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SMITHSONIAN
INSTITUTION

MUSEUM
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UNITED STATES NATIONAL MUSEUM BULLETIN 277

A Revision of the

Scenopinidae (Diptera)
of the World

L. P. KELSEY

University of Delaware

SMITHSONIAN INSTITUTION PRESS

CITY OF WASHINGTON
1969

Publications of the United States National Museum

The scientific publications of the United States National Museum
include two series, Proceedings of the United States National Museum
and United States National Museum Bulletin.

In these series are published original articles and monographs dealing
with the collections and work of the Museum and setting forth newly
acquired facts in the fields of anthropology, biology, geology, history, and
technology. Copies of each publication are distributed to libraries and
scientific organizations and to specialists and others interested in the
various subjects.

The Proceedings, begun in 1878, are intended for the publication, in
separate form, of shorter papers. These are gathered in volumes, octavo
in size, with the publication date of each paper recorded in the table of
contents of the volume.

In the Bulletin series, the first of which was issued in 1875, appear
longer, separate publications consisting of monographs (occasionally in
several parts) and volumes in which are collected works on related
subjects. Bulletins are either octavo or quarto in size, depending on the
needs of the presentation. Since 1902, papers relating to the botanical
collections of the Museum have been published in the Bulletin series
under the heading Contributions from the United States National Her-
barium.

This work forms number 277 of the Bulletin series.

FranK A. TAYLOR
Director, United States National Museum

U.S. GOVERNMENT PRINTING OFFICE
WASHINGTON : 1969

For sale by the Superintendent of Documents, U.S. Government Printing Office
Washington, D.C. 20402 - Price $2 (paper cover)

Contents

Introduction . :

Acknowledgments

Methods .

Biological notes

Classification . :
Systematic Arrangement. :
Key to Genera of Scenopinidae .
Genus Scenopinus Latreille

Fenestralis group .

Albicinectus group .

Brevicornis group .

Velutinus group é
Genus Caenoneura Krober .
Genus Pseudomphrale Krober
Genus Brevitrichia D. E. Hardy .
Genus Metatrichia Coquillett .
Genus Pseudatrichia Osten Sacken
Genus Belosta D. E. Hardy .
Genus Propebrevitrichia, new genus .
Genus Heteromphrale Krober .
Genus Prepseudatrichia, new genus .
Genus Seguyella, new genus ;
Genus Neopseudatrichia, new genus .
Genus Scenopinula Paramonov .
Genus Rezkiella Paramonov
Genus Stenomphrale Kréber
Genus Paratrichia, new genus
Literature Cited
Index .

Page

ee
PwWWAS BP wpe

16

45

93
110
162
164
165
214
221
274
281
286
286
294
299
304
304
318
320
323
329

' : 7 A a
OT As yes

A Revision of the Scenopinidae (Diptera) of the World

Introduction

Since Kréber’s (1937) last revision of the Scenopinidae, no attempt
has been made to classify this family on a worldwide scale. More
recent papers by Paramonov (1955) on Australian Scenopinidae and
D. E. Hardy (1944a) on North American species have treated with local
faunas and have made valuable contributions to our knowledge.
Krober (1913, 1914a, and 1925) included many illustrations of the
head and wings of previously described and new species. D. E. Hardy
(1944a, 1944b, and 1960) is the first author to illustrate extensively
the external genital structures as well. No attempt has been made, up
to the present, to make detailed comparative studies of both external
and internal structures of the male and female genitalia; only a single
illustration of the internal structure of the male was found, that of
Scenopinus fenestralis by Engel (1932).

The use of the genital structures has led to extensive revision of the
family, the erection of a number of new genera, and the synonymizing
of several which had no apparent justification. The use of the internal
genital structure has been invaluable in separating species whose
visible external features are so close that they could not otherwise
be separated.

The type and allotype of every available described species has been
figured and any discrepancies in the original descriptions have been
noted. Where types have been lost, topotypes agreeing with published
figures have been illustrated.

In the present classification, the family includes 16 genera, four
subgenera, and 214 species. The genus Lagarinus Enderlein (1913),
included by Kréber (1914a), was placed in synonymy with the genus
Chiromyza in the family Stratiomyidae by G.H. Hardy (1921) and Bezzi
(1922). Of the 13 genera listed by Kroéber (1937) only six remain. Two
new genera were erected by D, E. Hardy (1944a) and two by Paramonov
(1955), while an additional five genera and four subgenera have been
erected by me. The following genera have been removed from Kroéber’s
list: Cerocatus Rondani (1848) belongs in the Therevidae as indicated
by Rondani’s plates, and thus will not be treated in this paper.
Omphrale Meigen (1800) has recently been suppressed by the Inter-

1

2 U.S. NATIONAL MUSEUM BULLETIN 277

national Commission on Zoological Nomenclature (ICZN, 1963b) and
the species described in this genus revert to Scenopinus. Lepidomphrale
Kroéber (1913), with the single species niveus Becker, is returned to
Scenopinus where it agrees in all respects with other members of the
Albincinctus group. Archiscenopinus Enderlein (1914), with the single
species niger DeGeer, is in all ways, except for the widely separated
eyes of the male, similar to other members of the genus Scenopinus in
the Fenestralis group. Paromphrale Kréber (1937) containing the
species glabrifrons and antennatus show no genitalic characters that
would separate them from other members of the genus Scenopinus and
other members of the Fenestralis group. Omphralosoma Kréber (1937)
cannot stand because sguamosa and albifasciatus agree more closely
with the Albicinctus and Velutinus groups respectively, than with each
other, and the presence of scales on the body hardly warrants their
separation. Lucidomphrale Krober (1937), with the single species
lucidus, agrees with the members of the Albicinctus group except for
the long aedeagal parameres of the male.

Acknowledgments

This study has taken several years to accomplish and would have
been impossible without the cooperation and helpful suggestions of
many colleagues. I wish particularly to thank Dr. Willis W. Wirth
and Curtis Sabrosky of the U.S Department of Agriculture, Washing-
ton, D. C., for their enthusiastic encouragement of this study and for
their many helpful suggestions in the preparation of this manuscript;
Dr. George M. Worrilow, Dean, School of Agriculture, University of
Delaware, and Dr. Morris Cover, Director, Delaware Agricultural
Experiment Station, for their assistance in obtaining grants from the
General Faculty Research Funds and General Research Funds—
Agricultural Experiment Station, which made possible an opportunity
to visit museums in England, France and Belgium, to study the type
specimens not available on loan; Dr. Dale F. Bray, Chairman, Depart-
ment of Entomology and Applied Ecology, University of Delaware,
for his encouragement and support of this study, Publication No. 366
of the Department of Entomology and Applied Ecology.

I also wish to express my appreciation to the following persons and
institutions for the unselfish loan of material, including types with
permission to dissect genitalia, which helped to make this study possi-
ble. The abbreviations listed below are used in parentheses throughout
this text.

AMB Allan M. Barnes collection, State of California Department of Public
Health, Berkeley, Calif. (A. M. Barnes).

AMNH American Museum of Natural History, New York, N. Y. (J. G.
Rozen).

AMS
ANS
BMNH

CAS
CDA
CNC

CMP
CSC

CS1RO

CSU
CU
CUC
DEI

DZSA

IEE
INHS
IOC
IRSNB

KSU
MCSNG
MCSNM
MCZ

MNH
MNHN

MRAC
MTJ

MZU
NMB
NMP
NMYV
OSU
PAS
PDA

PHT
PU

SEE
SMIJ

SCENOPINIDAE OF THE WORLD 5

Australian Museum, Sydney, Australia (D. K. McAlpine).

Academy of Natural Sciences, Philadelphia, Pa. (H. J. Grant, Jr.).

British Museum (Natural History), London, England (J. P. Doncaster,
Keeper, H. Oldroyd, and J. C. Deeming).

California Academy of Sciences, Golden Gate Museum, San Francisco,
Calif. (P. H. Arnaud, Jr.).

State of California, Department of Agriculture, Sacramento, Calif.
(M.S. Wasbauer).

Canadian National Collection, Canada Department of Agriculture,
Ottawa, Canada (J. G. Chillcott).

Carnegie Museum, Pittsburgh, Pa. (G. Wallace).

Curtis Sabrosky collection, U.S. National Museum, Washington, D.C.
(C. Sabrosky).

Commonwealth Scientific and Industrial Research Organization,
Canberra, A.C.T., Australia (K. H. L. Key, Curator, and D. H.
Colless) .

Colorado State University, Fort Collins, Colo. (F. A. Lawson).

Cornell University, Ithaca, N.Y. (H. Dietrich).

Cairo University, Cairo, Egypt (M. Hafez).

Deutsches Entomologisches Institut, East Berlin, D.D.R. (W. Hennig
and G. Petersen).

Departmento de Zoologia, Secretaria da Agricultura, Sao Paulo,
Brazil (M. Carrera and N. Papavero).

Instituto Espanol de Entomologia, Madrid, Spain (E. M. Agacino).

Illinois Natural History Survey, Urbana, Ill. (L. K. Gloyd).

Instituto Oswaldo Cruz, Rio de Janeiro, Brazil (H. 8. Lopes).

Institut Royal des Sciences Naturelles de Belgique, Brussels, Belgium
(J. Verbeke).

Kansas State University, Manhattan, Kans. (N. Marston).

Museo Civico di Storia Naturale, Genova, Italy (D. Guiglia).

Museo Civico di Storia Naturale, Milan, Italy (C. Conci).

Museum of Comparative Zoology, Cambridge, Mass. (P. J. Darlington,
JT):

Musei Nationalis Hungarici, Budapest, Hungary.

Muséum National d’Histoire Naturelle, Paris, France (A. 5S. Bala-
chowsky, Director and L. Matile).

Musée Royal del’ Afrique Centrale, Tervuren, Belgium (P. Basilewsky).

Maurice T. James collection, Washington State University, Pullman,
Wash. (M. T. James).

Musée Zoologique de |’ Université, Strasbourg, France (F. Gouin).

Naturhistorisches Museum, Basel, Switzerland (F. Keiser).

Natal Museum, Pietermaritzburg, South Africa (B. R. Stuckenberg).

Naturhistorisches Museum, Vienna, Austria (M. Beier).

Oregon State University, Corvallis, Oreg. (J. D. Lattin).

Polish Academy of Sciences, Warsaw, Poland (A. Draber-Monko).

Pennsylvania Department of Agriculture, Harrisburg, Pa. (F. B.
Negley).

P. H. Timberlake collection, University of California, Riverside,
Calif. (P. H. Timberlake).

Purdue University, Lafayette, Ind. (L. Chandler).

Société Entomologique d’Egypte, Cairo, Egypt (A. Alfieri).

Science Museum, The Institute of Jamaica, Kingston, Jamaica
(THs Farr):

+ U.S. NATIONAL MUSEUM BULLETIN 277

SMNH Staatliches Museum fiir Naturkunde, Hamburg, Germany.
SMNS Staatliches Museum fiir Naturkunde, Stuttgart, Germany (E. Lind-

ner).

TAES Texas Agricultural Experiment Station, College Station, Tex. (H. P.
Burke).

UAriz University of Arizona, Tucson, Ariz. (F. G. Werner).

UCalB University of California, Berkeley, Calif. (P. D. Hurd, Jr. and J. A.
Powell).

UCalD University of California, Davis, Calif. (A. T. McClay).

UGa University of Georgia, Athens, Ga. (H. O. Lund).

UKan University of Kansas, Snow Museum, Lawrence, Kans. (G. W. Byers).

UNebr University of Nebraska, Lincoln, Nebr. (W. T. Atyeo).

UQue University of Queensland, Brisbane, Australia (F. A. Perkins and
T. E. Woodward).

USFS United States Forest Service; Berkeley, Calif. (C. B. Eaton); Moscow,

Ida. (M. M. Furniss); Albuquerque, N. Mex. (C. L. Massey);
Missoula, Mont. (R. E. Denton).

USNM United States National Museum, Smithsonian Institution, Wash-
ington, D.C. (J. F. Gates Clarke, Curator).

UZM Universitetets Zoologiske Museum, Kobenhavn, Denmark (8S. L.
Tuxen).

VMM Victoria Museum, Melbourne, Australia (C. W. Brazenor).

WSU Washington State University, Pullman, Wash. (M. T. James).

ZMH Zoologisches Museum der Humboldt Universitat, East Berlin,
D.D.R. (F. Peus and H. Schumann).

ZMUA Zoodlogisch Museum der Universiteit van Amsterdam, Amsterdam,

Holland (W. N. Ellis).

ZMUH Zoological Museum of the University, Helsinki, Finland (W. Hack-
man).

ZSBS Zoologische Sammlung des Bayerischen Staates, Munich, Germany
(W. Forster).

Methods

All figures were drawn using a binocular microscope provided with
an ocular grid. Scale marks on all original figures represent one-half
millimeter; the shorter of the two applies to the wing and lateral and
frontal aspects of the head; the longer applies to the genitalia.

The genitalia were removed from the specimens after the abdomen
was softened with water plus liquid detergent, boiled in KOH, and
drawn under water. Two or three views of each male genitalia were
made: lateral, ventral (physically dorsal), and posterior where this
would show added characters that would aid in identification; see
figure 1.

For the females, dorsal, ventral and lateral views of the 8th and
9th segments and roof of the bursa or other internal structures have
been illustrated where necessary; see figure 1.

All dissected parts have been stored in glycerin in microvials
attached to the specimen.

SCENOPINIDAE OF THE WORLD 5

Ficure 1.—Genitalic structures used in identification: aed., aedeagus; aed. pmr., aedeagal
paramere; acc. gl., accessory gland; ap., apodeme; bursa, bursal cavity of female; ej.
syr., ejaculatory syringe; 9th clsp., 9th segment clasper; 9th st., 9th sternum; 10th st.,
10th sternum; 9th t., 9th tergum; 10th t., 10th tergum.

Lateral and frontal aspects of the head and an enlarged view of
the antenna were drawn for each sex, and a typical wing figure was
made wherever possible.

Biological Notes

Very little is known of the biology of this interesting family. A few
of the well known, widely distributed species, such as Scenopinus
fenestralis and S. glabrifrons, have well documented life histories and
have been associated with their hosts. From the meager records that
are available, it is clear that the larvae of the Scenopinidae are all
predacious on other insects. A number of species are predacious on
the larvae of dermestids and have been commonly collected at the
windows of storehouses, feed mills, and museums. Members of the
genus Pseudatrichia have been reared from the nests of wood rats,
where they were feeding on the larvae of fleas and lice. Several species
have been taken as larvae or pupae from the nests of birds and reared
out. Members of the genus Belosta from the western United States
have been associated with bark-infesting beetles attacking pines and
hardwood, and one species has been taken from a termite nest. The

6 U.S. NATIONAL MUSEUM BULLETIN 2:77

members of the genus Prepseudatrichia from Africa have all been
reared from larvae collected in the wood of Acacia and Sterculia
attacked by buprestid larvae.

Information concerning the habitat of the larvae or the hosts on
which they feed has been recorded for less than ten percent of the
known species of Scenopinidae. From the meager records at hand,
however, some important clues to possible areas of concentration arc
indicated. Among the more obvious localities are: where dermestids
occur, such as grain storages, warehouses and museums; birds’ nests,
particularly those occupied for relatively long periods of time; the
nests or dens of animals; scar wood or under bark attacked by wood
boring larvae; and the nests of termites.

The larvae are elongate, light bodied, hard, smooth, and wormlike
with an elongate, pointed, strongly sclerotized, yellow head. When
these larvae are found they should be maintained with the host
culture, which should be held for a month or two after the emergence
of the host adult, as the Scenopinid adults do not emerge until the
next generation of the host has begun development. The pupae of the
Scenopinidae appear very spiny, particularly along the abdominal
segments.

The scarcity of specimens in most collections appears to be the
result of failure to collect in the right place at the proper time. Most
of the specimens observed have been obtained by sweeping foliage or
flowers which strictly limits the chances of contact. The adult period
appears to be relatively short in most cases and occurs at a regular
time each year. There is good evidence that in areas with uniform
plant or animal associations, large numbers of adults are present at
the same time. In one instance, in California, over 150 specimens of a
single species were taken on a two-day period along a distance of
more than 100 miles. Other extensive series have also been seen that
were taken in a short period by field crop inspectors sweeping the
same crop over an extensive area. In other instances, members of
the same species have been taken at the same locality in successive
years on about the same date. The use of some permanent type of
trap such as a Malaise trap would probably result in better sampling
than that accomplished by most collectors with hand nets.

Classification

Provisional keys have been made up for those genera of Scenopinidae
containing more than two species with the exception of Caenoneura
and Pseudomphrale. I believe that the species listed in this paper
comprise only asmall part of the ultimate known fauna of Scenopinidae.
For this reason, the simple characteristics of wing, halter and antennal
color have been used as the primary basis for separation to species.

SCENOPINIDAE OF THE WORLD 7

The taxonomist is, therefore, cautioned to check carefully the
figures and descriptions of specimens which run through the keys.
Unless the specimen comes from near the type area, it should be
checked critically to see that it fits all the characteristics, as most
species have only a limited distribution. For positive identification,
the internal structures both of male and female should be checked
against the figures.

In the following classification, the species have been grouped into
genera on the basis of all available characters with particular em-
phasis on the genital characteristics which have not previously been
used for identification of members of this family.

All males of this family examined to date have an internal ejacula-
tory syringe surrounded by accessory organs and apodemal structures
to support the muscles of the ejaculatory syringe and aedeagal extensors.

The females of Scenopinus are difficult to separate except by internal
genitalic structures or the conformation of the eighth sternum; the
shape of the bursal cavity is extremly helpful in identification.

In no case except S. niger has a male been examined where the eye
was not divided into an upper portion with larger facets and a lower
portion with smaller facets. In almost all cases, the lower portion of
the eye appears darker than the upper part. All females have eyes
with uniform facets in their make-up.

Systematic Arrangement

Genus Scenopinus Latreille

FENESTRALIS group

1. albicomus, new species Tunisia

2. antennatus (Kréber) 1914b, p. 74. (Omphrale) #7 Algeria

3. aquelonius, new species 9 British Columbia,
Canada

4. breviterminus, new species &@ 9 Calif., U.S.

5. cavifrons (Krober) 1937, p. 231. (Omphrale) Canada

6. efflatowni, new species 2 Egypt

7. fenestralis (Linnaeus) 1758, p. 597. (Musca) #9 Worldwide

8. glabrifrons Meigen 1824 p. 114. 7° Europe; N. Africa;
N. America;
Australia

9. griseus (Kroéber) 1913, p. 191. (Omphrale) & Hungary

10. japonicus (Séguy) 1920, p. 318. (Omphrale) 9 Japan

11. lesinensis Strobi 1902, p. 474. 9 Austria

12. microgaster (Séguy) 1948, p. 155. (Omphrale) @ 2 China

13. nidorcaupulus, new species 9 Colo., U.S.

14. niger (DeGeer) 1776, p. 188. (Nemotelus) #2 Europe

15. opaculus Loew 1873, p. 145. #7? Samarkand (U.S.8. R.)

16. physadius (Séguy) 1930, p. 111. (Omphrale) & 2 Morocco

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U.S. NATIONAL MUSEUM BULLETIN 277

saini, new species

schroedert (Kroéber) 1913, p. 201. (Omphrale) 9°
sinensis (Kréber) 1928a, p. 1. (Omphrale) ©
undulafrons, new species ©

varipes Loew 1873, p. 148. ¢
vitripennis Meigen 1824, p. 115. °

ALBICINCTUS group

aethiopicus (Lindner) 1955, p. 20. (Omphrale) 9°
albicinctus (Rossi) 1790, p. 69. (Afusca) #9

angustifrons (Kroéber) 1913, p. 197. (Omphrale) 9°

anthrax, new species o 9

balteatus Lamb 1922, p. 363. 9°
birot Kertész, 1899, p. 173. 7 2

bolgarti, new species oc @

bouviert (Séguy) 1920, p. 317. (Omphrale) °
brunneus (Kréber) 1913, p. 192. (Omphrale) °
bulbapennis, new species o 9

canarius, new species

curtiptlosus, new species 2

darwini, new species 9?

evansi, new species 9°

flavipes (Kréber) 1939, p. 398. (Omphrale) 9°
fragosifrons, new species o&* 9

gromier?, new species °

inquilinus Séguy 1931, p. 113. #7 9
katbergi, new species 9

ketserz, new species 9?

limpidipennis Loew 1874, p. 416. 7 9°
lucidus Becker, 1902, p. 39. & 9

madagascariensis Enderlein 1934, p. 429. °
minutus, new species o 9

monodi (Séguy) 1933, p. 75. (Omphrale) # 9
niweus Becker, 1907, p. 61. &

norrist, new species

oldenbergi (Kréber) 1913, p. 190. (Omphrale) 9?
opacus deMeijere 1924, p. 14. #@ 9?

palmulapal pus, new species °

papuanus (Kroéber) 1912, p. 25. (Omphrale) @ @

patrizi (Séguy) 1932, p. 499. (Omphrale) 9°
perkinst G. H. Hardy 1942, p. 202. 7 9
phaidimos, new species &@ Q

spurrelli, new species 9

squamosus Villeneuve 19138, p. 111. 9°

Egypt

Costa Rica

China

British Columbia,
Canada

Russia

Europe

E. Africa

S. Europe

S. Africa

Magadascar;
Mozambique

Seychelles

Australia; New
Guinea

W. Australia

Fr. Congo; ? Paris

E. Africa

Spain

Canary Islands

Egypt

N. Australia

N. Rhodesia

Kenya

W. Australia

Cameroun

Mozambique

S. Africa

Madagascar

N. Persia

Hawaii; Belg. Congo;
Egypt; Canary
Islands

Madagascar

Australia

Soudan

Tunisia

Australia

Germany; ? Africa

Sumatra

S. Africa

Hawaii; Formosa;
New Guinea; Ceylon

Libya

Australia

Turkey

Australia

Algeria

59.
60.
61.
62.
63.
64.
65.
66.
67.
68.
69.
70.
dale
d2.
73.

SCENOPINIDAE OF THE WORLD

BREVICORNIS group

brevicornis Loew 1873, p. 146. &@ 9°

fraternus (Kréber) 1924, p. 81. (Omphrale) # 9°

fryeri, new species 9

halteralis Frey 1936, p. 53. 0

lincinus, new species °

longiventris (Kréber) 1913, p. 206. (Omphrale) 9°

minusculus (Séguy) 1934, p. 74. (Omphrale) # 9°

nitidifrons (Kréber) 19138, p. 194. (Omphrale) °

nitidulus Loew 1873, p. 149. @ 2

parallelus, new species

pilosus (Séguy) 1930, p. 111. (Omphrale) ¢&

reduncus, new species 9

tarsalis (Kréber) 1913, p. 196. (Omphrale) # 9°

turneri, new species

unifasciatus (Kréber), 1913, p. 195. (Omphrale)
ore

VELUTINUS group

adventicius, D. E. Hardy 1960, p. 329. #7 @

alalacteus, new species o& 9

albifasciatus (D. E. Hardy) 1944a, p. 41. (Om-
phralosoma) &* @

barberi, new species @ 9°

beamert (D. E. Hardy) 1944a, p. 43. (Omphrale)
oe

beameri var. fuscus (D. E. Hardy) 1944a, p. 43.
(Omphrale) & 9°

bryanti, new species 9°

buscki, new species @& &

butleri, new species

chico, new species co °

cochisei, new species co 9°

cook?, new species

cretatus, new species o &

curticornis (Kréber) 1913, p. 202. (Omphrale) @
electus Adams 1904, p. 445. &

femoratus Macquart 1835, p. 7. 9

fijianus (Kréber) 1939, p. 399. (Omphro,le) 9

flandersi, new species @ 9°

gossypius, new species co 2

hagai, new species o 9

howden?, new species 9

Kuitertt (D. E. Hardy) 1944%, p. 46. (Omphrale)
eS

magdalenai, new species oj 9°

magnicornis (Krober) 4913, p. 202. (Omphrale) 9

Turkestan; Egypt
Egypt

Seychelles
Canary Islands
Madagascar
Ceylon
Madagascar

K. Africa

Iran; Crete; Egypt
Rio de Oro
Morocco
Madagascar
Eritrea

S. Africa

Greece

Hawaii

Ida., Nev., U.S.

Calif., Ariz., N. Mex.,
UES:

Ariz., U.S.

Calif., Nev., U.S.

Calif., U.S.

Ariz., U8.

Panama; J‘amaica

Ariz., U.‘s.

Calif., Ariz., Nev.,
U's.

Avjz., Calif., U.S.

Ariz., U.S.

Ida., Nev., U.S.

Brazil

Ariz., Calif., U.S.;
Baja Calif., Mexico

Brazil

Fiji Islands

Calif., U.S.

Southern U.S.

Ariz., U.S.

Ariz., U.S.

Texas, Ariz., Calif.,
N.M., U.S.

Mexico

Costa Rica; Peru;
Brazil

10

98.

99:
100.

101.

102.
103.
104.
105.
106.

107.
108.

109.
tO:

111.

112.
113.

114.
15,
Ig

1173
118.

LIS:
120.
121.
122.
123.
124.
125.
126.
127.
128.
129.
130.
131.

132.
133.
134.
135.

U.S. NATIONAL MUSEUM BULLETIN 277

mirabilis Adams 1904, p. 445. @ 9°

nubilipes Say 1829, p. 170. &
pecki, new species o @

pygmaeus Loew 1857, p. 90.

rossi, new species co P

schulzi Enderlein 1934, p. 429. @ 9°

seftont, new species o 9

serratus, new species

valgus (D. E. Hardy) 1944a, p. 50. (Omphrale)

oo?

vanduzeei, new species co
velutinus (Kréber) 1913, p. 203. (Omphrale) 7 9°

weemsi, new species co 9°

werneri, new species

whittakeri (James) 1955, p. 47. (Omphrale) & @
Genus Caenoneura Krober

robusta Kroéber (1924), p. 75. #7

nigra, new species 9

Western U.S.;
Canada

Eastern U.S.

Alberta, Canada;
Wyo., U.S.

Central America;
West Indies;
Surinam

Mexico

Brazil, S. America

Mexico

Ariz., U.S.

Calif., Wash., U.S.

Mexico

Costa Rica; Mexico;
Jamaica

Fla., U.S.

Ariz., U.S.

Wash., U.S.

Egypt
Spain

Genus Pseudomphrale Kroéber

clausa (Loew) 1873, p. 150. (Scenopinus) &
crenata (Becker) 1918, p. 550. (Omphrale) 9°
dubiosa (Becker) 1913, p. 551. (Omphrale) &
longirostris (Becker) 1913, p. 549. (Omphrale) 9
ralaestinensis Kréber 1937, p. 229. &

Genus Brevitrichia Hardy

addacifoxs, new species o
arnaudi, Inw species &
aspinosa, Nev species 2
beameri, new Species ot 2
boharti, new speGes 3 9
castanea, new spechs
coquilletiz, new specie g 9
davist, new species o*
dicksoni, new species ?
downeyi, new species

flocki, new species ?

griffini, new species o
griseola (Coquillett) 1900, p. 501. (Pseudatrichia)

oe

halli, new species & 9

helenae (James) 1938, p. 22. (Pseudatrichia) 3
hodgdeni, new species o& 9

insulana (Cole) 1923, p. 462. (Pseudatrichia) % 9

Tran
Tran
Tran
Tran
Palestine

Calit., U.S.

Baja Calif., Mexico
Ariz., U.S.

N. Mex., U.S.
Calif., U.S.

Ida., U.S.

Calif., U.S.

Utah, U.S.

Calif. URS:

Calif., U.S.

Ariz., U:S:

Calif., U.S.

N. Mex., Tex., U.S.

Calif., U.S.
Colo., U.S.
Calif., U.S.
Baja Calif., Mexico

136.
137.
138.
139.
140.
141.
142.
143.
144.
145.

149.

150.

Se eR
en cr tn
wm CO bO

-
ou
ao

166.
167.
168.
169.
170.
171.
172.
173.
174.
175.
176.

SCENOPINIDAE OF THE WORLD 11

kerni, new secies 9 Calif., U.S.
melanderi, ney species 9° N. Mex., U.S.
minuta, New Species oP Ariz., Calif., U.S.
oculwirida, newspecies @ Baja Calif., Mexico
ordwayi, new speies oc 2 Ariz., U.S.
pruinosa, new speties Baja Calif., Mexico
schlingeri, new speGes o 9 Calif., U.S.
scitulaesca, new speces 9 Texas, U.S.
timberlakez, new specie oY Calif., U.S.
yucatani, new species d 9 Mexico; Guatemala

Genus Metatrichia Coquillett

bulbosa (Osten Sacken) 1877, p. 275. (Scenopinus) U.S.; Mexico
ay

lophrysoma (Speiser) 1920, p. 216. (Pseudom- 8. Africa, Congo
phrale) °

robusta Kréber 1913, p. 180. 29 Paraguay; Brazil;
Argentina

stevenson? (Bezzi) 1925, p. 96. (Psudomphrale) 79 Rhodesia; Kenya;
S.W. Africa

waterhouset (Paramonov) 1955, p.640. (Pseudom- Australia
phrale) &

Genus Pseudatrichka Osten Sacken

albocincta Van Duzee, 1926, p. 164. ‘ Calif., U.S.
atombomba, new species o 9 N. Mex., U.S.
bakeri, new species 9° Calif., U.S.
barnesi, new species o 2 Calif., U.S.
biacristerna, new species 2 N. Mex., U.S.
bryanti, new species 9 Ariz., U.S.
bupennis, new species Calif., U.S.
cajoni, new species o ° Calif., U.S.
cloudcrofti, new species 9 N. Mex., U.S.
cockerellt, new species o 9 Colo., U.S.
concava, new species ° Calif., U.S.
convexa, new species Colo., U.S.
eaithales, new species 9 Colo., U.S.
evergreeni, new species Q Colo., U.S.
garretti, new species 9 British Columbia,
Canada
gracilipennis, new species Calif., U.S.
howdent, new species 9 Ariz., U.S.
jamesi, new species @ Colo., U.S.
leechi, new species 2 Calif., U.S.
longiventris, new species co ? Costa Rica
longurio (Loew) 1866, p. 42. (Atrichia) 9° Mexico
melanderi, new species oc 9 Calif., U.S.
morlani, new species o 9 N. Mex., U.S.
nordeni, new species 9 Calif., U.S.
parva LD. E. Hardy, 1944a, p. 36. 9 Ariz., U.S.
parvipennis, new species Calif., U.S.

313-168 O—69

9
“

12

Like
178.
L79.
180.
181.
182.
183.

184.
185.

186.

187.
188.

189.
190.

191.

192.

193.
194.
195.

196.
1G.
198.

199:
200.

202.

U.S. NATIONAL MUSEUM BULLETIN 277

peromysci, new species o Q Calif., U.S.
punctulata D. E. Hardy, 1944b, p. 104. #7 9 Brazil
sabroskyi, new species 9 Oreg., U.S.
saccharcupa, new species 9° Calif., U.S.
toupeta, new species Colo., U.S.
truncata, new species Mexico
unicolor Coquillett 1900, p. 500. 79 N. Mex., U.S.

Genus Belosta ID. E.. Hardy

albipilosa D. KE. Hardy, 1944a, p. 38. a9 Calif., Idaho, U.S.
pilosa (Coquillett) 1903, p. 102. /(Pseudatri- Ariz., Calif.,

chia) 78 Colo., U.S.
telfordi, new species 9 Ariz., U.S:
termitophaga, new species ° Ariz., U.S.
viticolapennis, new species ] Ariz., U.S.

Genus Propebrevitrichia, new genus

stuckenbergi, new species o S. Africa
turneri, new species oP / S. Africa

Genus Het¢ezromphrale Kréber
cyanops (Edwards) 1932, / p. 259. (Pseudom- Uruguay

phrale) & |
chilensis (Kréber) 1928b, p/. 31. (Pseudatrichia) 9 Chile

Genus Prejpseudatrichia, new genus

mateui, new species oP Sahara
stenogaster (Séguy) 1931, p. 114. (Scenopinus) Mozambique
violacea, new species ? Chad

Genus Seguyella, new genus

chinchona, new species 9° India
galactica (Séguy) 19°38, p. 334. (Omphrale) &# Kenya
turneri, new species oF S. Africa

Genus Neopseudatrichia, new genus

canei, new species S. Australia
kewi, new species oc 2 Australian Capital
Territory, Australia
mariaensis (G. H. Hardy) 1938, p. 419. (Pseuda- Tasmania
trichia) 3°

Genus Scenopinula Paramonov

pallidipennis Paramonov 1955, p. 651. 9 New South Wales,
Australia

203.
204.

206.

207.
208.
209.
210.

to

SCENOPINIDAE OF THE WORLD 13

Genus Riekiella Paramonov

bicornis Paramonov 1955, p. 653. 9° Queensland, Australia
collessi, new species co ° New South Wales,
Australia
complexa, new species Australian Capital
Territory, Australia
dycez, new species 9° New South Wales,
Australia
longiventris, new species ° W. Australia
quadrifida, new species o 9 Queensland, Australia
taylori, new species Victoria, Australia
uncata, new species o 9 New South Wales,
Australia

Genus Stenomphrale Kroéber

flavoscutellata (Kréber) 1929, p. 80. (Pseudom- Egypt
phrale) 3°

teutankhameni (Kroéber) 1924, p. 72. (Pseudom- Egypt
phrale) 3&7 °

Genus Paratrichia, new genus

lobosa, new species Australian Capital
Territory, Australia
westralica (Paramonov) 1955, p. 641. (Pseudom- W. Australia
phrale) &

Key to the Genera of Scenopinidae

Cell R5 open to the tip of the wing.
Cell R5 closed and petiolate .
Vein M1-+2 fading before edge of wing .
Vein M1+2 reaching edge of wing . .
Vein R4 branching before the middle of cell RE
Scenopinula Paramonov (p. 304)
Vein R4 branching near or beyond the middle of cell R5, vein M1+2 curving
toward vein R5. ... =... . . Riekiella Paramonov (in pete - ve
Vein M4+Cu normal, reaching edge of wing.
Vein M4+Cu fading just beyond the m crossvein
Seguyella, new genus (p. 294)
Head longer than high, thorax depressed, long slim flies resembling Pseuda-
trichia. . . . . . Prepseudatrichia, new genus (p. 286)
Head higher ‘aon long. body, not depressed, abdomen broad
Scenopinus Latreille, 6 (p. 14)
Large robust flies, with long slender antennae, vein R4 branching from R5
before middle of cell R5; male 9th tergum 4-lobed
Fenestralis group (p. 16)
Medium sized flies, variable antennae, vein R4 branching from near center
of cell R5; male 9th tergum 2-lobed, open ventrally
Albicinetus group (p. 45)
Medium sized flies, pear-shaped antennae, vein R4 branching from near
center of cell R5; male 9th tergum with basal portion extending ventrally,

wm Co ST bo

14

10.

11.

12.

13.

14.

16.

Wis

U.S. NATIONAL MUSEUM BULLETIN 277

open distally; female 8th sternum longer than tergum, 9th tergum usually

with short spines .. . ise. J) Brevicornis'groups(py 93)

Medium to small flies, short aniennee: vein R4 branching beyond the middle

of cell R5; male 9th tergum 2-lobed closed below

Velutinus group (p. 110)

Vein R5+M1-+2 ending at tip of wing. . . 3 Lie eer NE Lee TSS
Vein R5+M1-+2 bending sharply to end in bading edee

Caenoneura Krober (p. 162)

Longslender black ‘flies . \\. 4.0 iA goat at BR a OU ee.
Brown, tan or gray flies . . . Re ee, OF say Re Pr aa EY ie SER OL
Head longer than high, body Wabeoua ave Torii, Aceh, mae eemraten. NG

Head higher than long, mouthparts atrophied, Taek
Belosta Hardy (p. 274)
Ninth tergum of male about as long as high and as long as the 10th tergum.
Female 9th tergum and sternum subequal, tip of cell R5 blunt
Pseudatrichia Osten Sacken (p. 221)
Ninth tergum much taller than long, 10th tergum longer than 9th. Female
10th tergum with thick spines, tip of cell R5 sharp
Neopseudatrichia, new genus (p. 299)
Large robust bodied flies) «2 a) 5. «Bea fel pak Soli eee oe eng eons
Smaller flies... . Re. sae
Very large flies with broad blunt abdomen in both sexes, frontal area swollen
so that antennae project from middle of head; hairs often flat, scale-like
Metatrichia Coquillett (p. 214)
Smaller flies, with blunt abdomens, frons not swollen, male genitalia with

flange-like lobes on inside of 9th tergum . . Paratrichia, new genus (p. 320)
Small flies with pollinoseiseales on thorax. ............ ./14
Small flies, black, shining , . . by San ei ROR ilk aed
Wing with vein R4 branching fom cell R5 near mule, Hace 5 Mga euae one LO
Wing with vein R4 branching from the cell R5 near the middle . . . . 16

Males with two long aedeagal spines, females with 8th sternum excavated on
distal margin, 9th tergum often with row of stiff spines on posterior margin
Brevitrichia Hardy (p. 165)
Males with short aedeagal parameres, females with long pointed 8th sternum,
9th tergum with stiff spines. . . . . . . Heteromphrale Kroéber (p. 286)
Males with 9th tergum as two flaps covering genitalia, females with spines on
9th tergum .. . . .. . . . Propebrevitrichia, new genus (p. 281)
Males with 9th femeaan as four long lobes, females without spines on 9th
tergum .. . . . . Riekiella Paramonov (in part) (p. 304)
Antennae pointed: vein “R4 br anching from distal third of cell R5

Stenomphrale Kroéber (p. 318)

Antennae blunt, vein R4 branching from before the middle of cell R5
Pseudomphrale Kroéber (p. 164)

Genus Scenopinus Latreille

Omphrale Meigen, 1800, p. 29 (rejected name).
Scenopinus Latreille, 1802, p. 463.

Atrichia Schrank, 1808, p. 54.

Cona Schellenberg, 1803, p. 66.

Hypseleura Meigen, 1803, p. 273.

Scenopoeus Agassiz, 1846, p. 333.

SCENOPINIDAE OF THE WORLD 15

Astoma Lioy, 1864, p. 762.

Scaenopius Dalla Torre, 1878, p. 161.
Lepidomphrale Kroéber, 1913, p. 182.
Archiscenopinus Enderlein, 1914, p. 25.
Lucidomphrale Krober, 1937, p. 222.
Omphralosoma Krober, 1937, p. 222.
Paromphrale Kroéber, 1937, p. 222.

Type-species, Musca fenestralis Linnaeus (monobasic). This genus
is the largest, and by far the most cosmopolitan, genus, having repre-
sentatives in all geographic regions. Because of its size (111 species)
it has been subdivided into four infrageneric groups each with rather
distinct continental distribution: Fenestralis, Albicinctus, Brevicornis,
and Velutinus.

FENESTRALIS GROUP.—The members of this group are characterized
by a head that is higher than long, generally with long slender antennae,
R4 branching from R5 at or before the middle of the cell; generally
robust and of large size. The males have in common a four-lobed ninth
tergum and a three-pronged genital apparatus consisting of a median
aedeagus and two lateral aedeagal parameres. The females of this group
have a large membranous bursal area; see figures.

Geographically this group is Palearctic with an extension down the
west coast of the North American continent. Scenopinus fenestralis and
S. glabrifrons have become world-wide in distribution as a result of
transport in trade goods. S. niger, contrary to previous reports, is
strictly limited to the European continent.

ALBICINCTUS GROUP.—The members of this group are characterized
by having the head higher than long, the antennae only twice as long
as wide, the vein R4 branching from R5 near to or beyond the middle
of the cell and their size generally smaller than that of members of
the Fenestralis group. The males are characterized by a bilobed ninth
tergum which does not overlap or close on the ventral side (dorsal
on the specimen) leaving the genitalia exposed. The aedeagus is vari-
able, with the parameres often longer than the aedeagus proper. The
bursa of the female is generally as in the Fenestralis group but
somewhat wider than long.

The members of this group are found in the Ethiopian, circum-
Mediterranean area of the Palaearctic, the Oriental and Australian
regions including Hawaii.

BREVICORNIS GROUP.—The members of this group have the head
higher than long, the antennae generally pear-shaped. The wing has
R4 branching from R5 at or near the middle of the cell. M1 tends to
run parallel to R5 so that they are widely separated at the wing mar-
gin. The male terminalia, though open ventrally, have the proximal
portion of the ninth tergum extending ventrally to partially cover the

16 U.S. NATIONAL MUSEUM BULLETIN 277

venter. In the female, the eighth sternum extends posteriorly beyond
the margin of the tergum and the ninth tergum has randomly placed
coarse hairs; the bursal cavity is longer than wide.

The species placed in this group have been removed from the pre-
ceding group, in some cases on a tentative basis, particularly where
only the male is known. Other species presently included in the
Albicinctus group may belong here but until their females are found
their exact placement is questionable.

The members of this group are found in the circum-Mediterranean
part of the Palaearctic and in the Ethiopian region.

VELUTINUS GROUP.—The members of this group are small to
medium-sized flies with the abdomen broad and rounded. The head as
in the other members of the genus is higher than long, the antennae
are generally short, oval, rarely being over twice as long as wide. Vein
R4 generally branching from R5 well beyond the middle of the cell.
Male ninth tergum bilobed, generally extending ventrally to enclose
the genitalia from view in the pinned specimen. The aedeagus is
generally well developed and is flanked by two aedeagal parameres of
variable length, generally but not always shorter than the aedeagus.
The bursal cavity of the female is small and triangular.

This group is generally limited to the Nearctic and Neotropical
regions with an extension into the Pacific Island area.

FENESTRALIS group

Provisional Keys to Species of the FENESTRALIS Group

NEARCTIC SPECIES
1. Antennae long and narrow
Antennae short, pear-shaped SUD
2. Wings hyaline, halter knob white to brown. Was
Wings smoky gray, halter knob yellow-brown. . . . 9 S. achroederi ieroher
3. Frons shining, halter knob white, legs orange-brown
o' 9S. glabrifrons Meigen
Frons rugose, halter knob brown to white, legs red-brown
9S. fenestralis (Linnaeus)

cares

4. R4, branching before. the middle of cells Roy seca ye ecco = ee
R4 branching at the middle of cell R5. . . . . . 9S. aquelonis, new species
5. JR4 branching from:before'the middle oficell: Rd... 0. 4.224.) et 6

R4 branching from the basal quarter of cell R5
o' 9S. breviterminus, new species
6. Cell R5 narrowing gradually. . ... . . . 9 S. undulafrons, new species
Cell R5 narrowing abruptly attip. . . . . . 9? S.nidorcaupulus, new species

PALAEARCTIC SPECIES (MALES)

1. Wing hyaline ...@ y URE e, S NO GA Berea te vel Te Gene eee Ean ee eee
Wing brown or milky atte :
2. Frons shining, halter knob white .

oo

w

ou

10.

SCENOPINIDAE OF THE WORLD ry

Frons rugose, halter knob variable, white to brown
S. fenestralis (Linnaeus)
Vein R4 branching well before middle of cell R5. .S. glabrifrons Meigen

Vein R4 branching near middle of cell R56. . . . . .S. saini, new species
Wing brown or tinged brown .........2.2.2.2..2202.62.2242.2.5
Wie atic OF ravine tar raee ran, OS tsk tes, AMS i EU. 6 ne wT
Halter knob brown or black-brown. . . ce eens tO

Halter knob white, abdomen with 6 white oral eee . 8S. griseus Kréber
Halter knob black-brown, eyes widely separated, uniform facets, abdomen

normal length. ... . .. . . .S. niger DeGeer
Halter knob brown, eyes means fonehing, Aided into upper and lower facets,

abdomen short and broad. . ...... . . . .S. microgaster Séguy
Halter knob white ... . Seg ne Soy Use ce are GE ee gar at
Halter knob brown on top mente nclone .... . . .S. opaculus Loew
Third antennal segment long and narrow ..........2.2.2.2.2.9
Third antennal segment short, swollen at base. . .S. antennatus (Krdéber)

Abdomen with three stripes, first broad, second and third narrow
S. albicomus, new species

Abdomen with three broad white stripes. . . . . .S. physadius (Séguy)
PALAEARCTIC SPECIES (FEMALES)
Wing hyaline. . . gest Ses Bi: eA a eee Ge ee
Wing brownish or itlicy inte oy saree, ven Ge ee ee BE n ce oe we ee
Frons shining, halter knob white or brown. . . . , ...9
Frons rugose, halter knob variable brown to white . ‘Ss. fenestralis Mannenne)
Halter knob white. . ............ .. .S8. glabrifrons Meigen
granular. . .........4.2. 2... =. =. =. .8. lesinensis Strobl
Halter knob brown. ............. . .S8. vitripennis Meigen
Wiainp Smoky wihibe 4) fac ben eS re cas, OO de cee ee «6 aw oD
yas DIOS errs) ee ree leet a. Sal on ge ie ae Ste ee ede, SO
Halter knob white. ............ .. .8. physadius (Séguy)
Halter knob light tan... . ... . .S. opaculus Loew
Halter knob red-brown, brown or ane brows: antenna variable. . . .7
Halter knob light tan; antenna elongate, red- Soon . S. eflatouni, new species
Halter knob brown or black-brown; antenna variable. . . ...... .8
Halter knob red-brown above, cream below; third antennal segment black,
Srailats sae of we 2 ek oes wee) Ss Lesinensis: Strobl
HAM RnOD DIGWiad i ww 6 ic hn beh lage steel acetate S Phi & eS
Halter knob black-brown .. . le eek, etl ()
Antenna short, oval; vein R4 br ee foo near midale of melt R5, abdomen
ShOrt..+.. = = & ... . . .S. microgaster (Séguy)
Antenna long, narrow; vein RA branchine from basal third of cell R5,
abdomen normal. ... . . . . . .. sinensis (Kréber)
Cell R5 open at end of wing, margin: fel aoe nearly equal to length of r-m
cross vein. . . . . .S. japonicus (Séguy)
Cell R5 closed at end of wing, ee fenathi only half the length of r-m
CYOSS-VEINies Wels ws wer ss ow we 2 eee 2 Se iger (DeGeer)

1. Scenopinus albicomus, new species

FIGURE 2

Ma.r.—A moderately large black-brown fly typical of the group.

Head black-brown; eyes black, tinged with red, the upper two-thirds

18 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 2.—Scenopinus albicomus, new species, male: a, wing; b, c, head, lateral and frontal
aspects; d, enlarged detail of antenna; e-g, lateral, ventral, and posterior aspects of
terminalia.

SCENOPINIDAE OF THE WORLD 19

with large facets and the lower third with fine facets; frons small,
triangular, shining, with a shallow groove on the midline above the
antennae; ocellar tubercle not prominent, ocelli yellow to orange-
yellow. Antennae with first segment brown, short; second segment
twice as large and light brown; third segment brown, long and narrow,
over twice as long as first and second segments combined. Mouthparts
well developed.

Thorax dorsum black-brown covered with short silvery hairs;
posterior rim of humeral callus orange; supra-alar callus yellow.
Wing smoky white, veins tan; stem of halter tan, knob white. Legs,
femora and tibiae of all legs brown, tarsi yellow.

Abdomen first segment white to lateral margins, brown below
scutellum; second segment brown; third segment with a wide band;
fourth and fifth segments with narrower white bands with brown
between; genital segment brown with aedeagus projecting visibly;
see figure for details.

FrmaLe.—Unknown.

Length: Male body 4 mm., wing 3 mm.

Type-locality: Tunis (Tunisia); 2 August 1929 (C. Dumont).

Holotype: Male (MNHN).

2. Scenopinus antennatus (Kréber)
FIGURE 3

Omphrale antennata Kroéber, 1914b, p. 74.
Paromphrale antennata (Kréber).—Kroéber, 1937, p. 214.

Mauxe.—This species is known only from the type. A moderate
sized fly of a red-brown color; eyes divided, with larger upper and

Ficure 3.—Scenopinus antennatus (Krober), male: a, wing; b, c, head, lateral and frontal
aspects; d, enlarged detail of antenna; e-g, lateral, ventral and posterior aspects of ter-
minalia.

20 U.S. NATIONAL MUSEUM BULLETIN 277

smaller lower facets; frons granular with a median depression above
the antennae. The third antennal segment is broader at the base and
shorter than normal for the group.

Thorax red-brown, covered with a thin pile of short hairs. Wings
milky with light brown veins. Halter stem brown, knob cream. The
legs are light brown with the basal portion of the tarsi yellow darkening
distally.

Abdomen red-brown with three white bands. Genital segments as
figured.

Frmate.— Unknown.

Length: Male body 4 mm., wing 2.8 mm.

Type-locality: Biskra, Algeria; May 1891 (Handrich).

Type: (NMV).

3. Scenopinus aquelonius, new species

FIGURE 4

FremaLe.—A large fly with the typical S. fenestralis appearance
except for the antennae which are shortened and pear-shaped.

Head black, eyes red-brown, postocular flange broad and subshin-
ing, frons wide, subshining, with a median groove, a shallow depression
in the lower half. Back of the head concave; mouthparts brown, well
developed; a shiny band of integument between the silvery pilosity
lining the oral cavity and the lower eye margin. The ocellar tubercle
red-brown, set off slightly from the frons; ocelli tan. The first antennal
segment short, black; the second red-brown and twice as long as the
first; the third segment dark red-brown, pear-shaped, twice as long
as broad.

Thorax black; thinly covered with short silvery hairs, a patch of
silvery pile above the red-brown humeral callus; supra-alar callus
orange. Wings smoky brown, veins red-brown; halter stem brown,
knob red-brown. Legs: femora and tibiae red-brown, tarsi brown.

Abdomen broad, dark red-brown; see figure for distinguishing
characters of 8th sternum and bursa.

Maute.— Unknown.

Length: Female body 5 mm., wing 3.5 mm.

Type-locality: Trinity Valley, B.C., Canada; 8 August 1928 (R. T.
Turner).

Holotype: Female (CNC) 8632.

Paratype: Female, same locality, July 1937 (K. Graham) (CNC).

SCENOPINIDAE OF THE WORLD yA

Ficure 4.—Scenopinus aquelonius, new species, female: a, wing; b, c, lateral and anterior
aspects of head; d, enlarged detail of antenna; e, 8th sternum; f, lateral aspects of 8th and
9th segments; g, detail of 9th sternum and bursa.

22, U.S. NATIONAL MUSEUM BULLETIN 277

4. Scenopinus breviterminus, new species

Figure 5

Mate.—A large robust fly similar to S. fenestralis except for head
and genital structures.

Head black-brown; eyes red-brown, darker below; frons elongate,
separating the eyes above by at least the width of the median ocellus,
pitted and with three indistinct shallow grooves on lower frons;
ocellar tubercle black, distinct; ocelli clear in specimen. First antennal
segment small, brown, shining; second segment orange, larger than
the first; third segment red-brown, darker at tip, not twice as long
as wide.

Thorax dorsum black-brown, rugose, thinly covered with silvery
hairs; humeral callus red-brown, light yellow behind, supra-alar callus
red-brown; pleural region red-brown. Wings smoky hyaline, veins
brown; halter stem brown, knob red-brown. Legs all segments light
red-brown.

Abdomen red-brown, segments 3-5 banded with white on pos-
terior margin. Terminalia red-brown; see figure for details. Internal
structure lost through feeding by dermestids. The ninth tergum lobes
are shorter than in typical fenestralis.

Frmaue.—A large black fly resembling fenestralis except for the
antennae which are shorter.

Head black, eyes black; postocular flange flat, dull. Frons with a
median groove from the antennae to median ocellus, slightly swollen
above the antennae, granular, except smooth bands along eye margins.

Thorax black, rugose, subshining, humeral and supra-alar calli orange-
brown; halter stem brown, knob oxblood red, polished. Wing hyaline,
tinged brownish, veins brown. Foreleg with coxa red-brown, tro-
chanter orange, femur red-brown, tibia orange-brown, tarsus straw
colored; middle leg with trochanter orange-brown, femur dark red-
brown, tibia lighter and tarsus straw colored; hind leg with all seg-
ments dark red-brown except tarsus straw colored.

Abdomen dark red-brown; see figure for details of eighth sternum
and bursa.

Length: Male body 4.5-5.0 mm., wing 3.5 mm.; female body 5.7
mm., wing 3.7 mm.

Type-locality: Kerr Ranch, New Mexico; 23 April 1910 (J. D.
Mitchell), on Yucca.

Holotype: Male (USNM) 67453.

Allotype: Female, Carson Pass, California, in fungus (Mont. A.
Cazier) (USNM).

SCENOPINIDAE OF THE WORLD 23

Ficure 5.—Scenopinus breviterminus, new species, male, female: a, wing;

frontal aspects of male head; ¢, e¢, lateral and frontal aspects of female hea
details of male and female antennae; h

terminalia (internal structures destroy
lateral aspect of female 8th and 9th se

b, d, lateral and
d; f, g, enlarged
-J, posterior, ventral and lateral aspects of male
ed); k, ventral aspect of female 8th sternum; /,
gments; m, female 9th sternum and bursa.

24 U.S. NATIONAL MUSEUM BULLETIN 277

5. Scenopinus cavifrons (Kréber)
Figure 6 (after Kréber)

Omphrale cavifrons Kréber, 1937, p. 231.

The type of this species has been destroyed.

Length: Female body 4.1 mm., wing 3.7 mm.

Type-locality: Joinville, Algonquin, Canada?

Holotype: (SMNH).

A long series has been seen in the collection of the Illinois Natural
History survey with the label “Algonquin” (Northern Illinois) which
may actually have been the source of Kréber’s specimen. These
specimens all agreed with fenestralis (L.). However, since the type
cannot be checked, I have let the name stand.

G

Ficure 6.—Scenopinus cavifrons (Krober), female:
Details of frons and antenna (after Krober).

6. Scenopinus efflatouni, new species

FIGURE 7

Frema.Le.—A moderately large robust fly typical of the group. Head
red-brown; eyes red-brown; postocular ridge moderately broad; frons
subshining with small pits for fine hairs, swollen on either side of a
median groove extending from the upper half to the median ocellus,
flat below; the ocellar tubercle slightly swollen, ocelli orange-brown.
First two antennal segments orange-brown; third segment three times
as long as the first two combined, red-brown. Mouthparts well de-
veloped.

Thorax red-brown, with short recumbent white hairs, the posterior
margin of the humeral callus orange-brown; scutellum red-brown with
short erect white hair. Wings subhyaline, with brown cast, veins

SCENOPINIDAE OF THE WORLD 25

Ficure 7.—Scenopinus efflatouni, new species, female: a, wing; b, c, lateral and frontal
aspects of head; d, detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect of
8th and 9th segments; g, 9th sternum and bursa.

26 U.S. NATIONAL MUSEUM BULLETIN 277

brown; halter stem yellow, knob light tan. Legs with femora and
tibiae brown, tarsi straw yellow, darkening distally.

Abdomen red-brown, with short recumbent hairs, middle segments
with membranous posterior margins; see figure for details of eighth
sternum and bursa.

Matzr.—Unknown.

Length: Female body 4.0—-4.5 mm., wing 3.0-3.4 mm.

Type-locality: Abu Rowash, Egypt; 5 November 1924 (Efflatoun).

Holotype: Female (SEE).

Paratype: Female, Cairo; 4 July 1918 (Efflatoun) (SEE).

7. Scenopinus fenestralis (Linnaeus)

FIGURE 8

Musca fenestralis Linnaeus, 1758, p. 597.

Musca saltitans Scopoli, 1763, p. 350.

Musca spoliata Scopoli, 1763, p. 350.

Musca senillis Fabricius, 1794, p. 331.

Atrichia fasciata Schrank, 1803, p. 103.

Scenopinus pallipes Say, 1823, p. 100.

Scenopinus sulcicollis Meigen, 1824, p. 114.

Scenopinus domesticus Meigen, 1824, p. 116.

Scenopinus rufitarsis Meigen, 1838, p. 165.

Scenopinus scutellatus Macquart, 18438, pp. 6-8, pl. 1.
Scenopinus furcinervis Zetterstedt, 1844, p. 897.

Musca “‘tarda’’ Linnaeus.—Haliday, 1851, p. 138.1
Scenopinus graminicola Zetterstedt, 1859, p. 6045.
Scenopinus fuscinervis Schiner, 1862, p. 159.

Scenopinus scutellatus var. nigroscutellatus Frey, 1945, p. 36.
Scenopinus perkinst Paramonov (not Hardy), 1955, p. 651.

The type of this species has long been lost but it is a well established
and easily identified species which is distributed over all of the zoo-
geographic regions of the world. It is characterized by the rugose
frons in both sexes; antennae long, slender and dark brown; and
legs yellow-brown. The halteres vary from white to dark brown. The
male terminalia has a notch at the tip of the dorsal lobe that shows
considerable variation; see figure 8m. Specimens with these different

1 In 1851, Haliday published a paper in the Stettiner entomologische Zeitung,
vol. 12, p. 138, dealing with the Linnaean collection in London. The reference
to “tarda” is to a manuscript name in the Linnaeus collection which Haliday says
is a synonym of fenestralis.

ey

Ficure 8.—Scenopinus fenestralis (Linnaeus), male, female: a, wing; b, d, lateral and frontal
aspects of male head; c, ¢, lateral and frontal aspects of female head; f, enlarged detail of
antenna; g-1, ventral, lateral and posterior aspects of male terminalia; 7, ventral aspect
of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female 9th
sternum and bursa; m, various emarginations on ventral margin of male 9th tergum.

27

SCENOPINIDAE OF THE WORLD

313-168 O—€69——3

28 U.S. NATIONAL MUSEUM BULLETIN 277

patterns are widely distributed in North America and Europe and
are believed to be conspecific.

Length: Male body 4.0-4.5 mm., wing 3 mm.; female body
4.5-5.5 mm., wing 3.5-3.7 mm.

Type-locality: Europe.

Distribution: Worldwide.

8. Scenopinus glabrifrons Meigen
FIGURE 9
Scenopinus glabrifrons Meigen, 1824, p. 114.
Scenopinus halterata Meigen, 1824, p. 113.
Scenopinus orbita Meigen, 1824, p. 114.
Scenopinus laevifrons Loew, 1857, p. 88.
Omphrale glabrifrons var. surcoufi Séguy, 1920, p. 318.
Scenopinus civiculus G. H. Hardy, 1933, p. 418.
Paromphrale glabrifrons (Meigen).—Krober, 1937, p. 214.

These are large flies similar to Scenopinus fenestralis but are easily
separated on the basis of the shining frontal areas of both the male
and female. Antennae elongate orange-brown at the base and darker
at the tip. Halteres are white (similar appearing flies with dark
halteres are Scenopinus vitripennis Meigen). Male ninth tergum with
smooth margins; female eighth sternum and bursa with minor charac-
ters. Paramonov (1955, p. 649) places glabrifrons in synonomy with
cwiculus.

Length: Male body 4.0-4.5 mm., wing 3 mm.; female body
4.5-5.5 mm., wing 3.5-3.7 mm.

Type-locality: EKurope—type-specimen lost.

Distribution: Worldwide.

9. Scenopinus griseus (Kréber)

FicurE 10
Omphrale grisea Krober, 1913. p. 191.

Maur.—This small fly differs noticeably from most members of
this group, being considerably smaller and having very short swollen
antennae. It does, however, have the four-lobed condition of the
male genitalia placing it in this group. The abdomen has most of
the dorsal surface of segments 2-7 white. The wings have a brownish
cast and the halter knobs are white.

Frmate.— Unknown.

Length: Male body 2.5-2.7 mm., wing 1.8 mm.

Type-locality: Gy6én, Hungary.

Holotype: (MNH) destroyed.

Topotype: Male, Gyén (Kertész) (ZMUH).

Other specimen examined: Gyén (NMYV).

SCENOPINIDAE OF THE WORLD 29

Ficure 9.—Scenopinus glabrifrons Meigen, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; J, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

30. U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 10.—Scenopinus griseus (Krober), male: a, wing; b, c, lateral and frontal aspects of
head; d, enlarged detail of head; e-g, lateral, ventral and posterior aspects of terminalia.

SCENOPINIDAE OF THE WORLD 31

10. Scenopinus japonicus (Séguy)
Figure 11
Omphrale japonicus Séguy, 1920, p. 318.
FrmaLe.—This is a medium sized black fly; the head of the type
was crushed and the antennae which Séguy describes as being blackish
brown are missing. Frons rugose with pit above the antennae and

shining band extending up to ocellar triangle. Wing and terminalia as
figured.

Ficure 11.—Scenopinus japonicus (Séguy), female: a, wing; b, ventral aspect of 8th sternum;
c, lateral aspect of 8th and 9th segments; d, 9th sternum and bursa.

Ma.re.— Unknown.

Length: Female body 3.8 mm., wing 2.5 mm.

Type-locality: environs of Tokyo, Mt. Nikko, Japan; 1901 (J.
Harmand).

Type: (MNHN).

1l. Scenopinus lesinensis Strobl
Figure 12
Scenopinus lesinensis Strobl, 1904, pp. 533-34.

The type and allotype of this species are presumed to be lost.
The specimens illustrated are presumed to be the species but are
larger than the specimens of Strobl.

32 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 12.—Scenopinus lesinensis Strobl, female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect
of 8th and 9th segments; g, 9th sternum and bursa.

FrmaLe.—Head black. Eyes black-brown with a reddish cast,
postocular ridge moderate; frons rugose with median groove; sub-
shining, with a few sparse hairs; mouthparts dark brown, well de-
veloped; lower facial area subshining. Antennae elongate, first seg-
ment black, short; second red-brown; third elongate, granular, black.
Ocellar triangle not prominent, ocelli orange.

SCENOPINIDAE OF THE WORLD 33

Thorax black with short sparse hairs directed posteriorly, humeral
and supra-alar calli orange; posterior margin of scutellum orange.
Wings brown, veins dark brown; halter stem brown, knob red-brown
above, cream below. Legs orange-brown, except tips of tarsi brown,
hind legs darker.

Abdomen dark red-brown, hairy; see figure for details of 8th sternum
and bursa.

Length: Female 5.5 mm., wing 4.2 mm.

Type-locality: Verbosca and Brusji in Lesina, Yugoslavia, July.

Type: Lost.

Specimen figured, one of 4 females reared from larvae, Kitzbuhel,
Austria; end of May 1952 (NMV).

12. Scenopinus microgaster (Séguy)

FIGURE 13

Omphrale microgaster Séguy, 1948, p. 155.

This species from South China is characterized by the very short
abdomen of both male and female.

Ma.e.—Frons rugose with a median depression above a narrow
transverse ridge above the antennae. Antennae short and pear-shaped.
Wings smoky brown, veins brown; three narrow white bands across the
abdomen.

Frma.Le.—Frons with a slightly elevated central ridge with incurv-
ing lower side pieces along upper two thirds, a slight dimple in the
median groove above a central depression. For details of the eighth
sternum and bursa see figure.

Length: Male body 3.25 mm., wing 2.75 mm.; female body 4 mm.

Type-locality: Kouy (Kouri) Tcheou, China; 1921 (Cavalerie).

Type and allotype: (MNHN).

13. Scenopinus nidorcaupulus, new species

FicureE 14

This is a large species much like S. fenestralis except for the antennae
which are short and broad.

FrmMALE.—Head black, eyes black-brown; postocular ridge moder-
ately wide and polished; front roughened except shining at eye mar-
gins, shallow median groove above the antennae with a slight swelling
on either side; mouthparts dark brown, hairy; palpi less than half as
long as the oral opening, flattened, paddlelike. Ocellar tubercle black,
not prominent, median ocellus depressed into the front. Antennal seg-
ments black-brown, second shorter than first, segment 3 twice as long
as 1 and 2 combined, wide at base with a rounded tip.

34 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 13.—Scenopinus microgaster (Séguy), male, female: a, wing; 5, ¢, lateral and frontal
aspects of male head; c, d, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, lateral, ventral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

Thorax dorsum black, transversely striated; humeral and supra-
alar calli red-brown. Wings a dusky brown, veins brown; stem of
halter dark brown, knob shining, red-brown. Legs with femora dark
red-brown, tibiae brown and tarsi yellow-brown.

SCENOPINIDAE OF THE WORLD 35

Ficure 14.—Scenopinus nidorcaupulus, new species, female: a, wing; ), c, lateral and frontal
aspects of head; d, ventral aspect of 8th sternum; ¢, lateral aspect of 8th and 9th segments;
f, 9th sternum and bursa.

Abdomen dark red-brown to red-brown; see figure for details of
eighth sternum and bursa.

Mate.— Unknown.

Length: Female body 6+mm., wing 4.2 mm.

36 U.S. NATIONAL MUSEUM BULLETIN 277

Type-locality: Steamboat Springs, Colorado; 1 August 1943, 6800
ft. (Bryant).
Holotype: Female (CAS) 8926.

14. Scenopinus niger (De Geer)

FicureE 15

Nemotelus niger De Geer, 1776, p. 188.
Musca fenestralis Fabricius, 1781, p. 444.
Musca rugosa Fabricius, 1794, p. 330.
Nemotelus tarsata Panzer, 1809, p. 20.
Scenopinus atra Fallén, 1817, p 2.
Scenopinus nigripes Meigen, 1824, p. 117.
Scenopinus fasciata Walker, 1851, p. 85.

This well-known European species is distinctive in that the male is
one of the very few where the frons broadly separates the eyes and the
eyes are not divided into upper coarse facets and lower fine facets.
This species appears to be limited to continental Europe and is not
found on the Pacific Islands where other dark-winged species occur.

Length: Male body 4.45 mm., wing 4 mm.; female body 5.2 mm.,
wing 4.7 mm.

Type-locality: Europe.

Type: Lost?

Specimens present in most European collections.

15. Scenopinus opaculus Loew

Figure 16
Scenopinus opaculus Loew, 1873, p. 145.

This species from the trans-Caspian region looks like a small edition
of S. fenestralis with dark legs. The male genitalia, however, are quite
distinctly different. The abdomen of the female was lost so the details
of the eighth sternum and bursa could not be illustrated.

Length: Male body 1.25 mm., wing 1.25 mm.; female body 1.5-2
mm., wing 1.25-1.75 mm.

Type-locality: Samarkand.

Type: (ZMH).

16. Scenopinus physadius (Séguy)
FIGURE 17
Omphrale physadius. Séguy, 1930, p. 111.

This species from Morocco resembles S. glabrifrons in many ways.
The frons of the male is shining but does not protrude as much as
glabrifrons; the female frons is slightly granular and is covered with

SCENOPINIDAE OF THE WORLD 37

Ficure 15.—Scenopinus niger (De Geer), male, female: a wing; }, d, lateral and frontal
aspects of male head; c, ¢, lateral and frontal aspects of female head; f-h, ventral, lateral
and posterior aspects of male terminalia; 7, ventral aspect of female 8th sternum; J,
lateral aspect of female 8th and 9th segments; &, female 9th sternum and bursa.

38 U.S. NATIONAL MUSEUM BULLETIN 277

RaQ

SCENOPINIDAE OF THE WORLD 39

short hairs. The wings are a smoky white with yellow veins and the
halteres of both sexes are white. The terminalia of both sexes are
distinctive; see figure for details.

Length: Male body 4.5 mm., wing 3 mm.; female body 5.5 mm.,
wing 3.5 mm.

Type-locality: Bou Denib, Morocco (de Lepiney).

Type: (MNHN).

17. Scenopinus saini, new species

FiGurRE 18

Matr.—This moderately large dark-colored fly is closely related to
S. fenestralis. Head black-brown, eye facets brown above and black-
brown below; frons subshining, slightly smaller above the antennae;
ocellar tubercle prominent with a few anteriorly directed hairs, ocelli
clear; back of head concave; postocular ridge lacking. Antennae
elongate with subequal black-brown first and red-brown second seg-
ments, third segment red-brown, slender, about four times as long as
1 and 2 combined.

Thorax dorsum black, shining, with a thin pile of whitish hairs;
humeral and supra-alar calli red-brown; pleural areas red-brown, with
thin pile of hair. Legs red-brown throughout. Wings hyaline, veins
straw colored; halter stem red-brown, knob white.

Abdomen red-brown with broad white membranous borders on
segments 4-6; terminalia red-brown; see figure for details.

FremaLe.— Unknown.

Length: Male body 3.7 mm., wing 3 mm.

Type-locality: W. El Lega, S. Sinai, Egypt; June-August 1942
(Hafez).

Holotype: Male (CUC).

18. Scenopinus schroederi (Kréber)
FicurE 19

Omphrale schroederi Krober, 1913, p. 201.

This large Central American species from Costa Rica is placed in
the Fenestralis group on the basis of the long third antennal segment,
vein R4 branching from before the middle of cell R5 and the open

bursal cavity.
Mate.— Unknown.

Ficure 16.—Scenopinus opaculus Loew, male, female: a, wing; 6, c, lateral and frontal
aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged detail
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia.

40 U.S. NATIONAL MUSEUM BULLETIN 277

SCENOPINIDAE OF THE WORLD 41

Length: Female body 5 mm., wing 4 mm.
Type-locality: Costa Rica.
Type: Stettiner Museum. (Now housed in [PAS] Warsaw.)

19. Scenopinus sinensis (Kréber)

Figure 20 (after Kréber)
Omphrale sinensis Kroéber, 1928a, p. 1.

This species was known only from the type female which was
destroyed during World War II.

Mare.— Unknown.

Length: Female body 4.1 mm.

Type-locality: Canton, China (Helms).

Type: (SMNH) destroyed.

20. Scenopinus undulafrons, new species

FIGURE 21

This is another of the S. fenestralis type flies found along the west
coast of continental North America and is distinguished by its shorter
antennae and by its genital structures.

Frma.e.—Head black; eyes red-brown; the frons broad, slightly
swollen, subshining, with a definite longitudinal depression just above
the antennae, a slight median depression on the upper half; ocellar
tubercle slightly elevated, the median ocellus depressed; ocelli red-
brown to clear; postocular flange broad and elevated. Mouthparts well
developed, pale tan with numerous red hairs; palpi short, red-brown,
lighter at tip. Antenna with first segment dark brown, second red-
brown, third red-brown, less than twice as long as broad, see figure.

Thorax dorsum black, rugose, with short hairs; humeral callus
orange-brown, supra-alar callus red-brown. Wings smoky brown, veins
brown. Halter stem red-brown, knob red-brown. Legs with front coxa
through the tibia red-brown; the first tarsal segment straw yellow,
darkening distally.

Abdomen and terminalia red-brown; see figure for details of eighth
sternum and bursa.

Maxe.— Unknown.

Length: Female body 5.2 mm., wing 3.5 mm.

Type-locality: Terrace, B.C., Canada (Mrs. M. E. Hippisley).

Ficure 17.—Scenopinus physadius (S¢guy), male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, enlarged detail
of antenna; g-i, ventral, lateral and posterior aspects of male terminalia; 7, ventral aspect
of female 8th sternum; &, lateral aspect of female 8th and 9th sternum; /, female 9th
sternum and bursa.

42 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 18.—Scenopinus saini, new species, male: a, wing; b, lateral aspect of head; c, frontal
aspect of head; d, enlarged detail of antenna; ¢, lateral aspect of terminalia; f, ventral
aspect of terminalia; g, posterior aspect of terminalia.

SCENOPINIDAE OF THE WORLD 43

Ficure 19.—Scenopinus schroederi (Krober), female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect
of 8th and 9th segments; g, 9th sternum and bursa.

Ficure 20.—Scenopinus sinensis (Krober), female: Wing,
and lateral and frontal aspects of head (after Krober).

Holotype: Female (USNM) 67454.
Paratype: @ Oliver, B.C., Canada; 12 June 1958 (H. and A.
Howden) (CNC).

313-168 O—69——-4

44 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 21.—Scenopinus undulafrons, new species, female: a, wing; 5, c, lateral and frontai
aspects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

21. Scenopinus varipes Loew
Scenopinus varipes Loew, 1873, p. 148.

This species appears to be lost to science; at least all my efforts
to find its whereabouts have been fruitless.

Length: Male body 1 mm., wing 1 mm.

Type-locality: Balfrusch (Babul), Iran; Christoph Coll.

Type: Location unknown.

22. Scenopinus vitripennis Meigen
FIGURE 22
Scenopinus vitripennis Meigen, 1824, p. 115.—Trojan, 1956, p. 1-10.

This species has been synonymized with S. glabrifrons by a number
of the earlier workers. On the basis of the wing venation, however,
Trojan (1956) showed that vitripennis is a valid species. It is most
easily identified as a brown haltered glabrifrons-like species. The
formation of the eighth sternum and bursa further confirm Trojan’s
evidence that this is a different species; see figure.

The only known male of this species is presumed lost.

Specimens identifiable to this species have been found in a number
of European collections and come from Eastern Europe.

Length: Female body 4.5-5.5 mm., wing 3.5-3.7 mm.

Type-locality: Austria (Megerle).

Type-specimen: lost.

SCENOPINIDAE OF THE WORLD 45

Ficure 22.—Scenopinus vitripennis Meigen, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, left, ventral aspect of 8th sternum; right, dorsal aspect
of 8th and 9th segments; ¢, lateral aspect of 8th and 9th segments; f, ventral aspect of
9th sternum and bursa.

ALBICINCTUS group
Key to Species of the ALBICINCTUS group

PALAEARCTIC SPECIES (MALES)

1. Wing white orsmoky brown. . .. 2... ee ee ee ee ee
Wing hyaline, not definitely white or brown. . . . . 2... 2... + 78

46

bo

U.S. NATIONAL MUSEUM BULLETIN 277
Wing white, halter knob white. . . ...... ... .. .§S.niveus Becker
Wing smoky brown, halter knob brown .. . .S. bulbapennis, new species
Halter; knob: cream... vstonree nese errata ate tor oe cae oe eee ee ueae
Halter knob white ... . Reset opti satel weds te ee eee ae oh IO
Wing with whitish cast, pntenha Gros .... . . .S. albicinetus (Rossi)

Wing with brownish microtrichia, antenna black-brown
S. canarius, new species

Antenna red-brown or black-brown. ....... Tapecas 0
Antenna black ... ; ‘ é Ss. himpidipennis Loew
Antenna red-brown. Apdoinen att three white bance the first broad, termi-
nalia with two long aedeagal spines .:. . . . . . .S. lucidus Becker
Antenna black-brown, three narrow white sees on abdomen: terminalia with
aedeagus short. .......... ... . .S. phaidimos, new species

PALAEARCTIC SPECIES (FEMALES)

Wing brown or with brown, patches. .): 4... 40h 2 oa See eae ses ae
Wing hyaline ... si: 0<he -vedltey. Ss Csp@stwa ape ctetc: Magers ae
Wing brown, body without eeaielike has o} Nea te .3
Wing hyaline with brown patches in center of pelist body, clothed arith flat

seale-like hairs ... .... . . .S. squamosus Villeneuve
Halter knob white, matonnn brow. i ar ale @ ose | So See Datrize (Séguy)
Halter knob brown, antenna black-brown . . .S. bulbapennis, new species
Halter‘knob white: 0 <0 Sees) a) sileica ihe e ta eee we ees eet aan.
Halter knob cream or tan... . ea Ld ae Ne

Halter knob cream, antenna black, body, machen noseeable have
S. albicinctus (Rossi)

Halter knob tan, antenna brown, body with short hair
S. curtipilosus, new species
Antenna red-brown or black, M1+2 ending after the tip of the wing. . . 7
Antenna black-brown, M1-+2 ending before the tip of the wing
S. phaidimos, new species
Antenna with basal segments orange or red-brown, third segment brown or

red-brown .. . Aha eee ake as SoA COU Sy BB CCKEer

Antenna with third ceca black. ..... . . . §S. limpidipennis Loew
ETHIOPIAN SPECIES (MALES)

Wing hyaline, halter knob white, antenna black-brown or brown. . . . 2

Wing milky white, halter knob white, antenna orange-brown
S. monodi Séguy

Halter knob white, abdomen with one white band. .......... 3
Halter with base of knob red-brown, remainder white, no white stripes on the
abdomen . . . ... . . §. anthrax, new species
Antenna black- oe abdomen is one white band, aedeagal parameres
slender it scn2 ... . . S. balteatus Lamb
Antenna brown dackeneas distally, spdaiien with one narrow white band,
aedeagal parameres antler-like ....... . . . S. inquilinus Séguy

ETHIOPIAN SPECIES (FEMALES)

Wing hyaline. 3

Wing milky or brown.

Halter knob white or cream . :
Halter knob tan, brown or black browal.

oNWwW Mm wh

10.

Lf

12.

13.

14.

15.

SCENOPINIDAE OF THE WORLD AT

eal Gers kno Dr Chesiyey neers ewer 2. Sotre! Sore eet tee en we a Se
Halter knob white . .. . ee ee eee
Halter cream above, white below, ‘antenna Drown. . S. keiseri, new species
Halter knob cream, antenna black ...... . . . S. bouvieri (Séguy)
Halter knob brown or black-brown. . . . eee 40
Halter knob tan dorsally, whiter ventrally, eateniee orange at Base darkening
attip. ... ... . . §. brunneus (Kréber)
Halter knob onan anand iowa dertenaa distally
S. inquilinus (Séguy)
Halter knob black-brown above, white below, antenna black-brown
S. angustifrons (Krober)
Antenna red-brown, frons flat, wide, shining. . S. gromieri, new species
Antenna black-brown, frons narrow, rugose ... . . S. balteatus Lamb
Wing brown. . . ee ee ee iy fc) ee
Wing milky, halter white Loe ew ee eee eh. d6S. monodi Séguy
Halter knob red-brown to black. ......... 2 ee LO)
Halter knob yellow-brown .. . 4 oe Be ‘Ss. Ravipes (Kroéber)
Halter red-brown or base of knob meds prom remainder white . . a
Halter knob brown or black... . = & bes; eee po iA
Halter knob red-brown . . . ee ee eee
Halter knob with base red- ae a meaenen wihice.
S. anthrax, new species
Halter knob red-brown shining antenna orange, darkening at tip, palpus
broad and paddle-like. . . . . .. . . .S. palmulapalpus, new species
Halter knob red-brown, antenna red-brown, palpusnormal. .... . .13
Frons widening dorsally, cell R5 narrowing at tip . . S. evansi, new species
Frons widening ventrally, cell R5 nearly uniformly wide throughout
S. katbergi, new species
Halter knob brown, antenna dark, mouthparts normal ....... .15
Halter knob black-brown, antenna yellow, mouthparts atrophied
S aethiopicus (Lindner)
Halter knob brown above cream below, antenna orange basally darkening
distally, cell R5 parallel sided. . . . . S. madagascariensis Enderlein
Halter knob brown, antenna black, cell R5 broad, vein M1+2 ending in tip
Of Wine. 2) 2 2 eee. e hs ee. .... . . .S. oldenbergi (Kréber)
ORIENTAL AND PACIFIC SPECIES
Males . .2
Females . . no nee
Wing brown, halter eieb oe byow n or ohlaces aaiedun peek or cream. . .3
Wing hyaline, halter knob white, antenna yellow-brown, terminalia with two
long aedeagal spines... . .. . . . .S.lucidus Becker
Halter knob with base red- poe venmndes white, no white bands on the
abdomen, antenna black .... . . . . . .S. papuanus (Kroéber)

Halter knob black, antenna cream, Biaomien with white bands
S. opacus de Meijere
Wing brown, halter knob brown or with red-brown. . . . ee
Wing hyaline, halter knob white. . . . ae ae a, ineidus Began
Halter knob with base red-brown, fomnden Ww ite, antenna black
S. papuanus (Kroéber)
Halter knob brown, antennacream ..... .. . . .S. opacus de Meijere

48 U.S. NATIONAL MUSEUM BULLETIN 277

AUSTRALIAN SPECIES (MALES)

ee CS alah aaa eg id Sei oe can canes a a
Wing ‘hyaline ."=. ... SRE. ROR Gna tM: ce 0 2M,

2. Halter knob red-brown, eerie prom fons aia
S. bolgarti, new species
Halter knob red-brown with a cream band separating the upper and lower

halves, frons triangular, pebbly . . . . . . .S. fragosifrons, new species

3. Halter knob red-brown or brown, antenna brown or black-brown . . 4
Halter knob black, antenna brown. .... . .. S. perkinsi G. H. Hardy

4. Halter knob brown, antenna brown, normal sized fly. . . . S. biroi Kertész

Halter knob red-brown, antenna black-brown, very small fly
S. minutus, new species

AUSTRALIAN SPECIES (FEMALES)

1. Wing smoky brown or white kaa
Wing hyaline . go

2. Wing smoky brown . eo,
Wing white. : 4

3. Halter knob white, antenna medbrownlt veins s R5 onic M1 42 separated at ene
wing tip = .°.'. ‘ . . . §S. spurrelli, new species
Halter hnob red-brownl) smterinie black- eae wing with veins R5 and M1+2
meeting at the wingtip. . . . . . . . . §S. darwini, new species

4. Halter knob red-brown with a cream band dividing the upper and lower
halves, antenna red-brown, frons pebbly, swollen
S. fragosifrons, new species
Halter knob as above, antenna brown, frons rugose but not swollen
S. bolgarti, new species

5. Halter knob brown orred-brown ....... 6
Halter knob black .... . Dy, HT, 8) peehinci G. ‘H. Hardy
6. Halter knob brown, antenna Genngesbraea or brown. . . Olea cla

Halter knob red-brown, antenna black-brown, a very small ay
S. minutus, new species
7. Antenna orange-brown darkening to brown at tip. . S. norrisi new species
Antenna ‘brown iho) s26r Res Sk. RS. SE a. ee Sr ipopsKertesz

23. Scenopinus aethiopicus (Lindner)

FIGuRE 23
Omphrale aethiopicus Lindner, 1955, p. 20.

FrmMALE.—This appears to be a good species distinct from other
described African species. There are several characters that Lindner
overlooked; both specimens referred to in his description are females;
the type specimen had atrophied mouthparts as illustrated, while the
paratype had well developed mouthparts. Front with an elongate de-
pression above the antennae reaching over one-third the distance to
the ocellar triangle. Halter rounded at apex. The eighth sternum
appears to be characteristic but could not be illustrated in detail
along with the bursa as I was not granted permission to dissect.

Ma.e.— Unknown.

Length: Female body 3-4 mm.

SCENOPINIDAE OF THE WORLD 49

Type-locality: Kisangara, Kenya; 31 December 1951 or 1 January
1952 (D. O. Afrika Exp.).
Type: (SMNS).

Ficure 23.—Scenopinus aethiopicus (Lindner), female: a, wing; , c, lateral and frontal
aspect of head; d, sketch of atrophied mouthparts; ¢, enlarged detail of antenna; f, lateral
aspect of undissected 8th and 9th segments; g, ventral aspect of undissected 8th sternum.

24. Scenopinus albicinctus (Rossi)

FIGURE 24

Musca albicincta Rossi, 1794, vol. 2, p. 69.

Scenopinus zelleri Loew, 1845, p. 314; 1857, p. 89; 1873, p. 151.
Scenopinus albicinctus (Rossi).—Bezzi, 1898, p. 36.

Omphrale albicincta (Rossi).—Krober, 1913, p. 184.

This species is easily identified by its general light coloration.
Front white, dorsum of thorax and lower pleural areas dark with a
wide band of yellow extending from humeral area to posterior margin
of scutellum. Basal portion of the ninth tergum of male dark with
distal end white. Abdomen with a white band below scutellum;
second segment broad, remaining segments narrower and banded
white behind.

Length: Male body 3-4.5 mm., wing 3 mm.; female body 3.5-
4mm., wing 3 mm.

Type-locality: Italy.

Type: presumably lost.

Specimens examined: 207<7, 19, Djerba, Tunis (R. C. Shannon)
in (USNM); 17, Macedonia, Greece, September 1933 (R. C. Shan-
non) in (USNM).

50 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 24.—Scenopinus albicinctus (Rossi), male, female: a, wing; b, c, male head, lateral
and frontal aspects; d, ¢, female head, lateral and frontal aspects; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 51

25. Scenopinus angustifrons (Kréber)
FicureE 25 (after Kréber)

Omphrale angustifrons Kroéber, 1913, p. 197.

The type of this species has been lost and no specimen agreeing
with it has been seen.

Length: Female body 5.5 mm.

Type-locality: Willowmore, Cape Prov., [Republic of] S. Africa.

Type: (MNH) destroyed.

DN Figure 25.—Scenopinus angustifrons
(Krober), female: Wing and lateral

and frontal aspects of head (after
Krober).

26. Scenopinus anthrax, new species

FIGURE 26

Maue.—Head red-brown; eyes red-brown above, black-brown
below; frons very narrow, divided into a lanceolate upper portion and
a triangular lower portion by bands of plumose hairs that extend
dorsally from the oral margins to almost meet on the lower third of the
frontal area. Mouthparts normal, palpi flattened, broad, extending the
full length of the mouthparts; back of head concave, little or no
sclerotization showing even below; ocellar triangle prominent, ocelli
clear. Antennae red-brown, third segment three times as long as 1
and 2 combined.

Thorax dorsum dark red-brown, humeral and supra-alar calli
orange-brown; scutellum with orange-brown posterior rim, red-brown
above. Wings hyaline, anterior veins red-brown and straw colored
behind; halter stem red-brown, knob white with red-brown base.
Legs with red-brown femora and tibiae; first tarsal segment red-brown,
2-4 orange, 5 red-brown.

Abdomen red-brown with no white bands. For details of terminalia
see figure.

02 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 26.—Scenopinus anthrax, new species, male, female: a, wing; b, c, male head, lateral
and frontal aspects; d, e, female head, lateral and frontal aspects; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male ter-
minalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

FEMALE.—Head red-brown, eyes red-brown. Frons only as wide as
ocellar tubercle, punctured with a shallow median pit; ocellar triangle
not prominent ocelli orange; postocular ridge narrow; mouthparts as

SCENOPINIDAE OF THE WORLD 53

in male. Antennae as in male except last segment black-brown, polli-
nose.

Thorax dorsum black, transversely striated, with numerous short
hairs giving a silvery appearance; humeral and supra-alar calli red-
brown. Halter stem brown, knob red-brown with a brown pollinose
bloom. Legs short and stout, femora and tibiae red-brown, tarsi
orange-brown except last segment dark. Wing brown, fumose; veins
brown. Sternum of eighth segment orange-brown, sculptured as in
figure, bursa as figured.

Length: Male body 3.2-3.8 mm., wing 2.4-2.6 mm.; female body
3.2-4.5 mm., wing 2.3-3.2 mm.

Type-locality: Bekily, Reg. Sud de L’lle, Madagascar; March
1933 (A. Seyrig).

Holotype: Male (MNHN).

Allotype: Female Vila Pery, Mozambique; 24 November 1928
(P. Lesne) (MNHN).

Paratypes: o 2 copulatory pair, Bekily, Reg. Sud de L’lle, Mada-
gascar; April 1938 (A. Seyrig). 1 o& of same data but October 1936.
1 & Plaines de Fiherera; 1905 (F. Geay) in (MNHN). 2 9° Mada-
gascar Nord, Montagne d’Ambre 1000 M., det. Diego-Suarez; 23
November-—4 December 1957 (B. Stuckenberg) in (NMP).

27. Scenopinus balteatus Lamb

FIGurE 27
Scenopinus balteatus Lamb, 1922, p. 363.

The male is characterized by a smooth, sword-shaped central ridge
on the frons, a single white band on the abdomen and by the shape
of the terminalia. The female may be recognized by the configuration
of the eighth sternum and bursa. The halter of the male is tan, that
of the female white.

Length: Male body 3 mm., wing 2 mm.; female body 3.7 mm.,
wing 2.7 mm.

Type-locality: Mahe, Cascade Estate; 800-1500 ft., Seychelles
(Percy Sladen Exp.).

Types: (BMNH).

28. Scenopinus biroi Kertész
FIGURE 28
Scenopinus biroi Kertész, 1899, p. 173.
This species is common to New Guinea and Australia; may be
recognized by its dark body and bright yellow legs.
Ma.e.—Head black, frons small, triangular, eyes with large upper
and small lower facets. Thorax black; wings brown; legs yellow.

04 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 27.—Scenopinus balteatus Lamb, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; hf, 1, ventral and lateral aspects of male terminalia, cut
ejaculatory syringe suggested by dashed outline; 7, posterior aspect of male sternum; k,
ventral aspect of female 8th and 9th segments; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

Genital segments as figured.
Length: Female body 2.5 mm., wing 2.2 mm.; male smaller.
Type-locality: Friedrich-Wilhelmshafen and Erima, New Guinea.

SCENOPINIDAE OF THE WORLD 55

Type: (MNH) destroyed.

Paratypes: 2 99, (USNM) 24178, Kréber exchange. Male rede-
scribed and figured, National Park, Sidney, emerged 20 November
1922, from pupa found in Autechinus (Phascogale) flavipes (a small
mouselike animal) (CSIRO); 3 99, Cairns, Queensland, on window
(J. F. Illingworth) in (USNM).

Ficure 28.—Scenopinus biroi Kertész, male, female: a, wing; 5, lateral aspect of male
head; c, d, lateral and frontal aspects of female head; ¢, f, enlarged details of male and
female antennae; g-i, ventral, lateral and posterior aspects of male terminalia; j, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa.

29. Scenopinus bolgarti, new species

FIGURE 29

This species from West Australia is placed in the Albicinctus Group
on the basis of the male and female terminalia. It differs from the usual
members of the group, however, in the failure of M1+2 to attain the
tip of the wing or unite with R5, and in the shape of the male antenna.
Though presumed to be opposite sexes of the same species, lack of
wings on the female prevents positive identification.

Ma.e.—Head black, eyes with brown upper facets and black-brown
lower field; separated by a narrow band of frons which is rugose,
sunken below the eye margins, with a shallow pit above the antennae.
Ocellar tubercle elongate, dusted olivaceous; ocelli red-orange, back
of the head dusted with silvery gray pollen; mouthparts brown, well
developed, palpi short. Antenna with segment 1, dark brown, very

56 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 29.—Scenopinus bolgarti, new species, male, female: a, wing; b, d, lateral and frontal
aspects of male head; c, ¢, lateral and frontal aspects of female head; g, h, enlarged details
of male and female antennae; f, 1, 7, ventral, lateral and posterior aspects of male ter-
minalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

short; 2, brown, wider than long and twice as long as 1; 3, brown,
powdery, pear-shaped with two distinct points.

Thorax black, covered with dark brown scales; dorsum dark brown
with light brown lateral stripes, whitish above humeral callus and
posterior portion of mesonotum; humeral and supra-alar calli straw
colored; scutellum black; pleural areas black-brown, covered with
scales. Wings smoky white, veins very light brown anteriorly and
nearly clear posteriorly, vein M1+2 fading before joining R5 at the
tip of wing. Halter stem light brown, knob red-brown, scaly. Legs
with fore femur red-brown, scaly, flattened, fore tibia and tarsus
yellow-brown; middle and hind legs as above except femur normally
shaped.

Abdomen with second segment broad, red-brown, scaly, posterior
margin white; remaining segments with a thin dark band anteriorly,
white posteriorly, giving the appearance of a nearly white dorsum.
Sternites also with white posterior margins. Ninth tergum with
white margins and a red-brown scaled median area, and some long
hairs; for details see figure.

FEMALE.—Head black, eyes black-brown; frons divided by a median
groove, slightly elevated above the lateral margins which are sunken

SCENOPINIDAE OF THE WORLD 57

below the eyes; a triangular pollinose patch, covered with fine hairs,
having its base above the antennae and extending to the median
ocellus. Ocellar tubercle black, not prominent, ocelli orange-red.
Antenna with first segment short, black; second short black-brown;
third black-brown, over twice as long as first and second combined,
pear-shaped and widest at its base.

Thorax black, shining, with fine hairs; humeral cal!us orange-brown.
Halter with stem red-brown, knob dark red-brown with a creamy
brown band dividing the upper and lower halves. Wings brownish
at base but broken off, veins brown. Legs with femora and tibiae dark
red-brown, tarsi tan darkening at tips.

Abdomen red-brown; see figure for details of eighth sternum and
bursa.

Length: Male body 2 mm., wing 1.5 mm.; female body 2.5 mm.,
wing missing

Type-locality: Bolgart, W. Australia; 14 December 1961 (L. B.
Britton and A. Douglas).

Holotype: Male and female allotype (BMNH)

30. Scenopinus bouvieri (Séguy)

FIcurRE 30

Omphrale bouvieri Séguy, 1920, p. 317.

This species is quite similar to S. balteatus Lamb but differs in
details of the frons, the shorter length of the median crossvein, and in
details of the eighth sternum. The bursa have similar configurations
but differ in minute details.

The type was taken in Paris from the nest of a pigeon. A specimen
examined from Loango, French Congo, agrees in all details with the
type and, in line with the agreement in the shape of the bursal cavity
with the Albicinctus group, would suggest that this species is pri-
marily African in affinities.

Martre—Unknown.

Length: Female body 3.5 mm., wing 2.5 mm.

Type-locality: Paris (? introduced from Africa).

Type: (MNHN).

Material examined: Female Loango, Foret du Mayrombe, French
Congo; 1911 (Vodon) (MNHN).

31. Scenopinus brunneus (Kroéber)
FiGurE 31

Omphrale brunnea Kréber, 1913, p. 192.

This species, described originally from East Africa, appears to be
distributed across equatorial Africa from the Indian Ocean to the

58 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 30.—Scenopinus bouvieri (Seguy), female: a, wing; missing portions filled in by
dashed lines; b, c, lateral and frontal aspects of head; d, enlarged detail of antenna; ¢,
ventral aspect of 8th sternum; f, lateral aspect of 8th and 9th segments; g, 9th sternum and
bursa.

Atlantic. The type specimen is teneral and a good deal smaller than the
average. The ground color of other specimens is black or black-brown.
The antennae are colored as follows: first segment brown; second
brown-orange; third brown-orange at base darkening to black-brown
at tip. Halter tan dorsally and white ventrally. Front and middle legs
black-brown with first four tarsal segments yellow-brown and last
segment red-brown; hind leg with tarsi entirely red-brown.

Mate—Unknown.

Length: Female body 3-4.5 mm., wing 2.2-3 mm.

Type-locality: Waboniland, East Africa.

Type: (NMYV).

Material examined: 19, Njala, Sierra Leone, 1926 (E. Hargreaves)
(ZMUH); 19, Lihete Sllomele, Congo Belge, July 1936 (J. Ghesquiere)
(IRSNB); 499, Njala, Sierra Leone, 1926, March 1936, April 1936
(E. Hargreaves); Moramba, Sierra Leone, 27 February 1925 (K.
Hargreaves) ; 19, laboratory, Accra, Gold Coast, March 1920 (J. W. S.
Macfie); 19, Bembay, Senegal, February 1940 (J. Risbec); 19, Cico,
Nyasaland, April 1916 (R. C. Wood) ; 19, Chipinda Pools, S. Rhodesia
(lower Lundi River), 23 October 1960 (R. Goocher); all in (BMNH).

SCENOPINIDAE OF THE WORLD 59

Ficure 31.—Scenopinus brunneus (Kréber), female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect of
8th and 9th segments; g, 9th sternum and bursa.

313-168 O—69

5

60 U.S. NATIONAL MUSEUM BULLETIN 277

32. Scenopinus bulbapennis, new species
FIGURE 32

Matue.—Head black; eyes red-brown above, black-brown below;
frons narrow, triangular, extending two-thirds the distance from the
antennae to the ocellar triangle, rugose, with a median groove extend-
ing from the antennal base to an elliptical depression at the top of
the frons; ocellar tubercle red-brown, ocelli orange-red; back of head
dusted gray pollinose; oral opening bordered by silvery pile extending
one-third the distance up the frons and cutting off a triangular area
above the antennae. Mouthparts brown, medium size; palpi half the
length of the oral opening. Antennae black-brown, first segment short,
second wider than long, third oval, slightly broader at base, pollinose.

Thorax with dorsum black, rugose with a pollinose patch above the
humeral callus which is red-brown with a yellow posterior tip; supra-
alar callus yellow-brown; pleura black-brown. Wings smoky, veins
brown; halter stem brown, knob brown. Legs with all segments dark
red-brown.

Abdomen black-brown, second segment pollinose, third and fourth
with narrow white bands on posterior margin. Terminalia as in figures.

Frema.e.—Head black; eyes black-brown, the postocular ridge
narrow; frons rugose, with a groove extending between lateral swellings
on the lower frons through the flattened upper frons to the median
ocellus; silvery pile laterad of oral opening extending into the frontal
area above the base of the antennae. Ocellar triangle black, rugose,
with orange-red ocelli; back of the head convex. Antennae as in male.

Thoracic features as in male. Abdomen dark red-brown. See figure
for details of eighth sternum and bursa.

Length: Male body 2.7 mm., wing 2.1 mm.; female body 3 mm.,
wing 2.1 mm.

Type-locality: Sierra Nevada Highway, Spain 2200 m.; 27 July
1960 (J. R. Vockeroth).

Holotype: Male (CNC) 8631.

Allotype: Female and paratypes 2c’ same data as type (CNC).

33. Scenopinus canarius, new species

Figure 33
Scenopinus brevicornis Frey (not Loew), 1937, p. 53.

Both specimens of this species had been identified as S. brevicornis
Loew, but differed in the wing venation and shape of the ninth tergum.
Ma.e.—Head dark brown; eyes red-brown to black-brown above,
darker below; frons triangular with a brown pollinose covering, a
shallow groove located in its center; back of the head concave above,

SCENOPINIDAE OF THE WORLD 61

Figure 32.—Scenopinus bulbapennis, new species, male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, lateral, and posterior aspects of male
terminalia; k, ventral aspect of female 8th sternum; /, dorsal aspect of female 8th and 9th
segments; m, lateral aspect of female 8th and 9th segments; 7, female 9th sternum and
bursa.

convex below, dusted with gray pollen and bearing scattered golden
hairs. Oral cavity bordered with brown pile, mouthparts of medium
length, dark brown; palpi brown; ocellar triangle dark red-brown, sub-

62 U.S. NATIONAL MUSEUM BULLETIN 277

TS GC
hop)

Ficure 33.—Scenopinus canarius, new species, male: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of

terminalia.

SCENOPINIDAE OF THE WORLD 63

shining, ocelli red. Antennae dark brown, first segment small; second
twice as long and wide as the first; third pollinose and twice as long
as 1 and 2 combined.

Thorax dorsum black-brown, dusted with brown pollen, covered
with moderately long golden hairs of medium density; humeral callus
similar except posterior angle yellow, with several anteriorly directed
hairs; supra-alar callus yellow-brown; pleura red-brown dusted with
bronze pollen. Wings hyaline with brown microtrichia, veins brown;
halter stem brown, knob cream. Legs red-brown throughout, covered
with golden hairs.

Abdomen red-brown, segments 3—5 with wide white bands two-
thirds width of segment. Terminalia differing from brevicornis in size
and shape of ninth segment claspers, and shape of posterior margin
of ninth tergum—see figure.

Fremate.— Unknown.

Length: Male body 2.8-3.2 mm., wing 2.1-2.3 mm.

Type-locality: Gr. Canaria, Los Palmas (R. Frey).

Holotype: Male (ZMUBH).

Paratype: Male Cabrea Diaz, Canary Island (NMV).

34. Scenopinus curtipilosus, new species

FIGURE 34

FrMALEe.—Head black-brown; eyes black-brown; frons_ broad,
shagreened, with a central groove, sculptured as in the figure; ocellar
triangle not prominent, ocelli red; postocular ridge narrow. Mouth-
parts well developed, pubescence surrounding oral cavity extending
only halfway to eye, remainder shining. Antenna with first segment
red-brown, short; second segment orange-brown, twice as large as
first; third segment red-brown, three times as long as 1 and 2 com-
bined, narrow tapering.

Thorax with dorsum black, rugose with a thin covering of short
hairs; humeral and supra-alar calli orange; pleural areas black-brown.
Wings hyaline, veins straw yellow; halter stem brown, knob tan.
Legs with femora and tibiae red-brown with knee orange, tarsi
orange-yellow.

Abdomen red-brown, middle segments with thin white membranous
posterior margins. Eighth segment with red-brown sternum; bursa as
figured.

Mare.— Unknown.

Length: Female body 3.5 mm., wing 2.3 mm.

Type-locality: El Elisah (El Shectch), Egypt; 21-27 April 1939
(H.C.E.).

Holotype: Female (CUC).

64 U.S. NATIONAL MUSEUM BULLETIN 2.77

{ F

f. g.

Ficure 34.—Scenopinus curtipilosus, new species, female: a, wing; b, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

35. Scenopinus darwini, new species
Figure 35
Frmaue.—Head black-brown; eyes black-brown, extending to the

back of the head so there is no postocular ridge; frons granular,
narrower than the ocellar triangle, with a median depression one-

SCENOPINIDAE OF THE WORLD 65

Ficure 35.—Scenopinus darwini, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, ventral aspect of 8th sternum; ¢, lateral aspect of 8th and
9th segments; f, 9th sternum and bursa.

66 U.S. NATIONAL MUSEUM BULLETIN 277

quarter of the distance from antenna to ocellus; ocellar tubercle
black-brown, slightly raised; ocelli red-brown. Oral cavity bordered
by gray pollinose hairs; mouthparts black-brown, filling oral cavity.
Antenna with first segment brown, short; second segment yellow-
brown, short; third segment black-brown, elongate, three times as
long as 1 and 2 combined.

Thorax with dorsum black, covered with short hairs that give it a
granular appearance; humeral and supra-alar calli red-brown; scutel-
lum red-brown, hairy like the dorsum, not distinctly set off from
scutum by a deep groove; pleural areas red-brown. Wings brownish,
hyaline, veins red-brown, halter stem red-brown, knob red-brown,
large. Legs with femora and tibiae red-brown, tarsi orange.

Abdomen red-brown, subshining; see figure for configuration of
eighth sternum and bursa.

Matzr.— Unknown.

Length: Female body 3.5+ mm., wing 2.7 mm.

Type-locality: Darwin, N. T., Australia, 9 March 1916 (G. F.
Hill).

Holotype: Female (USNM) 67455.

36. Scenopinus evansi, new species

FIGURE 36

This specimen was identified as S. balteatus Lamb by van Emden,
but the resemblance is superficial and differences in the head, wings,
and particularly eighth sternum and bursa remove it from that species.

FrmaLy.—Head black; eyes black-brown, extending to back of
head; frons slightly ridged from laterad of the antenna bases to the
median ocellus, with a small triangular shining spot on the lower
third, as wide as ocellar tubercle at its narrowest point. Ocellar
tubercle distinct, ocelli orange-red. Mouth-parts two-thirds length
of oral opening; palpi round and slim, a little shorter than the rostrum.
Antenna red-brown, first segment short; second segment twice as
large as the first; third segment oval, 2.5 times as long as broad.

Thorax with dorsum black; humeral and supra-alar calli orange;
pleural areas red-brown. Wings brown-fumose, veins brown; halter
stem brown, knob red-brown, shining. Legs with coxae through
tibiae red-brown, tarsi orange-brown.

Abdomen red-brown; for details of eighth sternum and bursa see
figure.

Mate.— Unknown.

Length: Female body 3.5 mm., wing 2.9 mm.

Type-locality: N. Rhodesia (Sylvester Evans).

Holotype: Female (BMNH).

SCENOPINIDAE OF THE WORLD 67

Figure 36.—Scenopinus evansi, new species, female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect of
8th and 9th segments; g, 9th sternum and bursa.

68 U.S. NATIONAL MUSEUM BULLETIN 277

Paratypes: 42 9, Ruo, Nyasaland, 14 April 1916, (R. C. Wood);
2 2 @, Ruo, Nyasaland, 29 April 1916, (R. C. Wood); all in (BMNH).

37. Scenopinus flavipes (Kréber)

Figure 37
Omphrale flavipes Krober, 1939, p. 398.

This species is known only from the female type which is figured.
The third segment of the antenna was lost and Kréber’s detail is
included; see figure for details.

Mauzr.— Unknown.

Length: Female body 3 mm., wing 2.3 mm.

Type-locality: Rabai, Kenya; August 1930, (van Someren).

Type: (BMNH).

/
i cl
wt b.

Ficure 37.—Scenopinus flavipes (Kréber), female: a, wing; b, c, lateral and frontal aspects of
head; d, detail of antenna (after Krober); ¢, ventral aspect of 8th sternum; f, lateral aspect
of 8th and 9th segments; g, 9th sternum and bursa.

38. Scenopinus fragosifrons, new species
FiGgurRE 38
Matz.—Head black-brown; frons large, triangular, rugose, with a

faint median groove above antennae to the top of the anteriorly
sloping part of the frons; eyes red-brown, with large upper and

SCENOPINIDAE OF THE WORLD 69

smaller lower facets, protruding above antennae and extending to
the back side of the head (see figure). Ocellar tubercle slightly elevated
and distinct, ocelli red-brown. The antennae, located on the lower
receding portion of the face, with short red-brown first segment;
second orange-brown and twice as large as the first; third red-brown,
pear-shaped and twice as long as the first two combined. Mouthparts
well developed.

Ficure 38.—Scenopinus fragosifrons, new species, male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

Thorax black-brown, shining, with a few short hairs, humeral and
supra-alar calli orange-brown. Wings milky with anterior veins tan,
remaining veins very light and faint. Halter stem brown, knob red-
brown with a creamy band separating the upper and lower halves.
Legs, fore and middle, with coxa to tibia black-brown, tarsus red-
brown covered with short stiff spines; hind leg with coxa to tibia
black-brown; first tarsal segment orange-brown, three times as long as
second segment; remaining segments red-brown; all tarsal segments
densely covered with short stiff spines.

Abdomen with second segment red-brown with short pilose hairs
above and long white hairs laterally; segments 3-5 with broad white
bands medially, lateral portions completely sclerotized; remaining
segments red-brown. For terminal segments see figure.

70 U.S. NATIONAL MUSEUM BULLETIN 277

Fremate.—Head black-brown, flattened and receding below the
antennae; eyes red-brown; mouthparts well developed extending only
half the length of the oral cavity. Frons broad, swollen above the
antennae; nubbly portion narrowing dorsally to the width of the ocel-
lar tubercle, dorso-lateral portions with transverse ridges like a crackle
finish; postocular ridge moderate y wide; ocellar tubercle black-brown,
prominent; ocelli red-brown. Antennae as in male except all segments
red-brown.

Thorax as in male; tarsi uniformly light red-brown.

Abdomen red-brown; for details of eighth sternum and bursa see
figure.

Length: Male body 3.3 mm., wing 2.2-2.7 mm.; female body 4.5
mm., wing 2.8 mm.

Type-locality: Yanchep, W. Australia, 32 mi. north of Perth;
20-31 December 1935 (R. E. Turner).

Holotype: Male; allotype female; and paratype 16%’, 19, (same
data but 24 November and 2 December), (BMNH).

39. Scenopinus gromieri, new species

FIGURE 39

FrmMaLe.—Head red-brown; eyes brown, with a well defined
postocular rim; frons broad, nearly one-third the width of the head,
shining, reminiscent of S. glabrifrons but smaller in size yet larger
than S. phaidimos; ocellar tubercle prominent; ocelli orange. Anten-
nae red-brown, first segment short and small; second twice as large
as first; third segment four times as long as one and two combined, half
as broad as long. Mouthparts well developed, palpi short.

Thorax black-brown, granular; humeral and supra-alar calli yellow-
brown; pleural area red-brown. Wing hyaline, dusted with brown;
veins brown anteriorly, strawyellow posteriorly; halter stem brown,
knob white. Front leg with coxa, trochanter and femur straw colored,
tibia orange-brown, tarsus red-brown; middle leg with coxa, trochanter
and femur straw colored, remaining segments red-brown; hind leg
with all segments red-brown.

Abdomen red-brown; see figure for details of eighth sternum and
bursa.

Matrz.—Unknown.

Length: Female body 3.3-4.2 mm., wing 3.5-3.7 mm.

Type-locality: Cameroun N. O., Plateau de Dchany 1500 m.,
Saison Seche; 1 November 1942, (Dr. Gromier).

Holotype: Female (MNHN).

Paratypes: 3 2 9, Chilanga, N.W. Rhodesia; 2 2 9, 4 October
1913, in grain store in house 1 2, 20 November 1913, on window in
house; all (R. C. Wood) (BMNBH).

SCENOPINIDAE OF THE WORLD 71

Ficure 39.—Scenopinus gromieri, new species, female: a, wing; b, c, lateral and frontal as-
pects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

a2 U.S. NATIONAL MUSEUM BULLETIN 277

40. Scenopinus inquilinus Séguy

FiaurE 40
Scenopinus inquilinus Séguy, 1931, p. 1138.

This medium sized fly is easily distinguished in the male by the
single white band on the abdomen, the antlerlike shape of the male
aedeagal parameres, and shape of the ninth tergum. The female can
probably be positively identified only by the bursal cavity. Their
larvae have been found in Polyporus, in June and July, with adults in
July.

Length: Male body 3.3 mm., wing 2.3 mm.; female body 3.75 mm.,
wing 2.5 mm.

Type-locality: Nova Choupanga prés.Chemba, Mozambique; 24
August 1929 (P. Lesni), a copulating pair.

Type: (MNHN).

Material examined: 167,39 9, same data as type except 23 July;
1 @, Vila Pery, 24 November (MNHN); 1 oc, Luaba, Mozambique,
June and July 1957, (Usher and Stuckenberg); 1 @, Trelawney,
Southern Rhodesia, 14 December 1952, (N. J. Myers) (BMNH).

41. Scenopinus katbergi, new species

FicureE 41

FrmaLe.—Moderately large fly. Head black-brown, eyes brown; a
narrow postocular ridge; frons granular with a median shining band
widening on the upper part. White pilose bands bordering the oral
cavity extending to lower edge of frons above the antennal bases.
Ocellar tubercle prominent; ocelli clear; back of head covered with
gray pollen. Antennae with first segment black-brown, short; second
reddish brown, a little larger than first; third red-brown, elongate oval,
half again as long as wide.

Thorax with dorsum black, pitted, covered with short hairs;
humeral and supra-alar calli orange-brown; pleural areas red-brown,
pitted and covered with short hairs. Wings brown with brown veins,
appearing to have veins R2 as well as R3; R4 branches from basal
third of cell; halter stem red-brown, knob red-brown. Legs with femora
and tibiae red-brown; fore tarsus orange-brown, middle and hind
tarsi straw colored.

Abdomen red-brown; eighth sternum destroyed, bursal cavity as
illustrated.

Matrre.— Unknown.

Length: Female body 4.3 mm., wing 3.4 mm.

Type-locality: Katberg, east Cape Province, 4000 ft.; December
1932 (R. EB. Turner).

Holotype: Female (BMNH)

SCENOPINIDAE OF THE WORLD 73

Ficure 40.—Scenopinus inquilinus Seguy, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, remains of female 9th sternum and bursa.

74 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 41.—Scenopinus katbergi, new species, female: a, wing; b, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; e, bursa, 9th sternum lost.

42. Scenopinus keiseri, new species

FIGURE 42

FrEMALE.—Head black; eyes black-brown, extending to back of
the head so only the narrowest postocular ridge is formed. Frons
rugose, depressed above antennae and then swelling, a pit in center,
one-third the distance from the antennae to the median ocellus, median
portion raised above the eye margins. Ocellar triangle not prominent;
ocelli orange. Mouthparts brown, well developed, filling the oral
opening; palpi as long as the mouthparts, last segment broader than
the basal segments (see figure). First antennal segment brown, short;
second orange-brown, twice as large as first; third segment brown,
elongate oval, twice as long as broad.

SCENOPINIDAE OF THE WORLD 1

Ficure 42.—Scenopinus keiseri, new species, female: a, wing; 0, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral aspect
of 8th and 9th segments; g, 9th sternum and bursa.

Thorax with dorsum black; humeral callus orange along posterior
rim, supra-alar callus brown; scutellum black above with a brown rim;
pleural areas black-brown. Wings faint brown fumose; veins brown,
vein R4 branching beyond the middle, cell R5 tilted toward the front
of the wing at the tip. Halter with stem brown; knob cream above
and white below, to all white. Legs black-brown with orange-brown
tarsi, fore tibia a little lighter.

Abdomen dark red-brown; 8th segment red-brown, sternum and
bursa as figured.

Ma.re.— Unknown.

Length: Female body 3.2-4.4 mm., wing 2.3-2.8 mm.

Type-locality: Maroantsetra, Madagascar; 18-19 November 1957,
(F. Keiser).

Holotype: Female (NMB).

Paratypes: 6 99, same data as type (NMB); 1 9, same data as
type (USNM); 2 99, Perinet, Madagascar, December 1955, (B.
Stuckenberg) in (NMP).

313-168 O—69 6

76 U.S. NATIONAL MUSEUM BULLETIN 277

43. Scenopinus limpidipennis Loew
Scenopinus limpidipennis Loew, 1874, p. 416.

The type and allotype of this small species from Persia have not
been located so no additional information concerning the species can
be given.

Length: Male body 1% mm., wing 14 mm.; female body 2 mm.

Type-locality: Schahrud, Iran (Persia).

Type deposited: ? (not in Berlin).

44. Scenopinus lucidus Becker

Figure 43
Scenopinus lucidus Becker, 1902, p. 39.

This species has been returned to the genus Scenopinus from
Incidomphrale Kréber because of its complete agreement with the
members of the Albicinctus group. Its distinguishing feature, the
long aedeagal parameres of the male, serves as a good species char-
acter. The female agrees in all ways with Scenopinus and with the
characters of the Albicinctus group. One specimen was identified by
Kréber as S. glabrifrons, which it only superficially resembles; this
gives some idea of how difficult is the identification of females.

The details of the eighth sternum and particularly the bursa serve
to characterize the female. Just posterior to the membranous portion
of the bursa, on the midline, is a shell-like structure, that extends
down from the roof of the bursa, which is characteristic of this species.
(Hardy, 1960, p. 326-328 gives a good description of this species.)

Length: Male body 2.3-3 mm., wing 2-2.5 mm.; female body
3.5-4 mm., wing 2.5-3.3 mm.

Type-locality: Mariout, Egypt.

Type: (ZMH).

Material examined from: Hawaii (USNM) (PHT); Meadi, Egypt,
24 April 1917, (Efflatoun) (SEE); Christmas Island, November 1957,
(F. L. Hill), Elizabethville, Belgian Congo, (BMNH); Cape Verde
Islands (ZMUH); also from Algeria, Uganda, and northern Nigeria.

45. Scenopinus madagascariensis Enderlein

FIGURE 44
Scenopinus madagascariensis Enderlein, 1934, p. 429.

FrmaLe.—Head, cream below, black on upper frons and back of
head; eyes red-brown, a very narrow postocular rim fringed with
hairs; frons pollinose, granular, covered with short hairs, lower half
orange, extending dorsally between ventral extensions of black-brown

SCENOPINIDAE OF THE WORLD Ze

Ficure 43.—Scenopinus lucidus Becker, male, female: a, wing; b, lateral aspect of male
head; c, d, lateral and frontal aspects of female head; e, f, enlarged details of male and
female antennae; g-i, ventral, lateral and posterior aspects of male terminalia; j, ventral
left and dorsal right aspects of female 8th segment; &, lateral aspect of female 8th and
9th segments; /, female 9th sternum and bursa.

of upper frons; ocellar tubercle black-brown, granular, not set off,
ocelli orange; mouthparts brown, half the length of the oral cavity;
palpi shorter, cream-orange. Antennae with first segment orange,

78 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 44.—Scenopinus madagascariensis Enderlein, female: a, wing; b, c, lateral and frontal
aspects of head; d, enlarged detail of antennae; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, detail of posterior margin of 8th sternum; h, 9th sternum
and bursa; 1, lateral aspect of bursal area.

short; second orange, larger than first; third segment pear-shaped,
split at tip, orange basally darkening to black-brown distally.

Thorax dorsum with a black-brown ground heavily dusted with
yellow-brown pollen; humeral and supra-alar calli cream; scutellum
orange, granular; pleura red-brown, granular. Wings brownish hya-
line, veins brown; halter stem cream, knob brown above, cream below.
Legs with fore leg orange; middle leg orange except tarsus red-brown;
hind leg orange except coxa and tarsus red-brown.

Abdomen red-brown shining, base of segments black-brown, widest
at middle rounding posteriorly, flattened; see figure for details of
terminalia.

Mauzr.— Unknown.

Length: Female body 2.6 mm., wing 2.3 mm.

Type-locality: Madagascar, Diego-Suarez (S. V. Fredericks).

Type: (DEI).

46. Scenopinus minutus, new species

FiGure 45

Matxr.—Head black-brown; eyes black-brown above, black below;
frons short, subshining with a small dimple below the center; ocellar

SCENOPINIDAE OF THE WORLD 79

Ficure 45.—Scenopinus minutus, new species, male, female: a, wing; 6, lateral aspect of
male head; c, lateral aspect of female head; d, enlarged detail of male head and antenna;
e, enlarged detail of female antenna; f, g, enlarged frontal aspects of male and female
head; h-j, ventral, lateral and posterior aspects of male terminalia; k, ventral aspect of
female 8th sternum; /, lateral aspect of female 8th and 9th segments; m, female 9th
sternum and bursa.

tubercle elevated, black-brown, shining; ocelli red-brown; mouthparts
well developed, filling oral cavity. Antennae black-brown, third segment
broader at base, over twice as long as first two combined; see figure.

Thorax black-brown, shining; humeral and supra-alar calli red-
brown. Wing hyaline, whitish, veins tan to nearly clear; halter stem
black-brown, knob red-brown. Legs with femora black-brown,
remainder yellow-brown.

Abdomen black-brown, segments 3-5 with white bands, the first
indistinct. Terminalia black-brown; for details, see figure.

Frma.e.—Head red-brown; eyes black-brown; a well developed
postocular flange; frons shining with a median longitudinal depression.
Ocellar tubercle not prominent; ocelli red-brown. Mouthparts brown,
well developed, filling oral cavity. Antennae brown to black-brown.

Thorax as in male; the wings a little darker.

Abdomen all segments black-brown shining; for details of eighth
sternum and bursa, see figure.

Length: Male body 1.7 mm., wing 1.4 mm.; female body 1.7 mm.,
wing 1.4 mm.

80 U.S. NATIONAL MUSEUM BULLETIN . 277

Type-locality: Male, Bronte, New South Wales, Australia; 27
December 1954, (D. K. McAlpine).

Holotype: Male (AMS).

Allotype: Female, Como, New South Wales, Australia; December
1923, (H. Petersen) sweeping flowers, (USNM).

Paratypes: 1,1 2 (AMS);107,3 2 92, (USNM); (o'o’ same
data as holotype, 2 9 same data as allotype).

47. Scenopinus monodi (Séguy)

FIGURE 46

Omphrale monodi Séguy, 1933, pp. 122-127.

This species is known from the type male, allotype female and four
paratype females.

Mate.—With front depressed; antennal segments 1 and 2 orange,
the third segment missing but presumably orange-brown as in the
female. Eyes of male red-brown above and black-brown below. Body
covered with short white hairs; wings white, veins light yellowish;
knob of the halter white. Abdomen with three solid white bands.

Terminalia as figured.
c), (HAY
(ohGq GO: XK
SS

ay

—~

a.

Ficure 46.—Scenopinus monodi (Séguy), male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, enlarged detail
of female antenna; g-1, ventral, lateral and posterior aspects of male terminalia; 7, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa.

FrEmMALE.—With frons broad covered with white hairs pointing
ventro-laterally. Thorax as in male. Abdomen red-brown, eighth

SCENOPINIDAE OF THE WORLD red |

sternum short, broad; see figure for details of eighth sternum and
bursa.

Length: Male body 2.5 mm., wing 2 mm.; female body 3 mm.,
wing 2 mm.

Type-locality: Entre Tabankort et in Tassit, Sahara Soudanise;
28 December 1927, (Angieras and Draper).

Types: (MNHN).

48. Scenopinus niveus Becker

FIGURE 47

Scenopinus niveus Becker, 1907, p. 61.

This species is known only from the type-locality. The type was not
found, but a long series of over a dozen specimens, all males, from
the type-locality was seen in the Museum of Natural History, Paris.

Mate.—The male of this species is distinguished by the wing, where
veins R5 and M1 + 2 are parallel; by the five white bands on the
abdomen and the antennae.

FremaLe.— Unknown.

Length: Male body 2.5 mm., wing 2 mm.

Type-locality: Gafsa, Tunisia. .

Type deposited: Coll. Becker (not located).

Material examined: See above.

Ficure 47.—Scenopinus niveus Becker, male: a, wing; 6, c, lateral and frontal aspects of
head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of terminalia.

82 U.S. NATIONAL MUSEUM BULLETIN 277

49. Scenopinus norrisi, new species

Figure 48

FrmA.Le.—Head black; eyes black-brown; postocular ridge narrow;
front granular and no wider than the ocellar triangle, with a small
median pit located one-third the distance from the antennae to the
median ocellus. Oral opening bordered on each side by a band of gray
pubescence which begins on the lower frontal areas above and to each
side of the antennal sockets and extends ventrally between the eye
margin and the oral opening. First antennal segment very short,
black-brown; second segment orange-brown, two to three times size of
first; third segment elongate, more than twice as long as broad, orange-
brown basally darkening to brown distally, blunt on the end. Mouth-
parts well developed, nearly filling the oral cavity, dark brown; palpi
long, exceeding over one-half the length of the oral opening.

(FERS)

Frcure 48.—Scenopinus norrisi, new species, female: a, wing, b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, dorsal aspect
of 8th and 9th segments; g, lateral aspect of 8th and 9th segments; h, 9th sternum and
bursa.

Thorax black-brown, covered with a thin pile of short hairs. Wings
hyaline, with veins brown, space between R1 and R2 + 8 appears
filled, R4 branching beyond the middle of the cell; halter stem brown,
knob brown. Legs black-brown with tarsi orange-brown.

Abdomen uniformly dark brown, thinly covered with posteriorly-
directed hairs; see figure for details of the eighth sternum and bursal
plate.

Mate.— Unknown.

Length: Female body 4.5. mm., wing 3.5 mm.

Type-locality: Brisbane, Queensland, Australia; 14 January 1954
(K. R. Norris).

SCENOPINIDAE OF THE WORLD 83

Holotype: Female (CSIRO).
Paratypes: 1 @, Brisbane, 24 January 1960, (Hasler) (UQue);
1 2, Redlynch, Queensland, 26 September 1938, (BMNH).

50. Scenopinus oldenbergi (Krober)
Ficure 49
Omphrale oldenbergt Kréber, 1913, p. 190.

It is my opinion, based on the characters of the antenna and bursal
plates, that this species is not European in origin but is more likely
African and therefore must be placed in the Albicinctus group.

Ma.e.— Unknown.

Length: Female body 4 mm., wing 2.2 mm.

Type-locality: Berlin, 3 August?, Oldenberg.

Type: (DEI).

Figure 49.—Scenopinus oldenbergi (Krober), female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th and 9th segments; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

51. Scenopinus opacus deMeijere
Figure 50

Scenopinus opacus deMeijere, 1924, p. 14.

This species from Sumatra is one of the few from the Malaysian
Archipelago and is quite distinctive, particularly in the male termi-
nalia. These are moderately large black flies with orange antennae

84 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 50.—Scenopinus opacus deMeijere, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; J, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

and orange-brown to yellow tarsi; wings brown; see figure for details.
Length: Male body 3.5 mm., wing 2.7 mm.; female body 4.2 mm.,

wing 3.5 mm.

SCENOPINIDAE OF THE WORLD 85

Type-locality: Fort de Kock, Sumatra; December 1920 (E.
Jacobson).
Types and paratypes: (ZMUA).

52. Scenopinus palmulapalpus, new species

Figure 51

FrmMALe.—Robust; head black-brown; eyes black-brown, with
little or no postocular flange; frons much broader than the ocellar
triangle, slightly swollen above the antennae into two bulges separated
by a shallow groove leading to a shallow depression, appearing shining
but in reality slightly granular. Ocellar triangle not set off; ocelli
orange. Face silvery pilose laterad of antennae. Mouthparts well
developed; palpi over half as long as oral opening, broadly flattened
(paddle-like), covered with a velvety pile. First and second antennal
segments red-brown, third segment orange-brown at base darkening
at tip, about three times as long as broad.

Thorax dorsum black, rugose, transversely striated; posterior tip
of humeral callus and supra-alar callus yellow-brown; pleural areas
black-brown. Wings brown fumose, veins brown, vein M1+2 bending
forward to close cell R5 with a bottleneck; halter stem brown; knob
red-brown, shining. Fore leg with coxa to tarsus yellow-brown;
middle leg light brown; hind leg coxa to tibia red-brown, tarsus light
brown.

Abdomen red-brown, eighth segment hairy; see figure for details
of sternum and bursal plates.

Ma.re.— Unknown.

Length: Female body 3.4-3.8 mm., wing 2.3-2.7 mm.

Type-locality: Pietermaritzburg, Natal; 2 April 1918, (Harden-
berg).

Holotype: Female (NMP).

Paratypes: 2929, Pietermaritzburg, 17 November 1952, (B. Stuc-
kenberg) 16 February 1956, (B. Stuckenberg), (NMP); 1 @, Pieter-
maritzburg, 16 November 1954, (B. Stuckenberg) (USNM).

53. Scenopinus papuanus (Krober)
Figure 52
Omphrale papuana Krober, 1912, p. 25.

Scenopinus niger Grimshaw (not De Geer), 1901, p. 11.—D. E. Hardy, 1958, p. 11.
Scenopinus zeylanicus Senior-White, 1922-24, p. 205.

Specimens of this species from Hawaii were for a long time identified
as S. niger and it was not until 1958 that Hardy was able to clear up

86 U.S. NATIONAL MUSEUM BULLETIN 277

FicureE 51.—Scenopinus palmulapalpus, new species, female: a, wing; }, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, f, lateral and frontal details of palpus;
g, ventral left and dorsal right aspects of 8th and 9th segments; Ah, lateral aspect of 8th
and 9th segments; 7, 9th sternum and bursa.

the synonomy and describe the male. On examination of the type o
of S. zeylanicus in the British Museum, I found that this species is a
synonym of S. papuanus.

SCENOPINIDAE OF THE WORLD 87

Specimens have also been examined from Cocos (Keeling) Islands,
Christmas Island (BMNH), Guam (USNM), and Hawai (PHT)
establishing its distribution from Ceylon to Hawaii.

Length: of body 3.2-4 mm., wing 2.7-3.5 mm.

Type-locality: Kanshirei, Formosa; April (H. Sauter) (restricted
by Hardy, 1960, p. 331).

Type: (DED).

Ficure 52.—Scenopinus papuanus (Krober), male, female: a, wing; b, lateral aspect of male
head; c, d, lateral and frontal aspects of female head; e, f, enlarged details of male and
female antennae; g-1, ventral, lateral and posterior aspects of male terminalia; 7, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa.

54. Scenopinus patrizi (Séguy)
FicurE 53
Omphrale patrizi Séguy, 1932, p. 499.

This species is known only from the type. Frons broad and shining
black. Band of white present along the side of the thorax extending
from head to halteres which are white; see figure for details of the head,
wing, eighth sternum and bursa.

Mate: Unknown.

Length: Female body 2.5 mm., wing 2 mm.

Type-locality: Aujila [Augila], Libia; April 1931, (Patrizi).

Type: (MCSNG).

88 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 53.—Scenopinus patrizi (Séguy), female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antennae; ¢, ventral aspect of 8th sternum; f, lateral aspect
of 8th and 9th segments; g, 9th sternum and bursa.

55. Scenopinus perkinsi G. H. Hardy

FIGURE 54

Scenopinus perkinst G. H. Hardy, 1942, p. 202.—Paramonov, 1955, p. 651,
description of male.

This species is known only from Australia where it was originally
collected on the windows of the Brisbane Museum. The male was
taken as a pupa from a Spice Finch nest and another specimen has
been seen from a sparrow’s nest. The male is distinguished by a pair
of pubescent bulges on the insides of the tergal plates surrounding the
terminalia.

Length: Male body 3.5 mm., wing 2.5 mm.; female body 4-5 mm.,
wing 3 mm.

Type-locality: Brisbane, Queensland (F. A. Perkins).

Type: Female (UQue) destroyed.

Specimens examined: 1 <, Woolstoncraft, near Sydney, New
South Wales, emerged 12 June 1950 (K. A. Hindwood); 1 9, Brisbane,
Queensland (F. A. Perkins); (CSIRO).

56. Scenopinus phaidimos, new species
Figure 55

Ma.er.—Head black; eyes black, very distinctly divided into large
upper and smaller lower facets; frons small, reduced to a narrow

SCENOPINIDAE OF THE WORLD 89

Ficure 54.—Scenopinus perkinsi, G. H. Hardy, male, female: a, wing; b, c, lateral aspects
of male and female head; d, enlarged detail of female antenna; e¢, enlarged detail of inner
surface of male antenna; f, g, anterior aspects of male and female head; h-, ventral, dorsal
and posterior aspects of male terminalia (note swellings on inside dorsal surface of 9th
tergum); k, ventral aspect of female 8th and 9th segments; /, lateral aspect of female 8th
and 9th segments; m, female 9th sternum and bursa; n, female accessory organ.

Ficure 55.—Scenopinus phaidimos, new species, male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; /-j, ventral, lateral and posterior details of male
terminalia; &, ventral aspect of female sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

90 U.S. NATIONAL MUSEUM BULLETIN 277

triangle above the antennae, rugose with a median groove; ocellar
triangle elevated, ocelli orange-brown. First antennal segment short,
black; second larger, red-brown; third black-brown, pear-shaped, with
a distinct depression on the outer side at the tip.

Thorax black, shining; humeral callus with a small red-orange patch
on posterior edge. Wings hyaline, veins tan; halter stem tan, knob
white. Legs black with moderate bristles, tarsi orange-brown darkening
distally; middle and hind legs with the first tarsal segment subequal
to 2-5 combined.

Abdomen black, shining, segments 3-5 white along posterior mar-
gins. Terminalia black with a few long hairs; see figure for details.

Frma.Le.—Head black; eyes red-brown; postocular ridge broad
shining; frons swollen, shining, broader than either eye when viewed
from the front, with a broad median groove ending in a pit on lower
third of head. Ocellar tubercle distinct, set off, ocelli red-brown.

Thorax, legs, wing and halter as in male.

Abdomen, black, shining, flattened, eighth sternum longer than
tergum; see figure for details of eighth sternum and bursa.

Length: Male body 2.4 mm., wing 2.1 mm.; female body 3.2 mm.,
wing 2.5 mm.

Type-locality: 20 mi. N. of Sereflikochisar (Ankara area), Turkey,
24 June 1962, 900 m. (Guichard and Harvey).

Holotype: Male (BMNH).

Allotype: 2, and @2 paratype, Eskisahir, Kaymaz, Turkey, 800
m.; 27 July 1962, (Guichard and Harvey) (BMNH).

57. Scenopinus spurrelli, new species

FIGurE 56

Frema.Le.—Head black; eyes red-brown; postocular ridge narrow;
back of head pollinose; frons broad subshining, with sparse hairs
laterad of a median groove; ocellar tubercle shallow, ocelli whitish;
antennae red-brown, first segment short, second twice as large as first,
third soft pilose, tapering to tip from a broader base.

Thorax black, shining, with short hairs, humeral and supra-alar
calli red-brown on margins. Wing smoky brown, veins brown; halter
stem black-brown with a V-shaped bracket extending posteriorly to
hold knob which is white. Legs with coxae and trochanters dark red-
brown, femora orange-brown, tibiae and tarsi light orange-brown.

Abdomen black-brown; see figure for details of eighth sternum and
bursa.

Ma.e.—Unknown.

Length: Female body 4.6 mm., wing 3.3 mm.

SCENOPINIDAE OF THE WORLD 91

Figure 56.—Scenopinus spurrelli, new species, female: a, wing; }, c, lateral and frontal
aspects of head; d, ventral aspect of 8th sternum; , lateral aspect of 8th and 9th segments;
Ff, 9th sternum and bursa; g, lateral view of 9th segment and bursa, showing inner pocket.

313-168 O—69——_7

92 U.S. NATIONAL MUSEUM BULLETIN 277

Type-locality: Wilkur, Victoria, Australia; 2 December 1955,
(Spurrell).
Holotype: Female (VMM).

58. Scenopinus squamosus Villeneuve
FIGURE 57

Scenopinus squamosus Villeneuve, 1913, p. 111.
Omphralosoma squamosa (Villeneuve).—Krober, 1937, p. 222.

The presence of scales on the body is insufficient reason for erecting
a new genus for a single species, especially when the shape of the
antenna, wing venation, and bursal conformation agree with the other
members of the Albicinctus group. It is, therefore, being returned to
the genus Scenopinus.

Ficurr 57.—Scenopinus squamosus Villeneuve, female: a, wing; , c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

In addition to the scales, the presence of dusky patches on the center
of the cells of the wings helps to identify this species. The abdomen
has white posterior bands on the segments, the halteres are black.

For details of head and eighth segment and bursa, see figure.

Maue.— Unknown.

Length: Female body 2-3 mm., wing 2.2 mm.

SCENOPINIDAE OF THE WORLD 93

Type-locality: Ain-Sefra (Oran), Algeria; May and June [1897]

(M. Bleuse).

to

bo

Type: (IRSNB).
Material examined: 1 9, Ghardaia, Algeria; May 1897, (IRSNB).

BREVICORNIS group

Key to Species of the BREVICORNIS group

PALAEARCTIC SPECIES (MALES)

Wine smoky prowl OF WHILE. fu) soar ao eng aS ee le AEF eo 2
Wing hyaline. ...........2.... . . S. fraternus (Kroéber)
Wing milky white . a ee 3
Wing smoky brown . 3 5
Halter knob white; in black ‘call RS oom ae oe 4

Halter knob white; antenna black; cell R5 constricted at tip
S. brevicornis Loew

Antenna (spindle form) pointed. . ....... . . S. nitidulus Loew
Antenna oval . . SA ey ROO S. parallelus, new species
Halter knob white or with aa bar iy Be et sn eres 6
Halter knob red-brown. . . Oe anh S. hal¢epalis Brey

Halter knob white; antenna black, body clothed with medium long thin hairs
S. pilosus (Séguy)

Halter knob with a tan bar, antenna brown, body with sparse hairs
S. unifasciatus (Kréber)

PALAEARCTIC SPECIES (FEMALES)

WIED BING Yl OW. ace a Oh hee le oS 2S ee, ad a oR ee
Wing milky .... ... . . §S. brevicornis Loew
Halter knob white, erent Bincke or caw. oO rien iti mata” ad eat
Halter knob tan, antenna black-brown . . . . . . S. lincinus, new species
Antenna lacks yess ls eee es oe he ; 4
Antenna brown. . . a: S. Nalfascistus (Krober)
Halter knob long; tip of patentee pointed: vein R4 branching well beyond

middle of cell R5. . . . ... . . §. nitidulus Loew
Halter knob round; ee of aienne poceded: R4 branching from near middle

of ce] R5 .... . ... . . S. fraternus (Krober)

ETHIOPIAN SPECIES (MALES)

Winpsmoky whit@s i922 oe elton Ae ee eee . 2
Wing smoky brown. . . . ee ‘Ss. fryeri, new Papers

Halter knob white, antenna black, vein 1 M1- 42 ending before tip of wing. . 3
Halter knob red-brown, antenna ae -brown, vein M1+2 ending behind

tip of wing. . . ..... . . §, turneri, new species
Cell R5 constricted beyond! vein RA. ...... . S. minusculus (Séguy)
Cell R5 not constricted beyond vein R4. . . . . . . S. tarsalis (Kréber)

ETHIOPIAN SPECIES (FEMALES)

Wing hyaline . : ;
Wing smoky white or brown . cur
Halter knob yellow-brown, antenna beanie black, Aah s. ‘tarsalia (Kréber
Halter knob reddish yellow, antenna dark brown with yellow tip

S. nitidifrons (Kréber)

wm Wb

94 U.S. NATIONAL MUSEUM BULLETIN 277

See Wing whites ce 2 eo erreel eet ee 2 ie ee hy; ope rend Mee
Wan EOWAD «6 ray Sores sos ae San lomete are NBilnie pe oe elas Ne eatins (eu cic cen Senet teee

4. Halter knob white, antenna black, 8th sternum short
S. minusculus Séguy

Halter knob cream, antenna black-brown, 8th sternum long
S. longiventris (Kréber)

5. Halter knob tan, velvety, white below, antenna black-brown, velvety, cell R5

constricted at tip by R5 dipping down toward M1+2
S. reduncus, new species

Halter knob tan, antenna black-brown, cell R5 curving sharply anteriorly,
vein M1+2 ending well before tip of wing . . . .S. lincinus, new species

59. Scenopinus brevicornis Loew

FIGURES 58, 59

Scenopinus brevicornis Loew, 1873, p. 146.

This species is known only from the type-locality in Turkistan.
Loew described both sexes; however, I have seen only the type male.
Krober (1924) described and illustrated a female which is illustrated
here. On the basis of the wing venation, it is doubtful that this specimen
is actually the female of S. brevicornis as it does not agree very closely
with wings of the type. Two male specimens from the Canary Islands
which Frey identified as S. brevicornis are a new species, S. canarius.

Ficure 58.—Scenopinus brevicornis Loew, male: a, wing; 0, c, lateral and frontal aspects of
head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of terminalia.

SCENOPINIDAE OF THE WORLD 95

Figure 59.—Scenopinus brevicornis Loew, female: a, wing; b, c, lateral and frontal aspects
of head; d, ventral aspect of 8th sternum; ¢, lateral aspect of 8th and 9th segments; f,
9th sternum and bursa.

96 U.S. NATIONAL MUSEUM BULLETIN 277

Length: Male body 2.5 mm., wing 2 mm., female body 3 mm.,
wing 2.2 mm.

Type-locality: Warsaminor, Turkistan.

Type: (ZMH).

Specimen examined: 1 9 Mariout, Egypt; 1 September 1921
(Efflatoun) deposited in (SEE).

60. Scenopinus fraternus (Kréber)

Fiaures 60, 61

Omphrale fraterna Krober, 1923, p. 81.

Kréber remarks on the similarity of the male of this species with
that of S. nitidulus. Certainly in the details of the head, wing and
external genitalia they agree in most respects. The principal difference
lies in the internal genital structures where a marked difference in
the size of the ejaculatory pump is evident.

Ficure 60.—Scenopinus fraternus (Kréber), male: a, wing; b,c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of
terminalia.

There is some doubt, however, as to whether the female is of the
same species as the male. The wings do not agree at all well in the
two specimens; see figures.

SCENOPINIDAE OF THE WORLD 97

Length: Male body 2.5 mm., wing 2 mm.; female body 3.2-3.5
mm., wing 2—2.1 mm.

Type-locality: Male Saila, Egypt; 4 April 1923 (Fayoum).

Holotype: Male (SEE).

Allotype: Female, Ramleh, 25 June 1922.

Paratypes: 19, Nouzha, 5 July 1921; 1 9, Ezbet el Nakhl, 18 April
1922, (SEE).

Ficure 61.—Scenopinus fraternus (Krdber), female: a, wing; }, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e, dorsal left and ventral right aspects of 8th and
9th segments; f, lateral aspect of 8th and 9th segments; g, 9th sternum and bursa.

61. Scenopinus fryeri, new species

FIGURE 62

Scenopinus longiventris Lamb (not Kroéber), 1922, p. 362.

This specimen carries a determination label as S. longiventris Kréber
but details of the head and wing have led me to describe it as a new
species.

FremMaA.Le.—Head dark red-brown; eyes red-brown, with a prominent
postocular ridge; back of head concave; frons broad, finely rugose,
swollen basally and laterad of the ocellar tubercle, and with a median
depression. Mouthparts well developed; ocellar tubercle dark red-
brown, not sharply set off from the frons, ocelli orange-red. Antennae
with first two segments red-brown; first short, second twice as large;

98 U.S. NATIONAL MUSEUM BULLETIN 277

third segment black-brown, pear-shaped, as in figure, and covered
with golden hairs.

Thorax dark red-brown with a few sparse hairs; humeral callus
orange. Wings smoky brown; halter stem brown, knob tan. Legs dark
red-brown with lighter red-brown tarsi. Abdomen red-brown, sub-
shining; see figure for details of terminal segments and bursa.

C0

Ficure 62.—Scenopinus fryeri, new species, female: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e, dorsal aspect of 8th and 9th terga; f, lateral
aspect of 8th and 9th segments; g, ventral aspect of 9th sternum and bursa.

Matze.— Unknown.

Length: Female body 2.7 mm., wing 2 mm.

Type-locality: Aldabra Island, Seychelles; 1908-09 (J. C. Fryer).
Holotype: Female (BMNH).

62. Scenopinus halteralis Frey

FIGURE 63

Scenopinus halteralis Frey, 1937, p. 53.

This small fly from the Canary Islands is known only from the
male and is one of the smallest scenopinids seen. The specimen figured
is one of six paratypes.

Length: Male body 1.5 mm., wing 1.3 mm.

Type-locality: La Esperianza, Tenerife, Canary Islands; 16
August (P. Stora).

Type: (ZMUH).

SCENOPINIDAE OF THE WORLD 99

Figure 63.—Scenopinus halteralis Frey, male: a, wing; 6, c, lateral and frontal aspects of
head; d, enlarged details of head and antenna; e-g, ventral, lateral and posterior aspects
of terminalia.

63. Scenopinus lincinus, new species

FIGURE 64

This is one of several species which have female terminalia differing
from those of most of Scenopinus, in that the sternum is a little longer
than the tergum, rounded, and the ninth tergites have short stiff hairs
or spines. Based on the configuration of the bursal plates, all appear to
be distinct species. The present species is closely related to S. fraternus
and S. nitidulus as previously mentioned, as well as S. fryeri, S. re-
duncus and perhaps S. longiventris. The males of the last three are
unknown and the similarity of the males of the first two have already
been mentioned.

Frema.e.—Head black-brown; eyes dark red-brown; a broad, sunk-
en postocular ridge; frons broad, subshining, with a median dimple;
ocellar tubercle set off from frons; ocelli red-brown. First antennal
segment black-brown, short; second segment red-brown, twice as
large as first; third segment black-brown, pear-shaped, covered with
gray pollinose scales, with a ventro-lateral pit.

Thorax black-brown, subshining; humeral callus red-brown, supra-
alar callus tan. Wings smoky brown; veins brown, R5 and M1 + 2
both curving forward to end before the tip of the wing; halter stem
brown, knob tan. Legs black-brown; femora with sparse short hairs;
tibae and tarsi with short hairs.

Abdomen red-brown, subshining; for details of the terminalia see
figure.

Ma.r.— Unknown.

100 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 64.—Scenopinus lincinus, new species, female: a, wing; b, c, lateral and fronta
aspects of head; d, enlarged detail of antenna; e, dorsal aspect of 8th and 9th terga; f
lateral aspect of 8th and 9th segments; g, ventral aspect of 9th sternum and bursa.

Length: Female body 3 mm., wing 1.8 mm.

Type-locality: Ambobitantely, Madagascar; 8 June 1959, (F.
Keiser).

Holotype: Female (NMB).

64. Scenopinus longiventris (Kréber)
Figure 65 [after Kréber]

Omphrale longiventris Kréber, 1913, p. 206.

The type of this species has been lost and no material from the type
locality has been seen. One specimen from the Seychelles identified
as this species by Lamb (1922) has been described as a new species
S. fryeri on the basis of Kréber’s figures. See note under S. lincinus.

Length: Female body 3 mm.

Type-locality: Colombo, Ceylon.

Type: (MNH) destroyed.

65. Scenopinus minusculus (Séguy)
FicureE 66
Omphrale minuscula Séguy, 1934, p. 74.
Scenopinus longiventris Lamb (not Kroéber), 1922, p. 362.

Mauz.—This small fly is typical of members of the group. Frons
rugose with flat, slightly elevated shining ridge as in figure, last an-
tennal segment covered with short hairs.

Thorax shining; wings milky; halter white; legs dark except tarsi
light.

SCENOPINIDAE OF THE WORLD 101

O OIC

Ficure 65.—Scenopinus longiventris (Krober), female:
Wing and lateral and frontal aspects of head
(after Krober).

Abdomen with three white bands; terminalia as figured.

FrmaLe.—New. Head brown; eyes brown; frons broad shining with
a central depression; postocular ridge narrow; ocellar triangle indis-
tinct. Antenna as male.

Thorax brown, as male.

Abdomen brown; eighth sternum cleft along posterior margin; see
figure for details of eighth sternum and bursa.

Length: Male body 1.5 mm., wing 1.5 mm.; female body 2.7 mm.,
wing 1.9 mm.

Type-locality: Maromandia, d’Analalava, Madagascar; 1921, (R.
Decary).

Type: (MNHN).

Female description based on 9, Bikiley, Reo sud de I’lle; September
1926, (A. Seyrig) (MNHN).

Also examined: 1 co Aldabra Island, Seychelles (J. C. Fryer)
(BMNH).

66. Scenopinus nitidifrons (Kréber)
Omphrale nitidifrons Kréber, 1913, p. 194.

The type of this species has been lost. It is my opinion, however,
that this is in reality the female of S. tarsalis Kréber on the basis
of the descriptions, the locality and collection data, and general shape
of the wings. It is not uncommon to have slight differences in the
coloration of wings, halteres and legs in the members of this group.

See S. tarsalis (Kréber) and figure 71, p. 107.

102 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 66.—Scenopinus minusculus (Séguy), male, female: a, wing; b, d, lateral and frontal
aspects of male head; c, ¢, lateral and frontal aspects of female head; f, enlarged detail of
male antenna; g-i, ventral, lateral and posterior aspects of male terminalia; j, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 103

67. Scenopinus nitidulus Loew
FIGURE 67
Scenopinus nitidulus Loew, 1873, p. 149. Kréber, 1913, p. 193.

This species appears to be correctly identified in most collections.
Specimens have been seen from Egypt, Palestine, and Crete. The
type male comes from the Transcaspian Region and was examined
by me; the females agree. Kréber redescribed this species, including
the female sex from Crete.

Length: Male body 2 mm., wing 1.5 mm.; female body 2-2.5 mm.,
wing 2 mm.

Type-locality: Balfrush, Iran (Christoph).

Type: (ZMH).

68. Scenopinus parallelus, new species
FicurE 68

Matr.—Head black; eyes red-brown above, black-brown below;
frons small, triangular, shining, flat; ocellar tubercle elevated, shining;
ocelli clear. Mouthparts well developed; palpi short; oral cavity
bordered by gray pubescence. First antennal segment very short,
black; second larger, red-brown; third segment black-brown, pear-
shaped with a fine hairy covering.

Thorax dorsum black-brown with fine hairs; humeral callus red-
brown; pleural areas red-brown. Wings white, veins tan anteriorly,
faint posteriorly; veins R5 and M1+2 run parallel to each other to
tip of wing. Halter stem brown, knob white. Legs brown with lighter
tarsi.

Abdomen red-brown, segments 3-5 with broad white posterior
bands, with a distinct red-brown band between. Terminalia red-
brown; see figure for details.

FrmMALE.— Unknown.

Length: Male body 2.2 mm., wing 1.7 mm.

Type-locality: Villa Cisneros, Rio de Oro, Spanish Sahara;
June (L. Berland).

Holotype: Male and two paratypes (MNHN), one paratype
(USNM).

69. Scenopinus pilosus (Séguy)
FicurE 69
Omphrale pilosa Séguy, 1930, p. 111.

This species is known only from the type and is characterized by
the body being thinly covered with medium long hairs, and by the
wing, where M1+2 bends posteriorly to end well behind the tip of
the wing. See figure for details of male terminalia, head, and wings.

104 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 67.—Scenopinus nitidulus Loew, male, female: a, wing; }, lateral aspect of male
head; c, lateral aspect of female head; d, ¢, enlarged details of male and female antennae;
f-h, ventral, lateral and posterior aspects of male terminalia; i, ventral right and dorsal left

aspects of female 8th and 9th segments; j, lateral aspect of female 8th and 9th segments;
k, female 9th sternum and bursa.

FrmaLe.— Unknown.
Length: Male body 2 mm., wing 1.8 mm.

SCENOPINIDAE OF THE WORLD 105

Ficure 68.—Scenopinus parallelus, new species, male: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of termi-
nalia.

Figure 69.—Scenopinus pilosus (Séguy), male: a, wing; }, c, lateral and frontal aspects of
head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of terminalia.

Type-locality: Bou Knadel, Morocco (Athiery).
Type: (MNHN).

106 U.S. NATIONAL MUSEUM BULLETIN 277

70. Scenopinus reduncus, new species

Figure 70

This species is closely related to S. lincinus, S. fraternus, S. fryeri,
S. nitidulus and perhaps to S. longiwentris. It can be separated by
the wing venation where cell R5 is sharply narrowed at the tip of
the wing and by the shape of the bursal plate.

Ficure 70.—Scenopinus reduncus, new species, female: a, wing; b, c, lateral and frontal as-
pects of head; d, enlarged detail of antenna; ¢, dorsal aspect of 8th and 9th terga with
outline of 8th sternum; f, lateral aspect of 8th and 9th segments; g, ventral aspect of 9th
sternum and bursa.

Frma.tre.—Head black-brown; eyes black-brown, with a narrow,
sunken postocular ridge; frons broad subshining, with a median
depression on the lower half, ocellar triangle not set off, median ocellus
sunken; ocelli red-orange. Mouthparts well developed, brown; palpi
shorter than rostrum and labellum; back of head concave, pollinose.
Antennae black-brown; first segment short; second twice as large as
first; third pear-shaped, velvety with a distolateral pit.

Thorax black-brown, subshining, a few sparse hairs. Humeral
callus with a tan lateral spot. Wings smoky-brown, veins brown;
halter stem tan, knob tan, velvety above, white below. Legs black-
brown with red-brown tarsi.

Abdomen red-brown subshining. See figure for details of eighth
and ninth segments and bursa.

SCENOPINIDAE OF THE WORLD 107

Maue.— Unknown.

Length: Female body 3.4 mm., wing 2.5 mm.

Type-locality: Ambobitantely, Madagascar; 8 June 1958, (F.
Keiser).

Holotype: Female (NMB).

71. Scenopinus tarsalis (Kréber)

FiGurE 71 (after Kroéber)

Omphrale tarsalis Krober, 1913, p. 196.

I have not seen this species as the type has been destroyed. However,
it is my belief that S. nitidifrons probably is a female of this species.
From examination of the plate figures and collection data, it is hard
to see why these were identified as separate species. The slight dif-
ferences in wing venation, and coloration of the halteres and legs
is common among members of the Scenopinidae.

Ficure 71.—Scenopinus tarsalis (Krober), male, female: Wing and
head of male (tarsalis) and of female (as nitidifrons) (after Krober).

Length: Male body 1.5-2 mm., wing ?; 2 body 2.5 mm., wing ?.
Type-locality: Assab (on lower Red Sea), Eritrea (Katona).
Type: (MNH) destroyed.

72. Scenopinus turneri, new species

FIGURE 72

Maxe.—Head black-brown, eyes red-brown above, darker below;
frons elongate triangular, granular with a median shining groove;
ocellar tubercle red-brown, elevated, with redish ocelli. Mouthparts
normal. First two antennal segments black-brown; first short; second
flairing, twice as long as first; third segment red-brown, pear-shaped.

313-168 O—69 8

108 U.S. NATIONAL MUSEUM BULLETIN 277

Thorax black, granular with short white hairs. Humeral callus
red-brown; wings smoky hyaline, veins brown; halter stem black-
brown, knob red-brown. Legs with all segments dark red-brown.

Abdomen red-brown, segments 3-5 with wide white dorsal bands
separated by narrow strips of red-brown. Terminalia red-brown; see
figure for details.

Figure 72.—Scenopinus turneri, new species, male: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of termi-
nalia.

Frma.e.— Unknown.

Length: Male body 2.8 mm., wing 2.2 mm.

Type-locality: Aliwal, n. Cape Province, [Republic of] South
Africa; December 1922, (R. E. Turner).

Holotype: Male (BMNH).

73. Scenopinus unifasciatus (Kréber)
FIGurE 73
Omphrale unifasciata Kroéber, 1913, p. 195.

The type of this species has been lost. The specim en figured from
Cyprus agrees with Kroéber’s figures for S. wnifasciatus and is presumed

SCENOPINIDAE OF THE WORLD 109

to be that species. The previously unknown female is also believed to
belong to this pecies.

Maue.—Head black; eyes dark red-brown above, black below;
frons rugose with a median furrow ending in a shallow pit where the
eyes become contiguous. Ocellar tubercle prominent, with red ocelli.
Antennae with first segment short, brown; second red-brown, twice as
large as the first; last segment brown, pear-shaped, ending in a single
point (Kréber says a truncated cone).

Ficure 73.—Scenopinus unifasciatus (Kréber), male, female: a, male wing; b, female wing;
c, d, lateral and frontal aspects of male head; ¢, f, lateral and frontal aspects of female
head; g, h, enlarged details of male and female antennae; ik, ventral, lateral and posterior
aspects of male terminalia; /, lateral aspect of female 8th and 9th segments; m, dorsal
aspect of female 8th and 9th tergites with 8th sternum outlined; n, female 9th sternum and
bursa.

Thorax dorsum black, rugose with sparse white hairs; humeral and
supra-alar calli red-brown; pleura black-brown. Wings smoky brown,
veins brown; halter stem black-brown, knob white with a tan base.
Legs uniform red-brown throughout.

Abdomen black-brown, granulose, segments 3-5 with posterior white
bands. Terminalia black; see figure for details.

FrmaLre.—Head red-brown; eyes brown, with a medium broad post-
ocular ridge; frons rugose with a smooth median groove extending to
the median ocellus which is depressed, and swelling on either side of
ocellar triangle, hairs on lower frons directed ventrolaterally. Ocellar
triangle not set off, ocelli orange-red. Mouthparts brown, well devel-
oped; sides of oral opening lined with brown pubescence. Antennae

110 U.S. NATIONAL MUSEUM BULLETIN 277

red-brown, first segment small; second twice as large in all dimen-
sions, fringed with long hairs; third segment pear-shaped covered with
a dense pile of short golden hairs.

Thorax dorsum black-brown with a scattering of hairs; humeral
callus red-brown with white posterior border; supra-alar callus tan;
pleural areas red-brown; wings brown fumose, veins brown; halter
stem brown, knob white. Legs red-brown, femur with thinly scattered
hair; tibia densely haired, tarsi yellow-brown basally darkening
distally.

Abdomen red-brown, shining, with a few hairs; eighth segment with
short stiff hairs directed outward; ninth tergum with heavy spinelike
hairs. See figure for details of eighth sternum and bursa.

Length: Male body 2.2 mm., wing 1.8 mm. (Kréber gives length
2.5 mm., wingspan 3 mm.); female body 3.5 mm., wing 2.3 mm.

Type-locality: Island of Poros, Greece.

Type: (MNH) destroyed.

Male redescribed from Yermasoyia Hills, Cyprus; 11 September
1951, (G. A. Marromoustabis) in (BMNBH).

Female described from Adana, Karates, Turkey, sea level; 7 June
1960, (Guichard & Harvey) in (BMNH).

VELUTINUS group

Key to Species of the VELUTINUS group

NEARCTIC SPECIES (MALES)

1...» Wineihy Aline. ..g\0. 5, iscickvicay ol emyi dca ia hs gy een eae meee
Wing brown: Or white \. 5. iets <a divaid oltees We So eaion Samed eee Gare Ue
2... Walter knob; WHC 42. ¢ <7 intone sectey ations’ eeneunthne ba Woe bebeireadee a) heaae
Halter knob other color. .. . Saket sloth sida pAb et ath eee ceed tO
3. Abdomen with three white Herds hs bseeeh Raindal ll) Me aR aids el So eae
Abdomen with two white bands. . ... . . . S. werneri, new species
4. Posterior margin of 9th tergum excavated. ........4..... 9

Posterior margin of 9th tergum smoothly rounded
S. beameri (D. E. Hardy)
9th tergum with a flaplike notch on ventral posterior border, antenna long
S. electus Adams

on

9th tergum without a ventral notch. ... . . . S. whittakeri James
6; -;Halter:knob brown. Or'red- brow, david) alse 2) ie oeteme te seeker
Halter knob tan or yellow ... . Cae Micah (BE me ee ocala
7. Halter knob red-brown, 9th tergum eerie. WSR eos ce! Nal fe erties Mag

Halter knob brown, 9th tergum smoothly rounded
S. beameri fuscus (D. E. Hardy)

S:= Abdomen:with three whiteibandsie< i412. (i Bae Seay Sn
Abdomen with two white bands. . ... . . . S.gossypius, new species
9. First band narrow, second and third broad. . . . S. hagai, new species

First band broad, second andthird narrow. . . S. magdalenai, new species

10.

ll.

13.

14.

16.

7.

18.

19.

21.

22.

23.

SCENOPINIDAE OF THE WORLD EY

Halter knob tan, posterior margin of 9th tergum toothed ventrally
S. serratus, new species
Halter knob yellow, es ventral tip of 9th tergum long with a single

NOtCh yas ee a... eww ee ))6.)|CUS. «6mirabilis Adams
Wing brown or smoky Grom Wie cee. Ss. 21 ee Ow Th, © el ee Be we oes
Winw wiite-or milky Wwhit@;c Os «<a gol 4 of eee ele a oe ee ONT
Halter knob white. . .. . eerie ee vege ee mee ata ee nes ec
Halter knob black-brown or ea Gromit mew .... 14
Posterior margin of 9th tergum with a long Sorat et parameres long,

Wity: s ; .. . . . §.cochisei, new species
Posterior faarain) e 9th fercim Sik a fennel notch and point, parameres

long, wiry. . . . i.e ee ew 66h. «6S. Kuiteri (D. E. Hardy)
Halter knob red- Brew Tey ce a bee pa ees oe ee eee alo
Halter knob browntoblack. . ........ . S.valgus (D. E. Hardy)
Abdomen with three white bands . ......... eG
Abdomen with two white bands. ......... ‘S. mubilines Say
Bands on abdomen relatively broad. . . . . . S. flandersi, new species
Bands on abdomen narrow. ...... .. . . . S. rossi, new species
Halter knob white. . ... Bee ee cee beg a ee ls
Halter knob brown, tan, orcream. ..........2.2.2.+.. =. 24
Abdomen with three white bands . ........ See eae LO
Abdomen with two white bands. ......... S. aeukt, new species
All white bands broad. . . Ae | bese ee 20
First band narrow, second nad (aed mades Sse S. alalactews: new species
Body without ccales ees SS. -e: w a ha + ee |
Body with flattened scales. . . . 2... . . Ss. Saipaeetatna (D. EK. Hardy)
Small flies under 3 mm.; veins separated at tip of wing. . . .... . 22

Small flies over 3 mm.; veins R5 and M1-+2 close at tip of wing

S. cretatus, new species
Small flies under 3 mm., but over2mm...... . arcs. Ae Nes be
Small flies under 2 mm. long. . . ba tasa S. chico, new species
Veins R5 and M1+2 widely a pared A ena a wing

S. weemsi, new species
Veins R5 and M1+2 moderately separated=r-m cross vein

S. vanduzeei, new species

Halter knob brown or red-brown. ....... . CAs Ot}
Halter knob orange-yellow. ......... S. sation, new species
Halter knob red-brown. ........-.- 6+ ++8+ 2624+ ++ 26
Halter knob orange-yellow . . . . . . §. seftoni, new species

Halter knob brown, hind margin of the oth te ergum produced on the midpoint

S. cooki, new species
Third antennal segment short. . . . .. . . . . S. barberi, new species
Third antennal segment long and narrow. . . . . . S.butleri, new species

NEARCTIC SPECIES (FEMALES)

Wing hyslineta 20.0% oo 3 ew, 6 8 eee we He we ied Bue eB
Wing brown or white. . 2... 2... 2. ee eee ee ee ee ee el eG
Pinter enous). <4) pod hb ea Owe ee oe eS ed Se le
Halter knob other color. . . . eae ne ay Seer:
Lateral depressions on 8th fora nee seeds num. s. beameri (D. Ik. Hardy)
No lateral depression on 8th tergum . . ... . . . S. whittakeri James
Halter knob brown or red-brown. . .. 2... ee ee ee ee ee OD

Halter mob tancor yellows (2) 0/< <1 ecw es wee te ee ea ee eb aS

112

10.

tal

13.
14.
15.
16.
17.
18.
19.

20.

21.
22.

23.

U.S. NATIONAL MUSEUM BULLETIN 277

Cell R5 tapering to edge of wing. . . BP ena aa Ne ec ANG
Cell R5 sharply constricted before the ona of ihe Cale
S. gossypius, new species
Legs with tibia red-brown. . . . SAAS Shas Nace tis ta aghast ROA tee heh
Legs with tibia orange-brown to Beonne . . . §. magdalenai, new species
Eighth tergum flat with lateral depressions
S. beameri fuscus (D. E. Hardy)

Eighth tergum rounded without depressions. . . . S. valgus (D. i. Hardy)
Halter knob tan, tip of cell R5 sharply closed . . S. howdeni, new species
Halter knob yellow, tip of cell R5 open... . . . . S. mirabilis Adams
Wine brown - orsmoky brown sere h ota ta ba hy, Peeks (eee eet ea)
Wing whitevorimiliky whiten 2 iS) 75 ts jie fee ey ee tae aero aa deena IG
Haltemknolb: ligh tin face. x5) vias ee4 Sletten WS Seek, Rea sees EL
altentkmoOpyaatkey 235,35... Pu ea the, ha ny SRA ee Ee A Se ee een eres, lies
Halter ‘knob cream.or tan...) ..73 (0E"S oe STOR iE
Halter knob white ... . Sete Oe ‘Ss. vandtizeet new species
Halter knob cream, antenna ion! piratane . . . §. kuiterti (D. E. Hardy)
Halter knob tan, antenna short, pear-shaped. . . . S.cochisei, new species
Cell: Rb narrowed) atvedgevol withee ve) ee in een or eee ae eee,
Cell R5 open at edge of wing. . . cena) a hen Re ery
Halter knob red-brown to black Grows polished’ A S. nubilipes Say
Halter knob red-brown above white below . . . . . S. hagai, new species
Tarsi light red-brown 00) 30) 000.) fo ee Sip ryanti, new: species
Parsistraw: yellows) 60... 2° 8) ww SOY Ses S landers; newspecies
Halter knob: White + 4. a- 554s 0d-s6% Seo ce eek ey AN Ne ces amen Leh
Halter: knob: browmor tan .-. 2, .- oh oF ge EP ee de See
Halter knob: all: white 3) 7°08 OMe a a ls aby etna MES
Halter kaiob with base“tan- (325) 808 So. Ss. weemsi, new species
Body without flattened scales . . ... . eee BLO
Body with flattened scales... 2.2%, 2% s. ‘aihifacciatns (D. E. ard
Third antennal segment not parallel-sided. . . . . . BRE 22 ()
Third antennal segment uniform width. .... . Ss. johicas new species

Third antennal segment oval, widest at middle. .S. alalacteus, new species
Third antennal segment pear-shaped, widest at base
S. cretatus, new species

Halter knob brown orred-brown. ....... 5 PURE ai
Halter‘knob orange-yellow ))i5-).... - 2) 8 BPRS: ug: seftoni, new species
Halter knob red-brown . . . ae scweaee as oR ERES. LD
Halter knob brown above, white Below ve 3 Ss. peckd, new species
Antenna black-brown elongate oval, frons pwollén above antenna with a

broad median depression on lower fourth. . . . . S. rossi, new species
Antenna red-brown, short, frons swollen above antenna with a median

groove dividing lower two-fifths. . . . . . . . .§. barberi, new species

NEOTROPICAL SPECIES

Males . se
Females . Late IEICE MOOT sO
Wing hyaline, Aidamen wiht ce or three nies Shanice Boetet th, eS,
Wing milky white, two white bands on abdomen. . .S. ea omimens hoce
Halter knob brown or red-brown, abdomen with three white bands . 4
Halter knob white, abdomen with two white bands (Hawaii)

S. adventicius (D. E. Hardy)
Halter knob red-brown, abdomen with some narrow white bands . . . .5

SCENOPINIDAE OF THE WORLD 113

Halter knob dark brown, abdomen with three broad white bands
S. velutinus (Kréber)

5. Abdomen with three narrow white bands. . . . .S. buscki, new species
Abdomen with first two bands broad, third narrow. . S. schultzi Enderlein
Ce Ce yee eee ee sie a ie ete. bags amare. 2 eh oe
Wane Drown OD WHE! .c2 9. "ols Mes My « <hiag Ge Bow 2 bead oe a, SA
7. Halter knob red-brown or brown .........2.2.2.2.2.2.2.2..8
Halter knob white (Hawaii). . . . . . . . S.adventicius (D. E. Hardy)
Soma alver enO TeG-brOown ) gnu) 4. «es eek. Meth aoe ok oe. Bao G
Halter knob brown. . . . aoe oe a UO

9. Eighth sternum with a prominent nipple.ike swelling before end
S. schulzi Enderlein

Kighth sternum without suchswelling. . . . . . . .S.buscki, new species
10. Halter knob dark brown ......... .. .. . §. velutinus (Kréber)
Halter knob brown above, white below. . . . . . . S. fijianus (Kroéber)
11. Wing milky white, halter white. ....... . .. . S. pygmaeus Loew
Wing brown, baltver .browi 2M. e . katt alee ee eS ia ee ae
12. Antenna with third segment very small . . . . . S. curticornis (Kréber)
Antenna with third segment very large. . . . . S. magnicornis (Krober)

74. Scenopinus adventicius D. E. Hardy
Ficure 74

Scenopinus adventicius D. E. Hardy, 1960, p. 329.

This dark-winged species, along with S. papuanus, has been er-
roneously identified as niger (DeGeer), particularly in the more
frequently captured females. It can be distinguished by its smaller
size and by R4 branching well beyond the middle of the cell.

Ficure 74.—Scenopinus adventicius D. E. Hardy, male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; hA-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

114 U.S. NATIONAL MUSEUM BULLETIN 277

Length: Male body 3-3.7 mm., wing 2.3-2.6 mm.; female
body 4—4.5 mm., wing 3.5 mm.

Type-loca-ity: Honolulu, Oahu, Hawaii; January 1953, (D. E.
Hardy).

Types: B. P. Bishop Museum, Honolulu.

75. Scenopinus alalacteus, new species

FicurE 75

Mauxr.—Head_ black-brown; eyes red-brown above, black-brown
below, extending to back of head; frons subshining, narrow, tri-
angular, extending halfway to ocellar tubercle, with a median de-
pression; ocellar triangle black-brown, elevated, ocelli clear; back of
head concave, lower portion with long white hairs; mouthparts
brown, nearly filling oral cavity, palpi tan, nearly as long as rostrum.
Antennae red-brown: first segment short; second twice as large as
first; third oval, twice as long as first two combined.

Ficure 75.—Scenopinus alalacteus. new species, male, female: a, wing; b, d, lateral and frontal
aspects of male head; c, lateral aspect of female head; ¢, f, enlarged frontal and lateral
aspects of female head; g, enlarged detail of male frons; h-j, ventral, lateral and posterior
aspects of male terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of
female 8th and 9th segments; m, female 9th sternum and bursa.

Thorax dorsum black-brown with a patch of silvery pile above
the humeral callus, humeral and supra-alar calli tan; pleura red-
brown. Wings milky white, veins tan; halter stem brown, knob
white. Legs red-brown except tarsi straw colored at base, darker
at tip.

SCENOPINIDAE OF THE WORLD LE5

Abdomen red-brown, segment 3 with a narrow white band, broad
bands on segments 4 and 5. For details of terminalia see figure.

Frma.e.—Head dark red-brown; eyes red-brown with a broad
postocular flange elevated above the eyes on the top of the head;
frons slightly swollen above antennae, a median groove extending
nearly to median ocellus; ocellar tubercle broad, flattened and not
distinctly separated from the dorsal areas of the frons. Mouthparts
as in male; oral cavity surrounded by a band of silvery pile which
extends over the antennae in a narrow band; back of head concave;
extensive genae below eyes. Antennae red-brown, as in male.

Thorax, wings, and legs as in male.

Abdomen red-brown; see figure for details of segment 8 and bursal
plate.

Length: Male body 2.5-3 mm., wing 2-2.2 mm.; female body
2.6-3.1 mm., wing 1.9-2 mm.

Type-locality: Glenns Ferry, Idaho; 14 June 1927.

Holotype: Male (USNM) 67476.

Allotype: Female same but 8 July 1927, (USNM).

Paratypes: 2 co, 3 99, Glenns Ferry, Idaho, 8 July, and
5 September 1927, in (USNM); 1 <o, Sparks, Nevada, 28 June 1923,
(E. P. Van Duzee) (CAS); 1 o&, Adelaide, Idaho, 26 July 1929,
(UCalB).

76. Scenopinus albifasciatus (D. E. Hardy)

FIGURE 76

Omphralosoma albifasciata D. E. Hardy, 1944a, p. 41.

Except for the presence of scalelike hairs, this species agrees in all
other respects with other members of the Velutinus group.

Length: Male body 2.3 mm., wing 1.7 mm.; female body 2.66 mm.,
wing 2 mm.

Type-locality: Seeley, California; 17 July 1940, (R. H. Beamer).

Allotype: Female, Jacumba, California; 12 August 1935, (R. H.
Beamer).

Paratype: 1 co’, same data as type.

Types: (UKan).

I have seen a long series from Arizona in (USNM), (UAriz) swept
from alfalfa. Also 1, Mesilla Park, New Mexico (C. N. Ainslie)
(USNM).

77. Scenopinus barberi, new species

FIGURE 77

Maur.—Head dark red-brown; eyes brown above, red-brown below,
reaching the back of the head, contiguous for a greater distance than
the height of the frons, which is triangular with an elongate depression
in the upper half; ocellar tubercle red-brown, ocelli clear. Mouthparts

116 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 76.—Scenopinus albifasciatus (D. E. Hardy), male, female: a, wing; }, d, lateral
and frontal aspects of male head; c, ¢, lateral and frontal aspects of female head; f, g,
enlarged details of male and female antennae; h-j, ventral, lateral and posterior aspects
of male terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female
8th and 9th segments; m, female 9th sternum and bursa.

Ficure 77.—Scenopinus barberi, new species, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontai aspects of female head; f, enlarged detail
of male antenna; g, enlarged lateral view of female head; h, 7, ventral and lateral aspects
of male terminalia; 7, ventral aspect of female 8th sternum; &, lateral aspect of female
8th and 9th segments; /, female 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD LI7

brown, well developed, palpi orange-brown, two-thirds as long as
rostrum; an extensive postgenal area on lower part of the head.
Antennae red-brown; first segment small; second bell-like, much
larger; third pear-shaped, less than twice as long as broad.

Thorax red-brown, humeral and supra-alar calli red-orange. Wings
opaque, with milky microtrichia, veins light brown; halter stem brown,
knob red-brown. Legs red-brown with lighter tarsi.

Abdomen red-brown, segments 3-5 with broad white stripes sepa-
rated by thin sclerotized bands. Ninth tergum similar to S. electus but
more prominent and posterior emargination different; see figure for
details.

Frema.e.—Head orange-brown, eyes red-brown; frons very broad,
wider than the eye when viewed from the front, rugose, swollen over
the antennae and with a median groove dividing the lower two-fifths
of the frontal area; ocellar tubercle flattened and not distinctly
separated, ocelli clear. Mouthparts well developed but lighter in color
than the male; antennae as male except lighter.

Thorax dorsum orange-brown; humeral callus orange; pleura orange.
Wing as in male; halter stem brown, knob brown. Legs orange-brown
with straw-yellow tarsi.

Abdomen orange-brown, subshining; eighth segment red-brown. See
figure for details of sternum and bursa.

Length: Male body 2.2 mm., wing 1.7 mm.; female body 2.7 mm.,
wing 1.6 mm.

Type-locality: Bright Angel, Arizona; 10 July, (H. S. Barber).

Holotype: Male in copulo with allotype female (USNM) 67479.

Paratype: 192, same data (USNM).

78. Scenopinus beameri (D. E. Hardy)

79. Scenopinus beameri var. fuscus (D. E. Hardy)

FIGURE 78

Omphrale beameri D. E. Hardy, 1944a, p. 43.
Omphrale beameri var. fusca D. E. Hardy, 1944a, p. 43.

These two forms are similar except for the coloration of the halteres.

Length: Male body 2.3-2.5 mm., wing 2-2.3 mm.

Type-locality: of beameri, Olancha, California, 25 August 1940,
(R. H. Beamer); of var. fusca, Onyx, California, 23 July 1940,
(R. H. Beamer).

Types: (UKan).

I have seen a long series of undetermined specimens from the same
localities and collection dates as the types and paratypes in (UKan).
Also: 19, 1c, Fallon, Nevada, 12 August 1940, (L. Lipovsky & E.
Kenaga); 1 @, Oak Creek Canyon, Arizona, 9 July 1941, (L. H.

118 U.S. NATIONAL MUSEUM BULLETIN 277

Wage KK

Ficure 78.—Scenopinus beameri (D. E. Hardy), male, female: a, wing; b, d, lateral and
frontal aspects of male head; c, e, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; hA-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of female 8th and 9th segments (Dep.=depression); /, lateral
aspect of female 8th and 9th segments; m, female 9th sternum and bursa.

Banker); 1 o, 1 @, in copula, Prescott, Arizona, 20 June, (H. S.
Barber) (USNM).

80. Scenopinus bryanti, new species

FIGURE 79

FrmMaLe.—Head dark red-brown; eyes black-brown, with a thin
postocular rim; frons rugose, swollen above the antennae, with a
median elongate depression on the lower half, thinly covered with
short hairs; mouthparts well developed, palpi long but not as long as
rostrum; a band of silvery pile around the oral opening not extending
to eye margins or above antennal bases. Ocellar triangle two-thirds
width of frons and set off from frontal area; ocelli orange-red. Antennae
red-brown; first segment short; second twice as large; third not twice
as long as broad; see figure.

Thorax dorsum dark red-brown; humeral and supra-alar calli and
scutellum red-brown; pleurum red-brown. Wings hyaline, dusted with
brown, veins brown; veins R5 and M1 + 2 widely separated on wing
margin, cell R4 small. Legs with red-brown femora and tibiae, tarsi
light red-brown.

Abdomen red-brown, eighth tergum concave on posterior margin.
See figure for details of eighth segment and bursa.

Matr.— Unknown.

Length: Female body 2.8 mm., wing 2.2 mm.

SCENOPINIDAE OF THE WORLD 119

Ficure 79.—Scenopinus bryanti, new species, female: a, wing; 6, c, lateral and frontal
aspects of head; d, enlarged aspect of antenna; ¢, ventral left and dorsal right aspects of
8th and 9th segments; f, lateral aspect of 8th and 9th segments; g, 9th sternum and bursa.

Type-locality: Tucson, Arizona; 23 April 1937, (Bryant).
Holotype: Female (CAS) 8927.

81. Scenopinus buscki, new species

FIGURE 80

Maur.—Head black-brown; eyes bright red-brown above, black-
brown below, reaching the back of the head; frons small, triangular,
with a median groove extending from above the antennae to a median
pit in the upper fourth of the frons; postgenal areas small; mouthparts
red-brown; palpi very nearly as long as rostrum; silvery pile between
eye and oral cavity extending dorsally to lower angles of frons. First
antennal segment small, black-brown; second twice as large as first,
red-brown; third segment red-brown, over twice as long as broad,
square-tipped and covered with scales. Ocellar tubercle prominent;
ocelli red-brown.

Thorax dorsum black-brown, with silvery pile above the humeral
callus and extending in a thin band along the tergal margin behind
the callus; humeral and supra-alar calli red-brown; pleura red-brown.
Wings hyaline, veins light brown; R5 and M1 + 2 close together
bending forward to end in the margin of the wing before the tip;
halter stem red-brown, knob red-brown, shining. Legs with femora
and tibiae red-brown, fore and middle tarsi orange, hind tarsus
straw yellow.

120 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 80.—Scenopinus buscki, new species, male, female: a, wing; b, c, laterai and frontal
aspects of male head; d, e, frontal and lateral aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia
(apodeme of ejaculatory syringe lost on dissection); k, ventral aspect of female 8th ster-
num; /, lateral aspect of female 8th and 9th segments; m, female 9th sternum and bursa.

Abdomen red-brown; segments 3-5 with narrow white bands on
posterior half of segments. See figure for details of ninth tergum and
internal structures which are very large.

» OF THE WORLD 121

Frema.Le.—Head dark red-brown; eyes black-brown with a thin
postocular ridge; frons swollen above the antennae, with a median
groove extending into a large shallow median depression; genal area
small; mouthparts brown, large and well developed; oral cavity
bordered by a silvery pile that extends to base of antennae; ocellar
tubercle not prominent, median ocellus set in a depression. Antennae
as in the male.

Thorax, wing, halter and legs as in male except tarsi yellow-orange.

Abdomen red-brown. See figure for details of eighth segment and
bursa.

Length: Male body 3.5 mm., wing 2 mm.; female body 3.5-4.5
mm., wing 2.3-2.8 mm.

Type-locality: ua Chorrera, Panama; 17 May 1912, (August
Busck).

Holotype: Male, allotype female, same data (USNM) 67480.

Paratypes: 3 99, Cabima, Panama, 22 and 26 May, 28 August
1911 (A. Busck); 1 9, Corozal, C.Z., Panama, April 1911 (A. Busck) ;
1 9, Kingston, Jamaica, 8 October 1882 (M. Grabham); 1 9, in auto
from Mexico, 11 February 1952, at Brownsville, Texas; all in (USNM).

82. Scenopinus butleri, new species
FicureE 81

This species is close to S. mirabilis but differs in the length as well
as the shape of the third antennal segment and in the aedeagus and
ninth segment claspers.

Ma.e.—Head black; eyes red-brown above, dark red-brown below;
frons narrow, triangular, rugose, subshining, a few hairs on either
side of lower frons tend to make it look narrower; ocellar tubercle
prominent, ocelli orange-red. Antennae with first antennal segment
black-brown, moderately long; second segment black-brown, long,
the first two half as long as the third; third segment red-brown,
elongate, truncated, three times as long as widest point.

Thorax dorsum black, a deep lateral groove above the wing base;
humeral callus red-brown; pleural areas black-brown. Wings smoky
hyaline, veins light brown; R5 and M1 + 2 widely separated, M1 + 2
ending at tip of wing. Legs with femora and tibiae black-brown, fore
and middle tarsi dark red-brown, hind tarsi red-brown.

Abdomen black-brown, segments 3-5 with broad white stripes;
ninth segment black-brown; see figure for details. Aedeagus short
and stout, parameres long and stout, ninth segment claspers stout
with depressed area of tip next to aedeagus, longer tip lateral; cup of
syringe blackish.

FremaLe.— Unknown.

Length: Male body 2.7 mm., wing 2.5 mm.

122 U.S. NATIONAL MUSEUM ae 277

Figure 81.—Scenopinus butleri, new species, male: a, wing; b, c, frontal and lateral aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of
terminalia.

Type-locality: Cochise Stronghold, Dragoon Mts., Arizona; 24
April 1957, (F. Werner and G. Butler).
Holotype: Male (USNM) 68009.

83. Scenopinus chico, new species

FIGURE 82

This very small species from the Southwest has been given the
Spanish diminutive as its specific name.

Mauy.—Head black; eyes brown-black above, black below; frons
shining, swollen above the antennae and extending as a narrow band
to the ocellar tubercle, which is prominent, shining, with orange
ocelli. Mouthparts dark brown, well developed; oral cavity with silvery
pilosity extending to the base of the antennae; back of head concave.
First antennal segment short, black; second segment brown, larger
than first; third segment brown, long slender, twice as long as 1 and 2
combined.

Thorax dorsum black, with flat silvery scalelike hairs, a thick patch
on the posterior portion of the dorsum; humeral callus with a red-
brown margin; pleural areas dark red-brown. Wings smoky white,

SCENOPINIDAE OF THE WORLD 123

Figure 82.—Scenopinus chico, new species, male, female: a, wing; b, c, lateral and fronta!
aspects of male head; d, e, lateral and frontal aspects of female head; f, enlarged detail of
antenna; g-7, ventral, lateral and posterior aspects of male terminalia; 7, ventral aspect
of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female 9th
sternum and bursa.

veins straw yellow; R5 and M1 + 2 widely separated; halter stem
brown, knob white. Legs with basal portions through the femora
red-brown, tibiae light brown, tarsi straw yellow with darker tips.

Abdomen short, compressed, dark red-brown; segments 3-5 with
broad white bands dorsally, remainder red-brown. For details on
shape of ninth tergum and genital apparatus, see figure.

Fremate.—Head black-brown; eyes black-brown; with the narrow
postocular ridge depressed below the level of the eyes; frons broad,
subshining, rugose with a median groove extending from the antennae
to the median ocellus; ocellar triangle slightly elevated, ocelli clear;
mouthparts well developed. Antennae as in the male.

Thorax dorsum red-brown with thin scalelike hairs, sparse, no
dorsal patch as in the male. Other characters as in the male.

Abdomen red-brown, shining, with a few hairs. See figure for details
of eighth segment and bursa.

Length: Male body 2—2.2 mm., wing 1.6-1.8 mm.; female body
2.7 mm., wing 1.8 mm.

313-168 O—69—_9

124 U.S. NATIONAL MUSEUM BULLETIN 277

Type-locality: El Centro, California; 27 June 1917, (J. M.
Aldrich).

Holotype: Male (USNM) 67474.

Allotype: Female, Palm Springs, California, 10 April 1946,
(A. L. Melander).

Paratypes: 20’o’, same data as type; 1c’, Las Vegas, Nevada,
9 September 1930, (E. W. Davis) (USNM); 1c’, Palm Springs,
California, 3 April 1955, (W. R. M. Mason) (CNC); 10, Calexico,
California, 11 September 1952, (E. I. Schlinger); 1<7, Borrego, San
Diego Co., California, 3 May 1956, (M. Wasbauer) (UCalD); 107,
near Segurra, California, 14 April 1949, (Timberlake); 1c, 2 miles

16, Sentinel, Arizona, 23 July 1941, (R. H. Beamer) (UKan).

84. Scenopinus cochisei, new species

Figure 83

Maue.—Head black-brown; eyes dark brown-red above, black-
brown below; frons small, triangular, extending half-way to the ocellar
triangle, shining, margined with shallow pits along the eye margin;
ocellar tubercle prominent, ocelli clear. Mouthparts brown, well
developed, the palpi red-brown, shorter than the rostrum; antennae

Ficure 83.—Scenopinus cochisei, new species, male, female: a, male wing; b, female wing;
c, d, lateral and frontal aspects of female head; ¢, f, lateral and frontal aspects of male
head; g, A, enlarged details of male and female antennae; 7k, ventral, lateral and posterior
aspects of male terminalia; /, ventral aspect of female 8th sternum; m, lateral aspect of
female 8th and 9th segments; 7, female 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 125

with first segment very small, black-brown; second segment brown,
rounded, much larger than the first; third segment black-brown,
pear-shaped only a little longer than broad.

Thorax dorsum black-brown, humeral and supra-alar calli red-
brown; pleural areas dark red-brown. Wings smoky, veins tan; halter
stem red-brown, knob white. Legs dark red-brown, except. tarsi
which are orange-brown.

Abdomen short, red-brown, segments 3—5 extensively white dorsally ;
ninth tergum with dorsal points; aedeagal parameres long, like S.
mirabilis. See figure for details of terminalia.

FremMaLe.—Head red-brown; eyes black, with a narrow pollinose
postocular ridge; frons broad, subshining, red-brown below, black
above, with a broad shallow depression on the lower portion. Ocellar
tubercle flat but marked off from frontal area; mouthparts extending
only half the length of the oral cavity; palpi sub-equal; inner surface
of oral cavity pollinose; lateral portions to the eye broad, shining.
Antennae with first segment dark red-brown, subshining; second
segment red-brown, pollinose, as long as first segment but wider;
third segment red-brown, pollinose, about twice as long as broad, very
little taper.

Thorax dorsum dark red-brown; humeral callus with a posterior
orange patch; supra-alar callus orange-brown, wings hyaline, smoky
brown; veins brown; halter stem dark brown, knob creamy with tan
margins. Legs with femora dark red-brown, remainder red-brown.
Abdomen red-brown. See figure for details of eighth segment and bursa.

Length: Male body 2 mm., wing 1.7 mm.; female body 4 mm.,
wing 3 mm.

Type-locality: Chiricahua Mts., Arizona; 4 July 1940, (L. C.
Kuitert).

Holotype: Male (UKan) 6289.

Allotype: Female, Rustlers Park, Chiricahua Mts.; 5 July 1940,
(L. C. Kuitert) (UKan).

Paratype: Male Calexico, California; 8 April 1947, (A. L.
Melander) (USNM).

85. Scenopinus cooki, new species

FIGURE 8}

Matr.—Head dark red-brown; eyes red-brown above, black-brown
below; frons small, triangular, with a large elongate median depres-
sion, shining above, and an inverted triangle below. Ocellar triangle
red-brown, prominent, ocelli orange. Back of head concave, only a
narrow postgenal strip showing behind lower half of eye. Mouth-
parts large, brown; palpi red-brown long; white pubescence between
oral opening and eye margin extending into points above the antennae.

126 U.S. NATIONAL MUSEUM BULLETIN 277

FicurE 84.—Scenopinus cooki, new species, male: a, wing; J, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of ter-
minalia, severed portion of ejaculatory mechanism outlined.

SCENOPINIDAE OF THE WORLD 127

Antennae red-brown, first segment short, second lighter distally,
wider than long, third segment pear-shaped, over twice as long as
broad, over twice as long as the first two segments combined.

Thorax dorsum dark red-brown, rugose with transverse striations,
very thinly haired, a small patch of silvery pubescence above the
humeral callus; humeral and supra-alar calli orange; scutellum dark
red-brown above, lighter below; pleural areas red-brown. Wings
milky, veins tan, halter stem brown, knob dark brown. Legs with
femora brown, tibiae orange-brown; tarsi cream basally, orange-
brown distally.

Abdomen dark red-brown, segments 3-5 broadly white margined,
ninth tergum with a medioventral swelling on posterior margin; see
figure for details.

FremaLe.— Unknown.

Length: Male body 3 mm., wing 2.1 mm.

Type-locality: Patagonia, Arizona; 11 July 1950, (Paul P. Cook).

Holotype: Male (UKan) 6290.

86. Scenopinus cretatus, new species
FIGURE 85
Maue.—Head red-brown; eyes brown above, red-brown below,

upper area twice as extensive as lower; frons triangular, rugose,
extending half the distance to the ocellar tubercle; mouthparts

Ficure 85.—Scenopinus cretatus, new species, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, enlarged
detail of antenna; g-1, ventral, lateral and posterior aspects of male terminalia; J,
ventral aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments;
/, female 9th sternum and bursa.

128 U.S. NATIONAL MUSEUM BULLETIN 277

developed, but short, less than half the length of the oral opening;
palpi short; margin of face between oral opening and eye silvery
pilose to above the antennal base; back of head concave. Ocellar
tubercle not prominent, ocelli clear; antennae red-brown, first segment
short; second twice as large as first; third elongate, pear-shaped, over
twice as long as first two combined.

Thorax dorsum black, rugose with scattered light hairs; humeral
and supra-alar calli orange-brown; pleural areas red-brown. Wings
milky, veins tan; halter stem brown, knob white. Legs red-brown
except proximal portion of tarsi orange-brown.

Abdomen red-brown, segments 3-5 broadly white except at lateral
margins where triangular pieces of sclerotized tissue remain. Termi-
nalia red-brown; see figure for details of structures.

FrmMaLe.—Head black, eyes red-brown; a medium wide subshining
postocular band extending into a broad postgenal area below and
behind the eyes; frons broad, slightly narrower than the eye when
viewed from the front, swollen above the antennae, with a median
groove and depression above the swelling, groove extending to just
below the median ocellus, surface rugose subshining; ocellar tubercle
not distinctly separated or elevated, ocelli clear to amber; mouthparts
red-brown, short, with short palpi; silvery pile between oral opening
and eye margin. Antennae as in the male.

Thorax as in male; wing veins darker:

Abdomen red-brown subshining; for eighth segment details and
bursa, see figure.

Length: Male body 3.2 mm., wing 2.3 mm.; female body 3 mm.,
wing 2.3 mm.

Type-locality: 8 mi. s. Lovelock, Nevada; 30 May 1958, (T. R.
Haig).

Holotype: Male, allotype female, same data, (USNM) 67475.

Paratype: o Adelaide, Idaho; 7 July 1927, (UCalD).

87. Scenopinus curticornis (Kroéber)
FIGURE 86
Omphrale curticornis Kréber, 1918, p. 202.

This species from Brazil is known only from the type and is easily
distinguished by the extremely short third antennal segment.

Mauze.— Unknown.

Length: Female body 4.5 mm., wing 2.8 mm.

Type-locality: Brazil; in Winthems Coll. as pygmaeus Loew.

Type: (NMYV).

SCENOPINIDAE OF THE WORLD 129

Ficure 86.—Scenopinus curticornis (Kréber), female: a, wing; b, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

130 U.S. NATIONAL MUSEUM BULLETIN 277

88. Scenopinus electus Adams
FIGuRE 87

Scenopinus electus Adams, 1904, p. 445.

This species is noted by its elongate straight antennae and by the
fold in the ventral posterior margin of the ninth tergum. The type
and allotype have not been examined and no females fitting the species
have been seen.

Length: Male body 2.8-3.3 mm., wing 2—-2.3 mm.; female body
2.5 mm.

Type-locality: Bill Williams Fork, Arizona (F. H. Snow).

Types: (UKan).

The following specimens have been seen: 1c, Gilliland Ranch,
Phoenix, Arizona, 17 May 1940, citrus sweeps (Lot No. 40-12517)
(USNM); 127, Isla Partida, Gulf of California, Mexico, 23 March
1953 (P. H. Arnaud); 10, Los Frailes, Baja California, Mexico, 18
March 1953 (P. H. Arnaud) (CAS); 107, Cedros (island), Baja Cali-
fornia, Mexico, 3 June 1925 (H. H. Keifer) (UCalB); 107, Blythe,
California, 24 April 1955 (D. F. Hardwick) (CNC).

A
2),

Ficure 87.—Scenopinus electus Adams, male: a, wing; }, c, lateral and frontal aspects of
head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of ter-
minalia.

SCENOPINIDAE OF THE WORLD 131

89. Scenopinus femoratus Macquart
Scenopinus femoratus Macquart, 1835, p. 7.

Examination of the Macquart collection in the Museum of Natural
History, Paris, failed to bring this species to light although other
Scenopinidae identified by him were in the collection.

Length: Female body 2.25 mm.

Type-locality: Brazil.

90. Scenopinus fijianus (Kréber)

Figure 88

Omphrale fijianus Krober, 1939, p. 399.

The type was examined and figured as far as possible, the antennae
and abdomen of the type are missing so could not be figured; a copy
of Kréber’s detail of the antenna is included in the plates. A speci-
men in the (USNM), removed from an airplane in quarantine on
Oahu, Hawaii, which agrees with the wing of the type, is figured.

Length: Female body 3 mm., wing 2.5 mm.

Type-locality: Suva, Fiji Islands; 8 March 1928 (H. S. Simonds).

Type: (BMNH).

—

Ficure 88.—Scenopinus fijianus (Krober), female: a, wing (type); }, c, lateral and frontal
aspects of head (type); d,¢, lateral and frontal aspects of specimen agreeing with Krober
figures; f, detail of antenna (after Krober); g, detail of antenna of specimen agreeing
with type, (abdomen of type lost); h, ventral aspect of 8th sternum; 1, lateral aspect
of 8th and 9th segments; 7, 9th sternum and bursa.

132 U.S. NATIONAL MUSEUM BULLETIN 277

91. Scenopinus flandersi, new species
Figure 89
Ma.z.—Head black-brown; eyes red-brown above, dark red-brown

below; frons subshining, extending narrowly from antennal base
between silvery pile to broader area, then narrowing to a thin line

Figure 89.—Scenopinus flandersi, new species, male, female: a, wing; 6, c, lateral and
frontal aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 133

halfway to the prominent ocellar tubercle, ocelli orange-brown; post-
gena narrow below and behind the eyes; mouthparts brown, well de-
veloped, filling the oral cavity; palpi brown, nearly as long as the
rostrum; oral cavity bordered by silvery pile which extends onto the
frons well above the antennal base. Antennae dark red-brown, first
segment short, second twice as large as the first, third approximately
twice as long as broad, narrowing distally; see figure.

Thorax dorsum black with a patch of silvery pile above the humeral
callus; tips of humeral callus and the supra-allar calli orange; pleural
areas red-brown. Wings smoky hyaline, veins tan, fading before end
of the wing; halter stem black-brown, knob creamy white. Legs with
femora and tibiae red-brown, tarsi straw yellow.

Abdomen dark red-brown, segments 3-5 with medium broad white
bands along posterior edge. For details of terminalia, see figure.

Frema.e.—Head dark red-brown, eyes red-brown, with a narrow
postocular ridge; frons rugose, subshining, swollen above the antennae
and depressed medially just above swelling, extending as a slight
median depression or line to median ocellus; ocellar tubercle not
prominent, ocelli red-brown; oral cavity bordered by a silvery pile
which extends above the antennae; mouthparts large, well developed;
palpi long, genal area small. Antennae as in the male.

Thorax dorsum black-brown, with a silvery pilose area above the
humeral callus which is red-brown; supra-alar callus tan. Pleural areas
red-brown. Wings hyaline, brownish, veins brown; halter stem brown,
knob red-brown. Legs with femora and tibiae red-brown, tarsi straw
yellow.

Abdomen red-brown; see figure for details of eighth segment and
bursa.

Length: Male body 3 mm., wing 2 mm.; female body 3.4—4 mm.,
wing 2.2-2.4 mm.

Type-locality: Redlands, California; 1 May 1953 (S. Flanders),
ex elm branch.

Holotype: Male (CAS) 8916 (ex [PHT)).

Allotype: Female same data (CAS).

Paratypes: 1 o&, 12, same data as type (USNM); 2 99, same
data as type; 1 o, Sierra Madre, Monterey Co., California, 20
October 1923 (Armitage) (PHT); 1 &@, Long Beach, California,
14 August 1949 (A. L. Melander) (USNM).

92. Scenopinus gossypius, new species

Figure 90

This species is widely distributed across the southern United States,
from the Atlantic to the Pacific. Information appended to the speci-

134 U.S. NATIONAL MUSEUM BULLETIN 277

mens indicates that is has been swept from cotton and reared from
cotton squares; it may be a predator of the boll weevil or boll worm.

Maue.—Head red-brown to black-brown; eyes red-brown above,
black-brown below; frons narrow, rugose, with a median groove
ending in a shallow pit at the upper extremity; ocellar triangle only
slightly elevated, ocelli orange; mouthparts well developed, light
brown, palpi thin, elongate; oral opening bordered with silvery pile.
First antennal segment short, brown; second segment larger, light
brown; third segment brown, slightly tapering, 2% times longer than
one and two combined; see figure.

Figure 90.—Scenopinus gossypius, new species, male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged
details of antennae; h-j, ventral, lateral and posterior aspects of male terminalia; k,
ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

Thorax dorsum, red-brown to black-brown, striated, a thin patch of
silvery pile above the humeral callus which has a thin straw colored
posterior margin. Wings smoky hyaline, veins brown, R5 and M1 + 2
constricting distally and then widening slightly at wing margin to
give a bottle shape to cell R5; halter with stem dark brown to brown,
knob red-brown to orange-brown above, light below. Legs with coxae
through the femora ground color, tibiae grading from orange-brown
anteriorly to red-brown on hind pair, tarsi straw colored darkening
distally.

Abdomen, black-brown to red-brown with two white bands on
segments 4 and 5, sparsely covered with posteriorly directed hairs.
See figure for details of terminalia; close to S. velutinus but differs in
shape of aedeagus, ninth tergum, and ejaculatory process.

SCENOPINIDAE OF THE WORLD 135

FrmMaLe.—Head, black-brown to red-brown; eyes black-brown with
a very narrow postocular ridge; frons rugose, with two swellings on
either side of mid-facial groove above antennae extending into a median
depression, above which a faint median groove extends to median
ocellus; ocellar triangle shallow but set off from frons, ocelli orange.

Thorax, wings, halteres, and legs as in male.

Abdomen, black-brown to red-brown, thinly covered with posteriorly
directed hairs. For details of eighth segment and bursa, see figure.

Length: Male body 3-3.5 mm., wing 2.3-2.5 mm.; female body
3.5-4.2 mm., wing 2.5-3 mm.

Type-locality: Dallas, Texas; 24 April 1907 (W. W. Yothers)

Holotype: Male (USNM) 67473.

Allotype: Female Dallas, Texas; 5 July 1909 (W. W. Yothers)
(USNM).

Paratype males: 10, Dallas, Texas, 24 April 1907 (W. W.
Yothers); 1%, Kerrville, Texas, 11 June 1953 (L. J. Bottimer),
49°, Los Angeles, California, 1928 (H. M. Armitage) (on window in
propagation room) (USNM); 10, Jacksonville, Florida (Mrs. A. T.
Slosson) (AMNH).

Paratype females: 46 specimens from the following localities:
Alachua Co., Florida; Gainesville, Florida; Monticello, Florida;
Quincy, Florida; Tifton, Georgia; Jekyll Island, Georgia; Clark Co.,
Georgia; Selma, Alabama; Florence, Alabama; Rivers, Tennessee;
West Point, Mississippi; Belzono, Mississippi; Utica, Mississippi;
Ecuador Parish, Louisiana; Galveston, Texas; Coma Island, Texas;
Brownsville, Texas; Austin, Texas; Hearney, Texas; Kerrville, Texas;
Calvert, Texas; Mesilla, New Mexico; Nogales, Arizona; Riverside,
California; Los Angeles, California; Olancha, California; Claremont,
California; all in (USNM). 2 99, Jacksonville, Florida (Mrs. A. T.
Slosson) (AMNH). 1 9, Clarke Co., Georgia, 17 September 1937
(Horace O. Lund) (UGa). 1 9, La Jolla, California, 13 July 1941
(E. L. Todd). 1 9, Green River Camp, lower Santa Ana Co., California,
11 May 1933 (E. P. VanDuzee) (CAS). 1 9, Compton, Los Angeles
Co., California, 31 July 1952 (E. Swift) (UCalD). 1 9, Riverside,
California (Timberlake) (PHT). 1 2, Guamuchil, Sinaloa, Mexico,
8 May 1963 (R. C. Bechtel, E. I. Schlinger) (UCalB). 1 2, Manning,
South Carolina, 1 April 1945 (C. Sabrosky) (CSC).

93. Scenopinus hagai, new species
FIGuRE 91

Ma.r.—Head black-brown; eyes red-brown above, black-brown
below; frons subshining, nearly divided above the antennae by the
dorsal extension of silvery pilosity surrounding the oral cavity, a

136 U.S. NATIONAL MUSEUM BULLETIN 277

shallow shining depression on upper third of frons; ocellar tubercle
prominent, ocelli clear; mouthparts brown, well developed; palpi
nearly as long as labellum. First antennal segment brown, short;
second segment orange-brown, twice as large as first; third segment
brown, elongate oval, twice as long as wide.

Thorax dorsum black-brown with a patch of silvery pile above
the orange humeral callus; scutellum and pleural areas red-brown.
Wings hyaline, veins yellow; halter stem brown, knob creamy to dark
brown above, white below. Legs red-brown through the femora,
tibiae light brown, tarsi straw yellow.

Ficure 91.—Scenopinus hagai, new species, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia
(ejaculatory syringe apodeme missing); &, ventral aspect of female 8th sternum; /, lateral
aspect of female 8th and 9th segments; m, female 9th sternum and bursa.

Abdomen dark red-brown, segments 3-5 with thin white posterior
bands which increase in width toward the tip of the abdomen. For
details of terminalia, see figure. Differs from S. gossypius in larger
ejaculatory mechanism, parameres, and shape of ninth tergum.

FrEmMALE.—Head black-brown; eyes red-brown with a narrow post-
ocular rim; frons broad, swollen above the antennae, a median groove
through the lower third; back of head and rim of post-ocular flange
pollinose; mouthparts red-brown, well developed as in the male;
silvery pile bordering oral opening, extending above the antennae
onto the frontal area; ocellar tubercle depressed, not prominent,
ocelli red-brown. Antennae with first segment short, black-brown;

SCENOPINIDAE OF THE WORLD BY

second segment larger, red-brown; third segment, black-brown,
pilose, elongate oval, over twice as long as first two segments combined.

Thorax dorsum black-brown, with a pilose patch above the humeral
sallus which is red-brown anteriorly with an orange posterior border;
supra-alar callus yellow-orange; scutellum red-brown; pleural areas
dark red-brown; wings brownish, veins brown; halter stem brown,
knob red-brown above, white below. Legs: femora red-brown,
tibiae brown, tarsi straw yellow.

Abdomen dark red-brown. See figure for details of eighth segment
and bursa.

Length: Male body 2.7 mm., wing 2.1 mm.; female body 3.5 mm.,
wing 2.3 mm.

Type-locality: Yuma, Arizona; 6 July 1953 (Haga) on cotton.

Holotype: Male (USNM) 68007.

Allotype: Female Yuma, Arizona, 21 July 1953 (Haga) (USNM).

94. Scenopinus howdeni, new species

FIGURE 92

Frema.e.—Head black; eyes black-brown, with a narrow subshining
postocular band; frons rugose, with a narrow shining band along the
eye margins, swollen above the antennae, with a median groove
extending to the median ocellus, swollen between the ocellar tubercle
and upper corner of the eye; ocellar triangle flattened, cut off from
frons; ocelli orange. Mouthparts well developed; oral opening sur-
rounded by a white pubescence which extends over antennal bases;
genae shining. Antennae black-brown, first segment small; second seg-
ment twice as large, subshining; third segment granular, over twice
as long as wide.

Thorax dorsum black-brown, striated, with a white pubescent
patch above the humeral callus, which is orange-brown; pleural
areas red-brown, subshining; wings hyaline, veins brown, cell R5
narrowing sharply at tip of wing; halter stem red-brown, knob tan.
Legs with foreleg brown through tibia, tarsus cream, darkening
distally; middle leg with coxa and femur red-brown, tibia brown,
tarsus cream, darkening distally; hind leg red-brown except tarsus
cream, darkening distally.

Abdomen red-brown. See figure for details of eighth segment and
bursa.

Mare.— Unknown.

Length: Female body 4.4 mm., wing 3 mm.

Type-locality: S.W. Res. Sta., Portal, Arizona; 26 June 1956
(H. and A. Howden).

Holotype: Female (CNC) 8958.

138 U.S. NATIONAL MUSEUM BULLETIN 277

b, c, lateral and frontal

Ficure 92.—Scenopinus howdeni, new species, female: a, wing;
aspects of head; d, enlarged detail of antenna; e¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 139

95. Scenopinus kuiterti (D. E. Hardy)
FIGURE 95
Omphrale kuiterti D. E. Hardy, 1944a, p. 46.

The male of this species is distinguished by the long aedeagal
parameres and the point on the ventral margin of the ninth tergum.
It can be distinguished from S. cochisei by the shape of the ninth
tergum and the longer antennae. The female was heretofore unknown
and the description follows.

FrmaLe.—Head black; eyes red-brown with a narrow postocular
rim behind the eyes; frons smooth, shining along eye margins, genae
and swelling over the antennal base; transversely rugose over most
of frontal area with a shallow median depression; ocellar triangle
flat, set off from frons; ocelli orange. Mouthparts brown, well de-
veloped, palpi black-brown, half the length of oral cavity; no silvery
pile along oral opening. First antennal segment black-brown, short;
second, red-brown; third, rec-brown, slim, over twice as long as
first two segments combined.

Ficure 93.—Scenopinus kuiterti (D. FE. Hardy), male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, lateral and posterior aspects of male
terminalia; &, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

Thorax dorsum black, rugose, humeral and supra-alar calli red-
brown; pleural areas dark red-brown. Wing brown fumose, veins
brown; halter stem brown, knob cream. Legs red-brown, tarsi cream.

313-168 O—69—10

140 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen red-brown. See figure for details of eighth segment and
bursa.

Length: Female body 2.3-2.7 mm., wing 1.8-2 mm.

Type-locality: Buckeye, Arizona; 16 July 1940 (L. C. Kuitert).

Type: (UKan).

Female described from Andreas Canyon, Riverside Co., California;
10 May 1954 (Timberlake) (CAS) no. 8917 (Ex [PHT)).

Other specimens seen: 1, Black Mountain, Arizona, 13 July
1932 (R. H. Beamer) (UKan); 1 9, Morongo Valley, San Bernardino
Co., California, 30 August 1956 (Timberlake) (PHT); 299, Las
Cruces, New Mexico, 9 and 20 July 1960 (J. G. Watts) (USNM);
10°, Mesilla, New Mexico, September 1896, from granary (Th.
D. A. Cockerell) (USNM); 107, El Paso, Texas, 13 May, (Cockerell)
(USNM); 107, Mesilla Park, New Mexico (C. N. Ainslie) Webster
no. 5095 (USNM); 1c’, San Diego Co., California, 23 May, (E. P.
Van Duzee) (CAS); 1 &, Mesa, Arizona, 9 June 1955 (G. D. Butler)
swept/alfalfa (UAriz); 1 o&, Kyrene, Arizona, 11 July 1955 (O. L.
Barnes) swept/alfalfa (UAriz); 1 &%, Casa Grande, Arizona, 20 June
1956 (G. D. Butler) swept/alfalfa (UAriz); 1 o&, Helendale, California,
19 April 1959 (Timberlake) (PHT).

96. Scenopinus magdalenai, new species

Ficgure 94

Ma.zr.—Head black; eyes red-brown above, dark red-brown below;
frons subshining, extending one-third the distance from the antenna
to the ocellar triangle, with a converging groove above the antennae
and a pit at the apex of the frontal triangle; ocellar triangle set low at
back of head; ocelli red-brown. Mouthparts black-brown, well devel-
oped; back of head concave, subshining; little or no genal area.
Antennae with first segment black, short; second segment short, red-
brown; third segment black-brown with short hairs, elongate, over
twice as long as wide.

Thorax dorsum black, with silvery pilose area above the humeral
callus; humeral and supra-alar calli red-brown; pleural area black-
brown. Wings hyaline, veins brown; halter stem brown, knob red-
brown; legs with all femora dark red-brown; tibiae and tarsi of first
two pairs orange-brown; hind legs with tibiae brown, tarsi yellow.

Abdomen black-brown, segment 3 with broad white posterior
margin, segments 4 and 5 with very narrow bands. See figure for
details of terminalia. Primary difference is in structure of ejaculatory
syringe and accessory structures.

FrmaLe.—Head black, eyes dark red-brown with a narrow post-
ocular ridge and well-developed genae behind and below the eye; frons

SCENOPINIDAE OF THE WORLD 14]

Ficure. 94.—Scenopinus magdalenai, new species, male, female: a, wing; b, c, lateral
and frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g,
enlarged details of male and female antennae; h-j, ventral, lateral and posterior aspects
of male terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female
8th and 9th segments; m, female 9th sternum and bursa.

granular, subshining with the portion above the antennae bulging,
split by a median groove extending to a median depression on lower
third of the frontal area; antennae as in male.

Thorax as in male except halter knob light red-brown.

Abdomen black-brown. See figure for details of eighth segment and
bursa.

142 U.S. NATIONAL MUSEUM BULLETIN 277

Length: Male body 3 mm., wing 2.2 mm.; female body 3 mm.,
wing 2.1 mm.

Type-locality: Maria Magdalena (island), Tres Marias, Mexico;
20 May 1925 (H. H. Keifer).

Holotype: Male (CAS) 8922 on loan from (UCalB).

Allotype: Female same data (CAS).

97. Scenopinus magnicornis (Kréber)
FIGURE 95
Omphrale magnicornis Kréber, 1913, p. 202.

The type of this species is much smaller than other material seen
by me and is distinguished by its large bulbous third antennal segment.
As yet no male has been associated with it. It is close to S. velutinus
but separable by the shape of the antenna and bursal plate.

Length: Female body 3.5 (type)—4.4 mm., wing 2.6 (type)—-3 mm.

Type-locality: Amerika (Costa Rica) (in Winthems Coll. as
pygmaeus Lw.).

Type: (NMY).

Other specimens seen: 1 9, San Jose, Costa Rica; 1000-1200 m.,
15 November 1931 (F. Nevermann) in house (IOC); 4 99, La Suiza
de Turrialba, Costa Rica (Pablo Schild) (USNM); 1 9, Lima, Peru,
11 May 1961 (N. L. H. Krauss) (USNM).

<->

Ficure 95.—Scenopinus magnicornis (Krober), female: a, wing; , c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum; f, lateral
aspect of 8th and 9th segments; g, 9th sternum and bursa.

SCENOPINIDAE OF THE WORLD 143

98. Scenopinus mirabilis Adams
FIGURE 96

Scenopinus mirabilis Adams, 1904, p. 445.
Scenopinus nubilipes Cresson, [not Say] 1907, p. 111.
Scenopinus ramaleyz James, 1938, p. 22.

Close study of the terminalia of a series of males distributed over
a wide geographical area indicates that individuals with the char-
acteristic shape of the ninth tergum, excavated dorsal posterior
margin, and notch at the ventral tip, are apparently undergoing
speciation. I place S. ramaleyi in synonomy with S. mirabilis on the
basis of the short aedeagus with weak parameres and ninth segment
claspers with a long median knob on tip flanked laterally by an
equally wide depressed portion. The character given by Hardy for
separation is much too variable to be relied upon.

Ficure 96.—Scenopinus mirabilis Adams, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

The third antennal segment is oval, being about twice as long
as wide, cell R5 open at tip of wing by at least the width of the r-m
cross vein. Size of cell R4 variable.

Length: Male body 2.3-2.5 mm., wing 2-2.3 mm.; female
body 3.4-4 mm., wing 2.5 mm.

Type-locality: Bill Williams Fork, Arizona (F. H. Snow).

Types: (UKan).

144 U.S. NATIONAL MUSEUM BULLETIN 277

I have identified specimens from the following localities: 3°,
29 9, Escondido, California, 15 July 1941 (E. L. Todd) (UKan);
1, Albuquerque, New Mexico, 5 July 1941 (Burt Hodgden) (UKan);
1, Adelaide, Idaho, 21 July 1927 (USNM); 1c’, 12, Halsey,
Nebraska, 1 July 1957 (R. Henzlik) (USNM); 1<, Tucson, Arizona,
4 May 1942 (A. L. Melander) (USNM); 107, Calabasas, 6 mi. w.
Nogales, Arizona, 1 August 1961 (Werner, Nutting) (UAriz); 10,
Helendale, California, 21 May 1955 (W. R. M. Mason) (CNC);
1 o&,2 2 9, Medicine Hat, Alberta, 15 July 1956 (O. Peck) (CNC);
2 oo’, 2 9 9, Alamagordo, New Mexico, 25 April-15 May 1902,
(ANS).

99. Scenopinus nubilipes Say
FIGURE 97
Scenopinus nubilipes Say, 1829, p. 170.

The type of this specimen has been lost for many years and, as a
result, most American collections contain numerous specimens
identified as this species. My original aim was to study the pupal

Figure 97.—Scenopinus nubilipes Say, male, female: a, wing; 6, d, lateral and frontal
aspects of male head; c, e, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-j, ventral, lateral and posterior aspects of male ter-
minalia; &, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

skins of North American Scenopidae. This necessitated a more
detailed study of the genital structures where differences were more
obvious. This, in turn, necessitated descriptions of new species and
resulted in this paper. After due consideration of Say’s description

SCENOPINIDAE OF THE WORLD 145

and noting the type-locality, I have selected the following specimen
as the neotype of the species.

Matr.—Head black, eyes red-brown above, black-brown below;
frons small, triangular, extending less than half the distance from
the antennal base to the ocellar triangle, rugose with a median
groove entering a pit at the dorsal point; back of head concave;
mouthparts brown, well developed, filling oral cavity; palpi shorter;
ocellar tubercle prominent, ocelli orange-red. Antennae black-brown,
first segment short; second segment with an orange distal rim; third
segment elongate with a lateral distal pit; see figure.

Thorax dorsum black with transverse striations and a few sparse
hairs; humeral and supra-alar calli orange; pleural areas dark red-
brown. Wings dusty brown, veins brown; halter stem brown, knob
red-brown to black-brown, polished. Legs with femora black-brown,
tibiae red-brown, tarsi orange anteriorly to straw colored posteriorly.

Abdomen red-brown, segments 3 and 4 with broad white stripes.
For details of ninth tergum and accessory organs, see figure.

FremMALE.—Head black, eyes black-brown, with a narrow post-
ocular rim; frons broad, under one-third the width of the head,
slightly swollen, rugose with a median depression from the antenna
to the ocellar triangle, a pit in its lower third; mouthparts well
developed; back of head concave; ocellar tubercle flat, set off; ocelli
orange-red. Antennae as in male.

Thorax as in male.

Abdomen red-brown. For details of eighth segment and bursa
see figure.

Length: Male body 3.7 mm., wings 2.7 mm.; female body 4.3-
4.5 mm., wing 3 mm.

Neotype locality: Urbana, Illinois; 15 March 1959 (G. T. Riegel).

Neotype: Male (INHS).

Paratypes: 2 oo, 2 99, Urbana, Illinois, 15 March 1959
(G. T. Riegel) reared from granary trash, all with pupal skins, in
(USNM) and (INHS); 2 707, 399, Lafayette, Indiana, 24 June
1957 (L. Chandler) (PU); 1 o, Spencer Co., Indiana, 10 April 1953
(Montgomery) bean flowers (USNM); 1 @, 1 2, Lawrence, Kansas,
15 May 1896, 13 June 1898 (H. Kahl) (CNC); 1 9, Toloma, Illinois,
10 May 1927 (CAS).

100. Scenopinus pecki, new species

FIGURE 98

Ma.er.—Head black, eyes brown above, black below; frons slightly
swollen, shining, with punctures on either side next to the eye;
mouthparts brown, well developed; oral opening bordered by silvery
pile that extends to above the antennal bases; ocellar tubercle shin-

146 U.S. NATIONAL MUSEUM BULLETIN 277

ing, ocelli reddish; first two antennal segments black-brown, the
first smaller than the second, the third segment black, oval, twice
as long as the first two combined.

Thorax dorsum black, subshining; humeral and supra-alar calli
brown; wings smoky, veins brown to translucent; halter stem tan,
knob white; legs with femora and tibiae black, tarsi light brown.

Abdomen black, shining, segments 3 and 4 white above, black
below, ninth segment black, normally hairy. See figure for details
of ninth tergum and aedeagal structure.

Ficure 98.—Scenopinus pecki, new species, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, frontal and lateral aspects of female head; f, g, enlarged details
of male and female antennae; h-7, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

FrmaLn.—Head black, shining; eyes red-brown, with a narrow
postocular ridge; frons subshining with transverse striations, with
a slight swelling over base of antennae; ocellar tubercle set off, not
prominent; ocelli red; mouthparts normal; oral opening surrounded
by silvery pile. Antennae as in the male.

Thorax as in male except halter stem black, knob light brown
above, white below.

Abdomen black-brown; see figure for details of eighth segment and
bursa.

Length: Male body 2.3-2.8 mm., wing 2 mm.; female body
3.3 mm., wing 1.8 mm.

Type-locality: Scandia, Alberta, Canada; 10 July 1956 (O. Peck)
swept from range grass.

Holotype: Male (CNC) 8957.

SCENOPINIDAE OF THE WORLD 147

Allotype: Female, same data as type.

Paratypes: 12 oo", same data as type; 2 oo’, Lethbridge, Alberta,
14 July 1956 (O. Peck) (CNC); 3 #@o, 1 9, Thermopolis, Wyoming,
26 July 1960 (R. J. Lavigne) (USNM).

101. Scenopinus pygmaeus Loew
Figure 99

Scenopinus pygmaeus Loew, 1857, p. 90.
Scenopinus albidipennis Loew, 1869, p. 32.

This species, first described from Surinam, appears to be distributed
over the Caribbean region. Examination of the genitalia of the type
of albidipennis showed it to be identical with that of S. pygmaeus.
The type of S. albidipennis is somewhat teneral which may have
led Loew to describe it as a new species.

Ficure 99.—Scenopinus pygmaeus Loew, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, e, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-J, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

A redescription of the male follows with the description of the
hitherto unknown female.

Matr.—Head black, eyes red-violet above, blackish below; frons
small, subshining, with shallow pit at the upper two-thirds of frons;
the pile surrounding the oral opening extending above the antennae
and nearly reaching the midline from either side; ocellar tubercle
prominent, ocelli red. Antennae are red-brown; first segment short;
second, twice size of first; third elongate, over twice as long as the
first two combined, velvety.

148 U.S. NATIONAL MUSEUM BULLETIN 277

Thorax dorsum black, rugose, only a few very fine hairs along lateral
margin of tergum; humeral callus with a small posterior patch of
yellow-brown; pleural areas black-brown; wings hyaline to milky,
veins pale tan, faint; halter stem black, knob white. Legs with femora
red-brown, tibiae orange-brown, tarsi straw colored darkening
posteriorly.

Abdomen dark red-brown, shining, segments 4 and 5 with narrow
white posterior bands, all segments with short reddish hairs. See figure
for details of ninth tergum and genital structures.

FrmaLe.—Head black, eyes black-brown, with a narrow postocular
ridge; frons swollen above antennae, with swelling divided by a deep
median groove ending in a pit in the lower third of the frontal area,
remainder of frons rugose; ocellar tubercle flat, rugose; median ocellus
sunken, ocelli red-brown; mouthparts brown, well developed; palpi
black-brown, long; oral opening bordered by silvery pile which extends
over the antennal bases below the swellings on the frons. Antennae as
in male except third segment thickened at base, velvety.

Thorax dorsum black, rugose, a patch of white pubescence anterior
to the humeral callus, a few fine hairs on lateral margin; pleural areas
dark red-brown; wings brownish hyaline, veins brown; halter stem
red-brown, knob white to creamy. Legs with femora red-brown, tibiae
red-brown lightening posteriorly, tarsi light red-brown anteriorly
lightening to tan posteriorly.

Abdomen red-brown; see figure for details of the eighth segment
and bursa.

Length: Male body 2.3-2.8 mm., wing 1.7-2.3 mm.; female body
3-3.5 mm., wing 2.3-2.6 mm.

Type-locality: Surinam.

Type: (ZMH).

Material examined: 1, Cuba (Gundlach) as type of S. albidi-
pennis Loew (MCZ) no. 1086; 1, Mina Carlata, Trinidad Mt., Cuba,
11-16 July 1939 (C. T. Parson) in (MCZ); 107, 3 99, Mazaruni Clearing,
British Guiana, 9 July, 15 and 21 September 1937, in colony house
(Richards and Smart) (BMNH); 10, 19, La Suiza de Turrialba,
Costa Rica (Pablo Schild) Higuito, San Mateo, Costa Rica, (Pablo
Schild) (USNM); 77, 15 99, Palisades, Kingston, Jamaica, No-
vember 1957, February and September 1958, July and August 1964
(T. H. Farr) (SMIJ).

102. Scenopinus rossi, new species
Figure 100
Maure.—Head black-brown; eyes dark brown above, red-brown
below; frons with lower half smooth shining, bordered on each side by
the dorsal extension of lateral silvery pubescence surrounding the oral

SCENOPINIDAE OF THE WORLD 149

cavity, upper half of frons with an elongate median depression; ocellar
tubercle prominent, ocelli red-orange; mouthparts brown, well devel-
oped, extending the length of the oral cavity; palpi nearly as long as
rostrum; a narrow gena on lower part of the head. Antennae black-
brown, first segment small; second segment twice as large as first;
third segment twice as long as wide, slightly tapering, covered with
short golden hairs.

Thorax dorsum black, with a patch of silvery pile above the humeral
callus which has a red-brown posterior margin, supra-alar callus red-
brown; pleural areas black-brown. Wings smoky hyaline, veins light
brown anteriorly fading posteriorly; halter stem brown, knob red-
brown. Legs red-brown except tarsi straw yellow, fore tibia a little
lighter than others.

Ficure 100.—Scenopinus rossi, new species, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; A-j, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa; n, sketch showing central peg in membranous portion
of bursa.

Abdomen red-brown, segments 3-5 with narrow white posterior
bands. For details of ninth tergum and genital structure, see figure.

Fremate.—Head black-brown; eyes red-brown with a narrow
postocular band; frons broad, slightly rugose, subshining, swollen
above the antennae and with a broad median depression on the lower
fourth; ocellar tubercle broad and not prominent, median ocellus
sunken, ocelli whitish to orange. Mouthparts and antennae as in the
male.

150 U.S. NATIONAL MUSEUM BULLETIN 2:77

Thorax, wings, halter, and legs as in male.

Abdomen red-brown, eighth segment red-brown, sparsely covered
with hairs. See figure for details of sternum and bursa.

Length: Male body 2.7 mm., wing 2.1 mm.; female body 3.2 mm.,
wing 2.2 mm.

Type-locality: 6 mi. n. Victoria, Tampico, Mexico; 17 November
1946 (M.S. Ross).

Holotype: Male (CAS) 8923.

Allotype: Female, Revolution, 9 mi. s.e. of Buena Vista, Chiapas,
Mexico; 23 March 1953 (R. E. Bechtel, E. I. Schlinger) in (CAS).

Paratypes: 19, same data as allotype (UCalB); 1 6, 2 mi. s.e.,
of Revolution, Chiapas, Mexico, 22 March 1953 (R. E. Bechtel,
E. I. Schlinger) (UCalB).

103. Scenopinus schulzi Enderlein

Figure 101

Scenopinus schulzi Enderlein, 1934, p. 429.
Omphrale caenofrons Krober, 1937, p. 229.

This very common South American species was described from the
female; now the male has been found and is described below with a
redescription of the female.

Maur.—Head red-brown; eyes red-brown, darker below; frons
narrow triangular, striated; with white pilose bands laterad of oral
opening approaching above the antennal bases, area between antennal
bases and pilose bands smooth shining; ocellar tubercle red-brown,
prominent, ocelli orange; mouthparts well developed, orange-brown;
palpi nearly as long as rostrum, orange basally, black-brown distally.
Antennae red-brown; first segment short; second twice as large as
first; third elongate, three times as long as first two combined, darker
at tips.

Thorax dorsum black with a silvery pilose area above humeral
callus which is red-brown, as is the supra-alar callus; pleural areas
black-brown, shining. Wings hyaline, veins brown; halter stem brown,
knob mahogony brown; fore and middle legs orange-brown, hind
leg with coxa and femur red-brown, remainder orange.

Abdomen dark red-brown, segments 3 and 4 with moderately wide
white posterior margins, segment 5 with a very narrow white margin,
terminalia black, cleft so there is a wide dorsal and a narrow ventral
piece; see figure for details.

FrmaLe.—Head black; eyes reddish black, with a very narrow
postocular ridge; frons broader than the ocellar triangle, with a
shallow pit on the lower portion, subshining, with transverse rugose
striations below the pit, red-brown area anterior to the center of the

SCENOPINIDAE OF THE WORLD 151

pit on swelling above the antennae. Ocellar tubercle prominent, set
off from frons, ocelli yellow-orange. First antennal segment orange-
brown, very short; second segment twice size of first, orange; third
segment three times as long as broad, orange with a black tip.

Thorax dorsum black; humeral callus with a yellow spot, supra-
alar callus cream; pleura red-black. Wings brown fumose, veins
brown; halter stem brown, knob, dorsal surface red-brown, ventral
surface brown.

Ficure 101.—Scenopinus schulzi Enderlein, male, female: a, wing; b, lateral aspect of
male head; c, d, lateral and frontal aspects of female head; ¢, f, enlarged details of male
and female antennae; g-i, ventral, lateral and posterior aspects of male terminalia; J,
lateral aspect of female 8th and 9th segments; &, ventral aspect of female 8th sternum;
1, female 9th sternum and bursa.

Abdomen red-brown, eighth segment with concave posterior margin
on tergum, sternum with a raised bump before the end. See figure
for details of eighth segment and bursal plate.

Length: Male body 3-4 mm., wing 2.3-2.8 mm.; female body
2.5-2.75 mm., wing 2-2.25 mm. (actual range body 2.5-5.3 mm.,
wing 2—3.5 mm.).

Type-locality: Para, Brazil; 1 and 7 December 1893 (Albert
Schulz).

Type: (ZMH).

Other specimens seen: 2077, 399, Maracaju, Mato Grosso, Brazil,
February 1937 (USNM); Varneys, Saint Helena, 22 specimens cap-

152 U.S. NATIONAL MUSEUM BULLETIN 277

tured between 11 April and 26 November 1958, including a pair in
copula (A. Loveridge) (BMNH); 20°07, 19, Nova Teutonia, Brazil,
27°11’ S. 52°23’ W., 28 October and 3 November 1936 (Fritz Plau-
mann) (BMNH); 99°, Nova Teutonia, Brazil, November 1944, March
1946, November 1949, October 1956, October 1957, September and
November 1958 (Plaumann) (CNC); 107, Boa Vista Fundode Figueiras,
Cape Verde Islands, 30 January 1954 (University Exped.) in (ZMUBH).

104. Scenopinus seftoni, new species
Figure 102

This is another species similar to S. mirabilis but wing tip with
cell R5 more constricted, halteres orange-yellow to tan. Aedeagus
stronger and parameres weaker, ninth segment claspers pointed. Fe-
male eighth tergum and sternum excavated behind.

Ficure 102.—Scenopinus sefioni, new species, male, female: a, wing; ), c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, enlarged detail
of male antenna; g-1, ventral, lateral and posterior aspects of male terminalia; 7, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa; m, lateral sketch of bursa and central peg.

Ma.r.—Head black, eyes red-brown above, black-brown below,
frons subshining, swollen below, a median groove in upper half;
ocellar tubercle prominent, shining; ocelli red-brown; mouthparts
brown, moderately large, palpi half the length of oral opening which
is fringed by black-brown pollen; lower gena strongly developed,
shining. Antennae black-brown, first segment short, second larger,

SCENOPINIDAE OF THE WORLD Pas

third segment pubescent, oval, twice as long as wide and truncated
on the end.

Thorax dorsum black with a few sparse hairs, humeral callus red-
brown; pleural area dark red-brown; wing milky hyaline, veins tan;
cell R5 sharply narrowed at end (see figure), distance from r-m_ to
m cross vein twice as long as r-m; halter stem red-brown, knob tan
to cream yellow. Legs: femora and tibiae dark red-brown with yellow-
brown knee, tarsi straw yellow, darkening distally.

Abdomen dark red-brown, segments 3-5 with broad white stripes;
segments with a few lateral white hairs; ninth tergite black-brown,
cup of ejaculatory syringe blackish, aedeagus with a strong penis
flanked by weak parameres, ninth segment claspers pointed.

FremMALre.—Head black; eyes red-brown with a narrow postocular
rim; frons broad, slightly swollen above the antennal bases, rugose,
with shallow grooves either side of median ridge; ocellar tubercle
shallow, set off; ocelli red-brown. Mouthparts brown, as in male;
oral opening bordered by brown pile as in male. Antennae as in male.

Thorax dorsum and pleurum as in male; wings hyaline tinged
brownish, veins brown; halter stem red-brown, knob orange-yellow.
Legs with femora and tibiae dark red-brown, knees yellow, tarsi
orange-yellow, darkening distally.

Abdomen dark red-brown, eighth segment with posterior margins
of both sternum and tergum excavated; see figure for details of
eighth segment and bursa.

Length: Male body 2-2.5 mm., wing 1.5-2 mm.; female body 2.8
mm., wing 2.2 mm.

Type-locality: Espirutu Santo (island), Gulf of California, Mexico
(24°30’ N. 110°40’ W.); 23 March 1953 (P. H. Arnaud) Sefton Orca
Exped. to Gulf of California.

Holotype: Male (CAS) 8928.

Allotype: Female Isla Partida, Gulf of California, Mexico (24°30’
N. 110°40’ W.); 23 March 1953 (P. H. Arnaud) Sefton Orca Exped.
to Gulf of California (CAS).

Paratypes: 2 oo same data as type (CAS).

105. Scenopinus serratus, new species
FicurE 103

Ma.r.—Head black-brown; eyes amber brown above, brown below ;
frons small, triangular, extending over halfway to the median ocellus,
with a median groove below ending in a central pit, ocellar tubercle
red-brown, prominent, set on back top of head, ocelli clear; mouth-
parts creamy, palpi red-brown; lower part of gena narrowly developed.
First segment of antenna small, red-brown; second segment orange-

154 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 103.—Scenopinus serratus, new species, male: a, wing; 6, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects
of terminalia, missing portion of ejaculatory syringe outlined in dashed lines.

brown, twice as large as the first; third segment orange-brown dusted
with white, elongate oval, twice as long as wide.

Thorax dorsum black with a patch of silvery pile above the humeral
callus which is red-brown with an orange tip; pleural areas dark red-
brown. Wings hyaline, veins tan; halter stem tan, knob tan-brown

SCENOPINIDAE OF THE WORLD 155

above, white below. Legs with femora light red-brown, tibiae orange-
brown, tarsi straw yellow.

Abdomen with second segment black-brown, velvety; segments 3-5
with medium wide posterior white bands, remainder of abdomen
black-brown; ninth tergum with serrated edges; see figure for details.

FrmaLe.— Unknown.

Length: Male body 2.7 mm., wing 1.9 mm.

Type-locality: Casa Grande, Arizona; 20 June 1956 (G. D. Butler)
swept alfalfa.

Holotype: (USNM) 67477.

106. Scenopinus valgus (D. E. Hardy)
Figure 104

Omphrale valga D. E. Hardy, 1944a, p. 50.

None of the types or paratypes of this species has been seen, but
specimens which agree with Hardy’s figures have been found and are
illustrated. A series of three males and two females from Friday
Harbor, Washington, was found and is figured. The female is described
for the first time.

Mate.—Abdomen with segments 3-5 with every narrow posterior
white borders. Tergum of ninth segment as figured.

Ficure 104.—Scenopinus valgus (D. E. Hardy), male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, e, lateral and frontal aspects of female head; f, enlarged
detail of antenna; g-1, ventral, lateral and posterior aspects of male terminalia; 7, ventral
aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments; /, female
9th sternum and bursa.

313-168 O—69 11

156 U.S. NATIONAL MUSEUM BULLETIN 277

FrmauLe.—Head red-brown; eyes dark red-brown with a narrow
postocular rim; frons rugose, swollen above antennae, with shallow
ventral groove ending in a pit on the lower fourth of the frontal area;
ocellar tubercle indented, ocelli orange; oral opening bordered by
silvery pile ending squarely at base of swelling on front.

Thorax dorsum dark red-brown; humeral callus orange-tipped;
pleural areas red-brown; wing hyaline, veins brown anteriorly, faint
behind. Halter stem brown, knob red-brown. Legs red-brown with
tarsi yellow-brown basally, darkening distally.

Abdomen red-brown, shining. For details of eighth segment and
bursa, see figure.

Length: Female body 3 mm., wing 2.3 mm.; male body 2.6 mm.,
wing 2.1 mm.

Type-locality: Yosemite National Park, California; 1 August
1940 (R. H. Beamer).

Type: (UKan).

Specimens seen: 10, Clayton, Shasta Co., California, 15 July
1918 (EK. P. Van Duzee) (CAS); 20° 0°, 32 9, Friday Harbor, Wash-
ington, 25 July 1928 (M. Shackleford) (AMNH); 19, Victorville,
California, 30 May 1944 (A. L. Melander) (USNM); 19, s. fork
Santa Ana River, California, 30 July 1942 (A. L. Melander) (USNM).

107. Scenopinus vanduzeei, new species
Figure 105
Scenopinus nubilipes Cole (not Say), 19238, p. 462.

This species is closely allied to S. mirabilis and can be distinguished
by the shorter truncate antennae, the free portion of R5 being much
longer than R4, the portion of M between the r-m and m cross veins
longer, and the aedeagus which has a strong median lobe flanked by
shorter parameres; cup of ejaculatory syringe blackish.

Mauzr.—Head dark red-brown; eyes red-brown above, dark red-
brown below; frons narrow, triangular, without a median groove;
ocellar tubercle prominent, ocelli clear to red-brown. Antenna with
first two segments red-brown; first segment half as large as second;
third segment dark red-brown, less than twice as long as wide, flat-
tened, a dimple in the center of the tip.

Thorax dorsum black; humeral callus red-brown, pleura dark red-
brown. Wings milky white, veins light brown, halter stem red-brown,
knob white. Legs: foreleg with femur red-brown, tibia with distal
third orange-brown, tarsi orange; middle leg with femur and tibia
red-brown, tarsus orange-brown; hind leg with femur and tibia red-
brown, tarsus straw yellow darkening distally.

Abdomen dark red-brown, segments 3-5 with broad white bands.
' See figure for details of ninth segment and aedeagus.

SCENOPINIDAE OF THE WORLD 15%

FrmaLe.—Head dark red-brown; eyes red-brown with only a narrow
postocular rim; frons rugose, slightly swollen above the antennae,
no median groove; ocellar tubercle depressed, not cut off from frons,
ocelli orange. Mouthparts brown, not large; palpi short; pile around
oral opening brown, short. Antennae as in male.

Thorax dark red-brown; humeral callus red-brown. Wings smoky
brown, veins brown; halter stem red-brown, knob white. Legs with
femora and tibiae red-brown, tarsi light red-brown.

Abdomen red-brown. See figure for details of eighth segment and
bursa.

Ficure 105.—Scenopinus vanduzeei, new species, male, female: a, wing; }, c, lateral and
frontal aspects of male head; d, e¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; A-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of female 8th sternum; /, lateral aspect of female 8th and 9th
segments; m, female 9th sternum and bursa.

Length: Male body 2.3 mm., wing 1.8 mm.; female body 3.3 mm.,
wing 1.8 mm.

Type-locality: Mejia (island), Gulf of California, north of Angel
de la Guarda, 30 April 1921 (E. P. Van Duzee) (29°30’ N. 113°40’ W.).

Holotype: Male (CAS) 8929.

Allotype: Female, same data (CAS).

108. Scenopinus velutinus (Kréber)
Fiaure 106
Omphrale velutina Krober, 1913, p. 203.
Omphrale nubilipes Kroéber (not Say), 1928a, p. 2.
The type of this species has been lost. There is, however, a paratype
male in the U.S. National Museum and a long series of females from
the type-locality which have been used to figure this species.

158 U.S. NATIONAL MUSEUM BULLETIN 277

Length: Male body 3.5 mm., wing 2.5 mm.; female body 3.5-4
mm., wing 2.5-2.7 mm.

Type-locality: Costa Rica.

Type: Coll. Krober (lost).

Specimens seen: 21 99, Higuito, San Mateo, Costa Rica (Pablo
Schild) (USNM); 1 9, 2 mi. n.e. Guadelupe Atodia, Chiapas, Mexico,
24 March 1953, (R. C. Baker, E. I. Schlinger) (UCalB); 1 2, Constant
Spring, Saint Andrew, Jamaica, W.I., 23 August 1964 (T. H. Farr)
(SMIJ).

Ficure 106.—Scenopinus velutinus (Kréber), male, female: a, wing; 5, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, enlarged detail of
antenna; g—1, ventral, lateral and posterior aspects of male terminalia, ejaculatory mecha-
nism lost on dissection; 7, ventral aspect of female 8th sternum; &, lateral aspect of female
8th and 9th segments; /, female 9th sternum and bursa.

109. Scenopinus weemsi, new species

Figure 107

This small species appears to be closely related to S. mirabilis but
apparently has been isolated long enough to be recognized as a
distinct species.

Matzr.—Head black-brown; eyes black-brown above, black below ;
frons narrow triangular, slightly swollen in the middle, rugose but
subshining; ocellar tubercle black, prominent, ocelli orange; silvery
pile surrounding oral opening. First two antennal segments red-brown ;
first short; second about twice as large; third segment dark red-brown,
pear-shaped, wider at base, twice as long as wide, truncate at tip.

SCENOPINIDAE OF THE WORLD 159

Thorax dorsum black with a thin powder of white, humeral and
supra-alar calli red-brown to orange-brown; pleura red-brown to
black-brown, dusted with white in exposed areas. Wings milky white,
veins very faint, nearly white, R5 and M1+2 broadly separated,
M1+2 ending behind the tip of the wing; halter stem brown, knob
white. Legs with femora and tibiae dark red-brown, tarsi straw yellow.

Abdomen dark red-brown, segments 3-5 very broadly membranous
white, with thin sclerotized anterior bands. For details of ninth
segment, see figure.

Ficure 107.—Scenopinus weemsi, new species, male, female: a, wing; d, c, lateral and frontal
aspects of male head; d, e, frontal and lateral aspects of female head; f, enlarged lateral
aspect of female head and antenna; g, enlarged detail of male antenna; h-, ventral, lateral
and posterior aspects of male terminalia; k, ventral aspect of female 8th sternum; /, lateral
aspect of female 8th and 9th segments; m, female 9th sternum and bursa.

FrmaLe.—Head black-brown; eyes black-brown, with a very thin
postocular ridge fringed with hairs; back of head concave, covered
with a silvery pile, frons broad, rugose, slightly swollen, with a median
groove extending about halfway up the frontal area, lower fourth of
frons red-brown. Ocellar tubercle not set off from frons, ocelli orange.
Antennae as in male.

Thorax dorsum red-brown, otherwise as in male; pleura as in male,
halter stem brown, knob white with tan dorsal portion. Legs as in male.

160 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen red-brown, subshining. See figure for details of the eighth
segment and bursa.

Length: Male body 2—2.2 mm., wing 1.5 mm.; female body 2.8
mm., wing 1.7—2 mm.

Type-locality: Putnam Co., Florida; 6 July 1956 (H. V. Weems,
Jr.).

Holotype: Male (USNM) 67478.

Allotype: Female same data as type (USNM).

Paratypes: 3 o’o’, same data as type (USNM); 2 oa’, 1 Q,
Gainesville, Florida, 25 April 1952 (O. Peck) (CNC).

110. Scenopinus werneri, new species

FIGuRE 108

This species is of interest as it is one of the few recorded to have
been captured by ultra violet light traps.

Mauzr.—Head red-brown; eyes yellow-brown above, brown below
extending to but not around the back of the head; frons subshining,
swollen, extending nearly to ocellar triangle, with a shallow pit at the
midpoint. Ocellar tubercle black-brown, prominent, ocelli amber;
first antennal segment brown, short; second segment larger, yellow-
brown; third segment brown, pear-shaped.

Ficure 108.—Scenopinus werneri, new species, male: a, wing; b, c, lateral and frontal aspects
of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of
terminalia.

Thorax dorsum black-brown, subshining with faint transverse
striations; humeral callus tan; pleura red-brown; wings hyaline, veins
yellow-brown; halter stem brown, knob with base brown, remainder
white. Legs with fore and middle femora and tibiae orange-brown,
tarsi tan; hind leg darker red-brown, tarsus tan.

SCENOPINIDAE OF THE WORLD 161

Abdomen red-brown, segments 3 and 4 with white bands, remainder
red-brown. See figure for details of ninth segment.

FrmMALE.— Unknown.

Length: Male body 2 mm., wing 1.6 mm.

Type-locality: 8 mi. n. Vail, Pima Co., Arizona; 30 August 1962
(F. Werner and W. L. Nutting) u.v. trap.

Holotype: Male (USNM) 67481.

Paratype: Male same data (UAriz).

111. Scenopinus whittakeri (James)
FIGuRE 109

Omphrale whittakeri James, 1955, p. 47.
Dr. James was kind enough to submit two of his paratypes for
figuring. As James points out, this species is close to S. mirabilis

but the whole hind margin of the ninth tergum is more sloping and
the ventral extension is not prominent.

Bao DRA

Ficure 109.—Scenopinus whittakeri (James), male, female: a, wing; 4, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged details
of male and female antennae; h-, ventral, lateral and posterior aspects of male terminalia;
k, ventral aspect of female 8th sternum; J, lateral aspect of female 8th and 9th segments;
m, female 9th sternum and bursa.

Length: Male body 3 mm., wing 2.3 mm.; female body 3.5 mm.,
wing 2.5 mm.

Type-locality: Hanford, Benton Co., Washington; sagebrush, 640
ft., 6 May 1952, Ss8-5 (R. H. Whittaker).

Type: (WSU) 176.

162 U.S. NATIONAL MUSEUM BULLETIN 277

Allotype: Female, same data, 30 July 1952, Sc20-22.
Other specimen seen: 1 o’, Lowden, Washington; 22 June 1921
(A. L. Melander) (USNM).

Genus Caenoneura Krober
Caenoneura Krober, 1923, p. 75.

Type-species, Caenoneura robusta Kréber (monobasic). The genus
Caenoneura Krober is distinguished by the peculiar wing venation
which is crowded to the leading edge of the wing. Both members of
this genus come from the circum-Mediterranean area.

112. Caenoneura robusta Kroéber
Figure 110

Caenoneura robusta Kréber, 1923, p. 75.

This species is easily recognized by the peculiar venation of the
wing with all of the R veins and M1 + 2 ending in the leading edge
of the wing. The male has scalelike hairs on the frons and ocellar
triangle and five white bands on the abdomen.

Ficure 110.—Caenoneura robusta Krober, male: a, wing; b, c, lateral and frontal aspects of
head (note scale-like hairs); d, enlarged detail of antenna; e—g, ventral, lateral and posterior
aspects of terminalia.

SCENOPINIDAE OF THE WORLD 163

Length: Male body 3 mm., wing 2.4 mm.
Type-locality: Egypt.
Type: (SEE).

113. Caenoneura nigra, new species

Figure 111

Fremaute.—Head black; eyes black, with a narrow postocular
rim; frons broad, roughened, with a row of punctures along the
eye margin and a faint median groove. Mouthparts well developed,
black-brown; oral opening wider than long, fringed with silvery
pile. Ocellar tubercle prominent, ocelli reddish. First two antennal
segments black, short; third segment brown-black, over twice as
long as first two combined, truncate.

a
<\

OGY
b. Cc.

oH

Figure 111.—Caenoneura nigra, new species, female: a,
wing; b, c, lateral and frontal aspects of head; d, enlarged
detail of antenna.

Thorax black, rugose; humeral and supra-alar calli brown, scutellum
not sharply cut off from general tergal area; wings smoky brown,
veins dark brown, characteristic of genus. Halter stem black, knob
brown, round. Legs with femora and tibiae black, tarsi brown.

Abdomen black, broad, and rounded.

Mate.— Unknown.

Length: Female body 2.1 mm., wing 1.7 mm.

164 U.S. NATIONAL MUSEUM BULLETIN 277

Type-locality: Sierra Nevada Highway, Spain; 1600 m., 27 July
1960 (J. R. Vockeroth).
Holotype: Female (CNC) 8629.

Genus Pseudomphrale Kroéber
Pseudomphrale Krober, 1918, p. 180.

Typs-species, Scenopinus clausus Loew (monobasic). The genus
Pseudomphrale Krober is at present poorly defined as only one member
of the genus is available and the author was unable to obtain per-
mission to dissect the genitalia for study. The remaining species
have not been located so that the characters that could be used to
distinguish the members of this genus cannot be determined at
present. The only common character possessed by all the species
is the closed cell R5 and this character is common to ten other genera.
The members of this genus are found in the Middle East.

114. Pseudomphrale clausa (Loew)
Scenopinus clausus Loew, 1873, p. 150.

The type of this species could not be located.

Length: Male body 1.6 mm., wing 1.5 mm.

Type-locality: Samarkand; 19 June (Fedtschenko).
Type: Presumed lost as Kréber did not figure this species.

115. Pseudomphrale crenata (Becker)
Omphrale crenata Becker, 1913, p. 550.

The type of this species could not be found.

Length: Female body 3.5 mm., wing ?

Type-locality: vicinity of Tschachbar, Makran Coast of Indian
Ocean, Baluchistan; 12-16 March 1901 (Zarundy).

Type: ? Leningrad.

116. Pseudomphrale dubiosa (Becker)
Omphrale dubiosa, Becker, 1913, p. 551.

The type of this species could not be found.

Length: Male body 3 mm.

Type-locality: vicinity of Tschachbar, Makran Coast of the Indian
Ocean, Baluchistan; 12-16 March 1901 (Zarundy).

Type: ? Leningrad.

SCENOPINIDAE OF THE WORLD 165

117. Pseudomphrale longirostris (Becker)
Omphrale longirostris Becker, 1913, p. 549.

The type of this species could not be found.

Length: Female body 3.5 mm.

Type-locality: vicinity of Tschachbar, Makran Coast of the Indian
Ocean, Baluchistan; 12-16 March 1901 (Zarundy). Southern part of
the Mountain Cove Kuch-i-tuftan; 5 May 1901 (Zarundy).

Type: ? Leningrad.

It seems highly probable that one of the females belongs to P.
dubiosa and that only two species exist.

118. Pseudomphrale palaestinensis Kréber

Figure 112

Pseudomphrale palaestinensis Krober, 1937, p. 229.

The exact position of this species is in doubt at the present time
as I did not have permission to dissect the type specimen nor was
I able to compare it with other members of the group. The long
aedeagal parameres are reminiscent of North American Brevitrichia
but without a more detailed study of the ninth tergum and internal
structures little more can be said.

Length: Male body 3 mm., wing 2 mm.

Type-locality: Scopusberg, Jerusalem.

Type: (SMNS).

Genus Brevitrichia D. E. Hardy
Brevitrichia D. E. Hardy, 1944a, p. 32.

Type-species, Pseudatrichia griseola Coquillett (original designation).
The genus Brevitrichia Hardy is characterized by small flies with the
thorax covered with a fine granular dusting which is brown to gray.
Head higher than long and antennae generally pear-shaped and end-
ing in two points with a median sensory peg. Cell R5 of the wing
closed and vein R4 branched from the base of cell. In the male,
ninth tergum bilobed, open below and aedeagus in the form of two
elongate bristles. Female eighth segment elongate; sternum excavated
posteriorly. The members of this genus are found west of the Rocky
Mountains in the United States and Mexico.

Key to Species of Brevitrichia (males)

1. Wing hyaline . oe enn a2
Wing smoky white or brown . Bee eres ee ena. eae ete
Dae ELAlLel Men OU CAN ts 15 2 Mat cbr Cap bev “eR a ee he a ee,
Halter knobcream .............. . B. arnaudi, new species
3. Abdomen black-brown or red-brown. ... . Pa ne

Abdomen white. ........... .. .. .B. oculivirida, new species

166 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 112.— Pseudomphrale palaestinensis Krober, male: a, wing; 6, c, lateral and frontal
aspects of head; d, enlarged detail of antenna; ¢, lateral aspect of undissected terminalia.

10.

LL.

12.

13.

14.

15.

16.

Ti,

18.

19.

20.

SCENOPINIDAE OF THE WORLD 167

Abdominal segments black-brown with white posterior margins
B. halli, new species

Abdominal segments red-brown with creamy posterior margins
B. ordwayi, new species

Wine emiOlcys OrOWIs eehe! oly Sega) et wo ee Se ee we ee ee a eG
Wine sriakvwhites. 4-9 4c.4ted- 4 Le Src. o> Oe Se, gS ee oF
Halter knob tan... . . . . . . B. hodgdeni, new species
Halter knob tan with cream i wentral: surface. . . B. griffini, new species
TPGIUCY KOO WIC eich 68 ele wa: x Bs oe ewe Ee ea Oe Ss, SD
Halter knob cream ortan ...........4+++24-+-... 10
Abdomen pristine white. . .. . ee ee

Abdominal segments black with w hite Soetanies margins. B. insulana (Cole)
R4 branching beyond r-m cross vein equal length of r-m
B. addacifons, new species
R4 branching beyond r-m cross vein twice length of r-m
B. castanea, new species

Patter KnGptateed «oR 6... tw a ee oh ew le Ge ee Ae, OD
Halterknobcream. ...... 2.2.2.0 22 2 ee ee eee ees LD
Halter knob all tan. .. . ee ats ee lie
Halter knob tan above cream anal w, abdominal mens ae brown with

cream posterior margins. . .... . . . . . B. yueatani, new species
Abdomen white. .... . . . . . B. pruinosa, new species
Abdomen cream with Beiical rade ow n 1 Ds . . B. griseola (Coquillett)
Halter knob all cream... . ee ees eee
Halter knob cream, marked or fen below: or eo palldor. Gene ks
Abdomen marked with brown orcream ..... re ere
Abdomen white above. .. . 4 ay ee B. onan: new species
Abdomen white with narrow nafenes brown bands < tr47. 2 a 4 se 686
Abdomen red-brown. . . ws he em Sees ee ee
R4 branching beyond r-m apaat: fenseh of r-m. . . B. davisi, new species
R4 branching beyond r-m twice length of r-m. . . B. downeyi, new species

Abdomen with central red-brown ‘‘T’’s fading to white
B. coquilletti, new species
Abdomen with segments red-brown with posterior bands cream
B. beameri, new species

Halter knob white or yellow below .... . baal a a SOLS
Halter knob with tan margins. . ...... B. ‘ timberlakei, new species
Halter knob white below. ....... . . . . B. minuta, new species
Halter knob yellow below ... . ao ere cS grep. cee ee
Abdomen yellow with black basal paride. SAT ed B. helenae (James)

Abdominal segments brown with creamy margins. B. schlingeri, new species

Key to Species of Brevitrichia (females)

Wing hyaline .... ue ey ee ee ee ee
Wing smoky brown or wiilte Pr ee oe een a vec. oe ek
Halter knob tan, cream or orange .. . Aa te OR rs Ai) co a
Halter knob white, 7 spines on 8th fereum .. . . B. flocki, new species
Halter MOD GreaIyL s 4 w os 4s & acne Son ee) tee ee ee ae he

Halter knob tan or orange

Halter knob all cream :

Halter knob cream above tan bciow, 8th read witht 7 spines
B. downeyi, new species

or oO

168

10.

Tilt

12.

13.

14.

15.

16.

Alf

18.

19.

20.

21.

22.

U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen brown or orange .. . SOR STE Sh ota mine as oe eae tee
Abdomen brown grading to tan, ponecuonly CRSA at RoR with
Abdomen brown with white posterior bands; 8th eer Seth 7 spines
B. arnaudi, new species
Abdomen orange; 8th tergum with 7 spines and much hair
B. dicksoni, new species

Eighth tergum with 8 spines and a dorsal pit . . . B. boharti, new species
Eighth tergum with 6 spines . . .. . . . . . B.schlingeri, new species
Halter knob tan... . sieht Mang Cea oo eat
Halter knob orange, 8th fen aie an Ss spines . . . B. davisi, new species
Halter knob tan above cream below, 8th tergum with 8 spines and a ventral

pit . . . . . B. beameri, new species

Halter ienob eee fa sth feet sei 9 short stiff spines
B. halli, new species

iWin osm Oley: WiRTGE 8 pt ainph ae lam ecto ches mNrbenae! jad ie syedatn cain alle amu eecl aur NED
WAN RiSMmOlay DOW ss 9 ae iene ie BES Cee oer wale Tac eo minelel
Halter knobtan .. . ; AB ee at) seme
Halter knob cream, abdomen with saa Broun ups. 8th recone with 6 spines

andmuchhair. . . . os +) whe @ pi ee. Bcoquilletti, new species
Halter knob lighter belows SP LAte cai Palit ions:
Halter knob all tan, abdomen dark ede Braet "8th cae eit 1 dorsal

spine and7 pits. ............ . .. B. hodgdeni, new species
Halter knob cream below . . San a rae ca oA
Halter knob white below; 8th euenee vat 7 spines . B. ‘erilling new species
Abdomen brown; 8th tergum with 8 spines. . . B. melanderi, new species
Abdomen red-brown with white lateral patches grading to all red-brown; 8th

tergum with 4 spines. .... . .. . . B. yucatani, new species
Halter knob yellow-brown, cream or an Mesos Uw nual ey LO

Halter knob white, 8th tergum with 9 spines and a i earet tuft of hairs
B. insulana (Cole)
Halter knob cream or tan .. . oa ane A,
Halter knob yellow-brown, 8th cere arn eatnout spines aun th 8 pits
B. aspinosa, new species
Halter knob cream or tan .. . el Ce PLS
Halter knob tan; abdomen red- Broun Sth erent Ww ith By spines
B. ordwayi, new species
Halter knob entirely cream... . bel Ws Deere eet en a peanyal ee MS her O,
Halter knob cream with tan lateral bande, S toe She eo Lo
Abdomen brown with white lateral patches, 8th tora aa fee only
B. addacifons, new species
Abdomen with red-brown ‘‘T’’s cane to yellow-brown, 8th tergum with

6 spines. . . .. . . . B. timberlakei, new species
Abdomen brown eaiene cc jan or ea Ged-btown., ase so ora cae
Abdomen black and white or red-brown and white bende, poe es NaTs . 22

Abdomen brown grading to tan; 8th tergum with a fringe of hair oale
B. minuta, new species
Abdomen cream with red-brown central ‘‘T’’s grading to orange posteriorly;
8th tergum with 7 spines and very short hair. . . B. kerni, new species
Abdominal segments red-brown with white posterior bands; 8th tergum with
6G. Spiness 235% . . B. seitulaesca, new species
Abdominal Senta black mat white posterior bands; 8th tergum with
Si BPINESwsiha S18 SLAs Broa ee aac eae eninenla (Coquillett)

SCENOPINIDAE OF THE WORLD 169

119. Brevitrichia addacifons, new species
FicureE 113

Maue.—Head cream; eyes red-brown, metallic above, metallic
black-green below; frons small, triangular; ocellar tubercle brown,
pollinose; ocelli clear to red. Mouthparts well developed, haustellum
red-brown, remainder cream; antenna as figured; first segment brown,
second segment orange-brown, third segment brown.

Thorax dorsum black-brown with brown and tan pollinose scales,
lateral margins yellow; humeral and supra-alar calli cream; meso-
scutellum mostly black-brown and gray pollinose, lateral margins
yellow; pleural areas pollinose with yellow, red-brown and brown.
Wings smoky, veins tan, see figure; halter stem brown, knob cream
with a tan lateral band. Legs red-brown with pollinose scales.

Abdomen second segment with a broad anterior band of black
with a broad central posterior stem and narrow lateral extensions
like a “T,” remainder of posterolateral portion pristine white, remaining
segments white dorsally. Ninth tergum with base black, remainder
white pollinose. See figure for details of genital structures.

FrmaLe.—Head cream; eyes red-brown, metallic, with a moderately
broad postocular ridge without spots of color; frons broad, unmarked,
a few white hairs; ocellar triangle set off, red-brown, pollinose; ocelli
clear to red-brown; groove at upper angle of eye directed toward
back of ocellar triangle, fading, not reaching back of head, a central
notch in back of ocellar triangle. Antennae with first two segments
cream, third red-brown, shaped as shown in figure.

Thorax dorsum red-brown, covered with gray pollinose scales in
the center, with two lateral tan bands, area above humeral callus
and the lateral margins cream, area in front of scutellum orange
with two anteriorly directed arms; humeral and supra-alar calli
white. Mesoscutellum red-brown over middle half, lateral angles
cream pollinose. Pleura with prothorax white; anepisternum white,
with usual dark patch; katepisternum red-brown, with white dorsal
border. Wings hyaline, smoky, veins clear; halter stem brown, knob
white. Legs femora red-brown, tibiae and tarsi tan-brown.

Abdomen segments brown with white lateral patches. Eighth
segment yellow-brown, shaped as figured. Ninth tergum without
bristles, only long thin hair; see figure.

Length: Male body 2.7 mm., wing 2 mm.; female body 2.8 mm.,
wing 1.8 mm.

Type-locality: Antelope Springs, 8 mi. s.w. Deep Springs, Imperial
Co., California; 1 July 1961 (G. I. Stage).

Holotype: Male (UCalD) 75.

170 U.S. NATIONAL MUSEUM BULLETIN 277

Allotype: Female same locality, 29 June 1961 (J. Powell).
Paratype: 1 co’, same data as type (UCalD).

Ficure 113.—Brevitrichia addacifons, new species, male, female: a, male wing; b, female
wing; c, lateral aspect of male head; d, lateral aspect of female head; e, enlarged detail of
antenna; f, dorsal aspect of female head; g, h, ventral and lateral aspects of male terminalia;
i, ventral left and dorsal right aspects of female 8th and 9th segments; j, lateral aspect
of female 8th and 9th segments; &, bursal structure.

SCENOPINIDAE OF THE WORLD 171

120. Brevitrichia arnaudi, new species

FIGuRE 114

Maur.—Head brown; eyes black, with a green metallic glint
above and a red metallic glint below (brown in dried specimens) ;
frons tan, small, triangular; ocellar tubercle tan, pollinose; ocelli
orange. Antennae with all segments brown; see figure for details on
shape.

Thorax dorsum black-brown, covered with tan pollinose scales,
with a slight longitudinal striping; humeral and supra-alar calli
cream; mesoscutellum cream, with a brown patch occupying the
middle third. Pleura with prothorax cream; anepisternum cream,
katepisternum red-brown. Wings hyaline, veins tan; halter stem
cream, knob cream. Legs with femora light brown with a tan knee;
tibiae tan; tarsi tan, darkening distally.

Ficure 114.—Brevitrichia arnaudi, new species, male, female: a, wing; 6, lateral aspect of
male head; c, lateral aspect of female head; d, ¢, enlarged details of male and female
antennae; f, dorsal aspect of female head; g, h, ventral and lateral aspects of male termi-
nalia; i, ventral left and dorsal right aspects of female 8th and 9th segments, bursal
structure outlined; 7, lateral aspect of female 8th and 9th segments.

Abdomen segments brown with yellow-brown membranes along
posterior margins. Ninth tergum cream with a brown patch basally
which extends along the open margin; see figure for details.

FrmMaLye.—Head tan; eyes metallic green to brown in dried speci-
mens; postocular ridge broad, with a brown patch behind the eye;

313-168 O—69-—12

E72 U.S. NATIONAL MUSEUM BULLETIN 277

frons broad a little wider than the ocellar triangle, with a median
eroove running the entire length, a brown central band clothed with
a few sparse hairs, grooves from upper angle of eye extending to
back of head laterad of the midline. Ocellar tubercle red-brown,
with tan pollinose scales, ocelli orange. Antennal segments brown,
shaped as in figure.

Thorax dorsum red-brown ground, covered with longitudinal
red-brown pollinose bands; humeral and supra-alar calli orange to
cream, with a light area between. Pleural areas as in male; wings as
in male; legs as in male; halter cream.

Abdomen segments brown, with light membranous margins;
eighth segment orange-brown, ninth tergum with 8 stiff spines
sparse hairs, 10th tergum with a thick pencil of long hairs at base.

Length: Male body 2.5 mm., wing 1.8 mm.; female body 4 mm.,
wing 2.3 mm.

Type-locality: Los Frailes, Baja California, Mexico; 18 March
1953 (P. H. Arnaud) Sefton Orca Exped. to Gulf of California.

Holotype: Male (CAS) 8930.

Allotype: Female on same pin, sama data.

Paratypes: 1 o, Cabo San Lucas, Baja California, Mexico,
16 March 1953 (P. H. Arnaud); 2 2 @, Los Frailes, Baja California,
Mexico, 18 March 1953 (P. H. Arnaud) (CAS).

121. Brevitrichia aspinosa, new species

Figure 115

Frmate.—Head cream; eyes red-brown, probably metallic in
life; postocular ridge moderately wide, continuous behind ocellar
triangle, with a wide brown spot behind each eye; frons broad, with
a median groove bordered by bands of tan; sparse hairs over entire
frons; ocellar triangle red-brown, ocelli clear; grooves from upper
corner of eyes extending to lateral ocelli. Antennae with first two
segments brown; third segment orange-brown basally darkening to
brown distally; see figure.

Thorax tergum red-brown with lateral margins yellow, two yellow
streaks laterad of brown midline stripe posteriorly, all covered with
eray pollen; humeral and supra-alar calli yellow. Scutellum tan,
with a brown spot occupying the middle third. Pleura with prothorax
red-brown and cream, mesanepisternum cream with large red areas,
katepisternum red-brown with cream dorsal stripe, pollinose. Wings
smoky, veins brown; halter stem tan, knob yellow-brown. Legs with
femora red-brown, pollinose; knees yellow, tibiae and tarsi brown.

Abdomen red-brown with white membranous posterior margins.
Eighth segment red-brown, ninth tergite with no spines but 8 pits;
see figure.

SCENOPINIDAE OF THE WORLD 173

10)

47
eran
rN

Figure 115.—Brevitrichia aspinosa, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, dorsal aspect of head; e, ventral left and dorsal right
aspects of 8th and 9th segments, bursal structure outlined; f, lateral aspect of 8th and 9th
segments.

Mare.— Unknown.

Length: Female body 4.7 mm., wing 2.3 mm.

Type-locality: 7 mi. w. Seligman, Arizona; 1 July 1952 (R. H. &
L. D. Beamer, C. Weiner, A. Wolf, W. LaBerge, C. Liang).

Holotype: Female (UKan) 6291.

174 U.S. NATIONAL MUSEUM BULLETIN 277

122. Brevitrichia beameri, new species
FIGcurRE 116

Pseudatrichia griseola Cresson (not Coquillett), 1907, p. 113.

Mauy.—Head cream; eyes red-brown above, black-brown below;
frons tan, small, triangular; ocellar tubercle black-brown, covered
with gray pollen, ocelli red-brown. Antennae red-brown; see figure

for details.

Ficure 116.—Brevitrichia beameri, new species, male, female: a, male wing; b, female wing;
c, lateral aspect of female head; d, enlarged detail of male antenna; e, enlarged detail
of female antenna; f, dorsal aspect of female head; g, h, ventral and lateral aspects of male
terminalia; 7, ventral left and dorsal right aspects of female 8th and 9th segments; J,
lateral aspect of female 8th and 9th segments; &, bursal structure.

SCENOPINIDAE OF THE WORLD 175

Thorax dorsum black-brown, covered with gray pollen, hairs
sparse, short; humeral and supra-alar calli cream; scutellum broadly
red-brown, covered with gray pollen, lateral margins cream. Pleura
red-brown, with patches of yellow, covered with gray pollen; wing
smoky, veins tan; halter stem tan, knob creamy white; legs red-brown,
covered with gray pollen.

Abdomen red-brown, segments with narrow cream-colored mem-
branous posterior margins; ninth tergum distinctive, ending in points;
see figure for details.

FremMa.Le.—Head cream; eyes red-brown, with green metallic sheen;
frons broad with a median yellow-brown band covered by anteriorly
directed hairs; postocular ridge moderately broad, without a brown
patch, separated from frons by a shallow groove from the corner of
the eye to back of lateral ocellus. Ocellar tubercle red-brown, covered
with a gray pollen; antennae red-brown; see figure for details.

Thorax dorsum, pleura, and humeral callus as in male. Wing
hyaline, veins light brown; halter stem light brown, knob cream
below, tan above.

Abdomen red-brown, eighth segment red-brown, shining; ninth
tergum with 8 stiff spines and a ventral pit. See figure for details.

Length: Male body 2.5 mm., wing 1.7 mm.; female body 3.7 mm.,
wing 2.3 mm.

Type-locality: 25 mi. w. Tularosa, New Mexico; 1 July 1940
(R. H. Beamer).

Holotype: Male (UKan) 6292.

Allotype: Female, paratype female, Alamogordo, New Mexico;
15 May 1902 (Viereck and Rehn) (ANS).

123. Brevitrichia boharti, new species

FIGurRE 117

Maure.—Head cream; eyes metallic blue-green above, darker
below, frons small, triangular; ocellar tubercle red-brown, dusted
gray pollinose; back of head brown. Antennae with first segment
tan, second and third segments orange-brown; see figure for details.

Thorax dorsum black-brown, dusted with olive pollen, anterior
portion above humeral callus tan; humeral and supra-alar calli tan;
mesoscutellum tan, with brown central spot covering half of the
upper surface. Propleuron cream, mesanepisternum cream, with one
brown patch, mesokatepisternum red-brown. Wings smoky, veins
tan; halter stem red-brown, knob cream. Legs with femora brown
to black-brown, with light knees, tibiae and tarsi tan to brown,
darkening distally.

176 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen segment two with basal half red-brown, with a narrow
median band extending to posterior margin, remainder white; remain-
ing segments white above; ninth tergum white, with a brown spot at
base; see figure for details.

FrmaLe.—Head cream; eyes metallic blue-green to brown, with
a broad postocular ridge, with a band of brown behind eye; back of
head brown-black; frons very broad, with a brown band up the
middle, a faint median groove on lower half, a shallow groove ex-
tending from corner of eye to median ocellus. Ocellar tubercle red-
brown; ocelli orange. Antennal segments orange-brown. See figure
for details.

Ficure 117.—Brevitrichia boharti, new species, male, female: a, male wing; b, female wing;
c, d, lateral aspects of male and female heads; ¢, f, enlarged aspects of male and female
antennae; g, dorsal aspect of female head; h, 1, ventral and lateral aspects of male termi-
nalia; 7, ventral left and dorsal right aspects of female 8th and 9th segments; k, lateral
aspect of female 8th and 9th segments; /, bursal structure.

Thorax dorsum red-brown, with a dusting of tan pollen, a lighter
tan patch above humeral callus; humeral and supra-alar calli tan
to yellow. Scutellum tan, with a brown median spot. Pleural areas as
in male. Wings hyaline, veins brown; halter stem tan to brown,
knob cream; legs as in male.

Abdomen second segment broad, red-brown, with a light patch
on the mid-lateral portions; other segments brown basally, tan

SCENOPINIDAE OF THE WORLD Lz

distally; eighth segment red-brown; ninth tergum with 8 stiff spines
and a dorsal pit. See figure for details.

Length: Male body 2.7 mm., wing 2.2 mm.; female body 4.7 mm.,
wing 2.4 mm.

Type-locality: Anza, Riverside Co., California; 5 July 1956
(R. M. Bohart).

Holotype: Male (UCalD) 76.

Allotype: Female on same pin as type.

Paratypes: 1 9, Palm Springs, California; 4 May 1953 (W. R. M.
Mason) (CNC); 19,3 mi. s. of Palm Springs, 29 August 1951 (Timber-
lake) (PHT).

124. Brevitrichia castanea, new species
Ficure 118

Mauzr.—Head cream, eyes red-brown above, black-brown below;
frons small, triangular; mouthparts well developed; ocellar tubercle
red-brown, ocelli orange. Antennae with first two segments yellow-
brown, third segment missing.

Thorax dorsum red-brown, with granular pollinose, lateral margins
cream; scutellum all but the lateral corners red-brown; pleural areas
with prothorax cream; mesanepisternum cream with usual red-
brown spot; mesokatepisternum red-brown with cream band above.
Wings smoky, veins tan; halter stem brown, knob white. Legs with
femora red-brown, with yellow knees; tibiae and tarsi brown.

Abdomen second segment red-brown basally with median central
stem narrowing to a point posteriorly, with a narrow lateral base,
remainder pristine white; remaining segments white dorsally. Ninth
tergum red-brown, with a narrow posterior margin of white. See
figure for details.

FErMALE.— Unknown.

Length: Male body 2.5 mm., wing 1.7 mm.

Type-locality: Glens Ferry, Idaho; 5 August 1927.

Holotype: Male (USNM) 67487.

125. Brevitrichia coquilletti, new species
Figure 119
Pseudatrichia griseola Coquillett, 1900, p. 501, (in part).

Maue.—Head cream; eyes red-brown above, black-brown below;
frons very narrow, triangular; ocellar tubercle red-brown, covered
with granular gray pollen. Antennae with first two segments brown,
third yellow-brown to red-brown; see figure for shape.

Thorax dorsum red-brown; a median band extending two-thirds
of the way to the scutellum, bordered on either side by narrower

178 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 118.—Brevitrichia castanea, new species, male: a, wing; }, lateral aspect of head;
c, d, ventral and lateral aspects of terminalia.

SCENOPINIDAE OF THE WORLD 179

J.

Figure 119.—Brevitrichia coquilletti, new species, male, female: a, wing; 5, lateral aspect of
male head; c, lateral aspect of female head; d, e, enlarged details of male and female
antennae; f, dorsal aspect of female head; g, h, ventral and lateral aspects of male termi-
nalia; i, ventral left and dorsal right aspects of female 8th and 9th segments; J, lateral
aspect of female 8th and 9th segments; &, bursal structure.

bands extending from just behind the humeral callus, nearly attain-
ing the lateral margins of the scutellar sulcus, remaining areas yellow,
covered with gray pollen, hairs thin and indistinct; humeral and
supra-alar calli cream; scutellum yellow, with a median band of
red-brown which continues anteriorly across sulcus, fading along

180 U.S. NATIONAL MUSEUM BULLETIN 277

the midline. Pleural areas of prothorax cream, mesanepisternum
yellow with a red-brown patch; mesokatepisternum red-brown with
a yellow band above, all covered with granular pollen. Wings smoky,
veins tan; halter stem brown, knob cream; legs yellow-brown, becoming
brown distally.

Abdomen second segment with red-brown ‘‘T”’ with white lateral
and posterior margins, remaining segments with red-brown ‘“T’’s
fading posteriorly, mostly white; ninth tergum with red-brown basal
spot, remainder white, with long hairs; see figure for details.

Frma.Le.—Head cream; eyes red-brown to metallic, with a broad
postocular band extending across back of head, brown patches from
corner of eyes to back of head; frons broad, with a median groove
bordered by two bands occupying a little more than half of the
frontal width, separated from postocular ridge by a shallow sulcus
from the corner of the eye to the back of lateral ocelli, ocellar triangle
red-brown, pollinose; ocelli orange to red. Antennae (missing on
allotype) first two segments brown, third orange. See figure for shape.

Thorax dorsum with red-brown median and lateral stripes as in
male, separated by orange stripes, lateral margins yellow-orange,
all covered by gray pollen; humeral and supra-alar calli cream;
scutellum yellow, with a median stripe. Pleural areas as in male.
Wings smoky brown, veins tan; halter stem cream, knob cream.
Legs as in male.

Abdomen with median red-brown “‘Ts,”’ which become fainter pos-
terlorly; lateral and posterior areas tan. Eighth segment red-brown;
ninth tergum with 6 stiff spines and much hair; see figure for details.

Length: Male body 2.7 mm., wing 2 mm.; female body 4 mm., wing
2.1 mm.

Type-locality: Los Angeles, California (Coquillett) as paratype of
Pseudatrichia griseola.

Holotype: Male (USNM) 67483.

Allotype: Female on same pin.

Paratypes: 1 o, Corcoran, California, 22 July 1947 (W. W. Wirth);
2 oo", West Side Sta., California, 25 April 1930, on L. alyssoides; 1 2,
Victorville, California, 30 May 1944 (A. L. Melander) (USNM).

126. Brevitrichia davisi, new species

FiagureE 120

Ma.re.—Head cream; eyes red-brown above with blue-green metal-
lic luster, lower half more intensely green; frons narrow, triangular;
ocellar tubercle red-brown, frosted with gray pollen, ocelli orange.
Antennae with first two segments tan, third segment lost.

Thorax dorsum with black-brown mid-dorsal band, flanked by two
lateral bands, area above humeral callus tan, lateral margins and

SCENOPINIDAE OF THE WORLD 181

Ficure 120.—Brevitrichia davisi, new species, male, female: a, male wing; }, female wing;
c, lateral aspect of male head; d, lateral aspect of female head; e, enlarged detail of female
antenna; f, dorsal aspect of female head; g, A, ventral and lateral aspects of male termi-
nalia; i, ventral left and dorsal right aspects of female 8th and 9th segments, bursal
structure outlined with dashed lines; j, lateral aspect of female 8th and 9th segments.

anterior to scutellum yellow, frosted with gray and olive pollen;
humeral and supra-alar calli cream, scutellum cream, with red-brown

182 U.S. NATIONAL MUSEUM BULLETIN 277

median stripe extending across scutellar sulcus into a point. Pleural
areas with prothorax cream; mesanepisternum largely cream, with a
red-brown spot; mesokatepisternum red-brown, with a broad cream
band dorsally; all covered with gray pollen. Wings smoky, veins
cream; halter stem brown, knob cream. Legs: femora light red-brown,
pollinose; tibiae tan; tarsi tan, turning brown distally.

Abdomen second segment red-brown basally, with a central stem
and lateral extensions in the form of a ‘‘T,’’ remainder creamy white;
segment three with a narrow anterior band of red-brown, remainder
of segment and rest of abdomen white above. Ninth tergum yellow,
with a tan basal patch. See figure for details.

FrmaLe.—Head tan; eyes red-brown, with a metallic sheen;
postocular ridge medium broad, extending across back of head, brown
patches laterad of corner of eye; frons broad, with a darker tan median
band flanking a median groove, sulcus from upper angles of eyes to
back of lateral ocelli; ocellar tubercle red-brown, dusted with gray
pollen; ocelli orange-red. Antennae with first two segments brown,
third segment orange-brown. See figure for details.

Thorax dorsum with a black band on midline extending two-thirds
of the distance to scutellum, separated by thin orange-brown bands
from lateral black bands beginning behind the humeral callus but not
attaining the scutellar sulcus, remainder orange-brown, dusted gray
pollinose; humeral and supra-alar calli orange-brown; pleural areas
with prothorax orange, mesokatepisternum black, with orange dorsal
band, all dusted gray pollinose. Wings hyaline, veins brown; halter
stem brown, knob orange. Legs with femora red-brown, pollinose;
tibiae tan; tarsi tan basally, brown distally.

Abdomen with segments red-brown basally with ‘’T’”’-shaped mark-
ings, lateral areas and posterior margins tan; eighth segment orange-
brown; ninth tergum with 8 stiff spines and much fine hair. See figure
for details.

Length: Male body 3.2 mm., wing 2.3 mm.; female body 3.7 mm.,
wing 2.2 mm.

Type-locality: Leeds, Utah; 15 September 1931 (EK. W. Davis)

Holotype: Male (USNM) 67482.

Allotype: Female, Saint George, Utah; 15 June 1930 (E. W. Davis)
(USNM).

127. Brevitrichia dicksoni, new species

Ficure 121

Frema.e.—Head tan; eyes black-brown; postocular ridge narrow
separated on the midline behind ocellar tubercle, with faint brown
bands laterad of the midline; front broad, with two bands of brown
hairs on either side of a bare midline, not attaining ocellar triangle;

SCENOPINIDAE OF THE WORLD 183

sulcus from corner of eye to lateral ocellus separating frons from
postocular area; ocellar tubercle red-brown, pollinose; ocelli red.
Antennae with first segment tan, second light brown, third brown.
See figure for details.

Ficure 121.—Brevitrichia dickson1, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, dorsal aspect of head; e, ventral left and dorsal right
aspects of 8th and 9th segments, bursal structure outlined on ventral surface; f, lateral
aspect of 8th and 9th segments.

Thorax dorsum orange-yellow, a little darker on either side of the
midline anteriorly, with a thin lateral red-brown stripe extending
from the humeral callus to the scutellum, pollinose, dorsum with
short stiff hairs; humeral callus cream and orange with numerous
short hairs on the dorsal surface. Scutellum orange, with narrow
median red-brown band. Pleural areas with prothorax cream, mes-
anepisternum orange, with red-brown central patch; mesokatepister-
num with only the lowest portion red-brown, remainder orange,
pollinose. Wings hyaline, veins cream to brown; halter stem tan,
knob cream. Legs orange-brown, except tarsi tan.

184. U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen second segment broad, orange, with median brown spot,
granular; remainder orange, eighth segment orange, shining; ninth
tergum with 7 stiff spines and a ventral tuft of hairs; see figure.

Maue.— Unknown.

Length: Female body 4 mm., wing 2.1 mm.

Type-locality: 5.7 mi. w. of Glacier, California; 25 July 1960
(R. C. Dickson).

Holotype: Female (CAS) 8937 (ex [PHT)).

Paratype: 1 9, same data as type (PHT).

128. Brevitrichia downeyi, new species

FiGurE 122

Maue.—Head cream; eyes brown above, black below, may be
metallic in life; frons small, triangular; ocellar tubercle red-brown,
frosted tan pollinose, ocelli brown-red. Antennae with first two
segments brown, third segment yellow-brown. See figure for details.

Thorax dorsum black-brown, frosted gray pollinose, yellow bands
dividing dark area into three parts, pronotum cream above humeral
callus, with a black band separating it from the callus. Humeral
and supra-alar calli cream, scutellum cream with a narrow median
dark stripe. Pleura with prothorax cream; mesanepisternum cream,
with a black-brown spot; mesokatepisternum black-brown; all frosted
gray pollinose. Wings smoky, veins tan; halter stem brown, knob
cream. Legs femora light brown, frosted, with a light knee; tibiae
tan basally darkening to brown distally, tarsi brown.

Abdomen with second segment with a black-brown proximal band
with a central stem narrowing distally, with lateral extensions like
a “T,” remainder of margin white; third segment with a narrow
basal brown band, remainder white; remaining segments white.
Ninth tergum white, with a brown basal spot, many long hairs; see
figure for details.

Frma.te.—Head cream, eyes black-brown, may be metallic in life;
postocular ridge broad, with a brown band laterad of the corner of
the eye to the back of the head, sulcus from corner of eye extending
to back of head, separated by one-half the width of the ocellar tubercle
allowing front to reach the back of the head. Frons broad, with a
median groove dividing a band of light brown, covered with short
hairs; ocellar tubercle black-brown, frosted gray pollinose; ocelli
red-orange; antennae all segments brown. See figure for details.

Thorax dorsum black-brown, with two longitudinal yellow stripes
extending anteriorly between the yellow patch anterior to the
scutellum to the yellow patches above the humeral calli, granular
pollinose; humeral and supra-alar calli cream; scutellum cream-
yellow, with a narrow median brown patch. Pleural areas with

SCENOPINIDAE OF THE WORLD 185

jl
'

' mai 1
wir)
ae

Oe)

Ficure 122.—Brevitrichia downey?, new species, male, female: a, wing; b, c, lateral aspects
of male and female heads; d, ¢, enlarged details of male and female antennae; f, dorsal aspect
of female head; g, h, lateral and ventral aspects of male terminalia; 7, ventral left and
dorsal right aspects of female 8th and 9th segments, bursal structure outlined; 7, lateral
aspect of female 8th and 9th segments; k, bursal structure.

prothorax cream, mesanepisternum cream with a black-brown patch,
mesokatepisternum black-brown, granular pollinose. Wing hyaline,
veins tan; halter stem brown, knob creamy white. Legs with femora
brown; tibiae light brown, tarsi dark brown.

186 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen brown, mottled tan. Eighth segment brown, ninth
tergite with 7 stiff spines, moderately hairy. See figure for details.

Length: Male body 3.3 mm., wing 2 mm.; female body 3.5 mm.,
wing 2 mm.

Type-locality: Davis, California; 16 August 1955 (J. C. Downey).

Holotype: Male (UCalD) 76.

Allotype: Female, Davis, California; 14 August 1955 (A. T.
McClay).

Paratype:1 o, Davis, California; 14 July 1955 (EK. I. Schlinger)
(UCalD).

129. Brevitrichia flocki, new species

Figure 123

FrmMa.Le.—Head cream; eyes deep red-brown, medium broad post-
ocular ridge, with a narrow brown band laterad of the corner of the
eye, the two sides separated by median groove behind the ocellar
tubercle; frons broad, with a deep median groove extending from the
antennal base to the median ocellus, bordered by brown bands on
either side; frons separated from postocular area by deep grooves
extending from the corner of the eye to the median ocellus. Ocellar
tubercle red-brown, pollinose; ocelli red. Antennae with first two seg-
ments red-brown, third deep red-brown; see figure for details on
shape.

Thorax dorsum solid black-brown, with gray pollen and a yellow
patch behind the humeral callus; humeral and supra-alar calli white.
Scutellum red-brown with cream lateral margins. Pleura with pro-
thorax cream; mesanepisternum red-brown with dorsal portion under
wings orange; mesokatepisternum red-brown, with a dorsal yellow
band. Wings hyaline, granular, veins brown; halter stem brown,
knob white. Legs red-brown with knee of femur and base of tibia and
tarsus yellow, pollinose.

Abdomen red-brown, with lighter posterior margins; eighth seg-
ment shining, a row of spines along margin of tergum; ninth tergum
with 7 stiff spines, and long hairs over most of the surface. See figure
for details.

Ma.e.— Unknown.

Length: Female body 3.8 mm., wing 2.2 mm.

Type-locality: Santa Rita Mts., Arizona; 26 June 1940 (R. A.
Flock).

Holotype: Female (USNM) 67485.

130. Brevitrichia griffini, new species
Figure 124

Ma.er.—Head cream; eyes red-brown above, black-brown below;
frons extending only half the distance to the median ocellus, with

SCENOPINIDAE OF THE WORLD 187

Ficure 123.—Brevitrichia flocki, new species, female: a, wing; b, lateral aspect of head; c, en-
larged detail of antenna; d, dorsal aspect of head; ¢, ventral left and dorsal right aspects
of 8th and 9th segments; f, lateral aspect of 8th and 9th segments; g, bursal structures.

a small dark triangle at upper limit, flaring broadly around antennal
bases to lower part of head around oral opening; ocellar tubercle
black-brown, ocelli clear; back of head with central portion black-
brown, lateral areas cream. Antennae black-brown. See figure for
details. ;

Thorax dorsum black-brown ground, with an orange patch over
humeral callus, overlain by longitudinally striped brown and tan
pollen, lateral margins with more yellow than female; humeral and
supra-alar calli cream. Scutellum with a broad median band of

313-168 O—69_—13

188 U.S. NATIONAL MUSEUM BULLETIN 277

k.

Figure 124.—Brevitrichia griffini, new species, male, femaie: a, wing; b, d, lateral and
frontal aspects of male head; c, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; h, 7, lateral and ventral aspects of male terminalia;
j, ventral left and dorsal right aspects of female 8th and 9th segments; &, lateral aspect
of female 8th and 9th segments; /, bursal structure.

black-tan, lateral margins cream. Pleural areas of prothorax brown;
mesanepisternum cream, with a dark central patch, mesokatepis-
ternum black-brown, pollinose. Wings smoky brown, veins brown.

SCENOPINIDAE OF THE WORLD 189

Halter stem brown, knob tan above, cream below. Legs black-brown,
pollinose; knees of femora yellow-brown.

Abdomen segments black-brown, pollinose, with thin creamy
membranous posterior margins, ninth tergum with basal portion
red-brown, remainder creamy; see figure for details.

FrmMALe.—Head yellow; eyes dark red-brown; postocular ridge
moderately wide, brown bands from corners of eyes down over back
of head, lateral areas yellow-orange, covered with red-brown an-
teriorly directed hairs; frons broad, nearly twice the width of the
ocellar triangle, divided by a median groove which is bordered by a
band of brown, with short golden brown hairs. Ocellar tubercle black-
brown, pollinose, ocelli red-orange. Antennae with first two segments
brown, the second shorter than the first, third segment black-brown;
see figure for details.

Thorax dorsum black-brown ground, covered with longitudinal
bands of brown and tan pollen, an orange patch over humeral callus;
humeral and supra-alar calli cream; scutellum with a broad median
brown band, with lateral margins cream. Pleural areas with prothorax
brown; mesanepisternum cream with a large dark brown patch;
mesokatepisternum dark brown, pollinose. Wings smoky brown,
veins brown; halter stem brown, knob tan above, cream below.

Abdomen red-brown, pollinose; eighth segment red-brown, shining;
ninth tergum with 7 stiff spines, hairy; see figure for details.

Length: Male body 3.2 mm., wing 2.3 mm; female body 4.5 mm.,
wing 2.5 mm.

Type-locality: Mount Laguna, San Diego Co., California; 21
June 1963 (H. L. Griffin).

Holotype: Male (CAS) 8924, on loan from (UCalB).

Allotype: Female same data as type (CAS).

Paratype:1 2, Mt. Laguna, San Diego Co., California; 21 June
1963 (P. D. Hunt) (UCalB).

131. Brevitrichia griseola (Coquillett)
Ficgurr 125
Pseudatrichia griseola Coquillett, 1900, p. 501.

This species, as are most Brevitrichia, is limited in its distribution
to a small geographical area, in this case along the middle reaches of
the Rio Grande. Specimens were examined from near the type locality,
near Las Cruces, New Mexico, and Lajitas, Texas in the Big Bend
area, where wasps were provisioning their nests with them.

More detailed descriptions of the male and female follow. The para-
type of Coquillett is now the new species B. coquilletti.

190 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 125.—Brevitrichia griseola (Coquillett), male, female: a, male wing; b, female wing;
c, d, lateral aspects of male and female heads; ¢, f, enlarged details of male and female
antennae; g, dorsal aspect of female head; h, 1, ventral and lateral aspects of male ter-
minalia; 7, ventral left and dorsal right aspects of female 8th and 9th segments; &, lateral
aspect of female 8th and 9th segments; /, bursal structure.

Ma.r.—Head cream; eyes red-orange-brown, darker below; frons
triangular, slightly more than one-third the distance from the antennal

SCENOPINIDAE OF THE WORLD 191

base to medium ocellus, with a small brown dash at peak; ocellar
triangle black-brown, pruinose; ocelli red; a narrow postocular rim
laterad of ocellar tubercle. Antennae with first segment brown, second
segment red-brown, third segment brown. See figure for details.

Thorax dorsum black-brown, heavily dusted pruinose, a yellow
spot above humeral callus; sides of dorsum yellow, two crescent-
shaped yellow areas extending from lateral angles of scutellum,
nearly meeting but leaving a continuous black line down the midline;
humeral callus cream, with a dark band dorsally. Scutellum yellow
with a large median brown spot. Pleural areas dark brown ventrally,
yellow dorsally. Wings smoky, veins brown; halter stem brown,
knob cream to tan. Legs with femora red-brown, dusted pruinose;
tibiae and tarsi orange, darkening to red-brown distally.

Abdomen with terga red-brown anteriorly, with a central stem
and lateral arms in the form of a “T,’’ remainder of segments white;
this pattern narrows posteriorly; ninth tergum brown basally, cream
distally. See figure for details.

Frmate.—Head cream, eyes red-brown, postocular rim moderately
wide, unmarked, with short brown hairs, the two halves divided on
the midline behind the ocellar triangle; frons broad, with a brown
median patch on upper two-thirds, covered with short hairs; upper
part of frons cut off by shallow grooves extending from the corners of
the eyes to just behind the ocellar triangle; ocellar triangle black-
brown, dusted gray pruinose, ocelli red-brown.

Thorax dorsum red-brown, dusted gray pruinose, marked with
extensive areas of yellow above the humeral callus, along the margins
above the wing and the central postnotum, extending anteriorly as
two thin bands; humeral callus cream; scutellum yellow with a median
red-brown spot. Pleura red-brown ventrally, yellow above. Wings
hazy brown, veins brown anteriorly becoming faint posteriorly;
halter stem brown, knob cream. Legs with femora and tibiae red-
brown; tarsi with first segment orange, remainder red-brown, dusted
gray pollinose.

Abdomen with first three segments red-brown, with thin white
membranous margins, next three segments tan; eighth segment red-
brown; ninth tergum with 8 stiff spines; see figure for details.

Length: Male body 3.6 mm., wing 2 mm.; female body 4.1
mm., wing 2.5 mm.

Type-locality: Mesilla (near Las Cruces), New Mexico; 25 May
1897 (T. D. A. Cockerell).

Type: (USNM) 4712.

Material seen: 1 o, Las Cruces, New Mexico, 16 June 1917
(J. M. Aldrich); 1 o, 2 99, 1 mi. w. Lajitas, Brewster Co., Texas,

192 U.S. NATIONAL MUSEUM BULLETIN 277

23-30 April 1963 (H. E. Evans) collected from solitary wasp burrow;
all in (USNM).

132. Brevitrichia halli, new species

FIGuRE 126

Matur.—Head tan, back of head brown; eyes metallic blue-black,
darker below; front narrow triangular; ocellar tubercle black-brown,
frosted; ocelli clear; antennae brown. See figure for details of shape.

Thorax dorsum black-brown, granular, with a yellow-orange band
along lateral margins broken above wing bases; humeral callus yellow-
orange, scutellum with a broad red-brown triangle and yellow-orange
lateral angles. Pleural areas mixed red-brown and yellow-orange,
granular; wings clear, veins brown; halter stem tan, knob tan. Legs
with femora brown, granular, with yellow knee; tibiae and tarsi
brown.

Abdomen with segments black-brown, with thin posterior white
bands; ninth tergum tan, with brown basal patch. See figure for
details.

FrmaLe.—Head cream-yellow; back of head brown; eyes metallic
blue-black, postocular ridge moderately wide, brown patches laterad
of corner of eye; frons broad, with a median groove on lower half, a
tan median patch, forked ventrally, with short anteriorly directed
brown hairs. Antennae with first two segments brown, third segment
black-brown. See figure for details.

Thorax dorsum black, overlain with brown and tan longitudinally
striped pollinose scales; humeral and supra-alar calli tan; scutellum
with a narrow median red-brown patch, lateral areas tan. Pleural
areas black-brown covered with tan pollinose scales. Wings hyaline,
veins brown; halter stem tan, knob tan. Legs with femora red-brown,
knees yellow, covered with tan pollinose scales, remainder red-brown.

Abdomen red-brown, granular with white posterior membranous
margins; eighth segment red-brown, ninth tergum with 9 stiff spines.
See figure for details.

Length: Male body 2.6-3 mm., wing 1.9-2 mm.; female body
4.5 mm., wing 2.4 mm.

Type-locality: 4 mi. w. Pinon Flat, Riverside Co., California;
11 June 1954 (J. C. Hall).

Holotype: Male (UCalD) 77.

Allotype: Female on same pin with type.

Paratype: 1c, Verdemont, California; 25 May 1948 (A. L.
Melander) (USNM).

SCENOPINIDAE OF THE WORLD 193

Ficure 126.—Brevitrichia halli, new species, male, female: a, male wing; b, female wing;
c, d, lateral aspects of male and female heads; ¢, f, enlarged details of male and female
antennae; g, dorsal aspect of female head; A, 1, ventral and lateral aspects of male ter-
minalia; j, ventral left and dorsal right aspects of female 8th and 9th segments, bursal
structures outlined; &, lateral aspect of female 8th and 9th segments.

194 U.S. NATIONAL MUSEUM BULLETIN 277

133. Brevitrichia helenae (James)
FicurE 127
Pseudatrichia helenae James, 1938, p. 22.

I am indebted to Dr. James for the loan of the paratype male for
figuring. All attempts to locate the type have failed.

Length: Male body 2—2.5 mm.; wing 1.8 mm.

Type-locality: Roggen, Colorado; 29 May 1937 (Helen B. James).

Type: (CSU) lost.

Paratype: 107, same data (MTJ).

134. Brevitrichia hodgdeni, new species

FIGURE 128

Mate.—Head tan; eyes red-brown, with a metallic sheen, darker
below; frons narrow, triangular; ocellar tubercle red-brown, ocelli red-
brown; antennae black-brown. See figure for details.

Thorax dorsum black, covered with brown and tan pollinose scales
in longitudinal bands; lateral margins yellow-orange; humeral and
supra-alar calli orange; scutellum with a large central spot one-half
of its width, lateral angles yellow-orange. Pleural areas red-brown and
yellow in the usual arrangement, pollinose; wings smoky brown, veins
brown; halter stem brown, knob tan-brown. Legs: femora red-brown,
tibiae orange, tarsi orange basally to dark brown distally.

Abdomen dark red-brown, scarcely lighter along posterior margins;
ninth segment red-brown, with tan margins; see figure for details.

FrEmMALE.—Head tannish-brown, eyes dark red-brown, metallic, a
broad postocular rim with a red-brown patch laterad of the corner of
the eye; frons broad, with a median groove bordered by red-brown
bands; sulcus from corner of eye to lateral ocelli cuts off frons from
postocular ridge; ocellar tubercle black-brown, pollinose; ocelli black.
Antennae black-brown. See figure for details.

Thorax as in male.

Abdomen as in male, eighth segment red-brown, shining; ninth
tergum with one dorsal spine and 7 pits. See figure for details.

Length: Male body 3 mm., wing 2.1 mm.; female body 3.7 mm.,
wing 2.2 mm.

Type-locality: Lake City, California; 15 July 1941 (Burt Hodg-
den).

Holotype: Male (UKan) 6293.

Allotype: Female same data (UKan).

SCENOPINIDAE OF THE WORLD 195

Ficure 127.—Brevitrichia helenae (James), male: a, wing; , lateral aspect of head; ¢, d,
ventral and lateral aspects of terminalia.

196 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 128.—Brevitrichia hodgdeni, new species, male, female: a, wing; b, c, lateral aspects
of male and female head; d, dorsal aspect of female head; e, f, enlarged details of male
and female antennae; g, fh, ventral and lateral aspects of male terminalia; 1, ventral left
and dorsal right aspects of female 8th and 9th segments, bursal structure outlined with
dashed lines; 7, lateral aspect of female 8th and 9th segments.

135. Brevitrichia insulana (Cole)

FicurE 129

Pseudatrichia insulana Cole, 1923, p. 462.

The type and allotype are figured. Female ninth tergum with 9 stiff
spines.

Length: Male body 3 mm., wing 2.3 mm.; female body 5.8 mm.,
wing 2.6 mm.

Type-locality: Isla Raza, Gulf of California, Mexico; 21 April 1921
(E. P. Van Duzee).

Type: (CAS) 1339.

Allotype: Female (CAS) 1340.

Specimens seen: 1 9, Isla Partida, Gulf of California, Mexico,
23 March 1953 (P. H. Arnaud) (CAS); 1 9, Rialto, San Bernardino
Co., California, 24 July 1956 (J. C. Hall) (UCalD).

136. Brevitrichia kerni, new species

FiIGuRE 130

FrmMaLe.—Head cream; eyes deep red-brown, postocular ridge
moderately wide, a brown patch laterad of the corner of the eye; frons
with a median groove extending from the antenna to the median

SCENOPINIDAE OF THE WORLD 197

Ficure 129.—Brevitrichia insulana (Cole), male, female: a, wing; , c, lateral aspect of
male and female head; d, enlarged detail of antenna; ¢, f, ventral and lateral aspect of
male terminalia; g, ventral left and dorsal right aspects of female 8th and 9th segments,
bursal structure outlined; A, lateral aspect of female 8th and 9th segments; i, bursal
structure.

ocellus, a band of short hairs on either side; ocellar tubercle red-brown,
pollinose, ocelli clear; a groove from the corner of the eye to the
back of the head cutting off a central piece nearly the width of the

198 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 130.—Brevitrichia kerni, new species, female: a, wing; , lateral aspect of head;
c, dorsal aspect of head; d, ventral left and dorsal right aspects of 8th and 9th segments,
bursal structure outlined; ¢, lateral aspect of 8th and 9th segments; f, bursal structure.

ocellar tubercle; antenna with first segment tan, second segment
brown basally, tan distally, third segment missing.

Thorax dorsum with a median red-brown band extending two-
thirds the distance to the scutellum, bordered on each side by a red-
brown band extending from behind the humeral callus to the scutel-

SCENOPINIDAE OF THE WORLD 199

lum, separated from the median band by a strip of orange, the whole
covered with pollinose scales and a few short hairs; humeral and supra-
alar calli cream, with an orange patch between; scutellum cream, with
a narrow median red-brown band. Pleura with prothorax cream;
mesanepisternum cream, with a median red-brown patch; meso-
katepisternum red-brown, with a dorsal cream band. Wings smoky,
veins tan to brown; halter stem brown, knob cream. Legs with
femora light brown, tibiae tan, tarsi tan basally, becoming darker
distally.

Abdomen second segment cream, with a red-brown “T”’ in the
anterior two-thirds, posterior segments cream becoming orange with
red-brown ‘“T’’s becoming less distinct and constricted posteriorly.
Eighth segment red-brown; ninth tergum with 7 stiff spines and very
abundant hair; see figure for details.

Matze.— Unknown.

Length: Female body 5 mm., wing 2.3 mm.

Type-locality: Kernville, California; 20 July 1940 (R. H.
Beamer).

Holotype: Female (UKan) 6294.

137. Brevitrichia melanderi, new species

FicureE 131

FrEMALE.—Head cream; eyes brown in dried specimen, postocular
ridge narrow, unmarked by pigmentation, divided on midline behind
ocellar tubercle, covered with short hairs; frons wider than ocellar
triangle, with a ‘‘Y’’-shaped brown patch with the arms above the
antennae, middle of stem darker brown, brown area clothed with
short hairs; ocellar tubercle red-brown, ocelli clear; a groove from
corner of eye, extending just back of the ocellar triangle to meet the
groove from the opposite side and one from the back of the head,
anterior to the margin of the back of the head. Antennae with all
segments red-brown. See figure for details of the head.

Thorax dorsum red-brown, probably covered with tan pollen in
nongreasy specimens, an orange patch on prothoracic dorsum, humeral
and supra-alar calli orange, with a connecting marginal band; scu-
tellum orange, with a round red-brown spot covering the center.
Pleura with prothorax orange, mesanepisternum orange with a red-
brown patch, mesokatepisternum red-brown; wings brown, granular,
veins brown; halter stem tan, knob cream below, tan above. Legs
red-brown changing to brown distally.

Abdomen segments all brown with light tan posterior membranous
margins; eighth segment red-brown; ninth tergum with 8 stiff spines.
See figure for details.

200 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 131.—Brevitrichia melanderi, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, dorsal aspect of head; e,ventral left and dorsal right aspects
of 8th and 9th segments, bursal structure outlined; f, lateral aspect of 8th and 9th seg-
ments; g, bursal structure.

Mate.—Unknown.
Length: Female body 5 mm., wing 2.4 mm.

SCENOPINIDAE OF THE WORLD 201

Type-locality: Cloudcroft, New Mexico; 27 May 1902 (A. L.
Melander).
Holotype: Female (USNM) 67484.

138. Brevitrichia minuta, new species
F1iGuRE 132

Ma.e.—Head tan; eyes red-brown, with a metallic gleam above,
darker below; frons triangular, with a small separate triangular area
in front of median ocellus; ocellar tubercle red-brown, with a gray
pollinose covering; ocelli red-brown; antennae with first two seg-
ments orange, third segment brown. See figure.

Ficure 132.—Brevitrichia minuta, new species, male, female: a, wing; , d, lateral aspects
of male and female heads; c, ¢, enlarged details of male and female antennae; f, dorsal
aspect of female head, frons collapsed; g, h, ventral and lateral aspect of male terminalia;
.1, ventral left and dorsal right aspects of female 8th and 9th segments, bursal structure
outlined; 7, lateral aspect of female 8th and 9th segments.

Thorax dorsum black, densely covered with grayish white crustlike
pollen, a white spot above the humeral callus; humeral and supra-alar
calli white to yellow. Scutellum with black-brown dorsal surface,
lateroposterior surface white or yellow, all granulose. Pleura with
prothorax cream to white, mesopleuron black-brown covered with
gray pollen; wings smoky, veins cream; halter stem brown, knob

202 U.S. NATIONAL MUSEUM BULLETIN 277

cream to white. Legs: femora black-brown, tibiae brown, tarsi brown
darkening distally.

Abdomen red-brown with narrow cream to white bands along
posterior margins; ninth tergum white with red-brown basal spot;
aedeagal spines short, knobbed. See figure for details.

FrmMaLe.—Head cream; eyes red-brown, metallic; postocular ridge
moderately wide, unmarked, divided on the midline; frons broad,
unmarked; ocellar tubercle red-brown, frosted gray pollen; ocelli
orange; a groove running from corner of eye to back margin of ocellar
tubercle. Antennae with first two segments orange, third segment
brown. See figure for details.

Thorax dorsum red-brown, with a median and two lateral bands,
yellow above humeral callus, lateral margins and median area ahead
of scutellum tan, with two anterior bands nearly separating median
and lateral bands; scutellum yellow, with a central triangular patch
pointing posteriorly. Pleura with prothorax cream; mesanepisternum
cream, with red-brown patch; mesokatepisternum red-brown; wings
smoky white, veins cream; halter stem brown, knob cream. Legs as
in the male.

Abdomen red-brown with tan posterior margins grading to yellow-
brown posteriorly; eighth segment red-brown; ninth tergite with a
marginal fringe of hairs instead of spines. See figure for details.

Length: Male body 2.5 mm., wing 1.7 mm.; female body 2.5 mm.,
wing 1.7 mm.

Type-locality: Beaver Dam, Arizona; 20 June 1939 (Timberlake).

Holotype: Male (CAS) 8919 (ex [PHT]).

Allotype: Female, same data (CAS).

Paratype:1 &, Palm Springs, California; 24 June 1952 (Timber-
lake), (PHT).

139. Brevitrichia oculivirida, new species
FiguRE 133

Matz.—Head tan; eyes blue-green, metallic above, darker below,
very brilliant under the proper light; frons small, triangular; ocellar
tubercle red-brown, covered with gray pollen; back of head brown.
Antennae with first segment brown; second segment orange-brown;
third segment black-brown; see figure for details.

Thorax dorsum black-brown, except for lateral margins yellow,
covered with alternate longitudinal bands of tan and gray pollen,
humeral and supra-alar calli cream, scutellum cream with a broad
central spot of black-brown. Prothorax cream, mesanepisternum yel-
low with usual red-brown spot, mesokatepisternum red-brown, with
upper margin yellow, pollinose; wings hyaline, veins tan; halter stem
brown, knob tan. Legs red-brown except knees of femora yellow.

SCENOPINIDAE OF THE WORLD 203

6.

Ficure 133.—Brevitrichia oculivirida, new species, male: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, ¢, ventral and lateral aspects of terminalia.

Abdomen second segment with black-brown “T’’-shaped figure,
with short posterior lateral extensions at base of stem, lateral and
posterior margins pristine white as is the rest of the abdomen; ninth
tergum white, with dark basal band and spot. See figure for details.

313-168 O—69——14

204 U.S. NATIONAL MUSEUM BULLETIN 277

FremMA.Le.— Unknown.

Length: Male body 4.3 mm., wing 2.5 mm.

Type-locality: Isla Ildefonso, Gulf of Calif., Mexico; 30 March
1953 (P. H. Arnaud).

Holotype: Male (CAS) 8931.

140. Brevitrichia ordwayi, new species

FIGURE 134

Ma.ze.—Head cream; eyes black-brown above, darker below; frons
triangular, with a small tan triangle on the lower frontal area above
the antennae; ocellar tubercle black-brown, dusted gray pollen, ocelli
black-brown. Antenna with first segment red-brown; second segment
black-brown; third segment reddish-brown, covered with short golden
hairs. See figure for details.

Thorax dorsum black-brown, pollinose; a small yellow patch above
humeral callus; lateral margins yellow, two short bands of yellow
extending anteriorly from the lateral angles of scutellum; humeral
callus buff, with a red-brown patch above; scutellum with a large
red-brown patch medially, yellow on lateral angles. Prothorax yellow,
mesanepisternum yellow, with a red-brown patch, mesokatepisternum
red-brown, with a dorsal yellow band; wings hyaline, veins brown
anteriorly tan posteriorly; halter stem brown, knob tan. Legs black-
brown pollinose, knee of femur yellow.

Abdomen segments dark brown with light posterior margins;
ninth tergum tan, with a black-brown patch basally; see figure for
details.

FremaLe.—Head cream; eyes red-brown, metallic; postocular
ridges medium wide with red-brown median patches, the two sides
widely separated; frons tan, with a median groove bordered by a
brown band that narrows toward the ocellar tubercle; ocellar tubercle
red-brown, pollinose, ocelli red; a groove extending from the corner
of each eye cutting off a central piece as wide as the ocellar tubercle.
Antennae with first two segments brown, third segment orange-
brown; see figure for details of head.

Thorax dorsum red-brown, heavily covered with gray pollen,
spot above humeral callus yellow, lateral margins narrowly yellow
between humeral and supra-alar calli, orange stripes extending
anteriorly one-third the length of the dorsum from the scutellar
groove, bordering a central stripe of red-brown; humeral and supra-
alar calli cream; scutellum yellow, with a median band of red-brown.
Prothorax cream, mesanepisternum cream, with a central red-brown
patch, mesokatepisternum red-brown, with a dorsal yellow stripe.

SCENOPINIDAE OF THE WORLD 205

Ficure 134.—Brevitrichia ordwayi, new species, male, female: a, male wing; b, female wing;
c, d, lateral aspects of male and female heads; ¢, f, enlarged details of male and female
antennae; g, dorsal aspect of female head; A, 7, ventral and lateral aspects of male termi-
nalia; 7, ventral left and dorsal right aspects of female 8th and 9th segments; &, lateral
aspect of female 8th and 9th segments; /, bursal structure.

Wings smoky white, veins red-brown; halter stem tan, knob tan.
Legs with femora red-brown, pollinose; tibiae tan, tarsi tan darkening
distally to brown.

206 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen second segment red-brown, with yellow-brown lateral
patches, remaining segments red-brown; eighth segment red-brown,
shining; ninth tergum with 5 stiff spines and much anteriorly directed
long hair; see figure for details.

Length: Male body 3.5 mm., wing 2.3 mm.; female body 4.2 mm.,
wing 2 mm.

Type-locality: 2.5 mi. s. Willcox, Arizona; 29 May 1956 (S.
Ordway) on mesquite.

Holotype: Male (CAS) 8920 (ex [PHT)).

Allotype: Female 18.2 mi. n.w. Oracle Junction, Pinal Co.,
Arizona; 6 September 1958 (Timberlake) (CAS) (ex[PHT]).

141. Brevitrichia pruinosa, new species

Figure 135

Ma.zr.—Head tan; eyes black-brown, with a metallic green cast
above, darker below; frons small, triangular; mouthparts brown,
well developed; palpi cream, short; ocellar tubercle black-brown,
dusted gray pollinose; ocelli red. Antennae brown. See figure for
details.

Ficure 135.—Brevitrichia pruinosa, new species, male: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, ¢, ventral and lateral aspect of terminalia.

Thorax black-brown, dusted with a central band of brown and
lateral bands of gray pollen; humeral and supra-alar calli tan;
scutellum black-brown, pollinose above, lateroventral margins yellow.

SCENOPINIDAE OF THE WORLD 207

Prothorax tan, mesanepisternum red-brown, except below wing tan,
mesokatepisternum red-brown, all pollinose; wings smoky, veins tan
(faint); halter stem brown, knob tan; legs with femora red-brown
pollinose, tibiae tan, tarsi tan, except brown at tips.

Abdomen second segment with a broad red-brown anterior margin
with a thick median stem without substantial lateral extension,
remainder of segment pristine white including posterior margin;
remaining segments essentially white but with anterior and lateral
margins red-brown; ninth tergum white, with a red-brown basal
patch; see figure for details.

FrmALEe.— Unknown.

Length: Male body 2.6 mm., wing 1.9 mm.

Type-locality: Pulpito (anchorage), Baja California, Mexico;
20 March 1953 (P. H. Arnaud) Sefton Orca Exped. to Gulf of Cali-
fornia.

Holotype: Male (CAS) 8932.

Paratypes: 407<’, same data as type (CAS).

142. Brevitrichia schlingeri, new species

FIGuRE 136

Mave.—Head cream; eyes red-brown, with metallic blue-green
sheen, darker below; frons very narrow; ocellar tubercle red-brown,
pollinose, ocelli red; antennae red-brown; see figure for details.

Thorax dorsum red-brown, solid except sides, covered with longi-
tudinal bands of tan and brown pollen; humeral and supra-alar
calli yellow; scutellum red-brown, except lateral margins yellow.
Prothorax red-brown below humeral callus, then cream-yellow;
mesanepisternum yellow, with a median red-brown patch; meso-
katepisternum red-brown, with a yellow dorsal band, pollinose. Wing
smoky hyaline, veins tan; halter stem brown, knob cream-yellow.
Legs with femora red-brown, pollinose; tibiae yellow-brown; tarsi
yellow-brown, darkening to brown distally.

Abdomen segments all brown with narrow cream posterior bands;
ninth tergum cream, with a tan patch basally which extends along the
open margin; see figure for details.

Frmate.—Head cream; eyes red-brown, postocular ridge broad,
continuing uninterruptedly behind ocellar tubercle, a tan patch
along the mid-dorsal portion behind the eyes; frons broad, with a
“Y”’-shaped orange median stripe and a triangular brown area above
the antennae filling the arms of the ‘‘Y,” a groove extending from
the corner of the eye to the back edge of the lateral ocelli. Antennae
with two basal segments brown, third segment black-brown: see
figure for details.

208 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 136.—Brevitrichia schlingeri, new species, male, female: a, male wing; b, female
wing; c, d, lateral aspects of male and female head; e, f, enlarged details of antennae;
g, dorsal aspect of female head; A, 7, ventral and lateral aspect of male terminalia; 7, ventral
left and dorsal right aspects of female 8th and 9th segments; k, lateral aspect of female
8th and 9th segments; /, bursal structures.

Thorax dorsum solid red-brown with areas of orange above humeral
callus, margin and posterior third along the midline, covered with
gray pollinose, except on greasy specimens; humeral and supra-alar
calli cream; scutellum yellow, with a median triangular red-brown
patch. Prothorax cream, mesanepisternum yellow, with a red-brown
patch, mesokatepisternum red-brown, with a yellow dorsal margin;
wings hyaline, veins brown; halter stem brown, knob cream-yellow.
Legs with femora dark red-brown, pollinose; tibia yellow-brown;
tarsi yellow, darkening to brown distally.

Abdomen second segment red-brown, with orange lateral areas,
remaining segments red-brown anteriorly with orange posterior
margins turning to orange-brown; ninth segment orange, sternum

SCENOPINIDAE OF THE WORLD 209

with shallow excavation; ninth tergite with 6 spinelike hairs and
much long black hair; see figure for details.

Length: Male body 3.5 mm., wing 2 mm.; female body 4.5 mm.,
wing 2.6 mm.

Type-locality: Lancaster, Los Angeles Co., California; 22 July 1957
(E. I. Schlinger).

Holotype: Male (UCalD) 78.

Allotype: Female, Kramer Hills, San Bernardino Co., California;
19 April 1959 (E. I. Schlinger) (UCalD).

Paratype: 1 o,5 mi. s. Lancaster, Los Angeles Co., California,
15 May 1956 (E. G. Linsley, J. W. MacSwain) (UCalB); 1 ¢, Pinto
Flats, Imperial Co., California, 5 May 1958 (F. X. Williams) (CAS);
1 9, Blythe, California, 1 May 1955 (N. R. Richards) (CNC).

143. Brevitrichia scitulaesca, new species

FIGURE 137

FrmaLe.—Head cream, eyes dark red-brown; postocular ridge
moderately wide, with a brown patch on either side; frons broad,
with a median groove bordered by a tan band with sparse hairs;
grooves from the corner of the eyes to the back of the head cut off
a portion nearly as wide as the ocellar tubercle; ocellar tubercle
black-brown, pollinose, ocelli red. Antennae all segments brown; see
figure for details.

Thorax dorsum black-brown, dusted with brown pollen, an orange
patch over the humeral callus, lateral margins cream, and two orange
vittae, broader at base, extend from the lateral portions of the
scutellar groove to the anterior margin of the dorsum dividing the
dark areas into three fields; humeral and supra-alar calli white;
scutellum orange with a median red-brown spot continuous with
the median band of the dorsum. Prothorax white; mesanepisternum
cream, with a median red-brown patch, mesokatepisternum red-
brown, with a creamy dorsal band, all dusted gray pollinose. Wings
smoky white, veins brown; halter stem tan, knob creamy white.
Legs with femora red-brown, with yellow knees; tibiae tan-brown;
tarsi tan-brown, darkening distally. All segments pollinose.

Abdomen segments red-brown, with broad white posterior bands;
eighth segment red-brown; ninth tergum with 6 stiff spines, sparse
hair. See figure for details.

Maue.— Unknown.

Length: Female body 4 mm., wing 2.3 mm.

Type-locality: 1 mi. w. Lajitas, Texas; 23-30 April 1963 (H. E.
Evans) in nest of digger wasp Slenostictia scitula.

Holotype: Female (USNM) 67486.

210 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 137.—Brevitrichia scitulaesca, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, dorsal aspect of head; ¢, ventral left and dorsal right aspects
of 8th and 9th segments; f, lateral aspect of 8th and 9th segments; g, bursal structure.

144. Brevitrichia timberlakei, new species

Fictre 138

M atze.—Head cream; eyes red-brown, with a blue-green metallic
sheen, more intense below; frons narrow, triangular; ocellar tubercle
red-brown, with gray pollen; ocelli red. Antennae with first two seg-
ments tan, third segment tan basally, brown distally.

Thorax red-brown, covered with gray-green pollen, area above
humeral callus and in front of scutellum yellow, as well as marginal
areas; humeral and supra-alar calli cream; scutellum yellow, with a

SCENOPINIDAE OF THE WORLD Zit

Ficure 138.—Brevitrichia timberlakei, new species, male, female: a, male wing; ), female
wing; c, d, lateral aspects of male and female heads; e, f, enlarged details of male and
female antennae; g, dorsal aspect of female head; h, i, ventral and lateral aspects of male
terminalia; 7, ventral left and dorsal right aspects of female 8th and 9th segments; k,
lateral aspect of female 8th and 9th segments; /, bursal structure.

thin median red-brown stripe. Prothorax cream, mesanepisternum
yellow, with a median red-brown spot, mesokatepisternum red-brown,
with a dorsal band of yellow, all dusted pollinose; wings smoky,

242 U.S. NATIONAL MUSEUM BULLETIN 277

veins tan; halter stem brown, knobs cream, with a marginal brown
band. Legs: femora red-brown, pollinose; tibiae tan; tarsi tan, dark-
ening to brown distally.

Abdomen second segment with a red-brown “T,” with a broad
lateral base, remainder and posterior portion of segment white,
remaining segments with red-brown ‘“T’’s then fading out to all
white; ninth tergum white, with brown basal patch, hairs very long;
see figure for details.

FrmMauLe.—Head cream; eyes red-brown, probably metallic in life;
postocular rim broad, with thin brown patch behind eyes; frons
broad, with a tan ‘‘Y’’-shaped band extending from antennae to
ocellar triangle and the width of the latter; ocellar tubercle red-brown,
pollinose; ocelli red; a groove extending from corner of eye to lateral
ocelli and then nearly straight to back of head, cutting off a portion
nearly as wide as ocellar tubercle separating the two postocular ridges.
Antennae with first segment brown, second tan, third orange, dark-
ening at the tips; see figure for details.

Thorax dorsum with red-brown central and two lateral bands
separated by an orange stripe running between orange area above
humeral callus to central area on posterior third of dorsum, lateral
margins orange; all dusted with gray pollen; humeral and supra-
alar calli cream; scutellum yellow, with a narrow median band of
red-brown, pollinose. Prothorax cream, mesanepisternum cream with
red-brown patch, mesokatepisternum red-brown with a dorsal yellow
stripe, all pollinose. Wings smoky, veins brown; halter stem tan,
knob cream with tan marginal stripe. Legs with femora brown,
pollinose; tibiae tan, tarsi tan, darkening to brown distally.

Abdomen second segment with a brown ‘‘T” with a red-brown
central stem and broad base in a field of yellow-brown, red-brown
“T” on third and fourth segments, turning to yellow-brown beyond;
eighth segment yellow-brown, shining; ninth tergum with 6 spinelike
hairs, somewhat hairy. See figure for details.

Length: Male body 3 mm., wing 2 mm.; female body 4.5 mm.,
wing 2.2 mm.

Type-locality: Riverside, California; 19 September 1950 (Timber-
lake).

Holotype: Male (CAS) 8921 (ex [PHT)).

Allotype: Female, same data.

145. Brevitrichia yucatani, new species
FictrRE 139
Brevitrichia griseola D. E. Hardy (not Coquillett), 1944a, p. 34.

Maur.—Head tan; eyes red-brown, with a metallic glint above,
black-brown below; frons small; ocellar tubercle black-brown, ocelli

SCENOPINIDAE OF THE WORLD les

red. Antennae with first segment brown, second orange, third orange-
brown basally, darkening to black-brown, with short hairs; see figure.

Thorax dorsum black-brown, with yellow lateral margins; humeral
and supra-alar calli yellow, scutellum yellow, with black-brown area
covering central third. Prothorax yellow; mesanepisternum yellow
with red-brown spot; mesokatepisternum red-brown with yellow
dorsal band. Wings smoky, veins tan; halter stem tan, knob cream
below, tan above. Legs with femora red-brown, tibiae tan, tarsi tan
at base darkening distally.

Abdomen segments red-brown, with posterior white bands on seg-
ments; ninth tergum white, with red-brown spot on basal portion;
see figure for details.

Ficure 139.—Brevitrichia yucatani, new species, male, female: a, wing; b, c, lateral aspects
of male and female heads; d, e, enlarged details of male and female antennae; f, g, ventral
and lateral aspects of male terminalia; h, ventral left and dorsal right aspects of female
8th and 9th segments; 7, lateral aspect of female 8th and 9th segments; 7, bursal structure.

FEMALE.—Head lost before descriptions made, last antennal seg-
ment orange.

Thorax dorsum black-brown, heavily dusted with gray pollen, a
cream patch behind the humeral callus; humeral callus cream; scutel-
lum yellow, with red-brown spot occupying middle half. Pleural areas
as in male. Wings dusted brown, veins brown; halter stem tan, knob
cream below, tan above.

Abdomen first four segments red-brown with white posterior bands,
rest red-brown. Eighth segment red-brown, shining; ninth tergum
with 4 stout spines; see figure for details.

214 U.S. NATIONAL MUSEUM BULLETIN 277

Length: Male body 3.5 mm., wing 2.1 mm.; female body 5.2 mm.,
wing 2.5 mm.

Type-locality: Suchiapa, Chiapas, Mexico; 25 March 1953 (R. C.
Bechtel and E. L. Schlinger).

Holotype: Male (CAS) 8937.

Allotype: Female, El Rancho, Guatemala; 17 February 1932
(C. N. Ainslee) (USNM).

Genus Metatrichia Coquillett
Metatrichia Coquillett, 1900, p. 500.

Type-species, Scenopinus bulbosus Osten Sacken (original designa-
tion). The genus Metatrichia Coquillett is characterized by large ro-
bust flies with broad, flat, blunt abdomens. Cell R5 in the wing is
closed, vein R4 branches from various points along the cell. The heads
are higher than long and the front bulges at the middle projecting the
antennae forward. The genus is represented in all regions but the

Palaearctic.
Key to the Species of Metatrichia

Wing hyaline... . Sie Piece agree: se EE Se lng Mi BS OPRBEr hat a De ME Pana Ree
Wing brown or yellow Beeaa era. st os) at Sopnpne Ghd ey peed anima aie at ee
2. Halter knob brown .. . iy os SGA nS

Halter knob white, antenna vellow, ome digas oath winite hice abdomen
with white bars with anteriorly directed swellings laterally. (Africa)
o'M. stevensoni (Bezzi)
3. Body with flattened scale-like hairs (North America)
o'M. bulbosa (Osten Sacken)
Body only sparsely covered with microscopic scales (Australia)
o'M. waterhousei (Paramonov)

4. Wing brownish. .... Phsc oe rcetitO
Wing yellow-brown, halter imo cream, anu dibte eae Me body with-
out scale-like hairs (S. America). . . .... . &M. robusta (Kréber)

5. Halter knob brown or yellow-brown. . . NHN STRELA ny Selec MRE ©
Halter knob orange-brown or tannish yellow hal a ire a

6. Halter knob brown, body covered with white flattened seale: Ree Hairs (North
America) ... . .. . . 9M. bulbosa (Osten Sacken)
Halter knob pelinnebrovai aes black-brown, velvety, body without
flattened scales (S. America) . . . .. . . 9M. robusta (Kréber)

7. Halter knob orange-brown, a very lanes ty over 7.5 mm. (Africa)
2M. lophrysoma (Speiser)
Halter knob tannish-yellow a moderate sized fly under 5.5 mm., antenna
orange at base dark at tip (Africa). . . . . . 9M. stevensoni (Bezzi)

SCENOPINIDAE OF THE WORLD 215

146. Metatrichia bulbosa (Osten Sacken)
Ficure 140
Scenopinus bulbosus Osten Sacken, 1877, p. 275.

This is the most widespread North American species in the family
Scenopinidae.

It is a robust species covered with flat scale-like hairs.

Length: Male body 4-4.5 mm., wing 3-3.3 mm.; female body
3.5-5.38 mm., wing 3.2—4 mm.

Type-locality: Missouri, in July (C. V. Riley).

Type: (MCZ) 1057.

I have examined specimens from the following localities: California;
Isla San José and Pulpito (anchorage), Gulf of California, Mexico;
Texas; New Mexico; Washington; Idaho; Nevada; Kansas; Nebraska
(Scotts Bluff); Illinois; South Carolina; District of Columbia (Wash-
ington); Virginia (Mt. Vernon); West Virginia (Lost River State
Park); Pennsylvania; New Jersey; New York (Babylon, Long Island) ;
Connecticut; Massachusetts (Hollister); Vermont (Rupert Mt.);
and Delaware. One specimen from Nogales, Arizona, was taken
from a tunnel in a rose cane. No other clues as to their habits or
habitat are known.

147. Metatrichia lophrysoma (Speiser), new combination
Ficure 141
Pseudomphrale lophrysoma Speiser, 1920, p. 216.

The type of this species is presumed lost; a search of the material
from the Stettin Collection now housed in (PAS) failed to produce
it. However, two specimens exist in (AMNH) from the Belgian
Congo which agree closely with Speiser’s description and have been
illustrated. This is the largest of all the Scenopinids examined by me.

Length: Female body 7-7.5 mm., wing 5.5 mm.

Type-locality: Willowmore, [Republic of] South Africa; November
1907 (H. Brauns).

Type: deposited in Stettiner Museum (lost).

Specimens seen: 229, Banana, Belgian Congo, 6° S. 12°20’ E.;
July 1915 (Lang and Chapin) (AMNH).

148. Metatrichia robusta Kréber
FIGURE 142

Metatrichia robusta Kréber, 1913, p. 180.

The type of this species has been lost but I was fortunate to find a
number of specimens of both the male and female for figuring and
description. The female has not previously been described.

216 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 140.—Metatrichia bulbosa (Osten Sacken), male, female: a, wing; b,c, lateral aspects
of male and female heads; d, e, frontal aspects of male and female heads; f-h, ventral,
lateral and posterior aspects of male terminalia; 1, posterior aspect of female 8th and 9th
segments; 7, lateral aspect of female 8th and 9th segments; &, female 9th sternum and
bursa.

FreMALE.—Head black; eyes black-brown; a narrow postocular
band; frons rugose, with a smooth median band, lower portion juts
out extending the antennae outward. Mouthparts red-brown, well

SCENOPINIDAE OF THE WORLD 217

Ficure 141.—Metatrichia lophrysoma (Speiser), female: a, wing; ), c, lateral and frontal
aspects of head; d, posterior aspect of 8th and 9th segments; ¢, lateral aspect of 8th and
9th segments; f, 9th sternum and bursa.

developed completely filling oral cavity; palpi as long as oral cavity
with a fringe of tan hairs. Ocellar tubercle black, not elevated, median

918 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 142.—Metatrichia robusta Krober, male, female: a, wing; b, c, lateral and frontal
aspects of male head; d, ¢, lateral and frontal aspects of female head; f-h, ventral, lateral
and posterior aspects of male terminalia; 7, posterior aspect of female 8th and 9th segments;
j, lateral aspect of female 8th and 9th segments; k, female 9th sternum and bursa.

ocellus sunken, ocelli orange. Antennae black-brown, third segment
oval, velvety.

Thorax black with a few thin hairs, humeral and supra-alar calli
orange; an orange area below the wing. Wing brown, veins brown;
halter stem brown, knob orange-brown. Legs with femora red-brown,

SCENOPINIDAE OF THE WORLD 219

tibiae red-brown, tarsi light red-brown anteriorly, posterior legs with
yellow-brown tarsi.

Abdomen dark red-brown, with short reddish hairs, broad and thick,
tip of abdomen blunt; see figure for details.

Kréber gives the length of the male as 7 mm. and the wing-spread as
8 mm. Measurements made by me are as follows:

Length: Male body 5 mm., wing 3.6 mm.; female body 5.4 mm.,
wing 4 mm.

Type-locality: Paraguay.

Type: (MNH) lost.

I have seen several specimens from Nova Teutonia, Brazil, 27°11’ S.
52°23’ W.; 15 April, 30 November, and 18 December 1959 (Fritz
Plaumann) (CNC). Several from the same locality and collector in
(BMNH), as well as specimens from Bemberg, Misiones, Argentina,
14 March 1945 (Hayward, Willink, V. Golbach) and a specimen labeled
Patagonia (BMNH).

149. Metatrichia stevensoni (Bezzi), new combination
FIGuRE 143

Pseudomphrale stevensonit Bezzi, 1925, p. 96.
Pseudomphrale africana Lindner, 1962, p. 2.

This species from Africa is characterized in the male by the peculiar
markings on the abdomen, each segment having a posterior white
border and an anterolateral white spot. Lindner redescribed the
species as P. africana in 1962 and figured the abdomen. I am placing
the Lindner name in synonomy as it disagrees with Bezzi’s type only
in the detail of the tip of the wing which is more elongated than in
Lindner’s specimen.

The females I am assigning to this species come from western Africa.
On the basis of general wing characters, however, and using the dis-
tributional pattern of M. bulbosa in North America, I believe this
placement correct.

Frmate.—Head black, eyes black-brown; frons broad, granular,
covered with short anteriorly directed silvery scales, as is the mod-
erately wide postocular ridge. Mouthparts large and well developed,
postgenal area behind eyes orange, ocellar tubercle set off but not
prominent, ocelli red. Antennae with first segment brown, twice as
long as the second; second segment brown, short; third segment orange
at base, black at tip, truncated, twice as long as first two combined.
The frontal area does not project in this species as it does in MV.
bulbosa and M. robusta.

Thorax dorsum black, very granular with a close pile of narrow
scales, humeral and supra-alar calli orange, scutellum orange-brown,

313-168 O—69——15

230) U.S. NATIONAL MUSEUM BULLETIN 277

Figure 143.—Metatrichia stevensoni (Bezzi), male, female: a, male wing; b, female wing;
c, d, lateral and frontal aspects of male head; ¢, f, lateral and frontal aspects of female
head; g, enlarged detail of female antenna; h, i, ventral and lateral aspects of male ter-
minalia; 7, posterior aspect of female 8th and 9th segments; k, lateral aspect of female
8th and 9th segments; /, female 9th sternum and bursa.

lighter along the posterior rim. Pleural areas black and orange-red.
Wing brownish, veins brown; halter stem brown, knob tannish-
yellow. Legs red-brown except tarsi orange-yellow.

SCENOPINIDAE OF THE WORLD 221

Abdomen broad black-brown fading to red-brown posteriorly.
See figure for details of eighth segment and bursa.

Length: Male body 5-5.5 mm., wing 3.5-4 mm.; female body
4.5 mm., wing 3 mm.

Type-locality: Matapos, Southern Rhodesia; 21 September 1924
(R. H. R. Stevenson).

Type: (MCSNM).

Other specimens seen: 1, Dar es Salaam, Kenya, 18 November
1958-5 May 1959 (Lindner) Lindner’s type of P. africana; 1%,
Cholo, Nyasaland (R. C. Wood); 1c, Ruo, Nyasaland, 13 May 1916
(R. C. Wood) both in (BMNH); 2 99, Aus, South West Africa, Jan-
uary 1930 (R. E. Turner) (BMNH).

150. Metatrichia waterhousei (Paramonov), new combination

FIGURE 144
Pseudomphrale waterhousei Paramonov, 1955, p. 640.

This specimen from Australia definitely is a Metatrichia and not
Pseudomphrale as originally placed. It conforms in all ways to the
genus. The shape of the head, large mouthparts, scales on thorax,
robust body and genital structures, etc. leave no doubt as to its
position or identity.

Length: Male body 5 mm., wing 3.5 mm.

Type-locality: Killara, New South Wales; 20 January 1935
(D. F. Waterhouse).

Type: (CSIRO).

Genus Pseudatrichia Osten Sacken

Pseudatrichia Osten Sacken, 1877, p. 276.
Atrichia Schrank of Loew, 1866, p. 42.

Tyre-svecies <Atrichia longurio Loew (monobasic). The genus
Pscudatri-hia Osten Sacken is characterized by a head that is longer
then high, a long slender thorax and abdomen and the cell R5 closed
before the end of the wing.

The proper interpretation of this genus has been confused from the
beginning by Loew’s mistaking the sex of the type of P. longurio as
the male of the species, whereas he was dealing with a female. This
led to the concept of the genus as one where the males had the eyes
widely separated instead of contiguous on the midline, separating
the front from the ocellar tubercle. This necessitated the creation
of such genera as Pseudomphrale. The fact that no use was made of
genital structures in identifying the species of the family has led to
the creation of many unjustified genera in the past. The most useful
diagnostic characters are found in the shape of the head, wing venation,
the male terminalia, and the eighth sternum of the female.

222 U.S. NATIONAL MUSEUM BULLETIN 277

—

ae

Ficure 144.—Metatrichia waterhousei (Paramonov), male: a, wing; ), lateral aspect of head;
c, enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia.

The members of this genus are found only in the Nearctic and
Neotropical regions.

10.

1.

12.

13.

14.

15.

ou

SCENOPINIDAE OF THE WORLD 223

Key to Species of Pseudatrichia (males)

Wing browdl FUmMOSG 25s Gee ay a. 6s ele S--s e Sae S D
Wing hyaline es 5
Halter knob with dor al Surtate oe roe Brana: or ‘lacie. ee xa FO

Halter knob with base red-brown, remainder white
P. parvipennis, new species

Halter knob with dorsal surface brown or red-brown .. . . 4
Halter knob black with tip white (Brazil) . . P. otiictilata D, E. ede
Halter knob all red-brown (Costa Rica) . . . P. longiventris, new species

Halter knob with dorsal surface brown, ventral surface white
P. toupeta, new species

Halter knob white... . ona ey GW bet ak eo oe ee ce Ve aa
Halter knob with at least rie eee ae enue 2. ts ee
Antenna red-brown. .......... .... . P. morlani, new species
Antenna orange .. . . . . . . P. melanderi, new species
Halter knob with base red iawn, remade WiLG Coen ir empenes ie nie its, Sinemet
Halter knob with color at least along the dorsalrib. ........ #«I1
Antenna red-brown. .. . Seep eteiesates ate ose eoley oe ernie esc maee O)
Antenna black-brown or pone ae eee AO

Ventral margin of 9th tergum short, 9th pear mace a akin comb of hairs
P. atombomba, new species

Aedeagal structures very large, 9th sternum with a thick comb of hairs
P. cajoni, new species

Antenna black-brown. .......... .. . P. unicolor Coquillett
Antenna orange-brown . . . : . . . P. gracilipennis, new species
Dorsal rib of halter knob red- prow pleetine into white. ...... 12
Dorsal rib of halter knob brown with no bleeding or dorsalsurfacebrown. 13
Vein R4 ending well beyond end of cell R5 . . . P. barnesi, new species
Vein R4 ending equal to end of cell R56. . . . . P. truncata, new species
Doral surface of halter knob red-brown, brown or black-brown. . . . . 14

Only dorsal rib of halter knob red-brown, remainder white
P. bupennis, new species

Dorsal surface of halter knob brown or black-brown. . . . ney epee LO
Dorsal surface of halter knob red-brown ... .P. cockerelli, new species
Ventral surface of halter knob white. . . . . .P. punctulata D. E. Hardy

Ventral surface of halter knob black-brown, except a small white spot
P. peromysci, new species

Key to Species of Pseudatrichia (females)

Wing smoky brown .
Wing hyaline :
Halter knob partly ernite ;
Halter knob brown or red-brown . : :
Halter knob with base red-brown poeniee aie Pp. pammenitts new vepesies
Halter knob with base and dorsal rib red-brown, bleeding into white

P. cloudcrofti, new species

Be wm do

Antenna red-brown. . . . 5. de gO
Antenna black, short, 7th épdominal: segmerit! sharply donstricted at base
CBrazil) Pl oP OS, xo .. . . .P. punctulata D. E. Hardy

Abdomen with 8th ecenient lotte ail narrow (Costa Rica)
P. longiventris, new species

224

10.
ts
12.
13.
14.

15.

16.

i
18.
19.
20.

21.
22.

23.

24.
25.

26.

27.

U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen with 8th segment only slight longer than wide, marked with black

areas dorsally and ventrally (Mexico) . .... . . .P. longurio Loew
Halter knob white .... ERO nee nee eNT
Halter knob with at least ehe: Hage mraniced ite nea S Euan meat meet Tee
Antenna brown. . . ... . . P.albocinecta Van Duzee
Antenna red-brown, Black: ecere or orange 2 Ne ante ans nia nasser EineeN era EO
Antenna red-brown: or blaek-browa 2° 3445 (2-2 oe ee ee eee
Antenna orange .............. . P. melanderi, new species
Antenna black=browt, 2 ol0.co a te hrc eae Saat eel)
Antenna TEd-brown-s.. so eee ee ae P. aorlerae new species
Head longer than:high (7) 3" > a ee Pe parva IO Phe Liardy,
Head higher thanlong. . . . . P. bryanti, new species
Halter knob with base brown, remaiaes cahite: PEN Ts Men eee Late Btls Fa NSD,
Halter knob with more than just the base brown. . ........ 17
Antenna red-brown or black-brown. ...... Oe ete at ies
AMLCMMa lack aie (ee ay oyamios oe nla baie Ses P. arretcn new species
AMbeNMA TEG=brOwWmM' 20 5 ao uo ae ta) yh G oats eT ne nae Pe ce olen inl Mle
Antenna black-brown .. . ANN heey A Ce fate ete LL
Vein R4 ending even with the end “of call Re een He Cen eget ate

Vein R4 ending well beyond the end of cell R5 . P. biactisterna: new species
Vein R4 branching from just beyond the middle of cell R5
P. atombomba, new species
Vein R4 branching from near the distal fourth of cell R5
P. cajoni, new species
Eighthsternum notched on the posterior midline . P.evergreeni, new species
Eighth sternum gently rounded on posterior margin
P. unicolor, Coquillett

Halter knob with base and dorsal rib red-brown, remainder white. . . 18
Halter knob with dorsal surface cream, brown or black-brown .... 21
Red-brown not bleeding into'the white: 7) 9 Oe ee Cee og
Red-brown bleeding into:the whitey > cra) ae ne ee
Avitenna black-browm: 725 (sir ee eae 20
Antenna black. .. . ie Ap eee Pp. leechion new aaene
Vein R4 ending opens the oad of cell RS . . . . P. bakeri, new species
Vein R4 ending well beyond the end of cell R5 . . P. barnesi, new species
Antenna brown, black-brown or black . . . ee ee cued ae 2

Antenna orange, vein R4 ending before the end of cell R5
P. saccharcupa, new species
Antenna brown or black-brown . . . Scere oe
Antenna black, R4 ending before the ena of éell Re Ps convexa, new species
Antenna brown, R4 ending beyond the end of cell R5, small, under 5 mm.
P. concava, new species
Antenna black-brown, R4 ending beyond the end of cell R5, large, over 7 mm.
P. cockerelli, new species
Halter knob with dorsal rib red-brown, remainder cream... .. . . 25
Halter knob with dorsal surface brown or black-brown. ..... . . 26
Antenna black-brown. . ......... ... P.eaithales, new species
Antenna orange . . . . . . + . P. howdeni, new species
Halter knob with dorsal Pareace pean ventral surface white. . ... 27
Halter knob all brown except for a small white ventral spot, antenna black
P. peromysci, new species
Antenna orange-brown, frons narrow only as wide as the ocellar tubercle
P. sabroskyi, new species

SCENOPINIDAE OF THE WORLD 225

Antenna orange-brown, frons broader than the width of the ocellar tubercle
P. nordeni, new species

151. Pseudatrichia albocincta Van Duzee
FicurE 145
Pseudatrichia albocincta Van Duzee, 1926, p. 164.

Van Duzee’s description was based on a single female erroneously
considered to be a male. It is characterized by the silvery pile on back
of the head and the three-pronged eighth sternal plate.

Length: Female body 6 mm., wing 4 mm.

Type-locality: Bradley, Monterey Co., California; 23 May 1920
(E. P. Van Duzee).

Type: Correct to read ‘‘female” (CAS) 1874.

Other specimens: 1 9, Tehachapi Pass, California; 8 June 1929
(R. L. Usinger) (CAS).

Figure 145.—Pseudatrichia albocincta Van Duzee, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, ventral aspect of 8th and 9th sternum; e, lateral aspect
of 8th and 9th segments.

152. Pseudatrichia atombomba, new species
Ficgure 146
Pseudatrichia unicolor Cresson (not Coquillett), 1907, p. 1138.

Ma.e.—Head red-brown; eyes black-brown above, darker below;
frons small, swollen above antennae narrowing to less than the
width of the median ocellus; a narrow postocular ridge behind the

226 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 146.—Pseudatrichia atombomba, new species, male, female: a wing; 4, c, lateral
aspects of male and female heads; d, dorsal aspect of female head; e, enlarged detail of
antenna; f, g, ventral and lateral aspects of male terminalia; A, ventral aspect of female
8th sternum; i, lateral aspect of female 8th and 9th segments.

SCENOPINIDAE OF THE WORLD 221

eyes; ocellar tubercle prominent, ocelli brown. Mouthparts red-brown,
small, half the length of the oral opening, palpi as long as mouthparts.
Antennae red-brown, the second segment lighter. See figure for details
on shape.

Thorax dorsum black, transversely striated; humeral and supra-
alar calli red-brown; pleural areas dark red-brown. Wings clear,
veins brown; halter stem brown, knob with base red-brown, remainder
white. Legs with foreleg orange-brown with red-brown tarsus; middle
leg orange-brown, a little darker than foreleg; hind leg red-brown,
first segment of tarsus orange-brown, remainder darker.

Abdomen black, rugose, with a metallic sheen; remaining segments
black, shining; ninth tergum red-brown, truncated with the ventral
margin (upper) shorter; sternal comb sparse; aedeagal parameres
large, bulbous with two anteriomesal spurlike processes; see figure for
details.

FremaLe.—Head red-brown; eyes brown; postocular ridge broad;
frons swollen above antennae, with longitudinal and _ transverse
striations, covered with short hairs; ocellar triangle cut off, not
distinct, ocelli clear. Antennae as in the male.

Thorax as in male.

Abdomen first segment black, rugose, metallic; remaining segments
black, shining. Eighth sternum dark red-brown, convex on posterior
margins; see figure for details.

Length: Male body 4.7 mm., wing 3 mm.; female body 5.1 mm.,
wing 3.2 mm.

Type-locality: Alamogordo, New Mexico; 15 April 1902.

Holotype: Male (ANS).

Allotype: Female Alamogordo, New Mexico; 9 May 1902 (ANS).

153. Pseudatrichia bakeri, new species

FiGureE 147

FreMALE.—Head black-brown; eyes black-brown; with a broad
postocular ridge; frons striated, slightly wider than the ocellar tri-
angle; mouthparts very short, brown, not exceeding half oral opening;
palpi brown, shorter; ocellar triangle not prominent, cut off, ocelli
orange. Antennae with first two segments black-brown, third segment
missing.

Thorax dorsum black, transversely striated; humeral and supra-alar
calli red-brown; pleural areas red-brown; wings hyaline, veins brown;
halter stem red-brown, knob with red-brown dorsal rib bleeding into
white ventral portion. Legs: front legs, femora and tibiae orange,
tarsi red-brown; hind leg, basal half of femur orange, remainder of leg
red-brown.

228 U.S. NATIONAL MUSEUM BULLETIN 277

pee ee ey

Ficure 147.—Pseudatrichia bakeri, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, ventral aspect of 8th sternum; ¢, lateral aspect of 8th and 9th
segments.

Abdomen second segment black, punctured, with a purple metallic
glint, remainder of abdomen same except last two segments shining.
Eighth segment red-brown, concave on posterior margin of sternum
with two rounded lateral points; see figure for details.

Mauze.— Unknown.

SCENOPINIDAE OF THE WORLD 229

Length: Female body 5.3 mm., wing 3.2 mm.
Type-locality: Claremont, California (Baker).
Holotype: Female (USNM) 67466.

154. Pseudatrichia barnesi, new species

FiGureE 148

Mauz.—Head black; eyes black-brown above, darker below; frons
shining, with a protuberance above the antennal socket, very narrow
below the median ocellus; ocellar tubercle prominent, ocelli clear.
Mouthparts red-brown, filling three-fourths of the oral cavity; palpi
dark red-brown, half the length of oral cavity. Antennae black-brown;
see figure for details.

Ficure 148.—Pseudatrichia barnesi, new species, male, female: a, wing; b, c, lateral aspects
of male and female heads; d, dorsal aspect of female head; ¢, f, enlarged details of male
and female antennae; g-i, ventral, lateral and posterior aspects of male terminalia; ,
ventral aspect of female 8th sternum; &, lateral aspect of female 8th and 9th segments.

Thorax dorsum black, humeral and supra-alar calli orange-brown,
pleural areas red-brown. Wings clear, veins light brown; halter stem
brown, knob with dorsal edge brown bleeding into white ventral
portion. Front leg with femur red-brown, tibia yellow-orange, tarsus
brown; middle legs brown-orange; hind legs red-brown.

Abdomen second segment black, granular; remainder black; ninth
tergum pointed; ventral margin shorter; fringe of hairs on sternum

230 U.S. NATIONAL MUSEUM BULLETIN 277

moderately thick; aedeagus with bilobed parameres with median barb-
like projections; see figure for details.

FrmMaALe.—Head black; eyes black-brown; postocular ridge broad,
subshining; frons slightly wider than ocellar triangle, striated and
with a median depression; facial areas below antennae with silvery
pile; ocellar triangle set off, not prominent; antennae black-brown;
see figure for details.

Thorax as in male.

Abdomen as in male, except eighth sternum black-brown with a
shallow concave posterior margin with rounded lateral points; see
figure for details.

Length: Male body 5.2 mm., wing 3.2 mm.; female body 5.1-5.5
mm., wing 3.3-3.6 mm.

Type-locality: 3 mi. s. Glennville, Kern Co., California; 18
February 1959 (A. M. Barnes) reared from woodrat nest.

Holotype: Male (USNM) 67464.

Allotype: Female, same data as type (USNM).

Paratype: 10,5 2 2, same data as type; 1 o, 7 mi. s., 5.6 mi.
e. Tehachapi, Kern Co., California, 19 February 1953 (A. M. Barnes)
(AMB).

155. Pseudatrichia biacristerna, new species

Figure 149

FErMALE.—Head red-brown; eyes red-brown, with a wide post-
ocular flange behind the eyes; frons shining, with two diverging
bands of short hairs from median ocellus to antennal bases; ocellar
triangle cut off, not prominent; ocelli orange; mouthparts well de-
veloped, palpi half the length of the oral cavity. Antennae red-brown,
third segment covered with short hairs that reflect golden; see figure
for details.

Thorax dorsum black, with transverse striations; humeral and
supra-alar calli red-brown; pleura red-brown. Wings clear, veins
brown; halter stem brown, knob white except for base red-brown;
legs orange.

Abdomen black, metallic, intersegmental membrane brown medial-
ly, white laterally; eighth segment red-brown; sternum with two
sharp lateral points; see figure for details.

Mate.— Unknown.

Length: Female body 5.5-6.5 mm., wing 3.7—4.3 mm.

Type-locality: Santa Fe, New Mexico; August (T. D. A.
Cockerell).

Holotype: Female (USNM) 67469.

Paratypes: 2 992, Santa Fe, New Mexico, July, August (A.
Boyle; Cockerell nos. 1405, 1935) (USNM); 1 @, Santa Fe, New

SCENOPINIDAE OF THE WORLD Zo

Figure 149.—Pseudatrichia biacristerna, new species, female: a, wing; b, lateral aspect of
head; c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th ster-
num; f, lateral aspect of 8th and 9th segments.

Mexico, August (Cockerell no. 1710) H. Kahl Coll., C. M. Ace.
12676 (CNC).

156. Pseudatrichia bryanti, new species

FicurE 150

FremaLye.—Head black; eyes black, with a broad postocular flange;
frons only as broad as ocellar tubercle, with striations running from
eye margins toward the midline and median ocellus; ocellar tubercle
cut off, not prominent, ocelli clear. Mouthparts orange-brown,
nearly filling the oral cavity; palpi brown, half the length of the
oral cavity. Antennae black-brown, third segment with short re-
cumbent hairs reflecting a golden sheen; see figure for details.

232 U.S. NATIONAL MUSEUM BULLETIN 277

Thorax dorsum black, with transverse striations; humeral and
supra-alar calli red-brown; pleural areas red-brown. Wings clear,
veins brown; halter stem brown, knob white. Forelegs with femora
and tibiae orange, tarsi brown; middle leg with femur orange, tibia
red-brown, tarsus brown; hind leg with femur orange basally, red-
brown distally, tibia red-brown, first segment of the tarsus orange,
remainder brown.

Ficure 150.—Pseudatrichia bryanti, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum;
f, lateral aspect of 8th and 9th segments.

Abdomen second segment black, metallic, with a small lateral
white patch on the membrane; remaining segments black, metallic,
with larger white patches. Eighth segment red-brown, sternum blunt;
see figure for details.

Maxrze.—Unknown.

Length: Female body 6-7 mm., wing 3.6—4 mm.

Type-locality: Santa Catalina Mts., Arizona; 15 June 19388
(Bryant).

Holotype: Female (CAS) 8933.

Paratypes: 1 9, same data as type (CAS); 1 9, S. Catalina Mts.,
Arizona, alt. 9000 ft., 6 June 1926 (R. B. Streets) (USNM);
1 9, Cave Creek Canyon (S. W. Research Sta. of AMNH; 5000 ft.),
Chiricahua Mts. Cochise Co., Arizona, 21 April 1959 (S. M. and S. N.
Burns) (UCalD).

157. Pseudatrichia bupennis, new species

Figure 151

Mate.—Head black; eyes brown-black above, black below; frons
tapering from above the antennae to halfway to the median ocellus,
rest relatively broad, equal to half the width of the median ocellus,
relatively smooth with two rows of punctures; ocellar tubercle promi-
nent, ocelli clear; mouthparts red-brown, palpi very slender, short;
oral opening bordered by gray pubescence, laterad of antennae.

SCENOPINIDAE OF THE WORLD 200

= SS

| Z

Ficure 151.—Pseudatrichia bupennis, new species, male: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia.

234 U.S. NATIONAL MUSEUM BULLETIN 277

Antennae with first segment black-brown, second and third segments
red-brown; see figure for details.

Thorax dorsum black, striated; humeral and supra-alar calli
orange-brown; pleural areas black-brown; wings clear, veins dark
brown; halter stem black-brown; knob base and dorsal stripe brown,
dorsal rib and remainder white.

Abdomen segments 2—5 black, rugose with purple metallic sheen;
segments 6 and 7 black, shining, ninth tergite red-brown, pointed
dorsally, sloping ventrally, sternal brush thick; aedeagus orange, very
large, with two mesal peni prongs; see figure for details.

Frma.Le.— Unknown.

Length: Male body 5.5-6.5 mm., wing 3.7-4.5 mm.

Type-locality: Keen Camp, Riverside Co., California; 17 May
1959 (E. I. Schlinger).

Holotype: Male (UCalD) 72.

Paratype: 1 o&, Hastings Reservation, near Jamesburg, Monterey
Co., Calif.; 3 May 1958 (D. D. Linsdale) (UCalB).

158. Pseudatrichia cajoni, new species
Figure 152

Maur.—Head lost.

Thorax dorsum black, dusted with white hairs; humeral and
supra-alar calli red-brown; pleural areas black; wings clear, veins
brown; halter stem brown, knob white except base, brown. Forelegs,
coxa to tibia straw yellow, tarsus brown; middle leg, coxa to tibia
yellow-brown, tarsus brown; hind leg, coxa to tibia orange-brown,
tarsus brown.

Abdomen, segments 2-5 black, with a purple metallic sheen;
segments 6 and 7 black, shining; tergum of segment 9 red-brown,
smoothly rounded dorsally and sloping below; sternal comb of hairs
long and thick; aedaegus very large; see figure for details.

Frmaue.—Head black, eyes red-brown; postocular flange not
prominent, rounded shining; back of head convex, granulose; frons,
with a smooth median band bordered on each side by a broad band of
short hairs, eye margins smooth, wider than the ocellar tubercle.
Ocellar tubercle prominent, set off, ocelli red. Mouthparts well-
developed, red-brown; palpi half the length of the oral opening,
sides of opening silvery pilose. Antennae red-brown, second segment
shorter than first, third with silvery pubescence; see figure for details.

Thorax dorsum black, with pollinose patch above humeral callus
and covered with short hairs giving it a dusty appearance; humeral
and supra-alar calli orange-brown with a white membrane connecting
them; prothorax black with white pollen in front of coxa, remainder
of pleura black. Wings clear, veins brown, halter stem red-brown

SCENOPINIDAE OF THE WORLD 200

Ficure 152.—Pseudatrichia cajoni, new species, male, female: a, wing; b, lateral aspect of
female head; c, dorsal aspect of female head; d, enlarged detail of female antenna; e, ts
ventral and lateral aspects of male terminalia; g, ventral aspect of female 8th sternum
and 9th segment; 4h, lateral aspect of female 8th and 9th segments.

313-168 O—69

16

236 U.S. NATIONAL MUSEUM BULLETIN 277

knob white, except base red-brown. Front leg orange, except tarsus
darkening to red-brown; middle leg orange-brown, tarsus darkening
to red-brown; hind leg entirely red-brown.

Abdomen segments 2-5 black, with a metallic luster, covered with
short hairs, lateral portions interrupted with white membrane.
Eighth segment red-brown, eighth sternum deeply excavated along
posterior margin; see figure for details.

Length: Male body 6 mm., wing 4 mm.; female body 7.3 mm.,
wing 4.3 mm.

Type-locality: 1 mi. w. Cajon, San Bernardino Co., California,
24 December 1957 (A. M. Barnes).

Holotype: Male (USNM) 67472.

Allotype: Female, 1 mi. e. Cajon, California, 24 December 1957
(A. M. Barnes) reared from larvae in woodrat’s nest (USNM).

159. Pseudatrichia cloudcrofti, new species

Figure 153
Pseudatrichia unicolor Cresson (not Coquillett), 1907, p. 113.

Frema.te.—Head red-brown, longer than high; back of head ex-
cavated, pollinose; eyes black-brown; postocular ridge moderately
wide, subshining; frons dark red-brown, elongate, not wider than
ocellar triangle which is cut off but not prominent, ocelli clear. Mouth-
parts well developed, orange-brown, extending a little over half the
length of the oral opening; palpi orange, a little shorter. Antennae
with first two segments red-brown, third segment orange; see figure
for details.

Thorax dorsum dark red-brown, granular; humeral and supra-alar
calli orange-brown; pleura red-brown. Wings dusted brown, veins
brown; halter stem brown, knob base and dorsal rib brown, rest
white. Foreleg coxa straw colored, remainder of leg orange-brown;
middle and hind legs orange-brown.

Abdomen second segment red-brown, granular; remainder red-
brown, shining. Eighth segment light red-brown, sternum with three
points on posterior margin; see figure for details.

Mate.— Unknown.

Length: Female body 7 mm., wing 4.4 mm.

Type-locality: Cloudcroft, New Mexico; 16 June 1902.

Holotype: Female (ANS).

160. Pseudatrichia cockerelli, new species
Fiaure 154

Matz.—Head red-brown; eyes black, divided into larger upper and
smaller lower facets along the midline. Frons triangular, subshining,

SCENOPINIDAE OF THE WORLD 237

Ficure 153.—Pseudatrichia cloudcrofti, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢e, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

238 U.S. NATIONAL MUSEUM BULLETIN 277

vite Vi d

1
U

Ficure 154.—Pseudatrichia cockerelli, new species, male, female: a, wing; b, c, lateral aspects
of male and female heads; d, dorsal aspect of female head; e, f, enlarged details of male and
female antennae; g-i, ventral, lateral and posterior aspects of male terminalia; 7, ventral
aspect of female 8th sternum and 9th segment; &, lateral aspect of female 8th and 9th
segments.

with longitudinal striations; ocellar tubercle prominent, ocelli clear.
Antennae with first two segments red-brown; third black-brown,
pilose, twice as long as first two combined; see figure for details.

SCENOPINIDAE OF THE WORLD 239

Thorax dorsum black, with transverse striations; humeral and
supra-alar calli red-brown; pleural areas red-brown. Wings hyaline,
veins brown; halter stem brown, knob red-brown dorsally, white
ventrally. Foreleg orange, with tarsus darkening to red-brown distally;
middle and hind legs orange-brown, tarsi red-brown.

Abdomen second segment black-brown, with white lateral spots in
posterior membrane; remaining segments black-brown with nearly
equal white lateral margins. Ninth tergum concave along posterior
margin, ninth sternum with a sparse comb of hairs; aedeagus bilobed
with anteriorly directed medial spurs; see figure for details.

FremMALE.—Head red-brown; eyes black brown; broad postocular
ridge with transverse striations; frons wider than ocellar tubercle,
with longitudinal striations, swollen above antennal base; ocellar
triangle flat, set off, ocelli clear. Antennae as in male.

Thorax dorsum black, with transverse striations, humeral and
supra-alar calli red-brown with orange anterior and posterior spots;
pleura red-brown; wings hyaline, veins brown; halter stem brown,
knob with a thin red-brown dorsal rib, remainder white. Legs as in
male.

Abdomen second segment black-brown, granular; remaining seg-
ments black-brown with intersegmental membrane tan medially,
white laterally, eighth segment red-brown, sternum truncated; see
figure for details.

Length: Male body 7.2 mm., wing 3.7 mm.; female body 7.2 mm.,
wing 4.1 mm.

Type-locality: Florissant, Colorado; July 1919 (Cockerell).

Holotype: Male (USNM) 67470.

Allotype: Female, Florissant, Colorado; 7 July 1907 (S. A.
Rohwer) (USNM).

161. Pseudatrichia concava, new species

FicurE 155

FrmaLe.—Head red-brown; eyes black-brown, with a broad
postocular ridge; frons with a shallow median groove; ocellar tubercle
not set off, shallow; ocelli red. Antennae brown, third segment with
golden hairs.

Thorax dorsum black, transversely striated; humeral and supra-alar
calli red-brown; scutellum red-brown; pleural areas red-brown. Wings
clear, veins brown; halter stem brown, knob base and dorsal rib
brown, remainder white. First two pairs of legs straw yellow; hind leg
with orange-brown femur, straw yellow tibia, tarsus yellow basally,
darkening distally.

240 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 155.—Pseudatrichia concava, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

SCENOPINIDAE OF THE WORLD 241

Abdomen with segments 2—5 black, rugose with a purple metallic
sheen; segments 6 and 7 subshining; eighth segment black, sternum
hairy, concave on posterior margin; see figure for details.

Ma.r.— Unknown.

Length: Female body 4.5-5 mm., wing 3-3.2 mm.

Type-locality: 2 mi. n., 1.2 mi. w. Auberry, Fresno Co., California;
9 January 1958 (A. M. Barnes) reared from larva found in woodrat
nest.

Holotype: Female (USNM) 67462.

Paratypes: 1 9, same data as type; 1 9, Hackamore, Modoc Co.,
California, 24 May 1938 (A. M. Barnes) reared, nest of Neotoma
cinerea in (AMB); 1 9, Stanford Univ., California, 3 November, 1900
(Melander); 1 9, Yucca Valley, California, 5 October 1934 (A. L.
Melander) both (USNM).

162. Pseudatrichia convexa, new species

FIGURE 156

FEMALE.—Head black; eyes black-brown; postocular ridge with
a flat rim; frons swollen above antennae, striated as in figure; ocellar
triangle shallow, not cut off, ocelli red-brown. Mouthparts red-brown,
palpi half as long as oral opening. Antennae black; see figure for
details.

Thorax dorsum black, transversely striated; humeral and supra-
alar calli red-brown; pleural area red-brown. Wings clear, veins
brown; halter stem brown, knob base and dorsal rib brown, remainder
white. Front leg orange-brown, with black-brown tarsus, middle leg
red-brown, tarsus black-brown; hind leg red-brown, base of first
tarsal segment yellow-brown, remainder black-brown.

Abdomen segments 2, 3, 4 and the base of 5 black, rugose; remainder
black, shining. Eighth segment black, sternum broad, with rounded
posterior margin; see figure for details.

Mate.— Unknown.

Length: Female body 7 mm., wing 4.4. mm.

Type-locality: Lake City, Colorado; 21 August 1938, elev. 9000
ft. (Chas. L. Fluke).

Holotype: Female (USNM) 67458.

163. Pseudatrichia eaithales, new species
Figure 157

Frma.re.—Head black; eyes black-brown; postocular ridge broad,
striated; frons with a thin median groove bordered by transverse
striations; ocellar tubercle prominent, set off, ocelli brown; mouth-

242 U.S. NATIONAL MUSEUM BULLETIN 277

See

Ficure 156.—Pseudatrichia convexa, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum;
f, lateral aspect of 8th and 9th segments.

SCENOPINIDAE OF THE WORLD 243

Ficure 157.—Pseudatrichia eaithales, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral aspect of 8th sternum;
f, lateral aspect of 8th and 9th segments.

244 U.S. NATIONAL MUSEUM BULLETIN 277

parts dark red-brown, well developed; palpi over half the length of
oral opening. Antennae with first two segments black, third segment
black-brown; see figure for details.

Thorax dorsum black, with transverse striations; humeral and supra-
alar calli red-brown; pleural areas red-brown. Wings clear, veins brown;
halter stem brown, knob with base and dorsal rib brown, remainder
cream. Front and middle legs orange-brown with brown tibiae, hind
leg with all segments red-brown.

Abdomen segments 2-5 black, rugose, with a metallic purple sheen;
remaining segments black, shining. Eighth segment dark red-brown,
sternum slightly convex on posterior margin; see figure for details.

Mauze.— Unknown.

Length: Female body 6 mm., wing 3.8 mm.

Type-locality: Evergreen, Colorado; 5 August 1941 (M. T. James).

Holotype: Female (USNM) 67549.

Paratype: 1 9, Evergreen, Colorado; 21 July 1942 (M. T. James)
(CSU).

164. Pseudatrichia evergreeni, new species

Figure 158

FrmaLe.—Head black; eyes brown-black; postocular rim broad;
frons wider than ocellar triangle, swollen above the antennae, striated ;
ocellar tubercle prominent, set off, ocelli clear; mouthparts brown, well
developed; palpi very slender, half as long as the oral opening. Anten-
nae with first two segments red-brown, third segment blunt, black-
brown with a golden sheen from fine hairs; see figure for details.

Thorax dorsum black, transversely striated; humeral and supra-
alar calli red-brown; pleura dark red-brown. Wings clear, veins brown;
halter stem brown, knob base brown, remainder including dorsal rib
white. Front and middle legs red-brown, hind legs dark red-brown.

Abdomen segments 2-5 black, granular, with a metallic sheen;
6 and 7 shining; eighth segment black, posterior margin of sternum
convex with a shallow depression in middle third; see figure for details.

Maue.— Unknown.

Length: Female body 7 mm., wing 4 mm.

Type-locality: Evergreen, Colorado; 21 July 1942 (M. T. James).

Holotype: Female (USNM) 67461.

Paratype: 1 2, Evergreen, Colorado; 3 August 1941 (M. T. James)
(CSU).

165. Pseudatrichia garretti, new species

Figure 159

Frma.Le.—Head red-brown; eyes black-brown; postocular flange
flat, moderately broad; frons swollen, wider than ocellar triangle, with
grooves extending from laterad of the antennae to the median ocellus.

SCENOPINIDAE OF THE WORLD 245

—_——

SS
————

Ficure 158.—Pseudatrichia evergreeni, new species, female: a, wing; }, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

246 U.S. NATIONAL MUSEUM BULLETIN 277

4

Ficure 159.—Pseudatrichia garretti, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum;
f, lateral aspect of 8th and 9th segments.

Ocellar tubercle flat, not cut off, ocelli clear. Mouth parts red-brown,
well developed; palpi half as long as oral opening. Antennae with first
two segments red-brown, third segment black with golden pile hairs.

Thorax dorsum black, with transverse striations; humeral and supra-
alar calli red-brown; scutellum dark red-brown; pleural areas red-
brown. Wings clear, veins brown; halter stem brown, knob base brown
but dorsal rib and remainder white. Front leg orange, except last
tarsal segments brown; middle leg with femur orange, tibia orange-
brown, last tarsal segments brown; hind leg all red-brown.

Abdomen second through fifth segments dark red-brown granular,
submetallic, sixth and seventh segments dark red-brown. Eighth

SCENOPINIDAE OF THE WORLD 247

segment red-brown, sternum with posterior margin undulating; see
figure for details.

Matr.— Unknown.

Length: Female body 6.5 mm., wing 4.2 mm.

Type-locality: Keremeos, B.C., Canada; 26 June 1923 (C. B.
Garrett).

Holotype: Female (CNC) 8633.

166. Pseudatrichia gracilipennis, new species

Figure 160

Mauzn.—Head black, eyes brownish-black above, darker below, a
thin postocular flange; frons slightly rounded, with a narrow trans-
verse ridge just above the antennae, extending one-third the distance
to the ocellar triangle; ocellar triangle prominent, ocelli brown;
mouthparts red-brown, two-thirds the length of the oral opening,
palpi slender, short, half as long as the mouthparts. Antennae with
first segment red-brown, second and third orange-brown; see figure
for details.

Thorax dorsum black, transversely striated, humeral and supra-
alar calli with orange-brown spots; pleural areas a very dark red-
brown. Wings clear, veins brown; halter stem brown, knob base and
dorsal rib brown, remainder white. Fore and middle legs straw colored
with the tips of the tarsi red-brown, the second pair a little darker;
hind leg orange-brown.

Abdomen second segment to the base of fifth dull black, rugose,
remainder of fifth through seventh shining. Ninth segment black,
tergum very pointed, sternal comb of hairs straight; aedeagus without.
large knobs; prongs long, thin; see figure for details.

FrmMALE.— Unknown.

Length: Male body 6.5 mm., wing 3.5 mm.

Type-locality: Davis, California; 23 June 1952 emerged 27
September 1952 (KE. I. Schlinger and R. C. Bechtel).

Holotype: Male (UCalD) 73.

167. Pseudatrichia howdeni, new species

Ficure 161

Fremate.—Head black, longer than high; eyes black with red
glints; postocular flange narrow; frons half the width of the ocellar
triangle, transversely striated; ocellar triangle flat, not cut off, ocelli
clear. Mouthparts well developed, base brown, tip yellow; palpi
yellow. Antennae with first two segments red-brown, third segment
orange, notched at tip; see figure for details.

YAS U.S. NATIONAL MUSEUM BULLETIN 277

eas

i

Ficure 160.—Pseudatrichia gracilipennis, new species, male: a, wing; b, lateral aspect of
head; c, enlarged detail of antenna; d—f, dorsal, ventral and posterior aspects of terminalia.

SCENOPINIDAE OF THE WORLD 249

Ficure 161.—Pseudatrichia howdeni, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

250 U.S. NATIONAL MUSEUM BULLETIN 277

Thorax dorsum black, granular, transversely striated; humeral
and supra-alar calli orange-brown; pleural areas red-brown. Wing
clear, veins brown; halter stem brown, knob with base and dorsal
rib red-brown, remainder creamy white. Front leg orange; middle
lee orange-brown, except last segment of tarsus brown; hind leg,
coxa yellow, femur and tibia red-brown, tarsus yellow grading into
brown distally.

Abdomen black, rugose, with purplish metallic glints; eighth
segment red-brown, sternum with three points; see figure for details.

Mauz.— Unknown.

Length: Female body 5.5 mm., wing 3.3 mm.

Type-locality: Rustlers Park, Chiricahua Mts., Arizona; 8400 ft.;
11 June 1956 (H. and A. Howden).

Holotype: Female (CNC) 8634.

168. Pseudatrichia jamesi, new species

FIGURE 162

FrmMaLe.—Head deep red-brown; postocular flange narrow; frons
narrower than ocellar tubercle, transversely striated, a slight protu-
berance above antennae; ocellar triangle red-brown, distinct, cut off,
ocelli clear. Mouthparts well developed, red-brown; palpi brown, half
the length of the oral opening. Antennae with first segment red-brown,
second and third segments orange, third darker at tip, tip truncated.

Thorax dorsum dark red-brown; humeral and supra-alar calli red-
brown; pleural areas red-brown. Wings clear, with brown microtrichia,
veins brown; halter stem brown, knob with base and dorsal rib red-
brown, remainder white. Front leg, coxa tan, remainder of leg orange-
brown, middle and hind legs with coxae tan, remainder red-brown.

Abdomen red-brown, granular, except segments six and seven
shining, eighth segment red-brown, sternum deeply notched and 3-
pronged; see figure for details.

Maur.— Unknown.

Length: Female body 6.6 mm., wing 3.6 mm.

Type-locality: Evergreen, Colorado; 21 July 1942 (M. T. James).

Holotype: Female (USNM) 67460.

169. Pseudatrichia leechi, new species

Figure 163

FrmMa.Le.—Head black; eyes black-brown; postocular flange broad;
transversely striated; frons wider than ocellar triangle with a band of
striae passing from the anterior frontal area above antennae next to
the eye to the lateral edges of the median ocellus in a “V”’ shape,
remainder shining; ocellar triangle low, cut off, ocelli clear. Mouth-

SCENOPINIDAE OF THE WORLD 251

Ficure 162.—Pseudatrichia jamesi, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

313-168 O—69——17

952 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 163.—Pseudatrichia leechi, new species, female: a, wing; }, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum and
9th segment; f, lateral aspect of 8th and 9th segments.

parts brown, well developed, nearly filling the oral cavity, palpi two-
thirds the length of the oral opening. Antenna with first two segments
red-brown, third segment black, with short hairs that glint golden.
Thorax dorsum black, with transverse striations, humeral and
supra-alar calli red-brown, pleura largely black. Wing clear, veins

SCENOPINIDAE OF THE WORLD 253

dark brown; halter stem brown, knob with dorsal rib red-brown, re-
mainder white with some bleeding from dorsal rib. Front and middle
legs orange-brown, tarsi red-brown; hind leg red-brown, tarsus with
first segment orange, remainder red-brown.

Abdomen second segment black, granular, submetallic; membrane
black except for extreme lateral margins white; remaining segments
black, submetallic, posterior margins with large white patches on
lateral portions of membrane; eighth segment black, sternum broadly
excavated with two rounded points midlaterally; see figure for
details.

Ma.e.— Unknown.

Length: Female body 5.1-6.1 mm., wing 3.2-4 mm.

Type-locality: Mill Valley, Marin Co., California; 29 July 1948
(H. B. Leech).

Holotype: Female (CAS) 8934.

Paratypes: 4 99, Mill Valley, California, 20 September 1947,
10 May 1955, 10 June 1957, 10 July 1957 (H. B. Leech) on window in
house; 1 2, Mill Valley, September 1948 (E. S. Ross); all (CAS).

170. Pseudatrichia longiventris, new species
Ficure 164

Mauz.—Head red-brown, longer than high; eyes red-brown above,
black-brown below, with a very narrow postocular flange; frons
elongate, triangular, continuing as a thin band between the eyes to
the ocellar tubercle, swollen above antennae, granular below, shining
above; ocellar tubercle red-brown, granular; ocelli orange-red; back
of head concave in center below ocellar tubercle, swollen laterally,
dusted with gray pollen; antenna red-brown, first and second seg-
ments short, third segment oval, short.

Thorax dorsum black-brown, granular with a pollinose patch above
humeral callus; humeral and supra-alar calli red-brown, scutellum
dark red-brown; pleurum red-brown, shining. Wings brown, smoky;
veins brown, halter stem red-brown, knob red-brown. Foreleg with
femur red-brown, tibia and tarsus yellow-brown; middle leg with
femur red-brown, tibia light red-brown, tarsus yellow-brown; hind
leg with femur and tibia red-brown, tarsus yellow-brown.

Abdomen segments 2-4 dark red-brown, pebbly, with blue and
green metallic glints under light; remainder red-brown, subshining
without glints; segments 2 and 3 with white posterior membranes,
remainder with orange-brown posterior membranes, segment 7
sharply constricted near base; see figure. Ninth segment withdrawn
into seventh; for details see figure. Differs from Pseudatrichia
punctulata in the shape of the ninth tergum and aedeagal parameres.

254. U.S. NATIONAL MUSEUM BULLETIN 277

FrmaLe.—Head red-brown, longer than high; eyes dark red-brown,
postocular flange broad but abruptly ending at lateral margin of eye;
frons elevated above the antennae, narrow, no wider than the median
ocellus; ocellar tubercle prominent, not sharply cut off from frons;
ocelli red; back of head granular, subgenal and postlateral areas
extensive, shining; mouthparts red-brown, three-fourths the length
of the oral opening; palpi half as long. Antennae red-brown, the third
segment small, oval.

Ficure 164.—Pseudatrichia longiventris, new species, male, female: a, male wing; b, female
wing; c, d, lateral aspects of male and female head; ¢, dorsal aspect of female head; f,
lateral aspect of 7th, 8th, and 9th segments of male; g, lateral aspect of male terminalia;
h, posterior aspect of male terminalia; 7, ventral aspect of male terminalia; 7, ventral
aspect of female 8th sternum and 9th segment; &, lateral aspect of female 8th and 9th
segments.

Thorax dorsum red-brown, elongate, elevated into a blunt point
behind the head, depressed posteriorly in front of the scutellum;
humeral and supra-alar calli orange-brown; scutellum orange-brown ;
pleural areas orange-brown. Wings brown, smoky, long and narrow;
veins red-brown; halter stem brown, knob brown. Fore and middle
legs orange-brown; hind leg with coxa orange-brown, remainder
red-brown.

Abdomen red-brown, granular, subshining, with lateral margins
of tergites black. Eighth segment red-brown, shining, very elongate,
hairy, sternum ending in three points; see figure for details.

Length: Male body 8.5 mm., wing 4 mm.; female body 6.5 mm.,
wing 3.7 mm.

SCENOPINIDAE OF THE WORLD 250

Type-locality: Suiza, Costa Rica; July 1926 (Pablo Schild).
Holotype: Male (USNM) 67456.
Allotype: Female, same data except September (USNM).

171. Pseudatrichia longurio (Loew)
FIGURE 165
Atrichia longurio Loew, 1866, p. 42.

The type of this species was mistaken for a male, which resulted in a
serious misinterpretation of the genus by most European dipterists.
This species is one of the largest seen. The reader is referred to the
figure for pertinent details.

e.

Ficure 165.—Pseudatrichia longurio (Loew), female: a, wing; b, lateral aspect of head; c,
enlarged detail of antenna; d, ventral left and dorsal right aspect of 8th and 9th segments,
dotted lines outline a black patch on 8th tergum; ¢, lateral aspect of 8th and 9th segments.

256 U.S. NATIONAL MUSEUM BULLETIN 277

Mate.— Unknown.

Length: Female body 7.5-8.5 mm., wing 5.8 mm.
Type-locality: Mexico.

Type: (ZMH).

172. Pseudatrichia melanderi, new species

FIGuRE 166

Frmate.—Head black, longer than high; eyes black; postocular
flange wide at center; back of head narrower than the bulge of the
eyes; frons narrower than the ocellar triangle, with a median groove
and oblique striations on lower half; ocellar triangle prominent, set
off; ocelli orange. Mouthparts brown, shorter than oral cavity, palpi
half as long. Antennae orange; see figure for details.

Thorax dorsum black, humeral and supra-alar calli red-brown;
scutellum red-brown; prothorax light brown, remainder of pleura
red-brown. Wings hyaline, veins light brown; halter stem brown,
knob white. Legs orange, including coxae; hind legs slightly darker.

Abdomen segments 2—4 black, punctured, with a metallic purple
sheen. Segments 5-7 becoming red-brown. Segment 8 with tergum
red-brown, sternum orange, hairy, posterior margin two-pointed,
with a slight elevation on the excavated median portion; see figure for
details.

Ma.e.—Head red-brown, higher than long; eyes black-brown above,
black below; frons tapering to a point halfway to the median ocellus,
separating the eyes; remainder of the distance with a thin raised
ridge, lower portion with transverse striations; ocellar tubercle promi-
nent, ocelli clear; antennae orange; see figure for details.

Thorax dorsum black-brown; humeral and supra-alar calli red-
brown; scutellum red-brown; pleural areas red-brown. Wing clear,
veins brown; halter stem brown, knob white. Legs as in female.

Abdomen second segment as female, remainder lost.

Length: Male body ?, wing 4 mm.; female body 5.5-6 mm.,
wing 3.5-3.7 mm.

Type-locality: Clear Lake, California; 18 June 1935 (A. L.
Melander).

Holotype: Male (USNM) 67465.

Allotype: Female Santa Clara Co., California (Harkins Collection)
(CAS) 8925.

Paratype: 1 9, Lucerne, Lake Co., California, 17 August 1958
(Hugh B. Leech) on rotting scar wood of Umbellulasia californica
(CAS).

SCENOPINIDAE OF THE WORLD 257

Ficure 166.—Pseudatrichia melanderi, new species, male, female: a, wing; b, c, lateral
aspects of male and female head; d, dorsal aspect of female head; ¢, f, enlarged details of
male and female antennae; g, ventral aspect of female 8th sternum and 9th segment; /,
lateral aspect of female 8th and 9th segments.

258 U.S. NATIONAL MUSEUM BULLETIN 277

173. Pseudatrichia morlani, new species

Figure 167

Mauzr.—Head, red-brown; eyes red-brown above, black-brown
below; a narrow postocular flange behind eyes; frons narrow, shining,
but continuing uninterrupted to the median ocellus; ocellar triangle
prominent, ocelli clear. Antennae red-brown, last segment with
short hairs, which give a golden glint, cupped at tip.

Thorax dorsum black, metallic, transversely striated, humeral and
supra-alar calli orange-brown; scutellum red-brown; pleural areas
red-brown. Wings clear, veins brown; halter stem brown, knob white.
Forelegs yellow-brown with brown tarsi; middle legs, femora orange-
brown, tibiae red-brown basally, orange distally, tarsi brown; hind
legs with femora orange-brown basally, red-brown distally, tibiae
red-brown, tarsi, first segment orange, remainder brown.

Abdomen segments 2-4 black, rugose, with a purple metallic glint;
segments 5-7 shining, ninth tergum red-brown, generally withdrawn
into the seventh segment, posterior margin toothed dorsally and
concave on ventral half; spines of ninth sternum sparse and coarse.
Aedeagus bilobed with two median peni spurs. See figure for details.

FrmMaLe.—Head black, eyes black-brown; postocular flange smooth
and shining; frons red-brown, as wide as ocellar tubercle, with stri-
ations from the edge of the eye laterad of the antennae to the median
ocellus, a few white hairs set in these striations; frons swollen above
the antennae; ocellar tubercle flat, set off, ocelli clear; mouthparts
well developed, red-brown, filling three-fourths of the oral cavity;
palpi red-brown, half the length of oral cavity. Antennae red-brown,
third segment clothed in short hairs with a golden glint; see figure
for details.

Thorax as in male.

Abdomen black, metallic. Eighth segment red-brown, sternum
truncated; see figure for details.

This species is close to Pseudatrichia unicolor, but differs in many
ways.

Length: Male body 4.5-4.7 mm., wing 2.6-2.8 mm.; female body
4.2-4.8 mm., wing 3-3.2 mm.

Type-locality: Santa Fe Co., New Mexico; 13 January 1953
(H. B. Morlan) from Neotoma albigula nest.

Holotype: Male (USNM) 67468.

Allotype: Female, Santa Fe, New Mexico; 12 July 1952 (H. B.
Morlan) from Neotoma albigula nest, (USNM).

Paratypes: 167, Santa Fe, July (Cockerell 1377); 1, Pecos, New
Mexico, 17 June (Cockerell) at light; 3%, 6 99, 12 August, 18
November 1952, 13 and 17 January 1953, plus 7 pupal skins all with
same data as type and allotype, all (USNM).

SCENOPINIDAE OF THE WORLD 259

Ficure 167.—Pseudatrichia morlani, new species, male, female: a, wing; b, lateral aspect
of male head; c, dorsal aspect of male head; d, lateral aspect of female head; ¢, dorsal
aspect of female head; f-h, ventral, lateral and posterior aspects of male terminalia; 1,
ventral aspect of female 8th sternum and 9th segment; j, lateral aspect of female 8th and
9th segments.

260 U.S. NATIONAL MUSEUM BULLETIN 277

174. Pseudatrichia nordeni, new species

Figure 168

Frmaue.—Head black; eyes black-brown, postocular flange broad;
frons broader than ocellar triangle, swollen, with a long pit in front
of median ocellus, and with longitudinal striations that give it a dull
appearance; ocellar triangle flat, cut off; ocelli orange. Mouthparts
small, only half the length of the oral opening, palpi short. Antennae
with first two segments red-brown, third segment orange-brown,
tapering, forked at tip; see figure for details.

Thorax dorsum black, transversely striated; humeral and supra-
alar calli brown-orange; pleura dark red-brown. Wings clear, veins
brown; halter stem brown, knob with base and dorsal half brown,
ventral half white. Legs with all coxae straw colored; front legs
yellow-brown, middle legs orange-brown, hind legs red-brown.

Abdomen with segments 2-4 black, rugose, with a purple sheen;
seoments 5-7 black, shining; eighth segment black, hairy, eighth
sternum three-pointed with the median point truncated; see figure
for details.

Mauz.— Unknown.

Length: Female body 6.5 mm., wing 4 mm.

Type-locality: Norden, Nevada Co., California; 15 August 1951.

Holotype: 9? (CAS) 8935.

175. Pseudatrichia parva D. E. Hardy
Figure 169

Pseudatrichia parva D.E. Hardy, 1944a, p. 36.

This species is known only from the female, which is figured.
Length: Female body 3.7 mm., wing 2.7 mm.

Type-locality: Flagstaff, Arizona; 8 July 1941 (R. H. Beamer).
Type: (UKan).

176. Pseudatrichia parvipennis, new species

FicurE 170

Mawue.—Head black, eyes red-brown above, darker below; post-
ocular ridge very narrow; frons short, swollen, extending one-third
the distance to the ocellar triangle; ocellar tubercle raised, ocelli red;
mouthparts red-brown, short, half the length of the oral opening,
palpi shorter. Antennae orange-brown, third segment truncated; see
figure for details.

Thorax dorsum black, transversely striated; humeral and supra-alar
calli deep red-brown; pleura deep red-brown. Wings brownish, veins
brown; halter stem brown, knob with base brown, dorsal rib and

SCENOPINIDAE OF THE WORLD 261

Figure 168.—Pseudatrichia nordeni, new species, female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; ¢, ventral aspect of 8th sternum
and 9th segment; f, lateral aspect of 8th and 9th segments.

rest white. Fore and middle legs straw yellow; hind leg with basal
portion of femur straw yellow, remainder of leg red-brown.

262 U.S. NATIONAL MUSEUM BULLETIN 277

FIGURE 169.—Pseudatrichia parva,
c, dorsal aspect of head; d, enlarged detail of antenna; é,
9th segment; f, lateral aspect of 8th and 9th segments.

D. E. Hardy, female: a, wing; ), lateral aspect of head;

ventral aspect of 8th sternum and

SCENOPINIDAE OF THE WORLD 203

eG Se y
DPT BO
WR

Ficure 170.—Pseudatrichia parvipennis, new species, male: a, wing; b, lateral aspect of
head; c, enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia.

264 U.S. NATIONAL MUSEUM BULLETIN 277

Abdomen segments 2-5 black, rugose; segments 6 and 7 black,
shining; ninth tergum red-brown, pointed dorsally; aedeagus with
large lobes, but with two short wiry median peni; sternal brush of
hairs fairly thick; see figure for details.

FrmMALe.— Unknown.

Length: Male body 6 mm., wing 3.7 mm.

Type-locality: Linden, San Joaquin Co., California; 7 August 1953
(W. H. Lange).

Holotype: Male (UCalD) 74.

177. Pseudatrichia peromysci, new species

Figure 171

Maue.—Head black; eyes black-brown, black below; frons con-
tinuous to median ocellus but narrowed as eyes nearly touch three-
fifths of the way from antennae to median ocellus, widening above;
ocellar tubercle prominent, ocelli clear. Mouthparts red-brown,
three-fourths the length of the oral opening, palpi slender, half the
length of oral opening. Antennae black, third segment covered with
short pubescence that gives a golden glint.

Ficure 171.—Pseudatrichia peromysci, new species, male, female: a, wing; 0, ¢, lateral
aspects of male and female head; d, dorsal aspect of female head; e, f, enlarged details of
male and female antennae; g-i, ventral, lateral and posterior aspects of male terminalia;
j, ventral aspect of female 8th sternum and 9th segment; &, lateral aspect of female 8th
and 9th segments.

SCENOPINIDAE OF THE WORLD 265

Thorax dorsum black, transversely striated; humeral and supra-alar
calli red-brown; pleural areas black to red-brown. Wing clear, veins
dark brown; halter stem black-brown, knob with black-brown base
and dorsal rib, only a small portion of white below. Front and middle
legs, coxae to tibiae orange-brown, tarsi black-brown; hind leg, coxa
to tibia red-brown, first segment of tarsus orange-brown, grading into
black-brown.

Abdomen segments two through four black, punctured, with a
metallic sheen; remaining segments more shining. Ninth tergum
black, rounded but rough dorsally, receding ventrally; peni orange,
sternal brushes thick; see figure for details.

Fremaues.—Head black; eyes black-brown, postocular flanges
broad, shining; frons subshining, swollen above antennae, a median
cleft bordered by striations, striations on upper frons radiating
from median ocellus; ocellar triangle red-brown, not cut off, ocelli
clear. Antennae as in male.

Thorax as in male except knob of halter has much more white
than male.

Abdomen as in male; eighth sternum orange-brown, deeply concave
on posterior margin with two rounded lateral lobes; see figure for
details.

Length: Male body 4.5-5 mm., wing 2.5-3.1 mm.; female body
4.3-4.5 mm., wing 2.5-2.7 mm.

Type-locality: 3.5 mi. s., 10.2 mi. w. Arbuckle, Colusa Co.,
California; 15 January 1959 (A. M. Barnes) reared from larvae.

Holotype: o& (USNM) 67463.

Allotype: 2 same data (USNM).

Paratypes: 5077, same data as type; 107, 6.8 mi. s. 0.2 mi. w. Placer-
ville, Eldorado Co., California, 14 January 1959 (A. M. Barnes);
1 o&, 12.0 mi. s. Grass Valley, Nevada Co., California, 14 January
1959 (all collected as larvae from nests of Neotoma peromyscus and
reared out); 192, 3.5 mi. n.e. Lompoc, Santa Barbara Co., California,
9 February 1961 (A. M. Barnes) as third instar larva (AMB) and
(USNM).

178. Pseudatrichia punctulata D. E. Hardy
FIGURE 172
Pseudatrichia punctulata D. E. Hardy, 1944b, p. 104.

This species, described by Hardy, was only partially figured by him
and a complete set of figures of the male and female terminalia are in-
cluded here. The male of this species as well as P. longiventris is
notable for the sharply constricted seventh abdominal segment so
clearly illustrated by Hardy and in the sharply hooked aedeagal spines

266 U.S. NATIONAL MUSEUM BULLETIN. 277

Figure 172.—Pseudatrichia punctulata, D. E. Hardy, male, female: a, wing; ), c, lateral
aspects of male and female heads; d, enlarged detail of antenna; ¢, f, ventral and lateral
aspects of male terminalia; g, ventral aspect of female 8th sternum and 9th segment; h,
lateral aspect of female 8th and 9th segments.

they differ in the shape of the basal portion of the spines as well as in
other small details. The eyes are divided as in all Scenopinidae into
coarse upper facets and fine lower facets and the areas of union is
bandlike. The female eighth sternum is much like that of P. longiventris
but the female differs in other minor details particularly the eyes and
width of the postocular ridge.

Length: Male body 6.6-7.3 mm., wing 3.7-3.9 mm.; female body
7.5-8.3 mm., wing 4—4.3 mm.

Type-locality: Nova Teutonia, Brazil; 27°11’ S. 52°23’ W.;
20 February 1940 (Fritz Plaumann).

Type: (UKan).

179. Pseudatrichia sabroskyi, new species

FIGuRE 173

FEMALE.—Head red-brown; eyes black-brown, postocular flange,
narrow, rounded, depressed; back of head gray pubescent; frons
subshining with faint median longitudinal striations; ocellar tubercle
not prominent, cut off; ocelli orange; mouthparts a little over half
the length of the oral cavity; palpi red-brown, slender, subequal in
length to mouthparts. Antennae orange-brown; see figure for details.

Thorax dorsum black-brown, striated; humeral and supra-alar
calli red-brown tipped with orange; scutellum red-brown; pleura
red-brown. Wings clear, veins brown; halter stem brown, knob with

SCENOPINIDAE OF THE WORLD 267

Figure 173.—Pseudatrichia sabroskyi, new species, female: a, wing; b, lateral aspect of
head; c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral aspect of 8th
sternum; f, lateral aspect of 8th and 9th segments.

313-168 O—69——18

268 U.S. NATIONAL MUSEUM BULLETIN 277

base and dorsal half brown, remainder white. Front leg with coxa to
tibia straw yellow, tarsus darker toward tips; middle leg orange-
brown, hind leg red-brown.

Abdomen segment 2 dark red-brown, rugose, metallic; segments
3-5 red-brown, rugose, metallic; segments 6 and 7 red-brown, shining.
Eighth segment orange-brown, sternum with two prominent lateral
points and convex median swelling, nearly as long as the points; see
figure for details.

Matre.— Unknown.

Length: Female body 6 mm., wing 3.6 mm.

Type-locality: Antelope Mt. (6500 ft.), Harney, Lake Co.,
Oregon; 22 August 1931 (D. K. Frewing) ex Sabrosky Coll.

Holotype: Female (USNM) 67457.

180. Pseudatrichia saccharcupa, new species

Figure 174

Frema.te.—Head red-brown; eyes black-brown, postocular flange
medium wide; frons slightly swollen above the antennae, with faint
shining median groove bordered by longitudinal striations; ocellar
tubercle flat, not set off, ocelli red. Mouthparts brown; palpi red-
brown, half the length of the oral opening. Antennae with first two
segments red-brown, third segment red-orange with short hairs
glinting golden.

Thorax dorsum black, with transverse striae; humeral callus red-
brown with orange spot, supra-alar callus orange; pleural areas deep
red-brown. Wings clear, veins red-brown; halter stem red-brown, knob
base and dorsal rib red-brown, remainder white. Front leg orange
with distal tarsal segments darker; middle leg with base of femur
orange, remainder of leg red-brown; hind leg with all segments red-
brown.

Abdomen black, rugose, with a purple metallic sheen, membranes
brown on midline, white laterally; eighth segment red-brown, hairy,
eighth sternum with three points, median one shorter, rounded; see
figure for details.

Mauzr.—Unknown.

Length: Female body 6.5 mm., wing 4 mm.

Type-locality: Sugar Bowl Lodge, Placer Co., California; 26 July
1951.

Holotype: Female (CAS) 8936.

Paratype: @ same data but 19 August 1951 (CAS).

SCENOPINIDAE OF THE WORLD 269

Ficure 174.—Pseudatrichia saccharcupa, new species, female: a, wing; b, lateral aspect
of head; c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral aspect of 8th
sternum and 9th segment; f, lateral aspect of 8th and 9th segments.

270 U.S. NATIONAL MUSEUM BULLETIN 277

181. Pseudatrichia toupeta, new species

Figure 175

Ma.zr.—Head red-brown; eyes red-brown above a little darker
below, nearly meeting on upper part of frons; frons rugose above
antennae, continuing narrowly between eyes to ocellar triangle;
ocellar tubercle black-brown, ocelli orange. Antennae with first two
segments red-brown, third segment orange-brown, covered with short
hairs giving a golden glint; see figure for details.

Thorax dorsum black-brown, rugose, transversely striated; humeral
and supra-alar calli orange-brown; scutellum red-brown; pleural areas
red-brown. Wings brownish, veins brown; halter stem brown, knob
with dorsal portion light brown remainder white. Legs orange to
orange-brown.

Abdomen red-brown, rugose with purple metallic glints; ninth
tergum irregularly shaped; sternum with long black hair like a toupee,
ninth segment claspers hairy, aedeagus with two strong prongs; see
figure for details.

FrmaLe.— Unknown.

Length: Male body 6 mm., wing 3.6 mm.

Type-locality: “Colorado.”

Holotype: Male (USNM) 67467.

182. Pseudatrichia truncata, new species

Ficure 176

Mauy.—Head lost.

Thorax dorsum black, granular with fine transverse striations;
humeral and supra-alar calli red-brown; pleural areas black. Wing
clear, veins brown; halter stem brown, knob with basal portion red-
brown, distal portion white with a bleeding of brown into it. Legs
red-brown.

Abdomen second segment black with two small lateral white areas
on the intersegmental membrane; remaining segments black with
white areas on intersegmental membranes wider, central dark band
maintained. Ninth tergum brown-black; sternum with a sparse comb
of short thick hairs. Aedeagus bilobed with posteriorly directed spur-
like prongs; see figure for details.

Frma.Le.— Unknown.

Length: Male body 5.2 mm. est., wing 3.5 mm.

Type-locality: Mexico City, Mexico; 18 May (Juan Muller).

Holotype: Male (USNM) 67471.

SCENOPINIDAE OF THE WORLD 211

————

Soa)

Ficure. 175.—Pseudatrichia toupeta, new species, male: a, wing; 5, lateral aspect of head;
c, enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia.

272 U.S. NATIONAL MUSEUM BULLETIN 277

SSS

See

Figure 176.—Pseudatrichia truncata, new species, male: a, wing; b-d, ventral, lateral and
posterior aspects of terminalia.

SCENOPINIDAE OF THE WORLD 273

183. Pseudatrichia unicolor Coquillett

FIGURE 177

Pseudatrichia unicolor Coquillett, 1900, p. 500.

This species has previously been known only from the male, and
though many specimens have been identified to this species the number
remaining is limited to only a few from the immediate vicinity of the
type-locality. The female is described below.

Fremaue.—Head black-brown; eyes red-brown, postocular ridge
broad, fringed with hairs along posterior margin, transversely striated ;
frons wider than ocellar triangle, with median shining strip widest
above antennae, bordered by uniform bands of longitudinal striations

Ficure 177.—Pseudatrichia unicolor Coquillett, male, female: a, wing; b, c, lateral aspects
of male and female heads; d, dorsal aspect of female head; ¢, enlarged detail of male
antenna; f-A, ventral, lateral and posterior aspects of male terminalia; 7, ventral aspect
of female 8th sternum and 9th segment; 7, lateral aspect of female 8th and 9th segments.

from lower eye margin to median ocellus, bands with a few hairs;
ocellar triangle not elevated, set off, ocelli red-brown. Antenna with
first two segments red-brown; third segment black-brown, with a pile
of short hairs giving off a golden glint.

Thorax dorsum black-brown, transversely striated; humeral callus
red-brown, supra-alar callus orange; scutellum black with red-brown
lateral margins; pleural areas red-brown. Wings hyaline, veins brown ;
halter stem brown; knob with brown base, remainder white. Front
leg coxa through femur straw yellow, tibia yellow-brown, tarsus brown ;

274. U.S. NATIONAL MUSEUM BULLETIN 277

middle leg with coxa yellow, femur and tibia yellow-brown, tarsus
brown; hind leg with coxa orange-brown, femur red-brown, tibia brown,
first tarsal segment yellow-brown, remainder brown.

Abdomen black-brown, rugose, eighth segment red-brown, sternum
undulating along posterior margin, lateral lobes shallow; see figure for
details.

Length: Male body 6-6.3 mm., wing 4 mm.; female body 6.5
mm., wing 4.3 mm.

Type-locality: Las Vegas, New Mexico; July 19—(T. D. A.
Cockerell).

Type: (USNM) 4711.

Other specimens seen: 16, 19, Las Vegas, New Mexico, 3, 11
August (H.S. Barber); 1, Watrous, New Mexico, Cockerell no. 2503,
all (USNM).

Genus Belosta D. E. Hardy

Belosta D. E. Hardy, 1944a, p. 37.

Type-species Belosta albipilosa Hardy (original designation). The
genus Belosta D. KE. Hardy is closely related to the genus Pseudatrichia
from which it was removed because of the head being higher than long,
atrophied mouthparts and generally hairy condition. It has a closed
cell R5. The members of this genus are found in the western mountain
area of the United States where the larvae are predators of wood
attacking insects.

Key to the Species of Belosta

1. Males, eyes close together
Females, eyes widely separated .

2. Halter knob white SU ecaestiel achioucan EM RET MUMIA Me Ee WMG WUT ay sage eto ater
Halter knob tan, thorax with short white hairs . . . . B. albipilosa Hardy

3. Halter knob all white, thorax with long white hair. . B. pilosa (Coquillett)
Halter knob white above with a tan rim and ventral surface, thorax with

oe bo

0

very short scale-like hairs .... . . . .B. viticolapennis, new species
4;;Halter! knob: orange :.«) i ageciss) atts |e bee sack GE ie eto eee eo
Halter knob red-brown .. . aware KO

5. Thorax with mixed short red and white hairs, 7th sternum short, abrupt
B. albipilosa Hardy

Thorax with short white hairs only, 7th sternum long, sloping
B. pilosa (Coquillett)
6. Halter knob red-brown except ventral tip white. . . B. telfordi, new species
Halter knob red-brown above cream below . . . B. termitophaga, new species

SCENOPINIDAE OF THE WORLD 275

184. Belosta albipilosa D. E. Hardy
Figure 178
Belosta albipilosa D. E. Hardy, 1944a, p. 38.

This species, an active predator of wood boring insects, is adequately
described by Hardy. Sclerotized portion of female accessory gland
seven times as long as wide.

Length: Male body 4-5.5 mm., wing 3-3.8 mm.; female body
5-6 mm., wing 3.5-4 mm.

Type-locality: Riggins, Idaho; 18 May 1940 (T. Terrell) taken on
Pinus ponderosa.

Type: 2? (USNM).

I have seen all of the type-series in the (USNM), as well as material
from the following localities: 1 9, Wawona, California, on Thuja; 1 3,
Whitaker Forest, Sequoia Nat. Forest, California, 21 October 1940
(D. DeLeon) on Quercus kelloggi; 1 2, Stanislaus Nat. Forest, Califor-
nia, 27 May 1935 (W. H. Lange) on Pinus ponderosa; 1 9, Sierra Nat.
Forest, California (J. W. Bonberg) on Pinus ponderosa; 2 3A,
Bandelier, New Mexico, 1957; 1 9, Krassel, Idaho, 18 September 1957
(M. M. Furniss) on Pseudotsuga menziesii; all in (USNM); 1 @,
Junipero Sierra Pk., Santa Lucia Mts., Monterey Co., California, 3
July 1957 (H. B. Leech) on peak ca. 5800 ft. emerged from dead
branches Pinus coultert in (CAS); 4077, 19, Krassel, Idaho, emerged
January and 16 and 27 June 1960 (M. M. Furniss) ex Pseudotsuga
menziesii where they were found to be predaceous on the larvae of
Dendroctonus pseudotsugae, Hopk., U.S. Forest Serv. Res. Lab.,
Boise, Idaho.

185. Belosta pilosa (Coquillett)
FiGureE 179

Pseudatrichia pilosa Coquillett, 1902, p. 102.
Pseudatrichia flaviceps Coquillett, 1902, p. 102. New synonymy.

After a good deal of study on the paratypes of these two species as
well as considering where and when captured, I have concluded that
B. flaviceps is the female of B. pilosa. The wing venation corresponds
as closely as in most species; the presence of short but rather dense
pile on the female, the comparable size, etc., make it seem improbable
for them to be separate species. I did not observe the black hairs on
the male mentioned by Coquillett in his description. A beautiful male
was received, from Dr. Timberlake, which had the entire thorax and
much of the abdomen covered with long white hairs, giving it a truly
snowy appearance. Accessory gland of female has a flaring base,
sclerotized portion only as long as broad.

Length: Male body 4-5.5 mm., wing 3.6-4.5 mm.; female body
6 mm., wing 4.6 mm.

276 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 178.—Belosta albipilosa D. E. Hardy, male, female: a, wing; 0, c, lateral aspects of
male and female heads; d, frontal aspect of female head; e, f, enlarged details of male and
female antennae; g, h, ventral and lateral aspects of male terminalia; 7, dorsal aspect of
female 8th and 9th segments; j, ventral aspect of female 8th and 9th segments; &, lateral
aspect of female 8th and 9th segments; /, spermatheca.

xo
Ficure 179.—Belosta pilosa (Coquillett), male, female: a, wing of male; b, wing of female;
c, d, lateral aspects of male and female heads; e, f, frontal aspects of male and female
heads; g, h, enlarged details of male and female antennae; 7, 7, ventral and lateral aspects
of male terminalia; &, ventral left and dorsal right aspect of female 8th and 9th segments;

J, lateral aspect of female 8th and 9th segments; m, spermatheca.

SCENOPINIDAE OF THE WORLD Pt

278 U.S. NATIONAL MUSEUM BULLETIN 277

Type-locality: Williams, Arizona, 7 July (H. 8S. Barber) ; female as
Pseudatrichia flaviceps, Williams, Arizona, 15 July (H. S. Barber).

Types: (USNM) 6194 and 6193.

Specimens examined: 1c’, 19, paratypes of the above specimens
(USNM); 1 o&, Mannix California, 25 April 1951 (Timberlake)
(PHT); 1 o&, Strawberry Valley, San Jacinto Mts., California,
12 July 1912 (J. C. Bridwell) (USNM).

186. Belosta telfordi, new species
Fiaure 180

FremaLe.—Head brown; eyes black-brown; frons dark brown,
ocellar tubercle red-brown, mouthparts atrophied, palpi bulbous. An-
tennae black-brown; see figure for details. (The head was collapsed
so was removed, relaxed in KOH, and replaced on the specimen.)

Thorax dorsum black-brown, with medium long white hairs in
scattered patches; humeral callus black-brown; scutellum_ black-
brown. Pleural areas prothorax red-brown, mesanepisternum black-
brown with white hairs, katepisternum black-brown. Wings hyaline,
veins brown; halter stem red-brown, knob red-brown except ventral
tip white. Foreleg with femur black-brown with yellow-brown dorsal
surface and knee, remainder of leg black-brown; middle and hind legs
black-brown, femora with yellow knee.

Abdomen black-brown, intersegmental membranes with black-
brown midline and white lateral portions. Eighth sternum long,
sloping; accessory gland short with flaring base, sclerotized portion
twice as long as broad; see figure.

Mauzr.— Unknown.

Length: Female body 6.5 mm., wing 4.5 mm.

Type-locality: Tucson, Arizona; 17 March 1957 (A. D. Telford).

Holotype: Female (USNM) 68005.

187. Belosta termitophaga, new species
Fieure 181

FremMaLe.—Head cream; eyes dark red-brown; frons very broad,
tan, shining, with two longitudinal crescent-shaped marks, a brown
patch on gena below the base of the eye; head behind the eye cream,
center of back of head brown; mouthparts atrophied, palpi bulbous;
ocellar tubercle red-brown, ocelli clear. Antennae with first segment
short, brown; second segment tan; third segment brown, tapering,
over three times as long as 1 and 2 combined.

Thorax dorsum black, with two anterior tan patches above the
humeral callus, covered with medium long hairs, humeral and supra-
alar calli tan, scutellum red-brown. Propleura tan, mesopleural areas

SCENOPINIDAE OF THE WORLD 279

Ficure 180.—Belosta telfordi, new species, female: a, wing; b, lateral aspect of head; c»
frontal aspect of head; d, enlarged detail of antenna; f, ventral left and dorsal right aspect
of 8th and 9th segments; ¢, lateral aspect of 8th and 9th segments; g, spermatheca.

280 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 181.—Belosta termitophaga, new species, female: a, wing; b, lateral aspect of head;
c, frontal aspect of head; d, enlarged detail of antenna; e, ventral left and dorsal right
aspect of 8th and 9th segments; f, lateral aspect of 8th and 9th segments; g, spermatheca.

SCENOPINIDAE OF THE WORLD 281

red-brown with white hairs. Wing hyaline, veins brown; halter elon-
gate, stem red-brown, knob red-brown above, cream below. Legs with
yellow-brown femora, tan tibiae, darker tarsi.

Abdomen red-brown with white intersegmental membranes which
are dark along the midline. Eighth segment red-brown, sternum long,
sloping with a trilobed posterior margin. Accessory gland with a flat
flange and only about twice as long as wide; see figure for details.

Mauze.— Unknown.

Length: Female body 5.6-6.5 mm., wing 4.4-5.5 mm.

Type-locality: Tucson, Arizona; 12 April 1924 (A. A. Nichol)
from Kalotermes hubbardi nest, reared.

Holotype: Female (USNM) 68004.

Paratypes: 392 9, same data as type; 12, Tucson, Arizona, 13
April 1925 (C. T. Vorhies) from Kalotermes hubbardi nest. One para-
type (USNM), remainder (UAriz).

188. Belosta viticolapennis, new species

FIGURE 182

Maur.—Head red-brown; eyes brown above, black-brown below;
frons cream with a tan triangle above the antenna, area between oral
opening and eyes cream; mouthparts atrophied; palpi brown, bulbus;
ocellar triangle prominent, ocelli clear; antennae light brown, shorter
than other species, third segment diamond-shaped, over three times as
long as first two segments combined; see figure for details.

Thorax dorsum red-brown, shining, with very short scalelike hairs,
humeral and supra-alar calli cream; pleural areas red-brown. Wings
hyaline, veins brown, R4 branching narrowly from R5 before curving
to edge of wing; halter stem brown, knob white with tan rib and ventral
surface. Legs: femora brown with cream knees, remainder brown.

Abdomen segments red-brown with white posterior membranous
borders. Ninth tergum blocky, sternal spines sparse, stiff. Aedeagus
with parameres extremely long and tendril-like; see figure for details.

FrmaLe.— Unknown.

Length: Male body 3 mm., wing 2.2 mm.

Type-locality: Wiliams, Arizona; 17 July (H. S. Barber).

Holotype: Male (USNM) 68006.

Genus Propebrevitrichia, new genus

Type-species, Propebrevitrichia turneri, new species. The genus
Propebrevitrichia closely resembles the genus Brevitrichia; it differs in
the point of branching of vein R4 which forks from beyond the middle
of the closed cell R5 and by the elongate pointed female eighth ster-
num which is longer than the tergum. The members of this genus are
found in the Ethiopian region.

282 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 182.—Belosta viticolapennis, new species, male: a, wing; }, lateral aspect of head;
c, enlarged detail of antenna; d-f, ventral, lateral, and posterior aspects of terminalia.

SCENOPINIDAE OF THE WORLD 283

Key to Species of Propebrevitrichia

alent a RR: See MMe cB, ig ve ct at. om Ok ee SOR

Female frons broad, twice the width of the ocellar triangle, abdomen red-

brown covered with tan scales; segments 1—5 with white posterior bands;

8th sternum elongate, orange-red, exceeding the dorsum; 9th tergum with

3 stiff spines. ............... P. turneri, new species

2. Thorax black-brown dusted gray pollinose; abdomen dark red-brown, dusted

gray pollinose, posterior membranes white or tan; 9th tergite with basal
third dark remainder white with two dark spots, all dusted gray pollinose

P. stuckenbergi, new species

Thorax red-brown, covered with golden brown scales bordered on either side

of the midline by two brown bands, abdomen red-brown covered with tan

seales, half of segments 2 and 3 white, remainder of segments white dor-

sally; ninth tergum with basal portion brown, covered with brown scales,

remainder white ............. . .P. turneri, new species

189. Propebrevitrichia stuckenbergi, new species
Figure 183

Mae.—Head black-brown; eyes light brown above, dark brown
below; frons dusted gray pollinose, wide at the base narrowing rap-
idly, a shallow pit just below the halfway point between the ocellar

Figure 183.—Propebrevitrichia stuckenbergi, new species, male: a, wing; b, lateral aspect
of head; c, frontal aspect of head; d, enlarged detail of antenna; e, f, ventral and lateral
aspects of terminalia.

triangle and antennae; ocellar triangle black-brown, dusted gray pol-

linose, large and prominent; ocelli dark; back of head dusted gray

pollinose; mouthparts well developed, rostrum long and_ slender,
313-168 O—69-—19

284 U.S. NATIONAL MUSEUM BULLETIN 277

flabellum not expanded; palpi short. Antennae black-brown, first
segment short, second twice as long and wide as first, third segment
pear-shaped with a bitruncated terminal portion with a median peg;
see figure for details.

Thorax dorsum black-brown, dusted gray pollinose; humeral callus
brown with a cream lateral margin; scutellum black-brown; pleura
black-brown. Wings milky white, veins tan along anterior portion
nearly white behind; halter stem brown, knob immaculately white.
Legs all black-brown dusted pollinose, with tan joints.

Abdomen second segment dark red-brown dusted pollinose, with a
narrow white or tan posterior membranous margin, remaining seg-
ments dark red-brown dusted pollinose, with large crescents of white
membrane on dorsal portion occupying over half the length of the
segment, sterna sclerotized. Ninth tergum with dark brown basal
third, remainder white with two brown spots at about the midpor-
tion, aedeagus not exposed; see figure for details.

FrmaLe.— Unknown.

Length: Male body 2.2 mm., wing 1.7 mm.

Type-locality: Karroo, at Junction of Calvinia—Sutherland Rd.
near Inverdoorn Ceres, w. Cape Prov., [Republic of] South Africa;
23 October 1959 (B. and P. Stuckenberg).

Holotype: Male (NMP).

190. Propebrevitrichia turneri, new species

Figure 184

Matxr.—Head brown; eyes tan above, light brown below; frons
small, triangular, with a median groove, covered with light brown
scales; back of head convex laterally, covered with tan scales; genae
covered with brown scales and long hairs; mouthparts brown, long;
palpi tan, half the length of the rostrum; ocellar tubercle dark brown,
covered with tan scales; ocelli orange. Antennae with first two seg-
ments brown, short, second twice as large as the first; third segment
red-brown, latero-basal portion orange, covered with scales, ending
in two points.

Thorax red-brown, covered with golden brown scales, with two
brown bands laterad of the midline; humeral and supra-alar calli tan;
pleural areas red-brown, covered with tan scales. Wings smoky,
iridescent when reflecting light; veins tan; halter stem tan, knob
white. Legs with femora red-brown, covered with tan scales, knees
tan; tibiae and tarsi red-brown with tan scales.

Abdomen second segment red-brown, covered with tan scales, a
membranous white posterior border; third segment with anterior
half red-brown scaled, posterior half white, membranous; fourth
segment nearly all white, remaining segments white; ninth tergum

SCENOPINIDAE OF THE WORLD 285

white with basal portion brown scaled, aedeagal prongs not exceeding
side pieces; see figure for details.

FrmaLe.—Head red-brown; eyes orange-brown; postocular ridge
narrow; frons broad, swollen, with a median depression extending
between antennae and median ocellus, with scattered hairs; genal
areas to back of head yellow; other areas covered with golden scales.
Ocellar tubercle covered with golden scales, set off from frontal area;
ocelli red-brown. Antenna as in male.

Thorax as in male.

Abdomen red-brown covered with tan scales segments three to
five with posterior bands of white remainder without, eighth segment
orange-red, subshining, elongate; sternum longer than tergum, coming
to a point; ninth tergum with three stiff spines and a few hairs; see
figure for details.

Ficure 184.—Propebrevitrichia turneri, new species, male, female: a, wing; b, d, lateral
aspects of male and female heads; c, ¢, frontal aspects of male and female heads; f, g,
enlarged details of male and female antennae; h, i, ventral and lateral aspects of male
terminalia; 7, ventral left and dorsal right aspect of female 8th and 9th segments; k,
lateral aspect of female 8th and 9th segments.

Length: Male body 2.5 mm., wing 1.8 mm.; female body 2.8 mm.,
wing 1.9 mm.

Type-locality: Mossel Bay, s. Cape Prov., [Republic of] South
Africa; 4 December 1938 (R. E. Turner).

Holotype: Male (BMNH).

Allotype: Female Mossel Bay, s. Cape Prov., [Republic of] South
Africa; 20 October 1938 (R. E. Turner).

I86 U.S. NATIONAL MUSEUM BULLETIN 277

Paratypes: 7070", 52 9, Mossel Bay, s. Cape Prov., [Republic of]
South Africa, October, December 1938 (R. E. Turner) ; 1 2 , Worcester,
s.w. Cape Prov., [Republic of] South Africa, December 1933 (R. E.
Turner) all (BMNH).

Genus Heteromphrale Krober
Heteromphrale Krober, 1937, p. 221.

Type-species Pseudatrichia chilensis Krober (original designation).
The genus Heteromphrale is closely related to the preceding genus and
Brevitrichia but may be distinguished from Brevtrichia by the elongate
female eighth sternum and from Propebrevitrichia by the basal forking
of vein R4. The male genitalia is more robust than in the other two
genera. This genus is Neotropical in distribution.

191. Heteromphrale cyanops (Edwards), new combination
Figure 185

Pseudomphrale cyanops Edwards, 1932, p. 259.

This species is distinguished by the wing venation which is similar
to Brevitrichia and by the much heavier ninth tergum, short peni and
aedeagal parameres, and larger heavier internal structures associated
with the aedeagal syringe; see figure for details.

Length: Male body 2.5 mm., wing 2 mm.

Type-locality: Montevideo, Uruguay; 21 January 1927 (F. and
M. Edwards).

Type: (BMNH).

192. Heteromphrale chilensis (Kréber)
FIGurRE 186

Pseudatrichia chilensis Kréber, 1928b, p. 31.
Heteromphrale chilensis (Kréber), Kréber, 1937, p. 221.

This species is recognized on the basis of the wing venation which
is similar to that of Brevitrichia and on the basis of the spines on the
ninth tergum. The long eighth sternum separates it from species of
Brevitrichia; see figure for details.

Length: Female body 5 mm., wing 2.6 mm.

Type-locality: Concepcién, Chile (P. Herbst).

Type: (ZMH).

Genus Prepseudatrichia, new genus

Type-species Prepseudatrichia mateui, new species. The genus
Prepseudatrichia is most interesting in that the head and_ thorax
are distinctly Pseudatrichia-like, while the wings are typical of
Scenopinus. Both the male and female genitalia are distinctive.

SCENOPINIDAE OF THE WORLD 287

Ficure 185.—Heteromphrale cyanops (Edwards), male: a, wing; b, lateral aspect of head;
c, frontal aspect of head; d, lateral aspect of antenna; e-g, ventral, lateral and posterior
aspects of terminalia.

288 U.S. NATIONAL MUSEUM BULLETIN 277

ie

Ficure 186.—Heteromphrale chilensis (Krdber), female: a, wing; }, lateral a spect of head;
c, frontal aspect of head; d, enlarged detail of antenna; e, dorsal aspect of 8th and 9th
segments, sternum and bursal structures outlined; f, lateral aspect of 8th and 9th segments.

This genus has been reared from wood borers attacking Acacia
trees and is Ethopian in distribution.

SCENOPINIDAE OF THE WORLD 289

Key to the Species of Prepseudatrichia

1. Females ... Bae eee ay se hgh wees a eee epee

Males, halter feeb Bibi: row n, Phere black, striated, legs straw yellow

darkening to orange-brown ee .... . . P. mateui, new spccies

2. Thorax black, striated ..... 2% Faro
Thorax black with green and mnedte qicatine rates halter rane ode brown,
velvety, legs straw yellow with tarsi red-brown. . P. violacea, new species

3. Halter knob mahogany-brown, legs reddish, antennae orange, cell Cu ending
abruptly. . . ... . .P. stenogaster Séguy

Halter knob red- Baws n m Billick, Stans, legs straw yellow darkening to
orange-brown posteriorly, cell Cu long ending in a point
P. mateui, new species

193. Prepseudatrichia mateui, new species

FIGURE 187

Ma.e.—Head black, longer than high; eyes red-brown above,
black-brown below, close but not meeting on the midline; frons very
narrow, slightly swollen above the antennae, a median groove above;
ocellar tubercle prominent; ocelli clear; a white patch above the
antennal bases; oral opening large; mouthparts cream, well developed,
with long hairs on flabellum; palpi long and slender. Antenna black-
brown; first segment short; second longer than first; third granular,
truncated not twice as long as wide; see figure for details.

Thorax black, striated; dorsoventrally compressed, particularly in
prescutal area; a pollinose patch above the humeral callus; humeral
callus red-brown, narrow; pleural areas black-brown, shallow, the
front and middle legs more widely separated than normal. Wings
hyaline, veins brown, similar to Scenopinus; halter stem brown to
black-brown, knob red-brown to black, shining. Front leg straw
yellow with brownish yellow tarsus darkening distally; middle leg
with coxa and femur straw yellow tibia and tarsus brownish yellow
darkening distally; hind leg with all segments orange-brown, first
tarsal segment yellowish, rest red-brown.

Abdomen second segment red-brown to black, segments three and
four with white posterior bands extending to lateral margins. Ninth
segment red-brown to black, hairy. Both external and internal struc-
tures separate this species from all others; see figure for details.

Frematy.—Head black, rugose, longer than high; eyes yellow-brown;
with prominent postocular ridge, subshining; frons broad, rugose, with
a bulge above the antennae, a white patch above the antennal base,
wider than the ocellar triangle which is not prominent; ocelli brown,
mouthparts as in the male. Antennae black-brown, third segment
pear-shaped, darker at tip; see figure for details.

Thorax as in male; legs as in male but darker.

290 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 187.—Prepseudatrichia mateui, new species, male, female: a, wing; }, c¢, lateral
aspects of male and female heads; d, ¢, dorsal aspects of male and female heads; f, g,
enlarged details of male and female antennae; h, i, ventral and lateral aspects of male
terminalia; j, ventral aspect of female 8th sternum and 9th segment; &, lateral aspect of
female 8th and 9th segments; /, female 9th sternum and bursa, floor of cavity outlined.

Abdomen red-brown to black. Eighth segment elongate, tip of
sternum and ninth tergum with a few long hairs; bursa with a large
oval membranous area; see figure for details.

SCENOPINIDAE OF THE WORLD 291

Length: Male body 5 mm., wing 2.3 mm.; female body 4.5 mm.,
wing 2.3-2.7 mm.

Type-locality: Temba, Rgueibat, Sahara; 1 August 1958 (J.
Mateu) emerged in the laboratory from a tunnel in the wood of
Acacia raddiana Sgn. parasitized by wood-boring larvae.

Holotype: Male (MNHN).

Allotype: Female, Oued el Arfejd, Sahara Occ.; 11 April 1962
(J. Mateu) emerged in the laboratory, also from tunnels in wood of
Acacia raddiana.

Paratypes: 1 co, Mare (=pond) de Biliharo Ennedi (Mourdi),
North Chad, 12 January 1959 (J. Mateu); 1 o, Oued Trick (Osugarta
Mountains), 7 April 1961 (J. Mateu) parasite of Anthazia angusti-
pennis (Col. Buprestidae); deposited in (MNHN); 1 o, Beni Abbes,
Sahara (Algeria), 14 May 1962 (J. Mateu) retained by me (USNM).

All of these flies were reared from branches of Acacia which con-
tained wood-boring larvae, mainly Buprestidae, Bostrichidae and
Cleridae.

194. Prepseudatrichia stenogaster (Séguy), new combination
FiGuRE 188
Scenopinus stenogaster Séguy, 1931, p. 114.

This species, originally described in the genus Scenopinus, resembles
that genus only in the character of the wing, the head is longer than
high, the thorax dorso-ventrally compressed and the round slim
abdomen agrees with Pseudatrichia in appearance. It is placed in this
new genus on the basis of agreement with females of the other species;
see figure for details.

Length: Female body 5.5 mm., wing 2.25 mm.

Type-locality: Nova Choupanga prés Chemba, Zambezi, Mozam-
bique; July 1929 (P. Lesne) collected as a larva from the trunk of
Sterculia, pupated 6 July, adult emerged mid-July.

Type: (MNHN).

195. Prepseudatrichia violacea, new species

FiGurE 189

Frmate.—Head blackish brown, longer than high; eyes light brown,
probably darker in life; postocular ridge moderately wide, prominent;
frons narrow, only as wide as ocellar triangle, rugose, lower along
eye margin than center, reflecting purple glints, a flattened area
above antennae ending in a point one-third the distance to the
median ocellus. Ocellar triangle prominently set off. Mouthparts
light brown, palpi tan, well developed filling the oral cavity. Antennae
with first two segments dark brown, short; third segment lost.

292 U.§. NATIONAL MUSEUM BULLETIN 277

Ficure 188.—Prepseudatrichia stenogaster (Seguy), female: a, wing; b, lateral aspect of head;
c, dorsal aspect of head; d, enlarged detail of antenna; e, ventral left and dorsal right aspects
of 8th and 9th segments; f, lateral aspect of 8th and 9th segments; g, 9th sternum and
bursa, floor of cavity outlined.

SCENOPINIDAE OF THE WORLD 293

Ficure 189.—Prepseudatrichia violacea, new species, female: a, wing; b, lateral aspect of
head; c, dorsal aspect of head; d, ventral aspect of 8th sternum and 9th segment; e¢,
lateral aspect of 8th and 9th segments; f, 9th sternum and bursa, with floor of cavity
outlined.

294. U.S. NATIONAL MUSEUM BULLETIN 277

Thorax compressed, dorsum black with brilliant green and purple
metallic glints, prescutellar area depressed, lateral margins of dorsum
and pleural areas with sparse, white bulbous hairs particularly dense
above the humeral callus; humeral and supra-alar calli light brown;
scutellum black with purple glints. Wings hyaline, veins brown;
halter stem brown, knob red-brown, velvety. Front and middle legs
with femora and tibiae yellow, tarsi red-brown; hind legs missing.

Abdomen segments red-brown, eighth segment elongate, tapered;
see figure for details of sternum and bursal plate.

Maxzr.— Unknown.

Length: Female body 4 mm., wing 2.3 mm.

Type-locality: Temba, Tagant Mauriton (Chad), (J. Mateu) on
Acacia scorpiodes.

Holotype: Female (MNHN).

Genus Seguyella, new genus

Type-species Omphrale galactica Séguy. The genus Seguyella re-
sembles members of the genus Scenopinus but differs in venation of the
wing, male and female terminalia, and in having a notched third
antennal lobe. Vein R5 extending straight from the cross vein to before
the tip of the wing and vein M1 + 2 parallel or slightly diverging to end
in the wing margin well behind the tip of the wing. Vem M3 + Cul
short or missing beyond m cross vein. The members of this genus are
found in the Ethiopian and Oriental regions.

Key to the Species of Seguyella

1. Males, eyes contiguous along midline, wingshyaline. ......... .2
Females, eyes separated by broad frons, wings brown. ........ a

2. Halter knob brown, elongate, abdomen with dorsal portion of segments 3-7
white, terminalia black <0 etismeou 1 fais lence S. galactica (Séguy)

Halter knob red-brown, shining, rounded, abdomen with dorsal portion of
third segments red-brown, segments 4—7 white, terminalia red-brown

S. turneri, new species

3. Thorax black covered with small flat gray scales, halter knob orange-brown,

wing black-brown, abdomen red-brown covered with flat gray scales (India)

S. chinchona, new species

Thorax red-brown pollinose, halter knob tan pollinose, wing smoky brown,
abdomen red-brown, granular. ........ S. turneri, new species

196. Seguyella chinchona, new species

Figure 190

Frema.e.—Head black, higher than long; eyes black-brown; frons
shining, width of ocellar triangle below and flaring dorsally around
ocellar triangle to back of eyes; ocellar tubercle black, pollinose with
red-brown ocelli; oral opening covered with silvery pile which extends

SCENOPINIDAE OF THE WORLD 295

Ficure 190.—Seguyella chinchona, new species, female: a, wing; b, lateral aspect of head;
c, frontal aspect of head; d, enlarged detail of antenna; ¢, dorsal aspect of 8th and 9th
segments; f, ventral aspect of 8th and 9th segments; g, lateral aspects of 8th and 9th
segments.

296 U.S. NATIONAL MUSEUM BULLETIN 277

over the antennal bases, mouthparts brown, filling the oral cavity;
palpi two-thirds as long as mouthparts; back of head concave. An-
tennae red-brown; first two segments short subequal; third segment
pilose, cupped on end, less than twice as long as wide; see figure for
details.

Thorax dorsum black, covered with small flat gray scales, humeral
and supra-alar calli red-brown; pleural areas dark red-brown. Wings
black-brown, very dark, veins black-brown, M1 + 2 ending behind the
tip of the wing and M3 + Cul ending at the m cross vein; halter stem
red-brown, knob orange-brown. Legs orange-brown.

Abdomen red-brown covered with flat gray scales dorsally, eighth
segment with dorsum much longer than sternum, sternum with a
brush of long hairs from proximal margin, distal margin turning
upward. Ninth segment pendulous, hairy basally; see figure for
details.

Maztre.—Unknown.

Length: Female body 3.2 mm., wing 2.4 mm.

Type-locality: Chinchona, Anamali Hills, S. India; (8500 ft.)
May 1950 (P. S. Nathan).

Holotype: (CNC) 8630.

197. Seguyella galactica (Séguy), new combination
FicurE 191

Omphrale galactica Séguy, 1938, p. 334.

This species can be distinguished by its peculiar wing venation,
the sharply pointed ninth terga and the broadly flaring tenth sternum;
the penis consists of two short straight prongs with short constricted
tips; see figure for details.

Length: Male body 1.8 mm., wing 2.2 mm.

Type-locality: Elgon Saw Mill, Camp II on the east slope, Mt.
Elgon (2470 m.), Kenya (C. Aranbourg, P. A. Chappius and
R. Jeannel).

Type: (MNHN).

198. Seguyella turneri, new species

FIGURE 192

Frmate.—Head orange-brown; eyes red-brown; frons red-brown,
broad and shining, with a lighter median triangle, widest at the base
and narrowing to the median ocellus; frons continuous with postocular
flange and distance from eye margin to ocellar tubercle nearly as
wide as tubercle itself; ocellar tubercle orange-brown, pollinose;
ocelli red-brown. Mouthparts well developed; antennae red-brown,
first segment short; second twice the size of first; third twice as long as

SCENOPINIDAE OF THE WORLD 297

Ficure 191.—Seguyella galactica (Séguy), male: a, wing; b, lateral aspect of head; c, anterior
aspect of head; d, enlarged detail of antenna; e-g, ventral, lateral and posterior aspects of
terminalia.

298 U.S. NATIONAL MUSEUM BULLETIN 277

Ficure 192.—Seguyella turneri, new species, male, female: a, wing; b, lateral aspect of male
head; c, enlarged lateral aspect of female head; d, enlarged frontal aspect of female head;
e, dorsal aspect of female 8th and 9th segments; f, ventral aspect of female 8th and 9th
segments; g, lateral aspect of female 8th and 9th segments.

SCENOPINIDAE OF THE WORLD 299

first two combined, covered with short hairs, ending in two points;
see figure for details.

Thorax dorsum red-brown, pollinose; humeral callus orange;
scutellum orange-brown, pollinose; pleura red-brown. Wings smoky
brown, veins brown, faint posteriorly; vein M1 -+ 2 ending behind
the tip of the wing, vein M3 + Cul ending just beyond the m cross
vein; halter stem orange-brown, knob tan, pollinose. Legs light
red-brown throughout.

Abdomen red-brown, granular; eighth tergum and sternum sub-
equal, sternum with a basal band of long hairs. Ninth tergum four-
lobed, drooping; see figure for details.

Maure.—Very small, head black-brown; eyes red-brown, darker
below; frons small triangular, red-brown above antennae darkening
to black-brown; ocellar tubercle black-brown, prominent, granular.
Antennae with first two segments red-brown, first small, second
swollen, third segment dark red-brown swollen above and ending in
two points; see figure for details.

Thorax dark red-brown. Wings hyaline, veins light tan, faint
posteriorly; halter stem red-brown at base, cream below the knob
which is red-brown and shining, very round. Legs uniform red-brown
throughout.

Abdomen second segment red, segments three through six broadly
banded with white (four bands), ninth tergum red-brown. The
terminalia were lost on dissection so cannot be illustrated.

Length: Female body 2.5 mm., wing 2 mm.; male body 1.25 mm.,
wing 1.1 mm.

Type-locality: Mossel Bay, Cape Prov., [Republic of] South
Africa, January 1922 (R. E. Turner).

Holotype: Female (BMNH).

Allotype: Male, same data.

Genus Neopseudatrichia, new genus

Type-species Neopseudatrichia kewi, new species. The genus
Neopseudatrichia is closely related to Pseudatrichia which it resembles
except for the male and female terminalia. The male ninth tergum
short, not concealing the tenth segment, no comb of sternal hairs
as in Pseudatrichia, aedeagus consists of a short median penis flanked
by two short parameres, eighth sternum of female prolonged on the
midline, ninth tergum with a tuft of stiff spines. This genus is limited
to the Australian region.

Key to the Species of Neopseudatrichia

ee CYVISIC SMM 305) 8 eee eo es eae ay ye las Ae a eal cues dee nce wnt 2
NISICHMEI Coe ar fy sc) a Suse ee te ee eee es a ale
313-168 O—69 20

300 U.S. NATIONAL MUSEUM BULLETIN 277

2. Wing hyaline, halter knob black, stalk of vein R5+M1-+2 extending to
wing tip on the line of M1+2... .. . N. mariaensis (G. H. Hardy)

Wing hyaline, dusted brown granular, halter knob brown, elongate, granular,
stalk of vein R5+M1+2 more nearly on a line with R5 to tip of
Wings. fs ; BOs Pinte eae apes ...... . N. kewi, new species

3. Wing brown eoamalie snot Scee Se uelitre ree tee 4
Wing hyaline, halter knob finde stalk of vein Ree M1+2 extending foe tip

of wing on the line of M1+2. . ... . . N. mariaensis (G. H. Hardy)

4. Halter knob red-brown, egg-shaped, stalk of vein R5-++ M1+2, extending to
tip of wing on the line of vein R5, legs red-brown . . N. canei, new species
Halter knob brown, granular, elongate pointed, vein R5-+ M1+2 extending

to tip of wing, not on a direct line with vein R5, femora black-brown,
remainder lighter 2.0. 2 3/ss) a) Yaa e Ne kewlsmew-species

199. Neopseudatrichia canei, new species

FIGURE 193
Pseudatrichia mariaensis Paramonov (not Hardy), 1955, p. 639.

Maue.—Head black-brown, longer than high; eyes black-brown:>
darker below, occupying most of head from the side view; frons
shining, two fine rows of setae converging toward the midline from
antennal bases to middle of frontal area; eyes separated by width of
the median ocellus; ocellar tubercle prominent, black-brown, granular;
ocelli black; antennae with first two segments black-brown, first
twice as long as second, third segment missing; see figure.

Thorax dorsum black-brown, shining, with sparse white hairs;
humeral and supra-alar calli red-brown; scutellum red-brown, shining;
pleural areas red-brown, polished. Wings hyaline, with brown granu-
losis, veins brown; halter stem brown, knob red-brown, bluntly
rounded, egg-shaped. Legs red-brown.

Abdomen segments black-brown with narrow white bands posteri-
orly ; ninth tergum black-brown, shorter than tall, tenth segment much
longer; aedeagus with two blunt lobes; see figure for details.

FremaLe.—Unknown.

Length: Male body 4.2 mm., wing 2.4 mm.

Type-locality: Mt. Lofty (2334 ft.), South Australia; 15 January
1952 (H. M. Cane).

Holotype: Male (CSIRO).

200. Neopseudatrichia kewi, new species
FIGuRE 194

Maur.—Head black-brown, longer than high; eyes black, divided
into larger upper and smaller lower facets, from side view area behind
the eye extensive, back of head and back of ocellar tubercle silvery
pilose above; postgena shining; frons with a slight median bulge
bordered by two vittae which converge from above the antennal bases

SCENOPINIDAE OF THE WORLD 301

Ficure 193.—Neopseudatrichia canei, new species, male: a, wing; d, lateral aspect of head,
antennae lost; c-e, ventral, lateral and posterior aspects of terminalia.

to the midpart of the frons, area laterad of vittae sparsely covered
with brownish hair. Mouthparts well developed, filling the oral cavity,
oral opening surrounded by silvery pile; ocellar tubercle slightly raised,
subshining, with a few anteriorly directed hairs, ocelli clear; antennae
black, granular, third segment oval, twice as long as 1 and 2 combined;
see figure.

302 U.S. NATIONAL MUSEUM BULLETIN 277

FicureE 194.—Neopseudatrichia kewi, new species, male, female: a, male wing; b, female wing;
c,d, lateral aspects of male and female heads; ¢, dorsal aspect of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, lateral and posterior aspects of male
terminalia; #, ventral aspect of female 8th sternum and 9th segment; /, dorsal aspect of
female 8th and 9th segments; m, lateral aspect of female 8th and 9th segments.

SCENOPINIDAE OF THE WORLD 303

Thorax dorsum black, shining, punctured with bases of sparse
short whitish hairs; humeral and supra-alar calli red-brown; pleural
areas blackish, shining. Wing hyaline, granular, veins red-brown;
halter stem brown, pollinose, knob brown, granular, elongate, pointed.
Legs with femora black-brown, remainder lighter.

Abdomen second segment surface wrinkled, with blue-black
metallic sheen, remaining segments black, surface wrinkled, with
metallic sheen, posterior margins white. Ninth tergum black-brown,
shorter than high; tenth segment extending well beyond; aedeagus
flanked by two short parameres; see figure for details.

Frema.e.—Head black, elongate, flattened; eyes black; frons shining,
slightly broader than the ocellar tubercle, thinly covered with short
hairs; back of head silvery pilose above including the back of the
ocellar tubercle; postgenae shining; oral opening surrounded by silvery
pile; ocellar tubercle shining, with a few anteriorly directed hairs, ocelli
red-brown. Antennae black, granular; third segment twice as long as
first two combined; see figure for details.

Thorax dorsum black, shining, punctured with the bases of short
golden hairs, humeral and supra-alar calli red-brown; scutellum black,
polished; pleural areas black, shining. Wing hyaline, brown granular,
veins brown; halter stem brown, pollinose, knob brown, granular,
elongate, pointed. Legs black, tibiae and tarsi with golden hairs on
ventral surfaces.

Abdomen black, rough, with metallic reflection, white posterior
membrane between segments. Eighth segment black, very hairy;
sternum spoonlike on midline, ninth tergum with much ventrally
directed hair.

Length: Male body 4.7 mm., wing 3 mm.; female body 5 mm.,
wing 3.2 mm.

Type-locality: Lees Spring, A.C.T., Australia; emerged 22 Novem-
ber 1960 (I. F. B. Common and M. 8. Upton) reared from bark of
Eucalyptus.

Holotype: Male (CSIRO).

Allotype: Female Kew, New South Wales; 12 October 1962
(D. H. Colless) (CSIRO).

201. Neopseudatrichia mariaensis (G. H. Hardy), new combination
FiaureE 195 (after Hardy)
Pseudatrichia mariaensis G. H. Hardy, 1933, p. 419.

The type and allotype of this species have been lost. The species is
probably restricted to the Island of Tasmania.
Length: Male, female body 5 mm., wing ?.

304 U.S. NATIONAL MUSEUM BULLETIN 277

Wt
DO =

Ficure 195.— Neopseudatrichia mariaensis (G.H.
Hardy), male, female: Wing, lateral and dorsal
aspects of male and female heads, and enlarged
detail of antenna (after Hardy).

Type-locality: Maria Island, Tasmania; 29 December 1915 (G. H.
Hardy) pair in copula, sweeping along or near the shore.
Types: lost.

Genus Scenopinula Paramonov
Scenopinula Paramonov, 1955, p. 651.

Type-species Scenopinula pallidipennis Paramonov (original des-
ignation). The genus Scenopinula Paramonov is closely related to the
genus Scenopinus but differs in the strongly swollen frons of the
female and indistinct weak wing venation where vein M 1 + 2 runs

parallel to R 5 but does not attain the margin of the wing. This genus
is limited to the Australian region.

202. Scenopinula pallidipennis Paramonoy
FicurE 196

Scenopinula pallidipennis Paramonov, 1955, p. 651.

This species is figured from the type.

Length: Female body 2 mm., wing 1.5 mm.

Type-locality: 8 mi. s. of Lake Cobham, New South Wales;
18 November 1949 (S. J. Paramonov).

Type: (CSIRO).

Genus Riekiella Paramonov

Riekiella Paramonov, 1955, p. 652.

Type-species Riekiella bicornis Paramonov (original designation).
The genus Riekiella Paramonov resembles the members of the Brevi-

SCENOPINIDAE OF THE WORLD 305

Gq

SW
If

wid
“UY yo
hoe.

~

Astlhestin
Sos cere

SY
)
Ai

Iw

Ficure 196.—Scenopinula pallidipennis Paramonov, female: a, wing; 5, Jateral aspect of
head; c, d, enlarged Jateral and frontal aspects of head; ¢, ventral aspect of 8th sternum;
f, lateral aspect of 8th and 9th segments.

trichva complex and at present may be composed of as many as three
genera. Until more specimens are available for study the exact combi-
nations are uncertain. On the basis of wing venation, the condition
of the short M 1+2 would place bicornis and complera together while
the genitalia of the male would place complera with quadrifida which
has a closed petiolate cell. The female sternum of bicornis is quite
different from that of quadrifida and dycei which have short blunt
sterna, but agrees with longiventris which also has a closed cell. The
members of this genus are found in the Australian region.

306 U.S. NATIONAL MUSEUM BULLETIN 277

Key to Species of Riekiella

1. Vein M 1+2 not reaching vein R5 or edge of wing . 2
Vein M 14+2 ie in R5 before ae of wing. 6

2. Males poke . ; 3
Females 4

3. Wing smoky waits a seal ppntaraee. R4 bracing beyond tie mmadale of
vein R5... 4 3% . .. . « R.complexa, new species

Wing brownish, halter nok en a a brown rib, vein R4 branching before
the middle of vein R5, dorsal lobes of ninth segment ending in wiry

BOOKS... wo els. ee owe oe ae 3 Py RS cata, new, ie

4. Wing brownish .. 5
Wing hyaline, vein R5 hendiig: backward bean the beanehing of vein R4,
halter yellow, very large. . . . . .. . . R. bicornis Paramonov

5. Halter knob yellow-brown, darker at bese antenna black-brown, R4 branch-
ing from R5 near the center of the vein, running straight to the leading
edge of the wing . . ee . . R. uncata, new species

Halter knob orange- coer ae a tan paedeae baad R4 branching from near
the basal third of R5 and curving ‘‘S’’-like to the leading edge of

WIDE 5s 6 de ew 5 Sue Ss toe Boh we eyens Wee COMGRSI, new species

Ge VPales:. is 28, aa a Bos. cocincect a eee on Sta: Pe ou eee eee ee es ae
Beniales. a. jo. sl Mahe ca ek op ks Sek, Bee eb dos (Bye ak en te a Ieee

7. Wing brownish ..... i win fe Rate gS

Wing hyaline, halter knob Oro BS a tan caniteal Soe. ninth tergum
one-fourth length of body, with upper and lower sections equal in
length... . . . .R. quadrifida, new species

8. Halter knob dull allo ei a ‘dorsal eee rib to the tip, ninth tergum
one-fourth the length of the body, the dorsal portion pointed and nearly
twice as long as the rounded ventral portion. . . R. taylori, new species

Halter knob tan-brown, ninth tergum short, dorsal portion pointed, ventral
portion hooked, aedeagal parameres extending beyond ventral portion of
ninth tergum. . . .. . 4) se ee ay. OR. collessi, new.species

9. Winghyaline. ... 3 ca but cape

Wing smoky brown, ate halter fenoh Cree Tie ee vein R4 branching

from near center of cell R5, ending in leading edge of wing beyond the tip

of cell R5. 2... . . . . . R. dycei, new species
10. Halter knob brown rite a het aimpe palo, antenna black, tip of cell R5
pointed, small under 4 mm. ... .. . . R quadrifida, new species

Halter knob light brown above, cream below, antenna with third segment
orange at base, tip red-brown, cell R5 blunt at tip, eighth sternum pointed
much longer than tergum; a large fly over 6 mm.

R. longiventris, new species

203. Riekiella bicornis Paramonov

FIGURE 197

Riekiella bicornis Paramonov, 1955, p. 653.

The type of this species is figured for the first time. Of interest are
the venation of the wing and the structure of the eighth sternum which
far exceeds the dorsum and the extent of the ninth segment.

Length: Female body 3 mm., wing 2 mm.

SCENOPINIDAE OF THE WORLD 307

Ficure 197.—Riekiella bicornis Paramonov, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, dorsal aspect of 8th and 9th segments; ¢, lateral aspect of
8th and 9th segments.

Type-locality: 10 mi. s. Bowen, Queensland; 27 September 1950
(E. F. Riek).
Type: (CSIRO).

204. Riekiella collessi, new species
FIGURE 198
Ma.e.—Head black; eyes black, divided below the midline with

coarse upper and fine lower facets; frons very narrow, triangular, with
a median groove, the lower half white; ocellar tubercle black with gray

308 U.S. NATIONAL MUSEUM BULLETIN 277

eranulosis; ocelli orange-red; mouthparts well developed, black, with a
lighter tip; palpi tan; oral opening black posteriorly grading into white
which joins the area on the lower frons. Antennae black, granular;
first segment short; second segment twice as wide and long as first;
third segment pear-shaped, not twice as long as first two combined,
ending in two distinct points with a median peg.

Thorax dorsum black, covered with grayish brown pollen; humeral
and supra-alar calli tan; pleura black, with gray granulosis. Wings
brownish hyaline, veins brown, M 1 + 2 reaching R5 closing off a cell;
halter stem brown, knob tan-brown; legs black-brown, granular.

Ficure 198.—Riekiella collessi, new species, male, female: a, male wing; b, female wing;
c, d, lateral and frontal aspects of male head; e, f, lateral and frontal aspects of female
head; g, h, enlarged details of male and female heads; i, 7, ventral and lateral aspects of
male terminalia; k, ventral aspect of female 8th sternum and 9th segment; /, lateral
aspect of female 8th and 9th segments.

Abdomen first three segments black, with brown granulosis, remain-
ing segments more shining, no distinct broad band of white membrane.
Ninth tergum red-brown, granular, divided into four lobes; aedeagus
as two stiff spines extending beyond ventral lobe; see figure for details.

Frma.Le.—Head black, eyes black; postocular ridge narrow, cov-
ered with gray pollen, with sparse stiff hairs; frons gray pollinose,
with two median bands of brownish pollen on either side of a median
groove, a few sparse hairs; ocellar tubercle gray pollinose with a few
sparse hairs; ocelli red-orange; mouthparts brown, palpi cream-orange;
oral opening with anterior two-thirds cream-orange; antennae with
first two segments black with gray pollen; third segment pear-shaped,
pubescent, shining bronze or black depending on the light.

SCENOPINIDAE OF THE WORLD 309

Thorax dorsum black, covered with brown and tan pollen, with
evenly spaced short black hairs; humeral callus cream, supra-alar
callus orange, scutellum black-brown with a tan median pollinose
patch; pleura black-brown, dusted with gray pollen. Wings brown
fumose, veins brown, M 1+2 ending before reaching R5 or edge of
wing; halter stem yellow basally to brown distally, knob orange-
brown above and below with a tan median band. Legs with femora
red-brown, pollinose with orange knees; tibiae red-brown basally,
orange-brown distally; tarsi lighter basally, darker distally.

Abdomen first three segments dark red-brown, granular; remaining
segments red-brown, shining; eighth segment elongate; see figure.

Length: Male body 2.6 mm., wing 1.8 mm.; female body 4 mm.,
wing 2.3 mm.

Type-locality: Black Yamma State Forest, New South Wales,
Australia; 11 November 1964 (D. H. Colless).

Holotypes: Male (CSIRO).

Allotype: Female, same data.

205. Riekiella complexa, new species

Figure 199

Maue.—Head black, overlain with gray scales on the back of the
head; eyes black-brown above, black below, meeting on the midline;
frons narrow, with a median ridge, extending half the distance to the
median ocellus, ocellar tubercle black-brown with gray scales, ocelli
red-brown; mouthparts black, well developed, extending to the base —
of the antennae, oral cavity tan. Antennae black, the third segment
velvety giving off brassy sheen under light; see figure for details.

Thorax dorsum black, overlain with brown polkinose scales, sparse
erect black hairs; humeral callus tan; pleural areas as on dorsum.
Wings smoky, with scaly appearance; veins brown, M 1+2 not
reaching R5 or tip of wing; calypters large, fringed with long hairs;
halteres missing on both sides. Legs black-brown, with tarsi and knees
of femora red-brown.

Abdomen second segment black-brown, shining; remaining seg-
ments pollinose ventrally, shining dorsally; ninth tergum black-brown,
pollinose, hairy dorsally, tan ventrally, formed into four long lobes,
resembling those of R. quadrifida but shorter. The terminalia were
lost on dissection; a rough sketch is provided; see figure.

Fremate.— Unknown.

Length: Male body 2.5 mm., wing 2 mm.

Type-locality: Canberra, Australian Capital Territory, Australia ;
3 December 1962 (E. F. Riek).

Holotype: Male (CSIRO).

310 U.S. NATIONAL MUSEUM BULLETIN 277

—

Figure 199.—Riekiella complexa, new species, male: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, rough sketch of terminalia before loss on dissection.

206. Riekiella dycei, new species
FicvuRE 200

This and the following species are placed in this genus despite the
variance in the wing venation, pending the study of additional speci-
mens, particularly pairs.

Frma.te.—Head higher than long, pinkish brown; eyes brown;
frons broad, with a median groove extending from the antennae to
the median ocellus; ocellar triangle darker, set off from post-ocular
ridges by grooves from the corner of the eye to the lateral ocellus;
genal areas below and behind the eyes yellow-brown, unmarked;
mouthparts well developed; antennal segments dark brown, the first
and second short, third broad at the base truncated distally with two
points.

SCENOPINIDAE OF THE WORLD la

Thorax gray pollinose with darker setal pits. Wings smoky, granu-
lar, veins dark brown; halter stem brown, knob creamy white. Legs
dark brown basally, pollinose; femora with pale brown knees, re-
mainder of legs light brown.

Abdomen elongate, dark brown, eighth segment elongate; see figure
for details.

Mate.— Unknown.

Length: Female body 3.25 mm., wings 2.25 mm.

Type-locality: 12 mi. s. Gravesend, New South Wales, Australia;
30 September 1952 (A. Dyce) attracted to rabbit.

Holotype: Female (CSIRO).

———

SS

e.

Ficure 200.— Riekiella dycei, new species, female: a, wing; b, lateral aspect of head; c,
enlarged detail of antenna; d, dorsal aspect of 8th and 9th segments; ¢, lateral aspect of
8th and 9th segments.

312 U.S. NATIONAL MUSEUM BULLETIN 277

207. Riekiella longiventris, new species

Figure 201

FrmaLe.—Head yellow-brown; eyes dark brown; postocular ridge
with a raised vertical face behind the eye, to an elevated ridge; back of
head black, covered with gray pollen and with short black hairs;
frons broad, raised in a central band, covered with a band of short
black hairs, lateral margins next to eye depressed, a bare triangle above
the antenna: bases; mouthparts well developed; ocellar tubercle
black-brown, raised from a depressed base, with a few hairs; ocelli
dark brown; frons separated from postocular area by grooves that

Ficure 201.—Riektella longiventris, new species, female: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d, enlarged dorsal aspect of head; e, dorsal aspect of 8th

and 9th segments; f, ventral aspect of 8th and 9th segments; g, lateral aspect of 8th and
9th segments.

SCENOPINIDAE OF THE WORLD Slo

extend from the upper corner of the eye behind the ocellar tubercle
to the midline at the back of the head. Antennae with the first two
segments black-brown, subequal; the second with a fringe of hairs;
the third segment pear-shaped, orange basally darkening distally to
black-brown with a two-pointed tip.

Thorax dorsum black, covered with dark brown pollinose scales,
sparse short black hair; humeral callus tan; scutellum black; pleural
areas black-brown, pollinose, very little membrane or light color.
Wings hyaline, veins light brown; halter stem light brown, knob light
brown, creamy below. Legs with femora black-brown, pollinose;
remainder of legs light brown.

Abdomen very long, dark brown, grading to yellow-brown pos-
teriorly; eighth segment with dorsum shining mahogany brown,
sternum yellow-brown, keeled in posterior half, twice as long as ter-
gum. Ninth segment long, much like R. bicornis; see figure for details.

Mate.— Unknown.

Length: Female body 6 mm., wing 2.9 mm.

Type-locality: Applecross, West Australia; 25 February 1941
(K. R. Norris).

Holotype: Female (CSIRO).

208. Riekiella quadrifida, new species

FIGURE 202

Mae.—Head black, eyes red-brown above, black below; frons
small, granular, extending only one-third the distance from the anten-
nae to the median ocellus; ocellar triangle prominent, granular, ocelli
dark; oral cavity tan to cream, with a black roof; mouthparts tan,
short, palpi short. Antennae black, granular, third segment twice as
long as first two combined, pear-shaped, ending in two points with a
median peg; see figure for details.

Thorax dorsum black, with red-brown granular pollen; humeral and
supra-alar calli tan; scutellum black, subshining; pleura black, granu-
lar. Wing hyaline, veins tan, cell R5 petiolate; halter stem brown,
knob brown, with a tan band below. Legs with femora black-brown,
granular; tibiae tan; tarsi tan, darkening distally.

Abdomen black-brown, granular dorsally, shining below; ninth seg-
ment black-brown, four-lobed, elongate, being one-quarter the length
of the entire body; see figure for details.

FrmaA.Le.—Head black-brown, eyes black; postocular flange con-
tinuous with dorsal portion of frons; back of head granular; frons
broad, shining, over twice as wide as ocellar triangle, with a groove
extending from median ocellus toward the antennae where it becomes
an elevated swelling with sparse hairs; ora] cavity with tan sides and a

314 U.S. NATIONAL MUSEUM BULLETIN 277

ASS
= an SSN Ve

SN -
fa

Frcure 202.—Riekiella quadrisida, new species, male, female: a, wing; b, c, lateral and
anterior aspects of male head; d, ¢, lateral and anterior aspects of female head; f, g, enlarged
details of male and female antennae; h-j, ventral, dorsal, and lateral aspects of male
terminalia; &, ventral aspect of female 8th sternum and 9th segment; J, lateral aspect of
female 8th and 9th segments.

black roof; mouthparts brown, short; palpi tan, short. Antennae
black, granular, as in male.

SCENOPINIDAE OF THE WORLD So

Thorax as in male.

Abdomen black-brown, granular dorsally, shining below; eighth
segment elongate, tergum and sternum subequal; ninth segment
structures elongate; see figure.

Length: Male body 3 mm., wing 1.9 mm.; female body 3.5 mm.,
wing 2.2 mm.

Type-locality: N. Rosewood, Queensland; 29 December 1961 (R.
Lindsay) brigalow scrub.

Holotype: Male (CSIRO).

Allotype: Female, same data (CSIRO).

Paratypes: 167, 1 9, same data (CSIRO).

209. Riekiella taylori, new species

Figure 203

Maur.—Head black-brown, eyes red-brown with metallic glints,
darker below; no postocular flange; frons narrow, triangular, with a
median groove, lower frons granular; ocellar tubercle black-brown
dusted with gray pollen, a few anteriorly directed hairs from back;
ocelli orange. Antennae with first segment black-brown, small; second
segment black-brown, granular; third segment missing.

Thorax dorsum black-brown, heavily dusted with gray pollen, mar-
gin of tergum behind humeral callus with stiff hairs; humeral callus
cream with 8-10 stiff tan hairs, scutellum black with brown pollen;
pleural areas black-brown granular. Wings brownish hyaline, veins
yellow-brown, squamae noticeable; halter stem brown, knob dull
yellow with a dorsal brown rib to tip. Legs dark red-brown, granular,
femoral knees orange.

Abdomen black-brown, pubescent, giving a silvery-white appear-
ance in certain light, intersegmental membrane tan; terminalia
black-brown at base, tips and lower arms orange-brown, hairy, one-
quarter length of body; see figure for details of shape.

FremMALe.— Unknown.

Length: Male body 3.2 mm., wing 2.2 mm.

Type-locality: 20 mi. w. Mildura, Victoria, Australia; 22 October
1958 (K. L. Taylor).

Holotype: Male (CSIRO).

210. Riekiella uncata, new species

Figure 204

Ma.r.—Head black; eyes black, with reddish glints, divided on
midline into larger upper and smaller lower facets; frons narrow, tri-
angular, a dark median streak of pubescence on lower frons bordered

313-168 O—69__21

316 U.S. NATIONAL MUSEUM BULLETIN 277

———

Ficure 203.—Riekiella taylori, new species, male: a, wing; }, c, lateral and frontal aspects of
head; d-f, ventral, dorsal and lateral aspects of terminalia.

by tan pubescence laterally, upper half shining, continuing as a narrow
band between the eyes; ocellar tubercle black, covered with brown
pubescence, a few scattered black hairs on back margin; ocelli red-
brown; mouthparts black-brown, two-thirds the length of the oral
opening, palpi brown, half as long as mouthparts; back of head gray

SCENOPINIDAE OF THE WORLD ola

pubescent; antennae black, pubescent, third segment pear-shaped,
with a bronzy sheen.

Thorax dorsum ground black, surface covered with brown and tan
pollen, a thin scattering of short posteriorly directed bristles; humeral
and supra-alar calli cream; pleura with black-brown pollen. Wings
brown fumose, veins brown, M 1 + 2 fading before reaching R5;
halter stem orange-brown, knob tan, with a brown rib. Legs with

Ficure 204.—Riekiella uncata, new species, male, female: a, wing, upper dashed line indi-
cates line of vein on male wing, lower female; b, c, lateral and frontal aspects of male head;
d, e, lateral and frontal aspects of female head; f, g, enlarged details of male and female
heads; A-j, ventral, dorsal and lateral aspects of male terminalia; k, ventral aspect of
female 8th sternum and 9th segment; /, lateral aspect of female 8th and 9th segments.

coxae and femora black, pollinose; tibiae orange-brown, tarsi brown.

Abdomen black-brown with brown pollen, ninth tergum black-
brown to orange-brown, pollinose, hairy, tergum ends in a pair of
rounded hooks.

Frmate.—Head black-brown, eyes red-brown with red metallic
glints; postocular ridge narrow, with a band of hairs, granular; frons
with dark brown granulosis, with tan margins next to eyes, a median
groove from median ocellus, sunken for half the length of the frons,
continuing over the swollen lower half to antennal base, a few scattered
short hairs; back of head covered with dark brown pollen; mouthparts
brown, two-thirds the length of oral opening, palpi shorter, pale
orange, anterior half of oral opening light orange; ocellar tubercle
dark brown, granular; ocelli red-brown; antennae black-brown, first

318 U.S. NATIONAL MUSEUM BULLETIN 277

segment granular; second granular, with a few hairs; third segment
pear-shaped pubescent giving off bronze glints.

Thorax dorsum dark brown, granulose, with tan bands, with scat-
tered short, stiff, posteriorly directed bristles; humeral and supra-
alar calli yellow-brown; scutellum black-brown, granular; pleura
black-brown, granular. Wings brown fumose, veins brown, M 1 + 2
not reaching R5; halter stem orange-brown basally darkening to
brown distally, knob yellow brown, darker at base. Legs with femora
red-brown, granular; tibiae yellow-brown; tarsi with first segment
yellow-brown, remainder red-brown.

Abdomen first three segments black-brown, granular, remaining
segments black-brown shining, eighth segment elongate; see figure
for details.

Length: Male body 2.2 mm., wing 1.7 mm.; female body 3.3 mm.,
wing 2 mm.

Type-locality: Parkes, New South Wales, Australia; 12 November
1964 (D. H. Colless).

Holotype: Male (CSIRO).

Allotype: Female, same data.

Paratypes: 1 o, 19, Parkes, New South Wales, same data as
type; 1 &, 1 9, Eugowra, New South Wales, 15 November 1964
(D. H. Colless) (CSIRO).

Genus Stenomphrale Krober
Stenomphrale Kréber, 1937, p. 220.

Type-species Pseudomphrale teutankhament Kroéber. The genus
Stenomphrale Krober closely resembles the genus Pseudatrichia; the
male genitalia, however, lack the ventral brush of hairs and the
tergites are longer than those of Neopseudatrichia; the female termi-
nalia are very different from those of Pseudatrichia, having a brush
of hairs on the ninth tergum which is larger than those of Neopseuda-
trichia. This genus is found in the circum-Mediterranean area of the
Palaearctic region.

211. Stenomphrale flavoscutellata (Kréber)
Figure 205 (after Krober)

Pseudomphrale flavoscutellata Kréber, 1929, p. 80.
Stenomphrale flavoscutellata (Kréber), 1937, pp. 213, 220.

The only figures available for this species are those of Kréber
accompanying the original description and are among the least
adequate of the many figures he published. All attempts to locate the
types of this species have failed. The material supposedly is in the

SCENOPINIDAE OF THE WORLD 319

Figure 205.—Stenomphrale flavoscutellata (Kréber), male,
female: Anterodorsal aspect of head, detail of antennae
and dorsal aspect of abdomen (after Kroéber).

collections of the Royal Entomological Society of Egypt but the
curators of that collection were unable to locate it.

Length: Male body 3.5-3.6 mm., wing 2.1-2.4 mm.; female body
4mm., wing 3 mm.

Type-locality: Burg (Mariout), Egypt; 10 May 1927.

Type: (SEE) lost?

212. Stenomphrale teutankhameni (Kréber)

Figure 206

Pseudomphrale teutankhameni Krober, 19238, p. 72.
Stenomphrale teutankhameni (Kroéber), 1937, pp. 218, 220.

Kréber attributes the authorship of this species to Collin. However,
inasmuch as Kréber made complete original descriptions of both the
male and female, and no reference is found in the literature to a
description or even a reference to this species by Collin, I feel that
this species should be attributed to Kréber. This species appears
closely related to the Australian genera Neopseudatrichia and Para-
trichia in the form of the male ninth and tenth terga; it differs, how-
ever, in the large mouthparts and the venation of the wing. The female
eighth and ninth segments are quite different.

Length: Male body 3.5-3.7 mm., wing 2.4 mm.; female body 2.5-—
3.5 mm., wing 2 mm.

Type-locality: Mariout, Egypt; 16 March 1922.

320 U.S. NATIONAL MUSEUM BULLETIN 277

Figure 206.—Stenomphrale teutankhameni (Krober), male, female: a, wing; b, c, lateral and
frontal aspects of male head; d, ¢, lateral and frontal aspects of female head; f, g, enlarged
details of male and female antennae; A-j, ventral, lateral and posterior aspects of male
terminalia; k, ventral aspect of right half of female 8th and 9th segments; /, dorsal aspect

of left half of female 8th and 9th segments; m, lateral aspect of female 6th, 7th, 8th and 9th
segments.

Types: (SEE).
Allotype: Female, same data as type.
Paratypes: 1, same data as type; 1 9, Mariout, 5 April 1921 (SEE).

Genus Paratrichia, new genus

Type-species Paratrichia lobosa, new species. The genus Paratrichia
resembles the genus Pseudatrichia in wing venation; the head is higher
than long, the antennae short and forked at the tip. The male genitalia
have a pair of flaplike processes on the inside walls of the ninth tergite
which character separates it from other genera. The female is unknown.
Members of this genus are found in the Australian region.

213. Paratrichia lobosa, new species

FIGURE 207

Maure.—Head dark brown; eyes brown above, darker below; frons
narrowing to a point halfway to the median ocellus; ocellar tubercle
brown, ocelli clear; mouthparts well developed, filling half the oral

cavity, which is tan. Antennae brown, third segment oval, with a
bilobed tip.

SCENOPINIDAE OF THE WORLD ool

Ficure 207.—Paratrichia lobosa, new species, male: a, wing; b, lateral aspect of head; c,
enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia (note
flaps on either side of tergum).

Thorax dorsum dark brown, covered thinly with long hairs which
show white, a dusting of brown on the integument; humeral callus
with a small tan spot; scutellum solid dark brown, with erect white
hairs; pleura dark brown. Wings hyaline, veins brown, cell R5 closed
petiolate; halter stem light brown, knob chocolate brown. Legs dark
brown with tan knees.

Abdomen chocolate-brown, subshining. Ninth tergum small, blunt,
chocolate-brown, with sclerotized flaplike processes on inner face,
aedeagus short without visible parameres; see figure for details.

FrmaLe.—Unknown.

Length: Male body 3.3 mm., wing 2.2 mm.

Type-locality: Black Mt., Australian Capital Territory, Australia ;
30 November 1920 (A. Tonnoir).

Holotype: Male (CSIRO).

a22 U.S. NATIONAL MUSEUM BULLETIN 277

214. Paratrichia westralica (Paramonoy), new combination

Figure 208
Pseudomphrale westralica Paramonov, 1955, p. 641.

This species, though not agreeing in all respects with P. lobosa, is
placed with it for the present on the basis of similarity in the place of
branching of vein R4 from the middle of the cell and the presence of
sclerotized flaps on the inner surface of the ninth tergum.

Length: Male body 2 mm., wing 2.2 mm.

Type-locality: Zanthers, W. Australia; 21 August 1926.

Type: (CSIRO).

Ficure 208.—Paratrichia westralica (Paramonov), male: a, wing; b, lateral aspect of head;
c, enlarged detail of antenna; d-f, ventral, lateral and posterior aspects of terminalia (note
flaps on inside of tergum).

Literature Cited

Apams, C. F.
1904. Notes on and descriptions of North American Diptera. Kans. Univ.
Sci. Bull., vol. 2 (=whole ser., 12): 433-455. (=Kans. Univ. Bull.,

vol. 4 [6].)
Aaassiz, L.
1846. Nomenclatoris Zoologici. Index Universalis. . . . 393 pp. Solo-
dure [Solothurn], Switzerland.
BECKER, T.

1902. Aegyptische Dipteren. Berlin Zool. Mus. Mitt. vol. 2, (h. 2, art. 1)
pp. 1-66, 1 plate.

1907. Die Ergebnisse meiner dipterologischen Friihjahrsreise nach Algier
und Tunis, 1906 (Concel.). Z. system. Hym. Dipt., vol. 7, pp.
33-61, 97-128, 225-256, 369-407, 454, 12 figures.

1913. Persische Dipteren von den Expeditionen des Herrn N. Zarudny
1898 und 1901. (Akad. Nauk, S.8.8.R.) Imp. Akad. Nauk. St.
Petersburg, Zool. Mus. Ezheg. (Acad. Imp. des Sci. de St. Péters-
bourg, Zool. Mus. Ann.), vol. 17, pp. 503-654, 3 plates.

Bezzi, M.

1898. Contribuzioni alla fauna ditterologica italiana. Soc. Ent. Ital.
Bull., vol. 30, pp. 19-50.

1922. On the South American species of the Dipterous genus Chiromyza
Wied. (Diptera). Ann. Ent. Soc. Amer., vol. 15, pp. 117-124.

1925. Une Nouvelle Espéce du Genre Pseudomphrale de V Afrique du Sud.
Encyclopédie Entomol., sér. B, Diptera. Vol. 2, pp. 95-98.

Couez, F. R.

1923. Expedition of the California Academy of Sciences to the Gulf of
California in 1921. Diptera from the islands and adjacent shores of
the Gulf of California. II. General report. Proc. California
Acad. Sci., ser. 4, vol. 12, pp. 289-314, 38 figures, pp. 457-481, 16
figures.

CoquiL_LETT, D. W.

1900. New Scenopinidae from the United States. Ent. News, vol. 11, pp.
500-501.

1902. New Diptera from North America. U.S. Nat. Mus. Proce. (1903) vol.
25, pp. 83-126.

Cresson, E. T., JR.

1907. The North American species of the dipterous family Scenopinidae.

Trans. Amer. Ent. Soc., vol. 33, pp. 109-114, 1 plate.
Data Torre, K. W. von

1878. Beitrage zur Phyto-& Zoostatik des Egerlandes. Jehresb. naturh.

Ver. Lotos, vol. 27 (1877), pp. 7-208. Prague.
DE GEER, C.

1776. Mémoires pour servir 4 Vhistoire des Insectes. Vol. 6, 523 pp., 30
plates. Stockholm.

323

324 U.S. NATIONAL MUSEUM BULLETIN 277

Epwarps, F. W.
1932. Scenopinidae, Mydaidae. Diptera of Patagonia and South Chile,
part 5, fase. 3, pp. 258-260.
ENDERLEIN, G.
1913. Dipterologische Studien III. Uber Lagarinus nov. gen., eine isoliert
stehende Fliegengattung. Zool. Anz., vol. 42, pp. 250-253.
1914. Dipterologische Studien X. Zur Kenntnis der Stratiomyiiden mit 3
zastiger Media und ihre Gruppierung. B. Formen, bei denen der 1.
Cubitalast mit der Discoidalzelle eine Strecke verschmolzen ist
(Familien: Hermetiinae, Clitellariinae). Zool. Anz., vol. 44, pp.
1-25.
1934. Dipterologica. I. Ges. Naturf. Freunde, Berlin. Sitzb. (1933), pp.
416-429.
ENGEL, E. O.
1932. Familie: Omphralidae, pp. 123-126. Jn Dahl, F., ed., Die Tierweldt
Deutschlands. Vol. 26, Teil 5, 204 p. Jena.
Fasricius, J. C.
1781. Species insectorum exhibentes eorum differentias specificas, synonyma,
auctorum, loca natalia, metamorphosin. Vol. 2, 517 pp. Hamburg

and Kiel.
1794. Entomologia systematica emendata et aucta. Vol. 4, 472 pp. Copen-
hagen.
Fauu&n, C. F.
1817. Scenopinii et Conopsariea Sveciae. 14 pp. Lund.

Frey, R.

1937. Die Dipterenfauna der Kanarischen Inseln und ihre Probleme. Com-
ment. biol. Finska Vetensk. Soc., vol. 6, 237 pp., 10 plates.

1945. Tierogeographische Studien iiber die Dipterenfauna der Azoren. I.
Verzeichnis der bisher von den Azoren bekannten Dipteren. Com-
ment. biol. Finska Vetensk. Soc., vol. 8, 114 pp., 4 plates.

GrimsHaw, P. H.

1901. Part I: Diptera. pp. 1-77, plates 1-3. Jn Sharp, D., ed., Fauna

Hawaiiensis. Vol. 3, 704 pp., 19 plates. Cambridge, England.
Hauiway, A. H.

1851. Uber die Dipteren der in London befindlichen Linnéischen Sammlung.

Stettin ent. Zeitung, vol. 12, pp. 131-145.
Harpy, D. E.

1944a. A Revision of the North American Omphralidae (Scenopinidae).
Kansas Ent. Soc. J., vol. 17, pp. 31-51, 1 plate.

1944b. A new Pseudatrichia from Brazil (Omphralidae—Scenopinidae).
Kansas Ent. Soe. J., vol. 17, pp. 104-105, 1 figure.

1958. Diptera: Omphralidae (Scenopinidae). Jn Insects of Micronesia, vol.
13, pp. 11-18, 1 figure.

1960. Diptera: Nematocera—Brachycera (except Dolichopodidae). In
Zimmerman, E. C., ed., Insects of Hawaii, Honolulu. Vol. 10, 368
pp., 120 figures.

Harpy, G. H.

1921. Revision of the Chiromyzini. Proc. Linn. Soc., New South Wales
(1920), vol. 45, pp. 582-542, 2 plates.

1933. Miscellaneous Notes on Australian Diptera, I. Proc. Linn. Soc.,
New South Wales, vol. 58, pp. 408-420, 4 figures.

1942. Miscellaneous Notes on Australian Diptera, IX: Superfamily Asiloi-
dea. Proc. Linn. Soc., New South Wales, vol. 67, pp. 197-204.

SCENOPINIDAE OF THE WORLD oO

INTERNATIONAL COMMISSION ON ZOOLOGICAL NOMENCLATURE

1963b. Opinion 678. The suppression under the Plenary Powers of the
pamphlet published by Meigen, 1800. Int. Comm. Zool. Nomencl.,
Bull. Zool. Nomencel., vol. 20, pp. 339-342.

JAMES, M. T.

1938. The dipterous families Nemestrinidae, Cyrtidae and Scenopinidae in
Colorado. Kansas Ent. Soc. J., vol. 11, pp. 21-23.

1955. Two new Diptera from the Pacific Coast States. Kansas Ent. Soe. J.,
vol. 28, pp. 47-48, 1 figure.

Kertész, K.

1899. Verzeichniss einiger, von L. Bird in Neu-Guinea und am Malayischen
Archipel. Gesammelten Dipteren. Természetn. Fuz., vol. 22, pp.
173-195.

KROBER, O.
1912. H. Sauter’s Formosa-Ausbeute. Thereviden, Omphraliden (Dipt.).
Suppl. Ent., vol. 1, pp. 24-26.
1913. Die Omphraliden. Eine monographische Studie. Ann. Mus. Nat.
Hung,., vol. 11, pp. 174-210, 4 plates.
1914a. Diptera Fam. Omphralidae. Fasc. 161, pp. 1-16, 4 plates. In
Wytsman P., ed., Genera insectorum (q.v.). Bruxelles.

1914b. Beitrage zur Kenntnis der Thereviden und Omphraliden. Jahrb.
Hamburg. wiss. Anst. (1913), vol. 31 Beih. 2, pp. 29-74, 3 figures,
(=Zool. Mus. Hamberg Mitt., vol. 31).

1923. Aegyptische Dipteren aus den Familien der Conopidae, Omphralidae,
und Therevidae. Bull. Soc. Roy. Ent. d’Egypte, vol. 7, pp. 57-
116, 26 figures.

1925. Omphralidae (Scenopinidae). Jn Lindner, E., ed., Die Fliegen der
palaeartischen Region. Vol. 4, pt. 3, sect. 2, fam. 27, pp. 1-8, 2 plates.

1928a. Neue und wenig bekannte Dipteren aus den Familien Omphralidae,

Conopidae und Therevidae. Konowia, vol. 7, pp. 1-23, 17 figures.
1928b. Neue Dipteren des Deutschen Entomolog. Museums in Dahlem.
Ent. Mitt., vol. 17, pp. 31-41.

1929. Neue Dipteren aus Aegypten aus den Familien Tabanidae, Therevidae,
Omphralidae u. Conopidae. Bull. Soc. Roy. Ent. d’Egypte, n.s.,
vol. 13, pp. 73-84.

1937. Ein Beitrag zur Kenntnis der Omphraliden (Scenopiniden), Diptera.
Stettin ent. Zeitung, vol. 98, pp. 211-231, 2 figures.

1939. Beitrag zur Kenntnis der Therevidae und Omphralidae. Ann. Mag.
Nat. Hist., ser. 11, vol. 3, pp. 395-400, 5 figures.

LAMB, C. G.

1922. The Percy Sladen Trust Expedition to the Indian Ocean in 1905.
No. VIII.—Diptera: Asilidae, Scenopinidae, Dolichopodidae,
Pipunculidae, Syrphidae. Trans. Linn. Soc. London, vol. 18, pp.
361-416, 4 plates.

LATREILLE, P. A.

1802. Histoire naturelle, générale et particulitre, des Crustacés et des In-
sectes, Tome troisi¢me [vol. 3], (vol. 95), 468 pp. Jn Sonnini, C.5S.,
ed., Histoire naturelle par Buffon (q. v.). Paris, ““An X.”

LINDNER, E.

1955. Ostafrikanische Omphralidae, Therevidae und Conopidae (Dipt.)
Jahrb. Ver. vaterl. Naturk. Wiirttenberg, vol. 110, pp. 19-23, 3
figures.

326 U.S. NATIONAL MUSEUM BULLETIN 277

1962. Afrikanische Rhagionidae, Therevidae und Omphralidae (Dipt.).

Stuttgarter Beitr. Nat., no. 83, 3 pp., 1 figure.
LINNAEUS, C.

1758. Systema naturae per regna tria naturae. 10th ed., vol. 1, 824 pp.

Stockholm.
Lioy, P.

1864. I ditteri distribuiti secundo un nuova metodo di classificazione
naturale. Atti Ist. Veneto Sci., ser. 3, vol. 9, pp. 499-518, 569-604,
719-771, 879-910, 989-1027, 1087-1126, 1311-1352.

Loew, H.

1845. Ueber die systemat. Stellung von Scenopinus, und Beschreibung
einer neuen Art. Stettin Ent. Zeit., vol. 6, pp. 312-315.

1857. Die bis jetzt bekanntgewordenen Arten der Gattung Scenopinus.
Verhandl. Zool.-Bot. Ver. Wien, 7 (Abhandl.) pp. 87-90.

1866. Diptera Americae septentrionalis indigena. Centuria septima.
Berlin. ent. Z., vol. 10, pp. 1-54.

1869. Diptera Americae septentrionalis indigena. Centuria octava. Berlin.
ent. Z., vol. 13, pp. 1-52.

1873. Beschreibung europdischer Dipteren. Systematische Beschreibung der
bekannten europdischen zweifliigeligen Insecten, von Johann
Wilhelm Meigen. Vol. 3, Zehnter Theil oder vierter Supplement-
band, 320 pp. Halle.

1874. Diptera nova a Hug. Theod. Christopho Collecta. Ztschr. f. die
Gesam. Naturw., vol. 43, pp. 413-420.

Macaquvakt, J.

1835. Histoire naturelle des Insectes. Diptéres, Tome deuxié¢me. Diptera,
vol. 2, 703 pp., 12 plates. Jn Roret, N. C., ed., Collection des
suites 4 Buffon (q. v.). Paris.

1843. Diptéres exotiques nouveaux ou peu connus. Vol. 2, part 3, pp.
5-304, 36 plates. Paris.

MEIcEN, J. W.

1800. Nouvelle classification des mouches 4 deux ailes (Dipterea L.) d’aprés
un plan tout nouveau. 40 pp. Paris. This publication has recently
been suppressed by the International Commission on Zoological
Nomenclature (International Commission on Zoological Nomen-
clature, 1963b).

1803. Versuch einer neuen Gattungseintheilung der europdischen zwei-
fliigeligen Insekten. Mag. Insektenkunde, vol. 2, pp. 259-281.

1824. Systematische Beschreibung der bekannten europdischen zwei-
fliigeligen Insekten. Vol. 4, 428 pp., 9 plates. Hamm.

1838. Systematische Beschreibung der bekannten europidischen zwei-
fliigeligen Insekten. Vol. 7, 434 pp., 8 plates. Hamm.

MEIJERE, J. C. H. DE

1924. Studien iiber Siidostasiatische Dipteren, XV. Tijdschr. Ent., vol. 67,

pp. 1-64.
OsTEN SACKEN, C. R.

1877. Western Diptera: Descriptions of new genera and species of Diptera
from the region west of the Mississippi and especially from Cali-
fornia. Bull. U.S. Geol. Geog. Survey Ter., vol. 3, pp. 189-354.

Panzer, G. F. W.

1809. Faunae insectorum germanicae initiae oder Deutschlands Insecten.

H. 98, p. 20, plate 20.

SCENOPINIDAE OF THE WORLD 327

PaRAMONOY, S. J.

1955. A Review of Australian Scenopinidae (Diptera). Australian Journ.

Zool., vol. 3, pp. 634-653, 2 figures.
RONDANI, C.

1848. Esame di varie specie d’insetti ditteri brasiliani. Studi Ent. (Turin),

vol. 1, pp. 63-112.
Rosst, P.

1794. Mantissa Insectorum, exhibens species nuper in Etruria collectas,
adjectis faunae Etruscae illustrationibus ac emendationibus. Pisa,
Polloni, 1792-1794. 4. 2 vol., 148 pp., 154 pp., 8 colored plates.

Say, T.

1823. Descriptions of dipterous insects of the United States. J. Acad. Nat.
Sci. Phila., vol. 3, pp. 9-54, 73-104.

1829. Descriptions of North American dipterous insects. J. Acad. Nat.
Sci. Phila., vol. 6, pp. 149-178.

SCHELLENBERG, J. R.

1803. Genres des mouches Dipteéres représentés en XLII planches pro-
jettées et dessinées et expliquées par deux amateurs de l’entomo-
logie. 95 pp., 42 plates. Zurich.

ScHINnER, I. R.

1862. Die Fliegen (Diptera) Jn Redtenbacher, L. and Schiner, I. R., Fauna

Austriaca (q. v.). Vol. 1, 674 pp., 2 plates.
ScHRANK, F. von P.

1803. Fauna Boica. Durchgedachte Geschichte der in Baiern einheimi-

schen und zahmen Thiere. Vol. 3, 272 pp.
Scopout, J. A.

1763. Entomologia carniolica exhibens insecta carnioliae indigene et dis-
tributa in ordenes, genera, species varietates methodo Linnaeana.
421 pp. Vienna.

Sséeuy, E.

1920. Note sur quelques Omphrale (Dipt. Omphralidae) et descriptions
d’espéces nouvelles. Bull. Soc. Ent. de France, pp. 317-319.

1930. Contribution 4 l’étude des dipttres du Maroc. Mém. Sci. Nat.
Maroc, vol. 24, 206 pp. 115 figures.

1931. Contribution 4 l’étude de la Faune du Mozambique. Voyage de M. P.
Lesne 1928-1929. Bull. Mus. d’Hist. Nat., Paris, vol. 3, pp.
113-121, 1 figure.

1932. Spedizione scientifica all’oasi di Cufra (Marzo-Luglio 1931) Insectes
Diptéres. Ann. Mus. Stor. Nat., Genova, vol. 55, pp. 490-511,
3 figures.

1933. Mission Saharienne Augiéras-Draper 1927-1928. Insectes Diptéres.
Bull. Mus. d’Hist. Nat., Paris, sér. 2, vol. 5, pp. 122-127, 9 figures.

1934. Diptéres d’Afrique. Encyclopédie Entomol., sér. B, Diptera. Vol. 7,
pp. 638-80, 12 figures.

1938. Mission scientifique de ’!Omo. Diptera I, Nematocera et Brachycera.
Mém. Mus. Hist. Nat., Paris Mémoires, n.s., vol. 8, pp. 319-380,
55 figures.

1948. Diptéres nouveaux ou peu connus d’extréme-orient. Mus. Heude
Notes. Ent. Chin., vol. 12, pp. 153-172, 10 figures.

SenroR-Wuite, R. A.

1922/24. New Ceylon Diptera. Spolia Zeylan., vol. 13, pp. 193-283, 12

plates.

328 U.S. NATIONAL MUSEUM BULLETIN 277

SPEISER, P.
1920. Zur Kenntnis der Diptera Orthorrhapha Brachycera. Zool. Jahresb.,
vol. 43, pp. 195-220, 7 figures.
STROBL, G.
1904. Neue Beitrige zur Dipterenfauna der Balkanhalbinsel. Wiss. Mitt.
Bosn. Herz., vol. 9, pp. 519-581.
Trogan, P.
1956. Notes on the taxonomy of some European species of the genus
Omphrale Meigen (Diptera, Omphralidae). Annls. Zool. Warsz.,
vol. 16, pp. 147-156.
Van DuzeEx, M. C.
1926. A new species of Scenopinidae from California (Diptera). Pan-Pacific
Ent., vol. 2, p. 164.
VILLENEUVE, J.
1913. Diptéres nouveaux ou intéressants. Feuille des Jeunes Nat.; vol. 43,
pp. 111-113.
WALKER, F.
1851. In Walker, F., Stainton, H. T. and Wilkinson, S. J., Insecta Britannica
(q. v.). Vol. 1, 314 pp., 10 plates. London.
ZETTERSTEDT, J. W.
1844. Diptera Scandinaviae. Disposita et descripta. Vol. 3, pp. 895-1280.
Lund.
1859. Diptera Scandinaviae. Disposita et descripta. Vol. 13, pp. 4943-
6190. Lund.

Index

(Names of new species and genera in 7talics; page numbers of principal accounts in
italics)

addacifons, Brevitrichia,
(fig.)
adventicius, Scenopinus, 9, 113 (fig.)
aethiopicus, Omphrale, 48
aethiopicus, Scenopinus, 8, 48, 49 (fig.)
africana, Pseudomphrale, 219
alalacteus, Scenopinus, 9, 114 (fig.)
albicineta, Musca, 49
albicinctus, Scenopinus, 8, 49, 50 (fig.)
Albicinctus group, Scenopinus, 2, 8, 14,
45 (key), 57, 76, 83, 92
albicomus, Scenopinus, 7, 17, 18 (fig.)
albidipennis, Scenopinus, 147, 148
albifasciata, Omphralosoma, 115
albifaciatus, Scenopinus, 2, 8, 115, 116
(fig.)
albipilosa, Belosta, 12, 274, 275, 276
(fig.)
albocincta, Pseudatrichia, 11, 225 (fig.)
angustifrons, Omphrale, 51
angustifrons, Scenopinus, 8, 51 (fig.)
antennata, Omphrale, 19
antennata, Paromphrale, 19
antennatus, Scenopinus, 2, 7, 19 (fig.)
anthrax, Scenopinus, 8, 61, 52 (fig.)
aquelonius, Scenopinus, 7, 20, 21 (fig.)
arnaudi, 10, 171 (fig.)
Archiscenopinus, 2, 15
aspinosa, Brevitrichia, 10, 172, 173 (fig.)
atombomba, Pseudatrichia, 21, 225, 226
(fig.)
atra, Scenopinus. 36
Astoma, 15
Atrichia, 14, 22]
fasciata, 26
longurio, 221, 255

10, 169,

balteatus, Scenopinus, 8, 53, 57 (fig.)
bakeri, Pseudatrichia, 11, 227, 228 (fig.)
barberi, Scenopinus, 9, 115, 116 (fig.)
barnesi, Pseudatrichia, 11, 229 (fig.)
beameri, Brevitrichia, 10, 174, 175 (fig.)

170| beameri, Omphrale, 117

beameri, Scenopinus, 9, 117, 118 (fig.)
beameri var. fusca, Omphrale, 117
beameri var. fusca, Scenopinus, 9, 117,
118 (fig.)
Belosta, 5, 12, 274 (key)
albipilosa, 12, 274, 275, 276 (fig.)
flaviceps, 275, 278
pilosa, 12, 275, 277 (fig.)
telfordi, 12, 278, 279 (fig.)
termitophaga, 12, 278, 280 (fig.)
viticolapennis, 12, 281, 282 (fig.)
biacristerna, Pseudatrichia, 11, 230, 231
(fig.)
bicornis, Riekiella, 13, 306, 307 (fig.)
biroi, Scenopinus, 8, 53, 54 (fig.)
boharti, Brevitrichia, 10, 174, 176 (fig.)
bolgarti, Scenopinus, 8, 55, 56 (fig.)
bouvieri, Omphrale, 57
bouvieri, Scenopinus, 8, 57, 58 (fig.)
brevicornis, Scenopinus, 9, 60, 94 (fig.)
95 (fig.)
Brevicornis group, Scenopinus, 9, 15, 93
(key)
breviterminus, Scenopinus, 7, 22, 23 (fig.)
Brevitrichia 10, 165 (key), 189, 281, 286,
304
addacifons, 10, 169, 170 (fig.)
arnaudi, 10, 171 (fig.)
aspinosa, 10, 172, 173 (fig.)
beameri, 10, 174, 175 (fig.)
boharti, 10, 174, 176 (fig.)
castanea, 10, 177, 178 (fig.)
coquilletti, 10, 177, 179 (fig.), 189
davisi, 10, 180, 181 (fig.)
dicksoni, 10, 182, 183 (fig.)
downeyi, 10, 184, 185 (fig.)
flocki, 10, 186, 187 (fig.)
griffint, 10, 186, 188 (fig.)
griseola, 10, 189, 190 (fig.), 212
halli, 10, 192, 198 (fig.)
helenae, 10, 194, 195 (fig.)
329

330

Brevitrichia—Continued
hodgdeni, 10, 194, 196 (fig.)
insulana, 10, 196, 197 (fig.)
kerni, 11, 196, 198 (fig.)
melandert, 11, 199, 200 (fig.)
minuta, 11, 201 (fig.)
oculiverida, 11, 202, 203 (fig.)
ordwayt, 11, 204, 205 (fig.)
pruinosa, 11, 206 (fig.)
schlingeri, 11, 207, 208 (fig.)
scitulaesca, 11, 209, 210 (fig.)
tumberlaket, 11, 210, 211 (fig.)
yucatani, 11, 212, 218 (fig.)
brunneus, Scenopinus, 8, 57, 59 (fig.)
bryanti, Pseudatrichia, 11, 231, 232
(fig.)
bryanti, Scenopinus, 9, 118, 119 (fig.)
bulbapennis, Scenopinus, 8, 60, 61 (fig.)
bulbosa, Metatrichia, 11, 214, 215, 216
(fig.), 219
bulbosus, Scenopinus, 215
bupennis, Pseudatrichia, 11, 232, 233
(fig.)
buscki, Scenopinus, 9,
butlerz, Scenopinus, 9,

119, 120 (fig.)
127,122: (fig)
caenofrons, Omphrale, 150
Caenoneura, 6, 10, 162
nigra, 10, 163 (fig.)
robusta, 10, 162 (fig.)
cajont, Pseudatrichia, 11, 234, 235 (fig.)
canarius, Scenopinus, 8, 60, 62 (fig.), 94
canei, Neopseudatriacia, 10, 300, 301
(fig.)
castanea, Brevitrichia, 10, 177, 178 (fig.)
cavifrons, Omphrale, 24
cavifrons, Scenopinus, 7, 24, (fig.)
Cerocatus, 1
chico, Scenopinus, 9, 122, 123 (fig.)
chilensis, Pseudatrichia, 286
chilensis, Heteromphrale, 12, 286, 288
(fig.)
chinchona, Seguyella, 12, 294, 295 (fig.)
civiculus, Scenopinus, 28
clausa, Pseudomphrale, 10, 164
clausus, Scenopinus, 164
cloudcrofti, Pseudatrichia, 11, 236, 237
(fig.)
cochisez, Scenopinus, 9, 124, (fig.), 139
cockerelli, Pseudatrichia, 11, 236, 238
(fig.)
collessi, Riekiella, 13, 307, 308 (fig.)

INDEX

complexa, Riekiella, 13, 305, 309, 310
(fig.)

Cona, 14

concava, Pseudatrichia, 11, 239, 240
(fig.)

convexa, Pseudatrichia, 11, 241, 242
(fig.)

cooki, Scenopinus, 9, 125, 126 (fig.)

coquilletti, Brevitrichia, 10, 177, 179
(fig.), 189

crenata, Omphrale, 164

crenata, Pseudomphrale, 10, 164

cretatus, Scenopinus, 7, 127 (fig.)

curticornis, Omphrale, 128

curticornis, Scenopinus, 9, 128, 129
(fig.)

curtipilosus, Scenopinus, 8, 63, 64 (fig.)
cyanops, Heteromphrale, 12, 286, 287
(fig.)

darwini, Scenopinus, 8, 64, 65 (fig.)

davisi, Brevitrichia, 10, 180, 181 (fig.)

dicksoni. Brevitrichia, 10, 182, 183 (fig.)

domesticus, Scenopinus, 26

downey?i, Brevitrichia, 10, 184, 185 (fig.)

dubiosa, Omphrale, 164

dubiosa, Pseudomphrale, 10, 164

dycei, Riekiella, 13, 305, 310, 311 (fig.)

eaithales, Pseudatrichia, 243
(fig.)

electus, Scenopinus, 9, 117, 130 (fig.)

efflatouni, Scenopinus, 7, 24, 25 (fig.)

evansi, Scenopinus, 8, 66, 67 (fig.)

energreent, Pseudatrichia, 51, 244, 245
(fig.)

11,..241,

fasciata, Atrichia, 26

fasciata, Scenopinus, 36

femoratus, Scenopinus, 9, 131

Fenestralis group, Scenopinus, 2, 7, 13,
15, 16 (key), 39

fenestralis, Musca, 15, 26, 36

fenestralis, Scenopinus, 1, 5, 7, 20, 22,
24,096, 27 (fig.), 28,238,436, 39,41

fijianus, Omphrale, 131

fijianus, Scenopinus, 9, 131 (fig.)

flandersi, Scenopinus, 9, 132 (fig.)

flaviceps, Pseudatrichia, 275, 278

flavipes, Omphrale, 68

flavipes, Scenopinus, 8, 68 (fig.)

flavoscutellata, Pseudomphrale, 318

INDEX

flavoscutellata, Stenomphrale, 13, 378, | insulana,

319 (fig.)

flocki, Brevitrichia, 10, 186, 187 (fig.)

fragosifrons, Scenopinus, 8, 68, 69 (fig.)

fraterna, Omphrale, 96

fraternus, Scenopinus, 9, 96 (fig.), 97
(fig.), 106

fryeri, Scenopinus, 9, 97, 98 (fig.), 99,
100, 106

furcinervis, Scenopinus, 26

fuscinervis, Scenopinus, 26

galactica, Omphrale, 294, 296

galactica, Seguyella, 12, 294, 296, 297
(fig.)

garrettt,
(fig.)

glabrifrons, Paromphrale, 28

glabrifrons, Scenopinus, 2, 5, 7, 28, 29
(fig.), 36, 44, 70, 76

glabrifrons var. surcoufi, Scenopinus, 28

gossypius, Scenopinus, 9, 133, 134 (fig.),
136

Pseudatrichia, 11, 244, 246

gracilipennis, Pseudatrichia, 11, 247,
248 (fig.)

graminicola, Scenopinus, 26

griffint, Brevitrichia, 10, 186, 188 (fig.)

griseola, Brevitrichia, 10, 189, 190 (fig.),

212
griseola, Pseudatrichia, 165, 174, 177,
180, 189

grisea, Omphrale, 28
griseus, Scenopinus, 7, 28, 30 (fig.)
gromieri, Scenopinus, 8, 70, 71 (fig.)

hagae, Scenopinus, 9, 135, 136 (fig.)
halli, Brevitrichia, 10, 192, 193 (fig.)
halteralis, Scenopinus, 9, 97, 98 (fig.)
halterata, Scenopinus, 28
helenae, Brevitrichia, 10, 194, 195 (fig.)
helenae, Pseudatrichia, 194
Heteromphrale, 12, 286

chilensis, 12, 286, 288 (fig.)

cyanops, 12, 286, 287 (fig.)

hodgdeni, Brevitrichia, 10, 194, 196
(fig.)

howdeni, Pseudatrichia, 11, 247, 249
(fig.)

howdeni, Scenopinus, 9, 137, 138 (fig.)
Hypseleura, 14

inquelinus, Scenopinus, 8, 72, 73 (fig.)
313-168 O—69—— 22

331

Brevitrichia, 10, 196, 197
(fig.)

insulana, Pseudatrichia, 196

jamest, Pseudatrichia, 11, 250, 251 (fig.)
japonicus, Omphrale, 31
japonicus, Scenopinus, 7, 31 (fig.)

katbergt, Scenopinus, 8, 72, 74 (fig.)

ketserz, Scenopinus, 8, 74, 75 (fig.)

kernt, Brevitrichia, 11, 196, 198 (fig.)

kewi, Neopseudatrichia, 12, 299, 300,
302 (fig.)

kuiterti, Omphrale, 139

laevifrons, Scenopinus, 28

Lagarinus, 1

leechi, Pseudatrichia, 11, 250, 252 (fig.)

Lepidomphrale, 2, 14

lesinensis, Scenopinus, 7, 31, 32 (fig.)

limpidipennis, Scenopinus, 8, 76

lincinus, Scenopinus, 9, 99, 100 (fig.),
106

lobosa, Paratrichia, 13, 320, 321 (fig.)

longirostris, Omphrale, 165

longirostris, Pseudomphrale, 10, 164

longiventris, Omphrale, 100

longiventris, Pseudatrichia, 11, 253, 254
(fig.), 265

longiventris, Riekiella, 13, 305, 312 (fig.)

longiventris, Scenopinus, 9, 97, 99, 100,
101 (fig.), 106

longurio, Atrichia, 221, 255

longurio, Pseudatrichia, 11,
(fig.)

lophrysoma, Metatrichia, 11, 215, 217
(fig.)

lophysoma, Pseudomphrale, 215

Lucidomphrale, 2, 15, 76

lucidus, Scenopinus, 2, 8, 76, 77 (fig.)

221, 255

madagascariensis, Scenopinus, 8, 76, 77
(fig.)
magdalenat,
(fig.)
magnicornis, Omphrale, 142
magnicornis, Scenopinus, 9, 142 (fig.)
mariaensis, Neopseudatrichia, 12, 303,
304 (fig.)
mariaensis, Pseudatrichia, 300, 303
mateui, Prepseudatrichia, 12, 286, 289,
290 (fig.)

Scenopinus, 9, 140, 141

302 INDEX

melanderi, Brevitrichia, 11, 199, 200|nitidulus, Scenopinus, 9, 96, 99,

(fig.)

104 (fig.), 106

melanderi, Pseudatrichia, 11, 256, 257|nordeni, Pseudatrichia, 11, 260,

(fig.)
Metatrichia, 11, 214 (key), 221
africana, 219, 221
bulbosa, 11, 214, 215, 216 (fig.) 219
lophrysoma, 11, 215, 217 (fig.)
robusta, 11, 215, 218 (fig.), 219
stevensoni, 11, 219, 220 (fig.)
waterhousei, 11, 221, 222 (fig.)
microgaster, Omphrale, 33
microgaster, Scenopinus, 7, 33, 34 (fig.)
minuscula, Omphrale, 100
minusculus, Scenopinus, 9, 100, 102
(fig.)
minuta, Brevitrichia, 11, 201 (fig.)
minutus, Scenopinus, 11, 78, 79 (fig.)
mirabilis, Scenopinus, 10, 121, 125, 143
(fig.), 152, 156, 161, 168
monodi, Omphrale, 80
monodi, Scenopinus, 8, 80 (fig.)
morlani, Pseudatrichia, 11, 258, 259
(fig.)
Musca
albicincta, 49
fenestralis, 15, 26, 36
rugosa, 36
saltitans, 26
senilis, 26
spoliata, 26
“tarda; 26

Nemotelus
niger, 36
tarsata, 36

Neopseudatrichia, 12, 299 (key), 318, 319
canei, 12, 300, 301 (fig.)
kewi, 12, 299, 300, 302 (fig.)
mariaensis, 12, 300, 303, 304 (fig.)

nevius, Scenopinus, 2, 8, 81 (fig.)

nidorcaupulus, Scenopinus, 7, 33, 35
(fig.)

niger, Nemotelus, 36

niger, Scenopinus, 2, 7, 15, 36, 37
(fig.), 85, 113

nigra, Caenoneura, 10, 163 (fig.)

nigripes, Scenopinus, 36

nitidifrons, Omphrale, 101

nitidifrons, Scenopinus, 9, 101, 107
(fig.)

(fig.)
norrisi, Scenopinus, 8, 82 (fig.)
nubilipes, Omphrale, 157
nubilipes, Scenopinus, 10, 143,
(fig.), 156
nubilipes, Omphrale, 157

oculivirida, Brevitrichia, 11, 202,
(fig.)

oldenbergi, Omphrale, 83

oldenbergi, Scenopinus, 8, 83 (fig.)

Omphrale, 1, 14
aethiopicus, 48
angustifrons, 51
antennata, 19
beameri, 117
beameri var. fusca, 117
bouvieri, 57
brunnea, 57
caenofrons, 150
cavifrons, 24
crenata, 164
curticornis, 128
dubiosa, 164
fijianus, 131
flavipes, 68
fraterna, 96
galactica, 294, 296
grisea, 28
japonicus, 31
kuiterti, 139
longirostris, 165
longiventris, 100
magnicornis, 142
microgaster, 33
minuscula, 100
monodi, 80
nitidifrons, 101
nubilipes, 157
oldenbergi, 83
papuana, 85
patrizi, 87
physadius, 36
pilosa, 103
sinensis, 41
tarsalis, 107
valga, 155
velutina, 157
whittakeri, 161

103)

261

144

203

INDEX

Omphralosoma, 2, 15, 92, 115
albifasciata, 115
squamosa, 92
opaculus, Seenopinus, 7, 36, 38 (fig.)
opacus, Scenopinus, 8, 83, 84 (fig.)
orbita, Scenopinus, 28
ordwayi, Brevitrichia, 11, 204, 205 (fig.)

palaestinensis, Pseudomphrale, 10, 165,
166 (fig.)
pallidipennis, Scenopinula, 12, 304, 305
(fig.)
pallipes, Scenopinus, 26
palmulapalpus, Scenopinus, 8, 85, 86
(fig.)
papuana, Omphrale, 85
papuanus, Scenopinus, 8, 85, 86 (fig.)
parallelus, Scenopinus, 9, 103, 105 (fig.)
Paratrichia, 13, 319, 320
lobosa, 53, 320, 321 (fig.)
westralica, 13, 322 (fig.)
Paromphrale, 2, 15
antennata, 19
glabrifrons, 28
parva, Pseudatrichia, 11, 260, 262 (fig.)
parvipennis, Pseudatrichia, 11, 260, 263
(fig.)
bryanti, 11, 231, 232 (fig.)
bupennis, 11, 232, 233 (fig.)
cajoni, 11, 234, 235 (fig.)
chilensis, 286
cloudcroftt, 11, 236, 237 (fig.)
cockerelli, 11, 236, 238 (fig.)
concava, 11, 239, 240 (fig.)
convexa, 11, 241, 242 (fig.)
eaithales, 11, 241, 248 (fig.)
evergreent, 11, 244, 245 (fig.)
flaviceps, 275, 278
garretli, 15, 244, 246 (fig.)
gracilipennis, 11, 247, 248 (fig.)
griseola, 165, 174, 177, 180, 189
helenae, 194
howdeni, 11, 247, 249 (fig.)
insulana, 196
jamesi, 11, 250, 255 (fig.)
leechi, 11, 260, 252 (fig.)
longiventris, 11, 2538, 254 (fig.), 265,
266
longurio, 11, 221, 256 (fig.)
mariaensis, 257, 300, 303
melanderi, 11, 256, 257 (fig.)
patrizi, Omphrale, 87

333

patrizi, Scenopinus, 8, 87, 88 (fig.)
pecki, Scenopinus, 10, 145, 146 (fig.)
perkinsi, Scenopinus, 8, 26, 88, 89 (fig.)
peromysct, Pseudatrichia, 12, 264 (fig.)
phaidimos, Scenopinus, 8, 70, 88, 89
(fig.)
physadius, Omphrale, 36
physadius, Scenopinus, 7, 36, 40 (fig.)
pilosa, Belosta, 12, 275, 277 (fig.)
pilosa, Omphrale, 103
pilosa, Pseudatrichia, 275
pilosus, Scenopinus, 9, 103, 105 (fig.)
Prepseudatrichia, 6, 12, 286, 289 (key)
mateut, 12, 286, 289, 290 (fig.)
stenogaster, 12, 291, 292 (fig.)
violacea, 12, 291, 293 (fig.)
Propebrevitrichia, 12, 281, 283 (key), 286
stuckenbergi, 12, 283 (fig.)
turneri, 12, 281, 284, 285 (fig.)
pruinosa, Brevitrichia, 11, 206 (fig.)
Pseudatrichia 5, 11, 221, 223 (key), 274,
286, 291, 299, 318, 320
albocincta, 11, 225 (fig.)
atombomba, 11, 225, 226 (fig.)
bakerz, 11, 227, 228 (fig.)
barnesi, 11, 229 (fig.)
biacristerna, 11, 230, 231 (fig.)
morlant, 11, 268, 259 (fig.)
nordent, 11, 260, 261 (fig.)
parva, 11, 260, 262 (fig.)
parvipennis, 11, 260, 263 (fig.)
peromysci, 12, 264 (fig.)
pilosa, 275
punctulata, 12, 253, 265, 266 (fig.)
sabroski, 12, 266, 267 (fig.)
saccharcupa, 12, 268, 269 (fig.)
toupeta, 12, 270, 271 (fig.)
truncata, 12, 270, 272 (fig.)
unicolor, 12, 225, 236, 258, 273 (fig.)
Pseudomphrale 6, 10, 164, 221
africana, 219
clausa, 10, 164
crenata, 10, 164
dubiosa, 10, 164
flavoscutellata, 318
longirostris, 10, 165
lophrysoma, 215
palaestinensis, 10, 165, 166 (fig.)
stevensoni, 219
teutankhameni, 318, 319
waterhousei, 221
westralica, 322

304

INDEX

punctulata, Pseudatrichia, 12, 253, 265, | Seenopinus—Continued

266 (fig.)
pygmaeus,
(fig.), 148

Scenopinus, 10, 128, 147

quadrifida, Riekiella, 13, 305, 309, 313,
314 (fig.)

ramaleyi, Scenopinus, 143
reduncus, Scenopinus, 9, 106 (fig.)
Riekiella, 13, 304, 306 (key)
bicornis, 13, 304, 305, 306, 307 (fig.),
313
collessi, 13, 307, 308 (fig.)
complexa, 13, 305, 309, 310 (fig.)
dycet, 13, 305, 310, 311 (fig.)
longiventris, 13, 305, 312 (fig.)
quadrifida, 13, 305, 309, 313, 314 (fig.)
taylorz, 13, 315, 316 (fig.)
uncata, 13, 315, 317 (fig.)
robusta, Caenoneura, 10, 162, (fig.)
robusta, Metatrichia, 11, 215, 218 (fig.),
219
rossi, Scenopinus, 10, 148, 149 (fig.)

sabroskyi, Pseudatrichia, 12, 266, 267
(fig.)
saccharcupa, Pseudatrichia, 12, 268, 269
(fig.)
saint, Scenopinus, 8, 39, 42 (fig.)
saltitans, Musca, 26
Scaenopius, 15
Scenopinula, 12, 304
pallidipennis, 12, 304, 305 (fig.)
Scenopinidae, 1, 5 (fig.), 6, 7, 13 (key)
Scenopinus, 2, 7, 14, 76, 92, 99, 286, 289,
291, 294, 304
bulbosus, 215
clausus, 164
stenogaster, 291
Scenopinus, Albicinctus group, 2, 8, 15,
45 (key), 55, 57, 76, 83, 92
aethiopicus, 8, 48, 49 (fig.)
albicinctus, 8, 49, 50 (fig.)
angustifrons, 8, 51 (fig.)
anthrax, 8, 51, 52 (fig.)
balteatus, 8, 53, 54 (fig.), 57, 66
biroi, 8, 53, 55 (fig.)
bolgarti, 8, 55, 56 (fig.)
bouvieri, 8, 57, 58 (fig.)
brevicornis, 60
brunneus, 8, 57, 59 (fig.)
bulbapennis, 8, 60, 61, (fig.)

canarius, 8, 60, 62 (fig.), 94
curtipilosus, 8, 63, 64 (fig.)
darwini, 8, 64, 65 (fig.)
evansi, 8, 66, 67 (fig.)
flavipes, 8, 68 (fig.)
fragosifrons, 8, 68, 69 (fig.)
gromieri, 8, 70, 71 (fig.)
inquelinus, 8, 72, 73 (fig.)
katbergi, 8, 72, 74 (fig.)
keisert, 8, 74, 75 (fig.)
limpidipennis, 8, 76
lucidus, 2, 8, 76, 77 (fig.)
madagascariensis, 8, 76, 78 (fig.)
minutus, 8, 78, 79 (fig.)
monodi, 8, 80 (fig.)
niger, 85
niveus, 2, 8, 81 (fig.)
norrist, 8, 82 (fig.)
oldenbergi, 8, 83 (fig.)
opacus, 8, 83, 84 (fig.)
palmulapalpus, 8, 85, 86 (fig.)
papuanus, 8, 85, 87 (fig.), 113
patrizi, 8, 87, 88 (fig.)
perkinsi, 8, 88, 89 (fig.)
phaidimos, 8, 70, 88, 89 (fig.)
spurrelli, 8, 90, 91 (fig.)
squamosus, 2, 8, 92, (fig.)
zelleri, 49
zeylanicus, 85, 86
Scenopinus, Brevicornis group, 9, 14,
93 (key)
brevicornis, 9, 60, 94 (fig.), 95 (fig.)
fraternus, 9, 96 (fig.), 97 (fig.), 99, 106
fryeri, 9, 97, 98 (fig.), 99, 100, 106
halteralis, 9, 98, 99 (fig.)
lincinus, 9, 99, 100 (fig.), 106
longiventris, 9, 97, 100, 101 (fig.), 106
minusculus, 9, 100, 102 (fig.)
nitidifrons, 9, 101, 107
nitidulus, 9, 99, 103, 104 (fig.), 106
parallelus, 9, 103, 105 (fig.)
pilosus, 9, 103, 105 (fig.)
reduncus, 9, 99, 106 (fig.)
tarsalis, 9, 101, 107 (fig.)
turnert, 9, 107, 108 (fig.)
unifasciatus, 9, 108, 109 (fig.)
Scenopinus, Fenestralis group, 2, 7,
15, 16 (key), 39
albicomus, 7, 17, 18 (fig.)
antennatus, 7, 19 (fig.)
aquelonius, 7, 20, 21 (fig.)

INDEX

Scenopinus—Continued

atra, 36

breviterminus, 7, 22, 23 (fig.)

cavifrons, 7, 24 (fig.)

civiculus, 28

domesticus, 26

efflatount, 7, 24, 25 (fig.)

fasciata, 26, 36

fenestralis, 1, 5, 7, 15, 20, 22, 24,
26, 27 (fig.), 28, 33, 39, 41

furcinervis, 26

glabrifrons, 2, 5, 7, 15, 28, 29 (fig.),
36, 44, 70, 76

glabrifrons var. surcoufi, 28

graminicola, 26

griseus, 7, 28, 30 (fig.)

halterata, 28

japonicus, 7, 31, (fig.)

leavifrons, 28

lesinensis, 7, 31, 32 (fig.)

microgaster, 7, 33, 34 (fig.)

nidorcaupulus, 7, 33, 35 (fig.)

niger, 2, 7, 15, 36, 37 (fig.), 85, 113

nigripes, 36

opaculus, 7, 36, 38 (fig.)

orbita, 28

pallipes, 26

perkinsi, 26

physadius, 7, 36, 40 (fig.)

rufitarsis, 26

rugosa, 36

saint, 8, 39, 42 (fig.)

saltitans, 26

schroederi, 8, 39, 43 (fig.)

scutellatus, 26

scutellatus var. nigroscutellatus, 26

senilis, 26

sinensis, 8, 41, 43 (fig.)

spoliata, 26

sulcicollis, 26

“tarda,’’ 26

tarsata, 26

undulafrons, 8, 41, 44 (fig.)

varipes, 8, 44

vitripennis, 8, 28, 44, 45 (fig.)

Scenopinus, Velutinus group, 2, 9, 16,

110 (key), 115

adventicius, 9, 113 (fig.)

alalcteus, 9, 114 (fig.)

albidipennis, 147, 148

albifasciatus, 2, 9, 115, 116 (fig.)

barberi, 9, 117, 116 (fig.)

3395

Scenopinus—Continued
beameri, 9, 117, 118 (fig.)
beameri var. fuscus, 9, 117, 118 (fig.)
bryanti, 9, 118, 119 (fig.)
busckt, 9, 119, 120 (fig.)
butlerz, 9, 121, 122 (fig.)
caenofrons, 150
chico, 9, 122, 123 (fig.)
cochiset, 9, 124 (fig.), 139
cooki, 9, 125, 126 (fig.)
cretatus, 9, 127, (fig.)
curticornis, 9, 128, 129 (fig.)
electus, 9, 117, 130 (fig.)
femoratus, 9, 131
fijianus, 9, 131 (fig.)
flandersi, 9, 132 (fig.)
gossypius, 9, 133, 134 (fig.), 136
hagai, 9, 135, 136 (fig.)
howdeni, 9, 137, 188 (fig.)
kuiterti, 9, 139 (fig.)
magdalenai, 9, 140, 141 (fig.)
magnicornis, 9, 142 (fig.)
mirabilis, 10, 121, 125, 143 (fig.), 152,
156, 158, 161
nubilipes, 10, 148, 144 (fig.), 156, 157
pecki, 10, 145, 146 (fig.)
pygmaeus, 10, 128, 142, 147 (fig.)
ramaleyi, 143
rosst, 10, 148, 149 (fig.)
schulzi, 10, 150, 151 (fig.)
seftoni, 10, 152 (fig.)
serratus, 10, 153, 154 (fig.)
valgus, 10, 155 (fig.)
vanduzeei, 10, 156, 157 (fig.)
velutinus, 10, 134, 142, 157, 158 (fig.)
weemst, 10, 158, 159 (fig.)
werneri, 10, 160 (fig.)
whittakeri, 10, 161 (fig.)
Scenopoeus, 14
schlingeri, Brevitrichia,
(fig.)
schroederi, Scenopinus, 8, 39, 43 (fig.)
schulzi, Scenopinus, 10, 150, 151 (fig.)

11, 207, 208

scitulaesca, Brevitrichia, 11, 209, 210
(fig.)
scutellatus, Scenopinus, 26
scutellatus var. nigroscullatus, Sceno-

pinus, 26
seftoni, Scenopinus, 10, 152 (fig.)
Seguyella, 12, 294 (key)
chinchona, 12, 294, 295 (fig.)
galactica, 12, 294, 296, 297 (fig.)

336

Seguyella—C ontinued
turneri, 12, 296, 298 (fig.)

senilis, Musca, 26

serratus, Scenopinus, 10, 153, 154 (fig.)

sinensis, Omphrale, 41

sinensis, Scenopinus, 8, 41, 43 (fig.)

spoliata, Musca, 26

spurrelli, Scenopinus, 8, 90, 91 (fig.)

squamosa, Omphralosoma, 92

squamosus, Scenopinus, 2, 8, 92 (fig.), 97

stenogaster, Prepseudatrichia, 12, 291,
292 (fig.)

stenogaster, Scenopinus, 291

Stenomphrale, 3, 13, 318
flavoscutellata, 13, 318, 319 (fig.)
teutankhameni, 13, 318, 319, 320

(fig.)

stevensoni, Metatrichia, 11, 219, 220
(fig.)

stevensoni, Pseudomphrale, 219

stuckenbergi, Propebrevitrichia, 12, 283
(fig.)

sulcicollis, Scenopinus, 26

“tarda,’? Musca, 26

tarsalis, Omphrale, 107

tarsalis, Scenopinus, 9, 101, 107 (fig.)

tarsata, Nemotelus, 36

taylori, Riekiella, 13, 315, 316 (fig.)

telfordi, Belosta, 12, 278, 279 (fig.)

teutankhameni, Pseudomphrale,
319

teutankhameni, Stenomphrale, 13, 318,
319, 320 (fig.)

timberlakei, Brevitrichia, 11, 210, 211
(fig.)

toupeta, Pseudatrichia, 12, 270, 271 (fig.)

turneri, Propebrevitrichia, 12, 281, 284,
285 (fig.)

turnert, Scenopinus, 9, 107, 108 (fig.)

318,

INDEX

turnert, Seguyella, 12, 296, 298 (fig.)
truncata, Pseudatrichia, 12, 270, 272
(fig.)

uncata, Riekiella, 13, 315, 317 (fig.)

undulafrons, Scenopinus, 8, 41, 44 (fig.)

unicolor, Pseudatrichia, 12, 225, 236,
258, 278 (fig.)

unifaciatus, Scenopinus, 9,
(fig.)

108, 109

valga, Omphrale, 155

valgus, Scenopinus, 10, 155 (fig.)

vanduzeet, Scenopinus, 10, 156, 157 (fig.)

varipes, Scenopinus, 8, 44

Velutinus group, Scenopinus, 2, 9. 16,
110 (key), 115

velutina, Omphrale, 157

velutinus, Scenopinus, 10, 142, 157, 158
(fig.)

violacea, Prepseudatrichia, 12, 291, 293
(fig.)

viticolapennis, Belosta, 12, 281, 282 (fig.)

vitripennis, Scenopinus, 8, 28, 44, 45
(fig.)

waterhousei, Metatrichia, 11, 221, 222
(fig)

waterhousei, Pseudomphrale, 221

weemsi, Scenopinus, 10, 158, 159 (fig.)

werneri, Scenopinus, 10, 160 (fig.)

westralica, Paratrichia, 13, 322 (fig.)

westralica, Pseudomphrale, 322

whittakeri, Omphrale, 161

whittakeri, Scenopinus, 10, 161 (fig.)

yucatani, Brevitrichia, 11, 212, 213 (fig.)

zelleri, Scenopinus, 49
zeylanicus, Scenopinus, 85, 86

U.S. GOVERNMENT PRINTING OFFICE: 1969 O—313-168

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TTT,
3 9088 01 698

421 3