PPA ey IS ETT Err any Cain whee eed aA de we ne te itapeged A dagae eA tis Shae Deir etouets so dae vrai . oa ae) terrane ead Peet Bote ta us? Bice oe led neds ba 4 Pattee diy bye alk Pod Sess ae At hoahia irae he Tee vere rare ae Seay 4 aia nseiey rs Pe Rn ny ee Ams 8 hea HIRE be we ‘ate phot on Ripe) tet tore & ay rantras st ete ta a mon fearsome) att trl enn Pinas peter dy myn teh Rene Va tanee Vite cassie error Ty y ay es acptes eerste silytc tthe att a eset thre Peete te acetate Souk aa eee everest Peal WiAaah ia iennan®. : Ah NY tn ds yaaa eat dy hee +4 elesael of sees Tete a aes mar Hah + ee) ih ata ey es a via aah ee et ma Con z eichaen nbs phi na Sate oe 0 Ades, oP Dm Oni tae oua® absentee Cad wee erst: see a Paani ene cei pares ae nih gas ahead Ne aaa eer near oe be ee pean wa Se ee ey bb % BAA OF arAd. ‘sata see Boace nee haa! A Be Por wer raw opre gear no fe ina sone A a Sy ne + a ee oe a FO hanes Oe beast» ae. Pt se saad naceahecAcktihe sntenes, Pee reread 6 6 Saphcherde meeeste Reirreperners lata) 2 dw Bi pon mane cet tatectecete -b ebenebene oaane sty a 1D 8 Ae Oe 2m ae ese AR me Be wel ree tom Terry ores mene We Be «BEF reget p a oe ee eae mn Saba acorn aes he n ciaeaponsmransvipesmerssasee preeeeee aon 2) me ee mat oa naga ome et a She nde me wer BS abe ee tges, at Cnn Pilieleenetiel = athe itgme. SS esuebers SraNtgante! gamma Rie alene te ngteme se cae licateos Riedy ariel sttsa sted ater Pit shay Saetalbs bereins “har ere ee eae fat Ee Agha nae on Lahage fs te ee Ree See EE ot a EH ; 2072) conte a ELE Sees 8t ob nae ein RE eet OR Tee a ee Sst PIES fitout ry eae Phe ages ieee ile Sqtgnaigtg ee A fi erntetrte ete Ste sade 23 ieee’ gertet rsh meet paw eer, OS ge et piped ans ang rege tee Aer re gy mgere eae teaey Bethe eters se sre bety oepe sates ele igure gis enorme eaae Dictate SE gecte ag ermgten ine ene eee es frie Sy he ate rte shan tee bes “x it SUE aia B ohigl icap cet Se Pes Se ee aa Fe yen eene oe u alae abes, Retr acists wri cava vere aia peam reevsngee reg tat i ¢ Deel te soe peers tetavbaeese poo eepeeeeaage orion Pei eneat eet b 1 sega : Heri ide prwanoeetavgr seeeyogestar=* jae toe reecaeudesuwereretecnver ome ag bate ap ay) ta oats oy: aa Pu a tpeenetyeeenre renee eres ene va wer w ee ee eae _ eae era no wb cptemaera ree aren Pant as Cees Fach sesss Bree a rarer + ¥ vee etter sas Nuintayarnea ee erty RSD pewv ee aeigeree aber ere Riidgtneh lin ct) HARVARD UNIVERSITY e Library of the Museum of Comparative Zoology VOLUME 98, NUMBER 334 JUNE 6, 1990 Neogene Paleontology in the northern Dominican Republic 10. The Family Cancellariidae (Mollusca: Gastropoda) by Peter Jung and Richard E. Petit Paleontological Research Institution 1259 Trumansburg Road : Ithaca, New York, 14850 U.S.A. PALEONTOLOGICAL RESEARCH INSTITUTION Officers PRESIDENT) occ ct 8 cigs Sian oe ee en ee cee rene JAMES E. SORAUF VIGE=PRESIDENT, 25 3c sce ee roe HARRY A. LEFFINGWELL SEGRETARY 5.) oid. os Sees ho ee Se es eee HENRY W. THEISEN TREASURER? (0 5 tkcce tte Be a ee eee ae JAMES C. SHOWACRE ASSISTANT DREASURER) © ease) cece Cenc JOHN L. CISNE DIRECTOR So Societe) hac ee eee eee PETER R. HOOVER LEGAL GCOUNSED) 25 sci Cre OTS eee tee es HENRY W. THEISEN Trustees Bruce M. BELL (to 6/30/90) CATHRYN NEWTON (to 6/30/91) CARLTON E. BRETT (to 6/30/92) EDWARD B. Picou, JR. (to 6/30/92) RICHARD E. ByRrD (to 6/30/92) JAMES C. SHOWACRE (to 6/30/90) JOHN L. CISNE (to 6/30/91) JAMES E. SORAUF (to 6/30/91) J. THOMAS DuTRO, JR. (to 6/30/90) HENRY W. THEISEN (to 6/30/92) HARRY A. 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Petit Paleontological Research Institution 1259 Trumansburg Road Ithaca, New York, 14850 U.S.A. Library of Congress Card Number: 90-61317 Printed in the United States of America Allen Press, Inc. Lawrence, KS 66044 U.S.A. CONTENTS PATSSLLAC CRMC IC TTS PI Ter Roe rererenerar cca areas a lea cred sie Vie peeeaemeeaTs te ce meyers eve. i.e epay ay otmbep ayer sha.'s nn chesarelny oa] foi ohare yeteldvete sha yegeieyaons FRE SETNI CL PRCT RE TTI ITT Tee Mer ct Tees ect Rete roe Cun eel eres tete ce vest r ere A ree teva ts occ cus sin anteesoceyay ey photon naisuNloPay abe loops tha el shal Girn ov exe oU dgske hitiASHOM o oS oad owSRS wowed dona bob oOo od ducoDs CODD O sO Dotod dad deer orn noise aMEEeameasos Sen boo cnn on aonheEnoEon Der a NT MOMIGASACNG 's acide Ga edon debieddd ds oodeatic SUAw Eom ORM aocoBa mrad cHUenEBnaeesc niEadooopecamooomodaeocdnne Setene meteors ar TOS ICatipra DV pment er ttre eae er La verses crlscrcieeisiieresiys sot er = oicterspare: cists sre aiencel ava letninve siols bausieneral cetera eset San ard fondue orale ote Ope ORTApHyanGubaleOecOlOe yar se ec iaet stealer elaicevereters sferereyalere eee eledeete teeters ele etait tele) ctetiel=tn(e) )afeteiate|=l/atmlaie steln/=!=(a\- lpiats ASI EOTETONS OFCOM CH? INATMAONG soocoedccosudaulc onpdnaconn soso gdeéou cMuooDOdEdooe sean oecuSancoassgEccuonte Systematic Paleontology TiayiRaya Venn). dares tadol teow adleb o UU NE crock GUS giao Uaioe aioe Sern rca ian ee aiciin sea ara cin Gn oan GOeinn Gonna acm erin erecrimice Farvity Canedinniche Roressemal latins NOS os ocoeocduasconnmdueodds se soooncHconsonseovesenemsrs snesebseocsdssaudorcoee Gane Canaglaia tannins NO eee oes oe Rees one us b pad ont oo ebb ado conan poor eae nr uuon aR ocnds baos cn aen ae qansmc sean one Suissa Gace vdcaccp dois sees aaseooud as Sona Ss HoDsae sneSEe dononbn osanesau Baap onnaedconsocuUned Sone armod LEAs’ Cangdliznin (COGAERD) GAT Calley IBIS scocacs poco cocseonaroavonddocgand donpoogoHDoNEdASesWAD oSoodEa SeoEUdOoSS (Copealicnin (Calin Taine ORO, WO oo cascousoccce podudodcHos codebmaoeonad auonnnocosunoeoonnenaosepenoos Gancellarial(@ancellaria)junclasnesp See eee ee ee een noe Gee eee oe ieee eee eee ier ieee eee reer (Cirgea lenin (Came tienia naa erty, WONT socasescosossooswadaon oaageoonoaacoandecsoohyasbooddcnDH EUs eDSoHaNSSoL (Ceodichia (Career) (OCG DAN, WSO6: a coosaacoosuasacnsoodcesosenes saubdounsodnooonHbeoDDHoCooDD dees undoone Shizenre Arvada Oe, WORD. ocacccscccndscoccuscosoune seconde ous oosedonounsbosrESenesuaseboUurCcosbosoocudmedoce Caigcalienia CBT GHLY) TNE TL. Sthocwacccccscors pesado peasHus SoC oCeabo SAnabooEHoCODOnDUboUS EEF oDOSAdsCSDOmODDEe Shipanns Sma UOUEEEATIINA, WOOT ccdroossocoodduooo on ddedonucasbu spe ndospodaoneDHaDaduoodooOUSoUGeUENHoOC eran OOOe Cereal ne (SHED) TAG CA TIS, Ws TOs 090050 6a 00 os onde conic ODD OMGOEODEE SoA ENKeOoSDodoHOdaHeCObEDUHOOEsAddODoE SUE FRIUS JAGAN T7, NONE 2 os naan oonbn coos con codanesbenseasnueubs ceed enh oo dod doddoesodardoesanaopounnoedDacr Cancellaria (Bivetiella) gabbiana Pilsbry and Johnson, 1917............. 222.2020 cece eee es Gancellarial(Biveriella)vepistomsjerai Guppyanl Si Ole eercee cece nies cee acre ade eee tia eee (Gancellarial (BivetrellaioajOnensissnysp set eerie ec ce eee eee eee eee erect er erie eee eer SUID ERS JIN TODA OURS IO aoa onpooonguccoondddonsaoncconnDocoss so OooSeEEcoONEMaaDOAAmoqneMEnsoceD SAnctas Ciel icine WANGQ ISD ABATE, Ts Fy 540ganc8 caaccssdassoegsuon seh 4 deobouebodcodecaodsos oon camteaesacsosEEsaDseSS SHipenis ea Bia NEWS, MAD) 2 ssondosedccoondsoumonsevbneodesoasnoosEecuogeauenooeusHaososeDDUCdsoDOGesauEOdENE (Gancellarial(ertlernia)Vrriirandaanesp ee Peer ee en ee te eee eae ee ore eines siarereie eet severe eve shenetclie sarees Ire SubgenuseMfassylasAdamsyand Adams lS SA gyal crete ele ce ret ateie eye yey steteete ara ake eco er Vel oner ee ore tes a Ve tee Fal stsneiifof=\ole nf) ara) et stele taal (GancellarialMassyla)lopezanaanesp PEE TEE Eo eee ere nee nee hee er eee eee ee Cree eee eae een eer GenusrA pheravAGarmsrand vA Garnissa lS pes seyret arcane eects arte evened eve tel el baie ele veneer neteeelin otehcke stretch otetcdne ofemesT oe AVA OAD (MEM a MOU nroocanc cob oanocouus oncopeuedosns odnonasdcecsdoddsuosonadud To dcoccondodag2aged a soe Gane /Aailieciita DAL een ode sousaescesousdoporespudoudo tee cosh oceo saan taapEpe sou novooD sD noomuuodapaUodn acon RAAT Ny Wo blade ced bn Go OU aos OAltGe Hood GORE soe ACES nates On nuL QgUnoOh sb roe Duco cnavo peace (Genuseiriconostorma Blainvillerel 8257 vera acters svetet ates svelte ehctsercave ceeerer een sestere cules elec ole evel eased elisuel= lined ele\ Se) olay a= iotein ies =i+Ler tar = Chien aaniira yomeseanines Mi sacc¢ naan ge.coumecsasagduRsdcoobn osssebomocanascncsuncunnpbedosDaqaaooopAbAg ces Mrizonostomal (Ventrilia))eurabis\ (NAaUrys iL 917) ee oeiereneter sire cpetetetere re ere heer tote ete = ee eee ote eee Nett ated erate =e Erigonostoma (Ventrilia ?) insulare (Pilsbry and Johnson, 1917) .......--... 2-2. .e ese eee eee eee eee eee Gong bea AIGA RAE hone aeineedes sae dacbn odor ne uC On ear iden acon PuCeb OSA aan NO arabs OomnSoSS CORCoME One R OW eE Oe ABA AAEM Gh penbanocsontue due coeObeR co OCD SUN OOD ade cao D HO ASaaes SHAS baGonb addonoe onc epOr eacess meet Gaieuilgiigee Rane NOS ce secot ng coppacaeoodHDOdUODBOe Teo erooadosen ocopo cH DE SueOS ona nan aenaSsacuenadacsDmncscS oc Ligatinee lao paees (IN Eatinis ION) acoossacgoonbd doce ee eden ecoces eee DDE Eco o00or pocaboc nse ncuoonuooomdnoSoob Seco (Gie AVG Pieacoenin, ICs sass ecnp opto cos onan eaoos de gooeoeToSaEe oc ran ceusnbenednosGcUpss Goonasocostcocms conc ACH RATE FAG AGE TOO a ooee be cop Sa pu eeeobe beens UNO BUDO OSC Od ona Ren U MSD ADS Rpoatoos BbtaD poUC cease ceonR scones INTISSIGNE na dbi oo SO o bent eos UIaO a Deen a OOS On RUM anaaS aman ontacds Anaee a sae OMA doa SNE DIUe ro So oue cre tonaceena RGIS (CHG <5. esd sopeeha SEOs ob SB EABoEeeD HUOU oS Uadee LOTEE DCE Ace ene Saar OAR OMdcne Ant DRO Hoodoo Cdc Oo moses rae LIST OF ILLUSTRATIONS Text-figure Page 1. Index map showing location of investigated areas in the Cibao Valley, Dominican Republic. ....................000 0002000: 88 2. Columnar section of Rio Cana showing (discontinuous) “ranges” of cancellariid species. .................. foldout inside back cover 3. Columnar section of Rio Gurabo showing (discontinuous) “ranges” of cancellariid species. ................ foldout inside back cover 4. Columnar section of cliff exposures on Rio Yaque del Norte near Lopez, north of Baitoa, showing (discontinuous) “ranges” of cancellariid species and stratigraphic position of NMB localities. ........ 2.2.2.2... cece cece eee eee eee foldout inside back cover — 5. Section at the mouth of Arroyo Bajon on Rio Mao showing (discontinuous) “ranges” of cancellariid species and stratigraphic | position OF NMBJOCAaliGies. <5 2.:fce-cscis oie) vest ave tie) eh Shes en nsniresh fone rap eye, 006s foueier eh ou ere setts) S/o SvSIGh Ley a: Fans ey seetetene iste Lope tei saeS ee ee 89 6. Section exposed in Maury’s Bluff 2 on Rio Mao showing (discontinuous) “ranges” of cancellariid species and stratigraphic position OFINMB localities:. ©. n\.12)5.2).-0svelevere.chatancve cai sits cre foezetelicae ra vove OMuci evereuaye ete sare cereus te, Seep Stee vakore rave ieee eee RESTS OC kere 89 7. Section exposed at the downstream (eastern) end of Maury’s Bluff 3 on Rio Mao showing (discontinuous) “‘ranges” of cancellariid species and’ stratigraphic positions of NMBiand TU) localitiess 2 oe arc ciee cece ioe eiericrio ce ee inicio iaeisiee eerie retains 90 8. Schematic column for the central portion of Rio Amina showing (discontinuous) “ranges” of cancellariid species and relative stratigraphic positions of NMB and! TW) localities: .0....5c).-eceicises ores cee. cepepeuei ct ere sere 8 aint cueratey eects esegeiiner she oleosi Ta. ees eet ee ee 90 9. Schematic column for Rio Yaque del Norte showing (discontinuous) “‘ranges”’ of cancellariid species and relative stratigraphic positions ‘Of; NMBTOcalities.. zc. savers Sucre cececeleret Soke g «ofa erakers (edcr ere eens chek So sual.n HO ses fope vel a aise ee Senn os erees ee ae 90 10. (Restored) height/width diagram of Cancellaria (Cancellaria) guppyi Gabb. ... enn eee 95 11. (Restored) jheight/width diagram’ of Gancellaria (Gancellaria)junctaSDASp: ss eae = cies «eee celles eeieieieiel | Seiler 96 12. (Restored) height/width diagram of Cancellaria (Cancellaria) harrisi Maury. ...... 2.2... 2.00 e ccc ence eee eee eee eee eeees 97 13. (Restored) height/width diagram of Cancellaria (Cancellaria) rowelli Dall. .... 2... 0600000 99 14. (Restored) height/width diagram of Cancellaria (Bivetiella) epistomifera Guppy. ..... 2.2.66... e cece cc ee eee eee eee 104 15. (Restored) height/width diagram of Cancellaria (Bivetiella) bajonensis, n. Sp... eee eee eee 106 16. (Restored) height/width diagram of Cancellaria (Hertleinia) miranda, nN. Sp. ... 2... 6.6.62 eee eee eee 108 17. (Restored) height/width diagram of Aphera islacolonis (Maury): 223-22 <2 <= sees © ee see oe nee aes arae eee 110 18. (Restored) height/width diagram of Agatrix losquemadica (Maury). ........- 00 ccc cece cee nee ee tenet eee eee ete neeee 115 LIST OF TABLES Table Page 1. Numbers of specimens of the nineteen species of cancellariid gastropods collected in the Cibao Valley for this study. .......... 93 NEOGENE PALEONTOLOGY IN THE NORTHERN DOMINICAN REPUBLIC 10. The Family Cancellariidae (Mollusca: Gastropoda) by PETER JUNG! AND RICHARD E. PETIT? ABSTRACT Twenty species of Cancellariidae belonging to seven genera are described and figured. Their mode of occurrence and their stratigraphic ranges are discussed. Trigonostoma insulare is the only species described from the Dominican Republic which is not represented in the extensive collections studied and may not occur in the Neogene of the northern Dominican Republic. The following 10 species are described as new: Cancellaria (Cancellaria) juncta, Cancellaria (Pyruclia ?) uva, Cancellaria (Sveltia) inquilinus, Cancellaria (Bivetiella) bajonensis, Cancellaria (Bivetopsia) plectilis, Cancellaria (Hertleinia) miranda, Cancellaria (Massyla) lopezana, Perplicaria canae, Axelella emblema, and Admetula zalayana. The genus Cancellaria is represented by 13 species, which are assigned to seven subgenera. The genus Trigonostoma is represented by two species (including the enigmatic T. insulare), and the remaining five genera (Aphera, Perplicaria, Axelella, Agatrix, and Admetula) by a single species each. In terms of material, Aphera islacolonis is the most abundant species. Of the 19 species present in the collections under study only one is definitely known to occur outside of the Dominican Republic, and three others have questionably been recorded from other localities. No interpretation of this high degree of endemism is possible at this time. RESUMEN Veinte especies de Cancellariidae pertenecientes a siete géneros son descritas y figuradas. Su ordenamiento estratigrafico y sus modos de ocurrencia son discutidos. Trigonostoma insulare es la Unica especie que no esta representada en las amplias colecciones estudiadas y puede que no encuentre en el Nedgeno de la Republica Dominicana septentrional. Las siguientes 10 especies son descritas como nuevas: Cancellaria (Cancellaria) juncta, Cancellaria (Pyruclia ?) uva, Cancellaria (Sveltia) inquilinus, Cancellaria (Bivetiella) bajonensis, Cancellaria (Bivetopsia) plectilis, Cancellaria (Hertleinia) miranda, Cancellaria (Massyla) lopezana, Per- plicaria canae, Axelella emblema, y Admetula zalayana. El genéro Cancellaria esta representado por 13 especies, las cuales son asignadas a siete subgéneros. El genéro Trigonostoma esta representado por dos especies (incluyendo la enigmatica 7. insulare), y los restantes cinco géneros (Aphera, Perplicaria, Axelella, Agatrix, y Admetula) por una unica especie cada uno. En términos de material, Aphera islacolonis es la especie mas abundante. De las 19 especies presentes en las colecciones bajo estudio, solo una sola se conoce definitivamente representada fuera de la Republica Dominicana, y la ocurrencia de tres otras es cuestionable en otras localidades. Una interpretacion de tal alto grado de endemismo no es posible por estos momentos. INTRODUCTION This paper continues the series of taxonomic studies dealing with Neogene fossils from sections situated in the Cibao Valley, northern Dominican Republic (Text- fig. 1). The project within which these studies are being carried out has been outlined by Saunders et a/. (1982) and Saunders, Jung, and Biju-Duval (1986). Some comments concerning early collections of molluscs from this area have been given by Jung (1986, p. 5). We wish to state that the excessive number of ref- erences to previously published material which makes the text difficult to follow in some instances, and the excessive number of tables, were mandated by editorial dictate as is the following statement of authorship. This paper is the result of a joint effort in which both authors shared responsibility. The portions dealing with geology and stratigraphy are primarily the work of the senior author and the systematic portion is primarily ' Naturhistorisches Museum, Augustinergasse 2, CH-4051 Basel, SWITZERLAND. ? 806 St. Charles Road, North Myrtle Beach, SC 29582, U.S. A. the work of the junior author. However, both authors contributed to all sections and this is a true joint effort. ACKNOWLEDGMENTS The material on which this paper is based was col- lected during field work carried out in the years 1978, 1979, and 1980 as part of the project referred to above. The field work was made possible by a grant from the Swiss National Science Foundation (Grant 2.646-0.76). The financial help and the assistance in the field pro- vided by Institut Francais du Pétrole are gratefully acknowledged. We are indebted to the following persons for the loan of specimens under their care: Dr. Emily H. Vokes, Tulane University; Dr. Peter R. Hoover, Paleontolog- ical Research Institution; Ms. Jann Thompson and Mr. F. J. Collier, United States National Museum of Nat- ural History, Smithsonian Institution; Mr. C. P. Nut- tall, British Museum (Natural History); Dr. Robert Robertson and Ms. Elena Benamy, Academy of Nat- ural Sciences of Philadelphia; and Mrs. J. S. Lawless, Peabody Museum, Yale University. The manuscript 88 BULLETIN 334 was critically read by Dr. Matthew J. James, Sonoma State University and Dr. Gary Rosenberg, Academy of Natural Sciences of Philadelphia, and we are ap- preciative of their suggestions. In addition, we are es- pecially grateful to Mr. Wolfgang Suter, photographer at the Naturhistorisches Museum Basel as well as to Dr. Richard Guggenheim and Mr. Marcel Diggelin, both of the Scanning Electron Microscope Laboratory, University of Basel, Switzerland. BIOSTRATIGRAPHY The Cancellariidae are represented in the Neogene of the Cibao Valley by 19 species. These species occur in the areas shown in Text-figure 1 except that no cancellariids occur in Area 8 (Arroyo Punal). The “ranges” of the various species in the different sections are shown in Text-figures 2 to 9. The word “ranges” is put in quotation marks because the term is misleading. The species concerned do not, of course, occur continuously through a given sequence of sedi- ments. In fact, the occurrences are spotty. This is par- ticularly evident in the text-figures showing sections comprising a thick sedimentary sequence such as those of the Rio Cana, Rio Gurabo, and the Lopez section on Rio Yaque del Norte. The distributional patterns of the species discussed in this paper reflect a considerable degree of strati- graphic restriction. Twelve species (more than 60% of the cancellariid fauna) are restricted to single sections. This large percentage is surprising considering that the S RODRIGUEZ f sediments of all the sections in which cancellariid species occur are of similar age (late Miocene to early | Pliocene) with the exception only of the Lopez section — which is of late early to early middle Miocene age © (Saunders, Jung, and Biju-Duval, 1986, p. 30). Of the — 12 species restricted to single sections, four are known from only one locality each: Cancellaria (Pyruclia ?) uva, n. sp. is recorded only from locality NMB 17275: Arroyo Lopez on Rio Yaque del Norte (Saunders, Jung, — and Biju-Duval, 1986, text-fig. 26); Perplicaria canae, n. sp. has been found only at locality TU 1230: Cercado Formation (late Miocene) of the section on Rio Cana | (Saunders, Jung, and Biju-Duval, 1986, text-fig. 15); _ and Cancellaria (Bivetiella) gabbiana Pilsbry and Johnson, 1917 and Axelella emblema, n. sp. are known only from localities NMB 16938 and NMB 16942, respectively: both situated in the Baitoa Formation (late early to early middle Miocene) of the Lopez sec- tion on Rio Yaque del Norte (Saunders, Jung, and Biju- Duval, 1986, text-figs. 21, 25). The remaining eight species which are restricted to single sections are distributed as follows: Section on Rio Mao: Cancellaria (Cancellaria) maury- ae Olsson, 1922 is known from localities at Arroyo Bajon and Bluff 3 of Maury, whereas Cancellaria (Bivetiella) bajonensis, n. sp. has been found at lo- calities at Arroyo Bajon as well as at Bluffs 1, 2, and 3 of Maury (Saunders, Jung, and Biju-Duval, 1986, text-figs. 29, 30). ie} 10 20kmM 4 Rio Cana 2 Rio Gurabo 3 Rio Mao 4 Rio Amina 5 Canado Zalaya 6 Rio Yaque del Norte 7 City of Santiago 8 Arroyo Punal 9 Rio Verde =a] Upper Cenozoic f=) Oligocene - Early Miocene ? Eee] Mesozoic FERC BalT04 Sev gas Y 26 UNM as, PaO No OfP Oo aN. : ae Text-figure 1.—Index map showing location of investigated areas in the Cibao Valley, Dominican Republic (after Jung, 1986, text-fig. 1). DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 89 py, 1876, which occurs in beds exposed along the Rio Cana, Rio Gurabo, Rio Mao, Rio Amina, Rio Yaque del Norte, and Rio Verde; and Aphera islacolonis Section on Rio Gurabo: Cancellaria (Cancellaria) juncta, n. sp. and Cancellaria (Hertleinia) miranda, n. sp. are recorded from a number of localities within the late Miocene Cercado Formation; 7rigonostoma (Ventrilia) gurabis (Maury, 1917) is represented from two localities in the late Miocene part of the Gurabo Formation. Lopez section on Rio Yaque del Norte (late early to early middle Miocene Baitoa Formation): Cancel- laria (Cancellaria) rowelli Dall, 1896 has been found in many levels of the section, whereas Cancellaria (Massyla) lopezana, n. sp. is recorded from only two horizons. In the section exposed along Rio Cana, Cancellaria (Bivetopsia) plectilis, n. sp. is known from a single ho- rizon in the early Pliocene part of the Gurabo For- mation, but the species is also recorded from a nearby locality (TU 1422), the age of which has not been de- termined. In addition to the 12 species with restricted distri- bution as mentioned above, there are seven species which occur in several sections. The most widespread of these are Cancellaria (Bivetiella) epistomifera Gup- Young gravel terrace 2 Cc 2 20 28 on 2% fo fe ives oc wo Oxo ob es Od 16928 @ 8 —— 16918@ 16924@ 169250 > =| ——_16926@ 171778171788 3 ro) ee 16915@ 16923 @ 16927@ | | | | | | | | | | —y 169220 | | [o__—_—_—_—_~—~ 1m Text-figure 5.—Section at the mouth of Arroyo Bajon on Rio Mao showing (discontinuous) “ranges” of cancellariid species and strati- graphic position of NMB localities; # = NMB localities collected for microfossils and lithologic analyses; @ = NMB localities collected for macrofossils (after Saunders, Jung, and Biju-Duval, 1986, text- fig. 32). (Maury, 1917) which is recorded from the sections on Young gravel terrace ae = ae) ie) cS) & at a o LE Qa Oo a 2 =j {ok mn oO Clayey silt (Oy (©) ?TU1220 ee a CANADA DE MERA Sandy, clayey silt 16955m' 169548 L Sandy silt 16806 @ 16953 POTRERO DAM 20 16949 @ TU : 1218 16804 @ 16950 B10 Sandy silt Text-figure 8.—Schematic column for the central portion of Rio na showing (discontinuous) “ranges” of cancellariid species and relative stratigraphic positions of NMB and TU localities: # = NMB localities collected for microfossils and lithologic analyses; ¢ = NMB localities collected for macrofossils (after Saunders, Jung, and Biju- Duval, 1986, text-fig. 35). Rio Cana, Rio Gurabo, Rio Mao, Arroyo Zalaya, Rio Yaque del Norte, and Rio Verde. The sections of Text-figures 2 through 9 cover all the occurrences of cancellariid species except for three small areas: Bluff 1 of Maury on Rio Mao, Arroyo Zalaya, and Rio Verde (Saunders, Jung, and Biju-Du- val, 1986, text-figures 29, 36, 38). 17196 @ —| === | SANTIAGO Silts and clays 17315 a — [=== Clay with pteropods 2 1, ge 2 gig ARROYO HONDO cee a Silt with coral debris 1 = 2 ‘o — 5 wv | sg &Se 17268 @—p! 72068 | ; See 17305 \ ° Oe EEN = LA BARRANCA | | | 17267 @ —> =) Shelly sil 17302 «“/77 elly silts 17266 @——»! 7298 17299 17301 @ | | 17297 LA BOCA ca. base 17296 —=( Silt 4—PLIOCENE 17295 (| == 17294 = “ANGOSTURA GORGE” L Mainly limestone o > =) p ARROYO LOPEZ oO ae oon) 0% Shelly conglomerate = | LOPEZ SECTION = Shelly silts ete. -, = Aes Text-fig. 4 ag eS peeo Predominantly conglomerates e a 16909, 17168 17169 @ BAITOA CLIFF 17170 @ Predominantly 17328 @ conglomerates with silts + thin conglomerates at top 200 1004 m 17161 to 3 0 ines") Fomesa Text-figure 9.—Schematic column for Rio Yaque del Norte show- ing (discontinuous) “ranges” of cancellariid species and relative stratigraphic positions of NMB localities: # = NMB localities col- lected for microfossils and lithologic analyses; @ = NMB localities collected for macrofossils (after Saunders, Jung, and Biju-Duval, 1986, text-fig. 24). DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 9) The following three species occur at Bluff 1 of Maury on Rio Mao: Cancellaria (Cancellaria) guppyi Gabb, 1873, C. (B.) epistomifera, and C. (B.) bajonensis. Ac- cording to Saunders, Jung, and Biju-Duval (1986, p. 32), the age of the beds exposed at Bluff 1 is late Mio- cene. Along Arroyo Zalaya Cancellaria (Sveltia) inquili- nus,n.sp., Aphera islacolonis, and Admetula zalayana, n. sp. have been found. The age of the beds is early Pliocene (Saunders, Jung, and Biju-Duval, 1986, p. 34). Six species are recorded from Rio Verde, all from locality TU 1250 (for location see Saunders, Jung, and Biju-Duval, 1986, text-fig. 38): Cancellaria (Cancel- laria) harrisi Maury, 1917, C. (S.) inquilinus, C. (B.) epistomifera, Aphera islacolonis, Agatrix losquemadica (Maury, 1917), and Admetula zalayana. The age of the beds at this locality is N. 18, early Pliocene, according to Akers (in Vokes, 1989). Only four of the 19 species present in the collections under study have been reported from localities outside of the Dominican Republic, and three of these reports are based on questionable material as shown in our discussions. In the absence of critical monographs on the cancellariid fauna of the Caribbean area, no de- termination is possible as to the uniqueness of this endemism. Our opinion, based on our knowledge of the cancellariid fauna of the area, is that this endemism is not particularly unusual and that this family will prove to be of considerable importance in stratigraphy. Monographs on other families, based on these same Dominican Republic collections, are in preparation by specialists. When these are published, it may become possible to more objectively interpret our species con- cept, paleoenvironment, and the biostratigraphic use- fulness of the species. Trigonostoma (Ventrilia ?) insulare (Pilsbry and Johnson, 1917) is known only from its holotype which has been described from an unknown locality and stratigraphic horizon of “Santo Domingo”. As men- tioned under that species (p. 114, herein), it is possible that 7. (V. ?) insulare does not occur in the Neogene of the Dominican Republic. BIOGEOGRAPHY AND PALEOECOLOGY Twenty species of Cancellariidae have been reported from the areas studied, 19 of which are represented in our collections. These 20 species are grouped in 13 subgenera. All of these subgenera have at least one living representative in the Panamic—Pacific faunal province, but only seven of them have representatives living in the Caribbean faunal province. Lists of paciphile genera and subgenera of Cancel- lariidae were published by Woodring (1966, p. 428) and Vermeij (1978, p. 232). As those lists are now outdated, a current list is given below. An asterisk (*) denotes those taxa which occur in the Dominican Re- public. The first seven taxa listed are subgenera of Can- cellaria Lamarck, 1799, Perplicaria is a genus, and Extractrix is a subgenus of Trigonostoma Blainville, 1827. Euclia Adams and Adams, 1854 *Pyruclia Olsson, 1932 *Bivetiella Wenz, 1943 Narona Adams and Adams, 1854 *Hertleinia Marks, 1949 *Sveltia Jousseaume, 1887 *Massyla Adams and Adams, 1854 *Perplicaria Dall, 1890 Extractrix Korobkov, 1955 This list of nine genus-level taxa increases Vermeij’s 1978 list of six such taxa by 50%. Massyla was pre- viously known, but was overlooked in previous com- pilations; Bivetiella was known but was previously in- cluded in another subgenus; Hert/einia and Sveltia are here reported from the Neogene of the Caribbean for the first time; and Aphera Adams and Adams, 1854 was subtracted when Petuch (1981) reported a living species in the Caribbean. The relative abundance of the subgenera and species of Cancellariidae in the Tertiary of the Dominican Republic is very unequal. The 13 species placed in subgenera of Cancellaria account for just over 50% of the specimens examined. However, the single species of Aphera is represented by almost as many specimens (1,300+) as all other species of Cancellariidae com- bined. Species of Perplicaria, Trigonostoma, Axelella Petit, 1988, Agatrix Petit, 1967, and Admetula Coss- mann, 1889 are each represented by 16 or fewer spec- imens. Cancellariids inhabit subtidal to bathyal sand and mud bottoms in temperate and tropical regions. Depth and bottom condition records for some living relatives of the species under study are given in appropriate sections of the Systematic Paleontology portion of this paper. All the species treated in this paper (except 77ri- gonostoma (Ventrilia ?) insulare Pilsbry and Johnson, 1917, which is not represented in our collections) have been collected from silty sediments. The mode of oc- currence may vary somewhat within a single species. A large proportion of the collected shells occurred scat- tered in silts (/.e., at or close to the original place of burial). Many others, however, were found concen- trated in lenses, in silty bands, in shelly layers, in peb- bly beds, and in conglomeratic shell beds, lenses, and layers. These latter modes of occurrence point to prob- able transport over some distance before final depo- 92 BULLETIN 334 sition. Judging from the good state of preservation of the material, this transport must have been minor. Association of the species recorded herein with sandy sediments is rare. The following cases may be men- tioned: C. (C.) rowelli Dall, 1896 [scattered in pebbly, silty sands and in sands and silts of the Lopez section]; C. (B.) epistomifera Guppy, 1876 [in sand bed of the Rio Gurabo section]; Aphera islacolonis (Maury, 1917) [in sandy and pebbly silts of the Rio Cana section, and scattered in sands and silts of the Lopez section]; and Agatrix losquemadica (Maury, 1917) [in sand bed of the Rio Gurabo section]. These occurrences seem to be exceptions, and as a rule the species just mentioned also occur like the other cancellariid species. Regarding the early ontogenetic development of the species dealt with in this paper, it is noteworthy that all the species have blunt apices and that the orien- tation of the outer lip of their protoconchs is proso- cline. According to Shuto (1974), the blunt apices point to a lecithotrophic, direct type of development. This lack of a planktonic larval stage may account for the high degree of endemism previously mentioned. Little ecological data are available for cancellariids. Small juveniles of the Indo-Pacific species Cancellaria (Scalptia) contabulata Sowerby, 1832b [fig. 28] have been found on the spires of several species of living gastropods (Cernohorsky, 1972, p. 181). A Persian Gulf cancellariid, Nipponaphera paucicostata (Sowerby, 1894), was reported by Melvill and Standen (1901, p. 451) as being found “adhering to the upper part of Rapana bulbosa, 30-50 fathoms.” Cancellaria(S.) sca- lariformis Lamarck, 1822 [p. 113] has been found at- tached to living specimens of the bivalve Eucrassatella Iredale, 1924 in Australia (Garrard, 1975, p. 29). The reason for the association of these cancellariids with other molluscs is not known. The pallial complex and reproductive system of can- cellariids is similar to that of other neogastropods. The extreme anterior placement of the buccal ganglia, a simplified alimentary system posterior to the valve of Leiblein, and a highly specialized radula distinguish the Cancellariidae from other neogastropods. The can- cellariid radula is uniserial, each tooth being about 50 times as long as wide, with anteriorly directed cusps at the distal end of each tooth. This unique radula structure was the basis of the ordinal name Nemato- glossa (Olsson, 1970). For discussions of the soft parts and radulae of cancellariids, see Harasewych and Petit (1982, 1984). Based on the functional morphology of the alimen- tary system, it was speculated that cancellariids are suctorial fluid feeders (Harasewych and Petit, 1982; Petit and Harasewych, 1986). That this is true for at least one species was demonstrated by O’Sullivan, McConnaughey, and Huber (1987) who found that Cancellaria cooperi Gabb, 1865 [p. 186] feeds by suck- ing the blood of the Pacific electric ray, Torpedo cal- ifornica Ayres, 1855 [pp. 70, 71]. Neither food nor feeding habits are known for any other species of can- cellariid. Until more is known about the life history and ecol- ogy of Recent species of Cancellariidae, nothing can be inferred about their paleoecology. ABBREVIATIONS OF REPOSITORY INSTITUTIONS The following abbreviations for repository institu- tions are used in this paper: ANSP: Academy of Natural Sciences, Philadelphia, PA, U.S.A. BMNH: British Museum (Natural History), London, England, U. K. NMB: Naturhistorisches Museum Basel, Switzerland (the letter H after NMB stands for gastropods). PRI: Paleontological Research Institution, Ithaca, NY, U.S.A. TU: Tulane University, New Orleans, LA, U.S. A. USGS: United States Geological Survey, Washington, DC, U.S. A. USNM: United States National Museum of Natural History, Smithsonian Institution, Washington, DC, U.S.A. YPM: Peabody Museum of Natural History, Yale Uni- versity, New Haven, CT, U.S.A. SYSTEMATIC PALEONTOLOGY INTRODUCTION The base for the preparation of this paper has been the combined collections of the Naturhistorisches Mu- seum Basel and Tulane University. All of the type and figured specimens of Cancellariidae derived from these collections are deposited in the Naturhistorisches Mu- seum Basel. The type specimens of all species of Can- cellariidae occurring in the Dominican Republic, as well as the type specimens of species which were most important for comparative purposes, have been ex- amined and refigured. Although the amount of material available for this study is mentioned under each species, a summary of the number of specimens of each species is given in Table 1. The 20 species of Cancellariidae treated herein are grouped into 13 subgenera. Seven of these subgenera, containing 13 species, are assigned to the genus Can- cellaria Lamarck, 1799. The remaining seven species are placed in six genera. As is often the case with Ter- DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 93 Table 1.—Numbers of specimens of the nineteen species of can- cellarid gastropods collected in the Cibao Valley for this study. number of taxon specimens Cancellaria (Cancellaria) guppyi 75 Cancellaria (Cancellaria) mauryae 13 Cancellaria (Cancellaria) juncta 45 Cancellaria (Cancellaria) harrisi 528 Cancellaria (Cancellaria) rowelli 202 Cancellaria (Pyruclia ?) uva 4 Cancellaria (Sveltia) inquilinus 12 Cancellaria (Bivetiella) gabbiana 2 Cancellaria (Bivetiella) epistomifera 375 Cancellaria (Bivetiella) bajonensis 70 Cancellaria (Bivetopsia) plectilis 2 Cancellaria (Hertleinia) miranda 51 Cancellaria (Massyla) lopezana 2 Aphera islacolonis 1,300+ Perplicaria canae 2 Trigonostoma (Ventrilia) gurabis 10 Axelella emblema 3 Agatrix losquemadica 16 Admetula zalayana 5 tiary cancellariids, some species are represented by very few specimens. Fortunately, the species from the Do- minican Republic represented by few specimens are morphologically so distinct that they can be separated on a genus-level basis, eliminating any possibility that they are only aberrant specimens of common species. The extensive collections upon which this report is based have made it possible to examine large numbers of individuals of some species and thus obtain some indication of variation. Few species exhibit great vari- ability, even among those for which large numbers are available for study. Two notable exceptions are Can- cellaria (Cancellaria) harrisi Maury, 1917 and Aphera islacolonis (Maury, 1917). As mentioned elsewhere herein, genus-level division of the Cancellariidae has been inconsistent in pub- lished work, but probably no more so than in some other families. Subgenera are used herein to place to- gether species which share morphological characters that are not considered to be of sufficient importance to warrant separation as full genera. We consider these subgenera, most of which identify species groups with limited spatial and temporal distribution, to be of con- siderable potential value in better understanding the stratigraphy of the Caribbean Tertiary. Our use of gen- era and subgenera is subjective, but is based on pre- liminary results of a study of the phylogeny of the family (Harasewych and Petit, in preparation). In some instances cancellariids from the Dominican Republic treated herein as discrete taxa at the species level differ from populations from elsewhere in the Tertiary Caribbean faunal province in minor charac- ters that are, however, consistent within each popu- lation. We consider these consistent differences to be of systematic importance as it is possible that some of these, although seemingly minor (such as the persis- tence of surface ornamentation as discussed under Py- ruclia Olsson, 1932), may be of stratigraphic impor- tance. If they are, they might assist in establishing more precise temporal relationships between the various Tertiary Caribbean faunas. The Diagnosis is reserved for the description of su- praspecific categories. The Description is used only in species-level taxonomy, and is a description of the species, not of specimens. Several equivalent descrip- tive terms are used (e.g., volution and whorl, bifid and bifurcate; etc.), especially if they occur in the same sentence. Generic and subgeneric diagnoses begin with a gen- eral statement referring to size. These statements are not quantified as they are subjective and are based on work still in progress. No such general statement as to size is made in the descriptions of species as measure- ments are given under the heading Measurements. The measurements of many species are plotted in graphs. All measurements given for unnumbered specimens and all measurements plotted on graphs are of indi- vidual specimens and are not averages. Under the heading Material the number of speci- mens in the collections under study is given. The heading Occurrence is followed by detailed geo- graphic and stratigraphic information on a given species within the studied area of the Dominican Republic. Localities have been assigned to particular formations only in the sections on Rio Gurabo and Rio Cana, the Lépez section on Rio Yaque del Norte, and the lower part of the section on Rio Mao. In all the other areas such assignments are not possible as they are too spec- ulative at the moment (Saunders, Jung, and Biju-Du- val, 1986, p. 39). Under the heading Distribution there is general geo- graphic and stratigraphic information on a given species within the Dominican Republic p/us such information outside the Dominican Republic. Family CANCELLARIIDAE Forbes and Hanley, 1851 Remarks.—The neogastropod family Cancellariidae arose in the Lower Cretaceous, and was already well- developed and widely dispersed in the Early Tertiary. The cancellariids were highly experimental and de- veloped a wide variety of shell shapes (elongate to globose) and ornamentation (smooth to spinose). Many of these morphological forms, considered to be discrete taxonomic units, developed during the early history of the family, some being represented in the Recent fauna by only a few relict species. 94 BULLETIN 334 The family is usually divided into three subfamilies based on the genera Cancellaria Lamarck, 1799, Tri- gonostoma Blainville, 1827, and Admete Moller, 1842. Subfamily divisions are not utilized herein because a viable phylogeny for the family remains to be deter- mined. Preliminary results of work in progress indicate that several additional subfamilies may be required (Harasewych and Petit, in preparation). Genus-level division of the Cancellariidae has tra- ditionally been based on shell morphology, as has di- vision of most molluscan families. Cladistic analysis based on soft-parts morphology and cladistic analysis based on shell characters produce equally parsimoni- ous trees (Harasewych and Petit, in preparation) show- ing that there is a direct correlation between shell char- acters and soft-parts morphology. The division of the family into numerous genera and subgenera which seems necessitated by the wide variety of forms has been criticized by some authors. The problem was suc- cinctly stated by Woodring (1970, p. 334): “Cancel- lariids present a great variety of form and sculpture. Classification of such diverse species is dificult and so far unsatisfactory. Ample precedent is available for both conservative treatment and more narrowly re- stricted genera and subgenera.” Genus CANCELLARIA Lamarck, 1799 Cancellaria Lamarck, 1799, p. 71. Type species (by monotypy).—Voluta reticulata Linné, 1767. Recent, Caribbean. Cancellarius Montfort, 1810, p. 562. Type species (by original designation).— Voluta re- ticulata Linné, 1767. Recent, Caribbean. Exechoptychia Cossmann, 1903, p. 189. Type species (by original designation). — Cancellaria conradiana Dall, 1890. Pliocene, Florida and the Car- olinas. Diagnosis.—Shell of small to large size. General shape ovate, often high-spired. Outer lip prosocline with a weak stromboid notch. Interior of outer lip lirate. Col- umella with two or three folds, the adapical one being largest and overlying the prominent siphonal fasciole. Parietal callus weak but extending over the umbilical chink. Subgenus CANCELLARIA sensu stricto Diagnosis.—Shell of small to large size, generally ovate. Sculpture of axial ribs and spiral cords forming a cancellate pattern. Columella with three folds, the adapical one being largest and usually bifurcate. In- cremental growth evidenced by a thickening of the shell and slight lateral compression at intervals of approx- imately 120°. Remarks.—The nominotypical subgenus appears to be restricted to the later Tertiary and Recent faunas of the Western Hemisphere. The morphology of the shell and animal of Cancellaria reticulata (Linné, 1767) were described by Harasewych and Petit (1982). Cancellaria sensu stricto is usually restricted to those species with three distinct columellar folds, the adapical one being largest, broad, and bifid. A few species here assigned to Cancellaria s.s. have a sharp adapical fold, but possess the other characters of the subgenus. Cancellaria (Cancellaria) guppyi Gabb, 1873 Plate 15, figures 1-11; Plate 20, figures 1-3; Text-figure 10 Cancellaria guppyi Gabb, 1873, p. 236; Maury, 1917, p. 228, pl. 10, figs. 7, 8; Pilsbry, 1922, p. 333, pl. 22, fig. 7; Ramirez, 1956, p. Dieples ne le. ? Cancellaria guppyi Gabb. Anderson, 1929, p. 118. Description. —Protoconch smooth, of about two-and- one-half volutions. Teleoconch of about six whorls with finely reticulate sculpture of numerous spiral cords and collabral axial ribs. Shape globose with width being two-thirds of height. Suture impressed. Aperture sub- oval, elongate. Inner surface of outer lip with about 12 lirations which do not extend to outer edge, but which extend well into the aperture. Evidence of a slight stromboid notch present in growth lines. Parietal callus exists only as a thin coating on the adapical half of aperture but on the abapical half it is heavier and ex- tends as a shield over the umbilical chink. Columella with three folds, the bifid adapical fold overlying the siphonal fasciole and being much the largest; the de- scending center fold strong and often slightly bifurcate; the small abapical fold forms the edge of a short but pronounced siphonal canal. Lectotype.—ANSP 2990. This specimen has been figured by Pilsbry (1922, pl. 22, fig. 7) and is refigured here (PI. 15, figs. 1-3). Pilsbry (1922, p. 333) gave the measurements of “the type figured” and mentioned “the type and seven other specimens are no. 2990 A.N.S.P.” We take this designation of the figured spec- imen as “type” to constitute lectotype designation. Dimensions of lectotype.— Height, 28.1 mm; width, 18.6 mm. Type locality.—A type locality has not been desig- nated. Maury (1917, p. 64) cited the species from “Bluff 1, Cercado de Mao’. Bluff 1 of Maury is therefore considered the type locality (= TU 1293 = NMB 16910): late Miocene. For location see Saunders, Jung, and Biu-Duval (1986, text-fig. 29). Material.—Twenty-three lots with a total of about 75 specimens, most well-preserved, but almost always lacking the outermost part of the outer lip. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 95 Measurements.—The measurements of 42 speci- mens are plotted in Text-figure 10. Remarks.—These rotund shells usually exhibit flat- tened areas (see Pl. 15, fig. 2) reflecting rapid episodic growth as discussed by Harasewych and Petit (1982, p. 111) for C. reticulata (Linné, 1767). Anderson (1929, p. 118) reported Cancellaria guppyi _ from the Tubara Group of northern Colombia (late Miocene or early Pliocene) on the basis of a single specimen which he did not illustrate. Comparisons.—This distinctive species 1s easily rec- ognized by its fine cancellations, globose shape, deeply impressed suture, and thick shell. It can only be con- fused with Cancellaria mauryae Olsson, 1922 [p. 81, pl. 6, fig. 5] which also has fine cancellations but is much more attenuate, and C. juncta, n. sp., which has a thin shell, a proportionally longer and wider aperture, and a sharp adapical columellar fold. Occurrence.—This species is known from the fol- lowing localities (for locations see Saunders, Jung, and Biju-Duval, 1986, text-figs. 4-6, 29, 34, 35): Rio Mao: late Miocene: NMB 16910 and TU 1293 (both correspond to Bluff 1 of Maury). Cercado For- height in mm ft) iO fe Te Ws) we a SE i) ei) ea) width in mm Text-figure 10.—(Restored) height/width diagram of Cancellaria (Cancellaria) guppyi Gabb. mation (late Miocene): NMB 16916 and TU 1379 (both Arroyo Bajon); NMB 16913 (= Bluff 3 of Maury). Rio Gurabo: upper part of Cercado Formation (late Miocene): NMB 15898, 15899, 15900, 15902, 15903, 15906, 15907, 15910, 15911, 15912, 15916, and TU 1359, 1375. Lower part of Gurabo Formation (late Miocene): NMB 15869, 15871, 16809, and TU 1296. Rio Amina: probably late Miocene: TU 1219, 1220. Distribution.—Except for the report by Anderson (1929, p. 118) of a single specimen from northern Co- lombia, as mentioned above, the species has been re- ported only from the Dominican Republic. Cancellaria (Cancellaria) mauryae Olsson, 1922 Plate 15, figures 12-19 Cancellaria barretti Guppy. Guppy, 1866, p. 286 (list; not p. 289); Guppy, 1867, p. 157 (list, in part); Guppy, 1876, p. 520; Maury, 1917, p. 62, pl. 10, fig. 1; Maury, 1925b, pl. 9, fig. 17; Pilsbry, 1922, p. 332. Not of Guppy, 1866. Cancellaria reticulata Linné. Gabb, 1873, p. 236. Not of Linné, 1767. Cancellaria mauryae Olsson, 1922, p. 82, pl. 6, fig. 5. Description.— Protoconch smooth, of about two vol- utions. Both spiral and axial sculpture begin abruptly at the beginning of the first postnuclear whorl. Teleo- conch of about six whorls, slightly convex with an impressed suture. Finely reticulated sculpture consist- ing of about 17 primary spiral cords with one, or some- times two, weaker cords in the interspaces which begin on approximately the fifth teleoconch whorl, and axial ribs that are evenly spaced except for crowding behind the outer lip. Small nodes are formed at intersections of primary spiral cords and axial ribs. Outer lip slightly prosocline with a shallow stromboid notch. Columella with three sharp folds, the adapical one largest and abapical one forming the edge of the well-defined, but short, anterior canal. One or more short spiral ridges sometimes present between the extremities of the col- umellar folds. Weak parietal callus present on large specimens. Columellar callus strong, half covering the chink-like umbilicus. Holotype.—PRI 28661. This specimen was figured by Maury (1917, pl. 10, fig. 1), and by Olsson (1922, pl. 6, fig. 5), and is refigured here (Pl. 15, figs. 16-19). Olsson (1922, p. 83) selected Maury’s figured specimen as “type of this species’’. Dimensions of holotype.—Height, 36.7 mm; width, 22.7 mm. Type locality.—The exact locality from which the holotype was collected is not known. Maury (1917, p. 63) mentioned Bluffs 1, 2 and 3 on Rio Mao, Domin- ican Republic. We here restrict the type locality to locality NMB 16927 (Arroyo Bajon): late Miocene (Saunders, Jung, and Biju-Duval, 1986, text-figs. 30, 32). 96 BULLETIN 334 Material.—Five lots with a total of 13 specimens, most of them juvenile or incomplete. Measurements.— height width h/w (mm) (mm) — ratio PRI 28661 (holotype) [Pl. 15, figs. 16-19] 36.7 22.7 1.62 unnumbered specimen, from loc. TU 1294 29.8 17.9 1.66 unnumbered specimen, from loc. NMB 16922 38.7 23.3 1.66 unnumbered specimen, from loc. NMB 16923 25.0 14.5 D572. NMB H 17296, from loc. NMB 16927 (Pl. 15, figs. 12-15] 2522 16.4 1.54 Remarks.—Guppy (1866, p. 286; 1867, p. 157; 1876, p. 520) confused this Dominican Republic species with C. barretti Guppy, 1866 [p. 289, pl. 17, fig. 11] from the early Pliocene Bowden Formation of Bowden, Ja- maica, as did Maury (1917, 1925b) and Pilsbry (1922). Gabb (1873, p. 236) also misidentified this species as C. barretti, and placed it in the synonymy of C. retic- ulata (Linné, 1767). Olsson (1922, p. 82) rectified the situation by naming the Dominican species C. maury- ae. A discussion of C. barretti is given in the Appendix herein. Ramirez (1950, p. 12, pl. 3, fig. 3; 1956, pp. 12, 18, 19) reported C. barretti from the Dominican Republic. However, the identity of the specimen figured cannot be determined from the poor illustration which shows only a dorsal view. Ramirez does not mention colu- mellar dentition. It is probable that the species in ques- tion is C. mauryae. Comparisons.—Cancellaria (Cancellaria) mauryae differs from C. (C.) guppyi Gabb, 1873 in having a sharp, unbifurcated adapical columellar fold and in being less globose. A bifid adapical columellar fold is a characteristic of Cancellaria sensu stricto, and its absence in this species, which is otherwise much like many Caribbean Neogene species, is notable. Occurrence.—Rio Mao: Cercado Formation (late Miocene): TU 1294 (= Bluff 3 of Maury) and NMB 16915, 16922, 16923, 16927 (all mouth of Arroyo Ba- jon) (Saunders, Jung, and Biju-Duval, 1986, text-figs. 29, 30, 32). Distribution.—Except for Olsson’s (1922, p. 83) re- port of a single imperfect specimen from Water Cay, Panama (beds of late Miocene or early Pliocene age), the species is not known from outside the Dominican Republic. Cancellaria (Cancellaria) juncta, new species Plate 16, figures 10-16; Text-figure 11 Etymology of name.—L. juncta = associated. Description.—Slightly deviated protoconch smooth, prominent, of about two-and-one-half volutions. First teleoconch whorl with evenly spaced collabral ribs and fine spiral cords, the spiral cords quickly becoming equal in size to the axial ribs and forming typical can- cellate sculpture with small nodes formed where the cords cross the ribs. Teleoconch of about five whorls, the earlier having evenly cancellate sculpture. On the penultimate whorl the axial ribs lose prominence, ap- pearing more widely and sometimes irregularly spaced on the body whorl which has closely spaced growth lines. Spiral cords on the body whorl also become weak and more numerous than on earlier whorls. Suture slightly impressed. Varices, rarely formed, are indis- tinct. Prosocline outer lip thin with a distinct strom- boid notch. Inner portion of outer lip with about 20 short lirae which do not extend to the edge of the lip. Columella with three folds, the sharp undivided adap- ical one largest, descending, and overlying the siphonal fasciole. Center fold also descending and sharp. The abapical fold forms the edge of the short siphonal canal, and has a longitudinal crease making it bifid. No um- bilicus. Holotype.—NMB H 17297 (Pl. 16, figs. 10-13). height in mm Ww Ww 0 13 14 15 16 17 18 19 20 21 22 23 width in mm Text-figure 11.—(Restored) height/width diagram of Cancellaria (Cancellaria) juncta, n. sp. Dimensions of holotype.—Height, 33.8 mm; width, 23.5 mm. Type locality. —TU 1358: Rio Gurabo, upper part of Cercado Formation: late Miocene (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4-6). Material.—Nine lots with a total of 45 specimens; some of them juvenile or broken. Measurements.—The measurements of 20 speci- mens are plotted in Text-figure 11. Remarks.—Lateral compression, caused by incre- mental growth, is found only in very large specimens (over 40 mm) of C. juncta. Unfortunately, only in- complete specimens of that size have been found. Cancellaria juncta is here placed in Cancellaria sen- su stricto, even though its adapical columellar fold is not divided, due to its apparent close relationship to C. guppyi Gabb, 1873 and C. mauryae Olsson, 1922. Comparisons.— Young specimens of C. juncta may be distinguished from young C. guppyi by their less impressed sutures and proportionately wider aper- tures. Cancellaria juncta differs from C. guppyi in hav- ing a thinner shell, a wider aperture, an undivided adapical columellar fold, and a proportionately longer aperture. It differs from C. mauryae in having a round- ed tun-shaped shell that is more strongly constricted behind the siphonal fasciole. Occurrence.—Rio Gurabo: upper part of Cercado Formation (late Miocene): TU 1358, 1359, 1377, and NMB 15909, 15910, 15911, 15912, 15915, 15916 (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4-6). Distribution.— Not known from outside the Domin- ican Republic. Cancellaria (Cancellaria) harrisi Maury, 1917 Plate 17, figures 1-13; Plate 20, figures 4-6; Text-figure 12 Cancellaria harrisi Maury, 1917, p. 64, pl. 10, figs. 9, 10; Ramirez, 1956, p. 25. Not Cancellaria harrisi Maury. Li, 1930, p. 272, pl. 7, fig. 62 (= C. balboae Pilsbry, 1931; Recent). Description.—Smooth, erect protoconch of two-and- one-quarter to two-and-three-quarters volutions. Te- leoconch of about six or seven whorls. Sculpture of axial ribs crossed by spiral cords begins with onset of teleoconch. The prominent axial ribs decrease in num- ber on succeeding whorls. Number of axial ribs on body whorl of adult specimens of identical size ranges from 14 to 20, with as many as 30 on adults in some populations. Spiral cords cross the ribs, forming small nodes at intersections; two or three more prominent spiral cords below the shoulder forming larger nodes, making the teleoconch somewhat coronate. Spiral cords on early whorls coequal; intermediate cords appear on later whorls. On adults there are usually two inter- DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 97 mediate spiral cords which are not situated equidistant between the primary cords but crowd against the abap- ical side of the primary cords. Evidence of rapid epi- sodic growth sometimes indicated by flattened areas on the body whorl at approximately 120° intervals. Suture impressed. Outer lip thin, weakly crenulate, with a distinct stromboid notch. Interior of outer lip with about 15 short lirations which do not extend to lip edge. Columella with three folds, the adapical one overlying the siphonal fasciole being largest, and with distal portion slightly bifid. Abapical fold indistinct, forming the edge of the short, well-defined siphonal canal. Center fold descending, slightly bifid. All folds extend to the lip of the inductura. Umbilicus chink- like. Holotype.—PRI 28667 (Pl. 17, figs. 1-4). Dimensions of holotype.— Height, 29.5 mm; width, 17.5 mm. Type locality.—Maury (1917, p. 65) cited as occur- height in mm . 0) 8 Y 90 i We" CW i 8 i iB width in mm Text-figure 12.—(Restored) height/width diagram of Cancellaria (Cancellaria) harrisi Maury. 98 BULLETIN 334 rences her Zones H and I, Rio Cana at Caimito, Do- minican Republic. This being too imprecise, we here restrict the type locality to locality TU 1230: Cercado Formation (late Miocene) (Saunders, Jung, and Biu- Duval, 1986, text-fig. 15). Material.— Twenty-six lots with a total of 528 spec- imens. Measurements.—The measurements of 89 speci- mens are plotted in Text-figure 12. Remarks.— Although many specimens of C. harrisi are in our collections, it is common at only a few lo- calities with most lots consisting of five or fewer spec- imens. Of the total number studied, 56% are from a single locality (TU 1230), but only five specimens from that lot exceed 23 mm in height. Cancellaria harrisi was said by Maury (1917, p. 65) to be “the most beautiful of the Dominican Cancel- larias.’ Had she said “the most variable”, her com- ment would be less subjective. The number of axial ribs varies greatly, even within populations. This change in the number of axial ribs has a dramatic effect on the overall appearance of the shell. The shoulder nodes also vary in intensity, giving some shells a coronate aspect and others a round-shouldered appearance. Olsson (1932, p. 158), in a discussion of Euclia Adams and Adams, 1854 (type species: C. cassidifor- mis Sowerby, 1832a [p. 53]), placed C. harrisi in that subgenus. This placement was obviously based on the angled shoulder of C. harrisi, which seems exaggerated in Maury’s figure (the holotype is refigured herein, PI. 17, figs. 1-4). Marks (1949, p. 460) did not assign C. harrisi to a subgenus in his list of species. We prefer to place C. harrisi in Cancellaria s.s. until more is understood about the relationship between, and the necessity for, such genus-level taxa as Euclia and Py- ruclia Olsson, 1922. For further comments, see dis- cussion under Pyruclia [p. 100, herein]. Comparisons.— Young specimens are similar to C. metuloides Olsson, 1964 [p. 119, pl. 37, figs. 7, 7a] from the late Miocene Angostura Formation of north- western Ecuador, but that species has closely packed spiral cords and axial ribs which make the shell appear beaded instead of cancellate. In his discussion, Olsson (1964, p. 120) mentioned the similarity of the Ecua- dorian species to a “related undescribed Cancellaria” which “occurs in the Miocene beds at Baitoa, Santo Domingo.” This is probably a reference to young spec- imens of C. hAarrisi. \nother species from the Angostura Formation, Cancellaria maldonadoi Olsson, 1964 [p. 122, pl. 21, figs. 5, Sa] is close to C. harrisi, but differs in being less tabulate and in having fewer and weaker axial ribs. Woodring (1970, pp. 339, 340) considered C. mal- donadoi to be a nonspinose form of C. codazzii An- derson, 1929 [p. 116, pl. 14, figs. 4-7] from the Tubara Group (late Miocene or early Pliocene) of northern Colombia and placed it in the synonymy of C. codazzii. We consider the two to be distinct species. Cancellaria hettneri Anderson, 1929 [p. 114, pl. 10, figs. 5, 6] from the middle part of the Tubara Group (late Miocene or early Pliocene) of northern Colombia was stated by Anderson to be “allied to C. harrisi Mau- ry, but is more coarsely sculptured, larger, and more spinose.”” Woodring (1970, pp. 339, 340) justifiably placed C. hettneri in the synonymy of C. codazzii An- derson, a species more closely related to the Recent C. cassidiformis than to C. harrisi. Occurrence.— This species is recorded from the fol- lowing areas and localities (for locations see Saunders, Jung, and Biju-Duval, 1986, text-figs. 5, 15, 16, 21, 38): Rio Cana: upper part of Cercado Formation (late Miocene): TU 1230, 1282, 1301, and NMB 16835, 16837, 16838, 16839, 16842, 16844, 16852, 16854, 16855, 16856, 16986, 17003. Gurabo Formation (lat- est Miocene and early Pliocene): TU 1354 and NMB 16818, 16825, 16828, 16866, 16867, 16869. Rio Gurabo: Gurabo Formation (latest Miocene): TY) WAIT. Rio Yaque del Norte: early Pliocene: NMB 17268 (near La Barranca). Rio Verde: N. 18, early Pliocene according to Akers (in Vokes, 1989): TU 1250. Santiago de los Caballeros: early Pliocene: TU 1206. Distribution.— Not known from outside the Domin- ican Republic. Cancellaria (Cancellaria) rowelli Dall, 1896 Plate 18, figures 1-6; Plate 20, figures 7-9; Text-figure 13 Cancellaria rowelli Dall in Guppy and Dall, 1896, p. 307, pl. 29, fig. 1; Maury, 1917, p. 63, pl. 10, fig. 2; Pilsbry, 1922, p. 333. ? Cancellaria rowelli Dall. Olsson, 1922, p. 84, pl. 6, fig. 7; Weisbord, 1929, p. 50, pl. 6, figs. 9, 10. ? Cancellaria (Cancellaria) cf. rowelli Dall. Olsson, 1932, p. 156 ? Cancellaria (Cancellaria) aff. rowelli Dall. Jung, 1965, p. 551, pl. 75, figs. 7, 8. Description.—Protoconch smooth, of almost two- and-one-half volutions. Teleoconch of about seven whorls. Ornamentation begins with the formation of strong axial ribs at the onset of the teleoconch, with spiral cords being formed between the ribs before the onset of the second teleoconch whorl. Shell acute; su- ture impressed. On the body whorl there is a narrow, smooth channel between the shoulder and the suture. Sculpture of distinct axial ribs, about 35 on the body whorl. Fine spiral cords, of differing sizes, occur be- tween the ribs, crossing them only on the abapical half DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 99 of the body whorl, and at the shoulder where two or three are larger than others and form nodes where they cross axial ribs. Outer lip thin with about 12 short lirae within, which do not extend to the edge of the lip. Evidence of a shallow stromboid notch may be seen on the axial sculpture. Columella with three folds, the adapical one largest, overlying the siphonal fasciole. The descending abapical fold forms the edge of the short siphonal canal and parallels the center fold. Short secondary folds occur on the edge of the inductura. Holotype.—USNM 113762 (Pl. 18, figs. 1-3). Dimensions of holotype.— Height, 26.1 mm; width, 14.0 mm. Type locality.— Potrero, Rio Amina, Dominican Re- public. The type locality cannot be restricted at the moment because no material is available from the type area. Material.—Eighteen lots with a total of 202 speci- mens. Measurements.—The measurements of 49 speci- mens are plotted in Text-figure 13. 36 35 height in mm 34 33 32 31 30 29 te 28 27 i 26 25 - 24 23 22 bid 21 : as . co mes 19 0 11 12 13 14 15 16 17 18 19 width in mm Text-figure 13.—(Restored) height/width diagram of Cancellaria (Cancellaria) rowelli Dall. Remarks.— Maury (1917, p. 63, pl. 10, fig. 2) only published a copy of the original figure; she neither discussed the species nor gave any locality data. This most probably means that her field party did not collect the species at all. Olsson (1922, p. 84, pl. 6, fig. 7) reported two spec- imens of C. rowelli from the East Grape Creek, Costa Rica (age of beds not determined). His figured speci- men (PRI 20964) has a thickened terminal varix and generally coarser sculpture than specimens from the Dominican Republic, few of which reach the size of the Costa Rica specimens. It is possible that these two Costa Rica specimens are adults of C. anomoia Wood- ring, 1970 [p. 334, pl. 52, figs. 1, 2] from the lower part of the Gatun Formation (late Miocene) of Panama. Weisbord (1929, p. 50, pl. 6, figs. 9, 10) reported a single specimen of C. rowe/li from the Department of Atlantico, Colombia, from beds of undetermined age. This specimen (PRI 22950) has coarse sculpture and is possibly conspecific with the specimens from Costa Rica mentioned above. Olsson (1932, p. 156) compared some poorly pre- served material from the Montera Formation (age un- certain), Bayovar, Peru with C. rowelli, but pointed out differences which kept him from identifying his specimens as such, and stated that the Peruvian spec- imens may represent a different species. Jung (1965, p. 551, pl. 75, figs. 7, 8) compared spec- imens from the late early Miocene Cantaure Formation of the Paraguana Peninsula, Venezuela with C. rowelli, but pointed out consistent differences between them and Dominican specimens. As none of the above cited records can be deter- mined with certainty to be C. (C.) rowelli, they are omitted from our distribution records. Comparisons.—Cancellaria rowelli was compared by Dall (1896, p. 307), Maury (1917, p. 63), and Pilsbry (1922, p. 333) with the living Panamic—Pacific C. ur- ceolata Hinds, 1843 [p. 47]. It differs from that species in having a more rounded body whorl, in having finer sculpture, and in lacking an angled shoulder. Cancellaria rowelli differs from C. harrisi Maury in having less prominent axial ribs, and in being higher- spired. Also, in C. rowelli, not all of the spiral cords rise over the axial ribs. Occurrence.— As mentioned above, no material from the type area of this species is in our collections. All the studied specimens have been collected from the late early to early middle Miocene Baitoa Formation of the Lopez section on Rio Yaque del Norte (Saun- ders, Jung, and Biju-Duval, 1986, p. 30, text-figs. 21, 25), the only exception being locality TU 1226. Lo- cality TU 1226 is situated along the cliffon Rio Yaque del Norte, 500 m north of the bridge at the village of 100 BULLETIN 334 Baitoa (Saunders, Jung, and Biju-Duval, 1986, text- figs. 21, 28). The upper part of the cliff is inaccessible; it is made up of beds of the Baitoa Formation, which unconformably overlie beds of the Tabera Group. The fossils from locality TU 1226 are from the Baitoa For- mation; they were therefore not collected from out- crops, but picked up along the base of the cliff. The Lopez section has yielded material from the following localities: TU 1364, NMB 16935, 16936, 16938, 16940, 16942, 17265, 17280, 17281, 17282, 17283, 17284, 17286, 17287, 17288, 17289, 17290. Distribution.—Except for the questionable reports listed under Remarks above, the species is not known from outside the Dominican Republic. Subgenus PYRUCLIA Olsson, 1932 Pyruclia Olsson, 1932, p. 160. Peruclia Pilsbry and Olsson, 1941, p. 24 (error for Pyruclia). Type species (by original designation). — Cancellaria solida Sowerby, 1832a. Recent, Panamic—Pacific Prov- ince. Diagnosis.—Shell usually of large size, pyriform. Sculpture present only on early whorls; body whorl smooth or predominantly so. Columella with two pri- mary folds and a distinct siphonal fold. Of the two primary folds, the adapical one is much the larger and on large specimens is sometimes broadly divided, giv- ing the appearance of an additional fold. This division is not strictly bifid as in Cancellaria s. s., but is often widely U-shaped or shelved. Non-umbilicate. Remarks.—There are a number of nominal species of medium-sized to large cancellariids with sculptured early whorls and a smooth body whorl in the Neogene Caribbean faunal Province, and living in the Panamic— Pacific Province. They have similar characters, differ- ing primarily in shape, ranging from pyriform to barrel- shaped. The pyriform species can immediately be placed in Pyruclia, the subgenus erected for them. Those that are not pyriform cannot be placed, with any degree of certainty, in any named genus-level group. Some shells of similar shape differ in the number of volutions on which surface ornamentation persists, a difference which appears to be consistent within each lot. Although this seems to be a minor difference, it is thought advisable, in view of possible stratigraphic im- portance, to retain these as separate taxonomic units until more is known about their mutual relationship. Those species with sculpture on the early whorls and a smooth body whorl are: Cancellaria telemba Olsson, 1964 [p. 121, pl. 21, fig. 4; Angostura Formation, late Miocene, Ecuador]. This species can be distinguished by its high spire. The unnamed species figured by Marks (1949, pl. 78, fig. 9), tentatively referred to C. telemba by Ols- son, is a low-spired, barrel-shaped shell, the specific identity of which has not been determined. Cancellaria macneili Mansfield, 1937 [p. 609, pl. 85, figs. 1, 4; Choctawhatchee Formation, late Miocene, Florida] is similiar to C. telemba, with three or four spiral cords on the shoulder and numerous spiral cords on the anterior portion of the body whorl. Cancellaria lacondamini Olsson, 1964 [p. 121, pl. 21, figs. 1-lc; Picaderos Formation (but listed under Borbon Formation on p. 11), late Miocene or early Pliocene, Ecuador]. This is a massive, low spired, pyriform species which is easily separable from the others discussed here. Cancellaria scheibei Anderson, 1929 [p. 115, pl. 10, figs. 1-4; Usiacuri, Colombia; Tubara Group, late Miocene or early Pliocene]. This is also a large species, but it has a deeper and wider sutural channel than the other species. Cancellaria spatiosa Nelson, 1870 [p. 191; Tumbes Formation, late Miocene, Zorritos, Peru]. This is another large species which is similar to C. scheibei. Cancellaria diadela Woodring, 1970 [p. 338, pl. 53, figs. 7, 9; upper part of Gatun Formation, Panama; early Pliocene]. This species is close to C. scheibei in shape but differs in having an extremely low spire. Cancellaria auriculaperta Vokes, 1938 [p. 22, figs. 19, 20; Springvale, Trinidad; Springvale Formation, early Pliocene]. Rutsch (1942, p. 163) placed this in the synonymy of C. cibarcola Anderson, 1929, but it differs from that species in several aspects, one of which is its earlier loss of ornamentation. Woodring (1970, p. 338) recognized that Rutsch’s synonymy was incorrect. Cancellaria pycta Olsson, 1964 [p. 122, pl. 21, figs. 3, 3a; Angostura Formation, late Miocene, Ecuador] was considered to be a subspecies of C. cibarcola by Woodring (1970, p. 338). Woodring had two species combined as C. cibarcola (see discussion under C. cibarcola below), and the smaller of the specimens he figured differs from C. pycta only in the persis- tence of the sculpture. Cancellaria laevescens Guppy, 1866 [p. 289, pl. 17, fig. 12; Bowden Formation, early Pliocene, Jamaica] was reported from the Dominican Republic by Maury (1917, p. 64, pl. 10, fig. 6) and Pilsbry (1922, p. 333) from specimens collected by Gabb. These Domin- ican Republic specimens are Cancellaria (Pyruclia ?) uva, n. sp. Cancellaria (Pyruclia ?) laevescens dif- fers from most species discussed here 1n having strong spiral and axial sculpture persisting through the pen- ultimate whorl. For additional data on C. (P. ?) lae- vescens, see the Appendix herein. Cancellaria laevescens portoricana Maury, 1920 [p. 69, DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 10] pl. 7, fig. 10; Quebradillas, Puerto Rico; late Miocene or early Pliocene]. Described from an artificial cast of an external mold, this elongate shell cannot be identified with any of the species listed herein. See discussion under C. (P. ?) /aevescens in the Appendix herein. Cancellaria casicalva Marks, 1949 [p. 464, pl. 78, figs. 3, 10; Daule Formation, late Miocene, Ecuador] was described with the comment that “‘the subgeneric allocation is not known.” In general outline it re- sembles C. pycta Olsson, but differs in having a higher spire and in having two spiral bands which persist onto the body whorl. Cancellaria schucherti Olsson, 1932 [p. 162, pl. 17, figs. 3, 4; Tumbes Formation, late Miocene, Peru] is similar to the Recent C. obesa Sowerby, 1832a. It was considered to be a subspecies of that species by Pilsbry and Olsson (1941, p. 21) who placed C. schucherti in Cancellaria s. s. Olsson listed his de- scription of the species immediately following his description of the genus Pyruclia Olsson, 1932 and the listing of C. (P.) spatiosa Nelson, 1870. However, he did not assign it to a subgenus. Cancellaria cibarcola Anderson, 1929 [p. 116, pl. 14, figs. 1-3; Cibarco, Colombia; Tubara Group, prob- ably late Miocene] is close to C. auriculaperta. Woodring (1970, p. 338, pl. 52, figs. 9, 10; pl. 53, figs. 8, 10-12) combined two species from the lower part of the Gatun Formation of Panama (late Mio- cene) as C. cibarcola, neither of which are Ander- son’s species. The differences between the two species are consistent. The larger (Woodring, 1970, pl. 53, figs. 8, 10-12) has sculpture only on the first three teleoconch whorls and the plications on the inner portion of the outer lip are either weak or entirely absent. The smaller (Woodring, 1970, pl. 52, figs. 9, 10) has sculpture persisting through the fifth teleo- conch whorl, has strong lirations inside the outer lip, and is less broadly constricted anterior to the si- phonal fasciole. The relationship of these two species with others in this complex has not been determined. The smaller of the two species is obviously closely related to C. (P. ?) uva, n. sp. Cancellaria solida Sowerby, 1832a [p. 50], the type species of Pyruclia, is uncommon offshore from the Gulf of California to Peru (Keen, 1971, p. 654). It was reported from the Pliocene of Ecuador by Pilsbry and Olsson (1941, p. 24) and by Olsson (1964, p. 120). The living Cancellaria bulbulus Sowerby, 1832a [p. 55] has a limited range extending from El Salvador (Her- nandez, 1979, p. 204) to Panama (Keen, 1971, p. 654). It differs from C. solida in having a more at- tenuate spire, and its siphonal fasciole is either weak or absent. This species was also reported from the Pliocene of Ecuador by Pilsbry and Olsson (1941, p. 24). The living Panamic—Pacific species Cancellaria obesa Sowerby, 1832a [p. 52] and Cancellaria ovata Sow- erby, 1832a [p. 53] belong in Cancellaria s. s., but are mentioned here as both have almost smooth body whorls. Cancellaria (Pyruclia ?) uva, new species Plate 18, figures 7-9 Cancellaria laevescens Guppy. Guppy, 1866, p. 286 (list; not p. 289; Guppy, 1867, p. 167 (list, in part); Gabb, 1873, p. 236 (list); Guppy, 1876, p. 520; Maury, 1917, p. 64, pl. 10, fig. 6; Pilsbry, 1922, p. 333. Not of Guppy, 1866. Etymology of name.—L. uva = grape. Description.—Protoconch consists of a little more than two-and-one-half volutions. Teleoconch of about six or seven whorls. Sculpture on spire whorls of closely packed spiral cords and axial ribs of about equal size that form nodes at their intersections, making the sculpture appear pustulate instead of cancellate. The axial ribs disappear at the end of the penultimate whorl. The spiral cords weaken toward the end of the pen- ultimate whorl and on the body whorl are barely dis- cernible except in the depression above the siphonal fasciole. Suture impressed. Outer lip prosocline with a shallow stromboid notch and about 14 strong lirations within, which do not extend deeply into the aperture. Columellar callus thin but distinct. Body whorl con- stricted above the siphonal fasciole. Columella with three folds, the almost horizontal one largest, rarely slightly bifid, overlying the siphonal fasciole. The cen- ter fold and abapical fold descend sharply, the latter forming the edge of the short, well-defined anterior canal. Short secondary folds sometimes present on out- er edge of inductura. Holotype.—NMB H 17305 (Pl. 18, figs. 7-9). Dimensions of holotype.— Height, 30.0 mm; width, 19.5 mm. Type locality.— Locality NMB 17275: Rio Yaque del Norte, near mouth of Arroyo Lopez; probably late Miocene (Saunders, Jung, and Biyu-Duval, 1986, p. 30, text-figs. 21, 26, pl. 8, fig. 5). Material.— One lot containing four specimens, three of which are in good condition. Measurements.— height width h/w (mm) (mm) ratlo NMB H 17305 (holotype) [Pl. 18, figs. 7-9] 30.0 19.5 1.54 NMB H 17306 (paratype) 28.9 18.7 LESS NMB H 17307 (paratype) ANP? 18.3 1.49 102 BULLETIN 334 Remarks.—Specimens from the Gabb collection were cited by Maury (1917, p. 64, pl. 10, fig. 6) and Pilsbry (1922, p. 333) as C. laevescens Guppy, 1866. The spec- imen figured by Maury (PRI 28664) and the ones men- tioned by Pilsbry (ANSP 2985; three specimens) have been examined and prove to be C. uva. Cancellaria laevescens is discussed in detail in the Appendix herein. Comparisons.—Cancellaria uva differs from C. lae- vescens, with which it has been confused, in being smaller, in having weaker and more numerous axial ribs, in having a more rounded and less tabulate out- line, and in being less constricted above the siphonal fasciole. Also, strong sculpture persists on C. /aevescens until its shell is larger than that of C. uwva. It differs from the smaller specimen from the lower part of the Gatun Formation (late Miocene) figured by Woodring (1970, pl. 52, figs. 9, 10; not pl. 53, figs. 8, 10-12) as C. cibarcola Anderson, 1929, in having ornamentation persisting through the penultimate whorl. Occurrence.—Known only from locality NMB 17275, the type locality of the species. The provenance of the specimens referred to by Maury and by Pilsbry is not known. Distribution.— Not known from outside the Domin- ican Republic. Subgenus SVELTIA Jousseaume, 1887 Sveltia Jousseaume, 1887, p. 214. Type species (by original designation).—Sveltia var- icosa [sic] Brocchi [= Voluta varricosa Brocchi, 1814]. Pliocene, Italy. Diagnosis.—Shell of small to large size, normally high-spired with strong axial ribs which sometimes form spines at the shoulder. Shoulder usually angled. Columella with two folds, the adapical one slightly the larger, with a third incipient fold forming the edge of a short anterior canal. A chink-like umbilicus is present in most adults, as is a moderate parietal callus. Remarks.—Sveltia is represented in the Tertiary fau- nas of Europe, Chile, Ecuador, and Central America. In the Recent fauna it is represented off the West Coast of Africa and the West Coast of Central America. The species described below is the first Caribbean record of Sveltia, which should be added to the list of paciphile genera. Cancellaria (Sveltia) inquilinus, new species Plate 19, figures 1-8; Plate 20, figures 10-12 Etymology of name.—L. inquilinus = temporary in- habitant. Description.—Protoconch smooth, deviated, of one- and-three-quarters volutions. Teleoconch of about five shouldered whorls. Axial sculpture of strong collabral ribs, evenly spaced on early whorls but becoming less numerous on later whorls; about 11 ribs on penulti- mate whorl and nine or less on the body whorl. Axial ribs sometimes enlarged as varices. Spiral sculpture consists of cords of varying strength; about six primary cords on body whorl with three or four weaker cords in each interspace. The primary cords form sharp nod- ules where they cross the axial ribs; spiral cord on shoulder stronger than others and forming a short spine on each rib. Shoulder rounded back to an impressed suture. Aperture ovate. Slightly prosocline outer lip thickened with strong interior lirations which do not extend to the edge of the lip. Columella almost straight with two strong folds, the adapical one stronger; a third fold forms the edge of the short but distinct anterior canal. Columellar callus well-developed, sometimes not completely attached to the body whorl creating a pseu- doumbilicus. Umbilicus normally chink-like. Holotype.—NMB H 17309 (Pl. 19, figs. 5-8). Dimensions of holotype.— Height, 16.6 mm; width, 10.2 mm. Type locality.—Locality TU 1227, Arroyo Zalaya, Dominican Republic. Early Pliocene (Saunders, Jung, and Biyu-Duval, 1986, text-fig. 36 and table 3). Material.—Six lots containing 12 specimens. Measurements.—(complete specimens only): height width h/w (mm) (mm) ratio NMB H 17309 (holotype) [Pl. 19, figs. 5-8] 16.6 10.2 1.63 NMB H 17308 [Pl. 19, figs. 1-4] 16.7 10.8 1.55 unnumbered specimen, from loc. TOMI227: 8.2 Sal 1.61 unnumbered specimen, from loc. TU 1250 17.8 9.3 1.91 unnumbered specimen, from loc. TU 1250 12.0 8.0 1.50 unnumbered specimen, from loc. TU 1250 10.3 6.4 1.61 Remarks.—The Recent Panamic C. centrota Dall, 1896 [p. 13; Dall, 1908, p. 295, pl. 1, fig. 8] and C. gladiator Petit, 1976 [p. 35, pl. 1, fig. 2] live at a depth of 75 m or greater. Recent specimens of the West Af- rican C. /yrata (Brocchi, 1814) [p. 311, pl. 3, fig. 6] are recorded only from greater depths, the shallowest re- corded being 145 m (Barnard, 1959, p. 17.) Of the 12 specimens available. six are well-preserved adults, the others being juvenile or broken. Localities NMB 17267 and TU 1357 are each represented by a single badly broken specimen. Comparisons.—This new species is closest to C. zahni Bose, 1910 [p. 239, pl. 13, fig. 16] from beds of prob- ably late Miocene or early Pliocene age of the Isthmus of Tehuantepec. In comparison with C. inquilinus, n. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 103 sp., the Mexican species is more angular, has a deeper suture, possesses fewer axial ribs on early whorls, and is more umbilicate. The specimen figured by Toula (1911, pl. 29, fig. 12) as a “‘n. var.” of C. zahni differs from the Dominican species in being proportionally wider at the shoulder and in being less constricted behind the siphonal fasciole. Cancellaria inquilinus, n. sp., possesses shoulder spines proportional in size to those of C. centrota, but differs in most other respects. Occurrence.—This species is known from the fol- lowing six localities: TU 1227 (type locality): Arroyo Zalaya, early Pliocene. TU 1250: Rio Verde, N. 18, early Pliocene according to Akers (in Vokes, 1989); for location see Saunders, Jung, and Biju-Duval, 1986, text-figs. 3, 38. TU 1357: Rio Yaque del Norte, bluff on west side, just above new (1980) water plant at south edge of Bella Vista, 3 km (by road) south of bridge at Santiago de los Caballeros. Early Pliocene. NMB 17266 and 17267: La Barranca on Rio Yaque del Norte. Early Pliocene (Saunders, Jung, and Biju- Duval, 1986, text-figs. 21, 24 and table 3). NMB 17271: Arroyo Zalaya: early Pliocene (Saunders, Jung, and Biju-Duval, 1986, p. 34, text-fig. 36). Distribution.—Not known from outside the Domin- ican Republic. Subgenus BIVETIELLA Wenz, 1943 Bivetia Jousseaume, 1887, p. 193, non p. 163. Type species (by original designation). — Cancellaria similis Sowerby, 1833. Recent, northwest Africa. Bivetiella Wenz, 1943, p. 1356 (new name for Bivetia Jousseaume, 1887 [p. 193, non p. 163)). Type species (by original designation of Bivetia Jous- seaume, 1887 [p. 193].—Cancellaria similis Sowerby, 1833. Bivetiella Marks, 1949, p. 456. Type species (by original designation). — Cancellaria similis Sowerby, 1833. Recent, northwest Africa. Diagnosis.—Shell of medium size, stout, with a broad body whorl. Sculpture usually strongly and regularly cancellate. Varices present. Prosocline outer lip everted with a distinct stromboid notch. Inner portion of outer lip lirate. Columella with three folds, the adapical fold being sharp and the center and abapical folds either bifurcate or incipiently so. Umbilicus narrow or chink- like. Remarks.—Jousseaume’s 1887 monograph on Can- cellariidae was published in parts. He unfortunately used the name Bivetia for two distinct genus-level groups. His first usage (p. 163) of Bivetia was in the description of B. mariei Jousseaume, 1887 (= C. in- dentata Sowerby, 1832a) which established that species as the type species of Bivetia by monotypy. One month later he described the genus Bivetia, naming C. similis Sowerby, 1833 as type species, and not even mention- ing B. mariei in his list of included species. Recognizing that Jousseaume had used Bivetia in two different senses, Wenz (1943, p. 1356) proposed the replacement name Bivetiella. Marks (1949, p. 456) came to the same conclusion and proposed exactly the same name used six years earlier by Wenz. It should be mentioned here that in 1949 there were only one or two copies of Wenz’s work in the United States, and it was not available to Marks. That they should both propose the same name is not surprising, as Bivetia (of Jousseaume’s second usage) was derived from “Le Biy- et’, Adanson’s (1757, p. 123) name for the type species, and both Wenz and Marks evidently wished to retain this association. Three Dominican species are here placed in the sub- genus Bivetiella. Jung (1965, p. 549), in discussing his placement of C. epistomifera Guppy, 1876 in Cancel- larias. s., stressed the differences in the relative strength and number of axial ribs between the tropical Amer- ican species and the type species. Some species from the later Tertiary of Europe, almost certainly ancestral to C. similis, exhibit sculptural patterns similar to C. (B.) epistomifera. The primary consideration for place- ment in Bivetiella is, however, the columellar denti- tion. In Cancellaria s. s. the adapical columellar fold is almost always wide and bifurcate. In Bivetiella, the adapical fold is sharp, but the central and the abapical fold bordering the siphonal canal are either bifurcate or incipiently so. Also, species of Bivetiella have marked varices which do not occur in species of Cancellaria S. S. Cancellaria (Bivetiella) gabbiana Pilsbry and Johnson, 1917 Plate 19, figures 9-15 Cancellaria gabbiana Pilsbry and Johnson, 1917, p. 163; Pilsbry, 1922, p. 334, pl. 22, fig. 12. Description.—Smooth, slightly bulbous protoconch of about two volutions. Teleoconch of about six whorls. Spiral sculpture of raised cords, five visible on pen- ultimate whorl and about 15 on body whorl. Suture deeply impressed and narrowly canaliculate. Slightly rounded axial ribs, about 16-18 on body whorl. Var- ices present, weak on early whorls, occurring at ap- proximately 120° intervals. Outer lip somewhat everted, thickened by terminal varix, flaring abapically. Shal- low stromboid notch. Aperture wide, ovate, with about 104 BULLETIN 334 14 sharp interior lirations which do not extend to the margin of the lip, and extend inward only a short dis- tance. Parietal callus thin. Columella with three folds, the sharp adapical one being largest and almost hori- zontal; the center fold shorter and broader, descending sharply; the abapical fold borders the siphonal canal adaxially but becomes parallel to the center fold on the columellar callus. Short secondary folds and tubercules present on columellar callus. Umbilicus chink-like. Lectotype (here designated).—ANSP 3288. This is the specimen figured by Pilsbry (1922, pl. 22, fig. 12) and refigured here (Pl. 19, figs. 9-11). Dimensions of lectotype.— Height, 24.5 mm; width, 19.8 mm. Type locality.—Santo Domingo. No locality data were given by Pilsbry and Johnson, and the type lo- cality is here restricted to locality NMB 16938: Lopez section on Rio Yaque del Norte, Dominican Republic (Baitoa Formation, late early to early middle Miocene). Material.—Known only from the lectotype, one paralectotype, and two specimens from locality NMB 16938. Measurements.— height width h/w (mm) (mm) ratio ANSP 3288 (lectotype) [Pl. 19, figs. 9-11] 24.5 19.8 1.24 ANSP 67545 (paralectotype) 25.2 19.2 1.31 NMB H 17311 [PI. 19, figs. 12-15] 24.2 19.8 122 unnumbered specimen, from loc. NMB 16938 26.3 21.4 1.23 Remarks.—The wide, flaring aperture and squat form of C. gabbiana distinguish it from its congeners. This species is evidently quite rare, as only four spec- imens are known. It was not present in Maury’s col- lections. Occurrence.—Known only from locality NMB 16938: Lopez section on Rio Yaque del Norte; Baitoa Formation (late early to early middle Miocene (Saun- ders, Jung, and Biju-Duval, 1986, p. 30, text-fig. 21). Distribution.—Not known from outside the Domin- ican Republic. Cancellaria (Bivetiella) epistomifera Guppy, 1876 Plate 21, figures 1-11; Text-figure 14 Cancellaria moorei Guppy. Guppy, 1866, p. 286 (list, in part; not p. 289); Guppy, 1867, p. 157 (list, in part); Gabb, 1873, p. 236; Guppy, 1876, p. 520; Dall, 1903, p. 1583 (list, in part). Not of Guppy, 1866. Cancellaria epistomifera Guppy, 1876, p. 520, pl. 28, fig. 9; Maury, 7, p. 63, pl. 10, figs. 3, 4, 5 (in part: fig. 5 only; figs. 3, 4 = C. 1961, p. 52, pl. 14, figs. 1-9; Ramirez, 1950, p. 22, pl. 3, fig. 4; Ramirez, 1956, pp. 12, 18, 19, 23. Not Cancellaria epistomifera Guppy. Cossmann, 1913, p. 53, pl. 4, figs. 5, 6 ( dariena Toula, 1909). ? Cancellaria epistomifera Guppy. Olsson, 1922, p. 83 (?= C. epis- tomifera lipara Woodring, 1970, p. 337, pl. 52, figs. 7, 8; non C. lipara Woodring, 1951; = C. epistomifera sathra Woodring, 1973, p. 481). Not Cancellaria epistomifera Guppy. Maury, 1925a, p. 193, pl. 35, fig. 7 (= C. montserratensis Maury, 1925a). Not Cancellaria epistomifera Guppy. Maury, 1925b, pl. 9, fig. 11; (= C. (B.) bajonensis, n. sp.). Not Cancellaria epistomifera Guppy. Jung, 1965, p. 548, pl. 75, figs. 1-2. Description.—Protoconch slightly mammillated, of just over two-and-one-half volutions. Teleoconch of about six convex whorls, rounded at the periphery. Sculpture starts with a few weak spiral cords which are soon crossed by prosocline axial ribs. On later whorls the axial ribs, and the spiral cords which cross over them, are about equal in strength except on the later varices where the axial ribs are replaced by crowded growth lines. Development of varices variable, usually beginning on the third or fourth teleoconch whorl, with those on early whorls usually being weak. Varices spaced at approximately 120° increments. Penultimate whorl with five or six spiral cords in evidence, which are sometimes doubled on the sharply rounded shoulder behind which there is a narrow, smooth, channeled area abaxial to the deeply impressed suture. Prosocline outer lip strongly everted at the deep stromboid notch. Interior of outer lip with over a dozen strong lirations which do not extend deeply within. Parietal callus weak. 32 ° height in mm 0 13 14 15 16 17 18 19 20 21 22 width in mm Text-figure 14.—(Restored) height/width diagram of Cancellaria (Bivetiella) epistomifera Guppy. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 105 Abapical portion of columellar lip partially obscuring the narrow umbilicus. Columella with three folds, the adapical one almost horizontal and undivided. The center fold is descending, sometimes slightly bifurcate. The bifurcate abapical fold borders the siphonal canal adaxially but on the columellar callus swings out to the edge of the inductura, parallel to the center fold. Short secondary folds and tubercules usually present on the columellar callus. Lectotype.—BMNH G 83955 (figured herein). Se- lected by Pflug (1961, p. 53). Dimensions of lectotype.— Height, 28.3 mm; width, 19.7 mm. Type locality.—Yaque River (original label; see also Heneken, 1853, fig. 1), Dominican Republic. This in- formation is rather vague. The material of C. (B.) ep- istomifera used for this paper was collected from sev- eral sections (see under Occurrence) including the Rio Yaque del Norte section. However, specimens from that area are extremely scarce, and for this reason it is not advisable to restrict its type locality to a well- defined locality at this time. Material.— Thirty-seven lots consisting of over 375 specimens, many juvenile or broken, but a large num- ber of complete adult specimens. Measurements.—The measurements of 46 speci- mens are plotted in Text-figure 14. Remarks.—There has been considerable confusion regarding the Caribbean and Panamic Tertiary species of Bivetiella Wenz, 1943, probably due to the scarcity of sufficient material for comparison. Maury (1917) had both C. (B.) epistomifera and C. (B.) bajonensis, n. sp. and combined them. She divided her specimens into two sets: (1) with small protoconch; (2) with large protoconch. On the advice of Dall, she considered set (1) to be true C. epistomifera and suggested that set (2) might be a variety (her fig. 5 is indicated as “variety” on the plate caption). Unfortunately, this arrangement is reversed from the actual situation. Maury’s set (1), figured by her as C. epistomifera (pl. 10, figs. 3, 4; PRI 28662), is actually C. (B.) bajonensis. Maury’s figured “variety” (pl. 10, fig. 5; PRI 28663), is C. (B.) epistom- ifera Guppy. Jung (1965, pp. 548-550, pl. 75, figs. 1-3) identified one adult and nine immature specimens from the late early Miocene Cantaure Formation of the Paraguana Peninsula, Venezuela as C. epistomifera Guppy. The adult specimen is much more attenuate than any spec- imens of C. (B.) epistomifera we have examined from the Dominican Republic and, as pointed out by Jung, it has secondary spiral cords on the body whorl. For these reasons, we do not consider the Venezuelan spec- imens to be conspecific with C. (B.) epistomifera. Woodring (1970, pp. 335-337) referred a number of small cancellariids to C. epistomifera dariena Toula, 1909, and C. epistomifera lipara Woodring, 1970 (non C. lipara Woodring, 1951; = C. epistomifera sathra Woodring, 1973, p. 481). It is unlikely that all of the varieties figured by Woodring actually represent these two subspecies. However, none of the specimens from the Gatun Formation (late Miocene and early Pliocene) has an enlarged protoconch. Guppy (1866, p. 286; 1867, p. 157; 1876, p. 520) evidently confused C. moorei Guppy, 1866 [p. 289, pl. 17, fig. 7] from the Bowden Formation (early Pliocene) of Jamaica and C. (B.) epistomifera, as did Gabb (1873, p. 236). Their listing was repeated by Dall (1903, p. 1583). Cancellaria moorei is discussed in the Appendix herein. Comparisons.— Cancellaria (Bivetiella) epistomifera differs from the two subspecies recognized by Wood- ring (see above under Remarks) and other species of Bivetiella in having a large bulbous protoconch. Occurrence.— This species is recorded from localities of the following sections (from west to east): Rio Cana (Saunders, Jung, and Biju-Duval, 1986, text-figs. 15, 16): Cercado Formation (late Miocene): NMB 16857; lower part of Gurabo Formation (latest Miocene): NMB 16821. Rio Gurabo (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4-6): upper part of Cercado Formation (late Miocene): TU 1359 and NMB 15898, 15899, 15902, 15903, 15904, 15905, 15906, 15907, 15910, 15911, 15912, 15915. Gurabo Formation (late Miocene and barely into the early Pliocene): TU 1210, 1211, 1212, 1213, 1214, 1215, 1231, 1278, and NMB 15804, 15809, 15813, 15816, 15817, 15818, 15868. Rio Mao (Saunders. Jung, and Biju-Duval, 1986, text-fig. 29): late Miocene: TU 1293 (= Bluff 1 of Mau- ry); possibly early Pliocene: TU 1225 and NMB 16801. Rio Amina (Saunders, Jung, and Biju-Duval, 1986, text-figs. 34, 35): probably late Miocene: TU 1219, 1220. Rio Yaque del Norte (Saunders, Jung, and Biju-Du- val, 1986, text-figs. 21, 24, table 3): Baitoa Formation (late early to early middle Miocene): TU 1364 (Lopez section) and early Pliocene: NMB 17267 (La Barranca). Rio Verde (Saunders, Jung, and Biju-Duval, 1986, text-fig. 38): N. 18, early Pliocene according to Akers (in Vokes, 1989): TU 1250. Distribution.— Olsson (1922, p. 83) reported C. ep- istomifera Guppy from “Gatun Stage: C.Z.” on the basis of “a few, small and imperfect specimens.” As it was not found in Panama or the Canal Zone by Wood- ring, it is probable that Olsson’s specimens represent the subspecies C. epistomifera sathra, or some related species. Jung’s (1965) report from Venezuela is dis- cussed above under Remarks. The nominotypical sub- 106 BULLETIN 334 species is not known, with certainty, from outside the Dominican Republic. Cancellaria (Bivetiella) bajonensis, new species Plate 22, figures 1-7; Plate 26, figures 1-3; Text-figure 15 Cancellaria epistomifera Guppy. Maury, 1917, p. 63 (in part), pl. 10, figs. 3, 4 (not fig. 5); Maury, 1925b, pl. 9, fig. 11. Not of Guppy, 1876. Etymology of name.—Named after Arroyo Bajon, the type locality. Description.—Protoconch small, ofabout two smooth volutions. Teleoconch of about six slightly tabulate whorls. Spiral and axial sculpture begin immediately at onset of teleoconch. Axial sculpture of sharp ribs, closely and evenly spaced except on the varices where they are packed together and are barely visible as in- dividual ribs. Development of varices variable, usually beginning on third or fourth teleoconch whorl, with those on early whorls being weak. Varices spaced at approximately 120° increments. Spiral sculpture of evenly spaced spiral cords, about four visible on pen- ultimate whorl. On the body whorl there is usually a secondary spiral cord in the interspaces. Sharp nodes are formed where the spiral cords cross the axial ribs. The first spiral cord abapical from the crowded cords on the sharply rounded shoulder larger than the others, giving the shell a slightly angular, tabulate appearance. Suture impressed. Outer lip prosocline, everted at the distinct stromboid notch. Interior of outer lip with over a dozen strong lirations which do not extend deeply within. Columella with three folds, the adapical one sharp and almost horizontal. Center fold descending, usually unevenly bifurcate. The abapical fold forms the edge of the distinct siphonal canal and tends to be bifurcate. Short secondary folds and pustules present on the inductura. Umbilicus narrow but deep, partially obscured by the inductura. Holotype.—NMB H 17314 (PI. 22, figs. 1-4). Dimensions of holotype.— Height, 32.8 mm; width, 22.6 mm. Type locality. —NMB locality 16923: Arroyo Bajon, Rio Mao. Late Miocene (Saunders, Jung, and Biju- Duval, 1986, p. 32, text-figs. 29, 30, table 3). Material.—Sixteen lots consisting of 70 specimens, many juvenile or broken. Measurements.—The measurements of 14 speci- mens are plotted in Text-figure 15. Remarks.—For a discussion of the misidentification of this species by Maury (1917) see Remarks under C. epistomifera Guppy, 1876 [p. 105, herein]. Comparisons.—Cancellaria bajonensis differs from C. epistomifera, with which it has been confused, in having a smaller protoconch, a lower spire, and much sharper sculpture. The two species can be distinguished not only by sight, but by touch. Occurrence.—Rio Mao: Cercado Formation (late Miocene): TU 1294 and NMB 16913, 17269 (all cor- respond to Bluff 3 of Maury); TU 1379 and NMB 16916, 16917, 16918, 16923, 16924, 16927, 16928 (all Arroyo Bajon); NMB 16914, 16929, 16931, 16932 (all correspond to Bluff 2 of Maury). Late Miocene: NMB 16910 (= Bluff 1 of Maury). (See Saunders, Jung, and Biju-Duval, 1986, p. 32, text-figs. 29, 30, table 3.) Distribution.— Not known from outside the Domin- ican Republic. Subgenus BIVETOPSIA Jousseaume, 1887 Bivetopsia Jousseaume, 1887, p. 193. Bivetopsis Jousseaume (emendation by Cossmann, 1899, p. 9). Type species (by subsequent designation, Cossmann, 1888, p. 784).—Cancellaria chrysostoma Sowerby, 1832a. Recent, Panamic—Pacific Province. Diagnosis.—Shell small, stocky. Outer lip proso- cline, aperture rounded. Columella with three folds of about equal size. The almost horizontal adapical fold is slightly the largest. The descending abapical fold which borders the short siphonal canal is only slightly smaller, and the center fold, parallel to the abapical fold, is usually smallest. The body whorl is constricted above the strong siphonal fasciole. 39 38 height in mm 37 : 36 :. 35 34 33 0 19 20 21 22 23 24 25 26 27 width in mm Text-figure 15.—(Restored) height/width diagram of Cancellaria (Bivetiella) bajonensis, n. sp. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 107 Remarks.—We consider Bivetopsia to be an Amer- ican group, the few known species being confined to the later Tertiary of Florida, the Caribbean, and Ec- uador, and to the Recent fauna of the Caribbean and the Panamic—Pacific provinces. The relationship be- tween Bivetopsia and Bivetiella Wenz, 1943 should be investigated. As pointed out by Woodring (1928, p. 221) these two groups have much in common. Cancellaria (Bivetopsia) plectilis, new species Plate 22, figures 12-15 Etymology of name.—L. plectilis = complicated or intricate. Description.—Paucispiral protoconch smooth, of about two volutions. Teleoconch of about five shoul- dered whorls. Sculpture of strong spiral cords, about nine on the body whorl, which are each composed of two to five smaller cords. Axial sculpture consists of strong rounded ribs, about nine on the body whorl, and numerous fine, closely packed growth lines. Body whorl severely constricted behind the strong siphonal fasciole. Outer lip prosocline, crenate, with about 11 lirae extending well back into the aperture. Posterior canal small but distinct. Columella with three strong folds, the most abapical one forming the edge of the short upturned siphonal canal. Columellar callus well developed, tuberculose, extending in front of the small umbilicus. Holotype.—NMB H 17317 (PI. 22, figs. 12-15). Dimensions of holotype.—Height, 25.2 mm, width, 18.0 mm. Type locality.—Locality TU 1354, Canada de Zam- ba, Rio Cana, Dominican Republic; Gurabo Forma- tion (early Pliocene). (See Saunders, Jung, and Biu- Duval, 1986, text-figs. 15, 16.) Material.—Known only from the holotype and one paratype, both well-preserved. Measurements.— height width h/w (mm) (mm) ratio NMB H 17317 (holotype) [Pl. 22, figs. 12-15] 25.2 18.0 1.40 NMB H 17318 (paratype), from loc. TU 1422 20.3 16.5 1.23 Remarks.—There do not seem to be any published depth records for the Recent species of Bivetopsia Jous- seaume, 1887. We have examined specimens of the West Indian Cancellaria (Bivetopsia) rugosa Lamarck, 1822 [p. 115] which were dredged from a sandy bottom at moderately shallow depth. A specimen of this species (ANSP 272203) was taken in 25 m off Coral Bay, St. John, Virgin Islands (Rosenberg, written commun., 1988). Specimens of C. (B.) haemastoma Sowerby, 1832a [p. 54] are sometimes collected intertidally in the Galapagos Islands. Comparisons.—Cancellaria (Bivetopsia) plectilis is closely related to C. (B.) moorei Guppy, 1866 [p. 289] from the Bowden Formation (early Pliocene) of Ja- maica and its Floridian Pliocene subspecies, C. (B.) moorei pachia Smith, 1940 [p. 45], but differs in having rounded spiral cords subdivided by lesser cords and in having wider and more rounded ribs. The lectotype of C. moore is figured herein (Pl. 22, figs. 8-11). The Recent Caribbean species C. (B.) rugosa possesses sub- divided spiral cords but has weaker axial ribbing, a more angled shoulder, and is less constricted behind the siphonal fasciole. The Dominican species also dif- fers from the Recent Panamic and Galapagan species in details of sculpture and form. Occurrence.— Known only from two localities: TU 1354 (the type locality): Gurabo Formation (early Plio- cene) of Rio Cana (Canada de Zamba) and TU 1422: Arroyo Bellaco, a tributary of Rio Cana from the east, 3 km southwest of Las Caobas (age not determined: not plotted on any map in Saunders, Jung, and Biju- Duval, 1986). Subgenus HERTLEINIA Marks, 1949 Hertleinia Marks, 1949, p. 457. Type species (by original designation). — Cancellaria mitriformis Sowerby, 1832a. Recent, Panamic—Pacific Province. Diagnosis.—Shell of medium size, slender, mitri- form with an elongate aperture. Outer lip crenate with a prominent stromboid notch and a slightly recurved anterior canal. Columella fairly straight with two cen- tral folds, the adapical one largest. Sculpture cancellate. Remarks.—Hertleinia is treated here in its tradi- tional placement as a subgenus of Cancellaria La- marck, 1799, although it should probably be given status as a full genus. The Dominican species described below is the fourth known species of Hertleinia. Be- sides the Recent type species, the only previously known species are the Ecuadorian C. (H.) angosturana Marks, 1949 [p. 463, pl. 78, figs. 1, 2] from the late Miocene Angostura Formation, and C. (H.) marksi Olsson, 1964 [p. 125, pl. 37, fig. 6] from the early Pliocene Esmer- aldas Formation. Hertleinia must be added to the list of paciphile taxa. Cancellaria (Hertleinia) miranda, new species Plate 23, figures 1-5; Plate 26, figures 4-7; Text-figure 16 Etymology of name.—L. mirandus = wonderful, strange. Description.—Protoconch smooth, high-spired, of 108 BULLETIN 334 about two volutions. Attenuate teleoconch of about seven whorls. Sculpture of numerous spiral cords, about 17 on body whorl; evenly spaced except for slight crowding near suture. Sharp well-defined collabral ribs, about 20 on body whorl, evenly spaced except for crowding behind outer lip on adults. Outer lip slightly prosocline, rarely somewhat prosocyrt, crenate with a stromboid notch. Aperture ovate with strong but short interior lirations which do not extend to the outer lip. Columella almost straight with two sharp folds, the adapical one larger. Abapical portion of columella sloped to left and forming a short, well-defined, slightly upturned, anterior canal. Weak parietal wash some- times present. Holotype.—NMB H 17319 (Pl. 23, figs. 1-3). height in mm 0 11 12 13 14 15 16 17 18 19 width in mm Text-figure 16.—(Restored) height/width diagram of Cancellaria (Hertleinia) miranda, n. sp. Dimensions of holotype.— Height, 37.3 mm; width, 17.2 mm. Type locality.—Locality NMB 15903: Rio Gurabo, Dominican Republic; upper part of Cercado Forma- tion; late Miocene (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4, 6). Material.— Eight lots with a total of 51 specimens, most of which are complete and well-preserved. Measurements.—The measurements of 31 speci- mens are plotted in Text-figure 16. Remarks.—The Recent C. (H.) mitriformis Sowerby, 1832a [p. 51] ranges from Panama to Peru, “mostly offshore in depths to 37 m” (Keen, 1971, p. 653), al- though it is rarely collected intertidally in Panama. There is little variation among individuals of C. (H.) miranda. This is also true of C. (H.) mitriformis. Comparisons.— The Dominican species differs from the other species of Hertleinia Marks, 1949 in not being tabulate. The previously described species of Hertlei- nia all have one or two strong spiral cords forming a shoulder behind which the shell slopes to the suture. In C. (H.) miranda, shouldering is present in juveniles, but not in adults. The sculpture of C. (H.) miranda is much finer than that of C. (H.) angosturana Marks, 1949 and C. (H.) marksi Olsson, 1964 from Ecuador. It also has a wider, more flaring aperture. The holo- types of C. (H.) angosturana and C. (H.) marksi are figured herein on Plate 23, figures 6-8 and 9-11, re- spectively. Occurrence.—Rio Gurabo: upper part of Cercado Formation (late Miocene): NMB 15903, 15907, 15910, 15911, 15912, 15914, 15915, 15916 (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4, 6). Distribution.— Not known from outside the Domin- ican Republic. Subgenus MASSYLA Adams and Adams, 1854 Massyla Adams and Adams, 1854, p. 278. Type species (by monotypy).— Cancellaria corrugata Hinds, 1843. Recent, Panamic—Pacific Province. Diagnosis.—Shell of medium to large size. Whorls rounded, enlarging rapidly in early growth stages. Su- ture impressed. Sculpture consists primarily of spiral cords; axial folds and growth lines sometimes in evi- dence. Aperture ovate. Outer lip prosocline, lirate within. Columella with two strong folds, the adapical one being larger. Remarks.—Massyla occurs in the later Tertiary of the southeastern United States, Central America and northwestern South America, and in the Recent fauna of the Panamic—Pacific Province. Massyla is a paci- phile subgenus, although not listed as such by Wood- ring (1966, p. 428) or Vermeij (1978, p. 232). DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 109 Cancellaria (Massyla) lopezana, new species Plate 23, figures 12-18 Etymology of name.—Named after the Lopez sec- tion on Rio Yaque del Norte, Dominican Republic. Description.—Protoconch smooth, of about one-and- one-half volutions; border between protoconch and first postnuclear whorl marked by a slightly prosocline line, after which the sculpture consisting of six spiral threads and crowded, somewhat prosocline growth lines, starts immediately. There are about four-and-one-half teleo- conch whorls, well rounded with deeply impressed su- ture. Body whorl with about 18 evenly spaced spiral cords. Aperture ovate, outer lip prosocline, strongly and deeply lirate within. Columella straight with two strong folds, the adapical one larger, which extend to the edge of the thin parietal callus behind which there is a moderately small but deep umbilicus. Siphonal fasciole strong. The adapical portion of the outer lip bears a strong fold appressed to the body whorl, form- ing a small posterior canal. Holotype.—NMB H 17323 (PI. 23, figs. 12-15). Dimensions of holotype.— Height, 29.6 mm; width, 21.7 mm. Type locality.— Locality NMB 17288: Lopez section, Rio Yaque del Norte, Dominican Republic; Baitoa Formation, (late early to early middle Miocene) (Saun- ders, Jung, and Biju-Duval, 1986, p. 30, text-figs. 21, 25). Material.—Two lots with a total of only two speci- mens. Measurements.— height width h/w (mm) (mm) ratio NMB H 17323 (holotype) [PI. 23, figs. 12-15] 29.6 Pile 1.36 NMB H 17324 (paratype) [PIl. 23, figs. 16-18] 25.4 16.8 1.48 Remarks.—Only two specimens are available. The holotype is complete except for the extreme abapical portion of the outer lip; the paratype has the outer lip broken off the entire length of the aperture. Specimens are otherwise in good condition. Two living species of Massyla Adams and Adams, 1854 are known: C. (M.) corrugata Hinds, 1843 [p. 48; Hinds, 1844, p. 42, pl. 12, figs. 1, 2] and C. (M.) obtusa Deshayes, 1830 [p. 187]. A third possible Recent species is C. (M.) cumingiana Petit de la Saussaye, 1844 [pl. 112], although it is probably a junior subjective syn- onym of C. (M.) obtusa. These Recent species inhabit fairly shallow water in the southern part of the Pan- amic—Pacific Province. Massyla is represented in the Tertiary of the southeastern United States by a number of species, some still undescribed. Comparisons.—Cancellaria (Massyla) lopezana is closely related to C. (M.) jadisi Olsson, 1964 [p. 123, pl. 21, fig. 7] from the late Miocene Angostura For- mation of northwestern Ecuador but differs in having a greater whorl expansion rate, its shell being much larger although composed of the same number of whorls. Also, the Dominican species is constricted be- hind the siphonal fasciole whereas the siphonal fasciole of C. (M.) jadisi is much less prominent. It is also related to C. (M.) propevenusta Mansfield, 1929 [pl. 16, fig. 2; Mansfield, 1930, p. 47, pl. 17, fig. 2] of the Pinecrest Formation of Florida, which has more rap- idly enlarging whorls. The systematic importance, if any, of the adapical canal mentioned in the Description has not been established, but it also occurs in C. (AZ.) propevenusta. It has not been noted either in the Recent species or in C. (M.) jadisi. Occurrence.—L6pez section on Rio Yaque del Norte: Baitoa Formation (late early to early middle Miocene): NMB 17288, 16935 (Saunders, Jung, and Biju-Duval, 1986, p. 30, text-figs. 21, 25). Genus APHERA Adams and Adams, 1854 Aphera Adams and Adams, 1854, p. 277. Type species (by monotypy).— Cancellaria tessellata Sowerby, 1832a. Recent, Panamic—Pacific Province. Diagnosis.—Shell small. Shape elliptical, with an elongate, narrow aperture. Outer lip thickened and denticulate, lirate internally. Columella with two folds, the adapical one being larger and somewhat bifid or shelved in adults. A pronounced shield-like callus usu- ally present. Sculpture of evenly spaced spiral cords and axial ribs which form nodes at their intersections. Siphonal canal short but well-defined. Non-umbilicate. Remarks.—Aphera is known only from the Tertiary of Florida, the Caribbean, Central America, and north- western South America. Species from the later Tertiary of Italy placed in Aphera by Sacco (1894, pp. 66, 67) do not belong there. Formerly considered to be a paciphile genus (Woodring, 1966, p. 428; Vermeij, 1978, p. 232), a living species, A. lindae Petuch, 1987 [p. 109], has recently been discovered (Petuch, 1981, p. 333; 1987, p. 109) off Barbados. Golfo de Triste, Venezuela, the locality originally given for the only known specimen of Aphera lindae, is incorrect. The correct locality is 200 m depth off St. James, Barbados (Petuch, 1988, footnote on p. 160). This species from Barbados and the type species of the genus are the only living species of Aphera known. 110 BULLETIN 334 Aphera islacolonis (Maury, 1917) Plate 24, figures 1-9, 14-17; Plate 25, figures 1-8; Plate 26, figures 8-11; Text-figure 17 Cancellaria tessellata Sowerby. Gabb, 1873, p. 236. Not of Sowerby, 1832a. Cancellaria (Aphera) islacolonis Maury, 1917, p. 65, pl. 10, figs. 12, 12a. b; Olsson, 1922, p. 86, pl. 6, fig. 12; Ramirez, 1956, pp. 19, 22, pl. 2, figs. 4, 5; pl. 4, figs. 1, 2. Cancellaria ellipsis Pilsbry, 1922, p. 333, pl. 22, figs. 8, 9. {phera islacolonis (Maury). Perrilliat, 1973, p. 27, pl. 12, figs. 3-8. Not Aphera islacolonis (Maury). Woodring, 1970, p. 344, pl. 56, figs. 1, 2. (= unnamed species). Not Aphera islacolonis (Maury). Petuch, 1981, p. 333, figs. 81, 85, 86. (= Aphera lindae Petuch, 1987). Description.—Protoconch of just over two smooth, erect volutions. Elongate, elliptical teleoconch of about five whorls. Spiral sculpture of fine cords begins with onset of teleoconch, formation of axial ribs following shortly. The evenly spaced axial ribs, crossed by equal- ly evenly spaced spiral cords which form small nodes at the intersections, make the ornamentation finely cancellate. Spire high, often slightly concave. Aperture elongate, narrow. Outer lip thickened, most of the thickening occurring inside the aperture. Edge of outer lip with denticulations corresponding to the spiral cords. Inside of outer lip with about ten heavy, short lirations which reach only a short distance into the aperture. Near the adapical portion of the lip, one of these inner lirations is larger than the others. Columella with two strong, almost horizontal folds, the adapical one being larger, both incipiently bifid. Short secondary folds and pustules sometimes present on the inductura. Colu- mellar callus shield-like, covering most of the apertural side of the shell, the edge extending as a shelf, some- times everted abaxially. Siphonal canal short, well- defined. No umbilicus. Lectotype (herein selected).— PRI 28960. This spec- imen has been figured by Maury (1917, pl. 10, fig. 12) and is refigured here (PI. 24, figs. 1-3). Two paralec- totypes (PRI 28669, 28670) have also been figured by Maury (1917, pl. 10, figs. 12a, b) and are refigured here (Pl. 24, figs. 4-5). The incomplete specimen here se- lected as lectotype is the only one of Maury’s figured syntypes which has definite locality data. Dimensions of lectotype.—Height, 14.9 mm; width, 8.6 mm. Type locality.—Maury’s Bluff 2, on Rio Mao, Do- minican Republic; Cercado Formation (late Miocene) (Saunders, Jung, and Biju-Duval, 1986, p. 32, text-figs. 29, 30, table 3). Woodring (1970, p. 344) did not select a lectotype of A. islacolonis, but he designated Maury’s Bluff 3 as the type locality. As the lectotype was col- lected from Bluff 2, Woodring’s designation is sup- planted. Material.—This abundant species is represented in our collections by 64 lots containing over 1,300 spec- imens. The largest lots are from locality NMB 16923 (Cercado Formation of Arroyo BajOn on Rio Mao): 235 specimens, and from locality NMB 15903 (up- permost Cercado Formation of the Rio Gurabo sec- tion): 199 specimens. Measurements.—The measurements of 74 speci- mens are plotted in Text-figure 17. Remarks.—There are two distinct spire shapes in Aphera islacolonis, the prevalent shape being high- spired and almost, or slightly, concave (see PI. 24, figs. 1-4; Pl. 24, figs. 14-17; Pl. 25, figs. 1-8). The other, less common, form is “normal” or slightly convex, giving the entire shell a bullet-like shape (see Pl. 24, figs. 5-7). Both forms were figured by Maury (1917, pl. 10). Pilsbry (1922, pl. 22) had one specimen of each form when he described C. e/lipsis (see Pl. 24, figs. 6- 9). Perrilliat (1973, pl. 12, figs. 3-8) figured both forms from the Agueguexquite Formation of Mexico. That all three of these authors should have both forms is surprising, as in the Dominican Republic material, which we have examined, less than 1% of the speci- mens are of the “bullet’”” shape. These few specimens came from only six localities, with a maximum number of three specimens from one locality. At only two lo- calities was there a mixture of forms, and in both of these lots there was only one specimen of each form. 20 19 height in mm e 18 e = 17 11 10 0 5 6 7 8 9 10 11 12 width in mm Text-figure 17.—(Restored) height/width diagram of Aphera Is- lacolonis (Maury). DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 111 The Dominican Republic specimens of A. islacolonis are variable in sculpture, with some being more finely | sculptured than others. Also, some specimens are much ‘larger than others, with coarser sculpture in which the | axial ribs are sharp and prominent. All specimens in individual lots are remarkably uniform, but we find no basis for distinguishing these minor sculptural dif- ferences. The Recent Aphera lindae Petuch, 1987 [pl. 13, fig. 11; Petuch, 1981, figs. 81, 85, 86 (as A. islacolonis); Petuch, 1988, pl. 38, figs. 1, 2] is known from a single specimen. It has a “normal” spire which is neither concave nor convex, and is intermediate between the two forms discussed above. The width of this specimen is less than half its height, making it slimmer than any adult specimens of 4. is/acolonis which we have ex- amined. Aphera tessellata (Sowerby, 1832a) [p. 51] also has a slender shell. Approximately one-third of the specimens of A. is- lacolonis have been drilled, indicating that the species was subject to extensive naticid predation. Comparisons.—The specimen from the middle part of the Gatun Formation (late Miocene) figured by Woodring (1970, pl. 56, figs. 1, 2) is separable from A. islacolonis by its much thicker shell, its larger and heavier columellar folds, and coarser sculpture. Ols- son’s (1922, p. 86) reference to the occurrence of 4. islacolonis in the ““Gatun Formation” is omitted from our distribution records as it probably refers to the unnnamed species figured by Woodring, the only species of Aphera known to us from the Gatun Formation. Aphera wigginsi (Emerson and Hertlein, 1964) [p. 362, figs. Sd, e] from Pleistocene deposits of Isla Mon- serrate, Gulf of California, Mexico, was differentiated from A. islacolonis on the basis of its slender outline, the thicker parietal callus, the lack of grooving on the columellar plications, and in lacking a denticle on the upper portion of the columellar wall. The unique ho- lotype is much larger than any specimen of A. is/aco- lonis we have examined, but its height/width ratio does not serve to distinguish it. It does differ from A. is/a- colonis in the other characters mentioned, although few specimens of A. is/acolonis have a denticle on the adap- ical portion of the columella. The most obvious dis- tinction of A. wigginsi is its smooth, appressed, colu- mellar callus, which is not everted at the edges. Aphera peruana Nelson, 1870 [p. 190, pl. 6, fig. 3] from the Lower Zorritos Formation (early Miocene ?) and Tumbes Formation (late Miocene) of northern Peru, differs from A. is/acolonis in having much coarser sculpture and a fairly smooth, heavy, non-everted col- umellar callus. It also has a strong denticle on the adap- ical portion of the columella, and the adapical colu- mellar fold is broadly shelved. The holotype of A. peruana is refigured herein (Pl. 24, figs. 10-13). The living A. tessellata (Sowerby, 1832a) from the Panamic—Pacific Province differs from all other species of Aphera in being slimmer and in lacking a well-de- veloped columellar callus. Occurrence.—This species is recorded from the fol- lowing areas (from west to east): Rio Cana: upper part of Cercado Formation (late Miocene): TU 1230, 1282, and NMB 16835, 16837, 16838, 16839, 16842, 16844, 16857, 16986. Early Pliocene part of Gurabo Formation: TU 1354and NMB 16817, 16818, 16866. A single specimen (loc. NMB 16872) was collected from the base of the Mao Adentro Limestone Member of the Mao Formation (early Plio- cene)(Saunders, Jung, and Biyu-Duval, 1986, text-figs. 15, 16). Rio Gurabo: upper part of Cercado Formation (late Miocene): TU 1358, 1359, 1373, 1377, 1419, and NMB 15896, 15900, 15903, 15904, 15906, 15907, 15909, 15910, 15911, 15912, 15913, 15914, 15915, 15916. Late Miocene part of Gurabo Formation: TU 1210, 1211, 1212, 1231, and NMB 15806, 15807, 15871, 15878, 15882 (Saunders, Jung, and Byyu-Duval, 1986, text-figs. 4-6). Rio Mao: Cercado Formation (late Miocene): TU 1294 and NMB 16912, 16913, 17269 (all correspond to Bluff 3 of Maury); NMB 16915, 16916, 16917, 16918, 16922, 16923, 16924, 16926, 16927, 16928 (all Arroyo Bajon); NMB 16930 (= Bluff 2 of Maury) (Saunders, Jung, and Biju-Duval, 1986, text-figs. 29, 30, table 3). Arroyo Zalaya: early Pliocene: TU 1227A (Saunders, Jung, and Biju-Duval, 1986, p. 34, text-fig. 36, table 3). Rio Yaque del Norte: Baitoa Formation (late early to early middle Miocene): NMB 17286 (Lopez section); early Pliocene: TU 1403, 1405 (near La Barran- ca)(Saunders, Jung, and Biju-Duval, 1986, text-figs. PD |, P25). Rio Verde: N. 18, early Pliocene according to Akers (in Vokes, 1989): TU 1250 (for location, see Saunders, Jung, and Biju-Duval, 1986, text-fig. 38). Distribution.—Costa Rica: Uscari Formation (late Miocene ?). Mexico: Agueguexquite Formation (Plio- cene). Dominican Republic: Baitoa Formation (late early to early middle Miocene), Cercado and Gurabo Formations (late Miocene and early Pliocene), Mao Adentro Limestone (early Pliocene). Genus PERPLICARIA Dall, 1890 Perplicaria Dall, 1890, p. 90. Type species (by monotypy).—Perplicaria perplexa Dall, 1890. Caloosahatachee Formation (Pliocene), Florida. Daguinia Magne, 1966, p. 127. 112 BULLETIN 334 Type species (by monotypy).—Daguinia vigneauxi Magne, 1966. Burdigalian (late early Miocene), France. Diagnosis.—Shell small with rapidly enlarging con- vex whorls. Suture impressed. Aperture narrowly ovate, constricted adapically and expanded abapically. Outer lip thickened into a varix, internally lirate. Stromboid notch absent. Sculpture finely cancellate. Columella oblique but fairly straight, with three folds, the adapical one largest. The abapical fold borders the short si- phonal canal and is sometimes joined by the central fold. Columellar callus small but prominent. No um- bilicus. Remarks.—Originally described in the Mitridae, Perplicaria was first shown to be cancellariid by Wilson (1948). With the sole exception of the type species, specimens of Perplicaria are extremely rare. The only Recent species, P. clarki Smith, 1947 [p. 55], is found intertidally from Jalisco, Mexico to Ecuador, but is very rare. Daguinia has not previously appeared in the syn- onymy of Perplicaria, but we see little reason for sep- aration as the French species appears to differ only in being more attenuate and in having a structure on the adapical portion of the columella. The exact nature of this structure cannot be determined from the poor pub- lished figure, and the type has not been located, but it is probably a fusion of the adapical and central colu- mellar folds, resulting in a shelf-like structure. The known species of Perplicaria are: P. vigneauxi (Magne, 1966), Miocene, France. P. prior Maury, 1910, Chipola Formation, late early Miocene, Florida. P. canae, n. sp., Cercado Formation (late Miocene), Dominican Republic. unnamed species of Woodring, 1928, pl. 13, fig. 2, Bowden Formation (early Pliocene), Jamaica. P. perplexa Dall, 1890, Caloosahatchee Formation (Pliocene), Florida. P. clarki Smith, 1947, Recent, Panamic—Pacific Prov- ince. Perplicaria canae, new species Plate 25, figures 9-15; Plate 26, figures 12-14 Etymology of name.—Named after Rio Cana. Description.—Protoconch smooth, consisting of a little more than one volution; its outer lip prosocline. Four teleoconch whorls. Profile of teleoconch whorls nvex. First teleoconch whorl sculptured by four spi- ral cords. Just before the completion of the first teleo- conch whorl secondary spiral threads are introduced, and slightly prosocline axial riblets begin to appear. At the intersection of the spiral and axial sculptural ele- ments there are small knobs. Growth lines between the axial riblets are sometimes accentuated. Later spire | whorls gradually become proportionally higher, and | the number of spiral cords therefore increases as well. © On average the spiral cords are more prominent than — the axial riblets. Outer lip somewhat flaring near base, thickened, its inner surface with numerous lirae. Base — of aperture rounded. Columella with three oblique folds decreasing in size abapically. Columellar callus prom- inent; parietal callus less prominent. Holotype.—NMB H 17330 (PI. 25, figs. 9-11). Dimensions of holotype.—Height, 14.4 mm; width, 5.5 mm. Type locality.—Locality TU 1230: Rio Cana, east bank, just above the ford at Caimito on Los Quema- dos—Sabaneta Road (= loc. USGS 8534; Maury’s Zone H). Cercado Formation (late Miocene) (Saunders, Jung, and Biyu-Duval, 1986, p. 65, text-figs. 15, 16). Material.—Perplicaria canae is based on only two specimens: the holotype and one smaller paratype from the same locality. Measurements. — height width h/w (mm) (mm) ratio NMB H 17330 (holotype) [Pl. 25, figs. 9-11] 14.4 5.5 2.62 NMB H 17331 (paratype) [PI. 25, figs. 12-15] ES 4.4 297) Remarks.— Although slightly worn, both specimens of this species are well-preserved and appear to be adults, as they possess both a columellar callus and a thickened outer lip. The scarcity of material does not allow comments on variability. Comparisons.— Perplicaria canae is certainly closely related to P. perplexa Dall, 1890 from the Pliocene Caloosahatchee Formation of Florida. Perplicaria per- plexa is considerably larger than P. canae and its sculp- ture is coarser. On average, however, the axial riblets of P. canae are more accentuated. The protoconch of P. canae consists of a little more than one volution, whereas that of P. perplexa consists of one-and-one- half volutions. Occurrence.—Known only from TU 1230, the type locality of the species: Cercado Formation (late Mio- cene) of the Rio Cana section. Distribution.—Not known from outside the Domin- ican Republic. Genus TRIGONOSTOMA Blainville, 1827 Trigona Perry, 1811, expl. to pl. 51. Not Trigona Jurine, 1807 (Hymenoptera). Type species (by monotypy).—Trigona pellucida Perry, 1811 (= Buccinum scalare Gmelin, 1791). Re- cent, Indo-Pacific. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 13 Trigonostoma Blainville, 1827, p. 652. Type species (by monotypy).—Delphinula trigonos- toma Lamarck, 1822 (= Buccinum scalare Gmelin, 1791). Recent, Indo-Pacific. Subgenus VENTRILIA Jousseaume, 1887 Ventrilia Jousseaume, 1887, p. 194. Type species (by monotypy).—Ventrilia ventrilia Jousseaume, 1887 (= Cancellaria tenera Philippi, 1848). Recent, Caribbean Province. Arizelostoma Iredale, 1936, p. 318. Type species (by original designation). —Arizelosto- ma laseroni Iredale, 1936. Recent, Australia. Emmonsella Olsson and Petit, 1964, p. 541. Type species (by original designation).— 7rigonos- toma tenerum (Philippi, 1848). Recent, Caribbean Province. Diagnosis.—Shell of medium size, wide, openly and deeply umbilicate. Shoulder usually wide, angled, slop- ing down to the suture. Aperture trigonal. Columella with two descending folds, sometimes with a third weaker fold forming the edge of the short, notch-like anterior canal. Remarks.—The subgenus Ventrilia occurs in the lat- er Tertiary of Europe, southeastern United States, the Caribbean, northern South America and Central America, and in the Recent faunas of Australia and the Caribbean and Panamic—Pacific Provinces. Species of Ventrilia seem to be separable into two forms (lineages ?). One group is typified by a wide shell with a sloping columella and primarily spiral sculpture such as the Recent Caribbean species 7. (V.) tenerum (Philippi, 1848) [p. 24] and the living Eastern Pacific species 7. (V.) tuberculosum (Sowerby, 1832a) [p. 51]. The other group has an almost vertical columella and has spiral and axial sculpture with some of the axial ribs developed into varices as in the living Eastern Pacific species 7. (V.) breve (Sowerby, 1832a) [p. 52] and 7. (V.) goniostoma (Sowerby, 1832a) [p. 51], and T. (V.) gurabis (Maury, 1917) [p. 65, pl. 10, fig. 11] from the late Miocene part of the Gurabo Formation, Dominican Republic. Trigonostoma (Ventrilia) gurabis (Maury, 1917) Plate 27, figures 1-12; Plate 29, figures 1-4 Cancellaria (Trigonostoma) gurabis Maury, 1917, p. 65, pl. 10, fig. ie Description.— Large deviated protoconch of a little less than one-and-one-half volutions. Teleoconch whorls, about four in number, rounded with a pro- nounced excavated shoulder. Axial sculpture of weak prosocline ribs and fine growth lines with irregularly spaced varices. Spiral sculpture of strong cords, about nine on the body whorl, with weaker spiral cords mid- way of each interspace, and even weaker spiral cords between the primary and secondary cords. Aperture trigonal, outer lip lirate within. Columella almost ver- tical, with two strong descending folds and a third weaker fold forming the edge of the short notch-like anterior canal. Umbilicus deep. Holotype.—PRI 28668. This was apparently the only specimen Maury had when she described this species. It is refigured here (see Pl. 27, figs. 1—4). Dimensions of holotype.— Height, 11.0 mm; width, 8.6 mm. Type locality.—Rio Gurabo at Los Quemados, Do- minican Republic; Maury’s Zone D. Late Miocene part of Gurabo Formation. Material.—Two lots with a total of 10 specimens. Measurements. — height width h/w (mm) (mm) ratlo PRI 28668 (holotype) [PI. 27, figs. 1-4] 11.0 8.6 1.28 NMB H 17332, from loc. TU 1215; [Pl. 27, figs. 5-8] 11.4 9.0 E27 NMB H 17333, from loc. TU 1215; [Pl. 27, figs. 9-12] 13.2 9.6 1.38 unnumbered specimen, from loc. NMB 15848 11.6 8.7 1.33 unnumbered specimen, from loc. TU 1215 13.0 9.2 1.41 unnumbered specimen, from loc. TU 1215 8.0 6.3 V27 Remarks.— About half of the available material con- sists of complete, well-preserved adult specimens. Comparisons.— Trigonostoma (Ventrilia) gurabis 1s more closely allied to the Recent Panamic—Pacific species 7. (V.) breve (Sowerby, 1832a) than to the only living Caribbean species of Ventrilia, T. (V.) tenerum (Philippi, 1848). It differs from 7. breve in its much stronger spiral sculpture and in having more varices. It differs from 7. (V.) tenerum in having an almost vertical columella and axial sculpture forming varices. Occurrence.— Rio Gurabo: late Miocene part of Gur- abo Formation: TU 1215 and NMB 15848 (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4-6). Distribution.— Not known from outside the Domin- ican Republic. Trigonostoma (Ventrilia ?) insulare (Pilsbry and Johnson, 1917) Plate 27, figures 13-15 Cancellaria brevis Sowerby. Gabb, 1873, p. 236. Not of Sowerby, 1832a. Cancellaria (Trigonostoma) insularis Pilsbry and Johnson, 1917, p. 163; Pilsbry, 1922, p. 334, pl. 22, fig. 11. 114 Description.—Protoconch smooth, of about one-and- one-half volutions. Rapidly enlarging teleoconch of about five whorls, rounded at the periphery but with a rounded shoulder behind which there is a channel at the suture. Sculpture of about 14 weak spiral cords, sometimes with weaker spiral threads in the inter- spaces. Axial sculpture of rounded collabral prosocline ribs, about 12 on the body whorl. Aperture trigonal, outer lip heavy and lirate within. Columella sloping with two strong descending folds and a third fold form- ing the edge of the shallow anterior canal. Numerous tubercules present on the edge of the columella between the folds. Adapical portion of aperture with shaliow posterior canal, well-defined by lirations below the shoulder. Umbilicus deep. Holotype.—ANSP 2989 (Pl. 27, figs. 13-15). Dimensions of holotype.— Height, 24.2 mm; width, 21.0 mm. Type locality.—Santo Domingo, exact locality and stratigraphic position unknown. Material.—Known only from the holotype. Remarks.—The concave columella of this species and its sharply defined posterior canal below the shoul- der make placement in Ventrilia questionable. This species was described from a specimen in the Gabb collection and no precise locality data are avail- able. No additional specimens have been located. Woodring (1970, p. 345) lists a specimen from the Gatun Formation as 7. cf. insulare. A specimen in the Petit collection, from the lower part of the Gatun For- mation (late Miocene), is presumed to be the same species referred to by Woodring. This unnamed species is similar to 7. insulare, differing in being less umbil- icate, higher spired, and having a sharp shoulder. Trigonostoma (Ventrilia ?) insulare closely resem- bles 7. smithfieldensis Oleksyshyn, 1960 [p. 101, figs. 1, 2] from the Yorktown Formation (early Pliocene) of Virginia. Occurrence.— Not known (see above). It is quite pos- sible that this species does not occur in the Neogene of the Dominican Republic. Distribution.—As this species is known only from a single specimen from an unknown locality, its distri- bution is not known. Genus AXELELLA Petit, 1988 Olssonella Petit, 1970, p. 83. Not Olssonella Glibert and Van de Poel, 1967, p. 121. Type species (by original designation). — Cancellaria smithii Dall, 1888. Recent, North Carolina to Vene- zuela. ixelella Petit, 1988, p. 130 (new name for Olssonella Petit, 1970 non Glibert and Van de Poel, 1967) BULLETIN 334 Type species (by original designation of Olssonella Petit, 1970).—Cancellaria smithii Dall, 1888. Recent, North Carolina to Venezuela. Diagnosis.—Shell small, scalate, with a conical spire and rounded anterior. Teleoconch whorls rounded with — an impressed suture. Sculpture of spiral cords and pro- socline ribs. Columella almost vertical with two folds; a third weak fold sometimes forming the edge of the short siphonal canal. Degree of umbilication variable. Remarks.—Axelella is a compact genus of cancel- lariids which has been recognized in Miocene and later formations of the southeastern United States, Califor- nia, the Caribbean, and Central America, and Recent off the southeastern coast of the United States, the Gulf of Mexico, Venezuela, and in the Panamic—Pacific Province. One species occurs in the Recent fauna of the eastern Atlantic. The animal of A. smithii (Dall, 1888) [p. 70, fig. 292], the type species of the genus, has been described by Harasewych and Petit (1984). Axelella emblema, new species Plate 27, figures 16-19 Etymology of name.—Gr. emblema = inlaid work. Description.—Shell scalate. Protoconch smooth, of | about one-and-one-half volutions. Teleoconch whorls, about four in number, sculpture with seven to nine strong collabral ribs crossed by about 10 evenly spaced — spiral cords which rise over the axial ribs, usually with a single weaker spiral thread in each interspace. Ap- erture roughly trigonal, being rounded at the shoulder. Suture slightly impressed. Outer lip with about 12 in- terior lirations which extend well into the aperture. Columella with two broad horizontal folds, the adap- ical one being larger, and a barely perceptible third oblique fold forming the edge of a short siphonal canal. Umbilicus chink-like. Holotype.—NMB H 17335 (PI. 27, figs. 16-19). Dimensions of holotype.—Height, 10.7 mm; width, 7.1 mm. Type locality.—Locality NMB 16942: Lopez section on Rio Yaque del Norte, Dominican Republic; Baitoa Formation (late early to early middle Miocene) (Saun- ders, Jung, and Biju-Duval, 1986, p. 30, text-figs. 21- 25). Material.—Known only from the holotype and two paratypes. Measurements. — height width h/w (mm) (mm) ratio NMB H 17335 (holotype) (Pl. 27, figs. 16-19] 10.7 Wen LES NMB H 17336 (paratype) 9.3 5.7 1.63 NMB H 17339 (paratype) 8.2 Sez 1.58 DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 115 Remarks.—A\ll three available specimens are well- preserved, but probably immature. None of the three have protoconchs preserved well enough to be used for SEM photography. Recent species of Axelella live at depths from 20 to 110 m. Comparisons. —Axelella emblemais closer to the liv- ing A. smithii (Dall, 1888) than to any of its Caribbean Tertiary congeners, and differs from it in being pro- portionally broader with a much less impressed suture and in having coarser primary spiral cords. Both 4. thisbe (Olsson, 1964) [p. 126, pl. 22, fig. 6] from the Esmeraldas Formation (early Pliocene) of Ecuador and A. scalatella (Guppy, 1873) [p. 78, pl. 2, fig. 4] from the Bowden Formation (early Pliocene) of Jamaica have much finer spiral sculpture and deeply impressed su- tures. Axelella panamica (Petit, 1976) [p. 35, pl. 2, fig. 1] from the lower part of the Gatun Formation (late Miocene) of Panama has very prominent axial ribs with spiral cords that become prominent only where they cross the axial ribs. Occurrence.— Known only from locality NMB 16942: Lopez section on Rio Yaque del Norte, Baitoa For- mation (late early to early middle Miocene) (Saunders, Jung, and Biju-Duval, 1986, p. 30, text-figs. 21, 25). Distribution.—Not known from outside the Domin- ican Republic. Genus AGATRIX Petit, 1967 Agatrix Petit, 1967, p. 218. Type species (by original designation).—77rigono- stoma agassizii Dall, 1889. Recent, North Carolina to the Gulf of Mexico. Diagnosis.—Shell small, tabulate. Sculpture cancel- late. Suture impressed behind a rounded shoulder. Ap- erture ovate-trigonal, lirate within. Stromboid notch distinct. Columella sloping or concave with three folds, the abapical one forming the edge ofa short but distinct siphonal canal. A thin parietal callus becomes shield- like abapically and extends over the umbilical chink. Remarks.—Several Indo-Pacific Recent and Ter- tiary species have been referred to Agatrix in the past few years, but their true relationship to the species of the Caribbean and Panamic—Pacific provinces has not been determined. Agatrix losquemadica (Maury, 1917) Plate 28, figures 1-10; Plate 29, figures 5-8; Text-figure 18 Cancellaria (Narona) losquemadica Maury, 1917, p. 66, pl. 10, fig. 13. Cancellaria (Tribia ?) losquemadica Maury. Marks, 1949, p. 460. Description.—Pronounced protoconch smooth, of about one-and-one-quarter volutions. Teleoconch of about six whorls. On the first teleoconch whorl spiral cords appear a little before the axial riblets. Shell tab- ulate with deeply impressed suture behind a rounded shoulder. Sculpture of axial ribs, about 10 on body whorl, sometimes enlarged as varices, crossed by about 15 spiral cords. Spiral cords evenly spaced except for crowding on the shoulder, with occasional weaker spi- ral threads in interspaces. Spiral cords not present on inner half of shoulder. Aperture ovate-trigonal. Outer lip lirate within, with a noticeable stromboid notch in adults. Columella sloping, with three distinct folds, the abapical one forming the edge of a short but distinct anterior canal. A thin parietal callus, shield-like abap- ically, extends over the umbilical chink. Holotype.— PRI 28671. This was apparently the only specimen Maury had when she described this species. It is refigured here (see Pl. 28, figs. 1-3). Dimensions of holotype.— Height, 12.8 mm; width, 7.6 mm. Type locality.—Rio Gurabo at Los Quemados, Do- minican Republic; Maury’s Zone E. The type locality is here restricted to locality NMB 15863 which falls in the lower part of the Gurabo Formation (late Miocene) (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4, 6). Material.—Ten lots with a total of 16 specimens. Measurements.— The measurements of 10 complete specimens are plotted in Text-figure 18. Remarks.—Well-preserved adult specimens consti- tute only about half of the available material. There is height in mm 0 5 6 7 8 9 width in mm Text-figure 18.—(Restored) height/width diagram of Agatrix /os- quemadica (Maury). 116 BULLETIN 334 some variability among the specimens, a few having slightly weaker axial ribs which are not sharply round- ed at the shoulder. Comparisons.—Agatrix losquemadica, while related to A. epomis (Woodring, 1928) [p. 223, pl. 12, fig. 10] of the Recent fauna of Venezuela and the Bowden Formation (early Pliocene) of Jamaica, and to A. bea- trix (Olsson, 1964) [p. 128, pl. 22, fig. 9] from the Esmeraldas Formation (early Pliocene) of Ecuador, dif- fers from both species in having a greater number of spiral threads making the shell more finely sculptured. The living Western Atlantic species A. agassizii (Dall, 1889) [p. 130, pl. 35, fig. 4], the type species of the genus, and the Panamic—Pacific species A. strongi (Shasky, 1961) [p. 19, pl. 4, fig. 4] also have less prom- inent axial ribs and less deeply impressed sutures. Occurrence.—This species has been found in several areas (for locations see Saunders, Jung, and Biju-Du- val, 1986, text-figs. 4-6, 15, 16, 38): Rio Gurabo: late Miocene part of Gurabo Forma- tion: TU 1296 and NMB 15804, 15814, 15863, 15869. Rio Cana: upper part of Cercado Formation (late Miocene): NMB 16838, 16854: Gurabo Formation (late Miocene and early Pliocene): NMB 16821, 16866. Rio Verde: N. 18, early Pliocene according to Akers (in Vokes, 1989): TU 1250. Distribution.—Not known from outside the Domin- ican Republic. Genus ADMETULA Cossmann, 1889 Admetula Cossmann, 1889, p. 228. Type species (by original designation). — Cancellaria evulsa (Solander), (= Buccinum evulsum Solander, 1766). Eocene, England. Diagnosis.—Shell small, rounded, non-tabulate. Ap- erture broadly ovate, lirate within. Columella exca- vated with two strong descending folds and a third fold which forms the edge of the short but distinct anterior canal. Sculpture cancellate. Varices usually present at irregular intervals. Columellar callus not developed into a shield. No umbilicus. Remarks.—Two Caribbean and one Galapagan Re- cent species have been cited as Admetula, but they lack varices and may possibly be improperly placed in this genus. These species are: {. bayeri Petit, 1976 [p. 38, pl. 1, fig. 4], Gulf of Mexico; 1. vossi Petit, 1976 [p. 39, pl. 1, fig. 5], Bahamas; A. deroyae (Petit, 1970) [p. 85, pl. 1, figs. 3a, b], Ga- lapagos Islands. Admetula zalayana, new species Plate 29, figures 9-12 Etymology of name.—Named after Arroyo Zalaya, Dominican Republic. Description.—Protoconch large, of about one-and- one-half volutions, the first smooth, the last one-half with weak spiral threads. Postnuclear whorls begin abruptly with axial ribbing. Teleoconch of about five rounded whorls with deeply impressed suture. Sculp- ture of axial ribs, about 10 on body whorl, crossed by evenly spaced primary spiral cords, about 10 on the body whorl, with one or two secondary spiral threads in the interspaces. Axial ribs sometimes pronounced as irregularly spaced varices, the strongest varices on some specimens being at 90° intervals. Aperture ovate, lirate within. Columella excavated with two strong de- scending folds and a third fold which forms the edge of the short but distinct anterior canal. One or more tubercules sometimes present between the extremities of the columellar folds. No umbilicus. Holotype.—NMB H 17343 (PIL. 29, figs. 9-12). Dimensions of holotype.—Height, 11.2 mm; width, 7.7 mm. Type locality.—Locality TU 1227: Arroyo Zalaya; early Pliocene (Saunders, Jung, and Biju-Duval, 1986, p. 34 and text-fig. 36). Material.—Two lots with a total of only five speci- mens, all but one in an excellent state of preservation. Measurements.— height width h/w (mm) (mm) ratio NMB H 17343 (holotype) [Pl. 29, figs. 9-12] 11.2 7.7 1.45 unnumbered specimen, from loc. TU 1227 11.1 7.6 1.46 unnumbered specimen, from loc. TU 1227 8.9 5.7 1.56 unnumbered specimen, from loc. OM 227, 11.4 7.9 1.45 Remarks.— As mentioned above, the Recent species now placed in Admetula may represent a separate lin- eage as they lack varices. The two Caribbean species live at depths in excess of 500 m (Petit, 1976, pp. 38, 39) while the Galapagan species occurs at 150 m (Petit, 1970, p. 85). Comparison.—Admetula zalayana differs from A. zapoteca (Bose, 1910) [p. 240, pl. 13, fig. 17], which occurs in beds of probably late Miocene or early Plio- cene age of the Isthmus of Tehuantepec, in being pro- portionally wider with heavier sculpture and more pro- nounced varices. There are no other similar varicate species known from the Recent or Tertiary faunas of the Caribbean area. Occurrence.—Arroyo Zalaya: early Pliocene: TU 1227. Rio Verde: N. 18, early Pliocene according to Akers DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 117 (in Vokes, 1989): TU 1250. (For location see Saunders, Jung, and Biju-Duval, 1986, text-figs. 36, 38.) Distribution. — Not known from outside the Domin- ican Republic. APPENDIX In this Appendix we briefly treat species from the Tertiary Caribbean faunal Province that have been mistakenly identified with Dominican Republic species. Cancellaria (Cancellaria) barretti Guppy, 1866 Plate 16, figures 1-4 Cancellaria barretti Guppy, 1866, p. 289, pl. 17, fig. 11; Guppy, 1867, p. 157 (list, in part); Guppy, 1874, p. 438 (list, in part); Dall, 1903, p. 1583 (list, in part). Not Cancellaria barretti Guppy. Guppy, 1866, p. 286 (list; not p. 289); Guppy, 1867, p. 157 (list, in part); Guppy, 1876, p. 520; Maury, 1917, p. 226, pl. 36, fig. 1; Pilsbry, 1922, p. 332; Maury, 1925b, pl. 9, fig. 17; Ramirez, 1950, p. 22, pl. 3, fig. 3; Ramirez, 1956, pp. 12, 18, 19. (all = Cancellaria mauryae Olsson, 1922). Not Cancellaria barretti Guppy. Engerrand and Urbina, 1910, p. 125. Not Cancellaria barretti Guppy. Olsson, 1922, p. 81, pl. 6, fig. 6. (?= Cancellaria dariena Toula, 1909). Cancellaria (Cancellaria) barretti Guppy. Woodring, 1928, p. 219, pl. 12, fig. 6. Not Cancellaria (Cancellaria) barretti Guppy. Tucker and Wilson, 1932, p. 8, pl. 3, fig. 3. Not Cancellaria barreti [sic] Guppy. Gomez P. and Valerio G., 1971, p. 44, fig. 3. (= C. petiti Olsson, 1967). Holotype.—BMNH G 64069 (PI. 16, figs. 1-4). Dimensions of holotype.—Height, 34.6 mm; width, 18.8 mm. Type locality.—Bowden, Jamaica. Bowden Forma- tion (early Pliocene). Remarks.—The Dominican Republic references to C. barretti are discussed under C. mauryae Olsson, 1922 [p. 95, herein]. Engerrand and Urbina (1910, p. 125) reported the species from Zuluzum, Chiapas, Mexico, on the basis of two incomplete specimens which were not figured. We agree with Woodring (1928, p. 220) that it is 1m- probable that their material represents C. barretti. The record by Olsson (1922, p. 81, pl. 6, fig. 6) from the Rio Banano, Costa Rica (Rio Banano Formation, early Pliocene) appears to be based on a specimen of C. dariena Toula, 1909. Woodring (1928, p. 219) was also of this opinion. The Floridian shell reported as C. barretti by Tucker and Wilson (1932, p. 8, pl. 3, fig. 3) is one of a number of variable forms of Cancellaria sensu stricto in the later Tertiary of Florida which remain to be treated comprehensively. For comments on the record given by Gomez P. and Valerio G. (1971, p. 44, fig. 3), see Remarks under C. petiti Olsson, 1967 [p. 117, herein]. As mentioned by Woodring (1928, p. 219) the only known specimens of C. barretti are the holotype (fig- ured herein; Pl. 16, figs. 1-4) and the specimen figured by Woodring (1928, pl. 12, fig. 6). This latter specimen has a more rounded body whorl than the holotype. Cancellaria (Cancellaria) petiti Olsson, 1967 Plate 16, figures 5-9 Cancellaria (Cancellaria) cossmanni Olsson, 1922, p. 81, pl. 6, figs. 9, 11. Not C. cossmanni Morlet, 1888, p. 209, pl. 9, figs. 10, 10a, b. ? Cancellaria (Cancellaria) cossmanni Olsson. Anderson, 1929, p. 117. Not Cancellaria (Cancellaria) cossmanni Olsson. Oinomikado, 1939, p. 623, pl. 29, fig. 17. (? = juvenile Distorsio Roding, 1798). Cancellaria (Cancellaria) petiti Olsson, 1967, p. 44 (new name for C. cossmanni Olsson, 1922 non Morlet, 1888). Cancellaria barreti {sic} Guppy. Gomez P. and Valerio G., 1971, p. 44, fig. 3. Not of Guppy, 1866. Cancellaria cossmanni Olsson. Gomez P. and Valerio G., 1971, p. 44, fig. 4. Lectotype (herein selected). — PRI 20966. This is the larger of the two specimens figured by Olsson (1967, pl. 6, fig. 9) and is refigured herein (PI. 16, figs. 5—9). Dimensions of lectotype.— Height, 25.7 mm, width, 14.6 mm. Type locality.— Rio Banano, Limon Province, Costa Rica. Rio Banano Formation (early Pliocene). Remarks.—This species is included as it has been confused with C. barretti Guppy, 1866, from the Bow- den Formation (early Pliocene) of Jamaica, which in turn has been reported from the Dominican Republic. See Remarks under C. barretti Guppy, 1866 and C. mauryae Olsson, 1922 [pp. 117 and 95 herein, re- spectively]. As may be seen from the illustrations herein, C. barretti and C. petiti are distinct, differing in overall shape as well as in ornamentation. Cancellaria petiti was reported (as C. cossmanni Ols- son) from near Cibarco, Colombia (Tubara group; late Miocene or early Pliocene) by Anderson (1929, p. 117) on the basis of a single specimen which he did not figure. This species was also reported from southwestern Colombia (Cucurrupi River; beds of late Miocene or early Pliocene age) by Oinomikado (1939, p. 623, pl. 29, fig. 17). His poor figure of a small shell is not recognizable, but appears to be of a juvenile Distorsio. It is certainly not C. petiti. Gomez P. and Valerio G. (1971, p. 44, figs. 3, 4) figured two specimens, identifying the larger as C. bar- reti [sic]. Their specimens are clearly conspecific with each other and with C. petiti. Cancellaria (Pyruclia ?) laevescens Guppy, 1866 Plate 18, figures 10-12 Cancellaria laevescens Guppy, 1866, p. 289, pl. 17, fig. 12; Guppy, 118 BULLETIN 334 1867, p. 157 (list, in part); Guppy, 1874, p. 438 (list, in part); Dall, 1903, p. 1583 (list, in part); Maury, 1920, p. 69 (in part). Not Cancellaria laevescens Guppy. Guppy, 1866, p. 286 (list; not p. 289): Guppy, 1867, p. 157 (list, in part); Gabb, 1873, p. 236 (list); Guppy, 1876, p. 520; Maury, 1917, p. 64, pl. 10, fig. 6; Pilsbry, 1922, p. 333. (all = Cancellaria (Pyruclia ?) uva, n. sp.). Not Cancelaria [sic] laevescens Guppy. Guppy, 1910, p. 6; Guppy, 1913, p. 4. (= Cancellaria auriculaperta Vokes, 1938). Not Cancellaria laevescens Guppy. Hubbard, 1920, p. 157, pl. 24, figs. 5, 6. (= Cancellaria laevescens portoricana Maury, 1920); Li, 1930, p. 272, pl. 8, fig. 63 (= C. bulbulus Sowerby, 1832a; Recent). Cancellaria(Cancellaria) laevescens Guppy. Woodring, 1928, p. 220, pl. 12, figs. 7, 8. Cancellaria (Cancellaria) lavescens [sic] Guppy. Marks, 1949, p. 460 (list). Lectotype.—BMNH G 64070. This specimen is the syntype figured by Guppy (1866, pl. 17, fig. 12) and refigured here (Pl. 18, figs. 10-12). We consider Wood- ring’s citation (1928, p. 221) of this specimen as “ho- lotype”’ to constitute lectotype designation. Dimensions of lectotype.— Height, 42.6 mm; width, 27.2 mm. Type locality.—Bowden, Jamaica. Bowden Forma- tion (early Pliocene). Remarks.—Cancellaria laevescens portoricana Mau- ry, 1920 [p. 69, pl. 7, fig. 10] was described from an artificial cast of an external mold. It was stated by Maury to resemble ““Gabb’s Dominican specimen of C. laevescens [= C. uva, n. sp.] but is still smaller and is only a third the size of Guppy’s type.’’ We consider the specimens referred to C. laevescens by Hubbard (1920, p. 157, pl. 24, figs. 5, 6) to be conspecific with C. portoricana. Both Maury’s and Hubbard’s material came from Quebradillas, Puerto Rico (late Miocene or early Pliocene). Cancellaria portoricana cannot be con- sidered a subspecies of C. /aevescens as available ma- terial does not permit this determination. The species from Springvale, Trinidad (early Plio- cene) listed as C. laevescens by Guppy (1910, p. 6; 1913, p. 4) was described as C. auriculaperta Vokes, 1938 [p. 22, figs. 19, 20]. All reports of C. laevescens from the Dominican Republic are considered to be for C. uva, n. sp., and are discussed thereunder [p. 101, herein]. Cancellaria laevescens appears to be restricted to the Bowden Formation of Jamaica (early Pliocene) as all reports of it from elsewhere are referable to other species. Cancellaria (Bivetopsia) moorei Guppy, 1866 Plate 22, figures 8-11 Cancellaria moorei Guppy, 1866, p. 289, pl. 17, fig. 7; Guppy, 1867, p. 157 (list, in part); Guppy, 1874, p. 438 (list, in part); Dall, 1903, p. 1583 (list, in part). Not Cancellaria moorei Guppy. Guppy, 1866, p. 286 (list; not page 289): Gabb, 1873, p. 236 (list); Guppy, 1876, p. 520. (all = Can- cellaria (Bivetiella) epistomifera Guppy, 1876). Cancellaria (Bivetopsia) moorei Guppy. Woodring, 1928, p. 222, pl. 12, fig. 9. Cancellaria moorei (?) Guppy. Anderson, 1929, p. 117. Cancellaria (Bivetopsia ?) moorei Guppy. Marks, 1949, p. 460 (list). Lectotype.—BMNH G 64068. This specimen is the syntype figured by Guppy (1866, pl. 17, fig. 7) and is refigured here (Pl. 22, figs. 8-11). We consider Wood- ring’s citation (1928, p. 222) of this specimen as “ho- lotype”’ to constitute lectotype designation. Dimensions of lectotype.— Height, 19.0 mm; width, 13.6 mm. Type locality.—Bowden, Jamaica. Bowden Forma- tion (early Pliocene). Remarks.—This species was confused with C. ep- istomifera Guppy by both Guppy and Gabb, and thus appeared in their lists of Dominican Republic taxa. The only Dominican species of Bivetopsia known to us is C. (B.) plectilis, n. sp. [see Remarks on p. 107, herein]. Cancellaria moorei was reported from near Usiacuri, Colombia (Tubara group; late Miocene or early Plio- cene) by Anderson (1929, p. 117) on the basis of a single specimen which he did not figure. This is the only report of the nominotypical subspecies from out- side of the Bowden Formation (early Pliocene) of Ja- maica. The subspecies C. (B.) moorei pachia Smith, 1940 [p. 45, pl. 2, fig. 2] from near Belle Glade. Florida (probably Bermont Formation; Pleistocene) differs from the nominotypical subspecies in being less attenuate and in having fewer but larger axial ribs. REFERENCES CITED Adams, H., and Adams, A. 1853-1854. The Genera of Recent Mollusca, vol. 1. London, pp. 1-484. Adanson, M. Histoire naturelle des coquillages du Sénégal. Paris, pp. i- xcvi, 1-275, 19 pls Anderson, F. M. 1929. Marine Miocene and related deposits of north Colombia. Proceedings of the California Academy of Sciences, ser. 4, vol. 18, No. 4, pp. 73-213, pls. 8-23. Ayres, W. O. 1855. [Description of a new species of Cramp Fish]. Proceedings of the California Academy of Natural Sciences, vol. 1, pp. 70-71. Barnard, K. H. 1959. Contributions to the knowledge of South African marine mollusca. Part II. Gastropoda: Prosobranchiata: Rhach- iglossa. Annals of the South African Museum, vol. 45, pt. 1, pp. 1-237, 52 text-figs. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 119 Blainville, H. M. D. de 1825-1827. Manuel de Malacologie et de Conchyliologie. Paris. 2 vols., 190 pls. [Text pp. 1-647 issued 1825; pp. 649- 664 and plates issued 1827]. Bose, E. 1910. Zur jungtertiaren Fauna von Tehuantepec. I. Stratigra- phie, Beschreibung und Vergleich mit amerikanischen Ter- tidrfaunen. Jahrbuche der Kaiserlich-KOniglichen Geo- logischen Reichsanstalt, vol. 60, pp. 215-255, pls. 12-13. Brocchi, G. B. 1814. Conchiologia fossile subapennina con osservazioni geolo- giche sugli Apennini. 2 vols., Milano, 712 pp., 16 pls. Cernohorsky, W. O. 1972. Marine Shells of the Pacific, Vol. II. Sydney, Australia, 411 pp., 68 pls. Cossmann, A. E. M. 1888. Gastéropodes, in Dagincourt, E., Annuaire géologique universel...., vol. 4, pp. 765-785. 1889. Catalogue illustré des coquilles fossiles de l’Eocéne des en- virons de Paris, tome 2, fasc. 4. Annales de la Société Royale Malacologique de Belgique, vol. 24, pp. 1-385, pls. 1- 12. 1899. Essais de Paléoconchologie comparée, fasc. 3. 201 pp., 8 pls. Paris. 1903. Essais de Paléoconchologie comparée, fasc. 5. 215 pp., 9 pls. Paris. 1913. Etude comparative de fossiles Miocéniques recueillis a la Martinique et a l'Isthme de Panama. Journal de Conchy- liologie, vol. 61, pp. 1-64, pls. 1-5. Dall, W. H. 1888. Mollusks, in Agassiz, A., Three Cruises of the United States Coast and Geodetic Survey Steamer Blake. vol. 2, ch. 8, pp. 62-75, figs. 282-312. Boston and New York. 1889. /Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico (1877-78) and in the Caribbean Sea (1879-80), by the U. S. Coast Survey Steamer Blake, ..../ XXIV. Report on the Mol- lusca. — Part II. Gastropoda and Scaphopoda. Bulletin of the Museum of Comparative Zoology, Harvard, vol. 18, pp. 1-492, pls. 10-40. 1890. Contributions to the Tertiary fauna of Florida. Pt. 1. Trans- actions of the Wagner Free Institute of Science, vol. 3. pt. 1, pp. 1-200, pls. 1-12. 1896. Diagnoses of new species of mollusks from the west coast of America. Proceedings of the United States National Museum, vol. 18, pp. 7-20. 1903. Contributions to the Tertiary fauna of Florida. Pt. 6. Trans- actions of the Wagner Free Institute of Science, vol. 3, pt. 6, pp. 1219-1654, pls. 48-60. 1908. /Reports on the dredging operations off the west coast of Central America to the Galapagos, to the west coast of Mexico, and in the Gulf of California... XIV]. The Mol- lusca and Brachiopoda. Bulletin of the Museum of Com- parative Zoology, Harvard, vol. 43, No. 6, pp. 205-487, pls. 1-22. Deshayes, G. P. 1830. Encyclopédie Méthodique. Histoire naturelle des vers. vol. 2, pp. 1-144. Paris. Emerson, W. K., and Hertlein, L. G. 1964. Invertebrate megafossils of the Belvedere Expedition to the Gulf of California. Transactions of the San Diego Society of Natural History, vol. 13, No. 17, pp. 333-368, figs. 1- 6. Engerrand, J., and Urbina, F. 1910. Primera nota acerca de la fauna Miocenica de Zuluzum (Chiapas). Boletin de la Sociedad Geologica Mexicana, vol. 6, pt. 2, pp. 119-140, pls. 58-60. Fisher, N., and Tomlin, J. R. le B. 1935. The dates of publication of Forbes and Hanley’s Hist. Brit. Moll. Journal of Conchology, vol. 20, No. 5, pp. 150-151. Forbes, E., and Hanley, S. 1848-1853. detice me saci eeia eer eevee eee 9-11. Lectotype. ANSP 3288. Exact locality not known; 9. front view; 10. apical view; 11. rear view. Height, 24.5 mm; width, 19.8 mm. x2. NMB H 17311; from locality NMB 16938: Lopez section on Rio Yaque del Norte; Baitoa Formation (late early to early 12-15. middle Miocene); front view; 13. rear view; 14. apical view; 15. from right side. Height, 24.2 mm; width, 19.8 mm. x2. PLATE 19 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 98 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 98 PLATE 20 4 x 40 5 x 80 6 x 30 10 x 32 11 x 65 x 35 Figure 1-3. 4-6. 7-9. 10-12. DOMINICAN REPUBLIC NEOGENE. 10: JUNG AND PETIT 129 EXPLANATION OF PLATE 20 Page Cancellaria (Cancellaria) guppyi Gabb, 1873 .......... 06.0 eee 94 NMB H 17295; from locality NMB ioe Rio Gurabo; upper part of Cercado Formation (late Miocene). Wore ciasesitine outer lip of protoconch; 1. protoconch, x 14; 2. transition protoconch/teleoconch, x 70; 3. apical view, x12. Cancellaria (Cancellaria) harrisi a OT prepreg e ene oon ee tvexnus SO eee aerclok © wie henasbsisyjevathere 97 NMB H 17302; from locality NMB 16844: Rio Cana; upper part of Cercado Formation (late Miocene); 4. enioconch x40: & transition protoconch/teleoconch, x 80; 6. apical view, x 30. (Cancellarial(Gancellaria)irowelltiMalle SO Gy rier cscs ot sece 98,100,107,109 angosturana, Cancellaria (Hertleinia) 107,108 FBR DE, CR ICE EUTIT See ae Boer ee EO ERO a aon aotee 99 ANSP [Academy of Natural Sciences, Philadelphia, PA, U.S. A.] 87,88,92,94,102,104,107,114 Aphera Adams and Adams, 1854 ................2...... 87,91,109,111 slacolonisi(Malinys 1917) es.cesesene e- oaces se aaeasss a -2eas eaas sense acess ee nbd SOC AES OE ce EC EE EEE AEE 24,25,26 ...... 87,89-93,110,111,B lindaesPetuchipl\9 8i//mese eons. a cecte snes cos senessesaasce sce asees 109-111 DEFUANAINESOD WB O meen acces ccaaweasenec Se ocoasacewsstenonecineeeeens 111 LESSEN ALG (SOWETO S324) neeecne sane <2-nesease ene sstasceneeresareene 111 wigginsi (Emerson and Hertlein, 1964) 111 Arizelostoma Iredale, 1946 soon ulltts} VASCrONiblredale wl OS Optic: cos cco ee ea ee 113 Atlantic Ocean, (ESTEE aceset acs ean ecRO ALBEE CODE OCC EEE EECO ECO CE DEC nee CEE cece eeer eter 114 PVC SLO eaten re incvicsc asaiestacesissassavcscascedcecapercievcss (ovceetes 116 BEUTICUICPCT ICN GANCENGNIQ) ro: secs cnccenocaseoes nea eeasee ene 100,101,118 EMUIStr AL Aleee teen se ssi se sasnacsv see. vseainancroniss eases cass crcecedceuese 92,113 BEC EICNQUPCUIt 1988 rc ceec ens cetaesgens ceeeetesenteseceaecet 87,91,114,115 EINIDICMON MASP a vss cowseces dosent PA cocese 87,88,93,114,115,B EAHA URE WSNAD), Coccseceosostossauseececoscdedannaencseanieenecee 115 scalatella (Guppy, 1873) ................06608 115 SATOH (IDEAL TIRES) cococesaccesoncessoasseocse 114,115 EAISDEN Olsson eli9 G4) ere se ceec ace csesen ae snteas soesanses aces seeteaesere 115 BAW TEST (ISS) Merecescctecteee wrsoretckie ross deSse ec sstors daocee mo dnsdceseae os 92 BSAA AS Pee eee es see eee esate reece Ak eee sa Onie aes Seas os seta eee owe ene ee eae 116 Baitoa Formation 88,99,100,104,105,109,111,114,115 bajonensis, (COATT CATION Seba ascobspcceo DOO OTE OCER ECORI SEO 89,90,106 Cancellaria (Bivetiella) .. 22,26 ...... 87,88,91,93,104,105,106 FE] DOGO CONCEN ANI came nace creck Meee Tate ere ee Soe aes nte 97 BEAT DAG OS ecm ne 5 eee datos nee anes ce ost cies ta ce onc wanoeaee Senet eaten toe aane 109 WS EP AMES hp eeea ey coceecs are cee coeac hc sean ones oi nintes se teaa ie sabaae mesroes 109 Barnard (959): 2. 0... avuncccctieswasessvoecsescdsedseavesduewsereat iene tee 102 Darreti Cancel aria’ occas nccadcdessauaivscacdicescctavecessserestes eters 117 barretti, (GOT Te] 1s TPR RPE APPEEPECE PEEP ELT TEY EEE CEE TEER TOE E IED 95,96,117 Gancellarial(Gancellaria) sees lores 117 DAY CriSAGIMNCtUl a 522. soccscccesnissaoe desu suesseascvsocscccses cocuserssoeees 116 DEGLI VA BANU occ decscusa sides decsscuscasceacees scdtecsees tone otees LG Benamy, Elana Arent Th BermontshOrmatiOn i ecccccccccsscsvevessesestseesescatescstescstserccasvesy 118 BIVEH A VOUSSCAUIME lO OU ence etaceeaeeresceserantere seta eter ete er aeeeee 103 TRATIEDN OUSSCALLINE WO S/n oes ee ee eater sees ce ence saree sere en eeeneee 103 Bivetiel QaNiar ks 9 49) een ere eee ee ee eee nee 103 Bivetiella Wenz VQ43esrers-e-22e seen 91,103,105,107 IBIVELODSTORIOUSSCAUINIC ES Ot mene eeee ees eae eaeeee 106,107 IBIVELOPSISHNOUSSCAUIMEN LS OU ceeee tee necetee ieee cendancectessdcecersaeene 106 Blairivalles (US25—182i7))ere- sesso reece are ester ceceencerce eee 91,94,112,113 BMNH [British Museum (Natural History), London, England, UK] ee a eee ae ee a eae nN Seren ane ene 87,92,105,117,118 IBOEDONNBORMAtION cececcccecesecsese cence .-.c--52. seco cence ee 100-102,117,118 laevescens portoricana Maury, 1920 ...................0ee08+ 100,118 diparas Woodring, LOSI e. seksi ree nonce eee ee 104,105 LODEZANG= NESD\ pecres sac ae setenaee tee B Fyrata (Brocchis US14) ons scarce cence eee ele ee 102 macnerlisManstelds VOSMis.25.<-eeneneston oe nen eee eee 100 maldonador Olssons 1.964 acces ee cn see eaee eae see eae acess one eeare es 98 Maurya Olsson 922 reer, ce sncsacescswsceet seeaneane 89,95-97,117 metuloides! Olsson, 964 | ccscscessssiacts scx osase ooeenast eee eee ae eee 98 PIAL ATI VN SD) Sor cosa one se acres ee wae ERS RRR OTROS OE B PNILTI[ON IMS SOWETDY: 832A -o-2-nscencececeserens =< sescnseeereeseee 107 montserratensis Maury, 1925a 104 moorel Guppy, VS66) 22. 2.ssccneo ELE RESP ECECE ETE EER CCE Meee eR Deca soar 99 ROM ELOMD AM ei ereaetap ese eacrese acne techs deena seaaasesecsseas eae 90 RIO} Canal i... .235.60025 88-90,92,93,98,105,107,111,112,116,B GCaiMitO ye sececodoeecseck a ce sconeascey ees iveesssewsseenacceeteeeelee 98,112 RTOMGUAVUDIN eect terete so bees cect seca cows eaeaun aresmten eet ante 88 IRIOMGULADO! see se ees econ eee 88-90,92,93,95,97,98,105,108, 110,111,113,115,116,B RIO IMAG} .c20.<. ssevnderssnsesyans 88-90,93,95,96,105,106,110,111 Cercado de Mao, Bluff 1 of Maury (1917) ........... 88,90,91,94,95,105,106 Bluff 2 of Maury (1917) .............. 88,89,95,106,110,111 Bluff 3 of Maury (1917) ......... 88,90,95,96,106,110,111 RIO) Verde es cecssesnonsess steers scarves 88-91,98,103,105,111,116 RiosVaque del Norte: 2255, felicckc.tee- ecsscconces 88-90,98,99,105 ANS OSLUTAs GOLRE! fess asnies 114 smithii, VAX CTCL a eae cases cone ea ese 114,115 Cancellaria 114 Solander (1766) 116 SOliAG; ‘CANCCH ANIA vacseksccccee cn wee eae Jowno soe awe soon on TE Tee 100,101 Sonoma State University, Rohnert Park, CA .................:06255 88 South America, MOREM OTE sei aa oa code ew cee t conat oreo ee rete ee ae eens 113 TIGTENWESTEIM: oc ssce nc ones donee eden a svaduasaxt cosueecasseeawaneve 108,109 Souths Carolinas <. ciao s- acoocewewie os ccuavocucensuasecnscacsossetteceee costes 94 Sowerby (1832a) é Sowerby (1832b=11833)) =e tkecsatewco cee ccaecsewescezcsasctes 92,103 SOWELDY (S94) ices aera ceases oe one naan depeaneenee te neceraawenattenen ses 92 spatiosa, Gancellariar ives: Cancellaria (Pyruclia) .... Spring yvalevbormatonisee ee -esccc cece -a-soncsenssqcenseneseecenneneecee STRONGTWA RANT Ns act rensocens co oksene esse enc see he nota nae nee eee 116 Suter, Wolfgang Syeltia Jousseaume, 1887 VATICOSABLOGCCH I» -eecrecoceecicen aoe eco ec a 102 Swiss National Science Foundation ......................esceseeeeeeees 87 SPabera, Group correc ac SaesSev nee vctestasee sevice. evra ere 100 telermbas Gancellarias sci. ccnscoscrsee oot eee eee 100 LENCTA™ CANCEN AVIAN =. so csiociecedowsea teenie dncas Woe bee ta eden eORER ROCs 113 tenerum, IATA) TEDL OTE eR eRe CAE CoE OPO OER EEEEE COREE LOSER CIOS BOEECOCUOCEE HEC Trigonostoma (Ventrilia) tessellata, ADOT DP bs. eo encase nsenide love sdacniecaawtuvecusdasuaaaeseesteneeee oes 111 Cancellaria .... : PSHE AK CLE] a ene Rae are ane aoe TEE Sa 115 PPNOMPSONG) AN oc rscaceivessoa ve eseaeee reo eee oeecoeee war ssner ete ace ates Torpedo californica Ayres, 1855 .... Moula:CUSO9) i ccoen s- ves counweoawacecasdes MOUlaGLO UM) we Lovcccsaeuseteetcscesvess ois PVICONG SUTIN US Opa s cncisede te soe cea dye spensoe con aca nice seme edaeoee Trigona Perry, WSU: n. ossvec even saeensoodacssuvevs cevsdavensvensnosesecees pellucida Perry, 1811 112 Trigonostoma Blainville, 1827 ...................02+5 87,91,94,112,113 TAZ ASSEZIIID AUS 8 Die as edocs soe -voe te edoe wena sare ceer sant eaee renee 115 gurabis| (Maury, 1917): cv..22:.c0ssc0re.dsccoeve ser cevecasecsees sesense ste B insulare (Pilsbry and Johnson, 1917) ............2....0.:0000 87,114 cf. insulare (Pilsbry and Johnson, 1917) .... 114 smithfieldensis Oleksyshyn, 1960 ................cseccseecneseneceees tenerum (Philippi;.1848) |, scccsccccccsevsecss.csewssnvesvacsausterseestrs Trigonostoma (Extractrix) Korobkov, 1955 .... Trigonostoma (Ventrilia) Jousseaume, 1887 .... breve (Sowerby; 1832a)) <-s-ec2----0-cesscese=s es goniostoma (Sowerby, 1832a))) tcc: ce .s0neeeeeosccrecseoesse--¥- ee gurabis (Maury, 1917) ............... tenerum (Philippi, 1848) ...................-...+- tuberculosum (Sowerby, 1832a) iNT) Trigonostoma (Ventrilia ?) insulare (Pilsbry and Johnson, 1917) ... Be see Fars shh Poste Sead 5 ust DT ich pease Ris te OT AS AA trigonostoma, Delphinula 113 Sirinidad | Sprine Vale iecscceeacate- sa are canceseaesasseeseasee ase eneeee 100,118 TU [Tulane University, New Orleans, LA, U.S. A,] ..................- So Ley OS PPE CPT T Ere 87-92,94-103,105-107,111-113,116 PUI DATA KSTOUD co ose esscas crn sdacrinedacctouvees 95,98,100,101,117,118 BULLETIN 334 tuberculosum, Trigonostoma (Ventrilid) ...............02.00-00e-0e00 113 Buckerand Wilsoni(U932) <.ca.cccasesesccscosensececaeet ee ee eae eee 117 Mum bESHROnMAT ON esesesee cree cedseeeeeen eae ee eee eee 100,101,111 United States, offithesoutheasternicoast cess-1.-osdeeeee eeeence cre eee eee 114 SoutheaSstemn x: sccsicsCevsc ces. coecesazeasseeneon toons 108,109,113,114 urceolata, Cancellaria ... 99 Uscari Formation 111 USGS [United States Geological Survey, Washington, DC, U.S. A.] SE EEE ECOL EEE EEE PEPER eer pease eS sttccccnoeseecs 92,112 USNM [United States National Museum of Natural History, Smith- sonian Institution, Washington, DC, U.S. A.]........ 87,92,99 uva, GANCON ANIA seis cssescs seas cacsnsecdse snes ee 90,102 Cancellaria (Pyruclia ?) ......... 18:3 87,88,93,100,/01,118 VAFICOSG;, SVELLIG 2505. 5 oa Sea aioe Seo 102 VATTICOSG;, WOlUtA! < dexoeacs debe ee 102 Weénezucla: 2 Sie a ies cas acticeas scence ee 105,114,116 Golfo de Triste sae 109 Parapuana Peninsula)... < aa due dowasaeee dab'exan emastaseadsen deste vets oeee comet eee 87,92 wzahni: Cancellaridiess soscts ste, hnovsensceceese ee ee 102,103 2 OAIAV ANG) TA GINEHUI Dies teeaeeceseeeeareeeee 87,91,93,116 ZAPOLCCA; AAMELULA, vc. ..ss.ssnasasneawdacnascseavusssonswsescees sere teres Zone D of Maury (1917) Zone E of Maury (1917) Zone H of Maury (1917) Zone I of Maury (1917) ZOOS. FOFMALLON) ....620:50s0-ceconecensctesscesesscsasscassseasoaeseateee Z aunByaxay £ eunbiyp axes tp aunBijy-axay CERCADO FORMATION MAO FORMATION FORMATION GURABO edt 0) \) j Way We tdeee rat rn i t Hi H 2 ? Abe Hele Heiney ‘ ne LEE ft , i: Hi Ht ; a ni f , Hy i t) i ie = “4 uh " 1 --th- i C ian ea = —_-- — --- — C.harrisi an San a == — — — Aphera islacolonis Perplicaria canae — — — Aagatrix losquemadica Text-figure 2.—Columnar section of Rio Cana showing (discon- refer to thickness in m. CERCADO FORMATION GURABO FORMATION : tinuous) “ranges” of cancellariid species (after Saunders, Jung, and = ————— Biju-Duval, 1986, text-fig. 16). Numbers in second column from left - = re Ah 1 ets 5 c ; ra Arie 5 Spel] 5: 5 skies Sheed 55 ~ “shill HS SSS SSS = C. juncta —— C.harrisi — = — — = ——— _ =— — _ — ————_ C._ epistomifera C. miranda eS SS ES ee = = eo = = SS —— —— Aphera islacolonis Text-figure 3.—Columnar section of Rio Gurabo showing (dis- Trgonostomataursels continuous) “ranges” of cancellaniid species (after Saunders, Jung. eee i eas AY ny ee raeees Acatrin lnsqderneatea and Biu-Duvyal, 1986, text-fig. 6). Numbers in second column from sq) left refer to thickness in m Sa es pe 3s z a cE é E ie 3 —_—__—~ g 32 = oe FY 3 a3 S 3 2 e = & 3 = rs o = Sie 6 Car} 3 = 6 = = gssili ¢ 4,—Columnar section of cliffexposures on Rio Yaque ar Lopez, north of Baitoa, showing (discontinuous) ancellariid species and stratigraphic position of NMB fier Saunders, Jung, and Biju-Duval, 1986, text-fig. 25). C. rowelli C. gabbiana C. epistomifera C. lopezana Aphera islacolonis — Axelella emblema PREPARATION OF MANUSCRIPTS Bulletins of American Paleontology usually comprises two or more sep- arate monographs in two volumes each year. This series is a publication outlet for significant longer paleontological monographs for which high quality photo- graphic illustrations and the large quarto format are a requisite. Manuscripts submitted for publication in this monograph series must be typewritten, and double-spaced throughout (including direct quotations and ref- erences). All manuscripts should contain a table of contents, lists of text-figures and (or) tables, and a short, informative abstract that includes names of all new taxa. Format should follow that of recent numbers in the series. All measurements must be stated in the metric system, alone or in addition to the English system equivalent. 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Gilbert Dennison Harris (1864 - 1952) Founder of the Bulletins of American Paleontology (1895) ISBN 0-87710-4154 ate Wue reeiee ror e or i. Binekneenae 8 4e* eae ud nanaretes OSI Sh Beason Ppt r a 5 ata aye ‘7 oe Peres As Rolie 7H ar Theraeat ee Seat FOOwert ae 4, . nen aoa ol epee ee pea ec al ets ST "ATet ity . foie 4 : i epr corenrere ruse oe ere ee Soe aes