~» COMPOSITAE &
* NEWSLETTER

Number 18 June 1990

Scientific Editor: Bertil Nordenstam
Technical Editor: Agneta Lindhag
Department of Phanerogamic Botany,
P.O. Box 50007,

S-104 05 Stockholm, Sweden.
(Director: Prof. Bertil Nordenstam)

ISSN 0284-8422

CONTENTS

Editorial

C. Jeffrey: Recent References

C. Jeffrey: Generic Names of Compositae, Part VII

B. Nordenstam: Chromosome Number and Systematic
Position of Odontocline (Compositae-Senecioneae) 29

LIBRARY
JUL 9 1990

NEW YORK
BOTANICAL GARDEN

Comp. Newsl. 18. 1990

EDITORIAL

Dear Reader: The Compositae Newsletter is your forum - please use
it! Since the Recent References Series has now been closed down,
there is more room for other synantherological matters, such as ar-
ticles and reports, requests and announcements, news and notes, re-
views of literature, etc. Publication is free and the number of
subscribers now exceeds 500, so why don’t you take advantage of
this opportunity? Your contributions, suggestions and comments are
welcome.

Bertil Nordenstam

RECENT REFERENCES

Since the last list of Recent References in Comp. Newsl. 15, publi-
cation of annual volumes of the Kew Index of Taxonomic Literatu-
re for the years 1983-86 has been completed. From 1987 onwards,
the Kew Record is published quarterly. The need for further contri-
butions to this Recent References series has therefore been overta-
ken by events. Those wishing to obtain more recent references to
particular genera than are available in published issues of the Kew
Record should get in touch with me personally.

C. Jeffrey,

Herbarium,

Royal Botanic Gardens,
Kew,

Richmond,

Surrey TW9 3AE,

U.K.

2 Comp. Newsl. 18. 1990

GENERIC NAMES OF COMPOSITAE.
C. Jeffrey

Royal Botanic Gardens, Kew

Part VII. D

The following conventions are used in the list:

-, sometimes included in

=, a taxonomic synonym of

=, anomenclatural synonym of

Sp., number of species

M., monograph or revision

T., type

Tr., tribe

( ) Latin names enclosed in parentheses, name not validly published
[ ] enclosure in square brackets, reference not seen

As with previous parts of this serics, the author would be grateful for
notification of omissions and errors.

As with previous parts of this series, any indication of any type is not
intended as a formal act of lectotypification and is not to be construed as
such.

Dacryotrichia H. Wild, Garcia de Orta Ser. Bot. 1 (1-2): 67. 1973
T. D. robinsonii H. Wild (Zambia)

Sp. 1, Zambia

M.

Tr. Aster.

* Continued from Comp. Newsl. 16, 1989.

Comp. Newsl. 18. 1990 3

Dadia Vell., Fl. Flum. 347. 1825, genus inquirendum
T. D. lixa Vell. (Brazil)

Sp. 1, Brazil

M.

Tr. uncertain, perhaps not Compositae

Dahlia Cav., Ic. 1: 56, t.80. 1917

T. D. pinnata Cav. (Mex.)

Sp. 28, Mex., Guatem.

M. Sorensen in Rhodora 71: 309-365 & 367-416 (1969)
Tr. Helianth.

(Dalea P. Br., Civ. Nat. Hist. Journ. 314, 390 (sphalm. Delea). 1756)

Damatris Cass., Bull. Soc. Philom. Paris 1817: 139. 1817.
T. D. pudica Cass. (S Afr., Cape Prov.)
? = Haplocarpha Less.

Damironia Cass., Dict. Sc. Nat. 56: 224. 1828

T. D. cernua Cass. nom. illegit. (Xeranthemum variegatum L.) (S Afr., Cape
Prov.)

= Helipterum DC. nom. illegit. = Syncarpha DC.

Damnamenia Given, New Zeal. Journ. Bot. 11: 786. 1973 (- Celmisia Cass.)
T. D. vernicosa (Hook.f.) Given (New Zeal., Campbell I.)

Sp. 1, New Zeal. (Auckland & Campbell Is.)

M

Tr. Aster.
Danaa Colla, Mem. Reale Accad. Sc. Torino 38: 27, t.28. 1835 nom. illegit.

T. D. yegua Colla (Chile)
= Acrisione B. Nord.

4 Comp. Newsl. 18. 1990

Darwiniothamnus Harling, Act. Hort. Berg. 20(3): 108, 1962
T. D. tenuifolius (Hook.f.) Harling (Galapagos)

Sp. 2, Galapagos

M

Tr. Aster.
Dasiorima Rafin., Fl. Tellur. 2: 43. 1837

T. 3 syntypes, see Rafin., 1.c. (N Amer.)
= Solidago L.

Dasyanthus Bub., Fl. Pyrenaea 2: 198. 1899. nom. illegit. = Gnaphalium L.

Dasycondylus R.M. King & H. Robins., Phytologia 24: 188. 1972
T. D. lobii (Klatt) R.M. King & H. Robins. (Peru)

Sp. 7, Brazil to Peru

M.

T. Eupat.

Dasyphyllum Kunth in H.B.K., [Nov. Gen. Sp. 4 ed. fol. 13. 1818]
T. D argenteum Kunth (Ecuador)

Sp. 36, S Amer.

M. Cabrera in Rev. Mus. La Plata n.s. Bot. 9: 21-100. 1959

Tr. Mutis.

(Dasyranthus Rafin. ex Steud. Nom. ed. 2, 484. 1840 in syn.)

Daumailia Arenes, Bull. Soc. Bot. Fr. 95: 277. 1949
T. D. spinulosa Arenes (Syria)
= Urospermum Scop.

(Daveana Willk. ex Mariz corr. Mariz orth. mut. Dur. & Jacks., Ind. Kew. Suppl.

1:128, 1902 sphalm. = Daveaua Willk. ex Mariz corr. Mariz)

a

Comp. Newsl. 18. 1990 5

Daveaua Willk. ex Mariz, Bol. Soc. Brot. 9: 206,220, 243. 1891 (sphalm. Davea-
na) corr. Mariz, tom.cit. 258. 1891

T. D. anthemoides Mariz (Portugal)

Sp. 2, Portugal, N Afr.

M

Tr. Anthem.
(Davaélla Gandog., Fl. Europ. 14: 37. 1888)

Decachaeta DC., Prodr. 5: 133. 1836
T. D. haenkeana DC. (Mex.)

Sp. 7, Mex., C Amer.

M

Tr. Eupat.

Decaneurum DC., Arch. Bot. (Paris) 2: 516. 1833 nom. illegit. = Gymnanthe-
mum Cass. 1817 = Vernonia Schreb. sens. lat.

Decaneurum DC. ex Wight., Contrib. 7. 1834 nom. illegit.
T. D. reticulatum DC. ex Wight (India)
= Phyllocephalum Bl.

Decaneurum Sch. Bip., Tanacet. 44, 1844, nom. illegit. = Leucanthemella Tzve-
lev

Decastylocarpus Humb., Mem. Soc. Linn. Norm. (Comp. Madag.) 17, 30 cum
ic., 280, 1923.

T. D. perrieri Humb. (Madag.)

Sp. 1, Madag.

M

Tr. Vernon.

Decazesia F. Muell., Fragm. 11: 71. 1879
T. D. hecatocephala F. Muell. (W Austral.)
Sp. 1, Austral.

M.

Tr. Inul.

6 Comp. Newsl. 18. 1990

Deckera Sch. Bip., Flora 17: 478. 1834
T. Picris asplenioides L.
= Picris L.

Dectis Rafin., Fl. Tell. 2: 43. 1837
T. D. rugosa (Ait.) Rafin. (St. Helena)
= Commidendrum DC.

Deinandra Greene, FI. Franciscana 424. 1897
T. D. fasciculata (DC.) Greene (Calif.)
= Hemizonia DC.

Deinosmos Rafin., Fl. Tell. 2: 49. 1837 nom. illegit. = Dimorphanthes Cass.
= Pulicaria Gaertn.

Delairea Lem., Ann. Sc. Nat. Bot. Ser. 3, 1: 379. 1844 (= Senecio L. sens. lat.)
T. D. odorata Lem. (cult.)

Sp: 1,8. Afr.

M.

Tr. Senec.

Delamerea S. Moore, J. Bot. 38: 457, t. 416A. 1900
T. D. procumbens S. Moore (Kenya)

Sp. 1, trop. E Afr.

M

Tr. Inul.

Delilia Spreng., Bull. Soc. Philom. Paris 1823: 54, tab. II. 1823
T. D. berteri Spreng. (S Amer.)

Sp. 2, trop. Amer., Galapagos

M

Tr. Helianth.

Deloderium Cass., Dict. Sc. Nat. 48: 430. 1827
T. D. taraxacifolium Cass. (Crete)
= Leontodon L.

Delucia DC., Prodr. 5: 633. 1836
T. D. ostruthioides DC. (Mex.)
= Bidens L.

Comp. Newsl. 18. 1990

Demetria Lag., Gen. Sp. Pl. Nov. 30. 1816
T. D spathulata Lag. & D. glutinosa (Cav.) Lag. (syntypes) (Mex.)
= Grindelia Willd.

Demidium DC., Prodr. 6: 246. 1836
T. D. filagineum DC. (Madag.)
= Amphidoxa DC. - Gnaphalium L.

Dendranthema (DC.) Des Moul., Actes Soc. Linn. Bordeaux 20: [561]. 1858
T. D. indicum (L.) Des Moul. (China)

Sp. c.50, E & C Asia to Europe

M.

Tr. Anthem.

Dendrocacalia (Nak.) Nak. ex Tuyama, Bot. Mag. Tokyo 50: 129. 1936
T. D. crepidifolia (Nak.) Nak. ex Tuyama (Bonin Is.)

Sp. 1, Bonin Is.

M.

Tr. Senec.

Dendroseris D. Don, Philos. Mag. Ann. Chem. 11: 388. 1832
T. D. macrophylla D. Don (Juan Fernandez)

Sp. 710, Juan Fernandez

M.

Tr. Lactuc.

(Deneckia Thunb. orth. mut. Sch. Bip. in Walp., Rep. 2: 952. 1834 orth. var.
= Denekia Thunb. ex Schultes)

(Denekea Thunb. orth. mut. K. Dinter in Fedde, Rep. 16: 367. 1920 orth. var.
= Denekia Thunb. ex Schultes)

Denekia Thunb. ex Schultes, Fl. Cap., ed. 2, 665. 1823
T. D. capensis Thunb. (S Afr.)

Sp. 1, S trop. & S Afr.

M

Tr. Inul.

8 Comp. Newsl. 18. 1990

(Denekia Thunb., Prodr. Fl. Cap. 2: 153. 1800 nom. nud.)

(Denneckia Thunb. orth.mut. Steud., Nom. ed. 2: 491. 1840 orth. var.
= Denekia Thunb. ex Schultes)

Denira Adans., Fam. 2: 148, 549. 1763 nom. illegit. = Jva L.
(Dens Fabr., Enum. Meth. Pl. 67. 1759)
(Dens-leonis Seguier, Pl. Veron. 3: 264. 1754)

Derderia Jaub. & Spach, Ill. Pl. Or. 1: 129, t.67. 1843
T. Jurinea macrocephala DC. (Iran)
= Jurinea Cass.

Derouetia Boiss. & Bal., Diagn. Pl. Or. 3, ser. 2, 5: 114. 1856
T. D. frigida Boiss. & Bal. (Turkey)
= Crepis L.

Desmanthodium Benth. in Hook., Ic. 12: 14, t.1116. 1872
T. D. perfoliatum Benth. (Mex.)

Sp. 10, Mex. to Venez.

M.

Tr. Helianth.

Desmocephalum Hook., in Trans. Linn. Soc. Lond. 20: 208. 1843
T. D. inelegans Hook. (Galapagos)
= Delilia Spreng.

(Desmocephalus Hook. orth. mut. Benth. in Benth. & Hook.f., Gen.Pl. 2:347.
1873.
orth. var. = Desmocephalum Hook.)

Comp. Newsl. 18. 1990 9

Detridium Nees, Gen. Sp. Aster. 255: 1832
T. D. bergerianum (Spr.) Nees (sphalm. bergianum) (S Afr., Cape Prov.)
= Felicia Cass. nom. cons.

Detris Adans., Fam. 2: 135, 549. 1763 nom. rejic.

T. "Aster afric. flore coerul. fol. op. H.R. Par." (? = Felicia amelloides (L.) Voss)
(S Afr.)

= Felicia Cass. nom. cons.

Dewildemania O. Hoffm. in De Wild., Ann. Mus. Congo ser. 1, Bot. ser. 4, 1:X.
1903

T. D. filifolia O. Hoffm. (Zaire)

Sp. 3, S trop. Afr.

M

Tr. Vernon.

Diacantha Lag., Amen. Nat. Espafias 1: 40. 1811
T. Bucasia spinosa Ruiz & Pavon (Peru)
= Chuquiraga Juss.

Diacantha Less., Linnaea 5: 243. 1830 nom. illcgit.
T. D. arborea (Kunth) Less. (Ecuad.)
= Barnadesia Mutis.

Diacranthera R.M. King & H. Robins., Phytologia 24: 192. 1977
T. D. ulei R.M. King & H. Robins. (Braz.)

Sp. 2, Brazil

M.

Tr. Eupat.

Dialesta Kunth in H.B.K. Nov. Gen. Sp. [ed. fol. 4: 35] 1818
T. D. discolor Kunth (Colombia)
= Pollalesta Kunth

(Dianisteris Rafin., [Atl. Journ. 1: 153] 1832)

(Dianthoseris Sch. Bip., Flora 25: 439. 1842)

10 Comp. Newsl. 18. 1990

Dianthoseris Sch. Bip. ex A. Rich., Tent. Fl. Abyss. 1: 408. 1848
T. D. schimperi Sch. Bip. ex A. Rich. (Ethiop.)

Sp. 1, Ethiop.

M

Tr. Lactuc.

Diaperia Nutt., Trans. Amer. Philos. Soc. ser. 2, 7: 337. 1840
T. D. prolifera (Nutt. ex DC.) Nutt. (N Amer.)
= Evax Gaertn. = Filago L.

Diaphoranthus Meyen, Reise 1: 406. 1834
T. D. fuscus Meyen (Chile)
= Polyachyrus Lag.

Diaphractanthus Humbert

T. D. homolepis Humb. (Madag.)
Sp. 1, Madag.

M

Tr. Vernon.

Diaspananthus Miq., Ann. Mus. Bot. Lugd.-Bat. 2: 186. 1866 (- Ainsliaea DC.)
T. Ainsliaea uniflora Sch. Bip. (Japan)

Sp. 1, Japan

M

Tr. Mutis.

(Diaspanthus Miq. orth. mut. Kitamura, Mem. Coll. Sc. Kyoto Imp. Univ. ser. B,
15:304. 1940 orth. var. (sphalm.) = Diaspananthus Miq.)

(Diatonta Nutt. orth. mut. Walp., Rep. 2: 614. 1843 orth. var. = Diadonta Nutt.
= Coreopsis L.)

(Diatosperma A. Gray orth. mut. C. Muell. in Walp., Ann. Syst.
5: 211. 1859 orth. var. = Diotosperma A. Gray)

Comp. Newsl. 18. 1990 iL

Diazeuxis D. Don, Trans. Linn. Soc. Lond. 16: 251. 1830
T. D. trinervis D. Don (Peru)
= Lycoseris Cass.

Diacalymma Lem. ex Oerst., Ill. Hort. 2, misc. 37. 1855
T. D. fragrans Lem. (cult.)
= Podachaenium Benth.

Dibothrospermum Knaf, Flora 29: 298. 1843
T. Matricaria inodora L. (Europe)
= Tripleurospermum Sch. Bip.

Dichaeta Nutt., Trans. Amer. Phil. Soc. Soc.ser. 2, 7: 383. 1841
T. D. tenella Nutt. (Calif.)
= Baeria Fisch. & Mey. = Lasthenia Cass.

(Dichaeta Sch. Bip., Flora 33: 419. 1850 in syn.)

Dichaetophora A. Gray, Mem. Amer. Acad. Arts. ser. 2,4 (Pl. Fendl.) 73. 1849
T. D. campestris A. Gray (Mex. & Calif.)

Sp. 1,8 U.S.A., N Mex.

M.

Tr. Aster

(Dichotoma Sch. Bip. ex Benth. in Benth. & Hook-.f., Gen. Pl. 2: 364. 1873 in
syn.)

Dichrocephala L’Herit. ex DC., Arch. Bot. (Par.) 2: 517. 1833

T. D. bicola (Roth) Schlechtend. (=D. integrifolia (Linn.f.) O. Ktze)
Sp. 4, trop. & subtrop. Africa and Asia

M. Fayed in Mitt. Bot. Staatss. Munchen 15: 491-512. 1979

Tr. Aster.

12 Comp. Newsl. 18. 1990

(Dichropappus Sch. Bip. ex Kraschen., Not. Syst. Herb. Hort. Petrop. 4: 49.
1923 in syn.)

Dichromochlamys Dunlop, Journ. Adelaide Bot. Gard. 2: 235. 1980
T. D. dentatifolia (F. Muell.) Dunlop (Austral., N.T.)

Sp. 1, C Austral.

M.

Tr. Aster.

Dicoma Cass., Bull. Soc. Philom. Paris 1817:12. 1817
T. D. tomentosa Cass. (Senegal)

Sp. 35, trop. & S Africa, Madag., Arabia, India

Tr. Mutis.

Dicoria Torr. & A. Gray in Emory, (Rep. 143. 1858] & ex Torr. in Emory,
Rep. Bot. Bound. 86, t.30. 1858

T. D. canescens Torr. (Arizona)

Sp. 4, SW U.S.A., Mex.

M.

Tr. Helianth.

Dicranocarpus A. Gray, Mem. Amer. Acad. Arts ser. 2, 5: 322. 1854
T. D. parviflorus A. Gray (Mex.)

Sp. 1, Mex.

M.

Tr. Helianth.

(Dicroactis Rafin. [Herb. Raf. 67. 1833] nom. nud.)

(Dicrocephala L’ Herit. ex DC. orth. mut. Royle, Ill. Bot. Himal. Mts. 249. 1835
orth. var. (sphalm.) = Dichrocephala L’ Herit. ex DC.)

Comp. Newsl. 18. 1990

Didelta L’ Herit., Sturp. Nov. 55, t.28. 1786 nom. cons.

T. D. tetragoniifolia L’Herit. (S Afr., Cape Prov.)

Sp. 2, S Afr.

M. Roessler in Mitt. Bot. Staatss. Munchen 3: 304-312. 1959.
Tr. Arctot.

Dieteria Nutt., Trans. Amer. Phil. Soc. ser. 2, 7:300. 1840
T. 5 syntype species (N Amer.)
= Aster L.

Digitacalia Pippen, Contrib. U.S. Nat. Herb. 34: 378. 1968
T. D. jatrophoides (Kunth) Pippen (Mex.)

Sp. 4, Mex.

M.

Tr. Senec.

Diglossus (Cass.) Cass., Dict. Sc. Nat. 13: 241. 1819
T. D. variabilis Cass. (Peru)
= Tagetes L.

Dilepis Suesseng. & Merxm., Mitt. Bot. Staatss. Munchen 1: 14. 1950
T. D. dichotoma Suesseng. & Merxm. (Venez.)
= Flaveria Juss.

(Dimenostemma Cass. orth. mut. Steud., Nom. ed. 2, 1: 509. 1840 orth. var.
= Dimerostemma Cass.)

Dimeresia A. Gray, Syn. Fl. N Amer. ed. 2, 1(2) suppl. 448. 1886
T. D. howellii A. Gray (Oregon)

Sp. 1,W U.S.A.

M.

Tr. Helianth.

14 Comp. Newsl. 18. 1990

Dimerostemma Cass., Bull. Soc. Philom. Paris 1817: 11. 1817
T. D. brasiliana Cass. (Braz.)

Sp. 6, Braz., Boliv.

M.

Tr. Helianth.

Dimorphanthes Cass., Bull. Soc. Philom. Paris 1818: 20. 1818 nom. rejic.
T. D. sicula (L.) Cass. (Sicily)
= Pulicaria Gaertn.

Dimorphocoma F. Muell. & Tate, Trans. Proc. Roy. Soc. S Austral. 6: 107. 1883
T. D. minutula F. Muell. & Tate (Austral.)

Sp. 1, C Austral.

M.

Tr. uncertain, ? Aster., Anthem.

Dimorpholepis A. Gray in Hook., Ic. Pl. 9, t.856. 1851
T. D. australis A. Gray (Austral.)
= Helipterum DC. nom. illegit.

Dimorpholepis (G.M. Barroso) R.M. King & H. Robins., Phytologia 22: 118.
1971 nom. illegit. = Grazielia R.M. King & H. Robins.

Dimorphotheca Moench, Meth. 585. 1784 nom. cons.
T. D. pluvialis (L.) Moench (S Afr., Cape Prov.)
Sp.7,S Afr.

M. Norlindh, Studies in Calenduleae 1: 38 - 76. 1943
Tr. Calend.

Dinoseris Griseb., Symb. Fl. Argent. 213. 1879 & Abh. Konigl. Ges. Wiss.
Gotting. 24: 213. 1879

T. D salicifolia Griseb. (Argent.)

Sp. 1, S Boliv., N W Argent.

M.

Tr. Mutis.

Comp. Newsl. 18. 1990 15

(Dinosma Rafin. orth. mut. Post & Kuntze, Lex. 175. 1903 orth. var.
= Deinosmos Rafin. = Dimorphanthes Cass. = Pulicaria Gaertn.)

Diodonta Nutt., Trans. Amer. Phil. Soc. ser. 2, 7:360. 1841
T. 5 syntype species (N America)
= Bidens L.

Diodontium F. Muell. in Hook., Journ. Bot. Kew Gard. Misc. 9: 19. 1857
T. D. filifolium F. Muell. (Austral.)
= Glossogyne Cass.

Diomedea Cass., Bull. Soc. Philom. Paris 1817: 70.1817 nom. illegit. = Borri-
chia Adans.

(Diomedella Cass., Dict. Sc. Nat. 46: 398, 405. 1827 in syn.)

Diotis Desf. Fl. Atl. 2: 260. 1799 nom. illegit. non Schreb. 1791
T. D. maritima (L.) Cass.
= Otanthus Hoffmanns. & Link.

Diotosperma A. Gray in Hook., Ic. Pl. 9, t.855. 1851
T. D. drummondii A. Gray (W Austral.)
= Ceratogyne Turcz.

Diotostephus Cass., Dict. Sc. Nat. 48: 543. 1827
T. D. repens Cass. (patria ignota)
= Chrysogonum L.

Diplactis Rafin., Fl. Tell. 2: 45. 1837
T. several syntype spp., see Rafin., 1.c.
= Aster L.

16 Comp. Newsl. 18. 1990

Diplazoptilon Ling, Act. Phytotax. Sin. 10: 84. 1965 (? - Vladimiria Ijin)
T. D. picridifolium (Hand.-Mazz.) Ling (Sichuan)

Sp. 1, Sichuan

M.

Tr. Card.

Diplemium Rafin., Fl. Tell. 2: 50. 1837
T. Erigeron carolinum L. & E. quercifolium Lam. (syntypes, N Amer.)
= Erigeron L.

Diplocoma D. Don ex Sweet, Brit. Fl. Gard. 3, t.246. 1828
T. D. villosa (Sesse & Moc.) D. Don ex Sweet (Mex.)
= Heterotheca Cass.

Diplogon Rafin. [Amer. Monthly Mag. Crit. Rev. 4: 195. 1819] nom. rejic.
= Chrysopsis (Nutt.) Ell. nom. cons.

Diplopappus Cass., Bull. Soc. Philom. Paris 1817: 137. 1817
T. 3 syntype species (E North America)
= Aster L.

(Diplopapus Cass. orth. mut. Rafin., Fl. Tell. 2: 46. 1837 orth. var.
= Diplopappus Cass. = Aster L.)

Diplosastera Tausch, [Hort. Canal. 1, t.4. 1823]
T.D. tinctoria (Nutt.) Tausch (N Amer.)
= Coreopsis L

Diplostelma Rafin., New FI. 2:44, 95. 1837 nom. illegit. = Chaetopappa DC.

Diplostelma A. Gray, Mem. Amer. Acad. Arts ser. 2, 4: 72. 1849 nom. illegit.
T. D. bellioides A. Gray (Calif.)
= Chaetopappa DC.

Comp. Newsl. 18. 1990

Diplostemma Hochst. & Steud. ex DC., Prodr. 7: 75. 1838
T. D. alatum Hochst. & Steud. ex DC. (Yemen)
= Geigeria Griesselich

(Diplostephion Rafin., Fl. Tell. 2: 46. 1837)

Dipiostephium Kunth in H.B.K., Nov. Gen. Sp. [ed. fol. 4: 75. 1818]
T. D. lavandulifolium Kunth (Ecuad.)

Sp. 90, trop. Andes

M

Tr. Aster.

(Diplothria DC. orth. mut. Walp. Rep. 2: 620. 1843 orth. var. = Diplothrix DC.

= Zinnia L.)

Diplothrix DC., Prodr. 5: 611. 1836
T. D. acerosa DC. (Mex.)
= Zinnia L.

Dipterocome Fisch. & Mey., Ind. Sem. Hort. Bot. Petrop. 1: 26. 1835
T. D. pusilla Fisch. & Mey. (Iran)

Sp. 1, W Asia

M.

Tr. ?Calend.

Dipterocypsela S.F. Blake, Journ. Wash. Acad. Sc. 35: 36, fig. 1. 1945
T. D. succulenta S.F. Blake (Colombia)

Sp. 1, Colombia

M.

Tr. Vernon.

Dipterotheca Sch. Bip., Flora 25: 434. 1842
T. D. kotschyi Sch. Bip. (Sudan)
= Aspilia Thou.

18 Comp. Newsl. 18. 1990

Discomela Rafin., Neogenyton 3.1825
T. 3 syntype species (N Amer.)
= Helianthus L.

Discoseris (Endl.) O. Kuntze in Post & Kuntze, Lex. 181. 1903 nom. illegit.
= Richterago O. Kuntze = Gochnatia Kunth

Disparago Gaertn., Fruct. 2: 463, t.173. 1791 nom. cons.
T. D. ericoides (Berg.) Gaertn. (S Afr., Cape Prov.)

Sp. 7, S Afr.

M. Levyns in Journ. S Afr. Bot. 2: 95-103. 1936

Tr. Inul.

Dissothrix A. Gray in Hook., Journ. Bot. Kew Gard. Misc. 3: 223, t.5. 1851
T. Stevia imbricata Gardn. (N E Braz.)

Sp. 1, Brazil

M.

Tr. Eupat.

Distasis DC., Prodr. 5: 279. 1836
T. D. modesta DC. (Mex.)
= Chaetopappa DC.

Distegia Klatt, Bull. Herb. Boiss. 4: 463. 1846 nom. illegit. non Rafin. 1838
T. D. acida Klatt (S Afr.)
= Didelta L’ Hérit.

Distephanus (Cass.) Cass., Dict. Sc. Nat. 13: 361. 1819
T. D. populifolium (Lam.) Cass. (Mauritius)
= Vernonia Schreb. sens. lat.

Distoecha Phil., Anales Mus. Nac. Chile 1: 36, t.2, fig. 2. 1891
T. D. taraxacoides Phil. (Chile)
= Hypochaeris L.

Comp. Newsl. 18. 1990

Distreptus (Cass.) Cass., Dict. Sc. Nat. 13: 366. 1819 nom. illegit =
Pseudoelephantopus Rohr 1792 = Elephantopus L.

(Disynanthes Rafin. orth. mut. Reichb., Consp. 106. 1821 orth. var. =
Disynanthus Rafin. = Antennaria Gaertn.)

Disynanthus Rafin., [Amer. Monthly Mag. 268. 1818]
T. D. plantagineus Rafin. (N Amer.)
= Antennaria Gaertn.

Disynaphia Hook. & Am. ex DC., Prodr. 7: 267. 1838
T. D. montevidensis DC. (Urug.)

Sp. 12, Braz., Urug.

M. King & Robinson in Phytologia 22: 123-125. 1975
Tr. Eupat.

(Dithrichum Cass. orth. mut. DC., Prodr. 5: 619. 1836 orth. var. = Ditrichum
Cass. = Verbesina L.)

Dithyrostegia A. Gray in Hook., Journ. Bot. Kew Gard. Misc. 3: 97, 100. 1851
T. D. amplexicaulis A. Gray (Austral.)

Sp. 1, Austral.

M.

Tr. Inul.

Ditrichum Cass., Bull. Soc. Philom. Paris 1817: 33. 1817 nom. rejic. non
Hampe 1867 nom. cons.

T. D. macrophyllum Cass. (patria ignota)

= Verbesina L.

Dittrichia W. Greuter, Exsiccat. Genov. fasc. 4: 71. 1973 (- Inula L.)
T. D. viscosa (L.) W. Greuter (S Europe)

Sp. 2, Medit.

M.

Tr. Inul.

20 Comp. Newsl. 18. 1990

(Dizonium Willd. ex Schlechtend., Linnaea 5: 411. 1830 in syn.)

Doellia Sch. Bip. ex Walp., Rep. 2: 953. 1843
T. D. kotschyi Sch. Bip. nom. nud. (patria ignota)
= Blumea DC.

Doellingeria Nees, Gen. Sp. Aster. 177. 1832
T. 5 syntype species (4 N Amer., 1 Japan)
= Aster L.

Dolichlasium Lag., Amen. Nat. 33. 1811
T. D. lagascae Gill. ex D. Don (Argent.)
Sp. 1, Argent.

M

Tr. Mutis.

Dolichogyne DC., Prodr. 7: 256. 1838
T. D. staehelinoides DC. & D. gnaphalioides DC. (syntypes, Chile)
= Nardophyllum Hook. & Arn.

Dolichorrhiza (Pojark.) Galushko, Nov. Sist. Vyssh. Rast. 6: 210. 1970
T. D. renifolia (C.A. Mey.) Galushko (Caucas.)

Sp. 4, Caucas., Iran

M.

Tr. Senec.

(Dolicholasium Lag. orth. mut. Spreng., Gen. 2: 623. 1831 orth. var. =
Dolichlasium Lag.)

Dolichoglottis B. Nord., Op. Bot. 44: 33. 1978
T. D. lyallii (Hook.f.) B. Nord. (New Zeal.)
Sp. 2, New Zeal.

M.

Tr. Senec.

Comp. Newsl. 18. 1990

Dolichostylis Cass., Dict. Sc. Nat. 56: 138. 1828. nom. illegit. =
Fulcaldea Poir.

(Dolichotheca Cass., Dict. Sc. Nat. 51: 476. 1827)

Dolichotheca Cass. ex Less., Syn. Comp. 230. 1832 nom. illegit. =
Campylotheca Cass. - Bidens L.

Dolomiaea DC., Arch. Bot. (Paris) 2: 330. 1833
T. D. macrocephala Royle (Nepal)

Sp. 5, Tibet, Himal.

M

Tr. Card.

Dolosanthus Klatt, Bull. Herb. Boiss. 4: 472, t.5. 1896
T. D. sylvaticus Klatt (= Vernonia stenolepis Oliv.) (E trop. Afr.)
= Vernonia Schreb. sens. lat.

Donia R. Br. in Ait. & Ait., Hort. Kew. ed. 2, 5: 82. 1813
T. D. glutinosa (Cav.) R. Br. (Mex.)
= Grindelia Willd.

Doniana Rafin., [Amer. Monthly Mag. Crit. Rev. 2: 268. 1818] nom. illegit. =
Donia R. Br. = Grindelia Willd.

Doniophyton Wedd., Chloris And. 1: 7, t.4. 1855
T. D. andicola Wedd. (Chile)

Sp. 2, Chile, Argent.

M

Tr. Mutis.
Dontospermum Neck. ex Sch. Bip. in Walp., Rep. 2: 955. 1843

T. Buphthalmum maritimum L. (S France & Sicily)
= Asteriscus Mill.

21

22 Comp. Newsl. 18. 1990

Doratolepis (Benth.) Schlechtend., Linnaea 20: 593. 1847
T. D. tetrachaeta Schlechtend. (S Austral.)
= Leptorhynchus Less.

Doria Fabr., Enum. 73. 1759 nom. illegit. = Solidago L.

Doria Thunb., Nov. Gen. Pl. 167. 1800. nom. illegit.
T. 12 syntype species (S Afr., Cape Prov.)
= Othonna L.

Dorobaea Cass., Dict. Sc. Nat. 48: 447, 453. 1827 (- Senecio L.)
T. Senecio pimpinellifolius Kunth

Sp. 1, Colombia, Ecuador, Peru

M. Nordenstam in Op. Bot. 49: 51-53. 1978

Tr. Senec.

Doronicum L., Sp. Pl. 885. 1753 & Gen. Pl. ed. 5, 377. 1754
T. D. pardalianches L. (Eur. Alps)

Sp. 35, temp. Euras., N Afr.

M.

Tr. Senec.

Draconia Heist. ex Fabr., Enum. 79. 1759
T. Artemisia dracunculus L.
= Artemisia L.

(Dracontopsis Cass. orth. mut. Lem. in Orb. [Dict. 5: 173. 1849] orth.
var. = Dracopsis Cass. - Rudbeckia L.)

Dracopsis Cass., Dict. Sc. Nat. 35: 273. 1825 (- Rudbeckia L.)
T. D. amplexicaulis (L.) Cass. ex Less. (S U.S.A.)

Sp. 1,S U.S.A., N Mex.

M.

Tr. Helianth.

Comp. Newsl. 18. 1990 23

(Dracunculus Ledeb., Fl. Ross. 2: 566. 1846 in syn.) non Mill. 1754
Drepania Juss., Gen. 109. 1784 nom. illegit. = Tolpis Adans.

Dresslerothamnus H. Robins., Phytologia 40: 494. 1978
T. D. angusteradiatus (Barkley) H. Robins. (Panama)
Sp. 4, C America

M.

Tr. Senec.

Drozia Cass., Dict. Sc. Nat. 34: 217. 1825
T. D. dicephala Cass. (patria ignota)
= Perezia Lag.

Dubautia Gaudich. in Freycinet, Voy. Mond. Uranie Physic. Bot. 469, t.84. 1830
T. D. plantaginea Gaudich. (Hawaiian Is.)

Sp. 21, Hawaiian Is.

M. Carr in Allertonia 4: 62-107. 1985

Tr. Helianth.

Dubyaea DC., Prodr. 7: 247. 1838

T. D. hispida DC. (Himal.)

Sp. 10, Himal., W China

M. Stebbins in Mem. Torr. Bot. Cl. 19(3): 8-27. 1940
Tr. Lactuc.

Duchesnia Cass., Bull. Soc. Philom. Paris 1817: 153. 1817 nom. illegit.
= Francoeuria Cass. = Pulicaria Gaertn.

(Duchesnea Cass. orth. mut. Post & Kuntze, Lex. 188. 1903 orth. var.
= Duchesnia Cass. = Pulicaria Gaertn.)

(Dugaldea (Cass.) Cass. orth. mut. Meissn., Pl. Vasc. Gen. 2: 133. 1839
orth. var. = Dugaldia (Cass.) Cass. - Helenium L.)

24 Comp. Newsl. 18. 1990

Dugaldia (Cass.) Cass., Dict. Sc. Nat. 55: 264. 1828 (- Helenium L.)
T. Actinea integrifolia Kunth (Mex.)

Sp. 3, U.S.A., Mex., Guatem.

M. Biemer in Brittonia 26: 385-392. 1974.

Tr. Helianth.

Dugesia A. Gray, Proc. Amer. Acad. Arts 17: 215. 1886
T. D. mexicana (A. Gray) A. Gray (Mex.)

Sp. 1, Mex.

M.

Tr. Helianth.

Duhaldea DC., Prodr. 5: 366. 1836
T. D. chinensis DC. (China)
=InulaL.

(Duhaldia DC. orth. mut. Sch. Bip. in Walp., Rep. 2: 952. 1843 orth. var.
= Duhaldea DC. = Inula L.)

Duidaea S.F. Blake, Bull. Torr. Bot. Cl. 58: 496, pl. 42, fig. 8. 1931
T. D. tatei S.F. Blake (Venez.)

Sp. 3, Venez.

M.

Tr. Mutis.

(Dumantia DC. orth. mut. Farr et al., Ind. Nom. Gen. 1: 585. 1979 orth.
var. (sphalm.) = Dunantia DC. = IsocarphaR. Br.)

Dumerilia Lag. ex DC., Ann. Mus. Nat. Hist. Nat. 19: 64, 71. 1812
T. D. axillaris DC. (S Amer.)
= Jungia L.

Dumerilia Less., Linnaea 5: 13. 1830 nom. illegit.
T. D. humboldtii Less. (S Amer.)
= Perezia Lag.

Comp. Newsl. 18. 1990 25

Dunantia DC., Prodr. 5: 626. 1836
T. D. achyranthes DC. (Mex., Panama)
= IsocarphaR. Br

Durandoa Pomel, Mat. FI. Atl., 2. 1860
T. D. clausonis Pomel (N Afr.)
= Carthamus L.

Dusenia O. Hoffm. ex Dusén, Wiss. Ergebn. Schwed. Exped. Magell. 3:246.
1900. nom. illegit. = Duseniella K. Schum.

Duseniella K. Schum., Just’s Bot. Jahresber. 28(1): 475. 1902
T. D. patagonica (O. Hoffm. ex Dusén) K. Schum. (Argent.)
Sp 1, Argent. (Patagonia)

M.

Tr. Mutis.

Duttonia F. Muell., Linnaea 25: 409. 1853
T. D. sessiliceps F. Muell. (Austr.)
= Helipterum DC. nom. illegit.

Dymondia Compton, Journ. S Afr. Bot. 19: 110. 1953
T. D. margaretae Compton (S Afr., Cape Prov.)

Sp. 1, S Afr., Cape Prov.

M.

Tr. Arctot.

Dyscritogyne R.M. King & H. Robins., Phytologia 22: 158. 1971
T. Eupatorium adenospermum Sch. Bip. (Mex.)

Sp. 2, Mex.

M.

Tr. Eupat.

Dyscritothamnus B. L. Robins., Contrib. Gray Herb. ser. 2, 3, 65: 25. 1922
T. D. filifolius B. L. Robins. (Mex.)

Sp. 2, Mex.

M

Tr. Helianth.

26 Comp. Newsl. 18. 1990

(Dysinanthus Raf. ex DC., Prodr. 6: 270. 1838 in syn.)

(Dysodia Cav. orth. mut. DC., Prodr. 5: 639. 1836 orth. var. = Dyssodia Cav.)

(Dysodia Willd. orth. mut. Spreng. [Anleit. ed. 2, 2(2): 564. 1818] orth.
var. = Dyssodia Willd. = Adenophyllum Pers. - Dyssodia Cav.)

Dysodiopsis (A. Gray) Rydb., N Amer. Fl. 34: 170. 1915
T. Dyssodia tagetoides Torr. & Gray (Texas)
- Dyssodia Cav.

Dysodium L. C. Rich. ex Pers., Syn. Pl. 2: 489. 1807
T. D. divaricatum L. C. Rich. ex Pers. (Colombia)
= Melampodium L.

Dyssodia Cav. [Descr. 202. 1801]

T. D. papposa (Vent.) Hitchc. (Illinois)

Sp. 32 sens. lat. (SW U.S.A. to Colombia, Argentina), 8 sens. str. (SW U.S.A.,
Mex.)

M. Strother in Univ. Calif. Publ. Bot. 48: 1-88. 1969 (sens. lat.)

Tr. Helianth.

Dyssodia Willd., Enum. Pl. Hort. Berol. 900. 1809 nom. illegit.
T. D. porophyllum Willd. (Mex.)
= Adenophyllum Pers. - Dyssodia Cav.

Comp. Newsl. 18. 1990 oF

ADDITIONS AND CORRECTIONS

Acrisione B. Nord., Bot. Jahrb. Syst. 107: 582. 1985
T. A. cymosa (Remy) B. Nord. (Chile)

Sp. 2, Chile

M.

Tr. Senec.

Adenophyllum Pers., Syn. Pl. 2: 458. 1807 (- Dyssodia Cav.)
T. A. coccineum Pers. nom. illegit. (Mex.)

Sp. 9, SE U.S.A., Mex.

M.

Tr. Helianth.

Amphiachyris (DC.) Nutt., Trans. Amer. Philos. Soc. ser. 2, 7: 313. 1840
T. A. dracunculoides (DC.) Nutt. (Arkansas)

Sp. 2, U.S.A.

M. Lane in Syst. Bot. 4: 178-189. 1979

Tr. Aster.

Argyrovernonia MacLeish, Taxon 33: 106. 1984 (- Chresta Vell. ex DC.)
T. A. martii (DC.) MacLeish (Brazil)

Sp. 1, Brazil

M.

Tr. Vernon.

Askellia W. A. Weber, Phytologia 55: 6. 1984 (- Crepis L.)
T. Crepis nana Richardson (Canada)

Sp. 9, North Amer., N & C Asia, NW Himalaya

M.

Tr. Lactuc.

Billya Cass., Dict. Sc. Nat. 34: 38. 1825 non Billia Peyr. 1858 nom. cons. propos.
T. B. bergii Cass. (Gnaphalium asperum L.) (Cape Prov.) = Petalacte D. Don

28 Comp. Newsl. 18. 1990

Borkonstia Ignatov, Byull. Mosk. Obshch. Isp. Prirod. Biol. 88 (5): 105. 1983
nom. superfl. illegit. = Rhinactinidia Novopokr. (- Aster L.)

Brachythrix H. Wild & Pope, Kirkia 11: 25. 1978
T. B. stolzii (S. Moore) H. Wild & Pope (Tanzania)
Sp. 5, SE trop. Africa

M

Tr. Vernon.

Burkartia Crisci, Bol. Soc. Arg. Bot. 17: 242. 1976
T. B. lanigera (Hook. & Arn.) Crisci (Argent.)

Sp. 1, Argent.

M

Tr. Mutis.

Comp. Newsl. 18. 1990 29

Chromosome Number and Systematic Position of
Odontocline (Compositae-Senecioneae)

Bertil Nordenstam
Swedish Museum of Natural History
S-104 05 Stockholm, Sweden

Abstract

The somatic chromosome number 2n = 60 was counted in two species of Odon-
tocline B. Nord., a genus of the Compositae endemic to Jamaica. The systematic
position of the genus within the tribe Senecioneae is discussed in the light of
morphological, chemical and cytological evidence.

Introduction

The genus Odontocline B. Nord. contains six species, all endemic to the island of
Jamaica (Nordenstam 1978). The systematic position and affinites of the genus
within the tribe Senecioneae are somewhat obscure. Some important diagnostic
features such as the ’radial’ endothecium and ’senecioid’ filament collar indicate
a position in the subtribe Senecioninae. The continuous stigmatic surface of the
disc-floret style branches is a somewhat anomalous character in this context.
Other characteristic features are broad petiolate leaves, terminal compound syn-
florescences with yellow fragrant flowerheads, caudate anther bases, conical or
obtuse tips of style branches without a central tuft, and tooth-like receptacular
projections.

Material and methods

On the assumption that phytochemical and cytological data might shed some
light on the relationships of Odontocline, material for chemical analysis and for
cultivation was collected in Jamaica by the present author in 1980. The chemical
study was performed by Bohlmann et al. (1981), and plants were raised from
seeds in greenhouses in Stockholm. For chromosome counts root-tips were pre-
treated in colchicine, fixed in Cammoy’s solution, and squashed in aceto-orcein.

30 Comp. Newsl. 18. 1990

Results

Odontocline hollickii (Britton ex Greenm.) B. Nord. - Jamaica, Trelawney: 3,5
miles above Kinloss on road to Barbecue Bottom, limestone cliffs at roadside,
400 m s.m., 22.III.1980, Nordenstam 7901 (S voucher). - 2n = 60.

Odontocline glabra (Sw.) B. Nord. - Jamaica, Hanover: N slopes of Dolphin
Head, steep forested limestone slope, 400 m s.m., 31.III.1980, Nordenstam 7917
(S voucher). - 2n = 60 (Fig. 1).

Discussion

The ’senecioid’ complex comprises a large number of genera centered around the
core genus Senecio. This complex is now accomodated in the subtribe Senecioni-
nae (Jeffrey & Chen 1984), whcreas the ’cacalioid’ complex is recognized as sub-
tribe Tussilagininae. A number of Old World senecioid genera with tailed anthers
are referred to as the ’synotoid’ complex (Jeffrey & Chen 1984, Jeffrey 1986),
with Synotis as the largest genus.

The basic chromosome number in the Senecioninae is x = 10, many taxa be-
ing diploid with 2n = 20, but tetraploids (2n = 40) and hexaploids (2n = 60) as
well as higher numbers are frequent. Reduced numbers of 2n = 10 also occur, e.g.
in the Emilia group.

In Tussilagininae the basic number is x = 24-30, which can be interpreted as
a descending series from x = 30, an ancient polyploid level.

The number 2n = 60 found in Odontocline is thus not conclusive as regards
the taxonomic position of the genus. It can be interpreted as a hexaploid senccio-
id based on x = 10 or asa diploid cacalioid based on x = 30.

The morphological evidence indicates a position in subtribe Senccioninae,
although some characters are somewhat anomalous, as pointed out above. The
most closely related genus is probably Jacmaia, a monotypic genus likewise en-
demic to Jamaica. A striking difference is the appendaged Jacmaia style, which
accounts for the former placement of this taxon in Gynoxys, whereas Odontocline
until twelve years ago was retained in the over-expanded concept of Senecio. Se-
veral morphological characters indicate a close relationship between these two
genera, however, such as the tailed anthers, the continuous stigmatic areas of the
style branches, the receptacular projections, and the gencral habit with broad pe-
tiolate leaves. Jacmaia and Odontocline can be regarded as sister groups, but
their further affinities remain obscure. Possibly the South and Central American
genus Pentacalia is the best candidate as already suggested by Robinson & Cuat-
recasas (1978). The caudate anther base is one character in common, as well as

Comp. Newsl. 18. 1990 31

the fruticose habit with petiolate flat leaves, but there are also significant diffe-
rences in style and achene morphology.

The chemical evidence also indicates a close relationship between Odon-
tocline and Jacmaia (Bohlmann et al. 1981). The absence of furanoeremophila-
nes and the occurrence of acylpyrroles may seem to suggest relationship to some
senecioid groups now outside Senecio s.str. For example, some specialized Afri-
can ’synotoid’ genera (Mikaniopsis, Delairea) are also characterized by the lack
of furanoeremophilanes and the presence of acylpyrroles (Bohlmann et al. 1979).
This is probably not indicative of a close relationship to the Jamaican genera,
however, but more likely the result of parallel evolution. Bohlmann et al. (1979)
point out that several members of the Senecioneae have independently lost the
ability to synthesize furanoeremophilanes, and some of these instead produce va-
rious triterpenes or acylpyrroles. The latter are derived from pyrrolizidine alkalo-
ids, which are generally occurring throughout the tribe.

In summary, then, the chromosome number of 2n = 60 is inconclusive as re-
gards the affinities of Odontocline. The chemical evidence is difficult to interpret
at this stage. Morphology indicates a relationship of Odontocline and its sister
group Jacmaia to the more widespread Pentacalia. These genera may form a
New World ’pentacaloid’ complex distinct from the ’synotoid’ complex of the
Old World. The phylogeny of this complex needs to be further studied.

Acknowledgements
My thanks are due to Dr. Mari K4llersjé for skilful chromosome preparations. Fi-
nancial assistance from the Swedish Natural Science Research Council (grant B-
BU 2401-303 for Compositae taxonomy) is gratefully acknowledged.

32 Comp. Newsl. 18. 1990

References
Bohlmann, F., Zdero, C., Berger, D., Suwita, A., Mahanta, P. & Jeffrey, C. 1979.

Neue Furanoeremophilane und weitere Inhaltsstoffe aus siidafrikanischen
Senecio-Arten. - Phytochemistry 18: 79--93.

Bohlmann, F., Gupta, R. K. & Jakupovic, J. 1981. An acylpyrrole derivative and
further constituents from Jamaican representatives of the tribe Senecioneae.
- Phytochemistry 20: 831--832.

Jeffrey, C. 1986. The Senecioneae in East Tropical Africa. Notes on Composi-
tae: IV. - Kew Bulletin 41: 873--943.

Jeffrey, C. & Chen, Y.-L. 1984. Taxonomic studies on the tribe Senecioneae
(Compositae) of Eastern Asia. - Kew Bulletin 39: 205--446.

Nordenstam, B. 1978. Taxonomic studies on the tribe Senecioneae (Composi-
tae). - Opera Bot. 44: 1--83.

Robinson, H. & Cuatrecasas, J. 1978. A review of the Central American species
of Pentacalia (Asteraceae: Senecioneae). - Phytologia 40: 37--50.

SS)
S
BF SB ong eG
ioe ee
Bs Y = Si

Fig. 1. Somatic metaphase plate of Odontocline glabra,
2n = 60. The scale is 10 um.

Spangbergs Tryckeri AB, Stockholm 1990.