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CATALOGUE
OF THE
FOSSIL SPONGES
IN THE
GEOLOGICAL DEPARTMENT
OF THE
BRITISH MUSEUM
(NATURAL HISTORY).
WITH DESCRIPTIONS OF NEW AND LITTLE-KNOWN SPECIES.
(ILLUSTRATED BY 38 LITHOGRAPHIC PLATES.)
BY
GEORGE JENNINGS HINDE, Pu.D., F.G:S.
LON DON:
PRINTED BY ORDER OF THE TRUSTEES.
1883.
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PRINTED BY TAYLOR AND FRANCIS,
RED LION COURT, FLEET STREET.
PREFACE.
In submitting the accompanying Monograph to the scientific public, I feel assured
that the careful and conscientious manner in which Dr. Hinde has performed the
very arduous task entrusted to him will meet with the well-earned recognition of
all those who consult this work.
The earlier methods of classification of both the recent and fossil Spongide by
their external form and habit of growth having been thoroughly tested and proved
to be unreliable, the only sure method to be pursued—that of classifying them
according to their intimate microscopic structure—has been adopted by Dr. Hinde,
who, himself a pupil of the distinguished paleontologist Prof. Dr. Karl Zittel of
Munich (one of our highest authorities in this class of organisms), has followed
patiently the careful methods of research set forth in the ‘ Beitriige zur Systematik
der fossilen Spongien’ and other of Zittel’s works.
It is no small gratification to know that the determination of this very difficult
group of fossil organisms has now been so far completed, and that the types
of those species not heretofore described are now all carefully figured in the Plates
accompanying this Monograph, and the original specimens are preserved in the
British Museum of Natural History.
HENRY WOODWARD,
Keeper of the Department of Geology.
Britisa Museum (Narvrat History),
DercemBer 26rH, 1883.
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AUTHORS PREFACE.
Ar the request of the Keeper of the Geological Department in the Natural-History
Museum, I commenced, in the early part of 1881, the study of the collection of
Fossil Sponges in the Museum with the view of arranging them systematically and
preparing a simple Catalogue of their specific names and references. Hitherto no
attempt had been made to place the Fossil Sponges in scientific order, and with the
exception of the limited portion of the Collection formerly exhibited in the Museum
at Bloomsbury, which had been mostly included in the comprehensive genera of
Spongia, Scyphia, Manon, &c., the greater number of the specimens simply retained
the labels of the localities and horizons from whence they had been derived. A
preliminary examination made evident the fact that numerous specimens, more
particularly those from British strata, were either quite new to science or had been
described and figured in such an imperfect manner that their real characters were
unknown. Under these circumstances a mere catalogue of names and references,
such as had been at first proposed, would have been quite valueless; and, with the
consent of the Trustees, it was decided to enlarge the plan and embrace in the
Catalogue condensed descriptions of all the species from British strata and of the
new species from foreign localities, with figures of all the new forms as well as of
those which had been either inadequately figured previously, or of which it was
desirable to illustrate the minute structure. To carry out this plan involved a far
greater expenditure of time and work than was at first anticipated; but it is hoped
that the results of nearly three years’ study will be to place our knowledge of this
group of fossils on a more satisfactory basis than hitherto.
The Museum collection includes examples of a very large majority of all the
species of Fossil Sponges at present known, and, as may naturally be supposed, it is
especially rich in those occurring in the rocks of this country. It contains amongst
others the typical collections of William Smith, “The Father of English Geology,”
Toulmin Smith, Mantell, Bowerbank, and Cunnington. The fossil sponges from the
Jurassic and Cretaceous strata of France, Switzerland, and Germany are also well
represented. With a few unimportant exceptions, the Sponges from these countries
have been clearly figured in the works of Goldfuss, Roemer, Michelin, Quenstedt, Reuss,
Geinitz, de Loriol, Fromentel, and others; and as their minute spicular structure has
Vill PREFACE.
been already described and illustrated in the works of Prof. Zittel, it has only been
necessary for the purpose of the Catalogue to give references to the works in which
the description of these forms will be found. The Sponges from British strata, on
the other hand, have not received a similar amount of attention from the paleon-
tologists of this country; and, with the exception of the forms treated of by Toulmin
Smith and those lately described by Prof. Sollas, no systematic endeavour has hitherto
been made to describe these organisms in detail; and therefore it has been necessary
to supplement the meagre notices usually given, by such brief references to their
minute structure as will enable them to be recognized.
I have followed in this Catalogue the classification of Prof. Zittel, the only one,
in fact, which is at all applicable. Limitation of space has necessitated the omission
of generic diagnoses, save in those cases in which new genera are introduced, or in
which it has been needful to make emendations; and the student must therefore
refer for these to Zittel’s ‘Handbuch der Palaeontologie’ or to the “Studien tiber
fossile Spongien” (Abh. bay. Akad. d. Wiss., Bd. 153, 1877-78). An English trans-
lation of the main portion of the generic descriptions, by W. 8S. Dallas, F.L.S.,
is contained in the ‘Annals and Magazine of Natural History’ for 1877, 1878,
and 1879.
In a few instances I have described species of which the Museum does not at
present possess any examples; but these forms will be found in the Museum of
Practical Geology in Jermyn Street, and were kindly lent for examination by
Prof. Archibald Geikie, LL.D., F.R.S.
The illustrations of the minute structure will, in some cases, appear defective—a
result due, not to any lack of ability on the part of the artist, but to difficulties arising
from the state of preservation of the fossils themselves. In all cases the endeavour
has been to delineate the structure in its present condition.
I regret that it has not always been possible to give the exact localities of many of
the Sponges from the Upper Chalk. This arises from the fact that the greater part
of the collection made by Toulmin Smith is without any reference to the places from
which the Sponges were obtained; but it is definitely known that they were procured
from the counties of Kent, Surrey, and Sussex, and I have therefore catalogued such
specimens as from the South of England.
GEORGE J. HINDE.
DrrarrMent of Grotoey, British Museve
(Natura History),
Cromwett Roap, Lonpoy, 8. W.
INTRODUCTION.
A SPONGE, in the ordinary acceptation of the word, is the internal framework or
skeleton of an animal whose soft vital parts are composed of cellular protoplasm.
This protoplasm in the living sponge is differentiated into two layers—an inner of
ciliated cells, and an outer layer, named the syncytium, in which the cell-structure in
the ordinary condition is not recognizable. It is in this syncytium or exoderm that
the skeleton is secreted. In one small group of existing sponges the skeleton is
wanting and the entire animal consists of fleshy sarcode; but the rest of the class
secrete skeletons of very various composition and form. In the division to which
the ordinary sponge of commerce belongs, the skeleton is composed of elastic fibres of
a horny character, which anastomose together and build up those porous structures
with which we are all so familiar. In another group the horny fibres have a central
core composed of minute siliceous spicules usually secreted by the animal, though
occasionally the central portion of the fibre is filled with grains of sand and other
foreign particles which the sponge has selected from its surroundings. In yet other
sponges the skeleton consists almost exclusively of minute siliceous or calcareous
bodies, more or less intimately united together to form a resistant framework of a
porous character, which is generally traversed by numerous canals. The minute
mineral particles or spicules of the skeleton vary exceedingly in their form and
arrangement in different groups; they possess an interior canal, and are built up of
exceedingly thin concentric layers of silica or calcite mingled with organic material.
The classification of the Sponges, recent as well as fossil, rests upon the characters
of their skeletal structures. The existing forms of the class have been divided into
the following orders :—
1. Myxosponei_a, Haeckel.
Sponges destitute of a solid skeleton.
2. CeRratosponal#, Bronn.
Sponges with skeletons of horny fibres.
iy
7
B
INTRODUCTION.
Lo
3. MonactINELLIDa, Zittel.
Sponges with skeletons of horny fibres with cores of uniaxial siliceous spicules, or
built up wholly of uniaxial siliceous spicules.
4, TErRACTINELLIDZ, Marshall.
Sponges with skeletons of siliceous spicules, usually with four rays or arms, one
generally elongated to form a shaft, the other three disposed in pyramidal form ;
uniaxial and star-shaped spicules are also present.
5. Liraistip#, O. Schmidt.
Sponges with skeletons of siliceous spicules, either four-rayed or irregular in form,
which are intimately interwoven together into a continuous mesh.
6. HexactTINELLID®, O. Schmidt.
Sponges with skeletons of six-rayed siliceous spicules, either loosely interwoven
together, or organically united to form a continuous mesh.
7. CaLcIsponel&, Blainville.
Sponges with skeletons of calcareous spicules.
The first of these existing orders, the Myxospongie, is, of course, quite unknown
in the fossil state, and it is also very doubtful whether any remains of the next
order, the Ceratospongie, have been preserved. Certain casts of cylindrical bodies
from the Cretaceous system have been regarded as belonging to horny sponges; but
in the absence of all other characters but that of outward form these bodies cannot be
definitely placed in this group.
The Monactinellide have comparatively few representatives in the fossil state, and
they present a striking contrast to the abundance of this order in the present seas.
Their rarity as fossils, however, is not to be accepted as an indication of their scanty
existence in the past, but is more probably owing to the fact that the spicular
structure of these sponges is unsuitable to their preservation as fossils.
The structure of the Tetractinellide, like that of the order just mentioned, is also
but little adapted to the retention of the form of these sponges in the fossil state ;
but the constituent spicules of many of these sponges are relatively large and robust,
so that they are capable of preservation, and they are frequently met with detached
and scattered through the rocks. In some instances they are sufficiently numerous
to form thin beds, almost exclusively composed of spicules. It is therefore probable
that this order of sponges flourished as abundantly in the seas of the Neocomian
period as at the present day.
INTRODUCTION. 3)
Lithistid and Hexactinellid sponges, unlike those of the previous groups, are
more numerous and varied in the fossil than the recent state. The spicular com-
ponents of the skeleton in these sponges are firmly attached together, consequently
the form of the sponge is frequently preserved intact, even in cases where the spicules
themselves have subsequently been destroyed. The occurrence of detached spicules
and fragments of the skeleton scattered through the rocks plainly shows, however,
that only under favourable conditions of fossilization has the form of the sponges been
retained, and those now remaining probably comprise but a small proportion of the
number which previously existed.
Fossil Calcareous sponges are abundant in certain strata, but they belong to a
family which differs to such an extent from existing Calcispongie that the relationship
has been greatly doubted. Recent discoveries, however, prove that the component
spicules in the fossil Calcispongiz possess the closest resemblance to those of the
living examples of the order.
In respect to their form and dimensions, fossil sponges present as great diversity as
living ones. The most prevalent form is like a cup, funnel, or vase ; but cylindrical,
club-shaped, fan-shaped, and branching examples are also very abundant. The
smallest complete fossil specimen which has come under my notice is a Calcisponge,
belonging to the genus Peronella, which measures 5°5 millim. in length by 4 millim.
in width; whilst the largest is a Lithistid, belonging to the genus Doryderma, which
reaches a length of 590 millim. by 155 millim. in diameter.
Fossil sponges are of most common occurrence in calcareous and arenaceous rocks,
whilst they are rarely met with in shales and deposits formed from muddy sediments.
Strata containing masses and nodules of flint and chert frequently abound in fossil
sponges; indeed there is every reason to suppose that the flint and chert itself is
derived from the dissolved skeletons of siliceous sponges. Both arenaceous and
calcareous deposits seem to have been favourable to the existence of Tetractinellid
and Lithistid sponges, for they occur alike in the Lower and Upper Green Sands as
well as in the Chalk. The Hexactinellid sponges, on the other hand, favour more
particularly the deeper-formed deposits of limestone and chalk, though rarely forms
of this order are present in the Upper Green Sand ; and, according to Manzoni*, they
are comparatively numerous in Miocene strata in Italy, associated with shallow-water
organisms. ‘The Calcispongiz, on the other hand, are most abundant in arenaceous
or shallow-water deposits, and thus in their habitats resemble the existing members
of the order. Exceptions, however, occur in this group, for several Calcisponges are
found in Jurassic limestones associated with Hexactinellids, and in one example in
the Museum a species of Peronella is growing attached to the surface of a Hexacti-
nellid sponge.
Atthe present day, with the exception of the genus Spongilla, which inhabits fresh
* Spuene silicee del Miocene medio,
pug
B2
4 INTRODUCTION.
water, all the members of the class live in the sea. It is interesting to note that the
spicules of a fossil species of Spongil/a have been discovered in chert, of freshwater
origin, in beds of Purbeck age.
The Alterations produced by Fossilization in the Structure of Sponges.—The
changes to which fossil sponges have been subjected in the process of fossilization
have oftentimes resulted in the partial or complete destruction of their original
spicular structure and the replacement of the mineral composition of the skeleton to
such an extent that great controversy has arisen respecting its original nature. In
recent sponges the skeleton is composed either of amorphous silica or carbonate of
lime with a small quantity of organic material. In fossil sponges the organic material
would of course disappear, and only the mineral constituents of the spicules remain.
But even the mineral portion is seldom, if ever, in the same condition as in recent
spicules ; the amorphous silica and calcite have been replaced by crystalline silica
and crystalline calcite, as well as by peroxide of iron and iron pyrites; whilst not
infrequently the entire mineral structure has been dissolved and removed, leaving the
empty moulds of the spicular skeleton in the matrix. As a result of these changes,
siliceous sponges now occur with skeletons of calcite, and calcareous sponges with
fibres composed of silica. These changes are intimately connected with the character
of the strata in which the sponges are imbedded, but the causes producing them have
not up to the present been satisfactorily determined. In general the sponges in
calcareous strata have undergone the greatest alteration, the siliceous structures
being replaced either by calcite or iron peroxide, or dissolved away altogether, whilst
the structures of calcareous sponges, in common with the shells of mollusks in the
game strata, are oftentimes replaced by silica. In arenaceous or glauconitic strata, on
the other hand, the changes, whether of siliceous or calcareous sponges, have been
much less extensive than in strata of a calcareous character.
Changes produced in the Structure of Siliceous Sponges.—It may be affirmed that
no examples are known in which the siliceous spicules or fibres of fossil sponges retain
the perfect amorphous condition of the silica as in existing sponges. The nearest
approach to this original amorphous condition is shown in sponges preserved in the
glauconitic marls of Westphalia of the age of the Upper Chalk. The spicules of
these sponges have a smooth milky white aspect, and, when examined under the
microscope mounted in Canada balsam, exhibit the details of the interior canals and
the spinous adornment of their surfaces in beautiful preservation. When viewed by
polarized light they are either neutral or occasionally present faint prismatic tints.
The main feature which distinguishes these from recent spicules is their white porcel-
lanic aspect, which contrasts greatly with the glassy appearance of recent spicules.
‘The mineral condition of some of the siliceous sponges from the Upper Green Sand
of Warminster also resembles that of the examples from Westphalia mentioned above ;
but they present the interesting peculiarity of being inclosed ina matrix of translu-
INTRODUCTION. 5
cent glassy silica, which is shown to be crystalline by giving brilliant prismatic tints
under the polariscope ; whilst the spicules, on the other hand, are either neutral or
only give faint prismatic tints, thus indicating that the silica of which they are com-
posed is in a crypto-crystalline condition.
In the siliceous sponges from the Jurassic limestones of Franconia, which yet retain
a siliceous structure, the silica, seen by reflected light, has a white snowy aspect, and
is distinctly crystalline in character, In this condition the spicular elements are
nearly invisible when mounted in Canada balsam, but in glycerine or water show
perfect definition. The mineral change in these sponges, however, does not appear
to have affected the finer structural details of the skeleton, which are usually well
preserved.
Most of the siliceous spicules from the Lower and Upper Green Sand and from the
interior of chalk flints exhibit an exterior surface-aspect like that of ground-glass,
which is seen under the microscope to be owing to the minutely eroded condition of
their surfaces. The silica composing them is now crystalline, and frequently appears
like delicate fibres radiating from different centres. In these spicules the canals are
usually infiltrated with silica; in a few instances, however, they are filled with iron
peroxide.
Another modification of the silica is presented by sponges from the Upper Chalk
of Flamborough, in Yorkshire, which are so distinct in their mineral characters that
they can be at once recognized by this feature alone. On removing the chalky matrix
by acid the sponge itself is usually perfect in outer form, and the fibrous mesh has a
rough surface-aspect, as if composed of minute granular particles of a dull earthy
greyish aspect. ‘Thin sections of the fibre show that the silica is distinctly crystal-
line. In these sponges the spicular components of the skeleton, save in a few examples
where they possess relatively large dimensions, are not distinguishable; they appear
to have been, as it were, fused together, so that the individual forms are entirely
obliterated.
Siliceous Sponges inclosed in Chalk Flints.—The sponges met with in the interior
of chalk flints present very various conditions of preservation. In some examples the
flint is hollow and the interior cavity is partially or wholly filled with a fine powdery
material, largely made up of detached Tetractinellid sponge-spicules, which are now
composed of crystalline silica. In other cases the hollow flint incloses the perfect
form of a sponge, whose walls are either in immediate contact with the interior
surface of the flint, or separated therefrom by a small space usually filled with a white
siliceous powder. The sponges thus inclosed frequently retain the spicular structure
of the outer surface in perfect preservation, particularly those species with relatively
large spicules ; and even in some instances the minute spicules of the dermal layer
of the sponge yet remain intact and in their relative positions. This favourable state
of preservation, however, is limited to the outer surface of the sponge, for the spicular
6 INTRODUCTION.
structure of the interior is completely obliterated, and is now a mass of botryoidal or
porous crypto-crystalline silica, in which even the course of the canals has disappeared.
The surface spicular structure is usually of a snowy white aspect, and the silica is
crystalline.
In other examples, of somewhat rare occurrence, the hollow flint contains merely
the solid casts, in silica, of the central cloaca and canals of the sponge, whilst the
spicular skeleton itself has entirely disappeared.
In all the instances above quoted the sponges are contained in hollow flints; but
numerous examples occur in which the inclosing flint is quite solid. In some cases
the flint nodule has a rough conformity to the form of the sponge, in others no
relation is apparent. As a general rule the spicular structure in these solid ftints is
completely obliterated, and the interior of the flint merely exhibits the cloacal cavity
and canals, which are distinguishable from the circumstance that the silica which has
infilled them is of a different tint from that of the rest of the nodule.
Spicules of Tetractinellid and other sponges also occur scattered loosely in strata
of glauconitic sand and in places aggregated together into distinct beds. Sometimes
the individual spicules can be distinguished on the weathered surface of these sponge-
beds ; but in the interior of the beds the spicular forms are merged into glassy chert.
The silica of these spicules is usually crystalline. These spicules and spicular masses
occur in the Hils formation of Germany, in Neocomian sandstone at Haslemere in
Surrey, in the Kentish Rag quarries near Maidstone in Kent, at Folkestone, and in
the Upper Green Sand of Blackdown and the Haldon Hills in Devonshire. Spicules
of siliceous sponges in connexion with beds or nodules of chert have been met with
in Carboniferous strata in Ireland, in Lias strata from the Austrian Tyrol and
Glamorganshire, and in the Portland limestones of Upway in Dorsetshire.
Siliceous Spicules replaced by Peroxide of Iron and Iron Pyrites—Examples where
the original silica has been replaced by pyrites are not of very common occurrence.
A well-known instance is that of the earliest known sponge, Protospongia fenestrata,
Salter, from the Menevian of Wales, where the sponge is imbedded in black shale.
In a few sponges from the Jurassic limestones, the Grey Chalk, and the Upper
Chalk the same replacement occurs.
The replacement of silica by peroxide of iron is of extremely frequent occurrence,
particularly in the sponges from the Upper Chalk of this country. The peroxide is
present as a light reddish-brown powdery material, usually incoherent, though in a
few instances safficiently firm to retain the spicular mesh intact after the removal of
the chalky matrix. As a general rule, however, the sponge-structure when replaced
by peroxide is extremely indistinct and merely represented by a thin rusty film, in
which the character of the spicules is generally obliterated. Unfortunately most of
the siliceous sponges from the Upper Chalk of the south of England, which have not
been inclosed in flints, are in this very unsatisfactory condition; and the task of
INTRODUCTION. 7
discovering their true characters is rendered very difficult. But even in many of the
sponges inclosed in solid flints the silica has been replaced by peroxide of iron ;
and the former presence of the sponge is indicated by reddish markings in the flint.
These are particularly noticeable in the case of sponges belonging to the genus
Plocoscyphia, which are characterized by thin anastomosing walls. These walls
appear in section on the surface of broken flints as delicate labyrinthine lines of a
reddish tint. In a few cases where the spicular structure is of an open character it
has been replaced by peroxide of iron in such a manner that the form of the
spicules is perfectly preserved.
Siliceous Spicular Structure replaced by Calcite—The fact of calcite taking the
place of silica in fossil sponges has been greatly contested on account of the supposed
more stable character of silica in comparison with calcite. But the frequency of the
occurrence of undoubted siliceous sponges with skeletons in the condition of calcite
is a convincing proof of the instability of the colloid silica of sponge-skeletons. The
calcite which has replaced the silica is in a crystalline condition, and it appears to
have infiltrated the moulds after the siliceous skeleton has been dissolved and
removed. In the case of relatively large spicules, and where the skeletal mesh is of
an open character, the replacement has been effected without obliterating the cha-
racter of the structure; but in those instances where the spicules are minute and
they are united closely together, the finer details have disappeared, and only general,
ill-defined outlines of the original spicules and skeleton remain. As a rule the
spicular canals are obliterated where calcitic replacement has taken place; but occa-
sionally some other material has filled in the canals previous to the deposition of the
calcite, and in that case they can be distinguished.
Instances of the replacement of siliceous sponges by calcite occur in Silurian,
Devonian, Jurassic, and Cretaceous strata. The replacement appears to be confined
to sponges imbedded in calcareous or marly deposits. Iam not aware of a single
instance occurring in arenaceous beds. ‘The majority of the siliceous sponges from
the Jurassic limestone in certain localities are now calcareous, whilst in the same
strata elsewhere the same species of sponge remains siliceous. Not unfrequently too
the same individual specimen will be partly siliceous and partly calcareous ; in the
siliceous portion the spicules will retain their canals and other finer details, whilst
in the calcareous portion the spicules will be solid and the outlines of the skeletal
mesh confused and indefinite. In the sponges from the Upper Green Sand (so-called)
of Cambridge and the Grey Chalk strata of Dover and Folkestone the silica has been
almost entirely replaced by calcite. In some of the Upper Chalk sponges from
Flamborough, calcite has partially replaced the silica; but, as a rule, the skeletons
of the sponges from this locality remain siliceous. In only one instance have I found
detached spicules of siliceous sponges changed to calcite, and this occurs in the
spicules of Astrwospongia from the Silurian strata of Gotland.
8 INTRODUCTION.
Siliceous Spicular Structure dissolved, leaving empty Moulds.—This condition is
of frequent occurrence in beds where the matrix is suitable for retaining the form of
the spicular mesh, and it furnishes a clear proof of the ready solubility of siliceous
sponge-structure. The siliceous skeleton of these sponges has been entirely removed,
so that there are only empty cavities in the place of the solid spicular arms and
nodes. These moulds are sometimes present in a matrix of chert or flint, of which
there are examples in the Silurian strata of North America and in the Upper Green
Sands of Wiltshire ; they also occur in compact J urassic limestones, in the Cambridge
Green Sand, and even in the Upper Chalk of the south of England. It is somewhat
surprising that the empty moulds of the delicate sponge-skeleton should have been
retained in the soft Chalk of the south of England; but they have been so perfectly
preserved that in many instances the moulds of the delicate balks or rods which form
the lantern-like nodes of Hexactinellid sponges remain intact. Not only are the
connected skeletons of Hexactinellid sponges thus dissolved, but the detached spicules
of Tetractinellid sponges, though relatively much larger and more robust, have also, in
many instances, suffered a similar fate. A noted example occurs in beds of the Hils
sandstone in the north-west of Germany, where the moulds of the spicules remain in
a matrix of semi-translucent chert, giving to it a porous aspect. Other instances are
present in Lower Green Sand chert from near Haslemere in Surrey. In some of the
specimens from this latter locality the spicular canal had become infilled with silica
before the solution of the spicule itself, and this solidified canal now remains as a
delicate rod in the central axis of the hollow mould. In some sponges from the
Upper Green Sand the spicular skeleton seems to have been incrusted with a thin
delicate film of silica, which now remains after the solution and removal of the
skeleton. In some examples of Miocene Hexactinellid sponges described and figured
by Manzoni* the skeleton is similarly incrusted with a siliceous pellicle, but it has
not been subsequently dissolved.
Changes produced in Caleareous Sponges by Fossilization—As a rule the mineral
structure of fossil calcareous sponges has undergone fewer changes than the structure
of siliceous sponges, though, as a result of the small dimensions of the calcareous
spicules and their very intimate disposition in the fibre, a very slight amount of
alteration has been sufficient to obliterate the form of the individual spicules, and to
merge them into a common fibrous mass of amorphous or crystalline calcite. In
general the calcareous sponges imbedded in arenaceous strata are better preserved
than those from calcareous beds. In the numerous calcareous sponges from the
Neocomian gravels of Farringdon, in Berkshire, the sponge-fibre is enveloped in a
coating of dog-tooth crystals of calespar. The microscopic structure of these sponges
is very imperfectly preserved. ‘The fibre of the calcareous sponges from the Upper
Green Sand of Warminster is of a soft earthy character with a greyish-white tint, and
* Spugne silicee del Miocene medio, t. 5. f. 15.
INTRODUCTION. 9
in some cases so slightly altered that the minute component spicules can be detached.
The fibre of the calcareous sponges from Essen resembles in appearance that of the
Warminster sponges; but the individual spicules are, as a rule, not recognizable.
The fibre of the calcareous sponges from the Upper Chalk of the south of England
is in appearance smooth, white, and unaltered; but in thin sections the component
spicules are rarely visible, and the fibre presents either a homogeneous mass of
amorphous calcite or a very fine radiate crystalline structure.
Caleareous Sponges replaced by Silica.—The instances in which this change has
been effected are comparatively rare, and limited, with few exceptions, to sponges
imbedded in strata in which the remains of other calcareous organisms have likewise
been replaced by silica. I have not met with a single example of this change in any
of the numerous calcareous sponges from the Lower and Upper Green Sands of this
country ; but in the Jurassic strata of certain localities in Germany many of these
sponges have become silicified, and can be perfectly freed from the calcitic matrix
by acid. The fibre of such sponges after treatment with acid has a rough appear-
ance, and is of a snowy white tint; and, as may be expected, all traces of spicular
structure have disappeared. A partial replacement by silica is sometimes present in
examples of the well-known Pharetrospongia Strahani, Sollas, from the Upper Green
Sand of Cambridge. The fibre of these sponges is calcareous in the central portion,
whilst the exterior has been replaced by silica. This change is more remarkable
from the fact that the form of the individual spicules has not been destroyed, but
they can be distinguished in a siliceous condition. This same species of sponge
remains completely calcareous when imbedded in the Upper Chalk; but when in-
closed in solid flints the outer surface of the fibre has become silicified, whilst the
interior remains calcareous, in the same manner as in the Green-Sand specimens.
GEOLOGICAL DISTRIBUTION.
Fossil sponges make their appearance in some of the earliest fossiliferous rocks,
and they are present, though by no means in continuous sequence, in all the great
divisions of the geological scale to the Tertiary era. ‘They form, however, but a very
subordinate part of the fossil fauna of the Paleozoic and early Mesozoic groups, and
it is not until towards the close of the Jurassic and during the Cretaceous periods
that they occur in sufficient numbers to constitute a relatively important feature of
the fauna of these rocks. Judging by the constant association of sponges with beds
of flint and chert in the Mesozoic rocks, there is great reason to suppose that they
may have existed in strata of Paleeozoic age where beds of these materials are present,
though up to the present they have not been recognized in them.
Cambrian System.—The earliest known sponge is the Protospongia fenestralis, Salter,
belonging to the Hexactinellide. It is only known from fragments and detached
spicules, preserved in black shales in strata of Menevian age in Wales and in Sweden.
Cc
10 INTRODUCTION.
In the Quebec series of Canada there are numerous fossils which have been
referred to sponges by the late Mr. Billings; but as nothing definite is yet known of
their minute structure, the true character of these fossils remains doubtful. ‘They
have been placed in the genera Archwocyathas, Calathium, Eospongia, Rhabdaria,
and Trachyum.
Silurian System.—Sponges are by no means generally distributed in Silurian strata,
though they are not uncommon in beds of the Niagara series in North America and
in strata of corresponding age in the Silurian basin of the Baltic. They are also
present in the Boulder drift beds of North Germany. The order Monactinellide is
here represented for the first time by the genus Climacospongia, the Lithistide by
Aulocopium, Hindia, and the somewhat doubtful Aulocopina. Belonging to the
Hexactinellide are the genera Protachilleum, Astylospongia, Palwomanon, and the
remarkable genus Astrwospongia. The genus Amphispongia from the Upper Silurian
of the Pentlands isa dubious form, and also the genus Brachiospongia from the
Niagara group of North America.
Devonian System.—Very few sponges have as yet been discovered in rocks of this
age. ‘To the Monactinellide belongs the genus Lasiocladia from Belgium. No
member of the Lithistide has yet been discovered in this division. The Hexac-
tinellide are represented by Astrwospongia from Belgium and Illinois, and by the
peculiar genus Dictyophyton from the Upper Devonian strata of New York. Calca-
reous sponges belonging to the genus Peronella also appear here for the first time.
Carboniferous System.—From the Carboniferous Limestone of Scotland the Monac-
tinellid genera Pulvillus, Raphidistia, and Reniera have been described by Mr.
Carter, as well as the Hexactinellid Hyalostelia and Holasterella. Detached spicules
of Lithistid and Tetractinellid sponges also occur. Lately Steinmann has described
three genera, Sollasia, Amblysiphonella, and Sebargasia, of the family of the Phare-
trones, from the Carboniferous of Spain; but as no spicular structure has been pre-
served, the character of these forms is at present uncertain.
Permian System.—No well-ascertained sponges have been met with in this division.
According to Prof. Zittel, the genus Bothroconis, King, might prove to be a Hexac-
tinellid, and EHudea tuberculata, King, a calcareous sponge; but these forms have
not come under my notice, and, judging from the figures given of them, their true
characters appear to be very uncertain.
Triassie System.—No siliceous sponges have been recorded from this division, but
in the St.-Cassian beds in the Tyrol numerous calcareous sponges are present. Asa
rule, the fibre of these forms does not show any minute structure, and the sponge-
character of some of them is very problematic. These St.-Cassian forms have been
included in the following genera—EKudea, Colospongia, Verticillites, Celyphia,
Mimatella, Peronella, Corynella, Myrmecium, and Stellispongia. :
Liassic System.—It is only within the present year that fossil sponges of this
INTRODUCTION. 11
period have been discovered. In strata of this age at Schafberg, Austria, Prof.
Zittel discovered beds of agglomerated spicules of Monactinellid, Tetractinellid, and
Hexactinellid sponges, which have been described and figured by Dunikowski*. In
cherty strata in Glamorganshire there are also numerous spicules.
Jurassic System.—The sponges of this division are very numerous. The Monac-
tinellidee are represented by the single freshwater genus Spongilla, from Purbeck
strata in the south of England. The Lithistidee and Hexactinellide are met with
in great numbers in different zones of the Upper Jurassic, in South Germany,
Switzerland, Eastern France, and in the vicinity of Cracow; but they are entirely
absent in the strata of corresponding age in England. In some of the Portland beds,
however, there are large masses and nodules of flint, which probably indicate the
former existence of sponges in these strata; and in some of these flints detached
spicules of Tetractinellid sponges are still preserved. The Lithistids of the Upper or
White Jura of Germany belong to the following genera—Cnemidiastrum, Coral-
lidium, Hyalotragos, Pyrgochonia, Leiodorella, Platychonia, Placonella, Megalithista,
Cylindrophyma, Melonella, and Lecanella. ‘The Hexactinellids are included in the
genera Tremadictyon, Craticularia, Sphenaulax, Sporadopyle, Verrucocelia, Pachy-
teichisma, Trochobolus, Phlyctenium, Cypellia, Stauroderma, Porocypellia, Casearia,
Porospongia, and Toulminia. In most of these siliceous sponges the original skeleton
has been replaced by calcite; but in the strata of some localities the sponges yet
retain their siliceous structure in very perfect preservation. ‘There are but few
calcareous sponges in the English Jura; but they abound in the Jurassic strata of
France and Germany, and have been referred to the genera Hudea, Peronella,
Eusiphonella, Corynella, Myrmecium, Lymnorea, Stellispongia, Sestrostomella, Blas-
tinia, Oculospongia, Crispispongia, Pharetrospongia, and Protosycon.
Cretaceous System.—Both the Siliceous and Calcareous sponges may be said to
attain their maximum development in this period. In this country there are four
well marked zones in the Cretaceous rocks, each characterized by distinctive groups
of sponges. They are (I.) the Lower Green Sand, (II.) the Upper Green Sand and
Chloritic Marl, (III.) the Chalk Marl, Gray Chalk, and Lower Chalk, and (IV.)
the Upper Chalk, including the Maestricht Chalk.
(1.) Lower Green Sand.—Very few recognizable siliceous sponges occur in this
division ; but in sandstone strata at Haslemere, in Surrey, there are thin beds nearly
entirely composed of the spicules of 'Tetractinellid sponges, with an admixture of a
few Hexactinellid and Lithistid forms; some of these latter are sufficiently charac-
teristic to be referred to the genus Mastosia. Similar beds of sponge-spicules also
occur in the Hils sandstone in Westphalia. On the other hand, the calcareous
sponges are very numerously represented in this series, not only in this country, but
in France, Germany, and Switzerland. The gravels of Farringdon in Berkshire, and
* Spongien, Radiolarien und Foraminiferen der unterliassischen Schichten vom Schafberg. Wien, 1882.
cz
12 INTRODUCTION.
at Upware in Cambridgeshire, are filled with these sponges. The minute structural ©
characters of the sponges from these beds has only been partially preserved; but
traces of the spicules can usually be distinguished in thin microscopic sections. ‘The
following genera are represented—Verticillites, Peronella, Elasmocelia, Conocelia,
Corynella, Synopella, Oculospongia, Elasmostoma, and Pharetrospongia.
(1L.) Upper Green Sand and Chloritic Marl.—In beds of this series at Warminster
in Wiltshire, Blackdown and Haldon in Devonshire, the Isle of Wight, near Folke-
stone, and near Havre in France, sponges are very numerous. In some localities
the beds are largely made up of spicules of Monactinellid and Tetractinellid sponges
with an admixture of Lithistid and Hexactinellid spicules, and resemble those
already mentioned from the Lower Green Sand. These detached spicules have been
described and figured by Mr. Carter * and Mr. Parfitt. The Lithistid sponges of
this series are particularly abundant, and, as a rule, retain their siliceous composition ;
but as they are usually preserved in a very hard siliceous matrix, it is only by
preparing sections that the structure can be distinguished. The genera of Lithistids
comprise Chenendopora, Jereica, Doryderma, Holodictyon, Pachypoterion, Nematinion,
Carterella, Phymatella, Trachysycon, Siphonia, Hallirhoa, Jerea, Polyjerea, Kalpi-
nella, and Rhopalospongia. The Hexactinellids are less numerous in proportion to
the Lithistids, and belong to the genera Craticularia, Stauronema, Sestrodictyon,
Guettardia, Eubrochus, Plocoscyphia, and Sclerokalia. Calcisponges are numerous
in some localities: the most important place for them is Essen, in Prussia; they
are also present at Warminster and near Havre. ‘They mostly belong to the same
genera as those in the Lower Green Sand; but the majority of the species are
different.
(II1.) Chalk Marl, Gray Chalk, and Lower Chalk.—Beds of this series near Dover,
Folkestone, and in the Isle of Wight contain a well-marked group of sponges, for
the most part of the same genera as those in the underlying Upper Green Sand and
Chloritic Marl, but of different species. The siliceous skeleton of these sponges has
been replaced by calcite, and their present condition is very unfavourable for deter-
mination. ‘The sponges from the Chalk Marl of the Isle of Wight, however, remain
siliceous, and can be partially cleaned from the matrix by acid. The Lithistids of
this series of beds are included in the genera Stachyspongia, Phymatella, Siphonia,
Jerea, Nelumbia, Polyjerea, and Thamnospongia. ‘The Hexactinellids associated in
the same beds with the Lithistids belong to the genera Craticularia, Strephinia,
Verrucocelia, Stauronema, Leptophragma, Guettardia, Sestrocladia, Ophrystoma,
and Plocoscyphia. 1 do not know of any Calcisponges from this division.
(1V.) Upper Chalk, including Maestricht Chalk.—Siliceous sponges abound in this
division of the Cretaceous system. In this country they are of most frequent
* Annals and Mag. Nat. Hist. ser. 4, vol. vil. p. 112.
+ Transactions of the Devonshire Association, 1870.
INTRODUCTION. 13
occurrence in the Chalk near Flamborough Head in Yorkshire, and in the southern
and south-western counties. The strata of this age in Westphalia, Hanover, and
Brunswick contain large numbers of sponges, which retain their original structure
in most beautiful preservation. On the other hand, the Upper Chalk sponges of
this country are in a very unfavourable condition of preservation, and it is only in
those which have been preserved in the interior of flints that the spicular structure
can be satisfactorily ascertained. In the Flamborough forms, as already mentioned,
the siliceous spicules are fused together, whilst in the sponges from Surrey, Kent,
Sussex, and elsewhere to the South of England, the siliceous skeletons are either
replaced by peroxide of iron or completely dissolved away. The following genera
of Monactinellids are present in this division—Leniera, Scolioraphis, Dirrhopalum.
Acanthoraphis, Opetionella, and Clona. The Tetractinellid sponges have been
referred to the genera Ophiraphidites, Tethyopsis, Stelletta, Geodia, Thenea, and
Pachastrella. The Lithistid sponges include the genera Bolidium, Chonella, Selis-
cothon, Verruculina, Stichophyma, Jereica, Scytalia, Stachyspongia, Pachinion, Dory-
derma, Heterostinia, Carterella, Isoraphinia, Phymatella, Aulaxinia, Callopegma,
Trachysycon, Siphonia, Jerea, Nelumbia, Bolospongia, Thecosiphonia, Calymmatina,
Turonia, Thamnospongia, Pholidocladia, Ragadinia, Plinthosella, and Phymaplectia.
The genera of Hexactinellids comprise Craticularia, Verrucocelia, Leptophragma,
Pleurostoma, Guettardia, Coscinopora, Aphrocallistes, Ventriculites, Schizorhabdus,
Rhizopoterion, Sporadoscinia, Polyblastidium, Cephalites, Porospongia, Cincliderma,
Plocoscyphia, Tremabolites, Etheridgia, Toulminia, Camerospongia, Cystispongia,
Callodictyon, Porochonia, Becksia, Diplodictyon, Coloptychium, Stauractinella, and
Hyalostelia.
If the siliceous sponges are exceedingly numerous in the Upper Chalk, the
calcareous forms, on the other hand, are comparatively rare. In the South of
England examples of the genera Elasmostoma and Pharetrospongia are not uncommon
in the Chalk and in flints; and in the Chalk of Maestricht there are species of
Synopella, Oculospongia, and Elasmostoma.
Eocene System.—Numerous detached spicules of Monactinellid, Tetractinellid,
Lithistid, and Hexactinellid sponges have been discovered by Rutot * in the Eocene
Sandstone of Brussels.
Miocene System.—Lithistid and Hexactinellid sponges have been discovered by
Pomel* in strata of this age in the Algerian province of Oran, and by Manzoni in
beds of Molasse sandstone in the provinces of Bologna and Modena in Italy. ‘The
numerous genera which Pomel has constituted on the characters of the canal-
* Annales de la Société Malacologique de Belgique, tome ix. 1874; and Carter, Ann, and Mag. Nat. Hist.
ser. 4, vol. xix. p. 382.
+ Description des Animaux fossiles de la Province d’Oran, 5° fascicule, 1872.
+ La struttura microscopica delle Spugne silicee del Miocene medio, 1882.
14 INTRODUCTION.
structure are quite valueless. Manzoni has referred the Lithistids discovered by
him to Astrocladia, Siphonia, Jerea, and Chenendopora, and the Hexactinellids to
the genus Craticularia. Some of these Miocene sponges retain their siliceous
structure ; in others it has been replaced by calcite.
CLASSIFICATION.
To the obliteration and alteration produced by fossilization in the structure of
fossil sponges may be attributed the various conflicting opinions which have been
held respecting their characters and the relationship between them and existing
forms. It is unnecessary here to refer in detail to these opinions, or to the many
ineffectual attempts which have been made to bring the numerous fossil examples of
the class into a natural arrangement; it may suffice to state that all the systems of
classification, based on differences of external form, or the disposition of the internal
canals of the sponge, have proved utterly valueless, and the heterogenous forms
grouped together by the advocates of these systems clearly showed their artificial
character. The honour of discovering a natural principle of classification for fossil
sponges, and thus introducing order where previously a discreditable chaos prevailed,
is due to Professor Zittel, who adopted, as a primary basis of classification, the
characters of the minute spicular bodies of which the sponge-skeleton is composed.
As the result of a thorough microscopic research into the minute structure of most
of the known fossil sponges, Zittel brought these forms for the first time into a
definite systematic position ; so that students of this group have now no difficulty in
ascertaining the relative affinity of any specimen which retains even but slight traces
of its original structure. The first step in arranging a series of fossil sponges in
natural order is to ascertain the characters of the spicular skeleton; and as in the
majority of examples no spicular structure is preserved on the outer surface, it is
necessary to make a section through the sponge in order to discover, if possible, any
indications of structure in the interior. It sometimes happens that all traces of the
spicular skeleton have disappeared throughout the central portions of the sponge as
well as on the outer surface; and in this case the systematic position of the sponge
remains somewhat conjectural. But even when all structure has disappeared from
the sponges of certain horizons and localities, we oftentimes find the same sponges
from the corresponding strata in other places with their skeletal structures in good
preservation. Owing to this fact it is possible to ascertain the original character of
many of the sponges from the Upper Chalk of Flamborough and the southern
counties of England, in which merely the outer form and canal-structure is retained,
by comparing them with the sponges from the same geological horizon in Northern
Germany, in which the spicular skeleton remains intact.
The various modifications of the canal-system rank next in importance to the
spicular skeleton in affording characters for the minor subdivisions of the sponges ;
INTRODUCTION. 15
and, as a rule, these features are more frequently recognizable in fossil examples than
the spicular structures.
The following is a list of the different Orders, Families, and Genera in which the
fossil sponges referred to in the following pages are arranged. Genera of existing
sponges are only included when represented by fossil species.
Class SPONGLA.
Division I. SILICEOUS SPONGES.
Order MONACTINELLID2.
Climacospongia, Hinde.
Lasiocladia, Hinde.
Reniera, O. Schmidt.
Dirrhopalum, Ridley.
Acanthoraphis, Hinde.
Spongilla, Lamarck.
Cliona, Grant.
Order TETRACTINELLIDE.
Ophiraphidites, Carter.
Tethyopsis, Zittel.
Stelletta, O. Schmidt.
Geodia, Lamarck.
Thenea, Gray.
Pachasirella, O. Sch.
Order LItHISTID®.
Family Rutzomorina.
Cnemidiastrum, Zittel.
Corallidium, Quenst.
Hyalotragos, Zitt.
Pyrgochonia, Goldf.
Leiodorella, Zitt.
Platychonia, Zitt.
Bolidium, Zitt.
Chonella, Zitt.
Seliscothon, Zitt.
Chenendopora, Lam.
Verruculina, Zitt.
Stichophyma, Pomel.
Jereica, Zitt.
Seytalia, Zitz.
Stachyspongia, Zitt.
Pachinion, Zitt.
Family Mucamorina.
Placonella, Hinde.
Doryderma, Zitt.
Holodictyon, Hinde.
Pachypoterion, Hinde.
Heterostinia, Zitt.
Nematinion, Hinde.
Carterella, Zitt.
Isoraphinia, Zitt.
Family ANoMOcLADINA.
Cylindrophyma, Zitt.
Melonella, Zitt.
Lecanella, Zitt.
Mastosia, Zitt.
Hindia, Duncan.
Family Terraciapina.
Aulocopium, Oswald.
Phymatella, Zitt.
Aulaxinia, Zitt.
Callopegma, Zitt.
Trachysycon, Zitt.
Siphonia, Goldfuss.
Hallirhoa, Lame.
Jerea, Lame.
Nelumbia, Pomel.
Polyjerea, From.
Bolospongia, Hinde.
16
Thecosiphonia, Ziit.
Calymmatina, Zit.
Turonia, Mich.
Kalpinella, Hinde.
Thamnospongia, Hinde.
Pholidocladia, Hinde.
Ragadinia, Zit.
Plinthosella, Zt.
Phymaplectia, Hinde.
Rhopalospongia, Hinde.
Order HEXACTINELLIDZ.
Suborder DictTyoNniINa.
Family AstyLosponcip#&.
Astylospongia, emer.
Palezomanon, Remer.
Family Evreripe.
Tremadictyon, Ziét.
Craticularia, Z7tt.
Sphenaulax, Zit.
Sporadopyle, Zt.
Strephinia, Hinde.
Verrucoceelia, Htallon.
Stauronema, Sollas.
Sestrodictyon, Hinde.
Brachiospongia, Marsh.
Family Coscinororip&.
Leptophragma, Zitt.
Pleurostoma, Rem.
Guettardia, Mich.
Coscinopora, Goldf.
Family Mexxirionip2.
Aphrocallistes, Gray.
Family VenrRICcULITID2.
Pachyteichisma, Zit.
Trochobolus, Zitt.
Phlyctenium, Ziét.
Ventriculites, Mantell.
Schizorhabdus, Zit.
Rhizopoterion, Zitt.
Sporadoscinia, Pom.
INTRODUCTION.
Sestrocladia, Hinde.
Polyblastidium, Zitt.
Cephalites, Toulmin Smith.
Family SravropeRMID2.
Cypellia, Pom.
Stauroderma, Zitt.
Purisiphonia, Bowd.
Porocypellia, Pom.
Casearia, Quenst.
Porospongia, D’ Orbigny.
Ophrystoma, Zitt.
Cincliderma, Hinde.
Protospongia, Salter.
Dictyophyton, Hail.
Eubrochus, Sollas.
Family MranprosPoncip2#.
Plocoseyphia, Reuss.
Tremabolites, Zitt.
Etheridgia, Tate.
Toulminia, Zitt.
Camerospongia, D’ Orbigny.
Cystispongia, Ramer.
Family Catnopictyonip2&.
Callodictyon, Zitt.
Porochonia, Hinde.
Becksia, Schliiter.
Diplodictyon, Zitt.
Sclerokalia, Hinde.
Family Cwnorrycuipz.
Celoptychium, Goldf.
Suborder Lyssaxina, Zitt.
Family Monaxip, Marshall.
Astreospongia, Remer.
Stauractinella, Zt.
Family Potztakip#z, Marshall.
Hyalostelia, Zt.
Holasterella, Cart.
INCERT# SEDIS.
Amphispongia, Salter.
INTRODUCTION. G7
Division Il. CALCAREOUS SPONGES.
Order Catcispone1a, Blainv.
Family Puarerrones, Zitt.
Eudea, Lamez.
Colospongia, Laude.
Verticillites, Defrance.
Celyphia, Pomel.
Himatella, Zitt.
Peronella, Zitt.
Elasmoceelia, Rem.
Conoceelia, Zitt.
Eusiphonella, Zitt.
Corynella, Zitt.
Myrmecium, Goldf.
Lymnorea, Lamz.
Stellispongia, D’ Orig.
Sestrostomella, Zitt.
Blastinia, Zitt.
Synopella, Zitz.
Oculospongia, From.
Crispispongia, Quenst.
Elasmostoma, From.
Pharetrospongia, Sollas.
Family Syconrs, Haeckel.
Protosycon, Zitz.
18
Class SPONGLA.
Division I. SILICEOUS SPONGES.
Order MONACTINELLIDA, Zttéel.
Genus CLIMACOSPONGIA, Hinde, gen. nov.
Sponges subglobate, sessile, composed of elongate acerate spicules, which radiate
upwards from the base to the circumference, and are arranged so as form a closely
disposed series of radiating canals, which open at the surface. There are also acerate
spicules disposed horizontally so as to cross the vertical spicules at right angles, thus
forming an open tissue with rectangular interspaces.
The only fossil sponge with which this genus can be compared is the Pulvillus
Thomsonii, Carter (Ann. & Mag. Nat. Hist. ser. 5, vol. i. p. 137, t. x. f. 1-6), which
is also composed of long acerate spicules; but these do not appear to have the same
arrangement, or to be crossed horizontally by other spicules, as in the present genus.
The figure given by Carter of the vertical section of Pulvillus differs altogether from
the vertical section of Climacospongia.
CLIMACOSPONGIA RADIATA, Hinde, sp. nov. (Plate I. figs. 1, 1a.)
The only examples of this sponge in the Museum are portions of two individuals
which have been fractured in a vertical direction. The sponges are from 30 to
40 millim. in diameter. They are preserved in a silicified matrix; the exterior is
rough and weathered, so as to show the canal-apertures only in a few places. In
the vertical section the radiating spicules are in part siliceous, in part replaced by a
reddish earthy material, probably iron peroxide; the transverse spicules are mostly
replaced by the peroxide, or shown by the impressions in the matrix. The canals
are either subangular or circular in section, and 0°75 to 1°5 mm. in width. ‘They
are formed and bounded by long acerate spicules, disposed vertically, sometimes in
a single series; sometimes two or three spicules are side by side. ‘The ends of the
spicules overlap each other, but they do not appear in any way to be attached
together. ‘The vertical spicules are straight, or occasionally slightly incurved, and
nearly cylindrical. Near the extremities they taper very gradually. ‘Their surfaces
appear to be smooth. The longest measured is 5°5 mm. ‘The spicules of the
transverse series are not clearly shown in a vertical section; they appear to be of the
same character as the vertical spicules, and they cross these latter nearly at right
CLIMACOSPONGIA.—RENIERA. 19
angles; but their arrangement seems to be irregular, as in some places they are
nearly in juxtaposition, whilst in others they are 1 mm. apart.
The appearance of a vertical section of one of these sponges might, at first sight,
be easily mistaken for that of a tabulate coral, the vertical spicules representing the
walls of the coral, and the transverse spicules the tabule. The preservation of the
form of this sponge is very remarkable, as there is no indication that the spicules
were held together otherwise than by the sarcode. It is possible, however, that
other and smaller spicules may have been present; for the large acerates do not
completely fill the apertures in the silicified matrix, and some of the reddish material
present may be derived from the solution and replacement of smaller spicules.
Distribution. Silurian: Perry County, Tennessee. Judging from the mineral con-
dition of the specimens, they seem to have been derived from the same strata of the
Niagara series which have yielded numerous specimens of Hexactinellid sponges
belonging to the genera Astylospongia, Astrwospongia, and Paleomanon; and it is
interesting to discover a true Monactinellid sponge in the same low geological
horizon with these forms.
Genus LASIOCLADIA, Hinde, gen. nov.
LastocLaDia comPRessa, [Hinde, sp. nov. (Plate I. fig. 2.)
The single example of this sponge is of an elongate compressed form, 36 mm.
in length by 12 mm. in width, and appears to have formed part of a branching
sponge. This fragment is composed of stout, straight, acerate, fusiform spicules,
pointed at both ends, which are loosely arranged together in a generally upward
and outward direction. Here and there sheaves of spicules project very prominently
outwards. ‘The spicules appear to be generally equal in size; the longest measured
is 5 mm. by °25 in width.
The specimen is preserved on the surface of a fragment of olive-green shale. A
few spicules only now remain; and these are in the condition of crystalline calcite.
The larger portion of the specimen merely shows the empty well defined moulds of
the spicules in the shale, which, in the space occupied by the sponge, is of the rusty
tint usually present where siliceous spicules have been dissolved.
Distribution. Lower Devonian: Jemelle, Belgium.
Genus RENIERA, 0. Schmidt, 1862.
Reniera? Carteri, Hinde. (Plate I. fig. 8.)
1879. Spicule ? of a Renierid Sponge, Carter, Aun. & Mag. Nat. Hist. ser. 5, vol. iii. p. 144,
tis Colla Jie
Spicules smooth, cylindrical, with rounded obtuse ends; for the most part with a
straight central portion, and with both ends sharply incurved, but occasionally gently
curved throughout. Average length 1 mm., width 146 mm. .
D2
20 SILICEOUS SPONGES.
These spicules retain their siliceous condition; some are crypto-crystalline, others
are distinctly crystalline. In these latter the central canal is occasionally preserved.
They now occur detached. Mr. Carter regards them as probably allied to existing
large-spiculed Renierida; they are very much larger than any existing forms. For
the sake of reference I have placed them provisionally under the genus Reniera, and
associated them with the name of Dr. H. J. Carter, who first described and figured
them.
Distribution. Carboniferous Limestone: Dalry, Ayrshire.
Genus DIRRHOPALUM, Aidley, 1881.
DirrHoPALuM PLANUM, Hinde, sp.
1880. Reniera, sp., Hinde, Foss. Sponge-spicules, p. 21, t. 1. f. 18, 19.
Detached spicules of a conical form, widest at the summits, which are rounded,
and gradually tapering to a pointed or blunted extremity. The interior exhibits a
relatively large canal of a conical form. The surface appears to have been smooth.
In length these spicules vary between 0°495 and 0-832 mm., and the width at the
summit from 0°135 to 0:225 mm. Abundant in the interior of flints.
I had referred these spicules to a species of Reniera ; but Mr. Ridley* has suggested
their resemblance to the peg-top spicules of Dirrhopaluwm (Plocamia) clopetarium,
O. Schmidt; and though the fossil spicules are much larger than those of the
existing species, the similarity of form may indicate a relationship, and I therefore
place them in this genus until more is known of their affinities. Spicules of a
similar form, but with tuberculated or spimous surfaces, are also present in the
Chalk, and it is at present uncertain whether they belong to the same sponge as the
smooth forms. Prof. Sollas has given to the spinous forms the name of Rhopalo-
conus tuberculatus f.
Distribution. Upper Chalk: Horstead, North of Ireland; Coesfeld, Westphalia.
Genus ACANTHORAPHIS, Hinde, gen. nov.
ACANTHORAPHIS INTERTEXTUS, Hinde, sp. nov. (Plate I. figs. 3, 3a.)
Sponge apparently of an ovoid shape; the single example is 25 mm. in length
by 16 mm. in width. The only structure preserved is a thin delicate surface-
tissue with subangular apertures, about 0°75 mm. wide, and porous interspaces
between them. This tissue is formed by a layer of straight spicules, superposed over
each other in an irregular manner, but not apparently in any way attached to each
other. The spicules are fusiform, gradually tapering from the centre to each end ;
their surfaces are covered with minute blunted spines. They vary somewhat in
* Journ. Linn. Soc. vol. xv. p. 487. f+ Spong. Atlan. Gebiet. p. 63, t. 4. f. 18.
+t Ann. & Mag. Nat. Hist. ser. 5, vol. vi. p. 392.
ACANTHORAPHIS.—CLIONA. 21
dimensions; a fairly large spicule measures 2°25 mm. in length by 0°975 mm. in
width. The spicules are now composed of iron peroxide.
I am unable to determine whether this delicate film of large spinous spicules con-
stituted the entire skeleton of the sponge, or whether it formed merely the exterior
layer of a sponge whose interior skeleton has disappeared. Sufficient remains to
show that the form is entirely distinct from any hitherto known. The only specimen
is imbedded in soft chalk ; it formed part of the Bowerbank collection.
Distribution. Upper Chalk: Shortlands, Kent.
Genus SPONGILLA, Zam., 1816.
SPONGILLA PURBECKENSIS, Young. (Plate I. fig. 9.)
1878. Spongilla purbeckensis, Young, Geological Magazine, n.s. vol. v. p. 220, figs. a, 0.
The spicules of this sponge occur in great numbers in nodules or masses of chert ;
they are slightly curved acerates, fusiform, thickest in the centre, and gradually
diminishing to the extremities. According to Mr. Young the spicules are minutely
tuberculated; but this feature is not shown in those which have come under my
notice. They vary from 0-015 to 0:0255 mm. in width; the longest measured is
0°45 mm.
Some fragments of chert are principally composed of these spicules, which are
mingled together without any apparent arrangement. The specimen in the Museum
was presented by Mr. Young.
Distribution. In freshwater limestones of the Purbeck series: Stare Cove, Dorset.
Genus CLIONA, Grant, 1826.
Cuiona cretacea, Portlock, sp.
1843. Entobia cretacea, Portlock, Geological Report, p. 360.
1808. Parkinson, Organic Remains, t. 8. f. 10.
1851. Clionites Conybeari, Morris, Ann. & Mag. Nat. Hist. vol. viii. p. 89, t. 4. f. 8, 9, 10.
The examples of this species now occur as solid, spheroidal, ovate, or depressed
elongate siliceous bodies from 1:8 to 5°5 mm. in diameter, which are connected
together by numerous stolons into small groups. ‘These bodies are usually found in
flints, partially fillnmg the cavities formerly occupied by Belemnites, the tests of
Echinoderms, and the shells of Jnoceramus. The original cavities hollowed out by
the boring sponge in these organic bodies have first been filled with silica, so as to
form solid moulds of that material; and at a later stage the calcareous material of
the shells and tests has been dissolved away, leaving the siliceous infillings intact.
These vary considerably in size, and also in their distance from each other; but some
of the variations are probably owing to the thickness and character of the shells and
22 SILICEOUS SPONGES.
tests in which they have been excavated. In no instance have the spicules of the
boring sponge been met with. The examples are numerous. The originals of the
figures in the ‘Annals,’ vol. viii. t. 4. f. 9, 10, are in the Museum collection.
Distribution. Upper Chalk: Norwich, Ventnor, Kent, Surrey. Drift of the Haldon
Hills. Miocene: Madeira.
Ciiona GLomERATA, Morris, sp.
1851. Clonites glomerata, Morris, Ann. & Mag. Nat. Hist. vol. viii. p. 89, t. 4. f. 11.
This species consists of a single subglobular chamber, about 8°75 mm. in
diameter, which has been excavated in the guard of a Belemnite. This chamber is
connected with the exterior by two canals, about 2 mm. in width each, as well as
by numerous minute thread-like stolons. The original and only specimen is in the
Museum collection.
Distribution. Upper Chalk: Norwich.
Ciiona? Mante.ul, Wetherell, sp.
1852. Clonites Mantelli, Wetherell, Ann. & Mag. Nat. Hist. vol. x. p. 354, t. 5c. f. 1, 2.
This species is founded on small ovoid cavities in the shell of Jnoceramus. The
cavities have been infilled with silica; and by the dissolution of the shell they now
remain as solid bodies, of an ovoid form, 1:25 mm. in length and 1 mm. in
depth, disposed either irregularly or in concentric rows following the lines of growth
of the shell. There is no indication of any constriction at the aperture of the
cavities. ‘The chambers are quite unconnected with each other; the threads of flint
which Mr. Wetherell supposed to have been connecting stolons, are in reality nothing
more than fibres of silica which have infilled cracks in the shell of the Jnoceramus.
The form of the cavities and their isolation from each other make it very doubtful
whether they have been formed by a boring sponge. The original specimens are in
the Museum.
Distribution. Upper Chalk: near Bonchurch, Isle of Wight. Drift: Haldon, near
Exeter.
CLIONA, sp.
Perforations by boring sponges in the shells of Cretaceous and Tertiary Mollusca ;
but in the absence of spicules the excavated chambers are not sufficiently characteristic
to determine the species.
Distribution. Cretaceous: Pontotoc, Mississippi. Eocene, London Clay: Barton.
Miocene: Shurm, Sinaitic Peninsula; Las Palmas, Canary Islands; Porto Praya,
Madeira. Red Crag: Walton-on-the-Naze.
OPHIRAPHIDITES. 23
Genus TALPINA, Von Hagenow, 1840.
The simple tubular borings on which Von Hagenow has constituted this genus
differ so much from all those which we know to be produced by boring sponges, that
they can hardly be regarded as the work of sponges. The examples of 7. solitaria
and 7. ramosa, Von Hagenow, figured by Prof. Morris in the Ann, & Mag. Nat. Hist.
1851, vol. viii. t. 4. f. 4, 6@, are in the Museum.
Order TETRACTINELLIDA, Marshall.
Genus OPHIRAPHIDITES, Carter, 1876.
OPHIRAPHIDITES ANASTOMANS, Hinde, n. sp. (Plate I. figs. 4, 4a.)
Sponges growing in irregularly shaped masses, composed of loosely disposed tissues
which anastomose together. The tissues are from 2 to 4 mm. in width; they are
composed of irregularly curved acerate spicules which are, as it were, loosely felted
together. Mingled with the curved spicules there are also a few straight forms;
but I have been unable to detect any trifid spiculesin the general mass. An average
large spicule is 2 mm. in length by ‘09 mm. in width; spicules of much smaller
dimensions are mingled with the larger.
There are two examples which I refer to this species. One, preserved in soft
chalk (Plate I. fig. 4), shows, in what is apparently a vertical section, the disposition
of the tissues and interspaces of the sponge: the spicules in this example are now
replaced by iron-rust ; but their forms can be pretty clearly ascertained. The other
example is preserved in the interior of a chalk flint, and exhibits a flattened, uneven
mass of spicules of the same forms as those of the chalk specimen. From this mass
the forms figured (fig. 4a) have been selected.
This species differs from the Ophiraphidites cretaceus, Zittel*, in the open locular
character of the sponge. The spicules also are generally smaller; and no trifid
spicules have been detected. Detached spicules of a similar form to those of this
species are very common in the interior of flints from the Upper Chalk of this
country, and in strata of the same age in Westphalia and Hanover; they also occur
in the Eocene of Brussels; but it is extremely rare to find them still associated
together retaining their original positions in the sponge.
It seems somewhat anomalous to include this species in the Tetractinellide when
no four-rayed spicules have been detected in it; and I can only justify placing it here
from the close resemblance of its curved acerates to those of O. cretaceus, Zitt., in
which trifid spicules are also present, though rarely; and it seems to me not impro-
bable that with more perfect examples they would be found in this form as well.
Distribution. Upper Chalk: South of England.
* Studien, III Ab. p. 8, t. xi. f. 2a, b,c, d.
24 SILICEOUS SPONGES.
Genus TETHYOPSIS, Zitte/, 1878.
TETHYOPSIS CRETACEUS, Hinde, n. sp. (Plate I. figs. 5, 5a.)
Sponge either compressed or growing in small upright masses of irregular outline,
with interpenetrating canals and passages. The sponge-tissues are composed of
straight, smooth, acerate spicules, pointed at both ends, and from 2 to 3 mm. in
length. Mingled with the acerates are a few simple trifid spicules, with short
slightly recurved head-rays. ‘The spicules are disposed generally parallel with each
other in close juxtaposition. The surface-characters of the sponge are not preserved.
One example of this species is preserved in the interior of a flint, and displays very
clearly the form and arrangement of the spicules. Another specimen is a flattened
mass 110 mm. in length, 80 mm. in width, and 16 mm. in thickness, which has
been partially dissolved out of the Flamboro’ Chalk. This mass appears to be
entirely composed of spicules, which, however, are so altered by fossilization as to be
scarcely distinguishable.
From Tethyopsis Steinmanni, Zitt.*, the only other species of the genus, this form
differs in the smaller dimensions of the acerate spicules, and the shorter head-rays of
trifid forms.
Distribution. Upper Chalk: Flamboro’, Yorkshire ; South of England.
Genus STELLETTA, 0. Schmidt, 1866.
STELLETTA INcLUSA, Hinde, n. sp. (Plate I. figs. 6, 6 a.)
The only example of this species is preserved in the interior of a chalk flint, and
appears to have been originally hemispherical in form. The section shown is
61 mm. in length by 21 mm. in height. The sponge is composed of spicules in
close juxtaposition, parallel with each other, and disposed with an outward radial
direction.
There are three kinds of spicules present:—Ist. Straight, elongated acerates,
nearly cylindrical throughout: these do not appear to be very numerous. 2nd.
Robust trifid spicules with a straight shaft, which gradually tapers from the head to
the pointed extremity; the summit is somewhat flattened, the head-rays are short,
simple, somewhat sharply recurved, and with pointed ends. These spicules vary
between 2 and 4 mm. in length; an average specimen (2°475 mm. long) has the
shaft 0:15 mm. wide, and measures across the head-rays 0°412 mm. The main
portion of the sponge is composed of these trifid spicules, which are disposed in a
radial direction with the heads towards the exterior. 3rd. Anchor-shaped trifid
spicules, with a conical bullet-shaped head, and an elongated cylindrical or very
gradually tapering shaft. I have not obtained one of these spicules with an entire
* Studien, III Ab. p. 9, t. xi. £. 3.
STELLETTA.—THENEA. 25
shaft, so that I do not know their length. These anchor-shaped spicules appear to
be rare in proportion to the other trifids. The surface of the sponge has not been
preserved, so that it is doubtful whether it was furnished with disks or stellates.
The only specimen is from Sir P. Egerton’s collection.
Distribution. Upper Chalk: England.
Genus GEODIA, Zam. 1816.
Grop1a? cLavata, [Zinde.
Geodia? clavata, Hinde, Foss. Sponge Sp. p. 29, t. 2. f. 1-5.
Relatively large, straight, trifid spicules, with short rounded knob-shaped head-
rays, either simple or compressed. ‘The shaft constricted at the neck, and imme-
diately below slightly bulbous. Length varying from 7 to nearly 9 mm.; greatest
width of shaft 0°585 mm. These spicules have only been found detached in the
interior of flints.
Distribution. Upper Chalk: Horstead, Norfolk.
Geop1a? coronata, Hinde.
Geodia coronata, Hinde, Foss. Sp. Sp. p. 31, t. 2. f. 6-8.
Trifid spicules with small upright head-rays; the shaft is swollen at the summit,
and gradually tapers to the extremity. Average length 4 mm.; width of shaft
0-45 mm. ‘These spicules have only been met with detached in the interior
of flints.
Distribution. Upper Chalk: Horstead, Norfolk; Kent.
Geropia? Wricuttl, Hinde.
1880. Geodia? Wrightii, Hinde, Foss. Sp. Sp. p. 31, t. 2. f. 12.
Trifid spicules with blunted head-rays projecting forwards from the head of the
shaft. Rays and shaft with ring-shaped expansions. Average length 2 mm.; width
0-45 mm. At present they have only been found detached in the interior of flints.
Distribution. Lower Green Sand: Haslemere, Surrey. Upper Chalk: Horstead,
Norfolk; South of England; near Belfast (Wright); Coesfeld, Westphalia.
Genus THENEA, Gray, 1867.
THENEA, sp.
1880. Tisiphonia? sp., Hinde, Foss. Sp. Sp. p. 43, t. 8. f. 16-23. '
Trifid spicules with widely expanded, usually compound or bifurcate head-rays,
which are extended nearly at right angles to the shaft. The shaft varies greatly. in
length in different specimens; in some instances it is reduced to a mere rounded
E
26 SILICEOUS SPONGES.
central prominence. These spicules have only been found detached ; it is probable
that they were zone-spicules of one or more species of sponge allied to Thenea.
Distribution. Upper Chalk: Horstead, Norfolk ; Kent.
Genus PACHASTRELLA, O. Schmidt, 1868.
PACHASTRELLA PRIMAVA, Zittel.
1878. Pachastrella primeva, Zittel, Studien, III Ab. p. 9, t. xi. f. 4a, d.
Microscopic slides with detached spicules of this species. Prof. Zittel’s collection.
Distribution. Upper Chalk: Ahlten; Hanover.
PACHASTRELLA CoNVOLUTA, Hinde, n.sp. (Plate II. figs. 1, 1 a.)
Sponge growing in plate-like expansions of various forms and dimensions. It is
either fan- or ear-shaped, with rounded incurved margins, or folded so that the lateral
margins unite to become vasiform, or the walls are irregularly convolute. Some
examples are attached to the surface of other sponges; in others there is no indication
of any process by which they were fixed, and these forms may have been free. A
large specimen is 110 mm. in length by 100 mm. in width. The walls vary from
7 to 12 mm. in thickness. The surface of the sponge is rough, and frequently
uneven. The walls are in places penetrated by tubular apertures; but these are
very irregular in size and direction, and appear to be owing rather to extraneous
causes than to be of the nature of canals belonging to the sponge.
The walls are entirely composed of quadrifid spicules of various dimensions, loosely
mingled together, apparently without definite arrangement, and only held in position
by the interlacing of their rays with each other. The spicules of the interior of the
walls are only faintly recognizable; but those of the outer and inner surfaces are
better preserved, though even these are considerably altered by fossilization. The
rays of the spicules are robust, and apparently obtusely pointed. The length of an
arm of what appears to be an average spicule is 0°75 mm.
This species may be distinguished from Pachastrella primeva, Zitt., by its mode of
growth, and also by the dimensions of the spicules, which, so far as I can ascertain,
do not reach the size of the larger forms in Zittel’s species. In the general form
of the spicules, however, and in their disposition this species corresponds with
P. primeva, and with the existing forms of the genus, P. abyssi, O. Schmidt, and
P. interteata, Carter*. Detached spicules, apparently resembling those composing
this species in form and size, occur in hollow flints at Horstead ; but all the examples
in which the form of the sponge is retained are from Flamborough, and appear to be
not uncommon in the chalk of that locality.
Distribution. Upper Chalk: Flamborough, Yorkshire.
* Ann. & Mag. Nat. Hist. ser. 4, vol. xviii. p. 409, t. xv. f. 41.
PACHASTRELLA, 27
PACHASTRELLA PLANA, Hinde, n. sp. (Plate I. figs. 7, 7a.)
The only example of this species is contained in the interior of a flint, and is a
fragment of the wall of a sponge which appears to have been palmate in form. The
wall itself is about 3 mm. in thickness, the margins are attenuated. It is composed
of spicules loosely disposed over each other without being attached together. The
spicules appear for the most part to be only three-rayed; occasionally, however, a
fourth ray is represented by a very short process. ‘lhe rays of the spicules are
straight or slightly arched, nearly in a horizontal plane, and they spring from the
centre at nearly equal angles. Two of the three horizontal arms are not infrequently
longer than the third. The arms or rays are nearly cylindrical, their ends apparently
rounded. The arms vary from 0:6 mm. to nearly 1 mm. in length. ‘The spicules are
arranged parallel to the surface of the sponge-wall, so that there are small inter-
spaces between the rays.
The form and arrangement of the spicules readily distinguish this from the other
described species of Pachastrella. Similar spicules to those composing the walls
of this species are present detached in the Horstead flint. Foss. Sp. Sp. p. 48,
tii taal,
Distribution. Upper Chalk: Upware, Cambridgeshire.
Fraqments of Flint, Chert, and other Rocks composed of Sponge-spicules.
(Plate I. figs. 10, 11, 12.)
Flint or chert from the Portland Oolite of Upware near Weymouth, containing
trifid spicules. Fig. 10 is a magnified representation of a thin microscopic section.
Thin bands of rock from 25 to 75 mm. each in thickness, from the Lower Green
Sand near Haslemere in Surrey, are nearly entirely composed of sponge-spicules,
which form a compact mass now cemented together in a siliceous matrix. When
exposed to the weather the spicules stand out distinctly from the matrix. The
spicules are mostly acerates and trifids; a few forms of Lithistid spicules are also
present. Fig. 11 is a magnified representation of a minute portion of the weathered
surface of one of these sponge-beds. Nodules of rock from the Lower Green Sand
of Folkestone and from Badbury Hill near Farringdon are similarly made up of
spicules.
Bands of chert in the Upper Green Sands near Ventnor, Isle of Wight, are filled
with spicules, which, like those from the Lower Green Sand, are principally acerates
and trifids. Fig. 12 is a representation of a thin microscopic section of this chert.
Beds and layers of spicules also occur in strata of this age at Blackdown, and in the
Haldon Hills near Exeter. ;
Flints from the Upper Chalk sometimes exhibit spicules imbedded in the siliceous
mass; but more frequently the spicules occur infilling cavities in the interior of the
E 2
hl
28 SILICEOUS SPONGES.
flint nodules. A great variety of forms of trifid and acerate spicules are found
commingled together. Rarely, too, in the chalk itself similar spicules are found
grouped together; but in this case the spicules are replaced by iron peroxide.
Specimens from Norfolk and the south of England.
The spicules which thus occur in enormous numbers in the Oolite, the Lower and
Upper Green Sand, and the Upper Chalk are principally of Tetractinellid sponges,
and resemble those of the existing genera Geodia, Tethya, Stelletta, &c.; but their
forms are not sufficiently characteristic to allow a generic or specific determination
to be made from them.
Order LITHISTID &, O. Schmidt.
Family RHIZOMORINA, Zittel.
Genus CNEMIDIASTRUM, Zttel, 1878.
CNEMIDIASTRUM STELLATUM, Goldf. sp.
1833. Cnemidium stellatum, Goldfuss, Petref. Th. 1, p. 15, t. 6. f. 2.
1833. Cnemidium granulosum, Minst., Goldf. Petref. p. 97, t. 35. f. 7.
1878. Cnemispongia Goldfussi, Quenst. Petref. Bd. 5, p. 259, t. 126. f. 73, 74, and t. 127.
f. 1-16.
1878. Cnemidiastrum stellatum, Zitt. Studien, II Ab. p. 46, t. 3. f. 1, 2.
1879. Cnemidiastrum stellatum, Zitt. Handbuch der Paleont. Bd. 1, p. 150, f. 66.
Distribution. Upper Jura: Randen, Heuberg; Wiirtemberg.
CNEMIDIASTRUM STRIATO-PUNCTATUM, (roldf. sp.
1833. Cnemidium striato-punctatum, Goldf. Petref. Th. 1, p. 15, t. 6. f. 3.
1878. Cnemispongia Goldfussi, Quenst. p. p. Petref. Bd. 5, p. 268, t. 127. f. 19-22.
1878. Cnemidiastrum striato-punctatum, Zitt. Stud. II Ab. p. 46.
Distribution. Upper Jura: Randen, Heuberg ; Wiirtemberg.
CNEMIDIASTRUM CORALLINUM, Quenst. sp.
1858. Cnemidium corallinum, Quenst. Der Jura, p. 694, t. 84. f. 1.
1878. Cnemidium corallinum, Quenst. Petref. Bd. 5, p. 267, t. 127. f. 16-18.
1878. Cnemidiastrum corallinum, Zitt. Stud. Il Ab. p. 46.
Distribution. Upper Jura: Randen.
CNEMIDIASTRUM HoHENEGGERI, Zt.
1878. Cnemidiastrum Hoheneggeri, Zitt. Stud. II Ab. p. 46, t. 2. f. 8.
Microscopic slides with the spicules of this species, from Zittel’s collection.
Distribution. Upper Jura: Wodna, Cracow.
CNEMIDIASTRUM.—HYALOTRAGOS. 29
CNEMIDIASTRUM RIMULOSUM, Goldf. sp.
1833. Cnemidium rimulosum, Goldf. Petref. Th. 1, p. 15, t. 6. f. 4.
1878, Cnemidium rimulosum, Quenst. Petref. Bd. 5, p. 271, t. 128. f. 1-5.
1878. Tragos granulosum, Quenst. Petref. Bd. 5, p. 285, t. 129. f. 4, 5.
1878. Cnemidiastrum rimulosum, Zitt. Stud. IL Ab. p. 46, t. 3. f. 3.
Distribution. Upper Jura: Randen; Wiirtemberg.
CNEMIDIASTRUM PLURISTELLATUM, Zitt.
1878. Cnemidiastrum pluristellatum, Zitt. Stud. IL Ab. p. 46.
1833. Cnemidium stellatum, Goldf. Petref. Th. 1, p. 15, t. 30. f. 3.
1858. Cnemidium stellatum, Quenst. (non Goldf.) Jura, p. 676.
1878. Cnemidium stellatum, Quenst. Petref. Bd. 5, p. 272, t. 128. f. 6, 7.
Distribution. Upper Jura: Randen, Heuberg; Wiirtemberg.
Genus CORALLIDIUM, Zittel, 1878.
CoRALLIDIUM DICERATINUM, Qwenst. sp.
1852. Cnemidium diceratinum, Quenst. Handb. t. 61. f. 20.
1878. Cnemidium diceratinum, Quenst. Petref. Bd. 5, p. 275, t. 128. f. 10-12.
1878. Corallidium diceratinum, Zitt. Stud. IL Ab. p. 46.
Distribution. Upper Jura: Kelheim, Bavaria (Zittel’s coll.).
Genus HYALOTRAGOS, Zitte/, 1878.
HYALOTRAGOS PATELLA, Goldf. sp.
1833. Tragos patella, Goldf. Petref. Th. 1, pp. 14, 96, t. 5. f. 10, and t. 35. f. 2.
1878. Tragos patella, Quenst. Petref. Bd. 5, p. 283, t. 128. f. 26-28, and t. 129. f. 1-3.
1878. Hyalotragos patella, Zitt. Stud. IL Ab. p. 47, t. 3. f. 4, 5.
Distribution. Upper Jura: Hossingen, Heuberg ; Wiirtemberg.
TIYALOTRAGOS RADIATUM, (roldf. sp.
1833. Tragos radiatum, Goldf. Petref. Th. 1, p. 96, t. 35. f. 2.
1878. Tragos radiatum, Quenst. Petref. Bd. 5, p. 281, t. 128. f. 24, 25.
1878. Hyalotragos radiatum, Zitt. Stud. II Ab. p. 48.
Distribution. Upper Jura: Randen, Bohringen.
HYALOTRAGOS RETICULATUM, MJiinst. sp.
1833. Tragos reticulatum, Minst., Goldf. Petref. Th. 1, p. 96, t. 35. f. 5.
1878. Tragos reticulatum, Quenst. Petref. Bd. 5, p. 289, t. 129. f. 10-15.
1878. Hyalotragos reticulatum, Zitt. Stud. II Ab. p. 48.
Distribution. Upper Jura: Randen, Heuberg ; Wiirtemberg.
30 SILICEOUS SPONGES.
HYALoTRAGOS RUGOsUM, Miinst. sp.
1833. Tragos rugosum, Miinst. Petref. Th. 1, p. 96, t. 35. f. 4.
1878. Hyalotragos rugosum, Zittel, Stud. IL Ab. p. 48.
Distribution. Upper Jura: Streitberg.
HYALOTRAGOS PEZIZOIDES, Goldf. sp.
1833. Tragos pezizoides, Goldf. Petref. Th. 1, p. 18, t. 5. f. 8.
1878. Tragos fistulosum, Quenst. Petref. Bd. 5, p. 278, t. 128. f. 15-23.
1878. Hyalotragos pezizoides, Zitt. Stud. Il Ab. p. 48.
Distribution. Upper Jura: Randen, Heuberg; Wiirtemberg.
Genus PYRGOCHONIA, Zit. 1878.
PYRGOCHONIA ACETABULUM, Goldf. sp.
1833. Tragos acetabulum, Goldf. Petref. Th. 1, p. 13, t. 5. f. 9.
1833. Tragos verrucosum, Minst., Goldf. Petref. Th. 1, p. 96, t. 35. f. 6.
1878. Tragos acetabulum, Quenst. Petref. Bd. 5, p. 288, t. 129. f. 7, 8, 18.
1878. Tragos infranudatum, Quenst. ib. ib. p. 287, t. 129. f. 6.
1878. Pyrgochonia acetabulum, Zitt. Stud. II Ab. p. 48.
Instribution. Upper Jura: Randen, Heuberg.
Genus LEIODORELLA, Ztt. 1878.
LEIODORELLA EXPANSA, Zitt.
1878. Leiodorella expansa, Zitt. Stud. II Ab. p. 49, t. 2. f. 5, and t. 3. f. 11.
Distribution. Upper Jura: Wodna, Cracow (Zittel’s coll.).
Genus PLATYCHONIA, Zit. 1878.
PLATYCHONIA AURIFORMIS, Quenst. sp.
1878. Spongites auriformis, Quenst. Petref. Bd. 5, p. 319, t. 131. f. 1.
1878. Platychonia auriformis, Zitt. Stud. II Ab. p. 50, t. 3. f. 9.
Distribution. Upper Jura: Streitberg, Nattheim.
PLATYCHONIA VAGANS, Quenst. sp.
1858. Spongites vagans, Quenst. Jura, t. 82. f. 8.
1878. Platychonia vagans, Zitt. Stud. II Ab. p. 50, t. 3. f. 8.
Microscopic slides with spicules of this species (Zittel’s coll.).
Distribution. Upper Jura: Streitberg.
PLATYCHONIA.—SELISCOTHON.
(St)
an
PLATYCHONIA, sp.
Distribution. Upper Jura: Wurgau, Franconia (Zittel’s coll.).
Genus BOLIDIUM, Zitt. 1878.
BouipiuM PALMATUM, Reemer, sp.
1864. Amorphospongia palmata, Reemer, Paleont. Bd. 13, p. 55, t. 19. f. 8.
Microscopic slide with spicules of the species.
Distribution. Upper Chalk: Sudmerberg ; Goslar.
Genus CHONELLA, Zit. 1878.
CHONELLA TENUIS, Remer, sp.
1864. Cupulospongia tenuis, Roemer, Paleeont. Bd. 13, p. 51, t. 17. f. 7.
1878. Chonella tenuis, Zitt. Stud. II Ab. p. 52, t. 3. f. 6, 7.
Distribution. Upper Chalk: Sudmerberg, Biewende; Brunswick.
CHONELLA AURIFORMS, Remer, sp.
1840. Achilleum auriformis, ¥. A. Roemer, Nordd. Kreide. p. 2, t. 1. f. 3.
1878. Chonella auriformis, Zitt. Stud. II Ab. p. 52.
Distribution. Neocomian: Berklingen, Brunswick.
Genus SELISCOTHON, Zité. 1878.
SELISCOTHON PLANUS, Phillips, sp. (Plate II. figs. 2-4.)
1835. Spongia plana, Phill. Geol. of Yorkshire, p. 177, t. 1. f. 1.
1835. Spongia capitata, Phill. ib. p.177, t. 1. £. 2.
1878. Seliscothon planus et capitatus, Zitt. Stud. LI Ab. p. 54.
Sponge similar in form to an expanded mushroom. The body or upper portion is
plate-like, circular in outline, usually with a slight central depression immediately
above the stem. ‘The upper surface sometimes shelves gradually from the margin to
the centre ; more frequently, however, it is nearly flat or even slightly convex. The
margins are usually oblique, and form nearly a right angle with the under surface.
The lower surface is flat or slightly concave. The stem is inversely conical in form,
and gradually tapers to a blunted extremity. ‘There are no indications of its attach-
ment to a foreign body. It varies in length in different examples; an average
specimen measures 55 mm. Small examples of the species are not more than
27 mm. in width, whilst large forms reach up to 190 mm. ‘The thickness of the
body-plate varies between 10 and 14 mm.
52 SILICEOUS SPONGES.
‘The upper surface of the body is thickly covered as far as the marginal edge with
circular apertures ‘75 to 1 mm. in width; in specimens treated with acid these
openings appear to be oblique to the plane of the surface. ‘The under surface of the
body in the best-preserved specimens seems to be composed of a close dermal layer,
which, however, is usually absent, and then the vertical lamelle are exposed.
The sponge itself is composed of a series of delicate fibrous lamelle extending
from the centre to the circumference, having a general resemblance to the septa of
corals. These lamelle are about 0-4 mm. in thickness and about 1 mm. apart.
Numerous interstitial fibres connect the radial lamelle, so that the aspect of a
vertical section of the sponge-wall is that of a labyrinthine fibrous web. In all the
specimens the spicules forming the fibres have been so fused together that the
individual forms cannot be detected.
As no description whatever accompanies the figures of the sponges from the
Flamborough Chalk given by Phillips in the ‘Geology of Yorkshire,’ and as the
figures are very imperfectly drawn, it is often a matter of great difficulty to deter-
mine the forms which they are supposed to represent. Judging by the represen-
tation of Spongia capitata, plate 1. fig. 2, it would appear to possess but slight
relationship with the Spongia plana (fig. 1). In reality, however, the only difference
between the sponges thus designated consists in the outer form; the summit of
S. capitata has not been developed to the same extent as that of S. plana. The
examples in the Museum collection show a series of gradational forms between those
in which the summit of the sponge is scarcely more expanded than the top of the
stem, and those with very widely expanded bodies; and there can be little doubt that
Phillips’s S. capitata should be included in the same species as the S. plana.
This species seems to be the most common of any of the Flamborough sponges,
but it does not appear to have been recognized from any other locality in the
English Chalk.
Distribution. Upper Chalk: Flamborough, Yorkshire.
SELISCOTHON EXPLANATUS, Remer, sp. (Plate I. fig. 5.)
1864. Chenendopora explanata, Roemer, Palzont. Bd. 13, p. 44, t. 16. f. 3.
1878. Seliscothon explanatus, Zitt. Stud. II Ab. p. Bd, t. 4, f 2:
Body of sponge forming a flat plate-like expansion, apparently circular in outline.
No stem has been preserved. The upper surface has a few concentric, slightly
marked, rounded ridges. The margins rounded. ‘The only specimen is a portion of
a sponge, about 180 mm. in width across the summit and 5 mm. in thickness.
The upper surface is furnished with a dermal membrane in which are numerous
irregularly disposed circular apertures 0°75 mm. in width, and from two to three
diameters apart. The under surface appears to be entirely covered with the dermal
membrane.
9
SELISCOTHON.— CHENENDOPORA. By)
The vertical lamelle are 0°3 mm. in width and about 0°5 mm. apart. These
lamellz are formed of extremely delicate, elongated branching and spinous spicules,
which are interlocked together. In the Flamborough specimen these spicules are
undistinguishable ; those figured are from a specimen from Hanover.
The only example from Flamborough differs from the type of Reemer in possessing
rounded margins and a somewhat thinner plate, but resembles it in form, in the size
of the apertures of the upper surface, and in the furrowed dermal layer. From
S. planus this species is readily distinguished by its furrowed upper surface, the
smaller and less closely disposed apertures, and the more delicate character of the
vertical lamelle.
Distribution. Upper Chalk: Flamborough, Yorkshire; Ahlten, Hanover (Zttel’s
coll.).
SELISCOTHON MANTELLI, Goldf. sp.
1833. Scyphia mantelli, Goldfuss, Petref. Th. 1, p. 219, t. 65. f. 5.
1878. Scyphia mantelli, Quenst. Petref. Bd. 5, p. 375, t. 133. f. 4.
1878. Seliscothon mantel, Zitt. Stud. II Ab. p. 54, t. 4. f. 3.
Microscopic slides with spicules of this species.
Distribution. Coesfeld, Westphalia.
SELISCOTHON TESTA-FLORUM, Quenst. sp.
1878. Scyphia testa-florum, Quenst. Petref. Bd. 5, p. 377, t. 135. f. 7.
Distribution. Upper Chalk: Sudmerberg.
SELISCOTHON GIGANTEUS, Leemer, sp.
1864. Cupulospongia gigantea, Roemer, Paleont. Bd. 13, p. 51, t. 18. f. 1.
1878. Seliscothon giganteum, Zitt. Stud. II Ab. p. 54, t. 4. f. 4.
Distribution. Craie Chloritée: Vaches Noires, Havre.
SELISCOTHON, sp.
Distribution. Upper Chalk: Sudmerberg; Goslar.
Genus CHENENDOPORA, Lamz., 1821.
CHENENDOPORA FUNGIFORMIS, Lame.
1821. Chenendopora fungiformis, Lamx. Expos. méthod. des genres de l’ordre des polypiers,
p77, t. 75. £. 9; 10:
1808. Funnel-formed Alcyonite, Parkinson, Org. Rem. vol. ii. t. 11. f. 5.
1847. Chenendopora fungiformis, Michelin, Icon. Zooph. p. 130, t. 34. f. 3.
1861. Bicupula lata, Courtiller, Eponges fossiles, p. 139, t. 37. f. 1.
1878. Chenendopora fungiformis, Zitt. Stud. IL Ab. p. 55, t. 3. f. 13, 14.
4 SILICEOUS SPONGES.
(3%)
There are typical examples of this species in the Museum from the Craie of
Vaches Noires, near Havre, but only a single specimen from the Upper Green Sand
of Warminster, which I refer with some doubt to this species; for there are no
traces of the wrinkled dermal layer, and the margins of the cup are rounded, and
not flattened as in the French examples. ‘The spicular structure, however, appears
to resemble that of the typical forms; and I therefore leave it provisionally under
this species. According to Prof. Zittel the French examples are derived from the
Upper Chalk, or Senon of D’Orbigny; but I have not met with any from the same
horizon in England.
Distribution. Upper Green Sand: Warminster. Upper Chalk (Senon): Vaches
Noires, near Havre.
CHENENDOPORA MICHELINII, Hinde, sp. n. (Plate III. figs. 1, 1 a, 14.)
Sponges simple, with vasiform or cup-shaped bodies which gradually taper below
to an elongated cylindrical or compressed stem, which occasionally bifurcates in its
lower portion. The walls of the cup or vase are sometimes smooth and even, some-
times with longitudinal open folds; the margins are usually thin, rounded, and
occasionally with a slight inward or outward curve. ‘There is a considerable
variation in the thickness of the cup-walls in different specimens; the vasiform
examples range from 5 to 6 mm. in thickness, whilst the cup-shaped forms are even
9mm. thick. The stem is usually simple, but at its lower portion it either divides
into root-like extensions or becomes expanded into a hollow disk. In large speci-
mens the stem measures 260 mm. in length by 46 mm. in thickness; and the width
of the sponge at the summit varies in different examples between 55 mm. and
170 mm.
Both the exterior and interior surfaces of the cup are furnished with numerous
openings of canals about 0°85 mm. in width, and apparently similar on both surfaces.
The stems when weathered exhibit longitudinal branched canals.
The spicules are irregular branching bodies, covered with rounded tubercles, which
interlock and connect them with each other. ‘They are so intimately interwoven
together that in a thin microscopic section it is difficult to distinguish the individual
forms. Occasionally a sinuous canal can be seen in the axial line of the spicule.
The spicules are 0-042 mm. in thickness.
The sponges which I refer to this species are not uncommon in the Upper Green
Sand of Wiltshire. There is a considerable variety of form and dimensions in the
different examples; but I cannot discover any characters which would allow them to
be placed under more than a single species. ‘They seem to have been generally
referred to the Polypothecia infundibulum, Benett; but in the figure of this species
three or four cup-shaped sponges are represented as growing closely aggregated
together, whilst C. michelinii is uniformly simple. In the absence of any description
CHENENDOPORA.—VERRUCULINA. 35
of Miss Benett’s species, I am unable to compare it with the present one in other
characters.
The sponge figured in the frontispiece of the second volume of Parkinson's
‘Organic Remains’ is in the Museum collection, and appears to belong to this
species; but without making a section of it, I cannot satisfactorily determine the
point. The specimen is stated by Parkinson (/. c. p. 125) to have been found in
Wiltshire ; but its appearance differs so much from all the other examples from this
county, that it seems to me to have been brought from some other locality,
probably France.
Michelin has figured a sponge under the name of Chenendopora Parkinsonis, Icon.
Zooph. p. 131, t. 31. f. 1, which in outer form very closely resembles some of the
examples of C. Michelinii; but it differs in having numerous canal-apertures on the
flattened margins of the cup. Zittel has referred this form to the genus Margino-
spongia, D’Orbigny, and placed it in the Tetracladina family. Its spicular structure,
however, does not yet appear to have been ascertained.
Distribution. Upper Green Sand: Warminster; Vaches Noires ?
CHENENDOPORA PocILLUM, Michelin.
1847. Chenendopora pocillum, Mich. Icon. Zooph. p. 1382, t. 33. f. 5.
1861. Cupulina pocillum, Court. Epong. Foss. p. 18, t. 29. f. 1.
1861. Cupulina elata, Court. ib. p. 18, t. 29. f. 2.
1878. Chenendopora pocillum, Zitt. Stud. II Ab. p. 55.
Distribution. Craie Chloritée?: France (Bright collection).
Genus VERRUCULINA, Zitt., 1878.
Professor Zittel includes in this genus fan, palmate, or funnel-shaped sponges,
with projecting oscules on the upper or inner surface of the wall, and with small
pore-like openings on the lower or outer surface. For sponges similar in form, but
with oscules on both the upper and lower wall-surfaces, Zittel constituted the genus
Amphithelion, but at the same time acknowledges that the differences between it
and Verruculina are hardly more than of subgeneric importance. In practice,
however, I find that these differences are very difficult of application; for though in
typical forms the distinction between the oscules of the upper and the pores of the
lower surface of the sponge is sufficiently clear, there are many examples in which it
is difficult to determine whether the canal apertures of the lower surface have the
characters of pores or oscules. I therefore propose to relinquish the genus Amphi-
thelion and to include the sponges placed therein in the genus Verruculina. The
definition of this latter genus will thus have to be extended so as to embrace
F2
56 SILICEOUS SPONGES.
sponges which, in addition to the projecting oscules on the upper surface, are
furnished either with pores or oscules on the lower surface of the wall.
There are very numerous examples of this genus in the Upper Chalk of Flam-
borough and in the Middle and Upper Chalk of Germany and Bohemia. ‘They are
all similar in their mode of growth, but present distinct specific characters in the
dimensions and disposition of the canal-apertures of the upper and lower surfaces of
the wall, as well as in the thickness of the wall itself. I have found it a very
difficult task to identify any of the Flamborough examples with the figures given by
Phillips of the sponges from this locality ; for though there can be little doubt that
he has intended to represent one or more species of this genus, the figures given are
so imperfect in detail that, in the absence of any description, it is impossible to
recognize the species which they are supposed to indicate. For example, the
Spongia marginata, Phill. Geol. York. pl. i. fig. 5, belongs undoubtedly to the genus
Verruculina; but later authors, such as Reuss, Reemer, and Quenstedt, have each
relegated different forms of sponges to Phillips’s name, thus clearly showing the
insufficiency of his figure to establish the species. A somewhat similar difficulty
exists in respect to some of the species of this genus described by F. A. Roemer from
the North-German Chalk; for the descriptions are generally so meagre that they
would apply to more than one species.
It is only with reluctance that I have added four new species to the number
already included in this genus; but it seemed preferable to do this, than to place the
examples under specific names which have no definite characters assigned to them.
VERRUCULINA SERIATOPORA, Ramer, sp. (Plate III. fig. 4.)
1840. Manon seriatoporum, F. A. Roemer, Nordd. Kreide. p. 3, t. 1. f. 6.
1878. Verruculina seriatopora, Zitt. Stud. IL Ab. p. 59, t. 4. f. 1.
There is a fragment of a sponge in the collection which I refer, though not
without doubt, to the above species. The wall of the specimen is from 8 to 11 mm.
in thickness; the upper surface is furnished with projecting oscules, 2 mm. in
diameter near their bases, partly disposed in linear order. The lower surface appears
to be composed of closely interwoven fibres; but it is not sufficiently free from the
matrix to show if pores are present or not. The spicules of which the fibres of this
species are made up are minute irregularly branching bodies with spinous pro-
jections on their surfaces.
Distribution. Upper Chalk: Sudmerberg; Ahlten (Zittel’s coll.).
VERRUCULINA PLIcata, Hinde, n. sp. (Plate IV. figs. 2, 2a, 6, ¢, d.)
Sponges with plate-like walls, usually incurved, and occasionally, by the meeting
and union of the lateral margins, becoming open funnel-shaped. In a few specimens
the walls extend in a horizontal direction, and the sponge is platter-shaped. The
VERRUCULINA. 37
walls are from 6 to 9 mm. in thickness. The margins are somewhat attenuated and
rounded ; and the oscules extend close to the edge of the wall. The examples are of
various dimensions, a large specimen measuring 170 mm. in width.
The upper or inner wall-surface has numerous, irregularly disposed, slightly pro-
jecting oscules, 0°65 mm. wide, and about 2 mm. apart; the interspace between the
oscules appears to be composed of a compact membrane when perfect ; but usually
it exhibits closely arranged sinuous apertures. The lower or outer wall-surface is
roughened occasionally with shallow concentric furrows, and is traversed by thick-
set minute pores, 0°53 mm. each in width. In many examples the lower surface has
the appearance of a compact membrane ; but when treated with dilute acid the pores
become visible.
Traces of the spicules can be occasionally seen; but, as a rule, the spicular fibres
are destroyed, and the interior of the sponge-wall exhibits only a mass of porous
silica.
From Verruculina aurita, Rem. sp., Paleont. Bd. 13, p. 43, t. 16. f. 2, this
species is distinguished by the more numerous oscules on the upper surface, and the
very closely set pores on the lower, whilst the margins are not thickened as in
Reemer’s species. It differs from V. seriatoporum, Roem. sp., in the absence of a
linear arrangement and the smaller size of the oscules, and also by the pores of the
under surface. The V. Phillipsii, Reuss, sp., Boh. Ky. p. 77, t. 19. f..7, has much
stouter walls, and there no pores on the lower surface. It seems probable that
Reuss has included more than one species under V. Phillipsti ; for his fig. 9 has very
much thinner walls than fig. 7, and the oscules are not more than half the size.
V. plicata appears to have been abundant.
Distribution. Upper Chalk: Flamborough; Sudmerberg.
VERRUCULINA ASTRA, Hinde, n, sp. (Plate III. figs. 5, 5 a.)
Sponges fan-, cup-, or ear-shaped, with walls from 4:5 mm. to 6°5 mm. in thickness.
The margins rounded and of the same thickness as the wall. The specimens vary
from 40 to 70 mm. in lateral extension, and reach to 70 mm. in height.
The inner surface of the wall has irregularly disposed projecting oscules, each
about 0°7 mm. in width. From each of these oscules sinuous canals radiate in all
directions. ‘These canals extend apparently in a horizontal direction immediately
beneath the dermal layer. The under and outer surface of the wall has, in some
specimens, slight concentric ridges and furrows; it is covered with a dermal layer
which is pierced by minute pore-like apertures 0°3 mm. in width, which are either
irregularly scattered on the surface or disposed in reticulating lines.
This species approaches V’. plicata, but is distinguished from it by the radial
disposition of the canals round the oscules, and the irregular arrangement of the pores
of the under surface. ‘The walls are also thinner. In the general form and dispo-
38 SILICEOUS SPONGES.
sition of the oscules this species also resembles V. miliaris, Reuss, but is readily
distinguished therefrom by the different characters of the under surface.
Distribution. Upper Chalk: Flamborough.
VERRUCULINA CONVOLUTA, Quenst. sp. (Plate IV. figs. 1, 1 a, 4, ¢, d.)
1878. Spongia convoluta, Quenst. Petref. Bd. 5, p. 368, t. 132. f. 49, 50.
18385. Spongia convoluta?, Phill. Geol. York. t. 1. f. 6.
1870. Chenendopora tenuis, p. p., F. Roemer, Geol. Oberschl. p. 301, t. 31. f. 1.
1878. Amphithelion convoluta, Zitt. Stud. Il Ab. p. 60.
Sponge consisting of a convoluted plate, which by the union of the lateral margins
frequently becomes open funnel-shaped. The wall is nearly 7 mm. in thickness, the
margins rounded, and slightly thinner than the wall. Large specimens measure
115 mm. in width at the summit and 100 mm. in height.
Both the outer and inner surfaces of the sponge-wall are thickly covered with
minute apertures, varying from 0°3 to 0°5 mm. in width. Those of the upper or
inner surface have very slightly elevated margins, and are somewhat further apart
than those of the outer surface. These latter appear generally not to possess raised
margins, though in some cases they are present. The canal-structures of the interior,
as also the spicular fibre, have been completely obliterated in all the specimens.
In general form, thickness of the wall,.and the characters of the outer surface
these sponges resemble the V. convoluta, Quenst.; but in Quenstedt’s examples the
features of the inner surface had been obliterated, and I can only suppose that they
were originally similar to these forms. The figure given by Phillips of Spongia
convoluta, loc. cit. t. 1. f. 6, is altogether insufficient for recognition, and might be
applicable to two or three species of these Flamborough forms. F. Roemer has
figured a sponge under the name of Chenendopora tenuis, loc. cit. t. 51. f. 1, which
resembles the present form very closely; and if there had been any certainty that
this figure really represented the original type of his brother’s species, that name
would have had the priority. But judging from the figures and descriptions given
by F. A. Remer himself in the ‘ Nordd. Kreide’ and in the ‘ Palsontographica,’ there
is but little resemblance to the specimen figured by F. Roemer in the ‘Geol.
Oberschl.’ t. 31. f. 1. The identity of Chenendopora tenuis becomes still more
doubtful from the fact that the second figure given of it by F. Roemer, Geol.
Oberschl. t. 31. f. 3, appears to belong to a different species from the fig. 1 of the
same plate.
The similarity of the apertures on both surfaces of the wall and their small dimen-
sions very readily distinguish this from the other species of this genus.
Distribution. Upper Chalk: Flamborough.
VERRUCULINA. 39
VerrvucuLINA PustuLosa, Hinde, n. sp. (Plate III. figs. 2, 2 a.)
1845. Manon miliare, Reuss, p. p., Bohm. Kreide. p. 78, t. 19. f. 13.
1864. Chenendopora tenuis, F. A. Roemer, p. p., Paleont. Bd. 13, t. 15. f. 4, non Manon
tenue, ¥. A. Roem. Nordd. Kreide, t. 1. f. 7.
Sponges wide funnel-shaped, formed by a convoluted plate, which generally over-
laps at the margins. The walls are from 4 to 5 mm. in thickness; the margins are
rounded and of the same thickness as the walls. An average specimen is 64 mm. in
width at the summit, and 42 mm. in height.
The upper or inner surface is covered with small projecting oscules 0:5 mm. in
width, disposed in horizontal lines about 1 mm. apart. The oscules have thickened,
slightly projecting margins, and are about their own diameter from each other. The
under surface has similar but slightly smaller oscules, disposed either irregularly or
in sinuous lines.
There are several examples in the collection which have the same distinguishing
features, and thus appear to belong to a distinct species, characterized by thin
walls, the regular horizontal disposition of the oscules of the inner surface of the
walls, and the closely arranged oscules of the outer surface. The examples of
. this species are also, as a rule, smaller than those already described. Reuss has
figured (/oc. cit. t. 19. f. 13) a fragment of a specimen of this species, and included
it in his Manon miliare; but there is clearly a specific difference between this and
the other forms (figs. 10, 11) which he has placed under this name. The Chenen-
dopora tenuis, figured by F. A. Reemer in the ‘ Paleeontographica’ (/. ¢. f. 4), has the
same disposition of the oscules of the inner surface as the present form; but it is
different from the figure of the original type of his species in the ‘ Nordd. Kreide’
(7. c. f. 7), to which alone the name should be applied.
Distribution. Upper Chalk: Flamborough.
VERRUCULINA MILIARIS, Reuss, sp. (Plate III. figs. 3, 3 a.)
1846. Manon miliare, Reuss, p. p., Bohm. Kreide. p. 78, t. 19. f. 10-12.
1878. Amphithelion miliare, Zittel, Stud. II Ab. p. 60.
Sponges fan-shaped, or, by the coalescence of the incurved margins, becoming
funnel-shaped. The margins rounded, and of the same thickness as the walls. In
some examples the margins are extended into finger-like projections. ‘The walls are
from 5 to 8 mm. in thickness. The largest specimen in the collection is 100 mm.
wide at the summit, and 98 mm. in height.
The upper or inner surface with sparsely scattered oscules 1°2 mm. wide, which
strongly project, obliquely upwards, from the surface. ‘The apertures of the lower
surface of the wall are 0°5 mm. in width, with slightly projecting margins; they are
about their own diameter apart, and not infrequently disposed in sinuous lines. The
spicular structure and the interior canals are destroyed in all the specimens.
40) SILICEOUS SPONGES.
The Flamborough examples have the oscules of the upper surface more projecting
and less numerous than in Reuss’s figure 11a, which I take as the typical form of
the species; as, however, the similarity in other respects is very close, 1 did not
regard the difference in this feature as of sufficient importance to place them as a new
species. Reuss states in his description of this species that the openings of the outer
surface are larger and more projecting than those of the inner surface; but a glance
at his figures plainly shows that the surface which he terms “ outer” is in reality, in
the case of the funnel-shaped specimens, the inner surface. From the V. aurita,
F. A. Remer *, this form is distinguished by the projecting apertures of the under
surface.
Distribution. Upper Chalk: Flamborough.
Verrucunina Reussu, ‘Coy, sp. (Plate V. figs. 1, 1a.)
1848. Manon Reussii, M‘Coy, Ann. & Mag. Nat. Hist. vol. ii. p. 398.
1878. Manon circumporosum, Quenst. Petref. Bd. 5, p. 372, t. 132. f. 55.
Sponge forming large horizontal plate-like expansions, or in the form of a shallow
dish or cup, growing from a short peduncle. The margins are rounded, either even,
lobed, or occasionally digitate. The thickness of the wall is from 10 to12 mm. A
large example measures 190 mm. in width.
Both surfaces of the sponge-wall possess a thickened dermal layer. The oscules
are carried on the summits of small papille; they are 1°75 mm. in width each, and
are disposed irregularly on both surfaces, from one to three diameters apart. The
oscules of the under surface are somewhat less projecting than those of the upper,
but they are nearly equal as regards size.
The interior of the wall is composed of a labyrinthine web of very delicate
anastomosing fibres; there are no indications of distinct canals. The spicules
composing the fibre cannot be distinguished.
The thickness of the walls and the large size of the oscules on both surfaces
readily characterize this species. No figure accompanied M‘Coy’s description; but
there is no difficulty in recognizing the species from his description. The example
figured by Quenstedt under the name of Manon circumporosum appears to belong to
this species.
The examples are numerous and well preserved.
Distribution. Upper Chalk: Flamborough.
VERRUCULINA MACROMMATA, Rem. sp.
1864. Verrucospongia macrommata, Roemer, Palwon. Bd. 13, p. 45, t. 16. f. 4.
1878. Amphithelion macrommata, Zitt. Stud. IL Ab. p. 60, t. 3. f. 15.
Microscopic slide with detached spicules of this species.
Distribution. Upper Chalk: Ahlten, Hanover.
* Paleontographica, Bd. 13, p. 43, t. 16. f. 2.
VERRUCULINA.—STICHOPHYMA. 41
VERRUCULINA PAPILLATA, Hinde, n. sp. (Plate V. figs. 2, 2 a.)
Sponges either cup- or funnel-shaped, or growing in thick undulating expansions ;
the margins rounded and frequently digitate. The walls are massive, and vary in
thickness from 19 to 29 mm. _
The upper surface has numerous, irregularly disposed papilla, some of which
project 7 mm. above the surface. The oscules at their summits are 2°25 mm. in
width. These oscules are the apertures of cylindrical canals which are continued in
an oblique direction to the base of the sponge. The under or lower surface of the
wall has irregularly disposed apertures 1‘5 mm. in width, and from one to three
diameters apart, which project slightly above the surface. Both surfaces are provided
with a compact dermal layer.
The substance of the wall is composed of delicate anastomosing fibres; the spicules
of these fibres are obliterated.
A specimen in the collection measures 140 mm. in width and 110 mm. in height.
The distinguishing feature of this species is the presence of the oblique canals
which extend throughout the sponge. These canals resemble those of the next
genus Stichophyma; thus this form exhibits characters intermediate between Verru-
culina and Stichophyma. The only other species of Verruculina with which it can
be compared is the V. macrommata, Roemer; but in this form, so far as can be judged
from the figures, the oscules are not carried on the summit of papillee, and the inter-
mediate surface is perforated with small apertures. Nothing is stated respecting the
interior canal-structure of this species ; so that I cannot say if any resemblance exists
in this respect.
This species appears to be rare.
Distribution. Upper Chalk: Flamborough.
Genus STICHOPHYMA, Pome/, 1866.
STICHOPHYMA TURBINATUM, Ramer, sp.
1840. Manon turbinatum, ¥. A. Remer, Nordd. Kreide. p. 3, t. 1. f. 5.
1878. Stichophyma turbinata, Zitt. Stud. IL Ab. p. 61, t. 4. f. 5.
Distribution. Upper Chalk: Goslar.
STICHOPHYMA TUMIDUM, Hinde, n. sp. (Plate V. figs. 3, 4.)
1848. Rhizospongia polymorpha, p. p., Charlesworth, Proc. York. Phil. Soc. p. 73.
Sponges simple, elongate, club-shaped, or subcylindrical, usually widest near the
summit, and gradually diminishing towards the basal end. The stem exhibits
alternate horizontal swellings and contractions; near the upper portion in some
specimens it becomes nodose ; the summit is either conical, rounded, or subtruncate.
G
42 SILICEOUS SPONGES.
There is a great variety in the size, from small specimens measuring 150 mm. in
height by 37 mm. in greatest width to large individuals 390 mm. in length by
120 mm. in width.
The openings of the vertical canals usually project slightly above the summit of
the sponge; they vary between 3 and 7 mm. in diameter in different specimens.
There is an example of this species, now in the Jermyn-Street Museum, which has
the vertical canals partially separated from each other so as to form a group of
chimney-like elevations near the summit of the sponge. ‘The lateral surfaces are
covered with numerous circular apertures, with slightly projecting thickened margins.
In weathered specimens the margins are usually smooth. ‘These apertures are some-
times in close contact ; sometimes they are disposed in sinuous lines, which apparently
indicate the presence of canals immediately beneath the dermal layer.
The internal skeleton appears to be composed of anastomosing fibres; the spicular
structure has been destroyed.
This species may be readily distinguished by its large size, mode of growth, and
the dimensions of the vertical and lateral apertures. It appears to be a very common
form at Flamborough ; and there is a single specimen—now composed of iron
peroxide—from the Chalk of the south of England.
Distribution. Upper Chalk: Flamborough, Yorkshire; Bromley, Kent.
Genus JEREICA, Zittel, 1878.
JEREICA PuNcTATA, Miinst. sp.
1833. Siphonia punctata, Minst., Goldf. Petref. Ger. Th. 1, p. 221, t. 65. f. 13.
1878. Spumispongia punctata, p. p., Quenst. Petref. Bd. 5, p. 402, t. 134. f. 12.
1878. Jereica punctata, Zittel, Stud. IL Ab. p. 63, t. 5. f. 1.
Distribution. Upper Chalk: Sudmerberg, Goslar.
JEREICA POLYSTOMA, Lamer, sp.
1864. Jerea polystoma, F. A. Reemer, Palon. Bd. 18, p. 34, t. 12. f. 5.
1878. Jereica polystoma, Zitt. Stud. II Ab. p. 63, t. 4. f. 11, 12.
Microscopic spicules of this species from Zittel’s collection.
Distribution. Upper Chalk: Ahlten, Hanover.
JEREICA CLAVA, Lee, sp.
1839. Siphonia clava, Lee, Magazine of Nat. Hist. vol. iil. p. 12, f. 2, 3, 4.
1840. Siphonia ocellata, F. A. Roemer, Nordd. Kr. p. 5, t. 2. f. 2.
Sponge simple, straight, elongate, club-shaped, with a conical summit. Greatest
width a short distance below the summit; from thence it gradually tapers to the
JEREICA.—CCiLOCORYPHA. 45
narrow cylindrical base; occasionally slight constrictions are present. The largest
example in the collection is 140 mm. in length and 48 mm. in diameter.
At the summit is a group of closely arranged circular or polygonal openings of
vertical canals, each 2°8 mm. wide. The lateral surface of the sponge appears to
have been smooth, and thickly covered with minute pore-like apertures 0°3 mm. in
width. Beneath the dermal layer very delicate anastomosing fibres can be seen,
which are apparently composed of minute thorny spicules; but the forms of these
spicules are not sufficiently clear to be figured.
The S. ocellata, F. A. Reem., appears to be identical with this species. Roemer
states that only six canal-apertures are present on the summit of his sponge; but the
number of these canals evidently depends upon the size of the individual specimen,
and furnishes no ground for a specific character.
Distribution. Upper Chalk: Flamborough.
JEREICA cYLINDRICA, Hinde, n. sp. (Plate VI. figs. 1, 1a.)
Sponge massive, compressed, cylindrical, with an expanded flattened base; the
summit truncate, with a shallow basin-like depression. The only specimen is 72 mm.
in height and 84 mm. in width.
The surface characters are not well preserved, as the dermal layer has disappeared,
and only filled up sinuous canals running in a vertical direction can be distinguished.
In a vertical section through the central portion of the sponge, however, there are
exposed two sets of canals—one generally vertical, and the other extending from the
lateral surface to the central portion of the sponge. Both series of canals are minute,
and do not appear to exceed 0°35 mm. in width.
The spicules of this sponge are very irregular in form, with numerous thorny
and tubercular branches. They appear to be loosely connected together by their
twig-like extensions; and distinct spicular fibres are not apparent in the vertical
section.
In its general form and size, as well as in the delicate character and disposition of
the canals and spicules, this species may be readily distinguished from the others of
this genus.
Distribution. No label is attached to this sponge; but it appears to have been
derived from the Upper Green Sand of Wiltshire.
Genus CAUSLOCORYPHA, Zittel, 1878.
CLOCORYPHA, sp.
Distribution. Cretaceous: St. Adresse.
44 SILICEOUS SPONGES.
Genus SCYTALIA, Zité. 1878.
ScyTaLIA RADICIFORMIS, Phillips, sp. (Plate VI. figs. 4, 4a, 6.)
1835. Spongia radiciformis, Phillips, Geol. of Yorks. p. 90, t. 1. f. 9.
1864. Eudea annulata, F. A. Roemer, Paleont. Bd. 13, p. 26, t. 11. f. 2.
1878. Scytalia radiciformis, Zitt. Stud. Il Ab. p. 64, t. 5. f. 4.
Sponge either simple or occasionally two individuals growing in close contact,
partially coalesce together, elongate, cylindrical, with occasional swellings and
constrictions. The summit is conical; and the body tapers gradually below into
a cylindrical stem: the natural termination of this is not preserved. A small
specimen measures 65 mm. in length by 22 mm. in width; and a large example is
215 mm. long by 58 mm. wide.
The characters of the dermal layer cannot be distinguished in the Flamborough
examples. The cloacal tube is about 8 mm. in width, and extends about two thirds
the length of the sponge; from the summit-aperture sinuous canals 0°8 mm. wide
radiate for a short distance down the cone. In a horizontal section numerous canals
about 0°6 mm. in width can be seen extending from the cloacal tube towards the
exterior surface of the sponge. The surface beneath the dermal layer exhibits very
delicate reticulate fibres which, in specimens from Germany, are seen to be composed
of minute, irregularly branched, spinous spicules.
This species appears to be common.
Distribution. Upper Chalk: Flamborough; Ahlten, Hanover.
Scytania Fastierata, Lee, sp. (Plate VI. fig. 3.)
1839. Spongia fastigiata, Lee, Magazine Nat. Hist. vol. in. p. 14, f. 8.
Sponge simple, turbinate, with a very prominent conical summit, usually marked
off from the main portion of the body by a prominent collar-like ridge. The sponge
is widest at the junction of the cone with the body, and diminishes somewhat rapidly
to a slight cylindrical stem. The lateral surface exhibits shallow ridges and furrows.
An average specimen is 130 mm. in height by 70 mm. in width.
A smooth dermal layer appears to have covered the entire surface of the sponge.
The cloacal tube is 10 mm. wide, apparently cylindrical, and extends nearly to the
lower portion of the body. Numerous slightly arched canals 1 mm. in width extend
from the cloaca towards the exterior. Similar canals are also exposed on the conical
summit, radiating from the cloacal aperture, in specimens where the dermal layer
has been removed. These superficial canals by the further growth of the sponge
become inclosed, and then appear as interior canals.
The reticulate fibre of the interior can be distinguished in specimens treated with
acid; but nothing beyond the general Rhizomorine character of the spicules can be
recognized.
SCYTALIA.—STACHYSPONGIA. 45
There are several examples of this species in the collection, and they are very
constant in their outer form and other characters.
Distribution. Upper Chalk: Flamborough.
ScYTALIA TEREBRATA, Phillips.
1835. Spongia terebrata, Phill. Geol. Yorks. p. 90, t. 1. f. 10.
1878. Scytalia terebrata, Zitt. Stud. II Ab. p. 65.
Sponge massive, subcylindrical, with a flattened conical summit. The lower
portion of the body contracts to a cylindrical stem. The lateral surface carries
numerous concentric, slight, subangular ridges and shallow open furrows. The only
specimen in the collection is 220 mm. in length by 87 mm, in greatest width.
The surface, with the exception of the summit, is covered with a smooth dermal
layer. The straight cylindrical cloaca is 13 mm. in width ; from its aperture numerous
sinuous canals extend to the margin of the cone. In a vertical section thickly set
canals, 1:25 mm. in width, are exposed, extending from the cloaca to the exterior.
This species may be distinguished from the preceding by its different form and
larger dimensions. It appears to be the species indicated by Phillips's figure, which,
however, must have been drawn to the scale of one third the original.
Distribution. Upper Chalk: Flamborough.
Genus STACHYSPONGIA, Zittel, 1878.
STACHYSPONGIA sPIcA, Remer, sp. (Plate VI. figs. 2, 2 a.)
1864. Siphonocelia spica, F. A. Roemer, Palzont. Bd. 13, p. 30, t. 11. f. 5.
1878. Stachyspongia spica, Zitt. Stud. II Ab. p. 65, t. 5. f. 5.
Sponge massive, cylindrical, the surface covered with conical projections, disposed
either irregularly or rudely spiral, with their summits directed upwards. The
summit appears to have been truncate. An imperfect specimen is 210 mm. in
length and 73 mm. in diameter. The opening of the tubular cloaca is 31 mm.
wide at the summit, whilst at the basal end of the specimen it only measures
15mm. Thickly set minute sinuous canals extend from its inner surface towards
the exterior of the sponge, The outer surface appears to have been covered by a
dermal layer, in which are irregularly disposed minute apertures from °3 to ‘6 mm.
in width. Beneath this layer the surface is closely seamed by sinuous canals about
0:75 mm. in width. I have not detected any special apertures at the summit of
the conical elevations. The spicules of the interior are so poorly preserved that only
their Rhizomorine characters can be distinguished.
The example figured appears to belong to Reemer’s species, so far as can be judged
from his short description and diminutive figure. He states, however, that the
cloacal tube is only one fourth of the diameter of the sponge, but in the figure it
46 SILICEOUS SPONGES.
is represented as one third as large. In the Museum specimen the opening at the
summit is nearly one half the diameter, whilst at the basal end it measures only one
fourth the thickness of the sponge; so that at the base, at least, it agrees with
Reemer’s description.
This sponge appears to be a rare form in the Grey Chalk; there is a fragment of
a specimen in the collection inclosed in an Upper Chalk Flint, which also appears to
belong to this species.
Distribution. Gray Chalk: near Dover. Upper Chalk: South Coast of England,
probably.
Genus PACHINION, Zittel, 1878.
PacuInion soriptuM, Remer, sp. (Plate VII. figs. 1, 1 a, 4, c,d.)
1864. Jerea scripta, F. A. Roemer, Paleont. Bd. 13, p. 34, t. 138. f. 1.
1878. Pachinion scriptum, Zitt. Stud. II Ab. p. 66, t. 5. f. 2.
Sponges simple, cylindrical, or inversely conical, the lower portion of the body
gradually tapering to a cylindrical stem; the summit rounded or -depressed conical.
Very variable in size—a small specimen measuring 100 mm. in length by 55 mm. in
width, whilst a large cylindrical example attains a length of 380 mm. and a width
of 68 mm.
The cloacal tube is approximately cylindrical, and extends nearly the entire length
of the sponge; at the summit of a fairly large specimen it is 18 mm. in width. The
outer surface is covered by a smooth dermal layer. No distinctive canals are present,
the circulation apparently being carried on between the apertures of the spicular
fibre. ©
The interior of the sponge is formed by a relatively coarse fibrous network; the
fibres are about 0°315 mm. in width. On the surface of weathered specimens and in
sections the edges of the fibre have a rude resemblance to hieroglyphic characters.
The fibre is composed of irregular branching spicules with tubercled surfaces.
The dermal layer is formed of much smaller and more branched spicules than those
of the fibre. In the specimens from Flamborough only the general characters of the
spicules of the fibre can be recognized; but in examples from Germany the spicules
are beautifully preserved.
This species is very abundant in the Chalk at Flamborough; the fibre is, for the
most part, in a silicified condition, though occasionally the silica is replaced by
calcite. A few specimens are also met with in the Chalk of the south of England ;
but the siliceous spicular structure of these has been replaced by iron oxide, and is
quite as indistinct as in the Flamborough examples. This species can be readily
distinguished from the cylindrical specimens of Scytalia and Phymatella in the
Flamborough beds by its smooth surface, the coarse characters of the fibre, and the
absence of canals.
PLACONELLA.—DORYDERMA. AT
Distribution. Upper Chalk: Flamborough; south of England; Schweichelt,
Brunswick (Zittel’s coll.).
Family WEGAMORLNA, Zittel.
Genus PLACONELLA, Hinde, n. g.
Sponges sessile, growing in flattened, cake-shaped masses. The upper surface
uneven with shallow depressions, containing the apertures of several prominent
canals. The surface between the depressions faintly furrowed, and with openings
of numerous canals of different sizes. ‘The interior permeated irregularly by canals.
The skeleton consists of smooth, robust, irregularly branching spicules, which
closely intertwine together and form a reticulate mesh.
The spicules and the general structure of the skeleton of this genus are similar to
to those of Megalithista, from which it is distinguished by its mode of growth and
the absence of a central cloaca.
PLACONELLA PERFORATA, Hinde, n. sp. (Plate VII. figs. 2, 2a, 6.)
The only example of the genus has a circular outline about 68 mm. in diameter,
and is 35 mm. in thickness. ‘The upper surface is gently convex, the central portion
is elevated ; between this and the margins are three well-marked depressions, and one
or two minor ones; in these depressions are several irregularly disposed canal-apertures
about 2 mm. each in width. ‘The general surface is also covered with slight furrows
and irregularly disposed canal-apertures, in addition to the openings between the
spicular mesh.
The spicular skeleton is best shown on the surface of the specimen. A’ detached
spicule which I have been able to measure is 1 mm. in length by -15 mm. in width.
No distinctive dermal layer has been preserved ; there are, however, in the depressions,
fragments of long, straight, cylindrical spicules which may perhaps belong to the
sponge.
The specimen in the collection was labelled Achillewm tuberosum, Goldf.; but this
species has a skeleton of Rhizomorine spicules, and belongs to the genus Cnemidiastrum.
Distribution. Upper Jura: Nattheim, Wiirtemberg.
Genus DORYDERMA, Zittel, 1878.
DoryDrERMA DicHotomtM, Benett, sp. (Plate VIII. figs. 1, 1a, 10.)
1831. Polypothecia dichotoma, Benett, Catalogue of the Organic Remains of the County of
Wilts, pl. 13.
1864. Non Polyjerea dichotoma, F. A. Roemer, Palzont. Bd. 13, p. 36, t. 14. f. Le.
Sponge dendriform, growing from a simple upright cylindrical stem, which
48 SILICEOUS SPONGES.
divides and subdivides dichotomously; the branches are cylindrical or slightly
compressed, approximately of the same thickness throughout, and from 14 mm.
to 19 mm. in diameter in different specimens. The summits of the branches are
rounded. An average specimen is 250 mm. in height, with an expansion of 190 mm.
across the summit.
The branches are traversed longitudinally by a series of simple cylindrical tubes,
2mm.each in width. No regular canal-openings appear to be present on the lateral
surfaces of the branches; and the circulation seems to have been carried on between
the irregular interspaces of the mesh, which average ‘5 mm. in width.
The arms of the spicules are ‘09 mm. in thickness; they are very compactly inter-
woven together so as to form a very close mesh. No dermal spicules have been
detected in this species.
This species may be distinguished from D. ramoswm and D. Remeri by the great
regularity in its mode of growth and the close character of the spicular mesh. It
appears to be abundant.
Distribution. Upper Green Sand: Warminster, Wiltshire.
DoryDERMA RAMOoSUM, Mantell, sp. (Plate VIII. figs. 2, 2a.)
1822. Spongia ramosa, Mantell, Fossils of the South Downs, p. 162, t. 15. f. 11.
1808. A ramose Alcyonite, Parkinson, Organic Remains, vol. i. t. 7. f 6.
1878. Doryderma ramosa, Zitt. Stud. II Ab. p. 68.
Sponge branching, growing from an upright cylindrical stem, which divides and
subdivides occasionally, giving off short stumpy branches, from 12 to 18 mm. in
diameter. ‘The extremities of the branches are rounded. ‘The vertical canals are
about 15 mm. in width; they can hardly be distinguished, owing to the imperfect
preservation of the interior structure of the sponge. The interspaces of the mesh on
the surface are °6 mm. wide. The spicular arms are ‘1575 mm. in thickness. No
surface-spicules have been preserved.
The type of Mantell’s species is in the Museum collection. Its structure has been
so completely altered by fossilization, that it was only after repeated scrutiny that I
discovered a single spicule and was thus enabled to determine its true characters.
The branching sponges which, from their similarity of form, I refer to this species
are not uncommon in the interior of flints in the Chalk of Wiltshire. The surface-
structure is usually well preserved ; but the interior characters are mostly destroyed.
This species is less regular in its mode of growth, and the spicules and mesh-inter-
spaces are larger than in D. dichotonwwm, Benett, whilst the stems are usually smaller,
and the mesh-interspaces also, than those of D. Remeri, Hinde.
Distribution. Upper Chalk: near Brighton; Oare, Wiltshire.
DORYDERMA. 49
DoryperMA Ramer, Hinde. (Plate VIII. figs. 3, 3a, 3 3.)
1808. A ramose Alcyonite, Parkinson, Org. Rem. vol. ii. p. 92, t. 7. f. 7, 12.
1864. Polyjerea dichotoma, F. A. Reem. Paleont. Bd. 13, p. 36, t. 14. f. 1.
1878. Polyjerea dichotoma, Quenst. Petref. Bd. 5, p. 423, t. 135. f. 10, 11.
1878. Doryderma dichotoma, Zitt. Studien, II Ab. p. 67, t. 7. f. 1.
Sponge branching, consisting of a thick stem from which short branches are given
off in an irregular manner. The stem and branches traversed by numerous vertical
canals about 2 mm. each in diameter. The interspaces of the mesh at the surface
are from 1 mm. to 1-25 mm. in width. ‘The spicules are -1575 mm. in thickness,
and occasionally 2 mm. in length. A dermal layer is formed by numerous trifid
spicules, whose shafts fill the interspaces of the mesh, whilst their trifid summits
slightly project outwards.
F. A. Remer’s specific name, dichotoma, having been previously employed by
Miss Benett for a sponge of this genus, it is necessary to give another designation
to his species ; and I have therefore termed it Rameri, in memory of its discoverer.
From an examination of undoubted examples of this species, from the Upper Chalk
of Brunswick, presented to the Museum by Prof. Zittel, I am enabled to determine
its distinctness from either of the two foregoing species. It is much less regular in
its habit of growth, the stem is much thicker, and the mesh interspaces are notably
larger than in D. ramosum and D. dichotomum, Benett. The specimens appear to be
rare in the Chalk of this country, and are only met with in the interior of flints. It
is very seldom that the dermal trifid spicules are preserved ; but I have detected them
in one or two fragments in the Collection.
Distribution. Upper Chalk: Wiltshire; Brunswick (Zit#. coll.).
DoryperMA Benerti, Hinde, n.sp. (Plate IX. figs. 1, 1a, 18.)
1808. An Alcyonite, Parkinson, Org. Rem. vol. ii. t. 10. f. 6.
18381. Polypothecia, Benett, Cat. Org. Rem. Wilts. t. 10. f. 1.
1847. Non Jerea elongata, Mich. Icon. Zoophy. p. 134, t. 29. f. 4.
1854. Jerea Carteri, p. p., Morris, Cat. Brit. Foss. p. 28.
Sponges simple, massive, elongate pear-shaped, or with a cylindrical body sup-
ported on a relatively thin stem, occasionally also subglobate. The summit is
either rounded, horizontally truncate, or obtusely conical. In some examples the
body gradually tapers downwards to the stem, in others there is a distinct constriction
between the basal portion of the body and the top of the stem. There is great
variability in the size of different examples—the smaller forms measure 72 mm. by
42 mm., whilst the larger specimens reach a length of 390 mm. and a width of
125 mm.
The body of the sponge is penetrated by numerous straight vertical canals, 2°5 mm.
each in width, which open at the summit in close proximity to each other. A few
H
50 SILICEOUS SPONGES.
of these canals also extend into the stem. Numerous, closely set, arched canals,
‘5 mm. wide, extend from the lateral surface towards the central portion of the
sponge. The surface-openings of these lateral canals are formed by the irregularly
shaped interspaces of the mesh. There are also numerous horizontal canals in the
stem of the sponge.
The spicules of the body are robust, smooth, and irregularly branching, and about
‘135 mm. in thickness. Some of the spicular arms gradually taper towards their
extremities ; in others, however, the extremities are expanded in a similar manner to
those of the genus Lyidium. ‘The spicules of the stem are slender, elongate, thread-
like, with branching ends; they are loosely intertwined together in the direction of
their length. No surface-spicules have been noticed.
This species is apparently abundant and generally well preserved. It is very
probable that the figure given by Parkinson (/oc. cit.) is of a sponge belonging to this
species, though he makes mention of a large central cavity in the interior, of which
there is no evidence in the figure. Michelin refers the form figured by Miss Benett
to his Jerea elongata; but the spicular structure of the two is altogether different.
Some examples resemble in appearance the figure of Jerea pyriformis, Lamx. Exp.
Méthod. t. 78. f. 3; and they have apparently been referred to this species in Morris’s
Catalogue. There is, however, no resemblance in their spicular structure, as,
according to Prof. Zittel, Jerea pyriformis belongs to the Tetracladina family.
Prof. Morris also designates the sponges figured by Miss Benett in plate x. Cat.
Org. Rem. by the name of Jerea Carteri, but no reference is given to the author of
the specific name; and on this account, and on the further fact that in this plate two
different species are figured, whilst it is not stated to which the J. Carteri applies, I
have not retained that term for the present species.
Notwithstanding the variety of form and size in the examples which I refer to this
species, there exists such a close resemblance in their canal and spicular structures
that I do not see any ground for placing them under more than one species.
‘Distribution. Upper Green Sand: Warminster; Canamore quarry, near Crockerton.
Genus HOLODICTYON, Hinde, n. g.
Sponges inverted, conical, or irregular in form, with rounded or flattened summits,
and stems with branching processes. In the centre of the summit there is a shallow
cup-shaped or cylindrical cavity, which appears to extend only to a short distance into
the sponge.
The skeleton consists of a meshwork of smooth, irregularly branching spicules ;
the spicular arms either attenuated or with a spoon-shaped or elongated expansion
at their termination. No canals appear to be present.
I propose this genus to include a small group of sponges which closely resemble
HOLODICTYON.—PACHYPOTERION. ol
those of Doryderma in the character and disposition of the spicular mesh, but, unlike
that genus, appear to be destitute both of vertical and horizontal canals, so that the
circulation is wholly carried on between the interspaces of the mesh.
Hotopictyon capitatum, Hinde, n.sp. (Plate VII. figs. 3, 3a, 3d.)
The examples of this species vary from 100 to 150 mm. in length, and from 30 to
85 mm. in width at the summit. The cloacal aperture, if it may be so termed, is
from 6 to 9 mm. in width at the summit; in one example, which has been cut in two,
its depth is20mm. The spicules are robust, and measure about ‘2 mm. in thickness.
No dermal layer has been detected. ‘The spicules of the stem appear to be similar
to those of the body of the sponge. ‘The interstitial apertures are irregular in form,
and from ‘5 to ‘75 mm. wide.
Distribution. Upper Green Sand: Warminster, Wiltshire.
Genus PACHYPOTERION, Hinde, n. g.
Sponges simple, massive, with cup- or goblet-shaped bodies, frequently of great
thickness, supported on elongated cylindrical or compressed stems. ‘The body of the
sponge is traversed by a double series of canals—one, rising from the basal portion, is
either vertical or follows the contour of the cup and opens into it; the other,
and smaller, series commences on the exterior of the cup, and extends downwards in
an arched direction towards the centre of the sponge. The skeletal spicules of the
body of the sponge are smooth, curved, branching bodies of very irregular form; the
extremities of the arms are either blunted or slightly expanded. They form a compact
network both by the intertwining of the arms and by the attachment of the expanded
ends to the surfaces of adjoining spicules. The spicules of the stem are elongated,
thread-like, bifurcated at their ends, and loosely connected together. No dermal
spicules have been preserved.
From Doryderma this genus is distinguished mainly by its cup-shaped mode of
growth. It differs from Heterostinia, Zittel, in the apparent absence of those minute
spicular bodies in which, according to Zittel, the larger spicules of this latter genus
are imbedded, and which form the principal mass of the skeleton. It is possible
that these smaller spicular elements may have been originally present in the examples
of Pachypoterion; but, considering the state of preservation of these sponges, I think
that traces of them would have remained.
The examples of this genus at present known are limited to the Upper Green
Sand. On account of their resemblance in external form and in the canal-apertures
of the inner and outer surfaces to Chenendopora, these sponges have hitherto been
placed under that genus, and only by an examination of the spicular structure can
the differences be detected.
H2
SILICEOUS SPONGES.
Or
Lo
PAcHYPoTERION RoBusTUM, Hinde, n.sp. (Plate IX. figs. 2, 2a, 2 6.)
Sponges cup- or goblet-shaped, the walls very thick, so that in some examples the
cup is nearly filled with the siliceous skeleton. The body either gradually tapers to
the simple elongated, cylindrical stem, or there is a sharp constriction between the
base of the cup and the top of the stem. The specimens vary considerably in size:
small examples measure 110 mm. in length by 65 mm. in width, whilst larger forms
are 270 mm. long by 150 mm. in width.
The canal-apertures in the interior of the cup are, in some examples, arranged in
concentric circles; they are about 2 mm. in width, and either elliptical or circular in
section. ‘The apertures of the exterior surface vary between ‘5 and ‘9 mm. in width.
They appear to be disposed in a regular vertical series. In a horizontal section of
the base of the cup, the lateral canals appear as so many radiating lines extending
from the outer surface to near the central portion. ‘The arms of the spicules measure
*155 mm. in thickness.
This species appears to be abundant. The examples vary considerably in size,
thickness of the wall of the cup, and in the character of the stem; but as there are
numerous gradations between the extreme forms, I have included them under a
single species, which may be distinguished from the next by the thickness of the
walls and of the spicules and the dimensions of the canals.
Distribution. Upper Green Sand: Warminster, Pewsey, Sambourne, Folkestone.
PacHyPoTeRiIon compactuM, Hinde, n.sp. (Plate IX. figs. 3, 3 a.)
1816. Alcyonite (funnel-form), W. Smith, Strata identified by Organic Fossils, t. 5. f. 12.
Sponges cup-, vase-, or open funnel-shaped, with simple cylindrical stems. The
margins rounded. The walls of the body are from 5 to 9 mm. in thickness. The
width of the cup varies in different examples from 65 to 110 mm.
The canals opening into the interior of the cup are 1 mm. in width, inconspicuous,
and irregularly distributed. Those extending from the outer surface to the interior
are very numerous, about °3 mm. in width, and nearly horizontal in direction.
The spicules are of the same character as those of P. robustum, but of less thick-
ness; they measure ‘09 mm. in width, and about ‘225 mm. in length. ‘They are
very closely interwoven together, so as to form a compact network.
This species is distinguished from P. robustum by its thinner walls, smaller
spicules, and more compact spicular tissue.
The specimen figured is the original of William Smith’s Alcyonite, loc. cit. t. 5.
f. 12; but the lower portion of the stem is not represented. ‘The interior of this
specimen is filled with the hard matrix. The spicular structure (fig. 3 @) is from the
exterior surface of the same example.
Distribution. Upper Green Sand: Warminster; Saumur.
HETEROSTINIA.
or
wo
Genus HETEROSTINIA, Zittel, 1878.
HETEROSTINIA OBLIQUA, Genett, sp. (Plate X. figs. 2, 2a, 26, 2c.)
1831. Polypothecia obliqua, Benett, Cat. Org. Rem. Wilts. t. 8. f. 1.
Body of sponge consisting of a variously convoluted plate-like wall, either platter-,
ear-, or fan-shaped, and not infrequently, by the coalescence of the wall-margins,
becoming either cup- or open funnel-shaped. The body is usually supported by one
or more rod-like short processes, which are sometimes compressed so as to form
‘semipalmate expansions. The walls are from 7 to 10 mm. in thickness. The
examples vary greatly in size: a large cup-like specimen attains a width of 85 mm.,
and one platter-shaped form is 200 mm. in extension.
On both surfaces of the wall there are numerous irregularly disposed circular
apertures, about ‘7 mm. each in width, and from one to three diameters apart ;
between these circular apertures are the irregular openings formed by the spicular
mesh. No indications of canals in the interior of the wall have been preserved in
the specimens which I have examined, though occasionally the sponge-wall is pene-
trated by cavities which appear to be due to external causes.
The spicules of the body-skeleton are relatively large, vermiculate, branched, and
attached to each other by their slightly expanded extremities. In the best preserved
specimens the surface of the body-spicules exhibits a very fine, minute granulation ;
in other specimens the spicules appear to have smooth surfaces, but the smoothness
may be due to the wearing-off of the fine granulation by fossilization. In all other
respects the sponges with the granulated surfaces to the spicules and those with
smooth surfaces are precisely similar, so that I find it impossible to constitute two
species based on this feature alone. The body-spicules vary in thickness between
112 mm. and 1575 mm. The spicules of the stem are thinner and more elongated
than those of the body, and they are connected by being adpressed and intertwined
together. ‘The sponge appears to have possessed a dermal layer consisting of trifid
spicules with horizontally expanded, bifurcated, and pointed head-rays. This dermal
layer is but very rarely preserved. The minute irregularly shaped spicules which, in
the type of the genus, intervene between the larger mesh-spicules are also very rare,
and I have only detected their presence in portions of the stem.
In the mode of attachment of the body-spicules of this species by their slightly
expanded extremities, these sponges resemble those of the genera Lyidiwm and
Pachypoterion ; and I was at first disposed to place them under the former of these
two genera. ‘There are slight evidences, however, of the presence of the minute
spicular bodies which constitute the distinguishing character of the genus Hetero-
stinia, Zitt.; and Prof. Zittel, who has examined one of these sponges, pronounces’ it
a genuine example of this genus. In this case, the definition of Heterostinia will
54 SILICEOUS SPONGES.
have to be amended, so as to include sponges in which the body-spicules have
expanded extremities, as well as those with attenuated and pointed ends to the body-
spicules.
The figure given by Miss Benett, Joc. cit. t. 8. f. 1, leaves no doubt as to the
identity of the form represented, for there is no other sponge from the Wiltshire
Chalk with the pecuhar elongated compressed stem or root shown in her figure.
There is no certainty, however, that the figure of Chenendopora obliqua*, Michelin,
represents the same species as Polypothecia obliqua, Benett.
This species appears to have been very common in the Upper Chalk. As a rule
the specimens are inclosed in the hollows of flints, and exhibit, when thus preserved,
the spicular structure of the surface in a very beautiful manner. Generally the stem
of the sponge with only a small portion of the body is preserved, and in many
instances the stem alone remains in the interior of the flint. A few examples have
been preserved in the Chalk of Flamborough, and in these the body-wall remains and
the stem is usually wanting. In one example from this locality the sponge is nearly
flat and platter-shaped, and instead of a single central stem it gives off from different
parts of its under surface numerous small rod-like processes.
Distribution. Upper Chalk: Flamborough; Beckhampton, near Brighton; War-
minster, Stockton, Upware.
Genus NEMATINION, Hinde, n. g.
Sponges simple, elongate, rod-like in form, with a relatively small cup-shaped
summit, supported on a long cylindrical simple or bifurcated stem, which is either
branched or obtuse at its lower extremity. Below the cup vertical canals extend
throughout the length of the stem. The exterior surface is thickly covered with the
apertures of horizontal canals. The skeleton is composed of elongated, smooth,
thread-like spicules, which bifurcate near their extremities. These spicules are
interlocked together by their filiform extremities so as to form an open mesh-
work. 2
The peculiar feature of this genus is the small inconspicuous cup carried on the
summit of a very long stem. In its general form, and in the character of the
spicules, the genus approaches Carterella, Zitt.; but this latter genus has a rounded
summit, and the spicules, as a rule, are knobbed and blunted at their ends.
NEMATINION CALYCULUM, Hinde, n.sp. (Plate X. figs. 1, 1a, 1, 1c.)
The rod-like stem of the sponge is either straight or sinuous, usually simple,
though bifurcate examples occur. There is great variation in the length of different
specimens: the largest form which is in the collection is 310 mm. in length by
* Icon. Zooph. p. 182, t. 41. f. 2a, b.
CARTERELLA.—ISORAPHINIA. 55
45 mm. in width at the summit; a smaller form is only 20 mm. in width and
160 mm. in length. In some examples the width of the cup at the summit but
slightly exceeds the width of the stem.
The vertical canals penetrating the stem are ‘7 mm. in width and irregularly
disposed. In some examples the surface of the stem exhibits sinuous vertical canals.
The apertures of the horizontal canals, about ‘5 mm. in width, are present both on
the exterior of the cup and the stem; they are very closely set, oval or circular in
form, and bounded by the spicules of the mesh.
The spicules forming the cup and the eaterior portion of the stem are ‘0675 mm.
in thickness; they are disposed in close contact in the direction of the length of the
sponge. The spicules of the ¢nterior of the stem are much more slender than those
near the outer surface, and they are loosely connected together by their bifurcated
extremities. In length they vary between ‘787 and 1:575 mm.
The specimens are not uncommon, though perfect examples are rare.
Distribution. Upper Green Sand: Warminster, Wiltshire.
Genus CARTERELLA, Zittel, 1878.
CARTERELLA CYLINDRICA, Zitt.
1878. Carterella cylindrica, Zitt. Studien, II Ab. p. 68, t. 2. f. 7 and t. 7. f. 2.
Distribution. Upper Green Sand: Kelheim, near Regensburg (Zittel’s coll.).
CARTERELLA SPICULIGERA, Remer, sp.
1864. Jerea spiculigera, F. A. Roemer, Palont. Bd. 18, p. 34, t. 12. f. 6.
1878. Hulespongia, Quenst. Petref. Bd. 5, p. 414, t. 1385. f. 1, 2.
1878. Carterella spiculigera, Zitt. Studien, II Ab. p. 69, t. 7. f. 2.
Detached spicules of this species from Zittel’s collection.
Distribution. Upper Chalk: Ahlten, Hanover.
Genus ISORAPHINIA, Zttel, 1878.
IsORAPHINIA TEXTA, Ramer, sp. (Plate X. figs. 5, 3a, 3b.)
1864. Siphonocelia texta, Reemer, Paleont. Bd. 13, p. 29, t. 10. f. 11.
1878. Hulespongia texta, Quenst. Petref. Bd. 5, p. 417, t. 135. f. 3-7.
1878. Isoraphinia texta, Zitt. Studien, II Ab. p. 69, t. 5. f. 8 and t. 7. f£. 3.
Body of sponge cylindrical, the lower portion gradually tapering to a simple
rounded stem; the summit is truncate or depressed conical. Fairly large specimens
measure 230 mm. in length by 54 mm. in width.
The cloacal tube is nearly cylindrical, and extends to the lower portion of the body
56 SILICEOUS SPONGES.
of the sponge; ina large specimen it is 19 mm. wide at the summit. Specialized
canals do not appear to exist, and the circulation seems to have been carried on
through the interspaces of the spicular mesh.
The spicules are relatively large, straight, or slightly bent, cylindrical, with nodose
extremities; they attain a length of 1-25 mm. In the interior of the sponge these
spicules are united into small fascicles, and interwoven at their extremities to form
an open meshwork. On the exterior surface, however, they are disposed without
any definite arrangement, and cross each other in every direction, forming a compact
surface-layer.
This species is not uncommon in the Flamborough Chalk ; and may be distinguished
from the cylindrical forms of Pachinion and other genera occurring in the same beds
by the scattered layer of spicules on the exterior surface of the sponge, which have
been preserved, owing to their relatively large size.
Distribution. Upper Chalk: Flamborough. Planer-Kalk: Dohrnten, near Salz-
gitter (Zittel’s coll.).
Family ANOMOCLADINA, Zittel.
Genus CYLINDROPHYMA, Zittel, 1878.
CYLINDROPHYMA MILLEPORATA, (oldf. sp.
1833. Scyphia milleporata, Goldf. Petref. 1 Th. p. 8, t. 3. f. 2.
1878. Scyphia milleporata, Quenst. Petref. Bd. 5, p. 120, t. 121. £. 1-7.
1878. Cylindrophyma milleporata, Zitt. Studien, II Ab. p. 70, t. 5. f. 6.
Distribution. Upper Jura: Oerlingen; Wodna, Randen.
Genus MELONELLA, Zittel, 1878.
MELONELLA RADIATA, Quenst. sp.
1858. Siphonia radiata, Quenst. Der Jura, p. 679, t. 82. f. 18.
1833. Siphonia pyriformis, p. p., Goldf. Petref. 1 Th. t. 35. f. 10.
1878. Siphonia radiata, Quenst. Petref. Bd. 5, p. 249, t. 126. f. 60-72.
1878. Melonella radiata, Zitt. Studien, II Ab. p. 71, t. 5. f. 7.
Distribution. Upper Jura: Heuberg.
Genus LECANELLA, Zitéel, 1878.
LECANELLA PATEREZFORMIS, Zittel.
1878. Lecanella patereformis, Zitt. Studien, II Ab. p. 71, t. 6. f. 1.
Detached spicules of this species.
Distribution. Upper Chalk: Ahlten, Hanover.
MASTOSIA.—HINDIA. 57
Genus MASTOSIA, Zittel, 1878.
MAstosIA NEOcOMIENSIS, Hinde, n.sp. (Plate X. fig. 4.)
The only representatives of this species are detached spicules, which occur in green-
sand strata, mingled with spicules of tetractinellid sponges. The spicules in question
have a spherical or slightly elongated centre, from which three to five short cylindrical
arms project. The arms radiate irregularly from the central node, and are slightly
expanded at their ends. The central node averages ‘2 mm. in thickness, and the
arms are ‘225mm. in length. These spicules are distinctly typical of the Anomo-
cladine family; and though up to the present no sponge belonging to this family has
been recognized from the Cretaceous series, the occurrence of these spicules clearly
proves their existence in this period.
Some of the spicules described and figured by Mr. H. J. Carter from the Upper
Green Sand of Devonshire as “ knots or branching centres of silicified fibre like that
of Dactylocalyx”’* evidently belong to this species.
Distribution. Lower Green Sand: Haslemere, Surrey. Upper Green Sand: Devon-
shire (Mr. H. J. Carter).
Genus HINDIA, Duncan, 1879.
Hinpia Fisrosa, Ferd. Remer, sp. (Plate XIII. figs. 1, 1a, 14.)
1860. Calamopora fibrosa, F. Roemer (non Goldfuss), Die silurische Fauna d. westl. Tennessee,
p. 20, t. 2. f. 2, 2a, b.
1875. Spherolites Nicholsoni, Hinde, Abstract of the Proc. Geol. Soc. no. 305.
1875. Hindia spheroidalis, Duncan, Ann. & Mag. Nat. Hist. 5 ser. vol. iv. p. 84, t. 9.
Sponges globular in form, with an even rounded surface, and without peduncle
or any point of attachment. The specimens vary between 13mm. and 38 mm. in
diameter.
As seen in a section through the centre, the growth of the sponge commences with
a small foreign body or even loose tissue in the centre, from which extend minute
straight canals about *5 mm. wide, in close proximity to each other, to the outer
surface. ‘The canals are circular or polygonal in section; their walls are formed by
the spicular skeleton. The individual element of the skeleton appears to have a
thickened subspherical centre, from which four to six short arms radiate in different
directions, and by their attachment to the ends of adjoining spicular arms form a
continuous, open, very regular mesh. The spicular arms or rays appear to be either
smooth or tuberculated. No special surface-spicules have been preserved.
Examples of these sponges occur in different states of preservation. Those from
Tennessee, which F. Roemer placed under Calamopora, are silicified, the spicular
* Annals & Mag. Nat. Hist. vol. vii. 1871, t. 7. f. 13, 14, and t. 8. f. 23.
58 SILICEOUS SPONGES.
mesh is for the most part dissolved away, and only empty moulds remain; in a few
instances, however, the spicules remain, and are now of a reddish material. The
canals have been filled with cherty silica, and the minute projecting rods connecting
adjoining canals have been produced by the silica filling in the minute spaces between
the spicular mesh. ‘This appearance Remer mistook for the infilling of connecting
tubes in the wall of a coral. The condition of these sponges from Tennessee is
precisely similar to that of specimens of Astylospongia ; and no doubt can be enter-
tained that, like this last-named genus, they were originally siliceous. The examples
from New Brunswick, however, have had their original skeleton replaced by calcite ;
and this fact led Prof. Duncan to believe that they were originally calcareous, so that
‘“‘there must have been a former mimetic and calcareous group of Spongida.” The
character of the spicules cannot be so clearly seen in the calcite specimens; and after
a close examination of the forms from Tennessee and New Brunswick, I believe that
the spicular elements resemble those of the Anomocladine family rather than of the
‘Tetracladina.
The ‘Tennessee examples appear to have been completely spherical, those from
New Brunswick are oblate spheroidal in form; the internal structure and dimensions
of the canals appear to be the same in the sponges from both localities.
Distribution. Silurian (Lower Helderberg Group): near Dalhousie, New Brunswick ;
Perry county, Tennessee (Pearson’s coll.). I have also seen specimens of the same
species from the Lower Helderberg group at Schoharie, New York.
Family TETRACLADINA, Zittel.
Genus AULOCOPIUM, Oswald, 1846.
AULOCOPIUM CYLINDRACEUM, Ferd. Ramer.
1861. Aulocopium cylindraceum, Remer, Fossile Fauna von Sadewitz, p. 9, t. 3. f. 2a, 2b.
1878. Aulocopium cylindraceum, Zittel, Studien, II Ab. p. 73.
Distribution. Silurian: Gotland.
Genus PHYMATELLA, Zittel, 1878.
PHYMATELLA INTUMESCENS, Raemer, sp.
1864. Eudea intumescens, F. A. Roemer, Palzont. Bd. 13, p. 26, t. 11. f. 1.
1878. Scyphia intumescens, Quenst. Petref. Bd. 5, p. 392, t. 133. f. 23-26.
1878. Phymatella intumescens, Zitt. Studien, IL Ab. p. 74.
There is but a single imperfect example of this species in the Museum collection,
which is cylindrical in form, tapering near the base, with irregular hollows in its
PHYMATELLA. 59
lower portion. It is 43 mm. in width; the cloaca is cylindrical, about 12 mm. in
width, and extends nearly to the base of the sponge. The exterior surface is thickly
covered with numerous irregularly disposed canal-apertures 1 mm. wide; the canals
extend in a nearly horizontal direction towards the centre of the sponge. The four-
armed spicules are slender with inconspicuous nodules. The specimen is preseryed
in Chalk, and the spicular structure has been changed into reddish peroxide
of iron.
Distribution. Upper Chalk: South of England.
PHYMATELLA HETEROPORA, Ramer, sp.
1840. Scyphia heteropora, Roem. Nordd. Kreide, 18> Ce ne tie diet
1878. Phymatella heteropora, Zitt. Studien, IL Ab. p. 74, t. 8. f. 2.
Detached spicules of the body and surface-layer of this species.
Distribution. Upper Chalk: Biewende, Brunswick (Zittel’s coll.).
PHYMATELLA RETICULATA, Hinde, n.sp. (Plate XI. figs. 1, 1a, 10.)
Sponges massive, either cylindrical, club-shaped, or obliquely turbinate, with a
depressed conical or truncated summit. The lower portion of the body gradually
tapers to a cylindrical stem, which terminates in a branching root. In some
specimens nodose or spur-like projections are irregularly scattered over the surface.
The surface exhibits a coarse irregular reticulation. There is considerable variation
both in the size and form; cylindrical specimens reach a length of 320 mm. by
72 mm. in width; the turbinate examples, though not so long, reach to a greater
width—one of these is 105 mm. in diameter.
The tubular cloaca in a large specimen is 24 mm. wide at the summit; from its
margins branching canals radiate down the summit of the sponge. The canals
extending from the outer surface to the interior are 1 mm. in width.
The internal skeletal structure of these sponges has been so greatly altered by
fossilization, that it has been very difficult to determine the original character of the
spicules. I have, however, been able to discover in a thin transverse section traces
of the straight arms and junction-nodes of tetracladine spicules; the spicular arms
are ‘225mm. in length by ‘053 mm. in width. The outer surface, when treated with
dilute acid, also shows indistinct traces of similar spicules.
These sponges have been referred to Phymatella intumescens, Roem., sp., from
which, however, they are distinguished by their massive walls, the coarse irregular
reticulation of the surface, and their generally large proportions.
Distribution. Upper Chalk: Flamborough, Yorkshire; South of England.
60 SILICEOUS SPONGES.
PuyMATELLA Noposa, Hinde, nu. sp. (Plate XI. figs. 2, 2 a.)
Sponge nearly cylindrical or tapering in form, the basal end rounded and destitute
of any peduncle or point of attachment; the summit of the specimen has not been
preserved. The surface in the upper portion is covered with obtusely rounded nodes ;
the lower portion is nearly smooth. The single imperfect example is 165 mm. in
length, 60 mm. wide at the summit, and 40 mm. at the base. The wall is 20 mm. in
thickness at the summit.
The cloacal tube extends for a distance of three fourths the length ; it is 21 mm.
wide at the top, and 10 mm. at the lower end. The canals opening into the cloaca
are 1-2 mm. wide, and they have a downward curve. The exterior surface of the
sponge exhibits apertures of various sizes, which appear to be the openings of canals
which extend in a curved direction towards the interior of the wall.
The spicular structure of the interior has been obliterated, but on the outer
surface in some places small tetracladine spicules can be seen ; the arms are smooth,
about ‘2 mm. in length; the nodes at the junction of the arms are inconspicuous.
In outer form this species somewhat resembles Phymatella heteromorpha*, Roemer,
sp., but the walls are much thicker and the canal-apertures smaller than in that
species. In the curved direction of the canals this species differs from the typical
forms of the genus, but in other respects its characters are similar.
Distribution. Upper Green Sand: Warminster.
é
PHYMATELLA, sp.
Distribution. Upper Chalk: South of England; France?
Genus AULAXINIA, Zittel, 1878.
AULAXINIA SULCIFERA, Ramer, sp.
Siphonocelia sulcifera, F. A. Roemer, Paleont. Bd. 13, p. 380, t. 11. £. 7.
Aulaxinia sulcifera, Zitt. Studien, II Ab. p. 75, t. 8. f. 4.
Distribution. Upper Chalk ?
AULAXINIA costata, Hinde, n. sp. (Plate XII. figs. 1, 1a.)
Sponge elongated, inversely conical, with rounded summit (no stem preserved) ;
length 80 mm. by 35 mm. in width. The cloacal aperture is 9 mm. in width; from
its margin a series of eighteen nearly straight simple canals, 2°3 mm. in width, and
equidistant from each other, extends to the base. Between these vertical canals are
the apertures (about 1 mm. in width) of the horizontal canals. The spicules are
relatively large, the arms apparently smooth and about 5 mm. in length.
* Paleontographica, Bd. 13, p. 22, t. 8. f. 11.
CALLOPEGMA. 61
The only example of this species is a somewhat worn specimen from flint-gravel.
The canals in this specimen have been infilled with silica, and now project as so
many ridges down the sides of the sponge. The spicules have also disappeared, but
their moulds in silica are distinctly shown.
This species may be distinguished from A. sulcifera by the greater number of the
vertical canals, also by their straight direction, and the larger size of the skeletal
spicules.
Distribution. Gravel, Upper Chalk: Stanway, Gloucestershire.
Genus CALLOPEGMA, Zittel, 1878.
CALLOPEGMA SCHLONBACHI, Zitt.
1878. Callopegma Schlinbachi, Zitt. Studien, II Ab. p. 75, t. 9. f. 1.
Detached skeletal spicules of this species.
Distribution. Upper Chalk: Ahlten, Hanover (Zittel’s coll.).
CALLOPEGMA oBcoNIcUM, Hinde, n. sp. (Plate XI. figs. 3, 5a, 5b.)
Sponges small, inversely conical, apparently sessile. The upper surface is flat, or
even in some examples slightly convex; the basal termination is obtusely rounded.
An average specimen is 20 mm. in height and 22 mm. in diameter.
In the central portion of the summit are the apertures of several canals, about
2 mm. each in width, which extend downwards into the sponge; there is also a
system of branching sinuous canals, seen in weathered specimens, which radiate from
the centre of the upper surface towards the sides of the sponge. The lateral surface
appears also to be furnished with minute pores.
The spicular structure, as seen in a vertical section, is of an open, irregular
character. The arms or rays of the spicules are about ‘6 mm. in length. No
dermal layer has been preserved in the specimens.
This species is usually preserved in the interior of flints, and the spicular structure
is seldom retained. In some instances, however, the spicules remain in the form
of iron peroxide. This form may be distinguished from Callopegma acaule, Zittel *,
by its smaller size, the absence of a cup in the centre, and its even surface.
Distribution. Upper Chalk: South of England.
CALLOPEGMA FICoIDEUM, Hinde, n. sp. (Plate XI. figs. 4, 4a, 4 8.)
Sponges fig-shaped or subspherical, with flattened or slightly rounded summits,
and with one or more straight or curved peduncular projections at the base, and
occasionally slight nodosities on the surface. An average specimen is 60 mm. in
height and 50 mm. in diameter.
* Studien, II Ab. p. 75, t. 2. f. 6 a,
62 SILICEOUS SPONGES.
Several vertical canals, 1‘8 mm. wide, open at the summit; the lateral surfaces
have scattered apertures of canals, *8 mm. in width.
The body-spicules have prominent junction-nodes and relatively long smooth arms.
The dermal layer is composed of the horizontally-expanded heads of large trifid
spicules, some of which are 1 mm. in extension. There are traces of minute spicular
bodies between the arms of the larger spicules of the dermal layer, but their form
has been lost.
The specimens only occur in the interior of flints; they retain the outer form and
the spicules of the dermal layer, as well as of the body-skeleton immediately beneath ;
but the interior is a mass of porous silica in which even the canals are destroyed.
This species differs from C. acaule, Zittel, principally in its rounded summit and
the absence of a cup.
Distribution. Upper Chalk: Croydon, Guildford, Surrey.
Genus TRACHYSYCON, Zttel, 1878.
Tracuysycon nobosum, Hinde, n. sp. (Plate XII. figs. 3, 3a, 30.)
Body of sponge massive, barrel-shaped, with a truncated summit bounded by a
subangular margin; the stem has not been preserved. The lateral surfaces are
covered with rounded or obtuse elevations disposed irregularly. ‘The only specimen
is 155 mm. in height and 117 mm. in width.
The cloaca is subcylindrical, somewhat wider below than at its aperture ; it extends
rather more than one third the length of the body; in the present specimen it is
55 mm. in length by 51 in width. ‘The interior surface of the cloaca is thickly
covered with the canal-apertures, which are apparently disposed in vertical series.
These apertures are 1°75 mm. in width. The canals from the basal portion of the
sponge-body run nearly straight to the cloaca; those from the lateral portion follow
a winding course; whilst the canals springing from near the upper margin of the
sponge are bent downwards, following the contour of the summit so as to open into
the upper portion of the cloaca. There is also a series of numerous fine canals,
‘75 mm. in width, extending from the exterior surface in a curved direction towards
the centre of the sponge.
The body-spicules are moderately large; their arms, *3 mm. in length, are smooth
or sometimes tuberculated; they are much branched, and the junction-nodules are
swollen. No surface-spicules have been preserved.
The single example of this species in the collection is filled with translucent chert,
in which the body-spicules are fairly well preserved; the exterior surface is rough,
and shows no structure.
Distribution. Upper Green Sand: Warminster.
TRACHYSYCON.—SIPHONIA. 63
TRACHYSYCON suLCATUM, Hinde, n.sp. (Plate XII. figs. 2, 2a, 26.)
Sponge massive, fig-shaped, widest near the summit, which is depressed or slightly
rounded; the stem has not been preserved. The exterior surface, with the exception
of the summit, which appears to have been smooth, is furnished with sinuous, inter-
rupted, vertically disposed ridges and furrows, and also with conical, slightly curved,
obtusely pointed, spinous processes, about 15 mm. in length, which have no definite
arrangement. An average specimen is 110 mm. in height and the same in
thickness.
A wide funnel-shaped cloaca extends nearly to the base of the sponge; numerous
slightly arched canals, 15 mm. in width, open into this cloaca; there are also sinuous
branched canals exposed on the summit, radiating from the cloacal margin. The
spicular skeleton of the interior is completely obliterated. ‘The surface appears to
have been covered with a dermal layer of minute irregular spicules with horizontal
sinuously branching head-rays.
There are two examples of this form in the collection, both of which are com-
pletely infilled with silex as well as partially enveloped in it. The species is
distinguished from the Zrachysycon (Plocoscyphia) muricatum, Remer*, by its
depressed summit, wide cloaca, and its ridged surface.
Distribution. Upper Chalk: South of England.
Genus SIPHONIA (Parkinson), pars, Goldfuss.
The authorship of this genus is involved in obscurity. The name first appears in
the ‘ Petrefacta Germaniz,’ p. 16, and is attributed by Goldfuss to Parkinson; but I
have been unable to find the term employed anywhere by this last author, though
he refers at considerable length in the second volume of his ‘Organic Remains’ to
sponges of this genus under the general term of Fossil Alcyonia. Most later authors
appear to have followed Goldfuss in ascribing the genus to Parkinson without in-
dicating where the generic characters were defined. Under these circumstances
Goldfuss must be considered the author of the genus, and the first species which he
describes, Siphonia piriformis, will have to be regarded as its type. Goldfuss has
included in Siphonia the sponges which Lamouroux placed under the genera
Hallirhoa and Jerea; and many later authors, including Prof. Sollas, who has given
a very elaborate description of the structure and affinities of Siphonia, adopt the
Goldfussian extension of the genus. I purpose, however, to follow the proposal of
Prof. Zittel to retain the genus Siphonia for such typical forms as Siphonia piri-
formis, tulipa, ficus, &c., and to place the lobate forms under the genus Hallirhoa,
Lamx., and the forms without a cloacal tube under Jerea, Lamx. It is worthy of
notice that if Hallirhoa or Jerea are to be considered as synonymous with Siphonia,
* Palewontographica, Bd, 13, p. 28, t. 10. f. 9.
64 SILICEOUS SPONGES.
then, by the rules of priority, the term Siphonia will have to give place to one or
the other of the genera of Lamouroux which were published anterior to Goldfuss’s
Stphonia.
The genus Choanites, Mantell, ‘Fossils of the South Downs,’ 1822, p. 178, is of a
later date than the Hallirhoa of Lamouroux, besides which the term is inapplicable
to sponges like Siphonia, for the first two species placed under the genus by
Mantell, Choanites subrotundus and flexuosus, are not Lithistid sponges at all, and
the type of Choanites is in reality a Hexactinellid sponge.
SIPHONIA PIRIFORMIS, Goldfuss.
1853. Siphonia piriformis, Goldfuss, Petref. Th. 1, p. 16, t. 6. f. 7 a.
1878. Siphonia piriformis, Zittel, Studien, II Ab. p. 79, t. 9. f. 7.
Non Siphonia pyriformis, Sowerby, Geol. Trans. 2nd ser. vol. iv. t. 15 a.
Non Siphonia pyriformis, Sollas, Quart. Journ. Geol. Soc. vol. xxxiu. t. 25. f. 1,
3, 4, 6, 8.
Distribution. Craie: Vaches Noires, Ranville, St. Adresse.
SIPHONIA TULIPA, Zittel. (Plate XIII. figs. 2, 2 a, 2b, 2c.)
1878. Siphonia tulipa, Zittel, Studien, II Ab. p. 79, t. 9. f. 5.
1836. Siphonia pyriformis, Sowerby, Geol. Trans. 2nd ser. vol. iv. p. 340, t. 15 a.
1854. Siphonia pyriformis, Mantell, Medals of Creation, p. 231, f. 1-3.
1877. Siphonia pyriformis, Sollas, Quart. Journ. Geol. Soe. vol. xxii, t. 25. f. 1, 3, 4, 6, 8.
1878. Siphonia Websteri, Quenst. Petref. Bd. 5, t. 135. f. 15-19.
Sponges simple or occasionally bifid, with conical or subcylindrical bodies, either
sharply constricted beneath the body or gradually tapering to a cylindrical stem
which usually branches at its termination. There is great variation in the size of
different specimens: a small form in the collection has the body only 21 mm. in
length by 13 mm. in width; a large specimen measures 105 mm. in length by
46 mm. in width.
The cloaca is a narrow elongated tapering tube; the aperture is narrow, with a
sharp margin, and about 8 mm. in width. It extends to the central or even the
lower portion of the body of the specimen. The arched canals which open into the
cloaca are about 1:25 mm. in width, whilst those extending from the outer surface to
the interior are very numerous, closely set, and average ‘5 mm. in width. Branching
canals radiating from the cloacal margins over the upper surface can also be distin-
guished in some examples.
The spicules forming the interior meshwork of the body are mostly smooth-armed,
though near the extremities the arms, together with the minute branches, are
frequently tuberculated. The spicular arms are ‘135 mm. in length, measuring from
the centre to the point where they commence to divide, and ‘052 mm. in thickness,
SIPHONTA. 65
The spicules of the stem are elongated and thread-like. No dermal layer has been
preserved.
This species is extremely abundant in the Upper Green Sand of Blackdown and
Warminster. The Blackdown examples usually have a sharp constriction between
the basal portion of the body of the sponge and the stem, which also is of a slender
character. The Warminster examples, on the other hand, have more elongated
bodies, which not infrequently gradually taper below to form a stem of considerable
robustness. There are, however, both at Blackdown and Warminster, specimens
exhibiting characters intermediate between the two extremes, so that it is quite
impracticable to limit this species merely to forms which possess the marked con-
striction between the body and the stem. In all other features the specimens from
these two localities appear to be similar.
Distribution. Upper Green Sand: Blackdown, Warminster.
SIpHONIA IncrassaTa, Goldfuss.
1833. Siphonia incrassata, Goldfuss, Petref. Th. 1, p. 17, t. 30. f. 5.
1878. Siphonia incrassata, Zittel, Studien, II Ab. p. 79.
Distribution. Planer-Kalk: Bohemia. Craie Chloritée: Vaches Noires; Sud-
merberg.
SipHonia Ficus, Goldfuss. (Plate XIII. figs. 3, 3a.)
1833. Siphonia ficus, Goldfuss, Petref. Th. 1, p. 221, t. 65. f. 14.
1878. Siphonia ficus, Zittel, Studien, II Ab. p. 79, t. 9. f. 6.
Sponges fig-shaped or subspherical, with gently rcunded summits, either apparently
sessile or with short branching stems. An average specimen is 75 mm. in height
and the same in width. The cloaca is cup- or funnel-shaped; it extends to nearly
the centre of the sponge; the margins are sharp. In an average specimen it is
23 mm. in width at the summit. ‘The interior surface exhibits the closely disposed
apertures (2°25 mm. each in width) of the canals which open into it. Branching
canals also radiate from the cloacal aperture over the surface. The spicular mesh is
entirely replaced by calcite, and the forms of the individual spicules are nearly
entirely obliterated. The outer surface of the specimens is now smooth and only
exhibits a delicate fibrous network.
Distribution. Grey Chalk: near Dover.
SrpHon1a Koénie1, Mantell, sp. (Plate XIII. figs. 4, 4 a.)
1822. Choanites Konigii, Mantell, Geol. of Sussex, p. 179, t. 16. f. 19, 20.
1808. A Flint, from a gravel-pit, Parkinson, Organic Remains, vol. ii. p. 100, t. 9. f. 1.
1850. Choanites Kénigi, Dixon, Foss. Sussex, t. 17. f. 1, 3, 4.
1854. Choanites Kénigi, Mantell, Medals of Creation, p. 234, f. 1, 3, 4.
66 SILICEOUS SPONGES.
1877. Siphonia Kénigii, Sollas, Quart. Journ. Geol. Soc. vol. xxxii. p. 817.
1878. Siphonia Koenigi, Zitt. Stud. Il Ab. p. 79.
Sponges pear-shaped or subspherical, the summits usually truncate; stem cylin-
drical, apparently slender. An average specimen is 100 mm. in height by 70 mm.
in width.
The cloacal tube is cylindrical or funnel-shaped, and extends nearly to the basal
portion of the body. Its aperture is very wide, and the margins are rounded. Very
strongly marked branching canals, 2 mm. in width, radiate from the margin of the
cloaca down the sides of the sponge. The lateral surfaces exhibit circular or sub-
angular apertures of canals from | to 2 mm. in width. Some of these apertures are
bridged over by minute thread-like extensions of spicular fibre crossing them.
The spicules forming the mesh are relatively large, but they are so incrusted with
silica that it is impracticable to measure them. A dermal layer appears to have
extended over the outer surface of the sponge and over the interior of the cloacal
tube as well. The spicules composing it have horizontally expanded, slightly bifur-
cate head-rays; the shafts, if present, extended into the sponge.
Most of the examples of this species are preserved in the interior of flints, and
retain merely the form of the cloaca and the larger canals, the spicular structure
having entirely disappeared. There are, however, in the Museum collection a few
specimens from Flamborough which, in common with the other sponges from that
locality, are preserved in a chalky matrix, which can be removed by treatment with
dilute acid ; and these exhibit the form and canal-structure in a very perfect manner,
but the spicules for the most part are masked by siliceous accretions. They further
show the important feature of a dermal layer, which has hitherto not been discovered
in any example of the genus. This layer covered over the exterior branching summit
canals, and seems also to have covered the interior surface of the cloaca, for in a
well-preserved specimen the cloaca exhibits a smooth surface like that of the exterior,
and here and there the openings of the canals can be detected beneath this layer.
In some of the examples of this species, preserved in flint, there is a filled-up
spiral tube or canal, from 4 to 8 mm. in width, which winds through the interior of
the body between the cloaca and the outer surface, and opens out at the summit of
the sponge in the vicinity of the cloacal aperture. This tube has been described by
Cunnington * as forming part of the original fabric of the sponge, and probably
connected with the reproductive system. ‘The more probable suggestion is that the
tube has been formed by an annelid, which has built its shell pari passu with the
growth of the sponge; and it is not improbable that its constant proximity to the
cloacal aperture has arisen from the support which the animal inhabiting the tube
may have derived from the stream of water passing out of the cloaca. Examples in
the Museum show clearly that this spiral tube was not peculiar to this species of
* Report of British Association, Swansea, 1848, p. 67.
SIPHONIA.—HALLIRHOA. 67
Siphonia, for there are cylindrical sponges, probably belonging to Scytalia or Phy-
matella, in which it is also present.
This species appears to be very abundant in the Upper Chalk of this country. It
is very probable that most of the sponges so common in the flints at Brighton belong
to this species. ‘The flattened summit and the wide aperture and rounded margins
of the cloaca readily distinguish the form from the S. ficus.
Distribution. Upper Chalk: Flamborough, Yorkshire; near Brighton, and in other
localities in the South of England,
SIPHONIA, sp.
Distribution. Upper Chalk: Beckhampton. Chalk Marl: Ringmer.
Subgenus HaLuirnoa, Lamourour, 1821.
Hanurwoa costata, Lama. (Plate XIV. figs. 1, la, 10, 1¢, 1d, 1e.)
1821. Hallirhoa costata, Lamx. Exp. méthod. des Polypiers, p.72, t. 78. f. 1.
1831. Polypothecia bi-septemloba, KE. Benett, Org. Rem. Wilts. t. 1-5.
1847. Hallirhoa costata, Michelin, Icon. Zooph. p. 127, t. 31. f. 3.
1854. Siphonia lobata, Mantell, Medals of Creation, vol. i. p. 231, f. 4.
1878. Hallirhoa costata, Quenstedt, Petref. vol. v. p. 426, t. 135. f. 14.
1878. Hallirhoa costata, Zittel, Studien, Il Ab. p. 79.
Sponges simple, very variable in the form of their bodies or heads: the simpler
examples are conical and partially lobed at the base; the more complex are depressed
spheroidal, and divided vertically into a varying number (from three to seven) of
lobes, which occasionally show traces of subdivision. ‘The summit in the lobed
individuals is flattened or gently rounded. ‘The stem is relatively slender, crooked,
and with root-like processes at its extremity. The body of a small specimen is
42 mm. in height by 57 mm. in width, whilst a large individual measures 78 mm.
in height by 150 mm. in width. The longest stem which I have seen measures
180 mm.
The cloaca is cylindrical or funnel-shaped, with angular margins; its depth varies
in different individuals—in some examples it is not more than 15 mm. below the
surface, or about one fourth the vertical height of the body, whilst in others it
appears to penetrate to the central portion of the body. The larger canals are
15 mm. in width; those immediately beneath the cloaca are straight ; exterior to
these the canals follow a curved course to the cloaca. The smaller canals, which
extend in an arched direction from the exterior surface to the interior of the sponge,
are “7 mm. each in width. In weathered specimens numerous canals are exposed on
the upper surface, radiating from the margins of the cloaca.
The spicules forming the interior mesh have mostly smooth arms, but. their
K2
68 SILICEOUS SPONGES.
branching extremities are thickly covered with minute tubercles. The spicular
arms are ‘202 mm. in length by ‘067 mm. in thickness.
The extreme variability in the form of this species is well exemplified in the
numerous specimens in the Museum.
Distribution. Upper Green Sand: Warminster, Wiltshire. Red Chalk: Hun-
stanton, Norfolk (Prof. Seeley). Craie Chloritée: Vaches Noires, Havre.
HALLirHoA costata, var. BREVICosTATA, Michelin. (Plate XIV. fig. 2.
1847. Hallirhoa brevicostata, Michelin, Icon. Zooph. p. 127, t. 31. f. 2.
1878. Hallirhoa brevicostata, Zittel, Studien, II Ab. p. 79.
Sponge in form like a partially opened mushroom, the summit depressed, the sides
indistinctly divided into five or six shallow lobes, the stem thick at its junction with
the body and gradually tapering. The only example in the collection is 39 mm.
in height and 75 mm. in width. This form can hardly be regarded as more than
a variety of H. costata. It differs from the typical forms of the species in the slight
degree in which it is lobed and in the thickened stem. Michelin has erroneously
referred Hallirhoa (Polypothecia) agariciformis, Benett, to this variety, but the two
forms are very distinct from each other.
Distribution. Upper Green Sand: Warminster, Wiltshire.
HAauiruoa Costata, var. Tessonis, Michelin.
1847. Hallirhoa Tessonis, Michelin, Icon. Zooph. p. 128, t. 34. f. 1.
1878. Hallirhoa Tessonis, Zittel, Studien, II Ab. p. 79.
This variety only differs from the typical forms of H. costata in the gradual
tapering of the body of the sponge to the stem, instead of the sharp constriction
usually present between the base of the body and the summit of the stem.
Distribution. Craie Chloritée: Vaches Noires, near Havre (Tesson collection).
HALLIRHOA cosTaTa, var. ELEVATA, [/inde. (Plate XIV. fig. 3.)
In this variety the body of the sponge is formed by five subequal, elongated,
compressed lobes; the central portion of the summit is distinctly elevated, with the
cloacal aperture at the top. The upper margins of the lobes are separate from each
other, and more compressed than the sides. The cloacal aperture is 15 mm. in
width. The single example is 110 mm. in height and 100 mm. in width. This
variety conspicuously differs from the typical lobed forms of H. costata in the elevated
summit and in the narrow upper margins of the lobes.
Distribution. Upper Green Sand: Warminster.
HALLIRHOA. 69
HALLIRHOA AGARICIFORMIS, Benett, sp. (Plate XV. figs. 1, 1 a, 14.)
1831. Polypothecia agariciformis, Benett, Cat. Org. Rem: Wilts. t. 15. f. 1, 2.
q 1847. Siphonia acaulis, Michelin, Icon. Zoophy. p. 139, t. 38. f. 2.
1847. Non Hallirhoa brevicostata, Mich. Icon. Zoophy. p. 127, t. 31. f. 2.
1854. Hippalimus fungoides, Morris, Cat. Brit. Foss. p. 28.
Sponges more or less depressed, conical, or mushroom-shaped, with generally an
even slope from the cloaca to the exterior margin, which is frequently irregular in
outline, more or less circular, occasionally thin, and not infrequently with finger- or
spur-like processes extending outwards. The under surface of the sponge is hollowed
out and intersected with very prominent sinuous ridges, distributed in an irregular
manner. In many specimens there are no traces of a stem, and the sponge is
apparently sessile; in others, however, there is a small cylindrical process in the
centre of the underside, which may represent the upper portion of a slender stem.
There is considerable variation in the size of different examples—a small specimen
measures 40 mm. in height by 75 mm. in width, and a large form is 80 mm. in height
and 155 mm. across the base.
The cloaca is cylindrical and, as a rule, does not extend deep into the body. It
is situated at the summit of the cone, its margins are sharp, and its interior surface
is covered with the closely set apertures of canals, which, in some examples at least,
are disposed in vertical rows. The upper surface of the sponge is covered with
numerous, slightly sinuous, branching canals, 1-5 mm. in width, which radiate down-
wards from the margins of the cloaca towards the exterior edge. ‘These canals are
generally open, but were in all probability covered by a dermal layer. In a vertical
section canals similar in size and direction to those of the surface are exposed. ‘The
basal surface of the sponge exhibits numerous irregularly shaped canal-apertures ;
there are also indications of a dermal layer extending over the under surface.
The spicules of the interior mesh have robust smooth arms, with tuberculated
extremities which interlock together. The arms of the spicules are ‘38 mm. in
length and -078 mm. in thickness. A small portion of the surface of a specimen
belonging to the Jermyn-Street Museum is covered with a dermal layer of minute,
flattened, deeply laciniated spicules about °65 mm. in width, which are so arranged
that their arms overlap each other. ‘These spicules are furnished with shafts which
extend into the sponge. I have not been able to detach any of the spicules to
ascertain the length of the shaft, but its presence is clearly indicated by the minute
aperture which is seen where the heads of the dermal spicules have fallen off the
surface. Where this dermal layer is present the surface of the sponge is smooth,
and the underlying canals are quite concealed from view.
The examples of this species appear to have been abundant in the Upper Green
Sand of certain localities. Miss Benett’s figures very accurately represent the outer
surface of this species; notwithstanding this, Michelin relegated her forms to his
70 SILICEOUS SPONGES.
Hallirhoa brevicostata, which they do not at all resemble. On the other hand, this
same author has given the name of Siphonia acaulis to an example of this species
from Cap la Héve. Examples from this locality in the Museum collection distinctly
show the resemblance to specimens from Warminster, in spite of the different aspect
resulting from fossilization. This species has also been referred to Hippalimus
fungoides, Lamx.; but though [ have not seen an example of this form, the descrip-
tion and figure of Lamouroux represent an altogether different sponge from the
present species.
Distribution. Upper Green Sand: Warminster; Ventnor, Isle of Wight; Cap la
Héve, Seine-Inférieure.
Genus JEREA, Lamourouxr, 1821. ,
JEREA PYRIFORMIS, Lamz.
1821. Jerea pyriformis, Lamx. Exp. méthod. p. 79, t. 78. f. 3.
1847. Jerea pyriformis, Mich. Icon. Zooph. p. 133, t. 36. f. 3.
1878. Jerea pyriformis, Zitt. Studien, II Ab. p. 81.
Distribution. Craie Chloritée: Vaches Noires, Havre.
JEREA WepsteRI, Sowerby, sp., MS.?
1814. Tulip Alcyonium, Webster, Trans. Geol. Soc. vol. ii. p. 377, t. 28.
1854. Siphonia Websteri, Morris, Cat. Brit. Foss. p. 30.
Sponges pear-shaped or ovate in form, with a conical summit, and gradually
tapering to a somewhat slender stem at the base. In none of the Museum
specimens has the stem been preserved; but according to Webster’s figures it
appears to have been unusually long. The body of a small individual is 48 mm.
in length by 20 mm. in width, whilst a large specimen measures 81 mm. by 51 mm.
At the summit is a closely arranged group of canal-apertures 1-5 mm. each in
width, which open directly at the surface and not in any depression. Down the
sides are shown in some specimens longitudinal, sinuous, branching canals. The
spicular structure is not clearly exposed.
Prof. Morris quotes Sowerby as the author of this species, but I have not been able
to discover the published description.
Distribution. Upper Green Sand: Warminster; Isle of Wight.
JEREA RETICULATA, Hinde, n. sp. (Plate XV. figs. 2, 2a, 26.)
Sponges fig-shaped or subspherical, with slightly truncated summits, either with
a short peduncular stem or apparently sessile. A small specimen is 26 mm. in height
by 21 mm. in width, whilst a fairly large globular form is 35 mm. in height by 42 mm.
in width.
JEREA. 71
In the shallow depression at the summit of the sponge are the apertures of a group
of canals, about 1 mm. each in width. From the summit there extends over the
surface a series of canals which apparently anastomose with each other, and give a
reticulate appearance to the sponge. In a vertical median section the only canals
distinctly shown extend in an arched direction from the surface to the centre of the
sponge. These canals appear to be very wide near the outer surface, and to open
directly into the reticulate canals. The interspaces of the surface between the
reticulations appear to have no other apertures beyond the openings of the spicular
mesh.
The interior skeleton is built up of robust spicules with smooth arms and prominent
botryoidal nodes at their junction with each other. No dermal layer has been
preserved.
This species may be recognized by the reticulated surface and the peculiar character
of the canals extending from it to the centre of the sponge.
Distribution. Upper Green Sand: Warminster, Blackdown; Cap la Heve; Retsel,
Ardennes. Craie Chloritée: Honfleur, Vaches Noires. Gault? near Folkestone.
JEREA QuensteDtTI, Zittel.
1878. Jerea Quenstedti, Zitt. Studien, II Ab. p. 81, t. 10. f. 2.
1878. Siphonia ficus, Quenst. Petref. Bd. 5, p. 431, t. 185. f. 20-23.
Microscopic slides with spicules of this species.
Distribution. Quadraten-Kreide: Linden, Hanover.
JEREA CORDIFoRMIS, Hinde, n. sp. (Plate XV. figs. 5, 5a.)
Sponge obcordate in form, with an evenly rounded summit, constricted at the base,
and apparently supported on a stout stem. The only specimen in the collection is
93 mm. in height and 73 mm. in width.
The somewhat elongated rounded ridge at the summit is pierced by numerous
apertures of the vertical series of canals, 2mm. in width. From the summit also,
open, sinuous, branched canals extend down the sides of the sponge. The canals
extending from the surface towards the interior are greatly arched, and | mm. in
width.
As is usual in all the specimens from the Grey Chalk, no spicules have been
preserved in this form, but prominent junction-nodes can be recognized; and I have
no doubt that the sponge belongs to the Tetracladina family.
In its peculiar form and in the character of the summit this sponge is distinct from
any hitherto referred to this genus.
Distribution. Grey Chalk: near Dover.
~I
bo
SILICEOUS SPONGES.
JEREA EXCAVATA?, Michelin.
Jerea excavata, Michelin, Icon. Zooph. p. 135, t. 39. f. 2,
Jerea excavata, Zittel, Studien, II Ab. p. 81, t. 10. f. 1.
Rhysospongia pictonica, Court. Kp. foss. t. 1. f. 1, 2.
Sponge massive, hemispherical, the upper surface slightly concave, the under
surface rounded, apparently sessile. Height 70 mm., width 138 mm.
Numerous canal-apertures, 3°5 mm. in width, open into the shallow depression of
the summit and also on the sides. ‘The only specimen is a solid mass of flint which
merely retains the outer form and the canal-apertures. In the absence of the spicular
structure, it can only be doubtfully referred to Michelin’s species.
Distribution. Upper Chalk: Wiltshire.
Genus NELUMBIA, Pomel, 1872.
NELUMBIA TUBEROSA, Hinde, n.sp. (Plate XVI. figs. 1, la, 16, 1c.)
Sponge club-shaped, with an irregularly truncated summit, gradually tapering at
the basal end to form root-like processes. A few irregularly scattered nodose
projections are present on the surface. Length 180 mm., width 54 mm.
Numerous circular or polygonal canals, 3 mm. wide, open at the summit, and
extend vertically nearly to the base of the sponge. Another series of canals, 1 mm.
wide, extend from the lateral surfaces, in a slightly arched direction, to the central
portion of the sponge.
The spicules of the interior mesh, though now replaced by calcite, can be fairly
recognized. ‘They appear to possess smooth robust arms, -27 mm. in length by
045 mm. in width, and prominent junction-nodes. The surface appears to have
been completely covered with a delicate dermal layer principally composed of the
horizontally extended heads of trifid spicules with bifurcate, straight, pointed rays.
Some of the spicular heads are °6 mm. in width.
In the size and form of the specimen itself, and of the canals, this species differs
from the different forms figured by Courtiller* under the name of Polystoma, which
have been placed by Pomel as the types of the genus Nelumbia. In Pomel’st+
definition it is stated that the vertical canals do not penetrate deeply below the
summit of the sponge; but, as already indicated by Zittel, it seems very probable
that he is mistaken on this point, for in the figures of the type species the canals are
shown, in the section of at least one specimen, to extend to the basal portion of
the sponge.
Distribution. Grey Chalk: near Dover. Upper Chalk: South of England.
* Eponges fossiles, p, 10, t. 15.
+ Paléontologie d’Oran, p. 194.
=I
oo
POLYJEREA.—BOLOSPONGIA.
Genus POLYJEREA, Fromentel emend. Zittel.
POLYJEREA ARBUSCULA, Hinde, n. sp. (Plate XVI. figs. 2, 2 a.)
Sponge massive, growing from a simple, compressed stem, from which spring
upright, straight, or slightly curved, cylindrical, compressed, or club-shaped branches,
which also subdivide. ‘The branches are mostly separate, but occasionally coalesce
together; they average 36 mm. in thickness. At the summit of the branches are the
apertures of numerous canals, about 2 mm. in width, which extend vertically down
the branches. The lateral surfaces are covered with smaller apertures of canals,
1 mm. in width, which extend towards the centre,of the branch.
The spicules of the interior mesh have smooth arms and prominent nodes.
Occasionally traces of the dermal layer are preserved; it consisted, in part at least,
of trifid spicules, with horizontal heads of pointed rays, similar to those of Nelumbia
tuberosa.
I am unable to determine how far this species differs from Polyjerea ramifera,
Zitt.*, in its mode of growth. By the form and disposition of the branches it may
readily be distinguished from Polyjerea gregaria, Mich.} sp.
Distribution. Upper Green Sand: Warminster. Grey Chalk: near Dover.
POLYJEREA LOBATA, Hinde, n. sp. (Plate XVI. fig. 3.)
Sponge consisting of a compact group of fig- or barrel-shaped, or occasionally
cylindrical individuals, growing from a flattened base or from a slight peduncle.
The outer surface of the group is partially lobed, but in the central portion the
individuals are closely amalgamated together ; the summits are, however, free. The
number of individuals in each compound mass varies between six and twenty. In size
the compound form is from 90 to 105 mm. in width, and about 64 mm. in height.
The summit of each individual is truncate, or has a slight shallow depression ; the
canals opening at the summit are 2 mm. in width, and extend nearly to the base of
the sponge. The spicules of the interior mesh have been replaced by calcite in the
specimens from the Grey Chalk; in the forms from the Upper Green Sand they have
relatively short, smooth arms and prominent junction-nodes. Only traces of a dermal
layer have been preserved ; it resembles in the form of its larger spicules that of the
last species.
Distribution. Upper Green Sand: Warminster. Grey Chalk: near Dover.
Genus BOLOSPONGIA, Hinde, n. g.
Sponges subspherical or elongate and lobate. No regular series of canals is
present ; in the rounded forms there are interior loculi which communicate with the
* Studien, II Ab. p. 83. + Icon. Zooph. p. 134, t. 38. f. 1.
L
74 SILICEOUS SPONGES.
exterior by wide channels; in the elongate examples the lobes are deeply constricted,
and no definite canals are apparent. The spicular mesh of the interior is composed
of four-rayed spicules, with apparently smooth arms and prominent nodes at their
junction with each other. A dermal layer is partially preserved, but its component
spicules are not recognizable.
I propose this genus for a small group of sponges which, in external form, closely
resemble certain species of Astrobolia, Zittel. In this genus, however, the spicules
are of the Rhizomorine type, whereas in Bolospongia they are distinctly Tetracladine.
The specimens are from the Upper Chalk of Flamborough, and, in common with all
the examples from this locality, their spicular structure has been mostly destroyed.
Bo.osponeia eLopata, Hinde, n.sp. (Plate XVII. figs. 1, 1 a, 10.)
Sponges subspherical, closely resembling a potato in form, the surface uneven, with
slight rounded elevations and intervening depressions; in some examples there is a
minute peduncle not more than 5mm. in length; other specimens are apparently
entirely free. An average example is 45 mm. in diameter.
On the surface of specimens which have been treated with acid there are exhibited
in places small circular apertures about 2°5 mm. in width, and occasionally a few
canals radiate from these over the surface. In a vertical section irregular channels,
6mm. wide, and open spaces are exposed. The spicules have been replaced by
crystalline calcite and silica, but their rough outlines can be distinctly seen in thin
sections. ‘The arms are relatively large and robust, °315 mm. in length by ‘067 mm.
in width ; some of the nodes are ‘27 mm. in thickness.
Iistribution. Upper Chalk: Flamborough, Yorkshire.
Bo.osronela constricta, Hinde, n. sp. (Plate XVII. fig. 2, 2 a.)
Sponge elongate, consisting of a series of irregularly shaped lobes, the constrictions
between which extend to the central portion of the sponge. ‘There are a few traces
of surface-canals ; oscules are not apparent, and the circulation seems to have been
carried on in the channels between the lobes. The spicular mesh is of a loose,
open character ; the spicules are similar to those of B. globata. A compact dermal
layer apparently covered the outer surface, but the form of its spicules cannot be
recognized.
There is but one specimen of this singular sponge in the Collection. It is 108 mm.
in length by 42 mm. in width.
Distribution. Upper Chalk: Flamborough.
THECOSIPHONIA. 75
Genus THECOSIPHONIA, Zittel, 1878.
'THECOSIPHONIA NOBILIS, Remer, sp. (Plate XVII. figs. 3, 3 a.)
1864. Limnorea nobilis, F. A. Roemer, Paleont. Bd. 13, p. 37, t. 15. f. 1.
1878. Limnorea nobilis, Quenst. Petref. Bd. 5, t. 133. f. 8-11.
1878. Thecosiphonia nobilis, Zittel, Studien, II Ab. p. 84.
The only example of this species is a compound form, 72 mm. in height by 90 mm.
in width, consisting of three individuals closely aggregated together, and united by a
common dermal layer. ‘The individuals are roughly ovate in form; they appear to
have had one or two cloaca-like depressions on the summit; the canal-structure of
the interior has been destroyed. Some of the spicules of the interior mesh can be
‘ seen; they are relatively very large, the arms and nodes measuring ‘5 mm. in length ;
the nodes are very prominent.
The dermal layer is thin, and grows in concentric, wrinkled bands; it appears to
have originally extended all over the sponge, with the exception of the summit; its
surface is smooth, and contrasts strongly with the rough furrowed character of the
summit. I have been unable to determine the form of the spicules composing this
membrane.
Distribution. Upper Chalk: Wiltshire (Cunnington coll.). The example belongs
to the Jermyn-Street Museum.
THECOSIPHONIA TURBINATA, Hinde, n. sp. (Plate XVII. fig. 4.)
Sponge turbinate in form; the upper portion or body is depressed, conical, with a
wide base supported on a short, oblique, obtusely terminated stem. The base is
concentrically furrowed and wrinkled, and carries one or two short radical processes.
The single example is 85 mm. in height and 100 mm. in width.
The upper surface of the sponge is rough; the cloaca at the summit is 26 mm. in
width, and does not appear to have been deep; from its margins sinuous branching
canals radiate over the surface. The interior canals have been destroyed.
Only here and there can the four-rayed spicules of the mesh be detected; they are
smaller than those of 7. nobilis. The under surface and the stem appear to have
been completely covered with the dermal layer, but its spicular elements cannot be
recognized.
The only specimen is now a mass of flint, in which only the outer form, the dermal
layer, and a few of the interior spicules are preserved. In its form and in the
restriction of the dermal layer to the basal portion it differs from Thecosiphonia
(Tremospongia) grandis, Roemer*.
Distribution. Upper Chalk: Stockton, near Heytesbury, Wiltshire.
* Palwxontogr. Bd. 13, Taf. 15. fig. 3c.
L2
76 SILICEOUS SPONGES.
Genus CALYMMATINA, Zittel, 1878.
CALYMMATINA RIMOSA, Zittel.
1878. Calymmatina rimosa, Zittel, Studien, II Ab. p. 85, t. 2. f. 2, and t. 9. f. 8.
There is a single specimen in the Collection which, so far as can be determined
from its imperfect preservation, appears to belong to the above species. Nothing is
known as to the locality whence it comes; judging from the matrix it would seem
to have been derived from the Upper Chalk.
Genus 'TURONIA, Michelin, 1847.
TURONIA VARIABILIS, Michelin. (Plate XVIII. fig. 1.)
1847. Turonia variabilis, Mich. Icon. Zooph. p. 126, t. 35. £. 1-8.
1861. Turonia variabilis et sulcata, Court. Eponges foss. p. 25, t. 40. f. 1-3.
1878. Turonia variabilis, Zittel, Studien, II Ab. p. 86.
Sponges conical in form, with a flattened or rounded base, from which one or more
radical processes project. A small example is 21 mm. in height by 25 mm. in width ;
another specimen is 52 mm. in height by 42 mm. in width.
The upper, distinctly conical portion of the sponge has a tubular cloaca, which
opens at the summit of the cone; the sides are rough and deeply furrowed with
open canals, which radiate downwards from the edge of the cloaca; in these canals
are occasionally seen the apertures of smaller canals, which penetrate to the interior
of the sponge. The lower margin of the upper cone overlaps slightly the upper
portion of the base, and is usually distinctly marked off from it by a well-defined
furrow. The basal portion is smooth and compact.
‘The mesh-spicules have robust arms and branching extremities. The dermal layer
is partly composed of very minute irregular-shaped spicules, and partly of minute
trifid spicules with delicate horizontally expanded head-rays.
This species is rare in this country. I have only seen two imperfect examples; it
appears to be abundant in the neighbourhood of Tours. The specimen figured is
probably from France ; the particular locality is unknown.
Distribution. Upper Chalk: Flamborough, Yorkshire ; South of England; France.
Genus KALPINELLA, Hinde, n. g.
Sponges simple, cup- or vase-shaped, supported on a cylindrical or compressed
stem, with root-like processes at its termination. .Margins rounded. Both the
interior and exterior surfaces of the sponge-wall with numerous canal-apertures.
The canals are somewhat sinuous and extend through the wall at right angles or
obliquely to the surface. Vertical canals also extend down the stem.
KALPINELLA. 77
The mesh-structure of the interior is relatively close, composed of small four-
rayed spicules with short, robust, smooth arms, and branching tuberculated extre-
mities. No dermal layer has been preserved.
I place in this genus a group of sponges which in form closely resemble some
species of Chenendopora, and can only be distinguished therefrom by an examination
of the spicular structure, which is distinctly Tetracladine. The absence of the
dermal layer prevents a comparison with the recent genus Discodermia; there is
hardly a doubt that the fossil forms were originally furnished with a surface-layer ;
and should future discovery show that its spicular structure resembles that of Disco-
dermia, the present genus will have to be relinquished, as the interior spicular
structure is very similar to that of the recent genus.
With one doubtful exception from the Grey Chalk, all the sponges of this genus
come from the Upper Green Sand, and, in common with most of the examples from
the same beds, retain the spicular mesh of the interior in good preservation, but the
minute structure of the exterior surface is obliterated.
KALPINELLA PATER/FoRMIS, Hinde, n.sp. (Plate XVIII. fig. 4, and
Plate XIX. figs. 1, 1 a, 10.)
1831. Polypothecia expansa ?, Benett, Cat. Org. Rem. Wilts. t. 6. f. 2.
Sponges cup-shaped, in some cases shallow and expanded, in others moderately
deep. The stem is either short and compressed or elongated and cylindrical. The
margins of the cup are rounded, and sometimes exhibit open canals. The wall
varies between 12 and 18 mm. in thickness in different specimens. A small example
is 70 mm. in width across the cup, and a large specimen 110mm. The longest stem
preserved measures 93 mm.
The canal-apertures are from 1 to 15 mm. in width; they are nearly equal in
size, so far as can be ascertained, on both the upper and under surfaces of the cup-
wall; in some specimens the canals of the under surface open very obliquely
downwards.
The spicular rays of the interior are very irregular in length, and from °02 to
‘04 mm. in thickness. The extremity of the rays is developed into a bunch of
tubercles, and by the interlocking of the tubercles of adjoining rays the mesh is
built up. In some of the spicules the canals of the interior are still preserved ; they
extend from the centre of the spicule to about half or two thirds the length of the
arms, and appear as very minute cylindrical tubes.
In external form some of the specimens resemble the Polypothecia expansa, Benett ;
but in the absence of all description of the internal structure of this species, I have
deemed it best to place these forms under a fresh designation.
Distribution. Upper Green Sand: Warminster. Doubtfully from the Grey Chalk
near Folkestone. ©
78 SILICEOUS SPONGES.
KALPINELLA RUGOSA, Hinde, n.sp. (Plate XIX. fig. 2.)
Sponge massive, expanded cup- or vase-shaped, with corrugated and folded walls
in the upper portion of the cup, the under surface either smooth or with nodose
outgrowths. Only the upper portion of the simple cylindrical stem preserved. The
walls vary from 17 to 23 mm. in thickness. A fairly large specimen is 230 mm.
wide at the summit of the cup, and 110 mm. in height.
The canal and spicular structures resemble those of the last species, from which this
is mainly distinguished by its thicker and folded walls and its larger dimensions.
Distribution. Upper Green Sand: Warminster, Burbage, Wiltshire.
Genus THAMNOSPONGIA, Hinde, n. g.
Sponges growing in the form of simple upright branching stems, or forming bush-
like masses. The branches cylindrical or rarely compressed.
With the exception of one species, in which canals do not appear to be present,
these sponges are traversed longitudinally by one or more canals.
The interior skeleton forms a close mesh, composed of very minute spicules with
short stumpy arms. The arms are covered with relatively large prominent tubercles,
in such a manner that they appear in some instances to be made up of closely set
tubercles or rings. These spicules appear to be connected together either by the
interlocking of the tubercles of adjoining spicules or by the adpression of the
extremities of the spicular arms, but they do not form distinct nodes at their points
of junction with each other.
I propose this genus for a group of sponges which closely resemble, as regards
their form and mode of growth, some species of Polyjerea and Astrocladia, but
present a very different form of mesh-spicule. In both these last-named genera the
spicules of the mesh have smooth arms and form prominent junction-nodes; but in
the present genus the spicules are tuberculated throughout, and the nodes are incon-
spicuous. ‘The characters of the dermal layer also vary from those of Astrocladia,
in which no regular trifid spicules are present, whilst these form well-recognized
features in the dermal layer of this genus. The character of the interior spicules,
and their mode of union with each other, exhibit some similarity to the spicular
structure in Plinthosella ; but the mesh is much closer and the spicules are far smaller
than in any sponge of that genus, whilst the form of the spicules of the dermal
layers is altogether distinct.
The examples of this genus appear to be abundant in the Upper Chalk, and they
usually occur in the interior of flints, frequently loose, so that when the flint is
broken the sponge may be extracted. Asa rule, the interior skeletal structure is
destroyed, but the dermal layer is partially preserved, and the spicules in it can be
determined. I refer, provisionally, to this genus also a specimen from the Grey
THAMNOSPONGIA. 79
Chalk, which differs from the Upper-Chalk specimens in the characters of the
interior canals.
THAMNOSPONGIA GLABRA, Hinde, n.sp. (Plate XVII. figs. 5, 5a, 56, 5c.)
Sponge growing in bushy masses, consisting of cylindrical upright or bent stems,
which give off irregularly, at an open angle, short curved branches with conical oy
blunted extremities. A single mass attains a height of 160 mm.; the stems are from
8 to 11 mm. in thickness; the lateral branches are somewhat less. The surface of the
sponge in many specimens is extremely smooth, but in others, where the dermal
layer is absent, it is rough.
The canal-structure in all the specimens is very imperfectly preserved, and [ am
unable to affirm the presence of more than a single longitudinal canal, which is
faintly apparent in the centre of some of the stems, and is about 1:75 mm. in width ;
it is possible, however, that other canals would be seen in well-preserved specimens.
The terminals of most of the branches do not exhibit canal-apertures.
The interior skeletal mesh is formed of very minute tuberculated spicules, with
arms or rays about 16 mm. in length. The dermal layer is smooth and compact ;
the heads of the largest of the trifid spicules are *3 mm. in width; between the larger
forms are very numerous small ones. Asa rule, the larger spicules have fallen off
the surface of the dermal layer, leaving, however, distinct impressions of their form,
and of the shaft of the spicule, in the centre of each. In two small branches of a
sponge, apparently belonging to this species, which Mr. H. J. Carter, F.R.S., has
presented to the Museum, the smooth dermal layer exhibits in places minute groups
of circular pores. There are about ten of these pores in a group, which is only
-3 mm. across; the groups are once or twice their own width apart. The pores are
close together, and their margins are smooth and well defined.
This species differs from the next in its mode of growth and in the characters of
the dermal layer, the spicules of which are much smaller than in T. clavellata. It
appears to be not uncommon in the Upper-Chalk flints of the south-west of
England.
Distribution. Upper Chalk: Woodford, Beckhampton, Wiltshire; near Budleigh
Salterton, Devonshire, in superficial detritus (Mr. H. J. Carter).
THAMNOSPONGIA CLAVELLATA, Benett, sp. (Plate XVIII. figs. 2, 2a, 26.)
1831. Polypothecia clavellata, Benett, Cat. Org. Rem. Wilts. t. 12.
1854. Spongites clavellatus, Mantell, Medals of Creation, vol. i. p. 224, f. 2.
Sponge generally consisting of a single straight or slightly sinuous stem, which
gives off, mostly at right angles, short stumpy branches or spur-like processes, either
blunted or rarely bulbous at their extremities. In some cases the branches extend
to the adjoining stems and coalesce with them; but, as a rule, the stems are single.
80 SILICEOUS SPONGES.
The stems vary from 7 to 14 mm. in thickness, and reach to a length of 110 mm.
The branches are from 4 to 9 mm. in thickness.
The canal-structure is mostly obliterated; but in some specimens four or five
longitudinal canals, ‘6 mm. in width, are exposed in the broken ends of a branch. The
spicular mesh appears to be of the same character as in 7’. glabra, but the spicules
are somewhat larger. The dermal layer is only partially preserved; the larger trifid
heads are ‘6 mm. in extension, and thus double the size of those in 7. glabra.
This species is also very common in Chalk flints; the short branches, usually
single stem, generally rough exterior, and the larger size of the spicules of the
dermal layer readily distinguish it from 7. glabra.
Distribution. Upper Chalk: Chicklade, near Hindon; Stockton, near Heytesbury ;
Warminster, Wiltshire ; Dover.
THAMNosPonGIA? RETICULATA, Hinde, n.sp. (Plate XVIII. figs. 3, 3a, 30.)
Sponge consisting of a group of closely arranged, cylindrical or compressed stems,
growing in an upright direction, and frequently coalescing. The summits are conical.
The stems are from 10 to 16 mm. in thickness. An imperfect example is 100 mm.
in height and 80 in width.
Each stem is traversed longitudinally by a single cylindrical canal, 3-5 mm. in
width, into which sinuous radial canals, horizontal or slightly arched, and ‘5 mm. in
width, open. The exterior surface, when the dermal layer is absent, is covered
with a network of reticulated canals. At the summit of the individual stems are
the openings of the vertical canals, which thus appear to have served as cloacal
tubes.
The spicules of the interior have been replaced by calcite, and are too indistinct
for the presence or otherwise of the tubercles to be determined. The arms measure
185 mm. in length by ‘038 mm. in width. Traces of the dermal layer remain ; the
spicules composing it are of the same character as those of 7. glabra.
In the absence of a satisfactory determination of the mesh-spicules, I can only
refer this form provisionally to this genus. Its mode of growth readily characterizes
it from the previous species,
Distribution. Grey Chalk: near Folkestone.
Genus PHOLIDOCLADIA, Hinde, n. g.
Sponges growing in bushy masses, consisting of cylindrical branching stems. No
canal-structure apparent.
The interior skeletal mesh is composed of minute tetracladine spicules, with
strongly tuberculated or annulated arms, and apparently small twig-like extensions
at their ends, which grasp the arms of adjoining spicules without forming distinctive
PHOLIDOCLADIA., 81
nodes at their point of junction. The dermal layer is formed of minute irregular
scale-like spicules, which overlap each other to form a compact surface-covering.
This genus is allied to Thamnospongia in the general characters of the interior
spicular mesh ; but the spicular rays in this latter genus are not so strongly tuber-
culated, and the tubercles are not in a single series as in Pholidocladia. It is
further differentiated from Thamnospongia in the form of the spicules of the dermal
layer. The characters of the dermal layer ally the genus to Plinthosella, but the
interior spicules differ notably in their form, size, and disposition from those of
Plinthosella ; the mode of growth of the sponge itself is also very different.
The only examples of the genus are from the Upper Chalk, preserved in the
interior of flints.
PHOLIDOCLADIA DicHoTomuS, Hinde, n.sp. (Plate XX. figs. 5, 5 a, 5 6.)
Sponge growing in a bush-like form, with somewhat slender branches, from 5:5 to
85 mm. in thickness, which dichotomise at intervals. The branches frequently
coalesce when they come in contact with each other; they are rounded at their
terminations. The only specimen, which is incomplete, forms a mass about 110 mm.
in height and 75 mm. in width.
The spicular mesh of the interior is closely set; the spicular arms are about
*3 mm. in length and ‘07 mm. in width. ‘The dermal layer is only partially preserved ;
the scaly spicules composing it are very irregular in form—some are circular or oval
with smooth or indented margins, whilst others are partially elongated: they are
about ‘5 mm. in width. Where the dermal layer is preserved the surface of the
stems is smooth.
The specimen figured belongs to the Jermyn-Street Museum.
Distribution. Upper Chalk: Wiltshire (Cunnington coll.).
PHoLIDocLADIA RAMOSUS, Hinde, n. sp. (Plate XVIII. figs. 5, 5a, 5 6.)
Sponge growing in bushy masses of small irregularly branching stems, from 2°5 to
3°59 mm. in thickness; branches rounded at their ends. The spicular mesh of the
interior is formed of very nodose spicules, with arms about ‘25 mm. in length and
‘07 mm. in width. No dermal layer has been preserved.
This species generally occurs in the form of small empty branching tubes in the
interior of flints, which would hardly be recognized as casts of lithistid sponges.
The sponge-stem itself is very seldom preserved; but I fortunately found one or two
small fragments which left no doubt that these tubes were occupied by small
branching sponges, which had afterwards been dissolved away. ‘The character of
the interior spicular mesh of this species so closely resembles that of P. dichotoma
that it is highly probable that, similar to that species, it also had a dermal layer of
small scales. There are several flints in the collection with tubes even smaller than
M
82 SILICEOUS SPONGES.
in the one figured; but whether these were originally filled with sponges with yet
slenderer stems than the present form I am unable to determine, for no fragments of
the sponges remain.
Distribution. Upper Chalk: South of England.
Genus RAGADINIA, Zittel, 1878.
The typical and, up to the present, only described species of this genus, Ragadinia
rimosa, Reemer™*, sp., consists of ear- or platter-shaped sponges, with a short lateral
stem and thick walls. There are in the Museum collection several species of sponges
which correspond with the typical form of Zittel in the interior spicular structure,
and also in the spicular structure of the dermal layer and in the characters of the
canals, but differ in their exterior form. These sponges are either cup- or vase-
shaped, or irregularly club-shaped. As these sponges are similar in the important
features of their spicular and canal-structures, I propose to extend Zittel’s definition
of the genus so as to include sponges of the above-mentioned forms.
RaGaDINIA RIMOSA, Remer, sp.
1864. Cupulospongia rimosa, F. A. Roemer, Palzont. Bd. 13, p. 51, t. 17. f. 8.
1878. Ragadinia rimosa, Zittel, Studien, II Ab. p. 88, f. 4a-f; and Handbuch, p. 166, f. 79.
Microscopic slides of the interior spicules and of the spicules of the dermal layer,
from Professor Zittel.
Distribution. Upper Chalk: Ahlten, Hanover.
RaGADINIA COMPRESSA, Hinde, n. sp. (Plate XIX. figs. 5, 3 a.)
Sponges vase- or funnel-shaped, frequently compressed, though some examples
are open and shallow, supported on a short stout stem, which is attached to some
foreign object at its termination. Not infrequently there are nodose outgrowths in
the lower part of the cup. Walls comparatively thick, varying from 10 to 13 mm.
The margins are rounded, and occasionally show fine, transverse, open canals. The
specimens are from 80 to 113 mm. in height, and about the same in breadth at the
summit of the cup.
The upper or interior surface of the cup is furrowed irregularly by minute open
canals; the under surface exhibits also a series of delicate canals with a downward
direction. ‘These canals are only apparent in some specimens: usually the outer
surface has numerous apertures of canals, 1 mm. in width, which open very obliquely
to the surface.
The spicules of the interior mesh are minute and very characteristic in form. One
of the four arms merely consists of a prominent tubercle; the other three arms have
* Paleontographica, Bd. 13, p. 51, t. 17. f. 8.
RAGADINIA. 83
ring-like swellings near their centres, and at their distal ends split up irregularly
into small twigs, by which they are interlocked together, but they do not appear to
form prominent nodes at the point of junction of the arms. The outer surface of
the sponge, where the dermal layer is not present, shows, under a strong lens, the
tubercled spicular arm projecting outwards. The body-spicules of this species are
exactly similar to the detached spicules discovered in the Trimmingham Chalk by
Professor Sollas *, and named by him Compsapsis cretacea, and in the Chalk flint at
Horstead by myself, and named Ragadinia annulata +}. As, however, these remarkable
body-spicules are not peculiar to a single species of sponge, the names given to the
detached spicules cannot be applied to the present or the following forms in which
they occur.
A dermal layer is present in some portions of the outer surface of the specimens,
and probably originally extended over the entire sponge. It is composed of the
horizontally expanded heads of shafted spicules, about ‘3 mm. in width, which are
deeply lobed and laciniated. These heads overlap each other so as to form a con-
nected membrane with a smooth surface. Some of the detached dermal spicules
from the Horstead Chalk flint {, which I referred to Ragadinia annulata, probably
belong to the present species. ‘These dermal spicules also very closely resemble
those of the existing Corallistes asterodiscus, O. Schmidt; but from an examination
of microscopic specimens of this sponge, labelled by Schmidt himself, I have ascer-
tained that its body-spicules are different from those of the present species.
Distribution. Upper Chalk: Oare, Huish, Wiltshire (Cunnington coll.).
Ragapinia suncata, Hinde, n. sp. (Plate XX. figs. 1, 1 a.)
Sponges cup- or funnel-shaped, frequently with only a shallow interior depression.
They gradually taper to a short, straight or oblique, cylindrical stem, with root-like
processes at its termination. Sometimes there are small nodose projections near the
rounded margins of the cup. The walls of the cup vary from 6 to 8 mm. in
thickness. ‘The specimens are from 70 to 90 mm. in height and about 55 mm. in
width at the summit.
Both the outer and inner surface of the sponge is traversed by numerous open
canals, about ‘5 mm. in width, frequently disposed side by side, and running in an
oblique direction, thus giving the surface a furrowed aspect. In addition to the
surface-canals there are others extending into the wall and apparently opening into
the surface-canals.
The interior spicular skeleton is composed of spicules of the same annular cha-
racter, but larger than those of Ragadinia compressa; mingled with these there are
* Annals & Mag. Nat. Hist. 5th ser. vol. vi. 1880, p. 387, pl. 19. figs. 21, 22.
+ Fossil Sponge-Spicules, 1880, p. 58, pl. 5. figs. 1-4.
+ Fossil Sponge-Spicules, pl. 4. figs. 24-30.
M2
84 SILICEOUS SPONGES.
also exposed on the surface elongated irregular filiform spicules. ‘The entire sponge
appears to have possessed a smooth dermal layer of horizontal spicules with laciniated
arms; these heads are about ‘9 mm. in width. I have not been able to discover the
character of the shafts of these spicules.
This species differs from the two preceding, not only in its outer form, but in the
larger size of the body- and dermal spicules.
Distribution. Upper Chalk: Beckhampton, Wiltshire (Cunnington coill.).
Racaprnia cLavaTa, Hinde, n.sp. (Plate XIX. figs. 4, 4a, 4b, 4c.)
Sponges irregularly rod- or club-shaped, often with stumpy processes and nodose
outgrowths. Very variable in size, sometimes reaching to 100 mm. in height and
from 13 to 31 mm. in thickness.
The surface appears to have been covered with a smooth dermal layer; where this
is absent, surface-canals, about °6 mm. in width, are exposed, closely seaming the
exterior of the sponge.
The interior skeleton is composed of distinctly annulated spicules similar to those
of R. compressa. ‘The spicules of the dermal layer have narrow laciniated head-rays ;
they are about ‘5 mm. in extension, and so disposed that there are minute interspaces
between the rays or arms. When the spicules have fallen from the surface of the
sponge, it often happens that the material filling these small interspaces stands out
as sO many minute tubercles.
This species is readily distinguished from others of the genus by its peculiar form.
All the examples have been procured from the interior of flints, and only the spicules
of part of the dermal layer and the spicular structure immediately beneath this layer
have been preserved.
Iistribution. Upper Chalk: Wiltshire (Cunnington coll.).
Genus PLINTHOSELLA, Zittel, 1878.
This genus was founded on small, spherical, or irregular nodose sponges, from
5 mm. to 25 mm. in thickness, with an interior skeletal mesh of large irregularly
four-armed tuberculated spicules, and a dermal layer composed of small scaly disks
of irregular form. In the Museum collection and in the Jermyn-Street Museum
there are cup-shaped, rod- or club-like, and convoluted sponges with an interior
skeletal mesh closely resembling that of Plinthosella squamosa, Zitt., the type of the
genus. The spicules of the dermal layer of these sponges, however, vary in detail
from those of the type species, for they are usually disk-like with jagged edges, or
even occasionally lobate. ‘This variation is not sufficient, however, to be regarded as
a basis for generic distinction, for in a single specimen of the typical species there
may be found great diversity in the form of the disks of the dermal layer.
PLINTHOSELLA. 85
Relying on the similarity in the spicular structure of the interior mesh, and on
the general resemblance of the disks of the dermal layer of these cup-shaped, clavate,
and convolute forms to the same structures in the spherical P. sgwamosa, I shall
place them under this genus, and propose to extend the definition given by Zittel so
as to include sponges of these forms. The new forms, like the type species, have
been derived from the Upper Chalk.
PLINTHOSELLA squamosa, Zittel. (Plate XX. fig. 2.)
1878. Plinthosella squamosa, Zittel, Studien, Il Ab. p. 89, t. 2. f. 10 and t. 10. f. 5; also
Handbuch, p. 167, f. 80.
1880. Plinthosella squamosa, Hinde, Foss. Sponge-Spicules, p. 56, t. 4. f. 35-46.
Small spherical bodies, from 6 mm. to 23 mm. in diameter, apparently destitute of
any point of attachment.
In some examples there appear to have been a few canals immediately beneath
the dermal layer, but the circulation in the interior of the sponge seems to have been
carried on in the interspaces of the mesh.
The spicular structure of the interior is composed of relatively large, irregularly
four-armed spicules with tuberculated surfaces and minute twig-like extremities, by
which they are interlocked together. There is a dermal layer of small, irregularly
shaped disks or lamine which are arranged so as to overlap each other.
The examples of this sponge occur in this country as small globular bodies, from
the size of peas to that of marbles, inclosed in the interior of Chalk flints. In all
the specimens which I have seen the dermal layer is absent, and only the empty
cavities of the spicules of the interior labyrinthine mesh remain. When closely
examined with a strong lens, some of the moulds, still retaining the impressions of
the tubercles on the surface of the spicules, can be seen. In the Horstead Chalk I
discovered detached spicules both of the interior and of the dermal layer of this
species.
Distribution. Upper Chalk: South of England; Wiltshire (Mr. HZ. J. Carter) ;
Horstead, Norfolk (detached spicules); Ahlten, Hanover (Zittel’s coll.).
PLINTHOSELLA compacta, Hinde, n. sp. (Plate XX. figs. 3, 3a.)
Body of sponge cup-shaped, supported on a cylindrical stem. ‘The interior is
nearly filled with the spicular mesh, so that there is only a shallow depression at the
summit of the cup. ‘The only specimen is 50 mm. in height, and 72 mm. in thickness
near the summit.
The rounded margins of the cup are seamed with sinuous transverse canals; and
the outer surface is covered with apertures, 1 mm. wide, of canals which extend into
the sponge-wall. A few canals also extend into the stem.
86 SILICEOUS SPONGES.
The spicular structure of the interior closely resembles that of P. sguamosa. No
dermal layer has been preserved.
I have only seen one example of this species, which is from the interior of a flint.
Notwithstanding the absence of a dermal layer, I have but little doubt of the
propriety of placing it in this genus on account of the close resemblance of the
interior-mesh spicules to those of the type species. The specimen in question
belongs to the Jermyn-Street Museum.
Distribution. Upper Chalk: Wiltshire (Cunnington coll.).
PLINTHOSELLA Noposa, Hinde, n.sp. (Plate XX. figs. 4, 4 a.)
Sponges simple, rod- or club-shaped, with irregular nodose and spur-like projections.
A fairly large specimen is 100 mm. in height and 80 mm. wide.
The surface of the sponge, where the dermal layer is not present, is traversed by
sinuous canals from ‘5 to 1 mm. in width. The interior structure is obliterated, so
that I am unable to determine if interior canals are present.
The spicules of the interior mesh resemble those of P. sguamosa, but they are
somewhat smaller. The surface is covered with a dermal layer of disciform spicules
(about °6 mm. in width), very variable in figure, some being deeply lobed like those
of Ragadinia, whilst others are circular disks with merely jagged edges. These
spicules overlap each other in such a manner as to produce a compact, smooth
surface-layer.
In some respects this species exhibits characters similar to those of Ragadinia; and
without an examination of the spicular structure, it might readily be confounded with
R. clavata. As the spicules of the interior skeleton are of the same type as Plintho-
sella squamosa, it appears to me to belong rather to this genus than to Ragadinia.
The variable character of the spicules of the dermal layer is well exhibited in this
species, for in close proximity to each other on the surface there are nearly plain
disks and deeply-lobed branching spicules.
Distribution. Upper Chalk: Wiltshire (Cunnington coll.).
PLINTHOSELLA ConvoLUTA, finde, n. sp.
Sponge growing as an irregularly convolute plate from 5 to 5°5 mm. in thickness.
The only specimen, which is incomplete, is 110 mm. in height and 82 mm. in lateral
extension.
It is doubtful whether canals are present in the sponge-wall. The skeletal mesh
is composed of spicules closely resembling in size and form those of Plinthosella
compacta, The dermal layer is absent; but in one small portion of the outer
surface there are a few minute scaly spicules which may represent part of the former
external layer.
This species very closely resembles, in its mode of growth, Phymaplectia irregularis ;
PHYMAPLECTIA. 87
and in the absence of the dermal layer, the main point of distinction between these
forms consists in the much larger size of the interior spicules of the present species ;
when the dermal layer is present, the difference between the scaly spicules and the
trifids of P. crregularis at once distinguishes these forms from each other.
Distribution. Upper Chalk: Wiltshire.
Genus PHYMAPLECTIA, Hinde, n. g.
Sponges with plate-like walls, palmate or digitate, or, by the coalescing of the
margins of the convolute walls, becoming cup- or funnel-shaped. Canal-system but
slightly developed, the circulation apparently being carried on through the openings
of the spicular mesh. The interior skeleton is composed of irregular tuberculated
spicules of the same type as in Plinthosella and Spongodiscus, which are connected
together by the interlocking of minute twig-like extensions at the ends of the arms,
or by the apposition of adjoining spicules. A dermal layer is usually present, and is
chiefly composed of trifid spicules with minute, horizontally expanded, and slightly
bifurcated head-rays.
The character of the dermal layer distinguishes this genus from Plinthosella; in
form and mode of growth it differs from Spongodiscus, Zitt. At present no dermal
layer has been discovered in sponges of this latter genus, although there is hardly a
doubt that one was present originally.
The forms of the genus at present known are all from the Upper Chalk.
PHYMAPLECTIA IRREGULARIS, Hinde, n. sp. (Plate XXI. figs. 1, 1a.)
Sponges growing in upright fan-like, semipalmate, or digitiform expansions, or
occasionally, by the involution of the wall-plate and the coalescence of the margins,
becoming cup-shaped. ‘The margins of the wall are rounded ; at the base there are
traces of root-like extensions. The specimens are from 100 to 120 mm. in height ;
the walls are between 5 and 7 mm. in thickness.
No canal-system is apparent, but the condition of the specimens is such that it is
impossible to determine whether canals were originally present or not. The interior
spicular mesh is composed of irregular warty spicules, the arms of which are from
‘2 to 3mm. in length. A smooth dermal layer apparently covered the entire outer
surface of the sponge-wall, though only portions of it now remain. It is made up of
trifid spicules with horizontal heads; the rays are slightly compressed and regularly
bifurcated. ‘The heads of the larger spicules are between *4 and ‘5 mm. in expansion.
The character of the shafts is not shown.
All the examples of this species are preserved in the interior of flints. In outer
form they somewhat resemble the figures of Polypothecia palmata and P. fissa,
Benett*; but in the absence of all knowledge of the minute structure of these
* Cat. Org. Rem. County of Wilts, pl. 11. f. 2, and pl. 12.
88 SILICEOUS SPONGES.
sponges, I am unable to determine whether the resemblance is more than in outer
form.
Distribution. Upper Chalk: Wiltshire.
PuyMaPLectia spinosa, Hinde, n. sp. (Plate X XI. figs. 2, 2 a.)
Sponges with convolute walls, from 5 to 7 mm. in thickness, which coalesce at
their margins and become funnel-shaped. The margins are rounded; the basal
portion has not been preserved. The exterior surface is furnished in places with
small, straight spinous processes. A specimen is 92 mm. in height, and the same in
width at the summit.
Traces of sinuous canals, ‘5 mm. in width, are apparent in one part of a specimen,
but I cannot determine if these are generally present. The interior spicular mesh is
of the same character as in the preceding species, but the spicules are somewhat
more robust. The outer surface retains in places the smooth dermal layer; the
spicular heads of which it is formed are about ‘25 mm. in extension. The inner
surface of the cup is not exposed.
This species approaches very closely to P. irregularis, but the spicules of its interior
mesh are somewhat larger, and those of the dermal layer smaller, than in the last-
named species. ‘The specimen figured belongs to the Jermyn-Street Museum.
Distribution. Upper Chalk: Wiltshire (Cannington coll.).
PHYMAPLECTIA CRIBRATA, Hinde, n.sp. (Plate XXI. figs. 3, 3a.)
Wall-plate of sponge growing in wavy folds, showing also slight concentric,
rounded ridges on the outer surface. The wall is from 4 to 5 mm. in thickness.
The basal portion of the specimen is not preserved. The imperfect example is
82 mm. in height and 112 mm. in width.
The spicules of the interior mesh are very closely tuberculated; the arms are
about ‘4 mm. in length, and more robust than in the preceding species. The outer
surface of the sponge-wall is covered with a smooth dermal layer, which is penetrated
with numerous minute pores about 1 mm. in diameter, and only visible under a good
lens. The dermal layer between these pores is in part composed of extremely minute
trifid spicules, the head-rays of which are about ‘1 mm. in extension, in part of
apparently irregular spicular bodies, whose forms are not sufficiently well-preserved
for determination.
The mode of growth, and more particularly the characters of the dermal layer,
readily distinguish this species from the preceding, The only specimen known is
from the interior of a flint. It belongs to the Jermyn-Street Museum.
Distribution. Upper Chalk: Wiltshire (Cunnington coll.).
PHYMAPLECTIA.—RHOPALOSPONGIA. 89
PHYMAPLECTIA ScITULA, Hinde, n. sp. (Plate XXII. figs. 1, la, 10.)
Sponge funnel-shaped, open below, apparently formed by the coalesced margins of a
convolute plate. The walls thin and delicate, between 2°8 and 3:5 mm. in thickness.
The only specimen is 38 mm. in height, and 74 mm. in its greatest width.
The interior of the funnel is marked by very slight elevated spots about 5 mm.
apart, towards which closely set minute, radial, open canals, about -4 mm. in width,
concentrate. No distinct aperture is to be seen at the centre of these elevations; the
radial canals round some of them are but very faintly marked.
The spicular mesh is formed of relatively large, closely tuberculated spicules with
arms about ‘35 mm. in length. Very minute spicular bodies also appear to be present
between the mesh-pores of the larger spicules, but they are too indefinite for deter-
mination. No dermal layer has been preserved.
The thin delicate walls and the surface-canals of the interior characterize this
species. I have only seen a single example, preserved in flint, which belongs to the
Jermyn-Street Museum.
Distribution. Upper Chalk: Wiltshire (Cannington coll.).
Genus RHOPALOSPONGIA, Hinde, n. g.
Sponges simple or aggregate, club-shaped, with rounded or flattened summits;
stem simple, with, in some cases, root-like prolongations.
Neither cloaca nor prominent canals are present ; the sponge is traversed by curved
canals of moderate dimensions, which extend from the central portions and open at
the surface. Longitudinal canals are present in the stem.
The spicular tissue is composed, for the most part, of relatively large tuberculated
spicules, with straight or curved arms, which connect with adjoining spicules by the
interlocking of their tuberculated extremities. In these spicules the tetracladine
character can hardly be recognized, but mingled with them in the mesh there are
regularly four-armed spicules with smooth arms and tuberculated extensions. No
dermal layer has been preserved.
The mode of growth, character of the canal-system, and the ere spicular
structure readily distinguish this genus. In the tuberculate character of the
spicules, it resembles Plinthosella, Spongodiscus, and Phymaplectia; but, so far as
I am aware, in none of these genera are smooth-armed spicules mingled with
tuberculate forms.
At present I have recognized twe species belonging to this genus, both of which
are from the Upper Green Sand.
90 SILICEOUS SPONGES.
RHOPALOSPONGIA GREGARIA, Benett, sp. (Plate XXII. figs. 2, 2a, 2b, 2c.)
1831. Polypothecia gregaria, Benett, Cat. Org. Rem. County of Wilts, t. 14.
1847. Non Jerea gregaria, Michelin, Icon. Zooph. p. 134, t. 38. f. 1.
Sponges elongate, club-shaped, with rounded conical summits, either simple or
growing in groups of three or four individuals of different sizes, closely attached by
their lateral surfaces. Rarely, also, two individuals possess a common stem. The
outer surface of the body, or club-shaped portion, is either smooth and evenly
rounded, or with irregular nodose projections, or covered with small pustular
elevations. The cylindrical stem frequently exhibits parallel longitudinal furrows ;
it appears to have given off root-like processes at its termination. Small individuals
measure 55 mm. in height and 14 mm. in width; whilst a large example is 300 mm.
in length by 45 mm. in width.
The outer surface of the sponge is thickly covered with canal-apertures from 5 to
‘75 mm. in width. These canals apparently extend in an arched direction towards
the centre of the sponge; in a transverse horizontal section only a few of the larger
canals are distinguishable. Both in vertical and horizontal sections distinct lines of
growth parallel with the contour of the sponge are exposed.
The spicular mesh is relatively close; the smooth arms of the regular spicules are
‘06 mm. in width, whilst the tuberculated arms measure ‘(08 mm. wide. The regular
spicules are indiscriminately mingled with the others; they appear, however, to be
more frequent in the central portion of the sponge, whilst the irregular tuberculated
spicules prevail nearer the outer surface. Interior canals are preserved in many of
the spicules. This species is not uncommon in the Upper Green Sand.
There can be no doubt that Miss Benett’s figure represents this species, particularly
as there is no other sponge from the Green Sand with a corresponding form. Michelin
has referred this species to his Jerea gregaria, with which, however, it has no near
affinity.
Distribution. Upper Green Sand: Warminster and vale of Pewsey, Wiltshire.
RHOPALOSPONGIA OBLIQUA, Hinde, n. sp. (Plate XXII. figs. 3, 3a.)
Sponge compound, consisting of two short individuals, inversely conical in form,
growing from a single short, somewhat compressed stem. The summits are appa-
rently oblique.
A transverse section exposes the canals, 1 mm.in width, which extend to the outer
surface. ‘These canals are more numerous and distinct than those of the last species.
The spicular structure of the interior resembles that in R. gregaria.
I have only met with a single example of this species: it is 8) mm. in height and
51 mm. in width.
Distribution. Upper Green Sand: Warminster, Wilts.
ASTYLOSPONGIA. 91
Order HEX ACTINELLID &, Oscar Schmidt.
Suborder DICTYONINA, Zitéel.
Family ASTYLOSPONGIDA.
Genus ASTYLOSPONGIA, Ferd. Remer, 1860.
Ferd. Reemer described the minute structure of A. premorsa, the typical species
of the genus, as consisting of very regular six-rayed, star-shaped bodies, which are
connected together by the intermediate union of the rays of each body with the rays
of those immediately adjoining (‘Die silur. Faun. des west. Tenn. p. 8). In a
later definition of the genus (‘Lethea Palozoica,’ 1 Th. p. 307) the same author
states that the skeleton consists of a net-like web, composed of nodes, from which
six rays extend at right angles with each other, and unite with the rays of the
adjoining nodes. Prof. Zittel (‘ Studien,’ 1 Th. p. 45) states, in his diagnosis of the
genus, that the skeleton consists of amalgamated six-rayed spicules with solid nodes,
forming an irregular net-work with triangular, quadrangular, or polyangular meshes,
and that, asa rule, the arms of several adjoining spicules attach themselves to a
single node. In 1878 Dr. K. Martin published a very important memoir on the
structure of Astylospongia (‘Archiv des Vereins der Freunde der Naturgeschichte in
Mecklenburg’), in which he states that the number of the rays which radiate from a
single node vary from six to nine in number, and that they radiate in most variable
directions in order to unite with other rays to form a new node. He also made the
interesting discovery that the terminations of the rays divide into minute branches
(loc. cit. Taf. 1. f. 3). These facts led Dr. Martin to express the opinion that there
were wide differences between Astylospongia and typical Hexactinellids. In a short
comment on Dr. Martin’s memoir, Prof. Zittel (‘Neues Jahrbuch,’ 1877, p. 709)
states that the branching of the termination of the rays is entirely the result of the
state of preservation of the specimen, and that in unaltered examples the nodes are
always solid.
An examination of the microscopic structure of Astylospongia premorsa from
Tennessee leads me to confirm the observations of Martin that the spicules of this
species are composed of solid nodes with from six to nine smooth arms which radiate
from the centre in different directions and at varying angles. ‘The terminations of the
arms or rays of adjoining nodes unite together, and appear to form nodes by their union,
which, when the specimen is unaltered, cannot be distinguished from the nodes from
which the arms themselves originated ; but in specimens where a certain amount of
alteration has taken place, these secondary nodes can be seen to be composed of the
expanded terminations of the arms of several spicules united together. If this view
N2
92 SILICEOUS SPONGES.
is correct, Astylospongia premorsa can hardly be regarded as a hexactinellid sponge,
but shows a near relationship in its minute structure to the Lithistid genera Cylin-
drophyma and Melonella.
As, however, further observations are necessary to establish the true character of
the spicular structure of Astylospongia, I prefer in the meantime not to remove the
genus from the position assigned to it by Prof. Zittel.
ASTYLOSPONGIA PREMORSA, Goldfuss, sp.
1833. Siphonia premorsa, Goldf. Petref. 1 Th. p. 17, t. 6. f. 9.
1833. Siphonia excavata, Goldf. Petref. 1 Th. p. 17, t. 6. £. 8.
1860. Astylospongia premorsa, F. Remer, Silur. Fauna d. west. Tenn. p. 8, t. 1. f. 1.
1877. Astylospongia premorsa, Zitt. Studien, I Ab. p. 45; id. Neues Jahrbuch, p. 353, t. 2.
f. 1; id. Handbuch der Pal. p. 172, f. 88.
1878. Astylospongia premorsa, Martin, Archiv des Vereins d. F.d. N. Mecklenburg, p. 1, t. 1.
1878. Siphonia premorsa, Quenst. Petref. Bd. 5, p. 551, t. 141. f. 1-3.
Distribution. Silurian (Niagara group): Waldron, Indiana; Bath Springs, Perry
County, Tennessee.
ASTYLOSPONGIA STELLATIM-SULCATA, Roemer.
1848. Spongia stellatim-sulcata, F. Roemer, Leonh. u. Bronn’s Jahrbuch, p. 686, t. 9. f. 5.
1860. Astylospongia stellatim-sulcata, F. Roemer, Silur. Fauna d. west. Tenn. p. 11, t. 1.
£.2; 24. 25.
Distribution. Silurian (Niagara group): West Tennessee.
ASTYLOSPONGIA INCISO-LOBATA, Remer.
1848. Spongia inciso-lobata, F. Roemer, Leonh. u. Bronn’s Jahrb. p. 685.
1860. Astylospongia inciso-lobata, F. Reemer, Silur. Fauna d. west. Tenn. p. 11, t.1.£.3, 3a.
Distribution. Silurian (Niagara group): Bath Springs, Perry County, Tennessee.
ASTYLOSPONGIA IMBRICATO-ARTICULATA, Roemer.
1848. Stphonia imbricato-articulata, Roemer, Leonh. u. Bronn’s Jahrb. p. 685, t. 9. f. 3.
1860. Astylospongia imbricato-articulata, Roemer, Silur. Fauna d. westl. Tenn. p. 12, t. 1.
1 5) GIA
Distribution. Silurian (Niagara group): West Tennessee.
AstyLosponGia? Ramer, Hinde, n. sp. (Plate XXIII. figs. 1, 1 a, 10.)
A fragmentary specimen, having the spicular structure of the genus, but the
portion preserved is insufficient to determine the complete form and canal-structure.
The fragment is of a conical form, with open sinuous channels 1-75 mm. wide
PALZOMANON.—TREMADICTYON. 93
extending downwards from the summit of the cone. Circular canal-apertures,
1:25 mm. wide, are closely arranged in linear series in the channels, and also on
the summit. The canals appear to extend in a nearly straight direction to the
central portion of the sponge. The spicular nodes are about ‘16 mm. in thickness,
the arms about -15 mm. in length and ‘041 mm. in thickness. The minute expanded
terminations at the ends of the arms are very clearly shown in microscopic sections
of this form.
Nothing is known of the geological horizon or locality whence the specimen comes,
but the condition of preservation is precisely similar to that of specimens of Astylo-
spongia from the Silurian of North America.
Genus PALZOMANON, Remer, 1860.
PALEZOMANON CRATERA, Remer.
1848. Siphonia cratera, Roemer, Leonh. u. Bronn’s Jahrb. p. 685, t. 9. £. 4, 4a.
1860. Paleomanon cratera, Roemer, Silur. Faun. d. west. Tenn. p. 13, t. 1. f. 4, 4a.
1877. Paleomanon cratera, Zitt. Studien, I Ab. p. 45.
Distribution. Silurian (Niagara group): West Tennessee.
Family EURETID&, Zittel.
Genus TREMADICTYON, Zittel, 1877.
TREMADICTYON RETICULATUM, Goldfuss, sp.
1833. Scyphia reticulata, Goldf. Petref. 1 Th. p. 11, t. 4. f. 1. ‘
1833. Scyphia polyommata, Goldf. Petref. 1 Th. p. 8, t. 2. f. 16.
1833. Scyphia fenestrata, Goldf. Petref. 1 Th. p.7, t. 2. f. 15.
1833. Scyphia pertusa, Goldf. Petref. 1 Th. p. 6, t. 2. f. 8.
1877. Tremadictyon reticulatum, Zitt. Studien, I Ab. p. 46; Neues Jahrbuch, p. 355, t. 2.
f. 2, also p. 706; Handbuch d. Pal. p. 173, f. 89.
1878. Spongites reticulatus, Quenst. Petref. Bd. 5, p. 28, t. 115. f. 1-12, 14-23.
Distribution. Upper Jura, Spongitenkalk: Heuberg, Randen, Nattheim, Streitberg.
TREMADICTYON OBLIQUATUM, Quenst. sp.
1858. Spongites obliquatus, Quenst. Der Jura, p. 671, t. 81. f. 97.
1877. Tremadictyon obliquatum, Zitt. Studien, I Ab. p. 46.
Distribution. Upper Jura: Heuberg, Randen, Wiirtemberg.
94 SILICEOUS SPONGES.
Genus CRATICULARIA, Zittel, 1877.
CRATICULARIA CLATHRATA, Goldf. sp.
1833. Scyphia clathrata, Goldf. Petref. 1 Th. p. 8, t. 3. f. 1.
1877. Craticularia clathrata, Zitt. Studien, I Ab. p. 46; Neues Jahrbuch, p. 355.
1878. Spongites clathrata, Quenst. Petref. Bd. 5, p. 74, t. 117. f. 23, 25.
Distribution. Upper Jura: Heuberg, Randen, Wiirtemberg.
CRATICULARIA PARALLELA, (Groldf. sp.
1833. Scyphia parallela, Goldf. Petref. 1 Th. p. 8, t. 3. f. 3.
1877. Craticularia parallela, Zitt. Studien, I Ab. p. 46; Neues Jahrbuch, p. 355.
1878. Spongites cylindritextus, Quenst. Petref. Bd. 5, p. 65, t. 117. f. 9-15.
Distribution. Upper Jura: Heuberg, Wodna, Cracow (Zittel’s coll.).
CRATICULARIA DECORATA, Miinster, sp.
1833. Scyphia decorata, Minster in Goldf. Petref. 1 Th. p. 90, t. 33. f. 2a, b.
Distribution. Upper Jura: Randen.
CRATICULARIA PARADOXA, Miinster, sp.
1833. Scyphia paradoxa, Miinster in Goldf. Petref. 1 Th. p. 86, t. 31. f. 6.
1877. Craticularia paradoxa, Zitt. Studien, I Ab. p. 46; Neues Jahrbuch, p. 356; Hand-
buch d. Pal. p. 174, f. 90.
1878. Clathrispongia trochiformis, ventricosa, et perlata, Quenst. Petref. Bd. 5, pp. 75, 76,
80, t. 118. f. 2, 3, 6.
Distribution. Upper Jura: Streitberg (Z7tt. coll.), Randen, Wirtemberg.
CraticuLaria Firroni, Mantell, sp. (Plate XXIII. figs. 2, 2a, 2d.)
1833. Millepora Fittoni, Mantell, Geology S. E. of England, p. 378.
1822. Millepora , Mantell, Geol. of Sussex, p. 106, t. 15. f. 10.
1848. Brachiolites digitatus, Toulmin Smith, Ann. Mag. Nat. Hist. vol. 1. pp. 45, 365,
t. 16. f. 2.
1854. Brachiolites Fittoni, Morris, Cat. Brit. Foss. p. 26.
1864. Dendrospongia fenestralis, F. A. Roemer, Palont. Bd. 13, p. 21, t. 8. f. 6.
Sponges growing in masses of straight or slightly sinuous cylindrical branches,
from 22 mm. to 35 mm. in diameter and in some cases more than 220 mm. in
length. The branches are hollow tubes with walls from 2°5 mm. to 4 mm. in thick-
ness; the summits are open. ‘The canal-apertures, both on the outer and inner
wall-surfaces, are disposed in vertical and horizontal rows; the apertures are sub-
quadrate at the surface, and from 1 mm. to 1°5 mm. in width.
In most of the examples the spicular structure has been completely replaced by
CRATICULARIA.—SPHENAULAX. 95
calcite, and is therefore indistinct, but in one specimen from the Isle of Wight the
siliceous skeleton is partly retained. The interior mesh of the wall is more irregular
than in the Jurassic forms of this genus; the spicular arms or rays are about ‘05 mm.
in thickness, and the length from centre to centre of the nodes is‘25 mm. The
dermal layer of the outer surface of the wall has a stellate appearance ; the nodes of
the spicules are flattened, and numerous rays extend from each ; besides the inter-
spaces between the spicular arms, there are also minute circular apertures in the
dermal layer.
This species is abundant and occurs in all the divisions of the Chalk series, from
the Upper Green Sand to the Upper Chalk. Toulmin Smith describes this species
as “having a deep primary fold of regular quadrilateral and rectangular form,
usually more or less oblong, and arranged in tesselated figure.” I cannot ascertain
that the walls are at all folded, but merely penetrated with numerous canals on both
sides, which do not extend quite through the wall but terminate bluntly. T. Smith
does not mention the difference between the solid nodes of this species and the
lantern-nodes of Ventriculites.
Distribution. Craie Chloritée: Honfleur, Vaches Noires (Zesson coll.). Chalk
Marl: Ventnor, Isle of Wight. Grey Chalk and Lower Chalk: Dover, Folkestone.
Upper Chalk: South of England (Toulmin Smith and Mantell’s coll.).
CRATICULARIA SUBSERIATA, Rte@mer, sp.
1840. Seyphia subseriata, F. A. Roemer, Verst. d. Nordd. Kreide, p. 9, t. 3. f. 8.
Sponge consisting of bifurcating cylindrical branches, 15 to 22 mm. in thickness ;
the walls are 2 to 2°5 mm. thick. The canal-apertures are disposed in vertical and
transverse rows; they are ‘75 mm. wide, and nearly the same distance apart from
each other. Spicular mesh of the interior irregular in disposition. A single frag-
mentary specimen is in the collection, which appears to agree with Roemer’s species.
From Craticularia Fittoni it is distinguished by the smaller dimensions of the
branches and canals. .
Distribution. Upper Chalk: Norwich (Bayfield coll.).
Genus SPHENAULAX, Zittel, 1877.
SPHENAULAX cosTaTa, Goldf. sp.
1833. Scyphia costata, Goldf. Petref. 1 Th. p. 6, t. 2. f. 10.
1877. Sphenaulax costata, Zitt. Studien, I Ab. p. 47; Neues Jahrbuch, p. 356, t. 2. f. 3,
and p. 709.
1878. Sulcispongia incisa, colliciaris, rimosa, viaria, semiclathrata, Quenst. Petref. Bd. 5,
p- 83-90, t. 118. f. 8-16.
Distribution. Upper Jura: Engelshardt, Heuberg.
96 SILICEOUS SPONGES.
Genus SPORADOPYLE, Zttel, 1877.
SPORADOPYLE OBLIQUA, Goldf. sp.
1833. Scyphia obliqua, Goldf. Petref. 1 Th. p. 9, t. 3. f. 5a, b, d.
1877. Sporadopyle obliqua, Zitt. Studien, I Ab. p. 47; Neues Jahrb. p. 356, t. 2. f. 6.
1878. Favispongia obliqua, Quenst. Petref. Bd. 5, p. 118, t. 120. f. 29-53.
Distribution. Upper Jura: Randen, Streitberg, Bavaria.
SPORADOPYLE TEXTURATA, Goldf. sp.
1833. Scyphia texturata, Goldf. Petref. 1 Th. p. 6, t. 2. f. 9.
1833. Scyphia pertusa, Goldf. Petref. 1 Th. p. 6, t. 2. f. 8.
1877. Sporadopyle texturata, Zitt. Studien, I Ab. p. 47; Neues Jahrbuch, p. 356.
1878. Scyphia pertusa, Quenst. Petref. Bd. 5, pp. 126-129, t. 120. f. 61-69.
Distribution. Upper Jura: Randen, Wiirtemberg.
SPORADOPYLE RAMOSA, Quenst. sp.
1858. Spongites ramosus, Quenst. Der Jura, p. 683, t. 83. f. 1.
1877. Sporadopyle ramosa, Zitt. Studien, I Ab. p. 47; Neues Jahrb. p. 356.
1878. Ramispongia ramosa, Quenst. Petref. p. 140, t. 121. f. 11.
Distribution. Upper Jura: Heuberg, Randen.
SPORADOPYLE, sp.
Distribution. Upper Jura: Heuberg, Wodna, Cracow (Zittel’s coll.).
Genus STREPHINIA, Hinde, n. g.
Sponges growing in irregular, convolute, anastomosing expansions or open cup-
shaped. The wall on both sides is irregularly reticulate with circular, oval, or
irregular canal apertures of relatively large size. The canals terminate blindly. The
spicular mesh is small and somewhat irregular, the spicular arms apparently smooth ;
the nodes are solid. No special dermal layer is present.
In the mode of growth this genus differs from any other of this family. The
characters of the spicular mesh resemble those of Sphenaulax and Sporadopyle.
The only examples are from the Grey Chalk, and their skeletons have been com-
pletely replaced by crystalline calcite.
SrrEPHINIA convoLuUTA, Hinde, n. sp. (Plate XXIII. figs. 3, 3a, 36.)
The walls of the sponge are convolute and anastomose so as to form irregular
masses. ‘The wall is 4 mm. in thickness; the outer surface is reticulate, with closely
set, circular or oval canal-apertures, from 1 to 1-5 mm. in width, and about 1°3 mm.
STREPHINIA.—VERRUCOCGLIA. 97
apart; on the inner surface of the wall the canal-apertures, in some instances at
least, have a vertically linear disposition, and are somewhat larger than those of the
outer surface.
The mesh-spaces are quadrate; the distance from one nodal centre to another is
‘16 mm.; the spicular arms are apparently even, and ‘052 mm. in thickness.
There is only a single fragmentary specimen in the collection, which distinctly
shows the canal and spicular structures of this species; but from the same bed near
Dover there are large spreading masses of a sponge which has a similar mode of
growth, and probably belongs to this species, but the structural characters are
obliterated, so that it is impracticable to determine with certainty.
Distribution. Grey Chalk: Dover.
STREPHINIA RETEFORMIS, Hinde, n. sp. (Plate XXIII. figs. 4, 4a.)
Sponges either cup-shaped or growing in curved expansions. The wall is from
5-5 mm. to 7 mm. in thickness; the outer surface is reticulate, with oval or sub-
polygonal canal-apertures, from 2 to 3 mm. in width. The walls between these
apertures are from ‘75 to 1 mm. in width. The interior surface is not exposed.
The spicular characters resemble those of the previous species.
This species differs from S. convoluta by its thicker walls and the more open
characters of the reticulations of the outer surface.
Distribution. Grey Chalk: Dover.
Genus VERRUCOCCELIA, Ltallon, 1860.
VERRUCOCHLIA VERRUCOSA, Goldf. sp.
1833. Scyphia verrucosa, Goldf. Petref. 1 Th. p. 91, t. 33. f. 8, a, 8, ¢, e.
1820. Spongus botryoides ?, Konig, Icones fossiles, p. 4, t. 7. f. 82.
1858. Scyphia verrucosa, Quenst. Der Jura, p. 668, t. 81. f. 86.
1877. Verrucocelia verrucosa, Zittel, Studien, I Ab. p. 47; Neues Jahrbuch, p. 356.
1878. Mastospongia verrucosa, Quenst. Petref. Bd. 5, p. 146, t. 122. f. 3-6.
Distribution. Upper Jura: Randen; Heuberg, Streitberg.
VERRUCOC@LIA GREGARIA, Quenst. sp.
1858. Scyphia gregaria, Quenst. Der Jura, p. 668, t. 81. f. 85.
1877. Verrucocelia gregaria, Zittel, Studien, I Ab. p. 47 ; Neues Jahrbuch, p. 356, t. 2. £5.
1878. Mastospongia gregaria, Quenst. Petref. Bd. 5, p. 148, t. 122. f. 8-10.
Distribution. Upper Jura: Randen.
VERRUCOC@LIA TUBULATA, Toulmin Smith, sp.
1848. Brachiolites tubulatus, T. Smith, Aun. & Mag. Nat. Hist. vol. i. p. 366, t. 15. f. 7.
Sponges growing in the form of a hollow cone, which is studded on all sides by
fn)
98 SILICEOUS SPONGES.
small, upward projecting, cylindrical or compressed tubes, which are open above.
Neither the summit nor basal portion is preserved in the specimens. The example
figured by Smith is 61 mm. in height and 50 mm. in width. Another imperfect
specimen in the Museum is 86 mm. in height and 61 mm. in width. The projecting
tubes are from 5 mm. to 7°6 mm. in diameter, and in length from mere projecting
cups to tubes 17-6 mm. in length. The thickness of the spicular wall is about
12mm. The outer surface of the wall has a delicate lace-like dermal layer com-
posed of somewhat thickened spicular arms and nodes. There are minute circular
apertures, about ‘5 mm. in width, in this layer as well as the small interspaces in the
mesh itself. The mesh-spicules are small, with solid nodes. I have not been able
to determine the characters of the inner surface of the wall.
From the typical Jurassic forms of Verrucocelia this species differs in the possession
of a modified dermal layer; but I do not regard this feature as of sufficient importance
to place this form in a new genus, but propose to extend Zittel’s definition so as to
include sponges with a perforate dermal layer. The spicules of the interior mesh of
the wall, and the general mode of growth, resemble the typical Jurassic species.
The specimens are now in the condition of iron peroxide. The figure of this
species given by T. Smith represents it in a reverse position. The woodcut (fig. Q)
on p. 367, in the ‘ Annals,’ which, according to Smith, represents a transverse section
of this species near the top, does not appear to belong to this form at all.
Distribution. Upper Chalk: South of England.
VERRUCOCELIA VECTENSIS, Hinde, n. sp. (Plate XXIV. figs. 3, 3a, 3.)
Sponge generally conical in form, widest a short distance above the base, and thence
tapering to the summit. The only specimen is 152 mm. in height and 100 mm. in
greatest width. Numerous cylindrical tubes, from 20 to 35 mm. in length and from
11 to 20 mm. in width, spring from the central cavity and project upwards. These
tubes are open at the surface; the margins are rounded. The walls are from 2 to
2-5 mm. in thickness, and the interior of the tubes from 5 to 13 mm. in width. The
spicular structure of the outer surface-layer is slightly different from that of the
interior of the wall in that the arms and nodes of the spicules are somewhat thicker.
This reticulate dermal layer is perforated by numerous circular or ovate canal-
apertures, about °5 mm. in width. The spicular mesh of the interior of the wall is
somewhat irregular, and is composed of robust spicules with compact (?. e. not
lantern) nodes. The spicular arms are in places minutely spinous, in other places
they appear to be smooth; they vary from ‘045 to ‘067 mm. in thickness, and the
distance from one nodal centre to another is about °225 mm. There are also occa-
sionally present in the interstices of the mesh very minute delicate hexactinellid
spicules, which are only attached by the extremity of a single arm to the mesh
itself.
STAURONEMA. 99
The large dimensions of this species, and the size of the tubes and walls, readily
distinguish it from all others of the genus. The only specimen is in Chalk Marl,
and the spicular structure comes out very perfectly when the matrix is removed
by acid.
Iistribution. Chalk Marl: Ventnor, Isle of Wight (Mantell’s coll.).
Genus STAURONEMA, Sollas, 1877.
STAURONEMA Cartert, Sollas. (Plate XXIV. figs. 1, la, 16.)
1877. Stawronema Carteri, Sollas, Ann. & Mag. Nat. Hist. vol. xix. p. 1, t. 1-5.
1877. Stauronema Carteri, Zitt. Studien, I Ab. p.62; Neues Jahrbuch, p. 359.
Examples of this species in the Museum collection from the Chalk Marl of Barham
and the Isle of Wight, and from the Craie Chloritée of Cap la Heve, show the minute
spicular structures in a much clearer manner than the specimens from the Upper
Green Sand of Folkestone, on which Prof. Sollas based his descriptions; and they
enable me to add to, and partly to emend, the characters which he has assigned to
the species and genus.
The canals which pass into the true wall (oscular plate, So//as) from the inner or
concave surface of the sponge, do not appear to penetrate through it, as stated by
Sollas, but terminate blindly near the outer surface of the sponge, and, similarly, the
canals which enter the wall from the outer surface end blindly in the substance of
the wall. The canals in the supplemental skeleton (posterior mass) are altogether
irregular in their distribution. ‘The spicular mesh of the true wall is by no means
so regular in its disposition throughout as represented by Sollas in figs. 1—3, p. 7,
loc. cit., for though the mesh-interspaces are often circular, yet they are as often
quadrate, oval, or irregular in form.
Again, Sollas states (p. 7) that ‘“‘the outer margins of the fibres are so sharply
defined as to enable us to state with certainty that the fibres themselves are perfectly
smooth, and not in any way spined.” But in the example from La Heéve, not only
are the surfaces of the spicules bordering the canals thickly spined, which Sollas
observed in the Folkestone examples, but the spicular arms throughout the wall and
in the supplemental skeleton are clearly microspined. A remarkable feature in this
species is the supplemental skeleton (posterior mass, So//as), which is composed of a
spicular mesh, with arms or rays of about the same thickness as in the true wall, but
disposed in such a manner that the interspaces are extremely uregular, both in size
and form. This tissue appears to be of a similar character to that which in some
fossil sponges forms the stem and radical processes, and also grows over the lower
portion of the true sponge-wall. It is shown more particularly in the genus Plewrope,
Zittel, and is well developed in Craticularia (Laocetis) infundibulata*, Pomel. ‘Lhe
* Pal. d’Oran. 1866, p. 95, t. 1 bis, fig. 4.
02
100 SILICEOUS SPONGES.
dermal layer, which is formed of an extremely minute and delicate spicular reticu-
lation, extends alike over this supplemental skeleton as well as over the front surface
of the true wall.
The characters of Stawronema appear to me to exhibit a closer alliance to the
family of the Euretide than to the family of the Mellitionide, in which Prof. Zittel *,
relying upon the descriptions of Prof. Sollas, has placed the genus.
Prof. Sollas states that his specimens came from the Gault of Folkestone; but the
numerous specimens in the Museum collection from the same locality, are, according
to the opinion of Mr. R. Etheridge, sen., all derived from the Upper Green Sand f,
and as Sollas’s specimens were supplied to him by a local dealer, and were not
collected by himself, it seems probable that they also may have been obtained from
the Upper Green Sand, and not from the Gault.
Distribution. Upper Green Sand: Folkestone; near Eastbourne. Chalk Marl:
near Ventnor, Isle of Wight; Barham. Craie Chloritée: Cap la Héve.
STAURONEMA PLANUM, Hinde, n. sp. (Plate XXIV. figs. 2, 2a, 26, 2c.)
Sponge disk-shaped, with a circular outline; the upper surface nearly flat, the
under surface slightly convex, with concentric rounded ridges and furrows. ‘There
is no indication of a stem or point of attachment. ‘The margins are rounded and
slightly curve downwards. The width of the only specimen at present known is
70 mm. In the central portion the wall-plate is 12°5 mm. in thickness and near
the margin 6°3 mm.
The upper surface of the sponge is reticulate, with circular or ovate canal-apertures
about 1 mm. in diameter and about 1:3 mm. apart from each other. The canals are
slightly curved and appear to terminate blindly. No special canals are distin-
guishable on the under surface of the sponge.
The sponge-wall is composed of a regular meshwork of robust spicules with solid
nodes; the distance between the nodes is ‘35 mm. ‘The interspaces between the
mesh are subquadrate or nearly circular. The upper surface exhibits traces of a
finely reticulate dermal layer, which apparently extended over the canal-apertures.
The under surface also shows a delicate dermal layer of reticulate spicules, but its
state of preservation is insufficient for a close determination.
This species differs from S. Carteri, Sollas, in its form and in the absence of a
supplemental skeleton ; but it resembles that species in the character of the spicular
skeleton of the wall and the disposition of the canals.
The non-development in this species of the supplemental skeleton or posterior
* Neues Jahrbuch, 1877, p. 359.
+ Mr. F. G. H. Price also quotes this species from the Upper Green Sand or lower division of his Chalk
Marl at Folkestone, and he names this division the Stawronema-zone. No mention is made of its occurrence
in the Gault. (Quart. Journ. Geol. Soc. 1877, vol. xxxili. p. 434.)
STAURONEMA.—SESTRODICTYON. 101
mass, which is such a prominent feature in the type of the genus, does not appear
to me to be of sufficient importance by itself to justify placing this form in a separate
genus, and it would be -preferable to extend the definition of Stawronema so as to
include sponges in which a supplemental skeleton is not present.
The only example of this species has the spicular structure replaced by calcite,
and by a dark material, probably iron peroxide. ‘This dark substance has infilled the
canals of the hexactinellid spicules, which are thus distinctly shown, even when the
spicular arms themselves have been obliterated.
Distribution. Grey Chalk: Folkestone.
STAURONEMA comPactuM, /Tinde, n. sp. (Plate XXV. figs. 1, la, 14, le, 1d.)
Sponge apparently forming an ear- or fan-shaped expansion, growing from a small
rounded basal process; the wall is between 6:8 and 8 mm. in thickness. The only
specimen is the lower portion of an example, and measures 43 by 25 mm.
Both surfaces of the sponge-wall are furnished with oval canal-apertures from
‘75 to 1mm. in diameter, arranged in decussating lines. On what appears to be
the inner or upper surface these canal-apertures are about 2 lines apart, and the
interspaces between them consist of a thickened reticulate mesh with definite
circular pores. On the opposite surface the canal-openings are completely covered
in the lower part of the wall by a dermal layer of thickened irregular spicular
tissue ; somewhat higher the canals are only bridged over by this tissue, but not
concealed. ‘The canals are blind, and follow a slightly curved direction through
the wall.
The spicular mesh of the interior of the wall is composed of robust spicules with
compact nodes; the arms are apparently smooth; the distance between the nodal
centres is ‘33mm. The interspaces of the mesh are circular or oval.
Quenstedt* has figured a sponge from the Lower Planer of Bohemia, under the
name of Scyphia tenuis, which in the characters of the dermal layer and the canal-
structure closely resembles the present species; but according to Quenstedt its
spicular nodes are partly octahedral and partly solid, whereas in the present form
the spicular nodes are compact throughout.
Distribution. Grés vert (Upper Green Sand): France ?
Genus SESTRODICTYON, Hinde, n. g.
Sponge funnel-shaped, apparently by the infolding and coalescence of a plate-like
wall. The wall is perforated by numerous canals, arranged in a generally linear
direction, and occasionally decussating. ‘The skeletal mesh is composed of robust
spicules with compact nodes, which form a very regular quadrate mesh with small -
circular interspaces. No dermal layer appears to be present.
* Petrof. Bd. 5, p. 457, t. 137. f. 4.
102 SILICEOUS SPONGES.
This genus resembles Stawronema in the robust character of its spicular mesh, but
differs therefrom in the absence of a dermal layer and the perforate character of the
canals. The only specimen exhibits the spicular structure and the outer surface very
perfectly, but the interior is infilled, and quite concealed by the matrix.
SESTRODICTYON coNvoLUTUM, Hinde, n. sp. (Plate X XV. figs. 2, 2 a.)
The funnel-shaped sponge, which is imperfect at the base, measures 68 mm. in
height and 84 mm. in width at the summit. The canals are ovate in form, about
1:75 mm. in their greatest diameter; the distance between the rows is about 1:3 mm.
The spicules of the mesh have smooth arms; the distance between the nodal
centres is 3mm. Small pointed spines, apparently the free arms of the spicules
bordering the canals, project from the walls into the canals.
The outer surface characters of the specimen are so perfectly preserved, that if any
dermal layer had existed, one can hardly doubt but that traces of it would have
remained.
Distribution. Alpine Chalk (= Upper Green Sand): High Sentis, Canton Appenzell.
Genus BRACHIOSPONGIA, Marsh, 1867.
BRACHIOSPONGIA DiGiTaTa, D. Owen, sp.
1857. Scyphia digitata, D. Owen, Second Report on the Geol. of Kentucky, p. 111.
1838. Description dun fossile, Troost. Mém. de la Soc. Géol. de France, Tome 3, t. 11.
f. 8,9, 10.
1867. Brachiospongia Remerana et Lyonii, Marsh, Am. Journ. Science and Arts, 2 ser.
vol. 44, p. 88.
1878. Brachiospongia, Zitt. Handb. der Pal. Bd. 1, p. 173.
1880. Brachiospongia Remerana, F. Remer, Leth. geog. 1 Th. p. 319, fig. 61.
Up to the present nothing definite is known of the spicular structure of this extra-
ordinary sponge, and its true characters and position are therefore uncertain. By
Prof. Zittel it has been provisionally placed in the family of the Euretide. The
Museum only possesses a cast.
Distribution. Cincinnati Group: Kentucky.
Family COSCINOPORID4E.
Genus LEPTOPHRAGMA, Zittel, 1877.
Lepropuraema Mourcuisont, Goldf., sp.
1826-1833. Scyphia Murchisoni, Goldf. Petref. 1 Th. p. 219, t. 65. f. 8.
1877. Leptophragma Murchisoni, Zitt. Studien, I Ab. p. 48; Neues Jahrbuch, p. 357,
Vie aegale
LEPTOPHRAGMA.—PLEUROSTOMA. 103
Cup- or funnel-shaped sponges, with walls from 15 to 2 mm. in thickness.
Variable in size ; an average specimen measures 100 mm. in height and 75 in width
at the summit.
The canal-apertures of the outer surface are circular and very regularly disposed
in vertical and horizontal rows; they are about ‘5 mm. in width, and about the same
distance apart.
The spicular mesh has been replaced by peroxide of iron or dissolved away, and
is very indistinctly shown.
Distribution. Upper Green Sand (2): near Folkestone. Lower Chalk: South of
England. Upper Chalk: Haldem, Germany.
LEPTOPHRAGMA FRAGILIS, Remer, sp.
1840. Scyphia fragilis, F. A. Roemer, Nordd. Kreide, p. 8, t. 3. f. 11.
1877. Leptophragma fragilis, Zitt. Studien, I Ab. p. 48; Neues Jahrbuch, p. 358.
Sponge in the form of a shallow, open cup supported on an elongated cylindrical
stem. One specimen, imperfect at the margins, is 90 mm. in width; in another
specimen a stem 80 mm. in length and 65 in width, supports a portion of a cup.
The walls are only ‘5 mm. in thickness; their surfaces are closely covered with
minute canal-apertures ‘25 mm. wide, disposed in rows, or irregular in their distri-
bution. The mesh of the cup-wall has been changed into iron peroxide, and is not
recognizable. The spicules of the stem are elongate and irregular in form.
Distribution. Upper Chalk: South of England (coll. Bowerbank and Toul. Smith).
Genus PLEUROSTOMA, Rem. 1840, emend. Zitt. 1877.
PLEUROSTOMA RADIATUM, Ramer.
1840. Pleurostoma radiatum, Roemer, Nordd. Kreide, p. 5, t. 1. f. 11.
1877. Pleurostoma radiatum, Zitt. Studien, I Ab. p. 48; Neues Jahrbuch, p. 358.
The only specimen in the collection referable to this species is an elongated,
geniculate, much compressed stem 100 mm. in length, 14 mm. in width, and
3-5 mm. in thickness. At the basal end are several lateral root-like processes ;
the summit is imperfect. The walls of this hollow stem are about ‘75 mm. in
thickness. On both the lateral edges of the stem there are at irregular intervals
circular or ovate apertures from 2 to 3mm. in width, which open into the central
hollow. The outer surface of the wall is provided with numerous closely set circular
apertures -35 mm. in width, and about the same distance apart, and without definite
arrangement. ‘The spicular mesh has been replaced by iron peroxide.
Distribution. Upper Chalk: South of England (coll. 7. Smith).
104 SILICEOUS SPONGES.
PLEUROSTOMA BOHEMICUM, Zittel.
1877. Pleurostoma bohemicum, Zitt. Studien, I Ab. p. 48; Neues Jahrbuch, p. 358.
Istribution. Upper Chalk: Plauen, Bohemia (coll. Zittel).
Genus GUETTARDIA, Michelin, 1840-47.
GUETTARDIA STELLATA, Michelin (pars).
1840-47. Guettardia stellata, Michelin, Icon. Zooph. p. 121, t. 30. £. 8, 4, 6, 8-11, cet. excl.
1846. Guettardia Thiolati, D’Archiac, Mém. Soc. Géol. 2 sér. p. 197, t. 5. f. 15, and t. 8.
f. 5, 6, 7.
1848. Brachiolites angularis, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 357, f. O & P.
The number of the radiating flanges in this well-known sponge varies in different
examples from 3 to 10, the parallel walls are from 1 to 2mm. in thickness, and
about the same distance apart. The apertures on the outer lateral edges of the
flanges are ovate, and from 3 to 4mm. in width. The base is usually rounded or
with an imperfect stem; slender radical processes also appear to have originally
extended from the lateral edges of the flanges so as to serve in keeping the sponge
in an upright position. The central space appears to have been open above, and
there also appears to have been entirely free communication between the flanges and
the centre. There is great diversity in the size of different examples; a small form
is 28 mm. in height by 30 mm. in width, and some of the large examples, judging
by the extension of a single flange, must have been 240 mm. in width by 100 mm.
in height.
The walls carry thickly-set, minute, circular or oval canal-apertures, about ‘5 mm.
in width, disposed generally in quincunx. The spicules of the interior wall form an
irregular mesh with solid nodes. The dermal layer on the exterior surface, between
the canal-apertures, differs from the spicular mesh of the interior in a greater thick-
ness of the spicular arms, so that only minute circular pores remain between them.
The most complete description of this species is that given by ‘Toulmin Smith from
the examples in the English Chalk, which are either in the condition of rusty
peroxide of iron in the chalk itself, or preserved as reddish markings in flint. The
figure given by T. Smith, Joc. cit. p. 358, fig. O, is, to a certain extent, diagrammatic,
for the original in the Museum collection is by no means so complete as represented.
Michelin appears to me to have figured more than one species under G. stellata in
pl. 30, 7. ¢. Thus in the figures 2 and 5 the flanges grow upwards, and form
separate flattened branches, the same as in Plewrostoma, and only communicate with
the central cavity near their bases, and in this respect differ from the examples which
he has placed under figs. 3, 4, 6, 8, 9. For these latter forms I propose to retain
the name of G. stellata. D’Archiac has claimed one of these (fig. 6) as belonging to
his species G. Thiolat?, but he might equally well have claimed the other examples
GUETTARDIA.—COSCINOPORA. 105
(figs. 5, 4, 8, 9), as they clearly belong to the same species. The Museum possesses
several specimens from Biarritz which correspond very closely with D’Archiac’s figures
of G. Thiolati from that locality ; and I am unable to find any satisfactory differences
to distinguish them from the Chalk specimens of G. ste//ata, beyond those which may
arise from difference of preservation. Asa rule, however, they are smaller than the
Chalk forms.
The example figured by Mantell under the name of Ventriculites quadrangularis*,
is stated by T. Smith to be a fragment of this species; but neither Mantell’s figure
nor description is sufficiently clear for satisfactory recognition.
Distribution. Grey Chalk: Dover. Lower and Upper Chalk: Boxley, Kent;
Croydon, Shalford, Surrey; Glynde, Sussex; South of England (coll. 7. Smith).
Cretaceous: Biarritz (coll. Pratt).
GUETTARDIA RADIANS, Hinde, n. sp. (Plate XXYV. fig. 3.)
1840-47. Guettardia stellata, Michelin, pars, Icon. Zooph. pl. 30. f. 1, 2, 5.
Sponge consisting of a varying number, from 3 to 5 generally, of compressed
branches, which spring from a common, slightly inflated, hollow base, supported on
a short stem. ‘The branches are from 14 to 24 mm. in width; the parallel walls are
about 2 mm. in thickness, and about the same distance apart; circular apertures are
present on both the lateral margins. The wall-surface appears to be covered with
minute canal-apertures similar to those in G. stellata. No spicular structure is
present in the specimens.
I propose this species to include some of the forms included by Michelin under
G. stellata, but which differ from those to which I have restricted this term in the
fact that instead of forming flanges which open throughout their length into a central
cavity, the sponge divides into compressed branches, thus resembling a compound
Pleurostoma.
Distribution. Craie Chloritée: Vaches Noires. Biarritz? (coll. Pratt).
Genus COSCINOPORA, Goldfuss, 1826-33.
CosclNOPORA INFUNDIBULIFORMIS, (roldf.
1826-33. Coscinopora infundibuliformis, Goldf. Petref. 1 Th. p. 30, t. 30. f. 10.
1877. Coscinopora infundibuliformis, Zitt. Studien, I Ab. p. 49; Neues Jahrbuch, p. 359,
Up ree Ho
1878. Coscinopora infundibuliformis, Zitt. Handb. der Pal. p. 175, f. 91.
There are several examples from the Upper Chalk of the south ot England which
correspond in form and in the arrangement of the canal-apertures with typical
* Geology of Sussex, p. 177, t. 15. f. 6.
106 SILICEOUS SPONGES.
specimens from Germany; but as their spicular structure has been obliterated, a
satisfactory identification is impracticable.
Distribution. Upper Chalk: Croydon, Surrey; South of England; Coesfeld, Dahl-
feld, Westphalia.
CoscINoPoRA QUINCUNCIALIS, Toulmin Smith, sp.
1847-48. Ventriculites quincuncialis, T. Smith, Ann. & Mag. Nat. Hist. 1 ser. vol. xx. t. 7.
f. 7, and 2 ser. vol. i. p. 207.
Sponges narrow funnel-shaped, gradually tapering to a cylindrical stem, which
gives off horizontal root-like processes at its termination. An average specimen is
70 mm. in height, and 25 mm. in width at the summit.
The walls are from 2 to 2°5 mm. in thickness; the canal-apertures on the outer
surface are about ‘5 mm. in diameter, and disposed in very regular quincunx. Here
and there, in some specimens, traces of spicules can be detected ; they appear to have
had compact nodes. ‘The spicular structure of the stem is formed of elongated
irregular spicules; this structure overlaps the regular mesh-work of the wall in the
lower part of the body of the sponge.
This species is readily distinguished from C. infundibuliformis by its narrow
elongated form and the smaller size of the canals. Toulmin Smith states that the
walls of this species are folded the same as in Ventriculites, but I am unable to
recognize this character in his typical figured specimen. The specimens are preserved
in flint, and all appear to be incomplete at the summit.
Iistribution. Upper Chalk: Croydon, Surrey; Arundel, Sussex; Beckhampton,
Wiltshire.
Family MWELLITIONIDA, Zittel.
Genus APHROCALLISTES, Gray, 1858.
APHROCALLISTES ALVEOLITES, Ramer, sp.
1840. Scyphia alveoltes, Roemer, sp., Nordd. Kreide, p. 8, t. 3. f. 6.
1877. Aphrocallistes alveolites, Zittel, Studien, I Ab. p. 49; Neues Jahrbuch, p. 359.
The specimens which I refer to this species consist of two small fragments of the
sponge-wall, which is composed of prismatic canals 1-75 mm. in diameter, with very
delicate partitions. The spicular structure is now in the condition of iron peroxide ;
aud us the character of the mesh cannot be distinguished, there is some doubt as to
the correct identification of the specimens.
Distribution. Upper Chalk: South of England?
PACHYTEICHISMA.—TROCHOBOLUS. 107
Family VENTRICULITID A.
Genus PACH YTEICHISMA, Zittel, 1877.
PACHYTEICHISMA CarTERI, Zittel.
1877. Pachyteichisma Carteri, Zittel, Studien, I Ab. p. 50; Neues Jahrbuch, p. 360, t. 3.
f. 8; id. 1878, p. 59; Handbuch der Pal. p. 176, f. 92.
1808. An Alcyonite, Parkinson, Organic Remains, vol. 2, t. 11. f. 2.
1820. Spongus coarctatus ? Konig, Icones fossiles, p. 4, t. 7. f. 81.
1878. Lancispongia lamellosa tumulosa, Quenst. Petref. Bd. 5, p. 92, t. 119. f. 1.
Distribution. Upper Jura: Randen, Heuberg, Streitberg.
PACHYTEICHISMA LOPAS, Quenst. sp.
1858. Spongites lopas, Quenst. Der Jura, p. 685, t. 83. f. 5.
1877. Pachyteichisma lopas, Zitt. Studien, I Ab. p. 50; Neues Jahrbuch, p. 360.
1878. Lancispongia lopas, Quenst. Petref. Bd. 5, p. 95, t. 119. f. 3.
Distribution. Upper Jura: Heuberg, Wirtemberg.
PACHYTEICHISMA, sp.
Portions of the spicular structure of an undetermined species from the Jurassic
formation at Hohenpdlz, Bavaria (coll. Zittel).
Genus TROCHOBOLUS, Zittel, 1877.
TROCHOBOLUS CRASsICosTUS, Zitt.
1877. Trochobolus crassicostus, Zitt. Studien, I Ab. p. 50; Neues Jahrbuch, p. 360, t. 3.
f. 4; idem, 1878, p. 60.
1878. Scyphia barbata, Quenst. Petref. Bd. 5, p. 124, t. 120. f. 54.
Distribution. Upper Jura: Randen, Streitberg (coll. Zitt.).
TROCHOBOLUS LUCERNUS, [6nig, sp.
1820. Spongus lucerna, Konig, Icones fossiles, p. 4, t. 7. f. 83.
1820. Sponyus tuber, Konig, Icon. foss. p. 4, t. 7. £. 84.
1878. Muastospongia, sp., Quenst. Petref. Bd. 5, p. 148, t. 122. f. 7.
1878. Trochobolus, Zitt. Neues Jahrbuch, p. 61.
I am doubtful whether this peculiar sponge is rightly included in the genus
Trochobolus, for though it agrees with the type of the genus in its mode of growth,
I am unable to determine whether the spicular nodes are compact or lantern-like.
The spicular structure in all the specimens is replaced by either calcite or iron
peroxide, and in sections of calcite specimens the spicular nodes appear to be
compact; but it is doubtful whether the octahedral nodes can be detected in all
Pp 2
108 SILICEOUS SPONGES.
the cases in which replacement by calcite has occurred, though, as a rule, they are
pretty clearly shown. Should further research prove that the nodes are compact,
this species will have to be formed into a new genus, which will find its place close
to Verrucocelia.
Distribution. Upper Jura: Randen, Switzerland.
Trocuosotus constrictus, Hinde, n.sp. (Plate XXYV. figs. 4, 4a, 4 0.)
Sponge subcylindrical, surface uneven, with slight concentric swellings and depres-
sions, the summit truncate with roundea margins. Length of the only specimen
87 mm., width at summit 29 mm.
The thickness of the sponge-wall, measuring from the interior of the cloaca to the
outer surface, is 5 mm. ‘The outer surface exhibits a vertical series of ridges about
2mm. in width and 1 mm. apart. ‘The characters of the interior of the cloaca are
concealed by the matrix. The wall is built up of a regular spicular mesh with
lantern or octahedral nodes. The distance between the nodal centres is 312 mm.
The general appearance of this sponge so closely resembles the cylindrical examples
of Sphenaulax (Scyphia) costata, Goldf., that, judging from this character merely, it
would at once have been placed under this species ; but in a longitudinal section of
the wall the spicules exhibit distinct lantern nodes, and are similar to, though some-
what larger than, those of 7. crassicostus.
Distribution. Upper Jura: Randen, Switzerland.
Genus PHLYCTAINIUM, Zittel.
PHLYCTENIUM CONIFORMIS, Quenst. sp.
1878. Mastospongia coniformis, Quenst. Petref. Bd. 5, p. 152, t. 122. f. 13.
1826-33. Scyphia verrucosa, Goldf. p. p. Petref. 1 Th. p. 7, t. 2. f. 11.
1878. Phlyctenium coniformis, Zitt. Neues Jahrbuch, p. 61.
Distribution. Upper Jura: Heuberg, Wiirtemberg.
Genus VENTRICULITES, Mantel, 1822.
VENTRICULITES RADIATUS, Mantell, pars.
1822. Ventriculites radiatus, Mantell, Foss. of the South Downs, p. 168, t. 10, 11, 12, 14,
non t. 13. f. 2-6.
1815. Alcyonium chonoides, Mantell, Trans. Linn. Soc. vol. i. p. 402. t. 27-30.
1845-46. Scyphia radiata, Reuss, Ver. béh. Kreide, p. 74, t. 17. f. 14.
1854. Ventriculites radiatus, Mantell, Medals of Creation, 2 ed. p. 244, f. 81.
1870. Retispongia radiata, Ferd. Remer, pars, Geol. Oberschles. p. 302, t. 32, non t. 30. 1G
1878. Ventriculites radiatus discus, Quenst. pars, Petref. Bd. 5, p. 449, t. 136. f. 26.
VENTRICULITES. 109
Sponge consisting of a circular, flattened, or slightly concave, or occasionally
convex expansion, supported on a small funnel-shaped or inverted conical base, from
the extremity of which divergent root-like processes extend, which served to anchor
the sponge in the chalky mud.
The conical basal portion is from 20 to 30 mm. in height ; the summit expansion
attains, in fairly large examples, a diameter of 220 mm.
The wnder surface of the sponge exhibits a series of straight or slightly curved,
occasionally bifurcating ndges, which radiate from the centre to the margins. These
ridges are formed by the infolding of the thin wall-plaits of the sponge; they are
connected together at intervals by transverse extensions. ‘They are from 1:5 mm. to
2°5 mm. in thickness, and the intervening furrows are about 1 mm. in width.
The upper surface of the sponge has a smooth aspect ; it is composed of a reticu-
lated membrane with numerous circular or elliptical canal-apertures, from 1-5 to
3mm. in width, disposed either irregularly or in quincunx. ‘The total thickness of
the sponge-wall varies according to the amount of compression it has experienced ;
in some examples in the Chalk it is only 2 to 3mm., whilst in a flint-preserved
example it reaches a thickness of 6°5 mm.
In only a single specimen out of a large number of examples of this species have
I been enabled to discover traces of the spicular structure. The spicular mesh in
this specimen appears to be extremely regular; the interspaces are nearly quadrate,
and the distance between the spicular nodes is about 4 mm.
This species is probably the most abundant of any of the Ventriculites in the
English Chalk. It occurs either as mere impressions or as moulds filled with soft
ochraceous material in the chalk, or else as casts in flint. Very frequently the
basal or conical portion of a specimen is preserved in flint, whilst the expanded
portion remains as a mere impression in the soft chalk. Its appearance varies
greatly according to its mode of preservation, and the surface which is exposed
to view.
I have taken as the type of the species the specimens figured by Mantell on
tab. 14, ‘ Fossils of the South Downs.’ The figures, though roughly drawn, exhibit
very fairly both the upper and under surfaces of the sponge. The examples figured
by Mantell on tab. 13 belong, with one doubtful exception, to other species of
Ventriculites, and were so regarded by Mantell himself in the ‘ Medals of Creation,’
p- 245, f. 82.
Distribution. Upper Chalk: Norfolk; Surrey; Sussex; near Warminster, Wilt-
shire (coll. T. Smith and Mantel).
VENTRICULITES IMPRESSUS, Toulm. Smith.
1848. Ventriculites impressus, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. 1. p. 205;
and Ist ser. vol. xx. t. 8. f. 2, 3.
110 SILICEOUS SPONGES.
1848. Ventriculites muricatus, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 210, t. 18. f. 1.
1878. Ventriculites dilatatus, Quenst. Petref. Bd. 5, p. 460, t. 137. f. 6.
Sponge usually open funnel-shaped, gradually tapering to a slender stem with
divergent rootlets. In some instances the sponge forms flattened expansions with a
circular outline similar to V. radiatus. Height and width very variable; a large
specimen is 170 mm. in diameter.
The outer or under surface of the sponge-wall is reticulate, with circular or ovate
apertures, from *75 to 1-5 mm. in width ; the wall-fibres bounding the apertures are
about the same width as the interspaces. The upper or inner surface has numerous
apertures of a similar form to those of the lower, and disposed either irregularly or
in quincunx.
The total thickness of the wall is about 3mm.; but as all the specimens are in
chalk, and therefore compressed, it is impossible to determine the original thickness
with accuracy. No spicular structure is shown in any of the specimens.
This species appears to be nearly as abundant as the preceding. There is a great
amount of variation in different specimens, but I cannot find any characters which
would allow of a separation into distinct species. The V. muricatus, T. Smith,
appears to me, from a comparision of the original example in the Museum Collec-
tion, to belong to this species.
Distribution. Upper Chalk: South of England, Heytesbury; near Warminster,
Wiltshire (coll. T. Smith).
VenrRIcuLiTEs convoLutTus, Hinde, n. sp. (Plate XXV. figs. 5, 5 a.)
Sponges cup- or vase-shaped, with expanded margins and folded walls. No stem
preserved, and the species appears to have been sessile. The wall of a specimen
preserved in chalk is l-6 mm. in thickness. A fairly large specimen is 60 mm. in
height, and 120 mm. in breadth at the summit.
The outer surface is formed of flattened, bifurcating ridges about 1°5 mm. in wid th,
which run nearly parallel with each other from the base to the margins, and inter-
osculate so as to leave narrow, elongated interspaces. ‘The interior of the wall is
not exposed.
The spicular mesh, judging by the hollow casts in the Chalk, is irregular; the
distance between the spicular nodes is about 3mm. ‘This species may be distin-
guished by its folded thin walls and the disposition of the flattened ridges of the
outer surface.
In outer form and the disposition of the ridges of the outer surface of the wall,
this species resembles Cribrospongia subreticulata, Geinitz*; but the surface-apertures
* Paleontographica, Bd. 20, p. 23, t. 2. f. 2-4.
VENTRICULITES. pei!
in this latter species have a rectangular arrangement, and the wall is three times as
thick as in V. convolutus.
This species appears to be not uncommon in the Kentish Chalk.
Distribution. Upper Chalk: Gravesend, Broadstairs, Kent.
VENTRICULITES PocULUM, Zittel, MS.
Distribution. Upper Chalk: Libyan Desert (Zitt. coll.).
VENTRICULITES DECURRENS, Toulm. Smith.
1848. Ventriculites decurrens, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 215,
(ie IBY 1 fk
1848. Ventriculites decurrens, var. tenuiplicatus, T. Smith, ibid. p. 215, t. 13. f. 9.
1848. Ventriculites cavatus, T. Smith, ibid. p. 212, t. 13. f. 5.
1822. Ventriculites radiatus, Mantell, pars, Foss. South Downs, t. 13. f. 4.
1878. Ventriculites radidtus, Quenst. pars, Petref. Bd. 5, t. 136. f. 23.
Sponges varying in form, from subcylindrical to open funnel-shaped, gradually
tapering below to a slender cylindrical stem with branching roots. Small examples
measure 65 mm. in length by 25 in diameter, whilst a large specimen, without the
slender stem, is 120 mm. in height and of an equal width at the summit.
The outer surface of the sponge is formed by robust vertical or oblique ridges,
which occasionally bifurcate, but are rarely connected by lateral extensions. These
ridges are 2 mm. in thickness, and the intervening furrows about *75 mm. in width.
The interior surface exhibits circular apertures. ‘The entire thickness of the sponge-
wallis5 mm. The dermal layer of the exterior ridges of the wall is penetrated by
numerous minute circular pores, and minute spinous processes project outwards
from its surface. The spicular mesh of the interior appears to be of an irregular
character.
This species is also abundant. It somewhat approaches to V. radiatus in the
disposition of the ridges and furrows of the outer portion of the wall, but its mode
of growth readily distinguishes it from that species. The V. cavatus, T. Smith,
which is founded on a mere fragment of the wall of a sponge, appears to me to
belong to the present species.
Distribution. Upper Chalk: South of England; Bridgwick ; Monckton-Bassett,
Wilts (coll. T. Smith and Mantel).
VENTRICULITES MAMMILLARIS, Toulm. Smith.
1848. Ventriculites mammillaris, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 213,
flSee 7714:
1822. Ventriculites radiatus, Mantell, p. p. Foss. South Downs, t. 13. f. 2, 3, 5.
Sponges either elongated, narrow, funnel-shaped, or somewhat depressed, open
112 SILICEOUS SPONGES.
funnel- or cup-shaped. A slender cylindrical stem with branching roots is usually
present, but in some examples the root-processes spring directly from the base of the
funnel. The specimens vary from 50 to 120 mm. in height and from 20 to 75 mm.
in width. The outer surface of the sponge-wall is folded into vertical ridges about
2°8 mm. in width, and these are divided transversely so as to form so many rows of
rounded projecting wart-like bosses. The inner surface of the wall, according to
Toulmin Smith’s figure, appears to be composed of simple open furrows and ridges ;
but in none of the Museum specimens is the inner surface of the wall distinctly
shown, and it cannot be seen in the type specimen from which T. Smith’s figure has
been drawn, so that this portion of the figure is purely imaginary. In a specimen
preserved in flint the thickness of the wall is 4 mm.
‘The spicules of the interior mesh have conspicuous lantern or octahedral nodes ;
the spicular rays are about °25 mm. in length.
This species differs from V. decurrens mainly in the warty projections of the
outer wall.
Distribution. Upper Chalk: Sussex (coll. T. Smith and Mantell).
VENTRICULITES INFUNDIBULIFORMIS, S. Woodward. (Plate XXVI. figs. 1, 1 a.)
1833. Ventriculites infundibuliformis, 8. Woodw. Geology of Norfolk, t. 4. f. 20, 21.
1848. Ventriculites bicomplicatus, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i.
p- 219, fig. E.
1848. Ventriculites latiplicatus, T. Smith, ibid. p. 215, fig. D.
1848. Ventriculites striatus, T. Smith, ibid. p. 212, t. 13. f. 6, 15.
1848. Ventriculites radiatus, T. Smith (non Mantell), ibid. p. 218, t. 13. f. 10, 15.
Sponges elongated funnel-shaped, sometimes laterally compressed above, or slightly
expanding outwards; the divergent roots spring from the basal end of the funnel.
The total thickness of the wall is from 7 to 8:5 mm.; the plaits forming the wall are
about 1 mm. in thickness. I have not met with a complete specimen; the greatest
width at the summit of an individual is 83 mm.
‘The exterior surface is composed of rounded, occasionally bifurcating ridges, about
1-8 mm. wide, which extend vertically from the base to the summit, and are connected
laterally so as to leave narrow oval interspaces about 4 mm. in length by 1°25 mm.
in width, which are sometimes disposed in quincunx. ‘The inner surface of the wall
is rarely exposed ; it appears to be similar in character to the outer surface.
The spicular mesh is only shown by the empty moulds of the spicules in the
Chalk; it appears to be irregular, with prominent octahedral nodes; the distance
from node to node is *38 mm.
The aspect of this species varies greatly, according as it happens to be preserved
in the interior of flints or in chalk. Asa general rule only the lower portions of
the sponge are met with in flints, and these specimens do not show the oval inter-
VENTRICULITES. 113
spaces between the vertical ridges so conspicuously as in the Chalk examples. It is
probable that T. Smith’s figure of V. striatus has been drawn from one of the flint-
inclosed examples of this species.
This species is readily distinguished by the great thickness of the wall and the
characters of the outer surface. Though no description accompanies Woodward's
figure, the examples from Norwich in the Museum collection enable me to identify
the forms which he has placed under this name. Unfortunately the descriptions of
species given by Toulmin Smith are very meagre and incomplete, and many of his
figures are diagrammatic, so that reliance cannot be placed on them*. I have care-
fully examined Smith’s specimens in the Museum, and regard the forms referred by
him to V. bicomplicatus, V. latiplicatus, V. striatus, and V. radiatus as belonging to
the present species.
Distribution. Upper Chalk: Norwich; Clopton, Suffolk; Heytesbury, Beckhamp-
ton, Moncton-Bassett, Norton-Bavant, Wiltshire; Wolsk, Volga, Russia (coll. Bay-
field, Cunnington, T. Smith).
VENTRICULITES cRIBROSUS, Phillips, sp. (Plate XXVI. figs. 2, 2 a.)
1829. Spongia cribrosa, Phill. Geol. Yorkshire, t. 1. f. 7.
1864. Ventriculites multicostatus, F. A. Roemer, Paleont. Bd. 13, p. 19, t. 8. f. 1.
Sponges elongated, narrow, funnel-shaped, very gradually tapering from the summit
to the base; the root-processes usually spring from the basal end of the funnel, but
in some instances a short stem intervenes. ‘The wall is 6 mm. in thickness where it
has not been compressed. A fairly complete specimen is 240 mm. in length by
509 mm. in width at the summit.
The outer surface is reticulate; the ridges of the wall, about 1:2 mm. in width,
inosculate, so as to form elliptical interspaces, about 2°7 mm. in length by 1 mm. in
width, which, in some, though not in all the specimens, are disposed in quincunx.
The inner surface of the wall is concealed by the matrix. No spicular structure has
been preserved.
This species, so far as can be judged from the unfavourable condition of the
specimens, differs from V. infundibuliformis principally in its thinner wall and the
smaller size of the ovate apertures of the outer surface. Phillips’s figure is very
inexact, but there can hardly be a doubt respecting the form which he intended to
represent, since no other at all resembling it is found in the Chalk at Flamborough.
The sponge which F. A. Reemer has referred to this species (Nordd. Kreide, p. 9,
* Tt may also be mentioned here that the type of Toulmin Smith’s Ventriculites tessellatus (Ann. & Mag.
Nat. Hist. 2nd ser. vol. i. p. 211, t. 13. f. 2, 3, 4), now in the Museum, consists only of the impression of
the outer surface of a small fragment of a sponge on a small piece of chalk. No spicular structure can be
discerned, and it is doubtful whether the form even belongs to Ventriculites. The characters shown are
altogether insufficient for specific determination.
Q
114 SILICEOUS SPONGES.
t. 4. f. 2) differs from the Flamborough examples by its apparently thinner wall and
the circular form of the surface-apertures; but, on the other hand, the V. multi-
costatus of the same author, so far as can be judged from the descriptions and
figures, appears to be identical with Phillips’s species.
Distribution. Upper Chalk: Flamborough, Yorkshire.
VENTRICULITES aNGUSTATUS, Remer, sp. (Plate XXVI. figs. 3, 3 a, 3 0.)
1840. Scyphia angustata, Romer, Nordd. Kreide, p. 8, t. 3. f. 5.
1854. Scyphia angustata, Morris, Cat. Brit. Foss. p. 29.
Sponges narrow, funnel-shaped, occasionally compressed, gradually tapering from
the summit to the basal end. No stem or root has been preserved. ‘The thickness
of the wall in a silicified specimen is 5 mm., in specimens preserved in chalk it is
about 3°5 mm. The length of what appears to be an average specimen is 95 mm.,
and the summit width is 26 mm.
The outer surface has an irregular reticulate aspect; in places the ridges of the
wall, about °9 mm. in width, are disposed vertically, and connected by lateral exten-
sions, so as to leave transversely elliptical or irregularly subangular interspaces from
‘8 mm. to 1-2 in width; in other parts of the same specimen the disposition of the
ridges and interspaces is altogether irregular. The interior surface of the wall has
circular apertures, 1 mm. wide, and nearly the same distance apart, arranged regu-
larly in quincunx.
The spicular structure of the specimens is nearly obliterated, but I have ascer-
tained the octahedral character of the nodes. ‘The mesh appears to be irregular;
the distance from node to node is °35 mm. ‘The dermal layer, both of the outer and
inner surface of the wall, shows minute circular pores of different sizes.
This species can be readily distinguished from the funnel-shaped forms previously
described by the characters of the outer surface. This closely agrees with Roemer’s
figure of the type in the Nordd. Kreide; but it does not correspond with the forms
which the same author referred to the species at a later date under the name of
Cylindrospongia angustata*, nor with those which Ferd. Reemerf and Quenstedt f
have referred to the species. ‘This species appears to be rare.
Distribution. Upper Chalk: South of England.
VENTRICULITES ALCYONOIDES, Mantel.
1808. An Alcyonite, Parkinson, Organic Remains, vol. ii. p. 213, t. 10. f. 12.
1816. Flint Alcyonite, W. Smith, Strata Identified, t. 3. f. 1.
1822. Ventriculites alcyonoides, Mantell, Foss, South Downs, p. 176.
* Paleontographica, Bd. 13, p. 22, t. 8. f. 10.
+ Geol. Oberschlesien, p. 309, t. 30. £. 7, 8.
+ Petrefacten, Bd. 5, p. 457, t. 186. f. 2-14.
VENTRICULITES.—SCHIZORHABDUS. 115
1870. Cylindrospongia angustata, Ferd. Roemer, Geol. Oberschles. p. 309, t. 30. f. 7, 8.
1877. Ventriculites angustatus, Quenst. p.p. Petref. Bd. 5, p. 437, t. 186. f. 3, 11, 12, 14.
Sponges narrow, funnel-shaped or subcylindrical, sometimes with constrictions at
intervals. In some examples the aperture at the summit is contracted. The basal
end is seldom entire; the funnel appears to have been supported on a slender cylin-
drical stem. The wall appears to be about 4 mm. in thickness. In general the
specimens are about 50 mm. in length by 20 in width, but larger forms occur, which,
judging from fragments, would measure about 100 mm. long, with a summit-width
of 36 mm.
The outer surface of the wall is reticulate; the ridges or folds, about 1 mm. wide,
are disposed so as to leave circular or somewhat irregular interspaces, from ‘75 to
1 mm. in width, arranged either in quincunx or without definite order. The interior
surface of the funnel is not exposed.
The spicular mesh, as seen in a vertical section of a specimen, is irregular; the
distance between the spicular nodes is °375 mm.
It is sometimes difficult to separate some of the examples of this species which
have the surface-apertures of the wall arranged in quincunx, from specimens of
Coscinopora (Ventriculites) quincuncialis, 'T. Smith, sp. As a rule, however, the
canal-apertures in this latter species are much smaller, and the surface is much
more even than in V. alcyonoides.. In specimens which show the spicular structure
the difference is easily ascertainable, for in the type forms of C. guincuncialis the
nodes are compact, whilst in the present species they are clearly octahedral.
The figures of Parkinson and Smith referred to by Mantell must be accepted as
representing the type of the species, and not Mantell’s description, which embraces
more than one species. In the structure of the outer surface of the wall Ventri-
culites alcyonoides resembles V. impressus, T. Smith, and it is probable that the
disciform examples of Mantell’s description belong to this latter species.
This species appears to be abundant and generally distributed in the Upper Chalk.
Distribution. Upper Chalk: Wighton, Norfolk; Guildford, Surrey; Arundel, Lewes,
Sussex ; Boxley, Kent; Heytesbury, Norton-Bavant, Warminster, Wilts; Strehlen,
Germany (coll. Mantell, W. Smith, T. Smith).
Genus SCHIZORHABDUS, Zittel, 1877.
ScHIZORHABDUS LIBYCUS, Zitt.
1877. Schizorhabdus libycus, Zitt. Studien, I Ab. p. 51; Neues Jahrbuch, p. 361.
Distribution. Upper Chalk: Libyan Desert (coll. Zitt.).
Q 2
116 SILICEOUS SPONGES.
Genus RHIZOPOTERION, Zittel, 1877.
RHIZOPOTERION CERVICORNE, Goldf. sp.
1826-33. Siphonia cervicornis, Goldf. Petref. 1 Th. p. 18, t. 6. f. 11, and p. 98, t. 35. f. 11.
1877. Rhizopoterion cervicorne, Zitt. Studien, I Ab. p.51; Neues Jahrbuch, p. 362, t. 3. f. 6.
1878. Siphonia cervicornis, Quenst. Petref. Bd. 5, p. 422, t. 135. f. 9.
Two specimens of the stem and roots of a sponge preserved in flint probably
belong to this species. The spicular structure has not been preserved, but the
longitudinal canals very closely resemble those of typical examples from Germany.
Distribution. Upper Chalk: England; Bromberg, Haldem, Germany (coll. Zitt.).
Genus SPORADOSCINIA, Pomel, 1872, emend. Zttel, 1877.
SPORADOSCINIA MICROMMATA, Laem. sp.
1840. Scyphia micrommata, ¥. A. Roemer, Nordd. Kreide, p. 7, t. 2. fig. 11.
1872. Cribrospongia micromata, Schliiter, Spongit. des Miinsterlandes, p. 28.
1877. Sporadoscinia micrommata, Zitt. Studien, I Ab. p. 52; Neues Jahrbuch, p. 362,
lis Ws 1 Gs
Sponges cup-shaped or forming slightly concave expansions, supported on a short
inverted-conical, hollow, basal portion. No root-processes have been preserved.
The thickness of the wall is only 1 mm., but all the specimens have been replaced
by peroxide of iron, and it is probable that the wall has suffered from compression.
The height of a specimen is 45 mm. and the width across the summit is 105 mm.
The outer surface is furnished with irregularly disposed, circular or transversely
ovate apertures, about *9 mm. in width, and the same or a slightly greater distance
apart from each other; the inner surface of the wall has elliptical apertures, about
1:3 mm. in length, disposed regularly in quincunx. No spicular structure has been
preserved. The dermal layer appears to have been minutely porous.
Distribution. Upper Chalk; Bromley, Kent (coll. 7. Smith).
SporaposcintA Decuent, Goldf. sp.
1826-33. Scyphia Dechenii, Goldfuss, Petref. 1 Th. p. 219, t. 65. f. 6.
1877. Sporadoscinia Decheni, Zitt. Studien, I Ab. p. 52; Neues Jahrbuch, p. 362.
1878. Scyphia Dechenii, Quenst. Petref. Bd. 5, p. 454, t. 137. f. 2.
Microscopic fragments of the spicular structure of this species from the Upper
Chalk of Coesfeld, Westphalia.
SPORADOSCINIA CAPAX, Hinde, n.sp. (Plate XXVI. figs. 4, 4a, 40.)
Sponge open funnel-shaped, growing to a large size; no stem has been preserved.
The wall is 5 mm. in thickness; it is possible that it may have been originally of
SPORADOSCINIA.—SESTROCLADIA. Li
somewhat greater thickness, though I do not think that in this instance it has been
much compressed. The only specimen, which is destitute of the lower portion of
the funnel, measures 17() mm. in height by 168 in width at the summit.
The outer surface of the wall is pierced with irregularly disposed, transversely
elliptical or ovate apertures, from 1 to 1-8 mm. in length and about ‘6 mm. in width.
These apertures are only separated by slender wall-fibres, about *7 mm. in width.
The inner surface has circular or slightly ovate apertures, 1°6 mm. wide, and about
the same distance apart, regularly disposed in quincunx.
Only a portion of the wali has been preserved, and the spicular structure is either
in the condition of hollow moulds, or these are partially filled with peroxide of iron.
The interior mesh appears to be very irregular; the distance between the octahedral
spicular nodes is‘3 mm. The dermal layer appears to have been furnished with
stout spines, which projected from the surface and into the canals in the same
manner as represented by Zittel in S. micrommata*.
The main difference between this species and S. Decheni, Goldf., adenendent of
its much larger size, consists in the different form and arrangement of the apertures
of the outer surface of the wall.
Distribution. Lower Chalk: South of England.
Genus SESTROCLADIA, Hinde, n. g
Sponge ramose, consisting of cylindrical or compressed bifurcating branches. ‘The
branches are hollow tubes, open at their summits. The outer surface of the sponge
is formed by anastomosing ridges or folds with ovate interspaces. The interior
surface is concealed by the matrix, but, judging from vertical and transverse sections,
it appears to be penctrated by apertures similar to those of the outer surface. The
spicular mesh of the interior of the wall is irregular, the spicular nodes are octa-
hedral. The outer surface of the ridges or folds of the wall apparently possessed a
dermal layer, but its minute structure is not discernible.
This genus is distinguished from all others of the family of the Ventriculitide by
its dendritic mode of growth.
SESTROCLADIA FuRcaTUS, Hinde, n.sp. (Plate X XVII. figs. 1, la, 10.)
Sponge apparently growing from a simple stem, which gives off divergent branches
at irregular intervals. The branches have a generally upright direction of growth ;
they are from 15 to 28 mm. in diameter, usually cylindrical, but near the points of
bifurcation they are frequently compressed and expanded. The single example,
which is imperfect, is 155 mm. in height, and the lateral extension is 140 mm. ‘The
thickness of the wall is 3 mm.
* Neues Jahrbuch, 1877, Taf. 3. fig. 5a.
118 SILICEOUS SPONGES.
The ridges or folds of the outer surface are about 1 mm. in width, and the ovate
or somewhat irregularly elongate canal-apertures are from 1 to 2 mm. in length.
The octahedral nodes of the interior mesh are clearly shown; the distance from node
to node is about -4 mm.
There is but a single specimen in the Museum collection, which had been named
Spongia ramosa, Mantell; but beyond its mode of growth it has no other character
in common with that species.
Distribution. Grey Chalk: Dover.
Genus COXLOSCYPHIA *, Tate, 1865 (emend. Hinde),
This name was proposed by Tate as a substitute for Polycelia, Fromentel f,
which had been previously applied by Prof. King § to a genus of Actinozoa. As
representing a natural genus of sponges, Polycelia, From., is quite valueless, and
the forms included therein by Fromentel and F. A. Roemer have been placed by
Zittel in two or three genera. It is needless therefore to adopt Tate’s suggestion
and substitute another term for a genus which is now obsolete. “The particular
species, however, that Tate placed under Celoscyphia appears to me to belong toa
distinct genus, and I therefore propose to retain his name for sponges with the
following generic definition :—
Sponges consisting of simple cylindrical tubes growing from a common centre.
The walls of the tubes regularly folded so as to form vertical subparallel ridges with
intervening furrows. ‘The spicular mesh with octahedral nodes.
This genus is intermediate between Sestrocladia and Polyblastidium. 'The non-
branching character of the tubes and the vertical ridges of the wall distinguish it
from the former genus, whilst the absence of a common tubular axis separates it from
the latter.
Ca@.oscyPuia suLcata, Tate, sp. (Plate XXIX. figs. 1, 1a.)
1865. Celoscyphia sulcata, Tate, Quart. Journ. Geol. Soc. vol. xxi. p. 43, t. 5. £. 5.
1878. Celoscyphia sulcata, Zitt. Studien, III Ab. p. 31, note.
The only specimen consists of three short subequal cylindrical tubes, 17 mm. in
length and 9 in width, growing from a common centre. No stem has been preserved,
but it is probable that one was originally present. The walls of the tubes are
folded so as to form 12 to 14 vertical ridges; the intervening furrows are concealed
by the matrix, which also fills the cloacal tube. The cloacal aperture is circular,
and 3mm. in width. The total thickness of the wall is about 3 mm., whilst the
* This genus is unintentionally omitted from the list on p. 16.
t+ Quart. Journ. Geol. Soc. vol. xxi. p. 43.
{ Introduction a l’étude des Eponges fossiles, 1859, p. 32.
§ Annals & Mag. Nat. Hist. 1849, 2nd ser. vol. iii. p. 388.
C@LOSCYPHIA.—POLYBLASTIDIUM. 1g
wall-plaits are ‘5 mm. in thickness. The spicular mesh is small; the octahedral
nodes are *3 mm. apart. The characters of the dermal layer are obliterated, and the
interior spicular structure is only preserved in one or two places. The type speci-
men, the only one at present known, is in the Jermyn-Street Museum.
Distribution. Spongarian zone of the Chloritic Chalk (Zate) = Upper Chalk:
Island Magee, near Belfast.
Genus POLYBLASTIDIUM, Zittel, 1877.
POLYBLASTIDIUM LUXURIANS, Zitt.
1877. Polyblastidium lucurians, Zitt. Studien, I Ab. p. 52; Neues Jahrbuch, p. 363,
ney sini
Microscopic portions of the spicular structure of this species.
Distribution. Upper Chalk: Linden, Hanover (coll. Zitt.).
PoLYBLASTIDIUM RACEMOSUM, Toulm. Smith, sp. (Plate XXVII. figs. 2, 2 a.)
1848. Brachiolites racemosus, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 364, t. 15. f. 6.
Body of sponge club- or pear-shaped, supported on a slender stem, with divergent
rootlets at its termination. From the surface of the sponge short projecting hollow
tubes, from 8:5 to 14 mm. in diameter, project irregularly outwards. An average
specimen is 77 mm. in length by 40 in width.
The examples of this species are all in solid flint, and only transverse sections of
the tubular projections are shown on the outer surface of the flint. The walls of
the tubes are 2:5 mm. in thickness; they are formed of regular plaits, 6 mm. thick.
The spicular mesh appears to be irregular; the distance between the octahedral
nodes is ‘25 mm.
This species differs from P. luaurians, Zitt., by the larger size and lesser number
of the projecting tubes.
Toulmin Smith’s figure is diagrammatic, and a very inaccurate representation of
the original.
Distribution. Upper Chalk: Kent, Sussex (coll. T. Smith).
PoLyBLASTIDIUM TUBEROSUM, Toulm. Smith, sp.
1848. Brachiolites tuberosus, T. Smith, Ann..& Mag. Nat. Hist. vol. i. p. 354, t. 15. f. 3.
The type of this species is a fragment of a sponge, which has been partially cleaned
from a chalky matrix. Only the general form, indicated by markings in iron per-
oxide, can be distinguished. The body of the sponge appears to have been a thin-
walled, hollow, subcylindrical tube, with small bud-like projections on its exterior
surface. The tubular axis is about 46 mm. in length by 16 in width. The pro-
jecting buds are 8 mm. in length; their summits are expanded and slightly concave.
120 SILICEOUS SPONGES.
No aperture can be detected in the summit of the buds; but the condition of the
specimen is too imperfect for satisfactory determination. The figure of T. Smith’s
is to some extent a restoration of the original specimen. The closed summits of the
bud-like projections distinguish this species from P. luxurians, Zitt.
Distribution. Upper Chalk: Kent? (coll. T. Smith).
Genus CEPHALITES, Toulm. Smith, pars, 1848.
Toulmin Smith included in this species a variety of forms, which have been sepa-
rated by Prof. Zittel into two or three genera. As restricted by Zittel, the genus
only includes those sponges which resemble Ventriculites in the foldings of the plaits
of the wall, and possess the further characteristic that the summit of the sponge-
wall is truncate and covered with a delicate siliceous membrane.
CEPHALITES LONGITUDINALIS, Toulm. Smith.
1847-48. Cephalites longitudinalis, T. Smith, Ann. & Mag. Nat. Hist. Ist ser. vol. xx. t. 7.
f. 1, and 2nd ser. vol. i. t. 14. £. 1.
1848. Cephalites guttatus, T. Smith, ib. 2nd ser. vol. i. t. 14. f. 2.
1878. Cephalites longitudinalis and guttatus, Zitt. Studien, I Ab. p. 52; Neues Jahrbuch,
p- 363.
Sponges narrow, funnel-shaped, with a short slender cylindrical stem, terminating
in divergent rootlets. An average specimen is 68 mm. in length, and 25 in width at
the summit. The wall varies from 4 to 13 mm. in thickness in different individuals.
The exterior surface is formed by a series of longitudinal ridges about 3 mm. in width,
more or less sinuous, and occasionally interrupted transversely so as to form rows of
projecting bosses. ‘The inner surface has numerous circular canal-apertures, 1-5 mm.
wide, disposed in quincunx. The plaits forming the wall are about 1 mm. in thick-
ness. The spicular mesh is irregular; the distance between the spicular nodes is
about *25 mm.
I am unable to detect specific differences between C. guttatus and C. longitudinalis.
Even in the same specimen the outer surface will occasionally exhibit continuous
ridges and rows of projecting bosses. The examples of this species are common;
they are nearly all preserved in chalk, and the spicular structure is usually in the
condition of iron peroxide. In some examples of this and other species-of the genus
this replacing material is sufficiently firm for the spicular mesh to remain intact after
careful removal of the matrix, but it is of an extremely delicate character.
Distribution. Upper Chalk: Glynde, Westmeston, Sussex; Shalford, Surrey ;
Maidstone, Dover, Kent (coll. 7. Smith).
CEPHALITES. 121
CEPHALITES PARADOXUS, Toulm. Smith.
1848. Cephalites paradoxus, T, Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 283,
t. 14. f. 3.
1878. Cephalites paradoxus, Zitt. Studien, I Ab. p. 52; Neues Jahrbuch, p. 363.
Sponges funnel-shaped, supported on a short stem with diverging roots. An
average specimen is 80 mm. in length, and 34 in width, at the summit. The wall
varies from 4°5 to 8mm. in thickness. The wall-ridges of the outer surface are
2mm. in width; they anastomose with each other so as to leave irregular inter-
spaces. The interior surface appears to be formed by straight longitudinal ridges
and furrows. The disposition of the interior plaits of the wall appears to be the
same as in the preceding species.
Distribution. Upper Chalk: Kent (coll. T. Smith).
CrpHaites Benertia, Mantell, sp.
1822. Ventriculites Benettie, Mant. Foss. of the South Downs, p. 177, t. 15. f. 3.
Sponges funnel-shaped, with divergent roots, apparently springing from the base of
the funnel. ‘The type specimen is 59 mm. in length and 35 in width; the wall at the
summit is 9 mm. in thickness.
The outer surface is composed of anastomosing ridges, about 2 mm. in width,
disposed so as to leave oval or irregular interspaces from 2 to 6 mm. in length; the
interior surface of the wall appears to possess circular canals about 2 mm. in diameter.
The interior plaits of the wall are not shown.
So far as can be judged from the condition of the specimen, this species differs
from C. paradoxus principally in the characters of the interior surface of the wall.
Mantell’s figure is a very rude and not altogether accurate representation of the
original, which is in the Museum.
Distribution. Upper Chalk: Lewes, Sussex (coll. Mant.).
CEPHALITES ALTERNANS, Toulm. Smith.
1847-48. Cephalites alternans, T. Smith, Ann. & Mag. Nat. Hist. Ist ser. vol. xx. t. 7. f. 2,
and 2nd ser. vol. 1. p. 283, t. 14. f. 4, 5.
Toulmin Smith’s type of this species is a small funnel-shaped sponge, 28 mm. in
length by 20 in width; the thickness of the wall is 6mm. The outer surface is
apparently composed of reticulating ridges, about 1-5 mm. in thickness, with irregular
interspaces ; the interior surface has ovate canals, about 1:8 mm. in diameter, disposed
in quincunx. The plaits of the interior of the wall are disposed in somewhat
irregular folds, as shown in Smith’s figure, 7. c. vol. xx. t. 7. f. 2; the simple plait
is about 1 mm. in thickness. ‘The distance between the octahedral spicular nodes
R
122 SILICEOUS SPONGES.
is‘3mm. The ridges of the outer surface of the wall of this species are more slender
than in C. Benettiw, and the form itself is smaller.
Distribution. Upper Chalk: South of England.
CEPHALITES BULLATUS, Toulm. Smith. (Plate XXVII. figs. 5, 3a, 3b.)
1847-48. Cephalites bullatus, T. Smith, Ann. & Mag. Nat. Hist. Ist ser. vol. xx. t. 7. f. 3,
and 2nd ser. vol. i. p. 284, t. 14. f£. 6.
1816. Flint Alcyonite, W. Smith, Strata identified by Org. Fossils, p. 7, t. 3. f. 2.
Sponges with a funnel-shaped or subcylindrical body, about 56 mm. in length, and
from 21 to 56 mm. in width, supported on a delicate stem, with divergent rootlets at
its termination. The wall varies from 5-5 to 12 mm. in thickness.
The wall of the outer surface is folded in such a manner as to form, on the sur-
face of perfect specimens, small, flat-topped, lozenge-shaped or irregular projections,
about 4 mm. wide, which are disposed in quincunx. The dermal membrane of these
projections is minutely porous, and small, hollow, occasionally bifurcating, spines
spring from its surface. ‘The interior surface of the wall is perforated with circular
or elliptical canal-apertures about 2mm. in diameter. The plaits of the interior
portion of the wall are about ‘8mm. in width; they are disposed in deep folds
extending from the outer nearly to the inner surface, and wice versd; they also
form occasional loops.
This species is distinctly characterized by the features of the outer surface.
T. Smith’s figured specimen does not appear to be in the Museum Collection, but
there are several examples labelled by him from which the above description has
been taken. It appears to me very doubtful if the forms referred to this species by
Reuss* and Quenstedtt really belong to it.
Distribution. Upper Chalk: South of England (coll. 7. Sinith).
JEPHALITES CATENIFER, Toulm. Smith.
1848. Cephalites catenifer, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 286, t. 14.
f. 14, 15, 16.
1848. Cephalites compressus, T. Smith, ib. p. 287, t. 14. f. 10.
1877. Toulminia catenifer and compressus, Zitt. Studien, I Ab. p. 56; Neues Jahrbuch,
p. 368.
Sponges funnel-shaped or inverted conical, supported on a slender stem. An
average example is 48 mm. in length and 34mm. in width; the stem appears to
have been about 15 mm. in length. The wall is from 11 to 14 mm. in thickness.
The outer surface of the wall is composed of ridges about 1:8 mm. in width,
* Verst. d. bohm. Kreide, II Ab. p. 74, t. 18. f. 11.
+ Petref. Bd. 5, p. 484, t. 138. f. 10, 11.
CEPHALITES.—CYPELLIA. 123
disposed either in horse-shoe form or in irregular loops, so that the interspaces have
a generally sinuous character. The interior, or cloacal surface, is perforated with
cylindrical canal-apertures.
The interior of the wall is composed of: plaits, about 1:25 mm. wide; seen in
transverse section, they are arranged in deep folds with occasionally intervening
loops. The spicular mesh of the plaits is more regular than in the preceding
examples; the nodes are lantern or octahedral, and the distance from node to node
is 38 mm.
The C. compressus, T. Smith, appears to be merely the upper fragmentary portion
of a specimen of C. catenifer. The original of Smith’s figure, /. c. t. 14. f. 10, has no
trace of a stem or root as represented in the figure. The representation of a transverse
section of the plait in f. 15 is partly a restoration.
I am unable to follow Prof. Zittel in separating this species to form a distinct
genus. Seen in transverse section, the plaits of the wall do not appear to be more
complicated than in the typical forms of Cephalites, and they do not seem to possess
the distinctive characters of the Meandrospongide.
Toulmin Smith has given a description and figure of another species of Cephalites,
under the name of C. retrusus*; but the figured type in the Museum Collection is
merely a mould in wax, from which it is impossible to determine whether the
specimen has been a sponge or not.
Distribution. Upper Chalk: South of England (coll. T. Smith).
Family STAURODERMIDA, Zitt.
Genus CYPELLIA, Pomel (emend. Zitt. 1877).
CYPELLIA RUGOSA, Goldf. sp.
1826-33. Scyphia rugosa, Goldf. Petref. 1 Th. p. 9, t. 3. f. 6.
1877. Cypellia rugosa, Zitt. Studien, I Ab. p.53; Neues Jahrbuch, p. 364, t. 4. f. 1; Hand-
buch der Pal. 1 Bd. p. 179, f. 94.
Distribution. Upper Jura: Streitberg, Franconia.
CYPELLIA INFUNDIBULIFORMIS, (roldf. sp.
1826-33. Scyphia rugosa, var. infundibuliformis, Goldf. Petref. 1 Th. p. 87, t. 32. f. 2.
1878. Crucispongia annulata and cruciata, Quenst. Petref. Bd. 5, p. 165, t. 128. f. 2-5.
Distribution. Upper Jura: Heuberg, Streitberg, Randen. Oolite inferieure:
Verson, France.
* Annals & Mag. Nat. Hist. 1848, 2nd ser. vol. i. p. 285, pl. 14. f. 8.
Rez
124 SILICEOUS SPONGES.
CYPELLIA Ca&SPITOSA, Quenst. sp.
1878. Dolispongia cespitosa hexamera, Queust. Petref. Bd. 5, p. 311, t. 1380. f. 12.
Distribution. Upper Jura: Randen?
CYPuULLIA LIBERA, Quenst. sp.
1878. Nevxispongia libera, Quenst. Petref. Bd. 5, p. 162, t. 123. f. 1.
1878. Cypellia prolifera, Zitt. Neues Jahrb. p. 62.
Distribution. Upper Jura: Heuberg, Randen.
Genus STAURODERMA, Zittel, 1877.
STAURODERMA LocHENSE, Quenst. sp.
1858. Spongites Lochensis, Quenst. Der Jura, p. 669, t. 81. f. 96.
1877. Stauwroderma Lochense, Zitt. Studien, I Ab. p. 53; Neues Jahrbuch, p. 364, t. 4. f. 2.
Distribution. Upper Jura: Heuberg, Streitberg.
STAURODERMA CYLINDRATUM, Quenst. sp.
1858. Spongites cylindratus, Quenst. Der Jura, p. 686, t. 83. f. 8.
1878. Cavispongia cylindrata, Quenst. Petref, p. 158, t. 122. f. 19, 20.
As seen in a polished section, the interior structure of the wall of this species is
composed of relatively large spicules with solid nodes, forming an irregular mesh
similar to that of Stawroderma Lochense. 1 have not, however, detected any dermal
layer of distinctive cross-shaped spicules; so that it is doubtful whether this form
rightly belongs to the genus Stauroderma or not.
Distribution. Weuberg, near Balingen, Wiirtemberg.
Genus PURISIPHONLIA, Bowerd. 1869.
PURISIPHONIA CLARKEI, Bower.
1869. Purisiphonia Clarkei, Bowb. Proc. Zool. Soc. p. 342, pl. 25. f. 6, 7.
1870. Purisiphonia Clarkei, Moore, Quart. Journ. Geol. Soc. vol. xxvi. pp. 235, 240,
Dlelvepiemle
1878. Purisiphonia Clarkei, Carter, Ann. & Mag. Nat. Hist. 5th ser. vol. i. p. 376.
1878. Purisiphonia Clarkei, Zitt. Handbuch der Pal. vol. i. p. 179.
1878. Purisiphonia Clarkei, R. Etheridge, jun., Cat. Austral. Foss. p. 104.
Distribution. Lower Cretaceous?: Wollumbilla Creek, North Queensland, Aus-
tralia (coll. Bowerbankh).
POROCY PELLIA.—OPRHYSTOMA. 125
Genus POROCYPELLIA, Pomel (emend. Zitt. 1877).
POROCYPELLIA PYRIFORMIS, Goldf. sp.
1826-33. Scyphia pyriformis, Goldf. Petref. 1 Th. p. 10, t. 3. f. 9.
1877. Porocypellia pyriformis, Zitt. Studien, I Ab. p.53; Neues Jahrbuch, p. 364, t. 5.
ie Aa ee
Distribution. Upper Jura: Streitberg (coll. Zitt.).
Genus CASEARIA, Quenst. 1858.
CASEARIA ARTICULATA, Goldf. sp.
1826-33. Scyphia articulata, Goldf. Petref. 1 Th. p. 9, t. 3. f. 8.
1858. Casearia articulata, Quenst. Der Jura, p. 681, t. 82. f. 9.
1877. Casearia articulata, Zitt. Studien, I Ab. p. 54; Neues Jahrbuch, p. 365, t. 5. f. 2a, d.
1878. Spongites articulatus, Quenst. Petref. Bd. 5, p. 107, t. 120. f. 8-10, 12-21.
Distribution. Upper Jura: Streitberg; Heuberg ; Heiligenstadt; Giengen ; Randen.
Genus POROSPONGIA, D’Orbigny, 1852.
Porosroncia MARGINATA, Goldf. sp.
1826-33. Manon marginatum, Goldf. pars, Petref. 1 Th. t. 34. f. 9g, h.
1877. Porospongia marginata, Zitt. Studien, I Ab. p. 55; Neues Jahrbuch, p. 366.
Distribution. Upper Jura: Heuberg, Wirtemberg.
Porosponaia impress, MViinst. sp.
1826-33. Manon impressum, Mist. in Goldf. Petref. 1 Th. t. 34. f. 10.
1877. Porospongia impressa, Zitt. Studien, 1 Ab. p.55; Neues Jahrbuch, p. 366; Handbuch
der Pal. vol. i. (1878), p. 180, f. 95 a, 6, c.
Distribution. Upper Jura: Streitberg; Muggendorf, Franconia (coll. Zitt.).
Genus OPHRYSTOMA, Zitt. 1877.
OPHRYSTOMA MICROMMATUM, Ram. sp.
1864. Porospongia micrommata, Reem. Pal. Bd. 13, p. 9, t. 4. f. 14.
1877. Ophrystoma micrommatum, Zitt. Studien, I Ab. p. 55; Neues Jahrbuch, p. 366.
Sponge growing in flattened expansions of various dimensions. An imperfect
specimen is 80 mm. in width. The upper surface is smooth, and is provided with
numerous, irregularly disposed, circular or ovate apertures, from 2°5 mm. to 4mm. in
width and from 2°5 to7‘5 mm. apart. When perfect these apertures have distinctly
126 SILICEOUS SPONGES.
elevated, crater-like margins. The under surfaces in all the specimens is firmly
attached to the rocky matrix, and its characters are not visible. The thickness of
the sponge is about 5°5 mm.
The upper surface, or dermal layer, is composed of a very delicate, minutely porous,
siliceous network; the spicular structure is so minute that it is not distinguish-
able unless under the microscope; beneath this dermal layer the loose tissue of the
interior skeleton is exposed. This is formed of spicules with stout arms and lantern
nodes. The distance from node to node varies between *28 and :37 mm. In some
places the interspaces of the mesh are filled with minute slender hexactinellid
spicules, apparently disposed irregularly.
There are three examples of this species, which, so far as I am able to judge, corre-
spond with Reemer’s figure and meagre description. One specimen is from the Grey
Chalk, and is now calcareous in its composition ; the other two, from the Chalk Marl,
still retain in part their siliceous structure, and from these I have been able to
ascertain the spicular characters.
Distribution. Chalk Marl: Ventnor. Grey Chalk: Dover.
OPHRYSTOMA OCELLATUM, Seeley, sp.
1873. Porospongia ocellata, Seeley, MS. See Proceedings Geol. Soc. vol. xxix. p. 68, pl. 6.
f, 1-4.
1873. Ventriculites cavatus, Sollas (non T. Smith), Proc. Geol. Soc. vol. xxix. p. 68.
Sponges forming flattened or slightly curved expansions with rounded margins.
The wall is from 3°5 to 5mm. in thickness. The upper surface is perforated with
oval apertures from 2 to 3mm, in diameter, and about their own diameter apart,
disposed in diagonal lines or irregularly. The margins of these apertures do not
project as in the preceding species. The characters of the under surface are con-
cealed in the specimens which I have examined.
The spicular mesh of the interior of the wall is of an open irregular character.
The arms are robust, with prominent lantern nodes; the distance from node to node
is‘4mm. ‘The spicular structure of the dermal layer has been obliterated.
This species differs from the preceding in its smaller size and more regular form, also
in the generally smaller dimensions and regular disposition of the surface-apertures ;
the spicular structure of the interior is also more open, and the spicular arms are
larger than in 0. micrommatum. Only empty moulds of the spicular skeleton of the
wall are preserved ; good representations of these are given by Sollas (J. ¢.).
The name of this species was first given by Prof. Seeley to specimens in the
Cambridge Woodwardian Museum; and it was afterwards figured by Sollas under
the same designation, though in his description it is placed under the name of
Ventriculites cavatus, T.Smith. There is, however, no alliance, beyond the common
OPHRYSTOMA.—CINCLIDERMA. 127
possession of spicules with lantern nodes, between this form and the fragment of a
sponge which T. Smith named V. cavatus.
Distribution. Green Sand: Cambridge. According to Prof. Seeley it also occurs
in the Chalk at Hunstanton, Norfolk.
Genus PLACOTREMA, Hinde, gen. nov.
Sponges forming flattened expansions with rounded margins. ‘The smooth upper
surface is perforated with numerous apertures. The dermal layer is formed by large
cross-shaped spicules, disposed over each other without any regularity ; the interspaces
between their arms are occupied by a very minute spicular mesh. The interior
skeleton consists of a spicular mesh arranged so as to form delicate anastomosing
lamine. The nodes of the spicules are solid.
This genus differs from Porospongia, D’Orb., and Ophrystoma, Zitt., in the dispo-
sition of the interior mesh. From this latter genus also, which in general form it
closely resembles, it is further distinguished by the different character of the dermal
layer and the compact spicular nodes of the interior skeleton. It appears to be allied
to Placochlenia, Pomel; but as the characters of the spicular mesh of this genus are
not given by the author, it is not possible to compare this structure in the two genera ;
the dermal layer also differs from that of Placochlenia.
PLACOTREMA cRETACEUM, /Hinde, n. sp. (Plate XXVII. figs. 4, 4a, 46, 4c.)
This sponge appears to have grown in flattened masses of an elliptical or ovate
outline; the largest specimen is 60 mm. in length by 50 in width. ‘The wall varies
from 7 to 9-5 mm. in thickness. The apertures of the upper surface are ovate, with
depressed margins, and disposed either in diagonal rows or irregularly. They are
from 2°5 to 4 mm. in width.
The large cross-shaped spicules of the dermal layer are of various sizes; the ray
of a fairly large specimen measures 1'5mm. Some, if not all, of these surface-
spicules appear to have had a fifth ray, which penetrated the dermal layer at right
angles to the surface. The wall-plaits of the interior are -75 mm. in width; the
spicules of the mesh are relatively small, and the distance from node to node is
*25 mm.
The specimens are preserved in chalk; the spicules have been dissolved, and only
empty casts or infiltrations of iron peroxide remain.
Distribution. Upper Chalk: Kent and South of England (coll. Mantell and
Toulm. Smith).
Genus CINCLIDERMA, Hinde, gen. nov.
Sponges inverted conical or funnel-shaped. The exterior surface consists of a
very smooth dermal layer formed by large cross-shaped spicules, disposed in such a
128 SILICEOUS SPONGES.
regular manner as to leave quadrate or oblong interspaces like lattice-work. An
extremely delicate tissue, composed of much smaller and more slender spicules,
disposed without any regularity, fills the spaces formed by the regular framework
of the larger spicules, the whole forming an apparently compactly interwoven dermal
layer.
The interior of the funnel formed by this dermal layer is occupied by anastomosing
meandriform lamin or thin walls of spicular tissue similar to those which characterize
the genus Plocoscyphia. The spicular tissue is of a very irregular character, and the
spicular nodes are solid.
In this genus the features which distinguish the families of the Staurodermide and
the Meandrospongide appear to be present: the large cross-shaped spicules of the
dermal layer are essentially similar to those of the former family, and the disposition
of the internal spicular mesh is the same as in the latter family.
There is a certain resemblance between the markings on'the outer surface of the
small conical sponges of the genus Hubrochus, Sollas, and the present genus ; but so
little is known both of the exterior and interior structure of Hubrochus, owing to the
imperfect preservation of the specimens, that it is impossible to institute a comparison
between these genera. The spicular structure both of the dermal layer and of the
interior wall of Cincliderma is, however, altogether different from the magnified
representation of these structures in Eubrochus as given by Sollas*.
CINCLIDERMA QUADRATUM, Hinde, n.sp. (Plate XXVIII. figs. 1, 1a, 16, le, 1d.)
The broken-up fragments of this species appear to have formed part of a funnel-
shaped specimen, 46 mm. in width at its summit. The dermal layer is extremely
thin, and formed by a single layer of spicules; the rectangular spaces, bounded by
the larger spicules, vary in size between ‘75 mm. and 18mm. The rays of the large
spicules overlap each other, and they all possess a fifth ray extending into the wall.
In a few instances the smaller spicules of the interspaces of the lattice-work are
disposed so as to divide them further into smaller quadrate spaces; but, as a general
rule, these smaller spicules are disposed indiscriminately over each other. The
anastomosing wall-plaits of the interior of the sponge are about 1:8 mm. in thick-
ness ; the spicules of the mesh are relatively large, and very irregular in form.
The sponge is preserved in chalk; and the siliceous spicular structure has been
either replaced by iron peroxide or it has been dissolved, leaving empty moulds in
the chalk.
Distribution. Upper Chalk: South of England (coll. 7. Smith).
* Geological Magazine, Dec. 2, vol. iii. pl. 14. figs. 3, 4, 5.
EUBROCHTIS.—PROTOSPONGIA. 129
Genus EUBROCHUS, Solas, 1876.
Evusrocuus cuausus, Sollas.
1876. Eubrochus clausus, Sollas, Geol. Mag. vol. ili. n. s. p. 398, t. 14.
1877. Eubrochus , Zitt. Studien, I Ab. p. 46.
Small, inverted conical sponges, with truncated or rounded summits. The lateral
surfaces are covered with continuous fine linear grooves, which decussate with each
other in such a manner as to form small quadrate or oblong interspaces, the sides of
which are about ‘5 mm. in length. The summit of the sponge is also covered with
a similar lattice-work, but the lines forming it do not decussate as on the sides, but
are disposed at right angles to each other. The characters of the interior structure
of the sponge are too imperfectly preserved for satisfactory determination. Prof.
Sollas has figured fragments of spicular mesh (Joc. cit. figs. 4, 5) with which, he
states, the interior of the sponge was filled; but after a careful examination of the
type specimens in the collections of the Woodwardian Museum at Cambridge and of
the Jermyn-Street Museum, I could only detect in the sections of the interior a few
broken-up fragments of hexactinellid spicules, which were altogether insufficient to
give me a clue to the original structure of the interior skeleton.
Distribution. Cambridge Green Sand: Cambridge (coll. Tracy).
Genus PROTOSPONGIA, Salter, 1864.
PROTOSPONGIA FENESTRATA, Salter. (Plate XXVIII. fig. 2.)
1864. Protospongia fenestrata, Salt. Quart. Journ. Geol. Soe. vol. xx. p. 238, t. 13. f. 12a, b.
1871. Protospongia fenestrata, Hicks, Quart. Journ. Geol. Soc. vol. xxvii. t. 16. f. 20.
1877. Protospongia fenestrata, Zitt. Studien, I Ab. p. 45; Neues Jahrbuch, p. 354.
1880. Protospongia fenestrata, Ferd. Reemer, Leth. Geogn. 1 Th. p. 316, f. 59.
1880. Protospongia fenestrata, Sollas, Quart. Journ. Geol. Soe. vol. xxxvi. p. 364, f. 1.
Form of sponge unknown; the portions preserved consist either of detached
spicules or fragments of a delicate spicular framework formed of a single layer of
four-rayed spicules of various dimensions. The larger spicules are arranged so as to
form regular squares, which are divided by smaller spicules into smaller squares, and
‘these are again subdivided; so that the surface of a fairly complete specimen
resembles minute lattice-work. It is probable that a delicate spicular membrane
connected the entire framework together, as there is in many specimens a thin film
of iron pyrites between the interspaces of the lattice-work ; and as the larger spicules
are now in the condition of iron pyrites, this film of the same material may represent
a spicular membrane whose minute rays have been indistinguishably merged
together.
The not infrequent occurrence of portions of the spicular framework with the
S
150 SILICEOUS SPONGES.
spicules in their proper relative positions, is strong evidence .that they were held
together by a common membrane ; and the presence of such a membrane in sponges
with a similar lattice-like framework of large spicules, as in Cincliderma quadratuin
for example, also points to the same conclusion.
Whether this exterior spicular framework constituted the entire sponge is an open
question ; at present it is the only structure which has been discovered.
Prof. Zittel* is of the opinion that this and the following genus will probably
constitute a new family; but as nothing is at present known of the interior cha-
racters of these sponges, I prefer to place them provisionally in the family of the
Staurodermid on account of their resemblance to the dermal layer of some of the
sponges of this family.
Distribution. Menevian: St. David’s, South Wales. Lower Lingula Beds: Tyddyn-
gwladis. Upper Mawddach: North Wales. Also in black shales of Cambrian age
in Norway and Sweden.
Genus DICTYOPHYTON, Hail, 1863.
DicryoPHYTON TUBEROSUM, Conrad, sp. (Plate XXVIII. fig. 3.)
1842. Hydnoceras tuberosum, Conrad, Journ. Ac. Nat. Sci. Philad. vol. viii. Ist ser. p. 267,
togl Genta:
1863. Dictyophyton tuberosum, Hall, 16th Report of State Cab. New York, p. 90, t. 3. f. 1.
1879. Dictyophyton, sp., Schimper, Handbuch der Pal. 2 Bd. 1 Lief. p. 69.
1880. Dictyophyton tuberosum, Ferd. Roemer, Leth. Geogn. 1 Th. p. 127, f. 3.
1882. Dictyophyton, Zitt. Neues Jahrbuch, I Bd. p. 204.
Sponges inverted conical or elongated funnel-shaped, apparently subangular in
horizontal section when uncompressed, with projecting nodes disposed in horizontal
and vertical rows. Neither the basal extremity nor the summit has been preserved.
A fairly large, though incomplete, specimen is 190 mm. in length by 70 in width.
The surface of the sponge is divided into quadrate and oblong spaces by horizontal
and vertical grooves or raised lines, of varying depth and strength, crossing each
other at right angles. The stouter lines and grooves form the larger squares, the
sides of which are from 4 to 6 mm. in length, and these are divided and subdivided
by finer lines precisely in the same manner as in Protospongia. No structure can be
detected in the interior of the funnel.
Nothing beyond the outer form and quadrate impressions of the outer surface, in
a matrix of micaceous sandstone, remains of this sponge. The regular framework of
the surface, however, so closely resembles that of Protospongia that there is every
reason to suppose that it has been formed by the casts of spicules of different dimen-
sions, disposed so as to form a regular lattice-work, as in that genus. It is also
* Neues Jahrbuch fiir Mineralogie &c., 2 Bd., 1882, p. 204.
DICTYOPHYTON. all
probable that this framework was held together by a delicate spicular membrane,
for crushed examples occur in which portions of the framework are frequently
displaced without having the symmetrical arrangement destroyed; but in some
instances the spicules are broken up and mingled together, in the same manner as
in certain specimens of Protospongia.
This form was originally regarded by Conrad as a subgenus of Orthoceras; it was
afterwards referred by Hall to marine Alge. Still later Prof. Schimper expressed
doubts of its being a plant, and stated that the structure much nearer resembled the
skeleton of siliceous sponges. Prof. Ferd. Reemer rightly compared the structure
with that of Zetragonis Danbyi, M‘Coy, with which it is undoubtedly congeneric.
After this Mr. R. P. Whitfield *, without being aware of Schimper’s and Reemer’s
comparisons of the genus to sponges and allied forms, or at all events without
mentioning the fact, refers Dictyophyton and some other genera to sponges like the
recent Luplectella.
Hall enumerates eight other species of Dictyophyton, based on differences of outer
form; it is probable that they may be all included in the present species. The
Tetragonis Eifeliensis, Reemer, very closely resembles the present form, and there
can be hardly a doubt of its belonging to the same genus.
Distribution. Chemung Group, Upper Devonian: Cohocton, Steuben County,
Western New York.
Dicryopuyton Dansyi, M*‘Coy, sp.
1855. Tetragonis Danbyi, M‘Coy, Brit. Pal. Foss. p. 62, pl. 1D. £7 & 8.
1880. Tetragonis Danbyi, Ferd. Remer, Leth. Geogn. 1 Th. p. 304.
Sponge subconical or subovate, imperfect both at the basal end and at the summit.
Length about 25 mm. The surface is divided into larger and smaller squares by
apparently continuous lines, in the same manner as in the preceding species, but the
sides of the larger squares are not more than 1:5 to 2 mm. in length, thus very much
smaller than in D. tuberosum.
Both M‘Coy and Reemer have referred this form to the genus Tetragonis, Kichwald ;
but the typical species of this genus, 7. Murchisonii, Kichw. +t, though presenting a
superficial resemblance to the present form, possesses a structure of a quite different
character. From an examination of undoubted forms of Tetragonis from the Silurian
of Gotland, I believe that it is a sponge, and that it is very closely allied to Lscha-
dites, Murch.
Dictyophyton Danbyi differs from D. tuberosum, Conrad, and D. Eifeliensis, Remer,
in its smaller size and the slighter character of the spicular framework. The
only specimen which I have seen is M‘Coy’s figured type in the Woodwardian
* American Journal of Science, vol. xxii. p. 53.
+ Urwelt Russlands, Heft 2, p. 81, t. 3. f. 18.
132 SILICEOUS SPONGES.
Museum at Cambridge. The specimen is preserved in a matrix of micaceous
sandstone.
Distribution. Silurian. Upper Ludlow: Brigsteer, Kendal, Westmoreland.
Genus PLECTODERMA, Hinde, n.g.
Outer form of sponge unknown; the only structures preserved are fragments of a
thin spicular membrane, consisting of cruciform and five-rayed spicules of relatively
large size, which are grouped together into continuous rows by the apposition and
overlapping of several of their vertical axes, whilst the rays of their lateral axes
extend singly on either side of the vertical rows and overlap those from the adjoining
rows, thus forming an irregular framework. Smaller spicules are present between
the interspaces of the larger forms, but they do not appear to be arranged in definite
order.
This genus is closely related to Protospongia, Salter, and Dictyophyton, Hall, but
it may be readily distinguished from either of these genera by the less regular
character of the spicular framework and the absence of definite larger and smaller
squares. From Protospongia it is further distinguished by the disposition of the
spicules of the vertical rows in small bundles instead of in a single series. Whether
the spicules in Dictyophyton were arranged singly or in bundles is not known, as no
spicular structure has been retained.
PLECTODERMA scituLUM, Hinde, n.sp. (Plate XX XI. figs. 1, 1 a, 16.)
The examples of the species occur on the fractured surfaces of a soft micaceous
shaly rock. ‘The vertical rows are about 7°5 mm. apart from each other; they
consist of a variable number, generally from five to ten, of the spicular axes in pretty
close juxtaposition. ‘The spicules vary in size, from slender forms not more than
‘09 mm. in thickness, to large spicules with rays 35 mm. in thickness and 6°90 mm.
in length. The spicules are now mostly represented by empty moulds, and at first
sight appear as so many monaxial spicules crossing each other at right angles; but in
places where they are less thickly grouped together, the four rays, springing from a
common centre, can be clearly seen, and at the point of junction there is often a
small circular hole, indicating a fifth ray, extending inwards at right angles to the
other four. The spicular rays are straight, curved, or occasionally wavy ; they usually
taper from the central node. In some instances fragments of the original siliceous
spicules yet remain, but they are so far decayed as readily to fall into minute
pieces. In some cases the central canal can be seen in these fragments. Whether
the framework of larger spicules was held in position by a continuous membrane of
smaller spicules, as in the dermal layer of the Cretaceous genus Cincliderma, cannot
be so readily determined ; but traces of smaller spicules, irregularly disposed between
PLOCOSCYPHIA. 135
the larger rays, can be occasionally seen, so that there is great probability that a
continuous membrane was originally present.
Distribution. Silurian strata of Upper Ludlow age: Wetherlawlinn, Pentland
Hills, near Edinburgh. The specimens belong to =the collection of the Geological
Survey of Scotland, and were discovered by Mr. James Bennie.
Family MEANDROSPONGID 2, Zittel.
Genus PLOCOSCYPHIA, Reuss, 1846.
PLocoscyPHIA FENESTRATA, 7. Smith, sp. (Plate XXVIII. figs. 4, 4a, 45, 4c.)
1848. Brachiolites fenestratus, Toulm. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 367,
tel Gteae
Sponges subcylindrical, hemispherical, or irregular in form, oftentimes growing in
close contact with each other, so as to form masses of considerable size. The outer
surface is composed of numerous anastomosing, subcylindrical, open tubes, from 4 to
8 mm. in diameter. The central portion of the sponge, as seen in a transverse section,
appears to consist of open anastomosing folds of the sponge-wall, which, when simple,
are about 2 mm. in thickness; but the folds sometimes coalesce laterally, and are
consequently of increased thickness.
The spicular structure of the interior of the wall-plaits is generally regular; the
spicules form a rectangular oblong mesh with elliptical interspaces. The distance
between the nodes varies from *3 to’4 mm. The spicular arms are robust, smooth,
or with blunt spines, and about ‘08 mm. in thickness. The dermal layer is composed
of an irregular spicular network, with minute circular or ovate pores; the nodes of
the spicules are solid, whilst those of the interior of the wall are octahedral or
Jantern-like.
This is a well-marked species, and readily recognizable by the size and disposition
of the tubes of the exterior surface. In the numerous specimens in the Museum
from T. Smith’s collection, and labelled by him, I have not found one which corre-
sponds with the figure which he has given of this form. ‘This figure appears to me
to be diagrammatic merely, and altogether misleading.
Distribution. Upper Green Sand: Dover and Folkestone. Chalk Marl: Ventnor,
Isle of Wight; Norton Bavant, Wilts.
PLocoscyPHiA LABROSA, Toulm. Smith, sp. (Plate X XIX. fig. 2.)
1848. Brachiolites labrosus, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 368,
t. 16. £. 4.
1878. Antrispongia dilabyrinthica, Quenst. Petref. Bd. 5, p. 474, t. 137. f. 24.
154 SILICEOUS SPONGES.
1842. Compare Spongus meandrinoides, Leymerie, Mém. Géol. Soc. de France, vol. v. p. 1,
itp Us 15 2s
Sponges growing in subspherical, ovate, and irregular masses of very various
dimensions. A small specimen is 90 mm. in diameter, and a large example measures
190 mm. across. The mass is composed of anastomosing folds of wall, from 25 to
3 mm. in thickness, which in some specimens form wide anastomosing tubes, and in
other examples open meandriform folds. There are numerous intermediate examples
between the tubular forms and those with open folds, and frequently the two
characters are met with in the same specimen. ‘The surface of the wall is perforated
on both sides by the apertures of blind canals, from ‘5 to "75 mm. in width.
The dermal layer is composed of a minute spicular network with circular or ovate
pores; the spicular mesh of the interior of the wall is irregular in its disposition ;
the distance from node to node is ‘25 mm.; the lantern character of the nodes is
distinctly shown.
This species differs from the preceding in the disposition of the folds of the wall,
and in the closer and less regular character of its interior mesh. Mr. F. G. H.
Price * has named this form Plocoscyphia meandrina, Leymerie, which I presume is
the same as the Spongus meandrinoides of this author. Leymerie’s figure, however,
appears to me to represent a form differing from Smith’s in the thinner walls and
closer arranged folds. As the French typical specimen is only a cast in iron pyrites,
no other comparison beyond that of outer form and the thickness of the wall-folds is
practicable, and in both these features it differs from P. labrosa.
This species is extremely abundant in certain zones of the Upper Green Sand and
Chalk Marl.
Distribution. Upper Green Sand and Chalk Marl: between Folkestone and Dover.
PLocoscyPHIA PERTUSA, Geinitz.
1843. Tragos pertusum, Geinitz, Die Verstemerungen von Kieslingswalda, p. 19, t. 6. f.18.
1871. Plocoscyphia pertusa, Gein. Palzont. Bd. 20, p. 26, t. 2. f. 5a, 6, and t. 3. f. La, 6.
1878. Plocoscyphia pertusa, Zitt. Handb. der Pal. p. 181, f. 96.
1883. Plocoscyphia pertusa, Keeping, Fossils of Upware &e. p. 145, t. 8. f. la, 1b.
Sponge irregular in form, built up of convoluted walls, about 1°75 mm. in thick-
ness, disposed so as to leave wide channels or anastomosing cylindrical tubes, about
8mm. in diameter. The exterior surface is rough and uneven, and is furnished with
numerous irregularly distributed circular or elongate canal-apertures.
The dermal layer is composed of a stout spicular mesh; the spicular arms are
flattened and spinous, and the nodes solid; the interspaces of the mesh are mostly
circular. The interior spicular mesh has subquadrate interspaces. ‘The nodes are
lantern-like ; the distance from node to node is *25 mm.
* Quart. Journ. Geol. Soc. 1877, vol. xxxiil. p. 442.
PLOCOSCYPHIA. 155
The above description is taken from a beautifully preserved fragment of this
species presented to the Museum by Prof. Zittel.
According to Mr. Keeping this species occurs in the Neocomian beds at Brickhill ;
but the figure given is insufficient for me to determine whether the specimen is
rightly referred to this species.
Distribution. Plainer-Kalk (Upper Green Sand): Bannewitz, Saxony. Neocomian:
Brickhill, Bedfordshire (Aeepinq).
PLOCOSCYPHIA RETICULATA, Hinde, n. sp. (Plate X XIX. figs. 5, 3a, 30.)
Sponge growing in irregularly shaped masses; the convoluted walls, 3°5 mm. in
thickness, are disposed so as to form at the surface subcylindrical tubes from 11 to
16mm. in diameter. The outer surface of the wall has a reticulate aspect from
numerous circular or ovate canal-apertures, ‘75 to 1 mm. in width, and about their
own diameters apart. The canals are blind, and apparently are oftentimes curved in
their course in the wall.
The dermal layer is composed of robust spicules with flattened arms and compact
nodes; in places small spines project outwards; the interspaces are circular. ‘The
“spicular mesh of the interior of the wall is irregular; the spicular nodes are ‘22 mm.
apart; the mesh is subquadrate or circular.
This species approaches in general appearance very closely to P. pertusa, Gein.,
but the walls are much thicker and the canal-apertures larger. From P. labrosa,
T. Smith, it may be distinguished by its thicker walls and the disposition of the
canal-apertures.
Distribution. Upper Green Sand: Eastbourne. Chloritic Marl: Rocken End, Isle
of Wight.
PLOCOSCYPHIA SUBRUTA, Quenst. sp.
1878. Gyrispongia subruta, Quenst. Petref. Bd. 5, p. 480, t. 138. f. 2-6.
Sponges subcylindrical or club-shaped, with rounded summits. An apparently
average specimen is 110 mm. in length by 38 in width. The outer surface has
numerous circular or ovate apertures from 4 to 10 mm. in width.
The sponge is composed of convolute walls, 2mm. in thickness; the outer surface
consists of a spicular network, with definite canal-apertures ‘75 mm. in width; the
spicular arms are smooth, and the nodes compact. ‘The spicular mesh of the interior
of the wall has lantern nodes, the spicular arms are 2 mm. in length.
Quenstedt states* that Achillewm formosum, Reuss, probably belongs to this
species; but though there is a great resemblance in outer form, the spicular mesh
of A. formosum is composed of spicules with solid nodes, entirely different from those
of Quenstedt’s form.
Distribution. Chalk Marl: Rocken End, Isle of Wight.
* Bohm. Kreide, p. 79, t. 43. f. 7.
156 SILICEOUS SPONGES.
PiocoscypHia convotuTa, Toulm. Smith, sp.
1848. Brachiolites convolutus, T. Smith, pars, Ann. & Mag. Nat. Hist. vol. i. p. 355, t. 15.
f. 5, non fig. N. p. 357.
Sponge growing in small masses of irregular form, composed of delicate convoluted
walls about 1 mm. in thickness. Traces of a minutely perforated dermal layer are
present in one example; the interior of the wall is built of an irregular slender
spicular mesh; the distance between the nodes is°25mm. The spicules have been
replaced by iron peroxide, and the character of the nodes cannot be determined.
Toulmin Smith states that “the structure of the simple membrane (or wall) in
this species seems coarse—that is, the squares (of the mesh) are larger than in any
other species of the Ventriculitide.” Such, however, is not the case in the specimen
which he has figured, and which I have taken as the type of the species. ‘The inter-
spaces of the mesh in this example are relatively small. In the original specimen
there are no traces of the minute stem and root represented in the figure given by
Smith.
Distribution. Upper Chalk: South of England (coll. T. Smith).
ProcoscypHta FLEXUOSA, Mantell, sp. (Plate XXIX. fig. 4.)
1822. Choanites fleruosus, Mant. Foss. of the South Downs, p. 179, t. 15. f. 1.
1848. Brachiolites convolutus, T. Smith, pars, Ann. & Mag. Nat. Hist. vol. i. p. 357, fig. N.
1878. Gyrispongia labyrinthica, Quenst. Petref. Bd. 5, p. 485, t. 138. f. 12, 13.
Sponges inverted conical, or irregular in form, in some instances supported on a
cylindrical stem with divergent rootlets at its termination. ‘The body of the sponge
is composed of convolute anastomosing walls from 1:25 to 1-7 mm. in thickness. The
surface of the walls has numerous circular canal-apertures, ‘6 mm. in width. The
spicular mesh of the interior is somewhat irregular in disposition; the distance
between the octahedral nodes, or the diameter of the interspaces of the mesh, varies
between ‘35 and -4 mm.
The type of this species, now in the Museum, is the lower portion of a specimen
preserved in flint, showing a section of the convolute walls and traces of the spicular
mesh. Numerous other examples, both in flint and in chalk, exhibit walls of the
same thickness, and a spicular mesh with interspaces of the same size as those of the
type. In none of the specimens has the outer form been completely preserved ; so
that the only characters available for specific determination are the thickness of the
walls and the dimensions of the mesh interspaces. ‘The sponges figured by Quenstedt
under the name of Gyrispongia labyrinthica appear to correspond with the present
species; but I am unable to determine whether the Plocoscyphia labyrinthica of
Reuss* and F. Reemerf belong to it.
Distribution. Upper Chalk: South of England (coll. Mantell, T. Smith).
* Bohm. Kreide, II Ab. p. 77, t. 18. f. 10. + Geol. Oberschles. p. 309, t. 33. f. 7.
PLOCOSCYPHIA. 137
PLOCOSCYPHIA LABYRINTHICA, Mant. sp. (Plate XXIX. fig. 5.)
1822. Spongus labyrinthicus, Mant. Foss. of the South Downs, p. 165, t. 15. f. 7.
Sponges irregular in form, composed of anastomosing walls arranged in open
convolutions. The walls are from 2°5 to 3 mm. in thickness; their outer surfaces
are perforated by circular canal-apertures about ‘7 mm. in width. The distance
between the nodes of the interior mesh varies from ‘33 to ‘4 mm.
The examples of this sponge are usually preserved in hemispherical nedules of
flint, which exhibit on their flattened upper surfaces sections of the convolute walls.
This species is distinguished from P. flexuosa by the greater thickness of the walls
and the more open character of the convolutions, whilst the size of the spicular mesh
appears to be similar.
Distribution. Upper Chalk: Bridgwick; Offham, Kent (coll. Mantell).
PLocoscyPHIA VAGANS, Hinde, n. sp. (Plate XXIX. figs. 6, 6 a.)
Sponges irregular in form, with convoluted walls about 1:5 mm. in thickness. The
spicular mesh is composed of large spicules with lantern nodes; the distance between
the nodes varies between °625 and °8 mm.
The examples of this species are all preserved in chalk, and the spicular mesh is
now changed to iron peroxide or shown by empty moulds. ‘The species is readily
distinguished from those previously described by the unusually large size of the
spicules of the interior mesh of the wall.
Distribution. Upper Chalk: Maidstone, Kent, and other localities in the South of
England (coll. 7. Smith).
PLocoscyPHIA FOLIACEA, Toulm. Smith, sp.
1848. Brachiolites foliaceus, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 364, t. 16. f. 1.
Sponges club-shaped, usually curved in the lower portion, with a few small
divergent rootlets at the base, the summits rounded. The outer surface smooth,
with irregularly disposed rounded elevations and depressions. A fairly large example
measures 155 mm. in length, and 48 in width at the summit.
The body of the sponge is composed of anastomosing walls from 1 mm. to 1°75 in
thickness, disposed in open convolutions.
The examples of this species are preserved in chalk, and the spicular structure
is only indicated by markings in iron peroxide, in which the character of the mesh
has been obliterated. Its form and the features of the outer surface distinguish it
from other species of this genus. Toulmin Smith’s figure of this form is partly a
restoration.
Distribution. Upper Chalk: South of England (coll. T. Smith).
138 SILICEOUS SPONGES.
PLOcoScYPHIA ELEGANS, 7’. Smith, sp.
1848. Brachiolites elegans, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 355, t. 15. f. 4.
1878. Brachiolites elegans, Quenst. Petref. Bd. 5, p. 490, t. 138. f. 15.
Sponges with a subspherical body, supported on a slender cylindrical stem, termi-
nating in divergent rootlets. From the summit of the body an upright cylindrical
tube extends. The body of an average specimen is 47 mm. in height by 32 in width ;
the cylindrical tube measures 46 mm. in length by 23 in width; and the stem is
43 mm. long. The outer surface of the body presents rounded sinuous elevations
and depressions ; that of the tube is smooth and even.
The body is composed of anastomosing folds of wall about 1 mm. in thickness; the
tube appears to be hollow, and formed of a simple extension upwards of the body-
walls. ‘The dermal layer appears to have been furnished with minute circular canal-
apertures, the spicular mesh is regular, and the distance between the nodes is*35 mm.
The characters of the spicular nodes have been obliterated.
The peculiar form of this species readily distinguishes it from any other of the
genus. T. Smith’s figure has been drawn from two separate specimens, and is, to a
certain extent, a restoration of the originals.
Distribution. Upper Chalk: Brighton, Sussex ; Hollingbourne, Birling, near Maid-
stone, Kent (coll. T. Smith).
Genus TREMABOLITES, Ztt. 1877.
TREMABOLITES PERFORATUS, 7. Smith, sp.
1848. Cephalites perforatus, T. Smith, Ann. & Mag. Nat. Hist. vol. i. t. 15. f. 2.
Sponges subcylindrical or club-shaped, with rounded summits. ‘The sponge is
composed of anastomosing, convoluted walls about 1 mm. in thickness. The summit
is covered with an apparently delicate surface-membrane, which is perforated by
several circular or oval apertures from 3 to 7 mm. in width. ‘The spicular mesh of
the walls is irregular; the nodes are octahedral, and from -4 to ‘5 mm. apart.
Iam unable to determine the extent of the resemblance of the interior structure
of this species to that of Tremabolites (Manon) megastoma, F. A. Reem.*, but its
outer form, at all events, considerably differs. The forms figured by Quenstedtf
under the name of Cephalites polystoma differ from the present species in the
extension of the surface-membrane over the sides as well as the summit of the
sponge.
Distribution. Upper Chalk: South of England (coll. T. Sinith).
® Nordd. Kreide, p. 3, t. 1. f. 9. t Petref. Bd. 5, p. 503, t. 139. f. 8-10.
ETHERIDGIA.—TOULMINIA. 139
Genus ETHERIDGIA, Tate, 1865.
ETHERIDGIA MIRABILIS, Tate.
1865. Etheridgia mirabilis, Tate, Quart. Journ. Geol. Soc. vol. xxi. p. 48, t. 5. £. 4a, 46.
1877. Etheridgia mirabilis, Zitt. Studien, I Ab. p. 56; Neues Jahrbuch, p. 367.
Sponge conical with truncated summit, the base depressed, convex, with short,
projecting cylindrical root-processes. At the summit is a circular aperture 15mm.
wide. ‘The sides of the cone are provided with circular or elliptical apertures about
4mm. in diameter. The type specimen is 33mm. in height and 40 mm. in width.
A compact dermal membrane extended over the base and sides. The interior
structure has been completely obliterated. The only specimens known are in the
Jermyn-Street Museum.
Distribution. Spongarian zone of the Chloritic Chalk (Zate)=Upper Chalk:
Whitehead, near Belfast.
Genus TOULMINIA, Zttel, 1877.
TovuLMINIA OBLIQUA, Hinde, n. sp. (Plate XXIX. figs. 7, 7a, 76.)
Sponge funnel- or cup-shaped. The only example is 48 mm. in height, and 50 mm.
wide at the summit. The cloacal cavity is 20 mm. in width near the upper margin ;
it appears to be without any definite wall. The wall is composed of irregularly
anastomosing folds about 1 mm, in thickness. The dermal membrane of the summit
of the walls has nearly entirely been removed; the spicular mesh of the wall-plait is
irregular; the distance between the nodes is ‘25 mm.
Distribution. Upper Chalk: South of England (coll. T. Smith).
Toutminia Jurassica, Hinde, n. sp. (Plate XXX. figs. 1, 1a, 10.)
Sponge inverted conical in form, with an oblique summit. The single specimen
is 63 mm. in height, and 46 in width near the summit. The cloaca appears to be
funnel-shaped, and bounded by a definite wall; it is 23 mm. in width at the upper
margin. The sponge is composed of anastomosing convolute wall-plaits, 1 mm. in
thickness. The spicular mesh of the wall, as seen in a transverse section, is regular
in disposition; the nodes are octahedral, and ‘25 mm. apart from each other. No
dermal membrane has been preserved, probably owing to the weathering of the
surface.
This species differs from the preceding in the well-defined character of the cloacal
cavity as well as in the disposition of the wall-plaits, as seen in transverse sections.
This form is at present the only representative of the family of the Meandro-
spongide from the Jurassic system.
Distribution. Upper Jura: Randen, Switzerland.
T2
140 SILICEOUS SPONGES.
Genus CAMEROSPONGIA, D’Orbigny, 1847.
CAMEROSPONGIA SUBROTUNDA, Mant. sp. (Plate XXX. fig. 3.)
1822. Choanites subrotundus, Mant. Foss. of the South Downs, p. 179, t. 15. f. 2.
1831. Choanites subrotundus, Benett, Cat. Org. Rem. Wilts. t. 16. f. 1.
1848. Cephalites constrictus, Toulm. Smith, Ann. & Mag. Nat. Hist. vol.i. p. 292, t. 15. f. 1.
Sponges depressed globate in form, growing either singly or in small groups, and
not infrequently in a single series of two or three individuals closely attached
together. The sponges appear to have been attached by a few divergent rootlets
given off laterally; in the compound examples only the first sponge in the series
appears to possess the rootlets. Separate individuals vary between 15 and 35 mm.
in breadth, and 6 and 15 mm. in height. ‘The upper surface is slightly rounded,
oftentimes with a gentle depression towards the centre. ‘The aperture is circular or
oval, from 3:5 to 9 mm. in width, with an elevated margin, encircled by a shallow
furrow.
Nearly the entire upper surface and sides are covered by the smooth siliceous
membrane. The under surface is uneven, with a few circular apertures between
the ridges formed by the folded walls. The convoluted wall-plaits of the interior
are slender, and composed of a spicular mesh with lantern nodes; the nodes are
*375 mm. apart.
The subglobate form and the social mode of growth distinguish this species from
others of the genus.
Distribution. Upper Chalk: Lewes, Sussex ; Charing, Kent (coll. Mantell, T. Smith).
Also from the Spongarian zone of the Chloritic Chalk (Zate) = Upper Chalk: White-
head, near Belfast (Jermyn-Street Museum).
Camerosponata Capitata, Toulm. Smith, sp.
1848. Cephalites capitatus, T. Smith, Ann. & Mag. Nat. Hist. vol. i. p. 288, t. 14. f. 11.
1878. Cephalites capitatus, Quenst. Petref. Bd. 5, p. 498.
Sponges inverted conical in form, supported on a slender cylindrical stem, with
divergent rootlets. The summit is gently convex, the margin is subacute. From
the summit the body rapidly tapers to the base. The typical specimen is 21 mm. in
height by 31 mm. in width; the stem is 11 mm. in length. The summit-aperture is
circular, 11:5 mm. in width, its margin is thickened, and slightly elevated.
The smooth enveloping membrane only covers the summit; the sides exhibit a
reticulate aspect, formed by wrinkled ridges with circular or irregular interspaces.
A subcylindrical or funnel-shaped cloacal cavity extends from the summit to the
base, though it does not appear to have possessed a definite boundary wall. The
interior wall-plaits are loosely convolute; they are about ‘1 mm. thick. The nodes
of the spicular mesh are ‘45 mm. apart.
CAMEROSPONGIA. 141
This species has a great resemblance in outer form to C. (Scyphia) fungiformis,
Goldf.* ; but the summit is less elevated, and the margins do not project over the
sides as in Goldfuss’s form. ‘The spicular structure of the mesh is also much larger
than in C. fungiformis. The description and figures of C. (Manon) monastoma,
F. A. Reemerf, are too indefinite to permit of a comparison with the present species.
The specimen figured by Reemer cannot be distinguished from C. fungiformis, Goldf.
T. Smith’s figure is not an altogether accurate representation of the original, which
has a slender stem, and the lateral walls are much closer arranged than represented.
Distribution. Upper Chalk: South of England (coll. Mantell, T. Smith).
CAMEROSPONGIA FUNGIFORMIS, Goldf. sp.
1826-1833. Scyphia fungiformis, Goldf. Petref. 1 Th. p. 218, t. 65. f. 4.
1877. Camerospongia fungiformis, Zitt. Studien, I Ab. p. 57; Neues Jahrbuch, p. 368.
1878. Camerospongia fungiformis, Zitt. Handb. der Pal. p. 182, f. 97.
The Museum only possesses microscopic fragments of the spicular mesh of this
species. ‘The disposition of the mesh appears to be regular; the octahedral nodes
are ‘35 mm. apart: thus the mesh-interspaces are smaller than in C. capitata.
Distribution. Upper Chalk: Vordorf, Brunswick.
CAMEROSPONGIA TURBINATA, Giedel, sp.
1851. Ptychotrochus turbinatus, Giebel, Jahresbericht des naturhist. Vereins in Halle, p. 53,
fe Ph uly Hp
Microscopic fragments of the spicular mesh.
Distribution. Upper Chalk: Galgenburg, Goslar (coll. Zitt.).
CAMEROSPONGIA CAMPANULATA, 7. Smith, sp.
1848. Cephalites campanulatus, T. Smith, pars, Ann. & Mag. Nat. Hist. vol. i. p. 289, t. 14.
£. 12; non £13:
1877. Camerospongia campanulata, Zitt. Studien, I Ab. p. 57; Neues Jahrbuch, p. 368.
1878. Cephalites campanulatus, Quenst. Petref. Bd. 5, p. 498.
Sponges subglobate in form, supported on a short cylindrical stem with divergent
rootlets. The summit is dome-shaped, with slight elevations and depressions; the
central aperture is circular, and about 22 mm. in width. A typical specimen is
42 mm. in height and 54 in width, whilst the stem is only 8 mm. in length.
A wide subcylindrical cloacal cavity, apparently destitute of a boundary wall,
extends from the summit to the base. The smooth membrane covers the summit
and the sides to nearly two-thirds the height of the sponge. The under surface
exhibits the reticulations of the wall-plaits with irregular interspaces. The convolute
* Petrefacten, 1 Th. p. 218, t. 65. f. 4. + Nordd. Kreide, p. 2, t. 1. f. 8.
142 SILICEOUS SPONGES.
walls of the interior are about 1 mm. in thickness ; the octahedral nodes of the mesh
are ‘38 mm. apart.
The larger size and the dome-shaped summit distinguish this species from C. capi-
tata; the spicular mesh is also smaller. The figures H and I of T. Smith (Joe. cit.
p. 291), representing the disposition of the folds of the wall, are purely diagrammatic.
In the typical examples the wall-plaits do not exhibit the regularly arranged folds
depicted in these figures.
Distribution. Upper Chalk: South of England (coll. T. Smith).
CaMEROSPONGIA APERTA, Hinde, n.sp. (Plate XXX. figs. 2, 2a, 26.)
1848. Cephalites campanulatus, T. Smith, pars, Ann. & Mag. Nat. Hist. vol. i. t. 14. f. 3.
Sponges depressed, cup-shaped, supported on a short stem with diverging rootlets.
The interior exhibits a funnel-shaped cloacal cavity, bounded by a definite wall; the
margins are subacute. The lateral surfaces are smooth and slightly uneven; the
under surface is formed by uneven ridges with irregular interspaces. A few irregu-
larly convoluted wall-plaits are shown in a vertical section; the spicular mesh has a
regular disposition; the octahedral nodes are ‘25 mm. apart. The spicular mesh of
the walls of the cloaca is of the same character as that forming the interior plaits.
The body of the type specimen is 25 mm. in height by 36 in width.
This species differs from C. campanulata, as defined above, by its generally smaller
size, the widely expanded cloacal aperture, the definite cloacal wall, and the smaller
dimensions of the spicular mesh.
Distribution. Upper Chalk: South of England (coll. 7. Smith).
Genus CYSTISPONGIA, F. A. Remer, 1864.
CYSTISPONGIA BURSA, Quenst. sp.
1852. Cephalites bursa, Quenst. Handbuch d. Petref. p. 670, t. 60. f. 17.
1864. Cystispongia bursa, F. A. Roemer, Paleontographica, vol. xii. p.7, t. 4. f. 7.
1877. Cystispongia bursa, Zitt. Studien, I Ab. p. 57; Neues Jahrbuch, p. 368; Handb. d.
Pal. p. 182, £. 98.
1878. Cephalites bursa, Quenst. Petref. Bd. 5, p. 492, t. 138. f. 17.
Distribution. Obere Planer: near Vienenburg, Prussia (coll. Zitt.).
Family CALLODICTYONIDZ, Zittel.
Genus CALLODICTYON, Zittel, 1877.
CaLopictyon aneustatuM, Hinde, n.sp. (Plate XXX. figs. 4, 4a, 46.)
Sponges elongate, funnel-shaped, very gradually tapering from the summit to the
CALLODICTYON.—POROCHONIA. 148
base. The stem has not been preserved. An imperfect specimen measures 110 mm.
in length, and 40 mm. in width at the summit.
The wall of the sponge, about 2 mm. in thickness, is composed of an extremely
regular spicular mesh with oblong interspaces; the nodes are octahedral, and about
“6 mm. apart in a vertical direction and ‘45 mm. transversely. The outer surface is
formed of a very delicate open network, with circular or ovate apertures, ‘5 mm. in
width, disposed in vertical rows. ‘These apertures open directly into the interspaces
of the mesh, and do not appear to be connected with distinct canals. The interior,
or cloacal surface of the wall, does not appear to possess a modified spicular mesh.
The examples of this species are preserved in chalk, and the siliceous spicular
mesh has been replaced by iron peroxide, so that it is impossible to determine
whether the rays were smooth or spinous.
Cylindrospongia membranacea, Quenst.*, non Reemer, appears to belong to the
genus Callodictyon, and resembles the present form in the disposition of the apertures
of the outer surface ; its walls, however, judging from the figure, are not more than
one third the thickness of those of C. angustatum.
Distribution. Upper Chalk: South of England (coll. Bowerbank).
Genus POROCHONIA, Hinde, n. g.
Sponges funnel-shaped, with thin walls. The outer surface of the wall is formed
by a very delicate membrane, beneath which is a reticulate layer with circular
apertures. Below this layer is the regular spicular mesh of the wall; the nodes of
the six-rayed spicules are octahedral. The inner or cloacal surface of the wall is
formed by a reticulate layer similar to that overlying the spicular mesh. Definite
canals do not seem to be present, the circulation being carried on apparently through
the regular interspaces of the mesh.
I have based this genus on the characters of Ventriculites simplex, T. Smith, the
walls of which differ from those of typical forms of Ventriculites in being smooth
and not arranged in vertical folds. From Callodictyon this genus is characterized by
possessing a delicate surface-membrane (the polyp-skin of T. Smith) in addition to
the usual dermal layer (the underskin of T. Smith). The exterior membrane is only
present in some of the best-preserved specimens; and owing to the replacement of
its original spicular structure by iron peroxide, the minute characters cannot be
determined.
PorocHonta SIMPLEX, 7. Smith, sp. (Plate XXX. figs. 5, 5a, 50.)
1848. Ventriculites simpler, T. Smith, Ann. & Mag. Nat. Hist. 2nd ser. vol. i. p. 204, and
Istiserivolaxx. tac: fal!
® Petref. Bd. 5, p. 468, t. 137. f. 13.
144 SILICEOUS SPONGES.
1851. Spongia Townsendi, Morris, Catalogue, p. 31.
1878. Ventriculites simplex, Quenst. Petref. Bd. 5, p. 470, t. 187. f. 17.
Sponges funnel-shaped, either supported on a cylindrical stem with divergent
rootlets or having rootlets springing from the base of the funnel. Variable in size—
from small forms, 22 mm. in height by 30 in width at the summit, to large examples,
95 mm. in height and 92 in width.
The walls are frequently very compressed ; in the best-preserved specimens they
are from 1:5 to 2mm. in thickness. The exterior layer is smooth, and sometimes
exhibits concentric markings. The dermal layer is perforated with irregularly
disposed circular or oval apertures, about ‘5 mm. in width, and the same distance
apart. The spicular mesh is regularly subquadrate ; the octahedral nodes are -43 mm.
apart. The spicular structure of the stem and rootlets differs from that of the walls,
and consists of a network of elongate spicular fibres.
The examples of this species are abundant; in all cases the spicular structure is
either in the condition of iron peroxide or in hollow casts.
According to T. Smith, Spongus Townsendi, Mant.*, is nothing more than a
V. simplex in flint. Jf such were certainly the case the specific name of Mantell
ought to have been retained; but the type of Mantell, now in the Museum, is so
completely imbedded in flint that its spicular characters cannot be ascertained, and
it might belong to an entirely different form from Smith’s species.
The sponge figured by the late Sir Wyville Thomson, in the ‘ Depths of the Sea’,
as Ventriculites simplex, T. Smith, and from thence copied into other works, has no
resemblance to Smith’s species, though it may possibly be a genuine specimen of
Ventriculites.
Distribution. Upper Chalk: Lewes, Sussex; Charing, Kent (coll. Mantell and
£. Smith).
Genus BECKSIA, Schliiter, 1868.
Becksta S@KELANDI, Schliiter.
1868. Becksia Seekelandi, Schl. Sitzungsb. der nieder. Gesells. im Bonn, p. 93.
1872. Becksia Swkelandi, Schl. Ueber die Spongit.-Baenke des Miunsterlandes, p. 20, t. 1.
f. 5-7.
1877. Becksia Swkelandi, Zitt. Studien, I Ab. p. 58; Neues Jahrbuch, p. 369.
1878. Becksia Swkelandi, Quenst. Petref. Bd. 5, p. 489, t. 138. f. 14.
1878. Becksia Sekelandi, Zitt. Handb. der Pal. 1 Th. p. 183, f. 99.
Distribution. Upper Chalk: Coesfeld, Westphalia.
* Fossils of the South Downs, p. 154, t. 15. f. 19. t+ P. 483, f. 80; p. 484, f. 81.
DIPLODICTYON.—SCLEROKALIA. 145
Genus DIPLODICTYON, Zittel, 1877.
DIPLODICTYON HETEROMORPHUM, Leuss, sp.
1845. Scyphia heteromorpha, Reuss, Bohm. Kreide, p. 74, t. 18. f. 1, 2.
1877. Diplodictyon heteromorphum, Zitt. Studien, I Ab. p. 59. 4
Microscopic fragments of the spicular mesh.
Distribution. Plainer-Kalk: Schilling, Bohemia (coll. Zitt.).
Diptopictyon BayFriELpi, Hinde, n.sp. (Plate XXXI. figs. 2, 2a.)
Sponge compressed, triangular in outline, apparently supported on a short stem.
The walls, 3-8 mm. in thickness, run parallel with each other; their summit-margins
are rounded. On the compressed sides are two or three large ovate apertures. The
only example is 58 mm. in height and 50 mm. in width at the summit.
The outer surface has numerous circular, irregularly disposed apertures, about
1 mm. in width. ‘The thickness and spicular character of the dermal layer cannot
be determined owing to the condition of the specimen. The interior part of the
wall is formed of an apparently regular quadrate mesh; the octahedral nodes are
25 mm. apart.
This species may be distinguished from D. heteromorphum by its different form and
the smaller dimensions of the mesh-interspaces of the interior of the wall.
I have named this form after its discoverer, Mr. T. G. Bayfield of Norwich.
Distribution. Upper Chalk: Norwich (coll. Bayfield).
Genus SCLEROKALIA, Hinde, n. g.
Sponges cup- or nest-shaped, with very thick walls. Apparently free. The
interior surface of the cup is provided with circular or ovate canal-apertures disposed
in vertical rows. The exterior surface has no special canal-apertures. The canals
opening into the cloaca do not appear to extend far into the wall. A dermal layer,
composed of a thickened reticular mesh with irregular pores, forms the inner surface
of the wall; the outer surface does not possess a special dermal layer. The sub-
stance of the wall is formed of a robust spicular mesh with octahedral nodes. Near
the inner surface the mesh is very regularly disposed, but in the central and outer
portions of the wall the arrangement is of a less regular character. There does not
appear to be any definite boundary between the regular and irregular mesh ; and
the spicular rays are of the same thickness, and the nodes octahedral throughout
the wall.
The type of this genus is an imperfect example of a nest-shaped sponge preserved
in a siliceous matrix. The structure of the interior of the wall has been partly
obliterated, so that the character of the canals cannot be ascertained with precision.
U
146 SILICEOUS SPONGES.
The distinctive character of the sponge is the great thickness of the walls, which are
composed of a continuous mesh, not arranged in a series of folds as in Pachytei-
chisma. Neither can the walls be differentiated into a relatively thin true wall and
a posterior supplemental skeleton as in Stawronema, for the only difference between
the inner and the deeper portions of the wall consists in a gradual transition from a
regular to an irregular disposition of the spicular mesh.
The systematic position of this genus is uncertain. In the characters of the cloacal
surface it resembles sponges of the Euretidee family ; but the octahedral nodes and the
slight development of the canals exclude it from this group. In some respects the
wall-structure resembles that of the genus Diplodictyon ; and it may provisionally
be associated in the same family with this genus.
ScLEROKALIA CunnineToni, Hinde, n. sp. (Plate XX XI. figs. 3, 3a, 36, 3c.)
The only example resembles a bird’s nest in form. The walls are thickest in the
lower and basal portions, the upper margins are rounded. The specimen is 85 mm.
in height and 145 mm. in width. The extreme thickness of the wall is 50mm. The
apertures of the inner surface of the wall are about 1°5 mm. in length and about
2-5 mm. apart. The mesh-interspaces are circular or oval; the nodes are from °45 to
‘5mm. apart. The pores between the rays forming the spicular nodes are very
minute, so that their octahedral characters are not readily perceptible. The spicular
arms are about ‘12 mm. in thickness.
The spicular mesh is only partially preserved ; in some places the rays or arms
have been enveloped in a thin pellicle of silica, which, by the subsequent dissolution
of the inclosed spicules, forms hollow casts of the mesh.
With this species I have associated the name of Mr. William Cunnington, F.G.S.,
from whom the Museum has obtained its most valuable examples of sponges from
the Chalk and Green Sand of Wiltshire.
Distribution. Upper Green Sand: near Devizes, Wiltshire. The type specimen
belongs to the Jermyn-Street Museum.
Family CHLOPTYCHID, Zittel.
Genus CAALOPTYCHIUM, Goldfuss, 1826-33.
C@LOPTYCHIUM AGARICOIDES, Goldf. (Plate XXXI. figs. 4, 4a.)
1826-33. Cwloptychium agaricoides, Goldf. Petref. 1 Th. p. 31, t. 9. f. 20.
1833. Celoptychium agaricoides, S. Woodward, Geol. of Norfolk, t. 4. f. 19.
1841. Caloptychium agaricoides, F. A. Roemer, Nordd. Kreide, p. 10, t. 4. f. 5.
1876. Celoptychium agaricoides, Zitt. Ueber Coeloptychium, Abhand. k. bayr. Ak. II Cl.
Bd. 12, Ab. 3, p. 59, t. 3. f. 1, 4, 5, 6, 10, 11, 13; t. 4A.
C@LOPTYCHIUM. 147
1877. Ceeloptychium agaricoides, Zitt. Studien, I Ab. p. 59; Neues Jahrbuch, p. 371.
1878. Celoptychium agaricoides, Zitt. Handbuch der Pal. Bd. 1, p. 184, f. 100.
1878. Celoptychium lonyostium, Quenst. Petref. Bd. 5, p. 519, t. 140. f. 1, 2.
In addition to very perfect examples of this species from the Upper Chalk of
Westphalia, the Museum possesses a small specimen from the Upper Chalk of this
country which may be referred to this species, though, as the characters of the under
surface are concealed, it is doubtful whether it belongs here or to C. deciminum.
The specimen in question is without a stem; the body is 50 mm. in width and 8 in
thickness. The upper surface is flat with a central depression, the margins are
sharp and slightly elevated. The surface is for the most part composed of a coarsely
reticulate mesh with radiating bands of a finely punctate character. The interior
folds of the wall are about *9 mm. in thickness.
Distribution. Upper Chalk: Coesfeld, Westphalia; Lemforde, Hanover (coll.
Zitt.) ; South of England; Norwich (S. Woodward ).
Ca@LoptTycHium DEciMINUM, F. A. Ramer.
1841. Celoptychium deciminum, Roemer, Nordd. Kreide, p. 10, t. 4. f. 3.
1865. Celoptychium Belfastiense, Tate, Quart. Journ. Geol. Soc. vol. xxi. p. 43, t. 5. f. 7.
1876. Caloptychium deciminum, Zitt. Abhand. k. bayr. Ak. II Cl. Bd. 12, p. 62, t. 1. f. 6, 7,
and t. 3. f. 2.
1877. Celoptychium deciminum, Zitt. Studien, I Ab. p. 59; Neues Jahrbuch, p. 371.
The only example in the collection which I refer to this species is the upper
portion of a specimen from the Norwich Chalk, which is 114 mm. in width and 15
in thickness. The upper surface appears to have been flat or gently concave ; its
minute structure is not shown. On the ridges of the under surface are small circular
or oval apertures, 1‘2 mm. in width. The folds of the interior of the wall are
1:6 mm. in thickness; they are built up of an extremely regular spicular mesh, with
octahedral nodes -44 mm. apart.
The fragment of a specimen from the indurated chalk of Belfast, named by Tate
Celoptychium Belfastiense, appears to me to belong to this species. Only the
ridges of the under surface are preserved ; these are about 6 mm. in width; they are
furnished with small circular oscular apertures 1 mm. in diameter. The spicular
structure has been obliterated. The specimen is in the Jermyn-Street Museum.
Distribution. Upper Chalk: Norwich (coll. Bayfield); Belfast, Ireland.
Ca@LoPrycuiuM FurcATUM, Tate.
1865. Caloptychium furcatum, Tate, Quart. Journ. Geol. Soe. vol. xxi. t. 5. f. 6.
This species is founded on a fragment of the body of a specimen, showing the
ridges of the under surface, which openly bifurcate. A few oval oscular apertures
u 2
148 SILICEOUS SPONGES.
are present on the ridges. No spicular structure has been preserved; and the
specimen is too imperfect for comparison with other species.
Distribution. Spongarian zone of the Chloritic Chalk (Tate) = Upper Chalk: White-
head, near Belfast (Jermyn-Street Museum).
Ca@LOPTYCHIUM SEEBACHI, Zitt.
1876. Caloptychium Seebachi, Zitt. Abhand. k. bay. Ak. II Cl. Bd. 12, p. 68, t. 2. f. 5-7,
to. £1, 9,vand tt.b. tf. 2A
1878. Caloptychium Seebachi, Quenst. Petref. Bd. 5, p. 521, t. 140. f. 3, 4.
1879. Celoptychium Seebachi, Nicholson, Manual of Pal. vol. i. p. 143, f. 38.
Distribution. Upper Chalk: Lemforde, Hanover.
Ca@LoPTYCcHIUM SULCIFERUM, Ramer.
1841. Celoptychium sulciferum, F. A. Roemer, Nordd. Kreide, p. 10, t. 4. f. 4.
1876. Celoptychium sulciferum, Zitt. Abhand, k. bay. Ak. II Cl. Bd. 12, p. 72.
1878. Caloptychium percussum, Quenst. Petref. Bd. 5, p. 512, t. 139. f. 14.
Distribution. Upper Chalk: Coesfeld, Westphalia.
Ca@.optycuium LopatuM, Goldfuss.
1826-33. Celoptychium lobatum, Goldf. Petref. 1 Th. p. 220, t. 65. f. 11.
1864. Celoptychium lobatum, F. A. Roemer, Pal. Bd. 13, p. 4, t. 2. f. 12.
1876. Celoptychium lobatum, Zitt. Abhand. k. bay. Ak. IT Cl. Bd. 12, p. 73.
1878. Celoptychium sexlobatum, Quenst. Petref. Bd. 5, p. 509, t. 139. f. 13.
Distribution. Upper Chalk: Coesfeld, Westphalia.
Suborder LYSSAKINA, Zittel.
Family JONAKID, Marshall.
Genus ASTRAXOSPONGIA, F. Remer, 1854.
ASTRAOSPONGIA MENISCUS, F. Ramer.
1848. Blumenbachium meniscus, F. Roemer, Leonhard und Bronn’s Jahrbuch, p. 680, t. 9.
f. 1 a—c.
1852. Astreospongium meniscus, Bronn’s Leth. Geogn. 2 Th. p. 156, t. 5. f. 1 a-e.
1860. Astreospongia meniscus, F. Roemer, Die silur. Fauna des westl. Tenn. p. 14, t. 1. f. 6.
1878. Astreospongium meniscus, Zittel, Handb. der Pal. Bd. 1, p. 185.
1878. Astreospongia meniscus, Quenst. Petref. Bd. 5, p. 557, t. 141. f. 8.
1879. Astreospongia meniscus, Nicholson, Manual of Pal. vol. i. p. 135. f. 334, 6.
1880. Astreospongia meniscus, F, Remer, Lethea Pal. 1 Th. p. 314, t. 9. f. 2 a-e.
Sponges growing in the form of concavo-convex disks; sessile. Specimens vary
ASTRZOSPONGIA.—STAURACTINELLA. 149
from 40 to 90 mm. in width, and from 12 to 20 mm. in thickness. The entire
sponge is composed of star-shaped spicules, with six flattened or rounded rays in a
horizontal plane; the rays are usually obtuse at the extremities; the centre of the
spicule is slightly expanded. The spicules vary in size: large examples are 8 mm. in
diameter, and the rays are ‘75 mm. in width. Occasionally in the rays of some of
the larger spicules longitudinal furrows, representing the interior canals, are exposed.
The spicules do not appear to have been attached together. ‘The examples are now
partly altered to calcite; even where the siliceous composition has been retained, the
spicules are more or less obliterated.
Distribution. Silurian: Niagara group, Perry County, West Tennessee.
ASTRAOSPONGIA PATINA, [, Remer. (Plate XXXI. fig. 5.)
1861. Astreospongia patina, F. Remer, Die fossile Fauna d. Silur.-Gesch. von Sadewitz,
p- 14, t. 3. £. 5 a—d.
Sponges growing in concavo-convex disks, and sessile, the same as in the preceding
species. The star-like spicules vary from ‘36 to 1-5 mm. in diameter, and the rays from
‘033 to 112 mm. in width. In the centre of most of the spicules is a small protu-
berance, representing a seventh ray at right angles to the other six. This species
differs from A. meniscus in the much smaller size of the component spicules.
This species is only represented in the Museum by some detached spicules, which
I have discovered abundantly present in some decayed shales from the Isle of
Gotland, presented to me by Prof. G. Lindstrém of Stockholm. The spicules are
now composed of calcite; their rays are flattened and obtusely terminated; the
surface is rough and uneven, and no traces of canals have been preserved. These
spicules correspond in size and form with those of the unique example of this species
discovered by Reemer in the glacial drift of Sadewitz in Lower Silesia, and there can
be little doubt as to their belonging to a sponge of this species. The occurrence of
these spicules in situ further supports the conclusion, drawn from the other fossils in
the drift, that this deposit has been derived from the Silurian strata of the Baltic
basin.
Distribution. Silurian strata of Wenlock age: near Visby, Isle of Gotland.
Genus STAURACTINELLA, Zit. 1877.
STAURACTINELLA CRETACEA, Hinde. (Plate XXXI. fig. 6.)
1880. Stawractinella cretacea, Hinde, Foss. Sponge-Spicules, p. 70, t. 5. f. 9-11.
This species is based on detached hexactinellid spicules, with elongated rays
reaching to 2mm. in length. There is in the Museum a subcylindrical specimen
from the Upper Chalk, 42 mm. in length and 34 in width, which shows on the outer
150 SILICEOUS SPONGES.
surface numerous hexactinellid spicules apparently detached and irregularly mingled
together. Slender monactinellid spicules are also present. The specimen has been
treated with acid, and the spicules are now in the condition of iron peroxide. A
large spicule is 3mm. in length. The interior structure of the specimen, owing to
its condition, cannot be determined; but it is probably made up of spicules similar to
those on the exterior, and as these resemble the detached forms to which I have given
the name of S. cretacea, I refer this sponge to the same species.
Distribution. Upper Chalk: Horstead, Norfolk; South of England (coll. Bower-
bank).
Family POLLAKIDZ, Marshall.
Genus HYALOSTELIA, Zitt¢. 1878.
Hyatostetia Smita, Young and Young, sp. (Plate XXXII. figs. 1-1 4.)
1876. Acanthospongia Smithii, Young and Young, Cat. Western Scottish Fossils, p. 38.
1877. Hyalonema Smithii, Young and Young, pars, Ann. & Mag. Nat. Hist. vol. xx. p. 426,
t. 14. f. 1-3, 5-12, 14-17.
1877. Acanthospongia Smithii, Zitt. Studien, I Ab. p. 60.
1878. Hyalonema Smithii, Carter, pars, Ann. & Mag. Nat. Hist. vol. i. p. 129, t. 9.
f. 1-9, 12, 18.
This species was constituted by Messrs. Young to include detached spicules of
various forms which occur, freely intermingled together, in beds of decayed chert of
Carboniferous age in Scotland. Many of these spicules are simple Hexactinellids,
that is with six rays at right angles to each other, whilst others have a greater
number of rays, though they appear to be but varieties of the Hexactinellid type.
Messrs. Young suggested the probability that these various forms of spicules might
belong to more than a single species of sponge; and this was shown to be the
case by Mr. Carter, who subsequently described a sponge, Holasterella conferta, which
is composed of some of the abnormal hexactinellid spicules which Messrs. Young
had included in H. Smithii. I therefore propose to restrict H. Smithii to the
simple hexactinellid spicules, which are the most abundant forms in the beds at
Cunningham Baidland, and to the spicular rods with or without the four anchoring-
hooks at their termination. The simple spicules usually have a main axis, which
varies from 1°5 to 9 mm. in length; the other axes are generally very unequal, and
the rays are frequently inflated, and sometimes reduced to a mere rounded knob. The
spicular rods are met with singly or disposed parallel with each other in compressed
bundles; they are of indefinite length, and vary from ‘3 to 1:5 mm. in diameter.
When perfect they terminate in four short, recurved, anchor-like rays. These rods
are regarded by Messrs. Young as identical with those described by M‘Coy under the
HYALOSTELIA. 151
name of Serpula parailela, but if such were the case, the sponge ought, by the rule of
priority, to retain M‘Coy’s specific name. I do not think, however, that the spicular
rods of H. Smithii are similar to those described by M‘Coy, which, according to the
description, are only ‘5 mm. in thickness, whereas the majority of those found in the
Scotch beds are much thicker, and reach a maximum thickness of 1:5 mm.
Distribution. Lower Carboniferous : Cunningham Baidland, Dalry, Ayrshire. Pre-
sented to the Museum by Dr. J. Millar, F.L.S.
HYALOSTELIA PARALLELA, J/*Coy, sp.
1844. Serpula parallela, M‘Coy, Synop. Carb. Foss. Ireland, p. 169, t. 23. f. 30.
1880. Acestra parallela, F. Roemer, Leth. Geogn. 1 Th. p. 318, f. 60.
Straight cylindrical spicular rods, of indefinite length, and from -2 to 5 mm. in
thickness, disposed parallel with each other so as to form flattened bundles. ‘The
surface of the rods is smooth, and the interior canal is usually preserved. The
originally siliceous composition has been replaced by calcite. Though no anchor-
shaped terminations have yet been discovered in connexion with these rods, there
can hardly be a doubt that they are the root-like appendages of hexactinellid
sponges.
In addition to specimens from the Lower Carboniferous of Scotland, there is a well-
preserved example on a slab of rock, to which no label is attached, but which, from
its appearance and the character of the other fossils on it, seems to have been derived
from the ‘Trenton Limestone at Ottawa.
Distribution. Trenton Limestone: Ottawa, Canada. Lower Carboniferous: Cli-
theroe, Scotland.
HYALosteLia FascicuLus, JZ*Coy, sp.
1855. Pyritonema fasciculus, M‘Coy, Brit. Pal. Fossils, p. 10, t. 1 B. f. 13.
The only example of this species in the Museum is a thin compressed band of
spicular rods about 5°5 mm. in width. ‘The rods vary from °2 to °5 mm. in thickness ;
the best-preserved examples show closely-set concentric transverse wrinkles, but in
others the surface is quite smooth. The axial canals have not been preserved. The
spicules in this specimen are siliceous. The transverse wrinkles distinguish this
species from H. parallela.
No label is attached to the specimen ; but it appears to have been derived from the
Llandeilo district of Wales.
Hyatosteit rusirormis, Hinde. (Plate XXXI. fig. 7.)
1880. Hyalostelia fusiformis, Hinde, Foss. Sponge-Spicules, p. 71, t. 5. f. 12-16.
Free hexactinellid spicules, with an elongated main axis and inflated centre. The
152 SILICEOUS SPONGES.
rays are straight or slightly curved. The spicules vary from “45 to 1°05 mm. in
length.
Distribution. Upper Chalk: Horstead, Norfolk.
Genus HOLASTERELLA, Carter, 1879.
HoLASTERELLA CONFERTA, Carter. (Plate XXXII. figs. 2-2 f.)
1879. Holasterella conferta, Carter, Ann. & Mag. Nat. Hist. ser. 5, vol. ii. p. 141, t. 21.
f. 1-8.
1877. Hyalonema Smithii, Young and Young, pars, Ann. & Mag. Nat. Hist. vol. xx. p. 426,
t. 14. f. 20, 21, 22.
1878. Hyalonema Smithii, Carter, pars, Ann. & Mag. Nat. Hist. vol. i. p. 129, t. 9. f. ll a-e.
Sponge club-shaped, composed of abnormal varieties of hexactinellid spicules. The
simplest form is a spicule with five arms or rays at right angles to each other, and
the sixth ray at the summit of the vertical axis is replaced by several minute upright
or radiating processes. In more complex forms these processes at the summit are
elongated, and occasionally bifurcate so as to form a group of radiating rays, which,
in some cases, are sufficiently numerous to give the spicule a star-like form. The
most abnormal variety is a spicule with a flattened or platter-shaped summit composed
of numerous (18 to 25) fusiform, smooth, or tubercled rays, united laterally so as to
resemble the petals of a flower. The summit is supported on three or five smooth
rays which spring from its under surface. Mr. Carter does not appear to have met
with this variety; but in the collection made by Mr. Bennie it occurs in situ,
associated with the simpler spicules of this species, and it is not improbable that it
may have been a modified surface-spicule. There is a great variety in the dimensions
of the spicules of this species; the rays vary from ‘5 to 2°5 mm. in length.
Distribution. Lower Carboniferous: Law Quarry, near Dalry, Ayrshire. Collected
by Mr. James Bennie of Edinburgh.
HOo.astereLia Younal, Hinde, n. sp. (Plate XXXII. figs. 3-3 d.)
1877. Stellate Spicules, Young and Young, Ann. & Mag. Nat. Hist. ser. 4, vol. xx. p. 420,
t. 14. f. 13, 19, 23, 24, 27, 29.
1878. Hyalonema Smithii?, pars, Carter, Ann. & Mag. Nat. Hist. ser. 5, vol. i. p. 129, t. 9.
f. 10a, 8.
I propose this species to include detached spicules consisting of five to seven com-
pressed, straight, or slightly incurved, horizontally extended rays, springing from a
common expanded centre, with a single vertical ray beneath. ‘The upper surface of
these umbrella-shaped spicules is either smooth or tuberculated. The spicular rays
vary from °4 to 2 mm. in length, and from ‘07 to'7 mm. in width. In the same beds
with these detached spicules there are flattened fragments of spicular mesh composed
HOLASTERELLA. 153
of spicules of similar form to the detached specimens, but with their rays amalgamated
together, so as to form an extended wall with stellate interspaces. The tubercles on
the upper or outer surfaces of the spicular arms are much more developed than in
the detached forms, and the spicules themselves are more robust; but the resem-
blance in form and the association of these fragments with the detached spicules
lead me to regard them as portions of the dermal layer of the same sponge.
These umbrella-like spicules are rare in comparison with the simple hexactinellid
forms of Hyalostelia Smithii occurring with them. At first sight they might be taken
to be merely abnormal varieties of the simple forms ; but the suggestion of Mr. Carter
that they probably belong to a distinct species, appears to me to be justified by the
combination of other irregularly-shaped spicules in Holasterella conferta. I have
named them in honour of Mr. John Young, F.G.S., of the Hunterian Museum,
Glasgow.
Distribution. Lower Carboniferous: Cunningham Baidland, Law Quarry, Dalry,
Ayrshire. Collected by Mr. James Bennie.
HOoLastereLLA Wrieutil, Carter. (Plate XXXII. figs. 4-4 7.)
1880. Holasterella Wrightii, Carter, Ann. & Mag. Nat. Hist. vol. vi. p. 209, t. 14. B. f. 1-7.
This species is based on detached spicules occurring in decayed chert from the
Carboniferous strata of Ireland. The spicules vary much in size and form, but they
are all apparently modifications of hexactinellid spicules. ‘The simplest form is an
ordinary hexactinellid with three axes at right angles to each other. In other forms
one or more of the rays bifurcate, so that the extreme forms resemble small stars.
The spicular rays, both in the simple and complex varieties, are ornamented with
spiral rings. Rod-like spicules terminating in four recurved rays occur in the same
deposit, and may perhaps have formed the root-appendages of the sponge.
Distribution. Lower Carboniferous: Ben Bulbul, Sligo, Ireland. Presented to the
Museum by Mr. Joseph Wright, F.G.S., of Belfast.
HoLasteRELLA Benniet, Hinde, n. sp. (Plate XXXII. figs. 5-5 e.)
1877. Incrusting Sponge ?, Young and Young, Ann. & Mag. Nat. Hist. vol. xx. p. 429, t. 15.
f. 41.
Detached spicules of varied forms and relative large size. In the simplest variety
only five rays are present, four of which are horizontal and at right angles to each
other, and one vertical; the rays are either simple or divided at their extremities.
In more complex varieties there are six primary rays, which are either simple or
divide and subdivide in an irregular manner, so that the extreme forms have a stellate
or a ramose appearance. The spicular axes vary from 2°5 to 4mm. in length, and
from ‘5 to ‘9 mm. in width.
x
154 SILICEOUS SPONGES.
The spicules have a brownish granular appearance, and their surfaces resemble
frosted sugar. In cases where some of the rays are fractured the cavity of the
interior canal with a smooth wall is exposed, and the spicule at first sight looks like
an outer crust investing a foreign body, which probably gave rise to the suggestion of
Mr. J. Young that it might be an incrusting sponge. The spicules of this species
are rare, and as they are usually very fragmentary and possess an altogether anomalous
appearance, their hexactinellid affinities are not readily recognizable. In the peculiar
splitting up of the rays they resemble the complex forms of Holasterella Wrightii,
but they are almost gigantic in comparison, and do not possess the spiral sculpture
distinctive of this species.
‘The spicules figured have been kindly forwarded to me by Mr. James Bennie, and
I have great pleasure in naming the species after him.
Distribution. Lower Carboniferous: Law Quarry, Dalry, Ayrshire.
Incerte sedis.
Genus AMPHISPONGIA, Salter, 1861.
AMPHISPONGIA OBLONGA, Salter. (Plate XX XIII. figs. 12, 12a, 126, 12¢, 12d.)
1861. Amphispongia oblonga, Salter, Mem. Geol. Survey of Great Britain, 32, Scotland,
p. 185, t. 2. f. 3.
1877. Amphispongia, Zitt. Studien, I Ab. p. 45, note.
1879. Amphispongia oblonga, Nicholson, Manual of Pal. 2nd ed. vol. i. p. 185, f. 33.¢, d.
1880. Amphispongia, F. Roemer, Leth. Geogn. 1 Th. p. 317.
This species occurs as free, compressed, elliptical masses, rounded both at the base
and summit, from 30 to 60mm. in length and from 8 to 23mm. in width. The
thickness is inconsiderable. The lower portion of the sponge, from one-fourth to
one-half its entire length, is composed of closely approximated, straight, elongated,
conical spicules, about 3mm. in length, and from ‘75 to 1 mm. in width, arranged so
that their rounded summits form the outer surface of the sponge, whilst their obtuse
points reach to its central axis. The upper portion of the sponge, immediately above
the large conical spicules, consists mainly of cruciform spicules closely arranged in a
regular manner, so that one spicular axis is parallel to the compressed surface of the
sponge, whilst the other axis is at right angles to it. The rays of these spicules are
usually straight, from °6 to 9 mm. in length, and about ‘12 mm. in thickness. In
other spicules five rays(that is one ray at right angles to the other four) are present,
and possibly some may possess six rays, but from the condition of the specimens
the sixth ray, even if originally present, cannot satisfactorily be ascertained. Very
minute filiform monaxial spicules are also present mingled with the cruciform
spicules; and in one specimen there are indications of an exterior surface-layer of
AMPHISPONGIA. 155
filiform spicules regularly arranged in the direction of the length of the sponge.
Neither the spicules of the upper nor those of the lower part of the sponge appear
to have been organically attached together. No canals can be seen, nor in the
majority of the specimens is there any indication of a cloacal cavity ; but in one or
two examples the conical spicules of the basal portion are arranged so that a small
tubular cavity exists in the central axis, but this does not appear to extend into the
upper part of the sponge.
All the examples of this sponge are shown on fractured surfaces of a soft brown
micaceous shaly rock. The spicules have been completely dissolved away, and only
moulds in the matrix remain. These moulds are very clearly defined ; they are usually
empty, but occasionally they are filled with a powdery ochraceous material, probably
peroxide of iron. ‘This unfavourable condition of preservation and the close arrange-
ment of the spicules make it very difficult to discover their true forms. Thus Salter
regarded the radiating conical tubes of the lower part of the sponge as the moulds of
bundles of spicules, whereas I believe them to be the cast of simple conical spicules.
If Salter’s views were correct, the separate spicules of the bundles should have shown
some impression in the moulds, whereas the interior of these moulds is perfectly
smooth and even. The occurrence of these hollow conical moulds, detached from
the sponge and scattered singly throughout the rock, also shows that they were filled
by single spicules, and not by closely grouped bundles. ‘The smaller spicules of the
upper portion of the sponge usually present the appearance of so many minute linear
or fusiform hollows closely arranged side by side, with a minute circular hole in the
centre of each individual. In this position only casts of three rays are exposed, but
when detached from the mass, which frequently occurs, the spicules are seen to possess
four rays in a plane at right angles to each other, and in the meeting-point of the
rays a central hole is present in some instances, indicating the existence of a fifth
ray, and possibly a sixth ray may have projected outwards.
Amphispongia was regarded by Salter and also by Bowerbank as a Calcareous
sponge. ‘lhe supposed triradiate form of the spicules and their complete dissolution
supported this opinion. But the true cruciform figure of the majority of the spicules
and the hexactinellid characters of others, at once show the affinity of the sponge to
siliceous Hexactinellids. The proportions of the spicules also correspond closer with
those of Hexactinellid sponges than with Calcareous forms. ‘The complete dissolu-
tion of the spicules is no longer an argument in favour of their original calcareous
nature, for we now know that fossil siliceous spicules most readily undergo disso-
lution and replacement. The presence of peroxide of iron filling many of the casts
is also in favour of the original siliceous constitution of the spicules, as this material
is usually present in the moulds of undoubted siliceous sponges.
Whilst the form and proportions of the spicules in the upper portion of this
species clearly show a relationship to those of Lyssakine Hexactinellids, their
x2
156 SILICEOUS SPONGES.
peculiar arrangement and combination with the relatively large conical spicules of
the basal portion present characters so entirely different from those of any other
fossil siliceous sponge, that Iam unable to assign it to a definite position in this
order.
My study of this species has been greatly facilitated by the loan of specimens from
Prof. Nicholson, Dr. Traquair, and from the collection of the Geological Survey of
Scotland, through the kind permission of Prof. A. Geikie, Director-General of the
Survey.
Distribution. Silurian strata of Upper Ludlow age: Pentland Hills, near Edin-
burgh.
Genus MORTIERA, De Koninck, 1842.
MortTigeRA VERTEBRALIS, De Koninck.
1842. Mortiera vertebralis, De Kon. Anim. foss. carbon. Belg. p. 12, t. B. f. 3.
1846. Mortiera vertebralis, Mich. Icon. Zooph. p. 253, t. 59. f. la, 1 6.
1852. Mortiera vertebralis, Milne-Edw. & Haime, Brit. Foss. Corals, p. 209.
1872. Mortiera vertebralis, De Koninck, Nouy. Rech. sur les Anim. foss. carbonif. Belg.
p- 163, t. 15. f. 9, 9a.
1880. Mortiera vertebralis, F. A. Roemer, Leth. Geogn. 1 Th. p. 322, t. 39. f. lla, 116.
Sponges detached, growing in the form of short cylinders with biconcave ends, and
presenting a superficial resemblance to the centrum of a fish vertebra. The speci-
mens vary from 8 to 36 mm. in height and from 26 to 35 mm. in diameter. ‘The
wall of the sponge is composed of numerous vertical radiating lamelle, which
extend to the centre in the same manner as the septa of a coral. The specimens are
now calcareous and the minute structure has been obliterated, but traces of spicules,
apparently of a Tetracladine character, can be seen in transverse sections. In one
specimen, which has been less exposed to weathering influences, the exterior surface
is formed of a layer of elongate acerate spicules, about 4 mm. in length, disposed
parallel to the axis of the sponge.
This species was regarded by De Koninck as a coral allied to Cyclolites ; F. Roemer
placed it with the sponges from the presence of fragmentary spicules after treating
specimens with acid. There is no doubt that it is a true sponge, and it probably
belongs to the Lithistide, though its proper position cannot be determined until the
characters: of the interior spicules are ascertained. The septate disposition of the
wall is not peculiar to this species, for a similar arrangement of the wall-tissues also
occurs in the Lithistid sponges Cnemidiastrum stellatum, Goldf. sp., Corallidium
diceratinum, Quenst. sp., and also in sponges of the genus Seliscothon, Zitt.
Instribution. Carboniferous Limestone: Tournay, Belgium.
157
Division IT. CALCAREOUS SPONGES.
Order CALCISPONGIA, Blainville.
Family PHARETRONES, Zittel.
The structure of this group of fossil sponges has lately been studied independently
by several authors, and as the result of their investigations has an important bearing
on the true character and affinities of the family, which have hitherto been the
subject of much discussion, it is desirable to make a preliminary reference to its
history, and to the views lately published respecting it.
The family of the Pharetrones was constituted by Zittel to include a group of
sponges, very variable in outer form, but characterized by possessing a skeleton of
solid anastomosing calcareous fibres, so arranged as to form mesh-like walls of
varying thickness, either with or without distinct canals. The skeletal fibres, when
examined in thin transparent microscopic sections, were seen to be composed of
minute uniaxial or apparently three-rayed spicules; but as the true form of these
minute spicules could very rarely be clearly seen in thin sections, some authorities
disputed their resemblance to the spicules of recent. calcareous sponges, and regarded
them as siliceous spicules which had been changed to their present calcareous consti-
tution through fossilization. It was further maintained that, judging by the fragile
character of the spicules of existing calcareous sponges, it was altogether improbable
that similar forms would have been capable of preservation in the fossil state.
In the course of my examination of the Museum collection, I met with some
examples of these fibrous calcareous sponges which supplied the evidence needed to
determine their true characters. The specimens were in such a peculiar condition
of preservation that I was enabled to detach the component spicules of the fibres
and to ascertain their forms with as much precision as if they had belonged to recent
sponges. The majority of these spicules were three-rayed forms of very minute
proportions, and in addition to these there was on the surface of the sponge a layer
of relatively large three- and four-rayed spicules, strikingly similar in form, size, and
in their relative arrangement to those of existing calcareous sponges. Further, in
thin microscopic sections of other forms, I detected, besides the common three- and
four-rayed spicules, some peculiar fork-shaped spicules, of a form quite unknown in
any siliceous sponge, either fossil or recent, but resembling in the greatest detail the
spicules of some recent calcareous sponges from the Australian coast. I submitted
microscopic fragments of these sponges to Prof. Zittel, who stated, in a short com-
158 CALCAREOUS SPONGES.
munication to the ‘Neues Jahrbuch’*, that they fully proved the position of the
Pharetrones amongst Calcareous Sponges.
In the same number of the ‘ Neues Jahrbuch’ just referred to, there appeared a
lengthy article by Dr. G. Steinmann f on “ Pharetronen-Studien,” which professed to
be the results of five years’ investigation of this group. Steinmann agrees with Zittel
as to the originally calcareous nature of the fibre and to its being composed of small
simple calcareous bodies, which he states are, in their present fossil condition, im-
bedded in a calcareous mass, but originally the mass might have been either horny or
calcareous. In spite of the strong resemblances which Zittel had discovered between
this group of fossils and recent Calcareous sponges, Steinmann concludes that “The
Pharetrones are an extinct independent division of the Ccelenterata, whose skeletal
structures partly resemble those of Sponges, partly those of Hydrozoa, and in part
show completely peculiar characters. A resemblance to their dermal skeleton is
only found in septate Corals and Hydrozoa, and the structure of their skeletal fibres
can only and solely be compared with that of Alcyonaria” {. As, however, within a
few months after the publication of the paper the author himself, in a communi-
cation to the ‘ Neues Jahrbuch’ 4, withdraws his opinions, notwithstanding that they
had received the support of Prof. Moseley, it is not worth while to show their incon-
sistency with a true interpretation of the facts.
In the ‘Annals and Magazine of Natural History’ for January 1883 ||, Mr. H. J.
Carter published “ Further observations on the so-called Farringdon Sponges,” in
which he acknowledges the relationship of many of the Pharetrones to Calcareous
sponges; and he also gives a description and figures in the same paper of a recent
calcareous sponge possessing a skeleton of solid vermiculate fibres, composed of large
and small three-rayed spicules, thus distinctly similar to the skeleton of the fossil
Pharetrones.
Shortly after the appearance of Carter’s paper, Dr. Emil von Dunikowski contri-
buted to the ‘ Paleontographica’§ a highly important treatise on “ Die Pharetronen
aus dem Cenoman von Essen und die systematische Stellung der Pharetronen.” The
author fully corroborates the facts which I had already recorded** respecting the
spicules of the fibre of these sponges; and he adds the further interesting discovery
of traces of interior canals in some of the larger spicules, the supposed absence of
which led Steinmann to insist that these forms could not belong to Calcisponges.
Dunikowski treats fully of the anatomy of the group, and finds so close a resemblance
to that of the recent family of Leucones, Haeckel, that he includes therein the
Pharetrones as a subfamily. Iam unable to agree with this classification, which is
based on the opinion that the original constitution of the skeleton is essentially
* 1882, Bd. 2, p. 204. + Loc. cit. p. 139. + Loc. cit. p. 188.
§ Neues Jahrbuch, 1883, Bd. 1, p. 79. || 5th ser. vol. ii. p. 20. 4 Band 29, 1883.
** Annals & Mag. Nat. Hist. 1882, 5th ser. vol. x. p. 185.
PHARETRONES., 159
similar to that of the recent Leucones, and that its present fibrous character is
merely the result of a secondary alteration of the structure due to fossilization, as
originally it was neither calcareous nor horny, but consisted simply of spicules
imbedded in a mass of parenchym.
To determine the question of the original constitution of the fibrous skeleton of
the Pharetrones it is necessary to study the structure in those specimens which have
suffered least from the effects of fossilization. In the specimens from Warminster,
for example, which have been already referred to, the skeletal fibres have apparently
been altered to a very slight degree, and they are wholly composed of spicules in
close approximation to each other and as closely interwoven together as the strands
of arope. There is no indication of their having been originally loosely imbedded in
a mass of parenchym in the same manner as in recent Leucones, in which, according
to Haeckel *, the chief portion of the skeleton, consisting of the entire thickness of
the body-wall, is composed of irregularly scattered spicules. A similar structure to
that of the Warminster examples is also present in transparent sections of Sestro-
stomella from Vaches Noires. The entire fibre is composed of large and small
spicules in intimate contact with each other, and bordering the margins of the fibre
there is a definite layer of spicular bodies running parallel to it, thus showing very
clearly that the spicular constitution of the fibre is original, and is not due to
secondary alteration. It is true that in many specimens the fibre is now only
partially composed of spicules imbedded in a transparent ground-mass; but this
eondition results from a partial dissolution of the spicular components of the fibre.
Various stages of this alteration may be traced in a series of specimens: in the best-
preserved the fibre is entirely composed of spicules, whilst in those which have been
most changed no spicules at all are present, and the fibre is destitute of organic
structure. J do not think therefore that Dunikowski’s theory of the original simi-
larity of the skeleton of the Pharetrones to that of the recent Leucones can be
sustained ; and whilst admitting the great resemblance of the individual spicules in
these two groups, the fibrous arrangement of the skeleton in the Pharetrones appears
to me sufficient grounds for placing the group, as Zittel has done, in an independent
family.
A short reference may here be made to the characters of the Dermal layer (Dermal-
schicht, Zitt.; Deckschicht, Dunik.) in the Pharetrones, of which two distinct kinds
are present. One kind consists of a thin open layer of relatively large four-rayed
spicules, disposed on the outer surface of the sponge in such a manner that three of
the rays are parallel to the surface, whilst the fourth extends inwardly at right
angles to it. It is only under very favourable circumstances that this kind of dermal
layer has been preserved, and its absence in the specimens examined by Zittel led
him to suppose that it might be a distinguishing peculiarity of the group. It occurs
* Die Kalkschwimme, vol. i. p. 304.
160 CALCAREOUS SPONGES.
in Tremacystia (Verticillites) D’ Orbignyi, H., and Corynella rugosa, H., and traces of
it are present in some species of Eudea, Eusiphonella, and Lymnorea. ‘The other
kind of dermal layer has the appearance of a compact smooth or rugose membrane,
resembling in aspect the epitheca of a coral, which envelopes the sides and some-
times the greater part of the surface of the sponge. The structure of this membrane
is very rarely preserved; but, according to Dunikowski, it is mainly composed of
very irregular three-rayed spicules.
The genera into which Zittel has divided this family are mainly based on diffe-
rences in form and in the canal-structure, and not primarily, as in the case of the
Siliceous sponges, on the spicular characters. This is owing to the fact that in many
forms the spicular constituents of the fibres have been completely obliterated by
fossilization, and notably is this the case with most of the specimens from Triassic
strata; so that an attempt to establish the genera on similarity of minute structure
can be, at the best, only partially successful. Where, however, the spicular characters
of the type species of a genus can be ascertained, it is desirable to include in it only
those sponges with a similar minute structure, and thus constitute the genus on a
natural basis. Fossil calcareous sponges, like siliceous forms, very frequently are
similar in form and in the canal-structure, whilst the minute components of the
fibres, when examined under the microscope, are found to vary very considerably.
Dunikowski has proposed to subdivide the family according to the various combina-
tions of uniaxial, three-, and four-rayed spicules, in the same manner as Haeckel has
divided the recent Leucones; but I have found it extremely difficult in practice ta
determine in thin sections whether many spicules are uniaxial or three-rayed forms,
and it is.equally difficult to decide whether a spicule in which three rays are
shown in the section may not have possessed a fourth ray, so that this arrangement
appears to me not to be available for fossil sponges.
‘here are, however, in the Pharetrones which I have examined, some clearly
marked types of spicular structure which may be briefly mentioned. In the first of
these, which I propose to call the “Corynelia” type, the fibre is nearly entirely com-
posed of subequal, elongate, filiform, three-rayed spicules, arranged parallel to each
other in the direction of the axis of the fibre. In these spicules the basal ray is
so slightly developed that in transparent sections they may easily be mistaken for
uniaxial forms. Relatively large three- and four-rayed spicules are rarely present in
the fibre, though they form, in some species at least, an open dermal layer on the
surface of the sponge. This type of spicular structure is well developed in the
Cretaceous species of Corynelia and in Tremacystia. In a second type, which may
be designated the “Sestrostomella” type, the skeletal fibres are built up of relatively
large three- and four-rayed spicules disposed in the central portions of the fibre and
surrounded by smaller and very irregular forms, which frequently appear as sinuous
lines forming the exterior border of the fibre. ‘This structure is characteristic of
EUDEA. . 161
Sestrostomella, the Jurassic species of Stellispongia, Trachysinia, Oculospongia, Dia-
plectia, and Elasmostoma. In a third type, less clearly marked than the two pre-
ceding, the fibres are of a slender character, and mainly consist of relatively large
three- and four-rayed spicules, either heterogeneously mingled or sometimes disposed
in a single series. This structure is developed in Lymnorea, some Jurassic species of
Peronella, and in Inobolia. It may be named the Lymnorea type. In a fourth type
the fibre consists of uniaxial spicules arranged parallel to each other in the direction
of the fibre. No three-rayed forms appear to be present. The only genus in which
this structure exists is Pharetrospongia.
So far as the material at my disposition has permitted, I have endeavoured to
determine the spicular structure of the various species in the collection, but it will
require much more extended observations before a natural arrangement of this family
can be satisfactorily established.
Nore.—Since the first portion of this work was in type, the acquisition of fresh
specimens of Calcareous sponges has enabled me to make several additions to the
list of genera given on page 17.
Genus EUDEA, Lama. 1821.
EvpEa Manon, MMiinst. sp.
1841. Scyphia? Manon, Minst. Beitr. zur Petref. iv. p. 29, t. 1. f. 15.
1864. Epeudea Manon, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 231, t. 1. f. 2a, d.
1878. Eudea Manon, Zitt. Studien, III Ab. p. 27.
1878. Scyphia Manon, Quenst. Petref. Bd. 5, p. 540, t. 140. f. 31.
Distribution. Trias: St. Cassian Tyrol (coll. Klipstein).
EvpeEa potymorpua, Alipst. sp.
1843. Scyphia polymorpha, Klipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 284, t. 19. f. 12 a, b,c.
1864. Verrucospongia polymorpha, Laube, Denks. d. k, Akad. d. Wiss. Bd. 24, p. 236,
Up dle eel dA
1878. Hudea polymorpha, Zitt. Studien, III Ab. p. 27.
1878. Scyphia polymorpha, Quenst. Petref. Bd. 5, p. 550, t. 140. f. 53.
1879. Eudea polymorpha, Zitt. Neues Jahrbuch, p. 21.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
Eopra ciavata, Lame.
1821. Eudea clavata, Lamx. Expos. méthod. p. 46, t. 74. f. 1, 2, 3, 4
1834. Hudea clavata, Blainv. Man. d’Actin. p. 539, t. 64. £. 3, 3a.
1847. Eudea cribraria, Mich. Icon. Zooph. p. 25, t. 58. f. 8.
162 CALCAREOUS SPONGES.
1878. Eudea clavata, Zitt. Studien, III Ab. p. 27.
1879. Eudea clavata, Zitt. Neues Jahrb. p. 22.
In a vertical transparent section of a specimen from Ranville, the fibres, seen
under the microscope, are clearly defined from the matrix, but the spicular structure is
nearly entirely obliterated ; in one instance, however, a three-rayed spicule, of medium
size, could be distinctly seen. The fibre is about -12 mm. in thickness.
Distribution. Middle Jura: Couche a Polypiers—Ranville, Bénouville ; Calvados.
EvuDEA PERFORATA, Quenst. sp.
1858. Spongites perforatus, Quenst. Der Jura, p. 698, t. 84. f. 26, 27.
1878. Eudea perforata, Zitt. Studien, III Ab. p. 27.
1878. Orispongia perforata, Quenst. Petref. Bd. 5, p. 195, t. 124. £. 22-28.
1879. Eudea perforata, Zitt. Neues Jahrb. p. 22.
The fibre of this species, as seen in a transparent section, is about ‘25 mm. in
thickness; fragments of relatively large three- or four-rayed spicules can be distin-
guished in it, but for the most part the minute structure is destroyed. On the
summit of some specimens there are traces of a dermal layer of three- or four-rayed
spicules.
Distribution. Upper Jura: Randen; Nattheim, Wirtemberg.
EUDEA GLOBATA, Quenst. sp.
1878. Orispongia globata, Quenst. Petref. Bd. 5, p. 195, t. 124. f. 29-34.
1826-33. Manon peziza, Goldf. pars, Petref. 1 Th. t. 34. f. 8a.
1878. Eudea globata, Zitt. Studien, III Ab. p. 27.
Distribution. Upper Jura: Nattheim, Wirtemberg.
EvpEA Pisa, Quenst. sp.
1878. Orispongia pisum, Quenst. Petref. Bd. 5, p. 196, t. 124. f. 35, 36.
1878. Eudea pisa, Zitt. Studien, III Ab. p. 27.
Distribution. Upper Jura: Randen.
EUDEA HIRSUTA, Quenst. sp.
1878. Tubispongia hirsuta, Quenst. Petref. Bd. 5, p. 190, t. 124. f. 16-19.
Distribution. Upper Jura: Randen.
Genus COLOSPONGIA, Laube, 1864.
Conosponera DuBIA, Miimst. sp.
1841. Manon dubium, Miinst. Beitr. zur Petref. iv. p. 28. t. 1. f. 11.
1845. Manon pertusum, Kiipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 282, t. 19. f. 4a, 6.
VERTICILLITES.—CELYPHIA. 163
1864. Colospongia dubia, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 288, t. 1. f. 16.
1878. Colospongia dubia, Zitt. Studien, III Ab. p. 27.
1878. Manon pertusum, Quenst. Petref. Bd. 5, p. 549, t. 140. f. 50, 51.
1882. Colospongia dubia, Steinm. Neues Jahrb. p. 172, t. 6. f. 3 a-d.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
Genus VERTICILLITES, Defrance, 1829.
From the descriptions and figures of the type of this genus, Verticillites cretaceus,
Def., given by the author*, and more recently by Steinmann f, it would appear that
its minute structure is quite unknown; and the peculiar circumstance of its occur-
rence as merely empty moulds, leads me greatly to doubt whether it really is a
Calcareous sponge. Steinmann places it with the Pharetrones solely on account of
its outer form. Unfortunately the Museum does not possess an example of the type
species, so that I have not had an opportunity of seeing it; but until its minute
structure is known, it will be necessary to exclude from the genus other species with
a definite spicular structure hitherto placed under it by different authors. The genus
Verticillites, and its synonym Verticillopora applied by Blainville{ to the same
sponge, will therefore be restricted to its type species V. cretaceus.
Genus ENOPLOCGELIA, Steinmann, 1882.
ENOPLOCG@LIA ARMATA, ATipst. sp.
1843. Scyphia armata, Klipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 284, t. 19. f. 13, 14.
1864. Verrucospongia armata, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 236, t. 1.
f. 10 a, b,c.
1878. Verticillites armata, Zitt. Studien, III Ab. p. 28.
1878. Verrucospongia armata, Queust. Petref. Bd. 5, p. 550, t. 140. f. 55.
1879. Verticilites armata, Zitt. Neues Jahrb. p. 23.
1882. Enoplocelia armata, Steimmann, Neues Jahrb. Bd. 2, p. 166, t. 6. f. 4, 4a.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
Genus CELYPHIA, Pomel, 1872.
CELYPHIA SUBMARGINATA, Miinst. sp.
1841. Manon submarginatum, Minst. Beitr. zur Petref. iv. p. 28, t. 1. f. 9.
1841. Manon pisiforme, Minst. ibid. p. 28, t. 1. f. 10.
* Dict. scienc. nat. 1829, tome lviii. p. 5, Atlas, pl. 44.
T Neues Jahrbuch, 1882, Bd. 2, p. 174, t. 8. f. 2, 2a, 26.
~ Manuel d’Actinologie, 1834, p. 486, t. 66. f.1, la.
164 CALCAREOUS SPONGES.
1864. Verrucospongia submarginata, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 237,
its dle tt WN, The
1878. Celyphia submarginata, Zitt. Studien, III Ab. p. 29.
1878. Testaspongia craniolaris, Quenst. Petref. Bd. 5, p. 539, t. 140. f. 29, 30.
1882. Celyphia submarginata, Steinm. Neues Jahrb. Bd. 2, p. 158, t. 6. f. 6-10, and
tO. tao.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
Genus HIMATELLA, Zittel, 1878.
HIMATELLA MILLEPORATA, Jiinst. sp.
1841. Tragos milleporatum, Mist. Beitr. iv. p. 29, t. 1. f.17.
1864. Lymnoretheles milleporata, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 234, t. 1.
1b 7 Oy
1878. Himatella milleporata, Zitt. Studien, III Ab. p. 29.
1878. Tragos milleporatum, Quenst. Petref. Bd. 5, p. 538, t. 140. f. 25, 26.
1879. Himatella milleporata, Zitt. Neues Jahrb. p. 24.
Distribution. Trias: St. Cassian, Tyrol.
Genus PERONELLA, Zittel, 1878.
The definition of this genus by Prof. Zittel is based mainly on the form of the
sponge and the absence of any definite canal-system. The minute spicular structure
of the fibres is by no means uniform in the various species included in the genus ;
and Zittel suggested the probable necessity of splitting it up, when a better acquaint-
ance with the spicular structure of the different species would allow of a more natural
grouping based on the spicular characters.
Though with respect to the Triassic species no fresh grouping is practicable on
account of the complete obliteration of the minute structure of the fibres, yet in
the species from Jurassic and Cretaceous strata the fibrous structure is, as a rule,
sufficiently well preserved to permit its characters to be defined. It consists of
relatively large and medium-sized three- or four-rayed spicules, the rays of which are
disposed in the central portion of the fibre, and are surrounded by similar, but
smaller, spicules; in some instances the fibre is narrow, and almost exclusively
composed of the rays of the larger spicules. The borders of the fibre do not show
the sinuous spicules which characterize the genus Sestrostomella. I have not found
any simple acerate spicules in the fibres.
The above definition will include all the Jurassic species which I have examined,
but it will exclude for example Peronella multidigitata, Mich. sp., which resembles
typical forms of Peronella in the absence of definite canals, but its spicular structure
is distinctly similar to that of the genus Corynella. As, however, the presence or
PERONELLA. 165
absence of canals in the Pharetrones is but of secondary importance in comparison
with the fibrous structure, this species will require to be placed under Corynelia.
PERONELLA SUBC&SPITOSA, Miinst. sp.
1841. Scyphia subcespitosa, Minst. Beitr. iv. p. 28, t. 1. f. 14.
1878. Peronella subcespitosa, Zitt. Studien, III Ab. p. 32.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
PERONELLA Cymosa, Lamu. sp.
1821. Spongia cymosa, Lamx. Exp. méthod. p. 88, t. 84. f. 7.
1840-47. Scyphia cymosa, Mich. Icon. Zooph. p. 249, t. 58. f. 3a, b.
1854. Spongia cymosa, Morris, Cat. Brit. Foss. p. 30.
1878. Peronella cymosa, Zitt. Studien, III Ab. p. 82.
The only example of this species in the Museum is a small bushy mass, 69 mm. in
width by 40 in height, composed of radiating and occasionally bifurcating cylindrical
stems, about 8 mm. in diameter. ‘The stems are either single or amalgamated laterally.
The cloacal aperture is circular, about 1‘5 mm. in diameter. The anastomosing fibres
are about ‘13 mm. in thickness. I have not been able to ascertain the spicular
structure.
Distribution. Middle Jura: Couche 4 Polypiers—Ranville, near Caen. Great
Oolite: Hampton Cliff, Bath (Morris).
PERONELLA PISTILLIFORMIS, Lamxz. sp. (Plate XX XIII. figs. 1, 1a.)
1821. Spongia pistilliformis, Lamx. Exp. méthod. p. 88, t. 84. f. 6.
1840-47. Scyphia pistilliformis, Mich. Icon. Zooph. p. 250, t. 58. f. 4a, &.
1854. Scyphia pistilliformis, Morris, Cat. Brit. Foss. p. 29.
1878. Peronella pistilliformis, Zitt. Studien, III Ab. p. 32.
Sponges growing in small bushy masses of cylindrical or occasionally inflated
simple or branching stems, from 4 to 6 mm. in thickness. The cloacal aperture is
about 1-6 mm. in width. The exterior surface has a coarsely reticulate aspect.
The fibres, in a transparent section of a specimen from the Great Oolite near Bath,
appear as well-defined bands, from ‘12 to ‘5 mm. in width, of the same character in
the central portions as near the exterior of the stem. They are composed of three-
or perhaps four-rayed spicules, with rays varying from ‘05 to *18 mm. in length,
indiscriminately mingled together. ;
Distribution. Middle Jura: Couche 4 Polypiers—Langrune, Lebisey, Ranville,
Calvados (coll. Tesson). Great Oolite: Hampton Cliff, near Bath.
166 CALCAREOUS SPONGES.
PERONELLA CLAVARIOIDES, Lama. sp. (Plate X XXIII. figs. 6, 6 a.)
1821. Spongia clavarioides, Lamx. Exp. méthod. p. 88, t. 84. f. 8-10.
1878. Peronella clavarioides, Zitt. Studien, III Ab. p. 32.
In a transverse microscopic section of an example of this species from Ranville, the
fibres are clear, well defined, and equally disposed throughout the wall; they vary
from ‘07 to ‘2 mm. in thickness. They are composed of irregular three- and possibly
four-rayed spicules, heterogeneously mingled together. It is somewhat difficult, on
account of the close arrangement of the spicules in the fibre, to ascertain their dimen-
sions; they appear, however, to be relatively large; a single ray, partially visible,
measured *22 mm. in length and -056 mm. in thickness.
Distribution. Middle Jura: Couche a Polypiers—Ranville, near Caen.
PERONELLA MAMILLIFERA, Lamx. sp. (Plate XX XIII. fig. 3.)
1821. Spongia mamillifera, Lamx. Exp. méthod. p. 88, t. 84. f. 11.
1840-47. Spongia mamillifera, Mich. Icon. Zooph. p. 113, t. 26. f. 5.
1854. Spongia mammillifera, Morris, Cat. Brit. Foss. p. 30.
1878. Spongites mamillatus, Quenst. Petref. Bd. 5, p. 340, t. 181. f. 37, 38.
1878. Peroneila mamillifera, Zitt. Studien, III Ab. p. 32.
Sponges growing in depressed bushy masses, from 26 mm. to 60 mm. in width, and
about 32 mm. in height. The individual stems are subcylindrical or mammilliform,
from 6 to 8 mm. in diameter; the cloacal aperture at their summits is from 1°5 to
2 mm. in width, and in some instances three or four open furrows radiate from it.
The individual stems in some examples are nearly free from each other, whilst in
others only the coniform summits are free; in a few instances a corrugated dermal
layer is present on the under surface of the sponge.
The interior structure, as seen in a longitudinal section, is composed of narrow,
ill-defined fibres, about ‘1 mm. in width, which are arranged much closer near the
exterior than in the central portion of the stem. The fibres are now very crystalline,
and only show traces of three- or four-rayed spicules, with the rays disposed in the
central axis of the fibre. The spicular rays are from ‘1 to ‘2 mm. in length, and
about ‘03 in width.
Distribution. Middle Jura: Couche 4 Polypiers—Ranville, L’Amouroux, near
Caen (coll. Tesson); Mont Terrible. Great Oolite: Hampton Cliff, near Bath
(Morris).
PERONELLA TENUIS, Hinde, n. sp. (Plate XX XIII. figs. 2, 2a, 26.)
Sponges forming small bushy masses, of slender, cylindrical, bifurcating stems, from
3°5 to 5°5 mm. in thickness, either free or united nearly their entire length. The
cloacal aperture is about 1 mm. in width.
PERONELLA. 167
In a longitudinal section of a specimen, the fibres near the outer portion of the
stem appear as closely reticulate bands, about ‘1 mm. in width, whilst in the inner
portion they are more open and slender, and seem to be mainly composed of single
axial rays of three- or four-rayed spicules. Similar spicules are also present in the
axial line of the wider marginal fibres, but the other spicules of the fibre have been
obliterated. The spicular rays are from ‘1 to ‘2 mm. in length.
The delicate character and the spicular arrangement of the fibres distinguish this
species from P. pistilliformis, with which it agrees in the form and size of the
individual spongites. Though resembling P. mamillifera in the spicular characters
of the fibres, it is a distinctly smaller form.
Distribution. Inferior Oolite (Pea-Grit): near Cheltenham. Couche a Polypiers:
Ranville (coll. Tesson).
PERONELLA INFLATA, Hinde, n. sp. (Plate XXXII. figs. 6, 6a, 6 6.)
Sponge forming a bushy mass of upright stems or branches. The only specimen
is 100 mm. in height, and about the same in its greatest width. The stems are sub-
cylindrical, with occasional inflations and contractions, and also nodose swellings ;
they vary from 10 to 20 mm. in thickness. At the summit of each stem is a minute
chimney-like extension, from 3°5 to 7 mm. in width, penetrated by a circular cloacal
aperture about 3mm. in diameter. The outer surface of the sponge shows closely
arranged vermiculate fibres, with small irregular interspaces.
In a transparent transverse section of one of the stems there is shown a marginal
ring, about 3 mm. in width, of very closely arranged delicate fibres about -1 mm. in
thickness. hese fibres are formed of three- or four-rayed spicules, of relatively
large size,a single ray measuring ‘25 to ‘3 mm. in length, and from ‘04 to ‘05 mm. in
thickness. The spicular rays are usually arranged in the direction of the fibre. The
central portion of the stem, within the marginal ring, is filled with an open reticulate
fibre, which appears to be mainly composed of similar spicules to those of the marginal
portion, but arranged in a single series, so that the ends of the rays of adjoining
spicules only slightly overlap each other.
The spicular structure of this species is of the same character as in the two
preceding forms, P. tenuis and P. mamillifera; but it is readily distinguished from
these by the larger size and different form of the component stems.
Distribution. Middle Jura: Couche 4 Polypiers—Ranville, near Caen.
PERONELLA CYLINDRICA, Goldf. sp. (Plate XX XIII. fig. 4.)
1826-83. Scyphia cylindrica, Goldf. Petref. 1 Th. p. 5, t. 2. f. 3, and t. 3. f. 12.
1826-33. Scyphia elegans, Goldf. Petref. 1 Th. p. 5, t. 2. £. 5, 13.
1848. Scyphia cylindrica, M‘Coy, Ann. & Mag. Nat. Hist. 2nd ser, vol. ii. p. 418,
168 CALCAREOUS SPONGES.
1878. Spongites cylindricus, Quenst. Petref. Bd. 5, p. 172, t. 123. f. 6, 7, 9-15.
1878. Peronella cylindrica, Zitt. Studien, III Ab. p. 32.
1879. Peronella cylindrica, Zitt. Neues Jahrb. p. 25, t. 2. f. 4.
A transverse microscopic section of a specimen from Heidenstadt shows clearly
marked slender fibres, about ‘08 mm. in thickness, composed of relatively large
three- or four-rayed spicules, irregularly mingled together. A spicular ray measured
‘2 mm. in length and ‘02 mm. in thickness. In some cases the spicular rays project
beyond the margins of the fibre, though not to the same extent as in the specimen
figured by Prof. Zittel in the ‘ Neues Jahrbuch,’ 1879, t. 2. f. 4.
Distribution. Upper Jura: Randen; Heidenstadt, Wiirtemberg. According to
Prof. M‘Coy this species also occurs in the Coralline Oolite at Malton, Yorkshire.
PERONELLA RADICIFORMIS, Quenst. sp.
1878. Radicispongia radiciformis, Quenst. (non Goldf.) Petref. Bd. 5, p. 178, t. 123. f. 16-26.
1878. Peronella radiciformis, Zitt. Studien, III Ab. p. 82.
Distribution. Upper Jura: Randen, Switzerland.
PERONELLA MicHeE ini, Eéallon, sp.
1859. Parendea Michelini, Etallon, Lethea Brunt. p. 420.
1840-47. Spongia lagenaria, Mich. (non Lamx.) Icon. Zooph. p. 114, t. 26. f. 4.
Distribution. Upper Jura: Randen, Switzerland.
PERONELLA NODULOSA, Quenst. sp.
1858. Spongites nodulosus, Quenst. Der Jura, p. 698, t. 84. f. 24.
1878. Spongites nodulosus, Quenst. Petref. Bd. 5, p. 335, t. 131. £. 28-380.
1878. Peronella nodulosa, Zitt. Studien, III Ab. p. 32.
Distribution. Upper Jura: Streitberg; Nattheim, Wurtemberg.
PERONELLA CLAVATA, Rem. sp.
1839. Scyphia clavata, F. A. Roem. Ver. nordd. Oolit. Nacht. p. 10, t. 17. f£. 24.
1864. Siphonocelia clavata, F. A. Reem. Pal. Bd. 13, p. 29, t. 1. f. 2.
1878. Peronella clavata, Zitt. Studien, III Ab. p. 32.
Distribution. Lower Neocomian. Hils-Conglomerate: Berklingen, Brunswick.
PERONELLA TRUNCATA, Mrom. sp.
1861. Siphonocelia truncata, From. Cat. raisonné, p. 7, t. 1. f. 3, 3a.
Distribution. Lower Neocomian: Censeau, near Salins. Jura: Berklingen, Bruns-
wick.
PERONELLA. 169
PrRONELLA RAMOSA, /’. A. Roemer, sp. (Plate XX XIII. fig. 5.)
1839. Scyphia ramosa, I. A. Roemer, Verst. d. Nordd. Oolit. Nachtrag, p. 10, t. 17. f. 27.
1844. Scyphia ramosa, Mant. Medals of Creation, vol. i. p. 227.
1861. Discelia ramosa, From. Cat. raisonné, p. 9, t. 1. f. 5.
1878. Peronella ramosa, Zitt. Studien, ILI Ab. p. 32.
Sponge growing in small bushy masses of branching cylindrical tubes, from 4°5 to
6mm. in diameter. The summits are rounded, with a cloacal aperture about 1°5 mm.
in width. Occasionally the lower portion of the stems is enveloped by a compact
dermal layer; when this is not present, the surface shows only the irregular apertures
between the fibres.
A transparent section of a specimen from Farringdon exhibits indistinctly bounded
fibres about ‘15 mm. in width; the spicular structure has been mostly obliterated, so
that only rarely can small three-rayed spicules be detected.
Distribution. Lower Neocomian: Censeau, Jura. Lower Green Sand: Farringdon,
Berkshire.
PERONELLA GiLuimront, de Loriol, sp. (Plate XX XIII. fig. 10.)
1869. Discelia Gillieroni, de Loriol, Mon. de étage Urgon. p. 66, t. 4. f. 16, 17, 18.
1878. Peronella Gillieroni, Zitt. Studien, III Ab. p. 33.
Sponges growing in bushy masses, from 28 to 40 mm. in height, of cylindrical
branching individuals, from 5 to 7 mm. in diameter. The summits are truncate or
rounded, and the cloacal aperture is from 1:5 to 2mm. in width. There are traces
in some examples of an enveloping dermal layer in the basal portion.
The fibrous structure resembles that of P. ramosa, from which it is distinguished
by the larger size of the individual spongites.
Distribution. Lower Green Sand: Farringdon, Berkshire. Lower Neocomian:
Berklingen, Brunswick.
PERONELLA PROLIFERA, Hinde, n. sp. (Plate XX XIIL figs. 8, 8a.)
Sponges growing in bushy masses, from 40 to 50 mm. in height, of cylindrical
bifurcating stems, with rounded and occasionally inflated summits. The stems vary
in thickness from 6 to 10 mm., and the cloacal tube is from 1°75 to 2 mm. in width.
In a transparent section the fibres are seen to be relatively stout, from -2 to °3 mm.
in thickness; the structure has been largely destroyed by crystallization, but in places
small three- and four-rayed spicules are clearly shown. ‘The spicular rays vary from
04 to ‘1 mm. in length.
This species is distinguished from P. Gillieroni and P. ramosa, with which it occurs
in the same beds at Farringdon, by its larger size and more open mode of growth ;
the fibres also are thicker; but the spicular structure, so far as the condition of the
Z
170 CALCAREOUS SPONGES.
specimens allows of a comparison, appears to be similar, Mr. Davey * has referred
this form to Corynella (Scyphia) multidigitata, Mich.f, but the fibres are thinner,
and the spicular structure is also of a different character.
Distribution. Lower Green Sand: Farringdon, Berkshire.
PERONELLA FLABELLATA, D’Orbigny, sp.
1850. Hippalimus flabellatus, D’Orbigny, Prodr. de Pal. vol. i1. p. 97.
1861. Discelia flabellata, From. Cat. raisonné, p. 9.
1869. Discelia flabellata, de Loriol, Mon. de étage Urgon. p. 66, t. 4. f. 19, 20, 21.
1878. Peronella flabellata, Zitt. Studien, III Ab. p. 33.
Sponges consisting of several branching cylindrical stems, from 5 to 7 mm. in
diameter, attached laterally so as to form fan-shaped expansions. Cloacal tube about
2 mm. in width.
Distribution. Lower Neocomian: Berklingen, Brunswick.
PERONELLA FuRcATA, Goldf. sp. (Plate XX XIII. fig. 7.)
1826-33. Scyphia furcata, Goldf. Petref. 1 Th. p. 5, t. 2. f. 6.
1840. Scyphia furcata, F. A. Roemer, Nordd. Kreide, p. 5, t. 2. f. 6.
1864. Polyendostoma furcatum, F. A. Roemer, Pal. Bd. 13, p. 39, t. 14. f. 5.
1871. Epitheles furcata, Geinitz, Pal. Bd. 20, p. 34, t. 8. f. 7, 8.
1878. Scyphia furcata, Quenst. Petref. Bd. 5, p. 349, t. 182. f. 4-6.
1878. Peronella furcata, Zitt. Studien, IIT Ab. p. 33.
1883. Peronella furcata, Dunik. Pal. Bd. 29, p. 39, t. 39. f. 3, 4.
The examples of this species in the Museum Collection from Essen are small groups
of cylindrical tubes, from 5°5 to 7°5 mm. in diameter, and from 18 to 30 mm. in
height, usually springing from an extended base. The tubes generally bifurcate
near the basal portion, but they also divide near the summit as well. ‘The cloacal
cavity is about 1 mm. in diameter. The lower portion in some specimens shows
traces of a compact dermal layer. The fibres are about ‘12 mm. in thickness, and
closely anastomose together. There are no special apertures on the outer surface.
According to Dunikowski the fibres are mainly composed of relatively large three-
rayed spicules.
Distribution. Cenomanian: Essen. Upper Green Sand: Warminster.
PERONELLA RAMOsISSIMA, Dunik.
1888. Peronella furcata, Goldt., var. ramosissima, Dunik. Pal. Bd. 29, p. 39, t. 39. f. 6.
Distribution. Cenomanian (Upper Green Sand): Essen.
* Transactions Newbury Field-Club, p. 18.
+ Icon. Zooph. p. 217, t. 51. f. 9a, 6.
PERONELLA.—TREMACYSTIA. fal
PERONELLA OCELLATA, Hinde. (Plate XX XIII. figs. 9, 9a, 9b, 9c.)
Sponges mostly simple, subcylindrical stems from 4 to 6°5 mm. in thickness, and
from 15 to 22 mm. in height, apparently growing from a slightly expanded base,
In some cases one or two small lateral buds spring from the surface of the main stem.
The summits are truncate or slightly rounded, and the cloacal aperture is about
1:5 mm. in width. The outer surface is smooth, and is apparently furnished with a
dermal layer, perforated by numerous irregularly disposed circular apertures “4 mm.
in width, generally subdivided by cross bars of fibre. Very minute pores occur in
the interspaces between the larger apertures. The fibres, seen in a transparent
section, are about ‘14 mm. in thickness. Unfortunately no minute structure is
preserved.
This species is readily distinguished by its mode of growth and the characters of
the outer surface.
Distribution. Upper Chalk: Maestricht.
Genus TREMACYSTIA, //inde, n. g.
Syn. Veréicillites, auct. (non Defrance); Verticillopora, Mantell, Sharpe (non Blainville) ;
Ceriopora, pars, Goldf.; Thalamopora, F. A. Roemer, Mich., Pomel, Simonowitsch,
Reuss, Steinm., Dunik.; Discwlia, Loriol; Spherocela, Stemm., Dunik.; Barroisia,
Steinm., Dunik.; Verticillocelia, pars, From.
Sponges simple, or growing in bushy masses from a common base; the stems
occasionally give off lateral branches. Individual forms are cylindrical, inverted-
conical, or club-shaped, and are composed of a series of cylindrical, subspherical, or
cyst-like hollow segments superimposed over each other. ‘The segments are mostly
connected together by a central aperture in the roof of each, and by a more or less
developed axial tube usually perforated laterally with large apertures opening intbd
the different segments. The exterior wall of the sponge and the segmental divisions
are formed by a single layer of anastomosing fibres, penetrated by numerous minute
canals.
The skeletal fibres are composed of minute filiform three-rayed spicules in close
contact with each other, and disposed generally parallel to the axis of the fibre.
The basal ray of these spicules is generally so minute that it can scarcely be
detected in microscopic sections when the spicules are closely intermingled. Besides
these filiform spicules, the outer surface of the sponge is furnished with a dermal
layer of relatively large three- and four-rayed spicules. Possibly uniaxial spicules
may also occur in the fibres, though what appear as such in thin sections are, in my
opinion, portions of three-rayed spicules.
I propose this genus to include a group of well-known sponges, hitherto referred
to several genera, but which are naturally connected by their segmental growth,
Za
172 CALCAREOUS SPONGES.
the perforated walls of a single layer of fibre, and the similarity in the spicular
structure.
TremacystiA D’Orsicny1, Hinde. (Plate XXXIV. figs. 1, 1 a-1 0.)
1882. Verticillites D’ Orbignyi, Hinde, Ann. & Mag. Nat. Hist. 5th ser. vol. x. p. 192, t. 10.
f. 1, 2, 7, 8, and t. 11. f. 1-24.
1816. Alcyonite, W. Smith, Strata identified, t. 6. f. 12.
1882. Spherocelia Michelini, Steimm. pars (non Simonowitsch), Neues Jahrbuch, Bd. 2,
p- 162, t. 7. f. 4 (non 4a, 3).
Small club-shaped sponges, from 16 to 23 mm. in height, either simple or with
lateral branches; growing singly or in small groups from a common extended base.
The individual forms consist of a series, generally four to six in number, of sub-
spherical chambers, from 2 to 4°5 mm. in height and from 4 to 6°5 mm. in width.
The summit chamber is conspicuously larger than those beneath, and measures from
6 to 9mm. in height and from 9 to 14 mm. in width. The floor of each chamber
is formed by the depressed dome-shaped roof of the chamber below it, and thus there
is only a single partition-wall between the respective chambers. A central cylindrical
tube, about 2°25 mm. in diameter, connects the chambers, but I have not been able
to ascertain whether it is continuous throughout.
The walls are from -16 mm. to ‘2 mm. in thickness; they are perforated by
numerous minute circular or subcircular apertures, about *25 mm. in width and
the same distance apart. The outer surface is smooth. The fibre of the walls
is mainly composed of elongated, curved, filiform, three-rayed spicules, disposed
generally in the direction of the fibre, parallel with each other, and in close contact.
These spicules are from -15 to *3 mm. in length, and -006 mm. in thickness. The
basal ray is very slightly developed, and usually appears as a minute projecting knob
in the centre of the spicule. The outer surface of the fibres has a dermal layer of
three- and four-rayed spicules much stouter and larger than the filiform ones beneath.
The longest ray of the largest of these dermal spicules which I have yet met with
measures *3 mm. in length and ‘07 mm. in width. There are numerous gradational
forms between the dermal and the filiform spicules. I have not seen any simple
uniaxial spicules in the fibre.
This species is distinguished from Tremacystia (Thalamopora) siphonioides, Mich.,
by the spherical form of the chambers and its slightly thinner wall. From Zrema-
cystia (Thalamopora) Michelinii, Simonow.*, it differs in possessing a central siphonal
tube, and by the absence of a separate basal floor to each of the chambers. Its
form, slender walls, and the circular perforations in them distinguish it from
Tremacystia (Verticillopora) anastomans, Mant., and its allied species in the Lower
Green Sand.
= Beitr. zur Kennt. der Bry. des Essener Griinsandes, p. 31, t. 1. f. 2a, b,c.
TREMAOYSTIA. 173
Distribution. Upper Green Sand: Warminster, Wiltshire; Essen, Rhenish Prussia
(Steinmann).
‘TREMACYSTIA SIPHONIOIDES, Mich. sp. (Plate XXXIV. fig. 2.)
1840-47. Thalamopora siphonioides, Mich. Icon. Zooph. p. 210, t. 53. f. 9a, b.
1882. Spherocelia Michelinii, Steimmann, pars (non Simonow.), Neues Jahrbuch, Bd. 2,
p- 162, t. 7. f. 4a, b (non f. 4).
Subcylindrical sponges, from 13 to 26 mm. in height, and about 7 mm. in diameter,
with a truncated summit and a cloacal aperture about 1°75 mm. in width. Growing
singly or in small groups on an extended base. There are from 2 to 4 segmental
chambers, from 5 to 6 mm. in height, in the individual sponge. Slight constrictions
are shown on the outer surface, marking the commencement of fresh chambers.
The walls are from *2 to ‘5 mm. in thickness. The interior is not shown in the
specimens I have examined; but according to Michelin there is a continuous cloacal
tube. The walls are perforated by subcircular apertures about ‘2 mm. in width.
I have not been able to ascertain the internal structure of the fibre; but in a
specimen from La Heve there is on the outer surface a dermal layer of three- or
four-rayed spicules similar to those on the surface of T. D’Orbignyi.
This species differs from 7. D’Orbignyi in the cylindrical form of the sponge and
its individual segments. It is distinguished from 7. Michelinii, Simonow., by its
central cloacal tube and the single wall in the floor of the chambers.
Distribution. Craie Chloritée: La Héve. Upper Green Sand: Warminster, Wilt-
shire.
TremacystiA MIcHELINI, Stmonowittsch, sp.
1871. Thalamopora Michelinii, Simonow. Beitr. z. Kennt. der Bry. des Essener Griinsandes,
1s Gully tis JG ab PAS
I have not seen an undoubted example of this species; but according to the
detailed account and figures of Simonowitsch, the sponges consist of a series of four
to five depressed spherical chambers, gradually increasing in size. Each chamber
has a separate base, which rests on the roof of the subjacent chamber, so that there
are double walls between them. No cloacal tube is present, but each chamber com-
municates with that above by a central aperture with a raised margin. The walls are
relatively thin, and perforated by circular and elliptical apertures.
Relying on the accuracy of this description, there appears to me no doubt of the
distinctness of this species from the two preceding. The double partitions of the
chambers and the absence of a cloacal tube clearly mark it off both from 7. D’Or-
bignyt and T. siphonioides.
Distribution. Cenomanian: Essen (Simonowitsch).
174 CALCAREOUS SPONGES.
Tremacystia criprosa, Goldf. sp. (Plate XXXIV. figs. 3, 3a, 36.)
1826-1833. Ceriopora cribrosa, Goldf. Petref. 1 Th. p. 36, t. 10. f. 16 a—c.
1841. Thalamopora cribrosa, F. A. Rem. Nord. Kreide, p. 21.
1847. Monticulipora cribrosa, D’Orbigny, Prod. d. Pal. ii. p. 184.
1871. Thalamopora cribrosa, Simonow. Beitr. z. Kennt. der Bry. des Essener Griinsandes,
195 PAG) tt de tis Ce
1875. Thalamopora cribrosa, Reuss, Pal. Bd. 20, p. 137, t. 33. f. 11-15.
1882. Thalamopora cribrosa, Steinm. Neues Jahrbuch, Bd. 2, p. 167.
1883. Thalamopora, Dunik. Die Pharetronen, p. 43.
Sponges growing singly or in small colonies of separate or branching subcylin-
drical or club-shaped individuals, from 4 to 7 mm. in thickness and up to 27 mm. in
height. Each individual consists of a series of cyst-like chambers, about 1:25 mm.
in height and 2 mm. in width, disposed round a central cloacal tube, about 2 mm. in
diameter, with which each cyst connects by a relatively large aperture. The cloacal
tube is continuous throughout the length of the sponge. The summit of each cyst is
convex, and forms the floor of the cyst above, so that there is but a single partition-
wall between them. The surface of the sponge shows numerous vesicular inflations,
which indicate the exterior limits of the separate cysts.
The outer walls and the partitions between the cysts are very delicate, about
-25 mm. in thickness, and perforated by numerous subcircular apertures about
‘18 mm. in width. The walls are formed of a single layer of fibre, similar to that of
T. DP Orbignyi. The only specimen in the Museum is but a fragment of an individual,
from which I have not been able to ascertain the minute structure; but Dunikowski
states that three-rayed spicules are present in the fibre, and Steinmann asserts that
its spicular constitution resembles that of 7. anastomans, Mant. sp.
This species was placed by Goldfuss, Roemer, and Simonowitsch amongst the
Poiyzoa; Reuss regarded it as a Foraminifer, and Steinmann as an Alcyonarian.
Dunikowski agrees with me that it is an undoubted sponge, but states that it
altogether differs from other Pharetrones in the absence of fibres (Mangel der
Faserziige). I cannot, however, see that the walls of the specimen in the Museum
in anywise differ from those of 7. D'Orbignyi and T. siphonioides. They are formed
of a single thin layer of fibres; and the main difference between this and the species
above named consists in a greater subdivision of the interior chambers of the sponge.
This species is distinguished from 7. vesiculosa, Mich.*, which has a similar cyst-
like arrangement in its interior, by its form and mode of growth.
Distribution. Cenomanian: Essen an der Ruhe, Rhenish Prussia.
* Icon. Zooph. p. 209, t. 53. f. 8.
TREMACYSTIA., 175
TREMACYSTIA ANASTOMANS, Mant. sp. (Plate XXXIV. figs. 4, 4a, 46, 4c.)
1848. Verticillopora anastomans, Mant. Wonders of Geology, p. 636, f. 3.
1854. Verticillopora anastomans, Mant. Medals of Creation, 2nd ed. vol. i. p. 227, f. 4, and
p. 229, f. 3.
1854. Verticillopora anastomans, Sharpe, Quart. Journ. Geol. Soc. vol. x. p. 195, t. 5. f. 1.
1869. Discelia helvetica, Loriol, pars, Mon. Foss. de Landeron, p. 65, t. 5. f.4-7, 9-11.
1874. Verticillites anastomans, Davey, pars, Trans. Newb. Field-Club, p. 13.
1878. Verticillites anastomans, Zitt. Studien, IIL Ab. p. 28; idem, Hand. der Pal. p. 190,
f. 106.
1882. Barroisia anastomans, Steimm. Neues Jahrbuch, Bd. 2, p. 164, t. 8. f. 1.
1883. Verticellites anastomans, Keeping, Fossils of Upware &ce. p. 145.
1883. Verticillites anastomans, Carter, Ann. & Mag. Nat. Hist. ser. 5, vol. ii. p. 21; ib.
vol. ili. p. 26.
Sponges growing in bushy masses, up to 67 mm. in height and 70 mm. in width;
of cylindrical, branching, usually upright stems or tubes, from 4 to 6 mm. in thick-
ness, which are either free or coalesce laterally. The interior of the tubes is divided
into a series of cylindrical chambers by nearly horizontal partitions, about 1:5 mm.
apart. There is a continuous axial cloacal tube, about 1:5 mm. in diameter, which is
perforated by circles of relatively large lateral apertures opening into each chamber.
The outer surface is even, or faintly shows annular constrictions.
The walls vary, even in the same specimen, from ‘5 mm. to 1 mm. in thickness ;
they are perforated by very minute irregular pores, which are merely the interspaces
between the fibres; in some cases the perforations in the interior partitions are sub-
circular, and about ‘16 mm. in width. In the thin sections which I have examined,
only the larger three-rayed spicules of the dermal layer can be seen; but there are
traces of filiform spicules similar to those of 7. D’Orbignyi. Mr. Carter states that
the spicular structure resembles that of this last-named species, and that the surface
is confronted by a crust of pin-like spicules with their heads outwards.
This species is readily distinguished from others of the genus by its bushy growth,
the regular cylindrical form of the tubes, and more particularly by the thickness of
the walls.
Distribution. Lower Green Sand: Farringdon, Berkshire, and Upware, Cam-
bridgeshire.
TREMACYSTIA IRREGULARIS, Hinde, n. sp. (Plate XXXIV. fig. 5.)
Sponges forming colonies of straight or curved cylindrical tubes, from 7:5 to
9 mm. in diameter, and reaching to 62 mm. in height. ‘The interior is divided by
depressed convex partitions into a series of chambers, from 1:25 to 1:75 mm. in
height, some of which extend the width of the tube whilst others are cyst-like. The
siphonal tube is continuous, about 2 mm. in width, and provided with lateral aper-
176 CALCAREOUS SPONGES.
tures, as in the preceding species. The outer surface is even, or with slight swellings
and constrictions.
The walls vary from 24 to -4 mm. in thickness; the perforations and the spicular
structure of the fibres are the same as in 7. anastomans.
This species differs from 7’. anastomans in the larger size of the individual sponges,
the more delicate walls, and the irregular character of the interior chambers. Its
mode of growth differs from that of 7. annulatus, Keeping, sp.; but as the interior
characters of this species are not stated, it is not possible to make a comparison of
these features in the two species.
Distribution. Lower Green Sand: Farringdon and Upware.
TREMACYSTIA CLAVATA, Keeping, sp. (Plate XXXIV. fig. 6.)
1883. Verticellites clavatus, Keeping, Fossils of Upware &c. p. 146.
1869. Discelia helvetica, Loriol, pars, Mon. Foss. de Landeron, p. 65, t. 5. f. 8.
Sponges growing singly or in small colonies. The individual sponges are obconical
in form, from 25 to 52 mm. in height, and about 14 mm. in diameter. The partitions
between the chambers are nearly horizontal, and about 2°75 mm. apart. The central
cloacal tube is continuous, elliptical in section, and about 4 mm. in diameter. It is
perforated by a circular series of relatively large apertures, with collar-like projec-
tions, which open into each chamber.
The walls are from 25 to ‘5 mm. in thickness; the perforations in them are similar
to those of the two preceding species. The form and size of the sponge distinguishes
this from all other species of the genus. It is comparatively rare.
Distribution. Lower Green Sand: Farringdon and Upware.
Genus ELASMOCQCELIA, Ramer, 1864.
ELASMOca@LIa CRASSA, From. sp. (Plate XX XIII. fig. 11.)
1861. Elasmojerea crassa, From. Cat. raisonné, p. 10, t. 2. f. 10.
1878. Elasmocelia crassa, Zitt. Studien, III Ab. p. 34.
The anastomosing walls of the sponge are from 5 to 6-5 mm. in thickness; a single
row of cloacal tubes, about 1:75 mm. in width and the same distance apart, extends
the length of the crest of the walls. The fibres of the outer surface are so closely
arranged that there are only minute circular pores, about ‘16 mm. in width, between
them. ‘The fibrous mesh of the interior of the wall is relatively open; the fibres are
about ‘2 mm. in thickness.
There is but a single example of this species in the Museum; the walls are some-
what thinner than in Fromentel’s type, but the differences do not appear to me to
be sufficient to constitute a distinct species.
Distribution. Lower Green Sand: Farringdon, Berkshire.
ELASMOC@LIA.—CONOCG@LIA. 177
ELasMoca@siia Farrinaponensis, Mant. sp. (Plate XXXIV. figs. 7, 7a.)
1854. Tragos Farringdonensis, Mantell, Medals of Creation, vol. i. p. 229, f. 5.
1874. Tragos Farringdonensis, Davey, Transactions Newbury Field-Club, p. 14.
Sponges with vertically compressed, upright, or fan-shaped walls, from 30 to
45 mm. in height and the same in breadth. The flattened summit is from 10 to
17 mm. in width, and exhibits numerous circular canal-apertures, about 1:25 mm. in
width, irregularly disposed from one to three diameters apart from each other. The
lateral surfaces are smooth, and show irregular pores between the fibres. The canals
appear to extend throughout the sponge in a generally vertical direction.
The fibres of the interior form an open anastomosing mesh ; they are from 12 to
‘2mm. in thickness. The minute structure is not very distinctly shown in the
section examined; it appears to consist of three- and four-rayed spicules with
straight arms: slender filiform spicules of the Corynella type are also present.
The general form of this and the following species corresponds with that of
sponges of the genus Oculospongia, but the absence of a compact dermal layer
and the different characters of the spicular mesh separate them from this genus,
whilst in the large size and vertical extension of the cloacal tubes they resemble
typical forms of Elasmocelia.
Distribution. Lower Green Sand: Farringdon (coll. John Brown).
Evasmocastia MANTEL, Hinde, n.sp. (Plate XXXIV. fig. 8.)
Sponges growing in short, upright, or lobed masses from an expanded base. An
average specimen is 22 mm. in height and 31 in breadth. The upper surface is
flattened or slightly convex; it is penetrated by the apertures of numerous canals,
about 2°25 mm. each in width and about their own diameters apart. The fibres are
from ‘15 to 3mm. in thickness. This species may be distinguished from the pre-
ceding by its form and the much larger dimensions of the canals. It appears to
be rare.
Distribution. Lower Green Sand: Farringdon.
Genus CONOC(ELIA, Zittel, 1878.
Conoca:Lia CRASSA, From. sp.
1861. Stphonocelia crassa, From. Cat. raisonné, p. 7, t. 1. f. 1, 1a.
1878. Conocelha crassa, Zitt. Studien, III Ab. p. 34.
In a vertical transparent section of a specimen from Censeau the fibres exhibit a
somewhat similar structure to that of Sestrostomell/a; they are mainly composed of
filamentous wavy spicules, so closely arranged in the direction of the fibre that it is
impracticable to determine if they are uniaxial or three-rayed forms; in the central
2A
178 CALCAREOUS SPONGES.
portion of the fibres much larger three- or four-rayed spicules are in places distinctly
visible. The fibres vary from ‘14 to °3 mm. in thickness.
Distribution. Lower Neocomian: Censeau, Salins, Jura. Hils-Conglomerate:
Berklingen, Brunswick.
CoNOCHLIA CENTROLEVIS, Rem. sp.
1864. Limnorea centrolevis, F. A. Roem. Pal. Bd. 13, p. 37, t. 1. f. 18.
1878. Conocelia centrolevis, Zitt. Studien, III Ab. p. 34.
Distribution. Lower Neocomian. Hils-Conglomerate; Berklingen, Brunswick.
Genus EUSIPHONELLA, Zittel, 1878.
EvsIPHoNeLtA Bronnul, Miinst. sp.
1826-33. Scyphia Bronnii, Miinst. in Goldf. Petref. 1 Th. p. 91, t. 33. f. 9.
1854. Scyphia Bronnii, Morris, Cat. Brit. Foss. p. 29.
1859. Siphonocelia elegans, From. Introd. a l’étude des Ep. foss. p. 31, t. 1. f. 7.
1859. Parendea gracilis, Etallon, Leth. Brunt. p. 421, t. 58. f. 30.
1878. Scyphia Bronnii, Quenst. Petref. Bd. 5, p. 183, t. 124. f. 1-15.
1878. Eusiphonella Bronni, Zitt. Studien, III Ab. p. 385; id. Handbuch der Pal. vol. i.
pe LOL £109;
On the summit of the wall of an example of this species from Nattheim there is a
dermal layer of three-rayed spicules; possibly, however, the spicules may be furnished
with a fourth ray penetrating into the sponge-wall.
Distribution. Upper Jura: Nattheim, Wiirtemberg. Great Oolite: Minchin-
hampton (Jorvis).
EUSIPHONELLA INTERMEDIA, Miinst. sp.
1826-33. Scyphia intermedia, Minst. in Goldf. Petref. 1 Th. p. 92, t. 34. f. 1.
1878. Scyphia intermedia, Quenst. Petref. Bd. 5, p. 229, t. 125. f. 55-58.
1878. Eusiphonella intermedia, Zitt. Studien, III Ab. p. 35.
Distribution. Upper Jura: Randen; Nattheim.
EUSIPHONELLA PERPLEXA, Quenst. sp.
1878. Scyphia perplera, Quenst. Petref. Bd. 5, p. 230, t. 125. f. 59-63.
1878. Eusiphonella perplexa, Zitt. Studien, II Ab. p. 35.
Distribution. Upper Jura: Randen.
Genus CORYNELLA, Zttel, 1878.
The presence of a canal-system which, according to Prof. Zittel, constitutes the
main difference between this genus and Peronella, appears to me to be too inconstant
CORYNELLA. 179
a feature to serve for generic distinction; but the different characters of the spicular
structure in the forms referred to these genera afford a more satisfactory basis for
separation. Unfortunately, however, this test is inapplicable to the numerous
Triassic and Jurassic forms of the genus, the minute structure of which has not, up
to the present, been ascertained, and owing to their state of preservation is hardly
likely to be known; but if we accept as the type of the genus a Cretaceous species,
Corynella foraminosa, Goldf. sp., we find that the fibres are composed of minute
filiform three-rayed spicules disposed generally parallel with each other in the
direction of the fibre. On the outer surface of the sponge there is an open dermal
layer of relatively large three- and four-rayed spicules, but these are only seen in
specimens which are very favourably preserved. I have not met with any distinctly
simple, uniaxial spicules; those which at first sight appear to be such, and which
Zittel refers to as simple spicules, I have found to be portions of three-rayed spicules,
in which the unpaired or basal ray has but a very slight development.
The inconstancy of the canal-system in sponges of this genus is shown in speci-
mens of Corynella foraminosa from the Lower Green Sand at Farringdon. Whilst
in some individuals canals are present, in others, precisely similar in all other
respects, none are present, or they are restricted to the basal portion of the speci-
men where the walls are thickest. Again, in Corynella multidigitata, Mich. sp., no
canals are present, but the spicular structure resembles that of typical species of
the genus.
CoRYNELLA GRACILIS, Miinst. sp.
1841. Myrmecium gracile, Minst. Beitr. zur Petref. iv. p, 31, t. 1. f. 26, 27.
1864. Hudea gracilis, Laube, pars, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 232, t. 1.
f. 3a, b.
1878. Myrmecium gracile, Quenst. Petref. Bd. 5, p. 526.
1878. Corynella gracilis, Zitt. Studien, III Ab. p. 36.
The fibres of this species, as seen in a transparent section, are from ‘1 to ‘14 mm.
in thickness; in places a few wavy lines can be seen in them, but they are too
indistinct for their real characters to be determined.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
CoRYNELLA PYRIFORMIS, A{lipst. sp.
1843. Cnemidium pyriforme, Klipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 291, t. 20. f. 5.
1864. Hudea gracilis, Laube, pars, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 232.
1878. Corynella pyriformis, Zitt. Studien, III Ab. p. 36.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
180 CALCAREOUS SPONGES.
CorYNELLA ROSA, Laube, sp.
1864. Eudea rosa, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 282, t. 1. f. 4.
1878. Corynella rosa, Zitt. Studien, III Ab. p. 36.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
CoRYNELLA ASTROITES, Jiinst. sp.
1841. Cnemidium astroites, Miinst. Beitr. zur Petref. iv. p. 31, t. 1. f. 24.
1864. Epitheles astroites, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 235.
1878. Tragos astroites, Quenst. Petref. Bd. 5, p. 527, t. 140, f. 7-14,
1878. Corynella astroites, Zitt. Studien, III Ab. p. 36.
In a transparent microscopic section the fibres appear as well-defined, narrow,
anastomosing bands from ‘08 to 1 mm. in thickness; beyond traces of wavy lines no
spicular structure is discernible.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
CorYNELLA CAPITATA, Miinst. sp.
1841. Scyphia capitata, Minst. Beitr. zur Petref. iv. p. 28, t. 1. f. 12.
1864. Lpitheles capitata, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 235, t. 1. f. 8.
1878. Scyphia capitata, Quenst. Petref. Bd. 5, p. 526, t. 140. f. 6.
1878. Corynella capitata, Zitt. Studien, III Ab. p. 36.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
CoRYNELLA LYCOPERDIOIDES, Lama. sp.
1821. Hallirhoa lycoperdioides, Lamx. Exp. méthod. p. 72, t. 78. f. 2.
1840-47. Siphonia lycoperdioides, Mich. Icon. Zooph. p. 251, t. 58. f. 6.
1878. Corynella lycoperdioides, Zitt. Studien, III Ab. p. 36.
Distribution. Middle Jura: Couche a Polypiers—Ranville, Benouville, Caen.
CoryNELLA cosTata, Stahl, sp.
1824. Alcyonites costata, Stahl, Correspondenzbl. Wiirtem. landw. Ver. vi. p. 84, f. 29.
1878. Spongites astrophorus alatus, Quenst. Petref. Bd. 5, p. 207, t. 124, f. 54-57.
1878. Corynella costata, Zitt. Studien, III Ab. p. 36.
Distribution. Upper Jura: Randen; Giengen, Wiirtemberg.
CoRYNELLA QUENSTEDTI, Zitt.
1878. Corynella Quenstedti, Zitt. Studien, III Ab. p. 36.
1878. Spongites astrophorus caloporus et cornucopie, Quenst. Petref. Bd. 5, p. 208, t. 124.
f. 58-64.
Distribution. Upper Jura: Giengen; Oerlingen; Thurnau, Bavaria.
CORYNELLA. 181
CoRYNuLLA ASPERA, From. sp.
1864. Siphonocelia aspera, From. Polyp. Corall. de Gray. t. 15. f. 6.
1878. Corynella aspera, Zitt. Studien, IIT Ab. p. 36.
Distribution. Upper Jura.
CORYNELLA MADREPORATA, Quenst. sp.
1878. Madrespongia madreporata, Quenst. Petref. Bd. 5, p. 212, t. 124. f. 70-72.
1826-33. Cnemidium astrophorum, Goldf. pars, Petref. 1 Th. t. 35. f. 88.
1878. Corynella madreporata, Zitt. Studien, III Ab. p. 37.
Distribution. Upper Jura: Nattheim, Wiirtemberg.
CoRYNELLA FORAMINOSA, Goldf. sp. (Plate XXXIV. figs. 9, 9a, 95.)
1826-33. Scyphia foraminosa, Goldf. Petref. 1 Th. p. 86, t. 31. f. 4.
1851. Scyphia foraminosa, Morris, Cat. Brit. Foss. p. 29.
1854. Scyphia foraminosa et intermedia, Mant. Medals of Creation, vol. i. p. 227, t. 70.
It By (Be
1864. Hndostoma foraminosum, F. A. Roemer, Pal. Bd. 13, p. 39, t. 14. f. 6.
1871. Epitheles foraminosa, Gein. Pal. Bd. 20, p. 33, t. 8. f. 18.
1874. Scyphia foraminosa, Davey, Trans. Newb. Field-Club, p. 13.
1878. Corynella foraminosa, Zitt. Studien, III Ab. p. 37.
1883. Corynella foraminosa, Dunik. Pal. Bd. 29, t. 39. f. 5.
Sponges growing singly, or occasionally two or three individuals are united near
their bases. ‘The individuals are subcylindrical or subconical, usually widest below,
and gradually tapering to the summit. ‘The specimens vary from 12 to 50 mm. in
height, and from 10 to 21 mm.in width. The walls are from 4 to 9 mm. in thickness,
and the cloacal aperture is from 3 to 7 mm. in width.
A compact dermal layer covers the lower portion of the wall in some specimens ;
where this is not present, the exterior surface shows irregular apertures between the
fibres. ‘The interior surface of the cloaca is smooth and perforated with canal-
apertures, but in some examples canals are not apparent, or merely present near
the basal portion where the walls are thickest.
The anastomosing fibres, as seen in a transparent section of a specimen from
the Lower Green Sand at Farringdon, are from ‘18 to ‘24 mm. in thickness; they
are mainly composed of slender, filiform three-rayed spicules, with a few larger
forms intermingled. In the section the filiform spicules appear as if uniaxial,
but they probably all possess a third minute central ray, similar to the spicules
of C. rugosa. I have not detected any surface four-rayed spicules in the specimens
examined.
182 CALCAREOUS SPONGES.
This species is very abundant at Farringdon; the examples are usually larger than
the forms from the Cenomanian at Essen *.
Distribution. Lower Green Sand: Farringdon, Berkshire. Cenomanian= Upper
Green Sand: Essen, Rhenish Prussia.
CoRYNELLA TETRAGONA, Goldf. sp.
1826-33. Scyphia tetragona, Goldf. Petref. 1 Th. p. 4, t. 2. f. 2.
1826-33. Scyphia mamillaris, Goldf. Petref. 1 Th. p. 4, t. 2. f. 1.
1864. Endostoma tetragonum, ¥. A. Roemer, Pal. Bd. 13, p. 39, t. 14. f. 7.
1871. Epitheles tetragona, Gein. Pal. Bd. 20, p. 33, t. 8. f. 9-12.
1878. Scyphia tetragona, Quenst. Petref. Bd. 5, p. 352, t. 132. f. 13, 14.
1878. Corynella tetragona, Zitt. Studien, III Ab. p. 37.
1883. Corynella tetragona, Dunik. Pal. Bd. 29, p. 36, t. 39. f. 1, 2.
Distribution. Cenomanian: Essen, Rhenish Prussia.
CoryNELLA RUGOSA, Hinde. (Plate XXXIV. figs. 10, 10a.)
1882. Corynella rugosa, Hinde, Ann. & Mag. Nat. Hist. ser. 5, vol. x. p. 196, t. 10. f. 4,
and t. 11. f. 25.
Sponges simple, cylindrical, straight or curved, growing from a slender base ; the
summits truncate. The outer surface has concentric ridges and furrows, or tubercular
elevations. An average example is 48 mm. in height and 18 mm. in thickness. The
walls are from 4 to 7 mm. in thickness, and the tubular cloaca is 4 mm. wide at its
summit.
The outer surface exhibits closely arranged, approximately circular pores formed
by the interspaces of the fibre. The fibres are about ‘22 mm. in width, and are
composed of slender filiform three-rayed spicules, closely disposed parallel with each
other in the direction of the fibre. The outer surface is furnished with a dermal
layer of larger four-rayed spicules. ‘The spicular structure of this species closely
resembles that of Tremacystia D Orbigni, H.
Distribution. Upper Green Sand: Warminster, Wiltshire (coll. Cunnington).
* The sponges from Essen, figured by Quenstedt under C. foraminosa (Petref. Bd. 5, p. 301, t. 132.
f. 8-10), differ very materially from the typical examples of this species, and in fact resemble more nearly
some forms of Elasmostoma than Corynella. They are either simple or compound, subcylindrical or funnel-
shaped ; the interior surface is smooth and minutely cribrate, with circular, well-defined, closely approximate
apertures about ‘6 mm.in width. The exterior surface is partially covered with a dermal layer; where this
is not present, the naked fibres of the wall are exposed. Unfortunately the spicular structure is quite
destroyed in the thin section examined; and therefore I propose to allow these forms to remain under Cory-
nella until their minute structure is determined, and to designate the species C.? cribrata.
CORYNELLA.—MYRMECIUM. 185
CorYNELLA soctALis, Hinde. (Plate XXXIV. figs. 11, 11a.)
1882. Corynella socialis, Hinde, Ann. & Mag. Nat. Hist. ser. 5, vol. x. p. 197, t. 10. f. 3.
Sponge compound, consisting of three or more individuals growing from a common
cylindrical stem. The base is either widely expanded or contracted to a blunt point ;
the summits are wide, and obliquely truncate. The outer surface uneven, with short
blunted projections. The type specimen is 49 mm. in height and 45 mm. in its
greatest width.
The apertures of the outer surface are irregular in form. The fibres, as seen in a
transparent section, are from ‘12 to ‘2 mm. in thickness. The spicular structure
resembles that of the last species.
Distribution. Upper Green Sand: Warminster, Wiltshire (coll. Baker).
CoRYNELLA MULTIDIGITATA, Mich. sp.
1840-47. Spongia multidigitata, Mich. sp., Icon. Zooph. p. 217, t. 51. f. 9a, b.
1878. Peronella multidigitata, Zitt. Studien, III Ab. p. 33, t. 12. f.3; idem, Neues Jahr-
buch, 1879, p. 25,t. 2. f. 3.
Sponges growing in relatively large bushy masses, in one example 79 mm. in height
by 83 mm. in width; consisting of cylindrical tubes from 7 to 10 mm. in width,
frequently coalescing laterally. The cloacal aperture is 2mm. in width. The
outer surface exhibits only the irregular apertures between the fibres. No canals
are present.
The fibres, as seen in a transparent section of a specimen from Le Mans, are from
‘14 to ‘2 mm. in width, and are composed of filiform three-rayed spicules, with
slightly developed basal rays similar to those of the preceding species. The length
of the lateral rays is about ‘08 mm.
Distribution. Upper Green Sand: Le Mans, Sarthe.
Genus MYRMECIUM, Goldf. 1826.
Myrmecium? ueroGLypua, Alipst. sp.
1843. Scyphia hieroglypha, Kiipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 284, t. 19. f. 6a, 6.
1864. Epitheles hieroglypha, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 236, t. 1.
He 1G
1878. Scyphia hieroglypha, Quenst. Petref. Bd. 5, p. 549, t. 140. f. 52.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
MYRMECIUM HEMISPHERICUM, Goldf.
1826-33. Myrmecium hemisphericum, Goldf. Petref. 1 Th. p. 18, t. 6. f. 12.
1826-33. Cnemidium rotula, Goldf. ib. p. 16, t. 6. f. 6.
184 CALCAREOUS SPONGES.
1878. Spongites rotula, Quenst. Petref. Bd. 5, p. 234, t. 126. f. 1-41.
1876. Myrmecium hemisphericum, Zitt. Studien, IIT Ab. p. 38.
In a vertical microscopic section of a specimen from Randen the fibres appear as
narrow anastomosing bands from ‘07 to ‘1 mm. in width; the rays of irregular three-
or four-rayed spicules can be distinctly seen in the fibres. An average ray measures
‘2mm. in length by ‘04 mm. in width.
Distribution. Upper Jura: Heuberg, Nattheim, Thurnau; Heiligenstadt ; Basle ;
Randen.
MyrMeEciuM InDUTUM, Quenst. sp.
1858. Spongites indutus, Quenst. Der Jura, p. 698, t. 84, f. 21, 22.
1878. Spongites indutus, Quenst. Petref. Bd. 5, p. 245, t. 126. f. 42-46.
1878. Myrmecium indutum, Zitt. Studien, III Ab. p. 38.
Distribution. Upper Jura: Nattheim, Wurtemberg.
Genus LYMNOREA, Lamz. 1821.
LyMNOREA MAMILLOSA, Lamz. (Plate XXXV. figs. 1, la, 10.)
1821. Lymnorea mamillosa, Lamx. Exp. méthod. p. 77, t. 79. f. 2-4.
1878. Lymnorea mamillosa, Zitt. Studien, III Ab. p. 39.
Sponges growing in hemispherical or subspherical masses, from 15 to 25 mm. in
height and from 20 to 38 mm. in width. The basal portion is enveloped in a
concentric, wrinkled, compact dermal layer; in some instances the base is concave.
The rounded upper surface is furnished with small conical, more or less projecting,
eminences, with a circular aperture, about 1 mm. wide, at the summit of each. In
a vertical section there are traces of concentric layers of growth. The canals, in
some instances at least, extend irregularly through the sponge.
The fibres closely anastomose; they are about ‘12 mm. in width. In the transparent
sections which I have examined their structure is largely crystalline, but axial three-
or four-rayed spicules, with rays about °24 mm. in length, are clearly shown. In one
specimen the fibres on the summit of the sponge exhibit an open dermal layer of
spicules, probably four-rayed, similar to those of Zremacystia DL’ Orbigny?.
Distribution. Middle Jura: Couche & Polypiers—Les Moustiers, Ranville, near
Caen. Inferior Oolite (Pea-grit): near Cheltenham.
Genus INOBOLIA, /inde, n. g.
Sponges inverted conical, subspherical, or irregular in form, with convex summits.
The lateral surface in perfect specimens is inclosed in a compact wrinkled dermal
layer. No special canals appear to be present, and the summit only shows the
INOBOLIA.—STELLISPONGIA. 185
irregular interspaces between the fibres. The fibres are mainly composed of rela-
tively large three- and four-rayed spicules, the rays of which are disposed in the axis
of the fibre. The dermal layer is composed of minute, apparently three-rayed
spicules.
In its spicular structure this genus is allied to Lymnorea and some species of
Peronella; but its mode of growth and the absence of canals readily distinguish it
from both of these genera.
Ivopotia INcLusA, Hinde, n. sp. (Plate XXXV. figs. 2, 2a, 26.)
The examples of this species are from 20 to 42 mm. in height, and from 27 to
50 mm. in width. The fibres, as seen in a vertical transparent section, vary from
‘15 to -2 mm. in width; they anastomose so as to form a very irregular mesh. Their
condition is largely crystalline, so that only the rays of the large axial spicules can
be distinguished. A single ray of one of these measured *24 mm. in length by ‘036
in width. In one place the dermal layer, examined by a strong lens, shows numerous
minute three-rayed spicules, the rays of which closely intercross each other.
Whilst in the best-preserved examples no traces of canals are to be seen, there
are some specimens which show here and there the apertures of tubes resembling
those of Oculospongia. These tubes, however, have no regular course in the substance
of the sponge, and appear to me to result from extraneous sources. The spicular
structure of specimens possessing the tubes is similar to that of typical examples of
the species. ‘This species is apparently abundant.
Distribution. Inferior Oolite. (Pea Grit) near Cheltenham.
Genus STELLISPONGIA, D’Orbigny, 1847.
STELLISPONGIA ROTULARIS, Miinst. sp.
1841. Cnemidium rotulare, Minst. Beitr. zur Petref. iv. p. 31, t. 1. f. 25.
1841. Cnemidium Manon, Minst. ib. p. 30, t. 1. f. 20.
1841. Cnemidium astroites, Miinst. ib. p. 31, t. 1. f. 24.
1864. Stellispongia Manon, Laube, Denks. d. k. Akad. d. Wiss. Bd. 24, p. 238, t. 1,
ts Wey, Oy Ge
1878. Cnemidium rotulare, Quenst. Petref. Bd. 5, p. 527.
1878. Stellispongia rotularis, Zitt. Studien, III Ab. p. 40.
Distribution. Trias: St. Cassian, Tyrol (coll. Klipstein).
STELLISPONGIA VARIABILIS, Miinst. sp.
1841. Cnemidium variabile, Minst. Beitr. zur Petref. iv. p. 30, t. 1. f. 21-23.
1841. Cnemidium turbinatum, Miinst. ib. p. 30, t. 1. f. 19.
1843. Cnemidium stellare, Klipst. Beitr. z. g. Kennt. d. dst. Alpen, p. 291, t. 20. f. 6.
2B
186 CALCAREOUS SPONGES.
1843. Cnemidium concinnum, Klipst. ib. p. 292, t. 20. f. 7a, b.
1878. Stellispongia variabilis, Zitt. Studien, III Ab. p. 40.
1882. Stellispongia variabilis, Steinm. Neues Jahrb. Bd. 2, p. 180, t. 9. f. 2.
The fibres, in a transparent microscopic section, appear as well-defined anastomosing
bands about ‘1 mm. in width; no structure could be distinguished. According to
Steinmann they are composed of short, simple, sinuous spicules.
Distribution. Trias: St. Cassian, Tyrol (coll. Alipstein).
STELLISPONGIA HYBRIDA, Miinst. sp.
1841. Tragos hybridum, Miinst. Beitr. zur Petref. iv. p. 29, t. 1. f. 16.
1878. Stellispongia hybrida, Zitt. Studien, IIL Ab. p. 40.
Distribution. Trias: St. Cassian, Tyrol (coll. Alipstein).
STELLISPONGIA STELLATA, Lama. sp.
1821. Spongia stellata, Lamx. Exp. méthod. p. 89, t. 84, f. 13.
1840-47. Spongia umbellata, Mich. Icon. Zooph. p. 248, t. 58. f. la, 6.
1851. Spongia stellata, Morris, Cat. Brit. Foss. p. 30.
1878. Stellispongia stellata, Zitt. Studien, III Ab. p. 40.
1883. Non Séellispongia stellata, Dunik. Pal. Bd. 29, p. 38, t. 40. f. 5.
The spicular structure of the fibres of this species is clearly of the Sestrostomella
type. The fibres are from ‘15 to °24 mm. in thickness, and are composed of rela-
tively large irregular three- and four-rayed spicules, with rays from °3 to ‘53 mm.
in length. These spicules occupy the central portions of the fibre, and they are
surrounded by smaller sinuous, apparently three-rayed, spicules, which form the
exterior borders.
Distribution. Middle Jura: Couche a polypiers—Langrune, Ranville, near Caen
(coll. Tesson). Great Oolite: Hampton Cliff, near Bath (J/orris).
STELLISPONGIA CORALLINA, From. sp. (Plate XXXV. figs. 5, 3a, 5 6.)
1859. Enaulofungia corallina, From. Introduc. 4 VEtude des Ep. fossiles, p. 48, t. 3.
f.11, lla.
1859. Enaulofungia globosa, From. ibid. t. 4, f. 3.
1840-47. Cnemidium rotula, Mich. (non Goldf.) Icon. Zooph. p. 115, t. 26. f. 7.
1859. Astrospongia corallina, Ktallon, Leth. Brunt. p. 424, t. 59. f. 8, 9.
1878. Stellispongia corallina, Zitt. Studien, III Ab. p. 40.
Sponges usually simple, though sometimes two or three individuals are amalga-
mated together, subspherical, growing apparently free or attached to some other
organism. ‘The individuals vary from 9 to 21 mm. in diameter. The summit is
generally slightly depressed in the centre, from whence several well-marked open
canals, ‘6 to 1:25 mm. in width, radiate down the sides.
STELLISPONGIA.—SESTROSTOMELLA. 187
In a transparent section of a specimen from the Coral Rag at Suffield, in York-
shire, the fibres appear as narrow, open, anastomosing bands, from ‘14 to ‘3 mm. in
width; they are composed of relatively large three- or four-rayed spicules with an
exterior layer of smaller sinuous apparently irregular three-rayed forms.
Distribution. Coral Rag: Lyneham, Wiltshire; Suffield, Yorkshire. Upper Jura:
Randen.
STELLISPONGIA GLOMERATA, Quenst. sp.
1858. Spongites glomeratus, Quenst. Der Jura, p. 695, t. 84. f. 10, 11.
1840-47. Cnemidium stellatum, Mich. (non Goldf.) Icon. Zooph. p. 115, t. 26. f. 8.
1859. Didemospongia Thurmanni, Etallon, Leth. Brunt. p. 422, t. 59. f. 3.
1859. Stellispongia pertusa, aperta, hybrida, et glomerata, Etallon, ibid. pp. 423, 424, t. 59.
f. 4-7.
1878. Stellispongia glomerata, Zitt. Studien, III Ab. p. 40.
In a transverse microscopic section of a specimen from Basle the fibres very openly
anastomose ; they are about ‘28 mm. in width. ‘The minute structure resembles that
of S. corallina.
Distribution. Upper Jura: Giengen, Wiirtemberg; Basle.
STELLISPONGIA SEMICINCTA, Quenst. sp.
1878. Spongites semicinctus, Quenst. Petref. Bd. 5, p. 214, t. 125. £. 2-9.
1878. Stellispongia semicincta, Zitt. Studien, III Ab. p. 40.
In the Woodwardian Museum at Cambridge there is an imperfect specimen from
the Coral Rag at Scarborough, which, so far as Iam able to determine, appears to
belong to this species. It had been referred to Spongia floriceps, Phill.; but the
figure of this species in the ‘Geology of Yorkshire,’ pl. 3. f. 8, is insufficient for
comparison.
Distribution. Upper Jura: Randen; Giengen. Coral Rag: Scarborough, York-
shire.
Genus SESTROSTOMELLA, Zitte/, 1878.
SESTROSTOMELLA CRIBRATA, Quenst. sp.
1878. Spongites (Nudispongia) cribratus, Quenst. Petref. Bd. 5, p. 220, t. 125. f. 14-18.
1878. Sestrostomella cribrata, Zitt. Studien, III Ab. p. 41.
The fibres, as seen in a transparent section, are about ‘12 mm. in width; their
spicular structure is nearly entirely obliterated.
Distribution. Upper Jura: Randen, Switzerland.
188 CALCAREOUS SPONGES.
SESTROSTOMELLA RUGOSA, Hinde. (Plate XXXV. figs. 4, 4a, 4 b-d.)
1882. Sestrostomella rugosa, Hinde, Ann. & Mag. Nat. Hist. ser. 5, vol. x. p. 198, t. 10.
f. 6, and t. 12. f. 1-15.
Sponges growing in upright bushy masses, consisting of subconical and subcylin-
drical individuals, about 12 mm. in thickness, united at their bases and occasionally
laterally, but free at their summits. The type specimen is 68 mm. in height and
76 mm. in breadth. The lower portion of the sponge is enveloped in a compact
rugose dermal membrane. The surface exhibits an irregular open network of coarse
fibres. Where a cloacal aperture is present, it is about 2 mm. in width, but the
summits of some individuals do not show any distinct aperture ; in nearly all, open
canals extend from the summit down the sides of the sponge.
The fibres, as seen in a transparent transverse section, vary from ‘2 to ‘41 mm. in
width. They are composed of relatively large, axial or subaxial, three- and four-
rayed spicules, the rays of which, in some cases, reach to ‘5 mm. in length and
‘06 mm. in thickness. These central spicules are surrounded by small and minute
irregular three-rayed spicules, which present an appearance of wavy lines closely
following the contours of the fibre. ‘These smaller spicules are so closely interwoven
together that it is difficult to determine their individual forms; but amongst others
there is a very characteristic slender spicule, resembling in form a pitch-fork or
tuning fork. The basal ray or the handle is straight and tapering, whilst the paired
rays or tines of the fork are subparallel, and frequently one is slightly longer than
the other. The longest of these pitch-fork spicules which I have seen measures
altogether *3 mm.
Distribution. Cretaceous. Upper Green Sand?: Vaches Noires, near Havre.
SESTROSTOMELLA CLAVATA, Hinde. (Plate XXXV. figs. 5, 5a.)
1882. Sestrostomella clavata, Hinde, Ann. & Mag. Nat. Hist. ser. 5, vol. x. p. 201, t. 10.
£5, and t. 12. f. 16-25.
Sponges growing in bushy masses, consisting of numerous cylindrical individuals,
from 8 to 10 mm. in diameter, radiating from a common centre. No pedicel is
present, but a small smooth place indicates the spot on which the sponge rested
during its growth. The summits are rounded; the cloacal aperture is about 2°5 mm.
in width; in some cases only canal-apertures are present at the summit. Straight
open canals extend from the summit down the sides.
The minute structure of the fibres of this species is essentially similar to that of
the preceding, but it differs in form, mode of growth, and the somewhat more slender
dimensions of the fibres. In one part of a thin transverse section the characters of
the exterior portion of the fibre are clearly exhibited, and it is seen to be made up
TRACHYSINIA. 189
of minute irregular three-rayed spicules closely intermingled; amongst these the
pitch-fork forms are conspicuous.
Distribution. Cretaceous. Upper Green Sand: Vaches Noires, near Havre.
Genus TRACHYSINIA, Hinde, n. g.
Sponges either single or growing in bushy masses. The individual spongites are
depressed, cylindrical, with uneven, nodose surfaces; the summits generally inflated.
The cloacal tube is subcylindrical, shallow, or extending to some depth; in some
instances open radiating canals extend from its margins. ‘The interior canals appear
to be but slightly developed, the circulation taking place in the interspaces of the
coarse fibrous mesh. +
The fibres are composed of relatively large three- and perhaps four-rayed spicules
heterogeneously mingled with smaller forms. In some places the rays of the larger
spicules are in the axis of the ‘fibre, but this disposition is not so general as in
Sestrostomella. The margins of the fibre show lines of sinuous spicules as in this
last-named genus.
The minute fibrous structure of this genus shows its alliance to Sestrostomella,
from which it is distinguished by the tubular character of the cloaca and the nodose
mode of growth. In general form it resembles Corynella, but the spicular structure
distinguishes it from the typical forms of this genus. It appears to me not unlikely
that the minute structure of some of the Jurassic species of Corynella, such as
C. Quenstedti, Zitt., for example, will be found to correspond with that of the
present genus.
TRACHYSINIA ASPERA, Hinde, n. sp. (Plate XX XV. figs. 6, 6a, 6 0.)
Sponge forming a mass, 60 mm. high by 75 wide, of depressed cylindrical indivi-
duals, from 26 to 37 mm. in thickness. The outer surface is very uneven and covered
with nodose excrescences. The summits are inflated with occasionally slightly
elevated necks to the cloacal apertures, which are from 7 to 12 mm, in width. The
surface exhibits irregular apertures between the fibres.
The fibres, as seen in a transverse section, vary from ‘14 to ‘5 mm. in width; the
largest spicular ray observed is *2 mm. in length by ‘04 mm. in width.
Distribution. Middle Jura: Couche a polypiers—Ranville, near Caen (coll. Lesson).
TracuysiniA souirarta, Hinde, n. sp. (Plate XXXVI. fig. 9.)
Sponge simple, with a short cylindrical body supported on a compressed stem.
Surface with nodose excrescences. The summit is truncate, slightly concave, the
cloaca is shallow, with canals radiating from the margins; it is about 5 mm. wide.
‘The fibres are about -2 mm. wide. ‘The spicular structure has not been ascertained.
190 CALCAREOUS SPONGES.
This species differs from the preceding in its simple mode of growth and the more
delicate character of the fibres.
Instribution. Middle Jura: Couche a polypiers—Ranville (col/. Tesson).
TracnysiniA MINOR, Hinde, n. sp. (Plate XXXVI. fig. 10.)
Sponges forming small groups of subcylindrical individuals, from 10 to 18 mm. in
thickness, frequently united laterally. The outer surface is uneven and perforated
with large and small irregular apertures. The summits are usually inflated; the
cloacal aperture is from 3 to 5 mm. in width and has several well-marked open
canals radiating from its margin. The fibres, as seen in a transverse section, are
about ‘2 mm. in width; the spicular structure resembles that of 7. aspera, but in
the specimen examined marginal sinuous spicules are not present.
Distribution. Middle Jura: Couche a polypiers—Lebisey; Ranville, near Caen
(coll. Tesson).
Genus BLASTINIA, Zittel, 1878.
Bastia costata, Goldf. sp.
1826-33. Achilleum costatum, Goldf. Petref. 1 Th. p. 94, t. 34. f. 7.
1878. Spongites costatus, Quenst. Petref. Bd. 5, p. 221, t. 125. f. 19-23.
1878. Blastinia costata, Zitt. Studien, III Ab. p. 42.
Distribution. Upper Jura: Randen; Nattheim, Wiirtemberg.
BLASTINIA ALATA, Quenst. sp.
1858. Spongites alatus, Quenst. Der Jura, p. 699, t. 84. f. 28, 29.
1878. Spongites alatus, Quenst. Petref. Bd. 5, p. 222, t. 125. f. 24, 25.
1878. Blastinia alata, Zitt. Studien, III Ab. p. 42.
In a vertical microscopic section of a specimen from Nattheim the fibres bounding
the circular pores are from ‘09 to ‘12 mm. in thickness; their structure is, to a great
extent, obliterated, but here and there clearly marked portions of three-rayed spicules
can be distinguished.
Distribution. Upper Jura: Nattheim, Wiirtemberg.
Genus SYNOPELLA, Zittel, 1878.
SYNOPELLA PULVINARIA, Goldf. sp. (Plate XXXVI. fig. 1.)
1826-33. Manon pulvinarium, Goldf. pars, Petref. 1 Th. p. 2, t. 29, f. 7a, 5 (non t. 1.
f. 6 a, b).
1864. Tremospongia pulvinaria, F. A. Reemer, Pal. Bd. 13, p. 40, t. 14. f. 8.
1878. Synopella pulvinaria, Zitt. Studien, III Ab. p. 43.
The sponges from the Lower Green Sand at Farringdon, which I refer to this
SYNOPELLA.—OCULOSPONGIA. 191
species, are small, hemispherical, or amorphous masses, from 28 to 42 mm. in width
and about 26 mm. in height, frequently with a concave base, enveloped in a compact,
concentrically wrinkled dermal layer; the rounded upper surface is either smooth or
with slight elevations, on which several small canal-apertures are grouped together.
The interspaces between the canal-openings only exhibit irregular pores between the
fibres. The fibres are from ‘15 to 3 mm. in width. I have not been able to ascertain
the spicular structure.
The Farringdon examples are smaller than the specimen figured by Goldfuss from
Essen, but in other respects they appear to correspond. The forms from the
Maestricht Chalk, also referred by Goldfuss to the same, appear to me to belong to
a different species; and as Zittel has applied Goldfuss’s name to the Essen forms, I
propose to adopt another designation for those from Maestricht.
Distribution. Lower Green Sand: Farringdon, Berkshire.
SYNOPELLA SPHA&RICA, Mich. sp.
1840-47. Lymnorea spherica, Mich. Icon. Zooph. p. 216, t. 52. f. 16.4, b.
1878. Synopella spherica, Zitt. Studien, III Ab. p. 43.
Distribution. Cenomanian: Essen an der Ruhe.
SYNOPELLA Goupruss!, Hinde, n.sp. (Plate XXXVI. fig. 2.)
1826-33. Manon pulvinarium, Goldf. pars, Petref. 1 Th. p. 2, t. 1. f. 6a, b (non t. 29.
1 WG, Oy
Sponges inversely conical, subcylindrical, flattened, cake-shaped, or laterally com-
pressed, from 8 to 20 mm. in height and from 10 to 19 in width. The base and
sides are invested in a compact, concentrically wrinkled dermal layer ; the truncate or
rounded upper surface has from one to three small groups of canal-apertures, from
which several deeply marked open furrows radiate ; beyond these the surface exhibits
irregular apertures between the fibres. Minute structure unknown.
The radiating canals on the summit, which readily distinguish this from the two
preceding species, also give it the appearance of Stellispongia, and one can with
difficulty decide whether it should be ranged under that or the present genus.
Distribution. Upper Chalk: Maestricht.
Genus OCULOSPONGIA, From. 1859.
OCULOSPONGIA BINOCULATA, Quenst. sp.
1878. Spongites binoculatus, Quenst. Petref. Bd. 5, p. 248, t. 126. f. 59.
1878. Oculospongia binoculata, Zitt. Studien, III Ab. p. 43.
Distribution. Upper Jura: Giengen, Nattheim, Wiirtemberg.
192 CALCAREOUS SPONGES.
OCULOSPONGIA DILATATA, Ram. sp. (Plate XXXVI. fig. 3.)
1864. Tremospongia dilatata, F. A. Roem. Pal. Bd. 18, p. 40, t. 1. f. 24a, 6.
1874. Oculospongia ( flabellata ?), Davey, Transac. Newbury Field Club, p. 11.
1878. Oculospongia dilatata, Zitt. Studien, III Ab. p. 43.
Sponges growing in small hemispherical or laterally extended masses, from 15 to
20 mm. in height and 16 to 34 in width. ‘The base is inverted conical, flattened or
concave, and invested with a concentrically furrowed and wrinkled dermal layer.
The upper surface only exhibits irregular interspaces between the mesh-fibres and
irregularly scattered canal-apertures, with complete margins, about 1 mm. in width,
which, in the best-preserved specimens, slightly project above the general surface.
In a vertical section the fibres appear as so many radiating lines, with transverse
connections. Some of the canals extend from the base to the surface, whilst others
only reach a short distance into the sponge. The fibres are from ‘1 to *3 mm. in
thickness; the spicular structure, as seen in a specimen from Berklingen, is of the
Sestrostomella type. No structure is shown in examples from Farringdon which I
have examined.
Distribution. Lower Green Sand: Farringdon. Hils-Conglomerate: Berklingen,
Brunswick.
OCULOSPONGIA TUBULIFERA, Goldf. sp.
1826-33. Manon tubuliferum, Goldf. sp. Petref. 1 Th. p. 2, t. 1. f. 5.
1878. Oculospongia tubulifera, Zitt. Studien, III Ab. p. 43.
The fibres, as seen in a transparent microscopic section, are from "16 to 3mm. in
thickness ; no spicular structure has been preserved.
Distribution. Upper Chalk : Maestricht.
Genus CRISPISPONGIA, Quenst. 1878.
CRISPISPONGIA PEZIZOIDES, Zitt.
1878. Crispispongia pezizoides, Zitt. Studien, III Ab. p. 44.
1826-33. Manon peziza, Goldf. pars, Petref. 1 Th. t. 34. f. 86.
Distribution. Upper Jura: Nattheim, Wirtemberg.
CRISPISPONGIA EXPANSA, Quenst.
1878. Crispispongia expansa, Quenst. Petref. Bd. 5, p. 198, t. 124. £. 38-47.
1878. Crispispongia expansa, Zitt. Studien, III Ab. p. 44.
Distribution. Upper Jura: Nattheim.
DIAPLECTIA.—ELASMOSTOMA. 193
Genus DIAPLECTIA, Hinde, n. g.
Sponges growing in cup-, fan-, or platter-shaped expansions. The walls throughout
consist of the fibrous mesh. No canals are present; the surface on both sides of the
wall exhibits only irregular interspaces between the fibres. The spicular structure
of the fibres resembles that of Sestrostomella.
I propose this genus to include Spongia helvelloides, Lamx., and some other allied
forms which, from their similar mode of growth, have been included by Zittel in the
genus Pharetrospongia, Sollas; but they possess an entirely different spicular structure
from that of the type of this latter genus.
Drapiectia auricuna, Hinde, n. sp. (Plate XXXVI. figs. 4, 4a.)
Sponges forming fan- or ear-shaped expansions. The type specimen is 30 mm. in
height and 50 in width. ‘The wall varies from 5 to 10 mm. in thickness ; the margins
are obtusely rounded. ‘The fibres are from -2 to -4 mm. in thickness; they are made
up of relatively large three- and four-rayed spicules in the axis, and smaller sinuous
spicules bordering the fibre. Minute pitch-fork spicules, like those of Sestrostomella,
are also present.
Distribution. Inferior Oolite (Pea Grit): near Cheltenham. Middle Jura: Couche
a polypiers; Langrune (col/. Tesson).
DIAPLECTIA HELVELLOIDES, Lama. sp. (Plate XXXVI. fig. 5.)
1821. Spongia helvelloides, Lamx. Exp. méthod. p. 87, t. 84. f. 1-3.
1854. Spongia helvelloides, Morris, Cat. Brit. Foss. p. 30.
1878. Pharetrospongia helvelloides, Zitt. Studien, III Ab. p. 46.
Cup-shaped sponges supported on a small cylindrical stem. ‘The walls are from
3 to 5mm. in thickness. The fibres of the outer or under surface of the cup have a
generally vertical disposition; they vary from -14 to -4 mm. in thickness. The
spicular structure resembles that of the preceding species.
This may be distinguished from the preceding species by its form, thinner walls,
and the vertical disposition of the fibres of the outer surface.
Distribution. Middle Jura: Couche a polypiers—Langrune, Ranville, near Caen
(coll. Tesson). Great Oolite: Hampton Cliff, Bath (Morris).
Genus ELASMOSTOMA, From. 1859.
In the typical species of this genus /. acutimargo, F. A. Rem. (=£. frondescens,
From.), the spicular structure of the fibres is of the Sestrostomella type, consisting of
central, relatively large, three- and four-rayed spicules, surrounded by smaller sinuous
irregular spicules, also probably three-rayed. A similar spicular structure is shown
in E. Normanianum, D’Orbig., and E. consobrinum, D’Orbig. sp., as well as in some
2c
194 CALCAREOUS SPONGES.
new species which I have described below. In other forms, however, from the Lower
and Upper Green Sand which agree with the typical species of Elasmostoma in
general form, and in possessing a smooth surface penetrated with large apertures on
one surface of the wall, the spicular structure of the fibres more nearly corresponds
to the Corynella type, and I have therefore separated these into a distinct genus. In
some, if not all, of these latter forms distinct canals are present, whereas, according
to Zittel, canals are absent in typical species of Hlasmostoma.
ELASMOSTOMA ACUTIMARGO, Reem. sp.
1839. Tragos acutimargo, F. A. Roem. Verst. d. Nordd. Oolit. Nachtr. p. 10, t. 17. f£. 26.
1859. Elasmostoma frondescens, From. Introduction, p. 43, t. 3. f. 6, 6a.
1864. Elasmostoma acutimargo, Reem. Pal. Bd. 13, p. 45, t. 1. f. 21.
1878. Manon peziza, Quenst. pars, Petref. Bd. 5, p. 362, t. 182. f. 39.
1878. Elasmostoma acutimargo, Zitt. Studien, III Ab. p. 44.
Sponges growing in small fan-shaped expansions from 23 to 37 mm. in height and
from 24 to 41 mm. in width. The walls are from 2 to 4mm. in thickness. On the
upper or concave surface, though occasionally on the under surface, there is a smooth,
apparently compact, dermal layer perforated with circular or irregular apertures from
1-7 to 8 mm. in diameter. On the opposite surface are shown the reticulated fibres,
with small, irregular apertures between them. ‘The fibres, as seen in a transverse
microscopic section, are from ‘17 to “31mm. in thickness; the spicular structure is
but faintly shown, but it appears to be of the Sestrostomella type.
Distribution. Neocomian, Censeau, Jura: Nogerais; Berklingen, Brunswick.
ExasmostomMa Normanianum, D’Orbigny, sp.
1847. Cupulospongia Normaniana, D’Orbigny, Prodr. de Pal. vol. 11. p. 188.
1840-47. Spongia Peziza, Mich. (non Goldf.) Icon. Zooph. p. 143, t. 36. f. 5.
1864. Elasmostoma Normanianum, F. A. Roem. Pal. Bd. 13, p. 45, t. 16. f. 6.
1871. Elasmostoma Normanianum, Gein. Pal. Bd. 20, p. 36, t. 7. f. 7-12.
1878. Manon peziza macropora, Quenst. Petref. Bd. 5, p. 361, t. 132. f. 35.
1878. Elasmostoma Normanianum, Zitt. Studien, III Ab. p. 44.
1883. Elasmostoma Normanianum, Dun. Pal. Bd. 29, p. 41, t. 38. f. 4, 5.
Sponges fan-shaped, or by the coalescence of the margins becoming obliquely cup-
shaped. The examples vary from 24 to 51 mm. in width, and from 22 to 42 mm. in
height; the walls are from 3 to 5°5 mm. in thickness. One surface of the wall, either
the exterior or interior, is smooth, and perforated with circular apertures from 1 to
2mm. in width, and about the same distance apart from each other; the margins of
the apertures slightly project. The opposite surface only shows the irregular inter-
spaces between the fibres. According to Dunikowski, the fibres exhibit relatively
large axial three-rayed spicules, characteristic of the Sestrostomella type.
ELASMOSTOMA. 195
Owing to the meagre and unsatisfactory definition of this species by Michelin, very
various forms have been assigned to it by different authors, and it is difficult to deter-
mine which of these should properly be included in it. I have applied it to sponges
resembling the forms figured by Geinitz, though I do not agree with this author in
including therein the Manon Peziza figured by Goldfuss in Petref. 1 Th. t. 29. f. 8,
which appears to me to belong to a different species.
Distribution. Upper Green Sand: Warminster. Cenomanian: Kssen.
ELASMOSTOMA CONSOBRINUM, D’Orbigny, sp.
1847. Cupulospongia consobrina, D’Orbigny, Prodr. de Pal. vol. 11. p. 188.
1826-33. Manon peziza, Goldf. pars, Petref. 1 Th. t. 1. f. 7, 8.
1871. Elasmostoma consobrinum, Geinitz, Pal. Bd. 20, p. 38, t. 6. f. 8-10.
1878. Manon peziza, Quenst. pars, Petref. Bd. 5, p. 358, t. 182, f. 26-33.
1878. Elasmostoma consobrinum, Zitt. Studien, III Ab. p. 45.
1883. Elasmostoma consobrinum, Dun. Pal. Bd. 29, p. 41, t. 37. f. 4, 5.
Cup- or fan-shaped sponges with walls from 2 to 4mm. in thickness. One surface
of the wall, usually the outer or convex surface, is penetrated by minute circular
apertures about ‘5 mm. in width; the opposite surface has no special dermal layer,
and merely shows the interspaces between the fibres. The fibre, according to
Dunikowski, is composed mainly of relatively large axial three-rayed spicules.
D’Orbigny’s definition of this species is obviously insufficient for recognition, but
as the name has been applied by later authors to the sponges figured by Goldf. in
Petref. pl. 1, f. 7, 8, I have used it for these forms. As pointed out by Geinitz, the
sponges figured by Goldfuss were derived from Essen, and not, as erroneously stated,
from Maestricht.
Distribution. Upper Green Sand: Warminster. Cenomanian: Essen. Craie
Chloritée: Villers-sur-Mer.
Exasmostoma scrtutum, Hinde, n. sp. (Plate XXXVI. figs. 6, 6a, 6, 6¢, 6d.)
Sponges growing in flattened or slightly convolute fan- or plate-shaped expansions
from 40 to 100 mm. in height, and from 50 to 110 mm. in width. The base is
either expanded or variously curved. The wall is from 4 to 5 mm. in thickness.
The inner or concave surface of the wall has a smooth dermal layer, perforated by
circular apertures about 1:5 mm. in width, and from 1:5 to 3 mm. apart. The
margins of these apertures are thickened and, where best preserved, slightly elevated
above the surface. The surface between the larger apertures is also perforated by
minute irregular puncta. The under surface of the wall is rough; the fibres have
an indistinct radiate arrangement, and are connected by minute angular processes.
In a transverse section the fibres measure from ‘1 to °35 mm. in thickness. ‘The
2c2
196 CALCAREOUS SPONGES.
spicular structure is distinctly of the Sestrostomella type; a single ray of one of the
large axial spicules measures °24 mm.
In its general characters Manon Peziza, Goldf. Petref. 1 Th. t. 29. f. 8, resembles
this species; but the oscular apertures are uniformly single, and they are closer
arranged than in Goldfuss’s form.
Distribution. Upper Chalk: Charlton, Bromley, Kent; Sussex.
ELasmostoMa crassuM, Hinde, n.sp. (Plate XXXVI. fig. 7.)
Sponges growing in irregularly folded expansions; the only specimen is imperfect,
and measures 76 mm. in width and the same in height. ‘The walls are from 6:5 to
10 mm. in thickness; the inner or upper surface is smooth, and is perforated by
circular apertures from 1-6 to 2 mm. in width, and about the same distance apart ;
their margins are smooth and even. Between the larger apertures are smaller
irregular perforations. The under surface resembles that of the last species.
The fibres are from ‘13 to °4 mm. in thickness; in the specimen examined they are
entirely crystalline, and no structure has been preserved.
This species may be readily distinguished from the preceding by the much greater
thickness of the walls and the larger size of the oscular apertures.
Distribution. Upper Chalk: Bromley, Kent.
ExasMostoMa PLicatuM, Hinde, n. sp. (Plate XX XVII. fig. 1.)
Sponges with convolute walls, either fan- or open funnel-shaped. A fairly large
specimen is 95 mm. in width and 57 mm. in height. The walls are 4 mm. in thick-
ness. The inner surface has circular apertures 1 mm. wide, and from 1 to 2mm.
apart, with minutely punctate interspaces. ‘The under surface of the wall resembles
that of EZ. scitulwm, but the fibres are of a more delicate character.
In a thin transverse section the fibres measure about ‘17 mm. in width; their
spicular structure resembles that of H. scitwlum, and they are generally narrower,
and more evenly disposed than in that species.
This species approaches closely to E. scitulum; but the oscular apertures are
smaller, and the fibres are less robust.
Distribution. Craie Chloritée: Cap la Heéve, near Havre.
ELASMOSTOMA SUBPEZIZA, D’ Ord. sp.
1847. Cupulospongia subpeziza, D’Orb. Prodr. de Pal. vol. i. p. 288.
1826-33. Manon peziza, Goldf. pars, Petref. 1 Th. t. 5. f. 1.
1878. Manon peziza, Quenst. pars, Petref. Bd. 5, p. 363, t. 182. f. 42, 43.
1878. Pharetrospongia subpeziza, Zitt. Studien, III Ab. p. 46.
Sponges small, obliquely cup-shaped ; a fairly large example is 19 mm. in height
by 17 in width. The walls are from 2 to 2°5 mm. in thickness. The outer surface
ELASMOSTOMA.—RAPHIDONEMA. 197
is smooth, with minute circular apertures about -3 mm. in width, and from ‘6 to
1mm. apart; the interspaces are perforated with minute puncta. The inner surface
exhibits reticulate fibres so closely disposed that the interspaces can scarcely be
distinguished without a lens.
As seen in a microscopic section, the fibres are from ‘1 to °2 mm. in thickness; the
spicular structure is very indistinct, but it appears to resemble that of E. scitulum.
The form figured by Goldfuss is stated to have been derived from Essen; but, as
Geinitz has pointed out, this is an error, and there is no doubt that it came from
Maestricht, where these small sponges appear to be fairly abundant.
Distribution. Upper Chalk: Maestricht.
Genus RAPHIDONEMA, Hinde, n. g.
Sponges cup- or funnel-shaped, or forming convolute expansions. On one surface
of the wall, and in some forms on both, there is a modified dermal layer, either
compact or minutely porous. The dermal layer, either on the outer or inner surface
of the wall, is perforated with oscular apertures, except in one species in which it
has no oscular apertures, though it extends over the canal apertures of the wall.
Definite canals are usually present ; they penetrate the wall from one or both surfaces,
and in some instances extend quite through it.
The fibres are composed of filiform three-rayed spicules similar to those of Corynella ;
the basal ray of the spicules is but very slightly developed, so that in microscopic
sections they appear for the most part as simple uniaxial forms. Rarely are larger
three-rayed spicules to be seen in the fibres, though such may be present in the
dermal layer, the minute structure of which I have not been able to determine.
As already mentioned, this genus resembles Hlasmostoma in its mode of growth
and general structure, but the spicular constitution of the fibres is of a different
character, and resembles the Corynella type. It also includes some of the species on
which Prof. Sollas has based the genus Catagma, but the characters assigned to this
genus are not sufficiently definite, and its spicular structure, judging from the figures
and descriptions, is very different from that of Raphidonema, and resembles somewhat
that of Elasmostoma.
RaPHIDONEMA contortUM, Hinde, n. sp. (Plate XX XVII. figs. 2, 2a, 26.)
1874. Manon peziza, Davey, Trans. Newbury Field-Club, p. 10.
Sponges growing in convolute expansions ; the folds of the walls frequently coalesce
so as to form intricate masses of considerable size. The sponge is usually attached
by its base, not infrequently to a sponge of a different species. A small example is
29 mm. in height by 42 in width, whilst a fairly large specimen is 65 mm. in height
by 100 in breadth. The walls are from 4 to 6 mm. in thickness. Both surfaces are,
in the best-preserved specimens, smooth, and alike covered with a modified dermal
198 CALCAREOUS SPONGES.
layer of very closely arranged fibres, finer than those of the substance of the wall;
the interspaces between these surface-fibres are hardly visible without a lens. On
one surface of this wall, either the upper or under, are minute circular apertures,
from °5 to 85 mm. in width and from | to 3 diameters apart; these apertures are
connected with sinuous canals, which extend generally at right angles into the wall.
The fibres of the interior of the wall, measured in a thin section, are from ‘2 to
‘4mm. in thickness; the filiform spicules are slender, and closely arranged parallel
to each other in the direction of the fibre and round the margins of the canals.
In its mode of growth, and the size of the oscular apertures, this species corre-
sponds with El/asmostoma consobrinum, D'Orbig.; but besides the differences in the
spicular structure, which of course can only be ascertained in thin sections, the
present species is usually larger and the walls are thicker than in the examples of
E. consobrinum from the Cenomanian at Essen.
Distribution. Lower Green Sand: Farringdon, Berkshire.
RaPHIDONEMA PorcaAtTUM, Sharpe, sp. (Plate XX XVII. fig. 3.)
1854. Manon porcatum, Sharpe, Quart. Journ. Geol. Soe. vol. x. p. 196, t. 5. f. 2.
1874. Manon porcatum, Davey, Trans. Newbury Field-Club, p. 16.
1878. Catagma porcatum, Sollas, Ann. & Mag. Nat. Hist. vol. ii. p. 362.
1883. Catagma porcatum, Keeping, Foss. of Upware &e. p. 147.
Sponges either cup-shaped or growing in irregularly convolute expansions. The
specimens vary from 30 to 70 mm. in height and from 38 to 100 mm. in breadth.
The walls are from 4 to 5-5 mm. in thickness. The upper or inner wall-surface has
a smooth dermal layer penetrated by numerous, irregularly disposed, circular aper-
tures, about ‘85 mm. in width; the interspaces between these are minutely porous.
The under or outer surface of the wall is covered with sharply projecting sinuous
anastomosing ridges; the fibres of this surface in some examples are finer and closer
arranged than those of the interior of the wall, whilst in other specimens there is
but little difference between the surface and the interior fibres.
The interior fibres, as seen in a transverse microscopic section, are from *15 to
-3 mm. in width; the spicular structure resembles that of R. contortum. ‘This species
is characterized by the peculiar sinuous ridges of the under surface.
Sharpe states that the inner surface of this species is pierced by very numerous
ill-defined openings, but the oscular apertures only appear ill-defined when the
- surface is worn; in perfect specimens they are as clearly marked as in the preceding
species.
Distribution. Lower Green Sand: Farringdon; Upware.
RaPHIDONEMA PusTULATUM, Hinde, n.sp. (Plate XXXVI. figs. 8, 8a.)
Sponges either shallow cup- or funnel-shaped, or forming convolute expansions.
RAPHIDONEMA. 199
A small specimen measures 28 mm. in width by 13 in height, whilst a large form is
55 by 42 mm. The walls are from 3°5 to 7 mm. in thickness. One surface of the
wall, generally the upper, has a compact dermal layer, perforated by circular aper-
tures, from 1 to 1°6 mm. in width, with well-defined slightly elevated margins. The
opposite surface is smooth ; the fibres are, in some instances at least, more slender
and closer arranged than those of the interior of the wall.
The fibres, measured in a thin section, vary from ‘1 to ‘35 mm. in width; the
spicular structure is similar to that of R. contortum.
This species appears to have been referred by Sharpe* to Manon marginatum,
Phill.; Davey? has also placed it under the same designation, although he questions
the identity of Phillips’s species from the Upper Chalk with the Farringdon speci-
mens. Mantell{ has named it Tragos peziza, Goldf.
Distribution. Lower Green Sand: Farringdon.
RapPHIDONEMA MACROPORA, Sharpe, sp. (Plate XX XVII. fig. 4.)
1854. Manon macropora, Sharpe, Quart. Journ. Geol. Soc. vol. x. p. 195, t. 5. f. 3, 4.
1848. Chenendopora fungiformis?, Mant. (non Lamx.), Wonders of Geology, p. 637.
1874. Manon macropora, Davey, Trans. Newbury Field-Club, p. 15.
1878. Catayma macropora, Sollas, Ann. & Mag. Nat. Hist. ser. 5, vol. ii. p. 356, f. 1.
1878. Elasmostoma macropora, Zitt. Studien, III Ab. p. 44.
1883. Hlasmostoma acutimargo, Keeping (non Reem.), Fossils of Upware &c. p. 147.
Sponges cup- or funnel-shaped, or forming convolute expansions. A small speci-
men measures 25 mm. in height by 23 in width, whilst a large example is 100 by
110mm. The walls vary from 4 to 10 mm. in thickness. The inner or upper
surface of the wall is either even, or in some specimens is marked by concentric
rounded ridges and shallow furrows. It is furnished with a smooth, apparently
compact, dermal layer, which is perforated by circular or ovate apertures, from 1:5
to 3°5 mm. in width, disposed concentrically nearly in contact with each other.
Where the interior is ridged, the apertures occur in the intermediate furrows. Small
canals open immediately beneath these apertures. The outer surface of the wall is
uneven, sometimes with nodose excrescences ; only the naked fibres are shown.
In a transverse microscopic section of a specimen from Upware, the fibres appear
as closely reticulating bands, from ‘13 to -26 mm. in width. The filiform, sinuous
spicules are more robust and less regularly disposed than in R. contortum. Though
apparently uniaxial, here and there forms can be detected which show the stunted '
basal ray of three-rayed spicules, and it is probable that the majority of the spicules
are irregular three-rayed forms.
* Quart. Journ. Geol. Soc. 1854, vol. x. p. 189.
Trans. Newbury Field-Club, p. 15.
+ Wonders of Geology, 6th ed. 1848, vol. ii. p. 636; Medals of Creation 1854, vol. i. p. 229. f. 1.
200 CALCAREOUS SPONGES.
I have previously compared the structure of this species to that of Sestrostomella*,
but on further examination I find that the fibres of the interior of the wall are built
up of filiform spicules, though it is not unlikely that the dermal layer may be com-
posed of relatively large three- or four-rayed forms.
Distribution. Lower Green Sand: Farringdon; Upware.
RAPHIDONEMA STELLATUM, Goldf. sp.
1826-33. Manon stellatum, Gold. Petref. 1 Th. p. 3, t. 1. f. 9.
1871. Stellispongia Goldfussiana, Gein. Pal. Bd. 20, p. 31, t. 6. f. 4-7.
1878. Manon peziza stellatum, Quenst. Petref. Bd. 5, p. 361, t. 132. f. 34.
1883. Elasmostoma stellatum, Dun. Pal. Bd. 29, p. 40, t. 37. f. 1, 2, 6, and t. 38. f. 1, 2,3.
Sponges growing in thin expansions or funnel-shaped. ‘The walls are from 2 to
nearly 4mm. in thickness. The oscular apertures are about ‘5 mm. wide, and are
either on the upper or under surface of the wall. The fibres bordering the apertures
have a stellate arrangement. The opposite surface of the wall exhibits an open
network of fibres.
As seen in a transverse microscopic section, the fibres are from -12 to -25 mm. in
width ; they are mainly composed of slender, filiform, irregular, three-rayed spicules.
The forms which Dunikowski refers to as “Stabnadeln” appear to me to be more
probably fragments of three-rayed spicules.
The thin walls and stellate character of the oscular surface readily distinguish
this species.
Distribution. Cenomanian: Essen, Rhenish Prussia.
RAPHIDONEMA FARRINGDONENSE, Sharpe, sp. (Plate XX XVII. figs. 5, 5 a, 5d.)
1864. Manon Farringdonensis, Sharpe, Quart. Journ. Geol. Soc. vol. x. p. 196, t. 5. f. 5, 6.
1874. ? Chenendopora funyiformis, Davey, Trans. Newbury Field-Club, p. 18.
1878. Catagma Farringdonense, Sollas, Ann. & Mag. Nat. Hist. vol. 1. p. 362.
1878. Pharetrospongia Farringdonensis, Zitt. Studien, III Ab. p. 46.
Sponges cup- or funnel-shaped, free or attached by a spreading base, generally to
other sponges. Very variable in size; a small individual measures 30 mm. in height
and width, whilst a large specimen is 90 mm. in height by 120 in width at the
summit. The wall varies from 7 to 17 mm. in thickness. The outer surface of the
cup is very uneven, and frequently covered with large nodose excrescences. In some
examples the fibres of this surface are finer and closer arranged than in the interior ;
but, as a rule, there is scarcely any differentiation in their character. The interior of
the cup has a very varying aspect in different specimens. In some examples it is
coarsely fibrous and exhibits ill-defined apertures of numerous canals, about 1 mm.
wide, either irregularly distributed or with a partially horizontal arrangement.
* Annals & Mag. Nat. Hist. 5th ser. vol. x. p. 203.
RAPHIDONEMA.—PHARETROSPONGIA. 201
These canal-apertures are sometimes bridged over by the fibres. In other examples
a smooth, minutely porous dermal layer, consisting of very delicate, closely reticu-
lating fibres, extends to varying heights in the cup, and covers over the coarser fibres
and the canal-apertures. This dermal layer is very variously developed; in some
specimens it is limited to the lower portion of the cup or is entirely absent, whilst
in others it extends, like a thin film, quite to the margin, so that the interior of the
cup is perfectly smooth. The wall itself is perforated by numerous straight or
slightly sinuous canals, which run generally at right angles, but occasionally are
oblique to the surface.
The fibres vary from -]1 to ‘3 mm. in width, and are composed of filiform spicules
similar to those of &. contortum. In some cases the dwarfed basal ray of the three-
rayed spicules can be clearly seen; but, as a rule, this minute ray cannot be detected
in microscopic sections, and thus the spicules appear as if uniaxial. Very rarely is
a perfect spicule exposed in the section ; the entire length of an apparently complete
form is ‘25 mm.
The presence of minute three-rayed spicules in this and other species of this genus
distinctly marks them off from specimens of Pharetrospongia, in which only uniaxial
spicules have hitherto been discovered.
Distribution. Lower Green Sand: Farringdon, Berkshire. Very abundant.
Genus PHARETROSPONGIA, Sollas, 1877.
Professor Sollas has based this genus on the characters of the single species
P. Strahani, which is principally distinguished by the uniaxial form of the spicules
composing its fibres. Prof. Zittel proposed to extend Sollas’s definition so as to
include in the genus a group of cup- or funnel-shaped sponges having a general
resemblance in their mode of growth to P. Strahani. As, however, the spicular
structure of the fibres of some, if not all, of these sponges differs from that of
P. Strahani, it seems to me desirable to limit the genus to sponges which can be
shown to possess a similar spicular structure to that of the type species.
PHARETROSPONGIA STRAHANI, Sollas. (Plate XXXVIL. fig. 6;
Plate XX XVIII. fig. 1.)
1877. Pharetrospongia Strahani, Sollas, Quart. Journ. Geol. Soc. vol. xxxiii. p. 242, t. 11.
1878. Pharetrosponjia Strahani, Zitt. Studien, III Ab. p. 45.
1879. Pharetrospongia Strahani, Carter, Ann. & Mag. Nat. Hist. ser. 5, vol. iv. p. 432.
1882. Pharetrospongia Strahani, Steinm. Neues Jahrb. Bd. 2, p. 129.
Sponges having the form of variously convoluted plates, which sometimes anasto-
mose so as to become funnel-shaped, also occasionally subcylindrical. The walls are
from 7 to 13 mm. in thickness. The sponge does not appear to have been attached.
2D
202 CALCAREOUS SPONGES.
Small cylindrical forms are 47 mm. in length by 15 in thickness, whilst large
expanded examples reach the dimensions of 200 mm. in length and the same in
width. |
The outer surface of the wall is generally uneven and composed of a layer of
reticulate fibres, finer than those of the interior; the interspaces between them are
small, irregular in distribution, and either subangular or circular. The inner surface
of the wall is usually smoother than the outer, the fibres are thicker, and the inter-
spaces larger; in some cases definite circular apertures up to 1:3 mm. in width are
present, but generally the apertures are smaller and irregular in form. No definite
canals can be distinguished in the walls, even where they attain their greatest
thickness. In some instances the fibres have a general disposition in the direction
of the length of the sponge, with transverse connections; but, asa rule, no definite
arrangement is perceptible.
The fibres are from 13 to 55 mm. in thickness; they are composed of minute,
straight, or slightly curved, subcylindrical uniaxial spicules, closely arranged parallel
with each other in the direction of the length of the fibre. The longest of these
spicules which I have succeeded in measuring is ‘14 mm. and its thickness about
‘01 mm.
This species occurs in the Cambridge Green Sand, and it is also very abundant in
the Upper Chalk. The Chalk examples are for the most part larger and the walls are
thicker than in the Green Sand forms, but the differences do not appear to me to be
of specific value. Many of the examples from the Green Sand near Cambridge are
entirely calcareous, and the fibres completely dissolve when treated with acid; in
others the fibres have a thin surface-pellicle of silica and the interior is calcareous.
Attached to the siliceous pellicle are entire and fragmentary spicules which have
been replaced by silica, the spicules of the interior of the fibre retaining their
calcareous composition. A similar replacement has also taken place in some of the
forms from the Upper Chalk. Those which are imbedded in the chalk itself have a
smooth, waxy aspect, and when treated with acid completely dissolve; but not infre-
quently specimens are imbedded in flint, and in these cases the exterior portion of
the fibres becomes siliceous whilst the interior retains its calcareous condition. That
this partial silicification of the fibre is owing to its being surrounded by the flint is
clearly shown by the fact that some sponges are partly imbedded in flint and partly
in chalk, and only the portion in the flint has been replaced by a thin coating of
silica, that in the chalk being wholly calcareous.
By Prof. Sollas and Mr. Carter these sponges are believed to have been originally
siliceous, and, from the form and disposition of the spicules in their fibres, to be
allied to an existing species of Reniera. My reasons for regarding them as originally
calcareous are based upon the fact that in their general characters and structure
they closely resemble other Pharetrones occurring in the same strata, which by
PHARETROSPONGIA.—PACHYTILODIA. 203
=
Mr. Carter (though not by Prof. Sollas) are acknowledged to be calcareous sponges,
whilst their condition is altogether different from that of the undisputed siliceous
sponges in the same beds. As, however, some of the siliceous sponges from the
Cambridge Green Sand are probably derived from other deposits, the dissimilarity
existing between them and the specimens of Pharetrospongia may be alleged to be
owing to the different conditions to which they have respectively been exposed; but
the argument is quite applicable to the examples from the Upper Chalk; and if
there are important differences in the character and mineral structure of examples of
Pharetrospongia, and of undisputed siliceous sponges from the same beds and the
same localities, they may reasonably be supposed to be due to differences in their
original condition. Thus, for example, specimens of the calcareous sponges Elas-
mostoma scitulum and Pharetrospongia Strahani, from the Upper Chalk at Bromley
in Kent, alike retain the perfect form and calcareous structure of their component
fibres; but in the undoubtedly siliceous sponges in the same beds and locality, such
as Ventriculites or Scyphia, the fibres or mesh, instead of being perfectly smooth and
calcareous, as in the first-named forms, have been either completely dissolved, leaving
empty moulds, or else replaced by powdery iron peroxide. It is impossible to
explain these notable differences without supposing that the mineral structure of
these sponges was originally different, and it seems therefore reasonable to conclude
that the forms with the calcareous fibres retain their original constitution. Mr. Carter
readily acknowledges that Elasmostoma scitulum is a calcisponge; and Pharetro-
spongia Strahani so closely resembles it in general structure that, notwithstanding
the different form of its component spicules, .there can hardly be a doubt of its
calcareous origin.
Mistribution. Upper Green Sand: Warminster, near Cambridge. Upper Chalk:
Charlton, Bromley, Broadstairs, Kent; Ditchhampton, near Wilton, Norwich:
Pertwood, near Warminster.
Genus PACHYTILODIA, Zittel, 1878.
PACHYTILODIA INFUNDIBULIFORMIS, Goldf. sp.
1826-33. Scyphia infundibuliformis, Goldf. Petref. 1 Th. p. 12, t. 5. f. 2.
1874. Cupulospongia infundibuliformis, Gein. Pal. Bd. 20, p. 29, t. 4. f. 4, 5.
1878. Scyphia infundibuliformis, Quenst. Petref. Bd. 5, p. 347, t. 132. f. 1-3.
1878. Pachytilodia infundibuliformis, Zitt. Studien, III Ab. p. 46.
1883. Pachytilodia infundibuliformis, Dun. Pal. Bd. 29, p. 42, t. 40. f. 1, 2.
Sponges cup- or vase-shaped, supported on thickened nodose stems, occasionally
forming convolute extensions. An imperfect specimen measures 140 mm. in height
by 170 in width. The walls are from 8 to 25 mm. in thickness; the interspaces
between the fibres are relatively large. ‘The fibres are from ‘4 to ‘9 mm. in thickness :
2D 2
204 CALCAREOUS SPONGES.
according to Dunikowski they are composed of small uniaxial spicules associated
with large three-rayed forms.
Distribution. Upper Green Sand: Warminster. Cenomanian: Essen.
Family SYCONES, Haeckel.
Genus PROTOSYCON, Zitte/, 1878.
Prorosycon PunctaTuM, Goldf.sp. (Plate XX XVIII. figs. 2, 2 ae.)
1826-388. Scyphia punctata, Goldf. Petref. 1 Th. p. 10, t. 3. f. 10.
1858. Scyphia punctata, Quenst. Der Jura, p. 667, t. 81. f. 79.
1870. Scyphia punctata, O. Schmidt, Spongien-Fauna des atlant. Gebietes, p. 20, t. 2,
16 le
1878. Scyphia punctata, Quenst. Petref. p. 333, t. 131. t. 21-27.
1878. Protosycon punctatum, Zitt. Studien, III Ab. p. 48, t. 12. f. 7.
1879. Protosycon punctatum, Zitt. Neues Jahrb. p. 33, t. 2. f. 7.
1883. Protosycon punctatum, Carter, Ann. & Mag. Nat. Hist. ser. 5, vol. xi. p. 34.
Small subcylindrical sponges with truncated summits, and occasionally slightly
expanded bases. The usual dimensions are from 6 to 14 mm. in length, and from
2°5 to 4mm. in thickness; an unusually long form, figured by Quenstedt, measures
24 by 3'5mm. A funnel-shaped cloaca extends quite to the base of the sponge.
The walls are from ‘7 to 1mm. in thickness. The outer surface is apparently quite
smooth, but under a strong lens delicate vertical furrows are visible. In each of
these furrows is a row of minute circular or ovate canal-apertures, and still smaller
circular or subangular apertures are wedged in on the intervening ridges. The
interior surface of the cloaca is not exposed in the specimens which I have examined ;
but according to Quenstedt the canal-apertures are disposed in regular vertical and
horizontal series.
In a thin vertical median section the walls exhibit a series of horizontal canals,
about ‘17 mm. in width, some of which are apparently open throughout, while others
seem to be closed either at one or both ends. ‘These canals are bounded by three-
and four-rayed spicules, and sections of the spicular rays are clearly shown, though
the disposition of the spicules cannot be satisfactorily ascertained from a thin section.
In some instances the rays are arranged singly, whilst in others the rays of two or
three spicules are in close juxtaposition, but they are not apparently united into
fibres as in the Pharetrones. The spicular rays are about ‘04 mm. in thickness; I
have not been able to determine their length.
Distribution. Upper Jura: Streitberg, Franconia.
bo
SS
or
BACTRONELLA.
Incerte sedis.
Genus BACTRONELLA, Hinde, n. g.
Sponges simple, rod- or club-shaped, usually attached by a slightly expanded base.
The lower portion exhibits traces of a wrinkled compact dermal layer; the rest of the
surface is smooth and covered with minute circular and subangular apertures, which
appear to be bounded by a delicate open dermal layer of three- or four-rayed spicules.
The sponge appears to be throughout composed of spicules, not arranged in fibres,
but forming an open tissue with irregular interspaces. The exterior portion is
penetrated by definite cylindrical canals which can be traced for a short distance
into the interior, though they do not appear to extend into the central portion of
the sponge. The spicules are so closely intermingled in the exterior portion that
only circular sections of the rays, and rarely a portion of a conical ray, can be distin-
guished; but in the central portion the spicules are less intricately associated, and
definite three- and probably four-rayed forms can be distinguished. The rays of
some of these spicules are clearly microspined. Mingled with the definite spicules
are smaller forms which appear to be of an irregular character, though the condition
of the specimen prevents accurate determination. In transverse sections the interior
portion of the sponge is partially filled with a dark earthy matrix, and appears to
possess a different structure to the exterior, but this is probably owing to the more
compact agglomeration of the spicules near the surface.
This genus is based on the characters of the small Jurassic sponges which form
part of Goldfuss’s species Ceriopora clavata, Petref. 1 Th. p. 36, t. 10. f. 15 c—f (non
f. a, 6). The minute structure of these forms can only partially be made out in
thin microscopic sections, but sufficient is shown to indicate that it materially differs
from that of the fossil Calcareous sponges already described. ‘The absence of a fibrous
arrangement of the spicules distinguishes it from the Pharetrones ; whilst the structure
of the wall and the absence of a central cloaca separates it from the Sycones. ‘The
disposition of the spicules more nearly resembles that of the recent Lewcones; but in
none of the existing examples of this family are the three-rayed spicules microspined.
As the genus cannot suitably be included in any of the present divisions of the Calca-
reous sponges, I propose to leave it in an indefinite position until its characters are
more clearly ascertained.
BACTRONELLA PUSILLUM, Hinde, n. sp. (Plate XX XVIII. figs. 3, 5 a-q.)
1826-33. Ceriopora clavata, Goldf. pars, Petref. 1 Th. p. 36, t. 10. f. 15 c-f (non f. a, 6).
1858. Ceriopora clavata, Quenst. Der Jura, p. 665, t. 81. f. 59, 60.
1881. Ceriopora clavata, Quenst. Petref. Bd. 6, p. 245, t. 152. f. 63-88.
Subcylindrical or slightly inflated sponges, with conical summits, varying from 3°5
206 CALCAREOUS SPONGES.
to 10 mm. in length and from 2 to 3°5 mm. in thickness. The canals opening on the
outer surface are from ‘18 to’2 mm. in width; they can only be traced for a distance
of about ‘5 mm., where they appear to terminate in the interior spicular tissues. The
spicular rays are conical, acutely pointed, and occasionally with minutely spinous
surfaces. ‘The longest measured is‘14mm. In cross section the rays are circular,
with a maximum thickness of (085mm. ‘The specimens throughout are calcareous.
Distribution. Upper Jura: probably from Thurnau, Bavaria.
207
SUPPLEMENT.
Srnce the first part of the work was in type fresh additions have been made to the
Museum Collection, including some new species, the descriptions of which, as they
could not be inserted in their proper systematic position, are here appended.
Division I. SILICEOUS SPONGES.
Order MONA CTINELLID 4.
Genus HAPLISTION, Young and Young, 1877.
Hapuistion rractum, Hinde, n. sp. (Plate XX XVIII. figs. 4, 4a.)
In the fine collection of detached sponge-spicules which Mr. James Bennie very
generously forwarded to me for examination, I found several small fragments of a
fibrous sponge, resembling, as regards their siliceous condition, the spicules of the
Hexactinellid sponges associated with them. ‘The largest of these fragments only
measures 5 mm. in length and thickness; it consists of irregularly anastomosing
cylindrical fibres from -14 to *5 mm. in thickness, which form an uneven meshwork
with circular or elongated interspaces about ‘6 mm. in width. The fibres appear in
fractured surfaces to be solid throughout, and the interior portion only exhibits a
fine granular structure. The outer surface, on the other hand, is covered with a
layer of simple spicules, which, though generally arranged in the direction of the fibre,
are not parallel with each other. The spicules, when examined under the microscope,
present uneven, ill-defined outlines, but they appear to have the form of fusiform
acerates. The longest measured is 32 mm. in length by ‘03 mm. in width. Whether
their surfaces were primarily smooth or spinous it is impossible to determine.
There can scarcely be a doubt that the fibres were originally built up throughout
of spicules similar to those which are now exposed on their exterior surfaces, and
that by fossilization the interior spicules have been undistinguishably fused together.
Their resemblance in mineral condition to the spicules of undoubted siliceous sponges
in the same deposit points to the conclusion that they were also originally siliceous.
208 SUPPLEMENT.
I have placed these fragments with some doubt under the genus Haplistion, for
the type of the genus, H/. Armstrongi, Y. and Y.*, though possessing a fibrous
structure generally similar to these fragments, does not show any traces of spicules,
and its minute structure is therefore at present doubtful. Through the kindness of
Mr. John Young I have been able to examine a fragment of H. Armstrongi; and its
fibres, independently of the absence of the spicules, appear to me to be of a more
robust character than those of the present species.
Mr. H. J. Carter has also described a sponge from the same deposits under the
name of Raphidistia vermiculata}, which is probably allied to the present species.
Judging from the description and figures, 2. vermiculata is a fibrous sponge composed
of fusiform acerates of a sinuous form. Mr. Carter, however, regards the fibres as
belonging to a species of Hydractinia, and the sponge-spicules as parasitic on the
fibres. This view appears to me to be untenable; and Mr. Carter himself suggests
that the fibres might probably belong to the sponge. The sinuous form of the
spicules distinguishes this species from HH. fractum.
Distribution. Lower Carboniferous: Law Quarry, Dalry, Ayrshire. Collected by
Mr. James Bennie.
Order TETRACTINELLIDA, O. Schmidt.
Genus GEODIA, Lame.
Geopia? antigua, Hinde, n. sp. (Plate XX XVIII. tigs. 5, 5 a-e.)
Detached trifid spicules, with elongated cylindrical or subcylindrical shafts, from
‘2 to*3 mm. in width, and with simple, short, obtusely-pointed head-rays, which
project upwards at an acute angle with the head of the shaft. The shafts are all
incomplete; the longest preserved is 2mm. in length, and it is of the same thick-
ness at the end as at the summit. Associated with the trifid spicules are curved
acerates of somewhat similar dimensions, which may probably belong to the same
species. The trifid spicules are distinguished by the relatively short and obtuse
head-rays and the regular thickness of the shafts.
In the same deposit there are fragments of relatively large spicules, with robust
cylindrical shafts from -6 to ‘9mm. in thickness, and two short, stout, obtusely-
pointed opposite head-rays at the summit. The shafts are all incomplete; the
longest fragment measures 3mm. in length. These abnormal spicules may perhaps
pertain to a distinct species from the smaller forms.
These spicules occur in the same beds of decayed chert of Carboniferous age which
* Annals & Mag. Nat. Hist. ser. 4, vol. xx. p. 428, t. 15. f. 31-37.
7+ Annals & Mag. Nat. Hist. ser. 5, vol. i. p. 140, t. 9. f. 15-19.
SILICEOUS SPONGES. 209
have yielded such numerous spicules of Hexactinellid spicules of the genera Hyalostelia
and Holasterella, and they closely resemble these spicules in their condition of pre-
servation. Seen by reflected light they have a dull white aspect; when mounted in
Canada balsam, and examined by transmitted light, they are nearly entirely opaque.
Their surfaces are generally smooth, but in places they are pitted with minute rhom-
bohedral cavities. In comparison with the Hexactinellid forms they are extremely
rare.
Distribution. Lower Carboniferous: Law Quarry, Dalry, Ayrshire. From the
collection of Mr. James Bennie.
Genus PACHASTRELLA, O. Schmidt.
PACHASTRELLA veTuSTA, Hinde, n. sp. (Plate XX XVIII. figs. 6, 6a-f.)
Detached four-rayed spicules; three of the rays either nearly horizontal or forming
the outlines of a three-sided pyramid; the fourth ray is vertical to the other three,
and forms the apex of the pyramid; in rare instances the vertical ray is continued
below as well as above the pyramidal apex, and the spicule becomes five-rayed. The
rays are straight or slightly curved, cylindrical, or very gradually tapering to the
extremity, which is usually obtusely rounded, though in one instance the rays termi-
nate in minute digitations. Asa rule, the rays in each spicule are unequal in length,
the vertical ray being usually the shortest. ‘The spicules vary greatly in size; ina
large form the rays reach to 2°6 mm. in length by ‘47 mm. in thickness, whilst the
rays in a small spicule are not more than ‘54 mm. in length by ‘16 in thickness.
The close resemblance in the general characters of these spicules to those of the
Cretaceous species of Pachastrella, indicates that this genus dates back to the Carbo-
niferous period*. As a rule, the spicules are larger than in the forms from the
Upper Chalk. They occur under the same conditions as the forms described above,
and are comparatively rare.
Distribution. Lower Carboniferous: Law Quarry, Dalry, Ayrshire. From the
collection of Mr. James Bennie.
* The late Mr. Charles Moore, F.G.S., described and figured under the name of Grantia antiqua (Quart.
Journ. Geol. Soe. vol. xxiii. 1867, p. 538, pl. xvi. f. 33, 34) some detached triradiate calcareous spicules from
the Liassic Conglomerates at Brocastle, South Wales. It is very doubtful if these forms were originally
calcareous ; they may probably have been siliceous spicules allied to Pachastrella, the structure of which has
been replaced by calcite.
210 SUPPLEMENT.
Order LITHISTID &.
Family MEGAMORINA.
Genus DORYDERMA, Zittel.
DoryDERMA DaLRYENSE, Hinde, n. sp. (Plate XX XVIII. figs. 7, 7a-q.)
Detached spicules, usually more or less curved and irregularly branching; the
branches are cylindrical in section, and either obtusely rounded at their terminations
or more frequently expanded transversely, and concave so as to be adapted for
clasping the surface of adjoining spicules. A fairly large spicule is 1-1 mm. in
length and ‘18 mm. in thickness. The general resemblance in form and in the
termination of the branches of these spicules to those which form the skeleton of
the Cretaceous genera Doryderma, Pachypoterion, and Heterostinia, and of the
existing genus Lyidium is sufficiently close to prove that sponges with this type
of spicular structure existed as far back as the Carboniferous epoch.
Mr. Carter * has figured spicules of a similar character, but smaller than those from
Dalry, from strata of corresponding age at Ben Bulbul, near Sligo, and has referred
them as “in all probability surface-spicules like those of Corallistes aculeata.” Not-
withstanding the difference in size, these Irish forms may probably belong to the
same species as those from Dalry.
Distribution. Lower Carboniferous: Law Quarry, Dalry, Ayrshire. From the
collection of Mr. James Bennie. Ben Bulbul, near Sligo, Ireland. Presented by
Mr. J. Wright of Belfast.
Order HEX ACTINELLID &.
Family EURETID 2.
Genus SPORADOPYLE, Zittel.
SPORADOPYLE SANTANDERI, Hinde, n. sp. (Plate XX XVIII. figs. 8, 8a-d.)
The only example of this species is a fragment, 55 mm. in length, of a dichoto-
mously branching sponge. The branches or stems are subcylindrical tubes about
11mm. in diameter; the walls are from 2°5 to 3mm. in thickness; the cloaca
appears to be cylindrical, and continuous throughout the length of the sponge.
The outer surface exhibits depressed circular or ovate apertures of blind canals
about 1 mm. in width, which have a generally vertical arrangement, though not in
* Ann. & Mag. Nat. Hist. ser. 5, vol. vi. (1880) p. 212, t. 14B. f. 10, 11.
SILICEOUS SPONGES. 211
very distinct rows; the interspaces are rounded, and from 1 to 1:8 mm. in width.
The canal-apertures are in some cases partially bridged over by extensions of the
dermal layer. I have not been able to ascertain the characters of the interior or
cloacal surface.
The dermal layer is composed of spicules having a stellate disposition of the rays
so as to leave minute circular or irregular interspaces. The interior mesh is regular ;
the spicules are about ‘04 mm. in thickness, and the distance between the nodes is
about ‘25mm. The nodes are compact.
The regular mode of growth and the arrangement of the canal-apertures readily
distinguish this from the only other branching species of this genus, S. ramosus,
Quenst. sp.
The unique specimen is partially imbedded in a calcareous matrix; it retains its
siliceous structure in beautiful preservation.
Distribution. Neocomian: Santander, Spain.
TABULAR LIST OF SPECIES,
ARRANGED IN ZOOLOGICAL SEQUENCE.
The species marked by an asterisk to the left occur in British strata, whilst the asterisks in the
right-hand columns indicate the geological range of each form.
| *Spongilla Purbeckensis, Young
| *Geodia? Wrightii, Hinde
| *Geodia? antiqua, Hinde
Division I. SILICEOUS SPONGES.
Order MONACTINELLID &, Zttel.
Climacospongia radiata, Hinde
Lasiocladia compressa, Hinde
#Reniera) Oarteria Hande... e122. sesee cnn
*Haplistion fractum, UROL C MR eo ene te-, lIAE
*Dirrhopalum planum, Hinde ..............
* Acanthoraphis intertextus, Hinde
*Cliona cretacea, Portlock, sp.
*Cliona glomerata, Morris, sp. .............. oo wai = s\|le 6. ai[le ec lene sill}e « « «\lfe: stm ialfone weil leer
*Cliona? Mantelli, Wetherell, sp. ............ Shulevell oe ohel| Seca dental |e ersectl levevres ell Perereset | Poveneseel | emememet| eae
LAO jie aco penta UME tao On ou oOds omoa Ge Poocloweulleoccilococibo sols collec o0llbc solo ono
Order TETRACTINELLID &, Marshall.
*Ophiraphidites anastomans, Hinde
*Tethyopsis cretaceus, Hinde
*Stelleta inclusa, Hinde..................-
*Geodia? clavata, Hinde
*Geodia? coronata, Hinde
PAL OZOIC.
MESOZOIC.
KAINOZOIC.
Cambrian.
Ordoyian.
Silurian.
Devonian.
Carboniferous.
Triassic.
Jurassic.
CreracEous.
|
Neocomian or
Lower Green Sand.
Cenomanian or
Upper Green Sand and
Chloritic Marl.
Grey Chalk, Chalk Marl,
and Lower Chalk.
Gault.
* KKK KH
| Upper Ohalk.
Eocene.
| Miocene and Pliocene.
TABULAR LIST OF SPECIES.
2
a
18
PALASOZOIC. MESOZOIC. KAINOZOIC.
CRETACEOUS.
os g cr o
AE Seq |as 3
ie gaS/35 Pa
a 33 ‘Bg gio 3 8 5 Ay
write bee a6 Bee |45| ‘3 5
FI 5 A a | = s [28 gO2/S8H1 5 B
S/E| e/a] ee] 2 (2b| « |Sesl/Og| , | gs 5
g 3 Bs ° 2 Ks a 2 || Se! Qa Ba 5, ry Ss)
s|#|a | 2 | 8 |e | 8 Al a Si || 2 ai 8 aE}
On| Ola | a So feels Gi Pe iicce iets a a
‘MUNG TGE., SDS 6 6.0.0 kee Ci RED OO CEC or Ie ioe sil iebcetetcustesned lace srettsret ere lmuat alte ce o a arate lever tao) (2 b6 c *
Pachastrella primeeva, Zitt..............05- sovelscacleocslesoo|laone||lapblolla tard Sood ac da| acts on|lon.c x
Pachantrellarconvolutaaelanderasn.'-//4 ss sda to al ste talistererelas orci|ieiacellls ag aci[eo eci[le@ «le rorosei iors Wa|lelalm « *
*Pachastrella plana, Hinde ..............6. Pao [get coal etext allecelledrltseeies ee otee oe =) ~¥el|agciee *
*Pachastrella vetusta, Hinde..............-.|- *
Order LITHISTID &, O. Schmidt.
Family RHIZOMORINA, Zittel.
Cnemidiastrum stellatum, Goldf. sp. ........|..-. Sacile ci ollpeees tees eoll
Cnemidiastrum striato-punctatum, Goldf. sp...J....|....)....J..-s{....f]...-] x
Cnemidiastrum corallinum, Quenst. sp........ Senelles Boltsor elle o.cs bee ltecaiol eke
Cnemidiastrumebohener eer Zitt., «hoe cce ee \os celles colleen soelleeeclleesa| oF
Cnemidiastrum rimulosum, Groldf. sp......... 585 oenaes loool crake [eae | sence xs
Cnemidiastrum pluristellatum, Zit. ......../..-.|.... ee esee se es sr| es ell ae
Corallidium diceratinum, Quenst. sp. ........ aah esl eisnstall lee opet| ate) Z5cel|fena ail Rane a eco ailtadedanel||ls me oe
Platychonia auriformis, Quenst. sp........... Pa Sah dos aval Poder oh cll scores lee onsl|[ekoxee off
Platychonia vagans, Quenst. sp. ............ gerd etebeay cal fo ate all ache lvace eet | ep cael RIE
Hi @nonte, Ss ocosodpseato+ convo cocuud|leuad|les *
illolimiin jpplbarnnm, eas. ao0r doooeedellanee|ooo5|\aoeuleooo||\so05|||suclledocl|sona| oro) |uoct ol loo *%
Choniellaphen wise losntiBPsth cls ome chen t cate ots s Stes < alleletetelflsier evel swt stall's eters Dros ett [ae beeen ean ite * |
Chonella auriformis, Rem. sp...........++.. BRE otic tal (obec neni |ks.cracon | acura Sacral oesca.o| eco [eeeemea oy acter se |
*Seliscothon planus, Phill. sp. .............. epee | aeace ll fe | Forece|totewee | | rere [sree [iccenetey| Meemcnel ict eter raelllawanecs * | |
*heliscothoniexplanatus, Hem. 8p: «2460 ccles- =e selees ol... «le. Bone (asereacs ecto (a cece | ee eae Rnd x |
Seliscothon Mantelli, Goldf. sp. ............ cabs, Senate Steele esl: Bae faz a Mae *
Seliscothon testaflorum, Quenst. sp.........-- Pe eh sel le tanalieree tall loeram ah ita sos lees Se [teneueta |ietotena | ce eee eval een *
Seliscothon giganteus, Ram. sp...........-- ThE ee lteodetall fevered asseer al eRan ace [taes ws ciltoeareed tei rece herareterl™ © jae
NOMBCOLRONN Sai asks Lies ca aetiteer en ete cs sb gollocen|eamollocon|sacbiloo aolsoud|ne od poolaue unalloocc *
*Chenendopora fungiformis, Zama. .......... BF cee |e ale oecvel[egee oul etn apalltareellsta afl ltarstetep oeat seal a *
*Chenendopora Michelinii, Hinde............|..-- 5 real bearer 16-4. a0 | create eeeeed | ection Renee lama
Chenendopora pocillum, Mich............... obi c|laeopllnaes|(SAooltotac of|b orale dial Se mallccwral le ae
Verruculina seriatopora, Ram. sp. .......... 5 oe [8 ao lcatecallereve oll: arora (6 Gace a eters (eens acon mets thal aes * | |
“Wenning, iia, HOR aoc coeeanoesped|ooadl|os -ollsobd|loees|toclan|||s bers ppese arma loool scorn) {no or x |
SVierruculina astrseay Heder trie) e-cieise «ills « - 4 1:6 |Sboeeelleroce ol [o.6/6781 laleeckel haeteke! (eeconee| acntveN ses own, cl |didso *
*Verruculina convoluta, Quenst. sp. ........-- ppc) ester eed ee ll tee sch ae hotel [ton ectat eg: s\[for'stolas|tatataall aetcene etel| uote * |
Se Vertoaullunapis tn lOsay panel en vat tects ial. lta ai|er 21al| biel ctal| Sette ct-t-tlle eell|> 2 12 |)e-..>|'tets aif ators aoe 2 ally erer * |
*Verruculina miliaris, Hinde ss... 0.0.2.2 ee eleeee Lae H SrePeN ell icotet ected ollteh a g-atl aca SIA see ftetetcte) ar rere | emcee |
*Verruculina Reussii, M/‘Coy, sp. ............ stioillaced tifed00ocee ome] Bonellsoco) (ooo dloconlrosoo cto as * |
Verruculina macrommata, wm. sp. ........ ee lb ceil OGL eee (emer oneh | Cece (ee octet oth ail eubreaa foesteaomas aida * |
214 TABULAR LIST OF SPECIES.
PALEOZOIC. MESOZOIC.
Cretaceous. |
| |
ae Was)
SM eee |
58 aeai4s| |
ae Sua aS) |
a 858 Seago) 5! cal
5 = S| uate |
E B 3S sé SlaS| 3
tee alee ee eek ts: Seka goa|an| 5
Bolce a ca) alte ggelie | elo |O'S | eal ae
else Ba) i aM Tams | AS iment eel ee | Ree gs
s/é/a/a/é]a| 2 2 | SOR Se
*Verruculina papillata, Hinde .............. FoSdldoaalleadallesealoooollea gullacodllonacilos Sallaocdcd|leaco *
Stichophyma turbinata, Ram. sp. ........-- Be a| |b aael Pastel acco aro! tacdliio cals otclo4.onlleome aol oar *
*Stichophyma tumida, Anew oe Ar ee a faite el |lejetievellis: te te [el] o)e:.bilei|fexonoKel|||s eto pte|| (ettednitell iniialtetie\|te ceire\rsd |/o0allsocclaopaoolloaoc * ||
| Jereica polystoma, Ram. sp. ...... +++ eee poad|laoeadllora alacoollocos||aaolsoo coc oalle>oclloagedoloncs *
aviereicalclava, eee. Spracweier eit a ctepe elie ele Sooolsooulosoel|aooullsa col lococllaocoloussioouslanoo goon *
| *Jereica cylindrica, Hinde..........-..+-+.- Scoolled odes malo polrocul||o.oan) ton clloo nallocaa|\e =<
\WCoslocoryphasap: trent es fencnben sec ec efee ce lee ce fee eee ec eilee cefeeeelee selene e] #
| *Scytalia radiciformis, Phill. sp. ........---- eae Ara eee aera tered sl {cecal larcscokalleemeted IER A ea leEO ote *
| *Scytalia fastigiata, Lee, sp. ............--.. doooleop alloca ladec|loocllloccollacccllocoolltcica| poo os) aoe *
*Scytalia terebrata, Phill. sp. WA NE Ee! Fata noted load (sro. babckl||oo-w.alloo.oolloon.olleats.c teed ciealfo'o -| x
| *Stachyspongia spica, Ram. sp. .........--- aHudloconlloomoladacllooodl|lococ|4o 15) Aoosloa.calleogso- * | *
*Pachinion scriptum, Rem. sp..........-..-- Fooo|lodon||eabolloM coll>acclisooulcocolascollaccalasop o4o0 ce *
{Family MEGAMORINA, Zittel.
Placonella perforata, Hinde................ eee) echt laren koi crol|ioro oral llaa-aea|f se
*Doryderma dichotomum, Benett, sp. ........ Po Ee a alako a odalloooollac.colleooslamenteeamall, <2
*Doryderma ramosum, Mant. sp. ............ Do oo|lescolleas os 500)lo 0 Gallas os oe *
| *Doryderma Roemeri, Hinde ................ ree (meee Pane ley al leery dic.arc | bacec ol arated tere (cron ec.cl [Sseors * ||
*Doryderma Benethi, Hinde ............... Sod echoes laarea| lotsa clio allio oc *
*Doryderma Dalryense, Hinde ......:....... FOO a|Taallsoea||yoc}| <3
*Holodictyon capitatum, Hinde.............. eS mea (eee lato. dl ieee pall 3.003 ol oe eet loomed fie 6 |
_*Pachypoterion robustum, Hinde ............ Bees (eae ete yal so". 0-00) rd epee eee ce? |
| *Pachypoterion compactum, Hinde .......... FOP olleoe alles boo l|a oA slllvoc aellb * |
*Heterostinia obliqua, Beneté .............. Bee oct eters ova alloca! lo dois lo oneal mrkein parcrallaa o-oo. o)laa.7c * ||
*Nematinion calyculum, Hinde.............. oO (so aoe alles 66 |(no olla oc |lo ices *%
Carterella cylindrica, Zté. ................ ta yiteere esaices) [bisrorollc, “aral lororeralloronmtel| tears leno exe * |
Carterella spiculigera, Ht@m. sp. ............ | eee ere tocol iar Alla cect eae loeeeel Ieee eral loon c|lo ga: *
| *Isoraphimialtexta, seam. Sp. (0 .0....2-+-+- Bre (es ese | tore eae | ners eee a eeeeal lb etre dtr ons.d Sdllo 3:00 * |
Family ANOMOCLADINA, Zittel.
| Cylindrophyma milleporatum, Goldf. sp. ....|....J....|..--[e-..[oeeeffee ee] x
| Melonella radiata, Quenst. sp............-.- Beto itor tod (aaa aocalllaaaolt < |
| Lecanella paterseformis, Zitt. .............. peg aaa scoollaciaallsaiaial| sacto| odo olla cacllore cullo'sno.o0||o pion « ||
*Mastosia neocomiensis, Hinde.............. ont eae arco oollsacolllacacl|oc-o olf c=
i) duincdiamtibrosa,shcen. Spier ett tates res oo oallesoal| ez
Family TETRACLADINA, Zittel.
Aulocopium cylindraceum, Ram. .......... aacoloaooll €
*Phymatella intumescens, Ram. sp........-.. i 0.93 atO | eroeckGl Sea al loo: all [choral aow-alld oteel laaese lage. o|[a0c> *
| Phymatella heteropora, Ram.sp. .......... Jodalloo adllaogollecwallaooclosog|ao+5][a5 calles. slag cacollao4 &
*Phymatella reticulata, Hinde .............. 55.00l[n0 Gallaaqollance oalloo-cdlladeallo ehre|laacalloo0>|o0 +2 x
*Phymatella nodosa, Hinde 2... s2.- ss-- eee B evol lool Ora eel leona lota | (acto ans leuceeacd ereea| Rea
miebymatellasisps i: sare asccaietes see eee basdllearc|aeed|locnuloocollaseallsscqlp amd|gccolac0soq|laa05 *
KATINOZOIC.
Miocene and Pliocene.
TABULAR LIST OF SPECIES. 215
a
PAL.ZOZOIC. MESOZOIC. KAINOZOIC.
CRETACEOUS.
came
q sex (24 5
Sa ag5 (26 E
aq “aR | =
a 28 Ba elOs] u ie
. S ae gee l/4El 3 =z
ee | he Mme a se | Wee an ests Z62|sH| 4 a
oe 3 = a “4 3 c) oo Oo, 4 . oO
S(/E| 8/2) 8) a] 2 a8) /soee a | 8 g
Sele eel eee ica Go| wailcael) Mermipceen ee 28 aie ae
So on lre | eel ber [is ome 1S |e =
Aulaxinia sulcifera, Ram. sp............... Sonata s.colean ato ago] |e on cleans (ise ome cl oso tooo | [ox 0.c x
*Aulaxinia costata, Hinde... ........6see008 fd Bel heseihs cll ose oes each Or eeel (ec eel lake ol fae kate keer idl lo.c, cic *
Callopegma Schlénbachi, Zitt............... 98 Hallo nollen 6 doodle a ullas Se| to od|obeo eoue| oobo.cd|loc cc K
*Callopegma obconicum, Hinde.............. Fora d| looks olor o|forn'6 allosg cee lloreeal eenceel Iniekee Ie col eeiciso.a||o.r. oe *
*Callopegma ficoideum, Hinde ............../----|----|--- Peete Pe care | Sucre | Sie .cvell crew eeel| ch sshd | een steven eter ot *
*Trachysycon nodosum, Hinde ..............|.--- *
*Trachysycon suleatum, Hinde .............. Veal eactallece cl beta ola lta aaliboro ol eee eo acetic! lean c | [ar oc *
Siphonuiarpinttormis, Goldf. 5.20.20 2. -- 5 Oita lnxmecl lo eaclloe onl a erea ieee era latent | |e
“ASiplr@nin, nll, Aus on ao ede os B0ee BDUOBE B rollers Seecevel lo cue fa Ge | ceemeied leroo.a fp earl used Mies
Siphonia incrassata, Goldf. ................ By ah re eee ep eae eer Al eos cell meee alltcaspoylt es, © cI ae
“Wiligie wie Cueie eAdoepenedonesuedeee Eel cents Bester |S eaceol | Siento leeeteeel (eroeeo| auton premiere ee
*Siphonia Konigi, Mant. sp................. sonoleocbianoulssonlseimallladaslloe colloodelloccclloodoantocs
“ASA eONEy, F\Ooe coe oD DPBS OOOO DUE ODO ree Sobalooeallbece snes |aamalllamed> eco loos owollyeme.om rome
PHallinkoarcostata, WAMG ss. noses +e Soe |b Goa loreal to oe ol lorencee| llotogercl osc Bool eernell |e
*Hallirhoa costata, var. brevicostata, Mich... .. arora llero ool a ehoee| irc cle ces | eames Geos (Oineacl stowhel | a3
Hallirhoa costata, var. Tessonis, Mich. ...... *
*Hallirhoa costata, var. elevata, Hinde........ So éllovoalle sn oiom0 o|[b ote) ||eto aol [oar bcrac|[aos.oll = £4
*Hallirhoa agariciformis, Benett, sp........... ea S| levee eeaeeen Meso at Srenots! | loaned niecancl eters otolevel| ake
Jereaspyritormis, Lames 25.6.6 cee eee afer (i cnceel (eee ee eee fe ects [loatitcen betergial (ee. Crosorcl aune 2
*Jerea Websteri, Sowerby, sp. .........--+4- Pe ee ell dereiel [ns sll ees stall oc2-i| aoisel| estate llaatera| a ose
A VereakrenlCulatan wel sce =a s\n cree nine. 5 Obl Leese ln tecea |etee-cl In oterellipioka & *?| x
iereanQuenshedulay Atria sis) ta cisteversieisicysce visio “a bello po ollota loaselle-ccal lamer Sescollomrenlltacollo colon *
lena @ouibironnts) JEW oo o0 06 66 ob Ou bo Cob U|enee| |eood lacie) Serbs Spree | beeactel arriots||oicid) |jomreenl cabo orc *
CRD, Oa Widen san oooon Go oben do pollen eel lo clos| etercol ome! eae lio cacao acta loscac.d| a cron a coma acco
*Nelumbia tuberosa, Hinde .............--)---- Ex Sete Pe lead rol ke hao ol (okeuctcta © *
*Polyjerea arbuscula, Hinde................ * *
*Polyjerea lobata, Hinde ...............++. * *
*Bolospongia globata, Hinde................|.+-- ale oes
*Bolospongia constricta, Hunde ...... 10... c5 cece efe eee lee safes eens effec ec fee selene alee cefes eeu ale ee.
*Thecosiphonia nobilis, Ram. sp. ..........-. Bsere ell een ee a tehcwell zaxcest| arepee | nce aalicpnis al remenete ll nce ltaxeras tes ahs
@Miecosiphoniay unpiiatas) Ue one 2m ee eee llinieteiel|eon er =|erm tele etetelll esse llleperaic|[n ss a\lleRer=te[aw ee ellerw ere
(Chikainehnine, wis) Aki epee peoooenh oo colocor|soau|oavel|=s al boone |2o%o|onido boss jpeacamo
*[Tnronia variabilis, Mich...........-..0+ee- real eevee le neta Ste cel leeeea | levee end Ss capo Cetera ERB eee (onoute Edict [eerie
*Kalpinella paterseformis, Hinde ............|..-. *
*Kalpinella rugosa, Hinde..........+.:.+65 Re ctite | Bien seciiecal ecsccenl| "eta eal| heen tag a ess *
*Thamnospongia glabra, Hinde...........-.. See leper aaer eal [teene| eee ste steraccl focehiaral fee. |leteie elllayes hereto lovee
*Thamnospongia clavellata, Benett, sp......... Atel ko dig loca Kase Bie | eee en lores ey cee | eta (eacaaene rear cert | Aco
*Thamnosponpia reticulata, Hinde 2.2... .[es cess fee c eee ee fer eee ae fee selec ecloe calecse ee *
—Pnadligealh aka, Gi@aiguiak, JE 6s enep oo olluo colleobe|loo cols oo-dl lec 6 cl lio-cgo 6 Oaete| jaickor [exec | coroeetag| |p cic
“Dighitlheka, aaaatiey, MWe - occ cece Gobo bole so clog 6 ollotse ee bee esl lie eae Sesei oloedi hacker [oroeuioho| occa
InocaGhinie, mmngsp, Jigs Bt soooboeugeebeuleo edjoooclocballeaoojen ar
*Ragadinia compressa, Hinde .............. S iolal eicao Bl (o's 0 claw On |feta-c.h) ||aeretel | oyoto| eke occa) oboe orl |4 Oe
*Ragadinia suleata, Hinde.................. oc ciello co allote-etol eed ba |lescerct| Eroren Permes pacnta la 00foccaobicrc! low, Asc
*Ragadinia clavata, Hinde .......2...+..::|..%- > Sollow 0-6 lena c5| (aco 6 el lle] Roos laos | fo. oval lose/o.0 0] |t.0.00
*Plinthosella squamosa, Zitt................. spoulige bellacienl ecko aces | Sota eet loved [bcc fetceera tcl fors0.c
=P ritiosella compacts Lien wee teaiee terete evelciele slalelelae ee llleererellere otells acini ells ae «lenin alielawe
Hhaiinosallle, wepise, Ji ok bn ee oancon ce sslloe oc|boe Esso) /2016.0| Sold eo 0 bolo jo ridio| [om om) jor omonl nce
*Plinthosella convoluta, Hinde
*Phymaplectia irregularis, Hinde
*Phymaplectia spinosa, Hinde
*Phymaplectia cribrata, Hinde
*Phymaplectia scitula, Hinde
*Rhopalospongia obliqua, Hinde
Order HEXACTINELLID &, O. Schmidt.
Suborder Dicrronina, Zitt.
Family ASTYLOSPONGID.
Astylospongia premorsa, Goldf. sp.
Astylospongia stellatim-suleata, Ram. ......
Astylospongia inciso-lobata, Ram. ..........
Astylospongia Roemeri, Hinde..............
Palszsomanon cratera, Ra@m............+..-:
Family EURETIDA, Zitt.
Tremadictyon obliquatum, Quenst. sp
Craticularia clathrata, Goldf. sp
Craticularia parallela, Goldf. sp.............
Craticularia decorata, Miinst. sp.............
Craticularia paradoxa, Miinst. sp.
*Craticularia Fittoni, Mant. sp.
*Craticularia subseriata, Ram. sp.
Sphenaulax costata, Goldf. sp.
we eet tees
eee eee peewee
Sporadopyle texturata, Groldf. sp.
Sporadopyle Santanderi, Hinde
SporadopylesiSp wget armatures eterie rae re
*Strephinia conyoluta, Hinde
*Strephinia reteformis, Hinde ..............
Verrucoceelia verrucosa, Goldf. sp
Ca ee
*Verrucoceelia Vectensis, Hinde
*Stauronema Carteri, Sollas ...............--
*Stauronema planum, Hinde.......
Stauronema compactum, Hinde
Sestrodictyon convolutum, Hinde ..........
Brachiospongia digitata, D. Owen, sp
PALMOZOIO.
TABULAR LIST OF SPECIES.
MESOZOIC.
KAINOZOIC.
Cambrian.
Astylospongia imbricato-articulata, Rem. .... : ;
| Ordovian.
| Silurian.
see ele ee ele see
*Rhopalospongia gregaria, Benett, sp. ........ Reva oman sind scutaliaeon||le.2
*RR KX *
| Devonian.
| Carboniferous.
| Triassic.
Tremadictyon reticulatum, Goldf. sp. ........ Rae ete een isiaed clialataialli sats
Sporadopyle obliqua, Goldf. sp. ............ “canna ea 2 eo ae San
Sporadopyle ramosa, Quenst. sp. .......-..-- ioe eel | Eee ae
Verrucoceelia gregaria, Quenst. sp. .......... solallba calltroas
*Verrucoceelia tubulata, 7. Smith, sp. ........ Bere 825 dees Oe [bagel tapeetenl | emcee le
[kK KKK
Jurassic.
CRETACEOUS.
Neocomian or
Lower Green Sand.
Cenomanian or
Upper Green Sand and
Chloritic Marl.
Grey Chalk, Chalk Marl, |
and Lower Chalk.
Upper Chalk.
Miocene and Pliocene.
Hocene.
TABULAR LIST OF SPECIES. Zien
nnn
PALAOZOIC. MESOZOIO. KAINOZOIC.
CrETACEOUS.
a [a
3 H a és s nd 2
5 E aea\las 8
mn a3 Ss |\3o =
é a Bam loil . 4
5 EE 82 |se| = g
; . re go FEE |e 6! = a
akeehe | sis lela ise B02 |a5| 5 2
er Ee Se Me eee eM at ee et eels Sm renee li ost 8
Bee ree Wee sell Goi eh & I) eee leeeamcetaieames lip tee
Sricm vant ce op || eh) = 6) 2 le |S) 2
Family COSCINOPORID AS, Zittel.
*Leptophragma Murchisoni, Groldf. sp. ........ * x | x
*Leptophragma fragilis, Ram. sp. .......... spine cccilla. 6.016) |aco ella cena onal ail leona lento Ho tcuacc| loeicne *
*Pleurostoma radiatum, Rem. .........-.-.- USE| |qoee|| 4b wel\Zo call opis ol pcre! orem Iamacesl ine.d ol n Dfcecesal lars sto *
Pleurostoma bohemicum, Zitt..........-...- Bese eis cetete | eecaron| ivoreaestl Parental tse evenei|[ecsurerat|lecsrecel| cheer s|leneqereirane yaya isis
*Guettardia stellata, Mich. ...........-.. Se elle tell son allot gral earl Seve ol intern abestal Sea elcsrentc. 3 * | *
Guettardia radians, Hinde ..........--..-- sere SV aaraes | ace shail eicoteeell vee sucllltSye coral feratinsslliscecetsiliateucrallta Inge
«(Cena tunel plbO Is, (Cola > Se oolas collec sellaneo|oooclna bellseceloocolacouleoce|ococn *
= Coscmmoporalg WincoUn Cialis, ME SINE SPs 1) «lle « p-I|-12.21-]\0 ene! jllore. «*illov ep o1=|lf'> «1» elf mer ie|'2 = = =i loreter)io t= *
Family MELLITIONIDA, Zittel.
*Aphrocallistes alveolites, Ram. sp......-...- *
Family VENTRICULITIDA, Zittel.
Pachyteichisma Carteri, Zitt. .............. i *
Pachyteichisma lopas, Quenst. sp. ... ...... 2
IRE Gin ENG Mss [0b Gon Oey Ab pod aoe eee BOI *
Trochobolus crassicostus, Zitt............... *
Trochobolus lucernus, Konig ............-- *
Trochobolus constrictus, Hinde ............ a erelltewsvao si og oll Bkevezel| evoene!ll otercllh cle
Phlyctsenium coniformis, Quenst. sp. ......-. Ceeere || eee | eee *
*Ventriculites tadiatus, Mant. ...d-.... 06. o< La eerernn (ee ee Be aac cont ocaren lle rawat oll (aremaeel tokos aval Pe fate renel|lerenee *
*Ventriculites impressus, 7. Smith .......... Botte || seein | eae HESS al terra | ceeickol cacao! (crete la.oa cll breinanetic! jolt ao *%
*Ventriculites convolutus, Hinde ............ ows ell oes ai| Ree che Fee [Bae clll sree teked che |potouertelllsvicgers lionel stavatleveiens *
Ventriculites poculum, Zitt, .............. eesyena lta sical lteconsses Ba [te aivall lhe esirell eee cell Reheat eoteeeel lithe temo el| evte rele %
*Ventriculites decurrens, 7. Smith .......... EMSs los sPrate| Shssttese ec euskal Se stall ce nets [Pebcehiones faneeensl ca cteslfatieces eaarall se teen *%
*Ventriculites mammillaris, 7. Smith ........ Bese eens (ee eee || ee ree eae | eter || ola saillcrcees 6 llasaee ac llpche eee ree *
“ Vemiimenliities TimiminG hm iiormoey Sh MOM | go|ls000|a45 Saa9 455 0alb0onl|sacellsocalleccnleaollandoedeaos *%
*Ventriculites cribrosus, Phill. sp. .......... Pc A SOE Peale leeeeealee idle e [12 IE, *
*Ventriculites angustatus, Mam. sp...........|.... eee rs care | uavrern ll steal Levawekel nesRor ch ay grees) Mires clits cats ov srelltajs coc a
*Ventriculites aleyonoides, Mant.............|.... Be elec op | coat ell emer ica railewewcwalaeasvelli wl sie |fatava cntee | ree *
PehiwzorbapaUsMipyCuss ithe. 06-2 s- 2.7 cs clee«- BUA ea lyons lIAe ioe | leyercyell teeters ceaacelllcnteze| ve sree alliances %
*Rhizopoterion cervicorne, Goldf. sp. ........|.... Aedes vallo salle c alle .6 laa dle orm deck | aigicicies| [ao ican *
*Sporadoscinia micrommata, Ram. sp......... 66 alla o-o.0|eack asl locho alles c6l|la.0 6 olla Gann] /omonollone aa oem © see
Sporadoscinia Decheni, Groldf. sp. .......... s60||oasollag calla s5llo2 calllse collacweulloespeal acids) Herp cmc wave ae
*Sporadoscinia capax, Hinde................ Bro eve uses | eoctevel llevan ciel Mecttonel ll ersyerelieceetel|l ave. W'lloveetellla ave crete *
*Sestrocladia fureatus, Hinde ..............- eeu Case | Pec tee| tvoiees lteeeote'l|Psbenc|l ooree.cealltas «.2)[tom cone leerare sats *
*Coeloscyphia sulcata, Tate .............--- ES cereal me merall les caea sll eco uel |i evcocarat| et 8 availa id valle tqunnall ener tations terocd haat
Polyblastidium luxurians, Zitf. ............ Eee ec | eeu Metco | Pcnsancc | Moraitemtal Reicovayeil eee chelliav evauai| eretce,eil lotemsterar *
PRolyblastidimm racemosuna, Mee Sita Sen tee |e a -\|'82 eiell'eie as leteie l(c «llr o-|'s 2 rls = alee = elle ee «2 SERS ees
*Polyblastidium tuberosum, 7’. Smith, sp....... Pa Presi carte Pec sta este say sll'. cnavel [toca ie lfxe. cl Matar Morell”
*Cephalites longitudinalis, 7. Smith .......... oc lsc ee | pena ge tae Ai Uren ere Mea AN al ae tg
*Cephalites paradoxus, 7. Smith ............ Rese ese eet alta, 25 Oracles) eae olen cl neta lactows *
*Cephalites Benettie, Mant................. BRE wrslltes nae ll ease Sark Mltevcuctailteaahecclll coevel| Meare lee eects sete *
218 TABULAR LIST OF SPECIES.
PALMOZOIC. MESOZOIC. KAINOZOIC.
CRETACEOUS.
, Chalk Marl,
Chalk
Chloritic Marl.
and Lower
Cambrian.
Carboniferous.
Triassic.
Jurassic
Neocomian or
Lower Green Sand.
Cenomanian or
Upper Green Sand and
Grey Chalk
Miocene and Pliocene.
| Ordovian
| Silurian.
| Devonian
*Cephalites alternans, 7’. Smith
*Cephalites bullatus, 7. Smith
*Cephalites catenifer, 7’. Smith
* * * | Upper Chalk.
Eocene
Family STAURODERMIDA, Zittel.
Cypellia rugosa, Goldf. sp. ................ eh
Cypellia infundibuliformis, Goldf. sp. ........ Egerecd| eve datsl Gncaese terete
Cypellia ceespitosa, Quenst. sp..........+..-- Bey -r
Cypellia libera, Quensé. sp. ................ Ae:
Stauroderma Lochense, Quenst. sp Bis ences (eictora| otra lines:
Stauroderma? cylindratum, Quenst. sp. ...... Rates lacetaltocroltar oo
Purisiphonia Clarkei, Bow. ...............- Santi cove
Porocypellia pyriformis, Goldf. sp. .......... sero wae
Casearia articulata, Goldf. sp. .............. BE slspovlloaodl.ons|loc.csllle oes
Porospongia marginata, Goldf. sp. .......... Ls | atta
Porospongia impressa, Miinst. sp. ........--
*Ophrystoma micrommatum, /twn. sp.
*Ophrystoma ocellatum, Seeley, sp. .. .
*Placotrema cretaceum, Hinde ..............
*Cincliderma quadratum, Hinde
*Kubrochus clausus, Sollas
*Protospongia fenestrata, Salter
Dictyophyton tuberosum, Conrad, sp......... oe
*Dictyophyton Danbyi, J*Coy, sp.
*Plectoderma scitulum, Hinde
Family MEANDROSPONGID_E, Zittel.
*Plocoscyphia fenestrata, 7. Smith, sp.........)....)...
*Plocoscyphia labrosa, 7’. Smith, sp........... Peed mol Roesllbaal lc
*Plocoscyphia pertusa, Gemitz .............. basse Mee
*Plocoscyphia reticulata, Hinde ............ BO segidlls iad (5. clnsllooasl||aerco eae
*Plocoscyphia subruta, Quenst. sp. .......... Bee) Renn Bota bre eel salon allec-cclloaxd|lece vor *
*Plocoscyphia convoluta, 7. Smith, sp
*Plocoscyphia flexuosa, Mant. sp............. ea lees age (erty ane
*Plocoscyphia labyrinthica, Want. sp. ........ Bache ere eee
*Plocoscyphia vagans, Hides. -e 2 nies oie
*Plocoscyphia foliacea, 7’. Smith, sp........... Aaa SoG [8315-0| (DOSE) lc cal|lbaadlloaod|oosollocmalesdosallooac *
*Plocoscyphia elegans, 7’. Smith, sp.........-. Sere (Rarer riots (o-o1 t8] oes | oro ckol fo ot cic. cac04) getenel heise cred toner *%
AME OS eaten SISO Yh Ga so adlsan allo snallooacloosallbacalleooc
*Htheridgia mirabilis, Tate ............-... ore lation e.cioc| lotciec lOnce| hoi: .4| fara: [ao oi| Geeees eter Sioscilloiniors *
*Toulminia obliqua, Hinde ...............- 2a soloewols ofollootdnllbesl||oo salle. c-oa||o bon) |piama||bom olacilsac *
Toulminia jurassica, Hinde Be A |e secre | creo [ccna ellichere el eer
*Camerospongia subrotunda, Mant. sp......... Sifese lhagsas | cistinned|(orencns |iezevs:)|llorete ell mite te)\| okay bl] Sacer eeteg sll eeea *
*Camerospongia capitata, 7. Smith, sp. ...... {os0\soqclonscllooon eyesel| [lene fee
Camerospongia fungiformis, Groldf. sp.
TABULAR LIST OF SPECIES. 219
PALMOZOIC. MESOZOIC. KAINOZOIC.
Creracrous,
Ae
.¥ wea [as g
ces os ee 3
aa saa | 30 =
A a2] |e23|Ss! 2
5 86 gee (+8| 5
; ; ; 5 H S6o|s a ce
Alea lee |S leet. (eesleeonll = (us
Oo e aa} a io} n a Cc + o 29 =| oO r= o
BE -eiep ee ilfeee line, eel eye | a Unease Sales
So ja (eno e | 6 S Sy =
Camerospongia turbinata, Gebel, sp. ........ Eee eel ae | renee ears ave sel aise cell de Deal erate eae full cavres create tan *
*Camerospongia campanulata, 7’. Smith, sp.....|-..-|---.|.---|e-e-feee [less c[eeeefeeee|eceelee ees s|e-.. *
*Camerospongia aperta, Hinde .............. eal Neal epestete | psoas (iereeabe | [Mele | Bpsuara lie suet el ioict ral ieceseverecall oneal *
Cystispongia bursa, Quenst. sp. ............ pepe oeellot on or ealo So] | ete | ete ctl are ancl cael ororeaa | om cig eo
Family CALLODICTYONIDA, Zitztel.
*Callodictyon angustatum, Hinde ............ sccdlleocslloocalace cla acellec.culloouelloo ce ote looters pst lees
*Porochonia simplex, 7. Smith, sp. ........-- Mavaval|lerencsal ll osoterei etl [steers Pcl RecReae lice ster apa) tocanehap sie ecesete *
Becksia Scekelandi, Schlitter .............. Bee | creer | acteeteal | pears | cerepespel [tet cuciall Se chchellfecenretel | Uepeecekel|esvek sy store we *
Diplodictyon heteromorphum, Heuss, sp. ..--|...-[..- [eu efeeeefeo coffee sales eefee cafes ce[ecsses[eoes *
*Diplodictyon Bayfieldi, Hinde .............. vodulleoncliaacollaawellosealllocfclion me locpolebed|mocean Sel) ose
*Sclerokalia Cunningtoni, Hinde ............ ede eratecre |tceseere | tetes ical [a etrov ell ereseotcal[rsretcuctl exstaucll aaa
Family CAXLOPTYCHIDA, Zittel.
*Cceloptychium agaricoides, Goldf. .......... Se allo ectca ate alloc ote Seer eet ero loo ao llow mellbran ac trots eal
*Ceeloptychium deciminum, Rem. .......... dee alla ele veefeeeeleeccteeeeler eens *
*Coloptychium furcatum, Tate.............. 6h. dle oalhecn Rone os-erl soe lobebl to oallocp.c|looncne Berers||
Ceeloptychium Seebachii, Zitt............... seats coclaoecosollee odio cadlbeovllonéellooc ollanmects susnetel| Wak
Ceeloptychium sulciferum, Ram............. kano eorleepelreaslaeedlllecelee allan solomon bare cad coool £3
Ceeloptychium lobatum, Goldf. ............ 6 Sica itredae | Peiec| ea taelleenes eae leranctell eros Saagllooce oe veel
Suborder Lyssaxina, Zittel.
Family MONAKIDA, Marshall.
Astreospongia meniscus, Ram. fe) Ho colloid lcm | es
Astreospongia patina, Ram. ........+..-..Je+-e|eee- *
*Stauractinella cretacea, Hinde.............. soos sa nolsaed|coselleeos arodlécan| sone se oo] perio re
Family POLLAKID, Marshall.
*Hyalostelia Smithii, Young and Young, sp.....|....|.-..[----|---+]
*Hyalostelia parallela, M‘Coy, sp. .......... Sede | eras | epee al| seston *
*Hyalostelia fasciculus, M‘Coy, sp. ..........|+--- *
“Alabyalkosieliia aipcinqgnenibls), JOO oo ee Gombe ao|br ooo oe ollaahe||eaon| leon] oo aerate cos: sitll letareestens *
| Bl brings Conte, (Critter 66 cece anaoooed|lco colo oc dlodce||oe de *
Lll@unsigvuallliey AY@pmynry JS se Conb boas ne ueen||oe poloeooloood|oe ce *
“Aiglerreallles \Wantetaty CORP occas Eh adedl cleo Oobel omme lee ne *
mHolasterellay bennielseELcid camara ecetet se ay-ne gr'sy<||(26 -Fe'llri ea. ll(a1« areiliera = © *
IncERT# SEDIs.
*Amphispongia oblonga, Salt ..............|....|-+-- *
Mortiera vertebralis,, De Koninck: yeisine ooej-cllec are | 0 e- oo|-- > *%
bo
>|
bo
220 TABULAR LIST OF SPECIES.
PALZOZOIC. MESOZOIC. KAINOZOIC.
7 |
CRETACEOUS.
alae é
Bs sed |e g
aie #26 (35 2
3 28 a) aa S Bil oe Ay
5 S d 2 a. WS 2OS)/e&H|] & a
pees | Bi) SES eee eee ene icc | ales 2
ei cse | es] 8.) Paes a cedieenaie ee lle. |) 21 eens
Ele) 2 2 B = 5 Al Seles = 6 3
O/O;/a};/As;o} ays i) o) » 5 a
Division Il. CALCAREOUS SPONGES.
Order CALCISPONGIA, Blainville.
Family PHARETRONES, Zitécl.
Hudea Manon, Miunst. sp. ..............-- ox /cval| a tenes silts topexel| erektal| eed MES
Eudea polymorpha, Alipst. sp. ............ serine Reveille: ailee6.cll|
Hindeakelavabas Zemin 3 ths. clei oecicitsm cnt © stcel(oeadloucalcaccllocxolloo oll es
Hudea perforata, Quenst. sp. .............- apis at all Vetter evens ate ape eregrenel | eaeeedl ese
Eudea globata, Quenst. sp. ......0...e eee rosario evoneills geasnall iecelerel| Comme *
Budeaipisa Qwest. Spe. < stejes wists alo ae BascelllB ue ceiltenehsvl| 2 espe Mee *
Hudea hirsuta, Quenst. sp. -...........0.+6 Real eerste lee aeilgoaalilsa coll’ x
Colospongia dubia, Miinst. sp............... }' ——(Upper) species, Watstioterstresis1 vatale!sieleleseleieies« OY || ——S(Gusitity, JO gnogdoooncooondssooosnone. 67
BIOS PONPIA sD tla neloleliynielleleletaleteleet=lo/ebedstateleleysteye) sie 10 | —— , var. brevicostata, Mich............. 68, 69
Epeudea Manon, Laube ............. 0... ee eens 161 | —— » Var. Cleyata, HUNde . sac cecccer veces 68
Epitheles astroites, Laube ........-+-+eeeeeeeees 180 | —— SUV ATs RESSONIS, VLCCieataleleael wiolersielereshotelereit 68
WT anit WETS oso pooeDonaDDDooeoOrOOUDEE 180 | —— lycoperdioides, Lamx.............eeeee ee eee 180
foraminosa, (Gein. war. viele. 26 as» elele) einai joie ... 181 | Haplistion, Young and Young ...........0.0e0s 207
SPUR COLE, GID 00 ain .0/eie/eleis He ieieloirielere) s+)» elvis ie 170 | —— Armstrongi, Y. and Y..............00.005- 208
Hater og lip leew AVIDO ate parsteretsyeteiefelelsiete lleralershol el 183 PEA CHUN PARRA) oret nore oseselolereiclepctocasielotettataets 207
AAaRn pA EOM anacqnddenvoo rn ooddobddoOC 1S) |), lees “Aine Saanaoocoooboaodnoccs cones 18, 51
WCC Whar ooo SenooasAsanooS dopo aMOGoUCDO 139 ObLIiqua;WBEnets word. orci ~1olisiefotelslalalenel=tetelterar= 53
AMNITADUISs SHALE or. sernla a cpels, Sere te eros 139 | Hexactinellidee, O. Schmidt .......0.00eeeeeee 2, 91
rbrochus: Soll... 2: xis, sisioc015:< sista ee ebro ee IPP GR) |) Lethal, as) Baoanbooonosiossbdodooabs5eess 164
CIA WSUS; ISO coe oeisre's cversis.s Hoste eine asferemie nine 129 milleporata, Msinsts 5 nia 1>!aiaterie elelelelelletersietere 164
ONCE) EATS Sap eS OeDOnOS PDE ne ds SoG ic WS asa |) IebiGh IDET aa sap annnaoasnonoS Seadoo s So cgs 57
GIUMUPBEG PROB a s c/2oierofeta so ester etale Reker AAS) ——— fi prose, Peaeie ya) argesy el vioileieaehated cerehloreeteltaietst: 57
Clawaita ya trage or ats oles o1sts sisrstersie) sts SIeuareeneente 161 SPRETOULGIS, DUNG. areipicin/aideloie(olellsiokel=taloetatelote 57
CHLOE CT OP PMACH a eye talei Vote cenciaxeictele eos stoyaeuareien 161 | Hippalimus flabellatum, D’Orb..... 0... e eee eee 170
—— FL ODA QUENE. e (i) JSTOy BOSCO POCOUDD OUI OODOIEOUTOE 55 MACTOPONG, SRALDE.ssismici<\e ices cetera 199
—— marginatum, Goldf. ......... eee cece eeees 125
SFSIGH PLAGE: 5 ol cisiels o/a/els\0 wes ie 1s) elelalefolr sine) efeis 12, 63, 70 TCG ASCOMM, ER COIN» vvezeneieisie clei eRe 138
Gorter Move Anta) hors \a1e) leleieisisie!=I=rehats!alsts)2¥=/~/e 49 MURAI E; TROUSB 5, «,«sosevayors/av2s «1 re) Seo ace RVR NS 39
== Giniinigniit, WOES soonee coaqoaccdeduOCUOF 71 MONGSLOMG, HOBIE ojerereroelaeraeie eee 141
Selene CLG MICH) 101) s[aleue)eiciesaiore ofeleieiasialeivieisre ele 49 PEF USUME ND Ste ere s)-/etel eee ie ee eee 162
—— EXCavata, MUCK. ec ve vicicinics shee eesccewesies 72 | —— peziza, Goldf............0...- 162, 192, 194, 196
—— gregaria, Mich. .......+--seeerereeeeceeee 90 | —— macropora, Quenst. .......0.++.ss0000 194
MAI SLO EUOPTI olefalel\ateis}ntele\slalo)ul=/e/ejelettsleia)a 42 | — stellatum, Quenst. ...............-.--- 200
—— pyriformis, Lama... 1... ee ee cece cence 50, 70 OSEU OL e MUNA wrap hreyvor-Tae ares oe ree 163
—— Quenstedti, Ztt.......ccccccscveccserecees OD) == por cattint, SHATPG, «76% -c:-rols ale eiveveisis ose 6 see 198
ITE HN CHLA GA PETUTLE Wl cieicyo.cio,=| afelefursiaia\clafeis ol eel 70 | —— pulvinarium, Goldf. ..............000- 190, 191
——= Panis Veeditke eo ge deoG d000nUODOOOUOOUHOnE AGS | LEUSSIL IME CON: waaiered-psrc¥ssononeteretaisis cl elevere ae eave 40
SINCUAGETO, LORIN. 6.06 eciecicnsceecesersacss 55 SCCM GLUT COLI tase etoxareratatoielereyajeiei-fe isles nether ee 200
Went SIS Segenocoppecador OpocGopHG. 70 | —— submarginatum, Miinst..................... 163
TERE, Wis an dasaponooeapoon cpu oedoodd 12, 13, 42 | —— tubuliferum, Goldf...............000000000e 192
=== hye JED GeotoosucvanponsO mcd dpouscole 42 TAU PE TiDp IOS Seba6 podo og onOoIGOnos OOF 41
—— cylindrica, Hinde ..........ceseeceveveees Bor (Mastosla, Litt icec oascccades tacts se isieaciens 11, 57
—— polystoma, R@mM. .......0ccsersecereceses 42 MEO COMMENSIS, AENAL mise tere racine ceisie sities ee 57
—— punctata, Miinst.... 0.00 ccceescccceeereeas 42 | Mastospongia coniformis, Quenst................. 108
Jurassic species, Distiof . 0.2.0. cscs cece e ewe ees 224 GRegaria, QUCNst. wa sinish,6 8s Pecan eee eels 97
Hip CNGUNES da Sagens ens senoosanoosobosdaF 107
TiGiynGih, J7072 Shans sone onecon OOdOGUBOOUL L276: === wernucora, QuenBts 4) .i.:c:a}=¥elsrers Sm Me ralithistase72bLs. cm ativiece/eietels fers eta ceyere sts 11, 47
Lugs Ain & 5 aoa gouao onnvaGooRoDEsobad 47
Lancispongia lamellosa tumulosa, Quenst..........+ IG || MENGE Ais Gicaucnosdgecdnon bponocoodsod 11, 56
biarG, QWs onaa90n5 so sao gcoge dso DoNseT 107 TAGIA TAS LOQUEIST in areter cess nee 200
tubulifera, Goldf. +. cause swee nner eee 192 LOANS ES JANI Oiscan Ga one odyao vO O025 00. 193
(Ohanalliy i Aaeadeness soo s50 35 b0GR000005C 1S) |) —=—Strahant, Soll ry yeveretetonyoy-s yous te e's leis’ el 9, 201
QOphiraphidites, Cart. 2.5... s.000c0+-00 +00 => 13, 25 SUD DERICAD VAM UUs tee te rele tee leetanete r= lol ote rte ols) ave telotel= 196
AMASCOMBNG -EUNAe) | sa wale ee ice rieresteretcnernne 230) Pblycteontums Z7zts war tstieieisteisis wisielenst ose -7s1 11, 108
cretaceus, tits serree tic eee ehecer reer ere 23 (amimete, OE 35555 G00nsd0500" 00000 a 108
Ophirystomay Zita siete cer ere ttre 1251253)|| Bholidocladia whindem enn ml mtej-rrr ieee lee 13, 80
MIUCTOMMATUM, POM Aes eee ee eee 125 dichotomus eiinleeerrmenins cementite 81
ocellatums Seeley sector eeeie ieee 126 | ——ramosus, Hinde 20. ii. .cve ee cceessseeres 81
Ordovian species, List of.........0...--c+-.0-0+s- 993) PhymaplectiaHandenmemes annie) tetelteitaleeiele 13, 87
Orispongia globata, Quenst. ......... 0. ee evens 162 Oribrata, Heder enter telertatore/-< Claris oe 88
faa fart ONES Genaonononcca9ondodes ben 162 | —— irregularis, Hinde ..............00.-00000- 7
BCIbUIB CLEA meee lee ee cic kesh ofaral as. cole etciseie ieers 13, 26 | ——spinosa, Hinde ..........eececeee cece eee 8s
abyss On Schitan meter ei ser ie ieee 96) Bhiymatell ase Zezdapertatststetccoicteloietelelate eietalerevate 12, 15, 58
=== COnVOlUbarphLInde g.. wish oe SKEET Ee eee 96 | —— heteromorpha, Ram.,..............0000000- 60
=——AMLETLEK ban CO Otaesavelelsrelataiel slated toieiclelatei-falelal eles 96 | —— heteropora, Ram. ............ee eee cee eee 59
Ey JE Gegononesdo4a64oD0so0c07005" 27 | —— intumescens, Rem, ................+.---- 58
==" PTIMPOVAy ZU, one oy ojo ele eins olele wiieie winnie 96..\) == "nod ona, Wiendee, 6 er. ns wine 0 Rie see ee 60
WENO ITU) Aah amon cose toodeoosansouGs 209 TPO LACH EAE LTEAC. folk ere infeieheyefaresstniers) sions ee 59
MiNi Ab" sodas seossuobMaD0UCerex 13, 46, 56 Ah osu agen donobo cApouEaGWnoO oo DOR GOD 666 60
Seripbum Ren) sci) ce ate ieee eee eee AG))| Placochizenia; Pomel 2 yr). <1 icicie = ni yefoe sete ee 127
Rachy poten Onyel eiae we ve elses sieltesaisie terre 19,51 | Placonella, Honde ............000.eseeseenee 11, 47
compactum, Hinde. 2. 20, orci os slon snl) =e 52 POLLOLA LAs LUCE voqereyesoue ionofoteishaie lesen teyaae eee 47
robustum; conde 3...< ec vsisis Me wloareinies, te a PRLS: 52) | Placoirema, el inde cc icenrctrrs etic a retin 127
Rachyteichisma,t vite erryaniiee serine ss a 11, 107 CEbACE UMN, e Alte) rer erie\eleterernte le elects sa atrelt 127
@arteri-eZ7by pocicveys terete ao eieieietoie aint ee etetors ler Kye || boca, WG Snoomssodesosadoacods0cn0 11, 30
Topas, |Quensten 5 sors: seirtadeiaeen eee 107 | —— auriformis, Quenst..................2.00005 30
B\thi moocoue duadon soatomdaco OonsoE yen Onc 107 Ele baad coos Woiois orators Wanna Sem Renee 31
IBA Aim oagpasachudsnoocopoededeote% 203 VACATE, MOMENT ce ym ores ete = onal 30
—— infundibuliformis, Goldf. .................. 203 | Plectoderma, Hinde ......................44.. 132
PalesOM ANON LCC, cnsyers oke os, cis sue loses ioleheiee reas 10, 93 | Seitulums 2 rde 9.6 oases eons ee eee 132
(GRID JIS 55 aaaooaoddaqosse ood sanuoaesC 93m) eelenrostomas Hat... c..\ie seminar ee 13, 103
parengeavoraciisne ht. 12. ee mcl oleae tie 178 | bohemicnm) “uN pac.. ae eee eee 104
BeronellayeZattbarscrerce jo) a1s 2.1, te [peter ero 10, 11, 12, 164 TAGLALUIN ACG) pyaunisteloieiten et eee ie 103
ClavATIOIGES PMCID... oe eine ee wees 1GGh|) dalinthosellayeZ7eeeiy ons oleae leit terete ye eee 13, 81, 84
= Ih Jia ha go onde aeseesnAoddasoaoaG 34 168 COmpacta,wende ec. so emer eR cee seer 85
cylindrica (God eee ct ere ueer mies eee 167 | —convoluta, Hinde ,.................2.00-: 86
CYMOSA UAT ec eeie cis sme pote chaceulne 165 | NOdoOsa; Inde. sonektewe eGR EIEm teen 86
flabellata. 8/1 O7 Domenie taser creates sai 170 BOUAMOSA, ALC.) cine Bete toe etat aerate 85
Snide GULIE) soponce nos ges 9008S erat 170))|) Lliocene species, pLastiolnenyerieriteel-e ee ee eo
Gallieroni; e2orn ans eel Beek eee menen Gunite 169) || Blocoscyphia;-Aewse. lec eis leks le ons 12, 13, 188
inflata, sHende’,:. bare come eee oe owin s ovteics 167 Convoluta; els Smedaweer cries cn Seek eae 136
=> mamillitera Lame eee eee eye 166 ilies Ji SEIS" 5S oce couseeaOon coins asec 188
= Michelini Bets. Jas onion eer eee ieee 168 ‘uGenen, MISO saboooasedaossnososonc 133
Be rb aigitata, Litt. tree Cre C eer nec: 183 flexuosa, Mant. .............00..0.e see ees 136
=== nodulosa, (Quenst sa: Sten eee ee eee 168 —— foliacea, EN SINUM a ec ee 137
ocellnta, Hinde w.,...)-- | sree ee eet 171 JabrOsa, cli Ot/e fole)!sieleteletels 45
subcespitosa, Miimst. ....... 0.0. 222s seen ees 165 Sulcafer dy ROBIN ecw e+ -ie = m= +1e\sie\s(°le]8ielsiel=)= 60
—— subseriata; Roem. ........00eerecreeesecee 95 URED INS SOK 59 AU CHO NBO ODOODGOOOUUOOOr 55
—— tenuis, Quenst. ....0+5.sesrereceeereecces 101 UUMEGLA ECOG erayetaleicle/o 1s i= \-=\-vele /elelalelolalelolelale 168
testa-florum, Quenst. .....+. esses erence eee 33) j]) Sollastn, Stetanspeyerlotereret-|«1o\o1e1-el-.-\-\)o[ele -lnleieisieiay= 10
—— tetragona, Goldf. .......-- ees eee eee eeenes 182 | Spherocerlia, Steinm. ............-. see reece ees 171
texturata, Goldf........+.seeerevecccsseces 96 Michelimi, Steinm..........+..2eeeeees 172, 178
verrucosa, GOldf. ... 06... seeerereeeees 97,108 | Spherolites Nicholsoni, Hinde .....-.-+++++++++: 57
Scytulia, Zitt. 0.0... sce e cece eee eet e ee eeee 18, 44 | Sphenaulax, Ztt...........0.. sees eeee Gaone LUE,
fastigiata, Lee... sss ee see e eset tence eens 44 COStata, Gold fe o.oo. ewes snes ...- 95, 108
radiciformis, PA2ll.......-.0cccessesceceee 44 | Spongia capitata, Phill. ..........2e eee eee e ees 31
terebrata, Phill. ........0ecceee eee eee eens 45 Clavarioides, WAM... 0.0 2 seve vie « cinisinie 166
Sebargasia, Steimm......- +++ ees ee cere reece eens 10 convoluta, Quenst. 52.20. ecc see + enone 38
Seliscothon, Zitt........:e eee eect eee renee eens 18, 31 CT2OT OB MENA areyayers ole eleveiorereoeienesereker= lair -ieieks 118
explanatus, Ram. 00.1.0 6. cece reece eens 82 | —— cymosa, Lamx.........0-+.ct eee ress eee 165
—— giganteus, R@M.....-.+- esse eee nese eens 33 fastigiata, Lee «1.1... seen cence eee nee Ad
—— Mantelli, Goldf. ..........sseeecece secre 33 | —— helvelloides, Lamx......... Ad obdocoavmAS oud 198
—— planus, Phill. ... 0... ceeere cece renee ee ees 33 inciso-lobata, Roem........++e.eesererecees 92
BP ieee ees oe le tevaie oiee ise siclaisindjoleroreistcieteials 88 | —— lagenaria, Mich..........--+0++2+++eeeee 168
testa-florum, Quenst..........00eeee eee eeee 3 | —— mamillifera, Lamx. ............-.eeee0ee 166
Serpula parallela, M‘Coy ... 1. eevee ee eee ences 151 | —— marginata, Phill. ...........-+.2se0- eee 36
Sestrocladia, Hinde .........0+esee sees eeees 12,117 | —— mudtidigitata, Mich. ...........-s+ess.00-- 183
furcatus, Hinde ..........+-eeeeccrereenes N72 weeny NL Chir ejeyeatols\aletelotniel=tslsti=\alol oletelateteletetate 194
Sestrodictyon, Hinde ..........-+0sees see eeeees 101 pistilliformis, Lamx. ......+++++++++-+ee+>s 165
Conyolutims Eende terse orks elowistois sel bisisisiel= IO}, |) ees WM Shea anos ooo vod ede se onDS oon 31
Sestrostomella, Zitt.........2eeeece eer eeeees 11,187 | ——radiciformis, Phill. ......-+-+-++eeeeeeeeee 44
Gry auth Jl Bl Seaononaden opcodes us ouGKc 188 | ——‘ramosa, Mant... 20.05 ee eee ees ie 48
= Cribratay Quenst..... ojo. ascetics «oles viene eye || Ssaaens Wai bapa donocopoDodsOn oon cKo GoD 186
rugosa, Hinde. ......+..... see e eee e ee eees 188 | —— stellatim-sulcata, Roem. ........-+-+.-seeeee- 92
Siliceous spicular structure dissolved ............ 8 terebrata, Phill... 05.60 + ence cee sees 45
Siliceous spicular structure replaced by calcite .... 7 Townsendi, Morr. ......+0cece cesses eeees 144
Siliceous spicules replaced by peroxide ofiron .... 6 umbellata, Mich........+cecceeeeeeeeeveees 186
Siliceous sponges enclosed in flint................ 5 | Spongilla, Lam......-... see ee eee cece eee 3, 11
Silurian species, List of .......-.-.-.-0+++e sees 223 Purbeckensis, Young ......++.seeeeeeeeee 21
Siphonia, Goldf. .....-..0-0-0 esse ee este tees 12,63 | Spongites alatus, Quenst....++-0++eee ee ee renee 190
a ACHUUS; NICH G5. o)-\ ahas eteniehetaterere seedy ieyeleiale 0° 69 articulatus, Quenst. ........0+.se sees eeees 125
cervicormis, Goldf. .......-.0cee reer cece nes 116 | —— astrophorus alatus, Quenst. .......+++e+e00s 180
AMIE Sag Ganendo0 00 od 5016000700 OOMUE 42 | —— caloporus, Quenst........-eceseesenoee 180
CHGLCT.G-JEUCEING 05-2 os etal ayoe se eleteernrs aie sexsi oy} || ——— cornucopi@, Quenst. .....e++eeseeeeeeee 180
——— OLCAVULA, CROLALS «a5, oicis e's wiele el-eielsielafe)=iseverni 92 auriformis, Quenst. .....+-+.sseeeee sees 30
ma AICUSS) COLE fel olele el elera stele) iet =) teeter aelel= 65, 71 binoculatus, Quenst. .........eeceeeeeceens 191
imbricato-articulata, Roem. ........++++++++ 92 clathrata, Quenst. ©... +. 60s seers cere eee eee 94
incrassata, | GOld fan en ee seers 65 clavellatus, Mant. ......0++-+-+eeeseeeeess 7
KiGmigd, Wane. yee cele = « arereleteleeitnreserereel tole 65 CORLGHUS AG) MEUSten | stalol«)sininelreteieteh toll tacts 190
Lobabas Mant. ie «feiss late eitaleete -tateloreeteaetersielerel= 67 | —— cribratus, Quenst. ......6+seseseeeeeee soo HEY
INDEX. IAT
Page
Spongites cylindratus, Quenst. ..... 0+. ee ee ee eee 124 Stellispongia stellata, Lama@..........s0ceeeeeeee Page
cylindricus, Quenst. ......-..seee eee ee eee 168 Variabilia: Munsis sashihis seeds. duewens en 185
—— cylindritextus, Quenst. ........-..e.eee eee 94 | Stichophyma, Pomel ......00.cccccecceeeeees 13, 41
—— glomeratus, Quenst. ......... cece ee eee eee 187 turnida, » Bande - ce on nme hs eon A 41
ICCA AC) CUStaeteltrs tie wees sod ease aes 184 tunbineta; Roms cc. ss eee. vs Re 41
—— Fochensis, Quenst.. 0.00.6. cee cece eee eeee 124 | Strephinia, Hinde: x. 152 uose Wane e ntact 12, 96
= Las QUEMIS Ena wie) clei les ios ele sle'visi# os cis en oe 107 convolutesr inde... tas WME ae oe 96
mamillatus, Quenst. ....... 6660s. eee eens 166 reteformis; Fltyedel t,t... treat a temate reste sie 97
—— nodulosus, Quenst. .........-- see eee eee eee 168 | Sulcispongia colliciaris, Quenst.........++.00+e00: 95
obliquatus, Quenst..... 2.0... 60s sere seer eee 93 ¢Metaa, Quensts. 5 /2,2/)- 5 ee sachs orerereetenel rere 95
—— perforatus, Quenst............6+..ee seen ees 162 rimosa, Quenst. ........... fob ccono a0 cIodon0 95
—— ramosus, Quenst.. 6.2.0.0... cece eee eee ees 96 semiclathrata, Quenst. ........0ssseeeeeees 95
—— reticulatus, Quen8t....... 6.0. ce eee eee eee 93 Sarid, Quenst. \.268 sc lec ee 95
OLR UCUS Us atst teicictacisietste irae etevokeicisie eres USA Supplement syncs seis carw reese reset eee 207
semicinctus, Quenst. ........++++eee esse eee 18% || Sycones,«Hacckacetr. nakrnkck see eee ace eee 204
DAG ANS UCUSteye aiapelstcra cece s.<'eieecieratcrela ses ole Gy Sipe isc omeneorcucocebouvanseoc 12, 13, 190
SPONPOCTSCUSS Atte eyeierelefalsiele sie eislvlete s/e/elelele/s s/eln/le 87 (Collab, Je GooueoonoooeAb ooo onco one 191
Spongus botryoides, Konig... 2.2.06... eee Cy |) joni (AMlEE sooconsononceoneonodnsae 190
GAT ETT ey ONES Sogo Kn ObUeUOCCOROOOME 107 Spheaticas pill acini tomitscteierteitelsy )> 58
Sporsd dscininy omelet eee eee 18, 116 Tetractinellides, Marsh. ........0+5.s000++00 ss 2, 23
= Gilihy, Je C EEO San Gee HOpE Coos GaSoRGe 116 | Tetragonis Danbyt, M‘Coy..........s seers eee ees 131
=== IDuGall,. CHER, ondonononadoudumadocapaper 116 | Tetragonis Hifeliensis, Ram............++.++++> 131
micrommata, R@m. ......csceeecscceueees 116 Murchisoni, Hichw..... Sea loutsdeekve Boies aeeleuatole 181
Spumispongia punctata, Quenst. ............065- 42 | Thalamopora, Roem, .....-... + eee eee renee eee non
Stach yapOonpiay Attic. sergsonascacerddees 12, 13, 45 eribrosd, Roem. 1.1... 4.222 eee eee ee eees 174
== fla, 17 Pe apo enbeaoeoceouboonoonooooud 45 | —— Michelinwi, Simonow. .........-++eeeeeeee 173
Slotntint ICY 77214 a poe aoe aoe aE EEC goog 13, 149 Siphoniotdes, Mich. .... +... .+++00 0s se enees 178
CLOtACOaay ide lente fsofsruieraie ev ele oveeteselals Matas 149 pesiculosd, Mich... 6.60... e sence nee ecenanne 174
Staurod ermayt Zito). oa cto ctacs see ee detest as 11, 124 | Thamnospongia, Hinde ...........++..++- 12, 18, 78
—— (ginal, C775 no.0000 copo0D0CdUCaeS 194 | ——clavellata, Ben. ......... 66 ce seen e neces 79
mee ochenise: | Quenst.2). «ixesayeioe es Neetele chek 194 | —— glabra, Hinde.........cccee cece seen eens 79
Stanxodermidias Zttta..)ots% 2 celiststeicei tert sisteteta "2 123 P reticulata, Hinde .....2.+..0.ecerereeee 80
Shemmneney (Sa)! Sen paarae sepeaGie soe mopehese 12,99 | Thecosiphonia, Zitt, .........6+-ee seer eee 18, 75
—— (Chitin, SWS Goce boneraonboooucochnonte 99 | —— grandis, Ram. .........cececsece ee cnneees ie
—— compactum, Hinde. ............scceeeeeees 101 | —— nobilis, Ram........ 6. cece eee eee eens t 5
ee resinen iT IL EIEY ta ck) *..21sars Wena cle el 100 | —— turbinata, Hinde............eeeseee re neees 75
SbaMlettas Os tScherits craletavefs wie s/s vin ateie/deltelowta vib. 13, 24 | Thenea, Gray .......-..+++ee seen reer seen ees 18, 25
OWT, JHE oavcoopensnccnooes poomenae DA | —SapISPe open we ie we eee clei eicininin sr eswsine sass 25
Stellispongia, D’Orbig........s0..sseceeeaes 11, 185 | Toulminia, Zt. ........... see esse eee es 11, 18, 159
PUA, IAS ovancosuodousedapaddiecboooOUeE 187 | —— catenifer, Zitt........ 6. ccc e cece eee eee eees 122
Es corallinaeee nonin eteereyfamelsrtisatersteiate cteiele 186 | —— compressus, Zitt......ceecre eee eee eeeeeees 122
= rilmiernc, Aan - pooogacocugoDUOpouGoEC 187 | ——jurassica, Hinde. ..... 0.0 eee cece cence cues 139
= Gaiiieniim, CBid, orosoooonooncpogueener 200 obliqua, Hinde 1... .ssseee eee e cece eens 159
—— Ty, Wen coon tcnnor oonoeonoGOr 186, 187 | Trachysinia, Hinde.........6...eeee sees ee eeens 189
——=— Manon, Waube: ocicccccoceccer veces snare 185 aspera, Hinde.......++.0---ssetee ree neees 189
—— pertusa, Bt... ccc recs ecce sec e ee ee ee ereeens 187 | ——minor, Hinde 00.5.0 c0 ence cc eee ce see vanees 190
TO MLL ATIS VICIRE slajele «iclaleie s/s vicleleiclsine «eve sisi 185 Solitaria, Hinde ..ccce cess seer secescceees 189
—— semicincta, Quenst......ccseccssevcverrees 187 | Trachysycon, Zitt. ......+. iintege’sistameualalelsveraievese 12, 62
248 INDEX.
Page Page
Trachysycon nodosum, Hinde ....4....e++seseees 62 | Ventriculites Benettia, Mant...........++.sseeeee 121
Sulcatum, HInde . 2.00. cnecaw ven os esis 63 bicomplicatus, T. Smith .........+...-.-05- 112
Machi Um oB twats else = «+ ote evo elelareaeliel> isininialainy 10 cavatus, T. Smith .........-.--+s+.+0 111, 126
Tragos acetabulum, Goldf. .. 0.20.02 ceee cere cence 80 | —— convolutus, Hinde .........2.-+-+eseeeere 110
acutimargo, Roem.....-...+.2-5++++sersse 194 | ——cribrosus, Phil. 2... 00sec nsec nesses 113
Chunar, Anubis 3 eoqcndg0nnocorcdon9an0I0 180 | —— decurrens, 7. Smith .............-..22+-5- 111
Farringdonensis, Mant... . 0... 0000s eee eens 177 decurrens, var. tenuiplicatus, T. Smith ...... 111
—— fistulosum, Quenst...... +0000 ee cece eeeeees BY) | = ciintis, Qin oocceopnossboonebeaaacs¢ 110
granulosum, Quenst, ......2...0ce ec ceeeees 29 | ——impressus, 7. Smith............+++0055 109, 115
—— hybridum, Must. i....-/. <2 -- cielo vieisie eine 186 infundibuliformis, 8S. Woodw. ..........++-. 112
infranudatum, Quenst. .........0+.eee scene 30 | —— latiplicatus, T. Smith...................55 112
milleporatum, Miinst........+..--eseseerees 164 | —— mammillaris, 7. Smith ......--....++++..-- lll
nL EMU tsogoscodnduocdasesa050900a0€ 99 | —— multicostatus, Rem. ...............--200 118
MPROTSHTD CEG Ropnd BA SAAODS0.0500800 00000 134 | —— muricatus, T. Smith ........2-2+2 esse eee 110
REO CMG, .oapongocdn660G500000003 30 | —— poculum, Zitt.......... ccc eee ee eee eee eee iit
momo Clibin sanaanonooogsoqpa00000000 29 | —— quadrangularis, Mant........-0+++eee eee eee 108
—— reticulatum, Miinst.........00 ec ee eee eens 29 quincuncialis, T, Smith .......-+-..++-- 106, 115
PU OSMITAs INLATIS fale le afore lala) olafelelare lolelalerellnyelo\> 30 Tada tuss Cpt. apectotoiele/oln)e\«°)2es+e == 108, 111, 112
verrucosum, Miinst..........--.ssceceserss 30 radiatus discus, Quemst..........00+2+0ee0e 108
Prema holites.eZ1eee rites eleisls:siveteisievsiieierevere'eser> 18, 188 simplex, T. Smith .........+2+eee- eee eens 148
mepastoma, H@M.........0c.seseesceerece USS) || ———— strats eS /e, Petete eve 2 atnlotn tele tnlelelefolsVejnr=i-cae 112
—— perforatus, T. Smith .......0..ccceeeeeenee 138 tessellatus, T. Smith ...........21..++--05 118
Tremacystia, Hinde .......+.seceseseceerreees 171 | Verrucoccelia, Be. ........--. eee eens 11, 12, 13, 97
—— anastomans, Mant...........-+ece cece eens 175 gregaria, Quenst. .......-. sees eee ee eeee 7
=i CIAVALA REED: cn 1h tfs)aloicie a\aieie sleleieiel= «\isieie (sel 176 fubulata, 2 Smith... 20.21. .05seecee eee sne 7
—— cribrosa, Goldf. ........cceeeeeee eee eeees W7A. || ———=sVeectensis,. fide) e.e.0i oie 0 e1e)+1=1=)-10iclo le sri) @)= 98
—— D’Orbignyi, Hinde.............seeeee eee 172 verrucosa, Goldf. ........+2+-++-++eeeeee 97
drrepularis, Hinde... 5.000. 20cseererec ree 175 | Verrucospongia armata, Laube .........-.-.0e05- 168
—— Michelinii, Simonow. ...5... ese c cece nese 173 | —— macrommata, Roem. ...........+..-----++. 40
siphonioides, Mich. .........-++++-+e200e- 173 | —— polymorpha, Laube ......-.00-.+sseeeeees 161
Tremadictyon, Zit. 21... cceeee eee e erences 11, 93 submarginata, Laube.......++ +e ee ee evens 164
obliquatum, Quenst. .......+see+-eeeeeees 983 | Verruculina, Ziti. ......---d0.e ees e ee eeceees 13, 35
MO GLC UL cere) GOLA fai tai tayete leis ole! talololoseleitersieves 93 astreea, Hinde... ....sccc cee ces earn nec ren 37
Tremospongia dilatata, Roem. ......--eee eres ee 192 aurita, ROM. 0... cece cece te eee eens 37, 40
Ui, RAO Sesdqmocdodncsdrdob cadobus 75 convoluta, Quenst. 6.1... e cere eee eee ees 38
pulmmaria, Roem. 0.2... -enes2 ss ee eee 190 | —— macrommata, Ram. ......-.eseeee eee eeees 40
Triassic species, List of ......:22...+.-2+-2-+--5-- 994 | —— miliaris, Reuss ......-+.-e sees eeeeeeees 39
Mrochopoluss:Z221.)2/s,0 csine seis s sie = wietetiaielete ess 11, 107 papillata, Hinde. .....-.++ esses sense eens 41
GCOmimonihy, Je eoonosanagooogdaaangc0s 108 | —— Phillipsii, Rewss..........++ eee ee eee eee ees 37
I ETASSICOSINIS P00: Maiveiaterreniisic sie einetretsice 107 | —— plicata, Hinde..........sseeeeeee tere ences 37
= Ie awis, MONET Wadomocsuoaeamenoco0g 00000 107 pustulosa, Hinde ........eee sere sence ees 39
Tubispongia hirsuta, Quenst..........0ece ee ee eee 162 | —— Reussii, M‘Coy ......0..s2eevereeeseerees 40
Tulip Alcyonium, Webst. .......-.-++eceeeeeees 70 seriatopora, R@M. 1.1... eec eee c erence eee 36
Muroninsy telat tein sleletare eisletsietle elector 13,76 | Verticellites clavatus, Keep. ...-.+-++-++0seee ees 176
“leith (OOo an osvoddaseeda don souRuoonee 76 | Verticillites, Def......--+-+-eeeeee reer ees 163, 171
——— VATIADIUS) IMGCH os alc lleiseie swells ler v= «en 76 anastomans, Zitt. ......+-+seee ee ee seen eee 175
—— armata, Zitt. .....ceee reece eee cee eee eee 163
Upper Chalk species, List of...............+.... 997 | ——cretaceus, Def. ......2+.+ee-sredeseeeeees 163
D’Orbignyi, Hinde ....... +s sects eee ees 172
Ventriculites,eMantin csc eee aie cir 13,108 | Verticillocelia, From.......--++ sees rere eres 171
alcyonoides, Mant..........-...se+esneeees 114 | Verticillopora, Blainv.......-..++++.ss+e0s 163, 171
—— angustatus, Rem. ..... Siac ath eee raat 114 anastomans, Mant.......-.06++ sess eeeeees 175
PRINTED BY TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET.
, eho
whe 3
oii) > hii
Fig. 1
Fig. la
Fig. 2
Fig. 3
Fig. 3a.
Fig. 4,
Fig. 4a.
Hig. 5.
Fig. 5a.
Fig. 6.
Fig. 6a
Fig. 7.
Fig. 7 a.
Fig. 8.
Fig. 9.
Fig. 10.
Fig. 11.
Fig. 12. I
PLATE I.
. Climacospongia radiata, Hinde. Showing the interior of the sponge. Natural
size. Drawn from a fractured surface. From Silurian strata: Tennessee
. The same. Portion of the fractured surface of the same specimen, showing the
radiating and transverse spicules. Enlarged twelve times.
. Lasiocladia compressa, Hinde. Showing the spicules on the surface of a fragment
of shale. Natural size. From the Lower Devonian at Jemelle, Belgium
. Acanthoraphis intertextus, Hinde. Showing the outer surface of the sponge.
Natural size. From the Upper Chalk at Shortlands, Kent oe
The same. Portion of the outer surface of the same specimen, showing the dispo-
sition of the spinous spicules. Enlarged ten times.
Ophiraphidites anastomans, Hinde. Section of a specimen, imbedded in crate
showing the disposition of the skeletal tissues. The spicules have been replaced
by iron peroxide. Natural size. From the Upper Chalk: South of England .
The same. Portions of straight and curved acerate spicules, from a specimen
inclosed in flint, from the Upper Chalk. Enlarged thirty times.
Tethyopsis cretaceus, Hinde. A fragmentary specimen imbedded in flint. Natural
size. From the Upper Chalk: South of England . . ee
The same. Portion of the same specimen, showing the disposition of the seglen
Enlarged eight times.
Stelletta inclusa, Hinde. Filling a cavity in the interior of a flint from the Upper
Chalk. England. Natural size .
. The same. Acerate and trifid spicules from ie same specimen. Enlarged eighteen
times.
Pachastrella plana, Hinde. A fragmentary sponge inclosed in a hollow flint.
Natural size. From the Upper Chalk at Upware . . . .....- -
The same. Portion of the surface of the same specimen, showing the spicules.
Enlarged twelve times.
Reniera? Carteri, Hinde. Detached spicules from decayed chert of Lower Carbo-
niferous age. Dalry, Ayrshire. Enlarged twenty-seven times at tac
Spongilla purbeckensis, Young. Microscopic section of chert filled with the spicules
of this species. From Purbeck strata at Stare Cove, Dorset. Enlarged fifteen
WHEN 4 5 ss ee Ge ech rad a ee yraie “o a Ae Lys tee
Microscopie section of Ga from the Portland Oolite, at pears: near Weymontln
showing spicules of Tetractinellid sponges. Enlarged fifteen times
Portion of the weathered surface of a sponge-bed, showing detached spicules of
Tetractinellid and Lithistid sponges. Enlarged thirty times. From the Lower
Green Sand at Haslemere, Surrey .. . . . :
Microscopic section of chert, filled with spicules of Teteartineliid sponges. ‘Bale
fifteen times. From the Upper Green Sand, near Ventnor, Isle of Wight.
Page
18
19
20
23
24
24
27
19
21
27
27
Plate I.
FOSSIL SPONGES.
West, Newman & C? imp.
A Gawan del, et hth.
Fig
Fig. la.
Fig. 2.
Fig. 3
Fig. 4.
Mien co.
PLATE IL.
Page
Pachastrella convoluta, Hinde. A large ear-shaped example, showing the upper
surface. Natural size. From the Upper Chalk at Flamborough, Yorkshire . 26
The same. Portion of the outer surface of another specimen from the same
locality, showing traces of the spicules. Enlarged thirty times.
Seliscothon planus, Phill. sp. Showing the upper or interior surface of a small
specimen from the Upper Chalk at Flamborough. Naturalsize . . . . . 31
. The same. Showing the under surface and the stem of the same specimen.
The same. A specimen with a nearly plane upper surface. From the Upper Chalk
at Flamborough.
Seliscothon explanatus, Roemer, sp. Spicules of the interior skeletal fibres.
Enlarged sixty-four times. From the Upper Chalk at Ahlten, Hanover. . . 32
BOs tin ol ONG E'S). Plate I
A. Gawan del et hth West Newman & Ce
Fig. 1
Figs. 1 a, b.
Fig. 2.
Fig. 2a
Fig 3.
Fig. 32a
Fig. 4.
Fig 5.
Fig. 5a
PLATE Tit.
. Chenendopora Michelinii, Hinde. Natural size. From the Upper Green Sand
ataWarminster; Wiltshire” => —) 2) oe) eeeeneene ens oe aetna
The same. Spicules from the interior of the stem of another specimen from
the same locality. Enlarged one hundred times.
Verruculina pustulosa, Hinde. Showing the inner or upper surface. Natural
size. From the Upper Chalk at Flamborough, Yorkshire .
. The same. Portion of the outer or under surface of the same specimen.
Natural size.
Verruculina miliaris, Reuss, sp. A small fan-shaped specimen, showing the
inner or upper surface. Natural size. From the Upper Chalk at Flam-
borough .
. The same. Portion of the under surface of the same specimen. Natural size.
Verruculina seriatopora, Reemer, sp. Spicules of the interior skeleton. En-
larged sixty-four times. From the Upper Chalk at Ahlten, Hanover .
Verruculina astrea, Hinde. Showing the inner or upper surface. Natural
size. From the Upper Chalk at Flamborough
. The same. Portion of the under or outer surface of the same specimen,
showing the minute pores. Natural size.
Page
34
39
39
36
BOSSE pLRON GE
_ -
oe
(o:
Ao”.
A.Gawam del. et lth
x
pals
West Newman & C2? imp
ut
{eRe
PLATE IV.
Verruculina convoluta, Quenst. sp. Natural size. From the Upper Chalk at
Flamborough, Yorkshire . . . .
. The same. Portion of the surface of the interior of the cup, showing the minute
pores. Natural size.
The same. ‘The interior surface. Enlarged four times.
. The same. Portion of the surface of the exterior of the cup, from the same spe-
cimen. Natural size.
The same. Enlarged four times.
Verruculina plicata, Hinde. Natural size. From the Upper Chalk at Flam-
borough . EE Merci cl -5. on top aoe aks 64
The same. Portion of the surface of the interior of the cup. Natural size.
The same. Enlarged four times.
c. The same. Portion of the surface of the exterior or outer wall of the same spe-
cimen. Natural size.
The same. nlarged four times.
Page
38
36
HOssit SPONGES .
Plate IV.
al 3
a utaeeteew
watet
A. Gawan del et lith.
West, Newman & C? imp.
Fig.
PLATE VY.
. Verruculina Reussii, M‘Coy, sp. A medium-sized specimen, showing the upper
surface. ‘Natural size. From the Upper Chalk at Flamborough, Yorkshire .
. The same. /A portion of the under surface of the same specimen. Natural size.
; Verruculini papillata, Hinde. The upper surface of a funnel-shaped specimen;
the margins are partly broken off. Natural size. From the Upper Chalk at
Erm 9 o ao o @ ao &
. The same. A portion of the outer or under surface of the same specimen. Natural
size.
. Stichophyma tumidum, Hinde. Natural size. From the Upper Chalk at Flam-
INNA Ss 5 6 o 8
. The same. The summit of another specimen from the same locality, showing the
apertures of the vertical canals. Natural size.
Page
40
4]
4]
HOSSIE SPONGES Plate V.
"eas
j
I
ly
t abajay yee
iF ay be .
A.Gawan del litk West,.Newman &C*° imp
5 a al. et Jith
Fig.
Rig.
. 4a.
4b.
PLATE VI.
. Jereica cylindrica, Hinde. A median vertical section showing the shallow cup and
sections of the canals. Natural size. Probably from the Upper Green Sand of
Wiltshire
. The same. Spicules of the interior skeleton. Enlarged sixty-four times. Drawn
from a transparent microscopic section of the same specimen.
. Stachyspongia spica, Roemer,sp. The upper ea of a specimen from the Grey
Chalk, near Dover. Natural size
. The same. Transverse section of another specimen from the same iGcality, showane
the central cloacal tube and sections of the horizontal canals. Natural size.
. Scytalia fastigiata, Lee, sp. Natural size. From the Upper Chalk at Flam-
borough, Yorkshire
. Scytalia radiciformis, Phill. sp. ciarenl? size. ron the Upper Chalk at Flam-
borough . : i eRe reece Sc
The same. Transverse Seber of another ecmen from Flamborough, showing
the central cloacal tube and sections of the horizontal canals which open into it.
Natural size.
The same. Spicules of the interior skeletal fibres. Enlarged sixty-four times.
From the Upper Chalk at Ahlten, Hanover.
Page
45
44
FOSSIL SPONGES. Plate VI
G.M. Herschell del. et lith. West, Newman & (2 inp
Fig.
Fig.
Fig.
PLATE VII.
. Pachinion scriptum, Roem. sp. A medium specimen from the Upper Chalk at
Flamborough, Yorkshire. Natural size .
. The same. Transverse section of another specimen from the same docality,
showing the cloacal tube and sections of the skeletal fibres. Natural size.
. The same. One of the spicules of the fibre. Enlarged sixty-four times. From
the Upper Chalk at Schweichelt, Brunswick.
. The same. Spicules of the dermal layer. Enlarged sixty-four times. From
the Upper Chalk at Schweichelt, Brunswick.
. Placonella perforata, Hinde. Showing the upper surface. Natural size. From
the Upper Jura at Nattheim, Wiirtemberg .
. The same. A small portion of the upper surface of the same specimen, dhosine
the disposition of the spicules. Enlarged sixteen times.
. The same. A single spicule from the outer surface. Enlarged thirty-two times.
. Holodictyon capitatum, Hinde. Natural size. From the Upper Green Sand at
Warminster, Wiltshire
. The same. A vertical median section of another pearl ‘com the same locality,
showing a central cloaca. Natural size.
. The same. Portion of the outer surface of the same specimen, showing the
disposition and mode of attachment to each other of the skeletal spicules.
Enlarged thirty-two times.
46
47
Plate Vil.
SPONGES.
(Oeste A ill
a
West,Newman & C° imp.
M. Suft del. et lith.
i)
PLATE VIII.
Doryderma dichotomum, Benett, sp. Natural size. From the Upper Green Sand
at Warminster, Wiltshire . . . ..-.-. .+ + «@ ee ee eaake
. The same. ‘Transverse section of a branch of another specimen from the same
locality, showing sections of the longitudinal canals. Natural size.
. The same. Portion of the surface of 1a, enlarged eight times, showing three of
the canals and the disposition of the spicular mesh.
Doryderma ramosum, Mantell, sp. Portion of a specimen from the Upper Chalk
at Oare, Wiltshire. Natural size.
. The same. Portion of the outer surface of the same specimen, enlarged sixteen
times, showing the spicular structure and the interspaces of the mesh.
Doryderma Remeri, Hinde. A fragmentary specimen inclosed in a flint from the
Upper Chalk of Wiltshire? Natural size .
. The same. Portion of the outer surface of the same specimen, showing the dispo-
sition of the spicules and the mesh-interspaces. Enlarged sixteen times.
. The same. Two of the minute trifid spicules with which the interspaces of the
mesh at the surface are filled. Enlarged sixteen times. From the Upper Chalk
at Ahlten, Hanover.
Page
47
48
49
te VIIL.
Pla
SPONGES.
BOS Si
3
P|
fe
oO
a
‘|
Le}
‘3
2
3
ww
2
, ‘
allt |
:
‘
a soi hh
ei
ey
' ;
u
> Ay
hut ay:
WS
,
a, rw hn *
art sy
vd J i
Ae
_ ae.)
, ; ‘ > aliens * iv é
a”
Fig.
Fig.
Fig.
Fig.
Fig.
aulvae
. 10.
3a.
PLATE IX.
. Doryderma Benetti, Hinde. Natural size of the body and upper portion of the
stem of a medium-sized example. From the Upper Green Sand at Warminster,
Wiltshire US eas aoe ned Lb. GQ jo fer oe
The same. Part of the transverse section of the body of another specimen from
the same locality, showing the distribution of the longitudinal canals. Natural
Size.
The same. Portion of the outer surface of a specimen, enlarged eight times, showing
the spicules and the interspaces of the mesh.
Pachypoterion robustum, Hinde. Two thirds the natural size. From the Upper
Green Sand at Pewsey, Wiltshire .
. The same. Portion of a transverse section of the lower portion of the gay of
another specimen from Pewsey, showing in section the canals which run in a
generally vertical direction to the upper surface of the cup, and the appearance
presented by the radial canals which run in a slightly arched direction from the
outer surface to the centre. Natural size.
The same. Portion of the spicular mesh of the interior as seen in a polished
transverse section. Enlarged sixteen times.
. The same. Portion of the spicular mesh of the stem of another specimen from
Pewsey. Enlarged sixteen times.
Pachypoterion compactum, Hinde. The cup and the upper portion of the stem of
a specimen from Warminster. Natural size. From the collection of William
SHON” Gy 1G Sgn Op Gl eot yok SOG. 5 GG aio 6 Shae toe
The same. Portion of the outer surface of the same specimen, enlarged sixteen
times, showing traces of the spicules and the mesh interspaces.
Page
49
Plate Ix.
SPONGES.
FOSSIL
p
an &C°im
West Newm
M. Suft &2GMHerschell del.et Lith
5 ha,
ig. Le.
wo
. 26.
. od.
5 BD:
PLATE X.
. Nematinion calyculum, Hinde. A specimen, imperfect at the base, from the Upper
Green Sand at Warminster, Wiltshire. Natural size eae. 3
The same. ‘Transverse section of the stem of another specimen, showing traces of
the horizontal canals. Natural size.
. The same. Portion of the surface of fig. 1, enlarged eight times, showing traces
of the spicular structure.
The same. Portion of the imterior of the stem, enlarged sixteen times, showing
the character and disposition of the spicules. Drawn from the polished surface
of a vertical section.
Heterostinia obliqua, Benett, sp. The outer surface of an imperfect cup-shaped
specimen with numerous root-processes. Natural size. Inclosed in a flint from
the: Upper) Challsof Wiltshire =) stun ene
. The same. Showing the semipalmate stem and a portion of the expanded body-
plate of another flint-inclosed specimen from Wiltshire. Natural size.
The same. Portion of the outer surface of a specimen, in which the dermal layer
has been preserved, showimg its spicular structure. From the interior of a Chalk
flint from Wiltshire. Enlarged sixteen times. The specimen belongs to the
Jermyn-Street Museum.
. The same. Portion of the surface of a specimen, in which no dermal layer has
been preserved, showing the spicular structure of the interior of the wall of the
sponge. Enlarged sixteen times.
. Isoraphinia texta, Roemer, sp. From the Upper Chalk at Flamborough, Yorkshire.
Natural size 5 Slee 5) 358 nee tet. {reine eee: eee
The same. ‘Transverse section of another specimen from the same locality, showing
the central cloacal tube. Natural size.
The same. Portion of the outer surface of fig. 3, showing traces of the exterior
layer of spicules. Enlarged seven times.
. Mastosia neocomiensis, Hinde. Detached spicules of various sizes. Enlarged
twenty-six times. From the Lower Green Sand at Haslemere, Surrey
Page
54
53
or
HOS She “SPONGES. Plate X
ate
i Fielding &35.C.Woodward del et lth
man &C° im
Fig.
Fig.
Fig.
Fig.
Fig.
15.
.oa.
36.
4.
4a.
46.
PLATE XI.
. Phymatella reticulata, Hinde. A club-shaped example, imperfect at the base.
Two-thirds the natural size. From the Upper Chalk at Flamborough, York-
1 i er ae een ee hy OR
. The same. Portion of the surface of another specimen from Flamborough,
showing, after treatment with acid, apertures of the canals which open at the
surface and traces of the interior spicules, which have been in part replaced by
calcite. Enlarged eight times.
The same. Microscopic section of the interior, showing traces of the spicules.
The outline of the spicules is less definite than represented. Enlarged twenty-
four times. Drawn from a transparent section.
. Phymatella nodosa, Hide. An imperfect specimen from the Upper Green Sand
at Warminster, Wiltshire. Natural size
. The same. Portion of the outer surface, showing traces of Tetracladine winnie
Enlarged sixteen times.
. Callopegma obconicum, Hinde. A specimen inclosed in a flint. From the Upper
Chalk of the South of England. Natural size
The same. A vertical median section of a specimen, eee one an a half Ga
showing the general disposition of the canals and the spicular mesh.
The same. Portion of the interior spicular mesh of 3a. The spicules have been
replaced by iron peroxide, and the nodes formed by the junction of the spicular
rays are consequently imperfect. Enlarged thirty-two times.
Callopegma ficoideum, Hinde. A specimen inclosed in a flint. From the Upper
Chalk at Guildford, Surrey. Natural size :
The same. Portion of the outer surface of fig. 4, exbsroeil sixteen times, Hipeae
the extended heads of compound trifid spicules of the dermal layer.
The same. A fragmentary spicule of the interior skeleton. Enlarged sixteen
times.
Page
59
61
61
Plate XI.
FOSSIL SPONGES.
5\ SS)
“5
SHS esl
AGS
s)
st Newman & C® imp
Wes
G M. Herschel! del. et lith.
aelaas
.oa.
ig. 3b.
PLATE XII.
. Aulazxinia costata, Hinde. A silicified specimen in which the vertical canals have
been infilled with silica and now appear as solid ridges. Natural size. From
flint gravel at Stanway, Gloucestershire Sch ce ine cone cae
The same. Portion of the outer surface of fig. 1, shows casts in flint of the
spicular skeleton. Enlarged eight times.
Trachysycon sulcatum, Hinde. A vertical median section of a silicified specimen,
showing sections of the funnel-shaped cloaca and traces of the canals. Two-
thirds the natural size. From the Upper Chalk of the South of England
. The same. Portion of the outer surface of fig. 2, showing traces of the ridges and
broken spines. Natural size.
The same. Portion of the outer surface, enlarged sixteen times, showing traces of
spicules of the dermal layer.
Trachysycon nodosum, Hinde. A vertical median section of a silicified specimen,
showing the interior of the cloaca and the course of the canals opening into it.
Natural size. From the Upper Green Sand at Warminster, Wiltshire
The same. Portion of the outer surface of the same specimen, showing the dome-
shaped elevations and the apertures of canals. Natural size.
The same. Showing the interior spicular skeleton, drawn from a polished surface.
Enlarged thirty-two times.
Page
60
63
62
FOSSIL SPONGES.
Plate aly.
A.Gawan del. et lith.
West Newman & C°imp
Fig. 1
Fig. la.
Fig. 18.
Tangs,
Fig. 2a
Fig. 28.
Fig. 2c.
Fig. 3.
Fig. 3a.
=
Fig. 4.
Fig. 4a
PLATE XIII.
. Hindia fibrosa, Roemer, sp. The interior of a fractured specimen, showing the
radial disposition of the canals. Natural size. The sponge has been silicified,
and the canals now appear as minute straight rods. From Silurian strata of
the Niagara group in Perry County, Tennessee c — uae
The same. Portion of the interior of another silicified specimen ‘from the same
locality. Enlarged eight times. The rods represent the infilled canals, and the
tubercular projections on them are produced by the siliceous infilling of the inter-
spaces between the spicular arms of the interior skeleton.
The same. Portion of the spicular skeleton of the interior of a specimen from
Silurian strata of the Lower Helderberg group at Dalhousie, New Brunswick.
Enlarged seventy-six times. Drawn from a transparent microscopic section by
means of the camera lucida.
Siphonia tulipa, Zitt. Natural size. From the Upper Green Sand at Warminster,
Wiltshire
. The same. A vertical median section of another specimen from Warminster,
showing the cloaca, the course of the vertical canals which open into it, and
traces of the radial canals which extend from the surface towards the centre of
the sponge. Natural size.
The same. Transverse section of a specimen taken from the lower portion of the
body, immediately below the cloacal tube, showing sections of the larger vertical
canals and the smaller radial canals. Natural size.
The same. Portion of the interior spicular skeleton of 2a. Enlarged thirty-two
times. Drawn from a transparent section. The small square to the right of
the figure represents its natural size.
Siphonia ficus, Goldfuss. Natural size. From the Grey Chalk at Dover
The same. Portion of the outer surface of the same specimen, enlarged eight
times, showing the apertures of the radial canals and traces of the spicular
skeleton.
Siphonia Kénigi, Mant. sp. Natural size. From the Upper Chalk at Flamborough,
Yorkshire
. The same. Portion of the outer surface of the same specimen, enlarged thirty-two
times, showing the fragmentary heads of spicules of the dermal layer.
Page
64
65
65
FOSSIL SPONGES. Plate XII.
M. Sufts GM Herschel) del. et ith. West Newnan & C° imp
ei i
N
anne Miwiiig Vali
;
a
4 ; ate ¥ i
ig. 1d.
Fig.
Fig.
Fig.
5 Lae
~ Wa:
mule:
le.
w©
PLATE XIV.
Hallirhoa costata, Lamx. A small three-lobed specimen from the Upper Green
Sand at Warminster, Wiltshire. Naturalsize ......... =.
The same. A conical, partially five-lobed, example from the same locality, showing
the stem and the root-like extensions at its termination. Natural size.
The same. A four-lobed specimen from Warminster, showing a partial division in
each of the principal lobes. Two-thirds the natural size.
The same. A vertical median section, showing the shallow cloaca and the course
of the larger canals opening into it. Natural size.
The same. A transverse section of a five-lobed example, taken below the level of
the cloaca, showing in section the larger vertical canals. Drawn from a polished
specimen presented to the Museum by Robert Etheridge, sen., Esq., F.R.S.
The same. Portion of the interior spicular mesh, enlarged sixty-four times.
Drawn from a transparent microscopic section by means of the camera lucida.
The small square to the left indicates the natural size.
Hallirhoa costata, var. brevicostata, Mich. From the Upper Green Sand at War-
minster. Two-thirds the natural size
. Hallirhoa costata, var. elevata, Hinde. Two-thirds the natural size. From War-
minster
Page
67
68
68
BPOSSiIE SPONGES.
Plate XIV.
st Newman & C? ump
M. Suft & G M.Herschell del. et lith
F ee 0
et Los &
5 kG
BALE
PLATE XV.
. Hallirhoa agariciformis, Benett, sp. Showing the upper surface and the open
canals extending downwards from the cloaca. Natural size. From the Upper
Green Sand at Warminster, Wiltshire ; Pa e
The same. ‘Two spicules from the interior mesh of panier w arminster specimen.
Enlarged seventy-two times. Drawn from a transparent section by means of
the camera lucida.
The same. Portion of the outer surface of a specimen in which the dermal layer
has been preserved, showing the laciniated disciform heads of its component
spicules. Knlarged thirty-two times. The specimen is from Warminster, and
belongs to the Jermyn-Street Museum.
Jerea reticulata, Hinde. A specimen from the Grés Vert at Rettsel, Ardennes,
showing the reticulate disposition of the canals on the outer surface. Natural
size. The canals in this specimen have been infilled with siliceous material,
and now appear as solid fibres. . . . . .. . ae
1. The same. The summit of the same specimen, showing the close arrangement of
the canal-apertures. Natural size.
The same. A vertical median section of a specimen from the Upper Green Sand
at Warminster, showing the disposition of the radial canals. Natural size.
3. Jerea cordiformis, Hinde. From the Grey Chalk near Dover. Natural size
. The same. A vertical median section of the same specimen, showing the direction
of the vertical and radial canals. Natural size.
Page
69
70
71
HOSsil SPONGES.
re
*
*-
»
awam del.et hth
G
A.
PREAH eva:
Page
. Nelumbia tuberosa, Hinde. From the Grey Chalk near Dover. Natural size. . 72
. The same. A transverse section from the central portion of fig. 1, showing the
disposition of the vertical canals and faint traces of the radial canals. Natural
size.
. The same. A portion of la, enlarged thirty-two times, showing traces of the
spicular skeleton. The original siliceous skeleton has been replaced by calcite,
and where the spicules are in contact their forms have been indistinguishably
merged together ; occasionally, however, the separate rays can be recognized.
A section of a Foraminifer is also shown in the matrix of grey chalk.
. The same. Portion of the dermal layer of a specimen from the Upper Chalk,
enlarged thirty-two times, showing the heads of large compound trifid spicules
and traces of numerous smaller ones, and also of the mesh-spicules beneath.
. Polyjerea arbuscula, Hinde. From the Grey Chalk near Dover. Natural size. . 738
. The same. Transverse section of one of the branches of the same specimen,
showing the distribution of the longitudinal canals and faint traces of the spicular
skeleton, which is now calcareous. The sponge is preserved in a matrix of
Grey Chalk.
Polyerea lobata, Hinde. Natural size. From the Grey Chalk near Dover. . . 73
Plate XVI.
HOSSIL SPONGES.
West Newman & C? imy
A, Gawan del. et lith.
Fig.
Fig. 5a.
‘) 5
Fig. 5d.
5 War
14.
w
PLATE XVII.
. Bolospongia globata, Hinde. Natural size. From the Upper Chalk at Flam-
borough, Yorkshire’: 7 3002192)? eae) = ete ee ee oe
The same. Vertical section of another specimen from the same locality. The
dotted portions indicate the spicular structure, and the interspaces represent the
wide channels and loculi of the interior of the sponge. Natural size.
The same. Portion of 1a, enlarged thirty-two times, showing in the lighter
portions traces of the spicular structure. The spicules have been replaced by
crystalline calcite and silica, and are only recognizable where they were not in
close contact with each other originally. The square below the figure idicates
its natural size. Drawn from a transparent section by means of the camera
lucida.
Bolospongia constricta, Hinde. Natural size. From the Upper Chalk at Flam-
borough . tN tne Soe Pa aes sods ao ede eae
The same. A transverse section of the same specimen. Natural size.
Thecosiphonia nobilis, Reem. In the lower portion of the specimen traces of the
smooth dermal layer are shown. Natural size. From the Upper Chalk of
Wiltshire. The specimen figured belongs to the Jermyn-Street Museum
. The same. Portion of the interior spicular skeleton, enlarged thirty-two times, as
shown on the upper surface of the same specimen.
Thecosiphonia turbinata, Hinde. A silicified specimen, from the Upper Chalk at
Stockton, Wiltshire, showing some of the canals extending down the upper
surface, and traces of the corrugated dermal layer beneath. Natural size
Thamnospongia glabra, Hinde. A single branching stem, forming part of a large
bushy mass. Natural size. Inclosed in the cavity of a flint from the Upper
Chalk at Berkhampton . . ... . . : RPE te oh th, eG Voda.
The same. Portion of the interior spicular structure immediately beneath the
dermal layer, showing the tuberculated spicules. Enlarged sixty-four times.
The same. Portion of the dermal layer of fig. 5, enlarged thirty-two times. The
dark figures represent the hollow moulds of the larger spicules of the dermal
layer. The two lighter figures below represent the spicules themselves. The
entire dermal layer is formed of similar spicules; but, as a rule, only the larger
forms can be distinguished.
The same. Portion of the dermal layer of a specimen, enlarged thirty-two times,
showing slightly elevated spaces with groups of circular pores. From Upper-
Chalk detritus at Budleigh Salterton, Devonshire. Presented to the Museum
by Mr. H. J. Carter, F.R.S.
Page
74
74
75
75
79
Plate XVII.
IOVS Se, SHskonrmeaaisy,
E
West Newman & C° imp.
M. Suft & G M.Herschell del. et lith.
g. 3a.
ig. 3b.
PLATE XVIII.
. Turonia variabilis, Mich. sp. Natural size. Showing the furrowed upper surface
and the smooth dermal layer of the base. ee from the Senonian of
France
. Thamnospongia laneliaee: Benet sp. e specimen elneed in Ratar From the
Upper Chalk at Chicklade, Wiltshire. Natural size
. The same. Another flint-inclosed specimen, showing traces of ieee canals
on the fractured end of one of the branches. Natural size. From the Upper
Chalk at Stockton, Wiltshire.
. The same. Portion of the outer surface of a specimen from Stockton, showing
some of the larger heads of the compound trifid spicules of the dermal layer.
Enlarged thirty-two times.
Thamnospongia? reticulata, Hinde. From the Grey Chalk near Folkestone.
Natural size : - : : : oe i
The same. Transverse section of one of the stems, showing the central camel said
traces of the sinuous canals which open into it. Natural size.
The same. Portion of a transparent section from the interior of the sponge,
showing the disposition of the spicular skeleton. Enlarged thirty-two times.
The original silica of the spicules has been entirely replaced by crystalline
calcite.
. Kalpinella patereformis, Hinde. A vertical median section, showing the disposition
of the canals in the sponge-wall. From the Upper Green Sand at Warminster,
Wiltshire. Natural size
. Pholidocladia ramosus, Hinde. Showing the empty moulds of the branches of a
specimen in a flint from the Upper Chalk. Natural size .
. The same. A fragment of one of the branches from the interior of ge flint.
Natural size.
. The same. Portion of the surface of 5 a, enlarged sixty-four times, showing the
characters of the spicular skeleton, The small square at the top of the figure
indicates its natural size.
Page
76
79
80
77
81
1 OPS SE, SSHEHOMNKe date Plate XVII.
West, Newman & C?imp
M. Suft & G.M Herschell del. et ith
Fig.
Fig.
w
. 3a.
4b.
4c.
PLATE XIX.
. Kalpinella patereformis, Hinde. Two-thirds the natural size. From the Upper
Green Sand at Warminster, Wiltshire . . .. .
. The same. Transverse section of the stem, showing the distribution of the canals.
Natural size.
. The same. Portion of the interior spicular skeleton, Enlarged sixty-four times.
Drawn from a transparent section by means of the camera lucida.
. Kalpinella rugosa, Hinde. ‘Two-thirds the natural size. From the Upper Green
Sand at Warminster .
. Ragadinia compressa, Hinde. Natural size. From the interior of a flint from the
Upper Chalk at Huish, Wiltshire . Mere ee yo pS
The same. Two detached spicules, enlarged sixty-four times, from the Upper Chalk
at Horstead, Norfolk, The spicules on the outer surface of fig. 3 resemble these
forms, but they are not sufficiently clear to be shown.
. Ragadinia clavata, Hinde. Natural size. From the interior of a flint from the
Upper Chalk of Wiltshire. The specimen belongs to the Jermyn-Street
Museum .
. The same. A small club-shaped specimen, natural size, also from the interior of
a flint.
The same. A portion of the surface of fig. 4, enlarged sixty-four times, showing
the spicular skeleton immediately beneath the dermal layer.
The same. Portion of the surface of fig. 4 where the dermal layer has been pre-
served, showing the complete head of one spicule and fragments of others.
Enlarged thirty-two times.
Page
77
78
82
84
Plate XIX.
HOS Sta SPONGES.
West Newman & C° imp
A.Gawan del et lith.
Fig.
Fig.
.
Fig.
Vig.
Fig.
peli:
.oad.
4a.
5D.
PLATE XX.
. Ragadinia sulcata, Hinde. An imperfect specimen inclosed in a flint from the
Upper Chalk at Beckhampton, Wiltshire. Natural size
The same. Portion of the outer surface of another specimen in high Eile diewinal
layer has been preserved, showing its component spicules. Enlarged thirty-two
times.
. Plinthosella squamosa, Zitt. Natural size. From the interior of a flint. The
spicular structure has been completely dissolved, and only the empty vermi-
culate moulds of the sciaie remain. Presented to the Museum by Mr. H. J.
Carter, F.R.S.
. Plinthosella ee fails, Natural size. Preserved in the interior of a flint
from the Upper Chalk of Wiltshire. The specimen belongs to the Jermyn-
Street Museum . ; eet ig 8e,
The same. Portion of wee outer sires of the same specimen, exilataed chirty-tmp
times, showing the spicular structure.
. Plinthosella nodosa, Hinde. A specimen inclosed in a flint from the Upper Chalk
of Wiltshire. Natural size. The ie ae to the Jermyn-Street
Museum . : : : :
The same. Portion of tite outer ve of tite same specimen, os un pean
scales of the dermal layer. Enlarged thirty-two times.
. Pholidocladia dichotomus, Hinde. An imperfect specimen preserved in a flint from
the Upper Chalk of Wiltshire. Natural size. The specimen belongs to the
Jermyn-Street Museum .
. The same. Portion of the outer Bee ats a ee of fig. 5 where the dermal
layer is absent, showing the spicular structure of the interior skeleton. Enlarged
sixty-four times. The square beneath the figure represents its natural size.
The same. Portion of the outer surface of another branch of fig. 5, on which the
dermal layer has been retained, showing its component spicules. Enlarged
thirty-two times.
Page
83
85
85
86
81
Plate XX.
FOSSInN SPONGES.
Vest Newman & C°imp
—
M. Suft & G.M. Herschell del. at ith
7 oni? net
mee ee
AY r ty
Fig.
Fig.
Fig.
3a.
PLATE XXI.
. Phymaplectia irregularis, Hinde. Natural size. Inclosed in a flint from the Upper
Chalk of Wiltshire . .. .
. The same. Portion of the surface of the same specimen, enlarged thirty-two times,
showing the heads of trifid spicules of the dermal layer.
. Phymaplectia spinosa, Hinde. Natural size. Inclosed in a flint from the Upper
Chalk of Wiltshire. The specimen belongs to the Jermyn-Street Museum .
. The same. Portion of the surface of fig. 2 where the dermal layer is absent,
showing the spicular structure of the sponge-wall. Enlarged thirty-two times.
. The same. Portion of the surface of fig. 2 where the dermal layer has been
preserved, showing some of the heads of its component spicules. Enlarged
thirty-two times.
. Phymaplectia cribrata, Hinde. Natural size. From the interior of a flint from
the Upper Chalk at Oare, Wiltshire. The specimen belongs to the Jermyn-
StreetiMuseum:, ©. wa: foo see eeeet ke) SE eg ges Re te Ss eC
The same. Portion of the outer surface of fig. 3, enlarged thirty-two times, showing
some of the minute spicular heads and the pores of the dermal layer.
Page
87
88
88
FOSSIL SPONGES. eee
West, Newman & C? imp
Mi. Suft & GM. Hexschell del et lth.
hig.
Fig.
Fig.
la.
1b.
PLATE XXII.
Page
. Phymaplectia scitula, Hinde. Natural size. From the interior of a flint from the
Upper Chalk at Oare, Wiltshire. The specimen belongs to the Jermyn-Street
Museum. i. x Ps) ee co ate ee ee we eee
The same. Portion of the surface of the interior of fig. 1, enlarged six times,
showing open canals radiating from slightly elevated centres.
The same. Portion of the exterior surface of fig. 1, enlarged thirty-two times,
showing the spicular structure of the sponge-wall. The square on the left side
of the figure represents its natural size.
. Rhopalospongia gregaria, Benett, sp. Two-thirds the natural size. From the
Upper Green Sand at Warminster, Wiltshire. . . . a ait ee ae: CW
. The same. A vertical section of another specimen from Warmiaeter showing
traces of the radiating canals and concentric markings of growth. Natural
size.
The same. A transverse section of 2a. Natural size.
The same. he spicular structure of the interior. Enlarged sixty-four times.
Drawn from a transparent section by means of the camera lucida.
Rhopalospongia obliqua, Hinde. Natural size. From the Upper Green Sand at
Warminster. . . mutha oy oe 0 : c so ETD
. The same. A transverse section of fig. 3, own me attritnation of the oaihiala:
Natural size.
HOS | SiON GE Ss. Plate XXII.
rao uy fot
wman& C?
M. Suft & GM. Herschell del. et lith. West, Newman imp
g. 10.
rns)
g. 35.
PLATE XXIII.
. Astylospongia? Remeri, Hinde. An imperfect silicified specimen, showing the
disposition of the surface-canals and the apertures of the radial canals. Natural
size. Probably from Silurian strata .
. The same. A transverse section of fig. 1, showing the course sor ae radial earale!
Natural size.
The same. Showing the spicular structure of the interior, enlarged sixty-four
times. Drawn from a transparent section by means of the camera lucida.
. Craticularia Fittoni, Mautell, sp. An imperfect specimen from the Grey Chalk
near Dover. Natural size .
. The same. Portion of the outer saris of a specimen schon the Chalk Marl near
Ventnor, Isle of Wight, showing the character of the dermal layer. Enlarged
sixty-four times.
. The same. A fragment of the spicular mesh of the interior of the wall. Enlarged
sixty-four times.
Strephinia convoluta, Hinde. An imperfect specimen from the Grey Chalk near
Dover. Natural size.
. The same. A transverse section of a norton of the ea shone its ttiicknese
and the disposition of the canals. Natural size. The shaded portion of the
figure represents the spicular wall, and the lighter parts the canals. In places
the canals appear to extend quite through the wall, but this is owing to the
thinness of the wall at the extremity of the blind canals, so that it readily breaks
away when the surface is polished.
The same. Portion of fig. 3a, enlarged sixty-four times, showing the spicular
structure of the wall. The spicules have been entirely replaced by calcite of a
tint slightly darker than the chalky matrix.
. Strephinia reteformis, Hinde. Natural size. Showing the reticulate character of
the outer surface. From the Grey Chalk near Dover .
. The same. A vertical median section of fig. 4, showing the thickness of the wall
and the disposition of the canals. The spicular structure of this specimen, like
that of S. convoluta, has been replaced by calcite, and the walls at the extremity
of the canals have been partially broken away.
Page
94:
96
i
HOSSIN SPONGES. Plate XXII.
A.Gawan deL et lith
West, Nevanan & C° amp
—
— 7 ¥ se
0 a a ~
J
PLATE XXIV.
Page
. Stauronema Carteri, Sollas. The lower portion of an imperfect specimen from
the Upper Green Sand at Cap la Héve, near Havre, showing the inner concave
surface. Naturalsize . . . : Aa tetra memantine oo!)
. The same. Showing the convex outer surface of the same specimen.
. The same. Portion of the spicular mesh of the interior of the wall. Enlarged
thirty-two times.
. Stauronema planum, Hinde. The upper surface of a specimen, showing the canal-
apertures, which in places are partially obscured by a delicate dermal layer.
Natural size. From the Grey Chalk near Folkestone . . . . . . . . . 100
2a. The same. The under surface of part of fig. 2. Natural size.
2b, The same. A vertical median section of the same specimen, showing traces of the
5
35.
canals and the spicular mesh. Natural size.
. The same. Portion of the polished surface of 26. Enlarged thirty-two times.
The dark lines forming the squares represent the canals of the mesh-spicules,
which have been infilled with an opaque material, probably peroxide of iron.
. Verrucocelia vectensis, Hinde. From the Chalk Marl at Ventnor, Isle of Wight.
INatoralisizé. (<4) Gt pene’ . eS ee Se ee ce
The same. Portion of the outer surface of fig. 3, enlarged thirty-two times,
showing the structure of the dermal layer.
The same. Portion of the spicular mesh of the interior of the wall. Enlarged
thirty-two times. Drawn from a fractured surface of one of the projecting
tubes.
XIV,
Plate
FOSSIE SPONGES.
Slr
Newman & C? amp
West
. ot lith
+
M. Suft & G. M. Herschell
PLATE XXV.
Page
Fig. 1. Stawronema compactum, Hinde. The lower portion of a specimen from the Grés
vert, France ?, showing the front surface. Naturalsize . . .... . .4101
Fig. la. The same. Showing the outer surface of the same specimen. The canals in the
lower portion are concealed by the dermal layer. Natural size.
Fig. 14. The same. The fractured upper margin of the same, showing the thickness of the
wall and traces of the canals. Natural size.
Fig. le. The same. Portion of the dermal layer of the outer surface. Enlarged fifteen
times.
Fig. 1d. The same. A fragment of the interior spicular mesh. Enlarged sixty-two times.
Drawn by means of the camera lucida.
Vig. 2. Sestrodictyon convolutum, Hinde. Natural size. From the Alpine Chalk (= Upper
Green Sand) at High Sentis, Canton Appenzell, Switzerland. . . . . . . 102
Fig. 2a. The same. Fragments of the interior spicular mesh of fig. 2. Enlarged sixty-two
times. The dark lines in the lower portion of the right-hand fragment represent
canals which have been infilled with iron peroxide.
Fig. 3. Guettardia radians, Hinde. The lower portion of an imperfect specimen from
Cretaceous strata at Biarritz, France. Naturalsize ....... . . 105
Vig. 4. Trochobolus constrictus, Hinde. Natural size. From the Upper Jura at Randen,
Shisiy.od Chie hh ee oe a Pec oP o's 6 Go ge oo oo NOs
Vig. 4a. The same. Portion of a vertical median section of fig. 4, natural size, showing the
thickness of the wall.
Vig. 4b. The same. Portion of the polished surface of 4a, enlarged thirty times, showing
the spicular mesh and the octahedral character of the nodes. The spicules have
been replaced by calcite, and are consequently indistinct.
Vig. 5. Ventriculites convolutus, Hinde. An imperfect specimen, preserved in chalk,
showing both the outer and inner surfaces of the wall. Natural size. From
the Upper Chalk at Broadstairs, Kent sd 2 Be ps LO
Fig. 5a. The same. Portion of the outer surface of fig. 5, enlarged four times, showing the
flattened ridges of the wall and the narrow interspaces.
FOSSIL SPONGES. Plate XXV.
fy
ct a a IT cl.
West, Newman & 0° imy
M. Suft & G.M.Herschell del. et lith
ig. la.
. 4a.
45.
. The same. Part of a transverse section of another specimen a Tlaniberouees
PLATE XXVI.
. Ventriculites infundibuliformis, S. Woodward. An imperfect specimen, partially
imbedded in the chalky matrix. Natural size. From the Upper Chalk of the
South of England . Memoicr chile j :
The same. Transverse section of a specimen preserved 4 in flint, Shaun the ape
sition of the folds of the sponge-wall. Natural size. From the Upper Chalk of
the South of England.
. Ventriculites cribrosus, Phillips, sp. An imperfect a from the Upper Chalk
. 113
at Flamborough, Yorkshire. Natural size.
showing the folds of the wall. Natural size.
. Ventriculites angustatus, F. A. Roemer, sp. An imperfect silicified specimen from
4 . 114
. The same. The upper portion of fig. 3, showing the thickness of Re wal aa ihe
the Upper Chalk, South of England. Natural size .
canal-apertures of its inner surface. Natural size.
_ The same. A small portion of the outer surface of fig. 3, enlarged eight times,
showing the canal-apertures and the perforated dermal layer.
. Sporadoscinia capax, Hinde. An imperfect example from the Lower Chalk, two-
thirds the natural size, showing a fragment of the sponge-wall, with some oysters
attached to a part of its surface, and impressions of the ane of the
mner or cloacal surface. From the South of England .
The same. Portion of the outer surface of fig. 4. Enlarged eight times.
The same. Portion of the interior of the wall of fig. 4, enlarged thirty times, showing
traces of the spicular mesh. Drawn from a fractured surface. The spicules have
been completely dissolved, and only the empty moulds in the chalky matrix
remain.
Page
LG
Plate XXVI
IOUS SUE, 7 SWEIOENMe ase
West,Newman & C* imp
M.Suft & G.M.Herschell del. et lith
=) ie ie ‘"
par Peli” &
Fig.
Fig.
ig. 1d.
ig. 4c.
)
wa.
3b.
. 4a.
ig. 4b.
PLATE XXVII.
Sestrocladia furcatus, Hinde. Natural size. From the Grey Chalk, near Dover .
. The same. A transverse section of one of the branches, showing the thickness of
the walls. Natural size.
The same. Portion of the interior of the wall, showing traces of the spicular mesh
with the octahedral nodes. The spicules have been replaced by calcite, and are
partially obliterated. Enlarged thirty times.
Polyblastidium racemosum, T. Smith, sp. A specimen preserved in flint, from the
Upper Chalk of Kent, showing on the outer surface transverse sections of the
bud-like tubular projections. Natural size
the foldings of the sponge-wall and traces of the spicular mesh, which is par-
tially preserved in iron peroxide.
Cephalites bullatus, T. Smith. A specimen partially imbedded in Chalk, showing
the characters of the outer surface, and, where this has been removed, traces of
the interior folds. The structure is now preserved in iron peroxide. Natural
size. From the Upper Chalk of the South of England . oo:
The same. The summit of the same specimen, showing the cloacal aperture and
one or two of the canal-apertures which open into it, and also sections of the
wall-plaits in the interior of the sponge. The dermal layer which covered the
summit originally is not preserved. Natural size.
The same. Fragments of the spicular mesh forming the wall-plaits. The spicules
have been replaced by iron peroxide, and their outlines are very imperfectly
preserved. Enlarged thirty times.
Placotrema cretaceum, Hinde. A specimen partially imbedded in chalk, showing
the upper surface with the apertures in the dermal layer. Natural size. From
the Upper Chalk of Kent dG. SG. c Perc Fors
The same. Portion of a transverse section of another specimen, natural size,
showing the reticulate disposition of the interior wall-plaits, which are shown
by markings in iron peroxide.
The same. Portion of the dermal layer of the upper surface of the sponge, enlarged
fifteen times, showing the spicular structure. The spicules have been completely
dissolved, and are only represented by hollow moulds in the chalky matrix.
The same. Portion of the spicular mesh of the interior wall-plaits, enlarged
thirty times. Only the empty moulds of the spicules are preserved. Drawn
from a fractured surface.
Page
117
a 5 al tae oes - 119
. The same. Transverse section of one of the tubes, enlarged four times, showie
J
BOSS iin | SOG S . Plate XXVII.
West, Newman & C° imp
M. Suft & G.M.Herschell del. et lith
irons
( sity flu; ;
>” Bee
1
7 te iit ”
ae ay in 4a!
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Fig. la.
Fig. 1 8.
Fig. 1c.
Fig. 1d.
Fig. 40
Fig. 4c.
PLATE XXVIII.
Cincliderma quadratum, Hinde. The upper portion of a funnel-shaped specimen,
from the Upper Chalk of the South of cae showing the dermal layer.
Natural size . ee dp os,
The same. ‘The basal action os a specimen. oNataeal size.
The same. A median vertical section of 1 a, natural size, showing the disposition
of the anastomosing wall-plaits in the interior of the sponge.
The same. Portion of the spicular structure of the interior wall-plaits, enlarged
thirty times. The spicules have been dissolved and only empty moulds remain.
The same. Portion of the dermal layer of 1 a, enlarged fifteen times. The spicules
are only indicated by hollow moulds in the chalky matrix. The minute spicules
forming the layer between the framework of larger spicules are too small and
too thickly intermingled together to be clearly delineated.
. Protospongia fenestrata, Salter. A fragment of a specimen preserved in indurated
black shale. Enlarged five times. From Menevian strata at St. David’s, South
Wales. The spicular framework is now composed of iron pyrites; it has evi-
dently been distorted by pressure from its original rectangular form. The
figure is drawn from Salter’s type specimen
stone, natural size. From the Chemung group of the Upper Devonian at
Cohocton, Steuben County, Western New York .
at Folkestone, showing the disposition of the tubes at the surface. Natural
RIZE Nor ee eid els Cle toed to eee ae ne Eee
showing the arrangement of the wall-plaits of the interior of the sponge.
Natural size.
. The same. A fragment of the spicular mesh of the interior of the wall-plaits,
enlarged thirty times.
The same. A portion of the dermal layer of the wall, enlarged thirty times.
Page
- 128
=. is Soret 1)
. Dictyophyton tuberosum, Conrad, sp. An imperfect specimen nieeryea im sand-
. 130
. Plocoscyphia fenestrata, T. Smith, sp. A specimen from ae Upper Gere Seu
>: (Ee Sido Genet ee eel:
. The same. A transverse section of a smaller specimen from the same locality,
HOSSIM SPONGES. Plate XXVII.
West Newman & C° imp
M. Suft € G.M.Herschell del. et lith.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
3a.
36.
6.
6a.
PLATE XXIX.
. Celoscyphia sulcata, Tate. Natural size. From the Spongarian zone of the
Chloritic Chalk (Tate) (= Upper Chalk), Whitehead, near Belfast. The speci-
men belongs to the Jermyn-Street Museum
. The same. The upper surface of the central stem of fig, I, evil ged time times,
showing the foldings of the wall and traces of the spicular mesh.
Plocoscyphia labrosa, T. Smith, sp. Natural size. An average specimen from the
Upper Green Sand near Folkestone, showing the disposition of the folds of the
wall ea oe 5. os aS a eae!
Plocoscyphia ehinutaas Hinde. A specimen from the Upper Green Sand at East-
bourne, partially imbedded in a marly matrix, showing in the upper portion the
apertures of several tubes with the canal-openings in their walls, and in the
lower portion sections of the tube-walls. Natural size
The same. A fragment of the dermal layer of a specimen from ihe Chioritic Marl
at Rocken End, Isle of Wight, showing one or two of the canal-openings and the
smaller apertures of the spicular mesh. Enlarged thirty times.
The same. Three fragments of the spicular mesh of the interior of the wall of 3a,
enlarged thirty times, showing the octahedral or lantern nodes, and traces of
canals in some of the spicular arms. Drawn by means of the camera lucida.
Plocoscyphia flecuosa, Mant. sp. A fragment of the spicular mesh of the interior
of the wall, enlarged thirty times, showing the octahedral nodes. The siliceous
structure of the specimen has been completely replaced by iron peroxide. From
the Upper Chalk of the South of England. . 0
Plocoscyphia labyrinthica, Mant.sp. A specimen ‘muheadeae ina a flint Ean ee
on its upper surface, in section, the anastomosing folds of the wall, indicated by
markings in iron peroxide. Natural size. From the Upper Chalk at Offham,
Kent . Mee ee ee ee oe dc 86 ol
Plocoscyphia vagans, Hinde. A nodule of chalk, showing on its surface the
anastomosing wall-plaits. Natural size. te Coe, ee
The same. A fragment of the spicular mesh of fig. 6, enlarged thirty times,
showing the octahedral nodes. The spicular mesh is now in the condition of
iron peroxide, and its surface is rough and uneven.
Toulminia obliqua, Hinde. A median section of a specimen from the Upper Chalk,
showing in section the disposition of the interior wall-plaits. Natural size .
The same. A transverse section of fig. 7, showing the wall-plaits as seen from
above. Natural size.
. The same. The summit of fig. 7, showing the wall-plaits, which are exposed by
the removal of the dermal layer. The walls in this specimen are indicated by
markings in iron peroxide.
Page
. 118
133
. 136
. 139
Plate XXIX.
SHEN ONC ISS
FOSSIL
imp
st Newman & (
et lith
M. Suft & GM Herschell de
Reerh.1'
sf
cmaleae
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
Fig.
ig. 1b.
Aa.
4b.
OL
2 Das
PLATE XXX.
. Toulminia jurassica, Hinde. Natural size. Showing the weathered surface of a
- 139
specimen from the Upper Jura at Randen, Switzerland
The same. A transverse section of the same specimen, showing the cloacal cavity
and disposition of the anastomosing walls in the interior. Natural size.
The same. A portion of the polished surface of 1 a, enlarged thirty times, showing
the spicular structure of the walls. The spicules have been replaced by calcite,
and only faint outlines of the rays and the octahedral nodes are discernible.
. Camerospongia aperta, Hinde. Natural size. From the Upper Chalk of the South
of England. The same specimen has been figured by Toul. Smith in Ann. &
Mag. Nat. Hist. 1848, vol. i. t. 14. f. 18
cloaca.
. The same. A portion of the cloacal surface, enlarged thirty times, showing the
spicular structure. The spicules have been completely dissolved, and only their
empty casts in chalk remain. The octahedral character of the nodes is only
shown in a few places.
. Camerospongia subrotunda, Mant. sp. A small group of individuals, natural size,
from the Upper Chalk at Charing, Kent. The cloacal cavities are filled with
the chalky matrix .
summit, from the Upper Chalk of the South of England. Natural size .
The same. A fragment of the outer surface of the same specimen, enlarged thirty
times, showing the characters of the dermal layer. The spicules have been
dissolved, and are now shown by red markings of iron peroxide.
The same. Showing the structure of the interior of the wall. Enlarged thirty
times. The minute tubercles produced by the filled-in moulds of the octahedral
nodes are only partially preserved.
. Porochonia simplex, T. Smith, sp. A specimen from the Upper Chalk, which
retains the cast of the smooth exterior membrane. Natural size
The same. A cast of the dermal layer of another specimen from the Upper Chalk.
Enlarged thirty times. The apertures in the dermal layer have been infilled
with the chalky matrix.
_ The same. Showing the spicular structure of the interior of the wall of 5a.
Enlarged thirty times. The spicules are only indicated by empty moulds in
the chalk.
Page
. 142
. The same. A median section of the same specimen, showing fie fumed” ahaped
: fe erect ooh oe fe.
. Callodictyon angustatum, Fides n specimen, somewhat worn away near the
. 142
aa
adhe
SPONGE
IOS Siw,
anvp.
i
West,Newman & C°
M.Suft &G.M.Herschell del et lith .
a
ae a a
Fig.
Fig.
Fig.
ees Ea
lid:
i)
“I
. The same. Showing the Saniprelaed fies of fig g.2, ah the large lateral anetiees
. Hyalostelia fusiformis, Hinde. Free hexactinellid spicules with ater centres.
PLATE XXXII.
. Plectoderma scitulum, Hinde. A fragmentary specimen, natural size, from Silurian
strata of Upper Ludlow age, in the Pentland Hills, near Edinburgh. For the
most part only empty moulds of the spicules are preserved, but here and there
fragments of siliceous spicules yet remain. The specimen figured belongs to
the collection of the Geological Survey of Scotland. It was discovered by
Mr. James Bennie .... . ok Gh eae:
The same. A portion of fig. 1, enlarged five times, showite the arrangement of
the spicules. Owing to the uneven surface of the matrix the vertical axes of
the spicules appear as if frequently broken off.
The same. A single spicule, enlarged five times, showing the complete vertical
axis and one complete ray, and the commencement of the other ray of the
horizontal axis.
Diplodictyon Bayfieldi, Hinde. Natural size. From the Upper Chalk at Norwich.
The specimen is preserved in chalk, and only casts of the spicular structure are
preserved .
Sclerokalia Cunningtoni, Hinde. An imperfect specimen, natural size, showing
the interior, and sections of the walls. From the Upper Green Sand at War-
minster, Wiltshire. The specimen belongs to the Jermyn-Street Museum
thirty times.
. The same. Part of the spicular structure of the outer surface of the cup. Enlarged
thirty times.
. The same. A portion of the spicular mesh of the interior, immediately beneath
the dermal layer, showing its regular arrangement and the octahedral character
of the nodes. Enlarged thirty times.
. Celoptychium agaricoides, Goldf. Showing the upper surface of the body of a
small imperfect specimen from the Upper Chalk of the South of England. The
spicular structure is now only indicated by red markings of iron peroxide in the
chalky matrix. Natural size
of the walls in the interior of the sponge.
. Astreospongia patina, Remer. Detached spicules, enlarged twenty-eight times,
from decayed Silurian shale of Wenlock age, near Visby, in the Island of
Gotland. The spicules have been changed to calcite. A small quantity of the
shaly matrix is usually attached to the central portion of the spicule
Natural size. Traces of free hexactinellid spicules, now in the condition of iron
peroxide, are shown on the outer surface
From the Upper Chalk at Horstead, Norfolk. Enlarged twenty-eight times
Page
132
. 145
. 146
. The same. A portion of the dermal layer of the interior of the cup. Enlarged
Soak : ee See . 146
. The same. A vertical section of fig. 4, natural size, sWoWite in section the folds
. 149
. Stauractinella cretacea, Hinde. An imperfect specimen from the Upper Chalk.
. 149
. 151
HOS SLE ON GES.
Plate XXX1
M. Suft & C.M.Herschell del. et lith West Newm
. & CP amp
Figs.
Figs.
Figs.
Figs.
PLATE XXXII.
“Page
1-lg. Hyalostelia Smithii, Young and Young, sp. Detached spicules of this species.
Enlarged fourteen times. Figs. 1 to 1d are simple hexactinellid forms; in 1
and 1c one of the spicular rays is reduced to a rounded knob; figs. 1 e, 1 f, and
lg are terminal portions of the spicular rods with four recurved rays. From
Lower Carboniferous Limestone at Cunningham Baidland, near Dalry, Ayrshire
s. 2-2 f. Holasterella conferta, Carter. Detached spicules of the species. Enlarged
fourteen times. Figs. 2 to 2¢ are the simpler forms; fig. 2d is an imperfect
stellate spicule; 2e is the upper surface of one of the petal-like spicules, and
2 fis the under surface of a similar but larger spicule. These various forms
have been found in combination in fragments of the sponge. From Lower
Carboniferous Limestone, Law Quarry, Dalry, Ayrshire aoe ic
3-3 d. Holasterella Youngi, Hinde. Detached umbrella-shaped spicules. Enlarged
fourteen times. Fig. 8 shows the upper surface of a spicule which is smooth;
in 3a, 36 the vertical rays of the spicule are seen, and the upper surface is tuber-
culated; 3¢ shows the under surface and the vertical ray of a small spicule;
and 3d is the upper surface of a large spicule in which the rays are nearly in a
horizontal plane. From Lower Carboniferous Limestone at Law Quarry, near
Dalry, Ayrshire ee a ee eee wey ts a
4-49. Holasterella Wrightii, Cart ter. Detached spicules of the species. Enlarged
twenty-eight times. Figs. 4, 4a, 46 are imperfect hexactinellid spicules with
simple rays; 4c, 4d are fragments of hexactinellid spicules with bifurcated
rays; 4e, 4f are hexactinellid spicules in which the rays are subdivided so as to
give the appearance of stellates; and 4g is the terminal portion of a spicular
rod with four recurved rays, which may have formed part of the root-appendages
of the sponge. From Lower Carboniferous Chert, Ben Bulbul, Sligo, Ireland .
5-5 e. Holasterella Benniei, Hinde. Detached spicules of the species. Enlarged
fourteen times. Figs. 5 to 5e¢ are hexactinellid forms with the rays variously
subdivided; 5e is a five-rayed form with the rays simple; and 5 d is also five-
rayed, and the rays bifureate near their extremities. From Lower Carboniferous
Limestone, Law Quarry, Dalry, Ayrshire
ig. 6. Peronella inflata, Hinde. Natural size. From the Middle Furs Goueneh a Cake
piers: Ranville, near Caen. . . .
e. 6a. The same. Portion of the fibre in the aan cat of fie stem, ae yea seventy-
two times, showing traces of three-rayed spicules. Drawn from a transparent
microscopic section by means of the camera lucida.
ig. 60. The same. Showing the spicular structure in the interior portion of the stem.
Enlarged seventy-two times.
150
. 152
FOSSIL SPONGES. Plate XXXIL.
West.Newman & Cf nnp
M_Suft & G.M Herschell del et bith
PLATE XXXIII.
. Peronella pistilliformis, Lamx. sp. Natural size. From the Great Oolite at
. 165
. The same. Portion of the outer surface of the same specimen, enlarged fifteen
Hampton Cliff, near Bath . . . . .
. . . . . . . .
times, showing the irregular apertures between the fibres.
Peronella tenuis, Hinde. Natural size. From the Pea-grit beds of the Inferior
Oolite near Cheltenham .
the same locality, enlarged seventy-two times, showing the disposition of the
spicules of the fibre in the interior of the sponge-wall.
. The same. Showing the disposition of the fibres near the margin of the sponge-
wall, with an entire spicule and fragments of others. The fibre is largely
crystalline, and only the larger spicules have been preserved.
. Peronella mamillifera, Lamx. sp. Natural size. From the Middle Jura, Couche
. 166
. Peronella cylindrica, Goldf.sp. A small specimen from the Upper Jura at Randen,
3 ley
. Peronella ramosa, Roem. sp. Natural size. From the Lower Green Sand at
. 169
a Polypiers, Ranville, near Caen .
Switzerland. Natural size.
Hemant, lean), 5 8 4 96 0 om 6 a oO
Page
. 166
. The same. Portion of a transparent microscopic section of another specimen from
Fig. 6. Peronella clavarioides, Lamx. sp. Portion of a transparent section of the fibre,
enlarged seventy-two times, showing three- and four-rayed spicules. From the
Middle Jura, Couche & Polypiers at Ranville, near Caen . 166
Fig. 6a. The same. Another portion of the fibre, from the same slide as fig. 6, showing
traces of spicules.
Fig. 7. Peronella furcata, Goldf. sp. Natural size. From the Cenomanian or Upper
Green Sand at Essen, Rhenish Prussia . 2 170
Fig. 8. Peronella prolifera, Hinde. Natural size. From the Lower Green Sand at
Farringdon, Berkshire . 169
Fig. 8a. The same. Portion of a transparent microscopic section of another specimen from
Farringdon, enlarged seventy-two times, showing the relatively wide, radiately
crystalline fibre, with traces of small three- and four-rayed spicules. The greater
portion of the spicular structure in this specimen has been obliterated.
Figs. 9-9 6. Peronella ocellata, Hinde. Three examples of this species, natural size, from
the Upper Chalk at Maestricht . . 171
Fig. 9c. The same. USS
. 200
. 201
HO ows, | oLONGES . Plate XXXVI.
dl
ay
De
oY. «
ve
2
Ns
t
Weet Newman & C° imp
M. Suft& GM Herschell del.etlith. ks ¥ c :
Fig.
Figs.
Fig.
Fig.
Figs.
PLATE XXXVIII.
a0
1. Pharetrospongia Strahani, Sollas. A fragment of the interior fibre, enlarged a
seventy-two times, showing traces of the spicular structure. Drawn from a
transparent microscopic section of an entirely calcareous specimen. From the
Cambridge Green Sand . . 201
2, 2a, 2b. Protosycon punctatum, Goldf. sp. ‘Two specimens of average dimensions—
fig. 2 is enlarged twice; 26 shows the cloacal aperture at the summit. From
the Upper Jura at Streitberg . 204
2c. The same. A vertical median section of a specimen, natural size, showing the
thickness of the wall, and the cloacal tube filled with the matrix.
2d. The same. A portion of the wall of 2c, enlarged twenty-eight times, showing
sections of the spicules bounding the horizontal canals. Drawn from a trans-
parent microscopic section.
2e. The same.