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Xo dite ea Be es SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM BULLETIN 192 THE OPERCULATE LAND MOLLUSKS OF THE FAMILY ANNULARIIDAE OF THE ISLAND OF HISPANIOLA AND THE BAHAMA ARCHIPELAGO BY PAUL BARTSCH l. ay PHO agi rit Ser ; AAS 22200000 eo UNITED STATES GOVERNMENT PRINTING OFFICE WASHINGTON : 1946 For sale by the Superintendent of Documents, U.S. Government Printing Office, Washington 25, D.C. Price 75 cents ADVERTISEMENT The scientific publications of the National Museum include two series, known, respectively, as Proceedings and Bulletin. The Proceedings series, begun in 1878, is intended primarily as a medium for the publication of original papers, based on the collections of the National Museum, that set forth newly acquired facts in biology, anthropology, and geology, with descriptions of new forms and revisions of limited groups. Copies of each paper, in pamphlet form, are dis- tributed as published to libraries and scientific organizations and to spe- cialists and others interested in the different subjects. The dates at which these separate papers are published are recorded in the table of contents of each of the volumes. The series of Bulletins, the first of which was issued in 1875, con- tains separate publications comprising monographs of large zoological groups and other general systematic treatises (occasionally in several volumes), faunal works, reports of expeditions, catalogs of type speci- mens, special collections, and other material of similar nature. The ma- jority of the volumes are octavo in size, but a quarto size has been adopted in a few instances in which large plates were regarded as in- dispensable. In the Bidletin series appear volumes under the heading Contributions from the United States National Herbarium, in octavo form, published by the National Museum since 1902, which contain papers relating to the botanical collections of the Museum. The present work forms No. 192 of the Bulletin series. ALEXANDER WETMORE, Secretary, Smithsonian Institution. II CONTENTS Page Mae uniiamGae Of EIS PAtiOlde vaicn css. 0ip\s sie ajaia sie sits opie 62 5 gia qe-0¥ a9\5.6 1 Family Annulariidae Henderson and Bartsch ..............0ssee0s 3 Subfamily Chondropominae Henderson and Bartsch .......... 5 Genus \Chondropomat IP SCI eTE'.6 ites ot aloes oo cqeimsic's ¢a'e's 5 Wetmorepoma, new subgenus .........cccscsccccssscs 6 Subgenus Chondropomorus Henderson and Bartsch.... 7 Subgenus Chondropomium Henderson and Bartsch.... 22 Subgenus Chondropomella Bartsch ..........cccceceees 36 Articulipoma, NEW SUODSEMUS 5. .524.. 84.04.20. « Haitipoma Pseudolamella present. Peristome simple. SHellovard ie UAE are os cites care ete ie een oR ee ee Kisslingia Shellvelongate-conic) jes SU esis Gece Ae ee oe Klattea Peristome double. Axial and spiral sculpture covering entire surface............ Licina Axial and spiral sculpture not covering entire surface. Outer peristome very broadly expanded.............. Sallepoma Outer peristome not very broadly expanded...... Clydonopoma Genus PARACHONDRISCA Henderson and Bartsch 1920. Parachondrisca HENDERSON and BartscH, Proc. U. S. Nat. Mus., vol. 58, pp. 65, 66. Shell elongate-conic, marked by axial riblets only, which are fused into tufts at the summit. No trace of spiral sculpture, even on the umbilical wall, is present. Operculum with rather distantly spaced, retractively curved, strongly elevated, thin lamellae, which are fused on the inner edge of the turns. They are most elevated at the inner edge of the whorls and decrease outwardly, terminating before reaching the outer margin of the chondroid basal plate. Type species: Cyclostoma umbricola Weinland = Parachondrisca um- bricola (Weinland). 13 The term pseudolamelia is here applied to the flat plate formed by the expansion and fusion of the retractively curved opercular riblets on their outer edge, which produce an appearance simulating the decidedly reflected lamella of certain annularine mollusks. The latter, however, have the lamella free and not joined to the basal chondroid plate by riblets. The pseudolamella may cover the entire ribbed surface of the opercular whorls or be restricted to the inner margin, in which case the distal portion shows the ribbing only. Pseudolamellid Rhytidopomini- nae seem restricted to Hispaniola. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 73 PARACHONDRISCA UMBRICOLA (Weinland) Shell small, elongate-conic, brown, unicolor or with rather broad spiral bands of darker brown; interior of aperture pale brown, with the brown bands when present showing conspicuously. The expanded peri- stome is flesh-colored. Nuclear whorls 1.5, forming a rather blunt apex, strongly rounded, smooth, except for the last portion of the last turn, which shows the beginning of the postnuclear sculpture. Postnuclear whorls strongly rounded, marked by slender, rather distantly spaced, slightly retractively slanting axial ribs. At irregular intervals these riblets become expanded at the summit to form a broad, almost blisterlike denticle, which is always pale in color. Suture strongly constricted, almost channeled. Periphery strongly rounded. Base short, strongly rounded, openly umbilicated, and marked by the continuation of the axial riblets, which extend into the umbilicus where some at irregular intervals become expanded into broad lamellae. These expanded ele- ments are usually also stronger on the upper portion of the last whorl which they render slightly varicid. There is no spiral sculpture present, even on the umbilicus. Last whorl solute for about one-tenth of a turn, the solute portion conspicuously angulated behind the posterior angle of the aperture. Aperture subcircular; peristome double, the outer broadly expanded at the posterior angle where it forms a conspicuous auricle, while on the inner lip it is about half as wide as at the posterior angle; on the middle of the parietal wall and the outer lip it does not extend beyond the inner peristome, which is slightly exserted and re- flected. Operculum typically parachondriscoid. KEY TO THE HISPANIOLAN SUBSPECIES OF PARACHONDRISCA UMBRICOLA Axial ribs ane and closely spaced -26 036 or aes asans mnie the 4s cayemitensis Accial ribs not fine Or closely SPACE. < o <.<.c nye .cysce e's oueie ea aelie'sin omyes chow umbricola PARACHONDRISCA UMBRICOLA CAYEMITENSIS, new subspecies PLATE 13, FIGURE 4 This subspecies comes from Grande Cayemite, a large island lying off the north coast of the south peninsula, Haiti. Its larger size and finer and more closely spaced ribs, of which there are 144 on the last whorl in the type, as well as the less strong tufting at the summit, will easily distinguish it from typical P. «. umbricola. The type (U.S.N.M. No. 503997), collected by Eyerdam, has 8.4 whorls and measures: Length, 13.5 mm.; greater diameter, 6.1 mm. ; lesser diameter, 5.0 mm. U.S.N.M. No. 380314 contains 208 topotypes from the same source. PARACHONDRISCA UMBRICOLA UMBRICOLA (Weinland) Pate 13, FIGURE 7 1862. Cyclostoma umbricola WEINLAND, Malakozool. Blatter, vol. 9, p. 88. 1865. Tudora umbricola Preirrer, Monographia pneumonopomorum viventium, Suppl. 2, p. 137. 1920. Parachondria (Parachondrisca) umbricola HENDERSON and Bartscu, Proc. U.S. Nat. Mus., vol. 58, p. 66. 74 BULLETIN 192, UNITED STATES NATIONAL MUSEUM The typical subspecies ranges from Pestel, on the north coast of the southern peninsula, west to Anse du Clerc. Its much smaller size, less developed tufting at the summit, and much stronger and more distantly spaced axial ribs will readily distinguish it from P. u. cayemitensis. The specimen described and figured is one of 34 (U.S.N.M. No. 355166) collected by John B. Henderson and the author at the type locality, Jérémie, Haiti. It has 8.4 whorls and measures: Length, 10.8 mm.; greater diameter, 4.2 mm.; lesser diameter, 3.3 mm. There are 70 axial riblets on the last whorl. A hundred specimens yield the following comparative measurements: Length Greater diameter Lesser diameter Mm. Mm. Mm. GIreAtesticsca cad cele 12.9 Sf 4.9 east ails sd. SFA 2, 7.0 3.4 2.6 AAVeTapel cass cae Coe 9.7 4.4 35 U.S.N.M. No. 425689 contains 3 cotypes received from Weinland. U.S.N.M. No. 380323 contains 1 specimen collected by Eyerdam at Pestel. U.S.N.M. No. 355179 contains 4 specimens collected by Henderson and Bartsch at Corail. U.S.N.M. No. 355176 contains 4 specimens collected by Henderson and Bartsch on top of the range at Roseau. U.S.N.M. No. 355175 contains 29 specimens collected by Henderson and Bartsch at Roseau. U.S.N.M. No. 355174 contains 24 specimens collected by Henderson and Bartsch at Abricots, west of Jérémie. U.S.N.M. No. 355177 contains 14 specimens collected by Henderson and Bartsch at Jérémie. U.S.N.M. No. 162956 contains 32 specimens collected by Henderson and Simpson at Jérémie. U.S.N.M. No. 355168 contains 1 specimen collected by Henderson and Bartsch on the second ridge west of Jérémie. U.S.N.M. No. 355169 contains 5 specimens collected by Henderson and Bartsch on the shore cliffs at Jérémie. U.S.N.M. No. 355171 contains 27 specimens collected by Henderson and Bartsch at Debarras. U.S.N.M. No. 355172 contains 4 specimens collected by Henderson and Bartsch 5 miles west of Jérémie. U.S.N.M. No. 355173 contains 10 specimens collected by Henderson and Bartsch at Anse a Cochon. U.S.N.M. No. 403057 contains 17 specimens collected by Orcutt near the sea west of Jérémie. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 715 U.S.N.M. No. 403074 contains 12 specimens collected by Orcutt south of the road west of Jérémie. U.S.N.M. No. 402911 contains 155 specimens collected by Orcutt east of Trou Bonbon. U.S.N.M. No. 402653 contains 66 specimens collected by Orcutt be- tween Anse du Clerc and Trou Bonbon. U.S.N.M. No. 355178 contains 45 specimens collected by Dr. W. L. Abbott at Moron. CROSSEPOMA, new genus Shell elongate-conic, marked by axial ribs that fuse at the summit to form tufts. Spiral sculpture present in the form of threads or indicated by feeble nodules on the axial ribs, always stronger on the umbilical wall than on the spire. Umbilicus open. Aperture oval; peristome simple. Operculum with a heavy calcareous deposit, which is marked by retrac- tively curved axial riblets, leaving, however, a space between the turns showing the basal chondroid plate. Crossepoma is easily distinguished from Parachondrella by the strong tufting of the axial ribs at the summit. In Parachondrella they remain distinct. Type species: Cyclostoma emilianum Weinland = Crossepoma emili- anum (Weinland). KEY TO THE SPECIES OF CROSSHPOMA Asal tins: GIStanitly SPACEMT et het eee ee ck ae eee oe eee barnes hendersoni Axial ribs closely spaced. Axial ribs broad. Shelljarce, height more than. 15 mm ............066+ 93° emilianum Shel small) height less) than. LZ aii. ci e.. 22 torent «loc aie ere australe Axial Eri S* MOPIDTOAC see aia owe cee Mo eee let oleae ete o bate wore jacmelense CROSSEPOMA HENDERSONI, new species PLaTE 13, Ficure 9 Shell elongate-conic, pale buff, with interrupted spiral bands of brown, of which six are present between summit and periphery. A very broad seventh spiral band is immediately below the periphery and three smaller ones are anterior to this on the base. The columella also is pale brown. All these brown markings show within the aperture. There is also a small brown spot at the beginning of the nuclear turns. Nuclear whorls 1.5, inflated, well rounded, microscopically granulose. The ter- minal portion of the nuclear whorls gradually passes into the post- nuclear sculpture. The postnuclear whorls are moderately well rounded. The early ones less so than the later. They are crossed by strong sub- lamellar axial ribs, several of which may become fused into hollow cusps at the summit, or the individual ribs may become thus expanded. Of these axial ribs, 84 are present on the last turn. The spiral sculpture is indicated by feeble nodulations, which are stronger on the early turns. The intercostal spaces do not show spiral threads. Suture moderately 76 BULLETIN 192, UNITED STATES NATIONAL MUSEUM contricted. Periphery inflated, well rounded. Base moderately long, nar- rowly openly umbilicated, and marked by the continuation of the axial ribs and on the outer half of the umbilical wall by four spiral cords, which grow consecutively stronger from the outermost inward. Aper- ture oval, slightly angulated at the posterior angle; peristome simple. Operculum typically crossepomid. The type (U.S.N.M. No. 380353) was collected near Anse 4 Veau, Haiti, by Eyerdam. It has 5 whorls remaining and measures: Height, 16.1 mm. ; greater diameter, 7.3 mm. ; lesser diameter, 6.5 mm. Additional specimens in our collection show that the species ranges from Anse a Veau along the coast westward to Trou Rousselin at the western end of the peninsula. The species is easily distinguished from its neighbor, C. emilianum, by its much more distantly spaced axial ribs. U.S.N.M. No. 503998 contains 1 specimen collected by Henderson and Bartsch at Roseau. U.S.N.M. No. 383227 contains 4 specimens collected by Orcutt at Jérémie. U.S.N.M. No. 401730 contains 139 specimens collected by Orcutt between Anse du Clerc and Trou Bonbon. U.S.N.M. No. 404079 contains 2 specimens collected by Orcutt on a limestone cliff near the shore west of Trou Rousselin. CROSSEPOMA EMILIANUM (Weinland) Shell elongate-conic, varying in ground color from pale buff to brown, marked by interrupted, or sometimes solid, spiral bands of brown; the interior of the aperture may vary from pale buff to pale brown with the peristome usually soiled white. The nuclear whorls form a blunt apex that consists of 2.5 turns, which are strongly rounded and smooth except for the last portion of the last turn, which shows the beginning of the postnuclear sculpture. Postnuclear whorls well rounded, marked by closely spaced, retractively slanting axial riblets, which are usually a little wider than the spaces that separate them. The axial riblets are either expanded or several of them are gathered into hollow tufts at the summit. The spiral sculpture consists of obsolete ill-defined nodules, which render the axial riblets slightly wavy. Suture well constricted, rendered wavy by the tufts at the summit of the turns. Periphery strongly rounded. Base short, openly narrowly umbilicated, well rounded, and marked by the continuation of the axial riblets within the umbilicus and by moderately strong spiral threads. Aperture oval; peristome simple, slightly reflected. Operculum with the nucleus halfway between submarginal and subcentral, marked by slender, retractively slanting, ill- defined lamellae, which are fused on their inner and outer edges, which are slightly elevated, and which terminate about two-fifths from the outer margin of the chondroid basal plate of the turns, leaving a de- cided unribbed space between the turns. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 77 This species appears to be distributed over the coastal region of south- western Haiti. I am recognizing three subspecies, which the following key and descriptions will help to differentiate : KEY TO THE SUBSPECIES OF CROSSEPOMA EMILIANUM Shell: Stowe tH 6/0 ite eat. A Sa das Aa A A el gibbosum Shell not stout. Wutts-at,theisummiuty very Strongtiens | dais dened) cduowes emilianum Tatts atthe summit mot, very strong oe). 2. Se. fos. ans th ate oes insulanum CROSSEPOMA EMILIANUM GIBBOSUM, new subspecies Pate 13, Figure 8 This subspecies appears confined to the coast of the south side of the south peninsula where it extends from Damassins to Port Salut. The type (U.S.N.M. No. 536863) was collected by C. R. Orcutt on a hill north of Coteaux east of the river. This subspecies is easily differ- entiated from the other two by its much more gibbose form and by having the axial ribs finer and less expanded at their outer edge. In the type, which has 4.8 whorls remaining, there are 167 ribs upon the last turn and 3 obsolete spiral cords on the outer edge of the umbilical wall. The type measures: Height, 17.0 mm.; greater diameter, 10.0 mm. ; lesser diameter, 8.0 mm. U.S.N.M. No. 404032 contains 83 topotypes from the same source. U.S.N.M. No. 404900 contains 7 specimens collected by Orcutt at Coteaux U.S.N.M. No. 404826 contains 21 specimens collected by Orcutt west of Carpentier. U.S.N.M. No. 404096 contains 3 specimens collected by Orcutt on the first hill east of Damassins. U.S.N.M. No. 404127 contains 7 specimens collected by Orcutt on a hillside south of the road west of Coteaux. U.S.N.M. No. 403952 contains 6 specimens collected by Orcutt east of Riviere de l’Anse a Drick. U.S.N.M. No. 403984 contains 14 specimens collected by Orcutt at Point Sable west of Port Salut. U.S.N.M. No. 404761 contains 2 specimens collected by Orcutt on a hill north of Port a Piment. U.S.N.M. No. 402331 contains 91 specimens collected by Orcutt on the road between L’Acul and Port Salut. U.S.N.M. No. 403740 contains 28 specimens collected by Orcutt at Port Salut. U.S.N.M. No. 402121 contains 26 specimens collected by Orcutt on the north side of the Riviére du Port Salut near the mouth. U.S.N.M. No. 402159 contains I specimen collected at Acuzoo Village south of the Riviere de l’Acul. U.S.N.M. No. 403962 contains 4 specimens collected at Anse a Drick. 78 BULLETIN 192, UNITED STATES NATIONAL MUSEUM U.S.N.M. No. 404784 contains I specimen collected by Orcutt I mile west of Point Sable. CROSSEPOMA EMILIANUM EMILIANUM (Weinland) PLATE 13, Ficure 3 1862. Cyclostoma emilianum WEINLAND, Malak. Blatter, vol. 9, p. 87. 1862. Chondropoma emilianum PFEIFFER, Malak. Blatter, vol. 9, p. 94. 1920. Parachondria (Parachondria) emilianum HENDERSON and BartscuH, Proc. U.S. Nat. Mus., vol. 58, p. 66. This subspecies was collected by Henderson and me at various sta- tions along the coast between Jérémie and Anse du Clerc on the north- west coast of the south peninsula. Its slender form will readily distin- guish it from Crossepoma emilianum gibbosum, and its much more ex- panded ribs and stronger tufting at the summit and less gibbose whorls differentiate it from C. e. insulanum. The specimen figured (U.S.N.M. No. 124402) is one of two cotypes received from Weinland, that is, a paratype. It has 5 whorls remaining and 8g ribs on the last whorl and 4 moderately strong spiral threads on the outer portion of the umbilical wall and measures: Height, I5.2 mm.; greater diameter, 7.4 mm.; lesser diameter, 6.6 mm. One hundred additional specimens of this subspecies, also collected from Jérémie or the adjacent territory, yield the following average measurements : Length Greater diameter Lesser diameter Mm. Greatest : 6.9 WWOASE! <0 ciererevomte sie : 49 Average : 5.8 U.S.N.M. No. 425691 contains another specimen received by the Senckenbergische Museum from Weinland. U.S.N.M. No. 355348 contains 4 specimens collected by Henderson and Bartsch at Jérémie. U.S.N.M. No. 355350 contains 117 specimens collected by Henderson and Bartsch on the sea cliffs west of Jérémie. U.S.N.M. No. 355352 contains 1 specimen collected 5 miles west of Jérémie by Henderson and Bartsch. U.S.N.M. No. 355353 contains 143 specimens collected by Henderson and Bartsch at Anse a Cochon, U.S.N.M. No. 403056 contains 84 specimens collected by Orcutt near the sea west of Jérémie. U.S.N.M. No. 402924 contains 4 specimens collected by Orcutt at Trou Bonbon. U.S.N.M. No. 425374 contains 1 specimen collected by Orcutt be- tween Anse du Clerc and Trou Bonbon. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 79 CROSSEPOMA EMILIANUM INSULANUM, new subspecies PLATE 13, Ficure 10 This subspecies comes from Grande Cayemite Island off the north coast of the south peninsula. It most nearly resembles the typical race but is readily distinguished from that by its more inflated whorls and less strongly developed ribs, which are also less wide. The type (U.S.N.M. No. 425508) has 5.3 whorls remaining and bears 121 axial ribs on the last turn and 4 weak cords at the outer edge of the umbilicus. It measures: Height, 16.9 mm.; greater diameter, 8.0 mm.; lesser diameter, 7.2 mm. CROSSEPOMA AUSTRALE, new species Shell rather small, buff, unicolor, or with faint indications of inter- rupted spiral bands of brown. In the typical race there is also a strongly interrupted spiral band of brown on the base a little below the periph- ery. Nuclear whorls 1.5, well rounded, microscopically granulose. Post- nuclear whorls also well rounded and marked by slightly retractively curved axial ribs, which are gathered into tufts at the summit. On the early whorls spiral threads are indicated by rather strong nodules on the axial ribs; these are less apparent, although more numerous, on the later turns. Suture well constricted. Periphery inflated, strongly rounded. Base rather openly umbilicated and marked by the continuation of the axial ribs and several strong spiral cords on the outer edge of the umbilicus. Aperture broadly oval, slightly angulated at the posterior angle; peristome simple. Operculum typically crossepomid. The species is distributed over portions of the western part of the south peninsula of Haiti and the fle 4 Vache. It breaks up into two subspecies which the following key and descrip- tions will help to differentiate : KEY TO THE SUBSPECIES OF CROSSEPOMA AUSTRALE Umbilical wall with 3 strong and an outer feeble spiral thread..... vachense Umbilical wall with 2 strong and an outer feeble spiral thread...... australe CROSSEPOMA AUSTRALE VACHENSE, new subspecies PLATE 13, Ficure 6 This subspecies comes from the fle 4 Vache. It has finer ribs and feeble tufting thereof, as well as the possession of three strong spiral threads and a feeble cord on the outer portion of the umbilical wall, which readily distinguish it from the typical race. The type, U.S.N.M. No. 442849, has 103 axial ribs on the last whorl and measures: Height, 10.3 mm.; greater diameter, 6.0 mm.; lesser diameter, 4.7 mm. 80 BULLETIN 192, UNITED STATES NATIONAL MUSEUM CROSSEPOMA AUSTRALE AUSTRALE, new subspecies PLATE 13, FiGuRE 5 This subspecies ranges along the south coast of the south peninsula from Torbeck to Aquin, Haiti. It differs from C. a. vachense in having the axial ribs much stronger and the tufts at the summit much more strongly developed. There are also here only one feeble and two strong spiral threads on the outer portion of the umbilical wall. The type (U.S.N.M. No. 504027) was collected by C. R. Orcutt be- tween Vieux Bourg and Baie des Flamands. It has 4.3 whorls remain- ing, the last of which bears 110 axial ribs, and measures: Height, II.0 mm.; greater diameter, 6.2 mm.; lesser diameter, 5.0 mm. U.S.N.M. No. 402457 contains 5 topotypes from the same source. U.S.N.M. No. 403650 contains 1 specimen collected by Orcutt at Torbeck. U.S.N.M. No. 402796 contains 9 specimens collected by Orcutt at St. Louis. U.S.N.M. No. 402827 contains 9 specimens collected by Orcutt east of Aquin. U.S.N.M. No. 402679 contains 6 specimens collected by Orcutt east of Morne Rouge. U.S.N.M. No. 402696 contains 19 specimens collected by Orcutt at Morne Rouge. U.S.N.M. No. 402736 contains 2 specimens collected by Orcutt south of Morne Rouge. U.S.N.M. No. 403463 contains 1 specimen collected by Orcutt on the mountains east of Morne Rouge. CROSSEPOMA JACMELENSE, new species Shell varying from cylindroconic to broadly elongate-conic, marked by retractively curved axial riblets, which are gathered into prominent tufts at the summit. The spiral sculpture is indicated by feeble nodules on the axial ribs, which are stronger on the early turns than on the rest. Suture well constricted. Periphery well rounded. Base openly umbilicated and marked by the continuation of the axial ribs and on the umbilical wall by spiral threads. The last whorl is solute to a varying degree. Aperture oval, slightly angulated at the posterior angle; peristome simple and slightly expanded and reflected. Operculum with strong, rather distantly spaced, retractively curved, lamellar axial ribs, which are fused on the inner and outer terminations. These ribs do not extend to the outer limit of the turns of the basal chondroid plate but leave a space between them. The species appears restricted to the south coast of the south peninsula of Haiti. I am recognizing two subspecies, which the following key and de- scriptions will help to differentiate: HISPANIOLAN AND BAHAMAN ANNULARIIDAE 81 KEY TO THE SUBSPECIES OF CROSSEPOMA JACMELENSE DMCC ME ULOEGHIG™ cits. We yee as senses se axe elec bee ume peel 6 as jacmelense Shelti broad lylelosienatesconte cas OS a S280. LETRA de el cayesense CROSSEPOMA JACMELENSE JACMELENSE, new subspecies Piate 14, Ficure 14 This subspecies comes from Jacmel, Haiti, on the south coast of the island across the south peninsula from Port-au-Prince. The type (U.S.N.M. No. 355354) has lost the early whorls; the 4.4 remaining measure: Height, 14.8 mm.; greater diameter, 7.0 mm.; lesser diameter, 6.0 mm. The last whorl bears 117 axial riblets and has four spiral threads on the umbilical wall. This subspecies is easily distinguished from Crossepoma jacmelense cayesense by its cylindroconic shape, by having the axial ribs more dis- tantly spaced, and by having the last whorl much more solute and the peristome less expanded. U.S.N.M. No. 162928 contains 39 topotypes, 20 of which yield the following average measurements : Greater diameter Lesser diameter Greatest east Ooh On eae CROSSEPOMA JACMELENSE CAYESENSE, new subspecies Pate 14, Ficure 15 This subspecies ranges from Port a Piment, Haiti, eastward along the south coast to Aquin. We also have a number of dead specimens from fle 4 Vache, which I am unable to differentiate from the main- island form. The type (U.S.N.M. No. 504028), collected by C.R. Orcutt from near Les Cayes, is a complete specimen having 7.7 whorls and 192 axial ribs on the last turn and 4 spiral threads in the umbilicus. It measures: Height, 19.0 mm.; greater diameter, 9.3 mm.; lesser diameter, 7.3 mm. It differs markedly from Crossepoma jacmelense jacmelense in being much broader and in having a much larger number of ribs. U.S.N.M. No. 403344 contains 1,331 topotypes from the same source. U.S.N.M. No. 401756 contains 102 specimens collected by Orcutt on the road from Les Cayes to Laborde. U.S.N.M. No. 404130 contains 23 specimens collected by Orcutt on a hill near and west of Metesignix, east of Port 4 Piment. U.S.N.M. 404724 contains 8 specimens collected by Orcutt in the ravine west of Metesignix. 82 BULLETIN 192, UNITED STATES NATIONAL MUSEUM U.S.N.M. No. 403138 contains 183 specimens collected by Orcutt at Aquin. U.S.N.M. No. 402479 contains 70 specimens collected by Orcutt be- tween Vieux Bourg and Baie des Flamands. U.S.N.M. No. 404011 contains 9 specimens collected by Orcutt on the road between Torbeck and L’Acul. U.S.N.M. No. 401932 contains 7 specimens collected by Orcutt on the road between Les Cayes and Torbeck. U.S.N.M. No. 401783 contains 27 specimens collected by Orcutt at Les Cayes. HISPANIPOMA, new genus Shell ovate, marked by axial riblets and spiral threads whose junction forms weak nodules. Some of the axial ribs fuse to form tufts at the summit; others merely become thickened into cusps. Base openly um- bilicated, the umbilical wall always marked by spiral cords a little stronger than those of the spire. Aperture oval; peristome simple. Oper- culum heavily calcified with strong, retractively curved ribs, which do not extend to the outer edge of the whorls of the basal chondroid plate but leave a small channel between the turns. Type species: Cyclostoma quaternata Lamarck = Hispanipoma quater- natum (Lamarck). HISPANIPOMA QUATERNATUM (Lamarck) Shell elongate-ovate, varying decidedly in size in the different sub- species. The color ranges from unicolor flesh-color to pale rose; the in- terior of the aperture varies with the color of the exterior. The nuclear whorls form a blunt apex consisting of 1.5 turns, which are strongly rounded and microscopically granulose, with the last portion of the last turn showing indications of the postnuclear sculpture. The postnuclear whorls are well rounded, narrowly shouldered at the summit, marked by numerous, rather closely spaced, rounded, somewhat retractively slanting axial riblets, which vary in strength and spacing, heavier and more distantly spaced ones being usually succeeded by narrower and more closely spaced series; the spaces separating them are rarely as wide as the riblets. These riblets terminate either individually in little expanded cusps, or several of them become fused to form a conspicuous hollow denticle. The spiral sculpture consists of numerous slender, weakly developed threads, which are best seen in the intercostal spaces and which render their junction with the axial ribs weakly nodulose. Suture moderately well constricted. Periphery inflated and well rounded. Base moderately long, inflated, strongly rounded, narrowly openly um- bilicated, and marked by the continuation of the axial and spiral threads. The ribs and spiral threads are also present on the umbilical wall. The last whorl is solute for a fraction of a turn. Aperture oval; peristome HISPANIOLAN AND BAHAMAN ANNULARIIDAE 83 simple, slightly auriculated at the posterior angle, and a little more broadly expanded on the inner and basal lip than on the rest. Oper- culum as described in the generic definition. This species appears confined to Haiti. Representatives of it are known from the north coast of the island, the Cul-de-Sac region, and the island of Gonave. I am recognizing five subspecies, which the following key and descrip- tions will help to differentiate : KEY TO THE SUBSPECIES OF HISPANIPOMA QUATERNATUM Tufts at summit of whorls very closely approximated.......... quaternatum Tufts at summit of whorls not very closely approximated. Denticles at summit very strong. Suture stronely ehanneled {ac 0S sok. Seen eel. Be dentilobatum Suture!notistrongly channeled; . sed stews . pce. mee monticolum Denticles at summit not very strong. Sie lipeloicate-Omate occ cece se ctor sidlo o'er ais Gens = 5 anor cabaretense EMO VALOK. cee ee cron als oeonie rere ccreie ea ENS nals charmettense HISPANIPOMA QUATERNATUM QUATERNATUM (Lamarck) PLatTe 14, Ficure 13 1822. Cyclostoma quaternata LAMaRCK, Histoire naturelle des animaux sans vertébres, vol. 6, pt. 2, p. 147. 1842. Cyclostoma quaternata DELEssERT, Recueil de coquilles . . ., pl. 29, fig. 3. 1847. Cyclostoma quaternata Sowersy, Thesaurus conchyliorum, p. 149, pl. 28, figs. 178, 179. 1848. Cyclostoma quaternata PreirrER, Martini-Chemnitz Conchylien Cabinet, vol. 1, sect. 19, p. 81, pl. 10, figs. 12, 13. 1850. Cistula quaternata Gray, Nomenclature of molluscous animals and _ shells in the collection of the British Museum, p. 59. 1851. Tudora quaternata Preirrer, Zeitschr. Malak., vol. 8, p. 167. 1863. Chondropoma quaternata Reeve, Conchologia iconica, pl. 8, fig. 60. 1920. Parachondria (Parachondria) quaternata HENDERSON and BartscH, Proc. U. S. Nat. Mus., vol. 58, p. 66. This subspecies ranges from the Peterborough Mountains north of Gonaives, south through St. Marc through the Cul-de-Sac region to Fond Parisien and Fonds Verettes. It is the largest subspecies, usually with an orange tinge. Here, too, the tufts at the summit are more inclined to be appressed to the preced- ing turn than in the other races. The specimen figured (U.S.N.M. No. 355316) is one of 70 that Henderson and I collected at Trou Caiman near Thomazeau. It has a little over 4 whorls remaining and measures: Height, 19.7 mm. ; greater diameter, 11.0 mm.; lesser diameter, 9.8 mm. It has 150 axial ribs on the last whorl and 16 spiral threads between the summit and the periph- ery of the same whorl. The axial and spiral sculpture on the base of the last whorl in this subspecies is usually feebly developed, sometimes ob- 8&4 BULLETIN 192, UNITED STATES NATIONAL MUSEUM solete, and there are five threads on the umbilical wall. The strength of the spiral sculpture on the base, however, is variable in individuals even when gathered from the same place. Measurements of 100 specimens yield the following information: Greater diameter Lesser diameter Mm. Greatest us... sinc tc ciae east, #2 ete oe ee The large size and brighter coloration will readily distinguish this subspecies from the rest. U.S.N.M. No. 379974 contains 22 specimens collected by Eyerdam at Morne a Cabrits. U.S.N.M. No. 355309 contains 4 specimens collected by Herman Rolle at Port-au-Prince. U.S.N.M. No. 425386 contains 48 specimens collected by Orcutt at Morne a Cabrits. U.S.N.M. No. 355314 contains 2 specimens collected by Henderson and Bartsch in the Cul-de-Sac north of Port-au-Prince. U.S.N.M. No. 380436 contains 3 specimens collected by Eyerdam in the plain, Cul-de-Sac. U.S.N.M. No. 162951 contains 24 specimens collected by Henderson and Simpson at Thomazeau. U.S.N.M. No. 403099 contains 492 specimens collected at Fort Di- manche near Port-au-Prince. U.S.N.M. No. 355307 contains 231 specimens collected by Hender- son and Bartsch at Port-au-Prince. U.S.N.M. No. 403242 contains 11 specimens collected by Orcutt on the road to Damien. U.S.N.M. No. 403249 contains 13 specimens collected by Orcutt near Pont Bedette. U.S.N.M. No. 402436 contains 17 specimens collected by Orcutt near the Hasco Bridge, Cul-de-Sac River. U.S.N.M. No. 403394 contains 24 specimens collected by Orcutt to the east of the Haitian American Sugar Co. near Port-au-Prince. U.S.N.M. No. 355319 contains 3 specimens collected by Henderson on the plain east of Thomazeau. U.S.N.M. No. 401908 contains 3 specimens collected by Orcutt east of A. Triche’s house, Rue 4, Port-au-Prince. U.S.N.M. No. 380466 contains 15 specimens collected by Eyerdam 1 mile northeast of Port-au-Prince. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 85 U.S.N.M. No. 362391 contains 5 specimens collected by A. E. Vinson in the Cul-de-Sac Plain. U.S.N.M. No. 355313 contains 5 specimens collected by J. J. Brown at Port-au-Prince. U.S.N.M. No 162952 contains 66 specimens collected by Henderson and Simpson at Port-au-Prince. U.S.N.M. No. 355305 contains 71 specimens collected by Henderson at Port-au-Prince. U.S.N.M. No. 355321 contains 4 specimens collected by Henderson on the shore of Lake Assua. U.S.N.M. No. 162954 contains 3 specimens collected by Henderson and Simpson at Thomazeau. U.S.N.M. No. 162955 contains Io specimens collected by Henderson and Simpson at Port-au-Prince. U.S.N.M. No. 355320 contains 2 specimens collected by Henderson and Bartsch at Glore. U.S.N.M. No. 355315 contains 26 specimens collected by Henderson at Port-au-Prince. U.S.N.M. No. 355318 contains 2 specimens collected by Henderson at Morne a Cabrits. U.S.N.M. No. 355325 contains 1 specimen collected by W. C. Wood- ring 2 miles west of Gautier. U.S.N.M. No. 393241 contains 16 specimens collected by Bartsch 18 miles north of Port-au-Prince. U.S.N.M. No. 393776 contains 7 specimens collected by Bartsch 10 miles north of Gonaives. U.S.N.M. No. 393788 contains 6 specimens collected by Bartsch 15 miles north of Gonaives. U.S.N.M. No. 393803 contains 7 specimens collected by Bartsch on the Peterborough Mountain north of Ennery. U.S.N.M. No. 355323 contains 90 specimens collected by Henderson at Fond Parisien. U.S.N.M. No. 503999 contains 110 specimens collected by Henderson at Fonds Verettes. HISPANIPOMA QUATERNATUM DENTILOBATUM (Weinland) PLATE 14, Ficure 1 1880. Cyclostomus dentilobatus WEINLAND, Jahrb, deutschen malak. Ges., vol. 7, p. 344, in part. 1920. Parachondria (Parachondria) gonavicola HENDERSON and BartscH, Proc. U. S. Nat. Mus., vol. 58, p. 80. This race we have from various stations on the Island of Gonave, Haiti. Henderson and I failed to recognize its identity with the shell 36 BULLETIN 192, UNITED STATES NATIONAL MUSEUM we described as gonavicola. It is a stout race, smaller than typical quaternatum. The specimen figured (U.S.N.M. No. 504000) has 180 axial ribs on the last turn. It has 6 spiral threads on the first and 10 on the second whorl, while on the last they are too obsolete to be counted. There are 12 fairly strongly developed cords on the umbilical wall. The specimen figured is the type of gonavicola. It has a little over 3 whorls and meas- ures: Height, 14.9 mm.; greater diameter, 9.5 mm.; lesser diameter, 7.9 mm. Forty-nine additional specimens yield the following average measure- ments: Greater diameter Lesser diameter Length Mm. Greatest: cance ee ace eat mm meherore er oe AverTag@ecns 2 ce onecs U.S.N.M. No. 314944 contains 1 specimen collected by Dr. W. L. Abbott on Gonave Island. U.S.N.M. No. 355329 contains 32 specimens collected by Dr. W. L. Abbott on Gonave Island. U.S.N.M. No. 355330 contains 17 specimens collected by Dr. W. L. Abbott near Anse a Galets. U.S.N.M. No. 381015 contains I specimen collected by Eyerdam 1 mile northeast of Point-a-Raquette. U.S.N.M. No. 425368 contains I specimen collected by Eyerdam on Gonave Island. HISPANIPOMA QUATERNATUM MONTICOLUM, new subspecies PLATE 14, Ficure 3 This subspecies Henderson and I collected at Pétionville, and C. R. Orcutt gathered specimens at “Mount Traverse,” which is somewhere between Fonds Verettes and Saltrou, probably on the eastern side of the mountain. The type (U.S.N.M. No. 355338) comes from Pétionville. It has a little over 4 whorls remaining and measures: Height, 14.5 mm.; greater diameter, 7.5 mm.; lesser diameter, 7.2 mm. It has 137 axial ribs on the last turn and 11 spiral threads between the summit and the periph- ery on the last turn, and 12 on the base and 15 on the umbilical wall. The latter are less strong than those on the base. This race is readily distinguished from the other subspecies by its much stronger spiral sculpture. HISPANIOLAN AND BAHAMAN ANNULARIIDAE 87 Ten topotypes (U.S.N.M. No. 355339), collected at the same time, yield the following additional measurements: Length Greater diameter Lesser diameter U.S.N.M. No. 402075 contains 18 specimens collected by Orcutt at Mount Traverse between Fonds Verettes and Saltrou. HISPANIPOMA QUATERNATUM CABARETENSE, new subspecies PLATE 14, FiGurE 2 This subspecies was collected by Mr. and Mrs. E. C. Leonard on the summit of Morne Rouge, Cabaret, Moustique Bay, on the northwest coast of the island. It is a pale race of elongate-ovate outline with rather strong sculpture and very conspicuous tufts at the summit. In the type (U.S.N.M. No. 504018), which has 4 whorls remaining, there are 103 axial riblets on the last whorl, 11 spiral threads on this turn between the summit and the periphery, and 9 on the base and 9 on the umbilical wall. It measures: Height, 13.8 mm.; greater diameter, 8.0 mm.; lesser diameter, 6.7 mm. This race is most nearly related to its neighbor, Hispanipoma quater- natum charmettense, from which it can at once be distinguished by its much more elongate form. HISPANIPOMA QUATERNATUM CHARMETTENSE, new subspecies PLATE 14, Ficure 10 This subspecies was collected by Henderson and Simpson at Aux Charmettes, near Cap-Haitien. It is easily distinguished from the others by its short-ovate outline. The type (U.S.N.M. No. 355331) has 132 axial riblets on the last turn, II spiral threads between the summit and the periphery of the same whorl, 10 spiral threads on the base, and 3 heavy ones on the outside of the umbilical wall and a number of obsolete threads farther in. ORCUTTIPOMA, new genus Small rhytidopomid shells varying in outline from elongate-conic to ovate. The whorls are marked by slender sublamellar axial ribs, which may become expanded into auricles at the summit. These ribs may be straight, wavy, or even scalloped. The spaces between the axial ribs are marked by numerous very fine, closely spaced spiral lines. A series of scallops arranged in spiral order may be present or absent on the outer portion of the umbilicus. Aperture varying from ovate to almost cir- 88 BULLETIN 192, UNITED STATES NATIONAL MUSEUM cular; peristome double, the inner slightly exserted; the outer expanded. Operculum with subcentral nucleus, bearing many slender, retractively curved riblets, extending from the inner edge outward and growing weaker outwardly, vanishing before reaching the outer edge of the chondroid basal plate. Type species: Orcuttipoma orcutti, new species. KEY TO THE SPECIES OF ORCUTTIPOMA Outer peristome narrowly expanded... .<.ace. acne boon biceeae een chee orcutti Outer jpétistome» broadly expanded. 42 . ADsaIn/e) S266 .Ohe. 04.