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Harris Co. Cornell Univ., Ithaca, N. Y. WES. A.
CONTENTS OF VOL. IX
Buli. No. Page 37 —New Eocene Species from Alabama. Boye le el etic lies sl ond seen so Pl’s 1-3 1-32
38 —Recent Mollusca of the Gulf of Mexico
and Pleistocene and Pliocene
Species from the Gulf States.
Part 2.—Scaphopoda, Gastropoda.
Amphineura and Cephalopoda.
Bye Cee Maury correo oo. ye ae 33-173 39 —The Miocene of Northern Costa Rica
with Notes on its General Strati-
graphic Relations.
IBV. ae OSSOMny jc ote. 0 ent Pl’s 4-35 174-482
BULLETINS OF
AMERICAN PALEONTOLOGY
eS point ihd dl t
Vol. 9
No. 37
NEW EOCENE SPECIES FROM ALABANA
BY
hE VAT DRICH
March 6, 1921
*
Harris Co.
Cornell Univ., Ithaca, N. Y. OS Sy iA.
PREFACE.
The late Herbert H. Smith, curator of the Alabama State Museum of Natural History University, Alabama, made exten- sive collections in the Tertiary of Alabama, and at the time of his sudden and lamented death was engaged in classifying and arranging his material. The collections were made on joint ac- count for the Carnegie Museum of Pittsburg and the Alabama Museum of Natural History. The condition of the material made it necessary to go over all of it, and classify it for division as well as to describe what was considered new. The writer has also added a few new species from his collecting. It is regretted that Mr. Smith was not spared to complete his labors, because his ability in specific determinations was almost unique. The types of the new species collected by him are all in the Univer- Si buseum. We are indebted to Dr. W. H. Dall, Dr. Paul Bartsch, Prof. G. D. Harris and Dr. E. A. Smith for generous aid in both specific and generic determinations of doubtful forms.
NEw EoOcENE SPECIES FROM ALABAMA BY
TT. EAL DRICH
te,
Mitramorpha turriculata, n. sp. Plt, Rig
Shell ‘small, biconic, whorls eight, three embryonic and smooth, the fourth with strong longitudinal ribs, which continue over the other whorls, dying down some on the body whorl. Spiral strisze numerous, and impressed, a strong double band bor- dering the suture, aperture narrow, smooth within. Length 6mm.
Locality.—Choctaw Corner, Ala. Woods Bluff horizon.
Type.—My cabinet.
Remarks.—This little shell seems to be nearest to Mitramor- pha, it has a glazed appearance ; there is some evidence of a slight notch on and above the periphery of the body whorl.
Turris (Pleurotoma) nodoideus, n. sp. Pip ree igs 2, 3)
Shell small, spire pointed, whorls nine ; first four embryon- ic, the fifth bearing longitudinal ribs in part, and revolving lines in part with the ribs also present but without the subsutural col- lar below. ‘The sixth, seventh and eighth whorls with the bor- dering part of the ninth carrying first, a subsutural collar which becomes doubled and thicker towards the body whorl and is marked with very numerous raised lines belonging to the fasciole; second, just below is a rather broad concave revolving space reaching to the peripheral parts of the whorls carrying very numerous close set semicircular anal markings, these continue
6 BULLETIN 37 6
over the body whorl in curved raised lines : third, the periphery sometimes carrying a couple of coarse lines followed below by from four to six raised lines, also raised longitudinal ribs from the periphery to suture. Body whorl with raised revolving lines extending to the end of the canal, carrying a few alternating ones below the periphery ; aperture smooth within ; canal slightly FOStEate:
Locality.—Cave Branch, 6 miles E. of Thomasville, Ala. Woods Bluff beds.
Type.—Ala. Museum. University, Ala.
Remarks.—This species seems to differ in some respects from any heretofore described, but it may be only a variety of some existing species like 7uvris denticula Bast. et. al. The fact is thatthe Woods Bluff beds at Cave Branch contain so many numer- ous varieties that it is almost impossible to locate them properly. The whole subject of species in this family needs careful study. The young are often obtuse, and differ in shape and ornamenta- tion from the adult so that they could be called new species, and pass muster while a series would connect them all together. Am not prepared to go into details but hope this subject will be looked into by some of our younger students. Some specimens may be hybrids, and are altogether perplexing. One ought to have authentic specimens of the described species to even begin the examination. All the species of this group from this locality seem somewhat stunted.
Turris specus, n. sp., Pl. Pr. Bigs. 4,5:
Shell small, whorls probably seven (the embryonic tip is de- fective) first three embryonic, then four spiral whorls ornamented with a strong subsutural collar, then a slightly concave revolving fasciolar space covering about one fourth of the whorl, followed by a coarse cancellated surface ; the ribs cut by about four im- pressed revolving lines making a series of nodes, body whorl with the same ornamentation above followed by coarse revolving lines growing finer to the base ; outer lip smooth inside, columella thickened, base somewhat rostrate.
Locality —Cave Branch, Ala. Woods Bluff beds.
7 ALDRICH, HKOCENE MoLLUSCA 7
Type.—Ala. Mus. of Nat. History, University, Ala. Microdrillia rostratula Csy.
This species described by Col. Casey in 1903 bears a certain resemblance to Pleurotoma insignifica Heilpr. (Fusus nanas Lea) and has been referred to this species, but specimens before me show differences as follows: Col. Casey says the ‘‘body whorls are about four in number.’’ My specimens have five, also in the embryo there is sometimes but one costulate whorl, while Pleuro. insignifica Heilpr. has but three embryonic whorls, all smooth, and no costulate ones. ‘The two species seem to be quite distinct. The Texas form mentioned by Prof. Harris is probably the same as Heilprin’s species.
SPECIES TO BE ADDED TO THE Woops BLUFF FAUNA— FROM CAVE BRANCH, ALA.
Evia persracits Con. Cylichna deKayt Vea. Cancellaria alveata Con.
0 costata Lea. Pasithea guttula Lea (H. H. Smith). Cancellaria panones Harris. Ancillaria subglobosa Con. Levifusus sutert Aldr. Caricella dolita Con. Drillia lonsdalit Vea. Barnea alatoidea Aldr. Diplodonta nana Lea. Egerella subtrigonia Lea. Tellina leana Dall.
Turris bimoniatus, n. sp. Plot igi '6,
Shell small, whorls probably ten, spire on the type partially missing. Another specimen shows four embryonic smooth whorls. The four spiral whorls and part of the fifth ornamented with curved sinuosities covering the main part of the whorls, a
8 BULLETIN 37 8
strong collar below the sutures, the balance showing a double collar with a concave space between ; the whole of the surface with numerous spiral raised lines. Aperture about a third of the length of shell, lines on the base have a tendency to alter- mate.) Weneotlay aman:
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—Ala. Museum, University, Ala.
Remarks.— This seems to combine the characteristics of two or more species, and if there were only one specimen, could be considered a variety. It seems to be related to Z. moniliata Heilpr.
Pleurotoma (Peratotoma) Gardneri, n. sp. Pl. a.) Pigeyog a:
Shell medium, substance rather thin, broadly conic ; whorls nine to ten, embryonic ones four in number, the spire very small, but the whorls rapidly expanding, the permanent whorls profuse- ly ornamented with both longitudinal and spiral raised lines, the peripheral part and below carrying nodes while the spirals are alternate in size and thickness. Body whorl large, shouldered, profusely ornamented, the spirals nodular ; just below the suture there are a couple coarser than the others and also at base ; aper- ture half the length of the shell, smooth within, inner lip with a thin callus ; canal recurved and slightly spatulate. Length 21 mm.
Locality,—Pugh’s branch of Satilpa Creek, Ala., Gosport Sand.
Type.—Ala. Museum of Nat. History, University, Ala.
Remarks.—This species is on the borderland between /usus, Strepsidura and Pleurotoma. It isvery close to Pl. heilprinz, nobis from Jackson, Miss. The largest specimen is 25 mm. in length. The slit is not pronounced. Named in honor of Miss Julia A. Gardner of the National Museum.
Odostomia (Evalea) Bartschi, n. sp. Pls ity) Bie
Shell small, whorls six, spire almost totally immersed, shell rather rapidly expanding. Basal whorl large, suture deep, aper- ture oblong. One strong, straight fold on the columella placed
9 ALDRICH, EOCENE MOoLLUSCA 9
centrally; umbilicus open, inner lip somewhat recurved at base. Surface not smooth, carrying microscopic incised spiral lines. Length 3 mm. Locality. —Found at both Gregg’s and Bell’s Landings. Type.—¥rom Bell’s Ldg. Inthe Ala. Museum.
Turbonilla (Ptycheulimella) clinensis, n. sp. Pi ee hie 10:
Shell small, narrow, whorls eight, two embryonic, the next three longitudinally striated, balance smooth. Spire blunt, suture distinct, aperture small. Outer lip smooth within, inner lip with a strong fold curving up into the aperture and bordering the base. Length 3+ mm.
Locality.—Gregg’s Landing, Ala.
Type.—Alabama Museum, University, Ala.
Remarks.—Seems to be a distinct species. There are three other species of 7urbonzlla in the State Collection but too imper- fect to warrant description.
Turbonilla (Cingulina) tuscahomensis, n. sp. Play Pigs an: Shell small, whorls nine, the first three embryonic ; apex small, the fourth and fifth whorls with raised spirals, balance carrying five strong spirals, and somewhat cancellated ; suture deeply impressed. The space above the first or lowest spiral slightly wider than between the other four. Aperture broadly ovate ; inner lip slightly reflected ; base slightly spatulate. Length 4mm. Figured specimen from Bell’s Ldg. Locality.—Both Gregg’s and Bell’s Ldg., Alabama. Type.—Alabama Museum of Nat. History. Remarks.—tooks like a Bittium.
Turbonilla (Ptycheulimella) tardiusculus, n. sp. Pinte. cigs 12:
Shell with nine whorls, the first two embryonic, the balance smooth and of a dull brown color, substance of shell rather solid, suture lightly impressed, banded below. Aperture small, inner lip reflected at base, imperforate ; no fold showing on the col- umella. Two specimens found. Length4% mm.
Locality.—Cave Branch, Ala. Woods bluff beds.
Lype.—My collection.
IO BULLETIN 37 10
Pyramidella (iphiana) anita Ald.
The Nautilus, Vol, XXI, No. 1, p. 9, Pl. 1, Fie.\12)May.i1eo7:
This species was described as a 7urbonilla, but is now put into the Pyvamidellide by Dr. Bartsch.
Pyramideila (Syrnola) mitchelliana Aldr.
Bull. of American Paleontology, No. \22,'P.8,) Pl. 2 higea. ern.
Is also removed from Jurbonzlla.
The Ala. Museum has a number of new forms of small spe- cies, collected by Mr. Herbert A. Smith which are described herein. They have been submitted to Dr. Bartsch who has kind- ly indicated their generic position.
Pyramideila (Iphiana) obtusoides, n. sp. Pla) ie mes
Shell small, with four whorls capped by two embryonic ones turned abruptly to the back side of shell and partially immersed. The specimens, three in number, have a dull surface, apparently smooth, but under a glass numerous spiral lines are seen. Suture not deeply impressed ; aperture oblong-ovate ; outer lip sharp, smooth within ; inner lip reflected, somewhat twisted, carrying a single smalltooth. Length 3% mm.
Locality.— Found at both Gregg’s and Beil’s Landings, Ala. River, Ala:
Type.—Ala. Museum of Natural History.
Remarks.—A short, dull-looking species.
Epitonium munistriatum, n. sp. Pl 1.) igs anes
Shell small, aciculate whorls, probably 1oor more. Embry- onic ones three in number, apex pointed ; the next two or three with longitudinals only, balance cancellated. Whorls rounded, suture deep. Spirals 5 to 7, rather coarse, ribs raised with con- cave spaces between, base with smaller alternate spirals between the primaries ; intersections pointed. The base of this species (15) is in the Museum from Gregg’s Landing, Ala. The other specimen is from Woods Bluff, Ala. The aperture shows a a thickened inner lip where it joins the base, also a thin callus.
Locality.—As above.
Type—Ala. Museum of Nat. History.
XX ALDRICH, HOCENE MOLLUSCA II
Epitonium multiliniferum, n. sp. Pei HIS 16,
All the specimens are fragmentary, but well marked. Em- bryonic whorls missing ; suture distinct. Ribs on body whorl 34 in number, extending over the base, much weaker there ; very numerous close set spiral lines between the ribs. Aperture round, inner lip thickened, no umbilicus.
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—My cabinet.
Remarks.—This is a distinct species and it deserves differen- tiation from the other forms. The figure given by Prof. Harris in Bull. Am. Paleontology, No. 11, Pl. 12, Fig. 8, is probably a young example of this species. The sides are more nearly par- allel than in most Apztonzums.
Epitonium subacutum, n. sp. Pls t..) Bigs2 17, 13.
Shell thin, narrowly acute, whorls 12. The first five are em- bryonic and smooth ; the balance spirally striated ; cancellated. The vertical ribs on the body whorl about twenty-six in number, curving into the umbilicus. They are thin or acute ; the pri- mary spirals about ten in number with finer alternate lines be- tween ; inner lip with a slight callus. Length 10 mm.
Locality.—Cave Branch, Ala. Woods Bluff beds.
Type.—In Ala. Museum. One specimen shows the base and the other the embryonic shell.
Teinostoma subangulata Mr. var. Smithii, n. var. Pl. 1. Figs. 19, 20.
Shell small, flattened above, rounded below ; umbilicus not large ; a callus on the base which enters the umbilicus and con- tinues within. Bordering the suture is an appressed area raised above the whorls ; surface above with numerous close set revolv- ing lines ; periphery smooth with a few revolving lines below. Aperture circular ; outer lip somewhat thickened ; lines of growth perceptible ; interior somewhat pearly. Diameter 8 mm.
Locality.— Bell’s Landing, Ala.
Type.—Ala. Museum of Natural History.
Remarks.—Very much larger than the type of 7. subangu- lata Mr. Looks very much like a land shell. It differs some- what from Meyer’s species in the shape of the aperture.
12 BULLETIN 37 12
Teinostoma regularis, n. sp. Pl. 1, |, Pigs2a-¢.
Shell small but larger than the ordinary species of this genus ; whorls five, depressed above ; periphery rounded ; sur- face covered with fine lines which are crossed with lines of growth. Under a glass the intersections give the surface a beaded appearance ; a depressed space just below the suture. The base has coarser lines with a smooth space around the um- bilicus which is open ; aperture oval. Diameter 4 mm.
Locality.—Pugh’s Branch of Satilpa Creek, Clark Co., Ala. Gosport Sand.
Type.—Ala. Museum at University, Ala.
Remarks.—This little shell may possibly belong to the sub- genus Omphalius of Chlorostoma but it seems closer to Teznos- toma.
Ancillopsis Tuomoyi, n. sp. Plt: igs) 2a on.
Shell solid, thick, oblong, smooth and shining ; whorls 5-6 ; spire rather pointed ; the first four whorls small, the others rap- idly increasing ; a swollen callus reaching above the body whorl and also covering the front of this whorl and lapping over the back of this whorl. A deep suture reaches the aperture where it is rather wide ; a few irregular lines of growth are visible ; aperture about three-fourths the length of the shell, oblong-ovate; columella smooth, concave. The basal groove shows strongly at base, but only showing a short distance up where it is covered by the enamel. Length 28 mm ; breadth 21 mm.
Locality.—Bell’s Landing, lower bed, also one example from Gregg’s Landing, Ala. ‘The lower bed at Bell’s Landing is the same bed as at Gregg’s Ldg. Type from Bell’s Landing, lower bed.
Type.—Ala. Museum of Nat. History.
Remarks.—This species resembles /Zacron in some respects but is evidently an Anctllopszs.
Nassa pleona, n. sp. Pl. 1,, Figs) 25) 26:
Shell small, nuclear whorls broken off. Four remaining whorls carry longitudinal ribs that are crossed by two spirals,
53 ALDRICH, HOCENE MOLLUSCA 13
nodular at intersections. ‘Two close set spirals border the suture above and below same; suture channelled ; the last whorl car- ries about 10 widely spaced spirals, raised and nodular with con- cave spaces between. Outer lip thickened with the spirals ex- tending over same, denticulated ; inner lip enameled with the spirals showing through the same. Canal very short ending in a semicircular opening. Length 12+ mm. ‘Two specimens ob- tained.
Locality.—Ljisbon, Ala.
Type.—My cabinet.
Olivella semilignitica, n. sp. Pls apiiocu oy a2o.
Shell small, whorls six, four smooth ; the fourth and subse- quent ones bounded by a raised band just below the suture ; apex blunt, body whorl showing impressed lines of growth ; aperture asin the genus. Length 5% mm.
Locality.—Clark County, Ala. Woods Bluff horizon.
Type. —Ala. Museum of Nat. History.
Remarks.—The sutural callosity pronounced on the last whorl only. Differs from O. mediavia Harris in being smaller, more blunt, with fewer folds on the columella, and by a raised sutural band.
Gilbertia estellensis, n. sp. Bl 2), ABigs:/6,.% Shell globular, whorls three ; the first embryonic ; the sec- ond spirally striated but very narrow ; the third very globose and spirally striated ; outer lip thickened and broadly reflected ; a single tooth projecting into the aperture from it ; inner lip thickened with a broad spatulate tooth above, and two prominent teeth near the base ; the outer lip partially covering the second whorl. Under a glass the striz show that they are pitted as in wcddeon. Height 2% mm. :} breadth 2 mm. Locality.—Sucarnoochee beds, 3 miles south of Estella, Ala. Ty pe.—Ala. State Museum, University, Ala.
Cancellaria Giiberti, n. sp. Plgiie hie 620,
Shell with seven whorls remaining ; the spire is damaged ; cancellated ; suture deeply impressed and constricted, especially
14 BULLETIN 37) 14
on the lower whorls. Below the suture is a raised cord marked with a spiral line upon the lower whorls; above the suture is a wider space bounded by strong raised spirals with other raised spirals above. Body whorl expanded with a strong spiral at the periphery ; those above this are close together, those below al- ternated with a finer spiral ; lines of growth show on body whorl reversing near the base ; aperture oblong-ovate, the spiral lines rounding into same ; columella with two folds ; shell has a small umbilicus. Labrum lirate within. Length 9 mm.; breadth 4mm.
Locality.—Gregg’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—Only one specimen found. This species has many of the characters of ddmete.
Ovula regularoidea, n. Sp. Pl. 2.) ies Yo.
Shell small, elongate ; both anterior and posterior with fine revolving lines ; main body of shell smooth ; outer lip thickened and crenulated from beak to base ; inner lip also crenulated its whole length ; a heavy callus on the outer surface of the col- umella at the posterior end with the canal extending beyond the callus. On the smaller of the two specimens before me there are fine revolving lines half way down from the posterior ; aperture narrow, expanding back of the anterior canal ; ends somewhat flattened. Length of largest specimen 14 mi.
Locality.—Bell’s Landing marl, Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Cyprea estellensis, n. sp. Place
Shell small ; rotund ; smooth ; aperture narrow ; outer lip curving into the apex which is open. Lengthi2mm. Breadth 10 mm.
Locality. —Sucarnoochee beds, Pursley Creek, Wilcox Co., Ala.
Type.—Ala. State Museum.
Remarks.—This specimen has only preserved the shell in part but the interior is hard rock and gives the form quite accur- ately. The aperture is all filled with matrix, hiding the denti-
15 ALDRICH, EOCENE MOLLUSCA 15
tion. Prof. Harris mentions finding one species at Fort Gaines on the Chattahoochee and another (which is probably the one subsequently described % mile north of Ripley, Miss.) They are figured and named as they seem to be rare and peculiar to this horizon. Cyprea sp. ? Pia. Bigs. 2,, 4,
Shell small ; surface smooth ; basal callus extending par- tially over the sides ; outer lip expanded ; extremities somewhat pointed. One fragment shows the dentition, consisting of over 20 small teeth on the inner lip. Length of the larger fragment 15 mim.
Locality.—Sucarnoochee beds, one and three miles south of Estella, Ala. on Pursley Creek.
Types.—Ala. Museum of Nat. History.
Seems to be rather close to Cyprea Smitht nobts from Gregg’'s Ldg., Ala.
Siphonalia quadrilineata, n. sp. Pl, 2. Figs. 4, 5.
Shell resembling the Fuss subscalarinus Heilpr. Whorls eight, two embryonic, the balance more or less cancellated ; longitudinal costzee somewhat rounded; spiral stria very numer- ous, four fine ones between a coarser line ; the volutions are ap- pressed at the suture ; the spiral strize cross over the costee and the coarse lines make nodes of intersections ; canal reflected and twisted : the outer lip on the type is broken away but appears to have been smooth internally ; inner lip somewhat thickened. Length 17mm.
Locality.— Pugh’s Branch of Satilpa Creek, Clark Co., Ala. Gosport Sand.
Type.—In Ala. Museum, University Ala.
Remark.—One is rather rash to attempt a new form herewith but the surface ornamentation is peculiar and seems to be new.
Cerithiopsis estellensis, n. sp. Pla baoiro:
Shell minute ; whorls eleven or more, the first two smooth, the next two with raised ribs ; the balance showing two strong, heavily beaded spirals on the peripheral part of each whorl with
16 } BULLETIN 37 16
a third spiral much finer just below the suture ; one or more whorls are missing from the basal part.
Length 3 mm.
Locality.—Sucarnoochee clays, near Estella, Ala.
Type.—In Ala. Museum at University, Ala.
Levibuccinum (Euryochetus ?) harrisi, n. sp. Pl,.2, Bigs. 7-9.
Shell small ; whorls eight, three smooth, the other five be- low [with close-set raised lines ; outer lip smooth, slightly in- curved ; inner lip with a callus, thicker and slightly raised on the canal, which is short, rather wide ; shell somewhat truncated an- teriorly ; suture distinct. Length of largest specimen 4% mm.
Locality.—Cave Branch, Ala., about 6 miles east of Thomas- ville, Ala. Woods Bluff beds.
Type.—Ala. Museum of Nat. Hist., University, Ala.
Remarks.—This shell is quite fragile, the body whorl break- ing away from the spire. Prof. Harris figured the spire of this species under Aesopus erectus (Proc. Acad. Nat. Sciences, Phila., 1896, p. 476, pl. 21, fig. 3), which it resembles, but later on states it isnotthesame. ‘The Ala. Museum has three specimens, and there are two broken ones in my cabinet.
Type.—Ala. Museum of Nat. History, University, Ala.
TENUIACTALON, n. g.
Shell resembling Actgon but much more slender ; whorls nine ; aperture small, spire turned to the left and partially im- metsed ; suture deep; columella with one fold ; surface with numerous incised revolving lines.
Tenuiactzon pertenuis, n. sp. PL 2,.. Higy te;
Shell small, slender ; whorls 9 ; surface with numerous re- volving incised lines, stronger at base but almost disappearing at the shoulder of body whorl ; then just below the suture showing two or three stronger ones ; apex rounded and turned to the left, partially immersed ; outer lip somewhat incurved ; base round- ed ; columella with one strong fold ; the inner lip reflected over
the body whorl ; suture deep, giving each whorl the appearance of being inserted into the next succeeding one. Length 13 mm. ;
breadth of body whorl 4 mm.-; aperture 5 mm.
17 ALDRICH, HOCENE MOoLLUSsCcA 17
Locality.— Gregg’s Ldg. marl at Bell’s Ldg. and Gregg’s Rae. Ala. River, Ala. Type.—Ala. State Museum at University, Ala.
Teredo ringens, n. sp. Pion big. m2;
Shell substance thin, closed at the larger end like Auphus ; body of shell with raised, rather acute rings parallel to each other, and virtually at right angles to the longer diameter. Other fragments are not so stongly marked. Length of type specimen 37 mm. Average breadth 5 mm.
Locality.—Sucarnochee clay bed, 3 miles south of Estelle, Ala.
Type.—Alabama Museum, University, Ala.
Tellina estellensis, n. sp. Blo Ries, ssi.
Shell small, exceedigly thin, exterior smooth, except fine growth lines ; these are stronger near the ventral margin ; in- equilateral ; right valve with small laterals, larger in the oppo- site valve ; pallial sinus large, rounded ; anterior somewhat pointed ; posterior rounded. Length 13-15 mm.; height 9-11 mm.
Locality,—Sucarnochee beds, near Hstelle, Ala.
Type.—Ala. Museum of Nat. History, University, Ala.
Remarks.—This species is the same figured by Prof. Harris (eBulletins Am. Pal., Vol. 1, p. 182) asacast. It is rather com- mon but so very fragile it was almost impossible to procure a per- fect example.
Tellina cynoglossa Dall.
This is the common species in the Woods Bluff beds. It seems to be only avariety of 7ellina subtriangularis Aldr., which was described from half grown specimens, which is more regu- larly rounded posteriorly and also lacks the fold seen in older specimens. The full grown examples from Woods Bluff on the Tombigbee River measure as much as 25 mm. in length
Tellina bellsiana, n. sp. Plsa WRigs hits. Oy
Shell large, flat, thin ; nearly equilateral ; surface faintly concentrically striate ; lines growing coarser on the anterior end ;
18 BULLETIN 37 18
beaks very small ; the dorsal area strongly bent, pallial area gib- bous ; teeth normal. Height about 24 mm.
Locality.— Both Gregg’s and Bell’s Landings, Ala. River, Ala.
Type.—Ala. Museum of Nat. Hist.
Remarks.—One example in the Carnegie Museum. The type is broken some, shell thin for its size.
Tellina semirotunda, n. sp. Pl, 23° Pigin7,
Shell medium ; nearly equilateral ; anterior somewhat pointed ; surface smooth, except lines of growth irregularly spaced ; a slightly depressed area extending from beak to base behind the anterior fold. Cicatrices rather large, are well marked ; pallial area large ; cardinalsbifid. Length 16 mm. ; height 9 mm.
Locality.— Both Gregg’s and Bell’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Natural History.
Tellina semipapyria, n. sp. Pl..2.) \ Big ae,
Shell medium, resembling Tel/ina semtrotunda nobis , sur- face smooth except growth lines ; pallial sinus large, partially confluent with the pallial line ; only left valve known.
Locality.—Jackson’s Rockhouse branch. Woods Bluff beds. Ala. Length 23 mm. ; height 9mm.
Type.—Ala. Museum of Nat. History.
Remarks.—Bears a strong resemblance to the Claibornian species Tellina papyria Con. but is somewhat more rotund.
Semele langdoniana, n. sp. Pl. 2. Figs. up, ae.
Shell oblong, oval ; surface practically smooth, showing very fine lines that are coarser on the umbonal slope ; some spec- — imens show also fine radial lines reaching to the ventral margin ; posterior shortest ; ventral margin entire ; pallial sinus profound. Shell looks like a 7ellzna externally. Length of largest speci- men is about 27 mm.
Locality.—Bell’s Landing, Monroe Co., Ala. River, Ala.
Type.—(Left valve) Ala. Museum, University Ala.
Remarks.—There are several fragmentary specimens in the >
IQ ALDRICH, HOCENE MoOLLUSCA 19
lot. ‘This species is named after the late Dr. W. Langdon, an Assistant State Geologist of Alabama and the discoverer of the Floridian Miocene.
Semele monroensis, n. sp. Pl 2s, bigs. 20. 22:
Shell small, rather flat ; one right valve only in the collec- tion ; umbonal slope strongly bent ; surface smooth above with several widely spaced lines towards the ventral margin ; hinge stout ; ligament long for the genus ; the umbonal slope marked interiorly by a few radial lines. Length 5 mm.
Locality.—Bell’s Landing, Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—The specimen described is quite young but dis- tinct. Looks like a Weretrix in shape and ornamentation.
Lucina primoidea, n. sp. Plas. [Figs, 24/04:
Shell with many raised lines ; beaks recurved, higher than long. This species is figured because it is from an horizon close to the Cretaceous and is doubtless an ancestral form of Lucina cornuta Conr. of the Claibornian. The interior is not accessible.
Locality.—Black Bluff, Tombigbee River, Ala., about 20 feet above the base.
Type.—My collection.
Martesia recurva, un. sp. Pie, Sigs 25.26.
Shell small, short, rotund; with two accessory plates just forward of the umbones ; groove running nearly vertical from beaks to base and strongly marked ; concentric striz strongly marked on the posterior end, bending abruptly at the groove and thence running up behind the umbones and under the accessory plates ; the anterior end rounded and smooth ; cardinal margin bordered with a thickened and raised process. Length 8 mm. ; height 4% mm.
Locality.—Fleming’s Mill, Ala., on Pea River. Nanafalia beds. ,
Type.—My collection.
Remarks.—This species is more obtuse than the usual forms. Three examples found. The interior is not accessible.
20 BULLETIN 37 20
Rochefortia minuta, n. sp. Pl, 2: " Figs e728
Shell, small, ovate ; nearly equilateral ; periostracum still adherent, surface showing a few incremental lines ; beaks point- ed, with a resiliary pit underneath with short lamellee on each side, the posterior one the longest ; muscular scar slightly im- pressed ; the right valve has a space for the left lateral ; pallial line simple. Resembles somewhat Rochefortia Stimpsoni Dall. The space in our specimen under the beak looks as though it had been excavated out of the dorsal side, but such is not really the case. Prof. W. H. Dall has examined this specimen, and veri- fied its generic position. Am greatly indebted to him in a num- ber of cases, Length 3 mm. ; height 2mm,
Locality.—Bell’s Ldg., Alabama River, Ala.
Type.—Ala. Museum.
Montacuta bicuspidata, n. sp. PI.12.), ies. page 20:
Shell minute, oblong-ovate ; surface with incremental lines ; beaks carrying the prodissoconch, nearly central ; viewing the shell from the outside and above, it shows two tooth-like projec- tions, one on each side of the beak, rising from the dorsal mar- gin and another shorter one which does not show like the two first mentioned ; ends rounded ; base arcuate ; hinge with a small subtrigonal tooth and also a space for a lateral in the other valve. Pallial line simple ; adductors distinct but not deep. Length 234 mm ; height 2 mm.
Locality.—Bell’s Ldg., Ala.
Type.—Ala. Mus. of Nat. History, University, Ala.
Basterotia ? prima, n. sp. Pl 3. Sigs: aa ae
Shell small ; valves rather thick ; subquadrangular ; beaks subterminal ; shell profoundly gaping at the anterior ventral part ; a depressed space running from umbo to ventral margin ; surface nearly smooth, showing growth lines which also show in the interior. Anterior adductor scar deep, with a raised border buttressed to hinge plate ; posterior scar slightly im- pressed, pear-shaped and both rather large for the size of shell ; one cardinal tooth in right valve, corresponding to a deep socket
aI ALDRICH, EOCENE MOLLUSCA 21
in the other ; a longer tooth below. Three single valves in the collection. Length 8mm. ; height about 7 mm.
Locality.—Gregg’s and Bell’s Landings, Ala. River, Ala.
Type.—In Ala. Museum.
Remarks.—This species is doubtfully placed in Basterotia ; the large gap suggests an attachment to some other form, but is not irregular. Prof. Dall has kindly examined it, and suggests that it might be commensal.
Saxicavella alabamensis, n. sp. Blogs) Bigsias,
Shell inequivalve ; thin ; posterior much broader than the anterior ; outer surface showing irregular lines of growth, stronger over the umbonal slope ; an upright cardinal tooth directly under the umbo, with a short lateral on posterior side in the right valve ; anterior muscular impression distinct ; pallial line faint. Only one valve in the collection. Length 6 mm. ; height 4% mm.
Locality.—Bell’s Landing, Monroe Co., Ala.
Type.—In Ala. Museum.
Montacuta Herberti, n. sp. Bla Hage:
Shell small; inequilateral ; dorsal and ventral margins rounded ; each end rather blunt; the posterior somewhat angulated ; surface smooth with very fine concentric striations, and a few lines of growth near the ventral margin ; shell inflat- ed ; cardinals feeble, showing in left valve only ; beaks low, rather pointed, a small buttress under them. Length 9 mm ; height 6% mm.
Localéty.—Cave Branch, Ala.
Type.—Ala. Museum of Nat. History.
Remarrs.—One valve in my collection. Named in honor of the late Herbert H. Smith.
Ervilia lignitica, n. sp. Plaga bigs) G19!
Shell small, inflated ; nearly inequilateral ; rather solid ; the posterior slightly longer than the anterior ; both ends round- ed ; surface carrying a few irregularly spaced grooves of growth; interior polished, shining ; pallial sinus small, not reaching to
22 BULLETIN 37 22
center ; umbones small, dentition as in the genus with short and strong teeth. Length 6 mm. ; height 3% mm.
Locality.— Both Gregg’s and Bell’s Landing beds, Ala. River, Ala.
Type.—Ala. Museum of Nat. History, University, Ala.
Remarks.—This species has a strong resemblance to Arvzlia polita Dall from the Floridian Pliocene. The young seem to be higher in proportion to their length than the full grown spect- mens.
Panopea bellsensis, n. sp. Pl, 3. Figs, go, 07-
Shell rather large, thin, with a brown epidermis ; lines of growth irregularly spaced ; shell compressed, nearly equilateral ; beaks small, appressed to and partly covered by the reflected dor- sal margin; one large cardinal tooth ; ligamental attachment short ; interior porcellaneous with growth lines showing through; pedal scars small. Length 30 mm. ; height about 13 mm.
Locality.—Bell’s Ldg., Ala. River, Ala.
Type.—Ala. Museum of Natural History.
Remarks.—One valve and some fragments in the collection. This species has its epidermis preserved. The shell is extremely thin and delicate.
Donax acutangula, n. sp. Pl, 3.) sea ir:
Shell nearly equilateral ; pointed at both ends, the umbonal slope bent abruptly to almost a right angle in old specimens ; surface nearly smooth ; fine growth lines showing ; ventral margin entire ; beaks small, pointed, smooth ; hinge well devel- oped ; laterals very long, striated on the inner edges. Length about 31 mm. ; height 15 mm.
Locality.—Bell’s Landing, Ala. River, Ala.
Type.—Ala. Museum of Nat. History, University Ala. Remarks.—One whole valve and several fragments of both valves received.
Arca (Cucullaria) ozarkensis, n. sp. Pl, 3. Eiesr3, ag. Shell small, rounded at both ends, giving the shell a quad-
a
23 ALDRICH, HKOCENE MOLLUSCA 23
rangular outline ; beaks small, compressed and pointed ; surface with rather close set radial lines on the posterior slope ; balance of the surface smooth ; teeth as in the subgenus, three or four anterior and two posterior ; muscular scars shallow but plainly marked. Length 5 mm. ; height 3 mm. | Locality.—Woods Bluff beds near Ozark, Ala. Type.—My cabinet.
Ostrea intermedoides, n. sp. Pla. Pigs.154, 16:
Shell medium, valves rather thin ; surface in older speci- men with numerous raised concentric ribs ; the lower valve bent downwards near the beak ; the umbo shows radial strize ; mar- gins crenulated internally ; a fragment of the upper valve ap- pears to be flat. Length of lower valve 32mm. ; breadth about 20 mm.
Locality.— Bell’s Landing marl, Bell’s Landing, Ala. River, Ala.
Type.—Ala. Museum of Nat. History.
Remarks.—Have hesitated to add another oyster to our Eo- cene, but I have been unable to find a place for this species. Four lower valves and a fragment of the upper valve are in the collection. The ornamentation recalls O. falco Dall from the Jacksonian.
Psammobia Smithi, n. sp. PAs a ESS, ol LG,
Shell large ; substance of shell thin ; oblong-ovate ; rather compressed ;_ surface shining, with fine concentric sculpture medially ; both ends coarser and elevated ; on the posterior slope the laminze are rough and somewhat mammillated ; beaks very small ; muscular scars distinct but shallow. Length 60 mim ; height 32 mm. ; Breadth about 12 mm.
Locality.—Gregg’s Ldg. marl at Bell’s Landing, Ala. Riv- er, Ala.
Type.—Alabama Museum of Nat. History, University Ala.
Remarks.—Both valves of this splendid species are in the collection. It bears a general resemblance to P. flosa Conrad, but the dimensions are very different. Collected by the late H. H. Smith and named in his honor.
24 BULLETIN 37 24
Psammobia harrisi, n. sp. Pl. 3.\\ /Pigssimeyige
Shell nearly equilateral ; truncate behind, with fine concen- tric sculpture, coarser at each end ; ligamental area short ; cica- trices distinct ; lateral tooth long ; posterior one short. Length about 37 mm.
Locality.—Gregg’s Landing, Ala. River, Ala.
Remarks.—This species resembles P. ozarkana Harris but is _ more obtuse. The type is not perfect.
DIVERS * (Echinocyamus ?) meyeri, n. sp. Pi, 3.) PIGS.) 2m, 22:
Test small, ovate; flattened below ; deeply pitted with ovate scars above which become nearly square on the sides and below ; substance of the test of moderate thickness ; apicial system near- ly central ; ambulacral petals relatively short, straight, with numerous round pores. Peristome is medium circular, and slightly depressed ; periproct not small, closer to the peristome than the side. Length 5 mm. ; width 3% mm. ; height 3 mm.
Locality. —Gosport Sand, Claiborne, Ala.
Type.—My cabinet.
Remarks.—This species is peculiar in the deep pits covering the surface. It is larger than &. huxleyanus Meyer and more obese though egg-shaped. Received from Dr. Otto Meyer in whose honor I have named it.
NOTES BLACK BLUFF OR SUCARNOCHEEH HORIZON.
Prof. H. H. Smith and wife collected a number of species from these clays not far from Camden, Ala., in the vicinity of Estelle, Ala. The deposit consists of an aluminous clay, dark chocolate in color, and contains the greater part of the species mentioned by Prof. G. D. Harris from the bluff at Fort Gaines on the Chattahoochee River where they seem to be represented principally by casts. Those near Estelle are rather numerous, but very fragile. A few new species are described in this paper.
Ny i
25 ALDRICH, HOCENE MoLLusca
A surprising fact is that a minute Cordula, Corbula (Aloidis) miltum Dall, originally described from the Wilcoxian is extreme- ly common. ‘The supposed differences in the specific fauna is explained. The two localities are over 100 miles apart. ‘The list of species from near Hstelle is as follows :
Leda saffordana Harris. Vl Parva Rogers. guercollts Harris. milamensts ‘' ii Corbula ( Alotdis) miliune Dall. Glycymertis cf aviculoides Con. Pinna sp. ? Teredo ringens Aldr. Yoldta eborea Con. Venericardia wilcoxensis Dall. Tellina estellensts Aldr. Protocardia Harrist Dall. Pecten alabamensis Aldr. Ostrea ? young. Nucula ovula Vea. a mediavia Harris. Modiola sajfordi Gabb. Meretrix ripleyana Gabb. Cucullea saffordt Gabb. R macrodonta Whitt. Crassatellites sepulcollis Harris. fe gabbi Safford. wyca Sp. ? Corbula subcompressa Gabb. Strepsidura Fleilprint Aldr. Volutiithes rugatus Con.
ty limopsis Con. Lurritella tennesseensis Gabb.
66
66
(a
morton? Con.
humerosa Con. Turritela alabamtensts Whitt.
26
BULLE TING 27 29
Triton showaltert Con. Solarium periscelidum Dall. Solariella alabamensis Ald. Rtssoina alabamensts Ald. Pseudoliva unicarinata Ald. a scalina Heilpr. Pleurotoma guercollis Harris. 5 persa Whitt. medtavia Harris. a adeona White. longipersa Harris (var.) Calyptraphorus velatus, var. contpressus Aldr. Olivella mediavia Harris. Natica saffordia 2?“ OS << YeUer Sa NN With: onusta os eminula Con. LMU Anmaura tombigbeensis Harris. Mesalhia watsonensts Harris. ‘‘ pumila Gabb (var. ) aH alabamiensis Whitt. Levifusus pagoda Heilpr. ; dalez Harris. fusus guercollis Harris. ‘*’ ottonts, Aldr. ‘oS tortiles. WW ith. 8" “nonhri Alar Dentalium mediaviense Harris. Cadulus turgidus Mr. Atys robustotdes Aldr. Tornatella_quercollis Harris. Cylichna meyert Aldr. Gilbertia estellensts Aldr.
Cyprea 2 sp.
37
ALDRICH, EHOCENE MOLLUSCA
Extilia pergracits Con. Nautilus sp. Fragments.
Flabellum conoideum Vaughan. Balanophylla haleana M. Hd. & H. Stenocyathus n. sp.
— ——_—_—_ _—_—
Foraminifera, 12 species.
27
28
Figure
75 19,
21, 235 25> 27)
~
Il.
WP
BULLETIN 37
EXPLANATION OF PLATE 1
28
Page Mitramorpha turriculata, n, sp., Choctaw Cor. Length6mm... 5 Turris nodoideus, n.isp., Cave Branch. eigenen gmm... 5 Turris specus, n. sp.,'Cave Branch. j.t.5...0 sont nearer Om. <1) 46 Turris bimontatus, n. sp., Cave Branch: .icereeaneee TA HOM Sg Pleurotoma ( Peratotoma) Gardneri, n. sp., Pugh’s Br. 21 mm... 8 Odostomia ( Fvalea) Bartschi, n. sp., Bell’s Ldg............ 2) sama. ;)!) 8 Turbonilla ( Ptycheulimella) clinensts, n. sp., Gregg’s
TAM an ooo. sdlsac sansa sistenbielels/aacd see MGlee ee eee ema nee 3 IM 9 Turbonilla ( Cingulina) tuscahomensis, n. sp., Gregg’s
Tani ng’ ovcceioss cddeasiadesnade nap ranushacee aeeee ees aaa aaa 4mm... 9 Turbonilla ( Ptycheulimella) tardiusculus, a. sp., Cave
Brae ee eee ee RU eee a 4.5mm... 9 Pyramidella (Iphiana) obtusoides, n. sp., Gregg’s Ldg. 3.5 mm... 10 Epttonium munistriatum, n. sp., Jackson’s Rock House
Brancyy, isi se sass eel eed decay AU aes eee en 4 mm... 10 Epitonium munistriatum, n. sp., Gregg’s Landing...... 2 mit), 1G Epitonium multiliniferum, n. sp., Cave Branch......... 10mm... II Lipitonium subacutum, nun, sp., Cave Branch.,.......00..202 10 mm... II Teinostoma subangulata Mr., var. Smithi, n. var.,
Bellis: Landings eese i ae 8 mm... II Teinostoma regularis, n. sp., Pugh’s Branch.............+6 4mm... 12 Ancillopsts Tuomeyi, n. sp., Bell’s Landing............... 28 mm... 12 NG@SSa Alaona:.s Spe NMASWOR GC Cee eG slguveen ime anne 12+ mm... I2 Olivella semtlignitica: a. sp.)) Clark (Coli) (i520. 5.5 mm... 13 Cancellaria Gilberti, n. sp., Gregg’s Landing............... gmm... 13 Ovula regularoides, n. sp., Bell’s Landing........... ...... 14mm... 14
9, Pl. {
‘ >
ALDRICH, KOoCENE MOLLUSCA
PLATE 2 (2)
30 BULLETIN 37 30
a
EXPLANATION OF PLATE 2
Figure Page 1. Cyprea estellensis, n.sp., Pursley iCr, ...:....:a0seceeeent 12mm... 14 2.3... Cyprea sp., Pursley Cr .c...0so0s0c- ene sereneseeteseskne eee 15 mm... 15 4,5. Stphonalia quadrilineata, n. sp., Pugh’s Branch......... 17 mm... 15 Gilbertia estellensts, n: sp., Estelle........0.-3:50heeseee 2, gx2 mtn)... 13
7-9. Levibuccinum (Euryochetus ?) harrisi, n.sp., Cave Branch. ; ‘largest specimen, .....5.s.0ce,peceeeemeeee 4.5 mm... 16 10. Tenutiacteon pretenuts, n. gen. and sp., Gregg’s Ldg....13 mm... 16 Ii. Cerithiopsis estellensis, a. sp.,.Estelle...../2..51.5. eee 3mm... 15 12,. Leredo ringens, n. sp., Estelle -.27..0.):0 eee 37 tmim,,: \17 13, 14. Tellina estellensis, n. sp., Estelle......... largest sp....15XII mm... 17 15,16. Tellina bellstana, n. sp., Bell’s Ldg. ; height.............. 24 mm... 17
17. Tellina semirotunda, n. sp., Gregg’s Ldg. ; length......16 mm... 18 18. Tellina semtpapyria, n.sp., Jackson’s Rockhouse
Branch :; ‘lengthy... ....6.502[spenctslecabecvenesaes ae memeem 23 mm... 18 19, 20. Semele langdoniana, n. sp., Bell’s Ldg.. .........scseeceeee 27mm... 18 21, 22. . Semele monroensts, 0. sp., Bell’s Ude..: ..:......:2s:ceceeereeh 5 mm... 19 23, 24. Lucina primotdea; mn, sp., Black) Bluff. .0.......25):02 58 9x9x6 mm... 19 25, 26. Martesia recurva, n. sp., Fleming’s Mill, Nanafalia ©
DES les cca act arce tae sth neck attend socio seater ann emene 8x4.5 mm... 19 27, 28. . Rochefortia minutia, np. sp., Bells: Udg...... 00... .01-seeeeees 3 mm... 20
‘29, 30. Montacuta bicuspidata, n. sp., Bell’s Ldg............... 2x23{ mm... 20
Vol. 9, Pl. 2
Paleont.
Amer.
Pl. 2, No. 37
a
A LD RIC H , HOCENE M OI aU SCA
Ja ky 2)
aI
32
Figure E213. A, 5.
10;
20.
22),
BULLETIN 37
32 EXPLANATION OF PLATE 3
Page Basterotia ? prima, un. sp., Grege’s Ldg.........ccsedassens 8x7 mim...:20 Saxicavella alabamensis, n. sp., Bell’s Ldg.......... 6x45 mean MMontacuta Flerberii, un. sp., Cave’ Bro. aj. 9 m0... 28 Erulia lignitica, n..sp:, Bell’s dudes... (cau eee 6mm... 21 Fanopea bellsensts, nu: Sp., Bell's Wde. eu seas 30x13, mm...’ 22 Donax acutanzula,n. sp., Bells) Ldg.... 2.22 eae 31XI5 mm... 22 Arca (Cucullaria) ozarkensis, n. sp., near Ozark...... 5x2 MM. ee
Ostrea intermedoides, n. sp., Bell’s Ldg. No. 15, 24x19 mm. No, 16, 32x22) sn... 4234 Psammobia Smitht, n. sp., Bell’s Wdew...:. cc tceee 60x32 mm... 23 Psammobia harrisi, n. sp.; Gregg’s Ldg.. i...) s...geee 27 aia...) 24 (Echinocyamus ?) meyert, n. sp., Claiborne...... 5X344x3 mm... 24
Vol. 9, Pl. 3
Paleont.
Amer.
Bull.
Pl. 3, No. 37
x x
<
BULLETINS
OF
AMERICAN PALEONTOLOGY
Vol 9
*K
No 38 RECENT MOLLUSCA OF THE GULF OF MEXICO AND PLEISTOCENE AND PLIGCENE SPECIES FROM THE GULF STATES
Part 2
Scaphopoda, Gastropoda, Amphineura, Cephalopoda | BY
CARLOTTA JOAQUINA MAURY
January 31, 1922 [Part I, PELECYPODA is Bulletin No. 34, 1920.]
Harris Co. Cornell University, Ithaca, N. Y. ie San
BULLETINS
OF
AMERICAN PALEONTOLOGY
--3K DRS ee a Nee
Vol 9
*
No 38 RECENT MOLLUSCA OF THE GULF OF MEXICO AND PLEISTOCENE AND PLIOCENE SPECIES FROM THE GULF STATES
Part 2
Scaphopoda, Gastropoda, Amphineura, Cephalopoda BY
CARLOTTA JOAQUINA MAURY
January 31, 1922 [Part I, PELECYPODA is Bulletin No. 34, 1920.]
Harris Co. Cornell University, Ithaca, N. Y. Ue S.A.
4 MOLLUSCA OF THE GULF OF MEXICO a4
OHUIASS SCA rar ODA ORDER SOLENOCONCHA Genus DENTALIUM Linnzeus
lagueatum Verrill, Trans, Conn., Acad, Arts Sci., vol. 6. pt. 2, peua2ad,) pl. 44, £./18, 1885; Dall, Bull. Mus, Comp. Zool., eee 2 pei4z0, pl 27 ite Tt 889%) Ball a7. Uy S. Nat! Pieper 76, ‘pl. 27) 1. 1; pl’ 46, £..18, 1889; Pilsbty and Sano) lnyon s' Man. Conch:, 17, p. 10; ‘pl. 7; f..1,°2; pls) mug nuso7, lenderson, Bull. rrr, U.S. Nat. Mus., p) 23, Pert.) 6,/'7, 1920;
Distribution—Hatteras to Barbados, Gulf of Mexico: West Florida and off the Mississippi delta, 60-68 fms.
texasianum Philippi, Zeitschr. Malak, p. 144, 1848; Pilsbry & Suatp,|iryon’s Man. Conch., 17; p..22, 1897; Henderson, ements. IN. Mo, p: 28, pl. 1, f.'2, 4, 5). 9, 1920:
sexangulare Wilgard, House of Rep. Ex. Doc. 1, pt. 2, pp. Sa7ese, pl. 2. f..7) 1878.” Not of Lamarck.
wowlda \Singley, 4th Ann. Rept. ‘Texas Surv., p. 331, 1892; Dall, List Cameron Sh. Not the true gouldit of Dall. Note.—This may be identical with Chenu’s D. americanum 1843. Closely related to gouldzz Dall, with whichit has been confused.
Distribution.— N.Carolina to Yucatan, 3-9 fms. Gulf coast: Point au Fer and Cameron, La.; Galveston, Corpus Christi and Pass Cabello, Tex. Pleistocene, Lake Borgne bor- Mags,” La. texasianum cestum Henderson, Bull, 111, U.S. N.M., p. 29, eon tT) 4.) 6. O20:
Distribution.—Pass Cabello, Texas. Probably in very shallow water. |
rebeccaensis Henderson, Bull. 111, U. S. N. M. pp. 16, 31,pl. 3, Hoye. 1920.
BULLETIN 38 35
an
Distribution.— West Florida, 7-16 fms. and the Tortugas.
(Antalis) antiliarum d’Orbigny, In de la Sagra’s, Hist Pol. y Nat. Isla de Cuba, 2, p. 202, pl. 25, f. 10-13) 18die; Eilspes: & Sharp, Tryon’s Man. Conch:; 17, p. 57, pls aay gees, 1897; Henderson, Bull. 111, U.S. N. M.) pial 4, 6-8, 1920. Note.—Contined to shallow water. Deep water citations are referable to different species. Distribution.—Florida to Barbados. Gulf Coast: Charlotte Harbor and other stations off West Florida, 2-14 fms. (Antalis) pseudohexagonum Ihering. See Henderson Bull. rrr, Uns. NUM. 9. 46; ‘pl vO. tat -2 1926: Distribution.—Florida to Brazil. Gulf coast: Marco, Tampa and Charlotte Harbor, West Florida, 2-5 fms.
(Antalis) disparile d’ Orbigny, Hist. Pol. y Nat. Isla de Cuba, 2, p. 202, pl. 25, f. 14-17, 1846; (?) Vanattase Broa aeas, N.S. Phila, 55, p..757,:1903; Henderson, ‘Bulg ais INC os 7 be Os ei Oh O20:
Note.— Frequently confused with D. antillarum by authors although really closer to fexastanum.
Distribution.—Florida to Rio de Janeiro, Brazil. Gulf coast; Tampa, Fla. Littoral zone.
(Antalis) ceratum Dall, Bull. Mus. Comp. Zool., 9, p. 38, 1881; Idem, 18, p. 424, pl. 26, f..5, pl. 27, f. 2, 188op nollie S. N. M., p. 76, pl. 26, f: 5, 1889: Henderson) sai ore 5. NY Miipp. 40-5i,plygad.|2-7, 1o20: Distribution—Florida to Barbados. Gulf of Mexico: Off Mississippi delta, 321 fms.; Yucatan Banks, 84 fms.
(Anialis) taphrium Dall, Bull. M. C. Z., 18, p. 422, 1889; U.S. N.M.,p. 76, 1889; Pilsbry & Sharp, Tryon’s Man. 17, p. 58, 1897; Henderson, Bull. 111, U.S. .N. M., p. S3;0b gee 1920. :
@>)
36 MOLLUSCA OF THE GULF OF MEXICO
Distribution—Hatteras to Cuba, 22-182 fms. Type dredged between Mississippi delta and Cedar Keys in 30 fms.
(Antalis) bartletti Henderson, Bull. 111, U.S. N. M., p. 55, pl. eG 2.7), 1Q20.
peewerine, Wall: Bull. Mo. C. 2; 0, p. 37,).1881; Bull. 37, Persone Mi. p76. (gars), 1889. Not anidlarum. da’ Or- bigny, 1846.
Note.—True antillarum is semi-littoral while dartleffz is in deeper, colder waters.
Distribution.— Florida to Martinique. Gulf of Mexico: Be- tween Mississippi delta and Cedar Keys, 227 fms.; Yucatan Banks, 399 fms.
(Heteroschisma) callithrix Dall, Bull. M. C. Z., 18, p. 427, pl. 27, Potommosoo Bull a7. WU. SN. Mp. 76, pl.\27) f..10) 18389; Pilsbry & Sharp, Tryon’s Man. Conch 17, p. 62, pl. 7, f. 3, nea, ienderson, Bull. 111,.U..S. Ni M., p. 57, pl. 8, f. 3. 4010) O20.
Distribution—North Carolina to off Rio de la Plata, Ar- gentina. Gulf of Mexico: Between Mississippi delta and Cedar Keys, 1181 fms.; Yucatan Strait, 640 fms. (type lo- cality).
(Fissidentalium) meridionale Pilsbry and Sharp, Tryon’s Man. Borch 17, p. 73, pl.i1s, f. 32-34, 1897, (as variety of caz- ida) Wwenderson, Bull ri, U.S. N. M., ps 61, pl: o, f. ZN LQ20.
Ae Mall. Bull) Me ©: Z. 9, p. 37, .188t; 18, p. A225): T8o03 Broc, U.S. Ne M. 12;/p. 204, 1889. Not cevas Watson.
Distribution—Type dredged by the Albatross, 240 miles south- east of Rio de Janeiro, 671 fms., 37° temperature bottom. Also found Yucatan Banks, 1568 fms. and off Barbados. The largest Dentalium of the western Atlantic and charac- terized by its unique, ashen color.
7 BULLETIN 38 37
(Graptacme) ebcoreum Conrad, Proc. Acad. N.S. Phila., 3, p. 27, 1846; Pilsbry and Sharp, Tryon’s Man. ) Conch. 7. ip, 89, pl. 16, f. 47-49, 55, 56, 1897; Henderson) Balk rea:
©. N. M., p:.66, pl. 10, £; 3-5, 3, 9,. ple iy ieee es leptum Bush, Trans. Conn. Acad. Arts and ‘Sei sens
6, p. 470, pl. 45, f. 18, 18@, 1885; Dall) ABaie7 ee IN) ME pe 76 188o;
)
matara Dall, Bull» M. C:. Z., 18, pi 4e@0y asso." Bitianes, USS. Ne Moy pious:
Distribution.— Hatteras to Barbados. Gulf coast: Sarasota Pass (type locality), Charlotte Harbor, Fla. and between Mississippi delta and Cedar Keys. Beach to 111 fms.
Note.—Mr. Henderson unites matfara and Jlepbtum with choreum, noting that mazara is the perfectly smooth adult form of the species.
(Graptacme) semistriolatum Guilding, Trans. Linn. Soc. Lon- don, 17, pt. 1, p.34, pl. 3, f. 1-5, 1834; Chema) Concent p. 7, pl..4, f, £4, 15,,1843; Henderson, Bull’ ama a oe Mee 60. plait ft 1-5) 8,6. 5020.
Distribution.— An abundant littoral species. Florida to Barbados. Gulf coast; West Florida in shallow water, 7-12 fms.
(Graptacme) sericatum Dall, Bull. M. C. Z., 9, p. 37, 1881; 18, Dp. 423,-pl. 26, f..1,,1889; Bull. 37, U.S. Nigiy io eee 26, f. 1, 1889. Henderson, Bull. 111, U. Soe PLT 2, Onn mOZO:
Distribution.—Type only found. Dredged by the Blake, 640 fms., Yucatan Banks.
(Laevidentalium) callipeplum Dall, Bull. M. C. Z., 18. p. p. 419, pl. 27, £.126,:1889; ('Bull.27,°U. 8. N. Min eigee geg f. 12 6, 1880; Trans. Wagner Inst. Sci., 3, page yse2: Pilsbry and Sharp, Tryon’s Man. Conch., 17, p..teg,)p)a¢)
38 MOLLUSCA OF THE GULF OF MEXICO 8
i owree7: Henderson, Bulli1ir, U.S. N. M., p. 74, pl. 12, Pease 1O20.
Distribution.—Florida to Porto Rico. Gulf of Mexico, dredged to 169 fms. between Mississippi delta and Cedar Keys, (type locality).
(Laevidentalium) perlongum Dall, Bull., M. C. Z., 5, p. 61, 1878, Wonlen wudune): 9; p.'36, 1881; 18, p. 4109; ~pl. 27; f. 6, meson mmimbls 270) Wie. IN. Mi. p..76) pl..27) £26, 1889; Hen- Merson wmotiily ITs Wao.) IN My pe 75; ply oO, fl, 1920.
Distribution.—Hatteras to Rio de la Plata, t11—1330 fms. Gulf of Mexico between Mississippi delta and Cedar Keys at 940 and 1181 fms. also elsewhere in the Gulf at 1330 fms.
(Episiphon) sowerbyi Guilding, Trans. Linn. Soc. London, 17 Pear piiet.7. 1834: Pilsbry and, Sharp, Tryon’s Man.’ Women| 1 7. p.(1r7,\ pl. 20, f. 30, 1897; Menderson; Bull. 111, ee srNG VE) pp. 77-80, ple 13. f..2)'3, 10, 1920,
peewee) Bull, 37, U. 8. Nii M.,. p.<76, 1889. \ Pro- bably not jAlum of Sowerby, 1860, which was from the east Atlantic.
Distribution.—Hatterasto Barbados. Gulf of Mexico: Off _Mobile, Ala., 27 fms.; between Mississippi delta and Cedar Keys, 60 fms.
sowerbyi peiliceri Henderson, Bull. 111, U. S. N. M., p. 80, leaf 7335.94 LO20: Distribution.— Florida to Cuba, Cape Catouche, Yucatan, 25 fms.
(Bathoxiphus) ensiculus Jeffreys, Ann. and Mag. Nat. Hist. ser, AVEO P54 le i7; iroc. Zool. Soc.,) p. 660) pli 49; £. 4, 1882. Watson, Challenger Rept., (Scaphopoda), p. 12, pl. Pee oos Wally Bull! 37) U. SN.) My, pi 76;).1889.- Plenaerson bul iam NO SiN. Me pl Say ply) Tai) ie) iail5) PGi TOZO:
9g BULLETIN 38 39
Distribution.—EHast and West Atlantic. A deep.water species of wide range. Gulf of Mexico: Yucatan Strait, 640 fms.
(Bathoxiphus) didymum Watson, Jour. Linn. Soc. London, 14, p. 517, 1879; Henderson, Bull. 111, U. S. NM pe eee inion e20)
Distribution.--Culebra Isl. and Barbabos. Doubtful specimen, Yucatan Banks, 25 fms.
(Compressidens) pressum Pilsbry and Sharp, Tryon’s Man. Conch. 17, p. 124, pl. 7, f. 11; pl. 22, f. 50-52, 16GQ7. lemdeream, Bull, 111, U. S. N. M., p. 83, pl: 14, 1. ao eee
compressum Watson, Jour. Linn. Soc. London, 14, p. 516, 1879; Dall, Bull: M. C. Z., 9, p. 38, 1881; Watson) Challuikenr (Scaph.), p..9, pl. 1, f. 9, 1885; Dall, Bull 27." ssih ea p. 76, 1889. Not compressum of d’Orbigny, 1850.
Distribution.—Florida to the Antilles. Gulf of Mexico: Off Cedar Keys, 196 fms.; between Mississippi delta and Cedar Keys, Fla:
(Compressidens) ophiodon Dall, Bull. M. C. Z., 9, p. 38, 1881; 18, p, 427, pl. 26, £..9, 1889; Bull..27,"U. S. INU eee 26, f..9, 1889 ; Henderson, Bull! 111, U. Sv See LAE, 12) WTO2ZO:
Distribution.—Florida to Barbados. Gulf of Mexico: West Florida, 169 fms.
quadrangulare Harris, Bull. Amer. Paleontology, vol. 1, p. 95, 1895. Upper Miocene, Galveston well at 2465-2871 feet (Harris).
tetragonum Sowerby, Thesaurus Conch., 3, p. 103, pl. 224, f. 21, 22, 1860. Upper Miocene, Galveston well, at 2158- 2920 feet (Harris).
Remark.—A fragment of an undetermined, Pleistocene, Dez- talium was obtained from Knapp’s No. 2 well, Terrebonne Parish, La. at 1190-1430 feet.
40 MoLLUSCA OF THE GULF OF MEXICO IO
Genus CADULUS Philippi
(Polyschides) quadridentatus Dall, Bull. M. C. Z. 9, p. 36, 1881; rompeazeupliia7 if 5. 1eeoq; Bully 37) Ue S. NY) Me ps) 75; piven 5) 1830; lenderson, Ball, vir, U. SiN. M:, pp..97- TOO) pl 17, £2, 3,7 1920.
Massieusia. Ir Conn: Acad), 6, p. 471;pl. 45, 1.'20, 1885; Baum alan. 5. N/M, p. 76, pls 41,.t. 20, 1889. Distribution.—Hatteras to Rio de la Plata, 3-52, fms. Gulf
of Mexico: West Florida, 10-25 fms.; Yucatan Banks 25 fms.
(Polyschides) carolinensis Bush. In Verrill, Rept. Fish Com. (for Mose WP S87, 0oss- Lr, Conn, Acad... 6,) p.i471,° pl. 45, f. iomioon Wal Bull 37.00. Sy NM, p48) (pars)) pl. 41, f. 19, 1889; Pilsbry and Sharp, Tryon’s Man. Conch., 17, p. meal 25. fh 64, pl, 30; 1. 241-127, 1898; Henderson, Bull. pie Oo NE.) ps 102) pli p17, £:/6,''7, 1920: Distribution.—Hatteras to Key West, 3-87 fms. Gulf of Mexico, off Mobile, in 27 fms.
(Platyschides) poculum Dall, Bull., M. C. Z,, 18, p. 429, 1889; Blo Wow ING Me ps 76, 1889; Pilsbry and. Sharp, Muyonrs Nan. Conch). 17) p.i 172, pli 33) fe 565-/ 57, 1898. : Bleaderson,bull, rir, U.S. No M:, ps 108, pl. 17, f. 8,1920.
Distribution.— Type found off St. Vincent, W. I., 424 fms. Also dredged, 640 fms., Yucatan Banks.
(Platyschides) parvus Henderson, Bull. 111, U.S.N. M., p. 1173, Es fy 24 ToZO:
ameantus Dall, Bull. 37, U, S. N. M., p. 78, 1889. Net Dall’s
typical amzantus. Distribution.— Off Barbados, too fms. (type); Yucatan Banks, 1002 fms.
(Platyschides) elongatus Henderson, Bull. 111, U.S. N. M., p. B22 OV LO fet 5. 1O20,
It BULLETIN 38 41
Distribution.—Gulf of Mexico, off the mouth of the Missis- sippi River, 68 fms.
(Platyschides) arctus Henderson, Bull. 111, U. S. N. M., p. 124, MO £4 TOM TOZOs | Distribution.—Gulf of Mexico off Cape San Blas, Florida, 60 fms.
(Gadila) mayori Henderson, Bull. 111, U. S. N. M., p. 133, pl. LO 2, a O26.
Distribution.—Florida, Gulf of Mexico, off Cape San. Blas, Fla., 60 fms.
(Cadulus) transitorius Henderson, Bull. m1, p. 143, pl. Io, f. 6, 1920.
Distribution.—Florida to Old Providence Isl.,north of Colon. Gulf of Mexico, off Cape San Blas, Fla., 196 and 169 fms.
42 MOLLUSCA OF THE GULF OF MEXICO 12
aS) aA Sine ODA ORDER PTEROPODA
Genus CAVOLINA Abildgaard milexa Lesueur Nouv. Bull. Soc. Philom., 3, p. 285, pl. 5, f. 3, house wast/7yalea), Dall, Bully U. 5. Nat. Mus,, 37, p. 82, ooo noc, WU, 5. Nat. Mus:, 37, p. 197, 1910, Distribution.—West Atlantic, Lat. 42°N.—a2° S., East Pacific, Lat. 13° N.—42° 8. Gulf of Mexico, West Florida. Pelagic. Pleistocene to Recent.
uncinaia Rang, in d’Orbigny’s, Voy. Ame’r Me’r., p. 93, pl. 5, ion. te36, (as Afyalea); Wall Bull.37,U. S. N. Maus., Poel oo. L.) Ol. 18807 Proc.-U. 3. N. Mus... 27) p..198, feLe, eunson, Occ. Papers, Boston ‘Soc. Nat. Hist., 7,.p. 156, 1915.
Distribution.—West Atlantic, Lat. 40°N.-40°S. East Pacific, Lower California to Ecuador and Galapagos Isls. Gulf of Mexico; West Florida. Pelagic. Pleistocene to Recent.
(Diacria) trispinosa Lesueur, in Blainville; Dict. des Sciences Nat. g2me2, (1327) \(as ffyalea): Gould, Inv. Mass., p.. 504, 1870, (as Diacria): Dalle srs 10,182 pl), CO. fyi Ss, 188o: puniner, (Bull Bur, Hish., 21, pt. 2, p: 703, I913., Johnson, Dee Pap. Bost. Soc., 7, pi 157, 1015:
Distribution.— West Atlantic, Lat. 60°N.—40° S. Gulf of Mexico, West Florida. Pelagic. Pleistocene to Recent.
Genus CLIO Linnzeus
pyramidata Linnzeus, Systema Naturz, 12th ed., p. 1094, 1767; Date Bull. 27.400 5. IN. Mi, p. 80, 1889, (as Cleodora): men CLS. NM. 27, p. 198, 1910; Johnson, Occ. Pap: Bost. S0C-; 7, Pi-158, 1915.
Cleodora lanceolata Souleyet, Zool. Bonite, 2, p. 179, pl. 6, f 7-25, 1852. Distrib ution.—Atlantic, Spitzbergen to Lat. 40° S. South-
13 BULLETIN 38 43
eastern Pacific, Lat. 27° S. Gulf of Mexico, West Florida and Texas. Pelagic. Pleistocene to Recent.
recurva Children, Jour. Roy. Inst.,\15, p. 220, pl.7, 1.na7aeee) (as Balantium); Dall, Bull. 37, p. 82, 1889; Johnson, Occ. Pap. Bost. S0C.37,/p. 15a Trg,
Distribution.—Lat. 40° N.—33° S. Gulf of Mexico, West Florida. Pleistocene to Recent.
Genus CRESEIS (Rang) Sowerby
subula Quoy and Gaimard, Ann. des Sci. Nat., ser. 1, 10, p. 233, pl. 8 D, f. 2-3, 1827, (as Cleodora):;: Dall, Bal. waa aac: 1889; Dall and Simpson, Bull: U.S. Fish Com. for to00))7, p. 360, pl. 57, f. 1, 1901; Dall, Proc. U. S)\ Nei aa 1910; Johnson, Occ. Pap. Bost. Soc.,7,. puya5e meus icas Styliola); Dall, Bull. 112, U. S. Nat. Mus), (3) 50)ream
Distribution.—Atlantic, Lat. 41° N.—4o0° S. Type locality, Teneriffe. Pacific, Peruvian. Gulf of Mexico, West Florida and Texas. Pelagic. Pleistocene to Recent.
Genus STYLIOLA (Lesueur) Gray
virgula Rang, Ann. Sci. Nat., ser. 1,13, p. 316,072) Guea@en nas Creseis): Verrill, Trans. Conn. Acad., 5, .)557; sbeaaeun as Stvyliola); Dall, Bull. 37, U. S. N. Mus.) (piusain gaias Cvesets); Proc: U. S. N.M., 37, p. 198, 19o1ON(ae uae ae): Johnson, Occ. Pap. Bost. Soc., 7, p. 157, 1915, (as Cresezs).
Distribution.— Atlantic, Lat. 41°N. - 35° S. Pacific, off Juan Fernandez Isl. Gulf of Mexico, West Florida, Pelagic. Pleistocene to Recent.
Genus VAGINELLA Daudin chipolana Dall, Trans. Wagner. Inst. Sci.,3, pt. 2, p. 431, pl. 23, f- 4,5, 1802,
Lower Miocene, Chipola beds, Bailey’s Ferry, Calhoun Co., Florida. An ancient, Cretaceous to Recent, genus.
A4 MOLLUSCA OF THE GULF OF MEXICO 14
Genus CUViERINA Boas
columnella Rang, Ann. des Sci. Nat., ser. 1, v. 12, p. 323, 1827, Atlas 4°, pl. 45 B, f. 1-8. (as Cuvieria); Pelseneer, Challen- Pere veropoda, Zool. ,23,art. 1, p. 67, 1888; Dall, Bull, 27, U. Sewers... 82, pl: 66, f..117, 7°89; Tr.) Wagner Inst. Sete Pt, 2)'p) 431,'. 1892; Peck, Proc..U..S. N. Mus., 16, Peo2 wok hs5.t. 16, F892: Dall, ‘Proc. U.S. IN. M., 37, p: ies roro:) Folunson, Occ.) Pap. Bost. Soc.; 7; Pp: 159, 1915.
Tripiera columnella Verrill, ‘Trans.. Conn. Acad., 5, p. 557, £63216) pl 214, 1884. Distribution.—West Atlantic, Lat. 43° N.—4o0° S. East Pacific, Lat. 23° N.—42° S. Gulf of Mexico, West Florida
and Texas, recent. Lower Miocene Chipola bed at Alum Bluff, Calhoun Co. Florida.
Note.—The specific name is often, but erroneously, written columella. ‘The correct form is columnella since Rang prints it thus twice in the original description. The generic name Cuvierta is preoccupied.
Genus EMBOLUS Jeffreys
inflatus d’Orbigny, Voy. Ame’r. Me’r., p. 174, pl. 12, f. 16-19, pogou(as Avania); Dall; Bull:'.37, U. S..N. Mus., p. 80, nace, etoc. U.S: Ne Mus., 27, p..199, 1910; Johnson, Occ. A DOSt. SOC... 7, DP.) 155, 1915, (as Lzmacimne ).
Spirialis rostralis Verrill, Trans, Conn. Acad., 6, p. 431, 1885.
Distribution.—West Atlantic, Lat. 42° N.—40°S. East Pacific. Lat. 42° N.— 40° S. Gulf of Mexico, West Florida. Pelagic, Pleistocene to Recent.
Genus PERACLE Forbes
reticulata d’Orbigny, Voy. Amer, Me'r., p. 178, pl. 12, f. 32- Be. 1530, (as datlania). Wall, Bull) 37, U.S) N. Mus., feveo, 1889, Proc,\U. 5, N. Mus., 37,p- 199, 1910:
15 BULLETIN .28 45
Distribution.— West ‘Atlantic, Lat. 37°. Nia) Game et Pacific, Lat. 20° S. Mediterranean. Gulf of Mexico, West Florida. Pelagic. Pleistocene to Recent.
ORDER OPISTHOBRANCHIATA Genus ACTAESN Montfort
punctostriatus C. B. Adams, Jour. Bost. Soc. Nat. Hist., 3, p. 323, pl. 3, f. 9, 1840, (as Tornatella): Dall, Bilis ies: N. Mus., p. 84, pl: 41, f. 17, pl. 82, f. 22,786. oy ee Inst. Sci., 3. p. 14, ’90; Singley, 4th Anum Rept,/Geelisury: Texas, p. 332, ’92; Pilsbry, Tryon’s Maal @omemiecequm 15, p. 157, pl. 18, f. 98, 99, pl.-19, £. 22) 26.) 93 ene aad Simpson, Bull. U. S. Fish Com., 1, p. 362, 1901; Johnson, Occ. Papers, Bost..Soc. N. H.>:7, p. 947 emoms
punctatus d’Orbigny, In La Sagra, Hist. Pol) y Natiisiade Cuba, 1, p. 230, Atlas, pl. 17, f. 10-12, 1842; (as Pormen72 Dall, Proc. U. S. N. M., 6, p. 324, 1883. INGE Piipaeeare Ferussac, 1823. Not of Lea, 1833. Not of Piette, 1855.
cubensis Gabb, Tr. Amer. Phil. Soc., 15;'p) 245,meqe
Note.—The recent Cuban Actcon punctatus d@ Ormeny (preoccupied, renamed cubensis by Gabb) is now considered identical with the northern punctostriatus. Acteon pune- tatus Lea (Contr. Geol, p. 111, pl. 4, f. oG)aisn ainenge Eocene, Ala.; and, “de Dall, Pliocene; Caloosahatchie beds, Fla. Not Recent.
Distribution.—Cape Cod to the Antilles, 2-63 fms. Miocene (of Shiloh, New Jersey) to Recent. Gulf coast: Cedar Keys, Fla,, Corpus Christi Bay, Tex. Pliocene, Caloosahatchie beds, Fla.
exilis Jeffreys, Ann. & Mag. Nat. Hist., 4th, ser., 6, p. 85, 1870; Watson, Chall. Rept. Gastr., p. 624, ’86; Dall, Bull. M. C. Z. Harv. Coll. 18; p. 39);" 89; Bull. 37, U.S) ae
46 MoOLLUSCA OF THE GULF OF MEXICO 16
959, Pilsbry, Man. Conch.,1 ser., 15,p. 156, pl. 19, f. 4-6,’93.
Puueeniina tesculpia Verrill;' Proc. U. S. N, M:, 3, p. 381, 1880.
gees Werrill, Tr. Conn. Acad:, 5, p. 540, pl. 58, f.. 21, 1882.
Distribution.— West Atlantic, Martha’s Vineyard, Mass. to Florida. Also Mediterranean and Bay of Biscay. 92-1465 nose Cult of Mexico, Campeche Bank, at 200 fms. Pliocene to Recent.
pusillus Forbes, Rept. Brit. Assoc. Adv. Sci., p. 19I, 1843, (as Tornatella); Jeffreys, Ann. & Mag. N. H.,4th ser., 6, p. 84, 5th ser. 10, p, 34; Watson, Chall. Rept. Gastr. p. 627; Dall, Bull 37, U. S. N. M:, pr 84, 1889; Pilsbry, Man. Conch.., Msers, Ws) Dp. 150-1303:
An unfigured species.
Distribution.— Mediterranean, Madeira; West Indies. 20- 450 fms. Gulf of Mexico, off Sand Key, West Florida, at no fs,
perforatus Dall, Bull. M. C. Z., 9, p. 96, 1881; 18, p. 42, pl. 18, eenoo Eula W27S. ON. M,. pe 84; pl. 18, f. 3,789; Piicney. Man, Conch.,)t ser;°15,.p, 159, pl. 20, f. 36, 1893. Distribution.—Gulf of Mexico, 805 fms.
micisus Dall, MC. Z.\.9, p. 95, 1881; 18, p: 42, pl. 17,\f. 1, 1 8, merebull 275 Ut. Nooo, p: 84, pl. 17, £ 1,: 16, 789; Etisory, Man. (Coueh., 1 ser., 15, p. 161,.°92. Distribution.—Yucatan Strait, off Cape San Antonio, 640 fms. Also off Fernandina, N. EF. Fla., 294 fms.
@elicatus Dall. Bull. M. C. Z., 18, p. 41, pl. 17, f. 5, 1889: Proc, Bio Na Wi tah: 266,)'89; Bull. 37; U.S. N..M., p. 84, ple i7,t. 5,89: Pilsbry, Man. Conch.,; ser. 1, 15, p. 162, pl. BOM I? 5615 O32:
Distribution.—Gulf of Mexico, West Florida, region. Also Barbabos 73-400 fms.
17 BULLETIN 38 47
galvestonensis Harris, Bull. Amer. Pal., vol. 1, p. 96, pl. 3, f. 1, 1895.
Distribution—Upper Miocene, Galveston well at 2552-2871 feet.
Genus RNIGICULA Deshayes
nitida Verrill, Am. Jour. Sci., 3, ser., 5, p. 16, 1873; Tr. Conn, Acad., 3, p. 48, pl. 1,f. 2, 774; Dall, Bull ee ee 43,89; Bull. 37, p. 84, pl. 37, f. 2;)" 805 -hilspiye eta Conch., 1 ser., 15, p. 399, pl. 46, f. 38, 962; Jommsom med, Ra. Bost. Sec: Ni G7 apa ouse
leptocheila Brugnone, Misc. Mal., p. 41, plot, Gaye: Morelet, Jour. de Conch., pp. ‘131, 285," jilewsy tums Agassiz, Three Cruises of the Blake, 2, p. 70, f. 291, 1888.
Distribution.—Massachussetts to Pernambuco, Brazil. Also Pliocene of Italy (Brugnone) and recent in the Mediter- ranean. I9-1075 fms. Gulf of Mexico, Yucatan Strait, 640 fms.
Genus ACTEOCINA Gray
bullata Kiener, Sp. et Icon. Coq. Viv., p. 5, pl. 1, f. 4, (as Tornatella);~ Dall, Bull. 37, U.S. N. M. “pieaipese: Pilsbry, Man. Conch., 1 ser., 15; p. 183, pl. 50,44 90) eu 22, f. 17-19, ’93;) Harris, Bull. Am. Pal. vol) ji;{poeones: Dall. & Simpson, Bull: U.S. Fish Com., 1, p goa igplaag, ENON hOOr
Bulla canaliculata d’Orbigny, Hist. Pol. y. Nat. Isla de Cuba, Tp. rea pliable. ay ema NOt Of Say.
Tornatina olivula A. Adams, Thes. Conch., 2, p. 569, pl. 121, ey. Distribution.—Florida Strait to Trinidad Island. Upper Miocene, Galveston well at 2410-2425 ft. (Harris).
recta d’Orbigny, In Sagra’s Hist. Pol. y. Nat. Isla de Cuba, 1, p: 131, pl. 4, bis. f. 17-20) (as Bulla); Dall, Bulgin:
48
MOLLUSCA OF THE GULF OF MEXICO 18
IM p. 84; 1889; Pilsbry, Man.-Conch., 1 ser., 15, p: 184, pl. 22, f. 13-15, '93, (as Zornatina); Maury, Bull. Amer. ale INO) 29, p..r4,-pl--3, f. 3, 1917.
Distribution.—Florida Keys, Antilles and St. Helena. Gulf coast, Tampa. Also Lower Miocene Santo Domingo and Jamaica. A closely related, Older Pliocene, species is Acteocina wetherilli Lea, from Deal, N. J. and the Orbitolite bed, Tampa, Fla.
eanaliculata Say. Jour. Acad. Nat. Sci. Phila., 5, p. 211, 1822,
(as Volvaria); Holmes, Post. Pl. Foss. S. Car., p. 78, pl. 12, Ponca, (Gould, Binney’’ ed., p. 219, £. 510, 1870,-(Cas Waves) DalliProc, U.S: N. M.,.6, p. 324, 783; Bull. 37, ReroeeN NE Dn 4.0 ploy 52) f. 27).%80,; (as Lornatna); Singley, 4th Ann. Rept. Geol. Tex., p. 332, ’92; Pilsbry, Diam oneh.) (6) Ser, .L5,;p. "184, plii22) f..22; plu 50, f. 25,
or eecy pounson. ‘Occ. Pap, Bost.’s0c: N.) H.,''7, p. 147,
Hrs Mauny, bull: Am. Pal.,\ No.) 29, p..13, pl.'3, f..2, 1Or7,
obstricta Gould, Silliman’s Jour. Sci. 38, p. I96, I840, (as
mea iixdams, ! Thes). 9 2,..p.) (566, t. 121, f..\ 29, (as Tornatina).
Distribution.—Portland, Maine to Haiti, 0-63 fms. Lower Miocene (Santo Domingo) to Recent. Gulf coast.—Recent: Cameron and Point au Fer, La., (very abundant); Galves- ton and Corpus Christi, Tex. Pleistocene: North Creek, Manatee and Labelle, Fla.; Grand Chenier, La, Knapp’s wells, Terrebonne Parish, No. 2 at 1050-1790 ft., No. 3 at 258-1525 ft. Pliocene; Caloosahatchie River, Fla. Miocene; Gilbert well No. 10, Bateson, Tex., 323 ft.
candei d’Orbigny, Moll. Cuba, 1, p. 12%, pl. 4, f. 1-4, (as Bulla);
Bate bulla 7 Uy SN Mo: p. 84; “pho gy, -£.)..13) 1889; Wanatta Pr. AL NOS. Phila., 55,.p. 757, 1903.
Distribution.—Hatteras to Martinique, o-48 fms. Gulf
19g BULLETIN 38 49
coast, St. Joseph’s Bay and Crooked Isl., West Fla.
Genus RETUSA Brown
frieler Dall, Bull. M. C. Z., 9, p. tor, 1881, {as \C/e7remiee ye p. 47, pl. 17, f. 4, 89; Bull. 37, U. S34 Noy ieee eet f. 4,’89; Pilsbry, Man. Conch., 1 ser., 15, ps 20q)) pi eme 8, 11893. Note.—I follow Cossmann.in placing Ufriculus Brown, 1327 (not of Schumacher, 1817) im the syuonymiy or
Retusa.
Distribution.—Yucatan Strait, 640 fms.
Genus SCAPHANDER Montfort
punctostriatus Mighels, Pr. Bost. Soc. N. H., 1, p. 49, 1841, (as Gulta); Dall, Bull. 37, U.S. N. M., .p.186) ph ae se. Pilsbry, Man. Conch., 1. seri, 15; p. 246.001) oa) iouue eee Johnson, Occ. Pa. Bost. S..N: Hi, 7, pi a24¢pnon,.
fibrartus Love'n,Index Moll. Scand., p. 10, 1846.
Distribution.—Iceland & Norway to Bay of Biscay. Maine to Barbados, 46-1467 fms. Gulf of Mexico, Lat. 28° N., Lon. 88° W., 533 fms. Inhabits shallow water in its north- ern habitats.
watsoni Dali, Bull. M. C. Z., 9, p. 99, 1881; 18, @3aWoly Tas: 10; 89; Bull: 37,,U. S..N. M.; p. 86, pli 17, ele Distribution.—Hatteras to Barbados, 54-324 fms. Gulf of
Mexico, West Fla. region, dredged U.S. Fish Com., bot- tom temperature 46° F.
nobilis Verrill, Tr. Conn. Acad. Sci., 6, p. 200,Jjols a2. Dall, Bull. M. C. Z., 18; p: 53, 1889; Pilsbry,) Mam @ogene, L'SEn sy E55). 2405.03" Distribution.—Martha’s Vineyard 906-1309 fms., Delaware Bay 1091-1209 fms., Tobago 880 fms., Gulf of Mexico, U. S. Fish Com, Stationi2527 141.1030 fms.
50 MoLLUSCA OF THE GULF OF MEXICO 20
Genus SABATIA Bellardi
bathymophila Dall, Bull. M. C. Z., 9, p. 98, 1881, (as Atys?);18, Seer 17. £..97 9b, °89;\ Bull. 37;)'U: Se No M.)\p. 86; pl: Baie Oo, Ob,’ 89. Distribution.—Fernandina to Guadeloupe, 294-1568 fms. Gulf of Mexico, Yucatan Strait, 740 fms.
Genus CYLICHNELLA Gabb
bidentata d’Orbigny, Moll. Cuba 1. p. 125, pl. 4, f. 13-16, 1841, Kaswbulla): Gabb, Pr, A. N: S. Phila., 24, p. 273, pl..10, £. age Wall Bull. 37) Ui 5. N. M.,\p.86; pl. 41; f) 14, 789; Manmattay Pr AL NOS. Phila., 55, p..757, 1903.
Buna biplicata Wea. Pr. Bost, Soc..N. H., 1, p. 204, 1844.
Distribution.—Hatteras to Barbados, 7-168 fms. Pliocene to Recent. Gulf coast, Recent: West Florida and Texas. Pleistocene, Louisiana, Knapp’s No. 2 (well, Terrebonne Parish, 1519-1542 ft., Crowley No.4 well, Jennings, 1663- 1670) feet.
bidentata var. galvestonensis Harris, Bull. Am. Pal., vol. 1, p. CON ey 3) fy 2\) 1895. Distribution.—Upper Miocene, Galveston well at 2600-2733
eet, Genus BULLARIA Rafinesque
Bitiata Bruguiere, Ency. Meth., 1, p. 572, (as Bulla): Dall,
familie WS. IN. Mi.) p.'88,, 1889; Pilsbry,;-Man. Conch... 1, Sates, 2224 pli a7 i! 42-46, )O3. Distribution.—Recent, Mediterranean and Atlantic coasts of Portugal and Morocco. Also living at Clearwater Harbor, West Florida; New Harbor Isls. and Chandeleurs, Louisi- ana; Texas. Pliocene, Caloosahatchie marls, Fla.
occidentalis A. Adams, In Sowerby’s Thes., 2, p. 577, pl. 123, f. Poon too, (as) ude): Dall, Pr. U.S. N2M.). 6, py 324, Peroull: 27, U0. S-.N.M., p. 88, °89; Singley: 4th Ann.
21 BULLETIN 38 51
Rept. Geol. Surv. Tex., p. 332, °92; Watattayirr yen: Pitas 555 1P: 757), £906. Distribution.—Florida to St. Vincent, W. I. Gulf coast: Cedar Keys, St. Marks, Tampa, Crooked Isl (Calhoun Co.), Fla.; Corpus Christi and Matagorda Bays, Texas.
solida Gmelin, Syst. Nat., 13, p. 3434, (as Bulla); Dall, Bull. 37, U. S. N. M., p. 88, 1889, Pilsbry;< Man @omemiian seu: 15 P1335) Dla 4a, ee eae. Distribution.—West Indies, Vera Cruz and Maracaibo. Cited from Texas region, Gulf of Mexico (Dall, ’89).
abyssicola Dall, Bull. M. C. Z., 9, p. 97, 1881, (as Bulla); 18, p. 56, pl. 17, f. 1, 7°89; Bull. 37, U.S. Ne Mop peep 11, ’89; Pilsbry, Man. Conch., 1 ser., 15, (p. 336,"Rlepeoe 31; 1893.
pinguiscula Jeffreys, Ann. & Mag. N.H., p. 318, 1880, (zomen nudum); Watson, Chall. Exp. Gastr., p. 638, 1886.
Distribution.—Bay of Biscay, Azores, Antilles and Yucatan Strait, 450-640 fms.
Genus HAMINOEA Turton
elegans Gray. Ann. of Philos. new ser. 9, p. 408; 1825; Index Testac., Suppl., pl. 3, f. 2, (as Audla); Pilsbry Man: Conch., 1, ser.,\15; p. 355, pl. 41, 1: 37-30;)qply ais: 1893; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 364, I9Ol. | guiding? Swainson, Malacol, pp. 251, 360, f. 46, 1840; Adams, Thes., p. 580, pl: 124, f.87-89; Dall, Bull. 37, Basen ei Pp. 88, 1889.
Note.—Ffide Pilsbry, the true elegans of Gray is this species and not a European shell.
Distribution.—Antilles to Rio de Janeiro. Gulf coast, West Florida and Texas.
52 MOLLUSCA OF THE GULF OF MEXICO 22
Succmea Conrad, Pr. A. N. S. Phila., 3, p. 26, pl. 1, f. 5, 1846, peewee) Dall, Pr,,U.)S. N.. M., 6, p. 324, 1883; Bull. A NM cp. 88, 789; Mitchell; List Tex. Sh.; Faleoiy Man, Conch,, i Ser., 15; p: 357; pl. 48, f..18, 1893; Dalland Simpson, Bull. U. S. Fish Com:, 1, p. 364, I901. Distribution.—Indian River, E. Fla., to Texas. West Florida at Tampa (Conrad’s type locality), and Cedar Keys; Chandeleurs, La.
antillarum d’Orbigny, Moll. Cuba 1, p. 124, pl. 4, f. 9-12; Mesrem Nal: Bl.; 22;\p. 175; Dall; Bull. 37, U.S. N. M., py ooeacso;| Pilsbry, Man. Conch., 1 ser., 15, p. 358, pl: 41, f. 35, 36, 1893; Mitchell, List Texas Shells.
cerina Menke, Zeitschr. fur Mal., p. 142, 1853.
Distribution.—W. Florida to Porto Rico and St. Thomas. Gulf coast at Tampa, Fla., and Espiritu Santo Bay, Texas.
antillarum var. guadalupensis Sowerby, Conch. Icon., f. 14, 1868; Pilsbry; Man. (Conch., 1, ser.) 15,' p. 358, pl. 41, f. 30-34, 1893.
Distribution.—Guadeloupe, Cuba &c., Gulf coast at White Water Bay, West Florida.
petiti d’Orbigny, Moll. Cuba 1, p. 130, pl. 4, bis. f. 13-16, (as malar Wall Bully 27, U. S: N. M., p. 88, 1889: Pilsbry, Mater Conch... i Set.,0,5, p..359, pl..41, f. 23, 24).1892. Distribution.—St. Thomas, W. I. and Tampa, West Fla.
virescens Sowerby, Genera, No. 39, f. 2; Dall, Tr. W.I.S., 3, p us, 1800; Pilsbry, Man, Conch., 1 ser., 15; p. 360, pl. 40, top Pl. 43, f. 10, 1893.
Distribution.—Recent on the Pacific coast, California to
Mexico. Pliocene, Caloosahatchie beds, Fla. Not in the recent Atlantic fauna.
Genus CYLINDROBULLA Fischer beaui Fischer, Jour. de Conch., p. 275, pl. 8, f. 8, 9, 1856; Dall,
23 BULLETIN 38 53
Bull. 37, U. S. N M., p. 88, 1889; Pilsbry, Man. Conch., 1 Ser.,05; p.' 380, pl: 42° te) 2e-msaer
Distribution.—Guadeloupe (type locality), St. Thomas, W. I.; Gulf coast at Cedar Keys, 2-95 fms. ‘This species is the genotype.
Genus HYDATINA Schumacher
physis Linnzeus, Syst. Nat., ed. x., p. 727, 1758, (as Bulla): Dall, Bull. 37, U. S. N. M., \p..88, 1880; Pilsbry tant Conch., ser. 1, 15, p. 387, pl. 45, £. 14-07, seas) Marten Simpson, Bull. U. S. Fish Com., 1; p. 364)°1g0x:
Distribution.—Atlantic, Pacific and Indian ‘Oceans. Gulf coast: Sarasota, West. Fla.
Genus PHILINE Ascanius
flexuosa M. Sars, Nyt. Mag. f. naturvidens., 17, p. 181, pl. 11, f. 23-26; Christianiafjordens Fauna, pp. 69, 70, pl. 11, f. 23- 26; Dall, Bull. M. C. Z., 18, p. 59,1889; Balla ee M., p. 88, 1889; Pilsbry, Man. Conch,, 1 ser) 10, p2 ae 4, f, 86-89, 1896.
Distribution.—Christiania, Norway; Yucatan Strait, 640 fms.
Genus TETHYS Linnzeus
protea Rang, Hist. Nat. des Aplysiens, p. 56, pl. 10, f. 1-3,1828, (as Aplysta): d’ Orbigny, Moll. Cuba, 1, p. 117; Beau, Cat. Coq.’ Guadeloupe, p. 20; Dall, Pr. U.S. eee ’°83; Pilsbry, Man. Conch., 1 ser. 16, p. 78, ploig7,i- 0-22: 1895; Dall & Simpson, Bull. U. S. Fish Com. 1, p. 365, I9OI.
Distribution.—Florida to Carthagena, Columbia. Very abundant in the Antilles. Gulf of Mexico, West Florida (Dall).
willcoxi Heilprin, Pr. A. N. S. Phila., p. 364, 1886, (as Aplysia) ; Dall, Bull, 37, U.S. N,' M., \p. 90, ’80; Pileieny aaa:
54 MOLLUSCA OF THE GULF OF MEXICO 24
Women.) I Ser, 16, p..80, pl. 35, f. 30-32,.1895; Johnson, Occ. Bape Bost. Soc, N. H., 7, p..154, 1915.
Distribution.—Little Gasparilla Bay and Marco West Florida. Also Massachussetts coast.
ORDER PULMONATA
Genus AURICULA Lamarck
(Auriculastrum) pellucens, Menke, Syn. Meth. Moll., p. 131, 1830; ates We Su No Mo 8.) po. 275, pl. 18) £..8,1885;' Bull, Be on NM. ps) oo. pl. a7, £.:8y 18389.
Distribution.—Cedar Keys, Florida, to Demerara, British Guiana. Genus PEDIPES Scopoli
mirabilis Muhlfeld, Mag. Ges. Nat. Freude, Berlin, 8, p. 8, pl. Towra a, b 1818: Dall, Bull, 37, U. SiN: M,) p62, pl. 47, 1.17, 1889: Dall & Simpson, Bull. U.S. Fish Com., I,
We goe, plei'53) £.' 8) TOOL.
guadridens Pfeiffer, 1839: ovadis Adams, 1849, fvidens Pfeiffer, 1854.
naticoides Stearns, Pr. Bost. S. N. H. 13, p. 108, 1869; Dall, Ee Oo) NN 8a pe 279) \ pl. 18, F.°E7, 183 5%
Distribution.—Tampa, Florida,to Guadeloupe Island.
elongatus Dall, Pr. U. S. N. M.,8, p. 279, pl. 18, f. 4, 1885; Bull. Banos NG ME pe O2,, pl.ia7, f..4; 1889:
Distribution.—Marco, Southwest Florida. Genus MELAMPUS Montfort
coffeus Linnzeus,Syst. Nat. ed. x, p. 729, 1758,(as Bulla); Dall, Eos UN SN) May pl og, pli: 47. f.)\2, 18893 Wall) & Simpson, Bull:)U.'5 Fish Com. 1, p. 368, pl. 53, f..13, 1901.
contformis Bruguiere, Ency. Me’th. 1, p. 339, (as Bulzmus).
25 BULLETIN 38 55
Auricula ovula d’Orbigny, Moll. Cuba 1, p. 187, Atlas, pl. 13, i) 4-75 1853. Distribution.—Florida to French Guiana. Pleistocene to Recent. Gulf coast, Cedar Keys, Fla. and Texas.
coffeus var. gundiachi Pfeiffer, Vanatta, Pr. A. N.S. Phila., 55, P- 757, 1903. Distribution.—St. Andrews Bay, Washington Co., Fla. floridanus Shuttleworth, MSS., Pfeiffer, Mon. Aur. Viv., p. 36, 1856; Dall, Bull. 37, U. SN. M3, p: 92, .p19477 freee:
Distribution.—Tampa to Florida Keys.
flavus Gmelin, Syst. Nat., p. 3436, 1792, (as Voluta); Binney, Jour: Bost. Soc., Nat. Hist., 7, p. 166; 186247 ange Water Sh. pt. 2, p. 12,65; Dall, Pr. U. S. Nie eee pl. 18, f. 2, 785; Bull. 37, US. N. MJ spie2 ola 89; Dall & Simpson, Bull. U.S. Fish Com) 1.pygese. 54, f. 9, 1901; Mitchell, List Texas Shells.
Voluta flava Gmelin, Syst. Nat. p. 3436, 1792. Bulimus monile Bruguiere, Ency. Me’th., 1, p. 338, 1780.
torosa Moerch, Cat. VYoldi, 38, 1852.
coronatus Adams, 1849; corvonulus Adams, 1854. Distribution.—Florida to Guadelonpe. Gulf coast: Tampa, Cedar Keys, Fla.; Point au Fer, La.; Port’ Lawaea and Espiritu Santo, Tex. Note.—According to Mr. Mitchell, this species lives near shore, hiding in the day time under drift wood and sea weed and goes in search of food nocturnally.
lineatus Say. Jour. A. N.S. Phila., I ser. 2, p: 246, 1822; Dall,
Bull. 37,-U. S. Ny) Mi; 99.92) pl. 47, f.. 9; 12, Seamer 4th Ann. Geol. Rept. Texas, p. 352, ’92: Johnson, Occ. Pa. Bost: Soc. IN’ JH; 9, pgs.) tor 5.
brdentatus Say, Jour. A! N?)\S.)Phila., 1 ser., 2,’ aeeeee DeKay, N. Y:.,. Moll.,\ pi 57, pl. 5, £. 92) meng ies
56 MoLLUSCA OF THE GULF OF MEXICO 26
Binney, Land & Fr. W. Sh., 2, p. 10, ’65; Not dzdentatus Montagu, 1803, which is European, but introduced on New England coast.
beplicatus Pieiffer, Mon. Auric. Viv., p. 21.
Auricula cornea Deshayes, Ency. Me’th., 2, p. 90, 1830; jaumet Mittre’, 1841. Distribution.—Maine to Tortola Isl., Gulf coast: Tampa, Cedar Keys, St. Mark’s, Fla.; Point au Fer. Chaudeleurs, Belle Isle, La ; Galveston, Tex.
Note.—Northern specimens very distinct; southern grade towards flavus with which they may hybridize. Inhabits Salt marshes.
(Detracia) bulloides Montagu, Testacea Brit., p. 339, pl. 30, f. Amnsecancas. Volta); Dall, Pr: UliS.N. M., 3,.p..285, pl. er wens: Biull 37) Ui S. Ny M., p. 92, pl: 47,.4..7; 789.
Distribution—Cedar Keys, Fla. and the Antilles. Type of Gray’s subgenus Detracia.
(Sayella) hemphilli Dall, Pr. U.S. N. M., 6, p. 323, pl. 10, f. 6, nosawicas Jcuconia)* Pr). Ui-Si.N. M,. 8) p. 286, pl. 18, f. Bios bulk. WLS. Ni. Mow 37.) pi 92, pl 47, fp 41, 789% Singley, 4th Ann, Rept. Tex. Surv. p. 343; ’92.
Distribution.—Cedar Keys, Florida, and Corpus Christi, Texas. ‘Type of section Sayella Dall which is intermediate between Glauneria and Detracia.
(Sayella) crosseana Dall, Pr. U. S. N. M., 8, p.,286, pl. 18, f. 10, oy) bull 37) U. SiN Mop, 92, pl. 47,1. 10; ‘89; Singley, Arne Aun: Rept. lex.Surv., p. 343, 792.
Distribution.— West Florida at Egmont Key (Tampa Bay); Corpus Christi, Texas and the Bahamas.
Genus PLANORB1IS Guettard Gedis Wall) Proc..U. S. N. M., 46, p. 236-pl. 21, f. 3, 4, 1914:
27 BULLETIN 38 57
Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet. Brackish water formation.
Note.—The living species of this fresh water genus are not included in this Synopsis.
Genus TRALIA Gray
pusilla Gmelin, Syst. Nat. p. 3436, 1792, (as Volufa); Dall, Pr. U.S. N. M., 8, ps 276, pl. 18, £..5, (853 Bull e7 ee M., p. 92, pl. 47, f..5, > 89; Dall & Simpson, Balas: Kish ‘Com, 1, p.'360) pli.54. das room: triplicata Donovan, 1802; ovula Bruguiere, 1789; mztens Lamarck, 1822. Distribution.—Cedar Keys to Guadeloupe.
minuscula Dall, Bull. 37, U.S N. M., p. 92, 1889. Distribution.—Tampa to Bahamas.
(Alexia) myosotis Draparnaud, Tabl., p. 53, ’o1, (as Auvicula); Dall, Pr. U.S. N. M., 8, p. 277, °85; .Bull27; 0 See P02. Dk. 52,014 7G. 80:
denticulaia Montagu, ‘Test. Brit., p./234, pl. 20, au5oea as Voluta) Distribution.—Europe; Jamaica (introduced); West Florida, San Francisco (introduced).
Genus BLAUNERIA Shuttleworth
neteroclita Montagu, Test. Brit., Suppl., p. 469, ’08, (as Voluta); Arango, Moll, Cubana,\\p: 60, 778; Dall, Pro Wie aa 8p. 287, pli 17, £63,785; Buli. 37, U. S.No ee ae 47, f. 14, °89, Dall & Simpson, Bull. U. S. Fish Vom r a: 360, TQOL.
pellucida Pfeiffer,’40, cubenszs Pir.
Distribution.—Tampa and Marco, West Florida. Also Antillean.
58 MOLLUSCA OF THE GULF OF MEXICO 28
Genus SIPHONARIA Sowerby
alternata Say, Jour. A. N, S. Phila., 5, p. 215, ’26, (as Patella); iBiamey) Wand & FLW: Sh., 2, p. 153, f. 254, 765; Dall, Pr. im erING VN. 8) 9.) 287,85. (Bull. 37,0. 8) N. Mj) p.'92, 1889. Distribution.—The variety drunnea Hanley (Pr. Z. S., pp. 21,151,’58)is found at Sarasota Bay. West Fla.,and Bermuda. lineolata d’Orbigny, Moll. Cuba, 1. pl. 17, f. 13-15; Dall, Bull. Bye on IN Mp. 92,289. Distribution.—Fernandina, Fla., to Brazil and Texas region of the Gulf of Mexico. Genns WILLIAMIA Monterosato krebsi Moerch, Dall, Bull. 37, U.S. N. M., p. g2. ’89. Distribution.—West Florida to Barbados. Fenus VERONICELLA Blainville floridana Binney, Terr. Moll. U.S., 2, pl. 67, f. 17, ’51; Dall, Bik 27, U.S. N.-M., p. 90, 1880: Distribution.—Charlotte Harbor, West Florida to Florida Keys. Note.—Since Blainville erroneously attributed a shell to this
slug-like genus, Ferussac’s later name Vagznula is often used; but Veronzcella has priority.
ORDER CTENOBRANCHIATA
Genus TEREBRA Bruguiere
cinerea Born, Test. Mus. Vind., p. 267, pl. 10, f. 11, 12, 1780; even, Man), 7, p,\31, pl. 0; £.:67, 1885; Dall; Bull. M. C. Merion. 62.79, singley,, 4th Ann. Rept. Tex., p. 332, (oa) Dall S& Simpson, Bull: U. S. Fish Com., 1; p, 382, I9Ol.
salleana Deshayes, jamaicensis C. B. Adams.
29 BULLETIN 38 59
Distribution.—West Indies to Colombia. Gulf coast: Chandeleurs, La., Corpus Christi, Tex. Pleistocene, New Orleans Pumping Station, No. 7.
dislocata Say. Jour. A. N. S. Phila., 2, p. 236, 1825.) (as Cerithium); Hilgard, House of Rep., Ex. Doc. 1, pt. 2, p. 887, °78; Tryon, Man., 7, -p. 18, pl. 4, f)%6¢, 6575s: 4th Ann. Rept. Tex:, p. 332, ’92; Vanateay rie en ao: Phila. 1/55) P7576 oon. vudits Gray, petitit Kiener. Not petit? Maury, 1917.
Distribution.— Maryland to West Indies and Venezuela. Miocene to Recent. Gulf coast Recent; Cedar Keys, Fla.; Chandeleurs, La.; Galveston & Corpus Christi, Texas. Pleistocene of New Orleans pumping station No. 7; New Orleans artesian well of 1856; Lake Borgne borings; Knapp’s wells, Terrebonne Parish, No. I at 1600-1700, 2000-2150 ft., No. 2 at 1150-1290, 1731-1739, No. 3 at W150, We 25 acer. Also of North Creek, Manatee and Labelle, West Fla. Up- per Miocene, Galveston well, 2158-2920 ft.
concava Say. Jour. A. N. S. Phila., 5, p. 207, 1827; Dall, Tr. W. Inst. Sci., 3, p, 24, 1890; Singley, 4th Ann. Rept. Tex., p. 332, 92; Harris, Bull. Am. ‘Pal., vol: 1, progam
Distribution.—N. Carolina, Georgia and Texas, at Galves- ton . Pleistocene of North Creek, Fla. Pliocene, Caloosahat- chie, Fla. Upper Miocene, Galveston well at 2552-2920 ft. Note.—The variety vinosa Dall (Bull. M. C. Z., £8, p. 64, °89) is found on the Gulf coast at Sarasota Bay and Char- lotte Harbor, West Fla.
protexia Conrad, Pr. A. N. S. Phila., 3, p. 26) 99eaei"(as Cerithium); Tryon, Man. 7, p. 25, pl. 6, 1) G8): aee5- gall, Bull, 37, U. S. N. M.,- p.'94, °89; Tr. Wagner times: 25,90; Dall & Simpson, Bull, Fish Com. 1, p. 382, 1901; Vaughan, Publ. No. 133, Carn. Iust., p. 171, 1910; Maury, Bull. Am. Palii2q)p.2ey plat: Thon 7.
60 MOLLUSCA OF THE GULF OF MEXICO 30
Distribution.—Hatteras to Texas 2-50 fms. Lower Miocene to Recent. Gulf coast: Cedar Keys & St. Marks, F'la.; Cameron, La.; Galveston, Tex. Pleistocene, North Creek, Manatee and Labelle, Fla.. The variety Jdutescens E. A. Smith, ranges from N. Carolina to West Florida. It is more distinctly reticulated than the typical form.
fassila Dall, Bull. M. C. Z., 18, p. 66, pl. 36, f. 8, 1889; Dall ea simpson, Bull, U. S. Kish Com.; 1, p. 383, root.
Distribution.— Martinique. Also Gulf of Mexico, Lat. 23° N., Lon. 88° W., at 95 fms.; Yucatan Strait, 640 fms. limatula Dall, Bnll. M. C. Z.,18, p. 66, 1889; Bull. 37, U.S. N. M., p. 94, ’89. Distribution.—N. Carolina to Barbados. Also Gulf of Mex- ico between Mississippi delta and Cedar Keys at 111 fms. texana Dall, Nautilus, 12, p. 45, Aug., 1898; Pr. U. S. N. M., 24 Ps 502, pl.20, f..8, 1902. Distribution.—Matagorda Island, Texas. Collected by Hon. J. D. Mitchell. Very rare. galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 98, pl. 3, f. 5, 1895. Distribution.—Upper Miocene, Galveston well at 2387-2871 IgIO. , calhounensis Maury, Bull. Am. Pal., No. 21, p. 4, pl. 1, f. 3, IgIO.
Distribution.—Lower Miocene, Chipola marl, Bailey’s Ferry, Fla., and Bascom No. 2 well Mobile, Alabama, at a depth of 1241 feet, Chipola horizon.
indenta Conrad, Cited by Mr. Aldrich from the Bascom No. 1 well, Mobile, Ala. at 1500-1556 feet.
langdoni Dall, Pr. U. S. N. M., 18, p. 39, 1895; Tr. W. Inst., 3, pl. 59, f. 27, 1903.
ay BULLETIN 38 61
Distribution.—Lower Miocene, Chipola marl, Fla.; Upper Miocene Galveston well at 2158-2920 feet (Harris).
Genus CONUS Linnzus
proteus Hwass, Enc. Meth. vers, I, pt. 2, p. 682, 1789; Reeve, Conch. Icon.,f. 219, 4; Dall, Bull. 37, U.S. Neier 1889; Vaughan, Publ. No: 133, ‘Carn! instep mene, Maury, Bull. Am. Pal. No. 20, p. 42, plG ft: mm) sen
leontnus Wwass, 1791, spurius auctores, as of Gmelin.
Distribution.—West coast Florida, 19 fms. South to Vene- zuela. Pliocene, Caloosahatchie River, and Pleistocene, Labelle, Fla.
centurio Born, Mus. Vind., pl. 7, f. 10, 1780; Tryon, Man., 6, p. 33, f. 9, f. 68,°1884: Dall, Bull. M.. GC. Z1e eee ae:
Distribution.—Antilles. Also Gulf of Mexico between Mississippi delta and Cedar Keys at 25 fms.
delessertianus Recluz, Mag. de Zool., pl. 72, 1843; Tryon, Man., 6, p. 33, pl. 9, f. 67, °84; Dall (as delesserz7)) tai 27, USS eNe Me p94, OQ.
Distribution.—Hatteras to West Florida, 22-63 fms. Doubt- ful specimen, Porto Rico.
flavescens Gray, Sowerby, Conch. IIl., f. 68; Tryon, Man., 6, p. 36, pl. 10, £:\84, 1884; Dall, Bull. 37, U.S ee oe 1889.
Distribution.— Hatteras to Barbados, and West Florida, 15- 170 fms. Pliocene to Recent.
floridanus Gabb, Am. Jour. Conch., 4, p. 195, pl. 15, f. 4, 1868; Tryon, Man. 6, p. 38, pl. 11, f..4, 5, 1884;. Dall) Baliga, 94, 1889" Tr: Wi Iust. Seiei3))p, 27, 1890.
Distribution.—Hatteras to Florida Keys and West Fla. Pliocene of the Caloosahatchie beds.
62 MOLLUSCA OF THE GULF OF MEXICO 32
peali Green, Tr. Albany Inst.,1, p. 123,pl. 3, f. 3, 1830; Tryon, Mame, p36) pl. 10, f) 89-92, 18384; Dall, Bull. M. C. Z., 16).p. 68, 89. sear7ise Conrad, Aim. Jour. Conch., 5. p. 104, pl. 10, f. 1, 1869.
Distribution.—Hatteras to Isthmus of Panama. Also Cedar Keys, Florida, o-5 fms. Pliocene, Caloosahatchie beds. pygmaeus Reeve, Pr. Z. S., p. 179, 1843; Kiener, Coq. Viv. Conus, p. 174,pl. 111, f. 1 a-b,1848; Dall and Simpson, Bull.
Wes bish: Com, 1, p: 333, 1901. Distribution.—West Florida to TobagolIsl. Pliocene, Ca- loosahatchie beds. Pleistocene, North Creek, Fla.
amphiurgus Dall, Bull. M, C. Z., 18, p. 70, 1889. Distribution.—Coast of Yucatan, 27 fms.
chipolanus Dall, Pr. U. S. N.M., 18, p. 42, 1895. Distribution.—Lower Miocene, Chipola marls, Florida and
of the Bascom No. 2 well, Mobile, Alabama, at 1241 feet.
Genus TURRIS Bolten
albida Perry, Conch. Expl., pl. 32, f. 4, 1811, (as Pleurotoma); Darra sa Wes. NG Mp: 96, (895: Dall &' Simpson, Bally WU: S:-hish,Com., 1, p. 384, 1901; Maury, Bull. Am. Pal No.\29, p. 50, 1917.
Distribution.—Recent, Cedar Keys to Barbados, 26-100 fms. Oligocene of Yicksburg, Miss., and of Tampa, Fla.; Lower Miocene, Western Florida and Santo Domingo; Upper Mio- cene, Galveston well at 2158-2950 feet.
Note.—The variety ¢ellea Dall (Bull. M. C. Z., 18, p. 72, 1889) was dredged in the Gulf of Mexico between Missis- sippi delta and Cedar Keys. Genus LEUCOSYRINX Dall verrilli Dall, Bull, M. C. Z., 9, p. 57, 1881, (as Pleurotoma); 18,
23 BULLETIN 38 63
p..75, pl. ro, f. 5, 1889; Bull. 27, pli tefisyaoen
Distribution.—N. Carolina to Guadeloupe. Also Gulf of Mexico, Lat. 25° N., Lon. 84° W., 888 fms., and between Mississippi delta and Cedar Keys.
sigsbeei Dall, Bull. M. C. Z., 9, p. 57, 1881, (as Pleurotoma); 18, p. 76, pl. 11, f..-10, 789; Bull, 27, Ul Seine oe DEE, a LO, oes Distribution.—Yucatan Strait, 640 fms. Off Bequia, 1591 fms.
tenoceras Dall, Bull. M. C. Z.,—18; “p. 76, plo))36; 4 50eeeo Bull: 37,,.U. S: IN..Me) ps 06) pl 36) faa ee
Distribution.—N. Carolina to Guadeloupe. Gulf of Mexico between Mississippi delta and Cedar Keys at 724 fms.
subgrundifera Dall, In Agassiz’s Three Cruises of the Blake, 2, p. 66, f. 283, 1888, (as Pleuvetoma): Bult. C22 aren: 77, pl, 38, £. 1, °89;. Bull. 37, U. S. N. Ma poor mie $e rex Distribution.—Cape Fear, N. C., to St. Kitt’s, W. I. Gulf of Mexico, Yucatan Strait, 640 fms., and between Missis- sippi delta and Cedar Keys, 940 fms., U. S. Fish Com. St. 2384.
Genus ANCISTROSYRINX Dall
radiata Dall, Bull. M. C. Z., 18, p. 78, pl. 12, f. 12) 1880; M: 37, U. S. N. M., p. 96, pl. 12, f. 12, ’89; Dall & Simiesam: Bull. U2 S:; Fash Com:, 1, p. 284, 1901. Distribution. —Gulf of Mexico, Yucatan Strait,640 fms. and between Mississippi delta and Cedar Keys. Also Antillean.
Genus GENOTA Adams
mitrelia Dall, Bull. M.C. Z., 9, p. 56, Aug., 1881, (as Pleuro- toma); 18, p. 78, pl. 12, £.'5, 789; Bull. 37, Mesa. ies Di OOM Ol rein Sor
didyma Watson, 1881.
Distribution. Yucatan Strait, 640 fms. Also off Sombrero. Wit:
64 MoLLUSCA OF THE GULF OF MEXICO 34
Genus DRILLIA Gray
ostrearum Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Tryon, Miaime yp. 197, pl: 24, f..79; 784; Dall, Bull, 37, U.'S..N. Wien. (06, 789; Dall & Simpson, Bull. U. S. Fish Com., 1, Pp. 385, 1901. Distribution.—Hatteras to Cape Catouche, Yucatan and An- tilles. 15-170 fms. West Florida: Recent, Cedar Keys; Upper Oligocene, Tampa; Lower Miocene, Chipola beds; Pliocene, Caloosahatchie beds.
albicoma Dall, Bull. M. C. Z. 18, p. 83, pl. 10, f. 8, ’89; Bull. OS IN Mo) 27.0. 96, pl: 10). f. 8, 1889. Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W., 84 fms. Also off Barbados, and St. Thomas.
detecta Dall, Bull. M. C. Z., 9, p. 65, Aug., 1881: 18, p. 84, pl. nen meso: Ball, 27, p. 96, pl. 12, f. 11, ’89.
circumvoluta Watson, Jour. Linn. Soc., 15, p. 465, Nov. 1881;
Nept:' Chall, Gastr:, p: 357, pl.\21, f. 1, 1885: Distribution.—Gulf of Mexico, 339 fms. Also Culebra SY, a
alesidota var. macilenta Dall, Bull. M. C. Z., 18, p. 85, pl. 36, f. frsso.: Bull 37, Us S..N2 M», p: 96, pl. 36, £. 1, 780. Distribution.—Cape Fear, N. C. to Barbados. Gulf of Mexico between Mississippi delta and Cedar Keys, 111 fms. Typical form of species, off Hatteras.
eucosmia var. canna Dall, Bull, M. C. Z., 18, p, 86, 1889; Bull. Bons. ING Mi pr'96)." 80. Distribution.—Cape Lookout, N. C., to Grenada, W. I.
Also Gulf of Mexico, west of Florida, 50 fms.
leucomya Dall, Proc, U.S. N. M., 6, p. 328, pl. 10, f. 8, 1883;
Elina ran oG. pla 4s, it. 7, 1889; Ir. W. Imst.,))2)) ps, 36, 1890.
35 BULLETIN 38 65
Distribution.—Shores of Gulf of Mexico from Sarasota, West Fla., to Yucatan, 3-5 fms. Pliocene, Caloosahatchie beds, Fla.
albinodata Reeve, Zool. Proc., 6, 1846; Tryon, Man., 6, p. 197, pl. 14, f. 5, 1884; Dall, Bull. 37; U. S! Ni I pea ee Distribution.—Charlotte Harbor, West Fla., to Santo Domingo.
haliostrephis Dall, Bull. M. C. Z., 18, p. 86, pl. 13, f. 3, 1889; Bull. 37,.U.,85 Novos p96, pling iain es
Distribution.—Gulf of Mexico, West of Florida, 84 fms. acestra Dall, Bull. 37, U. S. N: M., p. 96, ple relmi ae, Distribution.— West Florida to Grenada 161-400 fms.
pharcida Dall, Bull. M. C. Z., 18, p, 88, pl. 12, f. 2, 1889; Bull. 87D OOyl ply ete 2 ese: exasperata Dall. Not of Reeve.
Distribution.—Gulf of Mexico, west of Florida, 229 fms. Also East Fla. to Barbados 150-1002 fms.
tristicha Dall, Bull. M. C. Z.. 18, p. 88, 1889; Bulli ay pwice, 1889. Distribution.—Three stations Gulf of Mexico between Mis- sissippi delta and Cedar Keys, 111-210 fms. ‘Temperature 52°-67°) Be
ebur Reeves, Pr. Zool. Soc., p. 116, 1845, (as Pleurotoma); Conch. Icon. pl. 30; f. 275, 945; Iryon, Mang oy eames: pl. 13, f. 56, ’84; Dall, Bull. 37, U. S. IN. M. -pitesiiee: Distribution.—Hatteras to Sombrero Isl. Also Gulf of Mexico, West of Florida, Blake Station 10, at 34 fms.
pagodula Dall, Bull. M. C. Z., 18, p. 90, pl. 13, f. 6, 1889; Bull. 27) D.O8) Pls) 13, fh. On oor Distribution.— Gulf of Mexico, West of Florida, 50 fms. Also Barbados.
66 MoLLUSCA OF THE GULF OF MEXICO 36
fea Mail Pr. U.'S.' N. M., 6, p. 328, pl. 10, f. 5, 1883; Bull, M. Gees) po) \OL, 803, Bull 37, Ul) S. N. 'M., p. 98) pl. 43, Pee OO).
Distribution.— Hatteras to Florida Keys, and West Florida at Cedar Keys, 3-15 fms. Note.—The variety carminura Dall (Bull. M. C. Z., 18, p. gI, 1889) was dredged in Gulf of Mexico, U. S. Fish Com. Station 2402 at 111 fms.
simpsoni Dall, Pr. Davenport Acad. Sci., 5, p. 54, 1887, (as Uveurorma): Bull: M. Cy Z., 68, p. or, 1889.
Distribution.— Hatteras to Tampa Bay, West Florida, 15- 18 fms.
lissotropis Dall, Bull. M.C. Z., 9, p. 58, Ang. 1881; 18, p. 9I, PUpiwnie 4, 1889; Bull 37,\p. 98; pl. 11, £.'3, 4; 789.
hypsela Watson, Jour. Linn, Soc., 15, p. 433, Oct. 1881; Chall. Ash On 240.) pl. 27,.\f) 4; 1835.
Distribution.—Gulf of Mexico, West of Florida at 220 fms. Also Antillean. Variety perfolita Dall (Tr. W. Inst., 3, p. 36, ’90), Caloosahatchie Pliocene.
dalli var. cestrota Dall, Bull. M. C. Z., 18, p. 92, 1889; Bull. 677-8 98, 89.
Distribution.—Between the Mississippi delta and Cedar Keys, at 196 fms. Typical dali Verrill, ranges from Martha’s Vineyard to Delaware Bay.
meeleata Dall, Bull. M.’C.'Z., 9, p.62, °81; 18, p. 92, pl. 11, f. H. LO8Q.
Distribution.—Gulf of Mexico, 229-339 fms. Also Antillean. Pleurotoma amblia Watson may be a synonym.
Genin) Wall, Bull. M.-C. Z., 9, p. 68, 1881; 18, p, 93, pl. 11, f.
eateoo:) Bull. 27D. 98.\ pl. 11, £2) 2,789.
Distribution.—Gulf of Mexico, West of Florida, 220-310 fms.
wn SJ
BULLETIN 38 67
havanensis Dall, Bull. M. C, Z., 9, p. 67, 1881; 18, p. 93, pl. 11, LAE MBG
Distribution.—VYucatan Strait, 640 fms. Also off Havana.
lithocolleta Watson, Jour. Linn. Soc., 15, p. 441, 1881; Chall. Rept. Gastr., p. 320, pl. 24, f..6, 1835 (as 7leu7arcn2): Dall, Bull. M. C. Z., 18, p. 95, pl. 11, 1, 6)) 780; aiee ep: 98 spl bl t.GWe eee
Distribution.—Hatteras to Guadeloupe,W. I. Also Gulf of Mexico between Mississippi delta and Cedar Keys, at 940 fms.
Genus CYMATOSYRINX Dall
centimata Dall, Bull. M. C. Z., 18, p. 95, pl. 36, f. 9, 1889; Bull. 39. US Se NOM. p98; pls 3865 taro. be"
Distribution.—Hatteras and Gulf of Mexico, Lat. 24° N., Lon. 84° W. at 1920 fms., and between Mississippi and Cedar Keys at 1181 fms.
moseri Dall, Bull. M. C. Z., 18, p. 97, pl..36, fi 2a) eso. Ba oy AN Me sO8.) Dl 26. seas ey
Distribution.—Cape Hatteras and Sarasota Bay, West Fla.
Genus PLEUROTOMELLA Verrill
packardi Verrill, Am. Jour. Sci., (3) 5, p. 15, 18733) {£<) @oums ACAG | 5. Parts, Pl agian. 9, pl. 57, themes
Distribution.—Gulf of Maineto Rhode Island. ‘The variety benedictz V. & S., (Wall, Bull. 37, U. S..NIAoi ees er: 14, f. 4, pl. 60, f. 7o a, ’89) ranges from Gulf ef Maine to Bequia, 1290-1507 fms.Gulf of Mexico, West Florida region. leucomata Dall, Bull. M. C Z., 9, p. 63, 1881, (as Drillia); 18,
p: 120, pl. ri, f..13,°"8093. Bull. 37, p. 102, Goh a eemeee 1889.
68 MOLLUSCA OF THE GULF OF MEXICO 38
Distribution.—Gulf of Mexico between Mississippi delta and Cedar Keys. Also Florida Strait. 533-940 fms.
agassizi Verrill and Smith, Am. Jour. Sci., 22, p. p. 394, 1880; pie eun Acad) 5. p.'454, pl. 57, f.'3, 3a, 82.
Distribution.—Rhode Island to Cape Fear, N. C. The wemlety mexicana Dall (Bull. 37, U. S. N:'M.; p. 104, pl. 11, f. 14, 1889) ranges from Western Florida and Yucatan re- gions of the Gulf to Martinique. 502-640 fms.
alifera Dall, Bull. M. C. Z., 9, p. 56, 1881, (as Bela); 18, p. 123, imme ion. SQy Bill. 27, +p. 104,.'p.: 12; £. 9, 89: Distribution.—Gulf of Mexico, West of Florida, 331 fis.
Megha Wall Bull, M.~C. Z., 18) p. 125, 1889; Bully 37, p. 104, 1889.
iewmowtons——Cape Kear, Ni C., 407 fms. Also’ Gulf of Mexico between Mississippi deita and Cedar Keys, at 1181 fms.
extensa Dall, Bull. M. C. Z., 9, p. 55, Aug., 1881, (as var. of Mrcurovma: blakeana): 18, p-126, pl. 10, f. 2,80; Bull. 37, aTOAy pl, 10, f.°2,, 80.
Fleurotoma streptophora Watson, Jour. Linn. Soc., 15, p. 464, Novwisse Chall) Gastr.,p. 366, pl. 19, f. 8 a-b,, 1885.
Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 88° W., 804 fms.; Yucatan Strait 640 fms. Also North Atlantic.
blakeana Dall, Bull, M. C. Z., 9, p. 54, 1881; 18, p. 126, pl. 10, ple oOs) bill. 37. pmo, ) pl. 10, fy 1; *89:
Oxers Veriitl, Tr. Conn. Acad., 6, p. 417, pl. 44,.f. 8,785.
Distribution.—Gulf of Maine to Santa Cruz, W. I., 100-1608 fms. Also Yucatan Strait, 640 fms.; Florida Strait, 339 Pimas,
39 BULLETIN 38 69
Genus GLYPHOSTOMA Gabb
gabbi Dall, Bull. M..C, Z., 18, p. 108; pl/13) £4 sis Bull. 37, U.'S..N. M., p.: 100, pl. 13) f./ 4, ene
Distribution.—Gulf of Mexico, Blake Station 36, 84 fms.; U. S.F.C.Sta. 2405, west Florida, 30-50 fms. Also off Barbados.
gratula Dall, Bull. M. C. Z., 9, p. 64, Aug., 1881, (as Pleuro- toma); 18, p. 110; pl. 12, f. 16; °89; Bul 29a. ie@pene. HeeOs 2380: znctits Watson, Jour. Linn: Soc., 15, p. 425, Oct., 18815) Rept. Chall. Gastr., -p./304)"pl 22.f gi) 7 aie
Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys, 227 fms. Also East Florida and Antilles. Pliocene, Caloosahatchie beds.
Genus MANGILIA Risso
balteata Reeve, Conch., Icon., 3, pl., 8, f. 57, 1846; Tryon, Man. Conch., 6, p. 247, pl. 24, f. 1, 84; Dall) Bullia7 Se M., p. 100, 789; Dall & Simpson, Bull. U.S) Pichon, TSP 300) (LOO):
Distribution.—Hatteras to Barbados and West Florida. Pliocene to Recent.
astricta Reeve, Pr. Z. S. p. 46,’46; Tryon, Man., 6,p.260, pl. 24, f. 26, “84; Dall Bullii27, UiS. IN. M., poeta: Distribution.—West Florida to Florida Keys.
biconica C. B. Adams, Conch. Contr., p. 65; Dall Bull. 37, p. ILOLOM tsiow Distribution.— West Florida to Jamaica. Unfigured.
plicosa C. B. Adams, Contr. Conch., p. 54, ’50, (as Pleurotoma) ; Dall, Bull. 37, U. S.. NOM), p. 100, pl. 50, £:) cay Bos sims ly, 4th Ann. Rept, Tex. Surv., p. 333, ’92; Johnson, Oce. Pa. Bost; Soc. IN. Ee ps as O15.
Pleurotoma plicata C. B. Adams, Bost. Jour. N. H., 3, p. 318,
70 MOLLUSCA OF THE GULF OF MEXICO AO
Piya. 6,40, Not of Lamarck. plicatum Kurtz, 1860; bvunnea Perkins, ’69.
WOlathurgella jewettd Stearns, Pr..A. N.S. Phila., fcr 1873, p. pe wally Pros Si IN. M6, "p. 329, 1883.
Distribution.—Massachussetts to Florida Keys and Gulf coast at Cedar Keys, Fla.; Corpus Christi Bay, Texas, 0-5 fms. Pleistocene, North Creek, Fla.; Pliocene, Caloosa- hat chie beds, Fla.
rubella Kurtz and Stimpson, Pr. Bost, Soc. N. H., 4, p. 115,’51; A atts, 277.) W. OS.) NM.) p. Too, "894. Dr. Wagner Inst, Sel 3, Pp.) 40,° 90;
Distribution.—Cape Lookout, N. C., to Charlotte Harbor, West Fla. Pliocene, Caloosahatchie beds. Pleistocene, Knapp’s No. 2 well, Terrebonne Parish, La., at 1800 feet.
Sieilata Stearns, Pr. Bost. Soc., N. H., 15, p. 22,.’72;\ Dall, Pr. NOS ING NEG) p.).328)).°83: Tryon, Man: 6) p.'246, pl. 34, Pree Wall. Bulln'29): p.\ TOO). Sos Pe. Wir ls S.,.3, 'p) AT; OO. 3 Distribution.—Cedar Keys and Tampa to Key West, Fla., Pliocene, Caloosahatchie beds.
eenestvid) Dall, Bull. 37) U. S...N. M., p. 102, ‘pl. 4a, f..4,:42, 1889.
Distribution.—Hatteras to Barbados. Also West Florida, 14-333 fms.
l imonitella Dalle Pr Wisin M6) pi 320; pli fo, f/110; 1883: Be a7 Pp. TO2 ph iast 2,80,
Distribution.—Cedar Keysto Tampa, Fla., o-5 fms. Pleis- tocene, Knapp’s| No. 3 well, Terrebonne’ Parish, La., at 1150-1200 feet. Perhaps this species is a dwarfed form of atrostyla Dall.
cerina Kurtz & Stimpson, Pr. Bost. Soc.,4, p. 115,1851;(as Pleuro- toma); Hilgard, Rept., Chief Engineers to Sec., War., p.
Al BULLETIN 38 eo. AE
358, 1870; House of Rep., Ex. Doc. 1, pt. (2; giisea rer Dall, Pr. U. S. N. M., 6, p.°328,'83; Dryem) (ayia meee: 310, pl. 22, f. 43,84; Dall, Bull. 37, U. S No eee pl. 44, f. 16a; 89; Vanatta, Pr. A. N.S. Phila esses. 1903; Johnson, Bost. Soc. N. H., ‘Occ) (Pas aa: fOrS.
Distribution.—Massachusetts to Texas. Gulf coast: Cedar Keys, St. Joseph’s Bay, Crooked Island, Florida; Galveston and Corpus Christi, Texas. Pleistocene, New Orleans well of 1856; Lake Borgne borings; Grand Chenier; Knapp’s wells, Terrebonne Parish, No. 2 well at 1190-1439, No. 3 at 1500-1525 feet.
cerinella Dall, Bull. M. C. Z., 18, p. 113, 1889; Bull. 27, Uo, Ny. MD OZ aoe
Distribution.—Hatteras to West Florida and Galveston, 14- 22 fms. Pleistocene, Knapp’s No. 3 well, Terrebonne Parish, La., at 1500-1525 feet; North Creek, Fla.
quadrata Reeve, Pr. Zool. Soc., p. 114, 1845, (as Clathurella); Tryon, Man., 6, p. 278, pl, 18, f. 25, 784: Dalijipaieia: UW2'S AN Mei, 102, 80s Distribution.—Hatteras to Yucatan. Dall regards dimznuta C. B. Adams (Conch. Contr., p. 63) as a wanlety nangine from Hatterasto Florida Keys and West Fla.
melanitica Dall, Bull. 37, U. S. N. M., p. 102, 1889. Distribution.—Hatteras to Haiti and West Florida.
antonia Dall, Bull. M. C. Z., 9, p. 59, 1881; 18, p. 116, pl. Io, f. 4, pl. 1, f: 11, °89; Bull, 37, U. S. N. M jp. mee, Woe aa: 4, pl. lL Lt 8SO, Distribution.—Yucatan Strait, 640 fms. Also Guadeloupe.
peripla Dall, Bull. M. C. Z., 9, p. 68, Aug., 1881; (as Drilla) ; 18; p; 115, pl. 11, £. 17, 980; , Bull. 37, p. 102,/ (plea ee 1889.
see eased halal
74 MoLLUSCA OF THE GULF OF MEXICO 42
chyta Watson, Jour. Linn. Soc., 15, p. 466, Nov., 1881 (as Devancia); Rept. Chall. Gastr., p. 358, pl. 18, f. 4, 1885.
Distribution.—Yucatan Strait, 640 fms.
elusiva Dall, Bull. M. C. Z.,9, p. p. 69, Aug., 1881, (as Drillza); Memes, pl. 12, 1. 7, 896; Bull. 37, U.S. °N. M., p.-102, pita. f 7, ’8o. Distribution.—Yucatan Strait, 640 fms. Perhaps identical with Defrancia perpauxilla Watson.
bandella Dall, Bull. M. C. Z., 9, p. 59, 1881; Verrill, Tr. Conn. meda.,.0, p. 250,) 84; Dall, Bull. M..Cy Z., 18, p. 116, pl. rome SOc cbUll 27S, N.. M., p. 102, pl. 10, £:.3, pl: 60,71. 173; "80.
Pleurotomella diomedee Verrill and Smith, Tr. Conn. Acad.,
Gupte 2, pl. 31, f. 5, 58, 1884. Distribution.—East coast United States and Gulf of Mexico, Blake Station 47, at 321 fms, Greatest depth 2100 fms.
comatotropis Dall, Bull. M.C. Z., 9, p. 58, 1881; Verrill, Tr.
Cammyeueade: p, 452,782. Dall) Bull). M.C. Z. ) 18,:p.) 116, ieee ie. Sor Balle 37S. NM. p. 102, plvi1, ff. 12; pliaa. ff) Spl: 61, £77, '89; Johnson, Occ. Pa. Bost. Soc., Nee 7, Pp. 146; TOTS.
tiara Watson, Jour. Linn. Soc., p. 440, 1881; Chall. Gastr., PGA Dl 2T ete 7 V7 5:
Larants pulchella Verrill, ’82 and ’84. Distribution.—Massachussetts to Barbados, 50-1075 fms. Also Yucatan Strait, 640 fms.
Belagia Dall Bull..M.C. Z.. 9, p. 61, 1881; 18, p. 117, pl. 11, f. Peco oull 27) UES. Nui M;. p. 102; pl. 11, f. 9,7 89:
Distribution. —Gulf of Mexico, Blake Station 44, in 539 fms., bottom temperature 39° F. Also Antillean.
43 BULLETIN 38 73
exsculpta Watson, Jour. Linn. Soc., p. 247, 1882, (as Drdllia); Chall. Gastr., p..371, Pl. 24, f..2,:°85 (as\Gzometa ar: Bull. M. C. Z., 18, p. 127, pl. 15, £. 9, 803 BU ay ees Ol Te he Oy SOK Distribution.—Yucatan Strait, 640 fms. Also Santa Cruz, Weal
subsida Dall, Bull. M. C..Z., 9, p. 62, 1881;,18; p, 118, pitiwena: 3,89; Bull. 37, U.S. IN. M.,. p. 102) pill are ee
Distribution.—Gulf of Mexico, West of Florida, 339 fms. Also Cuba. ipara: Dall, Bull, (Ms °@) Zis%e) p57) 1881; LS) oe Ee ee 1889. Distribution.—Yucatan Strait, 640 fms. Also Martinique, 502 fms, Genus CYTHARELLA Monterosato
cymella Dall, Bull. M. C. Z., 18, pl. 12, f. 4, 1889, (as Cythara) ; Bull. 37, U. S.).NioM,,p.\100, ly io) esas
Distribution.—Gulf of Mexico, 220 fms. Also Antillean. galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, 1895.
Distribution.—Upper Miocene, Galveston well, at 2236- 2871 feet. Genus DAPHNELLA Hinds jeucophlegma Dall, Bull. M. C. Z., 9, p. 70, 1881; 18, p. 102, pl. 9. (1F, 10, 1880) Distribution.—Gulf of Mexico, Blake Station 2, in 805 fms. (Eubela) limacina Dall, Bull. M. C. Z., 9, p. 55, Aug., 1881; Ver- rill, ‘Tt. Conn, Acad!).5, p..452, ’82; 6, p.- 205; (Ba. eam Bull. M. C. Z., 18, p: 106, pl. 9, f. 10, ’80; Bull. 27. aaa Mo ps 100;\pl out ONyse! hormophora Watson, Jour. Linn. Soc., 15, p. 457, Nov. 1881; Chall: (Gastr/)p) 5a sole ninti) oy, 85)
74 MOLLUSCA OF THE GULF OF MEXICO 44
Distribution.—Massachussetts to Pernambuco. Guli of Mexico, Yucatan Strait, 640 fms. and Blake Station 2, west of Florida, at 895 fms.
Genus CANCELLARIA Lamarck
reticulata Linnzeus, Syst. Nat., ed. XII, p. I190, 1767, (as Voluta); Sowerby, Thesaurus, 2, p..442, pl. 92, f. 17, 1848; Oa oun Man), ’7, p69, pl. 2) £.25,:26,'.785; Dall, Bull. M. LOZ. 120). 8o; Vanatta Pro AlN. 8.2, Phila.,..55, p: 757, 1903. Distribution.—Hatteras to Guadeloupe, 5-30 fms. Gulf coast: Crooked Isl., Charlotte Harbor and Sarasota Bay, West Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene, New Orleans Pumping Station No, 7.
Note.—The fossil species, C. convadiana Dall, (Tr. Wagner Tnsty, 32,'\p. 42, 1890), is closely allied to C. 7 etzczlata.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. 100, pl. 3, f. 11, 1895.
Distribution.—Upper Miocene, Galveston well at 2552-2600 feet.
(Norona) clavatula Sowerby, Pr. Zool. Soc,, p. 52, 1832; Conch. Piao ie bryony Mati 7) p.075, ply ial E 61) 785: Eanes (Salt Am.) Pal. vol) 1. py 99,")’953: Dall, Pry U.S. INE 27,D. 200, TOLO:
Distribution.—Recent, Panama to Paita, Peru. Upper Mio- cene, Galveston well at 2552-2600 feet, (Harris). Notin the recent Atlantic fauna.
(Trigonostoma) agassizi Dall, Bull. M. C Z., 18, p. 130, pl. 35, ioanmosos Salhi 27 5..N: Mi, pe 104, pl. 35, £.14,." 89: Distribution.—Cape Lookout, N. C., 18 fms. Gulf of Mex- ico between Mississippi delta and Cedar Keys, 25 fms.
(Trigonostoma) tenera Philippi, Zeitsch. fur. Mal., 5. p. 24, 48;
A5 BULLETIN 38 75
Dall, Bull: 37, U.S. N. M., p) 104) 7807, 0a ee Sap ALOE Ie Kel O}
stimpsont Calkins, Pr. Dav. A. 5., 2, p.. 250, pli at: (aaa oO.
Distribution.—Gulf of Mexico, Yucatan region. Pliocene, Caloosahatchie beds, Fla.
Genus ABMETE Kroyer
microscopica Dall, Bull. M. C. Z., 18, p. 131, 1889,(as 77#g07- ostoma?); Bull. 37, U. SN. M.; p. 106," 30,3 eee My, 24). 504, Ple2oueert) 1902: Distribution.—Campeche Bank, 200 fms. Also off Fernan- dina and Cuba. Genus OLIVA Martyn
circinata Marrat, Sowerby’s Thes. Conch., 4, Oliva, p. 21, pl. 17, f. 277, 1880; Johnson, Nautilus, 24, p. 123, Mar: naum
litterata Lamarck, Ann. du Mus., 16, p. 315, 1810; Not of Bol- ten, 1798. Hilgard, House of Rep. Hx..Doei sep gree. 886,.'78; Dall, Bull: 37, U. S. N. Mj po meg late ene: 8a, 89; Singley, 4th Ann. Rept. Tex: |\G.1S)3peazeree Vanatta, Pro A. N.S: Phila.) 55, 1.757) looce
carolinensis Conrad; Pr. A. N.S. Phila., 14, pegegiae
Distribution. —Hatteras to the West Indies, o-2 fms. Gulf coast: Franklin and Calhoun Counties, Fla., Chandeleurs, Point on Fer, Cameron, La.; Galveston, Corpus Christi, Matagorda, Tex. Pleistocene, New Orleans artesian well of 1856, Lake Borgne borings, Knapp’s wells, Terrebonne Parish, No. 1 well at 2000-2450, No. 2 at 1780-1800, No. 3 at 1150-2029 feet. Pliocene, Caloosahatchie beds, Fla.
liodes Dall, Tr. Wagner Inst. Sci., 3, p. 1576, pl. 58, 1903.
Distribution.— Lower Miocene, Chipola marl, West Florida; Bascom No. 2 well, Mobile, Alabama, at 1241 feet, Chipola horizon. ;
+ : ? a. + il
76 - MoLLUscaA oF THE GULF OF MEXICO 46
Genus OLIVELLA Swainson
Miltica Say, Jour. A. N.S. Phila., 2, p. 228, 1822; Hilgard, Rept.
Citietpaneineers to Sec..War, p.. 358, ’70, House of Rep., Eeewwoc 1, pt 2,px 886, 778; Dall, Pr. U.S. N. M., 6, p. gee oa) Bull, 327).U.S. N. M.;"p. 106, pl..34, f. 1, 2, 7895 em WS. p45) )90; Vanatta, Pr. A.'N. Si Phila., 55, P. 757, 1903; Deussen, U. S. G. S. Water-Supply Paper, Nie)3353 P- 77; 1914. Distribution.—North Carolina to Trinidad Isi. Gulf coast: Recent, Cedar Keys, St. Mark’s and St. Joseph’s Bay, Fla.; Galveston and Corpus Christi, Tex. Pleistocene, New Or- leans artesian well of 1856, Lake Borgne borings; Knapp’s wells, Terrebonne Parish, No. 1 at 1600-1700, No. 2 at 1050- 1842, No. 3 at 1150-1839 feet; North Creek, Manatee and Labelle, Fla. Pliocene, Caloosahatchie beds, Fla., and New Orleans Gymnasium well at 1200 feet. Miocene, Gilbert well No. 10, Bateson, Tex. at 323 feet.
pusilla Marrat, Thes. Conch., t. 21, f. 356-358, ’71; Tryon, Man. Bonet 5. p64) pl. 14, £.. 45, 46;°33;, Vanatta, Pr. A. N. Da tanilas 55.) p./7575, 03.
Distribution.—Cited by Vanatta from St. Joseph’s Bay and Crooked Isl., West Fla.
nivea Gmelin, Linn. Syst. Nat., ed. XIII, p. 3442, 1792; Reeve, ones licon!, f.\64-a) 750: Tryon, Man., 5, p67; pl: 15, f. fee Wall, Ball ey) U.S, N. M., p. 106, ?89;, Dall & eaupson, Bull. U. 5. Fish Com.;1, p. 393, Ol. Distribution.—Sarasota, West Fla. to Haiti, 27-805 fms.
jaspidea Gmelin, Syst. Nat.,ed. 13,p. 3442, 1792; Reeve, Conch. Wcon-.-6, p:\22, f. 58,1850; Dall & Simpson, Bull, U,'S. Fish Woitl. 1, p. 392, I9OT.
Distribution.—Hatteras to Brazil and West Florida, 27-805 fms.
floralia Duclos, in Chenu, Ill. Conch., p.6; Tryon, Man., 5, p.
A” BULLETIN 38 ae
68, pl. 16, f. 2, 3, 1883; Dall, ‘Ball 27.0) een FOG,.4/ 80; Distribution.—Hatteras to Tortola and West Florida.
eutacta Dall, Tr. Wagner Inst. Sci., 3, p. 1576, pl (Sats 03, £903. : Distribution.—Lower Miocene, Chipola marl, Fla, and Mo- bile Oil Co’s No. 2 Bascom well near Mobile, Alabama, at 1241 feet, Chipola horizon.
gaivestonensis Harris, Bull. Am. Pal., vol. 1, p. roo, pl. 3, f. 13, 1895. Distribution.—Upper Miocene, Galveston well at 2410-2871 feeb.
subtexana Harris,. Bull. Am. Pal., vol: 1, -p. Toro D, S05: Distribution.—Upper Miocene, Galveston well at 2552-2871 feet
Genus MARGINELLA Lamarck
carnea Storer, Jour. Bost. Soc., N. H.,1, p. /465) pliioygiay ia) 1837; Sowerby, Thes., f. 103; Tryon, Man. )i5)\@) 22a, f. 86, 1883; Dall, Bull. 37, U.S.,.N. M:) p.aoerree Distribution.—Charlotte Harbor, West Fla., Florida Keys and Antilles.
storeria Couthouy, Jour. Bost. Soc. N. H., 1, p. 440, pl. 9,f. 1,2, 1837; Dall, Bull 27 UiaS.’. Noo M. ‘p. 106 Wises
Distribution.—Gulf of Mexico, West Fla., to Colon.
cassis Dall, Bull. MC. Z., 18, p. 137, pl. 3. t./8)weeor, eae 27 Duin LOG! ply 25 TaN noo. Distribution.—Gulf of Mexico, Lat.25° N., Lon. 84° W., at tol fms. Also off Cuba.
apicinca Menke, Syn. Meth. Moll., p. 87, 1828; Tryon, Man., 5, Pp. 33, pl. 10, £..89,.’83;) Dall, Bull. 37, U.S) Nov Ree
#
Bie ee SES
aaa
MOLLUSCA OF THE GULF OF MEXICO 48
89; Singley, 4th Ann. Rept. Tex., p. 333, 92; Vanatta, Bia. S. Phila. 55, p.)757).1903; Vaughan, 2nd Ann. Rept. Fla. Geol. Surv., p. 148, 1909.
gonocdatss) Reeve, Conch. Icon., 15, pl. 18, £: 87.
Distribution.—Hatteras to Jamaica, Gulf coast: West Fla. and ‘Texas, Pleistocene, Manatee, Orient and Labelle, West Florida.
pellucida Pfeiffer, Wiegmann’s Archiv; 1, p. 258, 1840; Tryon,
amen spe 3, Dl. tO, t./9n, 83: Dall Bally 37.) U. So .N: M., p. 106, ’89. Distribution.—Sarasota, West Fla. and West Indies.
matsoni, Wall) Bull. M.C:\Z:, 9, p, 71, 18817 18, p. 137; pl. 19, f.
Bear) b'2,) 89% Bill. a7) U.S. Ne M, pe 106s" pl. 19), £: Boley ont. 2)? 8a.
Distribution.—Yucatan Strait, 640 fms. Also off Havana, 220-805 fms.
virginiana Conrad, Am. Jour. Conch., 4, p. 67, pl. 5, f. 4, 1868,
(asi2raunwem): Dall, Bull, 37, U.S. N..M., p, 106, ’89; Tr. YM S85) 54S, 00.
Distribution.—Recent, North Carolina to Yucatan, 10-294 fms. Pliocene, Caloosahatchie beds, Fla. Miocene, James River, Va.
denticulata Conrad, Jour. A. N. S. Phila., 6, p. 225, pl. 9, f. 21,
nozon) (Not of Tate,.’78); Dall, Bull..37, U.\S...N..M.) p. Mes) Son lime We TS. 3) p. sr, pl. 5 f. 8, 1890: Distribution.—Recent, Hatteras to Barbados. Also West Florida, 5-294 fms. Pliocene, Caloosahatchie beds, Upper Miocene, Maryland and Virginia.
Opalina Stearns, Pr. Bost. Soc. N. H., 15, p. 21, 1872; Dall, Pr.
ee VM Gp 324.783: Bull.'27, pi 108.) 89) Cas den ticulata var. opalina).
Distribution.— Hatteras to Barbados. Recent, Tampa (type
49 BULLETIN 38 7 79
locality) and Cedar Keys, 14 fms. Also Barbados, 100 fms.
aureocincta Stearns, Pr. Bost. Soc. N. H., 15, p. 22, 1872; Dall, Bull. 37, U.S. N. M., p. 108,789; Tr. Wiss eee, 1890. Distribution.—Virginia to West Florida, 3-44 fms. Pliocene, Caloosahatchie beds.
minuta Pfeiffer, Wiegmann’s Archiv. fur Naturg.1, p. 259, 1840; Tryon, Man., 5, p. 43, pl. 12; £.60, 783; Dall Baise. 57 ENG OMe. py Pom. eOr Distribution.—Mediterranean, West Indies, East and West Florida, 5 -294 fms. Miocene to Recent. Also Pleistocene, southern Europe and of North Creek, West Fla.
minima Guilding, Sowerby, Thes. Conch., 1, p. 388, pl. 78, f. 220, 1846; Dall,Pr. U. S, Ni M., 6, p. 324) 3a, ee oe Man., 5, p. 44, pl. 12, f. 62, °83- Dall, Bull) a7 seeeeyes: p. 108,’ 89.
Distribution.—Cape Lookout, N.C., to Haiti. Also West Florida at Cedar Keys, 0-22 fms. Note.—Tryon placed this species in synonymy of mznuta Pfr. According to Moerch minima is asynonym of lavalleana d’Orbigny, (Moll. Cuba, 2, p. rot, pl. 20, f. 36-38, 1842). succinea Conrad, Pr. A. N.S. Phila., 26, pl: 1) aijomy ee Tryon; Man., 5) p.34, pl. ro, f: 03,783; Dall Bai So N. Map. ros. pl amo is 6,80. Distribution.—Fernandina, East Fla., to Sombrero, W. I. Gulf coast at Tampa, 7o-1002 fms. Placed by Tryon in synonymy with zztzda Hinds.
fusina Dall, Bull. M. C. Z.)\9, p. 72; 1881: 18, p. 193) qolamuoame A, 803; Bull, 37.0. Ooo a 145580, Distribution.—Yucatan Strait, 640 fms. Also off Fernan- dina, Fla.
ni le
80 MOLLUSCA OF THE GULF OF MEXICO 50
Nucatecand Wall, Bull. M. C. Z., 9, p. 72, 1881; 18, p. 138, pl. foes 30° Bull--37,--U,, 5S; IN. M.,-p..,106, pl. 19; f. 5, 1889.
Distribution.—Yucatan Strait, 640 fms. Also Sand Key, Fla. 125 fms.
Seminula Dall, Bull. M. C. Z., 9, p. 72, 1881; 18, p. 139, pl. 19, meee so Bull37, U.S. N. M.,; ps 108, pl. 19, 1. 2, °89:
Distribution.—Yucatan Strait, 640 fms. Also Antillean.
rostrata Redfield, Cat. Marg., Am. Jour. Conch., 6, p. 246, Giove)/1870;. Dall, Bull. M. C).Z., 18,9137, 89; Ir. W: 1. B25 oop. 50,, 1890. oblonga Sowerby, Thesaurus, pl. 76, f. 106, I08. 40N oblonga of Swainson.
Distribution.—Gulf of Mexico, Lat. 23° N., Lon. 89° W., in 84 fms. Pliocene, Caloosahatchie beds.
(Volvarina) avena Valenciennes, in Kiener, Coq. Viv. Warginella; Deg pl. 6G, 1) 24,1841; Sowerby, Thes. Conch., 1, p. 391, pPengoireo, 4g, Dall, Bulli 37, U2 5. N. M., p..108;."89; Wall) s Simpson, Bull. U. S. Fish'!Com., 1, p. 394, 1907; Vaughan, Publ. No. 133, Carn. Inst., p. 171, 1910.
Distribution.—Key West to Colon. Also Texas region, Gulf of Mexico. Pliocene to Recent. Pleistocene, La- bella, West Fla.
(Volutella) ovuliformis d’Orbigny, Moll. Cuba. 2, p. ror, pl. 20, meee 1e45:) Wall Bull 27, U. S. N. M:, p. 108, ’89; DaligéeStimpson, Bully. 5. Fish’, Com), p.395; 1901; Vaughan, 2nd Ann. Rept. Fla. G. S., p. 148, I910.
Distribution.—Cape Fear, N. C., to Guadeloupe Isl. Also West Florida. Pleistocene, Manatee, West Fla.
(Volutella) hadria Dall, Bull. M. C. Z., 18, p. 142, 1889; Bull.37, OS:N. M., p. 108, 80.
55 - BULLETIN 38 $x
lacrimula Dall, Pr. U. S. N. M., 6,.p. 324; 1883: “NetVacmenwla Gould, Pr Bost. Soc) Ni 8,) pp: 28r.oneoe:
Distribution.—Cedar Keys, West Fla. Collected by Hemp- hill on mud flats. Also Charlotte Harbor, West Fla. |
Genus VOLUTA Linnzeus
virescens Solander, Dillw. Desc. Cat., 1, p. 562, Sowerby, Thes. I, pl. 52, f. 78; Tryon, Man:, 4, p. 84; pl 2a ee ieeae: Dall, Bull: 37, W..S2 UNaiMe ip.) 108780: Distribution.—Texas to Cartagena, Colombia.
Genus SCAPHELLA Swainson
junonia Hwass, Chemnitz, Conch. Cab. 11, p. 16, pl. 177, f. 1703, 1704, pub, 1795; Dall, Bull. M. C. Z., 18, pangs eee 5, c-e, 89; Bull. 37, U. S. N. M., ps 110, (plyigay eee 1889. Distribution.—North Carolina to Nassau. Also Gulf of Mexico,''Lat. 25° IN.; Lon.82°) Wat 2, imie)
Genus AURINIA H. and A. Adams
dubia Broderip, Zool. Jour., 3, p. 81, pl. 3, #. .1, 1828) (as Voluta); Reeve, Conch. Icon., Voluta, pl. 22, f. 59, 1849; Tryon, Man., 4, p. 90, pl. 27, f..77, 81, 1882; Bolmmnijetem, Mal. Ges., 6, p..150, pl. 4, f. 1-3, °79; Dall) Bali 18, ps. F51,.’?809; Bull. 37, U. S. N. M,p: 110) eae, WEL) 2A oe OnmlN2g ETT TGqo2:
fusus tesellatus Schubert & Wagner, Suppl. Bd. Mart. und Chemn., Conch. Cab., 1829; Kiener and Reeve; but not of Zekeli and Pictet, 1852.
mutabilis ‘Tuomey & Holmes, 1856. Not of Conrad, 1841.
Distribution.—Hatteras to Cape Florida. Gulf of Mexico between Mississippi delta and Cedar Keys, at 111 fms.
robusta Dall, Bull. M,C, Z.; 18, p. 153, pl. 35, 4: 2)So;)ea
37; Pe Alo, plu 25 Wh. one es
Fae es a Se a a
=
82 MOLLUSCA OF THE GULF OF MEXICO 52
Distribution.—Gulf of Mexico, Lat. 28° N., Lon. 86° W.., at 280 fms. Also off Cuba, 119 fms. Genus MITRA Lamarck
gemmata Sowerby, Thes. Conch., sp. 334, 1874; Tryon, Man., Meptos pl. 56, f.( 688, °82; Dall, Bull) 37, U. S. N. M., p. t10, ’89; (as var. of hanleyz Dohrn): Dall & Simpson, bulls U.S. Fish Com., 1; p. 396; 1901. Distribution.—Charlotte Harbor, West Florida, to Jamaica.
straminea A. Adams, Pr. Zool. Soc., p. 132, 1851; Sowerby, ites, Couch., f. 561;) Tryon, Man.,.4, p.140, pl. 41, 1.188, Hoare Wall Bull. 327, U5, N..M:, p. 110, °89: Distribution.—Gulf of Mexico, west of Florida.
wandoensis Holmes, Post-Plioc. Fos. S. Car., p. 77, pl. 12, f. 10, tod, 1860, (as Volutomitra); Dall. Bull. 37, U.S. N. M.,p. mMOnwsO- tt WW. I.:S., 3, p. 92; 1890. Distribution.—Cape Hatteras to Florida Straits and West Florida, 14-80 fms. Very closely allied to and probably identical with 17. rusht Dall, (Conch. Exch., 2. p. 9, 1887; Bull. M. C. Z., 18,.p. 160, ’89)- from the Gulf of Mexico, 27 mos) PWS, Kish Com Station 2372.
mea oa Ball, M.'C Z., 18, p. 159, pl. 15, f. 6, 1889; Bull. Boe ING MM.) p. rio; pl. is, f. 6, 789.
Distribution.—Gulf of Mexico, west of Florida, t19 fms. Also Antillean,
(Conomitra) blakeana Dall, Bull. M. C. Z., 18, p. 163, 1889; Bull. B.S, Ni. M:, p, 110; 789.
Distribution.—Recent, Vucatan Strait, 640 fms. Closely related to the fossil species, stamznca Conrad and /ausotdes
Lea. Genus MITROMORPHA A. Adams
pygmaea Dall, Tr. Wagner Inst. Sci., 3, p. 95, pl. 10, f. 3, 1890.
5D
BULLETIN 38 &3
Distribution.—-Type from Caloosahatchie marl, Fla., Plio- cene. Also Bascom No.1 well, near Mobile, Alabama, at 1500-1556 feet. Miocene. Not in the Recent fauna.
Genus FASCIGLARIA Lamarck
gigantea Kiener. Icon. Coq. Viv., p. 5. pl. 10, 11, 1840; Tryon,
Man., 3, p- 75, pl. 60, f. 14, °81; Dall, Bullog7 Sa: p. 112, °89; Tr. W.I1.S., 3, p. 104, ‘oo Singley, Agnes Rept. Tex. G.'S., p. 334, 792; Dall & Simpson, Bales: Fish Com., 1, p. 397, I901; Vaughan, 2d Ann. Rept. Fla. G. S., p. 149, 1909; Johnson, Nautilus, p. 44, Oct., 1919.
Distribution.—Hatteras to Brazil. o-1o fms. Gulf coast; West Florida; Point au Fer. La.; Corpus Christi and Mat- agorda Isl., Tex. Pleistocene, New Orleans pumping Sta- tion No. 7, North Creek and Manatee, West Fla. Pliocene,
Caloosahatchie River marl.
tulipa Linnzeus, Syst. Nat. ed X, p. 754, 1758, (as Murex);
Kiener, Coq. Viv., p. 2, pl. 1, 2, 1840; Tryon, Sizeen aya 74, pl. 50, f. 1-4, 781; Dall, Tr..W. 1. Si ao ema It, 90; Vanatta, Pr. A. N. $.. Phila, 535, im genus. Johnson, Nautilus, p. 46, Oct., 1919.
Distribution.—North Carolina to Colombia and Antilles.o-10 fms. Gulf coast: Cedar Keys, St. Joseph’s Bay and St. Marks, West Fla., Texas coast. Pleistocene, North Creek, Fla. Pliocene, Caloosahatchie beds.
distans Lamarck, An. s. Vert., 7, p.119, 1822; Tuomey & Holmes,
PI. Fos. 8. C., p. 151, pl. 30, f..7, 1857; Hileard, /Eiguscver Rep., Ex. Doc. 1, pt.' 2,p, 886, ’78; Dall, Tr) Walser aie 102, pl..7, f. 10, ’90;, Singley, 4th Ann. Rept, Geol) Surv: Tex. ,:p. 334, 92; Mitchell, List. Tex. Sh., p. 16; Vauehoam: Publ. No. 133, Carn. Inst., p. 171, I910; Johnson, Nautilus, Pp. 47, Oct.) roe:
Distribution.—Hatteras to Yucatan o-54 fms. Gulf coast: Cedar Keys, St, Marks, Fla.; Cameron and Chandeleurs,
ee eee es —_— a a tg
Sa
34 MOLLUSCA OF THE GULF OF MEXICO 54
La.; Galveston, Matagorda, Corpus Christi, and Mustang Isl., Texas. Pleistocene, New Orleans well of 1856, and pumping station No. 7; Orient and Labelle, West Fla. The Pliocene shell of the Caloosahatchie beds is the related, but slenderer, /. apiczna Dall.
(Mesorhytis) meekiana Dall, Bull. M. C. Z., 18, p. 172, pl. 36, f. Peso nroul 370. SiN. M..p. 112; pl. 36, £. 75. 89- Distribution.—Gulf of Mexico, Blake Stations 16 and 20, at 292 and 220fms. Also ofi Cuba, 400 fms. The first living species of Mesorhytis known, the type being the Cretaceous
_ species, gvaczlenits Meek. Genus FUSINUS Rafinesque
timessus Dall, Bull. M. C. Z., 18, p. 166, 1889; (as Fusus); Bull. eo NGM p. 112; So.
Distribution.—Gulf of Mexico, between Mississippi delta and Cedar Keys, 60 fms.; between Cedar Keys and Dry Tortugas, 27 fms. Also near Cuba.
eucosmius Dall, Bull. M. C. Z., 18, p. 167, pl. 35, f. 5, 1889; Nae PeTT 2, pl. 35, £25, 7.80, Distribution.—Gulf of Mexico, west of Florida. Also Bar- bados, 27-111 fms.
couei Petit, Journ de Conch., 4, p. 249, pl. 8, f. 1, 1853; Dall, Del ME) C.Z., 18, p. 167; 780: Distribution.—Gulf of Mexico, between Tampa and Dry Tortugas, 26 fms.
alcimus Dall, Buil. M. C. Z., 18, p. 170, 18809. Distribution.—Gulf of Mexico, 100 miles north of Yucatan, 95 fms.
amiantus Dall, Bull. M. C. Z., 18, p. 169, pl. 15, f. 11, 1889; Bull. Bein IN. Map. \rr2) pl. 15, f.. 11, '80:
Distribution.—Gulf of Mexico, Blake station 2, at 805 fms.
55 BULLETIN 38 85
aepynotus Dall, Bull. M. C. Z., 18, p. 169, 1889. Distribution.—Gulf of Mexico, 84 fms. Also Antillean. amphiurgus Dall, Bull. M. C. Z., 18, p. 171, 1889. |
Distribution.—Gulf of Mexico, Blake station 45, at 101 fms.
Genus LEUCOZONIA Gray
cingulifera Lamarck, 7, p. 107, 1822, (as Turbinella); Reeve, Conch. Icon., 4, pl. 3, f-17, 1847; Dall and Simpsoneial: U.S. Fish Com., 1, p. 398-1 90r:
Distribution.—Sarasota, Fla., to Texas and Brazil. Also Antillean. ;
ocellata Gmelin, Syst. Nat., p. 3488, 1792, (as Buccinum);Reeve, Conch. Icon., 4, pl. 8, f. 38, 1847; Dall and Simpson, Bull: U. S. Bish Com) 4, p. 308; Gor
Distribution.—Cedar Keys, Fla., to Guadeloupe, W. I.
Genus BUSYCGN Bolten
pyrum Dillwyn, Desc. Cat. Rec. Sh., 1, p. 485, 1817; Dall, Bull. 37, U. S. N. M., p. 112, '89; ‘Tr. Wool. Silay eee Singley, 4th Ann. Rept. G. S. Tex., p. 334;o2-) Vanaur Pr. A. N. S. Phila.’ p. 757, 1903; Vaugiamy aise eae Carn; Inst.) p:.17t; 1910,
Distribution.—Hatteras to Gulf of Mexico. Upper Miocene to Recent. Gulf coast: Cedar Keys, Fort Barranca, St. Mark’s, Fla.; Cameron, &c. La.; Galveston, Matagorda, ¢ Corpus Christi, &c., Texas. Pleistocene of Louisiana at 4 Grand Chenier and New Orleans pumping station No. 7. A high spired variety isfound in abundance at Cedar Keys, Point au Fer, Cameron and Galveston, and in the Pleistocene of Grand Chenier, La., North Creek, Labelle, Manatee and Orient, West Fla.
canaliculatum Linneeus, Syst. Nat., p. 753, 1758, (as Wurex); Holmes, Post-Pl. Fos. S.C. p. 66, pl. 11, £. 3) )7850;) tia
a at
ne he gas aa LD
86
MOLLUSCA OF THE GULF OF MEXICO 56
gard, House of Rep., Ex. Doc., 1, pt. 2, p, 886, ’78; Dall, mea Oo. N. Mi, p. 112, pl. 73, f..1, °89; Clark, Pleis- roeeue, Md., p. 180,pls; 46; 47, 48, 1906; Johnson, Occ. Pa. Busi Soc. N. H., 7, p. 139, 1915.
Distribution.—Cape Cod to West Florida and Texas. Pliocene (Carolinas). Pleistocene, New Orleans well of 1856 (Hilgard).
perversum Linnzeus, Syst. Nat., p. 1222, 1766, (as Murex);
Tuomey & Holmes, Pl. Fos. S.C, p. 145, pl. 29, f. 3,1856; Ebiigard: Elouse of Rep. Hx. Doc. 1, pt. 2, p. 886, '78; Dall, iy at, S.,'3,.p. 116, 90; Vanatta, Pr. A. N.S, Phila., BoD 7) LOO3 Vaughan, Publ. 133, Carn: Enst., p. 171, IQIO.
Distribution.—Hatteras to Cuba. Miocene (Carolinas) to Re- cent, o-3 fms. Gulf coast: Cedar Keys, St. Marks, &c., Fla.; Point au Fer, Cameron, La.; Galveston and Corpus Christi, Tex. Exceedingly abundant in the Pleistocene of Grand Chenier, La., and found by Hilgard in the Lake Bor- gue borings; also of Manatee, Orient, and Labelle, West Fla.; Pliocene, Caloosahatchie beds, Fla.
carica Gmelin, Syst. Nat., p. 3545, 1792, (as Murex); Holmes,
Bestel. Hos.S.-C.,p: 65, pl. 11, £.. 1, 1859; Dall, Bull. U.S, ee MiP. Wii2. Dl. 74,:f. 1; °89° Clark, Pleistocene, Md., p. 179, pls. 43-45, 1906; Johnson, Occ. Pa. Bost. Soc. N. H., 7) P- 139, 1915.
Distribution.— Massachussetts to the Antilles. o-10 fms. Gulf coast, West Florida and Gaiveston, Tex.
eliceans Montfort, Conch. Syst. 303; Dall, Bull. 37, U. S. N. M.,
De ir2, So. Distribution.—South Carolina to Campeche. o-5 fms. Genus MELONGENA Schumacher
corona Gmelin, Syst. Nat., p. 3552, 1768; Tryon, Man., 3, p.
Mon) 41 Nt. 199-202, 1681; Dall, “L...W..1.S., 3; p. 121,90;
a7 _ BULLETIN 38 87
Vanatta; Pr. A. N,'S. Phila.,\55, p. 757) 4028 Vaughan, 2d Ann. Rept. /G. 5S) Bla p.m s \toee!
Distribution.—Florida Keys to Yucatan and the Antilles. Gulf coast: Cedar Keys, St. Marks, St. Joseph Bay, Fla.; La.; and Tex. Pleistocene, North Creek, Orient and La- belle, West Fla.
melongena Linnzeus, Syst. Nat. ed XII., p. 1220; Tryon, Man.,
3, p. 107, pl. 41, f. 197, 198, 1881; Dall Balkis7 en ae M., p. 112, 89; Harris, Bull. Am. Pal.) volyn eaten nas:
Distribution.—Florida to Grenada. o-50 fms. Gulf of Mexico, Texas region. Also Upper Miocene, Galveston well at 2448-2465 feet (Harris).
Genus CANTHARUS Bolten
tinctus Conrad, Pr. A. N.S. Phila., p.:25, pla, fa) 1346 ioe Pollia); Dall, Bull. 37, U.'S..N. M.; p.) cr6 egies. tonidea); Dall and Simpson, Bull. U. S. Fish Com., 1, p. 399, IQOT. Distribution.—Hatteras to Vera Cruz. Gulf coast: Cedar Keys, Fla., and Matagorda, Texas. Pleistocene, North Creek, West Fla.
cancellaria Conrad, Pr. A. N. S. Phila., p. 25, pl. 1, f. 12, 1846;
Dall, Pr. U.S. N..M., 6, p. 327, °837 'Simetey lane f Rept... Tex. G. S.,'p. 334, ’92; Vanatta, Pr: ALN (Se alee q 55, P- 757, 1903; Vaughan, 2d Ann. Rept; Figgas, % 148, 1909. ; Distribution.—Florida to Isthmus of Panama. Gulf coast: i Cedar Keys, Fla.; Horn Isl., Miss.; Cameron, Pointau Fer, Chandeleurs, La.; Galveston, Tex. Pleistocene, Grand Chenier and New Orleans pumping station No. 7, and of f Orient, West Fla. ‘ ‘i
orbignyi Payraudeau, Cat. Moll. de Corse, p. 159, pl. 8, f. 4-6, i 1826, (as Buccinum); Dall, Bull. 37, U.S.N. M., p. 116, 7m
Hy Ni hi Wy,
38 MoLLUSCA OF THE GULF OF MEXICO 58
°89; Dall and Simpson, Bnil. U.S. Fish Com., 1. p. 400, IgOI. Distribution.—Florida to Yucatan. 22-25 fms. Also Medi- terranean.
limbatus Philippi, Abbild, 1, 3, pl. 1, f. 9, Tryon, Man., 3, p. Maorepin 7st) 257 .F88T:) Dall, Bull. 37, Uy So IN. M.,'p:; FLO, 730. Distribution.—Gulf of Mexico to Cuba.
Genus PHOS Montfort
candei d’Orbigny, Moll. Cuba, 2, p. 129, pl. 21, f. 23-25, 1845, (25 Cancellavia): Arango, Fauna Mal. Cub., p. 201, 1878; Dall, Bull. 37, p. 116, 1889; Dall and Simpson, Bull. U. S.
Bish) Com.) 1, p; 401, 1gor, Distribution.—Hatteras to Colon and the Antilles. Gulf of Mexico, between Mississippi delta and Cedar Keys, at 88 fms. Synonyms, fde Dall, are antillarum Petit, 1853 and grateloupensis Petit, 1853.
parvus C. B. Adams, Contr. Conch., p. 59, 1850, (as Z7rz¢oz) ; Pall bull27,°U. 8. N. M,. por, 1889.
peicoius Dall, Pr. U.S.N. M.,6, p. 325, pl. 10; f. 9, 783.
Distribution.—Charlotte Harbor, West Fla., and Texas. Also Antillean.
galvestonensis Harris, Bull. Am. Pal. vol. 1, p. 102, pl. 4, f. 4, 5, 1895. Distribution.—Upper Miocene, Galveston well at 2158-2871 feet.
Genus NASSARINA Dall
columbellata Dall, Bull. M. C. Z., 18, p. 182, 18809.
Distribution.—Cape Catoche, Yucatan. greyt Dall; Bull.-M. C. Z-, 18, p. 183, pl. 32, f. 12a, 1889; Bull. Be Ne Mi p) 116, pl..32)f. r2a, "89:
59 BULLETIN 38 89
Distribution.—Arrowsmith Bank, Yucatan, 130 fms. Also Antillean.
bushi Dall, Bull..M. C. Z., 18; p. 182, pli 15, £212, meee
Distribution.—Gulf of Mexico, Blake station 5, 152-229 fms. Also off Sand Key, Fla. and Barbados.
Genus ALECTRION Montfort vibex Say, Jour. A. N.S. Phila., (1), 2, p. 23%, 1822, (aswVassa)’- Am. Conch., pl. 57, f.\2; Dall, Bull. 27, Uns 116, pl. 50, f, 8, 1889; Singley, 4th Ann. Rept. Texts. Pp. 335, 1892; Johnson, Occ. Pa. Bost. Soc) Naao 7 ara IQI5.
Jretensts Perkins, Pr. Bost: Soc.,' 13, p. Ti7,jreeo: Distribution.—Cape Cod to Colon. Gulf coast: Cedar Keys and St. Mark’s, Fla., Point au Fer, La., Galveston, Corpus Christi, Tex. Pleistocene, New Orleans pumping station No. 7; Manatee, Orient and Labelle, West Fla. Pliocene, Caloosahatchie.
ambiguus Montagu, Test. Brit., pl. 9, f. 7, 1803, (as Buccinum);
Dall, Bull. M. C. Z., 18, p. 184, ’89; Not amdbigua Dunker (West African), which is zzcrassata Strom.
alba Say, 1826; candidissima C. B. Adams, Krebs Cat., p. 32.
Distribution.— North Carolina to West Florida and Barbados. Yucatan Strait, 640 fms. Variety anztillarum d’Orb., Ca- loosahatchie Pliocene.
acutus Say, Jour. A. N.S. Phila., 2. p. 234, 1822, (as Vassa); Helmes, Post-Pl. Fos. S. C., p..72;~pl. 12) faassen Buccinum); Ailgard, House of Rep. Ex: Docraipel 2 p 886, °78; Dall, Bull.37, U. S. N. M.,: p. 116,47) Bo.) eres, Bull. Am. Pal., vol. 1, p. 102, °95; Vanatta, (Braga: Phila., 55, p. 757, 1903; Deussen, Water Supply Pa. 335, p. 77, 1914. Distribution. —Carolinas to Barbados. Gulf coast: Recent, St. Joseph’s Bay and Crooked Isl., West Fla.;Point au Fer,
core) MOLLUSCA OF THE GULF OF MEXICO 60
Cameron, La.; Galveston and Corpus Christi, Tex. Pleisto- cene, Knapp’s wells, Terrebonne Parish, No. 1 at 1600-2443 feet, No. 2, 1050-1842, No. 3, 58-1839 feet, New Orleans ar.esian well of 1856, Lake Borgne borings. Pliocene, New Orleans Gymnasium Club well at 1200 feet. Upper Mio- cene, Galveston well, 2871 feet. Miocene, Gilbert well No. 10, Bateson, Texas at 323 feet.
consensus Ravenel, Pr. A. N.S. Phila., p. 43, 1861; Dall, Bull. MONO e218.) Dy 185, 89, Distribution.— North Carolina to Florida. 8-49 fms. Gulf coast 20 miles off Charlotte Harbor, W. Fla., at 13 fms.
hotessieri d’Orbigny, Moll. Cuba, 2, p. 142, 1845, Atlas, pl. 21, idie@-42, (as /Vassa): Dall, Bull. M. C. Z., 18, p. 185, 1889; Dal) és Simpson, Bull. U.S. Fish Com.,. 1, p. 403; 1901.
Distribution.— Gulf of Mexico, Blake station 36, at 84 fms. Also Hatteras and Sombrero, 30-85 fms.
galvestonensis Harris, Bull. Am. Pal., vol. 1, p. roi, pl. 4, f. 2,. 1895.
Distribution.—Upper Miocene, Galveston well at 2410-2871 feet.
trigalvestus Harris, Bull. Am. Pal., vol. 1, p. ror, pl. 4a, f. 3, 1895.
Distribution.—Upper Miocene, Galveston well at 2158-2871 feet.
gpseletus Say, Jour. A. N. S. Phila., (1) 2, p, 232, 1822, (as massa); Gould, Inv. Mass., pl. 15, f. 210; Dali, Bull. 37, Mis. IN: M., p.-116, pl. 50, f. 9, 1889; Johnson, Occ., Pa. eee Oe ONY 7, p. 135, 915; Dall; Bull)i112,,0.. 5. N: Man, >. 103, 1921.
Distribution.—Nova Scotia to Tampa, West Fla. Also San Francisco Bay, (introduced with oysters from the Atlantic).
61 BULLETIN 38. gt
beaumontensis Aldrich, Bull. No. 1, Univ. Texas, July, 1901; Nautilus, p. 74 and figs., Nov. 1go1, (as Wassa).
Distribution.—Pleistocene. Spindle Top Hill oil well, near Beaumont, Texas, at 390 feet.
Genus STROMBINA Moerch
gibberula var. galvestonensis Harris, Bull. Am. Pal,, vol. 1, p. TOS, Pl.) sO, OO ss Distribution.—Upper Miocene, Galveston well at 2410-2871 feet. Note—The typical form of gizbberula Sowerby, (Pr. Zool, Soc., p. 115, 1832), is recent on the Pacific coast of Central America southward to Paiti, Peru.
Genus ANACHIS H. and A. Adams
avara Say, Jour. A. N.S. Phila., 2, p. 230, 1882, (as Columbella); Tuomey and Holmes, Pl. Fos. 8. C. p,139;) pliesmis me, 1857; Dall, Bull. 37, U. S. N: M.,p./116, plo 50) se ee, Vanatta, Pr. A..N. S. Phila., 55, p. 757, reos-uie@masom: Oce. ‘Pa. Bosty oc N.. dal.) pul nao) oiess
Distribution. —Recent and Pleistocene. Massachussetts to Florida Keys. Gulf coast: Crooked Isl., West Fla., and Texas. Pleistocene, Lake Borgne borings, La., and North Creek, West Fla.
avara var. semiplicata Stearns, Pr. A. N. S. Phila., p. 344, 1873, (as Columbella); Tryon, Man., 5, p. 159, pl. 55, f. 70, 71, 83: Dall, Balloiag.a,) S.. N..M., ps aeons Distribution.—Cedar Keys and Charlotte Harbor, West Fla.; Louisiana; Corpus Christi, Texas.
avara var. similis Ravenel, Pr. A. N. S. Phila., p. 41, 1861, (as
Columbella; ‘Tryon, Man., 5, p. 159, pl. 55, f. 69, 83; Dall, Bull. 37, p. 116. ’89.
a)
Q2 MOLLUSCA OF THE GULF OF MEXICO 62
Distribution.—Carolinas to West Florida, Galveston and Yucatan.
avara var. translirataRavenel, Pr. A. N.S. Phila., p. 42, 1861; Mrvon, Man. 5, p. 150, °83; Dall, Bull! 37, p. 116, *89. Distribution.—New York to West Florida, Louisiana and Yucatan.
catenata Sowerby, Pr. Zool. Soc., p. 52, 1844; Conch. Icon., 11, Mien bs, Tho, 1850;. Lryon, Man.-' 5; p. £79, | pl. 58,2. 51, fogs Wall and Simpson, Bull. U.S. Fish Com,, 1, p. 404, LOOT,
Distribution.—Jamaica to Porto Rico. A shell closely re- sembling this species, and probably identical with it, is in our collection. As far as I know, this is the first time that catenata has been found on the Gulf coast.
obesa C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 2, 1845, (as ipiccnuMm): iryon, Man., 5, p. 169, pl. 57, f.7 and 20; 783; Dall Bull 37, p. 118, °89; Singley, ath Ann. Rept. Tex. G54. e25,,.92; Harris, Ball. Am, Pal., vol. 1, p. 102,’95; Wanatta) Pr. A. N.S. Phila, 55; p. 758, 1903.
ornata Ravenel, cancellata Gaskoin. A. ostreicola Melvellis a dark brown or black variety.
Distribution.—Hatteras to St. Thomas, W. I. and Vera Cruz. Phocene to Recent. Gulf coast: St. Joseph’s Bay,
- Crooked Isl., West Fla.; Point au Fer, Cameron and Chan- deleurs, La.; Galveston and Corpus Christi, Texas. Pleisto- cene, Grand Chenier, Teche No. 1 well, Jennings, La., at 1158-1199 feet, Knapp’s well, Terrebonne Parish, No. 1 at 1050-1800 feet, No. 3 at- 1150-1440 feet, Galveston well at 440-446 feet. Pliocene, New Orleans Gymnasium Club well abn iZ200 feet.)
amphisella Dall, Bull, M. C. Z., 9, p. 91, 1881; 18, p. 188, pl.
mM es) So: Bull) a7, Sy N. My, p.,118,, pl.).ro, f,) £00, 80.
63 BULLETIN 38 | 93
Distribution.—Vucatan Strait 43—640 fms.
hotessieriana d’Orbigny, Moll. Cuba, 2, p. 138, pl. 21, f, 37-39; Tryon, Man., 5, p. 144, pl. 52, £. 82. 1883; /Dall Bailes) Pent DS OG,
Distribution.—Tampa, West Fla., to Guadeloupe, W. I. Very closely allied to the following species.
pretri Duclos, Chenu, Conch. Il]., pl, 16, £. 7, 8; Tryon) Man.; Spat44, pl. 52k. 7 Ones Note.—C. B. Adams’ albella [Conch. Contr., p. 63, 1850, (as Pleurotoma); Dall Bull. U.S. N. M., p. 118, 1889] an unfigured species, is ide Dall and Simpson (1901) merely a mutation of pretrz, as shown by cotypes from Adams.
Distribution.—Variety albella, Cape Fear, N. C., to West Florida and Jamaica.
Genus COLUMBELLA Lamarck
mercatoria Linnzeus, Syst. Nat. ed. X, p. 730, 1758, (as Voluta); Kiener, Cog. Viv., Columbella, p. 23, pl. 542 0/2340; ) em Bull. 37. p. 116, ’89; Dall and Simpson, Bull. U.S. Fish Com) 1) piges s1oor. Distribution.—Cape Lookout, N. C., to Barbados and West Florida. Pleistocene of Costa Rica.
rusticoides Heilprin, Tr. Wagner Inst. Sci., 1, p. 81, pl. 8, f. 9, 1887? Pr. A. N.S.) Phila., p. 52, pl..3, 4.) 2,\?3 70 ibaa W.I1.S.,: 3, pi 135,790; Vaughan, Publ. (Noi 132 anae Tnst,))p.) 171, 19HO:
Distribution.—Cedar Keys and St. Marks, West Florida to Cuba. Pleistocene, Labelle, West Fla. Pliocene, Caloosa- hatchie beds, West Fla.
Genus ASTYRIS H. and A. Adams
lunata Say, Jour. A. N.S. Phila., 5. p. 213, 1826, (as Massa ) Tuomey aud Holmes, Pl. Fos. S.C. p. 136, pl. 28) £. am
04 MOLLUSCA OF THE GULF OF MEXICO 64
1857, (as Buccinum); Hilgard, House of Rep. Ex. Doc., 1, Bee. ps S86, 0878; Dall), Bull) 37). U.S. Ny M., p. 118, pl. powmeewo. 59: Johnson, Occ.’ Pa.’ Bost. ‘Soc. N.))H:, 7, p- 133, I915.
Bnccinum zonalis Linsley, Am. Jour. Sci., (1); 48, p. 285, 1845. Distribution.—Canada to Florida Keys. o-12 fms. Gulf coast: Cedar Keys, Fla.; Chandeleurs, La.; Galveston and Corpus Christi, Tex. Pleistocene. Lake Borgne borings, La., Knapp’s wells, Terrebonne Parish, No. 2 at 1050-1790, No. 3 at 258-1375 feet. Also of North Creek, Manatee and Labelle, West Fla. Pliocene of the Caloosahatchie, Fla.
duclosiana d’Orbigny, Moll. Cuba, 2 p. 136, pl. 21, f. 31-33, 745. (Not of Sowerby, 1847, which is a Javan shell); Dall, Bull 37, U.S.N.M.,p. 118, ’89, (as dunafa var.); Dall and Simp- eon) Bull U.S. Bish Com. , 1; p., 405, 1901. Distribution.—Hatteras to Barbados and Tampa, West Fla. 0-63, fms.
diaphana Verrill, Tr. Conn. Acad., 5, p. 513, pl. 58, f. 2, 1882; Weal Bally M | CZ. 5" 18) pe for, pls 35,’ f.:9),789; Ball. 37, oN MEL pw) DIS, ph. 35) "£9, 789: Distribution.—Rhode Isl. to Gulf of Mexico between Mis- sissippi delta and Cedar Keys, 64-487 fms.
Genus NiTIDELLA Swainson
parvula Dunker, Zeitschr, Mal., p, 64, 1847, (as Buccinum); peiom Man!) 5, p.n22,.pl.i48, £. 77;°83: Dall, Bull. 37, Wise N. M., p! 118, 789.
Distribution.—Gulf of Mexico, Texas region, to Barbados. Genus AESOPUS Gould
stearnsi Tryon, Man. Conch., 5. p. 179, 1883, (as Semznella); ait) Ball. Miu C.. 22-78, p. 194; pl..29)f..5, °89;, Bull. 37,
PEs, ploi29; £.) 5) 789. ,
65 BULLETIN 38 65
jilosa Stearns, Pr. A. N.S. Phila., p. 345° Gees Nitidella). Not Aesopus tlosus Angas, 1867, cf. Columbella peculiaris Guppy, Geol. Mag., London, pl. 18, f. 20, 1874.
Distribution.—Type locality, Tampa Bay, Fla. Also off Cape Fear, N. C. 15-17 fms. Pliocene of the Caloosahat- chie, Fla.
Genus MUREX Linnzeus
beaui Fischer and Bernardi, Jour. de Conch., 5, p. 295, pl. 8, f. I, 1856; Dall, Bull. M:-C3Z., 18, p. Tos Sa; 3R ame We NG Mp. ise sop
Distribution.—Gulf of Mexico between Mississippi delta and Cedar Keys, at 111 fms. Also Florida Reefs and Guade- loupe. 82-183 fms.
cabriti Bernardi, Jour. de Conch., 7, p. 301, pl. 10, f. 3, 1858; Dall, Bull.37,.W.48. N; (MoM. pi irs) ise;
Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake station 36, at 84 fms.
messorius Sowerby, Pr. Zool. Soc., p. 137, 1840; Conch. Illust., Murex, f. 93, 1839; Reeve, Conch. Icon., Wurex, f. 90, 745; Dall, Tr. W. I.S., 3, p. 139, ’90; Dall and Simpson; amit, U.S. Fish (Com) (1 \. 407, 1901,
Distribution.—Gulf coast, Cedar Keys, Fla. and Texas, to Colon. 2-30 fms. Pleistocene, New Orleans pumping station No. 7. Pliocene of the Caloosahatchie River, Fla.
(Chicoreus) rufus Lamarck, Anim. s. Vert., ed. 1, 7, p. 162,1882; Tryon, Man., 2, p. 90; pl. 15, f. 148, ’80; (Atle eae TO): Dall) Bull-377) UesvN.; M, p; 118788
sexcostata Emmons, Geol. N. Car., p. 248, f. 106, 1858.
Distribution.—Cape Fear, N. C., to Cartagena, Colombia, and Antilles. Also West Fla. 5-30 fms. Pliocene Ca- loosahatchie beds, Fla.
ce aisteal ave
96 MoLLUSCA OF THE GULF OF MEXICO 66
(Chicoreus) brevifrons Lamarck, Anim. s. Vert., ed. I, 7, p. 161, 7922: Reeye, Conch. Icon., 3, pl. 3, f.13, 1845: Tryon, Man., Peres tt) 172, 188o:-Dall, | Bull.’ 327, U.S. N...M., p:-118, 89; Ir. W.1.5., 3, p, 140,’90; Dall and Simpson, Bull. U. Se hish'Con:, 1, p. 407, 1901.
Distribution.—South Carolina to Cartagena, Colombia. Gulf coast, West Fla. Pliocene, Caloosahatchie beds.
(Phyllonotus) pomum Gmelin, Syst. Nat., p. 3527, 1792; Reeve, Wonehlicon.,.3, pl..o; £..35, 1845; Tryon, Man., 2, p.' 97, Wueze.if. 152, 80; Dall, Bull: 37, U.S: N. M., p..120, pl. 16, eee oor, br W.t. Si, 3, p..142; 90; Dall and Simpson, Bau. S. Fish Com., 1, p. 408; 1901.
oculatus Reeve, 1845; asperrimus d’Orbigny, 1853; mexicanus Petit, 1852; zmperzalts Swainson; globosa Emmons, 1858.
Distribution.—North Carolina to Venezuela. Gulf coast at various localities, West Fla.; Cameron and Chandeleurs, _La., and Texas. Pleistocene, New Orleans pumping station No. 7. Pliocene, Caloosahatchie River, Fla. Lower Mio- cene, Antilles. (Phyllonotus) fulvescens Sowerby, Conch. Illust., f. 30; Kobelt, Walia. Mal. Gesell, p..155, 277; Dall, Bull. 37;.U. S. N. M., p20, So, Vanatta, Pr. A. N.S. Phila., 55, p. 758, 1903.
Distribution.— Hatteras and the Gulf coast at Indian Pass, West Fla.; Point au Fer, Chandeleurs, La.; Galveston, Corpus Christi and Quintana, Texas. Pleistocene, Grand Chenier, La., Knapp’s wells, Terrebonne Parish, No. 1 at 2000-2150 feet, No. 3 at 790-830 feet.
Genus FAVARTIiA Jousseaume cellulosa Conrad, Pr. A. N.S. Phila., 3, p, 25, 1846, (as A/urex); Mapu M. C:.Z.,18,p: 210, pl. 16, f..1;.°89; (as Ocenebra) ; agg) Oo. No Me p..t2o0, pl,.16, f..1,.?89; Tr. W. 1. S:, a P: 150: 90. nuceus Moerch, Cat. Kierulf,p. 14, pl. 1,f. 9, ’50; Cat. Yoldi, p. 95, 1852, (as Ocinebra).
67 BULLETIN 38 97
Distribution.—Cape Lookout, N. C., to the Antilles and Vera Cruz. Gulf coast, Tampa Bay (typelocality), Cedar Keys and Texas. Pliocene of the Caloosahatchie.
cellulosa var. levicula Dall, Bull. M. C. Z., 18, p, 211, 1889; Bull. 37, p. 120, ’89, (as Octnebra).
Distribution.—Cape Lookout, N. C., to West Florida and Yucatan. 22-37 fms.
intermedia C. B. Adams, Contr. Conch., p.°60, 1850, (as Murex); Dall, Bull, 37, U. S. N. M., p. 220,80, (Gs Gemesra) al. and Simpsou, Bull. U. S. Fish Com., 1, p, 408, 1901.
Distribution.—Key West to St Thomas and Texas region of the Gulf. Variety a/ta Dall, Caloosahatchie Pliocene.
Genus UROSALPINX Stimpson
cinereus Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Fusus); Amer. Conch., 3, pl. 29, 1831; Gould, Inv. Mase pera, 637, (as Buccinum); ‘Tryon, Man., 2, p.2252)) ply go) tia ee. Dall, Pr. U.S. N. M., 6, p. 326, 1883; Buollitay i Sa M., p. 120, pl..50, f..6, ’89; Johnson, Occ.’ Pa, Bost pen: H., p. 132, 1915; Dall, Bull. 112,,U: S. No Mo ee ea,
Distribution.—Nova Scotia to Florida. Gulf coast, Cedar Keys (Dall); Pleistocene, Lake Borgne borings, La., (Hil- gard). Also on the Californian coast (introduced with oysters).
perrugatus Conrad, Amer. Jour. Sci., p. 397, 1846, (as /usus); Dall, Bull. M. C.° Z:, 18, p. 214, ’89; Bull.27)\pi kee eee Vatighan, 2d Ann. Rept. Fla. G.S., p. 148. 1900.
Distribution.—West Florida from Cedar Keys to Key West. Type locality, Manatee River, West Fla. Pleistocene, North Creek and Orient, West Fla.
tampaensis Conrad, Pr. A.N.S. Phila., 3, p. 25, 1846, (asMurex); Dall, Bull. M. C. Z.,.18, p. 214,789; Bull. 37, U.S eee Pure unise.
Q8 MOLLUSCA OF THE GULF OF MEXICO 68
Distribution.—West Florida from Cedar Keys to Sarasota. Pleistocene, North Creek, West Fla.
Genus MURICIDEA Swainson
multangula Philippi, Zeitschr. fur Malak, 5, p. 25, 1849, (as muss) Apbild. und’ Beschr , 3, p. 117, pl. 24,.f. 6, 1850; Dewmesimis7 Uys, N: Mi, p. 120,:pl. 76,4: 1) S80; “Fr. W. ese aeo. £40,1890° Pr: U. S. N. M., 24) p. 505, pl. 30, tel, 1QO2:
mempniiz Dall, Pr. U.S. N. M., 6, p. 327, 1883.
Distribution.—Cape Fear, N. C., to Yucatan and the Noith- ern Antilles, Gulf coast, Cedar Keys and Texas. 0-95 fms. Pliocene, Caloosahatchie beds.
estrearum Conrad, Pr. A. N.S. Phila., 3, p. 25, 1846, (asMurex); IPA es tWi . IN. Mi. 24, p.' 505, pl. 30,f. 2, 1902.
weort~aya Conrad, Am. Jour. Conch., 5, p. 106, pl. 12; f. 4, paoomtas, Cirosalpinx); Dall, Pr. U. S. N. M., 6, p. 326, °83; Balewlyee7,,.1S, Pp. 212, °8o;' Tr. W. 1. S.,: 3,'p. 140, °90, Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909.
Distribution.—LEast Florida south of St. Augustine and Gulf of Mexico, Cedar Keys, Fla., to Yucatan. 0-13 fms. Rocky shores. Pleistocene, Manatee and Orient, West Fla. Plho- cene, Caloosahatchie beds, Fla. Resembles Uvosalpinx perrugatus Conrad, but is slenderer and has a very different operculum.
hexagona Lamarck, Anim. s. Vert., 7, p. 169, 1822, (as Murex); iceve, Conch: [con:, 3; pl: 27; f. 120, 1845; Dall, Bull. 37, WerS.N. M., p. 120, '89: Dall & Simpson, Bull. U.S. Fish Com,, 1, p.\409, Ioor. Distribution.—Gulf of Mexico, Texas region to St. Thomas, MWit.25 fms. Genotype.
philippiana Dall, Bull. M. C. Z., 18, p. 213, 1889; Bull. 37, U.S. nee p. 120,89; Pr. US. N.'M., 24, ps 504, pl. 29,)\f: 5, 1902.
69 BULLETIN 38 99
Distribution.—Off Cape Catoche, Yucatan, and also Key West, Fla. 20-25 fms. Coral sand.
Genus EUPLEURA H. and A. Adams
caudata Say, Jour. A. N.S. Phila., 2, p. 236, 1822, (as Ranella); Stimpson, Am. Jour, Conch., 1, p.7 58, \ ply Jenene a) Hilgard, House of Rep. Ex. Doc..1, pt. 2; p)i886,/"7o oa, Pr. U.S. N. M., 6, p. 326, 383; Bull: 33. SS ee 120, pl. 50, f. 11, 789; Johnson, Occ. Pa. Bast/seche i, Tp BON MOE se Distribution.—Massachussetts to Florida. 1-3 fms. Gulf coast: Cedar Keys, St. Mark’s, Charlotte Harbor, Fla.; Chandeleurs, La. Pleistocene, Lake Borgne borings and New Orleans well of 1856 (Hilgard), and of Labelle, West Fla.
caudata var. sulcidentata Dall, Tr. W. I. S., 3, p. 144, 1890.
Ranella clathrata ‘“‘Gray’’ Calkins, Pr. Day. Beads seria: ZA LS 1or
muricitormis Tryon, Man., 2, p. 158, (in part). Not of Broderip. Distribution.—Gulf coast of Florida, Pleistocene of South- west Fla.
Genus TROPHON Montfort
(Boreotrophon) lacunellus Dall, Bull. M. C. Z., 18, p. 205, pl. 15, f..4,,1889; Bull, 27; U.S. N. M., p. 120, pl nse sey
Distribution.—Cape Fear, N. C., to Dominica. Also West Florida. 227-769 fms.
Genus THAIS Bolten
patula Linnzeus, Syst. Nat., ed. X, p. 739, 1758, (as Buccinum); Reeve, Conch. Icon., 3, pl. 1, f. 3, 1846; Tryon, Man., 2, p. 159, pl. 43, f. 19-22, 780; Dall, Bull. 37. (Ul Sie
XOO
MOLLUSCA OF THE GULF OF MEXICO 70
igenneg. (as Purpura); Dall, Pr. U.S. N. M.,. 37; p. 221, IQIO.
Distribution.—East and West Florida to Texas, the An- tilles and Brazil. Also Pacific, Gulf of California to Pana- ma and Peru.
haemastoma Linnzeeus Syst. Nat., ed. XII, p. 1202; Reeve,
Conmehumteon,, 3) 1.21, 1846;' Tryon, Man, 2,.p..167, pl. 49, Hose vos oso; Wall, Bull. 27, U.S. N. M., -p, 122, pl. 34, £2134 4).pl. 46," Ia-2b, ‘89; Singley ath: Ann. Rept, Tex. G. Sener ago..(o2* Vanatta, Pry A. N.S.) Phila., 55,.p.°758, 1903. (All as Purpura).
Distribution.—Cape Hatteras to Trinidad Isl. Gulf coast Cedar Keys and Ft. Barranca, Fla.; Horn Isl., Miss.; Point au Fer, La.; Galveston, Corpus Christi, Tex. Pleistocene, Grand Chenier, La., New Orleans pumping station No. 7, Knapp’s No. 3 well, Terrebonne Parish, at 1400-1440 feet.
haemastoma var. floridana Conrad, Jour. A. N.S. Phila., 7, p.
20 DE) 20,01. 21/1837; -bryom, Man., 2, p.' 167, -ply.40, f. 85, pl. 50, f. 86; Dall and Simpson, Bull. U. S. Fish Com., Dp ait, to0or.'’ CAll as Purpure).
Distribution.— Florida to the Antilles, Guif coast, West Fla.., Cameron, Chandeleurs, La.; Matagorda Bay, Tex. Pleisto- cene of Grand Chenier, La., (very abundant and showing the same mutations as the living shells).
deltoidea Lamarck, Anim. s. Vert., 7, p. 247, 1822; Reeve,Conch.
eons, 3;: pl.) 4, f. 18,1846; Kuster, Conch. Cab.,:pl.22, f. Pel ryon,) Man. 2)p. 163, ‘pl..47, £:.55,. 1880; Dall, Bull. Bas. | ONG May p. 22). 89;'-Dall and Stimpson, Bull, U. S, Fish Com:, 1, p.411, t901. (All as Purpura). Distribution.—Florida to St. Vincent, W. I., Gulf coast, West Fla. and Tex.
71 BULLETIN 38 IOE
Genus SISTRUM Montfort
nedulosum C. B. Adams, Pr. Bost. Soc. N. H., p. 2, 1845, (as Purpura): Tryon, Man.,-2, p. 190, pl So; 7s ose; Dall and Simpson, Bull. U. S: Fish Com., 1) ps ariesger
Distribution.—Cape Romano, southwest Florida, to Colon. Also Antillean.
roseum Reeve, Conch. Icon., sp. 46, 1846; Dall) Bullj27 i: NM po 522, Toso. Distribution.—Gulf of Mexico, Texas region, to Barbados. The rosy color is thought to be caused by an encrusting hy- drozoon.
Genus CORALLIGOPH{LA H. and A. Adams
abbreviata Lamarck, Enc. Me’th., pl. 435, f. 2a-b, 1795, (as Pyrula); Anim. si Vert... 7. p146, S22: galea Chemnitz. Distribution.—Cape Fear, N. C., to the Antilles. Gulf coast, West Florida. Pliocene, Caloosahatchie beds.
lactuca Dall, Bull. M. C. Z., 18, p. 220, pl. 16/4. 6, 188e)
Distribution.—Gulf of Mexico, Blake station 5, in 152-229 fms. Also off Fernandina, Fla.
deburghiae Reeve, Pr. Zool. Soc., p. 208, pl. 38, f. 3, 1857, (as Rhizochilus); Fryon, Man., 2, pl. 66, 1.) 382 )/iieeayaiae Latiaxis); Dall, Bull. U.S. N. M., p. 122)( plore) isesor
Distribution.—Cape Hatteras to Barbados. Also West Fla. 56-878 fms. Genus EPITOMIUM Bolten
angulatum Say, Amer. Conch., No. 3, pl. 27, (two upper figs) 1831, (as Scalaria clathrus var.); Hilgard, House of Rep. Ex. Doc. 1, pt. 2, ‘p. 887, 1878; Dall. Bully ig7 ese M., p. 122, °89; Singley, ath Ann. Rept. Vex: 3s aaa: 336, ’92; Johnson, Occ. Pa. Bost. Soc. .N. Hi,7)qpeiroas
IQI5.
r02 MOLLUSCA OF THE GULF OF MEXICO aie:
Distribution.—Connecticut to West Fla., Point au Fer., La., and Galveston, Tex. Pleistocene, Lake Borgne, La., bor- ings.
sayanum Dall, Bull. M.C. Z., 18, p. 313, 1889, (as Scala); Bull. Oe on IN. M., p. 122, pl. 50, f. 10, 789; Ir. W-- 1. Ss, 3, p- 158,90; Singley, 4th Ann, Rept. Tex. G. 5., p. 336, 1892.
clathrum Say (in part).
Distribution.—Virginia to Florida Keys, West Florida, Cam- eron, La., Corpus Christi and Galveston, Texas. Pleisto- cene, Grand Chenier, La. Pliocene, Caloosahatchie River, Florida.
tenue Sowerby, Pr. Zool. Soc., p. 110, 1844, (as Scala); Dall, Baie aU S.-N. M.; ps r22; 1880.
Distribution.—Gulf of Mexico, Texas and Cameron (?) La., to St. Thomas, W. I.
nitidellum Dall, Bull. M. C. Z., 18, p. 314, 1889, (as Scala); Bull. Be OemN. IM. p.9124, 89; Pr. U: S. N. Ms;.24, po.505, Die Zo; h. 8)) 1902. Distribution.—Hatteras to Cedar Keys, Fla. 32-63 fms.
Peng) Wall) (Bull) Mi .C. Z., 18, p. 310, 1889; Bull. 27, U. S. N, Mero 124.7 So, (as, ocala): Pr.. U.S. N..Mis (24, pe 506, piezo) f2_10, 1902.
Distribution.—Hatteras, N. C., to Texas and Vera Cruz, Mexico. 1I0-16fms. Pink with white varices.
denticulatum Sowerby, Thes. Conch., p. 87, pl. 32, f. 25, 26, ’47; Mevouy) Man 9, pl. 13/f., 100, 1887; Dall, Bull. 37, U. S. New ME. Dp, E24, ° 89.
Distribution.— Hatteras to West Florida and the Bahamas.
lineatum Say, Jour. A. N. S, Phila., 2, p. 242, 1822, (as Sca- laria); Wilgard, House of Rep. Ex. Doc. 1, pt. 2, p. 887, '78; aon.) Man 9, p79, pl. 16, f. 14, 87; Dall, Tr. W.1.S.,
ie! BULLETIN 38 I03
3, p..158, ’90; Johnson, ‘Occ.’ Pa.’ Bost. Sac IN aie ae TO2Q\ FOILS!
Distribution.—Massachussetts to Charlotte Harbor and Cedar Keys, West Fla.; Cameron, Point au Fer, La.; Gal- veston and Corpus Christi, Tex. Pleistocene. Grand Chenier, La., and Knapp’s No. 2 well Terrebonne Parish, at 1550-1570, 1731-1739 feet. Pliocene Caloosahatchie beds, Fla.
turriculum Sowerby, Thes. Conch., 1, p. 92, pl. 33, f. 61, pl. 34, f, 88, 1847; Singley, qth Ann. Rept. Tex. @)) Si) py 24a 1892.
Distribution.—Cape Hatteras to Haiti. 16-32 fms. Gulf coast at Galveston (Singley). Pliocene of the Caloosahatchie River, Fla.
mitchelli Dall, Nautilus, 9, p. 112, Feb. 1896; Pr. U. S. N. M., 24, 'P.) 506, pl 30; 78-3, )/4, oo2: Distribution.—Matagorda Island, Texas. Collected by J. D. Mitchell.
cochlea Sowerby, Thes. Conch., Scalaria, p. 103, pl. 35, f. 142 (only) 1847;Moerch, Jour. A. N. 5. Phila., 3, pi 205;2876, Tryon, Man.,9, p. 81, pl..17,#. 24,(1887;Dalli iPr We Mii245. Pp. SOON Db 20, fez, Toe2.
Distribution.—Hatteras, N. C., to Tortola. W.I. Gulf of Mexico near Cedar Keys, Fla., at 25 fms. Sowerby’s refer- ence to its occurrence at Loando, Portuguese West Africa, is questionable.
galvestonense Harris, Bull. Am. Pal., vol. 1, p. 103, pl. 4, f. 7, 1895.
Distribution.—Upper Miocene, Galveston well at 2552-2871 feet.
Note.—The following species of Afztomium have been re- ported by Singley from the Texas coast:—sevicifila Dall,
1O4 MoOLLUSCA OF THE GULF OF MEXICO 74
(Bull. M. C. Z., 18, p. 313, 1889); multisiviatum Say (Jour. Powe Nesow EF oila.,'5, p. 208, 1826; Dall, Bull. 37,,0.S..N.M., pl: 50, I. 5, 1889);contorquatum Dall, (Bull. M. C. Z., 18, p. 318, pl. 18, f. 9, 1889); and modestum C. B. Adams (Jamaica Shells). These species are not cited, as far as I am aware, by other authors from Gulf coast. They are not in our.col- lections from the Gulf and are noted here solely Ade Singley, (Fourth Ann, Rept. Tex. Geol. Surv., p. 336, 1892).
Genus JANTHINA Bolten
janthina Linnzeus, Syst. Nat., ed. X., p. 772, 1758, (Helix); adiOrbigny, Voy. Am, Me'r., p. 413, pl. 61,’ f. 8-10, £841; Waller. SN. M:, 37,°p. 223, ror0; Johnson; Occ. Pa. Bost. soci N,, H., 7, p., 101, 1915.
fragilis Lamarck, An. s. Vert., p. 89, 1801; Tryon, Man., 9,
p. 36, pl: 9, f. 4, 1887; Singley, 4th Ann. Rept. G. 8. Tex- aeup.ago, too2. Bartsch, Bull.’ or, U..S..-N...M.,. p. 65, one Distribution.—Nantucket, Mass., to Colon; and West Fla. to Quintana and Galveston, Texas. Pelagic. Also Pacific Ocean, Lat.42°N.to 36° S.,and Indian Ocean, Port Alfred, S. HK. Africa.
pallida Harvey, Thompson’s Ann. Phil. and Nat. Hist., 5, p. 96, plenie 2. tsar: Tryon) Man.,.9, p..37. pl. 10, 4.15, 1887. Distribution.—Europe, the West Indies and Gulf of Mexico, Cameron, La. (washed ashore). Also tropical waters of the Pacific Ocean. Pelagic, |
globosa Swainson, Zool. Illust. (1), 11, pl. 85, 1822; Tryon, Maw, 9, pl. 10; f.\124287;) Singley, 4th Ann. Rept. Tex., p. 336, .’92; Mitchell, List Texas Shells; Bartsch, Bull. 91, eS. NM: p..65,.1915.
Distribution.—Gulf Stream, Pelagic. Washed ashore at Quintana, Tex. and West Fla.Also Port Alfred, S.E. Africa.
communis Lamarck, An. s. Vert., ed, Desh., IX, p. 4; Tryon,
75 BULLETIN 38 Tos
Man., 9, pl. 9, f 99, 1887; Dall; Bull. 27, SU ase eo. 126, ’89; Mitchell, List. Tex. Sh.; Bartsch, Bull. 91, U.S. INGO cp. 65. OBS.
Distribution.—Nantucket to Colon, West Florida, Matagorda Bay and Mustang Isl., Tex. Pelagic. Also Indian Ocean, Port Alfred, S. E. Africa.
Genus MELANELLA Bowdich
(Eulima) conoidea Kurtz and Stimpson, Pr. Bost. Soc., N. H., 4, p. 115, 1851; Dall, Pr. U.S..N. M.,. 6, p: 336, 0882) Wao Man., 8, p. 273, pl. 49, f. 40, '86; Dall, Tr, Wie weee ee. 159, pl. 5, f. 11, 1890; Singley, 4th Ann. Rept. Tex., p. 336, ’92; Johnson, Occ. Pa. Bost. Soc. N. H1s,(7,.p: onmrong:
Distribution.—Hatteras to the Autilles. Gulf coast: Cedar Keys, Fla.; Galveston and Corpus Christi, Tex. Pliocene, Caloosahatchie beds, Fla.
(Eulima) gracilis C. B. Adams,Contr. Conch., p. t10, 1850; Dall, Bull 7; (US uN NS pe 2 oreo:
Distribution._-Hatteras to St. Thomas, W. I., West Fla. and Yucatan. Pliocene, of the Caloosahatchie.
(Eulima) intermedia Cantraine, Diagn, Moll., Bull. Ac. Brux., Pp. 14, 1835; Tryon, Man., 8,.p. 274, pl. 69,85 43) se, eau Bull.37; U.S.INWML) p. 126, pl: 52, faaawiee:
Distribution.—England to the Canary Isls., New England to West Florida and Barbados.
(Eulima) jamaicensis C. B. Adams, Pr. Bost. Soc., N. H., 2, p. 6; 1845;’ Tryon, Man, 8, p.°273, pl: 60,'f.. 4 eompae BulloMevC. 2. 18, (e277, 1589:
Distribution.—Gulf of Mexico, Blake station 20, in 220 fms. off West Fla. Also Barbados.
(Eulima) subcarinata d’Orbigny, Moll. Cuba, p. 217, pl. 16, f. 4- 6; Tryon, Man., 8, p. 273, pl. 69 f. 44, 1886;, Dalla Bales UL SANs M:,) p, 126,.7 80)
106 MOLLUSCA OF THE GULF OF MEXICO 76
Distribution.— West Florida to Haiti.
(Eulima) gibba De Folin, Les Me’le’agrines, p. 64, pl. 6, f. 4,’67; Poouyvian., 8p, 272) pl, 69). 1.35, °87; Dall, Bull. 37, Se ML. p.126).."8o;
Distribution.—Cape Hatteras to Campeche.
(Liostraca) bilineata Alder, Moll. Northumberland and Durham; wbovom. Man. 8) p. 279, pli 70, f: -72-74,-1887; Dall, Bull. Bp, £20,789.
Distribution.—Norway to Mediterranean, and West Florida GON Eat ti,
(Liostraca) hemphilli Dall, Pr. U. S. N. M., 6, p. 330, pl. to, f. Aerie. oul. 37.) U5. N./ Me pior26, ple 48, £: 11.789.
Distribution.—Cedar Keys to Marco, West Fla.
(iiostraca fisus Dall, Bull.M. C. Z., 18, p. 329, pl.19, £. 11b, open Ue). IN: M.. p..i126,) pl. 19, £11 by. 780:
Distribution.—Fernandina, East Fla., to St. Kitts, W. I. Gulf of Mexico, Yucatan Strait, 640 fms. Also off Hava- na, 400 fms.
Genus NiISO Risso
splendidufa Sowerby, Pr. Zool. Scc., p. 6, 1834, (Audima); A. Edams hes, Conch), /Vzs0,,p. 801, No. 4, pl, 170, 1.'8,°' 54: figron Mam:.18).1287,. pl. 71; f. 124, °86; Dall, Bull. 37, Pepe NN ep. Teg. Sor Pr Us S. Nw Mi, 37) pi.224, IQIO.
Distribution.—Cape Fear, N. C. to Grenada, and Gulf of Mexico, between Mississippi delta and Cedar Keys in 111 fms. Also Pacific Ocean, Sta. Elena, Guayaquil Bay, Ecua- dor (Type locality), in 6-8 fms. (Cuming).
aeglees Bush, Rept. U. S. Fish Com., p. 83, 1883-'85; Dall, Bull. Ga Se NY Me p..128) pl..41, f. 10a, 789, Cas zzterzupia Sowerby var. ).
TG; BULLETIN 38 Io7
Distribution.—Hatteras to Tampa, West Fla., 7-32 fms. Pleistocene, Knapp’s No. 3 well, Terrebonne Parish, La., at 1150-1200 feet.
Genus PYRAMIDELLA Lamarck
dolabrata Linnzus, Gmelin, Syst. Nat. p. 3585, No. 113, (as Trochus):: Tryon, Man., 8, p. 300, pl. 72) ¥) 71-72 ea Bully 27. USN ME. rae oa.
Distribution.—Sarasota, West Fla. to Barbados.
(Longchaeus) crenulata Holmes, Post-Pl. Fos. S. C., p. 88, pl. 13, f. 14, 14a, 1859, (as\Obelzscus); Dall, Bulli (iy aes.
Pp. 331, 1889; Tr. W. 1. S., 3, p. 247, 1892; Vamabtamien N.S. Philay 3.4758), 1Go2:
Obeliscus tesellatus Dall, Pr. U. S. N. M., 6, p. 330, TasauiNior of Adams, 1855. For full synonymy see Dall, 1889.
Distribution.—South Carolina to St. Thomas, W. I. Gulf coast, Cedar Keys, Tampa Bay. Fla., and Galveston, Texas. Pleistocene, North Creek, West Fla., and of the Lake
Borgne borings, La. Habitat, grassy or muddy flats to 2 fms.
(Longchaeus) candida (Meuschen) Moerch, Mal. Blatt., 22, p. 158, (as Obeliscus): Dall, Bull. 37, U.S. N.. M.) etree see: PW Oa eeepc T) lSO2.,
Distribution.—Hatteras to Barbados and West Florida. Pleistocene of Texas.
(Syrnola) smithi Verrill, Pr. U.S. N. M., 3, p. 380, 1881,(as Zur- bonilla): “Yr. Conn, Acad., 5, p.538, pl. 98p4tp meee: Bartsch, Pr. Bost. Soe, N. H., 34, p. 71, \pke aap a ea: 1909; Johnson, Occ. Pa. Bost. Soc. N. Ho+7; 92, ca snehioe of Tryon.
Fulimella unifasciata Dall (pars), Bull. M. C. Z., 18, p. 338, pl. 19, f. 11¢,. 1889; Bull. 27, U.S. N. M., p) 130) pleas 11c, 89. Not of Forbes (Rept. Aegean Inv., p. 188, 1843) which is a European shell.
108 MOLLUSCA OF THE GULF OF MEXICO 78
Distribution.— Massachussetts to Barbados and West Fla.
(Syrnoja) thelma Dall, Pr. U. S.N. M., 46, p. 254, pl. 20, f. 3, IQ14. Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., aty4o feet.
Genus TURBONILLA Risso
exilis C. B. Adams, Contr. to Conch., p. 74, 1850, (as Chemnit- oe an Bulle UW, So Ns) Miu p. 28)? 89°"Tr.. We LT S,, Be Pe i250, 92. Distribution.—Hatteras to Haiti and West Florida,3-63 fms. Pliocene of Caloosahatchie and Shell Creek, Fla.
Helotheca Wall,Bull. M. C. Z./ 18; p. 335, pl. 26, f. 7d, ’89; Bull. Bae Ne Mo, p. 128,’ pls 26,'f. 7d,’ 80.
Distribution.—Gulf of Mexico, Blake station 32, at 95 fms., west of Florida, 50 fms. Also Barbados, 100 fms.
interrupta Totten, Am. Jour. Sci., (1), 28, p. 352, f. 7, 1855, (as Geguera) Wall Bull, 37,°U. S. N. M.,..p.1128, pl. 26,.f. PMNS se lta VW) L: Os. 3,"Ps 259, 92; Bartsch, Pr: Bost. Pocw Ne UH 24) ps7. plea, f. 18) 23; 1900; Johnson, Occ. ar peosts soc! INOUE 7. p95, 191s.
uegaavin, Dall iPr. U..S.N. Mi,.6, p. 332, 1883.
Distribution.—Nova Scotia to Barbados. 2-107 fms. Gulf coast: Cedar Keys, Fla.; Galveston and Corpus Christi, Tex. Pleistocene, Lake Borgne borings, La., Knapp’s wells, Ter- rebonne Parish, No.2 at 1050-1790; No. 3 at 1150-1839 feet, also of North Creek, West Fla. Pliocene of the Caloosahat- chie beds.
fexaitis Kurtz, Cat. Rec. Mar. Shells, N.and 8S. Car., p. 8, ’60, faswencmiiieca yn Dall (er. W. L. S22; ps 257, 1892:
Distribution.—Charleston, S. C., to Marco, West Fla. Pliocene, Caloosahatchie beds.
i = =n +
79 BULLETIN 38 bcerze)
obeliscus C. B. Adams, Contr. Conch., p. 72, 1850, (as Chemnztt- zia); Dall, Bull. 37, U. S..N. M., p. 128,,7 80,3) re eee Bip 2Omn oa: Distribution.—Hatteras to Jamaica. 12-63 fms. Gulf coast,
West Fla. and Galveston, Tex. Pliocene, Caloosahatchie beds, Fla.
virga Dall, Pr. U.S. N. M., 6, p.'332, 1883: Bull a7) ee VE NO: AD28 Wie Distribution.—Hatteras to Florida Keys and Cedar Keys, Fla. 2-15 fms. Pliocene, Caloosahatchie River.
punicea Dall, Pr. U. S. N. 'M., 6, p. 332, 188324 De. Wye ee 3h De ZO MSG 2:
Distribution.—North Carolina to the Bahamas. 2-31 fms. Gulf coast, Cedar Keys, West Fla. Pliocene, Caloosa- hatchie beds, :
curta Dall, Bull, M.C. Z., 18, p. 337, pl. 26,1. 7e) teen ae SN pe mes pl 26, ci eae.
Distribution,—Yucatan Strait, 640 fms. Also off Cape Hat- teras and Antilles.
speira Ravenel, Pr. Elliott Soc. Nat. Hist, 1, p. 280, 1859, (as Chemnitzta) ;Holmes, Post-Pl. Fos. $.C.; p32) phi aging: 1a) 1859: Dall, Prowl. S.) 3) p..258, aeee
Distribution.—Pleistocene, S. Carolina, and #de Hilgard, of the Lake Borgne, borings, La. Pliocene, Coloosahatchie beds (?). Not reported in the recent fauna.
reticulata C. B. Adams, Contr. Conch:, p. 75, 1850: Dally Wa. WT Soap. GON wigan
Distribution.—North Carolina to Jamaica, o-60 fms. Re- ported by Singley from Galveston, Tex. Pliocene, Caloosa- hatchie beds, West Florida.
conradi Bush, Vanatta, Pr. A. N. S. Phila., 55, p. 758, 1903.
XIO MOLLUSCA OF THE GULF OF MEXICO 80
Distribution.—Cited by Vanatta from Calhoun Co., North- west Florida.
(Parthenia) cedrosa Dall, Pr. U.S. N. M., 6, 1883; Bull. 37, SUN.) Pp.’ E30; pl. 48, £.4,° 1889.
Distribution. —Cedar Keys, West Fla.
(Careliopsis?) styliformis Moerch, Malak,Blatt., 22, p. 169, 1874, DM Mea MCs Zi4. 18y pi 338, 1889; Bulls (37,0. S$.) N. NOT Os), SO.
‘Distribution.—Hatteras to Sarasota Bay, West Fla. Also St. Thomas, W. I. 2-52 fms.
Genus ODOSTOMIA Fleming acutidens Dall, Pr. U.S. Nat. Mus., 9, p. 331, 1883.
Distribution.—Hatteras to Cedar Keys, West Florida, 2-107 fms.
Note.—In 1892 (Tr. W. I. S., 3, p. 250), Dall regarded this shell as a variety of the European Miocene to Recent species, O. conotdea Brocchi (Conch. Foss. Subapp.,2, p. 659,pl. 16,f. 2, 1814). The latter species is recorded by Harris from the Pleistocene of the Galveston well at 440-458 feet.
impressa Say, Jour. A. N. S. Phila., 2,p. 244, 1822; Dall, Pr. Ce SeNY Nt 2. 6) pL 3st, 18835 Bull. 37; U.S: No M., p. 130, pleg2,f, 11,89; Singley, ath. Ann. Rept. Tex. G, S., p. Goroe wbartsch', Pr. Bost) Soc. N., H.,~34;. pl./13,' Ff. 51, REGO; Johnson, Occ. Pai Bost. Soc: N. H.,-7, p..98, 1915-
Distribution.—Massachussetts to Florida. Gulf coast: Tampa and Cedar Keys, Fla., Corpus Christi, Tex. Plio- cene, Caloosahatchie beds, Fla.
seminuda C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 280, pl. 4, MemawtS2o. (as /azi2a); Dall, Bull: 37; U.S. .N.M., p. MeO. pl. 52, 1.10, 89: Bartsch, Pr. Bost. Soc, N.H., 34, p.
er BULLETIN 38 Tit
97, pl. 13, f. 45, 48, 1909; Johnson, Occ. Pa. Bost. Soc. N. EL 5 7; P:\97,) LOLS:
Distribution.—Massachussetts to Florida Keys and West Florida.
Genus PERIST{CHIA Dall
toreta Dall, Bull. M,C. Z., 18, p. 340, 1889; Bull) 47) Wises M..p."130, plie4zi ts Toner
Distribution.—Cape Lookout, N. C., to Key West and Charlotte Harbor, West Fla. 2-22 fms.
agria Dall, Bull. M. C. Z., 18, p.’ 340, 1880; Bull, 49°00 ah Mp. 20,550!
Distribution.—Marco, West Fla. o-2 fms. Also Key West and off Hatteras.
Genus ATLANTA Lesueur
peroni Lesueur, Jour. de Physique, 85, p. 390, pl. 2, f. I, 2, 1817; d’Orbigny, Voy. Ame’r, Me’rid.,p, 171; pl) c2taee) 1836; Dall, Bull. 37, U. S. N. M., p. 136; (pl ae eee, pl. 66, f. Ir0a, 1889; Pr. U.S: N..M:, 37% pizegtaane- Bull) 112) USN. Mo ps 133. moor,
Distribution.—West Atlantic, Lat. 42° N. to Tropics. Gulf of Mexico, West Fla. and Texas. Also eastern Pacific. Pelagic.
Genus OXYGYRUS Benson
keraudreni Lesueur, Jour. de Physique, 85, p. 391, pl. 2, 1817; H. and A. Adams, Genera Rec. Moll., 2, p. 92, 1858, 3, pl. 69, f. 6, 6b; Verrill, Trans. Conn..Acad., 6). p ago i7es5- Dall, Bull. 37, U. S. N..M., p. 136, ’89; Johnson) (eeu ka: Bost? Soc. IN as 7) pio, rons.
Distribution.—West Atlantic, Lat. 39° N.to Tropics. Gulf of Mexico, West Florida and Texas. Pelagic.
le
X12 MOLLUSCA OF THE GULF OF MEXICO 82
Genus SIMNIA Risso
(Neosimnia) intermedia Sowerby, Spec. Conch., No. 1, p. 9, f. 32, 33, (as Ovulum), Thes. Conch., Ovulum, No. 40, p. 479, pl. HOON t OF, 62,1848; Reeve, Conch. Icon, Ovyluzz, pl. 7, f. goa, 1605: Tryon, Mani, 7, ‘p..251, pl. 3, °£.:85-86, (as Cova Mall Biull, MC) Z., 18, p. 235, 8889; Bull. 37, U- SoINY IM: p. 134, So.
Distribution.—Hatteras to Brazil. Gulf coast, Vera Cruz. 15-170 fms.
(Neosimnia) uniplicata Sowerby, Thes. Conch., p. 478, pl. 100, f. 30-32, 1848,(as Ovulum); Reeve, Conch. Icon., Ovulum, pl. Seema. 1865°)-Lryon) /Man),--7; p. 254, pls, £. 30, 40; feee Wall Buallies7) ps 134) '89;, Bull, r12,.U. 5S. N. M., p. £30, 1921.
Ovulum subrostratum Sowerby, 1848, antillarum and arcua- tum Reeve, 1865. carolinensts Moerch, 1877.
Amphiperas canadiensts Moerch, 1882 (typ. err. for carolinen- SES.)
Distribution.—Hatteras to Barbados and West Florida. 12- 121 fms. Also Pacific coast, San Pedro, Cal. to Lower Cal- ifornia.
Genus CYPHOMA Bolten
gibbosa Linnzeus, Syst. Nat., ed. X, p. 726 1758, (as Bulla); Reeve, Conch. Icon., 15, pl. 7, f. 32, 1865, (as Ovulum) ; iiwvou. Man. 7. p. 2505 pl..3, f, 81, 82,-1885; Dall, Bull. 37, i S.N)- M..p. 134, °89; Dall and Simpson, Bull. U. S. Fish Com., I, p. 420,1901, (as Ultemus).
f Distribution.—Georgia to Trinidad, 15-50 fms. Also West Florida.
83 BULLETIN 38 Itz
Genus PEDICULARIA Swainson
decussata Gould, Pr. Bost. Soc., N. H., 5, Pp. 127)) TSR eeadai. Bull. M. C.Z., 18, p. 237, pl. to, £. oa-b,) 88a iieiiny ay) U.S. N.M., piira4. pl 1o)-f gap eer
albida Dall, Bull, M. C. Z., 9, p. 39, 1881.
Distribution.—Georgia to Barbados, and Yucatan Strait, 640 fms.
Genus CYPRAEA Linnzeus
Spurca Linnzeus, Syst, Nat., ed. X, p. 724, 1758; Reeve, Conch. [con.. 3, pl. 14, f. 68, 1845; Tryon, Mani,.7° p) noses ong) f. 16; 17, 1885; Dall, Bull. 37, U, So Noa Pp. 136) eer
Distribution.—Cedar Keys, West Fla., to Barbados, 0-2 5 fms.
exanthema Linnzeus Syst. Nat., ed. XII, p. 1172, 1767; Reeve, Conch. Icon.,:3, pl.-5, f. 16, 1848; Tryon) Mam, vey ow
pl. 1, f. 4, 5, 1885; Dalland Simpson, Bull. U. S. Fish Com., 1\'p;, 420,) Toor.
Distribution.—Hatteras to Colon, West Florida and Texas. o-10 fms.
cinerea Gmelin, Syst. Nat., p. 3404, 1792; Reeve, Conch. Icon., 3, pl..22, f. 124,,1846; Tryon, Man., 7, p. 16G)ipl aie, 16,285; Dall, Balliia7,. UsS. IN. M., pi agerieer
Distribution.—Hatteras to Guadeloupe, West Florida and Texas. o-163 fms.
Genus TRIVIA Gray
pediculus Linnzeus, Syst. Nat., ed. X, p. 724, 1758,(as Cypraea); Reeve; Conch. Icon:, 3, pl: 23, f, 131, 1846; Tryon}iManii 7.
p. 201, pl. 21, f.'94,'95,/ 1885; Dall, Tr, (Wai aes 168, go?
—
WT4 MOLLUSCA OF THE GULF OF MEXICO 84
Distribution.—St Augustine, Fla., to Barbados and West Fla. Pliocene, Caloosahatchie beds.
suiusa Gray, Descr. Cat. Cypr. p. 16, 1832; Tryon, Man., 7, p. Pee mie2tet, 1. 2. T885° Dall, Bull, (Mi C. Z., 18) p. 240, 1880.
Distribution.—Cedar Keys, West Fla., to Barbados. 1-3 fms., on reefs. Pliocene, Caloosahatchie beds.
candidula Gaskoin, Pr. Zool. Soc., p. 200, 1835; Tryon, Man., Meeees) pl. 22). 33. 24..° 85: Dall; Ball. MC. Z., 18, p: ZaQ weg nicl. 37, .W. 5. NeiM:,: p. 136,780:
olorina Duclos, approximans Beck. Distribution.—Hatteras to Barbados. 18-140 fms. Gulf of Mexico, Yucatan Strait, 640 fms.
globosa Gray, Descr. Cat. Cypr., p. 14, 1832; Tryon, Man., 7, Deeo wpe amet o2-92,..7853) Dall, Ball. Mi Gi Zs, 18. p. Pao: mil. 27, Us. N.'M., p. 7136, 780;
pilula Kiener, sphaerula Mighels.
Distribution.—Gulf of Mexico, U. S. Fish Com. station 2373, west of Fla., at 23 fms. Also Key Westand Antilles. Pliocene, Caloosahatchie beds.
Genus ERATO Risso
maugeriae Gray, In Sowerby, Conch. Ilustr., p. 17,pl. 7, f. 47, mez lion. Man) 5,\p.9,\ pl. 4, f. 42, 43,83; Dall, Tr. Pere 3 253, Pp. 168, 1890.
faeces immons, N. Car. Geol: Rept., p. 262, f. 139, 1858. Not of Donovan.
Distribution.—Hatteras to Antilles and Colon; West Fla. and Vera Cruz. 0-63 fms,
Genus BURSA Bolten affinis Broderip, Pr. Zool. Soc., p. 179, 1832, (as Ranella);
oO
85 BULLETIN 38 IIs
Tryon, Man,, 3, p: 42, pl. 22, f. 38, a8812 alma U. 5. NoM.,; p. 132, (80; (as Gyxezewney,
Distribution.—Hatteras to the Antilles. Variety cubaniana a’ Orbigny, dredged on Arrowsmith bank, Yucatan, also off Key West and Cuba.
Genus DISTORTIO Bolten
reticulatus Link, Beschr. Rostock Samml., 3, p. 122, 1807, (as Mistoririx); Dall, Bull. 37, U: S. N. Mi, paiiag eee and Simpson, Bull. U, S. Fish Com., 1, p. 416, 1901; Mitch- ell, list Tex: Shells owia,
Distribution.—Hatteras to Barbados, West Florida and Mustang Island, Texas, 22-154 fms.
reticulatus var. clathratus Lamarck, An. s. Vert., 7, p. 186,1822,, Inc. Me’th., pl. 413. f. 4, (as Trzton); Dall and jaimipeou: Bull) U.S. Bish Conny pillars ream
Distribution.—Porto Rico and Gulf of Mexico near Key West (dredged by U.S. Fish Com.), also at Chandeleurs, La. Pleistocene, New Orleans pnmping station No. 7.
Genus COLUBRARIA Schumacher
lanceolata Menke, Synopsis, p. 87, 1828, (as Ranella); Reeve, Conch. Icon., 2, pl. 18, f. 79, 1844, (as 2720”) ¢ intra: Conch. Cab., pl.65, f..8; Tryon, Man, 3) pi 27s 162, 1881; Dall, Bull. 37,.U..S. N. M., 9p. iue2) senna and Simpson, Bull. U.S. Fish Com., 1, p. 416, 1901.
Distribution.—Hatteras to Barbados and Vera Cruz on the Gulf coast. Genotype.
testacea Moerch, Yoldi Cat., 107, (as77zton); Mal. Blatt., 24, p. 25; Tryon, Man., 3, pl. 14, f. 128, 1881; Dall Geen a S. N. M., p. 132, 80:1 r. W. 1. S..,..'3, Dp.) TOM
Distribution.—Hatteras to Sombrero, and Texas region of the Gulf of Mexico. Pliocene, Caloosahatchie River, West Florida.
r16 MOLLUSCA OF THE GULF OF MEXICO 86
Genus CYMATIUM Bolten
olearium Linnzeus, Syst. Nat., ed. XII, p. 1216, ( 77iton); Reeve, Moueny tcon!, 21, 32, 1844: Tryon, Man., 3, 7p. 11, pl. 5, f. emer Dall, Bull, 37, Ui SN.’ M.; p. 132,80, (as Lam- used); Bartsch, Bull. U.S. N.M., p. 94, 1915.
Distribution.—Hatteras to Cartagena, Colombia, and the Texas region of the Gulf of Mexico. Also Indian Ocean, Port Alfred, South Africa.
chlorostomum Lamarck, An. sans Vert., 7, p. 185, 1822, (as 777- Lom ramcicener, con. Coq), Viv:, p, 19, pl: 12, 1. 2; Tryon, Rice ts, pls 7) 147,48, T8813 Dall, Bull. 37, U.S, Mev ps 132) °89:' Dall and’ Simpson, Bull. U.S. Fish Com., I, p. 417, 1901,) as Lampusia).
Triton pulchellus C. B. Adams, Contr. Conch., p. 60, 1850.
Distribution.—EHast Florida to Barbados and Gulf of Mexico along the Mexican coast.
gracile Reeve, Conch. Icon., 77zton f. 58, 1845; Tryon, Man., 3, em2sto7) Fost; Dall, Bull. M. C. Z:, 18, p.'227, ‘pl. 20, f. eee Em 27 2° U.S, IN. Me, 132) “pl. 20, f.. 2; 789,/(as Lampusia).
Distribution.—Cape Catoche, Yucatan, 24 fms., U. S. Fish Com. Also Colon and Barbados. Max. depth roo fms.
cynocephalum Lamarck, An. sans Vert., 7, p. 184, 1822, (as 777- Vou); Reeve, Conch, Icon., 2, pl. 8, f. 26, 1844. Tryon, Man., POA eon pl. Lt, ft. 80,.-pl. 15,.f, 152, 81; Dall, Bull. 27, U. Syne Mp, 122,780; Dall and Simpson, Bull. U.S. Fish Some 1, p. 417, roor, (as Lampusia). Distribution.—Florida Straits to Margarita Isl. ‘Texas re- gion of the Gulf.
Genus NYCTILOCHUS Gistel
femorale Linnzeus, Syst. Nat., ed. X, p. 749, 1758, (as Murex);
87: BULLETIN 38 It7
Reeve, Conch., Icon., 2, pl. 7, £.. 22,) 1644 laa aay Tryon, Man., 3, p. 18, pl. 10, f. 70, "81; Dall Bua S. N. M., p. 132,.’89; Dall and Simpson) ‘Bal Veto Com., 1, p. 417, 1901, Cas Zoroz 22722).
Distribution.—Cedar Keys, West Fla., to Guadeloupe. Genus ASPELLA Moerch
scalarioides Blainville, Faune Francaise, p. 131, pl. 5,f. 5, 6,1826, (as Murex); Dall and Simpson, Bull. U. S. Fish Com., 1,p. AO. TOTO:
Distribution.—European and Antillean.
scalarioides var. paupercula C. B. Adams, Contr. Conch., p. 60, 1850, (as Murex); Dall, Bull. M. C. Z:, 18) po 2680 ee. Bully37, Us. NY MS pror2o)) 8a.
Triton cantraineit Recluz, Jour. de Conch., 4, pp, 246, 418, pl. 8, f. 10, 1853; Kobelt, Jahrb. Mal. Gesell, (4p) gaanimeg a, (as Ocinebra).
Distribution.—West Florida, at 50 fms.; ‘Texas and the Antilles.
scalarioides var. obeliscus A. Adams, Pr. Zool. Soc., p. 269, 1851; Tryon, Man., 2, p. 129, pl.\38, ff) 467, aese pian, Ball 27h SNe Nee. L201 50s Distribution.—Vera Cruz to St. Thomas. Texas region of the Gulf.
Genus OOCORYS Fischer
abyssorum Verrill and Smith, Tr. Conn. Acad. Sci., 6, p. 177, (not the fig, which is O. sulcata), 1884, (as Benthodolium) ; Dall, Ball.) M.C. Z.,)\\18) p. 220, 789; Bull. 370s ee DP) 132,190:
Distribution.—Northern part of the Gulf of Mexico, U. S. Fish Com. station, No. 2400, in 169 fms. Also off Caro- lina. Max. depth 2221 fms.
—>
x78 MOLLUSCA OF THE GULF OF MEXICO 88
Genus CASSIS Lamarck
cameo Stimpson, Silliman’s Am. Jour. Sci., p. 443, 1860; Tryon, Mama 70 D275. pls (2,1. 50, 7°85) Dall, Bull. 37,0U....5.' N. WEG D034, SQ. Distribution.—Hatteras to Barbados and West Fla.
tuberosa Linnzeus,Syst. Nat., ed. X, p. 735, 1758, (as Buccinum) ; ReevenvConuch: [con:,,/5,) pl, 3, 4.7, 1848; Tryon, Man. one pl 271) pleat. 51, 735; Dall Bull. (37,,0.52 N. May py t34, (So; Dall and Simpson, Bull. Ul S. Fish Com., Vs 4ALo,. LOOL. Distribution.—Hatteras to Barbados and West Fla.
(Cypraecassis) testiculus Linnceus, Syst. Nat., ed. X, p. 736, 1758, (as Buccinum); Reeve, Conch. Icon., 5, pl. 4, f. 10, Meee inves, Mati. 7) p. 273, pl. 2, f.64, pl.4;.f. 63, ’85; Dall, Bull. 37, U.S. N. M., p. 134, ’89; Dall and Simpson, Bate oY Nish’ Com., 1, p..418, T90r.
Distribution.—Hatteras to Trinidad, W. I., and the Texas region of the Gulf.
(Semicassis) inflata Shaw, Nat. Misc., 5, p. 22, pl. 959, 1812: piven, Man.,.7,)p. 274, pl:4, f.. 65, ’85; Dall, Bull. 37,. U. SyNM:, pp. 124) °89; Dall and Simpson, Bull U.S. Fish Com., I, p. 418, 1901; Mitchell, List Tex. Shells, p.6; Van- maoaniet. Pe.) NaS.) Phila. ,55;\p. 758, 1903.
Distribution.—Hatteras to Brazil, Gulf coast: Calhoun Co. West Fla.; Point au Fer, Chandeleurs, Lost Island, La.; Galveston, Tex. Pleistocene, New Orleans pumping sta- tion No. 7,Knapp’s No. 2 well, Terrebonne Parish, at 1791- 1342. tect.
Genus ONISC!BIA Swainson
dennisoni Reeve, Pr. Zool. Soc., p. 21, 1842, (as Oniscia); Tryon, Rom wip 282.0 plito. £, 20; °85: Dall) Bull. Mo'Gi"Z.j 108:
Hy) 231),'' 80.
&9 BULLETIN 38 lig
Distribution.—Arrowsmith Bank, Yucatan, at 130 fms. Also Guadelonpe Island. (Type locality).
Genus TONNA Brunnich
galea Linnzeus, Syst. Nat., ed. X, p. 734, 1758, (as Buccinum) ; Reeve, Conch. Icon:, 5, pl. 1, 1848; Tryon, Mani7 sem zer, pl. 1, f. 3,85; Dall and Simpson, Bull;.U. 5S. Bish Comisar, p. 419, 1901. (Allas Dolzum).
Distribution.—Hatteras to Trinidad, West Florida, Louisia- na, Galveston and Matagorda, Texas. Also Mediterranean.
perdix Linnaeus, Syst. Nat., ed. X, p. 734, 1758, (asBucciuum); Reeve, Conch. Icon., 5. pl. 6, f. 9, 1849, (as Dolzum) ; Tryon, Man., 7, p. 264, pl. 3, f. 15, pl: 4, & 23-25) 35; ame om, 37, U. SIND NE) py 34. iso:
Distribution.— West Florida to Brazil.
Genus PYRULA Lamarck
papyratia Say, Jour. A. N. S. Phila., 2, p. 238, 1382242 you, Man., 7, p. 266, pl. 6, f. 35,85; Dall, Tr) Wate ee 163, 90; Vanatta, Pr. A. N. S. Phila., 55, p> 753, 1902:
Distribution.—North Carolina to the West Indies. Gulf coast, Calhoun Co., Fla. and Texas. Pleistocene, New Or- leans pumping station No. 7 and of Fla. Pliocene, Caloos- ahatchie River.
Genus STROMBUS Linnzeus
pugilis Linnzeus, Syst. Nat., ed. X, p. 744, 1758; Reeve, Conch. Icon:, 6, pl. 16, f. 39-43, 1851; Tryon, Man., 7, p.,teG; me, f./13-15,.°85; Dall, Tr. W.1.S., 3, p. 177, °90; (Simalen aes Ann. Rept. Tex., p. 338, ’92; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 422, 1901; Vanatta, Pr. A) Ni Sea
120 MoLLUSCA OF THE GULF OF MEXICO QO
Bo Pe7 50, 1903: Vaughan, Publ. No: 133, Carn. Inst., p. E71) /TOIO.
Distribution.—Hatterasto Colon. Gulf coast; Cedar Keys and Calhoun Co., West Fla.; Point au Fer, Chandeleurs, Cameron, La., Galveston and Corpus Christi, Tex. Pleisto- cene, Grand Chenier, La., New Orleans pumping station No. 7; North Creek, and Labella, West Fla. Pliocene, Ca- loosahatchie River.
costatus Gmelin, Syst. Nat., p. 3520, 1792; Tryon, Man., 7, p. mose ress; Dall, Bull. 27,0. 5. N: M., p. 136, °89; Dall, and Simpson, Bull. U.S: Fish Com., 1, p. 422, 1901.
aecipurinus Reeve, Conch. Icon., 6, pl.'7, f. 12, 1850; Tryon, iManennger pl 1, 1. 7, 85. Distribution.—St. Augustine, East Fla. to Guadeloupe, W. I. Also West Fla.
Genus SEGUENZIA Jeffreys
monocingulata Seguenza, Boll. Comitato Geologico, 7, p. 188,’76; Miatson, Chall: Rept. Gastr:, p. 105, °86; Dall, Bull. M. C. epoch p 208, 69° Bull.a7, Uo. N. M., p./142; pl. 62,, f. 88, 89, 1880.
formosa Jeffreys, Rept. Valorous Cruise, p. 200, 1876; Dall, awe C.Z.; 9; Pp. 268, 89.
Distribution.— Western North Atlantic and Bay of Biscay. Gulf of Mexico, West Florida and Texas regions. 100-2033 fms. |
trispinosa Watson, Jour. Linn. Soc., 14, p. 591; Chall. Rept. Socom 5) Plight. 4.1886; Lryon, Man., 9, p. 47,. pl: Bie FOS, 1887, Wall. Bull.. 37, U..S. N. M.,.-p. 142, 1889. Distribution.—Hatteras to Pernambuco, Brazil and the Tex
region of the Gulf. 294-675 fms.
oe BULLETIN 38 Tat
iconica Watson, Jour. Linn. Soc., 14, p. 589, 1879; Chall. Rept. Gastr., 15; p. 107, pl: 7, f. 3, 1886> Tryon iawn, 47, pl. 8, £. 77, 78, 787; Dall, Bull. Ml Couz ia: 1889.
Distribution. —Gulf of Mexico, Florida Strait region, Culebra Isl., W. I., and off the Azores. 390-1568 fms.
Genus TRIPHORA Blainville
perversa var. nigrocincta C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 286, pl. 4, f. 11, 1839, (as Cerzthtum); Gould, Inv. Mass., Pp. 323, f.. 592, 1870; Dall, Pr."U. S: Ni MiG) eee Bull. 37, U.S. N. M., p..138, ?89; Singley; 44 Ann) Rent, G. S. Tex., p. 338, 792: Dall, Tr. Wood, Sap ieea ee Johnson. Occ. Pa, Bost. Soc. N.. H.,_7;. pa 2G eee
Distribution.—Massachussetts to Barbados. 0-30 fms. Gulf coast: Cedar Keys, Florida, Corpus Christi, Texas. Plio- cene, Caloosahatchie beds, West Fla.
decorata C. B. Adams, Contr. Conch., p. 177, 1850, (as Cerith- tum); Dall, atriw. E.1S., 3,99: 265, 192.
Distribution.—Gulf of Mexico, north and east shores, and the Antilles. Also Lower Miocene of the Chipola marl, West Florida. Variety olzvacea Dall, (Bull. M. C. Z., 18, Pp. 244; 1889; Bull. 327, U.S. N. M.,. p. 128) 7 Se) eae a Mexico, west of Florida, in 50 fms. Also Key West and Antilles.
colon Dall, Bull. M. C. Z.} 9, p. 86, 1881; 18, p. Zag pi ae ae 12,89; Bull.\37, Uo SAN. \M.,'p. 138) pl. 20; Ga geeiee:
Distribution.—Yucatan Strait, 640-1002 fms. Also off Havana, Cuba, 450 fms.
triserialis Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 246, pl. 20,
Sh
» Bea MoLLusca OF THE GULF OF MEXICO 92
meea, 16807, Bill) 37) Ul SiN. M.,:p. 138, pl. 20, f. 5a, 6a, 1898.
Distribution.— Yucatan Strait,640 fms. Also off Barbados, 154 fms.
bigemma Watson, Jour. Linn. Soc., 15, p. ror, 1880; Dall, Bull. Pee Aro. p.' Si." T88E- Watson, Chall. ‘Rept. Gastr., p. Beeld t) 6,785: Dall. Bull. M. C.’Z., 18, p: 248, °89.
Distribution.—VYucatan Strait, 640 fms. Also off St. Thom- asyW iL. 390 fms.
meus) Dall Bull. M. C. Z., 9, p. 83, 1881; 18, p. 240, pl. 20, f. boon bull. 27, U, SN. M:, p, 138) pl. 20, 1. 11, 789.
Distribution.—Yucatan Strait, 640fms. Perhaps a variety of digemma Watson.
a@urupta Dall, Bull. M. C. Z., 9, p. 84, 1881; 18, p. 249, ‘pl. 20, f. Geo mpulis: 37,0U.. 5S. N. M.,. p: 138, pl.2o, f.:9,.789.
Distribution.—Yucatan Strait, Cape San Antonio, 640 fms.
sorucula Dall, Bull. M.,C. Z., 9, p. 82, 1881; 18, p. 249, pl. 20, imeumov ceo. momll 37) UW. SecNi Maj p. 138, -pliizo, £.. 11b, 1889.
Distribution.—Yucatan Strait, 640 fms.
inflata Watson, Jour. Linn. Soc,, 15, p. ror, 1880; Dall, Bull. WE Ce. 2) 9, p81, 1881;, Watson, Chall. Rept. Gastr., p. Pomp AO, tit, 85° Dall, Bull. M:'C.., Z. 18, p.. 240, 1889.
Distribution.— Yucatan Strait, 640 fms. Also off Georgia, st. Augustine and Culebra Isl. 294-640 fms.
inflata var. ibex Dall, Bull. M. C. Z., 9, p. 86, 1881; 18, p. 249, pini20, £. 12b, 89.
Distribution.— Yucatan Strait, off Cape San Antonio, in 640 fms. Also off Havana, 450 fms.
93 BULLETIN 38 ¥23
cylindreila Dall, Bull. M. C. Z., 9, p. 83, 1881 18: po 250) pireo, f. 6, °89;.'Bull. 37, U. S. N. M., p: 138, pligerme sane
Distribution.—Cape San Antonio, Yucatan Strait, 640 fms.
Genus CERITHIOPSIS Forbes and Hanley
greenei_C. B. Adams, Jour. Bost. Sec. N. H., 2)p, 287. ipiaw 12, 1839, (as Cevzthzum); Dall, Bull. 39. U.S: ie: 138, pl. 52, f. 2, °80; Tr..W. 1. S., 3, p:)260)"e2a sg emaean: Occ. Pa) Bost: Soc, INCE 73 peo mens:
Distribution.—Prince Edward Island to Dominican Repub- lic and west of Cedar Keys, Fla., Corpus Christi and San Antonio, Texas. 3-10 fms. Pliocene, Caloosahatchie River, Florida.
crystallina Dall, Bull, M. C. Z.,-9, p, 89, 1881+-18,/pi 25a 0muese, f. 3, 789; Bull. 37, U.S. N. M., p. 138, ‘plage nee a ’89; Dall and Simpson, Bull. U.S. Fish ‘Com, a pnaea: IQOl. Distribution.— Washed ashore dead, Cedar Keys, Florida, dredged alive west of Florida in 50 fms. Common in An- tillean dredgings. 50-805 fms.
martensi Dall, Bull. M. C. Z., 18, p. 255, pl. 20, f. 2, ’89; Bull. 27) Ui os NZ aM o: 638) pl. 20. se 2 aor
Distribution.—Gulf of Mexico, west of Florida, at 1181 fms.
vanhyningi Bartsch, Pr. Biol. Soc. Washington, 31, p. 135, TOPS: Distribution.—Tampa Bay, West Fla.
burkevillensis Dall, Pr. U. S. N. M., 46, p. 231, pl. 22, f. 5, IQI4. Di stribution.—Upper Miocene or Lower Pliocene, Burke- ville, Texas.
Pe, Oe oe Re ie gee eee ee Se
- tot ae
ios cere oes
124 MOLLUSCA OF THE GULF OF MEXICO 94
(Eumeta) subulata Montagu, Test. Brit. Suppl., p. 115, pl. 30, iweweeos, (as A7urex):) Dall, Bull, M.-C. Z.,.18, ps 252, pl. ein oo: Bill. a7 Ua SUN.) MM. p. rao} pl.i20; f.4, Peet oot }ohnson.: Occ, ‘Pa. ‘Bost, Soc.: N. Hs,7, p. 126) LOS.
emersoni C. B. Adams, Jour. Bost. Soc. N. H., 2, p. 284, pl. 4, f. 10, 1838, (as Cerithtum) Verrill, Inv. Vineyard Sound, p. GAS mpl. 24. f. 151, 1373:
punctatum Philippi, 1848. Not of Bruguiere, 17809. - Distribution.— Massachussetts to Grenada, W.I. Gulf of Mexico, Cedar Keys, Texas and Yucatan. 2-15 fms. Plio- cene of the Caloosahatchie.
Genus SE{LA A. Adams
adamsi H. C. Lea, Tr. Amer. Phil. Soc. 2d ser. 9, p. 42, 1845, tasu@ereeune): Dall, Tr. W. 1..S.,3; p. 267,.1892; Vanatta Pee SL ila), 55), px 758, 1903; Jolnson Occ: Pa. OE SOC IN. Ely )i'7, Pp. 127, 19L5.
ynecurals ©. RB. Adams, Jour. Bost. Soc. N. H.,:32, p. 320, pl. Btn ito4o (as Cerzihkium); De Kay, N: Y. Moll, p..130, Creotvi7 2.42) Dali Ball MC. Z., 18, p. 250) 7863 Bull. U. See Mo pe 138, pl:.52, f.5,. °89., Not Cerzthinm, zerébrate Lamarck.
Distribution—Massachussetts to Samana Bay, Dominican Re- public. o-20fms. Gulf coast: Cedar Keys, Fla.; Chande- leurs, La.; Galveston and Corpus Christi, Texas, 0-20 fms. Pleistocene, Gulf coast. Pliocene, Caloosahatchie River, Fla. Lower Miocene, Chipola marl, West Fla.
Genus CERITH!UM Bruguiere
floridanum Moerch, Mal. Blatt., 23, p. 114, 1876; Dall, Bull. 37, Meo mNG MA ps 40, .89:\ lr. W. 1. Si, 3, p. 282, pl. 14, 4. mynHo2, 'Vaughan, Publ, No, 133; Carn. Inst.) p. 177; IgIO.
Distribution-—Hatteras to Cuba and West Fla. Pleistocene, Labella, West Fla. Pliocene, Calaosahatchie beds.
95 BULLETIN 38 125
algicola C. B. Adams, Pr. Bost. Soc. N. H., 2, p. 5, 48; Tryon, Man., 9, p. 129, pl. 22, f. 81, 82, °37; Dall)’ Tray, Dp. 252)o2.
Distribution.—Tampa and Charlotte Harbor, West Florida to Jamaica. Between tides. Pliocene, Caloosahatchie beds.
litteratum Born, Mus. Cees. Vind., p. 323, pl. 11, f. 14, 15, 1780; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 425, 1901.
Distribution.—East and West Florida, Antilles and Europe.
muscarum Say, Amer. Conch., 5, pl. 49, f. 1, 1832; Tryon, Man., 9, Pp; 132, pl. 23, f. 12, 13, °87; Dall, Tr. W iS ae Vaughan, 2d Ann. Rept. Fla. G. S., p. 148, 1909.
Distribution.—Florida to Jamaica. Gulfcoast: Cedar Keys and St. Mark’s, Fla.; Chandeleurs, La. Pleistocene, Mana- tee,'Labelle and North Creek, Fla. Pliocene, Caloosahat- chie beds.
variable C. B. Adams, Pr. Bost. Soc. N. H.. 2, p. 5, 1848; Sow- erby, Conch. Icon., 15, pl. 13, f. 91, 1865; Dall alae U. S. N. M., p. 140. 1889; Mitchell, List Texas Shells.
Distribution.—Tampa, Fla.; Galveston, Corpus Christi, Carancahua and Espiritu Santo Bays, Tex. South to the Antilles and Curacoa.
minimum Gmelin, Syst. Nat., p. 3564, 1792, (as Wurex); Tryon, Man., 9, p. 167, pl. 34, f. 9-11, 1887; Dall and Simpsen, Bull. W'S. Fish ‘Coma, p. 426) 100r,
Distribution.—Tampa, Fla., to Guadeloupe Isl.
minimum var. nigrescens Menke, Synopsis, p. 85, 1828; Dall, Bull. 327,,U. 3. N: MEG aAe. |) 80.
Distribution.—Tampa to Venezuela.
eburneum Bruguiere, Ency. Me’th., pl. 442, f. 1a, b; Tryon,
126 MOLLUSCA OF THE GULF OF MEXICO 96
Miaderor p.t2o, pl. 61, £.7h72,:1987;' Wall, Bull. 37, U.S. N. M., p. 140, 1889; Singley, 4th Ann. Rept. Texas Surv., P- 339, 1892. Distribution.—Florida to the Swan Isls. Cited by Singley from Corpus Christi, Texas.
galvestonense Harris, Bull. Amer. Pal, vol. 1, D. 164) pleas £27; ga, 1895. Distribution.—Upper Miocene, Galveston well at 2236- 2920 feet.
Genus CLAVA Martyn
chipolana Dall, Tr. Wagner Inst. Sci., 3, p. 290, pl. 22, f. 8, 1892. Distribution.—Lower Miocene of the Chipola marl, Alum
Bluff, Fla., and of the Bacom well, Mobile, Ala., at 1241 ft., Chipola horizon.
Genus CERITHIDEA Swainson costata Wood, Mal. Blatt., 23, p. 90; Dall, Pr. U. S. N. M., 6, pass, toes; bulk 37, Us Ss: N. M., p. 140, 7809. Distribution.—Tampa, West Fla., to St. Thomas and Ja- maica.
scalariformis Say, Jour. A. N. S. Phila., p. 128, 1825; Tryon, Miami. Too. pli 33. f78.°87> Dall, Bulli a7. U.S. N. Mj Dp. 140, °89. Distribtion.— Georgia to Cedar Keys, West Fla.
varicosa Sowerby, Genera Shells, No. 42, f. 5; Dall, List Came- fom Suells: Bull. M: C! Z., 18, p. 259,89
Distribution.—Texas, Breton Island, Chandeleurs, La., and south to Cuba and Jamaica.
turrita Stearns, Pr. Bost. Soc. N. H., 15, p. 24, 1872; Tryon, amy Ou pts) pl 2a f, 83.87: Dall, Tro W. 12S, '3) p, 290,1892.
97 BULLETIN 38 I27
Distribution.—Cedar Keys, Fla., to the Bahamas. Pliocene of Shell Creek, Fla.
hegewishi Philippi, Mal. Blatt., 23, p. ’89; Zeitschr. fur Malak., p. 19, 1841; Singley, 4th Ann. Rept. Tex.))p) 326, meo2: Mitchell, List Tex. Sh.
Distribution.—Cited by Singley and Mitchell from Matagor- da, Carancahua and Corpus Christi Bays, Texas.
Genus POTAMIDES Brongniart
matsoni Dall, Pr. U. S, N. M., .46,\p. 231, pl, 21h a ean, IQI4.
Distribution.—Well near Alexandria, La., at 49 feet; well at Pine Prairie, La., at 1540 feet; surface near Burkeville, Texas.
Note.—The brackish water molluscan fauna of Burkeville, Texas; Alexandria and Pine Prairie, La., was referred in 1914 by Dr. Dall to the Pliocene. Dr. W. D. Matthew thought the mammalian remains (tibia of a rhinoceros and upper molar of a horse, either Protohippus or Merychippus) indicat- ed a Late Miocene or Early Pliocene age. For discussion of the Burkeville beds (lower member of the Fleming clay) see Dumble, Univ. Tex. Bull. No. 1869,pp. 224-225, 1918. The stratigraphic relation of the Burkeville beds rather sug- gests an Upper Miocene horizon.
matsoni var. gracilior Dall, Pr. U.S. N. M., 46, p. 231, 1914.
Distribution.—Well near Alexandria, La., at 49 feet; well at Paririe Bluff, La.; near Burkeville, Tex.
Genus ALABiNA Dall
adamsi Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37) 03am, M.,; p..140; 893) Dr. We. TD.) S!,° 3, (pp. 276.) @ ae eees Bittium) .
128 MOLLUSCA OF THE GULF OF MEXICO 98
Distribution.—Hatteras to Haiti and West Florida, moder- ate depths. Pliocene. Caloosahatchie River and Shell Creek, West Fla.
cerithioides Dall, Bull. M. C. Z., 18, p. 258, ’89; Bull. 37, U.S. Meee I4o) So. dre We 1. Ss) 3, p..276; pl. 16; £8,925 Dall and Simpson, Bull. U. S. Fish Com., 1, p. 427, 1901. (All as Bzttzum cerithioide).
Distribution.—Cape Lookout, N. C.. to Samana Bay, Do- minican Republic. Singley lists with a question a shell from Corpus Christi, Tex. Pliocene, Caloosahatchie River, Fla.
Genus BITTIUM (Leach) Gray
varium Pfeiffer, Arch. fur Naturg., p. 256, No. 139, 1840, (as Sexuniunm):lryon,Man., 9; p. 152, pl. 29, f. 86,87; Dall, RW ely 5.3, p. 274, °92;° Dall and Simpson,, Bull; U. S. Mish Com!,; 1, p. 426, 1901.
eeupor eine CB. Adams, Pr. Bost. Soc. N)-H}, p..5, 1845; Sowerby’s Thes., p. 876, f. 210, 211, (as Cerzthtum).
Distribution.—Chesapeake Bay to St. Thomas, W. I. Gulf coast: Cedar Keys, Fla.; Chandeleurs, La.; Corpus Christi, Galveston and Laguna Madre, Tex. Pleistocene, North Beach. Fla. Pliocene, Caloosahatchie and Shell Creek, Florida.
Note.—Aittium nigrum Totten, synonym of &. alternatum Say, has been erroneously reported from Cedar Keys, Fla. and Chandeleurs, La. ‘These specimens were probably varium. "True alternatum, tide Dall, does not extend south- ward of New York Harbor.
boiplex Dall, Tr. W. Te San Dei27 50 Pl.v2T, aby G2)
Distribution.—Lower Miocene, Chipola marl, Fla., and of
the Basom No. 1 well, Mobile, Ala. at 1500-1556 ft., Chipola horizon, Aldrich’s collection. A varietal form.
99 BULLETIN 38 I29
galvestonense Harris, Bull. Amer. Pal.,vol. 1, p. 104, pl. 4, f. 8, 1895. Distribution.—Upper Miocene, Galveston well at 2550-2875 feet. Genus MODULUS Gray
modulus Linnzeus, Syst. Nat.. ed. X, p. 757, 1758, (as Zvochus); Dall, Bull. 37, U..S. N. M., p. 142, 1889; Singley; 4tm aan: Rept. Tex. G. S., p. 339, ’92; Dall and Simpson, Ball VU: S.' Bish Com.)t, p. 427, 1901:
lenticularis Chemnitz, Tryon, Man., 9, p. 261, ph 48, £. 91,92, 1887.
Distribution.—Hatteras to Brazil and to Cartagena, Colom- bia. Gulf coast: West Florida, Chandeleurs, La.; Galveston, Corpus Christi, and Espiritu Santo Bay, Tex. Between tides. Pliocene, Caloosahatchie and Shell Creek, Fla. Up- per Miocene, Galveston well at 2552-2871 feet (identifica- tion doubtful).
floridanus Conrad, Amer. Jour. Conch., 5, p. 107, pl. 12,f. 6, ’70; Hilgard, House of Rep. Ex. Doc., 1; pt. 2,) piies7amoge. Dall, Pr. U.S. N. M., 6, p. 335, 83; Bulle 2G aeons M.,’ p. 1442, 789; Tr. W. I. S., 3; ip. 205,%92.y eee Pabl., No.132, Carn: inst., p.. 171, 1910! Distribution.—Hatteras to St. Thomas, W.I. Gulf coast, Cedar Keys and St. Mark’s, West Fla. Pleistocene, Lake Borgne borings, La., and North Creek and Labelle, Fla. Pliocene, Caloosahatchie beds, Fla. and of the New Orleans Gymnasium club well at 1200 feet.
Genus PACHYCHEILUS Lea anagrammatus Dall, Proc. U. S. N. M., 46, p. 232, pl. 21, f, 5, 8, 1914. Distribution.—Upper Miocene or Pliocene. Well near Alexandria, La., at 49 feet. Also near Burkeville, T cxas,
30 MOLLUSCA OF THE GULF OF MEXICO 100
surface exposure. Brackish water formation.
satillensis Aldrich, The Nautilus, 24, pt. 11, p. 132, pl. 8, f. 1, Haewwrem, (as) Lotamzaes );) Dall, Pr: UW) SiN. M.;. p. 232, IgI4, Distribution.—Upper Miocene or Pliocene. Satilla River, Ga., and Burkeville, Tex. Brackish water formation.
smavis) Wall, Proc. U; SS. N. M., 46, p. 232, pl. 21, £..6, 9, 1614.
Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet, and well, Pine Prairie oil field, La., at 1540 feet. Also surface exposure, Burkeville, Tex.
Genus CAECUM Fleming
floridanum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851; Mcvony meade 8p. 2K7) pl.c66,.f.,,62,\'86; Dall, Bull, 37, ee NaMi) oO) 142, °89;. Fr, Wy L.'S:3, p. 208,' 92.
wreewlore ae Bolin, Jour. de Conch., 15, p. 46, pl..3, f. 6, 1867.
annulatum Emmons, Rept. Geol. N. Car., p. 274, f. 190,’ 58. Not of Brown, 1844.
Distribution.—Hatteras to Bahia, Brazil. Also West Fla. 2-18fms. Pliocene, Caloosahatchie, Fla.
instructum de Folin, Fonds dela Mer; Dall, Bull. 37, U.S. N. Miaip. 142, 780. Distribution.—Hatteras to Tampa, West:-Fla. Pliocene to
Recent. bipartitum de Folin, Fonds dela Mer; Dall, Bull. 37, U.S. N. M., Pu t42.)’ So.
Distribution.—Hatteras and West Florida.
Carolinianum Dall, Trans. Wagner Inst. Sci., 3, p. 300, pl. 22, f. Ssirog2:) Bull. .27,) U.S. N. Mp. 142; 1889, (name only).
IOI BULLETIN 38 131
Distribution.—Hatteras to Tortugas and to Egmont Key, West Fla. 2-63 fms. Pliocene, Caloosahatchie River, Florida.
glabrum Montagu, Test. Brit., 2, p. 497,1803, (as Dentalium); Tryon, Man., 8, p. 215, pl. 66, £. 57, 1886;) Dali gaanine 7, U.S. Ne aps aa so)
Distribution.— North Carolina to Tampa, West Fla. Also European. Pliocene of the Caloosahatchie and of England. Miocene of Yorktown, Va.
cooper! Sanderson Smith, Ann. & Lyc. N. Hist. N. Y., 7, pp. 154, 168, 1862; 9, p. 3093, f.' 3, 1870; Tryem,) iam: 221, pl. 67, -f. 84, °85; Dall, Tr. W. 1S.) 2oiizee usa: Vanatta, Pr. A. N. S. Phila., 55, p. 758, 1903, Joumeon, Occ. Pa. Bost. Soc. Ny EL )73) pi tea) ose
costatum Verrill, Am, Jour. Sci, (3) 3, p..283,. pieutno, LO72.
Distribution.— Massachussetts to the Antilles. Gulf coast: Crooked Island, Calhoun Co., West Fla. Pliocene of the Caloosahatchie and Shell Creek, West Fla.
puichellum Stimpson, Pr. Bost. Soc. N. H., 4, p. 112, 1851; Shells of New England, p. 36, pl. 2, f. 3; Hilgard, House ot Rep: Ex. Doc., 1, pt.'2, p. 886, 1878: “Tryon, Mant Ss paen 7 pl. 66, f. 63, °86>' Dall; Bull. 37, U. SS.) Nu pate 50, f. 22, *89; Vanatta, Pr. A. N. S. Phila es g oes 1903; Johnson, Occ: Pa“ Bost. Soc. N. H., 7, pimegaors.
Distribution.—Cape Cod, Mass., to the Tortugas. Gulf coast: Crooked Island, Calhoun Co., Fla. Pleistocene, Lake Dorgne, La., borings.
Genus MEIOCERAS Carpenter
deshayesi de Folin, Annales Soc. Linn. Maine et Loire, 11, ’69, Dall (Bull. 27. SN cel. 142. 7 8G,
Distribution.— Tampa, West Fla., to Jamaica, W. I.
os Ss
i324 MoLLUSCA OF THE GULF OF MEXICO 102
undulosum de Folin, Ann. Soc. Linn. Maine et Loire, 11, 1869; Dat eal) 27,0. S. Ni M., p. 142, So.
Distribution.—Charlotte Harbor, West Fla. to Jamaica. Pliocene to Recent.
niiduM Stimpson, Pr. Bost. Soc. N. H., 4. p. 112, 1851, (as Gacaum) Tryon, Man..8;p. 222,. pl. .69; £. 73, °86;. Dall, ee. S., 30-9302) 1892") Vanatta, Pr: A. N.S. Phila., 55) P- 758, 1903. Distribution.—West Florida, at Tampa and Crooked Isl. (Calhoun Co. ), to Jamaica.
Genus BIV@NIA Gray
exerta Dall, Bull. M. C. Z., Harvard Coll., 9, p. 39, 1881; 18, peas p26, 1.-6,’89; Bull. 27,0. S. N..M:, p. 144, pl. 26, PG 730)
Distribution.—Cape Lookout, N. C., to Barbados. Gulf of Mexico, West Florida and Texas regions, deep water. 3I1- 1oo2 fms.
Genus VERMICULARIA Lamarck
Spirata Philippi,Arch. fur Naturg. 2, pt.\1, 224, pl. 7, £. 1, 1836, Racin e7iicHius). tryon, Man, 8, p. 187, pl'55,.f: 99, ‘100, Soma pualli37,. 9 US. NoMa pp. 144,, pl.isa, ta 4,89; Mtr -S.,'3\)p. 204, 792° Sineley, 4th Ann, Rept..G.S. Pow) p)' 340,92, Jounson, Occ, Pa: Bost. Soc..N. H.,.7, p: 123, 1915, . Distribution.—Massachussetts to the West Indies. Gulf coast: West Florida; Matagorda and Corpus Christi Bays, and Miintana, Lex. | Pliocene of Shell Creek, &c, Fla:
Note.—Moerch described as V. melanosclera (Pr. Zool. Soc., p. 174, 1861) a form from Vera Cruz which Tryon regarded as a variety of sfzvaza, and on the same page of the Proceed- ings Moerch described V. quadrangularis from Yucatan. This was also regarded by Tryon as a variety of spirata (See
103 BULLETIN 38 133
Tryon’s Manuel, 8, p. 187, 1886). Shells from the Gulf, Blake station 36, at 24 fms., have been referred by Dall to V. lumbricalis Linn. (See Bull. M. C. Z., 18, p. 261, ’89) but the true Linnzean /uméricalis is Oriental.
nigricans Dall, Pr. U.S. N. M., 6, p. 334, 1883, (as lumbricalzs var.) Bullii37. SiN) cM an see
Distribution.— West Florida and Texas. 2-14 fms.
Genus PETALOCONCHUS Lea
irregularis d’Orbigny, Moll, Cuba, 1, p. 235, pl. 17, f. 16, 18,’42, (as Vermetus); Dall, Bull, 37, U. S) Ny (MG pi eso, Ar. Wi liso 32 pon ae:
Distribution.—Gulf of Mexico between Mississippi delta and Cedar Keys, Fla., at 27 fms. Also Antilles to Guadeloupe. Pleistocene of Fla. Pliocene, Caloosahatchie beds.
varians d’Orbigny, Voy. Ame’r Me’rid., p. 456, pl. 54, f. 7-10, 1843; Tryon,.Man., -8, p. 170, pl. 40, f.\22,!23))"865, ae Crew + TOs io eG. (G2).
Distribution.—Southwest Florida to Rio de Janeiro. Plio- cene, Caloosahatchie beds. Oligocene, Tampa silex beds.
erectus Dall, in Agassiz, Three Cruises of the Blake, 2, p. 71, f. 297, 88; Bull, M.C. Z:, 18, p: 262,’ pl... 28)) ae 37,0 25. NON EA 4p L Bikol AE ee:
Distribution.—Antilles and West Florida. 37-805 fms. Pliocene, Caloosahatchie River, Fla.
Genus SIPHONIUM
nebulosum Dillwyn, 2, p. 1076, No. 19; Tryon, Man., 8. p. 184, pl. 54, f. 87,1886; Dall, Bull. 37, U.S. N. Mi) piagaee:
Distribution.—East and West Florida to Tortola. Genus SILIQUARIA Bruguiere squamata Blainville, Dict. des Sci. Nat., 49, f. 213; Tryon, Man.,
134 MOLLUSCA OF THE GULF OF MEXICO 104
Seno, pl. 58, t..25.'786% Dall, Ball. 37, U.S. Nv M., p. 144, 89. Distribution.—Sarasota, West Florida, to Barbados. 2-163 fms.
modesta Dall, Bull. M. C. Z., Harv. Coll., 9, p. 39, ’81; Agas- siz, Three Cruises of the Blake, 2, p. 71, f. 296, ’88; Dall, Ele Os Z4).18)'p. 260, pl.26,'f., 4,789; Ball. 37; 0. 8. NM p..144, pl. 26; f:'4,\>89. Distribution.—Off Cedar Keys, West Florida, to Curacoa Isl. 94-805 fms.
Genus TURR!{TELLA Lamarck
(Haustator) variegata Linnzeus, Syst. Nat., ed. X, p. 767, 1758,
(as Turbo);Reeve, Conch. Icon., 5, pl. 5, f. 19, ’49; Tryon, ames WV LOSs Dl. Gn, £,.58,°86; Dall, Bulls 37.0. S. N. M3 pp.’ 144, *89. Distribution.—Matagorda and Espiritu Santo Bays, Texas to Cartagena, Colombia, and Antilles. Pleistocene Knapp’s wells, Terrebonne Parish, La., No. I at 1660-1700, 2250- 2450 (?), No. 2 at 1434-1800, No. 3 at 1330-1375 feet.
(Haustator) yucatecana Dall, Bull. M. C. Z., 9, p. 93, 1881; 18, pe2o5, pl 26, £. 3,°°39; Bully 37; Us S: No M., p: 144, pl. 20 0s.).3). 89, Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms.
(Torcula) exoleta Linnzeus, Syst. Nat., ed, X, p. 766, 1758, (as Turbo); Reeve, Conch. Icon., 5, pl. 6, f. 22, 1849; Tryon, Riaileus, Ps 205,. pl.i98, 09; 1886; Dull, Bull..37,,U. .S, N. M., p. 144, ’89; Dall and Simpson, Bull. U.S. Fish Com., i429, \\T QOL,
Distribution.— West Florida, in 50 fms.; Texas coast. An- tilles to Barbados. 45-170 fms.
(Torcula) acropora Dall, Bull. M. C. Z., 18, p. 264, 1889; Bull. a7 U.N. MM), p: 144,789.
105 BULLETIN 38 135
Distribution.—Hatteras to Grenada Isl. Gulf of Mexico, West of Florida, at 14-50 fms., and the coast of Texas, Vera Cruz and Yucatan. 3-413 fms. The Pliocene, closely relat- ed form is 7. subannulata Heilprin of the Caloosahatchie marl.
chipolana Dall, Trans. Wagner Inst. Sci., 3, p. 312, pl. 22, f. 24, 1302:
Distribution.—Lower Miocene of the Chipola beds, West Florida, and of the Bascom No. 2 well, Mobile, Ala., at 1241 feet, Chipola horizon. Aldrich’s collection.
subgrundifera Dall, Trans. Wagner Inst. Sci., 3. p. 313, pl. 22, f. 23) el OOD:
Distribution.—Lower Miocene, Chipola beds, Calhoun Co., West Fla., and of the Bascom No. 2 well, Mobile, Ala., at 1241 feet, Aldrich’s collection. A varietal form was record- ed by Harris from the Upper Miocene, Galveston well, at 2552-2920 feet.
terebriformis Conrad, Cat. Mio. Foss., Pr. A. N. S. Phila., p.
568, 1863, (Name only): Dall, Tr. W) 1. S230 ea. 1892. Distribution.—Lower Miocene, Chipola marl, Fla., and of the Bascon No. 1 well, Mobile, Ala., at 1500-1556 feet. Aldrich’s collection. Also Chesapeake Miocene of Easton, Md. (Conrad’s type locality).
satilla Dall, Proc. U.S. N. M., p. 233, pl. 22,4 eyaraese
Distribution.—Upper Miocene or Pliocene.. Well near Al-. exandria, La., at 49 feet. Brackish water formation.
Genus MATHILDA Semper yucatecana Dall, Bull, M. C. Z., 9,p. 90, 1881, (as Azttzum); 18, p. 266, pl. 20, f. 7, 789; Bull..37, U. S.No aie eres
7) 80: Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms.
136 MoLLUSCA OF THE GULF OF MEXICO 106
Also off Georgia and Fernandina, East Florida. Least dept, 2094 fms,
Genus LITTORINA Ferussac
ziczac Gmelin, Syst. Nat., p. 3587, 1792, (as Zvochus); Reeve, Sonem icon, 10,-/pl. 11,-f./ 57, 1857;' Fryon, Man.,.9; p, Boies 5, 1887") Dall) Bull. 37, U.S. N. M., p. 146; (ise) Wall aud Simpson, Bull. U.S. Fish,Com., 1, p. 429, Igol.
Distribution.—Florida Keys west to Texas and South to Barbados.
augulifera Lamarck, Anim. s. Vert., 7, p. 54, 1882, (as Phasian- gua) Wall, Bull. 327, U.S. N, M., p. 146, ’89; Dall and simpson, Bull..U.S. Fish Com., 1, p. 430, Igor.
Distribution.— West Florida and Texas to the Antilles and Cartagena, Colombia.
mespilum Muhlfeld, Verh, Berl. Ges. Nat., 1, p. 219, pl. 8, f. 8, Moan tas hdc) Lryon, Matn., -.9,p;) 252, pl. 45, f. 16, toe7,) Wall Bull. 27, U..S. N.M., p. 146, 789; Dall and Siapson, Bull) U.S. Fish Com., 1, p. 430, Toor.
Distribution.—Texas to Barbados.
ironaw: Say, Jour..A. N. S. Phila., (1), 2, p. 230, 1822, Tuo- Mevyuand Elolmes,, Pl, Poss. S. Car., ‘p.. 119), pl, 26, £. 5, 1857; Post.-—#Pl. Foss. S. Car., p. 91, pl. 14, f. 5, 1860; Giron, NMan,.O/p. 246, pl, 43,'f: 48,87: Dall, Bull, 37, U. SNe Ep. 146), ply) 69;, 1, 68 89; Tr. Wi 1. Si3)3,p. 320) 492; Singley, 4th Ann. Rept. G..S. Texas, p. 340, 92: Van- appar A ING) Phila.) 55, p. 758, 1903; Johnson, Occ. ae Ost 0c. INGE) 7) pyi21, 1915.
Distribution.— Massachussetts to Jamaica. Culf coast: Cedar Keys, and Calhoun and Washington Cos., West Fla., Chandeleurs, Weeks Island, Point au Fer, La.; Matagorda, Espiritu Santo Bays and Galveston, Tex. Pleistocene, New Orleans pumping station No. 7.
107 BULLETIN 38 137
Fenus LITIOPA Rang
bombix Kiener, Ann. des Sci. Nat.,p. 203, 1829; Ann. des Sci., 30, p.: 222, pl. 17, \B. .£.) 4-3, 18335) Dye ence 9, p. 281, p. 53,.£.:74, 1887; Dall, Bull. 37,0) Sate aae 148, ’89; Singley, 4th’ Ann. Rept: Tex.; p/ 240) aaa: Bull. 112, U. S. N.°M., p. 155, 1021, Johnson) (Oceana: Bost... Soc: IN] ea oad FO. Toms.
melanostoma (Rang) Sumner, Bull. U. S. Bur. Fish., 31, pt.
2p: 720) Lobe Distribution.—New England to Brazil. Gulf coast at Cam- eron, La.; Corpus Christi and Galveston, Tex. Also Pacific, California coast. Pelagic on floating Sargassum.
Genus TECTARIUS Valenciennes
muricatus Linnzeus, Syst. Nat., ed X, p. 1758, (as Zurbo); Reeve, Conch. Icon., 10, pl. 2; £. 11, 1857, (as Litorga) ea, Man., 9, p. 258, pl. 48, f. 68, 1887; Dall, Bull. 37, Ui vou M., p. 146, ’89; Dall and Simpson, Bull. U. S. Fish Com.,
I, Pp. 431, 1OOt. Distribution.—EHast Florida and West Florida and Antilles to Colon.
Genus ECHINELLA Swainson ld
nodulosa Pfeiffer, Tryon, Man., 9, p. 258, pl. 48, f. 75, 1887, (Excl. syn.); Dall) Bull. U. S. N.\M., p.\T46) eq, nat) and Simpson, Bull, U. S. Fish Com., 1, p. 431, 1gor.
Distribution.— North Carolina to Barbados and Texas.
Genus ISAPIS H. & A. Adams
ebsoleta Dall, Proc. U.'S...N. M., 46, p. 233, ol): 22 hie: IQI4. Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet.
738 MoOLLUSCA OF THE GULF OF MEXICO 108
Genus ALABA A. Adams aervaricesa C. B. Adams, Pr. Bost. Soc., N. H., 2, p. 6, 1845, Was veessoa); Moerch, Malak, Blatt., 23, p57, 1876; Dall, Baws S. N. M.,\p. 146, °89; Dall and Simpson, Bull. U: Sliema Com.) 1,)\p)' 431, 1901. Distribution.—Tampa, West Fla., to Haiti. Genus ARCHITECTONICA Bolten granulata Lamarck, Anim. s. Vert., 7, p. 3, 1822; Ency. Me’th., Peawon i. sa-b) 1792, (as Solerzum); Dall, Tr. W.1.S., 3,p. eee eae. Pm US. No M37, p. 232) 1910; Maury,’ Bull. men inal INO) 26, 9. 13't, pl. 23,'f, 3.) 1917: perspectiva Tuomey and Holmes, Pl. Foss. S. Car., p. 120, pl. Ze, o5y7. Not of Lanneeus nor Lamarck.
Distribution.—Hatteras to the Antilles. Gulf coast at Chandeleurs, a., Matagorda and other localities, Tex. Also Pacific Ocean, Lower California to Peru. Pleistocene, New Orleans pumping station No. 7; Upper Miocene, Gal- veston well at 2158-2871 feet. Lower Miocene, Chipola marl, West Fla., and Dominican Republic.
Genus TORINIA Gray
cylindrica Gmelin, Syst. Nat., p. 3572, 1792, (as 7vochus); Tryon, Viemven Dp. ty) ples, £82. 1887; Dall!) Bally 327, UL SiN. me. p. 148, '89; Dall and Sampson, Bull. U. S. Bish Com.,
BA 2, TOOL. Distribution.—Texas to St. Thomas, W. I.
Sanaitera C. B. Adams. Dall, Bull. 37; U. 5. N. M., p. 148, 1889.
Distribution.—Texas to Guadeloupe. Genus RISSOA Fre’minville
iecomtata Wall, Bally MC. Z.,. 18, p.,279, pl. 29) f. 1) 1889: Pee Ue a NM ep. T48)\ pls to, fh 2 80.
109 BULLETIN 38 139
Distribution.—Gun Cay, Hast Fla., 498 fms., Yucatan Strait at 640 fms.
acuticostaia Dall, Bull. M. C. Z., 18, p. 280, pl. 19, f. 10, 1889; Bull. 37, U. S. NoMa p. 150, pling, fi. 16)
Distribution.—VYucatan Strait, 640 fms., and between Mis- sissippi delta and Cedar Keys, 32 fms. Also off Hatteras and Barbados. Probably a variety of R. xanthtas Watson (Chall ‘Gastr:; p. (588, pliaa: £., 5) 1885).
Genus RISSGINA d’Orbigny
decussata Montagu, Test. Brit., p. 399, 1803, (as Turbo); Schwartz von Mohrenstern, Denkschr. d. Kais. Akad. d. Wissensch. Wien. 19, p. 80, f. 12, 1860; Tryon, Mam iayp. 385, pl. 58, f. 15, 1887; Dall, Bull. Ml Co Zee W303 ie Wi IS) Bam A aaa eae
Distribution.—Cape Fear, N. C., to Haiti and to West Fla. 2-17 fms. Also European. Variety planata Dall, Caloosa- hatchie Pliocene.
laevigata C. B. Adams, Contr. Conch., p. 114, 1850; Schwartz von Mohrenstern, Denk. Kais. Ak. Wien., 19, p/ 11", 4-776: 1860; Tryon, \Man., 9, p.. 390, pl. 59, f. 48, 87a AV ol Sl) BMD AD ie 2, Distribution.—Cape Lookout, N. C., to St. Thomas, W. I. Also West Florida. o-22 fms. Pliocene, Caloosahatchie beds. Lower Miocene, Chipola beds, West Fla.
chesneli Michaud, Descr. des Cog. Nouv., p. 17, 1832, (as Rés- soa)» Tryon, Man., 9, ‘p. 380, pl. 56, f:.73,) 87, ane W.1.S., 3, p. 343,92: Singley, ath: Ann. Rept) emi BAT Oe! Distribution.—Hatteras to Guadeloupe, Gulf coast: Cedar Keys, Fla.; Corpus Christi, Tex. Also Mediterranean and Mauritius. Pliocene, Caloosahatchie beds, West Fla.
browniana d’Orbigny, Moll, Cuba, 2, p. 28, pl. 12, f. 33, 35, ’42;
LAO MOLLUSCA OF THE GULF OF MEXICO iIO
Mine, Man. o, p; 390, pli’ 59,1.) 45, 87; Vanatta, Pr. A. Neo ena 55; Dp. 7585; bGO3.
Distribution. — West Indies and Gulf coast, Crooked Island, Calhoun Co., West Fla. (Vanatta).
elegantissima d’Orbigny, Moll. Cuba, 2, p. 26, pl. 12, f. 27-29, eae, ryen, Man.) 9, p. 374, pl, 56, f. 51. 1887. Distribution-—West Indies and Gulf coast, Chandeleurs, Was eDaily.
Sagraiana d’Orbigny, Moll. Cuba, 2, p. 25, pl. 12, f. 4, 5, 1852; (iyen. Mam, 9, pi 384, pl. 57, £. 86, 1887; Dall, Bull. 37, UL SVING NL. |p. 150,780.
Distribution.—Florida Strait, to Martinique. Also Gulf coast at Vera Cruz, Mex.
Genus BENTHONELLA Dall
nisoms Dall Bull, M.C. Z., 18, p. 283, 1889; Bull. 37, U.S.
NGgMEs p.:1 50,’ 80.- Distribution.—Gulf of Mexico between Mississippi delta and
Cedar Keys at 940 fms.
Caza all Ball M:.C. Z., 18, p. 282, 789; Bull. 37,U. S. N. M.,: eresO,) pl 42, 1.5, /8o, Distribution.—Georgia to Cuba, West Florida and Texas regions of the Gulf. 6-463 fms.
Heenan Dall, Bull. M. ©. Z.; 18, p. 282, 89; Bull. 37, U. S2 N. MEP, 252, 30, |
Distribution.—Gulf of Mexico between Mississippi delta and Cedar Keys at 940 fms., bottom temperature 39° F. Also off Cuba.
Genus SKENEA Fleming
planorbis Fabricius, Fauna Groenlandica, p. 394, 1780; Gould, Peavemntass)) ps) 200) 1.563, 1870; Ball, Bulll37...Us SiN: Mpa LO, ply 52) 0,18, 1889; Bush, ‘Tr. Conn. Acad! 'Sct,,
LE BULLETIN 38 I4I
10, p. 100, pl. 23, f. 5,8, ’99-1900; Johnson, Occ. Pal Bost: DOG. UNGAR Ca ao Ten ue ris! flelix depressus Montagu. Distribution.—Greenland (type locality) to Charlotte Har- bor, West Florida. ‘This species is the genotype. Genus AMPULLARIA Lamarck depressa Say, Dall, Bull. 37, U.S. N. M., p. 150, ’89. Distribution.—Georgia to West Florida, Texas and Mexico. caliginosa Reeve, Conch. Icon., Mon. Ampullaria, {. 118, pl. 25,
1856; Dall, Proc. U.S. N. M., 8, pi 255, 1885s ee UU. S:2NUM:S perso Taso.
Distribution.—Cedar Keys, West Florida to Texas, Mexico and Nicaragua. In freshwater swamps. pinel Dall, The Nautilus, p. 75, Nov. 1898.
Distribution.— Homosassa River, Fla.
Genus PALUDESTRINA d’Orbigny
aldrichi Dall, Proc: U.S. N. M:,'46, p. 234, pl. 22nt 7 owes Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet. Brackish water formation.
plana Aldrich, Nantilus, 24, pt. 12,' p. 139, pl S)0t "ee noun Dall; Pr. USS: ANMME 46) pp. 224) nora, Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at.49 feet.. Also Satilla River, Gal) (iype locality).
curva Dall, Proc, U.)'S))N.)M., 46, p. 235, ‘pli 22) bea me Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, Ga., at 49 feet.
cingulata Dall, Proc. U. S. N.'M., 46, p. 235, pl.\22) tf. a) 0am
Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet.
age AS Nee SS SS
r4zZ MOLLUSCA OF THE GULF OF MEXICO 112
merricusa: all, Proc, U.S: N..M., 46, p. 235, pl. 22. f..9, 1914.
Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet. Brackish water formation.
anton ail, Proc. U.S. N. M., 46, p. 235, pl. 22, f. 2, 1924.
Distribution.—Upper Miocene or Pliocene.. Well near Al- exandria, La., at 49 feet. |
Genus ASSIMINEA Leach
auberiana d’Orbigny, Moll. Cuba, 2, p. 8, pl. 10, f. 6, 7, 1842, (as Faludestrvina); Dall; Pr. U. S. N. M., 6, p. 335, 1883; Pani. ou Ni. Mi piso, 893. Pr. W. 1. S$.) 3) pr 247, 1892.
Distribution.—Cedar Keys, West Florida, and the Antilles. Pliocene, Caloosahatchie and Shell Creek, West Fla.
Genus TRUNCATELLA Risso
caribaeensis Sowerby (MSS.); Reeve, Conch. Syst. 2, pl. 182, f. 7, 1842; Binney, Land & Fr-Water Shells, N. A. Smith. Mise. Coll), No, 144) pt: 3, p. 98, pl. 198, 1865;: Dall, Bull. SW. IN Mi po 152) 89: Dall & Simpson, Bull, U.S, Pela Com 1, pi 426, pl) 153) 1:7 190T.
Distribution.—West Florida, Alabama, Texas (?) and Mex- ico. Also Greater Antilles.
bilabiata Pfeiffer, Wiegm. Arch., 1, p. 253, 1840; Binny, L. and BN oy ON AL pt. 3p. 99, £. 199) 1865; Dall Bull. .37, So ON. ME.) pe 152, 780. Distribution.—Sarasota, West Florida, to Honduras and Cuba.
pulchella Pfeiffer, Wiegm. Arch., 1, p. 356, 1839; Binney, L. and
Remeron. INO UA Ot! 3p. 99, f.'200,( 18657 Dall) Bull.)27, Wa ScONY NE. p.ms2)°8o; Dall & Simpson, Bull.) U..S.' Fish
Com, 15) py 426, 1901. Distribution.—Florida to St. Thomas, W. I. Gulf coast:
BULLETIN: 38 | I43
Tampa, West Florida; Galveston, Texas. (Dr. Gurley).
subcylindrica Linnzeus, Syst. Nat., ed. XII, p. 1248, 4767, (as
frelix); Gray; in Turton, Man., p. 295,t. 72, Beag ese y
Land and Fr. W. Sh>N. A., pt.93, pt) 100) a. y2eetmaeasen
Dail and Simpson, Bull. U.S. Fish Com., p. 436, 1901. Distribution.—Tampa, West Fla., to St. Thomas, W. I. Genus CHEILEA Modeer
equestris Linnzeus, Syst. Nat., ed. X, p. 780, 1758, (as Patella);
Tryon, Man., 8, p. 137, pli 41, f. 25, 26, 1886; Dale I. S., 3, p- 348, ’92, (as Afztrularza); Pro. SN eee 1D) 232) UnGrO:
Distribution.—Hatteras to Barbados, Gulf of Mexico, Blake
station 2. Also Pacific Ocean, Mexico to Chile. Pliocene, Caloosahatchie beds, Fla.
Genus CAPULUS Montfort
intortus Lamarck, Anim. s. Vert., 6, pt. 2, p. 18, 1822: d’Orbig-
ny, Moll. Cuba, 1, p. 186, pl. 24, s. 22; 23;)m@4e. tig Man., 8, p..13!, pl. 30, f. 75, 86; Dall, Bulk 270) ae Miu p54) eo: : Distribution.—Gulf of Mexico, Blake station 5, at 229 fms. Also off Florida reefs, Hatteras and Bahamas.
Genus CREPIDULA Lamarck
fornicata Linnzeus, Syst. Nat., ed. XII, 1, p. 1257, 1767, (as
Patella); Hilgard, House of Rep. Ex. Doc., 1, pt2, 9. 637, 1879; Dall, Bull. 37,,U: S..N. M., p.' 152) pli 42a 50, £.. 23, 24, 1880: ‘Ir. W..1.S., 3, p. 256) 702, eee 4th Ann. Rept. Tex., p. 341,:’92; Vanatta) Praia Phila., 55, p. 758, 1903; Johnson, Occ. Pa: Bost: Sec iy Ey 7, LOO), | TOTS.
Distribution.—Granada to Cartagena, Colombia. Gulf coast: Cedar Keys, St. Mark’s, Ft. Barranca, &c., West Fla., La., Galveston and Corpus Christi, Tex. Pleistocene, Lake
RS ee ee gee ee SS a eee ee ek : =
SS
144 MoLLUSCA OF THE GULF OF MEXICO II4
Borgne borings, New Orleans well of 1856, and New Orleans pumping station No. 7. Pliocene, Caloosahatchie beds. Lower Miocene Chipola beds, West Fla.
Hand Say jour. A. N.S: Phila., (1), 2; p. 226, 1822; ‘Tuomey enaubbvolimes, Pl. Hoss. S.\'Car., p. iri, pli25, f. 12, 1857; Paine 37, UO. Ne Miip) 152, pl. 48) £12, pl. 50, £. mom bn WTS. 30) ps 358,92: Vanatta Pr, A. N.S. Phila 55, p. 758, 1903; Jonnson,-Occ. Pa. Bost. Soc. N. H.,
G5 APs LO, HQT 5.
Distribution.—Canada to Trinidad, W. I. Gulf coast: Cedar ‘Keys, St. Mark's, Fla.; Chandeleurs, La.; Galveston, Corpus Christi and Matagorda Bays, Texas. Pleistocene, New Or- feaus | pumping ‘station No.7 and North Creek, Fla. Pliocene, New Orleans Gymnasium Club well at 1200 feet, and Caloosahatchie beds, Fla. Lower Miocene Chipola beds, West Fla. and of the Bascom No. 1 well, Mobile, Ala., at 1500-1556 feet, Chipola horizon.
aculeata Gmelin, Syst. Nat., 7, p. 3693, 1792, (Patella); Tryon, Manes) P.120, pl. 30, f. 61-62, 1886; Dall, Bull. M. C. Z., Heawpw2o0. 1589; Vanatta,.. Pr. Al) N.S. Phila., 55, p.758, naega Wall Bull i112, U. S. N. Mi, ps 162) 1921.
Distribution.—Cape Lookout, N. C., to Barbados. Gulf coast: Indian Pass, West Fla., and Texas. Usually 0-25 fms., but dredged in Gulf of Mexico, near Tortugas at 539 fms. Dwarfed and white at that depth. Also Pacific Ocean, California to Chile. Pliocene, Caloosahatchie beds, West Florida.
Penvexa say, Jour. AY Nz S. Phila., (1), 2, p. 227, 1822; Dall, Pees cose Mo pelrs2, pl. 50; .f..25, 1889; Tr), W. Seg DL 357" \O2!
Distribution.— Nova Scotia to Florida. Gulf coast: West Fla.; Chandeleurs, La.; Galveston, Tex. Pleistocene, North Creek, West Fla.
Its BULLETIN 38 X45
Genus CRUCIBULUM Schumacher
auricula Gmelin, Syst. Nat., p. 3694, 1780, (as Patella); Dil- lwyn’s Cat., 2, p.1017, 1617; Dall,: Bull) soo Gee 284, 7°89; Bull. 37, ers. IN. Me i. ges ee
planatum Schumacher, Essai, p. 182, 1817.
scutellatum var. auriculatum (Chemn.) Tryon, Man., 8, p. 118,
ply32)'f, 24,35. ese: Distribution. Cedar Keys, West Fla., and Texas, south to Barbados and Northern Brazil, 25-111 fms. Pliocene, Caloosa- hatchie beds, West Fla. The analogous Pacific species is C. spinosum.
striatum Say, Jour. A. N. S,. Phila., 5) p. 276) e826(ae Calypiraca); Tryon, Man., 8, p. 118,-pl. 23, fay ee Dall, Boll. M. C. Z., 18, p. 284, °80; Johnson) cea: Bost. Soc., Nj H.7, p, 109, 1055.
Distribution.—Nova Scotia to Florida. Cited by Dr. Dall from the Gulf of Mexico at Vera Cruz.
Genus CALYPTRAEA Lamarck
candeana d’Orbigny, Moll. Cuba, 2, p. 190, pl. 24, f. 28, 29,
1842, (as Lnfundibulum); Dall, Pr. U.S. N. M., 6, p. 335,
"83, (as Galerus); ‘Tryon, Man., 8, p. 121, pliysaige:
77, 1886; Dall, Bull, 37, U.. 5.) Noo Mi oe ee parvulus Dunker,1875.
Distribution.—Hatteras to Brazil. 6-52 fms. Gulf of Mex-
ico, Cedar Keys, West Florida, and Yucatan Strait, 640 fms., dead shell.
Genus XENOPHORA Fischer de Waldheim
conchyliophora Born, Index Mus. Cees., p. 333, 1778, (as Trochus) ; Tryon, Man., 8, p. 161, pl. 46, f. 80, °86;) Dall) daigaan U.S: N. M., p. 154, °80; Tr. W.1. S., 3; pp) gee geaniee 4,£.'Io, 10a,:’92; Bulli oo, U:'SuN: M., p. nos, eee
146 MOLLUSCA OF THE GULF OF MEXICO 116
Beto Maun. Bull, Aimer) Pal., INov'20,) p.\133, pl. 23, f. 7, 1917. agglutinans Lamarck.
Distribution.—Hatteras to the Antilles. Gulf of Mexico, West Florida and dredged at various stations, 14-229 fms. Usually not so deep as X. carvibaca and more heavily loaded. Pliocene, Caloosahatchie beds, West Fla. Lower Miocene, Chipola marl, West Fla. Oligocene, Tampa silex beds, Fla. Upper Eocene, Jackson, Miss. Eocene, Wood’s Bluff, Ala. Wpper Cretaceous, (Ripley) of Prairie Bluff, Ala. . The most ancient species of the Gulf Mollusca.
caribaea Petit, Jour. de Conch., 5, p. 248, pl. ro, f. 1, 2, 1856; Miyounian., 8, p. 162, pl. 47, 1-96, ’86; Dall} Bull. M. C. wetowon 201, 89; Dall) and Simpson, Bull. U.S. Fish Com ap) 438, LoOr.
Distribution.—Hatteras to Barbados and Gulf of Mexico, various stations, 14-274 fms.
Genus NATICA Scopoli livida Pfeiffer, Wiegm. Arch., 6, p. 254, 1840; Moerch, Malak. Bint 4) Dp, 6400773) Dall Bull) MoiCiZ.,°18.0p. 292, 80° end 2 Ue. IN. ML p. 154), 80: Youaicusts ©. B. Adams, Contr. Conch., p, 111, 1850.
proxtma Sowerby, Thesaurus, Vadica, pl. 8, f. 111. Not of C. B. Adams.
Distribution.—Hatteras to Barbados. Gulf of Mexico, be- tween the Mississippi delta and Cedar Keys, 26 fms.
maroccana Chemnitz, Conch. Cab., 5, p. 270, pl. 188, f. 1905-10; Diwver Cat. Rec oh.,2)p) 983, No. 13,1817; Dail,) Bull. PO 7. To pw202.' 1889; Bull) 37, Us 5. IN. Ma pirsa, 188oQ.
Distribution.—Hatteras to Barbados and Gulf of Mexico, west of Florida. Also European and West African.
rE BULLETIN 38 IA7
canrena Linnzeus (in part) Syst. Nat., ed. X, p. 776, 1758, {a> Nerita); Reeve, Conch. Icon., 9, pl. 4, f. 14) 18553) Paouiey and Holmes, Pl. Foss. S. Car., p. 115, ple) 25. tegege ea: Dall, Bull. 37, U.S).N. M., p. 154, 780; ie Nees p. 364,92; Maury, Ball.-Amer. Pal., Ne. Zone 22 TAO, TO D7,
plecatella Conrad, Pr. Ay N.S, Phila), p. 564) 1868. |
Distribution.—Hatteras to the Antilles, Pernambuco and Cartagena. Gulf coast, West Florida and Galveston. Plio- cene, Caloosahatchie beds, Upper Miocene, Galveston well at 2158-2920 feet. Lower Miocene, Bascom No. 2 well, Mo- bile, Ala., at 1241 feet, Chipola horizon.
alticallosa Dall, Trans. Wagner Inst. Sci.,'3, p. 365) pligau 23),) LOO? Distribution.—Lower Miocene, Chipola marl, Northwest Florida, and of the Bascom No. 2 well, Mobile, Ala. at 1241 feet, Chipola horizon.
Subgenus Cryptonatica Dall
pusilla Say, Jour. A. N. S. Phila., 2, p.257,; 1822 ;.iioane House of Rep. Ex. Doc.,1, pt. 2, p. 886, 1878;) Dalliieaim 37, U, S&S. N. M.j p. 154, pl. 50, £.:21, 7805) Simgter ae Ann, Rept. Tex., p. 341, ’92; Vanatta, Pr, Al Nis) aia 55, Dp. 758, 1903; Johnson, Occ. Pa. Bost: Soc, Nua aa 105, IQI5. Distribution.—Maine to Florida Keys. Gulf coast: St. Joseph’s Bay and Crooked Island, West Fla., Galveston, Tex. Pleistocene, New Orleans artesian well of 1866, Lake Borgne borings, Knapp’s wells, Terrebonne Parish, La., No. 3, at 1150-1200, 1330-1470, 1500-1525 feet.) Pitocene Caloosahatchie beds, and of the New Orleans Gymnasium 7 Club well at 1200 feet.
Genus POLINICES Montfort
Subgenus Euspira Agassiz
tenuis Recluz, Jour. de Conch., 1, p. 388, pl. 12, 27 eege:
L438 MOLLUSCA OF THE GULF OF MEXICO 118
PMiaroM Naan." 8). 38, ple TA) 4.25) 788: Dall, Bull. M,C. eon 265) Sow omlNa7) WU) 3.) NMG po. say T8890} CAI as Lunatia).
Distribution.—Gulf of Mexico, Yucatan Strait, 640 fms. Also of Cape Florida, Cape Fear and Cuba, 84-640 fms. Pulso(Pacitic Ocean,’ Valparaiso, Chile, Recluz’s type lo- eality:
lepialea Watson, Jour. Linn. Soc., 15, p. 261, 1880; Dall, Bull. MOOG.) 63), 81; Watson, Chaily Rept) Gastr., p. Aion ta 85. Wall Bulls) MC. Zu) \18, p. 205,’ 89. (All as Lunatia).
Distribution.— Yucatan Strait, 640 fms. Also off Fernandi- na, Fla., and Sombrero Isl,, 450 fms,
lningita Dall Bull. M.C..Z., 9; p. 93, 1881;, 18, p. 295, pl. 21, eee sereimil 27. WU. Sa ING Mo} ip. 154, pl. 2n, f.. 12,80. (All as Lunatia).
Distribution-—Yucatan Strait, 640 fms. Also near Old Providence, 382 fms. L.vadzata Watson, may be identical with this species.
Hennecrypia-Gabb, Jour. A. N.S. Phila.. (2); 4, p. 375, pl. Opts. too: (as /Vaizca): Wall. Bull./oo, U.S) Na Mp, HOO pl. .O, fF: QO, 1915.
ema ores Gabby jour, A. N.S. Phila., \(2)) "8, p:.339) pl: Aw fe 4, 1875. Distribution.—Upper Miocene, Galveston well at 2465-2733 feet, and of the Jennings-Heywood No. 29 well, Jennings, Ija., at 1960-1980 feet. Lower Miocene, Chipola marl, West Fla. Oligocene, Tampa silex beds.
Subgenus NEVERITA Risso
duplicata Say, Jour.,A. N.S. Phila., 2, p. 247, 1822, (as Watica); Sammi a7. Wo.) Ne. p. 154 ploisay fs 12). 80. an!
[eee
119 BULLETIN 38 I49
W..1S., 3, p.. 368,92: Vanatta, (Pr. Aes Phila., 55. Pp. 758, 1903; Johnson, Occ. Pa, Bost. Soc. NO 7) apiaes, 1QI5.
campechiensis Recluz, fossata Gould, texastana Roemer.
Distribution.—Massachussetts to Mexico. Gulf coast: Cedar Keys, and many other localities, West Fla., Horn Island, Miss.; Point au Fer, Cameron, Chandeleurs, La.; Matagorda Bay and Galveston, Tex. Pleistocene, Grand Chenier, New Orleans well of 1856, New Orleans pumping station No. 7, Zigler, No. 15, Jennings; La., at 1350-1481 feet, Jennings- Heywood No. 30 well at 1127-1169, 1169-1277 feet, Knapp’s wells, Terrebonne Parish, No. 2, at 1150-1200, 1330-1375, 1443-1618, 1700-1839 feet. Pliocene Gymnasium Club well at 1200 feet. Chesapeake Miocene, Maryland to Florida. Upper Miocene, Galveston well at 2158-2920 feet. Miocene, Crowley No. 25 well, Jennings, La., at 2468-2500 feet.
Subgenus Payraudeautia Bucq. Dautz. & Dollf..
nubila Dall, Bull. M. C. Z., 18, p. 294, 1889, (as WVeverita); Dall and Simpson, Bull. U.S. Fish Com., 1. p. 439, pl. 58, te 6, TOOT.
Distribution.—Gulf of Mexico, Blake station 23, at 190 fms. Also off Barbados, 140 fms, and Porto Rico.
Subgenus Mammilla Schumacher
uberina d’Orbigny, Moll. Cubana, 2, p. 31, pl. 17, f. I9, 1842, (as Natica); Moerch, Malak. Blatt., 24, p. 60, 1877; Tryon, Man., 8, pl. 16, f. 57, ’86; Dall aud Simpson, Ball, Uaiss, Fish’ Com,,. 1,9. 436. voor.
Distribution.—Gulf of Mexico, west coast of Florida, 14-40 fms., living. Also off Sombrero 54-70 fms. and Porto Rico.
lactea Guilding, Tr. Linn. Soc., 17, p. 29, ’33; (as Waticina); Tryon, Man, 8, p. 49, pl. 16; 1.57, 58, 86; Dall, Baltes,
Betas San aS
X50 MOLLUSCA OF THE GULF OF MEXICO 120
U.S.N. M,, p. 156, 89; Dall and Simpson, Bull. U. S. Fish Come 1, p. 439, Igol,
Distribution.—Florida Keys to Porto Rico and Brazil. Also Texas region of the Gulf of Mexico. West Fla. (?).
brunnea Linck, Beschr. der Rostock Sammlung,2, p. 140, 1807; Pa iwi) 27,.U.5. N..M., p..156,'’ 89.
mamitlaris Tamarck, Anim. s. Vert., 6, pt. 2, p. 197, ’22, (as Nawcay wiyon, Man, , 8, p; 43,-pl. 18, f.74, 786; Dall and simpson, Bull: U.S. Fish Com., 1, p. 439, Igor.
Distribution.—Florida Keys to Trinidad Island and Texas region of the Gulf of Mexico.
Genus SINUM Bolten
prespectivum Say, Amer. Conch., p. 175, pl. 25, (as Sigaretus); Holmes Post-Pl.) Foss. S. Car., p.’81, pl. 12, f. 16, >60;Tryon, MAae Spa157, pl. 24. f. 61,64, 7°86; Dall, Bull. 27; U.S. Neve on5O, So; Vanatta, Pro A, N.S. Phila.) 55, p. 758, 1903. Distribution.—New York to Martinique. Gulf coast: Cal- houn Co., Fla.; Chandeleurs, La.; Galveston, Mustang Isl. and Pass Cabello, Tex. Pleistocene, New Orleans pumping station No. 7, cited by Harris from Upper Miocene, Galves- ton well at 2252-2600 feet.
maculatum Say, Amer. Conch., p. 176, pl. 25, (as Szgarefus); Mimvou Mane (8, p./ 56) pl. 24, 1. 57, 58,7 86; Dall, Bull..37, eae IN Me, p4.156;)' 89. Distribution.—Hatteras to Guadeloupe and West Florida, shallow water. Closely related to martinzanum Phillip, Gub. uh Beschr 1, p.it44, pl. 1, f. 5, 1844).
mimon Dall, Bull. M,C. Z., 18, p. 297, °89, (as Szgaretus); Bull. Ba ato IN OMY Dp. 156,789.
Distribution.—West Florida, Florida Keys and Sombrero Isl., W. I. 54-84 fms.
121 BULLETIN 38 Ist
Genus EUNATICINA Fischer
semisulcata Gray, Zool. Beechey’s Voyage, p. 137, (as atica); Tryon, Man., 8, p. 32, pl. 9, f. 74, ?86;Dall, Bull ine a 13,)(p:296,) 89) (Drv Wet S23) p ia seuiaes
jfordiana and fordiz Simpson.
Distribution.—East Florida to Porto Rico. Gulf coast, Sarasota Bay, West Fla., between tides.
Genus LAMELLARIA Montagu
rangi Bergh, Mgr., p. 94, No. 8, ’53; Dall, Bull. 37,U. S. N. M., Pl 50; WeOe Distribution.—Texas region of the Gulf of Mexico. Ap- parently never figured.
Genus MARSENINA Gray
ampla Verrill, Pr. U. S. N. M., 3, p. 374, "805%\ ii eee Acad. Arts and Sci., 5, p. 518, pl. 42, 2. 3)\ga, (82) @20n Bull. 37, U. S.\N. M., p. 156, 789; Johnson, Occ, Payipece Soc. No Ei 7: poo, OLS!
Distribution.—Maine to Sarasota Bay, West Fla.
Genus ACMAEA Eschscholtz
candeana d’Orbigny, In de la Sagra’s, Hist. Pol. y Nat. Isla de Cuba, 2, p. 199, pl. 25, f. 1-3, 1845, (as Patella); Dall, Bull. 37,,U. 8S... Nv 'M., pp. 156, 89; Tryon,) Man! (meio: ’o1; Dail and Simpson, Bull. U..S. Fish Com 5 Veh ano, IQOI.
Distribution. —Florida Strait to Tobago, W. I. Also West Florida and Texas.
punctulata Gmelin, Syst. Nat., 13, pp. 3705, 3717, 1792, (as Patella); Pilsbry, Tryon,’. Man.,13, p. 37, ples) memes 1891;..Dall and Simpson, Bull. U.S. Fish Com) ja) pie IgOl.
O59 = Sa ee £-
MOLLUSCA OF THE GULF OF MEXICO 122
ie Ct is
Distribution.—Key West, Florida to Porto Rico, Guade- loupe, and the Gulf of Mexico at Vera Cruz, Mex.
leucopleura Gmelin, Syst. Nat., 13, p. 3699, 1792, (as Patella); Eucpny; iryon,,Man.; 13, 7; 40, pl. 5, f.) 16-21, *91; Dall .& Simpson) Bull. U.S. Fish 'Com., 1, p. 440, 1901.
geelanoicuca Wali, Bull. 37, U. S. N. M., p. 156, 1889. Not of Gmelin.
Genus LEPETELLA Verrill
iubicola Verrill and Smith, Amer. Jour. Sci., (3), 20, p. 396, sor Vemuill “Yr. Conn. Acad. Arts and)Sci., 5, p.534, pl. Bowme2ey 20a, 32;) Dall) Bull, M. CC. Z., 18)p.. 413, -pl. 25, Pe oecse mem 27 00S) IN NE. pt) r58)) pl) 25; £:6,. 80; Volusom @ec:\Pa, Bost..Soc.,N. H., 7, p. 86; 1915. Distribution.— Martha’s Vineyard, Mass., at 130-388 fms., to Gulf of Mexico between Mississippi delta and Cedar Keys, at 324 fms.
Genus PHASIANELLA Lamarck
pulchelia C. B. Adams, Proc. Bost. Soc. Nat. Hist., 2, p. 7, 45, tae e7bo): Vryon, Man.’ 10, pp) i264) )'’88; Dall, Bull. M, OZ ron 2351) Soo Bull 27) Wa S.No Mi) po 158), 1889.
brevis C. B. Adams. Not of d’Orbigny, 1842.
Distribution.—Cedar Keys, Fla., to St. Thomas, W. I. Plio- cene of South Carolina.
Genus TURBO Linnzeus
castaneus Gimelin, Syst. Nat., p. 3595, 1792; Pilsbry, Man., ro, mazes) 45) 0) 88-80,)'88; Dall, Bull. 37,0. SiN.) M., p: nee oo; Wallhiand Simpson, Bull. U.S: Kish Com., 1, p. A LOOT, Distribution.—Cape Hatteras to Trinidad, W. I. Also West Florida in shallow water. 25-295 fms, Pliocene, Caloosa- hatchie and Shell Creek, Fla.
123 BULLETIN 38 153
crenulatus Gmelin, Syst. Nat., p. 3575, 1788; Pilsbry, Man., Io, p. 203, pl. 45, f. 90, 88: Dall, Bull. 37, U. Sees) (89) Tr. Wis ain ase) “oe, Distribution.—Hatteras to Barbados. 2-30 fms. Also West
Florida at Crooked Island and St. Andrew’s Bay. Pliocene, of the Caloosahatchie, probably a variety cf crenulatus.
Genus LEPTOTHYRA (Carpenter) Pease
induta Watson, Jour. Linn. Soc., 14, p. 715, 1879 (as Zurbo); Chall. Gastr. pies 1G, fi, aee5.
albida Dall, Bull. M. C. Z., 18; p. 352, pl. say ieee
Distribution.—Hatteras to Martinique, 15-2805 fms. Yuca- tan Strait, 640 fms.
Genus LIVONA Gray pica Linnzus, Syst. Nat., 10, p. 763, 1758, (as Zurbo); Pilsbry, Man., 11, p. 277, pl. 41, f..24, °89; Dall, Bulli 37. Sa M., p. 160, °89; Dall and Simpson, Bull. U.S) Fish @om.: I, DP. 443,, £901. Distribution.—Charlotte Harbor, West Florida, to Colon. 37-160 fms. | Genus GAZA Watson superba Dall, Bull. M. C. Z., 9, p. 49, '81;. 18, Pp. 254.) Dr aan. 4, 4a,” 89; Bull, 37, U.S. N.°M., p. 160; plea 1889. Distribution.—Gulf of Mexico between Mississippi delta and Cedar Keys, 324 fms. Also Barbados.
fischeri Dall, Bull, Mus. Comp. Zool., Harv. Coll., 18, p. 355, pl. 37, f. 6, °89; Bull. 37,.8:S.N. M., p. m6@; 7s eg ete 18809.
Distribution.—Gulf of Mexico, west of Florida and St. Lucia, W. I., 423-426 fms.
ae flea
= Bg —s
154 MOLLUSCA OF THE GULF OF MEXICO 124
(Callogaza) watsoni Dall, Bull. Mus. Comp. Zool., 9, p. 50, ’81; Pee eS Oil. 22\t 70 7a!) pl.23) f. 0, Tay pl. 24, f), 2, 2a. 1889.
wrongarie jiosa Dall, Bull, M. CC. Z., 9, p. 42, 81. Distribution.—Yucatan Strait, 640 fms. Also off Cuba and Barbados.
Genus LIOTIA Gray
gemmia Tuomey and Holmes, Pleioc. Fos. S. Carolina, p. 118, pl. 26,, f. 4, °56, (as Zrochus);Dall and Simpson, Bull. U. S. Bish Com. ,1, p. 445, 1901.
tricarinata Stearns, Proc. Bost. Soc. Nat. Hist., p. 23, for ’72;
Was Aryeniectonzca):; Dall, /Bull..M.C. Z:, .18, p. 389, 89; Pela Ws 9. IN. M., |p.) 166, °89.
Distribution.—Hatteras to Porto Rico. Also off Tampa, West Fla., 15 fms. Pliocene, Caloosahatchie beds. Para Wa Bull M. C.Z., 18, p..389, pl. 33, £8, ’89-
Distribution-—Gulf of Mexico, Blake Station 2, at 805 fms. Also off Carolina, Key West and Havana.
Genus CALLIOSTOMA Swainson euglyptum A. Adams, Proc. Zool. Soc., p. 38, 54, (as Zizyphinus) ; Reeve. Mon. Z22yp., pl. 3, f..17, °63; Dall, Bull. M,C. Z., HremeZoau OG: Pilbry, Maniyi1t, p.374,- pl. 15,'f.137;, pl 57, 1. 9, 89. Distribution.—North Carolina to Florida, Texas and Vera
Cruz, Mexico. Erroneously referred by Reeve to Tasmania. 3-32 fms. Pliocene ancestor is C. philanthropus Conrad.
circumcinctum Dall, Bull. M. C. Z., 9, p. 44, ’81; 18, p. 364, pl. Pesan sa, Son Biull. 37, U.S. NM) py 162) pl, 22, £1 3;
3a, ’89. Distribution.—Florida Keys, Antilles and Yucatan Strait. 640-805 fms.
Sei
125 BULLETIN 38 I55
corbis Dall, Bull. Mus. Comp. Zool Harv. Coll., 18; p. 365, pl. 33, f: 1, °89; Bull. 37, .U.-S. N.M... py Woe Rie eee 1889.
Distribution.—Gulf of Mexico, west of Florida, to Jamaica. 220-450 fms.
tiara Watson, Jour. Linn. Soc. 14, p. 696, °79; Rept. Chall. Gastr., p. 60, pl. 6) f. 4, ’85, (as 7vochus); Dall - Baie, Deo IN ML paGongeG.
Distribution.—Gulf of Mexico, Texas region, to Dominica, W .. 1... 220-780 ims.
roseclum Dall, Bull. Mus. Comp. Zool, Harv. Coll., 9, p. 45, ’81; 18, p. 366, pl. 24, f. 6, 6a, ’89; Bulle 37, 0G Su ee: 162, pl. 24 fon Oa), 80,
Distribution.—North Carolina to Yucatan. Also Florida Straits and Havana. 21-200 fms.
pulcherum C. B. Adams, Contributions to Conch., No. 5, p. 60, "50; (as Zrochus); Pilsbry, Man., 11, p. 375, *69)) Daley 27, WU. SINT MY hs PO22\ 80).
Distribution.—Hatteras to St. Thomas, W. I. Texas region Gulf of Mexico. 15-63 fms. Type locality, Jamaica.
veliei Pilsbry, The Nautilus, p. 128, 1900. Distribution.—Caxambas Pass, Southwest Florida.
(Eutrochus) jujubinum Gmelin, Syst. Nat., p. 3570, 1792, (as t Trochus): Pilsbry, Man., 11, p. 404, pl. 40, #16) aan ane ‘ Bull. 37, U. S. N. M., p. 162, ’89; Dall and Simpson, Bull. ; U.S. Bish Com.) \ige.444) | r90%:
Distribution.—Hatteras to Cartagena, Colombia. Gulf coast, West Florida, Texas and Yucatan. Pliocene, Caloosahatchie $ beds, Fla. Ke (Eutrochus) jujubinum var. perspectivum Koch, Philippi, Abbild. u. ~ ;
Sie ea Se
x56 MOLLUSCA OF THE GULF OF MEXICO 126
Bese 1, pt. 2,
LTOMUS ples 1, L054) POA Se eispty, (Many, wee 405, pl..66, 1.35, 36, 89.
papacnss Courad, Pr. A. N.S. Phila., 3, p..26, pl. 2, f. 35, Monvasivociws):, Dall, Bull37, Us S: Ni.) Mp. 162, 89. Distribution.—Hatteras to Honduras. Also West Florida, shallow water.
(Eutrochus) jujubinum var. rawsoni Dall, Bull. M. C. Z., 18, p. goo) 80;Wall and Simpson, Bull. U. S. Bish Com., 1, p. 444, 1901. Distribution.—Cedar Keys, Fla. and the Antilles.
(Eutrochus) yucatecanum Dall, Bull. M. C.Z., 9, p. 47, 81; 18, PaO Pi 24,f. 4,44, 89. Distribution.—Yucatan Strait, 640 fms. Also off Carolina coast, 15-32 fms.
(Dentistyla) asperrimum Dall, Bull. Mus. Comp. Zool., 9, p. 40, (eMitas Wargariia): 18, p. 373; 89; Pilsbry, Man., 11, p mone oo. Wall) Bill. 37, U, 5. N. MM. p. 162, 7890,
Distribution.—Hatteras to Barbados. Also West Florida. 100-177 fms.
Genus SQLARTELLA S. Wood amabilis Jeffreys, British Conch. 3, p. 300; 5, pl. 61, f. 6, (as ijocus)\: Walt, Bull. M. C)Z., 18, p, 378,89; Pilsbry, Man., Mime etsy pl 57; £4 52, °8o.
Distribution.—Yucatan Strait, 640 fms. Also Antillean and European (North Sea off Shetland Isls, 85-95 fms).
scabriuscula Dall, Bull. M.C. Z., 9, p. 41, ’81, (as Margarita); Peo eo Pl 2h ite TO. 10a, So; Bull. 27 Ul S. ING M. «5 Meta ple 20," f. 10,.7Oa, 89.
Distribution.—Southern part of the Gulf of Mexico, 539 fms., bottom temperature 39° F. Also off Cuba.
127 BULLETIN 38 157
aeglees Watson, Jour. Linn. Soc., 14, p. 704, 779; Chall. Rept. Gastr., p. 81, pl. 5, f.. 10, ’85; (as Margariia).) Wangiea?: M. C. Z., 9, p. 40, ’81; 18, p. 379, ’89; Pilsbry, Unie aoe 215, pl, 66,1. 18) Tepe se. Distribution.—Gulf of Mexico, 287-888 fms. Also off St. Vincent, W. I.
aeglees var. clavata Watson, Jour. Linn. Soc. London., 14, p. 705, Chall. Rept. Gastr., p. 82, pl. 5, f. 8, (as Margarita); Dall, Bull. M. C. Z., 18, p. 380,89; Pilsbry, Manan oe 218, ple 66, fi OSG.) 8a.
Distribution.—Yucatan Strait, 640 fms.; Culebra Isl., W. I. 350 fms.; Pernambuco, Brazil, 675 fms. Dall and Pilsbry regard this as a variety of aeglees.
lubrica Dall, Bull. M. C. Z., 9, p. 44, ’81, (as Margarita); 18, p. 382, pl. 21, f. 9, "89; Pilsbry, Man., 11, p) 3240 25, 26, 89; Dall, Bull. 37, U. S. N. MM.) p. eater: 9, 9a, *89. Distribution.—Cedar Keys, West Florida, to St. Lucia, W. I. 116-805 fms.
lubrica var. iridea Dall, Bull. M. C. Z., 18, p. 382, ’89; Pilsbry, Man., 11,~p, 324, 89; Dall, Bull. 37);.U..S. Nae ae 1889. Distribution.—Cape Fiorida, Gulf of Mexico, West Florida region. 193 fms:
lissoconcha Dall, Bull. M. C. Z., 9, p. 41, ’81, (as Wargarita); 18; p: 381. pl. 21,:f..8, 8a, ’29; Bull. U. S. N. Me) ete ee: OI fi 1S) (oa. Ol Distribution.—Gulf of Mexico, Lat 28° N., Lon. 88° W., at 331 fms.
Genus BASILISSA Watson
alta Watson, Jour. Linn. Soc. London, 14, p. 597, 797 Rept. Chall. Gastr., p. 100, pl. 7, 1,8, °86; Pilsbry, Mamiya ima:
es
a i (ty
ir 1. i )
158 MOLLUSCA OF THE GULF OF MEXICO 128
Poe ol 2681s.) 189: Dall) Ball) 37) M.S. NN. M., p.) 164, 1889.
Distribution.—Gulf of Mexico, West of Cedar Keys; the Antilles and Brazil. 339-rorg fms.
alta var.delicatulaDall, Bull. M. C. Z., 9, p. 48, ’81, (as Seguen- ea uss ps! Zod, pl.) 22, f.'2)) 89; Pilsbry,;Mamn... 11, ps 421, pine a A 8% Dall) Bull. 37,U. SiN. M., p: 164) pl. BIW 2.) 2a. SO. Distribution.—Gulf of Mexico to Tobago, W. I. 805 fms. superba Watson, Jour. Linn. Soc. London, 14, p. 598, ’79; Rept. Oba Gort... 101, pl. 7, f. 10, °86;) Pilsbry, Man., 11, p. Ae Aonse, 1.6.7, 789; Dall, Bull. 37,,U. 5. N. M., p::164, 1889. Distribution.—Cape York, Australia, type locality. Also Gulf of Mexico west of Florida. 400-1400 fms. (Ancistrobasis) costulata Watson, Jour. Linn. Soc. London, 14, p. Geo, Veou"Rept. Chall, Gastr,, p. 103, pl7, f)11,:°°86; Dall. Me MC! Z) 18. p>, 284, pl.'23, 1.4) 4a, 89; Pulsbry, Man., MMO A2o pl. 26. 0,3, Soe Mall, Ball. 37. Uy SN.) M.,p. GA pl 23. f.'4,. 4a, °89. Distribution.—Georgia to Culebra, W. I.
costulata var. depressa Dall,Bull. M. C. Zool. Harv. Coll., 18, p- Bateeo. Pilsbry, Man., trp. 428, pl. 6, f: 14) 15, 789; Dall, Bana WS. Noo Mit. pb.) 164, pl. 23. 'f.. 4, 4ay-4 89. Distribution.—Yucatan Strait, 640 fms. Also Antillean.
Genus UMBONIUM Link
bairdi Dall, Bull. Mus. Comp. Zool., Harv. Coll., p. 359, pl. 21, Hnoniean So. Pisbry, | Mam, ).01,) ps 457) ple).60;)22\15,) 6; 18809.
Distribution.—Yucatan Strait, 640 fms. Also Florida reefs, 200 fms.
129 BULLETIN 38 159
Genus COCHLIOLEPIS Stimpson
parasitica Stimpson, Proc. Bost. Soc. Nat. Hist., 6, p, 308, and fig., 758; Dall, Bull. M.C. Z., 18, p. 360, °So;"auita S. N. M., p.. 162, °89;)Tr..W. 1. Set. | 3)! pl ae
Distribution.—Charleston Harbor, S. Car. to Florida Keys and Tampa, West Florida. At Charleston this species lives under the scales of a large annelid. Pliocene, Caloosahatchie beds, Fla., rare.
striata Stimpson, MS.; Dall, Bull. M. C. Z., 18, p. 360, 1889. Bull, 37, U. S. N. M., p. 162, *80; Ur. (Wagers: Sci., 3, p. 419, pl. 23, £. 16, 17,92; Jacot, Jomn) Bietamem Soc., 1p. .638, /Pebs toe: Distribution.—Beaufort, N. Car. (Jacot); Egmont Key, near Tampa, West Fla. (Colonel Jewett). Upper Miocene of North Carolina.
Genus ADEORBIS S. V. Wood
adamsi Fischer, Journ. de Conch., p. 173, pl. 10, f. 11, 1875; Dall, Pr. U.S.’ N. M:, 6. p. 335, °83; Bush) Wee Acad. Sci., 10, p. 104, 1899-1900. Distribution.—Cited by Dr. Dall from Cedar Keys, West Fla.
Genus VITRINELLA C. B. Adams
mooreana Vanatta, Proc. Acad. Nat. Sci. Phila., 55, p. 758, 1903. Distribution.—Crooked Island, Calhoun Co., Fla.
hemphilli Vanatta, Proc. Acad. Nat. Sci. Phila., p. 24, Piya. ae TACT Wot yy Distribution. —Cedar Keys, Florida.
interrupta C. B. Adams, Monograph of Vztrinella, p. 6, 1850;
Journ. de Conch.,.'6, p.176,; Dall, Bull. 27, U. so ie 166,. 1889; Bush, Tr, Conn. Acad. Sci., 10, pp tog iia
*99-I900.
160 MoLLUSCA OF THE GULF OF MEXICO 130
Distribution.—Tampa, West Fla., in shallow water, to Jamaica.
Genus CIRCULUS Jeffreys
liratus Verrill, Tr. Conn. Acad. Sci., 5, p. 529, ’82, (Omalaxis); Spets2, 85, (Seneca): Bush, of. czt., 6; p. 404, 7°85; Bull. Rime een 220) 0. 240, pl. 12, 12,’93>. Ers.Conn, Acad) Seto, Pp. 125, pl. 23, f..7, 12-12b, 99-1900.
Adeorbts supranttidus var. orbignyt Dall, Bull. M. C. Z., 18, p. 27a) (in part); Tr. W. 1. 'S., 2, p. 344,:'92; Johnson, Meeviea) Bost. Soc. Nv T,7, p. 120, 1915; Not orbzgnyz Piecher, Journ. de Conch. ,6, pp. 173, 286, ’57.
Distribution.—New England to Hatteras and West Florida. Lower Miocene, Chipola beds, West Fla.
Note.—The true C7zrculus orbignyi Fisher is West Indian and very minute, only 1% mm. in diameter,while dzvatus is 2% mm. in diameter.
trilix Bush, Rept. U. S. Fish Com., for 1883, p. 584, 1885, (as meeca) ir. Cann. Acad: Sci.,°10, p. 127, pl. 22, £: 6, 10, Oa, 02, a-2, pl, 23, 1. 10, 15, 1899-1900.
Adeorbts supranitidus Dall, Bull. M. C. Z., 18, p. 278, ’89; Poors. INSU MW perso. ply ated. 7, 7a so, (in part). Not supranitidus Wood, Cat. 1842, Crag. Moll., p. rage pl 15, £. 5a-b, 1848.
Distribution.—Massachussetts to Hatteras and to Cedar Keys, West Fla. 15-25 fms: Pleistocene, Knapp’s No. 2 well, Terrebonne Parish, La., at 1519-1632 feet.
Genus LISSOSPIRA Bush
proxima’ Iryon, Manual Conch., 10, p. 98, pl. 33, f. 4, 1888, CO eosivea) Bush) Ir. Conn, Acad) Sct., 10,’ p..130, f. 4; pH 220T 3) 18og-190a.
rugulosum Verrill, Tr. Conn. Acad.,5, p. 533,’82,(Cyclostrema). Not of G. O. Sars, 1878.
£35 BULLETIN 38 16x
affine Verrill, Tr. Conn. Acad., 6, p. 1909, pl. 32) f)4a5)a4.46 Cyclostrema). Not of Jeffreys, 1883.
trochoides Dall, Bull. M. C. Z., 18, p. 393, 1880; Dal Bais? U.S. N. M., p. 166, ’89, (as Cyclostrema). Not trochoides (Jeffreys) Sars which is a variety of pettersen7 Friele.
Distribution.—Lat. 41° N., Lon. 65° W. to:-Matia5 Lon. 74° W. (Bush); West Florida (Dall).
Genus CHORISTELLA Bush
pompholyx Dall, Bull. M. C. Z., 18, p. 394, pl 28, f. 9, ’89, (as Cyclostrema); Bull. 37, U. S. N. .M., ps 166) pleat: ’89; Bush, Tr. Conn: Acad. Sci., 10, pp. (oq) 14q\mean- 1900.
Distribution.—Blake Station 2, Gulf of Mexico, type locali- ty. Also Fernandina, Florida, to Cuba. 294-805 fms. Note.—This species is referred tentatively to Choristella as Miss Bush suggested that it might prove referable to this genus.
Genus EPICYN!A Moerch
multicarinata Stimpson, MS., Dall, Bull. M. C. Z., 18, pp. 273, 392,89; Bull. 37, U. S..N. M., p. 166; 780, (as) Ve7777-ae Bush, Tr./Conn, Acad.,10,' p, 112,’ go-1eec, Distribution.— Hatteras to West Florida. Pliocene, Caloosa- hatchie beds,’ Fla. Upper Miocene, Duplin Co., N. Car. Perhaps identical with the Avchitectonica gemma of Holmes listed by Hilgard from the Lake Borgne borings, La.
Genus ETHALIA A. Adams
The following species are of doubtful generic position. They were excluded from -thalia by Pilsbry (Man. Conch., 11, P- 457) when he restricted that genus and gave A¢halia guamensis Quoy and Gaimard, as the genotype. Dr. Dall has referred me to Miss Bush’s revision (Trans. Conn. Acad. Sci., 10, p. 416, ’89-I900). Apparently she tentative- ly retains these three species in £ihalza.
162 MOLLUSCA OF THE GULF OF MEXICO 122
suppressa Dall, Bull. Mus. Comp. Zool., 18, p. 362, ’89; Bull. 37, oe IN Mi) Dp.) 160, 80,
Distribution.—Goodland Point, West Florida. Collected by Hemphill.
fesse Wall, Bull: Mus. Comp. Zool., 18, p: 361, pl. 28, f. 7, 8, ene dl) 27). WS.) No M.,) p..160, "89, Distribution. —Yucatan Strait, 640 fms. North Carolina coast, 12-65 fms.
cold eal Bulli Mi.'C, Z., Harv. Coll., 18, pl. 28, £..3,'5, 789; Bale. 3. IN.) Mp. D600, pls 28) f. 3, 5;' 89.
Distribution.— West Florida and Cuba, 310 fms.
Genus TEINGSTOMA A. Adams
cryptospira Verrill, Trans. Connecticut Acad. Sci. and Arts.,6, p. Come gea (as /cvorella).: Wall, Bull. 37; Ui 5. N:M., p. 160, onauen iT, Conn, Acad, ‘Sci., (10, p. 118, £1, 25 1899- nooor Wall) Tr. Wagner Inst. Sci., 3; p. 414, 92.
istribution.—Hatteras to Florida. Gulf coast at Cronked
Hetand’) Calhoun Co., Fla. 30-150 fms. leistocene of North Creek, Fla. Miss Bush regards this species as a true Tetnostoma.
Genus NERITA (Linneus) Lamarck
pelorenta Linnzeus, Syst. Nat., 10, p. 778, 1758; Reeve, Conch. ewer a!) pli 2.618) Sbi1855: Lryon,, Man.)\70, pl 24, pl. ater 7) Toso Wall Balliie7. U.S.) NY. M., ps 166,,,1889; Palivand’ Simpson ibull, UU.’ >.) Fish. Com; -1,\p.) 445, IgOl. Distribution.—Florida, St. Vincent and Texas region of the Gulf of Mexico.
tessellata Gmelin, Syst. Nat., p. 3685, 1792; Reeve, Conch, Keon.) o) pli) t: 432) 43a,'1855; Iryon, Man., 10; p.,24, pl.
133 BULLETIN 38 163
4, f. 71, °88; Dall, Bull. 37, U.S. N. M:, p \6n Vee ee and Simpson, Bull. U. S. Fish Com., 1, p. 446, 1901,
Distribution.—East Florida to Colon. Gulf coast, West Florida and Texas, shallow water.
versicolor Lamarck, Anim. s. Vert., 6, pt. 2, p. 193, 1822; Reeve, Conch. Icon., 9, pl. 12,.f. 56, a-d, 1855; Daly aaa U.S. N. M., p. 166, 89; .Dall and Simpson, Balliyis, ish, Com. 1) p., 246,.1¢01
Distribution.—Florida Keys and West Florida to Colon, the Antilles and Bermuda.
Genus NERITINA Lamarck
reclivata Say, Jour. Acad. Nat: Sci. Phila.; 2, p. 257; 1822) (as Theodoxus); Reeve, Conch. Iren., f£.'34 a-b,- 7557 aly or U.S. N. M., 8; p. 259, pl. 17, £..8, ’85; Tryon) iam ame: 39, pl. 12, f. 25-27, ’88; Singley, Fourth Ann. Rept. Texas Geol. Surv., p. 342, 1892; Mitchell, List. Texas Shells, pe eae Distribution.—St. Augustine to Martinique. Gulf coast: Mouth of Manatee River, Tampa, Cedar Keys and St.Mark’s, Fla., Point au Fer and Chandeleurs, La.; Port Lavaca and Rockport, Tex. Pleistocene, Knapp’s No. 3 well, Terre- bonne Parish, La. from 1-700 feet.
Note.—The variety jalmae Dall is found at Palma Sola, Texas.
virginea Linnzeus, Syst. Nat., ed. X, p. 778, 1758, (as WVerzia); Reeve, Conch, ‘Icon., 9,, pl: 21, f. 92, a-d, 38559) Danie U.S N. M.,'8, p. 259, 85; Tryon, Man.) 10) p, goto as f. 31-35, 37-43; Dall, Bull. 37,.U. S. N.' Np. Ge eee. Singley, Fourth Ann, Rept. Geol. Surv. Texas, p. 342, 1892.
Distribution.—Florida to Brazil. Gulf coast: Various local- ities of West Florida; Corpus Christi and Rockport, Tex.
164 MOLLUSCA OF THE GULF OF MEXICO 134
pupa Linnzeus, Syst. Nat., ed. X, p. 378, 1758; Tryon, Man., Io, Pemomenniai 2. 73° 88 Mall Ball.) 27. Uu Ss: No My p: L696, So. Distribution.—Charlotte Harbor, West Florida to Jamaica. Shallow water.
viridis Linnzeus, Syst. Nat., ed. X, p. 778, 1758; Tryon, Man., Pop ate ole 1S.) ft. 88. 1888: Vall Bull, 27.0.5. Ni M., Pe TOS. SO. Distribution.—Florida Keys to Barbados, W. 1. Also Texas region of the Gulf of Mexico.
showalteri Lea, Pr. Acad. Nat. Sci. Phila., p. 55, 1861; Tryon, Ame hON D5 3..lo i781. 82,)/ 88s. Dall) Bull. 37, U.S. ING Es. 168,789.
Distribution.—Alabama and West Florida, Fluviatile. sparsilineata Dall, Proc, U.S. N. M., 46, p. 236, 1914.
ivemimia.) sp. idet., Aldrich, Nautilus, 24,) pt: 11, p. 131, NOR Werorl: 26, pl.) f. 3,4, 1912.
Distribution.—Upper Miocene or Pliocene. Well near Al- exandria, La., at 49 feet; Pine Prairie La., well at 1540 ft. and well % mile S. E. of Pine Praire depot, in dump. Also Satilla River, Ga., and Burkeville, Texas. One of the most widespread and characteristic species of the brackish water formation, extending from Georgia through Louisiana to Texas.
Genus PLEUROTOMARIA Sowerby
(Peirotrochus) quoyana Fischer and Bernardi, Journ. de Conch., SOD Os Ol Suk roe reso: Wall, Bully MO 20) o. p..73, fom Crosse, Journ.’ de Conch., 22, p. 14, 782; Dall, Bull. Pee Ze ton Dagon (pls 20,1. 1) pl. ant 1) 1b, fe, ply 37 it Beg. bm a7. SN.) M.,, p. 168 same) pls.) and) figs.) 1889.
Distribution.— Yucatan, near Arrowsmith Bank, at 130 fms.
135 BULLETIN 38 165
Also off Barbados, 73 fms., and Marie-Galante Island, near Guadeloupe.
Genus FISSURELLA Bruguiere (Cremides) barbadensis Gmelin, Syst. Nat., p. 3729, 1792, (as Patella); Dall, Bull. 37, U.S. N. M.. p. '170)°sa nian Man., 12, p. 164, pl. 37, f. 40-49, ’90; Dall and Simgean, Bulk. U.S. Fish Comin,’ p. 248) 190m
Genus LUCAPINA Gray
suffusa Reeve, Conch, Icon., in errata, 1851, (as Fissurella); Dall, Trans, Wagner Inst. Sci., 3, p. 424, pli) 2ai\\b) jaeeneee
hondurasensis Reeve, Conch. Icon., f. 70, 1851.
canccllata Sowerby, in part, Thesaurus Conch., p. 200, pl. 8, (243). f. 187 only, ’62; Dall, Bull. 37, U.S... Nee "89; Pilsbry, Man., 12, p. 200, pl. 63, f. 1-3olageeeee ’90; Dall and Simpson, Bull. U.S. Fish Com.; 1, pi 44o, IgOl.
Distribution.—Marco, West Florida to Vera Cruz, Mexico. Also Antilles and South America to the Island of Fernando Noronha, Brazil.
Note.—As there has been some question whether the name suffusa Reeve or cancellata Sowerby should be applied to this species, Dr. Dall’s opinion was asked. He replied, June, 1921, ‘‘I find nothing to change inthe synonymy of Z. suffusa as given in the Wagner memotr.’’
adspersa Philippi, Abbild. u. Beschr., 2, p. 34, pl. 1, f. 3, 1845; (as Fissurella): Pilsbry, Man., 12,\p. 199, pl, G2. eeG@) ’90; Dall and Simpson, Bull. U. S. Fish Como i py ane, IQOl.
fasctata Pfeiffer. Fissurellidea fasciata (Pfeiffer) Dall, Bull. WS. SU ONT EME i ou aes eo): Distribution.—West Florida region of the Gulf of Mexico to Cuba and: St. (Croix) Wy 2,
166 MOLLUSCA OF THE GULF OF MEXICO 136
Genus FISSURIDEA Swainson
alternata Say, Jour. Acad. Nat. Sci. Phila., 2. p. 281, 1822, (as iyssireia): Pilsbry,’Man., 12; p. 211; pl. 37, £. 50-53; pl. 61, Pen 25\7’90; Dall and Simpson, Bull. U. S. Fish Com., 1, p. 450, TQOI.
Distribution.—Chesapeake Bay through the Antilles to Trinidad Isl. and Fernando Noronha, Brazil. Gulf coast: Cedar Keys, Fla.; Galveston, Corpus Christi, Texas; Vera Cruz and Progreso, Mexico. Also south of Nicaragua. I-50 fms. Pliocene of South Carolina.
cayenensis Lamarck, Anim. s. Vert., 6, pt. 2, p. 12, 1822; Reeve, Gaven icon, f..82; Dall, Bull, 37,.U. S$. N.’M:., p. 170, hee, husbry, Man. 12, p: 212) pl. 37, f. above 60, 1890.
Distribution.—Cedar Keys, Fla. to St. Lucia, W. I. Placed by Pilsbry in synonymy of alternata, but is more elongated and with straighter sides than that species. Perhaps to be classed as a variety.
Genus PUNCTURELLA Lowe
tiohum Dall, Bull: M..C. Z., 9, p. 76, ’81; 18, p. 403, pl. 26, f. Sea yoo.) Bull) 37) Ui S)'M. IN). py 168)" ply 26, f. (8, 8b, 1889.
Distribution. —Yucatan Strait, 640 fms.
watson Dall, Bull. M. C. Z., 18, p. 403; ’89;-Bull. 37, U. S. INES) p. 168,)’8o.
Distribution.—Off Yucatan, at 200 fms. Also off Cuba and Barbados.
emecularis Dall, Bull, M. GC. Z., 9, p. 75, 81; 18, p. 403, pl. 26, Pa) oO Billy) 37. My S. NoiM.,\p:) 1638) pl. 26, £.7; 7D; 789. |
Distribution.—Florida Strait and the West Indies.
137 BULLETIN 38 167
(Fissurisepta) triangulata Dall, Bull. M. C. Z., 18, p. 404, ’89; Ball. OS. Ni. Mee pp iro, 89:
rostrata Watson, Chall.Gastr., p. 48, pl, 4, f. 10, ’85. Not of Seguenza.
Distribution.—Yucatan coast, 200 fms. Also off Fernan- dina, Fla., and Culebra Isl., W. I.
(Cranopsis) asturiana Fischer, Journ. de Conch., 30, p, 51, 1882, (as Rimula); Watson, Chall. Gastr., p. 45, pl. 4, f. 4, 85; Dall Bull: MoyGly Za 1 8h. 4oa veo!
Distribution.—Yucatan Strait, 640 fms. Also off Havana,
Martinique, and Cape Florida, and in the Gulf of Gascony at 2018 fms.
Genu EMARGINULA Lamarck (Rimula) frenulata Dall, Bull. M. C. Z., 18, p. 406, pl. 28, f. 4, "89; Bull. 37, U.S.N.'M., p. 170, pl 28;))fia ee Distribution.— West Florida, Florida Keys and Hatteras, 6- 52 fms.
(Emarginula) compressa Cantraine, Bull. Acad. Roy. Bruxelles, Q, p. 2. 1835; Jeffreys, Pr. Zool. Soc., p. 679, ’83;Dall, Bull. M.°C. Z., 18, °p; 404, 89.
Distribution.—Yucatan Strait, 640 fms. Also off Havana, Barbados, and coast of Portugal. ‘Tertiary of Sicily.
*
168 MoLLUSCA OF THE GULF OF MEXICO 138
CASS) AWW Nie OO RA ORDER POLYPLACOPHORA
Genus LEPIDOPLEURUS Risso
pergranatus Dall, Bull. M.C. Z., 18, p. 414, ’89, (as Leptochz- ton); Bull. 37, U.S. N. M., p. 172, ’89; Dall and Simpson, DeeeUns. Fish Com.) 1)" p: 451, pls 58) f. 115, a-c, T9OT.
Distribution.—Gulf of Mexico, western Florida region, Porto Rico and Dominica, 114-1181 fms.
Genus CHAETOPLEURA Shuttleworth
apiculata Say, Amer. Conch., app., pt. 7, 1830, (as Chzton); ae eile an UW. Su Ne tM. pl) 172) pl. 51; f. 10, 1889; Filsory, Man., 14, p! /35,.pl. 13, f. 75-79, 1892; Johnson, Wee Wa iBoest: Soc. N. H.,:7, p. 8; 1915: Distribution.—Massachussetts to Haiti, Gulf coast, various localities of northwestern Florida and Chandeleurs, La. 0-30 fms.
Genus ISCHNOCHITON Gray
limaciformis Sowerby, Proc. Zool. Soc., p. 26, ’32, (as Chzton); DalkvBull, M. C. Zi, 18, p. 415; 89; Pilsbry, Man., 14, p. 57, pl. 16, f. 9-16, ’92; Dall and Simpson, Bull. U. S. Fish @om. 1, p. 452, 1901. Distribution.—Gulf of Mexico, southeastern part, Lat. 24° N., Lon. 83° W., at 37 fms. Also Key West and the An- tilles, Central America and Peru. Under stones at low water. |
purpurascens C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9, Meaai(as, Onzom): Dall Ball.) 37,.U. Si: N..M., pi/172,.” 89; Pepi Man TA pan ss, pl: 17,, f:) 23,24, 1892; Dall and eimpson, Bull. U.S. Fish Com., 1, p. 452, 1901.
Distribution.—Texas region of the Gulf of Mexico, Florida Keys, Jamaica and Porto Rico.
139 BULLETIN 38 169
papillosus C. B. Adams, Proc. Bost. Soc. Nat. Hist., p. 9, 1845; Dall, Bull.: 37,,U. S. N. M.; p.\172, 789; Piulsbay ieee p. 114, pl. 21, f, 40, 41, 1892; Dall and Simpson, Ball yes: Hish Com., Ijp) asa.enget.
Distribution.—Tampa, West Florida, to St. ‘Thomas, Wik Genus CHITON Linnzeus tuberculatus Linnzeus, Syst. Nat., ed. X, p. 667, 1758; Pilsbry,
Man., 14, p. 153, pl. 33, f. 58-60, 1892; Dall and Simpson, Ball Us Si Bish Com. p.1453, 1s:
Distribution.—Texas and Florida to Trinidad and Ber- muda.
marmoratus Gmelin, Syst. Nat., p. 3205, 1792; Dall, Bull. 37, U. S. N. M., p.172, ’89; Pilsbry, Man, 14, pa w5eee es f. 72-76, 1892; Dall and Simpson, Bull. U. S. Fish Com., 1, P- 454, 1901. Distribution.—Texas, and the Antilles to Barbados. South to Cartagena, Colombia.
Genus ACANTHOPLEURA Guilding
granulata Gmelin, Syst. Nat., p, 3205, 1792, (as Chzton); Pilsbry, Man., 14, p. 227, pl. 50, 1892; Dall and Simpson) Balla: ©. Fish-Com., 1, p:454;' 1901.
ptcea Gmelin, Dall, Bull.\37, U.S. (N. M.) pi wgaiee:
Distribution.—Charlotte Harbor, West Fla., to Trinidad, Bermuda and Bahamas.
Genus ACANTHOCHITES Risso
spiculosus Reeve, Conch. Icon., pl. 9, f. 47, 1847; Dall, Bull. 37, U.S. N. M., p. 174,’ 780; Pilsbry, Man., 15, pizzas f. 60-62, 1893; Dall and Simpson, Bull. U.S. Fish Com., I. pi 455, 1908:
170 MOLLUSCA OF THE GULF OF MEXICO 140
Distribution.—Cedar Keys, West Florida, to Barbados. pygmaeus Pilsbry, Tryon’s Manual Conchology, 15, p. 23, pl 13,
fies SO.) 1893.
Distribution.—Cedar Keys and Key West, Fla.
I4t BULLETIN 38 171
CLASS GEPHAL GP am. ;
ORDER DIBRANCHIATA
Genus ARGONAUTA Linnzeus
argo var. americana Dall, Bull. 37, U. S. N. M., pp, 174, 200, pl. 43, f. 1 a-b, pl. 64, f. 142b, pl. 67, f. 2-3, 1880; ele om Occ. Pa. Bost. Soc.’ Nat. Hist.;\7; p.22e.smeiee
Distribution.— Massachussetts north to 43° N. Lat., and south to the Antilles and doubtfully to Brazil. Gulf of Mexico, West Florida region.
Note.—The typical Avgonauta argo Linnzeus, Syst. Nat., ed. X, p. 708, 1758, is Mediterannean. It has two or three times as many radial folds and carinal nodules as the varie- ty americana.
Genus SPIRULA Lamarck australis Lamarck, Anim.s. Vert. 7, p. 600, 1822; Encycl. Me'’th., pl. 465, £. 5; a,b; Dall, Science, n: s., 3, pp.1242- 245m,
Dall and Simpson, Bull. U.S. Fish Com., 1, p/ 250) piiaa: fA.) 00On.
Se ee ee ee
spirula Linneeus, in part.
peront (Lamarck) ‘Dall, Bull. 37, U. 5. No M., piiagaiee 63) Taro.
Distribution.—Pelagic. West Indies and the Gulf of Mex- ico, shells being washed ashore along West Florida coast and at Cameron, La., Galveston and Corpus Christi, Texas. A specimen with all the soft parts was also taken from the
172 MOLLUSCA OF THE GULF OF MEXICO 142
mouth of a fish trawled by the Albatross between the Mis- sissippi delta and Cedar Keys, at 324 fms. The Spzvula had just been seized alive by the fish and the soft parts, pre- viously imperfectly known, were complete. See Science, 1896.
Migere ‘are Several forms of Spzvula. S. spzrula, of Linnzeus was a composite, according to Dr. Dall, and since the species cannot be differentiated by the shells alone no one can tell exactly which species Linnzeus had although he gave America as the habitat. Huxley and Pelseneer (Chal- lenger Rept.) fixed the name australis Lamarck on the An- tillean species.
Note.—Argonauta and Sptzrula are the only shell-bearing Cephalopods reported from the Gulf of Mexico. Of other forms, the Brazilian, Lolzgo brevis Blainville is cited from Cameron and elsewhere on the Louisiana coast, and ZL. hemiptera Howell from Florida and the Gulf of Mexico.
BULLETINS OF
AMERICAN PALEONTOLOGY
Vol. 9
The Miocene of Northern Costa Rica WITH — On Its GENERAL STRATIGRAPHIC RELATIONS BY
A. A. OLSSON
. Harris Co. Cornell University, Ithaca, N. Y. ASS A"
sh Bie
®
ms
iE ie Ave EO aie
Maes iV Or COS hA “RICA
d : ‘ , . . ‘ oes * .
eee OG ENE OF NORTHERN
CEST A Elia A
WITH NOTES ON ITS GENERAL STRATIGRAPHIC
ALT,
EV
RELATIONS
BY
Ae Ae OLSSON
PRELIMINARY REMARKS
GENERAL STRATIGRAPHY OF NORTHERN CosTA RICA
THE CosTA RICAN MIOCENE A. THE USCARI STAGE B. THE GATUN STAGE C. THE USCARI - GATUN UNCONFORMITY
CORRELATION A. SANTO DOMINGO MIOCENE B. BOWDEN BEDS, JAMAICA C. MEXICAN MIOCENE D. THE East Coast MIOCENE
THE CosTa RicA GEOSYNCLINE
EF Costa RicA MIocENE—OLSSON 7
PRELIMINARY REMARKS
The fossil collections and field information, on which this monograph is based, resulted from over two years of field work by the writer in Panama and Costa Rica. The writer’s thanks are due to the Costa Rica Oil Corporation for permission to pub- lish; and he is most grateful for assistance, and encouragement received in the course of this work from Drs. A. C. Veatch and D. F. MacDonald of the Sinclair Exploration Company, to the Officials of the National Museum of Costa Rica and Professor J. Fidel Tristan of San Jose’.
The collections were studied in the Paleontogic Laboratory of Cornell University, the facilities of which, Professor G. D. Harris, most generously placed at my disposal. Cornell University, January, 1922: ALA OLSSON
181 Costa RicA Mr0ocENE—OLSSON 9
GENERAL STRATIGRAPHY OF NORTHERN Costa RICA
The northern or Atlantic slope of Costa Rica, the so-called “Tierra Caliente’’ is largely composed of Tertiary sedimentary rocks, which except for some folding, dip away from the slopes of the Cordillera Central towards the sea. ‘To the east, this sedimentary belt may be traced into the Province of Bocas del Toro of western Panama, where its upper and highly fossilifer- ous beds are well developed and exposed on the islands of the Chiriqui Lagoon. This Tertiary sedimentary belt ends some 35 to 40 miles east of the city of Bocas del Toro. ‘The Miocene sedimentary rocks of Batun and of the northern Canal Zone form a small independent unit, which begins about 30 miles west of Colon and ends close to the eastern limits of the Canal Zone.
In a general way, the stratigraphic succession in northern Costa Rica is as follows:
Pleistocene Slightly elevated coastal swamps, with depositsof cross-bedded sands, clay-marls etc.
Pliocene Certain reef-limestones composed of coral and hydrozea re- mains; also marls, blue lignitiferous clays, blue coarse sand- stones, and a predominating hard conglomerate, composed of large boulders of igneous rocks (the so-called boulder-clays of several writers on Costa Rican Geology).
Miocene The Gatun Stage or Formation. Coral-reef limestones, (near the coast), marls, more generally a blue or blue-black sand- stone, dark colored shales, sandy and impure limestones, ligni- tes and conglomerates, Generally very fossiliferous.
The Uscari Stage or Formation. Usually dark-colored shales but with some sandstones, limestones and conglomerates. The smaller types of foraminfera are usually quite abundant, and are associated with fish-scales, and small echinoid spines, with mollusks generally rare.
Oligocene Sandstones, shales and a complex of hard, often semicrystal- line limestones which pass through all possible gradations to coarse arkosic sandstones. The most abundant and characteris- tic fossils are the Orbitoidal foraminifera (Lepidocyclina). The
1O BULLETIN 39 182
age of these beds probably ranges from Lower to Upper Oligocene.
THE Costa RICAN MIOCENE
An important part of the stratigraphic succession of north- ern Costa Rica 1s composed of rocks of Miocene age, whose thickness varies according to locality and completeness of sec- tion, from a few hundred to several thousands of feet. ‘They are divisible into two stages or formations by an unconformity or disconformity of greater or less degree. In their natural order of sequence, these divisions of the Costa Rican Miocene are:
Gatun Stage or Formation: Middle and Upper Miocene
Uscari Stage or Formation: Lower Miocene
THE USCARI STAGE
The Uscari stage or formation derives its name from Uscari creek in the Talamanca valley of eastern Costa Rica. It con- sists principally of soft, dark-colored shales and because of their slight resistance to denundation, their outcrop frequently forms wide valleys and interior basins. These shales were deposited in quiet and moderately deep water, and their fauna is largely composed of the smaller types of foraminifera, such as Globig- evina, Textularia, Lagena, Nodosaria and Miliola, but the large Orbitoidal Lefzdocycline, abundant in the underlying Oli- gocene, are absent. Mollusks are very uncommon in this forma- tion. Thesmall fauna described by Gabb from Sapote, Costa Rica, belongs to the lower part of the Uscari formation.
THE GATUN STAGE
The upper division of the Costa Rican Miocene is equivalent in part to the Gatun formation of the Canal Zone. The typical Gatun, so named from its well-known exposures at Gatun and in which a part of the Panama Canal and the locks at Gatun were excavated, lies in an independent sedimentary basin across the
183 Costa Rico MIocCENE--OLSSON II
northern end of the Canal Zone and in the adjacent parts of the Province of Colon. There the Gatun formation consists of blue sandstones, dark-colored shales, fuller’s earth beds etc., generally less than 500 feet thick. Many of its beds are very fossiliferous. The mollusks of the Gatun stage of Panama have been studied by Dall, Toula, Brown, Pilsbry and Cossmann.
In Costa Rica, the Gatun is very much thicker than in the Canal Zone and represents a longer depositional period. Paleontologically, the Gatun of the Canal Zone, seems to repre- sent only the lower part of the formation as developed in Costa Rica. It is largely of sandy character, more resistant to denu- dation than the Uscari shales and gives rise to a more rugged and hilly topography. Although typically sandy, it also contains locally shales, lignites, conglomerates and coralline limestones. In some localities the limy members are of major importance. The formation is characterized by marked lateral variation in which respect it differs strongly from the much more uniform shales of the Uscari stage. ‘This feature of lateral variation gives rise to two important phases: that of the more typical Gatun, in which the beds are largely sandy in character, with some lignitic and coarsely conglomeritic members, and carrying the usual near shore Gatun fauna; and the coralline phase, in which the beds are more marine, with coralline limestone and richly fossiliferous marls perdominating, and with no lignitic beds.
Along the inner margin of the Miocene belt, the Gatun be- longs to the first and more typical phase; while the coralline type is more common along the coastal zone, as at Port Limon and Bocas del Toro. Coralline rocks may occur at any horizon in the Gatun succession or it may locally be composed of mas- sive coral-reef limestones. Such is the case in the Limon Penin- sula, and such coralline limestones, by most observers, have been regarded as of Pliocene or even of more recent age. At Limon,
T2 BULLETIN 39 184
this confusion has partly arisen from the fact that these Mio- cene limestones are overlain by the second series, of Pliocene age, which partly yielded the large fauna described by Gabb, published shortly after his death in 1881. Dall later contribut- ed to the discussion of this fauna. The truly Miocene limestones of Limon may be seen in good exposures along the sea-front west of the city. There they are massive in character, hard and recrystallized to the extent that all traces of organic remains have usually been obliterated or rendered uncertain of determin- ation. However, some sandy or marly beds are occasionally found intercalated in these coralline limestones, and from these layers we have collected a large fauna of unquestionable Mio- cene affinities, correlating closely in age with the Bowden beds of Jamaica.
The fauna of the Costa Rican Gatun is largely molluscan. The following paleontological study records 334 species. Future collecting will doubtless greatly add to this list, as several species comnion it the Canal Zone, still remains undiscovered in Costa Rica. ‘The smaller types of foraminifera abound in the more shaly phases of the formation, as well as several species of echin- oids, belonging to the genera Clypeaster and Eucope. Corals of simple and compound types occur principally in the coralline phase.
THE USCARI - GATUN UNCONFORMITY
At the close of Uscari time the Miocene sea was with- draw from the greater part of Costa Rica, but during the Gatun, it returned. These conditions are indicated by a stratigraphic break between the two stages, a complete change in their lithologic composition, the occurrence of Gatun beds in areas which had received no deposition during Uscari times and a change in their respective faunas. Although the full strati- graphic details of this important subject cannot be presented at the present time, attention is called to it, asthe outstanding fea- ture of the Miocene stratigraphy of Panama and Costa Rica; and because of its bearing on correlation.
185 Costa Rica MIocENE —OLSSON 13
In many parts of northern Costa Rica, the Gatun stage carries at its base a thick and persistent conglomerate, formed of coarse cobbles of igneous rocks, or of finer pebbles with alternat- ing layers of sandstone. This conglomerate generally rests dis- conformably on the Uscari, but in some cases is unconformable on these or older rocks. This basal conglomerate is generally firmly cemented and because of its superior hardness, may be- come an important ridge former. It marks the site of high water-falls along many streams. Fossils of common Gatun types are abundant at many localitiesin this basal conglome- rate’.
Gatun rocks are frequently found in areas which contain no Uscari beds. In such cases, the Gatun is found resting, with strong unconformity, on older rocks. This condition is seen in the Canal Zone and along the borders of the sedimentary belt of the western Panama. It indicates, either a complete erosion of the Uscari shales during the Uscari-Gatun interval or, what is more likely, a greater transgression of the Gatun sea. This uncomformable relation of the Gatun upon older sedimentary beds may be seen in the Chagres Spillway in the Canal Zone, where the underlying rocks are sandstones, probably equivalent to the Mt. Lirio of Lower Oligocene age. In the Chiriqui La- goon country, of western Panama, the base of the Gatun is frequently a coral-reef limestone, which rests directly tpon the older and more disturbed rocks.
Sen i CePA hee iy yf et mist
The present systematic study enumerates 334 species of molluska of which the majority belong to the Gatun stage. A few of these species have not been found in Costa Rica or west- ern Panama, but are common in the Canal Zone, and are includ- ed here for the sake of comparison with Costa Rican species. To these we may add the 18 species described by Gabb from Sapote, Costa Rica, but not included in our collections and the following species listed below from the Canal Zone. ‘This brings
14 BULLETIN 39 186
the total known Miocene fauna of Panama and Costa Rica to 396 species.
Volvulella micratracta Brown and Pilsbry
Ringtcula hypograpta Brown and Pilsbry
Conus haytensis Sowerby Corbula gatunensis ‘Toula Conus domingensis Sowerby Corbula sphenis Dall Cruns consobrinus Sowerby Corbula sericea Dall’
Conus emulator Brown and Pilsbry
Dritlia vaningenit Brown and Pilsbry
Drillia gatunensts 'Toula Solecurtus strigillatus Linné Drillia fusinus Brown and Pilsbry
Drillia Zooki Brown and Pilsbry
Drillia enneacyma Brown and Pilsbry Cythara heptagona Gagg
Marginella gatunensis Brown and Pilsbry Anachts fugax Brown and Pilsbry
Murex polynematicus Brown and Pilsbry Murex gatunsis Brown and Pilsbry
Typhis Gabbi Brown and Pilsbry
Malea camura Guppy
Cyprea Henekint Sowerby, var.
Littium nugatorium Brown and Pilsbry Natica bolus Brown and Pilsbry
Natica canalizonalis Brown and Pilsbry Sigaretus gatunensts Toula
Sigaretus (Eunaticina) Gabbi Brown and Pilsbry Chetlea princetonia Brown and Pilsbry Nucula (Acila) isthmica Brown and Pilsbry Pecten effosus Brown and Pilsbry
Pecten operculariformis ‘Toula
Pecten reliquus Brown and Pilsbry Amusium Toule Prown and Pilsbry
Ostrea gatunensis Brown and Pilsbry Cardium durum Brown and Pilsbry Cardium gatunensis Dall
Tellina equiterminata Brown and Pilsbry
187 Costa RicA M1ocENE—OLSSON 5
Chione tegulum Brown and Pilsbry Pitaria cora Brown and Pilsbry
Dosinia delicatissima Brown and Pilsbry Petricola mitlestriata Brown and Pilsbry Cycinella gatunensis Dail
Thracia gatunensis Toula
The Uscari fauna is still poorly known, but it includes sev- eral very distinctive species of which the following are most important:
Amusinm Lyonit Gabb
Arca dariensts Brown and Pilsbry also Lower Gatun Clementia dariena Conrad also Lower Gatun Cancellaria (Aphera) islacolonis Maury
Neverita nereidis Maury
Sconsta cocleana Olsson
Scaphella costaricana Olsson
Ptychosalpinx ? dentalis Olsson
Dentalium uscarianum Olsson
The Gatun is the great fossil bearing formation of northern Costa Rica and Panama, and is characterlzed by a rich and high- ly diversified fauna. This fauna at the present numbers nearly 400 species, but additional collecting will doubtless add consid- erably to this number. As we have already noted the Gatun beds present two important facies, the sandy and highly clastic phase of the typical Gatun, and the deeper water type, illustrat- ed by the coralline limestones and their associated marls. ‘These lithological types have their own more or less peculiar and dis- tinctive faunas.
The typical Gatun is the more common phase and is charac- terized by a molluscan fauna of heavy, thick-shelled species of littoral forms. In Costa Rica, the following are the most com- mon and characteristic:
Arca MacDonaldi Dall Marginella MacDonaltdi Dall Arca Pittiert Dall Voluta Alfarot Dall Pecten levicostatus Toula Strombus pugtloidis Guppy
16 BULLETIN 39 188
Cardium stirtatum Brown, Pilsbry Stormbus gatunensis ‘Toula
Chione Rowell Olsson Natica Guppyana Toula Chione mactropsis Conrad Turritella alithra Conrad and its Dosinia acetabulum Conrad varieties
Conus molis Brown and Pilsbry
The coralline phase is developed at several localities along the coast, most important of which are Port Limon and Bocas del Toro. Here the coralline limestones and associated marls form the greater part of the Gatun section, but coralline rocks may occur, interbedded with typical Gatun rocks, at any strati- graphical level.
These coralline limestones were formed, mainly as barrier reefs some little distance from the old Gatun shore-line. Natur- ally they offered very different habitat conditions from those of the typical near-shore and often lacustrine Gatun. Hence we find a fauna of deeper-water affinities with Pleurotomids and thin-shelled Cones, as the most important and characteristic element. In localities where these coralline rocks are exclusive- ly developed, this fauna bears very little relation to the typical Gatun fauna, although the two are contemporaneous. The following are species of the coralline phase:
Pecten coralliphila Olsson Mitra Alamagrensis vat. Antigona Harrisiana Olsson coralliphila Olsson Gafrarium limonensis Olsson Mitra poas Olsson
Large number of Pleurotomids Marginella avena Valennencies
Conus planiliratus Sowerby Olivella limonensis Olsson Conus limonensis Olsson Phos elegans, variety limonensts Conus stenostomus. Sowerby Olsson Turritella exoleta Linné Siliquaria modesta Dall,
var. /imonensts Olsson limonensis var. Olsson
About 13 per cent. of the Gatun fauna is identical or closely related to the recent species, the majority of which are found living along the present Caribbean coast, certain others only on the Pacific side in the Panama Province and a few common to both. ‘This large percentage of recent forms corresponds closely
189 Costa Rica MrocENE—OLSSON 7
with that characterizing the Chesapeake Miocene fauna of the eastern United States and shows their close agreement in age. The recent species and their. closely related forms in the Gatun fauna are as follows:
Arca occidentalis Phil. West Indian
Arca umbonata Lamarck West Indian
Arca candida Gmelin West Indian
Arca auriculata Lamarck West Indlan
Ostrea megodon Hanley Pacific
Anomia simplex d’Orb. West Indian
Phacoides vadians Conrad var. meditoamericanus Olsson West Indian
Phacoides trisutcatus Conrad West Indian
Glycymeris castaneus Lamarck West Indian
Glycymerts decussatus Linné West Indian
Divartcella quadrisulcata d’Orb. West Indian
Cardium medium Linné West Indian
Cardium spinosum var. Turtont Dall East Atlantic Cardium serratum Linné West Indian Pitaria circinata Born and its var. alternata Broderip West Indian and Pacific Aniigona multicostata Sowerby Pacific Antigona rugosa Gmelin West Indian and Pacific Macrocallista maculata Linné West Indian Chione grus Holmes West Indian Tellina crystallina Chem. West Indian and Pacific Strigilla pistformis Linné West Indian Mactra exoleta Gray Pacific Mactra alata Spengler West Indian Labtosa lineata Say West Indian Corbula Dietztiana C. B. Adams West Indian Tellidora cristata Recluz West Indian Conus proteus Hwass West Indian Conus floridanus Gabb var. costaricensis Olsson West Indian Terebra benthalis Dall var. bocasensts Olsson West Indian
18 BULLETIN 39 190
ZTurris albida Perry West Indian
Glyphostoma dentifera Gabb West Indian
Acteocina bullata Kiener var. costaricana Olsson West Indian
Acteocina recta d’Orb. West Indian
Volvula cylindrica Gabb West Indian
Ancistrosyrinx elegans Dall variety West Indian
Marginella avena Valen West Indian
Murex messorius Sowerby West Indian
Bursa crassa Dillwyn West Indian
Natica canrena Linne West Indian
Archtitectonica granulata Lamarck West Indian nnd Pacific
Astralium brevispinum Lamarck, variety basalts ; Olsson West Indian
Astralium caclatum Gmelin West Indian
Sistrum nodulosum C. B. Adams West Indian
Latirus tufundibulum Gmelin West Indian
Northia northig variety mtocentca Olsson Pacific
Crepidula plana Say West Indian
Xenophora conchyliophora Born West Indian
Oliva testicea Lamarck var. costaricensts Olsson
It is but natural to expect that the Miocene beds along the northern or Caribbean coast of Central America, should contain a fauna largely ancestral to that of the present Caribbean, but along with these strictly Caribbean types, we also find several species, identical or closely related to forms now resticted to the Pacific side. Their occurrence in Caribbean deposits points to a connection between the Atlantic and Pacific oceans during Mio- cend time and allows for a certain intermingling of their faunas. Geological observations indicate that there were at least two such straits crossing Central America during the Miocene, the main one across Costa Rica, and a second through the Darien of eastern Panama.
This Pacific element of the Gatun fauna comprises many of the most common and characteristic species. Certain of these species had a wide distribution in the West Indies during Mio-
Igt Costa Rico M1ocENE—-OLSSON 1g
cene time and have been noted by several writers. They comprise the following species from Panama and Costa Rica, listed together with the Pacific analogues:
Panama or Costa Rica Pacific Analogue
PAE ME OLFICIA SOWECIDY...0603.005 2200000 Arca grandis Sowerby Arca (Noetia) MacDonaldi Dall......Adrca reversa Sowerby Ostrea megodon Hanley
Mactra exoleta Gray
WAOGEV A) CSTFELLANG. OISSON...0.....06h eens ss Mactra (Harvella) elegans Sowerby Semele laevis Sowerby var. costaricensis Olsson........Semele laevis Sowerby
Macoma panamensis Dall variety canalts Olsson.../Jacoma panamensis Dall
Macoma gatunensis Toula.............. Macona elongata Hanley Telling crystallina Chemnitz.......2.... also West Indian Antigona multicostata Sowerby
LANSONA FWZOSG GMeliny...0..6. 0.02.2. also West Indian
Pitaria circinata Born variety alternata Broderip Cyclinella subquadrata Hanley variety guitana Olfson...... Cyclinella subguadrata Hanley
Cyclinella beteyensis Olsson..........06. Cyclinella Kroyert Philippi COMMS: FECOZNILUS ‘GUPPY..6.4....seecees- Couus pyriformis Reeve Turricula lavinoides Olsson............ Turricula lavina Dall Cancellaria islacolonis Mautry.......... Cancellaria tesselata Sowerby Cancellaria toroensis Olsson............ Concellaria tuberculosa Sowerby Cancellaria Plummeri Olsson.,......... Cancellaria bullata Sowerby Mitra Swainsoni Brod. variety
limonensis Olsson...... Mitra Swainsonit Broderip UNEOCHILA HEKCIAIS’ MAULy | s.4..4.'5) <o.d0s-: Neverita glauca Humbolt Northia northiae Gray variety
miocenica Olsson...... Northia narthiae Gray PAGED COMLUIA GUPPY so. dicrencdsleosesesees Malea ringens Swainson Oliva testacea Lam, variety costaricensts Olsson...... Oliva testacea Lam
With the closing of the Miocene straits, this Pacific element gradually had to give way before the development of the West Indian fauna, but it is still to be seen in such species as, 7ellina crystallina Chem. Architectontca granulata Lamarck, which have a distribution along both coasts. Moreover the Pliocene
20 BULLETIN 39 192
and Pleistocene beds of Central America contain a well-marked
Pacific element in such species as Vorthia northie Gray Oliva testacea Lamarck and FPecten ventricosus Sowerby.
CORRELATION
Miocene rocks are now known from a great many localities in the West Indies and the general Caribbean area of Central and South Anierica. In these areas they comprise a very important series of sedimentary formations. Beds equivalent to the Gatun, appear to be the most widespread and generally are highly fossil- iferous.
The following chart showing the equivalence of the Miocene horizons in the West Indies and elsewhere to those of Central America, does not differ materially from the more recent correl- ations proposed by Maury, Cooke or Vaughan, except in the ad- dition of the Uscari to the Lower Miocene of Panama and Costa Rica. The Gatun formation is here recognized as being of Mid- dle Miocene age with possibly some of its upper beds having Upper Miocene affinities. The Gatun, at its type locality in the Central Zone, is not more than 500 to 600 feet thick. In Costa Rica it is very much thicker, and its higher beds may range into the Upper Miocene. It is the exact equivalent of the Chesapeake Miocene of the eastern United States. In this correlation, I have been guided not only by the paleontologic evidence, but also by the field relations of the formations, studied over a wide rage of country in Costa Rica, and in Panama as far east as the Colombia frontier.
SANTO DOMINGO MIOCENE
In Santo Domingo and in Haiti, Miocene rocks are exposed in several areas, but attain their best development along the val- ley of the Rio Yayue del Norte in the northern part of the is- land. These beds are highly fossiliferous and because of the fine perservation of their fossils, have repeatedly attracted the atten- tion of paleontologists.
The earlier writers dealing with the paleontology of the Do- minican Miocene, believed that it represented but a single
193 Costa RicA MIOCENE—OLSSON oT
Stratigraphic unit, and Gabb insisted that no division was possi- ble. Dr. Maury however, as a result of paleontologic studies from collections carefully made in the field, was able to divide the Dominican Miocene into two formations, which she called the Gurabo and the Cercado. ‘To these Vaughan and his as- sociates, in more recent stratigraphic work on the island, have added 4 other subdivisions. The present recognized subdivisions of the Dominican Miocene are:
Mao clay MIDDLE Mao Adentro limestone Gurabo formation
(Sconsia laevigata zone)
Cercado formation (Aphera islacolon7s zone) Baitoa formation and Bulla conglomerate
MIOCENE
LOWER
The relation of the Costa Rican and Panama Miocene to that of Santo Domingo is extremely close; there being about go species in common. Of this number about 21 are recent, about 26 species appear to be restricted to the Gurabo, some 27 species are common to both the Gurabo and Cercado formations, and only r3rspecies are confined to the Cercado. The list of of Costa Rican and Panama species in the Dominican Miocene, is as follows:
Nucula tenutsculpta Gabb Cercado and Gurabo
*Arca occidentalis Phil Cercado and Gurabo * Arca umbonata Lamarck Cercado
Arca Maurye Olsson Gurabo
Arca Henekeni Maury Cercado
Arca goltoyaquensis Maury Cercado and Gurabo Arca chiriquiensis Gabb Gurabo * Arca auriculata Tamarck Gurabo
~ *Recent
22
BULLETIN 39 194
Glycymeris jamaicensts Dall Cercado Lime@a solida Dall Gurabo *Ostrea megodon Hanley Gurabo Spondylus gumancemocon Brown and Pilsbry, Gurabo *Anomta simplex d’Orb Gurabo Placunanomia lithobleta Dall Gurabo Pteria tnornata Gabb Cercado and Gurabo “Cardium serratum Linné Cercado and Gurabo Cardium dominicensis Gabb *Cardium medium Linné Gurabo Protocardia gurabica Maury Cercado *(Wactra alata Spengler Cercado Antigona tarquina Dall? Cercado Antigona Blandiana Guppy Gurabo *Pitaria circinata Born Cercado * Tellina crystallina Chemnitz Cercado Tellina cercadica Maury Cercado *Strigilla pistformts Linné Cercado Semele Claytont Maury Cercado Venericardia scabricostata Guppy Cercado and Gurabo Echinochama yaguensts Maury Gurabo Corbula vieta Guppy Cercado and Gurabo Corbula viminea Guppy Cercado *Acteocina recta d’Orb Cercado Acteocina triticum-tritonts Maury Cercado and Gurabo * Volvula cylindrica Gabb Cercado Bullaria pau percula Sowerby Cercado and Gurabo Ringicula domintcana Maury Cercado Terebra spirtfera Dall Cercado Terebra bipartita Sowerby Gurabo Terebra hattensts Dall Gurabo Terebva Berlinere Maury Gurabo Conus haytensis Sowerby Cercado and Gurabo >? Sewalli Maury Gurabo
*Recent
195
nN)
ws
Costa Rica MIocENE—OLSSON
Conus stenostomus Sowerby Gurabo
Conus proteus Hwass Cercado and Gurabo Conus recognitus Guppy Cercado and Gurabo Conus multiliratus Bose Gurabo
Conus gractlissimus Guppy Cercado and Gurabo Conus marginatus Sowerby Gurado
Conus consobrinus Sowerby Cercado and Gurabo Conus domingensis Sowerby Gurabo
* Turris albida Perry Cercado and Gurabo Drillia venusta Sowerby Gurabo
Drillia consors Gurabo
Cythara heptagona Gabb Gurabo
*Glyphostoma dentifera Gabb Gurabo
Cancellaria Maurye Olsson Cercado and Gurabo Cancellaria Rowelli Dall
Cancellaria epistomifera Guppy Cercado and Gurabo Cancellaria islacolonis Maury Cercado
Oliva brevispira Gabb Cercado and Gurabo Oltva cylindrica Sowerby Cercado and Gurabo Olivella muticoides Gabb Cercado and Gurabo Mitra rudis Gabb Cercado and Gurabo
Mitra longa Gabb Cercado and Gurabo *Latirus infundibulum Gmelin Gurabo Melongena consors Sowerby Cercado and Gurabo Alectrion losquemadica Maury Gurabo
Metula cancellaia Gabb Gurabo
Murex cornurectus Guppy Cercado and Gurabo "Murex messorius Sowerby Gurabo
Typhis alatus Sowerby Gurabo
NDistortio simillima Sowerby Gurabo
KBursa crassa Dillwyn Gurabo
Cassts sulcifera Sowerby Cercado and Gurabo Sconsia levigata Sowerby Gurabo
*Recent
24 BULLETIN 39
Dolium camura Guppy
196
Gurabo
Dolium elliptica Pilsbry and Johnson Gurabo
Cypraca Henekent Sowerby Cypraca dominicensts Gabb Strombus pugtloides Guppy Serpulorbis papulosa Guppy
Petaloconchus sculpturatus H.C.Lea *Architectonica granulata Lamarck *Xenophora conchyliophora Born
KNVatica canrena Linné Natica Youngi Maury Polinices subclausa Sowerby
Polinices Stantslas-Meuntert Maury
Neritina viridemaris Maury Neverita nereidis Maury
Cercado and Gurabo Gurabo | Gurabo
Cercado and Gurabo Cercado and Gurabo Cercado
Gurabo
Cercado and Gurabo Cercado and Gurabo Cercado and Gurabo Gurabo
Corcado and Gurabo Cercado
Most of the above listed species belong to the Gatun forma- tion. Cancellaria (Aphera) tslacolonis Maury, Neverita nereidis Maury and Corbula viminea Guppy occur in the Upper Uscari and in the Lower Gatun or transitional beds of eastern Costa Rica. This leaves but the following 8 Cercado species in the Gatun fauna, or a proportion of about 3.25 to 1. | Winter Cay Winter Cay
Arca Flenekent Maury Glycymeris jamatcensts Dall Protocardia gurabica Maury Antigona tarquinia Dall? Tellina cercadica Maury Semele Claytont Maury Corbula viminea Guppy
—— ee
*Recent.
Saury Saury Hotel
Water Cay and Gatun
Creek
Middle Creek Ringicula dominicana Maury Middle Creek, Red Cliff Creek and Hstrella River
197 Costa Rica M1ocENE—OLSSON 25 GENERAL, CORRELATION CHART NO. I. Costa Rica and Canal Santo Jama | Porio wesiern Panama Zone Domingo lca | Rico ee ee ea ae a ee ee Se a
v °
a Limon. Monkey Las Matas
S Point, Bocas Stage
ee Conglomerates | |
(boulder clays) | x *¥ & F & f /% Eh Mase ae) eae NA 64 Lal Cerros de A ? Toro Sal Stage | - Gatun Stage Stage V) | Quebra- \Bowden| dillas Stage | 1's,
Lil Mao Clay | |
Z Gatun | |
mA Stage |Mao Adeotro |
yo & Gatun Stage limestone
ere Gurabo Stage
|
A PUNE Ores eee ane OU eae gone a So a a *
Fa Cercado Stage = Uscari Stage ea fe) (Sapote ) o Baiton Stage iH * cn oan NE 3 Hes aera, (Get dae Feat 13 cae | a) * ‘
Y | |
O ‘Emperado l’s |
5 Undifferen- | Culebra Stage) Cevicos, 1's |
08 tiated Bohio cong. | Tabera Stage| |
\o)
26
BULLETIN 39
198
GENERAL CORRELATION CHART NO. II.
Colombia and Venezuela
RAC Plea PET sas nels wil Rae?
Cartagena and
Cumana % Dec Nase x x c x *
Contemporaneous
Mexico “é % Bs * Me “* xX * x * * &
Txuetepec, State of Oaxaca
Santa Rosa and Barranca de Santa
Maria Tatetla, State of Vera Cruz
*
Rio Coatzacoalcos, \Isthmus of Tehu-
antepec
Chesapeake Series st Lower Upper xx
Series _and Stage
Alum Bluff
*
Eastern United States
Caloosahatchie Waccamaw
Yorktown and Duplin Stage
Murfreesboro Stage
St. Mary’s Stage Choptank Stage
Calvert Stage
cP a DH Russa Gh i | age et
Shoal River marls Oak Grove marls Chipola marls
sk *k wi
Tampa Stage Vickburgs Stage
199 ~ Costa Rico MrocENE--OLSSON Dy)
The Arca Henckent, Glycymeris jamatcensis and Protocardia gurabica occur with a typical Gatun fauna at Water Cay, west- ern Panama, associated with Sconsza, and other Gurabo shells. With the exception of Aingicula dominicana,the other species are generally rare. The evidence is therfore strongly in favor of the equivalence of the Gatun beds with the Gurabo of Santo Domingo.
BOWDEN BEDS, JAMAICA
Next to Santo Domingo, the highly fossiliferous beds at Bowden, Jamaica, are the best known. From this locality, Dall has listed about 435 species, of which 12 per cent. appear to be identical with recent species.
Recent correlation by Cooke places the Bowden beds as equivalent to the Gurabo formation of Santo Domingo,and there- fore to the Gatun formation; a correlation which is borne out by our own studies. About 38 Gatun species, not including recent forms, are here considered equivalent to Bowden shells. In gen- eral, the Costa Rican localities which show the closest agree- ment with Bowden, are the upper coralline horizons, so that the Bowden beds may be a little higher than the main Gurabo and they are so considered here. The following Bowden shells oc- cur in the Costa Rican Gatun.
* Acteocina bullata ‘Kiener
*Acteocina recta d’Orb FPetaloconchus sculpturatus Wea *Volvula cylindrica Gabb Alaba turrita Guppy Bullaria paupercula Sowerby *Crepidula plana Say Terebra bipartita Sowerby *Nrtica near canrena Linne Conus planiliratus Sowerby Glycymeris jamaicensis Dall Conus stenostomus Sowerby *Arca occidentalis Philippi Conus granozonatus Guppy * Arca auriculata Lamarck Conus recognitus Guppy *Ostera megodon Hanley Conus interstinctus Guppy Limca solida Dall
Conus gracilissimus Guppy Placunanomia lithobleta Dall *Turris albida Perry Echinochama antiguata Dall
*Recent
28 BULLETIN | 20 200 Drillia consors Sowerby Venericardia scabricostata Guppy Drillia venusta Sowerby Myrte@a limoniana Dall Cancellaria Barrettt Guppy Phacoides actinus Dall
Oliva cylindrica Sowerby *Levicardium serratum inne *Latirus infundibulum Gmelin Cytherea Blandiana Guppy iilelongena consors Sowerby Tellina lepidota Dall
Metula cancellata Gabb *Strigilla pisiformis Linne Typhts alatus Sowerby Abra triangulata Dall
Distortrix simillima Sowerby Corbula viminea Guppy
* Bursa crassa Dillwyn Gastrochena rotunda Dali
Cassis sulcifera Sowerby Phalium moniliferum Guppy Sconsia sublevigata Guppy Malea camura Guppy
Strombus pugiloides Guppy Serpulorbis papulosa Guppy
MEXICAN MIOCENE
From Mexico, Bose} has described several small faunas from the Isthmus of Tehuantepec, and from the States of Oaxaca and Vera Cruz. These faunas, he considered as Upper Miocene and Pliocene.
The most interesting of these localities, in the present con- nection, are those at Tuxtepec in the State of Oaxaca, and Santa Rosa and the Barranca de Santa Maria Tatetla in the State of Vera Cruz. From these localities, we may note together with their Costa Rican and other affinities, the following species:
Fecten santarosanus Bose FPecten levicostatus 'Toula Gatun Amusium Mortont Ravenel Amusium luna Brown and Pilsbry Levicardium sublineatum Conrad Gatun Chesapeake Miocene Anomia simplex d’Orb Gatun-Chesapeake to recent Venus Ebergenvit Bose Chione ulocyma Dall Upper Chesapeake Solarium Villarellot Bose Architectonica granulata Wamarck Lower Miocene or recent Vermetus pulcher Bose Serpulorbis sculpturatus AH. C. Lea Gatun and Upper Chesapeake Miocene Strombus pugilis Linne Strombus pugtloidis Guppy Gatun Sconsia sublevigata Guppy Bowden fhos mexicanus Bose Gatun of Water Cay Melongena Mengeana Dall Solenosteira Vaughani Dall variety medioamericana Olsson,Gatun of Water Cay * (Recent,
1 Boletin del Instituto Geologico de Mexico, No. 22, 1919, pp. 11 - 88
a Cae aera
20x Costa Rica MIocENE —OLSSON 209
fleurotoma alesidota Dall var.
MEGA BOSE 02.2.0. Drillia consors Sowerby, Gatun Conus Agassizi Dall variety multilivatus BOSC........... Conus multiliratus Bose, Gatun OHMS “IBUPCRRAVTL BOSC...... ccccseecse. Gatun
The above species are from Tuxtepec and Santa Rosa. To these we may add, from Barranca de Santa Maria Tatetla, the following forms:
Ostrea ‘sculpturata Conrad............ Ostvea costaricensis Olsson, Gatun Lucina guadrisulcata Linné........... Divaricella quadrisulcata d’Orb Gatun, Upper Chesapeake Miocene to recent Levicardium serratum WLinné........ Gatun, Upper Chesapeake to recent Dostuia acetabulum Conrad......../... Gatun -Chesapeake Miocene
The above list is but a partial one of the entire faiina from these localities. However, it shows the close relationship be- tween these Mexican species and those of the Gatun and the Bowden, such as Sconsia sublevigata, Phos mexicanus, Conus multiltratus and others, witha few typical Chesapeake species such as Chione ulocyma Dall, Dosinia acetabulum Conrad and Cardium sublineatum Conrad. This illustrates the inte arming- ling of the Gatun-Bowden-Gurabo fauna with that of the Chesa- peake, and proves their equivalence in age.
THe Hast Coast MIocENE
A fine development of Miocene rocks is found along the Atlantic coastal plain and they extend more or less continuously from New Jersey south into Florida. They are continued around the borders of the Gulf of Mexico into Texas, but in thes gions, their marine phases are deeply covered by younger a, ma- tions, and they are known only from deep wells. These Mic- cene beds are divisible into two series, which early were classifi- ed as (a) the older or sub-tropical Miocene and (b) the younger and cold-water Miocene. These terms are descriptive of their main faunal differences. The upper division consisting of sever-
30 BULLETIN 39 202
al formations or stages is frequently known as the Chesapeake Miocene or Chesapeake series, and the lower consisting solely of the Alum Bluff formation, may be designated for sake of discus- sion, the Alum Bluff series. These Miocene formations or stages are the following:
| Yorktown, Duplin Stages |Murfreesboro Stage .
|
Upper
Aik * * * | Chesapeake Series|* * + i A, | | St. Mary’s Stage cage Nu GleKelke ye Choptank Stage 0 | Calvert Stage eS ee a ee ee eee ee to = Alum Shoal River marl | Bluff | Alum Bluff Series} Stage Oak Grove marl Lower | Chipola marl
|
This two-fold division isa very natural one. Each series represents a distinct trangression of the Miocene sea over parts of the Atlantic coastal plain, that of the Chesapeake being the more extensive. In Florida, where the Chesapeake Miocene is sometimes found resting upon the Alum Bluff beds, as at Alum and Jackson Blufis, the contact is seen to be an erosional uncon- formity. At these localities, the Chesapeake Miocene is equiva- lent to the Murfreesboro stage or formation of Virginia and North Carolina, and hence the time interval represented by this break is equivalent to the Lower Chesapeake or the Marylandian Miocene. Dall,* who has paid special attention to the relations of the Alum Bluff and the Chesapeake series, both stratigraphi- cally, and paleontologically, characterizes this break as ‘‘the most sudden, emphatic, distinct in the whole post-Cretaceous his-
* Trans, Wagner Free Inst., 1903, p. 1594
203 Costa RicA M1ocENE—OLSSON aT
tory of our southeastern Tertiary, and indicates physical changes in the surrounding region, if not in Florida itself, sufficient to alter the course of ocean currents and wholly change the temper- ature of the waters of our southern coasts.’’
The fauna of the Lower Miocene or Aluin Bluff series is of warm-water character and largely composed of Antillean types. In the Chesapeake series, the fauna is of cold or temperate-wat- er character and it finds its relations with the present Atlantic fauna north of Cape Hatteras. These faunal differences must be take into account in any correlation between the Miocene of the United States with that of the West Indies and Central America.
In the Correlation Charts,I have placed the Gatun formation as equivalent to the Chesapeake series, the Uscari formation with the Alum Bluff. The evidence upon which this correla- tion is based, may be summarized as follows:
1. The two-fold division of the Costa Rican and Panama Miocene, by an unconformity at the base of the Gatun forma- tion,
2. Wherever the Gatun beds have been found overlain by more recent formations, the age of these overlying beds is Plio- cene or younger.
3. The high percentage of recent species in the Gatun fauna, which agrees with the average as shown for the Chea- peake Miocene.
4. The Gatun fauna, contains a few species which appear identical with strictly Chesapeake species, such as the follow- ing:
Dosonta acetabulum Conrad
Petaloconchus sculpturatus H.C. Lea
Chama congregata Conrad
Plicatula marginata Say?
a2 BULLETIN 39 , 204
The Dosinia acetabulum Conrad is one of the most abundant and characteristic species of the Chesapeake Miocene and in its course through the Chesapeake formations, several varieties are developed, of great use in correlation. "The typical acetabulum is however confined to the Upper Chesapeake formations and it was figured by Conradin his Medial Tertiary, from a fine large example from Smithfield, Virginia. It is characterized by its large size, broad outlines and even, concentric banding. This Upper Chesapeake or typical acetabulum corresponds very close- ly in size, form and sculpture to the Costa Rican examples. It is very common at certain localities in Costa Rica, especially in the lower part of the Gatun formation. Bose records it from Mexico. The Fetaloconchus sculpturatus is a widely distributed species in the West Indies and Caribbean Miocene, from which it is more generally recorded as P. dominigensis, a name given to the Santo Domingian shell by Sowerby. Close comparative study of a large collection of typical sculptuvatus from Virginia and North Carolina, failed to show any real difference, either in details or sculpture or in nature of coiling. The Petaloconchus sculpturatus occurs in the Bowden beds of Jamaica; the Gurabo formation of Santo Domingo; the Quebradillas limestone of Porto Rico; the Springvale beds of Trinidad; and the Gatun of Panama and Costa Rica. It is also known from Cumana, V ene- zuela: Cartagena, Colombia; and Santa Rosa, Mexico. In the Chesapeake Miocene, it is restricted to the Murfreesboro, York- town and Duplin formations, that is, to the Upper Chesapeake. It does not occur in the overlying Pliocene so far as is known.
5. Possibly more significant because of their greater num- ber are the following species, which appear for the first time in the Atlantic coast Miocene in the Chesapeake formations. They also continue into the Pliocene and generally into the recent fauna.
205 Costa Rica M1ocENE—OLSSON 23
Anomia simplex d’Orb St. Mary’s to recent Cardium spinosum var. Tur- font Dall Murfreesboro to recent
Cardium serratum Linné Murfreesboro to recent Cardium medium Linné St. Mary’s to recent Labiosa lineata Say Duplin to recent Tellina umobra Dall Duplin to recent
Chione grus Holmes Duplin to recent Phacotdes radians Conrad Duplin to recent
Phacotdes trisulcatus Conrad | Chesapeake to recent Divaricella gquadrisulcata a’ Orb Murfreesboro to recent Oliva sayana var.immortua Pilsbry and Brown typical shell, upper Chesapeake to recent Fasctolaria tulipa Linné Pliocene
THE Costa RICAN GEOSYNCLINE
The wide distribution of Miocene beds around the borders of the Gulf of Mexico and Caribbean sea, indicates a condition of general deposition during Miocene time. Costa Rica was largely under water, at least during the Lower Miocene, and was the locus of a strait or geosyncline that then connected the Atlantie and Pacific Oceans; while Panama, parts of Nicaragua, and other ceutral American areas were mainly above sea level. This condition is shown in the general igneous character of the backbone of Panama, while in Costa Rica, sedimentary rocks of Miocene age occur even in the high interior and at elevations of several thousands of feet. This geosyncline covered the greater part of Costa Rica, a portion of western Panama, and western Nicaragua. It should be noted that this area, at the present time, is the locus of a large number of active volcanoes.
Certain changes of importance took place at the close of Uscari time, resulting in a brief withdrawal of the Miocene sea. The effects of these changes were widespread. These great land movements, which probably reached a culmination in the Plio-
34 BULLETIN 39 206
cene, had far reaching consequences and to them we must ascribe even the sudden and marked change between the fauna of the Alum Bluff and Chesapeake series of the eastern United States. It probably resulted in a partial closing of the Atlantic-Pacific straits and a disarrangement of the direction of oceanic currents.
‘he strongly clastic character of the Gatun formation as compar- ed with the more shaly Uscari, is also a result of the elevation of the land.
The Pliocene was a period of high elevation and, in Costa Rica, of intense volcanic activity. The coarse conglomerates, or the so-called boulder clays, which have perplexed most geo- logists who have visited Costa Rica, were formed from the erosion of these Pliocene lava flows and deposited as flood-plain material. In afew localities where these conglomerates reached the sea, they contain intercalated shale beds carrying marine fossils of late Pliocene age.
OS CN
CLASS GASTROPODA ORDER OPISTHOBRANCHIATA
SUBORDER TECTIBRANCHIATA
Genus ACTAEON, Montfort
Actzon costaricensis, n. sp.
Shell moderately large, ovate-cylindric; body-whorl large and more than 34 of the total length of the shell; spire pointed, conic; whorls about 6, convex and with deep sutures; spiral sculpture of fine, nearly regular, smooth threads separated by deep and somewhat wider interspaces; there are about 9 spiral threads on the spire-whorls and about 34 on the last whorl; the interspaces are crossed by fine, elevated, distant threads; aper- ture linear-lanceolate, rounded below; columella with a single large fold.
Height 19.5 mm, diameter 10 mm, aperture 14. 5 mm.
This isan unusually large and cylindrical species with a relatively high spire. Suggestion of the original coloration is indicated by a white band which encircles the middle of the last whorl commencing at the upper end of the aperture,— the general ground-color of the surface being a faint purple or red- dish brown. ‘The interspaces between the spirals are crossed by fine, elevated threads, but they appear merely punctate on the eatly whorls.
Gatun Stage: Hill No. 3, Banana River.
Middle Creek.
Genus ACTEOCINA, Gray
Acteocina bullata Kiener, var. costaricana, n. var. Plate 15 figures 20, 21 et: Jornatina bullata Kiener, Icon. Coq. Viv., p. 5, pl. I, fig..4.
206a 34a
34b BULLETIN 39 206b
Shell rather large, moderately solid; outlines subcylindrical but a little wider about the anterior half of the shell; the spire is short, composed of 4 or more whorls, and a small nucleus of the usual type, asin canaliculata,; sutures channelled; the sur- face appears smooth, until slightly magnified when it is seen to be covered with very fine spiral lines; columella carries a single strong fold.
Height 10, diameter 2)vapert ure o) mim:
The Costa Rican fossils are related to the recent A. bullata Kiener, of the West Indies but differ in their shorter spire and are somewhat more expanded about their anterior half. It is the largest species in the Costa Rican Miocene, often reaching a length of ro mm. Its surface is covered with fine, spiral lines and the columella carries a single, strong plication.
Gatun Stage: Middle Creek.
Coll..g, East Grape Point Creck:
Acteocina recta d’Orbigny
Bulla recta d’Orbigny, 1845, Delas Sagra, Hist. Pol. y Nat. Isla de Cuba, vol. 5, p. 67, no. 55; Atlas 8, pl. 4, 072s, figs, 16-20:
Tornatina recta Gabb, Trans. Phil. Soc., vol. 15, p. 246.
Lornatina coix-lacryma Guppy, 1876, Quart. Journ. Geol. Soc. Lon- don, vol. 22, p. 518. Probably not of Guppy, 1867.
Acteocina recta Maury, 1917, Bull. Amer. Pal., vol. 5, pi/17S,; pias,
fie 2. A small species, typically with a short exserted spire and broadly cylindrical form. ‘The Costa Rican examples are all less than 3mm in height. There is considerable variation in the height of the spire, which may be sunken in some cases as illustrated by the larger coix-lacryma Guppy from the Miocene of Venezuela and Martinique, to others with high spires and strongly descending whorls as in the recent canaliculata Say. However they all appear to belong to the same species. From the young of dudla/a var. costaricana, they are at once distinguish- ed by their smooth, plain surface.
Gatun Stage: Coll. 7, Hstrelia iver, Coll, 4, Red Cig Creek.
206c Costa RicA MIocENE—OLSSON 34c
Subgenus CYLICHNELLA, Gabb Acteocina triticum-tritonis Maury
Cylichnella bidtentata Gabb, 1873, (in part) Trans. Amer. Phil. Soc., Vol. 15, ps'273. | Not of d’Orbigny, 1845.
Cylichna bidentata Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. BZD. 519.
Acteocina (Cylichnella) triticum-tritonis Maury, 1917, Bull. Amer. Pale vole5, p: 178, pl. 3, fig. 4.
This shell was separated by Dr. Maury from the recent J7- dentata d’Orbigny, with which Gabb, Guppy and others had identified it. In ¢vztécum-tritonts the anterior of its plications is very small, the posterior one large, which is the reverse of the conditions as seen in the living d¢dentata. ‘The base of the shell is finely grooved with incised, spiral lines. The specimens from Costa Rica agree very closely with typical specimens from the Miocene of Santo Domingo.
Gatun Stage: Middle Creek.
Genus VOLVULA, A. Adams
Voivula cylindrica Gabb
Volvula cylindrica Gabb, 1873, Trans. Amer. Phil. Soc.. vol. 15, p. 246.
Vilvute cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London, WOl. 32, p..518.
cf. Bulla (Volvula) cf. oxytata Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt, Wien, vol. 58. p. 709, pl, 28, fig. 4.
Volvula cylindrica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 180, pl. 3, fig. 5.
This, is a small species of no unusual characters. It is nar- rowly cylindrical in form, shortly produced and pointed at its posterior extremity. It was described by Gabb from the Mio- cene of Santo Domingo, but Dall united with it, the oxytata Bush, in which case, it is also found in the recent fauna. Gabb records acuta d’Orb. from the Pliocene of the Limon Peninsula.
Uscart Stage: Middle Creek.
Genus BULLARIA, Rafinesque
Bullaria paupercula Sowerby Plate 4, figure 14
34d BULLETIN 39 : 206d
Bulla paupercula Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. '6,°p. 52. : Bulla paupercula Gabb, 1873, Tfans. Amer. Phil. Soc., vol. 15, p. 246. : x
Bulla paupercula Guppy, 1874, Geol. Mag. London, p. 437.
Bulla paupercula Guppy, 1876, Journ. Geol. Soc. London, vol. 32, p. 518.
Bulla paupercula Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3,
sO Oe hay On oe Bulla striata Dall, 1903, Trans. Wagner Free Inst., Sci., vol. 3, pt. 6,
S583. Bulle Re penae Maury, 1917, Bull. Amer, Pal., vol. 5, p. 182, pl. 3. fre. WS: Bullaria paupercula Maury, 1920, N. Y. Academy of Science, Tertiary Mollusca ‘of :/Porto.Rico!) vols 3, pt Lape. 75:
This species is rare in Costa Rica, and all our specimens are from the Lower Gatun rocks of Middle Creek. It is possibly identical with the recent Aullaria amygdala Dillwyn, abundant on the beaches along the northern coast of Panama ‘and Costa Rica.
Gatun Stage: Middle Creek.
Genus RINGICULA, Deshayes Ringicula dominicana Maury Ringicula semtstriata ? Gabb, 1875, Trans. Amer. Phil. Soc., vol. 15, p. 225. Not &. semistriata d’Orbigny. kRingicula dominicana Maury, 1917, Bull. Amer. Pal., vol. 5, p. 185, jn ae ui WEA OE
This is the most abundant of the Opisthobranch mollusks in the Miocene beds of Costa Rica. The whorls are in general more strongly sculptured with incised lines than typical domznicana from the Miocene of Santo Domingo, and the spiral lines extent over the greater part of the shell,except for a wide, smooth band about the suture.
The R. hypograpta Brown and Pilsbry,from Gatun is a larg- er shell with a heavier outer lip and more crowded spiral lines about the base. ‘The Costa Rican shells measure about 1.5 mm or less in height.
Gatun Stage, Middle Greck, Coll. 5, Red Clip Oya
Cowl. 7: fEsiveila \Rtver,
207 Costa Rico M1ocENE—OLSSON 35
ORDER CTENOBRANCHIATA CA SWPER-PAMILEY | TOXOGLOSSA
Genus TEREBRA Apanson
Terebra subsulcifera Brown and Pilsbry
Terebra subsulcifera Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Piilay vol 63. p.°339, pl. 22, figy: 7:
Terebra (Myurella) subsulcifera Cossmann, 1913, Jour. de Conchylio- Ioee VOL! 61, pl) T4, ‘pl. 1, He.) 25.
The single, fragmentary specimen comes from the Gatun beds of Mt. Hope in the Canal Zone. From dzfartita, which it resembles in its sculpture, this species and haztenszs Dall, differ in having only a single columellar fold.
Gatun Stage: Mt. Hope, Canal Zone.
Terebra cf. haitensis Dall Plate 1, figure 3
Terebra haitensis Dall, 1895, Proc. U. S. Nat. Mus., vol. 18, p. 35. Terebra haitensis Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt.
GD. "lO32.\ pl. 59, fig. 30. Terebra hattensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 194. pl. 4,
fio), 2,
The collection contains a single, imperfect shell from Sousci Creek, which should probably be referred to this species. From the subsulczfera, it differs in its more numerous and oblique ribs, and wider, less pronounced spiral bands. Like subsulcifera, the columella carries a single, sharp plication. Our specimen con- sisting of only 5 whorls measures 37 by 13 mm.
Gatun Stage: Sousct Creek, a branch of Hone Creek, C. R. Terebra bipartita Sowerby Plate 1, figure I, 2.
Terebra bipartita Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 5» P- 47.
36 BULLETIN 39 208
Terebra bipartita Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
225.
Terebra (Acus) bipartita Dall, 1895, Proc. U. S. Nat. Mus., vol. 18, p.#3e.
Terebra bipartita Maury, 1917, Bull. Amer. Pal., vol.5, p. 187, pl. 3 fig. 14.
This species, possessing much the same type of sculpture as sultifera, subsulcifera, haztensts etc., is recognized by having two, instead of one columellar plication. Full-grown specimens have a bipartite sculpture while in salcifera and the others it is tripartite.
The Costa Rican collection contains three specimens from Saury creek, near Cahuita, the largest of Io incomplete whorls mearures 77 by 19 mm.
Gatun Stage: Zone FE. Saury Creck
Terebra gatunensis Toula | Plate 1, figures 4-6
Terebra gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt, VOLU58, p.2705 "pl. 25; fie.) 12,
Terebra gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Bint vol: 63; "p: B39 ple 22) hie) 2!
Terebra wolfgangit Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phil., vol. 63, p. 340, pl. 22, figs. 1, 3-6, but not of Toula, 1908.
Terebra (Myurella) gatunensis Cossman,1913,Jour. de Conchyliologie, vol. 61, p. 13, pl. 1, figs. 26-29.
In the Canal Zone, the Gatun beds contain two common species of Zevebva, which were described by Toula as gatun- enstsand wolfgangi. ‘These species have been more or less con- fused by Brown and Pilsbry, so that their figures and description of wolfgangi, really represent gatunensis. Their figure 2 isa large and less common variety of gafunensis. 'Toula’s figure, al- though of a young imperfect shell, is clear and shows the main
209 Costa RicA MIocENE—OLSSON Cul
characters of the species, the spirally striated, faint sutural sul- cus and fine spiral scu]pturing of the whorls. He however in his notes, allies this species with the recent 7. dzslocata of Say, which is misleading. Toula’s figure can be duplicated by scores of specimens in our collection.
Terebra gatunensts is characterized by its large size, (length 50 mm or more), its faint sutural sulcus which may be smooth or with 2 or 3 faint spirals, showing best on young shells. The spiral cords of the main body of the whorl are variable in num- ber, spacing and strength, as well as the longitudinal ribs. A large coarse variety, with persistent sulcus, and heavier sculp- ture occurs rarely and is shown by Brown and Pilsbry as their figure 2. The pillar is nearly smooth, carrying a very broad fold above, and a small one below.
Gatun Stage: Gatun and Mit. Hope, C. Z. Terebra wolfgangi Toula Plate 1, figure 11, 12
Terebra wolfeangi Toula 19090, Jahrb. der K-K Geol. Reichsandstalt Jd < ) WOle/ 58) -p. 705, pl. 28; ig. 7-
Not 7erebra wolfgangi Brown and Pils. whichis 7. gatunensis Toula.
Although 7. gatunensis and wolfgang? have not been collect- ed by us except in the Canal Zone, where they are abundant, they are included here for the sake of completeness. It is also not unlikely that future collecting may still reveal their presence in the synchronous beds in Costa Rica and adjacent parts of Panama.
T. wolfgang? differs from the common 7. gatunenszs by its long, tapering form, deep, persistent sulcus anda sutural band generally free from spirals. It is also a much smaller species, the largest specimen in our collection measuring about 38 mm in length.
Gatun Stage; Mt. Hope, C. Z.
38 ? BULLETIN 39 210
Terebra gausapata Brown and Pilsbry Plate 1, figures 8, Io
Terebra gausapata Brown and Pilsbry, April 1911, Proc. Acad. Nat. Sci. Phila.) wol. 63. "p.3408 pl. 22, fos Sno:
Terebra acuaria Toula, Dec. 1911, Jahnb. der K-K Geol. Reichanstalt, VOL 62) pi) LO, spl. aeihig a Los
Terebvra (Myurella) acuaria Cossm., 1913, Journ. de Conchyliologie, VOl. 61 pe12,)\ pl Les. 19-242
The 7. gausapata, was described by Brown and Pilsbry, from young specimens, measuring only I9 mm in length. Later in the same year appeared Toula’s 7. acuaria, also based on young shells.
The important characters of this species, are its nearly straight ribs, deep sulcus, a sutural band with faint spirals and even sculpture. Figures 23 and 24 of Cossmann I believe represents wolfgangi. T. wolfgangt as will be noted from the accompanying figures is a much more slender species and usually with a smooth sutural band. ‘The pillar carries two broad folds, which are better developed than in wolfgang? or gatunensts.
Length 33, breadth 8.5 mm.
Gatun Stage ML) Tope, (Co. Z. Water Cay.
Terebra spirifera Dall Plate 1, figure 13 Terebra dislocata Gabb, 1873, Trans. Amer. Phil. Sci., vol. 15, p. 225, Not of Say.
Terebra (Acus) bipartita Sowerby, variety spirifera Dall, 1895, Proc. U. S. Nat. Mus., vol. 18, p. 38.
Terebra (Oxymeris) bipartita Sowerby, variety spirifera Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1632, pl. 59, fig. 13.
Terebra spirifera Maury, 1917, Bull, Amer. Pal., vol. 5, p. 188, pl. 3 igs. 415, )10:
In 7. Gatunensis and wolfgangi the pillar is nearly smooth
21r- Costa R1icA MIocENE—OLSSON 39
or with two board, sub-obsolete folds. In sfzri/era, the two folds are strong and recall those seen in the recent 7. aislocata. ‘The sutural band is strongly sculptured by the end of the ribs which are entirely dislocated by the deep sulcus. Typical specimens have three or four, even spirals, which are stronger than the longitudinal ribs. Variation occurs in the number, and regu- larity or these spirals,
Length 43, breadth 9 mm.
Gatun Stage: Banana River, Hill No. 1. Water Cay.
Terebra spirifera Dall, variety midiensis, n. var. Plate 1 fieure 7
This variety, represented only by a single specimen from Middle Creek, C. R., differs from typical sfzrzfera, in having on- ly three spirals, which are broad and strap-like in form.
Gatun Stage: Middle Creek. C. R.
Terebra Berlinerae Maury Plate 1, figures 17, 18
Tevebra Berlinere Maury, 1917, Bull. Amer. Pal., vol. 5, p. 198, pl. Ay fess 7. 3.
This beautiful species was described by Dr. Maury, from the Cercado (Miocene) formation of Santo Domingo. ‘The whorls are slightly convex, and its sculpture is closer, finer and more delicate than is seen in any of the associated Costa Rican Tere- bras. Our specimens, all imperfect, were collected from the hard Gatun sandstones of Boucary Creek. The most complete specimen measures 21 by 6.5 mm.
Gatun Stage: Boucary Creek.
4O BULLETIN 39 ZizZ
Terebra estreliana, n. sp. Plate 1, figures 9, 16
Shell small; nucleus of three smooth tapering whorls; post- nuclear whorls about 9, sculptured with a faint sutural sulcus, which is lacking or nearly so from the early whorls; axial sculp- ture of about eleven, straight or slightly oblique ribs which pass from suture to suture; but are nearly lacking from the base; spirals lacking or very faint on the spire-whorls, so that they ap- pear smooth and solid; the base of the last whorl is strongly con- tracted and shows four or five, very faint spirals; pillar straight and smooth; beak twisted.
Length 7.5, diameter 2.5 mm. This is a small species of rather solid appearance, strong ribs, but otherwise nearly smooth whorls. ‘The spirals are very
faint and are seen best only when viewed at right angles to the source of light. Our specimen came from the Estrella River.
Gatun Stage: Zone 7, Fistrella River, Gia.
Terebra costaricensis, n. sp. Plate 1, figure 19
Shell small and slender, with about three, long, tapering smooth, nuclear whorls; post-nuclear whorls eleven; sutural band rather large and prominent, strongly tuberculated; the spire-whorls are slightly concave and have at their base or just above the suture, a revolving series of tubercles, which are the enlarged bases of the longitudinal ribs; longitudinal ribs on the last whorl nuinber about twelve, are nearly straight and large and strong on each side of the suture, low and weak in the mid- dle of the whorls; the surface of the whorls are otherwise smooth or in some shells feebly spiralled; base slightly rounded with eight or more spirals which become finer as they approach the canal; the longitudinal ribs are continued across these spirals.
Length 13, diameter 3 mm.
The above description is based mainly on two specimens |
213 Costa RicA MIocENE—OLSSON AY
from Hill No. 3, of the Banana River. Our collection contains several smaller shells from Zone 7, of the Estrella River in which the base is somewhat more heavily sculptured and with very fine spirals over the main surface of the whorls.
Terebra benthalis Dall var. bocasensis, n. var. Plate 1, figures 25, 26
Cf. Zerebra (Acus) benthalis Dall, 1889, Bull. Mus. Comp. Zool., vol. ¥5,"p./05; pl. 29, fg. 6:
This species, fairly abundant in the shale beds, just north of the city of Bocas del Toro, Panama, agrees closely with Dall’s figure in the Blake Report, except that the revolving series of tuberclesin the fossil shells seem to be lower and broader. The whole surface on slight magnification is seen to be finely spirally striated.
Terebra benthalis was described by Dall from specimens dredged from Ico to 4oo fathoms of water off the Morro Light, Havana, Cuba. The species belongs to the section Fusoflerebra Sacco.
Dengthi 27) diameter) 6.5) mim.
Gatun Stage: Bocas del Toro, Panama. Terebra pumbriensis, n, sp. Plate 1, figure 27
Shell of medium size, composed of 9 whorls (type with a sinall portion of the apex lost); the early post-nuclear whorls (the rst 3 or 4) with two revolving rows of tubercles, formed by the intersection of fine, oblique riblets and two spiral bands; on the 4th whorl of our specimen, a median spiral makes its ap- pearance and rapidly increases in strength so that the later whorls have three rows of tubercles, the upper or most posterior one being somewhat the strongest; the longitudinal ribs are nar- row and oblique, and on the last whorl number about 20; the space between the ribs have three or four fine, incised growth- lines; periphery of the base has an additional tuberculated spiral,
42 BULLETIN 39 214
below which is a band composed of four smooth, close spirals; be- tween these spirals and the keel is a deep, concave zone; pillar nearly straight and smooth, with a single sharp keel on the back.
Length 20, diameter 4.75 mm.
Of this interesting and distinctive species, we have only a single, imperfect specimen from the Gatun beds of Pumbri Creek. The three rows of tubercles are evenly spaced and there is no sutural band.
Gatun Stage: Pumbri Creek, C. R. Genus CONUS Linné Conus molis Brown and Pilsbry Plate 2, figures 1, 2
Conus molis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila.,
VOL O27 LGAs, ipl. 230 uot oh. Conus molis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200.
This is the common Cone in the Gatun beds of Panama and Costa Rica and large specimens may reach a length of nearly 150 mm. The upper surface of the whorls carry 5 or more fine spirals and perfect examples of young shells have a few of the early post-nuclear whorls finely coronated. Described by Brown and Pilsbry from the Canal Zone, the species has also been col- lected by us at several stations along the Panama coast between Colon and Bocas, at Water Cay, and along the Banana River in Costa Rica. A large specimen from Water Cay measures: length 144 mm, width 81 mm.
Gatun Stage: Gatun, Water Cay, Banana River, ete.
Conus stenostomus Sowerby __ Plate 3, figure 2
Conus stenostoma Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 5 p. 44.
Conus stenostoma Guppy, 1866, /d., vol. 22, p. 287, pl. 16, fig. 2.
Conus stenostoma Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 230.
21s ! Costa Rico MIocCENE—OLSSON 43
Conus catenatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32, p.527; 1m part b
Conus stenostomus Dall, 1903, Trans, Wagner Free Inst. Sci., vol. 3, Be 6." p: 1583.
Conus stenostomus Maury, 1917, Bull. Amer. Pal., vol. 5. p. 203, pl. 6, fies A.
A. species characterized by its sharp angulated shoulder, concave upper surface of its whorls and attenuated anterior can- al. Well-preserved examples showing a highly polished sur- face and traces of yellow biotches of color markings were col- lected quite abundantly in the coralline of Port Limon. The species also occurs in Jamaica and Santo Domingo.
Gatun Stage: Port Limon.
Conus concavitectum Brown and Pilsbry
Conus concavitectum Brown and Pilsbry, 1911. Proc. Acad. Nat. Sci. Elia. vol. 63, p- 341, pl. 23, figs.:5,,6.
Conus (Lichoconus) concavitectum Cossmann, 1813, Jour. de Conchy- hologie, vol. 61, p43, pl. 4, figs. 3, 4.
In adult shells, the spire whorls become concave. The Species is also recognized by its sharp shoulder and low spire. It may be confused with varieties of proteus but the sharp should- er angle and spiralled spire whorls will separate it. Our speci- mens come from Water Cay, and measure 48 by 27 mm.
Gatun Stage: Canal Zone and Water Cay.
Conus proteus Hwass Plate 2, figures 3, 4
Conus proteus WHwass, 1789, Enc. Meth. vers, I pt. 2, p. 682.
Conus leoninus Wwass 1789, Enc. Meth. vers, 1 pt. 2, p. 683.
Conus proteus ? Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 232.
? Conus Berghausit ? Gabb, dem, p. 232. Not of Hoernes, Foss. Wien. Beckuvol) i) figs-z:
Conus leoninus Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 359.
44. BULLETIN 39 | 216
Conus proteus Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1, iD: 26:
Conus proteus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 206, pl. 6, Liga ie
This isa common species in the Miocene and Pliccene beds of Costa Rica and Panama. It is generally possible to distinguish two forms, a smaller one which agrees with typical examples of proteus in form and coloration (which is frequently preserved, ) and a larger and heavier type, like the so-called /eoninus. ‘This second variety may show several dark spiral bands in addition to the square yellow blotches. Such markings we have not observ- ed on recent examples of leonznus.
Gatun Stage: Banana River, Port Limon, Water Cay. FPlio- cene: Monkey Point; Bocas.
Conus Veatchi, n. sp. Plate 2, figures 5, 8
Shell rather large, heavy and broad, with a low, nearly flat spire, but pointed, slightly projecting apex; spire-whorls about Io in number, witha slightly raised border near the suture and marked with fine spirals which on the last whorl number 10 or 11; shoulder of the last whorl rounded or beveled; last whorl, large and broad, nearly smooth above, except for the very faint spirals, but with numerous, fine, irregular spirals on the canal; pillar straight; aperture straight and narrow, slightly wider below.
Length 43, breadth 28 mm.
The present collection contains two specimens, the larger of which from Water Cay, serves as the holotype. ‘The species is very unlike any of the associated Gatun species in its low, near- ly flat spire. The Conus demiurgus Dall, of the Chipola Mio- cene of Florida, is a related species.
It is named for Dr. A. C. Veatch, Director of the Explora- tion Work of the Sinclair Consolidated Oil Corporation, for his
217 Costa RicA MIocENE —OLSSON A5
interest and co-operation in obtaining the extensive collections which form the basis of the present work. Gatun Stage: Water Cay, Panania.
GOH CoD, Conus floridanus, var. costaricensis, n. var. Plate 3, figures 3, 9
Shell conic, witha moderately high spire of about eleven whorls; profile of the spire flat or slightly concave to the long projecting apex seen in well-preserved specimens; shoulder angle rather sharp with the whorls slightly concave above between the sutures; spire-whorls generally show 3 or more faint, raised spirals and strongly arcuated growth lines; the last whorl below the shoulder angle is usually nearly ‘smooth above (sometimes with faint spirals) but with heavy regular spiral groovings to the number of about fifteen on the canal; the canal is long, rap- idly attenuated and usually somewhat flexed below.
Height 44, breadth 21 mm. Type trom the Gatun beds of the Banana Rive
This fossil, quite common in Costa Rica, should probably be classed as a variety of the recent Conus floridanus Gabb, differing only in always being somewhat more slender. They also agree closely with Dall’s figure of Conus chifolanus Dall from the Chipcla Miocene of Florida.
Care should be taken in distinguishing this from C. emttator of the Canal Zone. In that species, the full-grown shell is larger, with nearly straight sides and finer irregular spirals on the canal.
Gatun Stage: Banana River; East Grape Potnt Creek.
Conus imitator Brown and Pilsbry Plate 2, figure 6
Conus tmitator Brown and Pilsbry, April, rgix, Proc, Acad, Nat. Sei, Phila.) vol. 63) p, 342, pl. 23, figy 4:
Conus DMallti Toula, Dec., igt1, Jahrb. der K-K Geol. Reichsanstalt, Wien, vol. 61, p. 508, pl. 31, figs. 23 a-d.
46 BULLETIN. 39 218
Conus (Lithoconis) Dallt Cossmann, 1913, Jour. de Conchyliologie, VOl..61.9 p.'41) pl.) 3.) figs.) 20) (30, pli Ay hes wa cee ? Conus , Dall Maury, 1917, Bull. Amer. Pal., woliS.p onal, LS; ah TS The Conus Dalli Toula is doubtless the young of zmztator described earlier in the same year by Brown and Pilsbry. Fig- ure 6 is of a specimen from Mt. Hope which measures 51 by 25 min. From Costa Rica, the collection contains a series of per- plexing Cones, which probably should be referred to this speces. From costaricensis they differ in their nearly straight sides, more irregular spiral groovingson the anterior canal, and they usually show faint spirals over most of the body-whorl, which I have not observed on costaricensis. A large shell from Zone C of Comadre Creek near Cahuita, measures 65 by 29 mm. Gatun Stage: Mt. Hope and Gatun C. Z. Water Cay, Panama. Banana River, East Grape Point Creek, Comadre, Mtiddle Creek, C. R.
Conus recognitus Guppy Plate 2, figure 9
Conus solidus Sowerby, 1849, Quart. Jour. Geol. Soc. London, vol. 6, Pp. 45, Not C. solidus Sowerby, 1841.
Conus solidus Guppy. 1866, Quart. Jour. Geol. Soc. London, vol. 22, p.1287. p16, tig. im.
Conus vecognitus Guppy, 1867, Proc. Sci. Assoc. Trinided, p. 171, Har- ris Reprint, Bull. Amer. Pal., vol. 8, p. 198.
Conus pyriformis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 229, Not of Reeve, 1843.
Conus vecognitus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol.
32, p- 527.
Conus recognitus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. Gi ps 1583)
Conus recognitus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 209, pl. 7, fie. 9.
This species is easily recognized by its well-rounded should- er and appressed sutures. It is closely related to the recent C.
Zig. Costa Rica MrocKNE—OLSSON 47
pyriformts Reeve of the west coast of Central America but the fossil shells have generally a lower and flatter spire. It is also known from the Miocene of Santo Domingo, Jamaica and Trinidad.
Gatun Stage: Banana River, Hill No. 3.
Conus limonensis, n. sp. Plate 3, figures 7, 8
Shell rather large, but delicate and rather slender; spire low, concave to the slightly projecting apex; spire whorls Io or more, nearly flat between the sutures or slightly concave due toa slightly raised border; spire-whorls with 3 or 4 low spirals which become faint on the later whorls so that they are nearly smooth; sutures deep; shoulders of the last whorl angled or somewhat raised; body-whorl long and slender, polished and smooth, ex- cept for a few faint spirals near the tip of the canal; anterior canal long and straight, evenly tapering.
Length 41, breadth 24 mm.
This species is based on 2 specimens from the coral lime- stones of Port Limon, where it is associated with Conus sfenost- omus and planiliratus. From stenostomus, its straight canal, low spire and nearly smooth, polished surface is most noticeable.
Gatun Stage: Port Limon, C. &.
Conus musaensis, n. sp. Plate 1, figures 22, 24
Shell small and solid; spire short, conic, composed of about 2 pellucid, globular and projecting whorls and 6 post-nuclear whorls; the spire-whorls are flat or slightly cancave and project slightly over the anterior sutures; body-whorl with the upper half smooth, but with 12 deep grooves on the anterior canal which produce wide, spiral bands: these grooves are delicately incised by raised longitudinal lines; color markings are some- times preserved, which appear as 8 narrow, gray bands on the upper half of the whorl.
Length 19, breadth 9 mm.
This small species, the Gatun analogue of the recent West
48 BULLETIN 39 220
Indian Conus Agasstzi Dall, is abundant along the Banana River and elsewhere in Costa Rica.
Gatun Stage: Banana River, Zone 6 Red Clif Creek, Wer.
Conus cf. interstinctus Guppy Plate 3, figure 12°
Conus inierstinctus Guppy, 1866, Quart. Jour. Geol. Soc. London, vol. 22,.0-P 7 Zoo. pl, MO Vitor ah,
A single specimen from Hill 1, of the Banana River, agrees closely with Guppvy’s figure of this Jamaican species. The shell measures 39 by zomm. The spire is moderate in height, with a slightly concave profile. The spire-whorls are smooth between the sutures but also show a slight coronation. ‘The last whorl carries about 15 narrow, widely spaced, spiral cords which an- teriorly are slightly beaded.
Dr. Maury unites zz¢erstinctus, with Sowerby’s catenatus, but that species is based on a very young and scarcely distin- guishable specimen. We therefore prefer to compare the Costa Rican shell with Guppy’s larger and well-figured zuzterstinctus.
Gatun Stage: Banana River, Hill No. rf. Conus cf. Sewalli Maury
Conus Sewatli Maury, 1917, Bull. Amer. Pal., vol. 5, . 201, pl.-5, fig. 3, pl. 6; fig:
The collection contains 2 young shells which may belong to this Dominican species. ‘They differ from the preceding zz/ter- stinctus in being shorter and broader, and the spire-whorls in place of being smooth are marked with 5 or 6 strong spirals. The last whorl is sculptured with 15 or more widely spaced beaded spirals. The dimensions of the largest shell are: Length or height 30, breadth 19 mm:
Gatun Stage: Banana River. Conus toreensis n. sp. Plate 2, figure 7
Shell of medium size, with a conic spire a little more than % the length of the aperture; the whorls of the spire numbering
7
221 Costa Rica M1rockRNE—OLSSON Ag
12 plu; are flat or slightly concave and bordered below by a low carina projectly slightly above the suture; this carina on all ex- cept the last whorl bears low nodes, about 20 to the later whorls; the spire-whorls are otherwise smooth except for the arcuate growth lines; the last whorl is a tapering cone, nearly smooth, except for the faint spirals about its lower one-third; the aper- ture is narrow, straight, with a thin arcuate outer lip.
Height 45, diameter 21, aperture 33 mm.
This shell belong to the Conus consobrinus group, agreeing in its form and strongly nodulated spire whorls. The main difference is that foroenszs is nearly smooth, the spirals showing only on the lower one-third, while cozsobrinus has most of the last whorl covered with beaded spirls.
Gatun Stage: Toro Cays.
Conus (Chelyconus) tortuosostriatus Toula Plate 1, figure 15
Conus (Chelyconus) tortuosostriatus Youla, 1911, Jahrb. der K-K. Geol. Riechsandstalt, Wien, vol. 61, p. 508, pl. 31, fig. 22.
Conus (Hemiconus ) tortuosostriatus Cossmann, 1913, Jour. de Conchy- ologie; yol, 61, p. 40, pl. 3, figs.. 28, 20:
Conus tortuosostriatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 205, pl. 6; fig. 9
This shell should probably be considered as a short variety of gractltssimus Guppy, differing chiefly in its broader form and shorter spire.
The body-whorl is sculptured with about 24 raised,even, spiral cords, separated by deep, even interspaces. ‘These interspaces are crossed by neat, incised lines, corresponding to the lines of growth. The nucleus consists of about 3, long, tapering smooth whorls, after which follow several turns of carinated and finely coronated post-nuclear whorls.
Bength 22) breadth, 9 mu Gatun Stage: Bocas del Toro.
59 BULLETIN 39 222
Conus granozonatus Guppy Plate’ 3) fig. 15
Conus granozonatus Guppy, 1866, Jour. Geol. Soc. London, vol. 22, p. 287, pliyT6. he. Ss. e
This is a rare species in Costa Rica, our collection containing.
only a single specimen from the Gatun beds of the Rio Blanco. It is characterized by a rather high, conic spire, with smooth or only slightly coronated whorls. The last whorl has about 18, wide, spiral bands, which are heavily granulated above. ‘The specimen agrees well with Guppy’s figure of the Jamaican shell. Length 25, breadth 11 mm. Gatun Stage: Rio Blanco, C. R.
Conus planiliratus Sowerby Plate 3, figures 10, 13
Conus planiliraius Sowerby, 1849, Quart. Jour, Geol. Soc. London, vol. 6, p. 44. .
Conus planiliratus Guppy, 1866, Jdem. vol. 22, p. 287, pl. 16, fig- ure 1/7,
Conus planiliratus Gabb, 1873, Trans. Amer. Phil. Soc., Pp. 230, in part.
Conus planiliratus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. B2 ups 6526.
Conus planilivatus Dall, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, Pyd5os:
Couus planilivatus Cossmann, Jour. de Conchyliologie, vol. 61, p. 48, ples hes 25.126, 27,
Conus regularis Gabb, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 359, pl. 46, figs. 45-48. Not of Sowerby.
Conus planilivatus Maury, Bull. Amer. Pal, vol 5;>p.) 209," pl 7, fig. 10.
The occurrence of this species in Costa Rica is limited to the coral limestones of Limon, Gabb had specimens from here which he identified with the recent West Coast Conus regularis of Sowerby. The Limon shells agree very well with Guppy’s figure in the Quarterly Journal and with Cossmann’s figures of Bowden and Mindi specimens. The species appears to be closely
223, Costa Rico MrocENE—OLSSON 51
related to the recent Conus Stimpsont Dall from the Florida eeraves. (Proc, U. S.: Nat. Mus., vol. 24, p.'503, pl. 20, fig. 7, 1903).
Surface of whorls are heavily sculptured with 25, 30 or more subequal spiral cords. Upper surface of whorls, flat or slightly concave and with 3 or more spirals. The shoulder is sharp and bounds a rather high, conic spire.
Length 31, breadth 16 mm.
Gatun Stage: Port Limon. Conus planiliratus, var. bocasensis, n. var. Plate 3, fig. 14,16
Shell of medium size, solid with a broad, stubby outline; spire low, conic with nearly straight sides and composed of seven or more whorls; the rst 4 or 5 post-nuclear whorls are coronated; spire-whorls with 3 faint spirals and curved growth-lines; body- whorl large and wide, with 25 strong but somewhat irregular simple spiral cords; their interspaces are wide and with crowded, raised lines of growth and sometimes intercalated spirals; anter- ior canal nearly straight.
ener) 32\ breadth 17 mim,
This Cone, which should probably be considered as distinct, is fairly common in the shale beds, belonging to the Gatun Stage on Bocas Island. From typical planzl7ratusit differs in its broad stubby form and coarse sculpturing.
Gatun Stage: Bocas del Toro, Panama. Conus marginatus Sowerby Plate 1, fig. 20
Conus marginatus Sowerby, 1849, Quart. Jour. Geol. Soc. London, VOlG) Dl .4A.
Canus marginatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 16, p. 230.
Conus marginatus Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32, p. 528, pl. 29, fig. 5.
Conus marginatus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 210, pl. 7, fi 2 ET
CR NX
BULLETIN 39 224
On the basis of Guppy’s figure inthe Quarterly Journal, this cone is seen to belong to the type exemplified by the recent C. verrucosus, with its sulcated sides and appressed or beveled
shoulder angle. Our collection contains a single specimen from ~
Zone 5 of Red Cliff Creek which is very close to Guppy’s figure. Cossmann’ S specimen of marginatus from Martinique should probably be referred to some other species.
Gatun Stage: Zone 5, Rea Clif Creek; (Gage Conus gracilissimus Guppy Plate 1, figure 14
Conus gracilissimus Guppy, 1866, Quart, Jour. Geol. Soc. London, VOL. 22) D. 26o, Dl. iG, toh Ae
Conus Orbignyi Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 230, Not C. Orbignyi Audouin, 1830.
Conus g1 acilissimus Guppy, 1876, Quazt. Jour. Geol. Soc. London, vol. 32, P- 527.
Conus gracilissimus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, PLO, i peeirs oc.
Conus gracilissimus Cossmann, 1913, Jour. de Conchyliologie, vol. 61, play, figs rs.
Conus gracilissimus Maury, 1917, Bull. Amer. Pal., vol. 5, p. 204, pl. 6, figs. 8.
A small, but long and slender cone, sulcated with 25 or more, strong, regular spiral cords. ‘The spire'is long and slend- er with slightly coronated whorls and the area between the su- tures is delicately decussated by 3 or more spirals crossed by the raised and arcuated lines of growth.
Our collection contains specimens fi ‘om Bocas and Limon, the largest measuring 26 by 9 mi.
Gatun Stage: Bocas del Toro, Pan., Fort.lLimon, ©. ie
Conus Burckhardti Bose Plate 3, figures 4, 5 Conus Burckhardti Bose, 1906, Bull. de Inst. Geol. de Mexico, numero 22, p. 50, pl. 5, figs. 39, 4o.
Shell of moderate size, with a rather high slightly concave spire and long, acute apex; sutural whorls smooth, except for the
SS po
Cat ie)
R25 Costa RrcA MIocENE —OLSSON
faint arcuate lines of growth and very indistinct spirals; the earlier whorls have a carinated stiture but are only very faintly coronated; last whorl heavily sculptured with about 20, wide, even spiral bands, separated by grooves of about 1-3 of their width; these grooves are faintly incised by the longitudinal lines of growth; the spiral bands are slightly wider above and smooth, becoming granulated on the canal; this granulation appears on the posterior side of the bands only; aperture long and narrow. Length 36, diameter 14.5 mim. 33 (apex broken) diameter 15 mm.
The original specimen of Conus Burckhardti was described from the Miocene of Mexico, State of Oaxaca. Our specimens from Panama and Costa Rica area little larger, but show the characteristic high, smooth, spire and strong,flat, spiral bands of the body-whorl.
Conus tortuosopunctatus Toula is a similiar but smaller species. The difference in sculpture of the two species may be seen in the accompanying figures, the spiral bands of /ortuoso- punctatus being double or incised, while in Burckhardti, they are broad and simple.
Gatun Stage: Water Cay, Panama. Zone 2, Last Grape Point Creek.
Conus Harrisi, n. sp. Plate 3, fig. 1
Shell of moderate size, subcylindric with a high, even, conic spire; spire-whorls about 6 plus, flat, with close sutures, except for the upper whorls which are slightly carinate; the surface of the spire-whorls is nearly smooth, except for a few faint spirals; the last or body whorl is strongly sculptured with widely spaced and sub-regular grooves; these grooves form about 14 wide, spiral bands about the lower 3-4th of the last whorl; the upper 4th below the shoulder angle forms a wide, smooth band; below, the spiral bands are generally smooth except those about the lower
54 BULLETIN 39 226
half of the shell, which may become, mesially divided and bear afew small granulations or beads about their upper half; aper- ture sublinear.
Height 51, diameter 23 mm.
A single specimen of this elegant species was collected from the Gatun of the Banana River. It is a species of the Conus Burckhardti type but much larger and when perfect reaching a length of nearly 60 mm. ‘The sculpture is of wide bands formed by deep, and regularly spaced grooves about the lower 3-4ths of the shell.
Gatun Stage: Hill No. 3, Banana River.
Conus multistriatus Bose Plate 1, figures 21, 23
Conus Agassizi Dall, variey multistriatus Bose, 1906, Bull. de Inst. Geol. de Mexico, numero 22, p. 49, pl. 5, figs. 34-38.
Conus gaza Johnson and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., WOl. 63% pli22), his 103): Conus gaza Maury, 1917, Bull. Amer. Pal., vol.) i5))p.) 20. 7 fo ens This is the Conus gaza of Johnson and Pilsbry described from the Miocene of Santo Domingo and the Isthmus. It isa small, pretty species, recognized by its nearly biconic outlines and neat sculpture. The spire is high; with nearly smooth, spire- whorls and carinated sutures. The last whorl is rather short and heavily sulcated with about 18, regular, spiral cords, separated by intervals of their own width. All our specimens come from Water Cay, where the species is fairly abundant. The largest specimen measures 20 by II nm. Bose’s original specimens are from the Miocene of the State of Oaxaca, Mexico. They are somewhat larger, measuring PRMMON MUCH cab eal Gatun Stage: Water Cay. Conus toriuosopunctatus Youla Plate 3, figures 6, II
Conus tortuosopunctatus Toula, 1911, Jahrb. der K-K. Geol. Reichsan - Stalt,;" Wien, vol. 61, pl ise7.ph iat ie. 121,
227 Costa Rica MrocENE—OLSSON 55
Shell small, with a high smooth, conic spire of about 12 whorls the earlier post-nuclear whorls are finely nodulated, the succeeding have the sutures carinated and then simple; body- whorl with 20 or more strong, beaded, spiral cords; three spiral cords are usually double or incised with a median line, especially on the anterior canal; the beading of the spiral cords is develop- ed best on their posterior portions; interspaces of the cords near- ly as wide as the cords themselves, smooth or with faint, longi- tudinal lines and frequently a smail median spiral.
bengtia) 20.) breadth 131mm. Gatun Stage: Mt. Hope.
FAMILY TURRITIDA
Genus TURRICULA Schumacher
SECTION SURCULA H. anp A. ADAMS Turricula favinoides, u. sp. Plate 4, figure 6
Shell rather large and moderately solid; whorls about 12 in number, heavily sculptured with ribs and spiral cords; nucleus of two smooth, slightly bulbous whorls; post-nuclear whorls in- creasing uniformily in size; concave above, widest below the per- iphery; suture strongly appressed with a spiral cord in front of it; anal fasciole flat or concave, appearing nearly smooth except for about three, small, widely spaced spirals; remainder of whorls strongly sculptured with 8 or 12 axial ribs which are lacking from the anal fasciole and from the base of the last whorl; spiral sculpture of 3 or 4 strong cords on the spire-whorls, becoming 10 or more on the last whorl in addition to those of the anterior canal; finer, intercalated threads are present between the main spiral cords on the later whorls; the interval between the spirals is finely, longitudinally striated by long growth lines: anterior canal long and nearly straight, slightly calloused along
56 BULLETIN 39 228
the inner lip; outer lip thin, with a shallow anal sinus, lying in the sutural fasciole.
Length 54, diameter 20, last whorl 38 mm.
This elegant species may be compared with the 7. jaguenszs Sowerby from the Santo Dominican Miocene and the recent 7. lavinza Dall from the West Coast of Mexico. The Dominican species lacks the sutural cord, as well as differing in details of its sculpture. 7. davinia, appears to be more closely related, differ- ing in its finer sculpture and proportions.
Gatun Stage: Hill ra, Banana River. Turricula favinoides var. limonensis, n. var. Plate 4, figure 12
Shell of medium size and with about 10 whorls; nuclear whorls about 2 in number, at first smooth, becoming finely rib- bed on the last half; post-nuclear whorls spirally and axially sculptured; anal fasciole concave, smooth, with a strong spiral cord, just in front of the appressed suture; axial sculpture of about 8 sharp or pointed ribs on the periphery of each whorl, lacking entirely from the anal fasciole and from the base of the last whorl; spiral sculpture of 3 cords on the early spire-whorls becoming more numerous on the later, due to the intercalation of additional spirals; on the last whorl, the spirals number about 22,including those on the canal; anterior canal long and straight; anal sinus shallow; outer lip thin.
Length 47, diameter 18, last whorl, 32 mm. 67 Dy 47
This shell when full-grown appears distinct from the /av- notdes but young specimens cannot be easily separated. This variety differs from the typical form, by its sharp, narrow, and widely spaced axial ribs, which produce a sharp, angled and not rounded shoulder to the periphery cf each whorl. The anal tasciole is also wider and more smooth.
Gatun Stage: Toro Cays. Port Liman, coralline limestones.
ee
229 CosTa Rica MIocENE—OLSSON
cA =]
Turricula taurina, n. sp. Plate 5, figure 3
Shell smali, slender and fusiform; mucleus of nearly two smooth, convex whorls; post-nuclear whorls about seven; su- tures appressed and bordered by a spiral cord; fasciole about one-half the width of the spire-whorls, smooth; the sculpture of the whorls is spiral, axial ribs lacking; on the spire-whorls there are four raised spiral cords with the flattened interspaces about three times the width of the spirals themselves; on the last whorl and anterior canal, there are 18 or 1:9 spirals; anterior canal, long, slender and strongly twisted or curved; outer lip simple; anal sinus as indicated by the growth lines is wide, moderate- ly deep and occupies the middle of the fasciole.
Length 25, diameter 8, last whorl 16 mm.
This isa small and delicate species, characterized by its slender, fusiform outlines and lack of an axial sculpture. The spirals are narrow, raised cords, rather widely spaced. The single specimen was collected at Bocas del Toro.
Gatun Stage: Bocas del Toro, Panama.
Turricula Terryi, n. sp. Plate 5, figure 4
Shell small, slender and fusiform, with a long, evenly taper- ing spire and slender, twisted anterior canal; nucleus of two, smooth, small whorls, followed by 8 sculptured post-nuclear; su- ture appressed, bordered by a low, but wide, spiral cord; upper half or more of each spire-whorl, flat or slightly concave and forms the fasciole; fasciole nearly smooth or sculptured with few, low, faint spirals; early spire-whorls strongly sculptured with 12 or 13 axial ribs which are most heavy on the periphery of each whorl; these ribs become progressively weaker and on the last whorlare only feebly developed on the shoulder or periph- ery; spiral sculpture persistant but weak, consisting on the spire- whorls of 4 cords on the lower half, alternating with much weak- er spirals; the fasciole as already noted is nearly smooth, or very weakly spiralled; on the last whorl, there are 20 spirals, includ-
58 BULLETIN 39 230
ing those on the anterior canal; aperture elliptical with a thin outer lip and long twisted anterior canal.
Length 23.5, diameter 7, last whorl 15 mm.
The small exposure of Gattn shales, on the east shore of Columbus Island, about one mile north of the city of Bocas dei Toro, has yielded a large number of small interesting gastropods. More than a dozen species of Pleurotomids were collected at this place by Mr. R. A. Terry and the writer during a short period of collecting.
The 7. faurina, also from this locality, is somewhat similiar but differs by its more convex whorls, deeper fasciole and prati- cally lacks all axial sculpturing.
Gaiun Stage: Bocas del Toro, Panama.
Genus TURRIS Bolton Turris albida Perry Plate 4, figres 1, 2
FPleurotoma albida Perry, 1811, Conch. Expl., pl. 32, fig. 4.
Lleusotoma virgo Lamarck, 1822, An. s. Vert., vol. 7, p. 94.
Pleurotoma haitensis Sowerby, 1849, Quart. Jour. Geol. Soc. London, VOL6)) p.'50:
Pleurotoma virgo Moore, 1853, Quart. Jour. Geol. Soc. Lond., vol. 9, ps h3o:
Pleurctoma barretti Guppy, 1866, Quart. Jour. Geol. Soc. London, vol. 22." p. 290) pl. 7, ae! 6.
Pleurotoma antillarum Crosse, 1865, Journal de Conchyliologie, vol. 13)\p.\34,0pl. 1, fe7/8. Not ot d’ Onbigny.
Turris (Surcula) virgo Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, PP: 206))\207)
Pleurotoma albida Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, PENT pA Zo. play digsksal,
Pleurotoma albida Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila: voly'632)\,p.)13243.
Pleuvotoma haitensis Cossmann, 1913, Jour. de Conchyliologie, vol. 61, (pi) TO) pl) a es. maAY
Pleuvotoma cf. antillarum Cossmann, 1913, Jour. de Conchyliologie, VOL NOT. LAU pla ies ie ey
Turris albida Dall, 1915, Bull. 90, U. S. Nat. Museum, p. 38, pl. 5, fig.
13, pl. 14, fig. 7.
231 Costa Rico MIOCENE—OLSSON 59
furris albida Maury, 1917, Bull. Amer. Pal., vol. 5, p. 214, pl. 8, figs. 4-8.
This is a common species in the Gatun beds of Central Amer- ica. ‘T‘he usual form is a large, strongly sculptured and should- er shell, which corresponds best to the variety described by Guppy from Jamaica as barrett, Our largest specimen from Toro Cays, Panama measures as follows: Length 86, diameter 27, last whorl 56 mm. 7. aldzda, is alsoa common fossil in the Miocene of Santo Domingo, Jamaica, Trinidad and Venezuela. Its geological range is from the Oligocene to the recent.
Gatun Stage: Gatun, C. Z. Toro Cay, Water Cay, Panama.
Genus LEUCOSYRINX Dall
Leucosyrinx chforis, n. sp. Plate 5, fignre 19
Shell subfusiform, thin, with an acute spire and a long, straight, anterior canal; nucleus small, globular of about 2, smooth, convex whorls; post-nuclear whorls about 7; the spire- whorls are strongly angled in the middle, leaving a wide, con- cave or sloping zone about the upper suture; the sculpture con- sists of small, more or less confluent tubercles on the periphery of the whorls and number on the last whorl about 14; in addi- tion, the whole surface is strongly sculptured with raised, sub- equal, spiral threads; base strongly contracted tothe long, straight, anterior canal; the outer lip is broken; anal sinus, as indicated by the growth lines, lies inthe concave zone, close to the upper suture.
Length 14.50, diameter 5.00, aperture 7.25 mm.
This appears to be a true Leucosyrinx, characterized by its thin shell, long, anterior canal and acute spire. It occurs in the Bocas shales with Drillia Itthocolletotdes, cocostna, Terebra benth- alts var. bocasensts and others, shells which are all closely related to recent deep-water species.
Gatun Stage: Bocas del Toro.
aS)
60 BULLETIN 39 23
Genus ANCISTROSYRINX Dall
Ancistrosyrinx elegans Dall, variety Plate 4, figure 17
Ancistrosyrinx elegans Dall, 1881, Bull. Museum Comp. Zool., vol. 9, P. 54. Anctstrosyrina elegans Dall, 1889, Bull. Museum Comp. Zool., vol. PS pt, p. 7Ols se, dace an The discovery of this elegant, recent species in the Gatun beds of Costa Rica is of more than usual interest, the record being based on a single, small but otherwise quite typical speci- men from Hill 1a of the Banana River. Comparison with Dall’s figure in the Blake Report, shows no important difference, ex- cept that the sculpture of the fossil shell is somewhat finer. A. clegans, is recorded by Dall from the Florida Reefs and from 805 fathoms of water off Havana, Cuba. The dimensions of our shell is as follows: Length 15, diam- efer 5:75, last wHorh1 1mm.
Gatun Stage: Hill ra, Banana River. Ancistrosyrinx Dalli, n. sp. Plate 4, figure 16
Shell larger than the preceding and very nearly smooth; nu- cleus very small and smooth; post-nuclear whor!s seven or more with a large and prominent, dentate or spinous keel; between the keel and the suture, there is a wide, nearly flat area, which carries a strong, smooth, median, spiral cord, behind which lies the small concave anal fasciole; the rest of the shell is smooth and without spirals;on the last whorl, the peripheral keel carries about 18 short, broad, spiny teeth; the canal is long, slender and straight; outer lip thin and fragile.
Length 14, diameter 7, last whorl 9.5 mm. 20 IO, 15 mi:
Two specimens are represented in our collection, and the smaller but more perfect specimen is selected as the Holotype.
It differs from the described species of Avxcistrosyrinx Dy its
233 Costa RicA MIocCENE—OLSSON 61
nearly total lack of spiral sculpturing. The species is named for Dr. W. H. Dall of the National Museum, the author of the genus Azcistrosyrinz.
Gatun Stage: Toro Cay, Water Cay, Panama.
Genus DRILLIA Gray Drillia venusta, Sowerby Plate 4, figure 9
fleurotoma venusta Sowerby, 1849, Quart. Jour. Geol. Soc. London, Wolpe 50; Pl. 10, fig.' 7.
Lleurvotoma venustuii, Guppy, 1856, Quart. Jour. Geol. Soc. London, VOL 22), pi 7289;
Turris (Drillia) venusta, Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 208.
Pleurotoma venusta, Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32, Pe 527-
FPleurotoma venusta, Guppy and Dall, 1896, Proc. U. S. Nat. Museum, vol. 19, p. 305.
Dyitiia venusia Maury, 1917. Bull. Amer. Pal., vol. 5, p. 216, pl. 8, 19S) 1.35514.
The Drillia venusta and consors of Sowerby, are the two commonest Pleurotomids inthe Gatun beds of Costa Rica. Both are characterized by a neat, regular and reticulate sculp- ture of spirals and axial ribs. In vwenusta, the anal fasciole is narrow, and appears asa deep constricted zone, encircling the upper part of each whorl. The spirals are narrow bands, pro- duced between deep, incised lines and in typical specimens nuim- ber on the last whorl about 18. The canal is rather long and usually more or less twisted.
Drillta venusta, occurs also in the Miocene of Jamaica, Trinidad and in Santo Domingo.
Gan Stage: Gatun, C.Z.
Banana River, Port Limon. Drillia consors, Sowerby Plate 4, figures 8, f0, i3
Pleurotoma consors, Sowerby, 1849, Quart. Jour. Geol. Soc. London, WO.) OF) p50.
62 BULLETIN 39 234
Fleurotoma consora, Guppy, 1866, Quart. Jour. Geol. Soc. London, VOL} 22)cpaBe:
Turris (Drillia) militaris, Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15). 207. (Not? of Hands, 1842)
Meurotoma coisors, Guppy, 1876, Quart. Jour. Geol. Soc. London, vol. 32, P- 527-
Drillia militaris, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, and SETIeSs) '(p..337.
Fleurotoma alesidota, Dall, var. magia Bose, 1908, Bol. Inst. Geol de Mexico, No? 22, p. 47, pl. 05, fies. 30, 31 sao. ele
Pleurotama sp. aff. Pl. alesidota (Dall) var. mactlenta Toula, 1911, Jahrb. der K-K Geol. Reichsanstalt Wien,vol. 61, p. 506, pl, 30, fig. Ir. Not alesidota or macilenta Dall, 1889.
Drillia consors Brown and Pilsbry, 1911, Proc. Acad Nat. Sci. Phila., VOLAOZ pena A 5s
Drillia consors, Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p. 20, pl. 2, figs: 8-14.
Dyillia consors, Maury, 1917, Bull.. Amer. Pal., vol.5\ pera) plc, figs: 15; 16.
This species is easily distinguished from the venusta, by its more slender form, wide but shallow anal fasciole, sharper sculp- ture and straight, longer, anterior canal. ‘The most common variety, illustrated by figure 10, averages in length about 35 mm. The axial ribs are numerous (about 28), straight .or slightly oblique and only slightly heavier than the raised spiral cords. In addition the surface is overrun by fine and almost mi- croscopic spirals. Figure 8 of a large variety from Grape Point, has about 9 whorls and measures 60 mm in length. ‘The early spire-whorls have the usual sculpture as seen on typical consors. On about the seventh whorl, the ribe increase suddenly to near- ly twice their normal number. ‘They gradually become obsolete, so that on the last whorl, pratically only the spirals are left. The fine, microscopic spirals seen on typical comsors are lack-
ing. The Drillia alestdota and its variety macilenta Dall of the recent West Indian fauna, are closely related to consors.
235 ~ Costa RicA M1ocENE—OLSSON 63
Caren Stage: Mt. Fope, C. Z. Banana River, Port Limon. Boras.
Dritlia cf. moensis, Gabb
Drillia moensis, Gabb, 1881, Jour. Acad. Nat. Sci. Phila., vol. 8, 2nd series, Pp. 351; pl.’ 46, fig. 33.
A single small speimen of a Drillza was collected from the coral limestones near Limon, which agrees partly with Gabb’s figure and description. Our specimen has 7 instead of 8 ribs, and lacks the sutural spiral mentioned by Gabb. The dimen- sions of this shell are: Length 12.5, diameter: 4.5, last whorl gy cat,
Gatun Stage: Port Limon. Drillia papaya, n. sp. Plate 4, figure 5
Shell of medium size, slender and subfusiform; whorls 1o (mucleus eroded); anal fasciole, a deep concave zone, occupying about 1-3 of a whorl interval and bordered above by a small su- taral cord; sculpture of heavy, knob-like ribs, crossed by spiral threads of different strengths; axial ribs about 6 on the last whorl, lacking from the fasciole and from the base and anterior canal; spiral sculpture of primary secondary and tertiary threads, there being about four primary threads on each spire-whorl, on the last whorl there are 8 or more primary threads, with an intermediate secondary and finer tertiary threads on each side; the anterior canal has numerous threads of varying strength; aperture long and subelliptical; anterior canal long and slightly bent backwards; anal sinus asindicated by the growth lines is deep and wide.
Length 32, diameter 10, last whorl 19 mm. A single specinien was coliected from certain sandstones of Lower Gatun age from a small tributary stream of the Rio
Cocles near Old Harbor, C. R. The recent Drillza haliostrephis Dall from the Gulf of Mexico appears closely related, but the fos-
64 BULLETIN 39 2436
sil shell has a more slender spire and slightly longer anterior canal,
Gatun Stage: Rio Coces, C. R.
Drillia coccos, nu. sp. Plate 5, figure 7
Shell small, slender and fusiform; whorls to or more, nu- cleus eroded; fasciole deep and concave, about % of the width of a spire-whorl, bordered above by a strong sutural cord, other- wise smooth; sculpture of strong axial ribs, which are lacking from the fasciole and from the base of the last whorl; these ribs number on the last whorl about 10; the ribs are crossed by strong, raised cord-like spirals, there being at first 2 on the earl- lest spire-whorls and on the later 3; the last whorl shows 8 spirals and the wide interspaces are occupied by 3 or more small- er ones; additional and more crowded spirals occur on the long slender canal; anterior canal long and slender, slightly bent.
Length 21, diameter 6, last whorl 12 mm.
A small, very slender and fusoid species with strong, persis- tent spiral and axial sculpture. A single specimen was collect- ed at Coco Plum, a small cocoanut plantation about 40 miles east of Bocas del Toro.
Gatun Stuge: Coco Plum.
Drillia chiriquiensis, n. sp. Plate 5, figure 2
Shell solid of medium size; whorls 7 (apex lost), of slightly convex profile; anal fasciole small but quite deep and nearly smooth; suture strongly appressed and bordered just below by a strong, elevated spiral cord; sculpture of numerous, somewhat oblique riblets and numerous weak spirals; the ribs commence at the lower edge of the fasciole, pass slightly obliquely across the whorls to the suture or on the last whorl across the base to the canal; the ribs on the last whorl number 15 or 16; spirals of numerous weak threads which are generally alernating in charac-
ter; the spirals are very small onthe fasciole, quite large and |
237 Costa RicA MirockENE—OLSSON 65
coarse on the anterior canal; the outer lip is heavily thickened by a large rib; anterior canal short and bent.
Length 25, diameter 8, last whorl 15 mm.
A single shell from Bocas, serves as type for this species. It recalls the D. consors Sowerby, but differs in its much finer sculpture.
Gatun Sage: Bocas del Toro, Panama.
Drillia citria, n. sp. Plate 5, figures 12, 13
Shell small; whorls about 8, of which the first 2 belong to the small, obtuse, smooth nucleus; the post-nuclear whorls are strongly sculptured with ribe and spirals; the upper 1-3 of the spire-whorls carries a concave fasciole, bordered posteriorly by a strong sutural cord to the rst post-nuclear whorl; the fasciole is smooth or only faintly sculptured with fine spirals; the sculpture consists on the last whorl of about 9 ribs which fade out rapidly on the base and on the fasciole; the ribs are crossed on the early spire-whorls by 2 spiral cords, becoming 3 on the later; on the last whorl there are 12 more of which 6 are on the anterior ca- nal; the intervals between the spirals, are 3o0r more times as wide as the spirals themselves and are sculptured with fine spirals threads; aperture sub-elliptical, with a short and slightly twisted anterior canal; anal sinus moderate.
Height 11, diameter 4 mim.
A small species, common in the coralline phase of the Gatun at Port Limon.
Gatun Stage: Port Limon.
Drillia aquanica, n. sp. Plate 5, figures 16, 17
Shell small, slender with a long spire and a shorter last whorl; whorls about 11, the nucleus composed of 2 small, smooth convex whorls and about 9 post-nuclear; whorls strongly sculp- tured with ribs and spiral cords; fasciole nearly 1-3 the width of the spire-whorls, marked with fine spirals and bordered posteri-
66 BULLETIN 39 23%
orly by a strong sutural cord; the spiral sculpture consists on the early whorls of 2, later 3 and finely 4 cords on the penulti- mate whorls; the last whorl has 8 spiral cords and 10 others on the canal; the spiral cords are separated by interspaces of slight- ly greater width; the interspaces are finely striated with small spiral threads; the axial sculpture consists of about 9 knot-like ribs over which pass the enlarged spiral cords; the ribs pass over the base of the last whorl to the canal and slightly over the fac- ciole; aperture elliptical with a short, straight canal.
Height 18, diameter 6, last whorl 9 mm.
This is a diminutive of Drillia fusiformis of the Miocene of Santo Domingo, the two species having practically the same sculpture. Drillia fusiformis of 11 whorls measures 35 mm in length, while aguanica of the same number of whorls has a length of only 19mm. The Maury collection contains a small
but quite typical specimen of aguanica from the Rio Gurabo, Santo Domingo.
Gatun Stage: Water Cay.
Drillia limonica, n. sp. Plate 4, figure 15
In general form like Drillia consors Sowerby, with a long spire and but slightly shorter anterior canal; whorls about 8% of which 1% belongs to the small, smooth, blunt nucleus; the post-nuclear whorls are sculptured with ribs and weaker spirals ; on the 1st three whorls, the anal fasciole is but slightly differ- entiated; it increases slowly in strength until on the later whorls it has become a concave zone asin comsors; a sutural cord is in- troduced on the 2nd post-nuclear whorl and is present on all the succeeding whorls; on the spire-whorls there are about 11, wide- ly spaced ribs; these ribs are lacking from the anal fasciole, ex- cept on the very earlest whorls, where they are faintly continued towards the upper sutures; on the last whorls, the ribs are small- er and number about 13; between these main ribe on the last whorl there appear occasionally smaller and shorter ribs which do not reach to the edge of the fasciole; there are 5 faint spirals
239 Costa Rico M1rocENE—OLSSON 67
crossing the ribs on the spire-whorls and very much finer ones in the anal fasciole; the last whorl shows about Io spirals together with others on the canal; aperture elongate with the anterior canal rather long and straight.
Benet ro, diameter '6;5 mm.
Like the Drillia consors in form, this species is distinguish- ed by its fewer ribs and shallower anal faciole. The ribs on the earlier whorls are oblique, but become very nearly straight and vertical on the later.
Gatun Stage: Port Limon. Drillia bocatoroensis, n. sp. Plate 5, figure 6
Shell of medium size, solid and strongly sculptured with ribs and spirals; whorls about 11, the nucleus eroded in the type specimen; fasciole occupying about 1-3 of the width of the spire- whorl, with a large sutural cord above which the edge of the su- ture is finely and regularly beaded or granulated; the surface of the fasciole is slightly undulated by the obsolete ends of the ribs, otherwise smooth; axial ribs about 8 on the last whorl, straight and nearly in line across the face of the spire-whorls to the apex, but interrupted and lacking from each fasciole; spiral sculpture of the few raised cords with wider interspaces; there are 3 spirals on the spire-whorls, about 6 on the last, with 9 more on the anterior canal; the interspaces in addition carry fine and sub- microscopic spiral lines; the base of the last whorl is somewhat constricted with a short anterior canal.
Length 23, diameter 8, last whorl 13.5 mm.
From the several species of fusoid Pleurotomids in the Gatun beds of Panama and Costa Rica, this species differsin its heavy, persistent ribs and sharp spiral sculpturing. ‘The fasciole is nearly smooth, bordered above by a heavy sutural cord.
Gatun Stage: Bocas del Toro.
68 BULLETIN 39 240
Driffia theobroma, u. sp. Plate 5, figure 1
Shell subfusiform; whorl 7 (apex broken); fasciole a narrow, concave zone, not strongly differentiated from the rest of the whorl, spirally sculptured and bordered above on the early spire- whorls by a sutural cord; sculpture consists on. the spire-whorl of about 10 low ribs, becoming more numerous and subobsolete on the last; the spiral sculpture consists of raised, alternating, revolving cords; there are 2 or 3 spiral cords on the early spire- whorl, becoming 5 or more on the later; the last whorl has about 1o spirals in addition to those on the anterior canal; between each pair of the main or primary spirals there is a smaller, sec-
ondary spiral; base rounded or contracted tothe slender anterior canal,
Length 21, diameter 7.5, last whorl 13 mm.
it is possible that this species, belongs to the genus Glypho- stoma, as the type specimen seems to show a slightly thickened and inflated outer lip. The aperture is however so completely filled with a rocky matrix, that this observation cannot be fully verified. The axial sculpture is practically lacking from the last whorl, leaving a surface simply marked with the rough spirals.
Gatun Stage: Bocas del Toro. Drillia aurantia, nu. sp. Plate 5, figure 15
Shel! small, solid; whorls about 9; the nucleus consists of about 1% smooth, convex whorls which through the gradual in- troduction of the ribs, followed by the spirals blend in with the post-nuclear whorls; the anal fasciole is rather wide (about 1-3 of the spire-whorl) and shallow; it uppears very early and car- ries from the start a strong, sutural cord; the sculpture consists of numerous closely spaced, linear ribs which are absent from the anal fasciole: these ribs number on the last whorl about 21; the spirals are uneven threads which are strongest below and on the base; the fasciole also carries one or more fine threads except
24l Costa Rica MIOCENE —OLSSON 69
on the base of the last whorl; the spirals do not cross the ribs so that the summit of the ribs are left nearly smooth; aperture small with a short, slightly twisted anterior canal.
Length 11, diameter 4 min.
A small species sculptured like the D. elegans Conrad of the Chesapeake Miocene of Virginia, but with more numerous and more closely spaced ribs.
Gatun Stage: Port Limon.
Drillia musacina, n. sp. Plate 5, figures 27, 2&
Shell small, slender; the nucleus of about 2 whorls, the 1st blunt, convex and smooth,followed by the 2nd which is strongly carinate; there are 7 post-nuclear whorls of slightly convex pro- file; the fasciole is a narrow band on which the ends of the ribs become strongly deflected to the left; the axial sculpture of about 16, narrow ribs on the last whorl; the ribs are continuous across the spire-whorls to the upper sutures but become bent and strongly deflected on the fasciole; on the last whorl, the ribs are continued to the base of the canal; the spirals vary somewhat ' in strength but are usually subregular spiral bands of which there are 7 or 8 on the spire whorls and about 14 on the last whorl and the canal; the back of the last whorl is strongly humped; the outer lip was probably thin (broken in all our specimens); canal short, straight.
Length 9, diameter 2.75, last whorl 5.00 mm. A small species abundant in the Gatun beds of the Banana River.
Gatun Stage. Hill ra, Banana River. Hill 3, Banana River.
Drillia estrellana, nu. sp. Plate 10, figures 31, 32
Shell very small, solid, porcellaneous; slender with a long spire and a short body-whorl; nucleus small of 2 smooth, convex
70 BULLETIN 39 Ee eg
whorls; post-nuclear whorls about 7; sculpture with strong ribs and impressed spiral lines in their interspaces; the anal fasciole is a constricted band about the upper :-4 of the whorl and which passing over the ends of the ribs dislocates them asin Zerebra, there are 11 ribs on the last whorl which commencing at the up- per suture pass across the sutural fasciole but somewhat dimin- ished in strength, continue across the face of the whorlina slightly oblique direction to the columellar region; the summits of the ribs are smooth but the interspaces are sculptured with impressed lines produced by spiral bands of regular width; there are 4 such bands on the spiral-whorls and about 120n the last whorl, including a few on the canal; aperture small, subellipti- cal and a short twisted canal.
Height 7.5, diameter 1.75 mm.
A small species of Terebroid appearance, the ends of the ribs being strongly dislocated as they pass across the small anal fasciole.
Gatun Stage: Coll. 7, Estrella River. Drillia limonetta, n. sp. Plate 5, figure 10
Shell small, solid, porcellaneous; whorls about 8, coarsely ribbed and polished; fasciole absent; the sculpture consists of moderately strong ribs which on the spire-whorls pass suture to suture but are only slightly flexed near the upper suture; on the last whorl, the ribs number about 11; the last rib is much en- larged and forms the outer lip; the ribs are but shortly continu- ed upon the base and do not reach the canal; in addition, the surface is covered with fine, irregular threads which are nearly obsolete on the upper portions of the whorls but are strong on the base and on the canal; aperture subelliptical with a short canal, somewhat bent or excavated ventrally; anal sinus simply as a vertically directed notch at the junction of the outer lip with the body-whorl and is bordered by a tooth-like denticle on its in- ner side.
Length 12.25, diameter 5.00, last whorl 7.25 mm.
243 Costa Rica MrocENE—OLSSON
NJ —
A small, solid and porcellaneous species, belonging to Dall’s Cymatosyrinx. The sutural fasciole being absent, the ribs are continuous to the upper suture and the surface is finely striated with small, irregular spiral threads.
Gatun Stage: Port Limon. Drillia carruca x. sp. Plate 4, figure 11
Shell of medium size, solid; whorls 7 plus (apex broken), rather rapidly tapering; no fasciole; axial sculpture of about 8, narrow ribs with deep and wider interspaces; the ribs commence at the top of the canal and pass across the whorls to the sutures; the ribs are nearly in line across the spire-whorls to the apex; spiral sculpture of slightly elevated cords, separated by wide, flat interspaces; there are about 7 spiral cords on the spire-whorl; with 12 on the last whorl; aperture sub-elliptical, the outer-lip somewhat thickened and a short canal.
Height 18, diameter 7.25, last whorl Ir mm.
This shell resembles Dreilia musa in its general sculpture of fine, widely spaced, spiral cords.
Gatun Stage: Coll.4, East Grape Point Creek. Driilia limonensis, n. sp. Plate 5, figures 8, 9
Shell of medium size and in form recalling D. venrusta So- werby; nucleus of nearly three, smooth convex whorls; post-nu- clear whorls 8; anal fasciole scarcely perceptible on the early whorls, becoming on the later, simply a narrow, constricted zone, bordering the suture; sculpture of rather numercus riblets, which are abruptly bent to the left near the posterior suture on crossing the constricted anal fasciole; three ribs number on the last whorl about 15; the spiral sculpture consists of very fine threads; the space between each pair of adjacent spiral threads is finely and mi- croscopically decussated by the raised lines of growth, producing a shagreening of the whole surface; in mature,shells, the last rib becomes unusually large and heavy, producing a large and strong-
72 BULLETIN 39 444
ly thickened outer lip; canal short, slightly bent inward or out- wards; callous on the inner lip.
Length.20, diameter 7.5, last whorl 11 mm.
The general form of thisshell is that of Drz//ia venusta Sow- erby, which it also recails in its constricted anal fasciole. The whole surface appears ininutely shagreened, an effect produced by the fine, sub-microscopic, spiral threads as well as a minute de- cussation oftheir interspace. This species is fairly abundant in certain sandy layers found intercalated in the hard and gener- ally barren coral-reef limestone along the sea-front at Port Limon.
Gatun Stage: Port Limon. Drillia Cristobali, n. sp. Plate 5, figure I
Shell of the general form of wenusta, rather solid; whorls 6, (apex broken in the type specimen); anal fasciole a narrow con- stricted zone, bordering the upper suture; no sutural cord; sculp- ture of numerous, straight, parallel ribs and fine spirals; ribs on the early whorls about 12, becoming about Ig on the last; on the anal fasciole, the ribs are shortly deflected or bent; spirals very numerous and on slight magnification seen to consist of narrow, even raised threads with interspaces of nearly twice the width of the spirals themselves; anterior canal rather short and slightly bent backwards; aperture sub-ellipitical with a thin outer lip; anal sinus small and shallow.
Length 20, diameter 8.5, last whorl 11.5 mm.
This species, like the /émonensis, has the general form and constricted fasciole of Drillia venusta Sowerby, but differs by its smaller size, short canal and quite different spiral and axial sculpture. D. limonensis, is more closely related, but that species differs in its more slender form and peculiar submicro- scopic sculpture.
The single specimen serving as the type, was collected from the Gatun shale, exposed about a mile north of the city of Bocas del Toro, on Columbus or Cristobal Island.
245 CosTa Rica Mr1ocKENE—OLSSON 73
Gatun Stage: Bocas del Toro, Panama.
Drillia bocasensis, n. sp. Plate 5, figure 5
Shell of medium size, solid; nucleus of about 2 smooth whorls; post-nuclear whorls about 10; anal fasciole a narrow con- stricted zone, nearly filled with a large, sutural cord; axial sculp- ture of large, heavy ribs, about 9 on each whorl; the ribs pass from suture to suture and on the last whorl across the base to the anterior canal; a large, hump-like rib is developed on the back of the last whorl, formed during a resting stage; no spirals; the growth-lines produce a minute but characteristic sculpture of heavy, raised threads, which pass obliquely across the surface of the whorls and ribs, and follow the curve of the anal sinus across the fasciole; the sutural cord is finely granulat- ed by the raised growth-lines; canal short but straight; outer lip large, with a deep anal sinus at its posterior union with the body-whorl.
Length 22, diameter 8, last whorl 12 mm.
This species should more properly be referred to the genus Cymatosyrinx Dall, based on the Pleurotoma lunata H.C. Lea, a Chesapeake Miocene fossil. The form and general sculpturing of docasensis is similiar to that of many species of this group but it may be recognized at once by its very peculiar, submicroscopic sculpture formed by the close, heavy growth-lines.
Gatun Stage: Bocas del Toro. Drillia aquaensis, n. sp. Plate 5, figure 25
Shell small and solid; nucleus of 1+ whorls (mostly broken on type specimen); post-nuclear whorls about 8; anal fasciole lacking; axial sculpture of about six, large, heavy and straight ribs,which pass from suture and across the base of the last whorl onto the anterior canal; each set of ribs is in a straight line from the canal across the spire-whorls to the apex; spiral sculp- ture of about 8 impressed lines which produce a series of flat but
74. BULLETIN 39 246
quite wide, spiral bands; on the last whorl there are about 22 spiral bands; much finer near the suture, larger and wider on the middle of the whorl! and on the canal; canal short and slight- ly bent to the left; inner lip calloused.
Length 15, diameter 5.5, last whorl 8.5 mm.
Characterized by its six axial ribs and finer sculpture of flat, spiral bands. It belongs to the genus Cymatosyrinx of Dall.
Gatun Stage: Water Cay, Panama. Drillla musa, n. sp. Plate 5, figure 26
Shell about the same size as the preceding but witha more slender spire; nucleus of 2 smooth whorls (broken); post- nuclear whorls about 8; no fasciole; axial ribs about 9, pass from suture to suture and across the base to the anterior canal the ribs are in line across the whole series of spire-whorls; spiral sculpture quite heavy, consisting of raised threads with wider in- terspaces; the spirals number on the spire-whorls 8 or 9, and on the last whorl about 22; aperture large and suboval; anterior canal short, narrow and heavily calloused to form the inner lip.
Length 17; diameter 6, last whorl 9 mm.
Related to the preceding, but differs by its more slender form; with 9 instead of 6 ribs and a much coarser spiral sculp- ture.
Gatun Stage: Banana River, C. R. Drillia lithocolletoides, n. sp. Plate 5, figure 18
Shell small, glassy or translucent; nucleus of about 3 smooth, convex and glassy whorls, followed by 6 post-nuclear whorls; the whorls are strongly angled in the middle by tubercular-like ribs but leaving a concave zone about the upper sutures which forms the anal fasciole; on the last whorl, the ribs or tubercles number Ii; are set obliquely and on the back of the whorl, they are con- tinued slightly over the base; the surface is smooth and polished; last whorl slightly constricted above the short, straight canal; aperture subovate.
247 Costa Rico MrocENE-—-OLSSON 75
Height 10.75, diameter 3.25 mm.
The Drillia lithocolleta Watson is a_ recent deep-water species, dredged by the Challenger and the Blake, at several stations in the West Indies and off the Florida coast, from depths of 400 to nearly 1ooo fathoms. ‘The Bocas shell seems to be very closely related to the recent species; differing only from Dall’s figure in the Blake Report (plate 11, fig. 61), in being somewhat more slender, with heavier tubercles and a longer base.
Gatun Stage: Bocas del Toro.
Drillia cocosina, n, sp. Plate 5, figure 14
Shell small, giassy or translucent in texture; spire twice or more the length of the last whorl and the canal; whorls 8 plus, the earlier ones missing; no anal fasciole; the sculpture consists at first of a lower row of small tubercles, but a second or upper row soon begins to appear and on the last whorl, this upper set of tubercles is very nearly equal to the lower; the last whorl has about 13 of these tubercles, in addition the whorl is finely sculp- tured with fine, regular, closely spaced spiral threads which cov- er the entire whorl and the canal, but leaving the tops of the tubercles smooth; aperture ovate, with a short, twisted canal.
Wength 12) diameter 2.5 mim:
This interesting species from the shales near the city of Bocas del Toro, is related to the recent Dyillia oleacitna Dall dredged from rather deep water in the Gulf of Mexico and else- wherein the West Indies. In differs from that species in being less slender and with a larger body-whorl.
Gatun Sage: Bocas del Toro.
76 BULLETIN 39 248
Genus GLYPHOSTOMA Gabb Glpyhostoma dentifera, Gabb.
Glyphostoma dentifera Gabb, 1872, Proc. Acad. Nat. Sci. Phila., vol. 2A, P..270, PIAL, HS. <Ae Glyphostoma dentifera Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p.
210. Glyphostoma dentifera Dall, 1889, Bull. Mus. Comp. Zool., vol. 18, Pe 2s\ pL LES:
Glyphostoma dentifera Cossmann, 1913, Jour. de Conchyliologie, vol. 61 pi 3k, ply 2. es. 55, O, 7A
Glyphostoma dentifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 225, pl. 9, fig. 16.
A single specimen of this species was collected from the cor- alline phase of the Gatun, near Port Limon. The shell showing 5 whorls (apex lost), measures 18 by 8mm. The pecular sha- greening, characteristic of the genus is strong and easily seen.
This shell agrees closely with the figure given by Cossmann, based on a Dominican specimen. It differs in several respects from the specimen figured by Dr. Maury in her Dominican Fos- sils from a metatype, sent by Professor Gabb to the Cornell Mu- seum. Dr. Maury’s shellis larger, has a longer spire, and the microscopic shagreening is much finer.
Dall in the Blake Report, doubtfully refers to this species, an imperfect specimen collected in 15 fathoms of water at Sand Keys.
Gatun Stage: Port Limon.
Glyphostoma moinica, n. sp. Plate 5, figures 29, 30
Shell small, solid with a coarse, tuberculate, subreticulate sculpture and a thickened, enlarged onter lip; nucleus small, pointed and tapering of 3 or more, smooth, convex whorls fol- lowed by about 4 post-nuclear; the sculpture consists of about 14 narrow ribs continuous from the canal to the upper suture and crossed by even, spiral cords; their intersection form small
249 Costa RicA MIocENE—OLSSON a7
rounded beds or tubercles; immediately after the nucleus, there is but one spiral, soon followed by two and on the penultimate there are three; the last whorl has five spirals and three or four more on the canal; aperture elliptical with a large expanded out- er lip, smooth within and a deep, anal sinus at its upper end; canal short and straight.
Height 5.75, diameter 2.75, last whorl 3.75 mm.
A small species doubtfully referred to Glyphostoma, having the outer lip smooth within and lacking the peculair, submicro- scopic structure of typical Glyphostoma.
Its sculpture is coarse, the intersection of the ribs and spirals forming small beads or tubercles.
Gatun Stage: Port Limon.
Genus CYTHARA Schumacher Cythara terminula var. costaricensis, n. var. Plate 5, figures 21, 22
cf. Cythara terminula Dall, 1890, Trans. Wagner Free Inst. Sci., vol. Supe, P30, pl. 2, fig. 5.
The Cythara terminula Dall, to which the present shell ap- pears closely related, was described from the Caloosahatchie beds of Florida, of Pliocene age. The Costa Rican shell with the same number of whorls is somewhat small _ (typi- cal teriminula of 7 whorls, 16 mm), (variety cosfaricensis 7 or 8 whorls, 13 mm), and has 8 instead of g ribs. The details of the spiral sculpturing is very similiar, consisting of flattened bands separated by sharp channels. The spiral bands carry a central incised line, which produce the appearance of being in pairs.
C. cercadica Maury from Santo Domingo, is a larger species with higher spire and simple and not banded spirals.
Length 13, diameter 5, last whorl 10 mm.
Gatun Stage: Hill ra, Banana River.
Cytharella limata, n. sp. Plate 5, figure 20
Shell small, nearly smooth and porcellaneous; spire slightly
78 BULLETIN 39 250
shorter than the narrow aperture; nucleus of about 3% smooth, convex whorls, the last 4% turn being very finely and closely rib- bed; there are 3% post-nuclear whorls; sculpture consists of narrow, slightly oblique ribs which pass across the spire-whorls from suture to suture and on the last whorl follow down on the anterior canal to its tip; there are 8 ribs on the last whorl; the spaces between the ribs are wide, flat and smooth; the tops and sides of the ribs themselves are carved or etched with fine, sub- obsolete spiral lines; aperture linear-elliptical, the outer lip some- what thickened by the last rib, but smooth within; anterior ca-: nal long straight.
Height 9, diameter 3.50, last whorl 7mm.
Gatun Stage: Port Limon,
Genus BORSONIA Bellardi Borsonia cocoensis, n. sp. Plate 5, figures 23, 24
Shell small, biconic; whorls about 7; nucleus of 2, nearly smooth whorls, followed by 5 strongly shouldered post-nuclear whorls, the angle of which on the spire-whorls comes just above the lower suture; above the shoulder, the whorls are flat or slightly concave to the upper suture; sculp- ture with a double or triple beaded cord on the peri- phery and with alternating beaded spirals both above and below; on the last whorl, the primary spirals number 15 or 16, above the shoulder 3 or 4; the suture is bordered in front by a strong cord; pillar long, slightly twisted in front and with 2 sharp, pli- cations; the anal sinus lies on the peripheral angle as in typical Turris.
Length 16, diameter &, last whorl 11 min.
The present species is based on two specimens from Coco Plum about 30 miles east of the city of Bocas del Toro. The B. varicosa Sowerby from the Dominican Miocene, is a related species, differing in its porportionately longer spire.
Gatun Stage: Coco Plum.
252 Costa R1cA MI0ocENE—OLSSON 79
Genus SCOBINELLA Conrad Scobinella Morierei (Laville) Cossmann Plate 4, figures 3, 4
Euchilodon Morierei Laville, in Cossmann, 1913, Jour. de Conchylio- logic, vol. 61, p. 34, pl. 3,’ figs 6, 7.
Shell of medium size, with spire and last whorl of nearly equal lengths; sculpture of numerous, close, bead-like spiral cords, 3 or 4 on the spire-whorls, 18 or more on the last whorl and canal; the sutural fasciole is rather narrow, shallow and concave and with 4 much finer beaded cords; the suture is bor- dered in front by a heavy beaded spiral, which on the early spire- whorls, nearly fills the entire fasciole ; columella with four sharp folds, the largest above; canal nearly straight; aperture narrow, with a lirated outer lip.
Length 49, diameter 15, last whorl 33 mm.
This is one of the most elegantly sculptured Pleurotomoids in the Gatun beds of Panama and Costa Rica. Described by Laville and Cossmann from Mindi in the Canal Zone, we have in addition collected the species from Toro and Water Cay, (where it is fairly common), Bocas del Toro and from Limon. ‘The pre- dominating spiral sculpture and columellar folds are very sug- gestive of Mitra.
Gain Stage: Mindi, C. Z. (Laville and. Cossmann)
Lloro aud Water Cays, Panama. Bocas del Toro, Panama. Port “Limon, coral limestones.
Genus HALIA Risso
ralia americana, n. sp. Plate 4, figure 7
Shell large, thin, buccinoid in form; spire elevated with shouldered whoris and deep sutures; whorls 3 plus, the earlier ones missing; the profile of the whorls is convex with a wide, shouldered, flattened or even slightly concave band just in front of the suture; the surface of the spire-whorls is smooth and un- sculptured but the early ones show very faint spiral bands; the
SO BULLETIN 39 252
upper half of the body-whorlis smooth but below, the whorl car- ties about 16, low, faint, spiral ribbons like those of MWalea but much less strong; these spiral ribbons average about 1.25 mm in width; the growth lines cross the face of the whorl from the tip of the canal to the shoulder angle in a slightly convex curve, the convex side of which is towards the aperture; from the shoulder angle to the suture the growth lines are reversed with the con- cave side facing the aperture; aperture broadly elliptical; the ca- nal was probably short and turned inward or ventrally.
Height 66, diameter 39 mm.
This truly remarkable species evidently belongs to the Pleu- rotomoid genus Afalza, of which the 4 known species are EHuro- pean and North African in their distribution. Its type species flalia Priamus Meuschen is living off the coast of Spain and North West Africa. According to Cossmann, there are three Tertiary speciesin Europe, WY. precedens Pant. from the Helve- tian of Italy, 7. Deshayeseana da Costa in the Tortonian of Portugal and Italy, and A. helicotdes Br. in the Plaisancian of the maritime Alps.
The European species are described as being entirely smooth, but the Gatun shell is sculptured on its lower half with ribbon- like spirals bands.
Gatun Stage: Wi. Hope, 0. 4
Genus CANCELLARIA Lamarck
Cancellaria dariena Toula Plate 6, figure 8
Cancellaria dariena Toula, !909, Jahrb. K-K. Geol. Reichsanstalt, Wien, vol. 58, p./ 31, pl. 25, fe! tg, (pl 2S, fesse:
Cencellaria dariena Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. PHila:;\wol. 63, p.345,\pl. 24) tgs! 3.) 4:
Canceliaria davienesis Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p. 51, pl. 4 figs. 9, Io. var. trachyostraca Brown and Pilsbry.
Cancellaria dartena, var. Toula, 1909, Jahrb. der K-K, Geol. Reich- sanstalt Wien, vol. 58, p. 37, pl. 28, fig. 11.
253 Costa Rica M1ocENE—OLSSON 81
Cancellaria dariena var. trachyostraca Brown and Pilsbry, tgtr, Proc. mead. Nat, sci, Phila., vol, 63, p:\ 345, pl. 23) figs. 1, 2:
This species is most abundant in the Canal Zone. It is dis- tinguished from the Barretét Guppy, which it most closely re- sembles by its shorter spire, larger body-whorl and a more irreg- ular spiral sculpture. <A large specimen from Mt. Hope, C. Z., has the following dimensions: Length 38, diameter 22; last whorl 30 mm. Shorter and more globose shells form the variety /fra- chyostraca Brown and Pilsbry.
Gatun Stage: Gatun and Mi, Flope, C. Z.
Water and Toro Cays, Panama. FAG COLES, OGTR,
Cancellaria Barretti Guppy Plate 6, figure 6
Cancellaria Barretti Guppy, 1866, Quart. Jour. Geol. Soc. London, VOL 722, p.. 289) pl. 17; fig. 11.
Cancellaria reticulata Gabb, 1873, Trans, Amer. Phil. Soc., vol. 15, p. 236. In part.
Cancellaria Barretit Guppy, 1876, Quart. Jour. Geol. Soc. London, Woll/3 2) |p. 520.
Not C. Barretti Maury, whichis C. Maury@, n. sp.
This species is the West Indian Miocene analogue or the re- cent Cancellaria reticulata Linné. ‘The most important differ- ence to be noted between thetwo shells is that in Barretiz the columellar plicee are more anteriorly situated and heavier. It is a Miocene fossil in the Bowden beds of Jamaica.
Gatun Stage: Banana River
Canceilaria Cossmanni, n. sp. Plate 6, figures 9, II
Shell of medium size, solid; nucleus of 3, small, smooth whorls; post-nuclear whorls 5, sutures deep; the whorls are slightly channeled or coronated just below the sutures; sculp- ture is evenly but coarsely reticulated by numerous, slightly ob- lique, narrow ribs, crossed by heavy, raised, narrow, spiral cords; the last whorl show 21 or 22 ribs; the spirals consists of narrow,
82 BULLETIN 39 254
raised cords, separated by wide, deep interspaces; there are 4 spirals on the spire-whorls, with 15 on the last whorl and the canal; base of last whorl rounded and contracted to the short, beak-like anterior canal; columella with 2, narrow, but strong plications, of which the upper is much the strongest; aperture sub-elliptical with the outer lip carrying 8, narrow, entering hire.
Length 25, diameter 14.5, last whorl 18 mm.
This shell should probably be considered as a variety of C. Barrett Guppy, but in the large series in our collection,the shell is always much smaller and with a much coarser and heaver sculp- ture. In some specimens, this sculpture becomes quite sharp and harsh, brought about by the crossing of the narrow spiral cords and equally narrow axial ribs.
It is an abundant and characteristic species of the Gatun beds along the Banana River, C. R.
Gatun Stage: Banana River. Cancellaria Mauryz, n. sp. Plate 6, figure 5
Cancellaria Barretti Maury, 1917, Bull. Amer. Pal., vol. 5, p. 226, pl. 10, fig. 1. Not of Guppy, 1866.
Shell large, ovate; an evenly conic spire in height less than the length of the aperture and a large, evenly convex last whorl; the nucleus is rather small, of 2 smooth, convex whorls followed by about 6 post-nuclear whoris; the profile of the spire-whorls is slightly convex with deep, distinct suture; the sculpture is can- cellate or reticulate, the spirals and ribs of very nearly equal strength; of the axial ribs there are 40 or more on the whorl, crossed by 24 spiral cords; on the spire-whorls there are 5 spiral cords becoming 7 on the penultimate; the spirals are fairly regu- lar, separated by interspaces 1% times their width, in which smailer spirals may occasionally appear; faint indications of rest- ing marks show as crowding of the ribs or as smooth, slightly humped spaces; aperture large, ovate; a thin outer lip with 17, long entering lirze; columella with 2 simple plicee, of which the
255 Costa Rico MroceENE-—-OLSSON 83
upper is much larger; a third is probably developed on the an- terior border of the canal; a well marked ridge about the base and a strong siphonal fasciole.
Height 36, diameter 23, last whorl 29, aperture 24 mm.
This shell was figured by Dr. Maury in her Dominican fos- silsas C. Barretti of Guppy, but it differs from that species in much shorter spire, more globose shell, larger aperture and by its columellar plicee. In the species closely related to the vefz- culata, such as Barretti, Cossmanni and others, the columellar plicee are heavy and the posterior one is usually more or less bifid. In this species, the posterior plication is large, wide and thin, especially in the interior of the sheil.
The Costa Rican collection is limited to a single, imperfect specimen, so we have used as the type of this species, the origi- nal figured specimen of Dr. Maury from the Miocene of Santo Domingo.
Gatun Stage: Water Cay.
Cancellaria epistomifera Guppy Cancellaria Mooreit Gabb. 1872, Trans. Amer. Phil. Soc., vol. 15, p. 236. Not C. Moorei Guppy, 1866.
Cancellaria epistomifera Guppy, 1876, Quart. Journ. Geol. Soc. Lon- don, ‘vol: 32, p: 520, pl. 28; fig. 9.
Cancellaria epistomifera Cossmann, 1913, Journ, de Conchyliologie, vols 61, p. 53, pl. 4, figs. 5, 6.
Cancellaria epistomifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. ‘te, figs. 3,4; 5.
For figures of this species, the reader is referred to Dr. Maury’s Dominican Fossils. We have collected a few, small and imperfect specimens at Gatun in the Canal Zone. Small shells are easily confused with the common C. dariena, but in gen- eral the spire is shorter, and the spiral sculpture is heavier and more regular. At maturity, the outer lip develops below a peculiar and characteristic sulcus or pout, which is not seen on any of the associated species of Cancellarta.
Gatun Stage: Gatun, C. Z.
84 BULLETIN 39 256
Cancellaria Rowelli Dall Plate 6, figure 7
Cancellaria Rowelli Dall, 1898, Proc. Acad. Nat. Sci. Phila., vol. 19, PP) 307, ply 20,25. Cancellaria Rowelli Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. 16; figs 2 Two specimens of a Cazcellaria which seem to be this spec- ies, were collected from the Gatun beds of East Grape Point Creek. The axial ribs are numerous, oblique and somewhat ir- | regular. The spaces between the ribs are finely spiralled with deep, incised lines, producing narrow and regular spiral bands. The columellar plications are large and more or less bifid. Shell measures: Length 30, diameter 18, last whorl 21 mm.
The Cornell Museum contains, in the Gabb collection, a single specimen which is probably this species. It agrees with the Costa Rican shells. This species was described by Dall from the Miocene of the Rio Amina, Santo Domingo.
Gatun Stage: Collection 2, East Grape Creek, C. R.
Subgenus TRIGONOSTGMA Blainville Cancelfaria toroensis, n. sp. Plate 6, figure 4
Cancellaria (Trigonostoma) aff. Cancellaria bullata Sowerby, Toula, 1911, Jahrb. der K-K. Geol. Reichsanstalt, vol. 61, p. 504, pl. 30, fH F2' TO:
Shell with 6 large, expanding whorls; the nucleus of 2% whorls is small, closely coiled and smooth; the succeeding 2 post-nuclear whorls are small, convex and finely reticulated with 6 spirals and fine ribs; the last two whorls are deeply channeled or excavated along the suture; this excavated band is smooth, except as crossed by oblique growth lines; the last whorl is somewhat contracted above and between this contracted zone and the sutural channel is a large, heavy, more or less bifid spirai cord. ‘This spiral cord is rendered coarsely and sharply nodulose by the axial sculpture; the sculpture consists of about 21, narrow, low, oblique riblets crossed by strong spiral cords; the spirals are
257 Costa Rica MIOCENE --OLSSON 85
rendered slightly nodulose by the crossing of the longit udinal ribs the penultimate whorl shows 6 or 7 spirals and 14 or 15 on the last whorl; the contracted band around the upper part of the whorl is nearly free from spirals, but is bordered above as already not- ed by the large, bifid and strongly nodulose spiral cord; base deeply but narrowly umbilicate, spirally sculptured within; col- umella with 3, oblique plications; inner lip with a wide callus, finely pustulated; the outer lip is broken in the type.
Length 29, diameter 20, last whorl 23 mm.
This is the Miocene analogue of the recent West Coast C. tuberculosa Sowerby. It is probably the species figured by ‘Toula from the Canal Zone and which he compared with C. budlata Sowerby. More recently Pilsbry and Johnson have described as C. insularis,a Trigonostoma from the Miocene of Santo Dom- ingo and which they consider as possibly identical with Toula’s Isthmian specimen. In their description they mention but 14 ribs while /ovoenszs has 21 and they relate their species with the Chesapeake Miocene perspecttva Conrad and the Tampa depressa Dall, species which belong to an entirely different group.
Gatun Stage:
Toro Cays, Providence of Bocas del Toro, Panama.
Canceliaria Plummeri, n. sp. Plate 6, figures 2, 3
Shell thin and delicate, with large, loosely coiled whorls; the whorls are prominently shouldered and deeply channeled or excavated along their upper sutures; nucleus of three small, smooth whorls; post-nuclear whorls 4; the sculpture consists of very fine and delicate spiral threads or lines and three rows of slightly elevated spine-like elevations or small tubercles; the larger of these rows is found along the shoulder of the whorls bordering the excavated zone, a smaller on the middle and a small scarcely noticeable one below; on the penultimate whorl, only two rows show and on the upper spire-whorls but one; the umbilicus is deep, but narrow; the interior is concealed in the matrix.
86 BULLETIN 39 258
Height 34, diameter 26, last whorl 17, aperture 19 mm.
This isa very distinctive species, characterized by its thin, delicate shell and fine sculpture. It may be compared with the recent 7. dullata Sowerby from the Pacific coast of Central America, which differs in its larger, more expansive body-whorl and coarser sculpture. The single specimen was collected from the lower part of the Gatun formation on Toro Cays, in the east- ern part of the Chiriqui Lagoon.
It is named for Mr. Fred Plummer of the Royal Dutch Oil Company of Hague, Holland.
Gatun Stage: Toro Cays.
Subgenus APHERA H. and A. Adams
Cancellaria islacolonis Maury Plate 6, figure 12
Cancellaria tessellata Gabb, 1873, Trans. Amer. Phil. Soc. vol. 15, p, 236. Not of Sowerby, 1832.
Cancellaria islacolonis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 229. plato whies <a ob:
The C. zslacolonis was selected by Dr. Maury, as the guide fossil for the lower of her Miocene formations, the Cercado. It is a very rare fossil in Costa Rica and we have collected it only along Red Cliff Creek, in the eastern part of the Republic. There it occurs in the upper part of the Uscari and in the lower part of the Gatun. .
It is closly related to the recent West Coast C. fesselata Sowerby.
Uscari tormation; Coll. 6, Red Cliff Creek. Gatun formation, Coll. 4, Red Cliff Creek.
= bre, ee ee
259 Costa Rica MIocENE—OLSSON 87
(B) SUPER-FAMILY RHACHIGLOSSA
Genus OLIVA Bruguiere
Olives are among the most common fossils in the Miocene becs of Panama and Costa Rica. ‘The various species, however show so few distinguishing characters, that their identification becomes at times, both difficult and uncertain. The following key has been prepared showing the characters which have been most relied upon for their separation. This key should be used in conjunction with the figures.
A. Aperture narrow and of about the same width along its whole length. Inner lip finely and regularly crenulated above.
B. Sutures deep or channelled, with the edge of the whorl projecting slightly above; spire of medium heteht, length 25 mm.
O, sayana var. tmmortua Pilsbry & Brown
BB. Sutures not channelled and with the upper edge of the whorl beveled or appressed.
C. Spire generally low, with concave profile and usually a projecting and globular nucleus. Length 35 mm. O. brevispira Gabb CC. Spire high or short, with conic sides; nu- cleus or protoconch not large or prominent. DP Spire hich, and? comic: shell large: Length 40 mm or more O. cylindrica Sowerby DD. Spire short, shell usually small. Length rarely over 30 mm. O. gatunensts Toula AA. Aperture wide and usually expanded and flaring in front; inner lip is not crenulated above. (Agaronza) B. Spire long, about 1-3 of the length of the shell;
88 BULLETIN 39 aor
inner lip not strongly calloused. Lenght up to 45 mm. O. mancinella, n. sp.
BB. Spire shorter, about 1-4 of the length of the shell; inner lip usually strongly callused. Length about 40 mm.
O. testacea var. costaricensts, n. var.
Oliva cylindrica Sowerby Plate 7, figure 1
Oliva cylindrica Sowerby, 1849, Quart Jour. Geol. Soc. Londou, vol. 6) pae4 5.
Oliva reticularis Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22),\p. 288... Notiot Lamarck.
Oliva cylindrica Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
215.
Oliva cylindrica Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 22.520:
Oliva cylindrica Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6) Papl5o3:
Oliva ovlindrica Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p. 57 lisse Se woe au
Oliva cylindrica Maury, Bull. Amer. Pal., vol. 5; p:\231, ple te,iee. I4, 14a.
A single, large specimen from Costa Rica, is doubtfully re- ferred to this Dominican Miocene fossil. It differs from the more common dorevispiva by its larger size and longer, more conic spire. This shell is also closely related to the recent O. avaneosa Tamarck of the West Coast.
Length 47, diameter 19 mm.
Gatun Stage: Upper Gatun beds, 1 mile south of shore, alony Old Man Sam Creek, C. R.
Oliva brevispira Gabb Plate 7, figures 2, 3, 4
Oliva brevispira Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 215.
Oliva brevispira Maury, 1917, Bull. Amer. Pal., vol. 5 p. 232, pl. fo, HS. LOM.
26% Costa Rica M1ockENE—OLSSON 89
variety Givaudi Cossmann
Oliva Giraudi Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p. 56, pl. 5, figs. 4-8.
A fairly common species in Cost Rica. Typically the shell is short; moderatly convex, a short projecting spire and concave spire-whorls.
Unusually broad forms as illustrated by figure 4 may be sep- arated by the name of Gzvaudz Cossmann. A large typical shell measures: Length 35, diameter 17 mm.
Gatun Stage: Coll.6, Red Clif Creek, C. R. fleadwaters of Middle Creek, C. R. Coll. 5, “Grape Point, Creck, Ci LR.
Oliva gatunensis Toula Plate 7, figure 5
Oliva gatunensis Toula, 1909, Jahrb. der K-K. Geol. Reicesanstaldt, VOL) 53)\p:'7o2, pl..25, fig. 12:
Oliva reticularis gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. sel. Phila... vol. 63, p. 348.
Oliva gatunensis Cossmann, 1913, Jour. de Conchyliologie, vol. 61, p. BO, pl, 5,2 11S. 9-2.
A comimon species at Gatun. The largest specimen in our collection measures 28 mm. in length but Brown and Pilsbry men- tion shells of an length of 35-38 mm. Oliva gatunensis, resem- bles somewhat the bdrevzrsoira of Gabb, but has a more even con- ic spire and lacks the projecting nuclear tip. The sutures are appressed. Brown and Pilsbry, place this shell as a variety of the larger, recent Oliva reticularts Lamarck.
ean Stage: Mt, Flope, C.Z. Oliva sayana var. immortua Pilsbry and Brown Plate 7, figs. 6, 7
Oliva sayana var. immortua Pilsbry and Brown, 1917, Proc. Acad. Mat Sel. Phita., p33, pl: 5,)f2.' 6.
The Oliva sayana Ravenel (Oliva literata of most writers)
90 BULLETIN 39 262
is acommon recent species along the eastern coast of United
Rar: becoming replaced in the West Indies by the Oliva reti-
cularis Lamarck. ‘These two species are principally distinguish- ed ae each other, in that the sutures of O. sayana are quite deeply channelled, above which projects the upper edge of the whorls while in reticularis the sutures are merely deep with rounded or beveied shell margins.
O. sayana var. tmmortua was described by Pilsbry and Brown, from Gatun beds in the vicinity of Cartagena, Columbia. What appears te be the same shell is common in the exposures found along the lower part of the Banana River. Like the drev- istyia it is rather darkly colored, but no trace of a pattern is pre- served. Our specimens averaged in length about 36 mm. ‘The largest specimen measures 4I by 19.5 mm.
Gatun Stage: Banana River, C. R.
Oliva testacea Lamarck, var. costaricensis, n. var. Plate 7, figs. 12, 13
Sheil rather solid, with a moderately projecting spire, about
1-4 or less of the total length of the shell; whorls about 5, the last, large, convex and widest about the middle; the spire-whorls appear as if mesially divided by an encircling weak callus about the lower half; aperture broadest below, with the inner lip heav- ily calloused above near its junction with the outer; the inner lip is non-denticulated above, along its lower part with 3 or more, irregular oblique, plaits; a broad band, arises from about the mid- dle of the inner lip, passes obliquely downward over the back of the shell to the lower end of the outer lip.
Rength 42, diameter 19 11m,
This Oliva is abundant in the Gatun beds of the Banana River, C. R. It is closely related to the recent O. festacea Lam- arck but seems to differ in being wider, shorter and with a lower spire.
The Oliva testacea Lamarck (in part O. /“zatula of some au- thors) is abundant along the Pacific coast of Panama, and in the Manuals, its range is given as the West Coast. Recently a close-
a lt oe
263 Costa Rico MiocENE—OLSSON QI
ly related if not identical form, was collected by us from the Pleistocene at Almirante, Panama, and several broken specimens from the beaches at Bocas del Toro, Panama, and from Manzan- illa, C. R. This recent Atlantic form differs only from the typ- ical West Coast festacea in being somewhat more slender and ap- parently differently colored.
Tryon in the Manual of Conchology unites festacea with hiatula Gmelin from West Africa.
Gamn stage» Banana River, Coll. 5, 6, of Red Clif Creer,
Oliva mancinella, n. sp. Plate 7, figures 8, 9
Shell slender and more delicate than the preceding; spire long and pointed and about 1-3 of the total length of the shell; 5 whorls; sutures deep; the spire-whorls have their lower half thickened by a smooth, encircling band of callus; aperture broad- est about the lower half; inner lip smooth above, with only a small and weak callus at its junction with the outer lip; the out- er lip carries below, 3 or 4 heavy and very oblique plaite; the outer lip 1s smooth and sharp.
Length 48, diameter 11 mm. This species differs from O. des‘acea and its variety costaricen-
sts by its much more slender and delicate shell. The porpor- tions of the spire are about 1-3 of the total length of the shell.
Corn stage: Coll. g, Last Grape Point, Creek, C.. R.
Genus OLIVELLA Swainson
A. Shelllarge, length 20 mm. or more; spire long and about ¥% the total length of the shell. Olivella goliath, un. sp. AA. Shell smaller; spire long or short, usually less than % of the total length. B. Shell short and stubby, spire rather short,
92 BULLETIN 39 204
bluntly pointed; 1-3 or less the total length. Olivella limonensts and variety bocas- ensts, n, Sp. audi vam
BB. Shell with a longer spire, oftensharply point- tol
C. Shell rather broad, with a heavy cal- lus about the upper part of the inner lip and extending partly over the pen- ultimate whorl. Length 12 mm. plus.
Olivella muticotdes Gabb.
CC. Shell slender, with a long, pointed spire; callus about the upper of the inner lip, slight.
Olivella Boussact Cossmann, var. Oliveila goliath, n. sp. Plate 7, figures 22, 23
Shell large, with a long, pointed spire of about % of the to- tal length of the shell; whorls about 6, with plain, nearly straight profile; sutures linear; last whorl with the greatest convexity about the middle; aperture subelliptical, broadest in front; in- ner lip with a thin, flat callus and with 3 small plaits below.
Length 26, diameter 10, last whorl 20, spire 13 mm 2a, 8.5 18 2 mm
An unusually large species, represented by two specimens from Red Cliff Creek and Banana River. Its general form is that of O. indevisa Guppy and O. Boussaci Cossmann, but nearly 4 times as large.
Gatun Stage: Banana River.
Coll) 5, Red Clafh\ Creeks) Cie Olivella muticcides Gabb Plate 7, figures II, 14, 17, 18
Oliva muticoides Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 215.
Olivella mutica variety muticoides Dail, 1890, Tarns. Wagner Free, Inst. Sci., (vol. 3, pt. Lp 45:
265 Costa RrcA MIoCENE—OLSSON 93
Olivella muticoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 232, pl. 1 an chee
M
A. broad, chubby species with a spire of moderate length. A large, thick callus is formed about the upper part of the inner lip, which spreads partly over the back of the penultimate whorl, giving to it a hump-back appearance. Measurements of our Costa Rican shell run as follows:
Length 17, diameter 7, last whorl 14, spire 6.5 mm. 13 6.5 ri 5-5 mm. Gatun Stage: Coll. 5, Red Cliff Creek. Coll. g, East Grape Point Creek.
Olivella Boussaci Cossmann, variety Plate 7, figures 15, 16
Olivella Boussaci Cossmann, 1913, Journ. de Conchyliologie, vol. 61, p. 60, pl. 5, figs. 16-19.
The Olivella indivisa Guppy (Proc. U.S. Nat. Mus., vol. 19, p. 308, pl. 30, fig. 10) from Jamaica and Olivella Boussact Cossmann from Martinique, belong to a closely related series characterized by their high and sharply pointed spire. The main difference seems to be that of size. The zzdivisa hasa length of 6.5 mm., the Boussact of 9g mm. The Costa Rican shells are generally larger and when full-grown, often 13 or more mm. in length.
Often very common. Length 13, diameter 5, last whorl 10, spire 6.5 mm (6 whorls)
Gatun Stage: Hill ra, Banana River. flill 3, Banana Rtver.
Olivelia limonensis, un. sp. Plate 7, figures 19, 20
Shell short and plump, witha small, broad, conic spire; whorls about 5, separated by deep sutures; last whorl very large, broadly cylindrical in form and slightly contracted in the middle; the inner lip has a wide callus, somewhat heavier above, with 5
94 BULLETIN 39 266
fine, denticles in the middle and few, more oblique plaits below;
the base is obliquely encircled by a lighter colored band; outer lip sharp, smooth within.
Length 10.5, diameter 4.75, last whorl 9, spire 2.25 mm. This is acommon shell in the coralline phase of the Gatun at Port Limon. The middle of the last whorl is usually dark col- ored, with the spire and the encircling basal band and callus on the inner lip white.
Gatun Stage: Port Limon. Olivella limonensis var.bocasensis, n. var. Plate 7, figures 24, 25
A much smaller shell than the /émoncnsis and possibly a dis- tinct species. The spire is somewhat higher and the shell less cylindrical and pump. Dimensions as follows:
Length 6.5, diameter 2.75, last whorl 6, spire 1.5 mm. ls 35253 6.5 1.75 mm. Gatun Stage: Bocas del Toro, Panama.
Genus ANCILLARIA Lamarck
Ancillaria aquaensis, n. sp. Plate 7, figure Io
Shell of medium size, solid; spire elevated, composed of about 5 whorls, the sutures of which are concealed by a wide band of callus this band of callus commences on the body-whorl, a short distance below the suture, and extends to the apex; where this band covers the sutures, it forms just above, a depressed or constricted band on the middle of each whorl; the last whorl is large, with a basal band of callus, which commences near the upper end of the inner lip, descends obliquely across the back of the last whorl to the anterior tips of the outer and inner lips; a short distance above this basal band, the shell carries a single, incised line; a small umbilical pit is found just behind the mid- dle part of the inner lip; aperture broadly sub-elliptical with a sharp outer lip.
Length 18.5, diameter 7.5, last whorl 12.5, spire 10 mm.
267 Costa RicA M1ocENE—OLSSON 95
The Ancillaria pinguis Guppy, from Jamaica is probably a closely related species. Guppy’s figure in the Geological Maga- zine, vol. 1, Decade 2, isvery poor and insufficient. Our shell seems to differ by its much longer spire and more slender shell.
The Ancillaria chipolana Dall, figured on Plate 41, fig. 3, of the Wagner Institute, is larger and has a longer spire. No um- bilical pit is shown in the figure.
Gatun Stage: Water Cay, (Isla de Aqua).
Genus MARGINELLA Lamarck Marginella MacDonaldi Dal! Plate 6, figures 14, I5, I7, 19
Marginella MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No. 2, p. 7.
This large MWarginella is one of the most common and char- acteristic species of the Gatun beds of Costa Rica. It is extreme- ly valiable in size and general from and heaviness of its shell, as may be seen in the accompanying figures of the more common varieties.
Typically the shell is oblong-cylindrical with a small spire of about 4 whorls, usually completely covered in front by a broad mass of callus, which spreads over the whole base of the shell and the outer lip, leaving an elevated ridge along each side as frequently seen in Cypr@a. From the back, the spire may be seen lying in the mass of callus.
The Marginella mindiensis Cossmann is a smaller related species from the Canal Zone. It differs also in having its outer lip finely denticulated, while in MacDonaldi the outer lip is us- ually smooth.
Length 28, diameter of base 15, vertical diameter Io mm.
24 14 g mm. 25 12 9.5 mm. L705 II 7.5mm.
Gatun Stage: Banana River.
96 BULLETIN 39 268
Marginella mindiensis Cossmann Plate 6, figures 16, 23
Marginella mindiensis Cossmann, 1913, Journ. de Conchyliologie, vol. 6L, p. 61, 7pl) 5 esa n5.
This shell is very closely related to WM. MacDonaildi. It dif- fers only in being smaller and usually with the outer lip more or less denticulated. The outer lip of WZ. MacDonaldi is generally smooth but occasional specimens occur in which the outer lip is faintly denticulated. It isa species of the Canal Zone.
Length 19.5; basal diameter 11, vertical diameter 8 mm. 16 9 Fpeonnn tl Gatun Stage: Mindi, (Cossmann) GAL OOS,
Marginella latissima Dall Marginella latissima Dail, 1896, Prac. U. S. Nat. Mus., vol. 19, p. 308 pl 2osnesur. This is a short and very broad species, described by Dall from Moen Hill, C. R. Dall remarks:‘‘This is perhaps the short- est and widest American species.”’
b)
Length 11, diameter 8.5 mm. Gatun Stage: Moen, Costa Rica, (Gabo).
Marginella latissima, var. pilsbryi, n. var. Plate 10, figures 1, 2
Marginella coniformis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 348, pl. 24, fig. 12. Not of Sowerby, 1849.
Like JZ. latisstma Dall, but less broad, more pointed anteri- orly, a slightly higher spire and a less heavy outer lip. In /afzssz- ma, the two posterior plications are shown as being very oblique, while in the present form they are nearly transverse as is seen in the recent cézcta. ‘The outer lip is finely granulated.
Length 14, basal diameter 9.5, vertical diameter 6.75 mim. Us II g mm.
This shell is figured asthe Marginella contformis Sowerby a common Dominican fossil, by Brown and Pilsbry. The Marg- |
269 Costa Rica M1ocENE—OLSSON 97
wnella coniformis has been figured by Guppy in the Quarterly Journal and similiar shells were collected in abundance by the Maury expedition to Santo Domingo. It isa larger and more cylindrical form.
It isan abundant shell in the quarries west of the Gatun locks at Gatun. A single, large shell figured as figure 1 was collected from the Banana River beds.
Gatun’ Stage:-Gatun, C.: Z. Banana River, C. R.
Marginella avena Valenciennes Plate 7, figures, 21, 28
Marginella avena Valenciennes, 1814, in Kiener, Coq. Viv. Marginella, Dura pl. 6, tes 24,
Marginella avena Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, and series, p. 355.
A common, recent species along the Caribbean coast of Panama and Costa Rica. Asa fossil, it occurs plentifully in the coralline phase of the Gatun at Port Limon and Bocas del Toro. The fossil shells do not differ materially from their recent repre- sentative.
The shell is elongate-cylindrical in outline, with a low, but slightly projecting spire. The aperture is norrowly linear, slightly wider in front. Outer lip thick, smooth within. The columella is provided with 4, obliquely descending plications.
A. series of specimens from Limon and Bocas measure as fol- lows. Length 11.75, diameter 4.25, spire 2. II.50 4.25 1.5, 12.50 4.50 2) Gatun Stage: Port Limon, C. R. Bocas del Toro, Panama.
Gou, 4, Red Clif Greek.
Marginalla collina, n. sp. Plate 7, figures 26, 27
Shell of the general form and size of A/. avena; but more
98 BULLETIN 39 270
solid and with a lower and less differentiated spire; whorls 4 or more with sutures scarcely distinguishable under their glaze or coat of callus; aperture linear-elongate, widest in front and slightly contracted in the middle; outer lip thickened, smooth within; pillar with 4, oblique plaits of nearly equal strength.
Length 11, diameter 4.5, spire 1.5 mm. 10 4.25 I mm, Lie25 ey I mim.
Closely related to the Marginella avena Val., this species differs in being more solid, less slender and with a lower spire. In avena the spire is clearly differentiated, with well marked su- tures to its spire-whorls. In the present shell, the spire is much lower, and the sutures are more thickly covered with glaze. The thick outer lip is carried across the end of the lip and firmly joined to the spire without any preciptible line of demarcation. In avena, the outer lip and spire are clearly differentiated from each other.
Abundant in the Gatun beds of the Banana River. Gatun Stage: fiill ra, Banana River.
Marginella leander Brown and Pilsbry Plate 6, figure 22
Marginella leander Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila vole 62.02.3247, pla 24y He is
Not known from Costa Rica, and the figure here given is from a specimen in our collection from Gatun. It is a broader and more cylindrical species than colliza, and with a much low- er and scarcely elevated spire. Our specimen measures 9 by 4 mim.
Gatun Stage: Gatun, C.OR.
Marginella musacina, n. sp. Plate 10, figure 25
Shell small and slender, with a high spire of about 1-4 of the total length of the shell; whorls about 5, with sutures light- ly concealed under a thin coat of glaze; last whorl narrowly cylin-
271 Costa Rico MrockENE—OLSSON 99
drical, widest about the upper half; aperture sublinear, with a
slightly thickened lip, somewhat contracted in the middle, smooth
within; inner lip smooth, or with only a thin wash of callus and
with 4, very oblique plications at its extreme anterior end. Weneth 9.25, diameter 3.25,-Spire 2:75 mm.
A small, slender species of unusual form. ‘Two specimens were collected in Costa Rica, the type from the Banana River, the other from Old Man Sam Creek, near Manzanilla Point.
Gatun Stage: Banana River,
Along Old Man Sam Creek, one mule south of the beach, C. R.
Genus VOLUTA Linnzeus Voluta alfaroi Dall Plate 8, figure 2
Voluta alfarot 1912, Smith, Misc. Coll., vol. 59, No. 2, p. 8.
This fine species described by Dr. Dall, from the Banana River, isa common and very characteristic fossil or the Gatun beds in Costa Rica and Western Panama, but it still remains to be recorded from the Canal Zone. Allied to the recent West Indian V. musica Linneeus it differs most importantly in its much smaller nucleus.
The shell is heavy and in its typical form the whorls are shouldered and carry about 12 ribs, which may be quite sharp and high on the shoulder angle. The whorls are sometimes smooth, without ribs and a shoulded angle. Spiral threads usu- ally occur on the lower one-quarter of the last whorl and on the canal, but in some cases on the spire-whorls as well. A large specimen from Water Cay measures;
Height 59 mm. diameter 34 mm.
Gaiun Stage: Water Cay Panama. Coll. 3, Red Cliff Creek; Old Man Sam Creek, E wile from shore; Goll... 2, Ourtana Créer; Comadre Creek, Soust creek; Coll. 5, Estrella River, Banana River, Rio Blanco;Port Limon.
100 ; BULLETIN 39 272
Genus SCAPHELLA Swainson Scaphella costaricana,n. sp. Plate 15, figure 12
Type fragmentary consisting of the nucleus and part of the three succeeding whorls; the shell is subfusiform; nucleus large, smooth and mamunilate at its apex; the three following whorls are slightly contracted about the upper sutures and strongly sculp- tured with coarse subregular, spiral threads separated by inter- spaces as wide or a little more; the spiral threads and their in- terspaces are crossed by fine lines of growth; sutures distinct; the columella with four plaits, the posterior one being the strong- CSU:
Length 29, diameter 18, diameter of nucleus 5.75 mm.
The unique type is unfortunately fragmentary and consists of the large, smooth, mammilate nucleus and part of the 3 suc- ceeding whorls. The shell is subfusiform and the whole surface is strongly sculptured with coarse, spiral threads. The columel- la is provided with 4 strong plaits. The type specimen was found in the Denialium zone of the Upper Uscari shales of Cocles Creek.
Uscari Stage: Cocles Creek.
Genus MITRA Lamarck Mitra Swainsoni Broderip, var. limonensis, n. var. Plate 6, fig. 1.
ef. Mitra Swainsoni Roderip, 1835, Proc. Zool. Soc., p. 193. cf. Mitra Swainsont Reeve,’ Conch. Icon.; A7ziva, pl. ee
ef. Wfitra Swainsoni, var. antillensis Dall, 1889, Bull. Mus. Comp. ZOOL Vols 18...) 158) pls) 38. tas 7s Shell large, of about 6+ whorls; the spire whorls are slight- ly convex, the last distinctly shouldered above, depressed or slightly concave about its middle; sculpture consisting on the spire-whorls of about 6, heavy, spirals cords with sculptured interspace as seen on Mittra Henekeni and longa; on the later whorls the spiral cords become more widely spaced and on the
273 Costa Rica MiIocENE—OLSSON Io!
last are in the form of broad, subobsolete, smooth bands without the sculptured interspaces; there are about 15 spiral cords on the last whorl in addition to those of the anterior canal; anterior canal of moderate length, strongly twisted; columella with 4 plaits, heaviest above; aperture narrow.
Length 74 (apex broken), diameter 23, last whorl 52, spire 35 mim.
A single, large shell with rudely sculptured whorls was col- lected from the coralline limestone near Port Limon. It agrees closely with Reeve’s figure of Af%itra Swainsont, except that the spiral bands are larger and heavier.
The Afitra Swainsonz is a West Coast shell belonging to the Panamic province. It is distinguished by its large size, rudely sculptured whorls and dark colored epidermis. Dall has des- cribed as the variety amzfillensis, specimens dredged off of Cape Lookout, N. C., Colombia and Yucatan.
Gatun Stage: Port Limon. Mitra longa Gabb Plate 6, figure Io
Mitra longa Gabb, 1873. Trans. Amer. Phil. Soc., vol. £5, p. 219. Mitra longa Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. Ganipe 446, pl: 24, fig. V1. Witra longa Maury, 1917, Bull. Amer. Pal., vol: 5, p. 238; pl. 11, figs. Tn i wea A long, slender species, sculptured with strong, revolving cords and fine, neatly engraved interspaces. We have collected this Dominican species only inthe Canal Zone. The figured spec- imen has the following dimensions:
Length 40, diameter 10.5, last whorl 26, spire 2I mm. Gatun Stage: Gatun, C. Z.
Mitra dariensis Brown ond Pilsbry Plate 6, figure 25
Mitra dariensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., OLIGO, 0. /340;))pl. 24, fos TL.
102 BULLETIN 39 274.
A single specimen from the Island of Bocas (Columbus or Colon Island) is here figured. It differs from the Mitra longa, in being shorter and broader. ‘The sculpture is less elegant.
Length 21, diameter 7, last whorl 14, spire 11 mm.
Gatun (Stage “Gatun Gan A, Bocas del Toro, Panama,
Mitra aff. rudis Gabb Plate 6, figure 13
Mitra rudis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 220.
Shell subfusiform, biconic; whorls about 7, coarsely sculp- tured with strong, revolving cords, separated by interspaces of about twice their width; These interspaces are finely longitudi- nally sculptured with raised threads above, obsoletely so below; the spire-whorls have 5 cords, the last whorl with 11 and several smaller ones on the canal; aperture subelliptical, with a thin out- er lip; columella with 4 plaits, largest above; anterior canal of medium length, strongly twisted below.
Length 33.5, diameter 12.5, last whorl 24, spire 16 mm.
It is with much uncertainty, that I have identified this rare species with Gabb’s unfigured Wtra rudis from Santo Domingo. Gabb,s descriptions and measurements indicate a shell of much the same characters as well as dimensions.
Gatun Stage: Coll. 3, fone Walk Creek) Ck: Mitra poas, n. sp. Plate 6, figures 20, 21
Shell small, nearly smooth and columbelloid in appearance; whorls 8, with straight sides; last whorl widest just above the base, which is contracted to the short anterior canal; the early spire-whorls have 5, low, smoothish spiral bands, with longitudi- nally sculptured, narrow interspaces; these spirals quickly be- come obsolete, leaving the whorls smooth except for one or two spirals bordering the upper sutures; the last whorl has about 4 spirals around the base and smaller ones on the canal; aperture subellipical, with a thin outer lip; columella with 4 plaits, larg-
275 Costa RicA MIocENE—OLSSON 103
est above; anterior canal of medium length and slightly bent to the left
Length 2:1, diameter 7.5, last whorl 13, spire II mm.
A smooth species of Columbelloid aspect. It is fairly abund- ant in the coral limestones near Port Limon.
Gatun Stage: Port Limon,
Mitra Almagrensis Toula var. coralliophila, n. var. Plate 6, figures 18, 24
of Mitra Alagrensts Toula, 1911, Jahrb, der K-K Geol. Reichsanstalt, vol; 62) p. 491, pk: 28, fig 12.
Shell small with a long spire and a shorter anterior canal; whorls about 8, with straight or slightly convex profile; sculp- ture of low, slightly elevated spiral cords, separated by inter- spaces of about their own width; these interspaces are finely sculptured by regular, raised, longitudinal threads; the spire- whorls have about 5 spiral cords, the last whorl with about 15; and smaller ones on the anterior canal; aperture subelliptical; a short canal, slightly bent to the left; pillar with 3 folds anda very small faint one below.
Length 17, diameter 6, last whorl 11, spire io mm.
This is a common species in the coral limestones of Limon. Toula’s Mitra Almagrensis was described from Tehuantepec. Our shells differ from Toula’s figure in being more slender and in having the body-whorl less contracted below.
Gatun Stage: Port Limon.
Genus FUSUS Lamarck Fuscs miocosmius, n. sp. Plate 8, figure 5
Shell long and slender, with the spire and canal of nearly equal length; whorls about 11 plus, very gradually tapering from the small nucleus to the body-whorl; whorls convex with indis- tinct, appressed sutures; the sculpture consists of large, swollen ribs, separated by equally wide interspaces, and the whole
104 BULLETIN 39 276
crossed by strong spirals with finer threads in between; on the Jast whorl there are 7 ribs, which commence on the base, just above its union with the long canal, and continue across the whorl to the suture; the early spire-whorls have 6 or 7 strong spirals with 1, 2 or 3, fine threads between; on the penultimate whorl, the earlier ‘spirals have become a primary set of 6 or 7, with their intervals occupied by a secondary set, nearly as large as the primary, and still smaller tertiary threads; the last whorl has about 18 strong spirals in addition to those on the canal; on the canal the division into primary and secondary spirals is bet- ter shown, there being about 16 primary and the same number of secondary spirals; on the extreme tip of the canal the spirals are very small and numerous; aperture small, rounded, with the out- er lip internally lirated and three or more oblique plaits on the columella.
Height roo, diameter 24 mm.
This fine species is closely related to the recent Fusus eucos- mutus Dall, from the Florida coast and the West Indies. The spire of ezcosmzus is more slender, has 8 ribs instead of 7, and the out- er lip is always smooth, while it is strongly lirated in mZzocos- Mtus.
Gatun Stage: Bocas del Toro, Panama.
Fusus honensis, n. sp. ' Plate 8, figure 6
Shell resembling in general features the /. mzocosmius, and with the same number of ribs; the spire is somewhat longer than the anterior canal; the whorls are somewhat more convex than those of mzocosmius due to the more strongly knobbed ribs; the ribs are crossed on the penultimate whorl by about 6, strong,even spirals, there being no secondaries and only occasionally a small thread may appear in their intervals; the last whorl, exclusive of the canalshows 10 or 11 spirals to which are added 17 or 18 on the canal; the canal is relatively short and not perfectly straight; the aperture is probably rounded (largely broken on the type specimen) and with its outer lip internally lirated.
Height 63, diameter 20 mm.
277 Costa Rica M1rocENE—OLSSON 105
Similiar in general form and in the number of its ribs to the fF, miccosmius from Bocas, this species differs in its smaller, heavier shell, and in the greater coarseness of its sculpture. The spirals are heavier and primary in character throughout and the ribs are more knobbed and sharp on their crests. From the Do- minican /. Henekent Sowerby, it differs in being much more slender and with fewer, persistent ribs.
Gatun Stage: Coll. 4, Home Walk Creek.
Genus FASCIOLARIA Lamarck Fasciolaria Gorgasiana Brown and Pilsbry Plate 8, figure 9
Fasciolaria gorgasiana Brown and Pilsbry, 1912, Proc. Acad. Nat. ci. Philia,, vol. 64,"p. 506; pl. 22;fg. 5.
This large Fasciolaria is fairly abundant in the Gatun beds of the Canal Zone and is recognized by its shouldered whorls and and strong knob-like ribs. The figured specimen is a large shell from the Gatun of Rio Betey, Costa Rica.
Gatun Stage. Gatun, C. Z. Water Cay. Rio Betey.
Fasciolaria tulipa Linnzeus, variety Plate 13, figure 4
Murex tulipa Linnzeus, 1758, Syst. Nat.,ed. 10, p. 754.
Fasciolaria tulipa Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 354.
Fasciolaria tulipa Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 3, pul, Ps LOI, pli 7 fo. 1k
The Tulip shell is the common, recent /asctolaria, in the Caribbean area, and it is distinguished from its more northern re- lative, the dzstans, by its darker coloration and sulcated sutural. band. It occurs as fossil in the Pliocene of Florida.
From the coralline limestones, near Port Limon we collected three young specimens and a portion ofthe spire of a large in- dividual of Fasciolaria. ‘The young shells show the nucleus and
106 BULLETIN 39 : 278
early sulcated whorls of typical fulipa, as fignred by Dr. Dall. They however become smooth somewhat earlier and the fragment of the spire shows the succeeding whorls to be smooth and lack- ing the sulcated sutural band as in dsfams,
The Fasctolaria semistriata Sowerby is an allied but distinct species from the Miocene of Santo Domingo, and likely to be dis- covered in the Gatun beds of Costa Rica and Panama. ‘This as may be seen from the new figure of Dr. Maury, (Bull. Amer. Pal., vol. 5, p. 244, pl. 13, fig. 1), differs conspicuously from the distans and tulipa, with which it was united by Gabb, by its deep sutural depressed band, which gives to the whorls, a very convex or even shouldered appearance. The early whorls are spirally sulcated as in ¢w/ifa, but have in addition knob-like ribs, like those seen on /. gorgasiana.
Uscarit Stage: Port Limon. Fasciolaria MacDonaldi, n. sp. Plate 8, figure 1
Shell large, (type specimen imperfect, with only the last two whorls preserved); number of whorls unknown; the spire-whorls are angled about the middle, forming a board, sloping shoulder above; the last whorl has the shoulder about the upper third; the sculpture is predominantly spiral, consisting of numerous, fine, alternating threads, which are slightly roughened by the growth lines; the angle of each whorl is longitudinally undulated by 12 faint ribs; anterior canal is nearly straight with two faint plice; outer lip sharp.
Length (2 whorls) 70, diameter 37 mm.
Of this large and elegant species, only a single imperfect specimen with less than two complete whorls preserved, was col- lected from the Gatun beds of the Banana River, Costa Rica. It differs from the more common /. Gorgasiana, by its persistent and rough spiral sculpture and less heavily tuberculated should- et.
This species is named in honor of Dr. D. F. MacDonald, well- known for his geologic work on the Isthmus, during the con-
279 Costa Rico MriocENE—OLSSON 107
struction of the Panana Canal, and former Chief Geologist of the geological force of the Sinclair Oil Corporation in Panana and Costa Rica.
Gatun Stage: Hill No. 2, Banana iver.
Genus LATIRUS Montfort
Latirus infundibulum Gmelin, variety Plate 8, figure Io
Latirus infundibulum Gmelin, Lamarck, Anim. sans, Vert. (ed, Desh. )
vol. 9, p: 386.
Latirus infundibulum Guppy, 1866, Quart. Journ. Geol. Soc. Loudon, vol, 22, p. 288.
Latirus infundibulum Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, pa 207:
Latirus infundibulum Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 354. Latirus infundibulum Maury, 1917, Bull. Amer. Pal., vol. 5, p. 246, pl. 13, ig. 3 A common recent species of the West Indian and Costa Rican coasts. Only an imperfent specimen was collected from the Gatun beds of lower Pumbri Creek, a small tributary of the Estrella River. From recent examples of zzfundibulum, it differs by its shorter canal, more numerous ribs (9 instead 6 or 7),and heavier spiral sculpture. The specimen is, however, to fragmentary to serve as a type for a new variety or related species.
Gatun Stage; Coll, 7, Pumbri Creck, C. R. Latirus irazu, n. sp. Plate 8, figure 12
Shell subfusiform; solid, with a long spire and shorter can- al; whorls about 9, with heavy, sharp, knob-like ribs and finer spiral threads; the last whorl shows 7, sharp, pointed ribs, which are continuous from suture to suture, but only feebly across the base of the last whorl; the sutureis bordered anteriorly by a prominent, frilled band or cord, formed by the elevated edges
108 BULLETIN 39 280
of the growth lines: spirals consisting of low, raised threads with wide interspaces; canal short, straight above but bent below, and with three small, smooth folds on the columella above; aperture with a thin outer lip.
Length 46, diameter 17, last whorl 29, spire 24 mm.
A smaller and more delicate species than the preceding ZL. zz- fundibulum Gmelin. In the present shell, the ribs are sharp- er and more pointed, and the canal is shorter and more strongly reflected backwards. The strongly frilled sutural band is lacking or only imperfectly developed on zxzfundibulum.
Gatun Stage: Port Limon. Latirus taurus, n. sp. Plate 8, figure 4
Shell large, slender, solid with the spire more than % the height of the shell; whorls convex, with indistinct sutures; sculp- ture of very regular, narrow ribs with deep interspaces, and cross- ed by strong, ridge-like spiral cords; whorls about 9; axial sculp- ture on last whorl with about 11 ribs; the ribs commence well down on the base and continue to the upper suture and are nearly in line acrossthe spire-whorls to the apex; the spirals consist of ridge-like cords and finer spiral threads; there are 2 principal spirals on the spire-whorls which cross the middle of the whorls; there are 4 principal spirals on the last whorl above the edge of the base, below which there are a few small- er threads; the anterior canal has 2 strong cords and finer threads are scattered over the whole shell; the anterior canal is straight and stocky and carries at its base a deep but narrow umbilical pit; the columella has 4 small folds; aperture subo- vate, anteriorly extended to form the long narrow canal; out- er lip smooth within.
Height 60, diameter 25, aperture 32 mm.
A large, solid and strongly sculptured shell. The straight, narrow ribs are spaced regularly over the whorls of the shell and are continuous from the base to the upper suture. They are
281 Costa RicA MIOCENE—OLSSON ete)
crossed by heavy, ridge-like, spiral cords, which on crossing the deep interspaces between the ribs, form small, sunken pits, so that the sculpture appears coarsely trellised.
Gatun Stage: Toro Cays.
Genus PTYCHOSALPINX Gill
Ptychosalpinx ? dentalis, n. sp. Plate 15, figures 14, 18
Shell buccinoid with convex, cancellated whorls; whorls about 6, with channelled sutures so that the whorls appear nar- rowly shouldered above; the sculpture is predominantly spiralled, the penultimate whorl with 7 strong spiral cords with interspaces 2 or 3 times their width; small spiral threads may appear in the interspaces; the last whorl has 14 or more spirals and the wide interspaces with one or more finer threads; the spirals are cross- ed by coarse wavy lines of growth, producing a sub-cancellate sculpture; the aperture is subovate; a well-developed siphonal sinus; the columella carries a single sharp plication at its lower end; the outer lip is slightly thickened and bears within about 12, small, narrow lire.
Height 28; diameter 11, aperture 16 mm.
The generic relation of this interesting species is very much in doubt. It has the general form of Comznel/a, but its colum- ella is provided with a strong anterior plication. In this regard it agrees with Ptychosalpinx of which there are several species in the Chesapeake Miocene. The apex of dentalts, appears to have been pointed and sharp, although all of our specimens have the tip of the spire very much weathered. True Ptychosalpinx is characterized by a large, convex and blunt nucleus and the outer lip is thin and smooth within. The outer lip of deztalzs is slight- ly thickened and internally finely lirated.
The P ? dentalts, is a very characteristic fossil of the Denta- lium zone of the Upper Uscari formation.
I1O BULLETIN 39 232
Uscari formation. Coco Plum, Panama. Rio Cocles. Comadre Creek, ete.
Genus PERISTERNIA Morch Peristernia insula, n. sp. Plate 8, figure I1
Shell small, with a sharp pointed spire, a little longer than the aperture; the general form and sculpture of the shell is like that of Uvosalpinx cinereus Say; nucleus of 2 small smooth whorls, followed by 7 post-nuclear; the profile of the spire-whorls is convex and strongly sculptured with ribs and sharp spiral cords; the last whorl has 8 ribs which are nearly lacking from the base of the whorl; the spiral sculpture consists of 2 principal cords about the middle of the earlier whorls above which lie smaller threads about the suture; on the later whorls, the spirals are somewhat heavier about the middle but irregular with finer, in- termediate threads in between the principal ones; base contract- ed; aperture subcircular; outer and inner lips crenulated or den- ticulated; canal short and bent.
Height 23, diameter 13, aperture 11 mm.
This and the following ¢ortugera seem correctly referable to the genus Peristernia Morch. The P. insula is somewhat like young specimens of /lzcata Conrad, from the Chesapeake Mio- cene of eastern United States, but has a longer and more pointed spire, and heavier sculpture.
Gatun Stage: Water Cay.
Peristernia tortugera, n. sp. Plate 8, figure 13
Shell elevated with a spire much longer than the aperture; nucleus of about 2% small, smooth whorls; the post-nuclear whorls about 7; sutures distinct; whorls strongly convex, should- ered; sculpture of narrow, heavy ribs, widely spaced and num- bering on the last whorl about 9g; the rib are continued across the whorls from suture to suture and for a short distance down on
hg
283 Costa R1cA MIOCENE— OLSSON a
the base; the spirals consists of irregular sharp cords; the early spire whorls carry 2 main spiral cords but bordered above and be- low by finer threads; the 2 principal spirals are continued on the later whorls but are nearly equalled in strength by the other spirals and hence lose their prominence; a short anterior canal; base strongly contracted.
Height 29, diameter 14.5, aperture 12 mm.
Distinguished from the preceding zzsw/a by its very much longer spire, more strongly contracted base and different spiral
sculpture.
Gatun Stage: Port Limon.
Genus XANCTUS Bolton Xancus scopulus, n. sp. Platei ii, ngure 7
Shell large, solid and heavy; spire nearly as long as the aper- ture; spire-whorls 6 plus (the tip broken), strongly coronate above and with large, wide, persistant ribs; the earlier spire- whorls are simply shouldered or angled about the middle, but the area about the upper sutures rapidly deepens and on the later whorls is a deep, excavated sutural zone, above which project the ends of the ribs; there is a strong sutural cord and a rather wide, ribbon-like band just above, forming at first a strongly ap- pressed suture; the last whorl has about 7 large, wide ribs, the areas between appearing as troughs or depressions; the sutural excavated zone carries several irregular spiral threads which are crossed by large growth-lines so that the resulting sculpture is more or less cancellate, more particularily on the earlier whorls; the growth-lines cross the sutural cord and upon the ribbon above, become much crowded and strongly bent forwards; the young shell. was sculptured over the whole shell with strong spirals, but with maturity, the spirals become obsolete and the shell is smooth and polished; columella with 3, strong plicz asin the recent scolymus; a long anterior canal, with a narrow, deep umbilicus behind the spreading inner lip.
Tine BULLETIN 39 asa
Height 265, diameter 136, aperture 158, last whorl 195 mm.
This very remarkable species is an extreme development of the X. scolymeus stock in which the upper portion of the whorl becomes a wide, deep, excavated sutural zone. Above this ex- cavated zone, project the high, rounded or appressed ribs and the carinate edge of the whorls. X. scolymus Gmelin, a recent spec- ice found plentifully along the north Panama coast, has the whorls simply shouldered, often merely rounded.
The Santo Domingan Miocene contains X. validus Sowerby, which has been identified by some with scolymus. In validus, the ribs are more numerous (about 10) and are sharper and more tub- ercular in form.
Gatun Stage: Banana River.
Genus MELONGENA Schumacher Melongena consors Sowerby Plate 9, figure I
Pyrula consors Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, Pp. 49.
Melongena melongena Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p- 205. Not of Linnzeus.
Pyrula melongena Guppy, 1874, Geol. Mag., vol. 11, p. 438.
Pyrula melongena Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, DP. 523.
Melongena consors Dall, 1900, Trans, Wagner Free Inst. Sci., vol. 3, pt Dips. 12)
Melongena consors Maury, Bull. Amer. Pal., vol. 5, p. 24y, pl. 14, fig. 5. The Melongena consors is not acommon fossil in Costa Rica. It is closely related to the recent JZ. corona Gmelin of the West Indies, the fossils shells differing mostly in having a longer spire and somewhat different sculpture above. The Costa Rican examples are exactly like Miocene speci- mens from Santo Doingo. As a fossil it occurs in the Miocene of Jamaica, Santo Domingo and Venezuela.
285 Costa Rica MIocENE—OLSSON 113
Gatun Stage: Coll. 5, Old Man Sam Creek. Cocles Creek.
Genus SOLENOSTEIRA Dall Solenosteira Dalli, Brown and Pilsbry Plate 8, figure 8
Solenosteira dalli, Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 348, pl. 24, fig. 14.
Coralliophila gatunensts Toula, 1911, Jahrb. der K-K Geol. Reichsan- stalt, vol. 61, p. 502, pl. 30, fig. 9.
A common species in the Canal Zone and figured here for comparision with the following species. The whorls are strong- ly shouldered, a feature accentuated by the few, but high,angled ribs. The spirals are heavy, raised cords, over and between which are finer secondary and tertiary threads.
Length 34, diameter 23, last whorl 27, spire 15 mm. Gatun Stage: Gatun, Mt. Hope, C. Z.
Solenosteira Vaughani Dall, var. medioamericana, n. var. Plate 8, fig. 7
cf. Solenosteira Vaughani Dall, 1903, Trans. Wagner Free Inst. Sci., VOL) Pt..O, p. 1633) pl. 6o, tig, 1S.
Shell solid, with a small pointed nucleus of 2 smooth whorls, and 6 post-nuclear; spire conic, of about % the total length of the shell; whorls convex, or bluntly shouldered about the mid- dle ; sculpture of numerous low ribs which are rounded, and heaviest on the shoulder angles; There are 7 to Io ribs on the last whorl; spirals of raised cords, with finer threads on the in- terspaces; the spire-whorls have about 6 primary spirals, the last whorl with 18, including those of the short canal; canal short with a deep but narrow umbilical pit; aperture oval, a thick in- ner and outer lip, the outer with about 10, long, entering liree and the inner with three small denticles above.
Length 42, diameter 24, last whorl 34, spire 17 mm. 34 oe Oe 28 13, mim.
The Solenosteira Vaughant is a Chesapeake Miocene species
I14 BULLETIN 39 286
from Jackson Bluff and Coe’s Mill, Florida. Dall’s figure inthe Wagner Free Institute, isso like our shell from western Panama and Costa Rica, that there can be but little doubt of their close affinities.
The present shell is larger than the S. Dall, of the Canal Zone, with more numerous, obtuse and not sharp ribs, and with a wider and deeper umbilical pit. A canal is developed at the posterior angle of the aperture as in the recent S. pallida Brod. while in S. Dalz, the posterior portion of the aperture is round- ed and there is no canal.
Gatun Stage: Toro and Water Cay, Panama. fiull 1a, Banana ‘River, Core
Solenosteira chiriguiensis, n. sp. Plate 8, figure 3
Shell large, heavy; spire about % the height of the shell; whorls about 7, strongly angled about the middle and carrying heavy knob-like ribs, crossed by heavy spiral cords; the last whorl has 8 ribs which are developed only on the middle of the shell, being lacking from the upper slope and from the base of the last whorl; the tops of the ribs are crossed by 2 strong,spiral cords with a wide, trough-like interval between; above the 2 principal spiral cords, there are 4 smaller cords on the upper slope and on the base and canal 10 or 11; base of the last whorl contracted to form the straight canal which carries a deep, narrow umbilius; aperture subelliptical.
Height 57, diameter 34, aperture 34 mm.
It is possible that this species belong to the genus Cymza rather than Solenosteira, but its aperture is so completely filled with a hard sandstone matrix that the presence or absence of a columellar fold cannot be determined. Its sculpture however is more like Solenosteira than Cymia. ‘The species will be recog- nized by its large size and characteristic sculpture.
Gatun Stage: Water Cay.
287 Costa Rico MIocENE—OLSSON 15
Genus METULA H. and A. Adams
Metula cancellata Gabb Plate 10, figure 12
Metula cancellata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, Pp. 205. Metula cancellata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 351. Metula cancellata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1584. Metula cancellata Maury, 1917, Bull. Amer. Pal., vol. 5, 249, pl. 14, fig. I9. Less common in Costa Rica and Panama than the following species and from which it is easily distinguished by its usual smaller size, more delicate shell, and fine sculpture.
The spiral threads on the last whorl number about 37 and are crossed by nearly as fine longitudinal ribs. The resulting re- ticulate sculpture is fine and neat. Our largest shell, a specimen from Bocas with 6 whorls measures:
Length 19, diameter 6.5, last whorl 14, spire 9, aperture IO mm.
Gatun Stage: Bocas del Toro, Panama, til 7a, Banana: River, Ce fe.
Metula Gabbi Brown and Pilsbry Plate 10, figure II
Metula Gabbit Brown and Pilsbry, tg911, Proc. Acad. Nat. Sci. Phila., WOL.62) p. 251, pl. 25, figs.4,.8.
This beautiful species is fairly common in the Canal Zone, but usually in a fragmentary condition. It related to the W/etula cancellata Gabb, differing mainly in its larger size and heavier sculpture. Ona speciman of 8 whorls from Gatun, the spirals on the spire whorl number about 8 and 39 on the last whorl and an- terior canal. The 2 upper spirals are noticeably stronger than the others. Aperture long and narrow and finely denticulated along the interior of the outer lip.
116 BULLETIN 39 288
The figured specimen from Gatun has the following meas- urements:
Length 29, diameter 10.5, last whorl 22, spire 12.5, aper- ture 16 mm.
Gatun Stage: Gatun and Mt.. Hope, C. Z. Metula Harrisi, n. sp. Plate ro, figure Io
Shell solid and more coarsely sculptured then MWetula Gabéz, and with a longer and broaded spire and shorter aperture; whorl 6 plus, (apex lost); sculpture of spiral cords and finer, curved longitudinal ribs; the spire whorls with 8 spiral cords which are finest next tothe lower suture, become progressively stronger above; on the last whorl and canal there are about 25 spirals; the spirals are slightly nodulated by the ribs of which there are about 4o on the last whorl; certain of the spiralson the middle of the whorl are double; aperture ovate-elliptical with a strongly thickened outer lip, internally with about 19 short lire; inner lip smooth; canal short and twisted.
Length 26.5, diameter 11, last whorl 19.5, aperture 13.5, spire 14 mm.
A solid and more coarsely sculptured species than the /Zetula Gabit. ‘The spire is broad and porportionately longer, due to the shorter aperture. In Gadi the spirals are of nearly equal strength over most of the shell surface with exception of those immediately bordering the upper suture. In the present species the spirals are finer below and progressively become stronger post- eriorly.
The type specimen was collected in the lower Gatun sand- stones of Coco Plum, Panana.
Gatun Stage: Coco Plum, Bocas del Toro, Panama. Metula Harrisi var. limonensis, n. sp. Plate 10, figures 5, 6
Shell more slender; a narrow and longer aperture more nearly the length of the spire; sculpture coarsely reticulate,
289 Costa RicA MIocENE—OLSSON Lay
with ro spirals on the penultimate whorl, about 26 on the last; aperture long, with a thickened outer lip, internally lirated in unison with the external spiral cords; inner lip smooth with a long, bent, beak-like canal.
Wenethy 27, diameter o.50, /last, wlorlii9, aperture 14, spire 13 mm.
Several specimens from the coral limestones of Port Limon. They are most closely related to Harrisz, in their heavy shell, coarse sculpture, and thickened outer lip. They differ in being more slender, a longer aperture and in minor details of their sculpture.
Gatun Stage: Port Limon, Genus PHOS Montford
Phos gatunensis Toula Plate 9, figures 4, 5
Phos gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt, Wein Vol38,(p. 701, pl. 28, fig 6? pl. 25, fig. 11.
Phos gatunensis Brown and Pilsbry, t9t1, Proc. Acad. Nat. Sci. Phila. Volk. 62, p) 240) pl 25. figs: 1, .2.
The Phos gatunensis is possibly the most common fossil in the Gatun beds of the Canal Zone and the adjacent sedimentary areas in the Province of Colon, but it still remains to be collected outside of this, its type area.
The shell is typically rather slender, with a sub-reticulate sculpture formed by the intersection of its numerous, straight longitudinal ribs, and the nearly equal, regular, strap-like, spiral - cords.
A typical specimen measures as follows: Length 30, diameter 13, last whorl 20, spire 15 mm. Cann stage: Gatun, C. 2.
Phos mexicanus Bose Plate 9, figures Io, I1
Phos mexicanus Bose, 1906, Bol. de Inst. Geol. de Mexican, numero 22,5030; plas oS, 18-27,
118 BULLETIN 39 290
Shell more broad and solid than gatunenszs of the Canal Zone; whorls about 8, those of the spire with nearly straight or only slightly convex sides; last whorl large, broad, convex and widest about the middle; the sculpture is much more coarsely re- ticulate than in gafunensts, with the intersection points of the ribs and spirals more or less subnodulose; the last whorl has about 18 ribs, which on the back of the outer lip are more close- ly spaced than elsewhere; the spiral cords are not so heavy as the ribs, separated by wider interspaces which carry one or some- times two very, fine, spiral threads; in addition, the spiral inter- spaces are very finely and neatly sculptured with fine, raised, longitudinal lines, corresponding to the growth-lines; on the spire-whorls there are 5 spiral cords, 13 on the last whorl and 3 more on the canal; canal short, twisted; aperture sub-ellip- tical, with a moderately heavy, but not thickened outer lip and internally lirated.
Length 30, diameter 15.5, last whorl 21, spire 14.5 mm.
A broader, more solid and more coarsely sculptured shell than the Phos gatunensis of the Canal Zone,
It was described from the Miocene of Paso Real, near Tux- tepec, in the State of Oaxaca, Mexico, by Dr. Bose, whose excel- lent figures agree exactly with specimens from Water Cay. The several species of Phos, are often extremely abundant but local in their distribution. Such is the case of Phos gatunensts of the Canal Zone, the Phos Mooret var. costaricenis of the Banana River. Likewise in the present instance; the mextcanus, 1s a very abundant fossil of the Gatun beds of the islands of the Chiriqui Lagoon and the adjacent Valiente Peninsula. A few specimens have also been collected in Costa Rica.
Gatun Stage: Water Cay. Toro Cay. Coco Plum, ete. Rito Cocles: (C.F.
Phos Moorei Guppy, var. costaricensis, n. var. Plate 9, figures 8, 9
Shell of medium size, solid; spire long about one-half the to- tal length of the shell; whorls 10, of which the first two belong
agt Costa Rica MIOCENE—-OLSSON 119
to a small smooth nucleus; succeeding whorls are sculptured with heavy persistent, straight ribs (13 or 14) on the last whorl; the ribs are crossed by even, raised, spiral cords; the spire-whorls have 4 spiral cords and a small one close to the upper suture; an- terior canal short and strongly twisted; aperture subelliptical, the outer lip with about Io, strong and regular internal lire.
Height 27, diameter 12, last whorl 17, spire 14 mm.
This is the common Phos of the Banana River. Although closely related to Moorez, from Jamaica, the Costa Rican fos- sils are smaller and more slender, as compared with Guppy’s figure in the Quarterly Journal, vol. 22, pl. 16, fig. 11.
Dr. Maury’s Phos Mooret, from the Miocene of Santo Do- mingo probably represents a distinct species, differing by its more slender spire, and in sculpture. The spiral intervals of costartcensis are smooth, while in the Santo Domingan shells, they carry fine spiral threads, which are best seen on the spire- whorls, sometimes becoming obsolete on the later.
Gatun Stage: Banana River, Old Man Sam Creek, 1 mile from shore or beach.
Phos elegans Gupp, var. limonensis, n. var. Plate 9, figures 12, 13
Shell usually smaller, more slender and delicate than the pre- ceding species; the spire is long, composed of about 9 whorls: the nucleus consists of 4 whorls, the first 2 are small, smooth and convex, the last 2, smooth, but sharply keeled about the periphery on the last 1-4 of the last nuclear whorl, faint curved lines appear above the peripheral keel, they gradually in- crease in strength and on the Ist post-nuclear whorl become the longitudinal ribs; the sculpture consists of fine, longitudinal ribs (23 or 24 on the last whorl); occasionally a rib may become greatly enlarged, forming varix-like thickenings corresponding to resting stages; the ribs are overrun by primary, secondary and tertiary spiral threads; the later spire whorls have about 6 spirals,
120 BULLETIN 39 292
with about 17 on the last whorl and the canal; aperture suboval, with a heavy but not thickened outer lip, internally lirated.
Length 27, diameter 11, last whorl 17, spire 4.5 mm.
This shell is related to the Phos elegans Guppy of the Mio- cene of Jamaica and Santo Domingo. Our shells differ from typical examples of edegans from Santo Domingo, in being larg- er, and more coarsely sculptured.
Varix-like thickenings occur scattered along the spire-whorls. The protoconch of typical elegans and /imonensts are practically identical. The recent West Coast Phos Veraquensis Hinds is a related but more finely sculptured species.
Gatun Stage: Port Limon. Bocas del Toro.
Phos estreilensis, n. sp. Plate 9, figures 17, 18
Shell small, slender, solid; spire much longer than the ap- erture, pointed; whorls of the spire 6 or more, convex in profile with deep sutures; sculpture of narrow, straight ribs which com- mence on the columellar area and pass across the whorl to the up- per suture; there are 12 or 13 ribs on the last whorl which are evenly and widely spaced except on the back of the outer lip where they are smaller and more numerous; the spirals consist of a primary set of threads or cords with a single, smaller second- ary thread in each interspace except on the base and the canal; the primary threads crossing the ribs form small, whitish tuber- cles; the spire-whorls have 4 primary spiralis and there are 11 on the last whorl, exclusively of the canal; aperture sub-elliptical, produced anteriorly to form the straight anterior canal; outer lip thickened and internally lirated.
Height 20, diameter 8, aperture 9 mm.
A small, slender species, coarsely sculptured like the larger and broader Phos Mooret variety costaricensts. It is limited in our collections to the Gatun beds of the Hstrella River.
Gainu Stage: Coll. 6, 7, Estrella River.
293 Costa Rica M1ocENE—OLSSON 123
Phos subsemicostatus Brown and Pilsbry Plate 9, figure 15
Phos subsemicostatus Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila; vol. 63, p. 350;-pl.25, fig. 3. A larger and finer sculptured species than the Dominican Phos semtcostatus of Gabb. A single imperfect specimen was col- lected from Zone E of Saury Creek.
Gatun Stage: Gatun (Brown and Pilsbry) one b, .saury Creek. (G.I.
Phos Veatchi, n. sp. Plate 9, figures 2, 3
Shell large and heavy, with a moderately long, coarse spire of about 9 whorls; nucleus small, of 2 smooth whorls; whorls of the spire moderately convex, with strong ribs continuous from suture to suture; sutures appressed and bordered below by a small spiral; the last whorl is widest about the middle and con- tracted below to the short anterior canal; sculpture of heavy, persistent ribs and regular raised spirals; the last whorl has 8 ribs; the spirals consists of raised threads, separated by wide, in- terspaces; an occasional secondary spiral will appear in the in- terspace but they are generally smooth or only obsoletely sculp- tured by the growth lines; there are 8 spirals on the spire-whorls, 16 or 17 on the last whorl; aperture ovate, with the outer lip thickened by the last rib, and internally with 9, long, entering liree; inner lip formed by a raised, spreading callus, armed above by a tooth-like denticle and a strong fold below on the edge of the canal.
Length 43, diameter 22, last whorl 28, spire 25 mm.
A large, robust species from Hill 1a of the Banana River. It differs from the sudbsemicostata by its strong persistent ribs and coarser spirals.
It is named for Dr. A. C. Veatch, Director of the Explora- tion work of the Sinclair Oil Corporation, under whose direction the Costa Rican collections were made.
122 BULLETIN 39 294
Gatun Stage: Hill ra Banana River. Phos beteyensis, n. sp. Plate 9, figure 6
Shell of the type of saésemicostata; whorls 3, moderately convex; sculpture with the axial element very weak but present on the penultimate whorl as fine, sub-obsolete, narrow ribs or ir- regular wrinkles on the shell surface; they are absent from the last whorl; the spiral sculpture consists of a broad zone, about the upper 1-3 of the spire whorl on which are irregularly distri- buted about 5 raised spiral threads; below this zone, the sculp- ture is largely in the formof broad uneven bands formed between incised spiral lines; on the last last whorl, this later zone occup- pies the middle of the whorl; the zone of raised spirals above the more smooth area of banded spirals gives to the whorls a slightly coronated appearance.
Length 27 (2% whorls), diameter 15, last whorl 21 mm.
The type specimen is very incomplete, but its characters are so distinctive and unlike any other species of Phos which we have seen that its discription is advisable. Like the sabsemztcost- atus the longitudinal ribs or coste fade out on the later whorls.
‘Gatun Stage; Betey Creek, C. R.
Phos metuloides Dall Plate 9, figure 16 Phos metuloides Dall, 1896, Proc. U. S. Nat. Mus., vol. 19, p. 310, pl. 28, fig. 15. Phos metuloides Brown and Pilsbry, t911, Proc. Acad. Nat. Sci. Phila., VOI. 63.) p.350. Phos metuloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 252, pl. 14, ASN 7.
A small species with a neat sculpture recalling that of M/e- tula cancellata and gabéi. It isnot rare in the Gatun beds of Mt. Hope in the Canal Zone. Dall also records it from the Mio- cene of Santo Domingo.
Length 19, diameter 9, spire whorl 13.5, spire 10 mm.
Gatun Stage: Mt. Hope.
295 Costa Rico MrockENE—OLSSON {27
Genus ALECTRION Montfort Alectrion ranuncula, n. sp. Plate 9, figures 20, 22
Shell of medium size, with a large body-whorl and a sharp pointed spire; the general form and sculpture recalls the recent A. acuta Say; spire about the same length as the aperture and canal and composed of about 7 whorls, of which the first 2 belong to the small, smooth nucleus; the profile of the spire-whorls is convex with deep, excavated sutures; sculpture of narrow ribs, the tops of which are crossed by spirals and separated by wide, smooth interspaces; the last whorl has about g ribs which are large and humped on the ventral face, finer and more numerous on the back; there are 3 spirals on the spire-whorls and 7 on the last with several more on the short canal; aperture subcircular with a large, heavy outer lip, internally denticulated.
The general form and sculpturing recalls the recent East Coast <Alectrion acuta Say, but the Costa Rican shell is much broader as indicated by the following comparative measurements with A. acuta Say, of the same height.
Height 14.00, diameter 6.25, last whorl 8, aperture 6 mm. (Alectrion acuta Say)
Height 14.50, diameter 8.25, last whorl 9, aperture 7.5 mm. (Alectrion ranuncula, n. sp.)
It is not uncommon in the Gatun beds of the Banana
River. Gatun Stage: Hill ra, Banana River. Coll. 7 Estrella.
Alectrion fosquemadica Maury Plate 9, figure 21
Alectrion losquemadica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 255, pls, Hgs: 22) 23;
The Costa Rica fossil is similiar in its sculpturing and form
to the common recent West Indian A. ambigua Montfort. As
Maury has pointed out in regard to the Doininican examples, the
124 BULLETIN 39 296
difference between losquemadica and ambigua, liesin the proto- conch, which in typical ambigua is short and blunt, but pointed and composed of several whorls in /osguemadica. Our shells are larger than typical losguemadica from Santo Domingo.
Height 11.50, diameter 7.00 mm.
Gatun Stage: Port Limon. Coll. 7, Estrella River.
Alectrion merenda, n. sp. Plate 9, figure 19
Shell small, a large body whorl anda sharp pointed spire; nucleus of 2 small, convex whorls followed by 5 post-nuclear whorls; sutures distinct but not excavated; the sculpture con- sists of about 12 ribs which are variable in their strength and spacing; the ribs are crossed by subequal spiral cords separated by wide, interspaces in which there may appear a smaller intermed- iate spiral; the suture is bordered by a set of 2 or 3 finer spirals; there are 3 spirals on the spire-whorls and 8 on the last whorl above the contracted zone of the canal; aperture subcircular with a thick, heavy outer lip.
Height 7, diameter 4.5 mm. The aperture is completely filled with a hard matrix which completely conceals the characters of the inner and outer lips. Uscart Stage: Old Man Sam Creek.
Genus NORTHIA Gray Northia northiae Gray, var. miocenica, n. var. Plate 9, figures 7, 14
ef. Northia northie Gray, 1834, in Griffith’s Cuvier, pl. 30, fig. 2. ef. Northia northie Kiener, 1832 Icon., Buccinum, p. 23, pl. 9, fig. 28.
The Northia northie Gray, (NN. serrata Dufresne) is a common and characteristic West Coast or Pacific species, ranging from the Bay of Panana to Guayaquil Ecuador. The fossils rep- resented by three incomplete specimens differ only from recent examples from Panama in lacking the hump-like thickening in
297 CosTA RicA MIOCENE ~- OLSSON 125
back of the outer lip and the corresponding shoulder to its whorls. But this is not an importnnt character and recent examples may sometimes lack this character.
But few investigators who have studied the West Iadian Miocene have failed to note the large percentage of species, close- ly related or even identical with recent Pacific species, a condition indicative of the union of the Atlantic and Pacific, permitting a free intermingling of their respective faunas. Late during the Miocene, this union was brought to a close or largely restricted, and since then the Pacific element has gradually given way before the encroachment and development of the West Indian. That this extinction has been a gradual one is shown by the Pleisto- cene deposits of Panama containg several comimon West Coast species not known from the recent Caribbean. Such species are the Pecten ventricosus Sowerby and Northia northie both record- ed by Dall from the Pleistocene of Panama or Costa Rica.
Gatun Stage: Middle Creek.
Banana River, Hill No. 3. Coll0) Red ‘Clap (Creek.
Genus COLUMBELLA Lamarck Columbella submercatoria, n. sp. Plate 10, figures 33, 34
Shell ovate, solid, a small conic spire and a large body- whorl; whorls about 7; spire-whorls but slightly convex so that the profile of the spire is nearly plane; last whorl large, broadly convex about the upper 1-3 and sloping evenly below to the short, produced anterior canal; sculpture of subobsolete spirals, there being about 6 on the spire-whorls and about 22 on the last whorl; aperture linear with a thickened outer lip, expanded in the middle and finely and evenly crenulated throughout; anteri- or canal short and armed with 8 strong denticles.
Length 17, diameter 10.5, aperture 11, spire 5.5 mm.
Closely related to the recent C. mercatoria Linn., common along the northern Costa Rican ccast, but difiers in its more
126 BULLETIN 39 298
globose form and finer spiral sculpturing. Traces of coloration are still preserved as faint blotches of yellow scattered over the back of the shell.
Typical Columbella has heretofore not been recorded from beds older then the Pliocene either in America or Europe.
Gatun Stage: Zone 5, Red Cliff Creek.
Genus STROMBINA Moerch
Strombina ambigua Guppy Plate to, figure 9
Columbella ambigua Guppy, 1866, Quart. Jour.Geol. Soc. London, vol. 22, Pp. 268, pl.16, fig S.
Strombina ambigua Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, PUG; py rS5oqe This species was described from the Bowden beds of Jamai- ca, of Miocene age. Our shells from Costa Rica agree exactly with Guppy’s figure in the Quarterly Journal.
A large species of Columbelloid aspect due in large measure to the slight thickening of its outer lip. The sculpture consists of about 25 ribs, which are smooth on the spire whorls and upper half of the last. The base of the last whorl and the canal have in addition about 14 spiral cords.
Length 27 (7 plus whorls), diameter 10, spire 15, aper- Cure 1 3) ia: Gatun Stage: Banana River.
Strombina Lessepiana Brown and Pilsbry Plate 10, figures 13, 20
Strombina lessepiana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila:, vol63;\p.\ 352) spl). 25, fies. 01, ha:
Columbella (Strombina) gatunensis Toula, 1911, Jahrb. der K-K, Geol. Reichsanstalt, Wien, vol. 61, p. 501, pl. 30, fig. 8.
An abundant species in the Canal Zone and with the excep- tion of S. ambigua, the largest species of S¢romézna in the Mio- cene beds of Panama and Costa Rica. The spire is long and smooth, with the exception that the upper sutural zone is usual-
299 Costa RicA MIOCENE—OLSSON 127
ly distantly raised into small, longitudinal plicaee which is most characteristic. An average specimen from the Canal Zone will measure: |
bength 25, diameter 10, spire 13, aperture 12.50 mm.
We collected from the lower Gattin sandstones of Cocles Creek, near Old Harbor, Costa Rica, several small Strombinas which except for size are typical Lessepiana.
Camm stage: Gatun and Mt, lope, C.' Z.
Cocles Creek. Cv R.
Strombina matima, n. sp. Plate to, figure 7
Shell ovate, moderately solid witha long evenly tapering and sharp-pointed spire; nucleus small, pointed and of three, smooth whorls; post-nuclear whorls 6; suture distinct and bordered an- teriorly by a small sutural ridge; the sculpture consists of about 18, smooth, narrow ribs, which are lacking from the back of the last whorl, which is smooth; the ribs are oblique in direction and along the upper sutures are slightly knobbed by the sutural ridge; the base of the last whorl and canal carries about 12 spirals; aper- ture subelliptical with a heavy outer lip internally provided with 3 or 4 denticles about the middle; anterior canal straight and smooth within.
Height 12, diameter 5.25, aperture 6 mm.
This is a species witha long, evenly-tapering spire and small nucleus. The spire-whorls are sculptured with numerous, small, smooth riblets. The riblets are obsolete on the back of the last whorl which is entirely smooth except for the spirals about the base and canal. S. matima resembles S. Basst Maury, from the Miocene of Santo Domingo but is more slender and has a great many more ribs.
Gatun Stage: Hill ra, Banana River.
Coll. 7, Estrella River.
Strombina cricamola, n. sp. Plate 10, figure 18
Shell ovate, solid, with a large, convex and spirally sculp-
128 BULLETIN 39 300
tured body-whorl and a long attenuated spire; whorls 6 plus (apex broken so that at least one complete whorl is lost from the type specimen); the first 4 whorls are smooth, after which a few incised spiral lines appear just below the upper suture; these spirals gradually increase in strength and in number and on the following whorls have become strong, raised spiral cords; the pen- ultimate whorl has 5 spirals, the last about 24 (including those of the anterior canal); the upper half of the last whorl and some- times the penultimate, are longitudinally plicated with 18 rib-like costez; aperture narrow, with a thickened outer lip, notched above and obsoletely denticulated below; the inner lip is smooth; canal short, twisted.
Length 16, diameter 8.5, aperture 9, spire 7 mm.
An unusually short and broad species, distantly related to the C. pulcherrima Sowerby, recent from the Gulf of Dulce. Two specimens were collected from the Gatun sandstones of Water Cay.
Gatun Stage: Water Cay.
Strombina costaricensis, n. sp. Plate 10, figures 8, 17
Shell solid, ovate, with a long attenuated spire; last whorl, large, flattened or contracted about the middle; whorls about 9%; the first 6 whorls are smooth and form the long attenuated spire; faint longitudinal plicee appear on the 7th, which increase in size and form the rib-like costee on the later whorls; axial ribs persist- ent and number on the last whorl about 15; the spirals begin shortly after the first appearance of the longitudinal plicee as faint incised lines near the upper suture; these spiral lines are further increased by the introduction of more lines below and be- come onthe following whorls raised spiral cords; the penulti- mate whorl has 5 or 6 spirals, the last with about 20; the spirals vary in strength on individual shells and on some specimens may be lacking from the middle of the whorl, forming a smooth me- dian band; aperture linear, small and coutracted and with a cal- loused inner and outer lip ; outer lip strongly thickened, notched above and feebly dunticulated below; canal short, straight.
ST en
301 CosTA RicA MIocENE—OLSSON 129
‘Length 15.5, diameter 7.5, spire 8, aperture 8 mm.
A species with an Azachis-like sculpture, the ribs and gen- erally the spirals being persistent. The body-whorl is flattened _ or slightly contracted about the middle, but less so than in the following variety. A long, tapering and nearly smooth spire as in the preceding .S. cricamola but much more slender and at- tenuated.
Gatun Stage: Headwater of Middle Creek, C. R. Strombina costaricensis, var. musanica, n. var. Plate to, figures 26, 27
Shell ovate, with a long, conic spire of 8 or more whorls; last whorl strongly contracted or indented about the middle; the first 3 whorls or more (apex broken) are smooth or sculpturless; on the following 3 whorls, the sculpture is largely of numerous, straight rib-like coste, the spirals being only very faint and in- distinct; on the last whorl the ribs number about 14; spirals about 9 on the penultimate whorl as faint raised threads and on the last whorl may be lacking from the middle or contracted zone; aperture linear-ovate, a thickened outer lip with 3 faint denticles below the upper notch; canal straight.
Length 15, diameter 6.5, spire 13, aperture 8 mm.
Probably a distinct species from the preceding and differing in its more evenly tapering and more strongly sculptured spire, and deeply contracted body-whorl. It resembles somewhat the figure of S. carfagensis Brown and Pilsbry from Colombia, but is smaller and has more numerous ribs. ‘The periphery of S. car- tagensis is rounded.
Gatun Stage: Banana River, C. R. Fast fork of Red Clif Creek.
Strombina sincola, n. sp. Plate 10, figures 19, 21
Shell small, with a large, conic spire and a dorso-ventrally flattened body-whorl; whorls about 8, the first 3 belonging tothe small, smooth, pointed nucleus; sculpture of the post-nuclear whorls of numerous, straight, strong, longitudinal ribs, which are
130 BULLETIN 39 302
persistent to the penultimate whorl, but are laking from the last whorl which is smooth and polished; the penultimate whorl has about 15 ribs; the ribs are generally beaded about their up- per extremities; sutures distinct; last whorl large and strongly laterally flattened between a prominent ridge or hump on the left- hand side of the whorl and the large outer lip; aperture sub- elliptical with a large, expanded outer lip, deeply indented be- hind; canal short and beak-like and spirally striated with about 8 threads on the back.
Length 7, diameter 3.25, spire 3.50, aperture 3.75 mm.
A small species abundant in the Gatun beds of Hiil ra, of the Banana River. The last whorl is rather small and strongly flat- tened dorso-ventrally. The spire is large and about % of the to- tal length of the shell.
Gatun Stage: Hill ra Banana River. fiill 37, Banana River.
Strombina chiriguiensis, n. sp. Plate 10, figures 14, 24
Shell small and in genral characters like the preceding but with a larger body-whorl and a shorter, more conic spire; whorls about 8, of which the first 3 are smooth and belong to the small pointed nucleus; the succeeding whorls until the penultimate are sculptured with fine,straight, longitudinal ribs,there being 19 on the penultimate whorl; the last whorl israther large, smooth and dorso-ventrally flattened between a hump-like ridge on the left side and the large, expanded outer lip; canal short, beak-like and spirally striated; aperture linear-ovate, deeply indented behind; inner and outer lips are both heavily calloused, especially about their posterior portions; over thiscallous and between the outer and inner lips passes the small posteriorly directed canal; lower half of the inner and outer lips denticulated with 8 or 9g strong, denticles, heaviest above. !
Length 7.50, diameter 4.25, spire 2.50, aperture 4.50 mm.
Of much the same size and general characters as S. szzcola of
OLSSON 132
303 CosTA Rico MIOCENE
Costa Rica, but with a larger and broader body-whorl, and a shorter, more conic spire. The porterior portion of the inner and outer lips are heavily calloused, a feature not developed in S. stncola.
Gatun Stage: Water Cay.
Genus MUREX Linnzeus
Murex messorius Sowerby
Murex messorius Sowerby, 1840, Proc. Zool. Soc. London, p. 137.
Murex recurvirostris Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.- 201. Not WZ. recurvirostris Broderip.
Murex messorius Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. VOl.,63, p. 353.
Murex messorius Maury, 1917, Bull. Amer. Pal., vol. 5, p. 265, pl. 16, MSS TD!
This is the common Murex in Costa Rica. It is generally small, seldom exceeding 40 mm in height. Each whorl carries three primary varices, between which are 3 or 4 smaller ribs or coste. It is arecent species in the West Indies.
Gatun Stage: Gainn, C. Z. Water Cay, Panama. ZOMe Fi nea, Clay Creer: Coll. 7, Pumbri Creek.
Banana River.
Subgenus PHYLLONOTUS Swainson
Murex cornurectus Guppy
Murex (Chicoreus) megacerus Gabb, 1873, Trans. Amer, Phil. Soc., vol. 15, p. 202. Not J/. megacerus Sowerby. ,
Murex cornurectus Guppy, 1876, Quart. Journ. Soc. London, vol. 32, Dw s2n, pli 28) fic. A:
Murex (Phyllonotus) cornurectus Maury, 1917, Bull. Amer. Pal., vol. 5. 207)pl. 16, figs..9, To.
132 BULLETIN 39 304.
A large species, possibly identical with the recent A7urex brevifrons Lamarck. ‘The varices are large and cross the spire whorls in a spiral direction. Between the varices, there is gen- erally a single rib-like elevation in the middle of the whorl.
Gatun Stage: Coll. 4, East Grape Point Creek. Coll. 5 Bed Oly Crear. Rio Betey. Sousz Creek.
Genus TYPHIS Montfort Typhis alatus Sowerby Plate 10, figure 15
Typhis alatus Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, p.45, pl to, aigs A.
Typhts alatus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 203.
Typhis alatus Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, D. 522.
Typhis alatus Brown and Pilsbry, 1911, Proc. Acad. Nas. Sci. Phila., vol. 63, p. 354.
A very characteristic species distinguished by the large wing- like expansion of its outer lip. It is the largest species in the Gatun and measures about 23 mm in height by 17 mm in diame- ter. It is found also inthe Miocene of Santo Domingo and in the Bowden beds of Jamaica.
Gatun Stage: Gatun, C. Z. (Brown and Pilsbry) Toro Cays, Panama.
Typhis linguliferus Dall, var. costaricensis, n. var. Plate 10, figs 22, 29
ef. Zyphis linguliferus Dall, 1890, Trans. Wagner Free Inst. Sci., vol. 2) Pe. Esp) 252 4 ols 12, dig. 7,
This is a smaller and more delicate speciesthan the 7. Gabdi Brown and Pilsbry from the Canal Zone. They agree with the Chipolan /énguliferus Dall in their general form but differ in their uniformly smaller size and in nearly lacking the spine-like pro-
305 Costa Rica MIocENE—OLSSON 133
cess on the shoulder of primary varices. It is fairly abundant in the Gatun beds of the Banana River.
Gatun Stage; Hill ra, Banana River.
Genus SISTRUM Montfort Sistrum nodulosum C. B. Adams Plate 10, figure 23
Purpura nodulosa C. B. Adams, 1845 Pree. Bost. Soc. Nat. History, pe) 3.
Ricinula nodulosa Tryon, 1880, Manual of Conch., vol. 2, p. 190, pl. 59, fig. 275.
Sistrum nodulosum Dall and Simpson, rgor, Bull. U. S. Fish. Com., VGls ty ps Arr:
Worn specimens of the species occur in the transitional beds of Red Cliff Creek. They cannot be separated from recent ex- amples from the present coast. It is very common as a recent species on the rocks and reefs along the northern Costa Rican and Panama coast, where it occurs about high-water level, asso- ciated with the equally as abundant Exgina turbinella Kiener and Phasianella affinis C. B. Adams.
Uscart and Gatun transitional beds; Zone 5 and 6,
Red Cliff Creek.
(OMS OPERA AMILY I eNIOGLOSSA
Genus DISTORTRIX Link
Distortrix simillima Sowerby
Triton simillimus Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, p. 48.
Persona simillima Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22. D2 286) pl.'17, hs. 13.
Distortia simillimus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 212)
Persona simillima Guppy, 1874, Geol. Mag., p. 439.
134 BULLETIN 39 306
Persona simillima Guppy, 1876, Quart. Journ. Geol. Soc. London, VOU 25.) 522°
Mstorsio ( Distortrix, Persona) gatunensis Toula, 1908, Jahrb. der K-K Geol, Reichsanstalt, Wien, vol, 58, p. 700, pl. 25, fig. 10.
Mstorsio gatunensis Brown and Pilsbry, 1Igt1, Proc. Acad. Nat. Sci. Phil Vvol62) 1 pare 56.pl. 20) hig aS, Distortrix simillima Maury, 1917, Bull. Amer. Pal., vol. 5, p. 271, pl. Types. 5: Not common in Costa Rica. It also occurs in the Miocene of Jamaica and Santo Domingo.
Catun Stage "Gann, (Cui Banana River. Port Limon.
Genus BURSA Bolten Bursa crassa Dillwyn Plate 15, figure 15
Ranella crassa Dillwyn, Reeve, Conch. Icon.,fig. 18.
Ranella crassa Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22, p.'288), pl. 18) fig. .o; é
Bursa crassa Gabb, 1873, Trans. Amer. Phil. Soc:, vol. 15, p, 212.
Ranella crassa Guppy, 1874, Geol. Mag., p. 438.
Ranella crassa Guppy, 1876, Quart. Jouru. Geol. Soc. London, vol. 32, pews 22
Bursa crassa Maury, 1917, Bull. Amer. Pal., vol.°5,/ 272.4? figs.16, 675
This is a recent species in the West Indies. It occurs also
as a Miocene fossil in Jamaica and Santo Domingo.
Gatun Stage: Banana River. Saury Creek.
Genus CASSIS Lamarck Cassis sulcifera Sowerby
Cassts sulcifera Sowerby, 1849, Quart. Journ. Geo 1. Soc. London, vol 6; p47. pli 10, er
307 Costa R1cA MIoCENE—OLSSON 135
Cassis sulcifera Maury, 1917, Bull. Amer. Pal., vol. 5, p. 274, pl. 18, HG! T, 2.1/2)
This is a large species very common in Miocene rocks of Santo Domingo but very rare in Costa Rica. Our record is bas- ed on asingle imperfect specimen from the Banana River and for a figure of this species the reader is referred to Dr. Maury’s Do- minican Fossils.
Gin stage: fill No. 3, Banana River.
Cassis fiammea Linnzeus
Buccinuim flammeum Linneeus, 1758, Syst. Nat., ed. 10, p. 736.
Cassis fiammea Reeve, 1848, Conch. Icon., vol. 5, pl. 5, fig. 12.
Cassis fammea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 356.
This is a recent species recorded by Gabb, from his so-called Pliocene clay beds of Costa Rica. Under this term, Gabb con- fused two formations, one composed largely of coralline hme- stones but with subordinate amounts of clays and sands equiva- lent in large measure to the Gatun formations,and an overlying, mostly sand and clays of true Pliocene age. Itis most likely that the single example of Casszs flammea collected by Gabb, was obtained from the upper or Pliocene formation.
Subgenus PHALIUM Link
Phalium moniliferum Guppy Plate 12, figure 11
Cassis monilifera Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 2390287, pl.i17, fig. 8:
Casidea granulosa Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 222. Not C. granulosa Bruguiere.
Cassis reclusa Guppy, 1874, Geol. Mag., London. pp. 434, 439.
Cassis reclusa Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, Pp. 525.
Cassis monilyera Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, P. 525-
Phalium moniliferum Maury, 1917, Bull. Amer. Pal.. vol. 5, p. 274, pl. LOmmMSS. A \5> pl. 19, fio.’ 1
136 BULLETIN 39 308
The Costa Rican specimens like those of Santo Domingo are extremely variable in their sculpture. ‘The typical monzliferum of Jamaica carries on the shoulder of the body-whorl one or two rows of large tubercles or granules and Guppy’s figure shows the spirals below as finely granulated.
In Costa Rica, the shells may vary from those with nearly smooth spirals, through transitional forms in which only the su- perior bandsare granulated,to the extreme in which all the spirals are finely granulated over the whorl shell. A typical specimen will average:
Length 25, \\diameter 22) mm)
Gatun Stage: Middle Creek. Rio Betey. — Zone 5. hed Cluj (Cream
Genus SCONSIA Gray
Sconsia laevigata Sowerby, var. Gabbi, n. var. Plate 12, figure 3
cf Cassidaria laevigata Sowerby, 1849, Quart. Journ. Geol. Soc. Lon- dons vol /6)pmA7. pl clo wie, 2.
cf Cassidaria sublaevigata Guppy, 1866, /dem, vol, 22, p. 287, pl. 27, fig 0:
Cassidaria laevigata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series; p. 356.
cf Sconsia sublaevigata Bose, 1906, Bol. Inst. Geol. de Mexico, Numero 22).D, (36, (pls iiA, TSS, 1, (LO.
Sconsta laevigata Prown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. V.Ol-637) 9.1250.
ef Sconsia laevigata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 275, pl. 10) 119.112.
As Gabb noted years ago,the Sconsias of Costa Rica are not the typcal /aevigata of Santo Domingo but appear to approach more closely in their striated whorls, the Jameican sudblaevigata of Guppy and the recent sfrzata of Lamarck.
Typical Jaevigata, has generally a distinct shoulder to its
309 Costa Rica MIocENE—OLSSON D7
body-whorl and a lower spire. The spire-whorls are striated but the spirals are lacking from the generally polished body-whorl. The aperture of /aevigata is subrectangular in outline, rather wide anteriorly and flaring posteriorly, while in Gabddz, the aper- ture is narrow and but little larger anteriorly. In /aevigata, the callus of the inner lip is large and spreads widely over the colu- mellar area, but its outer edge is raised and shelf-like, instead of rounded and appressed asin Gadbdz. Mature shells of laevigata, have three large, heavy varices, marking the resting stages but such varices are small and sometimes entirely lacking from the Costa Rican shells.
Typical laevigata will probably be found to be largely limit- ed to Santo Domingo. ‘The Jamaica sudlaevigata (possibly a var- iety) has more strongly sculptured whorls, a short spire and less complete varices. Its aperture is that of true /aevigata.
The Sconsia laevigata was found by Dr. Maury, in her work on the Paleontology and Stratigraphy of Santo Domingo to be limited to her upper or Gurabo formation (the-Sconsia laevigata zone). Ir Panama and Costa Rica, the Sconsia laevigata variety Gabbi, is frequently very abundant in the Gatun beds, but it is apperently lacking from the underlying Uscari shales where it is replaced by the new Scozsza cocleana and probably docasensts.
Cam Stage: Gatun, C. Z. Toro Cay, Water Cay, Panama. Comadre Creek near Cahuita C. R. FIO DIAN CON OO, A.
Sconsia bocasensis, n. sp. Plate 12, figures 12, 13
Shell like that of /aevzgata,but narrow and with coarsely sculp- tured whorls; spire more evenly conic, the apex of which is not produced or attenuated; whorls coarsely sculptured with heavy, persistent spirals threads, of which there are about 9 on the spire- whorls and nearly 50 on the last whorl; sutures appressed and bordered anteriorly by a strong ridge-like cord or by stronger spirals; the varices are but slightly developed; aperture subellip-
138 BULLETIN 39 310
tical, widest about the anterior 1-3; outer lip thickened and with about 18, fine, lirce-like denticles ; the inner lip with a thin spred- ing callus and with about 16 long, narrow rugations.
Length 35 (last 2 whorls only) diameter 25 mm.
Differs from the variety Gadd of laevigata by its more narrow shell, coarser sculpture and larger aperture. The suture is bord- ered by a cord-like ridge. .
Gatun Stage: Bocas del Toro. Sconsia cocleana, n. sp. Plate 12, figure 7
Shell globose; spire low of about 7 whorls, the apex slightly projecting; sutures distinct and deep, but not appressed; surface coarsely sculptured with large, irregular, but narrow spirals bands separated by deep interspaces which may in addition carry one or more finer spiral threads; there are 7 or 8 spirals on each spire whorl; on the last whorl the spirals are nearly twice as wide as the upper third of the whorl and more anteriorly; no varices; the outer lip is broken from the single specimen; inner lip is a wide but thin wash of callus, without rugations or denticles of any sort.
Length 46, diameter 32 mm.
The Uscari shales which everywhere in Costa Rica appear to underlie the Gatun sandstones is very rich in the smaller types of foraminifera, but mollusks are rare, fragmentary and difficult to collect. Fragments of Scozsta, were noted by us at several places which we believe belong to this species or to the Jocas- ensts.
This species will be recognized by its broad, globose shell and small spire. It lacks the sutural cord so strikingly develop- ed on bocasensts.
Uscari Stage: Rio Cocles near Old Harbor, C. R.
3ir Costa Rico MICCENE—OLSSON 139
Genus DOLIUM Lamarck
Subgenus MALEA Valenciennes Malea elliptica Pilsbry and Johnson Plate: 123). figure 2
Malea elliptica Pilsbry and Johnson, 1917, Proc. Acad, Nat. Sci. Phila., vol. 69, p. 169.
The true Malea camura Guppy, so common in the Miocene of Jamaica and Santo Domingo still remains to be collected in Costa Rica. It is usually a smaller species (height 60 mm) with a heavier shell and high, narrow, special bands numbering about 16 or 17 rather widely separated.
The 1. elliptica is larger (height 60-80 mm or more), more globose and thin. The spiral bands numbering about 22 are closely spaced and ribbon-like. The coiling is regular, while in camura the coiling is irregular and descending in the latter stages of growth.
The Malea elliptica was described but not figured by John- son and Pilsbry from the Miocene of Santo Domingo.
Height 63, diameter 54 mm. Uscart Stage: Red Cliff Creek. Gatun Stage: Bocas del Toro. Estrella River. flill ra, Banana River.
Genus CYPRAEA Linné Cypraea parisimina, n. sp. Plate 12, figure 10
Shell small, more or less depressed and with expanded, an- gulated sides; dorsal surface convex, but with a deep pit or depression at its posterior end, beyond which project the 2 short, pouting ends of the lips of the aperture; ventral sur- face flat, with a narrow aperture which is straight except near at its posterior end where it is slightly curved; lips strongly but regularly crenulated (about 21 on each lip) posterior sinus
140 BULLETIN 39 312
small and curved to the left; anterior sinus small and round- ed.
Length 39, basal diameter 25, height or vertical diame- tet, 1.7 mm: A small shel! with strongly depessed and angulated sides. Collected from the coral limestones of Port Limon. Gatun Stage: Port Limon, Cpyraea cf. dominicensis Gabb Cypraea Dominicensis Gabb, 1873, Amer. Phil. Soc., vol. 15, p- 236. Cypraeca dominicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 280, plop to} Se. oT Our collection contains a single, fragmentary shell from the coralline limestones near Limon. This specimen agrees closely with the Dominican shell figured by Dr. Maury as C. domztnicen- sts ?,_ It is characterized by its very thin shell, cylindroid form and straight, evenly denticulated apertrue. Length 28, height 12 mm. Gatun Stage: Port Limon.
Cypraea cinerea Gmelin Plate 12, figure 8 Cypraea cinerea Gmelin, 1792, Syst. Nat., p. 3404. Cypraca cinerea Reeve, \Icon., vol. 3, pl.22, fg. 124.
Cypraea cinerea Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 360.
The Cypraea cinerea isa common, small species of Cowry shell along the north coast of Panama and Costa Rica. Several small Cypraeas were collected from the Gatun sandstones of the Rio Blanco which cannot be distinguished from recent ex- amples of cizerea.
Gatun Stage: Rio Blanco.
Cypraea aimirantensis, n. sp. Plate 12, figure 9
Shell large, heavy and rather high; the outlines of the shell
313 Costa RicA MIocENE—OLSSON 14!
from below, the side and from above is nearly rectangular, but with the anterior extremity more pointed; the shell is high (about ¥% that of the length), with nearly flat or slightly convex sides; dorsal surface convex, a deep depression or pit about the posteri- or 1-4 and low, but large tubercles on each side of the posterior sinus; aperture narrow, curved, with the lips coarsely but regu- larly dentate (about 20 on each lip); posterior sinus is long and vertical; anterior sinus small and rounded; basal surface flat.
Length 60, basal diameter 39, vertical diameter 30 mm,
This is a species but distantly related to the C. Henkeni So- werby, differing by its longer and higher shell, straight sides and smaller dorsal tubercles. The aperture is strongly curved and very deep on the posterior extremity, ending in the long, vertical sinus.
Gatun Stage: Water Cay, Panama.
Genus STROMBUS Linnzeus
Strombus gatunensis Toula Plate 13, figures 5, 6
Strombus gatunensis Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt Wien; vol. 58, p. 673, pl..25, fig. 7:
Strombus gatunensis Brown and Pilsbry, 1gr1, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 355, pl. 26, figs. 3, 5.
But two species of Stromdus are known from the Gatun beds of Panama and Costa Rica, while five species occur in the synch- ronous Miocene strata of Santo Domingo and four in Jamaica. Other species may therefore be expected with further explora- tion work in Costa Rica.
In S. gatunensts, the coiling of the whorls follow closely the shoulder angle so that the last whorl appears very large and the spire low but wide. Young shells occasionally have the shoulder armed with few, large tubercles which later become nearly cov- ered by the close coiling and appressed sutures. The last whorl is large, nearly smooth and with a rounded shoulder angle.
Occasionally traces of the original coloration is preserved
142 BULLETIN 39 3r4
and consists of narrow, yellow V-shaped lines, the acute base of which are directed backwards or away from the aperture and lie along the center of the whorls. Similiar markings may be seen on Veatcht and Guppy* has described the same markings for pugiloides from Jamaica. Length 63, diam. 42 mm. Gatun Stage: Gatun, C. Z. Water Cay, Panama. fleadwaters of Middle Creek, C. R. Strombus pugiloides Guppy Gre! 13, figures E,\2\'5 Strombus pugilis Guppy, 1866, Quart. Geol. Soc. London, vol. 22, p. 287. Strombus pugilis Gabb, 1873, Trans. Amer. Phil) Soci, wel 3. p- 233. Strombus pugiloides Guppy, 1874, Geol. Mag., new series, vol. 1, p. 433. Strombus pugilis Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, p. 340. Strombus pugilis Bose. 1906, Boll. de Inst. Geol. de Mexico, numero 22, p. 35, pl. 4, figs. 1-6. Strombus pugiloides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 284, pl- 20, f1g))'G.
A smaller species than the gafunensis, from which it also differs by its longer spire and more strongly sculptured shell. The coiling follows below the shoulder of the whorls, which is either simply rounded or bears several large, spine-like tubercles asin the recent pugzlzs. Resting stages develop large, hump- like areas irregularly distributed on the spire-whorls. Varieties occur which may be spirally sculptured over the whole shell or the last whorl may be largely smooth. The original coloration is still preserved on some shells and consist as in gatunenszs of narrow, yellow, V-shaped lines, the apex of which lie about the middle of the shell and directed backwards. It is very distinct from the recent West Indian pugz/zs,which does not appear to de- scend below the Pliocene.
* Proc. Sci. Assoc, Trinidad, 1873, vol. 2, p. 82,(reprint: Bull. Amer Pal., vol:'3)'p! 214).
ae
315 Costa RicA MI0CENE—OLSSON 143
Uscari Stage: Sapote, C. R. (Gadd) Gatun Stage: Banana River. Zone Fi, Saury Creek. Zon 5). Bast Grape Fomt Cretk.
Genus CREPITACELLA Guppy Crepitacelia limonensis, n. sp. Plate 15, figure 17
Shell small, melanoid in form,with a large body-whorl and shorter conic spire; nucleus of 2 small, smooth whorls followed by 5 post-nuclear whorls; sutures distinct; the whorls are slight- ly coronated and crossed by oblique and slightly curved riblets which become headed on the coronate edge of the spire-whorls just below the upper sutures; on the last whorl, the ribs do not pass below the middle; the surface is covered with faint spiral threads, most distinct on the base of the body-whorl; aperture elliptical, with a thin, oblique outer lip and a subobsolete anteri- or canal; pillar straight.
Height 8, diameter 4.5, aperture 4 mm.
The Costa Rican shell, although possibly not mature,is much smaller than the C. cebula Guppy from the Miocene of Venezuela and Santo Domingo of the same number of ribs. It differs also in having but 2 nuclear whorls, while cepula has 3. The corona- tion of the whorls which is such a striking feature of the Domin- ican shell, is very much less and inconepicuous. The longitud- inal costee are more numerous and produce a fine circle of beads as they cross the small coronate edge of the spire-whorls.
Gatun Stage: Port Limon,
Boras del Toro.
Genus CERITHIUM Adanson
Cerithium costaricensis, n. sp. Plate 10, figure 28
Shell with a long, slender spire of about 12 whorls (tip brok- ed); sutures deep but somewhat appressed; the profile of the
144 BULLETIN 39 316
spire-whorls is convex with an occasional hump-like resting mark; the sculpture consists of primary spiral threads between which lie x, 3, or more finer secondaries; the earlier spire-whorls have heavy ribs but they become replaced on the later by simple gran- ulations on the primary spirals; a narrow, sloping band borders the anterior side of the suture and carries finer, non-beaded spiral threads: below this band, the later spire-whorls bear about 4 beaded, primary spirals and finer secondaries; the last whorl shows 6 primaries together with other spirals on the base and canal; aperture subcircular, with a moderately expanded outer lip, obli- quely produced anteriorly.
Height 28,. diameter 10.5, aperture 3) nam
The general form and sculpture of this species is that of the recent West Indian C. eburneum Bruguiere. ‘The fossil shell is larger, more slender and differently sculptured.
Gatun Stage: Coll. 6, Red Cliff Creek.
Genus CLAVA Martyn Clava costaricana, n. sp. Plate 10, figures 3, 4
Shell of medium size, cerithoid; whorls about 15, the early spire-whorls very small; the last whorl carries a large hump on the ventral side of the shell to the right of the aperture, so that the shell appears flattened on this side; the sculpture consists of strong spiral cords which are coarsely granulated by small, close- ly spaced longitudinal costze; the upper spiral cord bordering the suture is somewhat larger than the other; there are 3 spirals be- tween the sutures of the spire-whorl and 6 on the last whorl with additional smaller ones on the base of the shell; the outer lip is broken on all our specimens; the columella carries 2 strong folds.
Height 4o, diameter 15 mm. Ans I4 mm.
This species bears some resemblance to the C. caloosaénsts
Dall from the Pliocene of Florida. The Costa Rican shell is
317 Costa Rica MiocENE-—-OLSSON v:
ay tr
very much larger, broader and the longitudinal costee are more rib-like, continued across the whorls, crossing the interspaces as well as the spiral cords.
Gatun Stage: Banana River. Clava aiajuela, un. sp. Plate 10, figure 30
Shell small, slender; whorls 12 plus, the apex broken; the sides of the whorls are straight and form the narrow, tapering spire; sutures indistinct; the sculpture consists of nearly equal ribs and spiral threads; the early spire-whorls have 3, equal, spiral threads, separated by interspaces as wide as the spiral threads and deep; on the later whorls fine, intermediate threads appear in the interspaces and a fourth large spiral in the lower suture; the spirals are crossed and strongly beaded by straight or slightly curved, longitudinal ribs, which number on the last whorl about 25 or 24; heavy and thickened resting marks ap- pear on the later whorl on each volution; the aperture is broken, the columella straight and witha single small fold.
Height 17 mm, diameter 6 mm.
A small and finely sculptured species, represented by the single type specimen. Gatun Stage: Middle Creek.
Genus SERPULORBIS Sassi
Serpulorbis papulosa Guppy Plate 12, figure 1
Vermetus papulosus Guppy, 1866, Quart. Journ. Geol. Soc. London, MOM) 22s) Di 292, pl. 17) fig. 3. Vermetus papulosus Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, p. 519. Sespulorbis papulosa Maury, 1917, Bull. Amer. Pal., vol. 5, p. 291, pl. 22, fig. Io. Considerable variation may be seen in the Costa Rican shells, from specimens in which the sculpture is quite coarse and with the interspaces between the main cords carrying only one strong
146 BULLETIN 39 318
intermediate cord, to more delicately sculptured forms with finer tuberculated cords separated by wide and finely sculptured inter- spaces.
This species occur also in the Bowden beds of Jamaica and in the Gurabo formation of Santo Domingo.
Gatun Stage: Hill 1, 2, Banana River.
Genus PETALOCONCHUS H. B. Lea Petaloconchus sculpturatus, H. C. Lea Plate 14, figures Io, I5
Petaloconchus sculpturatus H. C. Lea, 1845, Trans. Amer. Phil. Soc., VO). /O}.p: 233) ple 24h fig ss.
Fetaloconchus domingensis Sowerby, 1849, Quart. Journ. Geol, Soc. London, vol. 5, p. 51, pl. 10, figs '8)’a ibe:
FPataloconchus sculpturatus Gabb, 1875, Trans. Amer. Phil. Soc., vol. 25D. 240,
Pataloconchus sculpturatus Guppy, 1876, Quart. Journ. Geol. Soc. Lon- don, vol, 32; p. 519.
Vermetus (Petaloconchus) sculpturatus Dall, 1892, Trans. Wagner Bree Inst: Ser, wol.35, pty 2:0p 6405:
Vermetus (Petaloconchus) pulcher Bose, 1906, Bol. Inst. Geol. de Mex- 1€0,, Numero 22.9.) 32) pling, diessi22, (23)
FPetaloconchus domingensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 359.
Petaloconchus domingensis Pilsbry and Brown, 1917, Proc. Acad. Nat. Sel. Phila), vol.6o.p.) 37.
Fctaloconchus domingensis Maury, 1917, Pull. Amer. Pal., vol., 5, p. 292, Olt 22. 0h oe rie
fataloconchus domingensis Wubbard, 1920, New York Academy of science; Sci. Sur. PLR.) vol.3; "pt, 2; p. ge:
The Antillean fossil was described by Sowerby as domingen- sis. It has been by some writers, considered synonymous with sculpturata, described a few years earlier by H. C. Lea from the Chesapeake Miocene of the eastern United States, while others have given domingensis the rank of a full species. A close com- parative study of the Costa Rica fossils with several sets of typi-
319 " Costa Rico M1ocENE—OLSSON 147
cal sculpturata reveals no distinction between them. Both are variable in their coiling from closely wound, narrow cylinders to more open, umbilicate coils. The sculpture is much the same.
In the United States, this fossil is limited to the Upper Chesapeake Miocene. In the Antillean and South American re- gions, itis found in the Bowden beds of Jamaica, the Gurabo formation of Santo Domingo, the Quebradillas limestone of Por- to Rico, the Springdale beds of Trinidad, from the Gatun beds near Cartagena, Colombia and Mexico.
Gatun Stage: Gatun Stage, C. Z.
Water Cay, Panama. COHN, Gi toast’ Grape Point | Creek. Comadre Creek.
Genus SILIQUARIA Lamarck
Siliguaria modesta Dall, var. limonensis, n. var, Plate 12, figure 4, 5, 6
cf Siliguaria modesta Dall, Mus. Comp. Zooi., vol. 9, p. 39.
ef Stliguaria modesta Dall, 1889, Bull. Mus. Comp. Zool., vol. 18, p. Z200s"pls'26, fig. 4.
This shell is very abundant in certain marly and sandy beds in the coralline limestones of the Limon Peninsula and elsewhere. Specimens in perfect condition are very difficult to obtain,the shell being very fragile and delicate. There are no longitudinal threads and the shell is smooth, except for the growth lines which are sometimes a little irregular.
We have associated this shell with the S. modesta of Dall, a recent species and recorded by Dr. Dall from several stations in the West Indies and the Gulf of Mexico, in waters ranging from 94 to 805 fathoms in depth. The fossils differ in being more loosely coiled and at the later stages long drawn out.
Dr. Maury’s S. guarabensis from the Miocene of Santo Do- mingo is more irregular in its habitus of growth and the surface of the shell carries faint longitudinal threads. Gabb has des-
148 BULLETIN 39 320
cribed as sculpturata a Siliquaria from the Limon Peninsula. This is united by Dall with the recent West Indian sguamata Blainville. I have seen no spicimens and Gabb may have collect- ed it from the true Pliocene beds of the Limon Peninsula. Gatun Stage, Island of Bocas del Toro. Port Limon.
Genus TURRITELLA Lamarck Turritella gatunensis Conrad Plate 14, figures 12, 13
Turritella gatunensis Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72, pl. 5, fig. 20. Turritella conradi Toula, 1909, Jahrb. der K-K Geol. Reichsanstalt, vol. 58, p. 694, pl. 25, fig. 4. Turritella gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol.'63, pl.,27) figs. °4) 5,9: The Zurritella gatunensis is a medium sized shell seldom over 40 mm in length. The base of the whorl overhangs the next, and each whorl is encircled by a median concave or constricted band. The whole surface it finely sculptured with small spiral threads. The spirals defining the median band are usually slight- ly heavier as well as a few about the upper part of the base.
The 7: gatunensts is a common and widely distributed species in the Gatun beds, but always less abundant than the I. altilira or its varities. Like the altiliva, zones of JT. gatun- ensis are frequent in some localities.
The 7: atacta Dall of the Tampa Silex beds, the 7. acropo- ra fossilin the Pliocene of Florida and recent along the east coast of United States and the West Indies, are related species.
Gatun Stage: Gatun, C. Z.
Water Cay, Panama. Boucary Creek, C. R.
321 Costa RicA M1ocENE—OLSSON 149
Turritella mimetes Brown and Pilsbry Plate 14, figure 5
Turritella mimetes Brown and Pilsbry, tIg911, Proc. Acad. Nat. Sci. Boilas) vol. 62, p. 357, pl.27, fg. t.
Turritella (flaustator) aff. Hanleyana Reeve- 7. lineolata (Kiener) Toula, 1911, Jahrb. der K-K Geol, Reichsanstalt, vol. 61, p. 401, pl. 30, figures 6a and 6b, A large, robust species in form and sculpture like the 7. variegata Linnzeus, recent in the West Indies.
The whorls are straight and usually slightly overhanging the lower sutures. Surface sculptured with primary, secondary and tertiary spiral threads. The primary threads are irregularly disposed and usually number about 7 or 8 to each whorl. Be- tween the primaries are the finer secondaries and tertiaries.
Small shells may be mistaken for the gazunensis, but the whorls are flat, without the median concave and constricted zone and the sculpture is more coarse. The figured shell of 14 whorls has a length of 70 mm and a diameter of 18 mm. Ai larger but more imperfect specimen of but 4 whorls measures:
Length 50, diameter 20 mm.
Gaim) Stage. Gatun, C. Z. Turritefla altilira Conrad, and varities.
Turritella attilira Conrad, 1857, Pacific R. R. Reports, vol. 6, p. 72, pl. 5; fig. 19.
Turritella gabbt Toula, 1909, Jahab. der K-K Geol. Reichsanstalt, p. 695, pl. 25, fig. 5.
Turritella altilira Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Bitla.;’ vol:63. p: 358) pl. 27, figs. 2, 3.
Of the species of 7urrztella in the Gatun beds of Costa Rica and Panana, the 7. altilzra of Conrad and its varieties is the most common and characteristic. The 7. fornata Guppy, from the Miocene of Trinidad and Venezuela is a closely related species.
These magnificent 7urritelle are preminently characteristic of the West Indian and Caribbean Miocene, but probably deriv- ed from earlier forms in the Oligocene. The stock continued in-
150 BULLETIN 39 322
to the Pliocene where its acme of development was reached in the elegant 7. preaticnuata Heilprin of Florida. It has left no descendants in the recent fauna.
The shell is long and tapering, each whorl sculptured with 2 prominent beaded spiral cords, between which the surface 1s con- cave or deeply channeled and may be smooth or with finer spiral threads. The upper spiral cord is simple or double. Several varieties may be recognized which differ in details of their sculp- ture, size and form of shell.
T. altilira Conrad, typical Plate 14 figures: 6,°7
This, the most elegant form is limited to the small Gatun sedimentary basin of the Canal Zone and the adjacent Province of Colon.
The shell is long, slender and with numerous whorls; each whorl is strongly sculptured by the two, primary spiral cords, each of which is bordered on adjacent faces by a smaller spiral, between which lies a deeply channelled groove; small- er beaded spirals occur in the median channel and are best devel- oped on the spire whorls; the beading or granulation of the spirals is produced by the arcuated lines of growth, the sinus of which lies a little above the middle of the shell; the upper spiral cord is larger than the lower and more strongly beaded and in general becomes double on the later whorls.
Gatun Stage: Canal Zone and Province of Colon. Var. chiriquiensis, n. var. Plate 14, figures 4, 8, 9, 14
Shell larger, less attenuated and more coarsely sculptured than typical altlira; the 2 spiral cords are less strong, of more nearly the same size and usually remaining simple; the 2 spiral cords enclose a wide, concave area of about % the width of the whorl; this concave band usually carries 2, 3, or more fine spiral threads; the beading or granulation of the spiral cords is feeble and best developed on the earlier spire whorls and in certain large,
333 Costa Rica MiocCENE—OLSSON 151
gerontic shells the spiral cords may become nearly smooth.
This is the common form on the islands of the Chiriqui La- goon, and of eastern Costa Rica. Likethe typical a/¢zlira it is often a very abundant fossil, forming zones of Zurrztella in the Gatun beds.
The sculpture is much less elegant than that of true a/¢zlzra, and consists mainly of the 2, primary spiral cords, between which lies a wide concave band. This band is but feebly sculptured by small, spiral threads, or in some cases, is smooth.
In small matters of detail of sculpture and in the form and taper of the shells, there is considerable variation. Such varia- tion is usually of but local signifieance and in general the shells collected at the same or nearby localities will agree closly with each other but differ more or less from those of more distant areas.
The 7. fornata Guppy from the Miocene of Venezuela and Trinidad should probably be considered asa variety of a/¢zlzra, most closely allied to the chizvzquensts. Its sculpture is like chzr- zquensts but with stronger spiral threads on its median band.
Gatun Stage: Water Cay. Grape Potnt and Red Cliff Creeks, C. R. Banana River. Rio Blanco.
Var. costaricensis, n. var. Plate 14, figure II
Shell small; whorls less numerous and more rapidly increas- ing so that the shell is porportionately broader and shorter; su- tures deep; the 2 primary cords are but weakly developed, be- tween which lies, the concave band as in chiriguzensis,; the band carries 3, 4, or more strong, irregular spirals, but slightly weak- er than the primaries; the granulation of the cords is weak and irregular.
Much smaller than the preceding varieties (usually about 40 mim) and with wider and more rapidly increasing whorls. The
152 BULLETIN 39 324
cords on mature shells are small and but slightly heavier than the spiral threads on the concave band.
This type replaces the chiriguiensis in the Gatun sandstones in the area of Upper Hone Creek and Boucary Creek. It is often extremley abundant, associated with the TZurritella gatunen- S725
Gatun Stage: Upper Hone and Boucary Creeks. Turritella exoleta Linnzeus, var. limonensis, n. var. Plate 14, figs 2, 3
Shell turreted, the whorls increasing more rapidly in diame- ter than typical evo/eza, the nucleus is small, immediately follow- ed by whorls carrying a median carina which gradually becomes more anterior in position and later the lower carina of the adult sculpture; the whorls are deeply concave or excavated about the middle between 2 principal carinz; on the last whorl there is seen to be a third carina which is concealed in the suture; the su- tures are excavated, becoming generally deeper with age; the surface is nearly smooth, except for very faint spirals and rough growth lines which may be raised and sublamellose on the medi- an concave zone.
Height 4o (6 later whorls) diameter 17.50 mm. Height 35 (10 earlier whorls) diameter 10 mm.
This is probably a distinct species from the recent West In- dian Zurritella exoleta Linneeus, differing in its more rapidly ex- panding whorls. Faint spirals may generally be seen covering the entire shell and the growth lines become raised and sublamel- lose in the concave medial zone. It is acommon shell in the cor- alline phase of the Gatun at Port Limon and elsewhere.
Gatun Stage: Port Limon. Bocas del Toro,
Turritella oreodoxa, n. sp. Plate 14, figure 1
Shell large, solid and very slender; whorls numerous and each but slightly larger than the preceding, slightly concave or constricted about the upper 1-3, above which the edge of the
325 Costa Rica MiockEN#—OLSSON 153
whorl is enlarged, forming a cord-like expansion about the upper suture; sutures indistinct and appressed; sculpture of fine, ir- regular and somewhat wavy spiral lines and indistinctly alter- nating in strength; growth lines as typical for HYaustator (not visible on the base of our specimen), commence at the lower suture and pass straight and parallel to the longitudinal axis of the shell to the middle of the whorl where they become bent™and continue obliquely to the left or towards the aper- iire to the upper suture. Length (2% whorls), 49, diameter 23 mm.
A very rare species and represented in our collection by only two imperfect specimens. It appears to be closely related to the Z. cartagensis Pilsbry and Brown from the Gatun of Colombia. The full-grown Costa Rican shell was probably longer and more attenuated and each whorl is prominently elevated or ridged just in front of the lower suture.
Gatun Stage: Rio Banana beyond Hill No. 5.
Genus ALABA A. Adams
Alaba turrita Guppy, 1896, Proc. U. S. Nat Mus., vol. 19, p. 321, pl. 2 Spy te 7).
Guppy’s figure of his type specimen from Jamaica agrees very closely with our specimen from the Estrella River. The whorls are finely spiralled with low ridges which are obsolete from the upper parts of, the whorls. Each whorl carries 2 or more broad, smooth varices formed during resting stages. The last whorl is strongly angulated by these varices.
Height 5, diameter 2.25 mm. Gatun Stage: Coll. 7, Estrella River.
154 BULLETIN 39 326
Genus ARCHITECTONICA Bolton
Architectonica granulata Lamarck Plate 13, figure 10 ,1I, 12
Solarium granulatum Yamrack, 1822, An. sans. Vert., vol. 7, p. 3,
Architectonica prespectiva Tuomey and Holmes, 1857, Pleioc. Fos. S. Cy, p:/ 120, ple 26, tre:
Solarium granulatum Dall, 1892, Trans. Wagner Free Inst. Sci., vol. 3, pt. 2, p. 329.
Solarium Villarellot Bose, Inst. Geol. de Mexico, Numero 22, p. 30, plea igs. 4-1
Solarium gatunense Toula, 1908, Jahrb, der K-K Geol. Reichsanstailt, Wien, vol. 58, p. 693, pl 25, fig. ¢.
Solarium granulatum Maury, 1917, Bull. Amer. Pal., vol. 5, p. 295, plies fgets:
The Costa Rican fossils are usually quite typical and not to be distinguished from recent examples of granulata. They vary somewhat in their height and persistence of their spiral sculpturing. The beaded and granulated character of the spir- als generally becomes obsolete on the later whorls so that the
spiral bands appear smooth. The umbilicus is deep but nar- row.
This is a stable species ranging from the lower Miocene to the recent. Its recent distribution extends from Cape Hat- teras on the north, southward through the West Indies and on the Pacific side from the Gulf of California to Peru.
Height 20, greater diameter 43 mm. 19, 29 mim. 1 20 mm. Galun Stage; Gatun, CZ. Water Cay, Panama. Coll. 2, Rio Cocles. Banana River. Rio Betey.
327 Costa Rico MiocENE—OLSSON 155
Genus NATICA Scopoli Natica canrena Linnzeus Plato 13, figure 9
Natica canrena VLinneeus, 1758, Syst. Nat., ed. 10, p. 776.
Natica canrena Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, 2nd series, p. 357.
Natica canrena Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., p. 508,
Natica canrena Maury, 1917, Bull. Amer. Pal., vol. 5, p. 298, pl. 23, fig. 10.
This is a recent species but also common as fossil in the Mi- ocene of the West Indies and the States. It is a warm water species so that in the Chesapeake Miocene of the eastern United States, it is found only in the warm, upper or Duplin stage. We collected several specimens at Gatun but none at Costa Rica. Gabb however has recorded it as abundant from the Limon Pen- insula but whether his specimens were obtained .from the Mio- cene or Pliocene is not known.
The species is distinguished by its strong, tangential, su- tural plicee, but the main surface of the whorl is otherwise smooth.
Gatun Stage: Gatun, C. Z. Natica Youngi Maury, var., cocleana, n. var. Plate 13, figure 8
cf, Natica Youngi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 299, pl.
2209S. UT, «k2:
Shell subgiobular with 5 rounded and convex whorls; sur- face smooth but with irregular, coarse growth lines about the su- tures; the surface is also faintly straited with weak spiral lines as may be seen on several otherwise smooth Naticas; aperture semilunar with a narrow callus on the inner lip, slightly thicker about opposite the middle of the umbilicus; umbilicus small, with the umbilical callus small and but poorly developed.
Length or height 25, diameter 34 mm.
156 BULLETIN 39 328
A single specimen from the Gatun of Cocles Creek. It agrees almost exactly with Dr. Maury’s type specimen of Watica Young? from the Miocene of Santo Domingo in size and form, but differs in its smaller umbilicus and smaller umbilical callus. The Costa Rican shell has the aspect of a Veverita,
Gatun Stage; Cocles Creek, near Old Harbor, C. R.
Section STIGMAULAX Moerch Natica Guppyana Toula Plate 13, figures 13, 14, 15
Natica (Stigmaulax) Guppyana Youla, 1909, Jahrb. der K-K Geol. Reichsanstalt, Wien, vol. 58, p. 696, pl. 25, fig. 6.
Natica Guppyana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 360.
This species stands intermediate in its sculpture and form to the VV. canrena Linnzeus and JV. sulcafa Born, both recent and fossil Miocene species. In Guppyana, the sculpture consists of even, wide and deep sulcations, which cross the face of the whorl from the upper suture to the umbilicus. They are in fact the continuation of the tangential sutural plicae of VV. canrena. With further growth these sulcations may become obsolete on the mid- dle of the whorls until they are lacking from the greater part of the shell except in the immediate vicinity of the upper sutures and on the umbilical angle. This change in sculpture is not seen to the same extent on all shells and some specimens may remain strongly grooved and sulcated even when large and mature. In other cases (in general with shells from the Banana River) only very young shells have the sulcation continuous across the face of the whorl, but soon become smooth, leaving the plica- tions only about the upper suture and on the umbilical angle. Such shells resemble closely MV. canrena, but in addition to the grooving of the umbilical angle, have a larger and more exanded aperture.
The WV. sxlcata Born still remains to be found in Costa Rica and Panana. It is abundant: as a Miocene fossil in Santo
329 Costa RicA MroceEne—OLSSON [57
Domingo, Jamaica and Venezuela. It differs from the Guppyana by its closer and more irregular longitudinal sculpture crossed by faint spirals, producing a cancellated appearance. ‘The umbill- cus is larger and more ample.
Gaiun Stage: Gatun, .C. Z. Water Cay, Panama. Banana River, C.F.
Genus POLINICES Montfort
Polinices subclausa Sowerby Plate 13, figures 16, 17
Natica subclausa Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, p. 51.
Natica subclausa Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22\'p. 200, pl. 18, fig. 8. Folinices subclausa Brown and Pilsbry, 3911, Proc. Acad. Nat. Sci. Philia., vol. 63, p. 560. Polinices subclausa Maury, 1917, Bull, Amer. Pal., vol. 5, p. 300, pl. DB ile? TAY This is a common fossil in the Miocene of Santo Domingo and Jamaica. The Costa Rican shells are small, the largest specimen in our collection measuring only 17 mm in altitute.
When viewed from the dorsal side, the shell has a distinct- ly Vertta-like aspect, due to its low spire, close coiling and rap- idly enlarging whorls and appressed sutures. The umbilicus is of medium size, expanding below. The inner lip is provided with a thick, heavy callus which joins above with the outer lip. At the upper edge of the umbilicus, the callus carries a deep, transverse groove, a most characteristic feature.
Gatun Stage: Gatun, C. Z. fiill No. 3, Banana River, C. R. Polinices Stanislas-Meunieri Maury Plate 13, figure 7
Polinices Stanistas-meuniert Maury, 1817, Bull. Amer. Pal., vol. 5, p. 300, pl. 23, figs. 15, 16.
158 BULLETIN 39 330
The Banana River beds contain a large species of Polinices which appears to be this Dominican species. It differs from the recent mammillaris Lamarck, of the Costa Rican and Panana coast by its much narrow umbilicus. Its surface is lined with very fine spiral threads.
Young shells may be mistaken for the swéclausa which oc- curs in the same beds, and careful study is sometimes necessary for their separation. The present shell is less heavy, with a higher spire and more convex spire whorls. The umbilicus is much narrower, anda smaller callus on its inner lip which shows but faintly the transverse groove. The surface of sudclausa is smooth, but in Stanzslas- Meuntert very finely lined with minute spirals.
Gabb has described from Sapote, Costa Rica, a Watica emi- nulordes which in formresembles young shells of the Stazzs/as- Meuntert. His figure shows a different shaped umilicus and no transverse groove on the callus of its inner lip.
Length or altitute 44, diameter 38 mm. Gatun Stage: fiill ra 2, 3, Banana River Goa
Subgenus NEVERITA Risso Neverita nereidis Maury
Neverita nereidis Maury, 1917, Bull. Amer, Pal., vol. 5, p. 301, pl. 23, HSS. eS.
Our collection from ths Lower Gatun of Red Chff Creek, contains a fragment of this very interesting species. The JV. nzeretdis described by Dr. Maury, from the Cercado formation of Santo Domingo, is closely allied to the recent West Coast /V. glauca Humboldt. The Miocene WV. zerezdis is less broad, with a narrower umbilicus and a larger umbilical callus.
Gatun Stage: Coll. 6, Red Clif Creek, (GC.
33% Costa RicA MrocKENE—OLSSON 159
SUB-ORDER SCUTIBRANCHIATA A. SUPER-FAMILY RHIPIDOGLOSSA Genus NERITINA Lamarck
Section SMARAGDIA Issel Neritina viridemaris Maury Plates 15, figure 22
Neritina viridis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 242, Not NV. viridis Linnzeus.
Neritina (Smaragdia) viridemaris Maury, 1917,Bull. Amer. Pal.,, vol. 5, De 316, pl. 24, fig. 11.
The Costa Rican specimens are a little smaller than typical viridemaris from the Miocene of Santo Domingo; but are identi- cal in other respects. They still retain their original coloration of a pale sea-green marked with straight and zigzag black lines. It is the Miocene precursor of the common, recent West Indian and Caribbean WV. viridis Linné.
Gatun Stage: Coll. 5, Red Cliff Creek. Middle Creek.
Genus CREPIDULA Lamarck
Crepidula plana Say Crepidula plana Say, 1822, Journ. Acad. Nat. Sci. Phila., vol. 2, p. 226, Crepidula plana Dall, 1892, Trans. Wagner Free Inst. Sci., vol. 3, pt. 2, p.. 358. Crepidula plana Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 360.
This common and well-known species has been recorded by Brown and Pilsbry from the Canal Zone. We have a single spec- imen from Water Cay, found in the interior of the specimen of Solenostetra Vaughant var. medioamericana. ‘This species first
160 BULLETIN 29 342
appears in the Lower Miocene and has continued into the re- Cent) fatinar
Gatun Stage: Gatun (Brown and Pilsbry).
Genus XENOPHORA Fischer de Waldheim Xenophora conchyliophora Born
Trochus conchyliophorus Born, 1778, Mus, Caes. Ind., p. 333.
Trochus agglutinans Lamarck, Anim. sans Vert., vol. 7, p. 14.
Phorus agglutinans Gabb, 1873, Trans. Amer. Phil. Soc., vol, 15, Diy24r.
Xenophora conchyliophora Dall, 1892, Trans. Wagner Free Inst. Sci., VOl. 3, pt. 2, pp: 360-262.
NXenophora conchyliophora, Maury, 1917, Bull. Amer. Pal. vol. 5, p. 297.
One immature shell was collected from the upper Gatun beds of Old Man Sam Creek. It is one of the few species which has continued pratically unchanged from Upper Cretaceous times to the recent. It is rare asa recent shell along the northern Panama and Costa Rican coast. Gatun Stage.
Genus PHASIANELLA Lamarck © Phasianella moilis, n. sp.
Shell small, solid, ovate with a short, blunt spire anda small narrow umbilicus; whorls about 4, somewhat convex; su- ture distinct; surface smooth and usually showing no features; in some cases, color markings are preserved as brown, wavy blotches of flammules and with very faint suggestions of fine re- volving lines which are so common on recent small Phaszanella.
Height 3.25, diameter 2.75, aperture 1.50 mm.
The Phasianclla punctata Gabb from the Miocene of Santo Domingo, has a longer spire and different color markings. It is much more like the recent ?. affinzs than the Costa Rican mollzs. P. mollis is related to P. umbilicata d’Orbigny but differs in its sculpture.
333 Costa RicA MIOCENE—OLSSON 161
The smaller forms of Phastanedlla, of which there are 2 or 3 species along the northern Costa Rican coast, are often extreme- ly abundant on the rocks and reefs exposed to the heavy, pound- ing surf. There they are associated with a host of other gas- tropods, principal among which are several species of Verzta, fissurella, Purpura and Chitons. The fossil shells therefore like the preceding Veritina viridemaris and several others which we have already called attention to, have drifted from some near-by rocky coast.
Gaiun Stage: Coll. 5, Red. Cliff Creek. Middle Creek.
Genus TURBO Linnzeus Turbo saltus, n. sp. Plate 15, figure 12
Shell perforate, ovate-conic; spire high of about 6 convex whorls, separated by deep, canaliculate sutures; the last whorl is large and perfectly convex in form; sculpture consists of irregu- lar, spiral cords or lirze; the earliest spire-whorls are strongly carinated by a single, strong spiral, above and below which the whorl is smooth; a second spiral appears above and the succeed- ing whorls of the spire have 2 principal spiral cords or lire, and g smaller ones; the last whorl has three spirals that are a little stronger and many smaller ones of different sizes; the spirals are smooth; aperture perfectly circular, a small, spreading callus on the body-whorl just above the perforate base.
Height 17, diameter 15.5, aperture 7.5 mm.
Among the recent species of 7urbo the salfus is related to the large 7. Spenglerianus Gmelin,on one hand and to the small- er 7. filosus Kiener on the other. The shell is perforate,in which character it approaches 7: /flosus, but the sutures are canalicu- late or channeled as in the non-perforate 7. Spenglerianus. The sculpture consists of irregular, smooth spiral cords or lire of which 2 or 3 about the middle of the shell are a little larger than the others.
162 BULLETIN 39 334
Gatun Stage: Coll. 5, Red Cliff Creek.
Genus ASTRALIUM Link
Astralium brevispinum Lamarck, var. basalis, n. var. Plate 15, figs. 4, §
The Astralium brevisphinum Lamarck is a recent species found living along the north coast of Panama and Costa Rica. It is most easily distinguished from the West Indian A. longzspr- num Lamarck by having its columellar area brilliantly stained with scarlet.
From upper Old Man Sam Creek in eastern Costa Rica, we collected a single specimen of an Astralzum, very similiar in most of its characters to drevespinum. It differs most importantly in having a more strongly sculptured base. This sculpture consists of a wide, flat band, inside of which are 3, heavy, nearly equal spiral cords. The whole surface is crossed by squamose lines of growth.
Height 20, diameter 29.50 mm. Gatun Stage: Coll. 6, Old Man Sam Creek.
Astralium caelatum Gmelin, variety
Trochus celatum Gmelin 1792, Syst. Nat., p. 3581.
Astralium celatum Pilsbry, 1888, Manual of Conch., first ser., vol. Io, p. 224, pl. 57, figs. 45, 46.
A large Turboid shell related to the recent 4. celatum Gme- lin was collected from the coral limestones near Port Limon. It is so firmly imbedded in its rocky matrix that only a smal! por- tion of its surface isexposed to view. Its surface sculpture is like that of celatum, but the radial costze on the upper part of its whorl are closely spaced and not far apart as in the recent shell.
ts height measures 55 mm, diameter 54.
Gatun Stage: Port Limon.
335 Costa Rico M10ocENE—OLSSON 163
Genus CHLOROSTOMA Swainson Subgenus NEQMPHALIUS Fischer
Chiorostoma costaricensis, n. sp. Plate 15, figure 16
Shell conoidal, solid, witha narrow, but deep umbilicus; su- ture distinct; the last whorlis strongly carinate or angled below; above this angle, the whorl is evenly convex, below, the base is nearly flat; the sculpture consists of beaded spiral cords, there being on the last whorl above the angled periphery 7 or 8 cords; the interspaces are nearly as wide as the cords, and are finely etched by oblique lines of growth and finer spirals; on the base, the spirals are irregular, the strongest about the middle; there are 2 or 3 cords about the upper part of the umbilicus, but the interior wall is smooth.
Height 10.5, diameter 16.5 mm.
The transitional beds between the Uscari and the Gatun of Red Cliff Creek, contain in addition to the usual fauna, several species, as certain Cerithoids, Szstrum nodulosum, Modulus mo- dulus, Nitidella and the present species, whose normal habitat is on the rocks and reefs of the extreme, upper limit of the lit- toral zone. They are always much broken and worn as if they had been rolled and washed about by the surf on a rocky or sandy beach and then drifted out to the deeper and quiter waters off shore where depostion was going on.
Although all of our specimens of C. costaricensis are very fragmentary and much worn, we have deemed it best to place the species on record, as this subgenus has been recorded only very doubtefully from beds older than the Pleistocene.
Uscart Stage: Zone 6, Red Cliff Creck. Gatun Stage: Zone 5, Red Cliff Creek.
164 BULLETIN 39 336
Genus CALLIQSTOMA Swainson
Calliostoma limonensis, n. sp. Plate 15, figures 8, 1
Shell of medium size, conic; whorls about 8, flat or but | slightly convex, so that the profile of the shell is a broad cone; |
sutures indistinct; the whorls are closely sculptured with fine, beaded spirals of primary and secondary orders; each spire-whorl has 7 or 8 primary spirals alternating with 1, 2 or 3 secondary; periphery sub-angular; base but slightly convex and closely spir- alled with flat or sub-obsoletely beaded spirals (about 20),umbil- icus narrow but deep, smooth within.
Height 19, diameter 18.5 mm.
Fairly abundant in the coral limestones along the shore west of Port Limon. Related to the recent C. jujubinum Gmelin, it differs by its more globose form and small, pointed attenuated apex.
Gabb hasalso described a small Calliostoma from the Limon Peninsular, as C. Guppyana. This shell of 8 or 9 whorls meas- ures but 5 mm in height. Itis a higher and more conic form with the whorls sculptured with beaded spirals.
Gatun Stage: Port Limon.
Calliostoma mancinella, n. sp. Plate 15, figures 9, 10
Shell of medium size; whorls about 8, with very indistinct sutures and a coarse sculpture of beaded spirals; the spire whorls are flat sloping evenly from the small projecting nucleus to the broadly rounded periphery of the last whorl; sculpture of the spire-whorls consists of about 6, strongly beaded spirals the largest of which boundsthe upper sutures; between these spirals are scattered a few smaller ones, the lower half of the peri- phery and the base have about 12 additional spirals; these spirals are larger, more widely spaced and are smooth except the 4 or 5 around the umbilicus; the interspaces of the basal spirals are finely incised by minute longitudinal lines, corres- ponding to the growth lines; umbilicus narrow, but deep,
337 Costa RrcA MIocKENE—OLSSON 16
mn
smooth within; aperture rounded, with the outer lip strongly oblique.
Height 15, diameter 21 mm.
Several specimens of this well-marked species were obtained from the Gatun beds of eastern Costa Rica, in the vicinity of Manzanilla Point. Inthe Santo Dominican Miocene it is relat- ed to the C. Grabaut Maury, but the Costa Rican shell is much lower and more depressed.
The C. Grabauz and the mancinella belong to the group which contains the large, elegant C. Sayana, described by Dal! from deep water off the Hatteras coast.
Gatun Stage: Coll. 4, East Grape Point Creek. Fleadwaters of Middle Creek. Coll 6 ,hed\ Clay” Creek: Z mtle south of shore, along Old Man Sam Creek, (Veatch)
Calliostoma castilla, n. sp. Plate 15, figure 6, 7
Shell rather large, in form and sculpture resembling the C. philanthropus of the Chesapeake Miocene of eastern United States, but umbilicated; whorls 5 plus (apex broken), flat and forming a broadly conic spire; periphery of the last whorl, subangular, due to a broad, cord-like ridge, which appears as if formed form an overlapping of the base; sculpture of the spire-whorls of fine, alternating and beaded spirals, there being about 13 on each whorl; the base is nearly smooth the spirals being nearly obso- lete or as broad low bands; the spirals about the umbilicus are slightly stronger and obscurely beaded; umbilicus deep and narrow.
i Height 20, diameter 25 min.
A large species with an angulated periphery, fine, beaded spirals above and a nearly smooth base. Buta single specimen
166 BULLETIN 39 33%
was collected from the coral limestones near Port Limon, asso- ciated with the common /émonensts.
Gatun Stage; Port Limon.
CLASS SCAPHOPODA Genus DENTALIUM Linnzeus
Subgenus DENTALEUM, s. s.
Dentalium bocasensis, n. sp. Plate 15, figures 2, 3
Shell rather larger, solid, gently curved and increasing grad- ually in size; the tip is more curved than the rest of the shell and strongly hexagonal in cross-section; the sculpture consists at first of 6 strong, primary ribs and wide, flat or slightly concave in- terspaces; the ribs are further increased by the introduction of 6 intermediate ribs and still later by 12 more; these ribs are sub- equal in size on the later portions of the shell; the interspaces are finely transversely lined and faintly longitudinally striated,
Length 52, diameter 6.25 ‘mim, tH) 4-75 mm.
The Dentalium bocasensis is fairly abundant in the shale beds of Bocas Island. It is recognized by its strongly hexagonal tip and finely transversely lined interspaces,and strong longitudi- nal sculpture.
Gatun Stage: Bocas del Toro.
Subgenus FISSIDENTALIUM Fischer Dentalium uscarianum, n. sp. Plate 15, figure I
Shell large, solid and heavy; gently curved, and the taper of the shell is slow and gradual; the tips are all broken on our specimens, but the later sections of the shell are circular or slight- ly compressed dorsal-ventrally; the surface is sculptured with about 24, low, subregular and rounded longitudinal ribs with
339 Costa Rica MrocknzE—OLsson 167
smaller intermediate ones in the interspaces; the ribs become gradually obsolete with age and the later portions of full-grown shells are nearly smooth; fine transverse lines of growth cross the surface and are best seen on the earlier portions of the shell; operture oblique.
Length 60, diameter 12 mim.
This large Dentalium is abundant and very characteristic of certain conglomeritic zones in the Upper Uscari shales. There it occurs associated with a small but very distinctive fauna, includ- ing the Pétychosalpinx? dentalis, Sconsia cocleana and Scaphella costaricana and other species. When complete, the Dentalium uscarianum frequently reached a length of 110 mm or more.
All our specimens are unfortunately lacking in the apical tip. In general sculpture, it recalls the Dentalium floridense Henderson dredged from 35 to 110 fathoms of water off Florida and in the West Indies, but the taper of the shellis more gradual and the sculpture more irregular, becoming obsolete with age. Bose* has described a large species as Dentalium rimosum from the Miocene of the Tehuantepec Peninsula. In that species, the ribs number about31, are very regular with deep interspaces, and ribs are finely etched by transverse lines of growth.
Uscari Stage: Coco Plum, Panama.
Rio Cocles.
Margarita—Old Flarber tratt. Comadre Creek.
Pumbri Creek, ete.
*Boll. Inst. Geol. do Mexico, No. 22, p. 55, pl. 3, figure 1, 1906.
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BULLETINS OF
AMERICAN PALEONTOLOGY
Vol. 9g
No. 39
The Miocene of Northern Costa Rica
WitH Notes ON Its GENERAL STRATIGRAPHIC RELATIONS BY A. A. OLSSON
Part 2 CLASS PELECYPODA
(Pp. r69-Finis)
fune 21, 1922
Harris Co. Cornell University, Ithaca, N. ¥. Wis. A:
NC Mee
343 Costa Rica MiocENE—OLSSON 171
ORDER PRIONODESMACEHA
-Genus NUCULA Lamarck Nucula orbiceila, n. sp. Plate 28, figures 19, 20
Shell of medium size, broadly elliptical or subovate in out- lines; obliquely truncated at its anterior end; gently convex; the surface is covered with close, subregular, concentric wrinkles and fine radial threads so that the sculpture is decussated; on the posterior dorsal slope, they are larger and pustular-like and their course 1s obliquely downward across the growth-lines; they are weak on the poorly defined lunule; the radical threads are main- ly between the concentric wrinkles but ventrally they cut into the wrinkles as well; interior of the shell, silvery white or faint- ly tinged with purple; the ventral margin is finely crenulated; chondrophore very small, oblique; anterior set containtng about 8 teeth, the posterior with about 17.
Length 9.50, height 7.50, semi-diameter 2.25 mm. II.00 8.50 2.50 mm.
From the recent WV. crenulata A. Adams, this shell differs by its larger size and more elliptical and depressed shell. Gabb has described from the Pliocene of the Limon Peninsula WVu- cula limonensts and moenensts, but both of these species are per- fectly smooth and sculptureless. The Nucula orbicella is a com- mon species in certain marly beds intercalated in the coralline limestones of Port Limon.
Gatun Stage: Port Limon. Nucula cahuitensis, n. sp. Plate 18, figures 21-24
Shell small, solid, obliquely subtrigonal in form; ventral margin widely rounded, a nearly straight posterior dorsal margin
172 BULLETIN 39 344
and a widely truncated anterior end; vaives gently convex; pro- toconch on the tip of the umbo small, smooth and white; surface polished; the sculpture of the disk of the shell is developed to a variable extent, but the umbos are always smooth; typically the lower half of the shell is covered with close, regular, concentric wrinkles which are slightly decussated by fine, even, and slight- ly raised radial threads; in other cases, the wrinkles may be pres- ent only on the extreme anterior or posterior ventral extremities and the radial lines showing only as a part of the shell substance and not as raised threads; the escutcheon is elliptical and thickly covered with raised wrinkles which lie at right angles to the dor- sal margin of the shell; they are similiar on the lunule; interior of the shell tinted with brown; ventral margin crenulated; chon- drophore small, oblique with the anterior set of teeth numbering about 7, the posterior about 14.
Length 3.50, height 3.00, diameter 1.60 mm.
This is a small species of subtrigonal outlines, suggestive of the Pacific V. exigua Sowerby in general form and size. On the few specimens of V. exivwa, which I have from the Bay of Pan- aina the sculpture is finely decussated over the whole shell with fine radial and concentric threads. The most striking features of cahuttensts are its strongly sculpture lunule and escutcheon.
Gatun Stage: Zone G, Saury Creek.
Nucula tenuisculpta Gabb Plate 18, figure 25 Nulua tenuisculpta Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 255. Nucula tenuisculpta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 325, pl. 26, figure 8. :
A species of the size and forin of cahuztensts but the valves are entirely smooth and polished. The lines of growth are fine and irregular. The radial lines are faint and sub-microscopic. Inner margin crenulated.
Height 3.25, length 4.00, semidiameter .75 mm. Gatun Stage: Coll 4g, Red Cliff Creek.
345 Costa RicA MIocCENE—OLSSON 173
Genus LEDA Schumacher
Leda Balboae Prown and Pilsbry Plate 18, figure 15
Leda Balboe Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., WOO 2.up. 302; )pl. 27, fig..8-
This species is fairly common in Costa Rica, but generally smaller than in the Canal Zone, where it frequently reaches a length of 22 mmor more. Its valves are moderately convex and regularly sculptured with even, close, concentric riblets which on the center of the shell disk may be slightly appressed. The escutcheon is well-defined and similiarly sculptured with the rib- lets running parallel to the hinge margin.
Cagmn Stage: Gatun, C. Z. Coll. 1, East Grape Potnt Creek. Zome G, Saury Creek. Ourtana Creek. fiill No. 3, Banana River.
Leda Davilae, n. sp. Plate 18, figure 20
Shell small, solid, strongly convex; the beaks nearly cen- tral; the rostrum is short, pointed and sharply defined from the rest of the shell disk by a high, cord-like ridge; the anterior end is bluntly rounded and carries 2, low flexures from the beaks to the anterior ventral extremity; lunule small and narow, lacking the concentric ribs and sculptured with small, pustular-like ridges passing at right angles to the hinge-line; the sculpture con- sists of about 30 concentric riblets which are somewhat larger and with deeper interspaces on the umbos; the interspaces show on the umbos and along the anterior slope, fine, radial lines or punctations; interior of the shell cavity deep, with a heavy hinge; the anterior and posterior sets of teeth number about 13 or 14, are high and V-shaped.
Length 5.75, height 4.00, semi-diameter 1.75 mm.
174 BULLETIN 39 346
A small, plump species, coarsely sculptured with even, con- centric riblets, separated by deep interspaces. These interspaces may carry fine, radial lines or punctations at the base of the con- centric ribs and are best seen on the umbones and the anterior slope. The ZL. indigina Dail (L. dzsulcata Guppy) from the Miocene of Jamaica is similiar but proportionately longer.
Gatun Stage: Coll. 4, Red Cliff Creek. Bocas Island.
Subgenus ADRANA, H. and A. Adams Leda quitanensis, n. sp. Plate 18, figure 19
Shell thin, elongate but slightly convex; beaks very low and scarcely distinguishable from above and situated at the anterior 1-3; ventral margin an even curve from the pointed extremities; dorsal margin straight; surface polished but covered with very fine, concentric lines which are crossed on the lower half of the anterior 2-3, by even, oblique lines (Sczssuda-like); these oblique lines are spaced about .25 mm apart; interior concealed.
Height 7.25, lenght 26.00, semidiameter .75 mm.
A much smaller species than the next (ezsnozdes), slightly more convex and with beaks situated more anteriorly. The sur- face is finely sculptured about the middle and the anterior por- tion by slightly oblique lines, as seen on certain Tellinoids such as Scissula. The Manzanilla beds of Trinidad contain Leda (Adrana) Guppyi Dall (Cercomya ledeformis Guppy). It is less elongate, more contracted posteriorly and has a different surface sculpture. Dall has also recorded Leda Guppyi from the Bow- den beds of Jamaica.
Gatun Stage: Outtana Creek. Zone F, Saury Creek. Coll. 1, East Grape Point Creek.
Leda ensincides, n. sp. Plate 18, figure 12
Sheli rather large, elongate, depressed and subequilateral;
347 Costa RicA MIOCENE—OLSSON 175
beaks very small and scarcely distinguishable from above, and situated nearly about the middle of the dorsal margin; extremi- ties bluntly rounded, with a broad, even curve to its base or ven- tral margin; the dorsal side is nearly straight but descending slightly towards its extremities; surface nearly smooth, the growth lines indistinct, but with fine, even very slightly oblique lines on the anterior two-thirds of the shell; interior con- cealed.
Length 54.00, height 15.00, diameter 4.50 mm.
Differs from the preceding species by its larger size, nearly central beaks and by its finer, more concentric sculpture. There are several species of this group recent in the West Indies and along the Pacific coast of Central America, but none appear at all closely related to the Gatun species.
Gatun Stage: Hill No. 3, Banana River.
Leda Dalliana, n. sp. Plate 28, figure 17
Adrana sp ? Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 1 4, Pp. 592.
Shell small, thin, elongate, depressed; the small, scarcely differentiated beak is sutuated about the anterior one-fourth of the shell; the anterior side is somewhat contracted and shortly rounded while the posterior side is produced and 3 times the length of the anterior and bluntly pointed at its extremity; the rostrum is long and narrow, formed by 2 radial, scabrous threads, separated by a wide interspace; concentric sculpture of thin, elevated threads or lamellae, regularly distributed on the lower half of the shell, but widely spaced on the umbonal area; the escutcheon is long and narrow, sculptured with threads parallel to the hinge-margin; interior of shell cavity shallow, the long hinge with an anterior set of 20 small teeth, the posterior with 26 OL 27.
Length 10.50, height 3.50, diameter of left valve .75 mm.
176 BULLETIN 39 348
A much smaller species than the preceding and differing in its more anterior beaks, depressed shell and heavier concentric sculpture as well as in other characters. It is probably the shell which Dall refers to as having fragments of from the Pliocene of Limon. But a single perfect specimen was collected
Gatun Stage; Port Limon.
Genus YOLDIA Moller Yoldia bocasensis, late 28, figures 21, 22
Shell small, thin, ovate-elliptical in form; beaks situated shghtly in front of the middle; valves moderately convex; both the anterior and posterior extremities are broadly rounded and counected by the gentle curve of the basal or ventral margin; the umbos are very small but distinct and he just above the greatest convexity of the valves; surface smooth, polished with no other sculpture than the incremental growth-lines; chondrophore dis- tinct, scarsely oblique and bordered on each side by an equal num- ber of V-shaped hinge-teeth; the anterior set contains about 14 teeth, which are very small next to the chondrophore becoming progressively larger distally, the posterior set with 13 or 14; the pallial sinus is indistinct, but rather large, broad and extends nearly to the middle of the valve.
Length 13.00, height 7.00, semidiameter 2.25 mm. 9.50 5250 1.50 mm.
A fairly common species in the clay beds of Bocas Island where it is associated with a fauna of rather deep water charac- ter. It has much the form of certain deep-water Yo/dzella but the interior of the valves show an indistinct but deep and wide pallial sinus.
Gabb has described from the Miocene of Santo Domingo, a Yoldia ovalis, which has never been figured and its description is very meager and insufficient.
Gatun Stage: Bocas del Toro.
349 Costa Rica M10ocENE—OLSSON 077
Genus LIMOPSIS Sassi Limopsis monilis, n. sp. Plate 26, figures 23, 24
Shell small, solid, moderately convex and subcircular in form but slightly oblique; the hinge-line is straight, the anterior dor- sal submargin very small, the posterior large undifferentiated from the rest of the shell ; the umbos are high and prominent, lying a little in front of the middle; exterior sculptured with strong radial ribs which are largest and strongest on the middle of the shell disk, finer on the dorsal submargins; these ribs are crossed by high widely spaced concentric lamellae, more closely spaced on the umbos; the lamellze on crossing the ribs are raised into short, spine-like fringes which are directed ventrally; inter- ior of shell deep; the hinge-line with a small cartilage pit at the anterior 1-3 of the shell; it is bordered on the anterior side by a set of 3 high, projecting teeth and 4 on the posterior side, the ventral and a part of the anterior margin is crenulated, most fine- ly in the middle and quite coarsely at the posterior portion.
Length 1.75, height 2.00, semidiameter .60 mm.
This is a small, subcircular species with a crenulated inner margin and a strong surface sculpture of ribs crossed by distant concentric fringes. It is fairly common at its type locality, giv- en below.
Gatun Stage: Old Man Sam Creek, 1 mile south of beach (A. C. Veatch, collector).
Genus GLYCYMERIS, Da Costa Glycymeris canalis Brown and Pilsbry late 18, figures 2-7
Glycymeris canalis Brown and Pilsbry, 1IgtI, Proc. Acad. Nat. Sci. Plata: vol, 63, p: 364, pl. 28; fig: 10,
Glycymeris trilobicostata Pilsbry and Brown, 1917, Proc. Acad, Nat. Sci. Phila,
Very abundant locally in the Gatun beds of Costa Rica, are
small and medium sized Pectunculids belonging to the group of
178 BULLETIN 39 350
G. acuticostata Sowerby of the Santo Dominican Miocene. They vary somewhat in size, (20 to 32 mm in height), are usually de- pressed and with an outline suggestive of the recent Hast Coast and Upper Miocene G. fectinata Gmelin. Typical acuticostata does not seem to occur in Costa Rica and I have adopted Brown and Pilsbry name of canalzs.
The G. canalis Brown and Pilsbry from the Canal Zone, and the /rilobtcostata Pilsbry and Brown from Colombia, are both bas- ed on young shells 15 mm or less in height. Both forms can be duplicated amongst the Costa Rican shells and are but varieties of the same species.
In its usual and more abundant form, the shell is depressed, with numerous (about 26), closely spaced rounded or angled ribs. These ribs are overrun by even, close, concentric threads. They represent the usual cazadzs or its variety ¢rilobicostata, and aver- age in height about 25 mm. More rarely, the shells are larger and as illustrated by figure 4 and 5 the ribs may become divided into 2 or 3, strong, radial cords. The following measurements will show the usual range in size.
Height 22, diameter 21, thickness 5.5
24 23 5:5 a2 32 8 29 29 8
Gatun Stage: Gatun, C. Z. | Comadre; Banana River; Coll. 4 East Grape Point Creek. Old Man Sam Creek, C. R.
Glycymeris jamaicensis Dall Plate 19, figures 3, 4
Pectunculus pennaceus Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22, p, 293. Not of Lamarck.
Axinea pennacea Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 255.
Pectunculus decussatus Guppy, 1876, Qart. Journ. Geol, Soc. London, vol. 32, p. 532. Not of Linné.
OLSSON 118)
351 Costa RicA MIOCENE
Glycymeris jamaicensis Dall, 1898, Trans. Wagner Free Inst., Sci., VOL. 3,, pt: 4,\ p., 608.
Glycymeris jamaicensis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 345, pl:)26, fig.) 13:
This large Pectunculid, externally resembles in its angular posterior side the Glycymeris pennacea with which it was confus- ed by Guppy and by Gabb., It differs in being somewhat less con- vex, less strongly sculptured and most importantly in its central beaks, located about the middle of the ligamental area. It is, on the other hand, much more closely allied with the recent West Indian G. wndatus Linné (G. limeatus Reeves) of which it is doubteless the Miocene ancestral form, the main difference being in details of its finer sculpture.
The Glycymeris jamaicensis occurs in the Miocene of Jamaica and Santo Domingo but there never seems to reach the large size that the species sometimes attains in Costa Rican. A large shell in our collection from Old Man Sam Creek measures as fol- lows:
Height 60, lenght 60, diameter of both valves 37 mm.
Gatun Stage: Water Cay. Cote 5. Rear Glif Creer: Coll, 4a, East Grape Point Creek. Colls. 5, 6, Old Man Sam Creek. Soust Creek of Upper Hone Creek.
Glycymeris carbasina Brown and Pilsbry Plate 19, figures I, 2, 5
Glycymerts carbasina Brown and Pilsbry, 1g11, Proc. Acad. Nat. Sci. Phila.) vol63; p: 363, pl. 28). fig: 9;
The Glycymerts carbasina was described by Brown and Pils- bry from a small individual from Gatun, measuring only 16 mm in height. It is closely related to the G. jamatcensts Dall and probably represents but a varietal form. Its main differences is its more circular and less convex shell, and less number of hinge- teeth which number about roto 12 on the anterior and posterior sides respectively.
180 BULLETIN 39 352
All our specimens were collected in the lower Gatun of E. Grape Pt., Cliff Creek in Eastern Costa Rica, the largest shell measuring as follows:
Height 56, length 55, diameter of both valves 32 mm. Gatun Stage: Coll. 4, East Grape Point.
Glycymeris castaneus Lamarck Plate 18, figures 13, 14 |
Fectunculus castaneus Lamarck, 1819, Anim. sans Vert., vol. 6, p Gee
Fectunculus castaneus Reeve, 1843, Conch. Icon., Fectunculus pl. 6, figs 32)
Our collection from the Banana River contains several small Pectunculids which differ from the jamazcensts and its allies by their less circular shell, greater convexity and coarser radial sculpture. These shells agree however very closely with a recent species which is abundant on the beaches of northern Costa Rica, which appears to be the Glycymeris castaneus of Lamarck. Glycymeris castaneus is distinct from undutus Linné (G. lineatus Reeve) with which it is sometimes united, differing by its form which is more produced posteriorly but not angulated and by its more pronounced radial costz. Both species are similiarly col- ored with chestnut markings.
Gatun Stage: Hill No. 3, Banana River.
Quttana creek.
Glycymeris decussatus Linnzeus Plate 19, figures 6, 7
Arca decussatus Linneeus, 1758, Syst. Nat., ed. 10, p. 694. Fectunculus pennacea Vamarck, 1819, An. s. Vert., vol. 6, p. 51. Pectunculus pennacea Reeve, 1843, Conch. Icon., Fectunculus, pl. die 42a Fectunculus decussatus Vinné. P. pennaceus Wamarck, (Lamy) 1911, Journ. de Conchy., vol. 59, p. 119, pl. 3, fig. 7. Of this rare shell, more commonly known as the G. penna- cea Lamarck we have several excellent specimens collected by
353 Costa RicA MIocCENE—OLSSON 181
Dr. A. C. Veatch from the Upper Gatun of Old Man Sam Creek in eastern Costa Rica and a few specimens from the coral-reef limestones of Port Limon. Although a very distinct species and not to be confused with any other Pectunculid recent or fossil, its characters have been generally misunderstood and its name has often been applied to shells of the G. /ineatus group.
The shell is of moderate size, strongly convex and inequila- teral. The beaks are slightly posterior of the middle, with the ligamental area entirely anterior to the beaks as seen in figure 7. The surface is sculptured with numerous, moderately coarse, radiating threads which are simple on the umbonal area but be- come divided by 3 or more finer threads ventrally.
Gatun Stage: Old Man Sam creek. Port Limon.
Glycymeris Lloydsmithi Pilsbry and Brown Plate 25, figures 8. 9, 10
Glycymeris Lloydsmithi Pilsbry and Brown, 1917, Proc. Acad. Nat. Sci. Phila., vol. 59, p. 39, pl. 6, fig. 6.
This is an abundant fossil in eastern Costa Rica, where it frequently forms zones in the lower and middle Gatun. Itstype of sculpture of broad, smooth, rounded ribs, is strikingly like that of G. subovata Say of the Miocene of eastern United States. It differs in its higher and narrower umbos and by its nearly smooth cardinal area.
The G.Lloydsmitht was described by Pilsbry and Brown, from beds equivalent to the Gatun, near Cartagina, Colombia.
WGawn stage. Zone > Last Grape Point Creek, C. &. Goll ign ed Chg Oreck. C. Fe. fleadwater of Middle Creek, C. R.
Genus ARCA Linnzus Arca occidentalis Philippi Plate 22, figure I
Arca occidentalis Philippi, 1847, Abbild. u. Beschr., 3, p. 14, pl. 17b, fig. 4a-c.
182 BULLETIN 39 354
Arca noe Guppy, Journ. Geol Soc. London., vol. 22, p. 293.
Arca occidentalis Sheldon, 1916, Paleont. Amer., vol. 1, p. 8, pl. 1, figs, 8-11.
Arca occidentalis Maury, 1917, Bull. Amer, Pal. vol. 5, p.327, pl. 29, hig.’ 3.
This isa common recent species of the West Indian and Caribbean fauna. Asa Miocene fossil it is found in Jamaica and Santo Domingo and it is here recorded from the Miocene of Costa Rica.
Gatun Stage: Port Limon.
Arca umbonata Lamarck Plate 22, figure 2
Arca umbonata Lamarck, 1819, An. s. Vert., vol. 6, p. 37.
Arca imbricata Gabb, 1873, Trans. Amer. Phila. Soc., vol. 15, p. 254, In part.
Arca umbonata Sheldon, 1916, Paleont. Amer., vol. I, p. 8. pl. 1, figs. 12-17.
Arca umbonata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 327, pl. 30, 10g tC
Arca mubonata Maury, 1920, New York Academy of Science, Sci. Sur. OLE. URE wot 2 pts wpenO!
The Arca umbonata and the preceding occidentalis area com- mon as recent shells along the entire northern coast of Panama and Costa Rica. The A. umbonata is recognized by its more convex and angular shell and finer sculpture of the middle of the shell disk. Asa Gatun fossil we have but a single small shell from the Kast Grape Point Creek.’ Dr. Maury records the species from the Miocene of Santo Domingo and from Porto Rico.
Gatun Stage: Coll. 1, East Grape Point Creek. Subgenus BARBATIA (Gray) Adams
Arca Mauryae, n. sp. Plate 22, figure 4, 7
Barbatia cf. Bonaczyi Maury, 1917, Bull. Amer. Pal., vol. 5, p. 329, pl. 30, fig. 15. Not of Gabb.
355 Costa RicA MrocKENE—OLSSON 183
Shell small, thin, convex, subrhomboidal in form; umbos wide and full; with the beaks at the anterior 1-3 of the shell; a — rounded posterior umbonal slope; the anterior extremity is brok- eninthe type specimen but from the earlier growth lines is simply rounded in form; the posterior side is obliquely truncate and meets the dorsal margin in a well-marked angle; the exteri- or is sculptured like that of Arca barbata, with the riblets group- ed in wide, regular bands which are defined by slightly deeper interspaces; the radial riblets are crossed and crudely granulat- ed by concentric threads; the sculpture on the posterior submar- gins is much heavier and predominantly radial that of the an- terior submargin is but slightly heavier than on the middle of the shell disk; interior of shell cavity deep, with astraight hinge- margin, the teeth arranged asin édarbata; cardinal area rather high, longitudinally striated posterior of the beaks, plain in front. |
Length 24? height 11.50, semidiameter 5.50 mm.
This species is closely related to the recent Avca barbata Linné. It differs from darbaza in its proportional shorter shell, greater convexity, less anterior beak and higher cardinal area. It is equivalent to the Dominican shell figured by Dr. Maury as Barbatia cf. Bonaczyt Gabb. Gabb in his remarks following his his description of Bonaczyz, compares his shell with darbata, and states that in Bonaczyz, the beaks are more anterior and the pos- terior end more produced and sloping. This is the reverse of the conditions as seen in the present shell. Dall on the other hand, considers Gabb’s shell as synonymous with Avca uwmbonta Lamarck.
Gatun Stage: Port Limon.
Section CALLOARCA Gray
Arca cf. candida Gmelin Plate 22. figures 5, 6
Arca candida Gmelin, 1792, Syst. Nat., p. 3311.
Barbatia (Calloarca) candida Dall, 1898, Trans. Wagner Free Inst., Sci. vol. 2.) pt. 4, p. 626.
184 BULLETIN 39 256
'
Arca cadida Shelton, 1916, Palaeont. Amer., No. 1, p. 16, pl. 3, figs. II, 42.
We have several small valves of a Calloarca from the Mio- cene limestones of Port Limon, the largest of which has a length of i mm. They belong to the Avca candida group and may possibly represent the young of that species. They seem to dif- fer in being more finely sculptured, especially about the middle of the shell disk. This sculpture consists of fine, beaded or granulated radial threads, which become coarser on the anterior and posterior submargins.
The Arca candida is recorded by Dall from the Bowden beds of Jamaica, the equivalent of the coral limestones of Limon. Dall also mentions from the same locality, the occurrence of a smaller and possible distinct species. The Avca candida is a common re- cent species of the West Indies and the Caribbean, and is abund- ant on the north coast of Panama and Costa Rica.
Length 18, height 11?, semidiameter 3.75 mm. Gatun Stage: Port Limon.
Subgenus SCAPHARCA Gray Arca dariensis Brown and Pilsbry Plate 22, figures 10-13
Arca oronlensis Dall, 1898, Trans, Wagner Free Inst. Sci., vol., 3, pt. 4, p. 658. Not of Gabb.
Arca dariensis Brown and Pilsbry, 1gt1, Proc. Acad, Nat. Sci. Phila., Vol. 63. p. 362, pli.22) fig” ao.
Arca gatunensis Toula, Dec. 1911, Jahrb. der K-K Geol. Reichsanstalt, vol,.61, p. 493, plio, fig, *4-
The common Ark of the Canal Zone. The shell is elongate in form with about 30 ribs. The ribs of the left valve are usually granulated over the whole shell, while those of the right valve are narrow and smooth on the middle of the shell disk. The ribs are characteristically divided on the anterior and posterior ends of the shell.
Gabb has described an Arca oronlensis from the black shales
HY
357 Costa Rica MIOCENE—OLSSON 185
of Oronli Creek in the Talamanca Valley (probably the Uscari shell). Itis like A. dariensis, in form, but with more central beaks and umbones and simple ribs. We have not seen this species. Gatun Stage: Gatun and Mt. Hope, C. Z. Water Cay.
Arca actinophora Dall Plate 23, figs..7,8; Plate 25, fig: 3
Scapharca (Scapharca) acitnophora Dall, 1898, Trans. Wagner Free Ist) Se1.. vol! 3. pt. 4) p. 647, pl: 33,-fig.: 26.
Scapharca (Scapharca) actinophora Sheldon, 1916, Paleont. Amer., VOLT, p. 50, pl. 11, fig. 13.
A larger species than the darvzensts, with broad, wide umbos. The ribs number 36 to 40, arenarrow and separated by inter- spaces of a little more than their own width. These interspaces are generally finely ornamented by regular, even, concentric threads. The ribs are simple and on the left valve, usually strong- ly beaded or granulated throughout, becoming wider and smooth- er on the posterior portion, or with the granulations only on their edges, so that the ribs appear as if mesially depressed. The ribs of the right valve are less strongly beaded and may become quite smooth, especially on the center of the shell disk.
Its ventral margin is gently curved to the rounded and not pointed posterior extremity.
It is a common species at Mt. Hope in the Canal Zone, but it also occurs but less abundant in Western Panama and in Costa Rica.
Gaiun, Siage: Mt, Fope, C. Z. Water Cay, Panama. Lone Ai jied ,Clojpjwtveck, OR. Coll puddone Wale Creek C, fF. IIL DANANGINIUCK Goh,
186 : BULLETIN 39 358
Arca Henekeni Maury
Arca consobrina Sowerby, 1849, Quart. Journ. Geol. Soc. London, vol. 6, p. 52, pl. ro, fig. 12. Not of d’Orbigny, 1844.
Arca consobrina Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 32, Pp. 531.
Arca consobrina Dall, 1888, Trans. Wagner Free Inst. Sci., vol. 3, pt. 4; p. 646.
Arca consobrina Sheldon, 1916, Paleont. Amer., No. I, p. 49.
Arca Henekeni Maury, 1917,Bull. Amer. Pal.,vol. 5, p. 331,pl. 29, fig. 2.
Arca (Scapharca) sobrina Pilsbry and Johnson, 1917, Proc, Acad. Nat.
sci. Phil, vol: 64; p:, 186.
Our shells from Water Cay are similiarin form and general sculpture to the 4. Henekent Maury (A. consobrina Sowerby), from the Miocene of Santo Domingo, the only noticeable differ- ence being the greater number of ribs on the Panama shell. The ribs of HYenckent from Water Cay average about 39 in number, and 35 to 36 on the Dominican shells. The ribs of the left valve are evenly beaded or granulated by concentric threads which pass across the interspaces as elevated threads. The ribs of the posterior-dorsal submargins are nearly smooth. They are more or less divided at the anterior extremity and very finely beaded.
Length 29, height 18.25, diameter 14.15 mm. aM 18, semidiameter of right value
7.25 mim. Gatun Stage: Water Cay.
Arca costaricensis, n. sp. Plate 25, figures 1, 2
Shell moderately large, elongate, convex, with the beaks sit- uated at the anterior 1-3;ventral margin but gently curved to the attenuated and pointed posterior extremity; right valve with about 30 ribs, of which the most anterior and posterior ones are rather wide, the others narrow; interspaces on the center or the disk 3 times or more the width of the ribs; the ribs of the right valve are obscurely granulated on the anterior extremity, more or less smooth on the center but on the posterior extrem- ity, they are double and peculiarly granulated along their sides;
Plate: 24, figs 92) 14
=
359 Costa R1icA MiocENE—OLSSON 187
this granulation begins on the sides and on the more anterior ribs, the center is smooth; this granulation is in the form of in- verted V’s; their acute apices lying along the center of the ribs; the ribs of the left valve are more or less granulated or beaded throughout and with the ribs on the posterior extremity, double and similiar to those of the right valve; hinge-line straight, with very numerous small, uninterrupted teeth; a rather wide cardinal area with 5 or 6 ligamental grooves; ventral margin in- ternally fluted in harmony with the external ribs.
Weneth) 52) height 28, diameter 34) mim: 52, 32 T7.mm (right valve ).
The form of this shell with its gently curved ventral mar- gin and attenuated posterior extremity is like the Arca actino- phora Dall, but is longer and with fewer and heavier ribs. On the other hand, this species may be compared with dartenszs Brown and Pilsbry, from which it differs most strikingly by its larger size, wider, higher and more central umbos and more pointed attenuated posterior extremity.
Gatun Stage: Hill ra, Banana River. Fiull.2, Banana, River. Zone 7, Pumbri Creek.
Arca honensis, n. sp. Plate 22, figures 8, 9
Shell rectangular, elongote, moderately convex; beaks situ- ated at about the anterior 1-4; right valve with about 26, nar- row, smooth or irregularly granulated ribs, separated by inter- spaces of twice their own width; the left valve with the same number of ribs and similiar to those of the right valve; the ribs of both valves are simple and undivided and if granulated, most heavily on the anterior half of the left valve; interspaces smooth or with irregular, raised concentric threads; cardinal area long and rather wide and with 5 or 6 linear grooves; hinge-lines straight with very numerous, small, uninterrupted teeth; ven- tral margin fluted in harmony with the external ribs.
188 BULLETIN 39 360
Length 39, height 23.5, semi-diameter 11 mm (lest valve) 2605 22 12 (left valve) 34 23 Il (right valve)
In general form, quite similiar to the Arca dariensis Brown and Pilsbry, but distinguished mainly by its simple, undivided ribs. Itis also amore convex shell, with higher and more prominent umbos. It differs from Gabb’s figure of 4. ovonlen- sts in its more anterior umbos.
It isa common species, occurring in zones in the Gatun sandstone of Hone Walk Creek of Eastern Costa Rica.
Gatun Stage: Hone Walk Creek.
Arca golfoyaquensis Maury, var. medioamericana, n. var. Plate 23, figures 4-6
cf. drca golfoyaquensis Maury,1917, Amer. Pal., vol. 5, p. 332, pl. 28, fig. 5.
? Arca (Andara) consobrina Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd series, vol. 8, p. 378.
The Arca golfoyaquensis was described by Dr. Maury from the Gurabo and Cercado formation of Santo Domingo. It is closely related to the Avca Henekent Maury, (the Arca consob- vina) differing mainly in form.
The Costa Rican shells are very similiar to Dr. Maury’s type specimens of golfoyaquensis, but differ in having 35 instead of 38 ribs, a higher shell, fuller umbos and sharper ribs on the posterior-dorsal area. ‘The central ribs of the left valve are ele- gantly beaded with their interspaces evenly sculptured with raised concentric threads. On the right valve, the ribs are more smooth but their interspaces are similiarly sculptured to those of the left. The ribs of the posterior-dorsal angle of both valves become divided ventrally by 2 or 3 fine threads. The posterior extremity is straight, meeting the hinge-line at an angle of nearly 120 degrees.
Length 54, height 34, diameter 34 mm.
361 Costa Rica MriocENE—OLSSON 189
Gatun Stage: Hill 1, Banana River. Colle. 5. dned «Clays Creer. Rio Blanco. Old Man Sam Creek, 1 mile south of beach.
Arca Veaichi, n. sp. Plate 23, figures 1-3
Shell large, moderately convex: in outline, the shell is near- ly square with the height about equal to its length; the anterior side is widely rounded to the ventral margin, the posterior side nearly straight, nearly at right angles to the hinge-line; umbos wide, with the unbonal angle widely rounded and with the ex- treme tip of the beaks situated at the anterior 1-3 of the total length of the shell; right valve with about 45 ribs, of which the posterior 14 are on the posterior-dorsal slope; the ribs are rec- tangular in section with the interspaces on the center of the disk about 3-4 of the width of the ribs; the ribs are finely scul- ptured but less strongly on the right valve; on the center of the shell disk, the ribs are finely nodulated or granulated but on the posterior-dorsal slope and on the anterior extremity, the granu- lations of the ribs is gradually overshadowed by the introduc- tion of 4 or 5 fine, threads on the top of each rib the interspaces on the center of the disk of the left valve are finely sculptured with even, raised concentric threads; the left valve is slightly larger and overlaps to a small extent the right; hinge-line straight with numerous small teeth, larger and higher at the ex- tremities; cardinal area that of typical Scapharca, and grooved with 3 lines; interior of the shell deep, with the posterior ad- ductor scar about twice the size of the anterior; basal margin fluted in harmony with the external sculpture.
Length 51, height 51, diameter 41.5 mm.
This isarather large, Cardium-like Arca of nearly square outlines and numerous finely sculptured ribs. Its form is also suggestive of Arvgzna, but its cardinal areais that of a true Scapharca. ‘The type specimen, a finely preserved individual
190 BULLETIN 39 362
with both valves comes from the Gatun of Water Cay. It is quite unlike any other known American species.
It is named for Dr. A. C. Veatch, Director of the Explora- tion Department of the Sinclair Consolidated Oil Corporation.
Gatun Stage: Water Cay. Mt. Flope, (C2:
Arca auriculata Lamarck Plate 22, figure 3
Arca auriculata Lamarck, 1819, n. s. Vert., vol. 6, p. 43.
Arca (Scapharca) auriculata Dall, 1898, Trans. Wagner Free Inst., Sci, vol. 3, pt.4, .p. 649.
Arca auriculata, Sheldon, 1916, Paleont, Amer., vol. I, p. 50, pl. II. fig. 19,
Scapharca auriculata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 339, pl. 28, fo. 3,
A common, recent species of the West Indies. Its hinge- line is generally produced or auriculated at its posterior extrem- ity.
It occurs in the Miocene of Jamaica and Santo Domingo. Dall also records it from near Limon.
The Costa Rican specimens in our collection are all small but otherwise typical.
Gatun Stage: Coll, 3, Hone Walk Creek. Old Man Sam Creek, 1 mile south of the beach.
Section CUNEARCA Dall Arca cacica, n. sp. Plate 24, figure I
Shell of moderate size, strongly convex and with high, in- flated umbos; beaks and umbos about the middle of the shell; the greatest convexity of the shell lying about the anterior 1-3, and with a narrow but deep, radial depressed zone extending from the beaks to the ventral margin, just anterior to the um- bonal angle; the anterior end is slightly rounded, the posterior
363 Costa Rica MrocENE—OLSSON 191
submargins strongly depressed or flattened with a straight post- erior margin; the left valve with 30 smooth and strongly nodul- ous radial ribs of which 9g are situated on the posterior sub- margin; the ribs of the posterior submargin are low, wide and nearly smooth; on the rest of the shell, the ribs are nodulose, very regularly in the middle of the valves more distantly on the anterior ribs; the interspaces between the ribs are narrow on the middle of the valves, become wider anteriorly and on the ex- treme anterior extremity equal the ribs in width; the cardinal area is high, as nearly % of the length. Length 39, height. 33, diameter of the left valve 15 mm.
The above description is based on a single left valve from the Upper Gatun beds near Cahuita. Distantly related to the Arca tncongrua Say and its southern form dbrasz/iana Lamarck, the fossil species differs in its higher and more convex shell, and in its high cardinal area. In this latter feature, this species is like the recent West Coast Arca labiata Sowerby, but has a greater number. of ribs and different form.
Gatun Stage: Across the divide trom Comadre Creek. Groups of Arca Pittiert Dall
This is a group of small and medium-sized Arks, containing 3 species, whose general appearance is that of the recent d7ca Chemnitzi Phil. The Costa Rican shells are usually abundant wherever they occur in the Gatun formation. Their relations and differences may be summarized as follows:
A. Both valves more or less similiarly sculptured, that is the ribs of the posterior half of the right valve, at least obsolete- ly beaded.
B. Ribs 25 to 28; umbos high and full; posterior-dor- sal slope, angular; posterior extremity somewhat produced, giving an elongate outline to the shell.
Area Pitttert Dall
AA. Valves not similiarly sculptured; posterior half of
192 BULLETIN 3g 264
the right valve with narrow ribs, separated by wide inter- spaces.
B. Theintervals between the ribs on the posterior half of the right valve with small (sometimes faint) interstitial threads. Ribs about 29.
Arca Lloydi, n. sp.
BB. The intervals between the ribs on the posterior
half of the right valve smooth; ribs 25, shell usually
larger. Arca) Finds. tn Sia Arca Pittieri Dall ' Plate 24, figures 2-6 Arca (Scapharca) Pittiert Dall 1912,Smith. Misc. Coll., vol. 59, No. 2, p. 9.
This is a common species in the Gatun beds of the Banana River. The ribs of the 2 valvesare similarly sculptured but somewhat less strongly on the right. The umbos are high. The posterior extremity is pointed and produced so that the out- line of the sheil is more elongate than in the following species:
Length 25, height 25, diameter 26 mm. | 29 28 semi-diameter 14.5 mm. Gatun Stage: fiill ra, Banana River, 3. Lone £, Saury Creek. Rio Betey. Coll. 7, Pumbri Creek.
Arca Lloydi, n. sp, Plate 24, figures Io-12
Shell small, cordiform, covex, subrectangular in outline but with the left valve somewhat more pointed at its posterior-ven- tral extremity; anterior side well rounded; posterior side straight and meeting the basal margin at an angle of about 75 degrees; umbonal angle rounded; the left valve has 29 ribs of which 9 are found on the posterior- dorsal slope; the ribs of the left valve are strongly beaded or granulated, and separated by inter- spaces of about their own width; on the right valve, the first 10 anterior ribs are strongly beaded; the next set as far as the um-
365 Costa Rica MiocENE—OLSSON 193
bonal angle are smooth, beyond which on the posterior-dorsal slope the ribs again become beaded as they approach the dorsal margin; the interspaces on the right valve are wide, and on the shell disk anterior to the umbonal angle they carry a fine, but often faint interstitial thread; the intervals are otherwise smooth or finely etched with concentric lines; hinge-lines straight with numerous small, vertical teeth and a wide, smooth cardinal area. Length 18, height 18, semi-diameter 9.5 mm.
A smaller species than the following (//zzdsz) and easily distinguished by the interstitial thread of its right valve. It is much less common than the other two species.
Named for Dr. E. R. Lloyd, formerly of the United States Geological Survey, and who was a member of the first Explora- tion Party of the Sinclair Oil Company in Panama and Costa Rica.
Gatun Stage: Hill 3, Banana River.
Zones, mea Clip Creer: Coll. 7, Pumbri Creek.
Area Hindsi, n. sp. Plate 24, figures 7-9
Shell small or of medium size, cordiform and similiar in out- line to the preceding species but larger; umbos high and wide, and but slightly anterior to the middle; the left valve has about 26 strongly nodulated ribs, separated by interspaces of slightly greater width; the first 9 or 10 ribs of the right valve are nodu- lated, followed by smooth ones to the umbonal angle, beyond which the remaining 7 become nodulated as they approach the dorsal margin; interspaces wider than the ribs, smooth and without any interstitial thread; hinge-line straight with numer- ous, small vertical teeth and a wide, smooth cardinal area; on some shells the ligamental area may carry one or two, small diamond-shaped grooves.
Length 24, height 25, semi-diameter 14 mm. A much larger species than the Zéoydz and without any in-
ig4 BULLETIN 39 366
terstitial thread in the interspaces of its right valve. It is nam- ed for Dr. Henry Hinds, formerly of the United States Geologi- cal Survey and the Exploration Department of the Sinclair Con- solidated Oil Corporation.
Gatun Stage: Hill No. 3, Banana River. Zone E, | Saury\ Greer. Coll. 1, East Grape Point Cree Coll. 3, East Grape Point Creek. Rio Betey.
Subgenus NOETIA, Gray
Arca MacDonaldi Dall Plate 25, figures 4-7
Area ( Noetia) MacDonaldi Dall, 1912, Smith, Misc. Coll., vol. 59, No. 2p. O:
This large Moétia, is very common and characteristic of the Gatun formation in many parts of Northern Costa Rica, and large specimens may reach a length and height of 55 mm or more. It was originally described by Dall from specimens col- lected by Pittier and MacDonald from the Gatun beds of the Banana River, where it is associated with Arca Pittieri and is very abundant.
Two forms or varieties may be recognized. In typical Mac- Donaldt, the shell is high, trigonal, with elevated, narrow umbos and a producted anterior extremity. The beaks are separated from the hinge-line by a wzde ligamental area. 'Thisis the common form along the Banana River. Frequently associated with typi- cal MacDonaldi are shells with a more vectangular outline, lower but wzde umbos. The beaks are close to the hinge line so that the 4agmental area ts narrow. ‘These two types have a very dif- ferent appearance but seem to be connected by transitional forms. To the last variety, I propose the name of subveversa.
Among recent shells, the dvca MacDonaldi finds its closest relation with the common Arca reversa Sowerby of the Pacific coast of Central America.
367 Costa Rica M1ocENE—OLSSON 195
Gatun Stage: Hill 1, ra, Banana River, typical and var. suberversa
Coll. 7, Pumbric Creek (typical) Goll:6, Ked Clif, Greek (typical) Betey (subreversa)
Zone G, Saury (subreversa)
Genus PTERIA, Scopoli
Pteria inornata Gabb Plate 28, figure 9
Avicula inornata Gabb, 1873, Trans. Amer. Phil. Soc., vol., 15, p.
253. Fteria inornata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 4, Pp. 669. ! Pteria tnornata Maury, 1817, Bull. Amer. Pal., vol. 5, p. 345, pl. 26, fig. 14.
This species is locally common in Costa Rica, but generally difficult to collect because of its thin and fragile shell. Its valves are as high as long, strongly convex and smooth. The anterior ear is of medium size, the posterior much larger and produced.
It was described as a Miocene fossil from Santo Domingo, but has also been collected at Bowden.
Uscari formation. Old Harbor. Gatun Stage. Zone 3; Red Cliff Creek.
Genus OSTREA, Linnzeus
Ostrea megodon Hanley Plate 18, figure 1
Ostrea megodon Hanley, 1845, Proc. Zool. Soc. London, p. 106.
cf. Ostrea cerrosensis Gabb, 1869, Geol. Surv. Calif., vol. 2, p. 35, pl. lie £0.61,
Ostrea megodon Dall, 1897, Trans. Wagner Free Inst., vol. 3, pt. 4, p. 685.
Ostrea megodon Maury, 1917, Bull. Amer. Pal., vol. 5, p. 347, pl. 34, fig: 3:
196 BULLETIN 39 268
This is a medium sized and sickle-shaped oyster with 4 or 5, large, tooth-like folds on the anterior and ventral margins. The beak and ligamental area are directed upwards and sometimes strongly to the posterior side. Its external surface, although strongly and deeply folded, is plain and nearly smooth.
The Ostrea megodon isa recent species along the west or Pacific coast of Central America. It occurs also as a Miocene fossil in Santo Domingo and in the Bowden beds of Jamaica.
Gatun Stage: Water Cay. Coll. ga, East Grape Point Creek. Fill ra, Banana River.
Ostrea costaricensis, n. sp. Plate 21, figure 2
Ostrea sculpturata Bose, 1906, Boll. de Inst. Geol. de Mexico, numero 22, p. 76, pl.\9, fg. 4. Not of Conrad, 1840:
Shell of medium size, subcircular in form but somewhat ob- lique; the shell is somewhat flexed or slightly saddle-shaped, the concave side of which corresponds to the left valve; both valves are strongly sculptured, the right with narrow, rib-like coste and wider interspaces, the left valve with the reverse of the sculpture of the right; the ribs are more or less nodose or knob- bed and are crossed by irregular concentric lines, which in some cases may become raised or lamellar-like; the hinge margin is ir- regularly crenulated:
Height 50, length 47, diameter Ig mm.
The type is a specimen from the Gatun of the Rio Blanco with both valve tightly closed so that the interior cannot be seen. What appears to be the same species, is represented by several sessile and free valves from East Grape Point Creek, and which are associated with O. megodon.
It differs from the Osirea gatunensts - haitensis group by its smaller size; less heavy shell and in that the ribs of the valves are narrow and coste-like and not long, strong plice. Bose has recorded and figured this species as Ostrea sculpturata Con-
|
369 Costa Rica MIoCENE—OLSSON 19
rad from the Miocene (as Pliocene) of the State of Oaxaca, Mexico. It is very distinct from the East Coast, Chesapeake Miocene Ostrea sculpturaia both by its form and sculpture.
Gatun Stage: Rio Blanco.
Coll. ga, East Grape Point Creek.
The collection also contains fragments or much decayed specimens of two or more large oysters, one of which is probably the gatunensis Brown and Pilsbry of the Canal Zone, and a large, heavy sedleformis-like species, possibly the Gzlbertharrisz Maury of the Dominican Miocene. ‘The specimens are however too fragmentary for positive determination.
Genus PECTEN, Muller Pecten gatunensis Toula Plate 16, figures 3, 4
FPecten (Flabellipectcn) gatunensis Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt Wien, vol. 58, p. 711, pl. 29, fig. 2.
Pecten gatunensis Brown and Pilsbry, 1911, Proe. Acad, Nat. Sci. Phila,, vol. 63, p. 365.
Shell rather large, subcircular; the left valve is nearly flat or slightly vaulted transversely across the middle of the shell; the right valve is moderately convex; the ears are small and nearly equal,those of the right valve slightly flaring at the sides; the right valve has 23 or 24 fairly strong ribs separated by inter- spaces a little more than % of their width; in large right valves, some of the central ribs become dichotomously or trichotomously divided but this feature is not always developed to the same ex- tent on all shells; the left valve has 18 or 19 ribs which fade out on the wide dorsal submargins; the ribs of the left valve are narrow with the interspaces nearly twice as wide; the surface is covered with fine, even, raised concentric threads best preserved on the left valves; interior fluted.
Length 62, height 60? diameter 12.5 mm. 62 60? 4 mm (left valve)
198 BULLETIN 39 370
We have redescribed this species as Toula had but a speci- men of the right valve. Itisa true Pecten, distantly related to the Pecten hemicyclicus Ravenel from the Chesapeake Miocene of Florida and the Carolinas, but is smaller and less broad. In both species, the ribs on the right valves of large specimens become divided in the same manner.
Garin Stages Gari. On 2, Water Cay,
Pecten MacDonaldi, n. sp. Plate 16, figures I, 2
Shell large, subcircular; ears of medium size and equal; left valve is slightly convex due tothe middle of the shell being transversely humped or vaulted and a depressed zone follows on the inner slope of each of the raised dorsal submargins; the right valve is slightly but evenly convex; sculpture of the right valve consists of about 26, low ribs which widen out as they approach the ventral margins; their interspaces are at first nearly as wide as the ribs themselves but become only % or % as wide ventrally; the left valve has 21 or 22 narrow ribs and wider inter- spaces; on the dorsal submargins, the ribs are small and fade away; surface with fine, even, raised lines best seen on the left valve.
Length 106, height 97, semi-diameter 12 mm. 83 80 mm.
The fragmentary Toro limestone at Gatun and Toro Point contains very few good fossils,although the rock itself is compos- ed almost entirely of broken and ground fragments of shells and barnacles. Dall has described an Zpztontum toroensis from the Toro Point limestone where it is fairly common. ‘The Pecten MacDonaldi is found in the Toro limestone which caps the hills just west of the locks at Gatun.
This species has much the same contour as the recent Pecten maximus Linné, from Europe. It isa larger species than the Pecten gatunensts 'Toula.
Toro Limestone. Gatun, west of the locks.
a7 Costa Rica MrocENE—OLSSON 199
Pecten coralliphila, n. sp, Plate 17, figures 5, 6
Shell small, very thin and generally translucent; broader than high; right valve moderately convex, the left concave with raised submargins; ears small and subequal; sculpture of the right valve of about 23 low, flat ribs, with flat interspaces a little more than % the width of the ribs; some of the ribs may become divided medially; the ribs of the left valve, about 20 in number, are narrow, usually somewhat irregularly spaced so that their interspaces are of variable width and new ribs may occasionally appear inthe wider interspaces near the ventral margin; surface in addition finely sculptured with even, raised concentric threads; interior of the shell fluted well into the cav- ity of the shell, in harmony with the external ribs.
Length 34, height 29, semi-diameter 4 mm (right valve)
This Pecten is fairly common in the coralline phases of the Miocene and its associated marls and sands. It is extremely thin and fragile in texture. Its sculpture resembles that of /ecten gatunensis Toula but is much more irregular. The interspaces of the left valve are of very variable width.
Cain) Stage: Port Limon.
Subgenus CHLAMYS, Bolton Section PLAGIOCTENiUM Dall
Pecten lfevicostatus Toula Plate 16, figures 5-7
Fecten tevicostatus Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, Malem: )ivole 53) p.. 7.03.0 ple 26; fies. 4) 154)6,
FPecten (Chlamys) santarosanus Bose, 1906, Boll. de Inst. Geol. de MEXICO, MUMero -22,p.\73) ply G6) figs. 2, 4,-° 2° Pecten (Chlamys } santarosanus Bose, op. cit. p. 23, pl. 1, figs. 1 and 4, 2 and 5.
A scallop of the general type of Pecten ventricosus Sowerby; both valves equally and strongly convex; ribs about 21, which are square or rectangular in section on the umbos and with deep interspaces; the ribs generally flatten or become more rounded in
200 BULLETIN 39 372
large shell near the ventral margins; the interspaces are covered with fine, raised, concentric lines, best seen on the umbos or on young shells; they become more crowded towards the ventral margins and may pass over the tops of the ribs as well; the dor- sal and posterior submargins are strongly flattened and sculp- tureless; the anterior ear is a little larger, projects forward and carries, in the right valve, a byssal notch; it is sculptured with several radial riblets; the posterior ear has its upper half nearly smooth or but faintly sculptured, but its lower half carries a band of 5 or 6 strong, radial cords. Height 29, breadth 34, thickness 10.50 mm (right valve) 34 38 22 (both vaives) 39 43 14 (left valve)
This is the common Fecfen in Costa Rica and frequently forms zones in the lower part of the Gatun formation. All our specimens have 21 ribs, which vary from strongly rectangular in section with deep interspaces to more flattened and rounded on the larger individuals.
Bose has described as /ecten santarosanus, a small scallop shell from the Miocene of the State of Oaxaca, Mexico. The ribs number 23 to 25. Later in the same volume, he figured a larger Pecten from the Miocene of La Barranca de Santa Marie Tatella, as Pecten santarosanus. This shell has but 21 ribs and is equivalent to Toula’s Jevicostata. If this larger specimen of Bose’s is equivalent to his typical sanx/arosanus, this name must of course take priority over Toula’s levzcostatus. As Bose’s typi- cal santarosanus has 23 to 25 ribs, while in /evzcostatus the ribs appear to beconstantly but 21 in number, they probably repre- sent two different species.
In the Dominican Miocene J/evicostatus is represented by the Pecten excentricus Gabb, which appears to differ constantly by its lower and more rounded ribs and smaller ears. ‘The recent Pacific Pecten ventricosus Sowerby is closely related and probably the direct descendent of /evicostatus. In that species, the ribs are low and rounded as in excentricus and as occasionally seen in
ee =
373 Costa Rica MIOCENE—OLSSON 201
large examples of Jevicostatus. The sculpture of the posterior ear is uniform. Inthe Chasapeake Miocene fauna the /ecten eboreus Conrad, variety comparilis Tuomey and Holmes is dis- tantly related, but is larger, less convex and generally with more numerous ribs.
Gatun Stage: Gatun, C. Z. Quitana Creek. Water Cay. flone Creck. Banana River. Listrella River.
Pecten costaricensis, n. sp. Plate 18, figures 8-11
| Shell small, inequivalve, oblique; the right valve is moder-
ately convex, the left but slightly; right valve with 17 flat-top- ped ribs, square in section and with deep U-shaped interspaces of a width equal to the ribs; these interspaces are sculptured with fine, concentric, raised lines which are quite far apart on the umbos, but become crowded ventrally or disappear entirely; the left valve has 17 or 18 ribs which are lower and more round- ed in section; the fine concentric lines on the left valve are con- fined to the umbos, the interspaces over the greater part of the shell being smooth; the anterior and posterior dorsal slopes are flattened and smooth, but sculptured below in each valve by strong radial threads on the lower side of the ears; ears subequal; the right valve with a deep, byssal notch in its anterior ear; the left valve has the lower half of each ear with 3 strong radial threads, while on the upper half, the radial threads are largely obsolete; valves internally grooved in harmony with the exter- nal ribbing to about the center of the shell.
Height 26.50, breadth 28, thickness 7.50 mm (right valve) 24 24 4.00 mm (left valve) 27 28 5.50 min (left valve)
Differs from the Fecten levicostatus ‘Toula, by its unequal valves, less convexity and fewer ribs. It is somewhat like the Pecten tnequalis Sowerby, from which it is distinguished by its
202 BULLETIN 39 374
more flattened left valve and lower, smaller ribs of its right. In emegualis, the fine, lamellose, concentric lines cover the entire interspaces between the ribs on both valves, while in the present species the interspaces of the left valve are largely smooth with the concentric lines confined to the extreme umbonal area.
Gatun Stage: Coll. 4, Red Cliff Creek. ZI mile south of the beach, along Old Man Sam Creek: Coll. 4, Last Grape Point Creek. Rio Blanco.
Section AEQUIPECTEN, Fischer Pecten (Aequipecten) preglyptus, n. sp. Plate 17, figures, 2, 7
Shell of moderate size, subcircular in form and convex; ears of usual size, subequal and with a straight hinge-line; sculpture of about 16 or 17 wide, little elevated ribs and narrower, scarce- ly distinguishable interspaces; the surface is otherwise smooth or with fine, concentric lines; interior of the shell with 29 or 30 Amustum-like internal liree which are evenly spaced and not in pairs; these liree are very faint in the middle of the shell cavity but strong distally.
Length 50, height 49, semi-diameter 6.5 mm.
An Amusium-like species found in the lower sandstones and conglomerates of the Gatun formation in upper Cocles Creek. It is closely related to the recent deep-water Pecten glyptus Ver- rill from the cost of Hatteras and Marthas Vineyard (see Dall Proc. U. S: Nat. Mus., vol. 12, p. 248, pl. 8; fiesta.) born species having about the same number of low, wide ribs and in- ternal liree. The fossil shell seem to have been nearly circular while glyptus is decidely oblique in outline.
Gatun Stage: Upper Cocles Creek.
375 Costa RicA MIocENE—OLSSON 203
Subgenus PSEUDAMUSIUM, H. and A. Adams Section PSEUDAMUSIUM, s. s.
Pecten almirantensis, n. sp. Plate 18, figure 16
Shell small, thin, compressed or but slightly convex; the posterior and anterior dorsal slopes are straight, meeting the beaks at an acute angle; the right valve is finely sculptured with close, regular, concentric lamella and wider interspaces; the posterior and anterior sides show a few, irregular, radial threads which cross the concentric lamellz; the ears are unfortunately broken; the interior of the shell cavity is filled with a thin de- posit of callus; ventral margin plain; hinge line minutely groov- ed with vertical lines.
Length 4.75, height 4.75, diameter of right valve 60 mm.
This small, elegant species is represented solely by an im- perfect right valve and represent the first true Pseudamusium to be discovered in the Miocene beds of Panama. Its sculpure is like the recent deep water Pseudamustum strigillatum Dall from the West Indies, but differs in having its anterior and posterior dorsal submargins meeting at the beaks at a much more acute angle.
Gatun Stage: Bocas del Toro
Section CYCLOPECTEN, Verrill Pecten oligolepis Brown and Pilsbry
Pecten aff. subhyalinus Smith, Toula, 1911, Jahrb. der K-K Geol. Reichsanstalt, Wien, vol. 61, p. 492, pl. 31, figs. 1a, b, c.
Pecten (Cyclopecten) oligolepis Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., vol. 64, p. 512, text-figure 5.
A very small species with broad, nearly smooth valves and large ears. The right anterior ear has a deep byssal notch and its surface is sculptured with fine, raised, concentric, lamellose lines. The ears of the left valve are similtarly sculptured but fin-
204 BULLETIN 39 376
er. The surface of the valves appear nearly smooth but in well- preserved specimens very faint concentric lines and fine radial striz may be seen. The left valve is more coarsely sculptured. The hinge is finely grooved with vertical lines. The measure- ments given by Pilsbry for his Gatun specimen, length 2.8, height 2.7, diameter of right valve .8 mm, are the usual dimen- sions for the Costa Rican specimens.
Gatun) Stage: Gar, G7 Coll. 4, East Grape Point Creek. Middle Creek.
Pecten aotus, n. sp. Plate 18, figures 17, 18
Shell small, thin, depressed or but slightly convex and Amusium-like; both valves except on magnification appearing smooth; ears of the left valve large and subequal; the anterior ear of the right valve about twice that of the posterior and car- rying a deep, byssal notch; the surface of the left valve on mag- nification is seen to be sculptured with fine, radial lines which often occur in irregular streaks or blotches, imparting a peculiar appearance to the shell; the right valve is perfectly smooth and featureless, except for a band of rough scales on its anterior ear above the byssal notch; interior of shell cavity shallow, a simple ventral margin and the hinge-line vertically grooved with fine lines.
Length 4.5, height 4.5, diameter .75 mm. left valve.
Differs from FPecten oligolepis Brown and Pilsbry in its larg- er Size, thinner and more Amusium-like shell, and strongly sculp- tured left valve. This sculpture consists of fine lines or camp- tonectes striations, which may be uniformly distributed but more usually occur in irregular streaks or blotches, imparting a dis- tinctive appearance to the surface. This sculpture is seen only on a slight magnification, the shell otherwise appearing smooth and featureless. It is quite common in the coralline limestones of Port Limon.
377 Costa RicA MIocENE—OLSSON 205
Gatun Stage: Port Limon. Collis, eed Clif (Creer
Genus AMUSIUM, Bolton
Amusium luna Brown and Pilsbry Plate 17, figure 1
Amusium Mortont Bose, 1906, Boll. de Inst. Geol. de Mexico, Numero 22, p. 24, pl. 1, figs. 3,6, 7,9. Not Amusium Mortont Ravenal,
1844, Amusium Mortoni Bose, op. cit. p. 74, pl. 8, figs. 1, 2; pl. 3, fig. 3. Pecten (Amusium) cf. Mortoni Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, Wien, vol. 58, p. 714, pl. 26, figs. 8, 9.
Pecten (Amusium) luna Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., vol. 64, p. 514, pl. 23. fig. 1.
This is the common Gatun Amustum in Panama and Costa Rica. Its valves are nearly equal and but slightly convex; ears equal and small, and defined from the rest of the shell by a sharp ledge; the umbos, as well as the rest of the shell disk are smooth, except for the faint lines of growth; the internal ribs are in pairs, numbering about 23 and spaced at intervals nearly twice the width of the pairs of ribs themselves.
Bose and Toula both referred this species to the Upper Chesapeake Miocene and recent Amustum Mortont Ravenel, from which it is very distinct. Amustum Morton? is not only much larger (height 170 mm), but is porportionately much broader, and with larger ears which are defined simply by a line from the rest of the shell.
The Dominican A. papyraceum Gabb has larger ears, the in- ternal ribs are more crowded and the left valve is generally strongly flexed along the anterior and posterior submargins. Amusium Toule Brown and Pilsbry, is found in the Gatun of the Canal Zone. Its surface is marked with dark-colored rays and is said to contain no internal ribs.
206 BULLETIN 39 378
Gatun Stage: Gatun, C. Z. Flotel Creek. Zone 6, Old Man Sam Creek. Coll. 4, East Grape Point Creek:
Amusium bocasensis, n. sp. Plate 17, figures 3, 4
Shell rather small, thin and subcircular in outline; moderate- ly convex; ears proportionately larger than in A. /una, and with their dorsal margins slightly inclined upwards from the beaks to their outer edges; the surface is smooth and polished, finely covered with concentric growth-lines and faintly showing the position of the internal liree; interior of shell with about 34 lirsze, which are usually nearly evenly spaced and not in pairs; the internal lire commence well in the interior of the shell, but are strongest distally.
Height 4o, breadth 43. 35 38, thickness 6 mm. (right valve)
A much smaller and more delicate species than the Amuszum luna Brown and Pilsbry. The internal lire are more or less equally spaced and not in pairs. It differs from Amuszum Lyonit Gabb from the lower Miocene of Sapota, Costa Rica by its much smaller ears.
Gatun Stage: Bocas del Toro.
Genus SPONDYLUS, Linnzeus Spondylus chiriquiensis, n. sp. Plate 20, figures I, 2, 5, 6
Shell large, ponderous, equivalve and strongly convex; both valves of the same convexity, but with the beak of the right, high and projecting above the strongly incurved beak of the left; in large shells both valves are provided with a cardinal area, that of the right being both high and wide and for the greater part straight but the beak curving over it toa small extent at its upper end; the cardinal area of the left vale is much smaller,
379 Costa RicA MIocENE—OLSSON 207
narrow and appears only with maturity and is more or less hid- den by the strongly incurved beak; the sculpture consists of strong radial cords and small ribs divisible into 3 sets; a primary set of 7, which in perfect specimens and in the young carry large, strong spines; a secondary thread in each of the wide interspaces and when perfect bearing smaller spines, and tertiary threads of a variable number between the secondaries and primaries; the whole surface is in addition, finely, longitudinally starited with minute, scale-bearing threads; young shells have a small foliace- -ous area on the umbo of the sessile or right valve and strong spines on both valves.
Height 151, length 120, diameter 115 mm.
A large, ponderous species with nearly equal and strongly convex valves which show no distortion due to attachment Young shells associated with the type have a small, somewhat foliaceous attachment area on the right valve, and each valve bears large, strong spines on its primary set of radial cords and finer spines on its secondary and even on some of the tertiary threads.
Of described species, the chiriguzensz?s seems nearest related to the dostrychites Guppy from the Miocene of Santo Domingo and Jamaica but differs by its much larger size, proportionately more convex whorls and by its cardinal area which is more up- right and with a more incurved beak at itsupper end. The sculpture of chiv7guiensts is more spiny,the spines being borne by the primary radials and to a less degree by the secondaries, and even by the tertiaries. In dostrychites only the primaries bear spines, the others appearing as large, nearly smooth, intermedi- ate threads and cords. We have specimens of a large, closely allied Spondy/us from the Chesapeake Miocene of Florida, dif- fering mainly in its cardinal area and in details of sculpture.
Gatun Stage: Water Cay.
Spondylus gumanomocon Brown and Pilsbry Plate 21 figure I
Spondylus americanus Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 257. Not of Lamarck. ;
208 BULLETIN 39 380
Spondylns gumanomocon Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., vol. 14, p. 514, footnote.
Spondylus gumanomocon Maury, 1917, Bull. Amer. Pal., vol. 5, p. 355-
This is a species with very unequal valves, that of the left or upper, being small, pectiniform and lacking ina cardinal area, while the right or lower valve is strongly convex, with a high produced beak and a high cardinal area. ‘The sculpture of the two valves is similiar, except that the umbo of the right is strongly foliaceous. It occurs in the Miocene of Santo Domin- go, where it was referred to the recent S. americanus by Gabb.
Gatun Stage: Coll. 2, Hone Walk Creek. Port Limon.
Genus PLICATULA, Lamarck
Plicatula marginata Say Plate 28, figures 6, 7
Plicatula marginata Say, 1824, Journ. Acad. Nat. Sci. Phila., vol. 4, Ppp. 136-137, pl. 9, fig. 4.
Plicatula marginata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 4, p. 764:
The Panama and Costa Rican examples are not typical but they approach more closely the Plicatula marginata Say than they do the densata which Dall has recorded from the Bowden beds of Jamica.
The shells vary from subcircular to elongate and in some cases carry the dark, marginal band so frequently seen in typical marginata. ‘The ribs number five to seven; the shells with more rounded form and more numerous ribs suggest the densaza, but the ribs are higher and more foliaceous, and can be exactly du- plicated by scores of true marginata in the Cornell collection. The above determination must however be considered as purely provisional at the present time.
The typical P. densata was described by Conrad from the lower Chesapeake Miocene of New Jersey, but it also occurs in
381 CosTaA Rica MiockKNE—OLSSON 209
the synchronous Calvert formation of Maryland. It is also found as a lower Miocene fossil in Florida and Dall recorded it from the Bowden beds of Jamaica and from the Oligocene of Gual- lava, Costa Rica. The P. marginata Conrad is an Upper Chea- peake Miocene and Pliocene fossil of the eastern United States. Gatun Stage: Water Cay, Old Man SamCreek, 1 mile south of the beach.
Genus LIMAEA, Born Limaea solida Dall Plate 28, figure Io Lime@a solida Dall, 1898, Trans, Wagner Free Inst. Sci., vol. 3, pt. 4, p. 769, pl. 35, figs. 4, 5. A small, solid shell, externally sculptured like a small Gly- cymeris. It has 12 rounded ribs and a coarsely crenulated basal
margin. Our single specimen from Costa Rica isa little smaller than Dall’s specimens from Bowden.
Gatun Stage: Coll. 4, Red Cliff Creek.
Genus ANOMIA, (Linné) Muller Anomia simplex d’Orbigny Plate 21, figure 6
Anomia ephippium Conrad, 1845, Fossils of the Medial Tertiary, p. 75, pl. 43, fig. 4. Not of Linné.
Anomtia simplex d’Orbigny, 1845, Hist. Pol. y Nat. Isla de Cuba, p.
367.
Anomia ephippium Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 257.
Anomia simplex Dall, 1898, Trans. Wagner Free Inst., Sci., vol. 3, pt. 4, p. 784.
Anomia simplex Bose, 1906, Boll. de Inst. Geol. de Mexico, numero 22). 25, pl. 2, figs. 18-23. Anomia simplex Maury, 1917, Bull. Amer. Pal., vol. 5, p. 355, pl. 26, fig. 15. The Anomia simplex is both common and characteristic in the Gatun of Costa Rica as well as in certain parts of the Prov-
210 BULLETIN 39 382
ince of Colton, Panama, near the Canal Zone. It frequently gives rise to important zones in the lower part of the Gatun for- mation, where it occurs in large numbers associated with Pecten levicostatus ‘Toula.
The specimens are generally large, thin-shelled, smooth and irregular inform, due to the object on which they were attach- ed. They average for the upper valves about 35 mm in height.
Anomia simplex appears as a Miocene fossil in the eastern United States for the first time in the St. Mary’s formation of Maryland in the upper part of the Lower Chesapeake. It occurs . throughout the Upper Chesapeake Miocene and Pliocene and passes into the recent fauna. Locally as a fossil it may become very abundant, as at the base of the Pliocene overlying Upper Chesapeake Miocene along the shores of Lake Waccamaw, North Carolina. It occurs rarely as a Miocene fossil in Santo Domingo, where it has been recorded by Gabb and Maury. In Mexico, it occurs in the Miocene of Santa Rosa, Vera Cruz as figured by Bose.
Gatun Stage: Several localities, Prov. of Colon, Panama.
Middle Creek. Comadre Creek. Hone and Hotel Creeks. Hill No. 2, Banana River, Soury Creek, etc.
Genus PLACUNANOMIA, Broderip Placunanomia lithobleta Dall Plate 21, figures 3, 4, 5
Placunanomtia lithobleta Dall, 1898,Trans. Wagner Free Inst. Sci.,vol. BPE A Peyae:
Placunanomia lithobleta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 356, pl. 34, eS.) 2.
The Costa Rican specimens are frequently widely plicate, nearly as much asin the Duplin Miocene P. plicata Tuomey and Holmes, but the surface is marked with fine, wavy, radial threads so that even very small fragments of this species are very easily recognized. It occurs in the Miocene of Jamaica, its type local- ity, and in the Miocene of Santo Domingo.
Gatun Stage: Hili 3, Banana River.
Sa
=e
383 Costa Rica MriocENE—OLSSON 211
ORDER ANOMALODESMACEHA
Genus THRACIA, Blainville Subgenus CYATHODONTA, Conrad Thracia Tristani, n. sp. Plate 20, fig. 3
Shell thin, subquadrate, the left valve slightly convex; the beaks situated at the posterior 1-3 of the shell, are low and in- conspicuous; anterior end, wide and broadly rounded, the poster- ior contracted, depressed and truncated at its extremity; the sur- face is sculptured with regular wave-like folds, which on the middle of the shell are concentric, but are oblique on the anter- ior portion; they are lacking from the posterior area; in addi- tion the whole surface is very finely pustulose and crossed by very indistinct growth-lines.
Length 42, height 30, diameter of the left valve 7.00 mm-
The Cyathodonta gatunensis 'Toula of the Canal Zone is a similiar but larger species, reaching a length of 52 mm. It dif- fers from the Costa Rican species in its greater proportional length, the more central beaks and in its sculpture, which ac- cording to Toula’s figure is strictly concentric.
The C. undulata Conrad, a recent species from the Gulf of California, is sculptured like 77istanz with oblique riblets but the left valve is depressed and strongly flattened.
This species is named for Professor J. Fidel Tristan, Director of the National Museum of Costa Rica.
Gatun Stage: Rio Betey.
212 BULLETIN 39 384
ORDER TELEODESMACEA
Genus CRASSATELLITES, Kruger
Crassatellites Reevei Gabb Plate 20, figure 4
- Crassatella antillarum Reeve? C. Reevei Gabb 1873, Trans. Amer. Phil.Soc.\ vol, 15.7p27252.
Crassatellites veevei Brown and Pilsbry, 1913, Proc. Acad. Nat. Sci. Phila.,\ vol. 64.\pi505 ipl. 2a) fis. 5)
The Crassatellites Reevet Gabb is not an uncommon fossil in the Gatun beds of western Panama and Costa Rica. Our specimens agree quite closely with Brown and Pilsbry’s figure of
the single right valve collected by Gabb in the Miocene of Santo
Domingo. Young and full-grown specimens show a strong posterior angulation which extends from the beak to the poster- ior ventral extremity. The posterior end is obliquely truncated but less so in young shells. The surface sculpture consists of even, concentric costeze which on the slightly flattened umbos are a little wider apart asin the Cvassatellites of the section Scam- ula Conrad. Inthis later character, our shells disagree with Brown and Pilsbry’s redescription of Gabb’s type specimen, in which the sculpture is said to extend without any irregularity upon the rather flattened umbos.
The C. mediamericanus Brown and Pilsbry from Sapote, Costa Rica is an earlier shell and is higher, less obliquely trun- cated behind, more coarsely sculptured and lacks the posterior angulation. In these characters except the lack of the posterior- umbonal angulation it is like the young shells of Aeevez.
Length 55, height 36.5, semi-diameter 10.5 mm. 50 35 diameter 21 mm, 8 42 semi-diameter 14 mm.
Or &
385 Costa RicA M1iocENE—OLSSON 212
Gatun Stage: Water Cay. Old Man Sam Creek, Coll. No. 6.
Banana River.
Subgenus CRASSINELLA, Guppy
Crassatellites bowdenensis Dall, variety costaricensis, n. var. Plate 29, figure 12
Cf. Crassatellites ( Crassinella) bowdenensis Dall, 1903, Trans. Wagner Preevlast, oct., Vol. 3, pt. 6, p.. 1476; pli 50, figs. 2; 3.
Shell small, triangular, slightly convex, subequilateral, the anterior end generally a little longer and more pointed; beaks triangular, slightly pointing forwards; lunule large, narrowly lanceolate, smooth; escutcheon small; surface sculptured with small, even, concentric riblets which on some shells may be strong and persistent over the greater part of the shell, or on some specimen sub-obsolete; interior of shell cavity shallow, with a moderately heavy hinge.
Length 3.00, height 2.75, diameter .75 mm. 3.50, 3550) By eyoudaouk:
The typical dowdenensis Dall from the Bowden Miocene of Jamaica is a more solid, more convex and triangular shell. The variety costaricensis is very abundant at several localities in Costa Rica in certain horizons. The sculpture varies from strong, even, concentric riblets covering the greater part of the shell, to forms which are nearly smooth.
Gatun Stage: Hill ra, Banana River.
Middle Creek. Zone 3, Outtana Creek.
Crassatellites midiensis, n. sp. Plate 29, figure 11
Shell small, trigonal, slightly convex and equilateral; the beaks are central and erect; the dorsal margins straight and meeting at the beaks nearly at right angles; the surface is sculp- tured with subregular, distant, concentric lamelle-like riblets,
214 | BULLETIN 39 386
numbering about 10; the interspaces carry irregular, finer, con- centric lines; lunule narrowly lanceolate, with the escutcheon narrow and smaller; interior of shell cavity shallow, with a mod- erately heavy hinge.
Length 3.25, height 3.25, diameter of one valve 1.00 mm.
A small, rare species, somewhat like the C. Guppyi Dall of the Miocene of Jamaica, but more trigonal, with more central beaks and somewhat different sculpture. The sculpture and form are very similar to specimens of C. galvestonensts Harris in the Cornell Collection from the Chesapeake Miocene of York- town, Va. This latter species is generally much worn but per- fect specimens have a coarse concentric sculpture like that of lunulatus Conrad. ‘The Costa Rican shells are however very much smaller.
Gatun Stage: Middle Creek, C. R. Coll. 4, East Grape Point Creek. Water Cay, Panama.
Genus CARDITA, (Bruguiére) Lamarck Cardita caribbeanensis, n. sp. Plate 26, figure 22
Shell small, moderately convex, elongate, its length a little less than twice its height; beaks very near the anterior end are small but distinct; an umbonal ridge extends from the beaks to the posterior-ventral extremity and along which and bordering the ribs are largest; the right valve with about 20, hight, narrow and sharply nodulose ribs, separated by wide, triangular, smooth interspaces; 6 of the ribs are situated on the posterior-dorsal submargins, and of which the 4th and 5th from the dorsal mar- gin are much smaller than the others; of the remaining ribs, the largest are found on the umbonal ridge, and become progressive- ly smaller anteriorly; lunule small but distinct; hinge of the right valve that of typical Cavdzta, with 3 cardinal teeth of which the middle one is much the largest, no laterals; ventral margin, internally fluted in harmony with the external ribs.
387 Costa RicA MIoCENE—OLSSON 215
Length 18, height 12.50, diameter of the right valve 5-50 mm.
This small species represented by a single right valve in our collection appears to be a true Cardzta, its hinge being essential- ly that of the recent West Coast Cardita laticostata Sowerby. With the exception of a few doubtful species in the Eocene of United States, true Cardifa has heretofore not been found as a fossil in the Americas and its recent American species are con- fined to the Pacific coast.
Compared with the recent Cardtta laticostata Sowerby, which is abundant along the Pacific Coast of Panama, the fossil shell differs in its smaller size, smaller umbos and inform. Its lunule is like that of /atzcostata.
Gatun Stage: Banana River.
Section CARDITAMERA, Conrad Cardita matima, n. sp. 7 Plate 32, figure 8
Shell robust, solid, strongly inequilateral with the beaks Situated at the anterior extremity of the shell; umbos large and full with strongly curved beaks above the small, sunken lunule; outline of the shell from within is subrectangu- lar with straight ventral and posterior sides which meet at nearly right angles; sculpture of about 18 strong, heavy cord- like ribs, separated by interspaces 1-3 of their width; the ribs are strongly nodulated on the disk and umbonal portion of the shell, but below they are simply irregularly wrinkled; a deep furrow extends from the beaks to the posterior extremity, just above the umbonal angle; the rib forming the summit of the ridge between the umbonal furrow and the hinge-line is larger than those on each side; shell cavity deep with a crudely den- ticulated ventral and posterior margin, hinge large (broken in our specimen), with a well-developed anterior lateral tooth in its left valve.
Length 30, height 25, semi-diameter 12 mm.
216 BULLETIN 39 388
But a single specimen of this very distinct spceies was col- lected by Dr. A. C. Veatch from the Upper Gatun of Old Man Sam Creek in eastern Costa Rica. It differs from the other known American Carditameras by its convex shell and strongly curved beaks. The ribs are large and strongly nodulated at first, becoming irregularly wrinkled near the ventral margin. The hinge is unfortunately broken on our specimen, a left valve, but the small anterior lateral still remains which shows this shell to belong to the section Carditamera
Gatun Stage: Old Man Sam Creck, 1 mile south from shore.
Genus VENERICARDIA, Lamarck
Venericardia scabricostata Guppy Plate 32, figures 14, 17
Cardita scabricostata Guppy, 1866, Quart. Journ. Geol. Soc. London, VOI’ 225.2203), pl 18, tig. a0;
Cardita scabricostata Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 252.
Venericardia scabricostata Maury, 1917, Bull. Amer. Pal., vol. 5, p.
362)-pl. 33. digi:
This species originally described by Guppy from the Bowden beds of Jamaica is common in the Miocene of Costa Rica and frequently forms small zonesin the Gatun beds. It occurs also in the Miocene of Santo Domingo.
The ribs numbering about 18 are high, narrow and strongly granulated. The interspaces are wide and generally smooth.
Gatun Stage: Zone E, G, Saury creek.
Zone 1, 3, Hast Grape Point opcep: Coll, a) Maa Clg) creer, fleadwaters of Middle creek.
Coll. C, Comadre creek.
Venericardia Terryi, n. sp. , Plate 32, figures 12, 13
Shell of medium size, solid, moderately convex and sub-cir-
339 Costa RicA MIOCENE—OLSSON 217
cular in form; beaks situated well forward and but slightly back of the extreme anterior end of the shell; the beak itself is small and loosely coiled over the small sunken lunule; sculpture consists of about 13, low, wide, smooth ribs which fit so closely together that there are no interspaces between them; the ribs are rounded in section on the umbos, wide and low ventally; the ribs are largest about the middle, small and fading away on the poster- ior and anterior dorsal submargins; interior concealed in a hard sandstone matrix. Length 22, height 21, semi-diameter 6 mm.
The first specimens of this interesting species was collected by Mr. R. Terry from some Gatun sandstones in Quitana Creek, a small tributary streams of the Rio Betey. In that locality, it is an abundant species and occurs in small zones.
It is related to the recent Pacific, V. crassicostata Sowerby but is a much smaller species. Gatun Stage: Quiiana Creek. ffill No. 3, Banana River.
Subgenus PLEUROMERIS, Conrad Venericardia Conradiana Gabb, var. limonensis, n. var. Plate 26, figs. 19, 20
Cardita Conradiana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., vol. 8, ZAG Series, p..377, pl. 47, fg. 79.
Shell small, subcircular, moderately convex; umbos high and prominent, with small, inconspicuous beaks at the anterior 1-3 of the shell; the sculpture consists of about I8 heavy, gran- ulated ribs, separated by somewhat narrower interspaces; interi- or of the shell deep, the hinge with 2 cardinal teeth and a small posterior and anterior lateral.
eneth!.7.00, height')7,.00;"diameter of the left valve 3.60 mi.
Gabb specimen described from the Pliocene beds of the Limon Peninsula, measures 1% mm in length. His enlarged figure is very similiar to the present shell, in its subcircular forin
218 BULLETIN 39 399
and in number of ribs. The figure of the hinge shows the two cardinal teeth but no laterals.
The common ¢videntafa Say of the Upper Chesapeake Mio- cene and recent fauna of the West Indies differs mainly in its more triangular form.
Gatun Stage: Port Limon.
Genus CHAMA, (Linné) Bruguiére
Chama congregata Conrad Plate 28, figure 11
Chama congregata Conrad, 1833, American Journal of Science, vol. 23, p. 341.
Chama congregata Conrad, 1838, Fossils of the Medial Tertiary, p. 32, Dil aoe:
Chama congregata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt..6; p. T400,
Chama congregatoides Maury, 1917, Bull. Amer. Pal., vol. 5, p. 200, pl. 23, fg. 8:
Our specimens from Costa Rica are all small and belong to the upper or free valve, which in this species is the right. These specimens cannot be distinguished from true congregaia of the saie size from the eastern United States, where it is an abund- ant fossil throughout most of the Chesapeake Miocene. The upper or right valve is finely, radiately frilled or fluted and these frills are cut by the concentric lamelle. ‘The attached or left valve is more coarsely sculptured,both radially and concentrical-
ly. Gatun Stage: Water Cay. Coll. a. Red Clif ‘Creek: Genus ECHINOCHAMA, Fischer Echinochama antiquata Dall Plate 28, figure 8
Chama antiquata Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22, p. 294. Not of Linné.
Echinochama antiquata Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1404, pl. 54, fig. 9.
39t Costa RicA MIocENE—OLSSON 219
Not “chinochana antiquata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 365, pl. 33, fig. 10. &. yaquensis Maury.
This species described by Dall from the Bowden beds of Jamaica differs from the common, recent arcinella Linné of the West Indies, by its more numerous, irregular and lower ribs which bear only small, short spines. In arcinella the middle ribs are heavy and bear large, stump-like spines and the wide interspaces are reticulated with a coarse, mesh-like sculpture.
Gatun Stage: Middle Creek, Coll. 6, Old Man Sam Creek. ffill No. 1, Banana River. Rio Blanco.
Echinocama yaquensis Maury Plate 28, figure 5
Chama arcinella Moore, 1853, Quart. Journ. Geol. Soc. London, vol. 9, p. 130. Not of Linné.
Chama arcinella Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p. 251.
Chama arcinella Guppy, 1874, Quart. Journ. Geol. Soc. London, vol. 32, p. 531.
Etchinochama antiquata Maury, 1917, Bull. Amer. Pal., vol. 5, p. 365, pla 3, fig. 10. ' Not of Dall.
Echinochama antiquata var. yaquensis Maury, 1917, op. cit. p. 365, pl. Co gS. bi 12,
Etchinochama trachyderma Pilsbry and Johnson, 1917, Proc. Acad. Nat. Sci. Phila., p. 197.
The LEchinochama yaquensis Maury from the Miocene of Santo Domingo, where it isthe prevaling species, differs from the antzquata by the reduced number of its ribs, which in some cases are nearly obsolete and in having its surface covered with a coarse, mesh-like sculpture of raised pustules.
In Costa Rica, the antzquata is the common species, but we have also a single species of yaguensts collected from the lower Gatun near Old Harbor.
220 BULLETIN 39 394
Gatun Stage: Old Harbor. Gain (CAF
Genus DIPLODONTA, Brown
Section DIPLODONTA, s. s. Dipiodonta collina, n. sp. Plate 32, figure 15
Shell small, subcircular, convex; umbos wide, with small inconspicuous beaks, slightly posterior of the middle; anterior and posterior extremities well rounded into the base forming a part of the same circle; surface sculptured with fine, even, con- centric threads which are nearly lacking from the smooth umbos; hinge normal, and a plain ventral margin.
Length 6.75, height 6.75, diameter of the rigitjvalve 1.60 mm,
A small shell resembling the Dzplodonta nucleiformis Wag- ner of the Upper Chesapeake Miocene and the recent West In- dian fauna, but less convex and more finely sculptured.
Gatun Stage: Middle Creek.
Section FELANIELLA, Dall Diplodonta insula, n. sp. Plate 32, figure 21
Shell small, slightly convex, subcircular; beaks slightly posterior of the middle so that the posterior is a little longer than the anterior; beaks low and inconspicuous; both anterior and posterior extremities widely rounded, the posterior and the ventral margin forming a part of the same circle; surface with fine, concentric threads heaviest on the extremities, but nearly lacking from the umbonal area; interior of shell shallow, the right valve with a strongly bifid posterior cardinal; ventral mar- gin plain.
Length 6.60, height 6.50, diameter of the right valve 1.25 mm.
393 Costa RicA MrIocKNE—OLSSON + 907
A small, subcircular shell, neatly sculptured with fine, con- centric threads. YY. mznor Dall from the Bowden beds is strong- ly oblique.
Gatun Stage: Water Cay.
Genus MYRTAEA, Turton
Myrtaea limoniana Dall Plate 32, figures II, 23
Myrt@a limoniana Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, Dt6,) p. 1358, pl. 52, fig. 10.
A small, thin and slightly convex shell, its surface sculptur- ed with fine, thin and sharp, elevated, concentric lamelle. We have several specimens from the coral limestones of Port Limon as well as alarger specimen from Zone 4, of Red Cliff Creek. This later shell measures: heigth 9.50, length 10.00, diameter of the right valve 2.50mm. It occurs also in the Bowden beds of Jamaica.
Gatun Stage; Port Limon.
Coll, 7, Red “Clay creer:
Genus PHACOIDES, Blainville
Subgenus CALLUCINA, Dall Phacoides radians Conrad, var. medioamericanus, n. var. Plate 32 fig. 16
The Phacotdes radians Conrad is a fossil of the Upper Ches- apeake Miocene of the Carolinas but has continued through the Pliocene to the recent fauna. The Costa Rican fossils differ from typical vadzans in having their umbos sculptured with about 6 ribs much stronger than the concentric, while in typical radians, the ribs are more numerous and about equal to the con- centric lines. The medtoamericanus is also smaller as may be seen in the measurements given below, while vadzans is large (20 mm) and often quite convex.
Length 9.5, height 9, semi-diametes 2.5 mm.
222 BULLETIN 39 3904
Gatun Stage: Headwater of Middle creek. Old Man Sam creek, 1 mile south of the beach.
Subgenus HERE, Gabb
Section CAVILUCINA, Fischer
Phacoides trisulcatus Conrad
Lucina trisulcata Conrad, 1841, Trans. Amer. Assoc. Nat. and Geol., VOL typ. Ero:
Lucina trisulcata Conrad, 1845, Fossils of the Medial Tertairy, p. 71, pl. 40, fig. 5.
Phacotdes (Cavilucina) trisulcatus Dall, 1903, Trans. Wagner Free Inst, Sci). vol, apt. 6,\p:11360.
Few specimens from the Gatun of Middle Creek. They are identical with typical specimens from the Chesapeake Mio- cene of the eastern United States. The species also occurs in the recent fauna.
Gatun Stage: Middle creek. Old Man Sam creek.
Subgenus LUCINISCA, Dall
Phacoides costaricensis, n. sp. Plate 32, figure 5
Shell subcircular, moderately thick and but slightly convex: beaks small and inconspicuous, a little in front of the middle; dorsal area impressed and well defined; sculpture consists of strong, regular, radial threads, widely separated by interspaces 2 or 3 times the width of the threads themselves; they are cross- ed by distant but regular concentric ridges, the intersection form- ing small beads or granulations, strongest on the anterior and posterior extremities; the lunule is narrow, lanceolate and with the concentric sculpture predominanting; interior of shell cavity rather shallow, the ventral margin coarsely serrated by the ex- ternal sculpture.
395 — Costa Rica MrocENE—OLSSON 223
Length 11.50, height 11.00, diameter of the left valve BeOS ) tw,
This species is recognized by its depressed or but slightly convex shell and coarse sculpture, of which the radial is a little stronger. Phacoides cribrarius Say and nassula Conrad are both more convex, with the concentric lamellae more distantly spaced on the umbos and dominating over the radial. P. muricatus Spengler of the West Indies is much more finely sculptured.
We have also a young shell from Water Cay which is pos- sibly P. nzassula, var. caloosana Dall of the Florida Pliocene. It is very close to specimens of that species in the Cornell collec- tion.
Gatun Stage: Middle creek.
Phacoides hocasensis, nu. sp. Plate 32, figure 6
Shell thin, subcircular, depressed; dorsal area well defined by a marked change in sculpture; beak small, but distinct, placed avlittle infront of the middle; the sculpture on the disk of the shell is fine, composed of subequal, radial threads, separated by natrow interspaces; the ribs and interspaces are crossed by fine, concentric threads or ridges which produce a finely scabrous surface; the concentric threads are a little more widely spaced on the umbos; thedorsal area is defined by a stronger radial, be- yond which follows a wide radial band, strongly sculptured with the concentric threads or lamellze but lacking in radials; the up- per half of the dorsal band bears in addition to the concentric lamelle, 2 or 3 weak but scabrous radial threads; the lunule is very narrow, lanceolate and with a few strong, concentric lame- llee; interior of the shell cavity shallow, with the ventral margin finely crenulated.
Length 10:00, height'9:25, diameter of the. left valve 2.00 mm.
A depressed and very finely sculptured species whose gener- al form is that of P. muricatus Spengler of the recent West In- dian fauna. It is more regularly sculptured than the P. Aispan-
224 BULLETIN 39 396
zolana Maury of the Cercado formation of Santo Domingo but the main difference lies in their dorsal areas, that of hzspantolana Maury being sculptured more or less like the rest of the shell; in bocasensts, as described above, the sculpture of the dorsal area is dominantly concentric, only the upper half bearing 2 or 3, low but scabrous radials.
Gatun Stage: Bocas del Toro. Subgenus PARV{LUCINA, Dall
Section BELLUCINA, Dall
Phacoides actinus Dall Plate 32, figure 24
Phacotdes ( Bellucina) actinus Dall, 1903, Trans. Wagner Free Inst. Scl., VOl.{3,, pt. 6, pi) k3S5, pls 62, oes
Dall gives for his typical specimens from the Bowden beds of Jamaica, a height of 4.50 mm, while the Panama shells, all from Bocas Island, do not exceed 3 mm in height but agree in other respects. It is also recorded by Maury from the upper part of the Cercado formation of Santo Domingo.
Gatun Stage: Bocas del Toro. Genus DIVARICELLA, von Martens
Divaricella quadrisulcata d’Orbigny Plate 32, figure 20
Lucina divaricata Conrad, 1840, Fossils of the Medical Tertiary, p. 39, pl. 20, fig. 3, but not of Linné.
Lucina quadrisulcata d’Orbigny, 1846, Voy. Am. Mer., p. 584.
Divaricella quadrisulcata Dall, 1903, Trans. Wagner Free Inst. Sci.,
voll3, pt. 6, p) 1389. Lucina (Divaricella) quadrisulcata Bose, 1906, Boll. de Inst. Geol. de Mexico) numero 22, p.77,/ pl. 1%, figs. 2,12:
This species is very rare in Costa Rica. The shells are rath- er thin and generally more finely sculptured and approach in this respect the D. compsa Dall of the Florida Pliocene. Our larges specimen measures: height 14 m, length 10 mm, diameter
397 Costa Rica MIocCENE—OLSSON 225
of left valve4.50 mm. The D. prevaricata Guppy of the Mio- cene of Santo Domingo and Jamaica measures only 8 mm in height.
The D. quadrisulcata is a recent species of the West Indies appearing in the east coast Miocene of the United States for the first time in the Upper Chesapeake.
Gatun Stage:Coll. 4,5, Red Clif creek. Middle creek.
Genus CARDIUM, Linné
Subgenus TRACHYCARDIUM, Moerch
Cardium stiriatum Brown and Pilsbry Plate 27, figures 4, 5
Cardium (Trachycardinm) stiriatum Brown and Pilsbry, 1911, Proc. Acad; Nat. Sci. Phlila., vol. 63, p. 366, pl. 28; fig. 11.
This is most elegant of the several species of Cardium in the Miocene of Panama and Costa Rica. The shell is high and inflated with about 30 radial ribs which are peculiarly sculptur- ed with twisted or obliquely projecting tuberclers placed on the posterior side of the ribs and overhanging the adjacent inter- space. Large specimens may reach a height 50 mm. It is re- lated to the recent West Coast Cardium belcheri Broderip.
Gotun Stage. Gatun, C. Z. Water Cay, Panama. fill No. 1, Banana River.
Cardium dominicense Gabb Plate 27, figure 2
Cardium )Trachycardium) dominicense Gabb, 1873, Trans. Amer. Bhi’ Soc., vol. 15, p. 25:
Cardium ( Trachycardium) dominicense Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd series, vol. 8, p. 344.
Cardium (Trachycardium) gatunensis Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt Wien, vol. 58, p. 720, pl. 27, fig. 4. Not Card- tum (Fragum) gatunensis Dall, 1900.
226 BULLETIN 39 398
Cardiuin ( Trachycardium ) dominicense Brown and Pilsbry, 1911, Proc. Acad, Nat: Sct. \Phila., vol. 63,\9.\.367:
A large species abundant in the Canal Zone but usually only as internal casts. It appears to be rare in Costa Rica, our col- lection containing only a few fragments of a large individual from East Grape Point Creek. It is distinguished not only by its large size, but very numerous (60-++) fine ribs.
Gatin, Stage: Gag C7,
East Grape Point Creek, C. R. SADE, Cites, \CGZoR)
Cardium costaricanum, n. sp. Plate 27, figure 3
Shell ovate, solid, inflated; umbos wide and full with high beaks; sculpture of about 24 strong ribs, rounded or subtrian- gular in section with their apex leaning posteriorly except those on the posterior slope which lean anteriorly; interspaces narraw, formed by the sides of the ribs; the ribs are mostly smooth ex- cept those on the anterior half of the shell which have their pos- terior side more or less granulated; this granulation is found al- so on a few of the central ribs near their ventral margin; hinge narrow; internal margin fluted in harmony with the external ribs.
Length 24.50, height 34, semi-diameter 15 mm.
This species is not uncommon in Costa Rica and may easily be confused with the C. dominicanum Dall. It differs in its larger side, narrow umbos, more pointed beaks and more flatten- ed ribs.
Gatun Stage: Coll. 4, East Grape Point Creek.
Subgenus FRAGUM, Bolton
Cardium medium Linnzeus Plate 27, figure 6
Cardinm medium Vinneeus, 1758, Sys. Nat. ed. 10, p. 678.
Cardium medium Reeve, 1844, Conch. Icon., Cardium, pl. 6. fig. 30.
Cardium (Fragum) medium Dall, 1900, Trans. Wagner Free Inst. Sciivols. 3) pts 5 peor
399 Costa Rica MiocENE—OLSSON 227,
A recent West Indian species but also common as an Up- per Miocene and Pliocene fossil along the Atlantic coastal plain of eastern United States. The ribs number 24 to 27 in front of the umbonal angle. Dall has described a Cardzwm (fragum) gatunensis from Gatun, on which the ribs number only 16 in front of the truncation and 10 behind.
Gatun Stage: Port Limon.
Section TRIGONIGCARDIA Dali Cardium heredium, n. sp. Plate 27, figure to
Shell small, solid, oblique, convex and subquadrate; strong- ly carinate on the posterior-umbonal slope and truncated; umbos high and prominent; shell sculptured anterior to the truncation with 12 or 13 strong ribs, separated by interspaces about % to Y% as wide; there are 7 or 8 smaller ribs on the posterior trun- cation; the interspaces are finely sculptured with small, elevated cross-threads; the anterior set of ribs on the type specimens are wide, subequal and smooth; the posterior are unequal, with the three, bordering the umboal angulation much the smaller and bearing small and scattered pustules; on the posterior ribs where the pustules are lacking, are small dot-like depressions.
Height 11.50, length 11.50, diameter 6.00 mm.
This species approaches the Cardium apateticum Dall from the Oak Grove sands of Lower Miocene age in Florida. It dif- fers by its more oblique umbos and proportionately higher shell. The pustules are variously developed and on some shells are more strongly developed and may also appear on some of the ribs anterior to the truncation. These more strongly scalptured shells may prove with more material to belong to another species.
Gatun Stage: Rio Betey.
Banana River. fistrella River.
228 BULLETIN 39 400
Subgenus PAPYRIDEA, Swainson
Cardium spinosum Meuschen, var. Turtoni Dall Plate 27, figure I
Cardium (Papyridea) spinosum var. Turtont Dall, 1ygoo, Trans. Wag- ner Free-Inst;, Sci., vol./3; pt. 5,p. 1108!
The fossil shells from Costa Rica, are referrible to this var- lety of spinosum, distinguished by having the interspaces be- tween the ribs marked with a flat thread, between sharp grooves. This variety is also found fossil in the Chesapeake Miocene at Jackson Bluff Florida and in the Pliocene Caloosahatchie beds of the same state. It the recent fauna it is confined, according to Dall, to the eastern Atlantic, with the typical sfzmosum in the West Indies and a third variety aspersum Sowerby along the Pacific side.
Gatun Stage: Port Limon.
Subgenus LAEVICARDIUM, Swainson €ardium serratum Linnzeus Plate 27, figures I1, 12
Cardium serratum Tinneeus, 1758, Syst. Nat., ed. 10, p. 680.
Cardium (Levicardium) serratum Dall, t900, Trans, Wagner Free Insti Sci nvol spt! 5) pull kO:
Cardium (Levicardium) serratum Brown and Pilsbry, 1911, Proc Acad. Nat. Sci. Phila., vol: 63; p. 367.
Cardium (Levicardium) serratum Maury, 1917, Bull. Amer. Pal., vol. 5, P-) 370; .Pl.. 36; me 8.
Most of the fossil shells are indistinguishable from recent examples of servvatum from the West Indies. Others shells are decidely oblique and approach in form the C. sudlineatum Con- rad of the Upper Chesapeake Miocene of eastern United States. The C. venustum Gabb from the Miocene of Santo Domingo is very distinct from serratum and characterized by its faintly flex- uous outline, strong radial striz, and in having the lower cardin- al tooth large and strong.
=
401 ~ Costa RicA M1ocENE—OLSSON 229
Gatun Stage: Gatun, C. Z. Middle Creek. C. R. URtO Belay, vO. ie.
Genus PROTOCARDIA, Beyrich
Protocardia costaricensis, n. sp. Plate 27, figure 13
Shell small, rather thin, convex; beaks a little anterior to the middle, with high, inflated umbos; the anterior end is wide- ly rounded, the posterior somewhat truncated; the posterior area is well differentiated and occupies the posterior one-third of the shell and is sculptured with numerous small radial threads sep- arated by wider interspaces; these interspaces are finely sculp- tured with cross-threads; anterior of the posterior area the shell surface is sculptured with small radial threads which are finely beaded; on the anterior end of the shell, the beading is more ir- regular and is seen to be made by the concentric sculpture.
Height 18, length 21, diameter of the left valve 7 mm.
This species differs from both P. jamazcensis Dall from the Bowden beds of Jamaica and FP. zslahtspaniole Maury from the Miocene of Santo Domingo by its smaller posterior area. Itap- proaches more closely the Dominican shell but is shorter and more coarsely sculptured.
The P. Newberryana Gabb from Gatun is based on a large internal cast which is entirely unrecognizable.
Gatun Stage: Comadre creek.
Subgenus LOPHOCARDIUM, Fischer
Protocardia gurabica Maury Plate 27, figures 7-9
Protocardia gurabica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 377, pl.
26, fie ALO; Shell thin, inflated, a little longer than high and with the greatest convexity about the center of the shell; the umbos are high, prominent and wide, and project well above the hinge
230 BULLETIN 39 402
margin; the anterior end is well rounded into the base; the pos- terior end is more contracted and gaping slightly at its extremi- ty; surface sculpture on the body of the shell, anteriorly con- sists of narrow, concentric wrinkles which are best developed on the anterior end but become obsolete on the middle of the shell and posteriorly; in addition the whole surface carries fine, sub-obsolete radial threads; on the posterior area, the sculpture is of heavier and coarser radial threads, lacking the concentric, except the irregular growth lines.
Height 56, length 32, diameter 20 mm.
The subgenus Lophocardium, of which there are two recent species along the West Coast of Mexico, differs from the true Protocardia, in its gaping posterior end and lack of lateral teeth as well-as in important difference in its anatomical structure. The shells are strongly convex, with high, inflated umbos and are extremily thin and fragile.
The P. gurabica was described by Dr. Maury from the Gur- abo formation of Santo Domingo, and compared by her with the Vickburgian P. diversa Conrad, but it appears to me to bea typ- ical Lophocardium and representing the first member of this in- teresting subgenus to be found in the fossil state. As her spect- men is very fragmentary, I have redescribed the species from better preserved Panama and Costa Rican material. The poster- ior end is gaping and the internal mold shows no impression of lateral teeth along the hinge margin. It approaches closely the L. Annette Dall, dredged from 8 to 27 fathoms in the Gulf of California, except that it is proportionately longer and lacks the concentric wrinkles on its posterior area. The L. Cumingz Brod- erip, the type of the subgenus, carries between its posterior area and the general surface of the shell, a thin radial lamina or crest, extending from the umbos to the ventral margin.
The fossil shell is extremily thin. It generally occurs simp- ly as internal molds which may still retains a part of the shell or impression of its sculpture on its surface.
403 Costa RicA M10cCENE—OLSSON 231
Gatun Stage: Gatun, and Mt. Hope, C. Z. Water Cay. Margarita Trail, GO. FR.
Genus DOSINIA, Scopoli
Section DOSINIDEA, Dall
Dosinia acetabulum Conrad Plate 31, figure I
Artemis acetabulum Conrad, 1833, Fossils of the Tertiary Formations, ps 20) pli6, fig. £,
Dosinia acetabulum Bose, 1906, Boll. de Inst. Geol. de Mexico, num- ELON22).p. OL, pls 11, 8S) 7p TL:
Dosinia (Artemts) cf. acetabulum Toula, 1908, Jahrb. der K-K, Geol. Retchsanstalts Wien, vol. 58, p..727, pl. 27, figs. 3, 8a.
The Dosinia acetabulum has been recorded by Bose from Mexico and by Toula from Panama, but their determinations were based on imperfect material or casts. The Costa Rican collection however contains a large series of finely preserved Do- sinias which are unquestionably the Dosznza acetabulum Conrad of the Chesapeake Miocene.
The surface of the Costa Rican specimens are generally neatly sculptured with even, concentric bands or ribbons which on the middle of the shell disk average 2 mmin width. The form of the shell is subcircular and like the typical D. acetabu- lum of the Yorktown formation of Virginia and North Carolina. The Costa Rican shells are equally as large as the Chesapeake shell measuring in height 70 mm or more.
The Dosinia acetabulum in the eastern United States, is a most characteristic fossil of the Chesapeake Miocene, where its range extends throughout most of the lower and upper Chesa- peake formations. It is lacking from the Lower Miocene or the Alum Bluff formation of Florida, as well as from the succeeding Pliocene. Inits range through the Chesakeake Miocene, it gives rise to several well-marked varieties of which the broad
222 BULLETIN 39 404
subcircular shell of the Upper Chesapeake is the typical form. The Costa Rican shells are most like the Upper Chesapeake typical acetabulum.
Brown and Pilsbry have described a smaller and more finely sculptured species from the Gatun Spillway as delicatissima. This spceies is represented in our collection from the same place and appears to be distinct. Toula’s specimen from Gatun, how- ever, is a true acetabulum. —
The Dosinia acetabulum is a common fossil in the Gatun beds of Costa Rica, where it occurs most abundautly in the lower part of the formation.
Gatun Stage: Gatun, C.-Z.
Hotel and Hone Cree k.. Pumobri Creek.
fiill No. 37, Banana River. Coll. 6, Estrella River.
Genus CLEMENTIA, Gray
Clementia dariena Conrad Plate 31, figure 4
Meretrix dariena Conrad, 1856, Pacific R. R. Reports, vol. 5, p. 328,
pl6, fig. 55.
Clementia dariena Gabb, 1881, Journ. Nat. Sci. Phila., vol. 8, p, 344, pl. 44. fig. 16.
Clementia dariena Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. .6, p:41235.
Clementia daviena Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt, VOL. +58, (0-17.25; Pli27s, 8S... 9, LO:
Clementia dariena Hubbard, 1920, Scientific Survey of P. R. and the Virgin Islands, vol. 3, pt. 2, p. 118, pl. 19, figs 10-12.
This species is very common in the lower Gatun of the Canal Zone. Itis very rare in western Panama and in Costa Rica, where it occurs in the lower part of the Gatun formation and in the Uscari shales. Romanes* has recorded it from Bar-
* Quart. Journ. Geol. Soc. London, 1912, vol. 68, p. 125.
405 Costa RicA M10CENE—OLSSON 223
ranca near Punta Arenas onthe Pacific coast of Costa Rica, from beds probably equivalent to the Lower Gatun or to the Uscari.
The shell is very thin and fragile, and hence the specimens are very frequently much crushed and distorted in shape. Typi- cally the shell is broad and high, the beaks anterior in position and the surface sculptured with narrow, wave-like undulations which are best developed on the umbos.
Length 70, height 60. diameter 34 mm (Sousi Creek).
Wscavt Stage: Sapote, C. Z. (Gabo)
Gurun Stage Gatun, C. Z.
Nancy’s Cay, Prov. of Bocas del Toro. Upper Hone Creek, and ws tributary Soust Creek.
Genus GAFRARIUM, Bolton
Section GOULDIA, C. B. Adams Gafrarium limonensis, n. sp. Plate 32, figure 18
Shell small, subcircular, moderately convex, umbos inflated, beaks anterior to the middle: surface finely sculptured with even, concentric riblets, more or less cancellated by fine, radial threads which are strongest on the anterior and posterior extremities, while the concentric riblets predominate on the middle of the shell; lunule distinct, narrowly lanceolate; interior of the shell deep, the hinge of both valves with 3 cardinal teeth and an an- terior lateral; ventral margin concentrically grooved but not tangentially asin 7vansenella.
Length 4.75, height 4.25, diameter of the left. valve 1.50:mm.
This small Gou/ldia, is fairly abundant in the Miocene coral limestones of Port Limon, and is characterized by its small, con- vex Shell and subcircular form. Gabb has described a small Gouldia from the Pliocene beds of the Limon Peninsula, but that
234 BULLETIN 39 406
species is more depressed, less circular and more trigonal in out- line. Gabb’s species may prove to be but a small form of the recent West Indien G. cerina C. B. Adams,
Gatun Stage; Port Limon. Old Man Sam creck, 1 mile south of the beach.
Gafrarinm alium Dall, va. costaricensis, n. sp. Plate 32, figures 19, 22
cf. Gafrartum altum Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt.6) Pp. 1249 3 pli ag7erhien 5.
Shell small, high, rounded trigonal, moderately convex; beaks small and pointed slightly forward, in young shell nearly central but becoming with maturity slightly anterior; anterior and posterior extremities nearly similiar and evenly rounded; surface with fine, concentric lines, most distinct and even to- wards the ventral margin, and with faint, radial strize on the anterior and posterior slopes; lunule large, lanceolate, defined by an impressed line; interior of shell deep, with a concentrically grooved margin.
Length 4.85, height 4.75, diameter of right valve 1.50 mim.
The Costa Rican shells differ from the typical altum, de- scribed by Dall from the Oak Grove sands of Florida, in being more trigonal in form and with higher beaks. The sculpture of its surface is mainly concentric, the radials showing only as very faint striz on the posterior and anterior extremities.
Gatun Stage: Middle creek. Coll. 6, Estrella River.
Genus MACROCALLISTA, Meek
Section CHIONELLA, Cossmann Macrocallista maculata Linnzeus Plate 31, figures 6, 7 Venus maculata Vinnzeus, 1758, Syst. Nat., ed. 10, p. 686, ed. 12, p. rio! Cytherea dariena Conrad, 1857, Pacific R. R. Report, vol. 6, p. 72, pl. 5 tee 2a
407 ~ Costa R1cA MIocENE—OLSSON pal
Callista maculata Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd series, BOS, Pi, 372.
Macrocatlista (Chionella) maculata Dall, 1903, Trans. Wagner Free Tmst, Sci., vol. 3, pt. 6, p. 1256.
Macrocallista maculata Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Piila:; vol,:63, p. 370.
The Macrocallista maculata is a common and widely dis- tributed fossil in the Costa Rican Miocene and is readily recog- nized by its depressed, porcellaneous and highly polished shell. In the usual sandy phases of the Gatun, the species is quite typi- cal, but becomes high and broad in the coralline or transitional phase.
A variety cuneata has been proposed by Gabb to include shells in which this posterior-dorsal slope is straight and not humped as usual. We have seen no examples of this type.
The Macrocallista maculata isa common recent species of the West Indies, appearing for the first time, in the lower Mio- cene beds of the Chipola River, Florida.
Gatun Stage: Hleadwaters of Middle creek. Rio Blanco.
Genus CALLOCARDIA, A. Adams
Subgenus AGRIOPOMA, Dall Callocardia gatunensis Dall Plate 32, figure 1
Callocardia (Agriopoma) gatunensis Dall, 1903, Trans. Wagner Free hse. Scr.) Vols 3) pt. 6) pp: 1260; pl. 54, figt1.
Callocardia (Agriopoma) gatunensis Brown and Pilsbry, 1911, Proc. mcad. Nat. Sci. Phila: vol.63,p.) 370:
A convex, cordate shell very abundant in the Gatun beds of the Canal Zone, especially in the quarries at Mt. Hope. It is a rare species in Costa Rica, but generally larger, more trian- gular in outline and somewhat coarsely sculptured. These shells are nearest to Dall’s variety meultefilosa.
236 BULLETIN 39 408
Dall also records the species from the Miocene of Santo Do- mingo and Jamaica.
Gabb’s C. sapotensts, from Sapote, Costa Rica, we have not seen. Itisa higher and more Dosinoid shell with nearly cen- tral beaks. It is possibly but a variety of this species.
The following measurements are of Costa Rican examples:
Length 46, height 28, thickness 35 mm. 53; 44, 16 mm (right valve)
Gatun Stage: Gatun and Mt. Hope, C. Z. ffill No. 1, Banana River. Soust Creek (Upper Hone creek).
Genus PITARIA, Roemer
Section LAMELLICONCHA, Dall
Pitaria circinata Born Plate 31 figures 3, 9
Venus circinata Born, 1780, Test. Mus. Vind. p. 61, pl. 4, fig. 8.
Cytherea alternata Broderip, 1835, Proc. Zool. Soc., p. 45.
Cytherea juncea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22, Pp. 632, pli 26. fig. 13.
Chione circinata Gabb, 1873, Tarns. Amer. Phil. Soc., vol. 15, p. 250.
Cytherea juncea Guppy, 1876, Quart. Journ. Geol. Soc. London, vol. 22. pisat
Pitaria (Lamelliconcha) ctrcinata Dall, 1903, Trans, Wagner Free Inst: Sciy,/ vol. 3, pt, 6p: 1269.
Pitaria circinata Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. vol. 63, p. 370.
Pitaria (Lamelliconcha) circinata Maury, 1917, Bull. Amer. Pal., vol. 5, P- 379, Pl. 37, fig. I.
The Pitaria circinata Born is a common, recent Veneroid along the Caribbean coast of Panama and Costa Rica. It occurs also on the west or Pacific side but ina slightly different form, as the variety alfernata Broderip. ‘This variety differs from the typical cévcinata by its larger, more convex shell and more dis- tant concentric ribs.
409 Costa RicA MIocENE—OLSSON 237
The fossil shells from Costa Rica are generally more like the West Coast alternata, but are smaller and with somewhat more pointed beaks. The species is also found as fossil, in Santo Do- mingo, Trinidad and Venezuela.
Geum Stage, Gatun, C. Z. Lonel. Saury Creek, Ch.
Pitaria Hiflij Dall, var. musanica, n. var, Plate 31, figure 5
Shell elongate, moderately convex and rather solid; beaks situated at the anterior 4%, with the anterior extremity widely rounded but obtusely pointed posteriorly; lunule small, ridged in the middle and defined by a faint line; posterior-dorsal area broad and flat; surface of the shell sculptured with low, irregu- lar, rounded, concentric ribs, which are persistent along the posterior-dorsal slope but lacking elsewhere near‘the ventral mar- gin; interior of shell unknown.
Length 42, height 27, thickness (left valve) 10.50 mm.
This species has a solid shell like that of Wacrocallista. It differs from the true //7z//zz described by Dall from the Gatun of the Canal Zone, by its more rude and irregular concentric sculp- tune.
The P. planivieta Guppy, from the Miocene of Jamaica and Santo Domingo is a much shorter and more regularly sculptur- ed shell.
Gatun Stage: Hill No. 2, Banana River, C. R.
Pitaria Guppyana Gabb Plate 31, figure 11
Caryatis Guppyana Gabb, 1881, Journ. Acad. Nat. Sci. Phila., 2nd series, vol. 8, p. 373, pl. 47, fig. 73.
Shell small, ovate, moderately convex and porcellaneous; dorsal side straight; ventral side gently convex; posterior ex- tremity sub-truncate but wide, meeting the dorsal side nearly at right angles; anterior extremity much narrower, produced and rounded; lunule narrowly lanceolate, sculptured with the con-
238 BULLETIN 39 410
tinuation of the concentric ribs and separated from the shell disk by an impressed line; surface of disk sculptured with rather wide, low or appressed ribs formed between incised lines, but the shell as a whole is porcellaneous and smooth; as the ribs pass over the dorsal-posterior area towards the dorsal margin they decrease in number through fusion; interior concealed in a hard matrix.
Length 32, height 18, diameter (right valve) 5 mm.
A small species of the coralline limestones and marls of the Limon Peninsula, from which it was described by Gabb. It approaches the P. planivieta Guppy of the Miocene of Jamaica and Santo Domingo but is smaller and more rectangular in outlines. It is rather convex with a smooth and polished surface and with regular, even, concentric ribs.
Gatun Stage: Port Limon.
Pitaria VanWinklez, n. sp. Plate 32, figure 2, 3
Shell elongate, depressed; anterior extremity broadly round- ed, posterior more acute; Iunule small, lanceolate and defined by an impressed line; escutcheon long and very narrow; surface sculptured with narrow, closely spaced and reflected ribs; these ribs which number about roo on the shell disk are even on the center of the shell, but generally uneven on the anterior extrem- ity; the ribs are continuous from the edge of the escutcheon to the anterior dorsal margin; the rounded posterior dorsal slope carries two sinall, faint rows along which the concentric ribs are slightly higher and occasionally raised into small incipient spines.
Length 41.5, height 32, diameter (left valve) 1o mm.
This species is quite similiar to the P. vosea Broderip and Sowerby of the West Coast in its depressed shell and general form. ‘The concentric ribs are very regular, even and continu- ous across the entire surface of the shell. Where these ribs cross the posterior-dorsal slope, they occasionally become raised into
AIL Costa Rica MIocCENE—OLSSON 239
two rows of incipient spines. In P. vosea there is only one row of incipient spines and the dorsal-posterior area is smooth.
This species is named for Miss Katherine Van Winkle of the Paleontological Department of Cornell University and who is at present engaged on a Monograph of the American species of Veneride.
Gatun Stage: Hill No. 37. Banana River, C. R. Pitaria boucaryensis, n. sp.. Plate 31, figure 10
Shell rounded quadrate, thin and moderately convex; pos- terior and anterior extremities rounded, umbos rather large; lunule small and feebly defined by a faint, impressed line; sur- face of disk sculptured with about 4o even, distant, concentric lamelle; interspaces generally smooth or irregularly sculptured with growth lines; interior of shell concealed.
Length 31, height 22, diameter (right valve) 10 mm.
As the single type specimen is fragmentary and imbedded in a hard sandstone, the above measurements are but approxi- mate. It isa very distinctive species, differing from the other fossil Costa Rican Pitarias by its more delicate and convex shell and more distant, even, concentric lamelle.
Gatun Stage: Boucary creek, C. R. Genus ANTIGONA, Schumacher
Antigona multicostata Sowerby Plate 30, figure I
Venus multicostata Sowerby, 1835, Proc. Zool. Soc., p. 22.
Venus multicostata Reeve, 1863, Conch. Icon., vol. 11, pl. 3, fig. 9.
Cytherea (Cytherea) multicostata Dall, 1902, Proc. U.S. Nat. Mus.,
vols) 26, Dp. 390.
This is a large species living on the south or Pacific side of the Isthmus. A perfect specimen was collected from the Gatun beds of Water Cay, identical in every respect with recent ex- amples from the Bay of Panama. It is another instance among the many which we have had occasion to note in the course of
240 BULLETIN 39 412
the present study of the occurrence of West Coast species in the Caribbean Miocene, proving conclusively from a purely paleon- tological standpoint the rather late separation of the Atlantic and Pacific through the final uplift of the isthmian lands.
The recent A. Lister? Gray is a smaller, more elongate species and with finer and sharper concentric lamelle.
Length 107 min, height 102 mm, diameter 69 mm. Gatun Stage: Water Cay.
Antigona tarquina Dall Plate 30, figure Io
Venus magnifica Gabb, 1673, Trans. Amer. Phil. Soc., vol. 15, p. 249. Not of Sowerby, 1853.
Cytherea tarquina Dall, 1900, Trans. Wagner Free I[nst., vol. 3, pt. 5, pl.'38; figs: 2;2a; vol. 3, -pt.c6s'pia127 4:
Antigona tarquina Dall, 1915, Bull. go, U. S. Nat. Mus., p. 147.
Antigona tarquina Maury, 1917, Bull. Amer. Pal., vol. 5, p. 381. pl. 37, fig. 4.
A small shell identical or closely related to 4. tarquina Dall was collected from Zone 3 of Saury Creek. It is similiar to Dr. Maury’s specimen of 4. targuina from Santo Domingo.
Typical A. farquina Dall is a common fossil of the Orthau- lax pugnax fauna of the Tampa Silex beds of Florida. It was also collected by Professor Gabb and Dr. Maury from the Mio- cene of Santo Domingo.
Gatun Stage: Zone FE, Saury creek, near Cahuita, C. R.
Section VENTRICOLA, Roemer
Antigona rugosa Gmeiln Plate 30, figure 4
Venus rugosa Gmelin, 1792, Syst. Nat., p. 3276. Venus rugosa Reeve, 1863, Conch. Icon., vol. 11, pl. 7, fig. 23.
Several specimens from the coralline limestones of Port Limon. They are practically identical with recent examples of A. rugosa from the West Indies. This species is distributed throughout the West Indies but is also found living in the Gulf
413 Costa Rica M1ocENE—OLSSON 241
of California. This distribution on both sides of the Isthmus is in harmony with its occurrence as a Miocene fossil.
Gatun Stage: Port Limon. Antigona Blandiana Guppy Plate 30, figures 5, 6
Venus Blandiana Guppy, 1874, Geol. Mag., London, Decade ii, vol. 1, p..436,: pl. 17, fig: 6.
Cytherea ( Ventricola) Blandiana Dall, 1903, Trans. Wagner Free Inst. Set, .VOl.-3) pty6; p. 1277.
Antigona ( Ventricola) Blandiana Maury, 1917, Bull. Amer. Pal., vol. 5, P. 381, pl. 37, fig. 5.
This species was described by Guppy from the Bowden beds of Jamaica but also collected by the Maury expedition in the Miocene of Santo Domingo. Dall also records it from the lower Miocene of the Chipola beds and from Curacoa, Dutch West Indies. Itis asmall species of nearly circular outlines and sculptured with distant primary lamellze and finer secondary threads in the interspaces.
Our Costa Rican shells consist of two specimens, one from the coral limestones near Port Limon, the other from the lower Gatun of Comadre Creek near Cahuita. They differ from Dr. Maury’s specimens from Santo Domingo in having the primary set of lamellee more distantly spaced and their interspaces more closely striated with the secondary threads.
Length 30, height 24, diameter 9 mm (left valves) Com-
adre. Length 24, height 23, diameter 8.5 mm (left valve) Port Limon. Gatun Stage: Comadre creek, near Cahuita. Port Limon. Antigona Harrisiana, n. sp. Plate 30, figure 3
Shell small, convex and nearly circular in outline; umbos full with the beaks situated a short distance in front of the mid- dle; lunule small but broadly cordate and sculptured with the
242 BULLETIN 39 4i4
continuation of the concentric ribs and lamelle; escutcheon lan- ceolate, bordered by an angled ridge from the shell disk; disk sculptured with about 28 even, concentric ribs which appear as close undulations of the shell surface; the ribs and inter- spaces are in addition sculptured with fine lamellz-like threads, each finely crenulated or radially striated; these lamelle vary in size, those of the interspaces or troughs between the ribs are fine and 5 in number, with 2 large ones forming the tops of the ribs themselves, so that the ribs appear as if mesially divided; the radially striated character of the ribs and lamelle is much strong- er on the anterior and posterior submargins; interior of the shell deep but concealed by matrix in the type specimens.
Length 21.5, height 19.5, diameter 6.25 mm (right
valve).
This elegant species occurs in the coralline phase of the Gatun formation in Costa Rica. Its relations are with the 4. Blandiana Guppy and the recent A. sévigillzna Dall, but differs from both in its more elegant sculpture. This sculpture consists of close, regular, wave-like undulations or ribs, the crests of which carry 2, low laminz, the troughs or interspaces, 5 or 6 finer threads. ‘These fine thread-like laminze are delicately etch- ed with fine radial striations.
It is named for Professor G. D. Harris of the Paleontologi- cal Department of Cornell University. . Gatun Stage: Port Limon.
Genus CYCLINELLA, Dall
Cyclinella beteyensis, n. sp. Plate 31, figure 2
Shell thin, Dosinoid or sub-circular in form, the margins of the shell nearly a perfect circle; slightly convex; beaks small but distinct, situated at the anterior % of the shell ; sculpture with very fine, concentric threads, which are a little elevated and strongest on the anterior end, nearly smooth on the middle; lu- nule small, lanceolate and defined by a small, incised line; hinge
415 Costa RicA MIocENE—OLSSON 243
normal, the left valve with 3, narrow; cardinal teeth and a wide ligament. Height 39, length 38? diameter 5 mm left valve.
Represented by a single, imperfect specimen from the Gatun of Rio Betey. It is but slightly convex, and in form approach- es very closely a species in the Newcomb collection from the Bay of Fonseca, which is probably the C. Avoyerd Philippi. The fossil shell is more perfectly circular in form.
Gatun Stage: Rio Betey.
Cyclinella subquadrata Hauley, var. quitana, n. sp. Plate 31, fig. 8
Shell small, thin, convex and subcircular in form; beaks small but distinct, with inflated tmbo and small beak situated at the anterior 4%; the anterior end is narrow and slightly pro- duced, while the posterior end is wide and with the hinge-margin is subquadrate in form; the surface is sculptured with fine, ir- regular, concentric growth-lines, which are strongest and more regular on the posterior dorsal submargin; the interior is con- cealed.
Height 28, length 31, diameter of the right valve 7 mm.
This is a smaller and more convex species than the beteyen- sts and like that species, it seems to find its recent analogue on the Pacific side, in this case the C. sabguadrata Hanley. On the Pearl Islands in the Bay of Panama I collected a few small valves which are probably the C. sudguadrata Hanley, but they are not so produced anteriorly as is shown in Reeve’s figure. These Pearl Island shells are very similiar to the Costa Rican fossil, differing mainly in their more inflated umbos and in slight difference in form.
From the Gatun beds of the Canal Zone, Dall has described C. gatunensts, but that species is very distinct from the two Costa Rican Cyclinellas.
Gatun Stage, Outtana creek.
244 BULLETIN 39 416
Genus CHIONE, Megerle von Muhlfeld Chione Rowleei, n. sp. Plate 30, figure 2
Shell rather large, convex, ovate; lunule broadly cordate and concentrically sculptured; escutcheon long, narrow, smooth and defined by a smail cord-like ridge; surface of the disk with about 30 slightly elevated concentric ribs which on the umbos are thin and lamellar but lower on the shell disk are wide and trianglar at the base, but with a thin zigzag or fluted lamellar ridge on top; the interspaces, much wider on the umbos is sculp- tured with even rib-like cords which also flute in harmony the dorsal face of the concentric ribs; the ventral face of the ribs are finely striated by radial incised lines which form cord-like ridges along % as wide as the radial cord of the interspaces; interior of the shell deep, with a small pallial sinus and subequal adductor scars; basal margin finely crenulated as well as the lunular mar- gin and the extreme posterior extremity.
Length 42, height 37.5, thickness (right valve) 17 mm.
This large, elegant Chione is quite common in the Gatun beds of Costa Rica. It also occurs in the lower Gatun at Gatun, usually in the form of casts. It recalls in a general way, the re- cent West Coast C. amathusia Philippi, but differs in its heavier shell, higher and fuller umbos and difference in details of its spiral and radial sculpture. In the Costa Rican shell, the radial cords are simple, relating it more closely with the small C. Woodwardi Guppy of the Miocene of Jamica and Santo Domin- go, while in C. amathusta, they are double and alternating.
The C. tegulum Brown and Pilsbry we have not seen. It is a smaller shell, related to the C. Woodwardi. In C. tegulum the concentric lamellz are much closer and the wide cordate lunule is sculptured only with radial lamellee.
It is named for Professor W. W. Rowlee, who collected a few specimens from the Banana River in the course of botanical investigation of the Balsa tree (Genus Ochroma), a very light wood used in the construction of life-buoys, ete.
417 Costa RicA MiocENE-—-OLSSON 245
Galun Stage: Gatun, C. Z. Banana River. Coll6," strella: River,
Chione costaricensis, n. sp. Plate 32, figure 4
Shell of moderate size, convex, ovate-trigonal; the dorsal portion of the type specimen is lacking; the surface is sculptured with fine, subregular, concentric lamellee spaced on the center of the shell disk about .75 mm apart; the interspaces and the ven- tral surfaces of the lamellz are marked with regular, incised lines forming radial bands of an average width of .40 mm; a faint rad- ial sinus extends from the beaks to the posterior; ventral margin of the shell is slightly arcuate at its posterior end; interior of ventral margin finely crenulated.
Length 34, height 29, diameter of the right valve Io mim.
This shell will be distinguished from the other Costa Rican Chiones by its more crowded, concentric, lamelle and regular, radial striz. The striz occur on the ventral faces of the lamel- lz and on their interspaces.
Gatun Stage: Hill No. 3, Banana River.
Section LIRGPHGRA, Conrad
Chione mactropsis Conrad Plate 30, figures 7, 8
Gratelupia? mactropsis Conrad, 1856, Pacific R. R. Reports, vol. 5, p. 329, pl. 6), fig.) 54.
Chione (Lirophora) mactropsis Dall, 1903, Trans. Wagner Free Inst. Sci, vol. 3, pt. 6, p. 1294.
Chione (Lirophora) mactropsis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p, 369.
Chione (Lirophora) ulocyma Brown and Pilsbry, 1911, /dem, p. 369. Not of Dell. 1903.
Until the construction work on the Panama Canal had com-
menced in earnest, the abundant and finely preserved fossils of
246 BULLETIN 39 418
the Canal Zone, so easily obtained at the present time, were not available, and the only fossil remains collected by the early trav- vellers were generally in the form of poorly preserved casts. In 1856 there appeared in the Pacific Rail Road Reports, the de- scription of an internal cast as Gratelupia? mactropsis, together with scarsely recognizable 7Jurritella gatunensis and altilira.
These specimens obtained by Mr. W. P. Blake from the Isthmus are the first noticed from the fossiliferous rocks in Panama and Central America.
The Livophora mactropsis described by Conrad from an in- ternal cast as Gratelupia ?, is one of the most abundant fossils in the Gatun beds of the Canal Zone, associated with the small- er and more finely sculptured holocyma Brown and Pilsbry. JL. mactropsts is a species of variable size, but moderately convex, longer posteriorly and usually with the basal margin more or less arcuated about the posterior extremity. The sculpture con- sists of irregular, more or less confluent concentric lamelle, which are overrun by fine radial lines. These radial lines may remain strong or become obsolete as usual with specimens from the Canal Zone. A narrow foliaceous band is developed on each side of the escutcheon and another about the anterior extremity below the lunule. The Costa Rican examples are often very much larger than those of the Canal Zone, the concentric lamel- le are finer and less confluent and with more persistent radial lines. ‘The following measurements will illustrate the range in size:
Length 30, heigth 23, thickness 15 mm, Central Zone.
6 66
37) 28, IQ
AN: Be. 9.5 mm (right valve) Old Man Sam Creek, C. R.
51.5, 38, 21.5 mm, Sousi, C. R.
Gatun Stage: Gatun, and Mt. Hope, C. Z. Old Man Sam creek, C. R. Soust creek (Upper Hone creck) Banana River.
419. Costa RicA MIOoCENE—OLSSON 247
Chione holocyma Brown and Pilsbry Plate 30, figure 9
Chione ulocyma Dall, variety holocyma Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 369:
Associated with mactropsis, in the Central Zone is a more convex and finely sculptured shell, allied to the ulocyma Dall of of the Floridian Miocene. It is the holocyma of Brown and Pils- bry. We have seen no typical wlocyma from Panama.
From mactropsis, the holocyma will be distinguished by its generally smaller, more convex and more circular shell. Its concentric lamelle are more numerous and less confluent and with fine but persistent radial strize. It is generally best distin- guished by its smaller and less clearly differentated foliaceous area on its anterior extremity, which in mactropsis is as wide an area as that along its posterior dorsal margin.
Heigth 29, length 36.5, thickness 19 mm. Gatun Stage: Mt. Hope, C. Z.
Chione hotelensis, n. sp. Plate 32, figure 7
Shell small, subtrigonal, heavy, but slightly convex; sculp- ture of about 15, thick, rounded and evenly spaced concentric lamellze; on the posterior-dorsal slope, the ends of the lamellz become attenuated, thin and somewhat appressed as in glyptocy- ma Dall of the Flordian Miocene; the lamelle themselves are smooth and rounded but with their interspaces finely radially in- cised by small lines; escutcheon rather large, smooth; lunule small, cordate and smooth except for the growth lines,
Length 23, height 19, thickness 12.5 mm.
This elegant little species is distantly related to the C. gly- plocyma Dall from the Oak Grove sands of Florida, but isa smaller and more compact shell. The radial striations commence on the ventral surface of the lamellz and are continued across the interspace to the base of the next rib.
All of our specimens were collected in a hard sandstone in the lower Gatun of Hotel Creek near Old Harbor, Costa Rica.
248 BULLETIN 39 420
Uscari Stage: Hotel creek, C. -R.
Chione chiriquiensis, n. sp. Plate 32, figure 9, Io
Shell small, solid, depressed; lunule rather large, cordate, smooth; escutcheon large, smooth and defined by an angled ridge; surface of the disk sculptured with about 8, large, irregu- lar ribs, like those of Chione latilirata: these ribs do not extend to the edge of the escutcheon, but commence a short distance in front leaving a narrow band which is smooth and sculptureless; the ribs are of variable size, flat-topped and smooth; they are rounded on their ventral. side, but shelving or recurved back- wards on the dorsal face; interspaces smooth or only sculptured with fine growth-lines; interior of shell of moderate depth, with a small pallial sinus and with a slightly larger posterior adduct- or scar; basal margin finely crenulated.
Length 13.75, height 12.50, diameter 9.50 mm.
The few specimens which we have from Water Cay, are probably immature, but they possess such distinctive characters that the species cannot be mistaken.
The heavy, irregular concentric ribs recall certain varieties of Chione latilirata Conrad. ‘They differ in being abruptly cut off or flattened a short distance behind the dorsal-posterior angle leaving a band asin C. paphia Linnzeus, but smooth and sculp- tureless. The interspaces are smooth,
Gatun Stage: Water Cay.
Section TIMOCLEA, Brown Chione grus Holmes, var. Plate 26, figure 21
Tapes grus Holmes, 1858, Post-Pliocene Fossils of S. Car., p. 37, pl. 7 fig. 5. Chione ( Timoclea) grus Dall, 1903, Trans. Wagner Free Inst. Sci., vol. 3, pt. 6, p. 1299. We have but a single and not entirely perfect specimen of a Timoclea closely related to C. gvus Holmes. It is somewhat
AAS st ee ne =
A@I Costa Rica M1i0ocENE—OLSSON 249
more convex than typical C. gvus from the Carolinas and more finely sculptured. More material is needed to show whether or not these charaters are constant. The C. gvus appears asa fossil in the Upper Chesapeake Miocene of the eastern United States and is recent from Cape Hatteras to Yucatan.
Gatun Stage: Coll. 4, Red Cliff creek.
Genus TELLINA, (Linné) Lamarck Tellina revala, n. sp. Plate 26, figures 7, 8
Shell elongate, slender, subequilateral with the beaks situ- ated near the center of the shell; anterior side long, with a straight dorsal margin and narrowly rounded at its extremity; posterior side much narrowed and rostrate, the left valve with 2 radial bands forming the rostrum, the dorsal one a fold, the oth- era wide sinus; ventral margin rounded anteriorly, nearly straight about the middle and a wide sinus posteriorly; the sur- face is sculptured with even, concentric lamellz; this sculpture on the rostrum is heavier and on crossing the fold and sinus be- come reversed in position.
Length 29, height 12, semi-diameter 2.5 mm. 207 14 3) iain,
We have two specimens of this typical 7eldina from Water Cay. The larger specimen is fragmentary but probably measur- ed when complete about 36 mm in length. The species recalls the recent West Indian 7. zuterrupta Wood (7. antonii Philippt) in its form and sculpture but lacks the finer radiating strie.
Gatun Stage: Water Cay.
Subgenus ARCOPAGIA, (Teach) Section MERISCA, Dall
Tellina crysiallina Wood Plate 26, figure 10
Tellina crystallina Wood, 1815, General Conchology.
iS) cn Q
BULLETIN 39 422
Tellina ( Merisca) crystallina Dall, 1901, Proc. U. S. Nat. Mus., vol. 23, DS hL Wi pliyi2) on, TO}
Tellina ( Merisca) crystallina Maury, 1917, Bull. Amer. Pal., vol. 5, p.- 387, pl. 38, fg. 4:
This 7ellina, the type of the section Merisca, is recognized by its thin, depressed shell, strongly sculptured with distant, but egular, raised, concentric lamelle. It is very rare as a fossil, our collection containing but a single specimen from the Gatun beds of the Banana River. It has previously been recorded by Maury as a Miocene fossil in Santo Domingo.
In the recent fauna, 7ellina crystallina is distributed along both the Atlantic and Pacific coasts of Central America. I have collected a few specimens from the beaches of Bocas Island and more abundantly at Bucaru, Province of Los Santos, on the Pacific side of Panama.
Gatun Stage: Hill No. 2. Banana River. Tellina nasua, n. sp. Plate 26, figure 17
Shell sub-rhomboid, slightly convex, thin; beaks small, in- conspicuous, posterior to the middle; the anterior end longer, the dorsal-anterior dorsal margin descending, the extremity rounded into the gentle curve of the base; (the posterior side broken in the type specimen); the greatest convexity of the shell lies along the broad, umbonal slope anterior to the beaks; the surface is finely sculptured with regular, sharp, concentric ridges, separated by narrow, band-like interspaces; interior of the right valve with a large pallial sinus which extends to the anterior % of the shell; hinge appearing normal for the section, but partly broken away, but showing a large anterior lateral tooth.
Length 20? Height 15.25, diameter of the right valve gwOe)) scalaoy
Our specimen if fragmentary, the posterior portion of the shell being broken away. There is however little doubt that the species is a true J/erisca, allied to 7. egutstriata Say and
423 | Costa Rica M1ocENE—OLSSON 251
T. lintea Conrad. It differs from eguzstriata in its greater dor- sal anterior slope and finer sculpture, and from 7. /zxtea of the recent fauna, in its somewhat greater proportional height and greater convexity.
Gatun Stage: Coll. 4, Red Cliff creek.
Section PHYLLODINA, Dall Tellina lepidota Dall Plate 26, figures 4, 5
Tellina (Phyllodina) lepidota Dall, 1900, Trans. Wagner Free Inst. Sci., vol 3, pt. 5, p. 1022, pl. 46, Go. 18.
Dall’s type of this species is based on a fragmentary right valve from Gatun. We have several complete and large speci- mens from Water Cay, where the species is fairly common. The right valve is flat and depressed, the left more convex. Both valves are nearly similarly sculptured with 2 or 3 wide undul- ations of the beaks, which are soon replaced by even, sharp, con- centric lamellz, of which 3 or 4 are higher and more lamellose onthe dorsal margin.
Length 24, height 12.5, diameter of the left valve 2.75
mm. Length 21.5, height 11.00, diameter of the right valve PiaG tii.
Gatun Stage: Water Cay.
Section EURYTELLINA, Fischer
Tellina costaricana, n. sp. Plate 26, figures 6, 9
Shell oblong, depressed; beaks small and inconspicuous about the middle of the shell so that the anterior and posterior ends are of nearly the same length; posterior end widely round- ed the anterior but slightly narrowed and less round- ed at its extremity; anterior and posterior dorsal margins nearly straight; surface of the disk finely sculptured with even, con-
252 BULLETIN 39 424
centric lines which are slightly lamellose on the posterior-dorsal margin; interior concealed.
Length 32, height 19, semi-diameter 2 mm.
This species is related to the recent West Indian 7. angulosa Gmelin and its Pacific analogue, the 7. rubescens Hanley. ‘The fossil shell differs by its more central beaks and sharper concen- tric sculpture, which is continued across the entire surface of the disk.
Gatun Stage: Hill No. 3, Banana River.
Tellina dariena Conrad Plate 26, figure 3
Tellina dariena Conrad, 1857, Pacific R. R. Reports, vol. 5, p. 328, Pl G6. tig 53:
Tellina semtlevis Gabb, 1861, Proc. Acad. Nat. Sci. Phila., vol. 12, p. 567.
Tellina Dariena Gabb, 1881, Journ. Acad. Nat. Sci. Pnila., vol. 8, p. 343, pl. 44 fig. 13.
Tellina Rowlandi Toula, 1908, Jahrb. der K-K. Geol. Reichsanstalt Vol. 58, p. 728, plu2zs.fe.-11.
Tellina Rowlandi Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila., vol. 63, p. 368.
Conrad’s figure of 7el/ina daricna is very poor but probably represents this species, the most abundant 7Zed/zma at Gatun. Conrad’s figure shows a subtrigonal shell, with a produced and truncated posterior extremity. In 1861, Gabb described TZel/ina semulevts together with Arca chiriqguiensts from the Gatun beds of the Chiriqui Lagoon. This species Gabb however later con- sidered synonymous with the earlier Zellina dariena of Conrad. Toula’s 7Zellina Rowland is certainly identical with Gabb’s Tellina semilevis as figured by Gabb.
Shell but slightly convex, subequilateral with the beaks sit- uated a short distance posterior to the middle of the shell; anter- ior and posterior dorsal slopes straight and evenly descending; posterior end rounded and subtruncate, the anterior rounded ; beaks low; surface striated with very fine concentric lines and
425 Costa RicA MI0CENE—OLSSON 253
the shell may sometimes appear nearly smooth and polished especially on the umbonal areas; hinge of the left valve with a bifid anterior cardinal, a small posterior and 2 laterals; the an- terior adductor is bordered on the posterior side by a thickened internal ray; pallial sinus long, ascending above in the middle of the shell and touching the anterior adductor scar. Length 34, height 19.5, semi-diameter 3.50 mm. Gaiun Stage. Gatun, C. Z.
Water Cay. Chiriqui Lagoon and Oronli creck (Gabo)
Subgenus ANGULUS, Mergerle
Section ANGULUS, s. s. Tellina umbra Dall Plate 26 figures 12, 15
Tellina (Angulus) umbra Dall, 1900, Trans. Wagner Free Inst. Sci., wok. 2, pt, 5,) p. 1033, pl. 46, fig. 13.
The Costa Rican specimen consists of a small individual with both valves. Itis somewhat smaller than typical umdéra, from the Upper Chesapeake Miocene of the Carolinas, but agrees in form and in the fine surfrce sculpturing.
The Zellina umbra is a common fossil in the Upper Chesa- peake Miocene of the eastern United States and in the Pliocene of Florida, It is closely allied tothe recent 7Tel/ina sybaritica Dall of the West Indies, differing most importantly in its less an- gular posterior extremity,
The Costa Rica specimens measures:
Length 8.00, height 4.15, diameter 2.25 mm.
Gatun Stage: Coll. 4g, Fast Grape Point creek. Section SCISSULA, Dall
Tellina cercadica Maury
Tellina (Scissula) cercadica Maury, 1917, Bull. Amer. Pal., vol. 5, p. 388, pl. 38, fig. 9.
254 BULLETIN 39 426
The Costa Rican specimens are fragmentary but there can be little doubt of their identity with the TZellina cercadica Maury from the Miocene of Santo Domingo.
The shell is thin and strongly depressed. The surface is polished and obliquely striated with fine lines. It is related to the recent Pacific 7ellina delicatula Deshayes.
Gatun Stage: Coll. G, Saury creek. Estrella River.
Genus TELLIDORA, Moerch Tellidora cristata Recluz Plate 26, figurce 1, 2
Lucina cristata Recluz, 1742, Revue Cuvier., p. 270. Tellidora cristata Dall, 1990, Trans. Wagner Free Inst; Set, vols)3: PENA) Pp. TOR]:
A recent species of the West Indies and previously known as a Pliocene and Pleistocene fossil from Carolinas. The fossil Costa Rican shells-are frequently larger than the recent form and with the dorsal margins more nearly in line. It is not rare in the form of impressions but the perfect shell is seldom found.
Gatun Stage: Hills ra, 2 and 3, Banana River.
Genus STRIGILLA, Turton Strigilla pisiformis Linné
Tellina pisiformis Linné, 1758, Syst. Nat., ed. 10, p. 677.
Strigilla pisiformis Dall, 1900, Trans. Wagner Free Inst. Sci., vol, 3, pt. 5, p. 1038. AUN Strigilla pisiformis Maury, 1917, Bull. Amer. Pal., vol. 5, p. 389, pl.
39, fig. 6.
This species is rare asa fossil in Costa Rica. It is very common asa recent shell on the sandy beaches along the en- tire north coast of both Panama and Costa Rica, where it occurs with Donax denticulata Linné and afew other species. It is
427 Costa RicA MIocENE—OLSSON 255
also found as a Miocene fossil in the Bowden beds of Jamaica and in Santo Domingo.
Gatun Stage: Middle creek.
Strigilla musanica, un. sp. Plate 26, figure 18
Shell subtrigonal, moderately convex, subequilateral; the umbos are broad and wide, situated about the middle of the shell, with the greatest convexity of the shell lying along the anterior- umbonal slope; the anterior end is broad and rounded, the pos- terior slightly narrow, but not produced; the sculpture is divid- ed between three areas, the posterior, central and anterior, each of which is set off from the adjacent one, by a marked change in sculpture along a well-defined line; on the anterior area, it con- sists of a concentric series of wave-like incised lines, the crest of
the wave lying near the posterior side and directed as usual dor-
sally; on the central area, the lines are vertically directed, com-
mencing on the lines bordering the posterior side; on the poster-
ior area, the lines commence on the anterior side and pass verti-
cally, forming therefore with the middle lines very acute an-
gles; the interior is concealed in our specimen. | Length 13.00, height 11.50, diameter of the right valve
3.00 mm.
Differs from S¢vigzlla carnaria Linné of the recent West In- dian fauna, by being more inequilateral, more convex and in de- tails of its sculpture whicn is sharply divided between three areas as described. The interior of the type specimen is concealed, so that the features of the pallial line and sinus cannot be deter- mined.
Gatun Stage: [fill No. 3, Banana River. Genus METIS, H. and A. Adams
Metis barba, n. sp. Plate 29, figures 3, 4
Shell subquadrate, inequilateral, with the beaks situated at the anterior two-thirds of the total length of the shell; anterior
256 BULLETIN 39 428
end longer with a sloping dorsal margin and rounded anterior extremity; posterior side shorter, with a dorsal margin sloping more rapiding to the wide and flexed posterior end; the left valve is convex but depressed along the narrow posterior submargin or wing; the right valve is less convex with a broad depressed area extending from the beaks to the ventral margin; this depressed zone is sharply separated from the posterior-dorsal slope or wing by a strong angle or flexure which corresponds to the sinus in the middle of the wing on the left valve; surface finely striated with concentric growth-lines.
Length 48, height 37, diameter 19 mm.
Pilsbry and Johnson have described without figuring, two species of Mefis from the Miocene of Santo Domingo (the M. efferta and postrema). These species, they compare with the M. chipolana Wall, from the Chipola beds of Florida and men- tion in their description the nearly central location of the beaks. In the Costa Rican shell the beaks are at the posterior one-third of the shell. It is very distinct from the AZ. chifolana Dall. JZ. trinttavia Dall from ‘Trinidad and Porto Rico differs in its more central beaks, fuller umbos and has the right valve much less flexed posteriorly.
Gatun Stage: Betey creek.
Genus MACOMA, Leach Macoma costaricana, n. sp. Plate 26, figures 14, 16
Shell broadly ovate, thin and slightly inflated; dorsal and ventral margin straight and parallel; the posterior dorsal side sloping to the rounded ventral extremity; anterior extremity rounded; the posterior side is very slightly flexed; surface smooth except for very fine, incremental growth lines; interior of shell concealed.
Length 29, height 19, semi-diameter 4.5 mm. The reference of this species is Wacoma is doubtful as the
429 Costa Rica MIocENE—OLSSON 257
hinge is completely concealed in all our specimens. It has the form of the J7. Conradi Dall, from the Chesapeake Miocene of United States but is higher and differs in many other respects. It is shorter and higher than IZ. gatunensts Toula with which it may occur.
Gatun Stage; Hill No. 3, Banana River. Rio Betey.
Subgenus PSAMMACOMA, Dall Macoma gatunensis Toula Plate 26, figure 13
_Macoma ( Tellina) gatunensis Toula, 1908, Jahrb. der K-K Geol. Reich- sanstalt, vol. 58, p. 729, textfigure Io,
Tellina gatunensis Brown and Pilsbry, 1Igt1, Proc. Acad. Nat. Sci. Whila,, vol. 63, p: 3268.
This shell is extremely similiar to the recent J. elongata Hanley of the Pacific coast of Panama and I have failed to find any character which will distinguish the fossil shells from the recent ones. Our shells were all collected loose near Hill 3, of the Banana River and were probably derived from beds low in the Gatun formation.
Gatun Stage: loose, near Hill 3, Banana River (Terry).
flacoma panamensis Dall, var. canalis, n. var, Plate 26, figure 11
Cf. Macoma (Psammacoma) panamensis Dall, 1900, Proc. U. S. Nat, Mus.) vol: 235 p. 324) pl. 4, fig. 3.
The MWacoma panamensis Dall was described from a left valve dredged from 33 fathoms of water in Panama Bay. The fossil shell, also a left valve from the Gatun of Mt. Hope, differs most importantly from Dall’s figure in being somewhat higher. The shell is thin and extremely fragile, with the surface simply scultured with the fine growth lines. The interior is concealed so that the characters of the pallial sinus cannot be seen. The specimens measures as follows:
258 |) S BULLE DING 39 430°
Length 30.50, height 15.00, diameter of the left valve 3.00 mm.
Gaiun Stage: Mt, fope; C. 2.
Genus SEMELE, Schumacher
Semele levis Sowerby, var. costaricensis, u. var. Plate 29 figure 1
Cf. Amphidesma levis Sowerby, in Reeves, 1853, Conch. Icon., pl. 7, fig. 50.
Shell large, subelliptical, but slightly convex; beaks very nearly central, the posterior side wide and well rounded, the an- terior somewhat narrowed, contracted and slightly the longer; the surface is nearly smooth and porcellaneous, faintly marked on the anterior and posterior extremities by fine, concentric lines.
Length 72, height 65.50, semi-diameter 8 mm.
This large Semele is quite common in the Lower Gatun beds of the Banana River. It is closely related to the recent Pacific — S. levis Sowerby, but differs in being slightly longer and more evenly rounded at its posterior extremity.
Gatun Stage: Hill No. 3, Banana River. fill ra, Banana River.
Semele Sayi Toula Plate 29, figure 8
Semele Sayt Toula, 1908, Jahrb. der K-K Geol. Reichsanstalt, Wien, vol. 58; p.'730, pl.i28, fig.) 17.
Semele Sayt Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila. vol, 63, p. 368.
Shell convex, rather thin, with the umbos nearly central. The anterior side is somewhat narrow or contracted, and round- ed at its extremity. The surface is sculptured with fine, hair- like, concentric threads. "The species was described by Toula from the Canal Zone. Our record from Costa Rica is based ona single, imperfect specimen from Sousi creek, where it occurs as- sociated with Clementia, Chione mactropsis and Turrttella altilira
431% Costa R1cA MI0CENE—OLSSON 259
variety costaricensis in the lower part of the Gatun formation.
Gatun Stage: Sousi creek.
Semele Claytoni Maury Plate 29, figure 2
Semele Claytont, Maury, 1917, Bull. Amer. Pal., vol. 5, p. 227, pl. 35, fig. 9.
The S. Claytont was described by Dr. Maury from the Cer- cado formation, Santo Domingo of Lower Miocene age. Itisa large species, reaching a length of 66 mm or more and charact- eristically sculptured with even, concentric lamellz like the re- lated species from the Tertiaries of eastern United States as chipo- lana Dall, Leana Dall and perlamellosa Heilprin. From these, the S. Clayfonz is distinguished in having its radial strize nearly obsolete. Our specimen from Costa Rica is fragmentary but probably measured when perfect 38 by 30 mm.
Uscart Stage: Hotel creek.
Genus ABRA, (Leach) Lamarck
Abra triangulata Dall Plate 29, figure 10
Abra triangulata Dall, 1900, Trans. Wagner Free Inst., Sci., vol. 3, pt. 5, p. 997, pl. 49, fig. 4.
A small, rotund or trigonal species abundant in the shale beds of Bocas Island. The valves are nearly equilateral, the an- terior end slightly the larger and inflated. Surface smooth and shining. The type was described by Dall from the Bowden beds of Jamaica. Our specimens are a little larger, measuring:
Weneth 7.25, herght 6:50, diameter of the left. valve He 75 Wi.
Gatun Stage: Bocas del Toro.
260 BULLETIN 39 432
Genus PSAMMOSOLEN, Risso
Psammosoien gatunensis Toula, var. costaricensis, n. var.
Plate 29, figure 7
Solecurtus gatunensis Toula, 1908, Jahrb. der K-K. Geol. Reichsan- Stalt, vol: 58; ~p. 732, pl: 28, fiz: 12.
Solecurtus gatunensis Brown and Pilsbry, 1911, Proc. Acad. Nat. Sci. Phila, vol. 58; p- 372:
Shell rather large, elongate, convex, the dorsal and ventral margins straight and parallel; the low, inconspicuous beaks sit- uated at the anterior %, the anterior end rounded, the posterior subtruncate; two low, radial depressed rays extend from the beaks obliquely to the ventral margin; the central part of the shell and the anterior end is sculptured roughly with irregular growth lines; the posterior dorsal slope has strong, vertical or slightly oblique lines which cross the finer concentric growth lines; in addition the whole surface of the shell is roughened by small, short threads as if the surface had been painted with a stiff brush.
Length 63, height 25.50, diameter of the right valve 14.50 mm.
Toula’s figure of an internal mold shows a smaller shell, a strong anterior-umbonal angle and a somewhat shorter anterior end. Itis possible that the two forms are distinct but we lack a good comparative series of the Gatun shell. A smaller and more delicate species of Psammosolen occurs at several localities in the Costa Rican Gatun. Itis not the P. strigdllatus (L) of Toula, differing in its pointed anterior end.
Gatun Stage: Rio Betey. Genus SANGUINOLARIA, Lamarck Section SAMMOTELLA, Blainville
Sanguinolaria alouatta, n. sp. Plate 29, figures 5, 6
Shell rather large, elongate; and rostrated behind; left
433 CosTaA RicA MIocCENE—OLSSON 261
valve nearly flat, the right valve much more convex; the right valve carries a strong umbonal keel trom the tip of the beaks to the posterior extremity; this keel is but poorly developed on the left valve; anterior end broadly rounded with the dorsal and ventral margins nearly parallel; the posterior side is much nar- rower and with its extremity pointed; the surface is smooth; the dorsal-posterior slope of the left valve is less smooth and carries coarse, irregular growth-lines; the interior is not known.
Length 54, height 30, semi-diameter 4 mm (a left valve) Length 60, height 27.5 semi-diameter 7 mm (a right valve)
This species stands intermediate between the S. Smzthwood- wardt Maury from the Miocene of Santo Domingo and the re- cent West Indian S. oferculata Gmelin. It differs from the Smzth- woodwardt, in being much larger with a less depressed left valve: From oferculata the Costa Rican shell has higher and more pointed beaks as well as being longer posteriorly. In full-grown specimens of oferculata, the surface generally shows radial lines and strize which may become quite strong near the posterior ventral margin. The Costa Rican shells are smooth except for the growth lines.
Gatun Stage: Hill No. 3, Banana River.
Genus TAGELUS, Gray Tagelus cebus, n. sp. Plate 29, figure 9
Shell elongate, moderately convex and solid; beaks central with the anterior and posterior ends nearly equal, and with the dorsal and ventral margins parallel and nearly straight; anterior and posterior extremities subtruncate; a broad depressed zone extends from the umbos to the ventral margin of the valves; surface marked with irregular concentric lines of growth; inter- ior concealed. |
Wength' 36, height 13, diameter jof the left valve 3.00 mim.
262 BULLETIN 39 434
The Szligua subegualis.Gabb, from the Miocene of Santo Domingo is doubtless a 7Zagelus, belonging to the Section JZes- opleura Conrad. It differs from the present shell in being thin- ner, more depressed and more pointed posteriorly. Gabb’s species approaches very closely the recent 7. dzvisus Spengler of the West Indian fauna, judging from the specimens in the Maury collection at Cornell, differing mainly in its larger size.
The interior of the Costa Rican shell is concealed so that the presence or absence of an internal rib cannot be determined. Its heavier shell and general form, is more that of typical Zage- lus differing from the recent gzjdus Spengler in its more central beaks.
Gatun Stage: Rio Betey. Genus MACTRA (L.) Lamarck
Subgenus MACTRELLA, Gray
Mactra exoleta Gray Plate 28, figure 2
Mactra exoleta Gray, 1837, Mag. Nat. Hist., p. 372. Mactra exoleta Reeve, 1854, Conch. Icon., Mactra, fig. 16.
Comparision of our fragmentary shell with recent examples from the Bay of Panama show no inportant differences. The species is notable for its high, narrow, inflated umbos, strongly depressed posterior-dorsal margin and alation of its dorsal-pos- terior angle.
At the present time, this species is restricted to the Pacific side, but represented on the Caribbean coast by the next, the Mactra alata Spengler, common along sandy beaches. It is therefore of more than usual interest to find these two species, together in the Miocene beds of the Caribbean slope of Costa Rica.
Gatun Stage: Boucary creek.
A435 CosTA RicA MIoCENE—OLSSON 263,
Mactra alata Spengler Plate 28, figure 3
Mactra alata Spengler, 1802, Skriv. Naturch. Selsk., vol. 5, part 2, Pp. 99.
Mactra alata Reeve, 1854, Conch. Icon., Wactra pl. 8, fig. 29.
Mactra (Mactrella) cf. alata Maury, 1917, Bull. Amer, Pal., vol. 5, Pp. 395-
A thin, fragile species distinguished by its sharp dorsal-pos- terior angle and hatchet-shaped shell. It is related to the exo- leta, already referred to, but is a very much broader shell. In exoleta, the dorsal-posterior area is flat or concave, with a slight raised line which is bounded on each side by a markly depressed or sunken band. In a/aza this area is convex.
It is living today on the Caribbean side of the Isthmus, be- ing quite common on sandy beaches.
Gatun Stage: Boucary Creek. flill No. 3, Banana River.
Section HARVELLA, Gray
Mactra estrellana, n. sp. Plate 28; figure 1
Shell rather large, ovate, thin, slightly convex; surface reg- ularly, concentrically plicated; the fragmentary type specimen about 38 mm in height has about 32 plicae which on the center of the shell disk are spaced about 2 mm apart; these plicze com- mence as in typical H/arvella at the dorsal-posterior carina which bounds the escutcheon and extends across the surface to the an- terior extremity; the surface is, in addition, finely, concentrically striated; lunule large elliptical, smooth or finely striated; escut- cheon smooth; interior not known.
Length 50, height 38, semi-diameter 8 mm, Although this is not a very rare species in the Costa Rican
Miocene, its thin, delicate shell renders the collecting of good specimens very difficult and it is usually secured only as distort-
264 BULLETIN 39 436
ed internals molds. Itis related to the recent Pacific “7. elegans Sowerby, but differs in being less convex, wider and with more numerous and closer concentric plicz.
Gatun Stage, Coll. 6. Estrella River. Coll. 2, QOuttana creek.
Mactra sincola, n. sp. Plate 28, figure 4
Shell thin, broadly subtrigonal, depressed, nearly equilater- al; beaks small and pointed anteriorly; lunule absent; escutche- on linear-lanceolate and defined from the shell disk by an elevat- ed carina; the surface is sculptured with strong, wave-like, con- centric plicee which are continuous from the anterior-dorsal mar- gin to the edge of the escutcheon; on the type specimen there are about 30 plicee on the center of the shell disk, spac ed about 1.75 mm apart; the intervals between the concentric plice are deep and trough-like and in width equal to the plicze them- selves; in addition the whole surface of the shell is finely sculp- tured with small, radial, vermiculate threads, best seen on the summits of the plicze; the escutcheon is ridged on the middle and smooth; the hinge, largely concealed in the sandy matrix, is mactroid.
Length 36, height 27, diameter of the right valve 5.5 mm.
The systematic position of this interesting shell is very doubtful. Its general surface sculpture of strong, concentric plicze and well-defined escutcheon resembles that of Harvella. The surface is covered with fine, vermiculate threads as seen on Labtiosa.
Gatun Stage: Saury creek. Genus LABIOSA, (Schmidt) Moller
Labiosa lineata Say Plate 21, figure II
Lutraria lineata Say, 1821, Journ. Acad. Nat. Sci. Phila., vol. 2, p. 310.
437 Costa Rica M10cENE—OLSSON 265
Mactra Nuttallii Reeve, 1854, Conch., Icon., Mactra, fig. 125. Labtosa lineata Dall, 1898, Trans. Wagner Free Inst. Sci., vol. 3, pt. 4, p. goo.
This species is represented in our collection by fragments of the hinge and a perfect mold of the interior which is figured. This mold compares very closely with recent specimens of Laé- osa lineata Say.
Dall records this species as fossil in the Pliocene Caloosa- hatchie beds of Florida and we have several fragments in the Cornell collection from the Upper Chesapeake Miocene of Nat- ural Well, North Carolina.
Gatun Stage: Saury creek. Coll, 6, Estrella River.
Genus MULINIA, Gray
Mulinia lateralis Say, var. sauryensis, n. var. Plate 28, figures 23. 24
The species of Mulinta are extremlly variable and we have therefore associated this form with the common J/ateralts of Say, although it may later prove to bea distinct species.
The general form of the shell is like corbulotdes Deshayes, the common Gulf variety of lateralzs of Say. It differs most im- portantly from /aferalis in having the beaks situated a short dist- ance above the hinge-line as is seen in the Chesapeake Miocene M. congesta Conrad. In other characters, the Costa Rican shells are more closely related to the J/ateralis, than to the congesta. The shell is fairly convex and thin.
Length 20, height 16, semi-diameter 6 mm. Gatun Stage: Zone FE, Saury creek.
Genus MESODESMA, Deshayes Mesodesma costaricensis, n. sp. Plate 28, figure 18
Shell ovate, moderately convex, inequilateral, thin; anterior
266 | BULLETIN 39 438
end about % again aslongas the posterior, rounded; beaks high, triangular; posterior and anterior dorsal margins straight; ventral margin evenly rounded; surface nearly smooth; but with the dorsal submargins finely concentrically grooved; interior concealed.
Length 22, height 16, semi-diameter 4 mm. A rather thin, Donax-like shell from the Lower Gatun beds
of the Banana River. Its surface is smooth and polished with concentric grooves on its posterior and anterior dorsal area.
Gatun Stage: Hill No. 3, Banana River. Genus GASTROCHAENA, (Spengler) Cuvier
Gastrochzna rotunda Dall
Gastrochena ovata Sowerby, variety rotunda Dall, 1898, Trans. Wag- ner Free Inst. .Sci., voli 3) pt. 4) p.1325.
Gastrochena rotunda Dall, 1915, Bull. N. S. Nat. Mus., No. go, p. 157, PlseEO, tie 2.
A single, small specimen of a Gastrochena was collected in the Gatun of Red Cliff Creek. It is widely expanded about the posterior portion and probably represents a young shell of G. vo- tunda which Dall records from the Bowden beds of Jamaica. This species is alsofound in the Tampa Silex beds and in the Lower Miocene of Florida.
Gatun Stage: Coll. 4, Red Cliff Creek. Genus CORBULA, Lamarck
Section ALOIDIS, Megerle
? Corbula vieta Guppy Plate 28, figures 15, 16
Corbula wieta Guppy, Quart. Journ. Geol. Soc. London, vol. 22, p. 580, pl. 26. fig. 8. Corbula disparilis Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15, p.
247. Corbula (Aloidis) vieta Dall, 1898, Trans. Wagner Free Inst. Sci.,
vol. 3, pt. 4, p. 859.
439 Costa R1icA MIOCENE—OLSSON 267
Corbula (Aloidis) vieta Maury, 1917, Bull. Amer. Pal., vol. 5, p. 395, pi 20, fig. 13.
The members of this group of Corbule are difficult to sepa- rate satisfactorily as all are quite similiar in their characters. The right valve is broad, high and coarsely sculptured with regular concentric costz, while the left valve is much smaller, longer than high, with its surface smooth or faintly lined with un- equally distributed radial threads.
The C. vieta was described by Guppy from the Manzanilla beds of Trinidad and most authors have regarded the E7ycina tensa described at the same time as its left valve, an indentifica- tion which is open to question. Guppy’s figure of ¢ezsa, shows a Veneroid shell strongly sculptured with even radials and fine concentric lines, and quite unlike the left valves of Corbule of the Alozdzs section.
? Corbula vieta from Costa Rica is identical with Maury’s specimens from the Miocene of Santo Domingo. It is a smaller and more circular shell than the Cordula heterogenea Guppy which we have from the Canal Zone.
Length 6.25, height 6.50, semi-diameter 3 mm. (right valve)
Gatun Stage: Water Cay. fill ra, 3, Banana River.
Section CUNEOQCORBULA, Cossman Corbula oropendula, n. sp. Plate 28, figures 12, 13, 14, 26, 27
Shell small, solid, elongate, both valves of nearly equal con- vexity but the the right valve somewhat larger and overlapping on the left; beaks nearly central, the anterior end well rounded, the posterior somewhat contracted, slightly rostrated and with a well-marked keel; ventral margin more or less rounded, ter- minating at is posterior end ina small sinus, just anterior to
268 BULLETIN 39 440
the keel; the basal margins of both valves are folded in and flattened as seen in C. contracta Say but toa less extent and least in the right valve; surface sculpture of rather heavy con- centric costz, most widely spaced on the umbos; the surface may also show fine radial lines of small pustules or puncte which are best developed on the rostrum; no lunular pit.
Length 6.5, height 4.25, diameter 4 mm (Banana River). 9 6 semidiameter 2 mm (Saury).
We have chosen as the type of this spceies, the smaller and more abundant form as found in the Gatun bed of the Banana River, rather than the larger shells from the more inaccessible Saury. It is related to the C. contracta Say, but differs in hav- ing the ventral margin more curved, and the posterior extremi- ty more emarginate. Its surface is sculptured with rather strong, concentric costz which are more widely spaced on the umbos, and submicroscopic radial lines of fine puncte or small pustules, best seen on the rostral areas.
Gatun Stage: Hills ra, 3, Banana river (abundant). Zone G, Saury creek. Coll. 6, Red Clef :creek.
Corbula Dietziana C. B. Adams Plate 21, figure 8 Corbula Dietziana C. B. Adams, Contr. Conch., vol. 12, p. 235.
Corbula (Cuneocorbula) Dietziana Dall, 1898, Trans. Wagner Free Inst, Sei,> vol; 2, pt:.4, p. 856. A recent species quite common in the coralline limestones of the Limon Peninsula, from which it has previously been re- ported by Dall from specimens collected by Hill.
In its earlier stages of growth, this species resembles the C. equivalvts Phil. .(C. cubaniana d’Orb) in its rectangular from and strong dorsal keel. Later in life, the shells become strongly distorted, the ventral margin is abruptly bent downwards nearly at right angles to the former plane of the shell as is seen toa much less degree infthe recent C. contracta Say. Its surface is
441 Costa Rica M1ocENE—OLSSON 269
coarsely concentrically grooved, the two valves being sculptured alike.
Gatun Stage: Port Limon.
Corbula hexacyma Brown and Pilsbry Plate 21, figures 12, 13
Corbula hexacyma Brown and Pilsbry, 1912, Proc. Acad. Nat. Sci. Phila., vol. 64, p. 518, pl. 26, fig. 4.
A species of the Gatun beds of the Central Zone. The shell is rather large, solid and with its valves similiary sculptured with a few, deep, wave-like, concentric ribs. Its right valve is slightly more convex than its left. Our specimens measure:
Length 20, height 14, diameter of the left valve 6.00 mm. Length 20, height 13.50, diameter of the rigth valve 5-75 mm. Gatun Stage: Gatun, C. Z.
Corbula orosi, n. sp. Plate 21, figures 9, 10
Shell elongate, convex with the low inconspicuous beaks about the middle of the shell, so that the posterior and anterior ends of the shell are about equal in length; the right valve is slightly larger and its basal margin folds over slightly that of the left valve; anterior end rounded, the posterior narrow and more pointed but with the keel only very poorly developed; the surface is finely and evenly concentrically ribbed and both valves are similiarly sculptured; no lunule.
Length 20, height 11.5, diameter of both valves 9.5 mm.
A rather large species with nearly equally convex valves; surface sculptured with fine, regular, concentric ribs. It resembles the C. sarda Dall, from the Chipola Miocene but is much larger.
Gatun Stage, Hill No. 3, Banana River.
270 BULLETIN 39 442
Section BOTHROCORBULA, Gabb
Corbula viminea Guppy Plate 28, figure 25
Corbula viminea Guppy, 1866, Quart. Journ. Geol. Soc. London, vol. 22, pii2Q3 spl. 23, io 0:
Lothrocorbula viminea Gabb, 1873, Trans. Amer. Phil. Soc., vol. 15 Pp. 247.
Corbula ( Bothrocorbula) viminea Dall, 1898, Trans. Wagner Free Inst. Sci, Vol. 2;\ pt. 4, p:°S50:
Corbula (Bothrocorbula) viminea Maury, 1917, Bull. Amer. Pal., vol. 5, (P:1397, DE. 39, f1SS. (20,121.
This is a very characteristic species, distinguished by its large, deep lunule and its surface strongly sculptured with coarse, concentric, wave-like coste. It is known from the Mio- cene of Jamaica and Santo Domingo.
The Corbula hexacyma Brown and Pilsbry, still known on- ly from the Gatun of the Canal Zone is similiar in form and sculpture but lacks the deep lunular pit.
Length 15, height 10, semi-diameter 4.85 mm (right
valve). Gatun Stage: Middle creek.
443
Fig.
CosTaA RicA MIo0CENE--OLSSON 27 EXPLANATION OF PLATES PLATE « (4) Page I. Terebra bipartita Sowerby, Saury Creek, height 77 mm........ 35 2. Terebra dipartita Sowerby, Middle Creek, height 53 mm. ao) ecebraict, haitensis Dall, Sousi Creek, height'37 mim......... 35 4, Verebra gatunensis Toula var., Gatun, height 58 mm............ 36 5. Terebra gatunensis Toula, Gat un, height 52 mm. 6. Terebra gatunensis Toula, Gatun, height 42 mm. 7. Terebra spirifera Dall, var. midiensis, n. var., Middle Creek 39 8. Terebra gausapata Brown and Pilsbry, Mt. Hope, height NAAM ee sete oc ulciet Uns eUeM Mekidua maleate tiara 38 g. Terebra estrellana, n. sp. Estrella River, height 7.5 mm...... 40 Io. Terebra gausapata Brown and Pilsbry, Mt. Hope, height DOW AME HULU AN de easiest site Ue cccbi SMeeiWieme ces es 38 nie) berebra, Wolfganei Toula,) Mt: Hope, height 38 1m.....2-.2..... 37 12. Terebra Wolfgangi Toula, Mt. Hope, height 26 mm. ign wlerepra spiniteray Dall, Banana RIVET. scercdcseierses see voulsasis bsetvete 27 Peyve onus Sracilissimus ~uppy, Bocas del Toro... 2.2) .... chess. sess 52 Heer Conus tortuosostriatus Loula, Bocas :delk Toro...) .0sccesnehn eek. 49 io Letebraectrellanayin.i spe, Pstrella (River .2020 its hs a 40 iy es, berebra’ Berlinera= Maury, Boucary Creek.» 1.2.2.3. 06s. cs..0846 39 19. . Terebra costaricensis, n. sp., Banana River, height 13 mm... 4o 20, (Couns marginatus Sowerby, Red: Chil Creek........... ieee tenes 51 aie onus multistriatus Bose, Water Cay. icici isbacnie cogusismciege wera tess 54 22 COMMS INUSESIS, t. Sp.) Banana! RAVER. sil) ledecs see se eed eden sea 47 Ze oonus multistriatus- Bose, Water Caync..i wl Wooo. 54 2 POMS MUUSTeNSIS. 11) Spi,Baianal RIVER ia ccateewnauceee ase oe 47
25- 26. Terebra benthalis Dall, var. bocasensis, n. var., Bocas
eyo betepra, puinbriensis; 1. sp., Pumbri, Creek...) ai ee ie AI
272
Fig.
Fig.
Fig.
BULLETIN 39 444 PEATE 2% (5) Page I-2. Conus molis Brown and’ Pilsbry, Banana, River... eee 42 3-4.'Conus proteus Hiwass,; Water Cay. ..).5.sc-2ce ce eee 43 5. - Conus Veatchi,) ‘in. spi, Water, Caye- uae Ja alin ble eve ese 44 6. Conus imitator Brown and Pilsbry, Mt, Hope... eee 45 7. Conus’ toroensis, n. sp., Toro.Cays......0.0..ssseee aoa 48 S$. | Conus, Veatehi,,micsps, Water(@ay..-11. seen alia oe 44 g. Conus recognitus Guppy, Banana River ..... cs paid soldat Hea Ree eee 46 PEADE 3.16) I. Conus Harrisi, n. sp., Banana River:..... 2s). 6 eee eee 53 2. Conus stenostomus Sowerby, Port Limon).:.0), 2). eee 42 3. Conus floridanus Gabb, var. costaricensis, n. var Banana R... 45 4-5. Conus Burckhardi Bose, Water Cay... ....c)cscemeneae ee eee 52 6. Conus tortuosopunctatus Toula, Mt. Hope:i:s:32 See 54 7-8. Conus limonensis,;n. sp:, Port Limon!):) 203 eer 47 g. Conus floridanus Gabb, var. costaricensis, n. var., Banana R. 45 to. Conus planiliratus Sowerby, Port Limon....20 cee uaeanene 50 II, Conus tortuosopunctatus Toula, Mt. Hope.s yc. cece 54 12.. Conus cf. interstinctus Guppy, Banana Rivero: sare ee 48 13. Conus planiliratus Sowerby, Port Limion..42.0 eee 50 14. Conus planiliratus, var. bocasensis, n. var., Bocas del Toro.. 57 15. Conus granozonatus Guppy, Rio Blanco... ....22).i ne ceeeseee cence 50 16. Conus planiliratus var., bocasensis, n. var., Bocas del Toro.. 51 PLADE 4,107) 1-2, Turrisalbida Perry, slightly reduced, Toro sCays io 58 3. Scobinella Morierei (Laville) Cossmann, length 48 mm, Toro CAYS. oc aheinciin sb oleboienibine eee eee ameter 79 4. Scobinella Morierei (Laville) Cossmann, length 52 mm., Toro Caysiicsis ss dipheesespennen poeeeeeee 79 5... Drillia papaya, n. sp., length 32 mm., Rio'@oeles.)..)2.) aoe 6. Turricula lavinoides, n. sp., length 54 mm., Banana River.... 55
Halia americana, n.sp., length 66 mm., Mt, Hope:.:.-...20..-.. 79
445
Fig.
Fig.
8.
OO St One NS
ey OS Lana!
Pa Pde
23.
Costa R1icA MIocENE—OLSSON agi
Page Drillia consors Sowerby, Grape Point Creek, length 60 mm... 61 Drillia venusta Sowerby, Banana River, length 30 mm........ 61 Drillia, consors Sowerby, Mt.. Hope, length 33 mm....0.... ..4.. 61
Drillia carruca, n.sp., East Grape Point Creek, length 18mm. 7I Turricula lavinoides, var. limonensis, n. var., length 97 mm.,
Loro Caysyisc.. Ma OM Maes Ratna ee 56 Drillia consors Sowerby, length 4o mm, Banana River,........ 61 Bullaria paupercula Sowerby, Middle Creek, length 22 mim... 34c Drilliatlimonica, n. sp.,\Port Lamon, length 19 mmi.........0./:. 66 Ancistrosyrinx Dalli, n. sp., Toro Cays, length 14 mm......... 60
Ancistrosyrinx elegans Dall, var., Banana R.,length 15mm _ 60
PLATE 5. (8)
Drillia Cristobali, n. sp., Bocas del Toro, heigth 20 mm........ 72 Drillia chirquiensis, n. sp., Bocas del Toro, height 25 mm.. 64 Murricula taurina, n. sp., Bocas del Tore, height 25 mm........ 57 Dursieula Terryi, n.sp., Bocas del Toro, height 23.5 mm..... 57 Prillia bocasensis, n.sp., Bocas del Toro, height 22 mm...... 73 Drillia bocatoroensis, n.sp., Bocas del Toro, height 23 mm.. 67 isa, COCccos: 1.) Sp.,, Coco Plum, height 21, mmM....2.....0..0.5: 64 Prillia limonensis, n. sp., Port Limon, height 20 mm.......:. 7p Drillia limonensis, n. sp., height 15 mm.
Drilha limonetta, n. sp., Port Limon, height 12.25 mm......... 70 Drillia theobroma, n. sp., Bocas del Tcro, height 21 mm...... 68 Dilla citria, mu: sp, Port Limon, height ri mlmi.s.2.22...0.25)24. 65 Drillia citria, n. sp., Port Limon, height 11 mm.
Wrbiia\cocosina, n.. Sp.) Bocas del Toro, height 12 mmi....... 75 Drilia aucantia, no sp.,/Port Limon) height 17 mins..5. +... 68 Dillidalaquanicay ni\sp.,. Water Cay, height mitiv.....:06...5..44 65
Drillia aquanica var. Drillia lithocolletoides, n. sp., height 10.75 mm.,
BOGaS, Gel hOnO sero eekeosuinwekeencentuns 74 Leucosyrinx chloris, n. sp., height 14.50 mm., Bocas Wel Moro rela: esas ak ass eaaiactet 69 Gytharella dimata, mn.) Sp. height \o mm Port Limon... 2.04.5. gig Cythara terminula Dall, var. costaricensis n. var., height 13 mm) alla; Banana Raverin::.. 77 Cythara terminula Dall, var. costaricensis n. var., height 13 mins) Hab ta Banana River ola oy) Borsonia cocoensis, n. sp., Coco Plum, height 16 mm............ 78
374
Fig.
Fig.
Fig.
BULLETIN 39 446
Page 24. Borsonia cocoensis, n.sp., Coco Plum, (14) mamibee ee 78 25. Dfrillia aqueensis, n. sp., Water Cay, height e5ammin ee 73 26. Drillia musa, n. sp., Banana River, heteitjx6 mimi eee 74 27-28. Drillia musacina, n.sp., Banana River, height 9 mm ..... 69
29-30. Glyphostoma moinica, n. sp., Port Limon, height, 5.75 TMM, J. ch vce cecees eee Sean eS eee nee 76
PLATE 6 (9)
I. Mitra Swainsoni Broderip, var. limonensis, n.sp., Port Limon, height 74 mati.) ca weer 100 2-3. Cancellaria Plummeri,.n. sp., Toro Cays, heightiga aime iies 4. Cancellaria toroensis, n. sp., Toro Cays, height 29 mim) eee: 84
5. Cancellaria Mauryze, n. sp., Cercado de Mao, Santo Domingo, height 36 mimo... ..00) j..dsasses ean Sena 82
. Cancellaria Barretti Guppy, slightly enlarged, Banana River. 81 7. Cancellaria Rowelli Dall, East Grape Point Cr., height 30mm. 84
8. Cancellaria dariena Toula, about natural size, Toro Cays........ 80 g. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R. 81 Io. Mitra longa Gabb, Gatun, height 4o mim: .20 liao eee IOI 11. Cancellaria Cossmanni, n. sp., somewhat enlarged, Banana R 81 12. Cancellaria isiacolonis Maury, enlarged, Red Cliff Creek........ 86 13. Mitra cf. rudis Gabb, Hone Walk Creek, height 33 'mm.......... 102 14-15. Marginella MacDonaldi Dall, Banana R., height 28. mm... 95 16. Marginella mindiensis Cossmann, Gatun, height 9 mm.......... 96 17. Marginella MacDonaldi Dall, Banana R., height 26 mm......... 95 18. Mitra Almagrensis Toula, variety coralliophila, n. var., Port Limon,’ ‘height 07 maar 103. 19. Marginella MacDonaldi Dall, Banana River, height 25 mm.... 95 20-21. Mitra poas, n. sp., about natural size, Port Limon.............06. 102 22. Marginella leander Brown and Pilsbry, enlarged, Gatun....... 2) OS 23. Marginella mindiensis Crossmann, Gatun, height 17.5 mm...... 96 24. Mitra Almagrensis Toula, variety coralliophila, n. var., Port Limon, height about 17 mm........ 103 25. Mitra dariensis Brown and Pilsbry, Bocas del Toro, height QU) MIT... bases ne:cwelgceas eee eeeteR eee meena IoI
1, Oliva cylindrica Sowerby,’ Old Man Sam Creeks, 3007 c2ecia 88 2-3. Oliva brevispira Gabb,|) Ried \Cliti Greek. fo. cs te amseeae 88
447
Fig.
Fig.
Costa Rica MIocENE—OLSSON 275
Page
4. Oliva brevispira Gabb, var. Giraudi Cossmann, Middle Creek. 88
PmoOliracatunensis, Doula, Gata rs. 1630 lacscocaee jaspaeecsaiebiwacceseeses 89 6-7. Oliva Sayana Ravenel, var. immortua Pilsbry and Brown
Banana weaver aw toscens seuss! nck oes 89
8-9. Oliva mancinella, n. sp., East Grape Point Creek.............. Ol
LomaMeiilatia aqucensis,(n. sp., Water Cay iti ueeste ieee. 94
Ir Olivella muticoides Gabb, East Grape Point Creek................ 92 12-13. Oliva testacea Lamarck, var. costaricensis, n. var., Banana
BRAVO ave -chactetetaaisatd iwuateaals ietiure sae chee st go
i) Olivella muticoides Gabb, Red Cliff Creele. on... eet ies eke ceees 92
15-16. Olivella Boussaci Cossmann, var., Banana River............... 93
17-18. Olivella muticoides Gabb. Red Cliff Creek. ......0.......0..00.... 92
te-20. Olivella limonensis, n.sp;,\ Port Timon)! iil. soo. e etek ee (62
ay. Marginella avena Valenctennis, Banana River .....i....2......+.- 97
Paw Olivella Goliath, n- sp. Red) Clit Créeks. i.ccsccs sisi sccss.ctesesecs 92
23. Olivella Goliath, n. sp., Banana River 24-25. Olivella limonensis, var., bocasensis n. var., Bocas del
| ER OLOU A meee erenr ca eenuleempotsves Se cuk iden zs 94 fo-27.) Marginella collina,’m. sp.,,|Bahana River, v.../.....cesseces coseeese 97 26, Marginella avena Valenciennes, Port Limon..........00.....00:0+-- 97
PHATE 8.5) (18)
1. Fasciolaria MacDonaldi, n. sp., Banana River, height 70 mm. 106 20) Soluta Alfaro Dall, Banana River, height 63,mm............1.. 99 3. Solenosteira chiriquiensis, n, sp., Water Cay, reduced........... TI4 A weaAticus taurus, n:isp.,, loro Cay, slightly reduced................. 108 5. Fusus miocosmius, n. sp., Bocas del Toro, height 100 mm..... 103 6. Fusus honensis, n. sp., Hone Walk Creek, height 63 mm ..... 104
7. Solenosteira Vaughani Dall, variety medioamericana, n. var. Moro Gays iam eer. eas eect 113
8. Solensoteira Dalli Brown and Pilsbry, Gatun, slightly en- Lae CUE een ones Meisseuteesscee 112
g. Fasciolaria Gorgasiana Brown and Pilsbry, Rio Betey, height ROMANUM ae eM aias cauntes aa aakatel ee atecu ees 105
Io. Latirus infundibulum Gmelin, Pumbri Creek, slightly re- CECE Ue oes NN, «viedo uve 107 Ir. Peristernia insula, n. sp., Water Cay, height 23 mm............ IIO
12. Latirus irazu, n. sp., Port Limon, height slightly reduced..... 107 13. Peristernia tortugera, n. sp., Port Limon, height 29 mm....... 110
276
Fig.
Fig.
BULLETIN 39 | 448
Page
Melongena consors Sowerby, Old Man Sam Creek, height
ND ReMi ooh 0a MEME erie ed Os 112 Phos Veatchi, n. sp., Hill 1a, Banana River, height 43 mm.. 121 Phos Veatchi, n. sp., Hill ra, Banana River, height 41 mm.. 121
Phos gatuuensis Toula,Gatun, height 30 mim... 2 ee Dry Phos..beteyensis, n. sp., Rio Betey, height 26 mm) 324249 122 Northia northize Gray, variety miocenica, n.var., tip, Banana River... oc c0k enue eee eee 124 . Phos Moorei Guppy, variety costaricensis, n. var., Banana River, height 26: times eee 118 Phos mexicanus Bose, Water Cay, height 29 mim..2.. .sse-s05 117
Phos mexicanus Bose, Water Cay, height 27.5 mm.
Phos elegans Guppy, variety limonensis, n. var., Port Limon length 27 MiWhalses enemy see teenaeee 119
Phos elegans Guppy, variety limonensis, n. var., Port Limon length 22.50 mm
Northia northie Gray, variety miocenica, n. var’, Banana
River. Gees seman PPO EN et 124 Phos snbsemicostata Brown and Pilsbry, Saury Creek, height :
322M cis flesh cues ee eee ee 121 Phos metuloides Dall, Mt. Hope, height 16.5) mimasy7 esses 122 Phos estrellensis, n. sp., Estrella River, height 19 mm......... 120
Phos estrellensis, n. sp., Estrella River, height 16 mm. Alectrion merenda, n. sp., Old Man Sam Creek, height 7 mm 124
Alectrion ranuncula, n. sp., Banana River, height 14 mm..... 125 Alectrion losquemadica Maury, Port Limon, height 11.50mm 123 Alectrion ranuncula, n. sp., Banana River, height 14 mm...... ee
PLATE, £0°+(135)
Marginella latissima, variety Pilsbryi, n. var., Banana R...... 96 Marginella latissima, variety Pilisbryi, n. var., Gatun . Clavarcostaricana,, m: sp.; Banana River, height 40 mum........ 144 Metula Harrisi, variety limonensis, n. var.. Port Limon, heigh. 27 Tul 02) eee eaten 116 Metula Harrisi, variety limonensis, n. var., Port Limon, height. 25 jmmmil isaac BP ANG bets PEO Strombina matima, n. sp., Banana River, height 12 mm........ 127
449
Fig.
Fig.
Costa Rica MIocENE—OLSSON DTT Page Strombina costaricensis, n. sp., Middle Creek, height 15 mm 128 9g. Strombina ambigua Guppy, Banana River, height 26 mm...... 126 TO: tetaila Warrisi, n.sp.,:\Coco Plum, ‘height 27 mim.-............2.. 116 11. Metula Gabbi Brown and Pilsbry, Mt. Hope, height 28.5 mm 115 12. Metula cancellata Gabb, Bocas del ‘Toro, height 18.5 mm...... 115 13. Strombina Lessepiana Brown and Pilsbry, Gatun, height PAs 0045 Metaes amen AS RENN SEL ir nay Canteen 126 14. Strombina chiriquiensis, n. sp.. ....... SUG Wt ta cla tena @ Meera ca 130 I5. Typhis alatus Sowerby, Toro Cays, height 23 mm............ deren Gee 1O-17s) strombina, ‘costaricensis, n. isp), Middle Creek..........5....... 128 18. Strombina cricamola, n. sp , Water Cay, height 16 mm......... 12 19. Strombina sincola, n. sp., Banana River, height 7 mm.......... 129 20, Strombina Lessépiana Brown and Pilsbry, Gatun....:............ 126 12, Strombina sincola, n. sp., Banana River, height 7 mm......... 129 22. Typhis linguliferus, variety costaricensis, n. var., Bannan R. 132 20) Sisttuim nodutosum C, B.°Adams, Red Cliff Creek..........2.... hee Vie SurOMloia, ChITIGUICHSIS, I's (SPisser.cauesislcers'. sechewwoesvasees celles ees 130 Pp MlapetMCha MmuUSaACINAa;, n, Sp., Batlana RAVE: ...0c: .sjcece e2t seeks 98
26-27. Strombina costaricensis, variety musanica n. var. Banana R 129 26.) Cerithium costaricensis, n. sp., Red Cliff Creek, height
PSMA. 1 eA ese dot seaeeaes sel 143 29. Typhis linguliferus, variety costaricensis, n. var., Banana R. 132 gon, Clava alajuela, m sp., Middle Creek, height 17. mmi........2.... 145 31-32. Drillia estrellana, n. sp., Estrella River, height 7.50 mm... 69 33-34. Columbella submercatoria, n. sp., Red Cliff Creek........ eet! 125
PLATE 11 (14)
I, Xancus scopulus, n. sp., Banana River, height 265 mm......... 11k
PLATE 12 (1{5)
se serpulorbispapulosa. Guppy, Banana: River: ).c.c.....c2+-cseeoee- 145
2. Malea elliptica Pilsbry and Johnson, Banana River, height Oa Tt TID Oe Mie ee einete koe seers eam 139
3. Sconsia levigata Sowerby, variety Gabbi, n. var., Toro Cays, Shiphithy seinlarseds, witist kazaa oes 136
4. Siliquaria modesta Dall, variety limonensis, n. var., Port Limon, somewhat enlarged......... 147
5. Siliquaria modesta Dall, variety limonensis, n. var., Port Lamon, slightly enlarved:. 212.1... 147
Fig.
BULLETIN 39 450 Page 6. Siliquaria modesta Dall, variety limonensis, n. var., Port
Limon, slightly enlarged............ 147
7. Sconsia cocleana, n. sp., Rio Coles, height nmi 2.23. ae 138
8. -Cyprea cinerea Gmelin, Rio Blanco... 1 oe ee T40
g. Cyprea almirantensis, n. sp., Water Cay, height 39 mm...... 140
10. Cypreea parisimina, n. sp., Port Limon, height 39 mm......... 139
11, Phalium moniliferum Guppy, Rio Betey, height 35 mm........ 135
12. Sconsia bocasensis, n. sp., Bocas del Toro, height 35 mm...... Laz, 13. Sconsia bocasensis, n. sp., Bocas del Toro.
PLATE 13. (16) I-2-3. Strombus pugiloides Guppy, Banana River.................-2-2. 142 4. Kasciolaria tulipa Linnzeus, variety Port Limon.?)..2) cies 105
5-6. Strombus gatunensis Toula, Water Cay, somewhat reduced.. 141
~
/-
Polinices Stanislas-Meunieri Maury, Banana River, slightly
reduced .....5.:.0ceceesss mene aeons 157 8. Natica Youngi Maury, varietv cocleana, n. var., Cocles Creek, slightly enlarged. 22). sr 155 9. Natica canrena Linnzeus, Gatuil......-...:2cc2.5sceneee ee eee 155 10-11-12. Architectonica granulata Lamarck, Banana River......... 154 13-14-15. Natica Guppyana Toula, Banatia River.2...0) 1. Ss-cemenenee 156 16-17. Polinices subclausa Sowerby, Banana Riwersi.scece-. sss 157 PLATE 14, ° (17) 1. Turritella oreodoxa, n. sp., Banana River, height 49 mm.. ..... 152 2. Turritella exoleta Linnzeus, variety limonensis, Port Limon, height 43 mimi... 0. 7/7 ees 152 3. Turritella exoleta Linnzeus, variety limonensis, n. var., Port Limon, height 36 mm 4. Turritella altilira Conrad, variety chiriquiensis, mn. var., Banana River, height 52 mm...... 150 5. Turritella mimetes Brown and Pilsbry, Gatun, height 72 mm 149 6. Turritella altilira Conrad, Mt. Hope, typical, height 50mm 150 7. Turritella altilira Conrad, Gatun, typical, height 36 mm...... 149 8. Turritella altilira Conrad, variety chiriquiensis, n. var., Wat- er Cay, height 116 mimi .5.2e eee 150 g. Turritella altilira Conrad, variety chiriquiensis, n. var. Ban-
ana River) height:4o mimi.) eeu 150
45I
Costa RicA MIOCENE—OLSSON 279 Page
Fig. ro. Petaloconchus sculpturatus H. C. Lea, Water Cay, height RAS MANNS ietiaais get ost Uae eae eeine Se atetiaci ts 146
11. Turritella altilira Conrad, variety costaricensis, n. var., Sousi Creek MMeroMe QA iat Cece wees class I5I 12. Turritella gatunensis Conrad, Water Cay, height 36.5 mm..... 148
Fig.
13. Turritella gatunensis Conrad, Water Cay, height 24 mm.
14. Turritella altilira, variety chiriquiensis, n. var., Red Cliff Creek hers htsavaitiien kite eee sess
I5. Petaloconchus sculpturatus H. C. Lea, East Grape Point Creek’; height): 34) mitt i. ols.s eats os
PLATE 15. (18)
I. Dentalium uscarianum, n. sp,, Rio Cocles, natural size......... 2-3. Dentalium bocasensis, n. sp., Bocas del Toro, somewhat CMIALVCE Liss. Soe cree ci eatesswennss vsedetes 4. Astralium brevispinum Lamarck, variety basalis, n. var., old Man Sam Creek, natural size...... 5. Astralium brevispinum Lamarck, variety basalis, n. var., Old Man Sam Creek, natural size......
6-7. Calliostoma castilla, n. sp., Fort Limon, slightly enlarged.....
8. Calliostoma limonensis, n.sp., Port Limon, about natural
SS VA CAE YEON REEL Ey gine Eade Pipe Pilea CE g-10. Calliostoma mancinella, n. sp., Red Cliff Creek, about nat- Mal SIZE) eas eeiaue kA bape eoteseeeaes eee It. Calliostoma limonensis, n. sp., Port Limon, about natural SA SNR DEER ie T/A [TW AUL IAL OS Re RRA nay
ia unbo saltussn. spi, Red. Clift,Creek, height 17 mms <j. .1<.). 13. Scaphella costaricana, n. sp., Cocles Creek, height 29 mm..... 14 Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural
15. <Actzeon costaricensis, n. sp., Banana River, height 19.5 mm. 16. Chlorostoma costaricensis, n. sp., Red Cliff Creek, diameter
17. Crepitacella limonensis, n. sp., Port Limon, height 8 mm..... 18. Ptychosalpinx ? dentalis, n. sp., Cocles Creek, about natural OYA SAN Nr eae Agi RR gre ae a Ig. Bursa crassa Dillwyn, Banana River, about natural size........ 20-21. Acteocina bullata Kiener, variety costaricana, n. var., Mid- die) Creek, “height! ro mim. .o 2) 6h:
150
146
280 BULLETIN 39 452
Page 22. Neritina viridemaris Maury, Middle Creek, enlarged............ 159 23. Alaba turrita Guppy, Estrella River; height 5 mm............... 153
PLATE 16 (19)
Fig. 1. Pecten MacDonaldi, n. sp., type, Gatun (Toro limestone)
right valve, height 96)u1mi...7..07.. 198
2. Pecten MacDonaldi, n. sp., cotype, same locality, left valve height 79 mm.
3. Pecten gatunensis, Chagres Spillway, right valve, height G2 WAM. 0... a serven aeaerate eee 197
4. Pecten gatunensis, same locality, left valve, height 48 mm.
5. Pecten levicostatus Toula, Hill No. 2. Banana River, slight-
ly reduced 2... 0.0 cece eee 199 6. Pecten levicostatus Toula, same locality, left valve, slightly reduced 7. Pecten levicostatus Toula, same locality, right valve slightly reduced
PLATE 17 (20)
Fig. 1. Amusium luna Brown and Pilsbry, Chagres Spillway, some- what reduced ii)... geen 205 Pecten preglyptus, n. sp., Cocles Creek, natural size (type).. 202 Amusium bocasensis, n.sp., Bocas del Toro, natural size,
COLY PE... . posse sccneueet ana ee epee ee penana 206 4. Amusium bocasensis, n. sp., Bocas del Toro, interior of right
valve. 5. Pecten coralliphila, n. sp., type, Port Limon, slightly en-
LAr Ged... vsvculseccceeneceeepeeeReEeeMersees 199 6... Pecten coralliphila, n: sp., Port Limon, left valye.7 7s. eae. 199
7. Pecten preglyptus, n.sp., interior view of type, natural size 202 PLATE 18 (21)
Fig. 1. Ostrea megodon Hanley, Banana River, natural size ........... 195 2. Glycymeris canalis Brown and Pilsbry, Comadre Creek, about natural G1ZC 1.2, .5.04s0cussuaweeessnebene 177
433
Fig.
‘Fig.
Costa RicA Mr1ocENE—OLSSON 281 Page 3. Glyc ymeris canalis Brown and Pilsbry, Banana River, about MACITAL SIZES nV eee ieeu a eace un etek twee 177, 4. Glycymeris canalis Brown and Pilsbry, variety, East Grape ‘PointiGreek cs cali eis Ms 177 5. Glycymeris canalis Brown and Pilsbry, variety, East Grape Bot Greeky j05 Vales ca seca hee ane 177 6-7. Glycymeris canalis Brown and Pilsbry, Banana River......... 177 8. Pecten costaricensis,n. sp., Red Cliff Creek, right valve........ 201 g. Pecten costaricensis, n. sp., Red Cliff Creek, left valve. 10. Pecten costaricensis, n. sp., Red Cliff Creek, right valve. Hr Pecten costaricensis, n. sp., Red Cliff Creek, left valve. 12. Leda ensinoides, n. sp., Banana River, length 54 mm........... 174 13. Glycymeris castaneus Lamarck, Banana River,length 30 mm. 180 14. Glycymeris castaneus Lamarck, Banana River,length 23 mm. 15. Leda Balboz Brown and Pilsbry, Banana River, enlarged..... 173 16. Pecten almirantensis, n. sp., Bocas del Toro,heigth 4.75 mm, 203 I7.ebectem actus; mn: sp. Port Limon, height 4.50 mim....... ..<.<... 204 18. Pecten aotus, n. sp., Port Limon, 4.75 mm. 19. Leda quitanensis, n. sp., Quitana Creek, length 26 mm........ 174 20. Leda Davile, n. sp., Bocas del Toro, length 5.75 mm........... 172 21-22-23-24. Nucula cahuitensis, n. sp., Saury, length 3.50 mm..... 171 25. Nucula tenuisculpta Gabb, Red Cliff Creek,length 4.00 mm. 172 PLATE 19 (22) 1. Glycymeris carbasina Brown and Pilsbry, East Grape Point Creek, about natural size........... 179 2. Glycymeris carbasina Brown and Pilsbry, same locality, AVOUT MALUTAl SIZE ss ccc sae'sscicceeatcwse 179 3-4. Glycymeris jamaicensis, Red Cliff Creek, about natural size 178 5. Glycymetris carbasina Brown and Pilsbry, East Grape Point Creek, small shell about natural SUZ AD AN ABOUT BY ert LAO pL cen cL AEE 179 6-7. Glycymeris decussatus Lamarck, Old Man Sam Creek, nat- TITAS ZS A an eete eet ce ee ces enle te 2 LOO PLATE 20 (23) 1. Spondylus chiriquiensis, n. sp., Water Cay, height 151 mm... 206 2. Spondylus chiriquiensis, n. sp., young shell to show the small foliaceous attachment area. ey aracia) bristani,-n. sp, Rio’ Betey, length 42 mats... cass 211
282
Fig.
Io.
BULLETIN 39 454
Page
Crassatellites Reevei Gabb, Banana River, slightly red uced.. 212 Spondylus chiriquiensis, n. sp., Water Cay, young right
Spondylus chiriquiensis, n. sp., Water Cay, young left valve. PLATH 21° (24)
Spondylus gumanomocon Brown and Pilsbry, Hone Walk Creek, shell greatly reduced....... 207 Ostrea costaricensis, n. sp., Rio Blanco, about natural size... 196 Placunanomia lithobleta Dall, Banana River, height 36 mm.. 210 Placunanomia lithobleta Dall, Banana River, height 49 mm Placunanomia lithobleta Dall, interior of same specimen as hourey a. Anomia simplex d’Orbigny, Middle Creek, height 34 mm..... 209 Corbula gatunensis Toula, Gatun, length 21 mm. Corbula Dietziana C. B. Adams,Port Limon, length 15.5 mm. 268 Corbula orosi, n. sp., type, Banana River, length 19.5 mm... 269 Corbula orosi, n. sp., type, Banana River, length tg mm. Labiosa lineata Say, Saury Creek, length 44 mm.......... ieeeeeee 264 Corbula hexacyma Brown and Pilsbry, Gatun,length 26 mm. 269 Corbula hexacyma Brown and Pilsbry, Gatun,length 18 mm.
PLATE 22 (25)
Arca occidentalis Phil., Port Limon, somewhat enlarged...... 181 Arca umbonata Lamarck, East Grape Point Creek, slightly enlarged) ...4 02.6 1o sb eee enna 182
Arca auriculata Lamarck,,Old Man Sam Creek, young shell.. 190 Arca Maurye, n. sp., Port Limon, right valve oftype, length
Area candida Gmelin, Port Limon, left valve, length 18mm. 183 Arca candida Gmelin, Port Limon, right valve length
13 mm. Arca Mauryce, n. sp), interior view of type. .i2...2) ee eee 182 Arca honensis, n. sp., Hone Walk Creek, left valve type, slightly reduced is. pecans 187 Arca honensis, n.sp., same locality, right valve of cotype, slightly reduced()o-ssi5.8 4. eaters 187
Arca dariensis Brown and Pilsbry, Water Cay, about natural SIZOY Cae dL banal earl 0 ue 184
455
Fig.
Fig.
Fig.
ws
53.
I.
2. Le
4-5.
6.
7-8,
I 2; 3-4 6
Z
3.
I.
Costa R1icA M1IccENE—OLSSON 283
Page Arca dariensis Brown and Pilsbry, Water Cay, about natural
Arca dariensis Brown and Pilsbry, Gatun, about natural size. Arca dariensis Brown and Pilsbry, Gatun, about natural size.
PUATE 23.):'(26))
Arca Veatchi, n.sp., Water Cay, left valve of type, length My ATVI sas eee cues tlem aisle idle eran 189 Arca Veatchi, n. sp., Water Cay, interior of left valve of type. Arca Veatchi, n. sp., Water Cay, right valve of type. Arca golfoyaquensis Maury, variety medioamericana,n. var., Hone Walk Creek, valve of type, Neral a all {oie Wie 00) 00 eee mee Ceara 188 Arca golfoyaquensis Maury, variety medioamericana, n. var., Hone Walk Creek, left valve of type, same locality. Arca actinophora Dall, Red Cliff Creek, nearly natural size.. 185
PLA THUAN V2 7)
Arca cacica, n. sp., type, Comadre Creek, length 36 mm...... 190 eed Pittiert Dall, Saury Creek, natural size... i... (o.t.ec se sas eee 192
-5- atca Pittiert) Dall, Hill 1a; Banana,River, natural size,
Arca Pittieri Dall, Hill 1, Banana River, natural size.
Arca Hindsi, n. sp., type, East Grape Point Creek, natural SUZ) cis i A allt nels oi giaintew ses ee au 193
Arca Hindsi, n. sp., co-type, Rio Betey, natural size.
Arca Hindsi, n. sp., co-type, Hill 3, Banana River, natural Size.
Arca Lloydi, n. sp., co-type, Red Cliff Creek, length 18 mm.. 192
Arca Lloydi, n. sp., type, same locality, diameter 18 mm.
Arca Lloydi, n. sp., co-type, same locality, length 15.5 mm.
Anca Henekent Maury, Water Cay, length 30 :mm.....:...:...-. 186
Arca Henekeni Maury, Water Cay, length 28 mm.
PLATE 25 (28)
Arca costaricensis, n. sp., type, Hill 1a, Banana River, nat- (Bb a) (ESD PACE BORE AE SIM TILA eR cing AT 186
284
Fig.
iS)
i)
16 be
2
130 14.
BULLETIN 39 4.56
Page
Arca costaricensis, n. sp., co-type, Hill 1a, Banana River nat- ural size. Arca actinophora Dall, typical form, Mt. Hope, slightly re-
Arca MacDonaldi Dall, Hill No. 3, Banana River, nearly nat- ural size, interior view to show high, cardinal aréayi-.4.7-.ese- eee
Arca MacDonaldi Dall, exterior of the same specimen.
185
194
Arca MacDonaldi Dall, variety subreversa, n. var., Rio Betey
exterior of left valve.
Arca MacDonaldi Dall, variety subreversa n. var., Hill No. 3 Banana River, interior of right valve, to show form and low card- inal area.
Glycymeris Lloydsmithi Pilsbry and Brown, Red Cliff Creek, right valve, slightly reduced.......
Glycymeris Lloydsmithi Pilsbry and Brown, same locality left valve.
Glycymeris Lloydsmithi Pilsbry and Brown, same locality right valve.
PLATE 26 (29) Tellidora cristata Recluz, Hill 3, Banana River, nearly nat-
ral SIZ@i5 Ls cisecesy estes ne eee eee Tellidora cristata Recluz, Hill 1a, Banana River, nearly nat- ural size. Tellina dariena Conrad, Water Cay, natural Size... 12s. cen-ene
Tellina lepidota Dall, Water Cay, length 24 mimi..)..21...c5-. pe Tellina lepidota Dall, Water Cay, length 21 mm. Tellina costaricana, n. sp., Banana River, about natural size Tellina rovala, n. sp., Water Cay, slightly enlarged............. Tellina rovala, n. sp., Water Cay, (type) slightly enlarged. Tellina costaricana, n. sp., Banana River, about natural size Tellina crystallina Wood, Banana River, enlarged nearly LWICEL «a lenewe'ssansebeloateceineeenpaeben sane Macoma panamensis Dall, variety canalis n. var., Mt. Hope, length, 15, tat) .ooscc: caress see tumor ent: Tellina umbra Dall, East Grape Point Creek, enlarged about 3% TIMIES 33 HLL ee eee eee eee Macoma gatunensis Toula, Banana River, about natural size Macoma costaricana, n. sp., Rio Betey, about natural size.....
181
251 249 257 253
257 256
457 Costa Rica M1ocENE—OLSSON 285
Page
15. Tellina umbra Dall, East Grape Point Creek, enlarged about BAU ATIM Si Wie, oceabdladies tercewo eects 252) 16. Macoma costaricana, n. sp., Banana River, about natural size 256 £7. , J ellina nasua, n.-sp:, Red Cliff Creek, about natural size...... 250 18. Strigilla musanica, n. sp., Banana River, length 13 mm....... 255
19-20. Venericardia Conradiana Gabb, variety limonensis, n. var., Port Limon, lenothi7y mms... occ: 217
21. Chione grus Holmes, variety, Red Cliff Creek, enlarged about A ELIMESLINY Mid oucnoe sada mlnlas Msaeesae 248
22. Cardita caribbeanensis, n sp., Banana River, length 18 mm. 214 23-24. Limopsis monilis, n. sp,, Old Man Sam Creek, length 1.75 D300 06 ON Par cL a oN iy ENON nA BE REE A FI Ly
Fig. 1. Cardium spinosum Meuschen, var. Turtoni Dall, Port Limon
AWOUL Abr AL SIZE jee. 5.2, ccecnaceesne ts 228
2. Cardium dominicense Gabb, East Grape Point Creek, slight- Tym TGA Ced eatin uns Gaide< occas ace wceeneaes 225
-3. Cardium costaricauum, n.sp., Kast Grape Point Creek height PY AES 6 glis CANS ene Peale eR ae eee 226
4-5. Cardium stiriatum Brown and Pilsbry, Banana River, slight- Lp PEOMICEG YW Mette, 2 ele eats at kets siian 225 6. Cardium medium Linné, Port Limon, about natural size....... 226 7. Protocardia gurabica Maury, Gatun, length 29 mm............... 229
8. Protocardica gurabica Maury, Water Cay, length 25 mm. g. Protocardia gurabica Maury, Water Cay, diameter 19 mm.
fo.) Gardium heredium, n,’sp/; Rio’Betey, length 11.50 mm........ 227
11-12. Cardium serratum Linné, Middle Creek, natural size......... 228
13. Protocardia costaricensis, n. sp., Comadre Creek, length 21 ATER NS arene pce cdl, amy mre ide phi 229
PLATE 28 (31)
Hig. +5.’ Mactra estrellana, n. sp, Estrella River, length 50 mm......... 263 2. _Mactra exoleta Gray, Boucary Creek, about natural size....... 262 3. Mactra alata Spengler, Banana River, about natural size...... 263 A ivactra sincola, m..sp., saury Creek, length 36)mmi |. .2.4.00.0..: 264 5.) chinochama yaquensis Maury; Old) Harbor, .......2.22,262.00.4h00 219
286 ; BULLETIN 39 458
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6. Plicatula marginata Say, Water Cay, the lower or attached valve, enlarged about 2% time.... 208
7. Plicatula marginata Say, Water Cay, the lower or attached valve, enlarged about 2% times. 208
8. _Echinochama antiquata Dall, Rio Blanco.....ul. eee 218 g. Pteria inornata Gabb, Red Cliff Creek, natural size. ........... 195 Io. Limeea solida Dall, Red Cliff Creek, height 2.5 mm.............. 209 11. Chama congregata Conrad, Red Cliff Creek, enlarged about wissen + a teihe eine deieeb eee eee se 218 12. Corbula oropendula, n. sp., Banana River, length 6.5 mm..... 267
13. Corbnla oropendula, n.sp., Banana River, same valve but less magnified, length 6.5 mm 14. Corbula oropendula, n. sp., same locality, right valve, 6.5
mm. 15. Corbula cf. vieta Guppy, Banana River, right valve, length 6.25 IMM...) cane eee ee 266 16. Corbula cf. vieta Guppy, Banana River, left valve, length 6.25 mm. 17. Leda Dalliana, n. sp., Port Limon; length: re. 50mm 175 18. Mesodesma costaricensis, n. sp., Banana River, length 22 TUTE 05.0 ois ninitie ale oe tek ot sR ee 265 19. Nucula orbicella, n. sp., Port Limon, length 9.5 mm............ I7I
20. Nucula orbicella, n. sp., Port Limon, length 11 mm.
21-22. Yoldia bocasensis, n. sp., Bocas del Toro, length 9.5 mm... 176
23. Mulinia lateralis Say, variety sauryensis, n. var.,-Saury Creek, about naturalssizess eae 265
24. Mulinia lateralis Say, variety sauryensis, n. var., Saury Creek, about natural size.
25. Corbula viminea Guppy, Middle Creek, length 15 mm......... 270
26-27. Corbula oropendula, n. sp., Saury Creek, length 9 mm...... 267
PLATE 29 (32)
Fig. 4. Semele levis Sowerby, variety costaricensis, n. var., Hill 3,
Banana River, length 72 mm...... 258 2. Semele Claytoni Maury, Hotel Creek, natural-size.............. 259 3. Metis barba, n.sp., Rio Betey (type), right valve, length
AS. MOI hoi Se ee eee eee ee 255
4. Metis barba, n. sp., Rio Betey (type), left valve, length 48 mim.
Fig.
Fig.
To, 1 Os
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he ce eae
Costa R1icA M1I1ocENE—OLSSON 287
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Sanguinolaria alouatta, n.sp., Banana River, right valve, HEME LEN OO IMM: 1.70 ck Weoceeetciule wet lakes 260 Sanguinolaria alouatta, n. sp., Banana River, left valve, length 54 mm. Psammosolen gatunensis Toula, variety costaricensis, n. var.
Rio, Betey length: 63 amo)... 260 Semele Sayi Toula, Sousi Creek, slightly enlarged .............. 258 Tagelus cebus, n. sp., Rio Betey, length 36 mm.................. 261 Abra triangulata Dall, Bocas del Toro, length 7.25 mm........ 259 Crassatellites midiensis, n. sp., Middle Creek, length 3.25
POAT Sh SA ee SEA AR UN A san aaah eds Suva 213 Crassatellites bowdenensis Dall, variety costaricensis, n. var.
Keto hve AO cae. hss ce uee vet oes tees 213
PLATE 30. (33)
mAovigona multicostata Sowerby, Water Cay. ....4-s.......i66 seed. 239 Chione Rowleéi, n. sp., Banana River, length 42 mm.......... 244 Antigona Harrisiana, n. sp., Port Limon, length 21.5 mm... 241 . Antigona rugosa Gmelin, Port Limon, about % natural size 240 Antigona Blandiana Guppy, Comadre Creek, length 30 mm.. 241 Antigona Blandiana Guppy, Port Lomon, length 24 mm.
Chione mactropsis{Conrad, Gatun, length 30 mml................. 245 Chione mactropsis Conrad, Sousi Creek, length 51 mm.
Chione holocyma Brown and Pilsbry, Gatun, length 36.5
PLATE 31 (34)
Dosinia acetabulum Conrad, Rio Betey, somewhat reduced... 231
Cyclinella beteyesis. n.sp.,; Rio) Betey, eight=79 mim... /..... 242 Pitaria circinata Born, Saury Creek, about natural-size......... 236 Clementia dariena Conrad, Sousi Creek, height 60 mm ...... 232 Pitaria Hillii Dall, variety musanica, n. var., Banana River,
MCAT CIN Are eT 2 ic di cic ea Maisiertoie’ Se 237
. Macrocallista maculata Linnzeus, Middle Creek, about nat-
PUTA ISTZ Wea sy Ratdeiowek aeuceni ani Rend oe taney 234 Cyclinella subquadrata Hanley, variety quitana, n. var., Quitana Creek, height 28 mm..... 243
288 BULLETIN 39 460
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9. Pitariacireinata Born, Saury Creek, about natural size......... 236 10, Pitaria boucaryensis, n. sp., Boucary Creek, about natural
SIZ ie a Se 239
II, Pitaria Guppyana Gabb, Port Limon, about natural size. ..... 237
PLATE 32) \(35)
Fig. 1. Callocardia gatunensis Dall, Banana River, about natural
SIZC...eceoas ssonesapaceaieatiepeepRi im emenas 235 2-3. Pitaria VanWinklee, n.sp., Banana River, about natural SIZE... essen dna iel eee ee eee a 238 4. Chione costaricensis, n. sp., Banana River, length 34 mm..... 245 5. Phacoides costaricensis, n.sp., Middle Creek, length 11.50 TATE 5 oo ois one's bint oly dae nee ala ea ae 222 6. Phacoides bocasensis, n. sp., Bocas del Toro, length 10.00 MMM) is oo. cslbied aie ee See miata Sere Ree 223 7. Chione hotelensis, n. sp., Hotel Creek, slightly reduced...... 247 8. Cardita matima, n.sp., Old Man Sam Creek, about natural SIZE acc esisine Sdacenmasiebhaoeeu Aiea eee eee 215
g-10. Chione chiriquiensis, n. sp., Water Cay, length 13.75 mm.. 248 11. Myrteea limoniana Dall, Red Cliff Creek, height 9.50 mm..... 221 12-13. Venericardia Terryi, n. sp., Quitana Creek, natural size,... 216 14. Venericardia scabricostata Guppy, East Grape Point Creek,
about natural size.i: sae 216 15. Diplodonta collina, n. sp., Middle Creek, height 6.75 mm..... 220 16. Phacoides radians Conrad, variety medioamericanus, n. sp.,
Middle Creek, height 9 mm........ 221 17. Venericardia scabricostata Guppy, Comadre Creek, about nat- Ural SIZE. i. dncet ete ee peeeneemees 216
18. Gafrarium limonense, n. sp., Port Limon, length 4.75 mm 233 Ig. Gafrarium altum Dall, variety costaricense, n. var., Estrella
River, enlarged about 3 times...... 234 20. Divaricella quadrisulcata d’Orbigny, Middle Creek, height
TA DATA, Lic sce syne apelin team enema 224 21. Diplodonta insula, n. sp., Water Cay, height 6.5 mm............. 220 22. Gafrarium altum Dall, variety costaricense, n.sp., Middle
Creek, height) 4.75 miMi,s5s27-,0-50 234 23. Myrtza limoniana Dall, Port Limomi)t, 0005, atone enema sella 221 24. Phacoides actinus Dall, Bocas del Toro, height 4.50 mm....... 224
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INDEX TO VOLUME 9
‘ote.—Figures refer to the continuous numbering of the volume (not of the separate Bulletins); Roman numerals refer to pages; Ltalics, to plates.
Abra triangulata 32........ 200, 431! Acanthochites pygmeeus........... 170 SPICUlOSUS ....2.... 4... OR ees 169 Acanthopleura granulata......... 169 PACS eset ss Sonina tis sss Sanicun sie ss 169 Acmea candeana..............0004.. 151 MEMCOPIEUNT A oils sc ssica'ss < nines 152 MmelanOleuca. «. 0.00.06 )s05 5c es 152 PUMICOMAG a oe. li sks cases shh I51 Actzon costaricensis /78........ 206a GABDETISIS aot en os osc eh stint 45 (SSCS 15 CS a ee 46 IRONS Matera eee Shedecsaeelsoc ds eas 45 PAIVESLOMESIG ©... c0006.200ces0055- 47 hak CCT S oe a Ate a eign 46 AMON Sy tee ciated). e's ise enigu bs «di 46 WEWIOLAUNS: 00 0 ces seh wasn cone et 46 | EVAL Ve ob tog Re ate ee 45 PUIMCTOSERIALUS. 060.0... 5 ede eee 45 PeOMINE SE ise ok Sa cdiw S cia'vcloisaala dae 46 Acteocina bullata......... 47, 190, 199 bullata, var. costa- BUCA CT O vice es cise dusoie nc 206a GAMA WAC Ae | Ms cases oceea see’ 48 BAAS oso netiecate ete teks 48 POCUA o.ceceteees AJ; 190, TO4, FOT, 206b thitiewm-tritonia. .......0.....0. 194 US DIMMU ELS Be oro i icicdde de vetlgns eae 48 @deorbis, adainsi.........2.0:......65 159 SUNORAMIGIGUS |. 2... ses. 2cu sens se 160 Admete microscopica................. 75 IMESOIIS SUCATHSI., .00.c.006 -esey senses 94 PME GEEVATICOSA, | ..):.:ssesesacdesses 138
CUTE evr ok hla edeecapes Bie 199, 325 AdlalinaracdagmSii suse, .aicce aie seeces 127 CERMEHIOIGES 4) Wah an aa eae 128 ICECRIOMIACUUUS an. o tee ctnc cee ners 89 ADAM S312 0o nds Gola tulebeusasiseaeetons 89 AMIUDIGUNS Voesces ca seves ena teet nest 89 atetublan yee ous an seca 89 DEatiMOMLENSIS: ised eee gt Candidissima yy fe soo. awe cs 89 CONSENSUS ..cad.ccs cs easc Me necaataae go PRELELISIS Tyco) Me eeheeseeu de ohoces 89 PAIVIESLOMENSIC csc. cascesessoses go WOLESSIETIN Zi Shwcs costes tees. go losquemodicaaczs. 0. acess k 195 losquemodica, 072 ).)s...0.h.6. 295 MIETEMG ae waco ca ente, 296 ODSOISUAS Ja lS rere) PAUUNCOIAL Muy oie le aks 295 ETIGAIVESTUS cease ceo ccnes go NEDER: Pgs tie leant Meee nan ae ahaa 89 Ava Bluth SEES sles ckedaseeaces 202 J Nico. 4: Woes eae A ibsalsame tate cuneate nals 57 Amaura tombigbeensis.............. 26 Amphiperas canadiensis........... Ti2 Ampullaria caliginosa. ............ 14! Gepressaci ce ley sate Saeed. I4I PUNST rast cee ak ame au cual. I4I Amusium bocasense 20.......... 378 | bk ba Mbp endear TIC NE A 200 iS ssa ob by An ADS) Nunn eeet MURAI Gee eng 187 AHOTCONMT hse Lee kee aL 2003377 [C0] 30 Ue CRE Ra Seger Came AUS Ae REE 186 Aneehisratbelia’ tc cce senna 93 am phisella gs.) iin ae ae he 92
290 BULLETIN 39 462
avara, var. semiplicata......... 92 dariensis ) 25 .V. ice ee 187, 356 VAT: HSIMALIIS oH uooceus sche teenth eek gI gatunéasis eee eee 356 WaT) thanSiitatarn.)acedeeets tee 92 golfoyaquensis:...aa.a 193 cancellatay ig ce esses, a uuacuas g2 golfoyaquensis, var. medio- Catenatal ie vekecsncer accesses 92 americana .|\ 265 4 u eee eee 360 PUPA eee e eee 186 PTAndiSiy... cece kee IQI INOLESSCRIA NA ees scence ee kuiee 93 henekeni |, 27.551), 193, \naey1a55 ODESSA ec sine nM ncaa LG a eeaeN sanity 92 hinds! \27..:.2) oasis gees 365 OLMaGA a ie ene wase nee ates 92 honensis | 255:53-<.ciee eee .359 ostrercolat cl Mies. EA neeuse 92 imbricata.:))j.tcossemaaeeeee Oe | DECECI Rie veer ce ee anise 93 lloyd \\27:ieneee ape Sanaa 364 Ancillaria acquaensis /0......... 266 macdonaldi .28...187, 191, 366 chipolana 5 oye e ase sae 267 MAUS. 25... cosas PIN SWISH siren cue entceeno ee aeets 267 occidentalis 25...........189, 193 subglobosarc ie) a ak ee spre cect Z. 199, 353 Ancillopsis tuomeyi .....:.......- 12 oronlensis’. 2. ..3a ee 356 ALACISthODASIS AN vibeowacn are meen ets ae 158 ozarkensiS: |) 9.543.) eee 22 Atleystosynitx dalliy 70. sacs 232 patricia..ccieeeee ee Wee te Ig! elegans yu lennetresee 190, 232 pittiert | 27s 187, 364 TAMIA tal se) eco aaceuneuee eee ees OB TEVETSA, .....eneeel eee eee IgI Anemia ephippium)) 4.2.0)... 381 SODL1M AL, :2) 4s saree eee 358 Simm plext wave scaicve 189, 194, 200, umbonata 25.....189, 193, 354 DOS s 27 secretes sutneeacneast 381 veatchi' 26). 2...) uae ee su Aumntalich lc. oi tan oe aceneeah eem nana es aah 35 Architectonica granulata 79...138 Antigona’ blandiana 33), 2200) 3 431 190, I9I, 196, 200, 326 Derrisiama 200) Jie aaa pe 188 perspectival.. >. 138, 326 Narrisianay (23. Masters. AI3 Aspella, var. obeliscus,:) 2222 117 multicostata!) 77).c:-cse« 189, I91 var. paupercula...2.) eens g ATI scalarioides.... cers aes 117 PUPOSA I Quil ew ie ass canal 189, 412 Assiminea auberiana,............... 142 tarquinia |.73)..<.% 196, 194, Astralium brevispinum /78...... 190 412 334 Aphera islacolonis zone............ 193 Ces lati 2. ccte sees 190, 334 ADL Sia ee ee ne een ake Beets Astyris diaphana .7.e i ee 94 Area‘actinophoray) 2Oieser. secre Chey duclosiana..")). 2 ae 94 aubicw@lata 250.52 189, 193, 199 leinata..).35.cse0 sc eee eee 93 362 Atlanta peroniy:. cee eee III CACICA Zee eee wecken ce aca 362 Atys robustoides,.... 0) sci4) ee 26 Candida 25 Wey ee ssean te 189, 355 Auricula Coronas: ince 5- eee 56 CHITIQUICHSIS! 2.5. eseccenunat 193 OVA: 0.06 een 55 Consobrinas ic.s2..c.csceses eens 358 pellucens,.o0 2c Ue 54 COStariCenSis (925) 44 ees slices 358 Auriculina inseniptass ie 46
463 Costa Rica MrocENE—OLSSON 291
PPEMMONNPAGUHIIT. ....0)cce0c. sevesesscs ee 54 isc) Sebi | ols oe 82 jou) 10) Cr oI ROIS LIC 2 | A ea 8I Amictiia imoriata.........2........+ 367 B Balanophylla haleana................27 Eee UMEIL EIN Eos a Snemnedee ve bowen e A3, Peete talc Oidea. 2)... ia cestesees 4 Basmiesa Bltan.......6.2.1.,<05 157 BOSMUPALAN Eke. csesiedesspbecds- cu 158 SIS CYEs 02 NA Ba SP Dl yn ae 158 Mate elteattila i... 0). Lc es 158 Mae ETSLCSSAL: lies ors qcsees se 158 BasterOtia ? prima, 37.-...........--. 20 BatwOxypWus...... 6.2.6 cm. Bie el 38 EVSUAY SEU NS) eg 68 Pe MrNOMOlNAI. oc... ss. oc. oc LT? Benthonella fischeri................. 140 PRA RPE Oran Sse tinle 2 sit Zete noe e Selec 140 MUR OMIS AN hea eet sel bs Saale des a 140 i 2) 0) Scie 01670 0) (=>. <a 128 PS TMMMIOUAE! 6d neceneteseh< +5 128 Ma VESUOMENSE): 60.6 nese edent 129 GN DSIUTAY . oS. sg oe eens 128 CHL ESC CCTM ae en ee 128 PAPUA MORALE oi corse bes seuss 186 VGECNLL CME OER 128 WANE ALCCATIUTO 0... see co decie cle 135 Bin @iata (OMETLA 2. 2 des se doce ne hss 132 PRM MMMCTM Ae cbs aicclc st vie cosksenosts cn nacehs 56 PE WELOC MICAS) 026. acne eens PORT TVS IOI ICE |G Ie RA ip alee ae a a 57 BOECOMOO NOM. oo.) ii sce sc.s cadences an 99 Borsonia cocoensis §&........ ie OG Bothrocorbula viminea............ 442 Bowden beds, Jamaica............. 199 UGCA ODES... .....c....secec00e 92 PN VAL eck an's coe soa lace shee von ss 87 (EIST 12) LISS etree a 99
LACIE SUAS SAS eee er 94
Bubimus) monte eeecckece: 55 182 OU V2 Bea SAAR eo uPE TS UR 48, 49 jeyRay toric: VAR DIG pena a als eaters ccm 50 Canta licwmlatas rye Us a uasen! 0 47 Paupercigla eres ase sees 206d MCU ters sate Mec eecboseesticast 206a Bullaria abyssicola:s 2c sessile ses 51 amyedalas Ni Av NG ah ER 206d OcCidienmbalis.S simi u ose verse 50 pauperculias)'7/.1G4, 169,)2C6C PINSUISCHMAAe Air RL ee ad 51 S@ lida: Maer ens nan Son NIV Ee oa 51 Strial care ey ee SNA MALLE 50 BuiSAy aE IS yoo 0 coe ON sh 114 crassa J8.....190, 195, 200, 306 Busycon canaliculatum ............. 85 CARICAL Stee ca enh Caen cet Mute! 86 SI (SS SIT Ue Wane ak ni 86 PEEVEESUMI oe ee ee soe 86 PYM mceeces sce suralsatew ements O5 C WaduuliiSy ar ChUSe. Gc ocsdiesc eet as At AMAL AILS Peete cecum ee eae 40 CATOMMENSISE aie eh el OD: 4O CIONBSAUUS 5 hiss awe yee eeeoeeoets 40 AAV OM Seek pose hs Nadel he een 4I PAEVUS ese le. ic ens decseueckaes 4o POCHME TNR ele oils a Rican: 40 Quadnidemtatus: li scu ios wees 40 ELAUSILOTMUS Hie ool ee Al WUE SUMS Hes. Masten eons se eeae isc. 26 Cacti annulate uae 130 bipartituiae, 0s... Wisse tasateee 130 CATOMMAMET S01) ce eetee see 130 COStAPUM tat iees selec e mete. 131 real kei ey ab 8 oc DAMN Wee nme SOL OLR 131 PUSUOUCLUIA. LNA ust wseetev ys. 130 TPE PATE rs Ue) crue oleae 130 CUE RCV IA LU cy Nea Se 122 pudehe tum ce ys e ence 131
292 Costa RicA MI0CENE—OLSSON 464
floridanunma uo as 130 Calliostoma asperrimumi........... 156 Castilla 7G kee Nal eer R27. CIfCUunICinCctum Mackie 154 CORDIS i Bessy asym ane 155 CuedypUUs se es ne see men ee 154 Fup rae ek ae 155 IIMOMEHSIS' ».78....0% essences: 336 mManeinellal jas. cenesst some ies 336 PHICHERU MI! eas7oe eee ace sees 155 WAT. HAWSOM Mac i)ctoacle (oleae 156 TOSCO WEA eek ane yh, 155 Lamipasnsis st eee eee bee 165 LS ote a Aig on 155 VOLRCT He iat Ce Seen Uta te 155 WIECAECCA NUN oc .': teninntecnae 156 Callocardia gatunensis §3?5....... 407 Cahlogagay. ls. UiN os sedi erate 154 Caly ptreeacan@eawa ste. eet 145 PAEVUINS oie o ee ete ene {45 Calyptraphorus valatus, var. com- PIOSSUSh ue) dhe ee use meen 26 Cancellarta agassiziii:.i2.is.smacewe 74 Al yeaa (yceead, talk autem ene secret 7 Danrettii ost eae 200, 253 Poullatar ls ccauiete nuns nome es 191 CANCE aureus weahe nua aa anne tas 88 Clavatullanny ae) 2e scene tte 74 CONLAdIA NAL .c. cache oee eee 74 COSSHMAMTA (Oo sua k sete: 253 COStAEA Le ie Sadesretiseeema amen 7 artema/ (Os Cc ueay eee 252 epistoumiitenay 2.4) 4c ee. 195, 255 PaAlVEStOMEMSIS:. aa. ciseueees bese 74 Silber (7c cere encore ee 13 islacolonis 9. .....187, I91, 195 196, 258 AMAURY) (QU ok. jen ences QS. Ve54 moorei... .... uae abaleMeirelemcutinaaton 255 PAMONES) 204) Lesa ads asocewsoiesee kobe 7
PlumImert: Oy.5605.5).0.8 TOU. 257
reticulata eee eee 74 rowelll . ‘@. gene 195, 256 SIMpsoniy 6400 dee 75 teneray ease 1 he bale eevanemvelets 74 tessellatay.: /.Jc-Useeanceanee IgI, 258 toroensis' 9:2). egies IQI, 256 var, trachyostraca.).cuecucrs 252 tuberculata.) 33 eeeeeeee IQI Cantharus limbatts)..5.2c eee eee 88 Orbig_gny! ¢'...nsveenteneneenes eee 87 tinctus...)., {Ree eeeepeneee ae 87 Capulus intertus.7)e) eee 143 Cardata caribbeanensis 29...... 386 laticostataJ:.:c. 3 eee 387 matima | 25020.) sadeaeue eee 387 scar bricostatas./.5.ees eee 388 Cardium, var, altermatai.o) 189 apateticumn: ie. sauces 399 costaficanum 3ous une 398 dominicense | 30)... 194, 397 GRD MUD s MME R MAP oN 186 PalliMeuse. -o.ccesee eee 186, 397 heredium 30) 4:3)-.e ee 399 medium 30...189, 194, 205, 398 Serratuit: | 3O-e css. 194, 205, 400 SP1INOSUML, (5.2 caveeseeeee 189, 205 var, turtonl,/ Joo 400 striatum 20 nj ieee 188, 397 sublineatuma 5.7.3.0 201 Careliopsis,:. 230. ieee eee ene 110 Caricella dolita. 2.2230 eee ene 7 Caryatis guppyana... {tf oee 409 Cassidaria leevigata..i0 2s eee 308 Cassis cameo...,:.i.-chen eee eee 118 flanimea,....2. 22.2208 eee 307 inflata. 2... c Lee eeeeaie eee 118 monilifera...\. deena 307 reclusas. 3) yA 5) see eee 307 suleiféral. sees 195, 200, 306 testiculus.:..(. 5.004 eee 118 tuberdsay..). 2.20. oases pee 118
465 BULLETIN 39 293
P Devo toe ibe t 8 (c>-¢: ae 42 PHUSTRITIO SA lal, oe osisss se dine voetlcl ees 42 PUMA AMR AA on. keen esaseces ce -ei42 Wercade formation. .............5.5.. 193 Cerithidea costata .:..... ........... 126
POE MASIAT, .ocicscicecn (2200s ileeees, E27 ScalarirOrmis..........-...-....-. 826 ECS AGC) SV eR OS TeX 6
Reem@e a. ld) svn y ied 126 Cerithiopsis burkvillensis......... 123 Bh sra ee ee ee eee cea ss 123 EMME TSOM Tesh st sc Sota ckeee eke a cece 124 LSU S1 90 1) ee BREECH CL. oo cs cob di nsecse essa sells 123 MUMPE ESET GH see's clive. dh sues 123 STEUER a ee A Be e101 ha 1h o a eA ee 122 Centhinm algicola .....7.........4..125 BOStAIICEUISE LP. .2.0.scve-cceee 315 CMISIMCAUHINS osc scuii ck. Seenen ses 59 PUG AE ETD Scr cs ch ce esse meiede's o 125 PUA ATT. 2255s vecceseei ses 124 PPIVESLODEHSE ia. c.0 doco.eleccees 126 PBC EMON bots oi s5- sco kneucelcse ce 123 DRDO APUMEN 02 eo) Loewe less 125 rash yoy 0d0 eta: el i a. 125 Waly MISTCSCENS!:.... 75h. ¢.2605. 125 GANT S CAMA Ss. oce sts cet cess. t=: 125 PROS ALIN oe es Sale velsie.! cis oe 59 IDCMCEAUUTAS ieee vescec ete’ 124 MERE DEALIG(s (2.5724 seb dit sew s'ns 124 WG Hee ee ee eee eae fe's 125 Cheetopleura apiculata.............. 168 Chama antiquata,.:..0...) ..cc.00.45 390 TOME Ai eesiccns sees d Mein t dua 391 COMPFEL ATA) F/...cc0se0e: 203, 390 eCongrevatoides,........-.....55. 390 @ireilea eGwestris. ../6.....5.....008. 143 BME CtONIA. occ... esl. eoues 186 SPATE 05) soon scesdecctocesuses 108 Chesapeake Miocene...... Weih omie 202
Chione chiriquiensis ?5.......... 420
COSEATICENSIS |) 75 sci cesteesseuer 417 SLUSH ZO Mice. eee 189, 205, 420 MOIOC YW IMA A ge eta fa cveuaueces 419 horelensisv wa sea wu. awe: 419 IMACLEOPSIS ope coease: 188, 417 TOMLCEL VF pRt owes een. 188, 416 LEU ee ea tees Sp eoaeke 187 WIOC Via se... 200) 201, AL7 COHTCO Me tc Mere kde re otuceren cea duals vendre 168 MVAGINOTACU SR sccvesasseteu esos 169 tuberculatusi we eck 169 CHICOTEUS ua Leese tamedded seeks ests 95 Chlorostoma costaricensis 78...335 C horistella pompholyx............ 161 CIneMbisy ITAtMStielase te nc cnetosdee. 160 jh bb aan as EAA Rer eee 160 Clase Tella JEwettl. sss. 0: secsees ce. as 70 Olavaralajwelay! (7p: koe, casceugs sone 317 Cipolanaryeuesteaccstcneesesces 126 COStaimcanaly, 77.0) toioc08 seuss 316 Clementia dariena }¥4....... 187, 404 Cleodoraylanceolatal eo cicciche esse 42 Chie, recunvals cceoses vee ous easee 43 PY LAMIdAtaea.deekeccen Men 42 Cochliolepis parasitica.............. 159 SUAL As. .acesnisleveveweehs sina sent 159 Columibellavambigua,.2c.ee. 0.027 298 MET CAtOFias....s40<00 Mage an, Tees 93 GUSCICOIMES © coe ec ray accee 93 SUD IMELCATOFIS Wic7frrccs ve-cts6e- 297 Columbraria , testacea....3....5..2.. 115 Compresside@nsr ves. ctescccnss 1 toe ek 39 GCOMORNERA eta esea use ekoacee ieee 82 Conusienmiulatone testo sure cet 186 APASSIZE Ree eens ook nes 200, 226 AU PHU SUS we easel ime 62 berg haste ss fsessccesees 3 paeed a2 105 VAT DOSCEMSIS), | (Ol sscnedeetecss 222 burckhartt). (10) c.0..20c8. 2OO 224 catenatus..... BAR Cae cence uBESS 215 CEMtunlOn estes. ae kee 61
204 BULLETIN 39 466
Conus chipolanus.......2...2... 62) (207 tortuosostriatus’ 7 J vas 221 CONCAVILCCHUII, 0 32ae aenceece 215 weateht (5 Geer srckcee a eee eee 216 COMSODIINUS Coo eeeeae 186, 195 Coralliophila abbreviata............ IOI war. eostaricensis: | 6..22..224. 207, deburginige 3) 0). 2 eee Oe Gallas toa kien e ae, 217 lactucas. 2.) 025.0 eee eee eee IOI delessertiamus, 3.022) eenue es aa 61 Corbula dietziana 24...... 189, 440 Gemstar gus]: ie) ceuecetest arse 216 disparilis::.. 0a eee 438 domingensis........ ...... 186, 195 gatunensis....5.2.05-9 eee 186 HAVeESCENS oases ieee 61 hexacyiiay 275,03 4A4I, 442 Horidantias: 5.32)... tenses 61, 189 milivi .c... 2:5 oe eee 25 DATA es shene seer tateiien a aeteaees 226 oropendula i972. eee: 439 gracilissimus 4...195, 199, 224 OFOS1 2.5.5 ae eye 441 granozonatus 6......... 1G9, 222 SCTICEA, ...540). 222 Se ee ee 186 Tamristt) (Ole) sc ues tease oe: 225 sphenis...3...5.40.9e eee 186 Haytensisi se scc es, 186, 194 Subcom presence eae 25 AMPitatOr: (054 Neon sh ee ee eta 217 vieta * 27). 2 See aeeeee 194, 438 {uterstinckuUs: 6. .2-2.cee 199, 220 viminea 37...194, 196, 200, 442 leOHemSIG2ieca.-teee See re 215 Costa Rican Geosyncline ........ 205 LeONIMUS A ica eee eee 61 Costa Rica Stratigraphy........... 181 limonensis: | 22:35... 2520: 188, 219 Cranopsis. ......0c2nau as) ean eeeeneean 167 Marsimatus| |) 70.03 sere! 195, 223 Crassatella antillaruim....5-240-.. 384 molis \5..2.,.ss0s basteeeehe 188, 214 Crassatellites bowdeninsis, var. Vary TMetirAatus. .,6s0. ste es 200 costaricensis! 7275. eee, eee HOD UREUG AV OS 2s CpseO RRA OLAS 195, 201 gabbi.. nici. l. ees 25 niultistmiatus 4 6 cscess sec ue: 226 mediensis. 32....-<ses eee 385 WNUSAEHSIS” 52.185 sess acess eee 219 mediamericanus: pew .seen er 384 pe LE ee CRN Ly PSNR DM NRE 62 FeeVel, 23/47--uneee Be Cee 384 planiliratus 6.....188, 199, 222 sepulcollis:....c/eceeemeae ene ee 25 proteus 5......61, 189, 195, 215 Cremides, .......u.ieseenaneneet en iemen 16 pyemiceds 10 uses SE eee 62 Crepidula aculeata 2 ee 144 Py tiformiis: 4/26).05. eng IQI, 218 CONVEX. »...0. ssn eeeeaee eee 144 recognitus 4...... IQI, 195, 199 fornicata.....31;.0esees seer 143
218 plana... 7.2.0. es 144, 190, 199, 331 Tes wlanis ye ake cee Mane 222 Crepitacella limonensis 78......315 Sete aly ATA EL 194, 220 Creseis subula... »nivinisidatlonne ata Mmannetag 43 Sabie) ma el ae ans Crucibulum auricula;: 227 145 ‘ planatemi. . oii: aedeche sees eee 145 SUEAHMS Ue. lock eeeacaewesacee ance ee 62 : scutillatumy:.: 22.22 eee eke 145 stenostomus 8,...188, 195, 199 : Le a alain’ a ees dai busi ebelctd on ale aretmearas 145 : Cryptonatica,.:.,.c./apeh2sce-aemeseeree 147 EOROEUSIS IS cre cyeareracsnedser cee 220
Cucullzea macrodonta...............25
YT RUE SE INS Glace oa zee Safford i vais si decnnds baer 25
467 Costa Rica M1ockENE—OLSSON 295
Cuvierina columnella............. 44 Cyathodonta gatunensis...........383 Cyclostrema affine............. AND 653 MOMPMOLYy X22. 2.2.6.2 22.06.0600. LOI BS MOST, oo is. soe eo ee ans ee 160 TEGEMOIGES 0.5). ..0 6 eect ees en ee: 161 Cylichua, bidentata.................205¢ BURGAS adeeb vt didlos bcloce 4] MSO es acite ls ahsnes setedet «« 26 Cyclinella beteyensis 37...191, 414 TAPIGIICTASIS, 2.2.05: ache ls vadedoees 187 PORE A ndcase nsts cuptdese LOI SMO MMAGTAtA,).......5206.secsecies 191 WAG OUAATIA 9 34: 2.250. ce cece AI5 Cylichnella bidentata................ 50 var. galvestonensis.............. 50 triticum-tritonis............... 206¢c Cylindrobulla beaui.................. 52 Cymatium chlorostomum ........ 116 eynocephalum......::........... 116 SASS avatar aisle baeieiaieicin cls doeins 116 CARING 2. Sone cincavcvacevessseees 116 Cymatosyrinx centimata............ 67 TOSS SCs ohee aaa ee 67 Gyphoma gibbosa........2.5.......-. 112 Cypreea alminatensis /75......... 202 CUE Toe) eee 1 wey bs domiinicensis.. ...........- 196, 312 See ietasiS 2 ers ie ue ca 14 SrA MEMO AL... 6 cvivc.tsv sec ese te: 13 J NEI EI <2) 0 A ee an P 186, 196 PAMISMMAINA, 75.2 bet denn. 311 SPU Yb Salo sks ook sosoes soe adie: 15 SSPUIUEA. SHAS hoa ciacescecps. uscd 113 OMT CASSIS! oes. iselivelescaescess 118 Cythara heptagona ........... 186, 195 terminula, var. costaricen- SIS OR Metis setyicsksn one) 249 ey euarelia cymella....)...2.....2..03- 78,
MAIVeStOMCMSIS,. 's.,.2202. 6008-73
Cy thereas alberim@ata |. Vauerscsccoersse 408 Iblanichiama ny, si esea ee ieee aces 200 GATE eee aac ec cemented 406 JR CCA eis lei teas ac ave Sues 408 LALQUUNAT i Lsciecenccdelmsecseceye 412
D
Daphnella hormophora.......... ..- 72 lewcophilegimanis bic ccsase sce. 0 73 PIMMACITIA estate cacreecta son 73
Weirancia chy tar cee doi sacs ude 72 Perpatctlla ey ees swawesacss 72
Dentalium antillarum........... 25, 36 DAREleth ie Crass Moni oea alae 36 DOCASENSIS | TO se0cc.sesieseeanices 338 Callipeplam a. ass escecss ae 37 Callichrixer ye. sicst heseassene are ds 36 COLAS ee cucaaden cece veenecoucees 36 CORAEWIMIN == 4 ieectesscissclsesces 35 COMIPFESSUMD Mok kes. 8) eece 39 CICS MAUI ee cert seacate neds 39 Gispartle:ssi AG ess. cad. secedaounsed 35 CDORCUMA ee. jee he sewtenie eee a7 SLISICUIUS fre foc ccilastaal/cadeete es £8 TUE h 0c apa tue Scr eet APNE eae oe AT 38 HOTIMESHSE sc secssccacesasere cues 339 SOULGM Gers. he. ohacenecscceecs cs 34 LAQULE ATI: 2550 oe euencesabae 34 RSD EUAN We Bene) Ged ai new Sales ae ess 37 TRALAT A iticsncescwosss ssisisiseisceees sor 37 MEdIaViaenSes:is.c estes eae 2O METIdOUAlS, okies ea ceae eee eee 36 Ophiodone. sera coketeees saws 39 PErOMM UIT. cot ech secsn oe 38 PRESSMAN 6 ies hus e ce souewnes 39 pseudokexagonumy.. -s..52.-22) 35 Quadrane ular co ascessec a cnseas 39 FEDECCACTISISN (Oka etic acon 34 Semistriolatumy...d.-..sstesetenk: 37, SERICA TIMI 2s. kok, eucideaeees sete a7, SEX ANGTMAT SC! vot ac. beeen te 34
296 Costa RIcA MIOCENE—OLSSON 468 Dentalium sowerby1.. 20-20-60) see + 38 CATTUCA" Zikiteees. sasek eee 243 Var. pellacert..7.42..05 Oe ee 38 chiriquiemsis) VS). :25-eaees 236
Cea lawyers ee eee te ete 35 chytai iia. eae eee ed tetragonum:!/..:::. RaAny Le gasadisle 39 circunivoluta... ds eaerecee 64 COXASIAM UM Sees! aoe 34 Citria 8. 237 Vat MCCSUITINY Lote al moat 34 COCCOS : Oy2.45). Lei eee ere 236 uSCarianunl 752) ee 187, 338 cocositia > 8... suadscur ene aeen 247 Dembistylansce ci. woes en meen 156 comatotropis:2cs..e eee G2 Webra ciate ilies. i) Un te ae veeeian 56 consors 7....195, 199, 200, 233 DIACHIA ee Pie ER PRE HNL SA ey od 42 cristobali’ 6473-3 244 Drplodonta collinay (255/205. a6.. 392 dalli, var. C€Strota yeasens 66 PSTN AE Giese cel araentes ge Nal 392 detecta .)onk. 2 eee 64 PUD AV AS, Leis hts VOW Ped eel e 7 CDUL... ceesclasie poe 65 Distortio, var. clathratus......... II5 ElUCIVA 0 loa ceek cath vee eee 72 SASUUMENSION nna eects 306 CNNEACYIMG, <\cm eeeastentieoeenene 186 reticulate. a wanes Date TI5 estrellana | 77i,25)uaseae seen 241 Simillima’....2:, sud one sesisesammene 195 exasperata:: ive. Sa athe Malate 65 DIStOTUD REE Luca Meme in aes II5 exscul pani Jc) 7 caterer eee 73 Stim ay ce eee en oe aie 305 eucosiMa, Var. Ca mmasisinisas 64 Divaricella quadrisulcata 35...189 fusifOrmiis. 2) leeastene eames 238 200, 205, 396 fuSinUs.. sa escent 186
Do lnm eke cee PAA hea I1g Patunensis.!).,ddecnan sees 186 COMTI ar a. Se ace» 196 haliostrephis juss... sours 65, 235 Sllipticay site ssayatteas ee eaten es 196 havanensis.. .useurececeye wee eeeae 67 ellipticusy she eet 3II IPATAss ....025s bee eseae mane eee 73, Donax acutaneula ip ila... 22 leucomya....0 he eeeen es eee 64 Dosinia acetabulum }34.....118, 201 limonensis S.L.cevscese coca 243 203, 403 limonetta 3.5372 aeeee 242 delicatissitiats <2 c8 ee!) 187 limoncia (7..! ./occsmerecmeaeeeeee 238 Drillia acestra. tees ah ene eae 65 liSSOtrOPIS..... coe meena eee 66 albICOMMarte: Pheu leumeeeaenues 64 lithocollecta.c ce. meseteeene 67 albiodata.. Js 008 na can 65 lithocollectoides | S.22) seu: 246 alesidota, var. macilenta...... 64 lonsdalli...:.) Dee eee 7 AQGUAEHSIS 6 Oe: ah SE a eae 245 MOENSIS: i... bee eee 235 AQMARICA:| (Si ssise (Ste cesar ec 237 THUS | Goin sande enyea seen aeeeee 246 AULAMUIA Nous eee cee ec neceacn 240 miusacina: |S iT eaee 241 amet 2 Cees s be eae, Saws 72 nucleata 20). .cuveeer kee waaeee 66 bocatercensis) i821. <ali as. 239 OStLEA TUM coe sneer see nee ae 64 DOScensis yi Qs ewes ee ee 245 pagodula joc scenes 65 VATS CAanMMaura. sd. ceacele selects 66 papaya \\)\ 7isdecton tee teen 235
469 Costa RicA M1ocENE—OLSSON 207
Mag elagia....... 0.6... ceceseeeee 72 EDN ee ee ee 71 (Sic) 0 Ga Vee eniee 65 SURI SOMIL cose ctsciidcee.seees santes 66 SPEUIO NIM ss N oide nb bo.s'snivctoe send 73 TAG U 1 se e e e 66 PME OO KOMI 1S .2.a.s secs sek sedess 240 Bea AM Ie SAS aic)s Wes es sabes he seiteds dee ve (ESS US) 0 ae ER oe ee 65 TMS MGs cscs sost let esos see 186 WEMUSGA, (7... 005 scc0s 195,, 200,.:233 “ier UB bE ae Pe MUN 5c 66 ZACYSI SUES AO RAE a EE 186
E
Behinella nodulosa..;............0. ley
Echinochama antiquata 37..199,390 Le (10 01 00 391 WaqGersis | 77.....-.<...6.194, FOI EOE LG ee 24
miaeeretla subtrigonia,..... ............ #]
Emarginula compressa............. 167 He UTM eh test, ell oc Velvwe anc 167
Ema bOUIS Utatus...i..c.cc. 2h. ..ccce es 4A
Hpicynia multicarinata............ 161
LD (SIS | CLIO cg ee 38
Epitonium angulatum....... abs IOI (SEL CIC 05 0d Ute ayaa Re a 102 COG MEAN ee bls ses lek se lasses des 103 COMTORG MALU, 3... si ecas cenne 104 ‘SISsOTG OLS) Gn a bh 50 beeen eee 102 PAIVESLONECTSE: (205.5. ods cs see 103 PPM SACUMM ss eee twesesaw tees esos 102 PROMS o's cries biaslnc dine v clei ete 103 PM OCESEUM 205505) vee ved ve secse' 104 Beiter |) 7... 00.6 s205.. il MMCUMEIS ETA TUT, ..03.2..0ccvedoese 104 PMUMISCRIATUIN | 7.0... sesec8cssce- IO HOUEIGE UNIT, (00 evecare erinnda eds 102 RVR TUUBERD Soh. Ls po a's odoin sclmowsmale in 102 BREA MG Pet iN alia os vaisceeecicwensess 102 BAI SLAP MAR Se's,o¢ dca sssie cle aivis'e con's 103 SUPACUCUIN 7.52.0: 2.2 sesboveee II BUMS G ss apicle sncee antiendteaeeoace tes 102
LOR IC UIT tae) Voewe yu sanenedese LOS
Bratomlsevis ies ul beret yee as II4 MIAN GETIA LL enione toe soesdos eee G 6 EVrvilia dig nitican gr edes.steectssess 21 Polka essence sens et eeoeeee sat 2 22 Bthaliarechisai su: teecteses soneees 162 SOG a Heh oe ccc aeeay s 162 SUP PTESSAs es swccetenousssese wots 162 Ep ioalany udiesan esa Poneeud arenes nade 72 J Bib Wo oa bz ny Punta ine ran prem ap Ae 105 Bulimellacunifasciatar..:.20.5.5.-.. 107 etarae bas he Mes wise eee demmenanmen’ 124 Eunaticina semisulcata............ 151 Huplenral caudatar icnsssoe- sesce: oe 99 var. sulcidentata;:.:::.-..2 99 AMUATCIPOLMISA |i coscscesstect ee: 99 EUS ital nw aeee seas cua tec soko. weatne sate 147 Butrochws..0 4)... Beale ten eesesea tau 155 bxcilia spereraciis 7.4.0.2. <ss--cs ie F Hasciolaria distans: 2.2.2... sp.sccsnces 83 CIGANCEAN ce servaceleweueaensais 83 POVSASIANa | h7 Ti seclasdapeitescssene 277 macdomaldd | P7725 eoc-satecnes 278 WNC CAA WA: Woe caieeteceee weeree core 84 CML Ai On tatoctneccss 93,205 1277, Mavartia, cellulosai cs ie. case sacs sae 96 Var OviICUlas cise sce: 97 Patermred iay. sca Cuter nests 97 WUTC CUS ce dae dans seca tels sep (locmactent 96 Missidemtaltuanii ii. ossetateesccceseeenes 36 Hissurella alternatai:: 22..7sssce-s- 166 ASLUTIATIAL ce shunts cece dneeteae 167 DAL DACEMSIS shane. stncdedee eames 165 Fissurellidea fasciata............... 165 Hissuridieavalternata...1: 27 1.cecss.54 100 CAVETIEHSIS. 4 osc sicosnseeticcsees ASSHTISCD tats. cedads Seog ce ethic ease 167 Flabellum conoideum.......... Saeed HUusinue:cepyMOtus;.....cc.0cvetoc se 85 SLCINUUIS coe iisesonded Cueaaaucenetaes 84 atiniamtisiaccr use secencess seee ee 84
298 CostTA RicA MIocENE—OLSSON 470
COMET NE a UR nea 84 CUCOSHEIISI Le Hees 84 EIMIESSWS) 25 Line. gs eee 84 HOMERSTS) i FT weit ees 276 IMIOCOSMIIUS TULF ales co ae ade: 275 MOHEK ccc. SL Eek aEL Ye Sa Rue 26 ORCORIS il 0 0s asennad ce 26 quercOllis: |. esec..6: ES Lateamasbe ete 26 tesellatusns Goose: seewsnense acest 8I COTES oie eteN een oie aeaen ee 26 G
Gadilawy salt eked ais se ee presen 41
Gafrarian altum, var. costari- COMSE\ WAG Cire ecn teas aioe 406 limonense)) 3521. .200 805) 405 Gastrochzena Ovata.............cceee 438 TOCUMEA iin ea eee 200, 438 Gatun'stage cite oe. Sates 181, 162 Gaza fischient) cites te sean hae 153 SUDEEDa a Gaius teotemeeee Deeipleiete 153 WAESOM Lil icen Honinsee ne boeaines 154 (Gemotadidiymiaytycices sae eee eee 63 mitre bla ie ease ae aaaiielsiss 63 Gilbertia estellensis | 2y2/2/...413, 26 Glycymeris aviculoides........... eZ 5 Canmalisi er cel eee eres 349 Carbasimay) 22 ks ee eecbi aes 351 Castameus (727 -.25/0.0/.48.. 189, 352 G@ecussatuss 122 024 wae vce 352 Glycymeris decussatus, .......... 189 jamaicensis 22...194, 196, I99
3a°
Hoydsmithty 25.054 02.0.. Pepa 352 trilobicostatwss. ccs ees 349 Glyphostoma dentifera 195,190,248 abi eee ees Raa es oa 69 Sratulany ese. peiveatdeee Poaceae --69 THORAC SU ae se ee wee gl 248 Graptacmer iia ihasee wi Se ee) a7 Gratelupia mactropsis....... Senet AIZ
Gurabo formation........ TEER Alea fee)
Gyrineum 1)... 2 ee II5
H Fialia americana \7a\ aa 251 Haminzea antillaraim..).nee eee 52
antillarum, var guadalup-
CHSIS... o4e gee ee 52 elegans... ince eee 5! Puildingi, :. ae eee ee 51 petitt. ..54i0C Ue eee mee 52 SUCCINEA: |: 5 eee VITESCENS 16.5 ee 52 THaustator, .:.iodics due 134 Helix depressus 2535s)... I4I Heteroschisma . 33.525). eee 36 Hydatina physis..2.)2 eee 53
l Infundibulum..) coe ee 145 Isapis ebsoleta/siia.ee eee 137 Ischnochiton limaciformis........ 168 papillosus,.. oe ee 169 puruprascens Gece 168
J Jaminia .....5.)¢:9.0 1 eee eee 110 Janthina comm ins 3 104 fragilis ..)15))/).0 eee 104 Plobosa.....3252..acr eae eee 104 janthina....... ee 104 pallida. ..:.. cee eee 104
L Labiosa lineata 25.....189, 205, 436 Leevibuccinum harrisiy i234). 3- 16 Levicardium serratum ............ 200 sublineatumi: (223 eee 200 Levidentalumy |. eee a7 Lamellaria raugi....2.. esse I51 Lampusia.... 154.05 ies eee eee 116 LatiaxiSyiiscio., 2.05 Sa eee eee IOI Latirus infundibulum /77...190,195
200, 279
47% BULLETIN, 26 299
MPURMSYMAZI 7 7......5.00.- 0000.0 00 279 MPU AN UTED ooo oo). sy coaceeeeces 280 JL 3 Orev)072) 010s ey re 345 S12 NE) eS a rr 347 22 Cane eee) se 245 EmeiMOIdeS 27 .6....... .e.es... 346 PATIAETISIS 26.000. 6 covece ces vosns. 25 EINE. AS So ae 2S BPE E CONSE ie Ges kese toca ck 25 GUIGAMENSIS 27.000... ss...ss, 346 SGI) GLE Ea i 25 Memetia tWbicola.. ..s........ 5.00006. 152 Lepidopleurus pergranatus....... 168 MUEHIEOCMILOU, 60... cbecc. ss. lsc occsoeees 168 Meprotityra induta................... 153 LU STINS C105 2 ae 56 Hevcesyrux chloris &............ 231 Sra Ole tater nd! Adcaec sax obelsee ses 63 SMI TWME META! 6.5262.....5 soe ee 63 WEMOCCEAG Sco ss edescseeiisas tics cscs 63 BRUM ere sal saelecicoliscies'ue decide 62 Leucozonia cingulifera... ........... 85 OS SWINE Sake Eee 85 ey WSUS! Gadel oo... sc eckee ones: 26 ECOG U Ce Re Arm 26 SICUUS TE EG A lee ee 7 Wetter sa SOU: es. o Si ee deacka sacs ss 194 HimMOpsis MONS 20...... ........ 349 Mimgea solida’ 77.........+.+.+s: 199, 381 HEAOR MUSE a. oo ells cas sseeee oes Pet. LOO ONO WAIEE Us vcs deceses ce we eseia ens: 154 OILS EOE: SO ee ae 154 Gone atMMAlA sc) See dese ness! 154 MisSOSpita PrOXiMa..........050..05. 160 MAAOM AM OOM DIX, . 64..6c002cec-cseesss ie 7 MPetATILOStOMA. ....6655600).00500- 137 Mattorina, auguilifera................. 136 PROT AU A Nols i Secs eieln sels ceessess 136 [AUS 3110000 0 oe 136 PEGE AAUS teil. cee ede eeven wee 12i7 ZNRTEN TS SS Ae a A onlee 136
PENA LCA os. ccs soesee cles vecestent 153
IOS COASTS caer toes ene ees 107 TOGO TUT TIA A ee es ea TN ee) UWi7 ucapina jadspersacest ssl ee. 165 Cancel Tatar sitet sion e emus. 165 PASCIAtasih) Sent sticsceseeccests tet 165 NONGUrASENSISY. ets te eee soe. 165 HEI B I-19, a) Al ogee 165 wena \enmStataw ts. jcee-. seem ee ee 426 Ginyari@atal ec. eile uk oc bal 396 PUIMIOIMSA ye ees siete suas 19 quadrisnlcatay Whiis 35, ease. 200 POUT LIAR teed beacons ek Hog oucume es 148 M
Mocoma costaricana 29........... 428 GlOND ALAIN Ine wel tmekecise 19i CALWHENSIS 1720 y.0c% 168s. sce 429 PATMAMeNSISH! Vo eon coves. sete. IGi
Vai, Canalis) ' 20.) 6tcee. 429 Macrocallista maculata 34...... 189,
406
Mactravalata, 197 isis. 189, 194, 435 GIECONGS Hedess gexec ceee sonar 19! estrehlamanp gm. seule I9QT, 435 EXO G7 e ans 189, 191, 434 TUR CAULER AEC tows smeneees tance ART SINCOlA Rieu cheese ne ees 436 Malea camutasssi:)s5.25 186, I9I, 200 GUDEICE R715 i SOL ei lire 2350 |
PAN EMS is rek ss eos ss cece we ecue pees 19f
WWE Wcak aan tee gh Aine emma np nee tN Pra EN 149 Mangilia. amtomiasi: if2)e.tac.teser 71 ASUMICE ANAS ticne esse rac duets ot 69 ALUOSUY lav asiscc as seorcos si eecuac gues 70 Dalt@ata ccs res.c:chetscbtes semcwaee 69 WICONICAS. Ao ete ot octet cate 69
el 2) 9101 ts NI ee momen Miami ee Rist Bad 70 Ceninella nesses ube ee 71 Timon tela yee) eae 70
He lAnItiCal youl cscceccauaos. 71 PETIPlA so esas eescuscecesseteees 71 DlicOSal wel ate eeeey ee eae ee 69
300 BULLETIN 39 472 Quadrata.}202 2. ec. iis ae Bus as ys Melampus bidentatus...../....2220 55 PUbe la ee eee eae care det 70 biplicatas 2.0 seen 56 stellata wow co neue ce er aaenead 70 bulloides cc eee 56
Mao Clayiiirstcsceiaen sce danesnmes 193 cironulus 3.05.0 2 55
Margarita flosaic ircktesaceeeee 154 COMMEUS 13.) aoueRe 54
Marcinellaapiciiaye.) occaatee eid var. gundlacH¥..) 2255 an 55 AULCOCIMCLA. 15 cecene see tee cee ek 79 coniformiis....;2sc.cee ee ee AVGWA)) WOL-es 80, 188, 190, 269 COTENALUS , «0. .ussaheseeeneeeeress 55 Cannas. hictes au Coassnseleoemecns ef | CTOSSEAN A. ba255, se eee EOP Ee 56 CASSIS ue hoe hoped tadecboesmngees 7 flaws ...i.).sseeneasce eee ene 55 COVINA Hh TOUe Reese een eiss 269 floridanus,....42 Ae eee 55 COMMOFMIS) <ess sees eet eeee es 268 hemphihlt. |). c:g eee 56 CONOIMA]IS HN. dabeseonaseee sates 78 jaumels Pron Ere Mele iy ily 56 denticulata.125-e et ees: 78 lineatus, diescisneseeee cee eee 55 EUS UIA ty Suen: Cetera eae 79 LOTOSA.. ais. hececck nese eee 55 PuULAHeNGiS7. se were een ee 186 Melanella bilineata................-- 106 hadhiay it eee eee a ee 80 Conoidea,....s525 veges 105 Lactimulac. ices een eenerse Pesce fUSUS, ..35. 54 eae eee 106 laticsiimay! WS tes merase 268 Zibba...s..5 tis heey de eee 105 leander Qe uaa scowe tase: 270 PTACIUS...0..casosmmcntecmsneeenetes 105 macdonaldi Dusvnccbeeree 187, 267 hemphilli. Jalbercb al Se Seeee creer 106 mindiensis ORs eee 268 intermedia jee eee 1 ar RNR 105 AMANITA al ee eke Setea oa 79 Jamaicensis -.)./:)2ree--s pee 105 PMIN EAC eaten ER 79 subcarinata), ..3..c0¢ceeeeeeee 105 musacina 73. -.sceee ee 270 Melongena consors 72...195, 200, OblOMga eee, baleen ae 80 284 Opalitia se ola weenie, 78 COPONA, oo 55.405 2sc2 eee pee Rene 86 ovaliformise.s.o-6s5cee ee 80 melongena............ 87, 200, 284 pelluciday. 3.0 sya 78 Meretrix dariena;22i)-2 5 eee 404
var. pilsbryi Vip eermoaes 268 ripleyana jaclselgue oidtarelele sievalereeceetaenine 25 TOSEEAbAL. es area beueenee sucess 80 Mesalia alabamiensis.................26 SEHIIHILAY a ee ee 80 pumila ..2.5....0/:.apeeere eens 26 StORETIA eae. ene 77 WwatsSONeENMSiS. 0.052. /2seae eee 26 SUCCINES, 2a co sais stdadsccnsbieeneee 79 Mesodesma costaricensis 3/.....437 virginiana. Big. TAR Reape ba hs 2 78 Mesorhiy tis, ...c..22:.30eo eee eee ee 84 WELLS ELIS MOON Weta UEC aS ey Gey 78 Metis barba 32......0..+-seeceeseeees 427 WIUCACeCAITA La Ne TNs i das 80 Metula cancellata 73...... 195, 200,
Marseninaiampla i cees I5I 287
Mantesia recurva (2.0040 b as oao 19 2410) 0) i & pee +++287
Mathilda) yucatecaiaw( Nee: 135 HaArriSt 13...secceerereerrer serene 288
Meioceras deshayest. 2.0). 131 var. limonensis 73......288 Sera LEED E Ds Pema Can Minny eine aKa a 132 Mexican Miocene ......... csp 200 TINGIULOSTIER eee ee ena 132 Microdrillia rostratula eee Ve
A723 Costa R1cA MiIocENE—OLSSON 301
Mitra alamagrensis............ 188, 275 Pe leMsee MINA). 2 vai ai at decaistine'd’es 82 war coralliophila, 9.......... 273 MEMTISIS Qe) oo. cock es yacasewes Page: SPCMUMAG ALA cis caiese'c «paced ov sncto's eo 82 Walt lIMONESIC....0.02. 00.665 IOI MOTUS iss. eb-intedse vos e LQ5; 273 OAS Ol encicleeinsinersiealhe TOS. 27.A. CUACUIS AOR RSMAS ee 195, 274 Apa IT INGA 2) ses tie ie ctdutltinn ece'ee elas 82 Saree acinar el baie elaicd 82 SUA SOMES. ssc soe cocgns oeies.o es 191 swainsoni, var. limonensis
(ON Mi le han SN A 272 DN AANGOCTISISH Stele case veteceecucess 82
Mitromorpha pygimea............... 82 PATER ALA sett eck s cus bees sinmenes sds 51
Moduliis floridanus.:............0... 129 GEO GIUM Gee ecco ioe sis lectietes ox « 129 SUMONGGH See se ytd t sytacs au danccens 25
Montacuta bicuspidata 2.......... 20
VCH MCRUT IS scusscsssisccruoeesss-02 1 Mulinia lateralis, var. sauryen-
SSG Lr tee ain slewiaid Deiaila S's 438 Pimmicidea Hem lili... 6.5..3.... 98 Miunicidea hexagona...... ........... 98
OULD} G08 aife4 by A eee ee ee 98 OSPLCAT UM, 250.255 cen0 000 bieialsiote o/s 98 OMIA A ete! cowabe sees 98 MRE XCA SWEHIMIMUS io. 6 os. she teoe ae 96 OSG UU a aN 0) Dente eer Ree 95 IaMeMTE ROHS Y, Va dc asl. sclevist sweaes 96 RPeM tA Bea eed, nb ree swieie,, ei ge mille nisinctele 95 BEM OSA ois tues er ssees tose 96 COPMURCCLUS. .. co cee sa. -o 195, 303 HUMES CONS cos sale vaca cals tactiielue on 96 PRAMUCT SIS 2/05 neo tsks's Eien enon 186 (21050) YOKE WAR Sea a 96 MeSSOTFiUS.. .... 95, 190; 1955303 MEM AEUS er), oe stccks wweteose cise veisiees 96 BOMVMEMAICUS 6.525.050 c..cc ues 186 ROOM IM gos oo econ a seauioe. seni e 96
Myrtea limoniana 35...... 200, 393 N Nassa, beaumontensis. ....2....25.26. gI HOCESSIETE siitsecesiee eg eVences sgt QO ODSOletas OR le vast oere esathueee rere) PUGOMEY WE eine ste ae ees tiwenies 12 IWiaSSaritia DSI. 6y 2 o oiceessbeselse ces = 89 colmmibeliataln vareate, seessmnc ees 88 DEAN Maia te cena Mamatainem ees rsa seo 88 Naticaalticallosaicc. )..J.2.s.ted.. 147 DOMISUIS ceases eetases camtentes 186 Camalizonalisn: wii stcieet aeons, 186 canrena 70..147, 190; 196, 199, 327 Gm ane aesstaa se nsesr eee: 26 POLGIA TA Seow aaeanecserened deen: I5I SUP PVA War a LOn eee 's-<: 188, 328 JAMMAICCO SIS wceescas sane ees 146 HACK CAMs: 2. sculede cositsjese aude od ee tne 149 VA Aan ea eres ae sas desees 146 [bb sohbl eRe O ME BemAM eetRere oer e 26 TIATOCCANG..2,.0.032.sesseses mets 146 OMS Cacia. eecedea oes eiae nets 26 piacavellare, cols ccs Me stor steels 146 IL ORCUINIAN 2 y. Panneeak duos cee te de 146 pUStla oie access wom te eee 147 PEVETSA se vacis tee. ose EOE ages 26 SadrOrciae oer ws onset eet ees 26 SUDCIAUS ARs. e: ties obi daslesnee 329 SUlCA tay. Maser sila ecstedeue teens 328 VOUMID Ge le oe ON Scat ataatauietns 196 youngi, var.cocleana 76 327 INaticinadacteay ys. Uo ecadet et ate 149 INCOSIMIMN TAG ras cets bua bea odu one LIZ INentta (pelOrontan;.. .cucasewsecar 162 tesseblatact ts odes ae ice 162 VELSECOVOL Hae eie de osname tien 163 INGTItHa PUPA... soe. be ueeucseebat 164 CHE CIHVabas {ccc taedsseeeneracan es 163, Showalteriky se sas necuaeee 164 SPatsi liMeatay nessa seseaecten: 164 VIDS INCA cc.ayeeaabesieeeaonties 165
302 CostTA RicA MIOCENE—OLSSON | 474 VITIGCMIATIS ZO. ueeuse se 196, 331 liodeéS: ...).ceaseeeeneeeteante NE 75 VATAGLIGE Ce iepatcoanecen tase tetris one 164 litterata..: iss lee eenber meee 75
Neverita brunnea)...2)25/ceeecore 150 maticinella) (703.302; oa eaee 263 CAMIPEChICMSIS ash -ueee sees 149 reticularis <..)c4a0-paeeee OU RN 260 duplicatasy sy passsccorerass tenes 148 sayana, var. immortua Zo CNA CAls eee ceaadaaaeae see accar reer 191 205, 261 SIMA TATATISH IE Pte clon Faceemmerte ne 150 testica, var. costaricensia Nereidis...:....187, 791, 196;2330 JO. 190, 262 Dla ee encebepest es soseenene 149 testaceéa’ :....orte sees 191, 192 EEOXASIAUA Dieesa ears «loos yrdietiines 149 Olivella, var. bocasensis 70.....266 WDerEMIA) Oy. ui Halos aisigee since 149 boussaci (JOlje-bessee ees 265
Niso eeglees..... Leet Ba Aa Sy A Wr Ae 106 Clitacta seme vals Gehigeaeeee anne 77 Splendiciitiaye ccaecserheerssssmes 106 floralia ....23...6h. Naneeneten near 76
Nitidellay parvilante.cscs-mechess-caae a 94 galvestomensis( eta. 77
IN OLOM Git senieress ioekceesaeenter ane 74 goliath | 7oye see. Lg SARA 264
Northia moerthice-32. to. ence 190, I9I jaspidea, . ... 22.8.) seyeeeeeeenes 76 northiz, var. miocenica limonensis, “7Os rare 188, 265
POE AE KSAT 296 Mediavia.. 2. dees eee eeneemee 26
Nucula cahuitensis 27........2+2-343 MUtICA..1...5 shennan ee esnee ne 76 ASU THICAy eesteeetae ate es wale meee 186 muticoides 70........... 195, 264 Peep ENEMAS iL Say te be icc AbE id 25 TMIVES 0. «ao eareieal anise eee 76 orpicellay car lure 343 pusilla. - oo... ie cueameeeveeeeae 76 OVA heen eas ae 25 semilignitica Z.s.sessesereees v3 tenuisculpta 2/......... 193, 344 subtexaua. |... yescsneme eae “wi
Nyctilochus femorale...... Nf oe 116 CESTACEA. ..o ii. ces scoeeeemninrmen ere 192
ONnISCia... ..046,. .2008/eselatner neat 118 0 Oniscidia dennisoni......c.scse000 118
Obeliseus) teselllatus. } to. eyes 107 Oocorys abySsorutl)..2.322-poenecne It7
Ociie bray ise H sacle wae eeee et 117 Ostrea costaricensis 2¢...201,.
Odestomia acutidenscoy-.-- i255. IIo 368 Hartschae ans Wee eee eee 8 gatunensis: .:. nse eee .186 COnotd eal. sleet Bat ine IIo intermedoides (9..ce upc 23 IMPTESSA Ni eceneeay sce Rew nce or 110 megodon 27/...... 189, I9I, Seminmida syne euectas ee IIo 194, 199, 367
Oligocene, \CostasRica.: 127. .1.5.) 181 sculpturata .2../chbe ey serene 201
Oliva brevispiray 70.92.25: 195, 260 Ovula reguiaroidea > 7 Wee 14 CATOHBENSIS Loe ee 75 Ovitlum 4.2.0 hee eeeee siigaien cee 112 CICA t ae ee ee POO ge 75 antillarum) 0. vee ee I1I2 cylidrica FAO 195, 200, 260 ATCWa tui. eee eae ine II2
Oliva gatunensis iy alee ees Wout 261 carolinensis..{) 223. I1I2
subrostratumal.)2..).yoe epee Ii2
PIFAUGR cs Mialsavenateceneer eens 261
475 BULLETIN 39 303
Omyeyrns iceraudrenii............... IIt P Pachycheilus anagrammatus.....129 SEMMETSIS. 62. cn arioisce ss .vencesenes 130 CIE aks 9 eek AA a 130 Paladestrina aldrichi......0..... .. 141 ALAS SIM ATEA cogs 0) casa eisiee ance es 142 CUO 12) 02 a an aD 141 MULTE ls osc cceauigeseocees'es 142 101205) Ra ae 14! MOMENI oe. 8 ss saudes-cuiecitiacte os 142 Eanopea belisensis 7........2...... 22 FAME HAA ee elcedesctdesse. ces esccss- IIO esiemed SUttUla:. 1.5... .s.cecece sess a JES) TSI G1 Soe 5 ee ae ee a cope tissa CRIIESUAS io cote wse'v sense sees sees 143 PQUUACAUAN cece’: soceees vende ss 143 Ve aiy inane (2201 0 ee ee 149 Recten alabamensis...........0...... 25 AIMMIGATITeNSIS 27........ se. 375 NOVELEIS. B12 AS a Aa 376 eoralliphila 20.........: FOO, A7t COSHATICENSIS 27.....)... 5005. girke 110 52) \Aa Ne Aelia ae ee 186 SaAvIMEHSIS-79..........20+.065- 369 levicostatus 79...187, 200, 371 AAI FON I rras'euials cicleche ul Mle de eee 377. MACH OMALAL!) 7O-..) 6.00 6..0205 370 OUTSOLE PIS! 62 sos. ses Pepa ete 375 opereulariformis .:.......:....: 186 PREM PtUS = 202... .3...kees. 2 374 ir) (5), DIU Rar ara wary CoE 186 SaHtarOSanus...:.:..-....- 200..27.1 MEMELICOSUS.,... sas 0dsectese esas 192 Pectunculus castaneus.............. 352 GeCuSsatus, fs... cscce ll 250,452 Pennaceus ....... Se 350, 352 Bedteularia albida....i.........05..0. Dia lGCUSSatarrs.:....... rodiectrs cutee 113 MEGIMESELOMCALUS....052..cccscsteoee 54 PAM MOUNG SMe. e | ccs seseceredac eset 54
ETE OIC. Se suecveedeuicen 54
OMS os eee eased A cisewoiate De 54 QaaGiiGensy ye a eee a ula. 54 BEIGOIMS) Ay ectek Meccan laos 54 Perache weticuilatar wuss. steve os cose 44 PReristeniia InSite sy ace tee 3 282 LONEU BELA WN si ovens ee tesa. ce 282 Reristichta acne yy. oise. cine sacs L0d CORCCA NG Ny tanec teaenwasarstecieee ae IIt Persona isimiillimiay.:,..c55.secsn 20 305 Petaloconchus domingensis 204,318 CLECCUSS a hoc e alias ace vetesstentee ces 133 HE RECUATISY. Wwe Lane, wie nas 132
sculpturatus 77...196, 199, 203, 204, 318
VaTAATISH ate ye ta Ueh tds cock agen m3 Petricola millestriatay. 5. .2.secs- 107 MetrotrochUsis teases Uetectoes 164 Phacoides-actinus: 35 ¢...... 200, 396
DOCASENISISH 15 eo. ctosteuleses 395
COSUATIC EMSIS oes. s estes 394
var. medioamericanus........159
TAGLANSIH 2 cca ve cseacueess 189, 205
radians, var. medioameri-
CAMS UR Sei aces sae coon dla 393
LrISul ea tus. esa. 189, 205, 394
TO OMUITe NWT, C75 ean 200; 307 Phasianella....... CAN cae te 136
ID ENAS i ran year amie sues ata, age 152
THOUS ea a ies Seeing toeLiacmcae eye
pulehellan iets sees osname. 152
DUNCTATA ee cess x RO EAS ey oe 333 Pitline tle xtOSaseiee teveur teal tee. eae 53 Phorus agglutinamsy icc j.ce8c sc: 332 Phosiantillarva es i eeeeaws 88
DELCVENSISHy U2) tio. dons tdtcee, 294
fab al(G hey Lae AWE Maree Re Re Ae 88
CLES AMS eee oweer arse na 188
elegans, var. limonensis
7 AE CLES aa SrA 291
ESELEMEMSICN 22 501 ula mates 292
PAlVeEStOMENSIS css ceca eee 88
PALWMEUSIS! | 72s eee 289
Prateloupensis. tase) ounce 88
304 BULLETIN 39 476
PCTACATMS 2 nese ay taser eee 88 meriloides) 72.0 Pee ea 294 IMMexiCanus) | 725....200\, 201/289 Moorei, var. costaricensis Dias Dhae Deus deiasisereoies 290 PAL VUS yas ssdostebooceucneeeacees 88 subsemicostatus 72........... 293 Wealtoht is 720s icuenccetce eases 293 PHylOUOtuUs sss .c.caea ee re ee ees 96 Pitaria boucaryensis }Z........... Ail CircwMata G77. .ctces I9QI, 194, 408 COTAINE el tate. coli Pe NB eerie sor 187 SUPPYVAMa: Wz... eee ee enone 409 hilli, var. musacina }34.....409 plamivietaas I Wie ces 409 Vatiwittkleze (96.0 os cos Ae 410 Placunanomia lithobleta 24...194, 199, 382 Rlanorbisophise ts). tyes 56 Platysehid Goon coulis Meek ete Gee, 4o Pleurotoival sae sree ere sea 63 BU COMA AMR Ad Sedeete at ee eos 26 abbnda {hic Pe gs Rae Beene 230 alesidotas. io ea ee 209, 234 Fela 0) BE een nae AEs He in 66 antillarnins dese eae 220 Date ital a seid Ore ee 230 blakseasaay Se et ear epaecun ws 68 Vers briminme@a ir. scne eee 70 CORIMAD Rava sob Saar No Una ral: 70 CORSOTS Veh ela Tune aan 234 ParaMends yey ee 8 PTAA Om aces se ene ieee 69 DAttetSis ne en ek ie 230 pa er Bh Ogg 4b UR eneb eb MAmnaaUn ahcdrapn ACen 8 INSIQMHICA LN SO venue ny 7 JOMNPIPErSAL Do ete iaen see 26 Vat. Miaemnaok a, ee 201 Meda. ee ened 26 | OF ok WIAD Nee rene 4 ere Rd Mest 26 plicataw i nee 69 QUSCCOMNS Ey i weet pu 26 Sim pPSOMdss La wae PA 6G
streptophoragigi..0.caa ane 68 venusta... ea ae 233 VITZO |. eee eee Bpa gs Bu §! 230 Pleurotomaria quoyana............ 164 Plaurotomella agassizi............... 68 blakeana (2.20, 2oe ian eee 68 brevis... 3h. Eee eee geen eee 68 diomedece.: nen ee eee 72 extensa...2usi een eee 68 filifera.. 0)... eee ee 68 hadria...... .i tape ee 68 incilis, S324 Lee 69 leucomata.:) 03 ee 67 var. mexicanas) 68 packardi:) ...2.435), ee ee 67 Plicatula densata 2a 380 marginata (oye 203, 380 Polinices eminuloides..............148 fringilla |... caine 148 hemierypta.2.i/i ee 148 leptalea si: sueeeeeeiee eee 148 stanislas-meunieri 76...196, 329 subclausa | Went 196, 329 tenuis; eso) 1. ee eee 147 Pollia 2... 00.360. 2S 87 Polyschides.......:cas-eaesahcoseee eee 4o Potamides, var gracilior-)...7). 127 matsoni. .... J: h eee 127 satillensis. .j... tee eee 130 Protocardia costaricensis 30...401 Surabica) Joy. ee 194, 196, 401 hharrisi.) .12).25.¢30e eee 25 Psammobia hartist “77. seeee 24 smithi 7... .cdsisoa oeeeeeeeeeeene 23 Psammosolen gatunensis, var. costaricensis 32)).....5. 432 Pseudoliva scalinal 2a... 26 unicarinata ...2... whee 26 Pteria tnormata i773 see 194, 367 Ptychosalpinx dentalis 78...... 187, 281 Puncturella cirenlaris:.) -es 166
triangulata...0. eee 167
477 Costa RicA MIOCENE—OLSSON 305
Pameturciial trifoltum............... 166 RS 210 oe eee 166 EAURNR OIA Se eh. sos ain ches dadence de I0O GCOS: ee IoI MOCMMOSUM oils cce cca one eye 305 Pyramidella candida..:............. 107 VRE SEB Ee an 107 CO Dea 1 107 MM TOMEULIATIA so. oly. 2 iors coven if) DUM SOUS, 6 Zs. esol veidiseae os vse if) SILOS) A Se 107 UG 0 a nS a 108 Pecan aObDreViata.... 0k i... IOI 1X25 0) 1020 eae 1 fe) R LE BSCE ASN AO ANE en 114 lereiiratans 0 oo i Sk 99 REA SUPE te aad ise ats Joie aes 306 1S G6 ie Gh (cl Cs a 49 elt Z@ CMS ii. vececk secs scsccesss IOI maAcimula moOdulosa.................... 305 UPSET UIGL AS a0 ee 167 SUMMA eee. eos a 167 Ringicula dominicana.......194, 196, 206d ID A CKCFe cc) 0) 0 er 186 |S oC S101 Ae 47 PHB nes ie ac an dcyuneses aed 47 Cie) TEE) Wa 2) 6: a ee 206d missoa acutiocostata.,............... 139 (S]0( E5015) RO eee oe 139 1X00 0) | | 138 Rissoina alabamensis................. 26 browniana ......... Ah rma 139 SM STONE ii uindvesec sac seeoes 139 GEGUSSAtA i is osu he doe ines 139 Ble PamtiSSiia .. 06... 66s sek. dee 140 VES CECT AA Re 139 AGAMA T A ee) eels cee one «oe-en: 140 mochetoruiaminuta 2.......52..6.5. 20 eereilaChy PLOSPITa... ....5. sedeese 5. 162 S
Sabatia bathymophila............... 50
Sanguinolaria alouata 32........ 432 Santo Domingo Miocene........... 92 Saxicavella alabamensis ........ 21 Sary ella ie ay ibe ola Aes aayey ROU ie 56 SSCALALIAL eam ML sea hs tire ete ee eee 102 Scaphancdernsibrarius. 2. ysseseee: 49 MODES Ab itts nse Uitee ut ue laa 49 PUNCtOSERIAEUS 22005 .ece dees cnsiece 49 Wi SO MiMi setae rcv ct auube esc ts 49 Scaphella) costaricana...:...1.5...0. 187 COSTATICCHSIS WTS: seeseselouices : 272 MOMMA nereeeec cas land laa 81 Scopinelilaimoriereis! 7... 00) 251 Sconsia bocasensis 75............ 309 GOCIE AMA WTF cess cuessiciet 157 \2210 PABNIG ALA cere es 195,200; 201 Vaile. Pa WM eT Sisco ser as 308 ZOTIE SENG Masia tava st os en 2 193 Suplasvisatan (Maile si hegers es 200 DE SUCH Ala nkeatiet decease eee ree 158 FORMOSA Sees Levees woseseeees 120 UC OMAC Atte tele ra, Naat oooaca iat ss 121 MOnOCiNeUlatays.\.c..cassleckos: 120 RilS PLU OSA tees teh lesen 120 Henle adams Meet ae Mae aay ee 124 Semelevelaytoni 72)...19/) 196, 1425 Ne EU ie tak ean eG a IgI var. costaricensis 32....430 hamodoiiataa yet wats tose: 18 HAOMTOCHSIS ne cee eet eetes 19 Saya MPR Ae ate louise ects 430 SEmICASSISH Ay eeecse cance tend? 118 Seminellla Steatnsive ss. tesscee eae 94 Serpulorbis papulosa 75...196, 200, 317 SCUlPCURAUUS sch ceeteceeenser es 200 SIGAHECUSI Ne ence teuce.N sos aeemeean. 150 PALM MEUSIS A ye sesemeesindeseuce es 186 eotehge bob YEN ohUR oo WSU mrmea Myr wes 2 uy 150 (O20) Of DARDEN BARRE RRR ER Bass GA 186 Siliquania squamata,. o,.se.cc 133 IASVOTMETISIS. 2). 9.25. see se esse 188 MIGUEStA: he eee, 134, 188
306 BULLETIN 39 478
Siliquaria modesta, var. limo-
TROIS EG) WEA Gt LES canes rites 319 Simmnia’ intermedia, j.445 wise MAGe WAT pieata wy 4 we ceae ee NSE Sa m2 Sinum: maculatvwme~. see 150 SALUT ATOR eae e as. eae cree eres 150 PELSPeCHiviUtia soa. Nacecee I50 Siphonalia quadrilineata 2...... nel Siphonariaaltermata../\s- ee. 58 limeolatar Je AUN Ey Misses, 58 Siphonium nebulosum.. ..... quatts 133 Sistrum nodulosum 7}......... ESOL 190, 305 POSSUM es chen aanem eh gees IOI okenea planorbis. ates ees deci 140 eolatiellavceglees {incite ommaesaeen 157 Ala DAMIEN SISA sy) umudae toe 26 Aiba Bilis. 2. Waesae eaten eee 156 Varo Clavatas sya wee ST Wars: Rete oP ncdon sees 157 MSSOCOMONA. 3c) eee ee 157 Wi bricas by eee Were bea ee 157 SCabrnyscwlaye i none eee cee 156 Solarium gatuneuse.. seeises BLO SranMilacuia assis Leese sees Svs 320 peniseehiduin Ue meres 226 willarellonietog ce cme 200, 326 Solecurtus satunensisi a ee: 432 strigillatus.......>:s SE ie ere Bape 6: Solenosteira chiriquiensis 77...286 dallas bg Sig Diy Matin et poo A AD 285 var. medioamericano...200 Vallis leeiind 1 nae aed 200 var. medioamericana TL Nias Soh melons a acne ome none 285 Spitialis rostralis 20.720 aes 44 Spondylus americanus.............. 379 ChirgquienSisn) 23 ua ese ie 378 gumanomocon 24...... 194, 379 Strepsiduta heuloriniiae eee 25 Strigilla musanica 29...........:. 427 pistiormis.e was 194, 200, 426
pisiformis::.o aa ROO Strombina ambigua /7}........... 298 chiriquieusis Vos). gi eee 302 costaricensis, nga 300 cricamola \) 77 se ieeae a cals 299 gibberula, var. galvestonen- SIS. vic kg ae eee ay eee oI lessepiatia 232A) ehe a 298 matima °73 2 ae 299 var, musamica, aan, 301 sincola | (7342) jeeaene eee een Strombus -accipitrimties.s ae. 120 COStatus ccswasnue-caneeneneeeeeeee 120 gatunensis. “7007 2u L3G.) Bike pugiloides 76.)...137) 296,364 pugilis Ve pee “L1G, 26a) ers Styliola......-/lusius seek aneeeeee eae 43 virgula®cyuee. sie sie aiemtelale aeRee 43 syrnola smithi, 2a ele a Li Tagelus cebus: (220 cea 433 Tapes Sr US) 1. ..c0dsng eee Pi AZO Taranis pulchella).)iiaeesna eee w2 Tectarius ‘muricatuss..4, valde C37 Teinostoma cryptospira....... Bay) regularisy: 7auem wahetroay any so aelvache subangulata, var. smithi 7...11 Tellidora cristata.) Zou 189, 426 Tellina bellsiana. 2.03 ee 17 cercadica, is iia 194, ne 425 costaricama | 29:25 24... Sella 423 erystallina)) 2o\sen ee 189, I9QI, 194, 421 cynoglossa.......... as aprtehece taeees 17 dariena))) Zou ain PAE Bee +424 equiterminata..../.2ei eee 186 estellensis(!) 2.058 c ciae tail RA gatunensis: S74 See PN) Dearria ii cL ae ae a ff lepidota)|2o2 7. area 200, 423 NASA Zoe aver tee a alee) 1 arate ieee
— ee
479 CosTA RIcA MIOCENE
AON EIS) 9220 ee Pe aR 421 HRONMAATAM (0 duce l aU siaek ede cdeeds 424 Sich ay Cae ee PAA, Se\i6h) OF Ohi Ree ee ae TS BemmITOtunda | 2:1... 02... 2060: 18 MONA ZO 2. Joules. 205, 425 Nemmiaectzeon pertenuis 2......... 16
Perconanbenthalis. 000. 1.0....60. 189 var. boscasensis 7......213
d (sic) agate ey ae eae TO4) 211 bipartita | 2.....s23. 194, 199, 207 Wale DOCASCHSICN: 00.824. Ne) 189 CAMWOMMEMSIS 6653. .cce es ce sae's 60 CHU TISIGE |. "Gea SR mh a ON a 58 COUGARS aD et En 59 BOStaniGeHSIS 4.5). 65.02 L.dece:! 2A (OInS] OYE) 021 a Be 59 SSS RMA 2.150, co cla sees slot wos 212 PalVESLOMENSIS! ..... Le. serene 60 PALUIMEMSIS (9)! oc. ioe ete 208 BAUISAD ATA Ie Wie is ile dekiecse.' 210 HAHUETISIS | Zyadeess...c.2h: 194, 207 MARMANEEASIS le oil. sas 58 2b a4 2 CKO) a) LN a a ee ee 60 Mormialaim Vi ea as 60 MRE SCERS 28h ou i si.e alse lave coe e 2 60
Wats MIICIENSIS! 7..0...2): 211 IESG UNE AR Sn aOR A Meese 60 Pert eee des eid. eG 59 OOD 4 0c RE GAS ee Ba EA 59 Pimapriensis;: 4.) 600) ne es SPIOVCLIS, ge es ARRAS a MRO 59 AMG MMM NL sell Laake 58 SPUGINCT AN A Ze occ. css. % wie 194, 210 Smmcmlemena, is .. 80.0. De 207 (ESE a alae Pe 60 WiHOSa.../.,..... LE Na ee) 59 wolfgangi 4.............. 208, 209
Meredo Tiacens\: 2..........0.4.. Win 25
UE DEBS. 5 S10 [SY er 53 SOLOS A Aa ae 53
Miltaic GelrOidea,.............505+..-0c% 100
OLSSON 307
Gera S WOM as Wea hastens access 100
VAT! OTA As Cay Ween ores = 100 ALUM aN eeviseadtantes re serinc se +s 99 WHE OCORUSH Aue mes ssuet cs uanenels 163 Thracia astuimensis...5.J--..-<.2-- 187 PHISTAMA 2721 si voceeescimeenteces 383
A OUT AN OAC AN Ge sack ctenteagiees dese ecto 119 PEFCUBE Me Bessel ewincanigeuian ets 119
SOT Cla ee cee Seas ares Sa Riese 134 Torithavcanaliterac ye. (1.60 vos. 138 Gy MMGICAl ole ccseyee<esee-necea == 138 Tormatella: quercollis:), 4.300.025: 26 Tormatina, oviulitiay.s..0...02. PO et Pas, er COIX=lACrY MIA i320. s.teee snes: 206b Tralia denticulata...... BUR AE 57 PLUINUSCUILARY cess teenie dscns ee anes 57 WMYOSOUSs.sceer ch escnssevsesetcesnss 57 PUGH Leas setens awoscescisiwas sce 57 PEIPLICAtas Wir ismeesrede denn ede comet 57 TrigonOstmMa .......0..secesseeeeoee e249 74 Triphora‘abrupta...:.-.4.......--++:- 122 bigemma.......... umeleaWlrreeraetts 122 COLO. ose ea Uae, peepee 124 cylindrella..........-s6.0e2-++- ieee GECOrataee Meus eo oN 121 TANEOUS) ene ct ee mune cana 122
Pt VAS LOCK ae Uo venel 22
LTA GUE CA sci seoste ws peetaatraltas teas 122 perversa, var. nigrocincta...12! COLI CUA ne ee eeosn: ste rekeemees 122 CEISCHIGS! Meus oe aces any 121
AT lL as.tawuessins sadeees o- eres tnlsecarian: 44 PL RAO ao ueer teh trae tet ate enters ns 115 CALLEATA CI. yan douse Weeeres 117 arene llats susie ee ascent sales 99 PALVUS Cate ce deletes cena seer te 88 pulcWelliseiy. se eastee sco teo .116 SHOW AELIY neh ect ma mcnaie 26 sphaovVUbb TOUT HAM ngre saoee Usb aman? 305 SetMtO MICA erate ceelentshesmeeeins Hida 87 Trivia approximans......... ..-,.- 114
CAMCHG Ua ee ete ane 114
308 BULLETIN 39 480
Dbivia elobOsas (2. 22.l2.-c0-1 esse cess 114 OLOEIMAN SA MUNEU Vin eure nes Il4 pediculus. 76 so sean 113 PULA ee cee eee gee II4 spheerula...... Woe see netaseaedoe! 114 SUTRISa AWA tii cs One cee Baie Sea a 114
Trochus'agelutinants: 2.2\-7.0...5- 332 AULADILIS Sc sacddaees sae teases 156 Nenticwlartses sss dien eso 129
Trpuncatella: bilabiata.t.3....s20. se 142 CATINGCTISISC ilo. S..c. ade ete LAS puller tices nse eee ies 142 subeylinidrical: tio. cc.ere sess tA3
Durbotalipida.os.bccsnseweenss cess 153 CASTAHCUS: (lien csadtnosmeecur enon 152 Crenmulatus .1.25...cseemeemeecae ae. 153 INUEICACUISS cscs (Renee ae 137 pulchellal 2322.25 2kceeececesaee 152 Saltust Sore Gace ease e ae 333 Spetigleriamus 2.220 vasees cases 333
Turbonilla...... 1h eye eo einen eal 107 belathieca 24.5 cee aan 108 CEGTOS: cca saiacdates cae mentees IIo CHMEHSIS 7 Eee neeeee see et 9 GONPrad dhe ec ee tere 109 feu) atts IEDR Pea boraar ne nieeico ccc 109 S@XILIS een ocho h nae ee aes 108 INterCUpla eyes eee eee 108 ObDElISCUSi-.2- 222586 Se CR aa 109 PUMLICSA Co ice toss obscn ae ponaeeseee 109 TEtiCula bay. yet ee iat nse emenene 10g SME a ee esses eteoneeseceeeaveee 109 Sty lifOraiis) 2.7. wssseeeres ase ee ate e218 00) CALC INGCULES <8 ie. enc= tk Neen 9 textiliss- cw 3 sy. secagee cee 108 tuscabOmensis:< 70540-2224 eee 9 VALG Ae . Phe atetecsceunseae ete 109 Vinidanialy slay eee 108
‘Purriculalavitia se Weis lores eto 191 LavInOIdeS } 72. csecssc ec. TOL, 227
var. limionensis)) 7.052.; 228 tavirina |) Secocsses yasacewosweeclos 229
LEGIT Gata teas ye eae ereone cs 229
Turris albida 7...62, 190, 194, 199,
230 var. tellliea.: cs eee 62 bimoniatys” 07.027 eeeeeeremeeee 7 denticulata cee eens 6 moniliatus . 5) 7 ee eee 8 nodoideus. #27. 2a eee 5 SPeCCUS. \ Z.iiccpeenaseeeeeeee Aes av 6 Turritella acroporayyc. eee 134 alabamiensis, 74.) .)ss aie 25 altilira 775.0 188, 327, 322
var. chiriquiensis 77...322 var. costaricensis 77...323
chipolana:; ..:.1072 cb eee 135 Couradi, «605035524. eae ee 320 exoletass..)))0 epee eee 134, 188 var. limonensis 77.....%24 PadDi......cacrtene nen eee eee 321 gatulensis | 77.07 eee 3,20 humerosa. ...))80. Site eee ee 25 var. limonensis,..J. 188 Mimetes 77.2) eens B20 mortoni.../ 0. c.see eee 25 oréodoxa if 7 \yis.uy ee 324 satilla ....4...)4 28 Gee 135 subgrundiferay yee eee ras terebriforniis) 72 ee r25 tennesseensis;...\issuceiee see 25 tornata... .).c.neseeasenpnen ae eee Bas variegata-.... yess 134 yucatecana..... 0 a aeeeeee eerie 134 Typhis alatus® 73.)..2 spose 195, 304 gabbl.........:.¢20-ssee Meee 186 linguliferus, var. costarien- SiS 3.420) RE eee 304 U Ultimus ie..0. 22h ee eee 1¥2 Umboniunm batrdia. 226 (eee 158 Urosalpinx, cinereus... OF, 282 florida. 0). 2..s.ceee eee 98 perrugatus.:. oan ress eeee 97 tam PeeMSIS. 5.0022. oreeee een epee 97
48x CosTA RicA M1ocENE—OLSSON 309
WOES | ns 48 V Warimellajchipolana’........ 2.2.6... 43 ‘yD SHURE i eee ee 58
Venericardia conradiana, var. ISHOMENSIS 29....c.2.366 3590 scabricostata 3?5...194, 200, 388 GETEP ALA 388 POO REMI SIS ellos, se diegeausecees 22 Reig DVamCIANA. 0. uses ccc cesses Ate | SIDE CSA En ena 200 He\eVC DU 400 he re es aa A406 UBS: CUO TCCE he oA ea 412 MM ICOStata.: sc... cscs sees AII PMAOSA!T....1....5 ESLER Sit CASCIO 4I2 Venmetus papulosus....72........... Bu 7 PICT A elses wdacisises's<ocee os 200 SCMAPEMFALUS, 15 6o.i ccc. ewe 05 2 318 Vermiculania nigricans............ 133 CARY le 8 aN a ee Wermiculis spirata........00...2.... 132 Weronieella tloridana................. 58 Watuvelila hemiplillt.., 0.6... es. 159 Me tIGAMEss CN Nec weed ce ee 159 EOFS 0 ge 159 TAMUNELCA TAMIA LAN Ji Shee dod): 161 Wolmta altaroi: £7..:...........187, 271
GP AIAN ner nea a cee arama ake, 55 GU Dian be eta enon send eiealamels 81 VALESCOMSS sescebiesansiesmacencnases SI Volutilithes Wim@psisy.cvcee ask seen 25 PU SACU Sh. uciec he ut helslnnes Meine apts 25 AVIOMMILS THe Mae ireciivat de naeielskteses noes 80 NOUV AIAN cei see eens bate ene a 48 NOMVaniae i iets UN EN den SO Volvula cylindrica.....190, 194, 199, 206c Volvulellasmicratractas)..0 52. 186 WwW Wailliamiivlerebsiie ect ce eee. 58 X IMATICUS /SCONVMUSIY, sees cenceteoe et. 284 SCOPUIUIS Hy 7 ices cose ieee toe 283 Xenophora agglutinans............146 CAPIDELA sacs dian a dwoasesaees 136 conchyliophora:...i.023.: 145, 332 Y Woldia; bocasensis 37.0. sans. 348 CDOPRCAT CA tee es hccers tae 25 Z PAZYAPUATUS iis dons aeestehs weltenre ema nee 154
ADDENDA ET CORRIGENDA
NovtTe.—-All page references are to the Volume and not the separate
188 206a 275
405
Bulletins.
Bor RNIGICULA read RINGICULA.
For Siphonaria lineolata d’Orb. (reme preccct fied ) substitute S. xaufraga Stearns.
For Oliva circinata Marrat substitute O. sayana Ra- venali(Cat. plo. tesa) and place) e7czaia in synonymy.
Kor Marginella apicinca read apicina.
For Scaphella junonia substitute AMaculopeplum junon- tum, this species being the type of Maculopeplum Dall, 1906.
For Epztomium read Eprtonium.
Janthina communts is regarded by Johnson (Nautilus,
1922) asidentical with /. jazthina. Bartsch, how- ever, considered communis and fragzlis (equals Janina) to be distinct (Bull U. S. N. M. 1915). The latter opinion is followed here.
For Petrotrochus read Perotrochus.
For Chione Rowell read ChtoneRow/eet.
iter Acicon costavicenszs, n. sp. insert Pl. 15, fig, 15.
For fuses nitocosmius read fusus miocosmtus.
mauiten Buw7sa crassa, tor Pl. 15, ig 15. read Pl..15 py. 19,
Roni, ip. Wearead. Hu'C. Tea:
For Gafrarium limonensts read Gafraritum limonense.
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