquely ectocaudad, a third weaker furrow being also more or less evident between this and the submarginal one. Not roughened. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. Ventral plates, excepting the first one, with a distinct median longi- tudinal sulcus impressed only at anterior end on the more cephalic sternites but becoming longer and more distinct on the caudal ones. On the more caudal sternites there is commonly a deep impression in the form of an inverted v which is most acute on the last plates, the mark becoming more obtuse and less and less distinet in going forwards. Coxal pores mostly small, pale edged, strictly circular, decreasing proximad on each coxa as usual: 3, 4,4,3; 4,4,5,4; 4, 5, 5, 4; Dy 55 485. 169/514 sete: Spines of first legs, (0:73:13 of the second, p53" tor 4 third, igs or PAE oe of the fourth and fifth, (P44 of the sixth, 0 0 ae 5; of the sev enth to the tenth, Oo a eleventh, ; er of the twelfth, 5 Os 3°5 OF Grys379; of the thirteenth, ai a5 35; of the penult, 4 13. ny 5, claws two; of the anal, aes ¥ 1, the claw single. Only the 1s pair of coxae laterally armed. Anal and penult legs in male short, moderately thickened, with the tarsi more or less abruptly more slender; fourth joint clavately thick- ened, dorsally a little complanate and longitudinally furrowed above, the corresponding article of penult legs being similarly furrowed but CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 291 not enlarged. These legs are similar in the female but are less thick- ened and have the furrows weaker. Claw of female gonopods rather slender and thin; tripartite, the lobes acute, with the median longest. Spines of each pair equal in size or nearly so; moderately narrowing from base to distal third and then more abruptly running to an acute point. Gonopods of male broad at base; distally broadly rounded to sub- conic. Length from 9.5 to 13 mm. Pseudomaturus.— Coloration similar to that of adult but prevail- ingly somewhat paler throughout. Antennae as in adult but articles relatively somewhat shorter. Ocelli 1 + 443; 1+ 5, 4, 3; larger than in the praematurus. Coxal pores: 3, 4, 4,3 to 4, 5, 5, 4, small. Claw of female gonopods smaller and paler than in adult, especially in specimens toward the praematurus stage, with lateral lobes relatively somewhat smaller. Basal spines as in adult, or the inner one of each pair a little more slender and shorter relatively. Spining of legs in older specimens as in adult while youngest ones 0,0, 2,21 may have formulae as follows :— fourth and fifth pairs, O13 OF 0, 0, 2, 2.2 0, 2, 2, 2 0, 0, 2, 2, 2 0, 2, 2,2 COs, seventh and eighth, p-9-4-3°3 OT o-0ra-2-33 ninth, o-9-5-5-3; tenth 0, 0, ¢ OSONSeibelee 1, 0, 3, 1, 20) and eleventh, ¢:p-3°5:3; twelfth, 9-p-3-3°33 thirteenth, oe penult, 1, 0,3, 1,0 . a 133-2; anal as in aoe aalt: Last pair of coxae laterally armed. Length 7.5 to9 mm. (Saluda, N. C.). Praematurus.— Dorsum brown. Head chestnut suffused with purple. Antennae dark like the head but more distinctly purplish proximally; light distally. Prosternum brown, the prehensors lighter, especially distad. Venter pale greyish yellow to yellowish brown, more brown caudad. Legs pale, the two last pairs darker and suffused with purple proximally, yellowish distally. Antennae composed of thirty to thirty-one articles which are mostly of moderate length as in adult, but shorter ones occur at intervals in pairs. Ocelli ten to thirteen in three series: e. g., 1 + 4, 3,2; 1+ 5, 3,3; 1+.5, 4, 3. Of the seriate ocelli the most caudal one of top series is relatively large. Coxal pores very smail, pale: 3, 4, 4, 3. Claw of female gonopods pale throughout, moderately curved, the three lobes present but the median one much exceeding the lateral ones, which appear as small but distinct teeth. Inner spine of each pair about one half the length of the outer one. OOS) at ORO ° Spines of first legs, 0:0: 10.10). i; of the second, ¢ 0:0,0-1-1; of the third and 292 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. of the fifth, fo:33 0-2. 34; of the sixth to eighth, 0, 0, 0, 2, 2, § fourth, > on m + L ) of the ninth, 1 OF 0,0,1,2.13 of the tenth and eleventh, 0,0, 21,1 of the twelfth, 245253 ‘of the thirteenth, ¢yso'2; of the penult, Bra op OF Goya i of the anal, (3:33, 4 oy OF es ro No lateral spine as yet manifest on last coxae. Length near 7 mm.; antennae 3.5 mm.; anal legs cir. 2.5 mm. (Linville Falls, N. C.). Type Locatiry.— North Carolina: Saluda. Found also in North Carolina at Catawba and Linville Falls. South Carolina: Landrum. Georgia: Tallulah Falls. Pairopius TaBrus (Chamberlin). Plate 5, fig. Lithobius tabius Chamberlin, Ann. Ent. soe. Amer., 1911, 4, p. 44. DescripTion.— Dorsum dark brown, either not at all or but slightly darker caudad. Head and first dorsal plate much darker, chestnut of a dull purplish red tinge. Antennae concolorous with head, pale distad. Venter light brown anteriorly, becoming darker and darker caudad; in some the entire venter is suffused with pur- plish. Legs pale brown, paler ventrally than dorsally, with tarsi yel- low; often suffused with purplish; legs becoming darker caudad, the two last pairs especially deep in color, brown of a dull purplish cast, with the tarsi orange or light ferruginous. Prosternum similar 1 color to the head but of a somewhat paler shade, the prehensors rites dull ferruginous. The characteristic reddish purple tint may be more or less evident throughout the body. Body considerably and uniformly narrowing from the tenth plate cephalad, with the head and the tenth plate of nearly the same width. Body typically near seven times as long as the width of the tenth plate. Widths of head and of first, third, eighth, and tenth, plates to each other as 438 : 36 : 36 : 42 : 43. Head suborbicular, with the sides conspicuously and rather evenly convex and the caudal margin weakly excurved. Equal in length and width or very nearly so, widest between eyes and marginal breaks. Head usually rather broadly and shallowly depressed a little each side of the median line from near caudal margin a variable distance cepha- lad. Not at all punctate or rugose, smooth and shining. Antennae short, reaching toward end of seventh segment or, occa- sionally, only to the fifth. Articles between the second and ultimate CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 293 moderately short, not much varying in proportions; in number mostly from thirty to thirty-four. Ocelli nearly always in four series, occasionally in but three; from fourteen to twenty in number: e.g., 1 + 4, 4, 3,3; 1+ 4, 4, 4, 3; 1+4,4,5,4; 1+4,5,4; 1+ 4,5,3. Single ocellus well separated, not larger than most caudal one of most dorsal series. Ocelli of caudal portion of most dorsal series much larger than the others, longitudi- nally more or less elongate. Ocelli decreasing markedly in size from above ventrad and in each series from caudal end cephalad. Prosternum 1.5 times wider than long. Distance between chitinous spots 2.25 times greater than width at level of bottom of median incision; 3,2 times the dental line. Line of apices procurved in the typical manner. Spine distally bristle-like. First dorsal plate smooth and shining; a transverse furrow a little in front of caudal margin curving cephalad at each side and extending to anterior border. Other major plates marked with a pair of distinct longitudinal furrows on the median portion, with, between them, a more shallow median furrow, all the furrows becoming deeper on the more caudal plates; a transverse furrow a little ways in front of the caudal margin, this laterally bending cephaloectad and entering a submarginal lateral furrow which on the more caudal plates is wide and deep, though only weakly developed on the anterior ones; a furrow running ob- liquely from each anterior corner to meet or nearly meet the longi- tudinal dorsal furrow of the corresponding side. Ventral plates mostly convexly curving from ends to middle. The usual longitudinal furrow mesad of each lateral margin mostly weak; a weak median longitudinal furrow often evident, this most distinct on anterior portion of plate; the usual transverse furrow in front of caudal border more or less distinct, while on some plates a second transverse furrow farther cephalad may be seen. d Coxal pores circular or a little transversely elongate, of moderate size. Porigerous area more or less depressed and marked off by lateral elevated rims: 4, 4, 4,3; 4,4, 4,4; 4, 5, 5,3 eee ; 0, 0, 2, 2,1 12, Spines of first and second legs, 0-0-1-3-13 of the rae’ a0 * eT ray 9 roe 3, the fourth to seventh, ¢¢4°3:3; of the eighth, 9-y:3: 3:3, one ventral OS of third joint very small or occasionally absent, giving formula 9:5; Onaaeea2 PONS R22 of ninth to eleventh, ¢°p:3:3-3; of the twelfth, (-j-3:3°3; of the thirteenth, 1,0,3,.2.2, 1,0, 3,21 1, 0, 3, 1,0 0,1,3,3,23 of the penult, 9-7:3:3:9, claws two; of the anal, ( Gaia OF, 5 ° OMe: in one specimen on one side only, observed as §-y-324) Claws two. Last two pairs of coxae laterally armed. The anal and penult legs are short in both male and female. In the ot ne 2 . 5 I. 25 [Spl 2, 3m 294 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. male the anal legs are moderately crassate, the thickening being most marked in the fourth joint which is typically wide and is deeply, longitudinally furrowed along the dorsal surface, the furrow being deepest and widest at the middle of its length; tibia also with a simi- lar, but shallower, longitudinal furrow as also, but less distinctly, the prefemur; tarsus abruptly more slender than the tibia. Penult legs somewhat less crassate; similarly furrowed. The immediately pre- ceding legs with prefemur and femur complanate above and with a dorsal longitudinal suleus toward each lateral edge, a ridge-like elevation between the two; sulci most marked on femur; more anterior legs similarly but less distinctly marked. Most of the legs somewhat compressed, the joints proximad of tarsus, especially tibia and femur, more or less longitudinally furrowed along each lateral surface. In the female the legs are similar but the posterior pairs are less inflated, with the femur not conspicuously broadened, the tarsi not abruptly more slender, and the sulci less distinet. All tarsi biartic- ulate. Claw of female gonopods moderately slender, thin, considerably curved; bipartite, the outer lobe usually obtusely angular and much longer than the more acute inner one. Basal spines subequal, rather slender, very acutely pointed. The statement in the original descrip- tion that the claw is tripartite is a typographical error. Length 9.5-10.5 mm. a claws two, with dorsal spine of coxa sometimes doubled. Anal and penult legs of male short; but little and uniformly thick- ened. ‘Third, fourth, and fifth articles longitudinally suleate dorsally, the fourth article most distinctly so. Length of type (c") 7.5 mm. Female unknown. Typre.— M. C. Z., No. 129. Alabama: Jackson. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 301 Parrospius ziINus (Chamberlin). Plate 5, fig. 2. Lithobius cantabrigensis zinus Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 41. Derscriprion.— Testaceous to brown, with the posterior segments darker, not of reddish tinge, the anterior plates sometimes tinged with violaceous. Head dark brown, not of reddish tinge. Antennae purplish brown, pale at tips. Prosternum brown of a lighter shade than head, the prehensors paler. Venter dilute testaceous, the caudal plates darker, burnt brown. Legs nearly like corresponding plates of venter; the posterior pairs brown, somewhat paler distally. Body moderately robust; 7.5 (Thomasville) to 8 times longer than width of tenth plate, conspicuously narrowed from eighth segment cephalad to first plate which is much narrower than the head. Widths of head and of first, third, eighth, tenth, and twelfth plates to each Other asi2os 2d 224-228 27 +26: Head well rounded, suborbicular, from about equal in length and breadth to a little longer (51 : 49). Caudal margin mesally straight. Hairs of moderate length, sparse. Antennae reaching sixth or seventh, or, sometimes, the beginning of the eighth segment. Articles twenty-eight to thirty-five which are mostly very short. Ocelli eleven to sixteen in three or four series: e¢.g., 1 + 4, 4, 3; 1+ 3,4, 3; 1+ 4, 4, 4; 14+ 4, 4, 4, 3. Single ocellus moderate, only a little largest. Ocelli of dorsal row larger than others of series as usual. Prosternum 1.47-1.48 times wider than long in types. Distance between chitinous spots 2.56-2.7 times width at mesal incision; 3.1- 3.34 times the dental line. Median incision rather narrowly v-shaped, with the sides nearly straight. Teeth of each pair well separated, the incision semicircular, the sides of teeth toward each other slanting more than the others; inner tooth larger than the outer but not borne farther forwards, so that the line of apices is a little recurved. Sides slanting directly ectocaudad from spine, and straight or nearly so. First dorsal plate 1.78-1.8 times wider than long; sides strongly convex anteriorly, the plate being widest some distance caudad of the anterior end; sides converging strongly caudad. Caudal margin of ninth plate straight. Posterior angles of eleventh and thirteenth dorsal plates weakly produced. Coxal pores: 2, 3, 3, 2; 3, 4, 4, 3. Small, the distal one on each coxa decidedly larger than the others which decrease proximad. 302 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. : 0,0, 21,1 0,221 Spines of first legs, (p24; of the second and third, } 0.0.12 D121; of the 0, 2, 2,2. 0,0; 2; 2,.2 OMRON fourth, (3434; of the fth, q-0as7 Thomasville) or a T1293; of the 2. 0, 0, 2, 2, 2 2 tl sixth and seventh, ° 0-0 nih eighth, 9-9-3-s"5 (Thomasville) or (Thomasville); of the tenth, 9 & 753; of the ninth, ¢ r5 0, 0, 3, 2, 2 ; of the eleventh, 9-9:5°3:3 (Thomas- 2 2 Ly 3, 2, g to 4 Py hy il ’ Ee y 2 Eigen.) or 0,0, 2,2 0, 0, 3, 22, : OF 0-833 of the twelfth, ° TO 22 tO ones OF the thirteenth, : Th (OS 8}, 9, ILOseh kal > (or o7:3°3-3, Thomasville var.); of the penult, 97333, claws two, or with minute anterior one also evident; of the anal, (-¢3-3-) claws two. Last one, or two (Thomasville), pairs of coxae laterally armed. Claw of female gonopods of moderate size; but little curved; acute, with a small acute tooth toward base on ectal side and a similar one on mesal side. Spines long and slender, but slightly or not at all converging from base to the relatively very short apical portion, which occupies only a fourth or a fifth of the total length. Mesal side of first article straight or only slightly incurved. Length 7.5 to 10 mm. A female 7.75 mm. long has antennae 3.75 mm. long; anal leg 3 mm. long; and tenth plate .98 mm. wide. Type Locauiry.— North Carolina: Brown’s Summit. Taken also in Virginia at Chatham, Natural Bridge, Lynchburg, Baleony Falls. Alabama: Anniston, Thomasville (variety). Ten- nessee: Johnson City. Georgia: Tallapoosa. Parrospius Ex1auus (Meinert). Lithobius exiguus, Meinert, Vidensk. meddel. naturhist. foren. Kjoben, 1886, 1884-1886, p. 110. (nee Chamberlin, Ann. Ent. soc. Amer., 1911, 4, p. 40; Can. ent., 1911, 43, p. 104). The original description is as follows: — Brunneus, corpore subtus pedibusque pallidioribus, capite fulvo; sat gracilis, sublaevis, capite suborbiculari, fere aeque longo ac lato. Antennae breviusculae, 31l-articulatae (articulis brevissimis). Oculi ocellis 7, in series 2 digestis. Dentes prosternales bini. Pori coxales 2, 3, 3, 2-3, 4, 4, 3, rotundati. [Laminae dorsales omnes angulis rectis.] Pedes corporis primi paris calearibus 0,0,1; pedes anales calcaribus 1, 2, 1, 0-1, 3, 2, 1, [unguibus binis], armati. [Pedes penultimi paris unguibus binis ental [Pedum analium coxae caleare singulo, parvo armatae]. Pedes postici breviusculi, crassiusculi. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 303 Genitalium femineorum unguis sat manifesto trilobus, aculeis tenuibus, interioribus quam exterioribus paulo tenuioribus. Long. 6.5 mm. (fem.) — 8 mm. (mas.). Locatity.— New York City. (L. Lund). Taryusius Chamberlin. Can. ent., 1912, 44, p. 176. Head with distinct lateral marginal interruptions. Antennae short; articles twenty-eight to thirty-five. Ocelli six to fourteen in three, or occasionally but two, series; single ocellus much largest. Prosternal teeth 2 + 2 (or very rarely 3 + 3 in individual cases) ; line of apices recurved or, at most, straight. Spines long, distally bristle-like. Posterior angles of the ninth, eleventh, and thirteenth dorsal plates produced. Coxal pores circular; uniseriate. Gonopods of male uniarticulate. Claw of female gonopods short; tripartite. First article with mesal side straight, not excavated mesally. Basal spines 2 + 2, or occasionally 3 + 3; conspicuously broad; usually widest at or near beginning of acuminate portion. Tarsi of all legs divided. Posterior legs of male without special modification. Either none or the last one or two pairs of coxae laterally armed. Third joint of all legs, or of all but first pair, with two or three dorsal spines. Fifth joint of all legs between second to fourth and twelfth pairs with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral 0, 1, 3, 2, 1 to 0, 1, 3, 3,0 and 0, 1, 3, 3,1; claws two. Dorsal spines of penult legs 1, 0, 3, 1, 1; ventral 0, 1, 3, 3, 1 or 0, 1, 3, 3, 2; claws 2 or 3. Dorsal spines of twelfth legs 0, 0, 3, 1, 2 or 0, 0, 3, 2, 2; ventral, 0, 0, 2,3, 2 to0,0,3,2,2. Dorsal spines of first legs 0, 0, 2, 2, 1 or rarely fewer; ventral 0, 0, 1, 2, 1 to 0, 0, 1, 3, 2. Length 9 to 15 mm. ‘Type.— T. angelus Chamberlin. This genus appears to be very close to Paitobius, the species of the two genera being much alike in habit, coloration, and general appear- ance. ‘Taiyubius is to be distinguished through its having the line of apices of prosternal teeth recurved and through the broad basal spines BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 304 Fic. 5.— Distribution of Taiyubius. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 305 of the gonopods in the females, these standing in conspicuous contrast to the slender ones uniformly present in Paitobius. 'Taiyubius is a prevailingly western genus, all the species occurring west of the Rockies excepting harrielae which is found commonly in Colorado but also in New England (Figure 5). Key to Species of Taiyubius. a. Each coxa of last one or two pairs of legs armed laterally with a spine. b. Ventral spines of penult legs 1, 3, 3, 2; of the anal, 1, 3, 3, 1; articles of antennae mostly 34 or 35. 7. satanus (Chamberlin). bb. Ventral spines of penult legs 1, 3, 3, 1; of anal legs 1, 3, 3, 0; articles of antennae 28-32... .. T. purpureus (Chamberlin). aa. None of the posterior coxae laterally armed. b. Third joint of about first nine pairs of legs with but two dorsal spines; ventral spines of second legs, 1, 3, 2; articles of an- tennae mostly 26 to 29... ...... T. angelus (Chamberlin). bb. Third joint of at most first three pairs of legs with two dorsal spines; others with 3; ventral spines of second legs 1, 2, 1; articles of antennae mostly 35 to 39. T. harrielae (Chamberlin). TAIYUBIUS ANGELUS (Chamberlin). Plate 5, fig. 4, 5. Lithobius angelus Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 155. Taiyubius angelus Chamberlin, Can. ent., 1912, 43, p. 176. Description.— Dorsum light brown to chestnut, the most caudal plates and the head with the first plate darker. Head often paler in front of the suture. Antennae concolorous with the head, pale distad. Venter pale testaceous to brown, the caudal plates con- siderably darker and more reddish. Prosternum and _ prehensors reddish brown, the latter paler distad. Legs colored like correspond- ing plates of venter, the posterior pairs being thus darker, brown to chestnut, uniform or sometimes a little paler distad. The body is often suffused in deeper tissues with a violaceous pigment which is commonly most evident in venter. Body considerably attenuated cephalad, with the tenth plate clearly wider than the eighth and than the head, and the first plate 306 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. narrower than the third. Body (in @ ) about 8 times longer than the width of the tenth plate. Width of head and of first, third, eighth, and tenth plates to each other as 40 : 36 :37 : 42 : 45. Head suborbicular, as long as wide or nearly so; widest at marginal breaks back of which the sides are convex and strongly converge, rounding about corners to the caudal margin which is nearly straight. Smooth and shining or under lens appearing very finely roughened over the middle portion. The usual median longitudinal sulcus beginning at a short transverse mark a little in front of the suture and extending cephalad. A wide, shallow longitudinal furrow each side of the middle running forward from the caudal margining sulcus. In some there is a more or less distinct transverse mark on each side a little ways caudad of the frontal suture. Hairs of moderate length, sparse. Antennae short or very short, reaching only to the fifth or sixth segment. Articles twenty-six to twenty-nine, but in most twenty- eight, which, beyond the first few, are short and decrease distad. Hairs rather long, not dense. Ocelli mostly ten or twelve in three series: e.g., 1 + 4, 3, 2; 1+ 4, 4,3; 1+.5, 3, 3. Single ocellus much largest, subvertically oval. Ocelli distinct and series regular; the ocelli of first and second series largest, decreasing ventrad and cephalad. Prosternum between 1.5 and 1.6 times wider than long. Distance between chitinous spots 4 times greater than length of the dental line and 2.25 times greater than width at level of bottom of mesal incision. Teeth 2 + 2, thee of each pair close together, of moderate size, the mesal tooth on each side directed somewhat mesocephalad and its apex caudad of the line tangent to apices of outer teeth. First dorsal plate smooth and shining like the head, under the microscope appearing very finely roughened; a transverse sulcus in front of caudal border usually continuous at each end with a longi- tudinal furrow parallel with the margin, while another longitudinal furrow may be more or less distinguishable each side of the median line. Other major plates also with a transverse caudal furrow and a lateral longitudinal furrow on each side together with a mostly more sharply impressed longitudinal furrow farther toward the middle, this diverging from the corresponding one of the other side caudad. Plates very finely rugose, becoming more distinctly roughened caudad. First ventral plate with a median basin-like depression, while the others show more or less distinctly the usual three longitudinal furrows, these becoming more distinct in going caudad; showing usually also two wide transverse depressions or furrows, one anterior CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 307 and one, more distinct, caudal, with sometimes finer lines and trans- verse rugae in the interval between the two. Coxal pores circular or slightly transversely elliptic: 4, 4, 4, 3; DONO WA Ora Osos OnO, Ol veLCS Spines of first legs, }-$-7°3°1 OF ococi- 5; of the second, - oe 353 of the third, (-4°3°3:3; of the fourth to ninth, §:}:5:3:3; of the tenth and eleventh, cosas; of the twelfth, [:333; of the thirteenth, ¢7°3:3°9 or g33c3 of the penult, 93:31, Claws two; of the anal, ¢y357,, claws two. None of the coxae laterally armed. Anal and penult legs short, moderately thickened, the femur and tibia weakly longitudinally suleate above. ; Gonopods of female of moderate size, but little or not at all extend- ing beyond caudal end of the last dorsal plate. Claw tripartite, the lobes acute, with the median one but little longer than the others. Mesal edge of first joint nearly straight. Basal spines 2 + 2, con- spicuously flattened above base, broad, widening distad to the begin- ning of the distal third of length from where narrowing to a point; edges often more or less minutely crenulate. Inner spine as usual, with broad surface set more or less at an angle with that of the outer one. Length 13 to15 mm. A female 13 mm. long has antennae 4.5 mm. long; anal legs 5 mm. long; width of tenth plate 1.6 mm. Tyre Locauiry.— California: Los Angeles. Taken also in California at Oroville and Santa Barbara. TaryuBius SATANUS (Chamberlin). Lithobius angelus satanus Chamberlin, Can. ent., 1911, 48, p. 380. Taiyubius satanus Chamberlin, Can. ent., 1912, 44, p. 176. Descrietion.— Dorsum brown, the caudal margins of the anterior major plates darker. Head dark brown, paler in front of suture. Antennae dark brown proximally, becoming pale distad. Prosternum dark brown with the prehensorial feet paler, orange. Venter pale brown, darker caudad, the last several plates being distinctly darkest; the anterior plates suffused with violaceous. Legs similar to venter and in part showing a violaceous tint as in most species of Paitobius; posterior pairs darker excepting the last two pairs which are orange. Body moderately attenuated cephalad; eighth and tenth plates nearly same in width, a little wider than head; first plate a little narrower than the third. Body in type (co) not quite eight times 308 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. longer than width of tenth dorsal plate. Widths of head and of first, third, eighth, and tenth plates to each other as 34 :31 :32 :36 :36. Head subrotund; wider than long (17 : 16). Widest near marginal interruptions, caudad of which the sides strongly converge. Caudal margin mesally ineurved. Two longitudinal sulci which, beginning close together at caudal margining sulcus, diverge cephalad and end abruptly a little caudad of the frontal suture. A transverse furrow is indicated a little in front of and parallel with the suture with the usual longitudinal median sulcus extending forward from it. Finely rough- ened. Hairs of moderate length, sparse. Antennae short; composed of about thirty-four or thirty-five com- pactly arranged articles of which the second is largest and those distad of the third short and very short. Hairs moderately long and sub- sparse. Ocelli six to nine in two or three series: e¢. g., 1 + 3, 2; 1+ 3, 3, 1; 1 + 3, 3,2 Prosternum between 1.6 and 1.7 times wider than long. Distance between chitinous spots three times the length of dental line and 2+ times greater than width at level of bottom of median incision. Median incision semicircular. Teeth large, acute. Excepting the first one, the dorsal plates are rather distinetly and considerably rugose. Each major plate with a median longitudinal suleus which is most distinct caudad; on each side a longitudinal sulcus diverges from the median one caudad, while caudad of its pos- terior end a shorter, sharply impressed one converges toward the corresponding one of the opposite side; farther ectad on each side a longitudinal sulcus extends parallel with margin for entire length of plate. Ventral plates with the usual three longitudinal sulci of which the median one is usually continuous entirely across the plate and is the most distinet, especially caudad. Anterior plates usually trans- versely broadly depressed. Spines of fir st legs, “ath 3 of the iwehal TT OF gyri; of the third, 0.04: asi; of the fourth, ¢ oe Le or ha aos of the fifth to ninth, PRES 25 of the tenth and elev enth, ° Oooo OF oF rea; of the twelfth, a a 3-5, the anterior dorsal spine of the last two articles minute; of the thirteenth, §:7: - 35 (left) or j4°3:375 (right); of the penult, mrs 3 — claws three; of the anal, 72353) 0 , Claws two. Last two pairs of coxae laterally armed. Posterior legs slender, of moderate length. In both anal and CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 309 penult pairs the prefemur, femur, and tibia are longitudinally weakly furrowed above. Gonopods of o clearly extended; narrow, distally rounded; pale. Length near 10 mm. In the male type, which is 10 mm. long, the antennae are 3.8 mm. long, the anal legs 4.4 mm. long, and the tenth plate 1.25 mm. wide Praematurus.— Coloration as in adult. Antennae very short. Articles twenty-eight (left) — thirty Sour (right), the two antennae differing in length correspondingly. Ocelli 1+ 3, 2. Single ocellus largest, the most caudal one of upper series next; ocelli of lower row smallest. Prosternal teeth 3 + 3, a small extra one being present at edge of incision on each side, the other two of usual proportions. Coxal pores small: 3, 3, 3, 2. Spines of first legs, ot'o'0-1; of the second, %°5°s*7; of the third, o-1; Of the fourth and fifth, (4:94:14; of the sixth, 45334"; of the seventh, OO of the eighth to eleventh, (4°33; of the twelfth, in) . iO, 8} 7,7 mo 12.9 of the thirteenth, ¢4°3'3-3 or 9-1°3°373; “of the penult, TOIT 1, 0, 3,120 0-13-32 claws ae ee; of the anal, 9-73-35, claws two. Last two coxae laterally armed. Gonopods of o projecting caudad beyond sternite as a pale conical process. Length 9 mm. Tyre Locatiry.— California: Oakland. TAIYUBIUS HARRIELAE (Chamberlin). Plate 5, fig. 6 Lithobius harrielae Chamberlin, Proc. Acad. nat. sci. Phil., 1906, p. 3; Ann. Ent. soc. Amer. 1909, 2, p. 191. Taiyubius harrielae Chamberlin, Can. ent., 1912, 44, p. 176. Description.— Dorsum light brown or testaceous, with a broad median longitudinal dark stripe extending the entire length or some- times evident only over caudal portion. Head testaceous, blackish or dusky in eye-region and sometimes in a curved band along the frontal suture. Antennae testaceous or yellowish. Prosternum and prehensors yellow or testaceous. Venter yellow, a few of the most caudal plates darker. Legs light yellow, with the caudal pairs some- what darker, sometimes dark brown or blackish on ectal surface espe- cially the joints proximad of tarsi. 310 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body moderately narrowed cephalad, the tenth plate being clearly wider than the head and slightly wider than the iar plate. Third plate a little wider than the first. Length, in type, 7.5 times greater than width of tenth plate. Widths of head and of ne third, eighth, and tenth plates to each other as 40 : 35 : 36:45 : 46. Head subcordate; wider than long in ratio 17 : 16, or sometimes equal in length and breadth. Caudal margin straight or but slightly incurved. A transverse furrow more or less evident caudad of the frontal suture, the furrow or branches from it usually angularly bent caudad at the middle. The usual median longitudinal sulcus in front of the suture, this sulcus commonly double with the parts diverging cephalad. A median longitudinal furrow on caudal portion of head while from one to three curved furrows may be traced more or less clearly on each side of it. Antennae short; composed of from thirty-three to forty-one articles of which those between the third or fourth and ultimate are short and very short and closely compacted. Hairs of moderate length, not dense. Ocelli ten to fourteen in three series: ¢.g., 1 + 4, 3,2; 1+ 4, 4,3; 1+ 5, 4, 3. Single ocellus much the largest, vertically oval. The most caudal ocellus of top series largest of the seriate ocelli. Prosternum 1.5, or a little more or less, times wider than long. Distance between chitinous spots 2.4 times width at level of bottom of mesal incision; from 4 to 4.5 times the dental line. Teeth 2 + 2, small, line of apices clearly recurved. Margin ectad of teeth convexly rounding ectad and then caudad as usual. First dorsal plate 1.8 times wider than long. Plates only very finely roughened, to the naked eye appearing nearly smooth and shining, especially so the first one. There may usually be traced on the major plates a longitudinal furrow on each side diverging from the median line caudad, with sometimes traces of a median longitudinal furrow; in some a second longitudinal furrow is evident ectad of the one first mentioned and toward which it converges caudad. Ventral plates without conspicuous markings, to the naked eye usually appearing smooth and shining. Coxal pores small and very small: 3, 4, 4, 3; 3, 4, 5, 3; 4, 5, 5, 3; 4,5, 5,4; 4, 4, 5, 4; 4, 6, 6, 5; ete. aie ae 0, 221, 0.2.21 0,0,3,21, opines of first legs, 9-y-1-9-1; of the second, (7 L21 OF 0,131; of the oe) KONO! 22, 2 0, eae , 0,0, 3,2, 2, third, a Tae oe ao tec) Ol Nan sads at the fourth and fifth, 99 9°9 Da 5 4 ha ae or ot: a. 31; of the sixth to eleventh, }°334; of the twelfth, TOS MOB wal 0, tee 4 pete aT OnSealoil Coes ose OF on3a1,. of the thirteenth) i734; of the penult, CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. oll isa; Claws two; of the anal, rat - 3 coxae laterally armed. Claw of female gonopods of moderate size, rather wide, tripartite; lobes acute, the median one not much exceeding the lateral ones. Basal spines 2 + 2 or, exceptionally 3 + 3, the innermost spine on each side in the latter case being much the smallest of the three; spines very broad, widest toward middle of length, acutely pointed, shaped somewhat like a broad arrow-head. Gonopods of male clearly exposed; subtruncate distally. Length mostly 10 to 12 mm. Type Locatiry.— Colorado: east of Glenwood Springs. Taken also in Colorado at Salina and in Massachusetts at Cam- bridge and Mattapan. 1, 0 >; claws two. None of the TAIYUBIUS PURPUREUS (Chamberlin). Lithobius purpureus Chamberlin, Proc. U.S. N. M., 1901, 24, p. 24. Taiyubius purpureus Chamberlin, Can. ent., 1912, 44, p. 176. The original description of the types, rearranged, follows: — Color dark purple-brown, the purple tint often conspicuous and unmixed. Head and legs yellowish brown, the tarsal joints of the last pairs of legs commonly lighter; antennae dark, yellow or rufous at ends. Antennae composed of from twenty-eight to thirty-two, articles which, excepting the first few, are short or very short; length 2.3- 3 mm. Ocelli mostly twelve or thirteen. Prosternal teeth normally 2 + 2, rather large and pale (rarely the teeth may be doubled). Coxal pores small or moderate: 2, 3, 3, 2; 3, 4, 4, 3. Ventral spines of first legs 0, 0, 0, 0, 1; of the penult, 0, 1, 3, 3, 1; of the anal, 0, 1, 3, 3, 0 (or, rarely, 0, 1, 3, 1, 0). Posterior coxae laterally armed. Claws of female gonopods rather long, tripartite, the lobes acute, with the median longest and the outer one smallest, sometimes near base and inconspicuous. Basal spines short and broad, widest at middle. Length 7 to 8.5 mm. Length of anal legs 3-3.3 mm.; of antennae 2.3-3 mm. Type Locaity. Utah: Salt Lake City. a2 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. The types (U. S. N. M. 786) were found under sticks, boards, and logs laid on fine loose soil in and near growths of willows on the banks of the Jordan River. Sonrspius Chamberlin. Can. ent., 1912, 44, p. 176. Head with distinct lateral marginal interruptions. Antennae short; articles normally twenty. Ocelli present; seriate; single ocellus enlarged. Prosternal teeth 2 + 2 or 3+ 3; spines slender. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced. Coxal pores circular or slightly elliptic; uniseriate. Gonopods of male uniarticulate. Claw of female gonopods short; tripartite, with median lobe not much longest. Spines 2 + 2; short and relatively very wide. Tarsi of anterior legs divided. Posterior legs of male without special lobes or modificatfons.. Third joint of all legs caudad of first or second pair with three dorsal spines or those cephalad of sixth with but two. Fifth joint of legs between third or fourth and twelfth with two dorsal spines. Dorsal spines of anal legs 1,,0,3, 1,0; ventral, 0; 1,3, 2, 0 or 0,1,3, 2, 1; cdaws two or three. Dorsal spines of penult legs, 1, 0, 3, 1, 1; ventral, 0, 1, 3, 3,1 or 0,1,3,3,2; claws two or three. Dorsal spines of twelfth legs 1, OF 3501; 1 tot, 0;.3;2,:25 ) ventral, (030; 393,.25 Dorsal vspmes of first legs 0, 0, 1, 1, 1 to 0, 0, 2, 2,1; ventral, 0,0, 1, 3, 1 and 0,0, 1, 3, 2 ColOROR 2, deand. On0502 257 Last two or three pairs of coxae laterally armed. Length 8-12.5 mm. Typr.— S. bius (Chamberlin). This genus seems to be very close to Nadabius but differs from it especially in the strongly produced angles of the ninth, eleventh, and thirteenth dorsal plates and in lacking a tibial crest in the anal legs of the male. It has thus far been found only in the section of the United States east of the Mississippi River and north of the Ohio River (Figure 6). Key to Species of Sonrbius. a. Anal legs bearing 3 distinct claws. b. Dorsal spines of twelfth legs 1,0, 3,2,2; prosternal teeth 3+3; length of type, o, 12.5 mm.)..S. biws (Chamberlin). 313 CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. Fic. 6.— Distribution of Sonibius. 314 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. bb. Dorsal spines of twelfth legs 1, 0,3, 1,1; prosternal teeth 2.2 lene ts: 5— Osama 6) 0e\scb a vee: S. parvus, sp. nov. aa. Anal legs bearing only 2 distinct claws. b. Penult legs with but 1 accessory claw; ventral spines of penult legs, normally, 0; 1,353; Ws0.>...: S. politus (MeNeil). bb. Penult legs with 2 accessory claws; ventral spines of penult legsmormallyaQil0,3.j3-2 cee ee S. numius (Chamberlin). Sonrisius Brus (Chamberlin). Lithobius bius Chamberlin, Can. ent., 1911, 48, p. 102. Sonibius bius Chamberlin, Can. ent., 1912, 44, p. 177; 1914, 46, p. 302. DescripTion.— Dorsum uniform brown. The head a little darker, with the antennae similar but becoming pale distad. Prosternum like head, the prehensors paler. Venter testaceous, the posterior plates darker. Legs light brown or testaceous dorsally, ventrally paler; caudal pairs dark proximally but distal articles yellowish or light orange. Body rather robust, being not quite seven times as long as width of eighth and tenth plates which are equal in width. Conspicuously narrowed cephalad, with first plate narrower than the third. Widths of head and of first, third, eighth, and tenth plates to each other as AG S405 42 2b 2e 52. Head wider than long in about ratio 23 : 21. Subrotund, the sides caudally strongly rounded. Caudal margin distinctly incurved mes- ally. Smooth and shining. Antennae short, reaching fifth or sixth segment. Articles twenty (left) — twenty-three (right), moderately long, cylindric, decreasing distad. In the antenna with twenty-three articles the distal ones are proportionately shorter. Ocelli twenty-two in five series: 1 + 5, 6, 5, 5; 1+ 5, 6, 5, 3, 2. Prosternum 1.5-+ times wider than long. Distance between chiti- nous spots two times, or a little more, greater than width at level of bottom of median incision; about 3.7 times the dental line. Teeth 3 +3; acute; subequal. Dorsal plates smooth and shining. Posterior angles of ninth, eleventh, and thirteenth acutely and strongly produced. Ventral plates sparsely hirsute; the more caudal ones very finely roughened. Longitudinal furrows mostly obscure. A wide, shallow, transverse furrow showing in some on caudal portion. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 315 Coxal pores small, circular: 4, 4,5, 3; 4, 6,5 foe Spines of first legs, (U4°3°33 of the second, (¢ 1a of the third, 5; of the fourth to ninth, ¢, 0° i >: 3.22. of the tenth, ¢°y; 3. 35; of the 3, 2 “a 3 TR SY O03, tot i enth and twelfth, -¢°3-3-9; of the thirteenth, 9°7°3°3:9; of the penult, 3,1, 3, 3, 0, ah 5, claws three; of the anal, . i * 2 Sorre 3. sy Claws three. Last hree pairs of coxae laterally armed. Anal legs in male short, the femur very moderately thickened, com- planate and longitudinally shallowly furrowed above; the tibia more slender; femur shallowly longitudinally furrowed above. Gonopods of male but little exposed; broad and short. Length of type (o7) 12.5 mm. Type Locauiry.— Michigan: Saunders. Taken also at Douglas Lake, Mich. SONIBIUS PARVUS, sp. nov. Plate.6, fig. 5: DescripTIoN.— Dorsum brown with first dorsal plate typically darker. Head darker, dusky brown, either with or without a dilute chestnut tinge. Antennae dark brown proximally, becoming orange or rufous at tips. Prosternum and prehensors testaceous. Venter light brown, the caudal plates darker. Legs of nearly same color as the corresponding ventral plates, the last pair thus darkest but tarsi of these orange or rufous. Body proportionately broad, being usually only about 6.25 times longer than width of tenth plate. Strongly narrowed cephalad. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 49 : 44:48 : 58 : 60 : 60. Head somewhat wider than long (49 : 46). Antennae short, the articles between the first few and the ultimate very short and closely united. Ultimate longer than the two preced- ing together. ey ee to seventeen in three or four series: e.g., 1 + 4, 4, 3; 1+5 4,2; 1+ 5, 4, 3, 1. Single ocellus considerably largest. Organ ” Tomésvary in usual position, of about same size in outline as the smaller ocelli. Prosternum with incision rather large, strictly v-shaped. Teeth equal or nearly so, small. Spine of usual form and in usual position. Sides of anterior portion of prosternum straight or nearly so from spine caudoectad, not convexly rounding out as more usual in the genus; 316 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 13 times wider than long. Distance between chitinous spots in type 2.6 times width at level of bottom of incision; 3.55-++ times dental line. First dorsal plate with sides moderately convex and caudally con- verging; 1.8 times wider than long. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly acutely produced, while in fully grown individuals the posterior angles of the seventh and also of the sixth may also be very weakly extended. Coxal pores small: 3, 4, 4, 3; °3, 4, 4, 4. Spines of first legs, 9-0) 0°24 or os , ‘134; of the second, 9 third, po: 73:1; of the fourth, } 0 02 2 1. of the fifth, ha ! of the sixth, » O° 2 3 5; of the seventh to eleventh, ¢: 0-0; m2 4 ;; of the twelfth, 1.0.3.1]. of the thirteenth, £9311; of the penult, 22:2 claws three; of the anal, (3:24 or a : 2 uy the claws three. Last two pairs of coxae laterally armed. Claw of female gonopods short; moderately curved; tripartite; median lobe somewhat longest, the lateral ones subequal and at same level or nearly so, all acute. Outer basal spine considerably larger than the inner one, broad and short, the sides subparallel from base to beginning of apical division which is very acute, denticulate on each side at base of apical portion and sometimes proximad of it. Inner spine in ventral view similar in shape but proportionately narrower (Plate 6, fig. 5). Length 8.5 to 9.5 mm. Typr.— M. C. Z., No. 146. Vermont: Grout’s Mill, near Mt. Stratton. Taken also in New York at Big Tupper Lake, St. Lawrence county, Thomas and F. Kk. Barbour. ae gaoa; of the 2,1 2, 0. 0, 0, 2, 2, 2. 3,2» OF 0, 0,2, 3,25 tO 11 3,2 Sonipius Potitus (McNeill). Lithobius politus McNeill, Proc. U. S. N. M., 1887, 10, p. 261. Chamberlin, Can. ent., 1911, 43, p. 103. Lithobius howet Bollman (ad. part max.), Proc. U. 8. N. M., 1889, 11, p. 409. Sonibius politus Chamberlin, Can. ent., 1912, 44, p. 177; 1920, 52, p. 95. Derscriprtion.— Dorsum brown, uniform or nearly so. Head either nearly same as dorsum or considerably darker. Antennae brown, having often a reddish or purplish tinge proximally, paler distad; in one specimen the first article is very much lighter than those immediately succeeding. Venter testaceous, the anterior plates almost clear yellow. Prosternum testaceous, the prehensors paler. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. SLT Legs like corresponding parts of venter, caudal pairs a little darker proximally but dilute orange distally. Body proportionately rather wide being mostly about 7 times longer than the width of the tenth plate. Body moderately narrowed cephalad and the first plate clearly narrower than the third. Widths of head and of first, third, eighth, and tenth plates to each other as 37:30:37 :43 : 48. Head wider than long in nearly ratio 14:18. Sides caudad of eyes nearly straight excepting in front of caudal corners, where more convex. Caudal margin mesally incurved. Marginal breaks not especially conspicuous. Smooth and shining. Antennae short, reaching in most only to the fifth or to edge of sixth segment. Articles twenty, mostly short, decreasing distad gradually, or, in some, those distad of the first six to eight abruptly shorter. Ocelli fifteen to twenty in mostly four series: e.g., 1 + 2, 3, 4, 5 1+ 5, 5,3, 2. Single ocellus much the largest, oval. Prosternum 1.5 times wider than long. Distance between chitinous spots 3.7 to a little less than 4 times the dental line; between 2.5 and 2.7 times width at level of bottom of mesal incision. Sides of anterior portion slanting almost directly from outer teeth. Teeth small, acute, with line of apices conspicuously reentrant; 2 + 2 or 3 + 3, in the latter case the most mesal tooth on each side commonly very small. Dorsal plates smooth and shining. Hairs short and sparse, more numerous on caudal plates. Ventral plates appearing smooth and shining, or under lens some are seen to be finely roughened, especially the more caudal ones. Hairs more numerous caudad. Coxal pores of medium size, circular or slightly transversely elliptic: 4, 63.6,/59 45 6,6) 65 5; 6, 6, 6; 4, 5, 5, 4. pel 0, 0; 1,.2, 1 ers ak 0, Spines of aa legs, 0-0 00-13-11 OF 00-1-3-2; Of the second, i371 OT 0: of the third, 9:9°3:3:3; of the fourth to tenth, 9-9-9°3"3; of the elev any 0, 0, 3, 2, 2 15,053,050 Bab ahs 00-33-23; Of the twelfth, 0-03-32; Of the thirteenth, } tress; of the 1,0,3.1,1 penult, 9°7°3°3°1, Claws three, of which the anterior accessory is very small; of the anal, }°32°¢ or ¢'4'3'2'1) Claws two or a third represented by an obsolete point. Last two or three pairs of coxae laterally armed. Posterior legs of male short and slender; the tibia and femur of both anal and penult pair shallowly longitudinally furrowed above. Gonopods of male exposed as moderately large rounded or hemi- spherical eminences bearing two or three bristles. 0, 0, 2 0, 0, s\e ’ 318 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Gonopods of female considerably exceeded by dorsal plate of anal segment. Claw short and relatively wide; tripartite; the lobes short and the median not much longer than the others. Basal spines short and proportionately very wide, with sides subparallel to the distal third or fourth which is acuminate with apical angle low or obtuse. Length 8 to 11 mm. Tyre Locauiry.— Indiana: Dublin. Taken also in Indiana: at Bloomington, Kokomo (Bollman coll.). Michigan: Ludington. Illinois: Peoria. Ontario, Ottawa, Quebec. Mr. Bollman appears undoubtedly to have applied his name Litho- bius howei to two entirely different species. The type of howe? is a male from Fort Snelling, Minn., and an examination of this shows it to be closely related to, and possibly identical with, L. mordax and to differ widely from the present species. In Bollman’s second and somewhat fuller account of the species (Proc. U. S. N. M., 1887, 10, p. 255 and 258) he gives the anal legs as having but one claw, with ventral spines 1, 3, 3, 1 and length 7 mm.; ventral spines of first legs 2, 3, 2; ocelli 25 in number; total length 15 mm., ete.; all of which points separate it clearly from politus; politus is first deseribed in this same paper under the heading Lithobius politus McNeill MSS., and is reported from Dublin and Bloomington, Indiana, and from Luding- ton, Mich. However, in Mr. Bollman’s Catalogue of the Myriopods of Indiana” (Proce. U.S. N. M., 1888, 11) and in his posthumous Cata- logue of North American Myriapoda, politus is not mentioned, while howev is listed from Dublin, Bloomington, and Kokomo. An exami- nation of the specimens from these localities in Mr. Bollman’s collec- tion labeled as L. howei show them to be the same as politus. It is thus evident that Mr. Bollman intended to drop politus as a synonym of howei; but that the two species are widely different is evident from the original descriptions, even though the presumed type of hower in the National Museum, which does not agree wholly with the description, might be regarded as possibly incorrectly labeled. Sonrpius NumIus (Chamberlin). Lithobius numius Chamberlin, Can. ent., 1911, 48, p. 102. Sonibius numius Chamberlin, Can. ent., 1912, 44, p. 177. Description.— Dorsum nearly uniform brown. Head darker. Antennae brown or reddish brown proximally, paler distad. Venter testaceous. Caudal legs dilute orange distad. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 319 Body in type seven times longer than width of tenth plate; more strongly narrowed cephalad than in politus. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 36 : 33 : 35 243 343. Head considerably wider than long (12 : 11); broadly subcordate; caudal margin mesally distinctly incurved. Antennae short, reaching the fifth segment. Articles of moderate length, decreasing regularly distad. Hairs of moderate length, sub- dense. Ocelli twenty to twenty-three in five series: e.g., 1 + 2, 5, 5, 5, 4; 1+ 3, 5, 5, 4, 2. Single ocellus oval, large. Prosternum about 1.5 times, or a little more, wider than long. Distance between chitinous spots 2.8 times width at level of bottom of incision, a little less than 4.5 times as great as the dental line. Teeth 2 + 2, small, acute, the line of apices reentrant. Spine almost contiguous with ectal tooth, bristle-like. Coxal pores in type: 4, 6, 6, 5 ; 0.0,1,2,1 (ONO Dae ), 2,.2,1 Spines of first legs, 0 C31 OT oo 131; of itae second, ? 0: 5 (3:1 OF 0,3, 2,1 agate 3,.2,2 paieiaaai of ithe third, } 00,231; Of the fourth, ? a (: 231; Of the fifth to eleventh, ? ORG oe 5; of the twelfth, a : - 33 of the thirteenth, a: t 2 Si of the penult, c33 sy, Claws three; of the anal, (433°, claws two Last two pairs of coxae laterally armed. Claw of female gonopods short; tripartite; lobes short with the median but little longer than the others. Spines broad; sides sub- parallel or a little diverging to acuminate apical portion, outer longer than inner; larger than in politus. Length of type (@ ) 10.5 mm. Tyre Locauiry.— Wisconsin: Haugen. Very close to the preceding species and may possibly prove to be only a variety of it. Napasius Chamberlin. Bull. M. C. Z., 1913, 67, p. 62. Head with distinct marginal breaks. Antennae short or very short; articles nineteen to twenty-three, but normally twenty. Eye consisting of ocelli arranged in from three to five series; single ocellus enlarged. Prosternal teeth 2 + 2, or occasionally 3+ 3 and 4+ 4; line of apices more or less recurved. 320 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Posterior angles of none of the dorsal plates produced or those of eleventh and thirteenth weakly so. Coxal pores circular; uniseriate: 3, 4, 4, 3 to 5, 6, 6, 5. Gonopods of male uniarticulate; bearing from two,to four long bristles. Claw of female gonopods tripartite, or rarely entire by obliteration of lateral teeth. Basal spines 2 + 2, typically short and rather broad with sides proximad of apical acuminate portion subparallel or slightly diverging from base distad. First article of gonopod not excavated proximally. Tarsi of all legs divided. In the male the anal legs are short and but little thickened; aoraie longitudinally more or less suleate; the fifth joint bearing a character- istic low crest at the distal end mesad of median line, the crest bearing a variable number of hairs, usually of special type. Last one to three pairs of coxae laterally armed. Third joint of all legs caudad of second with either two or three dorsal spines, the number bearing two variable. Fifth joint of all legs between first or second and twelfth with two dorsal spines.’ Dorsal spines of anal legs 15053, 1,,0-" ventral, Oreo ao, al torOsel oes Ole clawentone. Dorsal spines of penult legs 1,0, 3,1,1; ventral, 0,1, 3, 3,2; claws three or, less commonly, two. Dorsal spines of thirteenth legs IOS lordwOra, ou2-mventrale O01 3.5.2 1Or OMOnoe sD onsal spines of tw agen legs 1, 0, 3, 2, 2 or rarely only 1, 0, 3, 1, 1, the coxal spine sometimes missing; ventral 0, 0, 3, 3,2 or 0,0, 2,3, 2. Dorsal spines of eleventh legs 0, 0, 3, 2, 2; ventral 0, 0, 2, 3, 2 or 0, 0, 3, 3, 2 Dorsal spines of first legs 0, 0, 2, 2, 1 to 0, 0, 3, 2,1 and 0, 0, 3, 2, 2; ventrallO; 0-12 Ito 0) 02253. 2 Length 7 to 16 mm. but mostly above 10 mm. Typr.— JN. iowensis (Meinert). The species of Nadabius are for the greater part of medium size, prevailingly brown or chestnut in color with the head darker. In the north central states they are the most abundant in individuals of the Lithobiidae. Key to Species of Nadabius. a. Anal leg with 2 distinct claws. b. Dorsal spines of thirteenth legs 1, 0, 3, 2, 2; only the last pair of coxae laterally armed. c. Prosternal teeth 2 + 2; ventral spines of anal legs normally Oi A Oa Sell EET. aU Sees N. pullus (Bollman). CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. Syall Fic. 7.— Distribution of Nadabius. 322 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ce. Prosternal teeth 4 + 4 or sometimes 3 + 3; ventral spines of anal legs normally 0, 1, 3,3, 1...N. aristeus, sp. nov. bb. Dorsal spines of thirteenth legs 1, 0, 3, 1, 1; last 2 pairs of coxae laterally armed. c. Ventral spines of anal legs 0,1,3,2,0; dorsal spines of tweltthe legs (O803 3; lala een a N. oreinus, sp. nov. ce. Ventral spines of anal legs 0, 1, 3, 3, 0 or 0, 1, 3, 3, 1; dorsal spines of twelfth legs 1,0, 3, 1,2 or 0,0, 3, 1, 2. N. coloradensis (Cockerell). aa. Anal leg with but 1 claw. b. Dorsal spines of thirteenth legs 1, 0, 3, 1, 1. c. Dorsal spines of twelfth legs 1,0, 3, 1,1. N. mesechinus (Chamberlin). cc. Dorsal spines of twelfth legs 1, 0,3, 2, 2 to 0,0, 3, 2, 2 and (Oe Oaks 4 d. Prosternal teeth 3 + 3 or4 + 4....N.iowensis (Meinert). dd. Prosternal teeth 2-+ 2 or rarely 2+ 3. N. eigenmanni (Bollman). bb. Dorsal spines of thirteenth legs, 1,0, 3, 2, 2. N. holzingeri (Bollman). NADABIUS ARISTEUS, sp. nov. Plate 6, fig. 3, 4. Description.— Dorsum brown; a darker median longitudinal stripe more or less evident. Head and first dorsal plate much darker, almost black. Antennae very dark, brown to blackish, proximally, becoming paler and more or less rufous distally. Prosternum like head; prehensors rufous distally. Venter light brown or testaceous, darker caudally. Legs light brown, caudal pairs somewhat darker. Body robust only 6 to 7 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other as 90 : 78 : 83 : 102 : 101 : 98. Head subcordate, widest at level of breaks which are pronounced and rectangular. Wider than long in about ratio 15 : 14. Antennae moderately short. Articles twenty to twenty-one; of intermediate length, subeylindric, decreasing distad. Ultimate article shorter than the two preceding taken together. Ocelli twelve to eighteen in three or four series: e.g., 1 + 4, 4, 3; 14+ 4,3,3,1; 1+ 4,4, 4,3; 1+4, 5, 4,3. Single ocellus clearly CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE, 320 exceeding any other in size, or in largest specimens not greatly differing from most caudal ocellus of dorsal series. Prosternal teeth 4 + 4, rarely 3 + 3, acute, the most mesal on each side smallest. Incision small, between v- and u-shaped. Spine slenderly acuminate (Plate 6, fig. 4). In ine 1.6 times wider than long. Distance between chitinous spots 2 times width at level of bottom of median incision; 2.8 times the dental line. First dorsal plate widest anteriorly; sides convex, converging moderately caudad; 1.7— times wider than long. Posterior corners of ninth plate rectangular or a little rounded; those of the eleventh a little produced, those of the thirteenth more strongly so. Coxal pores: 4, 5, 5, 5; circular or in part transversely subelliptic. Spines of first legs, 0.0, re 4d op 0: i243 of the second, 9'9'3 oe) Poe ss of the third, 0.0.2, eo tO COLE _ of the fitth, ia a of the sixth, 00222 or Pee22. of the seventh, pee24, oS OF SS eighth and ninth, foes; of the tenth, no na or eleventh and twelfth, 0,0, 25; of the thirteenth, ; er Ta 3 e; of the anal, oO Pore the penult, j-{°3'3°2, or occasionally §}}3'3, Claws thre z 5. 7 1, claws two. Anal coxae laterally armed. Tibia of anal leg in male with a sharp edged dorsal longitudinal ridge showing at its distal end the usual crest which is low and not much elevated distad. Gonopods of male distinetly exposed; distally rounded; bearing two bristles. Claw of female gonopods short and broad, tripartite, with the median not much larger in size than the dorsal, these two bluntly rounded and short, the ventral tooth smaller and more proximal in position. Spines proportionately very short, broad, but little nar- rowed from base to distal division which is He and apically rounded. Length 11 we 16mm. A male 11.5 mm. long has antennae 4 mm. and anal leg 4.2 mm., with tenth plate cir. 1.9 mm. wide. Praematurus.— Coloration as in maturus, or a little lighter through- out. Antennae short; articles twenty (on one side in one specimen only seventeen). Ocelli 1 + 4, 3,3; 1+ 4, 4,3. Relations as in adult. Prosternal teeth small 3+ 3. Incision widely v-shaped. Coxal pores: 3, 4, 4, 3; very small. Claw of female gonopods very short, thin, bilobed or trilobed with the outer lobe very small and the inner lobe blunt. Spines 2 + 2, but the inner one of each pair minute, appearing, in specimens de- 324 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. scribed, as a mere point; the outer spine already rather robust though proportionately more slender than in the adult. ONO Ra . (O(a) pb Spines of first legs, ¢-o-0-a:43 of the second and third, 9:9: Sots of the ” 0;.0,.2,.2, 22 . ( 2 ake fourth and fifth, 00-231; of the sixth to eleventh, i eeereg Ge line 0, 0, ois 2 : PROevene tO soene twelfth, jps35; of the thirteenth, § + zo OF Gores; of. the anal, 1, 0, 3,1, 0 01331 tee pair of coxae laterally armed. Length 10-11 mm. (Ithaca, N. Y.). Type.— M. C. Z., No. 247, New Jersey: Macapin, November, 1913 (J. H. Emerton). Taken also in New York at Ithaca, and in Massachusetts at Wor- cester. Napapsius PuLLUS (Bollman). Plate 7, fig: 1-3: Lithobius pullus Bollman, Amer. nat. 1887, 21, p. 81. Proc. U.S. N. M., 1887, 105: 257. 1888-11 p. 409" Bulls 46) U. S25 Ne M., 1893; ps 195212 110, 184. Chamberlin, Can. ent., 1911, 43, p. 103. Lithobius elattus Bollman, Proce. U. 8. N. M., 1889, 11, p. 348. Chamberlin, Ann. Ent. soe. Amer., 1911, 4, p. 40. Descrretion.— Dorsum brown. Head darker, mostly somewhat chestnut. Prosternum dark brown, the prehensors lighter. Venter paler than dorsum. Legs similar to venter; posterior pairs light distally as usual. Body in types described 7.66 to 8 times longer than width of teeth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 33 : 29 : 31 : 39 : 39 : 37. Head precisely equal in length and breadth, or very nearly so. Lateral margin a little incurved midway between break and caudal corner, the latter widely rounded. Antennae short or very short, commonly reaching only upon fifth body segment. Articles normally twenty, these moderately short. Ocelli mostly near ten or eleven in three or four series: e. g., 1 + 4, 3,2; 1+ 4, 3,3; and 1 + 4, 3, 2, 1,a common arrangement. Eye- patch subtriangular, with apex ventrad. Of the seriate ocelli the first one of the top series is largest as usual, the others not much differing in size excepting the most ventral, which may be considerably reduced. Prosternal teeth 2-+ 2, of moderate size, equal, acute. Spine long, stout at base but distally slender and acute. In type measured, prosternum is 1.52 times wider than long. Distance between chitin- CHAMBERLIN: NORTH AMERICAN LITHOBITDAE. 325 ous spots 2.2— times width at level of bottom of median incision and 3.7 times the dental line. First dorsal plate 1.65 times wider than long. Posterior angles of none of the dorsal plates at all produced. Coxal pores rather small, circular: Si, 4b ak Bie 4, > erie Spines of first legs, a 1 3 i; of the second, 7 0-0, aa 4 or 0: o : a3 of the third to seventh, Oo 7 2 or? et a2; of the eighth, bas: 3 to 0, = et of the ninth, ¢ 0° “f a 2 Sor a . 253 of the tenth to twelfth, 5; oe 3, a 5; of the thirteenth, 7 : : ae of the penult, 0 t : 5, claws two, with the acces- UO Mosnls 0 oslenO sory one large; of the anal, 0:1°3'3'0, Very rarely 9°1°3-3-0, Claws two. Last pair of coxae laterally armed. Anal leg of male short, moderately thickened; fifth joint at distal end on dorsal surface and toward mesal side with a low keel-like crest which is most elevated distad; third, fourth especially, and the fifth joints longitudinally furrowed dorsally. In the female the last two pairs of legs are enlarged and furrowed as in the male but the anal pair lack the crest on fifth joint. Gonopods of male cuneate; each bearing three long bristles. Claw of female gonopods short and broad; distinctly tripartite, with the middle lobe longer and more acute than the lateral. Basal spines short and relatively very broad; distally acutely rounded; inner spine a little smaller than the outer, its principal plane set almost at right angles to that of the outer and its edge being thus presented ventrad. Length 9-12 mm. a male 11.5 mm. long, has antenna 5 mm. long, anal leg 3.6 mm. long, and tenth plate 1.4-+ mm. wide. Tyee Locauiry.— Indiana: Bloomington. Taken also in Illinois at Dwight. Tennessee: Johnson City, Russellville. Virginia: Chatham, Lynchburg, Natural Bridge, Bal- cony Falls, Marksville. West Virginia: White Sulphur. D. C.: Washington. The description above is based chiefly upon a pair of the cotypes from Bloomington. NADABIUS OREINUS, sp. nov. Plate 5, fig. 7; Plate 6, fig. 1. Description.— Brown. The head in front of suture paler. Antennae brown, yellow at tips. Prosternum brown, prehensors yellowish. Venter pale, of very dilute olivaceous cast, the last several plates much darker. Legs yellowish. 326 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 65 : 59 : 60 : 70 : 72 : 69. Head subcordate, conspicuously narrowed forward from a little behind eyes. Marginal breaks distinct but oblique. Wider than long in about ratio 65 : 60. Antennae short. The first four or five articles rather long, the others gradually decreasing in length distad. Ultimate article longer than the two preceding together. Ocelli in type fifteen or sixteen in three series: thus, 1 + 6, 5, 3; 1+ 6,5, 4. Single ocellus largest, subelliptic. Prosternum 1.62 times wider than long. Distance between chitin- ous spots 2.47 times width at level of bottom of sinus; 43 times the dental line. Sinus narrowly v-shaped. Line of apices of teeth nearly straight, the inner tooth being a little larger than the outer. Spine long and slender, bristle-like distally. Sides ectad of bristle convex. First dorsal plate only slightly narrowed caudad, the posterior corners strongly rounded as usual; caudal margin mesally incurved; 1.68 times wider than long. Caudal margin of ninth plate straight or with ends slightly bent caudad. Posterior angles of eleventh and thirteenth dorsal plates weakly produced. Coxal pores: 3, 4, 4, 4- 4, 4,4, 4. . . 0, 0, 2, 2, 0, 2,21, . Spines of first legs, (0-73 i; of the second, 9 C0132) Of the third to 0, 0,3, 2,2 0,0, 3, 1. Oua eleventh, j:j3'3'3; of the twelfth, o:0:3:3:2; of the thirteenth, 9:7°3:3°3; 0,31, AON of the penult, : 3-373, Claws three; of the anal, ¢73 sy, Claws two. Last two pairs of coxae laterally armed. Claw of female gonopods short and broad; tripartite, teeth distinct and acute, sometimes a fourth tooth present on mesal side. Basal spines relatively short and very broad; in ventral view the outer one is considerably broadest, widening from base to ‘the acutely acuminate distal portion (Plate 6, fig. 1). Immaturus.— Very pale, the head with the antennae and several most caudal plates yellowish. Posterior legs dorsally nearly white. Antennae already with the twenty articles present, those between the third and the ultimate relatively shorter than in the maturus. Ocelli 1 + 3, 3, pale. Single ocellus a little larger than the most caudal one of upper series. Seriate ocelli conspicuously decreasing in size cephalad. Prosternum as in adult. Coxal pores 2, 2, 3, 2. Gonopods of female short. The three articles developed. Claw appearing as a slender spine-like chitinous process which is entire CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 327 and slightly curved at tip. Distal article bearing a single ventral bristle, the second article two, and the first also two. Spines 1 + 1; moderately wide proximally, acuminate from near middle of Tenet Spines of first legs, (-p:g:3'13 of the second and third, }'h'¢-371; of the fourth and fifth, Push 1 of the sixth, (-¢¢o4; of the sev renth and eighth, (jt34; of the ninth (regenerated), vote4; of the tenth, foes 3; of the eleventh, 5 oa 2 of the twelfth, oo 3%5; of the thirteenth, j'1'233'2; of the penult, qirea's OF pega (on one side); of the anal, j oa eo Last two pairs of coxae laterally armed. Length cir. 5.5. mm. Typr.— M. C. Z., No. 271. California: Shasta Springs, August, 1909. The male of this species is unknown. p's NADABIUS COLORADENSIS (Cockerell). Plate 7, fig. 4. Lithobius kochii var. coloradensis Cockerell, Trans. Amer. ent. soc., 1893, 20, p. 370. Lithobius kochii Bollman (non Stuxberg), Proc. U. 8. N. M., 1889, 11, p. 349. Lithobius dopaintus Chamberlin, Can. ent., 1911, 43, p. 69. Lithobius coloradensis Chamberlin, Can. ent., 1911, 48, p. 70. DescripTion.— Dorsum testaceous to brown. Head dilute chest- nut, commonly darkest caudad of suture. Antennae brown proxi- mally, becoming rufous or yellowish distad. Prosternum brown, the prehensors paler. Venter greyish yellow to dilute brown, the caudal plates somewhat more deeply pigmented as usual. Legs yellowish grey or dilute greyish brown, the posterior pairs more strongly pig- mented, often light ferruginous or orange. Body from 8.5 to 9 times longer than width of tenth plate, commonly toward the latter limit. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 33 : 31 :33- :39 : 36. Head considerably wider than long (10 :9). Sides convex between eyes and marginal breaks, behind breaks conspicuously converging caudad, weakly excurved. Antennae short, mostly reaching to the fifth segment, more rarely to the sixth. Composed of nineteen to twenty-three articles, twenty being the usual number. Articles short, cylindric, or each only slightly enlarging distad. 328 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli ten to fifteen in three or four series: e.g., 1 + 4, 3, 2; 1+ 4, 3, oe) te ps oat ae 2d os) le roos on on alec lame 4, 3, 2. Prosternum in measured speciien 1.64 times wider than long. Distance between chitinous spots 2.4 times the width at level of bot- tom of sinus; 4 times the dental line. Teeth 2 + 2, rather small, acute; line of apices recurved. Spines much stouter proximally than the bristles, abruptly and characteristically narrowed near middle, the distal portion bristle-like. Median incision widely rounded at bottom, shallow. First dorsal plate 1.9-++ times wider than long; the sides convex, con- siderably converging caudad; caudal corners strongly rounded. Plates polished but under lens showing a very fine tuberculation or roughening. Angles of none of the dorsal plates at all produced. Anterior ventral plates nearly smooth, the plates caudad becoming more roughened and showing a more distinct median sulcus, this being deepest on the last two sternites. Coxal pores of moderate size to small, circular: 3, 4, 4, 4; 3, 4, 4, 3; 4,4,4,4; 4, 5, 5, 4; 4, 6, 6, 5. 0,0, 3, 2,1 0,0, 3, 2, 1 0, (Bs . oa - Ulites of first legs, ? COL 3D OL at or TO. 0, 2 3, 2) one second to eighth, Dye 2) 2 iy fa 0,0, 3 >; of the ninth to eleventh, ° 0: a 23,2 or 0.0.3, 3-9, Of the twelfth, TOs, gle ps al 10). O31": of the thirteenth and penult, {°1'3/3'3, claws two; of the anal, Lo, - 3p OF a . a4 31, claws two. Last two pairs of coxae laterally armed. Anal legs short and rather slender. In the male the third, fourth, and fifth articles are dorsally longitudinally furrowed; fifth article at distal end in usual place elevated into a low and rather broad crest similar to that of cowensis but lower and less conspicuous. Gonopods of male very short; bearing one or two bristles, the latter being apparently the normal “inal oe Claw of female gonopods short and relatively wide; tripartite, the three divisions distinct and subequal or the median usually a little longer than the others. Spines short and stout, the outer somewhat larger than the inner, in ventral view with sides parallel or a little diverging to about distal third or fourth of length, then abruptly narrowing and acuminate; wider surface of inner spine more or less oblique to that of the outer one. Length 10 to15 mm. A male 12.5 mm. long has antennae 3.5 mm. long; anal leg 4.25 mm. long; and tenth plate 1.42 mm. wide. Praematurus.— Color not evident in specimen described. Antennae very short; articles twenty. Ocelli 1 + 3,3; 1+ 4, 2. Caudal ocellus of upper row largest. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 329 Prosternum nearly as in maturus, teeth small and acute. 1.6 times wider than long. Coxal pores very small, circular: 2, 2, 3,3; 3, 3, 3, 3. Claw of female gonopods short and wide, three or only two teeth evident. Basal spines 2 + 2, the inner of each pair much shorter and more slender. Anal legs in male with fourth and fifth articles showing the longi- tudinal suleation dorsally but the tibial crest only faintly or not at all indicated. V entral “spines of first legs 0, 0, a a Spines of second and third legs, (tt24; of the fourth, 9 eee OF (ota; of the fifth, 7 2 of the sixth and sev enth, (os ae of the eighth to elev enth, (T3233; of the twelfth, } o- i 3353 of the thirteenth and penult, ¢ 0° is a z a: of the anal, 33 Coxae of anal legs armed laterally with a small pale spine. Length (oc) 6.5 mm.; anal leg 2.4 mm. long; antenna 1.6 mm. long. The praematurus form described is from West Cliff, Col. It is the specimen referred by Bollman to L. kochit Stuxberg, and is thus the type of coloradensis, the name having been proposed on the basis of a difference indicated by Bollman between this specimen and typical kochiv. Tyee Locauiry.— Colorado: West Cliff, T. D. A. Cockerell. Taken also in Colorado at Salina and Tolland (T. D. A. Cockerell), Durango (C. F. Baker), Manitou and Colorado Springs. Ipo NADABIUS MESECHINUS (Chamberlin). Plate 6, fig. 2 Lithobius mesechinus Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 158. DescripTion.— Brown. Head the same or but little darker, the anterior portion of frontal region pale. Antennae pale brown or yellowish. Prosternum like head, the prehensors lighter. Venter pale brown, the posterior plates darker. Legs pale brown or testa- ceous, the caudal pairs darker. Body robust, in types from slightly less than 6 to 6.4 times longer than width of tenth plate. Head considerably wider than long (40:37); widest caudally, narrowing slightly to eyes and then very strongly cephalad. Marginal breaks not conspicuous. Antennae short as usuat; composed of nineteen to twenty-one 330 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. articles of which the second is considerably longest and those distad of this short and nearly uniform, subeylindrie. Ocelli thirteen to sixteen in three series: e.g., 1 + 5,4,3; 1 +6, 5, 4. Single ocellus much largest. Others decreasing in size cephalad and ventrad; compactly aggregated. Prosternum 1.7 times wider than long. Distance between chitinous spots 2.2 times width at level of bottom of sinus; 4.6 times length of the short dental line. Teeth 2+ 2; very small, acute; inner larger than the outer. Spine small extending obliquely a little mesad of directly cephalad. Sinus rather large, strictly v-shaped. Sides ectad of spines broadly and widely rounded. First dorsal plate with sides moderately convex anteriorly and at caudal corners, but only little curved between these parts, a little con- verging caudad; about two times wider than long. Caudal margin of ninth plate straight or slightly extended at ends. Posterior angles of eleventh and thirteenth dorsal plates weakly produced. More caudal plates very finely roughened. Coxal pores of medium size or large, circular or a little transversely elongate: 3, 4, 4, 3. Spines of first legs, (REss or ci4e4ay; of thesecond, o-p°s doses; of the third io) Oredie eyeneee of the ninth, it of the tenth, i o eS 3 3; of the twelfth to penult, jy OL : : 5, the penult ak two claws; of the anal, (13:3, claw one. Last two pairs of coxae laterally armed. Claw of female gonopods short; tripartite; the lobes short, with the median one longest though not much so. Spines unusually short, wide; sides parallel from base to about middle of length at which the acutely acuminate apical division begins; inner spine somewhat smaller than the outer. Inner side of first article somewhat convex. Length 8-10 mm. A male 10 mm. long has antennae and anal legs 3.5 mm. long and tenth plate 1.57 mm. wide. Type Locanrry.— Oregon: Meacham, August, 1902. The male of this species is not known. FP NADABIUS IOWENSIS (Meinert). Plate 7, fig. 6; Plate 8, fig. 1. Lithobius iowensis Meinert, Proc. Amer. philos. soc., 1886, 23, p. 177. Boll- man, Proc. U.S. N. M., 1888, 11, p. 409. Chamberlin, Can. ent., 1911, 43, p. 70, 103. Gunthorp, Kans. univ. Sci. bull., 1913, 7, p. 164. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. Saal Lithobius trilobus Bollman, Proc. U. 8. N. M., 1888, 11, p. 409 (<7. in part). Lithobius bilabiatus Bollman (nee Wood), Proc. U. 8. N. M., 1887, 10, p. 256. Lithobius sexdentatus Kenyon, Can. ent., 1893, 25, p. 162. Lithobius bruneri Kenyon, Can. ent., 1893, 25, p. 162. Nadabius iowensis Chamberlin, Bull. M. C. Z., 1913, 57, p.62. Can. ent., 1914, 46, p. 302. Description.— Light yellowish or testaceous brown to dark brown, either uniform excepting the first plate, or with the caudal plates somewhat darker, the first plate in specimens in full color frequently clearly darker than the adjacent plates, approaching the head in color and sometimes fully as dark as the latter. Head from brown through reddish brown to deep brownish black or solid black. Antennae froma lighter brown to very deep brown, tips in the latter case paler. Pro- sternum and prehensors deep brown. Venter yellowish to brown with the most caudal plates always conspicuously darker brown. Legs yellowish to light brown, the caudal pairs usually more or less darker. Body varying from only 6.66 to 8 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth and twelfth dorsal plates to each other about as 41 :37 :40 :48 : 49 : 46. Head wider than long usually in ratio 22:21. Sides strongly rounded and converging from lateral breaks about corners to straight portion of caudal margin. Smooth and shining, punctae few. Hairs sparse, moderate. Antennae very short, reaching usually only to end of fifth segment. Articles twenty to twenty-five, twenty-three being not infrequent; subeylindrie or a little clavately widening; moderately short, with the second longest and the distal ones becoming very slender. Ocelli twelve to eighteen in four or five series: e.g., 1 + 4, 4, 4, 3; 1+ 4,4, 4,3, 1; 1+ 3, 4, 4,3, 2; 14+ 3,4,4,3,1; 1+ 4, 4, 3, 3,3. Single ocellus subvertically subelliptic, rather small though clearly larger than any of the seriate ocelli which are comparatively small and closely compacted. Prosternum between 1.45 and 1.5 times wider than long. Distance between chitinous spots 2.25— times width at level of bottom of sinus; 3.5-3.7 times the dental line. Teeth acute, well chitinized and usually dark, the innermost on each side often considerably smaller than the others; 3+ 3 or 4+ 4, very rarely only 2+ 2, and in one case observed as 6 + 4, the two extra teeth on the one side being separated by an interval from the others. Sinus wide, not acute at bottom. Spine stout at base, distally slender and usually somewhat curved mesad. 332 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. First dorsal plate 1.7— times wider than long; posterior corners well rounded, with the sides in front of them nearly straight and a little diverging cephalad to the anterior ends, in some appearing almost parallel. Plates smooth and shining or rarely obscurely some- what rugose. Hairs becoming more numerous and somewhat longer on the more caudal plates. Posterior corners of ninth, eleventh, and thirteenth plates not at all produced, being commonly obliquely excised, with line of excision making but small angle with the hori- zontal. Ventral plate clothed with rather numerous short hairs which become more dense on the caudal plates. Coxal pores rather small, especially the most proximal ones; cir- Culat ona oes 35 4, 4,4, 4; aN yp a Ste uaa Na aoe 0,0,2,2,1 0.0,2%1 0,0,22%1 0,022.1, 0, 0, 2,22 Spines of first legs, 0:0: : 21 0,0, ; 3, Jo, 0.0, 1,3, 21, 0,02, 2, 13 Or, 0, 0,2, 3,2 On ORON2N 252 o 20 0,0, Dae ONONS 22 of second to fifth, 0ra3; OL sixth, 0,0,2, 3,2 OF 0, 0,2, 3.23 of seventh and : OOS se }, 0, 3. 2, 2 0, 0, 3, 2, 2 eighth, j-¢-33-9; of ninth to elev enth, (5-35 OT 9-0-3379, being often of one formula on one side and of the other on the opposite; of the y 0, 0, 3, 1053522 ney a al twelfth, j-o:33°5 OF o7o-3'3'3; Of the thirteenth, ¢7@-3,3,2, the trochanter 5 ‘A OS el! spine being sometimes missing on one side; of the penult, (1353/3 claws three; of the anal, (33 p 0-1 a4 OF giesao Claw one! Last three pairs of coxae laterally armed. Anal legs short. In the female slender. In the male somewhat thicker than in the female; the third, fourth, and fifth joints dorsally longitudinally sulcate as usual, especially so the last two of these, the sulcus on the fifth joint being especially deep distad; fifth joint characterized by the usual low, flattened crest at the distal end; the crest highest distally, pilose with moderately long hairs which are curved at tips (Plate 8, fig. 1). Gonopods of male small, distally rounded, with sides also somewhat convex and the outer longer and more oblique than the inner; distally each gonopod bearing three or four long bristles. Claw of female gonopods of moderate size and curvature; usually strictly entire but not infrequently with a dorsal and a ventral denticle weakly indicated, the claw being evidently entire in a secondary way; acutely narrowing distad but apex usually narrowly rounded. Spines 2 + 2, or, occasionally, 3 + 3; of moderate length; when viewed at right angles to broad surface seen gradually to narrow distad with the sides slightly convex and the apex narrowly rounded; inner spine normally somewhat shorter than the outer but the difference appears 1 4 small accessory claw was observed on right leg in a male from Watersmeet, Mich., the claw of the left leg being single as usual. 906 CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE, 300 exaggerated because of the insertion of the inner spine farther proxi- mad. First joint not at all excavated on mesal side proximally. Length 12-16 mm. A female 12 mm. long has antenna and anal leg, exclusive of coxa, 4 mm. long and tenth plate 1.75— nm. wide. A female of same length may, however, have tenth plate only 1.5 mm. wide. A male 12.5 mm. long has tenth plate 1.75 mm. wide while a second one 13.5 mm. long has this plate of same width as in preceding. (Watersmeet, Mich., chiefly). Pseudomaturus.— Dorsum prevailingly light brown, with first plate either abruptly darker or not. Head darker brown. Antennae light brown to yellowish. Prosternum and prehensors similar to head or but little paler. Venter paler than dorsum, darker caudally as usual. Legs yellowish to light brown, the posterior pairs distally more pig- mented, somewhat light ferruginous or orange. Antennae as in maturus. Ocelligeegs t= 343; 3, 2301 4-4 353; 15 Ve 4.3.38; 2; 14, 4, 3, 2,1. Thus averaging fewer than in the ailille and more commonly in four series. Prosternum with teeth 3 + 3 or 2+ 2; when 3 + 3 the innermost often much smaller and close to side of sinus. Coxal pores very small; most frequently: 3, 4, 4, 3; also 3, 3, 3, 3 to 4, 4, 4, 3. Claw of female gonopods smaller and more acute and paler than in adult. Spines essentially as in maturus but distally more acute and the inner one proportionately somewhat smaller. Bristles fewer. Gonopods of male small and rounded; already bearing the three bristles. Spines of first legs Soto to gots of seventh may be, j'yo'3°2 oF ob a 35 of anal, } me ee aa Olausinas sat ch ae as in maturus. Last two pairs of coxae laterally armed or last three pairs as in maturus. Anal legs of male modified as in the maturus but the crest somewhat smaller and its hairs fewer. Length 10.5-13 mm. A female 11 mm. long has antenna 3.5 mm. and anal leg near 3 mm. long. (Mongona, Iowa). Praematurus.— Dorsum yellowish to testaceous and brown. Head brown, always darker than dorsum of body. Antennae light brown proximally, paler distally. Prosternum and prehensors testaceous to brown, usually distinetly darker than the venter which is yellow to dilute brown, tenth caudal plates darker as usual. Legs like venter, but posterior pairs distally light, subferruginous or orange. Antennae nearly as in older stages. Rate 9 ras wr 0, 304 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli nine to fourteen in three to five series, but mostly in three or four: seg 3; 35/250 loans 1-4 oon ones T= ara eee ele All ocelli deeply pigmented. Prosternal teeth 2 + 2, equal or very nearly so, rather pale. Sinus wide, subacute at bottom. 1.47 times wider than long. Distance between chitinous spots aie times width at sinus, 3.3 times dental line. Coxal pores very small: 2, 3, 3,2; 3, 3, 3, 3; 3, 4,4, 3. Claw of female gonopods small, entire, slender and acute, pale. Spines 2 + 2, acutely acuminate, the inner much shorter and more slender than the outer. Distal article mostly with but one bristle; the second with two or three; and the first with three. ONO Maer ONO asl * Pope of first legs, o0121 OF 0-00-21; Of the second and third, Ds OOP ees 0. 2, 2, 2 ° “TOF Tos21; Of the fourth to seventh, eos; of the eighth, 2 Drees, 050) 34242, iO the ninth to eleventh, ¢ a: a 33-9, Of the twelfth, 9-9-3-3'5; of 2 . nee aoa a 83 of the penult, ( u 33-3; of the anal, a) - a1 OF oy. Only the last pair of coxae laterally armed. Lea 9.5-11 mm. A female 10.5 mm. long has antenna 4 mm. and anal leg 3+ mm. long and tenth plate 1.3 mm. wide. ‘The maxi- mum length overlaps the minimum for the pseudomaturus. (Mon- gona, Ia.) Immaturus.— Dorsum pale brown, most plates showing three paler longitudinal lines, of violaceous tinge throughout. Head and posterior plates darker. Prosternum and prehensors similar to head. Antennae light brown, paler distally. Venter and legs pale, the posterior pairs usually more brightly pigmented. Antennae in most composed of nineteen articles of which those distad of the third are mostly very short, though varying considerably in length, as usual in yet developing antennae. Ocelli seven to ten in two or three series: e.g., 1 + 4, 2; 1+ 3, 3; 1+ 3, 3, 2; 1+4, 3, 2. First ocellus of top series as large as or Jarger than the single ocellus. Prosternal teeth 2 + 2, equal, pale. Sinus somewhat more acute at bottom than in maturus. Coxal pores very small, the most proximal one on each coxa espe- cially minute: 2, 2, 2,2; 2, 3, 3, 2. In the smaller specimens short and pale but the three articles al- ready present; of these the distal is short and simply rounded with no indication of a claw and is glabrous; second article with one bristle and first one with one longer bristle in addition to the minute pale chitinous point representing the spine; sternite with two bristles. In larger specimens the claw is present, being small, slender, and acute; o\os\ . Pee tr |= ct —* Blo B S\os|o EB Se =) CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 300 distal article with one bristle, second mostly two and first usually four while sternite may bear as many as ten. Spines of first legs, (0:0 71to “ene zt; of thesecond, ooo ito OCT of the third, (+t: a to a oon oe of the fourth and fifth, 9°9°9- 2 ; to oor 0 Vs 2 and pee: of ne ao OOO tO gora-373; Of the seventh to ninth, (137 to nysge; of the tenth, 4 O12 b Otte 21 tO joes; of the eleventh, Poe o a OF O° 5 3 35; of the twelft] ; 1 oF? too 4, 34 3 3. of the thirteenth, ¢ 0: So OF : 1 - 333 of the penult, 7 aes 373; of the anal, SHORE INO, rs to aay Last pair of coxae alone laterally armed. Dea ath 5.6-9 mm. In specimen 5.6 mm. long widths of head and of first, third, eighth, tenth and twelfth plates are to each other as 20.5 : 18.5 : 20 : 23:24:22. (Earliest stage, Watersmeet, Mich.; later, Mongona, Iowa.) Type Locauiry.— Iowa: McGregor. Found also in Iowa at McGregor (Davis), Mongona, Tama, Boone, De Witt. Nebraska: Omaha, Fremont. Kansas: Anderson, Bourbon, Coffey, Cowley, Douglas, Franklin, Graham, Jefferson, Labette, Linn, Pottawatomie, Reno, Sumner, and Trego counties (fide Gunthorp). Illinois: Rock Island, Franklin Grove, ea Peoria, East Peoria. Indiana: Bloomington (Bollman coll.), L Fayette, Richmond, Greencastle, Salem, New Providence, Wy Jaceiatts (fide Bollman). Michigan: Watersmeet, Saunders, Menominee, Ludington. Wisconsin: Beloit, Haugen, Marinette. Colorado: Manitou (Hayden Exp. 1873, Camp). Wyoming: Cheyenne. Variation in this species from place to place is large, the place- modes for size and other characters often differing considerably. In some places the ventral spines of the anal legs would seem to be the more frequently 1, 3, 2, 1 (some places in Illinois and Wisconsin) whereas in others the formula is almost exclusively 1, 3,3, 1. Segrega- tion of a distinct local variety, however, seems nowhere to have been clearly effected. Specimens cannot be referred to the different stages of postembryonal development on the basis of size because of frequent and wide overlapping. The range in size of the ¢mmaturus, as limited in the usual way, is very large as it is also, e.g., in the spining of the legs as noted above. An examination of specimens in several vials from Bloomington, Ind., (Coll. U.S. N. M.) labeled by Bollman as L. trilobus shows them for the most part to be referable to the present species, some speci- mens, however, are 7’. pullus. As these specimens do not very well conform to the original description, it may be supposed either that they were incorrectly referred or that there has been a transfer of 336 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. labels or an incorrect reference by number, as many of Mr. Bollman’s specimens were indicated only by numbers referring to his personal catalogue. NADABIUS EIGENMANNI (Bollman). Plate 7, fig. 5. Lithobius eigenmanni Bollman, Proc. U. 8. N. M., 1888, 10, p. 625. Description.— Dorsum testaceous to brown. Head in long preserved specimens seems also to be brown with frontal region paler, of orange tinge. Antennae yellowish, darker proximally. Proster- num like head, or a little paler. Venter pale, yellowish or testaceous, with the caudal plates more deeply pigmented as usual, brown of orange tinge. Most legs pale, the posterior pairs bright yellow or orange. Body from 7.25 (in type specimen) to 7.75 or 8 times longer than width of tenth plate, being a robust form. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other about as 28 : 26 : 27 : 31 : 32 : 30.5 to considerably wider. Head from equal in length and breadth; of usual general shape; caudal margin weakly and widely incurved. Antennae short; articles short and of usual arrangement. Hairs long. Each eye consisting of from eight to eighteen ocelli arranged in two or three to four and occasionally five series: e.g., 1 + 4, 3; 1+ 4, 4, 3,1+ 4, 3, 3; 1+ 4, 3, 1; 144, 3, 2; 1 +4, 4, 3; 1; 1+ 4, 4, 4,3; 1+ 3,4,5,4,1. Single ocellus often small, those of most dorsal row, or at least most caudal one, the largest of all, as well as most caudal of second row, commonly exceeding it in size; or, in largest specimens, the single ocellus is distinctly largest. Ocelli com- monly not crowded, deeply pigmented, and rather irregular in form and size. Prosternum 1.5 (type) or mostly near 1.6 times wider than long. Distance between chitinous spots 2.4, or a little less, times the width at level of bottom of sinus; 3.5 to 2.8+ times the dental line. Mesal incision of moderate width, its sides recurved. Teeth equal in size or nearly so, rather small, line of apices a little recurved. Spine not very close to tooth; much stouter at base than the bristles, narrowing rapidly distad and above level of apex of tooth slender and bristle-like. First dorsal plate 1.8 times wider than long; sides strongly diverg- ing cephalad. Dorsal plates rather strongly arched. Smooth. Posterior inner plates with corners obliquely truncate. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 33 Coxal pores circular: 2, 3,38,2; 2,3,3,3; 2,4,4,3; 3,4, 4, 3; 4,5; 5, 9 a OW yal On 0h 25:2501 0, 0, 2, 2, 1 ee of first legs, o-o:i34 and 0:0: I om (pseudomaturus) to 9: 0-334 0, 0, 3,2, 1 f 1, 2,2 0, 0, 22,2 and 0-0-1.3-2 (maturus); of the second, oo 13-1 and 9:9: 1.3-9 (pseudomat- 0, 0, 2, 2, 2 0, 0, 2 0,0, 2, 2, 2 w of the third to sixth, 99-735 0,0, ; of the seventh to eleventh, 0: 05 0, 8, 1, 2 0, 0, 3,2, 2 : g.3 OF 0,3, 3, 2 the coxa ae eect 1, 0,3,1,1 33-9; Of the penult, 9 73°35, 0 wee a claw single. Last two urus), to fysas and pos >in fully grown speci imen 9723-9; Of the twelfth, OS 3 armed; of the thirteenth, ¢ claws three; of the anal, 7 pairs of coxae laterally armed. Anal legs in male considerably inflated, the fourth joint most noticeably so. Third, fourth, and fifth joints longitudinally fur- rowed above, the fourth the most distinctly so. Fourth joint on mesodorsal side toward distal end subdensely clothed with long hairs which at the tips are curved somewhat caudad; joint elsewhere only sparsely clothed with short hairs; fifth joint at distal end above with the usual crest which is low and often inconspicuous excepting in fully grown specimens. Penult legs a little less inflated; similarly suleate; the fourth joint without the special growth of hair. Gonopods of male with outer side more oblique than the mesal; distally transversely straight or nearly so; thin, edge-like, distad. Claw of female gonopods short and proportionately wide; tripar- tite, with the median lobe little longest and the outer often somewhat smaller than the inner. Basal spines short and very broad, with sides subparallel or somewhat diverging distad to the acuminate apical portion which is rather short and acute with its sides more or less incurved; inner spine considerably and constantly smaller than the outer (Plate 7, fig. 5). Mesal edge of first article straight, as usual in the genus. Length from 7 to 12.5 mm. A male 9 mm. long has antennae 3.25 mm. and anal leg 3 mm. long with tenth plate 1.17 mm. wide. Type Locauity.— British Columbia: Glacier (C. H. Eigenmann). Taken also in British Columbia at Kaslo. Oregon: Portland, August, 1902. The description above is based in part upon the types of the species; but these are not fully developed and are in bad condition so that some features could not be made out with certainty. The main part of the account was consequently drawn from specimens collected at Portland, Oregon, and Kaslo, B. C. The species in the region about Kaslo varies much, but intergrades seem too frequent for the separation of a distinct variety. € 2 Ps 0, = 1 0, 1 308 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. NADABIUS HOLZINGERI (Bollman). Lithobius holzingert Bollman, Entomol. Amer., 1887, 3, p. 83. Chamberlin, Can. ent., 1911, 48, p. 104. DerEscripTIon.— Brown to chestnut, the head darker. Prosternum nearly like head, the prehensors somewhat paler. Venter paler brown, the caudal plates darker, somewhat chestnut. Legs yellowish brown or brown, the caudal pairs, as usual, clearly darker. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 44 : 44 : 45 : 52 : 54 : 52. Head wider than long in ratio 22:19. Widest at marginal breaks, which are pronounced. Caudal margin straight. A transverse impression a little in front of caudal marginal thickening, the impres- sion deepest at middle. Antennae short. Articles twenty to twenty-three (twenty-eight seq. Bollman); short. Ocelli, in types examined, sixteen or seventeen in four or five series: g, 1+ 4,4,4,38; 1+3,4,4,4; 1+ 3,4, 3, 3,3. Single ocellus elliptic or oval, large. Prosternum 1.5 or 1.6 times wider than long. Distance between chitinous spots 2.25-2.5 times the width at level of bottom of sinus; 3.4-3.5 times length of dental line. First dorsal plate with sides rounded and diverging cephalad; 1.76 times wider than long. Posterior angles of eleventh and thirteenth dorsal plates with a slight tendency toward production as usual. Coxal pores circular, moderately small: 3, 4, 4, 4; 4, 5, 5, 4; D050; 0. Spines of first legs, & 03373; of the second to fourth, 9 0 33; of the fifth to tenth, 3 220 ‘of the elev enth and twelfth, (o73 a2 5; of the thirteenth, (13:35; of the penult, } eres claws three, the two acces- sory claws being large and distinct; of the anal, §:°3:3:4, claw one, long. Last pair of coxae laterally armed. Anal legs of male short and moderately thickened; fourth and fifth articles dorsally rather strongly longitudinally suleate and ‘the fifth joint having at its distal end a low, thin crest in usual position, with its dorsal line convex and highest distad. Penult legs similarly some- what thickened and dorsally furrowed, but with no crest. Gonopod of male subcuneate, distally subtruncate; bearing three bristles. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 339 Claw of female gonopods short, wide, tripartite. Spines short and stout, subequal. Length 10 to 14mm. A male 14 mm. long has antenna 5.5 and anal leg near 4.5 mm. long. Type Locatiry.— Minnesota: Winona. (J. M. Holzinger). The description above is drawn from three of the original types (7), the statement as to the female gonopods, however, being taken from the original description. Bollman gives length of males as from 16 to 21 mm., but this is much too large as is frequent with Bollman’s measurements,— possibly the anal legs are included in them. Poxasius Chamberlin. Poabius Chamberlin (nee Koch), Ann. Ent. soc. Amer., 1912, 5, p. 153. Pokabius Chamberlin, Can. ent., 1912, 44, p. 316. Head with lateral marginal interruptions distinct (Pokabius sens. str.; Anobius) or sometimes scarcely evident (Lophobius). Antennae short or very short; articles normally twenty, occasion- ally varying a little from this in individual cases. Eye composed of seriate ocelli which are from six to twenty-five in number and are arranged in from two to six series. Single ocellus enlarged. Prosternal teeth 2 + 2; line of apices more or less recurved. Spines slender, distally bristle-like. Posterior angles of none of the dorsal plates.or those of the ninth, eleventh, and thirteenth produced. Coxal pores uniseriate, circular. Gonopods of the male uniarticulate. Claw of female gonopods normally tripartite, occasionally only bipartite, the median lobe commonly not much longest. Basal spines 2+ 2, usually intermediate in length and stoutness, but varying in both directions. First article of gonopods not excavated proximally. Tarsi of all legs divided. Anal legs of the male with fourth joint characterized by a dorsal longitudinal ridge more or less conspicuously elevated into a lobe at or near the proximal end, and the third joint with a dorsal ridge usually elevated at the distal end (Pokabius sens. str.; Anobius, Plate 8, fig. 4; Plate 9, fig. 1, 3, 4; Plate 10, fig. 1, 3; and Plate 11, fig. 1, 2); or in some species the ridge of fourth joint less developed and more or less elevated at distal end or in some cases practically absent (Lophobius). Penult legs never with any special lobes (excepting in P. (Anobius) centurio). COMPARATIVE ZOOLOGY. BULLETIN: MUSEUM OF 040) NS A, Meat va } Fic. 8.— Distribution of Pokabius. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 341 From one to three last pairs, or rarely none, of coxae laterally armed. Fifth joint of all legs between first and twelfth with two dorsal spines. Dorsal spines of anal legs in female varying from 1, 0, 2, 1, 0 and 1, 0, 3, 1,0, the most common formula, to 1, 0, 4, 1, 0, and in the male fron als 0, PoalrOntoule 0. on lb uOss ventralOnle 3 2)(0:0r 0; 1.3, 2, 1 claw one (or in socius only). Dorsal spines of penult legs 1, 0, 3, 1, 1 or rarely 1,0, 2,1,1; ventral, 0,0, 2,3, 2 to0, 1,3,3, 1 and0, 1, y Berg claws two or een Dorsal spines of thirteenth ne 11053; 1,1 or rarely (utahensis) 1, 0, 2, 1, 1; ventral, 0,1, 3, 3, 2 or 0, 0, 3, 3, 2, rarely only 0, 0, 2, 3, 1 (utahensis). eee spines of pve legs ivOcor ls, Jeonrarelyols O72 ai ileto ly OS. 18 2: penal) OZ ee ONO 3, 332. Worsaliispines of “first legs, 0; 0, 2,25 1 to 0,0; 3, 2; 2; ventral 0:0, 1 2,1 to/0; 0; 2,3, 2. Length 6.5 to 17.5 mm. but mostly from 9 to 15 mm. Type.— P. verdescens Chamberlin. Variation among the species embraced in the genus as here defined is in many features large; but there seems no reason to doubt its entire naturalness excepting only the portion designated as the sub- genus Lophobius. The latter may have to be withdrawn and possibly subdivided when the region over which it ranges is more thoroughly explored and the species now recognized and others yet to be found are better known in both sexes. Pokabius is a comparatively large genus having its center in the southwest, ranging up the west coast to Washington and British Columbia and up the Mississippi valley to southern Minnesota and Wisconsin (Figure 8). In the upper Mississippi valley it is represented chiefly by P. bilabiatus a very abundant spécies. The distribution of the subgenus Lophobius is separately shown (Figure 9). The species of Lophobius are found under stones over foothills and in similar open, dryer and less wooded areas in largely semi-desert regions. The key does not include P. carinipes (Daday), which is insuffi- ciently known. (Cf. p. 367). Key to Subgenera and Species of Pokabius. a. Posterior angles of ninth, eleventh, and thirteenth dorsal plates WEOUUICCEE te Sees aos heey «ez Anobius, subgen. nov. b. Dorsal spines of twelfth legs 1(0), 0, 3, 1, 2; head wider than long; penult leg of o with a lobe on the fourth joint; length LO Prim Onslesshytaes ean eae ceuete P. centurio (Chamberlin). 1 P. (Anobius) centurio (Chamberlin). Type. 342 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. bb. Dorsal spines of twelfth legs 1, 0, 3, 1, 1; head as long as wide; penult leg in o with no such lobe; length 13-16 mm. P. gilae, sp. nov. aa. Posterior angles of none of the dorsal plates produced. b. Anal leg of o with a dorsal ridge on fourth joint elevated into a conspicuous lobe at or near the proximal end; marginal interruptions of head conspicuous..... . Pokabius sens. str. c. Dorsal spines of anal legs of o 1, 0, 4, 1, 0 to 1, 0, 6, 1, 0; of (2 .1,10;,4,, 1,0: d. Body 8.75-9.5 times longer than width of tenth plate; ventral spines of thirteenth legs usually 0, 1, 3, 3, 2. P. bilabiatus (Wood). dd. Body only 6.7-8 times longer than width of tenth plate; ventral spines of thirteenth legs usually 0, 0, 3, 3, 2. P. bilabiatus verdescens Chamberlin. ce. Dorsal spines of anal legs 1, 0, 3, 1, 0 or 1, 0, 2, 1, 0 in both o' and 9. d. None of the coxae laterally armed. Ventral spines of penultiilessi0;e 33) lin tana. P. disantus, sp. nov. dd. One or more pairs of posterior coxae laterally armed; ventral spines of penult legs 0, 1, 3, 3, 2. e. Only the last pair of coxae laterally armed. f. Distance between chitinous spots 3.5 times the dental line and near 2.33 times width at sinus; anal leg of o (Plate 10, fig. 3)....P. utahensis (Chamberlin). ff. Distance between chitinous spots near 4.4 times dental line and 2.9 times width at sinus; anal leg of on @blate te ches?) 02. wae P. sokovus (Chamberlin). ee. Last 2 or 3 pairs of coxae laterally armed. f. Anal legs of co" with dorsal spines normally 1, 0, 2, 1, 0 and lobe of fourth joint at extreme proximal end (Plate, Os itie walters: P. clavigerens (Chamberlin). ff. Anal leg of both co and 2 with dorsal spines 1, 0, 3, 1, 0 and lobe of fourth joint cephalad of extreme proxi- mal end. g- Dorsal spines of first legs normally 0, 0, 3, 2, 2; claw of 2 gonopods subtruneate with lobes but weakly indicated; anal leg of (Plate 9, fig. 4). P. pitophilus (Chamberlin). gg. Dorsal spines of first legs normally 0, 0, 2, 2, 2; claw of 2 gonopods distally acute with 3 lobes CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 343 distinct and the median much longest; anal leg of o& (Plate 8, fig. 4). .P. tginus (Chamberlin). bb. Anal leg of & with dorsal ridge elevated, if at all, at distal end, never with lobe at proximal end, sometimes not evident; marginal interruptions of head usually weak or scarcely THMEUININES Ge, ena A SS cue eae ol Lophobius, subgen. nov. ce. All of anterior legs, or all but first 1 or 2 pairs, with third joint bearing 2 ventral spines. b. Ventral spines of penult legs 0, 1, 3, 3, 2. e Annales wath: Zuclawseind sion ioss P. socius (Chamberlin). cc. Anal leg with the claw single. d. Dorsal spines of twelfth legs 1, 0, 3, 1, 1. e. Ventral spines of anal legs 0, 1, 3, 2,0; of the twelfth and thirteenth 0, 0, 3, 3, 2. P. collium (Chamberlin). ce. Ventral spines of anal legs 0,1, 3, 2,1; of twelfth and thirteenth, 0, 0, 2,3, 2. .P. pungonius (Chamberlin). dd. Dorsal spines of twelfth legs 1, 0, 3, 2, 2 or 1, 0, 3,.1, 2. e. Ventral spines of anal legs 0,1,3,2,1; head a little longer than wide.......... P. arizonae, sp. nov. ee. Ventral spines of anal legs normally 0, 1, 3, 2,0; head widerh than longs). 20 eed ac P. helenae, sp. nov. bb. Ventral spines of penult legs 0, 1, 3,3, 1 (lobe at distal end of fourth joint of anal leg in o strongly developed). P. castellopes (Chamberlin). aa. First 7 pairs of legs with the third joint bearing but a single TILT ANS [ITI © Seer DN enn aN ye lou Rol ar P. eremus, sp. nov. ANOBIUS, subgen. nov. PoKABIUS CENTURIO (Chamberlin). Plate 10, fig. 1, 2. Lithobius centurio Chamberlin, Proc. Acad. nat. sci. Phil., 1905, 56, p. 651. Poabius nankus Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 153, pl. 12, fig. 4. Descrirtion.— From light orange-brown to dark brown in speci- mens in full color; with typically a darker median longitudinal band more or less evident, and with some of the major plates in some degree darkened along caudal border. Head nearly same as dorsum or a little lighter, paler in front of suture and darkened over middle of 344. BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. caudal portion. Antennae light brown. Venter a lighter brown than dorsum, the posterior sternites darkest, all excepting the latter with a paler central circular area. Anterior legs yellowish, the pos- terior pairs becoming brown, uniform. Body very robust, being typically only about 5.5 times longer than width of tenth plate. Twelfth plate as wide as, or wider than, the tenth, and the tenth wider than the eighth. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 49 : 46: olFOLe 6870! Head considerably wider than long (about 10:9). Caudal margin widely and moderately incurved. Width not much varying in region between eyes and a level a little caudad of marginal breaks, which are weak and very oblique, the sides caudad of this region strongly con- verging to caudal corners. A deep and conspicuous but rather short transverse sulcus parallel with and a little in front of median portion of frontal suture. A shallow curving longitudinal furrow each side of middle in front of caudal border. Finely punctate and but slightly weakly roughened. Antennae short; articles normally twenty but varying from nine- teen to twenty-two, moderate, decreasing distad. Ocelli fifteen to twenty-two in usually four series: e. g., 1+ 5, AA Al Se On, 45 25 ise O50; 45°35) tO Oo, 48 Ocellrclosely: curved and in consequence commonly hexagonal. Single ocellus large, elliptic, lying obliquely beneath and contiguous with caudal ends of series. Prosternum about 1.6 times as wide as long. Width between chitin- ous spots 2.75, or somewhat more, times as great as width at level of bottom of sinus; 5.2 times as great as dental line. Teeth 2+ 2; small, acute, with apical portion sometimes extended into a bristle- like tip which is easily broken off. Spine as usual, distally bristle-like, borne on a small tubercle ectad of outer tooth, about three somewhat similar bristles ectad of the spine. All dorsal plates finely roughened but the caudal ones more dis- tinctly so. Most plates with a median furrow, sometimes replaced by two, and on each side one or two approximate ones diverging from it caudad. Ninth, eleventh, and thirteenth dorsal plates with the caudal angles strongly produced, but apices of processes a little rounded rather than acute. Posterior angles of seventh plate very slightly extended caudad, the mesal side of extended portion long and making but slight angle with median portion of the margin. Caudal corners of sixth plate a little obliquely excised. Lateral and caudal margins of CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE, 345 plates bearing numerous very short, acutely tipped, hairs upon minute serratures. Ventral plates with two main wide transverse furrows between which there may be finer impressed lines. The last plate with two sharply impressed longitudinal sulci converging from the anterior margin and ending abruptly at about middle of the plate, where their ends are united by a less strongly impressed transverse sulcus; the preceding plate similarly but rather less conspicuously marked. Last ventral plate with adjacent coxae and succeeding segment subdensely pilose, the anterior plates being only very sparsely clothed. Coxal pores small and circular, with those of each coxa widely separated: 2,3,3,2; 3, 3, 3,3. Tee) 0, 0,3, 2 ONON See Spines of the first legs, (oias; of the second to tenth » 0, 0, 2, 3,23 9 of the eleventh, (::3:3°3; of the twelfth, 1.221; of the thirteenth, 0 t 23 geas; of, the penul same as of thirteenth, claws three; of the anal, r 210 1, 0, 2, 1,0 . a a (H) or g1321 (2, also co’), claw one. Last pair of coxae laterally armed. Anal legs of male short and rather slender; the third joint abruptly produced at extreme distal end subdorsally into a high lobe, the caudal face of which is vertical or at right angles to axis of joint and the anterior face but little slanting, the lobe bearing two (or one) of the dorsal spines; fourth joint produced into a larger conspicuous lobe at proximal end and contiguous with the lobe of the preceding joint, the lobe somewhat plate-like, continuing caudad as a lower ridge along mesal side of dorsal surface (Plate 10, fig. 1). In the penult legs of male the prefemur is elevated into a low, inconspicuous lobe at distal end, while the femur or fourth joint bears at its distal end a conspicu- ous lobe which is rather narrow from side to side; this lobe projects conspicuously dorsocaudad over plane of the articulation with tibia, both its distal and its proximal surfaces slanting in this direction but the proximal surface at an angle with the distal so that the lobe decreases distad in thickness as well as in width. Gonopods of male small, low conical, distally rounded, pale, extended caudoectad. Claw of female gonopods tripartite; median lobe largest though blunt and comparatively short. Spines moderate in length and stoutness, narrowing a little from base to short apical portion, with sides a little incurved; inner spine shorter and a little more slender than the outer (Plate 10, fig. 2). Length 13-15 mm. A male 14.25 mm. long has anal leg 5 mm. long and tenth plate 2.6 mm. wide. 346 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Type Locatiry.— New Mexico: Las Vegas. Found also in New Mexico at Las Valles (T. D. A. Cockerell), Santa Fe (H. S. Barber). This is the only member of the genus thus far noted to have the penult, as well as the anal legs of the male pronouncedly modified. POKABIUS GILAE, sp. nov. Plate 11, fig.” 1: Description.— Dorsum light brown or testaceous, with anterior plates of more greyish cast; posterior borders of major plates in middle and anterior region with a blackish caudal border. Head somewhat ferruginous, darker caudad of suture and especially in a median black- ish spot contiguous with it. Antennae brown proximally, yellow at tips. Prosternum and prehensors light testaceous. Venter dilute greyish brown. Legs dilute brown, the caudal pairs darker brown the tarsi and mesal surface of anal pair lighter. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 60 : 51 : 58 : 70 : 76 : 75.5. Head subcordate, but with caudal margin nearly straight. Equal in length and width. Marginal breaks weak. Antennae as usual. Ocelli fourteen to sixteen in four series in type: 1 + 2, 5, 4, 2; 1-65, 593},2. Prosternal teeth small. Median sinus small, shallow, widely v- shaped. Spine slender, straight and long. Prosternum 1.45 times wider than long. Distance between chitinous spots 2.8-+ times the width at level of sinus; 4.6 times the dental line. First dorsal plate very moderately narrowed caudad; 1.64 times wider than long. Posterior angles of eleventh and thirteenth, and of ninth on one side rather weakly produced (probably both normally produced in ninth). Coxal pores small: 2, 2, 2, 2; 2, 3, 3, 2 0, 0, 221 oe 0, 0, 2, 2, 1 Spines of first legs, 0-0; T21 OT 00,1315 of the second, 9:9: Or Oris Ny PP Pp pas; of the third and fourth, oss; of the fifth to sleventhe ; ao 3 nai, of the twelfth, §? Bul OL the thirteenth, } css; of the penult, * a4 “43, Claws three; of the anal, a i33-) claw one. Last three pairs of coxae laterally armed. Penult legs of male without special lobes or other modifications. Anal legs of male with the third and fourth joints dorsally ridged and CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 347 produced into lobes at their contiguous ends in the general way usual in the genus, the third bearing a longitudinal dorsal ridge elevated distally into a lobe which at base is contiguous with a lobe at proximal end of a longitudinal ridge on fourth joint (Plate 11, fig. 1). Length between 9 and 10 mm. in type. Type.— M. C. Z., No. 184. Arizona: Thatcher. Known only from the male. Collected under willow leaves at edge of Gila River, April, 1913. POKABIUS BILABIATUS (Wood). Plate 8, fig. 2. Lithobius bilabiatus Wood, Proc. Acad. nat. sci. Phil., 1867, p. 130. Stuxberg, Ofvers. Vet.-akad. Férhandl., 1875, 32, no. 3, p. 18, 31, 32. Chamber- lin, Can. ent., 1911, 43, p. 103. Archilithobius bilabiatus Stuxberg, Proce. Cal. acad. sci:, 1876, 7, p. 138, 139. Lithobius tuber Bollman, Proc. U.S. N. M., 1887, 10, p. 256. Lithobius malterris Kenyon, Can. ent., 1893, 25, p. 161. Poabius bilabiatus Chamberlin, Ann. Ent. soc. Amer., 1912, 6, p. 153. Kan. univ. Sci. bull., 1913, 7, p. 167. Descrietion.— Dorsum from light brown or testaceous to dark brown in specimens in full color, with the posterior plates usually somewhat darker than the others, while the plates generally may be paler along caudal border and in a median longitudinal line; first dorsal plate either the same as adjacent ones or it may be, in fully colored specimens, decidedly darker and not much paler than the head. Head from brown of a dilute chestnut tinge to deep brownish black. Antennae like the head or nearly so, tip rufous. Prosternum in paler specimens nearly like the head, in darker ones paler than head, dark brown but not blackish; prehensors may be a little paler than prosternum. Venter testaceous to brown of lighter shade than dorsum, caudal plates darker as usual. Legs testaceous to clear brown when in full color; posterior pairs much darker, usually much like head in color, their tarsi rufous. Body typically long and slender being from 8.75 to 9.5 times longer than width of tenth plate from which it is only moderately narrowed cephalad. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 47 : 44 : 46 : 52:53:51. Head considerably wider than long (e.g., 47 : 43). Caudal margin widely but only weakly incurved, laterally widely curving to and 348 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. about the corners. Sides between posterior corners and marginal breaks usually substraight. On caudal portion of head a transverse semicircular impression is usually conspicuous, a distinctly impressed transverse sulcus on each side being also evident in front of caudal border. Finely and rather densely punctate. Antennae very short, reaching only to fourth and fifth body-seg- ments. Moderately attenuated but not very slender distally. Com- posed of the usual twenty, or occasionally of twenty-one or twenty- three articles, which are short but vary considerably in different speci- mens. Ocelli thirteen to seventeen in four, or less commonly in three, series: e.g..1 + 5,4,3; 1+ 4, 5, 3, 1, a frequent arrangement; 1 + 3, 4, 3, 1; 1+ 3,4,4,1; 14+ 3,4,3,2; 1+ 4,4,3,1; 1+ 4,4,3,2; 1+ 45,4, 2; 1+ 5, 4, 3, 2; 1+ 5, 4, 4, 3. Single ocellus moderate in size. Series usually distinct and regular. Ocelli of top row not much larger, the decrease ventrad being gradual and moderate. Prosternum between 1.57 and 1.7, times wider than long. Distance between chitinous spots in an average specimen measured 2.4 times the width at level of bottom of median sinus and 4 times the dental line. Teeth 2+ 2; line of bases forming a weakly reentrant angle and line of apices straight or nearly so, the inner tooth being usually larger; teeth acute. Spine stout proximally, slender distally and usually somewhat curved; attached a little caudad of margin. First dorsal plate about 1.7 times wider than long; surface finely punctate and roughened. Major plates mostly showing the short transverse sulcus toward each lateral margin near caudal third and usually one or two dot-like impressions on each side farther cephalad. Under lens showing a fine roughening which is somewhat more pro- nounced on caudal ones. Posterior corners of all plates distinctly and more or less uniformly simply rounded. Ventral plates with three longitudinal sulci deeply and distinctly impressed, a submedian transverse sulcus also evident on most plates. The median longitudinal sulcus less distinct on the anterior plates. On last two plates a v-shaped impression with apex caudad, this most distinct on the ultimate one, the two lateral longitudinal sulci being evident in addition, at least on the penult plate. Coxal pores moderate, the most proximal one on each coxa often very small, all circular or subcircular: 3, 5, 4,3; 3, 5, 5, 3; 3, 5, 5, 4; 3,0, 45 444 5 4 An OND, os 40s Ones 4, Oy Oy Om Ay ON OA) 410 Oa Os Ooo. Os OD. 4 OO. 6, Be oe a 0, 0, 0; 0525.25.22 PAI) Spines of first legs, poy oa to 0 0, 1, 3.23 of the second, o: 132 OF O05 ane ONONSaoca2 v-0-23-3; Of the third and fourth, ?: Hees OP ate 33; of fifth to tenth, CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 349 2 0,.0, 3, 2, 2 Us OD) Qe , By 2 oy ale 3-9, Of the eleventh, (:? oe or o0-2-3-9; Of the twelfth, 9-933 1, Ons 1e thirteenth and penult, ; ae? 4 those of penult less often 9° 0: 3: 3 s ited ONG: Ik OF 0; 1,0 three; of the anal, in the male, §-3'5') o:a-3-2.0 oF rarely 0,0, 3, 2, th claw Piso but in the female nearly always, i. v 4 ca rarely Oa: a claw one. Last three or two pairs of coxae later ally armed. In female the posterior legs are short and slender, wholly without special lobes or furrows. In the male the third article of anal legs is produced at distal end mesad or dorsomesad into a large lobe the caudomesal side of which is convexly rounded and bears above five or six, or rarely in small specimens (pseudomaturus) but four, stout spines of which the most mesal is largest, and immediately ectad of these spines a subconic or rounded process which projects dorsad or dorsocaudad and bears distally a number of very short acute bristles; ectad of hinge on dorsal side a single spine in usual position. Fourth joint at a little distance caudad of its anterior end produced meso- dorsad into a very large and conspicuous flattened lobe which has its anterior side nearly straight and transverse to axis of joint; caudad the lobe lowers obliquely to a low ridge which at distal end of joint is extended into a small, conical, distally strongly rounded, process bearing several aculeate hairs, this process being larger than the simi- lar one on third joint; lobe densely clothed at its rounded apex with short aculeate bristles (cf. var. verdescens Plate 8, fig. 3). Penult legs without lobes but the third, fourth, and fifth articles, the fourth most distinctly, dorsally longitudinally sulcate. Gonopods of male small and wart-like; commonly distally sub- truncate; bearing usually a robust bristle inserted in a distinct in- dentation and two smaller bristles. Claw of female gonopods short and relatively wide; but little curved; tripartite, with the median lobe largest, acute; lateral divi- sions small, or very small, and tooth-like, occasionally the inner one absent and rarely the claw appearing subentire while in other cases it may show two short, blunt and nearly equal divisions. Basal spines of moderate size, with the outer considerably larger than the inner: narrowed uniformly from base distad to beginning of about distal fifth of length, where it is very obliquely truncate on exterior side, apex acute (Plate 8, fig. 2). Length 11 to 17.5 mm. A male 17 mm. long has anal leg 4.7 mm. long; antenna 5 mm. long; and tenth plate 1.93 mm. wide. A female 15 mm. long has tenth plate 1.57 mm. wide. Pseudomaturus.— Coloration as in maturus or somewhat paler in average specimens. Antennae as in maturus. 300 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ocelli somewhat fewer and frequently in but three series: e. @., Ay 4A: Claw of female gonopods smaller and often with only one, the ventral, of the lateral teeth distinct. Bristles of joints somewhat fewer. Spines as in maturus, or the inner relatively a little smaller. Spining of legs as in adult, but dorsal spines of anal legs in male more often only 1, 0, 4, 1, 0. Relatively somewhat wider in proportion to length than fully matured specimens Length 9.5-12 mm. Praematurus (late).— Dorsum light brown or testaceous, the pos- terior plates darker and sometimes verging toward chestnut. Head chestnut. Antennae at base brown or light chestnut but paler over most of length with tip lightest. Venter dilute brown, with the anterior plates somewhat darkened and the posterior ones more strongly so. Legs pale yellowish to pale greyish brown, the pos- terior pairs in some degree darker but pale distally. Prosternum similar to head but more dilute, the prehensors somewhat lighter. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 29 : 25:27:31 :32:30. Proportionately broader than in adult, sometimes being only eight times longer than width of this plate. Antennae essentially as in adult. Ocelli small, well separated; in number and arrangement: e.4., fic So3cas Olay Ae 2. Prosternum nearly as in adult, but in the one case measured only 1.5 times wider than long with distance between chitinous spots 2.73 times width at sinus and 4.1 times dental line. Coxal pores: smalli:h373,13;3;) 3) 4,433) dy O45 OF Claw of female gonopods small, thin, and pale, acute, the exterior or ventral tooth small and the inner or dorsal one not at all or but weakly indicated. Basal spines 2 + 2, the same form as in adult or nearly so, but the inner relatively much shorter and more slender, acute. Spines of first legs, (7-4 ORO 3! seventh to eleventh, 9 °0°5 LORS as Ee of the second to Sixth, rae #2. of the twelfth, j-¢-3-3 OF i t a3; Of the thirteenth, j3-3°5 or = : x 2 to (-res2 OF i : a of the penult, iss; of the anal (9), ae ‘;. Last two pairs of coxae laterally armed. (De Witt, Ia.). Immaturus.— Dorsum pale brown, with caudal plates darker. Head brown or dilute ferruginous brown, paler in front of the suture. ee o' CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. Sill Antennae pale greyish brown or yellowish, brighter distad. Pro- sternum yellowish brown, prehensors lighter. Venter pale, caudal plates more densely pigmented, yellowish to orange. Legs pale, brighter at tips, the posterior pairs more pigmented, brightest at tips like the others. Body only 7.5 times longer than width of tenth plate thus still wider proportionately than in praematurus. Widths of head and usual series of plates to each other as 22 : 18 :19 : 22.5 : 23 : 22. Head relatively longer than in adult in specimens examined (<<) being but little wider than long. Antennae very short with 20 articles already present, these short and very short; antennae submoniliform. Ocelli mostly small and well separated; single ocellus relatively large; ocelli of top row largest of seriate ocelli. Ocelli in three series: 6. G51 4 4, 2; 23 1.43, 4,.2. Prosternum 1.54 times wider than long. Distance between chitin- ous spots 2.85 times width at sinus; 4.44 times dental line. Teeth 2+ 2, rather small, line of apices a little recurved. Spine and sinus as In praematurus. Coxal pores: 2, 2, 3,2; 2, 3, 3, 2; 2, en Spines of first legs, oe a 1 to aN oT and o0-0-21; of the second, 1 1 0, 0,.0, 2, 2 0, 0, 2, 2 0:10.41, 22 ery” Lee. Wane MO Honea nd 70,3521 23 of and fourth, (yoo 252 = — eke and 4 ’ ae, ), 1, 2, +3; of the fifth to seventh, op 0; 0, 0, 2, 2, 2 0, 0, 3, 2, 2 nUgSs ), 0. 1, 2,2 to 0, 0, 2, 3, 29 of the tel ath, Osos Sicventh: ieaa; of the twelfth, oorsa-3; of the neat Pose ) cosas; of the penult, 7-333; of the anal (SM), Raaay OF ee (observed on one side). ee pair of coxae laterally armed. Anal legs of male with third joint but little produced mesad at distal end. Fourth joint already showing in a distinctly developed form the flattened mesodorsal process toward its proximal end, but this process relatively much shorter than in the maturus and the angle formed by anterior and posterior edges much more obtuse, the anterior side oblique, not nearly at right angles as in adult, and the lobe not showing the acuteate hairs; the smaller subconical processes at distal ends of third and fourth joints of adults not yet at all indicated. Length 6.5-7.5 mm. A male 7.5 mm. long has antennae 3 mm. and anal leg 2.6 mm. long with width of tenth plate 1 mm. (DeWitt and Tama, Iowa). Pullus IV (Larva IV).— Dorsum pale qellowish white with the head more pigmented, yellowish. Antennae and legs whitish. Prosternum and prehensors like head. Venter nearly same as dorsum of body. oon BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body more strongly narrowed caudad of tenth plate than in adult. Ratio of widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 32 : 26.5 : 28 : 30 : 28 : 23. Head equal in length and breadth. Caudal margin straight or a little excurved. Antennae composed of seventeen articles which, excepting the first three and the ultimate one, are very short; submoniliform; sub- sparsely hirsute. Ocelli 1 + 2,1. The first, or caudal, ocellus of upper series relatively very large, as large as or larger than, the single one and much larger than others of series. Prosternum 1.54 times wider than long. Distance between chitin- ous spots 2.55 times width at sinus and 3.53 times dental line. ‘Teeth 2 + 2, small, subequal with line of apices straight or slightly recurved. Spine and sinus as in older stages, or former, proportionately to teeth, somewhat stouter. A single coxal pore present on each twelfth coxa. ‘ho 2 0, 0, 0, 0, 1 . 0, 0, 0, 0, Spines of first legs, 0,0, 0,0, 1; of the second to eighth, Coord; of the . 00,00, 0,.0,0,0, , 0, 0, 0, 0, . ninth, 9-001 OT oo Lia; Of the tenth to twelfth, goa o 6(De Witt, ’ as): Last three pairs of legs showing only as short pale buds in which the joints, excepting the coxa, are not as yet distinctly indicated. Type Locaurry.— Illinois: Rock Island. Found also in Illinois at Franklin Grove, Peoria, Sterling. Iowa: De Witt, Tama. Nebraska: Grand Island. Kansas: Cowley, Douglas, Graham, Sumner, and Trego counties (fide Gunthorp). Indiana: Bloomington (Bollman). Minnesota: Winona (Holzinger coll., seq. Bollman). Wisconsin: Devil’s Lake. Mississippi: Byram. This is apparently a very common species in the states of the northern Mississippi valley, ranging also southward toward the Gulf as indicated by its occurrence in Mississippi. It varies much in many of its features and, from material available for study, seems to have a considerable tendency toward the formation of minor forms or varie- ties; but intergradation is frequent and, in spite of differences in local modes, it seems inadvisable to attempt their separation as varieties, except in the case of verdescens, until the species is more thoroughly known and the significance of the differences indicated more clearly understood. The species offers excellent material for a careful study in variation. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 353 POKABIUS BILABIATUS VERDESCENS Chamberlin. Plate 8, fig. 3. Poabius verdescens Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 154, pl. 13, oye; Ih DeEscrIPTION.— Dorsum sometimes from clear brown to light chestnut with first plate and caudal borders of others dark but more often brown of a pronounced olivaceous cast. Head either like dorsum but usually ferruginous brown or chestnut, dusky caudad of the suture. Prosternum olivaceous to dark brown. Venter typically light to dark olivaceous though sometimes clear brown. Legs pale olivaceous to brown with posterior pairs darker as usual. Body more robust than in bilabiatus proper, varying from only 6.7 to 8 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth and twelfth plates (in co”) to each other as 64 :- 63 :63-+ : 71:71 : 64. Antennae typically somewhat longer than in bilabiatus, reaching seventh segment. Ocelli as in the species. Prosternum typically relatively narrower than in the species, being from 1.5 to 1.57 times wider than long. Distance between chitinous spots 2.5 to 2.8 times width at sinus and 4 to 4.5 times dental line. Line of apices of teeth straight or very nearly so. Spining of legs very nearly as in bilabiatus, excepting that the ventral spines of the thirteenth legs, in all specimens studied, is 0, 0, 3, 3, 2, a formula sometimes present but not usual in the species. Anal legs of male as described for bilabiatus proper. Female gonopods essentially as in the species. Length 11-18 mm. Tyre Locauiry.— Illinois: Peoria! Taken also in Iowa: Tama! Mongona! Boone! Mississippi: Canton! This varietal form occurs in the same general region as the species proper. Judging from limited number of specimens of the immature stages examined, the ratio of length to width of tenth plate increases with development in bilabiatus, the slenderness becoming more and more pronounced; whereas, on the other hand, in verdescens it de- creases. Study of more extensive collections may break down the seeming differences between the two forms; but if so the range of variation in the species will be found to be unusually great. 354 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. POKABIUS DISANTUS, sp. nov. Plate 9, fig. 3. Lithobius clavigerens Chamberlin, Pomona coll. journ. ent., 1910, 2, p. 373. DEscripTion.— Dorsum testaceous. Head dusky behind suture and abruptly lighter in front of it. Antennae light distad. Pro- sternum dusky testaceous or yellowish; prehensors yellow. Venter dilute yellow, caudal plates dusky yellow. Legs very pale, the caudal pairs more densely pigmented, bright yellow. Body 9 to 10 times longer than width of the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 48 : 48 : 49 : 54:54:52.5 (o”). Head (in o”) slightly longer than wide (49:48). Weakly sub- sparsely punctate. Bristles moderately long, sparse. Antennae short; articles twenty, short, subeylindric, uniform. Ocelli ten to twelve in three series: e.g., 1 + 3, 4, 3; 1+ 4, 3, 2; pte yee Prosternum 1.48 times wider than long. Distance between chitin- ous spots 2.7 times width at level of bottom of median sinus; 4.56 times the dental line. Teeth small, acute, with line of apices recurved. Spine moderately stout at base, bristle-like distally, straight. First dorsal plate 1.7 times wider than long; sides gently convex, considerably converging caudad to and about the rounded caudal corners; widest just cephalad of middle. Caudal corners of ninth, eleventh, and thirteenth dorsal plates obliquely excised. Coxal pores small, eireular: 2 222M woven ae ] my) 1.22 0.0,2,2,2 0, 0,1, 2,2 Spines of first legs, P0734 to ; of the second, o-9-7-3-7 OF a » 0,0 121 FO 901,315 second’, 0,0,1.3,1, 9 ONOM2N 22 2,2 Q 2,2 , 0, 1, 2, 2 0, 0; 1, 3, 15 of the third to fifth, ? “0, a BY aL of the sixth a 0.0. Pe 3,1 or OW), ab 3, 29 of the seventh and eighth, } 0,0, mo Ol the ninth, * 0; + T 39; of th ne t te end) 0,0, 2,22 0,°0,'2,'2,'2 0, 0,3, 2,2 NOsaete Oa so; Ol the eleventh, 0°0,2,3,2 OF 0,0;2, 35 of the twelfth, 0-9:3'3'3; ale -3'3'1, Claws three; of the 2, 1, 0 T 3-9. None of the coxae of the thirteenth, foes; of the meu anal, (13:3) or, less commonly, (<7), armed laterally. Anal legs of male with the third joint elevated on dorsomesal side into a conspicuous, strongly chitinized ridge which is highest a little caudad of middle of its length, its edge rounding down evenly both ways; bearing two stout dorsal spines on caudal slope of ridge one behind the other, not in a transverse line at end of joint as in claviger- ens. Fourth article crassate, longitudinally furrowed along dorso- CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 300 mesal surface, the furrow limited on mesal side by a low ridge which is elevated at proximal end into a more transverse conspicuous lobe, thin in a nearly cephalocaudad direction but wider transversely, con- cave on caudal side and projecting cephalomesad; third, fourth, and fifth joints longitudinally furrowed on ectal side toward dorsal surface (Plate 9, fig. 3). Claw of female gonopods slightly curved; distinctly tripartite, with the median lobe much longest; the dorsal and ventral at about same level but the ventral smaller and more acute. Basal spines moderately long and slender; sides incurved between base and acuminate distal portion; a denticule at base of acuminate portion on each side with sometimes a second one farther toward apex. Length 9-11 mm. A male 9 mm. long has antenna 3.25 mm. and anal leg 3— mm. long, with tenth plate 1 mm. wide. Tyre Locatity.— M. C. Z. No. 276.— California: Laurel Canyon, Los Angeles Co., June, 1909. Found also in California at Eaton’s Canyon, April, 1913. This species much resembles P. clavigerens but is clearly distinct -from that species in spining of legs as well as in the form of lobes of anal legs and in other features. PoKABIUS UTAHENSIS (Chamberlin). Plate 10, fig. 3, 4. Lithobius utahensis Chamberlin, Proc. U. 8. N. M., 1901, 24, p. 22 (nee Ann. Ent. soc. Amer., 1909, 2, p. 187). Lithobius obesus Bollman (nec Stuxberg), Proc. U. 8. N. M., 1888, 11, p. 347. Poabius utahensis Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 153. Description.— Dorsum yellow, frequently of a dilute orange tinge. Head pale orange, commonly with the frontal region somewhat paler. Antennae like the head proximally, yellow distally. Prosternum and prehensors like the head. Venter pale yellow, darker, dilute orange, caudally. Legs pale, the posterior pairs more densely pigmented as usual, bright yellow or dilute orange. Body in females on average shorter and relatively broader than in males, being mostly from 8 to 8.5 times longer than width of tenth plate in the former, and in male often near 9.5 times longer. Widths of head and of first, third, eighth, tenth, and twelfth plates in an average individual to each other as 23 : 20 : 21 : 24 : 24.5 : 28. Head commonly slightly longer than wide (45 : 44). Caudal margin 356 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. straight or very weakly excurved. Head in front of eyes rather broadly rounded, subsemicircular. Smooth. Hairs moderately long, sparse. Antennae very short, reaching fourth or beginning of fifth segment. Articles twenty or sometimes twenty-one or twenty-two; mostly very short, decreasing in size from the second to the ultimate. Ocelli five to eleven, but mostly five to eight, in two longitudinal series: ¢.g.,1 + 1,3; 1+ 3,3; T+ 2,251 + 2,3; 1+ 5,2; 1+ 5,3. ‘Ocelli often irregular in form, size and arrangement and varying considerably in size in different individuals. Single ocellus of about same size as the larger ones in series. Prosternum 1.5 times wider than long. Distance between chitin- ous spots 2.33 times width at level of bottom of sinus; 3.5 times the dental line. Median sinus moderately wide, v-shaped. Teeth small and pale or scarcely darkened. Spine moderately stout at base, bristle-like distally, nearly straight. First dorsal plate about 1.66-++ times wider than long; sides cepha- lad of the rounded corners nearly straight and also nearly parallel. Posterior angles of none of the plates produced. Plates smooth and shining. Coxal pores small, with the most proximal one on each coxa often TUNE 2, he, eee 3,2; DSO MONO OR 3. ONONZS 22: 0, 0, 2, 2, 2 Spines of first legs, pp-q'3-2 Or sometimes 9° 9,1": of the second to 0,125.25.2), 0, 0, r e sev enth, °: O13-9; Of the eighth to eleventh, 9-9-3:3-3; of the twelfth, a ale 1,031.1 S 1 ONOaIT 23-2 2 OF sometimes 9.9°3°3°33 ot the thirteenth, 9:9°3'3'7 or, less com- aie ah al On; 1, 0, 3, 1, 1 monly, . 02:31; Of the penult, | 0133-1 OF 0133-1 claws three; of the anal, } 1 es 0 or : aa 3 = > Claw one. Last pair of coxae laterally armed. Anal legs of male with the third article typically produced at distal end mesad or dorsomesad into a conspicuous lobe which is thin in cephalocaudal direction and is contiguous with a corresponding lobe at proximal end of fourth joint which bears below apex a patch of short spinules projecting caudad; third joint typically with a wide longitudinal furrow on mesal or mesodorsal surface, this limited dorsally or ectally by a longitudinal, ridge-like, elevation; the two mesal spines of dorsal surface of third joint short, distally rounded, and finely serrulate, projecting cephalad from lobe; fourth joint longi- tudinally furrowed along dorsomesal surface, the furrow ending anteriorly at base of lobe previously mentioned, deepest cephalad; fourth joint more or less constricted or excavated just caudad of the lobe and then thickening caudad, swollen (Plate 10, fig. 3). The lobe formed by the extension of third and fourth joints at their con- CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 357 tiguous ends subject to considerable variation in size. Penult legs of male slightly swollen but not otherwise specially modified. In the female the posterior legs are moderately slender. Gonopods of male pale; moderately narrowed distad with apex rounded; bearing one moderately long bristle. Claw of female gonopods short and relatively wide, rather strongly curved; claw tripartite, all lobes distinct, the middle one a little larger than the others. Spines 2+ 2; rather robust, and typically more or less clavately widening from base to the acuminate apical portion, the outer side of which is the longer and is finely crenulate or serrulate (Plate 10, fig. 4). Inner side of first article straight or nearly so. Length of females 7.5-9.5 mm.; of males 8.5-10.5 mm. A male 8.5 mm. long has antennae 2.4 mm. “ond anal leg 2.6 mm. long; width of tenth plate .896 mm. Pseudomaturus.— Yellow, with less of orange tinge than in maturus. Ocelli averaging in number toward lesser limit given for maturus. Claw of female gonopods paler and smaller with lobes more slender and acute. Spines a little less robust with inner relatively a little shorter. Length 6.5-7.5 mm. Be eiine a Coloration nearly as in older stages but lighter yellow with less of orange tinge. Widths of head and of first, third, eighth, tenth, and twelfth as SOL OUI col 2360: 382 37. Antennae very short; full number of articles present. Ocelli 1 + 1, 2; 1+ 3, 2. Single ocellus variable in relative size. Prosternum nearly as in older stages. 1.6 times wider than long. Distance between chitinous spots 2.36 times width at sinus; 3.7 times dental line. Coxalspores $25.2) 25:25°2) 2513.25) 25351352 Female gonopods with fone short, ae the second and third. Claw small, thin and pale; the three teeth evident, though slender and acute. Third joint mostly with one short bristle; second with two or three; first with four or five. Spines 2-2; the inner much shorter and more slender than the outer which is much as in maturus but smaller and often not at all clavately widened distad. Gonopods of male small and wart-like; as yet glabrous. . 0,0,1,1,1 0,0, 1, 2, : 0,0, 1, 2,2 Spines of first legs, (0-2-4 OT oororai; Of the second, o'p'o:21 OF 0,0, 1, 2.1, 0,0,1,2°2' 0,0,4,3,3 0, 0,1, 2,2 0, 0, 0, 2, 1) of the third, 0, 0, 0, 1, 2 OF 0, 0, 0, 2, 25 of the fourth, 0, 0, 0, 2, 25 of the 22,3 . O02) 2) et 050,212) 12 fifth and sixth, ¢ 0, a 022; of the seventh and eighth, ¢; 9°0;2°2 OF 0,0:1.3:55 358 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 2, 2,.2 of the ninth to elev enth, 5, paa3; of the twelfth, 9 toa; Of the thir- LON Zee LU Aa lea be teenth, 10,0,3,3, 2 OF 0,0, 2 3 23 of the penult, (334, claws three; of the anal, g-a-3-3" o Last pair of coxae laterally armed. Anal leg of male modified as in maturus but lobe at junction of third and fourth joints somewhat less developed with the spinules on its caudal tae EWEN: Length 5. 5mm. A female 6.5 mm. long has tenth plate .73 mm. wide, thus ae nine times longer than width of this plate. Immaturus.— Dorsum very pale, whitish, the caudal plates tinged with yellow. Head dilute orange. Prosternum and prehensors like head. Venter whitish, caudal plates tinged with yellow. Legs and antennae whitish. Antennae already with twenty articles but, especially the median and distal ones, considerably shorter than in the praematurus. Ocelli 1 + 2. The single ocellus smaller than the other two, the caudal one of which is largest. Pale. Prosternum with teeth small and pale. Spines at base relatively to teeth somewhat stouter than in maturus, abruptly narrowed near middle to bristle-like form. In specimen measured only 1.45 times wider than long. Distance between chitinous spots 3 times width at sinus; 4.25 times dental line. Coxal pores very small: 1, 2,2, 2; 2, 2,2, 2. Gonopods of male small, whitish, glabrous. Gonopods of female short and pale. Claw appearing as a small, acute pale point with no indication of lateral teeth or in earlier stages claw wholly absent. Third and second articles: glabrous; the first with one or two bristles. Spines 1 + 1 or 1 + 2, small and acute, pale. Spines of first legs, 0. ° o 0: i; of the second to fourth, 0: 3 eel 11; of the fifth, o-o-oaci OT oo; of sixth to tenth, o> O41 113 or in part, especially . 0, 0, 0, 2, 2 0,0, 1,22 in the more anterior ones, 90-011; Of the eleventh, 9-907 4 of the aD 16,341 twelfth, 0 vied; of the thirteenth same as oo OF 90.22.13 of the AOS aI st tO I (spa aUal, 0, 3, 1, 6 1,0, 2,1, 0 penult, o-13°31 0/1231 OF o-ne21; Of the anal, oy 3r0 Or O31 None of coxae as yet laterally armed. Anal legs of male with fourth joint constricted, or excavated, near proximal end and the third and fourth joints more or less produced at junction into a lobe similar to, but usually less marked than, that in older stages, the spinules also present. Dorsal spines of third joint less modified. Length of principal specimen described, 5.4 mm. Agenitalis II. Whitish, of faint yellowish tinge caudad. Head dilute orange or yellowish. Antennae and legs whitish. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 309 Antennae composed of seventeen articles which, excepting the first few and the ultimate one, are very short. Bristles proportionately long and straight, not dense. Ocelli 1 + 1 or 1 + 2. First ocellus in series largest. Prosternum with teeth, sinus and spines as in 7mmaturus or nearly so. 1.425 times wider than long, thus narrower than in immaturus, the proportionate width increasing with development throughout. Distance between chitinous spots 2.66-++ times width at level of bot- tom of median sinus; 3.42 times dental line. Coxal pores minute: 1, 1, 1, 1. Gonopods not yet exposed. . 0, 0, 0, 0, 0, 0, 0, 0 Spines of first legs, (i t'o4; of the second to twelfth, o:oo:115. of the 0,0, 1, 1,1, Quote Ai 0,0, 1, 0, 0 thirteenth, ¢oo'11; of the penult, ¢7ia3; of the anal, gyoro or 0, 0, 0, 0, 0 o12to- No lateral spine on any coxae. Anal legs already nearly as in older stages but lobes weaker. Length 3.5-4.5 mm. | Tyre Locauiry.— Utah: Neff’s Canyon, Salt Lake Co.! Taken also in Utah in Mill Creek Canyon, City Creek Canyon, Farmington Canyon, Ogden Canyon, Cottonwood Canyon, American Fork Canyon, Provo Canyon; Spanish Fork Canyon, and _ other canyons of the Wahsatch Mts., where it is common under damp leaves along the streams. | PokaBius sokovus (Chamberlin). Plate ties 2; 3: Lithobius sokovus Chamberlin, Ann. Ent. soc. Amer., 1909, 2, p. 189. Poabius sokovus Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 153. Description.— Dorsum and venter very pale testaceous to whitish yellow. Head and antennae with prosternum and prehensors pale brown. Legs from whitish to pale yellow. Body of type 8.8 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each Othereasy Zor 22-Des2o 225 2252 24, Head well rounded behind but the caudal margin straight. A little wider than long (23 : 22.5). Antennae short, reaching the fifth segment as usual. Articles nineteen to twenty, short, with ultimate equalling the two preceding together. Ocelli six to eight arranged in two series: e.g., 1 + 8, 2; 1+ 4, 2; 1+ 4,3. Single ocellus small, not much differing in size from caudal ocelli of series. 360 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Prosternum 1.55 times wider than long. Distance between chitin- ous spots 2.9 times width at sinus, the anterior portion of prosternum being narrow with sides strongly converging; 4.37 times the dental line. Teeth very small and pale, the inner the larger and the line of apices thereby nearly straight. Median sinus rather small, v-shaped. First dorsal plate with sides in front of the well-rounded caudal corners nearly parallel. Posterior angles of none of the plates pro- duced. Coxal pores small, circular: 1,2, 2,2; 2,2; 2,2. ® 0, 0, 2, 2, 0,.0, 2;.2, 2 : ue of first legs, ¢--237 or goed; of the second and_ third, 0,0, 22, 2 0, 2,22 : iE of the fourth and fifth, ¢ 01,3) or Ges; of the sixth, 0,2,3,15, of the seventh to dleventh ah ial Eee eeeas; of the penult 13. cessory minute; of the anal, $7 ee laterally armed. Anal legs of male short and moderately inflated. Third joint ele- vated dorsally in a longitudinal ridge and at distal end in a transverse one much as in utahensis; fourth joint dorsally with a conspicuous longitudinal furrow which is widest and deepest proximally, the furrow being limited along the sides by a low ridge-like elevation and at the anterior end by the usual more conspicuously elevated lobe which extends dorsomesad and which is continuous on each side with the low longitudinal ridges, the lobe concave on its caudal surface, the concavity being a continuation of the dorsal furrow (Plate 11, fig. 2). Claw of female gonopods tripartite. Spines moderately long and slender, with the sides subparallel or a little incurved between base and short, acuminate, distal portion (Plate 11, fig. 3). Length 6.5-9 mm. Type Locatiry.— Nevada: Las Vegas (1909). Dug up in loose soil in a garden. The female is not quite fully mature. 0, 0, O05 ees 0, 239; Of the twelfth and thir- , claws two or an anterior ac- 3, 0 9, claw one. Last pair of coxae POKABIUS CLAVIGERENS Chamberlin. Plate 9; fig. 1; 2 Lithobius clavigerens Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 159. Pomona coll. journ. ent., 1910, 2, p. 373 (in part). Lithobius manni Chamberlin, Pomona coll. journ. ent., 1910, 2, p. 373 (pseudo- maturus var.). Poabius clavigerens Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 153. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 361 DEsScRIPTION.— Testaceous or light brown, or rarely of greyish cast suffused with violaceous (freshly moulted specimens) to darker brown with caudal plates commonly somewhat darker, chestnut or subferruginous; posterior borders of plates often darker and also in part with an interrupted longitudinal median stripe. Head mostly of a dilute ferruginous tinge, darker caudad of suture. Antennae similar to head, but typically pale proximally and at very tips. Pro- sternum usually much lighter than the head. Venter pale, the more caudal plates more deeply pigmented, yellow to somewhat ferruginous. Legs pale, the caudal pairs more deeply pigmented, brown, often lighter distally and along mesal surface where color may be yellow or some shade of orange. Body moderately robust, being typically from 7.75 to 8.5 times longer than width of tenth plate, but sometimes only 7 times longer (var. manni). Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 44 : 41 : 43 : 53 : 54:51, the first plate being clearly narrower than the third. Head from considerably wider than long, as in the type (44 : 41) to nearly equal in length and breadth or but slightly wider (only about 29.5 :29 in var. manni). Cordate, considerably narrowed caudad of breaks and strongly so in front of eyes. The most distinct sulcus is the median longitudinal one in front of the suture which unites with a transverse one parallel with and a little in front of the suture. A short oblique sulcus, not very deeply impressed, each side of the middle just back of the suture the inner end of which is near a small, pit-like impression. The usual semicircular depression on caudal portion shallow and wide. Irregularly punctate. Antennae short, the usual twenty articles short, decreasing distad; reaching to fifth or sixth segment. Ocelli twelve to twenty-five but mostly sixteen to twenty-five (clavi- gerens, forma typica), arranged in three or four, or, more rarely, even in five or six, series: ¢.g., 1 + 5, 5, 3; 1+ 5, 5,4, 2; 1-4 5,5, 4, 1; ie oon e lo oy ono, 25 ltd, Oy Onose 6,15, 0,88) fc-656; 6, 5, 1; 1+1,5,6,6,5,1. Ocelli frequently well separated, not contiguous, regular in form and arrangement, decreasing cephalad, black. Single ocellus large. Prosternum 1.53 to 1.58 times wider than long. Distance between chitinous spots 2.3+ to 2.55 times width at level of bottom of sinus; 4.2 to 4.4 times as long as the dental line. Sinus wide, strictly v- shaped (manni), or with sides concave and widely rounded at bottom (forma typica). Spines moderately stout at base but attenuated and 362 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. bristle-like distad, straight. Teeth small, the mesal one of each pair the larger so that line of apices is commonly almost straight. First dorsal plate moderately narrowed caudad; about 1.85 times wider than long; sides, except at ends, nearly straight or but gently convex, moderately converging caudad. Dorsal plates mostly show- ing two short, deeply impressed, diverging, longitudinal lines at ante- rior end close to median line. Weakly roughened. Posterior corners of ninth, eleventh, and thirteenth dorsal plates mostly obliquely and somewhat concavely excised; others with corners less extensively excised or else subrectangular. Ventral plates with each lateral longitudinal furrow on anterior por- tion ending abruptly in a deeper pit or bending directly ectad, espe- cially so on the anterior plates. "The median longitudinal sulcus represented on anterior plates mostly by a pit-like depression, but, more elongate on posterior plates. Last two plates of leg-bearing segments with corresponding coxae and also succeeding sternite sub- densely pilose. Coxal pores small, with the most proximal on each coxa often minute, with series sometimes irregular and even appearing double on some coxae: 3, 4, 4,3, the most usual numbers and arrangement, to 4,4, 4, 4 and 3, 4,3 + 3, 3. Spines of first legs, Put32 to judged of the second and third, 39.9 f 0, 0, 3, 2, 2 FON Ss2; ae 0, 0, 3,-2, 2 00-23-92 to oye: 3-4; of the fourth to eleventh, 99-33-53; of the twelfth, ORs Pehl oy OSS 032 sora cos 32; of the thirteenth and penult, 01,33, 1 Claws 3; of the 1,0, 2,1, 0 0,3, 1 WaNAgO anal, j-7-3'2°5 (male), or -4'3'5-p, Sometimes g4y'33-1 (var. mannt). Last two pairs of coxae laterally armed. Anal and penult legs short and but moderately thickened in female. In the male the fourth joint has the usual ridge-like elevation along mesodorsal surface, this increasing in height proximad and terminat- ing in a conspicuous flattened lobe lying oblique to the rest of the ridge, with the longitudinal furrow widest and deepest just caudad of this lobe; the rounded summit of lobe densely clothed with short, straight, fine hairs (Plate 9, fig. 1). Claw of female gonopods moderate in size, only slightly curved; tri- partite, with the lobes subacute, the middle one being but little longer than the inner or dorsal, the outer, or ventral, smallest. Spines moderately slender. Outer basal spine of each pair a little longer than the inner; in ventral view sides of spines subparallel from base to the acuminate apical portion or gently incurved between these limits; apical portion short and but moderately acute. Mesal side of first article straight and parallel with that of other gonopod or nearly so. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 363 Length 11-16 mm. A female 15.5 mm. long has antennae 5.5 mm. and anal legs 5.25 mm. long, with tenth plate 1.97 mm. wide. Pseudomaturus.— Coloration as in adult or but, on an average, a little paler. Antennae same as in maturus. Ocelli 1 + 5, 6, 3,1; 1+ 5, 5, 4, 2; fourteen to eighteen in three or four series. . Claw of female gonopods a little smaller and paler but otherwise as in the maturus. Spines not differing but bristles. somewhat fewer. Coxal pores a little smaller relatively: 3, 4, 4, 3. Length 9-11 mm. Tyre Locatiry.— California: Pacific Grove. Taken also in California at Monterey, Carmel, and Stanford. This is a very common species in the woods near Monterey and Pacific Grove where it is found under fallen logs and the bark of the same. PoxKaBIUS PITOPHILUS (Chamberlin). Plate 9, fig. 4, 5. Lithobius pitophilus, Proc. Acad. nat. sci. Phil., 1903, 55, p. 157. Chamberlin, Pomona coll. journ. ent., 1910, 2, p. 373. Poabius pitophilus Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 153. DeEscriptTion.— Brown to chestnut. Head somewhat darker with frontal region paler than region caudad of suture. Antennae chestnut, rufous at tips. Prosternum brown or pale chestnut, with the prehen- sors lighter. Venter testaceous, the caudal plates darker as usual. Legs pale brown, the posterior pairs light distally. Body in types 7.3-7.5 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 32 : 31 :33 :38.:38 or 39 : 36. Head considerably wider than long (8 : 7). Caudal margin widely and considerably incurved. Of about uniform width in region be- tween breaks and eyes. Antennae short, reaching fifth segment; articles twenty, short and approximately uniform. Ocelli mostly twelve to twenty in four, or occasionally in three, longitudinal series: e.g., 1 + 5, 4, 2; 1+ 3, 5, 3,4; 1+ 4, 5, 5, 4; 1+ 4, 5, 4,5; 1+ 5, 5, 5, 3. Single ocellus much largest. Other ocelli regular in form and arrangement, with the most ventral smallest as usual. 364 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Prosternum typically about 1.48 times wider than long. Distance between chitinous spots 2.7 times width at level of bottom of median sinus; 4.4 times the dental line. Sinus widely v-shaped and rather shallow, its sides a little concave. Teeth small, the inner one on each side distinctly and considerably larger than the outer and the line of apices thus straight or sometimes even somewhat procurved. Spine small and slender. First dorsal plate about 1.75 times wider than long; sides but moderately excurved and converging caudad. Posterior minor plates with caudal corners abruptly rounded, not excised. Coxal pores small, circular: 3,4,4,4; 4,4,5,4; 4,5,5,4; 4,5, 5 Spines of first legs, 0 0: a 3 3 5 Or 0: of = of th a of the tenth, j-7-33 a 3; of the elev enth, 9-9@ and thirteenth, §}} a of the penult, L ~ a anal, j4°3'2-5 OF o° a, 3. ou p Claw one. Anal legs of male short with the fourth joint moderately inflated. This joint dorsally with the usual conspicuous, longitudinal, ridge-like elevation which is much highest at beginning of middle third of length of article where it is produced dorsad or mesodorsad into a rounded lobe between which and the anterior end the ridge is lowest; at caudal end the ridge is produced into a small, subconic process which extends dorsocaudad. Penult legs rather slender, weakly longitudinally furrowed above. Claw of female gonopods moderately long and curved; tripartite, with the lobes rather short and often blunt, or sometimes almost obliterated and leaving the claw subentire. Spines short and rela- tively wide, the sides somewhat diverging from base to acuminate apical portion which is short and, especially in the outer spine, ob- tusely angular with outer slope much more oblique than the inner and somewhat crenulate; inner spine inserted farther dorsad so that in ventral view it is often partly covered by the outer one. Mesal sides of first article of the two gonopods moderately diverging cephalad but not at all excavated. Gonopods of male small, sides convex and strongly converging to the short, subtruncate, distal end which bears in an indentation a very stout bristle, a second and sometimes a third smaller bristle also present. Length 10-13 mm. A male 10.25 mm. long has anal legs 3.5 mm. long and tenth plate 1.39 mm. wide. Tyrer Locaurry.— California: Truckee (1902). The types were taken in the pine woods about Truckee, where the species is common. V5), 2,2, he second to ninth, ? 0-0°2:3-33 2 1,0,3, 22 2, OF 0,0, 2, 3, s. of the twelfth 1 5, Claws three; of the CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 365 PoxKasius 1GINus (Chamberlin). Plate 8, fig. 4; Plate 10, fig. 5. Poabius iginus Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 154. DescrieTion.— Dorsum brown, the posterior plates mostly darker than the others; in some the major plates have caudal borders dark- ened. Head brown of dilute ferruginous cast with typically a dark, smoky, or blackish, median area between frontal suture and caudal margin. Antennae similar to head proximally, becoming paler distad, the ends being yellowish. Prosternum and prehensors similar to head but lighter. Venter yellowish to dilute brown, the last few plates much darkest. Legs nearly like corresponding plates of venter or a little paler; the caudal pairs lightest distally as usual. Typical full grown males average near 7 times longer than width of tenth plate but younger males and females are more slender, being as much as 8.4 times as long as the tenth plate. Widths of head and of first, third, eighth, tenth and twelfth plates to each other about as 39 :39 :41 :45 :45 : 43.5. Head only slightly wider than long (39 : 38). Portion back of eyes subquadrate, the sides between eyes and lateral breaks nearly straight and parallel, from breaks rounding about corners caudad as usual. Caudal margin straight. A transverse sulcus in front of caudal margin which is united or continuous with two straight longitudinal sulci moderately diverging from it cephalad. Hairs short and sparse. Antennae very short, reaching only to the fifth segment, attenuated distad in the usual degree. Articles twenty; sides of each article straight, but little diverging distad. Ocelli twelve to fifteen in three, or sometimes in four, series: ¢.g., 1+ 5, 5, 3, a common arrangement; 1+ 5, 4, 3; 1+ 5, 5, 3, 1. Single ocellus of moderate size. Ocelli of series small and nearly uniform, compact, black. Prosternum 1.53 times wider than long. Distance between chitin- ous spots 2.85 times width at sinus; 4.6 times dental line. Sinus rather wide, sides a little concave. Teeth small; line of apices a little recurved. Spine short and slender, not bristle-like. First plate 1.85 times wider than long; sides straight and only slightly converging caudad. Plates smooth or under lens appearing only slightly obscurely and finely roughened. Caudal margins of all plates straight, the caudal corners simply rounded or subrectangular. 366 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Coxal pores very small or minute, circular: 2, 3, 4, 2; 2, 4, 3, 2; Des By by, OD oh, ee ley Spines of first legs, 7 a0 oo 4 to ae 5; of the second to eleventh, oo >: 3°53 of ile twelfth, teeta fe! of the thirteenth and penult, ¢-34 i, of the anal, §° 1° 9 Claw one. Last two pairs of coxae laterally armed. In the anal legs of male the third joint, or prefemur, is strongly clavately thickened distad but is not extended mesad in a special lobe as in most related species, and the conical process is only ob- scurely, or not at all, indicated. The fourth joint bears a little caudad of its anterior end the usual conspicuous lobe extending in a.meso- dorsal direction, but this is smaller than usual and its caudal slope is steeper and shorter, the usual ridge-like caudal extension of the eleva- tion absent or but obscurely indicated but the process at the caudal end of the joint is well developed, subeylindric, bearing a few aculeate hairs, a small number of which also occur on the main lobe. Third, fourth, and fifth articles longitudinally suleate dorsally, the sulcus of the fourth lying ectad of the process (Plate 8, fig. 4). Corresponding joints of the penult legs also similarly suleate. Claw of female gonopods very short and relatively broad, thin and nearly straight; tripartite, the lobes rather obtuse and distally more or less rounded, the median but little longer than the others. Basal spines short and stout, in surface view with sides parallel or a little diverging distad to the acutely pointed apical portion, broad surface of inner spine nearly at right angles with that of the outer one (Plate 10, fig.5). Inner sides of first article of the two gonopods a little diverging from each other cephalad as usual. Gonopods of male very short; distally bearing a bristle which arises from a slight indentation, no other bristles being observed in type. Length of types 11-12mm. A male 11.5 mm. long has antennae 5 mm.long and anal leg 4.5 mm. long, the tenth plate being 1.64 mm. wide. Pseudomaturus. ration as in maturus or a little paler. Specimens studied considerably more slender than the typical adults, being near 8.5 times longer than width of tenth plate. Antennae as in maturus. Ocelli 1 + 4, 5, 3; 1+ 3, 4,3; 1+ 3,4, 2 Coxal pores: 2, 3, 3, 2, minute. Specimens of this stage studied cannot be separated on basis of length from the true maturus, being 10-11.5 mm. long. Type Locatiry.— Washington (E. Bergroth). This species seems to be nearest to P. pitophilus, which it probably replaces in the north. CHAMBERLIN:, NORTH AMERICAN LITHOBIIDAE. 367 POKABIUS CARINIPES (Daday). Lithobius carinipes Daday, Term. fiigetek., 1889, 12, p. 153, pl. 5, fig. 31. Chamberlin, Pomona coll. journ. ent., 1910, 2, p. 372. The original description, somewhat rearranged, is as follows: — Brunneo, medio dorsi fusco fasciato. Corpore gracili. Antennis sat longis, 22- articulatis, dense crinitis, in apice violaceis. Oculis utrinque 15 in seriebus tribus positis. Coxis pedum maxillarium 4-dentatis. Laminis dorsalibus inermibus, sparsim ciliatis. Poris coxalibus uniseriatis 2, 3, 4, 3. Pedibus analibus [maris] sat crassis, articulo tertio quartoque supra suleatis, articulo tertio sulco profundiore carinaque in apicem rotun- data aculeataque instructo. Ungue pedum analium simplici; calcaribus 0, 1, 3, 2, 0; articulo primo calcari laterali armato. Longit. corp. 15 mm. Latit. corp. 2.5 mm. Femina ignota. Locatity.— Northern California: One male (Joanne Vadona). The type is in the National Museum at Budapest. It will probably be found that the process referred to the third joint of the anal legs involves also a contiguous lobe at proximal end of the fourth and that the species is a true Pokabius. LopHoBIus, subgen. nov. PoxKaBius socrus Chamberlin. Plate 12, fig. 2. Lithobius socitus Chamberlin, Proc. U. 8. N. M., 1901, 24, p. 23. Descrietion.— Dorsum yellowish brown to clear brown, some- times with a paler median longitudinal line. Head of same shade as dorsum or darker brown, sometimes of dilute chestnut cast, frontal region somewhat paler. Antennae light brown, paler distally. Prosternum and prehensors dark brown or dilute chestnut. Venter yellowish brown, the posterior plates deeper brown. Legs light brown, the posterior pairs darker like the corresponding plates of venter. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 80 : 73 : 74 : 85 : 85 : 84. COMPARATIVE ZOOLOGY. MUSEUM OF BULLETIN: 368 Fic. 9.— Distribution of Lophobius. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 369 Head subcordate, considerably wider than long (8:7). Marginal breaks slight or scarcely evident. Caudal margin widely incurved. Antennae short as usual, reaching fifth or sixth segment. Articles normally twenty, occasionally up to twenty-five; first seven to twelve moderately long, the others short. Ocelli thirteen to twenty-two, mostly in four series, occasionally in three or five: e.g., 1 + 5, 4,3; 1+ 5, 4,3,1; 1+ 5, 4,3, 2; 1-+-5, 5, 3, 2; 1+ 5, 5, 4, 3, 2. Single ocellus much the largest. Organ of Tomosvary well separated from the ocelli. Prosternum proportionately very wide, being near 1.72 times wider than long. Distance between chitinous spots 2.1 times width at level of bottom of median sinus; 4.9-+ times the dental line which is proportionately very short. Teeth small, normally 2-+ 2, occa- sionally extra ones occurring. Margin extending from ectal tooth almost directly ectad for a considerable distance before abruptly bending back caudad. Sinus rather wide, somewhat v-shaped, narrowly rounded at bottom. Sides of first dorsal plate convex at ends but nearly straight be- tween, only weakly converging caudad. Posterior angles of the eleventh and thirteenth dorsal plates weakly produced; those of the ninth straight or somewhat excised. Coxal pores moderate in size and circular or sometimes somewhat transversely oval: 3, 4, 4, 3; 4, 5, 5, 45 5,60) Gi5: spines of first pair of legs, 9°p't:3:1) less commonly jji34; of the second, ¢'t'3'3; of the third to eighth, f eg3; of the hath to tenth, eS: of the eleventh, ¢: osss; of the twelfth, res; of the thir- teenth and penult, j-4°3'3'3, claws three; of the ana , Claws two, the accessory claw small. Anal legs of male rather slender; the fourth joint a little clavate, with a distinct dorsal longitudinal furrow mesad of which the joint is commonly a little thickened or elevated in ridge-like form, this highest at caudal end but not truly lobate or cristate. Claw of female gonopods short and broad, tripartite, with lobes acute or rarely with four or five more obtuse teeth or crenulations while, in the other direction, one of the three lobes normally present may be obsolescent. Spines broad, with sides more or less diverging from base to beginning of the acuminate portion (Plate 12, fig. 2). Length 12-14 mm.; of anal legs 5-5.6 mm. Tyrer Locatity.— U.S. N. M. No. 784. Utah: near Salt Lake City. Found also in Utah at various points in Salt Lake, Davis, and Utah counties. L ONONS Tose al, Po310 D (habehepal 370 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Under stones in hollows and gulches over the foothills and in lower portions of canyons. Often in association with P. colliwm. Poxkasius cottium (Chamberlin). Plate 11, fig. 4-6. Lithobius collium Chamberlin, Proce. U. 8. N. M., 1901, 24, p. 23. Nadabius colliwm Chamberlin, Bull. M. C. Z., 1913, 57, p. 62. DescrIpTIon.— Yellow to light brown, or, more commonly, with a decided tendency to ferruginous or orange; in some part of plates may be dusky. Head sometimes like dorsum but more typically a brighter lemon-yellow, or more reddish orange, or light ferruginous. Antennae mostly bright yellow or orange proximally. Body in most, slender and 8.5-9 times longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 39 : 37 : 38 : 45 : 45 : 44. Head wider than long in ratio (average) of 39 : 36.5. Sides only slightly excurved between eyes and marginal breaks, which are weak or obscure, and the diameter over this region nearly uniform. Caudal margin weakly widely incurved. A distinct transverse sulcus in front of median portion of caudal margining rim with two sublongitudinal sulci diverging cephalad from this. A median longitudinal sulcus as usual in front of suture. Antennae short, reaching only to the fifth segment. Strongly attenuated distad, the tips being very slender. Articles normally twenty, sometimes up to twenty-three, short, decreasing distad. Ocelli mostly fifteen to twenty-two in number; arranged mostly in four series: ¢.g., 1 + 5, 5, 5,3; 1-+ 5, 5, 3, 1. Prosternum 1.45 times wider than long. Distance between chitin- ous spots 2.64 times greater than width at level of bottom of median sinus and 5.38 times the dental line which is thus proportionately very short. Median sinus rather shallow and narrow, well rounded at the bottom. Spine small, slender and hair like. Teeth small; line of apices recurved; in some the teeth may be in part almost obsolete (Plate 11, fig. 4). First dorsal plate 1.7— times wider than long; sides convex, con- verging caudad of anterior third. Caudal corners of ninth, eleventh, and thirteenth dorsal plates obliquely excised. Smooth. Coxal pores small, the most proximal one on each coxa often minute and the most distal one much largest: 2, 3, 3, 2; 3, 4, 4,3; 3, 4, 4, 4. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. ofl Spines OL first and second legs, o: pees or oo oe 33; of the third to eleventh, », Ooa3¢3; of the twelfth and thirteenth, i o 2 x9; of the penult, Sa claws three; the anterior accessory comparatively long; of the anal, j7'3:3'¢, claw one. Last three pairs of coxae laterally armed. Anal legs short; slender distad of the fourth article which is thick- ened; fourth article clavately widening distad, considerably thicker in dorsoventral direction than in lateral, dorsally with a conspicuous ridge-like elevation which, beginning even with the general surface at the proximal end, rises in height to the caudal end where it termi- nates as asomewhat rounded prominence or crest, which is toward the mesal side and projects dorsomesad, the ridge curving somewhat mesad of directly caudad in its course from anterior to posterior end, the joint longitudinally furrowed mesad of the ridge (Plate 11, fig. 6). Gonopods of male small and rounded, wart-like, each bearing dis- tally two long bristles. Claw of female gonopods of moderate length, considerably curved; tripartite, the three lobes typically distinct with the median largest, but claw occasionally subentire. Spines moderately stout, the outer a little the larger; in view of broader surface the sides are subparallel or slightly divergent to the acutely acuminate apical portion (Plate 11, fig. 5). Length 13-16 mm. A male 15 mm. long has antennae 5.5 mm. and anal leg about 4.7 mm. long with the tenth plate 1.64 mm. wide. Pseudomaturus.— Coloration as in adult or a lighter yellow. Head, antennae, and posterior legs lemon-yellow. Gonopods of female with claw nearly as in adult or a little thinner. Spines slightly more slender, with the inner one sometimes relatively somewhat smaller. Dorsal spines of first legs mostly, 0,0, 2, 2,1. Ventral spines of twelfth and thirteenth legs more commonly 0, 0, 2, 3, 2. Anal legs of male as in maturus but ridge and crest a little less pro- nounced. Length 10-12 mm. Praematurus.— Yellow, head and posterior plates brightest. Ocelli 1 + 4, 3, 1; 1+ 4, 3, 2. Ocelli well separated, first of top series large. Coxal pores: 2, 3, 3, 2, minute. Claw of female gonopods with claw nearly same as in older stages. Third article with two bristles, second with three or four, and first with four to six. Spines 2 + 2, the inner much the smaller, close together, acutely acuminate from base. Sie BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Spines of first legs, i ??34; of the second, j472°3-2; of the twelfth, " S33; of the thirteenth, 3 pe3'3; of anal and penult as In adult. Length of specimen descr ibed 9.25 mm. Late immaturus.— Yellow with last plates and the head _ brightest. Antennae like head. Venter paler. Last pair of legs bright yellow. Antennae with twenty articles which are short and closely com- pacted. Ocelli 1 + 4, 3, small, well separated, the single ocellus smaller than first one of upper series. Prosternum nearly as in adult. Line of apices of teeth more nearly straight. 1.48 times wider than long. Distance between chitinous spots three times width at level of sinus; 4.3 times the dental line. The anterior portion proportionately less wide than in adult and the dental line relatively longer. Coxal pores minute: 2, 2, 2, 2. Gonopods of female with a claw present, thin, small, acute and pale with lateral teeth already indicated. Spines 1 + 1, acute, attenu- ated from base to tip. Second article with two bristles, the first with three or four. Spining as given for pracmaturus, which is early in that stage as the present is late in the immaturus. Length 9 mm.; antennae 2.6 mm.; anal leg 2.9 mm.; width of tenth plate 1 mm. Widths of head and of the first, eighth, tenth and twelfth plates to each other as 47 : 43 :46 :55 :55 : 53. Type.— U.S. N. M. No 7838. Utah: Salt Lake Co. Found also in Utah in Salt Lake City, Farmington, Ogden, Provo, Lake Point, Antelope Island, Fremont Island, ete. This species is very common in Utah under stones over the foothills during the damper months in spring and fall and in the lower parts of the canyons. It was found comparatively abundant under stones on the islands of the Great Salt Lake. PoKaBIus PUNGONIUS (Chamberlin). Sozibius pungonius Chamberlin, Ann. Ent. soc. Amer., 1912, 5, p. 152. DeEscrIPTION.— Dorsum light testaceous, with the caudal plates and the head darker. Antennae yellow proximally and at tips, the remaining portion somewhat darker. Prosternum and caudal ventral plates yellowish, the remaining portion of venter a paler yellow. Posterior pairs of legs bright yellow. Body of type 8.6— times longer than width of tenth plate. Widths CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. aie of head and of first, third, eighth, and tenth plates to each other as 33 730-35 335 : 34. Head in type precisely equal in length and breadth. Caudal margin straight. Margination distinct and continuous cephalad to eyes, there being no distinct marginal breaks though there is evident the very oblique suture through the marginal rim. Hairs sparse and moderately long. Punctae irregular and weak. Each antenna in type composed of twenty-one articles of which all but the most proximal and the ultimate are very short and closely united, decreasing distad. Antennae very short, reaching the fifth segment. Ocelli twelve to fifteen in four series: e.g, 1 + 2, 4, 4,4; 1+ 3, 4, 3,1. Ocelli small, black, irregular in form and size, with the series also irregular. Single ocellus small like the others. Prosternum only 1.4 times wider than long. Distance between chitinous spots 2.65 times as great as width at level of bottom of sinus; 4.24 times length of dental line. Teeth small, black, narrowly rounded or subacute distally; in type 2 + 3; line of apices distinctly recurved. Sinus narrow, sides acutely converging. Spine slender and acute, close to outer tooth. First dorsal plate 1.66-+ times wider than long; sides rounded, conspicuously narrowed caudad of middle. None of plates with caudal angles produced. Posterior plates weakly finely roughened and punctate. Coxal pores very small or minute: 3, 3, 3, 2. 0,0,2,2,1 0, 0, 2,22, Spines ot first legs, o'0,2:3-23 of the second to sixth, 0,0, 2.3.33 of the 02.22 0,033.3. 22. seventh, ? 0, 0, 2,3, 2 or 0, 0, 2, 3 2 3, 2 3, of the eighth to eleventh, ees; Of the penal al twelfth and thirteenth, Foes 2) 0! the penult, ¢-4°3:3:2, the claw one or 1, 0, 3, 1, 0 accessory claw obsolescent; of the anal, 0(7°3:o'7, claw one. Last two pairs of coxae laterally armed. Anal legs of male short and slender, not at all specially modified. Gonopods of male pale, rounded, bearing two bristles. Length 11 mm.; antennae and anal legs 4 mm.; width of tenth plate 1.28 mm. Tyre Locatiry.— Colorado: Marshall (T. D. A. Cockerell). Only the male of this interesting species is known. POKABIUS ARIZONAE, Sp. Nov. Description.— Light orange-yellow with the head and the poste- rior plates a deeper orange, the frontal region of head palest. An- tennae like head but lighter yellow distad. Prosternum and _ pre- 374 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. hensors light orange. Legs yellow, the posterior pairs more brightly pigmented, light orange. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 61 : 56 : 58 : 65 : 67 : 62. Head in type slightly longer than wide (63:61). Rather strongly narrowed cephalad. Marginal breaks weak, oblique. Caudal margin weakly widely incurved. Antennae short. Articles twenty-four; first four moderately long, the others decreasing gradually distad; ultimate shorter than the two preceding ones taken together. Ocelli 1 + 4, 4,2. Single ocellus largest though not greatly so. Prosternum 1.55 times wider than long. Distance between chitin- ous spots 2.68 times width at level of sinus; 4.25 times dental line. Teeth apically narrowly rounded; distance between teeth of each pair about equal to that between the two mesal ones. Sinus broadly v-shaped, the sides straight. Spine clearly stouter than the bristles, straight, regularly acutely acuminate. First dorsal plate widest a little cephalad of middle, not strongly narrowed caudad; 1.6 times wider than long. Posterior corners of the ninth, eleventh, and thirteenth dorsal plates obliquely excised. Coxal pores small, widely separated: Dearie Spines of first to fourth pairs of leg rS, wy 2 3-3; of the fifth to eleventh, 0, 0, 3, 2 12:03;3, 19:2 et. 3 0, 3, 1,1 00-2372; of the welneh 0-03-35 OF e 0-3°3,9; Of the thirteenth, 1.0. 8.1.1, of the penult, 9° a3 3-9, Claws three, ae the accessor y claws small but distinct. Anal legs of male slender, not modified. Gonopods of male well exposed; bearing two bristles. Length 12 mm. ‘Typr.— M. C. Z., No. 285, Arizona: Tucson (W. M. Wheeler). ‘This species is known from a single male. It is aberrant. POKABIUS HELENAE, sp. noy. Plate 12, fig. 3. Description.— Dorsum yellowish brown to clear brown, with sometimes part of the plates dusky. Head mostly more or less ferruginous with frontal region a little paler. Prosternum and pre- hensors testaceous of a dilute ferruginous tinge. Venter yellow to yellowish brown, with caudal plates ferruginous. Legs like corre- sponding plates of venter. CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 375 Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 76 : 72 : 74 : 85:86 : 84. Head widest immediately caudad of eyes, a little wider than long. Caudal margin mesally incurved. Marginal breaks slight. Antennae short and slender, usually reaching to the fifth segment. Articles normally twenty, moderate, sometimes the first seven to ten markedly and abruptly longer than those succeeding. Ultimate article longer than the two preceding together. Ocelli fifteen to eighteen in three or Ere frequently in four series: Eglo, 4; 1-5, 4.4.3; 14-5)0,38,2; 1-5, 5,42; Is 5; 6, 5,1; 1+ 6, 5,4, 2. Single ocellus eae largest, elliptic. Organ of Témosvary relatively small. Prosternum 1.65 times wider than long. Distance between chitin- ous spots 2.5 times width at level of sinus; 4.33-+ times the dental line. Sides of anterior portion convex and rounding out considerably laterad. Teeth very small, narrow at base, distally rounded. Sinus strictly v-shaped. Spines slender, distally bristle-like, straight or nearly so. First dorsal plate a little widest anteriorly; sides substraight and not strongly converging caudad to the rounded posterior corners; in type near 1.9 times wider than long. Posterior corners of the thir- teenth plate weakly produced, those of the ninth and eleventh typi- cally, somewhat concavely, excised. Coxal pores: 3, 4, 4, 3; A444. 4,5, 5,3; 4, 5, 5, 43 sete: t Spines of first legs, ¢ 02 33; of che fied to ninth, $3 i 23 of the tenth, $5; ae 233 of the eleventh, {>} a 25 of the twelfth, } 0-0: ogg OF ng a3 ; of the thirteenth and penult, oe = }3°2, Claws three; of the anal, bse Claw one. Last two pairs of coxa laterally armed or spine of penult coxae sometimes absent. Anal legs of male rather slender; the fourth joint a little crassate, a little incurved or depressed above and weakly elevated at distal end. Claw of female gonopods short, tripartite, the lobes all acute and distinct, the median longest though not greatly so. Spines moder- ately long and of moderate proportionate breadth, sometimes appear- ing bent mesad near middle of length, the sides typically subparallel from base to apical portion. Length 9-13 mm. ~ Typr.— M. C. Z., No. 185. Montana: Helena (W. M. Mann). 376 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. POKABIUS CASTELLOPES (Chamberlin). Plate 12, fig. 1. Lithobius castellopes Chamberlin, Proc. Acad. nat. sci. Phil., 1903, 55, p. 158. Pomona coll. journ. ent., 1910, 2, p. 373. DerscrieTion.— Dorsum testaceous; head but little paler. An- tennae light; paler distad than proximad. Legs yellowish, the caudal pairs more deeply pigmented and brown proximad, yellowish distad. Body in type 7 times, or a little less, longer than width of tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 28 : 26 : 27.5 : 32 :35 : 33. Head a little longer than ae (30 : 28.5). Cordate. Caudal margin mesally incurved. Head considerably narrowed cephalad. Sparsely clothed with rather long hairs. Smooth. Marginal breaks distinct. Antennae short; slender distad. Articles short and very short, decreasing distad as usual. Ocelli twelve in three series: 1+ 5, 4, 2. Patch elongate. Ocelli separated, not contiguous, those of top series considerably largest. Prosternum 1.4 times wider than long. Distance between chitin- ous spots 2.38 times greater than width at level of bottom of median sinus; 3.58 times the dental line. Median sinus strictly v-shaped, with sides straight. Teeth small and pale; line of apices recurved. First dorsal plate 1.8 times wider than long, widest anteriorly and considerably narrowed caudad. Caudal minor plates with posterior corners obliquely excised, but line of excision short. Smooth. Coxal pores very small: 2, 3, 3, 2. Spines of first to third pairs of legs, eS ee 25 of the twelfth, 77335; of the thirte penult, } o1-3-3-4 Claws three; of the anal, j-1-3-3"0» pairs of coxae laterally armed. Anal and penult legs in the male considerably swollen. Anal legs short; fourth joint thick, somewhat bowed ventrad with dorsal surface concave, saddle-like and at distal end elevated into a con- spicuous lobe which narrows to an edge dorsad and is truncate caudad on level with end of joint or distally a little extending beyond it; lobe on caudal surface near apex bearing a caudally directed spine; dorsal surface of joint cephalad of the lobe longitudinally suleate (Plate 12, fig. 1). of the four th to eleventh, enth, °° 0 z 3; of the claw one. Last two CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. BMiTh Gonopods of male well exposed; distally truncate; sides sub- parallel. Length of type (co) 8.8 mm.; antennae and anal legs 3 mm.; tenth plate cir. 1.28 mm. wide. Type Locatiry.— California: Shasta Springs (1902). POKABIUS EREMUS, sp. nov. DescriptTion.— Brown, with the head darkest. Antennae light brown. Venter light brown. Legs pale, posterior pairs darker, light distally. Body moderately narrowed cephalad from the tenth plate with the first dorsal plate slightly narrower than the third and much narrower than the head. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 47 : 50 : 51 : 62 : 64 : 64. Head slightly wider than long (57:56 or 55). Widest a little caudad of eyes from where narrowing regularly to the caudal corners, a little indented or incurved at level of suture where the marginal break would be if present. Caudal margin straight. Conspicuously narrowed cephalad. Antennae of intermediate length, reaching to sixth segment. Articles twenty to twenty-one, of intermediate length to rather long, subeylindric, each only slightly widening distad, of nearly uniform proportions; ultimate shorter than the two preceding ones taken together. Ocelli in type twelve in three series: 1-+ 5, 4, 2. Those of the upper series large, regular, deeply pigmented. Single ocellus not larger than the caudal ones of series, or but little so, and not otherwise sharply differentiated. Ocelli of most ventral series considerably smaller and paler than the others in the type. Prosternum in type 1.425 times wider than long. Distance between chitinous spots 2.8 times width at level of bottom of median incision; 4.5 times length of dental line, the anterior portion of the prosternum being especially narrow at its cephalic end and the dental line corre- spondingly short. Median incision large; sides concave; narrowly rounded at bottom; wide, the distance between inner teeth being greater than that between the teeth of each pair. Spine rather stout at base; long and slenderly acuminate; inserted a considerable dis- tance caudad of level of teeth. Margin sloping directly ectocaudad from base of outer tooth. 378 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. First dorsal plate with sides only weakly convex, regularly and only moderately converging caudad; in type cir. 1.78 times wider than long. Posterior corners of ninth, eleventh, and thirteenth dorsal plates obliquely excised (the right corner of ninth plate in type is not thus excised) but the caudal margin is a little angularly extended caudad just mesad of inner end of the excision. Coxal pores small, especially so the proximal ones: 4, 4, Spines of first legs, (oae4; of the second to seventh, 0-0" 2 eighth and ninth, ae ee of the tenth, OG eS Or Ba eleventh, §:¢:3'3°5; of the twelfth, CaS 3, 5 or o, : : th >; of t ores; of the penult, @333, claws three, the seues accessory being long and slender; of the anal, {'3°2'p, the claw single. Last two pairs of coxae laterally armed. Claw of female gonopods long and rather slender, strongly curved; tripartite, with the lobes all acute and the median not greatly exceed- ing the lateral ones. In ventral view the outer basal spine is narrowest near the middle, the sides incurving; the acuminate apical portion is very short, with one or both edges denticulate and the tip truncate. The outer spine is larger than the inner one which appears acuminate from very base and is distally acute. Length of type (@ ), 8.5 mm. Typr.— M. C. Z., No. 187. British Columbia: ‘Kaslo. The spining of the legs of this species at once differentiates it from the others at present known. It is the only one in which the tibia of the anterior legs bears two dorsal spines, and in which the tibia of the penult leg is armed dorsally. : "5; of the : of the Boe SIMOBIUS gen. nov. Marginal interruptions of head slight and oblique or sometimes scarcely manifest. Antennae short; composed in type species of twenty-one or twenty- two articles. Eyes composed of seriate ocelli; single ocellus distinctly differen- tiated, large. Prosternal teeth 2+ 2; line of apices recurved. Spines slender, bristle-like. Posterior corners of 11th and 13th plates (in type-species) weakly produced. Coxal pores small, circular; uniseriate. Claw of female gonopods tripartite, the median tooth not much CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 379 Fic. 10.— Distribution of Simobius. 380 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. exceeding the lateral ones. Spines 2 + 2, of intermediate length and breadth. First article of gonopods not excavated proximally. Tarsi of anterior legs divided. Anal legs of male short and moderately inflated; without special processes or modifications. Penult legs in male with fifth article bearing at the distal end on the dorsal surface a conspicuous process. Last two pairs of coxae laterally armed. Third joint of all legs caudad of the second pair normally with three spines. Fifth joint of all legs between first or second and twelfth with two dorsal spines. In type species, dorsal spines of anal legs 1, 0, 8, 1, 0; ventral, 0, 1, 3, 3, 1; claw single. Dorsal spines of penult legs 1,0, 3, 1,0; ventral 0, 1, 3, 3, 2; claws three. Dorsal spines of twelfth legs 1, 0, 3, 1, 2; ventral Op Unmaiose. Length 11 mm. and over. Typr.— S. ginampus (Chamberlin). The type species is the only one thus far known. In the character of the modification of the penult leg in the male, the genus suggests Guambius but is very readily distinguished from that genus in other characters. SIMoBIUS GINAMPUS (Chamberlin). Plate 12, fig. 4, 5. Lithobius ginampus Chamberlin, Ann. Ent. soc. Amer., 1911, 1909, 2, p. 187. Description.— Dorsum yellowish brown or testaceous to dark smoky brown, or somewhat mahogany-brown, when in full color show- ing a longitudinal darker stripe on all or most of the major plates. Head from light chestnut, in paler specimens, to mahogany or almost black in the darker ones. Antennae light brown to chestnut proxi- mally, paler distally. Prosternum from light brown to dark smoky brown; prehensors lighter, rufous, distad. Venter yellowish to brown, the last several plates darker. Legs yellowish to brown, nearly uni- form. Body proportionately wide, being commonly only 7, or less, times longer than width of the tenth plate. Body considerably narrowed cephalad with the first plate narrower than the head and than the third, the latter being but little narrower than the head. Widths of head and of first, third, eighth, and tenth plates to each other as 48 : 45 247 : 55 : 50. Head wider than long in about the ratio 48 :45. Rather strongly CHAMBERLIN: NORTH AMERICAN LITHOBIIDAE. 381 narrowed cephalad from caudal edge of eyes. Immediately in front of the suture is a median depression and a little in front of this a dot- like second impression; immediately caudad of the suture is a dot-like impression each side of the median line and a second similar pair farther caudad; the usual semicircular impression just in front of the caudal margin. Head otherwise nearly smooth, shining. Antennae very short; reaching fifth or sixth segment. Articles twenty-one or twenty-two, short and subcylindric; ultimate rather long. oe of each eye mostly ten or twelve in three series: e¢.g., 1 + 4, 3,2; 1+ 4,4,2; 1+ 4,4,3. Single ocellus large, ovate; of the other ocelli, the most caudal of the dorsal series are largest, the others decreasing cephalad and ventrad. First dorsal plate near 1.8 times wider than long. Most major plates showing a pair of furrows diverging from the anterior end caudad and the usual short, transverse, submarginal sulcus on each side near beginning of caudal third of length. Moderately rough- ened, the caudal plates more distinctly so. Posterior angles of ninth plate with caudal edge straight or a little bent cephalad; those of eleventh and thirteenth a little produced caudad. Ventral plates with the usual three longitudinal sulci. A submedian transverse impression which is most distinct on plates toward caudal end. Coxal pores small, decreasing proximad: 3, 4, 4, 3; 3, 4, 5, 3; 4,5, 4, 3; 4, 5, 5, 4; et e 0, 2, 2, 1 O05 25125 OROR2 Noa 0, 0, 2, 2, 1, Spines of feces 0 0, 01,2, 2 OF eS of thesecond, 0-071, 3-2 OF 0,0,1-3°33 EPae ae of the third to tenth, ? 0. ane 3-9; of the elev enth, ¢ 0° 3 3-9; of the twelfth, 1, 0,3,1,2 Kelair 0, 3180 003-372; of the thirteenth, ; any i + 3; Of the penult, 9 0-1,3,3,2, Claws three; of the anal, j° 1! 324, claw one. Last two pairs of coxae laterally armed. Anal and penult legs of male short and moderately inflated with the tibial joints abruptly more slender. Anal legs without special lobes. Penult legs with the fifth article having dorsocaudal corner of distal end obliquely excised as in Arenobius, ete., and bearing upon the oblique surface a conspicuous subcylindric but laterally compressed, process which is sublongitudinal in position and is as high, or nearly as high, at proximal as at distal end; free surface of lobe obliquely truncate, bearing but few bristles (Plate 12, fig. 4). Gonopods of male well exposed, with sides convex and the distal edge oblique and nearly straight, the ectal side being shorter than the mesal; bearing two bristles. Claw of female gonopods tripartite, the lobes or teeth short and 382 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. usually rather blunt, the median being longest though not greatly so. Outer spine longer than the inner; a little acuminate from base distad with the sides more or less incurved near the middle; apical portion short, narrowly rounded at tip (Plate 12, fig. 5). Length of types 11 to14mm. A male 13.5 mm. long has antennae 5 mm. long, anal legs 4 mm. long, and the tenth plate 2 mm. wide. Type Locauiry.— Washington: Pullman (W. M. Mann). Found also in Alaska on Forrester Island (Harold and R. W. Heath). PLATE 1. CHAMBERLIN.— Lithobiid Genera. PEATE: Sozibius pennsylvanicus Chamberlin. (Upsal, Pa.). Fig. 1. Fourth joint of anal leg, mesal view. Fig. 2. Ocelli of right eye and organ of Témésvary, showing typical relations. Fig. 3. Ocelli of left eye of another specimen, showing larger actual and relative size of ocelli. Fig. 4. Claw of gonopod of 9 viewed at right angles to its concave surface. Fig. 5. Basal spines of left gonopod of 9, ventral view. Sozibius tuobukus (Chamberlin). (Hot Springs, N. C.). Fig. 6. Third and fourth joints of right anal leg of <, dorsal view. BULL. MUS. COMP. ZOOL. LITHOBIIDAE. PLATE 1 PLATE 2. C HAMBERLIN.— Lithobiid Genera. PLATE 2. Sozibius tuobukus (Chamberlin). (Hot Springs, N.C.). Fig. 1. Claw of right gonopod of 2, subventral view. Fig. 2. Basal spines of left @ gonopod, ventral view. Sozibius providens (Bollman). Fig. 3. Anterior portion of prosternum. (Russellville, Tenn.). Fig. 4. Claw of right gonopod of 9¢, subventral view. (Russellville, Tenn.). Fig. 5. Basal spines of left gonopod of 9, ventral view. (Russellville, Tenn.). Fig. 6. Basal spines of right gonopod of 2. (Type, Bloomington, Indiana, showing, in comparison with Fig. 5, extent of variation in form). PLATE 2 LITHOBIIDAE, BULL. MUS. COMP. ZOOL. PLATE 3. CHAMBERLIN.— Lithobiid Genera. PLATE 3. Pampibius paitius (Chamberlin). (Unaka Springs, Tenn.). Fig. 1. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6. Fig. 7. Anterior portion of prosternum. Ocelli of left eye and organ of Témésvary. Third and fourth joints of left anal leg of o, mesal view. Basal spines of right gonopod of 2, subventral view. Typhlobius kebus Chamberlin. (Santa Barbara, Cal.). Anterior portion of prosternum. Claw of right gonopod of 2, subventral view. Typhlobius coecus (Bollman). (Saluda, N. C.). Basal spines of left gonopod of 9, ventral view. BULL. MUS. COMP. ZOOL. LITHOBIIDAE. PLATE 3 PLATE 4. CHAMBERLIN.— Lithobiid Genera. ioe) Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6. PLATE 4. Typhlobius coecus (Bollman). (Saluda, N. C.). Anterior portion of prosternum. Claw of left gonopod of 9, ventral view. Paitobius naiwatus (Chamberlin). (Landrum, N. C.). Anterior portion of prosternum. Basal spines of left gonopod of @, ventral view. Paitobius carolinae (Chamberlin). Anterior portion of prosternum. Basal spines of left gonopod of 9, ventral view. BULL. MUS. COMP. ZOOL. LITHOBIIDAE. PLATE 4 PLATE 5, CHAMBERLIN.— Lithobiid Genera. PLATE 5. Paitobius tabius (Chamberlin). (Johnson City, Tenn.). Fig. 1. Basal spines of left gonopod of 9, view a little ectad of ventral. Paitobius zinus (Chamberlin). Fig. 2. Basal spines of left gonopod of 92, ventral view. Paitobius adelus Chamberlin. (Upsal, Pa.). Fig. 3. Basal spines of left gonopod of 9, ventral view. Taiyubius angelus (Chamberlin). (Santa Barbara, Cal.). Fig. 4. Anterior portion of prosternum. Fig. 5. Basal spines of left gonopod of @. Taiyubius harrielae (Chamberlin). (Colorado). Fig. 6. Basal spines of left gonopod of 9, ventral view. Nadabius oreinus Chamberlin. (Santa Barbara, Cal.). Fig. 7. Anterior portion of prosternum. LITHOBIIDAE. PLATE 5 BULL. MUS. COMP. ZOOL. PLATE 6. CHAMBERLIN.— Lithobiid Genera. PLATE 6. Nadabius oreinus Chamberlin. (Santa Barbara, Cal.). Fig. 1. Basal spines of left gonopod of 2, ventral view. Nadabius mesechinus (Chamberlin). (Meacham, Oregon). Fig. 2. Basal spines of left gonopod of @. Nadabius aristeus Chamberlin. (Macapin, N. J.). Fig. 3. Basal spines of left gonopod of @. Fig. 4. Anterior portion of prosternum. Sonibius parvus Chamberlin. (Grout’s Mill, Vt.). Fig. 5. Basal spines of left gonopod of 9, ventral view. Tidabius cantabrigensis (Meinert). (Type, Cambridge, Mass.). Fig. 6. Anterior portion of prosternum. Fig. 7. Basal spines, right gonopod of 9, ventral view. BULL. MUS. COMP. ZOOL. PLATE 6 LITHOBIIDAE, SP A gre OT Fann PLATE 7. CHAMBERLIN.— Lithobiid Genera. PLATE 7. Nadabius pullus (Bollman). Fig. 1. Anterior portion of prosternum. (Paratype, Bloomington, Ind.). Fig. 2. Basal spines of left gonopod of 9, ventral view. (Paratype, Bloomington, Ind.). Fig. 3. Portion of tibia and first tarsal joint of anal leg of co’, mesal aspect, showing tibial crest. (Lynchburg, Va.). Nadabius coloradensis (Cockerell). (Manitou, Col.). Fig. 4. Basal spines of right gonopod of 9, ventral view. Nadabius eigenmanni (Bollman). Fig. 5. Basal spines of left gonopod of 9, ventral view. Nadabius iowensis (Meinert). Fig. 6. Basal spines of left gonopod of 9, ventral view. PLATE 7 LITHOBIIDAE. BULL. MUS. COMP. ZOOL. eS Z PINT reyes Tm eee reer an, TSE en Sa PNP ean : or i = \ a eT eo re = vy. Pi a i ~ ee ata hte a a rb PLATE 8. CHAMBERLIN.— Lithobiid Genera. PLATE 8. Nadabius itowensis (Meinert). (Boone, Iowa). Fig. 1. Portion of tibia and tarsus of anal leg of <, mesal view, showing crest. Pokabius bilabiatus (Wood). (DeWitt, Iowa). Fig. 2. Basal spines of left gonopod of 9, ventral view. Pokabius bilabiatus verdescens Chamberlin. (Type, East Peoria, IIl.). Fig. 3. Femur and portion of the two contiguous joints of left anal leg of o' showing the characteristic lobes. Pokabius iginus (Chamberlin). (Washington). Fig. 4. Femur and portion of prefemur and tibia of right anal leg of &, dorsal view. BULL. MUS. COMP. ZOOL. LITHOBIIDAE. PLATE 8 PLATE 9. CHAMBERLIN.— Lithobiid Genera. PLATE 9. Pokabius clavigerens (Chamberlin). (Pacific Grove, Cal.). Fig 1. Femur and part of prefemur and tibia of left anal leg of , sub- dorsal view. Fig. 2. Basal spines of left gonopod of 9. Pokabius disantus Chamberlin. (Type, Laurel Canyon, Cal.). Fg. 3. Femur and portion of the two contiguous joints of the left anal leg of the o’, subdorsal view. Pokabius pitophilus Chamberlin. (Truckee, Cal.). i Fig. 4. Femur and portion of the two contiguous joints of the left anal leg of o’, mesal view. Fig. 5. Basal spines of right gonopod of ¢, ventral view. on BULL. MUS. COMP. ZOOL. PLATE 9 LITHOBIIDAE. PLATE 10. CHAMBERLIN.— Lithobiid Genera. PLATE 10. Pokabius centurio (Chamberlin). (Las Valles, N. M.). Fig. 1. Femur and portion of prefemur and tibia of anal leg of <, view a little ectad of dorsal. Fig. 2. Basal spines of left gonopod of 9, ventral view. Pokabius utahensis (Chamberlin). Fig. 3. Femur and portion of the two contiguous joints of the anal leg of 0’, subdorsal view. (Provo Canyon, Utah Co., Utah). Fig. 4. Basal spines of left gonopod of 2, ventral view (Little Willow Canyon, Salt Lake Co., Utah). Pokabius iginus (Chamberlin). (Paratype, Washington). ‘Fig. 5. Basal spines of left gonopod of 9, ventral view. BULL. MUS. COMP. ZOOL. LITHOBIIDAE. PLATE 10 PLATE 11. CHAMBERLIN.— Lithobiid Genera. PLATE 11. Pokabius gilae Chamberlin. (Thatcher, Arizona). Fig. 1. Femur and portion of prefemur and tibia of anal leg of <, dorsal view. Pokabius sokovus (Chamberlin). (Cotype, Las Vegas, Nev.). Fig. 2. Femur and portion of the two contiguous joints of anal leg of 0, mesal view. Fig. 3. Basal spines of right gonopod of 9, ventral view. Pokabius collium (Chamberlin). Fig. 4. Anterior portion of prosternum. (Antelope Is., Gt. Salt Lake, Utah). Fig. 5. Basal spines of left gonopod of 9. (Antelope Is., Gt. Salt Lake, Utah). Fig. 6. Femur and portion of the two contiguous joints of anal leg of ¢, dorsal view. LITHOBIIDAE. PLATE 11 BULL. MUS. COMP. ZOOL. Ee Weer OC Tt} PLATE 12. CHAMBERLIN.— Lithobiid Genera. PLATE 12. Pokabius castellopes (Chamberlin). (Shasta Springs, Cal.). Fig. 1. Femur and portion of the two contiguous joints of anal leg of @, lateral view. Pokabius socius (Chamberlin). (Provo, Utah). Fig. 2. Basal spines of left gonopod of 2, ventral view. Pokabius helenae Chamberlin. (Helena, Montana). Fig. 3. Basal spines of left gonopod of 2, ventral view. (Helena, Montana). Simobius ginampus (Chamberlin). (Pullman, Washington). Fig. 4. Portion of fifth and sixth joints of penult leg of <, dorsal view, showing the characteristic tibial process. Fig. 5. Basal spines of left gonopod of 2, ventral view. BULL. MUS. COMP. ZOOL. PLATE 12 LITHOBIIDAE. Bulletin of the Museum of Comparative Zodlogy AT HARVARD COLLEGE. Vou, LVIE. No. 7. THE ETHOPOLIDAE OF AMERICA NORTH OF MEXICO. By Raven V. CHAMBERLIN. Wits Six Piates. CAMBRIDGE, MASS., U.S. A.: PRINTED FOR THE MUSEUM. APRIL, 1925. No. 7.— The Ethopolidae of America North of Mexico. By Rauew V. CHAMBERLIN. Tue Ethopolidae (Chamberlin, Bull. M. C. Z., 1915, 59, p. 531) embraces a group of genera in the Lithobioidea characterized chiefly by having the coxal pores in several series. Bothropolys (B0@pos, pit, and roAvs, many) was proposed by Wood in 1863 for several American species having the characteristic men- tioned. Later Stuxberg (Ofvers. Kongl. vet. akad. Férh., 1875, 32, no. 3, p. 5-22) merged the group again with Lithobius which he sub- divided into subgenera according to the angulation of the dorsal plates, a system which, though manifestly in large part artificial, has been followed by nearly all subsequent students. Latzel in his work on Die Myriopoden der Osterreichisch-Ungar- ischen Monarchie (1880, 1, p. 35) regarded Wood’s grouping as natural. In speaking of it he writes:— er, was nur loblich ist, alle Arten der Gatt. Lithob. mit zahlreichen, ungeord- neten oder in unregelmassige Reihen vertheilten Hiiftporen unter dem gemein- samen Gruppennamen Bothropolys zusammenfasste. Leider ist daran doch zweierlei zu tadeln: einmal ist der Name Bothropolys entschieden falsch gebildet, da er richtig Polybothrus heissen sollte; dann hat Wood unterlassen, auch consequenterweise den Rest der Gattung zu benennen, der etwa mit dem Namen Oligobothrus zu belegen gewesen wire. Ich bin geneigt zu glauben, dass diese Zweitheilung der Lithobius-Arten im Sinne Wood’s sehr natiirlich ist, da man beziiglich der Hiiftporen nie, beziiglich der Zahnbildungen an den Riickenschilden, besonders beim Subgenus Neolithobius und Hemi- lithobius leicht im Verlegenheit gerath. Die Gruppe Polybothrus umfasst bisher nur (18 bis 45 mm) grosse Formen, die sich auch durch zahlreiche (namlich 12-20) Hiiftziihne auszeichnen. In die Gruppe Oligobothrus gehéren Arten, deren K6rperlinge von 31 mm bis unter 8 mm heruntersinkt, und deren Hiiftzahne meist in der Zahl 10-16 auftreten, oder andererseits fast vollig verkiimmert sind. Ferner haben die Arten der Gruppe Polybothrus in der mannlichen Form gut entwickelte Genitalanhinge, wihrend die Mannchen der Gruppe Oligobothrus stets, bis auf 2 kleine behaarte Wiarzchen, verkiimmerte Genitalanhainge haben. But while thus recognizing the naturalness of Wood’s system and urging objections to that of Stuxberg, Latzel adopts the latter through- out “wegen ihrer sonstigen Bequemlichkeit,’’ a procedure which it seems difficult to justify. Of course the suggestion of a change from Bothropolys to Polybothrus on etymological grounds cannot be fol- 386 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. lowed; while the name Oligobothrus was wholly unnecessary as its function was already filled by Lithobius sens. str., whether the groups were to be recognized as genera or as subgenera. In 1896 Brolemann (Ann. Ent. soc. France, 65, p. 45) recognized Bothropolys as a subgenus but unnecessarily used Latzel’s Oligo- bothrus for Lithobius sens. str. In 1901 Pocock (Ann. mag. nat. hist., ser. 7, 8, p. 448) considered Bothropolys as proposed by Wood a valid genus, and Verhoeff (Bronn’s Thierreich, 1907, 5, p. 234 et seq.) also accepts it and at the same time separates from it the European species as a distinct genus p. 237 :— k. Antennen fast konstant aus 20 Gliedern bestehend-(selten 17 bis 23). Nordamerikanische und orientalische Formen.... Bothropolys. 1. Antennen aus 30-80 Gliedern bestehend. Paliarktische Formen. Polybothrus. Polybothrus is attributed to Latzel; but manifestly this name can- not be thus used, inasmuch as it was proposed, as seen above in the passage quoted from that author, strictly as an etymological reforma- tion of and substitute for Bothropolys and must be wholly eliminated from use as a synonym of the latter. Verhoeff divides Polybothrus into subgenera by applying the Stuxberg system; but unfortunately he lists no species for any of these groups and the names given are therefore difficult to apply and also really without standing. We may, however, use the name Eupolybothrus for the European genus of which Lithobius grossipes C. Koch may be specified as the type. It is quite possible that there will be found to exist among the Eurasian species other genera. In North America the species of the Etho- polidae fall into three genera. It will be noted that the American gen- era differ from the European not only in the smaller number of antennal joints, a character not applicable in younger specimens, but especially in the presence of a stout ventral spine on the posterior coxae. The latter character is developed in very young specimens. ‘This spine is present while the coxal pores are still uniseriate; and as a ventral coxal spine is present in no other American genera excepting the rare Zinapolys, its presence may serve in separating the young of these genera from the other lithobioids (cf. Can. ent., 1911, 43, p. 379). Among European Lithobiidae, the species Lithobius dalmaticus Latzel also has the coxae of the anal legs, though not of the penult, similarly armed with a stout ventral spine. This species is evidently generically distinct from the American Zinapolys and may stand as the type of a new genus, Abrotobius. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 387 Key to Genera of Ethopolidae. a. Posterior coxae unarmed ventrally; articles of antennae 30 or more. Eupolybothrus, gen. nov. aa. Coxae of last one or two pairs of coxae armed ventrally with a stout spine; articles of antennae normally 20 (rarely varying to as many as 27). b. Lateral margining of head continuous, not interrupted or broken (Plate 1, fig. 1); basal spines of 2 gonopods 2+2. ce. Prosternum with a distinct diastema in dental line on each side in which the spine is inserted; only anal coxae ventrally armed. Zygethopolys, gen. nov. cc. Prosternum without diastema in dental line, the spine inserted at each anteroectal angle; last 2 pairs of coxae ventrally armed. Bothropolys Wood. bb. Lateral margining of head discontinuous or interrupted laterally (Plate 4, fig. 1); basal spines of 9 gonopods 3 + 3 or 4 + 4. Ethopolys Chamberlin. Among other characters presented by Eupolybothrus it may be mentioned that the gonopods of the male are biarticulate as in Etho- polys; that no diastema is present on the prosternum; and that in most the claw of the gonopods of the female is entire and the basal spines 2 + 2; margin of head discontinuous as in Ethopolys. Exami- nation of European species not accessible may reveal other genera. The genus has not been found in America. The North American species of the Ethopolidae with the single exception of B. multidentatus (Newport) are found in the region west of the Rocky Mountains. In this western region they oceur abun- dantly from Mexico to Alaska and in most parts of this territory are the largest of the Lithobioidea. East of the Rocky Mountains, and par- ticularly in the southeastern states, the Ethopolidae are replaced by the large species of Lithobius proper which are comparatively rare, or in many parts wholly absent from, the western states. Boruropotys Wood (emend. auct.). Journ. Acad. nat. sci. Phil., 1863, new ser., 5, p. 15. Trans. Amer. philos. soc., 1865, 13, p. 152. Verhoeff, Bronn’s Thierreich, 1907, &, p. 24. Chamber- lin, Can. ent., 1912, 44, p. 173. Polybothrus Latzel, Myr. Osterreich.-Ungar. monarch., 1880, 1, p. 35. Marginal thickening of head continuous forward to eyes, not inter- rupted or broken at sides (Plate 1, fig. 1). Antennae short, mostly not reaching caudad of the seventh segment; 388 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. composed of twenty articles, or occasionally of as many as twenty- four. Prosternal spine well chitinized, situated at or near each anterior angle ectad of all the teeth, the latter being more or less uniformly spaced with no diastema separating them into two groups on each side (Plate 2, fig. 2, 3). Claw of anal legs single; that of the penult legs either single or with a single accessory aces minute or obsolete. Ventral spines of anal legs normally 1, 1, 3, 2, 1. Coxae of last two pairs of legs each bearing a stout ventral spine as well as being armed laterally and dorsally. Gonopods of male uniarticulate. Gonopods of female with the claw tripartite, the lateral divisions being situated but little proximad of the median, the three often on a level or nearly so; basal spines 2 + 2, or, occasionally, 2 + 3. Typr.— Bothropolys multidentatus (Newport). In the species of this genus there is particularly well marked on each side of the head a semicircular submarginal impression (line of muscle attachment), the anterior end of which is near the eye, the caudal end curving mesad and appearing as a short transverse sulcus at about one third the length from the caudal margin. There is but little variation in the spining of the legs from species to species within the genus. The dorsal spine of the anal legs are um- formly 1,0,3,1,0; the ventral 1,1,3,2,1, rarely increased to 1, 1, 3, 3, 2 or falling to 1, 1, 3, 2,0 on one side; claw single or with a slight point representing an accessory claw. Dorsal spines of the penult legs 1, 0, 3, 1,1; ventral, 1, 1, 3, 3, 1, rarely 1, 1,3, 3,2; claw one, or with a single ACCESSORY, claw. Dorsal spines of thirteenth legs 1,0, 3,1, 1 toy 15,0;3,.2;.2;\ ventral uniformly 0,15°3;.3;.2. 9 Dorsal spines of twelfth legs: 1,0; 315dh-to: 15053325 2;\- ventral ae 0, 1,3, 3,2. Dorsal spines of Re enth legs 0, 0, 3, 2, 2 or 1,0, 3,,2;.2: ventral 0, 0, 3, 3, 2 or 0, 0, 2, 3, 2. Dorsal spines of first and second legs 0, 0, 3, 2, 1 or 0, 0, 3, 2,2; ventral always 0, 0, 2, 3, 2. The prosternal spine in the immature stages is attenuated and dis- tally slender and bristle-like (Plate 2, fig. 3); but in the great majority of specimens, the slender apical portion is lost in the adult, leaving the spine blunt at tip and proportionately short (Plate 1, fig. 6, 7 and Plate 2, fig. 2). In variations in the claw of the female gonopods resulting in partial or complete obliteration of teeth or lobes, the result is due to an appar- ent filling in of the short incisions normally present between the teeth. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 389 — This gives a distally broad, subtruncate claw (Plate 2, fig. 7; Plate 3, fig. 1); cf. further under Ethopolys. In the known species of the genus as here restricted, the posterior angles of the ninth, eleventh, and thirteenth dorsal plates, or of those together with those of the sixth and seventh, are produced. However, in Bothropolys as conceived by Verhoeff, there is greater variation in this respect, species with the angles of more of the dorsal plates or with those of the fourth, sixth, seventh, ninth, eleventh, thirteenth, and fourteenth, produced being also included. Verhoeff (Bronn’s Thier- reich, 1907, 5, p. 240) proposes a purely artificial division of the genus, which he attributes to the North American and Oriental regions, as follows:— a). Tergite ohne Fortsitze am Hinterrande. Coxosternum der Kieferftisse mit 6 +6 Zihnchen. 7-9 Ocellen jederseits. Probothropolys n. subg. 8). 9., 11., 13. Tergit hinten mit Fortsitzen. Coxosternum der Kiefer- fiisse mit 7 +7 bis 10 + 10 Zahnchen. 10-21 Ocellen jederseits. Allobothropolys n. subg. y). 6., 7., 9., 11., 18. Tergit hinten mit Fortsitzen. Coxosternum der Kieferfiisse mit 6 + 6 bis 9 + 9 Ziihnchen. 19-35 Ozellen jederseits. Eubothropolys n. subg. 6). 4.,6., 7., 9., 11., 13. und 14. Tergit hinten mit Fortsitzen. Coxoster- num der Kieferfiisse mit 7-8 Zihnchen. Ocellen ca. 30 jederseits. Telobothropolys n. subg. Of the characters used in this key, those drawn from the prosternal teeth and eyes are, as given, wholly worthless since not even the species can be separated by them. This leaves only the one character of the angulation of the dorsal plates, the exclusive use of which gives an artificial grouping, species of really different genera, ¢.g., of Bothropolys sens. str. and Ethopolys being brought together and at the same time separated from others truly congeneric with them. It will be seen that Verhoeff ignores the rule that the subgenus containing the type of the genus must bear the name of the latter. However, under any conditions, his subgeneric names have no standing, since he lists no species under any of them. Because of their composite character it would be difficult to apply them with any certainty; and it therefore seems the only course open is to regard them as nomina nuda in accord with a strict application of the rules of nomenclature. Of the two groups into which the North American species of Bothropolys as here restricted fall, Bothropolys sens. str. must be applied to that containing B. multidentatus (Newport), while the second may stand as Poropolys, subgen. nov., with B. permundus Chamberlin as its type. 390 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Key to Species of Bothropolys. a. Posterior angles of the sixth and seventh dorsal plates not produced. Poropolys, subgen. nov. b. Ninth dorsal plate with the posterior angles only very slightly pro- duced, those of eleventh and thirteenth with processes small. Body near 9.5 times longer than width of tenth plate. B. victorianus, sp. nov. bb. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced. Body mostly from 7 to 8, or but little more, times as long as width of the tenth plate. c. First dorsal plate of the male as wide as the eighth and as wide as or mostly a little wider than the tenth; ocelli 27 to 37 in mostly 5 series; length of maturus from 23 to 40 mm...B. hoples Brolemann. cc. First dorsal plate of the male distinctly narrower than the eighth and tenth plates; ocelli 15 to 25, mostly in 3 or 4 series; length of GMS WE) AY WW 5 Sooo coat acsos B. permundus Chamberlin. aa. Posterior angles of the sixth, seventh, ninth, and eleventh dorsal plates PLOCUCE AY fe eirs co Motes Ree ae NE SEN Bothropolys sens. str. b. Head wider than long; prosternum about 1.8 times wider than long; body not more than eight times longer than the width of the tenth POLAT Aes race uc ae ee ae eee eee B. multidentatus (Newport). bb. Head as long as wide or slightly longer; prosternum between 1.5 and 1.6 times wider than long; body nearly nine times longer than the WACO Woe ten NOs 6 stunceslooeoadese B. columbiensis, sp. nov. BOTHROPOLYS sens. str. BOTHROPOLYS MULTIDENTATUS (Newport). Plate 1, fig. 1-8. Lithobius multidentatus Newport, Trans. Linn. soe. London, 1845, 19, p. 365. Gervais, Hist. nat. ins. Aptéres, 1847, 4, p. 236. Newport, Cat. Myr., 1856, p. 17. Chamberlin, Ann. Ent. soc. Amer., 1909, 2, p. 190. Bothropolys nobilis Wood, Journ. Acad. nat. sci. Phil., 1863, new ser. 5, p. 15. Bothropolys multidentatus Wood, Trans. Amer. philos. soc., 1865, new ser., 1865, 13, p. 152. Chamberlin, Can. ent., 1911, 48, p. 98. Ann. Ent. soc. Amer., 1911, 4, p. 48. Can. ent., 1914, 46, p. 301. Psyche, 1918, 25, p. 24. Lithobius (Eulithobius) multidentatus Stuxberg, Ofvers. Kongl. vet. akad. Forh., 1875, 32, no. 3, p. 10, 26. Bollman, Proc. U. 8. N. M., 1887, 10, p. 263. DescripTion.— Dorsum varying from light yellowish brown, with the first and last several segments often darker, through chestnut to CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 391 Fic. 1.— Distribution of Bothropolys Wood. deep mahogany in specimens in full color. The head varies from reddish brown or chestnut to almost black. Venter but little paler than dorsum. Legs like venter, but tarsi distinctly paler, in the last pair, especially, usually distally rufous with the tibia also often affected. 392 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae mostly concolorous with head, distinctly rufous distally. Prosternum a little paler than head, the prehensorial feet paler, in darker specimens rufous especially ectally and distally. Body attenuated caudad, but only slightly so cephalad, the sides being nearly parallel from the head to the tenth plate. Body from seven to eight times longer than the width of the tenth plate. Head much wider than long, the ratio usually near 7:6.5; widest just caudad of eyes, the sides from these first running nearly parallel, converging but slightly, and these over the caudal third of length con- verging a little more strongly to the rounded caudal corners; caudal margin mesally nearly straight, curving forwards on each side. Mod- erately roughened. Ocelli arranged in an elongate area in which mostly five to seven, often irregular, series can be detected, from twenty-five to forty in number; e¢.g., 1+ 4, 6,7, 6, 5,4; 1-+ 4, 5, 6,6,5,4. Single ocellus large, subcircular. Antennae short, usually reaching to about the middle of the seventh segment. Articles typically twenty, varying also, on the one hand, to nineteen, and, on the other, to twenty-three or twenty-four; all articles long and differing slightly in proportions, or, sometimes, a number of those immediately preceding the ultimate shortened, the ultimate usually long and slender but also sometimes shortened. Articles densely clothed with moderately long hairs, less densely so proximad. The prosternum mostly near 1.8 times wider than long, being thus unusually wide. Mesal incision rather deep, its sides subparallel, rounded at bottom. Teeth uniform in size and spacing, mostly from 7+ 7to9+9innumber. Spine stout, in younger specimens always drawn out to a slender acute apical portion, but in older ones this slender apical portion is commonly lost, leaving the spine blunt distally and proportionately short (Plate 1, fig. 7). The form of the spine in younger specimens or occasionally even in large adults is shown (Plate 1, figs. 8). Dorsal plates not much roughened, the caudal ones more so than the anterior. The usual submarginal transverse sulcus on the third, fifth, seventh, eighth, tenth, twelfth, and fourteenth plates extending from margin mesad or somewhat obliquely mesocephalad a little back of middle of length. Posterior angles of sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced, the processes of the sixth sometimes evenly rounded distad rather than acute, and the inner side of those of the seventh sometimes long, the angulation of these two plates varying considerably. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 393 Typically the first ventral plate is broadly mesally depressed; the other plates show three impressions, a median and a lateral on each side, the latter usually farther cephalad and often replaced by a pair of impressions of which one lies caudad of the other; there these im- pressions in some specimens pit-like, with the median one tending to elongate transversely on the anterior plates and longitudinally on the posterior ones; often all three impressions take on the character of longitudinal furrows, of which the median one is especially conspicu- ous on the more caudal segments. In the male the lateral furrows on the fourteenth plate converge caudad and are united by a transverse impression immediately in front of the smooth caudal portion, but the median furrow often causes this impression and extends to or nearly to the caudal margin; the lateral furrows on the fifteenth plate also united transversely but less converging than those of the fourteenth; the smooth pilose border, especially laterally, less sharply defined than in Ethopolys and the last two plates less abruptly differentiated from those preceding it. In the female the lateral furrows of the fourteenth and fifteenth plates are either not evident or weakly impressed, and more widely separated and less converging than in the male; but a transverse impression is sometimes conspicuously developed. Pos- terior plates in male shortly subdensely pilose especially on borders of last two; less as in the female. Coxal pores numerous, arranged in three to five series, which are commonly irregular. Pores mostly small, increasing in size as usual from most anterior series to those of most posterior in which they are of moderate size and often elliptical in shape. Spines of first and second legs, 0 ees} 0 of the third to ninth, 00 2 2,2 of the tenth: and eleventh, oye go OF us ees; of the twelfth, (3372, 1, 0, 3, 2,1 1, 0,3, 2,2 0133-2, OF o1:3-3-2, the first formula being commonest; of the thir- teenth the same as of the twelfth excepting that the ventral spines 103.11 OAs ct may be 0, 1, 3,3, 1; of the penult, 773-34, rarely {73'3'5, the claw ib 0, 3, 1, 0 armed with a very Sill accessory aay: of the anal legs, 74°3°31, rarely 0, 3, 1, 0 : 5 to 332, the claw single or else with an accessory claw Pereeenicd by a minute or obsolete point. Last two pairs of coxae laterally armed. Anal legs in female slender, rather short. In the male they are somewhat thicker; tibia a little clavately thickened distad and at the distal end the dorsal surface is raised into a slight rounded elevation which is more densely pilose; the first tarsal joint is also similarly more densely pilose on the dorsal surface at the distal end than elsewhere and sometimes is weakly clavate. The femur somewhat thicker than in the female and also clavately widened distad. 394 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Gonopods of female with claw but moderately bent; the two lateral lobes but little proximad of level of median; occasionally one of the lateral lobes may be replaced by two so that there are four divisions instead of the usual three. The outer of each pair of spines consider- ably larger than the inner and often bifid distally; spines subevlindric with acuminate distal portion short and commonly blunt; the inner spine may be bent mesad at tip. For appearance of gonopods of male, see Plate 1, fig. 2 Length 15 to 27mm. The antenna in specimens from 24 to 26 mm. long is from 11 to 12 mm. long, the anal legs exclusive of coxa being from 9 toll mm. long. A 2 26 mm. long has the tenth (and eighth) dorsal plate 3.8 mm. wide; a o 19 mm. long has the tenth (and eighth) dorsal plate 2.8 mm. wide. Immaturus.— Light brown in life, often having something of a vio- laceous tinge. Head light reddish brown or chestnut. Antennae and last pair of legs pale, vellowish distad. Antennae with the twenty articles present, of nearly same propor- tions as in adult. Eyes composed of seventeen to nineteen ocelli in 4 series; ¢.g., INE sd, OMA Ze Prosternal teeth 6 + 6. Spines slender and pale. Chitinous lines distinct, complete or nearly so. Three joints of female gonopods present, the claw indicated merely as a chitinous point to the distal joint. Second joint with two or three bristles, the first with from five to seven, the last with one or two Coxal pores small, mostly in but two series, those of caudal row largest, six to twelve in number. — Spines of first legs, 5 i332 ; spines of second legs, {s°3°4"3-4 ; of third tosev enth, 95 0° BS 3 i, of eighth to tenth, 5 2 3: a3 the eleventh, ne: of the twelfth, u a - 233 of the thirteenth, (7 3. 5; of the anal, 7 r et Ventral spines of last two pairs of coxae strongly developed as also the lateral and dorsal. Length of specimen described, 12 mm. Agenitalis (o").— Light violet-brown. Head more reddish. An- tennae and last legs paler, yellowish. Prosternal teeth 5 + 5. Spines pale, slender. Ocelli about eight in two series; e.g., 1 + 4, 3. The twenty articles of antennae already present, varying in length, the shorter ones occurring at intervals. Angles of dorsal plates produced as in adults, but the processes of the sixth weaker. 9; of 1, 0, 3, 1, ne Onuegegs 3,2 9 Sy CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 395 Coxal pores on anal coxae two in one series; on others in two series, four or five in number, three of caudal or ventral row largest as usual. Gonopods small, wart-like, glabrous. 0, . 0, 0,1, 1 . 00111 0, 0, 2,21 Spines of first legs, -¢t@- i; of second, 5 1; of third, 9 0°3°11; , 0, 0,1, 2.1 NCE ava of fourth, j-o-323; of fifth to ninth, ? 00-2213 “ tenth and eleventh, 0, 0,3, 2,2 Chinese tele yas italy Toei; of twelfth, pyse73 of thirteenth, tiie21; of fourteenth, AO atel 1.0.3.1), 0 aly Sh 2a of anal, 11,320 ° Length of specimen described cir. 9 mm. Type Locatity.— New York. Known Locauitres.— Massachusetts (Warwick!). New Hamp- shire (Marlow). New York (Ithaca! Penn Yan! Albany! Thousand Is.!). Ohio (Columbus! Kelly’s Island!). Indiana (abundant throughout the state). Michigan (Ludington, Ann Arbor! Lansing! Douglas Lake!). Illinois (Franklin Grove! Peoria!). Pennsylvania (Philadelphia!). New Jersey!. District of Columbia ieshington’): Maryland. Delaware. Kentucky (near Mammoth Cave! Osceola! Greensburg!). Tennessee (Beaver Creek, Mossy Creek, Nashville! Knoxville! Russellville!). Virginia (Machaville, Natural Bridge! Chatham! Balcony Falls!). West Virginia (White Sulphur!). North Carolina (Catawba! Chapel Hill! Morgantown!). South Carolina. Georgia!. Florida (Palatka!). Alabama (Maplesville! Jackson!). Mississippi (Canton! Fernwood! Byram!). Arkansas (Little Rock!). Louisiana!. In this widespread species, considerable variation occurs. Varia- tion in the spining of the legs is apparently more frequent than in related species, as noted particularly on the twelfth and thirteenth pairs where an extra or caudal spine occurs on the fourth or on the fourth and fifth articles raising the dorsal formula to 1,0, 3, 2, 1 or 1, 0, 3, 2, 2 in place of the typical and much more frequent 1, 0, 3, 1, 1. Specimens collected at Franklin Grove, IIl., have a considerably larger proportion with the increased number of spines on these legs than in those from other localities studied, this variation in most places appearing to be rare. BOTHROPOLYS COLUMBIENSIS, sp. nov. Descrietion.— Dorsum brown, with the caudal plates light fer- ruginous. Head also of pale ferruginous cast with the frontal region lightest. Antennae pale brown, lightest distally. Prosternum pale ferruginous, with the prehensors lighter, more yellowish. Venter brown, the caudal plates light ferruginous. Legs yellowish brown, brighter distad; caudal pairs light ferruginous. 396 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Body rather slender, being nearly nine times longer than the width of the tenth plate. Ratio of width of head to that of first, third, eighth, and tenth dorsal plates as 44 : 40 : 39 : 44 : 44. Head slightly longer than wide (45:44); widest immediately caudad of the eye, the sides incurved at level of suture and then mod- erately converging caudad to the well-rounded corners. Caudal margin straight or nearly so. On caudal region a narrowly V-shaped impression with the apex caudad and more or less extended as a median sulcus; ectad of each arm of the V-shaped impression are two or three longitudinal sulci. A V-shaped impression on frontal portion of head with the point a little in front of suture and the arms diverging cepha- lad A weaker transverse constriction extending straight across frontal region from near lateral ends of frontal suture Eyes composed of sixteen ocelli in three series: thus, 1 + 6, 5, 4. The single ocellus not very large. Ocelli well spaced, regular, decreas- ing in size cephalad and ventrad as usual. Antennae short, reaching to the seventh segment. Articles twenty, moderate in size and uniform. Prosternum in type 1.54+ times wider than long. Distance be- tween chitinous spots 1.9 times width at mesal incision, 2.53 times the dental line. Mesal incision moderate V-shaped, with the sides some- what flaring. Teeth of moderate size, nearly uniform, the outer ones a little larger and a little more widely spaced; 6 + 6 in number. Spine moderately stout at base a little above which it is constricted then running out into a fine bristle-like distal portion. First dorsal plate widest at anterior end with the sides moderately converging to the rounded caudal corners; 1.8 times wider than long; with two pairs of longitudinal furrows ending about one third the length of plate from caudal margin; a weak transverse furrow near caudal ends of longitudinal ones. Other plates mostly with two sub- median longitudinal sulci and each side of these two deeper furrows which converge caudad and commonly meet at an acute angle. Pos- terior angles of sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced, but those of the sixth only weakly so, those of the seventh more distinct but with the mesal edge long. Coxal pores in three or four series of which the most caudal is long- est and composed of largest pores as usual. 2 a . . . . Spines of first and second legs, ¢'t'2'3'2; of third to ninth inclusive, 0.0, 3. 2 0, 3, 2,2 . 1 (0), 0, 3, 1,1 00,2, z >; of tenth and eleventh, Aan “33-3; of the twelfth, Se Sao 0. OS sel sel. 0,3, 1,1 the thirteenth, O13 323 of the penu ult, ¢4'3-3-1, a very small accessory 1, 0 claw present; of the anal, ¢3:3:5:], the claw single. Last two pairs of coxae laterally armed. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 397 Anal legs in female of moderate length, slender, dorsally longitudi- nally suleate. Claw of female gonopods tripartite as usual, the lobes of usual relations. Basal spines conical, somewhat rounded at tips. Length 15 mm.; length of antennae 6.6 mm.; of anal legs 6 mm.; width of tenth plate, 1.68 mm. Locatity.— British Columbia (Kaslo!). The type (M. C. Z. 672) isa female in the late pseudomaturus stage. This species is very close to multidentatus (Newport) but differs es- pecially in proportion of body, head, and prosternum. The head is longer than wide whereas in the other species it is conspicuously wider than long; and the prosternum is but 1.54 times wider than long as against 1.8 times in multidentatus. The coloration of the two species is different. PoROPOLys, subgen. nov. BoTHROPOLYS HOPLES (Brolemann). Plate 2, fig. 9. Plate 3, fig. 1-4. Lithobius (Bothropolys) hoples Brolemann, Ann. Soe. ent. France, 1896, 665, p. 45. Lithobius hoples Chamberlin, Ann. Ent. soc. Amer., 1909, 2, p. 191. Bothropolys hoples Chamberlin, Can. ent., 1912, 44, p. 173. DerscripTion.— Dorsum from brown, with the head and the first and last segments reddish, to light chestnut in specimens in full color, the head being then not at all or but little darker. A median longi- tudinal stripe with on most of the major plates one or two fainter somewhat oblique marks each side of it testaceous, these light lines extending about two thirds the length of the plate; caudal borders and minor plates often also testaceous. Antennae brown to light chestnut proximally, becoming yellow distad. Prosternum and pre- hensorial feet colored like the head. Venter and legs testaceous, the last segments darker, often chestnut, the anal pair of legs also darker, in fully colored specimens light chestnut. Robust, being from 7 to 8.25 times longer than width of tenth plate. The first dorsal plate in the male is as wide as or mostly a little wider than the eighth and tenth, but in the female it is always distinctly narrower; the ratio of width of first, eighth, tenth, and twelfth plates is typically near 68 : 67 : 67:64 in the male, and 64 : 68 : 68 : 66 in the female. 398 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. The head is wider than long in about the ratio 5 : 4.5. Sides nearly straight from immediately back of the eyes to the rounded caudal corners, slightly converging; caudal margin straight between the rounded corners or mesally very weakly incurved. HOS ai0 anal, j-7°3°3'1, Claw single. ono S\es2 - ) By De ) By By Bye |, By By fd keno RS 400 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae composed of the full twenty articles which are mostly uniform and relatively somewhat shorter than in the adult. Prosternal teeth 7+ 7. Spine stout at base, running out to a bristle-like apical portion. Spines of first legs, P23; of the second Toa; Of the thindeto ninth, 3 0.3, ae off the tenth and elev enth, 5 ae 133-5; of the twelfth and thirteenth, i T3332; of the penult ¢q-334; of the anal, p>raa@a- Coxal pores in two or three series, ten to fifteen in number, those of most anterior row commonly very small. Claw of female gonopods very short and pale; but slightly bent; tripartite, the lateral teeth weak and much shorter than the acute median division. Basal spines 2 + 2, but the inner one of each pair much more slender than the stout conical outer one and only about half as long. Length cir. 15 mm.; length of antennae 7 mm.; of anal leg cir. 6.5 mm. Tyre Loca.tiry.— Washington. Known Locauities.— Washington. Oregon (Portland!). British Columbia (Kaslo!). Montana (Flathead Lake!). Idaho (Kootenai Co.!). This large species is common in the Northwest. The marked sexual dimorphism shown in the proportions of the body is very interesting and is not paralleled in any other known North American species either of Bothropolys or of Ethopolys. As indicating the extent to which the antennae may occasionally vary, it may be mentioned that in a specimen from Idaho the left antenna reaches only to the fifth segment, while the right one extends to the eighth segment, the number of articles, nevertheless, being the same on the two sides. BoTHROPOLYS PERMUNDUS (Chamberlin). Plate 2, fig. 1-8. Lithobius permundus Chamberlin, Proc. Acad. nat. sei. Phil., 1902, p. 42. Lithobius xanti Chamberlin (ex parte), Proc. U. S. N. M., 1901, 24, p. 24. Bothropolys permundus Chamberlin, Can. ent., 1912, 44, p. 173. DescripTion.— Dorsum brown, the head with the last few and also often the first or first few dorsal plates rust-brown or ferruginous; middle portion of dorsum sometimes very dark, dusky, and part of the plate often with a median longitudinal blackish stripe with some- CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. AO] times, especially on the more caudal plates, a pale oblique line on each side. Legs brown to testaceous, the tarsi, as usual, more brightly colored, light orange; last pairs of legs light orange in color. Antennae brown to orange, usually lighter distad. Venter brown, with the last several plates ferruginous. Prosternum and prehensorial feet pale ferruginous. Body from 7.6 to 8 times longer than the width of the tenth plate. Widest as usual at eighth and tenth plates; narrowed conspicuously caudad and but little cephalad; the first plate in both male and female clearly narrower than the eighth and tenth. The width of first, eighth, tenth, and twelfth plates in a male are to each other as 58 : 63 : 63:60; ina female as 61 : 62 : 62 : 62, the female being more uniform. Head wider than long in about ratio 13:12. Of usual shape; sides converging considerably from immediately back of eyes caudad. A rather sharply impressed curved furrow a little caudad of and parallel with the frontal suture, this furrow not extending over the lateral portions of head. The usual short transverse furrow over each lateral border at beginning of caudal third of length. Head polished, the puncta few and scattered. Eyes composed of fifteen to twenty-five ocelli in three to five series. Single ocellus of moderate size, subcircular, typically contiguous with first one of dorsal row. Seriate ocelli small, compactly grouped. Examples of arrangements are 1 + 6, 5,3; 1+ 5,5, 4,3; and 1 +6, 5, 4, 3. Antennae usually reaching to the seventh segment. Composed of from twenty to twenty-three articles, all of which are moderately long and do not vary much among themselves in proportions. Hairs moderately long. Prosternum near 1.6 times wider than long. Chitinous lines very faint. Mesal incision with sides considerably converging, the sinus at bottom narrowly rounded. Teeth nearly uniform in size and spacing, or the outer ones more widely separated than the inner. Teeth 7+ 7 to 9+ 9. Spine of usual acuminate form, the bristle- like tip commonly lost in adults leaving the basal portion bluntly tipped. Dorsal plates polished, not rugose being roughened only by the fur- rows mentioned below. The first, third, fifth, eighth, tenth, twelfth, and fourteenth plates with the usual short submarginal sulcus on each side near beginning of caudal third of length, the seventh with similar _ impressions at each caudal and each anterior corner. Each major plate excepting the first or the first and third with a median and on 402 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. each side a lateral longitudinal furrow which are least distinct on the anterior plates; often also an oblique furrow on each side running from near the anterior end of median furrow ectocaudad and meeting the outer longitudinal furrow at an angle. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced, those of sixth and seventh obliquely truncate or excised. Ventral plates with the usual three longitudinal furrows which in- crease in distinctness from anterior to posterior regions and may be almost obliterated on first plates; the median impression on the more anterior segments is often pit-like; a transverse furrow a little in front of caudal margin may or may not be evident. Typically in the male the three furrows on the last three plates extend distinctly across the plate, the two lateral converging and meeting upon the median one in an acute angle in front of the caudal end of the latter or else connected across it by a transverse furrow. Coxal pores varying in size as usual with those of the most caudal series and at the distal end largest; arranged in three or four series and mostly from twelve to twenty-seven in number on each coxa. Spines of first to fifth legs, te 39 of sixth and seventh, wees ; of the eighth and ninth, 9 osas ; of the tenth, 1 0375 ; of the elev enth, it US a4 1, 0, 3, 2, 2 ILS eae al iE he Bir Coes OF Gee 25 of the twelfth and thirteenth, 0) 73°3'9 or 0°4°3°3°3 3 1 053,100 of the penult, ¢ 73°33, with one small accessory claw; of the anal, 1, 0, 3, 1, 0 T1321, Claw 1. Last two pairs of coxae laterally armed. In the female the anal legs are of medium length and slender. In the male the fourth, fifth, and sixth articles are somewhat more thickened, the sixth or first tarsal joint being commonly conspicuously thicker than the seventh joint; the fourth and fifth joints are slightly complanate above and more densely clothed with the fine short hairs on dorsal side toward distal end. Gonopods of male consisting of one subcylindric, distally rounded or sometimes obliquely subtruncate article which bears seven or eight bristles. Claw of female gonopods of moderate length, not strongly curved, tripartite, with the lobes nearly equal in length and form. Basal spines subuniformly acuminate from very base the apical portion longer and more slender and more acute than usual. Outer spine largest (Plate 2, fig. 8). Length from 19 to 27 mm. Ina female 20 mm. long the antennae are 9.8 mm. long; the anal legs, exclusive of coxa, 8 mm.; width of first dorsal plate 2.4 mm.; of the tenth plate 2.6 mm. Pseudomaturus.— Dorsum brown with margins of major plates often CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 403 darkened; head and caudal segments light ferruginous, the former somewhat dusky caudad of the frontal suture. Antennae brownish or somewhat ferruginous proximally, yellowish distad. Prehensorial feet pale orange, the prosternum a little darker. Venter light brown, the last few segments subferruginous. Legs light brown, pale orange at tips. Last pair of legs light orange excepting proximad. Ocelli fewer in number than in maturus but of same proportion and arrangements such as | + 5, 5, 4. Antennae composed of twenty to twenty-two articles; when of the larger number some of the more distal articles short, apparently from subdivision of larger articles; when twenty moderately long and decreasing in length distad from the second to the penult. Prosternal teeth mostly 6+ 6. Spine as in adult, but tip more usually present. Chitinous lines distinct. Coxal pores eight to fourteen in two or three series, the more cephalic ones very small or some minute. Spines of first legs, ¢ 0 ae of the eighth and ninth, ()}: Ousci ct bd to Nobo bo , the row bo 3; of the tenth and eleventh, of the twelfth, 9-7°3'3'5 or 4 313 of the thirteenth, a t > 3 50 penult, 7 : - z i; of the anal, an » ie Gonopods of female nearly as in adult, but claw less strongly chitin- ized; of about equal width distad and proximad; teeth appearing as crenations of about same length, but middle one broader. Spines nearly as in adult but the inner proportionately to the outer one a little more reduced. Length 14 to 17 mm. long. A female 15.5 mm. long has antennae 6.5 mm. long, and anal legs 7 mm. long. Praematurus.— Light brown commonly of more or less violaceous tinge, head light ferruginous, the caudal segments more yellowish. Prosternum and prehensorial feet and the most caudal ventral plates yellowish or weak orange tinge. Antennae yellow distad. Legs pale, the caudal pairs yellow or yellowish orange. Ocelli 1 + 5, 3,2; 1+ 5, 4,2. Compact, the single one contiguous, not much different in size. Articles of antennae already twenty, a little shorter than in adult. Prosternal teeth 5 + 5, uniformly spaced, lips narrowly rounded, black. Spine long, bristle-like at tip. Chitinous lines very distinct. Coxal pores in two rows, those of the anterior one often relatively very small or minute, the distal pore of the caudal row being much the largest. Spines of first and second legs, ())'33'2; spies of third to eighth . 404 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Ooo ee 0, 0,3, 2, 2 . . pairs, 9023.23 Of ninth to eleventh, o'9,3°3'2; of the twelfth, and thir- 10,34, TOSI 1,0, 3, 1, 0 teenth, (3-333; of the penult, ¢ 334; of the anal, Ppse4. Gonopods of male appearing as pale rounded processes bearing two or three bris‘!es. Claw of femnsle gonopods brown, short and thin; the middle lobe clearly longer than the lateral ones, acute; the lateral teeth small, acute, and toward base of claw. Basal spines 2 + 2, with the inner one of each pair much shorter and more slender than the outer. Length of a female 14 mm., with anal legs 5.5 mm., and antenna 4 mm. long. A male is 12 mm. long. Immaturus. n, frequently with a strong violaceous tinge showing through from beneath. Head and last segments above and beneath rust-yellow or light orange. Last pairs of legs yellowish. Ocelli 1+ 5,4 to 1+ 5,4, and 1+ 5, 4, 2 or 1+ 5,3,1; the anterior ones pale, whitish. Prosternal teeth 4 + 5 or 5 + 5; the outermost on each side largest. Spine slender, bristle tipped. Chitinous lines strongly evident. Articles of antennae twenty, shorter than in the adult; often in part somewhat moniliform. Posterior angles of ninth, eleventh, and thirteenth dorsal plates already strongly produced. Coxal pores mostly in a single series, but sometimes with one or two in a second more anterior row; small, margins pale; from four to seven on each coxa. . Spines of first legs, “2237347 ; of the second, §-q-232 ; of the thi fifth, $3 335 or sometimes, Coes ; of the sixth faa sev enth, ¢, = of the eighth and ninth, (-p°3:3'5 OF oo2s3; of the tenth, 00, = BY io 3-gro 3 of the eleventh, HE a aot of the twelfth and thirteenth j of the penult, 73:34 ; of the anal, 73344 « In the gonopeds of the female the three joints are distinct. The distal article short and apically more or less rounded, the claw being either not evident or else indicated merely as a minute point. The first article with three or four bristles, the second with two and the third with one on the ventral surface. Basal spines 1 + 1, very small, acutely conical, pale. Length8.5tollmm._ A specimen cir. 9.8 mm. long has antennae 3.7 mm. and anal legs 4 mm. long. Agenitalis.— Very pale yellowish, conspicuously violaceous. Head brighter yellow, in front of suture. Anal legs whitish at distal ends. Ocelli 1 + 3,2; well separated from each other; pale. Single ocellus but slightly larger than the adjacent one. Organ of Tomdésvary 9 d to 0,0,3,22 , DRON2 5S: 6, 1 2, 29 2, rc 2, 23 23 OF neal 1 ? Sule , 1, 3, ’ CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 405 elliptical, with outline as large as an ocellus; situated cephaloventrad of anterior eye of ventral row. Articles of antennae twenty, shorter than in the older stages, sub- moniliform. Prosternal teeth 5 + 5; small, the innermost smaller than the others; but little darkened at tips. Spine slender, bristle-like above base. Chitinous lines indistinct. Coxal pores pale and small, increasing in size distad. In a single series; ¢€.g., 2, 3, 3, 2. Spines of first legs, 9 fourth and fifth, 0 Boa OF : of the twelfth, nie 47 or ; of the thirteenth, o-o34; of the penult, 4 0, ae 3° . te i g oF of ihe anal, ? ?: z ae Gonopods of the female biarticulate; the distal article conical. Each article with one bristle. The distal article may show trace of subdivision line for formation of the third article. A specimen 7.5 mm. long has the anal legs cir. 3.5 mm. and the antenna cir. 4 mm. long. Known Locauities — Utah (canyons of the Wahsatch Mountains at lower and middle elevations throughout the range: the Weber! Farmington! City Creek! Mill Creek! Little and Big Cottonwood! American Fork! Provo! Spanish Fork! ete.). In and about the more thickly populated districts this species tends to be wholly replaced by Lithobius forficatus (Linné). The species sometimes occurs in association with E. awanti (Wood). This species is closely allied to B. hoples Brolemann,; but aside from its smaller size and usually different color, the ocelli are fewer, being from 15 to 25 in number in three or four series whereas in B. hoples they are from twenty-seven to forty in mostly five series; the spines of the female gonopods are typically characteristically different as shown in the figures; and the male lacks the marked difference in proportions presented by that of hoples, the first dorsal plate being always distinctly narrower than the eighth and tenth. 0, 0, 2, 2, 2 th ues : cee 3 cle is f ie 3 - of eo Fh to fhe cleve el : 1 ‘i ,0,2,2,1 7 uA 1 abal mal Boils a2 soy BoTHROPOLYS VICTORIANUS, Sp. Nov. Plate 3, fig. 5, 6. Description.— Brown, the caudal plates darker and of a ferrugi- nous cast. Head paler over the frontal region. Antennae brown, yellow distally. Legs similar in color to corresponding dorsal plates. 406 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Prosternum light brown of ferruginous tinge. Venter brown with the caudal plates more deeply pigmented, of ferruginous cast. Body slender, in type being about 9.6 times longer than the width of the eighth plate. The first dorsal plate in the female type not much differing in width from the tenth plate. Head wider than long in the ratio 63 : 61. Widest as usual immedi- ately behind eyes from where the sides converge a little to the rounded caudal corners, a little bent in at level of suture but lateral margin unbroken. Caudal margin slightly incurved at middle. An im- pressed dot each side of the median line a little caudad of the frontal suture continuous with a fainter furrow extending ectocephalad from it. A small V-shaped impression in front of caudal margin and cephalad of each of its ends a short oblique furrow which indistinctly unites with its fellow also to forma V-shaped outline; there is also a short oblique impression ectad of each arm of the V-shaped mark. A median longitudinal furrow extends forward from the frontal suture. Ocelli nineteen or twenty in four or five series; ¢.g., 1 + 5, 5, 5,3, 1 and 1 + 6, 5, 4, 3 Antennae short, reaching only to the fifth segment. Composed of twenty articles which are moderately long and subclavate, not much varying among themselves in proportions. Hairs of medium length. Prosternum between 1.5 and 1.6 times wider than long. Distance between chitinous lines twice the width at level of bottom of median sinus, 2.3-+ times the dental line. Teeth 8 + 8, uniform, apically rounded. Spines a little longer than the teeth, stout basally, distally attenuated, bristle-like. Posterior angles of eleventh and thirteenth dorsal plates distinctly produced, but the processes short; those of the ninth (Plate 3, fig. 5) but weakly and inconspicuously produced, the apex of the process scarcely extending caudad of median portion of the plate. Plates all strongly rugose over the median as well as the lateral portions. Most showing a median longitudinal furrow and a deeper, irregular, longi- tudinal furrow between the median one and the lateral edge on each side. The major plates with the usual submarginal transverse short sulci near beginning of the caudal third, each of these connecting with a furrow extending obliquely cephalomesad from its inner end. First and second ventral plates broadly depressed as usual. Others with the three longitudinal impressions much as in hoples. Spines of first legs, ¢ 0-0: ae 33 of the second to the sixth, (eee of the sev enth, 9 pees OF i oea2; of the eighth, ih 3 33; of the ninth and tenth, {9 = 33; of the eleventh, a: R = 35 OF G75 33-5; of the twelfth ’ 's CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE, 407 : 1,0) 35101 p TO ese tel: ze g and thirteenth, 613-32; Of the penult, 77-357, claw single; of the 0, 3,1 Tee Aa ; aa anal, ¢-3°3°9°] OF ¢-43-3'p, the claw single. Coxae of last two pairs of legs laterally armed. (Note.— The right penult leg of the type specimen has the ventral spines 1, 1, 3, 3, 1; but the lateral spines of the fourth joint are both on the cephalic side, one being thus clearly accessory and not to be reckoned in the normal formula). The anal legs of the female are moderate in length and slender. The prefemur and femur are weakly longitudinally sulcate dorsally. The claw of the female gonopods moderately wide; the lobes are acute, the middle one a little the largest, the apices of the lateral ones being but little proximad of that of the median one. Basal spines 2+ 2 or 3+ 2; not acuminate from base, narrowest near middle of length; apical portion very short (Plate 8, fig. 6). Length 25 mm.; width of eighth plate 2.6 mm.; length of antenna, 10 mm.; of anal ies exclusive of coxa, 8.4 mm. Pseudomaturus.— A little paler throughout than the maturus. Ocelli fourteen in three series; thus, 1 + 6.4, 3. Antennae as in maturus excepting that the ultimate article in speci- men studied is proportionately considerably longer. Prosternal teeth 6 + 6. Coxal pores fewer than in maturus; in two or three series. Spines of legs as in maturus. Claw of female gonopods as in maturus excepting that it is a little more slender. Spines 2+ 2 with the inner one of each pair only about two thirds as long as the outer one and correspondingly more slender. Of same form as in the maturus. Length 15.5 mm.; width of eighth plate 1.6 mm.; length of an- tennae 6.8 mm. Type Locauiry.— British Columbia, Vancouver Island (Victoria). Known Loca.itres.— Vancouver Island (Victoria! J. E. Benedict coll.) Alaska (Forrester Island! Harold and Ronald Heath). This species is allied to B. permundus and B. hoples, being appar- ently closest to the latter. So far as indicated by the type specimens (M. C. Z. 675) the species may be very readily distinguished through the character of the processes of the ninth, eleventh, and thirteenth _ dorsal plates, these being but weakly developed and those of the ninth in particular being almost obliterated. The spines of the female gonopods are of a different form. The species is also distinctly more slender and smaller than the characteristically robust B. hoples which is common in the neighboring parts of Washington and British Co- lumbia. The specimens from Forrester Island agree fully with those from Victoria. 408 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ZYGETHOPOLYS, gen. nov. Marginal thickening of head smoothly continuous forward to the eyes as in Bothropolys, not interrupted or broken at sides. Antennae short, reaching seventh segment; articles twenty. A diastema present in the dental line on each side of prosternum and separating the teeth into two groups. Spine situated in the diastema, well chitinized, stout at base. Claw of anal legs single; that of the penult legs with a well-devel- oped accessory claw. Ventral spines of anal legs 1, 1, 3, 3, 1. Coxae of only the anal legs bearing a spine ventrally. Last two pairs of coxae armed laterally. Gonopods of female with claw bipartite, the lobes equal. Basal spines 2 + 2. Male unknown. Type.— Z. nothus, sp. nov. As the genus is known from only the female of a single species, it is not possible to speak with entire certainty as to its position other than to remark that it combines features of Bothropolys and Ethopolys with apparently more in common with the former, under which it might seem justifiable to range it as a subgenus. It is better and clearer, however, at present to keep the group apart. From Bothro- polys it may be separated at once through the presence of the prosternal diastema. It is also different in having the spining of the anal legs ventrally 1, 1, 3,3, 1 instead of the 1,1, 3,2,1 (rarely 1, 1,3, 3,2 present in all species of Bothropolys, in lacking a ventral spine on the penult coxae, and in having the claw of the penalt legs with a well- developed accessory spine or claw at base. The claw of the female gonopods is also characteristic in being bipartite with the lobes equal, it being tripartite in all known species of Bothropolys. The first dorsal plate also is narrower than the third, an interesting condition not noted in any other species of the Ethopolidae. Outside of the Ethopolidae, the only genus of Lithobioidea known to the author to have a diastema in the prosternal dental line is Ere- mobius, a new genus typified by Lithobius provocator Pocock of the Bermuda Islands. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 409 ZYGETHOPOLYS NOTHUS, sp. nov. Plate.s, fig. 7,8: DescrieTion.— Dorsum brown, dusky toward edges, the first one and the last several plates a little darker, of slightly reddish or dilute chestnut cast. Head dilute chestnut, dusky especially in a median longitudinal band continuous anteriorly with a narrow curved trans- verse one a little back of and parallel with the frontal suture between which and the dark band there is a clear area. Antennae dark brown excepting the ultimate article which is abruptly brighter yellow in color. Legs light brown excepting the last two pairs, which are dark, the anal pair especially so but pale along mesal surface. Prosternum and prehensors testaceous. Venter light brown, excepting fourteenth and fifteenth sternites which are much darker, chestnut or dark reddish brown. Body only about 6.5 times longer than the width of the tenth plate. First dorsal plate narrower than the head and the third plate, the latter distinctly narrower than the eighth and tenth plates. Widths of head and of first, third, eighth, tenth, and twelfth plates to each other as 49 : 46 :.49 : 56 : 57 : 55. Head wider than long in the ratio 49 : 46. A deeply impressed curved transverse sulcus between eyes and a little distance caudad of the suture, this sulcus consisting of three curved sulci united at ends, each with convexity cephalad. A short and deep semicircular or U-shaped transverse impression a little in front of the caudal marginal thickening. A median longitudinal furrow extending from frontal suture cephalad and bifureating at anterior end. Surface not punc- tate. Eye-patch oblong, composed of twenty-three ocelli arranged in four series: thus, 1 + 6, 6, 6,4. Single ocellus circular, contiguous with seriate patch. Seriate ocelli more nearly uniform in size than usual, distinct, black. Antennae short, reaching to the seventh segment. Articles twenty, of moderate size or short. _ Ultimate article a little longer than the two preceding taken together. Prosternum 1.9 times wider than long. Distance between chitinous lines 1.9 times the dental line. Spine stout at base, narrowed to an acute point, inserted contiguously to margin. Teeth 2 — 6+ 6 — 2, the most ectal and the most mesal on each side smallest, the others nearly uniform in size. Mesal incision very narrow, acute. 410 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. i» : Fic. 2.— Distribution of Zygethopolys nothus Chamberlin. First dorsal plate 1.7 times wider than long. A characteristic semi- circular impression having an end on the anterior margin at about two thirds the distance from the median line to the lateral margin. Other plates moderately and more or less uniformly roughened over the CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 411 entire surface, the major ones with the short submarginal transverse sulci found in other species of the tribe but without distinct longitudi- nal or oblique furrows; a semicircular impression like that of the first plate faintly indicated, this more distinct on the eighth and tenth plates, the impression on the tenth terminating on a second similar but more convex impression. Posterior angles of the ninth, eleventh, and thirteenth dorsal plates produced strongly, others straight. Ventral plates with three longitudinal furrows, the median one especially becoming more distinct caudad and crossing entire length of plates, the lateral furrows on the fourteenth and fifteenth plates indistinct. ‘ Coxal pores eight to fifteen in two or three series, there being two series on the anal coxae and three on the others. . 0,0,3,2.1 0,0, 3, 2 . Spines of first legs, (tas; of the second, j-o'33°5; of the third to . 0, 3, 2 2 W002 2. ae ninth, 9:9/9°3,3; of the tenth and eleventh, 9:93:32; of the twelfth to penult inclusive, ¢{'3:3:3, the claw of the penult legs with a well- developed accessory claw; of the anal }7:3:3:4, the claw single. Last two pairs of coxae laterally armed. Anal legs in female moderately long, slender. Gonopods of female with the claw small; not at all or but slightly curved; rather deeply bipartite with the lobes acute, equal (Plate 3, fig. 7). Basal spines 2 + 2, with the inner one of each pair distinctly smaller than the outer; spines in surface view showing a short, not at all acute, terminal portion and a proportionately long subcylindric basal portion that is somewhat broader distally than proximally (Plate 3, fig. 8). . Length 15 mm.; width of tenth plate, 2.3 mm.; length of antennae 7mm.; of anal legs, 7.8mm. Type, M.C. Z. 687. Locauity.— Alaska (Forrester Island!). Collected by Harold and Ronald Heath, May 20-26, 1913. {THOPOLYS Chamberlin. Can. ent., 1912, 44, p. 173. Lateral marginal thickening of head abruptly discontinuous or broken a little back of middle or near caudal end of the longitudinal curved impression back of each eye (Plate 4, fig. 1). Antennae usually, but not always, rather long and reaching to or beyond the eighth segment. Composed of twenty or rarely of as many as twenty-eight articles. 412 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. In the prosternum a well-marked diastema separating an outer group of teeth on each side from a larger inner group with a basally stout but distally slender spine situated in the diastema (Plate 4, fighe2>) Claw of anal legs armed with a minute accessory claw or this obso- lete or absent; that of the penult legs with two accessory claws. Coxae of last two pairs of legs each bearing a stout ventral spine as well as a dorsal and a lateral one. Gonopods of male distinctly biarticulate. Claw of female gonopods tripartite or entire; when tripartite with the lateral divisions small and situated considerably proximad of the median one. Basal spines 3 + 3 or 4 + 4 or occasionally 4 +- 5. Typr.— Ethopolys xanti: (Wood). In this genus a semicircular impression caudad of each eye is distinct as in the preceding genus. As in Bothropolys also there is not great variation in the spining of the legs within the genus, this largely being the same also as for that genus. The dorsal spines of the anal legs are always 1,0, 3, 1,0; the ventral 1, 1, 3, 2,1, the claw with a very small accessory spine or single. Dorsal spines of the penult legs 1,0, 3,1, 1, ventral 1, 1, 3, 3, 2, two accessory claws present. Dorsal spines of the thirteenth legs 1, 0, 3, 2, 2, or, more rarely, 1, 0, 3, 1, 1; the ventral, 0, 1,3, 3,2. Dorsal spines of the twelfth legs, 0, 0, 3, 2, 2 or 120:'3,-220remore’rarely..12/0;.5 leat ventral alieys Onl. 383,022 Dorsal spines of the eleventh legs, 0, 0, 3,2 2) 2 or l.\Os3) 2,257 ventura: 0, 0, 3, 3, 2 or rarely 0,0, 2,3, 2. Dorsal spines of the second legs, 0, 0, 3, 2, 2; ventral 0, 0, 2,3, 2. Dorsal spines of first legs 0, 0, 3, 2, 1; ventral 0, 0, 2, 3, 2. In the species known to belong to this genus either none of the dorsal plates have their posterior angles produced or else those of the ninth, eleventh, and thirteenth are extended. The name Ethopolys sens str. may cover the latter group which contains the type, E. xanti (Wood), while for the former Archethopolys, subgen. nov., with E. integer Chamberlin as the type, may be used. The prosternal spine in the species of this genus seems practically always to retain its slender terminal portion throughout life (Plate 4 fig. 2,3, 4; Plate 5, fig. 7; Plate 6, fig. 6). In variations in the specimens of species of this genus resulting in entirety or a marked tendency toward entirety of the claw of the female gonopods, the result is due apparently to reduction in or dis- appearance of the lateral teeth with codrdinate strengthening of the median one, to a removal of the lateral teeth rather than to an evening CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 413 Fic. 3.— Distribution of Ethopolys Chamberlin. up such as occurs in Bothropolys. The consequence of this is a long, distally acute claw (Plate 5, fig. 8), contrasting with the blunt claw in Bothropolys (Plate 2, fig. 7; Plate 3, fig. 1). 414 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Key to Species of Ethopolys. a. Posterior angles of none of the dorsal plates produced. Archethopolys subgen. nov. b. Only one tooth ectad of the diastema on prosternum; dorsal spines of twelfth legs, 1, 0, 3, 1, 1; antennae short; length 14-18 mm. E. pusio (Stuxberg). bb. Two or more teeth (at least on one side) ectad of diastema on pro- sternum; dorsal spines of twelfth legs, 1, 0,3, 2,2, or 0, 0, 3, 2, 2; antennae long; length, 19-35 mm. c. Prosternum normally with only two teeth ectad of diastema, rarely with three; claw of female gonopods always tripartite. d. Antennae reaching to or beyond the ninth segment; color yellow to light amber-brown, the head not reddish. E. bipunctatus (Wood). dd. Antennae not extending beyond the eighth segment; color darker brown, commonly in part dusky, head of reddish cast, typically chestnultarsecc: ene ee eee E. sterravagus (Chamberlin). cc. Prosternum normally with three or four teeth ectad of diastema, only occasionally with but two; claw of @ gonopods entire or bipartite. d. Head coarsely and conspicuously punctate; claw of female gono- pods entire, with no distinctly developed lateral teeth. (Wash- IN FON ACTCs) Fata hee Ce ea eee oe E. integer Chamberlin. dd. Head not punctate, or punctae scarce and fine; claw of 2 gono- pods bipartite (Alaska)....... E. integer alaskanus Chamberlin. aa. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced. : Ethopolys sens.str. Claw of @ gonopods tripartite; prosternum with three or two teeth ectad oL.diastema.. voc tek ah Oe ee ee E. xanti (Wood). ARCHETHOPOLYS, subgen. nov. ETHOPOLYS BIPUNCTATUS (Wood). Plate 5, fig. 5. Bothropolys bipunctatus Wood, Journ. Acad. nat. sci. Phil., 1863, new ser., 5, p. 16. Trans. Amer. philos. soc., 1865, 18, p. 153. Lithobius (Archilithobius) bipunctatus Stuxberg, Ofvers. Kongl. vet. akad. Férh., 1875, 32, no. 3, p. 14, 30. Lithobius bipunctatus Chamberlin, Proc. U. 8. N. M., 1901, 24, p. 22. Ethopolys bipunctatus Chamberlin, Can. ent., 1912, 44, p. 174. DescripTion.— This is the palest species of the genus, varying from light amber-brown, with the first one and the last several seg- CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 415 ments lighter, to almost clear yellow and uniform or with the last few segments a little darker. Head mostly very dilute orange, faintly dusky caudad of the frontal suture. Antennae very light brownish yellow or yellow proximad and darker distad of proximal third or halt of length excepting sometimes at very tip. Prosternum and _pre- hensorial feet and last ventral plates dilute orange or rusty yellow. Last pairs of legs yellow of weak orange or rusty tinge or the tarsi darker and more distinctly orange. Venter over median portion often brownish or dusky, usually clear, vellow cephalad. Gradually and uniformly narrowed caudad from the tenth plate and also narrowed cephalad of the tenth plate considerably more than in the related species. | Head with sides convex, a little converging from the eyes to the marginal breaks and then more strongly than usual rounding and converging mesocaudad to the caudal corners. The usual short median longitudinal impression contiguous with and immediately in front of the median angle of the caudal marginal thickening, with from its anterior end two weak furrows diverging from it cephalad. Eyes composed of from thirteen to twenty-two ocelli, which are distinct and regular. Series three or four but mostly the latter. Single ocellus much larger than the others, paler, broadly elliptic, not contiguous with other ocelli. The number of ocelli is frequently seventeen in the following arrangement: 1 + 5,5, 3, 2; 1+ 5, 5,5, 4; and | + 5, 4, 3. Antennae long, usually reaching the ninth segment. Composed of twenty long, subcylindric articles, of which those distad of the proxi- mal three to five are usually more strictly cylindric. Hairs rather short. Prosternum 1.7 times wider than long. Chitinous lines absent or obscure. Spine attached a little caudad of anterior edge of diastema; attenuated and becoming distally bristle-like, pale. The most fre- quent number and arrangement of the teeth is 2 — 6+ 6 — 2, ve., with two teeth ectad of and six teeth mesad of the diastema on each side. Other arrangements are: 1—6+6—2; 2—5+6-— 3; 1 —5+ 5 — 3; and in once case even 4 — 7 + 6 — 4. Posterior angles of none of the dorsal plates produced, those of the ninth, eleventh, and thirteenth being obliquely excised. All major plates roughened, the more posterior ones most strongly so, the first plate, however, being smooth or nearly so. Major plates with one or two longitudinal furrows on each side subparallel with the margin and which may be more or less broken and a third oblique furrow running 416 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. from near middle of anterior margin caudoectad and meeting the lateral furrows at an angle; these furrows absent from or very obscure on first plate. First, third, fifth, eighth, tenth, twelfth, and fourteenth plates with the usual short transverse submarginal sulcus on each side near beginning of caudal third of length and all of these plates except- ing the first also with a distinct subtranverse impression near each anterior corner; the seventh plate also with two corresponding pairs of impressions but, as usual, the posterior pair of these is near the caudal corners. The minor plates, especially the more caudal ones, with a curved, subtransverse, sulcus at each anterior corner, the impression running somewhat caudomesad with the convexity caudad and mostly covered by edges of major plates; also with a short curved mark at each caudal corner adjacent to marginal thickening. First ventral plate with two fovea near anterior margin and cepha- lad of the broad-depression on the more caudal portion of plate, a short transverse deeper furrow occurring in the latter depression. Of the usual three longitudinal impressions the lateral ones show in varying degrees of distinctness; the median may be only obscurely indicated on the anterior plates, while it becomes more distinct on the caudal. On most a transverse depression in front of caudal margin which is commonly distinctly furrow-like on at least the last two plates where it terminates. Coxal pores twelve to twenty-five in number on each coxa arranged in three or four series. As usual, of varying sizes, those of the most anterior row much smallest and often minute; all circular or subel- liptie. Spines of first legs, i332; of the second to the tenth, (33:3; of the elev enth, 9 0 ie ae 2, 2 or aOR 33; of the twelfth, a: + 5 353 of the thir- teenth, ¢ OL eh ae or HES: ol the penult, 7 CHE ee 3-9, two accessory claws present; of the anal, 74°3:9'1, claw single. Anal legs long and slender; those of male not manifestly modified. Claw of female gonopods long and stout, considerably curved, tri- dentate distally, the lateral lobes less proximad than usual, the median much stoutest. Basal spines 3+ 3, the outermost a little stouter than the median and the innermost considerably less robust than the median, all often bent a little ectad at tips, black above bases. The two articles of male gonopods together subconical, the distal article apically rounded. Proximal article mostly with near four bristles and the distal with two. Length from 19 mm. to 29 or 30 mm. A female 24.5 mm. long has the tenth plate 3.1 mm. wide; the third 2.5 mm. wide; the first at CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 417 widest level 2.8 mm. wide; the antennae 15 mm. long; and the anal legs 13 mm. long. Pseudomaturus.— Coloration somewhat paler than the average in adult, the head being more yellow without or with but little of the orange or rusty tinge. Antennae yellow. Anal legs yellow. Other legs often pale dusky brown proximally with tibiae and tarsi yellow. Caudal ventral plates often dusky. Ocelli about twelve in three series; ¢. g., 1 + 4,4, 3; not contiguous. The most caudal ocellus in each of the two upper series considerably larger than others. Antennae very long, reaching to or a little beyond the ninth segment. Articles twenty, long and mostly strictly cylindric as in the adult. Prosternal teeth 2 — 5+ 5 — 2, mostly. Spine slender and pale. Teeth regular and uniform, darkened distally. Chitinous_ lines faint. Coxal pores in two series, those of the more cephalic one much smaller and generally fewer; six to twelve on each coxa, the distal one of caudal row often very much larger than others. 0, 0, 3, 2, 1 Spines of frst to sixth legs, 90-333; of sev enth to ninth, (0:3 2 2 2: DISH 9 g Dy 25 2 0, 0, 3, 2, 2 p i, Wh ze 2 of the tenth, ¢°0'3°3°3 OF p-4-3:3-3; of the eleventh, o. 933.53 of the twelfth, 0; 0, 3, 2, 2 | ee esi a 5; ot the thirteenth, >; of the penult, ; 3-9; Ol the anal, 0133.2 0, 1, 3, 3,2 13.3.2 Elst Distal article of male gonopods broadly subconic, in specimen described glabrous; first article with two or three bristles. Length of specimen described 15.5 mm.; of antennae 9.5 mm.; of anal leg 9 mm. Immaturus.— Coloration very similar to that of the pseudomaturus but still paler; the antennae palest proximally and at tips, somewhat orange elsewhere. Legs more uniform in color, the tarsi a little brighter; anal legs yellow. Ocelli six or eight in two or occasionally in three series; e¢. 9., 1+ 3,2; 14+ 3,3,1. The first ocellus of the upper row rather larger than the single ocellus, much larger than other seriate ones. Organ of Tomésvary a little smaller in outline than the smallest ocellus. Antennae proportionately shorter, reaching only to the eighth seg- ment. Articles twenty or twenty-one all relatively shorter than in the adult and some, occurring mostly in two’s at intervals, much shorter, or as many as six of these may be grouped immediately proximad of the - ultimate one. Hairs relatively longer than in the adult. Prosternum with chitinous lines weak but rather more distinct than in the older stages. Teeth 1 —4+4-—1 or 2—4-+ 4 — 2 with 418 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. the outermost one on each side in the latter case very small. Spine slender and bristle-like above base. Coxal pores in one series, small with very pale margins; 2, 3, 3, 2 OLieowone: OS 002,051 Spines of first legs, goii4 Ko 0 ( 0, 0, 3, 2, 1 i, 2,4 OF 2 OOmh eA 0221; of the second, 6: 0, 1 32,2 3; of the sixth to eighth, 9 0. 2313 0,0,3,2.2 239, the spine of ; of the penult, § a 2 e of the third to fifth, 7: , 3, © of the ninth to elev enth, ? 0: a aaa Omune twelfth the trochanter minute; of the thirteenth, ot Pisa2 OF Tr 332; of the anal, ese Second article of male gonopods faictner rather short, rounded apically, pale and glabrous. Proximal article narrowed from base to second article, where it is of same width as the contiguous portion of the latter, bearing one bristle. A male 9 mm. long has antennae 5.75 mm. and anal legs 4.25 mm. long. Agenitalis. 5 @ il Onoene Bone Pale brown mesally, whitish towards ends with the first and last segments yellow. Head yellow, whitish caudally. Antennae whitish vellow. Legs pale, the caudal pairs of a yellowish tinge. Ocelli five in two series; thus, 1 + 2,2. All pale, the single ocellus smaller than the proximate one of the upper row, all separated from each other. Organ of Tomésvary very small in outline. Antennae reaching to the seventh segment. Articles seventeen short, the shortest ones occurring at intervals. Hairs relatively longer than in the adult. Prosternal teeth 1 — 4-+ 4 — 1, spine bristle-like above base. Teeth brown above bases, distally rounded, uniform; or the outermost and innermost on each ade a little smaller than the others. Coxal pores 1, 1, 1, i or 2 my Nile Spines of fat legs, fot : to ftaii; of second to seventh, ane the ventral spine of third article minute; of eighth and ninth, 9° of the tenth, ('¢-7-3.4; of the eleventh, 00, 721, of the twelfth, pe of the thirteenth, ee of the penult, = of or terri; of ‘the anal, {-;q-9- The spine on the ventral surface of the third article becomes smaller and smaller cephalad, on the first pair being minute and difficult to detect or quite absent. Gonopods very short and mostly concealed; undivided. Anal glands still evident. Length 7.5 mm.; antennae 4.6 mm. long; anal leg 2.8 mm. long. Type Locauiry.— “ United States West of the Rocky Mountains” (George Suckley, collector). CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 419 Known Locauiries.— Utah (Emigration Canyon! Wildwood, Provo Canyon! Clinton’s Cave, near Lake Point! Uintah Moun- tains! Daniell’s Creek!). Nevada! ETHOPOLYS SIERRAVAGUS (Chamberlin). Plate 5, fig. 1-4. Lithobius sierravagus Chamberlin, Proc. Acad. nat. sci. Phil., 1903, p. 154. Bothropolys monticola Chamberlin (ad part. max. non Stuxberg), Pomona coll. journ. ent., 1910, 2, p. 369. Ethopolys sierravagus Chamberlin, Can. ent., 1912, 44, p. 174. Description.— Dorsum light to dark brown, commonly dusky and sometimes very dark, the first one and commonly the last two plates commonly of dilute chestnut cast like the head; nearly always on the major plates a pale median longitudinal stripe with one on each side of it extending obliquely ectocaudad; sometimes some or all of the principal plates bordered caudally and laterally with a distinct stripe of blackish. Head from brown of slight reddish cast to dilute or clear chestnut, dusky immediately caudad of the frontal suture. Antennae proximally like the head, distally light brown or yellowish. Legs brown or testaceous, the posterior pairs darker, often of dilute chest- nut tinge. Venter brown, the fourteenth and fifteenth plates of chestnut cast. Prosternum and prehensors dilute chestnut. Body from 7.75 to 8.5 times longer than wide. The first plate in both male and female considerably narrower than the tenth. Widths of head and of first, third, eighth, and twelfth plates to each other in © as 80:81 :75:85:87:82; ina o as 69:70:65:74:75:73. The first plate is slightly wider than the head. Head wider than long (67 :65 ina o*; 77:75 ina 9). Of the usual general form, being widest near or in front of lateral marginal breaks. A U-shaped impression, which is not very deep, on caudal ‘portion, the mark being connected at its middle with a short longi- tudinal furrow extending to the caudal marginal thickening; a rather deep short transverse submarginal sulcus a little caudad of level of marginal break on each side. Surface a little uneven, subsparsely and rather finely punctate. Eyes consisting mostly of from thirteen to twenty ocelli arranged in three or four, but much most commonly in four series; ¢.g., 1 + 4, 5, a2 le Ona s LS OnOy4nas © I -i6rb; 4294 de 5,554, 2; 1+ 5,5,4; 1+.6,5,3; 1+.5,4,3. Seriate ocelli distinct, decreas- 420 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ing gradually and but little cephalad and ventrad, those of the most ventral row often very small. Single ocellus separated by a space from the others; pale; subtriangular, with the sides convex and the angles rounded. Antennae of moderate length but varying considerably, mostly reaching the seventh segment, but sometimes the eighth and rarely even the ninth on one side. Articles from twenty to as many as twenty-eight; mostly long and subeylindric, distal ones slender; the ultimate much shorter than the two preceding taken together, or when the articles are twenty often but little longer than the penult or even shorter than it. In antennae with the larger numbers of articles a variable number of those preceding the ultimate have become divided, giving correspondingly shorter articles; thus in an antenna with twenty-four articles the first eleven may be of the usual long form while the next six have divided, giving twelve decidedly shorter articles which with the ultimate increases the number from twenty to twenty-four, ete. (Plate 5, fig. 3, 4). Often the antennae with the larger number of articles are relatively shorter than those with fewer. Hairs very short, dense. : Prosternum between 1.66-++ and 1.8 times wider than long. Dis- tance between chitinous spots 2 to 2.25 times the dental line, 1.8 or 1.9 times width at level of bottom of mesal sinus. Spine situated very close to or nearly upon anterior margin; stout at base and distally acuminate as usual. Nearly always two teeth ectad of each diastema, sometimes but one on one or both sides or with these on one side, much more rarely upon both. Much the most frequent arrangement noted is 2 — 7+ 6 — 2, with the larger number on the left side. Other arrangements observed are as follows, the left side being represented first: 2—6+7—2; 2—6+6—3; 2—6-+6—T1; 2—6+6-2; 1—6+6-—1; 2—64+7-1; 2—114+ 6-2; Bie) Op LOt 2h = 6 OL or First dorsal plate of the usual form; surface like that of head; twice as wide as long. Other plates rugose and tuberculate laterally, the: median portion remaining smoother, the roughening becoming more and more pronounced in going caudad; the roughening clearly less than in FE. xanti. The short submarginal transverse sulci near be- ginning of the caudal third evident on the first, third, fifth, eighth, tenth, twelfth, and fourteenth plates as usual but apparently less pronounced than in FE. integer; from the inner end of each of these extends a furrow mesocephalad and another directly cephalad to the anterior margin, the latter furrow on the more caudal plates extending through to or nearly to the caudal margin. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 4?1 _ Posterior angles of none of the dorsal plates produced; those of the ninth, eleventh, and thirteenth straight to considerably obliquely excised, occasionally those of the thirteenth very slightly produced. Ventral plates with the usual three longitudinal furrows of which the median one is mostly pit-like and confined to the caudal part of plate but may be more indistinctly extended cephalad and caudad. In the female the fourteenth and fifteenth plates are transversely depressed across the middle with the furrows obscure; but in the male the fur- rows are strongly impressed, the lateral ones on the fourteenth con- verging caudad and commonly ending abruptly at beginning of caudal fourth of length, while on the fifteenth the lateral furrows are sub- parallel or a little excurved and terminate abruptly as on the four- teenth, while the median furrow on both plates is typically reduced to one or a row of a few impressed dots or pits but may appear as a con- tinuous sulcus, especially on the fourteenth; the lateral furrows also may appear as rows of dot-like pits. A wide lateral and caudal border on both of these plates is smooth and densely clothed with fine short hairs. Coxal pores mostly twenty-five to forty-five in number on each coxa, the smallest number being on the twelfth pair. Series three to five, the most anterior of which is often irregular or confused. Spines of first legs, i223 of the second, jy ,2:5,2 oF o:j2's.23 of the third to tenth, 002 3.23 of the eleventh, 0, O33 2; of the twelfth, ¢ 1332 OF 0 13,3,9; Of the thirteenth, 09-73-35; of the penult, 7 7°3°3:2, with two accessory claws; of the anal, 77°34) toy 3'3 1, the claw single or with a minute accessory claw. Last two pairs of coxae laterally armed or, occasionally, only the last pair. Claw of gonopods of female of moderate size, comparatively broad distally, distinctly tripartite, the median lobe largest, the lateral proximad of the median (Plate 5, fig. 1). Occasionally one lateral tooth may be replaced by two or more minute denticles or may be almost obliterated. Basal spines 3 + 3 to 4+ 4, typically gradually and uniformly narrowing from base to the subacute distal point. When a fourth spine is present it is commonly small and inserted on mesal side of article where it is easily overlooked (See Plate 5, fig. 2). Length from 20 to 28 mm. A female 28 mm. long has the anal leg 12 mm. long and the tenth dorsal plate 3.6 mm. wide. Pseudomaturus.— Light brown, the major dorsal plates with the three longitudinal pale lines well marked, last segment darker. Head pale orange, dusky, clearer in color immediately along the frontal suture. Prosternum and prehensorial feet pale orange. Antennae yellowish or light orange-brown, paler distad. Legs pale, the tarsi © 492 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. brighter. Last pairs light orange, somewhat duller in color proximally. Venter pale brown, the last segment a little darker. Ocelli 1 + 5,5, 3,2 in specimen described. Single ocellus very much larger than any other, suborbicular or broadly suboval with the narrower end ventrocephalad. Organ of Tomésvary in outline smaller than smallest ocellus, in line with ventral row and close to anterior ocellus of same. Antennae, in specimens examined, composed of from twenty to twenty-two articles, when twenty-two the extra ones apparently resulting from subdivision of the penult and antepenult articles; the ultimate article relatively shorter than in earlier stages. Prosternum with chitinous lines distinct. Teeth 2 — 6+ 6 — 2 Spine as in maturus. Coxal pores eleven to nineteen in number, in three series, sizes vary- ing as In maturus. Mes : Dae Hal 0, 0. 3. 3 1 0;:0;.3,.2, 2 Spines of first to third legs, g 9.9-3-2; Of the fourth, 5 0,2 Bee OF 0-0,23,3 3 2 03:0, 3, 2, .2 0:3; '2, 2 of the fifth ito ninth, 6 oes; of he tenth, 9-9-3°3°2 to 03,3, 2% of the 0).3, 2,.2 150,35 25.2: WON 2 eleventh, j-:3°3°2; of the twelfth, 0133-23 of the thi ec Os a5 OF ¥10).35:2).2 1, 0;.35 15.1 1,0 0, 1, 3, 3,2) of the penult, iL abe} 3,2) ot the anal, 7 1, ? * 2 41° Fifteenth and succeeding segment subdensely pilose on ventral surface, hair of the fourteenth fewer. Claw of the female gonopods shorter and thinner and less strongly chitinized than in the adult, the teeth as usual. Basal spines 3 + 3, but the innermost on each side only about one half the height of the median one and much more slender. Length 19 mm.; antennae 10 mm. Praematurus.— Pale brown, with violaceous tint showing through from beneath. Head in front of suture, light orange, elsewhere dark- ened or smoky. Antennae pale brown to yellowish, tending towards orange, paler distad. Ocelli 1 + 4,3,2; 1+ 4,4,2. Rather pale. The single ocellus larger than the first one of the upper row, the anterior ones of ventral row very small. : Antennae reaching well along on eighth segment. Articles twenty, all of moderate length or long, the last one much longer than the penult. Prosternal teeth 2 — 5 + 5 — 2. Coxal pores very small, the distal one of caudal series conspicuously largest; arranged in two series, four to nine in number on each coxa, the smallest number occurring on the twelfth and the fifteenth coxae. 0, 0, 3, 2,1 5 ; : Spines of first and second legs, o-9-9:3:1; of the third to the sixth, CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 423 3 2 . OOS ese g (0335; of the seventh to the ninth, ¢'o/932; of tenth and eleventh, oe q q 1), Shoe See miny iko TRONS lal eve 5 5; of the twelfth, ¢73:3:3; of the thirteenth, 9-73'33; of the d im 3 BSS 5 . eg 2 ’ ,riage- (Anal missing). Coxae of last two pairs armed as in the adult. Fifteenth ventral plate subdensely pilose, especially over the caudal portion. Claw of female gonopods pale, short, and thin, not curved, the lateral teeth appearing only as points at base. Spines 2 + 2, the innermost on each side short and very slender, almost bristle-like. Length of male 13 mm.; antennae 7.6 mm. Length of female, 15 to 16 mm.; antennae 7.5 to 7.8 mm. Tyre Locauiry.— California (Shasta Springs). Known Locauitrres.— California (Truckee! Shasta Springs! Hay- ward). Oregon (Portland! Oregon City!). Ranging from Portland southward along the Sierra Nevada Moun- tains. It seems not to occur in the coastal mountains or the region south of San Francisco Bay. North of Oregon it is replaced by E. integer Chamberlin, a closely related species. THOPOLYS INTEGER Chamberlin. Plate 5, fig. 6-8. Plate 6, fig. 1. Ethopolys integer Chamberlin, Rept. Can. Arctic exped., 1913-18, 1919, 3, Tre Jal, jo), 720) Jel _ Description.— Dorsum mostly dark brown with the caudal plates darker; major plates often darkened over the lateral and caudal borders; major plates of posterior half of body typically with a distinct longitudinal median black stripe, this becoming indistinct on the more anterior ones; main plates commonly with a light stripe extending obliquely caudolaterad on each side from near middle of anterior margin. Head, as also in part the first dorsal plate in some, reddish brown to chestnut; a blackish median longitudinal stripe extending from caudal margin to frontal suture caudad of which it expands laterad. Antennae brown to light chestnut, paler, light brown, distad. Venter light brown, the fourteenth and _ fifteenth sternites commonly of a chestnut tinge. Prosternum and prehensors brown of a light chestnut tinge. Legs testaceous, the posterior pairs darker, brown to light chestnut-brown. Body of same form in male and female; or nearly so. About eight times longer than width of the tenth plate. Head and first dorsal 424 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. plate of same width, narrower than the tenth plate. Widths of head and of first, third, eighth, tenth, and twelfth dorsal plates to each other IM Gt ASS. Osage (42 (eS: On. 19) el SOlenio eg fo Oe orn Omar and in @ as, e.g., 84 : 84 : 78 :87:: 87 : 85 or 84 : 84 : 76 : 86 : 87 : 85. Head distinctly wider than long (86 : 79); widest a little caudad of lateral breaks. A V-shaped impression on caudal half of cephalic plate. Entire surface subdensely punctate, the puncta distinct. Ocelli from twelve to nineteen in three or four series, but by far most commonly in 4; ¢.g., 1 + 5, 6, 4,3; 1+ 6, 5, 4,3; 1+ 5, 5, 4, 2; 1s 0ON3; 2a Ld, 5, Ano as Oso dl olathe Acre Dg el I etye 4,2; 1+ 5,5,4; 1-+6,5,3. Simgle ocellus much largest, sub- orbicular to somewhat angular, clearly separated by a space from the others. Seriate ocelli distinct, regular, decreasing moderately ventrad and cephalad. Antennae reaching to from the fifth to the eighth segments, but mostly to the sixth or the seventh. Articles twenty to twenty-four mostly long and cylindric or nearly so, mostly not clavate or but slightly so. Ultimate article long and slender, a little shorter than the two preceding together. Prosternum c7r. 1.7 times wider than long. Distance between chitinous spots in specimen measured 1.97 times the dental line and 1.75 times the width at level of bottom of median sinus. Chitinous lines well developed excepting toward caudal ends. Finely, densely punctate. Spine inserted on ventral surface a little caudad of the anterior edge; moderately short, uniformly attentuated to an acute point; much stouter than the bristles (Plate 5, fig. 7). Teeth conical, subacute or apically narrowly rounded; those adjacent to diastema ‘ on each side largest, others very gradually decreasing ectad and mesad from these. Most commonly three or four teeth ectad of diastema but also in some only two. Examples of arrangement are as follows, the left side being indicated first in the formulae: 3 — 7 + 6 — 3, the commonest of all arrangements and numbers; 3 — 6 + 6—3; 3—5+6—3; 4—6+6—-—4; 2—6+6—3; 2—6+ 6—2;2-—6+ 5-2. First dorsal plate narrowed from anterior end caudad as usual, wider than long nearly in ratio 21:11; surface nearly like that of head, punctate but not rugose. Major plates with the short trans- verse Impressions at caudal third of length deeply marked. A furrow extending from inner end of each of these submarginal sulci obliquely mesocephalad to near middle of anterior margin and also an irregular deep furrow extending directly cephalad to anterior margin. All CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 425 plates distinctly and rather coarsely punctate, and, especially the more caudal ones, subfinely rugose and irregularly tuberculate toward lateral borders, the median portion remaining nearly smooth excepting on the fifteenth plate. Ventral plates, excepting last one or two, densely finely punctate over entire surface. .A lateral longitudinal impression in front of the short suture at each caudolateral corner, these being more narrowly and sharply impressed on caudal plates. On the caudal portion be- tween the lateral impressions is a short median pit or depression which, however, is sometimes elongate. The fifteenth sternite is wholly smooth over a wide lateral and caudal border and is here densely clothed with very fine, short hairs. In the male the fourteenth plate has a V-shaped impression formed by two distinct sulci converging from the anterior margin to a point at or a little in front of the caudal margin or these converging sulci sometimes ending farther cephalad and not meeting; also a median longitudinal sulcus usually of same distinctness as the lateral ones. In the male the fifteenth sternite also is marked with three deeply impressed longitudinal sulei which terminate at the beginning of the smooth caudal border; of these the lateral converge a little but less than those of the fourteenth plate. ew 0, 3,2 0. 0, 3, 2,2 : d Spines of first legs, § 04.0, 35 to pores (at least on one side); of the 0; 0,.3,°2,.2 BUS R-Fr > y second to tenth, pe oe On tle elev enth, 10-330; of the twelfth, . ; oe or o: L ay 3 >; of the thirteenth, a: : - 233 of the penult, i s a e55 with two accessory claws; of the anal, 7133], claw single or with a very minute accessory claw. Last two pairs of coxae, or occasionally only the last pair, laterally armed. Anal legs of male moderately long, slender, the fourth joint with a longitudinal dorsal sulcus over the median portion and ectad of this a longer sulcus extending the full length of the joint; corresponding to the latter there is an ectodorsal longitudinal sulcus on the fifth joint. Third joint with a longitudinal dorsal sulcus. Claw of female gonopods long and well curved; entire, with no trace of lateral teeth (Plate 5, fig. 8). Basal spines 3 + 3; acuminate from near middle of length, apically narrowly rounded. Length 20 to 30 mm. A female 29 mm. long has anal leg 12 mm. long, antennae 15 mm. long and the tenth dorsal plate 3.6 mm. wide. A male 25 mm. long has the tenth plate 3.2 mm. wide. Tyre Locauity.— Washington (Pullman!), Known Locauities.—Washington (Wenatschee! Pullman!). Oregon (Corvallis!). This species is very close in general appearance and structure to 496 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. E. sierravagus Chamberlin; but the females are easily separated by the character of the gonopods. They differ ordinarily in the relative length of the ultimate article of the antennae; in the proportionate length of the prosternal dental line and the modal number of teeth ectad of the diastema; and in various other minor points. ETHOPOLYS INTEGER ALASKANUS Chamberlin. Plate 6, fig. 2. Rept. Can. Arctic exped. 1913-18, 1919, 3, pt. H., p. 21 H. While in E. integer the head and first dorsal plate are strongly and rather coarsely punctate, especially over the anterior portion of the head, these parts in the Alaskan specimens representing the present subspecies are smooth and wholly without punctae or very nearly so. Posterior angles of thirteenth dorsal plate weakly produced, those of the eleventh showing slightly a similar tendency. The claw of the female gonopods in the two type females has a dis- tinct tooth on the inner side toward the distal end but none on the outer, being thus bipartite instead of essentially entire as in integer, or tripartite as in sverravagus. In the two female types from Forrester Island a median dorsal black stripe is distinctly marked from the caudal end of the fourteenth plate cephalad to the frontal suture of the head. In one specimen the dor- sum is dark brown, dusky about edges of plates and with the head and most caudal plates of a dilute chestnut cast; the posterior legs testa- ceous, streaked with dark. The other specimen is much darker with the chestnut tinge affecting all the plates. The specimens from Sitka (males), presumably of the same sub- species, as they are at present wholly lack any reddish tinge, the color being a dull, nearly uniform, dusky olive brown. Dorsal spines of first legs in Sitka specimens 0, 0, 2, 2,1. Dorsal spines of second legs in specimens from both localities may be 0, 0, 3, 2, 1 OnOsOa, on 2 Length of max. 2, 23 mm. Type Locariry.— Alaska (Forrester Island!). Collected by Harold and Ronald Heath. Also taken in Alaska (Sitka!). CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 427 ErnHopo.tys puso (Stuxberg). Plate 6, fig. 3-6. Lithobius pusio Stuxberg, Ofvers. Kongl. vet. akad. Foérh., 1875, 32, no. 2, p. 66. Ann. mag. nat. hist., 1875, ser. 4, 15, p. 188. Lithobius (Archilithobius) pusio Stuxberg, Ofvers. Kongl. vet. akad. Forh., 1875, 32, no. 3, p. 16, 30. Proc. Cal. acad: sci., 1876, T. Bothropolys pusio Chamberlin, Can. ent., 1911, 43, p. 378. Ethopolys pusio Chamberlin, Can. ent., 1912, 44, p. 174. DescripTion.— Dorsum brown commonly of a reddish tinge or chestnut, head usually not darkened, being concolorous with dorsum; some of major plates in same specimens with the caudal border very dark and some with a median dark stripe. Antennae reddish brown, pale distad. Prosternum brown, with the prehensors lght, often rusty, yellow. Posterior ventral plates like the prosternum, the rest of the venter and the legs lighter brown, the caudal pairs of legs being commonly lighter, yellow, distad. The body from 7.6 to 8+ times longer than width of tenth plate, apparently more robust in the male. In the male widest at tenth plate, the body attenuated cephalad to the second segment; first plate narrower than the eighth. In the female the eighth and tenth plates are rather more nearly equal in the width and the attenuation of the body slighter. Head subcordate, the sides between the eyes and the rounded caudal corners weakly excurved and slightly converging caudad. Anterior margin arcuate, being convex at middle, then on each side concave, and then again convex adjacent to the antennae. Wider than long in about ratio 47 : 43 or nearly 12:11. The usual short, sharply im- pressed, curved sulcus extending mesad or cephalomesad from each lateral margin near the break. A short furrow near each caudolateral corner. Head finely and lightly punctate. Eyes consisting of from eleven to thirteen ocelli in three series: e.g. 1+ 5,3,2; 1+ 5,4,3; 1+ 4,4,2; 1+ 4, 4,3, which is fre- quent. Ocelli usually not deeply pigmented, greyish; decreasing in size ventrad. Single ocellus subcircular, but little larger than the adjacent one of the first row. Organ of Tomésvary in outline much smaller than the smallest ocellus, at anterior end of most ventral row of ocelli. Antennae short, reaching the sixth dorsal plate. Composed nor- mally of twenty articles which are moderately long and nearly uniform 428 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. in proportions. Clothed with moderately long and sometimes some- what rufous hairs. Prosternum one and two thirds times wider than long. Chitinous lines distinct. Mesal incision narrow, acute. Spine slender, distally bristle-like. One tooth ectad of diastema. Teeth mostly 6+ 6 or 6 + 7. Posterior angles of none of the dorsal plates produced. Third, fifth, eighth, tenth, twelfth, and fourteenth with the usual short, submargi- nal sulcus on each side near beginning of caudal third of length; the seventh with a similar but less distinct impression farther caudad and a second similarly weak one toward anterior corner. Plates finely rugose, most with two sublongitudinal and often broken furrows, one a little each side of middle, and sometimes, especially on the caudal segments, also with a median longitudinal furrow which may be doubled. First ventral plate with entire surface concavely depressed. Others with three longitudinal furrows, of which the median is rather short and often pit-like and lying near the middle portion of plate; some- times with two poorly defined transverse furrows which on the an- terior plates lie one near the anterior margin and one near the caudal, the anterior one lying farther caudad and the posterior one being more deeply impressed on the more posterior segments. On the posterior segments the median furrow becomes longitudinally elongate. On the fifteenth plate, especially in the male, the three longitudinal furrows are most distinct and converge commonly over nearly entire length of plate caudad to a less sharply defined transverse furrow in front of the caudal margin. Coxal pores as usual of various sizes, mostly small and very small. Arranged in two or three series and usually from seven or eight on the twelfth to twelve on the other coxae. Spines of first legs, o ° > z 5; of the second to the tenth, EEE of 1,0,3,2,2 g 0 ie Oe leat P ° the eleventh, 0:0°3'2'2; of the twelfth, ¢°7'333; of the thirteenth, 1, 0, 3, ONS sao : 3s q ie ea 3; of the penult, j 133-2, With two accessory claws; of the anal, LO NSs if 1 [1321- Last two pairs of coxae laterally armed. Anal legs of moderate length and slender in both sexes. Distal article of male gonopods considerably narrower than the first, apically rounded, with mostly two bristles; proximal article with four or five bristles. Claw of female gonopods rather short and broad, tripartite, moder- ately curved. Basal spines 3 + 3 or sometimes only 2 + 2, cylindrical proximally and conical in outline distad, flattened or excavated on CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 429 dorsal side and sometimes with small accessory points at junction of cylindrical and distal portions. Length mostly between 14 and 18 mm. A female 17.5 mm. long has the antennae 6.5 mm. and the anal leg cir. 5.5 mm. long, with the tenth plate 2 mm. wide. A male 15.2 mm. long has antennae 6 mm. and anal leg between 5.5 and 6 mm. long, with the tenth dorsal plate ~2 mm. wide. Praematurus.— Light brown or yellowish. Head light orange or hight brown. Antennae orange or brown but paler distad. Legs whitish yellow to pale brown with caudal pairs more deeply pigmented, clearer yellow. Prosternum darker than venter with prehensors in darker specimens testaceous. Ocelli mm most cases 1+ 4,3,2; also 1+ 3,3,3, etc. Single ocellus suborbicular or broadly elliptic, paler than the others, of same size as most caudal ocellus of the upper row. Ocelli decreasing in size cephalad and ventrad. Antennae composed of twenty long articles proportioned nearly as in the adult. Prosternum with chitinous lines obscure. Spine as in maturus. Teeth mostly 6 + 6; distally well rounded. Ectal ones more widely spaced than others as in the maturus. Impressions of head and dorsal plates already developed or indi- cated as in the adult. Coxal pores small, usually in one series, which may be irregular, on the twelfth coxae, and in two series on the others; from three or four, on twelfth coxae, to seven or eight on each coxa. . o ; 3,2,1 : : Spines of first and second legs, (7333; of the third to the ninth, 0,0, 3. 2, 2 1,0, 3,2, 2 ION SeINony eet(0, 3 1,2 0-02-32; Of the tenth, 99335; of the eleventh, (9-335 or ¢9:3°3°3; ie of the twelfth, eT of Te thirteenth, >? : ee of the penult, 3, 1, 3, +3; of the anal, Pes Eh 3. Claw of female gonopods rather slender, pale, distally a little curved, entire or with lateral teeth appearing as mere points at base. Basal spines 2 + 2, the inner one on each side scarcely one half the length of the outer Sud very much more slender. Length of specimen described 11.8 mm.; of antennae cir. 6 mm.; of anal legs czr. 4.8 mm. Immaturus.— Pale above and below, light violaceous brown, the anterior and the caudal segments orange. Head and antennae orange, the latter more yellowish, pale distad. Legs pale, brighter, yellowish, distad; caudal pairs light orange or yellow. Ocelli six to eight in two series: e¢.g., 1 + 3,2; 1+ 3,3; 14+ 4,3. Pale. The single ocellus larger than the adjacent ones. 1,0, 3 ‘ ibae . 430 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Antennae composed already of the twenty articles, of which part are relatively shorter than in the adult. Prosternal teeth 4 + 4 with the spine on the extended edge ectad of outermost tooth, no tooth ectad of spine as yet developed. Chitinous lines very distinct, complete. Coxal pores pale, small, in one series on the fifteenth or on this and the twelfth and in two on the others; in number from two (fifteenth) and four (twelfth) to six or seven (thirteenth and fourteenth) on each COxa. OOO 0, 0, x , . pS Wa Spines of first legs, 0-0-0:11; of the second to eighth, p-o-0-1-1; of 212 0,0, 2,2 0,212 00,3, 172 the ninth, 9 o, pest to Corsi; Of the tenth, ¢-g-7-7 OF ong 224; of the BS 0.03 4,5 DiOTes ste tO easied eleventh, ¢-y: ie aa OF 00.22.13 of the twelfth, 9-9-3354 or o-o-s31; of the i, 6,2, 1,1 IONS Isle Ube iki thirteenth, pg3t7 to gaoo3; of the Seals 0-13-29; of the anal, UO; Sek 10 . 0 13-10, the lateral and dorsal spine of coxa also present, the lateral being small. Gonopods in male appearing as simple, smooth, rounded prominences which are not yet biarticulate. Gonopods of female triarticulate. Claw short, very pale, acutely pointed, with no signs as yet of lateral teeth. Basal spines 1 + 1, small; slender. Bristles very few. Length, 6.5 to 8 mm. Type Locauiry.— California, near San Francisco (probably Sausa- lito). Known Locauities.— California (Sausalito! Mill Valley! Brook- dale! Santa Cruz Island! Capitola! Felton Big Trees!). Stuxberg’s types are evidently of the 7mmaturus stage, a little later than those above described, as elsewhere indicated (Chamberlin, Can. ent., 1911, 43, p. 379). ETHOPOLYS MONTICOLA (Stuxberg). Lithobius monticola Stuxberg, Ofvers. Kongl. vet. akad. Férh., 1875, 32, no. 2, p. 65. Ann. mag. nat. hist., 1875, ser. 4, 15, p. 188. Lithobius (Archilithobius) rontieole Stuxberg, Ofvers. Kongl. vet. akad. Forh., 1875, 32, no. 3, p. 14, 80. Proc. Cal. acad. sci., 1876, 7, p. 137. Bothropolys monticola Chamberlin (ex part. minor), Pomona college journ. ent., 1910, 2, p. 369. Ethopolys monticola Chamberlin, Can. ent., 1912, 44, p. 174. DerscriptTion.— Lamina cephalica subcircularis, eadem fere latitu- dine ac longitudine, margine postico subrecto, setis punctisque impressis sparse praedita. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 431 Antennae mediocres, articulis 20 maximam partem cylindraceis, setis rigidis vestitis compositae. Oculi longitudine triplo majore quam altitudine, ocellis 7-9 in 2 series longitudinales digestis. Coxae pedum maxillarium secundi paris dentibus 6 + 6 conicis, acutiuculis, nigerrimis armatae. Scuta dorsualia rugulosa, sparsissime pilosa, 2°, 4°, 6°, 7°, 9°, 11°, 13° margine postico recto, angulis posticis rectis vel rotundate rect- angulis, 1°, 3°, 5°, 8°, 9°, 12°, 14° margine postico elevato sinuato, angulis parum productis, rotundatae acuminatis. Seutum ventrale 15 mm. fovea longitudinale profundiore, cetera omnia plana. Pori coxales numerosi, in 3-4 series irregulares dispositi. Pedes primi paris calearibus 2,3, 2. Pedum analium articulus primus calearibus binis, altero majore inferiore, altero minore laterali armatae. Pedes anales ungue singulo, calearibus 1, 4, 3, 1 — 1, 4, 3, 2 armati. Color non manifestus. Longitudo corporis 18 mm. Stuxberg. Hasirat.— Sierra Nevada (G. Eisen). The identity of this species is at present a matter of considerable doubt. When it appeared that only one species of the Ethopolidae with none of the dorsal plates produced occurred in California, its identity with that form seemed probable; but the discovery of adult E. pusio makes this doubtful. The ocelli are given as seven to nine in two series. This agrees neither with adult pusio nor young sverra- vagus. If it be assumed that Stuxberg failed to note the most ventral row of eyes, then the agreement is close with adult pusio while the number would be still too small for s¢erravagus. The number of pro- sternal teeth agrees with pusio but is too small for sterravagus of the length given, so far as noted by the author. The spine-formula as given for the anal legs does not agree with that of either species as it stands. It is quite possible, indeed, that monticola is not the same as either of the species, and the only course open at present, therefore, is so to regard it. Should pusio and monticola prove to be the same, the latter name would have precedence. 432 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ErTHopotys (?) CALIFORNICUS (Daday). Lithobius californicus Daday, Termes. fiizetek., 1889, 12, p. 153. DeEscrRIPTION.— Corpore sat robusto, flavescenti; capite satura- tiore colorato; antennis pedibusque dilute flavescentibus; oculis utrinque 14 in seriebus tribus positis; coxis pedum maxillarium denti- bus 8 armatis; laminis dorsalibus inermibus, leviter punctatis; poris coxalibus multi seriatis, parvis rotundis; pedibus analibus sat crassis, ungue singulo; infra calcaribus 0, 1, 4, 3, 1, in articulo primo calcari- bus duobus lateralibus armatis; calcarium genitalium femineorum tribus paribus. Longit. corp. 20 mm. Latit. corp. 3 mm. Patria: California borealis. Specimen unicum in spiritu. vini rectif. conservatum a D. Joanne Vadonna collectum. This species is placed under Ethopolys rather than Bothropolys because of the presence of 3 + 3 basal spines on the female gonopods. However. if a ventral spine is absent from the anal coxae as represented, it probably belongs to neither of these genera; but it is possible that one of the two lateral spines mentioned is the real ventral spine. The species is readily separated by its robustness and the small number of prosternal teeth 4-+ 4. It is impossible, however, on account of the insufficiency of the description and of our knowledge of the myriopod fauna of a broad strip of country along the coast of California north of San Francisco, to say anything more definitely of this form without study of the type. The unusual spining given for the anal leg is probably due to including a dorsal spine on third and fourth articles, as it is probable Stuxberg did in the case of monticola. ETHOPOLYS sens. str. ErHopoLys XANTI (Wood). Plate 4, fig. 1-6. Bothropolys xanti Wood, Journ. Acad. nat. sci. Phil., 1863, new ser., 5, p. 15. Trans. Amer. philos. soc., 1865, 18, p. 152. Chamberlin, Pomona coll. journ. ent., 1910, 2, p. 369. Chamberlin, Can. ent., 1911, 48, p. 378. Lithobius xanti Stuxberg, Ofvers. Kongl. vet. akad. Férh., 1875, 32, no. 3, p. 10, 27. Proc. Cal. acad. sci., 1876, 7, p. 185. Bollman, Proc. U. S. N. M., 1887, 10, p. 261. Chamberlin (ad part. min.), Proc. U. S. M., 1901, 24, p. 24. Proc. Acad. nat. sci. Phil., 1902, p. 42. Ann. Ent. soe. Amer., 1909, 2, p. 190. Ethopolys xanti Chamberlin, Can. ent., 1912, 44, p. 173. CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 433 DescriptTion.— Brown to rusty brown and ferruginous, sometimes very dark, in part suffused with blackish. The head and the first one and usually one to several caudal plates darker and more reddish. Venter and most legs clear brown, the tarsi of the legs more brightly colored, the posterior sternites and legs darker, ferruginous or ferru- gineotestaceous. Prosternum and_ prehensorial feet somewhat ferrugineotestaceous. Antennae usually dark brown, sometimes of a reddish tinge, a little paler at tips. Body mostly from 8 to 8.6 times longer than width of tenth plate, ot nearly same size and proportions in both sexes. Eighth and tenth plates of same width, with the first but little narrower. Head wider than long in about ratio 73:70. Of usual shape, the sides a little converging from behind eyes caudad. Caudal margin straight or very slightly incurved mesally. Anterior margin in form of three convex lines or crenations of which the median is much smaller. Nearly smooth, polished, very finely punctate; the usual distinct, short, transverse submarginal sulcus on each side near marginal break. Ocelli from ten to eighteen in mostly four rows: e. g., 1 + 5, 5, 4, 2; 1-5, 4,4,.2; 14+ 4,4, 3,1; 1+ 5,4,38,1. Single ocellus very large, obliquely elliptical, pale. Seriate ocelli decreasing in size ventrad; series very regular. Organ of Tomésvary a little in front of anterior ocellus of most ventral row and very small in outline. Antennae long, reaching usually to the eighth segment of body. Articles normally twenty, all long or very long, not densely clothed with rather short hairs. Prosternum cir. 1.67 times wider than long. Chitinous lines in- complete below. Mesal incision narrow and rather deep, the sides < little converging caudad, and the bottom narrowly rounded. Spine acuminate, slender distally as usual, inserted on very anterior edge of diastema in adults. Either two or three teeth normally ectad of diastema. Teeth varying in number from 6 + 7 to 10 + 10 (Plate 4, fig. 2). The first, third, fifth, eighth, tenth, twelfth, and fourteenth plates with the usual short submarginal sulcus on each side at about beginning of the caudal third; on the seventh plate a similar impression near each caudal and each anterior corner, the tenth having also a mark at each anterior corner but this absent or obscure on the others. Except- ing the first all the dorsal plates are very much roughened with tubercu- lar and rugose elevations. Each major plate excepting the first with a median longitudinal furrow and on each side one running from clese to anterior end of this obliquely ectocaudad and further laterad another 434 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. furrow subparallel with margin and meeting or crossing the oblique furrow at an acute angle. Posterior angles of ninth, eleventh, and thirteenth dorsal plates strongly produced. Ventral plates with the usual three furrows which are most distinct on the caudal plates, the median mostly deepest, but all, and especially the median, becoming less and less distinct cephalad. On the fifteenth plate the three furrows are very deep and end abruptly in front of the smooth, pilose caudal border which covers about one fourth the length of the plate; the furrows not converging. On the fourteenth plate there is a transverse furrow in front of the caudal border at which the longitudinal furrows, which are less distinct than on the fifteenth, and especially so in the female, terminate or the medium sometimes crossing to the caudal margin. Coxal pores small, less varying in size than in related species; those of the caudal row frequently but slightly larger than the others, the distal largest. From nine to thirty in number, and mostly in three, but often in four series, the number of pores in the rows not so much differing as usual. Spines of first legs, 0 333; of the second to the ninth, (9-33; of the tenth, °° c oe a8 Bo of the eleventh, } 0: 0 3 ee 2 of the twelfth, ; 0, 3, 2 1 TONSHON2 : 50.8) Bale . 013,32; Of the thirteenth, 9-73:3:3; of the penult, 7 + 33-2, with two small accessory claws; of the anal, t ae , the claw single. Last two pairs of coxae laterally armed. Anal legs long and slender in both sexes. The third and more distal articles longitudinally suleate dorsally; ventrally the articles may show a longitudinal furrow each side of middle leaving a broad area between them. Distal article of male gonopods much narrower than the proximal, long and cylindric in shape, distally broadly rounded; bearing mostly five or six bristles. Proximal article bearing on ventral surface seven or eight bristles. Claw of female gonopods strongly curved, moderately long, acute, the lateral teeth small and borne well proximad, sometimes almost obliterated. Basal spines 3 + 3 or 4 + 4, or very rarely only 2 + 3 or 2 + 2; stout, conical, decreasing in size mesad on each side; when 4 + 4 the innermost on each side may be attached on mesal side where it is easily overlooked. Length from 19 to 26 mm. in both males and females. A male 25 mm. long has the eighth and tenth plates 3.1 mm. wide, with the first 3 mm. wide; anal legs 10 mm. long; and the antennae 11 mm. long. or CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE. 435 Pseudomaturus (early).— Brown. Head and antennae orange, the latter paler at tips. Legs lighter over tarsi as usual, the anal and penult pairs testaceous or pale ferrugineotestaceous. Markings of head and dorsal plates fully developed. Ocelli 1 + 4,4,2; 1+ 4,3,2. Single ocellus large and elliptic. Antennae long; composed of twenty long articles as in the maturus. Prosternal teeth 2 —5 + 5 — 2 Coxal pores all small and very small toward, those of caudal side exceeding the more cephalic ones more than in the maturus, especially on the coxae with the fewer pores. Mostly from six to twelve in number in two (twelfth coxae) or three rows 0,0, 3, 2,1 Spines of first to tenth legs, 0:0:2'3°2; oP the eleventh, 4 0,2, 3 OF ; (te 35 3-5; of the twelfth and thirteenth, ise; of the ee i e533 of the anal, }):-3:3-4. Claw shores. and paler, especially proximally, and thinner than in the maturus, the lateral teeth about the same. Articles with some- what fewer bristles. Basal spines 3 + 3, but the most mesal on each side very slender, attached on mesal surface and more dorsad than the others; the median spine relatively somewhat less robust than in maturus. Gonopods of male pale, the basal joint cylindric and the distal one conic. Distal article with one or two bristles, the proximal with two or three. Length 15-17 mm. Praematurus.— Light violaceous brown throughout, the head and the first segment and the last several segments darker; dorsal plates usually darker along the caudal and sometimes the lateral borders. Head lighter in front of frontal suture, dusky caudad of suture and about the eyes. Prosternum and prehensors somewhat orange, the latter distally somewhat rufous. Legs and venter paler than dorsum; anal legs rusty yellow. Ocelli 1 + 4,3, 2. Distinct; the most ventral much smaller; the single one as usual. Antennae long. Articles 20, long. Prosternum with chitinous lines very distinct. Teeth already —5+5-2o0r2—-5+6- Coxal pores six to eight in two series on each coxa, those of cephalic row few and often minute, sometimes close and nearly merged. Spines of first legs, $331; of the second to fourth, ¢°0°3°3'2; of the fifth to tenth, ? Oo os 35; of the eleventh, 0-0, 3,23 of che twelfth and thirteenth, sa: of the penult, ¢- 7:33; of the anal, Pr3°s4- 436 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Claw of female gonopods pale, short and thin; median lobe acute, the lateral plainly indicated. Basal spines 2 + 2; the mesal one on each side slender, acute, and not more than one half the length of the outer one; the outer one moderately stout; both pale. : Gonopods of male with distal article very small, conical, the proxi- mal rounded and narrowed distad; distal articles with one or two bristles, the proximal with two or three. Length 12-15 mm. Immaturus.— Pale yellowish brown, with violaceous showing through from beneath. Head and caudal segments pale orange or rusty yellow, the head darker caudad of the frontal suture. Legs pale, light tipped, the last pairs more yellowish. Ocelli 1 + 4, 3,1, the one in the most ventral row very small. Single ocellus scarcely larger than adjacent one of first row. Antennae already with twenty articles, which are long as in the adult. Prosternal teeth 1 —5-+5—1. Spine attached a little caudad of edge on ventral surface. Coxal pores small, with pale margins, four to eight on each coxa in two series. Ls : 0,0,1,1.1 0,0,2.1,1 0, 0.2, 2,1 Ke joe of first legs, 9°9:2°974 a 0.9.2 37; Of the second, 9 0,3 oo 0-0'2,3,1; Of the third to sev enth, | peas; of the eighth to tenth, j-3:3:5; 5 10,322 eee of the eleventh, 9:9:3'3' 23, of the twelfth, 94°3:3:3; of the thirteenth, 1, 0, 3, 1,2 0,3, 1,1 10/321, 0 OmINge3: 25 of the penult, | iE L 8, 3,20 ol the anal, Tae a lc Claw of female gonopods yellow, small, nearly straight, the lateral teeth indicated as acute points near its base. Basal spines | + 1, straight, pale. Length cir. 11 mm. (Laurel Canyon, Los Angeles Co., Cal.). Agenitalis IT.— Violet-brown. The frontal region of head pale orange; caudal portion dusky. Legs pale; caudal pairs, especially distally, bright whitish yellow. ~ Ocelli 1 + 3, 2, the single ocellus equal in size to the adjacent one of the upper row. The twenty articles of antennae, but few of the articles not already of the full proportionate size. Prosternal teeth 1 — 5 + 4 — 1, the extra tooth on side having six teeth located on upper part of mesal incision and reduced in size. Coxal pores from three to five in number; in mostly one, but some- times in two, series. Opagaivek Sus of the first legs, f-G4274 to pest of the second to the fifth, ee e 3 31; Of the sixth to the tenth, ; oft the eleventh, eee . 0,0, 3, 1, 2 of t he twelfth, 9-9-3313; of the MGateen h, egg OF eee; of the 0, 0, 0, 0, CHAMBERLIN: NORTH AMERICAN ETHOPOLIDAE, 437 ONOMS alent 0, 0, 3, 1, penult, 9, IEHEE 10 OL; (misgt952 5 of the anal, two pairs of coxae ev ident. Anal ae no longer evident. Length cir. 9.5 mm. (Laurel Canyon, Los Angeles Co., Cal.). Agenitalis ferns The head in front of suture and the caudal segments yellowish. Legs pale, commonly brighter at tips; caudal pairs from white to yellow. Prehensors yellowish, brighter distad. Antennae violaceous, yellowish at tips. Ocelli 1 + 2, 2, the more caudal one in each of the two series con- siderably larger than the anterior one, dusky or brownish in color. Single ocellus not larger than the adjacent one of dorsal row. Organ of Tomésvary minute in outline, well separated from the ocelli. In antennae of three specimens examined twenty articles are already present; in one specimen seventeen articles are present in right and twenty in left; and in another specimen sixteen are present in right and twenty in left. Articles varying in length, articles short and intermediate in size occurring at intervals. Coxal pores small and pale margined; two on twelfth coxae, and one each of the others. Dorsal plates with posterior angles of ninth, eleventh, and _ thir- teenth dorsal plates produced, but those of the ninth only slightly so and the processes distally rounded. Hairs of body and legs very sparse, the seriately arranged ones of ventral surface of tarsi more numerous. 108 351.10 = 1321. Lateral spines of last ae 0,0, 1, 1 a 0, 2,21 Spines of first legs, 9°0°0'1:1 OF oc-at-2 ; of the secon d to sixth, 99-343 29 2, 2, 2 of the seventh and eighth, 5 0,0, 2, 2.4 - of the ninth, ? ; of the tenth, ee ae yee 0011 2 hue eleventh, :}:7474 ie Oo ins twelfth, j-o-¢:74; of the thir- teenth, oo-i11; of the penult, COLL1, the two Aa claws dis- . 0, 0, 0, 0, 0 . : . " . . tinct; of the anal, ;°9'7:19, the trochanter being without spine or bristle. Anal glands still manifest. Length 7-8 mm. Type Locauiry.— California (Fort Tejon). Known Locatities.— California (San Bernadino Co.! Los Angeles! Laurel Canyon! Santa Monica! Claremont! Santa Barbara! Monterey! Pacific Grove! Cypress Point! ete., Stanford! Sausalito! Brookdale! Santa Barbara! Santa Cruz Island! ete.). Utah! This is an abundant species in California along the coast and in the coast mountains from San Francisco to the southern part of the state, where it ranges in part with FE. pusio. In the Sierras and northward it is replaced by E. sierravagus and E. integer; while in the Wahsatch Mountains of Utah it is almost wholly replaced by Bothropolys per- mundus, but there maintains a foothold in a number of limited areas. EXPLANATION OF THE PLATES. " eh 2 ie CHAMBERLIN.— North American Ethopolidae. Fig. 1. PLATE 1. Bothropolys multidentatus (Newport). Part of border of head, showing the continuous lateral margination. (o Ithaca, IN. Y-)- Fig. 2. Portion of second maxillae showing character of median sinus and membranous bridge. Fig. 3. Fig. 4. Fig. 5. Fig. 6. Fig. 7. Fig. 8. (Illinois). 3’ gonopod, lateroventral view. (Kentucky). Claw of @ gonopod. (Illinois). Basal spines of 2 gonopod. (Illinois). Spine and most ectal tooth of prosternum of adult 9. (Illinois). Same of an adult o. (Ithaca, N. Y.). Same as an immaturus showing relatively greater length of spine. BULL. MUS. COMP ZOOL. ETHOPOLIDAE. PLATE 1 PLATE 2. CHAMBERLIN.— North American Ethopolidae. PLATE 2. Bothropolys permundus (Chamberlin). Fig. 1. Prosternal teeth and spines of adult, the latter with persistent apical portion. (Provo Canyon, Utah). Fig. 2. Prosterna] spine and outermost tooth of an adult co, showing ordinary form of spine in the maturus. Fig. 3. Prosternal spine and outermost tooth of an immaturus, showing the tip still present. Fig. 4. Proximal portion of anal leg in adult, showing ventral coxal spine and coxal pores. Fig. 5. Same in an immaturus, showing the pores still uniseriate. Fig. 6. Claw of @ gonopod, ordinary form. Fig. 7. Claw of 2 gonopod with teeth obliterated. Fig. 8. Basal spines of @ gonopod, typical form. Bothropolys hoples (Brolemann) Fig. 9. Claw of 9 gonopod. PLATE 2 ETHOPOLIDAE. BULL. MUS. COMP. ZOOL. Ns re Py par a) rt ae A ees ae | CuHamBERLIN.— North American Ethopolidae PLATE 3. Bothropolys hoples (Brolemann). Fig. 1. Claw of 2 gonopod, showing teeth practically obliterated. (Brit- ish Columbia). Fig. 2. Basal spines of a 9 gonopod. (Flathead Lake, Montana). Fig. 3. Same, more robust form. (Portland, Oregon). Fig. 4. Outline of one side of ninth dorsal plate showing form of posterior process. Bothropolys victorianus Chamberlin. Fig. 5. Outline of one side of ninth dorsal plate showing form of posterior process. Fig. 6. Basal spines of Q gonopods. (Type). Zygethopolys nothus Chamberlin. Fig. 7. Claw of 2 gonopods. Fig. 8. Basal spines of 2 gonopod. BULL. MUS. COMP ZOOL. ETHOPOLIDAE. PLATE 3 > bd ye z nm et C#amBertin.— North American Ethopolidae. PLATE 4. Ethopolys xanti (Wood). Fig. 1. Border of head, showing discontinuity or “break” in lateral margining. (Pacific Grove). Fig. 2. Prosternal teeth and spine of maturus. (Pacific Grove). Fig. 3. Same of immaturus, same magnification as for Fig. 2. (Laurel Canyon). Fig. 4. Prosternal teeth and spine of left side more highly magnified. (Monterey). Fig. 5. Basalspines of 2 gonopod. (Santa Cruz Island). Fig. 6. Gonopod of o. (Pacific Grove). BULL. MUS. COMP ZOOL. ETHOPOLIDAE. PLATE 4 —~ = Ure re a ow oh - : = ae! ® a I 7) 7 ={ ea - a 7 ; “, - : — 7 —— ior . . J > Tos ve a eed eee taeda tl aie, os me _ _ al AL 4 a OLE el, ; _ =, ” i a 7 vs wn r , : a Ba as <7 CuamBertin.— North American Ethopolidae. PLATE 5. Ethopolys sierravagus (Chamberlin). Fig. 1. Claw of @ gonopod. (Shasta Springs, Cal.). Fig. 2. Basal spines of 2 gonopod. (Portland, Oregon). Fig. 3. Terminal portion of an antenna composed of twenty-four articles, showing form of distal articles. (Shasta Springs, Cal.). Fig. 4. Same of antenna composed of the normal twenty articles. (Shasta Springs, Cal.). Ethopolys bipunctatus (Wood). Fig. 5. Claw of @ gonopods. (Utah Co., Utah). Ethopolys integer Chamberlin. Fig. 6. Terminal portion of normal antenna showing form of distal articles. Fig. 7. Prosternal spine and teeth of left side. Fig. 8. Claw of 2 gonopod. (Pullman, Wash.). ETHOPOLIDAE. PLATE 5 BULL. MUS. COMP. ZOOL. CHAMBERLIN.— North American Ethopolidae. PLATE 6. Ethopolys integer Chamberlin. Fig. 1. Outline of right end of thirteenth dorsal plate showing usual form of posterior corner. Ethopolys integer alaskanus Chamberlin. Fig. 2. Outline of right end of thirteenth dorsal plate showing form of posterior corner. (Sitka, Alaska). Ethopolys pusio (Stuxberg). Fig. 3. Entire gonopod of 9. (Topotype. Sausalito, Cal.). Fig. 4. Gonopods of &. (Topotype. Sausalito, Cal.). Fig. 5. Same, right side, of immat., more highly magnified, ectal tooth not yet developed. (Mill Valley, Cal.). Fig. 6. Prosternal teeth and spine, left side, of maturus. (Topotype. Sau- salito, Cal.) BULL. MUS. COMP. ZOOL. ETHOPOLIDAE. PLATE 6 Bulletin of the Museum of Comparative Zotlogy AT HARVARD COLLEGE. Vout. LVII. No. 8. THE GENERA LITHOBIUS, NEOLITHOBIUS, GONIBIUS, AND ZINAPOLYS IN AMERICA NORTH OF MEXICO. By Raureu V. CHAMBERLIN. With THREE PLaAtTEs. CAMBRIDGE, MASS., U. S. A.: PRINTED FOR THE MUSEUM. JUNE, 1925. | ere - aniiizoo. yes oe : . | CorKam ata) urson a nasa yi 3 iM | “lsiahn eMC Fanaa ht ve oe all whi /? j 5 a LA PU Ae DOM A MUU Pah Rod Ca we Sout eA No. 8.— The Genera Lithobius, Neolithobius, Gonibius, and Zinapolys in America North of Mexico. By Ratpen V. CHAMBERLIN. THE genera revised in this paper include all the species of larger size in the Lithobiidae in the restricted sense occurring in America north of Mexico. In the West and the Southwest these forms seem nearly wholly to be replaced by the similarly large species of the Ethopolidae and Gosibiidae. The paper was in manuscript in its present form eleven years ago, and is published unchanged excepting for the addition of the descrip- tion of Neolithobius helius Chamberlin, the omission of diagnoses. preceding the descriptions, and minor additions to locality records, and bibhography. Key to Genera. a. Coxae of last two pairs of legs ventrally armed; (posterior angles of none of the dorsal plates produced); claw of female gono- pods entire and the basal spines numerous (e.g., 6-+6); articles Omantennae 20.0.5. .28 Ace ee Mes Zinapolys Chamberlin. aa. None of the coxae ventrally armed; posterior angles of three or more of the dorsal plates produced; claw of female gonopods partite, the basal spines 2 + 2 or 3 + 3. Gs, Aviicles ofantennae 20k. 0.5 ¢s.% Ve. ee. Gonibius, gen. nov. bb. Articles of antennae more than 20, mostly 25-45. c. Posterior angles of only the ninth, eleventh, and thirteenth dorsal plates produced; anal legs of male not conspicu- ously moditied. 0) H... Mee. Lithobius Leach (emend.). ce. Posterior angles of seventh, ninth, eleventh and thirteenth dorsal plates produced; fourth article of anal legs of male specially thickened and with its dorsal surface or else the entire article bowed ventrad. Neolithobius Stuxberg (emend.). . ZINAPOLYS Chamberlin. Can. ent., 1912, 44, p. 14. Antennae short; articles twenty. Eyes composed of seriate ocelli. Organ of Témésvary large in outline. Prosternal teeth 5+ 5 to 7+ 7, without diastema on each side. 442 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Fic. 1.— Distribution of Zinapolys Chamberlin. Spine slender, situated caudad of ectal tooth on each side. Posterior angles of none of the dorsal plates produced. Coxal pores uniseriate; present on last four pairs of coxae. CHAMBERLIN: GENERA OF LIFHOBIIDAE.!. : 443, Claw of female gonopods strictly entire, deeply excavated on mesal side. Basal spines in type 6 + 6(5). Gonopods of male large; biarticulate. Tarsi of all legs bearing three claws. Coxae of last two pairs of legs armed ventrally and laterally as well as dorsally. Third joint of all legs armed with three dorsal spines. Fifth joint of first thirteen pairs of legs with two dorsal spines. _ Dor- sal spines of anal legs 1, 0, 3,1, 0; ventral 1, 1,3,3,1(2). Dorsal spines of penult legs 1, 0,3, 1,1; ventral 1, 1, 3,3, 2. Dorsal spines of twelfth and thirteenth legs 1, 0, 3, 2, 2; ventral 0, 1, 3, 3, 2. Length of type species 17-20 mm. Typre.— Z. zipius Chamberlin. Outside of the Ethopolidae this is the only known American member of the Lithobiomorpha having any of the coxae ventrally armed. It is unique in the large number of basal spines borne by the female gonopods. The precise affinities of the genus seem at present quite problematical. Only the type species is known. ZINAPOLYS zIprus Chamberlin. Plate 1, fig. 1-3. Zinapolys zipius Chamberlin, Can, ent., 1912, 44, p. 14. DescripTion.— The original color of types, now largely bleached, seems to have been as follows:— dorsum ferruginotestaceous with the caudal segments somewhat darkest; venter and most of the legs more yellowish, the last pairs of legs darkest, the last ventral plates orange or light ferrugimous; prosternum and head nearly like last ventral plates; antennae a little darker than legs but paler than head. Body narrowed considerably cephalad from the tenth to the second segment and conspicuously caudad as usual. Widths of first, third, eighth, and tenth plates to each other as 19 : 18 : 21 : 21.5. Head wider than long in about ratio 15:13. Anterior margin between antennae nearly straight. Head back of eves subquadrate, the sides a little converging caudad to the rounded posterior corners. The elevated caudal marginal thickening not. widening or extending cephalad at middle. Head depressed along frontal suture. A furrow a little caudad of and parallel with the frontal suture. Commonly a median furrow is present with one diverging from it on each side cephalad from the caudal margin. Ocelli mostly about twenty in four series: e.g., 1 +5, 5, 5, 4; 444 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 1+ 5, 5, 5, 3. Single ocellus not very large. Organ of Témésvary large, in outline exceeding an ocellus; not contiguous with eye-patch. Antennae short, reaching the fifth or the sixth segment; composed of twenty articles of medium or short length, the second one being much the largest and those immediately proximad of the ultimate one often very short. Hairs not dense; longer than usual. Prosternum wider than greatest length in about ratio 5:3; 7.¢., 1.66 times wider than long. Anterior margin extending cephalad of ectad from middle on each side. Prosternal teeth 5 + 5 to 7+ 7; large, decreasing in size from the most ectal to the most mesal on each side. Spine rather stout at base but acuminate and distally long and slender as shown in the figure. Major dorsal plates mostly showing a median longitudinal furrow and a little ectad of it on each side one diverging from it from the anterior margin caudad. A depression or wide furrow within each lateral marginal thickening and from the anterior end of this furrow usually a branch extends mesocaudad. A distinct transverse depres- sion a little distance in front of caudal margin, this uniting laterally with the longitudinal furrow on each side. Caudal margin of ninth plate at each end bent obliquely somewhat cephalad; posterior margins of eleventh and thirteenth dorsal plates straight. Ventral plates with each lateral border bent somewhat ventrad, a furrow marking the line of bending; at least on caudal plates a median longitudinal furrow may be detected. Most plates showing several transverse, irregular impressed lines or furrows of which one or two are ordinarily more pronounced than others. Coxal pores rather large, very distinct; from circular to transversely elliptic or suboblong in outline;. arranged mostly in order 4, 5, 5, 5 or 4,4, 5,5 Gonopods of male moderately large; the proximal article thick, cylindric, with four or five ventral bristles in a transverse row near distal edge. Distal article much narrower than the proximal, conical; bearing mostly two or three long bristles. Claw of female gonopods proportionately short and broad; deeply excavated or hollowed out on mesal side. Basal spines 6 + 6 or 6 + 5; stout but rather short, subconical. Spines of first to seventh pairs of legs, (is 323 of the eighth to eleventh, }y733'3;_ of the twelfth and thirteenth, j-7:3:3:3; of the penult, 130.8515 0, 3, 0, 3, 1, 0 : i333; of the anal, ps3} or peeeoy- Cl legs with three claws. Last two pairs of coxae laterally armed. Length 17-20 mm. A male 20 mm. long has the tenth plate 2.8 mm. CHAMBERLIN: GENERA OF LITHOBIIDAE. 445 wide; the antennae 8 mm. long; and the anal leg (exclusive of coxa as usual) 7.8 mm. long. Locauiry.— Idaho: Kootenai Co.! The types of this interesting species embrace five adults. GONIBIUS, gen. nov. Antennae moderate to long; articles twenty. Ocelli numerous, seriate; single ocellus differentiated. Prosternal teeth numerous (6 + 6 to 9 + 9). Spine ectal in po- sition. No diastema in dental series of each side. Posterior angles of fourth, sixth, seventh, ninth, eleventh, and thirteenth, or of only the last three mentioned, dorsal plates produced. Coxal pores large, uniseriate. Claw of female gonopods tripartite. Basal spines 2 + 2; broad. Gonopods of male uniarticulate (gly ptocephalus). None of coxae ventrally armed; some of posterior pairs armed laterally as well as dorsally. Third joint of all legs with three dorsal spines, that of some of anterior ones with but one or two ventral ones, some following with two and only the mere caudal with three. Fifth joint of legs from second or about fourth to thirteenth or fourteenth pairs of legs with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0; ventral, 0, 1, 3, 3, 2 to 0, 1, 3, 2, 1; claw one. Dorsal spines of penult legs Ta) 3,.2, 2 or 1 0, 3,1, 1: ventral 0; 1, 3,/3; 22 claws two or three. Dorsal spines of thirteenth legs 1, 0, 3, 2, 2; ventral 0, 1, 3, 3, 2. Ventral spines of twelfth 0, 0, 3, 3, 2 or 0, 1, 3, 3, 2; dorsal 1, 0, 3, 2, 2 or 0; 043,.2, 2: Length 18-20 mm. GEnotyPE.— Lithobius rex Bollman. Of the two species grouped in this genus, one, the type, is known only from the female and the other only from the male, an unfortunate circumstance. Analogy makes it seem likely that the two species should go into separate subgenera on the basis chiefly of the difference in angulation of the dorsal plates, the type constituting Gonibius sens. str. while glyptocephalus may be included in a new subgenus Tambius. 446 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Fic. 2.— Distribution of Gonibius Chamberlin. Key to Species. a. Posterior angles of fourth, sixth, seventh, ninth, eleventh, and thirteenth dorsal plates produced; ventral spines of anal legs 0, 1, 3, 3, 2; dorsal spines of penult legs 1, 0, 3, 2, 2. G. (Gonibius) rex (Bollman). CHAMBERLIN: GENERA OF LITHOBIIDAE. ‘ 447 aa: Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced; ventral spines of anal legs 0, 1,3, 2, 1; dorsal spines ~ of penult legs 1, 0, 3, 1, 1. G. (Tambius) gly ptocephalus (Chamberlin): GONIBIUS REX (Bollman). Plate 1, fig. 4, 5. Lithobius rex Bollman, Proc. U.8. N. M., 1888,11, p.350. Bull. 46,U.S.N.M., 1893, p. 102. DescripTion.— Original color difficult to make out from type. Apparently greyish brown with head, antennae, and the first dorsal plate and margins of the others darker. Legs light brown or testaceous. Body robust, in the type being at present only 6.25 times longer than the width of the tenth plate (while the body may have shrunk through drying, the length of 25 mm. given by Bollman in the original descrip- tion was doubtless too large, as seems to have been true of most meas- urements given by that author for lithobiids). Widths of head and of first, third, eighth, and tenth plates to each other as 88 : 80 : 76: 88 : 88. Head distinetly wider than long (ratio 11 :9), considerably wider than first dorsal plate but same as width of eighth and tenth plates. Caudal margin nearly straight. Surface uneven, densely, finely punctate. Ocelli 18-21 in four or five series: e.g., 1 + 5, 5, 4, 3(4), 1+ 5, 6,5, 3,1. Single ocellus large, subcircular. Other ocelli also com- paratively large and distinct, those of the most dorsal row being largest as usual. Antennae long, reaching well upon the tenth body segment. Com- posed of twenty articles all of which are long or very long. Prosternum about 1.87 times wider than long. Distance between chitinous spots only 1.25 times width at level of bottom of median sinus; two times the length of the dental line. Teeth rather small, nearly uniform, distally narrowly rounded, 9+ 9 in number. An- terior margin wide; each side weakly convex, with mesal end a little farther caudad than the ectal. Spine ectal in position as usual, slender and straight, acute. First dorsal plate like head; the others very strongly roughened. Posterior angles of the fourth, sixth, seventh, ninth, eleventh, and 448 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. thirteenth dorsal plates strongly produced, .those of the last three mentioned being especially long and acute. Coxal pores large, strongly transverse, being narrowly oblong or sometimes keyhole shaped : 8, 8, 8, 7. Claw of female gonopods short and broad; tripartite, the two more mesal lobes nearly equal in length with the median one but little the longer, the ectal lobe smallest. Basal spines 2 + 2, very wide, some- what clavately widening to distal third which is abruptly narrowed to an acute point (Plate 1, fig. 5). Mesal side of proximal article straight or nearly so. Anal legs of female long and slender, the distal articles especially slender. Fifth article or tibia much longer than the fourth article or femur. Ratio of lengths of fourth, fifth, sixth, and seventh articles being to each other as 66 : 79 65): 40. Spines of first and second legs, 9 oia3; of the third either the same or Feige; of the fourth, (p54; of the fifth to eighth, oees; Of the ninth to eleventh, i AB. at of the twelfth, ETE of the thir- 2. teenth and fourteenth, j{' 3 ee the latter with three claws of which the anterior accessory is very small; of the anal, ('{'3'3°s, Claw one. Last four pairs of coxae laterally armed. Length of type (2 ) 20 mm.; width of tenth plate 3.2 mm.; length of antennae 13 mm.; length of anal legs 12 mm. Locauiry.— Georgia: Tallulah! Only the single type female above described is known. Gontipius (TAMBIUS) GLYPTOCEPHALUS (Chamberlin). Lithobius glyptocephalus Chamberlin, Proc. Acad nat. sci. Phil., 1903, p. 35. Description.— Dorsum more or less chestnut with a median longi- tudinal pole stripe. Head more reddish, paler along margins and frontal suture. Antennae reddish brown, paler distad. Prosternum and prehensors brown, the latter somewhat paler ectad. Venter light brown, the caudal plates darker. Legs testaceous, the caudal pairs darker. Body about seven times longer than the width of the tenth plate. Widths of head and of first, third, eighth, and tenth plates to each other as 65 : 64 :59 : 65:65. The head is thus equal in width to the eighth and tenth plates as in G. rev. Head wider than long in about ratio 13:12. The short transverse sulcus near each marginal break is deep with its mesal end curving CHAMBERLIN: GENERA OF LITHOBIIDAE. 449 cephalad. A short sulcus extending mesocaudad from above eye on each side. A short but well-impressed transverse sulcus a little in front of median portion of caudal margin. A median longitudinal furrow on caudal portion with another one each side of it may be traced. Head a little elevated immediately cephalad of frontal suture, and on this elevation a median pit-like depression. Surface appearing smooth and shining, being only obscurely uneven and punctate. Margining continuous forward to the eyes but the thickening narrower cephalad of the lateral break. Head widest a little caudad of the eyes from where the sides converge to the caudal corners. Ocelli mostly near fifteen in three series: e.g., 1 + 5,5,4; 1+ 5, 5,3; 1+ 7,6,4; 1+ 5,5,5. Single ocellus large, obliquely oval. The patch elongate. The antennae moderate in length, reaching upon the seventh body segment. Articles normally twenty, but may number up to twenty- three on one side; slender and long; the ultimate long and pointed. Prosternum a little less than 1.6 times wider than long. Distance between chitinous spots about 1.9 times width at level of bottom of sinus; 2.5 times dental line. Teeth mostly 6+ 6 or 7+ 7; stout, acute, decreasing in size and extent of separation mesad. Spine a little ectad and caudad of ectal tooth; bristle-like distad. First dorsal plate nearly like head, but obscurely uneven, while the other plates are more markedly rugose and subgranulose, the caudal ones, as usual, most so, though none of them are strongly so. The major plates with a short submarginal sulcus on each side at about one third the length from the caudal margin and also a second similar one toward anterior end, there most conspicuous on more posterior plates. A pair of diverging longitudinal sulci on plate with a wider median furrow between them; on each side a more ectal furrow parallel with margin and usually uniting with the oblique sulcus near its middle. First ventral plate broadly mesally depressed, a sharply impressed median longitudinal sulcus crossing the broader depression. Other plates showing three distinct longitudinal furrows of which the median one on some of the posterior plates may become shortened and even pit-like. Hairs sparse, most abundant on the Caudal plates. Coxal pores rather large, subcircular or transversely elliptic: in number, e.g., 6, 7, 7, 6-6, 7, 8, 8. Gonopods of male well exposed; broad, distally subtruncate; bearing six or seven bristles. Gonopods of female with claw tripartite, the lateral lobes relatively much reduced. Basal spines 2 + 2; stout, clavately widening above base. 450 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Anal legs in the male moderately long; slender. Neither the anal nor the similarly slender penult legs bearing any special lobes or furrows. Spines of first legs, §°0°> 3-9; of the second to the tenth, OS of the eleventh, (03:33; of the twelfth, ress; of the thirteenth, o133-3; of the penult, }$ yaa; claws two, of the anal, Oy ~ goes at claw one. Last three pairs of coxae laterally armed. Length 18-22 mm. A male 18 mm. long has anal legs (exclusive of» coxae) 9 mm. long; antennae 9 mm. long; and tenth plate 2.6 mm. wide. Type Locatiry.— New Mexico: Beulah! Known Loca.ities.— New Mexico: Beulah! Chicario Canyon near Raton! Las Vegas! (T. D. A. Cockerell). 5 , LirHosius Leach (emend.). Trans. Linn. soc. London, 1815, 11, p. 381. Lithobius (ex part) Gervais, Hist. nat. ins. Aptéres, 1847, 4, p. 229. Newport, Cat. Myr., 1856, p. 2, 15. Meinert, Nat. tiddskr., 1868, 5, p. 258. Latzel, Myr. Osterreich-Ungar. monarch., 1880, 1, p. 31. Verhoeff Bronn’s Thierreich, 1907, 5, p. 239. Antennae varying in length in ours from short to moderately long articles always more than twenty, and in American species under fifty, being mostly from twenty-five to forty-five. Seriate ocelli present, mostly numerous. Single ocellus clearly differentiated. Prosternal teeth 5+ 5 to 10+ 10, or rarely but 4+ 4. Spine ectal in position, slender. No diastema in dental line of each side. The ninth, eleventh, and thirteenth dorsal plates with posterior angles produced. Coxal pores uniseriate, circular or, as mostly, transverse; present on last four pairs of coxae. Claw of female gonopods tripartite. Basal spines 2 + 2 or 2 + 3. Gonopods of male mostly uniarticulate, but they may be biarticu- late. None of coxae armed ventrally; some of posterior pairs armed dorsally and usually, but not always (e.g., L. forficatus), armed laterally. Third joint of all legs armed with three dorsal spines. Fifth joint of legs from first or second to eleventh or in most to thirteenth and sometimes to fourteenth with two dorsal spines. Dorsal spines of anal legs 1, 0, 3, 1, 0 (or very rarely varying to 1, 0, 3, 1, 1 or 1,0, 3, 2, 0 CHAMBERLIN: GENERA OF LITHOBIIDAE. 451 Fic. 3. — Distribution of Lithobius Leach. in L. forficatus, (according to Latzel); ventral 1, 3, 3, 1 or 1, 3, Bh eae claws one or two. Dorsal spines of penult legs 1,0, 3, 1, 1 or 1,0, 3, 2, 2; ventral mostly 0, 1, 3, 3, 2, less commonly 0, 1, 3, 3, 1 or 0, 1, 3, 3, 3. Dorsal spines of twelfth and thirteenth legs 1, 0, 3, 2, 2 to 1,0, 3, 1, 1 or coxal spine of twelfth missing. 452 ' BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Posterior legs of male essentially unmodified, not lobate or of conspicuously different form. Length of our species 11.5-30 mm., mostly above 15 mm. GEnoTYPE.— L. forficatus (Linné). In common with the other genera discussed in this paper and, in fact, with all members of the Lithobiidae as restricted by the author, the mouthparts of Lithobius have the following characteristics: — Labrum tridentate, a large median one in the sinus and a smaller one on each side. Laterally the labrum is clothed along margin and over border on the dorsal side with numerous bristles which are in large part ramose or laciniate, but in part are simple. The first maxillae have the outer branch biarticulate; distal article of this branch densely clothed with simply ramose hairs. Inner branch undivided; set off by a suture from the coxal division; densely clothed over the distal portion with simply ramose hairs similar to those of the outer branch. Second maxillae with claw of palpus normally five partite. Distal article complanate dorsomesally and clothed over this area with many penicillate and ramose setae similar to those of the first maxillae. Palpus triarticulate with the usual traces indicating two additional proximal articles. Coxosternum commonly with the median portion curved caudad and also bent dorsad; less strongly chitinized than the lateral portions which it widely separates. The distribution of the genus in North America is shown in Figure 3. It is rare in the Pacific Coast region and in the Southwest, where it is represented by but a single known species, L. chumasanus Chamberlin. Key to Species of Lithobius. a. None of the posterior coxae laterally armed. . L. forficatus (Linné). aa. Coxae of the last three pairs of legs laterally armed. b. Claw of anal legs single. c. Penult legs with the claw single; dorsal spines of thirteenth leeesrale Or eoyl Nel 4 erent ier a ante L. atkinsoni Bollman. cc. Penult legs with three claws; dorsal spines of thirteenth lees Oooo reo aa te Re ee L. celer Bollman. bb. Anal leg with two claws. c. Ventral spines of anal legs 0, 1, 3, 3, 1. L. chumasanus Chamberlin. CHAMBERLIN: GENERA OF LITHOBIIDAE. 453 LITHOBIUS FORFICATUS (Linné). Plate 2, fig. 1-3. Scolopendra forficata Linné, Syst. nat., ed. 10, 1758, 1, p. 638. DeGeer, Mém. ins., 1778, 7, p. 557. Fabricius, Spec. ins., 1781, 1, p. 532. Latreille, Gen. crust. ins., 1806, 1, p. 78. Sill, Verh. mitth. Siebenb. ver. nat., 1861, 12) pele Lithobius forficatus Leach, Edinb. encye., 1815, 7, p. 408. Zool. misc., 1817, 3, p. 39. C. Koch, Deutschl. Crust. Myr. Arach., 1844, hft. 40, p. 20. New- port, Trans. Linn. soc. London, 1845, 19, p. 367. Gervais, Hist. nat. ins. Aptéres, 1847, 4, p. 230. Newport, Cat. Myr., 1856, p. 18, 19. L. Koch, Myr. gatt. Lithobius, 1862, p. 39. C. Koch, Die Myr., 1863, 1, p. 113, fig. 104. Palmberg, Sverig. Myr. ord. Chilop., 1866, p. 15. Eisen and Stuxberg, Ofvers. Kongl. vet.-akad. Foérh., 1868, 25,no. 5,p.376. Meinert, Nat. tidsskr., 1868, 5, p. 259. Stuxberg, Ofvers. Kongl. vet.-akad. Forh., 1871, 28, no. 4, p. 496. Meinert, Nat. tidsskr., 1872, 8, p. 315. Fickert, Myr. arach. kamme Riesengeb., 1875, p. 7. Stuxberg, Ofvers. Konegl. vet.-akad. Férh., 1875, 32, no. 3, p. 10,27. Rosicky, Zool. landesd. Bohemen, 1876, p. 16. Latzel, Jahrb. nat. landesmus. Karnten, 1876, 12, p. 97. Fedrizzi, Atti. Soc. Ven.-Trent., 1877, 5, p. 205. Stuxberg, Proc. Cal. Acad. sci., 1877, 7, p. 1386. Haase, Schlesiens Chil., 1880, 1, p. 19. Latzel, Myr. Osterreich-Ungar. monarch., 1880, 1, p. 57. Sograf, Bull. Friends, nat. hist., 1880, 32, pt. 2, p. 1-33, pl. 1-3. Toémdésvary, Zool. Anz., 1880, 3, p.617. Fanzago, Boll. Soc. ent. Ital., 1882, 14, p. 48. Latzel, Boll. Soc. ent. Ital., 1882, 14, p. 366. Meinert, Proc. Amer. philos. soc., 1886, 23, p. 176. Bollman, Proc. U.S. N. M., 1887, 10, p. 260; 1888, 11, p. 409. Evert’s, Tijdschr. ent., 1888, 32, p. 41. Daday, Term. fuz., 1889, 12, p. 86, 107. Dalla Torre, Zool. jahrb. Syst., 1889, 4, p. 80. Pocock, Ann. mag. nat. hist., 1889, ser. 29, p. 62. Ellingsen, Forh. Vid.- selensk. Christiania, 1891, no. 10, p. 5. Bollman, Bull. 46, U.S. N. M., 1893, p. 200. Daday, Term. fiizetek., 1893, 16, p. 112. Kenyon, Can. ent., 1893, 25, p. 162. Pocock, Inst. nat., 1893, 2, p. 310. Brolemann, Boll. Soc. ent. Ital., 1895, 27, p.81. Attems, Sitzb. Kongl. akad. wiss. Wien, 1895, 104, p. 19. Simacovies, Verh. Siebenb. ver., 1896, 46, p. 100. Chamberlin, Proc. U. 8. N. M., 1901, 24, p. 24. Rothenbiihler, Rev. Suisse zool., 1901, 9, p. 358. Verhoeff, Boll. Soc sci. Bucarest-Roumanie, 1901, p. 168, 172. Tonniges, Zeits. wiss. zool., 1902, 71, p. 328. Williams, Amer. nat., 1903, 37, p. 299. Verhoeff, Zool. jahrb. Suppl., 1905, 8, p. 216. Evans, Proc. Roy. phys. soc. Edinb., 1906, 16, p. 408. Chamberlin, Ann. Ent. soc. Amer., 1909, 2, p. 190; 1911, 4, p.42. Can. ent., 1911, 43, p. 69, 101. Gunthorp, Kans. univ. sci. bull., 1913, 17, p. 165. Chamberlin, Bull. M. C. Z., 1914, 58, p. 220. Can. ent., 1920, 52, p. 166. Lithobius vulgaris Leach, Trans. Linn. soc. London, 1815, 11, p. 382. Johnston, Mag. nat. hist., 1835, 8, p. 491. 454 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Lithobius laevilabrum Leach, Trans. Linn. soc. London, 1815, 11, p. 382. Lithobius americanus Newport, Trans. Linn. soc. London, 1845, 19, p. 365, pl. 33, f. 29. Gervais, Hist. nat. ins. Aptéres, 1847, 4, p. 236. Newport, Cat. Myr., 1856, p. 17. Wood, Journ. Acad. nat. sci. Phil., 1863, new ser., 5, p. 14; Trans. Amer. phil. soc., 1865, 18, p. 148. Lithobius leachit Newport, Trans. Linn. soc. London, 1845, 19, p. 368; Cat. Myr., 1856, p. 19. Lithobius forcipatus Gervais, Hist. nat. ins. Aptéres, 1847, 4, p. 229. Lithobius muscorum L. Koch, Myr. gatt. Lithobius, 1862, p. 43. Lithobius forficatus villosus L. Koch, Myr. gatt. Lithobius, 1862, p. 41. Lithobius trilineatus L. Koch, Myr. gatt. Lithobius, 1862, p. 37. Lithobius hortensis L. Koch, Myr. gatt. Lithobius, 1862, p. 45. Palmberg, Sverig.-Myr., ord. Chilop., 1866, p. 17. Lithobius coriaceus L. Koch, Myr. gatt. Lithobius, 1862, p. 51. Meinert, Nat. tidsskr., 1869, ser. 3, 5, p. 260. Porat, Ofvers. Kongl. vet.-akad. Forh., 1869, p. 637. Lithobius multidentatus Wood (non Newport), Journ. acad. nat. sci. Phil., 18638, n.s., 5, p. 13. Lithobius curtirostris Eisen and Stuxberg, Ofvers. Kongl. vet.-akad. Férh., 1868, 25, p. 376. Description.— In general color varying from brownish yellow to chestnut; dorsal plates showing often. A median dorsal paler line, as also sometimes an oblique line on each side. Head brown to chestnut like the dorsum, with the frontal region sometimes paler. Antennae brownish yellow to chestnut, the tip usually rufous. Prosternum somewhat paler than the head but darker than the venter; prehensors mostly rufous or pale ferruginous distally. Venter brownish yellow to brown; frequently dusky, with the most caudal plates reddish to subferruginous. Body widest at tenth segment; very gradually parrowing cephalad and more decidedly so caudad as usual. Widths of first, third, eighth, and tenth dorsal plates to each other as 54 : 52 :57 : 59. Head a little wider than long (ca. 24:23); the portion caudad of eves subquadrate, the sides a little converging caudad of the marginal breaks. Caudal margin slightly incurved mesally. Immediately in front of median portion of caudal marginal thickening a curved sulcus, the free ends of which appear as diverging shallow furrows; this impression sometimes very vague, at others distinct. Surface somewhat uneven; punctate, the puncta being usually more distinct and dense in the frontal region. Ocelli twenty-two to forty, but mostly twenty-six to thirty-four; in five to eight series, but mostly in six. Single ocellus large, sub- CHAMBERLIN: GENERA OF LITHOBIIDAE. 455 ov Fic. 4. — Distribution of Lithobius forficatus (Linné). circular to oval in outline. Seriate ocelli closely arranged but those of the most dorsal one or two rows often with ocelli somewhat sepa- rated, especially so the more caudal ones. Examples of arrange- ment are as follows: 1 -+ 6,6,6,6,5,3; 1+ 4, 5, 5,5,4,2;. 1-+-5, 456 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 5 (aloo ae L +5, 05 O.0, 4,05 Lt OF OMG, O45 Meat ional, Oe 1 + 3, 6, 6, 5, 5, 4, 3. Antennae short, normally reacting at most to the anterior portion of the seventh segment, consisting of from thirty-six to forty-nine articles, but the number mostly from thirty-nine to forty-five of which those between the second and ultimate are short and very short and closely compacted. Hairs dense, short, straight. Prosternum wide, being mostly nearly 1.7 times wider than long. Anterior portion narrowed considerably cephalad; anterior margin a little convexly curving from each end to the median sinus. Teeth a little more widely separated ectally than toward median line; stout, moderately subacute: 5-+ 5; 5+ 6; 6+.6; 6+ %;.7 + 7. Spine in usual position at ectal end, a little caudad of most ectal tooth; slender (Plate 2, fig. 1). Dorsal plates well arched; surface but little uneven, appearing mostly smooth and shining; the short submarginal transverse im- pressions or sulci near beginning of caudal third of length commonly weak or obscure. Posterior angles of ninth, eleventh, and thirteenth dorsal plates produced, the processes stout and subacute. Ventral plates with a longitudinal median sulcus, which is more distinct on the caudal ones. A transverse sulcus on caudal portion of plate which is commonly obscure on anterior plates. In addition there is a weakly impressed line or furrow running obliquely ecto- cephalad on each side from near the junction of the two previously mentioned sulci, these oblique lines being most distinct on the last plate where they unite at the median furrow, with the caudal portion of which they compose a Y-shaped impression. Caudal plates and genital segment more densely pilose than the others. Coxal pores large, strongly transversely elongate, the most proximal ones on each coxa smaller and often subcircular; the most distal one mostly distinctly broader distoproximally, broadly elliptic to circular. Sometimes all pores are elliptic or almost circular as is normal in immature specimens. Examples of arrangement: 6, 6, 6,5; 7,8, 8, 6; S, OF0,,55 S10) LO S10, 107 10,9 OTS Sl O) Oe 9 HOR Onoe ld, Loris ae 125 105210: Gonopods of male biarticulate; first article broad, bearing 4-6 bristles; second article small and wart-like or broadly subconie, usually with a single long bristle. Claw of female gonopods short and rather wide; the three lobes mostly blunt, the median considerably exceeding the lateral ones which are subequal, but these always well developed (Plate 2, fig. 2). CHAMBERLIN: GENERA OF LITHOBIIDAE. 457 Basal spines 2 + 2 or 2+ 3, stout, in outline subconical (Plate 2, fig. 3). Posterior legs rather short, slender, being only slightly thickened. Femur of anal and penult legs with two parallel and rather weak, longi- tudinal furrows along the dorsal surface; the prefemur similarly but more indistinctly furrowed. Wholly without special lobes or modi- fications in the male. Spines of first three pairs of legs, §-}:3:3:5; of fourth to eighth pairs the same or ¢'f'3°3-2; of the ninth to eleventh, }133'3; of the twelfth, 1, 0,3, 1,2 . 1, 0, 3,1, 1 10,3,1,1 0-03-32; Of the thirteenth, ¢°7°3'3'3;. of the penult, 9'7°33'5, claws three, 1, 0, 3.1.0 . the anterior accessory pone minute; of the anal, 0°1:3,3,2, Claw single. . Length 18-30 mm.; width of tenth plate 2.25 to 3.5 mm. A o&@ 22.5 mm. long has the ae plate 2.8 mm. wide; antennae 10 mm. long and anal legs (exclusive of coxae) 9.6 mm. long. Pseudomaturus.— Coloration like that of paler adults. From light to dark brown; head darker brown to light mahogany. Antennae hike head, except at tips which are pale and usually rufous. Venter and legs paler brown; the caudal pairs of legs mostly darker than the others, often rufous distad. Antennae reaching from the fifth to seventh segment: Articles in specimens examined varying from thirty-six to forty-four. Ocelli mostly in five series and twenty to peny enn in number: PALS 66,5330 dG nGy a4 8: WPS, 4, 4.3-, Wi, 5, Onan: Prosternum as in maturus. Teeth mostly 5 + 5 Coxal pores 5, 6, 6, 5; 6, 6, 7, 6. Gonopods of male biarticulate; distal article in younger specimens glabrous, in older ones with one bristle and the first article mostly with two. Claw of female gonopods nearly as in adult, but a little thinner and somewhat shorter and paler with the teeth or lobes rather more acute. Spines about as in maturus or the mesal one of each pair in smaller specimens may be relatively shorter. Spining of legs as in maturus excepting that the dorsal spines of first legs are prevailingly 0, 0, 3, 2, 1 instead of 0, 0, 3, 2, 2 and the ventral spines of the ninth legs are more commonly 0, 0, 2, 3, 2 instead of 0,0, 3, 3,2, a formula in the adults also commonly holding as well for a number of the more anterior legs. Length 14-18 mm. A o 16.5 mm. long has antennae and anal legs both very near 6 mm. in length. (Watersmeet, Mich.). Praematurus.— Pale brownish yellow to light brown, the caudal 458 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. segments, especially in pale specimens, darker. Head and antennae usually a darker brown with the frontal region frequently paler and the antennae lighter at tips. Venter and most of the legs paler than the dorsum, the caudal plates and legs darker, the latter with tarsi often rufous. Prosternum a darker brown than the venter; the pre- hensors rufous distally. Antennae reaching fifth body segment. Articles thirty-four to thirty-eight. Ocelli twelve to nineteen in three or four series: e¢.g., 1 + 5, 6, 4, 3. Prosternal teeth 4+ 40r5 + 5 Coxal pores from circular to broadly elliptic: in number mostly 5, 5,5, 4. Porigerous area only slightly depressed. Gonopods of male biarticulate, the distal one small and glabrous, the proximal one with usually but one bristle. Claw of female gonopods short, moderately curved, thin and pale; the three lobes distinct, acute. Spines 2 + 2; the inner one of each pair much smaller than the outer, being only about one-half as long and less than half as broad. Bristles more sparse than in the pseudo- maturus. D, OF 2, 1, 0, 0, 0,0,2,2, 1 Spines of first legs, 979-934 1 to 0° >; of the second, p'p'3'3:4 to adult number; of the iehed and fot rth, ooe5 to adult number; of the fifth and sixth, (23:35 to p'p:33:3; of the seventh to eleventh, eer or ventral spines of tenth and eleventh may be 0, 0, 3, 3, 2, and in one specimen ventral spines of eleventh on one side were 0, 0, 2, 2, 2; of the remaining four pairs as in the older stages. Length 11 to 13.5 mm. A @ 13.5 mm. long has antennae 5 and anal legs 5.5 mm. long. Immaturus.— Brownish yellow to light brown, the caudal segments, especially in paler specimens, darker. Head usually a little darker than dorsum with frontal region paler, often yellowish. Antennae usually of same color as frontal region of head. Venter and legs pale with caudal plates darker; tarsi of last pairs of legs often pale rufous, the whole body in life may be suffused more or less with a violaceous color. Antennae composed of twenty-nine to thirty-six articles, of which the first three are long and the others mostly short, but commonly showing longer ones at intervals. Ocelli ten to eighteen in three or four series: e¢.g., 1 + 4, 4,3; a common arrangement: 1 + 5, 5,4, 3. Single aaa leipes elliptic. Prosternal teeth nearly always 4 + 4; rarely 5 + 5 Coxal pores all circular or in larger specimens transversely some- what elliptic: mostly 4, 4, 4, 3, but varying from 3, 3, 3, 3 to 4, 4, 4, 4. 0, 3, 2, 0, 2, 3, 0, 0, CHAMBERLIN: GENERA OF LITHOBIIDAE. 459 Male gonopod appearing as a small rounded tubercle, often con- cealed, which is undivided and may bear a single bristle; or in some a trace of the second article. Claw of female gonopods small, pale, appearing at most as a small acute spine without trace of lateral lobes or in smallest specimens claw wholly absent and then often only two articles developed in gonopod. Basal spies | + 1 or occasionally with the merest trace of a second one on one side; pale, often appearing as mere points. The second and third articles may be entirely glabrous or the second may have two or three and the third one or two bristles, the first having from one to four. 0,0,1,1,1 0,0, 21,1 . 0,.0,.2, 2,1 Spines of first legs, 9-0-4214 to 0,0,%3a),13 of the second, 0-o-334 or ey ‘ ‘ s Ones 2 1 with ventral spines 0, 0, 2,3, 2; of the third, 9-344 or ventral spines ; PAT 2D 0101.25 9 2d, 7 . to 0, 0, 2,3, 2; of the fourth, $:9:7°5 1 to o-o-2"35; Of the fifth and sixth, 10, 0,2,2.2 0, 0. 3, 2, 2 MERON O S24 orale : : 0-333 tO o-9-3:3-0; Of the seventh, 3, 30), 2 of the eighth and ninth, — in ). _ i 25 1, 0, & o 3, 2:2. 10-232; of the tenth and elev enth, tes 2Or ventral spines 0, 0, 3, 3, 2 10), 0, 3, 1, 2 Pal ORO oaks L 1,03, 1, 813, 2 of the twelfth, “G05, 52 3 of the thirteenth, Fee) Cag gs OT 453,53, of the penult, o, : 3 10 to & ; z 5; of the anal, pet As b 3 = tod 00; - : 0 Length 9-11 mm. A o& 10.5 mm. long has antennae 5 mm. and anal legs 3 mm. long. (Watersmeet and Marinette, Mich.). Agenitalis IT.— Dorsum pale; dilute brownish to pale testaceous. Head light brown, the frontal region lighter. Venter and legs a little paler than dorsum, the caudal segments usually darker. In life the entire body is suffused with violaceous. Articles of antennae twenty-five to thirty-three; those distad of the third varying considerably in size in different individuals, often sub- moniliform. Ocelli nine to twelve in two to four series: e.g., 1 + 2,3, 2;.1-+ 4, Syl; f= 453,2; 1+ 4,4,2; 1+ 4,3;2,1; 1+ 5,4. Ocelli pale. Single ocellus much largest, elliptic. Prosternal teeth by far most frequently 4 + 4; occasionally 4 +5 or 5+ 5. Coxal pores’ 2; 3,3, 2-2; 2) 2;'2; cireular. Gonopod of male very small, glabrous; commonly wholly concealed. Gonopods of female appearing as subeylindric processes which are undivided and entirely glabrous. Spines of first legs, a ae t : Soto tO Woe et; of the fo f ita 24; of the fifth, ¢¢> of the sixth the same to }}-?3:7; of the seventh, sy to jt ae of the eighth and ninth, a, 3. 75 to 10.2 : 2 ate the tenth and ae a ) 4; of the second and third, be BBS Le re is 0.0, 3, 2, 2 0, 0, 3, 2, 2 : : 0, 0, 3, 1, 2 . . 0-02-99 tO o0-2°3-2; Of the twelfth, to o-03:3-2/; of the thir- 1Tn a regenerated leg of one specimen the formula is o: a, : . i: 460 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ONON2@) deel ee OOS alee 0, 0, 3, 1,1 NOMS Adsel teenth, 0, 0, 2, 2, 2a) to 9 13.3.2) of the penult, hikerenn to 0, 1, 3, 3,2) of the if 0, 0, 2), 1,0 + 1,03. 1.0 ana Leki 0,1,3, 2, 1° Length 7-9 mm. A specimen 7.5 mm. long has antennae 3.25 mm. and anal legs about 2.8 mm. long. (Watersmeet, Mich., July, 1910). Agenitalis I.— Very pale in color. Violaceous in Es Articles of antennae twenty-one to twenty-seven; those distad of the third short, moniliform. Ocelli five to seven in two or eceionally in hte series: €.9., 1+ 3,2, the commonest formula; 1 + 2,2; 1+ 2,2,1; 1+ 3,3. Pale. The single ocellus and the caudal one of top series ee than the others. Prosternal teeth mostly 4 + 4; more rarely 3 + 4or3 + 3. Coxal pores 1, 1, 1, 1, each toward distal end of its coxa, to 1, 1, 1, 2 amg 2s Alt be, ihe Moderates in size, pale edged. Anal glands still distinct and rather large. Caudal angles of ninth, eleventh, and thirteenth dorsal plates pro- duced, but the processes short and somewhat rounded. Gonopods not evident in either sex. ane ‘oh On0,0eit) 0,0 ON ab ra snes of first legs, o:o:4°471 OT 0:0; 0 ag, of the second, 9” 0,0, 0), 1,1 to Corii; of the third and fourth, oi71; of the fifth and sixth, 00.121, of . 0, 0. 24),2,1 0, 0, 2.2.2 o01-3-7; of the seventh and eighth, 9°94. 2, 12) 5 of the ninth, 9, 0, 1, BOE 0, 0, 3! . 0, 6,2, 1,2 eset of the tenth, 9 6.1 3 ie} ae eleventh, 9.6.4. 3a), 13 of the twelfth, 9 OnOedzas 5 ) 5 of the thirteenth, po 41; of the penult the same or Guamaity) OL the al 0, 0, 0, 0, 0 anal, 0, 0,1,1,1- Length 6-7 mm. (East Peoria, Ill., July, 1910). Pullus IV (Larva IV).— Very pale testaceous to dilute ferruginous, with head and antennae a little darker. Antennae composed of twenty-one to twenty-two short articles, moniliform. Ocelli three to five in two series: e¢.g.,1 + 1,1; 1+ 2,1; 1+ 2, 2. Prosternal teeth mostly 4 + 4 with the most mesal one on each side very small; occasionally 4 + 5, or 3 + 3. A single pore on each coxa of twelfth legs.! Anal glands large, distinct. we pf QeOnONt 1.4 ve QUONO Sant ead Spines of first legs, o-p-oaa; of the second to the fourth, 9°¢:0:4.4; of aye Os Onde ie . Ce ae er : 0; 0, 2(0), 2, 2 the Heth, 2. of the ninth and tenth, eae of the eleventh, 0, 2,1, 2 0, 2,1,1 1, 0,2,1,1 ey ree of the twelfth, (iyo; of the thirteenth, 5 eess; of the Oz el 1, 6, 2, 1,0 penult, 4 0, Bee EEL bay 3, 1) ot the anal, (hal 3.2 1- Length of specimen described 7 mm.; antennae 3.5 mm. Tyre Locauiry.— North Carolina: Jackson Co., Balsam! Known Locauities.— Georgia: Macon (L. M. Underwood), Atlanta! Lula! Tallulah Falls! Bremen! North Carolina: Balsam (G. F. Atkinson); Saluda! Hot Springs! South Carolina: Taylors! Greenville! Seneca! NEOLITHOBIUS STUXBERG (emend.). Ofvers. Kong. vet.-akad. Férh., 1875, 32, no. 3, p. 8, 10,26. Proc. Cal. acad. sci., 1877, 7, p. 135. Latzel (in part), Myr. Osterreich-Ungar. monarch., 1880, 1, p. 35. Bollman (in part), Proc. U. 8S. N. M., 1887, 10, p. 262, 626. Bull. 46, U.S. N. M., 1893, p. 29, 43, 129, 164. Eulithobius Verhoeff (in part), Myr. Bronn’s Thierreich., 1907, 5, p. 240. Antennae moderate or long; articles thirty to fifty. Ocelli numerous, seriate. Single ocellus clearly differentiated. Prosternal teeth 5 +°5 to 10 + 10. Spine ectal in position, slender and mostly bristle-like distally. No diastema in dental line on each side. Posterior angles of seventh, ninth, eleventh, and thirteenth dorsal plates produced. Coxal pores uniseriate, mostly transverse, but sometimes circular. Claw of female gonopods tripartite. Basal spines 2 + 2 or, rarely, 3+ 3. Gonopods of male uniarticulate or mostly so. None of the coxae of legs armed ventrally. In all known species some of the posterior coxae are armed both dorsally and laterally. Third joint of all legs armed with three dorsal spines. Fifth joint of legs from first or second to eleventh or sometimes to thirteenth armed with two dorsal spines. Dorsal spines of anal legs mostly 1,0, 3, 1,0, more rarely 1, 0, 3, 2,0 or even 1, 0, 3, 2, 2 (sometimes in mordax); ventral 0, 1, 3, 3, 1 or 0, 1, 3, 3, 2; claw one or two. Dorsal spines of penult legs 1, 0, 3, 1, 1 to 1,0, 3, 2,2; ventral 0, 1, 3, 3, 2 or 0, 1,3, 3,3; claws one, two, or three. Dorsal spines of twelfth and thinieenth legs 1, 0, 3, 2; 2 or, less often, 1,0; 3, 1; I. Anal legs of male with fourth joint enlarged, at least distally, being in most conspicuously widened, and especially bowed ventrad, or at least with dorsal surface depressed and incurved, rarely with dorsal surface not depressed but elevated at distal end. Length 17-30 mm. CHAMBERLIN: GENERA OF LITHOBIIDAE. 471 Fic. 5.— Distribution of Neolithobius Stuxberg. GENoTYPE.— Lithobius vorax Meinert.! Neolithobius includes a compact and homogeneous group of species having a seemingly unbroken distribution (Figure 5). 1 Stuxberg specified no type. The three species originally included by that author are vorar Meinert, mordar L. Koch, and éransmarinus (L. Koch). The first mentioned is here selected as the type. 472 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. Key to Species of Neolithobius. a. Anal legs with the claw single. b. Penult legs with but one claw. c. Tibia of all or of nearly all legs excepting the anal with three ventral spines, dorsal spines of penult legs (usually also of twelfth and thirteenth) 1, 0, 3, 1, 1. N. transmarinus (LL. Koch). ce. Tibia of none of legs normally with more than two ventral spines d. Dorsal spines of penult legs 1, 0, 3, 2, 2 or 1, 0, 3(2), 2(3), 1 e. Ventral spines of anal legs 0,1,3,3,1, the dorsal normally, 1,0,2,0; dorsal spines of penult legs 0 aco hep ee Re EAs ome N. mordax (L. Koch). ee. Ventral spines of anal legs 0, 1, 3,3, 2, the dorsal nor- mally 1,0,3,1,0; dorsal spines of penult legs DFO, Sn 2 Oya atc ae ee N. helius Chamberlin. dd. Dorsal spines of penult legs 1, 0,3, 1,1; ventral spines of anal and penult legs 0, 1, 3, 3, 2. e. Fourth jomt of anal leg of male strongly produced dorsad at distal end (Plate 3, fig. 5). N. xenopus Bollman. ee. Fourth joint of anal leg of male moderately bowed ventrad but not conspicuously produced at distal end: above yA ae aon ee N. suprenans, sp. nov. bb. Penult legs with two or three claws. c. Dorsal spines of penult legs normally 1, 0, 3, 2, 2. d. All legs from the thirteenth forward at least as far as the sixth pair with three ventral spines at distal end of the tibia! 51 ine tern Meneame N. latzelii (Meinert). dd. Either none of the legs with tibia bearing three ventral spines or at most the tibiae of thirteenth and of one or two preceding pairs thus armed. . NV. tyrannus Bollman. ce. Dorsal spines of penult legs normally 1, 0,3, 1, 1 or at most WO on Zeke d. Tibiae of all legs from second or first to thirteenth armed with three ventral spines.......... N. vorax (Meinert). dd. Tibiae of all legs from first to thirteenth with but two ventral spines, none with three. e. Dorsal hairs of anal segment of o strongly curled or UNCIM ATE.) Mae ya: eR N. devorans (Chamberlin). CHAMBERLIN: GENERA OF LITHOBIIDAE. 473 ce. Dorsal hairs of anal segment of co straight, not at all curled or uncinate........ N. voracior (Chamberlin). aa. Anal leg with two claws. Dorsal spines of penult legs 1, 0,3, 1,1; tibiae of twelfth and thirteenth and sometimes of one or two preceding pairs of legs with three ventral spines............ N. underwoodi (Bollman). NEOLITHOBIUS MORDAX (L. Koch). Plate 3, fig. 1, 2 Lithobius mordax L. Koch, Myr.-gatt. Lithobius, 1862, p. 34. “Wood, Trans. Amer. phil. soc., 1865, 18, p. 149. Meinert, Nat. tidsskr., 1872, 8, p. 294. Lithobius (Neolithobius) mordax Stuxberg, Ofvers. Kongl. vet.-akad. Forh., 1875, 32, no. 3, p. 10, 26. Proc. Cal. acad. sci., 1877, 7, p. 185. Bollman (in part), Proc. U.S. N. M., 1887, 10, p. 262, 627. Ent. Amer., 1888, 4, p. 8. Bull. 46, U.S. N. M., 1893, p. 29, 44, 46, 146, 185. Kenyon, Can. ent., 1893, 25, p. 162. Chamberlin, Can. ent., 1911, 43, p. 101. Psyche, 1918, 25, p. 24. ? Lithobius howet Bollman, Amer. nat., 1887, 21, p.81. Proc. U.S. N. M., 1887, 10, p. 255, 259. Bull. 46, U.S. N. M., 1893, p. 19, 22, 26. Lithobius spinipes Bollman (in part), Bull. 46, U. 8. N. M., 1893, p. 146. Lithobius mordax louisianae Brolemann, Ann. Ent. soc. France, 1896, 65, p. 48 (07). Lithobius transmarinus perarmatus Brolemann, Ann. Ent. soc. France, 1896, 65, p. 48 (2). Lithobius transmarinus Chamberlin (in part), Ann. ent. soc. Amer., 1911, 4, p. 45. Derscription.— Dorsum from light to dark brown and chestnut; nearly uniform, or, in the paler specimens, with the first plate and several moot caudal ones darker than rest of dorsum; plates fre- quently paler along lateral and caudal margins and in a median longi- tudinal line, with sometimes one on each side of it. Head a little darker, and more reddish than the dorsum. Antennae chestnut, paler, often yellow, distad. Prosternum and prehensors chestnut. Venter testaceous to dark brown; the posterior plates darker, reddish or chestnut. Legs testaceous or brown, the posterior pairs darker, mostly same shade of chestnut like corresponding plates of venter. Body moderate, commonly 8 or 8.25 times longer than width of the tenth plate. Nearly parallel-sided, but little narrowed cephalad from the eighth plate to the third, which is narrower than the first. First plate wider than head. Widths of head and of first, third, eighth, and tenth plates to each other as 94 : 96 : 88 : 94 : 95. Head wider than long in ratio near 11.75 : 10.75; widest immedi- 474 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. ately caudad of eyes. Caudal margin mesally a little, widely incurved. Marginal breaks abrupt and distinct. Sides caudad of breaks moderately converging caudad to the rounded corners, nearly straight; between breaks and eyes convex. Surface rather finely but distinctly and uniformly punctate; not rugose or roughened otherwise. The usual semicircular impression a little in front of caudal marginal thickening with one or two pit-like depressions on median line in front of suture and usually a similar impression a little each side of the median line just caudad of the suture. Ocelli twenty-seven to fifty, mostly in from six to eight series: e¢.g., 1 FE 583.7, 6; OVS 16.6; 6,593) 25 FE by ONS 8) Gib. lee 7,9, 10,9, 6,5. Single ocellus very much larger than any other; subvertically narrowly oval with the narrow end ventrad; often paler than the other ocelli. Seriate ocelli compactly arranged, the series often confused; an ocellus (usually the most caudal one of the fourth row) immediately in contact with single ocellus always considerably larger than any other of the seriate ocelli but smaller than the single one. Organ of Témésvary often in contact with most anterior ocellus of bottom row and in some line or ventrad of it. Antennae long, reaching the eighth body segment. Articles mostly thirty-five to thirty-nine.! Articles mostly moderately short, de- creasing distad. Hairs rather short, dense. Prosternum 1.8 times wider than long. Distance between chitinous spots 2+ times width at level of bottom of median sinus and up to as much as three times the dental line. Median sinus rather deep, its sides subparallel and its bottom rounded. Line of apices of teeth mesally distinctly reéntrant, each side being slightly convex. Outer teeth on each side more widely separated from each other than the more mesal ones which are often closely crowded: 6 -+ 6, perhaps most common; 6+ 7; 7+ 7; 7+8; S+8; 7+ 9. Spine in usual ectal position; slender and acute, stouter than the bristles. First dorsal plate smooth and shining like the head. Other plates finely roughened with tubercular elevations, the plates becoming more and more strongly tuberculate toward caudal end of body. Posterior angles of seventh, ninth, eleventh, and thirteenth plates strongly produced. Dorsal hairs of anal segment sparse, straight. Ventral plates each with a distinct transverse furrow which reaches on each side only part way to the lateral margin and which is a little caudad of the middle of the plate. The usual lateral and median longitudinal furrows distinct. 1 Reported with as few as thirty-two (Meinert) and even thirty (Bollman); but probably the specimens were immature. CHAMBERLIN: GENERA OF LITHOBIIDAE. 475 Coxal pores elliptic to strongly transverse, slit-like or keyhole- shaped muchas m L. forficatus: 6,6; 6, 5; 7,8, 75:0; 898,70; 8, 8, Sone bt sl 4 Oy Gonopods of male wide; caudal margin more or less. convex, bearing mostly five or six bristles. Claw of female gonopods long, but little curved; tripartite with the median lobe long and acute, but the lateral teeth small, far proximad, and often almost obliterated. Basal spines 2 + 2; subequal, stout, a little widening from base to beginning of acuminate distal portion, the edges of which are commonly minutely serrulate. In the female the anal and penult legs are moderately long and slender; the femur of each is distinctly longitudinally suleate along dorsal surface; the prefemur and femur less distinctly so; ectal surface of all articles longitudinally suleate but most distinctly so on prefemur, femur, and tibia; also mesal surface, especially of more distal articles longitudinally suleate; tibia and first tarsal joint of anal legs slightly bowed ventrad. In the male the femur of anal legs is thickened with the dorsal surface flattened and depressed or more or less bowed ventrad, longitudinally sulcate, more thickened at distal end; tibia slender and unmodified. In the male the same sulcations are present on the anal legs as in the female but those of the mesal surface are deeper and more distinct. Tibia of anal legs always con- siderably longer than the femur. Spines of first and second legs, oe 23) 5; of the third to eighth pairs, 0, “| 2 > . > o232; of eighth or with ventral spines of seventh and eighth or of eighth alone 0,0, 3, 3,2; of ninth, 00 0 oe 35 or with ventral spines 0, 0, 3, 3, 2; of the tenth and twelfth, ¢ 0:0, 03. Be of the thirteenth, y , a3; 1, 0, 3, 2, 2 i‘ : g : of the penult, 9°7°3°3'2 (also noted on one side of one specimen as 0° 13, aD the leg possibly regenerated) ; of the anal, #34, the dorsal spining occasionally varying to 1, 0, 3, 2, 2 and 1,0, 3, 1,0; claw one. Last three or four pairs of coxae laterally armed. Length 20-30 mm. A 2 29 mm. long has antennae and anal legs each 14 mm. long; and tenth plate 3.5 mm. wide. Pseudomaturus.— Dorsum testaceous to dark brown, marked with the usual longitudinal pale lines Head brown to dull chestnut, contrasting more with the dorsal plates than in adults; frontal region paler. Antennae brown or light chestnut proximally, elsewhere light. chestnut. Prosternum dull brown. Venter paler than dorsum, darker caudad, the legs like corresponding plates. Antennae as in maturus. Ocelli noted as 1 + 6, 6,5, 4, 1; 1 + 6,6, 6, 5,4; 1 + 3, 5, 6, 6, 4, 2; 476 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. 1+ 3, 6, 6, 5, 5, 3, 1; ete. Then in five to seven series, the larger number less common. Prosternal teeth 5 + 6 to 8 + 8; as in maturus. Coxal pores less strongly transverse than is usual in maturus, being more often elliptic to subcireular. Claw of female gonopods shorter and less deeply colored than in maturus with lateral teeth more distinct and the median one relatively shorter. Spines nearly as in maturus or somewhat narrower proximad of apical division. Spines of legs as in maturus. One specimen was noted with spines of both penult legs, (337 In one the ventral spines of tenth legs were 0, 0, 2, 3, 2. As many as the last seven pairs of coxae may be dorsally armed. Posterior legs of male as in maturus but femur mostly less enlarged. Length 16-19 mm. Praematurus — Testaceous or brown; showing the usual pale lines. Head a little darker; frontal region paler. Antennae brown, yellow distad. Prosternum and prehensors brown, a little darker than venter. Venter testaceous to brown, posterior plates a deeper brown. Poste- rior legs dilute ferruginous, often dark brown proximally. Articles of antennae thirty-two to thirty-four; mostly short, in distal portion longer single articles alternating with groups of shorter ones. Ocelli thirteen to twenty-three in three to five series: ¢.g., | + 5, 4, 3; 1+ 3, 4, 3, 2; 1+ 3, 6, 5, 5, 3. All distinct. Single ocellus largest but the first one of second series, which is contiguous with it, not much smaller. Prosternal teeth 5+ 5 to 7+ 7. Sides of median sinus con- siderably converging caudad. Coxal pores circular to elliptic: 5, 5, 5,4; 5, 5, 5, 5. Claw of female gonopods small, pale, narrowly acute with no indi- cations of lateral teeth or one indicated as a small acute pomt. Spines 2+ 2; slender, conical, sides converging from base to apex; inner spine much shorter and more slender than the outer. ONON Snes = 9 5 ‘ Spines of first legs, 0-022 ‘+ or ventral spines 0, 0, 2, 3, 2; of the 0, 0, 3, 2, 2 second and third, ? ee of the fourth and fifth the same OF 0,0, 2, 3,23 of the sixth to eighth as In maturus; of the ninth and tenth as in maturus or oe iH aS 39; of eleventh, twelfth, and thirteenth as in maturus; of penult, } ° a z i}, sometimes with mere traces of accessory claws; of the anal, } 0; or ae D Last five to seven pairs of coxae dorsally armed, last three or Hee pairs laterally. (Jackson, Alabama). CHAMBERLIN: GENERA OF LITHOBIIDAE. 477 Immaturus.— Dorsum light brown, nearly uniform, the light lines much as in older stages. Head darker, somewhat reddish brown, with frontal region paler. Antennae light brown proximally, the distal two thirds lighter. Prosternum dusky brown. Posterior legs light, yellowish distad. Articles of antennae thirty-one; beyond the first few short and very short ones are intermingled. Ocelli twelve to sixteen in three or four series: e¢.g., 1 + 3, 5, 5; 1 + 4, 5, 4, 2. Prosternal teeth 5 + 5. Coxal pores 4, 4, 4, 35 circular or a little elliptic. 2, 2,1, ° 0, 0, 3, 2, 1 Spines of first legs, (-¢-3:3:4; of the second and third, }'33'4; of the 0. 0 3.2.2 ONO S252 0, 3, 2, 2 fourth to ninth, 9-9-3373; of the tenth, 00233; Of the eleventh, C023: of the twelfth, 3 ee 1, 0,3,2.1 3; of the thirteenth, 9°7:3:3:3; of the penult, oss Claw dguasemied or with mere traces of accessory claws; of the anal 9°7°3°3'7. Last two pairs of coxae, at least, laterally armed. Length of specimen described 10 mm.; antennae and anal legs 5 mm. long. Type Locatiry.— Louisiana: near New Orleans. Known Locauitires.— Louisiana: New Orleans! Mississippi: Biloxi! Holly Springs! Fernwood! Canton! Brookhaven! Ala- bama: Jackson! Selma! Mobile! Tennessee: Nashville! Arkansas: Little Rock to Ultima Thule (Bollman) Iowa! Kansas! Nebraska! Minnesota: Fort Snelling! Winona! Lithobius howet Bollman was based upon a single male from Fort Snelling, Minn. The specimen in the U.S. N. Museum labeled type is a male of N. mordax Koch, differing so much from the original descrip- tion that it is very doubtful whether some transfer of specimens may not have been made. It differs in size, being 20 mm. long instead of 15 mm.; in having the posterior angles of the seventh, ninth, eleventh, and thirteenth dorsal plates produced instead of only the ninth, eleventh, and thirteenth; and in having the prosternal teeth twelve (six on each side) instead of six all together (or three on each side). Bollman said that his type was in bad condition, a statement true of the specimen under consideration. The antennae are broken off; but the original description states that twenty articles are present which, of course, could not apply to a specimen of mordax. Later Bollman dropped the name S. politus (McNeil) and applied the name L. howei to the same specimen that had previously borne the former name; but there are some differences between the original description and S. politus, so that if the specimen above mentioned be rejected as 478 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. the type, it is difficult to apply the name L. howei with certainty. Bollman himself, however, in his later writings, without any explana- tion of the change, applied it to Sontbius politus (McNeil). NEOLITHOBIUS TRANSMARINUS (L. Koch). Plate 3, fig. 3, 4. Inthobius transmarinus L. Koch, Myr. gatt. Lithobius, 1862, p. 33, 34. Lithobius (Neolithobius) transmarinus Stuxberg, Ofvers. Kongl. vet.-akad. Forh., 1875, 32, no. 3, p. 26, 32. Bollman (in part), Proc. U.S. N. M., 1887, 10, p. 626. Ent. Amer., 1888, 4, p.8. Bull. 46, U.S. N. M., 1898, p. 43, 80. Chamberlin (in part), Ann. Ent. soc. Amer., 1911, 4, p. 45. Psyche, 1918, 25, p. 24. Gunthorp (in part), Kans. univ. Sci. bull., 1913, 17, p. 166. Lithobius spinipes Bollman (in part), Bull. 46, U.S. N. M., 1893, p. 146. Description.— Dorsum from yellowish brown to deep brown; in darker specimens the major scuta often more deeply colored along caudal borders, blackish; a median pale line with one each side of it, as in mordax, more or less traceable; nearly uniform, the posterior plates commonly not darker. Head with a more reddish cast; frontal region paler. Antennae brown or dilute chestnut proximally, be- coming pale distad. Prosternum light chestnut, the prehensors commonly more reddish. Venter yellow or light brown, sometimes almost slate colored, the last few plates a very little darker. Legs in general like corresponding parts of venter; the tarsi in specimens in full color more or less orange; posterior pairs from yellowish or very dilute orange in paler specimens to deep orange or somewhat ferrugi- nous in the darker ones. Body in an average male 8.5 times longer than width of tenth plate. First plate widest widths of head and of first, third, eighth, and tenth plates to each other as 78 : 82 : 76 : 79 : 79. Head a little wider than long (ad 78 : 75). Distinctly and uniformly punctate, the puncta fine. Not rugose; shining. 5, the thirteenth being the same; of the penult, *52425'32”, or a gn A the anterior or ectal dorsal spine of the fourth joint in the latter case being mostly very small, claws two or three, in latter case the anterior accessory being very small; of anal, o133-2, the claw single. Pseudomaturus.— Testaceous with the first plate and the last few darker or not; major plates showing three longitudinal light lines of which the median is more obscure. Head light chestnut-brown. Antennae brown or light chestnut proximally, light brown to yellowish elsewhere. Prosternum dilute chestnut, the prehensors paler. Venter brown to pale brown, posterior plates darker, light chestnut to deep brown, not reddish. Legs fulvous, the posterior pairs usually not darker than the others. Antennae as in maturus; articles thirty-two or thirty-three short and very short ones intermingled. Ocelli twenty to twenty-eight in five to seven series. Prosternal teeth, in all specimens examined, 5 + 5. Coxal pores in most 5, 5, 5, 4 but also 6, 6, 6, 5; small, circular or in part somewhat elliptic. Gonopods of male small; bristles one or two Claw of female gonopods smaller than in maturus with divisions more slender and acute. Spines as in matur us. Spines of first legs, 9°: 3:771) 0073-22 OF brorar373; of the second, “0.28.1, to oo-233; of the third to fifth, ¢: degra; of the sixth to eighth, 9 0, aS or ventral spines 0, 0, 2, 3, 2 or even 0, 0, 2, 2, 2; of the remaining pairs uniformly as in the maturus. Length 14-18 mm. A female 15 mm. long has antennae 8.5 mm. long; and anal legs 6 mm. long. Widths of head and of first, third, eighth, tenth and twelfth plates to each other as 53 : 48 : 47.5 : 57: 57 : 52. Immaturus (early).— General color pale yellowish white, the head 492 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. scarcely darker than rest of dorsum and venter also nearly same as dorsum. Legs light, uniform. Antennae slightly darker than the legs, palest distally. Head wider than long, in ratio 32 : 30. Articles of antennae thirty-one; short and very short, the shortest ones often occurring in pairs among the somewhat longer ones in distal portion. Ocellitpale; V=Fi2, 5) 47271 Sa 2 by 35ale Prosternal teeth 4 + 4, sstoenit in size and spacing. (In a meas- ured specimen 1.66 times wider than long. Distance between chitinous spots 1.9 times width at sinus and 2.77-+ times the dental line). Coxal pores small, circular; 3, 3, 3, 2. Gonopods of female as yet consisting of but two articles but of these the distal one shows slight indications of subdivision and at its apex the claw appears as a pale, acute point. A single basal spine on each side, this being very small and appearing as a mere acute point. First article bearing one or two bristles, the distal one glabrous. Spines of first legs, 9-0-7" ti; of the second, §°o'4 é iii; of the third and fourth, tg-tei; of the fifth, Ot Lp the posterior dorsal spine of the tibia care very small; of the sixth the same or with ventral spines 0,0, 1, 2, 2; of the seventh and eighth, (:?: 73:13 of the ninth to eleventh, 3 6 322, of the twelfth, §:3:33-3; of the thirteenth, sas; of the penult, 91°33, as in adult; and of the anal, gaa Last pair of coxae laterally armed. Length of specimen described 9.5 mm.; length of anal legs 4.25 mm. ; of antennae 5.5 mm. (Widths of head and of first, eighth, tenth, and twelfth plates to each other as 32 : 29 :33 :33 : 29). Locatiry.— Alabama: Jackson! This species is very close to voracior. Usually both sexes of fully grown specimens of these species are to be distinguished quite readily by the form of the coxal pores, these in voracior being circular or broadly elliptic whereas in devorans they are larger and mostly strongly transverse. In some cases, however, it is difficult to separate the females; but the males are always distinguishable. NEOLITHOBIUS VORACIOR (Chamberlin). Lithobius voracior Chamberlin, Ann. Ent. soc. am., 1912, 5, p. 150. Descrietion.— Dorsum brown, the first plate and the several most caudal ones commonly darker and more reddish or chestnut; plates often showing three longitudinal pale lines. Head conspicu- CHAMBERLIN: GENERA OF LITHOBIIDAE. 493 ously darker than the dorsum, cherry-red or chestnut, or in others often very dark, nearly mahogany. Antennae chestnut, much lighter distad. Prosternum dark brown or dilute brownish chestnut, with the prehensors paler. Venter yellowish brown, the posterior plates much darker, burnt brown. Legs yellowish brown, the posterior pairs darker, being often somewhat chestnut with distal articles a little paler. Body more robust than in devorans, the length being mostly less than seven times as great as width of tenth dorsal plate (6.8). Con- siderably attenuated cephalad from eighth plate, with the third plate narrower than the first and the latter clearly narrower than the head. Average ratio between widths of head and first and tenth dorsal plates 68 : 65 : 74.3. Widths of head and of first, third, eighth, tenth, and twelfth plates in a o to each other as 68 : 65 : 63 : 75 : 75 : 68. Head wider than long. Widest immediately back of eyes. Sides rather strongly convex and converging caudad of the well-marked marginal breaks to the rounded posterior corners. Caudal margin nearly straight or but slightly incurved mesally. The usual subcircular impression on caudal portion. Surface subsparsely punctate, the puncta varying in size but mostly fine; obscurely uneven. Eyes composed mostly of from thirty to thirty-five ocelli, rarely as few as twenty-three, arranged in from five to seven longitudinal series; 2g: = 3602155") 49> 394,53"65'5, 4927 I= 5,6, 6, 6, 4,13: 1 + 3, 5, 6, 6, 6, 4, 3; 1+ 4, 6, 7, 6, 5, 4, 1. Single ocellus large subvertically oval. Ocelli of most dorsal row larger and paler as in related species. Antennae of moderate length, reaching mostly to the end of the seventh or beginning of eighth segment. Attenuated considerably but the distal portion not very fine. Articles beyond the first several rather short. Hairs moderately long. Articles thirty-two to thirty- six In number. Prosternum 1.6 times wider than long. Distance between chitinous spots 1.9 times wider than long of prosternum at level of bottom of mesal incision; 2.3 or 2.4 times as great as length of dental line usually, this varying with number of teeth present Prosternal teeth 6 + 6 to 8 + 8, other numbers noted being 6+ 7, 7+ 7, and 7+ 8; either uniform or varying irregularly in size; subacute, being but narrowly rounded at tips. Spine proximally clearly stouter than the hairs, but distally bristle-like, curved; situated immediately ectad of outer tooth. First dorsal plate with surface similar to that of head. All finely 494 BULLETIN: MUSEUM OF COMPARATIVE ZOOLOGY. punctate. Plates roughened, more especially the caudal ones as usual, the elevations or irregular tubercles small. Posterior angles of seventh, ninth, eleventh, and thirteenth dorsal plates strongly pro- duced, processes of the seventh much as in devorans. Ventral plates densely punctate and with numerous fine impressed lines which mostly run out from the puncta. The usual longitudinal furrows clearly developed. Hairs of posterior plates more numerous and longer. Anal segment in male dorsally densely clothed with rather long and strictly straight hairs, these not being at all distally curved or uncinate. In the female the dorsal hairs of this segment are but sparse. Coxal pores transversely elliptic or in part subcircular; of medium size; in largest specimens more strongly transverse as in the preceding species. In number from 6, 6, 6, 4 to 8, 8, 8, 6, other arrangements frequently noted, being 6,).75. 74.55) Gist, 6) 53 6,16) 6,53 77 Gand ‘, U8; Srat Gonopods of male rather small; distally strongly rounded or broadly subconic; bearing six or seven long bristles. Gonopods of female with claw long and rather strongly bent or curved near middle; tripartite, the median division long and acute, the lateral small and also usually acute, the inner or more dorsal one con- siderably more distal in position than the outer one which is near the middle of length of claw. Basal spines long and stout, subequal, attenuated uniformly from base distad. Basal article with inner side nearly straight. Anal legs in female short and rather slender; the third, fourth, and fifth articles longitudinally suleate dorsally, the fourth being most deeply so; the third and fourth articles suleate ventrally. Penult legs similarly but less strongly suleate. In the male the penult legs are as in the female; but the anal legs are more strongly modified. Fourth article with dorsal surface depressed or bowed ventrad over middle and proximal portion; relatively thicker than in female; articles longitudi- nally suleate along mesal surface, especially so in the more distal ones. ee! three pairs of coxae laterally armed. Spines of first legs, ; of second, o " 5S 3 3; of third, 0: o og 3 3 of fourth to eighth, 23,2 OF . - = 5; of ninth to eleventh, 0: o. 33 —t of twelfth, a ae 2 or ie a 3-9, a Spine sometimes borne by trochanter of one leg while 0, 3, 2,2, pestoals absent from the other; of thirteenth, 9°{'3'3°5; of penult, 9'y°3°3°3 or 0, 3,2 = — », the anterior spines of fourth joint in latter case being mostly very small, tarsi ending in three claws but the anterior accessory one CHAMBERLIN: GENERA OF LITHOBIIDAE. 495 mostly minute as in related species; of anal 5 gs t, or, less often, ventral spines 0, 1, 3, 3, 2, the claw single. Length from 17 to 24mm. 0 i , .) PLATE 1, CHAMBERLIN.— Genera of Lithobiidae. Fig. 1. Fig. 2. Fig. 3. Fig. 4. Fig. 5. Fig. 6. PLATE A. Zinapolys zipvus Chamberlin. (Kootenai Co., Idaho. Type). Anterior portion of prosternum showing teeth and spines. Claw of 9 gonopods, subventral view. Basal spines of right gonopod of 9°, ventral view. Gonibius rex (Bollman). (Tallulah Falls, Ga. Type). Claw of left 9 gonopod, ventral view. Basal spines of right @ gonopod. Lithobius atkinsoni Bollman. (Taylors, S. C.). Basal spines of left 2 gonopod. BULL. MUS. COMP. ZOOL. LITHOBIUS. PLATE ets ECS ort pe PLATE 2. CHAMBERLIN.— Genera of Lithobiidae. Fig. Fig. Fig, Fig. Fig. Fig. a ar PLATE 2. Lithobius forficatus (Linné). (Philadelphia, Pa.). Anterior border of prosternum. Claw of @ gonopod, submesal view. Basal spines of left gonopod of 9, ventral view. Neolithobius vorax (Meinert), Left anal leg of o, mesal view. (Jackson, Miss.). Claw of right gonopod of 9, ventral view. (Ocean Springs, Miss.). Basal spines of left gonopod of 9. (Ocean Springs, Miss.). BULL. MUS. COMP. ZOOL. LITHOBIUS. PLATE 2 i a fa RAS F CHAMBERLIN.— Genera of Lithobiidae. Fig. Fig. Fig. Fig. Fig. Fig. bo PLATE 3. Neolithobius mordax (L. Koch). (Jackson, Ala.). Claw of right gonopod of °. Basal spines of left gonopod of ¢. Neolithobius transmarinus (L. Koch). Anal leg of o’, mesal view. (Canton, Miss.). Basal spines of left gonopod of 9. (Brookhaven, Miss.). Neolithobius xenopus (Bollman). (Macon, Ga. Type). Anal leg of <’, lateral view. Neolithobius underwoodi (Bollman). (Selma, Ala.). Basal spines of right gonopod of 9°. PLATE 3 LITHOBIUS. BULL. MUS. COMP. ZOOL. ee eee =, ACME BOOKBINDING CO., INC. NOV 2 9 1983 100 CAMBRIDGE STREET CHARLESTOWN, MASS. wi 3 2044 066 Gk aoa hee sey bre Shy ra ee B qine Ryans vary é rays a, cane pig's Nas “ a wa Be Bow Qt Mag a Vea BA ry et USA Bee Pe ae rt, ky My HARM INES ae ene} » vu Ai ga ee ON Sols A AAR IO AY “ PEL yet gee Wash wash cis Pes past 42 AUS Myla: Le gety Aaa i Soh ALO, ‘. ue eis Ws deacder eh at 1 a ays) be Got yd ke y be i,