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CATALOGUE

OF THE

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IIAN DECAPOD CRUSTACEA %

IN THE

COLLECTION

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INDIAN MUSEUM.

PART I. BRACHYURA.

FASCICULUS I. INTRODUCTION AND DROMIDES OR DROMIACEA

(BRACHYURA PRIMIGENIA).

BY

A. ALCOCK, M.B.. LL.D., F.R.S.

■ • 1

INDIAN MEDICAL SEKVICE, COKRESPONDING MEMBER OK THE ZOOLOGICAL SOCIETY, FELLOW OF THE GEOLOGICAL SOCIEXr.
SUFEIUNTENDENT OP THE INDIAN MUSEUM AND PROFESSOR OP ZOOLOGY AND COMPARATITE ANATOMY
IN THE MEDICAL COLLEGE, CALCUTTA. SOMETIME SUKOEON-NATURALIST
\ TO THE INDIAN MARINE SURVEIT.

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1901.

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CATALOGUE

OF TlfK

INDIAN DECAPOD CliUSTACEA

IX THE

COLLECTION

OF THE

INDIAN MUSEUM.

PART I. BRACHYURA.

FASCICULUS I. INTRODUCTION AND DROMIDES OH DROMIACEA

(BRACHYURA FRIMIGBNIA).

BY

A. ALCOCK, M.B.. LL.D., F.R.S.

INDIAN MKIilCAL SRKVICE, COIiRESPONDING MEMBER OF THE ZOOLOGICAI, SOCIETY, FELLOW dV THE i; tUI.DIilc'Al, SOrlECV.

SUPICRINTENDENT OF THE INDIAN MUSEDM AND PUOFESSOR OF ZOOLOGY AND COMPAnATIVE ANATOMY

IN THE MEDICAL COLLEGE, CALCL'ITA. SOMETIME SUItGEON-NA M' 1,'A 1 I ST

rO THE INlilAX -MAI.'INE smVKV.

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CALCUTTA:

PKIXIED BV URUKR UF TIIK TKUSrEKS OF I'llK INDIAN MUSKU.M.

1901.
Price Seven Rupees.

CONTENTS OP PART I FASCICULUS T.

Index of Authors Quoted ...

InTRODDCTION : —

Morpliology of tlie Deoapoda
Classification of the Decapoda ...

Brachyora: —

General Considerations

Classification of tlie Bracliyura ...
Dromides or Dromiacea ...

Droroiidea

Homolidea
Table of the Genera and Species of Recent Dromiacea

pp. v-ix.

pp. -2-12.
jip. l:{— 1.5.

]i|i. 1(3—25.
pp. 26—28.
pp. 28—29.
pp. 29—58.
pp. 58—73.
pp. 74—80.

Phefack.

The Catalogue of [udian Decapod Crustacea, of which this is the first instal-
ment, occnpied in its early preparatory stages inucli of the attention of my
predecessor Mr. James Wood-Mason, and has in execution taken up the greater
jiart of my own time, particularly in leisure honrs, during the last eight years.

At the outset T looked foi'ward to signalizing on the title-page my late frientl's
connexion with this undertaking; but 1 now Hnd it necessary to take the entire
responsibility of the work as it stands, and to state that the idea of codifying oin-
knowledge of Indian carcinology originated many years ago Avith Wood-Mason,
and that by collecting and commencing to index all the classical literature of the
subject, by sorting into great-groups that part of the Museum collection of
Ci'ustacea tliat was made independently of the Survey Ship " Investigator," by
gradually ac(|uiring a considerable nnnil)er of authentically-named species of
Crustacea, and by provisionally naming a good uiany Tndo-Pacific species himself,

Wood-Mason had gone no inconsiderable way towards eariying his idea into
effect.

It is pi-0]»osed to issue the ('ataloguc in three collateral and iiulependent
series, one for the llraehijiira, anotluM' for the Marriiru, :ind a third for the
AitoTihiira.

— I

iisr

THE INDIAN MUSEUM

A. Ai.cocK, Major, I.M.S.,

Sn'iierinh'ndcnt of tlir Indian Mhschih.

INDEX OF THE WORKS QUOTED IN THIS VOLUME.

1. AnAMS, A. and White, A. — Zoology of the Voyage of H.M.S, Samarang, Crustacea.
London, 1850.

tl. Alcock, a. — Natural History Notes from the Royal Indian Marine Survey Ship
" Inve.stigator." Series II, No. 1. Annals and Magazine of Natural
History, May, 1894.

3. ,, ,, Materials for a Carcinological Fauna of India. No. .5. The Brachyura

Priniigenia, or Dromiacea. Jo%irnul of the Asiatic Society of Bengal,
Vol. LXVIII, Pt. 2, 1899.

4. ,, ,, An Account of the Deep Sea Brachyura collected by the Royal Indian

Marine Survey Ship " Investigator." Calcutta, 1899.

5. Alcock, A. and

Anderson, A. R, S. — Natural History Notes from the Royal Indian Marine Survey Ship
" Investigator." Series II, No. ll. Jotirnal of the Asiatic Society of
Bengal, Vol. LXIII, Pt. 2, 1891.

6. „ ,, Do. Do. Series III, No. 2. Annals and Magazine of Natural History,

January, 1899.

7. Bate, C. Spence. — Report on the Crustacea Macrura collected by H.JI.S. Challenger.

London, 1888.

8. Bei.l, T. — A History of the British Stalk-eyed Crustacea. London, 1853.

9. BoA.s, J. E. V. — Studier over Decapodernes Slaegtskabsforhold, avec ua resume eu

francais. Danske Videnskab. Selskab Skrivter, 6 Eiekke, luiturviden-
skahelig og mathematisk Ajd. I, 2, 1880.

10. BonuAPAll.E, L. A. — Crustaceans from the South Pacific. Part IV, Crabs. Proceedings

of the Zoological Society of Loudon, for 1900.

11. Bosc, L. A. G. — Histoire Naturelle des Crustaces. Paris, 1803.

12. BouviEi;, E, L. — Sur rorigine Homarienne des Crabes. Bulletin de la Societe Pliiloma-

thique de Paris: ser. 8, torn. VIII, 1895-yG.

13. ,, ,, Crustaces Anomoures et Brachyures recuellis en Basse Califoruie.

Bulletin die Miiseurr,. d'Hlstoire Naturelle. Paris, 1898.
1-1. Buoccul— Sur les organes genitaux males des Crustaces Decapodes. Annates des Sciences

Naturelles, ser. 6, torn. II, 1875. Zoologie.
15. Cano, G. — Viaggio della R. Corvetta Vettor Pisaui attorno al Globo. Crostacei

Brachiuri ed Anomouri. Boll. Soc. Nat. Napoli, III, 1889.

hj. Cakbington, J .T. and

LovETT, E. — Notes ou British Stalk-eyed Crustacea. The Zoologist : ser, :;, Vol. VI,
1882.
17. Cakcs, J. V. — Prodronuis Fauna; Mediterranean. Vol. I. Stuttgart, 1885,

VI

18. Clads, C. — Untersucliuniien zur Erforscliuiifj tier gpnealogisclicn Grundlage des Crus-

taceensystems. Vienna, 1876.

19. „ Grundziige der Zoologie, 4tli Edit. Waibuig, 1880.

20. Costa, 0. G.— Fauna del Regno di Napoli— Naples, 1836-1870. Crostacei.

21. Dana, J. D. — United States Exploring Expedition. Vol. XIII, Crustacea, pait I.

Philadelphia, 1852.

22. De Br^BISSON. — Catalogue des Crustaces du Calvados. Memoires de la Societe Liuneenne

du Calvados. 1825, Caen et Paris.

23. De Haan, W. — Fauna Japonica, auctoie P. F. De Siebold, Crustacea. Leyden, 183.5-

185C.

24. De Man, J. G. — Ueber die im indischeu Archipel gesammelten Decapoden. A-rchic fiir

Naturgeschichte, Jahrgang LIII, Bd. i. 1887. Berlin.

23. ■„ Report on the Podoplitlialmous Crustacea of the Mer^ui Archipelago.

Journal of the Linneaii Society, Zoology. Vol. XXII, 1887-1 888.

26. ,, Ueber neue order seltene iudopacifiselie Brachyuveii. Zoologische

JaJirbiicher. Ahtheiluny fiir Systematik, etc. Bd. IV, 1889.

27. „ Decapoden uud Stomapoden von Malakka, Borneo, und Celebes.

Zooloyische Jahrbiccher. Ahtheilmuj fiir Systematik, etc. Bd. IX, 1897.

28. Desmarest, a. G. — Article Malacostraca in Dictionnaire des Sciences Naturelles. Tom.

XXVIII, 1823. Paris.

29. ,, Considerations Generales sur la Classe des Crustaces. Paris, 182.5.

30. Eydoux et Sodleyet. — Voyage autour du Monde sur la corvette La Bonite. Zoologie,

torn. I. Paris, 1841.

31. Pabricius, J. C. — Entomologia Systematica (et Supplementum). Copenhagen, 1792-

1798.

32. Gerstaecker, A. — Bronn's Klassen und Ordnungen des Thier-Reichs. Crustacea

Malacostraca.

33. Gray, J. E. — The Zoological Miscellany. London, 1831.

34. GuiRiN-MtNEViLLE, F. E. — Iconographie du Regue Animal du Cuvier. Paris, 1829-

1844. Crustaces.

S5. ,, Description du genre Hypoconcha. Jieviie et Magasin du Zoologie : ser. 2,

tom. VI. 18.54. Paris.

~i6. GuiLDIKG, L. — An Account of some rare West-Indian Crustacea. Transactions of the
Linnean Society of London. Vol. XIV, 1825.

37. Hasweli., W. A. — Description of some New Specie.s of Australian Decapoda. Proceed-
ings of the Linnean Society of New South Wales. Vol. VI, 1882.

;i8. „ Catalogue of the Australian Stalk- and Sessile-eyed Crustacea.

Sydney, 1882.

39. Hei.leb, C. — Crustaceen-Fauna des rothen Meeres. SitzmigshericMe der Mathematisch.

Naturivissenschafllichen Classe der K. Akademie, Wien. Bd. XLIV. Abth.
1, 1862.

40. „ Die Crust aceen des siidlichen Europa. Vienna, 1863.

41. „ Reise der Oesterreichi.schen Frigate Novara uni die Erde. Crustaceen

Viennii, 1865.

Vll

42. Hendeusos, J. R. — Report on tlie Aiioniiira collected by H.M.S. Challenger. Loudon-

1888.

43. ,, A Contribution to Indian Carcinolog-y. Transactions of the Liiincau

Society of London. Zoology, ser 2, Vol. V, 1893.

4^. Hehbst, J. F. W. — Ver.such einer Naturgeschiclite der Fvtjibben und Krebse. Berlin,
1790-1804.

45. HiLGENDORF, F. — Mozambique Crustaceen. Moudtsherichte der K. Preiissischen Akademie

der Wissenschaften zu Berlin, 1879.

46. HnxLEY, T. H. — On tbe Clas.sification and the Distribution of the Crayfishes. Proceed-

ings of the Zoological Society of London, 1 878.

47. „ The Crayfish. London, 1880.

48. Ilh:str.\tions of the Zoology of the Rotal Iniuan ^Iarine Sdrvey Ship " Investi-

gator." Calcutta, 1892-1900.

49. Ives, J. E. — Echiuoderms and Arthropods from Japan. Proceedings of the Academy of

Natural Sciences of Philadelphia. 1891 (1892).

.50. KiNGSLEY, J. S. — On Crustacea from Virginia, North Carolina, and Florida. Proceed-
ings of the Academy of Natural Sciences of Philadelphia, 1879 ( 1880).

.jl. KossMANN, R. — Zoologisehe Ergebnisse einer Reise in die Kiistangebiete des rotheu
jNIeeres. Malacostraca. Leipzig, 1877.

52. „ Kurze Notizen iiber ncue Crustacccn. Archie fiir Natiirgeschichte, -Jahr-

gang, XLIV. Bd. i., 1878,

53. Kradss, F. — Die siidafi-ikaniscUen Cr-ustaceen. Stuttgart, 1843.

54. Lamarck, J. B. P. A. — Histoire Xaturelle des Animaux sans Vertebres. Paris, 1815-

1822.

55. Lati;eille, P. A. — Histoire Xnturelle des Crustace.s et des Insectes. Vols. V-VI.

Paris, 1803.

66. ,, Geneiu Crustacfonini et Insectorum. Paris, 1806.

57. ,, Articles in Nouveau Dictionnaiie d'Histoire Naturelle. Tom. IX.

Paris, 1817.
68. Lea.ii, W. E.— The Zoological Miscellany. London, 1814-1817.

59. ,, A Tabular view of the characters of the Insecta, etc. Trancactiins of

the Linnean Society. Vol. XI, 1S15.

60. ,, Malacostraca Podophthalmia Britannia". Loudon, 1815.

61. LiNNfi, C. — Systema Naturas, torn. 1. pars. ii. Editio Duodecima Reformata. Stock-

holm, 1767.

62. Lucas, H. — Exploration Scientiiique de TAlgerie. Histoiro Naturello dc3 Auimaus Arti-

cules. Part I. Paris, 1849.

63. „ Histoire Naturelle des Crustaces, des Arachnldes, et des Myriapodcs.

Paris, 18.J0.

64. Maci.eay, W. S. — Illustratious of the Aunulosa of South Africa. London, 1838.

65. MiERS, E. J. — Catalogue of the Stalk- and Sessile-eyed Crustacea of Ne\T Zealand.

London, 1876.

66. „ On Crustacea from the Corean and .lapauesp. Seas. Proceedings of the

Zoological Society of London, 1879.

vni

f>~. MiERS, E. J. — On Crustacea from tlie Malaysian Region. Ammh and Magazine of Natural
History, ser. 5, Vol. V, 1880.

68. „ Crustacea from Goi'ee Island, Senegarabia. Annals and Magazine of Natural

History, ser. 5. Vol. VIII, 1881.

69. „ Ou Some Crustaceans from Mauritius. Proceedings of the Zoological Society

of London, ]884.

70. „ Eeport on the Zoological Collections made in the Indo-Pacific Ocean during

the voyage of H.M.S. Alert. Crustacea. London, 1884.

71. „ Report on the Bracbyura collected by H.M.S. Challenger. London, 1886.

72. Milne Edwards, A. — Crustacea nouveaux provenant des Voyages a Zanzibar et a Mada-

gascar. Nouvelles Archives da Museum d'Histoire Naturelle de Paris.
Tom. IV, 1868.

78. „ Memoire siir les Crustaces Decapodes du genre Dynomeue. Annales des

Sciences Naturelles. Zoologie. Ser. 6, Tom. VIII, 1879.

74. „ Reports on tlie Results of Dredging by the United States Coast Survey

Steamer Blake. Bulletin of the M-iiseum of Comparative Zoology at
Harvard College in Cambridge. Vol. VIII, ISSO-Sl.

75. „ Recueil de Figures de Crustaces nouveaux ou peu counus. Paris, 1883.

70. Milne Edwaei-s, A.

et BiuviEi;, E. L. — Crustaces Decapodes provenant des campagnes du yacht I'Hirou-
delle. Part. I. Brachyures et Anomoures. Monaco, 1894.

77. ,, Cru.staces Decapodes piovenaut des campagnes de I'Hirondelle (Sup-

pU'inent) et de la Princesse Alice. Monaco, 1899.

78. „ Expeditions Scientifiques du Travailleur et du Talisman. Crustaces

Decapodes: part I. Brachyures et Anomoures. Paris, 1900.

79. Milne Edwaiids, H. — Histoire Naturelle des Crustaces. Paris, 1834-1840.
SO. „ La Regne Animal par Georges Cuvier. Les Crustaces. Paris.

81. MiJLLER, F. — Zur Crustuceenfauna von Trincomali. Verhandlungen der Naturforschenden

Gesellesrhaff in TiaseL Till, 18S6.

82. Ortmann, a. — Die Decapodeiikreb.se des Strassburger Museums. ZoologischeJakrbucher.

Ahtheilnng fiir Systematik, etc. Bd. VI, 1892.

83. „ Crustnceen in Semon Zoologische Forschungs-reisen in Australien und

dem Malayischen Ai-chipel. Jenaische Denhschriften. VIII, 1895.

84. ,, Malacostraca, in Bronn's Klassen und Ordnungen des Thier-Reichs.

8.5. Peters, W. — Ueber das System der Dekapodeu. Monntsherichte der K. Preussischeu
Ahademie der Wissenschaften zu Berlin, 1861 (1862), p. 717.

86. Rathbun, Makt J. — Decapod Crustacea of Jamaica. Annals of the Institute of Jamaica.

Vol. I, 1897.

87. ,, Decapod Crustaceans of West Africa. Proceedings of the United

States National Museum.. Vol. XXII, 1900,

88. RiCHTEKs, F.— Meeresfauna der Insel Mauritius und der Seychellen. Decapoda.

Berlin, 1880.

89. Risso, A. — Histoire Naturelle des Crustacea des environs de Nice. — Paris, 1816.

90. ,, Histoire Naturelle des Principales Productions do I'Europo Meridionale, etc.

Paris, 1826.

1*1 Riiix. P.— Crustaces de la Mediterraneo et de son littoral. Paris, 1828.

02. RcMPHiUiS, G. E. — D'Amboinsche Rariteitkamer, eti3. Amsterdam, 1705.

93. RuPPELL, E. — Beschreibung und Abbildung von 2i Arten kurzschwiinzigen Krabbeii
des rothen Meeres. Frankfurt, a. M., 1830.

'J4t. Sep.a, a. — Thesaurus Rerum Naturalium. Amstei'dam, i7i>8.

9-5. Smith, S. J. — Preliminary Report on the BracLyura and Anomura dredged in deep
water off the south coast of Now England. Proceedings nf the United
States National Museum for 1883, Vol. VI.

96. „ Reports on the Decapod Crustacea of the Albatross Uredgings. Aninutl

B'eports of the United States Commissioner of Fish and Fisheries fur 1882
(188-1) and 1885 (1886).

97. STE:;i;i.\ti, T. R. R. — A History of Crustacea. Recent Malacostraca. London, 1893.

9g. „ ., ilarine Investigations in South Afiica, Crustacea. Cape Town, 1900.

99. SiiMPtioN, W. — Prodronious descriptionis Animaliuni evertebratorunr, etc. rruceedimjs
of the Academij of Natural Scienas of Philadelphia, 1858 (1859).

100. „ Notes on North American Crustacea in the Museum of the Smithsonian

Institution. Annals of the Lijceuni of Natural History of New . York,
Vol. VII, 1862.

101. Studeu, T. — Crustaceen der Gazelle von Westafrika. Ahhandlungen der Koniglichen

Akademie der Wissenscluijtvu :u Berlin, 1882 (1883).

102. Thomson, G. M. — A Revision of the Crustacea Anomura of New Zealand. Transactions

and Proceedings of the Neu- Zealand Institute, 1898 (1899).

103. To/./.ETTi, A. T. — Zoologia del Viaggio Intorno al Globo della R. Pirocorvetta ilagenta.

Crostacei Brachiuri 0 Anomouri. Florence, 1877.

104. Veurili,, a. E.— On the parasitic habits of Crustacea, llie American Naturalist, Vol. Ill,

iy70.
lO.j. W.4LKEI!, A. 0. — On Crustacea from Singapore. Journal of the Linncan Sociclg, Zoologg.

Vol. XX, 1886-1900.
106. "White, A. — List of the specimens of British Animals in the collection of the British

Museum. Part IV, Crustacea. London, 1850.
1U7. WnnELEflGE, T. — The Crustacea of Funafuti. Memoir III. Australian Museum, Sydney,

1897.

108. „ Scientific Results of the Trawling expedition of H.M.C.S. Thetis,

Crustacea. Memoir IV. Australian Mttse^^m, S'/dnei/, 1900.

109. WooD-M-\soN, J. — Natural History Notes from the Royal Indian ilarine Survey Ship

■' Investio-ator." No. 21. Anmils and Magazine of Natural History.

March, 1891.

110. ZiETz, A. — New species of Austialian Crustaceans. Transactions aiul Proceedings of the

Foyal Society if South Australia, Vol, X, 18S6-I887.

IXTRODUCTION.

This volume is the First Part of a monograph of the Decapod Crustacea of
that part of the Oriental Eegion of Wallace which Kes within the political
boundaries of British India.

The Decapoda belong to the subclass Malacosfraca of the Crustacean kind,
the Crustacea being one of the main branches of the great phylum Arthropoda.

In the members of this phylum, which also includes the Xipliosura, the
Arachiida, the Millipedes, the Centipedes, and the Insects, the body consists of a
series of segments or somites carrying jointed or segmented appendages. Though
the segments — with the pair of appendages which each one typically carries
— may vary greatly in actual form, yet they are all built on one and the same
fundamental plan, and the superficial layers of the integument, both of the
somites and of the appendages, are more or less completely hardened, by chitin
or by salts of lime, so as to form a rigid exoskeleton, which during growth is
periodically moulted.

Among the Arthropoda the Crustacea are recognized by the fact that while
they possess antennae, as do the air-breathing Insects, Centipedes and MilUpedes,
they typically breathe in water by means of gills connected with the appendages,
as do the non-antennate king-crabs {Xiphosura) . Their spermatozoa are non-
motile, like those of Millipedes.

Among the Crustacea the Malacostraca are distinguished by the fact that
the body is made up of a definite and constant number of segments.

The characters that separate the Order Becapoda from the other ordei's of
Malacostraca will be understood if we examine, in some detail, a typical member
of the Order.

The type usually selected is the European Crayfish, Astacus fluviatilis : the
type here chosen is the Andaman Lobster, Nephrops andamanica, as belonging to
the local fauna.

As far as tbe general construction of the body, the general modifications of its appendages,
and the general relations of the viscera go, any of the common species of freshwater or murine
prawns (Palsemon, Feneus) may be used for dissection if Nejihrops is not procurable.

I. MOEPHOLOGT OF NePHEOPS ANDAMANICA.

1. Of the general construction and segmentation of the Body.

In a typical Decapod Crustacean, such as Nephrops andamanica, the body
is elongate and is manifestly divided into two regions, an anterior region or
cephalothorax and a posterior region or abdomen, of which the latter is the longer.

The cephalothorax is covered, as to its back and sides, by a shield or
carapace, so that its segmentation — or subdivision into somites — is not immediately
apparent ; but the abdomen obviously consists of a series of seven segments, and
therefore bespeaks our attention first.

Any one of the seven freely-movable abdominal somites, except the last, is
seen to be a ring, with a pair of appendages attached to its under surface. The
dorsal arc of the ring is wide fore-and-aft and is known as the tergmn, the
ventral arc is narrow fore-and-aft and is known as the sternum, while the over-
lapping sides of the ring are called the plexira : furthermore, the narrow corner
between the articulation of the appendage and the pleurum of either side is
termed the epimerum.

On turning to the cephalothorax, the most that we can distinguish, before
the removal of the carapace, are the narrow sterna of some of the thoracic
somites ; and even when we have removed the carapace, and have cleared away
the gills that lie beneath it, we shall at first sight find, instead of the succession
of rings that the abdomen shows, merely the greatly-extended epimera of some
of the thoracic somites.

The cephalothorax is, however, as an examination of its sterna and append-
ages reveals, made up of 14 segments, all of which are immovably fused together,
many of which are contracted and foreshortened and obscured, and none of which
— with the exceptions presently to be noticed — possess identifiable terga like those
of the abdomen. The place of terga and pleura is taken by the carapace, the
main dorsal mass of which is generally supposed — though it is not certain that
this explanation covers all the facts — to be an enormous extension backwards of
those elements of the 3rd (antennal) and 4th (mandibular) somites of the body.

The carapace, it is to be observed, does not merely serve as a cover to the
united segments of the cephalothorax, but it overlaps those segments so as to
enclose on either side of them a space in which the gill-plumes he, this space
being known as the branchial chamber. Moreovei-, it is produced forwards in the
middle line, to form a stout bayonet-like organ of defence — the rostrum.

Although the cephalothorax is one composite whole, it consists of two ill-
defined regions, an anterior cephalic region or head and a posterior region or
thorax. The boundary line between the two regions is marked on the carapace
by a deep semicircular groove — the cervical groove ; and when the carapace is
removed, the division between the two regions is still obscurely indicated by a

fold or a pair of lateral hollows in which a pair of organs known as scapliorinatliites
(to be described along with the appendages) lie.

We shall find, on examining the appendages, that the head consists of six
segments fused together, and the thorax of eight. Add to these the seven
abdominal segments, and we get twenty-one as the number of segments that
make up the body of Nephrons.

It has been questioned wlietlier the last segment of the abdomen, known as the telson, is really
a somite, since among other reasons it carries no appendages. But it seems probable, from the
form and sculpture of this part of the body in certain deep-sea Decapods— notably the J? rf/o)U(f«
and some GalatheiJse — that the telson consists of a modified somite and a pair of biramous appeud-
iiges all fused together. Moreover it is traversed by the intestine.

It has also been objected that the eyes are not truly homulogous with appendages and that
the part from which they spring is not therefore a somite. But apart from other considei-ations
tlie fact that, as a "substitution" abnormality, the' eyestalk may be replaced by a recognizable
appendage, is enough to dispose of this objection.

On the other hand it has been argued that the pair of fleshy lobes that form the posterior
boundary of the mouth-slit and are known as the metastoma or loiver lip, .are true appendages
and therefore represent an additional somite ; though there are no facts to support this view.

The cephalothorax is held together not merely by fusion of the sterna and

epimera of its constituent somites, but also by an internal lattice-work of thin
hard obliquely-transverse partitions known as apodemes.

Although the apodemes are internal and serve to protect and support the
viscera as well as to give attachment to muscles, they are merely infoldings of
the cuticle of the constituent sterna and epimera. They are only found behind
the mouth and in the ventral half of the somites.

Before further considering the segmentation of the cephalothorax it is
necessary to speak of the appendages.

2. Of t]te Appendages in general.

The recognizable appendages of Nephrops andamanica are twenty pairs.

The first six pairs belong to the head : three of them lie in front of the
mouth and are organs of the senses, while three are crowded together on either
side of the mouth and are biting and chewing organs, the last pair also
contributing to the respiratory mechanism.

The next eight pairs belong to the thorax : the first three of them, which
are crowded together in close succession to the mouth-parts of the head, ari>
organs partly of manducation and partly auxiliary to respiration ; while the
other five are legs in the ordinary sense, though some of them are modified
for prehension, and most of them take a certain part in respiration. It is from
these 5 pairs of large legs that the name of the Order is derived.

The last six pairs belong to the abdomen and are primarily swimming-
paddles, some of which, in the male, are modified to subserve reproduction,
and in the female subserve nutrication or the care of the developing embryo.

Just as the abdomen furnishes the type of the unmodified Crustacean seg-
ment, so also it gives us the type of the Crustacean appendage.

An appendage taken from, say, the 3rd abdominal somite consists of a
stalk, or protopodite, to the end of which two leaf -like branches are articulated,
the inner branch being known as the endopodite the outer as the exopodite. The
stalk itself consists of two segments, a short basal piece or coxopodite, by means
of which it is articulated to the somite, followed by a larger sub-cylindrical
hasipodite'.

Although a minority of the appendages of a Decapod Crustacean exactly con-
form to this description, yet they all can be referred to this general plan, or type.

3. Of the Segments and Appendages of the Head.

The first segment, or ophthalmic somite, is hidden by its appendages, the
eyes. If the eyes are pushed aside, we shall find, between and just below
them, a small plate of leathery consistence : this is the sternum of the ophthal-
mic somite, and is all that we can with safety recognize as belonging to that
somite. Its appendages, the eyestaJks, each consist of a simple two-jointed pro-
topodite, the large compound facetted eye terminating and forming the chief
part of the second joint.

The second segment, or antenmdar somite, is also a good deal hidden. All
that we can recognize as certainly belonging to it is its sternum, which is a
narrow partition between the antennules ; but the " orbits," or facets on either
side of the carapace on which the eyes rest, are very probably its epimera. Its
appendages are the antennules, each of which consists of a thickish stem or
peduncle (protopodite) to the end of which two long slender t&:^eYmg flagella
(exopodite and endopodite) are articulated. The protopodite is abnormal in
consisting of three segments : the proximal segment contains the avditori/ sac,
the small orifice of which is seen at the inner border of the upper surface of
the segment, in its posterior half. The flagella consist of a series of little rings
with fine setae : the outer, or olfactory flagellum, is the thicker of the two, and
its setae are larger and more thick-set, especially in its distal part.

The third segment is the antennal somite and its appendages are the antennse.
The broad space between the bases of the antennte, known as the epistome, is the
sternum of this somite, behind the outer ends of which the epimera are visible.
The pleura and the tergum of this somite probably form all that part of the
carapace that lies in front of the cervical groove.

Perhaps it would be better to say tbat, whatever other elements maj- enter into the formation
of that part of the carapace which lies in front of the cervical groove, the pleura and terga of the
third or antenuular somite can, in many Macrarous and Braohyurous Crustacea, be recownizod as
forming the preponderating element.

Each antenna consists of a peduncle, a scale (exopodite), and a lash or
fiagellum (endopodite). The peduncle is constituted of a rather zig-zag series of
five joints, the first two of which correspond with a protopodite : on the ventral
surface of the first joint is a prominent orifice, the renal tubercle, at which the
green gland, or excretory organ, opens. The antenna! scale, which is a broad
thin plate, is articulated to the end of the 2nd joint of the peduncle. The
fiagellum, which is slender and of great length, is made up of a succession of
little rings, and articulates with the end of the peduncle : it is an organ of touch.

The fourth segment is the mandibular somite and its appendages are the
viandihles : its modifications take the same turn as do those of the antennal somite.
Its sternum is recognizable as forming the posterior part of the epistome ; its
pleura and tergum probably constitute the preponderating elements of that part
of the carapace which lies behind the cervical groove.

It is difficult to avoid the inference that in the carapace of most Macrura and Bracltyura
the part whicli lies behind the cervical groove belongs to the mandibular somite. In most
Anoviura, however, and in some Macrura, the mandibular soniite seems to enter into the
formation of the sidewalls of the carapace only. Furthermore, the sutures of the carapace of
Thalasdna forcibly suggest that other terga and pleura, besides those of the antennal and mandi-
bular somites, may take a considerable share iu the formation of both the roof and the sidewalls
of the carapace.

Each mandible consists of (1) a strongly calcified protopodite placed obliquely,
and (2) a cm"ved three-jointed palp (eiidopodite). The body of the mandible,
or protopodite, is shaped something like a tooth : its distal end is broadened and
is grooved fore-and-aft, the ventral edge of the groove forming a salient and
trenchant incisor process, while the concealed dorsal edge of the groove forms a
broadened molar process.

The fifth and sixth segments are only visible after the anterior appendages
of the thorax ai'e removed, and even then all that is seen of them is a little plate
lying in the middle line, behind the mouth, and representing their conjoint sterna,
to which they are reduced. It is also possible that their pleura are represented
— though their individuality is, of course, completely lost — by a part of the edge
of the carapace in their own immediate neighbourhood. Both these somites are,
however, represented by appendages.

The appendages of the fifth somite are the \st maxillx! They are thin
foliaceous chewing-organs, and lie in contact with the mandibles. Each consists
of (1) a leaf-like coxopodite and basipoditc, both of which have their long axis
nearly transverse and their free (distal) end fringed with set«, and (2) a two-
jointed endopodite or palp. The exopodite is not present.

The appendages of the sixtli somite are the 2iid maxillse. All their parts
are thin and foliaceous. The coxopodite and basipodite resemble those of the
1st maxillge, except that each is deeply cleft into two lobes. There is a tapering
endopodite, and a leaf-like exopodite, which last, being used to bale out the
branchial chamber in the course of respiration, goes by the special name of
scaphognathitn.

4. Of the Segments and Appendages of the Thora,r.

The individuality of all eight thoracic somites is maintained by the sterna,
which can be seen without removal of the carapace, and by the epimera which
can only be seen after the carapace is removed. The individuality of their
pleura and terga is entirely lost, but it is possible that the slightly-raised lateral
and posterior borders of the carapace represent these elements. We need,
therefore, consider in detail only the appendages.

The fate of the terga and pleura of the thoracic somites has ahead}' been incidentally con-
sidered in dealing with the composition of the carapace.

I have adopted with slight modification the views of Milne Edwards I., that the carapace is
formed by the terga and pleura of the aptenno-mandibular somites, and that these elements of the
thoracic somites are practically unrepresented in the carapace.

It must be admitted however that the sculpturing and sutures of the carapace admit of other
equally reasonable interpretations. For instance, Huxley {The Crayjlah, p. 166) taught that that
part of the carapace which lies in front of the cervical groove represents the terga and pleura of
the head ; while of that part which lies behind the cervical groove, the median region represents
the terga, and the lateral regions represent the pleura, of the thoracic somites. The sutures
of the carapace of Thalassina certainly support this interpretation, and even in the carapace of
Nephrops there are grooves and spines which hiirmonize with it.

The first pair of thoracic appendages are the 1st maxillipeds. They consist
of a protopodite, endopodite and exopodite, with, in addition, a leaf -like appendix
having the same function as the scaphognathite and known as an epipodite. The
protopodite is formed of a leaf-like coxopodite and basipodite similar to, only
larger than, the same elements of the 1st maxillte : the endopodite is a thin two-
jointed leaf ; the exopodite, which is longer and slenderer than the endopodite,
ends in a truncated y?a(7e//«<Jrt ; the epipodite is attached to the outer end of the
coxopodite. Their function, like that of the next two pairs of appendages, is
partly manducatory, partly respiratory, and in structure they are intermediate
between maxillae and ordinary maxillipeds.

The second pair of thoracic appendages are the 2nd maxillipeds. They
are foot-jaws in which their function as organs of mastication has not disguised
their resemblance to the ordinary appendages of the thorax. They consist of a
protopodite, an endopodite, an exopodite, and an epipodite to the base of which
a rudimentary gill-plume or podobmnch is attached. The protopodite consists
of a short coxopodite and basipodite : the endopodite, which is larger and longer

than the exopodite, is made up of the following five successive pieces, (1) a short
ischiopodite, which is immovably united with the basipodite, (2) a long meropodite,
(3) a short carpopodite, (4) a short and broad piroi^odite, and (5) a semioval
dadylopodite. The slender exopodite resembles that of the 1st maxillipeds.

The third pair of thoracic appendages are the third or external maxillipeds :
they are foot-jaws in which the resemblance to the ordinary appendages of the
thorax is more decided than in the case of the other maxillipeds. Except that
their endopodite is longer and their exopodite shorter, and that their ischiopodite
is longer and their meropodite relatively shorter, and that their podobranch is a
functional gill-plume, they do not essentially differ from the 2nd maxilhpeds.
The inner edge of their ischiopodite is, however, furnished with an elegant row
of calcareous teeth.

The fourth to the eighth pairs of thoracic appendages, which are commonly
known as " legs," are modified for locomotion and, in part, for prehension, and
though they are not all exactly alike, have all an essential similarity of structure.
None of them have an exopodite, and in all of them the endopodite consists of
five subcylindrical segments, or joints— ischiopodite, meropodite, carpopodite,
propodite, and dactylopodite — which form, in succession, a continuous series with
the coxopodite and basipodite, all of these joints being capable of independent
motion, except the basipodite and ischiopodite which are firmly united and move
as a single piece. All of these appendages except the last have, articulated to
the dorsal angle of the coxopodite, an epipodite and a podobranch.

The fourth pair, which are known as the great chelipedfi, are much the
longest and vastly the most massive of the appendages of the thorax : their
propodite is of enormous size and has its distal angle produced far beyond the
articulation of the dactylopodite as a long process, which is a counterpart of
the dactylopodite itself and forms with the dactylopodite a pair of strong
pincers — the chelae.

The fifth and sixth pairs are slender and subcylindrical : they also end in
chelae, which are small.

The seventh and eighth pairs are simple legs for crawling, the seventh are
a little longer than, and the eighth are about the same length as, the fifth and
sixth pairs.

5. Of the Segments and Appendages of the Abdomen.

The typical abdominal segment, and its appendages, have already been
described. The segments are all freely movable : the tergum of the first seg-
ment is overlapped by the carapace and overlaps the tergum of the second, the
tergum of the second is overlapped by that of the first and overlaps that of
the third, and so on in succession.

The first abdominal somite differs from the others in the large extent of

8

its epimera and tlie small and rudimentary character of its pleura, in these
respects approacliing to the thoracic somites. Its appendages also differ from
those of the other abdominal somites in being uniramous. The first pair of
abdominal appendages are alike in both sexes, in so far as they consist of a pro-
topodite followed by a single branch, probably the endopodite ; but whereas in
the female they are slender, in the male they are stout and are deeply grooved
along their inner face so as to form, by their apposition, a canal. Their apposition
is rendered as intimate as possible by the interlocking of a little roughened
facet on the inner edge of each. The canal forms a passage for the sperm, and
these appendages in the male are commonly spoken of as copidatory organs.

The second abdominal somite is distinguished by the large size of its pleura,
but is in all respects typical. Its appendages in the female are quite typical ;
but in the male they are characterized by the presence of a little lobe on the
inner edge of the base of the endopodite, which little lobe can be securely
apposed to its fellow of the opposite side, so that both can play in the canal
formed by the apposition of the first pair of appendages, and they must there-
fore be regarded as accessory organs of copulation.

The third to the fifth abdominal somites are quite typical, as also are their
appendages, which are biramous setose swimming-paddles constructed on the
plan already described. In the female these appendages, as also those of the
2nd somite, are used at the breeding-season for carrying the fertilized ova and
the newly-hatched young.

The sixth abdominal somite has the pleura less well developed, and the
sternum more strongly developed, than those of the other abdominal somites.
Its appendages, which are known as the caudal sivhnmerets, are strongly calcified
and of large size, and they form with the last abdominal somite the great tail-
fan, which is the principal organ of aquatic locomotion. In these appendages the
protopodite is short and broad, the exopodite and endopodite are broadly foliaceous,
and the exopodite is transversely divided in its distal moiety into two segments.

The seventh, abdominal somite is the telson, which differs from all the other
somites in being a broad thin ^jlate without a^^pendages. On this account, and
also because it is destitute of a nerve-ganglion, it is sometimes regarded as
merely an outgrowth of the sixth somite. But the facts that the alimentary
canal perforates it, and that in many deep-sea species of Decapoda it can
apparently be resolved into a somite and a pair of appendages all fused together,
are held sufficient to justify its inclusion in the series of somites.

6. Of the Ees])iratory Organs.

The organs of respiration of the Crustacea are so intimately connected with
the appendages, that they must be treated in immediate sequence to them.

The gills of Nephrons andamanica are attached on either side to the epimera
and appendages of certain of the thoracic somites. The space on either side in
which they lie is known as the gill-chamber : its inner wall is formed by the
epimera of the thoracic somites, its outer wall is formed by the side-wall of the
carapace, or hrancliiostegite.

In respiration water is drawn under the edge of the branchiostegite into
the branchial chamber, and streams forwards over the gills, until it passes out
again between the anterior edge of the branchiostegite and the epistome, the
current being maintained chiefly by the movements of the scaphognathite,
already described in connexion with the 2ud maxillfe.

There are twenty gills on either side, though the most anterior of them,
which is attached to the 2nd maxillipeds, is a rudiment.

They are arranged in three series in either branchial chamber : those of
the outer series, which with the epipodites are attached to the coxopodites of
the appendages, are known as podohrancUas ; those of the middle series, which
are attached to the membranes that connect the coxopodites with their somites
are known as artlirohranch'm ; while those of the inner series, which are attached
to the epimera of the somites, are known as phmrolrancMse.

The podobranchise are six in number on either side, including the rudi-
mentary one, and are attached one to each of the following appendao-es : the 2nd
maxillipeds (rudiment), the external maxillipeds, and the first four thoracic legs.

The arthrobranchiffi are ten on either side, and are attached in pairs to the
external maxillipeds and the first four thoracic le^s.

The pleurobranchiaj are four on either side, and are attached to the epimera
of the last four thoracic somites.

The number and disposition of the gills is expressed in the following
branchial formula., in which the thoracic somites are numbered in serial succes-
sion to the six somites of the head : —

The Branchial Formula of Nephrops andamanica.

Somites and

Podobi-aucliire

Artl

irobianchite.

tlieir

Pleuro-
brancliiic

appendages.

Epipodites. Anterior.

Posterior.

VII

VIII

IX

X

XI

XII

XIII

XIV

0 ep.
*r + ei).

1 + ep.
1 4 ep.
1 + ep.
I +ep.
l+ep.
0

0
0

0
0

0

0
0
0
0

1
1

I
1

=

0 + op.
1- + ep.
3 + ep.

3 + ep.

4 + op.
4 + ep.
4 + ep.
1

Total

6 + r+7 ep.

6

*

Very

small.

5

4

=

19 + r+7 ep.

10

Each gUl consists of a multitude of stiffisli vascular filaments, arranged
round a central stem, to form an elongate-pyramidal plume or brush. Very
long thread-like setx are also connected with the gill-plumes, and are numerous
enouo-h to form by their interlacing a sort of adjustable straining apparatus to
keep the gills from getting clogged by dirt.

The structure and arrangement of the gills are of the highest value in the classification of
the higher Crustacea. On this subject see Huxley, on the Classification and Distribution of the
Crayfishes, Proceedings of the Zoological Society for 1878, pp. 752-788.

7. Of the Digestive Si/stem.

The dio-estive system of Nephrops consists of the alimentary canal and of a
pair of large digestive glands.

The alimentary canal is a tube, of which the anterior part is greatly ex-
panded to form a stomach : it is for the most part thin-walled, and it passes w^th a
single curve, but without any coiling, straight through the body from the mouth,
which lies behind the 4th (mandibular) sternum, to the anus, which perforates
the ventral surface of the telson.

The mouth is a longitudinal median chink, bounded in front by an over-
lapping fold of the integument termed the labrum or upper lip, and behind by a
pair of fleshy leaves known as the metastoma or lower lip. On either side of the
mouth are the powerful mandibles, upon which, like the leaves of a book, lie the
two pairs of maxillae and the 1st pair of maxillipeds, these being closely followed
by the last two pairs of maxiUipeds.

A short " gullet " leads upwards from the mouth into the capacious
stomach : like the stomach, it is lined by an invagination of the cuticular exoskele-
ton which is cast off every time the animal moults.

The stomach is divided into two chambers — a wide anterior one in the roof of
which is a powerful crushing-apparatus formed by three gastric teeth, one median
and two lateral, and a narrower posterior chamber furnished with hairs which
interlace to form a filter.

From the posterior or jiyloric chamber the narrow intestine passes straight
back to the vent.

On either side of the stomach and anterior part of the intestine, the cavity
of the body is filled by a large digestive gland. This paired gland consists of a
vast number of tubules, which at length discharge into a duct that opens into
the anterior part of the intestine on either side.

8. Of the Circulation of the Blood.

The blood is colourless, and the circulatory system consists of a heart and
thin-walled blood vessels.

11

Tlie heart lies just beneath the dorsal surface of the cai'apace, in the middle
line, behind the cervical groove. It is enclosed in a semi-transparent sack known
as the pericardium, which also contains blood. The cavity of the heart com-
municates with the cavity of the pericardium by three pairs of valvular openings
{ostia), so that blood can pass from the pericardium into the heart, but not in
the reverse direction.

From the heart seven main arteries carry pure arterial blood to all parts of
the body. The ultimate ramifications of these arteries open freely into the
interstices of the tissues, whence the blood, after bathing the tissues, is at last
collected in vemus sinuses by which it is transported to the gills. After becoming
aerated in the gills, the blood passes by definite vessels into the pericardium and
so returns to the heart through the ostia in its walls.

9. Of the Benal Excretory Si/ste

m.

The organs corresponding in function with the kidneys of higher animals
are a pair of tubular coiled green-glands. These are lodged in the antero-lateral
corners of the cephalothorax and open at a tubercle on the basal joint of the
antennal peduncle.

10. Of tlie Central Nervous System.

The central nervous system is of the ordinary Arthropod type. It consists
(L) of a large composite cerebral ganglion placed immediately in front of the
gullet and behind the eyes in the middle line of the b'ody, (2) of a continuous
ventral cord or chain of ganglia running along the whole length of the A^entral or
sternal aspect of the body, in the middle Une, as far as the penultimate abdominal
somite, the ganglia of the chain corresponding in a general way with the segmen-
tation of the body, and (3) of a pair of comnmsures which form a long narrow
collar round the gullet and connect the cerebral ganglion with the ventral chain.

The cerebral ganglion, which is really double, gives off nerves to the eyes,
antennules, and antennae, and to the muscles which move those appendages.
The ganglia of the ventral chain send off nerves to all other parts of the body.

[The Voluntary Muscles of Nephrops, like those of Arthropoda in general
are made up of multi-nucleate striped fibres, and their tendons are derived from
invaginations of the integument (exoskeleton)].

11. Of the Organs of Reproducticrn.

These in the male are coiled tubular glands, which open upon the coxopodites
of the last pair of thoracic appendages (5th pair of legs).

12

In the female the oviducts opeu upon the coxopodites of the 3rd or ante^
penultimate pair of thoracic legs.

As already mentioned the 1st pair of abdominal appendages are modified to
assist the reproduction of the species.

The eggs when laid are attached to the 2nd-5th pairs of abdominal
appendages of the female, and the young when hatched do not differ greatly from
the adult in form.

We have no evidence on this point, but from the observations of Spence
Bate upon the common shore-crab of England, of Coste upon the crayfish and
certain crabs, of Andrews on Cambarus, of Wilson on Cancer pagimcs, and of
Brandes on Galathea and Eiqmgitrus, we may probably conclude that the eggs
are fertilized before they are laid, as in the case of the air-breathing Arthropods.

12. Of Moulting or Exuviation.

In its growth from the young to the adult stage and even beyond it, the
exoskeleton is periodically moulted.

At each moult the entire cuticle of the body and its appendages is shed
bodily like a mould, and with it the lining membrane of the stomach and the
gastric teeth.

After casting its exoskeleton, the animal remains for some days soft and
flabby, until the new cuticle hardens.

It seems probable that although this process of moulting takes place
periodically throughout the greater part of the life of the animals of this class,
yet that at last a limit of growth is reached and exuviation ceases : for we
often find old crabs encrusted with barnacles and tubicolous annehds which have
evidently been attached for a long time.

The Decapod Crustacea have also a certain power of renewing appendages,
or parts of appendages, that have been accidentally lost. From the stump of
the lost part a bud grows out, which in course of successive moults acquires
the form of the lost part. In consequence of this facility of repair, these
Crustacea will often voluntarily, on sufiicient occasion, discard an appendage.

On tbe general morphology and organization of the Decapod Crustacea the following works
must be consulted : —

H. Milne Edwards, Histoire Naturelle des Cnistaces, Vol. I.
De Haan, Fauna Japonica, Crustacea.

Dana, Uuited States Exploring Expedition, Crustacea, Part I.
Bell, History of British Stallc-eyed Crttstacea, Introdaction.
HcxLET, THE CRAYFISH (International Scientific Series).

Boas, Stndier over Decapodernes Slxgtskabsforhold (Vidensk. Selsk. Skr., 6 Ra?kke, naturvid-
og matheraat. Afd. I. 2. Kjobenhavn, 1880).

13

Gerstascker and Ortmann, Malacostraca iu Bronn's Klassea und OrJnungen des Thier-ReichS'
Spence Bate, Beport on the Challenger Crustacea Macrura, Introduction.
Stebbing, A History of Crustacea (luternational Scientific Series).

II. Classification of the Decafoda.

Altlioufjli the views of Boas, who divides the Crustacea of the Malacos-
tracous Order Decapoda into two suborders Xatantia and Rejifantia, have much
to commend them, yet I prefer to follow Milne Edwards I. and Dana (from
whom also DeHaan is not essentially divergent) who recognize three principal
groups of Decapoda, namely, Macrura, Anomura or Auomala, and Brachijura.

Of these three suborders the Macrura occupy the middle place, having
intimate connexions on the one hand with the AiiomaJa {Anomura) and on the
other hand with the Brachyura, while so far as can be seen at present, the two
latter groups {Anomala and Brachyura) are not connected with one another in
any other way but through the Macrura.

The most obvious links between the Macrura and the Anomala (Anomttra) are (1) the
Thalassinoids, which are almost uuited with the Paguroids through the curious symmetrical
hermit-crab Pijlocheles, and (2) the TJroptychoids, which, through Ptychogaster, have a less
striking connexion with the Astacoids. On the other hand the interval between the Macrura and
Brachyura is almost bridged by the deep-sea Dromioid genera Homolodromia and Archnodromia.

The three suborders of Decapoda are thus defined : —

Suborder I. Macrura. Body elongate. Abdomen as well developed as,
or even better developed than, the cephalothorax, behind which it is more or
less completely extended and beneath which it is never permanently flexed, and
endinof iu a large symmetrical tailfan the lateral lobes of which (caudal swim-
merets) are almost without exception foliaceous. Front not joined with the
epistome. Thoracic sternum usually narrow : if broad, the last segment is not
independently movable. The genital ducts never open on the sternum. The
antenna! peduncle commonly, though not universally, carries a movable exopodite,
which is generally foliaceous.

Suborder II. Anomala or Anomura. Abdomen less well developed than
the cephalothorax : although in a few exceptional forms it is elongate and
symmetrical and extended in a straight line behind the cephalothorax, it is as a
rule either (1) flexed or rolled on itself, or (2) flexed against the stenuim, or (3)
coiled in a spiral, in which last case it is more or less soft and asymmetrical. It
is only exceptionally that the 6th abdominal somite is destitute of appendages
homologous with caudal swimmerets. The front is not united with the epistome
across the ophthalmic somite. The last thoracic segment is independent, the
last segment of the thoracic sternum, when not atrophied, being movable. The
genital ducts never open on the sternum. If the autenual peduncle carries an
exopodite, this is never foliaceous.

14

Suborder III. Brachyuha. Abdomen much less well developed than the
cephalothoi'ax beneath which it is folded : its 6tli somite is almost always destitute
of appendages, but if, as is rarely the case {Dromiidea), they ai'e present, they
appear in a modified form as a pair of small lateral plates intercalated between the
6th and 7th somites. The front is joined with the epistome across the hidden
ophthalmic somite to form an interantennular septum. Thoracic sternum usually,
but not universally, broad, the last segment is fused with the others. The antenna!
peduncle never carries a movable exopodite. The genital ducts of the female
usually, but not always, open on the sternum ; those of the male sometimes do.

15

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16

THE BRACHYURA.

I, The Beachyuea in General.

1. Of the General Form of the Body in the Bmchyura.

In the Crabs, as in Nephrops, the body is divided into two regions — a
cephalothorax consisting of 14 somites fused together, and an abdomen consist-
ing of 7 ; but the abdomen of the crab is very much smaller than the cephalo-
thorax, beneath which it is usually folded out of sight, being often, in fact, in
the male little more than a rudiment.

As in Nephrops, the cephalothorax is covered by a large overlapping cara-
pace, which, however, is usually broader than long.

Moreover, although the abdomen as a whole is moveable, yet it is very
common to find some of its somites fused together : the abdominal terga do not
overlap one another, hardly ever are any of the sterna calcified, and hardly ever
are there recognizable pleura.

There are certain undoubted crabs (e.g., the Raninidm) in' which the cephalothorax is much
longer than broad and the abdomen, though reduced in size, is by no means folded out of sight.

Moreover there are other Deoapoda besides crabs {e.g , the Lithodidse &nd Porcellanidse) in
which the carapace may be broader than long and the abdomen is folded beneath it.

In the great majority of crabs the anterior end of the carapace is not pro-
longed to form a rostrum, but is, on the contrary, truucated and often deflexed.
As a result of this the eyestalks, instead of lying close together in the longi-
tudinal axis of the body, are forced asunder and pushed back, until they lie
almost in the transverse axis of the body. A similar but less marked change
of position is usually seen in the antennules and antennge.

Moreover, both the eyes and the antennules become more or less enfolded
or embedded in pits. When these pits are shallow and the second joint of the
antennal peduncle remains free, as in the Dromides, we find a pair of common
orbito-antonuular fossee into which the eyes and antennules are, or are not,
completely retractile. But when, as in most crabs, the second joint of the
antennal peduncle becomes more or less fixed, we find distinct antennular fossar
into which the antennules can bo folded, and distinct orhits for the eyes, the
second joint of the antennal peduncle forming the partition between the two
cavities.

There are numerous crabs (e.g , the Oxyrhyncha) in which tlie carapace is produced anteriorly
to form a rostrum, but in this case the rostrum always sends down from its under surface a
process that separates the antennules from one another.

Again tliere are many crabs, besides the Uomolidea, in which the eyes are not concealed in
orbits, and there are some ci'abs in whieli the eyes, though broadly separated from one another,
. are not pushed out at right angles to the sagittal axis of the body.

17

On tlie other hand tlieie are numerous Decapoda, besides ciabs, in which no rostrum is pre-
sent ; and in some Loi-icate Macrura the eyes lie at right angles to the long axis of the body and
are embedded in perfect orbits, thougli these are not formed in the same way as the Brachyurous
orbit.

The segments of the ceiihalothorax are immovably united together, as in
Nepliro])s, but the sterna of the last 5 or G segments are of great breadth.

The relative breadth of the Brachyurous " sternum " is very variable. In the Baninido}
the posterior thoracic sterna are even narrower than in Nephropa. On tho other hand there are
many Decapods besides crabs, e.g., the Loricates, the Galatheoids, the .Porcellanoids, and the
Lithodoids, in which there is a broad " sternum."

As in Neplirops, the cephalothorax is strengthened, without unnecessary
increase of weight, by an internal framework of ajwdemes. In most crabs in
addition to the transverse apodemes there is a longitudinal apodeme — the median
apodeme — formed by an infolding of the cuticle of some of the posterior thoracic
sterna in the middle line, and united posteriorly with the fu,sed and infolded
endosternal apodemes of the last thoracic sternum.

In the Bromides alone there is no median apodeme ; but the inner or endo-
sternal apodemes of the last thoracic sternum are strongly infolded longitudinally,
so as to lie close together in the middle line of the body and thus to form the
beginnings of a median apodeme.

The carapace of the Brachyura and its sutures and grooves will l)e con-
sidered subsequently.

2. Of the Ajppendagcs of the Brachyura.

The appendages of the cephalothorax correspond very closely with tliose of
Nephrons from which they differ only in certain details of form.

The 1st pair of aj)]JCiidages of the head are the ri/es : these are borne on
protopodites, of which the coxal joint is usually concealed, although in some
crabs (e.g., Homohi) it is of considerable length and is fully exposed, and they
are commonly lodged in orhits.

The simplest form of orbit (seen in the Dromiacea) is a shallow cavity into
which both the eye and the antennule are retractile : the most perfect form of
orbit is a deep pocket complete on all sides and completely separated from the
antennular fossee. In a considerable number of crabs the orbits are wanting.

The 2nd pair of appendages of the head arc the antennules, which resemble
those of Nejihrops in composition, but differ conspicuously in the extreme short-
ness of their flagella, and in being as a rule retractile into pits {anteaaular fossx)
beneath the front.

There are other Decapod Crustacea, besides crabs, in which the antennules are more or less
folded and their flagella much abbreviated. On tho other hand, there are not a few crabs in
which antennular fossa; are either altogether absent, or are not distinct from the orbits.

3

18

In all Bracliym-a, liowever, the basal joints of the antennulai* peduncles are
separated from one another by an inter-antennular .se])twm formed by the union of
the front with the epistome.

The 3/T? pair of appendages of the head are the antcnnse, which differ from
those of Neplirops in never having a movable antennal scale, and in the reduced
leno-th of the flagellum : moreover owing to complete fusion between the 2nd and
3rd joints (which form the so-called " basal joint ") the antennal peduncle consists
of only four joints, of which the 1st, or true basal joint, is, in crabs above the
level of the Bromides and Raninidse, reduced in size, fixed, and more or less con-
cealed. There are a good many crabs, especially among the Oxystomes, in which
the antennae are quite rudimentary.

There are certain Maciuia {e.g., t.he Loiicates) and plenty of Anomura, in whicli the 2ncl and
3rd joints of the antennal peduncle are fused to form a single piece and the size and mobility of
the 1st joint are considerably decreased.

The 4>th, Uh and Qtli pairs of appendages of the head are, respectively,
mandibles and two pairs of maxillge not essentially different from those of
Nephrops. The fold of the carapace in Avhich these appendages, along with the
three pairs of maxillipeds, lie is deep and is, usually, very distinctly dehmited both
at the sides an'd in front : it is known as the buccal cavern, and its roof — i.e., the
part that Hes between the epistome and the labrum — is known as the cndostonte
or " palate."

The appendages of the tliora.v do not essentially differ from those oi Nephrops :
the first three pairs are foot-jaws or maxillipeds, the fourth pair are great
chelipeds, and the last four pairs are , ambulatory legs which are never truly
chelate though sometimes the last pair or last two pairs may be subchelate.

The !.•</ and 2nd maxillipeds resemble those of Nephrops, but in the 1st
maxilhpeds the endopodite and epipodite are usually much larger, and in the
2nd maxilhpeds the exopodite is usually longer and broader and the podobranch
is usually well developed.

The 3rf?- or external maxillipeds have, in the great majority of crabs, quite
lost their pediform character and become broad valves or folding-doors that
completely close the broad buccal cavern in which all the other mouth-parts lie,
so that these are all concealed from view. This change in the character of the
external maxillipeds is mainly brought about by a broadening and flattening of
the ischiopodite and meropodite : the three terminal joints ttsually, but by no
means always, remain small and form a sort of "flagellum" or "palp" to the
enlarged meropodite. The exopodite, which is sometimes six>ken of as the
exognath, may either remain comparatively slender or may become as much
broadened as the ischiopodite and meropodite.

There are, however, many crabs in which the external maxillipeds are almost as distinctly
pediform as they are in Nephrops, and do not conceal the other appendages of the mouth : in

19

fact, we meet with every gradation between pediform and opercular external-maxillipeds. On
the other liand, there are a few Decapods outside the limits of the Brachyura, in which the
external maxillipeds are opercular.

In no crabs above the level of the Dromides are epipodites or podobranchia?
found on any of the last five thoracic appendages.

N.B. — In the larger chelipeds of crabs the meropodite is frequently spoken
of as the " arm," the carpopodite as the " wrist," and the propodite and dactyl-
ns together as the " hand," the body of the propodite being the " palm," its
process being the " fixed finger," and the dactylus being the " movable finger."

The appendages of the abdomen, like the abdomen itself, are of a rundimen-
tary or, at least, subordinate character.

In the male they are present on the first two abdominal somites only, and
are completely modified to form uniramous organs of copulation.

In the female they are present on the 2nd, 3rd, 4th and 5th somites as
slender setose biramous appendages to which the eggs, when laid, are attached.
In the Dromides alone the 1st abdominal somite has a pair of appendages, which
are slender short and uniramous.

In no crab does the 6th abdominal somite carry appendages : in the Uromn-
dea alone are these appendages represented— and then in a much modified form
— by a pair of little plates intercalated between the 6th and 7th abdominal
somites.

In certain other Decapods, besides crabs, the abdominal appendages are reduced in size and
number in one or both sexes. Instances occur among the Loricata, the Pagurids, and the
Galatheids ; but here we always tind the appendages of the 6tli somite present. Among the
Lithodidse however even the appendages of tlie 6ih somite are absent.

3. Of the Oir/uns of Uespiration of the Brachyura.

As in Nephrops, the gills are attached on either side of the body to the epi-
niera and appendages of certain of the thoracic somites, and lie in a branchial
chamber formed by the overlapping of the carapace on either side. Into this
chamber the water used in breathing finds its way, usually but not invariably
by a valvular orifice situated between the basal joints of the chelipeds and the
neighbouring margin of the carapace, and from it the water, when done with, is
expelled by a channel running in either side of the buccal cavern, the current
being maintained by the scaphognathite of the 2nd maxilla and the epipodites of
the maxiUipeds, especially those of the 1st pair. The entry to the chamber
is the afferent branchial orifu-r, the exit from it is the eff'erent branchial canal.
In the Oxystome crabs the efferent branchial canals unite and traverse the endos-
torae in the middle line, and in many Oxystomes the afferent branchial canals
lie on either side of the endostome.

20

The number of gill-plumes in either brancliial chamber varies from 20 to 6,
but the typical number is 9, arranged as follows : —

The Branchial Formula of Scylla serrata.

Somites and

their
appendages.

VII

Podobranchiffi.
0 ep.

Arthrobranchia?.

1 _

Pleuro-
branchia'.

0

r —
An

iterior.
0

Posterior.
0

0 ep.

VIII

1 + ep.

1

0

0

=

2 + ep.

IX

I+ep.

1

1

0

=

3 + ep.

X

0

1

1

0

=

2

XI

0

0

0

1

=

1

XII

0

0

0

1

=

1

XIII

0

0

0

0

=

0

XIV

0

0

0

0

^

0

Total 2+ Sep. 3 2 2= 9 + 3ep.

In the typical crabs the gills are phyllobranchise ; that is to say, the gill-
elements are broad thin leaves arranged in two series along a central stem, like
the barbs of a feather-vane. There are however, among the Bromides, to be
found trichobranchise and gills of an intermediate character.

In certain land-crabs although functional gills are present, aerial respiration
is in part carried on by the remarkably thick and vascular lining-membrane of
the branchial chambers.

4. Of the Vinceral Systems.

Except for the better definition of the buccal cavern and for the valviform
modification of the external maxillipeds, the Digestice System, of the Brachyura
is not essentially different from that of Nephrops.

Crabs feed in a very human fashion. Holding the food in one of the large chela% they tear
ofi pieces with the other and carry the morsels to the mouth.

The Circulatory and Renal Excretory Systems do not differ essentially from
those of Nephrops. Nor does the Central Nervous System, except that, in cor-
respondence with the shortening-up of the body, the ganglia of the ventral chain
are all united into a central mass or ring from which the nerve-cords radiate.

In the Bromides, however, this concentration of the ventral ganglia is not
quite complete.

5. Of Beiyrudnction, among Brachyura.

The reproductive organs resemble those of Nephrops. In the male their
ducts usually perforate the coxopodite of the last pair of thoracic legs, but

21

sometimes they perforate the corresponding segment of the sternum. In the
female they usually open, near the middle line of the body, on the sternum cor-
responding mth the ante-penultimate pair of legs, but occasionally they perforate
the coxopodites of these appendages themselves.

The ova are impregnated internally, the organs of insemination being the
modified 1st and 2nd pairs of abdominal appendages of the male. Of these the
1st pair are the largest, and are made up of a propodite followed by a long
rigid exopodite which is rolled like a scroll to form a hollow tube of considerable
length. The 2nd pair are slender and consist of a propodite followed by a
styliform endopodite which is either closely applied to the preceding appendage
or is actually lodged in its channel.

The extruded ova, which are small and very numerous, are attached to the
setose abdominal appendages of the female until they are hatched.

In all the Brachyura except a few species of land-cralis the young leaves
the egg as a zoasa.

In the typical crab-zoa?a the body consists of a cephalothorax enclosed in
a carapace, followed by a slender six-jointed abdomen that terminates in a fork.

The carapace is usually armed with toui- large spines, namely, one pointiu"-
forwards and corresponding with the rostrum, one very large one pointing back-
Avards, and one on either side.

The abdomen is without appendages, but the following cephalothoracic
appendages are present : — large stalked eyes, uniramous antennules, non-flagel-
late antenna? provided with an antennal scale, Uvge mandibles consistino- of
propodite only, two pairs of maxilky differing little in composition from those
of the adult, and two pairs of large biramous maxillipeds which are the prin-
cipal organs of locomotion at this free-swimming stage.

In the course of a series of moults the remaining appendages are gradually
acquired and at the last moult the zoa?a changes into a Merjalopa.

In the Megalopa the cephalothorax is much broader than in the zoxn and its
spines are usually much reduced in size. The abdomCn though relatively shorter
than in the zofea is still of considerable size, and though curved is carried out
behind the cephalothorax : it consists of 7 segments and the terminal fork is
replaced by an oval telson, and the abdominal appendages are present.

By a further series of moults the Megalopa gradually changes into the adult
form.

0. Of flic Braclujurovs Carcqjace.

In the typical crabs the carapace is a transversely-oval shield with sharply
defined lateral borders that separate the dorsal surface from the sidewalls, anil
marked by certain mtiires and groores which delimit definite areas or regions.

The most constant suture is one that runs along either sideviall, in a more
or less sinuous course, from the antero-lateral angle of the buccal cavern to the
postero-lateral angle of the carapace. This, which is known as the cjnmeral
fiuture, cuts off on either sidewall a ventral piece which may with much reason be
regarded as, in the main, the pleurum of the 4th or mandibular somite of the head.

The cervical groove, which is so conspicuous in Nephrops, can generally he
made out, though obscurely, on the Brachyurous carapace. It is most distinctly
traceable in the Dromides. Wlien present, it helps to define the gastric region
of the carapace : its anterior loop is 2^i'obably represented on the Brachyurous
carapace by a groove that is sometimes found marking off a suborbital areola.

Certain grooves or depressions are commonly noticeable which map out the
surface of the carapace into regions. The regions form three dorsal series — a
median and two lateral.

The median series includes the frontal region, which lies between the orbits;
the (jastric region, which occupies the greater part of the middle of the carapace
behind the front and coincides in a general way with the boundaries of the
stomach ; the eardiac region, a small area which lies in the middle line imme-
diately behind the gastric region and indicates the position of the heart ; and the
intestinal region, which follows the cardiac and overlies the intestine just where
it curves over the endosternal apodemes to enter the abdomen.

The lateral series include, on either side, the hepatic region, which occupies
the antero-lateral corner of the carapace behind the orbits ; and the hrancMal
region which takes up all the wing of the carapace outside the gastro-cardiac
region and behind the hepatic region.

On the sidewalls of the carapace the triangular space on either side of the
buccal cavern is known as the pterygostomian region, between which and the
hepatic region lies the subhepatic region.

Any or all of the dorsal regions of the carapace may be further divided
symmetrically into areolae. The gastric region is fairly constantly subdivided
into three areoliB — one of which is postero-mediau, the others being antero-
lateral. Again a narrow S23ace fringing the upper margin of the orbit is some-
times separated as the orbital region, while the space between the lower margin
of the orbit and the pterygostomian region is sometimes wide enough to be
distinguished as a suborbital region.

Mention must here bo made of tlie linese anomtcricse, whicli though characteristic of the sub-
order of Anomura {Anortiala) are also found among the B/af/i//«m of the Hqmolidean families.
They are the homologues of the epimeral sutures of the Brachyuran carapace, but they are found
high up on the sidewalls of the carapace or even on the dorsum itself.

7. Of the General Bionmiy of the Brachynra.
The majority of crabs are marine, but some live in brackish water, others

23

entirely in fresh water, wliile a considerable numljer are amphibious, and not a
few live entirely on land, only visiting the sea periodically.

The marine crabs are found everywhere, but are most numerous in wnrm
seas. Most of them are inhabitants of the littoral and shallow water, but the
moderate depths up to 50 fathoms are also rich in species. A fair number of
species are found in -50-100 fathoms, while the depths between 100 and 500
fathoms are by no means poor in crabs. Below 1000 fathoms, however, crabs
are not met with in abundance, although a species of Etlium. w^as dredged h\
the " Challengel- " at the enormous depth of 1875 fathoms. In these seas the
greatest depth at which a crab {Ethusa investigator is) has been found is loOO
fathoms.

I^he habits of the littoral and shallow-water species are extremely variable.
Many species shelter in the rocks and reefs, and some burrow in sand and shingle
and mud. Many sluggish and inoffensive species are curiously coloured and
sculptured so as to resemble the broken shells and bits of eroded and incrusted
coi-al-rock among which they live, while other such species actually deck them-
selves mth a patchwork of pieces of seaweed and fragments of shell so as to
escape notice. Numerovis small species pass their lives esconced among the
bi'anches of living zoophyte-colonies, or among the spines of Echinoderms, and
are coloured exactly like their hosts. Many Bromiidas carry about a hoijse of
their ovm., in the form of a sponge which fits tightly over their carapace ; whih'
other species of this family, and of the Borlppida', roof themselves over "with the
dead valve of a Lamellibranch shell, upon which it is common to find a sea-
anemone growing. Other small species (notably among the T'innoteridse) live as
commensals in the mantle-chamber of living bivalve moUusks, or in the cloaca of
Holothurians, or in the tubes of Annelids, or in the atrial cavity of Asgidians.
Some bold and active species, such as the Grapsi, expose themselves fearlessly
on the rocks between tide-marks, trusting to their speed and craft to escape
their enemies ; while other active species, such as the PortunidsB and Matididx,
have their legs modified to form paddles by means of Avhich they swim with
much ease and speed. A few species {Plagvsia, Planes, Yaruna) make their
home on drift timber, drift seaweed, hulls of ships, etc., and so may be regarded
as belonging to the oceanic fauna.

The majority of freshimin- crabs belong to the CycloniGtope family of
Telplmsidx, although there are numerous Catometopes that also live in fresh-
water. During the hot and dry season some of these are accustomed to aestivate,
burrowing deep into the moister layers of the soil.

The ampldbions crabs are for the most jjart Catometopes. They are
generally gregarious and live close to the seashore, or among estuarine swamps,
in biirrows. They obtain their food along shore, and can breathe air so long as
their gill-chambers are kept moist. Although their burrows are usually flooded

24

at high tide, this is by no means always the case. Some of the most intelligent
of all the crabs are of amphibious habit.

Most of the Imid-crahs also belong to the Gaiomeiopa, though a few species
of Telphuddai are terrene. They prefer damp jungle, where they either burrow
in the earth or crouch under roots and fallen timber. Though their gills are
well developed, they also in part respire by means of the thickened linmg-
membrane of their greatly-enlarged branchial chambers ; but it is essential that
the air should be moist. Though they are typical inhabitants of the moist
jungles of the islanda and seaboard, yet they are also found at elevations of
several thousand feet in places where the rainfall is heavy.

Of the deep-sea crah.-i, those that live at moderate depths, to which it is
possible that a certain amount of light penetrates, very often have the eyes
enlarged; while those that live in the uuilluminated abysses very commonly
have the eyes small and deficient in pigment.

Although no existing members of the Arthropod phylum exceed the largest
Brachyura in size, yet the size of the body in this suborder is extremely variable.
At "the one extreme we find GoUodes malabaricus, an inhabitant of these seas, of
which the carapace, in an adult and egg-laden female, is less than one-sixth of
an inch in its greatest diameter, and Ehalia diadmnena, another inhabitant of
these seas, the greatest diameter of the carapace of an egg-laden female of which
is less than one-fifth of an inch : at the other extreme we have the Australian
Pseiulocarcinus gicjas whose carapace is said to reach a breadth of about two feet,
and the Japanese Kaempferia (Macrochira) Kaempferi the span of whose cheli-
peds attains to ten feet.

The Brachyura as a whole are the most highly organized and most intellig-
ent of the Crustacean" class. The Cyclometopes and Catometopes are particu-
larly alert and active, and among several of the gregarious species of amphibious
Catometopes life appeal's to be almost as complex as it is among many insects.

Although parental care does not go beyond the simple and almost mechanic-
al stage of carrying the eggs until they are hatched, yet in some Brachyura the
sexual feelings are sufficiently sustained and complex to give rise to well marked
" secondary " differences between the sexes and between the adults and immature
young of the male sex.

If the males are not larger than the females, yet, at any rate, it is very
common to find the male having one or both of its chelipeds very much larger
than those of the female, the difference in size being usually most marked in the
chelse. We know that in some cases the males use their enlarged chelipeds for
fighting with each other. On the other hand, it must be remembered that there
are many species of crabs in which the chelipeds show no sexual differences
whatever.

25

Differences in coloin- between the sexes have also been noticed, though they
do not appear to be of common occurrence. In the Oelasimi one cheliped of the
adult male is not only enormously enlarged, but its chelaj are also brilhantly
coloured. In one species of this genus the brightly coloured chelaj are obtru-
sively displayed by rival males, apparently to attract the female. And in more
than one species of the same genus the carapace of the adult male is known to
be more brilliantly coloured than that of the female, at any rate under the
influence of sexual rivalry and excitement.

Several Brachyura {e.g., species of Matnfri, Oci/jwda and Metaplax) possess
organs for the production of sound. These organs consist of two opposable file-
like ridges, the ^^brations of which, when they are rhythmically rubbed against
one another, give rise to grating and whirring sounds of considerable power.
The more perfect of the two ridges is placed on some part of the flexor aspect of
the cheliped ; the opposed ridge may either be placed on another part of the
cheliped or below the orbit.

In Mai Ufa and Ocijpoda both sexes possess these organs and have been
observed to use them when moved by fear or anger. In MefapJa.c the male alone
has them, so that in this case they are probably used to express sexual emotions.

As regards the part they play in Nature, the crabs, in common with other
Crustacea, are generally regariled as scavengers. They are more particularly
the scavengers of the sea-shore, and in tropical islands land- and strand-crabs
are of much importance as land-scavengers also.

On the other hand the marine species are one of the principal sources of
food of numerous fishes, especially of Rays, and remains of small crabs have
been found in the stomach of starfish. Among the chief depredators of the
shore and freshwater and amphibious species in this country may be mentioned
water-lizards {Varanus) and ci'ocodiles and occasionally frogs, swimming- and
Avadiug-birds of all kinds, kites and crows, jackals and other small carnivorous
mammals and sometimes pigs and monkeys.

[Ou the general bionoiiiy of N[alapo.straca see Ortmanii, in Bioim's Tliier-Rcicli, Malacos-
traca,pp. llSl-1262].

II, Gexeeal Classification' of thk Bi;A('uvri;A.

Class CRUSTACEA.

Order MALACOSTRACA.

Suborder Beachyuka, Leach, Boas, Stebbing.

Caneri Brachyuri, Liuneus, SyBtema Naturu-, ed. xii. torn. I, para ii. p. 1038 (1767).
Kleistagnatha {paxt), Fabrioius, Entomologioe Systcmaticiu Supplementum, p. 307 (ITli'^l.
Caneri Brnchyuri (part), Lamarck, Syat. des Animaux sans vertebres, p. 148 (1801).

4

26

Brachyuri, Latrcille, Hist. N:it. des CrastacL'3 efc des Insectes, Vol. V, p.. 3-17 (1803): Genera Cra8caceorum et
Insectoi-am, Vol. 1, p. 35 (1806) : Fam. Nat. du Regne Animal, p. 267 (1825)— exc. Lithodcs.

Brachyura, Leach, Transactions of the Linnean Society, Vol. XI, 1815, p. 307.

Brachyures, Desmarest, Considerations Generales sur la Classe des Crustacea, p. 84 (1825) — exc. Litliodes.

Brachyures, Milne Edwards, Hist. Nat. des Crustaces, Vol. I, p. 247 (1834), plus Atiomoures Aptenircs,
Vol. II, p. 168.

Brachyura, Macleay, Illustrations of the Annalosa of S. Africa, p. 54 (1838).

Brachygnatha et Brachyura Oxystomata, De Haan, Crustacea in Siebold's Fauna Japonica (1850).
' Brachyu.r<i, Dana, United States Exploring Expedition, Vol. XIII. Crustacea, Part I, p. 58 (1852), plna
Anomoura S^operiora, p. 400,

Brachyura, Heller, Crustaceea des siidlichen Enropa, p. 17 (1863), plus Ano/iiura Aptcrura, p. 143.

Brachyura, Sliers, Catalogue of the Crustacea of New Zealand, p. 1 (1876): Report on the Gliallenger
Brachyura 188G p. 1 : [plus Anomura Droinidea et Raninidea, Henderson, Report on the Challenger Anomura, 1888].

CAUfiNOMORPHA : Hdxley, Pboc. Zool. Soc, 1878, p. 785.

Brachyura, Clans, Grundziige der Zoologie, 4th ed., Vol. I, p. 632 (1880)— exc. PorceUanid:e et Lithodidie.

Brachyura, Boas, Studier over Decapodernes Sl^gtskabsforhold (Vidensk, Selsk. Skr., 6 Rajkke, natar-
videnskabelig og niathematisk Afd. I. 2) Kjobenhavn 1880, pp. 159, 200.

Brachyura, Haswell, Catalogue of the Australian Crastacea, p. 1 (1882) : plus Anomoura superiora, p. 138.

Brachyura, Stebbing, A History of Crustacea, p. 51 (1893).

Dromiiden, Oxystomnta and Brachyura, Ortmann, Malacostraca, pp. 1153, 1150, 1163, iu Broun, Klasseu und
Orduungeu des Thier-Ueichs, Bd. V, new series.

Carapace with a well defined lateral border to the greater part of its dorsal
surface, and with an epimeral suture (or its homologue, a liaea anomurica) on
either side.

" Front " always in contact with the cpistome, and often broad.

Abdomen much smaller than the cephalothorax, beneath which it is folded :
the abdominal pleura, when distinguishable, are in the same plane and in the
same straight line with the abdominal terga.

Thoracic sternum usually broad, the last segment never independently
movable : usually channelled in the middle line for the lodgment of the male
abdomen.

The eyes in repose are commonly carried at right angles to the longitudinal
axis of the body, and are often lodged in orbits.

The antennules are always separated from one another by a septum derived
from the " front : " their basal joint is swollen and enlarged, their flagella are
extremely short, and they are commonly retractile into fosste beneath the front.

The antennal peduncle consists of 4 joints, but the first joint is often reduced
in size, cramped up, and concealed : there is never a movable antennal scale, and
the antennal flagellum is usually shorter than the carapace.

All the mouth-parts lie in a well defined fold of the carapace or buccal
cavern. The mandibles consist of a strongly calcified protopodite, the free end
of which forms a triangular trenchant incisor process, and of a three-jointed
incurved palp.

The coxa and basis of the 1st and 2nd maxillae arc well developed, those o£
the 2nd maxillge being deeply cleft. The endopodite (palp) of the 1st maxillos
usually curves obliquely outwards. The scaphognathite is a very broad plate.

27

The flagella, when present, of the exopodites of the maxilhpeds are ahnost
always bent inwards nearly at a right angle, and the epipodites of the maxillipeds
are ahnost always very large,

Tn a great majority of crabs the ischium and merus of the external maxilli-
peds are broad plates, which completely close the buccal cavern like doors and
conceal the other mouth-parts.

The thoracic legs are six-jointed, their basipodite and ischium being fused
to form a single piece. None of these legs have exopodites, and it is only in one
small primitive group that any of them carry epipodites or podobranchifp. The
1st i^aii', which ax-e often enlai'ged, are chelate; the 2nd and 3rd pairs are mono-
dactylous ; the 4th and 5tli pairs may either be monodactylous, or sometimes one
or both of them may be subchelate and reduced in size.

Tn the male, modified uniramous appendages are present on the 1st two
abdominal somites. In the female biramous setose appendages are present on
the 2nd-5th abdominal somites. In the Divmidcx alone a pair of small uni-
ramous appendages are also present on the 1st abdominal somite, and in the
Bromiidea alone a pair of plates intercalated between the 6th and 7th abdominal
somites are the modified horaologues of apj^endages of the Gtli somite.

The ducts of the male reproductive organs open either on the coxopodites
of the last pair of thoracic legs, or, less commonly, on the corresponding seg-
ment of the sternum itself. The oviducts almost always open on the sternal
segment corresponding with the antepenultimate pair of thoracic legs, rarely
upon the coxjb of these legs themselves.

The gills are almost always phyllobranchias : in the Dromiidea alone they
may sometimes be trichobranchia?, or of an intermediate character.

Five main sections of the Brachyura are generally recognized, as is shown in
the following table : —

I. The efferent brancliial channels run along the middle line of the endo-
stome, so that the buccal cavern or, at any rate, that part of it which
corresponds with the endopodites of the external maxillipeds, is
elongate-triangular, its apex traversing or invading the extremely
narrow epistome. The 1st abdominal somite of the female is without
appendages. No appendages of any sort to the 6th abdominal somite OxvaiOMA.
II. The efferent branchial channels run along either side of the endostome,
so that the buccal cavern is more or less quadrangular, never acutely
triangular : —

1. The 1st or true basal joint of the antennal peduncle is a large,
exposed, and generally a freely-movable joint. Either
Unem anovuiriae are present on the carapace, or the append-
ages of the 6th abdominal somite are present as a pair of
plates intercalated between the 6th and 7th somites. The
1st abdominal somite of the female carries a pair of slender
uniramous appendages. The oviducts open on the coxo-

28

podites of the antepenultimate pair of legs. No median
apodeme
2. The 1st or true basal joint of tlie nntennal peduncle is small
and almost immovably embedded. Line.v anomnricie are
not present, nor are there any sort of representatives of
appendages on the 6th abdominal somite. No appendages
on the 1st abdominal somite of the female. The oviducts
open on the sternum. A median apodeme is present
[Brachyuea vera] : —

i. The vasa effereniia of the male pierce the coxo-
podites of the last pair of thoracic legs : —

a. Carapace narrowed anteriorly, the front
produced to form a distinct rostrum
which may be pointed or deeply bifid or
trifid : epistome usually long fore-and-
aft ...
p. The front is truncated and never forms a
rostrum: epistome short foie-and-aft ...
ii. The vasa efierentia of the male open either on the
last segment of the thoracic sternum, or if they
perforate the coxopodites of the last pair of
thoracic legs, then they pass obliquely forwards in
a special groove in the sternum

Dromides.

OXTRHTNCHA.

Cyclometopa.

Catometopa

BROMIDES or DROMIACEA.
[Dromiacea, De Haan).

Apterurcs et Homoliens (partim), Milre Edwards, Hist. Nat. Crust. II. 168, 180.
Dromiacca, De Haan, Fann. Japon. Crust., p. 102.

Dromidia vel Anomoura Maiidica Superiora, Dana, V. S. Expl. Exp., Crust, pt. I, p. 4O0.
Apterura, Heller, Crust. Sudl. Europ., p. 1-13.
Anomoura Dromidea, Miers, Cat. Crust. New Zealand, p. 57.

Dromiacea, Boas, Reoherohes sur les affinites des Crustacc-s Dccapodes (Vid. Selst Skr., 6 Eaokke, nat. (i;j;
math. Afd. I. 2, 1880, p. 202).

Anomoura Dromidea, Haswell, Cat. Austral. Crust., p. 138.

Anomura Dromidea, Henderson, Challenger Reports, Zoology, Vol. XXVII., p. 2.

Dromiac^s, Bouvier, Bull. Soc. Philomath, de Paris, (8) VIll. 189(5, pp. 34-108.

Drominea, Stebbing, Hist. Crust., p. 133.

Dromiidea, Ortmann in Bronn's Thier-Reieh, Malacostraca, p. 1153.

Dromiacea vel Brachyura Primigenia, Alcock, Journ. Asiatic Soc. Bengal, Vo'. LXVIII. pt., 2, 1899, p. 12J.

Jhomiaceie, Milne Edwards et Bouvier, Crust. Decap. de I'Hirondelle (suppl.jet dela Princesse Alice, Monaco,
1899, p. 8, and Exp. Sci. du Travaillenr et du Talisman, Crust. Decapod, pt. 1, p. 5.

Carapace seldom broader tlian long, often— as also the appendages — pilose.
Front narrow.

Either orbits and antennular pits are wanting, or there are common orbito-
antennular pits into which both the eyes and the antennules are retractile.

The antennal peduncle consists of four separate and freely-movable joints,
in the basal one of which the renal tubercle is prominent and conspicuous : tlie
antennal flagellum is comparatively long.

29

The epistome is triangular or truncate-triangular, and is well delimited from
the palate.

The buccal cavern is quadrilateral, but is usually broader in front than
behind. The external maxillipeds may ])o pediform, or sub-pediform, or com-
pletely opercular.

The last pair of legs arc dorsal in position, and, with few exceptions, are
prehensile slender and reduced in size, or even sometimes rudimentary. The
penultimate pair sometimes resemble the last pair.

The abdomen in both sexes is large, and usually consists of seven separate
seo-ments : in the male it has the usual anterior two pairs of modified copulatory
appendages, behind which, very occasionally, are 3 other pairs of rudimentary
appendages : in the female it has the usual four pairs of ovigerous appendages
and, in addition, a pair of smaller uniramous appendages situated on the first
segment.

The oviducts pei'forate the coxopodites of the antepenultimate pair of legs,
and the seminal ducts perforate the coxopodites of the last pair of legs.

The gills are usually phyllobranchife, but may be trichobranchias or may be
intermediate in character : they vai-y in number from 20 to 8 on either side.

All the Dromides are marine and many of them are found only in deej)
water.

I have proposed to divide the Drouiiacea into two tribal groups, namely : —

I. Dromiipea, in which the carapace has no liiiesi anomuricse, the eyes and
antennules are almost always retractile into orbito-antennular pits, the Gth pair
of abdominal appendages are commonly represented by a pair of small plates
intercalated between the Gth and 7th abdominal somites, and the sternum of the
female is traversed in more or less of its extent by two obliquely-longitudinal
grooves.

II. HoMOLiDEA, in which linrcS aiKiiii urine are usually present, the eyes and
antennules arc not retractile into ca^uties, the Gth pair of abdominal appendages
are not in any Avay represented, and the sternum of the female is not traversed
by any special grooves.

I. Dkomiiuka, Alcock.

Vrnmicns, Jlilne Edwards, Uist. Nat. Crust. 11. lliS.
DromicJa;, Henderson, Challenger Anomnrn, p. 2.

Dromidx et Dynomenidw, Ortmann in Bronn's Thicr-Reicli, ilalacostraca, p. 1155.

Promiiii.E et Dynomcnin^; A. Slilne Edwards & E. L. Bouvier, Crust. Decap. HirondoUe et PrinccBSe .\iice
Monaco, 1899, p. 9.

Dromiidii', Alcock, .Tourn. Asiatic Soc. Bengal, Vol. LXVIll, pt. 2, 1S',(9, p. 12.").

Carapace sometimes longer than broad, often broader than long, without
liiK'X auoiiiurica;.

30

Eyes and antennules almost always {nomoJodromia is the only exception)
retractile into common orbito-antennulary pits, the lower wall of which is formed
about equally (1) by the basal joint of the antennule itself, (2) by the basal joint
of the antenna, and (3) by a sub-orbital spine or dentiform lobe.

These orbito-antennulary pits very often show traces of a sub-division into
two fossae, one for the antennule the other for the eye — the boundary between
the two fossa? often being a tooth or a sort of fold in the upper margin of the
" orbit."

Bye of the ordinary form, situated at the end of a short stout eye-stalk, the
basal joint of the eye-stalk being inconspicuous.

Epistome triangular, its apex in close contact with the deflexed tip of the
front. Vault of the palate of good depth.

External maxillipeds iisually opercular, sometimes subpediform.

Fingers of the chelipeds generally short, stout, channelled along their
opposed surfaces, and strongly calcified in their distal half.

Sternum of the female traversed longitudinally, in part or in almost all of
its extent, by a pair of special grooves that sometimes end in special tubercles.

The abdomen of both sexes consists of seven separate segments. Very
often a pair of small lateral plates — the rudiments of the 6th pair of abdominal
appendages — is intercalated between the Gtli and 7th somites.

The gill-plumes vary in number from 20 to 14 on either side, and are either
trichobranchife or phyllobranchife.

Many of the species are protected by a commensal Sponge or Ascidian, or
by an empty valve of a Lamellibranch shell, carried over the back.

The Dromiidea are divided into three subfamilies which are distinguished as
follows : —

Synoims of the families of Dromiidea.

Tlie last two pairs of legs are reduced in size, subdorsal, and
preliensile : —

1. Lateral borders of carapace ill defined. Tlie appendages
of the 6th abdominal somite are not in any way represented.
Epipodites are present on the chelipeds and next two pairs

of legs. Gill-plumes 19 or 20 on either side, trichobranchia^ Homolodromid^.

2. Lateral borders of carapace well defined. Tlie appendages
of the 6tli abdominal somite are represented by a pair of
plates intercalated between that somite and the telsoii.
Epipodites may be present on tlie chelipeds, but not on any

■ ; of the legs. Gill-plunies 14 or 10 on either side, phyllo-

branchia? ... ... ... ... ... Dromid.k.

31

II. Only the last pair of less reduced iu size and subdorsal. Lateral
borders of carapace well defined. The appendages of the 6th
abdominal somite are represented as in the Bromidm. Epipodites
are present on the chelipeds and next two pairs of legs. According
to A. Milne Edwards and Bouvier the gill-plumes are 20 on either
side ... ... ... ... ... ... Dyxomenidj.

Of the nineteen or twenty known species of Indian Dromiidea only ten or
eleven are likely to fall in the way of ordinary collectors, the other eight being
known only from considerable depths.

Family Homolodromid.i:, Alcock.

Alcock, Jouru. Asiatic. Soc. Bengal, Vol. LXVIII, pt. 2, 1899, p. 127.

Carapace longer than broad, convex in both directions, the lateral borders
hardly defined.

Front cut into two prominent teeth, between which, but on a much lower
23laue, a third small tooth is sometimes present.

Antennal flagclla longer than the carapace.

External maxillipeds opercular but with a pediform cast.

Chelipeds equal, slender, though stouter tlian the legs.

First two pair of legs much longer than the chelipeds : last two pair much
shorter than the first two pair, subdorsal, prehensile or chelate.

The abdomen in both sexes consists of 7 separate segments : there are no
lateral platelets intercalated between the Gth and 7th segments.

The gills are trichobi'anchiie, or are intermediate between trichobranchia?
and phyllobranchiaj : the gill-plumes are very numerous — there may be as many
as 20 on either side.

Foliaceous epipodites are present on the chelipeds and next two pairs of legs.

The sternal grooves of the female are short, ending at the level of the
genital openings.

This family contains one Indian genus, ArarJniodrumia, which differs from
the other two known genera of the family as follows : from Homoludromia A. M.
Edw. in having the eyes and antennules retractile into orbito-antennular cavities,
and from Dicraaodromia A. M. Edw. in having a small arthrobranch above the
2nd maxillipeds.

All the members of the family belong to the Deep-sea fauna.

32
Aeachnodkojiia, Alcock.

Amchnodromia, Alcock, Investigator Deep Sea Brachyura, p. 17, and Joarn. Asiatic Soc. Beugal, LXVJII., pt.
2, 1899, p. 130.

Carapace elongate-oblong but somewhat broader behind than in front, deep,
inflated, tomentose, its texture thin but well calcified : two creases break either
lateral border, the posterior one being the more distinct and being continued to
the cardiac region ( = branchial groove), the anterior one, or true cervical groove,
not proceeding far on to the dorsum of the carapace. The front is horizontal,
prominent, and deeply bifid.

The antennule and eye of either side are completely retractile into a common
deep fossa (just as in Droviia) which affords them complete protection. As in
Dromia, the floor of this common antennular-orbital fossa is formed by a sub-
ocular (" antennal ") tooth in contact with the basal joint of the antenna, and, as
in Dromia, the outer wall of the orbit is breached by a wide gap. The orbital
portion of the fossa, Avhicli is loosely filled by the eyes, has the hollow for the
eyes much deeper than the hollow for the eyestalk. The eyestalks are slender,
the eyes small but jDerfectly formed and well pigmented.

The two basal joints of the antenniB, which are (juite freely movable, largely
fill the gap in the lower wall of the orbit, and lie in the same plane with the
antennules ; the second joint has its antero-external angle produced to form a
coarsish spine : the antennal flagella are longer than the carapace.

The palate is particularly well demarcated from the epistome and is rather
broader in front than behind : the ridges that define the expiratory canals are
very distinct. The epistome is in the closest possible contact mth the front, but
without complete fusion. The external maxillipeds are distinctly operculiform,
but owing to the moderate expansion of the merus and to the coarseness of the
palp, they have a slight pediform cast : they close the buccal cavern, but not so
tightly as in Dromia.

The chelipeds are equal and are rather slender, though considerably stouter
than the legs : the fingers are well calcified and are hollowed oi cuiJJere, the tip
of the dactylus shuts into a notch in the tip of the opposed finger.

The legs are cylindrical : the first two pairs are very long, the last two are
short, subdorsal in position, and cheliform rather than subcheliform.

The sternal grooves of the female end opposite the openings of the oviducts,
without tubercles.

The abdomen of both sexes consists of seven distinct segments. In both
sexes the pleura of the 3rd-6th abdominal somites are remarkably free and in-
dependent {i.e., not in contact with those in front and behind) and the last abdo-
minal tergum is nearly as long as the preceding five combined.

33

The branchial formula is as follows ;

Somites aud

their

Podobiaiichia\

Ai

;-tlii

I'obranchiiv.

Pl(

jurobraiicliije.

nppeudasjes.

VII

ep.

0

0

=

ep.

VIII

1+ep.

1

0

=

2 + ep.

IX

1 +ep.

2

0

=

3 + ep.

X

1+ep.

2

0

=

3 + ep.

XI

1+ep.

2

1

=

4 + ep.

XII

I+ep.

2

1

=

4 + ep.

XIII

0

2

1

=

3

XIV

0

0

1

=

1

Total

6 + 6ep. +

11

= 20 + (3 ep.

Araeliuixhwnia Baffiiu, Alcock and Anderson. Plato I., figs. 1, \ii-c.

Ararlitindrnmia Buffini, Alcock and Anderson, Ann. Blapr. Nat. Hist., Jan. 1899, p. 7.
For references see Alcock. J. A. S. B., Vol. LXVIII., [it. 2, 1899, p. 132.

Body and appendages tomentose.

Carapace deep — nearly as deep as broad — convex, from a fourth to a fifth
longer than broad, its greatest breadth being just in front of its posterior border :
except for a few sharp granules antero-laterally and along the ill-defined lateral
borders it is smooth, and the regions are hardly indicated. The front is deeply
cleft to its base, into two acutely triangular teeth.

Upper margin of orbit notched near its outer angle which is dentiform, the
outer angle of the lower margin of the orbit is much more strongly dentiform,
and the (outer) orbital wall between the two spines is deficient.

Antennal flagella longer than the carapace. The antennal and antennulai-
peduncles both surpass the tip of the front by their terminal joint.

Chelipeds rather slender, unarmed except for a few granules seen on denuda-
tion, about If times the length of the carapace: fingei's strongly hollowed 'en
cuiUrre^ especially the fixed one, which alone has teeth : wrist not elongate,
about as long as the dactylus and a little shorter than the palm.

First two pairs of legs more than twice the length of the carapace : their
dactyli are about two-thirds the length of the preceding joint, are stout, are
sharply spinate along the posterior edge, and end in a claw. The last two pairs
of leo-s are about the same length as the carapace : their small claw-like dactyli
shut down on a ring of spines at the end of the preceding joint.

Colours in spirit : dirty white, with a bluish tinge on the carapace and a faint
reddish tinge elsewhere ; eyes chocolate.

34

In the largest specimen, which is a female, the carapace is 30 milhm. long
and 2-i millim. broad, and the extended abdomen is 35 millim. long.

2371-2373 | Off tlie Travaneore coast T 430 fathoms.

" — Y^ Types of the species , ^at. 7° 17' 30" N. Long. 76^ 54' 30" E.J' "Investigator."

2^. Bay of Bengal, oS W. coast of Amlanians. 238-290 fathoms. " Investigator."
6

Named after the famous navigator William Baffin, who though best known

as an explorer of Arctic Seas, was, like his brilliant contemporary John Davis,

connected with the early history of our Indian empire. Both Davis and Baffin

lost their lives in Indian waters, Davis being treacherously murdered by jMalay

Pirates, Baffin being killed in action against the Portuguese at the entrance to

the Persian Gulf. " In the Indies he dyed, in the late Ormus businesse, slaine

in fight with a shot."

Family Dya^omenidjh, Ortmann.

Dynomenidie, Ortmann, in Bronn's Thier Reich, Malacostraca, p. 1155: Alcock, J. A. S. B., Vol. LXVllI., pt. 2,

1899, p. 127.

Dynomeniiu-e, A. Milne Edwards and E. Bonvier, Crust. Decap. Hirondelle et Princesse Alice, Monaco, 1899,

p. 9.

Carapace either longer than broad and convex, or broader than long and
flattish, the lateral borders well defined.

Front broadly triangular, sometimes notched at tip. Antennal flagella not
so long as the carapace.

External maxilhpeds typically opercular, completely closing the buccal cavern.

Chelipeds equal or slightly unequal, generally much stouter than the legs.

First three paii' of legs stout, about as long as the chelipeds. Fourth (last)
pair of legs dorsal and rudimentary.

The abdomen in both sexes consists of 7 segments, and there is a pair of
lateral platelets intercalated between the last two segments.

The (''ills are phyllobranchife but sometimes show the transition from tricho-
to phyllobranchia?. The branchial formula is said by A. ]\Iilne Edwards and
Bouvier (Crust. Decap. Hirondelle et Princesse Alice, Monaco, 1899, p. 10), to
be the same as that of Eomarus milcjari^.

The sternal grooves of the female end at the level of the genital openings.

This family contains two genera, both of Avhich are represented in the Indian
fauna.

Kiy fo the genera of Dynomenid^e.

I. Carapace flattish, broader than long, pilose ... ... ... Dtxomene.

II. Curapace convex, longer than broad, spiiiose ... ... ... Acanthodromia.

35

DvNOMENE, Latreille.

Diynomcne, Latreille in Cuvier'8 Rogne An. (nouv. eJ., 1820), p. 69: Desmareat (29), Milne Edwards (79),
Lamarck (54), De Hann (23), Dana (21), A. Milne Edwards (73), Ortmann (84).

Milne Edwards and Bonvier, Crust. Decnp. Hirondello et Princesse Alice, Monaco, 1899, p. 10.
For references see J. A. S. B., LXVIII., pt. 2, 1899, p. 133.

All parts usually tomentose.

Carapace subcircular or polygonal, flattisli, slightly broader than long.

Front broadly triangular, dorsally grooved, more or less distinctly notched
or divided at tip.

Palate well delimited from epistome : efferent branchial channels well
defined.

The chelipeds usually do not differ greatly in size from the first 3 pair of
legs : these are stout and of about equal length.

The 4th (last) pair of legs are quite rudimentary and alone are dorsal in
position.

The branchial formula, according to Milne Edwards and Bouvier, is the
same as that of Hoiwrnis vulgaris.

In the male (though not in the case of any known Indian species) in addition
to the usual appendages of the first two abdominal somites, rudimentary append-
ages may be present on the next 3 somites.

Dififrihition : Indo-Pacific, from Mauritius and Madagascar to California:
Tropical Atlantic in the neighbourhood of C. Verde Is. Occasionally descending
to depths of about 1-iO fathoms.

Bynomenc pilummiidcs, Alcock. Plate I., fig. 2.

Dynomenc ■pihnmioides, Alcock, J. A. S. B., LXVIIT., pt. 2, 1899, p. 133.

Body and appendages covered with a thick coat of stiff club-shaped hairs.

Carapace subcircular, flattish, the regions ill defined : a few inconspicuous
symmetrically disposed lumps on the gastric and anterior part of the branchial
regions.

Front broadly-triangular, deeply grooved in the middle line. Upper ])order
of orbit oblique, smooth, notched near the middle ; lower border smooth, notched
at the inner angle as in D. nrfula.

Lateral borders of carapace cut into 5 spiniform teeth, the last of which is
the smallest.

Chelipeds in the male a little unequal, the larger one about 1^ times as
long as the carapace and about a dactylus longer than the longest legs : except
for a few denticles along the outer edge of the wrist they are, when denuded.

36

smooth : the inner angle of the wrist is produced, and the fingers, which are
arched and spooned at tip, are shorter than the palm.

When the legs are denuded they are smooth except for some coarse and
inconspicuous serrations of the anterior border of the merus and carpus and for
a series of small teeth along the posterior border of the stout dactylus.

The last pair of legs are slender rudiments.

There are no vestiges of any appendages on the 3rd-5th abdominal somites
of the male.

In the single specimen the carapace is 10 millim. long and a little over 11
millim. in greatest breadth.

-— — . (Type of the species). Off Kiltan I., Laccadives. 50-30 fathoms. " Investigator."

Differs from D. hisjnda in the form of the carapace, the outline of which is
nearly a circle ; in the non-serrated orbital edges, and the non-spinulose chelipeds
and legs : from D. prxdator in having the antero-lateral borders of the carapace
cut into 5 spiniform teeth, and the wrist of the chelipeds of normal form : from
D. Ursula, in having the pterygostomian regions smooth and the legs and chehpeds
non-spinulose : from D. filhoH in having no rudimentary appendages on the 3rd-
5th abdominal somites of the male : and from D. pugnatrix in having the carapace
almost circular, the lower edge of the orbit non-serrated, the legs non-spinose,
and the chelipeds of normal form.

AcANTHODROMiA, A. Milne Edwards.

^ca;i»ioc!ramia, A. Jlilne Edwards, Bull. Jlus. Comp. Zool., 1880, p. 31 : E. L. Bouvier (12), Ortmann (84),
Alcock (3).

For references see Alcock, J. A. S. B., LXVIII., pt. 2, 1899, p. 13-1.

Differs from Dijnomene in having the carapace longer than broad, convex,
and closely covered with spines instead of hairs.

Distribution: Caribbean Sea and Andaman Sea, in depths of 75 to 150
fathoms.

Acanthodromia margarita, Alcock. Plate I., figs. 3, Sa.

Dynomcne margarita, Alcock, Investigator Deep-Sea Brachyura, p. 19, pi. ii„ fig. 3,

The whole carapace and dorsal surfaces of the chelipeds and legs are as
closely as possible covered with spines and spinules : the under surfaces of the
body and legs, the eye-stalks, antennte, and external maxillipeds are closely and
crisply granular.

On the middle of the fourth abdominal tergum is a pair of large smooth
tubercles, exactly like pearls, in the closest contact with one another.

37

Carapace sub-cylindrical, longer than broad ; the regions hardly indicated,
though the branchial groove is fairly plain.

Front triangular, deflexed, dorsally concave ; its apex is in close contact
with that of the epistome, and is surmounted by a horizontal spine similar to the
larger spines of the surface of the carapace. Supraorbital borders tumid.

Antennal flagellum nearly as long as the carapace.

Chelipeds equal, a little longer and stouter than the first three pair of legs,
and not much longer than the carapace. The fingers are short and stout, and
meet throughout their extent.

The last pair of legs are slender rudiments, hardly longer than the basal
joints of the other legs.

Colours in spirit, milk-white ; eyes deeply pigmented.

In the single small male the length of the carapace is 5 millim., the greatest
breadth a little over 4 millim.

^ , r Andaman Sea. "1 _. , ,

(Type of the spec.es). | ^^^^ ^g, ^^, ^_ ^3, g, ^_j. .0 fathoms. '• Investigator.

Family DkomiidJ'; ■■^.r.

Ccomuiie (partim), Dana, U. S. Expl. Exp. Crnst , pt. ii., p. 1428: Henderson, Challenger Anomnra, p. 2 :
Ortmann in Bronn's Thier Reich, Malacostraca, p. 1154.

Dromiime (partim), Milne Edwards and Bouvier, Crast. Decap. Hirondelle et Princesse Alice, Monaco, 1899
p. 9.

Dromiidie, Alcock, J. A. S. B., LXVIIL, pt. 2, 1899, p. 128.

Carapace variable, sometimes as long as or even a little longer than broad,
sometimes slightly broader than long; generally strongly convex in both direc-
tions, sometimes flat ; commonly ovoid or subcircular, occasionally pentagonal :
the lateral borders well defined in most of their extent.

Front usually cut into 3 teeth, the middle one of which is always on a much
lower plane than the others and is often of insignificant size or even absent : the
front is rarely triangular, \vithout lateral teeth. Antennal flagella shorter than
the carapace.

External maxillipeds typically opercular, completely closing the buccal cavern.

Chelipeds equal, generally much stouter than the legs.

First two i^air of legs generally stout, not much shorter than the chelipeds.

Last two pair of legs, subdorsal and prehensile, generally much reduced in
length and slender. There is a tendency for the fourth (last) pair to be a little
longer than the third pair, and occasionally the fourth pair are as long as either
of the first two pair.

The abdomen in both sexes consists of 7 segments, and there is a pair of
lateral platelets intercalated between the last two segments.

38

The gills are pliyllobrancliise and are either 14i (or 16) in number on either
side.

An epipodite of small size may be'present on the chelipedsbut not on any of
the legs.

The sternal grooves of the female are variable : they may end at the level
of the genital openings, or at the bases of the first pair of legs, or at the bases
of the chehpeds.

This family is represented in the seas of India by six genera.

Key to the Indian Genera of Dromiidfe.

I. Front much as in Dynomene, broadly triangular, dorsally grooved, notched
at tip. The sternal gi-ooves of the female do not quite reach to the
level of the genital openings on the antepenultimate pair of legs
II. Front usually cut into 3, sometimes into 2, teeth, rarely entire and
triangular. The sternal grooves of the female reach tlie level of tlie
bases of the 1st pair of true ambulatory legs : —

1. Third pair of ambulatory legs, though shorter, not less stout

than the first two pair, ending in a huge talon-like dactylus :
fourth (last) pair of legs short and very slender. Carapace
flat and pentagonal

2. Third pair of ambulatory legs similar to, though sometimes

shorter than, the fourth (last) pair. Carapace usually
convex : —

a. Legs smooth, the meropodites not specially dilated

b. Legs nodular, the meropodites not specially dilated ...

c. Legs nodular ; the meropodites of the chelipeds and

first or first two pair of ambulatory legs dilated,

petal-like
III. Front deeply cut into two broad lobes, each of which is broadly bifid : the
divergent sternal tubercles of the female are at the level of the
chelipeds ...

SPH-ERODnOMlA.

CONCHOECETES.

Dromia.
Cryptodeomia.

Petalomera.

Lasiodromia.

Sph.ekodeojiia, Alcock.

Sphs^rodromia, Alcock, Investigator Deep-Sea Brachyura, p. 16, and J. A. S. B., LXVIIl., pt. 2, 1899, p. 152.

All parts except the tips of the dactyli and fingers tomentose.

Carapace sub-globose. Front broadly triangular, grooved and notched in
the middle line as in Di/nomeitr.

Palate well delimited from the epistome : efPerent branchial channels | well
defined.

Chelipeds longer and stouter than the first two pairs of ambulatory legs,
which are not nodose. Last two pair of legs nearly similar to one another in
length and thickness, which are much less than those of the first two pair.

39

In tlie male, in addition to tlie usual appendages of the first two abdominal
somites, rudimentary appendages may be present on the next three somites.

The sternal grooves of the female are ividely separated and inconspicuous
and do not quite reach to the level of the genital pores, as in Dynomenc.

Branchial formula not known.

VMi-ihiitiou : Bay of Bengal, Andaman Sea, 70 to 112 fathoms.

Keij to flic sjh'cii-s of Sphferodromia.

I. The antei-o-laferal borders of tlie carapace run nearly up to the outer

orbital angles ... ... ... ... ... S. hnidalli.

II. The antero-liiteral borders of the carapace run nearly up to the angles

of the buccal cavern... ... ... ... ... S. mix.

SpJixroih-omia Krudalli, Alcock ^t Anderson. Plate IV., figs. IS, 18'r.

Dromidla KendalU, Alcock & Aiuleraon, J. A.S. B., LXIII.. pt. 2, 18!)1-, p. 173: Ulastriitioiis of the Zoolniry
of the Investigator, Crustacea, plate xxiv, figa. 1, lu.

All parts covered with a thick, yellowish, velvet-like tomentum.

Carapace subcircular, globose, smooth when denuded, cxcejit for some
scattered vesiculous granules on the pterygostomian regions and sidewalls ; the
cardiac region well defined and the " cervical " gi'oove distinct : the antero-
lateral borders, which are subcristiform and entire (except, perhaps, for a vesi-
culous granule or two) end at the orbital angle.

The front consists of two blunt triangular teeth. The upper border of the
orbit is smooth and slightly notched near the middle : the lower wall of the orbit,
outside the basal joints of the antennules and antenna; consists of a bluntly
triangular tooth : the outer orbital angle is rounded off.

The external maxillipeds when closed leave a wide gap between their
anterior border and that of the buccal cavern.

The chelipeds, which are about a dactylus length longer than the longest
legs, arc about 1^ times the length of the carapace : vesiculous granules are
present on all the edges of the arm, on the upper and outer surface of the wrist,
and everywhere on the hands except the lower part of the inner surface : the
wrist is a longish joint, not much shorter than the arm or palm, and has not its
inner angle well defined.

The last two pairs of legs are about half tli(^ length of the other two : eacli
ends in a small claw-hke dactylus which is opposed to 2 or 3 tiny spinules at the
end of the projiodite.

40

Carapace of the unique specimen 18 raillim. in either diameter.

Q'«2 C Bay of Beiio-al. 112 fathoms ) ^, , ,._, „

?±^. Ttp. OF T„K SPECIE.. I J.i4O5'55"Long.80'25'20" I' "I-est.gator.

Named after Commander C. J. Kendall, R.I.M., sometime surveyor on board
the Investigator.

Sphxrodromia nv.r, Alcock. Plate IV., fig. 19.

Sph^mdromia nux, Alooek, J. A. S. B., LXVIII., pt. 2, 1899, p. 151.

Carapace globose, finely frosted with granules when denuded: the well
defined lateral borders run down to the angles of the buccal cavern, and not to
the orbits.

The tooth which forms the lower wall of the orbit is very broadly triangular
and non-prominent.

When the external maxillipeds are closed there is a mere chink between
their anterior border and that of the buccal cavern.

The edo-es of the arm and the outer surface of the wrist and hand arc finely
granular, the granules of the wrist and hand having a tendency to run in lines :
the upper border of the wrist and palm are coarsely subcristiform.

Ill the male rudimentary appendages are present on the 3rd, 4th and 5th
abdominal somites.

Carapace of the largest specimen nearly 10 millim. long and nearly 11
millim. in greatest breadth.

3540

10

Types op the species. Gulf of Alartaban. 70 fathoms. " Itivesticrator."

Coxciiu:cETES, Stimpson.

Concluecetes, Stimpson, Proc. Acad. Kat. Sci. Pliilad. 1838(1859), p. 226: Alcock, J. A. S. B., LXVIH., pt. 2,
1899, p. 150.

All parts, except the dactyh (which are more or less hairy) and the tips of
the fingers, covered with a velvety tomentum.

Carapace flat, subpentagonal, non-elongate in the adult. Front cut into 3
teeth, of which the middle one is much smaller and is on a very much lower
plane than the others.

Palate well delimited from the epistome. Efferent branchial channels well
defined.

Chelipeds in the male much more massive and much longer than any of the
legs.

The third pair of ambulatory legs though shorter are not less massive than

41

the first 2 pair, and end iu a powerful talon-like dactylus. The fourth (last)
pair of legs are short and slender.

The sternal grooves of the female do not meet ; they end in scroll-like
tubercles on the second segment of the sternum, between the bases of the 1st
pair of ambulatory legs.

Tlie branchial formula is as follows : —

Arth

robraucbii'B.

Soniif'Ps fiTin f iiPiT*

PnnoriT'fl itf ni'K'

A

PleurobrancliifB.

appendages.

X \JUU>Jl(lll(.,rLlllt.<

Anterior.

Posterior.

VII

ep.

0

0

0

^

ep.

VIII

l+ep.

0

0

0

=

1 + ep.

IX

ep.

1

0

=

2 + ep.

X

ep.

i

0

=

2 + ep.

XI

0

1 (small)

1

=

3

XII

0

1 (sms

ill)

1

=

3

XIII

0

0

1

=

2

XIV

0

0

0

1

-

1

Total 1 + 4 ep. 5 4 4 = I4 + 4ep.

Distribution : Indo-Pacific from the Indus Delta on the west to Hons-kono-
on the east and to the coasts of Queensland on the south, in shallow water
usually.

Conchwcetes artificiosus, (Fabr.) Stimpson. Plate III., fig. IG.

Dromia artifciosa Fabriciug, Ent. Syst. Suppl., p. 360, Bosc (U), Latreille (55). •

Cancer arfificiosu^, Herb-st (-14). Conchcecetoi a)-h>'cio.su.s Stimpson (99), Headeraon (43), Alcock (3). Dromia
conchifera, Haswell (37 & 38).

For references see Journ. Asiatic Soc. Bengal, LXVIIL, pt. 2, 18P9, p. 151.

Carapace pentagonal, with the posterior border of the pentagon curved, its
dorsal surface quite flat, its greatest length (in the adult) about eqnal to its
greatest breadth, its regions all well defined by grooves, the cervical and bran-
chial furrows both equally well cut. There are sometimes a few granules near
the borders of the carapace.

Front cut into 3 teeth with granular edges, the middle tooth being smaller
and on a much lower plane than the others.

Upper border of orbit very oblique : a granidar spine or tooth marks the
true inner supra-orbital angle of higher Brachyura. Outer border of orbit
appai-ently wanting, but on denudation it is found to be defined by a concave
row of granules. Sub-orbital lobe granular and dentiform.

On the lateral borders of the carapace are usually two teeth, one immediately

behind the cervical groove, the other immediately behind the branchial groove :

one (the posterior) or both of these teeth may be nearly worn away, but usually

they are both very distinct. Between the first spine and the orbital tooth is a

6

42

(sometimes broken) row of granules, and between the same spine and the outer
angle of the buccal cavern is a row of granular tubercles : the surface of the
subhepatic region between these two rows of granules may, when denuded, be
granular or not.

The chelipeds of the adult male are, especially in the male, much more
massive, compared with the legs, than is usual among the DrQuiiidse: they are
also much longer than any of the legs. The outer (exposed) surfaces of all the
joints are more or less granular, some of the granules on the palm being very
large and visible without denudation : in addition, the upper border of the arm
is denticulate, there are 2 coarse tubercles at the far end of the outer surface of
the wrist, and 2 on the palm just behind the finger-joint.

The first 3 pair of ambulatory legs are short, and some of their joints are
granular and have a tendency to be nodulai', a nodule on the carpus being very
constant. Of these legs the 3rd pair ends in a characteristic stout talon-like
dactylus the tip of which bends towards a stout lobe at the proximal end of the
posterior border of the propodite.

The 4th (last) pair of legs are very slender : they reach to the far end of
the carpus of the 3rd pair, and end in a tiny claw-like dactylus.

In both sexes the abdomen has a convexity along the middle line.

This species protects itself with the valve of a Lamellibrancli shell, which is
held, as in a frame, by the strong hook-like dactyli of the third jiair of legs.

Distribution: as for the genus,
a 125

o

689-690

7 ■

1206-1207

7

4532
7 ■

3727-3728

y

7906

9
7946-7947

9

235-236
10 ■

Sandheads, R. Hooglily. Branch Pilots Milner and Barnett.

Madras. Purchased.

Port Blair, Andamans. G. H. Booley.

Madras. " luvestigator."

Orissa coast 7 t'atlioms. "luvestigator."

Sandheads, R. Hooghly. Branch Pilot Milncf.

Sandheads, Jt. Hooglily. Branch Pilot Daly.

Karachi. " Investigator."

-3593 Lat. 23° 33' N. Long. 67" 27" E.

^0-- 62 fathoms. 'Investigator.

43

Conchcpcetes andamaiiirm, Alcock. Plate III., fig. 17,

Conchrj'cetes nndamanicus, Alcock, Journ. Asiatic Soc. Bengal, LXVIII., pt, 2, 1899, p. 152,

As to the validity of this species, which is founded on three very small
specimens, I am by no means sure.

It differs from adults of C. arfiflciosus in the following particulars : —
The carapace is more depressed and a little more elongate. Though the
front is cut into 3 teeth the lateral ones alone are large and conspicuous, the
median one being minute and invisible in a dorsal view. There is no spine or
tooth on the upper border of the orbit. The antero-lateral borders though
granular are thin and overhanging and have no traces of spines or teeth. The
subhepatic regions are granular, but are not bounded by distinct rows of
granules.

Instead of two blunt tubei-cles behind the finger-joint of the chelipeds there
is a single large subacute tubercle.

In the largest specimen the carapace is 7"5 millim. long and 7 raillim.
broad.

2859

-= — . Port Blair, Aiidnmnns. G. H. Booley.

This species also roofs itself in with a valve of a dead Lamellibranch.

Dromia, Fabr.

Dromia, Fabricins, Ent. Syst. Suppl., p. 359. Latreille (55, 56 & 57), Leach (60), Riaso (89 & 90), Desmarest
(28), n. Milne Edwards (79), Lamarck (54), De Haan (23), Dana (21), Stimpsoa (99), Henderson (42), Ortmann
(84), Alcock (3).

For references see J. A. S. B., LXVIII., pt. 2, 1899, p. 136.

All parts except the tips of the fingers and of the dactyli are, generally,
tomentose.

" Carapace not elongate in the adult, strongly convex or subglobose.

Front cut into tlu'ee teeth, of which the middle one is on a lower plane than
the others and is often so much smaller than them and so much deflexed as to be
hardly visible from a dorsal view.

Palate well delimited from the epistome : efferent branchial channels well
defined, but not always bounded by distinct and unbroken ridges.

The chelipeds may have some of the joints nodose, but the ambulatory leo-s
are smooth.

None of the legs have tlie merus dilated. The last two pair of legs are
distinctly subcheliform, the spine at the end of the propodite against which the
dactylus closes being well developed.

The sternal grooves of the female do not meet, and they end opposite the
bases of the 1st pair of true (monodactylous) ambulatory legs.

u

The branahial formula of -Drowa'rt. E('!/;p/(M and D. cranioides is as follows,
but iu D. unidentata the ejpipodite is absent from the chelipeds :—

Arthrohranchise.

Somites and their
appendages.

Podobranchife.

^

T* 1 pn vnlimn

icliisB.

Anterior.

Posterior.

X ICUX U Lf 1 Of 1 1

• VII

ep.

0

0

,0

=

0 ep.

VIII

1 + ep,

0

0

0

=

1 + ep.

IX

0 ep.

1

0

=

2 + ep.

X

0 ep.

1

0

= .

2 + ep.

XI

0,

1 (small)

1

=

3

XII

0

1 (small)

1

=

3

XIII

0

0

1

=

2

XIV

0

0

0

1

=

1

Total l+-lep. f) 4 4 = 14+4 ep.

The posterior arthrobranchitB of somites XI and XII are short and abruptly
truncated.

Distribution : off the shores of all tropical and Avarmer temperate seas round
the globe, from the West Indies eastwards to Australia, and extending as far
north as the English Channel and Japan, in shallow water usually.

Key to the Indian species of Dromia.

I. An epipodite on the chelipeds : antero-lateral border of the carapace
cut into teeth : —

1. Carapace in the adult broader than long, the median frontal

tooth prominent in a dorsal view : last two pairs of legs of

nearly equal size ... ... ... ... D. ruvipJai.

2. Carapace in the adult at least as long as broad, the median

frontal tooth very inconspicuous in a dorsal view : last pair

of legs much longer than the penultimate pair ... D. cranioides.

II. No epipodite on the chelipeds: antero-lateral border of the carapace,
between the orbit and the cervical groove, entire : both the anterior
and the posterior borders of the propodite of the last pair of legs
end in large spines ... ... ... ... ... D. uiAdentata.

Dromia Bumpliii, Fabr. Plate II., fig. 4.

Dromia Rumphii, Fabricius, Ent. Syst. Suppl., p. 360.

[Ramph (92), Seba (94) ] Milne Edwards (79), De Haan (23), Stimpsou (99), Tozzetti (103), Hilgendorf (45),
Miers (67), Walker (105), Ortmann (82), Henderson (43), Alcock (3).

For references see J. A. S. B., LXVIII.. pt. 2, 1899, p. 137.

All parts, except the tips of the fingers and dactyli thickly covered with a
harsh tomentum, with sometimes scattered tufts of longer hair on the carapace.

Carapace in adults broader than long, strongly convex, smooth ; the cardiac

45

region and the branchial or " cervical " groove on either side of it plainly marked,
the gastric region faintly indicated.

Front cut into 3 nearly horizontal teeth of nearly equal, size, the middle one
on a lower plane and slightly the most prominent.

In young specimens a projection of the upper edge of the " orbit " marks
the position of the true inner supra-orbital angle of the higher Brachyura, liut in
large specimens tliis is obsolete.

The true antero-lateral borders of the carapace are cut into 3 sharp but
coarsish spines, the 2nd of which often has a small secondary denticle at its base.
The postero-lateral borders are convergent and have one large coarse spine,
placed immediately behind the cervical groove. In addition there is a spine on
the summit of the infra-orbital lobule, and another at the outer angle of the
buccal cavern.

The borders of the arm are dentate, especially the upper border, and there
are 2 or 3 teeth at the distal end of the upper border of the wrist and also alone-
the upper border of the hand : all these dentations tend to disappear with ao-e,
but two tubercles at the distal end of the outer surface of the wrist are persistent.

The last two pair of legs are about equal in lengtli, being hardly half as lono-
as either of the first two pair : their propodites are much shortened and their
dactyli are claw-like, forming chete with the opposing sjiines at the end of the
propodites.

Abdomen of male with a broad convex ridge down the middle line.

Sternal tubercles of female very pi'ominent.

An epipodite is present on the chelipeds.

Distribution : Indo-Pacific Seas from the Red Sea, Mozambique, and IMauri-
tius, to Japan.

1120. Persian Gulf. . . \V. T. Blauforcl, F. R. S.

1171. Tuticorin. 5 fathoms. "Investigator"

1527. Madras. Purchased.

785G

573-571>

7 ■

091

• T"'

3729
9 ■
5635-5636
9

1579

JO ■

3.539

10

Off Malabar coast. 29 fathoms. '• Investigator."

Ceylon. Colombo Museum.

Madras. Purchased.

Orissa coast. 25 fathoms, " Investigator."

Off Malabar coast. 2S fathoms. " Investigator."

G. of Martabau. 67 fathoms. " Inyestigator."

Off Malabar coast . ■1-9 fathoms. " Investigator."

46

Dromia cranioides, de Man. Plate II., fig. 5.

Dromidia cranioides, de Man, Journ. Linn. Soc, Zool., XXII., 1887-88, p. 208, pi. xiv., figs. 6-8 : Alcock,
.T. A. S. B., LXVIII., pt. 2, 1899, p. 138.

This species may perhaps prove to be identical with the Dromia incUca
briefly characterized by Gray (Zool. Miscell., p. 40).

Carapace globose, a little longer than broad, perfectly smooth except for the
cervical groove and for two small faint elevations side by side just behind the
front.

Front cut into 3 teeth, the middle one of which is so small and on a plane
so much lower than the others that it is hardly seen in a dorsal view.

A strongly-marked acuminate tooth near the middle of the upper border of
the orbit is equivalent to the inner supra-orbital angle of higher Brachyura.
Sub-orbital lobe dentiform, very prominent. Outer orbital angle well defined,
sub-dentiform. A tooth, Inxt not a strongly pronounced one, at the outer angle
of the buccal cavern.

True antero-lateral borders of the carapace cut into 3 or 4 teeth ; when 4, it
is by intercalation of a little tooth close to the base of the 1st.

Postero-lateral borders slightly convergent, Avith one tooth placed im-
mediately behind the branchial or " cervical " groove.

Borders of arm granular or obtusely denticulate, as also are the upper
border of the wrist and of the hand : two tubercles at the distal end of the
outer surface of the wrist.

The last two pair of legs have a claw-like dactylus which meets, in a cheU-
form manner, a spine at the end of the corresponding propodite. The last pair
are much longer than the last pair but one, being, in fact, very little shorter
than either of the first two pair.

Abdomen as in D. Enmi^lm.

The sternal grooves of the female approach one another closely, but do not
actually meet, on the segment of the sternum corresponding with the 1st pair of
ambulatory (monodactylous) legs, near the anterior end of which segment they
terminate, without tubercles.

The chelipeds carry an epipodite.

The length of the carapace of the largest specimen is 28 millim., its greatest
breadth is 26 milUm.

697. Port Blair, Andamans. Wood-Mason Coll.

^ili^ (Type of the species). Mergui. Anderson Coll.

6

^ and — Port Blair, Andamans. G. H. Boolej.

9967 W. Coral Bank, Andamans. „ j^^ggticator."

6 ■ 15 fathoms.

47
Dromi'i unidenfata, Riippell. Plate II., fig. 6.

Dromia unidentata, Kuppell, 2-t Krabben rotli. Meer., p. 16, pi. iv. fijr. 2, pi. xi. fif?. 9- H. Milne Edwards (79),
A. Milne Edw.irds {72J, Hilgendorf (45), Muller{81), Alcock (3). Dmmidia unidentata, Kosaraann (51), .le
Man (25), Cano (15), Heuderson (43), Ortmann (83).

For references see J. A. S. B., LXVIII., pt. 2, 1899, p. 139.

Carapace about as long as broad, strongly convex, with some dimples when
denuded, two of which, separating the post-gastric from the branchial regions,
are specially conspicuous. Cervical groove well marked.

Front cut into two broadish Init sharp teeth, between which, but on a voiy
much lower plane, is an extremely inconspicuous denticle.

A broad tooth near the middle of the upper border of the oi-bit. Outer
orbital angle prominent but not dentiform. Suborbital lobe bluntly dentiform,
but not prominent.

Antero-lateral borders entire, rather sharp. A slight projection, hardly
amounting to a tooth, on the postero-lateral border, immediately behind the
branchial or " cervical " groove.

Chelipeds smooth, except for two tubercles at the far end of the outer
surface of the wrist.

The fourth (last) pair of legs are not so very much shorter than either of
the first two pair and are very much longer than the ord pair. The propodites
of the last two pair are broader than long and are very spiny, one of the spines
in the case of the last pair being at least as long as the spine against which the
claw-like dactylus closes — so much so, that the last pair of legs appear to end in
o claw-like spines the middle one being the dactylus.

The abdomen of the male, when denuded, has a broad convex ridge down
the middle line ; but when not denuded, the terminal segments of the male
abdomen form with the basal joints of the chehpeds and first two pair of, legs a
remarkably fiat surface, owing to the abrupt angular bending up of the last three
abdominal segments.

The sternal grooves of the female approach one another closely, but do not
actually meet, on the segment of the sternum corresponding with the 1st pair of
monodactylous (ambulatory) legs, near the anterior end of which segment they
terminate, but without tubercles.

No epipodites on the chelipeds.

The carapace of the largest specimen is 24 millim. long, and 22*5 millim. in
greatest breadth.

In one of the male specimens, in which the vasa deferentia are, as usual,
wonderfully prominent, there are also openings in the basal joints of the ante-
penultimate pair of legs, corresponding with the genital openings of the female.

48

Distribution : Red Sea and East coast of Africa, Persian Gulf, Ceylon,
Coromandel coast, Andaman Sea, Malay Archipelago.

711. Port Blair, Andamans. Wood-Mason Coll.

8113
6 •

7K5

10 ■

Mergui. Anderson Coll.

Persian Gulf. F. W. Townsend.

Cryptodromia, Stimpson.

Crj/pfodromia, StimpaoD, Proc. Acad. Nat. Sci. Philad., 1858 (1859), p. 225. Miers (65), Haawell (38),
Henderson (42), Ortmann (84), Thomson (102), Alcock (3).

Epidromia, Kossmaun, Raise roth. Meer. Crust., p. 69.

For references see J. A. S. B., LXVIII., pt. 2, 1899, p. 140.

As Dromia, but the legs, or at least the first two pairs of them, as well as
the chelipeds, are nodular or verrucose. The ridges that define the efferent
branchial channels are distinct. The chelipeds may carry a small epipodite, but
not in any of the Indian species.

All the species are small.

Distribution. : Indo-Pacific, from the Red Sea, East coast of Africa, and
Mauritius, eastwards through the coasts of India and the East Indian Archi-
pelago, to China and Japan and Australia and New Zealand. Occasionally found
in moderate depths up to 150 fathoms, or even nearly up to 500 fathoms.

Kvij to the Indian species of Cryptodromia.

Carapace smooth (noii-gianular) : —

1. No .spines on dorsal surface of carapace : —

i. Front cut into 3 teeth, all of which are plainly vibible in a
dorsal view : autero-lateral borders of carapace witli
inoie than one tooth : legs nodular : —

((. Antero-lateral borders with 3 teeth (not including
I lie outer orbital angle and some teeth on the
subhepatic region) ... ... ... C. pileifera.

h. Antero-lateral borders with 2 teeth (not including
the outer orbital angle, etc.) : —

tt. Regions of carapace well defined : no
; tubercle on the surface of the maxilli-

peds ... ... ... C. canalicnhita.

r p. A pearl-like tubercle in the middle of the

exposed surface of the merus of the
external niaxillipeds ... ... 0. liiilliferu.

49

ii. Front cut into 3 teetli, tlie middle one of which is hardly
seen in a dorsal view : antero-lateral horders of carapace
with a single tooth, at their anterior end : legs hardly
nodular ... ... ... ... ... C. Hilgendorfii.

2. A dorsal spine on the hepatic region of the carapace, just behind

the outer orbital angle ... ., ... ... C. de Manii.

II. Carapace (and appendages also) profusely granular : the regions of the
carapace well defined and areolated : —

1. Carapace subcircular in outline, its antero-lateral borders non-

dentate, the median frontal tooth the most prominent ... G. ebaltoides.

2. Carapace pentagonal in outline, its antero-lateral borders dentate,

the lateral frontal teeth the most prominent ... ... C. Gilesii.

Cryptodromia pihifera, Alcock. Plate II., fig. 7.

Cryptodromia tuberculaia var. jnlcifem, Alcock, J. A. S. B., LXVIII., pt. 2, 1899, p. 141.

Body covered with a short scanty scurfy tomentura which does not conceal
the texture beneath.

Carapace broader than long, convex, smooth, without distinction of regions :
the cervical groove broad, shallow.

Front cut into 3 broad triangular teeth of about equal size, the middle one
of which is on a lower plane than the others and is deflexed.

A sharp tooth near the middle of the upper border of the orbit marks the
true inner supra-orbital angle. Outer orbital angle dentiform. Suborbital lobe
dentiform and very prominent.

True antero-lateral border cut into 3 or 4 blunt teeth : in the gap between
the 1st tooth and the outer orbital angle two subhepatic teeth — one of which is
large — may show up and may, from a dorsal view, look as if they belonged to
the antero-lateral border : there are two similar teeth, one alone of which is
conspicuous, at the outer angle of the buccal cavern. On the postcro-lateral
))order, at the branchial or cervical groove, is a denticle.

Wrist and palm, and corresponding joints of first two j^fiir of legs, sharply
and profusely nodular or tubercular on the outer surface : fingers compressed,
meeting only at tip.

The third pair of legs, though much slenderer and less nodular than the first
two pair and only about half their length, are fashioned on much the same plan,
except that the propodite is much shortened : the spinule at the end of the
propoditc of this pair is not always big enough to form a chela with the claw-like
dactylus.

Last (4th) pair of legs slender and smooth, a good deal longer than the
l>enultimate pair: their propodite has spines at the end of both borders, the

7

50

spine at the end of the anterioi- border being large enough to form a chela with
the dactylus.

Abdomen of the male slightly convex along the middle line, the 4th and 5th
terga with some little nodules : in the female the 3rd-5th terga have the surface
a little uneven, but not distinctly nodular.

Every specimen has a commensal sponge which covers it completely like a
cap.

In the Indian Museum are 70 specimens from the Andaman reefs.

The carapace of a large egg-laden female is 9 millim. long and 11 millim.
broad.

705. Port Blair, Andamans. Wood-Mason Coll.

19-tl-1950~1

7 '

^^-,o/,o, >■ Types op the species. Great Coco I. A. Alcock.

2272-2281 .'

~~7 J

55^. Off Little Andaman, ]0 fathoms. "Investigator."

9

Gryptodromia canalicidata, Stimpson. Plate II., fig. 8.

Cryptodromia cancdiculatn, Stimpson, Proc. Acad. Nat. Sci. Philad. 1858 (1859), p. 240. de Man (2-i),
Ortmann (82), Alcock (3).

Dromia tomenfosa, Heller (39). Cryptodromia fomenfosn, Hilgendorf (45), Kossmann (51). ? Cryptodt-nmia
penfagonalis, Hilgendorf (45;, Henderson (43), Ortmann (83).

For references see .T. A. S. B., LXVIIL, pt. 2, 1899, p. 142.

Tomentum short, close, velvet-like.

Carapace not quite as long as broad, only moderately convex, its surface
smooth, its regions very fairly indicated : the cervical groove is distinct, tlie
fronto-orbital region is marked off by a shallow transverse groove that runs
forwards from one antero-lateral angle of the carapace round to the other, and
the front itself is longitudinally grooved.

Front cut into 3 broad triangular teeth of nearly equal size, the middle
one nearly horizontal, but on a much lower plane than the others, whicli are
somewhat upcurved.

A tooth near the middle of the upper border of the orbit marks the position
of the true inner supra-orbital angle. Outer orbital angle dentiform. Infra-
orbital lobe dentiform and prominent.

True antero-lateral borders with 2 teeth of which the anterior is the moi-e
prominent and acute: in the well-marked gap between the 1st (large) tooth and
the outer orbital angle a stout subhepatic tooth shows up : below this again is a
tooth at the outer angle of the buccal cavern.

51

On the postei'o-lateral border, immediately behind the branchial or cervical
groove, is a tooth.

Outer surface of wrist nodular : a few nodules on upper liorder of palm :
fingers short and stout.

The carpus and propodite of the first 2 pair of legs are nodular.

Last 2 pair of legs short and slender, not nodular, not much more than half
the length of the first 2 pair : the 4th (last) pair very little longer than the 3rd,
Both end in a strong claw-like dactylus, but are hardly cheliform, although there
is a small spine at the end of the propodite of each.

Abdomen of male with a convex ridge down the middle line.

The carapace of the largest specimen is U millim. long, and nearly IG
millim. in breadth.

Distiibidion : Indo-Pacific Seas from the Red Sea and east coast of Africa to
the Loo-choo Is.

349. Persian Gulf. " W. T. Blanford.

■ Great Coco I., Andamans. A. Alcock.

Cryptodroiitia huUifcra, Alcock. Plate II., figs. 9, 9a.

Cryptodromia bullifera, Alcock, Journ. Asiatic Soc. Bengal, LXVIII., pt. 2, 1S99, p. 143.

Body covered with a short tomentum.

Carapace about as long as broad, convex, smooth, the cervical groove distinct.
Front cut into 3 acute rather slender teeth, the middle one of which is on a
lower plane and slenderer than the others. An acute spine near the middle of
the upper border of the orbit marks the position of the true inner supra-
orbital angle. Outer orbital angle spiniform. Suborbital lobe dentiform, faii-ly
|)rominent.

True antero- lateral borders of the carapace cut into 2 teeth, the anterior
being much the larger and spine-like. In the gap between the 1st tooth and
the onter orbital angle two small smooth subhepatic tubercles are visible, one
below the other.

An elegant pearl-liJce tubercle below fhem(b-orbif 1(1 lolii', a similar but .smnthr
tnhcrde in the middle of the ventral surface of the merm of, the external maxillipeds
and another in the middle of the ventral surface of the second joint of the antennal
peduncle, are characteristic.

An extremely inconspicuous denticle on the postero-lateral border, ])ehiud
the branchial or cervical groove.

Outer surface of wrist and upper surface of hand nodular, two of the
nodules on the wTist being particularly acute.

52

Outer surface of carpus and propodite of first 2 pair of legs broken but not
nodular.

Last 2 pair of legs slender and very short, ending in claw-like dactyli, but
not cheliform.

Abdomen of male convex along the middle line.

The carapace is between 5 and G millim. long, and about 5 millim. broad.

— -— . Off Ceylon. 34 fathoms. "Investigator."

y

52

— . (Type OF THE species). 7^ ni. E. of N. Cinque I. Andamans. 490 fathoms. "Investigator."

Gryptodromia de Mavii, Alcock.

Cryptodromia de Mani, Alcock, Journ. Asiatic Soc. Bengal, LXVIII., pt. 2, 1899, p. 144.

Carapace etc. tomentose.

Carapace as long as broad, convex, smooth, the cervical groove rather
indistinct. Front cut into 3 teeth, the middle one of which is the smallest and
is much deflexed.

A tooth near the middle of the upper border of the orbit (true inner supra-
orbital angle). Outer orbital angle dentiform. Suborl)ital lobe dentiform, but
not very prominent.

True antero-lateral border with two blunt teeth : two more blunt teeth on
the subhepatic border and one at the angle of the buccal cavern are continued on
from the antero-lateral border. A tiny denticle on the postero-lateral border,
just behind the branchial or cervical groove.

A tooth on the hepatic region, dorsad of the antero-lateral border, and just
behind the outer orbital angle, is characteristic.

Outer surface of wrist and upper surface of hand nodular ; outer surface of
hand granular.

Outer surface of carpus and propodite of first two pairs of legs uneven.
Last two pairs of legs short, ending in claw- like dactyh, notchehform; the
penultimate pair shorter than the last pair.

Carapace 5 millim. long and not quite 5 millim. broad.

— ^— . (Type op the species). Mergui. Anderson Coll.

The unique specimen is in too bad a state to be decently figured.

Cryptodromia Hilgendorji, de Man. Plate III., fig. IL

1 Hilgendorji, de Man, Arcbiv f. Natnrges. LI II., 1887
nil., pt. 2, 1899, p. 145: Nobili, .Ann. Mas. Genov. (

Carapace etc. with a short velvet-like tomentum.

C*yptod(07mo Hilgendorji, de Man, Arcbiv f. Natnrges. LIIl., 1887, i., p. 404, pi. xviii., fig. 'S : AIpi>ck,
J. A. S. B., LXVIII., pt. 2, 1899, p. 145 : Nobili, .Ann. Mas. Genov. (2) XX., 1899, p. 249.

Carapace longer than broad, convex, smooth, without distinction of regions.
€ervical groove broad and shallow.

Front cut into 3 teeth, the lateral ones broad and triangular, the middle
one so small and deflexed as to be little visible in a dorsal view.

There is no distinct tooth in the upper border of the orbit, but only an
angular bulge, to mark the position of the inner supra-orbital angle. Outer
orbital angle and sub-orbital lobe subdentiform.

The antero-lateral borders of the carapace are smooth and entire, but as
they bend sharply inwards towards the orbits their anterior angle forms a
forwardly-directed tooth, the space between which and the outer-orbital angle
is concave.

A very small prominence on the postero-lateral border, just behind the
branchial or cervical groove.

The chelipeds and legs have an uneven surface, but are not really nodular,
though both the inner and outer angles of the winst are strongly pronounced.

The last 2 pair of legs are short and slender, the 4th (last) pair being very
little longer than the 3rd ; both end in stout claw-like dactyli but are not
markedly cheliform.

The abdomen bends in very sharply from tlie -Itli segment, making the
under surface of the body very flat.

The carapace of the larger of the two Indian Museum specimens is
12 millim. long, and nearly 12 milhm. broad.

Distribution : Persian Gulf, East Indian Archipelago.

!^ Persian Gulf. A. R. S. Anderson.

10 ■

CnjpfoJromia chdlioides, Alcock. Plate III., fig. 12.

Dromia {Cnjptodromm) chalinide^, Alcock, Journ. Asiatic Soc. Bengal, LXVIII., pt. 2, 18^0, p. U5.

Carapace hardly at all tonientose : a few hairs on the borders of some of
the leg-joints.

Carapace subcircular with projecthig hont, convex, it.'< -vuf ace, I ilr I Imt <>/
the appendages, ecenjwhere do^ehj and rrisphj ,jranvhir : not only arc all its
regions well defined, but they ai-e also areolated, aV llw arn,l;r heiug convex and
particularly icell circumscribed.

Front longitudinally grooved, cut into 3 serrulate teeth, the middle one of
which is slender and prominent, while the lateral ones are broadly-triangular
and little prominent.

Upper border of orbit serrulate, very oblique, devoid of any tooth or spine.
Outer orbital angle and suborbital lobe not salient.

54

The lateral borders of the carapace, which run below the orbit to the angles
of the buccal cavern, and the suborbital lobes are serrulate but are not cut into
teeth, though there may be a granular tubercle in front of the cervical groove
and another behind it.

The chelipeds and legs are crisply and closely granular, the chehpeds and
first two pairs of legs being also nodular. In the wrist the inner, and even more
the outer, angle are dentiform.

Last two pairs of legs slender, hardly half the length of the first two pairs,
ending in claw-like dactyli, but not cheliform : the last pair are hardly longer
than the peniiltimate pair.

The abdominal terga are granular and more or less sculptured.

The carapace of the largest specimen is 7 millim. long and 8 millim. broad.
182-183

10

(Types of the species). Karachi. "Investigator."

Cryptodromia Gilesii, Alcock. Plate III., fig. 13.

Drcmla (Cryptodromia) Gilesii, Aloock, Joarn. Asiatic Soc. Bengal, LXVIII., pt. 2, 1899, p. 146.

No tomentum : only a few hairs on some of the leg-joints.

Carapace pentagonal, convex, its greatest length about equal to its greatest
breadth, the greater part of its surface covered with vesiculous granules : not
only are all the regions very distinct, but they are also areolated — the areolae
however not being so individually convex as they are in D. ehalioides.

Front cut into 3 triangular teeth, of which the middle one is the smallest
and is on a lower plane and obliquely deflexed.

Upper orbital border very oblique : a hardly noticeable angulation — not a
distinct tooth — mai'ks the true inner supra-orbital angle. Outer orbital angle
not pronounced. Suborbital lobe dentiform but inconspicuous.

Antero-lateral borders of the carapace cut into 5 small granular lobules or
tubercles, of which only 2 belong to the true antero-lateral border, the other 3
lieing on the subhepatic border and at the outer angle of the buccal cavern. A
granular tubercle on the postero-lateral border, just behind the cervical groove.

Legs and chelipeds crisply granular, the chelipeds and first 3 pair of legs
being also nodular : the nodules on the carpal joints being prominent and acute.

Last 2 pair of legs very slender, hardly half the length of the first 2 pair,
ending in hook-like dactyli, not cheliform.

All the abdominal terga are symmetrically sculptured and granular.

The carapace of an egg-laden female is 8 millim. long and 8^ millim. broad.

This species which is closely related to D. sculpta Haswell, is easily distin-

55

guished from I), eballoides (1) by the sharply pentagonal carapace and less-coin-
pletely isolated areolae, (2) by the much more prominent front, (3) by the ant(}ro-
lateral borders being broken by irregular tubercle-like lobules, and (4) l)y the
more abundant sculpture of the abdominal terga : in everything but the form of
the meropodites of the chelipeds and first pair of legs it strongly resembles

Fefalomera.

— ^ — '—. (Types OP THE si'ECiEs). Off Malabar coast, 28 fathoms. " Investigatoi'."

Named after Lt.-Col. G. M. J. Griles, I.M.S., who was Surgeon-Naturalist
to the Investis:ator from 1884-1888.

^t)

Petalomeua, Stimpson

Pefalomera, Stimpson, Proc. Acad. Nat. Sci. Philad. 1858 (1859), p. 226: Alcoek, J. A. S. B , LXVJII.
pt. 2, ]89y, p. 147.

Closely resembles Crypiodromia, especially those species (e.g. G. rbalioidf's
and Gilrsii) in which the carapace is granular and areolated, and indeed only
differs from Cri/ptodromia in having the upper border of the meroj^odites of the
chelipeds and next, or next two, pair of legs cristiform.

The sternal grooves of the female are widely separated and end on the
posterior part of the sternum corresponding with the 1st pair of (mono-
dactylous) ambulatory legs. '

The chelipeds carry a small epipodite.

Distrlhnfiou : Ceylon, Andamans, Japan, N. Australia.

Pctalomem imika, Alcoek. Plate III., figs. 14, 14(6.

Fetnlomcra granulata var. indicn, .\Icock, Joarnal Asiatic Soc. Bengal, LWIII., pt. 2, 1899, p. 148.

Carapace etc. hardly at all hairy : edges of the legs with some hairs.

Carapace a little longer than broad, convex in both directions, with very
numerous unevenly distributed vesiculous granules all the regions are distinct,
but are not all equally well defined. The cervical and branchial grooves are
both present.

Front dorsally grooved in the middle line, cut into 3 serrulate teeth, of
which the lateral ones are large and triangular, while the middle one is small
and is on a much lower plane.

Upper border of orbit serrulate : a tooth near its middle marks the true
inner supra-orbital angle. Outer orbital angle pi-onounced but not dentiform
The suborbital lobe forms a granular tubercle or denticle.

Antero-lateral borders of the carapace cut into 3 granular teeth the first
being subhepatic.

56

Chelipeds very mucli more massive than the legs : they and the first pair of
legfs have the merus petaloid, owing chiefly to the thin expanded crest-like upper
border of that joint. The merus of the next pair of legs is not petaloid, though
its upper border is sharp. In the chelipeds the inner border of the wrist and
the upper border of the palm are prominent and, like the upper and outer sur-
faces of those joints, are granular : there are also two sharp tubercles at the
distal end of the outer surface of the wrist.

The first two pair of legs have a few small granules on some of the joints.

The last two pair of legs are slender and end in small claw-like dactyli,
which are opposed to a very small spine at the end of the corresponding pro-
podites : the last pair of legs is very slightly longer than the penultimate pair.

In both sexes the abdomen has a convex ridge down the middle line and the
2nd-5th or 6th terga have a few scattered granules on their surface.

The largest specimen is slightly over 15 millim. long, and is 15 millim. broad,
but in young specimens the carapace is more elongate.

Colours of fresh spirit specimens : yellow with some reddish markings.

In the Indian Museum are 22 specimens, from the Andamans and from off
Ceylon 28 and 34 fathoms.

This species is to be distinguished from P. granulata only in not having the
merus of the second pair of legs (3rd pereiopods) petaloid.

From P. pnlchra Miers (Zool. H. M. S. " Alert," p. 2G0, pi. xxvii. fig. A), it

differs in having a tooth on the supra-orbital border, which border is serrulate,

not entire, and in being more granular.

713. (Types of the species). Port Blair, Andamans. Wood-Masou Coll.

8699 & 2839 , , p „ r„„i.„

Andamans. tr. H. Booley.

Off S. coast Ceylon, 34. fatlaoms. " Investigator."

Off N.-E. coast Ceylon, 28 fathoms. " Investigator."

Andamans. " Investigator."

Lasiodromia n.n.

Somalodromia, Miers (nee Homolodromia A. M. Edw,, 1S80), Zool. H. M. S. Alert, p. 553, 1884.
Pseudodromia, Alcock {nee Stimpson, Henderson, Ortmann), J. A. S. B., LXVIII., pt. 2, 1899, p. 149.

Body and appendages remarkably tomentose.

Carapace somewhat elongate. Front broad, deeply cut into two prominent
lobes, each of which again is broadly bifid : the gap between the two lobes is in
contact with the epistome.

6 7

5352-53.50

9
11.34

lo"'

3512
10 ■

57

Palate well delimited from the epistome : efferent branchial channels well
defined by low ridges.

Chelipeds not longer, and but little more massive, than the first two pair of
ambulatory legs. None of the legs verrucose or dilated. The last pair of legs
are slender, and are much longer than the penultimate pair and little shorter
than any of the others.

The sternal grooves of the female end in a pair of prominent juxtaposed
divergent tubercles placed at the level of the base of the chelipeds.

No epipodite on the chelipeds.

Distrihdiou : Seychelles, Laccadives, Ceylon.

The peculiar form and breadth of the front is due to the prominence of the supra-orbital
eave and especially of the supra-orbital tooth, which have fused with the front.

I am still in doubt on the question of separating Lasiodromia { = Eomalodromia) from

Pseiiidodromia.

The name Eomalodromia is so extremely similar in sound to Homolodromia (A. M. Edw., 1880),
that it would be very inconvenient to retain it. I therefore propose a new name Lasiodromia in
allusion to the long shaggy fringe of the anterior border of the carapace.

Lasiodromia Goppingeri (Miers). Plate III., figs. 15, 15a.

Eomalodromia Coppingeri, Miers, Zool. H. M. S. Alert, p. .55i, pi. L., fig. B.

Pseudodromia quadricornU (an Eomalodromia Coppingeri ?) Alcock, J. A. S. B., LXVIII., pt. 2, 1899, p. 149.

Body and appendages tomentose, with numerous long hairs: a hue of
peculiarly long silky hairs forms a fringe or false anterior border to the carapace
behind the deflexed front.

Front deflexed, dorsally grooved in the middle line, cut into two broad
teeth, each of which is fused at base with a broad and peculiarly prominent
supra-orbital tooth ; so that the front appears to consist of two widely-separated
lobes each of which is bifid.

Carapace in the adult longer than broad, slightly convex from side to side,
almost flat fore and aft behind the line of long hairs that marks the frontal
declivity : its surface, when denuded, is quite smooth : only the branchial or
cervical groove and the cardiac region are distinctly marked.

Lateral borders of carapace entire, except that there may be a tiny denticle
behind the branchial groove, slightly convergent posteriorly.

Outer orbital angle dentiform. Sub-orbital lobe dentiform, deflexed.
Chelipeds and legs comparatively slender, the chehpeds shorter and hardly
stouter than the legs. Two acute tubercles on the outer surface of the wrist.

Fourth (last) pair of legs little slenderer and about as long as either of the
first two pair, ending in a slender claw-Hke dactylus to which a spinule at the
end of the propodite is opposed to form an almost perfect chela.

8

68

Third pair of legs not less stout tlian, but only about half the length of,
the first two pair ; ending in a claw-like dactylus.

Length of carapace of an adult female 7 milUm., greatest breadth — in fe-ont
of the branchial groove — 6 millim.

8885

— 7-. OfE Pedro Shoal (N. o£ Laccadives). " Investigator."

— . Off S. coast Ceylon. 34 fathoms. "Investigator."

II. HOMOLIDEA, Alcock.

Somoliens (partim) Milne Edwards, Hist. Nat. Craat. II. p. 180.

Eomolidx, Henderson, Challenger Anomura, p. 18 : Ortmann in Bronu'a Thier- Reich, Malacostraca, p. 1155.
Eomolinee, A. Milne Edwards & E. L. Bouvier, Crast. Decap. Hirondelle et Princesse Alice, Monaco, 1899, p. 9.
Homolidea, Alcock, Journ. Asiatic See. Bengal, Vol. LXVIIL, pt. 2, 1899, p. 126.

Carapace longer than broad, linese anomuricx usually present. The eyes
are not retractile into orbits nor the antennules into pits. Basal antennular joint
subglobular.

The slender basal joint of the eyestalk is almost always plainly visible, and
is sometimes of great length.

The antennal flagella are, except in the Latreillidae, much longer than the
carapace.

The interantennulary septum is a distinct vertical process, and is not formed
merely by the close apposition of the apex of the epistome to the front.

The front forms a slender triangular prominent rostrum, which may be bifid
at tip and often has a spine on either side of its base.

The division between the epistome and palate is distinct, but the vault of
the palate is shallow.

External maxillipeds pediform or sub-operculiform.

The chelipeds and legs are long and slender : the fingers are not channelled
en cuillere. Only the last pair of legs is dorsal and reduced in size.

Sternum of the female broad, without any special longitudinal grooves.

The abdomen of the male, and usually but not always of the female also,
consists of seven separate segments. There are no lateral platelets intercalated
between the 6th and 7th segments.

The gills are phyllobranchijB, and the gill-plurpes vary in number from
14 to 8 on either side.

The species are, for the most part, confined to deep water.
The Homolidea are divided into two families as follows : —

59

Key to the families of Homolidea.

I. The slender basal joint of the eyestalk is not mucii longer, or may
even be much shorter, than the terminal joint. The gill-plnmes
are 13 or 14 on either side and epipodites are present on the
chelipeds and often also on the first two pairs of ambulatory legs Homolid^.

11. The slender basal joint of the eyestalk is much longer than the
terminal joint. The gill-plumes are 8 on either side and no epi-
podites are present on the chelipeds or legs ... ... Lateeillidj;.

The genus Hoviologenus Henderson = Hoinolopsis A. M. Edw. (A. Milne Edwards, Bull. Mus.
Comp. Zool. VIII. 1880, p. .34 : Hendei-son, Challenger Anomura, p. 20 : Bouvier, Bull. Soc.
Philomath. Paris (8) VIII. 1896, p. 63 : Ortmann, Bronn's Thier- Reich, Malacostraca, p. 1156 :
Milne Edwards and Bouvier, Crust. Decap. Hirondelle et Princesse Alice, Monaco, 1899, p. 12j,
appears to represent a di.stiuct section of the HomoUJse, differing from all other known Eomolidia
in the following characters: — the epistome is reduced and is hidden between the basal joints of
the antennules, the 2nd maxillipeds have neither an epipodite nor a gill-plume (podobranch), and
the external maxillipeds cany a podobranch.

Family HoMOLiDiE, Alcock.

Homolidse sensu restricto, Alcock, Jonrn. Asiatic Soc. Bengal, LXVIII., pt. 2, 1899, p. 130.

Carapace elongate-quadrangular, or ovoid, or urn-shaped. Antennal flagella
longer than the carapace. The slender basal joint of the eyestalk is either very
little longer, or a good deal shorter, than the terminal joint. The external
maxillipeds are pediform or subpediform. In all the Indian species that belon'r
to this family the gills are 1-i on either side and there are epipodites on the
chelipeds and first two pairs of ambulatory legs, the branchial formula being as
follows : —

Somites and

Arthi

obranchite

their
appendages.

Podobranchia;.

A

Pleurobranchias

c
Anterior.

Poster

or.

VII

ep.

0

0

0

=

ep.

VIII

1 + ep.

0

0

=

2 + ep.

IX

0 ep.

1

0

=

2 + ep.

X

0 ep.

1

0

=

2 + ep.

XI

0 ep.

1

1

=

3 + ep.

XII

0 ep.

1

1

=

3 + ep.

XIII

0

1

0

1

=

o

XIV

0

0

0

0

=

0

Total

1 (6ep,)

6

5

3

=

14 (Gep.)

Kei/ to tlie genera of Homolidae.

Carapace deep. External maxillipeds decidedly pediform. Terminal
joint of the eyestalk very much longer tban the basal joint, which
is entirely hidden by the basal joint of the antennule. Dactylus
of last pair of legs not nearly half the length of its propoditc and
closing against the iisiai end of the latter joint

Hypsophrts

60

II. External maxillipeds sabpediforni, the antero-external angle of the
menis being: dilated. Dactylus of last pair of legs about half as
long as its propodite and opposed to a tubercle on the posterior
border or proximal end of the latter joint. Basal joint of eyestalk
not entirely hidden : —

1. Carapace not depressed, its hepatic spine some distance

behind the level of the supra-orbital spine ... ... HoMOLi.

2. Carapace depressed, its hepatic spine almost on the same

level as the supra-orbital spine ... ... ... Paromolopsis.

None of the eiglit Indian species of Eomolidse are likely to fall in the way
of ordinary collectors, since five of them inhabit depths between 100 and 1000
fathoms, and of the other three only one has, so far, been taken in less than 50
fathoms of water.

HoMOLA, Leach.

Eomola, Leach, Trans. Linn. Soc. XI. 1815, p. 324, and Zool. Miscell. Vol. II., p. 82. Latreille (57),
Desmarest (29), Risso (90), Ronx (91), Milne Edwarda (79), de Haan (23), Dana (21), Heller (40), Henderson (42),
Ortmann (82 & 8i), A. Milne Edwards and Boavier (76), Alcock (3 & 4.)

For references, see J. A. S. B., LXVIIL, pt. 2, 1899, p. 154.

Carapace deep, longer than broad, quadrilateral or urn-shaped, with deep
vertical sides, the gastric region well demarcated and occupying the anterior
half of the carapace, the linea anomurica distinct and dorsal.

Front narrow, forming a rostrum, which is either entire or bifid at tip and
has a spine, often of large size, on either side of its base.

The orbits are quite incomplete and do not even conceal the eyestalks, and
the eyes, which project far outside them, are retractile against the sides of the
carapace. The eyestalks are long and the slender basal joint is slightly longer
than the terminal joint.

The epistome is fairly or very distinctly marked off from the palate. The
expiratory canals are very well defined. The external maxilHpeds are subpedi-
form.

The chelipeds are rather slender and generally somewliat spiny. The legs
are long and more or less compressed and spiny, the last pair are subcheliform,
but have the propodite dilated near the basal end and never twice the length of
the dactylus.

The abdomen of both sexes consists of seven separate segments and is rather
broad.

Distribution : West Indies and Atlantic coasts of North America, Azores
and Atlantic coasts of Europe, Mediterranean, East Indian Seas from Cape
Comorin to the Philippines : commonly at considerable depths up to nearly
500 fathoms.

61

The species of Homola may be groiiped in three subgenera, as foUo-ws : —

(1) Homola (Type H. harhata). Carapace quadrate, its broadest part being
in front, across the middle of the gastric region : the lines^ anomuricse keep close
to the lateral borders and are rather inconspicuous. Rostrum a bifid tooth, with
a small spine or tooth on either side of its base. The last pair of le^s reach to
the end of the preceding pair.

(2) HoMOLAX (Type H. megalops). Carapace urn-shaped, its broadest part
being behind, across the middle of the branchial regions : the linex anomuricse
run well inside the lateral borders and are conspicuous. Eostrum as in Soimla.
The last pair of legs reach beyond the carpus of the preceding pair.

(3) Paeomola Wood-Mason (Type H. cnvieri). Carapace decidedly macru-
rous in form, its greatest breadth being behind : i\\e liacse anomuricse yxvo. ^\e\l
inside the lateral borders and are very conspicuous. Rostrum a simple cylindrical
spine, flanked on either side by a spine of equal or greater size. The last
pair of legs do not reach beyond the end of the merus of the preceding pair.

Subgenus Homola.
Homola andamanica, Alcock. Plate IV., fig. 20.

Homola andamanica, Alcook, Investigator Deep-Sea Brachyura, p. 7, anil Illustrations of the Zool. of tlie
Investigator, Crust., pi. xl., 6g. 1 : J. A. S. B., LXVIII., pt. 2, 1899, p. 156.

Carapace elongate-subquadrilateral, its greatest breadth is across the middle
of the gastric region, behind which point its sides are quite straight and vertical :
it is well calcified, and, hke all other parts except the antennary flagella, is
covered with short soft but stiff hairs that are not thick-set enough to form a
coat of concealment.

Rostrum a depressed dorsally-grooved tooth, bifid at tip. Four spines on
the anterior border of the carapace, namely, one on either side of the rostrum
and one at either supra-orbital angle.

Lateral borders of dorsum, of carapace straight, very slightly convergent,
spinate ; the first spine, which stands alone on the hepatic region, is of pre-
eminent size, the second though much smaller than the first is much larger than
any of the others.

Gastric region very well demarcated, armed with nine large spines — three
in a triangle on either median area, one on either lateral area, and one on the
hinder part of the central area.

Some spines on the subocular, subhepatic, and pterygostomian regions —
lai'gest on the subocular region, where they are definitely arranged in two
crescentic rows. Two spines on the carapace outside the antenna-peduncle, in
addition to the spinuliform suborbital angle.

62

Eyes somewhat reuiform : the joint on whicli tliey are borne is shorter
than the slender basal joint.

Chelipeds slender, but distinctly stouter than the legs, more hairy than the
carapace, especially along the edges of the joints. Upper and lower borders
of arm spiny ; wi'ist with rows of spines on the outer surface and a spine or
two at the inner angle ; lower border of hand dentate, upper border of hand
denticulate, cutting edges of fingers sharp, entire.

Legs compressed, their edges plumed with short bristles, with long bristles
interspersed. The second and third pair, which are a dactyl-length longer than

the first, are not quite 2- times the length of the carapace : in all three pairs
both edges of the merus are armed with stout spines — at least in the distal half,
and the posterior border of the propodite and dactylus with compressed articu-
lated spines which are distant and acicular on the propodite, but stout very
regular and close-set on the dactylus.

The subcheliform fourth pair of legs reach very slightly beyond the end of
the carpus of the preceding pair : the merus has 3 or 4 spines on the lower
border and a terminal spine on the upper border, the claw-like dactylus closes
against a bunch of spines on the near end of the propodite.

In a female the carapace is about 27 millim. long, and about 21 miUim. wide.

1384

Andaman Sea 13° 16' 30' N. 93' 8' E. 79 fathoms. " Investigator."

Andaman Sea 13M 7' N. 93° 7' E. 90 fathoms. "Investigator."

10

2691
10

Types of the species

This species closely resembles the Atlantic and Mediterranean H. harhata,
from which indeed it differs only m the following particulars : —

The eyes are more reniform. The second spine of the lateral border of the
carapace is just behind the hepatic region, instead of being on that region.
There are spines on the posterior border of the meropodites of all four pairs of
walking legs.

Subijenus Homolax.
Eomola {Homolax) megalops Alcock. Plate. IV., fig. 21.

Homola megixlops, Alcock, Ann. Mag. Nat. Hist., May 1894, p. 408 : Illustrationa of the Zool. of the Investi-
gator, Crust., pi. xiv., figs. 1, la: Investigator Deep Sea Brachyura, p. 9 : J. A. S. B., LXVIII., pt. 2, 1899, p. 158,

Carapace urn-shaped, its greatest breadth is across the middle of the
branchial region ; its sides, and still more the spinulate lateral borders of its
dorsum, are elegantly curved ; the hairs that cover it are so inconspicuous as to
be recognizable only with a lens.

68

Rostrum a depressed dorsally-grooved tootli, entire, or emarginate at
tip. Four spines on the anterior border of the carapace arranged as in
if. andamanica.

The only enlarged spine of the lateral border stands alone on the hepatic

region.

Nine spines on the gastric region— two immediately behind the spines at
the base of the rostrum, the other seven in an open S-shaped curve across the
middle of the region.

A single row of spines on the subocular region, which region is remarkably
hollowed for the reception of the retracted eye. Two spines, one above the
other, on the carapace beside the antenna-peduncle, in addition to the blunt
suborbital tooth.

Eyes reniform, very large, their major diameter being one- sixth the breadth
of the carapace : the joint on which they are borne is as long as the slender
basal joint.

Chelipeds slender, their arms and wrists distinctly slenderer than the
meropodites of the ambulatory legs : in the adult male they do not reach half-
way along the merus of the first pair of legs : they are covered with a short
inconspicuous velvet, with hardly any long bristles on the edges of the joints:
they are armed much as in H. andamanica, but the upjyer border of the hand is
spiny and the lower border faintly denticulate. The fingers, which have a sharp
entire cutting-edge, are as long as the rest of the hand.

The ambulatory legs have the surface— especially the dorsal surface— of
most of the joints covered with a close short velvet, but have few or no bristles
alono- their edges. The 2nd and 3rd pair, which are nearly a dactylus longer
thaa°the first, are nearly three times as long as the carapace : the subcheliform
4th (last) pair reach beyond the end of the carpus of the preceding pair. The
first three pair have the anterior edge of their greatly compressed meropodite
closely spinate, and the posterior edges of that joint and the ischium closely
spinulate; their last three joints have the edges unarmed, except for a series of
fine capillary spinelets hardly distinguishable from bristles along the merus and
carpus, and a few small jointed spinules at the base of the posterior border of
the dactylus. The last pair of legs have the posterior edge of their subcylindrical
meropodite closely spinate and have only a single terminal spine on the upper
edo-e, the carpus has a strong terminal spine on its posterior border, and the
propodite has a salient group of spines behind the middle of its posterior border
forming a subcheliform stump for the serrated posterior edge of the claw-like
dactylus.

Colour in life salmon-pink.

64

Dimensions of carapace of a full-grown specimen 41 millim. long, 36 millim.
broad.

6928-6929 _ , . , „ IT-Sl' 40" N. 92° 46' 40" E

8832
9 ■

8958-8960

9

2580-2585

10

, . ^ ^ ir-3r40"N. 92°46'40"E.^„^ ^. ^ „

(Types OP THE SPECIES). Andaman Sea, ,o^ «^,^ , , >-" Investigator.

188-220 fathoms J

Off Colombo, 142^00 fathoms. " Investigator."

. Off Madras coast, 13° 51' 12" N. 80' 28' 12" E. 145-250 fathoms. " Investigator."
. Andaman Sea, 14° 13' N. 93' 40' E. 370-419 fathoms. " Investigator."

Subgenus Paromola Wood-Mason.
Homola (Paromola) profundorum, Ale. & And. Plate V., fig. 22.

Homola profundorum, Alcook and Anderson, Ann. Mag. Nat. Hist., Jan. 1899, p. 5 : Aloook, Investigator Deep
Sea Brachyura, p. 10, pi. i. fig. 2: J, A. S. B., LXVIII., pt. 2, 1899, p. 159.

Carapace very decidedly macrurous, deep, ovoid-triangular, broadest abaft
tbe middle of the branchial region, tapering to an acutely-spiniform rostrum of
which the length is about a third that of the rest of the carapace. Diverging
from either side of the base of the rostrum is a spine of similar form and size.
The only other elevations on the carapace are a hepatic spine just behind the
hollow for the retracted eye, an antennal spine just outside the antennal base,
and a blunt denticle near the middle of the ill-defined lateral border.

The gastric region is well delimited, and the linea anomurica is broad
conspicuous and dorsal.

The stout cylindrical terminal joint of the eye-stalks is longer than the
slender basal joint, the eyes are of good size, well pigmented, and hemispherical.

The chelipeds are slender but are stouter than the legs ; the arm has the
outer lower border spinate and, on the upper border, a few spinules and a strong
terminal spine ; both the inner and the outer angles of the wrist are armed
with a strong spine, the fingers are much shorter than the hand and have the
cutting-edge entire.

The ambulatory legs are slender and subcylindrical, the 2nd and 3rd pair,
which are slightly longer than the first, are at least three times the length of
the carapace. In the first 3 pair there are a few distant spines and a strong
terminal spine on the anterior border of the merus, a few articulating spinules
at the far end of the posterior border of the propodite, and a comb of articulating
spines along the posterior border of the dactylus — the last joint being but half
the length of the last but one. The dorsal fourth pair of legs are far slenderer
than the others and do not reach the end of the merus of the preceding pair :
their propodite is triangular, owing to the expansion of its posterior border, and

65

opposes a sharply-serrated edge to the less strongly toothed posterior border of
the short dactyhis— the parts being cheliform rather than subcheliform.

The body and appendages are coated with very short distant bristles which
do not conceal the surface : there are some longer and thicker bristles along the
edges of the chelipeds, and a very few scattered hairs along the edges of the legs.

The carapace is about 13 millim. long, and about 9 millim. in greatest
breadth, in a young female.

•'S74-'2S7G , rOfi Travancore coast, 430 fathoms. ),, ,. . ,,

-"4 ^-'^., (TV..BS OP THC SPECIES).!^, ^^, 3^,, ^_ ^g. ^^, 3^,, ^ j " Invest.gator.

10

Paeomolopsis, Wood-Mason.

Paromolop^i^, Wood-Mason, Ann. Mag. Nat. Hist., March 1891, p. 268 : Alcock, Investigator Deep Sea
Brachyura, p. 11, and J. A. S. B., LXVIII., pt. 3, 1899, p. 160.

Differs from Homola in the following characters : —

The carapace, which is urn-shaped, is depressed, its side- walls being far from
vertical and being overhung by the sharply-defined lateral borders. The hepatic
region is elongate and advanced, so that the hepatic spine is on a level with the
spines of the anterior border of the carapace and helps to form a very distinct
" orbit."

Distribidion : Seas of India from 300 to 600 fathoms.

Paromolopsis Boasi, "Wood-Mason. Plate V., fig. 23.

Paromolofnis Boasi, Wood-Mason, Ann. Mag. N.at. Hist., March 1891, p. 268 and fig. 5 : Alcock, U. cit.

Every part of the body and appendages, except the antennal flagella,
covered with a short close downy tomentum.

Carapace ending in a short triangular rostrum with an upturned tip ; its
greatest breadth, which is across the middle of the branchial regions, is equal
to its length without the rostrum : the lateral border is well-defined throughout,
is carinated, is co-extensive with the length of the carapace, and ends in a large
triangular hepatic spine the tip of which is on a level with the tips of the spines
of the anterior border : these are four in numl3er, one on either side of the
rostrum and one at either outer orbital angle.

There are two small antennal spines, there are some definitely-placed nodular
swellings on the well defined gastric region, and the surface of the denuded
carapace is granular, but there are no spines other than those mentioned.

The swollen terminal joint of the eyestalk is rather longer than the slender
basal joint : eyes of good size, well pigmented, hemispherical, retractile into a
very decided hollow in the front wall of the hepatic region.

9

6Q

The 2nd joint of tlie antenna-peduncle is not produced or acute at the
antero-external angle ; the antennal flagellum is much longer than the carapace.

Chelipeds (in the adult female and young male) short, just reaching beyond
the end of the carpus of the first pair of legs : the arm is slenderer than the
corresponding joint of the first three pair of legs : the fingers are longer than
the hands : none of the joints are spinate.

The second and third ambulatory legs, which are longer than the first by
their dactylus, and longer than the fourth by their merus and dactylus, are 3
times the length of the carapace. In the first three pair the anterior border of
the meropodite is armed with large spines, but the other joints are unarmed :
the dactylus is slender, curved, and of great length, being hardly shorter than
the preceding joint.

In the subchehform, dorsal, fourth (last) pair the anterior border of the
merus ends in a spine and the posterior border of the merus is spiny through-
out, the propodite is much dilated and toothed at its basal angle posteriorly, so
as to be ^shaped, and has one or two spines on the undilated portion of its
posterior border, and the dactylus is short and is toothed along the posterior
border.

The carapace of an adult female is 45 millim. long and 43"5 millim. broad.

The colours in life vary from red to bluish-pink.

3735,^ . f Bay of Bengal, about 30 miles W. of Middle >, ^ ^. ^ „

—• (^"^ '' '^' ^^^"^^>- 1 Andaman I. 480-500 fathoms. I Investigator.

3178

'—^. G. of Manar, 6° 29' N., 79° 34' E. 597 fathoms. " Investigator."

g- . B. of Bengal, 13' 47' 30" N., 92° 36' E. 561 fathoms. " Investigator."

107 787-789
10 ' 10 *

2378

10 ■

Arabian Sea, 9° 34' 57" N., 75" 36' 30" B. 406 fathoms. " Investigator."

Andaman Sea, 13° 50' 30" N., 93° 26' E. 498 fathoms. " Investigator."

2388

-^. Arabian Sea, 7° 17' 30" N., 76° 54' 30" E. 480 fathoms. " Investigator."

3743

-^. Arabian Sea, 10° 8' 43" N., 75° 33' 30" E. 295-360 fathoms, "Investigator.".

Htpsophrts, Wood-Mason.

Eypsophrys, Wood-Mason, Ann. Mag. Nat. Hist., March 1891, p. 269; Alcook, Investigator Deep Sea
Brachynra, p. 12, and J. A. S. B., LXVIII., pt. 2, 1899, p. 162.

Carapace deep, longer than broad, quadrilateral or ovate-oblong, with deep
vertical parallel sides, the gastric region well delimited and occupying its anterior
half, the linea anomurica dorsal, distinct or indistinct.

\ %

67

Front narrow, forming a simple or bifid rostrum wliicli has a spine on either
side of its base.

The orbits do not afford any concealment to the eyes, but form, on either
side of the rostrum, a broad concave facet sharply marked off from the rest of
the carapace by a ridge that arches round dorsally from the rostrum to the
antennal spine : at the upper and inner angle of this facet is a well defined
hollow that catches the knee of the 2nd and 3rd joints of the antennulary
peduncle when flexed. The eyes are well formed : the terminal joint of the eye-
stalk is barrel-shaped much as in Homola, but the slender basal joint is short or
obsolescent, so that the eyes do not appreciably project beyond the edge of the
orbital facet. •

The mouth-parts are very like those of Homola, but as the outer border of
the merus of the external maxillipeds is hardly at all expanded these appendages
are even more pediform than in Homola.

Chelipeds slender, spiny, equal. Legs of the first three pair long, with
broad compressed meropodites. Fourth (last) pair of legs short, very slender,
cheliform, their dactylus, which is many times shorter than their propodite,
shuttincr down against and co-termiuous with the slightly expanded distal end
of the propodite.

The abdomen of both sexes consists of seven separate segments.

Distrihufion: Arabian Sea and Bay of Bengal between 400 and 1000
fathoms.

Eeij to the species of Hypsophrys.

I. Rostrum simple: gastric region veiTUCOse ... ... ••• H. supercihosa.

II. Rostrum bifid: gastric region spiuose ... ••. ••■ H. longipes.

Hiipsophrijs supercUiosa, Wood-Mason. Plate VI., fig. 2-i.

E,jvsovhrv. suferciUosa, Wood-Masoa. Aan. Mag. Nat. Hist., March 1891, p. 263: 111. Zool. Investigator,
Crast., pi. xiv., figs. 4, ia : Aloock II. cit.

Rostrum simply pointed. Linea anomurica rather indistinct.

Four small spines or teeth on the anterior (orbital) border of the carapace,
two being far apart at the base of the rostrum and one at either outer orbital
ano-le. Two, or all four, of these teeth may be obsolescent or obsolete.

Lateral borders of dorsum of carapace not defined, except by a single
isolated spine on the hepatic region. Gastric region sharply subdivided into
three subregions, of which the lateral are somewhat nodular. Two or three
spines on 'the subhepatic and suborbital region, the innermost of which is
" antennal," also sometimes a few spinules.

68

Eyes well formed and facetted, but pale. Antennal fiagella about half again
as long as the carapace.

The pediform external maxillipeds have their surfaces and edges devoid of
spines.

Chelipeds slender, but much more massive than the ambulatory legs, about
half a hand-length shorter than the first pair of ambulatory legs in the adult
male : spines and spinules in rows on edges and on both inner and outer surfaces
of arms, wrists and hands : fingers about three-fourths the length of the palm.

The second pair of legs, which are slightly longer than the first and third
and considerably more than twice the length of the fourth, are slightly more
than three times the length of the carapace. In the first three pair the
meropodites are compressed, with the anterior border spiny and the posterior
border much less strongly and profusely spiny ; the other joints are slender
and unarmed, except for a few articulating spinelets at the far end of the posterior
border of the propodite and in the basal half of the posterior border of the
dactylus ; the dactylus is slightly shorter than the propodite.

The fourth (last) pair are very slender and are unarmed except at their
cheliform ending : their propodite is many times longer than the dactylus.

The terminal joint of the male abdomen is bluntly triangular.

There are some soft bristles on the chelipeds, and a few on the legs, and
some very short and inconspicuous hairs on the carapace.

Colours in life, pink ; fingers blackish.

The carapace of a large egg-laden female is 19 millim. long and 15 millim.
broad.

6225-6226 (Types of the"1
9 species). [

6227-6228 [

9 J

8523-8532

Arabian Sea, 15' 2' N., 72' 3i' E. 740 fathoms. " Investigator."

y

69-87
10 ■

88-91
10 ■

92
lO"

93
10'

3742

1o~'

Arabian Sea, 12° 5' 35" N., 71° 35' 50" E. 805-880 fatboms. " Investigator."

Arabian Sea, 15° 11' N., 72" 28' 45" E. 912-931 fathoms. "Investigator."

Arabian Sea, IS" 47' N., 72° 3' 45" E. 891 fathoms. "Investigator."

Arabian Sea, 22° 14' 25" N., 67° 8' 55" E. 947 fathoms. " Investigator."

Arabian Sea, 23° 8' 22" N., 65° 49' 45" E. 890 fathoms. " Investigator."

Bay of Bengal, 10° 50' 30" N., 80° 44' 30" E. 987-900 fathoms. " Investigator.','

69
Eyiisophrys longijoes, Alcock & Anderson. Plate VI., fig. 25.

Hypsophrys longipes, Alcock and Anderson, Ann. Hag. Nat. Hist., Jan. 1899, p. 6 : Alcock, Investigator Deep
Sea Brachyura, p. 15, pi. i., fig. 1 : J. A. S. B., LXVIII., pt. 2, 1899, p. 164.

Rostrum deeply bifid. Linea anomurica distinct.

Four large spines on tlie anterior border of tlie carapace — two close
together at the base of the rostrum, one at cither orbital angle.

Lateral borders of dorsum of carapace well defined, spinulate ; the ridge
on the side-wall of the carapace that defines the branchial regions anteriorly is
also spinulate. A row of spines on the hepatic region, the largest of which is
on the lateral border of the carapace and has a spine dorsad of it.

Gastric region obscurely subdivided, each lateral subregion is armed with 5
or 6 large spines, while on the median region there is a central spine sometimes
followed by a row of spinules. Subhepatic and suborbital region with numerous
large spines, one of which is " antennal."

Eyes well pigmented. Antennal flagella more than twice the length of the
carapace.

Rows of spinules on the exposed surface of the ischium merus and exopo-
dite of the external maxillipeds, and a row on the basal joint of the antennules.

Chelipeds slender, reaching not far beyond the end of the carpus of the
first pair of ambulatory legs, the arm and wrist not stouter than the meropodites
of the first 3 pair of legs ; spinate and spinulate as in the preceding species ;
fingers as long as the hand.

The second and third pair of legs, which are slightly longer than the first
and three times as long as the fourth, are four times the length of the carapace.
In the first three pair of legs the merus is compressed and has its anterior
border spinate and its posterior borders spinulate, the jiosterior border of the
propodite carries a few distant articulating spinelets, and the dactylus — which
is about two-thirds the length of the preceding joint — has a close comb of arti-
culating spines along its posterior border.

The fourth (dorsal) pair, which are extremely slender, have the posterior
border of the merus strongly spinate : the propodite is several times longer
than the minute dactylus.

The terminal joint of the male abdomen ends acutely.

Hairs and bristles are sparsely present just as in the preceding species.

The carapace of a large egg-laden female is 38 millim. long and 30 millim.

broad.
23G5-2369

(Types of the species).

1

Arabian Sea, 7° 17' 30" N., 76° 54' I ^ .. . ,
- , „ , . r Investigator.

2887 j 30" E. 430 fathoms. | °

10
2887

lo" L» J

70

Family Latreillid^, Alcock.

Latreillidae, Alcook, Journ. Asiatic Soo. Bengal, LXVIII., pt. 2, 1899, p. 130.

Carapace either elongate-quadrangular or piriform. Antennal flagella
shorter than the carapace. The slender basal joint of the eyestalk is very much
longer than the terminal joint. The external maxiUipeds are suboperculiform.
The gills are 8 on either side and there are no epipodites on the chelipeds or
ambulatory legs.

The branchial formula is as follows : —

isomites ana tnei:
Appendages.

Podobranchia).

Artlirobranchiae.

Pleurobranchias.

VII

ep.

0

0

=

ep.

VIII

1 + ep.

0

0

=

1 + ep.

ix

ep.

2

0

r=

2 + ep.

X

0

2

0

=

2

XI

0

0

1

==

1

XII

0

0

1

=

1

XIII

0

0

1

=

1

XIV

0

0

0

=

0

Total

1 + 3 ep.

= 8 + .3 ep.

Key to the genera of Latreillidge.

I. Carapace subquadrilateral. Antennas long. All seven abdominal
segments distinct in both sexes

II. Carapace piriform, its anterior poi-tion forming a long snbcylindrical
" neck." Anteunse short. The 4th, 5th and 6th abdominal seg-
ments of the female are fused together ...

Latreillopsis,

Latreillia.

Latreillia, Roux.

Latreillia, Rons, Crnst. Medit. pi. xxii. and text. Milne Edwards (79), DeHaan (23), Heller (40), Henderson (42),
A. Milne Edwards and Bonvier (76), Bouvier (12), Ortmann (84), Alcock (3).

For referencea see J. A. S. B., LXVIII., pt. 2, 1899, p. 167.

Carapace elongate-piriform, not covering the basal joints of the legs, its
anterior part prolonged to form a subcylindrical " neck " at the end of which
are the spiniform rostrum (lying deflexed between two long slender divergent
" supra-ocular " spines) the eyes, the antennules, and the antennse. The regions
are fairly well indicated, and there is no linea anomurica.

The eyes are large, and the slender basal joint of the eyestalk is of extra-
ordinary length, being several times longer than the terminal joint. Antenna?
short, filiform.

Epistome of great length fore and aft, corresponding with the " neck " of

71

the carapace. Buccal cavern well demarcated from the epistome, the efferent
branchial channels well defined. External maxillipeds not completely closmg
the buccal orifice : they have a pediform cast, the ischium and merus bemg
rather narrow and the flagellum coarse.

Chelipeds long and slender, but always much shorter than the first three
pairs of ambulatory legs: all the joints are slender, except the palm, which m
one or both sexes is club-shaped. Fingers shorter than the palm.

First three pair of ambulatory legs very long and slender ; some of their

joints are spiny.

Fourth (last) pair of legs more or less reduced in length, subdorsal in

position.

The abdomen of the male consists of seven separate segments ; that of the
female consists of five segments-the 4th, 5th and 6th being fused together.

Distribution: Atlantic coasts of North America between 38° and 40° N. :
off the Canaries and Azores : Mediterranean Sea : Bay of Bengal and Andaman
Sea : Japanese Seas : New South Wales coast.

Latreillia jjennifera, Alcock. Plate VII., fig. 27.

Latreillia pennifera, Alcock, Journal Asiatic Soo. Bengal, LXVIII., pt. 2, 1899, p. 168.

Very closely related to L. elegans, Koux.
- Carapace smooth, without spines, though the hepatic regions have a strong
bulge : the "neck" is rather slender (equally so in both sexes) and is nearly as
long as the rest of the carapace measured in the middle line.

°Kostral spine short, acute, strongly deflexed. Supraocular spines as long
as the eyestalks, about half the total length of the carapace ("neck" included)
measured in the middle Une; occasionally bearing some tiny secondary spmules.

Antennules slightly longer than the eyestalks: the outer flagellum longer
and very much coarser than the inner.

The chelipeds, which are slightly longer in the male than in the female, are
between 3^ and 4 times the total length of the carapace: their joints are long,
slender, and cylindrical, except the palm of the male, which is club-shaped :
there are a few spines on the arm, but the other joints are smooth : the fingers
are not half the length of the palm.

The first 8 pair of ambulatory legs, though they increase slightly in length
from before backwards, are not very dissimilar in length, the first pair being
nearly 8 times the total length of the carapace. All their joints are slender :
the merus is spinate, the carpus sparsely spinate, and the propodite is shghtly
dilated at the far end of the posterior border where there are a few spmes.

72

The last pair of legs are between 4^ and 5 times the total length of the
carapace and reach almost to — in the female even beyond — the end of the carpus
of the last pair but one : the merus is rather sparsely spinate, chiefly on the
posterior border, and the propodite is plumed on both sides so as to exactly
resemble the vane of a feather : the dactylus is extremely short.

In both sexes the last abdominal segment is shaped like a spearhead : in
the female the 2nd and 3rd abdominal terga have a median spine and the 4th
has a spine at the proximal end of either lateral border.

Colours in spirit yellow. In life the carapace is reddish with longitudinal
stripes of dark red, the eyestalks chelipeds and legs are closely cross-banded
with red, and the eyes are purplish black.

The carapace of an adult female, with eggs, is 11 millim. long.

i-ip . Bay of Bengal, S° 51' 30" N., 81° 11' 52" E. 28 fathoms. " Investigator."

1713-1718

lo

3538 ,_, , 67 fathoms.

(Type of the species). I

1

G. of Martaban, 14' 26' N., 96" 23' E. I ^ .

f- ' Investigator.

JO ^ L J

OQQf

' ' ■■ Ofi Torres Is., Mergui Archipelago. 40 fathoms. " Investigator.'

Latreilloims, Henderson.

Latreillopsis, Henderson, Challenger Anomura, p. 21 ; Ortmann in Bronu's Thier Reich, Malacostraoa, p. 1156 :
Aloook, J. A. S. B., LXVIIl., pt. 2, 1899, p. 165.

Carapace subquadrilateral, deepish, with vertical side-walls, not entirely
concealing the basal joints of the legs : the regions fairly well indicated. Front
of moderate width, ending in a spiniform rostrum on either side of which is a
long slender divergent supraocular spine. Linea anomurica present, most dis-
tinct posteriorly.

Eyes as in Latreillla, large and borne free at the end of slender eyestalks of
remarkable length. Antennaa long, the peduncle slender and cylindrical.

Epistome well demarcated from the palate. Buccal cavern much broader in
front than behind, the efferent Ijranchial channels very well defined. Though
the external maxillipeds do not quite meet across the buccal cavern they are dis-
tinctly operculiform, owing to the expansion of their merus.

Chelipeds long and slender but much shorter than the first 3 pairs of ambu-
latory legs : their joints, like those of the legs, are cylindrical, and the palm in
the male is enlarged and club-shaped.

Ambulatory legs slender, the first 3 pairs very long ; the fourth pair reduced
in length, and subchelate.

73

The abdomen in both sexes consists of 7 separate segments.

DL^trihution : Oriental Seas at moderate depths up to nearly 100 fathoms.

Latreillopsis hisphivm, Henderson. Plate YII., fig. 26.

L'ltreiUopsis hispinosa, Uenderson, Cluilkfiiger Anomura, p. 22, pi. 11, fig. 3, J : Aloock, J. A. S. B., LXVIII.,
pt. 2 , 1890, p. 166.

Carapace longer than broad, shaped much as in Homola : frontal region with
three sharp slender spines, the middle one — which is the shortest and is slightly
deflexed — being the rostrum, the other two — which are about a third the length
of the carapace and are slightly up-tilted — being placed above the bases of the
eyestalks. Gastric region tumid, with a tubercle posteriorly and a curved
transverse row of tiny tubercles anteriorly. Cardiac region small, tumid,
culminating in two tubercles placed side by side or confiueut. Branchial regions
with an irregular surface, and with one or two tiny spinules on the side wall.
Hepatic regions standing out like a pair of little wings, with two spines — the
foremost of which is nearly as long as the rostrum — projecting obliquely for-
wards from their prominent outer angle, and with one or two small spinules on
their under surface. Eyestalks nearly as long as the supraocular spines.
Antennal peduncle about as long as the eyes and eyestalks combined, the
flagellum more than three-fourths the length of the carapace.

Chelipeds and legs slender, cyhndrical, practically smooth, except for a spine
at the far end of the anterior (extensor) border of the merus. The chelipeds in
the male are just over twice, in the female less than twice, the length of the
carapace without the rostrum. In the female they are hardly stouter than the
legs ; but in the male they are distinctly stouter, especially as regards the palm,
which is club-shaped : the palm is much longer than the fingers.

The first three pairs of ambulatory legs increase in length, gradually but
slightly, from before backwards, the 3rd pair being between 4 and 4- times the
leno'th of the carapace : the dactyli are long and curved. The fourth (last) pair
of legs are a little longer than the male chelipeds : their last two joints are short,
and the dactylus folds down, like a knife-blade, on a double row of spines along
the posterior border of the propodite.

In both sexes the last abdominal tergum is shaped like a spearhead, and the
2nd, 3rd, 4th and Gth terga have an acute tubercle in the middle line.

The carapace of an egg-laden female is 8 millim. long.

Colours in spirit yellow, the fingers and eyes dark brown.
32S4-3285 3494

lu 10

10

Off the Audamaus, 53 fathoms. " luvestiVator.'

74

TABLE OF THE GENERA AND SPECIES OF BROMIDES.

The names of the Indian genera and species are printed in small caiiitals.

DROMIIDEA.

Family Homolouhomid^.

Ball. Mus. Comp. Zool. VIII. 1880, p. 3i

Somolodromia

A. Milne
Edwards.

„ paradoxa, I Bull. Mus. Comp. Zool. he. cit. and Rec. Fig. ^

A Milne ) Crust. Nouv. ph 39, tig. 2: Bouvier, Bull. Soc. ^ Caribbean .Sea.
Edwards. ( Philom. Paris (8) VIII. 1895-96. )

Dicranodromia ( Bull. Mus. Comp. Zool. VIII. 1880, p. 31 : A. Milne

A.Milne ^ Ed\vard,s and Bouvier, Crust. Decap. Hirondelle et
Edwai'ds. (. Princesse Alice, Monaco, 1899, p. 11.

" T'MilnP ^ Loc.cU.v 32: Bouvier, Bull. Soc. Philom. (S) j c.nbl,..n Sea.
J} ■, ; VIII. 1895-96. (

Edwards. (. ^

„,,.. ( Zool. Jahrb., Syst. VI. 1892,p.549,pl. xxvi.fig. 4:)
„ Boderleini, \-q^^^.^ 5^11. Qoo. Philom. Paris (8) VIII. 1895- [ Japan Sea.
(Jrtmann. ) or )

f Rec. Fig. Crust. Nouv. pi. 10: Bouvier, Bull. Soc.l
,^ , . I Philom. fS) VIII. 1895-96, p. 50: A. Milne Ed- ]
„ Maylimxi, \ ^^^^.^^ ^^^ Bouvier, Crust. Decap. Hirondelle et ! E. Atlantic (Cana
A. Milne "i, Princesse Alice, Monaco, 1899, p. 14, and Exp. Sci. f ries and Azores).
Edwards, m .>, , rr.ii n t t-i.;.,„„ t3+ t ^ it I

I Travailleui' et Talisman, Crust. Decap. Pt. I. p. 14, |
l^pl. iii. fig. 4, pi. ix, figs. 1-11. J

Aeachnodkomia r

Alcock. \

„ Baffini,

Alcock and
Anderson.

Ante,

Ante.

Indian Seas.

Dynomene Latreille.

Family DYNOMENii'.i:.
Ante.

f Consid. Gen. Crust, p. 133, pi. xviii. fig.-2 : Guerin,^

ilcon. Regno Anim. pi. xiv. fig. 2 : Milne Edwards
in Cuvier, Regue Anim. \A. xl. tig. 2 : and Hist. Nat.
Crust. II. p. 180: A. Milne Edwards, Ann. Sci.
hisfida, I Nat. (6) VIII. 1879, Art. 3, p. 5, pL xii, ph xiii.

Desmarest. 1 figs. 1-15: de Man, Archiv f. Naturges. LIII.
1887, i. p. 408: Ortmann, Zool. Jahrb., Syst., VI.
1892, p. 543. l)ynoinene latreilli, Eydoux and
Souleyet, Voy. Bouite, Zool. I. p. 239, pi. iii. figs
L3-5.

r Ann. Lye. Nat. Hist. New York VII. 1862, p.
Ursula, J 239 . ^ ^iiug Edwards, Ann. Sci. Nat., Zool ""

ludo-Pacific (Jlauri-
tius to Sandwich
Is.).

J

Stimpson.^yjjj_ jgyg^ ^^.j._ 3_ p 9^ p]_ ^- ggg i^j.ig.

2, P.^

California.

75

f Aun. Sci. Nat., Zool., (G) VIII. 1879, Ait. 3, p. 8,^
DYNOMENEpr^-Za^or, )pl. xiv. fig.s. 20-26 : Micrs P. Z. S. 1884 p 13 | j^^^^^p^^^jg g^^^^

) Man, Arcliiv f. ^aturges. LTII. 1887. i. J- , ,. H<,n.,«n«r \

A. Milne -{ (note) : de

Edwards. | p. 409: Ortmaun, Zool. Jalu-b., Sy.st., VI. 1892, p.
L543, pi. xxvi. fi-. 3. J

to Madagascar. )

f C. R. de la Soc. Pliiloiii. Paris, 1894-9.5, p. G, and"!

„.„ ,. I Bull. Soc. Pliilom. (S) VIII. 189.5-96, pp. 58, 59, -r. /,,,^.-,r
FiUioh. ,. ->-. .T., -vi-i 1711 ] 3 ij> ■ T? o • I >^- Atlantic (Ij.

' <{ figs. 22, 23: Jihlne Edwards and Bouvier, Exp. bci. }-

Bouvier.

I Travaillour et Talisman, Crust. Decap. Pt. T, p. 5,
l^pl. iii. pi. viii. figs. 1-18. J

Verde Is. &c.).

,, pngnnfrlx,
de Man.

,, FILUMNOIDES,

Alcock.

ACANTHODROMIA

A. Milne
Edwards.

„ ermivea,
A. Milne
Edwards.

,, .MAU(,;4H.1TA,

Alcock.

Zoiil. .Tahi-b., Syst., IV. 1889, p. 444, pi. x. tig. 13. Mauritius.

Ante. Arabian Sea.

Arite.

Bull.
Bouviei'
pp. 56,

Mas. Comp. Znol. VIII. 1880, p. 31
, Bull. Soc. Pliilom. (8) VIII. 1895-96, } Caribbean Sea.
57. Bgs. 18-21.

Ante.

Andaman Sea.

Dromia Fabr.

„ RuMi'Hii, Fabr.

Family Dromudj:.

Ante.

Ante.

C Milne Edwards, Hist. Nat. Crust. II. 173, pi. xxi. ^
figs. 5-8: Bell, Brit. Stalk-eyed Crust, p. 369:
Heller, Crust. Siidl. Europ. p. 145, pi. iv. figs.
10-1 J : Broccbi, Ann. Sci. Nat., Zool., (6) II, 1875,
Art. 2, p. 106 (male appendages) : Carrin^ton
and Lovett, Zoologist (3) VI. J882, p. ^259
(habits): Cams, Pi-odr. Faun. Medit. I. p. 498:

E. Africa to Japan.

E. Atlantic from the
English Channel

„ vuhjai-if:, Edw. ■{ Ortmaun, Zool. Jaliib., Syst., VI. 1892, p. 547 : }- to the W. coast of

latar, Fdw.

Bouvier, Bull. Soc. Philom. i 8) VIII. 1895 : Rath-
bun. P. U. S. Nat. Mus. XXII. 1900, p. 300:
A. Alilne Edwards and Bouvier, Exp. Sci. Travail-
leur et Talisman, Crust. Decap. i)t. I. p. 17, pi. ix.
fig. 15. (Broiiiia medilerranea Leach, Malac. Pod.
Brit. pi. xxivd : Dromia cmnmunis, Lucas, Hist.
^Nat. Anim. Artie, in Fxpl. Sci. Alg^rie, pt. I. p. 20).

r Milne Edwards, Hist. Nat. Crust
J en/thropus, Ratlibuu, Ann. I
(1897, p. 39).

Africa,
rauean.

Mediter-

. Crust. H. 174 {D.'^
nst. Jamaica, No. I. >

nodosa,
A. M. Edw.
& Bouvier.

Exp. Sci. Travailleur et Talisman. Ciust. Decap.'
pt. I. p. 18, pi. ix. figs. 12-14. C

ciliata, r C'hallenir,., Anonnirn, p. 3, pi. L fi-. 1.

Henderson. ^^ '

Milne Edwards, Hist. Nat. Crust. II. 175

W. Indies.

C. Verde Is.

Australia.
Loc. ?

,, gibbosa, Edw.

„ octodentafa, f P.L.S., N. S. Wales, VI. 1S81-82, p. 7.55 and Cat.^^^ Antvalio
Haswell.l Austral. Crust, p. 140. J^' ^"^^'^''^

16

Dromia fulvo- f A.m. Mag. Nat. Hist. (5) VIII. 1881, p. 270, pl.\^y_ j^^^.-^.^_

hispida, Miers. \xvi. fig. 1. J

spinirostris, f Ann. Mag. Nat. Hist. (5) VIII. 1881, p. 271, pl-\^-. Africa.
Miers. \xvi. fig. 2. J

w, f Trans. Roy. Soc. S. Australia, X. 1886-87, p. 299,\g ^^^fj.^u^
z. \pl- xiv. figs. 6-6. J '

hicavernosa,
Zietz.

Stimpson. r Proc. Acad. Nat. Sci. Pliilad. 1858, p. 225 ; Hender-
^eniis of < son. Challenger Aiiomura, p. 12. (Dromidiopsis,

Dromidia
(subge
Dromia). (L. A. Borradaile, P. Z. S., 1900, p. 672).

/ Milne Edwards, Hist. Nat. Crust. II. 176: Dana,^
hirtissima, ) U. S. Expl. Exp. Crust, pt. I. p. 403 : Brocclii, Ann. f S. Africa and Saud-
'' Sci. Nat. Zo "
appendages).

Lamarck, j Sci. Nat. Zool. (,6) II. 1875, Art. 2, p. 105 {male ( wicli Is
I appendages). )

, , r Milne Edwards, Hist. Nat. Crust. II. 177 : de Man,^

giobosa, J ^r^,,,i.p f Naturges. LIII. i. 1887, p. 396, pi. xviii. yindo-Pacific:

Lamarck, ^j^^ ^ . Bgi-radaile' P. Z. S. 1900, p. 671. J

TOIDENTATA, f ' , .^ \Africa to Malay

Riippell.\ J Archipelago.

rotunda f Smith's lllustr. of S. Africa, Annulosa. p. 71. Ts. Afrlcn.

Macleay.\_ J

INDICA, Gray. Zool. Miscell., p. 40. " Indian Ocean."

^''^^d ^ M**' ! (probably identical with D. indica Gray. ) ^Andaman Sea.

, Milne Edwards, Hist. Nat. Crust. II. 178 : de Man,"^
caput-mortu- I ^^.^.j^^^ ^._ jsfaturges. LIII. 1887, i. p. 393, pi. xvii. Undian Ocean.

urn, Latr. i „ ,

„ spongiosa, f Proc. Ac. Nat. Sci. Pliilad. 1858, p. 238 : Henderson, \g ^^^.-^^
Stimpson.^ Challenger Anomura, p. 12. J

var. r
III, <
liers. l^

stimpsonii, -l Zool. H. M. S. Alert, pp. 519, 552, pi. L. fig. A. >Mozambique.
Miers. l^ J

„ excavata, f Proc. Ac. Nat. Sci. Philad.11858, p. 239 : Haswell,\^^g^^,^j.,^_
Stimpson. \ Cat. Austral. Crust, p. 140. J

f P. L. S., N. S. Wales, VI. 1881-82, p. 755, and Cat. ^
^ ,. . I Austral. Crust, p. 139 : de Man, Archiv f. Naturg. J y , „ .^ „
" ^^^stral^ens^s )^ ^111. 1887, i. p 396, pi. xvii. fig, 6: ? Henderson, ^"^"-P^^^fi^ ^^^
HaswelL] ^^^^^_ Linn. Soc. Zool. (2) V. 1893, p. 406: Borra- ( ^^^^°" *° ^'l'"
Ldaile, P. Z. S., liOO, p. 572. /

var. de Alan, Zool. .lahrb., Syst., IX. 1896, p. 372. Celebes.

„ orientalis, /Ann. Mag. Nat. Hist. (5) V. 1880, p. 370, i>l. xv.\" Indo- Malayan
Miers. \ figs. 1, 2. J Seas."

„ hicornis, f Abh. Ak. Berlin, 1882 (1888), p. 20, pi. i. fig. 9\S. of Cape of Good
Studer. \a, fc: Henderson, Challenger Anomura, p. 13. J Hope.

„ spinosa, /Abh. Ak. Berlin. 1882 (1883), p. 22, pi. i. fig. 10 a.\H. of Cape ot Good

Studer. \ b. J Hope.

f Ann. Lye. Nat. Hist. New York, VII. 1862, p. 71:)
„ antiUensis, J Henderson, Challenger Anomura, p. 12, pi. i. ^ G. of Mexico: Brazil.
Stimpson.-^^ fig. 5. 3

CEyPTODROMlA Ante (=Epidromia, Kossmau).

Stirapson.

r Proc. Acad. Nat. Sci. Philad., 1858, p. 239 : de Man, )
„ coronata, \ ^j,^.,,;^ j- jjaturg. LIII. 1887, i. p. 39H, pi. xviii. [ Bonin Is. : Moluccas,
istimpson. ^jj ._,. ortman.i, Zuol. Jahrb. Syst. VI. 1892, p. 5i3, )

Crtptodkomia

Zool. Miscell., p. 40: Stiinpsoii, Proc. Ac. Nat. Sci.
Philad., 1858, p. 239: Heller, Novara Ciusfc., p. 71:
Jliers, Cat. Crust. New Zeiiland, p. 57: Haswell,
Cat. Austral. Crust., p. 139 : Miers, Zool. H. M. S.

Australia, New Zea-

lateralis, ■{ Alert, p. 259 : Henderson, Challenger Anomui-a, f- land : Japanese

(Gray). I p. 5: Cano, Boll. Soc. Nat. Napol. III. 1889, p. and Cliina Seas.
255: Thomson, Trans. New Zealand Inst., 1898,
I p. 170, pi. XX. figs 1, 2: Whitelegge, Mem. Aus-
Ltral. Mns. IV. pt. 2, 1900, p. 163. J

,. arnboinensiii, Archiv f. Naturg. LIII. 1887, i. ji. -106, pi. xvii. Moluccas,

de Man. fig. 4.

,, iaponim, C Challenger Anonaura, lb8S, p. 6, pi. i. fig. 2:"\ t , ^^ ,. ,.

■^ fr J -< Tin-i -i 1 Tvf \ i 1 Ai TTT 100-7 iln >-Japan and Funa-tuti.

Heuderson.\ Whitelegge, Mem. Au.stral. Mus. 111. 1897, p. 140. J '

,, stearnsii, ( Proc. Ac. Nat. Sci. Philad., 1891, p. 216, pi. xii-\T

Ives. I figs. 1-3. J

„ fallax, Milne Edwards, Hist. Nat. Crust. II. 176. Mauritius.

(Lamk. ).

„ tuhercnlata, ( Proc. Ac. Nat. Sci. Philad., 1858, p. 239: de Man, 7 Cliina and E. Indian
Stimpson. | Archiv f. Naturges. LIII. 1887, i. p. 40 J. ) Seas.

„ I'lLEiFEUA Ante. Andaman Sea.

Alcock.

„ tumida, ( Proc. Ac. Nat. Sci. Philad., 1858, p. 240 : Ortmann, \ China and E. Indian

Stimpson. { Zool. Jahrb. Syst. VI. 1892. p. ,544. | Seas.

„ sp. Miers. Proc. Zool. Soc, 1879, p. 41. Japanese Seas.

„ CANALici'LATA, ( =Bromui iomeutosa, HeWev) Ante. Red Sea to Japan.

Stimpson.

„ PENTAGONALis, { = / canitUcidata, Ht.) Ante. Red Sea, India.

Hilgendorf.

„ EULLiFEKA, Ante. Indian Seas.

Alcock.

,, yrnitulata f Archiv f. Naturg. XLIV. 1878, p. 256 and Reisc\p i o
(Kossman).\ roth. Meer., Crust., p. (ly. J

„ lamellata, f Zool. Forschungsr. Malii_y. Arch, (in Jena. Denk-'lrp q,

Ortmann.\schr. VIIL), 1894, p. 34, pi ii. fig. 8. j^iorres fetr.

„ HILGENDORFI, Ante. Indian Seas,

de Man.

„ incisa. Challenger Anoniura, 18^8, p. Ri, pi. i. fig. 4. Australia.

Henderson.

„ nndi}"-s, Milne Edwards, Hist. Nat. Crust. II. J77. Loc. ?

(Lamk.).

„ sculpla, P. L. S., N. S. W. VI. 1881-82, p. 756, and Cat. .Australia.

Haswell. Austral. Crust , p. 141.

„ nodulifera, Cliallengei' Anoniura, 1888, p. 8, pi. i. fig. 3. Australia.

Henderson.

„ GiLESii, Ante. Indian Seas.

Alcock.

„ EiiALioiiiES, Ante. Indian Seas.

Alcock.

Eiidromia Cliallengei- Aiiomiira, p. 13, 1888.

Henderson.

„ frontalis; Op. cit., {). 11, pi. i. fig. 7. S. Africa.

Hendcr.son.

78

Petalomeka ^^,^^^

JscJiiipson.

„ granulahi. Proc. Ac. Nat. Sci. Philail., 1858, p, 240. Jajianese Seas.

Stimpsoii.

„ pulchm, Zool. H. M. S. Alert, p. 260, pi. xxvii. fii^. A. Torres Straits.

Mieis.

IKLlICA,

Alcock.

Aule. Indian Seas.

T. r Proc. Ac. Nat. Sci. Pliilad., 1858, i>. 226 : Heiider-

PsEUDODUOiiiA I challeno-er Anomiu-a, p. l'> : Stel.biu-, S African

ht„np.son. ^g^.^,,,t,_ |,, 24.

r Proc. Ac. Nat. Sci. Pliilad., 1S58, p. 240: Hender-"^

" '",;"*' ■{ son, Clialleno-er Anomura, p 16, pi. i. fay. 8 : Stebb- VS. Africa.

St.mpson. y^^^^ g_ ^^f^.^- ,_ c^,^^^^ j,_ 24. J

„ INTEGRIFUONS, r Trans. Linn. Soc. Zool. (2) V. 1893, p. 406, pl-\g i^jia
Heudei'soii.\xxxviii. tijifs. 7-9. J

Lasiodkomia

Alcock. {— Hoiiiiih'drdviia, Jliers nee Edw.). Ante.

,, copprNGRKi, Ante. Indian Ocean.

Miei-s.

CONCHOECETES Ailte.

Stimpson.

,, AHTinciosos, Ante. Indo-Pacific.

Pabr.

,, ANDAMANICUs, Ante. Andamans.

Alcock.

r Rev, et Mag. Zool. (2) VI, 1854, p. 833 : Stimp-^

Hypoconcha Guerin- 'son, Proc. Ac. Nat. Sci. Philad., 1858, p. 226: Heu- f

Menevillo. ") der.son, Clialleuger Anomui-a, p. 17 : Bouvier, Bull. C

(.Mas. d'Hist. Nat., Pari.s, 189S, p. 374. )

r Krabbeii und Krebse, III. i. 57, pi xlviii. figs. 2,^
„ sahulosa, \ 3 : Guerin-Meneville, Rev. et Mag. Zool. (2) VI. ( -^ Indies

Herbst., 1 1854, p. 333 pi. v : Stimpson I. c. : Henderson, I.e. : ( '
V, Bouvier I.e. J

„ digneti. Bull. Mus. d'Hist. Nat., Paris, 1898, p. 376. California.

Bouvier.

„ californiensis, Bull. Mus. d'Hi.st. Nat., Paris, 1898, p. 375. Califoi'uia.

Bouvier.

,, iianamensis, f Ameiican Naturalist III. 1869, p. 249 : Bouvier, \p

S. 1. Snuth.\Bull. Mus. d'Hist. Nat., Paris, 1S9S, p. 375. j-f anaraa.

r Ann. Lye. Nat. Hist. N. York, VII. 1862, p 72 :^
„ arcnaia, ) Kingsley, ' Proc. Acad. Nat. Sci. Pliilad, 1879, r W. Indies, Florida,

Stimpson. ) p. 404; Bouvier, Bull. Mus. d'Hist. Nat., Paris, C &c.
C 1898, p. 375. ;

Sphjirodrohia Ante.

Alcock.

„ KENDALLi, Alc. A>ife. B. of Bengal.

& Anders.

,, Nux, Alcock. Ante. G. of Martaban.

\_AsctdiopMlus, Richters, iu Miibius Meeresfauna Mauritius u. Seycliellen, p. 158, is said to
belong to the Bromidm. One species, Ascidiophilus caphyrseformis.']

HO. M GLIDE A.

HoMOLA Leaeli.

Iiarhnfa
(F.abr.

■{

vigil, A.
Milne
Ecbvaids

orientalis,
Henderson,

ANDAMANICA,

Alcock.

Family Ho.MOi.iDj;.

Ante.

f Fabi'icius, Ent. Sj-st. emend. II. 4.30 : Herb.st,^
Krablieii und Krebse II. iv. 1C6, pi. xlii. fij?. 3 :
Latreille, Nouv. Diet. d'Hist. Nat. XV. 1817, p.
281 : S. I. Smitli, P. U. S. N. M. ]883, p. 24, and
Albatross Crust., p. 7 in Rep. U. S. Fish. Comm.
1882 (1884) and 1881. (1886), p. 33, pi. ii. fi^. 1 :
A. Milne Edwards and Bouvier, Ciust. Decap.
Hirondclle et Princesse Alice, Monaco, J899, p. 11.
Homola spinifrons Leacb, Tr. Linn. Soc. XI. 1815,
p. 324, and Zool. Miscell. II., p. 82, pi. Ixxxviii.
Desmarest, Consid. Gen. Crust., p. J34, pi. xvii.
figs. 1, la : Risso, Hist. Nat. Europ. Merid. V., p.
34 : Milne Edwards in Cuvier Resjne Anim. pi. xxxix.
fig. 2, and Hist. Nat. Crust. II. 183, pi. xxii. figs.
1-4 : Costa, Faun. Regn. NapoL, Crust. Brach., p. I.
Heller, Crust. Sudl. 'Europ. p. 149, pi. iv. figs. 12,
13 : Brocchi, Ann. Sci. Nat. Zool. (6) II. 1875, Art.
2, p. 107 (male iiy'peiidaqex) : A. Milne Edwards,
Hull. Mus. Comp. Zool VIII. 1880, p. 33: Carus,
Prodr. Faun. Jledit. I., p 498 : Ortmann, Zool. Jahrb.
Syst. VI. 1892, ]>. £42. ? Honiola t-pinijjes Guilding,

I^Tr. Linn. Soc. XIV. 1825, p. 334.

Bull. Mus. Comp. Zool. VIII. 1880, p. 33:
Bou\ier, Bull. Soc. Philoni. Paris, (S) VIII.
1895-96, p. 62.

Botli sides of N.

(• Atlantic. Azores,

^Mediterranean.

{

Caribbean Sea.

{Challenger Anomura, 1888, p. 19, pi. ii. fig. 1 ;\Philippines
Whitelegge, Mem. Austral. Mas. IV. 1900, p. 163. J Is. Austra

Andaman Sea.

s and
Australia.

Ki

Subgenus
HCJIOLAX Alcock.

„ MEGALOl'S,

Alcock.

Subgenus
Pakomola Wood-

ilason.

Ante.
Ante.
Ante.

Ante.

Indian Seas.

riivieri
I Risso

f Risso, Hist. Nat. Europ. Merid. V. 1826, p. 34:"]

Roux, Crust. Medit. pi. vii : Guerin, Icon. R^gne

Anim. pi. xiii. fig. 1 : Milne Edwards, Hist. Nat.

Crust. II. 183 : Costa, Faun. Regn. Napol., Crust.

Brach., p. 2 : Lucas, Hist. Nat. Anim. Artie. Crust.
, ■{ pi. svi. tig. 1 : Heller, Crust. Siidl. Europe, p. 151 : }>
'• Carus, Prodr. Faun. Medit. I. 499 : Ortmann, Zool.

Jahrb., Syst., VI. 1892, p. 542: Bouvier, Bull. Soc.

Philom. Paris, (8) VIII. 1895-96, pp. 60, 77, fig.s.

24, .34: A. M. Edw. and Bouvier, Travailleur et
^Talisman, Crust. Decap. pt. 1., p. 10.

Mediton-anean : At-
lantic nr. Canary
Ls.

„ rROPUNDORUJI,

Ale. & Anders.

Paromoi.oi'sis

Wood-Mason.

„ BOASI, Wood-
Mason.

Ante.
Ante.
Ante.

Arabian Sea.

Indian Seas.

80

Htpsophrys

Wood-Mason.

„ SUPERCILIOSA,

Wood-Mason.

I.ONGIPES, Ale.

& Anders.

Ante.

Ante.

Ante.

Arabian Sea.

Arabian Sea.

Homologenus

Henderson

rostratus,
(A. M. Edw.).

f ( = Eomolopsis, A. M. Edw., Bull. Mus. Comp."\
1 Zool. VIII. 1880, p. 341, Henderson, Cliallencrer |
-{ Auomura, y>. 20 : A. Milne Edwards and Bouvier, J-
I Crust. Decap. Hirondelle et Princesse Alice, Monaco, |
L.1899, p. 12. J

f Bull. Mus. Comp. Zool. VIII., 1880, p. 34, and^
I Rec. Fio-. Crust. Nouv., pi. 39, fig. 1 : Bouvier, Bull.
I Soc. Pliilom., Paris (8) VIII., p. 62. fig. 2.r, : Milne
■{ Edwards and Bouvier, Crust. Decap. Hirondelle et |-

Priucesse Alice, Monaco, 1899, p. 13, and Exp. Sci.

Tiavailleur et Talisman, Crust. Decap. Pt. 1, p. II,
^pl. i, fig. 1, pi. viii. figs. 19-21.

Both sides
Atlantic.

of N.

fp. Henderson. Challenger Anomura, p. 21, pi. ii, fig. 2.

Kear the Philippines,

Family Latreillid^.

Latkeillia Roux.

Ante.

' Crust. Medit. pi. xxii : Milne Edwards, Hist. Nat.^
Crust. I. 277 : Lucas, Espl. Sci. de I'Algerie, Zool.,
pt 1, p. 3, Crust, pi. i. fig. 1 : Heller, Crust, siidl.
Europ., p. 147, pi. iv. fig. 14 : Smith, P. U. S. N. M.,
1883, p. 23, and Albatross Crust, (in Rep. U. S. Fish.
Comm., 18S4), p. 7, pi. ii. figs. 2, 2a, pi. iii. fig. 1 :
„ elegans, Roux. -^ Carus, Prodr. Faun. Medit. I. p. 498 : Milne Edwards ^
and Bouvier, Crust. Decap. Hirondelle, Pt. I. Mo-
naco, 1894, p. 59, pi. vi. figs. 13-15, and Crust.
Decap. Hirondelle et Princesse Alice, Monaco, 1899,
p. 13, and Esp. Sci. Travailleur et Talisman, Crust.
Decap., pt. 1, p. 13: Bouvier, Bull. Soc. Philom.,
^Paris (8) VIII. 1896, p. 64, fig. 26. J

Both sides of N.
Atlantic : Medi-
terranean.

„ PENNIPEEA,

Alcock.

„ australiensis,
Henderson

Ante.

Indian Seas.

,, ralida.

DeHaan

,, phalangium,

DeHaan

Latreillopsis

Henderson.

„ HISPINOSA,

Henderson.

r Ciiallenger Anomura, 1888, p. 24
\ Whitelegge, Mem. Austral. Mus. IV

{Faun. Japon. Crust., p. 107, pi. xxx, fig. 1 : Adams"
and White, Samarang Crust., p. 5 : Henderson,
Challenger Anomui-a, p. 24.

(Faun. Japon. Crust., p. 108, pi. xxx. fig. 2 : Adams"^
and White, Samarang Crust., p. 5 : Ortmann, Zool. Vjapan.
Jahrb., Syst., VI. Is92, p. 542, pi. xxvi. fig. 2. J

Ante.

4, pi. ii. fig. 4:"l , , ,.
,1900, p. 165. |'^"s*^'''^l>

>s->|

J, >Japaii.

A)iti:

Audamans and
Philippines.

BRACHYURA

PLATES A AND I-VII.

-1^

m

-V\,

r>

.^

a-

c

o

S

tt!

§

o

a:

2:

-T3
C

c«

08

X.
-a

?

o

liMt' .Museum Ocu.sl/acea.

Brachyura Plate I .

|^«i^*" V'

-#•■?* ''Hi,;

0"'

V--^

:U8

V5

A.U. Dhowdhary S: '^.C.Mo'iHui dpi.ei l.ilh.

1. ;.-j.lKJc. A I

i.-jriljli V 2 R; .

Pj. Dynomene pilunitioif!''.-: x b.

Jhy.:i.

/cx3.

■ '>..'>.'<. AcTinLhoclriMiiia ■maTS^aril.-i x 8.

Indian Museum (Jrnslacea.

-Braohyiira Plate H.

^' v--=-

4

"1 m

vT

M

•t

*;yEf.

/v6'.

^1'

^

•"v;

..«:,

S7

ri

/

I.e. Chowdhary- &= o. C. Morid\il del.et Lilh.

4. Dn-imia rumptiu.

5. Dromia oramoideH x 1^.

Dromia unidentata ' 2.

7. Cryplodromia pileifera x6. 8. Cryptodromia canalicuiata x 2. 9.9a.Crypt.odromia bullifera x 6.

Indian Museum Crustacea.

Brachyura Plate III.

// \.3 (f.

/3 X -^ J .

W^

Si '%;'

m

'%.

%i

/Sx*.

S. C.Mondul & A.C. Chowdhary del.et LiUi.

11. Crypt, odromia hil^endorfl x3. 12. C'ryptodromia ebalioides x 5, 13. Cryplodromia gilesii x 4.

14,14a. Petalomera indica x S. 15.15e. Lisiodromia coppingeri x 4, 15. Chonohoecele.q artiliniosus.

Indian Museum Crustacea

Brachyura Plate IV.

^

19 x3.

/8a X3.

Mondui & A. G. Chowdhary del. et Lith

18. Spliaerodromia kendalli x2.

19. Sphaerodroniia n\ix x3.

Indian Mi]:-;cum IJi-uataaea.

A. 0. Chowdh.'-iiy & o.C.Monckil dal. el Litii.

2ij. Paromola [jrofundorum x 2.

i'lC.iH., MuiiCL.iu C;-LV.'.a('

Bi'aohyiira. Plate VI.

A.C. Chowdhar-y & 3. i.. Mondul del el; l.itli.

C4. I lypiK.'plify;' pupcrciUo.-jn x^.

25. llvpt.ophryH lon^ipe;^

Indian MuKOuin t^ruHtfetoec

..'lira Piaie Vi..

"=^

i\

\^

W

'^^

^ X "^,

X

-Si ^

\

}

Z7b xj.

S.C.MoTidui del. el Lith.

^6. L'S.tTC-illopcifc' bispinosa .•, 4

. 37. Latreillia penmfjr'

LIST OF PUBLICATIONS ISSUED BY THE TRUSTEES OF THE

INDIAN jyrUSBUM.

Rs. As. P.
Account of the Deep-Sea Bracliyura collected by the R.I.M.S. Investigator. By

A. Alcock, M.B., C.M.Z.S., F.G.S. ... ... ... ... ... C () 0

Account of the Deep-Sea Madreporaria collected by the R.I.M.S. Investigator. By

A. Alcock, M.B., C.M.Z.S., F.G.S. ... ... ... ... ... 4 0 0

Catalogue of the Archfeological Collections in the Indian Museum, Parts I and II. By J.

Anderson, M.D., F.R.S., etc. ... ... ... ... ... 4 12 0

Catalogue of Coins of the Indian Museum, Parts I to IV. By C. J. Rodgers, M.R.A.S.,

M.N.S. ... ... ... ... ... ... ... 24 0 0

Catalogue of Indian Crustacea. Part I. Introduction and Brachyura Primigenia. By

a": Alcock, M.B., LL.D., F.R.S, etc. ... ... ... ... .'.700

Catalogue of Indian Deep-Sea Crustacea Decapoda Macrura and Anoraala in the Indian

Museum. By A. Alcock, M.B., LL.D., C.M.Z.S.... ... ... ... 10 0 0

Catalogue of Indian Deep-Sea Fishes in the Indian Museum. By A. Alcock, M.B., C.M.Z.S.,

F.G.S. ... ... ... ... ... ... ... 5 0 0

Catalogue of Mammalia in the Indian Museum, Parts I and II. By J. Anderson, M.D.,

F.R.S., and W. L. Sclater, M.A., F.Z.S. ... ... ... .. GOO

Catalogue of Mantodea in the Indian Museum, Parts I and II. By J. Wood-Mason,

F.Z.S., etc. ... ... ... ... ... ... ... 2 0 0

Catalogue of Moths of India, Parts I to VII. By E. C. Cotes and C. Swinhoe, F.L.S.,

F.Z.S., etc. ... ... ... ... ... ... ... 5 12 0

Echinoderma of the Indian Museum : Deep-Sea Ophiuroidea collected by the R.I.M.S.

Investigator. By R. Koehler ... ... ... ... ... 10 0 0

Echinoderma of the Indian Museum : Shallow- Water Ophiuroidea collected by the R.I.M.S.

Investigator. By R. Koehler ... ... ... ... ... 400

Figures and Descriptions of Nine Species of Squillidfe from the Collection of the Indian

Museum. By J. Wood-Mason, F.Z.S. , etc. Edited by A. Alcock, M.B., C.M.Z.S.,

F.G.S. ... ... ... ... ' ... ... ... 2 0 0

Guide to the Zoological Collections Exhibited in the Bird Gallery of the Indian Museum.

By F. Finn, B.A., F.Z.S. ... ... ... ... ... ... 0 12 0

Guide to the Zoological Collections Exhibited in the Fish Gallery of the Indian Museum.

By A. Alcock, M.B., C.M.Z.S., F.G.S.... ... ... ... ... 0 8 0

Guide to the Zoological Collections Exhibited in the Invertebrate Gallery of the Indian

Museum. By A. Alcock, M.B., CM Z.S., F.G.S.... ... ... ... 0 10 0

Guide to the Zoological Collections Exhibited in the Reptile and Amphibia Gallery of the

Indian Museum. By A. Alcock, M.B., C.M.Z.S., F.G.S. ... ... ... 0 3 0

Hand List of MoUusca in the Indian Museum, Parts I and II, and Fasciculus E. By G.

Nevill, C.M.Z.S., etc. Index, Parts I and IL By W. Theobald ... ... 7 4 0

List of Batrachia in the Indian Museum. By W. L. Sclater, M. A., F.Z.S. ... ... 1 0 0

List of Birds in the Indian Museiun. Part I. Corvidse, Paradiseidge, Ptilonorhynchidas,

and Crateropodidse. By P. Finn, B.A., F.Z.S. ... ... ... ... 10 0

List of Snakes in the Indian Museum. By W. L. Sclater, M. A., F.Z.S. ... ... 10 0

Monograph of the Asiatic Chiroptera and Catalogue of the Species of Bats in the Indian

Museum. By G. E. Dobson, M.A., M.B., F.R.S., etc. ... ... ... 3 0 0

Monograph of the Oriental Cicadidas, Parts I to VII. By W. L. Distant, F.E.S. ...£.2 2 6

The above can be obtained from the Superintendejit of the Indian Museum, Calcutta, and from Messrs.
Friedlander Sf Sohn, 11, Garlstrasse, Berlin.