MT eae eet erin? ae ¢ ty Raa ona et RSS ats a Maan tines eens Se RNG S a > 4 % Siac Va a Ran eae Gy SECT tS Sauer Rect et iN shoes Sea Rea Sat att oa beat Petit ne) SS sak toe Beans mye Pen Ar ex xt’ Les re Aman Pm cee Lay RINT : Hesttg ‘ MY ASE hs hae ED SAA C tHe a Eeisestc \ aoa mets ni i a cn ‘ : ie pies pee oat Pe ne a iv ta i » q A ie Pee’ ‘ gy nr . w Le 4 R ° : ‘ ‘ , * \ ' « . { i . | ) oar he i fy i ; _. | , i) | ik | (a 1 uy Ve my me as ‘ y ‘ ! q — Va a 1) } hee is hi és i i , a. I, f ; \ % i A J . It n ia ; os i i) ie . } i, , t , h , 1 , ¥ | i | os f . SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM | Bulletin 63 ~ A MONOGRAPHIC REVISION OF THE COLE- OPTERA BELONGING TO THE TENEBRI- ONIDE TRIBE ELEODIINI INHABITING THE UNITED STATES, LOWER CALI- FORNIA, AND ADJACENT ISLANDS BY FRANK E. BLAISDELL, SR. Of San Francisco, California SESE ING RE oo Soe Ase x 2 yO E , | WASHINGTON b GOVERNMENT PRINTING OFFICE 1909 SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM Bulletin 63 A MONOGRAPHIC REVISION OF iia, COLE OPTERA BELONGING TO THE TENEBRI- ONIDE TRIBE ELEODIINI INHABITING THE UNITED STATES, LOWER CALI- FORNIA, AND ADJACENT ISLANDS BY FRANK E. BLAISDELL, SR. Of San Francisco, California W ASHINGTON GOVERNMENT PRINTING OFFICE 1909 A0TS4A BULLETIN OF THE UNITED STATES NATIONAL MUSEUM. IssuEpD JUNE 24, 1909. IL ADVERTISEMENT. The scientific publications of the National Museum consist of two series—the Bulletin and the Proceedings. The Bulletin, publication of which was begun in 1875, is a series of more or less extensive works intended to illustrate the collections of the U. S. National Museum and, with the exception noted below, is issued separately. These bulletins are monographic in scope and are devoted principally to the discussion of large zoological and botanical groups, faunas and floras, bibliographies of eminent natural- ists, reports of expeditions, etc. They are usually of octavo size, although a quarto form, known as the Special Bulletin, has been adopted in a few instances in which a larger page was deemed indis- pensable. This work forms No. 63 of the Bulletin series. Since 1902 the volumes of the series known as ‘‘Contributions from the National Herbarium,” and containing papers relating to the botan- ical collections of the Museum, have been published as bulletins. The Proceedings, the first volume of which was issued in 1878, are intended as a medium of publication of brief original papers based on the collections of the National Museum, and setting forth newly acquired facts in biology, anthropology, and geology derived there- from, or containing descriptions of new forms and revisions of limited groups. A volume is issued annually, or oftener, for distribution to libraries and scientific establishments, and in view of the impor- tance of the more prompt dissemination of new facts a limited edition of each paper is printed in pamphlet form in advance. RicHarp RaATHBUN, Assistant Secretary, Smithsonian Institution, In Charge of the United States National Museum. WasHInGTON, U.S. A., May 15, 1909. iil PREFACE, In 1901 when work was first seriously begun on the Eleodiini, a bibliography was prepared. The great conflagration in San Francisco that followed the earthquake of 1906 destroyed the California Academy of Sciences with its library, so that when the present monograph was about completed there were no library facilities at hand, nor was it possible for me to review the literature elsewhere. The manuscript was therefore submitted for publication without a final review. Accordingly Mr. E. A. Schwarz, the eminent coleopterist at the U.S. National Museum, most kindly undertook the task for me, and to him I extend my most sincere thanks for the correction of the bibliography, for many suggestions, and for rearranging certain parts of the work. In the text I have applied the term ‘‘heterotype”’ to the individuals that form the extremes of a specific or a varietal series. The hetero- types therefore, represent the extreme modifications in size, form, sculpturing, and color. Theindividuals connecting the extremes of a series constitute the intermediates or ‘‘mesotypes.” Frequently individuals of one species are observed to simulate those of another species in form and sculpturmg; an individual of this character has been called an ‘‘amphitype.” A unique, when serving as a type, is spoken of as a ‘“‘monotype.”’ When more than one or a series of idividuals serve as a basis for a specific or varietal description, they are referred to as ‘‘polytypes;” if a male and female serve they are ‘“‘sexitypes;” otherwise they are ‘‘co-types.”’ Furthermore, in order to systematically deal with the many minor degrees of divergencies exhibited by individuals that are presumably the progeny of parents specifically or racially identical, I have deemed it conservative and scientific to.recognize incipient races and incipient subraces. In accordance with that view I have used the name forma to make it possible to relatively define aggregations of individuals possessing some particular or salient characteristic; it is believed that these divergent characters have arisen through local or general cli- matic or environmental conditions acting upon the progeny of parents specifically or racially identical. AVall PREFACE. In other words, the several formx enumerated under Fleodes den- tipes may arise from eggs deposited by a single typical female, under the influence of climatic, geographical, or environmental conditions capable of producing such divergencies. A Latin name has been used to express the salient characteristic of any given forma, and these characteristics may be similar for each species or variety. A species or variety may have smooth and rough, short and elongate, small and large, caudate or ecaudate forms, consquently a repetition of Latin names is called for. These terms are absolutely synonymous with the specific or varietal terms and must not be perpetuated as distinct grades, but simply used as an aid in recording data, and as a compromise between unscientific lumping and splitting, or the difference between 47 and 400 species. Logically the present treatment ought not to seriously collide with the rules of the International Code governing nomenclature. Frank E. BLAISDELL, Sr. TABLE OF CONTENTS. Rein WIG tO E per ae ee pee eee) eee ote Re re le ee iS Hue Katt OWL CPO GEE en sae Sete ek pe ie ee HES ue eye. Paul urs eee 2 af aye - Meet ocis7Os BU Gagee ee te eh ere ee yk ee cae ee Ses anes Seen Cralge x UCEH 2 WATT LOMUY Set. cyst Sage hah 2 ioe, elena S eee nate eS Aaa lata jopew ls - DinprmicaiiGn Alldesv Masia a) oss eee ans See heey on aE onetime arnt p inl yincalkey to penera,ot Hleodiini. 23. S25. os. Dy eee a cle oe 22 oss iGemuse leat Comet tees et ok Fee et Gere as See aay Sea Se Analytical key to subgenera of Picadas AEE Cpe Sn Se are eed eee hes nls Synopsis of general subgeneric characters...5........-.-...-.-----+------ Subgenus Melaneleodesisan2 tcc sigs bac ees ook ese ease ea he oe Growpel:— Carbonara SeChlONe ce 4-= oes pane ook Giga toe Soke oe ae Bleodesid eo iliste seen eee ere ee ie oe eats a Se Blead CarCar bomen stp ccepte mente oi: Aor ee rer wey auiers aicn, Se Bleodestcanbonattanvanrs SOLOG=] 94-45 ee ae) eee nee Ne odeaieini tare cten ie oe eye nthe Ga Conk ae ot i oak Blecdestampla war: Golosaa- tor. s sah See eine Sas 8: BNO ESO DSOLC LA ae ee ele Um NE od ac eee ee a Rleodes.obsoleta: var. poreatacs =. 22 55.52 Seo S82 see ReGen Ka LVRS ee iC airs Se RE ate ANC ne EN At cc Te Hleocdessomnlasaes sae Geer scctee es eee wee ine Se BeeS te adit Hileodesiomuissan vars py omecdee senate aes oe ee ae Hleodes:omissaqvars peningulariss. 22.6.2. 22-22-25 52042-+-- Group I.—Quadricollis section... . . Fest teen Lee Se se es Eleodes quadricollis. . ae TT ee AW SS ee Na Epipleure very narrow, not attaining the humeral angles______ Hmbaphion. 3. Epipleurze broader at base, more or less gradually narrowing to apex. Epipleurze occupying only a part of the inflexed portion of the elytra ; buccal processes of the genz not produced________________ Hleodes. Epipleure occupying the whole of the inflexed portion of the elytra; buccal processes of the gene acutely produced________ EBleodimorpha., 30 - BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The genera can also be defined by the male genital characters as follows: Sternite (fig. 6, Plate 6) with lobes fully and strongly connate; clave dorsal to the edeagus, connate forming a pallium (similis, mucronata) (Blaptini) Blaps. Sternite distinctly bilobed, lobes connected by a membrane that at most is but feebly chitinized, lobes capable of abduction (EHleodiini). Clavie ventrad to the edeagus. Clavee spiculiform, notched at side of the tips; alee more or less strongly inflexed, not prominent ventrally, plane or somewhat concave, never strongly (convex {CB late te etic iy) es ee ee Eleodes. Clavie flattened (liguliform), of equal width throughout, tip obtuse and rounded; alze short, not strongly inflexed, giving attachment to subale; \(PlGtEr, es Sas a eee Eleodimorpha. Clave laterad to the edeagus, truncate at tip, more or less concave beneath ; ale moderately and evenly inflexed, rather prominent ventrally (Plate Cg BRS TY sc a ge ye ee wee Hmbaphion. Trogloderus has the lobes of the sternite united by membrane and capable of abduction. Unfortunately the clave could not be studied without destruction of the fragile material at. hand. Genealogy—The antecedents of the present tribe are not suggested to my mind by the data at hand. The male sexual characters are the most distinctly differentiated for the generic subdivision of the tribe. The above synoptic table indicates these relationships just as forcibly as a diagram could do where there are so few elements involved. The genera are related, first, by the bilobed sternite, the lobes being capable of abduction; second, by the relation of the clave to the edeagus; third, by divergence having taken place along the lne of degree of development in the ale. In Eleodimorpha the subale have been differentiated as distinct sclerites from the ale. In Lleodes and Embaphion the extrusion of the edeagus with simultaneous abduction of the clave, is brought about by muscular action—certain extrusor muscles; the return of these structures is accomplished by the inherent elasticity of the ale, the latter being the continuously produced and inflexed sides of the apicale. It is logical to suppose that accompanying the differentiation of the subale as distinct exoskeletal elements that there has been a corresponding muscular differentiation; for when the continuity of the ale underwent solution, they in all probability were no longer capable to functionate by their own inherent resiliency—of course, it is possible for the continuity of the elements to be restored by becoming connate at a sutural line; but this would evidently still be a weak point and correspondingly diminish the functional activity of the part. i i it i i all ail ee maa REVISION OF ELEODIINI—BLAISDELL. aL In all probability there has been a compensatory development of retractor muscles to overcome this loss of power. Genus ELEODES Esehseholtz. Mentum variable in form, more or less trilobed, the middle lobe larger and more convex; lateral lobes frequently inflexed and more or less concealed. Last joint of labial palpi triangular, narrower than the maxillary, which are broadly triangular or securiform. Lead not deeply inserted, suture between epistoma and front fre- quently distinct; epistoma truncate or feebiy emarginate, sides straight or sinuate and converging anteriorly. Eyes narrow, subreni- form. Antenne eleven jointed, the last three or four usually notably compressed, last joint frequently though not always smaller than the preceding. Prothorax variable in form, usually applied against the bases of the elytra. Scutellum always present, distinct and equilateral. Elytra of variable form and sculpture, in many species prolonged into a cauda behind; epipleure always distinct, broader at base and always visible at the humeral angles, gradually narrowing to apex. Legs moderately long, but somewhat variable in this respect; femora never strongly clavate, profemora most so, the mesofemora slightly, the profemora frequently armed in one or both sexes with a tooth of variable form, rarely all the femora armed; tarsi usually channelled and setose beneath, first and second joints of the protarsi frequently slightly thickened beneath and often the first two or three joints are clothed with a tuft of silken hairs, or dilated and densely covered with a spongy pubescence beneath. Spurs of the meso- and metatibie well developed, moderately slender, narrowing from base to apex, acute, the internal more or less longer than the external. Varying more or less in the different species. The above characters are those applicable to the whole genus; characters peculiar to certain groups will be given as of subgeneric value. The genus /leodes is one of the largest of the Tenebrionide and contains many more species than all of the other genera of the present tribe put together (United States). It is also the most polymor- phous and has been the most difficult to subdivide of any in our fauna. The study of the genitalia has greatly reduced this difficulty, but extreme variations in many individuals of each species will yet be a source of great perplexity. The most salient extreme characters observed in the genus may now be reviewed : Color.—Black, sometimes reddish along the elytral suture as in acuta, obsoleta, and hispilabris,; rarely fusco-piceous as in longicollis 32 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. and énsularis, probably from immaturity, or from aging in the cabinet ; in one instance the elytra are described as castaneous (sub- cylindrica Casey) ; this unusual coloration is undoubtedly due to im- maturity. Dead specimens from long exposure to climatic inclemency become fuscous and even castaneous. In a lmited number of species the legs are dark brown throughout, as in brunnipes Casey. Form.—Notably large and elongate as in the males of gigantea, oblong as in tricostata, elongately oblong as in acuta and texana, oval in veterator, inflated as in ventricosa and depressed in tibialis. Size.—Largest in acuta, texana, and grandicollis; smallest in clavi- COrNIS. Surface.—Glabrous and shining, as in dentipes, gigantea, and omissa,; opaque as in tricostata and opaca. Sculpturing—Whule usually simple it becomes tuberculate in gra- nosa and granulata; rugoso-muricate in cordata and muricate to a varying degree in many. Pubescence.—Body clothed with long flying hairs as in longipilosa, pilosa, and hirsuta, or confined to the elytra as in letcheri; hairs short, recumbent and abundant in veterator and opaca, Characters that vary within specific limits ——These are the factors which make the limitations of species extremely difficult. Each species may have its extreme large (gigantism) and small (nanism) forms; its smooth and rough forms, elongate and robust forms, while the sculpturing varies from comparatively smooth to rough inde- pendently of size or form. This can be accounted for by environment to a great extent and to certain inherent factors. With robustness there is a relative shortening of the appendages and also of their segments, as for instance the joints of the antenne from the fourth to the seventh or eighth tend toward equality of length, while in the elongate forms there is a more or less inequality of length evident, the joints decreasing in length more or less from the fourth to the eighth, these joints being comparatively and rela- tively longer as a part of the general habitus. These remarks apply most emphatically to the tarsi and their joints as well; the antenne and legs as a whole exhibit this variation of length; the head varies in size and relative length and breadth even in any particular form. So it can be seen why in this particular genus (and this fact is worthy of broader application) that the relative length of parts of any appendage can not always be given as evidence of specific dis- tinctness. These conclusions have been arrived at after a long and tedious recourse to micrometrical treatment of hundreds of specimens. In many specimens, especially in the subgenus /eodes, there is to be observed a more or less evident smooth median pronotal line REVISION OF ELEODIINI—BLAISDELL. 30 that may extend from the apical to the basal margins; this line is most pronounced as a rule in 2. acuticauda var. laticollis, where it often becomes a distinctly impressed line; usually it is only observable as a line in surface luster. The significance of this character is obscure. Another very interesting character is that observed in the hind tibiz of the males of certain species of the subgenus H'leodes. The metatibie are slightly arcuate and narrow in basal half, rather ab- ruptly dilated and straight in apical half. In the literature I find this character mentioned by Doctor Horn as one of the male char- acters of . wickhami. This is not a staple nor a specific character, but 1s observed as peculiar to the male of certain species, but not in all forms of those species; it is seen in the caudate forms of F’. esch- scholtzi and luce, in armata and in the elongate form of dentipes; other males of the same species have the metatibiz perfectly straight, some have them arcuate in basal half but not abruptly dilated apically. Individuals of many species exhibit what must be considered as a normal tibial arcuation, which must be distinguished from a common fortuitous arcuateness; the latter usually appears exaggerated and is generally a part of a general abnormal habitus, and it frequently accompanies unusual shortness of the legs. ANALYTICAL KEY TO SUBGENERA OF ELEODES. The genus Hleodes may be subdivided by clearly defined female genital char- acters into the following subgenera: Internal lobe of the valvular apex well developed, membranous, although fre- quently chitinous; external lobe obsolete; appendage directed backward ANTON CH SOULE eRNTS Cl mee aes ee es ee ae nm eae ee ee SE ee fk ee ee be Bee 1 Wxternal apical lobe well developed, always membranous; internal lobe feebly developed; appendage directed backward and inward____-~~~1 Melaneleodes. 1. Superior pudendal membrane not broadly exposed________________________ 2 Superior pudendal membrane broadly or fully exposed; body and apical lobe of valve fully chitinized; appendage minute _____________ Steneleodes. 2. Appendages present, apical lobe defined from body of the valve____________ 3 Appendages obsolete; apical lobe not defined from body of the valve. Heteropromus. 3. Appendages moderate in size, subconical to semiellipsoidal and flattened; inferior margins of the valves not contiguous throughout their entire length but separated by a fusiform genital fissure___________ 4 Appendages minute and punctiform; inferior margins of the valves con- tiguous throughout; genital fissure closed____________________ Discogenia. 4, Apical lobe subacute, triangular, membranous or more or less semichit- MGI AS lag sk ae Se ee Se te a eee es sees 5 59780—Bull. 68—09 3 34 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Apical lobe rounded at apex, depressed, well chitinized, more or less concave beneath conmyexinboVves 2:25 25 te ee Se eee \Wetablapylis. 5. Dorsal plate variable in form, not broadened externally, finely, very sparsely punctate, more or less feebly and very sparsely pubescent Dorsal plate broadened externally, margin strongly arcuate, rather coarsely punctate and sparsely clothed with rather long flying hairs on outer MOLety Ve! ee ee ee ee Promus. 6. Submarginal groove not visible from above; surface of dorsal plate at most but) feebly. concave: generally” planes a ee 1 Submarginal groove dorsal or visible at some point from above; surface of dorsal plate more or less deeply concave__—________-__= Eleodes. 7. Superior outline of genital segment quadrate, triangulo-parabolic to para- DOC 2s See Pelee ee Ee ee en ee ee ee 8 Superior outline more or less triangular. Outline triangular; dorsal plate oblong-triangular, external border at most but slightly arcuate, internal border not strongly arcuate dit) WaS@S ssc. es Fe ee aoe eee Blapylis. Outline subequilaterally triangular; external border of dorsal plate arcuate, angle obsolete; superior membrane reaching to middle Of dorsal™ plates a =e ee ee ee Tricheleodes. 8. Superior outline of genital segment quadrate to triangulo-parabolic; superior membrane variable in length; dorsal plate oval-oblong, internal border re- flexed and impunctate; external border more or less arcuate, angle evident; apex small and membranous; appendage large, more or less semieliiptical to“subconical= depressed 2. 222 oe ee ee a ee ee eee Litheleodes. Superior outline parabolic, short; dorsal plate subovate, extending upon the apical lobe beyond the appendage, sides not reflexed; appendage short, pro- jecting from beneath the outer border of the apical margin of dorsal plate; genitale fissure Subapi Callas ts a ee ee ee ee Pseudeleodes. The student must bear in mind in resorting to the use of the above table that care must be taken to familiarize himself with the general structure of the genital segment. Besides the distinct generic differences there are in each subgenus specific characters which can only be worked out positively with large series, and, while of scientific interest, would not be of practical im- portance in the separation of species. The male genital characters may be summarized as follows: Apicale of the edeagophore broadly triangular, depressed on dor- sum and more or less broadly membranous centrally; apex more or less rounded =22 3234523 eee eee eee eee Apicale triangular and slightly elongate, convex dorsally; apex more or less produced and subacute. Steneleodes. Metablapylis. Melaneleodes. Dorsum with an evident median, more or less membranous | Hleodes. STOOVE) beurd Soak ee ee a ee et ar cen al Litheleodes. Promus. Dorsum without median groove, except in tibialis___.____________ Blapylis. REVISION OF ELEODIINI—BLAISDELL. 35 SYNOPSIS OF GENERAL SUBGENERIC CHARACTERS, Anterior femora at least, armed in both sexes (except in caudifera and longi- PiUOsMaN Were wine n Lectin aren VoUTaHvie) l= sw n-.2 = et hee Pt ae Hleodes. Anterior femora armed only in the male or mutic. Anterior tibial spurs dissimilar in the sexes; femora mutic__J/elaneleodes. Anterior tibial spurs similar in the sexes. Tarsi similar in the sexes, or nearly so. Middle lobe of the mentum large, lateral lobes rudimentary, in- visible without dissection ; anterior tarsi with first joint more or less slightly thickened at tip beneath; anterior femora armed OL SUI AC ees Ae = re SS ee Oe Sere EN De eee. Discogenia. Middle lobe of the mentum small; anterior tarsi comparatively simple beneath, groove entire. . Lateral lobes of the mentum fully exposed; sculpturing com- paratively simple; femora mutie_____________1 Vetablapylis. Lateral lobes moderately exposed; species opaque to shining; elytra tuberculate; anterior femora not sinuate, mutic. Pseudeleodes. Anterior tarsi dissimilar in the sexes. Species pubescent throughout. Hairs long and flying; femora mutic; anterior tarsi with first joint scarcely produced ventro-apically, ventro-apical spinules noticeably produced in the female, not so in the male plantar Zroovesidistineta..— === ea Tricheleodes. Hairs short, yellowish and recumbent; form oval, opaque, Sul Ge WRES Se Cees eee ae ee ee Heteropromus, Species not usually pubescent, rarely so. Form elongate, usually large; first joint of the anterior tarsi slightly thickened at tip beneath, bearing a small trans- verse tuft of yellowish or brownish modified spinules which interrupt the groove in the male; simple in the female. Steneleodes. Form ovate or fusiform; the first one or two joints of an- terior tarsi in the male more or less thickened and some- times flattened beneath, densely clothed with fine pubescence ; tarsi simple and unmodified in the female; anterior femora ArMeds OLA MUbC2=—— sans s Ss Se es ee ee Promus. Form short ovate, moderate in size to small, robust (elon- gate and depressed in tibialis); anterior tarsi of the male with first two or three joints feebly thickened at tip be- neath and clothed with dense silken or brownish pubescent tufts, obliterating the groove; joints simple with grooves entire in female; femora mutic___________________ Blapylis. Form ovate, moderate in size, less robust; first joint of the anterior tarsi more or less thickened and slightly more prominent ventrally than the others, pubescent tuft variable, most evident in extricata; in the male first joint with a minute tuft of silken pubescence at tip beneath __Litheleodes. 36 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Subgenus MELANELEODES, new. Anterior spurs of the anterior tibie dissimiliar in the sexes, and more or less curved. Longer and slightly thicker than the posterior in the male; notably longer, much broader and stouter and most strongly curved in the female. z In the males of humeralis and rileyi the anterior spurs are much more strongly developed than in the same sex of the other species of the subgenus. The anterior femora are mutic and the first joint of the anterior tarsi is more or less thickened and feebly produced at tip beneath in both sexes. The male generally has a minute pencil of yellowish or brownish modified spinules on the produced tip; this is surrounded by a few closely placed, ordinary piceous spinules; in the female the produced tip bears only a few ordinary spinules. Subgeneric genital characters—M ale—— torte) Seis |e |'aisjers|| eisrts [fear ation | voters NE len stetesey| hak=rel sere aver fateea feces etenerel erate rece acer : | | var. anthracinad ....|....|----| om | eoratelteeoie || si eerie ISBScleosal anos sacdlecad sessseas sb.qelece sect tate ep eaae | | | | | Var. lsivans 22 sa\eat- Sai eal oleosole leer tepreelt eats shearers seeeleeee) sees] eee e|e eee lee eres A : | | | | | | CUNEAECONIS xe ace leeea |e see | See al eetes eee eaeel eee CT eS raiall avers Seiste|lewele|ioeie| Stee eis sic] sata sefee| Sere eres ; | | | humeralis 6......--.- tera seemlstos lerseelea Ee Scapa se | eee eel Iteesce [ocala ected | ene | ee Meron | | 2 | LOUIE ae se atatctare wiaretars |= == a|oaee | X |escs}-ee-|oeee|an=- |n2--|--2- [feted ter =ts event seen fe cheater eelae erevet fet ofe ey tal aioe Ree ies laral SS R UMCOSLALG sa s-seenes x XK je---] X J----] X |----]---- as sities Ho isratetelh) CMa II ete tmlell eveimtn || BOS en RO Be | PEMINOTAES rs <:-)2-)-)a'=,- | ance ceisler cel see |e eer | Sach eee eee y= /ao| earn [leis | rovers mallets NEOMERICANG ...---- io x ce See cera Sere joes | Se ee alficr eee iS aes | | | “ Quadricollis—Sonora, Mexico. In the Biologia Cent. America (Vol. IV, Pt. 1, p. 80) the author writes that he had seen specimens from Mexico. They probably belonged to a race or a closely related species. I have not seen an authentic example from Lower California. A specimen so given by Dr. George Horn—and examined by me—is probably a closely related species (Proc. Cal. Acad. Sciences, 2d ser., IV, Pt. 1, p. 350). The specimen was anatomically defective and has since been destroyed. » Humeralis.—Mexico ? In the Biologia (reference as above) the author writes: “There is an example of this species in Mr. F. Bates’s collection labeled as from Mexico ; possibly some mistake about the locality.” I agree with this. Dr. George Horn, in the Proc. Cal. Aead. Sciences, 2d ser., IV, Pt. 1, p. 350, gives this species as occurring at Sierra Laguna, San Francisquito, La Chuparosa, and San José del Cabo, Lower California. I have studied these specimens in the collection of the academy and find them not humeralis at all. (See tibialis.) Genealogy—In the accompanying diagram I have presented my present views regarding the descent and relationships of the species and races of the subgenus J/elaneleodes. From the subgeneric trunk divergence occurred along the lines of Groups I and II. In Group I the principal divergence is manifest in the calcarine development as set forth for the Carbonaria and Quadricollis sec- tions. In the Quadricollis section the calearine modification is most strongly perpetuated, the specific differentiation resulting in the mod- ification of surface sculpturing, cuneaticollis being intermediate be- tween guadricollis and humeralis, but retaining greater affinity with REVISION OF ELEODIINI BLAISDELL. 39 the former; antennal modification resulted in racial divergence in quadricollis, while greater calcarine development characterized the humeralis ramus. ii In the Carbonaria section crural development seems to have taken place, not affecting the main carbonaria stem where surface seulptur- ing has undergone modification along two lines, the smooth and the asperate as indicated in the synopsis of the species. Group I appears to be very plastic and reactive to modifying forces; Group IT less so. In Group I there is evidence of greater specific and racial differentiations. one Obsoleta hao Onmtissa. oo a Peninsularis ee fe Debilis Tustrans, Anthracing Quadricollis aN Cuneaticollis Be Humeralis Ampla eta Pedinoides ANG Quadricollis Section /_—— N°omeXicana Section on Oy D. olosa oD SN < Z cok Tricostata, Subgeneric Trunk Fic. 1.—GENEALOGICAL DIAGRAM OF THE SUBGENUS MELANELODES. The species of the present subgenus may be divided into two groups as follows: Elytra elongate oval, convex, comparatively narrow, feebly emarginate at base, humeral angles not rectangular; thorax not conspicuously broadened at base, angles obtuse, not overlapping humeral angles of elytra, more or less arcuately declivous at the sides, marginal bead rarely entirely visible when viewed TGC EN I iyamebors TIN END OC eat ree re ee ee a ee Group I. Elytra broad, flattened, truncate at base, humeral angles rectangular; thorax conspicuously broadened at base, angles rectangular, overlapping the humeral angles of the elytra; pronotum but slightly arcuately declivous at the sides, marginal bead distinctly visible from above._-—-—~~-____--~_---=-_ Group II. Group I may again be conveniently divided into two sections: Anterior spur of anterior tibiz: more or less gradually narrowed from base to ERG NEE ATTN ORME RECS ce Reka thas Lie Peg ee ie pel ete thnk Fe ey aE Carbonaria Section. Anterior spur of anterior tibiz: more or less decidedly thickened in both sexes; gradually narrowing from base to apex in the male; much thickened and sub- parallel, narrowed more or less gradually beyond the middle third or basal half, subacute or obtusely rounded when worn__--__--~ Quadricollis Section. 40 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. s) 1 6. 9 -~* GROUP I.—CARBONARIA SECTION, Legs moderate in length ______________ eee ee ee ee ei al begs lone; elytra rather, more pointed: behind? =e 6 Surface smooth; shining; punctures simples. 22232)" > 2 ee 7 Surface more strongly sculptured, punctures more or less muricate______ 5 Smaller: species... 2 221 ee ee ee eee ee 2 eareer ‘SPECIES’ 2-2 = oe a a ak eee nee eer 4 Elvtra with distinct and rather distant series of moderately sized punctures, with a single interstitial series of irregularly spaced, fine punctures (form SEMANA FOX CAEL COL) Sa eae ae ap a debilis. Elytra with distinct series of rather coarse punctures, punctuation never diffuse. Pronotum usually widest just before the middle (females at least). East of the. Colorado River ee eee eee carbonaria. Pronotum quite quadrate, sides evenly rounded, equally narrowed before and behind (form somewhat similar to a smooth, finely punctate Joryty... “Dexas ees Se. Se ee eee ee var. soror. Elytra usually more or less diffusely punctate, series frequently more or less evident, punctures generally subequal in size. West of the Colorado River. Larger size: Apical angles of thorax obtuse, apex feebly emarginate or truncate; elytra -widest= at: 1d dil esa sa ee eee ee omissa, Apical angles subacute, apex moderately emarginate; elytra usually widest behind the middle; surface polished___—__ var. peninsularis. Smaller size. Thorax more or less quadrate; males slender__var. pygmed. Elytra more or less subasperately sculptured; surface moderately shining, punctures irregularly, rather densely placed and submuricate, rarely sub- SUT ek A Si ea 2 es en ee ee SG knausii. Elytra muricately punctate; surface dull. father robust. Elytra striato-punctate; sometimes estriate and dif- fusely punctate; intervals more or less moderately convex____obsoleta. More elongate. Elytra sulcate, intervals strongly Convex___var. porcata. Surface more or less opaque througheut; pronotum broader ____-~-—__~ ampla, Surface more or less shining, pronotum quadrate and polished____var. dolosa, GROUP 1I.—QUADRICOLLIS SECTION, Anterior spur of the anterior tibise, moderately produced in both sexes__--1 Anterior spur of the anterior tibise, strongly produced in both sexes (less in the male) 2222 222 ee ee ee ee ae eee 2 Klytra striato-punctate to diffusely punctate. Antenne moderate in length; surface more or less shining; epistoma coarsely and. contiuently punctates 2 22e ee eee quadricollis, Antenne long. Epistoma more finely and not confluently punctate. Marcer- species: surtace: dull= =. 2- see var. anthracina. Smaller species; surface polished and shining ___—___ var. lustrans, Elytra with punctures irregularly distributed, without serial arrangement, but producing a strongly rugulose appearance by mutual semicoales- cenee! 22-222 2 = ee oo Se es Se eS CUneaticollis: Elongate species___---~~ eae oe 3 Shorter; more? robust species =). = 23 =. 2 ee eee ee eee 4 REVISION OF ELEODIINI—BLAISDELL. 41 3. Elytra opaque, densely muricately punctate (Washington); frequently granulato-muricately (Nevada and California) or tuberculo-muricately (Colored 0) Ap UN CLUE Cees tS Be See ee es ee humeralis. 4, Elytra shining, subasparately punctate, punctures rather sparse, often with M@oteehlehlinesimarranecemen t=. ieee eee ee es rileyi. Virals COStAtCl= = es ee eee Pee Pe a ee elect Eh A aA Se eer 2 1 Hilyiraenoteinechemicas tecostates- se ae reer ee OR ee eee 1. Elytra with alternate interspaces more elevated, at times all quite evenly SOP Se SULLA Ce yO DA QM C2 eae ee ees et a ek ee ae ee ee tricostata. 2. Elytra glabrous and shining, with regular strize of moderately large punctures, interstices equally and very feebly convex ; pronotum with basal GEDEES SONS Rae eens os eee wet ee aes Merry mney nantes ke pedinoides. Elytra opaque, diffusely punctate; basal depressions of pronotum obso- LOU Giese stk bt Bir. Tee ee. ee Se airk abies PW Peele ye var. neomexicana. GROUP I.—CARBONARIA SECTION. ELEODES DEBILIS LeConte. Eleodes debilis—LEContTrE, Proc. Nat. Sci. Acad. Phila., 1858, p. 185.— Horn, Trans. Amer. Philos. Soc., XIV, 1870, p. 308.—Casry, Annals N. Y. Acad. Sci., V, Nov., 1890, p. 396. Ovate, moderately robust, glabrous and shining. Head twice as wide as long, feebly convex, rather coarsely punctate, punctures very sparse on the vertex, denser on the epistoma; surface more or less impressed laterally and along the frontal suture. An- tenne moderate, reaching to the base of the prothorax, subequal in the sexes, outer four joints very feebly compressed, scarcely dilated, third joint about equal to the next two taken together, fourth hardly longer than the fifth, the latter, sixth and seventh subequal, eighth feebly triangular, ninth and tenth suborbicular, scarcely transverse, eleventh ovate. Pronotum subquadrate, widest at the middle, about one-fifth to one-third wider than long; disc rather strongly and evenly convex from side to side, feebly so antero-posteriorly and more or less de- clivous at the apical angles, finely and sparsely punctulate, rather more strongly so laterally; apex teebly and evenly emarginate, usu- ally obsoletely margined; sides converging a little anteriorly, rather evenly and moderately arcuate in apical two-thirds, thence to base obliquely straight or feebly sinuate, finely margined; base rather broadly rounded and finely margined, about a third wider than the apex; apical angles subacute; basal angles obtuse. Propleure smooth, sparsely and more or less obsoletely punctulate, slightly rugulose. Elytra oval, widest at the middle, about a third longer than wide; base broadly and very feebly emarginate, scarcely wider than the AQ BULLETIN 63, UNITED STATES NATIONAL MUSEUM. contiguous base of the prothorax; hwmeri more or less acute; sides evenly arcuate, apex obtusely and narrowly rounded; disc evenly and moderately convex, laterally more strongly rounded, evenly and quite strongly arcuately declivous posteriorly, punctate, punctures shghtly separated, moderate sized and arranged in rather distant series; intervals flat with a single series of irregularly spaced and usually fine punctures, both series becoming more or less confused at the sides and apex. Epipleure moderate in width, superior margin feebly sinuate be- neath the humeri, very gradually narrowing to apex; surface gla- brous and more or less finely and sparsely punctulate. Sterna more or less glabrous, irregularly and more or less densely punctate, rugulose. Parapleure rather coarsely and somewhat densely punctate. Abdomen shining and glabrous, sparsely, and finely punctulate, more or less rugulose. Legs moderate. Anterior femora mutic in the sexes; anterior tibial spurs slightly dissimilar; first jomt of the anterior tarsi produced at tip beneath and feebly dissimilar in the sexes. Male.—Slightly robust. Thorax quite quadrate; abdomen sub- horizontal, first two segments more or less broadly impressed at middle, intercoxal process slightly concave; anterior tibial spurs comparatively small and slender, the anterior one scarcely widened. First joint of the anterior tarsi with a minute pencil of brownish modified spinules surrounded by dark spinules on produced tip be- neath. Female.—Moderately robust.. Thorax slightly transverse. Abdo- men horizontal and rather strongly convex. Anterior tibial spurs a little larger and longer, the anterior spur shghtly widened but grad- ually narrowed from base to apex. First joint of the anterior tarsi with dark spimules on the produced tip beneath. Measurements.—M ales: Length, 12-13 mm.; width, 5.5 mm. Fe- males: Length, 12.8-14.8 mm.: width, 6—6.5 mm. Genital characters, male. Edeagophore elongate oval, acutely pointed at the apical extremity and shghtly arched. Basale oblong, evenly convex, sides shghtly arcuate. Apicale triangular, rather evenly convex above, with a narrow median membranous groove at the middle third; sides feebly arcuate at base, thence broadly and more or less strongly sinuate to apex, the latter appearing attenuated; base strongly and moderately narrowly arcuate at middle, laterally broadly sinuate. Sternite rather short and transversely parabolic. Each lobe with the external border evenly arcuate to apex, the latter narrowly rounded; internal border oblique; surface feebly convex, shining, REVISION OF ELEODIINI—BLAISDELL. 43 very sparsely punctate and setose apically, setae moderately short. Sinus triangular, membrane slightly setose at bottom. Female.—Genital segment quadrate, valves reflexed at the sides, setose, glabrous and shining. Valvula (Plate 4, fig. 26).—Dorsal plate oblong, explanate exter- nally and slightly deflexed, internally reflexed and impunctate in apical half; surface convex externally, very sparsely punctate, punc- tures denser on convexity, each with a long seta; external margin straight ; angle rectangular and prominent, sometimes feebly rounded, with a few moderately long setxe; apical margin squarely truncate to moderately and arcuately produced in outer half, more or less sin- uate over the appendage; internal margin more or less arcuate. Apex short, triangular and acute, set with a few moderately long sete. Appendage (cercopod) short and moderate in size, conical or feebly depressed, bearing moderately long sete. Basal promimences not evident. Superior pudendal membrane feebly and distinctly longitudinally rugulose, reaching to about the middle of the dorsal plate. Ventrolateral surface moderately convex, scarcely concave before the apex; submarginal groove broad and shallow beneath the ex- planate external border of dorsal plate; surface finely punctate and setose, setze minute. The internal borders of the valvule are con- tiguous in basal sixth; genital fissure fusiform and moderately wide, with the inferior pudendal membrane visible in basal half. Habitat—New Mexico (Santa Fé, C. V. Riley) ; Arizona (Sulphur Spring Valley, Hubbard and Schwarz); Texas (Alpine, elevation 4,400-6,000 feet, H. F. Wickham). Number of specimens studied, 3 males, 6 females. Type in the LeConte collection. Type-locality—New Mexico; collected by Mr. Fendler. Salient type-characters——Thorax subquadrate, slightly narrowed anteriorly; sides broadly rounded; apical angles acute, the basal ob- tuse; base broadly rounded. Elytra oval and convex, strongly de- clivous posteriorly, not acuminate at apex; base broadly emarginate; humeri acute; disc quite strongly striato-punctate, intervals with a single series of fine punctules (LeConte). Diagnostic characters——In size and shape debilis resembles a smooth extricata, from which it differs in having the anterior femorz unarmed in both sexes, and by the distinct and rather distant series of moderate punctures, with a single interstitial series of irregularly spaced fine punctures. The punctuation in debilis somewhat resembles that observed in carbonaria, from which it can be separated by its smaller size and female genital characters; the latter also separates it from all other members of the subgenus. 44 BULLETIN 638, UNITED STATES NATIONAL MUSEUM. A specimen before me has been compared with the LeConte type and pronounced typical by Mr. F. Blanchard. | This species has been considered as identical with carbonaria, and I agree with Col. Thomas Casey “ in considering it a distinct species. The genital characters are somewhat aberrant to the present sub- genus, but on account of the greater development of the external lobe of the valvular apex, I have placed it here; the apical margin of the dorsal plate is quite squarely truncate and the appendage more strongly exposed. In all specimens examined the female genital characters were constant. . On account of the homogeneous structure exhibited by the small series before me, I am unable to indicate with any feeling of certainty, its relationships and descent. A large series illustrating the lines of variation is necessary for this purpose. I might tentatively suggest that debidlis may tend toward the subgenus Promus. I would about as soon accept this postulate as to consider it identical with car- bonaria. The mentum? is moderate in size, and more or less triangulo- trapezoidal in outline; surface rather coarsely and somewhat densely punctate, laterally with rather broad, shallow, and subfoveate im- pressions, with a median longitudinal subcariniform convexity ; each puncture bears a short inconspicuous seta. The prosternum between the coxe is rather prominent ventrally. convex to nearly horizontal and mucronate behind. The mucro is variable in size, sometimes pointed and at others broad and obtusely rounded. The mesosternum is more or less arcuately oblique and concave. The intercoxal process of the first ° abdominal segment is shghtly transverse and equal in length to the post-coxal portion. The meta- sternal salient is about two-sevenths narrower than the abdominal process. In the male the second abdominal segment is equal in length to the post-coxal part of the first and about a third of its length longer than the third segment, the latter being about a fourth of its own length longer than the fourth segment. In the female the second segment is noticeably longer, being about a fourth of its own length longer than the post-coxal part of the first. The fourth is about half as long as the second, while the third is about a fourth of its own length longer than the fourth. The profemora are somewhat tumid externally, the tibial grooves are well developed, concave, and glabrous, margins subcariniform, converging and meeting at basal third. “VII. Coleopterological Notices, II, Oct. 6, 1890, p. 396. >The mentum refers to the middle lobe unless otherwise stated. © In all descriptive remarks only the visible segments are considered. REVISION OF ELEODIINI—BLAISDELL. 45 The meso- and metafemora are each subequal in width, the former is scarcely compressed, the latter distinctly so. The tibial grooves have their margins feebly subcariniform and gradually converging to become contiguous and evanescent at basal thirds. The protibiz are feebly compressed ; all the tibiz are without tarsal grooves, and muricately sculptured. The tarsi are moderate in length and comparatively slender. The protarsi are about a fourth of their length shorter than a mesotarsus; first four joints subequal in width; the second, third, and fourth are about as long as wide, the fourth shghtly smaller than the third; the fifth is about as long as the preceding three taken together; the first is longer than wide, and rapidly narrowing to the base. The mesotarsi are about a fifth of their length shorter than a metatarsus. First four joints are subequal in width, gradually and shightly diminishing in length from the second to the fourth. Joints one and five are each separately subequal to the combined lengths of the second and third. The metatarsi are about two-fifths of their length shorter than their metatibie. The fourth joint is about equal to the combined lengths of the second and third; the first is a little longer than the same, and the second is just a little longer than the third. The comparative tarsal measurements were made on the female. ELEODES CARBONARIA (Say). Blaps carbonaria Say, Journ. Acad. Nat. Sci. Phila., IIT, 1828, p. 260. Eleodes carbonaria LECoNntTE, Complete Writings of Thomas Say, II, 1859, p. 125; Proc. Acad. Nat. Sci. Phila., 1851, p. 181.—Horn, Trans. Amer. Phil. Soc, XIV, 1870, p. 308.—CHAMPION, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 8308.—Casry, Annals N. Y. Acad. Sci.. V, Nov., 1890, p. 395. Hleodes soror LECoNTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 185. Eleodes immunis LECon?TrR, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.— Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 308. Oblong-ovate, moderately elongate, smooth, more or less shining, estriate, elytral punctures arranged in series. Head twice as wide as long, more or less convex, rather densely and irregularly punctate, punctures denser anteriorly, sparser about vertex, surface more or less impressed along the frontal suture and laterally. Antenne moderately long, reaching to or very slightly beyond the base of the prothorax, subequal in the sexes, outer four joints feebly compressed, just perceptibly dilated, third joint about equal to the next two taken together, fourth just noticeably longer than the fifth, the latter, sixth and seventh subequal, eighth triangular and as wide as long; ninth and tenth more or less feebly transversely oval; eleventh ovate. Pronotum subquadrate, widest at or a little in advance of the middle, and about one-fifth to one-third wider than long; désc evenly 46 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. and moderately convex, more or less arcuately declivous laterally and at the apical angles, finely and sparsely punctulate, punctures slightly denser and coarser along the margins laterally; apex slightly emargi- nate or truncate, finely or more or less obsoletely margined; sides evenly and more or less moderately arcuate from base to apex, or more strongly arcuate in the anterior two-thirds, and thence to base oblique and convergent, or very feebly arcuate, sometimes obsoletely sinuate, finely beaded; base feebly arcuate, finely to more or less obsoletely margined, and one-fifth to one-fourth wider than the apex; apical angles distinct, scarcely at all rounded; basal angles obtuse. Propleurw finely and very sparsely submuricately punctate and more or less distinctly rugulose. Elytra oval, widest at the middle, less than twice as long as wide; base slightly emarginate, more or less feebly sinuate laterally, a little wider than or equal to the contiguous base of the prothorax; humeri subacute or feebly obtuse; sides evenly arcuate, apex obtuse; disc moderately convex on the dorsum, at times slightly depressed, later- ally quite strongly and evenly rounded, evenly and arcuately decli- vous posteriorly, serially punctate, strial punctures usually rather large and somewhat deeply impressed, at times rather small and not impressed, the individual punctures are usually more distant from each other than their own diameters; intervals flat, frequently feebly convex, with a single series of very small, distantly and irregularly spaced punctures; laterally the interstitial punctures. become larger, and both series more or less irregular and somewhat confused. Epipleure gradually narrowing from base to apex, superior margin feebly sinuate beneath the humeri; surface finely to obsoletely or dis- tinctly and sparsely punctate. Sterna shining to subopaque, finely and rather densely punctate, at times rugulose. Parapleure rather coarsely and quite densely punctate. Abdomen more or less glabrous, finely and sparsely punctulate, first segment more strongly sculptured; surface obsoletely to dis- tinctly rugulose. Legs moderate. Anterior femora mutic in the sexes; anterior tibial spurs and tarsi slightly dissimilar in the sexes. Male.—Body comparatively slender. Elytra rather gradually narrowed and evenly arcuato-obliquely declivous posteriorly. Ab- domen slightly oblique, very moderately convex, first two segments more or less impressed at the middle. Anterior spurs of the anterior tibie shghtly the stouter, almost twice as long as the posterior and gradually narrowing from base to apex, both comparatively slender and acute. Tarsi distinctly longer than in the female, the anterior with first joint bearing a minute pencil of brownish modified spinules, surrounded by ordinary piceous spinules on the thickened tip beneath. REVISION OF ELEODIINI—BLAISDELL. 47 Female.—Body quite robust. Elytra rather broadly oval, rounded and more or less arcuately and vertically declivous posteriorly. Ab- domen horizontal, evenly and rather strongly convex. Anterior spurs of the anterior tibiz distinctly the stouter, more gradually narrowing from base to apex and about a third longer than the posterior. First joint of the anterior tarsi transversely clothed with ordinary spinules on thickened tip beneath. The material before me and referred to the present species is quite variable and presents a number of incipient races, of which three may be recognized to aid in the placing of specimens: Forma glabra.—Smooth and shining. Elytral punctures fine and not impressed, with a tendency to some irregularity on the disc, in- tervals flat. Forma typica—FElytra with rows of rather large, impressed and distinctly defined punctures, intervals flat. Forma interstitialis——Elytra with very strongly impressed punc- tures, intervals feebly convex. The above forms or incipient races are connected by an abundance of mesotypes and when properly arranged make an instructive morphological series. Measurements.—Males: Length, 15.8-18 mm.; width, 5.5-6.5 mm. Femates: Length, 17-20.5 mm.; width, 7.8-9 mm. Genital characters, male-—Kdeagophore rather elongate, fusiform, somewhat strongly arched and well chitinized. Basale oblong, rather strongly convex and more or less gibbous basally; sides evenly and rather moderately arcuate. Apicale triangular and feebly elongate; surface evenly convex, with a median longitudinal groove extending from near the apex to near the base, membranous in apical half, linear towards base; s?des broadly and very feebly sinuate at middle third, apex subacute: base broadly and arcuately lobed at middle, very feebly sinuate laterally. Sternite parabolic in outline and slightly transverse. Each lobe with the external border more or less evenly arcuate, apex more or less evenly rounded, angle feebly evident at times; internal border short; surface rather evenly convex and glabrous, quite coarsely and densely punctate apically, more sparsely so towards base, setose, setae moderate in length, longer about apex, and not very dense. Sinus short, with the membrane very sparsely setose at bottom. Female.—Genital segment quadrato-trapezoidal, somewhat de- pressed and setose. Valvula (Plate 3, fig. 16).—Dorsal plate oblong, sometimes slightly widened at base, frequently deflexed apically, gradually explanate externally from basal third to apical margin with which it is directly continuous; surface plane, finely to rather coarsely and sparsely punctate, setose, setee rather short; external border feebly sinuous to 48 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. scarcely arcuate; apical margin not defined from the external apical lobe. Apex with the outer lobe well developed, the latter three-fourths as wide as the dorsal plate, rather broadly rounded and slightly sem1- membranous at tip; internal lobe small and membranous; both lobes set with long flying sete. Appendage short and more or less conical, sometimes slightly visible from above, directed backwards and shghtly inwards. Fossa moderate and fringed with moderately long hairs. basal prominences moderate. Superior pudendal membrane longitudinally rugulose and reach- ing to the side of the internal lobe of the apex. Ventrolateral surface rather short, not strongly convex, but more or less gradually sloping to the apex, not usually transversely impressed but sometimes more or less so just behind the narrow transverse basal convexity; submarginal groove well defined beneath the explanate external border of the dorsal plate, not markedly bending inwards at apex to fossa; surfaces finely and sparsely punctate, setose, sete very short, at apex short and flying. Internal margins of the val- vule contiguous for a very short distance both at base and apex; genital fissure broadly fusiform, with basal half closed by the inferior pudendal membrane. Remarks on female genital characters—Forma glabra has the genital segment quadrate in outline and more thickly setose at apex, with the basal convexities more strongly developed. Forma intersti- tialis has the dorsal plate more strongly and coarsely punctate. Habitat—F¥orma typica.—Texas (Alpine, June, elevation 1,400- 6,000 feet, H. F. Wickham; Uvalde, elevation 930 feet; Granjino, May; Marfa, elevation 4,600-4,800 feet, July, Wickham); New Mex- ico (Albuquerque, March, Wickham; Santa Fe Canyon, August, elevation, 7,000 feet, F. H. Snow); Arizona (Fort Grant; Oak Creek Canyon, Snow) ; Colorado (La Junta, Garland, Pleasant Val- ley, on the Arkansas River, Snow; Wickham’s List). Forma glabra.—Texas; New Mexico (Highrolls, May and August; Cloudcroft, June, Warren Knaus); Colorado (Garland, June; Salida); Arizona (Chiricahua Mountains, May; Sulphur Springs Valley, June). Forma interstitialis.—Texas (Kl Paso, July, elevation 3,700-3,800 feet, Wickham) ; New Mexico (Deming, July, Hubbard and Schwarz; Luna; Albuquerque, March); Arizona (San Simon, July; Sulphur Spring Valley, May; Tucson, April; Chiricahua Mountains, May, Hubbard and Schwarz). Number of specimens studied, 100. Type destroyed. ee REVISION OF ELEODIINI—BLAISDELL. 49 Type-locality as given by Thomas Say: “ Region of the Arkansas River and near the Rocky Mountains.” Salient ty pe-characters.—Thorax subquadrate; sides regularly arcu- ate; apex very slightly emarginate; base nearly rectilinear, a little arcuate each side near the angles, the latter obtuse. Elytra without impressed strive; six regular series of punctures, which are more dis- tant from each other than the length of their diameters; interstitial | lines with a few remote, acute, minute punctures; sides rounded and irregularly punctured (Say). Diagnostic characters —Easily recognized from the other members of the subgenus by the smooth polished surface, the elytra being sculptured with rather distantly placed series of coarse punctures; the intervals are flat, with a single series of small distantly spaced punctures. At times the punctures of both series may be equal in size, and the species is then with some difficulty separated from omssa (see p. 72 and forma glabra) ; typically the intervals are flat, but often become feebly convex and approach porcata (see p. 63 and forma interstitialis). The extreme heterotypical specimens are apt to have a subasperate punctuation. Immunis LeConte appears to be a true synonym, probably more closely related to var. soror. (See p. 50.) The mentum is moderate in size, triangulo-trapezoidal to parabolo- triangular in outline; surface rather broadly foveate laterally with a median longitudinal ridge, sometimes feebly sculptured, more or less strongly punctate and not noticeably bearded. The prosternum is variable; frequently continuously rounded be- tween the coxe and behind, but not with the precoxal portion, not mucronate; often almost horizontally produced, at other times convex between the coxee and with a more or less strongly developed mucro behind. The mesosternum at times is quite vertically declivous, at others oblique and more or less feebly concave. The metasternal process is as wide as the abdominal salient is long. The abdominal intercoxal process (male) is shghtly transverse, a little wider than the metasternal salient, and equal to the post- coxal portion; the latter is also equal in length to the second seg- ment; the third segment is one-third of its length longer than the fourth. In the female the abdominal process is quadrate, and about a fifth of its width wider than the metasternal salient, and equal in length to the second segment; the latter is about twice as long as the fourth; the third segment is equal in length to the post-coxal part of the first. 59780—Bull. 63—09——4 50 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The profemora (male) are clavate, gradually tumid externally and subeylindrical in section; less so in the female and frequently somewhat compressed. The tibial grooves are more or less well defined by subcariniform margins which become contiguous and evanescent at basal fourth; the floors of the grooves are usually con- cave, glabrous or sometimes minutely subasperate. In both sexes the mesofemora are subtumid at middle, with the grooves rather narrow and limited by subasperulate margins, which are more or less evanescent before becoming contiguous at basal third. The sexes have the metafemora usually just noticeably widening from base to apex; grooves as on the mesofemora. The protibie are frequently subarcuate, feebly compressed. All the tibiae are without tarsal grooves and more or less muricate; the articular cavities are closed. The tarsi are moderate in length and stoutness. The protarsi are about a half of their length shorter than a mesotarsus. Joints two, three, and four are subequal, and combined subequal to the fifth; the first is longer than wide. . The mesotarsi are subequal to (male) or a seventh of their length shorter than (female) a metatarsus. Joints one and five subequal in length, either one is subequal to the combined lengths of the third and fourth; jomts two to four subequal in width and length, or just feebly diminishing in length in the order named. The metatarsi are about a third of their length shorter than a meta- tibia. The third joint is slightly shorter than the second, both com- bined about equal in length to the fourth; first jomt equal to the combined lengths of joints two, three, and four of a mesotarsus. ELEODES CARBONARIA var. SOROR LeConte. Form as in carbonaria, smooth and more or less feebly shining, sides of the pronotum apparently evenly arcuate; elytral punctures usually moderately small. [Head twice as wide as long, more or less moderately convex, some- what coarsely, irregularly, and sparsely punctate, punctures shehtly denser on the epistoma; surface usually impressed laterally. An- tenne moderate in length, reaching to the base of the prothorax, sub- equal in the sexes, outer three joints scarcely compressed or dilated, third joint about equal to the next two taken together, the fourth very slightly longer than the fifth, the latter, sixth, and seventh sub- equal, eighth subtriangular, and slightly shorter than the seventh, ninth and tenth suborbicular, eleventh subovate. Pronotum quadrate, widest at the middle, where it is one-eighth to one-sixth wider than long, slightly narrowed before and behind; disc moderately convex, quite strongly and arcuately declivous along ed REVISION OF ELEODIINI—BLAISDELL. 51 the lateral margins, finely and sparsely punctulate; apex subtruncate and more or less obsoletely margined; sides evenly, broadly and moderately arcuate from base to apex when viewed vertically from above, more strongly so in the anterior three-fourths when viewed obliquely from the side and obsoletely sinuate before the basal angles, margin finely beaded; base broadly arcuate, quite finely margined, and about one-fourth to one-third wider than the apex; apical angles more or less subacute; basal angles obtuse. Propleure smooth, finely and sparsely punctulate, more or less sparsely rugulose. Elytra oval, widest at the middle, rather strongly emarginate at the base and equal to or a little wider than the contiguous base of the prothorax; humeri acute and slightly prominent anteriorly ; sides evenly arcuate, apex obtuse and rather narrowly rounded; disc mod- erately convex on the dorsum, strongly and evenly rounded laterally, evenly and rather strongly arcuately declivous posteriorly, punctate, punctures moderate in size and arranged in moderately distant, scarcely impressed series; intervals more or less irregularly and sparsely punctulate, especially along the suture, laterally the series usually remain distinct. E'pipleure rather wider than in carbonaria, and gradually narrow- ing from base to apex; superior margin feebly and broadly sinuate beneath the humeri; surface smooth, sparsely and irregularly punc- tulate. Sterna shining, irregularly punctate and more or less rugulose. Parapleure irregularly and more or less closely punctate. Abdomen shining, evenly and sparsely punctate, usually obsoletely rugulose; intercoxal process more strongly sculptured. Legs moderate as in carbonaria. Anterior femora mutic in the sexes; anterior tibial spurs comparatively small and slightly dissimi- lar in the sexes, the anterior spur but slightly longer than the pos- terior. Anterior tarsi with the first joint feebly thickened at tip beneath and shghtly dissimilar in the sexes. Male.—Body moderately slender, elongate-subovate. Elytra grad- ually narrowing, arcuately and somewhat obliquely declivous pos- teriorly. Abdomen slightly oblique, moderately convex, more or less broadly impressed at middle of the first two segments, intercoxal process distinctly concave. Anterior spur of the anterior tibia grad- ually narrowing from base to apex. First joint of the anterior tarsi with a small pencil of modified spinules surrounded by ordinary spinules on the thickened tip beneath. : Female.—Body moderately robust. Elytra rather broadly oval, arcuately and somewhat vertically declivous posteriorly. Abdomen horizontal and somewhat strongly convex. Anterior spur of the anterior tibiz slightly thickened, acute, and gradually narrowing . 52 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. from base to apex. First joint of the anterior tarsi set with ordinary spinules on tip beneath. Measurements—Males: Length, 16.5 mm.; width, 6.5 mm. Females: Length, 18.3-19 mm.; width, 8-8.5 mm. Genital characters.—Male characters as in carbonaria. Female.—As in carbonaria, except that the dorsal plate of each ralve is narrower, with the sides more parallel and less explanate externally; the outer lobe of the apex is less broadly rounded. [Tabitat—Texas (Eagle Pass and San Antonio, LeConte; Browns- ville, Wickham. ) Number of specimens studied, 8. Type, a female in the LeConte collection. Ty pe-locality.—Kagle Pass; Texas. Salient type-characters.—Thorax quadrate, slightly narrowed an- teriorly and posteriorly ; sides broadly rounded; anterior angles acute, the posterior obtuse. Elytra strongly striato-punctate, intervals sparsely punctate (LeConte). Diagnostic characters.—On aceggnt of the form of the prothorax it somewhat resembles a smooth goryi, and LeConte differentiated it from that species as follows: “ The thorax is more quadrate and not more narrowed towards the apex than towards the base; the elytra are more deeply emarginate at base, and transversely much less convex, and those of the female are much less dilated; the punc- tures are more regular and never have the appearance of fovee.” All the specimens that I have received have either been labeled gory? ov wudentified, and it is undoubtedly confused with that spe- cies in collections generally. In all cases of doubt the subgeneric and genital characters must be carefully studied. The scuplturing 6f soror is quite like that of a typical carbonaria, except that the serial punctures on the dorsum of the elytra are smaller, although as large or slightly larger and more distinctly de- fined laterally, the surface is feebly duller, the pronotum more quad- rate on account of the strongly deflexed sides of the dise and conse- quently less strongly rounded when viewed from above, and usually not more narrowed toward the apex than the base, but this character may be variable. The actual form of the pronotal margin in both sexes is like that of a male carbonaria, widest at the middle and not noticeably more strongly arcuate in the anterior moiety, as in the female of car- bonaria. From the members of the guadricollis section of the present sub- genus it 1s to be known by the form of the anterior spurs of the anterior tibie. I am indebted to Professor Wickham for the specimens in my collection, REVISION OF ELEODIINI—BLAISDELL. 53 LeConte separated the species described by him as ¢mmunis from soror as follows: “ Of the same form as sovor, but the thorax is more narrowed behind, almost as in #’. guadricollis; the punctures of the rows of the elytra are sometimes placed in distinct grooves, while in other specimens the surface is even.” (See carbonaria.) Mr. Blanchard has pronounced my specimens undoubtedly the same as LeConte’s type from Eagle Pass, Texas. He also writes that the type and a male from “ Texas” have the elytra and under side shining, and two other females from “ Texas” are dull. The mental, sternal, abdominal, and crural characters are as in carbonaria, except that the meso- and metafemora appear to be slightly more slender. The protarsi appear to be subequal in length in the sexes, and the mesotarsi appear relatively slightly shorter. ELEODES AMPLA, new species. Subfusiform to fusiform-ovate, subopaque, and smooth. Head twice as wide as long, more or less moderately convex between the eyes, rather broadly flattened anteriorly, usually feebly impressed laterally, sometimes across base of the epistoma; frontal suture dis- tinct and frequently bisinuate; evenly and sparsely punctate, pune- tures coarser on the epistoma and finer on the vertex. Antenne reaching a little beyond the base of the prothorax, very feebly com- pressed in outer four joints, scarcely dilated, third joint scarcely as long as the next two combined; the fourth, fifth, sixth, and seventh subequal in length; eighth triangular and as long as wide; ninth suborbicular; tenth slightly transversely oval; eleventh oval to ovate, and generally distinctly obliquely truncate at tip. Pronotum widest at or a little in advance of the middle, about one- fourth wider than long; moderately and evenly convex from side to side, feebly so antero-posteriorly, more or less arcuately declivous laterally along the sides; finely and sparsely or almost obsoletely punctulate; apex slightly and evenly emarginate, more or less obso- letely beaded; sides almost evenly arcuate from base to apex when viewed vertically from above, more strongly so in the anterior three- fourths, and obsoletely sinuate at basal fourth when viewed obliquely from the side, finely beaded ; base quite broadly rounded and more or less feebly sinuate at middle, finely beaded, and about a fourth to a third wider than the apex; apical angles subacute; basal angles obtuse. Propleurw rather smooth, finely and sparsely punctulate, usually distinctly rugulose. Elytra oval, less than one-half to about a third longer than wide; base more or less emarginate, frequently sinuate laterally, scarcely to a little wider than the contiguous prothoracic base; huwmeri subacute, scarcely prominent anteriorly; sides evenly arcuate, apex obtuse and 54 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. eather narrowly rounded; disc more or less moderately convex, some- times slightly flattened on the dorsum, more strongly and rather evenly rounded laterally, arcuately declivous posteriorly, punctate; the strial punctures are rather small to moderate in size and some- what closely placed in moderately distant series, usually not at all, but sometimes moderately impressed; intervals flat, rarely slightly convex, and with a single series of rather widely spaced, very fine punctures which at times become slightly irregular, especially later- ally and about apex. Epipleurw woderate in width, gradually narrowing from base to apex, Superior margin very broadly and slightly sinuate beneath the humeri; surface smooth, obsoletely to finely and sparsely punctulate. Sterna more or less shining, strongly punctate, and rugulose. Parapleure rather densely, finely, and subscabrously punctate. Abdomen usually shining, sparsely and quite regularly punctulate, more or less rugulose; intercoxal process and fifth segment more strongly sculptured. Legs long, comparatively moderately stout. Anterior femora mutic and the anterior tibial spurs moderately long, slightly dissimilar in the sexes. Anterior tarsi with the first joint feebly thickened at tip beneath. MJale—Body moderately slender and more or less fusiform, an- tenn reaching a short distance beyond the base of the prothorax. Elytra quite gradually narrowed in posterior fourth, arcuately and obliquely declivous. Abdomen distinctly oblique, moderately convex, first two segments more or less impressed at middle, process concave. Anterior spur of anterior tibize about a third longer and a little stouter than the posterior, and narrowing moderately from base to apex. First joint of the anterior tarsi with the minute pencil of modified spinules on tip beneath scarcely evident and surrounded by the ordinary spinules. ; Female.—Robust. Antenne reaching just beyond the base of the prothorax. Elytra broadly oval, somewhat gradually narrowed and arcuately declivous posteriorly. Abdomen horizontal and rather strongly convex. Anterior spurs of the anterior tibize about a half longer than the posterior and feebly broadened, with sides rather slowly but evenly narrowing to apex, the latter acute. Anterior tarsi with ordinary spinules on the thickened tip beneath, the latter trans- verse and narrow. Measurements —M ales: Length, 19-23 mm.; width, 6.5-8.5 mm. Females: Length, 23-25 mm.; width, 6.2-9.5 mm. Edeagophore elongate, flaxseed-shaped (flattened, oblong-ovate) and more or less arched. Basale oblong; surface strongly convex towards base; sides feebly arcuate. Genital characters, male. REVISION OF ELEODIINI—BLAISDELL. 55 Apicale triangular and slightly longer than wide; surface evenly convex, with a narrow median groove in apical half; sides broadly sinuate at the middle, apex gradually narrowed and subacute; base broadly lobed and feebly sinuate laterally. Sternite transversely parabolic in outline. Each lobe subtriangular with the external border evenly and broadly arcuate, apex rounded but not narrowly; internal border rather short; surface evenly con- vex, glabrous at basal third, sparsely punctate and setose apically, setee moderate but longer at apical border. Membrane sparsely setose across the bottom of the sénus. Female.—Genital segment subquadrate, sometimes shghtly deflexed at apex, setose. Valvula (Plate 4, fig. 27).—Dorsal plate oblong, moderately ex- planate externally; sides subparallel, and shghtly sinous; surface nearly plane, slightly longitudinally concave, glabrous and im- punctate in basal third, elsewhere coarsely, sparsely, and irregularly punctate, setose, sete rather long and reclining; apical margin on external lobe strongly arcuate in outer three-fourths, internally sinuate. Both apical lobes set with long hairs; external lobe large, the internal very small. Appendage submammilliform and scarcely projecting beyond the fossa, hardly visible from above. Fossa rather large. Basal prominences scarcely evident. Superior pudendal membrane longitudinally rugulose and reach- ing about to the base of the internal lobe. Ventrolateral surface rather strongly convex at base, scarcely to feebly transversely concave before the apex; submarginal groove broad extending beneath the apex to the margin of the fossa. In- ternal margins of the valves contiguous at basal eighth; fissure broadly fusiform and closed in basal half by the inferior pudendal membrane. Habitat—Arizona (Oracle, July 15, Hubbard and Schwarz; Pinal Mountains, collection, University of Nebraska; Santa Rita Mountains, elevation 5,000-8,000 feet, July, F. H. Snow); New Mexico (Santa Fé Canyon, August, elevation 7,000 feet, F. H. Snow) ; Texas (Brownsville, June, F. H. Snow). Number of specimens studied, 13. Type in my own collection. Ty pe-locality—P inal Mountains, Arizona. Salient type-characters.—Subopaque. Prothorax widest at (male) or in advance (female) of the middle. Elytra noticeably narrowing at apical fourth in both sexes; striato-punctate, striae not impressed, strial punctures rather small, the series rather distant, interstitial punctures very fine. Legs long. 56 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Diagnostic characters.—In collections the females of the present species are usually associated with swbnitens, while the males are placed with guadricollis or carbonaria. On account of its large size and dull integuments ampla has to be carefully differentiated from swvbnitens, from which it differs in the sides of the pronotum, being more strongly rounded and widest at or in advance of the middle, by the less prominent apical angles, and the subgeneric characters. Ampla bears no resemblance to a typical carbonaria,; the dull luster, fine and unimpressed elytral punctuation readily separates it from the latter species, and besides the males are quite fusiform in outline and the legs are distinctly longer, characters never observed in car- bonaria. A male from Santa Fé canyon, New Mexico, has the strial punctures of the elytra moderately large and slightly impressed, but the form is elongate and fusiform. From quadricollis it differs by the anterior spurs of the anterior tibie of the females, being much narrower and evidently narrowed from base to apex, and by the long legs. Specimens from about Brownsville, Texas, are more shining than those observed from elsewhere. For characters separating it from its race dolosa, see below. In fact, ampla is the largest species in the present section of the subgenus. The mentum is very moderate for so large a species and varies quite a little in form—triangulo-trapezoidal to trapezoido-parabolic; the surface is moderately strongly punctate, and the sete are very small, laterally with shallow foveate impressions, rather broadly and longi- tudinally convex at the middle. The prosternum is variable, usually feebly convex between the coxe, rounded behind and very feebly mucronate at middle, or with a mod- erate mucro; sometimes horizontally produced, compressed, and ob- liquely truncate. Mesosternum more or less declivous and feebly concave. The metasternal process is about as wide as the abdominal salient is long. The abdominal intercoxal process is slightly transverse (male) or subquadrate (female), and about a fifth of its width wider than the metasternal process, and subequal in length to that of the post-coxal part of the same segment, the latter being quite equal to the second (male), the second segment a little longer (female) than the former. The third segment is about a third longer than the fourth. The profemora are feebly tumid to rather strongly so (male), less strongly and as variable in the female, in both sexes usually more or less shgehtly compressed, or subcylindrical, in transverse section in certain specimens (males) ; tibial grooves not strongly limited by the a a REVISION OF ELEODITNI—BLAISDELL. 57 moderate subcariniform margins that converge, become contiguous, and then evanescent at basal fourth. The grooves are not notably concave. The mesofemora are moderately compressed, subfusiformly and not strongly tumid (male), or gradually wider to near apex in outer half (female) ; grooves not usually well defined, margins rather feeble. The metafemora are feebly widened outwardly with sides sub- parallel; grooves less defined, margins more or less asperulate and evanescent near the middle. The protibiew are scarcely compressed; all the tibiz are without tarsal grooves. The tarsi are moderately long and comparatively stout. The protarsi in the male are about a third longer than in the female. In the male the protarsi are about a fifth of their length shorter than a mesotarsus. Joints subequal in width, two, three, and four subequal in length and width, the fifth scarcely as long as the three preceding taken together; the first not as long as the two following combined. In the female the joints are relatively and proportionally the same. The mesotarsi (male) are a little shorter than a metatarsus. Joints one and five subequal in length; two and three are subequal in length, the fourth a little shorter. A metatarsus (male) is about a fourth of its length shorter than its metatibia. Joint one, three times as long as the third, the fourth quite equal to the combined lengths of the second and third, the latter two are subequal in length. In the female the joints are proportion- ally the same. ELEODES AMPLA var. DOLOSA, new. Elongate-ovate, more or less shining, pronotum polished. Head with the frontal suture obsolete or more or less well marked. Pronotum quadrate, widest at the middle in both sexes, and a little wider (;4) than long; disc polished and shining, very finely and sparsely punctulate, punctures a little denser laterally; sédes rather evenly and not strongly arcuate from apex to base, obsoletely sinuate at the basal fourth; base more or less finely and obsoletely beaded and about a fourth wider than the apex. Elytra with the disc striato-punctate, strial punctures feebly im- pressed, each puncture moderate in size and separated by about their own diameters; interstitial punctures small, distantly spaced, both series more or less irregular at the periphery. Legs long. Anterior tibial spurs moderately short. Otherwise as in ampla. 58 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Male—Body moderately narrow, elongate-ovate. Elytra moder- ately narrowed in posterior fourth. Otherwise as in ampla. Female.—Moderately robust. Elytra less gradually narrowed in the posterior fourth. Otherwise as in ampla. Measurements.—Males: Length, 18-21 mm.; width, 7—-7.5 mm. Females: Length, 21 mm.; width, 9 mm. Genital characters, male —Edeagophore scarcely at all arched. Basale elongate and suboval. 5 A picale elongately triangular; surface extremely sparcely and finely punctulate. Sternite somewhat transverse. Each lobe triangulo-quadrate, with the external border broadly and feebly sinuate at basal three-fourths, thence arcuate to apex, the latter more or less rounded; internal border short and oblique. Otherwise as in ampla. Female.—Genital segment quadrate. Valvula.—Dorsal plate scarcely concave. Appendage very small. Ventrolateral surface quite broadly concave before the apex. Otherwise as in ampla. Habitat—Arizona (along the Colorado River, Beverly Letcher). Number of specimens studied, 7. Sexitypes in my own collection. Type-locality.—Western Arizona. Salient type-characters—More or less shining, elongate-ovate. Pronotum quadrate, polished, widest at the middle. Elytra striato- punctate, strial punctures not impressed (male) or feebly impressed and coarser (female). Legs long. Diagnostic characters—M surface lustre and sculpturing resembles carbonaria; by its larger and more elongate form and long legs it is most closely related to ampla. Mr. Blanchard writes me that it is not to be referred to any species in the LeConte collection. It has heretofore been referred to car- bonaria, but the long legs will quickly separate it from that species. A specimen in Mr. Blanchard’s collection has the sides of the pro- notum more strongly arcuate. The mental, sternal, abdominal, and crural characters are practi- cally the same as in ampla. The anterior tarsi are apparently less elongate in the male, and in both sexes for that matter; in the female joints two, three, and four are comparatively a little smaller. ELEODES OBSOLETA (Say). Blaps obsoleta Say. Journ. Acad. Nat. Sci. Phila., III, 1823, p. 261.— LECONTE, Complete Writings of Thomas Say, II, 1859, p. 1538. Eleodes obsoleta Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 308. Eleodes obsoleta var. porcata Casry, Ann. New York Acad. Sci., V, Nov., 1890, p, 396. REVISION OF ELEODIINI—BLAISDELL. 59 Oblong-ovate, slightly elongate, black, feebly shining to subopaque; elytra striate and slightly scabrous, frequently reddish along the suture. Head twice as wide as long, moderately convex, frontal suture usually distinct, more or less feebly impressed laterally, rather finely and not very densely punctate, punctures slightly sparser on the ver- tex. Antennw moderate and subequal in the sexes, about reaching to the base of the prothorax, outer four joints feebly compressed, slightly and gradually dilated, third joint equal in length to the next two taken together, fourth just perceptibly longer than the fifth, the latter, sixth and seventh subequal, eighth slightly triangular, ninth and tenth irregularly orbicular in outline, eleventh oval and very slightly longer than wide. Pronotum widest at or slightly in advance of the middle, sub- quadrate one-fourth to one-third wider than long; disc moderately and quite evenly convex, more or less declivous at the sides, finely, usually irregularly but quite evenly punctate; apex feebly and evenly emarginate to subtruneate, finely to obsoletely beaded; sides nearly evenly arcuate from base to apex when viewed vertically from above, or more strongly so in the anterior three-fourths, thence oblique and more or less feebly sinuate to base, finely beaded; base more or less evenly and shghtly rounded, finely margined and about a third wider than the apex; apical angles distinct and subacute; basal angles usually obtuse. — Propleure finely, more or less irregularly and submuricately punc- tate, rugulose. Elytra oval, less than twice as long as wide, usually widest at about the middle; base feebly emarginate, scarcely to slightly wider than the contiguous prothoracic base; humeri subobtuse, scarcely at all prominent anteriorly; s¢des evenly arcuate, apex obtusely rounded; disc rather evenly convex, slightly flattened on the dorsum, laterally more strongly and rather evenly rounded, arcuately declivous pos- teriorly; striate, rarely estriate or smooth, strive impressed and rather coarsely, uniseriately punctate near the suture to more irregularly so externally; intervals flat to moderately convex, sparsely and irregu- larly punctate, punctures more or less feebly scabrous; striae less de- fined and punctures more irregular and denser laterally. Hpipleure moderate in width, gradually narrowed from base to apex, superior margin beneath the humeri very feebly and broadly sinuate; surface finely and muricately punctate. Sterna finely to obsoletely punctate. Parapleurw rather densely and finely punctate. Abdomen more or less polished, finely, sparsely punctulate, and more or less rugulose; intercoxal process more strongly sculptured. 60 BULLETIN 63,.UNITED STATES NATIONAL MUSEUM. Legs rather slender and moderate in length; anterior femora mutie in the sexes; anterior tibial spurs dissimilar; anterior tarsi feebly dissimilar in the sexes, first joint slightly thickened at tip beneath. Male.—Elongate, elytra rather gradually narrowed posteriorly, quite evenly and arcuately declivous behind. Abdomen moderately oblique, not strongly convex, more or less feebly impressed at middle on first two segments; intercoxal process somewhat concave. Anterior spurs of the anterior tibiz about twice as long as the posterior, shghtly curved, just feebly widened and gradually narrowed from base to apex, the latter acute. First joint of the anterior tarsi with the minute tuft of modified spinules scarcely evident, ordinary spinules present on the thickened tip beneath. Female.—Robust. Elytra somewhat broadly oval, and slightly narrowed posteriorly, usually arcuately and rather vertically decli- vous behind; abdomen horizontal and strongly convex. Anterior spurs of the anterior tibiz usually about a third to a half longer than the posterior, moderately curved and rather gradually narrowed from base to apex, noticeably widened. First joint of the anterior tarsi with ordinary spinules on the thickened tip beneath. Four forms may be recognized : Forma glabra.—Elvtra estriate and more or less smoothly sculp- tured. Forma typica.—KElytra striato-punctate, intervals slightly convex, subasperate. Forma annectans.—Elytra rather strongly striato-punctate, inter- vals strongly convex; sculpturing subasperate. a ees Forma punctata.—Elytra estriate, irregularly and more or less muricately punctate. Measurements.—Male: Length, 12.5-16 mm., width, 5-7 mm. /e- male; Length, 14.5-18 mm.; width, 6.5-7.5 mm. Genital characters, male—Edeagophore (Plate 3, fig. 1) flax-seed- shaped (flattened oblong-ovate), scarcely arched. Basale oblong-oval, glabrous, moderately convex; sides more or less feebly arcuate. Apicale slightly elongate and triangular; surface in basal moiety evenly convex, in apical half a median longitudinal membranous groove; sides moderately arcuate in basal half, sinuate at middle, hence feebly or scarcely arcuate to apex, the latter acute but not acuminate; base broadly lobed at middle two-fourths, broadly and feebly sinuate laterally. Sternite (Plate 3, fig. 2) parabolic in outline. Each lobe with the external border more or less evenly arcuate from base to apex, fre- quently slightly sinuate in basal half; apex evenly rounded, angle usually not evident; internal border more or less oblique and feebly sinuous; surface rather sparsely punctate in apical moiety, setose, a eh te REVISION OF ELEODIINI—BLAISDELL. 61 sete not dense and moderate in length, longer on apical border. Membrane very sparsely setose across the bottom of the sinus. Female.—Genital segment (Plate 3, fig. 3) quadrate, somewhat de- pressed and setose. Valvula.—Dorsal plate oblong, explanate externally from base to apex, glabrous; sw7face more or less plane, never excavated, some- times with the apical portion slightly deflexed, sparsely punctate, setee moderate in length, longer apically; external margin feebly sinuate towards base, feebly arcuate towards apex, the latter with the external lobe strongly developed, three-fourths as wide as the dorsal plate and broadly rounded; internal border straight to feebly sinuous, the inner lobe of apex minute and separated from the ex- ternal by a small sinuation; apex set with long flying sete. Appendage minutely mammilliform, usually invisible from above; fossa fringed with fine and rather long hairs. Basal prominences not conspicuous. Superior pudendal membrane longitudinally rugulose, reaching to ° the base of the internal apical lobe. Ventrolateral surface (Plate 3, fig. 4) with the basal swollen por- tion quite short, more or less broadly and transversely concave before the apex; surface glabrous, sparsely punctate and minutely setose ; submarginal groove broad and shallow beneath the expanded sides, curving inward beneath the external lobe to the base of the internal lobe. Internal margins of the valves not contiguous at apex; genital fissure broadly fusiform, and nearly closed by the longitudinally rugulose inferior pudendal membrane. Habitat—Forma_ glabra.—Arizona (Williams, Barber, and Schwarz) ; Colorado (Arboles, C. F. Baker). Forma typica—Texas (Mobeetee, H. S. Barber); Nebraska (Grant, July, C. V. Riley; Pine Ridge, Sand Hills, Sioux City, col- lection University of Nebraska); Kansas (F. H. Snow); Colorado (Golden, E. J. Oslar; Colorado Springs, June, elevation 6,000—7,000 feet, H. F. Wickham; Sterling, collection University of Nebraska; Platte Canyon, October, Greeley, H. Soltau; La Junta, Durango, South Park, San Luis Valley, Pueblo, West Cliff, Canyon City, Den- ver; Wickham’s List) ; Wyoming (Cheyenne, C. V. Riley and H. F. Wickham); Oklahoma (Fort Supply, October, H. S. Barber) ; Mon- tana (Assiniboine, Hubbard and Schwarz; Moose Jaw, August, A. N. Caudell; Helena); South Dakota (Badlands, Pine Ridge, September, L. Stejneger; Hot Springs, collection University of Nebraska). Forma annectans.—Colorado (Gulnare, Las Animas County, July 8). Forma punctata.—New Mexico (Coolidge, Wickham) ; Colorado (Edith; Denver, collection of E. C. Van Dyke). 62 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Number of specimens studied, 200. Type destroyed. Ty pe-locality.—Say received his specimens from the “Arid plains of Arkansas and Missouri, in the vicinity of the Rocky Mts.” Salient type-characters.—Body oblong-subovate, elytra with im- pressed striae, which are slightly scabrous with minute elevated points and impressed punctures; interstitial lines also punctured; sutural margin obsoletely reddish brown (Say). Diagnostic characters——This common species is to be recognized from the other members of the group by its duller integuments and striato-punctate elytra. In the typical form the elytra are feebly scabrous at the central part of the disc and more strongly so at the periphery; the striz are very obvious; in some specimens the striz are obsolete or feebly evident and the sculpturing slightly more scabrous (forma punctata), while in some there is a more or less dis- appearance of the asperate punctuation so that they are quite smooth and feebly shining and the striz are feebly marked (forma glabra) ; this form appears to be rare, as I have only seen four samples, and has no doubt heretofore been referred to guadricollis. The small and shghtly more robust form, reddish along the elytral suture, is to be considered the typical form. Say’s measurements are one-half to three-fifths of an inch in length. This form in Colorado and elsewhere has very convex elytral intervals and is here spoken of as forma annectans, and is the homologue of porcata. The var. porcata is a larger and much more elongate form with strongly convex elytral intervals. (See p. 63.) The mentum is moderate in size and varies in form from triangulo- trepezoidal to trapezoido-parabolic; the surface is moderately punc- tate, sete minute, convex at middle, and more or less foveate laterally. The prosternum is rather short before the coxe and protuberant ventrally with the coxw, usually rounded antero-posteriorly between the same; in the small forms not mucronate, but in a few specimens there was a very slight mucro. In the larger individuals and in the punctate form the mucro becomes evident, but I have never seen it well developed. The mesosternum is at times feebly convex, never strongly concave, more or less oblique, but at times it is rather vertically declivous. The metasternal process is about as wide as the abdominal salient is long (male) or slightly narrower (female). The abdominal process is quadrate (male) or shghtly transverse (female) and about a fourth of its width wider than the metasternal salient, equal in length to the post-coxal part of the first segment, the latter being equal in length to the second (male); the second is a little longer in the female, where it is a half longer than the third; REVISION OF ELEODIINI—BLAISDELL. 63 in the male the third is a little longer than the fourth, and in the female a little less than twice as long as the fourth. The metasternum laterally between the cox is as long as the width of a mesotibia at apical third. The femoral and tibial characters are practically as in ampla, ex- cept that the legs are much shorter. The tarsi are moderate in length and rather slender. The protarsi in the male are a little stouter than in the female; the difference in length is less marked, those of the male being just a little longer. The protarsi are about a third (male) or a little less (female) of their length shorter than a mesotarsus. Joints two, three, and four are subequal in size, each about as wide as long and together scarcely as long as the fifth; the first is a little longer than wide. The mesotarsi are about a third (male) or a sixth (female) shorter than a metatarsus. Joints two to four diminish just a little in length in the order named; the fifth is about equal to the combined length of the second and third, and subequal to the first. The metatarsi are about a fourth of their length shorter than a metatibia. Joint three is just noticeably a little shorter than the second, and the two together are subequal in length to the fourth, the latter subequal with the first. ELEODES OBSOLETA var. PORCATA Casey. Oblong-ovate, moderately elongate, black, elytra suleate. Head twice as wide as long, frons slightly flattened to feebly con- vex, faintly impressed laterally, frontal suture usually finely im- pressed ; surface somewhat densely punctate, more coarsely so on the epistoma and finely and sparsely on the vertex; autennw moderate in length and stoutness, outer four joints very slightly compressed, more or less feebly dilated, third joint about equal to the next two taken together, the fourth slightly longer than the fifth, the latter, sixth, and seventh subequal, eighth feebly triangular, ninth and tenth circular in outline, eleventh oval to ovate and very slightly longer than wide. Pronotum subquadrate, one-fourth to one-third wider than long, widest in advance of the middle as viewed vertically from above, at the middle when viewed obliquely from the side; disc gabrous and shining or feebly alutaceous, moderately and evenly convex, more or less arcuately declivous laterally, finely and rather sparsely punctate, the punctures becoming a little coarser, but not very dense laterally ; apex sightly emarginate in circular arc, finely or obsoletely mar- gined; sides quite evenly and moderately arcuate in anterior two- thirds, thence less arcuate and oblique to base as viewed vertically 64 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. from above, or more strongly arcuate at the middle, thence oblique and moderately convergent, feebly sinuate to base as viewed obliquely from the side, finely beaded; base quite evenly and feebly rounded, finely to somewhat obsoletely margined and about one-fifth wider than the apex: apical angles distinct, not acute, frequently narrowly rounded; basal angles obtuse, sometimes more or less rounded. Propleure finely, sparsely to rather densely submuricately punctate and more or less rugulose. Elytra oval, usually widest at about the middle, less than twice as long as wide; dase broadly and feebly emarginate, as wide as or slightly wider than the contiguous prothoracic base; humeri more or less feebly prominent: s¢des evenly arcuate, apex not very narrowly rounded; disc quite evenly convex, occasionally slightly flattened on the dorsum, laterally rather evenly and strongly rounded, arcuately declivous posteriorly, deeply sulcate, the sulci finely, uniseriately to irregularly and muricately punctate; intervals about equal in width to the sulci, very convex, usually strongly defined, finely, sparsely, and irregularly punctate; inflexed sides feebly sulcate, intervals feebly convex, irregularly and rather more densely punctate. Epipleure moderate in width and gradually narrowing from base to apex, superior margin feebly and broadly sinuate beneath the humeri; surface finely, more or less sparsely, submuricately punctate. Sterna more or less finely, densely, submuricately punctate and rugulose. Parapleuvre finely and rather densely punctate. Abdomen somewhat shining, sparsely to more or less densely punec- tate, especially on the first and last segments, more or less rugulose. Legs moderate, comparatively rather longer than in obsoleta. Anterior femora mutic in the sexes. Anterior tibial spurs dissimilar. Anterior tarsi dissimilar in the sexes, first joint shghtly thickened at tip beneath, with a small tuft of spinules. Male—Body somewhat slender. Elytra rather gradually nar- rowed posteriorly, evenly arcuately and somewhat obliquely declivous at apex. Abdomen slightly oblique, moderately convex and more or less broadly flattened at middle on first two segments, intercoxal process more or less concave. Anterior spurs of the anterior tibiz about a half longer than the posterior, distinctly wider and gradually narrowing from base to apex. Anterior tarsi longer than in the female, first joint with a minute and inconspicuous tuft of modified spinules, surrounded by ordinary spinules on the thickened tip beneath. Female.—Body robust. Elytra rather broadly oval, rather rapidly narrowed, arcuately and vertically declivous posteriorly. Abdomen horizontal and strongly convex. Anterior spurs of the anterior tibiz about a third longer than the posterior, distinctly broadened, REVISION OF ELEODIINI—BLAISDELL. 65 gradually but less rapidly narrowing from base to apex. First joint of the anterior tarsi with tuft of ordinary spinules at tip beneath. Measurements.—Males: Length, 12-18 mm.; width, 6-7 mm. Fe- males: Length, 18.5-21 mm.; width, 7.5-9.5 mm. Genital characters, male-——Edeagophore flaxseed-shaped (flattened oblong-ovate), slightly elongate and not usually arched. Basale oblong, glabrous, evenly convex; sides more or less parallel. Apicale triangular, shghtly elongate, apical region more or less deflexed; surface glabrous, rather strongly convex at middle, with a long median membranous groove reaching nearly to the base, groove linear in basal third; s¢des broadly sinuate at middle third, slightly arcuate toward base; apex acute and rather gradually narrowed; base very broadly lobed at middle and very feebly sinuate laterally. Sternite slightly transverse. Each lobe with the external border feebly arcuate and oblique, becoming arcuate at apex, the latter broadly rounded or feebly subtruncate, angle scarcely evident; inter- nal border nearly straight; surface rather sparsely punctate in apical two-thirds, setose, setee moderately long on apical border, shorter toward base. Membrane sparsely setose across the bottom of the sinus, which is narrow and triangular. Female.—Genital segment quadrate and setose. Valvula (Plate 3, fig. 14).—Dorsal plate somewhat broadly oblong, moderately explanate externally and at apex, frequently with a marked antero-posterior convexity; surface plane, sparsely punctate along the internal moiety and at apex, sete not long; external and internal borders more or less straight and quite parallel; apex with the external lobe broadly rounded and apparently semi-chitinous, the internal lobe small and membranous as usual and separated from the external by a small sinuation at junction of the first and second inner fourths of the apical margin of the dorsal plate. External lobe set with long flying sete, a few on the internal lobe. Appendage small, inferior, and more or less compresso-mammilli- form. Fossa rather large and fringed with sete. Basal prominences not conspicuous. Superior pudendal membrane as in obsoleta. Ventrolateral surface feebly convex toward base, broadly and shal- lowly concave before the apex, glabrous, finely and very sparsely punctate, sete: minute, denser at apex; submarginal groove shallow and not well defined at the apex, not reaching the internal lobe. Internal margins of the valves contiguous at basal sixth. Genital fissure quite broadly fusiform. Inferior pudendal membrane regulose and largely visible. It is to be noted that the sides of the dorsal plates are not as strongly explanate as in obsoleta, 09780—Bull. 683—09——5 =. 66 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Habitat—Arizona (Fort Apache, Casey; Galiuro Mountains, May ; Fort Grant, July; Chiricahua Mountains, May, Hubbard and Schwarz; Williams, July; Ash Fork, June; Flagstaff, July, Barber and Schwarz; Kearn’s District, Navajo Indian Reservation, April A. W. Barber; Peach Springs, Walnut, Winslow, July; Seligman, July, H. F. Wickham; Prescott, H. C. Fall); New Mexico (Pecos, July). Number of specimens studied, 85. Type is in Col. Thomas Casey’s collection. Type-locality—F ort Apache, Ariz. Salient type-characters.—Prothorax with the dise evenly convex, finely, sparsely punctate, the punctures becoming rather coarse later- ally but not very dense; apex feebly emarginate; sides more strongly arcuate before the middle, thence moderately convergent and gradu- ally feebly sinuate to the basal angles, which are very obtuse but not distinctly rounded; base feebly and evenly arcuate. Elytra with the dise very deeply sulcate, the sulci finely, rather sparsely and muricately punctate, the intervals equal in width to the sulci, very convex, finely and sparsely punctate (Casey). Diagnostic characters.—The strongly sulecate elytra separates this race from all others of the group. The individuals are unusually larger and more elongate than obsoleta. Well developed specimens appear quite distinct from obsoleta, but they mark a heterotypical variation which is connected to the typical form of obsoleta .by an abundance of mesotypes of all sizes and sculpturing, so that it can only be considered as a good variety. Some individuals are nearly as large as ampla, notably a series from Fort Grant, Arizona, collected by Hubbard and Schwarz. I have authentic examples. before me kindly contributed to Mr. Charles Fuchs and myself by Colonel Casey. The mentum is moderate and varies in form from trapezoidal to trapezoido-parabolic; surface is rather strongly punctate, sete small, moderately convex and more or less strongly foveate laterally and narrowly impressed along the apical margin, the latter being fre- quently deflexed. In Casey’s types the prosternal process is prolonged and prominent. In the series before me it is very variable—from a small mucro to well developed, or compressed and subtruncate behind. The mesosternum varies relatively with the foregoing, concave to convex, shghtly oblique to vertically declivous. The intercoxal process of the first abdominal segment is subquad- rate (male) or slightly transverse (female), and about a third of its width wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a mesotibia at apex. REVISION OF ELEODIINI—BLAISDELL. 67 The post-coxal part of the first segment is equal in lengtn to the process, equal in length to the second segment in the male, and the third segment in the female. In the male the third segment is a third of its length longer than the fourth; in the female the second is twice as long as the fourth. The legs are usually quite strongly sculptured. The profemora and the metafemora are as in carbonaria. The mesofemora are usu- ally, gradually and feebly widened from base to apex. The protibize are subcylindrical in transverse section. All the tibie are without tarsal grooves and the articular cavities are closed. The tarsi are moderate in length, comparatively a little stouter than in obsoleta. A protarsus in the male is about a sixth of its length longer than in the female, and subequal in stoutness in the two sexes. The relative proportions of the tarsi to each other, and the con- stituent joints of each one to each other is practically the same as in obsoleta. ELEODES KNAUSII, new species. Oblong-ovate to ovate, more or less shining, estriate and moder- ately convex. Head twice as wide as long, feebly convex, frontal suture feebly marked, frons very slightly impressed laterally, somewhat coarsely, irregularly and more or less densely punctate, especially on the epis- toma. Antenne moderate in length, outer four joints feebly com- pressed, distal three slightly dilated, third about as long as the next two taken together, fourth scarcely longer than the fifth, the latter, sixth and seventh subequal in size and about as wide as long, eighth shorter and apparently a little wider than long, ninth subtriangular and shghtly transverse, tenth transversely oval, eleventh ovate. Pronotum subquadrate, widest just in advance of the middle when viewed vertically from above, at the middle when viewed obliquely from the side, and about a third wider than long; disc evenly and moderately convex, rather arcuately declivous at the sides and apical angles, quite finely, irregularly and somewhat sparsely punctate. punctures shghtly denser at the sides; apea more or less feebly emarginate in circular arc, finely and more or less obsoletely beaded ; sides moderately arcuate in anterior three-fourths, thence obliquely convergent to base as viewed vertically from above, or more strongly arcuate anteriorly, thence convergent and more or less sinuate to base when viewed obliquely from the side, finely beaded; base feebly arcuate, sometimes feebly sinuate at the middle, finely and more or less obsoletely beaded, one-fifth to one-third wider than the apex; apical angles distinct, not at all prominent anteriorly, sometimes very feebly and narrowly rounded; basal angles obtuse. 68 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Propleurw obsoletely to finely, sparsely submuricately punctulate and rugulose. Elytra oval, distinctly less than twice as long as wide, widest at the middle; dase more or less emarginate, usually feebly sinuate each side of the middle, and generally shghtly wider than the contiguous prothoracic base; humert more or less exposed and obtuse; sides evenly arcuate, apex obtuse and rather narrowly rounded; disc mod- erately convex on the dorsum, quite evenly and strongly rounded at the sides, arcuately declivous posteriorly; almost coarsely, irregu- larly and rather densely punctate, the punctures are subequal in size and obsoletely muricate, frequenty there is a tendency to an arrange- ment into rather distant rows on the dorsum; laterally there is a shght tendency to rugulosity. Epipleurw moderate in width and gradually narrowing from base to apex; superior margin feebly and broadly sinuate beneath the humeri: surface dull, finely and irregularly punctate. Sterna obsoletely to rather densely and finely punctate, more or less rugose. Parapleure finely and quite densely punctate. Abdomen finely, sparsely, and irregularly punctate, more or less rugulose, more coarsely sculptured on the first segment, the fifth coarsely punctate. Legs rather short and somewhat slender, more or less strongly sculptured. Anterior femora mutic; anterior tibial spurs compara- tively small and feebly dissimilar in the sexes. Anterior tarsi with the first joint shghtly thickened at tip beneath and a httle dissimilar in the sexes. JMale.—Oblong-ovate, moderately slender. Antenne reaching to the base of the prothorax. Elytra moderately narrowed in apical fourth, arcuately and scarcely obliquely declivous posteriorly. Ab- domen moderately oblique and convex, more or less broadly im- pressed at middle of the. first two segments. Anterior spur of the anterior tibie about a half longer than the posterior, rather slender and gradually narrowing from base to apex, acute and feebly curved. First joint of the anterior tarsi with a small in- conspicuous pencil of modified spinules, surrounded by ordinary spin- ules on the thickened tip beneath. Female.—Ovate and robust. Antenne not quite reaching to the prothoracic base. Elytra feebly narrowed and arcuately, vertically declivous in apical fourth. Abdomen horizontal, strongly convex. Anterior spurs of the anterior tibie about a half longer than the posterior, slightly broadened, gradually narrowed from base to apex; both spurs rather strongly curved. First joint of the anterior tarsi with ordinary spinules on the thickened tip beneath. REVISION OF ELEODIINI—BLAISDELL. 69 Measurements—Males: Length, 14.5 mm.; width,6mm. Females: Length, 14-16 mm.; width, 7-8 mim. Genital characters, male—Kdeagophore subfusiform and rather elongate, not arched. Basale oval-oblong, evenly convex, with sides feebly arcuate. Apicale triangular, slightly elongate; surface evenly convex with a median membranous groove extending from near the tip almost to the base, where it becomes shghtly less membranous; s¢des arcuate in basal half, thence more or less sinuate to apex, the latter very gradually and slightly decurved and very narrowly rounded. _ Sternite quite parabolic in outline. Each lobe with the external border quite evenly arcuate; apex rather narrowly rounded, angle not evident; internal border short, straight and oblique; surface slightly convex, glabrous, rather coarsely punctate in apical half, setose, setae moderate in length; punctures denser and sete longer at apex. Membrane slightly setose at bottom of the sinus. Female.—Genital segment quadrate, somewhat depressed, setose and not strongly chitinized. Valvula (Plate 4, fig. 24.)—Dorsal plate oblong and glabrous; surface plane, finely and sparsely punctate, with rather short reclin- ing sete arising from the same; external border straight or feebly sinuous, gradually explanate from the basal third to the apical margin, the latter rounded and explanately produced upon the ex- ternal lobe in outer three-fourths, directly and arcuately continuous with the external border; external apical lobe rather feebly mem- branous beneath and concealed from view from above; internal lobe moderate and separated from the external by a small sinuation in the apical margin; internal border straight or sinuous; apical region set with a few long, flying, soft sete. Appendage slightly visible from above, directed backward and feebly inward, conico-mammilliform. Fossa inferior, rather large, and fringed with long, flying sete. Superior pudendal membrane longitudinally rugulose and reaching to the base of the internal apical lobe. Basal prominences not noticeable. Ventrolateral surface not strongly convex in basal moiety, more or less concave laterally before the apex, but not transversely so; sub- marginal groove broad and shallow beneath the explanate external border of the dorsal plate and apical lobe, meeting the fossa at its external edge; external apical lobe not strongly developed beneath ; surface finely and irregularly punctulate and very finely setose. Inter- nal margins of the valves contiguous for a short distance at base and apex, between which the fissure is rather broadly fusiform, and closed in basal two-thirds by the inferior pudendal membrane, the latter feebly rugulose. 70 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Habitat—New Mexico (Cloudcroft, James Canyon, June, Warren Knaus). Number of specimens studied, 8. Sexitypes in my own collection. Ty pe-locality.—Cloudcrott, New Mexico. Collector, Warren Knaus. Salient type-characters.—More or less shining, estriate and moder- ately convex. Pronotum subquadrate; disc rather finely, irregularly and sparsely punctate, punctures denser at the sides; apex very feebly emarginate; apical angles distinct and not at all prominent ante- riorly; basal angles obtuse. Elytra with the humeri more or less exposed; dise almost coarsely, irregularly, and rather densely punctate, punctures subequal in size and obsoletely submuricate; laterally denser, with a slight tendency to rugulosity. Legs rather short and somewhat slender. Diagnostic characters.—Resembles the robust form of extricata or vileyé in general outline. If it inhabited the region where parvicollis is found it might be mistaken for that species; some forms of leconte? resemble it. First of all, the genital characters distinguish it from all the above and associates it with obsoleta which it resembles, but less strongly in general habitus. It is usually confounded with extricata, from which it differs in the mutie anterior femora of the male, the more strongly arcuate sides of the pronotum at anterior two-fourths, and besides the punctuation is quite different. In rileyi the female has the anterior spurs of the anterior tibiv strongly developed, while in nausti they are but feebly broadened. The spurs have greater development in the male of 7i/eyi also than in the male of Anausi. In parvicollis and lecontei the lateral margins of the pronotum is distinetly visible from above, while in Anausi the pronotal sides are arcuately declivous. The genital and tarsal characters are also dis- tinctive. Anausii differs from typical obsoleta in its smooth and shining sur- face and estriate elytra. It is also more robustly ovate than in the smooth form of obsoleta. IT am indebted to Mr. Warren Knaus for this interesting species, and I take pleasure in dedicating it to him. Nnausii appears to bear the same relation to the Carbonaria section that ri/ey7 bears to the Guadricollis section. In this respect the two species are analogous. There is a specimen before me from Cananea, District of Arizpe, State of Sonora, Mexico, that resembles /nausc in form, but it is more alutaceous and smoother, the striae of punctures are more evident and the interstitial punctures smaller and less conspicuous. The anterior tibial spurs are also feebly developed, and while the study of the | | | | i | REVISION OF ELEODIINI—BLAISDELL. ll United States specimens leaves the relationships in doubt, a knowl- edge of the Mexican contingent would undoubtedly be more elucida- tive. At present /:nausii appears as a derivative of obsoleta. The mentum is more or less trapezoido-parabolic, rather densely punctate and subfoveate laterally, somewhat convex at the middle; the sete are scarcely evident. The prosternum is moderately produced behind, feebly convex or horizontal between the coxe and triangularly dilated behind the median transverse axis of the acetabula; the mucro is small, at times the process is compressed and more or less vertically truncate behind. The mesosternum is quite vertically declivous and more or less feebly concave. The intercoxal process of the abdomen is subquadrate (male) or shghtly transverse (female), and about a fourth (male) or a third (female) of its width wider than the metasternal salient. The metasternum laterally between the cox is as long as the width of a mesotibia at apical third. The post-coxal part of the first abdominal segment is subequal (male) or equal (female) in length to that of the process. In the male the second segment is equal to the length of the process; the third is about equal to the post-coxal part of the first; the fourth is about two-thirds as long as the second. In the female the second segment is about twice as long as the fourth, the third is equal to the post-coxal portion of the first. The legs are noticeably short. The profemora of the male are moderately clavate and feebly com- pressed; in the female feebly tumid and moderately compressed ; the tibial grooves are limited by feeble but distinct subeariniform mar- gins which meet and become evanescent at the basal fourth. In both sexes the meso and metafemora are quite similar. The mesofemora have the surface lines feebly but distinctly arcuate, and the femora in outline may be said to be subfusiform. The meta- femora have the superior and inferior surface lines quite parallel. The grooves are limited by asperulate margins which converge at basal third on the mesofemora and become evanescent, while on the metafemora they become evanescent about the middle without be- coming contiguous. The tibizw are all more or less feebly arcuate, and rather more strongly widened apically than is usually observed; the tarsal grooves are absent and the articular cavities are closed. The protibixe are observed to be feebly compressed or subeylindrical. The tarsi are moderately long and rather stout when considered in a comparative sense. The protarsi of the male are about a fourth longer than those of the female. Tq BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The protarsi are about a fourth (male) or two-thirds (female) of their length shorter than a mesotarsus. The mesotarsi are about a ninth—less in the female—of their length shorter than a metatarsus. The metatarsi are about a seventh (male) to two-fifths (female) of their own length shorter than their respective metatibia. The relative proportions of the joints of each tarsus are practically the same as in obsoleta. ELEODES OMISSA LeConte. EHleodes omissa LECONTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.—Horn, Trans. Amer. Philos. Soe., XIV, 1870, p. 308. Hleodes interruptad BLAISDELL, Ent. News, III, Dec. 1902, p. 241. Oblong-ovate to ovate, elongate, and more or less shining. Head twice as wide as long, more or less convex, feebly impressed laterally, frontal suture fine and usually evident, rather finely and not densely punctate, punctures sparser on vertex. Antenne moderate, outer four joints very feebly compressed, scarcely dilated, third joint equal in length to the next two taken together, fourth shehtly longer than the fifth, the latter, sixth, and seventh subequal, eighth just the least shorter, subtriangular, and longer than wide, ninth and tenth somewhat circular in outline, eleventh ovate. Pronotum subquadrate, widest at or in front of the middle as viewed vertically from above, one-fourth to one-third wider than long; disc evenly and moderately convex, more or less arcuately declivous at the sides, finely, sparsely, and more or less irregularly punctulate, punctures just a little coarser and denser laterally; apex feebly emarginate or subtruncate in circular arc, finely or obsoletely beaded; sides evenly and moderately arcuate in the anterior two- thirds, thence to the base more or less straight and converging (viewed vertically from above), or evenly and rather broadly arcuate, converging and more or less feebly sinuate (viewed obliquely from the side), finely beaded; base shghtly rounded to subtruncate, finely margined, one-ninth to a fourth wider than the apex; apical angles subacute to obtuse, rarely feebly prominent anteriorly; basal angles obtuse, frequently apparently rounded when viewed vertically from above. Propleure finely and very sparsely punctate, more or less rugulose, rarely rugose. Elytra oval, widest at the middle, less than twice as long as wide; base feebly emarginate to truncate, usually scarcely wider than the contiguous prothoracic base; humeri obtuse and not prominent ; sides evenly arcuate, apex obtusely rounded; disc moderately convex on the dorsum, rather strongly and evenly rounded laterally, arcuately declivous posteriorly, punctate, punctures frequently of the same REVISION OF ELEODIINI—BLAISDELL. (o size, fine, diffusely arranged, not dense, generally some evidence of a serial order; the strial punctures are frequently slightly the larger, the series are moderately distant and rarely impressed. Epipleure moderate in width, gradually narrowing from base to apex, very feebly sinuate along the superior margin beneath the humeral region; surface finely and sparsely punctulate. Sterna more or less shining, rather densely punctate and more or less rugose. Parapleurw quite densely punctate. Abdomen glabrous, finely and very sparsely punctulate, usually more or less distinctly rugulose—almost rugose at times. Legs moderate in length and stoutness, sometimes moderately thickened; anterior femora mutic and the anterior tibial spurs slightly dissimilar in the sexes; anterior tarsi with the first joint shghtly thickened at tip beneath and slightly dissimilar in the sexes. Male.—Oblong-ovate, rather slender and elongate. Elytra grad- ually narrowing in the posterior third, arcuately and somewhat obliquely declivous posteriorly. Abdomen oblique, moderately con- vex, more or less flattened at middle of the first two segments. Anterior spurs of the anterior tibiwe slightly thickened, nearly twice as long as the posterior, both comparatively short. First joint of the anterior tarsi set with a small tuft of spinules, the modified spinules scarcely evident. Female.—Ovate and robust. Elytra broadly oval, and somewhat eradually narrowing in the posterior fourth, arcuately and vertically declivous behind. Abdomen horizontal and strongly convex. An- terior tibial spurs rather variable in length, the anterior distinctly broadened and gradually narrowing from base to apex, sometimes with sides slightly arcuate in basal half, and about a third longer than the posterior, both acute, tapering, and stouter than in the male. First joint of the anterior tarsi set with a small transverse tuft of ordinary spinules on the thickened tip beneath. T have observed only four forms worthy of distinction in the hun- dreds of specimens before me of this variable species. Forma typica.—Thorax somewhat transverse, apex feebly emar- ginate or truncate, angles somewhat obtuse and not at all prominent anteriorly. Femora distinctly thickened. Male.—¥lytra rather more elongate than usual. Female.—FElytra with the sides more strongly arcuate anteriorly than usual. Forma cataline.—Pronotum more strongly punctate, especially laterally. Femora more or less thickened. Forma communis.—Thorax as in the typical form. Elytra with the sides less strongly and suddenly arcuate anteriorly. Femora not noticeably thickened. 74 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Forma emarginata.—Thorax quadrate, apex distinctly emarginate, angles subacute and somewhat prominent anteriorly. Measurements.—M ales: Length, 16-19 mm.; width, 6-7 mm. Fe- males: Length, 18-23 mm.; width, 8-10 mm. Anomaly.—The specimen described by me“ as interrupta belongs here. The prothorax has the side margin at middle rather abruptly interrupted. The elytra has the disc sulecate posteriorly. Length, 16.6 mm.; width, 7.1 mm. Genital characters, male-—KEdeagophore (Plate 2, fig. 1) elongate- ovate to fusiform, and more or less arched. Basale feebly oblong-oval to oblong; surface evenly convex; sides feebly arcuate, sometimes converging slightly toward the apex. Apicale triangular, slightly elongate; swrface moderately and evenly convex, with a median longitudinal grove in apical half, sometimes extending from apex to base, shehtly dilated and mem- branous apically, gradually narrowing to become linear basally; sides moderately and evenly arcuate in basal half, thence rather feebly sinuate to apex, the latter subacute; base rather broadly lobed at middle and feebly sinuate laterally. Sternite (Plate 2, fig. 2) transversely parabolic. Each lobe more or less subparabohe in outline; external border straight to evenly arcuate, the apex being continuously but more narrowly rounded, angle sometimes evident; internal border quite short and_ feebly arcuate; surface scarcely convex, sparsely punctate and setose in apical three-fourths, sete rather long on the apical border. Mem- brane distinctly and sparsely setose at the bottom of the sinus, the latter short. Female.—Genital segment (Plate 2, fig. 3) quadrate, moderately explanate laterally and setose. Valvula—Dorsal plate oblong, moderately narrow, sides subpar- allel; surface plane, glabrous, sparsely punctate, impunctate at basal third, setose, setae moderate and reclining; external border straight to feebly arcuate, continuously so with the more strongly and evenly rounded apical border, which is more or less inwardly oblique; in- ternal lobe short and separated from the external by a small sinua- tion; external lobe set with rather long sete, the internal with a few rather short ones; internal border slightly sinuous. Appendage just visible from above, short mammilliform, with a pencil of sete at tip. Fossa moderate and in the internal wall of the external apical lobe. basal prominences scarcely evident. Superior pudendal membrane longitudinally rugulose, reaching about to the internal lobe of the apex. 4¥Ent. News for Dec., 1902. REVISION OF ELEODIINI—BLAISDELL. 15 Ventrolateral surface (Plate 2, fig. 4) moderately convex, slightly concave laterally before the apex; surface sparsely punctate and finely setose; submarginal groove well developed beneath the explanate ex- ternal border of the dorsal plate, passing obliquely inward beneath apex to the inner angle of the fossa, ending at the internal apical lobe. Internal margins of the valves contiguous at basal sixth; geni- tal fissure quite broadly fusiform, and closed at basal half by the inferior pudendal membrane. Mabitat.—Southern California; Santa Catalina Island. Number of specimens studied, 500. Type is in the LeConte collection. Ty pe-locality—San Diego, California. Salient type-characters—Longer than usual. Thorax slightly con- vex, sides broadly rounded, somewhat narrowed posteriorly, finely and sparsely punctulate. Elytra declivous and obliquely narrowed behind, sparsely, finely, and very irregularly punctate. Femora un- usually thick (LeConte). Diagnostic characters.—The most common species in Southern Cali- fornia and always heretofore referred to qguadricollis, from which it differs in the smaller and more slender anterior spurs of the anterior tibiz (females) ; from carbonaria, to which it is most closely related, by the punctures of the elytral dise having a less distinct serial ar- rangement, and by the serial punctures being scarcely larger than those of the intervals, the tendency being toward diffuse punc- tuation. Omissa is with greater difficulty separated from the smooth form of carbonaria, where the elytra punctures are small and of the same size; here locality must be relied upon for their separation. All those specimens from the eastern side of the Colorado River are forms of carbonaria, while all those from the western side are to be referred to the present species. I do not know of a single specimen taken west of the Colorado River ever having been identified as an authentic carbonaria. In fact I consider omissa as a western modification of carbonaria. Omissa is a variable species in regard to the form of the pronotum, but in the several hundreds which I have studied I found an abun- dance of all necessary mesotypes, both as regards the form of the pronotum and elytral punctuation. In a large percentage of those individuals that have the sides of the pronotum evenly arcuate from apex to base, it will be found that they really present a more normal form of the side margin than do those that are widest in front of the middle; for here it will be seen that the sides of the pronotal disc are most strongly declivous behind the middle and that the true margin is displaced downward and not visible when the pronotum is viewed vertically from above; 76 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. when viewed obliquely from the side the normal marginal curve will be seen and that in reality the pronotum is widest at the middle. It must also be borne in mind that the pronotal dise is normally moderately declivous laterally in the present subgenus. In many specimens (mostly male, but some females as well) the pronotal sides are less strongly deflexed and appear quite evenly rounded from apex to base, and as a result the pronotum is broader as compared to those with strongly deflexed sides where the thorax is less rounded and more narrowed to the base. Although the punctuation is variable, usually there is always some evidence of a serial arrangement; rarely, the strial punctures are rather large and even moderately impressed; these specimens ap- proach carbonaria in this respect, but oméssa is as a rule more elongate in the male and more broadly ovate in the female. The strongly punctate individuals are not necessarily confined to the region bounding the Colorado River on the west, but are to be taken at any part of the distributional area. The larger Southern California specimens are to be referred to omissa, as the characters exhibited by the anterior tibial spurs are those characteristic of the present section of the subgenus; with these conceptions of oméssa and the elimination of qguadricollis everything becomes clear. Ampla is a larger, more elongate species with elongate legs. For the differential characters of the varieties pygmaa and penin- sularis see below. I am indebted to Prof. H. C. Fall for notes on comparisons of specimens with the LeConte type. The mentum is variable, rather small to moderate in size, and tri- angulo-trapezoidal to trapezoido-parabolic; surface rather strongly punctate, more or less foveate laterally and convex at middle.) The prosternum is convex or horizontal between the coxe and mucronate behind; the mucro may be small and subacute; larger and conical in contour, or compressed and obliquely to nearly vertically truncate behind. The mesosternum is more or less concave, and varies in the degree of its obliquity. The intercoxal process of the abdomen is quadrate (male) or a little transverse (female), and about a fifth (male) or a sixth (fe- male) of its width wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a mesotibia at the middle. The post-coxal part of the first abdominal segment is equal (male) or subequal (female) in length to that of the process. In the male the second segment is about a half longer than the third and twice as long as the fourth, and equal to the length of the process. REVISION OF ELEODIINI—BLAISDELL. a In the female the second segment is about a third of its length longer than the third and twice the length of the fourth. The profemora in the typical form are distinctly tumid and clavate, and in the males of the other forms and all females more or less mod- erately so. The grooves are rather broad and the margins are feebly subeariniform, converging quite close to the base. The mesofemora are gradually and feebly thickened externally ; the metafemora have their superior and inferior borders subparallel. The grooves are dull and feebly subasperate, the margins finely asperulate and more or less evanescent before becoming contiguous. The protibie are more or less feebly arcurate and slightly com- pressed. All the tibie are usually without tarsal grooves, although an occasional specimen exhibits rudimentary grooves on the protibiz with the articular cavities shghtly open, otherwise the cavities are closed. The tarsi are moderate both as to length and stoutness. The protarsi in the male are just noticeably longer than in the female, and specimens of corresponding size must be selected, as a large female compared with a small male would not show the true difference. The protarsi are about two-fifths (male) or three-sevenths (female) of their length shorter than a mesotarsus. Joints two, three, and four are subequal, just noticeably smaller in the female, and together about equal to the fifth in length; the first is about equal to the combined lengths of joints three and four. The mesotarsi are about a seventh (male) or an eighth (female) of their length shorter than a metatarsus. Joints one and five are subequal in length, and the combined lengths of two, three, and four just a little longer than either. The metatarsi are about three-fourths (male) or a half (female) of their length shorter than a metatibia. Joint two is a little longer than three; the first is as long as the combined length of third and fourth. ELEODES OMISSA var. PYGMAZA, new. Oblong-ovate to ovate, more or less shining and smooth. Head finely punctate, punctures denser on the epistoma, frons feebly convex to flat. Antenne moderate, outer four joints scarcely dilated, fourth, fifth, sixth, seventh, and eighth subequal in length, ninth and tenth suborbicular and slightly transverse. Pronotum quite quadrate, widest at or just in front of the middle, one-fifth to one-fourth wider than long; disc evenly and moderately convex from side to side, finely and sparsely punctate; apex truncate in circular arc; sides evenly and not strongly arcuate in anterior two- thirds, thence more or less straight and converging to base, or some- 78 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. times apparently quite evenly arcuate from apex to base, the latter quite truncate, equal to the length, a seventh to a fourth wider than the apex; apical angles obtuse, not in the least prominent anteriorly, sometimes feebly rounded; basal angles obtuse. Elytra oval, quite evenly and more or less moderately convex on the dorsum, more strongly rounded laterally; dase scarcely emarginate, equal to or slightly wider than the base of the prothorax; humer? obtuse, scarcely prominent or rounded; sides evenly arcuate, apex obtusely rounded; dsc finely and diffusely punctate, a serial arrange- ment usually evident, punctures generally of the same size, serial punctures sometimes slightly larger. Otherwise as in omissa. Male——Usually quite slender and oblong-oval, frequently more ovate, about three times longer than wide. Elytra scarcely wider than the prothorax in the typical form, at other times wider. Both sexes otherwise as in omissa. ; Measurements —Males: Length, 13-15 mm.; width, 4.5-5.2 mm. Females: Length, 13-16.5 mm.; width, 6-7 mm. Genital characters as in omissa (Plate 2, figs. 1, 9, and 10.) Habitat—Southern California (San Diego to Kaweah, Tulare County). Number of specimens studied, 1,000. Sexitypes in my own collection. Type-locality.—San Diego, Cal. Salient type-characters—Much smaller than omissa. Thorax quite quadrate. Elytra at the widest point scarcely wider than the pro- thorax in the male. Diagnostic characters.—Typical specimens are much smaller than omissa, the larger individuals becoming indistinguishable from it. Pygmea is of especial interest from the fact that it has been generally distributed as gentilis. The female type before me was identified as gentilis by Colonel Casey, and is the species referred to by him in his Coleopterological Notices, VII,“ where he writes “that a very large series of gentilis which I took at San Diego shows quite clearly that this species should be associated with guadricollis and vicinus, and is out of place in the present arrangement.” Gentilis is a different species altogether and is a race of gigantea. - Pygmea is the most abundant species about San Diego, where I have seen it so abundant beneath boards, tin cans, sacks, and cow- chips, that they actually carried these things about over the ground. These remarks apply to the early “ seventies,” when San Diego con- sisted of scattered houses and the saline flats, covered by ice plants (Mesembryanthemum crystallinum) formed the greater part of the “Annals N, ¥Y. Acad. Sci., V, Nov., 1890, p. 395. REVISION OF ELEODIINI—BLAISDELL. 79 maritime landscape, then these insects were gregarious in untold hundreds. I have never found them so plentiful in recent years. Specimens from Tulare and Kern counties represent a slightly. different incipient race, which may be characterized as follows: Forma borealis.—Males less oblong and a little more ovate. I am indebted to Mr. Ralph Hopping for the privilege of studying a large series. In Professor Fall’s collection there is a female that is more opaque than usual, but otherwise it does not differ to any extent; it was col- lected in the San Bernardino Mountains, California. Var. peninsularis is larger, glabrous, and shining, found farther south upon the peninsula of Lower California (See below). I can not determine that Pygmea has any other characters worthy of mention as distinguishing it from omdssa, with which it agrees in mental, sternal, abdominal, and crural characters. It might be mentioned that the prosternum is more or less convex between the coxe and mucronate behind; the mucro is small. The mesosternum is feebly oblique and slightly concave. ELEODES OMISSA var. PENINSULARIS, new. Suboblong-ovate to ovate, glabrous and shining, elytra obsoletely subsulcate. Head quite finely punctate; antennw somewhat slender, eighth joint scarcely shorter than the seventh. Pronotum subquadrate, widest at the middle, a fifth to a fourth wider than long; disc evenly and moderately convex from side to side, scarcely more arcuately declivous laterally, but distinctly so at the apical angles, somewhat obsoletely, finely and sparsely punctu- late; apex truncate to feebly emarginate in circular arc; sides moderately and rather evenly arcuate, somewhat oblique posteriorly, briefly and very feebly sinuate before the base, as seen obliquely from the side; base feebly rounded, and scarcely to one-fifth wider than the apex; apical angles subacute, with a tendency to become feebly prominent anteriorly; basal angles obtuse and not rounded. Propleure obsoletely punctulate and more or less rugulose. Elytra oval, usually widest behind the middle; désc¢ striato-punc- tate, punctures generally not distinctly defined, more or less eroded, striae somewhat impressed, intervals apparently somewhat feebly con- vex; strial punctures rather moderate in size and rather closely placed, intervals minutely, irregularly and obsoletely punctulate, punctules rather more distinctly defined laterally, but not usually confused. Epipleure obsoletely punctulate. Sterna obsoletely punctate and more or less rugose. 80 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Legs rather somewhat slender. Femora moderately and rather sparsely punctate; anterior tibial spurs rather feebly developed. First joint of the anterior tarsi very feebly thickened at tip beneath. Otherwise as in omissa. JMale.—More or less oblong-ovate and somewhat slender. Abdomen feebly oblique. First joint of the anterior tarsi with a minute and pointed tuft of modified spinules on tip beneath. Female.—Ovate, anterior spur of the anterior tibiz not strongly differentiated and not elongate as usual. Measurements —Males: Length, 15-17 mm.; width, 6—-6.5 mm. Females: Length, 16-18 mm.; width, 7-8 mm. Genital characters.—M ale.—Edeagophore as in omissa. Sternite parabolic in outline, slightly transverse. Each lobe sub- triangular; external border quite evenly arcuate, with apex introrsely placed and narrowly rounded; internal border straight; surface shghtly convex, glabrous and not densely punctate in apical half, setae moderate in length and not dense. Membrane sparsely setose at bottom of the sinus, the latter rounded and subparabolie. Female.—Genital segment quadrate; dorsal surface plane and setose. Valvula.—Dorsal plate oblong, aoderntely wide and explanate apically; surface nearly plane, very glabrous, sparsely and _ finely punctate, setae moderate, longer towards apex; borders subparallel, the external sinuous at basal third, thence feebly arcuate to and con- tinuously so with the broadly and strongly rounded apical margin, which is minutely sinuate at internal fourth; internal margin sinuous. Apical external lobe inferior, the internal small, both with a few long flying hairs. Appendage small mammilliform, not visible from above, with few long hairs on tip. Fossa moderate and fringed with long soft sete. Basal prominences not visible laterally. Superior pudendal membrane longitudinally rugulose and reaching to the base of the internal apical lobe. Ventrolateral surface strongly convex at the basal third, thence to apex broadly concave; submarginal groove broad and well marked basally, thence passing into the general concavity; surface in apical two-thirds sparsely and finely punctate and setose. - Internal mar- gins of the valves contiguous at basal sixth and subcontiguous in apical two-sixths; fissure broadly fusiform and closed at the basal half by the inferior pudendal membrane, the latter rugulose. fabitat.—Lower California (Sierra San Lazaro). Number of specimens studied, 20. Types in the collection California Academy of Sciences; co-sexi- types In my own. rra San Lazaro, Lower California. Ly pe-loc REVISION OF ELEODIINI—BLAISDELL. 81 Salient type-characters—Glabrous and shining; elytra obsoletely subsulcate. Pronotum widest at the middle; disc scarcely more arcu- ately declivous laterally than on the dorsum, distinctly declivous at the apical angles, obsoletely and sparsely punctulate; apical angles subacute with a tendency to be feebly prominent anteriorly. Elytra usually widest behind the middle; dise striato-punctate, punctures more or less eroded, strie somewhat impressed, intervals apparently somewhat feebly convex and obsoletely punctulate. Legs somewhat slender. Diagnostic characters —The salient type characters are sufficient to differentiate peninsularis from the other races of oméssa. It is very interesting to note that it is necessary to use care in recog- nizing it from the peninsular form of inswlaris, which occurs in the same region. In ‘nsularis the pronotum is more quadrate and the apical angles are more prominent anteriorly, and the anterior femora are. feebly armed in both sexes. The first joint of the anterior tarsi of the male is clothed with a brush of golden pubescence beneath—the latter often dark in old specimens. The genital characters are quite different and subgenerically so in the two species. In the peninsular form the elytra are obsoletely striato-punctate as in omissa var. peninsularis. Dr. George Horn, in his report of the Coleoptera of Baja Cali- fornia,” referred the specimens here described, as peninsularis to gen- tilis, and these were confused with the peninsular form of énsularis which were also referred to gentilis. (See Promus insularis. ) Three specimens in the academy’s collection differed from the type series of peninsularis in being intensely black and highly polished. One female was slightly aberrant; serial arrangement of the elytral punctures distinct, strial punctures distinctly larger than the inter- stitial; the pronotum had the sides quite strongly margined, the bead thin and reflexed. . These specimens were destroyed in the great confla- gration of April 18, 1906. GROUP I. QUADRICOLLIS SECTION. ELEODES QUADRICOLLIS Eschscholtz. Eleodes quadricollis Escuscuourz, Zool. Atlas, III, 1833, p. 12, pl. xiv, fig. 5.—MANNERHEIM, Bull. Soe. Nat. Moscow, XVI, 1843, p. 268.—LE Contr, Proc: Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 888.—Casry, Annals N. Y. Acad. Sci., V, Noy., 1890, p. 395.—CHAMPION, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 80. Eleodes tarsalis Casry, Annals N. Y. Acad. Sci., V, Nov. 1890, p. 399. CPRLOCE Cll ACAGs SClew Veet dup 349> 59780—Bull. 63—09-——6 82 BULLETIN 638, UNITED STATES NATIONAL MUSEUM. Oblong-ovate to ovate, moderately elongate, shining, sometimes. subasperulate at the sides of the elytra. Head scarcely twice as wide as long, more or less broadly and feebly impressed laterally, frontal suture usually evident and more or less bisinuate; frons broadly flattened to feebly convex, coarsely punctate, punctures finer and sparser on vertex, more coarsely, densely, and somewhat confluent on the epistoma. Antenne mod- erate in length and comparatively stout, outer four joints very feebly compressed and not dilated, third joint subequal to the combined lengths of the next two, the fourth just noticeably longer than the fifth, the latter, sixth, seventh, and eighth subequal in length, the eighth subtriangular, ninth and tenth suborbicular, eleventh short ovate. Pronotum more or less subquadrate, usually widest before the middle, occasionally at the middle, from one-sixth to one-third wider than long; disc evenly and moderately convex, arcuately declivous laterally, rather coarsely, almost evenly and rather thickly punctate, punctures generally denser at the sides where it is also more or less rugulose on the declivity; apex subtruncate to very feebly emargi- nate, finely and more or less obsoletely beaded; s/des rather strongly arcuate anteriorly, thence straight and obliquely converging to the base as viewed vertically from above, more broadly and evenly arcu- ate in anterior three-fourths, thence more or less feebly sinuate to base as viewed obliquely from the side, finely beaded; base subtrun- rate to feebly rounded, finely margined, one-fifth to one-third wider than the apex; apical angles obtuse and slightly rounded; basal angles obtuse. Propleure usually glabrous, sparsely punctulate, and more or less irregularly rugulose. Elytra oblong-oval to oval, frequently strongly inflated, widest at the middle; base truncate to feebly emarginate; hwmeri obtuse, at times shghtly prominent, narrowly rounded; sédes evenly arcuate, apex obtusely rounded; dsc moderately convex on the dorsum, strongly and quite evenly rounded laterally, inflexed sides broadly and feebly concave before the apex, dorsum arcuately declivous posteriorly, strongly, quite evenly and diffusely punctate to striato- punctate, the punctures generally simple but shghtly asperate lat- erally and on apex. Epipleure moderate in width, gradually narrowing from base to apex, very feebly and broadly sinuate beneath the humeral region; surface finely and sparsely punctate. Sterna quite strongly and densely punctate, rugose. Parapleure strongly and densely punctate. Abdomen glabrous and shining,. rather thickly punctate and rugulose, REVISION OF ELEODIINI—BLAISDELL. 83 Legs moderate, strongly punctate, anterior femora mutic; anterior tibial spurs dissimilar; first joint of the anterior tarsi slightly thick- ened at tip beneath. Male——Oblong-ovate, comparatively narrow, sometimes subcylin- drical.. Antenne reaching to the base of the prothorax. Elytra rather narrowly oblong-oval, gradually narrowed in apical fourth, arcuately and slightly obliquely declivous behind. Abdomen dis- tinetly oblique, very moderately convex and broadly impressed at middle of the first two segments. Femora sometimes slightly thick- ened; anterior tibial spurs rather short, the anterior about one-half longer than the posterior, feebly thickened and gradually narrowed trom base to apex, acute. Anterior tarsi with a tuft of spinules on the thickened tip beneath, the pencil of modified spinules scarcely evident. Female.—Body robust, ovate, antenne not reaching the base of the prothorax. Elytra broadly oval, frequently inflated, moderately nar- rowed at posterior sixth, arcuately and vertically declivous behind. Abdomen horizontal and strongly convex. Anterior spurs of the anterior tibie robust and broad, at least twice as long as the posterior, sides arcuate and parallel, narrowed in about apical fourth and sub- acute. Anterior tarsi with a small transverse tuft of ordinary spi- nules on the thickened tip of the first joint beneath. Measurements—Males: Length, 16.5-21 mm.; width, 6-7.5 mm. Females: Length, 17.5-24 mm.; width, 8-12 mm. Genital characters, male—Edeagophore (Plate 2, fig. 8) fusiform to flattened oblong-ovate, rather stout and arched. . Basale oblong; surface rather strongly convex, sides more or less arcuate. Apicale triangular, slightly elongate; surface somewhat strongly and evenly convex, with a median membranous groove in apical half; sides feebly arcuate in basal half, thence broadly and feebly sinuate to apex, the latter somewhat narrow and acute; base with a small rounded lobe at middle and broadly sinuate laterally. Sternite (Plate 2, fig. 5) transverse. Each lobe slightly quadrate, with the external border quite straight in basal half, thence broadly arcuate to apex, angle narrowly rounded; internal border quite straight; surface moderately convex, sparsely and strongly punctate, setose, setae moderate, longer and denser in apical area. Membrane sparsely setose at bottom of the sinus, the latter broad. Female.—Genital segment (Plate 2, fig. 6) quadrate, not deflexed apically and setose. Valvula—Dorsal plate oblong, slightly narrowing toward apex, explanate externally and at apex; surface nearly plane, impunctate in basal half, elsewhere sparsely punctate and setose, sete rather short and reclining; external border nearly straight, converging 84 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. apically; apical border strongly arcuate on the external lobe, the latter occupying the entire apical margin, with a small sinuation sep- arating it from the minute internal. lobe; internal border nearly straight, at times more or less sinuous. Apical margin clothed with rather long flying soft sete. Appendage not visible beyond the margins of the fossa, short, mammilliform. Basal prominences not evident. Superior pudendal membrane longitudinally rugulose, and reach- ing to opposite the internal apical lobe. Ventrolateral surface (Plate 2, fig. 7) moderately convex basally and feebly concave before the apex, punctuate and setose, sete rather short, long about the fossa; submarginal groove moderate beneath the somewhat explanate external border of the dorsal plate, curving inward beneath the external apical lobe to the fossa. Internal mar- gins of the valves contiguous in basal eighth; fissure rather broadly fusiform, closed in the basal half by the inferior pudendal membrane. Variations in genital characters.—In the most ventricose specimen that I examined the entire apical border was taken up by the exter- nal lobe; the internal lobe was not evident from above; the dorsal plate was narrower than usual. The genital segment viewed ven- trally showed the inferior border of the valvular membrane as form- ing the internal wall of the fossa, and that the internal lobe was therefore small and ventral. Appendage slightly compressed. In a specimen exhibiting marked variation, I found the sete shorter, and the external lobe of the apex more membranous; the sides of the dorsal plate were quite straight and parallel; the internal lobe was ventral as above, and the inferior pudendal membrane was longer than usual. Habitat—California (about the Bay of San Francisco, Esch- scholtz; Mount Diablo, Thomas Casey; Monte Bello, near Mountain View, Santa Clara County, Edward Ehrhorn; San Emigdio Canyen, Kern County, Fordyce Grinnell). Number of specimens studied, 500. Type, location unknown to me. Type-locality—F oothills near San Francisco. Salient type-characters.—Thorax punctulate, subquadrate, broadest anteriorly ; elytra punctato-striate. Body of the male subcylindrical, of the female obovate (Eschscholtz). Diagnostic characters —TYhe strongly developed anterior spurs of the anterior tibize (females) characteristic of the present section, separate quadricollis from all the members of the Carbonaria section. But it will be necessary to actually scrutinize the anterior spurs of the very robust examples of omissa, where the anterior spurs are cer- REVISION OF ELEODITNI—BLAISDELL. 85 tainly strongly developed for that section, and while it will be seen that the spurs are distinctly broadened, they will also be observed to taper more gradually from base to apex, and neyer so parallel nor so suddenly or apically narrowed as in guadricollis. Unworn spurs must always be taken as the criteria. The var. anthracina described below is duller in surface luster, less convex, not inflated, and in typical specimens have the elytra just noticeably obsoletely subsulcato-striate, characters never observed in the true guadricollis; the latter by its quite simple elytral punctua- tion is easily separated from humeralis; in cuneaticollis the elytral sculpturing is distinctly muricato-rugulose. Aéeyi is more robustly ovate. Quadricollis has been up to the present time confused with omissa. Reference to the original description of this species as given by Eschscholtz should quickly close all controversy. That author writes that the elytra are punctato-striate (elytris punctato-striatis) and that this species is plentiful in the foothills about San Francisco (“ Bei St. Franziseo auf den Anhohen hiufig”). These remarks can only be applied to the present species. What has previously been said in regard to the variation of the elytral punctuation in the different species must be recalled at this point. All those specimens with the elytra punctato-striate are to be re- garded as typical. Fully two-thirds of the large series before me have the “ sculpture consisting of punctures sometimes fine, at others rather coarse, rather densely but irregularly placed, and never muri- cate; never arranged in rows” (Horn). Doctor Horn undoubtedly included omissa in the above definition, but then a large percentage of the examples of that species exhibit a serial arrangement of the punctures. After all, the characters apply to guadricollis, for the punctures are sometimes fine, but gen- erally rather coarse; punctato-striate specimens are not common, and never really muricate in the sense as it is applied to humeralis and its races. Quadricollis and omissa as well have a feeble type of sub- muricate punctuation at the elytral sides and apex. I have a male guadricollis at hand, kindly transmitted to me a number of years ago by Colonel Casey. It is not at all inflated, was taken near San Francisco, and is diffusely punctured; bears the label quadricollis in Casey’s handwriting. I consider it a typical specimen, except as regards the punctuation. Tarsalis is only an inflated form (forma tarsalis) of quadricollis. Colonel Casey himself has made this fact known." 4 Coleopterological Notices, V, p. 597, Annals N. Y, Acad. Sci., Dec., 1895. 86 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The student will at times find it very difficult to decide whether a given specimen (especially males) shall be referred to guadricollis or Omissa. In my experience the epistoma in quadricollis is always more coarsely punctate than in omissa. Specific differentiation is no more difficult in this instance than is met with in other sections of the genus Eleodes. The extreme forms or heterotypes of carbonaria and obsoleta are as difficult of separation as are the above, in fact, all the constituents of the present section of the subgenus J/elaneleodes might be consid- ered as races and forms of a single species if viewed from a broad evolutionary standpoint. The mentum is usually triangulo-trapezoidal, strongly punctate and foveate laterally within the more or less strongly defined margins; the surface is longitudinally ridged at the middle. The prosternum is more or less strongly protuberant ventrally with the coxee, and more or less longitudinally convex between the same, usually grooved at the middle and rather distinctly margined around the acetabula; frequently submucronately produced behind and at times nearly vertically subtruncate. The mesosternum more or less concave and arcuately declivous. The intercoxal process of the abdomen is quadrate (male) to feebly transverse (female) and about a third of its width wider than the metasternal salient. The metasternum laterally between the coxz is as long as the width of a mesofemur at the base. The post-coxal portion of the abdomen is equal in length to that of the process, and also to that of the third segment. In the male the second segment is about equal to the width of the process, and the third is a half longer than the fourth. In the female the second is twice as long as the fourth segment. The profemora are usually moderately clavate in the male and less so in the female. The grooves are limited by rather strong margins, which are more or less asperulate and converge near the base. The meso- and metafemora have their superior and inferior sur- faces quite parallel. The margins of the grooves are not strongly marked, are finely muricate, and become evanescent before becoming contiguous. The tibize are frequently more or less arcuate and without tarsal grooves; the articular cavities are closed. The tarsi are somewhat variable as regards to stoutness, fre- quently rather more slender in the female. The protarsi are usually slightly thicker in the male and apparently subequal in the sexes. The protarsi are about two-fifths of their length shorter than a mesotarsus, Joints two, three, and four are subequal in the male; REVISION OF ELEODIINI—BLAISDELL. Oi the fourth a little smaller than the preceding in the female; the fifth is quite equal to the combined lengths of the preceding three; the first about a half longer than wide. The mesotarsi are about two-sevenths (male) or a little less (fe- male) of their length shorter than a metatarsus. Joints two to four are just a little progressively shorter or subequal, and together a little longer than the fifth, which is quite equal to the first. A metatarsus is about five-ninths of its length shorter than a meta- tibia. Joint three is just noticeably shorter than the second, both together equal to the length of the first, the fourth a httle shorter. ELEODES QUADRICOLLIS var. ANTHRACINA, new. Oblong-ovate to ovate, surface subopaque, elytra usually obsoletely subsuleate; antenne reaching beyond the base of the prothorax in the male. Head somewhat coarsely punctate, the coarser punctures on the epistoma not confluent nor crowded. Antenne rather long, shghtly stout, feebly clavate, outer four joints slightly compressed and feebly dilated, third joint as long as the next two combined, fourth a little longer than the fifth, the latter, sixth, and seventh subequal, the eighth shorter and subtriangular, ninth and tenth suborbicular, eleventh ovate to oval. Pronotum subquadrate, usually widest at about the middle, near a sixth wider than long; disc evenly and moderately convex, arcuately declivous laterally, finely and rather sparsely punctulate, scarcely coarser or denser laterally, where it is frequently slightly rugulose ; apex subtruncate in circular arc, and finely or obsoletely margined ; sides evenly and moderately arcuate in the anterior three-fourths. thence converging to base, frequently rather evenly arcuate from apex to base (viewed vertically from above), margin finely beaded; base slightly rounded, often feebly sinuate at the middle, finely and more or less obsoletely beaded, about one-fourth wider than the apex and equal to the length; apical angles distinct and very narrowly rounded ; basal angles obtuse. Elytra oval, widest at the middle, less than twice as long as wide; base more or less emarginate, frequently slightly sinuate laterally, usually a little wider than the contiguous prothoracic base; humeri obtuse, narrowly rounded and searcely at all prominent anteriorly ; disc moderately convex on the dorsum, more strongly and almost evenly rounded laterally, arcuately declivous posteriorly, surface usually striato-punctate, intervals apparently very feebly convex, giving an obsoletely and faintly subsulcate appearance, strial punc- tures somewhat small, series moderately distant, rarely impressed, interstitial punctures more or less irregular on the dorsum, quite equal in size to the strial; frequently the punctures of both series are 88 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. arranged without trace of order, nearly always so laterally and on the apical declivity, where they are scarcely ever subasperate. Otherwise as in guadricollis. Male.—Antenne reaching a short distance beyond the base of the prothorax. Pronotum usually widest at the middle. Female.—Anienne reaching to the base of the prothorax. Prono- tum not usually very noticeably widest before the middle. Otherwise the sexual characters are as in quadricollis. Measurements—M ales: Length, 18-19.5 mm.; width, 6.5-7.5 mm. Females: Length, 18-22 mm.; width, 7.5-10.5 mm. Genital characters, male.—Edeagophore flaxseed-shaped and mod- erately arched. Basale oblong, quite evenly convex above; sides shghtly arcuate. Apicale triangular and moderately elongate; surface moderately and evenly convex, with a median semimembraneous groove in apical half; sides nearly straight, apex subacute; base rather broadly and arcuately lobed at the middle, and slightly sinuate laterally. Sternite parabolic in outline and shghtly transverse. Each lobe with the external border quite evenly arcuate to apex, the latter nar- rowly rounded; internal border short and nearly straight; surface feebly convex, impunctate in basal half, sparsely punctate apically, setose, sete moderate in length, longer about the apex, not dense. Membrane sparsely setose across the bottom of the sinus, the latter: short. Female.—Genital segment quite quadrate, explanate externally in apical two-thirds. Valvula.—Dorsal plate oblong, moderate in width, sides subpar- allel; surface quite plane, glabrous, coarsely and sparsely punctate, each puncture with a moderately long reclining seta, impunctate at base; external border more or less straight to feebly arcuate to the apical border, which is strongly rounded on the external lobe and shehtly sinuate just external to the minute internal lobe; external lobe about as wide as the dorsal plate, both lobes sparsely set with long sete; internal margin more or less sinuous. Appendage not visible from above, short mammilliform and scarcely projecting beyond the margins of the fossa. Basal prominences scarcely evident. Superior pudendal membrane longitudinally rugulose, reaching to the internal lobe. Ventrolateral surface as in quadricollis. Habitat—Arizona (Fort Grant, July 10; Hot Springs, June; Oracle, San Simon, Galiuro Mountains, May; Catalina Springs, April; Tueson, Hubbard and Schwarz; Nogales, September, Albert Koebele and F. W. Nunenmacher). REVISION OF ELEODIINI—BLAISDELL. 89 Number of specimens studied, 150. Types (Cat. No. 12202) are in the U. S. National Museum collec- tion; cotypes in my own. Type-locality.—F ort Grant, Arizona. Salient type-characters—Integuments dull in luster. Epistoma somewhat coarsely punctate, punctures not crowded nor confluent. Antenne in the male reaching beyond, and in the female to the pro- thoracic base. Thorax in both sexes widest at about the middle and ° not more strongly arcuate before the same. Elytra striato-punctate, obsoletely and faintly subsulcate. Diagnostic characters——Anthracina differs from quadricollis im its salient type-characters. For its separation from carbonaria, humeralis, and cuneaticollis, see differential diagnosis under guad- ricollis. I have seen specimens of this race labeled carbonaria, and it is generally labeled guadricollis in collections. In its dull luster and long antenne it might be confused with ampla, but the shorter legs and large anterior spurs of the anterior tibiz in the female will separate it. Mr. Blanchard has compared it with the LeConte types, especially vicina, and says that it differs from all and is most closely related to gquadricollis. Prosternum as in guadricollis, but less inclined to be submucronate and less frequently grooved at the middle between the coxx. The mesosternum is frequently shghtly more strongly concave. Otherwise as in quadricollis. ELEODES QUADRICOLLIS var. LUSTRANS, new. Oblong-ovate to ovate, moderately elongate, very glabrous and shining. Head comparatively rather small, frons moderately and quite evenly convex, scarcely to distinctly impressed laterally, frontal suture usually not evident, almost coarsely, rather thickly and evenly punctate, punctures usually not denser on the epistoma. Antenna rather long, somewhat slender, outer four joints scarcely at all com- pressed or dilated, third joint quite equal in length to the next two taken together, the fourth just visibly longer than the fifth, the latter, sixth and seventh subequal, eighth just the least shorter and subtri- angular, ninth and tenth suborbicular, eleventh short ovate or oval. Pronotum subquadrate, widest at or just a little in front of the middle, one-fourth to one-third wider than long; disc evenly and moderately convex, more or less arcuately declivous at the sides and apical angles, finely and sparsely punctate, sometimes a little more 90 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. coarsely so laterally; apex truncate in circular arc, finely and more or less obsoletely beaded; sides evenly and not strongly arcuate in anterior three-fourths, thence convergent to the base, or more or less arcuate from apex to base, as viewed vertically from above, more broadly arcuate and just the least sinuate before the base as viewed obliquely from the side, rather distinctly and finely beaded; base feebly rounded and more or less finely margined, a seventh to a fifth wider than the apex; apical angles obtuse and not in the least promi- nent anteriorly; basal angles obtuse. Elytra oval, widest at the middle, scarcely twice to one-fourth longer than wide; base truncate to feebly emarginate, scarcely to a little wider than the contiguous prothoracic base; humeri obtuse ; sides evenly arcuate, apex not very narrowly rounded; disc slightly flattened and moderately convex on the dorsum, rather evenly and not very broadly rounded at the sides, arcuately declivous posteriorly, punctate, punctures rather fine and more or less irregular, usually with a serial arrangement on the dorsum, interstitial punctures more or less irregular, those of both series equal in size or the strial are shghtly larger; both series always confused at the sides and on the apex; not at all submuricate. Legs rather slender. Otherwise as in quadricollis. Male somewhat depressed. Female.—Convex and comparatively less elongate than in an- thracina. Other secondary sexual characters as in quadricollis. Measurements.—M ales: Length, 16-17 mm.; width, 6-7 mm. /e- males: Length, 16.5-17 mm.; width, 8-8.2 mm. Genital characters, male—Sternite with each lobe more or less parabolic in outline. Otherwise as in guadricollis. Female—As in quadricollis, except that the ventrolateral surface is strongly convex at basal fourth and broadly concave before the apex, sete rather long internally. Habitat—Arizona (Chiricahua Mountains, June, Hubbard and Schwarz; Catalina Springs, April; Tucson.) Number of specimens studied, 9. Types (Cat. No. 12203) in the U. S. National Museum collection. Ty pe-locality Chiricahua Mountains, Arizona. Salient type-characters.—Glabrous and shining. Antenne reaching beyond (male) or to (female) the base of the prothorax. Body in the female less elongate than in guadricollis. Punctures on the epis- toma not coarser than on the frons. Pronotum widest at the middle; ise finely and sparsely punctate, punctures not coarser nor denser laterally. Elytra with the discal punctures arranged in series on the oc a iit i a A i i te le REVISION OF ELEODIINI—BLAISDELL. 9] dorsum, laterally confused and not submuricate; punctures of both series fine and equal in size. Diagnostic characters —Ditters from both quadricollis and its race anthracina by the salient type characters. In both sexes the form is smaller and less stout and the female is decidedly less elongate and more robustly ovate. The punctuation is noticeably finer in all the specimens that I have seen, the epistoma is more sparsely and finely punctate, the punctures being not at all confluent. A striking feature in the sculpturing is the very smooth integuments. The large anterior tibial spurs separate it from all the constituents of the Carbonaria section. Mr. Blanchard, after careful comparisons made with the LeConte material, writes: “ I do not recognize this as belonging to any of the named forms; it is nearest to qguadricollis.” The prosternum is more often decidedly mucronate, although not strongly produced. Otherwise as in quadricollis. ELEODES CUNEATICOLLIS Casey. Eleodes cuneaticollis Casry, Annals N. Y. Acad. Sci., V. Nov. 1890, p. 397. Oblong-ovate to ovate, elongate, rather shining; elytra coarsely rugulose. Head twice as wide as long, feebly convex or flattened between the eyes, coarsely and strongly, irregularly and more or less confluently punctate, punctures finer on the vertex. Antennw moderate in length and somewhat stout, scarcely compressed and not dilated, third joint fully as long as the next two combined, the fourth but slightly longer than the fifth, the latter, sixth, seventh, and eighth subequal, ninth suborbicular, tenth transverse, eleventh short ovate. Pronotum widest at the junction of the anterior and middle thirds, one-third to two-fifths wider than long; disc broadly and moderately convex, more or less declivous laterally and at the apical angles, rather sparsely, coarsely, and deeply punctate, the punctures about twice as large and distinct as those of humeralis, frequently more or less rugu- lose at the sides; apex truncate or broadly and feebly emarginate in circular arc, very finely and more or less obsoletely beaded; sides strongly arcuate anteriorly, conspicuously convergent and almost straight in basal half, viewed vertically from above, or more broadly arcuate in anterior three-fourths, thence convergent and more or less sinuate to the base, as viewed obliquely from the side, finely beaded ; base truncate to slightly rounded and finely margined, about equal to the length and one-tenth to one-fifth wider than the apex: apical angles obtuse and more or less narrowly rounded; basal angles obtuse. 92 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Propleure sparsely punctate and more or less strongly rugulose. Elytra oval, widest at the middle, less than twice as long as wide; Jase truncate to feebly emarginate, sometimes sinuate laterally, equal to or slightly wider than the contiguous prothoracic base; humeri obtuse, at times more or less rounded, again slightly prominent; s¢des evenly arcuate, somewhat pointed at the apex, the latter not broadly rounded; disc more or less moderately convex on the dorsum, quite strongly and evenly rounded laterally, arcuately declivous posteriorly ; inflexed sides somewhat oblique; very coarsely, deeply, and densely punctate, punctures irregularly distributed without trace of a serial arrangement, not muricate, but producing a strongly rugu- lose appearance by mutual semi-coalescence. HE pipleurw moderate in width and gradually narrowing from base to apex, superior margin very broadly and feebly sinuate beneath the humeri; surface sparsely punctate. Sterna somewhat shining, densely punctate and more or less rugose. Parapleure coarsely and densely punctate. Abdomen glabrous, sparsely punctate, punctures coarser and denser on the fifth segment; surface more or less rugulose. Legs short and rather robust. Anterior femora mutic; anterior tibial spurs dissimilar in the sexes; first joint of the anterior tarsi thickened at tip beneath, feebly dissimilar in the sexes. Male.—More or less slender. Antenne reaching to the base of the prothorax. Elytra arcuately and more or less obliquely declivous behind and gradually narrowed in the posterior fourth. Abdomen more or less distinetly oblique, moderately convex, broadly im- pressed on first segment, less so on the second. Anterior spurs of the anterior tibiew slender, acute, and about a third longer than the posterior. First joint of the anterior tarsi with the tuft of spinules obsolete at tip beneath, groove slight and not interrupted on the thickened tip. Female.—Ovate, robust, antenne not quite reaching to the pro- thoracic base. Elytra broadly oval, arcuately and more or less ver- tically declivous and less strongly narrowed behind. Abdomen hori- zontal and more or less strongly convex. Anterior spurs of the anterior tibiz moderately broadened, sides more or less parallel, narrowing in apical fourth, rather acute or blunt if worn. First joint of the anterior tarsi distinctly thickened at tip beneath, tuft of spinules not evident. Measurements.—Males: Length, 14-17.5 mm.; width, 5.5-6.8 mm. Females: Length, 15-18.8 mm.; width, 7-9.2 mm. Genital characters, male-—Edeagophore elongate and acutely ovate, and rather strongly arched. REVISION OF ELEODIINI—BLAISDELL. 93 Basale rather short, suboblong oval; sides feebly arcuate; surface evenly convex. Apicale elongately triangular and rather arched; surface evenly convex, with a rather narrow longitudinal groove in apical two- thirds; sides nearly straight, apex acute; base with a rather small rounded lobe at middle and feebly sinuate laterally. Sternite short and transversely parabolic in outline, each lobe somewhat transverse with the external border broadly arcuate; apex shghtly prominent and rather narrowly rounded; internal border rather straight: surface feebly convex, strongly and sparsely punctate, setee rather long. Membrane very sparsely setose across the bottom of the sinus, the latter short. Female.—Genital segment quadrate, valves with their external borders feebly converging, setose. Valvula (Plate 3, fig. 18).—Dorsal plate oblong, feebly explanate externally and nearly plane; surface glabrous, finely and very sparsely punctate, sete short and reclining; external border more or less feebly arcuate; internal border straight or sinuous; apical margin moderately arcuate in external three-fourths on the external lobe; internal lobe not prominent and separated from the external by a feeble sinuation. Appendage sometimes visible from above, rather short mammilli- form, with a few short sete at tip. Fossa large and quite terminal, the inferior margin of the valvular membrane forming the internal margin, the latter more prominent than the internal lobe above. Basal prominences not in the least evident. Superior pudendal membrane longitudinally rugulose, long, reach- ing to the internal apical lobe. Ventrolateral surface moderately convex, not prominent at the base and not transversely concave before the apex; submarginal groove very broad and shallow, forming a lateral concavity before the apex. Habitat—California (San Francisco, Marin, Alameda, and San Mateo counties, August). Specimens have been distributed as having been collected in San Diego County, but all of those were undoubtedly collected by Mr. G. W. Dunn and sent out without true locality labels—at least that is the case with a series that I have had for years. Personally I have collected them only in the above-named localities and have no authentic data of their having been taken elsewhere. Colonel Casey’s type specimen was without an exact locality label. Number of specimens studied, 30. Type in Colonel Casey’s collection. Lype-locality—California; probably from some place about the Bay of San Francisco, 94 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Salient type-characters.—Head rather large, coarsely and deeply punctate. Antenne short and robust. Prothorax with apex nearly as wide as the base; sides strongly arcuate anteriorly, conspicuously convergent and almost perfectly straight in basal two-thirds; disc widest at apical third, rather sparsely, coarsely and deeply punctate, punctures about twice as large and distant as those of humeralis. Elytra distinctly less than twice as long as the prothorax; disc rather depressed above, very coarsely, deeply, and densely punctate, the punctures irregularly arranged without trace of impressed strie, not muricate but producing a strongly rugulose appearance by mutual semi-coalescence. Legs short and robust (Casey). Diagnostic characters —Cuneaticollis differs from all the other species of the present subgenus in having the elytra strongly rugulose from semi-coalescence of the coarse punctures. As Casey writes, it belongs near humeralis, but differs in four important characters, namely: * The much shorter and robust antenne and legs, the very much coarser and deeper elytral sculpture, and coarser, sparser pro- notal punctures, and finally the smaller and much less unequal spurs of the anterior tibie in the male, the larger spur in humeralis being nearly four times as long as the smaller one.” In the largest females before me the humeri are exposed. The most noticeable disparity in the elytral characters of Casey’s type and the series before me, is that, while in the type the elytra are distinctly less than twice as long as the prothorax, in all the examples before me the elytra are distinctly a little more than twice as long as the same. My series agrees in all other particulars with the original description. I have not seen any other species that could be referred to cuneaticollis; Casey does not mention the number of specimens he had before him at the time of writing his description, but specimens do occasionally have unusually short elytra in other species. In some examples the elytra are vaguely substriate when viewed longitudinally. The individual specimens of cuneaticollis exhibit considerable variability, more so than in the other related species. The largest female before me has abnormally short legs, another has the sides of the prothorax unusually and strongly rounded in the anterior third; the antenne are noticeably more robust and shorter in some specimens than others. I beheve the present species to be more closely related to quadricollis than to humeralis; the anterior tibial spurs suggest this relationship. The mentum is subparabolic to triangulo-parabolic, not coarsely but densely punctate, and not noticeably setose; surface more or less foveate laterally and broadly convex centrally. The prosternum and mesosternum are variable, as in humeralis. REVISION OF ELEODIINI—BLAISDELL. 95 The intercoxal process of the abdomen is subquadrate (male) or shghtly transverse (female) and about one-fourth of its width wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a mesotibia at the extreme apex. The post-coxal part of the first abdominal segment is equal in length to the process, and the latter to that of the third segment. In the sexes the second segment is twice as long as the fourth. The profemora are more or less distinctly clavate; the grooves are moderately broad and the margins subeariniform, more or less asperu- late, becoming contiguous and evanescent at basal third. The mesofemora are scarcely or very feebly clavate, grooves quite plane, margins feeble and asperulate, evanescent at middle before becoming contiguous. The metafemora are not at all widened externally and the superior and inferior surface lines are quite parallel; grooves as on the meso- femora. The tibize may be more or less feebly arcuate, tarsal grooves absent, articular cavities closed. The pro- and mesotibiz are somewhat inwardly produced at apex. The protibiz are sometimes quite distinctly carinate externally in basal half, noticeably so in the largest female before me. Tarsi variable in stoutness. The protarsi are about one-half (male) or one-fourth (female) of their length shorter than a mesotarsus. Joints two, three, and four subequal in size, and together a little longer than the fifth; the first shghtly shorter than the third and fourth taken together. The mesotarsi are about a ninth (male) or a fifth (female) of their length shorter than a metatarsus. Joints two, three, and four feebly decreasing in length in the order named; the fifth subequal to the combined lengths of the second and third, the first to the third and fourth. A metatarsus is about a half (male) or a third (female) of its length shorter than its metatibia. Joints two and three are subequal, and together about equal to the fourth, which is subequal to the first. ELEODES HUMERALIS LeConte. Eleodes humeralis LEContTE, Reports of Explor. and Surveys .. . 47th and 49th Parallel, XII, Appendix No. 1, 1857, p. 50; Proc. Acad. Nat. Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 809.—CHAMPIoN, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 80. Oblong-ovate to ovate, elongate, more or less opaque; elytra densely, rather finely and muricately, or granulato-muricately pune- tate. 96 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Head rather less than twice as wide as long, frequently feebly and vather broadly impressed laterally, frontal suture obsolete or feebly evident and scarcely sinuate; frons feebly convex, usually more or less flattened, densely and finely, sometimes a little coarser and more or less confluently punctate, punctures fine on the vertex increasing in coarseness to the epistoma. Antenne long and somewhat slender, feebly compressed and scarcely dilated in the outer four joints, third joint as long as the next two taken together, fourth shghtly longer than the fifth, the latter, sixth, and seventh subequal, eighth a little shorter and more or less subtriangular, ninth and tenth suborbicular, eleventh short ovate. Pronotum subquadrate, widest at or just in front of the middle, about one-third wider than long; disc moderately and quite evenly convex, feebly and rather arcuately declivous laterally, rather finely and densely punctate, usually more or less narrowly subgranulate along the sides; apex truncate or very feebly emarginate in circular are, usually finely beaded; sides rather broadly and moderately arc- uate in the anterior two-thirds, thence to the base convergent and straight or feebly sinuate, marginal bead rather thin, finely reflexed and visible throughout its entire length from above, base truncate or feebly rounded, finely margined, and about one-sixth wider than the apex, about equal to the length; apical angles obtuse, scarcely rounded; basal angles obtuse, not rounded, almost rectangular at times. Propleure quite densely punctato-rugulose. Elytra oval, not twice as long as wide, usually widest at the middle; base truncate or more or less feebly emarginate, as wide as or sheghtly wider than the contiguous prothoracic base; hwmeri obtuse or shghtly prominent; sé¢des evenly arcuate, apex somewhat narrowly rounded; disc more or less slightly depressed on the dorsum, some- times slightly ogival, arcuately declivous posteriorly, and quite strongly but not broadly rounded laterally, very densely and rather finely muricately punctate or almost granulate, punctures more simple centrally along the suture, the latter frequently impressed ; surface at times is obsoletely striate, the inflexed sides are inwardly declivous and with the surface lines somewhat straight. Epipleurw somewhat narrow, gradually narrowing from base to apex, the superior margin is broadly and rather feebly sinuate be- neath the humeri; surface densely and subasperulately punctate. Sterna more or less shining and very densely punctato-rugulose. Parapleure densely punctate. Abdomen more or less shining, rather densely punctato-rugulose. The first segment is more strongly sculptured, and the fifth quite densely punctate, REVISION OF ELEODIINI—BLAISDELL. 97 Legs moderate in length, comparatively somewhat slender. Iemora mutic and not thickened; anterior tibial spurs dissimilar in the sexes; anterior tarsi with the first joint shghtly thickened at tip beneath, groove interrupted at the ventral apical margin, tuft of spinules feebly developed and slightly dissimilar in the sexes. Male-——Oblong-ovate to ovate, not slender. Antennx reaching a short distance beyond the prothoracic base. Abdomen feebly oblique, ‘moderately convex, more or less impressed at middle on segments one and two. Anterior spurs of the anterior tibie usually about three times longer than the posterior, shghtly broadened and slender, acute, gradually narrowing from base to apex. Female.—Ovate, robust. Antenne reaching at least to the protho- racic base. Elytra arcuately and vertically declivous behind. Abdo- men horizontal, strongly convex. Anterior spurs of the anterior tibiee much broadened and produced, about three times as long as the posterior, sides subparallel and narrowed in apical fourth, subacute, worn spurs are obtusely rounded at tip. In the material before me three forms may be recognized: Forma typica—Elytra generally flattened on the dorsum, sides of the disc are not broadly rounded, and the inflexed portions are ob- liquely and inwardly declivous. Elytral dise very densely and rather finely muricately punctate. Forma granulato-muricata——KElytra generally moderately convex on the dorsum, sides more or less broadly rounded, dise moderately densely granulato-muricate, granules fine and shining. Forma tuberculo-muricata.—Pronotum more arcuately declivous laterally, with the margins more or less invisible from above. Elytra more coarsely and strongly sculptured, subtuberculately muricate, the punctures much sparser than in the typical form. Measurements.—M ales: Length, 14.2-17.5 mm.; width, 6.2-7 mm. Females: Length, 16-20.2 mm.; width, 7.6-8.5 mm. Genital characters, male-—KEdeagophore elongately ovate, pointed and slightly arched. Basale oblong, evenly convex above; sides feebly arcuate. Apicale triangular, twice as long as wide, evenly convex above, with a rather wide median membranous groove in apical half; sides straight and convergent to the subacute tip; base with a rounded lobe at middle, feebly sinuate laterally. Sternite subparabolic, almost as wide as long. Each lobe with the outer border more or less straight in basal two-thirds, thence obliquely subtruncate to apex, tip narrowly rounded; internal margin feebly arcuate; surface scarcely convex, glabrous, very sparsely punctate in apical half and setose, sete moderately long, longer on apical margin and rather sparse. Membrane with scattered setx across the bottom of the sinus, the latter quite deep. 59780—Bull. 68—09——7 98 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Female—Genital segment trapezoidal, valves rather distant and convergent apically, setose; superior pudendal membrane rather widely exposed. Valvula (Plate 3, fig. 17)—Dorsal plate oblong, feebly explanate externally, moderately narrow; surface plane, finely and sparsely punctate, also setose in apical half, sete fine and short; borders parallel, straight or feebly sinuous; apical margin feebly arcuate in outer three-fifths on the external apical lobe, the latter slightly defined from the internal lobe by a slight sinuation; internal lobe not prominent. Apical parts set with moderately long sete. Appendage not visible from above, short mammilliform, not ex- tending beyond the margins of the fossa, the latter rather large. Superior pudendal membrane longitudinally rugulose and reaching to the internal lobe. Ventrolateral surface rather evenly convex, not protuberant, sur- face lines quite straight when viewed longitudinally, not transversely concave before the apex; submarginal groove broad and _ shallow, forming a lateral concavity that does not bend inward beneath the apex; surface finely and sparsely punctate and setose, sete very short. Internal margins of the valves contiguous in basal eighth; fissure rather broadly fusiform, open apically, closed in basal half by the inferior pudendal membrane. In the granulato-muricata form the genital segment of the female is more quadrate, valves quite parallel. The dorsal plate is explanate externally in apical two-thirds. Otherwise as in the typical form. Forma typica—//abitat.——Washington (Walla Walla; Yakima, C. V. Piper) ; Oregon (George Horn, G. W. Dunn) ; California (north- ern, George Horn) ; Idaho (Hubbard and Schwarz). Forma granulato-muricata.—Nevada (Verdi, April, F. E. Blais- dell; Carson City, July; Reno, H. F. Wickham) ; Utah (C. V. Riley) ; California (Lassen County). Forma tuberculo-muricata—Colorado (C. V. Riley; South Park, elevation, 8,000-10,000 feet, Bowditch; Buena Vista, Wickham’s List). Number of specimens studied, 105. Type in the LeConte collection. — Type-locality—* Wenass River to Fort Colville.” Salient type-characters—Opaque, head and thorax densely punc- tate. Thorax subquadrate, narrowed behind, sides rounded, base truncate, posterior angles obtuse. Elytra depressed on the dorsum, obsoletely striate, sculptured with subacute granules that are more or less thickly and irregularly placed, punctate on each side of the suture. Prosternum perpendicular behind (LeConte). Diagnostic characters.—Humeralis exhibits a remarkable develop- ment in the anterior tibial spurs. In both sexes the anterior spur is REVISION OF ELEODIINI—BLAISDELL. 99 about three or four times longer than the posterior. In this it differs from all of the other species of the subgenus, except ri/eyi. The integuments are more or less opaque, and in the typical form from Washington the elytra are densely muricately punctate and more or less flattened on the dorsum. Specimens from Lassen County, California, have the elytra dis- cretely granulate; each granule under low power of magnification is polished and shining, under high power minutely pointed. These constitute the heterotypical western form; in the Nevada specimens the granules are smaller, less shining, but discrete. In the Colorado examples the granules are much larger and sparser, and characterizes the incipient race here designated as forma tuberculo-muricata. The Idaho specimens are feebly shining and more convex and resemble the species described as schwarzii. The form of the anterior tibial spurs will separate the two. These specimens and those from Colorado have the sides of the pronotum more strongly declivous, so that the marginal head ean not be seen from above when viewed vertically. The relationships of these forms are really dubious, but the material at hand does not warrant their separation as distinct species. I have seen specimens of the granulato-muricata form labeled granulata; the latter species belongs to another subgenus and_ is different altogether. Rileyi is smaller, more sparsely punctate, more convex in form, and undoubtedly more closely related to Awmeralis than to any other species of the subgenus. The mentum is of moderate size, rather parabolic in outline in the typical form, and more triangulo-trapezoidal in the other forms; the surface is coarsely but not very densely punctate, moderately convex and scarcely foveate laterally; sete very short. The prosternum is usually quite semicircularly convex antero- posteriorly between the coxxe; sometimes feebly submucronate or rarely vertically subtruncate behind. The mesosternum is more or less feebly oblique and concave. The intercoxal process of the abdomen is slightly transverse in both sexes, and about one-third (male) or a sixth (female) of its width wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a metatibia at apex. The post-coxal portion of the first abdominal segment is equal in length to that of the process, and in the female to the length of the third segment. 100 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. - In the male the second segment is a little longer than the post-coxal part of the first and twice as long as the fourth. The third is about one-half longer than the fourth. In the female the second segment is about two and one-half times longer than the fourth. The profemora are rather feebly and evenly tumid, scarcely clavate ; the grooves have well-defined subasperulate margins that become contiguous at basal fourth. | The meso- and metafemora are scarcely at all widened externally. The superior and inferior surface lines are quite parallel. The tibial grooves of the mesofemora are quite well defined by asperulate mar- gins that become evanescent before becoming contiguous. On the metafemora the grooves are similar, but evanescent at middle. In the typical form the tibiez are rather slender, and stouter in the other forms. The tarsal grooves are obsolete and the articular cavi- ties closed. The tibie are usually not at all arcuate. The tarsi are moderate in length and stoutness. The protarsi in the male are apparently a httle longer than in the female; about two-sevenths of their length shorter than a mesotarsus. Joints two, three, and four are subequal, and together about equal to the fifth; the first is scarcely as long as the second and third taken together. The mesotarsi are about a fifth of their length shorter than a meta- tarsus. Joints two, three, and four slightly decrease in length in the order named, and together a little longer than the fifth; the first is quite equal to the combined lengths of the second and third. The metatarsi are about five-eighths of their length shorter than a metatibia. Joints two and three are subequal, and together equal to the first, the fourth being shghtly shorter. ELEODES RILEYI Casey. Eleodes rileyi Casry, Annals N. Y. Acad. Sci., VI, Nov., 1891, p. 57. Rather robust and convex, somewhat strongly shining, intense black throughout. /7Zead scarcely twice as wide as long, feebly convex, rather broadly and more or less impressed laterally, frontal suture usually distinet, bisinuate, and more or less impressed. coarsely and rather closely punctate; punctures sparser on the vertex, rather coarser on the epistoma. Antenne subequal in length to the head and prothorax, the third joint three times as long as wide and rather longer than the next two combined; the outer joints but moderately robust. Pronotum from one-fourth to three-fifths wider than long, and widest a little before the middle; disc moderately and evenly convex, arcuately declivous at the sides, rather coarsely, deeply punctate, sit ae ane Se a ee ~~. REVISION OF ELEODIINI—BLAISDELL. TOW densely so toward the sides, sparsely and unevenly, with large im- punctate patches toward the middle; apex broadly and feebly emar- ginate, more or less obsoletely and finely margined; sédes strongly arcuate at apical third, thence strongly convergent and nearly straight to the apex; distinctly sinuate before the base, finely and distinctly beaded; base transverse, generally very broadly, feebly sinuate toward the middle, two-ninths to one-fourth wider than the apex, rather coarsely margined laterally, more or less obsoletely so at middle; apical angles obtuse, but not at all rounded; basal angles sheghtly obtuse, not much rounded and rather prominent. Propleure sparsely, irregularly punctate and more or less rugulose. Elytra two and one-half times longer than the prothorax, and, in the middle, from one-fifth to nearly one-half wider than the fatter, widest at the middle; dase truncate or feebly emarginate; humeri narrowly rounded, slightly prominent, and usually distinctly exposed ; sides evenly arcuate, apex narrowly rounded; disc moderately convex on the dorsum, rather evenly rounded laterally, and arcuately decliv- ous behind, rather coarsely, feebly and subasperately punctate, some- times with very feebly impressed distant lines, the punctures rather sparse, not much denser laterally, confusedly disposed, but often hav- ing a feeble neal arrangement. Epipleure moderate in width, gradually narrowing from base to apex, superior margin feebly and broadly sinuate beneath the humeri; surface finely, sparsely, and rather obsoletely punctate. Sterna densely punctate and rugulose. Parapleure densely punctate. Abdomen more or less shining, sparsely punctate and rugulose; first segment densely and strongly sculptured, and the fifth more coarsely and densely punctate. Legs rather short, anterior femora mutic; anterior tibial spurs extremely unequal, the anterior spur robust, long, subparallel, and the posterior spur very small and acute. Anterior tarsi with the first joint feebly thickened at tip beneath, with tuft of ordinary spinules more or less feebly developed. Male——Oblong-ovate, not noticeably slender. Elytra arcuately and somewhat obliquely declivous behind. Abdomen distinctly oblique and moderately convex, more or less broadly impressed on first two segments. Anterior spur of the anterior tibie about one- half as wide as that of the female, and somewhat gradually narrowed in apical half, and acute. Female.—Robust, ovate. Elytra arcuately and vertically declivous behind. Abdomen horizontal and rather strongly convex. Anterior spur of the anterior tibize much broadened, and narrowed in apical fourth, and subacute. Measurements —Length, 12-15 mm.; width, 5.2-7.2 mm. 102 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Genital characters, male. feebly arched. Edeagophore elongately ovate, very Basale oblong, about three times longer than wide; surface evenly and moderately convex; sides feebly arcuate. Apicale triangular; surface evenly and moderately convex, groove semi-membranous and rather broad, extending from near tip to base; sides arcuate in basal third, thence broadly and moderately sinuate to apex, the latter narrowly rounded; base subtruncate. Sternite subparabolic. Each lobe with the apex somewhat ex- trorsely terminal; external border rather straight at basal third, thence arcuate to apex, the latter somewhat rounded; internal border arcuate; surface more or less convex, punctate and setose in apical half, setee moderate in length and sparse. Membrane sparsely setose across the bottom of the sinus, the latter deep. Female.—Genital segment moderate in size, more or Jess trapezoi- dal, valves more or less reflexed externally, somewhat feebly deflexed at. apex, setose; external border rather strongly explanate, often less so toward apex. Valvula.—Dorsal plate oblong, sides slightly converging apically ; surface rather plane, glabrous, sparsely punctate and setose, sete longer in apical moiety; external border usually feebly and broadly sinuate from the rather prominent basal angle; apical margin not distinct from the external lobe, the latter broadly and not very evenly arcuate, very feebly defined from the internal lobe by a minute sinu- ation; apex clothed with rather long sete; internal border more or less sinuous; internal lobe small. Appendage minute within the transverse fossa, Superior pudendal membrane longitudinally rugulose and not quite reaching the internal apical lobe. Ventrolateral surface moderately convex in basal two-thirds, thence broadly and feebly concave to apex; submarginal groove broad be- neath the explanate external border of the dorsal plate and becoming continuous with the general concavity; surface sparsely punctate and finely setose, setae longer on apex. Internal margins of the valves contiguous at basal fifth, subcontiguous apically; fissure rather broadly fusiform, and closed in basal half by the inferior pudendal membrane. Habitat—Arizona (Thomas Casey) ; Idaho (?). Number of specimens studied, 2 (male and female). Type in Colonel Casey’s collection. Ty pe-locality.— Arizona. Salient type-characters.—Rather robust. Head coarsely and rather closely punctate. Prothorax with apex broadly, feebly emarginate; base transverse, very broadly, feebly sinuate toward the middle; sides strongly arcuate at apical third, thence strongly convergent and REVISION OF ELEODIINI—BLAISDELL. 103 nearly straight to the apex, distinctly sinuate before the base; disc rather coarsely, deeply punctate, densely so at the sides, with large impunctate patches toward the middle. Elytra with disc rather coarsely, feebly, and subasperately punctate, punctures rather sparse, not much denser laterally, confusedly disposed (Casey). Diagnostic characters—The structure of the anterior tibial spurs allies this species with Aumeralis alone. The two specimens before me are smaller and shorter ovate, the punctation is sparser and much less asperate. The female resembles a large cordata in outline. T am indebted to Colonel Casey for the authentic male in my col- lection. I have before me a small series of specimens collected at Soda Springs, Idaho, which I refer to the present species until a larger series shall have been studied. One of the specimens, a male, is more strongly convex than the male received from Colonel Casey, while the females have the facies of a moderately smooth cordata, and the punctuation is slightly coarser, the integuments duller and shghtly more asperate. One male from Idaho has the anterior spur of the anterior tibie much longer and stouter than in that sex of rileyi; in a single female from the same region the anterior spur is unusually prolonged and attenuated, being about four or five times longer than the posterior, which is very short and unusually slender, in the other females the anterior spurs are at least three times longer than the posterior, much broadened and obtusely rounded at tip; these are undoubtedly worn, and those in the female mentioned above with unusually prolonged spurs represent the normal condition. I believe that a sufficiently large series of ri/eyi would show what relation exists between these forms; they can be geographical races of either humer- alis or rileyi. The genital characters are the same as in rileyi. In the authentic male of 7/eyi the mentum is moderate and sub- triangular in outline, the surface not coarsely punctate, feebly convex, and scarcely foveate laterally. The prosternum is not at all produced and vertically truncate be- hind. The mesosternum is obliquely arcuate and very feebly and broadly concave. The intercoxal process of the abdomen is evidently slightly trans- verse and about a third of its width wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a mesotibia at basal third. Post-coxal part of the first abdominal segment is equal in length to that of the process, and also equal to the third segment in length ; the second is about one-half longer than the fourth. The profemora are clavate; the grooves have distinct margins that are asperulate and become contiguous at basal third. 104 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The mesofemora are feebly tumid, and the grooves have feebly formed margins that are finely muricate, becoming contiguous and evanescent at middle. The metafemora have subparallel surface lines, and the grooves are as on the mesofemora. The tibiee are without tarsal grooves and the articular cavities are closed. All of the tarsi are missing on the authentic specimen under consideration, except a single metatarsus and that is proportioned as in humeralis. GROUP II.—TRICOSTATA SECTION. The species of this group differ from those of Group I in the broad, flattened form, with the elytra truncate at base and the humeral angles rectangular; the prothorax is likewise broadened at pase, with the angles rectangular, overlapping the humeral angles of the elytra. The pronotal disc is much less declivous laterally and the marginal bead is always visible when the surface is viewed verti- pally from above; there is consequently scarcely any variation in the characteristic pronotal outline. The anterior spurs of the anterior tibiz are distinctly thickened and broadened in both sexes, less so in the male where they are always more so than in the same sex of the preceding group. The first joint of the anterior tarsi 1s always more or less thickened at tip beneath and even shghtly produced; a tuft is also evident. In the females the dorsal plates of the genital valves are broad and oblong. (See Plate 1, fig. 11, and Plate 3, fig. 20.) There is also a development of pubescence in one species at least. The hairs are setiform. ELEODES TRICOSTATA Say. Blaps tricostata Say, Journ. Acad. Nat. Sci. Phila., III, 1823, p. 262. EKleodes tricostata LECoNTE, Proc. Acad. Nat. Sci., Phila., 1858, p. 181.— Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 307. Bleodes planata Souter, Studi Ent., 1848, p. 366. Pimelia alternata IKirpy, Fauna Boreali-Amer., IV, 1837, p. 232. Hleodes robusta LEContTr, Proc. Acad. Nat. Sci. Phil., 1858, p. 183. Oblong to oblong-oval, black, opaque, clothed with short setiform hairs, each arising from a puncture. ITecad moderate, feebly convex, rather finely, more or less densely and evenly punctate, punctures more or less sparse on the vertex; frontal suture usually defined, surface more or less broadly impressed across the fronto-epistomal junction. Antenne moderately short, not reaching to the prothoracic base, outer three joints very feebly com- pressed, scarcely dilated, third joint hardly as long as the next two ahs REVISION OF ELEODIINI—BLAISDELL. 105 taken together, fourth very slightly longer than the fifth, the latter with sixth, seventh, and eighth subequal in length, the eighth feebly triangular, ninth and tenth suborbicular, eleventh oval. Pronotum widest at the middle, a little less than one-half wider than long and at least twice as wide as the head ; disc moderately con- vex, with broad, shallow basal impressions, finely, evenly and rather densely punctate, laterally feebly asperate and rugulose; apex evenly and somewhat deeply emarginate, marginal head rather fine; s¢des broadly, evenly, and rather strongly arcuate in the anterior four-fifths, thence to base oblique or more or less sinuate, marginal bead some- times fine, at other times rather coarse; base squarely truncate to feebly emarginate, rather coarsely margined and about two-thirds wider than the apex; apical angles distinct, not acute, sometimes shehtly rounded; basal angles usually rectangular, rarely shghtly rounded or feebly obtuse. Propleurw quite evenly and rather finely muricately punctate, at times more or less rugulose. Elytra oblong-oval, usually widest at the base, sometimes at the middle; base truncate, slightly overlapped by and a little wider than the base of the prothorax; humeri quite rectangular, overlapped by the basal angles of the prothorax; sides subparallel to feebly arcuate, more or less arcuately converging in apical third, apex obtuse; disc usually flattened, very feebly to moderately convex, laterally sub- acutely margined by the subhumeral costa, arcuately declivous pos- teriorly, more or less costate, there being a common sutural, two discal, a humeral and subhumeral costa on each elytron, alternating coste being usually more or less rudimentary or obsolete, sulci more or less shallow ; surface scabrous from rather fine, densely placed muricate punctures, which become slightly tuberculiform on the slightly ele- vated intervals; inflexed sides inwardly and obliquely declivous, sur- face plane and muricately scabrous; pubescence dual, consisting of nearly erect brownish sete, and subfusiform, paler, flattened, more fragile and semi-reclining subsquamiform hairs. The costv are usu- ally more coarsely muricate. Epipleure rather narrow and gradually diminishing in width from base to apex, the superior margin rather strongly and broadly sin- uate beneath the humeral region; surface evenly and muricately punctate. Sterna more or less evenly and muricately punctured. Parapleure rather finely and asperately punctate. Abdomen somewhat shining, finely, submuricately, and rather sparsely punctulate, sometimes rugulose. Legs somewhat slender. Anterior femora mutic; anterior tibial spurs dissimilar in the sexes, the anterior spur curved, larger, and longer than the posterior. Anterior tarsi slightly dissimilar in the 106 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. sexes, first Joint shghtly thickened and feebly produced at tip be- neath with a tuft of spinules interrupting the groove. Male.—Oblong-oval, somewhat elongate. Antenne reaching to the posterior fifth of the prothorax. Elytra usually widest at the base and thence arcuately narrowing to apex. Abdomen slightly oblique, moderately convex, distinctly impressed at base of the first segment and between the coxe. Anterior spur of the anterior tibia about one- half to one-third longer than the posterior, gradually tapering from base to apex, and acute. First joint of each anterior tarsus with a small, subacute tuft of yellowish modified pubescence on the produced tip beneath; second joint slightly thickened at tip beneath with a similar and rather inconspicuous tuft; groove interrupted. Female.—Oblong, robust. Antenne reaching to the posterior fourth of the prothorax. Elytra broadly oval and usually widest at the middle, scarcely narrowed but more or less arcuately rounded pos- teriorly. Abdomen horizontal and rather strongly convex. Anterior spur of the anterior tibia about twice as long as the posterior, curved, distinctly broadened, with sides quite parallel, somewhat narrowing at tip. First joint of an anterior tarsus with a tuft of ordinary pic- eous spinules on the thickened tip; second joint unmodified. The present species is quite variable, and I deem it proper to indi- cate four incipient races, as follows: Forma typica.—Slheghtly elongate in form, pubescence usually pale flavate in color. Elytra distinctly flattened, normal costz well devel- oped, the intermediate completely obsolete or very feebly indicated, intervals broad and flat. Forma ovalis—Oval, short and robust in form. Pubescence usually black. Elytra rather strongly convex, normal cost well developed and somewhat glabrous and shining, intermediate coste more or less evident. Forma costata—General form as in typica but smaller. Elytral coste more or less strongly and quite equally developed. Forma robusta.—Form broadly oblong, large. Elytra more or less convex, all of the costey more or less developed, rather coarsely muricate, at apex becoming subspiculiferous. Measurements —Males: Length, 18-22.5 mm.; width, 6.5-10.5 mm. Females: Length, 14-22.5 mm.; width, 7-11.5 mm. Genital characters, male-—Edeagophore flaxseed-shaped, slightly elongate and not arched. Basale oblong, evenly convex and more or less gibbous in basal half; sides evenly arcuate. Apicale triangular, slightly elongate, feebly arched; dorsal surface evenly convex, with a moderately wide median membranous groove in apical half; sides feebly arcuate or nearly straight; apex subacute ; base more or less broadly arcuate, scarcely sinuate laterally. REVISION OF ELEODIINI—BLAISDELL. 107 Sternite transversely parabolic. Each lobe with the outer border evenly arcuate; apex more or less rounded ; internal border more or less straight; surface feebly convex, shining, sparsely to rather densely punctate, setose, sete not dense and moderate in length. Membrane slightly setose across the bottom of the sinus, the latter moderate in size, Female.—Genital segment quadrate, shghtly longer than wide, with a shght antero-posterior convexity, setose. Valvula (Plate 3, fig. 20).—Dorsal plate oblong, varying more or less in width. Surface glabrous and shining, plane to feebly concave, sparsely punctate in apical half and along the internal moiety to base, setee rather short; sides subparallel; outer lobe of apex not differentiated from the apical margin of the plate, evenly rounded, well developed, scarcely defined from the very small and hardly visible internal lobe. The valvular apex obliquely and inwardly truncate, sometimes very shghtly sinuate at the inner third, apical area set with rather long and flying sete. Appendage small and conical, partly visible from above, directed backward and more or less inward. Superior pudendal membrane long, reaching to the base of the in- ternal lobe of the apex, and longitudinally rugulose. Ventrolateral surface more or less evenly and moderately convex, sparsely punctate and setose, seta rather short; submarginal groove arcuately bending inward to the fossa, well defined beneath the explanate external border of the dorsal plate and external apical lobe, the latter set with quite long flying hairs. Internal margins of the valves contiguous in basal fifth, genital fissure broadly fusi- form, margins of the valves not closely approximated at apex. In- ferior pudendal membrane visible in basal half of the genital fissure. Habitat.—Texas (Clifton, Bosque County, G. W. Belfrage: San Diego, E. A. Schwarz; San Antonio, H. Soltau; Alice, E. A. Schwarz; Mobeetee, October); New Mexico (Santa Fé Canyon, 7,000 feet, August, F. H. Snow; Albuquerque, February, collector, H. Soltau; Las Vegas, August, Barber and Schwarz); Oklahoma (South Mc- Alester, June, H. F. Wickham); Kansas (Topeka, Hubbard and Schwarz; Riley County, May and September, Popenoe; Finney County, September, H. W. Menke, collection of Ralph Hopping; Onaga) ; Colorado (Colorado Springs, June and April, H. Soltau; Glenwood Springs, June, Hubbard and Schwarz; Denver, H. Soltau; Gulnare, Las Animas County, collection of California Academy of Sciences; La Junta, Golden, Poudre Canyon, near Long’s Peak, Estes Park, Lamar, Sterling, Holly, Fort Collins, La Veta, Veta Pass, Wickham’s list) ; Iowa (Sioux City, April, H. Soltau) ; “Taken at Ames by Professor Osborn, while Professor Shimek has taken it in Lyon County (June); eastern Emmet County (August); southern 108 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Dickinson County (August) and eastern Woodbury County (Sep- tember) ; it extends as far east as Independence, Buchanan County, September, on a broad, dry sand flat, along theWapsipinicon. Bot- tom” (H. F. Wickham, Iowa Acad. of Sci., 1899); Wyoming (Cheyenne, May; collectors, Hubbard and Schwarz); Montana (Helena, May; Assiniboine, August; collectors, Hubbard — and Schwarz) ; Southern Dakota (Sioux Falls, May, H. Soltau) ; British America (Medicine Hat, September, C. V. Riley). Number of specimens studied, 110. Type destroyed. Type-locality given by Say, Missouri and Arkansas. Salient type-characters.— Body oblong-oval, with numerous minute prostrate hairs. Thorax transversely subquadrate, anterior edge concave; basal edge rectilinear, covering the base of the elytra; lateral edge arcuate; anterior angles rounded; posterior angles excurved. Elytra scabrous, with three elevated lines, a common sutural, and a lateral one on the edge, intermediate spaces broad, flat, not concave, and sometimes with the appearance of a slightly elevated longitudinal line alternating with the others (Say). Diagnostic characters.—A. distinct species, dull in color, sparsely pubescent, and with more or less costate elytra. In the typical form there is a humeral, a subhumeral, sutural, and two discal coste, the intervening costx obsolete or very feebly indi- ‘ated, with the intervening spaces flat and wide. In the costate form all of the cost are developed and of about equal prominence; the individuals are usually under the average size and decidedly oblong in form; males more frequently observed. The individuals of the oval form are more or less oval and rather convex, the cost are normal, strong, and frequently glabrous, the pubescence may be blackish or of the normal fulvous color. The robust form is a larger, stouter, and decidedly more coarsely sculptured variety. Laterally and at apex the murications are quite spiculiform. The mentum is comparatively small, triangular to trapezoido- triangular in general outline; surface feebly convex, more or less shghtly impressed along the sides at times, rather finely punctate, each puncture with a short seta. Prosternum comparatively broad between the coxe, usually convex antero-posteriorly and more or less arcuate or vertical behind, rarely subtruncate or submucronate. The mesosternum is usually arcuately vertical and more or less moderately concave, sometimes feebly so. The intercoxal process of the first abdominal segment is quadrate and equal in length to the post-coxal part of the same segment; it is also equal to the second in length; the third segment is one-half J renee nit gh cal a aa amr rare amare ©.” REVISION OF ELEODIINI—BLAISDELL. 207 ELEODES GRANDICOLLIS Mannerheim. Eleodes grandicollis MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 266; Mag. Zool., XIV, 1844, No. 180 (fig.).—Horn, Trans. Amer. Phil. SOCs ONG Ved SOs ps alae Hleodes valida BoHEMAN, Freg. Eugen. Resa, Ins., 1858, p. 90. Large and robust, oblong oval, black, and shining. [ead twice as wide as long and more or less feebly convex, quite coarsely and rather irregularly, sparsely, and at times densely punc- tate; punctures denser at the periphery. Antenne rather short and stout, reaching to about the posterior fourth of the prothorax and sub- equal in the sexes; outer joints shghtly compressed and not noticeably dilated; third joint equal in length to the next two taken together ; the fourth shehtly longer than the fifth; the latter, sixth, and seventh subequal in length; eighth and ninth about as long as wide; tenth slightly wider than long; eleventh smaller, trapezoidal and longer than wide. Pronotum widest at about the middle, less than one-third wider than long, and more than twice as wide as the head; dise smooth and shining, quite strongly convex, and rather strongly declivous at the angles; surface finely, evenly, and rather sparsely punctate, narrowly opaque, and asperate along the lateral margins; apex trun- cate or feebly emarginate and rather finely beaded; s¢des broadly and evenly arcuate, somewhat straight, and converging posteriorly, scarcely ever sinuate, finely beaded; base slightly arcuate and rather coarsely margined; apical angles distinct and deflexed, at times more or less rounded; basal angles obtuse and never prominent. Propleure smooth, more or less strongly rugulose and at times sparsely punctate. Elytra oval, usually estriate, smooth, and shining, about twice as long as wide, widest at the middle; not emarginate nor abruptly declivous at the base, the latter slightly narrower than the con- tiguous prothoracic base; humeral angles absent; s¢des evenly arcu- ate, apex obtuse and rounded; disc very convex, suddenly and rather _ obliquely declivous posteriorly ; surface finely and quite evenly pune- tate, punctures arranged in moderately distant, unimpressed series, the individual punctures not closely placed; intervals with a series of distantly and regularly, placed punctures, the punctures becoming subasperate at the sides, and more strongly so on the apex. E’pipleure moderately narrow and very gradually and evenly narrowing from the base to the apex; surface smooth and sparsely punctate. Sterna moderately and quite densely punctate and rugulose. Parapleure rather strongly and quite evenly punctate. 208 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Abdomen sparsely and quite evenly punctate, more or less irregu- larly and reticulately rugulose; punctures denser on the fifth segment and finely setose. : Legs moderate in length and very stout. Anterior femora in both sexes, armed with an acute tooth near the external fourth; tibial spurs slightly broadened, those of the protibiz equal and acute. Tarsi similar in-the sexes and rather strongly spinulose. Male—Robust. Elytra oblong oval and at least twice as long as wide, nearly vertically declivous posteriorly. Abdomen moderately oblique, not strongly convex; segments one and two broadly and more or less strongly impressed. Tibial spurs feebly broadened. Female——Very robust. Elytra broadly oval and less than twice as long as wide, vertically declivous posteriorly. Abdomen hori- zontal rather strongly convex, and not impressed. Tibial spurs noticeably broadened. In the material before me two forms can be recognized, notably those from central California, and those from southern California, Arizona and Lower California. Mannerheim’s type was undoubtedly collected somewhere in the vicinity of San Francisco. Forma typica——Thorax narrower with the sides not strongly rounded, and the apical angles frequently less strongly marked. Elytra less broadly oval, so that the general form is less robust and narrower. | Measurements.—Males: Length, 27-29 mm.; width, 10.5-11.5 mm. Females: Length, 29 mm.; width, 11.5-12 mm. Forma valida.—Thorax broader and more strongly rounded at the sides; elytra broadly oval, and the general form much more robust. Measurements.—M ales: Length, 29.-32. mm.; width, 10.5-11.5 mm. Females: Length, 32.5-34. mm.; width, 14.-15.5 mm. Genital characters, male——Kdeagophore comparatively small and elongate oval. Basale suboval, widest in basal half, evenly convex from side to side, slightly arched; dorsum with a rather strong antero-posterior convexity and not gibbous in basal half, apically much narrower than at middle and the sides rather strongly arcuate. Apicale small and rather elongately triangular; surface evenly convex, not grooved; sides nearly straight, scarcely sinuate, the apex being shghtly produced and very narrowly rounded at tip; base with the median third obtuse, not rounded nor acute and feebly sinuate laterally. Sternite slightly transverse. Each lobe rather long, broadly rounded at tip, angle scarcely evident; external border nearly straight and rather strongly convergent apically; internal margin shghtly arcuate or straight; surface quite densely punctate in apical os bundle eae ees Urea) mah Saxe eS AAR ace a) REVISION OF ELEODIINI—BLAISDELL. 209 two-thirds, setose, setee dense at tip and rather long. Membrane not setose across the bottom of the narrow sinus. Female.—Genital segment triangulo-trapezoidal, rather robust, comparatively small and setose. Valvula (Plate 1, fig. 8).—Dorsal plate oblong, glabrous; surface broadly concave, more deeply so along the strongly reflexed inner side, sparsely punctate, each puncture with a long flying seta; exter- nal border rather straight, and the internal somewhat sinuous; apical margin truncate, angle quite rectangular and set with quite long sete. Apex rather short, semi-chitinous and finely setose, with a tuft of rather long sete at tip. Fossa large and fringed with rather long sete. Appendage conical, truncate, moderate in size and setose, with a loose pencil of hairs at tip. Basal prominences rather strongly developed. Superior pudendal membrane not reaching to the middle of the dorsal plate. Ventrolateral surfaces rather strongly and transversely prominent in the basal moiety, broadly and rather deeply concave before the apex; apical region punctate and rather thickly set with quite long flying sete. Submarginal groove feeble. Internal margins of the valves contiguous in basal fifth. Forma typica.—Habitat—Central California (San Francisco, Van Dyke; Marin County, Charles Fuchs; Calaveras County, elevation 2,000 feet, F. E. Blaisdell; Monterey County; Tulare County, Ralph Hopping). ° Forma valida.—Southern California (San Diego, Los Angeles, Monterey, San Bernardino, and Kern counties); Arizona (Yuma Desert, March; Santa Maria River). Lower California (San José del Cabo; Lower Paraiso; San Pedro Martir). Number of specimens studied, 59. Location of type unknown to me. Ty pe-locality—* California ” (Mannerheim). Salient type-characters.—Elongate, convex, and shining. Head deeply punctate. Thorax with sides strongly rounded, margin re- flexed, finely punctulate. Elytra oblong, moderately narrowed each side of the apex, obsoletely striato-punctate. Anterior femora den- tate (Mannerheim). Diagnostic characters.—The only species in the subgenus with obso- lete humeri. One of our largest and finest species, very distinct and with obscure relationships. I believe that a study of the Mexican species is necessary before much light can be thrown upon the origin of not only grandicollis and dentipes, but armata. 59780— Bull. 68—09——_14 910 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. In the series of examples before me the mentum is variable. Although large, it is moderate when taken in a comparative sense, varying from trapezoidal to quadrato- or parabolo-trapezoidal; apex truncate to evenly arcuate; surface scarcely convex, strongly punctate and almost scabrous, feebly foveate at the basal angles. The prosternum is variable and strongly protuberant ventrally with the coxe, being almost declivous from the anterior margin. It is usually longitudinally grooved between the coxee, rounded antero- posteriorly, and not mucronate behind, or with an abruptly developed mucro; at times convex between the coxe and subtruncate behind, with a deflexed mucro. The mesosternum is arcuately or obliquely declivous and more or less strongly concave. The abdominal process is about a fourth of its width wider than the metasternal salient, subquadrate, and equal in length to the post- coxal portion of the first segment, which is also equal to the third, the latter one-sixth of its length shorter than the second; the fourth is one-half of its length shorter than the second. Metasternum laterally between the coxe is about as long as the width of a mesotibia at apex. Femora stout. The profemora are thickest at the junction of the outer and middle thirds, from that point gradually narrowing each way. Tibial grooves moderately large, edges rather thick, subearini- form, very feebly arcuate, and converging to become contiguous a short distance before the base; floors very feebly concave, subopaque, and extremely, finely, and reticulately rugulose. Anterior margin dentately laminate at the outer fourth. Meso- and metafemora of nearly the same width throughout; sides quite parallel. Mesofemora with the grooves moderate in width, margins feebly cariniform, converging very gradually to become evanescent a short distance from the base; floors of the grooves scarcely concave, opaque as on the profemora. The metafemora have the grooves about moderate in width, with the sides feebly cariniform, gradually converging to become evanes- cent at basal third; the floors are as on the profemora. The protibiz are shghtly compressed, carinate in basal half exter- nally, with the tarsal grooves more or less feebly developed, asperate and limited posteriorly by a muricato-serrulate margin, articular cavities closed. Mesotibiz more or less rounded, briefly or obsoletely carinate exter- nally just before the base; tarsal grooves simply indicated by a smooth opaque line, or feebly channeled surrounding surfaces muri- cate; articular cavities closed. REVISION OF ELEODIINI—BLAISDELL. OA. The metatibize rounded, rather small at base and gradually thick- ening to apex; tarsal grooves obsolete or feebly indicated, articular cavities closed. Tarsi rather long and stout. The protarsi are about a fifth of their length shorter than a meso- tarsus. Joints two, three, and four equal in length, rather short and distinctly wider than long, and together equal to the length of the fifth ; first about as long as wide. Mesotarsi just a little shorter than a metatarsus. Joints two, three, and four equal in length, scarcely as long as wide, and together about equal to the length of the fifth; first joint a little longer than wide. The metatarsi are about half as long as their metatibia. Joints two and three equal in length and scarcely longer than wide, and together about as long as the fourth; first joint about one-and-a- half times longer than wide. DENTIPES GROUP, Section A. The constituents of the present section have the integuments more or less opaque, more or less asperately and muricately sculptured ; the surface is never normally entirely glabrous nor shining; further- more, the species are undoubtedly genetically related by having the inferior surface of the cauda, when present, formed by the dilated apical extremity of the epipleure. A part of the species are ecaudate, the elytral apices being but shghtly produced, and in the indifferent stage of development. There is also a tendency to hirsuteness. With the development of the elytral cauda, the femoral teeth become abortive. The protibial spurs are acute and quite similar in the sexes, not strongly divergent as in the Obscura Group, and instead have a tend- ency to become parallel and backwardly curved. The mentum is distinctly bearded in a part of the species, but here it occurs as a part of a tendency to general pilosity. The head is moderate; the articular cavities of the tibiz are nearly closed, and as a rule the tarsal grooves of the tibiz are scabrous. The abdomen is horizontal in both sexes, and the intercoxal salient is always dis- tinctly wider than the metasternal process. The genital characters of this section are not sufficiently character- istic to be of practical use when tabulated. The male genital characters present nothing more than those enu- merated as subgeneric. The most important character is the presence of the membranous groove on the apicale. In the female the genital segment is triangular or trapezoidal in outline, the dorsal plate of the valves is concave and not retracted, the submarginal groove is scarcely visible from above, the appendage flattened or conical, the superior pudendal membrane does not reach 22 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. beyond the middle of the dorsal plates, and the genital fisstire is open and fusiform. Our species may be separated as follows: Species ecaudate. Elytra suleate; intervals more or less convex =2=22—= 222-5) ses hispilabris. Hlytra: substrate; intervals stlats sss eee Sponsa. Species caudate. Dorsal surface of body not clothed with long, flying hairs________ caudifera, Dorsal surface clothed with long, flying hairs’ 2-2 longipilosa. The female genital characters may be tentatively formulated as follows: Genital segment more or less quadrato-trapezoidal; appendage flattened and subtriangular. Dorsal plate oblong, sides parallel; apical margin subtruncate_____ Sponsa. Dorsal plate irregularly oblong; apical margin more or less arcuate. hispilabris. Genital segment more or less triangular; appendage conical. Dorsal plate oblong, sides subparallel; apical margin more or less oblique. caudifera, Dorsal plate oblong-oval, sides arcuate; apical margin more or less arcu- ate eS ee we A Rh ee ee ee eee longipilosa, Genealogy.—An examination of the accompanying diagram will elucidate the apparent relationships and lines of descent. The lines of divergence are apparently the ac- quisition of the elytral cauda on the one hand and the elytral apices re- maining indifferent on the other; in the former the development of an Sculptiis unusual degree of pubescence with the disappearance of the femoral Elongeta teeth are very striking morpholog- ical coincidents along the lines of correlated characters. Tracing the divergencies along FIG. 4.—GENBHALOGICAL DIAGRAM OF THE the ecaudate ramus of the senea- DENTIPES GROUP, Secrion A, SUBGENUS logical tree, Tee swallll appear as if ee sponsa diverged, retaining a com- paratively simple form of elytral sculpturing and has apparently reacted less than hispilabris to its environment. Hispilabris has, indeed, undergone a wonderful amount of differ- entiation into a number of incipient races, primarily by the sculp- turing remaining comparatively simple with rather fragile integu- ments, and, secondarily, by a more decided character of sculpturing having been acquired along with an increased thickness of the exo- skeletal wall. In the former instance forma typica and nupta were evolved, the latter by increased ventricosity of the body; in the latter Sponsa Longipilosa Caudate Forms REVISION OF ELEODIINI—BLAISDELL. Alay case sculptilis took on a stronger form of elytral sculpturing, e/ongata an elongate form, and /wvis by robustness and a less pronounced form of sculpturing. This method of intuitively working out and presenting these gene- alogical data—possibly more theoretical than real—may at least lay a foundation upon which to erect a more substantial knowledge by stimulating biological research and criticism. Distribution—The four species under consideration are distributed through the following States: Texas, Arizona, New Mexico, Oklahoma, Colorado, Kansas, Ne- braska, Nevada, northeastern California, Oregon, Washington, Mon- tana, and Wyoming. Sponsa is southern in habitat, occurring in Texas, Arizona, New Mexico, and Colorado. Hispilabris has the most extensive distribution, the typical form occurring in Texas, New Mexico, Oklahoma, Kansas, Colorado, Ne- braska, and Wyoming; forma nupta has thus far been found in Texas and Oklahoma; forma convewa in Texas; forma sculptilis in Arizona, Colorado, northeastern California, Wyoming, Montana, and south into Mexico; forma elongata in Arizona; forma levis in Utah, Ore- ~ gon, and Washington. Caudifera from Texas, Arizona, New Mexico, and Colorado. Longipilosa thus far from the desert regions of Nevada and north- eastern California. ELEODES SPONSA LeConte. Eleodes sponsa LECoNTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 184.—Horn, Trans. Amer Eh iSoc. MV SiO: ps sl3e Subovate, elongate, black to nigro-piceous. Head twice as wide as long, shghtly convex, and more or less im- pressed laterally and along the frontal suture; swrface more or less evenly punctate, punctures moderate, sometimes with small impunc- tate areas. Antennw more or less slender and quite long, reaching shehtly beyond the base of the prothorax, outer four joints slightly compressed, feebly and gradually dilated; third joint about equal to the next two taken together, fourth slightly longer than the fifth, the latter, with sixth and seventh subequal, eighth about as wide as long and triangular in outline, ninth and tenth wider than long and trans- versely oval, eleventh subovate. Pronotum widest scarcely in advance of the middle, about one- fourth wider than long, less than twice as wide as the head, and with the length equal to the apex; disc smooth and feebly shining, mod- erately and quite evenly convex, more or less declivous at the apical angles, frequently very narrowly impressed laterally along the mar- 214 BULLETIN 63. UNITED STATES NATIONAL MUSEUM. gins in anterior half, scarcely opaque, rather finely and more or less evenly and sparsely punctate; apex feebly emarginate or subtruncate, generally finely margined laterally and obsoletely so at middle; sides evenly and moderately arcuate anteriorly, less so or rather straight and convergent from middle to the base, sometimes feebly sinuate in front of the basal angles, marginal bead fine; base feebly arcnate, finely beaded, and equal to the apex; apical angles acute, prominent anteriorly and frequently everted; basal angles subrectangular or shghtly obtuse. Propleure smooth, obsoletely punctulate and rugulose, or impunc- tate. Elytra elongate oval, feebly shining to slightly opaque, about twice as long as wide, usually widest at the middle; base very feebly emargi- nate and equal to or slightly wider than the contiguous prothoracic base; Aumeré obtuse, sometimes shghtly prominent; s¢des evenly arcu- ate, sometimes narrowing towards base, apex more or less obtuse, never produced; disc more or less evenly convex from side to side, or somewhat flattened, more or less rapidly and arcuately or obliquely declivous posteriorly ; surface feebly substriate, the strie with rather coarse, closely placed, and more or less impressed punctures, the inter- vals flat or at times slightly convex and with a more or less uneven series of submuricate punctures; sometimes the punctures are in- dented, both series becoming more strongly muricate at the sides and apex. E'pipleure moderately narrow, with a broad and gentle upward curve at base, thence narrowing gradually to apex; surface smooth and obsoletely punctulate. Sterna usually more or less feebly punctate and rugulose. Parapleure more or less strongly punctate. Abdomen horizontal, usually smooth and obsoletely punctulate, fifth segment rather more strongly punctate. Legs moderate in length and rather slender, profemora more or less armed in the sexes; protibial spurs quite equal and acute; tarsi similar in the sexes. Male.—Somewhat narrow. Antenne with about three joints reaching beyond the prothoracice base. Elytra rather rapidly, arcu- ately, and obliquely declivous posteriorly, apex moderately acute. Abdomen moderately convex. Profemora with an acute or obtuse tooth at outer fourth. Female——Somewhat robust. Antenne with about one joint ex- tending beyond the prothoracic base. Elytra arcuately and nearly vertically declivous posteriorly ; apex more or less obtuse. Abdomen strongly convex. Profemora mutic or with a very obtuse tooth at about the outer fourth. REVISION OF ELEODIINI—BLAISDELL. Ores Two forms are to be recognized, namely: Forma typica.—Thorax moderate in width, sides not very strongly rounded; elytral punctuation more or less muricate, disc striato- punctate, interstitial punctures smaller or at most subequal to the strial punctures, and usually not at all indented. Forma convexa.—Thorax wider, form more convex, punctuation not at all muricate, elytral punctures moderately coarse, the strial closely placed, the interstitial larger or subequal and more or less in- dented, and more or less irregular in some intervals or laterally. Measurements—Males: WLength, 22-25 mm.; width, 8-10 mm. Females: Length, 22.5-26 mm.; width, 9.5-10 mm. Genital characters, male.—Kdeagophore moderate in size and of the usual oblong-ovate form. Basale oblong and moderate in length. Apicale about twice as long as wide at base, triangular and evenly convex above, with a feebly impressed linear depression in middle third; sides very feebly arcuate from base to apex, the latter nar- rowly rounded and obtuse; base broadly and subacutely lobed at the middle and sinuate laterally. Sternite transverse. Each lobe triangular with the external bor- der evenly arcuate from base to apex; internal margin straight; apex subrectangular and narrowly rounded; surface sparsely punctate and setose in apical two-thirds, sete: moderately long and not very dense. Membrane not setose across the bottom of the sinus. Female—Genital segment subquadrato-trapezoidal and moderately setose. Valvula (Plate 4, fig. 8)—Dorsal plate oblong, with the surface more or less broadly concave, margins more or less reflexed and sub- parallel; finely and sparsely punctate, setose; external margin feebly sinuate to slightly arcuate; internal margin arcuate; apical margin subtruncate or feebly rounded, with a few short sete. Angle more or less rounded. Apex short and triangular, acute at tip and very finely setose. Appendage short and depressed, wider at base than long, broadly rounded at tip, setose, setze rather short forming a loose tuft; external angle of the fossa fringed with a few quite long sete. Submarginal groove small and sometimes visible from above. Superior pudendal membrane reaching to about the middle of the dorsal plate and longitudinal rugulose. Ventrolateral surfaces ——Body transversely inflated and with the basal prominences moderate; quite strongly concave laterally and less so centrally in front of the apices; quite strongly punctate in apical three-fourths, each puncture with a rather short semi-erect hair. Genital fissure moderately narrow. 2916 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Habitat—New Mexico (Santa Fé, Hubbard and Schwarz; Cool- idge, Coll. E. C. Van Dyke; Aztec, April, Coll. C. F. Baker; High- rolls, June 11); Colorado (southern, George Horn) ; Texas (Marfa, elevation 4,600-4+,800 feet, and at Alpine, July, 4,400-6,000 feet, H. F. Wickham); Arizona (M. L. Linell). Number of specimens studied, 20. Type in the LeConte collection. Type-locality—New Mexico; collected by Mr. Fendler. Salient type-characters.—Thorax a little convex and somewhat rounded, sparsely punctate, anteriorly broadly emarginate, apical angles subacuminate; sides rounded; base subrotundate with the angles subrectangular. Elytra at apex obtusely acuminate; disc punctato-striate, laterally and at apex muricate, intervals sparsely uniseriately punctate (LeConte). Diagnostic characters.—S ponsa is to be separated from hispilabris by the elytra being feebly substriate and with the intervals flat, rarely feebly convex and rather coarsely muricately punctate. In some specimens the interstitial punctures are in slightly larger dents and the sculpturing is not at all muricate but smooth. The thorax is usually less rounded on the sides and nearly quadrate, but there is considerable variation. The elytral sculpturing sometimes resembles specimens of /ispil- abris which have the intervals scarcely convex. In a complete series the transition from the one species to the other is evident. In sponsa the elytral disc is usually flattened. LLeConte mentioned a specimen collected in New Mexico that had the elytra not at all muricate. The mentum is variable. In size it may be comparatively large or small, varying in outhne from parabolo-trapezoidal, trapezoidal to trapezoido-triangular, rarely the apex is subacute; surface coarsely punctate to almost impunctate, evenly convex or more or less foveate laterally. The prosternum and mesosternum as in héspilabris; the latter is sometimes quite vertical and deeply concave at middle. The abdominal process is about as wide as the third segment is long, the post-coxal part of the first is shorter than the second seg- ment, and the latter is twice as long as the fourth; the third is about one-fourth of its length shorter than the second. The abdominal salient is but slightly wider than the metasternal process. . The metasternum laterally between the cox is scarcely as long as the width of a mesofemur at base. The tibiz and femora are about the same as in hispilabris. The articular cavities of the tibiwe are very narrowly open as a rule or closed. 5 REVISION OF ELEODIINI—BLAISDELL. Oe The tarsi are moderate, more slender in the smaller individuals and stouter in the larger. The protarsi are about three-fifths of their length shorter than a mesotarsus. Joints two, three, and four are subequal, and just a little wider than long, the three combined about equal to the fifth; the first is distinctly longer than wide. The mesotarsi are about as long as a metatarsus. Joints two, three, and four subequal and about as long as wide; together about equal to the fifth; the first joint is scarcely twice as long as wide. The metatarsi are about one-half as long as a metatibia. Joints two and three are subequal and distinctly longer than wide, together subequal to the fourth; the first is scarcely as long as the fifth. ELEODES HISPILABRIS Say. Blaps hispilabris Say, Journ. Acad. Nat. Sci. Phila., III, 1823, p. 259; Amer. Entomology, I, 1824, p. 32.—LeContTr, Complete Writings of Thomas Say, I, 1859, p. 32. Hleodes hispilabris LeContr, Complete Writings of Thomas Say, II, 1859, p. 152.—Horn, Trans. Amer. Phil. Soc., IV, 1870, p. 313.—CHAM- PION, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 79. Eleodes sulcata LEeContTE, Proc. Acad. Nat. Sci. Phila., 1852, p. 67; 1858, Os asi Eleodes lecontei GEMMINGER, Coleopt. Hefte, VI, 1870, p. 122 (new name). Eleodes connerxa LECONTE, Reports of Explor. and Survey, ... 47th and 49th parallel, XII, Pt. 3, 1857, p. 49. EHleodes nupta LeEContTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 183. Eleodes compositus Casry, Annals N. Y. Acad. Sci., VI, Nov., 1891, p. 58. Elongate ovate, somewhat shining, more or less convex and sulcate ; color black, frequently reddish along the suture. Head about twice as wide as long, feebly convex, finely and not densely punctate, feebly impressed laterally. Antenne long, mod- erate in stoutness, reaching to the prothoracic base, outer four joints shehtly compressed and noticeably dilated; third joint equal in length to the fourth and fifth combined, fourth a ttle longer than the fifth, the latter, sixth, and seventh subequal, eighth equilaterally triangular in outline, ninth and tenth transversely oval, eleventh irregularly ovate to subtrapezoidal. Pronotum widest at about the middle, about one-fifth to less than one-third wider than long, and less than twice as wide as the head; disc smooth and more or less shining, quite evenly and moderately convex, more or less declivous at the apical angles, finely, sparsely, and quite evenly punctate, very narrowly opaque and granulate later- ally; apex more or less feebly and broadly emarginate, finely beaded laterally and obsoletely so at middle; sides broadly and evenly arcuate anteriorly, converging and more or Jess straight posteriorly, -or shehtly sinuate before the basal angles, finely to rather coarsely 918 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. beaded, and very narrowly or more or less reflexed; base feebly arcu- ate and more strongly margined, a little wider than the apex and equal to the length; apical angles acute and more or less everted ; basal angles obtuse. Propleure smooth, more or less rugulose and nearly impunctate. Elytra oval, sulcate, usually less than twice as long as wide, widest at the middle; base very feebly emarginate, usually very little wider than the contiguous prothoracic base; humeri obtuse and not promi- nent; sides evenly arcuate, apex more or less acute and usually very shghtly produced; disc more or less convex, frequently depressed, and more or less suddenly obliquely declivous posteriorly ; surface usually deeply suleate, sulei deep and about as wide as the intervals, each with a series of more or less evenly and closely placed, small asperate punctures; the intervals are more or less strongly convex, smooth and shining, each with a single irregular series of distantly placed punc- tures. The deflexed sides are more coarsely, irregularly punctate and subsuleate. Epipleurew moderate in width, gradually narrowing from base to apex, where they may be feebly dilated beneath the elytral apices and sightly excavated at that point; surface smooth, frequently im- punctate, at other times very sparsely and finely punctulate. Sterna moderately punctate and rugulose. Parapleure usually quite smooth and sparsely but not deeply punctate. Abdomen horizontal, smooth, very finely punctulate and rugulose ; fifth segment more strongly punctate. Legs somewhat slender. Anterior femora armed in the sexes; protibial spurs subequal, frequently the posterior appears to be just the least longer and stouter than the anterior. Tarsi grooved beneath and similar in the sexes. Male-—Somewhat narrow and elongate. Antenne reaching slightly beyond the base of the prothorax. Length of the pronotum slightly greater than the width of base and about one-fifth wider than long. Elytra arcuately and obliquely declivous posteriorly. Abdomen moderately convex from side to side. Anterior femora with a moderate and acute tooth at about the outer third. Female.—More or less robust. and rather broad. Antenne reach- ing to or shghtly beyond the prothorax, the latter with the base equal to the length and about one-third wider than long. Elytra rather suddenly and arcuately declivous posteriorly. Abdomen strongly convex. Anterior femora with a more or less obtuse tooth at about the outer third. Genital characters, male-—KEdeagophore of the usual oblong-ovate form and small. REVISION OF ELEODIINI—BLAISDELL. 219 Basale rather short, not a great deal longer than the apicale, not usually arched nor strongly convex. Apicale elongately triangular, quite evenly convex above, with a linear impression in middle third; sides feebly arcuate from base to apex, the latter subacute; base rather acutely lobed at middle and scarcely sinuate laterally. Sternite transverse. Lobes rather small; each with the external border straight and converging from base to apex, the latter rounded, angle introrse and scarcely developed; internal margin straight; surface punctate and moderately thickly setose, setae moderate in length and not extending upon the membrane across the bottom of the sinus, the latter short but rather broad. Female—Genital segment quadrato-trapezoidal, width at base about equal to the length, surface rather conspicuously setose. Valvula (Plate 3, fig. 19)—Dorsal plate more or less irregularly oblong, quite deeply concave, with margins reflexed; surface very finely and sparsely punctate, setee rather long and flying; external margin feebly sinuate to slightly arcuate; internal border rather broadly arcuate, sometimes subangulate near base; apical margin evenly rounded and set with quite long flying hairs, angle rounded. Apex short and triangular, acute at tip, finely setose; external angle of the fossa quite thickly set with rather long hairs. Lateral plate visible from above external to the dorsal plate, with submarginal groove between the two and more or less visible. Appendage depressed, more or less triangular and rounded at tip, not longer than the apical lobe; surface set with quite long sete. Superior pudendal membrane reaching to about the middle of the dorsal plate, and longitudinally rugulose. Ventrolateral surfaces—Body rather strongly convex at base, not inflated, glabrous, more or less strongly and transversely concave before the apices; apical moiety sparsely punctate, with sete rather long. Internal margins of the valves contiguous at base for a short distance. Genital fissure rather wide and fusiform, closed in basal half by the inferior pudendal membrane. Salient type-characters.—More or less shining. Elytra frequently more or less depressed, and somewhat narrowly rounded laterally, frequently more or less reddish along the suture; sulci moderate in depth, intervals more or less convex. The apical angles of the pro- notum are acute and more less everted. Say’s description is too brief as to the essential characters. The following incipient races or forms should be recognized: Forma typica.—See salient type-characters. Measurements —Males: Length, 20-25.5 mm.; width, 8.5-10.5 mm. Females: Length, 18-28 mm.; width, 8.8-12 mm. 220 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Number of specimens studied, 150. Type destroyed. Pype-locality. —* Inhabits Missouri,” a badly defined geographical region in Say’s day. Forma nupta LeConte.—Elytra with the dorsum slightly and the sides strongly convex (LeConte). The elytra are more or less evenly convex from one epipleura to the other, usually somewhat inflated, sometimes quite strongly so. Some of the smaller examples are quite ceychriform. Measurements. les: Length, 21-27 mm.; width, 9.5-11.2 mm. Females: Length, 19-29 mm.; width, 9.8-14 mm. Number of specimens studied, 15. Type in the LeConte collection. Type-locality— Laredo to Ringgold Barracks, Texas. Forma sculptilis—Integuments dull and rather thick. Head and thorax more or less shining. Elytra more or less convex, sulci usually opaque and deep, with the intervals strongly convex, shining and conspicuous, quite abruptly limiting the grooves. Measurements —Males: Length, 19-28 mm.; - width, 1828.54 mm. Females: Length, 19-24 mm. ; arate 8-10 mm. Number of specimens sahieds 40. Forma elongata.—General form distinctly elongate and narrower as compared to the other forms; black. Elytra are sulcate, but less strongly so than in sculptilis; the intervals at times are feebly convex. The prothorax is noticeably larger and the legs stout. Measurements—M ae ee 26.5 mm.; width,9 mm. Females: Length, 29 mm.; width, 10.2 mm. Number of specimens ete 6. Forma convexa LeConte.—Elongate, black and bright. Thorax moderately punctulate, sides rounded, anterior angles acute and shghtly prominent; basal angles obtuse. Elytra elongate oval, in- tervals subconvex and rugose ; subacute posteriorly. Measurements.—Length, .9; width of elytra, .382; thorax, .23 (Le- Conte). Specimens not at hand. Type is in the LeConte collection. Type-locality.—Prairie Paso, Texas; collector, Dr. Cooper. Forma levis.—Robust, integuments decidedly black, thick and alutaceous, also quite smooth. Thorax rather large. Elytra usually evenly convex from side to side, sulci very shallow, intervals very feebly convex, subobsoletely sculptured. Legs somewhat slender. Measurements.—M ales: Length, 23 mm.; width, 8mm. Females: Length, 23-26 mm.; width, 9.2-10 mm. Number of specimens suidied. 2 REVISION OF ELEODIINI—BLAISDELL. Ot The following species described by Colonel Casey evidently belongs here: Forma composita Casey.—Moderately robust, convex, intense black throughout, the pronotum dull and strongly alutaceous; elytra shin- ing, very feebly alutaceous, glabrous. [ead densely, finely, rather sparsely punctate and dull; antennee rather long and stout, fully as long as the head and thorax, third joint between three and four times as long as wide, outer joints shehtly broader, the ninth and tenth somewhat transversely oval. Pronotum about as long as wide; disc evenly convex, very minutely, sparsely punctate; apea broadly, very feebly emarginate; sides paral- lel, broadly arcuate or subangulate just before the middle, very feebly sinuate just behind the apical angles, feebly convergent and nearly straight to the base; base arcuate and equal to the apex in width; apical angles right, slightly blunt, not prominent; basal angles broadly obtuse. Elytra about two and two-thirds times as long as the prothorax, and, in the middle, one-half wider than the latter, about twice as long as wide; humeri not at all exposed; sides broadly arcuate, apex acutely angulate; disc strongly declivous behind, deeply sulcate, the ridges strongly convex and shining, each with a single rather uneven series of small, moderately distant punctures, the intervals as wide as the ridges, deep, opaque, each with a single series of rather small, close-set, asperate punctures. Legs slender, the femora shining, rather finely but strongly punc- tate, the anterior alone with a strong acute tooth near the apex; tibie strongly rugose except toward base, the spurs of the anterior small, acute, and nearly equal. Prosternum with the apex slightly reflexed, acute and tuberculiform. The above is Casey’s original description. Measurements of the type.—J/ale: Length, 16 mm.; width, 6.6 mm. Unknown to me. Type: a wnigue in Colonel Casey’s collection. Ty pe-locality.—Texas. Forma typica.—//abitat.——Missouri (Say). Colorado (August, C. V. Riley; Denver, April and June, Coll. H. Soltau; May, Colls. Dyar and Caudell; Colorado Springs, June, elevation 6,000-7,000 feet, H. F. Wickham. In his List of the Coleoptera of Colorado, Pro- fessor Wickham gives the following localities, the forms are not differ- entiated: La Junta, West Las Animas, Canyon City, San Luis Valley. Fort Garland, Trinidad, Holly, Fort Collins). Kansas (collections of Chas. Fuchs, C. V. Riley, F. H. Snow, and F. E. Blaisdell). New Mexico (Las Vegas, August, Colls. Barber and Schwarz; Albuquerque, Coll. E. C. Van Dyke). Texas (Mobeetee, October, Coll. H. S. Bar- pay BULLETIN 63, UNITED STATES NATIONAL MUSEUM. ber.) Oklahoma (Fort Supply, Coll. H. S. Barber.) Wyoming (Cheyenne, June, Colls. Hubbard and Schwarz). Nebraska (Sand Hills, September, Coll. C. V. Riley). Forma nupta.—Oklahoma (Fort Supply, Coll. H. S. Barker; Texas (Alice, E. A. Schwarz; Fort Creek, E. A. Mearns; San Diego, April and May, Hubbard and Schwarz; Goliad County, October, J. D. Mitchell). Forma sculptilis—Arizona (Walnut, July, H. F. Wickham; Oracle, July, Coll. Hubbard and Schwarz; Williams, May, Coll. Barber and Schwarz; Fort Grant, July, Coll. Hubbard and Schwarz; Ash Fork, June; Yuma, Chas. Fuchs). New Mexico (Las Vegas, August, Coll. Barber and Schwarz). Colorado (Denver, May, Coil. Dyar and Caudell). Utah (Salt Lake City, April, Coll. H. Soltau). Calli- fornia (northeastern, Coll. Hubbard and Schwarz). Wyoming (Cheyenne, May, H. Soltau; June, Hubbard and Schwarz). Mon- tana (Helena, May, Henry Ulke, Hubbard and Schwarz). Mexico ~(Cananea, district of Arizpe, State of Sonora, elevation 4,500-6,000 feet, Dr. W. C. Alvarez). Forma elongata.—Arizona (Coll. Beverly Letcher, Prescott). Forma convexa.—Texas (Prairie Paso, Doctor Cooper). Forma levis.—Utah (Salt Lake, June, Coll. Hubbard and Schwarz). State of Washington (Walla Walla, Charles Fuchs). Diagnostic characters —Hispilabris is recognized from the mem- bers of the Obscura Group by the prominent dentiform apical angles of the prothorax and by the simple tarsi; from all species of the subgenus /’/eodes by the more or less deeply sulcate elytra with con- vex intervals, and in never being caudate. I consider the typical form to have more or less depressed elytra, the latter with the intervals feebly to moderately convex and fre- quently reddish along the suture; the integuments are generally thin- ner than in the other forms. The five incipient races which I have recognized are simply ex- tremes along particular lines of divergence in the ample series before me, and they are all united into one continuous series by an abundance of mesotypes. They ought to be readily recognized by the brief diagnoses that I have given. Their salient characteristics may be epitomized. as follows: Nupta with its strongly convex and more or less ventricose elytra. Integuments as in typical hispilabris. Sculptilis with the elytral intervals very strongly and prominently convex, with the sulci deep and opaque. Denser integuments. Elongata by its elongate and moderately convex form, elytral intervals more or less moderately convex. Black and denser integu- ments. | REVISION OF ELEODIINI—BLAISDELL. 293 Convexa by its elongate form. Elytra with subconvex and rugose intervals. Levis by its robust convex form; integuments black and aluta- ceous; large prothorax and somewhat eroded sculpturing. Composita Casey has evidently the elytral sculpturing of forma sculptilis and the general structure of Aispilabris, except that it has an aberrant prothorax. But here the difference is no proof that it is specifically distinct. Analogous aberrations are observed in dentipes in particular, and Aispilabris is fully as variable as that species. The unique described by me as interrupta in the subgenus JMelane- leodes was simply an example of symmetrical fortuitous variation of sculpturing, and was an anomalous oméssa. I must hold that the present form, which is a unique and undoubtedly related to a very variable species, is an aberration, and the collection of others like it must give it a greater claim to more than an incipient race. The mentum is moderate in size and quite constant in form, and more or less trapezoidal; the sides are quite straight and converging to the apex, the latter distinctly truncate, or slightly arcuate; the sides are not noticeably curved. In fact, I have not yet seen an ex- ample with the middle lobe evenly rounded anteriorly. The prosternum is variable. Usually moderate in width behind the equator of the cox, evenly convex, or precipitously so behind; fre- quently submucronate, vertically truncate, and feebly compressed behind in nupta. Mesosternum variable; oblique to arcuately declivous and more or less concave. The metasternum laterally between the coxe is about equal in length to the width of a mesofemur at base. The abdominal process is subquadrate, slightly transverse, and in width about equal to the length of the post-coxal portion of the same segment; second segment slightly longer than the post-coxal part of the first and about twice as long as the fourth; the third is about one-third of its length shorter than the second. The abdominal salient is about one-fourth of its width wider than the metasternal process. It is to be observed that the elytral apices have a tendency to be slightly produced, and especially in forma sculptilis, and that the epipleure dilate at tip to form the inferior surface of this feeble but evidently produced elytral apex. This character becomes of impor- tance when taken in relation with the caudal characters of caudifera and longipilosa. The tibial grooves of the femora are not usually quite entire and rather narrow. Those of the profemora may have their margins feeble or strongly defined; the floors are glabrous and rather flat or 2294 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. feebly concave. The margins are scarcely arcuate, and converge a short distance from the femoral base; the anterior margin is dentately laminate at about the junction of the outer fifth and sixth sevenths. The grooves of the meso- and metafemora are more or less defined by sharp margins, which converge to become contiguous somewhere near the inner third, less frequently near the middle. The tibiz are variable. Protibizx more or less feebly compressed, obsoletely to distinctly carinate externally; the external edge scarcely arcuate. The tarsal grooves are sometimes feebly marked and asperate, or at other times quite well defined and more or less gla- brous. The articular cavities are scarcely open, or feebly so at the most. The articular cavities of the meso- and metatibiz are usually quite closed. The tarsal grooves of the mesotibiz are sometimes defined for nearly the whole length of the external surface, floor of each quite flat and more or less glabrous; usually the grooves are poorly defined and asperate. The external surface of each metatibia is more or less flattened and the groove generally feebly defined. The tarsi vary in stoutness according to the general habitus of the individual. In some examples of the typical form the tarsi appear compata- tively longer than in the more robust forma lwvis. In the typical form a protibia is 6 mm. long and its tarsus 3.5 mm. in length; in forma levis the protibia is 6 mm. and the tarsus 3 mm. in length. The actual lengths of parts are of no value, but only when taken relatively. It appears as if the shortening is done at the expense of the length alone. In the typical form joints two, three, and four are evidently longer than wide; in forma lwvis the same joints are distinctly wider than long. In each instance they are subequal to each other, and combined equal to the length of the fifth. The first joint is about one-half longer than wide. In the typical form the protarsi are three-sevenths of their length shorter than a mesotarsus and in the forma levis one-third shorter. The mesotarsi in the typical form are about one-ninth of their length shorter than a metatarsus—to the unaided eye they appear subequal. Joints two, three, and four are longer than wide and shghtly decreasing in length in the order named; together they are a little longer than the fifth; first joint about equal in length to the second and third taken together. In forma levis the meso- and metatarsi are subequal, and the meso- tarsi have joints two to four, inclusive, quite equal. REVISION OF ELEODIINI—BLAISDELL. 995 Forma typica has a metatarsus half as long as a metatibia; in forma levis it is distinctly less than one-half as long. In the former type form joints two and three are subequal in length and about one and one-half times longer than wide, and together equal to the fourth ; the first joint is a little shorter than the fourth. In forma levis the relative lengths are practically the same. Tt appears that with each sight variation in size the relative pro- portions between the tarsi and their joints also correspondingly vary, and at the same time the essential specific characters were homoge- neous in the examples tested. Of course, these variations must be within normal specific limits. These facts become taxonomic pabulum for those students who recog- nize a new species every time certain structural elements present slight differences in their relative proportions. There are times, of course, when these characters are of great value, but specific instabil- ity must determine their value in a given case, and normal specific variations must be recognized. ELEODES CAUDIFERA LeConte. Hleodes caudifera LEContr, Proc. Acad. Nat. Sci. Phila., 1858, p. 184.— Horn, Trans. Amer. Phil. Soec., XIV, 1870, p. 314. Elongate, ovate, caudate. Head about twice as wide as long, feebly convex and more or less impressed laterally along the frontal suture; surface irregularly and rather finely punctate, sparsely so with impunctate areas at center, and densely punctate laterally. Antenne moderate, scarcely com- pressed, last four joints very slightly and gradually dilated, third joint very slightly longer than the fourth and fifth combined; fourth to the seventh, inclusive, subequal; eighth shghtly shorter and broader; tenth and eleventh about as wide as long, the latter ovate. Pronotum widest in front of the middle, about one-third wider than long and about twice as wide as the head; disc rather shining, quite smooth, moderately convex, and slightly declivous at apex and basal angles, more or less impressed and opaque laterally along the margins in the anterior two-thirds, finely and very irregularly punc- tate with impunctate areas at middle; laterally the opaque area is more or less transversely rugulose and granulate; apex feebly emarginate or truncate, very finely or obsoletely margined; sédes evenly and quite strongly arcuate in anterior one-half, quite straight or feebly sinuate and convergent from middle to base, marginal bead fine or obsolete; base slightly arcuate and very finely margined, wider than apex; apical angles rounded, obtuse or minutely denti- form and everted ; basal angles obtuse. 59780—Bull. 68—09——15 226 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Propleure smooth and opaque, rugulose, with few scattered” granules, especially upon the acetabular convexities. Elytra oval, shining, and more or less suleate, less than twice as long as wide—apical prolongation not included—widest at middle and truncate at base, where they are slightly wider than the contigu- ous prothoracic base; humeri obtuse and more or less rounded; sides evenly arcuate, apex horizontally produced; disc moderately convex, slightly flattened and arcuately rounded laterally, obliquely declivous posteriorly passing into the cauda; surface striate, striz more or less impressed and strongly punctate, the punctures muricate and closely placed, intervals more or less convex but not strongly so, uniseriately punctate, the punctures distantly placed and muricate, those of the outer intervals more or less confused and at times rather densely and muricately subrugoso-punctate; inflexed sides obsoletely striate and quite densely and irregularly muricately punctate and at times slightly concave at base of apex. Cauda moderately long, granulo-muricate, above with the suture impressed, tip emarginate; beneath concave, sparsely muricato- granulate, the surface of each lateral half being a dilatation of the apical epipleural surface of the corresponding side; in the median line the two inferior epipleural margins are contiguous. Epipleure rather narrow and gradually narrowing from base to apex, where they dilate to become the inferior surface of the cauda; surface opaque, smooth, and sparsely muricately punctulate, except at base. Sterna more or less irregularly, submuricately punctate and rugulose. Parapleure more or less irregularly punctate. Abdomen horizontal, rather finely, sparsely, and evenly submuri- cately punctate. Legs moderately long and more or less slender. Femora rather — thickly clothed with moderately long appressed setee, with an occa- | sional long flying hair on the external surface. Tibie densely muri- cate and rather sparsely set with long flying hairs; anterior spurs quite equal in length and size and more or less backwardly curved. Tarsi similar in the sexes and clothed with rather long hair-like setee, those on the last joint of each tarsus projecting noticeably be- yond the ungues. Male—Somewhat narrow. Antenne reaching shghtly beyond the prothoracic base. Elytra rather gradually and obliquely declivous posteriorly, becoming sinuate in passing into the horizontal cauda; the latter may be equal to about one-sixth of the entire elytral length. Abdomen feebly ventricose to the fourth segment. Anterior femora with a broad and very obtuse tooth or mutic, REVISION OF ELEODIINI—BLAISDELL. Bor Female.—Rather robust. Antenne reaching scarcely to the protho- racic base. Elytra somewhat suddenly obliquely declivous posteriorly and sinuate at base of apex. Cauda equal to about one-eleventh of the entire elytral length. Abdomen quite strongly ventricose to the fourth segment. Anterior femora quite mutic. Measurements without and with the cauda—Males: Length, 21 and 24; 25 and 29.5 mm.; width, 8-10 mm. Females: Length, 22 and 94; 26.5 and 29.5 mm.; width, 9.5-12 mm. Genital characters, male.—Kdeagophore of the usual oblong-ovate form. | Basale short and oblong, subequal in length to the apicale. Apicale elongate and suboblong-triangular, moderately convex and slightly flattened, with a linear membranous groove in middle third; sides parallel in basal half, thence arcuate, becoming slightly sinuate at apex, the latter acute and slightly produced; base arcuate, scarcely sinuate laterally. Sternite subquadrate. Each lobe about twice as long as wide, broadly rounded at apex; external margin nearly straight; internal margin rather arcuate; surface glabrous, not very thickly punctate and setose, sete moderately long. Female. ena segment Ree subtruncate, length about equal to the width at base. Valvula (Plate 4, fig. 7).—Dorsal plate oblong with the surface broadly concave, margins reflexed and nearly parallel; surface finely punctured in apical two-thirds, each puncture bearing a short semi- recumbent seta; external border nearly straight; internal margin slightly sinuous: apical margin more or less oblique, asperate, set with moderately long and evenly distributed sete, which are not longer at the angle, the latter more or less rounded and never promi- nent. Apex membrano-chitinous, scarcely longer than the append- age, acute; surface set with fine setee, apical tuft rather short. 7 Appendage chitinous, conical, quite long and nearly as long as the apex, blunt, and asperate in apical half, set with rather long sete, tuft at tip quite long. Superior pudendal membrane scarcely reaching one-third from the base. Ventrolateral surfaces—Body moderately convex but not notice- ably inflated, slightly and transversely concave before the apices, finely punctate and set with very fine, short sete in apical two-thirds. Submarginal groove obsolete. External margin of the dorsal plate not prominent laterally. Genital fissure long and fusiform, with the inferior pudendal membrane visible in basal half. Habitat—New Mexico (Aztec, April, C. F. Baker; Coolidge, Coll. E, C. Van Dyke and Hubbard and Schwarz; Santa Fé, C. V. Riley; 228 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Luna, Hubbard and Schwarz) ; Arizona (Adamana, May, H. S. Bar- ber; Winslow, July, Coll. Barber and Schwarz; Holbrook, Coll. E. C. Van Dyke, Charles Fuchs, and Beverly Letcher) ; Colorado (Henry Ulke; Eastern, George Horn); Texas (El Paso, March, Coll. H. Soltau). Number of specimens studied, 40. Type in the LeConte collection. Ty pe-locality—New Mexico; collector, Mr. Fendler. Salient type-characters.—Thorax scarcely convex, slightly punc- tate, laterally a little concave and transversely rugose; anteriorly broadly emarginate, apical angles not acuminate; sides strongly rounded, narrowed posteriorly; posterior angles obtuse. Elytra - slightly flattened, substriate with seriate punctures, intervals uni- seriately and sparsely punctate, at the sides and apex muricate, pos- teriorly obliquely declivous and prolonged. Anterior femora mutic (female) (LeConte). The following forms are before me: Forma glabra.—More or less shining and glabrous, not at all muri- ‘at, except on the cauda. Elytra striate, strize impressed, with quite coarse and closely placed punctures; intervals moderately convex, with a single series of small, distantly placed punctures. Legs quite slender. Cauda gradually narrowing from base to apex. Measurements——Male: Length, 25 mm.; with cauda, 31 mm.; width, 9.5 mm. Female: Length, 20 mm.; with cauda, 22.5 mm.; width, 9 mm. Habitat—Texas (El Paso), Arizona. Forma sublevis.—Subopaque to feebly shining. Antenne slender and rather long. Elytra with series of unimpressed punctures; in- tervals flat with a series of distantly spaced punctures, which are as large as those of the striz. Sculpturing submuricate. Legs rather slender. Measurements.—Female: Length, 25 and 28 mm.; width, 10 mm. Habitat— Arizona. Forma typica.—See salient type-characters. Forma scabra.—More or less shining. Elytra with unimpressed strive of very closely placed and minute muricate tubercles; intervals with a single series of distantly placed and larger muricate tubercles, which become distinctly larger laterally and more tuberculiform. In- flexed sides of the elytra rugoso-muricate. The general surface is very minutely rugulose. Size of forma typica. Legs rather short. Habitat—New Mexico (Aztec). Diagnostic characters——Both sexes are caudate. Thorax widest just in advance of the middle; body not noticeably pubescent, tibiee and tarsi with long flying hairs. These characters serve to differen- tiate caudifera from sponsa, hispilabris, and longipilosa, ] ' EO teas 2 tt ss - REVISION OF ELEODIINI—BLAISDELL. 229 Doctor Horn in the Revision failed to mention that the female was caudate. Forma glabra is a rather striking variation, on account of the smooth integuments and in having the cauda longer than usual in the male, and moderately arcuately decurved in its apical third, being gradually narrowed from base to apex. Forma sublevis has the sculpturing of sponsa, and forma scabra has the elytra tuberculo-scabrous. The mentum is comparatively rather small and triangulo-trapezoidal in outline. The sides con- verge anteriorly and are scarcely at all arcuate; the apex is more or less rounded and the surface is more or less foveate as well as coarsely punctured. The mentum and sides of the epistoma are set with rather long black hairs. The prosternum is variable, moderately wide between the coxe and somewhat triangularly dilated behind the equator of the coxe; more or less convex antero-posteriorly, or feebly convex to subhori- zontal between the acetabula; more or less moderately produced and compressed, oblique to vertically truncate behind, at times sub- mucronate. The mesosternum is variable as to the degree of obliquity and more or less concave. The abdominal process is subquadrate, slightly wider than long and about as wide as the post-coxal portion is long, the latter equal in length to the second segment; the third is twice as long as the fourth, and about one-third of its length shorter than the second. The abdominal salient is about one-seventh of its width wider than the metasternal process. The metasternum laterally between the cox is about as long as a mesofemur is wide at base. The tibial grooves of the femora are not entire, the floors are glabrous, quite plane, and limited by rather well defined edges. Those of the profemora are moderate in width; the anterior margin is scarcely dentately laminate at the outer third in the male and less so in the female; the posterior margin is obsolete internally before becoming contiguous with the anterior, neither one reaching the femoral base. The mesofemoral grooves are obsolete atthe internal third of the femora and the margins do not become contiguous; on the meta- femora they scarcely extend basally beyond the middle, where they become evanescent before becoming contiguous. The protibiz are somewhat compressed and distinctly carinate externally; the tarsal grooves are scarcely developed and muricate. 230 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The mesotibizwe are feebly compressed and the external surface is ridge-like, and the tarsal grooves are evident, although more or less feeble and muricate. The metatibiz are flattened externally, obsoletely grooved, and muricate. Both the meso- and metatibiz are more or less carinate in basal third externally. The tarsi are quite stout. The protarsi are scarcely one-third of their length shorter than a mesotarsus. Joints two, three, and four are subequal, just a little wider than long, and together about equal to the fifth; the first is a little longer than wide. The mesotarsi are about one-seventh of their length shorter than a metatarsus. Joints two, three, and four are subequal, a little longer than wide, and together just the least longer than the fifth; the first — is about one-half longer than wide. The metatarsi are scarcely one-half as long as a metatibia. Joints two and three are subequal, distinctly longer than wide, and together scarcely as long as the fourth; the latter is a little longer than the first. ELEODES LONGIPILOSA Horn. Eleodes longipilosa Horn, Trans. Amer. Hnt. Soc., XVIII, Feb., 1891, p. 42, pier, fie: As: Elongate oval, moderately shining, surface sparsely clothed with long, black, flying hairs; caudate. Head twice as wide as long, more or less moderately convex, coarsely and more or less irregularly punctate, hairs long and flying. Antenne moderate in length and thickness, scarcely compressed or dilated in outer four joints, third joint equal to the next two taken together, fourth to the eighth inclusive subequal, seventh conico- triangular, eighth triangulo-quadrate, ninth and tenth suborbicular, eleventh ovate and pointed; clothed with rather long hairs. Pronotum widest at or slightly in advance of the middle, about one- fourth wider than long, and twice as wide as the head; désc mod- erately convex, coarsely, irregularly, and more or less sparsely punc- tate, rather narrowly opaque and granulate along the sides, fre- quently with impunctate areas at center; apex slightly emarginate, finely or more or less obsoletely margined; sides arcuate in front (occasionally subangulate) and obliquely converging posteriorly, marginal bead fine or obsolete and replaced by minute granules; base more or less arcuate, finely margined, one-tenth to one-fourth wider than apex and about equal to the length; apical angles small, sub- acute, not everted or subdentiform and feebly everted; basal angles obtuse and scarcely rounded. Propleure sparsely muricate and rugulose. * | REVISION OF ELEODIINI—BLAISDELL. Jot Elytra elongate oval, widest at middle and more attenuated pos- teriorly; base truncate or feebly emarginate, shghtly wider than the contiguous prothoracic base; hwmeri obtuse and more or less rounded ; sides evenly arcuate and slightly sinuate at apex, the latter pro- duced, forming a moderately long, horizontal, or shghtly down- wardly oblique cauda; disc feebly convex on the dorsum, rather strongly rounded laterally, and somewhat gradually, arcuately de- clivous posteriorly ; surface more or less striato-punctate, very vaguely suleate, punctures near the suture scarcely muricate, more strongly so laterally and on the apex, serial punctures of the strize moderate in size and closely placed, the interstitial series similar and more dis- tantly spaced, both series becoming irregular laterally and on the apex; suture impressed on the cauda, the latter emarginate at tip; each puncture with a long flying hair. Epipleure moderate in width and gradually narrowing from base to apex, where they dilate to form the inferior surface of the cauda; surface opaque and obsoletely punctulate. Sterna more or less punctate and rugose, sparsely clothed with long brownish hairs. Parapleure coarsely punctate and not noticeably pubescent. Abdomen horizontal and more or less ventricose in both sexes, quite closely punctate, most densely so on the first and fifth segments. Legs moderate in length and thickness, densely sculptured and clothed with long, flying, sparsely distributed hairs. Anterior femora mutic in both sexes; anterior tibial spurs acute, subequal, moderate in length, and more or less curved. Tarsi similar in the sexes. Male.—Somewhat narrower than the female. Antenne reaching to the prothoracic base. Elytra shghtly narrowly oval. Abdomen very feebly flattened on first two segments. Female.——Rather robust. Antenne scarcely attaining the pro- thoracic base. Elytra quite broadly oval. Abdomen strongly convex. Measurements without and with the cauda—Males: Wength, 19 and 21.5; 19.5 and 22.5 mm.; width, 7.2 mm. /emales: Length, 19 and 21; 20 and 22.2 mm.; width, 8.2 mm. Genital characters, male-—Kdeagophore of the usual oblong-ovate form, rather elongate, and scarcely arched. Basale rather short oblong, evenly convex; sides feebly arcuate. Apicale elongate, nearly as long as the basale, and triangular; surface quite evenly and moderately convex, with a narrow median and more or less obsolete groove in basal two-fifths; sides moderately arcuate in basal half, thence more or less sinuate to apex, the latter produced and acute, not deflexed; base broadly rounded at middle, and more or less sinuate laterally. 232 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Sternite quadrato-parabolic and slightly transverse. Lobes slightly elongate, each with the external border feebly sinuate in basal half, thence evenly and rather broadly arcuate to apex, the latter rounded; internal border membrano-chitinous and not well defined in basal half; surface more or less convex, from the middle gradually sloping to the sinus, rather coarsely punctate and setose; sete rather long at apex and moderately dense; membrane not setose across the bottom of the sinus; the latter broad. Female.—Genital segment somewhat equilaterally triangular, shehtly flattened, and setose. Valvula (Plate 4, fig. 23).—Dorsal plate oblong oval; sides more or less reflexed; surface more or less concave, smooth, and shining; finely and sparsely punctate, each puncture with a fine and rather short seta; external margin more or less feebly arcuate; apical margin feebly arcuate, angle scarcely evident; internal margin more or less arcuate, feebly sinuate in basal half; apex short, triangular, subacute at tip, and finely setose. Valvular membrane visible in apical half. Appendage about as long as the apex, conical, with a loose tuft of fine and not very long hairs at tip. Superior pudendal membrane finely and not closely longitudinally rugulose, not quite attaining the middle of the dorsal plate. Ventrolateral surfaces——Body evenly but not strongly convex, and not transversely concave before the apex, shining and_ glabrous, finely, sparsely punctate and setose; submarginal groove obsolete; internal margins of the valves contiguous in basal fifth. Genital fissure long, fusiform, and rather wide, closed by the inferior puden- dal membrane in basal half, apically by the valvular membranes. Habitat—Nevada (Humboldt Valley, Mr. Krause), California (Amedee, Lassen County, July, elevation 4,200 feet., H. F. Wickham). Number of specimens studied, 16. Type in the Horn collection. Type-locality—Humboldt Valley, Nevada; collector, Mr. Krause. Salient type-characters.—Clothed with long, black, flying hairs. Thorax about a fourth wider than long; sides arcuate in front. obliquely narrowed posteriorly; disc moderately convex, coarsely, sparsely, irregularly punctate; apex slightly emarginate, anterior angles subacute, but not everted. Elytra with dise feebly convex, very vaguely sulecate; surface irregularly, sparsely muricate, the two grooves nearest the suture coarsely punctate. Elytra prolonged at tip in a cauda. (Male.) (Horn.) Diagnostic characters Remarkably distinct among its congeners on account of being clothed throughout with long black or brownish flying hairs; caudate in both sexes with the anterior femora obso- letely dentate. —— pe Se a he ee ce we oe, Piet lee 6am, 1p eri PS REVISION OF ELEODIINI—BLAISDELL. 2933 The mentum is quite trapezoidal in the females, and also rather broad; triangulo-trapezoidal and smaller in the males. The sides are more or less straight, and converge to apex, the latter arcuate or truncate; surface more or less strongly punctate, scarcely foveate, and set with long hairs. The prosternum is somewhat variable, usually feebly convex and longitudinally grooved between the coxxe, compressed and vertically truncate behind, with the angle more or less prominent and _ sub- mucronate; sometimes oblique and feebly mucronate. Mesosternum scarcely to strongly concave, and more or less arcu- ately oblique. The abdominal process is about equal in width to the length of the post-coxal part of the same segment, and subquadrate. The second segment is longer than the post-coxal portion of the first; the third is half of its length shorter than the second, and about twice as long as the fourth. The abdominal salient is a little wider than the metasternal inter- coxal process. The metasternum laterally between the coxve is as long as the width of a metafemur at base. The tibial grooves of the femora are not entire, their margins are distinct and granular, their floors flat and very sparsely granulate at the sides. On the profemora the sides of the grooves become contiguous a short distance from the femoral base; on the meso- and metafemora they are somewhat contiguous near the middle, thence to base the femora are rounded beneath. On the profemora the anterior margins of the grooves are not visibly laminate. The protibiz are quite distinctly carinate externally ; the meso- and metatibiz are also in basal fourth externally; tarsal grooves on the protibiz evident, but feebly and coarsely sculptured; the grooves are less evident on the mesotibiz and densely sculptured; metatibiz quite circular in transverse section and not grooved, coarsely sculptured. The protibie are slightly compressed, with the articular cavities more or less feebly open; on the meso- and metatibiz the cavities are closed. The tarsi are moderate in length and rather stout, with joints rather closely articulated. The protarsi are about a fifth of their length shorter than a meso- tarsus. Joints two, three, and four rather short and subequal, dis- tinctly wider than long, and together about equal in length to the fifth; the first about as long as wide. The mesotarsi are about equal in length to a metatarsus; joints two, three, and four about as long as wide and subequal, together 234 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. about equal to the length of the fifth; first joint just a little longer than wide. The metatarsi are about one-half as long as a metatibia; joints two and three subequal in length, a little longer than wide, and together shorter than the fourth; the first about a half longer than wide. DENTIPES GROUP, Section B. The present section contains those species that have the integu- ments glabrous and more or less shining, and never distinctly as- perately sculptured. The caudal characters, as already defined, show, without doubt, a genetic relationship not heretofore recognized. As in Section A some of the species are ecaudate, but unlike those in that section, there is here no tendency to hirsuteness among its constituent species; on the other hand there is a tendency to complete non-devel- opment of such epidermal structures; spines and spinules are de- veloped here as in the species of the preceding Section. The anterior tibial spurs are similar in the sexes, acute and mod- erately divergent, at times feebly curved backward. The abdomen is horizontal and the intercoxal salient of the first segment is always wider than the metasternal process. The genital characters are the same in this section as in the pre- ceding one. The species may be tabulated as follows: All of the femora armed. Femoral teeth strongly developed, long and acute; elytra more or less vertically: declivous ‘behind=222.. * Sans e sale eee ee armata, Femoral teeth rudimentary but distinct on the mesofemora and obsolete On: the: meta femora. Se ee ee ee var. impotens. Femoral teeth quite equilaterally triangular, shorter, broader, and less acute; elytra evenly and arcuately declivous behind and more attenuate. militaris. Femoral teeth smaller and obtuse, otherwise as in militaris__var. femorata, Anterior femora alone armed. Thorax strongly constricted at base. Thorax strongly transverse and very strongly rounded at the sides. Dise with sides narrowly impressed and granulate, more or less evenly and moderately convex; elytral apex more or less attenuately pro- duced or caudate 2 2 eae eee et acuticauda, Dise with sides broadly impressed and granulate, less convex; pune- tuation finely muricate; elytra more or less attenuate and not CAUC A TCE Oa ae SRY ae Se eee oe ae ee var. laticollis. Thorax moderately strongly and evenly rounded at the sides; elytra not attenuate behind with apex obtusely rounded, never acute nor pro- Quced: «2 Se ee 2 a a ee ee ee ee dentipes. Thorax feebly constricted at base. Thorax transverse; elytra striato-punctate, series quite closely placed and impressed, intervals more or less convex__-__--~---- subpingwis. la a el Mla a eal a ia i i at i 3 ¢ : 1 7 4 3 REVISION OF ELEODIINI BLAISDELL. 235 Thorax subquadrate, sides not strongly rounded; elytra smooth, not at all muricately punctate. Hivtrale stricomathersclosely placed]. 2-2 — 222 gracilis. IDA mee WTEC INS CO pe ee ee eee var. distans. Thorax subcylindrical, sides evenly and feebly arcuate throughout ; elytra with unimpressed distant rows of nearly simple punctures. subcylindrica. Thorax not constricted at base; males more or less caudate. Elytra strongly ventricose; apical angles of prothorax moderately prominent anteriorly; first joint of the protarsi (male) clothed at apex beneath with a transverse tuft of golden modified spinules. ventricosa, Elytra feebly ventricose; apical angles of prothorax larger, more prom- inent anteriorly, acute and more or less reflexed; first joint of ante- rior tarsi without transverse tuft of spinules_____________ var. falli. Elytra rarely ever ventricose and distinctly striato-punctate. Apical angles of prothorax feeble and not prominent anteriorly. eschscholtzii. Apical angles usually well developed, acute and anteriorly prom- TVG) Garp ee cee ne tet ue artes Sale te Ree a nd ee ed ee var. luce. Elytra not distinctly striato-punctate and elongate oval. Apical angles of prothorax everted_—_-— 2) es tenuipes. ApicalvanglessnoOt-evertedes 2 2 Se ee wickhami. The student must bear in mind that in all probability he will find the above table inadequate as an aid in the determination of all specimens. If the examples be more or less typical there should be no trouble, but among species so polymorphous, puzzling forms will constantly be found. In the text I have given attention to the most marked incipient races as well as to uniques and aberrations when at hand. Genealogy—TVo speculate upon lines of descent of the species con- stituting the present section involves great doubt as to the correctness of the assumptions, but may be a useful working hypothesis. Never- theless I must present my convictions; for there is no doubt in my mind but that the ancestral stock had the femora strongly armed as in armata, and that this species is a direct and somewhat modified modern exponent of that stock. A typical armata does not have the sides of the prothorax sinuate or constricted before the base. Now, I have before me examples of armata which have the sides of the prothorax as strongly sinuate as in any dentipes, and all of the femora are well armed. I advance the theory that all of the species now constituting the present section of the Dentipes Group in our fauna—which includes Lower California as well as all of that territory north of Mexico— have had their origin from the ancestral avmata stock. It may be asked when did this divergence take place? I would answer, first, that which took place in early times, and, second, that which has taken place in more recent times. 236 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. To the first class belong those forms which have had origin from the primitive eschscholtzii, gracilis, and acuticauda rami. The accompanying genealogical diagram will elucidate my meaning. The more recent derivatives of the eschscholtzii ramus are our pres- ent eschscholtzii, luca, and tenuipes-wickhami,; of the gracilis ramus, our present gracilis, distans, and subcylindrica (+); of the acuti- cauda ramus, the present acuticauda, and laticollis. Around each of these more recent forms are the present incipient species and races which render our species so polymorphous. Armata Impotens Femorata Eschscholtzi Gracilis Distans Subcylindrica Laticollis -Ventricosa Fic. 5..—-GENBALOGICAL DIAGRAM OF THE DENTIPES GROUP, SECTION B, SUBGENUS ELEODES. The recent derivatives of armata, are militarts and femorata by the modification of general form and slow atrophy of the femoral teeth; there are edentate forms derived from these species at the present time in Lower California, and two of them are before me. An example of this loss of the femoral teeth is our impotens. I have studied very instructive series of armata from Fresno and Merced counties. ‘These series show all intermediate stages between a typical armata and a typical dentipes. These remarks are sufficient to indicate my ideas on this subject. Distribution.—The species of Section B are found inhabiting six States or regions: Mexico, Texas, New Mexico, Arizona, California, and Lower California. REVISION OF ELEODIINI—BLAISDELL. Table of Distribution. | | POLIS ocse toe Se | Arizona. .| New Mex- | Texas. .| Mexico .|.-.--..--- | | ico. | ; VES CLTISEOLNS See eee eee [epee

a seine We ety REVISION OF ELEODITNI—BLAISDELL. 951— The metatarsi are equal in length to half the length of a metatibia. Joints two and three are subequal and distinctly longer than wide, and together equal to the fourth; the first is sightly shorter. ELEODES DENTIPES Eschscholtz. Eleodes dentipes WscuscuHoutz, Zool. Atlas, Til, 18338, p. 10, pl. xiv, fig. 4.— MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1848, p. 267; Mag. Zool., 1834, p. 31, pl. cxvii.—LeEConTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horwn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. Hleodes prominens Casry, Annals N. Y. Acad. Sci., V, Nov., 1890, p. 401. Hleodes elegans Casry, Annals N. Y. Acad. Sci., V, Nov., 1890, p. 401. Eleodes confinis BLAISDELL, Ent. News, VI, no. 7, Sept., 1895, p. 237. Elongate oval or ovate, smooth, and convex. Head twice as wide as long, feebly convex, very feebly and vaguely impressed laterally, rather coarsely, irregularly and densely punctate. Antenne usually moderate, but variable in length, outer four joints compressed, slightly and gradually dilated, third joint quite equal to the next two combined, fourth slightly longer than the fifth, the latter, sixth, and seventh subequal, eighth subtriangular in outline and very slightly shorter than the seventh, ninth, and tenth trans- versely oval or suborbicular, eleventh trapezoido-ovate. Pronotum widest at middle, about one-third wider than long, less than twice as wide as the head; disc more or less bright and shining, moderately convex, rather finely, sparsely, and irregularly punctate, usually obsoletely impressed, narrowly opaque, and granulo-punctate laterally along the margin in middle four-sixths, rather declivous at apical and basal angles; @pew moderately and rather arcuately emar- ginate, obsoletely margined; sides broadly and evenly arcuate, dis- tinctly sinuate in basal fifth, and briefly subparallel before the angles, more or less finely margined, margin reflexed; base feebly arcuate and very finely margined, slightly wider than the apex; apical angles moderately acuminately produced and more or less everted; basal angles subrectangular or shghtly prominent. Propleure more or less finely and irregularly granulato-muricate with a large smooth area posteriorly. Elytra oval, smooth and shining, slightly more than twice as long as wide, widest at the middle; base feebly emarginate, scarcely wider than the contiguous prothoracic base; huwmeri obtuse, not prominent nor rounded; sides evenly arcuate, apex obtuse and not rounded; disc evenly and rather strongly convex from side to side, more or less gradually arcuately declivous posteriorly ; swrface rather finely punc- tate, punctures arranged in moderately distant unimpressed series and quite closely spaced, intervals with a series of finer punctules, which are more distantly spaced, and become more irregular and denser along the suture; the punctures become more confused and 252 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. very feebly muricate laterally and on apex; sometimes the punctures are subequal and diffuse. Epipleure moderately wide at the humeri and thence gradually narrowing to apex; surface very finely and obsoletely punctate. Sterna more or less densely punctate and rugulose. Parapleurw rather coarsely punctate. Abdomen horizontal, rather finely and evenly punctate, rugulose, the punctuation denser on the fifth segment. Legs variable in length, form and stoutness. Anterior femora armed in both sexes; tibial spurs similar in the sexes, the anterior equal and acute; tarsi similar in the sexes. Male.—Elongate, subfusiform oval, and somewhat narrow. 3 ¥ REVISION OF ELEODIINI—BLAISDELL. 263 about equal in length to the post-coxal part of the first; the fourth is half as long as the third. The abdominal salient and that of the metasternum are subequal in width. The metasternum laterally between the coxe is about equal in ' length to the width of a mesofemur at base. The tibial grooves of the femora are strongly developed and de- fined; the margins are more or less cariniform; the floors are gla- brous and more or less concave. The profemoral grooves nearly attain the femoral base, while those of the meso- and metafemora become more or less obsolete at the basal fourth or third, respectively. The anterior margin of each groove is more or less laminate, and dentately so at about the outer fourth. The femora are distinctly sinuate external to the teeth, and the anterior surface is longitudinally and inwardly impressed from the bases of the teeth. It must also be noted that the metatibie, while arcuate in basal half, may also be straight and slightly widened in the apical half (see wickhami). The tibie are distinctly arcuate in basal half. The articular cavities are more or less open and the tarsal grooves more or less well developed, although rather narrow and usually defined by asperate margins. The floors are quite smooth. The protibize are distinctly carinate externally, and the meso- and metatibie are scarcely at all carinate externally. The tarsi are usually quite stout and moderate in length. The protarsi are about a sixth of their length shorter than a meso- tarsus. Joints two, three, and four are short, subequal, and distinctly wider than long, and together distinctly shorter than the fifth; the first is slightly longer than the second. Each mesotarsus is about a seventh of its length shorter than a metatarsus. Joints two, three, and four are subequal and evidently just a little wider than long, and together about equal to the fifth; the first 1s about as long as wide. The metatarsi are distinctly less than half as long as a metatibia. Joints two and three are subequal and longer than wide, and together decidedly shorter than the fourth; the first is about one-half longer than wide. ELEODES ARMATA var. IMPOTENS Blaisdell. Elongate, subovate, convex, and moderately shining. Head about twice as wide as long, moderately convex, irregularly and rather sparsely punctate, punctures moderate in size and denser at the periphery. Antenne short, moderately slender, subclavate, outer four joints compressed and rather moderately dilated, third joint equal to the next two combined, fourth slightly longer than the 264 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. fifth, the latter, sixth, and seventh equal and cylindrical, eighth tri- angular, ninth and tenth transversely oval, eleventh ovate. Pronotum widest a little in front of the middle and slightly wider than long; disc evenly and a little more than moderately convex, more or less declivous at the apical angles, finely and sparsely pune- tate; apex truncate to feebly emarginate, finely margined laterally; sides evenly arcuate in the anterior two-thirds, thence less so or some- what straight and converging to base, sometimes feebly sinuate, very finely margined; base feebly arcuate, not coarsely margined and shehtly wider than the apex; apical angles small and acutely denti- form, at times strongly everted; basal angles subrectangular and not prominent. Propleure opaque, more or less finely and very sparsely punctate, frequently the acetabular convexities are strongly rugulose. Elytra oval and more cr less elongate, smooth, widest at the mid- dle; base truncate or feebly emarginate, and about equal to the con- tiguous prothoracic base; humeri obtuse and not prominent; sides evenly arcuate, apex obtuse; disc evenly and quite strongly convex or somewhat depressed, more or less arcuately declivous posteriorly ; surface finely punctate, punctures closely and quite regularly placed in the rather close series, intervals with a single series of more dis- tantly and regularly placed punctures, that often become larger in the outer intervals; both more irregular and denser at the periphery. Epipleure moderately wide and gradually narrowing from base to apex; surface obsoletely and sparsely punctulate. Sterna usually rather finely punctate and frequently quite strongly rugose. Parapleurw rather opaque, more or less sparsely, irregularly, and ‘rather coarsely punctate. Abdomen finely and sparsely punctate, punctures denser on the fifth segment, the first frequently rugulose. Horizontal. Legs slender to moderately stout, and also moderate in length; femora quite densely punctate, and all armed; the anterior with an acute tooth, the middle with a smaller and obtuse salient, the pos- terior with a still smaller and very obtuse salient; tibial spurs rather slender and moderately long and acute, the anterior subequal; tarsi similar in the sexes. Male.—Elongate and rather narrow. Antenne reaching to about the posterior fourth of the prothorax. Elytra rather gradually arcuato-obliquely declivous posteriorly. Abdomen moderately con- vex and broadly impressed and with a stronger longitudinal median impression. Posterior tibiz more or less arcuate in basal half. Female.—Flongate and rather robust, antenne usually not attain- ing the posterior fourth of the prothorax. Elytra more or less broadly oval and somewhat suddenly arcuato-obliquely declivous REVISION OF ELEODIINI—BLAISDELL. 265 posteriorly. Abdomen strongly convex, not impressed. Posterior tibie scarcely arcuate. Measurements.—Males: Length, 21.5-28 mm.; width, 7-9.5 mm. Females: Length, 21-28 mm.; width, 7.5-10.5 mm. Genital characters, male—KEdeagophore of the usual depressed oblong-ovate form, and slender. Basale not arched, sides nearly parallel; surface evenly convex, scarcely gibbous. Apicale triangular, slightly elongate, feebly decurved; surface evenly convex, feebly depressed, with a linear groove in apical half and not extending upon the apex; sides evenly and very moderately arcuate; apex very narrowly rounded and subacute; base broadly lobed at the middle, scarcely sinuate laterally. Sternite moderately transverse. Each lobe subparabolic with the external border evenly arcuate and moderately converging to apex; apical margin rather broadly rounded, angle scarcely evident; 1n- ternal border subarcuate; surface feebly convex and rather densely punctate and setose; setee long and quite dense at apical margin. Membrane not setose across the bottom of the sinus. Female. setose. Valvula—Dorsal plate oblong, concave internal border rather strongly reflexed in apical two-thirds and impunctate, longitudinally subcanaliculate external to the same; surface glabrous, finely, evenly, and sparsely punctate and setose in apical two-thirds; sete reclinate and rather short; external border feebly arcuate; internal margin more or less arcuate, sinuate near base; apical margin evenly arcuate, angle rounded. Apex rather long, subacute at tip, not densely punctate, sete small and slightly denser at tip; superior margin of the fossa set with rather long sete, not denser at angle. Appendage flattened, moderately wide, subconico-semiellipsoidal, and covered with rather long sete in apical half. Superior pudendal membrane reaching to the middle of the dorsal plate and longitudinally rugulose. Basal protuberances moderate. Ventrolateral surfaces.—Body slightly inflated, not strongly con- cave laterally, but feebly transversely so before the apices; surface finely and sparsely punctate, setose, setze short, longer on the inferior margin of the fossa. Submarginal groove small; external margin of the dorsal plate slightly prominent laterally. Internal margins of the valves contiguous in basal third. Genital fissure narrowly fusi- form, inferior pudendal membrane visible in the basal half. Variations —Apex small; appendage as long as the apex and sub- conical; apical margin of the dorsal plate less arcuate, with angle Genital segment subtrapezoidal, moderate in size, and 266 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. narrowly rounded. Dorsal plate with a shght antero-posterior con- vexity. Habitat—California (Livingston, Merced County, July, F. E. Blaisdell, Charles Fuchs; Fresno, collection of the California Acad- emy of Sciences, and the collection of E. A. Schwarz; San Joaquin County, D. W. Coquillett). Number of specimens studied, 22. Type in my own collection, female. Ty pe-locality—Livingston, Merced County, California. Salient type-characters—Pronotum moderately convex; disc smooth, very finely sand sparsely punctate; apex truncate; apical angles dentiform, small, acute, and strongly divergent; sides moder- ately arcuate in the anterior two-thirds, thence feebly convergent to base, scarcely sinuate; basal angles almost rectangular and_ not prominent. Elytra with punctures moderately fine, arranged in rather closely placed, feebly impressed series, intervals with similar, sparsely, and irregularly placed punctures. The anterior femora with an acute tooth, the middle with an obtuse process, the posterior with a very small and obtuse salient. Diagnostic characters—Form variable. It differs in a general way from armata by being more slender and usually considerably smaller, and especially in having the teeth of the hind femora rudimentary or obsolete, while those of the mesofemora are distinct or rudimen- tary, and no doubt obsolete at times, and such individuals (forma subedentata) are scarcely separable from dentipes. The thorax may be as in typical armata or more or less constricted at base as in dentipes. Individuals of armata with normal femoral teeth occasion- ally have the prothorax constricted nearly as in dentipes and can be spoken of as forma sinuata. No doubt but that ¢mpotens is a transitional form between armata — and dentipes, as it occurs chiefly at the periphery of the area of distribution of armata. Dentipes appears to be found mainly be- yond the area inhabited by armata. Some examples of ¢mpotens have the general form of gigantea. General observations.—The middle lobe of the mentum is rather large, trapezoidal and slightly broader than usual, apex more or less arcuate, and the surface evenly and feebly convex. The sterna and abdomen as in armata. The articular cavities of the protibiz are more or less open. The other tibiz and femora are as in armata. The tarsi may be slender or stout and moderate in length. The protarsi are scarcely a fourth of their length shorter than a mesotarsus. First four joints subequal, the first apparently as long as wide, the others a little wider than long, and together about equal to the fifth. 7 REVISION OF ELEODIINI—BLAISDELL, 267 The mesotarsi are about equal in length to a metatarsus. First four joints apparently subequal and just a little longer than wide; joints two to four, inclusive, together a little longer than the fifth. The metatarsi are a little less than a half as long as their metatibie. Joints two and three subequal and distinctly longer than wide, and together shorter than the fourth: first joint the least bit longer than the second. ELEODES MILITARIS Horn. EHleodes militaris Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 310. Elongate oval, moderately shining and convex. [ead twice as wide as long, usually moderately convex, and more or less impressed about the eyes, not coarsely and rather sparsely punctate at middle, punctures becoming denser at the periphery. Antenne moderate, outer four joints feebly or not at ‘all dilated, shghtly compressed, third joint nearly twice as long as the fourth, the latter subequal to the fifth, sixth, and seventh individually, the latter and eighth about as wide as long, ninth and tenth suborbicular, eleventh truncate ovate. Pronotum subquadrate, widest at the middle, a fifth to a third wider than long; disc evenly and rather strongly convex, more or less declivous laterally at apex, finely and evenly punctate, punctures but slightly denser at the sides; apew rather deeply emarginate, finely margined; stdes evenly and moderately arcuate, scarcely oblique posteriorly or at times converging to base, finely margined; base truncate or feebly arcuate and not very coarsely margined; apical angles acute, not dentiform and moderately advanced, or rarely sub- dentiform and feebly everted ; basal angles obtuse and not prominent. Propleurw smooth, with a few scattered punctures posteriorly and on the acetabular convexities, more or less rugulose. Elytra elongate oval, smooth, moderately attenuated behind, widest at the middle; base subtruncate or feebly emarginate, and equal to the contiguous base of the prothorax; huwmeri subobtuse, not rounded nor prominent; s¢des evenly arcuate, apex more or less produced and rather narrowly rounded; disc strongly convex, gradually arcuately declivous behind; surface finely punctate, punctures arranged in dis- tant unimpressed series, those of each series rather closely placed, intervals with a single series of distantly placed punctures, punctua- tion becoming denser and irregular at the sides and apex. Epiplure rather wide and gradually narrowing from base to apex; surface smooth and obsoletely punctate. Sterna not densely and more or less coarsely punctate; prosternum anteriorly finely and sparsely punctate and more or less transversely rugulose. Parapleure moderately sparsely and not coarsely punctate. 268 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Abdomen horizontal, rather finely and sparsely punctate, denser on the fifth segment. Legs moderate, sometimes quite robust; all of the femora armed with an acute triangular tooth in both sexes, and coarsely, densely punctate; all of the tibiz more or less arcuate in basal half; tibial spurs acute and subequal on their respective tibiae; tarsi similar in the sexes. Male—Antenne reaching to the posterior fifth of the prothorax, the latter with base one-half wider than the apex and a fourth greater than the length, a third wider than long. Abdomen moder- ately convex, broadly and not strongly impressed. Femora robust, very coarsely and rather strongly punctate, punctures often deeply impressed. Sometimes the femoral teeth are very strong. Female.—Antenne scarcely reaching to the posterior fifth of the prothorax, the latter with the base about equal to the length, about a fifth wider than long, and a fourth wider than the apex. Abdo- men evenly and strongly convex. _ Measurements— Male: Length, 22.5-28 mm.; width, 7.5-9.5 mm. Female: Length, 22-25 mm.; width, 7.5-8.5 mm. Genital characters, male——KEdeagophore of the usual flattened oblong-ovate form, and somewhat oblong. Basale with the sides scarcely parallel, about two and a half times longer than wide, shghtly or not arched, very strongly and quite ab- ruptly convex from side to side, with rather a strong antero-posterior convexity, the disc appearing gibbous at middle. Apicale triangular, shghtly elongate and decurved apically; sur- face evenly and moderately convex, and with a linear groove in apical half, that does not extend upon the apex; sides arcuate at base, feebly sinuate towards apex, the latter shghtly produced and nar- rowly rounded at tip; base broadly lobed at middle, scarcely sinuate laterally. Sternite moderately transverse. Each lobe subtriangular; exter- nal border nearly evenly arcuate; internal margin nearly straight, with apex rounded; surface strongly punctate in apical half, setose, setee not dense, and moderate in length. Sinus rather broad, mem- brane not setose across the bottom of the same. Female.—Genital segment trapezo-triangular, moderate in size and rather strongly setose. Valvula (Plate 1, fig. 20).—Dorsal plate oblong, concave, sides evenly reflexed; surface finely, evenly, and not very densely punctate throughout, sete short and reclinate; external border quite straight ; internal border more or less sinuous; apical margin more or less arcu- ate, angle obtuse. Dorsal margin of the fossa conspicuously fringed with rather long sete, those at the angle a little longer and denser. Apex moderate and triangular, tip acute with few short hairs. REVISION OF ELEODIINI—BLAISDELL. 269 Appendage flattened, semi-ellipsoidal, comparatively large, punc- tate, and strongly setose at apical moiety, sete long. Superior pudendal membrane attaining the middle of the dorsal plate and longitudinally rugulose. Basal protuberances rather well developed. Ventrolateral surfaces —Body not inflated and very feebly con- cave laterally and before the apex; finely and sparsely punctate, setose throughout, setee small and reclinate; inferior margin of the fossa with a few rather long sete. Submarginal groove not well de- veloped, dorsal plate scarcely expanded laterally. Internal margins of the valves contiguous in basal third. Genital fissure narrowly fusiform, closed in basal half by the inferior pudendal membrane. LHabitat—Lower California (collection of Hubbard and Schwarz; Comondu, San José del Cabo, San Quintin, collection of California Academy of Sciences; Santa Margarita Island, March, collected by Charles D. Haines, collection of Charles Fuchs). Arizona (Charles Fuchs). Number of specimens studied, 12. Type in the Horn collection ; collected by William M. Gabb. Ty pe-locality—Lower California. Salient type-charactcrs.—Male, thorax subquadrate, sides rounded, feebly converging at base; apex rather deeply emarginate, angles acute, not dentiform, basal angles not prominent. Female, thorax a third broader than long, moderately rounded at the sides, not con- verging at base, apex as in the male, base notably broader than apex. Elytra strongly attenuated behind, with distant rows of rather fine and distinct punctures (Horn). Diagnostic characters. —Militaris is separated from armata by the femoral teeth being less produced, broader and more equilaterally tri- angular. The anterior angles of the prothorax are less prominent and less strongly dentiform, everted or not. The elytra are more attenuated behind. The insect appears more obtuse in front on account of the stronger discal convexity of the pronotum. *- * » femoribus * * * breviter dentatis” (femorata). Femorata is quite distinct from armata, the chief differences being the form of the elytra and the femoral teeth—both very varia- ble characters. General observations.—The mentum is subparabolic. In one speci- men it is subtrapezoidal;- surface scarcely convex and not foveate laterally. The prosternum in the specimens at hand is arcuately rounded antero-posteriorly, with a small and abruptly developed mucro be- hind, which may be horizontal or deflexed. Mesosternum as ie armata. The abdominal process is subquadrate, feebly transverse; the post- coxal portion of the first segment is equal to the second in length; the third is as long as the width of the metasternal salient ; the fourth is about one-half as long as the second. The intercoxal process of the abdomen is about a fourth wider than the metasternal salient. The metasternum laterally between the coxe is as long as the width of a mesotibia at apex. The tibial grooves of the femora are well developed and distinctly defined; their floors are opaque and quite plane. Those of the pro- femora are subentire, those of the meso- and metafemora nearly attain the femoral base, and the carinate margins are somewhat eva- nescent internally. The anterior margin of each groove is more or less dentately laminate externally. The protibiz are somewhat compressed and distinctly carinate externally; their tarsal grooves are more or less developed, opaque, and asperately sculptured. The articular cavities are more or less open. | The mesotibiz are carinate externally in basal third; the grooves are more or less indicated, opaque, and muricate. The articular cavi- ties closed. The metatibiz are more or less finely subcarinate externally, and the grooves are scarcely evident, muricately sculptured; articular cavities closed. Both the meso- and metatibie are slightly com- pressed. Tarsi moderate in length and rather stout. The protarsi are about _asixth of their length shorter than a mesotarsus. Joints one to four, inclusive, subequal and a little wider than long; joints two to four combined equal in length to the fifth. The mesotarsi are about a seventh of their length shorter than a metatarsus. Joints two to four inclusive subequal and very little 276 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. longer than wide; together scarcely longer than the fifth; the first joint is about one and a half times longer than wide. The metatarsi are a little less than half as long as a metatibia. Joint two is evidently just the least longer than the third, both are longer than wide, and together distinctly shorter than the fourth, also a little longer than the first. ELEODES ACUTICAUDA LeConte. Fileodes acuticauda LEContTE, Ann. Lye. Nat. Hist. N. Y., V, 185%, p. 135; Proc. Acad. Nat. Sci. Phila., 1858, p. 181.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 314. Var. laficollis LEContTE, Ann. Lyc. Nat. Hist. N. Y., V, 1851, p. 135. Klongate, more or less shining, strongly convex, elytra more or less strongly produced and not suddenly caudate. [ead slightly more than twice as wide as long, quite evenly convex, moderately finely and evenly punctate. Antenne rather stout, mod- erately short, outer four joints slightly compressed, more or less feebly and gradually dilated, third joint equal in length to the next two combined, fifth, sixth, and seventh subequal in length and each shghtly shorter than the fourth, eighth trapezoidal in outline, ninth and tenth wider than long, eleventh subtrapezoidal. Pronotum widest at about the middle, about four-sevenths wider than long and more than twice as wide as the head; dsc more or less bright and shining, moderately convex, noticeably declivous at base and apex, rather narrowly and more or less impressed and opaque laterally, finely, irregularly and somewhat densely punctate, some- times granulate laterally; apex moderately emarginate, obsoletely or distinctly margined; sides very strongly arcuate im anterior two- thirds, thence to base less so or somewhat straight and converging, and strongly sinuate in front of the basal angles, more or less finely margined; base moderately arcuate and rather coarsely margined, a little narower than the apex; apical angles small, acuminately pro- duced and dentiform, very acute and more or less everted; basal angles obtuse, sometimes quite prominent. Propleure smooth, obsoletely punctulate and rugulose. Elytra elongate oval, smooth and more or less shining, widest at the middle; base feebly emarginate and not wider than the contiguous prothoracic base, humert obtuse and not at all prominent; sides evenly arcuate, frequently broadly and feebly sinuate at sides of apex, the latter gradually produced; disc quite evenly and strongly convex from side to side, more or less gradually and arcuately de- clivous posteriorly, or sometimes more or less broadly sinuate before the apex; surface smooth, punctate, the punctures closely placed in longitudinal series, the latter moderately closely placed, intervals ~~ REVISION OF ELEODIINI—BLAISDELL, Gi irregularly and move or less sparsely punctulate, punctures usually not noticeably denser laterally. Elytral suture on apex more or less deeply impressed. Apex or cauda concave beneath, the surface of which is formed by the internal elytral surface. Epipleurw slightly widened at base, thence gradually narrowing to apex, where it narrowly margins the inferior surface of the pro- duced apex or catia; surface smooth and usually impunctate. Sterna more or less punctate and rugose. Parapleure more or less punctate. Abdomen horizontal, obsoletely or rather strongly punctate, rugu- lose. Legs more or less moderate, anterior femora armed in the sexes; anterior tibial spurs equal and acute. Tarsi similar in the sexes. Male.—Elongate. Antenne attaining the posterior fourth of the prothorax. Elytra usually more than twice as long as wide and more or less strongly and gradually produced at apex. Abdomen rather strongly convex, scarcely at all flattened or impressed. Legs some- what stout, especially the anterior femora, which are armed with a very acute tooth; posterior tibiz more or less arcuate. Female.—Robust. Antenne attaining the posterior third of the prothorax. Elytra suboval, and usually less than twice as long as wide, rather moderately obliquely declivous posteriorly ; apex more or less produced. Abdomen strongly convex. Legs moderate; ante- rior femora with a more or less acute tooth; posterior tibixe straight. Genital characters, male—Edeagophore of the usual oblong-ovate form and rather elongate. Basale slightly or not arched, quite evenly convex. Apicale triangular, shghtly elongate, not decurved at tip; surface rather evenly convex more strongly so apically where the sides appear compressed, groove at middle two-fourths, fine toward apex and shghtly dilated toward base; sides feebly sinuate in apical two- thirds; apex rather acute and* gradually narrowed; base broadly lobed at middle, lobe rounded, laterally sinuate. Sternite transverse, sometimes quite oblong, lobes longer than wide. Each lobe with the external border evenly and broadly arcuate apic- ally, quite straight toward base; internal margin nearly straight; apex narrowly rounded; surface punctate and setose. Membrane not setose across the bottom of the sinus, the latter rather broad. Female.—Genital segment trapezoidal, moderate in size, glabrous and setose. Valvula (Plate 3, fig. 7)—Dorsal plate oblong, shghtly concave, sparsely punctate and setose in apical two-thirds, setze moderately short, semirecumbent; external border straight or feebly and broadly sinuate; internal margin quite strongly reflexed in apical half, more 278 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. or less sinuate toward base; apical margin squarely truncate and transverse; angle rectangular or narrowly rounded, with a few rather long setw that are also sparsely distributed along the apical margin. Apex small, subacute, with a tuft of a few rather short sete. Fossa rather large. Appendage quite large, flattened and triangular or semi-elliptical, broad as long, setose toward tip, with a tuft of quite long sete. Superior pudendal membrane attaining the middle of the dorsal plate and quite distantly longitudinally rugulose. Ventrolateral surfaces (Plate 3, fig. 8).—Body slightly inflated at base, rather strongly and transversely concave before the apex, rather strongly and sparsely punctate; sete along the ventral margin of the fossa rather long and flying. Apex quite densely setose beneath. Submarginal groove well developed beneath the projecting external border of the dorsal plate. Internal margins of the valves contiguous in basal third, genital fissure rather broadly fusiform and closed in basal half by the inferior pudendal membrane. Two forms may be recognized as follows: Forma typica—Glabrous. Elytra with striz of small unimpressed punctules or punctures which are frequently not very evident, inter- vals somewhat or very irregularly punctulate. Males dimorphic. The typical males are very strongly but not suddenly caudate; the atypical males with the females have the elytral apices moderately or searcely produced, and not caudate. Measurements.—M ales: Length, 16.5-30 mm.; width, 5-10.5 mm. Females: Length, 24.5-28.5 mm.; width, 8-9.5 mm. Forma punctata—Glabrous. Antenne stout, attaining the pos- terior third of the prothorax. Elytra with serial punctures moder- ately large, often eroded and feebly impressed, the series rather closely placed and often irregularly duplicated, intervals irregularly and finely punctulate, rather more densely so laterally, males apparently homomorphic and with the females*have the elytral apex moderately produced and not strongly caudate. Legs rather stout. Measurements.—Males: Length, 22-28 mm.; width, 8-8.5 mm, Females: Length, 22.5-26.5 mm.; width, 8.5-10.5 mm. Tlabitat.—Forma typica.—California (San Diego County; San Diego, elevation 50-300 feet. F. E. Blaisdell, E. C. Van Dyke, Charles Fuchs, and collection U. S. National Museum; Poway, eleva- tion 700 feet, F’. E. Blaisdell; Los Angeles County, E. C. Van Dyke; Santa Barbara; Fort Tejon, April 19-21, Fuchs and Hopping; Tulare County; San Bernardino County); Lower California (San Pedro Martir, collection California Academy of Sciences). Forma punctata.—California (Los Angeles County, E. C. Van Dyke). Number of specimens studied, 120. a! REVISION OF ELEODIINI—BLAISDELL. 279 Type in the LeConte collection. Type-locality—San Diego, California. Salient type-characters—Thorax finely punctate, broad and greatly rounded at the sides, which are subdepressed, narrowed behind; basal angles obtuse, somewhat more prominent than usual. Elytra strongly convex, finely and seriately punctate, posteriorly declivous, strongly attenuated, and produced (LeConte). Diagnostic characters —The prothorax is usually at least one-half wider than long, sides very strongly rounded and strongly~ con- stricted at base; anterior angles acute, prominent and dentiform, but generally smaller than in /aticollis, and frequently everted. The ely- tra in the typical form are elongate and strongly produced in the males, not suddenly caudate, and directed obliquely downward. In examples from Fort Tejon the males have the elytra distinctly caudate, the cauda horizontal and formed nearly as in ducw. In the other forms the elytra are less elongate and moderately produced. The elytra are never sulcate, but simply with strize of small punc- tures that are at times rather coarse and rarely impressed. The elytral surface is never muricate, and is in this character sepa- rated from /aticollis, the latter having the pronotum more strongly punctate, duller, and rather broadly impressed and granulate later- ally; in acuticauda the pronotum is narrowly or not at all impressed nor granulate laterally. From dentipes it must be separated by the form of the pronotum. In acuticauda the sides of the pronotum are more strongly and less broadly arcuate and the constriction is more rapidly formed and larger; in dentipes the sides of the pronotum are less strongly and more broadly arcuate, the sinuation is shorter and more gradually formed. In some examples of acuticauda the pronotum is not wider than in dentipes, and these are the difficult specimens to place, but the form of the arcuation still remains the same, and a quick eve can readily catch the difference. In forma typica the males may have the find tibiee arcuate in basal half and more or less gradually thickened apically. General observations —The middle lobe of the mentum is moder- ate and rather wide, subparabolic, sometimes subtrapezoidal ‘in out- line, the apex is seldom truncate; surface slightly convex centrally and more or less feebly foveate laterally within the margin. The prosternum is variable, usually convex or horizontal between the coxxe and with a moderate mucro behind; rarely rounded or sub- truncate behind. The mucro may be horizontal or deflexed. Mesosternum is more or less oblique and concave. The abdominal process is quite quadrate and in width equal to the length of the third segment; the post-coxal part of the first segment 280 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. is twice as long as the fourth; the second is a little longer than the post-coxal part of the first. The abdominal salient is about a fourth of its width broader than that of the metasternal process. The metasternum laterally between the cox in length is about equal to a mesotibia at middle. The tibial grooves of the femora are well developed and scarcely entire, the margins are cariniform and become evanescent, usually at the internal fourth; their floors are smooth and quite flat. The anterior margin of the profemoral grooves become quite sud- denly dentately laminate at about the external fourth. The protibiz are more or less distinctly carinate externally and feebly compressed, their tarsal grooves are more or less evident and scabrous and the articular cavities are closed. The mesotibiz have the grooves more or less developed and muri- cate and the articular cavities are closed; externally obsoletely cari- nate in basal fourth. The metatibiz are more or less flattened exter- nally and roughly sculptured, not at all carinate and the articular cavities closed. The tarsi are moderate in length and quite stout. The protarsi are about a third of their length shorter than a meso- tarsus. Joints two, three, and four are quite closely articulated, sub- equal, wider than long and together about equal to the fifth; the first is about as long as wide. The mesotarsi are very little shorter—about an eighth of their length—than a metatarsus. Joints two, three, and four are subequal, as long as wide, and together quite equal to the fifth; the first is shghtly shorter than the next two taken together. The metatarsi are a little less than half as long as a metatibia. Joints two and three are subequal and distinctly longer than wide, together just the least shorter than the fourth, and a little longer than the first. ELEODES ACUTICAUDA var. LATICOLLIS LeConte. Elongate oval, more or less shining and smooth, strongly convex and not caudate, interstitial punctures of the elytra more or less muricate. Head rather large, twice as wide as long, rather moderately con- vex, very feebly and broadly impressed laterally, finely, irregularly and rather sparsely punctate, quite densely so laterally, punctures finely muricate, frontal suture obsolete. Anfennw moderately stout, reaching to the base of the prothorax, outer four joints feebly com- pressed and very shghtly dilated, third joint very shghtly longer than the next two combined, fourth slightly longer than the fifth, the —< ° = Sey EAR ei tte ere lo" od ee ee i ad REVISION OF ELEODIINI—BLAISDELL. 981 latter, sixth, and seventh subequal, eighth triangular to triangulo- conical, ninth and tenth suborbicular in outline, eleventh subtrape- zoidal. Pronotum widest at about the middle, almost a half wider than long; disc moderately convex, more strongly so at the angles, rather finely, distinctly, irregularly and not densely punctate, rather broadly impressed and granulo-punctate laterally; apex broadly and quite evenly emarginate, distinctly margined laterally, and obsoletely so at the middle; s/des greatly rounded in anterior three- fourths, thence strongly sinuate and converging to base, rather coarsely margined; base feebly arcuate, equal to the apex and coarsely margined; apical angles strong, acuminately produced and dentiform, more or less everted, and somewhat reflexed at times; basal angles subrectangular and more or less prominent. Propleurew rather smooth, irregularly and sparsely granulo-muri- cate, obsoletely rugulose. Elytra oval to elongate oval, widest at the middle; dase subtrun- cate, more or less slightly wider than the contiguous prothoracic base; Aumeri obtuse, not rounded nor prominent; sides evenly arcuate, apex gradually narrowed, rarely attenuated and not pro- duced; disc moderately convex on the dorsum and more strongly so laterally, arcuately declivous posteriorly ; surface punctate, punc- tures arranged in rather distant series, moderately small and rather closely placed, intervals with a series of similar punctures, slightly more distantly spaced, on the dorsum very faintly muricate, laterally distinctly so and frequently more distantly spaced, both series be- coming more irregular and finely muricate on the apex, punctures minutely setigerous. EL pipleure moderate at the humeri and gradually narrowing to apex; surface sparsely and finely submuricately and_ obsoletely punctate. Sterna more or less densely, submuricately punctate, and rugose. Parapleure rather coarsely and irregularly punctate, punctures rather shallow. Abdomen horizontal, rather densely, somewhat finely, submuri- cately punctate and rugulose. Legs rather long and somewhat stout; anterior femora armed in both sexes; anterior tibial spurs subequal and acute. Tarsi similar in the sexes. Male—Somewhat narrow. Elytra arcuately and rather obliquely declivous behind. Abdomen moderately convex and slightly flat- tened at middle on the first three segments. Anterior femora armed with an acute tooth. Metatibie more or less arcuate. 282 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Female—Somewhat robust. Elytra arcuately declivous posteri- orly. Abdomen evenly convex. Anterior femora with an obtuse or subacute tooth. Metatibiz straight. Genital characters, male. not arched. Basale oblong; surface moderately convex and the sides quite parallel. Apicale subequilaterally triangular; surface evenly convex, coarsely and sparsely punctate, median groove linear and fine in middle three-fifths; sides feebly arcuate from base to apex, the latter subacute and not produced; base broadly and arcuately lobed at middle, feebly sinuate laterally. Sternite transversely subcrescentiform. Each lobe wider than long; external border broadly and evenly arcuate from base to apical angle, which is introrse at the internal margin, the latter short; angle rounded ; surface moderately convex, quite densely punctate in apical three-fourths; setose, setee quite long and quite dense. Membrane not setose across the bottom of the sinus, the latter short. Female.—Genital segment rather small, setose, and triangularly trapezoidal in outline, moderately short. Valvula (Plate 3, fig. 12).—Dorsal plate oblong with sides shghtly reflexed and moderately concave; surface glabrous, very sparsely punctate and setose, sete moderate in length and somewhat erect ; external border straight or feebly sinuate; internal margin feebly arcuate; apical margin subtruncate, angle narrowly rounded, both fringed with rather long hairs, which are somewhat dense at the angle; apex small and short, triangular, finely setose, with a tuft of shghtly longer hairs at tip. Appendage rather small and conical, equal in length to the apex, finely setose, with a tuft of longer hairs at tip. Basal prominence small and not very evident. Superior pudendal membrane attaining the middle of the dorsal plate; finely but not strongly longitudinally rugulose. Ventrolateral surfaces—Body scarcely swollen in basal half. broadly concave laterally before the apices; setose, sete rather long, especially upon the inferior margin of the fossa and at angle; surface glabrous. Internal margins of the valves contiguous in basal fourth. Genital fissure fusiform, widest in apical half, closed by the inferior pudendal membrane in basal moiety. It is proper to recognize two forms, as follows: ‘longate, usually large and more or less shining. Interstitial punctures finely muricate and moderately distant. the punctures laterally and on the apex more decidedly prickly, each with a very small stiff seta. Elytral apex gradually narrowed and slightly ral, depressed and / REVISION OF ELEODIINI—BLAISDELL. 283 produced in the male; scarcely produced and obtuse in the female. Legs long and rather stout. Measurehents—Males: Length, 24.5-34 mm.; width, 9-10 mm. Females: Length, 26-82 mm.; width, 10.5-10.5 min. Forma minor.—Somewhat robust and more or less shining. Inter- stitial punctures quite distant and very distinctly prickly laterally and on the apex. Sexes quite homomorphic. Elytral apex scarcely produced and more or less abtuse. Legs moderate. Measurements.—Males: Length, 22.5-27 mm.; width. 8-9.5 mm. Females: Length, 21.5-26.5 mm.; width, 9.2-8.5 mm. Habitat—Forma typica.—Califormia (San Diego County, Los Angeles County; collections Charles Fuchs, E. C. Van Dyke, F. E. Blaisdell, and U. S. National Museum; Fort Tejon, April 19-21, Fuchs and Hopping) ; St. Nicholas Island (collection U. S. National Museum). Forma minor.—California (Poway, elevation 700 feet, Ff. EK. Blais- dell; Fort Tejon, Fuchs and Hopping; collection U. S. National Museum). Number of specimens studied, 125. Type in the LeConte collection. Type-locality—San Diego, California. Salient type-characters.—Thorax broad, sides greatly rounded and subdepressed ; narrowed behind; basal angles rectangular, somewhat unusually prominent; anterior angles acuminate. Elytra finely and seriately punctate; apex declivously attenuated (LeConte). Diagnostic characters.—This race 1s separated from acuticauda by the form of the pronotum, elytral apices, and sculpturing. Acuticauda has the pronotum very narrowly granulate and very feebly impressed laterally along the margin; the disc is nearly evenly convex from side to side; in /aticollis the sides are quite broadly impressed and granulate; consequently the surface is less evenly and less strongly convex. In acuticauda the punctuation is simple, while Jaticollis is remark- able for having the interstitial punctures very finely muricate; on the elytra laterally the interstitial punctures are distinctly prickly, on the apex irregular and likewise muricate. In laticollis the elytra are never caudately produced, although at times somewhat gradually narrowed, but never as strongly as in acuticauda. Dentipes forma elongata’ may have the pronotum somewhat im- pressed laterally and more or less granulate, but here the form of the prothorax must be the diagnostic criterion. (See acuticauda and synopsis. ) / 284 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. A series of laticollis from St. Nicholas Island is almost distinctive of another race (forma insularis) or insular form. The pronotum is notably bright and shining, and the general form is rather more elongately robust. General observations.—The middle lobe of the mentum is moderate and subparabolic or subtrapezoidal in outline; the surface is very feebly convex and coarsely punctate, rarely feebly foveate laterally. The prosternum is but moderately prominent ventrally with the cox, evenly arcuate antero-posteriorly and mucronate behind, or subhorizontal with the tip mucronate; frequently deeply and longi- tudinally grooved between the coxee. ; The mesosternum is more or less arcuately declivous and more or less broadly concave. The abdominal intercoxal salient is quite quadrate and in width is a little shorter than the third segment; the post-coxal part of the first is a little longer than the third; the second twice as long as the fourth, and the third is a third of its length shorter than the second. The abdominal process is about a fourth of its width wider than the metasternal salient. The metasternum laterally between the coxe is quite short and in length is about equal to the width of a mesotibia at middle. The tibial grooves of the femora are well developed, and the mar- gins are quite cariniform; the floors are quite flat, opaque, and fre- quently slightly granular, especially on the meso- and metafemora. The profemoral grooves are subentire, the margins slightly evanes- cent before the base, and the anterior margins are dentately laminate near the outer third. The mesofemoral grooves are evanescent at basal fourth, those of the metafemora at basal third. The external surface of the protibiz is more or less distinctly cari- nate and the tarsal grooves are more or less evident, sometimes feebly glabrous toward apex, but generally scabrous; the articular cavities are not strongly closed. The meso- and metatibiz have the articular cavities strongly closed and margined; the tibiz are not carinate. Externally both have the grooves evident in apical two-thirds, with rest of the surface very densely, strongly, and muricately sculptured. The tarsi are quite stout. The protarsi are about two-sevenths of their length shorter than a mesotarsus. Joints two, three, and four are subequal and shghtly wider than long, together about equal to the fifth; first joint about as long as wide. The mesotarsi are about a ninth of their length shorter than a meta- tarsus. Joints two, three, and four are subequal and about as long as wide, together longer than the fifth; the first is about one and a half times longer than wide. REVISION OF ELEODIINI—BLAISDELL. 285 The metatarsi are not quite equal to half of the length of a meta- tibia. Joints two and three are subequal, together shorter than the fourth and longer than the first. ELEODES ESCHSCHOLTZII Solier. Hleodes eschscholtzii Souirr, Studi Entomol., II, 1848, p. 254.—CHAMPIoN, Biol. Centr.-Amer., IV, Pt. 1, 1884, p. 77, pl. 111, fig. 24, male. Var. luce LECONTE, Smithsonian Miscell. Coll., 167, Pt. 1, 1866, p. 114.— Horn, Trans. Amer. Phil. Soe., XIV, 1870, p. 312. Elongate, fusiform oval, smooth and more or less shining, strongly and more or less caudate. Head about twice as wide as long, more or less convex, feebly im- pressed within the sides of the frons, rather finely, more or less irregu- larly, and not very densely punctate. Antenne long and moder- ately slender, outer four joints feebly compressed, gradually and fee- bly dilated, third joint scarcely as long as the next two combined, fourth to the seventh inclusive subequal, eighth subtriangular, ninth and tenth more or less orbicular to feebly oval, eleventh ovate and obliquely truncate at tip. Pronotum quadrate, widest at the middle, an eighth to a fourth wider than long; disc moderately and evenlyconvex, declivous at the apical angles, finely to obsoletely, evenly and rather sparsely punctu- late; apex truncate to more or less evenly emarginate, finely margined, obsoletely so at the middle; s#des evenly and feebly to moderately arcuate, finely margined; base feebly arcuate and finely beaded, and about a fourth wider than the apex; apical angles subrectangular to obtuse and not rounded, or acute and more or less advanced; basal angles obtuse and not rounded. Propleure obsoletely punctulate and more or less rugulose. Elytra oval, widest at the middle; base truncate to feebly emargin- ate, very slightly wider than the contiguous prothoracic base; humeri obtuse, not rounded nor prominent; sides evenly arcuate, apex more or less produced; disc evenly and strongly convex from side to side, obliquely to arcuately and more or less gradually declivous posteri- orly; surface striato-punctate, punctures moderate in size, closely and regularly placed, in moderately distant and more or less im- pressed series, the intervals with a series of irregularly and more or less distantly spaced, very fine punctures; punctures not becoming confused at the sides or apex, the intervals scarcely convex, but becoming slightly so on the apex, where the suture is more or less impressed. ; Epipleure rather narrow and gradually narrowing from base to apex, where they extend narrowly along the sides of the inferior sur- 286 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. face of the cauda or produced apex; surface finely and obsoletely punctulate. Sterna more or less finely, densely, obsoletely punctate and rugulose. Parapleure more or less finely and rather densely punctate. Abdomen horizontal, finely, obsoletely punctulate and rugulose, punctures denser on the fifth segment. Legs quite slender and moderately long, usually not strongly sculp- tured; anterior femora armed in both sexes; anterior tibial spurs subequal and acute; tarsi similar in the sexes. Male.—Elongate and somewhat slender. Antenne attaining a shght distance beyond the prothoracic base, the latter a little greater than the width of the prothorax. Elytra very obliquely declivous, then subhorizontally produced at apex into a long cauda, the latter frequently with the sides more or less converging to tip, which is slightly deflexed. Abdomen quite strongly convex, feebly, longi- tudinally, and narrowly impressed at middle on first three segments. The anterior femora armed with a moderate and acute tooth; pos- terior tibie quite strongly arcuate to slightly beyond the middle, then rather suddenly dilated and straight to apex. Female.—Quite robust. Antenne not quite attaining the protho- racic base, the latter equal to the length. Elytra moderately broadly oval, arcuately declivous posteriorly, apex slightly produced and sub- acute. Abdomen strongly convex. Anterior femora with a more or less obtuse tooth, and the metatibie straight without being widened apically. | Measurements.—M ales: Length, 27-29 mm.; width, 9-9 mm. /’e- males: Length, 24—25.5 mm.; width, 9-9.5 mm. Genital characters, male-—Edeagophore elongately subfusiform and feebly arched. Basale more or less oblong, evenly convex above, and the sides evenly and moderately arcuate. Apicale elongately triangular and very feebly decurved; surface moderately and evenly convex above, with a narrow median longi- tudinal groove; sides feebly arcuate or straight; apex not produced and narrowly rounded; base acutely lobed at middle, broadly and feebly sinuate laterally. Sternite subparabolic in outline. Each lobe triangular; external border more or less evenly arcuate; apex rather broadly rounded; in- ternal margin more or less sinuate; surface glabrous, rather evenly, moderately densely punctate and setose in apical three-fourths, sete moderate in length, becoming long on the apex. Membrane not setose across the bottom of the sinus, the latter rather triangular. Female-—Genital segment robust, subquadrate and strongly chiti- nized, REVISION OF ELEODIINI—BLAISDELL. 287 Valvula.—-Dorsal plate more or less oblong, glabrous and shining, sides very strongly reflexed; surface deeply concave, very finely, sparsely punctate and finely setose ; external border rather strongly arcuate; internal margin more or less sinuous; apical. margin promi- nent, evenly and moderately arcuate, angle rounded. Apex short and triangular; valvular membranes strongly exposed between the apices caudad to the pudendal membrane. Appendage subsemielliptical, rather equal in length and size to the apex. Appendage, apex, and margin of the fossa finely punctate and setose, setee short and inconspicuous. Superior pudendal membrane attaining the middle of the dorsal plate and longitudinally rugulose. Ventrolateral surfaces—Body moderately inflated and glabrous, transversely and broadly concave before the apices, the latter finely punctate and setose. Submarginal groove more or less obsolete. In- ternal margins of the valves contiguous in basal third; genital fissure narrowly fusiform and closed in basal half by the visible inferior pudendal membrane. Habitat—Mexico (Sinaloa, coll. Chas. Fuchs; Alamos, Buchau- Hepburn; Presidio, Forrer). United States (Texas). Number of species studied, 5 (87). Type is probably with the Solier collection. Type-locality—Mexico (Alamos?), Salient type-characters.—Thorax quadrate; disc very finely punc- tulate; apex feebly emarginate; apical angles subacute and slightly prominent anteriorly; sides evenly but not strongly arcuate; basal angles obtuse. Elytra striato-punctate, punctures moderate and closely placed in the series, intervals scarcely Convex and with a series of rather distantly spaced punctules; cauda horizontal. Diagnostic characters—The figure given in the Biologia I take to represent what 1s probably a typical form of this species. I can not see that it differs very much from some of the less strongly de- veloped forms of ducaw. The prothorax is quadrate, with the sides feebly and. evenly rounded from base to apex; apical margin feebly emarginate, angles small, acute, and just feebly prominent anteriorly. In the specimens before me the sides of the prothorax are rather less rounded, apical margin very feebly emarginate or truncate and the apical angles scarcely at all prominent. The general form of the body is rather more slender, with outline more evenly fusiform. The cauda narrows somewhat to tip. I do not believe that a large series could be separated from a simi- Jarly large series of luce. In a series of five examples (4 males and 1 female) mentioned in the Biologia, the largest male measures 35 mm., including the 288 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. caudal prolongation of 6 mm.; similar large males are common in the series of duce taken in Lower California. The hind tibie are curved in basal half, straight and thickened toward apex as in luce. General observations.—The middle lobe of the mentum in the ex- amples at hand is triangulo-trapezoidal, more or less arcuately rounded at apex; surface is coarsely punctate, just noticeably convex and scarcely at all foveate ijaterally. The prosternum in the series under examination is rounded antero- posteriorly, feebly mucronate or not. In the larger examples it would no doubt be strongly mucronate as in duce. Ina large male without locality and in the U. S. National Museum collection, the prosternum is horizontally and triangularly mucronate, exactly as in the large specimens of /ucw,; the specimen measures 33 mm. Two accompanying females have similar prosterna. These specimens are without doubt from Mexico. The mesosternum is the same as in /wew, except that it 1s less ver- tically oblique. The abdominal process is subquadrate and feebly transverse, in width equal to the length of the third segment; the post-coxal por- tion of the first segment is equal in length to that of the second; the third is scarcely twice the length of the fourth and about a third of its length shorter than the second. The abdominal process is about a fourth of its width greater than that of the metasternal salient. The metasternum laterally between the coxe is about equal in length to the width of a mesotibia at apex. The tibial grooves of the femora are well defined, margins carini- form, floors plane, opaque and smooth. Those of the profemore nearly attain the femoral base and the margins become contiguous. The margins of the grooves on the mesofemora become evanescent at the internal fourth before becoming contiguous. On the meta- femora the margins are evanescent at internal third. The meso- and metafemora have the grooves somewhat narrow. On the profemora the anterior margin of each groove is dentately laminate at external fourth. The protibiz are more or less carinate externally and not com- pressed; the articular cavities are more or less open, and the tarsal grooves are more or less developed and scabrous. The meso- and metatibiz are not carinate externally and the artic- ular cavities are closed. The tarsal grooves of the mesotibiz are quite well defined in apical two-thirds; the sides of the grooves are muricate and the floors smooth and opaque. The metatibie are sim- ply flattened externally or feebly grooved and muricate. The tarsi are moderate in length and stoutness. REVISION OF ELEODITNI—BLAISDELL. 289 The protarsi are about a third of their length shorter than a meso- tarsus. Joints two, three, and four subequal, apparently very slightly wider than long, and together about equal to the fifth; the first about as long as wide. The mesotarsi are very slightly shorter than a metatarsus. Joints two, three, and four subequal, scarcely longer than wide, and together just a little longer than the fifth; the first about one and a half times longer than wide. The metatarsi are a little shorter than half of the length of a meta- tibia. Joints two and three subequal, distinctly longer than wide and together scarcely as long as the fifth, a little longer than the first. ELEODES ESCHSCHOULZII var. LUCA LeConte. Elongate, more or less shining, convex and more or less caudate. Head twice as wide as long, moderately and evenly convex, rather finely, quite evenly and not densely punctate. Antenne long and slender, subequal in length in the sexes, outer four joints feebly com- pressed and slightly dilated, third joint subequal in length to the next two combined, fourth very slightly longer than the fifth, the latter to the seventh inclusive subequal, eighth a little shorter and obeonical, ninth and tenth scarcely wider than long, eleventh more or Jess ovate. ~ Pronotum more or less quadrate, widest at the middle, usually about a sixth to a third wider than long, and less than twice as wide as the head; désc smooth, moderately and evenly convex, slightly declivous at the apical angles, very finely, evenly and sparsely, or obsoletely punctulate; apex deeply emarginate, finely or obsoletely margined; s¢des evenly and not strongly arcuate, very finely mar- gined; base feebly arcuate, finely beaded, and about a sixth wider than the apex; apical angles porrect and acute, at times dentiform and more or less everted; basal angles obtuse. Propleure smooth, more or less rugulose and finely punctulate anteriorly. Elytra more or less oval, smooth, usually widest at the middle; base subtruncate to feebly emarginate and a little wider than the contiguous prothoracic base; humeri obtuse, not rounded nor promi- nent; s¢des evenly arcuate, apex more or less caudate; disc evenly convex and punctato-striate, strie shghtly impressed with the punc- tures rather coarse and closely placed, intervals scarcely convex with a series of extremely fine punctules that are more or less irregularly placed, not becoming asperate at the sides and apex. Cauda slightly deflexed at tip, above with the elytral suture impressed, beneath con- cave, the concavity being formed by the general internal surface of the elytra. 59780—Bull. 683—09——19 290 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Epipleure quite wide at the humeri, thence gradually and rather ‘apidly narrowing for a short distance, then less rapidly to apex, where they narrowly margin the sides of the inferior surface of the rauda: surface smooth, generally impunctate. Sterna more or less punctate and rugulose. Parapleure not strongly punctate. Abdomen horizontal, finely, sparsely, quite evenly punctate and more or less rugulose. Legs rather long and quite slender; anterior femora armed with a straight, acute tooth in both sexes; tibial spurs similar in the sexes and rather small, the anterior being equal and acute. Tarsi similar in the sexes. Male—FElongate and dimorphic. Pronotum nearly quadrate, about a sixth to less than a third wider than long. Elytra gradually de- clivous posteriorly; apex briefly produced or strongly and horizon- tally caudate, and accordingly two forms may be recognized: . Forma typica with the cauda equal to about a fourth of the entire length of the elytra. Abdominal segments one to three usually broadly and longitudinally impressed; posterior tibiz quite strongly areuate in basal half, thence to apex straight and distinctly widened. Forma ecaudata with the elytral apex simply obliquely and briefly produced, acute at tip. Abdomen evenly convex from side to side; posterior tibiwe not arcuate and not more suddenly widened toward apex, Female.—Robust. Pronotum slightly transverse and about a fourth to a third wider than long. Elytra more or less broadly oval and slightly ventricose, quite strongly, suddenly and obliquely decliv- ous posteriorly: apex simply acute. Abdomen evenly convex. Pos- terior tibie straight. Genital characters, male. and shghtly arched. Basale evenly convex, oblong, with the sides subparallel, and about twice as long as the apicale. Apicale longer than wide at base, triangular; surface evenly con- vex, with a very fine median groove extending from near the base to the apex, where it becomes wider; sides nearly straight or feebly arcuate; apex acute and not produced; base subacutely lobed at the middle and sinuate laterally. Sternite truncately subtriangular. Each lobe with the external border straight in basal two-thirds converging toward apex, thence evenly arcuate to angle; internal margin rather arcuate, apex nar- rowly rounded; surface rather strongly and densely punctate throughout, setose, sete quite long and dense at tip. Membrane not setose across the bottom of the sinus, the latter rather broad. Edeagophore oblong-ovate, rather small REVISION OF ELEODIINI—BLAISDELL. 291 Female.—Genital segment subquadrate, not strongly developed at base, moderate in size, and setose. Valvula (Plate 1, fig. 21).—Dorsal plate, oblong, with sides more or less reflexed; surface moderately concave, finely and sparsely punc- tate, and with rather short sete; external border moderately arcuate ; internal margin quite straight and parallel; apical margin subtrun- cate and fringed with a few very short hairs; angle subrectangular and without conspicuous sete. Apex short, stout, and triangular, with few very short hairs at tip. Appendage moderately large, as long as the apex, flattened, semi- elliptical, and broadly rounded at tip, with two or three long sete, others short and scattered over the surface. Fossa moderate in size and not conspicuously fringed with sete, situated beneath the ex- ternal half of the apical margin of the dorsal plate. Superior pudendal membrane attaining about the middle of the dorsal plate and longitudinally rugulose. Valvular membranes visi- ble between the apices caudad. Ventrolateral surfaces.—Body shghtly swollen at base, moder- ately and transversely concave before the apex, the latter covered with short sete. Submarginal groove normal and linear, more or less visible from above, especially in dried specimens; lateral plate not visible externally when viewed from above. Internal margins of the valves contiguous for a short distance at base; genital fissure long and fusiform, closed in basal half by the inferior pudendal mem- brane. The three following forms are to be recognized: Forma typica.—Moderate in size. Males more or less caudate. Elytra striato-punctate and not noticeably inflated. Measurements.—Males: Caudate form: Length, 20-28 mm; with cauda, 23-31.5 mm.; width, 8-10.5 mm. Ecaudate form: Length, 24-28 mm.; width, 9-9.5 mm. Females: Length, 22-27 mm.; width 9.2-11.5 mm. Forma inflata—Quite robust. Elytra striato-punctate and more or less distinctly inflated (Plate 1, fig. 22). Measurements.—Males: Not at hand. Females: Length, 26.5-82 mm.; width, 11.5-13.2 mm. Forma grandis.—Very robust and elongate, smooth, body very evenly and gradually narrowed behind. Elytral strive more or less obsolete. Measurements —Male: Length, 37 mm.; width, 13.5 mm. Habitat Lower California. Forma typica.—Cape St. Lucas; Santa Rosa, Gustav Beyer; San Jose del Cabo, collections of Charles Fuchs and California Academy of Sciences 292 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Forma inflata.—San Jose del Cabo, collection Charles Fuchs. Forma grandis.—Calmalli mines, collection California Academy of Sciences. Number of specimens studied, 140. Type in the LeConte collection. Type-locality.—Cape St. Lucas, Lower California. Salient type-characters.—Thorax quadrate, apex deeply emargi- nate, apical angles porrect and acute, but not acuminate; sides mod- erately rounded, basal angles obtuse; dise moderately convex and sparsely punctulate. Elytra oval and convex, acutely attenuate be- hind, strongly striato-punctate, strie shghtly impressed, intervals punctulate (LeConte). Diagnostic characters—The characters that separate this race from eschscholtzii are very feeble. In ducw the apical angles of the pro- thorax are more strongly prominent anteriorly and the apical mar- gin more strongly emarginate. The prothorax is less typically quad- rate and the cauda is less attenuated toward tip. From the differ- ences presented by the series before me, I have no doubt but that a larger series from the different geographical regions would prove these differential characters of no value. In the strongly developed males with long caude the hind tibiz are more or less arcuate in the basal half and rather abruptly dilated in the apical moiety, these characters are not evident in the ecaudate form. From the forms of acuticauda it is separated by the striato-punc- tate elytra, by the sides of the pronotum being quite feebly rounded, and by the elytral apices being more abruptly attenuate, the cauda being horizontal. From dentipes it 1s readily known by the quadrate prothorax with feebly rounded sides. Forma inflata has to be separated from ventricosa by the sides of the pronotum being less convergent anteriorly and by the greater width of the apex as compared with the base; the elytra are not so decidedly inflated and the apices are not produced in the females. This form appears to lead toward ventricosa—in fact may be inter- mediate, but undoubtedly has great affinity with duce. A large series of both species would no doubt demonstrate this relationship. The males are probably ecaudate. Forma grandis is a wonderful specimen of gigantism—such phe- nomenal development makes the limiting of species exceedingly diffi- cult in the absence of an ample series. The epipleure are wide at the humeri and rather rapidly narrowing’ to opposite the meta- sternal epimera, thence narrower, with margins gradually converging to apex. Fourteen series of very fine punctules are clearly indicated upon each elytron. eet tan tt ee i ek ily REVISION OF ELEODITINI—BLAISDELL. 293 General observations —The middle lobe of the mentum is variable, but nearly always triangular, in the large examples quite equilaterally so; the apex is scarcely truncate, generally distinctly rounded; sur- face more or less irregular from the broad, shallow, lateral fovee. The prosterna in the series before me are exceedingly variable. In the more strongly developed examples the prosternum is horizontal and triangularly, strongly mucronate; in the smaller specimens it is rounded antero-posteriorly and without a mucro, at other times it is more or less mucronate behind and more or less convex between the coxe. The prosternum is not strongly prominent ventrally with the Coxe, The prosternum in forma grandis has an extraordinary develop- ment. It is horizontal between the coxe and uncinately produced and slightly but evenly decurved, arcuately sinuate above the mucro. The mesosternum is more or less oblique and concave. The abdominal process is subquadrate, slightly transverse, with its width about equal to the length of the post-coxal part of the first segment, and also to the length of the third; the second is about twice the length of the fourth. The abdominal salient is about a fourth of its width broader than the metasternal process. The metasternum laterally between the coxe is about as long as the width of a mesotibia at middle. The tibial grooves of the femora are well developed, with the margins more or less distinctly cariniform. The profemoral grooves are moderately wide, with the sides feebly arcuate or quite straight. the latter becoming contiguous at the femoral base; the anterior margins are quite acutely dentately laminate at the outer third; the floors are nearly flat, opaque, and smooth. The mesofemoral grooves have the sides evanescent at about basal fifth, while those of the metafemora become evanescent at basal fourth. The floors of the grooves on both are flat and glabrous. The protibiz are feebly compressed and the mesotibie just notice- ably so. The protibiz are distinctly carinate externally; the tarsal grooves are more or less well marked and scabrous; the articular cavities are nearly always closed. The metatibiz are variable in form and carinate to a shght extent the grooves are more or less well marked and at times rather deep, subglabrous, and opaque; the articular cavities are closed. The metatibie are variable in form and carinate to a slight extent externally in basal fourth; the grooves are more or less obsolete or feeble and the articular cavities are closed. The tarsi are more or less variable in length and thickness; usually moderate. The protarsi are about a third of their length shorter than a mesotarsus. Joints two, three, and four are subequal, the 294 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. fourth is at times very shghtly longer than the second, all are wider than long and together a little shorter than the fifth; the first is about as long as wide. The mesotarsi are about an eighth of their length shorter than a metatarsus. Joints two, three, and four are subequal, scarcely longer than wide and together shghtly longer than the fifth; the first is about one and a half times longer than wide. The metatarsi are slightly shorter than half the length of a meta- tibia. Joints two and three are subequal and a little longer than wide, together scarcely as long as the fourth, and noticeably longer than the first. ELEODES TENUIPES Casey. Eleodes tenuipes Casry, Ann. N. Y. Acad. Sci., V. Nov., 1890, p. 399. Elongate-ovate, rather smooth and shining. Caudate. [lead twice as wide as long, finely and sparsely punctate, very shghtly convex. Antenne moderate, scarcely compressed and not dilated in outer four joints, third joint a little longer than the next two taken together, fourth just a trifle longer than the fifth, the lat- ter to the eighth, inclusive, subequal, eighth subtriangular, ninth and tenth suborbicular, eleventh ovate. Pronotum quite quadrate, about a seventh wider than long, widest just at or a little in advance of the middle; disc evenly and moder- ately convex, minutely and very sparsely punctulate; apex quite truncate, more or less finely and obsoletely margined; sides quite evenly and not strongly arcuate from base to apex, finely beaded ; base broadly and very feebly arcuate, about equal to the apex and about a sixth shorter than the length; apical angles very acute, small, denti- form, and everted; basal angles obtuse, feebly or scarcely rounded. Propleure quite smooth externally, more or less obsoletely punctate and rugulose. Elytra elongate oval, widest at the middle; dase feebly emarginate, and equal to the contiguous prothoracic base; humeri obtuse, not in the least prominent; s¢des evenly arcuate, apex more or less pro- duced; dise moderately convex on the dorsum, quite evenly rounded laterally, arcuately and obliquely declivous behind, punctate, punc- tures arranged in distant unimpressed rows of fine, simple, moder- ately approximate punctures, the intervals each with a single series of still finer and widely spaced punctures, which are generally simple, but which laterally toward apex become very coarse sparse asperities. L'pipleurw moderate in width beneath the humeri, thence gradually narrowing to apex, where it narrowly margins externally the infe- rior surface of the elytral prolongation; surface smooth, sparsely and very finely punctate toward apex. REVISION OF ELEODIINI—BLAISDELL. 295 Sterna more or less obsoletely punctate and very feebly rugulose. Parapleure coarsely and somewhat obsoletely punctate. Abdomen horizontal, quite smooth, very finely and sparsely punc- tulate, finely rugulose. Legs rather long and slender. Anterior femora more or less armed ; anterior tibial spurs subequal and rather stout. Tarsi similar in the sexes, with the spinules beneath quite slender. Male—Body moderately slender, antenne attaining the base of the prothorax. Elytra produced at tip forming a cauda, which is nearly horizontal and 4 mm. more or less in length, with its inferior surface formed by the general internal surface of the elytra. Abdomen moderately convex, more or less feebly impressed along the median line on the first three segments. Anterior femora armed with a slender and acute tooth at the outer third. Measurements.—Males: Length, with cauda, 30.5-31 mm.; width, 10-10.8 mm. Females: Unknown to me. Genital characters, male—Edeagophore elongately fusiform, not arched. Basale elongate oblong, very moderately convex, with sides shghtly arcuate. Apicale elongate, triangular, nearly twice as long as wide, mod- erately convex above, groove very fine and linear; sides feebly arcuate, just noticeably sinuate in apical half, tip subacute; base with a broad rounded lobe at central two-fourths, laterally feebly sinuate. Sternite transversely parabolic. Each lobe rather short, with the external border evenly arcuate and continuously so with the apical border, which is rather broadly arcuate, angle not evident; internal border short, feebly arcuate or straight; surface feebly convex, sparsely punctate and setose, setee quite long, longer and denser en the apical margin, impunctate in basal fourth. Membrane not setose across the bottom of the sinus, the latter rather broad. Habitat—Texas (El Paso). Number of specimens studied, 1 male; type not seen. Type a male in the collection of Colonel Casey. Collector, G. W. Dunn. Ty pe-locality.—E]1 Paso, Texas. Salient ty pe-characters—Prothorax with the apex very nearly as wide as the base, broadly, feebly emarginate in circular are, the apical angles very acute and in the form of small everted teeth; base very feebly arcuate; basal angles extremely obtuse; sides almost evenly and distinctly arcuate; disc widest just visibly before the middle, rather strongly and evenly convex, minutely and_ very sparsely punctate. Elytra with humeri not prominent; disc with distant unimpressed rows of fine simple moderately approximate punctures, the intervals each with a single line of still finer and ex- 296 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. tremely widely spaced punctures, which are generally simple, but which laterally toward apex become very coarse sparse asperities (Casey). Diagnostic characters—In tenuipes the apical angles of the pro- thorax are strongly everted, the sides are quite evenly arcuate; in wickhami the apical angles are not everted and the sides are oblique posteriorly; in fenwipes and in a specimen received from Professor Wickham—the latter I refer to wichhami—the elytra are similar and more elongately oval than in /ucw. In the specimen of tenwipes before me the elytra are very shghtly widened behind the middle and this character is less evident in Wickham’s specimen. In both the length of the prothorax is comparatively greater as compared to the width than in duce; the latter when well developed have the apical angles of the prothorax acute and prominent anteriorly, with the apex distinctly and evenly emarginate, and in the ecaudate form these characters are less strongly marked. Professor Wickham’s specimen is of interest, combining the characters of tenuipes and wickhami. Some specimens of Jucw have the apical angles of the prothorax more or less strongly everted. In the specimens of eschscholtzii before me the prothoracic apex is quite truncate and the angles are subacute and not prominent an- teriorly. General observations.—The middle lobe of the mentum is moderate in size, with the apex arcuate, the surface is moderately coarsely punctate and feebly scabrous, not noticeably setose. The prosternum is arcuately convex between the coxe, and with them protuberant ventrally. It is not mucronate nor longitudinally grooved in the specimen before me. The mesosternum is arcuately and vertically declivous, broadly and moderately concave. The abdominal intercoxal salient is quadrate and about a third of its length shorter than the post-coxal portion of the segment, the lat- ter being slightly shorter than the second, which is about a fourth longer than the third, the latter being less than twice as long as the fourth. . The abdominal salient is about a sixth of its width wider than the metasternal process. The metasternum laterally between the cox is about as long as the width of a mesofemur at base. The tibial grooves of the femora are well defined by cariniform margins, the floors are glabrous and shining, and nearly plane. Those of the profemora are moderately wide and nearly attain the femoral base; the margins are scarcely arcuate and gradually converge to the base, where they become contiguous; the anterior margins are acutely and dentately laminate at the apical fourth. On the mesofemora the 2. Ge * yn) wales Abd oe atten mio * REVISION OF ELEODIINI—BLAISDELL. 297 grooves are narrower and gradually converge to become evanescent at basal fourth without becoming contiguous. The metafemora have the grooves moderately narrow and eyanes- cent at basal third, where the margins are distant. The protibiz are slightly compressed and distinctly carinate exter- nally in basal two-thirds; the tarsal grooves are not strongly devel- oped and limited posteriorly by a series of rather coarse and sparse denticulations, and the floors are subscabrous; the articular cavities are more or less closed. The mesotibiz are scarcely compressed and feebly carinate exter- nally in basal fourth; the tarsal grooves are rather well developed, rather broad apically, gradually narrowing toward base, and their ‘floors are subglabrous, being limited by muricate margins; articular cavities closed. The metatibiz are somewhat rounded, obsoletely carinate in basal third externally, thence to apex more or less flattened and concave; surface of the grooves more or less shining and sparsely subasperate ; articular cavities closed. The tarsi in the specimen at hand are moderately long and rather stout. The protarsi are about a third of their length shorter than a meso- tarsus. Joints two, three, and four are subequal, wider than long, and together equal to the fifth; the first is about as long as wide. The mesotarsi are subequal in length to a metatarsus. Joints two and three about as wide as long, the fourth scarcely as wide as long, apparently very slightly decreasing in length from the second to the fourth, together subequal to the fifth; the first is a little longer than wide. The metatarsi are about four-tenths as long as a metatibia. Joints two and three about equal in length and size, a little longer than wide and together scarcely as long as the fourth; the first is about one and a half times longer than wide. ELEODES WICKHAMI Horn. Eleodes wickhami Horn, Trans. Amer. Ent. Soc., XVIII, Feb., 1891, p. 41, pl. Teeatios 2: Elongate, more or less shining and caudate, Head twice as wide as long, rather opaque, feebly convex, finely and very sparsely punctate, frontal suture distinct. Antenne rather long, reaching to or slightly beyond the prothoracice base, not com- pressed nor dilated in outer three joints, third joint a little longer than the next two taken together, fourth very little longer than the fifth, the latter. to the seventh inclusive subequal, eighth slightly shorter and a little more robust, ninth and tenth orbicular, eleventh ovate. 298 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Pronotum subquadrate, one-sixth wider than long, slightly nar- rowed behind, widest at or just in advance of the middle; dise mod- erately convex, sparsely, very finely and indistinctly punctulate; apex slightly emarginate to subtruncate, finely and more or less ob- soletely margined; sides slightly and evenly arcuate anteriorly, and more or less oblique posteriorly, sometimes slightly sinuate before the basal angles, finely margined; base feebly arcuate and finely beaded, about one-seventh wider than the apex; apical angles more or less distinct, acute, not everted although sometimes feebly reflexed; basal angles obtuse, not rounded. Propleurw smooth, obsoletely punctate and more or less rugulose. Elytra elongate oval, widest at the middle; base feebly emarginate, very slightly wider than the contiguous prothoracic base; Aumert obtuse, not prominent nor rounded; sédes evenly arcuate or subpar- allel, attenuate posteriorly and arcuately declivous with the apex more or less caudate; disc shghtly and evenly convex on the dorsum, more strongly so and broadly rounded laterally, punctate, punctures fine and more or less indistinct and arranged in series, sometimes feebly impressed, intervals with a single row of fine, widely spaced punctures, which are generally simple, but may become small asperi- ties on sides about apex. Epipleure moderate in width beneath the humeri and gradually narrowing to the apex; surface smooth and more or less obsoletely punctulate. Sterna rather smooth and more or less obsoletely punctate and rugose. Parapleure more or less coarsely and obsoletely punctate. Abdomen horizontal, somewhat shining, indistinctly, sparsely punctulate, and more or less transversely rugulose. Legs rather long and moderate in thickness; anterior femora armed in one sex at least; anterior tibial spurs subequal and rather stout. ‘Tarsi similar in the sexes with the spinules beneath moder- ately stout. Male.—Body slightly robust. Elytra arcuately and obliquely de- clivous behind, prolonged into a narrow, horizontal cauda, whose under surface is a part of the general internal surface of the elytra, the epipleure passing narrowly along the sides to tip, the latter shghtly decurved. Abdomen feebly and broadly impressed at middle, especially on the second segment. Anterior femora with an acute and rather stout tooth at outer fourth; anterior tibize more or less con- stricted at base. Middle and posterior femora simple; posterior tibie frequently slightly arcuate and narrow in basal half, generally more or less abruptly dilated in apical moiety; both the middle and hind tibiwe generally with the surface quite asperate in apical half. REVISION OF ELEODIINI—BLAISDELL. 299 Measurements —Males: Length, 27-84 mm.; including cauda, 33-39 mm.; width, 12 mm. Females: Unknown to me. Genital characters, male-—Kdeagophore elongate oblong oval, not slender. Basale oblong, about twice as long as wide, moderately convex, sides subparallel. A picale rather broadly triangular, a little longer than wide, feebly convex, without median groove; sides nearly straight to tip, the latter acute; base with a rather narrowly rounded lobe at middle third, laterally broadly and feebly sinuate. Sternite as in tenuipes. Habitat—Arizona (Tucson, H. F. Wickham); New Mexico (Las Cruces). Number of specimens studied, 1 male. Type, a male in the Horn collection. Type-locality—Tueson, Arizona; collector, H. F. Wickham. Salient type-characters.—Thorax subquadrate, a little wider than long, slightly narrowed behind, sides slightly arcuate in front, oblique posteriorly, anterior angles not everted; disc moderately con- vex, sparsely very finely and indistinctly punctate, elytra elongate oval, attenuate posteriorly; disc convex with rows of extremely fine indistinct punctures (Horn). Diagnostic characters.—(See tenuipes.) Without doubt, to my mind the two species just described are extreme forms (heterotypes) of the same species, but on account of the paucity of material known to me, I have deemed it best to retain them separate until a larger series shall have been collected. At most wickhami can only be a race of tenuipes, Casey’s name having priority of three months (tenuipes, November, 1890; wich- hami, February, 1891). Unfortunately only males are known to me. Both forms have been compared with duce by their authors. They do show greater affinity to that species than to any other, except gracilis. From luce they can be recognized by the elytra being sculp- tured with rows of fine punctures, and not strongly striate nor with the intervals at all convex; the elytra are slightly more inflated and more suddenly declivous behind. The prothoracic characters are too variable to be relied upon. By direct comparison I can not say that the legs are always longer than in luce. The form of the hind tibize in wickhami is not peculiar to that species, as the same character is seen in the males of a number of other species, notably in the caudate form of /ucw. It is strange that such characters should have been overlooked or ignored by pre- vious writers. 300 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. From gracilis the two species under consideration differ in the caudate elytra of the males and in having the prothorax slightly longer. The prothorax in tenuipes is indeed almost as in the typical gra- cilis. The type of tenuipes and wickhami were practically collected in the same geographical region, the latter extending into New Mex- ico (Wickham’s specimen) ; gracilis occurs in the same area. My specimen of tenuipes was obtained from Mr. G. W. Dunn, who collected it at El Paso, Texas; it was probably one of the same lot from which Colonel Casey obtained his type. General observations—The following analytical remarks are founded upon Professor Wickham’s New Mexico specimen : Mentum moderate in size and triangular, with apex rounded; sur- face very feebly convex and quite coarsely punctate. The pro- and mesoternal characters are identical, as are also the tarsal characters, with those of tenuipes. The abdominal and metasternal salients and abdominal segments do not differ relatively from those of tenuipes. The metasternum laterally between the coxe is about as long as the width of a mesofemur at base. The femoral and tibial characters are the same as in fenuipes, ex- cept that the protibiz have the articular cavities widely open. The metatibize are just noticeably arcuate in basal half and gradually but distinctly dilated toward apex. ELEODES VENTRICOSA LeConte. Eleodes ventricosa LECoNTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 186.— Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 311.—CHAMPION, Biol. Centr. Amer., IV, Pt. 1, 1884, p. 78. Fusiform ovate, robust, elytra usually strongly inflated, smooth, and more or less shining. Head twice as wide as long, more or less moderately convex, fre- quently slightly and transversely impressed between the eyes, with the vertex more strongly convex, frontal suture distinct, finely, sparsely, and irregularly punctate. Antenne moderately long, at- taining a slight distance beyond the prothoracic base, comparatively slender, very feebly compressed, and slightly clavate; third joint about equal to the next two combined, fourth a little longer than the fifth, the latter to the seventh, inclusive, subequal in length, eighth shorter and slightly subtriangular, ninth and tenth suborbicular, eleventh ovate. Pronotum slightly transverse, widest at the middle, one-fourth to one-third wider than long, narrowing moderately anteriorly, slightly posteriorly; disc quite evenly and moderately convex, finely and sparsely punctate; apex slightly emarginate, finely and more or less Li ncaa Mek nine nelle Ste ee i a Se ae ee et REVISION OF ELEODIINI—BLAISDELL. 301 obsoletely margined; s¢des moderately arcuate from base to apex, finely margined, the margin frequently more or less narrowly re- flexed ; base moderately arcuate and rather coarsely margined, about one-third wider than the apex and distinctly greater than the length; apical angles distinct and more or less prominent anteriorly, gener- ally acute and not dentiform; basal angles distinctly rounded, fre- quently subobtuse. Propleure sparsely punctate and more or less rugulose, frequently smooth externally, Elytra broadly oval and usually strongly inflated, always less than twice as long as wide, widest at the middle; base moderately emarginate, usually as wide as the contiguous prothoracic base; humert obtuse, not prominent nor rounded; sides evenly and quite strongly arcuate, apex more or less produced; dise quite evenly and strongly convex from side to side, at times more or less feebly de- pressed on the dorsum, and more strongly rounded laterally, evenly and arcuately declivous posteriorly ; sw7face punctate and feebly sul- cate, punctures more or less coarse and arranged in moderately dis- tant series, intervals at times feebly convex, at others flat, with a single series of fine punctures, which are more or less regularly or irregularly arranged; laterally and on the apex the punctures be- come much coarser and the intervals at times strongly convex. | Epipleure moderately narrow at base and very gradually narrow- ing to apex, superior margin near base broadly and feebly sinuate; surface smooth, very finely and distinctly or obsoletely, sparsely punctate. ; Sterna more or less shining, moderately densely punctured and rugose. Parapleure more or less coarsely punctate. Abdomen horizontal, moderately convex, shining, and glabrous, very sparsely punctulate and more or less rugulose. Legs rather long and moderate in thickness. Anterior femora armed in one sex. Anterior tibial spurs similar in the sexes, the anterior slightly longer than the posterior, both acute. Tarsi nearly similar in the sexes; each joint, except the last, of all the tarsi with a slender and acute tuft of golden-yellow and somewhat modified spines on apical angles beneath. (See male.) Male—Elytra rather gradually and arcuately declivous behind and produced at tip into a moderately long cauda, which is not hori- zontal; elytral sides and dorsum feebly and broadly sinuate before the produced apex, the latter with its inferior surface a part of the general internal elytral surface and narrowly margined at the sides by the epipleure. Abdomen more or less broadly impressed in the median line on first three segments. Anterior femora armed with a rather long, acute, and rather strongly curved tooth at outer fourth. 302 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. First joint of the anterior tarsi with the tip slightly and transversely produced beneath, the same bearing a tuft of modified spines which obstructs the groove. Female.—Elytra quite evenly arcuately declivous behind, with the apex feebly produced or simply subacute. Abdomen not impressed. Anterior femora more or less sinuate in outer fourth. Measurements.—Males: Length, 30-83 mm. (with eauda) ; width, 13-14 mm. Females: Length, 26-33.2 mm; width, 13.5-15 mm. Genital characters, male.—Kdeagophore elongately oval, depressed and not arched. Basale oblong, very moderately and evenly convex, sides feebly arcuate. Apicale triangular, very slightly longer than wide and rather broad; surface rather feebly convex and not grooved; sides evenly and not strongly arcuate to tip, the latter subacute; base with a sub- triangular lobe at middle, which is narrowly rounded at tip, feebly sinuate laterally. Sternite transversely parabolic. Each lobe with the external mar- gin evenly arcuate to apex, the latter somewhat broadly rounded ; in- ternal border straight to feebly arcuate; surface very feebly convex, shining, not very strongly chitinized at inner and basal third, sparsely punctate, sete not long, those on the apical margin longer and not very dense. Membrane not setose across the bottom of the sinus, the latter broad. Female.—Genital segment robust, trapezoidal, valves reflexed ex- ternally, strongly chitinized and setose, Valvula (Plate 5, fig. 1).—Dorsal plate oblong, strongly concave, margins more or less reflexed ; surface glabrous, sparsely and strongly punctate, setose, sete rather short and inconspicuous; external mar- gin straight or feebly sinuate; apical margin inwardly obliquely truncate, angle acute and as prominent apically as the apex, the lat- ter quite short, robust and triangular, both sparsely set with short sete; internal margin arcuate or straight. Submarginal groove more or less visible from above and quite moderate. Appendage short and flattened, semielliptical, projecting but a short distance beyond the margin of the fossa and eccentrically placed. Fossa terminal, rather large and transverse. Superior pudendal membrane attaining the middle of the dorsal plate and feebly longitudinally rugulose. Basal prominences quite well developed. Ventrolateral surfaces strongly convex, glabrous basally and more or less concave before the apex laterally, very sparsely and coarsely punctate. Internal margins of the valves contiguous in basal and apical fourths. Genital fissure rather broad and fusiform. Inferior pudendal membrane closing the fissure in basal half. AE ll Fs = \. SS {re ere oe REVISION OF ELEODIINI—BLAISDELL. 303 Habitat—Texas (Brownsville, Charles Drury, H. F. Wickham; Uvalde, Wickham; Wades, Alice, and San Diego, Hubbard and Schwarz, collection U. S. National Museum). Mexico (coll, F. Bates). Number of specimens studied, 15. Type in the LeConte collection. Type-locality.—Texas, near the Rio Grande River. Salient type-characters.—Thorax finely punctate, anteriorly mod- erately and posteriorly slightly narrowed; sides rounded; anterior angles acute; base broadly rounded, humeri scarcely acute. Elytra striato-punctate, intervals uniseriately punctulate (eConte). Diagnostic characters.—Quickly recognized by its robust form, the elytra being strongly ventricose in both sexes, moderately caudate in the males, the cauda being oblique and not horizontal; in the females the elytral apex is shghtly produced. The apical angles of the prothorax are distinct and not dentiform ; the basal are rather broadly rounded, the angles may be sometimes feebly evident; the sides narrowing moderately anteriorly. As LeConte writes the punctures composing the striz of the elytra vary very much in size; sometimes they are very large, while at others they are hardly different from the more distant interstitial punctures. Two examples in the series before me have the elytral intervals per- fectly flat and the strize not in the least impressed. The antenne are noticeably feebly clavate on account of the joints one to eight, inclusive, being elongate and retaining the same width throughout, while joints nine to eleven, inclusive, are slightly widened at the middle and more or less transverse. The same structure is observed in eschscholtzii and luce, only that the eighth joint is also feebly widened. This may also be the case in ventricosa. This species is separated from luca forma inflata by the basal angles of the prothorax in the latter being obtuse, distinct, and not at all rounded. See var. falli, p. 306. General characters.—The mentum is large, more or less trapezoidal, rarely subquadrate; apex truncate or feebly arcuate, slightly reflexed, and frequently impressed at middle so as to appear more or less emar- ginate; surface coarsely punctate, sete very short, subfoveate at the basal angles; inflexed lobes rather stout. The prosternum is quite variable. Convex between the cox, with which it is quite strongly protuberant ventrally and also moderately short before the acetabula; compressed and perpendicularly truncate behind, rising into a small mucro at the ventral angle; at times sub- truncate behind, with a moderate mucro, at others rounded and not at all mucronate. 3804 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The mesosternum is quite short, obliquely declivous, and rather strongly concave. The abdominal process is about one-fourth of its width broader than the metasternal salient, nearly quadrate, slightly transverse, and equal in length to the post-coxal portion of the same segment, the latter being equal to the third in length; the second is twice as long as the fourth. - The metasternum laterally between the coxe is about as long as the width of a mesotibia at apex. The tibial grooves of the profemora are well developed and defined by smooth cariniform margins, that are scarcely at all arcuate, quite evenly and gradually convergent, to become contiguous a short distance from the base; floors glabrous and shining, feebly concave; the anterior margins at the outer fourth are dentately laminate in the males; in the female more or less sinuately atrophic in the outer fourth. On the mesofemora the sulci are less strongly defined by cariniform margins, that become quite evanescent at the inner third; the floors are glabrous, shining, and very feebly concave. The metafemoral tibial grooves are rather narrow and with rather feeble margins, that become evanescent without becoming contigu- ous at about the middle. The protibiz are scarcely compressed, not carinate, but smooth ex- ternally ; tarsal sulci more or less feebly developed and subasperate ; posterior surface muricately sculptured, but not coarsely so; articular cavities usually closed. The mesotibiz are more or less rounded and not coarsely muri- cately sculptured, smooth, and not carinate in basal third externally; tarsal grooves more or less obsolete and subasperate ; articular cavities closed. Metatibiz quite cylindrical, not coarsely muricately sculptured, more or less feebly flattened and rarely grooved externally; articular cavities distinctly closed. The tarsi are moderately long and stout. The protarsi are about one-fourth of their length shorter than a mesotarsus. Joints two, three, and four are subequal in length and slightly wider than long, together about equal to the fifth; the first is slightly longer than wide. The mesotarsi are just slightly shorter than a metatarsus. Joints two, three, and four subequal in length and very shghtly longer than wide, together equal to the length of the fifth; the first is about a half longer than wide. The metatarsi are equal to half the length of a metatibia. Joints two and three subequal in length, distinctly longer than wide and REVISION OF ELEODIINI—BLAISDELL. 305 together scarcely as long as the fourth; first joint with a total length of about twice as long as wide. The ungues are comparatively long. ELEODES VENTRICOSA var. FALLI, new. Elongate-ovate, smooth and shining, elytra scarcely inflated, simply punctate and more or less caudate in the male. Antenne as in ventricosa. Prothorax quadrate, about one-eighth wider than long, not notice- ably more narrowed anteriorly than posteriorly; a@pea broadly and quite strongly emarginate; s¢des evenly and quite moderately arcuate from base to apex; base one-fifth to one-fourth wider than the apex and more or less greater than the length; apical angles strongly prominent anteriorly, acute, rather large, slightly reflexed and not everted; basal angles obtuse and not rounded. Propleure quite smooth and quite obsoletely sculptured. Elytra rather broadly oval, scarcely inflated, sides eveniy and moderately arcuate, with the apex more or less produced; disc quite evenly arcuate from side to side, sometimes slightly depressed on the dorsum, not suleate, punctate, punctures fine and usually of the same size; the serial punctures may be coarser, scarcely more strongly sculptured laterally. Cauda feebly impressed along the suture and a little longer than in ventricosa. Sterna and parapleure more or less obsoletely sculptured and glabrous. Legs usually rather slender; anterior tibiae slightly constricted at base, otherwise as in ventricosa. Male-—Body somewhat robust. Elytral cauda about 4 mm. long and less oblique than in ventricosa. The tuft of spinules on the first joint of the anterior tarsi is interrupted at middle and does not close the groove. Otherwise as in ventricosa. Female——Body moderately robust. Elytral apex more or less shghtly produced and subacute. Otherwise as in ventricosa. Measurements—M ale: Length, 32.2 mm. with cauda; width, 12.5 mm. females: Length, 30-83 mm.; width, 13.2-14 mm. Genital characters, male.—As in ventricosa. Female-—Compares with ventricosa as follows: The genital seg- ment has the apical margin of the dorsal plate of a valve less oblique and sometimes feebly arcuate; angle about as or less prominent. The ventrolateral surfaces are less strongly but more evenly convex, and the surface lines straight when viewed longitudinally. The appendage is larger, subconico-semiellipsoidal, completely fill- ing the fossa and projecting beyond the margins of the dorsal plate, not as long as wide. 59780—Bull. 683—09 20 306 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Fissure narrower, otherwise as in ventricosa. One of the speci- mens from E] Paso, Texas, has the apical margin of the dorsal plate as in ventricosa. Habitat—Texas (El Paso, March, H. Soltau; Fort Bliss, collection OFLC. Ball). Number of specimens studied, 3 (1 male and 2 females). Co-types distributed as follows: Male in the collection of H. C. Fall; female types in the collection of the U. S. National Museum. Type-locality.—Texas (El Paso and Fort Bliss). Diagnostic characters —Falli is separated from ventricosa by the less transverse prothorax, the very acute, anteriorly prominent, and more or less reflexed apical angles of the same; the basal angles are distinctly obtuse and not rounded; the elytra are much less inflated, and consequently appear more elongate, and the cauda in the male is longer. From luce forma inflata it is recognized by the more strongly de- veloped apical angles of the prothorax, the oblique cauda of the male, and the mutic anterior femora of the female. From tenuipes, which it approaches, it is quickly recognized by the well-developed anterior prothoracic angles; the cauda in tenuipes is shghtly oblique at times, but more horizontal in wickhami. Tenuipes is less inflated and therefore narrower and more elongate. Falli is undoubtedly intermediate between tenuipes and ventricosa, and differs with the latter from all other members of the subgenus by the character of the vestiture of the anterior tarsi and the mutic anterior femora in the female. General characters—The mentum is exactly as in ventricosa, and the prosternum is more or less rounded behind and feebly mucronate. The mesostermum is as In ventricosa. The relative size of the metasternal and abdominal salients are the same as in ventricosa. The post-coxal portion of the first segment is a little longer than the process and quite equal to the third in length; the second is about a fourth of its length longer than the third, the latter being about twice as long as the fourth. The metasternum laterally and the femora are as in ventricosa. The protibiv are more or less distinctly carinate externally, and the articular cavities are more or less open; the tarsal sulci are quite well developed, as they also are on the mesotibie. Otherwise the tibie and the tarsi are relatively as in ventricosa. Subgenus BLAPYLIS Horn. The present subgenus is based chiefly upon the character of tarsal pubescence. The males have two or three basal joints of the anterior REVISION OF ELEODIINI—BLAISDELL. 307 tarsi feebly thickened at tips beneath, and there clothed with quite dense, silken, yellowish or brownish pubescence. The first joint of the middle tarsi also bears a small tuft of similar pubescence at tip beneath, and at times the second exhibits a very small one. The species naturally associate themselves into groups, the relation- ships by which this is accomplished result from their genealogical development; and, accordingly, the pubescent tufts may be flat, shghtly longer and truncate, or still longer, usually smaller and subacute. In most females the first joint of the anterior tarsi is more or less transversely thickened at tip beneath, and simply clothed with ordi- nary spinules across that point where the groove is more or less interrupted. In the males the pubescent tufts always interrupt the grooves which are evident on the remaining joints, and in all except the first in the female. At times there is a slight tendency to a lateral compression of the tarsi, but this is not always evident. The femora are always mutic; at times the anterior femora are slightly sinuate at apex beneath. The pronotum is never declivous at the sides; the disc is always evenly convex from side to side, but in the parvicollis section there is some flattening along the lateral margins corresponding to the granu- late area, the sides appearing somewhat dilated as a result. The marginal head is always distinctly visible from above, entire in those species which are very slightly sinuate or constricted at the base, and obsolete or simply continuing as a mere line to the basal angles in those species which are more strongly constricted. The base is at times so strongly constricted as to appear almost pedun- culated. The general form is oval to ovate, some species are slightly more elongate and oblong. The elytra are oval, with sides more or less strongly rounded, apex never produced; the humeral angles are always more or less rounded, except in ¢¢bzalis, where they are dis- tinctly angulate. There is in this subgenus a tendency for the anterior tibie to be- come constricted at base, most evident in ¢ébialis. The tibial spurs are quite similar in the sexes. Among the species of Blapylis are to be found our smallest Eleodes. With the subgeneric tarsal characters are to be correlated the fol- lowing: Genital characters, male—Apicale of the edeagophore triangular and without a median groove, except in tibialis (see p. 316) for re- marks upon this species). 308 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Female.—Geuital segment triangular or triangulo-paraboliec in out- line. Valvule.—Dorsal plates oblong, with a few scattered punctures and’ sete, apical margins not always well defined (see Synopsis) ; apices membranous or chitinous, always finely setose, generally with a pencil of longer hairs at tip (penicillate). Appendage small, mammilliform and penicillate. Superior pu- dendal membrane nearly attaining the apical fourth of the dorsal plate, or only the middle (cordata) ; genital fissure in the apical third of the segment. General characters —The crural characters are quite variable with- in certain limits in each species. The profemora are more or less clavate, the mesofemora at times feebly so in external two-thirds; the metafemora have their opposed surfaces quite parallel. The protibize are at times subcarinate in basal half externally and the tarsal grooves are not developed, although sometimes indicated ; the mesotibiz are frequently obsoletely grooved on their external sur- faces, and the metatibiz more or less flattened on those surfaces. The articular cavities are closed on all the tibiee. The relative lengths of the tarsi and of the joints in each are prac- tically the same as in the other subgenera. The protarsi of the male are a little longer than in the female. The stoutness and length of the legs differ very much in the same species, but for purposes of comparative measurements, normal ex- amples have been selected. Distribution.—Of the seventeen species given in this paper all but three occur in California; out of six races only two are not found in that State. Tibialis 1s apparently peculiar to Lower California; /econte? to Colorado and New Mexico; snow?7i to Colorado, New Mexico west- ward to the Colorado River in Arizona; tenebrosa and its var. nana to Nevada and eastern central and northern California. Inculta may possibly occur on the mainland in California; it is authentically abundant on Santa Rosa Island off the Santa Barbara coast; consobrina occurs from Los Angeles County to Siskiyou County; scabripennis is from the Fort Tejon region; blanchardii at Indio and the mountains in eastern San Diego County; neotome in southern California, and I have taken it in numbers from the wood- ‘ats nests on the hillsides of Mission Valley, San Diego County. Fuchsii oceurs in the high mountains and forests of eastern Tulare County; hornii from the eastern central and northern parts of the State. Parvicollis from ledges on the hills near the coast in central parts of the State; var. planata in the western central, and producta in the v REVISION OF ELEODIINI—BLAISDELL. 309 ‘mountains and foothills from the southern to the northern parts of California. Hoppingt is at present only known from Eldorado County ; clavi- cornis and scabrosa maritime and lovers of sand-dunes. Cordata occurs from Los Angeles to northern parts of the State and at all extreme points of its northern distribution passes into pimelioides, which has the largest distribution of any species of Bla- pylis, occurring in Colorado, Montana, Wyoming, Idaho, Utah, Ne- vada, Oregon, Washington, Vancouver Island, British Columbia, its var. brunnipes in Idaho, Wyoming, Colorado, and Nevada. Most of the species are developing into a considerable number of incipient races. Consobrina Blanchardii ¥Fuchsii SX bape ee \ Oe Lore! re oe Nenioans a Scabrosa lanat % ee ee Hornii a oN ° om a ee ee ‘Prodncta | ae oe: Rotundicoltis Hoppingii Lecontei ee os Os nipes Subgeneric Trunk I'1G. 6.—GENHALOGICAL DIAGRAM OF THE SUBGENUS BLAPYLIS. Genealogy.—lt is doubtful if by any herculean effort a biologist could in the period of a few years, with the comparatively small series of specimens at hand from the different regions which they inhabit, work out a correct history of descent. The individuals of the species making up the present subgenus are very protean in their divergences. They have evidently diverged along three main lines from the an- cestral subgeneric trunk (see diagram). It is quite hypothetical whether ¢bialis belongs to Blapylis at all; if not, it must have con- geners in Mexico. If it does belong here it diverged early. Taking the degree of constriction of the prothoracic base as the criterion of divergence, two lines of descent are evident—cordata and pimelioides with a cordate type of prothorax and the lateral margin 310 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. obsolete upon the strong basal constriction, and the other line con- tinuing reactive to a greater degree, again may have diverged dichot- omously, evolving the fenebrosa strain, with the sides of the pro- thorax evenly or nearly arcuate from apex to base; the other being the consobrina-parvicollis ramus. Parvicollis 1s not only plastic, but prolific, populating the regions it inhabits with puzzling varia- tions, many of them at the beginning of their incipient racial devel- opment. In these forms the sculpturing is rather fine, abdomen not coarsely sculptured, pronotal base quite strongly constricted, the pro- thorax is broader and less strongly convex and more or less impressed along the sides. The consobrina ramus apparently departed from an interesting strain; that is, ¢wculta, an insular species apparently, for if it be found on the mainland it is rare; in énculta the thorax is broad, very feebly rounded at the sides, and scarcely at all sinuate near the base, with the marginal bead strong and reaching the basal angles—these characters in contrast to the more elongate consobrina, with narrower prothorax which is distinctly constricted at base, the marginal bead becoming a fine line upon the constriction, and the elytra more tuber- culately sculptured. Here, again, are evidences of plasticity to en- vironment. Let the student correlate the above characteristics with the genital characters as presented below in the synopsis, and I believe the above relationships will become more evident. SYNOPSIS OF GENITAL CHARACTERS. Males. Apicale of the edeagophore with a median groove; sides scarcely arcuate. tibialis, Apicale without a median groove. lecontei. snow ii. tenebrosa, inculta, consobrina. blanchardii. neotome. fuchsii. hornii. parvicollis, hoppingii. clavicornis, scabrosda, cordata, pimelioides. Apicale larger, sides more or less broadly sinuate, apex appear- ing attenuate 2-2 ee eee eee Apicale rather narrow; sides feebly arcuate_______--------_-. REVISION OF ELEODIINI—BLAISDELL. sll Females. Superior pudendal membrane long, attaining the apical fourth of the dorsal plate. Apical margin of the dorsal plate not defined from the apex; appendage fackenedasmammiliitorm ands subapical] 2-22 ee tibialis. Apical margin usually well defined from the apex and more or less arcuate; appendage true mammilliform and more at base of the apex. lecontei. snowiti. tenebrosa. inculta. consobrina. blanchardit. neotome, fuchsii. hornii. Dorsal plate rather broad, not more than three times as TONS RAST Wil CC eee ae ee ee ee Se ee ee clavicornis. scabrosa. Apical margin of dorsal plate not well defined from the apex, {parvicollis. the latter more frequently Semichitinous_________________ ieee Superior pudendal membrane short, attaining the middle of the dorsal plate. Apical margin of the dorsal plate more or tess truncate; anglef{cordata. evident and more or less rounded ; appendage at base of rea melioides. Dorsal plate elongate, four times as long as wide______ Caseyi is aberrant. In the male the apicale is grooved and in the female the superior pudendal membrane is long, attaining the base of the apex. The material is not sufficient to indicate the relation- ships. ANALYTICAL KEY TO SPECIES OF SUBGENUS BLAPYLIS, Lateral marginal bead of prothorax always distinct and entire, attain- ATS pe RD Seale ST CS ee ace ee PS te a as al Lateral marginal bead more or less obsolete on basal constriction _______ 5 1. Thorax transversely suboblong-oval, lateral margins feebly or not at all SSL TNT EAC cls teas 1 AS Cee ee Nee ee ee eee as 2 Thorax quadrate, not at all sinuate at base; form very elongate and de- pressed: surrace smoovhvandeshimin cee ae eee eS eee tibialis. PEE SUETACEHINOLEe On. LESSi'Shimin ees ers See Se RS st Ss ee ee 3 Surface opaque; elytra sculptured with small granules, densely and irregu- larly placed. Size larger, length 14-18.5 mm.; granules shining and rounded_tenebrosa. Size smaller, length 10-12 mm.; granules coarser and submuricate laterally, more evidently setigerous___. = ~_______________ var. nana. 3. Thorax moderately transverse, sides rather strongly rounded_____________ 4 Thorax distinctly transverse, sides very feebly arcuate, scarcely sinuate at ASCE OTIV ATO DUES tate = eed ek a eee ek a inculta. bs Sculpturing rather finely muricato-granulate laterally; sides of the pro- notum evenly rounded from apex to base, sometimes briefly sinuate at base-oronrmnelongatemee meas sews See Se een EP ete aa al) ee hea snowti. Sculpturing more coarsely muricato-granulate ; pronotum briefly, feebly, and broadly sinuate at base; elytra flattened (or convex) on the dorsum; SSUES CC es CHU] a ae ee a Ey ed ee lecontei. 312 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. 10, aah 13. 14. Thorax transversely suboblong-oval; somewhat coarsely and discretely punctate ..4. 23.2 Bae ee ee ee eee 6 Thorax subcordate and coarsely, rugoso-punctate___—____-__--=--___=__ = Ae Propleure: not? hairy 22255520 ea ee ee ee iG Propleurs hairy: 2 soe ee Be ee ee eee 14 Genz: not, Produced 22. 2 ee ha ee ee eee 8 Genz produced ; sculpturing more or less coarse First three abdominal segments densely, coarsely punctate and more or less PUGOSE 252 ee ee 9 Virst three abdominal segments more finely and sparsely punctate, the first somewhat .Buculosecc 23 es oe we ae a ee eee 10 Pronotum and elytra moderately convex, surface dull; elytra [consobrina. more or less tuberculate laterally, asperate eee Pronotum and elytra very strongly convex; surface shining; elytra subas- perately sculptured =. Se ee ee ee ee ee neotome. Humeri more or less obtusely rounded, frequently prominent anteriorly___11 Humeri obsolete; elytra equal to the prothoracic base, very gradually widen- ing to middle, causing the thorax to appear somewhat remote; surface smooth, shining Jor dull’: antennze long] S22 2 ee eee hornii. Humeri obtusely rounded and not prominent to prominent anteriorly ; thorax noticeably, transverse. 2s 2c ES ee ae ee) ee 12 Humeri obtusely to broadly and evenly rounded, not at all prominent an- teriorly; thorax not noticeably transverse, more as in consobrina, fre- quently: subeordate (female) surface dulle—2s2*2- sees ee fuchsii. Surface muricato-granulate laterally and on apex; epipleural margin at humeri more or less) visiblestrom above: 2 = See ee eee 13 Surface finely and densely muricato-granulate; thorax somewhat finely punctate; form oblong-oval and depressed; antennz short; humeri TOUT Cc 8 ee See A ee PES ee ee ee eee hoppingii. Surface shining; elytra short oval, convex, rather rapidly declivous behind ; humeri obtusely rounded, not at all prominent; pronotum rather coarsely PUPA CG: ee as ee Sh a Se eee parvicollis, Surface more or less dull. = Elytral base not broad; humeri rounded, feebly obtuse and not conspic- uous; elytra somewhat attenuate at apex, sides evenly rounded, dise gradually declivous behind (male). *Elytral base broad, humeri rather broadly rounded and somewhat obtuse, Conspicuous and not prominent anteriorly; apex less attenuate; disc more rapidly decli- yous: behind.(temale) {222 32" = eee eee var. planata. Elytral base broad; humeri conspicuous and more or less prominent anteriorly and margined by the epipleurie; elytra oblong, disc flat- tened, narrowing behind the middle, sides parallel; prothorax strongly tranS Verses is Sea ee ee ee ee eee ieee var. producta. Elytral base broad; humeri as in producta; elytra oblong, disc flattened usually, coarsely and densely rugoso-punctate centrally, muricato- tuberculate laterally and on apex; thorax more sparsely and coarsely punctate; female often with the form of cordata_____- var. constricta. Pronotum not noticeably transverse; disc with rather long erect hairs lat- erally in the granulate area; propleural hairs rather long__-_clavicornis. Pronotum distinctly transverse; disc with very short, erect setz laterally ; propleural hairs shorter than in clavicornis == scabrosa. 5s horax strongly constricted atibase=—-=-—= === = ee 16 Thorax less strongly and more briefly constricted; sides broadly, strongly, and evenly rounded as in scabrosa_------------------ var. rotundipennis, uP es Ps o < REVISION OF ELEODIINI—BLAISDELL. yO) 16. Sides of pronotum more strongly arcuate in front of middle, behind narrow- ing rather suddenly and more or less subangulate, or more evenly rounded and less gradually narrowing behind. Mivitran coarsely LUZOsO-pUNChlCa= = nae te ee cordata, Elytra sculptured with distinct tubercles, which vary in size, rounded or submuricate. Tubercles distinct, rounded, not noticeably setigerous; legs black aie Sh pep SOREN. Rie OR SSE Beh Pw Dele Corn Bet) carte ea ee dS pimelioides. Tubercles more evidently setigerous; legs brown____var. brunnipes. Sculpturing finer, distinctly pubescent throughout, somewhat like hirsuta, Nairs shorter than in that species; legs reddish or IO TO yt Ss Hr ee eR ee ee ls Dee es caseyi. The reader will no doubt appreciate the difficulty of formulating a table that will make it possible to place all specimens, especially in a genus so protean as leodes. I desire to impress on the student the importance of first determining the subgenus to which the specimen under consideration actually belongs, then try to determine whether it is typical specifically or exhibiting considerable variation. The most important diagnostic tests are those considering the general form, form of thorax with its sculpturing, form and sculpturing of the elytra. Color plays very little part in /leodes, although surface luster is often helpful. ; ELEODES TIBIALIS, new species. Klongate, oblong-ovate, nearly three times longer than wide, very smooth and shining, strongly depressed. Head about twice as wide as long, feebly convex, more or less obso- letely impressed laterally, frontal suture obsoletely indicated, very finely, sparsely, and irregularly punctate, punctures just noticeably denser laterally and on the epistoma. Antennw moderate in length and stoutness, outer three joints very feebly compressed, feebly and very gradually dilated, third joint about equal to the next two taken together, fourth slightly longer than the fifth, the latter, sixth, and seventh subequal in length, eighth more or less triangular, ninth and tenth orbicular, eleventh short ovate. Pronotum quadrate, widest at or just a little in front of the middle, about a third wider than long; disc evenly, very moderately convex. frequently somewhat depressed, more or less declivous at the apical angles, very minutely punctulate, obsoletely so at times, punctules sparse ; apex truncate to moderately emarginate, very finely and more or less obsoletely margined; sides quite evenly and not strongly arcuate, slightly less so and at times somewhat straight in the pos- terior third, marginal bead entire and fine; base truncate, sometimes feebly sinuate at middle, margin fine, a little wider than the apex; apical angles more or less prominent anteriorly, obtusely and nar- rowly rounded; basal angles obtuse and not in the least rounded. 314 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Propleure smooth, obsoletely punctulate, and more or less feebly rugulose. Elytra elongately oval, smooth, widest at the middle, sometimes more posteriorly; dase truncate and at times just noticeably wider than the contiguous prothoracic base; humeri obtusely angled, not rounded nor prominent; s¢des evenly arcuate, obliquely narrowing in apical third, apex not very narrowly rounded; disc depressed, feebly convex, sides more or less suddenly inflexed and almost evenly rounded, quite evenly and arcuately declivous posteriorly; surface usually obsoletely punctulate or finely and irregularly so, a serial arrangement often evident, the interstitial punctures of the same size as those of the series, often very feeble and eroded striz are evident ; punctules usually denser on the sides and apex, those about the latter with small brownish hairs. Epipleure moderately wide, gradually narrowing from base to apex, superior margin rather strongly defined, especially toward apex, beneath the humeri feebly and broadly sinuate; surface at times feebly concave, smooth, obsoletely punctulate, and more or less feebly rugulose. Sterna rather smooth, more or less obsoletely and sparsely punctate, rugose, Parapleure rather evenly and not densely punctate. Abdomen finely and sparsely punctulate, more densely so on the fifth segment, where it is also more or less finely setose along the margin, Legs moderate in stoutness and somewhat long; femora mutic, finely and sparsely punctate, the profemora very feebly sinuate in apical fifth; anterior and middle tibiz more or less strongly con- stricted at base; anterior tibial spurs similar in the sexes, the ante- rior a little stouter and just a little longer than the posterior, both acute. Anterior tarsi dissimilar in the sexes. Male.—Rather narrow, elongately oblong-ovate. Antenne attaining the base of the prothorax, the latter with base equal to the length. Elytra with the sides not strongly arcuate. Abdomen somewhat oblique apically, rather feebly convex, broadly, more or less strongly impressed on the first two segments, and the metasternum feebly con- ‘ave, the mesosternum rather prominent. Anterior tibizxe suddenly constricted at base; first joint of the anterior tarsi feebly thickened at tip beneath and clothed with golden hairs. Female.—Rather more robust. Antenne not attaining the base of the prothorax, the latter with the base less than the length. Elytri more broadly oval, sides more strongly arcuate. Abdomen horizontal and moderately convex. Anterior tibiz moderately and rather grad- ually constricted at base; first joint of the protarsi scarcely thickened at tip beneath, groove entire. a tt ores re ete wy SY pRB PES a: REVISION OF ELEODIINI—BLAISDELL. 315 Two forms are recognized— Forma typica.—Elongate-ovate, strongly depressed and very finely punctulate; anterior tibiz quite strongly constricted at the base. Forma oblonga.—Elongate oblong-ovate, not strongly depressed, vather strongly punctate; anterior tibie not constricted at base. Measurements.—Males: Length, 20-22 mm.; width, 6.5-8 mm. Females: Length, 20-23 mm.; width, 9—-9.2 mm. Genital characters, male-—Edeagophore of the usual oblong-ovate form and depressed. Basale oblong; sides broadly and evenly arcuate; surface rather evenly convex, not gibbous nor noticeably arched. Apicale triangular; surface rather evenly convex, with a median shallow and rather broad groove extending to the apical fourth; sides scarcely arcuate; apex acute and slightly produced; base with a rounded lobe at middle and feebly sinuate laterally. Sternite feebly transverse and subparabolic in outline. Each lobe with the external border evenly arcuate; apex obtuse; internal mar- gin straight to feebly arcuate; surface glabrous and shining, not densely punctate, setose, setee moderately long and not dense. Mem- brane at bottom of the sinus, not setose. Female.—Genital segment elongately triangulo-parabolic in out- line. Valvula (Plate 4, fig. 16)—Dorsal plate elongate oblong, moder- ately narrow, sides subparallel and not reflexed; surface more or less feebly concave, glabrous, finely and sparsely punctate in apical third ; external margin nearly straight in basal three-fourths and rather moderately arcuate in apical fourth; internal border more or less sinuous; apex very short, subacute, very feebly chitinized and setose ; setee moderately long and fine, those on apical fourth of the dorsal plate sparsely distributed. Appendage short, more or less flattened and somewhat semicircular in outline, sete at tip rather short, and subapical in position upon the apex. Superior pudendal membrane longitudinally rugulose, attaining the apical fifth of the dorsal plate. Ventrolateral surfaces not inflated, surface lines straight when viewed longitudinally, regularly convex from side to side, glabrous and very sparsely punctate. Apex finely and sparsely setose. Sub- marginal groove rather broad beneath the more or less explanate external border of the dorsal plate, the latter sometimes more or less translucent. Internal margins of the valves contiguous in_ basal three-fourths; fissure apical and triangular between the slightly diverging ventral plates. Inferior pudendal membrane attaining the base of the apices and visible in the apical fissure. Habitat—Lower California. 316 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Forma typica.—Sierra Laguna and La Chuparosa. Forma oblonga.—San Francisquito. | Remarks.—Tibialis has been heretofore wrongly identified, and un- recognized as a distinct species. The following will correct the erroneous report as given by Doctor Horn. The specimens were in the collection of the academy, but were destroyed in the recent con- flagration (April 18, 1906) in San Francisco. Forma typica.—Female from Sierra Laguna labeled and reported as humeralis; two females from La Chuparosa reported as gracilis, and a female from the same place was also given as gracilis. Forma oblonga.—A male and female from San Francisquito, re- ported as humeralis. Number of specimens studied, 6. Type in my own collection (male), co-types in that of Mr. Fuchs. Ty pe-locality.— Sierra Laguna, Lower California. Salient type-characters.—Surface smooth and more or less aluta- ceous. Body elongate'and depressed. Prothorax widest at the mid- dle, sides not strongly arcuate, less so posteriorly, marginal bead en- tire; apical angles shghtly prominent anteriorly and narrowly rounded; basal angles obtuse and not rounded. Elytra with the humeri obtusely angled; surface finely, irregularly and_ obsoletely punctulate. Anterior tibiz strongly and suddenly constricted at base. Diagnostic characters —Tibialis is a unique and very distinct species in the United States fauna. I could not recognize it among the Mexican species given in the Biologia. Although aberrant it will have to constitute a group in the present subgenus until more material can be studied; it is not far out of place, as the first joint of the protarsi is clothed with yellow pubescence beneath and the genital characters are more in harmony here than elsewhere. It differs from all other species in its elongate, depressed, and subnyctobatoid form ; the protibiew are constricted at base in the typical form, but not in the oblong form. General observations —The mentum is moderate in size, parabolo- quadrate in form; inflexed lobes small, apex truncate or more or less arcuate and more or less deflexed; surface moderately densely pune- tate, punctures not crowded nor noticeably setigerous, laterally more or less feebly foveate and feebly convex centrally; base of the ligula rather more exposed than usual. The prosternum is not strongly prominent ventrally, evenly convex antero-posteriorly between the coxee and also behind, smooth and in the specimens before me not in the least produced posteriorly. 4 Proceedings, California Academy of Sciences, 2d Ser., IV, Pt. 1, pp. 849-350. REVISION OF ELEODIINI—BLAISDELL. 317 The mesosternum is subvertical in its anterior two-thirds and more or less arcuate; surface smooth, broadly and more or less concave. The abdominal process is quadrate (male) or slightly transverse (female) and about a fourth of its width wider than the metasternal salient. The post-coxal part of the first segment is longer than the process and equal in length to the third (male) or a little longer in the female; second segment is about twice as long as the fourth. The metasternum laterally between the coxe is as long as the width of a metafemur at base. The profemora are moderately clavate; the tibial grooves are rather wide externally, gradually and quite evenly narrowing in basal two- thirds; floors quite concave externally, margins distinct, irregularly and unevenly, minutely subcrenulate, becoming contiguous at base; both margins are more or less obsolete opposite the tibial constriction. The mesofemora are scarcely subclavate, although feebly and evenly tumid; grooves rather narrow, with margins as on the profemora, but less strongly marked and less sinuate at apex. Metafemora very feebly increasing in width from base to apex; grooves less developed than on the mesofemora, margins obsolete at middle and continued to base as lines of asperities. The tibize may be more or less feebly arcuate, most noticeably so in the protibie, the latter more or less carinate externally in basal half or two-thirds; tarsal grooves not present and the articular cavities are closed. The tarsi are apparently slightly elongate and moderate in stout- ness, The protarsi in the male are just noticeably stouter and slightly longer than in the female. They are about a third (male) or a fourth (female) of their length shorter than a mesotarsus. The mesotarsi are about an eighth (male) or a seventh (female) of their length shorter than a metatarsus, the latter about a half of its own length shorter than its metatibia. ELEODES SNOWII, new species. Oblong-ovate, more or less shining, about twice as long as wide; sides of the pronotum evenly rounded from apex to base; elytra more or less muricately granulate and not distinctly flattened on the dorsum. Head twice as wide as long, more or less feebly convex, more or less feebly impressed laterally, frontal suture more or less evident, rather finely and irregularly punctate, shghtly denser and coarser laterally and on the epistoma, sparser on the vertex. Antennw moderate, very ed feebly compressed and not noticeably incrassate, third joint about 318 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. equal in length to the next two combined, fourth to the seventh inclu- sive subequal, eighth subtriangular, ninth and tenth suborbicular, eleventh short ovate. Pronotum transversely oval, about one-half wider than long, widest at the middle; disc moderately and evenly convex, not very finely and quite densely punctate, denser and granulate laterally; @pea more or less feebly emarginate and’ more or less obsoletely margined; s?des quite evenly and moderately rounded, rarely just the least feebly sin- uate before the basal angles, marginal bead fine and entire; base quite truncate, more or less finely and obsoletely margined, subequal to or about a seventh wider than the apex, frequently a little shorter than the length; apical angles obtuse, narrowly rounded at tip and not prominent; basal angles obtuse and not rounded. Propleure more or less shining, irregularly muricato-granulate and rugulose. Elytra oval, less than twice as long as wide, widest at the middle; base not emarginate and evidently wider than the contiguous pro- thoracic base; Awmeri usually broadly rounded, sometimes obtusely and less broadly so; sides evenly arcuate, apex not very narrowly rounded; disc more or less feebly or moderately convex on the dorsum, laterally more strongly rounded, not suddenly inflexed, although somewhat so occasionally, arcuately declivous posteriorly; surface rather finely and submuricately punctate centrally, laterally and on apex muricato-granulate, granules and punctures irregularly but closely placed; punctures usually more or less simple at centre of the disc. Epipleure at middle as wide as the mesofemur at base, gradually narrowing from base to apex, superior margin straight beneath the humeri when viewed longitudinally from the apex; surface smooth, and obsoletely punctate. Sterna more or less densely punctate. Parapleure quite densely punctate. Abdomen shining, more or less densely punctate, more or less rugulose, frequently reticulately so; last segment occasionally rufous. Legs moderate in length and stoutness; anterior femora more or less sinuate in outer fifth and mutic; anterior tibiz more or less con- stricted at base, spurs quite similar in the sexes, the anterior slightly longer and stouter than the posterior; anterior tarsi with the first joint feebly thickened at tip beneath, and dissimilar in the sexes. Male.—Body somewhat slender, antenne attaining the base of the prothorax; elytra quite evenly, arcuately and feebly obliquely de- clivous posteriorly. Abdomen slightly oblique, not very moderately convex; first two segments broadly and not strongly impressed at middle. Anterior tarsi with the first two joints clothed with golden REVISION OF ELEODIINI—BLAISDELL. 319 pubescence at tips beneath, tuft of the second joints about one-half the size of the first. Female.—Rather robust, antenne scarcely attaining the base of the prothorax. Elytra somewhat broadly oval, arcuately and more or less vertically declivous behind. Abdomen horizontal, and rather strongly convex. Anterior tarsi without golden pubescence on the first two joints beneath, the produced tips clothed with the ordinary ferrugineous spinules that may interrupt the groove. Measurements— Males: Length, 14-17.5 mm.; width, 6-7 mm. Females: Length, 15.2-18 mm.; width, 7-8.5 mm. Genital characters, male-——KEdeagophore oblong fusiform; not arched. Basale oblong, scarcely three times as long as wide, evenly convex; sides quite parallel. Apicale small, nearly equilaterally triangular; surface moderately convex, without groove; sides more or less sinuate, tip acute; base broadly lobed, feebly sinuate laterally. Sternite transversely quadrate. Each lobe long and narrow, tri- angular; external margin sinuate basally, obliquely arcuate apically ; apex very narrowly rounded; internal border quite straight; sur- face very feebly convex, rather densely punctate in apical half, setose, setae moderately long. Membrane not setose across the bot- tom of the sinus, the latter broad. Female.—Genital segment triangular, slightly longer than wide, superior surface plane. Valvula (Plate 4, fig. 12).—Dorsal plate very slightly narrowed from base to apex, sides feebly convergent or parallel; surface plane, glabrous, sparsely punctate, sete not evident; margins nearly straight ; -apical margin more or less evenly rounded, often rather prominent internally. Apex long and slender, sometimes moderate, subacute, external lobe obsolete, finely setose. Appendage short mammilliform, clothed with a few short sete, three or four longer ones at tip. Fossa at the base and external sur- face of apex. Basal prominences not evident. Superior pudendal membrane reaching to near the apical margin of the dorsal plate, longitudinally rugulose. Ventrolateral surfaces as in tenebrosa. Habitat.——Colorado (Ouray, elevation 7,500-8,000 feet, July, H. F. Wickham); New Mexico (Santa Fé Canyon, elevation 7,000 feet, August, F. H. Snow; Cloudcroft, March and August, Warren Knaus); Arizona (Williams and Flagstaff, Barber and Schwarz; Oak Creek, August 2, Eug. Smith, from Knaus; along the Colorado River, Charles Fuchs). Number of specimens studied, 26, 320 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Sexitypes in my own collection. Ty pe-locality—Santa Fé Canyon, New Mexico; Prof. F. H. Snow, collector. : Salient type-characters——More or less shining. Sides of the pro- notum evenly arcuate from base to apex, surface not very finely punc- tate, punctures rather densely placed, denser laterally, where it is also granulate. Elytra convex not distinctly flattened, although somewhat so in the male; not very suddenly rounded, and inflexed laterally ; rather finely and submuricately punctate centrally ; muricato-granu- late laterally and on apex. Diagnostic characters.—In snowii the sculpturing is not strongly muricato-granulate as in deconte?, where the elytra are more muri- cato-tuberculately sculptured. In snow7i the tendency is toward a simpler form of muricate punctuation, and, besides, the form is more elongate and the elytra usually moderately convex. Snowii has not the facies of a planipennis, typical leconteé has. The side margins of the pronotum are at times similar in snowi?, inculta, tenebrosa, and horniit. In inculta the body is much more robust and the pronotum distinctly transverse, the elytral punctua- tion is rather finely muricate; fenebrosa is opaque and the elytra sculptured with fine shining granules; horni has the pronotum more narrowed at base, antenne longer, humeri absent, and the elytra slowly widening from the base. A series of specimens collected at Williams and along the Colorado River in Arizona are more elongate than the types or other New Mexican specimens, and several of them have the pronotal sides dis- tinctly and briefly sinuate in front of the basal angles. Others from the same place, and taken at the same time, have the sides evenly arcuate from apex to base; in every other respect they are identically the same; one female is more robust and quite like a female from Cloudcroft, New Mexico. Specimens from Colorado and New Mexico also show this variation; tenebrosa and inculta present analogous variations. A female example collected at Oak Creek, Arizona, and before me, is more robust, somewhat like a large variety of parvicollis, but from some specimens of nigrina of similar facies it 1s almost impos- sible to separate it, in fact it was impossible until I examined the genital segment, which agreed exactly with those from Williams and along the Colorado River. I have no hesitation in referring the above specimens to the present species, as there is nothing to warrant me in separating them; the genital characters agree. It might be suggested that those specimens with sinuate side pronotal margins approach consobrina, but the latter is more robust in form and with coarser sculpturing and more strongly punctate abdomen. I have never seen a true consobrina rts Aan ee anette Crema bee aga . ae REVISION OF ELEODIINI—BLAISDELL. ook east of the Colorado River. I would sooner believe that they were variations of tenebrosa. A single specimen taken in New Mexico and in the U. S. National Museum collection is robust and resembles consobrina in this respect, but I would still refer it to snowéi, even if I did not know the locality from which it came. A much larger amount of collecting must be done and larger series obtained before the relationship can be worked out, and besides we must learn something about hybrids and atavistic reversions. The Arizona specimens are not as strongly sculptured, and more shining, with the humeri more obtusely angled than the New Mexican examples, which have the humeri broadly and evenly rounded. The typical, sinuate, and robust forms must be recognized and so labeled in our collections. General observations—The mentum is somewhat small and para- bolic or trapezoido-parabolic in outline, the apex may be slightly deflexed ; the surface is feebly convex, very slightly foveate laterally and not strongly punctate, the punctures are not noticeably setigerous. The prosternum is more or less prominent ventrally with the coxe, sometimes very slightly so; horizontal between the coxee, submu- cronate to vertically truncate behind, or arcuately convex antero- posteriorly ; surface more or less longitudinally grooved between the coxe. The mesosternum is arcuately convex and more or less broadly concave. The metasternum laterally between the coxe is as long as the width of a mesotibia at apex. The abdominal process is quadrate (male) or very shghtly trans- verse (female) and from a fourth to a third of its width broader than the metasternal salient. The post-coxal part of the first abdominal segment is equal in length to the process and aiso to that of the third; the second is about one-half (male) or not twice as long (female) as the fourth. The tarsi are moderate, the anterior in the male is noticeably shehtly longer than in the female. The protarsi are about one-half (male) or a fourth (female) of their own length shorter than a mesotarsus. The mesotarsi are subequal to (male) or a little shorter (female) than a metatarsus. The metatarsi are about two-thirds of their length shorter than a metatibia. ELEODES LECONTEI Horn. Eleodes lecontei Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 316. Hleodes subaspera LECONTE, Smithson. Miscell. Coll., 167, Pt. 1, 1866, p. 114. Ovate, about twice as long as wide, surface rather dull, elytra dis- tinctly flattened and muricately punctured, 59780—Bull. 68—09——21 322 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Head about twice as wide as long, scarcely convex between the eyes, very feebly impressed laterally, punctures rather fine, some- what densely placed and quite evenly distributed. Antenne long, rather slender, outer three joints very feebly compressed and not dilated, third joint scarcely as long as the next two combined, fourth just noticeably longer than the fifth, the latter, sixth, and seventh subequal in length, eighth just the least shorter, slightly wider and subtriangular, ninth and tenth suborbicular, eleventh irregularly short-ovate. Pronotum somewhat transversely oval, about a third wider than long; disc moderately and evenly convex, rather coarsely and densely punctate, the punctures crowded and granular laterally ; apex scarcely emarginate and more or less obsoletely beaded; sides rather strongly rounded, scarcely briefly sinuate before the basal angles and finely beaded; base truncate to feebly arcuate, obsoletely margined, about a seventh wider than the apex and quite equal to the length; apical angles distinct and subacute, not in the least prominent anteriorly; basal angles obtuse. Propleure rather densely punctate and rugulose. Elytra oval, about a half wider than the thorax, widest at the middle; base truncate; humeri very obtuse and rounded; sides evenly arcuate, apex rather narrowly rounded; disc very feebly convex, typically depressed, rather suddenly and somewhat narrowly rounded laterally and inflexed, dorsum frequently more strongly convex, arcuately declivous posteriorly; surface muricato-granulately punc- tate, punctures irregularly and quite densely placed, becoming dis- tinctly granular and setigerous laterally and on apex, setee short and inconspicuous, the granules are small and_ scarcely subseriately arranged. Epipleure moderate in width, gradually narrowing from base to apex, superior margin quite straight beneath the humeri; surface more or less obsoletely punctate. Sterna rather densely punctate and more or less rugose. Parapleure rather densely but not strongly punctate. Abdomen more or less shining, quite evenly and not very densely punctate, punctures denser on the fifth segment, the first more strongly sculptured. Legs moderate in length and somewhat slender; anterior femora mutic; anterior tibial spurs rather short and apparently subequal in length and stoutness, and probably similar in the sexes; anterior tarsi dissimilar in the sexes. Male-—Unknown to me. Female—Body rather broadly ovate, antenne reaching a little beyond the base of the prothorax. Elytra arcuately and more or less vertically declivous behind; abdomen horizontal and rather REVISION OF ELEODIINI—BLAISDELL. a2 strongly convex; anterior tibize somewhat constricted at base; an- terior tarsi with the first joint shghtly thickened at tip beneath, and there clothed with a small tuft of piceous spinules. Measurements —Female: Length, 16.2 mm.; width, 8 mm. Genital characters, male.-—Not studied. Female.—Genital segment triangular and somewhat depressed, glabrous, very sparsely setose, and well chitinized. Valvula (Plate 5, fig. 21).—Dorsal plate suboblong, slightly widest at base, external border slightly convergent apically and slightly explanate; surface somewhat outwardly declivous, feebly convex but shghtly concave before the apex, glabrous, with a few coarse, scattered, setigerous punctures, sete short; external border feebly arcuate or straight, passing arcuately into the apical border, the latter defined from the apical surface, angle not evident; internal border more or less feebly sinuous. Apex fully chitinized, feebly convex above, scarcely excurved, rather broadly rounded at tip, with a number of moderately short sete over the base along the apical margin of the dorsal plate; in- ternal valvular membrane densely clothed with very short sete; fossa an excavation at the base of the external surface of the apex and beneath the apical margin of the dorsal plate. Appendage small and mammilliform, not conspicuous, with a tuft of moderately long sete at tip. Superior pudendal membrane longitudinally rugulose and attaining the apical third of the dorsal plate. Ventrolateral surfaces flattened centrally with the surface lines straight when viewed longitudinally, laterally moderately arcuate and sparsely setose; surface glabrous; submarginal groove distinct beneath the slightly explanate external border of the dorsal plate and terminating in the fossa. Apex somewhat concave beneath and somewhat densely set with short sete. Internal margins of the valves contiguous at basal two-thirds; fissure apical, apices not con- tiguous. Inferior pudendal membrane not visible. Habitat.—Colorado (no special locality indicated by the labels). Number of specimens studied, 1 female, which was carefully com- pared with type and pronounced typical by Mr. Blanchard; the specimen is in his collection. Type in the LeConte collection. T ype-locality— Colorado. Salient type-characters—Somewhat shining. Head and thorax thickly and finely punctate, the latter with apex scarcely emarginate; apical angles acute, not prominent; sides finely margined and strongly rounded, not sinuate posteriorly; base strongly truncate and scarcely narrower than the apex; basal angles obtuse. Elytra with the dor- sum slightly convex, apex strongly declivous, sides suddenly inflexed, 324 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. with subseriately placed small granules, on each side and apex briefly piliferous. Antenne slightly incrassate externally. Length 60 (subaspera) (LeConte). Diagnostic characters —The present species wes originally de- scribed by LeConte as suwbaspera; this name having been used by Solier at an earlier date. Doctor Horn renamed it in honor of Doctor LeConte. The single female specimen before me has been kindly loaned for study and pronounced by Mr. Blanchard as being exactly typical. In its general facies it is suggestive of planipennis. The elytra are distinctly flattened on the dorsum, and the sides of the pronotum are evenly rounded nearly to the basal angles where they become slightly subsinuate. Lecontei is much less shining than planipennis and the sculpturing is distinctly more asperate; the head and thorax are more coarsely punctate. Although the male is unknown to me, it is safe to as- sume that the secondary sexual characters are the same as In snowwiél. I believe that the elytra are more fortuitously depressed than nor- mally so and that the average specimens will have the elytra nor- mally convex. I surmise that I have examined such examples and erroneously referred them to snow before I fully recognized the species under consideration. The genital characters in the female are the extreme for the sub- genus Blapylis, the valvular apex is fully chitinized, it is also setose and not produced, characters not observed in planipennis from which it is perfectly distinct and not at all closely related. From snow?, which appears to be worthy of specific standing—if not, at least it would be a good race of decontec—it must be carefully differentiated. For a long time I have confused the two species; the sculpturing is quite similar in the two, but much less strongly and coasely granulate and more of the muricate type in snowii, where also the elytra are less broadly oval and more elongate and shining. In parvicollis and cordata and their races the pronotum is more or less strongly constricted at the base, less so in consobrina, where the body is more robust. In lecontei the marginal bead of the pronotum attains the base. General observations—The mentum is moderate and rather para- bolic in outline, finely and not deeply punctate, very feebly convex, and obsoletely foveate. The prosternum is moderately prominent ventrally with the coxe, arcuately convex antero-posteriorly, and not in the least mucronate behind. The mesosternum is feebly arcuate and oblique, very slightly con- cave, REVISION OF ELEODIINI—BLAISDELL. 325 The metasternum laterally between the cox is as long as the width of a mesotibia at the apex. The intercoxal process of the abdomen is slightly transverse and about a third of its width broader than the metasternal salient. The post-coxal portion of the first abdominal segment is equal in Jength to the process and also to that of the third; the second is about twice as long as the fourth. The protarsi are about a fourth of their length shorter than a meso- tarsus; the mesotarsi are about a fifth of their length shorter than a metatarsus; the metatarsi are about three-eighths of their length shorter than a metatibia. ELEODES TENEBROSA Horn. Eleodes tenebrosa Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 316. Oblong-oval, about two and a third times longer than wide; elytral sculpturing consisting of very small shining tubercles arising from a very opaque base. Head twice as wide as long, feebly convex, moderately impressed laterally and frequently along the frontal suture, which is usually not evident, not coarsely and more or less densely punctate, punctures distinctly crowded laterally and on epistoma. Antenne moderate in length, slightly robust, outer four joints very feebly compressed and just noticeably wider, third joint equal to the next two taken together, fourth just the least longer than the fifth, the latter, sixth, and seventh subequal in length, eighth about as wide as long and subtri- angular, ninth subtrapezoidal, tenth suborbiculo-trapezoidal, eleventh frequently shghtly smaller and short-ovate. Pronotum transversely oval, widest at the middle, a third to a half wider than long; disc moderately and quite evenly convex, rather coarsely and densely punctate, granulate at the sides; apex slightly emarginate, and more or less obsoletely margined; sides evenly and strongly arcuate from apex to the basal twelfth, then rather quickly sinuate and nearly straight and parallel to basal angles, more rarely evenly arcuate from angle to angle, marginal bead moderate, feebly reflexed and entire; base subtruncate in circular are and finely mar- gined, slightly wider than the apex, and not quite equal to the length; apical angles obtuse and more or less narrowly rounded; basal angles shghtly prominent and quite rectangular, sometimes subobtuse. Propleure smooth and opaque, sparsely and quite evenly granulate, acetabular convexities frequently rugulose. Elytra oblong oval, about a half longer than wide, scarcely at all wider than the prothorax, widest at the middle; base not emarginate; humeri quite broadly rounded, angle sometimes obtusely indicated ; sides evenly and not strongly arcuate, frequently subparallel, apex ob- 326 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. tusely rounded and not very narrowly so, not in the least produced ; disc more or less feebly to modertaely convex on the dorsum, sides evenly and moderately rounded, arcuately declivous behind; surface ‘rather densely and irregularly covered with small, rounded, shining tubercles, arising from a very opaque base. Epipleure smooth, opaque, sparsely and evenly subasperately punctate, moderate in width, superior margin scarcely to feebly and broadly sinuate beneath the humeri, as viewed longitudinally. Sterna quite densely punctate. Parapleure quite densely and coarsely punctate. Abdomen somewhat shining, smooth, and rather densely punctate ; fifth segment frequently more or less rufous; intercoxal process at times quite rugose, Legs moderate and quite strongly sculptured; anterior femor: mutic; spurs of the anterior tibiz quite similar in the sexes, the anterior slightly longer than the posterior; anterior tarsi dissimilar in the sexes, middle tarsi similar. Male—Somewhat narrow. Antenne attaining the prothoracic base. Elytra arcuately declivous behind. Abdomen slightly oblique and quite moderately convex, broadly impressed at middle of the first two segments. Anterior tarsi with the first jot slightly thick- ened at tip beneath and invested with a moderately obtusely pointed tuft of golden, silken pubescence, which is inclosed by a few ordi- nary spinules; second joint very feebly thickened with tuft not evi- dent, or at most scarcely half as large as that of the first. Female.—Rather broadly oblong. Antenne not quite attaining the prothoracic base. Elytra arcuately and vertically declivous posteri- orly. Abdomen horizontal and rather strongly convex. Anterior tarsi with the first jomt shghtly thickened at tip beneath, groove interrupted by ordinary piceous or ferruginous spinules. Measurements.—M ales: Length, 14-16 mm.; width, 5.5-6.5 mm. Females: Length, 15-18.5 mm.; width, 7-7.5 mm. Genital characters, male. not arched. Basale oblong oval; surface rather feebly convex, with the sides moderately arcuate. Edeagophore flattened oblong-ovate and Apicale rather small, triangular, and about a half longer than wide; surface evenly convex, not grooved; sides somewhat arcuate in basal sixth, thence feebly and very broadly sinuate to apex, the latter subacute; base broadly lobed at middle and feebly sinuate laterally. Sternite transversely parabolic. Each lobe with the outer border more or less evenly arcuate; apical margin broadly and evenly rounded; internal margin feebly arcuate; surface very feebly con- vex, very sparsely punctate and setose, setee moderately long, shghtly REVISION OF ELEODIINI—BLAISDELL. 321 longer on apical margin, impunctate in basal fourth. Membrane not setose across the bottom of the sinus, the latter narrowly triangular. Female.—Genital segment elongately triangular, superior surface plane. Valvula (Plate 1, fig. 9).—Dorsal plate elongately oblong; surface plane, glabrous, obsoletely punctate and sparsely setose; sides more or less feebly arcuate and subparallel; apical margin feebly arcuate or subtruncate, scarcely rounded internally over the apical lobe, angle evenly rounded. Apex rather long and slender, subacute and finely setose, with a few long sete at tip; external apical lobe not in the least evident. Appendage short mammilliform, with about two to four long sete at tip. Fossa in basal and external surface of apex. Superior pudendal membrane reaching to the apical fourth of the dorsal plate and longitudinally rugulose. Basal prominences not evident, evenly rounded at base. Ventrolateral surfaces (Plate 1, fig. 10).—Body quite triangular and evenly convex from side to side, glabrous, surface lines straight as viewed longitudinally, very feebly punctate and setose about the apex. Submarginal groove moderate beneath the slightly explanate ex- ternal border of the dorsal plate and attaining the fossa. Internal margins of the valves contiguous in about basal three-fourths; genital fissure very narrow in apical fourth; the inferior pudendal mem- brane is not visible. Habitat—California (Owens Valley, Dr. George Horn; Bodie, elevation 8,475 feet, July, Wickham; Lundy, 7,000-8,000 feet, July) ; Nevada (western, Mr. Gabb; Verdi, April, Blaisdell). Number of specimens studied, 125. Type in the Horn collection. Type-locality—Owens Valley, California; collector, George Horn. Salient type-characters—Opaque. Head rather coarsely punc- tured. Thorax one-third broader than long, very coarsely punc- tured, becoming granular at the sides; sides strongly rounded, mar- gin entire, basal angles moderately prominent, rectangular. Elytra elongate oval, moderately convex, sides moderately rounded; basal angles obtuse; surface rather densely and irregularly covered with small, rounded, shining tubercles, arising from a very opaque base (Horn). ; Diagnostic characters—The salient type characters differentiate tenebrosa from all other species of the subgenus—in fact it is quite unique among its congeners by not having the elytra broader at the widest part than the thorax. Occasionally specimens are observed with the sides of the pro- notum evenly rounded from base to apex and in this character resem- ble snowii, but here the surface is more or less shining and the 328 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. pronotal dise is less coarsely punctured. These remarks apply especially to the Arizona or elongate form of snowi7i. I have seen a few specimens that were rather ovate in form, with the elytra showing a tendency to become less elongate and wider than the prothorax. Such variations are expected and must be recognized. General observations —The mentum is usually quite parabolic in outline, sometimes subtriangular; surface very shghtly convex, not coarsely but quite densely punctate laterally, where it is sometimes subfoveate; the punctures are not noticeably setigerous. The prosternum is variable, more or less prominent ventrally with the cox, and moderately widened behind the axes of the acetabula, frequently grooved at the middle. In the more strongly developed individuals it is horizontal between the coxe and mucronate behind; in others, arcuate antero-posteriorly and with a more or less well- developed mucro at the middle of the posterior border; again, it is evenly arcuate and not at all mucronate. The mesosternum is vertically arcuate, or more obliquely so, deeply concave, appearing somewhat bilobed; the concavity varies in depth. The metasternum laterally between the cox is as long as the width of a protibia at apex. The abdominal process is quadrate and about a fourth of its width wider than the metasternal salient, and equal in length to the post- coxal portion of the first segment; also about equal to the length of the second segment (male). In the male the second segment is one-half longer than the fourth, and the third about a third longer than the same segment. In the female the second is twice as long as the fourth and about a fourth longer than the third. The tarsi are moderate in length and somewhat stout. The protarsi are about two-fifths (male) to three-eighths (female) of their length shorter than a mesotarsus. The mesotarsi are about a seventh (male) to an eleventh (female) of their length shorter than a metatarsus. The metatarsi are about three-eighths (male) to two-thirds of their length shorter than a metatibia. ELEODES TENEBROSA var. NANA, new. As in tenebrosa, except smaller, more strongly and densely sculp- tured. Sides of the pronotum slightly tuberculate, margin more strongly sinuate before the basal angles. Elytral tubercles larger and more distinct, more evidently setigerous at the sides; humeri obtuse, scarcely rounded to subrectangular, with the epipleural margin often visible at the angle. Measurements.—M ales: Length, 10.5-12.5 mm.; width, 5-5.5 mm. Females: Length, 12 mm.; width, 6 mm. REVISION OF ELEODIINI—BLAISDELL. 329 Genital characters as in tenebrosa. Habitat.—Nevada (Carson City, July, Wickham; Verdi, April, Blaisdell) ; California (U. S. National Museum collection—no exact locality given). Diagnostic characters——Nana is more coarsely sculptured; the tu- bercles of the elytra are denser, confluent, and submuricate, but most of them still remain bright and shining. The humeri are more often subangulate. This form inhabits the same region as the typical race. ELEODES INCULTA LeConte. Eleodes inculta LEContTE, Proc. Acad. Nat. Sci. Phila., 1861, p. 352, male.— Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 315. Robust, oblong ovate, about twice as long as wide, black, often somewhat piceous, feebly shining to somewhat dull; elytra irregu- larly and subasperately muricately punctate. Head about twice as wide as long, feebly convex, more or less broadly impressed laterally and along the frontal suture, the latter more or less evident; irregularly and not very finely punctate, rather densely so laterally and on the epistoma. Antenna moderately long, somewhat slender, outer four joints feebly compressed and feebly incrassate, third jeint scarcely as long as the next two taken together, fourth just the least longer or subequal to the fifth, the latter, sixth, and seventh subequal, eighth just a little shorter and more or less subtriangular, ninth and tenth suborbicular in outline or very feebly transverse, eleventh short-ovate. Pronotum more or less transversely quadrate, a third to a half wider than long, widest at the middle; disc moderately and very evenly convex, distinctly, irregularly, and not coarsely punctate, punctures separated by an interval equal to or twice their own diame- ters, narrowly subgranulate laterally; apea broadly, evenly, but not strongly emarginate, finely and somewhat obsoletely margined; sides evenly, broadly, and not strongly arcuate, very feebly subsinuate before the basal angles, and less so behind the apical angles, marginal bead fine and entire; base very feebly and broadly rounded, sub- truncate, rather coarsely margined, one-sixth to one-third wider than the apex, equal to or a little greater than the length; apical angles subacute, shghtly prominent anteriorly, and with a slight tendency to eversion; basal angles obtuse and not in the least rounded, just in the least prominent. Propleure smooth and shining, sparsely and distinctly submuri- cately punctate and rugulose. Elytra oval, less than a third longer than wide, widest at the mid- dle; base subtruncate. Zumeri more or less broadly rounded, epi- pleural margin not visible from above; sides evenly arcuate, apex 330 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. not very narrowly rounded; disc moderately to rather strongly convex on the dorsum, sides not broadly rounded, inflexed portions scarcely arcuate, arcuately declivous posteriorly; surface irregularly and quite densely, more or less distinctly subasperately and muri- rately punctate, sculpturing not strong, simple centrally and not distinctly granulate laterally and on the apex. Epipleure moderately wide, gradually narrowing from base to apex, superior margin not strong and almost straight beneath the humeri; surface smooth, impunctate toward base, obsoletely and sparsely punctate toward apex. Sterna.—Prosternum scabrously rugoso-punctate; meso- and meta- sterna more or less shining, glabrous and rugoso-punctate. Parapleure rather densely, more or less evenly, and somewhat coarsely punctate. Abdomen shining, sparsely and finely punctate, more or less rugulose. Legs rather short and somewhat slender; anterior femora mutic and not noticeably sinuate; tibial spurs rather slender and moderate in length, similar in the sexes; anterior spurs of the protibie shghtly longer than the posterior; anterior and middle tarsi dissimilar in the sexes. Male.—Oblong-ovate, somewhat slender. Antenne reaching a little beyond the base of the prothorax. Elytra evenly and arcuately de- clivous posteriorly. Abdomen slightly oblique apically, moderately convex, segments one and two broadly impressed at middle. First three joints of the protarsi not noticeably thickened and clothed with golden pubescence beneath, surface of the pads flat; first two Joints of the mesotarsi similarly pubescent, first joimt in apical half, the second with a much smaller tuft at tip. Female.—Distinctly robust. Antenne about attaining the base of the prothorax. Elytra broadly oval, arcuately and more or less verti- cally declivous behind. Abdomen horizontal and strongly convex. First two joints of the anterior and middle tarsi a little more pubes- cent than usual beneath, grooves not evident. Measurements —Males: Length, 13.5-14 mm.; width, 6.5-6.5 mm. Females: Length, 14-17 mm.; width, 7.5-8 mm. Edeagophore flattened oblong-ovate, Genital characters, mate. acutely pointed at apex and scarcely arched. Basale oblong-suboval, moderately convex, sides feebly arcuate and nearly parallel. Apicale triangular. Surface very moderately convex, without groove; sides briefly arcuate at base, thence broadly and more or less strongly sinuate, so that the apex appears more or less attenuated in apical half; base evenly rounded and scarcely lobed at middle. REVISION OF ELEODIINI—BLAISDELL. 831 Sternite parabolic and slightly transverse. Each lobe triangular; external border slightly oblique, and more or less feebly sinuate in basal half, thence subtruncate and feebly arcuate to apex, the latter moderately rounded; internal border more or less sinuate; surface feebly convex, rather coarsely and densely punctate in apical three- fourths, sete rather long, denser on apical margin. Female.—Genital segment (Plate 5, fig. 18) triangular, about a third longer than wide; surface about plane, finely setose on apex. Valvula—Dorsal plate oblong, elongate, about three times longer than wide; surface plane, more or less declivous laterally at base, smooth and shining, very sparsely and finely punctulate in apical half, setee very fine and not distinct; external border about straight, feebly sinuous and very feebly converging apically; apical margin more or less arcuate and rather distinct from apex; internal margin quite straight. Apex short and scarcely more prominent than the appendage, subacute, membranous, finely setose, with a few rather long sete at tip. Appendage short mammilliform, with a pencil of quite long sete at tip. Tossa nearly occupying the external surface of the apex. Basal prominences not evident. Superior pudendal membrane scarcely longitudinally rugulose and attaining the apical fourth of the dorsal plate. Ventrolateral surfaces.—Body triangular, surface lines straight as viewed longitudinally, smooth and shining, feebly punctate and setose in apical third; submarginal groove distinct beneath the shehtly explanate external border of the dorsal plate. Internal margins of the valves contiguous in basal three-fifths, fissure in apical two-fifths, narrow and nearly closed. Habitat.—Nearly all of the specimens before me were taken on Santa Rosa Island, which hes off the coast of Santa Barbara County, California. One specimen bears a “ Santa Barbara Co.” label and another simply a “ Cal.,” while in Mr. Fuchs’ collection is one labeled “Alameda County.” There are no differences noticeable in the speci- mens. Iam not sure that this species is found on the mainland. Number of specimens studied, 19. Type a male in the LeConte collection. Ty pe-locality—Santa Rosa Island; collector, Mr. C. M. Bache. Salient type-characters——Subopaque. Head and thorax rather densely punctate; thorax with the sides strongly rounded, briefly sinuate behind; basal angles obtuse and slightly prominent; base broadly rounded. Elytra oval, apex attenuate and strongly de- clivous; humeri broadly rounded, dorsum deplanate, punctures small and granular, irregularly placed, granules on each side briefly pilifer- ous. Antenne a little incrassate at tip (LeConte). 332 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Diagnostic characters—Inculta is a distinct species, with a shorter, more robust form, distinctly transverse prothorax (female), and feebly subasperate (subgranular) punctuation. The sides of the prothorax are scarcely at all sinuate before the basal angles—in some specimens not in the least and in others feebly so; the basal angles are obtuse, distinct, and with the appearance of being just a little prominent; the lateral bead is distinctly entire. I do not believe inculta to be a synonym of scabripennis. The type of the latter is a female and was taken at Fort Tejon. Doctor Horn made so many wrong determinations in /’/eodes that I can not be- heve him right in this instance. Mr. Blanchard has very carefully compared a female inculta (which is before me) with the female type of scabripennis. His com- ments are as follows: ‘ Shorter and less convex than the type of scabripennis, thorax much broader. It seems nearer inculta of which the type is a male with base of the prothorax more constricted.” Later and in another sending I submitted a male specimen, which he unhesitatingly pronounced inculta. I sent the female first inten- tionally with a request to compare it with scabripennis. The form of the thorax in snow7di (male and female) and the male of inculta is quite similar, but the apical angles in the latter are more acute and prominent, and in every example studied the sides are just the least subsinuate behind the angles so as to give them an appearance of having a slight tendency to eversion. The form of the prothorax should readily separate inculta from hornii, neotome and consobrina—none have it so transverse in the female, nor the sides so feebly sinuate at base as a constant character. In consobrina the apical angles are as prominent, but scarcely as acute, the sides may be feebly sinuate behind the apical angles and more or less strongly sinuate before the base. General observations.—The mentum is moderate in size, sometimes rather small in the male, subtruncate and parabolically rounded; surface feebly convex, scarcely subfoveate laterally, finely punctate, the punctures not distinctly defined nor noticeably setigerous. The prosternum is moderately prominent ventrally with the cox, moderately arcuate antero-posteriorly between the coxee and usually with a small mucro behind at middle; at other times more feebly arcuate and more or less vertical behind, with the angle mucroid. The mesosternum is arcuately declivous and more or less deeply and not broadly concave. The metasternum laterally between the coxve is as long as the width of a mesotibia at apex. The abdominal intercoxal salient is subquadrate (male) or slightly transverse (female) and a little (male) to a fourth (female) of its own width broader than the metasternal process. REVISION OF ELEODIINI—BLAISDELL. 333 The post-coxal portion of the first abdominal segment is equal in length to the process, also to that of the third (male). In the male the second segment is a third of its length longer than the fourth; in the female the second is twice as long as the fourth and a little longer than the third. The tarsi are slender and moderate in length. The protarsi are about two-fifths (male) to a fourth (female) of their length shorter than a mesotarsus. The mesotarsi are about a seventh (male) to a fifth (female) of their length shorter than a metatarsus. The metatarsi are about a fourth (male) to two-thirds (female) of their length shorter than their respective metatibie. ELEODES CONSOBRINA LeConte. Eleodes consobrina LEConTE, Ann. Lye. Nat. Hist. N. Y., V. 1851, p. 185.— Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 316. Hleodes veseyi LECoONTE, Proc. Acad. Nat. Sci. Phila., 1858, p. 187. Oblong-ovate, robust, elytra coarsely and submuricately punctate centrally and tuberculate laterally; surface dull or feebly shining. Head about twice as wide as long, more or less feebly convex, slightly impressed laterally and often along the frontal suture, which is generally evident as a more or less smooth line, sides of the frons not usually prominent, more or less evenly, rather densely and some- what finely punctate, punctures slightly denser laterally. Antenne rather stout, moderate in length, scarcely compressed, very feebly and gradually widened in outer four joints, third joint comparatively short and scarcely as long as the next two combined, fourth to the eighth, inclusive, quite equal, ninth and tenth feebly transversely oval, eleventh short-ovate. Pronotum somewhat transversely’ oblong-oval in outline, two- sevenths to a third wider than long, widest at the middle; disc mod- erately to rather strongly and evenly convex, quite densely, evenly, or irregularly punctate, punctures moderately coarse, becoming granu- late at the sides; apex evenly and moderately emarginate, obsoletely margined; sédes more or less strongly arcuate and constricted at basal twelfth or eighth, thence briefly straight or feebly oblique to the basal angles, marginal bead rather thin and fine, at times somewhat coarser and more or less entire and reflexed; base slightly rounded and not very coarsely beaded, equal to or a fourth wider than the apex and about equal to the length; apical angles subacute to feebly rounded and rather prominent anteriorly; basal angles minute, obtuse or rec- tangular, feebly prominent at times. Propleure opaque, not densely muricato-granulate, and more or less rugulose about the acetabular convexities, 3384 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Elytra suboblong-oval, about a half longer than wide, widest at the middle; base rather truncate and more or less sinuate. Each side of the middle equal to or distinctly wider than the contiguous prothoracic base; Awmeri more or less obtusely rounded; sides evenly and moderately arcuate, apex more or less feebly attenuate, not very narrowly rounded; disc moderately convex, not noticeably flattened, evenly and strongly rounded laterally, arcuately declivous posteriorly ; surface irregularly, densely and coarsely, slightly submuricately punctate on the dorsum, sides and apex densely sculptured with rec- linate granules or tubercles, rarely the punctures are feebly subseriate on the dorsum each side of the suture. Epipleure comparatively moderate in width, gradually narrowing from base to apex, superior margin nearly straight—at most but feebly sinuate beneath the humeri, somewhat decurving apically, rarely visible from above at the humeri; surface very sparsely sub- muricately punctulate. Sterna densely punctato-rugulose. Parapleure more or less shining and coarsely punctate. Abdomen more or less shining, more or less densely and coarsely punctate, more strongly so and rugulose to rugose about the coxe, less densely and more finely sculptured on the last two segments. Legs moderate in length, somewhat stout, and densely but not very coarsely sculptured; anterior femora mutic; tibial spurs similar in the sexes, the anterior of the protibize slightly longer than the pos- terior; tarsi dissimilar in the sexes. Male—Oblong-ovate, somewhat narrow. Antenne scarcely reach- ing beyond the prothoracic base. Elytra somewhat attenuate in apical fourth, arcuately and rather obliquely declivous behind. Ab- domen more or less oblique and moderately convex, more or less broadly impressed at middle on the first two segments. First two joints of the protarsi slightly thickened at tip beneath, and there clothed with blunt tufts of yellowish pubescence, the second smaller than the first; first jot of the mesotarsi with a similar but moderate tuft, obliterating the groove. Female.—Robust, ovate. Antenne scarcely attaining the protho- racic base. Elytra arcuately and quite vertically declivous behind. Abdomen horizontal and rather strongly convex. First joint of the protarsi slightly thickened at tip beneath, groove almost entire. Measurements.—Males: Length, 15-16.5 mm.; width, 6-6.5 mm. Females: Length, 16-19 mm.; width, 7.5-9 mm. Genital characters, male.—Kdeagophore flattened oblong-ovate, very slightly arched. Basale oblong, slightly more than twice as long as wide; sides feebly arcuate, subparallel; surface moderately convex. REVISION OF ELEODIINI—BLAISDELL. 33D Apicale triangular, moderately convex above, without groove; sides slightly prominent at base, thence feebly sinuate to tip, the latter subacute; base with a moderate and rounded lobe at middle, feebly arcuate laterally. Sternite transversely parabolic. Each lobe triangular; surface glabrous, feebly convex, impunctate in basal third, punctate else- where, more densely so about apex, setose, setee moderately long; ex- ternal border obliquely arcuate; apex narrowly rounded, and con- tinuously so with the internal border which becomes sinuate. Mem- brane not setose, sinus rather broad and triangulo-oval. Female.—Genital segment parabolo-triangular, rather depressed, dorsal surface quite plane and not noticeably setose, except at apex, glabrous. Valvula (Plate 4, figs. 9 and 10).—Dorsal plate oblong, quite hori- zontal, with the sides subparallel; surface plane or very slightly concave, obsoletely punctulate; internal and external borders shghtly arcuate; apical border more or less evenly rounded, angle not in the least evident. Apex triangular, rather broad, very finely setose, sete at tip moderately long. Fossa at base of the external surface of the apex behind the dorsal plate. Appendage short mammilliform, with a pencil of quite long sete at tip. Superior pudendal membrane longitudinally rugulose and rather broadly exposed, attaining the apical fourth of the dorsal plate. Basal prominences very smal). Ventrolateral surfaces triangularly plane at center to base; surface lines quite straight as viewed longitudinally and moderately convex transversely; surface smooth and impunctate. Submarginal groove well developed beneath the distinctly explanate external margin of the dorsal plate, extending obliquely across the base of the apex as a shallow concavity; apex finely setose. Internal margins of the valves contiguous in basal three-fourths; genital fissure in apical fourth and rather narrowly fusiform. Inferior pudendal membrane not visible. Habitat—California (San Diego County; San Bernardino Moun- tains; Los Angeles County; Santa Barbara County; Fort Tejon, Fuchs and Hopping; San Francisco County; Shasta and Siskiyou counties). Number of specimens studied, 45. Type.—A male in the LeConte collection. Type-locality—Mountains about Santa Isabel, California. Salient type-characters—Oblong. Thorax rounded, apex emar- ginate, thickly punctate, sides granulate, margin reflexed, posterior angles minute. Elytra thickly and coarsely punctate, sides and pos- teriorly rough and reclinately granulate, apex attenuate (LeConte). 336 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Diagnostic characters —It would appear from a careful study of LeConte’s description that the type (male) had the sides of the pronotum rounded nearer to the base than in the majority of the specimens before me, in consequence there is a less degree of con- striction and the angles smaller; such is the case in a specimen which has been compared with the type by Professor Fall, and in his col- lection, and which I take to be quite typical. The example was col- lected in the San Bernardino Mountains. In all of the specimens making up a small series taken at Fort Tejon by Fuchs and Hopping, the sides of the pronotum are more strongly constricted, straight, and parallel in about the basal eighth; the angles are therefore larger and rectangular, but some of the specimens approach Professor Fall’s specimen, so that this character is simply a variation and of no specific value. The Fort Tejon specimens no doubt belong to the form described by LeConte as veseyi, the type of which is a large male and remark- able for the attempt at strie of large punctures near the middle of the elytra (LeConte). Mr. Blanchard writes me that veseyi is so near the type of consobrina that he is surprised that LeConte should have given it a name; it is no doubt a heterotype and therefore a true synonym of consobrina. In veseyi the pronotum is more strongly constricted and the basal angles larger and subrectangular; therefore it 1s related to the specimens above mentioned from Fort Tejon. An interesting specimen from Lower California was referred to consobrina by Doctor Horn; it was destroyed with the Academy in the recent disaster before I had an opportunity to study it. It is very difficult oftentimes to satisfactorily separate this species from some forms of parvicollis; as a rule in consobrina the pronotum is more strongly and evenly convex from side to side, and not at all impressed or flattened within the lateral margins; in parvicollis, when the pronotal surface is evenly arcuate from side to side, it generally arises more suddenly and strongly from the margins. In consobrina the humeri are more rounded and the superior epipleural margin not visible from above; the elytral sculpturing is also coarser and tuberculate. These characters are valuable when the pronotal ones are weak or exhibit an approach to parvicollis, as they do sometimes. The heterotypes of both species are at times no doubt amphitypical. In consobrina the marginal bead is not so reliable a character as in tenebrosa, inculta, and snowii, for instance; the species is always more robust. It is often necessary to examine the genital segment of the female to more satisfactorily place some particular specimen, and males from the same region can be placed with the females. @Pproc. California Acad. Sciences, 2d ser., IV, Pt. 1, p. 350, REVISION OF ELEODITNI—BLAISDELL. 837 Specimens from the Farallone Islands have been determined as consobrina by Doctor Horn, but I prefer to consider them as a form of parvicollis, for the following reasons: The general facies is more like the latter species, although in some specimens the marginal bead is entire, the specimens are decidedly less convex, and the pronotal dise is feebly impressed laterally, the basal constriction is stronger, and the sculpturing is plainly more muricate laterally than tuberculate. Again, parvicollis is much more abundant along the coast on the mainland at points opposite the islands than is consobrina, so that in all probability when that body of land became separated from the mainland it was populated with the form of parvicollis existing at that time, and as this change occurred in comparatively recent times, the species has undergone but sheht change since and is practically identical with that which is now found along the coast on the mainland. I have collected for several years about San Francisco Bay and have not taken a single specimen of consobrina, while I have found parvicollis abundant. Consobrina is surely more abundant south than north of San Francisco. In consobrina the gene are not produced as in blanchardii, and for remarks on scabripennis see p. 338. In fuchsi the prothorax is com- paratively smaller and less convex. In some specimens (females from Los Angeles County) the elytre are densely and distinctly tuberculate throughout. General observations.—The mentum is triangulo-parabolic to para- bolic in outline; surface convex at middle and more or less foveate -along the sides, rather densely but not distinctly punctate and not noticeably setose. The prosternum is not strongly prominent ventrally with the coxee, coarsely sculptured, usually distinctly grooved along the mid- dle, horizontal; rarely feebly arcuate antero-posteriorly between the cox, strongly and horizontally mucronate behind; often vertically truncate posteriorly with the angle more or less well developed as a mucro, which may be deflexed or oblique. - The mesosternum is arcuately and obliquely declivous, deeply and rather broadly concave, so that it appears somewhat bilobed. The metasternum laterally between the coxe is as long as the width of a mesotibia at middle or base. The abdominal process is subquadrate (male) to moderately trans- verse (female) and a third (male) to a sixth (female) wider than the metasternal salient. In the male the post-coxal part of the first abdominal segment is equal in length to the process, and also to that of the second; the latter is twice as long as the fourth; the third is a half longer than the fourth. 59780—Bull. 68—09 22 338 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. In the female the second segment is equal in length to the process; the third is equal to that of the post-coxal part of the first and about a third longer than the fourth. The tarsi are moderate in length and more or less stout. Tarsal formula: Pro. Meso. Meta. Metatibia. Male —27 es 4 6 Female.—2 24 33 54 ELEODES SCABRIPENNIS LeConte. Eleodes scabripennis LECONTE, Proc. Acad. Nat. Sci. Phila., 1859, p. TT, female.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 315. The original description is as follows: | “Atra, ovata, subnitida, thorace latitudine paulo breviore, subquadrato, lateri- bus rotundatis, postice sensim paulo angustato, angulis posticis obtusis dense punctato, ad latera subasperato, elytris ovalibus, thorace latioribus, dorso parum convexis, postice valde declivibus, granulis parvis inordinatis dense exasperatis, et versus suturam punctatis, tibiis muticis, prosterno postice oblique submu- cronato, antennis extrorsum parum incrassatis. Long. .65.” Habitat—California (Fort Tejon). Diagnostic characters —My. Blanchard writes me that the type is a robust female, with the following remarks upon the same: Stout, convex. Thorax convex, wider than long, sides very briefly sinuate at hind angles, coarsely, densely punctate, more closely and asperately on sides, base margined; flanks opaque, rather closely and distinctly granulate. Elytra convex, sides strongly rounded, coarsely granu- late; epipleurzee broad, sparsely granulate, antenne rather stout, joints 4-7 subelongate, Sth scarcely longer than wide. LeConte compares it with veseyi, saying that it has the same sculpturing and size, but differing by the thorax being much less rounded on the sides, by the posterior angles not being at all promi- nent. I have not seen a single specimen in the material at hand which I could refer to this species, and personally I have no idea of its habitus. Fortunately I can rely upon the observations of such careful workers as Mr. Blanchard and Professor Fall. Mr. Blanchard has compared all likely specimens with the type at Cambridge and has failed to refer any examples to the present species. For remarks upon dnculta see p. 332. Professor Fall compared a series with the LeConte type and labeled a male example as scabripennis. It was collected at Santa Barbara, California. This specimen has the form of a male conso- brina, from which it differs no more than the males of that species differ among themselves. The elytral sculpturing is more like sharp- ened granules than tubercles, finer and dense, with punctures simple rach side of the suture; the pronotal punctuation is denser than in cece tare me Ge a Rt ORR fee Rein — > REVISION OF ELEODIINI—BLAISDELL. 3909 consobrina. The specimen is distinctly consobrina-like in general, and not at all like znculta. If this specimen is to be considered as an exponent of scabripennis, I would have no hesitation in pronouncing it as a synonym of con- sobrina. One fact is evident, that no student besides LeConte ever had any true conception of this species. Doctor Horn did not, for he referred the Santa Barbara specimens (énculta 7) to this species. I know of only one authentic specimen, and that is the type. A specimen in Doctor Van Dyke’s collection was doubtfully re- ferred to scabripennis; 1t was collected at Indio, California. It is very opaque and sordid in appearance; the sculpturing is rather coarse and dense, but upon examination I found the gene produced, and it is therefore blanchardii, although rather strongly sculptured and larger (16.5 mm.—8 mm.) than any example of the series studied. It makes a good companion for Casey’s specimen referred to under blanchardii. Until more collecting has been done about Fort Tejon and Santa Barbara, I recommend that scabripennis be retained on our lists. LeConte writes that the prosternum is a little more prominent behind than in vesey?. ELEODES BLANCHARDII, new species. Oblong-ovate, somewhat robust at times, scarcely twice as long as wide, elytra coarsely punctate and subtuberculate laterally ; gene dis- tinctly produced; surface somewhat dull. Head scarcely twice as wide as long, feebly convex, more or less feebly and broadly impressed laterally, sometimes feebly so along the frontal suture, the latter more or less evident as a smooth line; rather thickly punctate, punctures rather coarse and somewhat granulate, denser laterally and on the epistoma. Antennw moderate in length and stoutness, feebly compressed and slightly dilated in the outer four joints, third joint about equal in length to the next two combined, fourth just the least longer than the fifth, the latter to the eighth inclusive subequal in length and slightly longer than wide, the eighth sometimes subtriangular, ninth and tenth orbicular to slightly transverse in outline, eleventh truncate-ovate and about as long as wide. Pronotum somewhat transversely suboval, about a half wider than long, widest at the middle; disc moderately evenly convex, rather densely and more or less evenly punctate, punctures moderate in size, becoming granulate at the sides; apex subtruncate in circular are, obsoletely beaded ; sides evenly and rather strongly rounded, more or less constricted or sinuate at basal ninth, thence straight or oblique 340 BULLETIN 63, UNITED STATES. NATIONAL MUSEUM. to angles, marginal bead more or less reflexed and somewhat entire; base feebly rounded and feebly beaded, scarcely a seventh wider than the apex and equal to the length; apical angles obtuse, sometimes distinct and at others feebly rounded; basal angles subrectangular and not at all prominent. Propleurew more or less coarsely and rather sparsely muricato- granulate, more or less rugulose on the acetabular convexities. Elytra oval to somewhat oblong-oval, about a third longer than wide and widest at the middle; dase subtruncate, wider than the con- tiguous prothoracic base; humeri more or less obtusely rounded ; s¢des evenly arcuate, apex not broadly rounded; disc more or less mod- erately convex on the dorsum, rather strongly, evenly, and not very broadly rounded at the sides, arcuately declivous posteriorly ; surface coarsely punctate, punctures irregularly placed, with a tendency to coalesce in twos and threes, rather simple and with a feeble corroded appearance on either side of the suture, becoming muricato-tubercu- late laterally and on the apex. Epipleure moderate in width, gradually narrowing from base to apex, superior margin rather strong, scarcely sinuate beneath the humeri, where it is sometimes slightly visible at the angle from above; surface usually impunctate and smooth. Sterna densely and rather scabrously sculptured. Parapleure coarsely and rather densely punctate. Abdomen more or less shining, quite coarsely and densely rugoso- punctate, especially about the coxe; last two segments more finely, less densely punctate, and not rugose. Legs moderate in length and stoutness; anterior femora mutic; anterior tibial spurs rather small and similar in the sexes, the an- terior slightly longer and just noticeably stouter than the posterior ; anterior and middle tarsi dissimilar in the sexes. Male-—Oblong-ovate, somewhat narrow. Antenne attaining the prothoracie base. Elytra arcuately and somewhat obliquely decli- vous behind. Abdomen slightly oblique, moderately convex, broadly and rather strongly impressed at middle of the first two segments. Anterior tarsi with the first two joints not noticeably thickened at tip beneath, each with a small tuft of yellowish pubescence, tufts subacute ; first joint of the middle tarsi bearing a small tuft. Female.—Ovate, somewhat robust. Antenne not attaining the prothoracic base. Elytra arcuately and almost vertically declivous posteriorly. Abdomen horizontal and rather strongly convex. An- terior tarsi with the first joint slightly thickened at tip beneath, groove scarcely interrupted by the apico-marginal spinules. Males: Length, 15 mm.; width,6.3 mm. Females: Length, 14.5-17 mm.; width, 7.3-7.5 mm. Measurements. REVISION OF ELEODIINI—BLAISDELL. 341 Genital characters, male—Edeagophore elongately oblong-ovate and slightly arched. Basale elongate oblong, shghtly narrowing apically ; surface evenly convex and the sides feebly arcuate. Apicale triangular; surface moderately convex, without groove ; sides quite strongly sinuate in apical two-thirds, apex appearing attenuate and acute; base obtusely lobed at the middle and sinuate laterally. Sternite transversely parabolic in outline. Each lobe subtriangu- lar; surface feebly convex and glabrous, punctate and setose in apical three-fourths, punctures rather dense, especially about apex, sete moderate in length, longer about apical margin; external margin more or less arcuate to apex, the latter introrse and rather narrowly rounded ; internal margin quite straight. Membrane not setose across the bottom of the sinus, the latter rather deep and triangular. Female.—Genital segment triangular, obtuse at apex and moder- ately depressed; surface nearly plane, well chitinized, and sparsely setose. Valvula (Plate 6, fig. 1). Dorsal plate oblong, four times as long as wide; surface glabrous and shining, shghtly concave, very sparsely punctate and setose in apical moiety, sete fine; sides parallel and more or less feebly arcuate or sinuous; apical margin evenly rounded. Apex short and more or less chitinized, finely and rather densely setose, sete slightly longer at tip. Appendage short mammilliform with pencil of moderately long sete at tip; fossa in the external surface of apex and not covered by apical margin of the dorsal plate, sete just noticeably longer about its margins. Superior pudendal membrane longitudinally rugulose and attain- ing the apical fourth of the dorsal plate. Basal prominences scarcely evident. Ventrolateral surfaces flattened centrally to base, triangular, sur- face lines straight, viewed longitudinally; laterally moderately rounded. Submarginal groove well developed beneath the slightly explanate external border of the dorsal plate, and attaining the fossa, surface scarcely at all concave before the apex. Internal margins of the valves contiguous in basal two-thirds; genital fissure narrow in apical third, inferior pudendal membrane not visible. Habitat—California (San Diego County). I have taken this species upon the Bolean Mountain, which borders Warner’s ranch. It probably occurs throughout the Cuyamaca Range, also at Poway (elevation, 700 feet). Examples are before me from the Blanchard collection, and others received from D. W. Coquillett a number of years ago, all, however, from the above-named county. 342 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Number of specitnens studied, 9. Types in my own and Mr. Blanchard’s collections. Type-locality—San Diego County, California. Salient type-characters—The buccal processes of the gene are pro- duced and anteriorly prominent. Pronotum moderately convex; apex subtruncate in circular arc; sides rather strongly rounded, con- stricted at basal ninth, thence straight to the basal angles, the latter subrectangular; apical angles obtuse and distinct. Elytra about a third longer than wide; humeri obtuse and rounded, epipleural margin just visible from above; disc coarsely punctate, punctures somewhat eroded, noticeably coalescing in twos and threes, simple about the suture, muricato-tuberculate laterally and about apex; the most peripheral are distinct and shining tubercles. Diagnostic characters—Smaller and similarly sculptured to con- sobrina, to which it is closely related, and easily separated by the unusually developed buccal process of the gene that are anteriorly produced and somewhat acute. An example before me, and kindly given me for study by Colonel Casey, apparently belongs here. It has the thorax very coarsely and densely punctate. There is also a tendency for the punctures to coalesce. In fact, it is more strongly sculptured throughout than any other specimen of blanchardii at hand. A female taken on the Bolean Mountain approaches it and connects it with the typical speci- mens. Casey’s specimen has the buccal processes very shghtly promi- nent and not narrowed. It is without doubt heterotypical of the present species; in form it agrees with a male in Mr. Blanchard’s collection. It should be noted that the elytra are shorter than in consobrina. I take great pleasure in remembering the kind and painstaking aid accorded me by Mr. Frederick Blanchard. General observations —The mentum is comparatively large, but variable in size, slightly transverse and parabolic in outline, some- times less strongly rounded at the sides; surface rather broadly convex at the center and narrowly subfoveate within the lateral mar- gins; not noticeably setose. Pro- and mesosternum as in consobrina. The metasternum laterally between the coxe is as long as the width of a mesotibia at the middle. The abdominal process is subquadrate, and a fifth (female) to a third (male) of its width broader than the metasternal salient, also equal to the post-coxal part of the same segment, equal to the length of the third (female). In the male the second segment is twice as long as the fourth, the third being a third of its own length longer than the fourth. In the female the second is twice as long as the fourth. The tarsi are of moderate length and not as stout as in consobrina. = She ee eee een ie le i ie Oe” 4 O'R’: REVISION OF ELEODIINI—BLAISDELL. 343 Tarsal formula: Pro. Meso. Meta. Metatibia. Male.—2 25 yakt a : FELD Female.—2 24 23 5 ELEODES FUCHSII, new species. Ovate, subopaque, prothorax comparatively small. Elytra sub- asperately granulate laterally and on apex. //ead about twice as wide as long, more or less feebly convex, impressed laterally and usually along the frontal suture, the latter more or less bisinuate and fine, sides not very prominent, more or less evenly punctate, punc- tures moderately fine and not very densely placed. Antenne rather long, feebly compressed and slightly widened in the outer four joints, third joint scarcely as long as the next two combined, the fourth to the eighth inclusive subequal in length, the eighth slightly widened, ninth as wide as long and subtriangular, tenth orbicular or trian- gulo-orbicular, eleventh ovate. Pronotum scarcely to moderately transverse, a fourth to a half wider than long, widest at about the middle; disc moderately and evenly convex, rather densely and somewhat finely punctate, becoming more or less granulate along the sides; apea slightly and evenly emarginate or truncate in circular arc, obsoletely margined; sides rather strongly arcuate, less so behind the middle, where they appear somewhat oblique to the rather strong constriction at basal seventh, which may not be very suddenly or at other times rather gradually formed, thence to the angles more or less straight and parallel; mar- ginal bead rather fine and somewhat obsolete at basal constriction ; base shghtly rounded, finely to obsoletely margined, very shghtly wider than the apex and about equal to the length; apical angles dis- tinct and subacute, or slightly obtuse; basal angles subrectangular or obtuse and not in the least rounded. Propleurw smooth and subopaque, sparsely muricato-granulate and more or less rugulose on the acetabular convexities. Elytra oval, about a third longer than wide, widest at the middle; base equal to or distinctly wider than the contiguous prothoracic base ; humeri evenly or obtusely rounded; s¢des broadly and moderately arcuate, moderately attenuate at apex, the latter not broadly rounded ; disc moderately convex, not depressed, quite evenly and not very broadly rounded laterally, arcuately declivous posteriorly; surface with the punctures rather coarse, densely and irregularly placed, at times vaguely seriate when viewed longitudinally, punctures simple on each side of the suture, asperately granulate laterally and on the apex, minutely setigerous. Epipleurw moderate in width, gradually narrowing from base to apex, superior margin straight or very feebly sinuate beneath the 344 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. humeri, where it is at times somewhat visible from above; surface smooth, opaque, sparsely and quite obsoletely punctate. Sterna irregularly and densely punctato-rugulose. Parapleure rather coarsely but not densely punctate. Abdomen more or less glabrous, variable in dullness, rather sparsely and somewhat finely punctate, more coarsely, densely, and rugulosely sculptured about the coxw, last two segments more finely so. Legs moderate in length and stoutness; anterior femora mutic and more or less sinuate in outer fifth; tibial spurs similar in the sexes and rather small, the anterior shehtly longer than the posterior; tarsi dissimilar.in the sexes, Male.—Moderately narrow. Antenne reaching a little beyond the base of the prothorax. Elytra a little narrowed posteriorly, arcuately and rather obliquely declivous apically. Abdomen shghtly oblique and moderately convex, impressed on the first two segments at mid- dle. First two joints of the protarsi shghtly thickened at tip beneath with pubescent tufts upon the same; the tufts are truncate, that of the first moderate in size and about twice as large as that of the second. First jomt of the middle tarsi with a very small tuft at tip; all the tufts golden yellow. Female.—Robust. Antenne just slightly reaching beyond the base of the prothorax. Elytra arcuately but scarcely vertically declivous posteriorly. Abdomen horizontal and rather strongly convex. An- terior and middle tarsi simple beneath, grooves distinct. Measurements.—Males: Length, 13-16 mm.; width 6-7 mm. /e- males: Length 14.5-17 mm.; width 7-8.2 mm. Genital characters, male—KKdeagophore somewhat flattened ob- long-ovate, moderately elongate and slightly arched. Basale oblong, sides feebly arcuate, surface moderately convex. Apicale triangular, rather strongly convex and without a groove; sides rather strongly sinuate so that the apex appears produced, the latter acute; base with a small rounded median lobe and feebly sin- uate laterally. Sternite small, shghtly transversely oblong. Each lobe small and subtriangular; surface almost impunctate in basal third, thence to apex sparsely to densely punctate at apical margin, setose, sete small, increasing in length to apex where they are moderately long; sides sinuate and somewhat oblique in basal two-thirds, thence to apex truncately subarcuate; apex introrse and subacute; internal margin straight or feebly sinuate. Membrane not setose, sinus triangulo- oblong. Female-—Genital segment trapezo-triangular, dorsal surface very shghtly ogival, very feebly setose and moderately depressed. Valvula.—Dorsal plate slightly oblique, being externally declivous, elongately oblong, sides subparallel and moderately explanate exter- ol REVISION OF ELEODIINI—BLAISDELL. 845 nally; surface scarcely concave, very sparsely punctate and setose, setae very small; external and internal borders straight; apical mar- gin oblique externally and arcuate internally, angle broadly rounded, apex small and short, triangular and finely setose, without longer setee at tip; fossa at middle of the external surface. Appendage small and short mammilliform, with a pencil of moder- ately long sete at tip. Superior pudendal membrane more or less longitudinally rugulose and attaining the apical fourth of the dorsal plate. Basal prominences not evident. Ventrolateral surfaces flattened centrally to base, rounded laterally ; surface smooth and not noticeably setose; sides feebly concave later- ally before the apices, and the submarginal groove broad beneath the explanate external border of the dorsal plate. Internal mar- gins of the valves contiguous in basal two-thirds; genital fissure in apical third, moderately wide and fusiform. Inferior pudendal mem- brane not visible. Habitat.—California (Tulare County, at Colony Mills, elevation 5,415 feet, Giant Forest; Marble Fork, Kaweah River, elevation 5,200 feet, Kaweah to Progress Springs, elevation, 1,000-1,740 feet. Col- lected by Charles Fuchs and Ralph Hopping). Number of specimens studied, 90. Sexitypes in my own collection; co-types in the collections of Van Dyke, Fuchs, Hopping, and my own. Type-locality—Colony Mills, Tulare County, California. Salient type-characters.—Pronotum comparatively small, appar- ently subcordate, the sides rather strongly arcuate anteriorly, behind the middle less arcuate and very slightly converging, constricted in basal seventh, thence quite straight to the basal angles, which are subrectangular; disc rather finely and densely punctate, becoming granular laterally; apical angles distinct and subacute; apex feebly and evenly emarginate. Elytra about a third longer than wide, in the female broadly oval and apparently inflated in contrast to the small prothorax; dise mod- erately convex on the dorsum, punctures rather coarse, densely and irregularly placed, simple about the suture and exasperately granu- late laterally and about the apex. Diagnostic characters.—This species is variable in form, typically the females are subcychriform; the heterotypes resembling conso- brina on the one hand and parvicollis on the other. In typical specimens the prothorax is comparatively small and apparently subcordate, the elytra broadly oval in the female, giving the effect of inflation, when there is none the dorsum is moderately convex; the humeri are broadly rounded in both sexes. Such examples resemble consobrina. 346 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. In other specimens the humeri are obtusely rounded and the su- perior epipleural margin is often visible from above, the thorax wider and more evenly rounded at the sides, and the general form resembles parvicollis. ‘The integuments are more or less opaque. Fuchsii appears to be intermediate between consobrina and parvi- collis. 1 at first associated. it with the former and then with the lat- ter as a race, but could not satisfy myself as to which species it 1s most closely related, and therefore determined on the intermediate course and give it specific standing. It can be recognized from consobrina by the more finely and much less densely sculptured abdomen and more cordate prothorax; from parvicollis by the narrower prothorax and the more consobrina-like habitus. From neotome it is separated by the much less convex form and less strongly sculptured abdomen. Producta has a much more transverse prothorax, the humeri strongly prominent with the superior epipleural margin very dis- tinctly visible from above, the elytra very evidently flattened, form oblong-oval and broad at base. The males are more difficult to sepa- rate, the sides of the pronotal disc are usually distinctly flattened and granulate; in fuchsii the pronotal disc is not usually depressed at the sides and quite strongly and evenly convex. Hornii is more depressed, with very oval elytra, the humeri being obsolete. General observations—The mentum is usually parabolic in out- line, although at times somewhat triangular; the surface is rather narrowly convex at middle from apex to base and more or less broadly foveate laterally, finely scabrous, punctures not distinctly defined and not noticeably setigerous. The prosternum is quite gradually prominent from the anterior margin so that the intercoxal process is not strongly nor suddenly protuberant ventrally; at times it is evenly arcuate antero-posteriorly or feebly arcuate between the coxee and subtruncate behind, with the mucro more or less produced or small and more or less deflexed. Mesosternum more or less vertically arcuate and with a varying concavity. The metasternum laterally between the coxz is as long as the width of a mesotibia at middle. The abdominal process is more or less quadrate and a third (male) to a fourth (female) of its width broader than the metasternal salient, as long as the second segment in the mate and to the post- coxal part of the first segment, as well as the third, in the female. Tn the male the post-coxal part of the first segment is equal to the leneth of the third, the latter about a fourth longer than the fourth. In the female the second is twice as long as the fourth. REVISION OF ELEODIINI—BLAISDELL. 847 The tarsi are moderate in length and not very short; there is eyi- dent variation in the individuals of the series before me. Tarsal formula: Pro: Meso. Meta. Metatibia, Male.—23 33 34 6 Female.— 2 22 35 cig! ELEODES NEOTOMA, new species. Ovate, moderately robust, shining, and smooth, strongly convex; Elytra scarcely asperately muricately punctate. Head twice as wide as long, feebly convex, more or less impressed laterally and along the frontal suture, the latter usually evident as a glabrous line, quite evenly and not very densely punctate, punctures moderate, sides scarcely at all prominent. } i“ we tm REVISION OF ELEODITNI—BLAISDELL. 879 first segment is slightly shorter than the process; the second segment is about twice as long as the fourth. In the male the third segment is about equal in length to that of the process and about a third of its own length shorter than the second. In the female the second segment is about equal in length to the process, and the third is about a fourth of its length shorter than the second. The protarsi are about three-eighths (male) or a third (female) of their length shorter than a mesotarsus. The mesotarsi are about four-elevenths (male) or a fourth of' their length shorter than a metatarsus. A metatarsus is about a third (male) or three-fifths of its length shorter than its metatibia. ELEODES CORDATA Eschscholtz. Eleodes cordata EscuscnoutTz, Zool. Atlas, III, 18383, p. 12—MANNERHEIM, Bull. Soe. Nat. Moscow, XVI, 18438, p. 272.—LEeConrr, Proc. Acad. Nat. Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, Durolive Hleodes tuberculata EscuscHoutTz, Zool. Atlas, III, 1833, p. 12.—MANNER- HEIM, Bull. Soc. Nat. Moscow, XVI, 1848, p. 274. Fleodes intricata MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1843, p. 273. Eleodes stricta LeContTrE, Reports of Explor. and Surveys, 47th Parallel, XII, Appendix No. 1, 1857, p. 50. var. rotundipennis, LECoNtTE, Reports of Explor. and Surveys, 47th Parallel, XII, Appendix No. 1, 1857, p. 50. Moderately robust, subovate, more or less shining, about twice as long as wide, prothorax strongly constricted at base, pronotum con- fluently and very coarsely punctate; elytra rugosely muricato-tuber- culate. Head about twice as wide as long, feebly convex, lateral impres- sions not strongly marked, frontal suture not evident, very coarsely and confluently punctate. Antennw moderate and somewhat stout, outer four joints very feebly compressed and very slightly widened, third joint equal in length to the next two combined, fourth scarcely longer than the fifth, the latter to the eighth inclusive subequal, eighth subtriangular, ninth transversely oval and frequently slightly robust, tenth transversely oval, eleventh short ovate and usually slightly smaller than either of the two preceding joints. Pronotum subcordate, about a half wider than long, widest at about the middle; disc moderately convex, more or less declivous at apical and basal angles, coarsely, densely, and more or less confluently punctate, interstices more or less convex; apex feebly emarginate to truncate, obsoletely margined; sides more strongly rounded in front, narrowing rather suddenly behind the middle, causing them to appear somewhat angulate, frequently more evenly arcuate to the 380 BULLETIN 638, UNITED STATES NATIONAL MUSEUM. constriction, which is usually at basal sixth, thence straight and parallel to the basal angles; the fine margin is very feebly reflexed and obsolete on the constriction; base truncate in circular are, finely or obsoletely margined, about equal to the length and slightly to a fourth wider than the apex; apical angles obtuse, not in the least prominent anteriorly and more or less rounded; basal angles rec- tangular and not prominent. Propleurw more or less sparsely granulate and rugulose. Elytra rather broadly oval, about a fourth longer than wide, widest at the middle; base truncate and wider than the contiguous pro- thoracic base; humeri rounded, scarcely at all angulate; stdes evenly arcuate, apex obtuse and moderately rounded; disc moderately convex on the dorsum, rather strongly and evenly rounded laterally, strongly and arcuately declivous posteriorly, generally quite vertically so; surface densely, irregularly, and rugosely muricato-tuberculate, less strongly sculptured along the suture; punctures minutely setigerous. Epipleure moderate in width, shghtly dilated beneath the humeri, superior margin feebly and very broadly sinuate in basal fourth and scarcely visible at humeri when viewed from above; surface smooth, sparsely and rather coarsely punctate. Sterna more or less densely punctato-rugose, punctures setigerous, noticeably so on the mesosternum; sete short, black; and rather stiff. Parapleure densely punctate. Abdomen smooth, sparsely and distinctly punctate; first segment punctato-rugose, the fifth rather densely punctured. Legs moderate in length and thickness and quite strongly sculp- tured, the punctures bearing short black sete. Anterior femora mutic in the sexes. Anterior tibial spurs quite similar, the anterior shghtly longer than the posterior. Anterior and middle tarsi dis- similar in the sexes. Male—Moderately robust, somewhat narrow, antenne reaching to or just beyond the base of the prothorax. Elytra about a fourth longer than wide. Abdomen moderately convex, slightly oblique, broadly impressed at middle of the first segment. Spurs of the pro- tibize quite slender and acute. First two joints of the protarsi clothed with flattened tufts of golden pubescence beneath; first joint of the middle tarsi with a well-developed and similar tuft at apical third beneath; pubescent joints not noticeably thickened at tip. Female.—Robust. Antenne about reaching to the prothoracic base. Elytra broadly oval and about a fifth longer than wide. Abdomen quite strongly convex. Spurs of the protibie slightly thickened and acute. First joint of the protarsi slightly thickened at apex beneath, groove more or less distinct. Measurements.—Males: Wength, 13-15.2 mm.; width, 6-6.8 mm. Females: Length, 12-17 mm.; width, 6.8-9 mm. ws pi a 1m Cx at ade ne aa “allel ended REVISION OF ELEODIINI—BLAISDELL. 381 Three forms may be recognized according to the character of the seulpturing: Forma sublevis.—Similar in form to the typical form, but smoother, with the sculpturing somewhat obsolete. Forma typica.—Strongly and roughly sculptured; elytra rugosely punctate, the interstices between the punctures elevated and convex, and more or less confluent transversely. Forma intermedia.—Sculpturing more strongly muricato-tubercu- late, with the intervals between the punctures less convex and conse- quently less rugose, approaching pimelioides. Genital characters, male—Kdeagophore stoutly fusiform, feebly arched. | Basale oblong, about twice as long as wide; surface evenly convex; sides feebly arcuate, and the apex quite deeply and triangularly emarginate. Apicale triangular, rather narrow, usually distinctly narrower at base than the basale at apex; surface strongly convex, without groove; sides nearly straight or feebly arcuate, slightly prominent at base; apex acute; base acutely lobed at middle, laterally feebly sinuate. Sternite subparabolic. Each lobe triangular; external border more or less straight or feebly and broadly sinuate in basal two- thirds, thence quite evenly arcuate to apical margin, angle nar- rowly rounded; internal margin feebly sinuate; surface very feebly convex, punctate and setose in apical half, setee moderate in length, not dense, longer toward and on apical margin. Membrane not setose across the bottom of the sinus, the latter more or less triangular and apparently not closed by the membrane. Female——Genital segment triangulo-trapezoidal, surface of the valves slightly declivous, setose. Valvula (Plate 4, fig. 6).—Dorsal plate oblong, feebly narrowed from base to apex. Surface plane or feebly concave, smooth, punc- tures scattered, each with a rather long flying seta; borders quite straight; apical margin oblique, angle obtuse and feebly rounded ; internally not distinctly defined from the apex, the latter rather short and membranous, rounded at tip and clothed with moderately long setae. Appendage mammilliform, with a pencil of quite long hairs at tip. Fossa rather large at base of the external apical surface. Basal prominences not evident. Superior pudendal membrane longitudinally rugulose, and attain- ing the middle of the dorsal plate. Ventrolateral surfaces.—Body triangular, surface lines straight when viewed longitudinally, quite evenly convex laterally, sparsely setose in apical half. sete moderate and more or less flying. Sub- 382 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. marginal groove well developed beneath the shghtly expanded ex- ternal border of the dorsal plate and terminating at the fossa beneath the angle. Internal margins of the valves contiguous in basal two- thirds; fissure narrowly fusiform in apical third, and the inferior pudendal membrane is not visible. Habitat.—California. Forma sublaevis, about San Francisco Bay; Forma typica, central California, along the coast; San Joaquin and Sacramento valleys; Calaveras County; Tulare County, Ralph Hop- ping; Los Angeles County, Doctor Van Dyke; Forma intermedia, northern California, southern Oregon, western slopes of the Sierra Nevada Mountains. Professor Wickham in his “ Coleoptera of Colorado” says that Professor Snow has reported one specimen from Colorado. It is in all probability a variation of pémelioides. Number of specimens studied, 500. Type in the Eschscholtz collection ? T ype-locality.—* California.” Salient type-characters.—Ovate, robust, shining. Head and pro- notum coarsely and confluently punctate; thorax subcordate, apex feebly emarginate, angles obtuse; sides more strongly rounded in front, narrowing rather suddenly behind the middle, causing them to appear somewhat angulate; strongly constricted at base; angles rectangular. Elytra oval; humeri much rounded and obtuse; disc arcuately and vertically declivous posteriorly; surface densely and rugosely muricato-tuberculate, less strongly so along the suture. Diagnostic characters.—Cordata ditters from all other species in the subgenus Blapylis, except pimelioides, in having the sides of the prothorax more or less subangulate. This character is especially marked in the males. The prothorax is more evenly rounded behind the middle in most females, but always strongly constricted at base, giving it with the above characters a subcordate appearance. The marginal bead does not reach the base. The coarse sculpturing will cause it to be frequently confused with scabrosa, but the setose propleure of the latter will readily separate the two, besides the pronotal sculpturing is quite different in the two species. Clavicornis is much less strongly sculptured, and the sides of the pronotal dise are distinctly setose along, with the pro- pleure. In pimelioides and brunnipes the elytra are sculptured with rows of small and rounded tubercles; in the latter species the legs are brown or reddish. In studying a large series of cordata it is very evident that the sculpturing varies toward pimelioides, so that in the heterotypes it is quite difficult to tell where one species leaves off and the other begins; : REVISION OF ELEODIINI—BLAISDELL. 383 cordata acquires a rugoseness of sculpturing and pimelioides a tuber- culate form of elytral markings. They are to be considered as two divergent forms derived from a common ancestral ramus. Specimens from Oregon have the prothorax rounded on the sides, as in parvicollis and scabrosa (see rotundipennis). Specimens from southern California collected by Doctor Van Dyke are remarkable among those of the large series before me on account of their elongate form, measuring: Length, 15 mm.; width, 6.5 mm.; elytra more elongately oval (forma elongata). These specimens approach an unique male from Amador County, in which the form is elongate oblong, thorax as in cordata, antenne long and stout, and the elytra are distinctly tuberculate and not rugose. The whole habitus is that of a stout, oblong cordata (forma oblonga). Tength, 16 mm.; width, 7 mm. These are remarkable variations from the monotonous short, robust, normal form. Mannerheim’s types of éntricata were taken about Fort Ross, Cali- fornia, by Doctor Fischer, and I can not see that it differs in any essential characters to give it even varietal standing. Many females have the * ¢thorace transverso, lateribus valde rotundatis.” Ina large series of cordata a variation in the length and stoutness of the an- tenne and legs, independent of sex, may frequently be observed. The humervi vary considerably also, never prominent, but frequently subangulate and always more or less rounded. Tuberculata Eschscholtz is to be disposed of in the same way as intricata. For sticta, see p. 384. General observations —The mentum is usually more or less trian- gulo-trapezoidal and rather small, finely sculptured, feebly foveate laterally, and very shghtly convex at middle. The prosternum is variable, usually rather prominent ventrally with the coxe, and quite strongly arcuate antero-posteriorly, without a mucro; frequently a small mucro is present at middle of the pos- terior edge; rarely horizontal, subtruncate behind, and more or less feebly mucronate. The mesosternum is vertically arcuate and more or less broadly and deeply concave. The metasternum laterally between the coxe is as long as the width of a mesotibia at middle. The abdominal process is subquadrate, equal in width (female) or a third (male) of its width broader than the metasternal salient. In the male the post-coxal part of the first segment is equal in length to the third; the second to that of the process and twice as long as the fourth and about a fifth longer than the third. In the female the post-coxal part of the first segment is equal in length to that of the process and also to the second, the latter being 384 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. about twice as long as the fourth; the third is about one-half longer than the fourth. The tarsi are usually moderate in length and stoutness. Tarsal formula: Pro. Meso. Meta. Metatibia. Male.—2 23 a 43 Female.—1#4 23 3 4h ELEODES CORDATA var. ROTUNDIPENNIS LeConte. Pronotal sides evenly and broadly rounded (as in parwicollis), basal constriction short and the angles rectangular. Otherwise as in cordata. LeConte was very much in doubt as to the propriety of separating this form even as a race. The constancy of form in the prothorax, which, taken in conjunction with the coarse elytral sculpturing, make it appear at first sight as a northern modification of scabrosa; this is not so; the propleure are without distinct sete, and besides the pronotal punctuation is quite different, being as in cordata; this is in fact the only superficial character which separates and pre- vents it from being naturally associated with the parvicollis section. I consider it a good race, and the series before me shows conclusively that it varies more toward typical cordata than pimelioides. Habitat—Oregon (Koebele); British Columbia (Victoria and North Bend, Hubbard and Schwarz) ; Washington (H. K. Morrison, Easton and Olympia). Number of specimens studied, 11. Type in the LeConte collection. Ty pe-locality — Oregon.” T have taken this variety at Oregon City, near Portland, Oregon. Stricta LeConte, according to the author, has the “ thorax less sud- denly but very much constricted at base, fully one-half wider than long; elytra more coarsely and less densely punctured,” and is to be taken as a synonym of the present race. ELEODES PIMELIOIDES Mannerheim. Eleodes pimelioides MANNERHEIM, Bull. Soc. Nat. Moscow, XVI, 1845, p. 274: Mag. Zool., XIII, 1848, no. 129, fig. — LeContTr, Proc. Acad. Nat. Sci. Phila., 1858, p. 182.—Horn, Trans. Amer. Phil. Soc., XIV, 1870, p. 318. Eleodes viator LEContE, Proc. Acad. Nat. Sci. Phila., 1858, p. 188. Eleodes subligata LeConte, Reports of Explor. and Surveys, 47th Parallel, XII, Appendix no. 1, 1857, p. 50. var. brunnipes Casry, Ann. N. Y. Acad. Sci., V, Nov., 1890, p. 402. . Moderately robust, ovate, feebly shining to opaque, about twice as long as wide; prothorax more or less strongly constricted at base, densely rugoso-punctate; elytra sculptured with small tubercles, which may be rounded or reclinate and more or less piliferous, nT REVISION OF ELEODIINI—BLAISDELL. 385 Head densely punctate, antenne somewhat slender, ninth joint tri- angulo-orbicular to transversely oval, tenth more or less transversely oval. Pronotum subcordate to transversely suboval, widest near the mid- dle, a fourth to scarcely a half wider than long; sides evenly and quite strongly arcuate to basal seventh, or subangulate at middle, rounded in front and quite rapidly converging posteriorly and sin-. uate at basal fourth, thence in each instance quite straight and par- allel to the basal angles; base equal to the length or in some males shorter than the length; apical angles obtuse, frequently not in the least rounded, at other times more or less so. Elytra quite broadly oval to subquadrate, widest at or behind the middle, a fourth to a third longer than wide; disc more or less deplanate on the dorsum, strongly, arcuately, and vertically declivous posteriorly; surface densely tuberculate, tubercles apparently ar- ranged in rows on the dorsum or irregular throughout; each bears a very short, black seta near apex; when arranged in rows there are very small muricate punctures scattered sparsely and irregularly ‘between, always less distinct along the suture centrally; the tubercles are more or less rounded and shining, the interstices between more or less opaque. Otherwise as in cordata. Male.—First two joints of the protarsi with tuft of yellowish pubescence near tip beneath; that of the second joint is rather small; tuft on the first joint of the mesotarsi quite small. Tufts somewhat long and truncate at tips. Otherwise as in cordata. Female.—First joint of the anterior tarsi distinctly thickened at tip beneath. Otherwise as in cordata. Measurements.—Males: Length, 12-14.5 mm.; width, 6-7 mm. Females: Length, 13-15.2 mm.; width, 6.5-8 mm. Genital characters, male—As in cordata. Female.—Genital segment triangular, surface quite plane and shghtly setose. Valvula (Plate 5, fig. 5).—Dorsal plate oblong, feebly or scarcely narrowed apically, slightly explanate externally; apical margin nearly transverse to feebly oblique, inwardly not defined from the surface of the apex, angle obtuse and more or less feebly rounded. Ventrolateral surfaces.—Submarginal groove distinct and well defined beneath the expanded external border of the dorsal plate. Otherwise as in cordata. Habitat—California (Shasta, Siskiyou, Butte, Lake, and» Hum- boldt counties) ; Oregon (The Dalles, Hubbard and Schwarz; Clacka- mas County, Charles Fuchs) ; Utah (Alta, Wasatch in July, Amer- ican Fork Canyon in July, Hubbard and-Schwarz; Park City); Ne- vada (Elko, H. F. Wickham); Washington (Easton, C. V. Riley; 597SO— Bull. 68—09— oF -) 386 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Pullman, Palouse, A. M. Warren; Olympia, E. C. Van Dyke; Walla Walla, Charles Fuchs; Everett, July, Wickham and Van Dyke) ; Idaho (Camas Prairie, C. V. Riley; Cceur d’Alene, June, H. F. Wick- ham); Montana (Helena, August, Hubbard and Schwarz) ; Colo- rado (Central City, Garland, C. V. Riley; Buena Vista, July, Hub- bard and Schwarz; Golden, Glenwood, Horse Fly Peak, Idaho Springs, Aspen, Fort Collins, Leavenworth Valley (9,000-10,000 feet), Roaring Fork, Monument Gulch to Montrose, Little Willow Creek, Canyon of Big Blue, Montrose and adjacent mountains, H. F. Wickham in the “ Coleoptera of Colorado”); Wyoming (Natural Park, July) ; British Columbia (Vancouver Island, collection of Dr. E. C. Van Dyke). Number of specimens studied, 90. Type in the Mannerheim collection. Type-locality.—* California.” Salient type-character—Opaque. Thorax with the sides rounded, densely rugoso-punctate. Elytra subquadrate, apex strongly decli- vous, dorsum deplanate, densely granulate, granules reclinate, rough and piliferous (Mannerheim). Diaqnostic characters.—In general form like cordata, from which it differs in having the elytra sculptured with small rounded tuber- cles. The form of the tubercles differs, in some specimens distinctly rounded, in others reclinate and often more or less muricate. In some males there is evidence of rows. The series from Walla Walla, Washington, have the elytra depla- nate and subquadrate; these characters are less marked in the other specimens. In the race brunnipes the legs are brownish and more distinctly piliferous, at least in those specimens from Nevada. Viator, described by LeConte from Fort Bridger, is a male, with the elytral tubercles large and extending almost to the suture. Colonel Casey also considers that this form does not differ from ordinary males of pimelioides. Subligata is one of the forms associated with cordata by LeConte; he obtained specimens from Oregon. Doctor Horn considered it a synonym of the present species, and which is undoubtedly correct. For remarks upon variation in sculpturing see cordata. The legs are variable, usually more slender in the males and stouter in the females. The thorax and humeri vary as in cordata. ELEODES PIMELIOIDES var. BRUNNIPES Casey. Rather robust, moderately convex, coarsely, densely sculptured and dull, black; legs dark brown throughout. Head large, fully two-thirds as wide as the prothorax, coarsely, extremely densely punctate and scabrous. Antenne longer than the : SN ee eee. REVISION OF ELEODIINI—BLAISDELL. 387 head and prothorax, rather slender, third joint about four times as long as wide. Prothorax scarcely a third wider than long; disc moderately, evenly convex throughout, very coarsely, deeply, and confluently punctate; apea subtruncate; sides strongly angulate at the middle, thence very feebly arcuate to the apex and broadly sinuate to the base; base subtruncate and nearly equal to the apex in width; apical angles obtuse, not distinctly rounded, not at all prominent; basal angles right and not rounded. Elytra at base nearly a third wider than the contiguous protho- racic base, very slightly more than twice as long as the latter, ab- ruptly and obtusely rounded behind when viewed vertically; sédes strongly arcuate behind, gradually convergent and straighter thence to the humeri, which are slightly obtuse but scarcely at all rounded; disc widest behind the middle, where it is from a third to two-fifths wider than the prothorax, feebly convex on the dorsum, strongly so laterally; surface coarsely, densely, asperately punctate, the asperi- ties arranged without trace of order. Legs moderate in length, somewhat slender. Male and female secondary sexual characters as in pimelioides. Measurements.—Length, 9.2-9.5 mm.; width, 4.3-4.6 mm. ip REVISION OF ELEODIINI—BLAISDELL. 455 Elytra elongate oval, fully twice as long as wide; base subtruneate, feebly sinuate laterally; humeral angles more or less distinct ; margin acute, not foliaceous nor suddenly formed, evenly and gradually reflexed and attaining the apew, sides feebly arcuate, gradually nar- rowing to apex and attenuately converging, apex acute and not pro- duced ; disc more or less concave, suture frequently raised posteriorly : here the dise is more or less biconcave, gradually, evenly arcuately declivous behind, acutely inflexed laterally, the inflexed sides moder- ately oblique and plane; surface sculptured with fine, irregularly, sparsely, and evenly placed, subasperate punctures, the inflexed sides similarly sculptured; each puncture bears a very minute decurved seta. Epipleure narrow, scarcely at all differentiated from the inflexed sides of the elytra and similarly punctured; distinctly defined at apex and feebly, minutely dilated beneath the elytral apices. Sterna opaque and finely, irregularly sculptured. Parapleure opaque and obsoletely punctate. Abdomen opaque, finely, rather evenly and not densely punctulate ; first two segments horizontal, third shghtly oblique, last two much less convex. Legs long and rather slender; posterior femora usually as long as the first four abdominal segments; profemora mutic; tibial spurs quite similar and the protarsi simple. Male—Elongate and rather narrow. Abdomen quite strongly convex, median surface of the first three segments more or less broadly and feebly impressed. Anterior tibiae suddenly and_ briefly con- stricted at base. Measurements.—Males: Length, 15-20 mm.; width, 6-8 mm. Genital characters, male —Edeagophore elongately subfusiform, six to eight times longer than wide, more or less arched. Basale elongate, about five times longer than wide; sides sub- parallel to slightly arcuate; surface transversely and moderately convex, distinctly membranous centrally at the distal extremity. Apicale rather slender, elongately triangular; dorsal surface moder- ately convex, with a median linear groove at middle two-thirds, very sparsely and finely punctulate: sides briefly arcuate at base and middle, with intervening edge and side of apex feebly sinuate; apex subacute and shghtly produced; base distinctly bisinuate with a median subacute lobe. Sternite subparabolic in outline, circularly concave at middle like a watch-glass between the bases of the lobes. Each lobe triangular and distinctly longer than wide; external border evenly arcuate, apex subobtusely rounded, and the internal border straight or feebly sinuate; surface more or less convex, concave internally to sinus, basal third smooth and almost impunctate, apical two-thirds densely 456 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. punctured and clothed with quite long sete. Sinus triangular, two- thirds closed by membrane, the latter setose. Habitat.—Nevada (Horn; Hubbard and Schwarz); California (Lassen County, Charles Fuchs). Number of specimens studied, 2 (males). Type in the Horn collection; collected by Mr. William M. Gabb. Type-locality—Western Nevada, Salient type-characters——The most elongate of our species. Pro- thoracic margin acute, moderately explanate and reflexed. Elytr: with margin simply acute and extending to apex. Epipleure indis- tinctly defined except at apex. Diagnostic characters —The elongate form, with elytra gradually and arcuately narrowing to apex give a characteristic facies. Separated from contuswm by its narrower form, besides the body has a greater dorso-ventral thickness, and the males are not caudate. In planuwm the elytral margins do not reach the apex, in clongatum they do. From depressum by its elongate oval elytra, obsoletely defined epipleuree and by having the basal joints of the antenne less pubescent. General observations—The mentum is rather small, middle lobe variable, sides converging anteriorly, straight or more or less arcuate, with apex rounded or subacute; surface very slightly convex and more or less feebly foveate laterally. The prosternum is arcuately prominent ventrally with the coxe, feebly compressed and sometimes subvertically truncate posteriorly. Mesosternum arcuately declivous and rather deeply concave. The abdominal intercoxal salient is small and quite quadrate, one- half of its width shorter than the post-coxal portion of the same seg- ment, the latter subequal in length to the second; third segment a little shorter than the second; the fourth is one-half the length of the second. The abdominal and metacoxal salients are quite equal in width. The metasternum laterally between the coxe is short and equal in length to the width of a mesotibia at middle. The tibial grooves of the femora are well developed, more or less concave externally, becoming flat internally, glabrous and impunc- tate; margins distinct, more or less acute and gradually converging to the femoral base. The protibiz are distinctly compressed, and carinate externally ; carina quite entire, surface longitudinally grooved behind the same, groove asperately sculptured. Meso- and metatibize subcylindrical in section, flattened or grooved externally, grooves opaque, glabrous and more or less punctate. The tarsi are moderate in length and stoutness. REVISION OF ELEODIINI—BLAISDELL. 457 Protarsi about one-fifth of their length shorter than a mesotarsus. First four joints short, subequal in size and length, the first not noticeably longer, each just a little wider than long; the fifth equal to about the preceding three in length. Mesotarsi about one-fourth of their length shorter than a meta- tarsus. The first joint just the least shorter than the next two com- bined; joints two to four inclusive about equal in length and just a httle longer than wide; the fifth about equal to the preceding two combined. The metatarsi are slightly more than one-half as long as a meta- tibia. The first joint is just a little shorter than the next two com- bined; the second and third subequal and distinctly longer than wide, both together equal to the fourth. EMBAPHION GLABRU\M, new species. Suboblong oval, twice as long as wide, nigro-piceous, surface more or less shining, quite smooth, obsoletely sculptured, margins of the pronotum moderately reflexed. [ead about twice as wide as long, plane or feebly convex, slightly impressed laterally, sometimes along the frontal suture, sides not strongly prominent, obsoletely punctulate. Antennw rather slender and equal in length in the sexes, reaching slightly beyond the pro- thoracic base, very shghtly compressed in outer four joints, which are also feebly dilated, third joint equal to the next two taken together, fourth scarcely longer than the fifth, sixth and seventh very slightly shorter and subequal, eighth subtriangular, ninth and tenth sub- orbicular, eleventh ovate. Pronotum proper slightly wider than long and about as wide as one elytron, margins thin and moderately foliaceous, reflexed and concave; disc moderately convex, smooth and obsoletely punctulate, laterally obsoletely granulate and rugulose; apex moderately deeply emarginate, the emargination about four times wider than deep, obsoletely margined; sides evenly rounded from base to apex, not noticeably widening at base; base truncate and about equal to the apex proper, finely margined; apical angles moderately rounded, reaching to the eyes; basal angles rounded and slightly more promi- nent posteriorly than the base. Propleure smooth and impunctate. Elytra oval, width equal to about two-thirds of the length, margins acute, slightly reflexed and not quite attaining the apex; base sub- truncate; humeri proper obtuse and rounded, not anteriorly promi- nent; sédes quite evenly but not strongly arcuate, apex simply sub- acute; disc smooth, more or less transversely concave or feebly bi- concave, sometimes plane between the reflexed margins, frequently 458 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. slightly and briefly arcuately declivous at base, and arcuately de- clivous posteriorly as usual; éufleved sides not strongly oblique, quite plane at the middle, feebly concave beneath the humeri and distinctly so where it passes into the apex; surface smooth and obso- letely punctate, inflexed sides smooth and impunctate. Epipleure smooth, impunctate and obsoletely defined, except at apex; surface on the same plane as the inflexed sides of elytra. Sterna and parapleurew smooth and more or less obsoletely seulp- tured. Abdomen obsoletely sculptured and smooth; third segment more or less oblique to the fourth and fitth segments which have a feeble transverse convexity. Legs moderate in length and thickness. Anterior femora mutic, tibial spurs quite similar and the tarsi are alike in the sexes, and simple. Male—Elytral disc rather obliquely arcuately declivous pos- teriorly, apex acute and just noticeably produced. Abdomen moder- ately convex and broadly impressed on first two segments. Protibive briefly and moderately constricted at base. Female.—Elytral dise arcuately and more or less vertically decliv- ous behind, apex simply subacute, abdomen rather strongly convex. Protibiz gradually and feebly narrowed at base. Measurements.—Male: Length, 17 mm.; width, 8.2 mm. /emales: Length, 15-17 mm.; width, 7.5-8.5 mm. Genital characters, Male—Edeagophore elongate subfusiform, six to eight times longer than wide and more or less arched. Basale elongate and about five times longer than wide, sides more or less feebly arcuate, surface moderately convex. Apicale elongately triangular; dorsal surface moderately convex, without median groove (in the few specimens examined), impunctate 3 sides broadly and feebly sinuate; apex subacute and rather slender; base bisinuate with a median triangular lobe. Sternite as in elongatum. Female.—Genital segment triangular in outline, rather elongate, surface quite plane and not setose. Valvula (Plate 5, fig. 14).—Dorsal plate oblong-triangular; surface very slightly concave, smooth, obsoletely and sparsely punctate; ex- ternal border more or less sinuate; internal border very feebly arcu- ate; apical margin short and more or less feebly rounded or truncate, directly continuous with the dorsal surface of the apex in the internal third, angle rounded, not setose. Apex chitinous, produced and arcuately everted, convex above and coneave beneath, tip rounded, excavated externally at base for the appendage, the latter rather short mammilliform, with a few moderately long setz at tip. Basal prominences obsolete. REVISION OF ELEODIINI—BLAISDELL. 459 Superior pudendal membrane attaining the middle-of the dorsal plate, finely and longitudinally rugulose. Ventrolateral surfaces.—Body somewhat depressed, surface lines straight when viewed longitudinally, moderately convex transversely, smooth and shining; submarginal groove distinct beneath the not ex- planate external margin of the dorsal plate. Internal margins of the valves contiguous in basal four-fifths. Fissure very narrowly fusi- form in apical fifth, membrane not visible. Habitat—Arizona (Winslow, Barber and Schwarz; Holbrook, H. F. Wickham) ; New Mexico (Coolidge, Wickham). Number of specimens studied, 7. Sexitypes (Cat. No. 12235) in the U. S. National Museum collection. Type-locality.—W inslow, Arizona. Salient type-characters.—Surface smooth, more or less shining and obsoletely sculptured. Pronotum with the lateral margins thin and moderately foliaceous, reflexed and concave. Elytral margins acute, shghtly reflexed and not quite attaining the apex; disc more or less shghtly concave. Diagnostic characters—Conspicuous on account of its smooth in- teguments and entire absence of the short sete that are so noticeable in all of the other species. The body has a much greater dorso- ventral thickness than any of the species before me. The sexes are also less differentiated than in any other. It differs from contusum, elongatum, and depressum in not having the marginal lines of the elytra attain the apex. From planum by its moderately foliaceous prothoracic margins, more or less concave elytral disc and in the smooth and obsolete sculpturing. Glabrum appears more robust than any other form on account of the unusual dorso-ventral thickness. General observations—Mentum with the middle lobe slightly smaller than usual, subtruncate anteriorly and feebly straightened laterally, the sides oblique and converging to apex, the outline is more semicircular than otherwise; surface smooth and somewhat punctate laterally. The prosternum is quite prominent ventrally with the coxe, arcuate antero-posteriorly and rather vertical posteriorly, not in the least mucronate. Mesosternum arcuate and more or less vertically declivous, longi- tudinally concave. The abdominal process is slightly transverse, in width scarcely as long as the third abdominal segment; the post-coxal portion of the first about equal to the second, the latter is twice as long as the fourth; the third is about one-half of its length shorter than the second segment: The abdominal and metasternal salients are equal in width. 460 BULLETIN -63, UNITED STATES NATIONAL MUSEUM. The metasternum laterally between the coxe is short, in length about equal to the width of a mesotibia at middle. The tibial grooves of the femora are entire, glabrous, flat to more or less concave, their margins moderate to feeble and acute, converg- ing to the femoral base in each, where they are scarcely contiguous. The external surfaces of the tibize are as in planum. The tarsi moderate as usual. The protarsi are about one-fourth of their length shorter than a mesotarsus. First four joints small and subequal, the first distinctly longer than wide, second, third, and fourth as wide as long; the fifth about equal in length to the preceding three combined. The mesotarsi are about one-fifth of their length shorter than a metatarsus. The first joint is about one and a half times longer than the second ; joints two to four inclusive are subequal in length and a little longer than wide; the fifth about equal in length to the pre- ceding three taken together. The metatarsi are about two-thirds as long as their respective meta- tibia. The first joint is about equal to the second and third combined ; the fourth equal to the first; the second and third are subequal in length and about one and a half times longer than wide. EMBAPHION CONTRACTUM, new species. Oblong-oval, piceous black, feebly shining to subopaque, sides of the body not foliaceous, acute elytral margin not attaining the apex; pronotum constricted in basal eighth; each puncture with a very short, semierect seta. /Tead moderately large, more or less plane to feebly convex between the eyes, rather broadly and distinctly impressed within the supra- antennal convexities, frontal suture more or less feebly evident; surface sparsely, irregularly, and somewhat finely punctate, punctures shghtly denser at the periphery. Antenne long and moderately slender, outer four joints feebly compressed and scarcely dilated; third joint equal to the next two taken together; fourth scarcely longer than the fifth; the latter to the seventh, inclusive, subequal in length: eighth subtriangular and longer than wide; ninth and tenth very feebly transversely oval; eleventh pointed ovate and scarcely longer than wide. Pronotum with the total width about one-half greater than the length, margin acute and not foliaceous; disc proper about as wide as long, moderately, broadly, and evenly convex, sparsely, irregularly, and not strongly punctate, with a more or less feeble transverse anti- basal impression, laterally rather broadly impressed or feebly con- cave within the sheghtly reflexed acute margins, surface here more opaque and obsoletely sculptured; apex broadly, evenly, and moder- REVISION OF ELEODIINI—BLAISDELL. 461 ately emarginate, evidently not beaded; s¢des evenly and moderately arcuate to the basal seventh or eighth, there quite abruptly con- stricted with sides parallel or shghtly convergent to base, the latter slightly arcuate and rather coarsely but not strongly beaded, equal in width to the apex; apical angles very narrowly rounded; basal angles subrectangular. Propleure smooth, opaque, and obsoletely sculptured. Elytra oblong-oval, widest at the middle, a third of the length longer than wide; base feebly emarginate between the rather broadly rounded and slightly more anteriorly prominent humeri, sides evenly and moderately arcuate, more strongly convergent in apical fourth where they are feebly and broadly sinuate, the apex appearing some- what produced and subparabolically rounded; dése flat to feebly con- vex, limited laterally from the obliquely inflexed sides by an acute margin, that does not attain the apex, arcuately declivous pos- teriorly in apical third; surface irregularly and not densely subas- perately punctate, at times there is a distinct tendency to a serial arrangement, the punctures are shghtly denser and a little more muricate at the periphery, the acute margin is subserrulate; inflexed sides more or less obsoletely punctate. Epipleure narrow and not dilated at the base, gradually narrow- ing to apex; surface smooth and impunctate; superior margin not defined in basal half, thence to apex well developed. Sterna quite smooth and more or less obsoletely sculptured. Parapleure rather coarsely and more or less obscurely punctate. Abdomen smooth, very sparsely and finely punctulate, more or less obsoletely rugulose; punctules more distinct and denser on the fifth segment. Legs slender and moderate in length, femora mutic, anterior tibial spurs subequal and the tarsi similar in the sexes. JMale.—Unknown to me, the secondary sexual characters are prob- ably as in planum. Female—Moderately broad. Abdomen not very strongly but evenly convex. Anterior tibie quite gradually and slightly sinuate at base. Measurements —Males: Not studied. Females: Length, 1821.5 mm; width, 9-10 mm. Genital characters, male—Not studied. Female—Genital segment elongate and triangular in outline, dor- sal surface more or less transversely ogival in section, impunctate and not setose. Valvula (Plate 5, fig. 20).—Dorsal plate oblong-triangular, about four times longer than wide; surface feebly convex basally and concave apically; external border more or less sinuous; internal bor- der nearly straight and somewhat beaded; apical margin arcuate in 462 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. external two-thirds, becoming continuous internally with the inner margin of the fossa. Apex produced and not everted, sides sub- parallel, externally continuing the line of the dorsal plate, tip rather broadly rounded, dorsal surface concave; internally the valvular membrane is finely setose and semi-chitinous caudad to the pudendal membrane. Appendage short and nearly hemispherical, projecting very little beyond the fossa, setose at tip, sete long forming a loose pencil. Fossa rather broad and not sharply defined. Superior pudendal membrane reaching to the apical fourth of the dorsal plate and longitudinally rugulose. Basal prominences scarcely at all prominent. Ventrolateral surfaces transversely convex, surface lines straight when viewed longitudinally, concave apically before the apices. Sub- marginal groove distinct beneath the feebly explanate external margin of the dorsal plate. Internal margins of the valves contiguous in basal two-thirds; genital fissure very narrow in apical third. Inferior membrane not visible. Habitat—New Mexico (Cloudcroft, elevation 9,000 feet). Number of specimens studied, 7 (females). Types in my own collection. Ty pe-locality.—Cloudcroft, New Mexico; collector, Warren Knaus. Diagnostic characters.—The seven specimens studied are homomor- phie and distinct from their congeners. The shortening of the ex- panded pronotal margin is quite remarkable, and the basal constric- tion causes the prothorax to appear more distant from the elytra than in depressum. : In fact, the form of the prothorax is an approach to that observed in Hleodes planipennis. Contractum appears as a step toward the bridging over of the gap between Lleodes planipennis and ELm- haphion. Wt is also significant that both should inhabit the same geographical region. Some specimens of planipennis exhibit a ten- dency to a thinning and reflexion of the pronotal margins, but never to an acuteness of the elytral margins. When such examples are found planipennis must be referred to Lmbaphion. In depressum the acute elytral margin attains the apex, in contrac- tum it does not. In planum the acute and expanded pronotal margins reach to and slightly beyond the base, while in contractum they are abbreviated posteriorly and not extending beyond the basal seventh or eighth, the true basal angles being exposed. Contractum is without doubt an extreme and perpetuated modifi- eation of planum. Tn the series of contractum examined the abdo- minal segments are not oblique. REVISION OF ELEODIINI—BLAISDELL. 463 I am indebted to Mr. Warren Knaus for this interesting addition to our list. General observations—The mentum is rather small, feebly trans- verse, more or less arcuate anteriorly, although at times apparently subtruncate, sometimes feebly impressed at apex so as to appear some- what emarginate; surface feebly convex, glabrous, and obsoletely punctate at center, narrowly impressed peripherally within the mar- gin, there opaque and distinctly punctate. Prosternum less suddenly protuberant ventrally than in planwm, otherwise the same. The mesosternum is quite vertical and arcuately declivous, more or less concave as usual. The metasternum laterally between the coxe is as long as the width of a mesotibia at apex. The metasternal and abdominal salients are quite equal in width. The abdominal process is a little longer than wide. The post-coxal portion of the first abdominal segment is about equal to the third in length; the second is about twice as long as the fourth segment. The tibial grooves of the femora are well developed, more or less concave, but not strongly so. The margins are subcariniform exter- nally, becoming more or less evanescent internally. On the profemora the grooves have the margins slightly arcuate and about attaining the femoral base; those of the meso- and metafemora have the mar- gins quite straight, scarcely attaining the base on the former and evanescent at basal fifth on the latter. The protibiz are feebly compressed and distinctly carinate ex- ternally: tarsal grooves more or less obsolete, articular cavities closed. On the meso- and metatibie the grooves are more or less de- veloped and narrow on apical two-thirds, shghtly glabrous, margins asperate, and the articular cavities are closed. The tarsi are the same as in planum. The protarsi are about two to three-fifths of their length shorter than a mesotarsus, joints two to four subequal in length and scarcely as long as wide; the first joint is a little longer than wide; the fifth is Just the least longer than the three preceding joints taken together. The mesotarsi are about two-fifths of their length shorter than a metatarsus. Joints two to four, inclusive, are subequal in length and slightly longer than wide; the first joint is about one-half longer than wide; the fifth is shghtly shorter than the preceding three joints taken together. The metatarsi are distinctly more than a half longer than their respective tibia. Joints two and three are subequal in length and distinctly more than one-half longer than wide, and together as long as the fourth; the first is scarcely as long as the next two taken together. 464 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. EMBAPHION PLANUM Horn. Bmbaphion planum Horn, Trans. Amer. Phil. Soc. Phila., XIV, 1870, p. 321. Oblong-oval, about twice as long as wide; margins of body not foliaceous, acute elytral margins not attaining the apex; surface more or less dull. Head moderate, about twice as wide as long, quite plane between the eyes, feebly impressed laterally; sides of the frons slightly promi- nent, finely, rather sparsely punctate, each puncture with a minute curved seta, frontal suture at times distinct, sometimes slightly impressed. Antenne quite long; outer four joints scarcely com- pressed or widened; third joint about equal to the next two taken together; fourth to the seventh, inclusive, subequal; eighth triangu- lar; ninth and tenth suborbicular; eleventh ovate. Pronotum one-sixth to one-half wider than long and slightly nar- rowed behind; margin acute, not foliaceous nor suddenly formed ; disc feebly convex, finely and rather sparsely punctate, punctures becoming shghtly denser laterally, each with a small curved seta; apex evenly but not deeply emarginate, the emargination four or five times wider than deep, usually very finely margined; sides evenly and moderately arcuate from base to apex; margin slightly explanate, widening a little posteriorly, feebly reflexed; base distinctly shorter than apex, transverse, and obsoletely margined ; apical angles obtusely rounded, attaining the eyes; basal angles obtuse, rounded, and pro- jecting shghtly beyond the middle of the base. Propleure smooth, impunctate, and obsoletely rugulose, defined from the inferior surface of the pronotal margin by a shallow groove. Elytra elongate oval, base at middle shghtly prolonged and feebly sinuate laterally; Awmeral angles distinct and rounded ; margin acute, feebly reflexed and not attaining the apex; sides evenly arcuate, grad- ually converging apically; apex acute and not caudate; disc flattened, at times feebly convex, never concave, evenly and rather gradually arcuately declivous posteriorly ; acutely inflexed laterally, the inflexed sides feebly convex; surface sculptured with subasperate punctures, that may be serially arranged or irregular and somewhat denser laterally ; the inflexed sides are irregularly and rather sparsely punc- tate. Each puncture bears a short curved seta. E'pipleure narrow and not dilated at base, gradually narrowing to apex; superior margin distinetly defined; surface glabrous and im- punctate. Sterna more or less feebly punctate and rugulose, feebly shining. Parapleure quite glabrous, feebly and obsoletely punctate. Abdomen horizontal, more or less shining, sparsely or not very densely punctulate, obsoletely rugulose. Legs moderate and rather slender, anterior femora mutic, tibial spurs similar and the protarsi simple in the sexes. o & PS. REVISION OF ELEODIINI—BLAISDELL. 465 Male—Somewhat narrow. Antenne reaching considerably be- yond the prothoracic base. Abdomen moderately convex, first three segments rather broadly impressed along the middle. Protibiz suddenly and briefly sinuate at base. Female.—More robust. Antenne reaching slightly beyond base of the prothorax. Abdomen rather strongly convex. Protibie gradually narrowing at base. Measurements.—M ales: Length, 14-19.5 mm.; width, 6-9 mm. Fe- males: Length, 13.2-17 mm.; width, 7-8.5 mm. Genital characters, male-—Edeagophore very slender, elongate, and moderately arched. Basale about five times longer than wide, rather strongly convex from side to side, the latter feebly arcuate. A picale small, triangular, shghtly elongate; surface rather strongly convex and without groove; sides feebly and broadly sinuate; apex gradually attenuated and acute; base with a rather short, subacute lobe at middle and feebly sinuate laterally. Sternite transversely triangulo-parabolic in outline. Each lobe triangular, with external border more or less arcuate to apex, the latter subacute and rather gradually narrowed; internal margin nearly straight and oblique; surface more strongly convex along the external border, thence sloping inward, rather densely punctate in apical two-thirds, setose, sete long and rather dense, especially in apical third. Sinus more or less semicircular, membrane more or less setose across the same. Female.—Genital segment (Plate 5, fig. 9) triangular, surface quite plane, impunctate and not setose. Valvula.—Dorsal plate elongately subtriangular, moderately nar- rowed from base to apex; surface plane and glabrous; external border more or less feebly sinuate; apical margin truncate at middle, internally continuous with surface of the apex, angle evenly rounded; internal margin very feebly arcuate. Apex produced and chitinous, more or less everted, convex above and concave beneath, tip rounded. Fossa at base of the external surface of apex and beneath apical mar- gin of dorsal plate. Appendage short mammilliform, with a pencil of four or five rather long setee at tip. Basal prominences moderately developed. Superior pudendal membrane attaining the apical fourth of the dorsal plate, and longitudinally ruguiose. Ventrolateral surfaces——Body slightly protuberant laterally in basal half, feebly concave laterally before the apices; surface lines straight. Submarginal groove shallow beneath the feebly expanded external border of the dorsal plate. Internal margins of the valves contiguous in basal three-fourths; genital fissure in apical fourth 59780—Bull. 68—09——30 466 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. and nearly closed, margins fringed with very small sete. Inferior membrane not visible. Habitat—Kansas (George Horn): Colorado (Denver, April, H. Soltau); Wyoming (Cheyenne, June and July, Hubbard and Schwarz). Number of specimens studied, 34. Type in the Horn collection. Ty pe-locality.—Kansas. Salient type-characters—Thoracic margin acute, not foliaceous nor suddenly formed, not deeply emarginate at apex. Elytral margin acute, feebly reflexed, and not attaining the apex; disc flattened, and never concave. Epipleural limits distinctly defined (Horn). Diagnostic characters —The thorax is distinctly wider than long, the apical emargination is shallower and broader, the elytral dise is flat or feebly convex, and the apices never caudate; the margin does not reach the apex. The epipleurz are usually well defined and even feebly convex. From contusum it differs in the shallower apical emargination of the prothorax, and by the elytral margin not reaching the apex. ‘The prothoracic margin is distinctly narrower. It is recognized from elongatum by the shorter and broader form and by the elytral margin not attaining the apex. From depressum by the wider thoracic and elytral margins, and in not having the marginal line reaching the apex. Planum is undoubtedly very closely related to contusum and a very large series might show it to be simply a subspecies. I deem it best at present not to unite the two until further collecting has been done. The explanate thoracic margin varies in width and the marginal line is often indicated to apex by a line of asperities. General observations—The mentum is moderate, middle lobe more or less semicircularly arcuate anteriorly from the subangulate sides. The apex may be subtruncate and the sides straight and oblique; surface feebly convex and more or less obsoletely foveate laterally. Prosternum and mesosternum as in contusum. Ina few specimens there is a very slight tendency for the prosternum to become feebly prominent posteriorly. The abdominal process is rather small and nearly quadrate, its width is slightly less than the length of the post-coxal portion of the same segment, the latter is about equal to the third in length; the second is one-fifth of its length longer than the third, the latter scarcely twice as long as the fourth. The abdominal and metasternal salients are equal in width. The metasternum laterally between the coxz is short, in length about equal to the width of a mesotibia at middle. ” « 7 : : f : REVISION OF ELEODIINI—BLAISDELL. 467 The tibial grooves of the femora are well developed and defined; surface of each plane to concave, glabrous and at times feebly rugu- lose; margins well defined and more or less acute, gradually con- verging to the femoral base in each instance, those of the profemora are shghtly arcuate. The protibiewe are slightly compressed and distinctly carinate ex- ternally, carina more or less entire, surface behind it rather narrowly and longitudinally grooved, groove more or less asperate. Tarsal grooves of the mesotibiz rather more or less evident ex- ternally in apical two-thirds, grooves more or less narrow, sub- glabrous or subasperate. External surfaces of the metatibie are more or less flattened, scarcely grooved and asperate. The tarsi are moderate in length and stoutness. The protarsi are about one-fourth of their length shorter than a mesotarsus. The first four joints are subequal in length, the first is shghtly longer than wide, the succeeding three feebly wider than long. The mesotarsi are about one-fifth of their length shorter than a metatarsus. First joint scarcely as long as the next two combined; the second, third, and fourth subequal in length and_ distinctly longer than wide; fifth a little shorter than the preceding three taken together. The metatarsi about three-fifths as long as their tibie. First joint quite equal to the succeeding two taken together, the fourth just a little longer than the second and third combined, the latter two are equal to each other. EMBAPHION CONTUSUM LeConte. Embaphion contusum LeContre, Journ. Acad. Nat. Sci. Phila., 2d ser., IV, 1858, p. 20; Smithson. Contrib. to Knowledge (Coleopt. Kans. and east- ern N. Mex.), 1859, p. 15, pl. 1, fig. 8—Horn, Trans. Amer. Phil. Soe. Phila) XLV; 1870; p. 321: Var. laminatum Casry, Ann. N. Y. Acad. Sci., V, Nov. 1890, p. 408. Elongate oblong, piceous black, more or less opaque, sides of the prothorax rather broadly foliaceous, elytral margins less strongly so, margins distinctly attaining the apex. Head small, less than twice as wide as long, nearly plane, broadly and feebly impressed laterally, sides of the frons rather prominent ; surface very finely, sparsely, and quite indistinctly punctate, each puncture bearing a small, short, and curved seta. Antenne long and rather slender, very feebly compressed in outer four joints, which are feebly widened; third joint a little shorter than the next two taken together; fourth shghtly longer than the fifth; the latter, sixth, and seventh subequal; eighth shghtly shorter and subtriangular; ninth and tenth suborbicular; eleventh ovate. 468 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Pronotum proper about as wide as long, lateral margins thin and moderately foliaceous, more or less reflexed and concave; disc rather moderately convex, finely and rather sparsely punctate, laterally more densely so, each puncture with a short curved seta; apex rather deeply and somewhat semicircularly emarginate, the emargination about three times wider than deep, not noticeably margined; sides evenly rounded, margin not widening at base; base quite truncate, and about equal to the apex, with a distinct marginal bead; apical angles obtuse, formed by the moderately advanced foliaceous mar- gins, and are about half as long as the head: basal angles broadly rounded, slightly more prominent posteriorly than the middle of the base. Propleure smooth and quite impunctate. Elytra elongate oval, nearly twice as long as wide, margins acute, slightly reflexed and always attaining the apex; base quite truncate, much wider than the contiguous prothoracic base proper; humeral margins rather broadly rounded, not anteriorly prominent, humeri proper obtuse ; sides feebly arcuate or subparallel in basal half, thence to apex quite evenly and broadly arcuate, apex more or less acute and produced ; disc quite plane or feebly concave, sometimes biconcave by elevation of suture in posterior half, more or less arcuately de- clivous posteriorly, acutely inflexed laterally, inflexed sides quite straight and oblique; surface sculptured with subasperate punctures which may be rather densely placed and more or less irregularly arranged, at times somewhat serially so, occasionally the punctures are not asperate and rather coarse; inflexed sides irregularly and more or less densely punctate. E pipleure narrow, not in the least dilated at base, very gradually narrowed to apex, smooth and impunctate; superior margin well defined in apical fourth, thence to base by a series of punctures, surface on the same plane as the inflexed sides of the elytra; widening at apex to become the inferior surface of the more or less produced apex. Sterna quite smooth and more or less impunctate. Parapleurw rather sparsely and more or less indistinctly punctate. Abdomen smooth, very sparsely and more or less indistinctly pune- tulate, obsoletely rugulose; third segment scarcely oblique, others horizontal. Legs moderate in length, rather slender. Profemora mutic as usual, tibial spurs similar and protarsi simple in the sexes. Male.—Somewhat narrow. Antenne reaching considerably beyond base of the prothorax. Elytral disc more or less gradually declivous behind, apex produced into a short cauda, which may be angulate at the sides where the inflexed sides terminate; cauda more or less abruptly produced, rather oblique, and more or less asperate, with REVISION OF ELEODIINI—BLAISDELL. 469 tips separated. Abdomen moderately convex and more or less im- pressed on first two segments. Protibize suddenly and briefly sinuate at base. Female—Rather broad. Antenne reaching beyond the protho- racic base, but shorter than in the male. Elytral dise arcuately de- clivous posteriorly, not produced at apex, but more or less acute. Abdomen quite strongly convex. Protibie gradually narrowing at base. Measurements —Males: Length, 15-20.5 mm.; width, 7.2-8 mm. Females: Length, 14-19.5 mm.; width, 7-10 mm. Genital characters, male.—Edeagophore slender, elongate oblong- ovate. Basale elongate oval, surface moderately convex, sides moderately arcuate, apex deeply and triangularly emarginate to receive the base of the apicale. Apicale slender and elongately triangular; surface evenly and rather strongly convex, without groove; sides nearly straight, gradu- ally converging to apex, the latter acute; base strongly and acutely lobed at middle, oblique and broadly sinuate laterally. Sternite subparabolic in outline. Each lobe subtriangular, external border quite evenly arcuate to apex, the latter subacute, prominent and shghtly produced; internal border sinuate; surface more strongly convex along the external border, thence sloping inward, densely punctured in apical two-thirds, especially internally, setose, sete quite long and dense, more so on apex. Sinus semicircular, membrane not setose across the same. Female.—Genital segment (Plate 5, fig. 8) elongately triangular with the apices, or triangular-trapezoidal without the same, superior surface plane, not setose. Valvula—Dorsal plate suboblong to feebly and elongately tri- angular, slightly narrowing from base to apex; surface plane, smooth, and not noticeably punctate; external border angulate at basal fourth, thence more or less sinuate and arcuate to apical margin, the latter subtruncate at middle two-fourths, angle rounded, in inner fourth continuous with the surface of apex; internal margin quite straight or more or less feebly arcuate. Apex chitinous, produced, and everted, convex above and concave beneath, rounded at tip; fossa at base of the external apical surface, beneath apical margin of dorsal plate. Appendage mammilliform, glabrous, with a pencil of few quite long setee at tip. Basal prominences feebly evident, lateral surfaces of valves slightly visible from above. Superior pudendal membrane attaining the apical fourth of the dorsal plate, finely and longitudinally rugulose. 470 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Ventrolateral surfaces—Body with surface. lines straight when viewed longitudinally, quite evenly convex from side to side, smooth and shining. Submarginal groove rather small beneath the external border of dorsal plate, which is feebly explanate in apical half, termi- nating at the apical margin. Internal margins of the valves con- tiguous in basal four-fifths; fissure quite short, nearly closed, mar- gins finely setose. Inferior membrane not visible. Habitat—Wyoming (Cheyenne, June, Hubbard and Schwarz; April, H. Soltau); Colorado (Salida, Fort Collins, Clear Creek, South Park Region, San Luis Valley, H. F. Wickham) ; New Mexico (Deming, July and August, Hubbard and Schwarz; Las Vegas, August, Barber and Schwarz; Coolidge, Wickham) ; Arizona (George Horn; Winslow, Hubbard and Schwarz; Santa Rita Mountains, July, Hubbard and Schwarz; Tucson, Charles Fuchs) ; Kansas (F. H. Snow; George Horn). Number of specimens studied, 48. Type in the LeConte collection. Type-locality.—F ort Laramie and Santa Fé, New Mexico. Salient type-characters—Opaque. Thorax with deeply emargi- nate apex, margin broadly explanate, moderately reflexed, apical angles subacute, basal angles broadly obtuse. Elytra with the dorsum plane, acutely margined at apex, margin reflexed, strongly so, seri- ately, finely and rather densely muricato-punctate. Male with elytral apices briefly prolonged (LeConte). Diagnostic characters.—In this species the thorax proper is broader, the prothoracic margins are moderately foliaceous and reflexed, the integuments are more or less opaque, finely and subasperately sculp- tured, the elytral margins attain the apex. The males are briefly caudate. From muricatum it differs by not having the basal angles of the prothorax prolonged backward over the humeri. The elytral margin is thinner, narrower, and usually scarcely reflexed unless it be near the humeri. The sides of the pronotum are rounded from apex to base and not broader behind, the basal angles are broadly rounded and shghtly more prominent posteriorly than the middle of the base. In contusum the males have the inferior surface of the elytral cauda form by the dilated apices of the epipleural surface. From glabrum it is recognized by the dull luster and finely sub- asperate sculpturing, also by the acute elytral margins attaining the apex. In the U. S. National Museum collection there are two males and two females which were collected in New Mexico. They are the largest and most strongly developed specimens that I have seen. The elytral apices are distinctly caudate, and the elytral dise quite evenly concave transversely in the males and the suture is feebly raised Sas te SSRN, ee Nel Sl ie ey Ly SP Sr 1 Valo REVISION OF ELEODIINI—BLAISDELL. 471 posteriorly, indicating an approach to laminatum Casey. The females have the elytral dise broadly oval and quite flat. The epi- pleure are not at all defined from the inflexed sides except at apex, they are glabrous. In these specimens the sexes have the prothoracic margin slightly broader and more decidedly reflexed than the average number of specimens of the typical form (forma typica); in fact, some specimens have the margins scarcely at all reflexed. These large specimens may be placed as an extreme form in the cabinet, and labeled forma grandis. General observations—Mentum more or less variable and moderate in size. Middle lobe arcuate anteriorly, or subtruncate with the sides straight and converging to apex; surface very feebly concave, ‘ather longitudinally prominent at middle and broadly, feebly con- cave laterally and more or less punctate. The prosternum between the cox is quite suddenly prominent ventrally, antero-posteriorly arcuate, frequently more or less grooved. Feebly widened behind the equator of the coxe. The mesosternum is more or less obliquely and arcuately declivous, more or less broadly and feebly concave. The abdominal salient subquadrate and scarcely as wide as the third segment is long; post-coxal part of first about equal in length to the second segment, the latter twice as long as the fourth; third about one-third of its length shorter than the second. The abdominal and metasternal processes are subequal in width. The metasternum laterally between the coxe is very short and about equal to a mesotibia at base. The tibial grooves of the femora are entire, well defined and gla- brous; their margins are subacute and smooth. The profemoral grooves are more or less strongly concave; the margins converge to become contiguous at the femoral base. The mesofemora have the grooves scarcely concave; the margins converge to become feebly distant at femoral base. The grooves on the metafemora are not concave, quite flat, with margins gradually converging and slightly distant at base. The protibiz are slightly compressed and carinate externally, the carina is entire, the tarsal groove is immediately behind it and is frequently obsolete, the surface always asperate. The mesotibie are more or less distinctly grooved externally in apical two-thirds; the surface of the groove is glabrous, with sparsely placed asperate punctures. The metatibiz are simply flattened externally and asperate. Tarsi moderate as usual. The protarsi are scarcely one-fourth of their length shorter than a mesotarsus; joints two, three, and four subequal in size and very slightly wider than long and together about equal to the fifth; the 472 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. first is a little longer than the second; mesotarsi about two-sevenths of their length shorter than a metatarsus; joints two, three, and four subequal, a little longer than wide, and together just a little longer than the fifth; the first about one and a half times longer than wide. The metatarsi are about. one-half of their length shorter than their metatibia; first and fourth joints equal in length and either one is equal to the second and third taken together; the latter two are sub- equal in length, each about twice as long as wide. EMBAPHION CONTUSUM var. LAMINATUM Casey. Moderately robust, black and dull throughout. Head small, very finely, sparsely, and subasperately punctate. Antenne slender, third joint from four to nearly five times as long as wide, eighth one-half longer than wide. Pronotum about one-half wider than long; disc proper very feebly convex, fully as long as wide, and having two feeble, subparallel, and sinuous impressions near the middle; surface throughout finely, ex- tremely sparsely, and subasperately punctate; apex strongly emargi- nate, the emargination not quite three times as wide as deep; side margins very widely and strongly reflexed, the outer edges parallel, strongly and evenly arcuate throughout; base transverse; basal angles broadly rounded and projecting beyond the median portion of the base. Elytra from one-half to four-fifths longer than wide; base trans- versely truncate; humeri rounded; sides very thin and broadly but not abruptly reflexed, the acute lateral edges parallel and feebly arcuate in basal two-thirds, then rounded to apex and slightly pro- longed, uniting in a prolongation of the suture; dése distinctly wider than the prothorax, each elytron broadly concave, the suture elevated ; inflexed sides nearly flat, somewhat coarsely and sparsely but un- evenly punctate; surface of disc with approximate, imperfectly defined and feebly impressed rows of rather coarse, impressed punc- tures, also finely and sparsely asperate. Epipleure very imperfectly defined except near the apex. Legs very slender. Male.—Somewhat slender. Female——Moderately robust. Measurements.—Lenegth, 14-15 mm.; width, 6.8-7.5 mm. Genital characters not studied. Habitat—Texas (El Paso, G. W. Dunn). Unknown to me in nature. Type in Colonel Casey’s collection. T'ype-locality.—K | Paso, Texas. REVISION OF ELEODIINI—BLAISDELL. 473 Casey writes that this form is related only to contusum, and I can not help deciding that it is only one of the extreme forms of that species. I do not consider that the peculiar form of the upper sur- face of the elytra can give specific standing in this instance. It is true that in the typical contusum the elytral dise is plane, but in the series before me are examples with flat elytral discs; others have them slightly concave, with margins more strongly reflexed, until finally the extreme forms approach the one here described under contuswm as forma grandis, for example; the latter is much larger than dami- natum, the male having a length of 20.5 mm., width 10.5 mm.; the female, length 19 mm., width 10.5 mm., and is distinctly a much larger form. The elevation of the elytral suture is observed in some examples of all of the species known to me. The sculpturing is quite variable and in the extremes of a series quite in contrast. Furthermore, the elytral punctures referred to in the description are merely impressed fovew, and according to the original remarks do not appear to be in the least setigerous: “ the finer, but strong and sparse, asperities are distributed over the sur- face without regard to the punctures, and each bears a strong, thick, semi-erect seta.” After a careful examination of the sculpturing under high power, I consider that the asperities represent the true punctures, whose margins have been produced into the minute eminences, each bearing a seta. The minute fovez or dents are not the true punctures. The sculp- turing is quite similar in all of the species before me; even the minute fovezx are scattered here and there. The sculpturing of laminatwm is peculiar only in the apparent abundance of fovex. A comparative study of the elytral sete shows that in glabrum the setee are the most minute and the stoutest in contuswm, the longest and most slender in planum. Not having seen the types nor an authentically identified specimen, my views are, of course, founded upon the series at hand and the laws governing homology and analogy. EMBAPHION MURICATUM Say. Akis ’? muricata Say, Journ. Acad. Nat. Sci. Phila., III, 1824, p. 251. Embaphion muricatum LECoNTE, Complete Writings of Thomas Say, IT, 1859, p. 149; in Thomson’s Arcana Nat., I, 1859, pl. x11, fig. 10.— Horn, Trans. Phil. Soc. Phila., XIV, 1870, p. 320. Embaphion concavum LeContr, Proc. Acad. Nat. Sci. Phila., VI, 1853, p. 446. Oval to oblong-oval, brownish to piceous black, thoracic and elytral margins very broad and foliaceous, strongly reflexed. 474 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Head small, less than twice as wide as long, plane, sides of the frons shghtly prominent, punctate, punctures very feebly subasperate, fine, not dense, each with a small curved and short seta, frontal suture usually not visible. Antenne rather long, quite slender, outer four joints very shghtly compressed and scarcely widened, third joint shorter than the next two taken together, fourth scarcely longer than the fifth, the latter and sixth subequal, seventh shorter, eighth feebly shorter than the seventh and slightly triangular, ninth and tenth suborbicular, eleventh subovate. Pronotum with margins very broadly foliaceous, the margin more than one-half wider than the disc, the latter comparatively narrow, longer than wide at middle, very feebly convex, usually with irregular depressed areas; finely, more or less subasperately and sparsely punc- tate; reflexed margins wider posteriorly and more or less concave, a little more distinctly punctate, punctures less sparse, each with a short curved seta; apex deeply and feebly subquadrately emarginate, the emargination about one-half wider than deep, sides almost parallel, and scarcely margined; s¢des evenly but not strongly arcu- ate, moderately converging from base to apex; base proper feebly arcuate, not margined and about equal to the length, laterally sinuate ; apical angles rather narrowly rounded and formed by the advanced foliaceous margins and nearly as long as the head; basal angles are posteriorly prominent, subacute, and projecting backward over the basal angles of the elytra. Propleurw smooth and impunctate; inferior surface of the foli- aceous margins obsoletely punctate. Elytra oval to elongate oval; margins broad and reflexed, angles at humeri nearly rectangular and more or less truncate at base, pos- teriorly extending beyond the apex, the two meeting on a line with the suture above the true elytral apex, and defined from the same by a sight groove, borders evenly arcuate from base to apex or more or less parallel basally; base evenly but not strongly emarginate;” humeri proper broadly rounded and not prominent; s¢des proper more or less evenly arcuate; a@pea proper not produced and narrowly rounded; disc plane, feebly convex, at times slightly concave, the inflexed sides nearly straight and oblique, gradually and not strongly arcuately declivous behind; surface sculptured with approximate series of fine asperate punctures, which become more irregular and slightly denser laterally. Each puncture bears a short and rather robust curved seta. Epipleure narrow, not attaining the humeral margin and not dilated, but gradually narrowing to apex, not defined from the in- flexed sides of the elytra and on the same plane; superior margin obsolete, except near apex; elsewhere represented by a line of punc- tures or a faint groove. REVISION OF ELEODIINI—BLAISDELL. ale Sterna more or less dull, finely and not distinctly sculptured. Parapleure smooth, rather sparsely but not very distinctly punctate. Abdomen horizontal, very finely and sparsely punctulate, obso- letely rugulose and quite evenly convex. Legs rather slender, moderate in length. Anterior femora mutic, protibial spurs similar in the sexes, the anterior slightly longer than the posterior. Protarsi simple. Male——Abdomen moderately convex and not noticeably impressed ; anterior tibiz quite suddenly and briefly constricted at base. Female.—Abdomen rather strongly convex and the protibie grad- ually narrowed at base. Measurements —Males: Length, 16-18 mm.; width, 9.5-10 mm. Females: Length, 15-18 mm.; width, 9—10.5 mi. Genital characters, male—KEdeagophore elongate oblong-ovate, rather slender, feebly arched and slightly depressed. Basale oblong, very moderately convex, sides feebly arcuate. Apicale triangular, shghtly depressed; surface rather feebly con- vex and without groove; sides nearly straight or feebly and broadly sinuate to apex, the latter acute and not attenuated; base distinctly subacutely lobed at middle, feebly and broadly sinuate laterally. Sternite quite parabolic in outline. Each lobe rather long, with external border more or less evenly arcuate or straight in basal half, thence feebly arcuate to subtruncate to apex, the latter narrowly rounded and rather prominent; internal border oblique and short; surface scarcely convex, rather densely punctate im apical two-thirds, setose, setee rather long and not dense. Female.—Genital segment (Plate 5, fig. 7) triangular, rather elon- gate, including apices, or subtrapezoidal without the same, surface quite plane and not noticeably setose. Valvula.—Dorsal plate rather elongate-oblong or oblong-oval ; sur- face scarcely concave, smooth, very sparsely and somewhat coarsely punctate; external border more or less feebly arcuate and _ slightly explanate; internal border arcuate; apical margin more or less trun- cate, angle rather broadly rounded, internally continuous with the surface of apex, not setose. Apex chitinous, produced, slightly longer than the width of dorsal plate at basal third, everted, convex above and concave beneath, tip rounded, external border feebly ex- cavated at base and slightly beneath the apical margin of dorsal plate, forming the fossa. Appendage somewhat spherically mammilliform, glabrous, with a pencil of quite long sete at tip. Basal prominences obsolete. Superior pudendal membrane attaining the middle of the dorsal plate, finely and longitudinally rugulose. 476 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Ventrolateral surfaces—Body somewhat elongately triangular, obliquely truncate laterally at base, quite acute at the internal mar- gins of valves in the median line at base; surface lines straight when viewed longitudinally, evenly convex from side to side, smooth and shining. Submarginal groove well defined beneath the feebly ex- planate external border of the dorsal plate, terminating at the apical margin. Internal margins of the valves contiguous in basal four- fifths; fissure narrowly fusiform in apical fifth, margins finely setose, inferior membrane not visible. Habitat.—Texas (Mobeetee, July, H.S. Barber) ; Colorado (Pueblo, October 27, H. Soltau; Colorado Springs, Greeley, June, elevation 6,000-7,000 feet, Wickham; Fort Collins, Bellevue, West Las Animas, Wickham); Kansas (Wallace County, elevation 3,000 feet, F. H. Snow; Rice County, Clark County, June, elevation 1,962 feet, F. H. Snow); Nebraska; South Dakota (Alexandria). Number of specimens studied, 40. Type destroyed. Ty pe-locality — Arkansas at the Rocky Mountains,” (Say.) Salient type-characters.—Margins of the thorax and elytra broadly foliaceous and strongly reflexed; basal angles of the prothoracie mar- gins projecting strongly backward over the basal angles of the elytra. Diagnostic characters.—Distinct from all other species as indicated by the sahent type-characters. The concavum of LeConte is simply a larger, longer, and narrower form, with the broad elytral margins more suddenly and almost perpendicularly reflexed. General observations —The mentum is moderate in size and vari- able. The middle lobe may be more or less arcuate anteriorly, or truncate, evenly slightly emarginate at apex; sides more or less straightened, and converging anteriorly. The form anterior to the lateral and subbasal angles is subparabolic or subtriangular. Prosternum arcuate antero-posteriorly and somewhat prominent ventrally with the coxee; sometimes more or less oblique and feebly compressed posteriorly. Mesosternum variable as to the degree of obliquity; usually sub- vertically arcuate and broadly, feebly concave. The abdominal process is subquadrate and just the least transverse, and equal in width to the length of the third segment; the post-coxal part of the first segment about equal in length to that of the third; the second is twice as long as the fourth; the third is about one-third of its length shorter than the second. The abdominal and metasternal salhents are quite equal. The metasternum laterally between the coxz is about equal in length to the width of a mesotibia at apex. REVISION OF ELEODIINI—BLAISDELL. 477 The tibial grooves of the femora are entire and well defined, gla- brous, concave on the profemora, less so on the meso- and meta- femora; the margins are acute and smooth, gradually converging to the femoral base, where they meet. The protibiz are scarcely at all compressed nor carinate exter- nally, simply flattened or feebly concave in apical half, forming a shght tarsal groove, which is moderately asperate. The meso- and metatibiz are very feebly flattened externally in apical half. The tibiz appear to be quite variable as to length. In some ex- amples the protibiz are scarcely more than one-half the length of the metatibiea; while in others they are at least three-fifths as long. This seems to be an individual variation observed more or less throughout the tribe Eleodiini. The protarsi are about one-fourth of their length shorter than a mesotarsus. The four basal joints are small, the second, third, and fourth subequal in size and length and about as wide as long; the first is about one-half longer than wide; the fifth about equal to the three preceding joints taken together. The mesotarsi are about one-fifth of their length shorter than a metatarsus. Joints two, three, and four are subequal in length and shghtly longer than wide; the first is distinctly more than one and a half times longer than wide; the fifth about equal in length to the preceding three combined. The metatarsi are about half as long as a metatibia. The first joint is about equal to the fourth; the second and third are subequal, the second apparently just a little longer than the third, and both taken together about equal to the fourth. Genus ELEODIMORPHA, new. . Body rather robust and blapyliform. Head not deeply inserted, epistoma distinctly emarginate, sides of the front straight and converging anteriorly and not in the least dilated, frontal suture scarcely evident; eyes narrow, transverse, and subreniform; antenne 11-jointed, outer three joints feebly com- pressed. Mentum moderate, trilobed, lateral lobes rather small and inflexed, middle lobe trapezoidal, narrowing at base as usual and attached to a short gular peduncle, surface concave. Maxillary palpi moderate in length, last joint triangular and mod- erately broad; third subtriangularly obconical, a little longer than wide and about two-thirds as long as the second, the latter elongate and obconical. Labial palpi small, last joint more pyriform than triangular, con- stricted at base on internal and lateral surfaces, a 478 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Buccal processes of the gens strongly and subacutely produced anteriorly over the base of the maxille. Mandibles emarginate at tip, cusps apparently equal. Prothorax woderately transverse and margined; pronotal apex deeply and broadly emarginate (Plate 12, fig. 89), base truncate; pro- sternum longitudinally impressed before the cox, prominent in the median line and continuously so with the intercoxal portion, which is mucronate posteriorly ; anterior margin of the prosternum arcuate at middle and broadly sinuate laterally. Elytra with the disc not strongly deflexed laterally, distinctly margined by the superior border of the wide epipleuree which occupy the whole of the inflexed side, the superior epipleural margin visible from above throughout its entire length. Mesosternum and metasternum short and not prominent ventrally. Anteriorly the mesosternum is concave, posteriorly meeting the meta- sternal intercoxal salient at middle of the cox, suture shghtly arcu- ate. Metasternal epimera very distinct. Intercoxal process of the first abdominal segment rectangular and broad, as usual. Legs rather short, femora moderately compressed and widest near the middle; tibie slightly but distinctly dilated apically, feebly but characteristically arcuate, spurs small and normal; tarsi moderate in length and thickness, joimts proportioned as in the other members of = _ the tribe, spinous beneath with the plantar grooves distinct. The tribal characters are distinct and the species has a very Blapylis-like form and the general structure is the same. The types were for years associated with Hleodes parvicollis, scabrosa, and err MOLE - i, 1h arate ah Ra Pe Tt a Ne te Ag ae “ip 2eee cordata in my collection. | The produced buccal processes of the gene are as marked as in Nyctoporis. Eleodes blanchardi presents the same condition, but to a much less degree. The maxillary galea is evidently quite different from that observed in Lleodes and apparently two-jointed. I could not determine posi- tively without dissection. . Salient generic characters—Epistoma emarginate, sides of front not dilated, buccal processes of the gene produced (Plate 8, fig. 24), mentum concave; impressed sides and prominent middle of the pro- sternum, the latter with the anterior margin arcuate at middle and sinuate laterally (Plate 8, fig. 24%); wide epipleure, elytral disc not inflexed at the sides; femora compressed and widest at middle, tibiz arcuate and evidently dilated apically. These characters define a very distinct-genus, very unlike anything else found in the United States. From its southern habitat I suspect that it may be related to some Mexican form and have carefully searched all the literature to de- termine that fact, but without avail. At present I have no way of REVISION OF ELEODIINI—BLAISDELL. 479 verifying the results of my previous search and consider it a new and very remarkable addition to our fauna. Type of the genus.—Kleodimorpha bolean. ELEODIMORPHA BOLCAN, new species. Feebly ovate, somewhat robust; a little more than twice as long as wide; black, somewhat opaque; feebly setigerous beneath; dorsally the very minute sete scarcely project out of the punctures. Head somewhat wide, almost plane, punctate, punctures coarse, confluent on the epistoma and laterally, irregular and sparse centrally, ‘seattered on the vertex; sides of the frons scarcely prominent and feebly impressed within; frontal suture slightly evident laterally. Antenne moderate in length and stoutness, outer three joints feebly compressed and slightly dilated, third joint not five times as long as the second, scarcely as long as the fourth and fifth taken together ; joints 4 to 8 rather short, somewhat obconical, the fourth just the least longer than the fifth, the latter, sixth, seventh, and eighth sub- equal; ninth and tenth slightly transverse and oval; eleventh short, ovate. Pronotum widest just in advance of the middle, transverse, about one-half wider than long; disc evenly and moderately convex, rather coarsely punctate, punctures simple, irregular, and sparse centrally, becoming denser and submuricate laterally ; apex broadly and deeply emarginate, bottom of the emargination rather straight at middle three-fifths, about equal to the length, marginal bead not very coarse ; sides evenly and broadly arcuate, moderately sinuate in basal fifth, very briefly and feebly so just behind the apical angles, marginal bead fine; base truncate, rather coarsely beaded, and about one-fifth wider than the apex; apical angles anteriorly prominent and subacute; basal angles rectangular. Propleurew sparsely and submuricately punctate, and more or less rugulose, moderately convex, not in the least concave beneath the pro- notal margins, except at the apical angles. Elytra oblong oval, widest at the middle, a little more than one-half longer than wide; base truncate, scarcely to a little wider than the contiguous prothoracic base; Awmert not in the least prominent, obtuse, angle almost distinct; s¢des evenly, moderately, and broadly arcuate; apex obtuse; disc evenly convex from side to side, less strongly so toward the base, more strongly so toward apex, where it is arcuately declivous, not inflexed laterally; surface evenly and not very densely muricately punctate, punctures evenly arranged in evi- dent series, each puncture distinctly defined from the others; the strial are smaller and less muricate, the interstitial are larger and more evident as small shining muricate tubercles, both series less strongly 480 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. developed centrally, at the periphery both stronger and subequal in size, becoming irregular about the apical declivity. Epipleurw quite broad, occupying the whole of the inflexed sides of the elytra; surface smooth and opaque, sparsely and finely sub- asperately punctate; scarcely narrowing in basal two-thirds, thence gradually to apex; superior margin visible from above throughout its entire length, forming a distinct and subacute bead for the elytra. Sterna—Prosternum impressed before the coxee, prominent at middle and continuously so with the intercoxal portion, continuing forward to the apical arcuation and posteriorly ending in a conical mucro; surface sparsely and subasperately punctate and feebly rugulose laterally, on the central prominent portion coarsely muricato- tuberculately punctate, and with erect brownish hairs. Meso- and metasterna irregularly and = sparsely punctate and more or less rugose. Mesosternum with brownish hairs. Parapleure coarsely but not densely punctate. Abdomen somewhat shining, first three segments rather densely punctate and more or less rugose, the small asperities are shining and larger centrally; last two segments more finely and densely punctate in their apical moieties, quite impunctate basally. Legs moderate in length and thickness. “Femora moderately com- pressed, widest near the middle, mutic in both sexes; tibiae moderately arcuate in both sexes; spurs small and similar. Tarsi with the plan- tar grooves entire, marginal and apico-marginal spinules as usual and ferrugineous in color. Male—Slightly narrower. Elytra arcuately and just the least obliquely declivous posteriorly. Abdomen subhorizontal, moderately convex, rather strongly, broadly and longitudinally impressed at middle of the first three segments; intercoxal process concave; punc- tuation finely muricato-tuberculate. Legs stouter, and the protarsi moderately stout. Female.—Slightly broader and a little more robust. Elytra arcu- ately and vertically declivous posteriorly. Abdomen evenly convex, intercoxal process feebly impressed, segments rather sparsely and finely subasperately punctate. Legs less stout, anterior tarsi slender. Measurements—Male: Length, 13 mm.; width, 5 mm. Female: Length, 13 mm.; width, 5.8 mm. Genital characters, male-—Edeagophore (Plate 6, fig. 11) obtusely oblong-ovate in form, well chitinized, moderately elongate. Basale oblong, about two and a half times longer than wide, rather evenly convex, slightly depressed along the middle, with a faint indi- cation of a median longitudinal linear impression. Apicale vather broadly subpyriform; apical third strongly and quite suddenly deflexed, becoming immediately horizontal and lami- nate, broadly rounded at apex; dorsal surface glabrous with basal REVISION OF ELEODIINI—BLAISDELL. 481 two-thirds somewhat depressed, laterally more arcuate and declivous, circularly membranous at center; sides arcuate at basal third, thence sinuate to apical third, where it is arcuate and continuously so with the broadly rounded apex; base feebly lobed at middle, broadly and feebly sinuate laterally. Clava (Plate 7, fig. 8) comparatively broad, flattened, and of nearly equal width throughout, obtuse and rounded at apex, slightly arcuate, with convexity outward, not. contiguous in the median line, except at base, ventrad to the edeagus; ale of the apicale evidently differentiated as separate elongate sclerites (swbalw), but united by a suture to the apicale. Sternite transversely parabolic in outline, bilobed. Each lobe sub- triangular; surface not strongly but slightly unevenly convex, gla- brous, rather sparsely punctate and pubescent in posterior two-thirds, pubescence denser in the apical area, hairs yellowish; external bor- der straight and oblique in basal three-fifths, thence straight and obliquely subtruncate to apex, the latter narrowly rounded; internal border quite straight; interlobar membrane not pubescent. Sinus triangular. Female.—Genital segment (Plate 6, fig. 12) triangular, about as long as wide at base, superior surface quite plane, chitinized through- out, apically very finely pubescent. Valvula.—Dorsal plate not defined from the apex; surface gla- brous, plane, very sparsely and finely pubescent apically; external border rather straight, rounded at the angle and directly continuous with the feebly and broadly sinuate superior margin of the fossa; internal border broadly sinuate in apical half, thence feebly arcuate to the base. Apex short, rounded at tip, chitinous, fossa on its external surface, finely pubescent, hairs short but noticeable. Appendage mammilliform, short, penicillate, hairs quite long. Superior pudendal membrane long, reaching to the base of the apex, very minutely longitudinally rugulose. Basal prominences noticeable. Ventrolateral surfaces quite evenly convex in basal two-thirds, glabrous, here and there a minute hair, very feebly impressed before the apex. Submarginal groove distinct beneath the feebly explanate external border of the dorsal plate, subobsolete in basal fourth. Apex feebly convex beneath. Appendage and fossa distinctly visible from below. Internal margins of the valves contiguous, genital fis- sure apical, small, and fusiform. Habitat—Bolcan Mountain, between Warner’s Ranch and Julian, San Diego County, California. Taken from beneath a log. Number of specimens studied, 2 (male and female). 59780—Bull. 68—09 ol 482 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Sexitypes in my own collection. Ty pe-locality.—Bolean Mountain. Salient type-characters—Body blapyliform, surface dull, buccal processes of the genx produced; prothorax deeply emarginate at apex, angles anteriorly prominent and subacute, base truncate; pro- sternum impressed before the coxx, prominent at middle and pubes- cent. Elytra truncate and more or less coarsely margined at base, adapted to the prothorax; sculpturing of evident series of small shining muricate tubercles; epipleuree wide, forming the inflexed sides of the elytra. Legs short, femora compressed, tibiz arcuate, tarsi simple. Diagnostic characters.—Distinct as defined by the salient type characters, and can not be confused with any other species in our fauna. General observations—The mentum is trapezoidal, coarsely punctate and setigerous, surface concave. Prosternum between the cox feebly grooved, not prominent ventrally and with the anterior median prominence forming a con- tinuous and feeble arcuation from the apical margin to the tip of the conical mucro. Mesosternum deeply concave to receive the prosternal mucro, sides prominent around the acetabula, pubescent. The intercoxal process of the first abdominal segment is rectangu- lar, slightly transverse, equal in length to the post-coxal portion, its width scarcely equal to the combined lengths of the third and fourth segments. In the male the post-coxal part of the first segment is about equal in length to that of the third, the second is a little longer than the third. In the female the post-coxal part is about equal in length to the third, the second scarcely twice as long as the fourth. The metasternal salient is about a fourth (male) to a third (female) narrower than the abdominal salient. The metasternum laterally between the coxe is about as long as the width of a mesotibia at middle. The superior and inferior surface lines of the femora are distinctly arcuate, most noticeable in the inferior at middle; the tibial grooves are well defined and subentire, the floor of each is flat, smooth, and abruptly limited by cariniform margins, they are longitudinally convex, the convexity is adapted to the tibial arcuation. In the male the lateral surfaces are longitudinally impressed along the inferior margins, not so in the female where the femora are a little less stout and the grooves limited by thinner margins. REVISION OF ELEODIINI—BLAISDELL. 483 The tibiew are arcuate and each is somewhat rounded in basal half and thence to apex somewhat compressed and distinctly but not strongly dilated. The protibiz are the most compressed and scarcely carinate exter- nally, the tarsal grooves are evident but feeble and the posterior border of each more prominent than the anterior. The tarsal grooves of the mesotibiz are the best developed, their margins are equal, The external surface of the metatibize are flattened and more or less grooved. | The articular cavities of the protibiz are nearly open, on the other tibiz they are shut off from the grooves by well marked but somewhat atrophic edges. The relative lengths of the tarsi and also of their joints are quite similar in the sexes. In the protarsi joints two to four inclusive are quite equal and short, the first is longer than wide, the fifth about equal in length to the preceding three taken together. In the mesotarsi joints two to four inclusive are subequal and short, the first is scarcely as long as the second and third taken together, the fifth is not as long as the combined lengths of the preceding three. In the metatarsi joints two and three are equal and together about equal to the fourth; the first is a little longer than the fourth. Tarsal formula: Pro. Meso. Meta. Metatibie. Male.—2 Zhe a 28 45 Female.—12 21 23 43 Genus TROGLODERUS LeConte. Trogloderus LEContTrE, N. Amer. Entomologist, I, no. 1, July, 1879, p. 2. Body elongate, rough, opaque, eyes slightly oblique; sides of the frons moderately dilated, distinctly more prominent laterally than the eyes; epistoma moderately dilated laterally, scarcely in the least reflexed, margin arcuate laterally and emarginate anteriorly; labrum moderately small, transverse, somewhat short, sinuate, strongly re- tractile beneath the epistoma ; mandibles not at all visible from above; antenne with the third joint as long as the two following combined, eighth, ninth, and tenth distinctly transverse, somewhat perfoliate, eleventh narrower than the tenth; mentum rounded in front, trape- zoido-triangular to parabolic in outline, surface more or less im- pressed laterally; prothorax deeply emarginate in front, rounded at the sides, then briefly and strongly constricted or simply sinuate, apical angles stout and anteriorly prominent, basal angles small and more or less prominent and acute, disc more or less foveate in the 484 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. median line; scutellum very feebly entering between the elytra, the latter not margined at base nor adapted to the prothoracic base, suture and four coste each side acutely elevated; epipleuree narrow, entirely overlapped by the metasternal side pieces and first three abdominal segments, exposed and horizontal opposite the last two segments; prosternum quite strongly protuberant ventrally before the coxe and produced behind: mesosternum deeply concave; metaster- num and first two abdominal segments at middle, broadly and shal- lowly concave; intercoxal process of first abdominal segment broader than long; profemora more or less armed with a tooth at’ outer fourth; anterior tibize more or less compressed and arcuate. The above diagnosis differs in several important points from that given by LeConte or Horn. The laterally dilated epistoma with sides arcuate is in ‘contrast to the condition in the other genera of the present tribe, where the sides of the epistoma are sinuate or straight and convergent ante- riorly. The epistoma in the present genus appears longer on account of the dilatation and less broadly sinuate anterior margin. In most of the specimens before me the labrum is not visible, but in others it is entirely uncovered as in /’Jeodes, and in all probability it is capable of greater retraction than in the other genera of the tribe, where it is retractile to a less degree. The mandibles are not visible from above—except slightly in the sinuation of the labrum—as in LVeodes, Embaphion, and Eleodi- morpha, and this is no doubt due in part to the fact that the mouth parts are not so strongly developed or prognathic as in the above genera. In the Seaurini (Cerenopus and Argoporis) the labrum is entirely invisible from above and apparently deeply concealed beneath the produced epistoma, at least I have never seen it protruded. The scutellum does not enter between the elytra to any extent in costatus, but more so in tuberculatus. The epipleure while narrow, are nearly entirely inflexed above and overlapped by the metasternal side pieces and first three abdominal segments, but they are visible beneath the humeri opposite the meso- sterum, horizontal and exposed near the apex opposite the last two abdominal segments. I have not observed such an extreme condi- tion elsewhere in the Eleodiini, although it is more or less indicated in sporadic instances. All specimens that I have examined are identical in this respect. The habitus of the insect might be considered somewhat discordant in the present tribe, but no more so than in the cases of Lleodes granosa and veterator in that genus. er a een I ee REVISION OF ELEODIINI—BLAISDELL. 485 I do not consider it in harmony with the Seaurini as defined by Casey.*. On account of the short, transverse head, with eyes near the prothorax, it is more closely related to the Eulabini. In many respects it is more closely in harmony with Eleodimorpha than any other heretofore mentioned. The genital characters appear to show greater affinity with the Eleodiini. (renital characters, male—Kdeagophore oblong-ovate and acutely pointed. Apicale triangular, evenly convex above and acute at apex. Sternite bilobed, lobes separate and united by membrane. Female.—Genital segment moderately depressed, subequilaterally triangular, valves more or less reflexed and fully chitinized. The dorsal plate is not at all defined from the apex, surface more or less concave, with scattered sete in apical moiety. The appendage is very small, semi-oval in outline, subapical and feebly penicillate. Superior pudendal membrane moderately exposed and reaching at least to the apical third of the dorsal plate. Ventrolaterally the submarginal groove is very feebly defined, the surface lines are straight longitudinally. Internal margins of the valves contiguous in basal half, fissure narrow in apical half; inferior membrane not visible. It can be seen by comparing figs. 4, 7, 8, 18, and 14 of Plate 6 with those of the different species of /Veodes, and then with figs. 5 and 6 of the same plate, that 7rogloderus is more Eleodiinine than Scaurinine. Hulabis agrees with Cerenopus in having the two lobes of the sternite coalescent into one piece, emarginate at apex, and this is distinctly in accord with what appears as true Scaurinine charac- ters. This investigation is in its infancy and not mature for dis- cussion at the present time. For the present, at least, I agree with Dr. George Horn in re- taming 7rogloderus in the Eleodiini. Future studies might indicate the advisability of separating it, possibly with Lleodimorpha, as a distinct tribe, and it will indeed be interesting to discover some other closely related species that might indicate its genealogical history. General characters.—The profemora are stout and subclavate, oval in transverse section; the tibial grooves are shallow and feebly defined by asperate margins, best defined in the males, but in either sex more or less evanescent toward the base, narrower in costatus, especially in the males. The meso- and metafemora are comparatively slender, slightly and very gradually increasing in width from base to apex, but almost parallel, quite straight, except for the normal curve by which they are adapted to the sides of the body, oval in transverse section; tibial “Annals N. Y. Acad. Sci., V, Nov., 1890, p. 391. 486 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. grooves more or less evanescent. in basal half and feebly defined apically, very shallow but rather smooth. Protibize distinctly compressed, especially along the inner side; external border arcuate in basal half, thence straight and with a tarsal groove, that evenly and gradually widens to the articular cavity with which it is continuous; the anterior margin of the groove is ‘ather smooth, the posterior more or less denticulate; the anterior and posterior sides of the articular cavities are feebly dilated. Le- Conte says that the apical angle is produced, but I do not see that it is in the least so. In the male of costatus the protibie are briefly and very abruptly constricted at base so that a right angle is formed; the tibizv thence to apex quite even in width, as a whole slightly arcuate, ach internal edge very broadly and feebly sinuate. The meso- and metatibie are slender, straight, gradually and feebly increasing in size to apex, where they are somewhat. dilated, circular in transverse section, and more or less grooved or flattened externally for the tarsi; articular cavities closed. The protuberant prosternum is very obvious in costatus, less so in tuberculatus, 1 each it is deeply grooved around the anterior border of the acetabula; in ¢vberculatus ‘the anterior margin is distinctly deflexed. Distribution.—Confined as far as known to the arid regions of eastern California and western Nevada, northward into Idaho. The two known species may be separated as follows: Pronotum cribrately Uncrate eet oe) ee awe SS ee ee ee costatus. Pronotum -tuberculate-—2= 22.22 2 ee oe ee eee tuberculatus. The synopsis of genital characters may be formulated as follows: Superior pudendal membrane long, reaching to the apical fourth Of thie; dorsal plates 2222s 22 a ee eee Stele oe re costatus. Superior pudendal membrane shorter, reaching to apical third__-___ tuberculatus. TROGLODERUS COSTATUS LeConte. Trogloderus costatus, LECONTE, N. Amer. Entomologist, I, No. 1, July, > 1879, p: 3, pl. 1, figs 3. Elongate, oblong-oval, opaque, dirty black, integuments dense; thorax cribrately punctate; elytral coste acutely elevated. Head \ess than twice as wide as long, feebly convex, more or less irregularly impressed, frontal suture not evident, but indicated by the abrupt termination of the frontal impression, at times slightly elevated; coarsely and confluently cribrate, the punctures at times more or less corroded. Antennw short, scarcely reaching beyond the middle of the pro- thorax, somewhat clavate, feebly compressed, second joint nearly one- half as long as the third and not quite as long as wide, third joint about equal to the next two taken together, fourth the least longer “a” es Oe ‘ | | A et oy REVISION OF ELEODIINI—BLAISDELL. 487 than the fifth, the latter, sixth and seventh subequal and each about as long as wide, eighth, ninth, and tenth transverse and gradually increasing in width, eleventh narrower than the tenth, triangulo- ovate and about as long as wide. Pronotum one-third to one-half wider than long, widest at the middle; disc rather strongly convex and moderately declivous at the apical angles, coarsely, more or less confluently and cribrately punc- tate, with two deep oval fovex occupying nearly the whole of the median line, sometimes forming a deep median groove, or frequently entirely absent; apex deeply and broadly emarginate, beaded laterally, obsoletely so at middle three-fifths; s¢des moderately strongly arcuate to basal ninth, thence briefly and strongly sinuate to base, here the sides are divergent or subparallel, scarcely at all sinuate behind the apical angles, marginal bead distinctly serrulate anteriorly, posteriorly obsoletely so; base transverse, scarcely beaded; apical angles large, prominent anteriorly, subacute; basal angles minute, acute, and promi- nent externally or rectangular. Propleure more or less convex, but not prominently so, smooth, more or less granulate on the acetabular convexities and posteriorly, sometimes rugulose. EHlytra elongate oval, less than twice as long as wide, widest at the middle; base not margined nor adapted to the contiguous prothoracic base, which is slightly wider; hwmeri obsolete; sides evenly arcuate and quite evenly and gradually narrowing from the middle to apex, the latter shehtly produced and rather narrowly rounded; disc quite evenly and rather strongly convex from side to side, somewhat quickly and arcuately declivous posteriorly, but shghtly and broadly sinuate before the moderately oblique apex (viewed from the side), the suture and four smooth costz each side acutely elevated, sutural costa feeble and evahescent on apical declivity, the humeral and subhumeral meet- ing anteriorly on humeral region, the subhumeral free and obsolete before the apex, first discal free at both extremities being evanescent on apical declivity, second discal uniting with the humeral before the apex, point of union not tuberculate nor prominent; each in- terval broad and apparently biseriately subfoveolate, the foveole more or less indistinctly marked, more or less feebly transversely coalescent, irregular in the outer two intervals, the minute subsquami- form setz scattered independently of the pits. Epipleuiw narrow, overlapped at middle by the metasternal pieces and first abdominal segment, horizontal in apical two-thirds; surface smooth, obsoletely punctate. Sterna sparsely, coarsely, and more or less moderately punctate ; mesosternum deeply and broadly concave, sides prominent; metaster- num more or less feebly and broadly concave. Parapleure coarsely and more or less obsoletely punctate. 488 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Abdomen horizontal, moderately convex, coarsely and more or less obsoletely punctate; imtercoxal process and first two segments at middle third, distinctly and broadly concave, the concavity extending more or less upon the third segment, where it is rather semilunar in outline, the concavity is lmited laterally by a convex ridge, well marked on first segment and there continuing the internal line of the acetabula, less evident on the second segment. Legs vather short and scabrous; femora punctate, tibiae muricate; anterior tibia more or less curved and shghtly compressed; tibial spurs similar in the sexes. Tarsi rather short; first joint of the ante- rior with a distinct process beneath, plantar groove not well defined; apical margin of each joint somewhat prominent beneath; spinules flavate, short and stout. Male.—Somewhat narrow. Elytra arcuately and shghtly obliquely declivous behind, abdomen strongly concave. Profemora with a small and broad tooth at outer fourth. Protibie distinctly and briefly constricted at base, distinctly angulate at point of constriction. First joint of the protarsi with a distinct and more or less rounded process beneath, which is about a third longer than wide, truncate at tip, where it is very inconspicuously spinulose. Female.—Slightly broadened. Elytra arcuately and rather verti- ‘ally declivous, thence feebly and broadly sinuate before the moder- ately oblique apex. Abdomen less strongly concave. Profemora obsoletely dentate, tooth sometimes minute. Protibie feebly and briefly but not suddenly constricted at base. First joint of the pro- tarsi with a distinct process beneath which is about a fifth longer than wide. Measurements.—Males: Uength, 11-12 mm.; width, 44.5 mm. Female: Length, 12.8 mm.; width, 5 mm. Edeagophore (Plate 6, fig. 7) *oblong- Genital characters, male. 2 ovate, acutely pointed, not arched. Basale oblong-oval, twice as long as wide, evenly convex: sides feebly arcuate. Apicale triangular, slightly elongate; dise evenly convex, without a groove; sides feebly arcuate at basal third, thence slightly and broadly sinuate at tip, the latter acute: base arcuately lobed at middle, laterally feebly sinuate. Sternite transversely trapezoidal. Each lobe triangular and rather long; sides rather gradually and mutually converging to apex, the latter subacute although narrowly rounded; surface more or less evenly convex, glabrous at base and impunctate, apical two-thirds more or less densely punctate and setose, setae quite long, denser and longer about apical margin. Sinus broad, membrane not setose. Female.—Genital segment (Plate 6, fig..13) subequilaterally trian- gular, valves reflexed and fully chitinized, moderately depressed. + q . ee a “ATE ret? REVISION OF ELEODIINI—BLAISDELL. 489 Valvula—Dorsal plate triangulo-oblong and not at all defined from the apex, external margin moderately convergent apically ; surface more or less concave and with scattered punctures externally and apically, each puncture with a moderately short seta; external margin straight to feebly and broadly sinuate, continuous with the subacute margin of the apex, the latter shghtly emarginate exter- nally for the minute fossa; internal margin quite straight; apical margin not in the least evident, apex not at all defined from the rest of the valve, obliquely conforming to the plane of the dorsal plate, more or less rounded at tip, which is subacute and finely setose within. Appendage minute and punctiform, apparently slightly com- pressed, calvous. Superior pudendal membrane long, attaining the base of the apex, finely longitudinally rugulose, narrowly exposed. Basal prominences not evident. Ventrolateral surfaces not longitudinally convex, moderately so laterally; surface very glabrous and not defined from apex; sub- marginal groove very feebly defined for a short distance cephalad to the appendage; apex with few scattered sete. Internal margins of the valves contiguous in basal moiety, fissure narrow in apical half, membrane not visible. Habitat.—Western Nevada and the contiguous eastern California ; Idaho (Rock Creek). Number of specimens studied, 3. Type in the LeConte collection. Ty pe-locality.—Rock Creek, Owyhee County, Idaho. Salient type-characters—Elongate, dirty black, opaque. Head and thorax coarsely and confluently cribrate, the latter with two deep oval fovew occupying nearly the whole of the median line. Elytra elongate oval, with suture and four discoidal cost acutely elevated, intermediate furrows deep and broad, subreticulate (LeConte). Diagnostic characters—The distinctly cribrately punctate prono- tum distinguishes costatus from tuberculatus; in the latter species the elytral coste are asperate and not smooth as in the former. The mentum is somewhat small and more or less trapezoido-trian- gular, foveate laterally, not noticeably strongly sculptured. The prosternum is strongly protuberant ventrally before the cox, arcuately declivous anteriorly and quite horizontal between the acetabula, feebly longitudinally grooved at middle from anterior margin; surface asperately punctate and sparsely setose, sete short ; produced and acute posteriorly. Mesosternum deeply and broadly concave, sides prominent. Metasternum laterally between the coxe as long as the width of a mesotibia at middle. 490 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. The abdominal intercoxal process is slightly transverse, about a third of its width broader than the metasternal salient and equal in length to the post-coxal part of the first ventral segment; second seg- ment is as long as the third and fourth taken together, and nearly three times as long as the fourth. The tarsi are rather short and slender. ml we Parsal formula: Pro. Meso. Meta. Metatibice. Male.— 1} 12 ioe es Female.— 1} 14 13 32 TROGLODERUS TUBERCULATUS, new species. Elongate, oblong-oval in outline, twice as long as wide, integuments dense and opaque, pronotum tuberculate; elytral coste acutely ele- rated and subserrulate, apex bituberculate. Head \ess than twice as wide as long, plane, interocular region with three impressions, two lateral and one median, intervening sur- face convex; epistoma moderately prominent and feebly convex, con- tinuously so with the supra-antennal convexities, frontal suture not evident, but indicated by the rather suddenly convex epistoma; sur- face punctato-tuberculate. .lntenne short, evidently not reaching be- yond the prothorax, feebly compressed and somewhat clavate, third joint about equal to the next two taken together, fourth and fifth about as wide as long, sixth and seventh evidently just the least wider than long, eighth feebly transverse, ninth and tenth distinctly so, eleventh small, narrower, and about as long as wide, orbiculo-ovate. Pronotum widest at the middle, somewhat explanate at the sides, one-half wider than long; disc moderately convex, distinctly arcuately declivous at the apical angles, rather densely tuberculate, tubercles small and rounded, with two deep oval fovex oecupying the whole of the median line and about a third of the width, the fovez limited laterally by raised and rounded sides and separated by a similar transverse convexity, the concavity of each reaching the apical and basal margins, respectively; apea deeply emarginate in circular are (viewed from the front), the emargination appearing angulate laterally when viewed from above, without a bead; sédes evenly, broadly, and rather strongly arcuate, quite suddenly sinuate at basal tenth, thence briefly straight and almost parallel to base, margin subserrulate; dase, transverse, without bead; apical angles subacute, triangular, stout, and anteriorly prominent; basal angles minute, subacute and feebly prominent posteriorly. Propleure moderately convex antero-posteriorly and more or less concave between the somewhat explanate sides of the pronotum and the acetabular convexities, irregularly granulate and more or less rugulose. ote St ee ee -, Rae eh 2. nee ee yz REVISION OF RLEODIINI—BLAISDELL. 491 Elytra oval, about a third longer than wide, widest at the middle ; base not margined nor adapted to the contiguous prothoracic base, not narrower; Auwmeri rounded or feebly subangulate at the meeting of the humeral and subhumeral cost; s¢des evenly and moderately arcuate, apex not noticeably produced when viewed from above, rather broadly rounded; disc rather strongly convex from side to side, slightly depressed on the dorsum, arcuately declivous poste- riorly, but sinuate before the shghtly oblique apex; surface sculptured with a sutural and four coste on each side, each acutely elevated, sutural costa feeble and evidently distinct to the apex, subhumeral, humeral and outer discal costx unite on each elytron near apex in a distinct tubercle, the internal discal costa becoming evanescent on the declivity; each costa is distinctly subserrulate on crest from a series of impressed and yather distantly spaced punctures, each puncture with a minute seta; intervals broad and apparently biseri- ately subfoveolate, the foveole rather distinctly marked and more or less transversely coalescent, giving a subreticulate appearance ; each interval with sparsely placed minute elevated granules, which bear a minute seta projecting from their base behind; sculpturing irregular in the outer intervals. Epipleure narrow and entirely overlapped by the sternal side pieces and first three abdominal segments, exposed apical portion horizontal, also briefly exposed beneath the humeri. Sterna and parapleure more or less scabrous and somewhat tuber- culate. Sterna feebly concave. Abdomen horizontal, moderately convex, first segment distinctly flattened and slightly concave between the coxa, second segment very feebly flattened at middle third; surface more or less scabro- tuberculate. Legs short and seabrous. Femora tuberculately punctate, with a minute tooth at outer fourth on the profemora. Tibie muricato- tuberculate, protibiz somewhat constricted at base. Tarsi short and feebly grooved, first joint of the protarsi moderately produced at tip beneath. Measurements. width, 6 mm. Genital characters, male——Not studied. Female.—Genital segment (Plate 6, fig. 14) triangular, a little longer than wide, fully chitinized, valves shghtly reflexed. Valvula.—Dorsal plate elongate, oblong-triangular; surface gla- brous and slightly longitudinally concave, very sparsely punctate and setose apically, sete short; external border more or less feebly sinu- ate, somewhat prominent toward base; internal margin straight. Apex not defined from the dorsal plate, briefly sinuate externally, forming a fossa, tip rounded, but not broadly so; internally finely setose on the valvular membrane. Male: Unknown. Female: Length, 12 mm.; 492 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Appendage small, flattened, and semi-elliptical, with two or three short setv at tip. Superior pudendal membrane attaining the apical third of the dorsal plate and not rugulose. Basal prominences not evident. Ventrolateral surfaces with surface lines straight when viewed longitudinally, moderately convex transversely; surface glabrous, sparsely punctate apically and feebly concave laterally before the apices, the latter concave beneath. Submarginal groove scarcely evi- dent. Internal margins of the valves contiguous in basal two-thirds, fissure subapical, with margins feebly setose. Membrane not visible. fabitat——California (Los Angeles County). Number of specimens studied, 1. Monotype a female (Cat. No. 12236) in the U. S. National Museum. Ty pe-locality—tLos Angeles County, California; collector, D. W. Coquillett. Diagnostic characters—TVhe tuberculate pronotum, which is some- what dilated at the sides, subserrulate costs of the elytra, outer three coste meeting in a tubercle upon the apex, sufficiently char- acterize this species as distinct. The female genital characters, if constant in a series, will also prove of value, as the superior pudendal membrane is much shorter and not longitudinally rugulose in twberculatus. General observations—The mentum is moderate in size and sub- parabolic in outline, obsoletely foveate laterally, and not noticeably scabrous. The anterior margin of the prosternum is quite strongly deflexed, the surface is gradually and not strongly arcuate between the coxe, grooved only between the same and mucronate behind. The surface is transversely flattened in front of the coxz at middle and very decidedly in contrast to the protuberant condition observed in costatus. The mesosternum is rather narrowly and deeply concave and moderately prominent laterally. The metasternum laterally between the cox is about as long as the width of a mesotibia at middle. The abdominal intercoxal process is subquadrate and about as wide as the metasternal salient, and as long as the post-coxal part of the first ventral segment, quite equal to the length of the second, which is subequal to the third and fourth segments taken together; the fourth is about one-half as long as the third. The tarsi are rather short and somewhat stout, at least stouter than in the female of costatus before me. Tarsal formula : Pro. Meso, Meta. Metatibia. Male.— ue Vie : Female.— 14 14 13 3s ~ > £. 7 }. REVISION OF ELEODIINI—BLAISDELL. 493 SPECIES NOT RECOGNIZED. I have not been able to recognize nor to correlate the following species in the ample material at hand: ELEODES VICINA LeConte. Nigra, nitida, thorace subtiliter punctulato, latitudine haud longiore, lateribus modice rotundatis, postice subangustato, elytris discrete minus subtiliter seria- tum punctatis punctis paucis interjectis, femoribus muticis. Long. .7. Habitat ad flumen Gila. BH. quadricollis aftinis, at elytris punctis majoribus magis discretis distinetius seriatis differt. Mas, subeylindricus, elytris dorso convexis, postice attenuatis. Femina, elytris latioribus, dorso minus convexis, postice vix attenuatis. The type is from Sonora, which is the same as that of Arizona. Mr. Blanchard writes me that the two males in the LeConte collee- tion are more like carbonaria than quadricollis, the side margins of the elytra being stronger and the punctures at the sides more substriate. Colonel Casey, who has no doubt seen the types, writes: ¢ Vicina is distinct from quadricollis in having much finer punctures, which are sparser and much less asperate towards the sides, and also in several other characters, among which may be mentioned the form of the prothorax, widest at anterior third in quadricollis and just before the middle in vicinus, the much longer posterior tarsi in the male of vicinus. Vicinus is peculiar to the Gila Valley of Arizona. A male from M. L. Linell’s collection, bearing a label with vicinus written upon it, evidently in his handwriting, corresponds to Le- Conte’s description, and very closely to two males in my own col- lection from Arizona and without definite locality. Linell was considered a careful student and his label bears some weight. His specimen is quite different from one which Fall ques- tionably refers to vicina. I do not believe vicina to be distinct, and it must be a race or form of carbonaria, or guadricollis near the varieties anthracina and lustrans: the form of the protibial spurs in the female must decide which. ELEODES NITIDUS Casey. The following is Colonel Casey’s description : Rather slender, convex and subeylindrical, black throughout and strongly shining, glabrous. Head rather sparsely punctate, moderately coarsely so toward apex; an- tenn robust, nearly as long as the head and prothorax, the third joint four 7Ann. New York Acad. Sci., V, Nov. 1890, p. 395. 494 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. times as long as wide and fully as long as the next two together, last three joints distinctly dilated, forming a club, the ninth and tenth strongly trans- verse. Prothoraxr quadrate, very nearly as long as wide; dise just visibly wider at apical third than at base, evenly convex, externally, minutely, and sparsely punctate; apex transverse, broadly and feebly bisinuate, equal in width to the base; sides parallel, very feebly arcuate; base broadly and rather strongly arcuate; apical angles right, narrowly rounded; basal .angles broadly obtuse and not prominent nor at all rounded. BHlytra two and three-fourths times as long as the prothorax, and in the middle nearly one-half wider, about twice as long as wide; base broadly emar- ginate; humeri right, not noticeably exposed; sides broadly arcuate; apex acutely rounded; disc obliquely declivous behind, having feebly marked, distant, completely unimpressed series of extremely minute punctures; intervals mi- nutely, sparsely, and more or less confusedly punctate, the punctures not distinctly larger or denser laterally. : Femora all slender and completely unarmed; spurs of the anterior tibize very unequal, the anterior slender, acutely pointed, and moderate in size; posterior tarsi slightly compressed, nearly three-fourths as long as the tibiee. Prosternum slightly prolonged but not at all reflexed, the apex vertical. Length, 18 mm.; width, 6.7 mm. Habitat—Arizona. Casey’s specimen was a unique and he compares it with longicollis. He writes that it differs from that species in its flatter, rather shorter prothorax, which is less rounded on the sides, and in its shorter, broader elytra, broadly and rather strongly emarginate at the base and having widely distant rows of punctures, and in having a shorter, stouter, and much more clavate antenne. It is possible that the species described by me as dissimilis may be nitidus. Dissimilis is very variable. (See p. 398.) ELEODES REFLEXICOLLIS Mannerheim. I have failed to satisfactorily place this species of Mannerheim’s, the type of which was taken at Ross, California. This locality is undoubtedly Fort Ross, near the mouth of the Russian River on the coast. Mannerheim says that it is close to Hleodes planata Eschscholtz, and that the sides of the thorax are unusually reflexed, and the base abruptly narrowed. Peculiar and aberrant forms are occasionally met with. (See scabricula for remarks upon a unique specimen taken in Napa County, California, and in the collection of A. Koebele. ) HARLY STAGES OF THE ELEODIINI. Comparatively little has been written upon the early stages of the Tenebrionide of the United States, and also little upon the tribe, ware ws > ’ > REVISION OF ELEODIINI BLAISDELL. 495 treated in this paper. The only articles accessible are those of Carl F. Gissler*. Some of the figures are so very poor in these papers before me that they are of little use. Mr. Gissler writes that in studying the larval states of the Tene- brionide we find that there is scarcely a coleopterous family whose members are homologically so closely allied and correlatively alike, and are all referable to three typical larval forms: 1. Those having homogeneous corneous segments, cylindrical in longitudinal series, the stigmata situated in the pleurites. Example, that of Tenebrio. 2. Form more compact, with only the prothoracie segments corne- ous and abdominal segments slightly arcuate. The dorsal segments do not overlap, forming a marginal bead, above which (on the tergal part of the segment) the abdominal stigmata are situated. Example, that of Bolitophagus and Bolitotherus. 3. Clypeus not entirely separated, mandibles more strongly dentate, terminal joint of labial palpi large and truncate, pygidium without articulated spines. The larvee of the /Veodiini belong to the first form. They are elon- gately cylindrical, resembling those of the Elateride, which are com- monly called wireworms. The former differ in having the cardo of the maxillze connate and a transverse impression immediately behind the clypeus, thus appar- ently separating the same from the front, and also by the better de- veloped labrum (Gissler). Very young larve have their antennal joints more globular than the older ones. In full-grown larve of the Eleodiini the first joint of the antennx is much shorter than the second, its substance soft, non- chitinized and retractile; its position is in a minute cavity, which is surrounded by a semi-circular ridge (Gissler). William R. Erichson has described this as the first joint in the larva of Tenebrio.® From my dissections I regard this simply as the basal articulation with its membrane. Compare figs. 9 and 10, Plate 13. It is quite evident that the antenne are retractile, but when dissected out it is composed of only three joints. Mr. Gissler obtained eggs of leodes gigantea from specimens kept in captivity. These were discovered in October, and measured in length 0.8 mm.; width, 0.85 mm.; elongate oval and milk white in «Bull. Brooklyn Ent. Soe. for 1878-1879, I, No. 1, p. 11; No. 3, p. 18; No. 11, p. 85d. >See Archiy fiir Naturgeschichte, 1841, p. 365. 496 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. color. In November (20th, 1877) he noticed the first larvee; none appeared after December. Mr. Gissler describes the larva of 'Jeodes gigantea and dentipes as follows: Larva (Lleodes gigantea).—Head convex above, gular region con- cave; all the mouth parts like Zenebrio; antenne nition by the stouter second and third joints. Pygidium with 18 lateral spines, terminal obtuse tip, sparsely frimbriate. Color of small specimens (6 mm.) entirely white. No trace of ocelli or even dark spots on the ophthalmic region. The small larve have the abdominal segments nearly all flat and gradually after several exuviations acquiring the typical wire-shape and darker color. The legs, thoracic and abdominal segments as in Tenebrio. Larva (Lleodes dentipes).—Pygidium with but ten spines, the two terminal are slightly longer and run parallel over the tip. The front legs also differ from those in the larva of gigantea, the head, mouth parts, antenne, middle and posterior legs are as in the latter species. A larva but a few hours old (Plate 13, fig. 15) has two small blunt terminal spines, at the sides of which are two stout, long bristles; there are no lateral spines. The latter appear after the first moult. Length, 2-5 mm. Mr. Gissler also writes that a number of H/eodes dentipes began to deposit their eggs in April, and the larvee grew to 30 and 35 mm. The above descriptions are very unsatisfactory and ac no clear nor complete idea of the structural characters of these larve Mr. Gissler kept his larve in a breeding jar with earth a decayed wood. I have kept in captivity the several species of H/eodes found about San Francisco and succeeded in obtaining the eggs of several species. In general form and color they were all alike, milky white and oval in form, differing only in size and this in a few fractions of a milli- meter; the majority were about a millimeter in length. I examined them carefully under high magnifying power, but the enveloping tunie was alike in all, no sculpturing. The following were exam- ined: Eggs of dentipes, quadricollis, cordata, parvicollis, and scabrosa. I succeeded in obtaining larve from only one species and that was dentipes; they perished during the excitement following the San Francisco disaster. The larve had reached the length of 9.5 and 10 mm. I discovered them in July and I did not know how long they had been dead. I had not discovered them before April 18. 3etween these larvee and those mentioned by Mr. Gissler I find very little resemblance. This probably depends upon a different age of the larve, My study and dissections were made upon two REVISION OF ELEODIINI—BLAISDELL. 497 larve 10 mm. in length (Plate 13, figs. 4-14), supplemented by eight others. The illustrations give the characters better than a labored description. LARVA OF ELEODES DENTIPES. Length, 10 mm.; color, castaneo-testaceous, at times fuscous. Sur- face glabrous. Form elongately cylindrical, flattened ventrally. The ultimate and posterior border of the penultimate segments, both above and beneath, sparsely clothed with long flying hairs; the under surface of the thoracic segments are similarly clothed, the hairs being a little longer and rather denser; the sides of the head beneath are rather more conspicuously pubescent. Similar but shorter hairs are scattered here and there over the dorsal surface of the tergites, apparently without a definite orderly arrangement, ex- cept laterally just a little in advance of the posterior border on each segment there is a longer hair, and these are observed on all the seg- ments, forming a series on each side; on the prothorax there is a hair at each angle. There is also an evident series on the sternites just within the lateral border on each side; four or five hairs were counted; the number varied, possibly from being broken off; the series are definite and continuous on the abdominal sterna. Head (Plate 13, fig. 9) somewhat circular in outline, but truncate at base, transverse without the clypeus and labrum, slightly de- pressed; just behind each antenna are two ophthalmic spots; all are small and transverse, the anterior a little more external than the posterior, separated by about five times their own length; imme- diately in front of the anterior are three or four setigerous punctures. Antenne three-jointed, situated immediately behind the mandibles in articular cavities with rather prominent borders; joints one and two about equal in length and about twice as long as wide, terminal joint very small and bearing a seta at tip; retractile. (See figs. 10 and 9, Plate 13.) | Mandibles short, somewhat irregularly triangular in outline, apex emarginate, superior cusp longer than the inferior; superior surface feebly convex, slightly irregular; external surface arcuate as viewed from above, more strongly so just apicad to the middle when viewed from below; external margin of superior surface with a cluster of some three or four sete at middle, a couple of setee more basad; in- ternal margin with a small triangular and acute dentiform process shghtly basad to the mid-point between apex and molar. Inferior surface irregularly convex, excavated within the cuspi- date apex, distinctly evident as viewed from below. Internal surface with a prominent molar (figs. 13 and 14, Plate 13). Membranous lobes so evident in the adult, obsolete. 59780—B ull. 68—09 32 498 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Maxille vather stout. Cardo and stipes rather distantly sepa- rated from the mentum by a membranous structure. Palpi three- jointed, second joint slightly broader than the first, third distinctly smaller. Mentum subquadrate, sides slightly converging to base, apex sin- uato-truneate, the articulating membrane attaching it to the pal- piger more or less distinct; ligula a delicate median process between the palpi, bearing two sete at tip. Labial palpi short and two-jointed; first jot about as wide as long, possibly a little longer if fully extended; second joint much narrower and smaller. Submentum (figs. 8s and 11s, Plate 13) trapezoidal, base moderately arcuate, apex truncate, sides feebly, broadly but distinctly sinuate. Both the mentum and submentum are sparsely clothed with rather long flying hairs. Gula subquadrate, sides somewhat arcuate basally and feebly sin- uate apically. Suture distinct. Labrum transverse, a little more than twice as wide as long, anterior margin shghtly arcuate, more strongly rounded laterally. There is an apical series of about eight sete, and a basal series apparently the same in number. Clypeus with a feebly transverse impression which shows at the sides as an emargination, just back of which is a setigerous puncture, another more internal, the sete of the two sides being in line; sepa- rated from the frons by a distinct transverse groove which ter- minates at the anterior border of the antennal socket. Prothorax nearly quadrate on the dorsum, slightly transverse, in some specimens transversely and feebly concave. J/eso- and meta- thorax much shorter and together scarcely as long as the prothorax Abdominal segments subequal in length. Spiracles—There are nine pairs. The anterior is situated under the margin of the prothorax in a depression external to the coxa, larger than the others and with the margins more prominent and chitinous. The abdominal are in segments 1-8 inclusive and situated at the anterior margin of the pleurites; they are circular and flat, margins not prominent. Pygidium (Plate 13, fig. 5) triangular, sides moderately arcuate, apex rather broadly rounded. The marginal series of spines con- sists of eight on each side, the terminal two are just noticeably little stouter and longer than the others. The specimen from which fig. 5 was drawn shows three spines dis- placed inward from the proximal extremity of the marginal series; another specimen had them present only on one side and in others they were entirely absent. I was unable to determine if this was the result of wear. REVISION OF ELEODIINI—BLAISDELL. 499 Mr. Gissler notes that in one specimen studied by him there were two spines articulated in one cavity, while in another specimen they were asymmetrically placed, nine on one side and eight on the other. In the sculpturing of the pygidia a few smooth spots were observed in Eleodes. Legs similar in form, the anterior larger and stouter, claws of the middle and posterior are distinctly more slender; the anterior are more strongly chitinized. The coxe are quite conical and transverse, also prominent. The anterior has spines arranged as follows: At inner side of base of claw a spine points inward, tibiz with three spines on inner sur- face, femur with three, trochanter with two, each podomere with scattered flying hairs of varying lengths. The middle and posterior legs also have spines somewhat differ- ently arranged and more difficult of description. Mr. Gissler writes that in Z’enebrio and Hleodes (Plate 13, fig. 15) the first post-embryonic stages are alike in the form of the pygidium. The integumentary appendages of larval pygidia may be regarded as the homologues of the dorsal parapodia of their ancestors—the Annelides. Two very minute tubercles can be seen under strong microscopic power at the middle of the base of the pygidial tips in Lleodes. Whether they are orifices or not I can not decide. The larve of /e- odes when disturbed and handled violently jerk their abdomen and spurt a turbid, odorless, watery fluid out of two lateral orifices situ- ated under the tergal fold near the posterior margin of the seventh abdominal segment. These structures are a peculiar adaption to the larval body, since the excretion of the repugnatorial glands of the imago is spurted out through the anal opening. The higher develop- ment of this gland is certainly attained during the pupal period (Giss- ler). Mr. Gissler found that in Blaps mortisaga the pygidium had usually twelve lateral and two latero-terminal spines (Plate 13, fig. 18). Out of ten specimens two had sixteen in all, the rest only fourteen. In Eleodes gigantea and E. dentipes he gives as the usual number of spines sixteen lateral and two latero-terminal. A number of either species had only sixteen in all. (See Plate 13, fig. 17.) The larve of /Jeodes are abundant, but I have never had time to successfully rear them, to determine the species to which they be- longed. From the sand dunes along the coast the larvee of L’. clavi- cornis and scabrosa have been taken. Two pup were obtained; one developed a clavicornis, the other being identical was studied, figured, and preserved in spirits, 500 BULLETIN 638, UNITED STATES NATIONAL MUSEUM. PUPA OF ELEODES CLAVICORNIS. (Plate 13, figs. 1, 2,"and. 3.) Length, 10.5 mm.; width of body, 4.5 mm.; greatest width from knee to knee, 7 mm. Oblong-oval, moderately arcuate, head flexed against the prosternum ; abdomen moderately depressed ; color yellow- ish white, appendages semitranslucent. Head exposed, vertex visible from above, clypeal region convex, frons somewhat transversely impressed, vertex rather more promi- nent; antenne curving backward against the sides of the prothorax, over the profemora, at which point they are not visible when viewed from below. Pronotum evenly convex; base almost truncate, basal angles rather broadly rounded, sides arcuate, apex feebly and broadly emarginate, with angles somewhat narrowly rounded. Mesonotum short, transverse, and moderately convex; elytral pads passing obliquely backward and ventrally above the pro- and meso- crura, apical fourth between the meso- and metacrura, visible ven- trally only at this point. -~Epipleural margins apically contiguous to basal half of the mesotarsi. Metanotum a little longer than the mesonotum, moderately convex, and broadly sinuate behind. Legs prominent laterally, not appressed against body, distinctly compressed and comparatively broad; tarsi distant from each other in median line, except the metatarsi, which are in contact in apical half; coxee and sterna visible in the median line. Abdominal segments convex dorsally, less so ventrally; pleural region of segments one to seven produced laterally into subquadrate laminiform processes; last two segments without lateral processes and more evenly convex from side to side. Fourth ventral segment deeply and broadly emarginate at apex; fifth smaller, rather arcuate at apex and within the emargination of the fourth; third segment broadly and less deeply emarginate. Abdomen at apex terminating in two elongate processes, each grad- ually tapering from base to apex, the latter chitinous; beneath the bases of these cerci there is a small segment which is deeply emar- ginate, with angles prominent posteriorly. The lateral processes of segments one to seven are limited at base dorsally by a longitudinal impression on either side of the dorsum ; the impression begins on segment one and ends on base of the seventh ; ventrally the impressions are less strongly marked. In the specimen at hand the processes of segments two to six are deeply and semicircularly emarginate at middle third, each anterior and posterior third is squarely truncate at apex, with angles sub- acute, those at the emargination are chitinous and denticulate, each RA ee ee ae OR eek feet TS teed PO SS eee Sit rye ar ae oe Saree With Pet, p ek aa ie, Wee ere eee REVISION OF ELEODIINI—BLAISDELL. 5O1 dentate angle has a small seta at middle of its side toward the emargination; contiguous sides of the processes of each adjoining segment narrowly chitinous, very minutely and irregularly denticu- late; posterior third of the process of first segment not developed; processes of seventh segment imperfectly developed. Anal cerei with a few scattered sete. Pronotum with a lateral marginal row of comparatively closely and evenly spaced chitinous sete, internal to which is a submarginal series of distantly spaced sete, often broken off and appearing as minute chitinous tubercles; there is also a post-apical series of very distantly spaced and similar setee, as well as an antebasal series simi- larly spaced. In Plate 13, fig. 2, it will be seen that the post-apical series appear as marginal when viewed from above. On the dorsal surface of the head there are four to six sete; dor- sal surface of abdominal segments three, four, five, and six with a seta each side near basal and external angle. Legs with a short seta on outer anterior femoral region near the knee: fourth ventral ab- dominal segment with a subapical row of sete. SPribel Bie ASP aaaeNer. In (Blaps) the globular outer antennal joints, wide epipleure which terminate upon the external surface of the cauda when present, the intercoxal tubercle of the first abdominal segment and the pubes- cent pit at base of the second abdominal segment the present tribe stands in marked contrast to anything observed in the genus Jeodes, and in fact all of the genera of the Hleodiini, except in the wide epipleure which forms the whole inflexed sides of the elytra in Hleodimorpha. The genital characters are certainly very distinct and more primi- tive than is to be seen anywhere in the Hleodzini. Genital characters, male-—tLobes of the sternite strongly connate, clave connate, lying dorsad to and forming a pallium for the edeagus, more obsolete ale, more embryonic condition of the edeagophore, there being a complete solution of continuity in the chitinization along the median line of both the basale and apicale, these segments being longitudinally subdivided by an evident membranous line into lateral sclerites. Female.—Valves of the genital segment (Plate 5, figs. 16 and 17) completely chitinized, apex of each undifferentiated, appendage ab- sent. The facies of the segment is quite different from that observed in the Hleodiini, but a phylogenetic relationship is evident. I am not desirous to repeat what may already be known in Europe, but simply to indicate my reasons for not retaining the genus Blaps in the same tribe with the others dealt with in this paper. 502 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. It is only by the assiduous working of data and in the making of deductions in accordance with the laws governing ontogenesis and variation that our knowledge will be complete or as full as scientific progress will permit. The student must bear in mind that the characters of Blaps given have been drawn up from a very limited number of species, and that it is to be expected that a genus so rich in species will present varia- tions analogous to those observed in /leodes. I wish to point out that the genus is undoubtedly a divergent stem from the ancestral phylogenetic trunk from which /7eodes also had its origin. A de- tailed study along the lines which IT have laid down might be the means of greatly modifying our view of relationships as at present accepted. Genus BLAPS, Fabricius: This well-known genus of the Old World needs no treatment at my hands and is given a place here on account of two species having been introduced into the United States and therefore are to be con- sidered as a part of our fauna. In many characters it resembles “7eodes and also presents several very distinct and divergent characters. Mentum transverse, plane and more or less variable in form, some- times trapeziform, sometimes rounded at the sides and in front, at times subtruncate, rarely sinuate, always narrowing at base for a short distance. The lateral lobes are small and invisible. Mawillary palpi elongate, last joint securiform, longer than wide. Labrum prominent, transverse, rectangular or subcordiform, angles rounded and more or less sinuate at middle of the anterior border. Head prominent, not deeply inserted, frontal suture distinct or obsolete; epistoma with sides more or less straight and convergent anteriorly, apex truncate or feebly sinuate. Hyes transverse, narrow, subreniform or sublunate. Antenne moderate, 11-jointed, third joint long, 4 to 7 of variable length, obconical, 8 to 10 globular, 11th oval, pointed at tip. Prothorax variable; sometimes almost plane and rectangular, at others more or less convex and rounded at the sides, the latter always margined. Scutellum of variable size, sometimes not visible. Elytra at base not as wide or wider than the contiguous protho- racic base; form and length variable, frequently attenuated poste- riorly and caudate at apex. Epipleure attaining the humeral angle where the superior margins are visible from above, gradually narrowing to elytral apex. Legs generally long, especially the posterior; anterior femora more or less thickened; tarsi moderate, spinous beneath, plantar grooves distinct. Prosternum mucronate or not. REVISION OF ELEODIINI—BLAISDELL. 503 Male characters.—Narrower, finer punctuation, greater length of elytral cauda when present. Abdomen more or less moderately convex, first abdominal suture more or less impressed with base of second segment and apical moiety of the first, giving the abdomen the appearance of being oblique and the sterna somewhat promi- nent; first suture with a fossa at middle, densely clothed with golden pubescence on the segmental margins; sometimes a truncated tubercle on the intercoxal process of the first abdominal segment. Female characters —Broader form, somewhat coarser punctuation. Abdomen rather strongly convex. Genital characters—The characters observed in the specimens at hand are as follows: Male.—Edeagophore, length 5.5 mm.; edeagus 1.5 mm. Very elongate, somewhat flattened, strongly arched, somewhat oblong- ovate and well chitinized. Basale, elongate oblong, about four times as long as wide, mod- erately convex from side to side, divided by a median membranous groove which extends from base to apex, apically continuous with the groove of the apicale. Apicale elongately triangular, at least twice as long as wide, some- what depressed, dise feebly convex, narrowly and more strongly so laterally, a long median groove extending from base to apex: base quite broadly and strongly lobed at middle three-fifths, laterally broadly and feebly sinuate; sides slightly and evenly arcuate, regu- larly converging to the subacute apex, the latter rendered minutely emarginate by the median groove. Sternite transversely suboblong, lobes strongly connate and con- tinuously chitinized, transversely prominent at middle, the surface sloping apically and basally; apical moiety broadly concave; basal half less broadly concave at about middle third, laterally the surface is convex and gradually diminishes toward the lateral angles of the apex; apical margin» broadly and evenly emarginate; external borders moderately arcuate; apical angles somewhat prominent and rather narrowly rounded. Apical moiety densely pubescent, hairs of moderate length and rather coarse, basal portion glabrous. Clave absolutely not evident as distinct structures. They are present but connate forming a dorsal pallium for the edeagus. The margins of the pallium gradually attain the midline ventrad to the base of the edeagus, becoming contiguous but not connate; beyond the middle of the edeagus the pallium is entirely dorsal, with sides distinctly chitinous and semimembranous at middle. The apex of the pallium projects just a little beyond the caput edeagi. Ale strongly inflex and somewhat obsolete. 504 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Female.—Genital segment (Plate 5, figs. 16 and 17) triangular, elon- gate, general dorsal surface plane and glabrous, appendages absent: Valvula fully chitinized, apex not differentiated from the dorsal plate, surface slightly irregular, at middle minutely and ocellately punctured, a minute seta arising from each puncture, apex slightly deflexed and subacute, very feebly convex above; external border broadly and feebly sinuate at middle, arcuate basally and apically; internal margins rather straight in apical two-thirds, arcuate and converging from base where they are moderately distant to become contiguous at middle third. Superior pudendal membrane long, reaching to the base of the apical region, very finely longitudinally rugulose. Ventrolateral surfaces convex basally, less so towards apex, mod- erately concave beneath the slightly deflexed apices; internal margins of the valves contiguous basally and apically, genital fissure fusiform and just basad to the middle: submarginal groove obsolete, some- what evident beneath the apical margin. Dorsal plate not at all explanate. Apical two-thirds very sparsely and finely pubescent. The following two species can be credited to our fauna: Blaps similis Latreille. Blaps mucronata Latreille. T shall not attempt a specific diagnosis. Doctor George Horn com- pared native specimens with the European and dispelled any previous doubts about their correct identification. [abitat.—A few specimens of mucronata occurred near Baltimore, Maryland. S/milis occurs abundantly at Alexandria, Virginia. I have had the privilege of studying a series in the collection of the U.S. National Museum, said to have come from West Virginia (Riley Coll.) ; figs. 16 and 17, Plate 5, were taken from a broad female from that series. A small series was kindly loaned me by Mr. Charles Waterhouse, of the British Museum, including what is supposedly correctly identified specimens of similis, mortisaga, and mucronata. 3esides these, IT examined a series rather recently obtained from London (British Museum), England, by Miss Juha Wright. The specimens in our American collections do not seem to be defi- nitely labeled. Mr. Liebeck, of Philadelphia, writes me that in his collection is a specimen from Alexandria, Virginia, labeled as gibba; one from Boston, Massachusetts, taken in a warehouse, identical with a specimen (European) in the Horn collection of similis; two labeled mucronata, supposedly from Canada. In the Horn collection (Academy of Natural Science, in Phila- delphia) there are three forms—one a broad form taken by Doctor Castle at Alexandria, Virginia, and labeled gibba,; two forms have @See Can: lnt., XVI, p. 3%, pl: xxi: ERDARS 7 ue RS en rt REVISION OF ELEODIINI—BLAISDELL. ~ 505 the label sémné/is—one a broad form, single specimen from Europe, and the other an elongate slender form, of which some are labeled ce Md.” A study of the genitalia of Blaps and their careful comparison with typical forms of the genitalia of Hleodes, Embaphion, and Eleodimorpha leaves no doubt in my mind that they are not in any way directly related and that it is absolutely necessary to erect a new tribe (HVeodiini) to receive our species. No species of the Blap- tint (Blaptida), except Blaps, has yet been discovered in the United States fauna. With Blaps eliminated from the Hleodiini the tribe becomes com- posed of homogeneous elements, with possibly the exception of Trogloderus, as T could not study the genitalia in all points of detail for scarcity of material. APPENDIX. In May, 1908, there appeared in the Bulletin of the Department of Geology, Vol. 5, No. 12, of the California University in Berkeley, a paper on the Quaternary Myriopods and Insects of California, by Fordyce Grinnell, jr. In this paper the author recorded seven species of /’leodes, three being described as new, all having been taken from asphalt beds at Rosemary, near Los Angeles, California. They are supposedly the fossil remains of recent or older species. These are the fossils referred to on page 29 of this monograph. I had the pleasure of identifying the species for Mr. Grinnell. The following is a list of the species: EHleodes acuticauda WeConte, also the punctate form; LE /codes acuticauda var. laticollis, Hleodes consobrina LeConte. The following were described as new: Hleodes (subgenus Eleodes) elongata Grinnell; Hleodes behrit Grinnell and intermedia Grinnell. The last two belong to the subgenus Blapylis. In order to more fully correlate these new forms with the more recent forms considered in the present monograph, I carefully reexamined the types and record the following facts: All of the specimens are very brittle, deep black in color, and shin- ing; they appear as if the asphalt had exerted some chemical effect upon them that to a certain extent had modified the structural details. ; These specimens are interesting and present some points of struc- ture and seulpturing which can not be correctly described until a larger or more perfect series shall have been collected. The heads, pronota, and appendages are missing from all the speci- mens, and the elytra alone can not tell the whole story. 506 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. ELEODES ELONGATA Grinnell. The type is a right elytron, fully exposed except the epipleural margin, which is sunken into the asphalt. The humerus and epipleura immediately beneath are free. The specimen is too poor and fragile to give a clear idea of the sculpturing. The elytron is in form like that of “'Jeodes grandicollis ; the punctu- ation is similar, very sparse and small, scarcely at all subasperate, and quite sharply defined; general surface glabrous and shining. Humerus rounded, and the superior epipleural margin is thin and sharp. Length, 21.5 mm.; width, 9 mm. Type-specimen.—Number 10025, University California Collection of Invertebrate Paleontology. This species really appears to be a form of grandicollis, it differs but slightly from some of those of to-day. Grandicollis is so distinct that an elytron or thorax ought to be readily recognized. ELEODES BEHRII Grinnell. One elytron (left), apical sixth buried in the asphalt, dorsal sur- face and epipleura exposed, and in a fairly good state of preservation. The elytral dase is truncate, humerus rounded and not in the least prominent ; séde evenly arcuate; dish; moderately convex, broadly and evenly arcuately declivous laterally, not striate, punctuation diffuse, rather dense, subasperate, and with a tendency to coalesce in twos and threes, the intervals more or less convex and forming slight trans- verse rugule. E’pipleura rather broad and concave, superior margin moderately prominent and rather sharp, gently curving upward anteriorly to the humerus, thence caudad more broadly and arcuately curving downward to apex; surface smooth, sparsely and distinctly pune- tate, punctures moderately small; there is also a trace of transverse rugule. This species is without a doubt closely related to consgbrina or parvicollis,; the elytral sculpturing is more like that observed in the latter. The epipleural characters are unlike anything known to me; the marked coneavity is rarely observed in sporadic instances in individ- uals of to-day; the curve of the superior margin is more strongly downward to apex than in any species. It is possible that these well- marked characters are the result of warping in the asphalt. This species is the only one of the three under consideration that probably deserves a name. It is surely a true leodes (Blapylis) and not at all worthy of a generic standing, as the author suggest- ively wrote. ES Sea se eee eee REVISION OF ELEODIINI—BLAISDELL. 507 Length, 10 mm.; width, 4.5 mm.; greatest width of epipleura, 1.5 mm. Type-specimen.—Number 10023, University California Collection of Invertebrate Paleontology. The type, an elytron which is “narrowly oval” with its fellow of the opposite side would give a more broadly oval figure, like that observed in a small parvicollis, the dorsal surface being somewhat flattened. ELEODES INTERMEDIA Grinnell. A complete abdomen, meso- and metathorax, with elytra, consti- tutes the type; the specimen is quite strongly flattened. The form of the body is broadly oval and truncate anteriorly. Elytra truncate at base; humeri rounded and not prominent; sides evenly arcuate; apex not narrowly rounded; disk evidently flattened on the dorsum, and moderately convex, more strongly and arcuately rounded: laterally; surface quite densely and submuricately punc- tate, punctures coalescing more or less, the interstices convex, forming slight transverse rugule; sculpturing denser and coarser laterally and on apex, not in the least striate; surface evidently evenly and moderately arcuately declivous posteriorly. EL pipleure rather broad at base, gradually narrowing to apex; sur- face distinctly concave throughout, finely, subasperately, and sparsely punctate; superior margin strong, quite sharp and prominent, not sinuate beneath the humeri, but straight in the anterior moiety, and thence quite evenly and gradually arcuately descending to apex as viewed longitudinally. Mesosternum evidently feebly arcuately declivous, deeply concave at middle between the prominent and rounded acetabular borders; very densely, rather finely and confluently punctate. Metasternum and episterna densely and rather finely punctate. The metasternum laterally between the cox apparently about two-thirds as long as the post-coxal part of the first abdominal segment. Abdomen densely and moderately coarsely punctate. First segment at middle evidently as long as the second and third segments com- bined; intercoxal salient quadrate and distinctly transverse, surface concave and evidently with glabrous areas, equal in length to the post- coxal part of the same segment; second and third segments evidently equal in length and each subequal to the length of the post-coxal part of the first. Length, 12 mm.; width of elytra, 8.5 mm. Ty pe-specimen.—Number 10026, University California collection of Invertebrate Paleontology. This specimen is related to parvicollis, and I believe specifically identical with behrii,; the latter is smaller and probably a male, while intermedia is the female. 508 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. There is scarcely any difference at all except in size and the epi- pleural curve. The age of the asphalt beds at Rosemary, near Los Angeles, is Quaternary. The remains of the saber-toothed tiger were also taken from this deposit. The remains of the Coleoptera were found in the neighborhood of the bones. My colleague, Dr. E. C. Van Dyke, who has devoted time to the study of the origin of the Californian entomological fauna, consid- ers that the above data substantiate the statements that have been made from time to time that a great deal of the so-called Upper Sonoran or Californian fauna found its way into this State, from the south, at a very early period, probably during early Tertiary times. In this list are to be included most of the typical genera and sub- genera of the most southern parts of the State. Already at the time at which these deposits were made, most of .the species had assumed the appearance that we see in their descendants, the forms living at the present day. a ae ECM AE RR Oh Or ae eee ee > ret Me ee ee A le a ee See IXPLANATION OF PLATES. PLATE 1. Fig. 1. Dorsal view of the right valve of the genital segment of Hleodes nigrina; showing the chitinous, moderately produced apex, evenly and broadly rounded at tip, with the valvular membrane rather largely exposed within and caudad to the superior pudendal mem- brane. 2. Ventral view of the left valve of Hleodes nigrina; showing submar- ginal groove, concave ventral surface of apex and part of the genital fissure. F 3. Dorsal view of the apicale of the edeagophore of Hleodes nigrina; show- ing the depressed and broadly membrano-chitinous area at middle. 4. Dorsal view of the genital segment of Hleodes gigantea; showing the fully exposed superior pudendal membrane, falciform dorsal plates, chitinous and everted apices, and minute appendages. 5. Ventral view of the left valve of the genital segment of Hleodes gi- gantea; showing the broad submarginal groove continuous with the concavity of apex, straight internal margin of the valve, and absence of the genital fissure. 6. Dorsal view of the apicale of the edeagophore of Hleodes gigantea; showing the broadly and strongly depressed membranous surface and slightly recurved apex. 7. Dorsal view of the left valve of the genital segment of Hleodes longicol- lis; showing its resemblance to that of gigantea, but more strongly punctate. 8. Dorsal view of the right valve of the genital segment of Hleodes grandi- collis ; note the truncate apical margin of the dorsal plate, rectangular angle, truncate appendage, and short pudendal membrane. 9. Dorsal view of the right valve of the genital segment of Hleodes tene- brosa; note the plane dorsal plate, long, slender, subacute, and mem- branous apex; compare with figs. 1 and 2. 10. Ventral view of the left valve of the genital segment of Hleodes tene- brosa; showing evenly convex surface and subapical fissure. 11. Dorsal view of the right valve of the genital segment of Hleodes pedi- noides, typical form; note the explanate and plane dorsal plate which is evanescent apically into the well-developed external apical lobe, also the small rudimentary internal lobe and long pudendal membrane. 12. Dorsal view of the right valve of the genital segment of Hleodes ex- tricata, typical form; note the oval-oblong dorsal plate, acute and small apex, large and flattened appendage. 13. Dorsal view of the right valve of the genital segment of Hleodes ob- scura, typical variety; note the dorsal position of the submarginal groove, dorsal plate apparently contracted with sides arcuately re- flexed, surface deeply concave, basal prominences strongly developed. 14. Dorsal view of the right valve of the genital segment of Hleodes sutura- lis; compare with figs. 28, 13, 15, 16, and 17. 510 Fig. 15. 16. 17. 18. 19. Bagey al Ol 6. 10. Lele BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Dorsal view of the right valve of Hleodes dispersa; compare with figs. 18, 16, 14, and 23. Dorsal view of the right valve of HWleodes sulcipennis; compare with hes 13) 15; 14 ands23: Dorsal view of the right valve of Hleodes acuta; compare with figs. 14 and 28. Dorsal view of the right valve of Hleodes armata; note the nearly plane dorsal plate, reflexed internally, the flattened subconical ap- pendage, and that the basal prominence is not well developed. Lateral view of the apicale of the edeagophore of Hleodes terana; note the moderately protruded edeagus between the apex above and the abducted claya beneath. Dorsal view of the right valve of the genital segment of Hleodes mili- taris; note that the dorsal plate is concave with sides reflexed, sub- marginal groove somewhat visible from above, appendage large and flattened, the basal prominences evident. . Dorsal view of the right valve of Hleodes luce; compare with figs. 18, 20, and 22. Dorsal view of the right valve of Hleodes luce, forma inflata ; compare with figs. 20, 21, 14, 17, and 23. Dorsal view of the right valve of Hleodes texana; note the resemblance between figs. 22, and 21 as compared with 13, 14, 15, 16, and 17. PLATE 2. Dorsal view of the edeagophore of Eleodes omissa; note the membra- nous groove of the apicale, the fully abducted clave and the extruded edeagus. Sternite (male) of Hleodes omissa, forma communis, exhibiting varia- tion from the typical form; note the sparsely setose interlobar mem- brane; a, the produced internal basal angle of the triangular chitinous plate of the sternite (see p. 18). Dorsal view of the genital segment of Hleodes omissa, common form; compare with figs. 6 and 9 of this plate and fig. 11, Plate 1. Ventral view of the genital segment of Hleodes omissa, common form; note the surface contour, the direction of the appendages and the broad fusiform genital fissure, the visible inferior pudendal mem- brane. Sternite (male) of Eleodes quadricollis, forma tarsalis; compare with fig. 2: Dorsal view of the genital segment of Hleodes quadricollis, forma tar- salis; note the differences about apices of the valves. Ventral view of the genital segment of Hleodes quadricollis, forma tar- salis ; compare with figs. 4 and 10. Dorsal view of the edeagophore of Hleodes quadricollis ; compare with ne 1: Dorsal view of the genital segment of Hleodes pygmea; valves slightly abducted: note the resemblance to fig. 3. Ventral view of the genital segment of Hleodes pygmea; valves moder- ately abducted. Sternite (male) of Hleodes marginata; note the rather rounded lobes and long sete; compare with fig. 2; Plate 8, fig. 2, and Plate 4, fig. 15. Dorsal view of the genital segment of Hleodes marginata; note the nearly fully chitinized valves, the produced and everted apices, and punctiform appendages. Compare with fig. 16, Plate 5. b xs ee Caer Fig. 15. on ~ LO? ile 12. 9 vo. 14, fh I REVISION OF ELEODIINI—BLAISDELL. Sieh Ventral view of the genital segment of Hleodes marginata; note the contiguous internal margins of the valves and compare with fig. 16, Plate 5. PLATE 3. Dorsal view of the edeagophore of Hleodes obsoleta; see explanation of fig. 1, Plate 2. Sternite (male) of EHleodes obsoleta; compare with figs. 2 and 5 of Plate 2. : Dorsal view of the genital segment of Hleodes obsoleta; note the very explanate dorsal plates, especially at apex; compare with figs. 3, 9, and 6 of Plate 2. Ventral view of the genital segment of Hleodes obsoleta; compare with figs. 4, 10, and 7, Plate 2; a, paranal plate (see p. 19). Dorsal view of the right valve of Hleodes letcheri; note resemblance to fig. 12, Plate 1. Ventral view of the left valve of EHleodes letcheri. Dorsal view of the right valve of Eleodes acuticauda; compare with figs. 12, 18, and 9 of this plate, and figs. 8 and 18 of Plate 1. Ventral view of the left valve of Hleodes acuticauda; note the large appendage and the concave surface before the apex. Dorsal view of the right valve of Eleodes dentipes; compare with figs. 11 and 13 (type); note the variability of the apical margin of the dorsal plate and form of the appendage. Ventral view of the left valve of Hleodes dentipes. A Dorsal view of the right valve of Eleodes dentipes, showing variation. Dorsal view of the valve of Hleodes laticollis ; compare with fig. T. Dorsal view of the right valve of Mleodes dentipes, forma confinis (type) ; compare with figs. 7, 9, -and 12. Dorsal view of the right valve of Hleodes porcata; note the broadly rounded external apical lobe; compare with fig. 8; the explanate mar- gin of the dorsal plate is less diaphanous, so that the body of valve does not show through it. Dorsal view of the right valve of Eleodes obsoleta; variety showing the obsolete explanate margin toward base. Dorsal view of the right valve of Hleodes carbonaria; compare with figs. 3, 14, 17, 18, and 20; also figs. 3, 6, and 9, Plate 2. Dorsal view of the right valve of Hleodes humeralis; see for com- parison fig. 16. Dorsal view of the right valve of Hleodes cuneaticollis; see for com- parison fig. 16; note the more membranous external apical lobe. Dorsal view of the right valve of Hleodes hispilabris; compare with figs. 7, 8, 19, and 23 of Plate 4. Dorsal view of the right valve of Hleodes tricostata; note the distinctly oblong dorsal plate and compare with fig. 11, Plate 1. PLATE 4. Dorsal view of the right valve of Hleodes scabricula; compare with fig. 12, Plate 2; the notch defining the apex from the dorsal plate appears to be constant. Dorsal view of the right valve of Hleodes neotome; note the subapical position of the appendage and compare figs. 9, 10, and 16. 512 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Fig. 8. Dorsal view of the right valve of Hleodes (Promus) fusiformis; note the broad and setose dorsal plate as compared to the species of the other subgenera; compare figs. 4, 5, and 22; also figs. 2 and 8, Plate 5. 4. Dorsal view of the right valve of Eleodes opaca; note the less setose condition. See for comparison fig. 8. 5. Dorsal view of the right valve of Hleodes goryi. See for comparison figs. 3 and 4. 6. Dorsal view of the right valve of the genital segment of Hleodes cor- data; note the oblong dorsal plate with apical angle evident; com- pare with figs. 9, 10, 11, 12, and 16, also observe the shortness of the superior pudendal membrane as compared to figs. 9, 10, 11, 12, and 16. 7. Dorsal view of the right valve of EHleodes caudifera; compare with igs. 8 and 23. 8. Dorsal view of the right valve of Hleodes sponsa. | 9. Dorsal view of the right valve of Hleodes consobrina; compare with ig. 10, which is a little less swollen at apex, but the same. 10. Dorsal view of the right valve of Hleodes consobrina; compare with fig. 9, which is more swollen at apex, but the same. 11. Dorsal view of the right valve of Hleodes clavicornis; note the elon- gate form and narrow dorsal plate ayd small membranous apex; compare with fig. 22, Plate 5. 12. Dorsal view of the right valve of Hleodes snowii; compare with fig. 9, Plate 1, and fig. 21, Plate 5. 13. Dorsal view of the right valve of Hleodes granosa; compare with figs. 14 and 28, noting the short superior pudendal membrane. 14. Dorsal view of the right valve of Hleodes pilosa; compare with figs. 25 and 28; also with fig. 12, Plate 1; fig. 5, Plate 3. 15. Sternite (male) of Hleodes sulcipennis; compare with fig. 5, Plate 2; also fig. 2, Plate 8, and fig. 11, Plate 2. Note the circular membranous area at base between the lobes, the latter densely pubescent. 16. Dorsal view of the right valves of Elcodes tibialis; note the elongate form, short appendage, and apex; compare with figs. 2, 10, and 11. 17. Dorsal view of the right valve of Hleodes veterator; note the undiffer- entiated apex from dorsal plate, and absence of the appendage. 18. Dorsal view of the right valve of Hleodes femorata; compare with figs. 17 and 20, Plate 1. 19. Dorsal view of the right valve of Eleodes gracilis; compare with fig. 23° also fics. 9 and i Plate. 3: 20. Hleodes innocens.—Dorsal view of the apicale of the edeagophore; note resemblance to fig. 6, Plate 1. 21. Hleodes innocens (female).—Dorsal view of the genital segment; com- pare with figs. 4 and 7, Plate 1. 22. Eleodes insularis (female).—Dorsal view of the right valve; com- > \ pare with figs. 3, 4, and 5, Plate 4. 25. Eleodes longipilosa (female).—Dorsal view of the right valve; com- pare with figs. 7 and 8, Plate 4. 24, Hleodes knausii (female).—Dorsal view of the right valve; compare with figs. 16 and 3, Plate 3. 25. Hleodes arcuata (female).—Dorsal view of the right valve; compare with fig. 12, Plate 1. 26. Hleodes debilis (female).—Dorsal view of the right valve; note the less rounded and more strongly chitinized dorsal plate and short membrane; compare with fig. 16, Plate 8. These characters were present in all specimens examined. Wig. 27 28. ele REVISION OF ELEODIINI—BLAISDELL. 518 . Eleodes ampla (female).—Dorsal view of the right valve; compare with fig. 16, Plate 3. : BHleodes granulata (female).—Dorsal view of the right valve; compare with fig. 12, Plate 1, and fig. 5, Plate 3. PEATEs . Bleodes ventricosa (female).—Dorsal view of the right valve; note the oblique apical margin and prominent angle; compare with figs. 8, 21, and 22 of Plate 1; figs. 12 and 13 of Plate 3. Hleodes striolata (female).—Dorsal view of the right valve; compare with fig. 3; also, figs. 3, 4, 5, and 22 of Plate 4. Elecodes subnitens (female).—Dorsal view of the right valve; com- parisons the same as for fig. 2. Eleodes parvicollis (female; typical).—Dorsal view of the right valve compare with fig. 22; also, figs. 2, 6, and 10, Plate 4. Hleodes pimelioides (female).—Dorsal view of the right valve; compare with fig. 6, Plate 4. Eleodes planipennis (female).—Dorsal view of the genital segment: note resemblance to figs. 7, 8, 9, 14, and 20; also, compare with figs. 6 and 10, Plate 4. Embaphion muricatum (female).—Dorsal view of the genital segment; comparisons are to be made with figs. 6, 8, 9, 14, and 20. Embaphion contusum (female).—Dorsal view and comparisons the same as for fig. 7. Embaphion plunum (female).—Dorsal view and comparisons the same as for fig. 7. Eleodes schwarzii (female).—Dorsal view of the genital segment; com- pare with figs. 11 and 12, noting that the valvular apices are quite different, but fully chitinized; compare also with fig. 1, Plate 1. Eleodes nevadensis (female).—Dorsal view of the right valve; compare with fig. 12; and also fig. 1, Plate 1; note the produced parallel apices with tips broadly rounded, convex above and concave beneath. Eleodes dissimilis (female).—Dorsal view of the right valve. Eleodes inculta (female).—Dorsal view: of the genital segment; com- pare with figs. 9 and 10, Plate 4. Embaphion glabrum (female).—Dorsal view of the right valve; com- pare with figs. 6, 7, 8, 9, and 20. Eleodes hornii (female).—Dorsal view of the genital segment: note the tumid apices and smooth membrane; compare with fig. 9, Plate 4 Blaps similis (female; West Virginia).—Dorsal view of the genital segment; note the fully chitinized and coalesced condition of the parts; compare with fig. 12, Plate 2. Blaps similis (female; West Virginia).—Ventral view of the genital segment; note the position of the genital fissure and compare with fig. 18, Plate 2. ; Hleodes subpinguis (female).—Dorsal view of the right valve; compare with fig. 19, Plate 4, and fig. 9, Plate 3. . Hleodes distans (female).—Dorsal view of the right valve; compare with fig. 19, Plate 4. Embaphion contractum (female).—Dors o view of the apical moiety of the right valve; compare with figs. 6, 8, 9, and 14. 59780—Bull. 68—09 3 3 514 BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Fig. 21. Eleodes lecontei (typical; female).—Dorsal view of the right valve; compare with figs. 10 and 12, Plate 4. 292. Bleodes scabrosa (female).—Dorsal view of the right valve; compare with fig. 11, Plate 4; also fig. 4 of this Plate. PLATE 6. Fig. 1. Eleodes blanchardii, female—Dorsal view of the right valve; compare with fig. 10, Plate 4. 2. Eleodes hoppingii, female.—Dorsal view of the genital segment; compare with fig. 4, Plate 5. 8. Bleodes caseyi, female.—Dorsal view of the genital segment; compare with fig. 14, Plate 4, and fig. 5, Plate 5. Dorsal view of the genital segment of Cerenopus concolor. Edeagophore of Cerenopus concolor, dorsal view. Sternite of Cerenopus concolor. . Dorsal view of the edeagophore of Trogloderus costatus. Sternite of Trogloderus costatus. . Dorsal view of the edeagophore of Nyctoporus @quicollis. 10. Dorsal view of the genital segment of Nyctoporus aquicollis. 11. Edeagophore of Eleodimorpha bolean, dorsal view. 12. Dorsal view of the genital segment of Hleodimorpha bolcan. 3. Dorsal view of the genital segment of Trogloderus costatus, female. 14. Dorsal view of the genital segment of Trogloderus tuberculatus, female. 15. Last two joints of the maxillary palpi of T'rogloderus tuberculatus. 16. Mentum of Eleodes nigrina, female. 17. Labrum of Hleodes nigrina, female.* 18. Inferior interungual process of a metatarsus in Hleodes dentipes, male. 19. Unguis from metatarsus in FH. dentipes, male. 20. Mentum of Discogenia marginata, male. SD UR A —~ PLATE Ti Fig. 1. Ventral view of the male genitalia of Hleodes cordata, edeagophore partly extruded. 2. Dorsal view of the same, edeagophore completely extruded. 3. Ventral view of the genital segments of Hleodes cordata, female, valves abducted. 4, Dorsal view of the same. 5. Ventral view of the sternite of Eleodes parvicollis, apicale slightly visible. 6. Ventral view of the apicale of the edeagophore of Eleodes cordata, show- ing clave partly abducted and edeagus nearly protruded. Ventral view of the apical half of the edeagophore of Embaphion con- tusum, showing parts in place. 8. Ventral view of the apical half of the edeagophore of Eleodimorpha bolcan, showing relative position of edeagus, Clavee, and subalze; com- pare with figs. 7 and 1. =~ Explanation of figures.—a, apicale; b, edeagus; c, clava; d, basale; e, sternite ; f, apex of right lobe of sternite; g, membranous interlobar region of ster- nite; #, membranous apex of the valve; 7, appendage; /, apical angle of the dorsal plate; k, submarginal groove; /, explanate external border of the dorsal plate; m, ventro-lateral surface; ”, membranous Sth segment, with sclerites of the paranal region and their articulation with base of the ventro- Spb ple Er rile REVISION OF ELEODIINI—BLAISDELL. 515 lateral lamina showing through; o, sternite of the 7th segment; 0’, median basal process of the same; p, sternite of the Gth segment, figs. 1 and 3; tergite of the 6th segment, figs. 2 and 4; g, supra-anal plate; 7, tergite of the 7th seg- ment; s, fossa for the appendage; ¢, superior pudendal membrane; wu, dorsal plate; v, valvular membrane; w, inferior pudendal membrane; «#, genital ori- fice; y, inflexed ale of the apicale; 2, inflexed sides of the basale. XN PLATE 8. Fig. 1. Head of Eleodes dentipes, showing lines of measurements: ad, median line; b, post-marginal ocular line; ¢, post-ocular line. 2. Left mandible of Hleodes quadricollis (female), internal surface show- ing: molar (c), lacinia (b), and post-molar membranous lobe (d). 3. Labium of Hleodes dentipes: a, the chitinous paraglossa which is connate with the body. 4. Diagram showing the relative lengths of the different segments of the legs and abdomen in Hleodes sulcipennis: A, femora: a, metafemora; b, mesofemora; ce, profemora. B, tibiw: d, metatibia; ec, mesotibia; f, protibia. C, tarsi: g, metatarsus; h, mesotarsus; i, protarsus. D, abdominal segments: j, first visible; k, second; 7, third; m, fourth; n, fifth. 5. Mentum and labium of Hleodes scabrosa. 6. Mentum and labium of 2. clavicornis, slightly extended to show relation between labium and lateral lobes of the mentum, the middle lobe free at apex. 7. Protibial spurs of Hleodes quadricollis, female: a, posterior; b, an- terior. 8. Side view of the first two joints of the protarsus of Eleodes sulcipennis, female: «a, tuft of modified spinules at apex beneath, the latter thickened and produced; b, apico-marginal tuft of spinules bounding groove at apical margin of second joint; c, apical spines. 9. Outline of side view of first two joints of the protarsus of ZH. sulci- pennis, male, showing the greatly thickened (@) and produced apices beneath, spines not shown, groove obliterated. 10. Inferior surface of right mandible of HE. dentipes, female, X 28. 11. Maxilla of H. dentipes, female: a, palpus; b, galea; ¢, lacinia. 12. Superior surface of right mandible of #. dentipes, female, X 28. 138. Hypopharynx of HE. dentipes, female, X 28. 14. Protibial spurs of H#. sulcipennis, female: a, anterior; posterior without letter. 15. Inferior view of the mouth parts of H. dentipes, female, showing men- tum, submentum, gular sutures, labium partly exposed, maxilla, and mandibles. 16. Ventral view of the first two joints of a protarsus of H. dentipes, male, showing entire plantar groove. 17. Ventral view of first two protarsal joints of H. dentipes, female, showing plantar groove entire. 18. Lateral view of first two protarsal joints of HL. scabricula, male, showing first joint moderately produced at tip beneath, groove entire on the second. 19. Lateral view of first two protarsal joints of H. scabricula, female, show- ing tip of first joint more produced with tuft of moderately fine modi- fied spinules, 516 Fig. 20. 21. 99 ——s BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Outline of side view of first two protarsal joints of BE. quadricollis, male: tip of first joint scarcely at all produced, with apico-marginal tufts. Outline of side view of first two protarsal joints of H. quadricollis, female; first joint distinctly produced at apex beneath, with tuft of modified spines, groove interrupted. Outline of side view of first two protarsal joints of H. longicollis, male: a, tuft of golden and rather soft pubescence, groove interrupted; b, second joint with pointed apico-marginal tufts, groove entire between the same. Outline of side view of first two protarsal joints of E. longicollis, female; joints partly tipped to one side showing the entire plantar groove. 2 Bleodimorpha bolcan.—a, mentum, buceal fissures and produced buccal processes; the submentum is subobsolete, gular sutures distinct as in Hleodes; b, outline of the ventral apical margin of the prothorax, arcu- ate at middle, sinuate laterally. PLATE 9. Dorsal view of the edeagophore of Hydrophilus triangularis; a, and d, the inner and outer lobes of Packard. Ventral view of the same. Dorsal view of the edeagophore of Blaps similis, showing the apicale as formed by the uniting of the inner lobes of Packard. Ventral view of the same, showing the pallium (a) and edeagus (d). Dorsal view of the edeagophore of Eleodes dentipes. Ventral view of the same. Ventral-view of edeagophore Hleodes planipennis. Dorsal view of the edeagophore of Hleodes scabricula. Ventral view of the same. . Dorsal view of the genital segment of Hydrophilus triangularis, female. Ventral view of the edeagus (@), basal chitinous.processes (@) and clave abducted (d) of Hleodes dentipes. Dorsal view of the same. PLATE 10. Notr.—The bilateral asymmetry is intentional to illustrate variations which are constantly observed, when the pronota are viewed from above. ioral ~ Or CO tf ~ 9 10. salt: 12. 13. Pronotum of Hleodes debilis, female. Pronotum of Eleodes omissa, male. ; Pronotum of Lleodes omissa, female: variations a and Db. Pronotum of Hleodes omissa, female; variations @ and Db. Pronotum of Fleodes pygmea, female type. Pronotum of Hleodes pygmaa, female, variation. Pronotum of Hleodes ampla, male. Pronotum of Hleodes ampla, female; a, marginal curve as viewed obliquely from the side and from above; b, as viewed vertically from above. Pronotum of Lleodes carbonaria, forma glabra, female. Pronotum of Lleodes carbonaria, forma typica, female. Pronotum of Lleodes carbonaria, var. soror, female. Pronotum of Hleodes obsoleta, var. porcata, male. Pronotum of Lleodes obsoleta, male; a and b, showing variations in form. REVISION OF ELEODIINI—-BLAISDELL. 517 Fig. 14. Pronotum of Hleodes obsoleta, female; a, as seen vertically from above; b, as seen obliquely from above and to the side. 15. 16. Nice 18. DE 20. Zl Pronotum form. Pronotum form. of Hleodes knausii, male; of Hleodes knausii, female; illustrating two variations in illustrating two variations in Pronotum of Hleodes quadricollis, male; illustrating two variations in form. Pronotum of Hleodes quadricollis, female; illustrating two variations in form: a, viewed vertically from above; b, as seen obliquely from above and to the side. Pronotum Pronotum Pronotum form. Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum forin. Pronotum Pronotum reversion to the ancestral of Eleodes humeralis, female. of Hleodes of Bleodes of Elesdes of Bleodes of Eleodes of Bleodes of Eleodes of Eleodes of Eleodes of Hleodes of Eleodes of HLleodes of Bleodes tricostata, female. neomexicana, female. pedinoides, male. obscura, female. hispilabris, female. sponsa, female. sponsa, female; a variation. caudifera, female. caudifera, male; a variation. longipilosa, female. gracilis, female. humeralis forma granulata-muricata. cuneaticollis, male; illustrating two variations in of Hleodes dentipes, female; illustrating two variations in of Llecdes dentipes, female; taken from an aberration. of Eleodes dentipes, female; drawn from an aberration in my own collection; the right side is that of confinis, and probably a armata, Pronotum of Hleodes dentipes forma confinis, female; drawn from the type in my own collection; probably a reversional form. of Bleodes armata var. impotens, female; drawn from the type. Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum . Pronotum of Eleodes armata, female; a variation. of Eleodes armata, male. of Eleodes militaris, female. Bran lale of ELleodes femorata, male. of Eleodes laticollis, male. of Eleodes acuticada, male; typical. of Eleodes luca, caudate form. of Eleodes eschscholtzi, female. of Eleodes tenuipes, male. of EHleodes wickhami, male. of EHleodes ventricosa, male. of Eleodes ventricosa, female; variation. of Eleodes falli, male; drawn from type. of Bleodes grandicollis, male. of Hleodes insularis, female. of Hleodes subnitens, female. 518 Fig. 14. Fi ev > 15. 16. aT. 18. 19: 20. Zl 22 23. Pie a. 26. al. 28. 29. aa ov. 31. » 32. oo ov. 9 ot. OF ow. 36. 97 ol. 38. 16. i 18. 9; 20. Zi; Ppa 99 amt De 24. 20. BULLETIN 63, UNITED STATES NATIONAL MUSEUM. Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum lating h Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum of Eleodes fusiformis, female. of Hleodes striolata, male. of Hleodes goryi, female. of Eleodes opaca, female. of Hleodes veterator, female. of Eleodes arcuata, female. of Lleodes extricata, male; variation. of Lleodes extricata, female. of Eleodes granulata, male, forma tuberculata. of Eleodes vandykei, male. of Eleodes vandykei, female. of Eleodes letcheri, male. of Elcodes letcheri, female. of Eleodes pilosa, male. of Hleodes pilosa, female, variation. of Hleodes pilosa, female. of Hleodes hirsuta, female. of Hleodes granosa, male. of Hleodes granosa, female. of Eleodes tibialis, male. of Eleodes inculta, female. of EHleodes snowiti, female, New Mexico. of Hleodes snowii, female, Arizona. of Rleodes tenebrosa, female. of Hleodes clavicornis, female. PLATE a2: of Hieodes hornii, female. of Eleodes neotome, female. of EHleodes consobrina, female, typical form. of Eleodes scabrosa, male. of Hleodes rotundipennis, male. of Bleodes rotundipennis, female. of Lleodes parvicollis, male, typical form. of Eleodes producta, male. of Hleodes fuchsii, female. of Bleodes cordata, female. of Eleodes pimelioides, male, two variations. of Bleodes caseyi, female. of Hleades nigrina, male, showing two slight variations. of Eleodes nigrina, female, drawn from an aberration umeralis, of Hleodes perlonga, male. of Hleodes nevadensis, male. of Hleodes dissimilis, male. of Lleodes schiwarzti, female. of Eleodes gigantea, male. of Bleodes gigantea, female. of EBleodes estriata, female. of Bleodes longicollis, male. of Eleodes longicollis, male, elongate cylindrical form. of Hleodes innocens, female. of Bleodes marginata, male. simu- @, r - t Fig. 26. 20. So bh bo S 0 9 ie wo oo t OD © om 0 oo oo Ce DAD = Se eee eae So Ol ® OO =] at 15. 14. Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum Pronotum . Pronotum Pronotum 519 REVISION OF ELEODIINI—BLAISDELL. of Eleodes scabricula, male. of Eleodes planipennis, female. of Embaphion muricatum, male. of Hmbaphion contusum, female. of Hmbaphion planum, male. of Hmbaphion glabrum, female. of Embaphion elongatum, male. of Lleodes hornii, male. of Hleodes distans, male. of Eleodes subpinguis, female. of Eleodes gentilis, male, drawn from a LeConte type. of Hmbaphion contractum. of Trogloderus tuberculatus, female. of Lleodimorpha bolcan, wale. PLATE 13. Pupa of Eleodes clavicornis, ventral view. Pupa of Hleodes clavicornis, dorsal view. Pupa of Eleodes clavicornis, lateral view. Larva of EHleodes dentipes, dorsal view, 10 mm.; x 10. Dorsal view of the pygidium of Hleodes dentipes. Ventral view of the last five abdominal segments of the larva of EHleodes dentipes, showing position of stigmata on the pleurites. Ventral surface of the right prothoracie leg of the larva of Hleodes dentipes. Ventral view of the maxilla of the larva of Eleodes dentipes; lettering same as on fig. 11. Dorsal view of the head of the larva of EB. dentipes. Antenne of the same, X 28. Gula (g), mentum (mm), submentum (s), a membranous lobe (a), and labium of larva of EH. dentipes. Dorsal surface of the left mandible of the larva of Eleodes dentipes. Ventral surface of the right mandible of the larva of Eleodes dentipes. Internal or oral surface of the left mandible of the larva of Hleodes dentipes, showing molar. Pygidium of Hleodes larva shortly after hatching, x 60. Antenna of Blaps larva. After Gissler. Pygidium of Hleodes larva. After Gissler. Pygidium of Blups larva. After Gissler. After sissler. AND Sexe The black-faced numbers indicate generic or specific descriptions. A. Page. AeUibayss laps see ne ae ce pAseree = cl-sci-laisis leila 195 BCOd CSke ii naeacroseae soe tas semen 185,195 acuticauda, Eleodes........-......---- 234, 276, 505 PAIS (f2) BINT CR Ae Se eet isiein = aateeieie leer 473 altematassbimeliay. 325.2. 5- cose eeiesei= ce se 104 Am pPhid Onaemasese ees saosin cfominiseeeisis 3 amiplaHleOd @Seacae nee sa -5 5-3 =eeem soci 38, 40,53 amthracina elCOCeS = arse nis seis —...--.-.- 310, 311, 312,339 IPRISID Shaan oct eshte ate Se tecisiee ee ete 3, 25, 29, 30, 502 CU ae eee ae sete lot nine cae eee (eee 195 Car bonablianers —secie- ccc aoceeeae aoe acacia 45 Cin CAL Mee eee eee COSe Reece ae cecmeas 121 NISPUADLISss cee Sases Ses eee esse saeee = 217 PLC EON Aba eee ee eee aed = eee ene a 504 ODSCUNAS Ee See eee eee eee erase eeee 187 ObSoleta Sasa sae t Ses ae eh Bn ee Sa 5s (OWEN bc Soaedomap ole adadeaenrapasdonGody 175 Sinan Sees ee eo ecto Ste eet encores 504 SUUMAIS hare ane Ae ae eis cenisieeeeee 199 WI COSTA TAS Sse ce Soe sects Sartore eislociestne asp LOS BIRD i Gesy wsaeesce= 2 Seen. eee eee eee 38, 40,58 GISNE TSH aeeee aeee oe eee Sse 185, 187,190,191 UO POL CALA =2 = ee eae eee 38, 40, 58, 63 GIS SIMS! eee nna noe cee nate 393,298 obtusa. 25 Saiacosas eee cee ee 127,130 VOTANEVAGENSIS 2 epee ae seesaw eee 393, 402 OMUSS@is!3 cs eae ee See ee Ae ee ee 38, 40, 72 GIStEMS Sep Sac can ete eeae re hoe cece ees 242 V7; PCNINSUATiISSs Sess ee seen eee 38, 40,79 GoloSates kee So eee ol ee ree yee 38, 40,47 UU APYEIN = ae re Ae ae 38, 40,77 elegans) scat En Son cee ae oe eerie 251,255 ODAC Sete eS ele eee eee 156,175 ClOM PATS 354 veces 55s ce eee Ss oene 505, 506 Narvicollisesee eas Meee se ee 310, 311, 312,354 ESCHSCHOIMZIE Sepanc ed secs See eee ee 235,285 UGT\COUSTICLA= eee Ee eneee cee tee 312,355,365 DUNS LUGS: ees 7a ee ere ee 235. 285,289 Dlanatatce ssa: aoe eee eee 312, 355,359 eStridtars . cot oceno se wes ae eases 412,421 PLOGUCtaS pea sae eee 312,355,362 ORG CAG ae hee eae COR eee ee 116,124 edinoidess.= yee oes ere 38, 41, 109 VOT ATIZONENSIS$ 2-2 soe secon eee eeeae cee 125 UOT RCOMEK CANA ass =e. eee 38, 41,118 fea ieee ee ee ena Se aC en ee nee 235, 305 Peninsulanise-2 ese See eee Cee 38, 40,79 fomOratacss Boss vse ne ee eee ccaes 272 | DELON Pass n ee Se Wes a eee eee 393,398 fischeriss 2, ppc se eee oan ee Sees ASO iy = MOS oes codec cme seat San eee 138, 142 fl CHSH sant az senor ee 310, 311,312,343 | pimlelioidesssce. sete seaeee eee 310, 311,313, 384 FUSWORMISSa-¢ acces ee eee eee 156,172 | UT sOLUNTLPES. semen ease eee 313,384, 386 Pentilisus cesses Seok eee oc See eros 412,416 planatancc osc ace snes eee eee 104 piganiteaes east nas Reo ose sees teas 411,412,496 planipenniss<-.ese cca accesses eee eee 435, 445 VAT ACSETIA bo te acta te arate eee 412.421 POL CHE Rie ws eietm areas eee ee 38, 40, 63 SOM Soseee see Ee eet aes 41284169) productasees s-- see mee _cemeee nee eee eee 312, 362 POLY eset s eee nee See eee eee 156, 164 | pLominens? soc see ee eee eee 251, 254 Sracilist se so oor Seok coc eee seer 235,237 Py euNse a A see sete tee eecie e eer 38, 40, 77,418 BOR CIStANS ser Se aaa eee 235, 242 | QUadricolliss:.5- 2h ci. l eee. Sosa ee see oor aOHOn STANGICONIS!.\: ners: aoe AS eee Races 207 VEr ANLNTACIN A=. a= sta asso ee Sees 38, 40,87 STAN OSA ott setae na ee ee nea eee eee 147 | LUSEPANS - -stacine ee oe eaeeiscenie = 38, 40,89 STANDS Je Se = ae eee ateeeecone 116,127 TOME KICOMIS hs sss e-sneeees sacs eee ee 450,494 Naydeni: tse. 2 sse she oe ooee eee ene oeee ears 25 WUC Vis eck clesa cee see see ce eae ee 38, 41, 100 HITSUba Sse. thn Sse ee Soe eee 138,139 MODUSUM eck. = = S epaare Oe eee ie ae i eee 104,106 hispulabnisens seer ses eee sene Cee eee 212,217 scabrictlas. 2.2. se ot aoe ee ae eee 435,440 Hoppin giiees eee esse ee 310,311, 312,368 Scabripennis),..- 35. .-.---e cides = 5 $ 33 a > 2 2 : : i ei : we GENITAL CHARACTERS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 509, 510. a ; U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 2 . . | Phe! |e il nid i fH ; \ GENITAL CHARACTERS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 510, 511. U. S. NATIONAL ‘MUSEUM BULLETIN NO. 63 PL. GENITAL CHARACTERS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGE 511. 597S0—Bull. 68—09——34 RE eee YF ores Pr ye U. S. NATIONAL MUSEUM GENITAL CHARACTERS OF THE ELEopIINI. FOR EXPLANATION OF PLATE SEE PAGES 911, 512, 513, BULLETIN NO. 63 PL, 4 U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 5 4 "4 + ‘ ; ‘ ‘ ‘4 Je le ' =e j GENITAL CHARACTERS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 513, 514. U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 6 aa \ j Te aE fe o ay WIP, iN fi COMPARATIVE AND GENITAL CHARACTERS OF THE ELEODIINI AND RELATED TRIBES. FOR EXPLANATION OF PLATE SEE PAGE 514. U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 7 TOPOGRAPHY OF THE GENITAL SEGMENTS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 514, 515. Xf Sav VARY «4, Pet ape 2X" U. S. NATIONAL MUSEUM ANATOMY OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 515, 516. BULLETIN NO. 63 PL. 8 U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 9 Seer HOMOLOGY OF THE GENITAL SEGMENTS OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGE 516. wre ee ee U. S. NATIONAL MUSEUM PRONOTA OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 516, 517. BULLETIN NO. 63 PL. 10 U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 11 PRONOTA OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 517, 518. 59780—Bull, 63—09——35 U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 12 - AC a a oe r S é » i PRONOTA OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGES 518, 519. U. S. NATIONAL MUSEUM BULLETIN NO. 63 PL. 13 EARLY STAGES OF THE ELEODIINI. FOR EXPLANATION OF PLATE SEE PAGE 519, | ee » = SONIAN INSTITUTION LIBRAR ‘Will sll | I! i