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BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

FOUNDED BY MARSHALL FIELD, 1893

VOLUME XVII

NUMBER 4

A CONTRIBUTION

TO THE

FLORA OF HONDURAS

BY

T. G, YUNCKER
PROFESSOR OF BOTANY, DEPAUW UNIVERSITY

B. E. DAHLGREN

CHIEF CURATOR, DEPARTMENT OP BOTANY
EDITOR

PUBLICATION 405

CHICAGO, U.S.A.
JANUARY 31, 1938

BOTANICAL SERIES

FIELD MUSEUM OF NATURAL HISTORY

FOUNDED BY MARSHALL FIELD, 1893

VOLUME XVII NUMBER 4

A CONTRIBUTION

TO THE

FLORA OF HONDURAS

BY

T. G, YUNCKER
PROFESSOR OF BOTANY, DEPAUW UNIVERSITY

B. E. DAHLGREN

CHIEF CURATOR, DEPARTMENT OF BOTANY
EDITOR

PUBLICATION 405

CHICAGO, U.S.A.
JANUARY 31, 1938

THE LIBRARY OF THE

^ 3 19 1938

UNIVERSITY 'OF ILLINOIS

PRINTED IN THE UNITED STATES OF AMERICA
BY FIELD MUSEUM PRESS

- 5
FB
V. 17 4

LIST OF PLATES

00 I. Typical view along the road in the interior of Honduras.

*o

*

* II. a. View of Siguatepeque, Honduras; 6. Hand-sawing pine

logs into boards for use in local construction.

oo HI. Lycopodium arcanum Maxon. Type specimen.

IV. Anthurium basiattenuatum Standley. Type specimen.

V. Anthurium lucens Standley. Type specimen.

VI. Anthurium Yunckeri Standley. Type specimen.

VII. Thecophyllum montanum L. B. Smith. Type specimen.

VIII. Tillandsia glossophylla L. B. Smith. Type specimen.

IX. Tillandsia lampropoda L. B. Smith. Type specimen.

X. Tillandsia Yunckeri L. B. Smith. Type specimen.

XI. Piper faviculiferum Trelease. Type specimen.

XII. Calliandra arborea Standley. Type specimen.

XIII. Desmodium luteolum Standley. Type specimen.

XIV. Ilex hondurensis Standley. Type specimen.
XV. Begonia Yunckeri Standley. Type specimen.

XVI. Achyrocline Yunckeri Blake. Type specimen.

XVII. Verbesina vicina Blake. Type specimen.

XVIII. Zexmenia perymenioides Blake. Type specimen.

287

A CONTRIBUTION
TO THE

FLORA OF HONDURAS

T. G. YUNCKER

INTRODUCTION

In 1883 William Botting Hemsley published in the Biologia
Centr all- Americana a catalogue of Central American plants. This
publication, which lists all the plants then known from Central
America, gives only about 160 species for Salvador and Honduras
combined. Hemsley states that he included these countries in his
list primarily to emphasize the great lack of knowledge regarding
the plants of that area. Other than this very inadequate enumera-
tion, practically nothing has been known of the flora of the Republic
of Honduras until comparatively few years ago.

During the winter of 1927-1928 Paul C. Standley made a bo-
tanical exploration of the northern coastal region, and in 1931
published the Flora of the Lancetilla Valley,1 a descriptive catalogue
of the plants which he had found in the valley and in the vicinity of
the nearby coastal town of Tela. This Flora, which covers a very
limited territory, is the only general treatment which we now have
of the plants of Honduras. He also has written The Woody Plants
of Siguatepeque2 from specimens and observations which he obtained
on a brief collecting trip to the interior. E. B. Bartram has pub-
lished a list3 of the species of mosses collected by Standley. In
1927 Professor Samuel J. Record visited Honduras, making col-
lections in the northern part of the country, and later published a
list4 of the trees. Other than these and a small number of additional
papers mostly describing occasional new species discovered by va-
rious collectors, comparatively little work has been done toward
producing a comprehensive flora of the country, and much of Hon-
duras remains as yet a terra incognita, botanically speaking.

During July and August, 1934 collections were made by the
writer in the Lancetilla Valley and Tela regions and to a lesser
extent in the interior about the village of Potrerillos and in the

1 Field Museum of Nat. Hist. Bot. Ser., vol. 10. 1931.
2Journ. Arnold Arboretum, vol. 9, no. 1. 1930.

3 Field Museum of Nat. Hist. Bot. Ser., vol. 4, pp. 349-364. 1929.

4 Tropical Woods, no. 10, June, 1927.

289

290 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

vicinity of Lake Yojoa. From June to the middle of August, 1936,
with the assistance of Ray Dawson and Howard Youse, collections
were made about the village of Siguateqepue and adjacent moun-
tains in the Department of Comayagua. Siguatepeque was selected
as the base from which to do our collecting mainly at the suggestion
of Standley, who believed that many species new to science
would be found in the mountains near there, and also because the
climate and living conditions at that altitude are generally agreeable.
The period of our visit coincided with the rainy season, but the rain-
fall was not sufficiently heavy to interfere greatly with our work.
It is believed that a fairly representative collection of those plants
in collectable condition during the period of our stay was obtained.
Many plants both herbaceous and woody, especially numbers of
orchids and bromeliads, were in neither flower nor fruit, and a period
of collecting later in the season would undoubtedly yield a large
number of additional species.

To reach the interior of the country one ordinarily lands at
Puerto Cortes on the north coast. The fastest and most comfortable
means of travel in the country is by airplane. The only other way
of reaching the interior is to go by train to the inland terminus of
the railroad at Potrerillos and then by motor-bus over a road inde-
scribably poor.1 The road goes by way of the beautiful Lake Yojoa,
across which one must ferry. If one can overlook the annoyances
occasioned by the poor condition of the road, the journey is of
considerable interest, especially to a botanist, because of the great
variety of vegetation through which one passes.

Siguatepeque is a quaint town of low, white-washed, Spanish-
type adobe houses with weathered, red tile roofs, situated at 1,100
meters altitude near the eastern edge of a small, more or less circular
plain. This plain, which is somewhat uneven and has occasional
rocky outcroppings, is surrounded by mountains which to the south-
west rise to an altitude of 2,000 or more meters. It is drained by
exceedingly crooked streams which during the rainy season carry
large amounts of water from the mountain slopes and have in many
places cut gorges several feet in depth like miniature grand canyons.

When there is sufficient rain, the plain is covered with abundant
short grass which affords excellent pasturage for considerable num-
bers of cattle, mules, and horses. Scattered here and there are farm

1 Standley gives an interesting description of his journey over this road
(Jour. Arn. Arb. 9:15. 1930). Our own experiences convince us that his comments
are not exaggerations.

FLORA OF HONDURAS 291

houses with gardens and small plantations of bananas and corn,
but in the main the plain is devoted to the grazing of stock. During
the rainy season numerous species of small herbaceous flowering
plants, weedy and otherwise, are very abundant in the grassland.
Scattered trees and a few small, shrubby species such as Psidium,
Lantana, and Acacia occur in neglected pastures, marshy areas, and
along occasional fence rows. Generally speaking, however, large
woody plants are not abundant except as planted about dwellings
or along the stream courses where a considerable number of woody
as well as herbaceous species grow well. During the dry season,
which ordinarily occurs between January and April, the soil is said
to become very dry and powdery and to be blown about in clouds
by any slight breeze. During this period the grass also dries up, and
it becomes necessary to drive the stock to meadows at higher alti-
tudes in the surrounding mountains to secure sufficient pasturage.
Here and there on the plain small, marshlike areas are found where
during the wet season occur many grasses, sedges, and other marsh
species.

Rising from the edge of the plain and extending to 1,300 or 1,400
meters altitude is a range of more or less rocky foothills forested
with a well-marked zone of Pinus oocarpa Schiede. The trees in
this zone are almost exclusively coniferous with occasional deciduous
trees or in some places are mixed with considerable numbers of oak.
The stand is open and parklike, and to ascend through it, especially
in early morning, is a delightful experience. The pines are mostly
fine, tall, straight specimens occasionally reaching a diameter of a
meter or more. During the wet season the forest floor is carpeted
with various species of flowering plants and short grasses which
provide excellent pasturage to large numbers of animals. It is
reported that in the dry season this region is burned over each year,
which, it is claimed, serves to destroy certain insect pests of the
cattle and keep undershrubs from becoming established. This
annual burning, together with the intensive grazing which is prac-
ticed, prevents any natural reforestration. Few young trees were
observed and some of the old trees have succumbed to repeated fire
wounds. The trees produce a good grade of timber which is used to
a limited extent locally for corner posts, beams, etc. in construction
of the adobe houses characteristic of the region. A small sawmill
runs when required, though much of the material is cut on location
by hand. The log is placed on a scaffolding, and one operator pulls
the saw from beneath while another stands on the log. A small

292 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

amount of resin which is distilled and used locally is collected from
the trees.

Originally a dense growth of deciduous forest extended from the
upper limits of the pine zone to the summit of the principal mountain
range. Much of the original forest has been destroyed, however,
and at the present time, between the lower pine-clad ridges and the
main range capped with deciduous virgin forest, lie gently to steeply
sloping pastures and cornfields cut here and there with deep, steep-
sided ravines through which swiftly descending mountain streams
dash over rocks as steep cataracts. In these gorges, densely
shaded by overhanging tree tops and wet with the spray of water-
falls, one finds the original vegetation surviving. Giant tree ferns,
Heliconias, arums, bromeliads, orchids, and many other herbaceous
and woody species grow in great profusion to produce a dense en-
tanglement through which one often needs to cut one's way. In
this intermediate zone one also discovers occasional subalpine
boggy areas exhibiting characteristic and interesting species.

This intermediate cleared zone, fringed principally with pine and
scrub oak at the lower edge, gives way to the more steeply rising
ridges and gulches of the highest range of peaks still covered for the
most part with deciduous forest. Each year sees additional areas
being fired and cleared for planting, with the result that in a com-
paratively short time most of the original deciduous forest as well
as the lower pine belt will have been destroyed, with the original
vegetation surviving only on the highest and steepest mountain
peaks and in ravines unsuited for agricultural purposes. Such
deforestation will likely bring about extensive erosion and serious
destruction of land now useful for agricultural purposes, as well as
have an effect on the rainfall.

In the forest at the summit giant, smooth-barked trees with
wide buttresses predominate. Their tops, which often meet far
overhead to produce a canopy through which the sun rarely pene-
trates, carry great numbers of epiphytes on their branches. These
ordinarily are accessible to collectors only when a branch breaks off
in a storm because of the excessive load which it carries. The under-
story is made up of many species of small trees, shrubs, palms, arums,
lianas, etc., which often form an entanglement of considerable
density, or in some places the forest may be more open. The forest
floor is overspread with deep, damp humus, rotting logs, and en-
tangled brush and vines. The logs and tree trunks are covered with
ferns, Peperomias, arums, and other low or creeping plants forming

FLORA OF HONDURAS 293

a matted growth with numerous species of mosses, liverworts, and
lichens.

The list of plants presented in this paper includes, in addition
to the plants collected about Siguatepeque in 1936, those obtained
in 1934 about Potrerillos and Lake Yojoa. Included also are those
species found in the Lancetilla-Tela region not previously reported
by Standley in his Flora. A list of over one hundred species cul-
tivated in and about Siguatepeque is not included, and a number of
common garden and street weeds observed about the village is also
omitted.

In the list those species collected on the plains about Siguatepe-
que are commonly cited without any indication of locality. El
Achote, given as the station for many species, is an area in the
mountains near the upper edge of the pine forest belt at about 1,350
meters altitude. The summit of the range above El Achote, which
is frequently enveloped in clouds, rises to an altitude of about 2,000
meters. El Rincon is a region about 12 kilometers west of Siguatepe-
que at the edge of the summit forest about 1,400 to 1,500 meters
altitude.

In the preparation of this list I have become greatly indebted
to the following specialists who have assisted by identifying and
verifying specimens as well as generously allowing the inclusion of
the descriptions of new species which were discovered: Mr. A. H. G.
Alston (Selaginella} ; Prof. Oakes Ames (Orchidaceae) ; Mr. E. B.
Bartram (Musci) ; Dr. S. F. Blake (Compositae) ; Mrs. Agnes Chase
(Gramineae); Dr. C. W. Dodge (lichens, excluding Cladonia);
Prof. A. W. Evans (Cladonia); Mr. E. P. Killip (Passifloraceae) ; Mr.
E. C. Leonard (Acanthaceae) ; Dr. W. R. Maxon (Pteridophyta) ;
Mr. C. V. Morton (Smilacaceae and Gesneriaceae) ; Dr. L. B. Smith
(Bromeliaceae) ; Dr. William Trelease (Piperaceae, Phoradendron,
and Quercus). I am especially indebted to Mr. Paul C. Standley,
who not only identified or verified most of the remaining numbers
and contributed critical notes to many species as well as the descrip-
tions of several new species, but who also gave much helpful advice
and information about the country and the conditions prevailing
there. To Messrs. Ray Dawson and Howard Youse who assisted
me on the 1936 expedition I wish to express my sincere appreciation.
One could not wish for more capable assistants and more cheerful
companions. I am also grateful to Dr. Wilson Popenoe and other
officials of the United Fruit Company who freely gave advice and
assistance whenever possible.

294 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

An almost complete set of the specimens upon which this paper
is based is deposited in the Herbarium of Field Museum.

LIST OF GENERA AND SPECIES

LICHENES
CHIODECTONACEAE

CHIODECTON Ach.

Chiodecton sanguineum (Sw.) Vainio. On tree trunk at El
Achote.

PANNARIACEAE
COCCOCARPIA Pers.

Coccocarpia cronia var. isidiophylla (M. A.) Dodge. On tree
trunk.

PANNARIA Del.

Pannaria lurida (Mont.) Nyl. On tree trunk, summit of the
range above El Achote.

STICTACEAE

LOBARIA (Schreb.) Hue

Lobaria conformis Vainio. On tree trunk near El Achote.

STICTA Schreb.

Sticta clathrata Dutr. On tree trunk.

Sticta laciniata (Sw.) Ach. On log, near the summit of the
range above El Achote.

Sticta Pickeringii Tuck. On tree trunk.

STICTINA (Nyl.) Hue

Stictina carpoloma (Del.) Nyl. On tree trunk near the
summit of the range above El Achote.

PELTIGERACEAE
PELTIGERA Willd.

Peltigera polydactyla (Neck.) Hoffm. On a moist bank in a
ravine near El Achote.

CLADONIACEAE
CLADONIA Hill

Cladonia chlorophaea (Floerke) Spreng. Wet soil, river bank.
Cladonia corymbulosa Nyl. On rotten log, ravine at El Achote.
Cladonia cristatella Tuck. On fence slats.

FLORA OF HONDURAS 295

Cladonia didyma (Fe"e) Vainio. On log near El Achote.

Cladonia fimbriata (L.) Fries. On log near El Achote.

Cladonia fimbriata var. Balfourii (Cromb.) Vainio. On a
tree trunk near El Achote.

Cladonia fimbriata var. chondroidea Vainio. On soil.

Cladonia Floerkeana (Fries) Sommerf. On tree trunk.

Cladonia furcata var. pinnata (Floerke) Vainio, f. truncata.
On log, top of the range above El Achote.

Cladonia pityrea (Floerke) Fries. On tree trunk near El
Achote.

Cladonia subcariosa Nyl. On dry soil.

PARMELIACEAE
PARMELIA Ach.

Parmelia americana Mey. & Flot. On log near the summit of
the range above El Achote.

Parmelia capitata Lynge. On fence pickets.

Parmelia cristifera Tayl. On tree trunk.

Parmelia latissima Fe"e. On fence post.

Parmelia pseudoflavescens Zahlbr. On fence pickets.

Parmelia subrugata Nyl. On tree trunk near El Achote.

Parmelia tinctorum Despr. On fence slat.

USNEACEAE
RAMALINA Ach.

Ramalina alludens Nyl. On tree trunk near El Achote.
Ramalina attenuata Pers. On tree trunk.
Ramalina yemensis (Ach.) Nyl. On tree trunk.

PHYSCIACEAE
ANAPTYCHIA Koerb.

Anaptychia leucomelaena (L.) Vainio. On fence post.
Anaptychia leucomelaena var. albociliata Nyl. On tree
trunk.

Anaptychia podocarpa var. conferta Vainio. On tree trunk.

PHYSCIA Ach.

Physcia segilata (Ach.) Nyl. On tree trunk.
Physcia syncolla Tuck. On tree trunk.

296 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

MUSCI

FISSIDENTACEAE
FISSIDENS Hedw.

Fissidens reclinatulus C. M. On moist bank, ravine near El
Achote.

LEUCOBRYACEAE
OCTOBLEPHARUM Hedw.
Octoblepharum albidum Hedw. On rotten log near El Achote.

POTTIACEAE
GLOBULINA C. Muell.

Globulina globulifera (Hpe.) C. M. New for Honduras. On
a dry bank, plains east of Siguatepeque.

FUNARIACEAE
FUNARIA Schreb.
Funaria calvescens Schwaegr. On soil.

BRYACEAE
BRYUM Dill.

Bryum andicola Hook. On moist bank, in a ravine near El
Achote.

Bryum argenteum Hedw. var. lanatum Bry. Eur. On rotten
log.

BARTRAMIACEAE
PHILONOTIS Bridel
Philonotis tenella (C. M.) Besch. On moist bank.

ORTHOTRICHACEAE
MACROMITRIUM Bridel

Macromitrium longifolium (Hook.) Brid. New for Hon-
duras. On a rotten log near El Achote.

Macromitrium mucronifolium (Hook. & Grev.) Schwaegr.
On tree trunk.

Macromitrium Podocarpi C. M. On log near El Achote.

Macromitrium Sumichrasti Dub. New to Honduras. On a
tree trunk near El Achote.

MICROMITRIUM Aust.

Micromitrium fragile (Mitt.) Jaeg. On log.

FLORA OF HONDURAS 297

SCHLOTHEIMIA Brid.

Schlotheimia Mohriana C. M. On bank of stream.
Schlotheimia Sartorii C. M. On tree trunk.

RHACOPILACEAE
RHACOPILUM P. de Beauv.
Rhacopilum tomentosum (Hedw.) Brid. On wet bank.

RHIZOGONIACEAE
RHIZOGONIUM Bridel

Rhizogonium spiniforme (Hedw.) Bruch. On a rotten log
near El Achote.

METEORIACEAE
PAPILLARIA (C. Muell.) C. Muell.
Papillaria nigrescens (Sw.) Jaeg. On tree trunk.

FABRONIACEAE

FABRONIA Raddi

Fabronia flavinervis C. M. On fence rails and tree trunks.

THUIDIACEAE

RAUIA Aust.
Rauia subcatenulata (Schp.) Broth. On tree trunk.

THUIDIUM B. & S.

Thuidium delicatulum (Hedw.) Mitt. On a log, summit of
the range above El Achote.

ENTODONTACEAE

ERYTHRODONTIUM Hamp.

Erythrodontium longisetum (Hook.) Par. On tree trunk.

SEMATOPHYLLACEAE
RHAPHIDORRHYNCHIUM Besch.
Rhaphidorrhynchium Lindigii (Hpe.) Broth. On log.

SEMATOPHYLLUM Mitten

Sematophyllum caespitosum (Hedw.) Mitt. Common on soil.
Sematophyllum cuspidiferum Mitt. New to Honduras. On
rock.

Sematophyllum galipense (C. M.) Mitt. On tree trunk.

298 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

HYPNACEAE

MITTENOTHAMNIUM Hennings
Mittenothamnium diminutivum (Hpe.) E. G. Britt. On

wet banks and rotten logs.

Mittenothamnium Lehmanii Besch. On tree trunk.

ISOPTERYGIUM Mitten

Isopterygium tenerum (Sw.) Mitt. On rotten log.

CTENIDIUM (Schimp.) Mitten

Ctenidium hondurense Edwin B. Bartram, sp. nov. — C.
mollusco (Hedw.) Mitt, forsan proximum, differt foliis breviter
acuminatis, fortiter plicatis, tenuibus denticulatis.

Probably dioicous, male flowers not seen. Rather robust, soft,
laxly tufted plants, green with a tinge of brown, slightly glossy.
Stems irregularly pinnately branched. Leaves crowded, strongly
falcate-secund with circinate tips; stem leaves up to 1.5 mm. long
and 0.7 mm. wide, lanceolate from an auriculate, broadly cordate
base, slightly decurrent, short-acuminate, deeply plicate, denticulate
all around; costa double, well defined, extending about one-third of
the way up the leaf; cells linear with rounded ends, 3-4/x wide and
8-12 times as long, with firm walls, shorter and irregularly rec-
tangular at the basal angles and across the leaf insertion; branch
leaves slightly smaller, from an oblong-ovate, scarcely cordate base,
short-acuminate, strongly plicate. Perichaetium large; inner peri-
chaetial leaves erect, ecostate, not plicate, up to 4.5 mm. long,
gradually narrowed to a long, filiform, slightly denticulate point;
seta 12 mm. long; capsule horizontal, oblong, about 2 mm. long.
Peristome, lid, and calyptra unknown.

HONDURAS: On a log near El Rincon, about 10 miles west of
Siguatepeque, 1,400-1,500 meters altitude, Department of Coma-
yagua, July 24, 1936, T. G. Yuncker, R.F. Dawson & H. R. Youse 6608.

While evidently near C. molluscum, this species differs in several
noteworthy particulars. Both the stem and branch leaves are
rather broadly short-acuminate and very strongly plicate; the costa
is well developed with the forks extending some distance up the leaf,
and the perichaetium is large and conspicuous.

POLYTRICHAGEAE
ATRICHUM Beauv.

Atrichum angustatum (Brid.) Bry. Eur. var. Mulleri (Schp.)
Bartr. On soil.

FLORA OF HONDURAS 299

OPHIOGLOSSACEAE
OPHIOGLOSSUM L.

Ophioglossum crotalophoroides Walt. On a moss-covered
rock in a wet ravine near El Achote.

Ophioglossum reticulatum L. Rich soil, river bank, about
6 kilometers west of Siguatepeque.

MARATTIACEAE
MARATTIA Sw.

Marattia excavata Underw.? In forest, summit of the range
above El Achote.

OSMUNDACEAE
OSMUNDA L.

Osmunda cinnamomea L. (form). Subalpine bog near El
Achote.

Osmunda regalis L. Subalpine bog near El Achote.

SCHIZAEACEAE
ANEMIA Sw.

Anemia adiantifolia (L.) Sw. On a rocky cliff near Potrerillos.
Anemia humilis (Cav.) Sw. On a rocky cliff in pine forest.
Anemia pastinacaria Moritz. In wet soil, pine forest.

Anemia Phyllitidis (L.) Sw. Terrestrial, wet ravine near El
Achote.

LYGODIUM Sw.

Lygodium polymorphum (Cav.) HBK. Climbing in a thicket
near Potrerillos.

GLEICHENIACEAE
DICRANOPTERIS Bernh.

Dicranopteris pectinata (Willd.) Underw. Scrambling over
shrubs in thicket on river bank in pine forest.

HYMENOPHYLLACEAE
HYMENOPHYLLUM J. E. Sm.

Ilymenophyllum asplenoides Sw. On a tree in forest at the
summit of the range above El Achote.

Ilymenophyllum fucoides Sw. On a tree in forest at the
summit of the range above El Achote.

300 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

Hymenophyllum polyanthos Sw. On a tree in forest at the
summit of the range above El Achote; also on trees and logs in a
wet ravine near El Achote.

TRICHOMANES L.

Trichomanes capillaceum L. On a tree in forest at the sum-
mit of the range above El Achote.

Trichomanes montanum Hook. On a tree in forest at the
summit of the range above El Achote.

Trichomanes radicans Sw. var. On a tree in forest at the
summit of the range above El Achote.

Trichomanes rigidum Sw. On side of a wet ravine near El
Rincon.

DICKSONIACEAE
CIBOTIUM Kaulf.

Gibotium regale Versch. & Lem. Leaves up to 2 meters in
length, the base covered with abundant, golden-brown hairs. Bottom
of a dark, wet ravine near El Achote.

CYATHEAGEAE
ALSOPHILA R. Br.

Alsophila mexicana Mart. Trunk up to 4.5 meters tall and 12-
14 cm. in diameter. Leaves 2-3 meters in length, the base covered
with thin, brown scales. In forest at the summit of the range above
El Achote.

Alsophila Schiedeana Presl. Trunk about 2 meters tall and
10-15 cm. in diameter. In forest at the summit of the range above
El Achote; also in a wet ravine near El Rincon.

CYATHEA Sm.

Cyathea mexicana Schlecht. & Cham. Trunk about 2 meters
tall. Leaves up to 2 meters in length, armed and with narrow,
brown scales at the base. In spray of waterfalls, bottom of a wet
ravine near El Achote.

HEMITELIA R. Br.

Hemitelia costaricensis (Klotzsch) Mett. Trunk about 1 meter
tall and 7 cm. in diameter. In a dry gulch, hills east of Siguatepeque.

FLORA OF HONDURAS 301

POLYPODIACEAE
ACROSTIGHUM L.

Acrostichum daneaefolium Langsd. & Fisch. In water,
edge of a marsh near Tela.

ADIANTOPSIS F<§e
Adiantopsis radiata (L.) Fe"e. On a wet cliff, river bank.

ADIANTUM L.

Adiantum concinnum H. & B. On a wet cliff in a ravine near
El Achote.

Adiantum tenerum Sw. In crevices of a rocky cliff near
Potrerillos.

Adiantum trapezoides F£e. In a wet ravine near El Achote.

ASPLENIUM L.

Asplenium auritum Sw. On a tree near the shore of Lake
Yojoa; also in a wet ravine near El Achote.

Asplenium cristatum Lam. Bottom of a wet ravine near El
Achote.

Asplenium formosum Willd. On a tree near the shore of
Lake Yojoa.

Asplenium miradorense Liebm. On a rotten log in forest at
the summit of the range above El Achote.

Asplenium praemorsum Sw. On a tree, river bank.

Asplenium radicans L. var. partitum (Klotzsch) Hieron.
Terrestrial. In a wet ravine near El Achote.

Asplenium serra Langsd. & Fisch. On a log in deep forest near
the summit of the range above El Achote.

Asplenium serratum L. On a tree near the shore of Lake
Yojoa.

BLECHNUM L.

Blechnum occidentale L. In a wet ravine near El Achote.
Blechnum unilaterale Sw. Rock hillside, beneath pine trees.

GHEILANTHES Sw.

Cheilanthes angustifolia HBK. In wet soil in pine forest,
above the plains of Siguatepeque.

302 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

DENNSTAEDTIA Bernh.

Dennstaedtia globulifera (Poir.) Hieron. On a wet cliff,
ravine near El Achote.

Dennstaedtia rubiginosa (Kaulf.) Moore. In a forest on a
ridge above Lancetilla Valley.

DIDYMOCHLAENA Desv.

Didymochlaena truncatula (Sw.) J. Sm. In a wet ravine near
El Rincon.

DIPLAZIUM Sw.

Diplazium Donnell-Smithii Christ. In a wet, shady ravine
near El Achote.

DRYOPTERIS Adans.

Dryopteris denticulata (Sw.) Kuntze. In cloud-zone forest
at the summit of the range above El Achote.

Dryopteris effusa var. divergens (Willd.) Hieron. In a ravine
near El Achote.

Dryopteris Hemsleyana (Baker) C. Chr. In cloud-zone forest
at the summit of the range above El Achote.

Dryopteris interjecta C. Chr. In a wet ravine near El Achote.

Dryopteris Linkiana (Presl) Maxon. In a wet ravine near El
Achote.

Dryopteris melanochlaena C. Chr. In a wet ravine near El
Achote.

Dryopteris nicaraguensis (Fourn.) C. Chr. Bottom of a wet
ravine near El Achote.

Dryopteris normalis C. Chr. Side of a ravine near El Achote.

Dryopteris oligocarpa (H. & B.) Kuntze. In a subalpine bog
near El Achote.

Dryopteris patula (Sw.) Underw. On rocky hillsides near El
Achote.

Dryopteris pseudosancta C. Chr. On river bank.

Dryopteris resiliens Maxon, sp. nov. — Sect. Goniopteris. Certe
D. paucijugae proxima, a qua differt: rhizomatis paleis fere glabris;
lamina ambitu late lanceolata; pinnis multijugis profundius pinnatifi-
dis, superioribus gradatim minoribus sed conformibus et sessilibus,

FLORA OF HONDURAS 303

laminae apice breviter acuminato paucilobato; segmentis longi-
oribus et plerumque falcatis, mmquam truncatis; segmentorum
majorum venis 10-13-ju.gis, basalibus non conniventibus; soris
indusiis fulvis nee fusco-vinaceis praeditis; parenchymate mem-
branacea et verruculosa, nee herbacea et laevi.

Rhizome horizontal or creeping, woody, 5-8 cm. long, about 1 cm.
thick, bearing numerous stout roots throughout, obscurely paleaceous
at apex, the scales very few, cucullate, coherent, subovate, 3-5 mm.
long, yellowish brown, thin, entire, nearly glabrous. Fronds 4-10,
subfasciculate, 40-70 cm. long, arching, the stipes about as long as the
blades, slender, dull stramineous, with a thin covering of minute,
stalked, stellate hairs; blades 20-35 cm. long, 12-20 cm. broad,
lanceolate to lanceovate, pinnate nearly throughout, only the short,
abruptly acuminate apex pinnately lobed ; pinnae numerous (15-19-
jugate), sessile, the 2 or 3 lowermost pairs sharply deflexed, these the
largest, 8-12 cm. long, 1.5-3 cm. broad, lance-attenuate, obliquely
pinnatifid two-thirds the distance to the costa or more, inequilateral
at base, subauriculate, the distal segment often nearly free, the seg-
ments in general mostly oblong or linear-oblong, falcate or subfalcate,
4-5 mm. broad at base, distally acutish; other pinnae narrowly
oblong, mostly from a slightly broader base, the upper ones gradually
smaller but conform, one or more often viviparous at base; costae
and costulae minutely puberulous beneath, the hairs both simple and
stellate; veins of larger segments 10-13 pairs, distant, oblique,
elevated, glabrate, simple or a few of the distal ones acutely forked
midway to the margin, the arcuate basal ones running out nearly or
quite to the sinus but never connivent; sori medial, small, distant;
indusia yellowish brown, firm, rigidly ciliolate, persistent. Leaf
tissue membranous, grayish green, verruculose throughout.

Type in the U. S. National Herbarium, No. 1,309,047, collected
in Lancetilla Valley, near Tela, Department of Atlantida, Honduras,
altitude 20-600 meters, in wet forest, Dec. 26, 1927 to March 20,
1928, by Paul C. Standley (No. 52,893). Other material at hand is
as follows:

HONDURAS: Lancetilla, alt. 20-600 meters, Yuncker 4609;
Standley 52894, 53977, 54578, 54873; Chickering 142; Pauline Ames
52, 61, 74, 77. Puerto Arturo, near Tela, Pauline Ames 177. Near
Puerto Castilla, Pauline Ames 87. Rio Pelo, Cordillera de Mico
Quemado, Dept. Yoro, alt. 60-380 meters, Pauline Ames 135, 138,
139. Near Ceiba, alt. 120 meters, Dyer A223. Rio Bermejo, Dept.
Santa Barbara, alt. 180 meters, J. D. Smith 5698 (leg. C. Thieme).

304 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

MEXICO: Ixtacomitan, Chiapas, alt. 120 meters, Sept. 27, 1887,
Rovirosa 51 .

Most of this material belongs to recent collections. Of the
specimens cited only the last two were seen by Dr. Christensen, and
not unnaturally were referred by him to D. paucijuga (Kl.) C. Chr.,
a species properly to be restricted to Tobago, Trinidad, and Vene-
zuela. The relationship is close, yet in the light of recent material
D. paucijuga may always be distinguished by its relatively broad,
subdeltoid blades, which also are gradually narrowed to an elongate,
lobate apex, its much fewer and less deeply pinnatifid pinnae, its
closer and often truncate segments, its subfuscous indusia, and its
smooth, thicker leaf tissue. Moreover, the rhizome scales are easily
seen and are conspicuously stellate-pubescent, whereas in D. resiliens
it is hard to find so much as a single stellate hair upon the scales,
which are few, closely packed, and nearly concealed at the apex of
the rhizome. Habitally the two species are alike, especially in their
several very strongly deflexed basal pinnae. In D. resiliens the
viviparous condition is more pronounced. A single frond often
bears 2 or 3 buds upon the upper rachis, these producing good-sized
plants, in one instance a young plant (still attached) of 9 fronds
measuring up to 12 or 13 cm. long.

Dryopteris resinifera (Desv.) Weatherby. On rock out-
cropping.

Dryopteris subincisa (Willd.) Urban. In a wet ravine near
El Achote.

ELAPHOGLOSSUM Schott

Elaphoglossum rampans (Baker) Christ. In a wet ravine
above El Achote.

Elaphoglossum rubescens (Kuhn) Christ. On a tree in forest
near El Achote.

Elaphoglossum scolopendrifolium (Raddi) J. Sm. On a
rotten log in forest at the summit of the range above El Achote.

Elaphoglossum tectum (H. & B.) Moore. In a subalpine
bog near El Achote.

HECISTOPTERIS J. Smith

Hecistopteris pumila (Spreng.) J. Sm. On a tree in forest above
Lancetilla Valley.

HEMIONITIS L.
Hemionitis pinnatifida Baker. In shade, river bank.

FLORA OF HONDURAS 305

LINDSAYA Dryander

Lindsaya quadrangularis Raddi. In a wet ravine near El
Rincon.

MAXONIA Christensen

Maxonia apiifolia (Sw.) C. Chr. var. dualis (Donn. Sm.) C.
Chr. Bottom of ravine near El Achote.

NEPHROLEPIS Schott

Nephrolepis cordifolia (L.) Presl. In a dark, wet ravine near
El Achote.

Nephrolepis pendula (Raddi) J. Sm. On a dead tree near the
shore of Lake Yojoa.

ODONTOSORIA (Presl) F£e

Odontosoria Schlechtendalii (Presl) C. Chr. Fronds 1-1.5
meters long, arching over a stream in a wet, shady ravine near El
Achote.

PITYROGRAMMA Link

Pityrogramma calomelaena (L.) Link. In thicket on river
bank.

Pityrogramma Schaffneri (Fe"e) Weatherby. On river bank
in shade.

Pityrogramma tartarea (Cav.) Maxon. On side of a wet
ravine near El Achote.

POLYPODIUM L.
Polypodium angustifolium Sw. On a tree along river bank.

Polypodium angustum (H. & B.) Mett. On a tree in wet
ravine near El Achote.

Polypodium apiculatum Kunze. On a tree in dense forest at
the summit of the range above El Achote. A species of northern
South America, previously known otherwise only from Hispaniola.

Polypodium asplenifolium L. On a tree in wet ravine near
El Rincon.

Polypodium astrolepis Liebm. On a tree, river bank near
Siguatepeque; also on a tree near the shore of Lake Yojoa.

Polypodium carpinterae Rosenstock. In a wet ravine near
El Achote; also at El Rincon. Known previously only from Costa
Rica.

306 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Polypodium crassifolium L. In a wet ravine near El Achote.
Polypodium cryptocarpon Fe"e. On a tree near Lake Yojoa.

Polypodium cultratum Willd. On a tree in forest at the
summit of the range above El Achote.

Polypodium delitescens Maxon. On a tree in dense forest at
the summit of the range above El Achote. New to Honduras ; common
in the high mountains of Jamaica, and known previously also from
Alta Verapaz, Guatemala.

Polypodium dissimile L. On the ground in forest at the
summit of the range above El Achote.

Polypodium fraternum Schlecht. & Cham. On a rock in pine
forest near El Achote.

Polypodium induens Maxon. On a tree in forest, summit of
the range above El Achote.

Polypodium Lindenianum Kunze. On a tree near Lake
Yojoa.

Polypodium loriceum L. On a rotten log in forest near the
summit of the range above El Achote.

Polypodium montigenum Maxon, sp. nov. — Sect. Eupolypo-
dium. Epiphy ticum, e turma P. plebeii, facie P. plebeium ipsum praeci-
pue simulans, a quo differt: rhizomatis paleis admodum obliquis et
adpresso-imbricatis, non anguste et crasse atro-carinatis, marginibus
pallidis teneris angustioribus et planis numquam complicato-crispis ;
laminis pinnatis, nee pinnatifidis; stipite et rhachibus non foliaceo-
marginatis subpersistente adpresso-paleaceis. Fortasse P. leu-
costicto propius, sed rhizomate longe repente libero paleis non atro-
carinatis angustioribus onusto, frondibus distantibus nee fasciculatis,
laminis pinnatis nee pinnatifidis, segmentis non dilatatis inter alia
sat recedit.

Epiphytic; rhizome rampant, loosely attached by a few distant,
wiry roots, largely free, 30-40 cm. long, 3-5 mm. thick, flattish and
shallowly sulcate in drying, divaricately branched, persistently
paleaceous; scales very oblique, closely appressed-imbricate, lance-
subulate, 3-4 mm. long, 1-1.5 mm. broad, dark castaneous, with
thin, pale, plane, fimbriate borders, or the underlying ones ovate-
attenuate, with broader borders, the scales thus bicolorous, the
broad, subopaque, median stripe percurrent. Fronds conform,
mostly distant, 20-50 cm. long, straight; stipes mostly a little

FLORA OF HONDURAS 307

shorter than the blades, 1-2 mm. thick, bisulcate, dull castaneous,
thinly and deciduously appressed-paleaceous; blades mostly 15-30
cm. long, 6-14 cm. broad, imparipinnate, the pale castaneous rachis
lightly marginate, deciduously paleaceous; pinnae chartaceo-cori-
aceous, 10-16 pairs, spreading, straight or subarcuate, adnate
(upper ones slightly decurrent), the fertile ones linear, 2-4 times their
width apart, mostly 4-7 mm. broad, acutish, the sterile ones closer,
broader (7-10 mm., rarely 15 mm. near middle), narrowed toward
base, sublanceolate, attenuate toward apex; margins cartilaginous,
faintly and remotely notched or, in large, sterile pinnae, sometimes
broadly serrulate; veins free, concealed, oblique, twice-forked;
pinnae punctate-paleaceous beneath, the scales distant, appressed,
0.5-1 mm. long, long-attenuate from a subtriangular base, light
castaneous, with deeply toothed, pale margins; sori 10-16 pairs,
uniserial, yellowish brown, globose, large (2-3 mm. in diameter),
medial, usually contiguous, often touching both midrib and margin.

Type in the U. S. National Herbarium, No. 1,182,092, collected
from a tree trunk at edge of forest, along cart road from Vara Blanca
(between Poas and Barba volcanoes) to La Concordia, Costa Rica,
alt. 1,600-1,950 meters, July 23, 1923, by William R. Maxon and
Alfred D. Harvey (No. 8479). A second specimen of this collection
has afforded data regarding the sterile fronds. Additional material
examined is as follows:

HONDURAS: Summit of range above El Achote, Dept. Comayagua,
alt. 1,800 meters, Yuncker, Dawson & Youse 6140.

COSTA RICA: Volcan de Barba, at 2,400-2,800 meters, M. Valeria
34, A91, 197. Volcan de Irazu, at 2,300 meters altitude, Pittier 840;
crater of volcano, Jimenez 1154 (greatly reduced). Volcan de
Turrialba, alt. 2,000-2,500 meters, Pittier 13257 (J. D. Smith 7487);
Standley 35059. Santa Clara de Cartago, alt. 1,950 meters, Lan-
kester 743, 901 ; Maxon & Harvey 8200. Las Nubes, Prov. San Jose",
alt. 1,500-1,900 meters, Standley 38781.

PANAMA: Cordillera above Camp I, Holcomb's Trail, above El
Boquete, Prov. Chiriqui, alt. 1,800-2,000 meters, Kittip 5301.

Habitally P. montigenum suggests P. plebeium Schlecht. & Cham.,
but that species is at once distinguished by its pinnatifid blades, its
closer and truly dilatate segments, and its shaggy, squarrose-pale-
aceous rhizomes, upon which the scales are densely packed, broad,
with a thick, very narrow, sharply elevated percurrent, black median
stripe, and with very broad, pale, ruffled borders. The relationship
to P. leucosticton Kunze is perhaps somewhat closer, but aside from

308 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

scale characters, that differs notably in its dimorphic, fasciculate
fronds, green-margined stipes, dilatate segments, and short, thick,
densely felted-radicose rhizomes.

Polypodium plumula H. & B. On a rotten log in a wet ravine
near El Rincon; also at the summit of the range above El Achote.

Polypodium polypodioides (L.) Watt. On a tree in a wet
ravine near El Achote.

Polypodium rhodopleuron Kunze. On a tree in a ravine near
El Achote.

Polypodium trichomanoides Sw. On a tree in dense forest
at the summit of the range above El Achote. New to Honduras;
common in the higher mountains of Jamaica, but otherwise known
previously only from a single collection from Alta Verapaz, Guate-
mala.

Polypodium triseriale Sw. On a tree near the summit of the
range above El Achote; also on a tree near Lake Yojoa.

POLYSTICHUM Roth

Polystichum platyphyllum (Willd.) Presl. In a wet ravine
near El Achote.

PTERIDIUM Scop.

Pteridium aquilinum (L.) Kuhn, var. Feei (Schaffn.) Maxon,
comb, nov.— Pteris Feei Schaffn. ex Fe"e, Me"m. Foug. 8: 73. 1857.

The present form, which was briefly described by Fee from
material collected at Huatusco (S chaff ner 138, 141) and on Popocate-
petl (Schaffner 186), is represented in the U. S. National Herbarium
by the following specimens:

MEXICO: Jalapa, Veracruz, Pringle 8342. Cordoba, Veracruz,
Finck 18; Kerber Exsicc. 79a. Huatusco, Veracruz, Mohr in 1857.
Guanajuato, Duges 6. Alvarez, San Luis Potosi, E. Palmer 67 in
1902. Teziutlan, Puebla, Orcutt 4029. Between Somoriel and Las
Lajas, Hidalgo, Rose 9204- Below Ajusco, Mexico, Rose & Painter
7214.

GUATEMALA: Laguna de Ayarza, Dept. Jalapa, alt. 2,400 meters,
Heyde &Lux (J. D. Smith 4080). Chichavac, Dept. Chimaltenango,
alt. 2,400-2,700 meters, Skutch 12. Bananera, Dept. Chimaltenango,
J. R. Johnston 512; Mrs. B. B. Lewis 448.

HONDURAS: Near Siguatepeque, Dept. Comayagua, alt. 1,050-
1,400 meters, Yuncker, Dawson & Youse5600, 5781; Standley 56034.

FLORA OF HONDURAS 309

These agree essentially and are characterized, briefly, by their
short, deltoid or broadly pentagonal blades, their extraordinarily
copious, pilose-villous, hairy covering (much less pronounced upon
the upper surface), their relatively broad and flat outer indusium,
and the invariable presence of many long, slender cilia fringing the
outer indusium.

Further studies may show this form to be specifically distinct.
Its relationship is with the variable P. aquilinum var. lanuginosum
(Bong.) Fernald (Rhodora 37: 247. 1935) of western North America,
with which it appears to be connected through two or three transi-
tional specimens from northern Mexico; but that so-called variety,
remarkably varied as to shape of blade, has a much narrower outer
indusium, which though hairy upon the surface is almost without
exception wholly devoid of cilia. In this last respect var. lanugino-
sum differs widely from most European specimens of P. aquilinum,
in which as a rule the outer indusium is conspicuously long-ciliate,
the cilia often reaching nearly to the midrib of the segments (as in
var. Feei)-, but even in Europe the cilia are sometimes altogether
wanting, though rarely so, and in leaf shape, hairiness, and other
characters European specimens show almost as great diversity as
is exhibited by American material.

PTERIS L.

Pteris orizabae Mart. & Gal. In forest near the summit of the
range above El Achote.

Pteris propinqua Agardh. In deep forest, Lancetilla Valley.
Pteris quadriaurita Retz. In a ravine above El Achote.

RHIPIDOPTERIS Schott

Rhipidopteris peltata (Sw.) Schott. On a tree in forest at the
summit of the range above El Achote.

STENOCHLAENA J. Smith

Stenochlaena vestita (Fourn.) Underwood. Climbing on a
tree in a wet ravine, El Achote.

STRUTHIOPTERIS Willd.

Struthiopteris polypodioides (Sw.) Trev. On a tree in dense
forest near the summit of the range above El Achote.

Struthiopteris varians (Fourn.) Broadh. In a wet ravine
near El Achote. Known previously only from Mexico.

310 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

TECTARIA Cav.

Tectaria dilacerata (Kunze) Maxon. In a wet ravine near El
Achote; also in forest above Lancetilla Valley.

VITTARIA J. E. Sm.

Vittaria dimorpha K. Muell. On a tree in a wet ravine near
El Achote.

Vittaria h'lifolia Fe"e. On a tree near the shore of Lake Yojoa.

MARSILIACEAE
MARSILEA L.

Marsilea sp. In water about 30 cm. deep in a marsh near
Siguatepeque.

LYCOPODIACEAE
LYCOPODIUM L.

Lycopodium arcanum Maxon, sp. nov. — PI. III. Sect. Urosta-
chys. Planta epiphytica, ca. 20 cm. longa et lata; caulis ascendens,
quinquies bipartitus, basi 2 mm. diametro, lignosus, angulatus; folia
numerosa, 10- vel 12-faria, patentia, plerumque arcuata, acicularia,
10-17 mm. longa, basi 0.5-0.7 mm. lata, sursum angustiora, usque
ad apicem subpungentem sensim attenuata, integra, longe decur-
rentia, sordide viridia, tenero-herbacea, siccitate per longitudinem
carinata vel bisulcata latere abaxiali, latere adaxiali profunde con-
cava vel subplicata, interdum ut videtur subtriquetra; sporangia
pauca, in parte superiori zonis disposita (sporophyllis conformibus),
rotundato-reniformia, 1.5 mm. lata, 1.2 mm. longa.

Plant epiphytic, 20 cm. long and broad, the stem 5 times divar-
icately bipartite, 2 mm. in diameter at base, about 1 mm. at apex,
rigid, not flattened in drying; leaves crowded in attachment, borne
in 10 or 12 ranks, acicular, spreading, mostly arcuate, 10-17 mm. long,
0.5-0.7 mm. broad at extreme base, 0.3-0.5 mm. broad below middle
when plane, tapering to a subpungent apex, entire, delicately her-
baceous, long-decurrent and sharply carinate upon the stem, on the
abaxial side carinate throughout or often bisulcate by shrinkage of
leaf tissue, on the adaxial side deeply concave or subplicate in drying,
the leaf sometimes appearing sub triquetrous; plant sparingly fertile
in the apical third, the sporangia borne in short, alternating zones,
the sporophylls identical with foliar leaves; sporangia rounded-
reniform, 1.5 mm. broad, 1.2 mm. long, with a deep sinus, protruding
far beyond the sides of the sporophyll.

FLORA OF HONDURAS 311

Type in the U. S. National Herbarium, No. 1,637,830, collected
at summit of range above El Achote, above the plains of Siguatepe-
que, Department of Comayagua, Honduras, at 1,800 meters altitude,
from the stem of a tree fern in rain forest, July 28, 1936, by T. G.
Yuncker, R. F. Dawson, and H. R. Youse (No. 6149). No other
material has been seen.

The present species is most nearly related to L. stamineum
Maxon (Smiths. Misc. Coll. 56, No. 29: 2. pi. 2. 1912) of Panama and
Costa Rica, which it resembles habitally. But in L. stamineum the
leaves are more numerous, invariably bright red at base, and of
great delicacy, being truly capillary. In the present plant there is
no trace of red.

Lycopodium cernuum L. In a subalpine bog near El Achote.
Lycopodium reflexum Lam. On a wet cliff, river bank.

Lycopodium Tuerckheimii Maxon. On a tree fern, in forest
at the summit of the range above El Achote.

SELAGINELLACEAE
SELAGINELLA Beauv.

Selaginella cuspidata Link. On a dripping cliff, river bank in
pine forest above Siguatepeque.

Selaginella cuspidata var. elongata Spring. Wet, shady
river bank.

Selaginella guatemalensis Baker. In wet ravines at El
Achote; also at the summit of the range in cloud zone.

Selaginella huehuetenangensis Hieron. On a cliff along river.

PINACEAE
PINUS L.

Pinus oocarpa Schiede. A tree 18 meters tall and 45 cm. in
diameter or sometimes larger. Forming a belt of pine forest about
the base of the mountains which rise from the edge of the plains of
Siguatepeque.

Pinus pseudostrobus Lindl. A tree 12-15 meters tall and 60-
75 cm. in diameter. In open pine forest above El Achote.

ALISMACEAE
ECHINODORUS L. Rich.

Echinodorus tenellus (HBK.) Buch. In a subalpine bog near
El Achote.

312 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

GRAMINEAE
ANDROPOGON L.

Andropogon bicornis L. In shallow water, margin of a marsh.
Andropogon furcatus Muhl. In old, wet pasture.
Andropogon leucostachyus HBK. In wet pasture.

ARUNDINARIA Michx.

Arundinaria sp. Two species were collected, both of which were
sterile. One had stems about 6 meters tall and 2.5 cm. in diameter.
The other had stems 3.5 meters tall, light green mottled with darker
green spots. In wet ravines at El Achote and El Rincon.

AXONOPUS Beauv.
Axonopus rhizomatosus Swallen. In old, wet pasture.

BAMBUSA Schreb.

Bambusa sp. Stems about 6 meters tall and 5 cm. in diameter,
yellow. Large clump on river bank.

ELEUSINE Gaertn.
Eleusine indica (L.) Gaertn. River bank.

ERAGROSTIS Host

Eragrostis hirta Fourn. River bank, vicinity of Siguatepeque.
"Very rare. New for Honduras" (Chase).

Eragrostis tephrosanthos Schult. On a wet bank.

ERIOCHRYSIS Beauv.

Eriochrysis cayennensis Beauv. In a subalpine bog near
El Achote.

LASIACIS (Griseb.) Hitchc.

Lasiacis divaricata (L.) Hitchc. Scrambling over shrubs in a
thicket, edge of forest near the summit of the range above El Achote.

Lasiacis procerrima (Hack.) Hitchc. Side of a wet ravine near
El Achote.

LEERSIA Sw.

Leersia hexandra Sw. In a marshy pasture.

FLORA OF HONDURAS 313

PANICUM L.

Panicum arenicoloides Ashe. On a rocky hillside.
Panicum laxum Sw. In old, marshy pasture.

Panicum maximum Jacq. Stems up to 1.2 meters tall. River
bank.

Panicum sphaerocarpon Ell. On a rocky hillside.

PASPALUM L.

Paspalum notatum Fliigge. In old pasture.
Paspalum plenum Chase. River bank.
Paspalum virgatum L. In a marsh near Tela.

PENNISETUM L. Rich.

Pennisetum setosum (Sw.) Rich. In wet ditch about 30 km.
west of Tela.

PENTARRHAPHIS HBK.

Pentarrhaphis scabra HBK. Inflorescence and upper parts of
the stems red. In old, wet pasture.

PHARUS L.
Pharus latif olius L. In the forest above Lancetilla.

ROTTBOELLIA L. f.
Rottboellia aurita Steud. var. stigmosa Hack. In a marsh.

SETARIA Beauv.
Setaria geniculata (Lam.) Beauv. In marshy pasture.

SORGHASTRUM Nash

Sorghastrum nutans (L.) Nash. Inflorescence reddish brown,
attractive. In an old banana plantation.

SPOROBOLUS R. Br.
Sporobolus indicus (L.) R. Br. In old, wet pasture.

TRICHOLAENA Schrad.
Tricholaena rosea Nees. River bank.

314 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

TRIPSAGUM L.

Tripsacum dactyloides L. In rich soil, river bank, 6 km. west
of Siguatepeque.

UNIOLA L.
Uniola Pittieri Hack. In sand near the beach west of Tela.

CYPERACEAE
CAREX L.

Carex Humboldtiana Steud. Side of a wet ravine near El
Achote. The species is new for the flora of Honduras.

Carex polystachya Sw. River bank.

CYPERUS L.

Cyperus cayennensis (Lam.) Britton. In a low, wet area on
plains. The species is not reported from Honduras, although
naturally to be expected there.

Cyperus haspan L. In old, wet pasture.

Cyperus incompletus (Jacq.) Link. River bank. Not re-
corded previously from Honduras.

Cyperus prolixus HBK. In a wet ravine near El Achote. A
new species record for Honduras.

Cyperus surinamensis Rottb. In old, wet pasture.
Cyperus tenerrimus Presl. In pine-forest belt in wet soil.

Cyperus unioloides R. Br. In old, wet pasture. A species
new for the Honduran flora.

ELEOCHARIS R. Br.
Eleocharis flaccida (Reichenb.) Urban. In a marsh.

Eleocharis geniculata (L.) R. & S. In a subalpine bog near
El Achote.

Eleocharis mutata (L.) R. & S. In a marsh. New for the
Honduran flora.

Eleocharis nodulosa (Roth) Schult. In a wet field.

Eleocharis sulcata (Roth) Nees. In a subalpine bog near El
Achote.

FIMBRISTYLIS Vahl
Fimbristylis diphylla (Retz.) Vahl. In an old, wet field.

FLORA OF HONDURAS 315

FUIRENA Rottb.
Fuirena simplex Vahl. In a subalpine bog near El Achote.

LIPOCARPHA R. Br.

Lipocarpha macula ta (Michx.) Torr. In a subalpine bog near
El Achote.

RYNCHOSPORA Vahl

Rynchospora barbata (L.) Kunth. Plants solitary or in small
clumps, scattered on a rocky hillside.

Rynchospora cephalotes (L.) Vahl. Margin of a marsh, 12
km. east of Progreso.

Rynchospora corymbosa (L.) Britton. In old, wet pasture.
The species is new for the flora of Honduras.

Rynchospora cyperoides (Sw.) Mart. In a marshy field. Not
reported previously from Honduras.

Rynchospora dives Standl., sp. nov. — Perennis ca. 1 m. alta
fere omnino glabra, rhizomate ut videtur brevi, culmis laevibus
acute triquetris sat dense foliatis; folia longa ad 9 mm. lata, margini-
bus scabfis; paniculae laxae elongatae, corymbis paucis axillaribus
compositis usque 6 cm. latis laxe multifloris, spiculis oblongo-
lanceolatis brunneo-stramineis ca. 5 mm. longis sessilibus vel breviter
pedicellatis, ramis hispidulis, bracteis linearibus vel fere filiformibus,
infimis longissimis, glumis oblongo-lanceolatis attenuatis glabris;
setae 6 nucem subaequantes retrorse scabrae pallidae; nux ca. 1 mm.
longa rotundato-ovalis pallide brunnescens lucida obscure longi-
trorsum costata minute reticulata, rostro cum tertia parte nucis
aequilongo viridi conico. — Honduras: Side of ravine near El Achote,
hills above the plains of Siguatepeque, Dept. Comayagua, 1,350
meters, July 15, 1936, T. G. Yuncker, R. F. Dawson & H. R. Youse
5900 (type in Herb. Field Mus.).

The relationship of the plant is with R. locuples Clarke, a species
of Costa Rica, which has wider leaves and larger achenes with a
relatively much longer beak.

Rynchospora polyphylla Vahl. Moist river bank about 5 km.
northwest of Siguatepeque.

Rynchospora robusta (Kunth) Boeckl. In a subalpine bog
near El Achote. An addition to the known flora of Honduras.

316 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

SCLERIA Berg

Scleria pterota Presl, var. melaleuca (Schlecht. & Cham.)
Standl. On a ledge, river bank, about 5 km. northwest of Sigua-
tepeque.

Scleria secans (L.) Urban. Scrambling over bushes in a thicket
near Lake Yojoa. A common species of Central America, but not
recorded heretofore from Honduras.

STENOPHYLLUS Raf.

Stenophyllus hirtellus (Schrad.) Standl. In an old, wet
field. In Central America listed previously only for Guatemala.

PALMAE
CHAMAEDOREA Willd.

Chamaedorea geonomaeformis Wendl. Stems 1-1.5 meters
tall and 2.5 cm. in diameter. Fruit becoming purplish black; rachis
orange-red. In a wet ravine near El Achote.

Chamaedorea graminifolia Wendl. Stems up to 6 meters
tall and 10-12 cm. in diameter. In forest near the summit of the
range above El Achote.

Chamaedorea Lindeniana Wendl. Stems about 2 meters
tall and 1-2 cm. in diameter. In a wet ravine near El Rincon.

ARACEAE
ANTHURIUM Schott

Anthurium basiattenuatum Standl., sp. nov. — PI. IV.
Epiphytica, caudiculo ut videtur brevi, internodiis brevissimis;
petiolus 9-12 cm. longus gracilis supra canaliculatus, dorso obscure
tricostatus geniculo ca. 1 cm. longo tumido instructus; lamina
anguste oblongo-lanceolata ca. 29 cm. longa atque 8 cm. lata, apicem
angustum versus longe sensim attenuata, basi acuta, crasse chartacea,
opaca, supra in sicco cinereo-viridis, nervis non elevatis, subtus fere
concolor, basi trinervia, costa crassa elevata, nervis lateralibus
utroque latere ca. 12 angulo semirecto adscendentibus teneris
prominentibus fere rectis 6-7 mm. a margine in nervum collectivum
undulatum conjunctis, venis prominulis laxe reticulatis; pedunculi
gracillimi petiolis paullo longiores; spatha late oblonga acutiuscula
2.5 cm. longa vel ultra ca. 1 cm. lata elevato-puncticulata; spadix
sessilis 2 cm. longa vel paullo ultra 3 mm. crassus cylindricus; sepala

FLORA OF HONDURAS 317

longitudine sua latiora. — Honduras: On fallen log in deep forest
near the summit of the ridge above El Achote, in cloud zone, above
the plains of Siguatepeque, Dept. Comayagua, 1,850 meters, August
1, 1936, T. G. Yuncker, R. F. Dawson & H. R. Youse 6268 (type in
Herb. Field Mus.).

"Spadix deep maroon; spathe, peduncles, petioles, and midrib
beneath pinkish."

Anthurium concinnatum Schott. On a rotten log near the
shore of Lake Yojoa.

Anthurium lucens Standl., sp. nov. — PI. V. Caudiculus ut
videtur brevis crassus, internodiis brevissimis; petiolus ca. 40 cm.
longus gracilis teretiusculus dorso grosse costatus; lamina chartacea
sagittato-triangularis ca. 37 cm. longa atque basi 22 cm. lata, apicem
acuminatum versus sensim angustata, basi late profunde (ca. 7 cm.)
cordata, lobis posticis obtusis, supra in sicco sublucida, venulis
prominulis, subtus fere concolor, basi 7-nervia, nervis infimis latere
exteriore nervos 3 emittentibus, costa utroque latere nervos ca. 4
adscendentes remote a margine junctos emittente; pedunculus ca.
43 cm. longus gracilis teretiusculus; spatha reflexa lineari-lanceolato-
oblonga 9.5 cm. longa 12 mm. lata apice angustissime filiformi-
attenuata; spadix gracilis sessilis 12 cm. longus basi 4 mm. crassus.
—Honduras: In wet ravine on bank of stream near El Achote, hills
above the plains of Siguatepeque, Dept. Comayagua, 1,350 meters,
July 13, 1936, T. G. Yuncker, R. F. Dawson & H. R. Youse 58U
(type in Herb. Field Mus.).

"Petioles and peduncles up to 1 meter long; spathe pale maroon;
spadix deep red to maroon."

Anthurium pallens Schott. Spathe and spadix green. On a
rotten log in a wet ravine near El Rincon; also in deep forest near
the summit of the range above El Achote.

Anthurium parvispathum Hemsl.? Spadix maroon to brown;
spathe green to light red; berries greenish red. On a log in forest
near the summit of the range above El Achote.

Anthurium scandens (Aubl.) Engler. Spathe and spadix
green; berries white. Bottom of a ravine near El Achote; also on a
tree near the shore of Lake Yojoa.

Anthurium Yunckeri Standl., sp. nov. — PL VI. Caudiculus
crassus ut videtur brevis, internodiis brevissimis, cataphyllis latis
ca. 4 cm. longis mox decompositis fibrosis; petiolus gracilis ca. 26

318 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

cm. longus subteres, geniculo vix incrassato ca. 1.5 cm. longo;
lamina triangulari-oblongo-cordata 23-25 cm. longa prope basin 9
cm. lata, e basi apicem versus sensim angustata, apice longe angu-
stissime acuminata, basi anguste profunde (ad 5 cm.) cordata, lobis
posticis apice rotundatis, supra in sicco opaca griseo-viridis, costa
elevata, venis prominentibus, subtus brunnescens, basi 7-nervia,
costa gracili elevata utroque latere nervos ca. 7 angulo angusto
adscendentes fere rectos remote a margine in nervum collectivum fere
regularem conjunctos emittente, nervo tenuiore inter pares primario-
rum parallelo interposito, venulis prominentibus reticulatis; pe-
dunculus gracilis petiolum aequans teretiusculus; spatha oblongo-
lanceolata 5.5 cm. longa 1 cm. lata apicem versus longissime anguste
attenuata; spadix sessilis vix ultra 4 cm. longus 3 mm. crassus apicem
versus paullo angustatus. — Honduras: Bottom of ravine near El
Achote, hills above the plains of Siguatepeque, Dept. Comayagua,
1,350 meters, July 17, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 5935 (type in Herb. Field Mus.).

"Spadix dark red; spathe and peduncle light red."

MONSTERA Adans.

Monstera pertusa (L.) de Vriese. On trees in a wet ravine
near El Achote.

SPATHIPHYLLUM Schott

Spathiphyllum Friedrichsthalii Schott. Spathe green ; spadix
whitish. Stream bank in a ravine near Achote.

BROMELIACEAE
CATOPSIS Griseb.

Catopsis brevifolia Mez & Werckle". On a tree near the shore
of Lake Yojoa.

Catopsis nitida (Hook.) Griseb. On a tree in a wet ravine at
El Achote.

Catopsis nutans (Sw.) Griseb. On a tree near bank of river.

Catopsis Oerstediana Mez. On a tree in forest at the summit
of the range above El Achote.

Catopsis subulata L. B. Smith. On a tree, thicket on plain.

PITCAIRNIA L'He>.
Pitcairnia sp. On ground near El Achote

FLORA OF HONDURAS 319

THECOPHYLLUM Andre"

Thecophyllum montanum L. B. Smith, sp. nov. — PL VII.
Acaule, 4 dm. altum; foliis dense crateriformi-rosulatis, 25 cm.
longis, obscure punctulato-lepidotis, dorso lineis latis undulatis
purpureis transverse pictis; vaginis ellipticis, 1 dm. longis; laminis
ligulatis, acutis, 30-35 mm. latis, planis; scapo erecto, medio 5 mm.
diametro, glabro; scapi vaginis erectis, quam internodia paulo
longioribus, late ellipticis, breve triangulari-laminatis; inflo-
rescentia pauciflora, subdensa, 5 cm. longa; bracteis primariis ex
elliptico acuminatis, flores subaequantibus, nullo modo imbricatis,
flores infimos 3 alteros 2 collaterals in axillis gerentibus; bracteis
florigeris suborbicularibus, obtusis, 6-9 mm. longis, quam sepala
bene brevioribus, late convexis, nullo modo carinatis, coriaceis,
laevibus, obscure punctulatis; floribus subsessilibus; sepalis late
ellipticis, obtusis, 12-15 mm. longis, coriaceis, extus glabris laevi-
busque; cet. ignotis.

HONDURAS: Comayagua: Epiphytic in virgin forest, summit of
range above El Achote, above the plains of Siguatepeque, alt. 1,850
meters, 1936 Yuncker, Dawson & Youse 6243 (Herb. Gray, type).

Thecophyllum montanum is most nearly related to T. Wercklea-
num of Costa Rica. Unlike the latter it has the leaf sheaths of the
same color as the blades, the primary bracts small and remote, and
the floral bracts little if any broader than long.

TILLANDSIA L.

Tillandsia Butzii Mez. Leaves mottled red and green. On a
tree in a ravine near El Achote.

Tillandsia cyanea E. Morr. On a tree in forest near the sum-
mit of the range above El Achote.

Tillandsia excelsa Griseb. Leaves thin, green with purple
spots above, purple beneath; inflorescence branching, reddish. On
a tree in forest near El Rincon.

Tillandsia fasciculata Sw. Leaves greenish purple. Bracts
and peduncle red. In forest near El Achote.

Tillandsia festucoides Brongn. On a tree near shore of Lake
Yojoa.

Tillandsia glossophylla L. B. Smith, sp. nov.— PI. VIII.
Acaulis, 4 dm. alta; foliis dense cyathiformi-rosulatis, 2 dm. longis,

320 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

minutissime obscureque brunneo-lepidotis; vaginis ellipticis, amplis,
1 dm. longis; laminis ligulatis, acutis, 25 mm. latis, planis, viridibus
vel purpureo-afflatis; scapo erecto, 5 mm. diametro; scapi bracteis
erectis, dense imbricatis, late ovatis, apiculatis vel infimis acuminatis,
glabris, nervatis; inflorescentia simplicissima, lineari-lanceolata,
acuta, 18 cm. longa, 25 mm. lata, complanata; bracteis florigeris
erectis, dense imbricatis, late ellipticis, obtusis, 3 cm. longis, sepala
subaequantibus, ecarinatis, carneo-rubris, glabris, nervatis, sub-
coriaceis; floribus subsessilibus; sepalis anguste ellipticis, obtusis,
subchartaceis, valde nervatis, glabris, liberis; petalis linearibus,
laminis anguste ellipticis, 1 cm. longis; staminibus inclusis; stylo
elongate; capsulis quam sepala paulo longioribus.

HONDURAS: Comayagua: In ravine near El Achote, above the
plains of Siguatepeque, alt. 1,350 meters, 1936, Yuncker, Dawson &
Youse 5934 (Herb. Gray, type).

Tillandsia glossophylla appears to be more nearly related to T.
Walteri Mez of Peru and Bolivia than to any North American
species. T. Walteri differs in its triangular leaf blades and generally
larger flower parts.

Tillandsia grandis Schlecht. In forest near El Achote.

Tillandsia juncea (R. & P.) LeConte. Peduncles and leaf tips
blood-red. On a tree near bank of river.

Tillandsia lampropoda L. B. Smith, sp. nov. — PI. IX. Acaulis,
ad 43 cm. alta; foliis multis, substrictis, anguste cyathiformi-rosu-
latis, inflorescentiam subaequantibus; vaginis oblongo-ellipticis, 1
dm. longis, dense obscureque punctulatis, fulgide purpureis; laminis
angustissime triangularibus, caudato-acuminatis, base 15-20 mm.
latis, densissime adpresseque cinereo-lepidotis; scapo erecto, 6 mm.
diametro, glabro; scapi bracteis erectis, imbricatis, late ovatis,
apiculatis vel infimis anguste caudato-laminatis, perobscure punctu-
latis, laevibus, lucidis, vivo rubris; inflorescentia simplicissima,
elliptica, acuta, valde complanata, 15 cm. longa, 7 cm. lata; bracteis
florigeris distichis, dense imbricatis, quam internodia 8-plo longiori-
bus, late ovatis, acutis, 5 cm. longis, 4 cm. latis, valde carinatis,
coriaceis, laevibus, lucidis, perobscure punctulatis, ad apicem versus
cinereo-lepidotis, apice citrinis, base rubris, anguste atro-mar-
ginatis; floribus solum imperfecte cognitis, subsessilibus; sepalis
anguste lanceolatis, acutis, 32 mm. longis, carinatis, coriaceis,
nervatis, glabris, liberis; petalis ca. 5 cm. longis; stylo elongato.

FLORA OF HONDURAS 321

HONDURAS: Comayagua: On tree near El Achote, above the plains
of Siguatepeque, alt. 1,350 meters, 1936, Yuncker, Dawson & Youse
5895 (Herb. Gray, type).

It is not possible to see whether the stamens are exserted or not,
but the style is much too long for the section Phytarrhiza. Conse-
quently it must belong to either Platystachys or Allardtia, and I am
inclined to feel that it is probably the latter. In habit T. lampropoda
resembles the larger varieties of T. fasciculata, but the purple leaf
sheaths and lustrous, mostly apiculate scape bracts readily distin-
guish it. Also in T. fasciculata the scales are uniform and completely
cover the sheath and blade, while in T. lampropoda there is an abrupt
transition from the inconspicuous, separated, punctiform scales of
the sheath to the much larger, overlapping ones of the blade.

Tillandsia Leiboldiana Schlecht. Leaves green above, purplish
beneath ; inflorescence bright red. On a tree in ravine near El Achote.
An exceptionally attractive species.

Tillandsia Makoyana Baker. On a tree near Siguatepeque.

Tillandsia multicaulis Steud. Leaves green, becoming purple
toward the base; bracts yellow. On a tree in forest near El Achote.

Tillandsia polystachya L. Leaves green with faintly purple
tips. On a tree in thicket near bank of river.

Tillandsia pruinosa Sw. On a tree near the shore of Lake
Yojoa.

Tillandsia punctulata Cham. & Schlecht. Leaves green, be-
coming purplish at the base; inflorescence red. On a tree in forest
at the summit of the range above El Achote.

Tillandsia recurvata L. Flowers white to lavender. On a
tree in a thicket on plain.

Tillandsia Schiedeana Steud. Leaves reflexed; peduncles red.
On a tree in forest near El Achote.

Tillandsia Standleyi L. B. Smith. Peduncles and bracts
scarlet. A very attractive species. On a tree in forest near El Achote.
Known only from this region.

Tillandsia usneoides L. Common on trees on the plains and
hills; also about Lake Yojoa.

Tillandsia vestita Cham. & Schlecht. On a tree near the shore
of Lake Yojoa.

322 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Tillandsia Yunckeri L. B. Smith, sp. nov.— PL X. Acaulis,
verisimiliter 5 dm. alta; foliis multis, crateriforme rosulatis, rectis,

4 dm. longis, valde purpureo-suffultis; vaginis late ellipticis, ca. 15
cm. longis, dense punctulato-lepidotis; laminis ligulatis, acutis, ad

5 cm. latis, planis, supra glabris, subtus obscure punctulato-lepidotis;
scapo erecto, valido, glabro; scapi bracteis erectis, dense imbricatis,
foliaceis, magnis; inflorescentia composita, anguste pyramidata, 2
dm. longa, ad apicem versus densa; bracteis primariis suborbiculari-
bus, breve foliaceo-laminatis, spicas axillares superantibus amplexan-
tibusque, vivo fulgide rubris, laevibus, glabris, lucidis; spicis ses-
silibus, late lanceolatis, acutis, dense 4-floris, basi bractea sterili
praeditis, 65 mm. longis, 25 mm. latis, complanatis, supremis sub-
erectis, infimis patentibus; bracteis florigeris ovato-lanceolatis,
acutis, ad 45 mm. longis, sepala multo superantibus, acute carinatis
et lateribus planis, coriaceis, laevibus vel obscure nervatis, glabris;
floribus subsessilibus; sepalis lanceolatis, acuminatis, 3 cm. longis,
coriaceis, glabris, acute carinatis, posterioribus alte connatis; petalis
tubuloso-erectis, 5-6 cm. longis; staminibus verisimiliter exsertis.

HONDURAS: Comayagua: Epiphytic in forest at summit of range
above El Achote, above the plains of Siguatepeque, alt. 1,800 meters,
1936, Yuncker, Dawson & Youse 6432 (Herb. Gray, type); 6031
(Herb. Gray, cotype).

The material at hand does not show with certainty whether the
stamens are exserted or included, but the probability is that Til-
landsia Yunckeri belongs to the section Platystachys, where it is
most closely related to T. prodigiosa Baker. It differs from this
latter species in its much fewer-flowered spikes and very short-
bladed primary bracts as well as in its ligulate, acute leaves.

COMMELINACEAE
CAMPELIA L. Rich.

Campelia Zanonia (L.) HBK. An herb, about 1.5 meters tall.
Flowers pinkish violet; fruit violet. In sandy soil, bottom of a wet
ravine near El Achote.

COMMELINA L.

Commelina coelestis Willd. An herb about 50 cm. tall.
Flowers blue. In old, wet pasture.

TINANTIA Scheidw.

Tinantia erecta Schlecht. Stems very succulent, up to 1 meter
tall and 1.5 cm. in diameter. Petals deep blue. River bank about

6 km. north of Siguatepeque.

FLORA OF HONDURAS 323

TRADESCANTIA L.

Tradescantia elongata Mey. Petals violet to pink. In a wet
ravine near El Achote.

PONTEDERIACEAE
EICHHORNIA Kunth

Eichhornia azurea (Sw.) Kunth. In a stream between Tela
and Lancetilla Valley.

HETERANTHERA R. & P.

Heteranthera limosa (Sw.) Willd. Flowers pale lavender,
delicate. In wet ditches.

Heteranthera reniformis R. & P. Corolla white. In a sub-
alpine bog near El Achote.

LILIACEAE
SCHOENOCAULON Gray

Schoenocaulon officinale (Schl. & Cham.) Gray. An herb
with white flowers. In moist soil, river bank.

SMILACINA Desf.

Smilacina paniculata Mart. & Gal. Flowers white; branches
of the inflorescence and the berries reddish. In forest near the sum-
mit of the range above El Achote.

SMILACACEAE
SMILAX L.

S mi lax subpubescens A. DC. A woody vine. In a thicket at
the margin of forest near the summit of the range above El Achote.

Six additional species of Smilax were collected in sterile condition,
of which four are believed by C. V. Morton to be new species
though not describable in their sterile condition.

AMARYLLIDACEAE
BOMAREA Mirb.

Bomarea ovata (Cav.) Mirb. Sepals yellow flushed with pink;
corolla greenish with maroon spots. Climbing in a thicket near
river bank about 6 km. north of Siguatepeque.

324 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

CRINUM L.

Crinum cruentum Ker. In a low, marshy area between Tela
and Lancetilla Valley.

IRIDACEAE
SISYRINCHIUM L.

Sisyrinchium alatutn Hook. Flowers bright yellow. In wet
soil.

MUSACEAE

HELICONIA L.

Heliconia librata Griggs. Plant about 3 meters tall; bracts
golden yellow. In rich, moist forest near Lake Yojoa.

Heliconia pendula Wawra. An herb about 2 meters tall;
peduncle and bracts bright red. Bottom of a wet ravine near El
Achote; also near Tela River, Lancetilla Valley.

ZINGIBERACEAE
HEDYCHIUM Koenig

Hedychium coronarium Koenig. In moist soil between Tela
and Lancetilla Valley.

RENEALMIA L. f.

Renealmia aromatica (Aubl.) Griseb. Stems about 1 meter
tall; inflorescence scarlet. In a wet ravine near El Rincon.

MARANTACEAE
CALATHEA Meyer

Calathea verapax Donn. Sm. An herb about 1 meter tall;
bracts scarlet. In a wet ravine near El Rincon.

ORCHIDACEAE

BRASSAVOLA R. Br.

Brassavola cucullata R. Br. On a tree in Lancetilla Valley.

CATASETUM L. Rich.

Catasetum maculatum Kunth. On a tree near the shore of
Lake Yojoa.

COMPARETTIA Poepp. & Endl.

Comparettia falcata Poepp. & Endl. Flowers pink. On a tree
near El Achote.

FLORA OF HONDURAS 325

CRANICHIS Sw.

Cranichis pseudociliata Schltr. In humus in deep forest near
the summit of the range above El Achote.

DICHAEA Lindl.

Dichaea muricata (Sw.) Lindl. On a tree near the shore of
Lake Yojoa.

Dichaea trichocarpa Lindl. On a tree in a wet ravine near El
Rincon.

EPIDENDRUM L.

Epidendrum Brassavolae Reichb. f. On a tree near the sum-
mit of the range above El Achote.

Epidendrum centropetalum Reichb. f. On a tree in cloud
zone near the summit of the range above El Achote.

Epidendrum Chloe Reichb. f . On a tree near the summit of the
range above El Achote.

Epidendrum cochleatum L. On a tree near the shore of Lake
Yojoa.

Epidendrum cobanense Ames & Schltr. Flower parts fleshy,
golden-yellow turning to brown. On a tree in wet ravine near El
Achote.

Epidendrum difforme Jacq. On a tree near the shore of Lake
Yojoa.

Epidendrum Lindleyanum (Batem.) Reichb. f. On a tree in
a subalpine bog near El Achote.

Epidendrum luteoroseum A. Rich. & Gal. On a tree in pine-
oak association near El Achote.

Epidendrum ochraceum Lindl. On a tree near the summit of
the range above El Achote.

Epidendrum polyanthum Lindl. On a tree near the shore of
Lake Yojoa.

Epidendrum polyanthum var. myodes (Reichb. f.) Ames,
Hubb. & Schweinf. On a tree in a wet ravine at El Achote.

Epidendrum polybulbon Sw. On a tree in a ravine near El
Achote.

Epidendrum pygmaeum Hook. On a tree in forest near the
summit of the range above El Achote.

326 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

Epidendrum radicans Pav. ex Lindl. In a subalpine bog near
El Achote.

Epidendrum ramosum Jacq. var. mixtum (Schltr.) Ames,
Hubb. & Schweinf. On a pine tree in forest above El Achote.

Epidendrum rhynchophorum A. Rich. & Gal. On an oak
tree in forest near El Achote.

Epidendrum teretifolium Sw. On a tree near El Achote.

Epidendrum Tuerckheimii Schltr. On an oak tree near El
Achote.

Epidendrum virgatum Lindl. Rocky river bank.

ERYTHRODES Blume

Erythrodes Tuerckheimii (Schltr.) Ames. Terrestrial. In
forest near the summit of the range above El Achote.

GONGORA Ruiz & Pav.

Gongora cassidea Reichb. f. On a tree in a ravine near El
Achote.

GOVENIA Lindl.
Govenia Powellii Schltr. Flowers white. On river bank.

HABENARIA Willd.

Habenaria clypeata Lindl. Lactiflora form. On ground in
pine forest near El Achote.

Habenaria crassicornis Lindl. On ground in a wet ravine near
El Achote.

Habenaria novemfida Lindl. On ground in grass above El
Achote.

Habenaria setifera Lindl. On ground in grass near El Achote.

IONOPSIS HBK.
lonopsis utricularioides (Sw.) Lindl. On a tree near river.

ISOCHILUS R. Br.

Isochilus linearis R. Br. On a tree in a wet ravine near El
Achote: also near the shore of Lake Yojoa.

FLORA OF HONDURAS 327

LEPANTHES Sw.

Lepanthes acuniinata Schltr. On a rotten log in forest in
cloud zone near the summit of the range above El Achote. First
report of this species from Honduras.

Lepanthes Dawsonii Ames, sp. nov. — Herba epiphytica, dense
caespitosa. Caules secundarii gracillimi, plus minusve quinque-
vaginati, apice unifoliati, vaginarum ostiis obscure hispidis. Folium
elliptico-lanceolatum, breviter petiolatum. Inflorescentiae folio
breviores. Flores succedanei. Sepala lateralia anguste lanceolata.
Sepalum dorsale simile. Petala late reniformia. Labellum triloba-
tum, lobis lateralibus linearibus, lobo medio minute glanduloso.

Epiphytic herb, up to 4 cm. tall. Secondary stems erect or
obliquely ascending, 2-2.5 cm. long, concealed by about six closely
appressed, obscurely hispidulose sheaths which terminate in infundi-
buliform openings. Leaf 1-1.5 cm. long, about 6 mm. wide, elliptic-
lanceolate with the apex minutely tridentate. Flowers few, opening
in succession in abbreviated racemes. Lateral sepals 4 mm. long,
about 1 mm. wide, united to about the middle, narrowly lanceolate,
acuminate, acute, conspicuously 1-nerved through the middle.
Dorsal sepal similar and about equal to the laterals in length. Petals
reniform, 1 mm. in the longest dimension. Labellum 3-lobed; lateral
lobes hardly 1 mm. long, linear, sharply incurved at the distal end;
mid-lobe much shorter than the laterals, narrowly triangular,
minutely and obscurely glandulose. Column dilated upward.

Lepanthes Dawsonii is an ally of L. acuminata Schltr. from which
it differs in having smaller flowers in which the labellum has a pro-
nounced mid-lobe. In dried specimens the sepals are orange-yellow,
with an amethyst-purple labellum.

HONDURAS: Above the plains of Siguatepeque, on tree trunk in
forest near the summit of the range above El Achote, in cloud zone,
at 1,800 meters altitude, July 28, 1936. T. G. Yuncker, R. F. Dawson
& H. R. Youse 61 52 A (type in Herb. Ames No. 43769).

Lepanthes Yunckeri Ames, sp. nov. — Herba epiphytica, dense
caespitosa. Caules secundarii gracillimi, plus minusve quinque-
vaginati, apice unifoliati, vaginarum ostiis hispidis. Folium ellip-
ticum, marginatum, breviter petiolatum. Inflorescentiae singulae
vel duae, folio breviores. Flores succedanei. Sepala lateralia ovato-
lanceolata, acuminata, valde acuta. Petala rhomboidea. Labellum
integrum, cordato-suborbiculare, leviter retusum. Columna generis.

328 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Epiphytic herb, up to 4.5 cm. tall. Secondary stems 0.5-2 cm.
long, concealed by about five closely appressed, hispid sheaths which
terminate in infundibuliform openings. Leaf about 1 cm. long, up to
6 mm. wide, elliptic, marginate, coriaceous, with the apex minutely
tridenticulate. Peduncles about 7 mm. long, shorter than the leaves,
arising in the axil of a leaf, emerging singly or in pairs and bearing
a several-flowered, abbreviated inflorescence, the flowers opening in
succession. Sepals 4 mm. long, about 2.5 mm. wide with the dorsal
sepal 3-nerved and the laterals 2-nerved. Petals rhomboidal, later-
ally prolonged into subcaudate tips, 3 mm. in the longest dimension.
Labellum cordate-suborbicular, retuse in front, 2 mm. long, 2 mm.
wide, 3-nerved. Column dilated upward.

Lepanthes Yunckeri resembles L. eximia Ames in the vegetative
structures and in having the labellum simple rather than bilobed, but
differs markedly in the outline of the petals and in the more abbre-
viated peduncles. From dried specimens the sepals appear to have
been yellowish and the lip purplish with a tawny margin.

HONDURAS: Above Siguatepeque, on tree trunk in forest near
the summit of the range above El Achote, in cloud zone, at 1,800
meters altitude, July 28, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 6152 (type in Herb. Ames No. 46887).

LEPANTHOPSIS Ames

Lepanthopsis floripecten (Reichb. f.) Ames. Flowers light
brown. On a tree in wet ravine near El Achote.

MALAXIS Sw.

Malaxis aurea Ames. Flowers greenish. On ground, stream
bank.

Malaxis Maxonii Ames. Flowers greenish. On a rotten log in
forest near the summit of the range above El Achote.

Malaxis Parthonii Morr. On ground in a wet ravine near El
Achote.

MAXILLARIA R. & P.

Maxillaria uncata Batem. On a tree above Lancetilla Valley.

Maxillaria variabilis Batem. On a tree near the shore of Lake
Yojoa.

NOTYLIA Lindl.

Notylia trisepala Lindl. & Paxt. On a tree, hills above Lance-
tilla Valley.

FLORA OF HONDURAS 329

ONCIDIUM Sw.

Oncidium carthaginense Sw. On a tree, plains near Sigua-
tepeque.

Oncidium graminifolium Lindl. On rocks in a ravine near El
Achote.

Oncidium leucochilum Batem. On a tree, rocky river bank.

PELEXIA Poit.

Pelexia hondurensis Ames. On ground in a wet ravine near
El Achote.

PLEUROTHALLIS R. Br.

Pleurothallis Carioi Schltr. On a tree, bank of river.

Pleurothallis fuegii Reichb. f. On a tree near the summit of
the range above El Achote. Very rare.

Pleurothallis hirsuta Ames. On a fallen tree trunk near El
Achote.

Pleurothallis Johnsonii Ames. On a rocky cliff near El Achote.

Pleurothallis pansamalae Schltr. On a tree, summit forest
above El Achote.

POLYSTACHYA Hook.

Polystachya luteola Hook. On a rotten log, hills above
Lancetilla Valley.

Polystachya minor Fawc. & Rendl. Flowers fleshy, rich yel-
low. On trees in a wet ravine near El Achote.

RESTREPIA HBK.
Restrepia xanthophthalma Reichb. f. On tree near El Achote.

SARCOGLOTTIS Presl

Sarcoglottis picta Klotzsch. Flowers greenish white. In wet
soil under trees.

SCHOMBURGKIA Lindl.

Schomburgkia tibicinis Batem. On coconut palms near San
Juan Carib village, west of Tela.

SOBRALIA Ruiz & Pav.

Sobralia macrantha Lindl. On ground in a wet ravine near
El Achote.

330 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

STANHOPEA Hook.

Stanhopea ecornuta Lem. Flowers large, white. On a tree
near the shore of Lake Yojoa.

STELIS Sw.

Stelis hymenantha Schltr. On a tree near the summit of the
range above El Achote.

Stelis thecoglossa Reichb. f. On a tree near the summit of the
range above El Achote.

VANILLA Sw.

Vanilla pompona Schiede. Fruit triangular in section, up to
15 cm. in length. Climbing in a thicket, river bank.

PIPERACEAE
PEPEROMIA Ruiz & Pavon

Peperomia aggravescens Trelease, sp. nov.— Herba late
prostrata et radicans stolonifera, caulibus modice brevibus adsur-
gentibus rubris foliatis sparse minute hispidulis ad glabrescentibus
vix 2 mm. crassis; folia alterna, caulium sterilium orbicularia et 1
cm. lata, alia subrhombeo-elliptica et plus minusve obtuso-acumi-
nata vel lanceolata atque acuta, basi acuta, 2-2.5 cm. longa, 1-1.5
cm. lata, 1- vel obscure 3-5-nervia, subtus pallidiora, in sicco sub-
coriacea opaca; petiolus 3-5 mm. longus amplexicauli-decurrens;
spicae terminales 50-60 mm. longae 1 mm. crassae, pedunculo
gracili 10 mm. longo; bracteae rotundo-peltatae, ovario suboblique
apiculato, stigmate subapicali.

HONDURAS: On rotting wood, cloud zone above El Achote, near
Siguatepeque, Department of Comayagua, 1,800 meters, Yuncker,
Dawson & Youse 5990 (type in herb. Univ. Illinois; duplicate in Herb.
Field Mus.), also No. 6188 from the same locality.

Peperomia amnicola Trelease, sp. nov. — Herba majuscula
repenti-adsurgens epiphytica glabra, caule 3 mm. crasso; folia
alterna elliptico-oblanceolata vel subobovata subobtuso-acuminata,
basi cuneata, 4-7 cm. longa 2-3 cm. lata, 5-7-nervia, nervis basin
versus confluentibus, in sicco firma, subtus pallidiora, petiolo circiter
5 mm. longo; spicae apicales, immaturae tan turn visae.

HONDURAS: On a tree, river bank near Siguatepeque, Department
of Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5799 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

FLORA OF HONDURAS 331

Peperomia argumentosa Trelease, sp. nov. — Herba parvula
rhizomatosa glabra, caule vix 2 mm. crasso; folia alterna elliptica
vel elliptico-subobovata abrupte breviter acuminata 3-5 cm. longa
2-3 cm. lata 3-5-nervia, in sicco coriacea, subtus flavescentia, petiolo
gracili ca. 10 mm. longo; spicae terminales et ex axillis superioribus
nascentes vix 60-70 mm. longae et 2 mm. crassae, pedunculo 10-15
mm. longo; bracteae rotundo-peltatae; baccae rotundato-ovoideae
subapiculatae, stigmate obliquo.

HONDURAS: On a log in ravine near El Achote, above Siguatepe-
que, Department of Comayagua, 1,350 meters, Yuncker, Dawson &
Youse 5966 (type in herb. Univ. Illinois).

Peperomia caducipilosa Trelease, sp. nov. — Frutex radicanti-
pendula arboricola evanescente subconspicue molliter pilosa, caule
2-3 mm. crasso; folia lanceolato-elliptica vel ovato-elliptica acute
acuminata, basi subacuta vel latiora basi obtusa, versus basin
pinnatinervia, 4-11 cm. longa 2-4.5 cm. lata, petiolo 1.5-2.5 cm. longo
subpersistente pubescente; spicae terminales breves solitariae,
pedunculo aequilongo medio bracteato, bracteis rotundo-peltatis.

HONDURAS: Lancetilla Valley, alt. 30 meters, Yuncker 4544 (type
in herb. Univ. Illinois).

Peperomia chucanebana Trelease. On tree near Lake Yojoa,
at 630 meters.

Peperomia circulifolia Trelease, sp. nov. — Herba parvula plus
minusve repens caespitosa arboricola, caule 1-2 mm. crasso primo
velutino; folia saepe 5-6-verticillata elliptico-obovata emarginulata,
basi acuta, ca. 10 mm. longa atque 5 mm. lata, vix plus quam 1-
nervia, margine ciliata, ciliis rectis, subtus tantum transiente fla-
vescentia, subtus hispidula, supra minus dense hispidula, petiolo
ca. 2 mm. longo hispidulo; spicae terminales 20 mm. longae 1 mm.
crassae, pedunculo gracili 10 mm. longo sparse minute hispidulo;
bracteae rotundo-peltatae.

HONDURAS: In a wet ravine on a tree trunk near El Achote, above
Siguatepeque, Department of Comayagua, at 1,350 meters, Yuncker,
Dawson & Youse 6361 (type in herb. Univ. Illinois).

Peperomia circulifolia var. flava Trelease, var. nov. — Folia
magis obovata ca. 10 mm. longa et 8 mm. lata subtus subpersistente
opaco-flavescentia; planta terrestris.

HONDURAS: In the same region as the typical form of the species,
at 1,350 meters, Yuncker, Dawson & Youse 5869 (type in herb.
Univ. Illinois; duplicate in Herb. Field Mus.).

332 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Peperomia circulifolia var. eciliata Trelease, var. nov. — Var.
flavae valde similis, foliis 3-nerviis tenuioribus minus flavescentibus
non radiato-ciliatis.

HONDURAS: Region of the typical form of the species at 1,350
meters, Yuncker, Dawson & Youse 6360 (type in herb. Univ. Illinois).

Peperomia collocata Trelease, sp. nov. — Herba parvula
rhizomatoso-adsurgens glabra, caule 1 mm. crasso; folia alterna
elliptico-subspathulata plerumque obtusa, basi subcuneata, 3.5-7
cm. longa 1.5-3 cm. lata, ca. 5-nervia, nervis 10-15 mm. longe
confluentibus, in sicco subfirma, subtus pallidiora, petiolo gracili 5
mm. longo; spicae terminales et ex axillis superioribus nascentes 70
mm. longae 1 mm. crassae, pedunculo gracili fere 25 mm. longo;
bracteae rotundo-peltatae; ovarium oblique apicatum, stigmate sub-
apicali.

HONDURAS: On a login aravine at El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6106 (type in herb. Univ. Illinois).

Peperomia condormiens Trelease, sp. nov. — Herba modice
alta sed delicata subsimplex glabra, caule gracillimo; folia alterna
orbiculari-ovata emarginata basi rotundata vel ad petiolum abrupte
angustata, 1.5-2.5 cm. longa et aequilata, 5-7-nervia, nervis superior-
ibus in foliis longioribus e basi distincte separatis, supra intense
viridia, subtus pallidiora, petiolo gracili vix 10 mm. longo; spicae
terminales 70 mm. longae 1 mm. crassae, pedunculo 25 mm. longo;
bracteae rotundo-peltatae; ovarium ovoideum, stigmate apicali
stylum brevem terminante.

HONDURAS: Summit in forest above El Achote, above Siguate-
peque, Department of Comayagua, 1,800 meters, Yuncker, Dawson
& Youse 6237 (type in herb. Univ. Illinois; duplicate in Herb. Field

Mus.).

Peperomia conocarpa Trelease, sp. nov. — Herba modice parva
glabra rhizomatosa, ramis fertilibus erectis simplicibus, caule vix 2
mm. crasso sulcato; folia 3-4-verticillata elliptico-obovata emargi-
nata, basi cuneata, 6-12 mm. longa 4-8 mm. lata, in sicco viridia
subtenuia sed opaca, vix plus quam 1-nervia, petiolo 1-2 mm. longo;
spicae terminales subfiliformes 15-25 mm. longae, pedunculo fili-
formi aequilongo; bracteae rotundo-peltatae; baccae late conicae
pseudocupula basali stipatae, stigmate apicali stylum brevem
terminante.

FLORA OF HONDURAS 333

HONDURAS: On a tree trunk in forest, cloud zone above El Achote,
near Siguatepeque, Department of Comayagua, 1,800 meters,
Yuncker, Dawson & Youse 6189 (type in herb. Univ. Illinois; dupli-
cate in Herb. Field Mus.); also No. 6010.

Peperomia Dawsoni Trelease, sp. nov. — Herba subparva longi-
repens lignicola, caule 2 mm. crasso plus minusve minute hispidulo;
folia alterna orbicularia vel rotundo-ovata, rare sublanceolata, apice
rotundata vel obtuse breviter acuminata aliquanto emarginulata,
1-1.5 cm. longa et aequilata, obscure trinervia, coriacea, subtus
flavescentia, petiolo filiformi 2-7 mm. longo sparse minute hispidulo;
spicae terminales 25 mm. longae 1 mm. crassae, pedunculo vix 5
mm. longo, bracteis rotundo-peltatis; ovarii apex obliquus, stigmate
subapicali.

HONDURAS: On a rotten log in forest, cloud zone above El Achote,
near Siguatepeque, Department of Comayagua, 1,800 meters,
Yuncker, Dawson & Youse 6009 (type in herb. Univ. Illinois).

Peperomia defluens Trelease, sp. nov. — Herba majuscula
radicanti-pendula glabra, caule 2 mm. crasso; folia subelliptica vel
ovata acuminata, basi obtusa, 11 cm. longa 3-5 cm. lata, basin versus
pinnatinervia, petiolo 1 vel in foliis inferioribus 4 cm. longo; spicae
terminales statu fructifero 80-100 mm. longae 4 mm. crassae, pe-
dunculo 20-30 mm. longo prope medium bracteato; baccae anguste
oblongae oblique acuto-scutulatae, stigmate in scutulo centrali.

HONDURAS: In forest, Lancetilla Valley, 27 meters, Yuncker 4916
(type in herb. Univ. Illinois) ; also at 30 meters, Yuncker 1+630.

Peperomia eripipunctulata Trelease, sp. nov. — Herba sub-
parva stolonifero-adsurgens glabra non nigro-punctulata, faciem P.
rubefactae simulans, caule 2 mm. crasso; folia alterna lanceolata vel
elliptica subobtuso-acutata, basi acuta, 2.5-5 cm. longa 1-2 cm. lata,
3-nervia vel minus conspicue 5-nervia, nervis interioribus inferne
confluentibus, in sicco subcoriacea, subtus pallidiora, petiolo vix 10
mm. longo; spicae terminales vix 40 mm. longae et 2 mm. crassae,
pedunculo 5 mm. longo.

HONDURAS: On a rotten log in ravine near El Rincon, about 10
miles west of Siguatepeque, Department of Comayagua, 1,400-1,500
meters, Yuncker, Dawson & Youse 6079 (type in herb. Univ. Illinois) ;
in a ravine near El Achote, above Siguatepeque, 1,350 meters, No.
6104.

Peperomia filici-decorans Trelease, sp. nov. — Herba sub-
parva repenti-adsurgens arboricola, caule 2 mm. crasso transiente

334 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

minute crispo-puberulo; folia ternata vel quaternata lanceolata
obtuse protracto-acuminata, basi acuta, vix 2.5-3 cm. longa atque
1-2 cm. lata, 3-nervia, sparse crispo-puberula vel glabrescentia,
firma, subtus pallidiora, .petiolo 3-4 mm. longo; spicae terminales
20 mm. longae 1 mm. crassae, pseudo-pedicellis sparsis, pedunculo
gracili 15-20 mm. longo, bracteis rotundo-peltatis.

HONDURAS: On the trunk of tree fern, ravine near El Achote,
above Siguatepeque, Department of Comayagua, 1,350 meters,
Yuncker, Dawson & Youse 6105 (type in herb. Univ. Illinois).

Peperomia flagitans Trelease, sp. nov. — Herba subparva
simplex sed plus minusve caespitosa glabra, caulibus vix 2 mm.
crassis; folia alterna elliptico-subobovata apice obtusa et emarginu-
lata, basi cuneata, 2.5-3.5 cm. longa 1.5-2 cm. lata, 5-nervia, in sicco
coracea atque opaca, petiolo 5 mm. longo amplexicauli decurrente;
spicae terminales 60 mm. longae 2 mm. crassae, pedunculo 10-20
mm. longo.

HONDURAS: River bank 6 km. west of Siguatepeque, Department
of Comayagua, 1,200 meters, Yuncker, Dawson & Youse 6383 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Peperomia inaudax Trelease, sp. nov. — Herba modice parva
simplex erecta glabra, caule vix 2 mm. crasso; folia alterna elliptica
vel inferiora subobovata emarginulata, basi acuta, 2.5-3.5 cm. longa
1.5-2 cm. lata 1-nervia vel obscure 3-nervia, subtus pallidiora,
petiolo 5 mm. longo amplexicauli-decurrente; spicae terminales 50
mm. longae 2 mm. crassae, pedunculo 20 mm. longo, bracteis ro-
tundo-peltatae; baccae 1.5 mm. longae ovoideo-cylindraceae, pedi-
cellis in maturitate gracilibus aequilongis, stigmate in apice scutelli
oblique concavi inserto.

HONDURAS: On a log in forest, cloud zone above El Achote, near
Siguatepeque, Department of Comayagua, 1,850 meters, Yuncker,
Dawson & Youse 621*2 (type in herb. Univ. Illinois).

Peperomia lancetillana Trelease, sp. nov. — Herba modice
alta diffusa et radicans glabra, caule ca. 3 mm. crasso; folia alterna
obovata et obtusa vel subrhombica et subulato-apicata, basi cuneata,
5-9 cm. longa 2.5-3.5 cm. lata pinnatmervia, in sicco coriacea,
petiolo 1-2 vel 3.5 cm. longo; spicae terminales et sympodiales 70-
120 mm. longae 1-2 mm. crassae, pedicellis 2-3 cm. longis in apice
pedunculi 1-bracteati duplo longioris geminatis, bracteis minutis
rotundo-peltatis; baccae ellipsoideae rostratae, stigmate ad basin
rostri anteriore.

FLORA OF HONDURAS 335

HONDURAS: Lancetilla Valley, near Tela, 60 meters, Yuncker 4943
(type in herb. Univ. Illinois); at 30 meters, Yuncker 4629; at 75
meters, Yuncker 5084-

Peperomia lancetillana var. spathifolia Trelease, var. nov.
— Caulis brevissimus, internodiis brevibus; folia elliptico-spathulata
vel obovato-spathulata obtusa basi longe cuneata 7-10 cm. longa
2.5-4.5 cm. lata; spica solitaria 50-75 mm. longa 2 mm. crassa,
pedunculo longiore.

HONDURAS: Lancetilla Valley, 60 meters, Yuncker 5040 (type in
herb. Univ. Illinois).

Peperomia laudabilis Trelease, sp. nov. — Herba subparva
simplex stolonifera, foliis decidue puberulo-ciliatis exceptis glabra,
caule 1 mm. crasso; folia alterna elliptico-oblanceolata vel sub-
obovata apice rotundata vel subtruncata, basi subcuneata, 1.5-4.5
cm. longa 1-1.5 cm. lata, 2-4-nervia, in sicco viridia tenuia, petiolo
5 mm. longo; spicae terminales et axillares 50 mm. longae 1 mm.
crassae, pedunculo filiformi 20 mm. longo et ultra, bracteis rotundo-
peltatis; baccae ovoideae apice obliquae, stigmate subapicali.

HONDURAS: On a tree base, El Achote, near Siguatepeque, De-
partment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6298 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Peperomia longiacuminata Trelease. On a log in forest,
Lancetilla Valley, at 30 meters.

Peperomia mala Trelease. sp. nov. — Herba subparva rhizo-
matosa adsurgens glabra, caule 2-3 mm. crasso 4-lineato; folia
alterna elliptica subacuta, basi acuta, 20-55 mm. longa 10-23 mm.
lata obscurissime 3-5-nervia, coriacea, subtus pallidiora, petiolo 5-10
mm. longo amplexicauli-decurrente; inflorescentia?

HONDURAS: On a tree trunk, river bank, Siguatepeque, Depart-
ment of Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5737
(type in herb. Univ. Illinois).

Peperomia mollipubis Trelease, sp. nov. — Herba modice alta
adsurgens ubique gracili-pilosa, caule 1-3 mm. crasso; folia alterna
subrhombice elliptica vel obovata obtusa interdum obscure emar-
ginulata, basi acuta, 3.5-4 cm. longa 1.5-2.5 cm. lata 5-nervia, subtus
pallidiora, in sicco tenuia sed opaca, petiolo gracili 20-30 mm. longo;
spicae terminales et ex axillis superioribus nascentes in statu juvenili
25 mm. longae 2 mm. crassae, pedunculo brevi crasso, bracteis
rotundo-peltatis margine hispidis.

336 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

HONDURAS: In a ravine at El Achote, near Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6367
(type in herb. Univ. Illinois).

Peperomia montis-verticis Trelease, sp. nov. — Herba modice
alta adsurgens glabra, caule 2 mm. versus basin crasso; folia alterna
subelliptica obtuse breviter acuminata basi acuta, 4-7 cm. longa
2-3 cm. lata 5-nervia, in sicco subcoriacea, petiolo gracili circiter
10 mm. longo amplexicauli-decurrente; spicae 60 mm. longae 2 mm.
crasse apiculatae, pedunculo 5 mm. longo, bracteis rotundo-peltatis.

HONDURAS: Cloud zone near summit of the range above El
Achote, near Siguatepeque, Department of Comayagua, 1,850
meter, Yuncker, Dawson & Youse 6244 (type in herb. Univ. Illinois).

Peperomia nebuligaudens Trelease, sp. nov. — Herba parva
stolonifero-adsurgens glabra, caule 2-3 mm. crasso, internodiis
brevibus; folia alterna elliptica vel subobovata, majora subacuta,
basi acuta, 2-6 cm. longa 1-3 cm. lata 3-nervia, subtus pallidiora,
in sicco coriacea et opaca, petiolo 5-10 mm. longo amplexicauli-
decurrente; spicae terminales in statu juvenili 30 mm. longae 2-3 mm.
crassae, pedunculo 10 mm. longo, bracteis rotundo-peltatis; baccae
sunglobosae, stigmate in apice obliquo subapicali.

HONDURAS: Cloud region above El Achote, near Siguatepeque,
Department of Comayagua, 1,850 meters, Yuncker, Dawson &
Youse 6186 (type in herb. Univ. Illinois).

Peperomia novae-hispaniae Trelease. Near Paral, shore of
Lake Yojoa, Department of Cortes, at 630 meters.

Peperomia novella Trelease, sp. nov. — Herba modice alta
lignicola adsurgens, superne foliata et ramosa, caule 3 mm. basin
versus crasso subpersistente puberulo; folia vulgo ternata vel qua-
ternata subspathulata obtusa vel subtruncata, basi cuneata, 10-25
mm. longa 4-10 mm. lata, in sicco firma, subtus flavescentia, vix
plus quam 1-nervia, supra ad costam puberula, subtus sparse pu-
berula, petiolo 2-3 mm. longo puberulo; spicae immaturae, ter-
minales, bracteis rotundo-peltatis.

HONDURAS: On a rotten log in forest, cloud zone above El Achote,
near Siguatepeque, Department of Comayagua, 1,800 meters,
Yuncker, Dawson & Youse 6139 (type in herb. Univ. Illinois).

Peperomia opaca Trelease, var. ciliata Trelease, var. nov.—
Herba modice alta rhizomatosa caespitosa faciem P. reflexae simu-
lans, caule primo rigido-pubescente, foliis plus minusve rigido-

FLORA OF HONDURAS 337

ciliatis; spicae in statu juvenili 20 mm. longae 2 mm. crassae,
pedunculo 10-12 mm. longo rigido-pubescente, rhachide pubescente;
baccae subcylindraceae, pseudocupula basali, apice acutatae, stig-
mate apicali.

HONDURAS: Cloud zone above El Achote, near Siguatepeque,
Department of Comayagua, 1,850 meters, Yuncker, Dawson &
Youse 6187 (type in herb. Univ. Illinois); also at 1,800 meters, No.
6008; at 1,500 meters, No. 6078; and at 1,350 meters, No. 6366.

Peperomia perplexa Trelease. On tree near Lake Yojoa,
Department of Cortes, at 630 meters.

Peperomia petenensis Trelease, var. hondurensis Trelease,
var. nov. — Folia 7 cm. longa, 4 cm. lata et acuta, vel 12 cm. longa, 7
cm. lata et profunde emarginata; spicae juveniles 90 mm. longae
3 mm. crassae, in statu adulto ad 290 mm. longae.

HONDURAS: El Achote, near Siguatepeque, Department of Co-
mayagua, 1,350 meters, Yuncker, Dawson & Youse 5922 (type in
herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Peperomia praeteruentifolia Trelease, sp. nov. — Herba modice
alta adsurgens, caule basin versus 3 mm. crasso transiente puberulo,
superne ramoso; folia alterna, longiora rhombico-ovata, basi et
apice longe acuta, 2.5-6 cm. longa 1.5-4 cm. lata, 3-nervia vel
propter costam obscure ramosam obscure 5-nervia, in sicco mem-
branacea, plus minusve persistente sed sparse adpresso-pilosa,
petiolo filiformi puberulo ad 1-3 cm. longo; spicae axillares in statu
juvenili 30 mm. longae 1 mm. crassae, pedunculo filiformi puberulo
5 mm. longo, bracteis rotundo-peltatis.

HONDURAS: River bank near Siguatepeque, Department of
Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5614 (type in
herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Peperomia rhodophlebia Trelease, sp. nov. — Herba modice
alta epiphytica glabra, caule 3-4 mm. crasso in sicco valde angulato ;
folia opposita vel ternata rhombice lanceolato-elliptica acuminata,
basi cuneata, 25-40 mm. longa 15-18 mm. lata, in vivo rubro-venosa,
in sicco crassa dura opaca, petiolo 5 mm. longo; inflorescentia?

HONDURAS: On a tree near Potrerillos, Department of Cortes,
Yuncker 4896 (type in herb. Univ. Illinois).

Peperomia rinconensis Trelease, sp. nov. — Herba stolonifera,
ramis breviusculis foliosis minute puberulis circiter 1 mm. crassis;
folia alterna subrhombeo-lanceolata, obtuse protracto-acuminata,

338 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

basi acuta, 3.5 cm. longa 1.5 cm. lata 1-nervia vel obscure 3-5-nervia,
in sicco opaca subtus pallidiora, petiolo 2-4 mm. longo amplexicuali-
decurrente aliquanto sparse minute puberulo; spicae terminales 40-
50 mm. longae 1 mm. crassae, pedunculo vix 10 mm. longo, bracteis
rotundo-peltatis; ovarium apice obliquum, stigmate subapicali.

HONDURAS: On a rotten log in a wet ravine near El Rincon, west
of Siguatepeque, Department of Comayagua, 1,400-1,500 meters,
Yuncker, Dawson & Youse 6080 (type in herb. Univ. Illinois).

Peperomia rotundifolia (L.) HBK. On stones and trees in
forest above Lancetilla Valley, 60-90 meters.

Peperomia rubefacta Trelease, sp. nov. — Herba modice alta
rhizomatosa nigro-punctulata, caule 1-2 mm. crasso glabro vel
superne pilis paucis sparsis onusto; folia primo crassa, lucida vel
supra pilis paucis conspersa, lanceolata acuminata, basi acuta, 4-6
cm. longa 1.5-2 cm. lata, 5-nervia nervis interioribus basi aliquanto
confluentibus, in sicco opaca subcoriacea, petiolo 5-10 mm. longo
amplexicauli-decurrente plus minusve canaliculate, marginibus
aliquanto ciliatis; spicae 100 mm. longae 2 mm. crassae, pedunculo
gracili 10 mm. longo, bracteis rotundo-peltatis; baccae ovoideae
apice obliquae et plus minusve rostratae, stigmate subapicali.

HONDURAS: El Rincon, west of Siguatepeque, Department of
Comayagua, 1,400-1,500 meters, Yuncker, Dawson & Youse 6081
(type in herb. Univ. Illinois).

Peperomia suavis Trelease, var. hondurensis Trelease, var.
nov. — Characteribus speciei, odorem Zingiberis exhalans; folia ellip-
tico-ovata breviter acuminata, basi rotundata vel peltata, 16-19
cm. longa 10-13 cm. lata, ubique obscure nervosa, petiolo 9-12 cm.
longo; spicae 220 mm. longae 3-4 mm. crassae pedunculum bibrac-
teatum ca. 7 cm. longum solitarie terminantes; baccae rotundato-
ovoideae glandulosae rostratae.

HONDURAS: Ravine near El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 5878
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Peperomia subgeminispica Trelease, sp. nov. — Herba ma-
juscula diffusa radicans glabra rupicola, caule 3-4 mm. crasso; folia
alterna oblanceolato-obovata subobtuse acuminata, basi cuneata,
8.5-13 cm. longa 3.5-5.5 cm. lata, e tertio vel medio inferiore pin-
natinervia, petiolo 2-3 cm. longo; spicae terminales apice pedunculi
brevis geminatae 128 mm. longae 2 mm. crassae, pedunculo 1-2.5

FLORA OF HONDURAS 339

mm. longo, bracteis rotundo-peltatis parvis; stigma prope basin
scutelli longiacuminati insertum.

HONDURAS: On a rock, hills above Lancetilla Valley, at 90 meters,
Yuncker 4987 (type in herb. Univ. Illinois).

Peperomia tressis Trelease, sp. nov. — Herba modice alta repens
glabra, caule 2 mm. crasso; folia alterna late elliptica ad rotundo-
obovata vel subrhombeo-obovata, apice rotundata vel obtuse sub-
acuminata, basi acuta, 2-4 cm. longa 2-2.5 cm. lata, 3-nervia vel
obscure 5-nervia, subtus paullo pallidiora, in sicco chartacea,
petiolo ca. 10 mm. longo; spicae ex axillis superioribus nascentes
35 mm. longae 2 mm. crassae, pedunculo ca. 5 mm. longo.

HONDURAS: In a ravine near El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson &
Youse 6001 (type in herb. Univ. Illinois).

Peperomia Trumani Trelease, sp. nov. — Planta parva epi-
phytica rhizomatosa, caule filiformi, internodiis brevibus puberulis;
folia inferiora rotundata vel elliptica, superiora lanceolata atque
basi et apice acuta, 8 mm. longa et aequilata vel superiora 15-20 mm.
longa atque 5 mm. lata, 1-nervia vel obscure 3-nervia, puberula vel
glabrescentia, subtus pallida, petiolo vix 5 mm. longo; spicae termi-
nales vix 10 mm. longae et 1 mm. crassae, pedunculo 1 mm. longo.

HONDURAS: On a rotten tree trunk near Lake Yojoa, Department
of Cortes, 630 meters, Truman G. Yuncker 4847 (type in herb. Univ.
Illinois).

Peperomia yojoana Trelease, sp. nov.— Herba modice parva
repens radicans glabra, caule 1-2 mm. crasso; folia alterna lanceo-
lata ad subrhombeo-oblanceolata, apice attenuate subobtuso, basi
acuta vel cuneata, 1-3.5 cm. longa 0.5-1.25 cm. lata, 1-nervia vel
plus minusve obvie 3-5-nervia, coriacea et opaca, subtus flavescentia
et concolore vel obscure granulosa, petiolo vix 5 mm. longo amplexi-
cauli-decurrente; spicae vix 60-70 mm. longae et 1 mm. crassae
laxiflorae, pedunculo fere 2 cm. longo, bracteis rotundo-peltatis;
baccae subglobosae, stigmate in apice obliquo aliquanto acutato
subapicali.

HONDURAS: Lake Yojoa, Department of Corte"s, 630 meters,
Yuncker 4913 (type in herb. Univ. Illinois), 4823, 4902.

Peperomia Yousei Trelease, sp. nov. — Herba majuscula
stolonifero-adsurgens fragrans glabra, caule 4-5 mm. crasso; folia
alterna elliptica vel obovata obtusa vel fere truncato-emarginulata,

340 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

basi cuneata, 5-7.5 cm. longa 4-5 cm. lata graciliter pinnatinervia,
in sicco coriacea, petiolo 2-3.5 cm. longo; spicae pedunculos 1-2
graciles axillares 1-bracteatos 4-5 cm. longos terminantes, initio
anthesis 115 mm. longae et 2 mm. crassae, pedicellis ca. 2 cm.
longis sparse minute puberulis, bracteis magnis granulosis paucis
pseudo-verticillum efformantibus; baccae subcylindraceae, scutello
obliquo attenuato-acuminato, stigmate subapicali.

HONDURAS: In a wet ravine near El Achote, above Siguatepeque,
Comayagua, 1,350 meters, Yuncker, Dawson & Youse 5995 (type in
herb. Univ. Illinois), 6134- Duplicate of the latter number in Herb.
Field Mus.

Peperomia Yunckeri Trelease, sp. nov. — Herba modice alta
sed delicata aliquanto laxe ramosa, pagina superiore folii obscure
viridi excepta plus minusve rubescens, caule 1 mm. vel basin versus
ad 3 mm. crasso persistente minute puberulo; folia alterna ovata
vel ovali-ovata basi et apice acuta 1.5-2.5 cm. longa 1-1.5 cm.
lata multiplinervia, nervis basalibus 2-jugatis, addito prope medium
costae ramo alio, subtus albescentia, vulgo plicata, utrinque minute
puberula vel glabrescentia, petiolo gracili ca. 10-15 mm. longo
puberulo; spicae terminales et ex axillis superioribus nascentes
filiformes 60-100 mm. longae laxiflorae, pedunculo ca. 5-15 mm.
longo sparse puberulo; bracteae rotundo-peltatae; baccae rubrae
ellipsoideae, apice conico-obliquo, stigmate subapicali.

HONDURAS: Wet soil along bank of a stream northwest of Siguate-
peque, Department of Comayagua, 1,100 meters, Yuncker, Dawson
& Youse 6392 (type in herb. Univ. Illinois; duplicate in Herb. Field
Mus.),

PIPER L.

Piper actae Trelease, sp. nov. — Frutex 2-3-metralis nervis folii
subtus puberulis exceptis glaber, ramulis nigrescentibus, inter-
nodiis gracilibus modice elongatis; folia lanceolato-elliptica acute
acuminata, basi subobtusa, latere altero paullo breviore, 12-13 cm.
lata e medio inferiore pinnatinervia, nervis ca. 5x2, utrinque sub-
lucida, petiolo vix 8-12 mm. longo infra medium vaginante; spicae
50 mm. longae 3 mm. crassae, pedunculo 5 mm. longo.

HONDURAS: Tela, in sandy soil along the beach, Yuncker 4657
(type in herb. Univ. Illinois).

Piper adornatispicum Trelease, sp. nov. — Frutex 3-metralis
facie P. ornatispico arete similis, internodiis superioribus breviusculis
gracilibus primo cinereo-tomentosis; folia elliptica vel sublanceolata

FLORA OF HONDURAS 341

acute acuminata, basi oblique subcordulata, 16 cm. longa 5.5 cm.
lata, e medio inferiore pinnatinervia, nervis ca. 6x2, supra minute
pubescentia serius minute granuloso-asperata, nervis subtus apicem
versus puberulis sed basi pubescentibus, rugescentia, petiolo 5+3-
10+5 mm. longo pubescente; spicae 100 mm. longae 4 mm. crassae,
pedunculo 10 mm. longo pubescente; bracteae rotundo-subpeltatae
ciliolatae subzonatae.

HONDURAS: In a ravine at El Achote, above Siguatepeque,
Department of Comayagua, 1,050 meters, Yuncker, Dawson &
Youse 5885 (type in herb. Univ. Illinois; duplicate in Herb. Field
Mus.).

Piper allatum Trelease, sp. nov. — Frutex 1.5 m. altus, inter-
nodiis superioribus subgracilibus elongatis transiente puberulis;
folia lanceolata ad subrhombeo-elliptica acuminata, basi angustata
obtusa atque altero vel utroque latere cordulata, 15-20 cm. longa
5.5-10 cm. lata, e medio inferiore pinnatinervia, nervis ca. 5x2,
subtus plus minusve puberulis, petiolo ca. 8+2 mm. longo minute
pubescente; spicae in sicco viso valde juveniles et parvae.

HONDURAS: In a ravine at El Rincon, 10 miles west of Siguate-
peque, Department of Comayagua, 1,400-1,500 meters, Yuncker,
Dawson & Youse 6068 (type in herb. Univ. Illinois).

Piper apertum Trelease, sp. nov. — Frutex metralis, internodiis
floriferis gracilibus brevibus primo hispidis; folia subelliptica acute
acuminata, basi subcordulata altero latere paulo breviore, 12-13
cm. longa 6 cm. lata, e medio inferiore pinnatinervia, nervis ca.
5x2, supra albo-granulosa, nervis subtus minute adpresso-pubes-
centibus, petiolo 8+2 mm. longo subhispido; spicae rectae 60 mm.
longae 3 mm. crassae, pedunculo gracili fere 15 mm. longo glabrato,
bracteis transverse subpeltatis ciliolatis; baccae teretes truncatae,
stigmatibus 3 parvis sessilibus.

HONDURAS: Lake Yojoa, Department of Cortes, 630 meters,
Yuncker 4821 (type in herb. Univ. Illinois).

Piper apertum(?) var. nodosum Trelease, var. nov. — Nodosus;
folia basi angustata inaequaliter cordulata, nervis subtus hispidis;
inflorescentia?

HONDURAS: With the type, Yuncker 4821 in part (type in herb.
Univ. Illinois). ,

Piper arenicola Trelease, sp. nov.— Frutex vix 2-metralis,
internodiis modicis transiente hirsutis; folia elliptica acute acumi-

342 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

nata, basi aliquanto inaequaliter obtusa, 12-16 cm. longa 6-7 cm.
lata, e medio inferiors pinnatinervia, nervis ca. 5+4, vix rugosa,
supra glabra sublucida, subtus pallida subpuberula, nervis hirsutis,
petiolo vix 10 vel 5+5 mm. longo subhirsuto basi vaginante; spicae
vix 40 mm. longae et 3 mm. crassae, pedunculo 5 mm. longo puberulo;
bracteae parvae transverse subpeltatae ciliolatae.

HONDURAS: Sandy soil, Tela to Lancetilla Valley, at 27 meters,
Yuncker 4499 (type in herb. Univ. Illinois).

Piper auritum HBK. f. rinconense Trelease, forma nov.—
Frutex vix 1.5 m. altus, ramulis gracilibus cito glabrescentibus;
folia elongato-ovata subacuta, basi oblique auriculata, petiolo latere
altero paullo longiore, 22-25 cm. longa 11-12 cm. lata, nervis subtus
basin versus ut petiolo subvillosis, petiolo 50+2-5 mm. longo;
inflorescentia?

HONDURAS: El Rincon, Siguatepeque, Department of Comayagua,
1,400-1,500 meters, Yuncker, Dawson & Youse 6064 (type in herb.
Univ. Illinois).

Piper auritum HBK. var. amplifolium C. DC. Wet ravine
near El Achote, above Siguatepeque, Department of Comayagua,
at 1,350 meters.

Piper calvescens Trelease, sp. nov. — Internodia florifera sub-
gracilia brevia glabra ; folia ovata subacute acuminata, basi rotundata
vel acute contracta, modice parva 10-13.5 cm. longa 5-8 cm. lata,
palmatim 5-nervia vel obscure 7-nervia, nervis prope basin aliquanto
pubescentibus, petiolo brevi 5-10 vel ad 15 mm. longo, primo
aliquanto villoso glabrescente; spicae oppositifoliae graciles sed sub-
longae 60-70 mm. longae 3^4 mm. crassae, pedunculo gracillimo
brevi 10 mm. longo sed petiolum oppositum superante aliquanto
pubescente; bracteae subpeltatae ciliolatae; flores sessiles perfecti;
ovarium glabrum, stigmatibus 3 sessilibus.

HONDURAS: San Pedro Sula, Department of Santa Barbara,
Thieme 5461 (type in U. S. Nat. Herb. No. 796,474 and, in part, No.
796,493).

Piper calvescens var. pedunculatum Trelease, var. nov. —
Folia rotundo-elliptica subabrupte breviter acuminata, basi acuta,
13-14 cm. longa 9-10 cm. lata, 7-nervia, petiolo et pedunculo 10-15
mm. longis; spicae in statu fructifero 75 mm. longae 5 mm. crassae.

HONDURAS: Potrerillos, Department of Cortes, Yuncker 4882
(type in herb. Univ. Illinois).

FLORA OF HONDURAS 343

Piper calvescens var. portrerillosense Trelease, var. nov. —
Gracilis nodosus nigrescens essentialiter glaber; folia elliptica
graciliter acuminata, basi acuta, 7-10 cm. longa 4-6 cm. lata, 5-
nervia, petiolo 5-10 mm. longo; spicae 50-70 mm. longae 5 mm.
crassae densiflorae, pedunculo 5 mm. longo; bracteae subcrescenticae
griseae; baccae ovoideae, stigmatibus parvis erectis.

HONDURAS: Potrerillos, Department of Cortes, Yuncker 4894
(type in herb. Univ. Illinois).

Piper comayaguanum Trelease, sp. nov. — Frutex 2-metralis,
internodiis gracilibus cito elongatis sordido-villosis; folia rotundo-
ovata aromatica acuminata, basi rotundata vel inferiora cordata,
12-19 cm. longa 9-14 cm. lata, 5-7-nervia, primo supra pubescentia,
nervis subtus pallidis hispidis, petiolo 10-15 mm. longo vel in foliis
inferioribus ad 28 mm. subvilloso; spicae in siccis juveniles, pe-
dunculo sordido-subvilloso.

HONDURAS: River bank about 6 km. north of Siguatepeque, De-
partment of Comayagua, 1,300 meters, Yuncker, Dawson & Youse
6310 (type in herb. Univ. Illinois).

Piper cristicola Trelease, sp. nov. — Frutex vel suffrutex 2-
metralis, internodiis floriferis subgracilibus sed elongatis hirtellis sed
ad et infra nodos glabrescentibus; folia ovata obtuse sed apiculate
breviacuminata, basi fere aequilaterali cordata, 33 cm. longa 19-20
cm. lata, e tertio infimo subpinnatinervia, nervis longioribus ca.
6x2, intermediis interpositis, supra glabra viridia lucida, subtus
pallidiora primo ad nervos hirtella, petiolo 5 cm. longo rigido-
hirtello; inflorescentia?

HONDURAS: In forest at summit of the range above El Achote,
near Siguatepeque, Department of Comayagua, 1,800 meters,
Yuncker, Dawson & Youse 6247 (type in herb. Univ. Illinois).

Piper Dawsoni Trelease, sp. nov. — Frutex vix 2-metralis, ramis
gracilibus, internodiis elongatis evanescente villosis; folia lanceolata
sensim subacuminata, basi obtusa, 10-19 cm. longa 3-4 cm. lata, e
medio inferiore pinnatinervia, nervis 4-5x2, supra basi bus pilorum
brevium granuloso-scabra, subtus ad nervos subvillosa, petiolo 5 mm.
longo subvilloso; spicae rectae apiculatae 80 mm. longae 3 mm.
crassae, pedunculo 10 mm. longo villoso; bracteae rotundo-sub-
peltatae ciliatae.

HONDURAS: In a ravine at El Achote, above Siguatepeque, De-
partment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6283 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

344 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

Piper Deamii Trelease, sp. nov. — P. decrescens var. ovatum C.
DC. in DC. Prodr. 16, pt. 1: 251. 1869. Frutex ca. 2-metralis
velutino-hirsutus, internodiis floriferis gracilibus brevibus; folia
rotundo-ovata breviter acuminata, basi aperte cordata et intra
sinum breviter cuneata, 8-10 cm. longa 6-8 cm. lata, 7-9-nervia,
petiolo 10 mm. longo; spicae ca. 30 mm. longae et 2 mm. crassae,
pedunculo 5 mm. longo.

NICARAGUA: "Guatemala," Friedrichstahl (type in Herb. Kew.);
also in herb. Mus. Vienna as No. 905, from Granada. Known also
from Salvador. [The locality is evidently Nicaraguan rather than
Guatemalan, as listed in Dr. Trelease's original manuscript. Ap-
parently all the Friedrichsthal plants at Vienna are labeled as from
Guatemala, but fortunately the locality name usually is written on
the label, from which it appears that a large number if not the
majority of them are really from Nicaragua and Costa Rica.—
P. C. Standley.]

Piper Deamii var yojoanum Trelease, var. nov. — Frutex
nodosus 2-metralis, internodiis floriferis subgracilibus brevibus
primo minute crispo-pubescentibus; folia ovata acuminata, basi
cordulata vel breviter cordata, 7.5-9 cm. longa 4.5-5.5 cm. lata, 7-
nervia, nervis praesertim subtus hirtellis, petiolo 10 mm. longo
crispo-hirtello; spicae 35 mm. longae 5 mm. crassae, pedunculo 10
mm. longo plus minusve hirtello; baccae ovoideae, stigma tibus
brevibus.

HONDURAS: Paral, Lake Yojoa, 630 meters, Yuncker 4875 (type
in herb. Univ. Illinois).

Piper deltoideocarpum Trelease, sp. nov. — Frutex 3-metralis,
internodiis superioribus subgracilibus brevibus breviter molliter
pubescentibus; folia ovata sensim subacuminata, basi paullo in-
aequilatera obtusa, ca. 16 cm. longa et 9 cm. lata, e tertio infimo
pinnatinervia, nervis ca. 5x2, supra primo minute pubescentia
serius paullo asperata, nervis subtus crispo-puberulis, petiolo 1-2
cm. longo; spicae 80 mm. longae 3 mm. crassae, pedunculo 5 mm.
longo; aliquanto pubescente;bracteae triangulari-subpeltatae ciliatae;
baccae plus minusve acute triquetrae.

HONDURAS: River bank near Siguatepeque, Department of
Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5654 (type in
herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper epilosipes Trelease, sp. nov. — Frutex nodosus glaber vix
2-metralis facie spec. P. atrichopus similis, internodiis superioribus

FLORA OF HONDURAS 345

brevibus gracilibus; folia anguste lanceolata sensim attenuata, basi
angustata obvie obtusa, 10-12 cm. longa 3-4 cm. lata, e tertio infimo
pinnatinervia, nervis longioribus 3x2, nervis brevibus sursum curvis
fere ad apicem insertis, in sicco chartacea et paullo undata, petiolo
5 mm. longo vel in foliis inferioribus 10 mm. longo; spicae 40 mm.
longae 2 mm. crassae, pedunculo 5 mm. longo; bracteae subcrescenti-
cae; baccae depressae subrotundae, stigmatibus sessilibus.

HONDURAS : In a ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6296
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper epilosipes var. heterophyllum Trelease, var. nov. —
Folia majora, 12-15 cm. longa 3-4 cm. lata, inferiora ovata atque
10 cm. longa et 4 cm. lata, petiolis infimis 15 mm. longis; spicae 50
mm. longae 3 mm. crassae, pedunculo 5-10 mm. longo.

HONDURAS: With the type, Yuncker, Dawson & Youse 6097 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper exigens Trelease, sp. nov. — Frutex glaber vix 2.5 m. altus,
internodiis superioribus gracilibus breviusculis; folia lanceolato-
ovata acuminata, basi subobliqua acuta vel abrupte acutiuscula,
16-19 cm. longa 7.5-8 cm. lata, multiplinervia, nervis basalibus 2
paribus, utroque latere infra tertium supremum nervis 3 separatis
additis, lucida, subcoriacea, petiolo 15 mm. longo; spicae 100 mm.
longae 3 mm. crassae, pedunculo 10 mm. longo.

HONDURAS: Ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6124
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper exigens var. subovatum Trelease, var. nov. — Folia
elongato-ovata ad 20 cm. longa atque 8.5 cm. lata, vel late ovata
atque 9-12 cm. lata; spicae 110 mm. longae.

HONDURAS: With the type, Yuncker, Dawson & Youse 5997 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper faviculifemm Trelease, sp. nov. — PI. XI. Frutex ali-
quanto nodosus 1.5 m. altus, internodiis subgracilibus elongatis
subpersistente molliter cinereo-pubescentibus; folia ovata subacumi-
nata, basi breviter cordata, 12-20 cm. longa 6-13 cm. lata infra
tertium supremum pinnatinervia, nervis ca. 6-7x2, bullato-
alveolata, supra granuloso-scabra, nervis subtus breviter pubescenti-
bus, petiolo 15-20 mm. longo griseo-pubescente; spicae paullo curvae
75 mm. longae 2 mm. crassae, pedunculo 8 mm. longo; bracteae
subcrescenticae dense pilosae.

346 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

HONDURAS: Ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6368
(type in herb. Univ. Illinois).

Piper firmifolium Trelease, sp. nov. — Frutex nodosus glaber
3-4-metralis, internodiis superioribus brevibus gracilibus; folia
lanceolata obtuse subacuminata, basi acuta, 6-8 cm. longa 2-3.5 cm.
lata, saliente 3-nervia, foliis interdum ovatis, basi rotundato-cordatis,
8 cm. longis, 4 cm. latis atque 5-nervis, subtus paullo pallidiora, in
sicco firma, petiolo 3 mm. longo; spicae vix 50 mm. longae atque 2
mm. crassae, pedunculo 5-10 mm. longo; bracteae rotundo-sub-
peltatae lanosae; baccae rotundae, stigmatibus sessilibus.

HONDURAS: Near El Rincon, about 10 miles west of Siguatepeque,
Department of Comayagua, 1,400-1,500 meters, Yuncker, Dawson
& Youse 604-8 (type in herb. Univ. Illinois; duplicate in Herb. Field

Mus.).

Piper fraguanum Trelease, var. viride Trelease, var. nov.—
Folia 17-21 cm. longa 8-11 cm. lata subtus laete viridia; spicae 75
mm. longae 4 mm. crassae.

HONDURAS: Lancetilla Valley, at 27 meters, Yuncker 4918 (type
in herb. Univ. Illinois).

Piper inconsumptum Trelease, sp. nov. — Frutex 2-3-metralis
glaber vel in nervis paginae inferioris folii puberulus, internodiis
breviusculis gracilibus; folia lanceolato-elliptica plus minusve falcate
breviacuminata, basi acuta latere altero solemniter breviore, 16-18
cm. longa 6-7 cm. lata, infra tertium supremum pinnatinervia, nervis
5x2, subtus pallidiora, petiolo fere 15+5 mm. longo basi tantum
vaginante; spicae suberectae 90-125 mm. longae 3-4 mm. crassae
pallido-zonatae, pedunculo 10 mm. longo; bracteae crescenticae vel
triangulari-subpeltatae ciliolatae; baccae subteretes truncatae, stig-
matibus parvis sessilibus.

HONDURAS: Forest near Paral, Lake Yojoa, Department of Corte*s,
630 meters, Yuncker 4871 (type in herb. Univ. Illinois), 4874-

Piper laceratibracteum Trelease, sp. nov. — Frutex glaber
nodosus ca. 2-metralis, ramis nigrescentibus, internodiis gracilibus
striatis cito elongatis; folia lanceolata obtuse subacuminata, basi
acuta, 9-10.5 cm. longa 3.5-5 cm. lata, 3-nervia vel obscurissime
5-nervia, subtus paullo pallidiora, in sicco crasse papyracea, petiolo
vix 5 mm. longo; spicae 50-90 mm. longae 2 mm. crassae, pedunculo
filiformi 10 vel usque ad 25 mm. longo; bracteae triangulari-subpel-

FLORA OF HONDURAS 347

tatae, medio fuscae, margine lacerate pallido; ovarium depressum,
stigmatibus sessilibus.

HONDURAS: Ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6226
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper laedans Trelease, sp. nov. — Frutex 1.5 m. altus, ramis
geniculatis nodosis, internodiis brevibus gracilibus primo valde
ferrugineo-villosis; folia lanceolata anguste acuminata, basi angustata
obtusa latere altero breviore, 13-17 cm. longa 4.5-5.5 cm. lata, e
medio inferiore pinnatinervia, nervis 4+3, albido-granuloso-scabra,
nervis subtus villosis, petiolo 5-7+3 mm. longo villoso; spicae 60 mm.
longae 3 mm. crassae fere rectae, pedunculo 10 mm. longo ferrugineo-
villoso.

HONDURAS: Ravine near El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 5821
(type in herb. Univ. Illinois).

Piper monteluctans Trelease, sp. nov. — Frutex humilis plus
minusve caespitosus glaber, internodiis superioribus gracillimis
breviusculis; folia late lanceolata obtuso-acuminata, basi uno latere
obtusa altero acute protracta, 12-16 cm. longa 4-6 cm. lata, e medio
inferiore pinnatinervia, nervis ca. 4x2, in sicco tenuia viridia,
petiolo 3-5+2 mm. longo; inflorescentia?

HONDURAS: Creeping over logs in forest, cloud zone on range
above El Achote, near Siguatepeque, Department of Comayagua,
1,800 meters, Yuncker, Dawson & Youse 6176 (type in herb. Univ.
Illinois; duplicate in Herb. Field Mus.).

Piper multinervium Mart. & Gal. Artanthe Galeottii Miq.;
P. aduncum, P. angustifolium, P. elongatum of authors, as to
Mexico and northern Central America; P. xanthocladum Trelease
ex Standl. & Calderon, Lista PI. Salv. 62. 1925, nomen. River bank,
vicinity of Siguatepeque, 1,050 meters; also at 1,200 meters.

Piper multinervium var. amplum Trelease, var. nov. —
Frutex evanescente puberulus; folia elliptico-lanceolata vix cordu-
lata, 20 cm. longa 8 cm. lata, petiolo ca. 5+3 mm. longo; spicae
curvae 90 mm. longae 2-3 mm. crassae, pedunculo 8-10 mm. longo.

GUATEMALA: La Libertad, Pet£n, Lundell 2556 (type in herb.
Univ. Michigan).

Piper multinervium var. cayoense Trelease, var. nov.—
Frutex 1.5 m. altus persistente aliquanto puberulus; folia elliptico-

348 P'IELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

lanceolata acuminata, basi inaequaliter cordulata, 18 cm. longa 6-7
cm. lata, petiole 3+2 mm. longo; spicae curvae fere 150 mm.
longae 2-3 mm. crassae, pedunculo ca. 10 mm. longo.

BRITISH HONDURAS: El Cayo, H. H. Bartlett 11570 (type in herb.
Univ. Michigan).

Piper multinervium var. flavicans Trelease, var. nov.—
Frutex nodosus 3-metralis, internodiis in statu juvenili brevibus et
modice gracilibus rubescentibus dense velutinis; folia lanceolata
sensim longiattenuata, basi inaequaliter obtusa, latere longiore ad
petiolum acute decurrente, ca. 10-12 cm. longa atque 4 cm. lata,
nervis adpresso-puberulis, petiolo 4+4 mm. longo primo velutino;
spicae curvae 90 mm. longae 3 mm. crassae, pedunculo 15 mm.
longo glabro.

HONDURAS: Siguatepeque, Department of Comayagua, 1,200
meters, Yuncker, Dawson & Youse 5859 (type in herb. Univ. Illinois;
duplicate in Herb. Field Mus.).

Piper multinervium var. hirsuticaule Trelease, var. nov.—
Frutex 4-metralis, ramulis primo molliter hirsutis; folia lanceolata
acute acuminata, basi paullo obliqua cordulata, 15-20 cm. longa
4.5-5.5 cm. lata, petiolo 5+1 mm. longo molliter piloso; spicae
curvae 50 mm. longae 2 mm. crassae, pedunculo 15 mm. longo sparse
velutino.

HONDURAS: Siguatepeque, Department of Comayagua, 1,050
meters, Yuncker, Dawson & Youse 5724 (type in herb. Univ. Illinois;
duplicate in Herb. Field Mus.); also, at 1,200 meters, No. 5699.

Piper multinervium var. kantelolense Trelease, var. nov.—
Frutex valde nodosus transiente aliquanto puberulus, internodiis
brevibus; folia lanceolata acuminata, basi inaequaliter obtusa vel
subcordulata, 13-15 cm. longa 4-5 cm. lata, petiolo ca. 4+4 mm.
longo; spicae curvae 100 mm. longae 2-3 mm. crassae, pedunculo
10 mm. longo.

GUATEMALA: La Libertad, Pete"n, Lundell 3008 (type in herb.
Univ. Michigan). Also at Kantelol, Lundell 3168.

Piper multinervium var. paralense Trelease, var. nov.—
Arbuscula nodosa 5-metralis, trunco basi 5 cm. diam., ramulis
evanescente puberulis serius minute infra nodos granulosis; folia
elliptico-lanceolata graciliter acuminata, basi uno latere tantum
cordulata, 15-19 cm. longa 5-6 cm. lata, petiolo 3+3 mm. longo;
spicae curvae 110 mm. longae 3 mm. crassae, pedunculo 10-55 mm.
longo glabrato.

FLORA OF HONDURAS 349

HONDURAS: Paral, Lake Yojoa, Department of Corte"s, 630
meters, Yuncker 4877 (type in herb. Univ. Illinois).

Piper multinervium var. peracutum Trelease, var. nov.—
Frutex 2-3-metralis transiente velutinus, nodis denudatis minute
granulosis; folia late lanceolata vel elliptico-lanceolata sensim longi-
acuminata, basi inaequaliter cordulata, 20 cm. longa 7-7.5 cm. lata,
petiolo 4+3 mm. longo; spicae curvae 100 mm. longae 2 mm. crassae,
pedunculo 12 mm. longo glabrato.

HONDURAS: Lancetilla Valley, near Tela, Department of Atlan-
tida, 20-600 meters, Paul C. Standley 54921 (type), 53083.

Piper multinervium var. productipes Trelease, var. nov.—
Frutex nodosus 3-metralis transiente aliquanto velutinus; folia
lanceolata acuminata, basi inaequilaterali latere longiore ut in typo
subauriculato-cordulato, 17 cm. longa 5.5 cm. lata; spicae curvae
90-100 mm. longae 3 mm. crassae, pedunculo 15 mm. longo.

HONDURAS : Ravine near El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 5819
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper multinervium var. protractifolium Trelease, var. nov.
- — Frutex vix 3-metralis, ramulis glabratis purpurascentibus; folia
elliptica atque 15 cm. longa et 5.5 cm. lata, vel elongata, 18 cm.
longa 5 cm. lata, acute acuminata, basi subobtusa latere altero
breviore, nervis subtus aliquanto tomentulosis, petiolo 3+3 mm.
longo glabrato; spicae curvae 100 mm. longae 2 mm. crassae,
pedunculo gracili 15 mm. longo glabrato.

HONDURAS: Pine forest above Siguatepeque, Department of
Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5668 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper multinervium var. puberulipedunculum Trelease,
var. nov. — Frutex altus 5-metralis, trunco 7.5 cm. diam., ramulis
transiente adpresso-pilosis; folia subovato-lanceolata acuminata,
basi obliqua obtusa latere breviore subcordulato, 14-19 cm. longa
5-7 cm. lata, nervis puberulis, petiolo ca. 3+3 mm. longo glabrato;
spicae curvae vix 90 mm. longae et 2 mm. crassae, pedunculo gracili
velutino 6-10 mm. longo.

HONDURAS: El Achote, above Siguatepeque, Department of
Comayagua, 1,350 meters, Yuncker, Dawson & Youse 5876 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

350 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Piper multinervium var. pubescenticaule Trelease, var.
nov. — Arbor 10-metralis in statu juvenili rubescens, ramulis primo
dense subtomentosis; folia elongato-lanceolata acuminata basi
oblique subacuta, 12-20 cm. longa 4-6 cm. late, aliquanto ruges-
centia, nervis subtus prominentibus pubemlis, petiolo 5+2 mm.
longo; spicae ca. 50 mm. longae et 2 mm. crassae, pedunculo velutino
10 mm. longo.

HONDURAS: Siguatepeque, Department of Comayagua, 1,050
meters, Yuncker, Dawson & Youse 5731 (type in herb. Univ. Illinois;
duplicate in Herb. Field Mus.).

Piper multinervium var. Skutchii Trelease, var. nov. —
Frutex 5-metralis evanescente sparse sericeus; folia late lanceolata
17-26 cm. longa 5.5-7.5 cm. lata basi inaequaliter cordulata, petiolo
vix 3+3 mm. longo; spicae 90 mm. longae 2-3 mm. crassae, pedun-
culo 15 mm. longo.

GUATEMALA: Colomba, Department of Quezaltenango, 900 meters,
A. F. Skutch 1299.

Piper multinervium var. tamashense Trelease, var. nov. —
Frutex 2-metralis, trunco 3 cm. crasso, evanescente puberulus; folia
lanceolata vel elliptico- vel subovato-lanceolata acuminata, basi
inaequaliter cordulata, 15-18 cm. longa 4.5-7 cm. lata, petiolo ca.
5+5 mm. longo; spicae in sicco viso juveniles et brevipedunculatae.

BRITISH HONDURAS: Tamash River, 30 meters, W. A. Schipp 1382.

Piper multinervium var. telanum Trelease, var. nov.—
Frutex 2-3-metralis subpersistente puberulus; folia late lanceolata
vel elliptico-lanceolata acuminata, basi inaequaliter cordulata, 16-18
cm. longa 5-6.5 cm. lata subrugescentia, petiolo ca. 3+3 mm. longo
glabrato; spicae curvae 90 mm. longae 2-3 mm. crassae, pedunculo
10 mm. longo puberulo.

HONDURAS: Tela, Department of Atlantida, 20-600 meters, Paul
C. Standley 53264 (type in Herb. Field Mus.); Tela River, Yuncker
4512; Tela to Lancetilla Valley, 30 meters, Yuncker 4498 (folia 16
cm. tantum longa).

Piper mystaciferum Trelease, sp. nov. — Frutex 2-metralis,
internodiis superioribus modice gracilibus et elongatis primo dense
curvo-pilosis; folia lanceolata sensim subacuminata, basi obtusa
latere altero breviore, 15-18 cm. longa 5-6 cm. lata e medio inferiore
pinnatinervia, nervis 4X2 venis transversis connexis, supra dense
adpresso-pubescentia et asperata, costa barbata, petiolo ca. 6+4 mm.

FLORA OF HONDURAS 351

longo piloso; spicae in sicco viso juveniles atque 35 mm. longae, 2
mm. crassae, pedunculis pilosis 10 mm. longis.

HONDURAS: Ravine at El Rincon, 10 miles west of Siguatepeque,
Department of Comayagua, 1,300-1,400 meters, Yuncker, Dawson

6 Youse 6054 (type in herb. Univ. Illinois; duplicate in Herb. Field

Mus.).

Piper nubigaudens Trelease, sp. nov. — Frutex vix 3-metralis,
internodiis superioribus gracilibus substrigosis brevibus; folia lan-
ceolata sensim acuminata, basi paullo inaequilaterali obtusa, 9-11
cm. longa 3.5-4 cm. lata, e medio inferiore pinnatinervia, nervis ca.
3X2, glabra, in sicco chartacea fragrantia, petiolo vix 5 mm. longo
substrigoso; spicae 55 mm. longae 3-4 mm. crassae, pedunculo ca.

7 mm. longo; bracteae crescenticae; baccae depressae cum rhachi
elongatae, stigmatibus sessilibus.

HONDURAS: Cloud zone forest above El Achote, near Siguatepeque,
Department of Comayagua, 1,800 meters, Yuncker, Dawson & Youse
5991 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper ornatispicum Trelease, sp. nov. — Frutex 3-metralis,
internodiis floriferis breviusculis modice crassis primo griseo-tomen-
tosis; folia elliptica subacuminata, basi inaequilateralia uno latere
obtusa, 14-16 cm. longa 7-8 cm. lata, e medio inferiore pinnatinervia,
nervis 5X6, primo supra minute pubescentia serius granuloso-aspe-
rata, subtus puberula, non rugosa, petiolo 10 mm. vel 15+5 mm.
longo primo pallido-tomentoso; spicae 80 mm. longae 3 mm. crassae,
pedunculo 5 mm. longo pubescente; bracteae triangulari-subpeltatae
pallidae, ovariis fuscis.

HONDURAS: Thicket on river bank near Siguatepeque, Depart-
ment of Comayagua, 1,050 meters, Yuncker, Dawson & Youse 5618
(type in herb. Univ. Illinois).

Piper paludicola Trelease, sp. nov. — Frutex 3-metralis inferne
nodosus, internodiis floriferis brevibus modice gracilibus primo
molliter pilosis serius breviter velutinis; folia lanceolato-elliptica vel
ovato-elliptica acute acuminata, basi aliquanto inaequaliter cor-
dulata, 11-17 cm. longa 5-7 cm. lata, e medio inferiore pinnatinervia,
nervis 4-5x2, aliquanto rugosa, supra valde lucida obscure viridia
glabra, subtus pallidiora subpuberula, nervis breviter hirsutis,
petiolo 10-15 mm. longo subhirsuto infra medium vaginante; spicae
50-60 mm. longae 3-4 mm. crassae, pedunculo 5 mm. longo glabrato;
baccae rotundae griseae.

352 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

HONDURAS: In marsh near Tela, Department of Atlantida, Yun-
cker 1+977 (type in herb. Univ. Illinois).

Piper petens Trelease, sp. nov. — Frutex paullo plus quam 2 m.
altus petiolis et nervis subtus obscure puberulis exceptis glaber,
internodiis gracilibus primo brevibus; folia elongato-elliptica acute
acuminata, basi paullo inaequilateralia et vulgo acuta, 11-18 cm.
longa 5.5-7 cm. lata e medio inferiore pinnatinervia, nervis 5+4, in
sicco papyracea, petiolo 10+3 mm. longo; spicae in sicco viso juve-
niles vix 10 mm. longae et 1 mm. crassae apiculatae griseae breviter
pedunculatae.

HONDURAS: Ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6107
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper pictamentum Trelease, sp. nov. — Frutex nodosus 1-2-
metralis, internodiis crassis brevibus pallidis pallido-tomentosis;
folia elliptica vel oblongo-elliptica acuta, basi inaequaliter cordulata,
6-12.5 cm. longa 3-5.5 cm. lata, supra manifeste granulosa, subtus ad
nervos pallido-villosa, e medio inferiore pinnatinervia, nervis 6x2,
petiolo 5 mm. longo tomentoso-subvilloso; spicae rectae vel curvae
80 mm. longae 3 mm. crassae, pedunculo 10 mm. longo tomentoso;
bracteae rotundo-subpeltatae ciliatae pallidae et manifeste ut in P.
Yunckeri zonatae.

HONDURAS: Rocky hillside, Siguatepeque, Department of Co-
mayagua, 1,100 meters, Yuncker, Dawson & Youse 5902 (type in
herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper pictamentum var. fluminis Trelease, var. nov. —
Forma multo gracilior minus tomentosa, aliter ut in forma typica
speciei.

HONDURAS: River bank, Siguatepeque, Department of Coma-
yagua, 1,050 meters, Yuncker, Dawson & Youse 5718 (type in herb.
Univ. Illinois; duplicate in Herb. Field Mus.).

Piper praemollitum Trelease, sp. nov. — Frutex vix 3-metralis,
internodiis superioribus gracilibus brevibus primo tomentulosis;
folia ovato-elliptica acuminata, basi rotundata altero latere paullo
majore, 15-17 cm. longa 6.5-9 cm. lata e medio inferiore pinnati-
nervia, nervis ca. 6+5, supra puberula, subtus pallidiora aliquanto
tomentulosa, petiolo ca. 10+2 mm. longo tomentuloso; spicae 70 mm.
longae 2 mm. crassae, pedunculo vix 5 mm. longo puberulo.

HONDURAS: Near shore of Lake Yojoa, Department of Cortes,
630 meters, Yuncker 4825 (type in herb. Univ. Illinois).

FLORA OF HONDURAS 353

Piper rinconense Trelease, sp. nov. — Frutex vix 3-metralis,
faciem P. Yousei simulans, internodiis superioribus subgracilibus
brevibus primo subsericeis non nigrescentibus; folia elliptica vel
sublanceolata acuminata, basi inaequilaterali altero vel ambobus
lateribus rotundata, 14-15 cm. longa 6 cm. lata e medio inferiore
pinnatinervia, nervis 5+6, subtus puberula, petiolo 5-7+3 mm.
longo aliquanto pubescente; spicae in sicco viso juveniles vix 15 mm.
longae et 2 mm. crassae, pedunculo curvo glabrato vix 5 mm. longo.

HONDURAS : Ravine near El Rincon, about 10 miles west of Sigua-
tepeque, Department of Comayagua, 1,400-1,500 meters, Yuncker,
Dawson & Youse 6052 (type in herb. Univ. Illinois).

Piper roseovenulosum Trelease, sp. nov. — Frutex vix 3-
metralis, internodiis floriferis graciusculis brevibus primo griseo-
pubescentibus cito glabrescentibus; folia lanceolato-elliptica sub-
falcate longiacuminata, basi inaequilateralia latere breviore saepe
acuto, 10-15 cm. longa 4-8 cm. lata e medio inferiore pinnatinervia,
nervis 4x2, subtus griseo-pubescentia, supra lepidoto-asperata et
subtus scabrida, aliquanto rugosa, subtus pallidiora, venis in vivo
roseis, petiolo ca. 6+4 mm. longo griseo-subscabrido; spicae 70 mm.
longae 3 mm. crassae, pedunculo vix 10 mm. longo scabrido.

HONDURAS: Near Paral, shore of Lake Yojoa, Department of
Cortes, 630 meters, Yuncker 4873 (type in herb. Univ. Illinois).

Piper scabriseptum Trelease, var. reductum Trelease, var.
nov. — Caule magis glabrescente; folia 12-14 cm. longa 5.5-7 cm.
lata, petiolo ad 10 mm. longo.

HONDURAS: Lancetilla Valley, Department of Atlantida, 27
meters, Yuncker 4773 (type in herb. Univ. Illinois).

Piper scalarispicum Trelease, sp. nov. — Frutex subnodosus
ramulosus vix 2-metralis, internodiis floriferis graciusculis brevibus
primo velutinis; folia anguste lanceolata longiattenuata, basi plus
minusve obtusa, 10-13 cm. longa 3-3.5 cm. lata e medio inferiore
pinnatinervia, nervis longioribus ca. 3x2, subtus crispo-velutina,
petiolo vix 5 mm. longo; spicae 60-80 mm. longae 4 mm. crassae,
pedunculo 10 mm. longo puberulo; bracteae triangulari-subpeltatae
ciliatae scalariformi-zonatae; baccae depressae rotundatae cum
rhachi elongatae, stigmatibus sessilibus.

HONDURAS: Cloud zone at summit of the range above El Achote,
near Siguatepeque, Department of Comayagua, 1,800-2,000 meters,
Yuncker, Dawson & Youse 6156 (type in herb. Univ. Illinois; dupli-
cate in Herb. Field Mus.)..

354 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Piper siguatepequense Trelease, sp. nov. — Frutex divaricate
ramosus 2.5 m. altus, internodiis superioribus gracilibus breviusculis
persistente molliter velutinis; folia elliptico- vel rhombeo-subobovata
acute acuminata, plerumque basi obtusa, 8-10 cm. longa 4.5-5 cm.
lata 5-nervia, supra plus minusve persistente minute pubescentia,
subtus pallidiora ad nervos velutina, petiolo vix 10 mm. longo
velutino; spicae curvae 70 mm. longae 3 mm. crassae, pedunculo 5
mm. longo velutino; bracteae subcucullatae pilosae; baccae ovoideae,
stigmatibus sessilibus.

HONDURAS: River bank near Siguatepeque, Department of Coma-
yagua, 1,050 meters, Yuncker, Dawson & Youse 6042 (type in herb.
Univ. Illinois; duplicate in Herb. Field Mus.).

Piper siguatepequense var. subrhombifolium Trelease, var.
nov. — Folia rhombeo-ovata 12-13 cm. longa 6-7 cm. lata basi sub-
truncata; spicae 80 mm. longae.

HONDURAS : Region of the type, Yuncker, Dawson & Youse 561 7
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper tenuiculispicum Trelease, sp. nov. — Arbuscula 5-
metralis, internodiis floriferis breviusculis gracilibus primo breviter
pubescentibus nodis exceptis glabrescentibus; folia elliptica graciliter
acuminata, basi obtusa latere altero paullo breviore, 13-17 cm. longa
5.5-9.5 cm. lata e medio inferiore pinnatinervia, nervis ca. 5+6,
manifeste scabrida, nervis subtus ferrugineo-substrigosis, petiolo vix
10 mm. longo hispido; spicae 60 mm. longae 2 mm. crassae, pedunculo
5-6 mm. longo hispido.

HONDURAS: Bank of Tela River above Lancetilla Valley, Depart-
ment of Atlantida, 27 meters, Yuncker 4509 (type in herb. Univ.
Illinois).

Piper tesseraespicum Trelease, sp. nov. — Frutex 3-metralis,
ramulis ut petiolis nervisque subtus puberulis, internodiis floriferis
gracilibus subelongatis; folia subrhombeo-obovata vel elliptica obtuso-
acuminata, basi obtusa latere altero breviore, 24 cm. longa 13 cm.
lata e medio inferiore pinnatinervia, nervis 5+6, in sicco. tenuia
viridia venosa minute granuloso-asperata, petiolo ca. 15+3 mm.
longo; spicae 60 mm. longae 3 mm. crassae, bracteis deltoideis mani-
feste pilosis, pedunculo ca. 5 mm. longo; baccae triquetrae.

HONDURAS: River bank near Siguatepeque, Department of Coma-
yagua, 1,050 meters, Yuncker, Dawson & Youse 5662 (type in herb.
Univ. Illinois; duplicate in Herb. Field Mus.).

FLORA OF HONDURAS 355

Piper viae-marginis Trelease, sp. nov. — Frutex nodulosus
ramosus 2-metralis, internodiis superioribus gracilibus breviter
albido-tomentosis; folia late vel subrhombice elliptica obtuse acu-
minata, basi inaequaliter subcordulata, 9-15 cm. longa 4.5-7 cm.
lata e medio inferiore pinnatinervia, nervis 4-5X2, utrinque pube-
rula, subtus pallidiora et primo albo-tomentulosa, subrugulosa,
petiolo 10-15 mm. longo; spicae 60 mm. longae 4 mm. crassae,
pedunculo 5 mm. longo velutino; bracteae subcrescenticae ciliatae;
baccae triquetrae truncatae, stigmatibus sessilibus.

HONDURAS: Side of trail, near Siguatepeque, Department of
Comayagua, 1,050 meters, Yuncker, Dawson & Youse 6044 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper Yousei Trelease, sp. nov. — Frutex glaber vix 2-metralis,
internodiis floriferis gracilibus brevibus sed cito elongantibus nigres-
centibus; folia subelliptica acuminata, basi oblique acuta, 9-14 cm.
longa 5.5-6.5 cm. lata e medio inferiore pinnatinervia, nervis 4x2,
in sicco papyracea, subtus pallidiora, petiolo 5 vel 5+3-5 mm. longo;
spicae in sicco viso juveniles vix 10 mm. longae et 1 mm. crassae
pendulae, pedunculo minus quam 5 mm. longo.

HONDURAS: El Achote near Siguatepeque, Department of Coma-
yagua, 1,350 meters, Yuncker, Dawson & Youse 5999 (type in herb.
Univ. Illinois; duplicate in Herb. Field Mus.). El Rincon, 1,400-
1,500 meters, Yuncker, Dawson & Youse 6070.

Piper Yunckeri Trelease, sp. nov. — Frutex nodosus ramosis-
simus 1.5 m. altus, internodiis brevibus crassis pallidis persistente
tomentulosis; folia subelongato-elliptica acuta basi cordulata 9-10
cm. longa 3.5-4 cm. lata e medio inferiore pinnatinervia, nervis ca.
5x2, utrinque tomentulosa vel glabrescentia, petiolo 5 mm. longo
tomentuloso; spicae rectae vel curvae 60-110 mm. longae 3 mm.
crassae, pedunculo 10 mm. longo puberulo; bracteae conspicuae
rotundo-subpeltatae, margine pallido ciliato; baccae rotundatae,
stigmatibus sessilibus.

HONDURAS: Subalpine bog near El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6331 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Piper Yunckeri var. latum Trelease, var. nov. — Forma humilis
immatura vix metralis, foliis inferioribus latioribus.

HONDURAS: With the type, Yuncker, Dawson & Youse 6333 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

356 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

CHLORANTHACEAE
HEDYOSMUM Sw.

Hedyosmum Artocarpus Solms. A tree 6-7.5 meters tall and
12-15 cm. in diameter; aromatic. In a wet ravine near El Achote.
The tree is new to the flora of Honduras.

MYRICACEAE
MYRICA L.

Myrica mexicana Willd. A shrub about 1 meter tall, aromatic.
Edge of a ravine near El Achote.

BETULACEAE
OSTRYA (Michx.) Scop.

Ostrya virginiana var. guatemalensis (Winkl.) Macbr. A
tree about 9 meters tall and 15 cm. in diameter. Ravine near El
Achote.

FAGACEAE

QUERCUS L.

Quercus achoteana Trelease, sp. nov. — Leucobalanos-Macro-
phyllae(1}. Ramuli crassiusculi 4-5 mm. crassi substriati glabres-
centes grisei, lenticellis pallidis; gemmae ovoideae magnae glabrae
lucidae rubescentes; stipulae plus minusve persistentes setaceae;
folia decidua subobovata obtusa basi cordulata mucronulato-crenata
16 cm. longa 8 cm. lata, supra viridia glabra, subtus cito glabres-
centia, nervis 12-13x2 non arcuato-conjunctis; petiolus glabrescens
brevis crassus; flores et fructus? — A small tree, 5 meters tall, the
trunk 15 cm. in diameter.

HONDURAS: In pine-oak association near El Achote, above Sigua-
tepeque, Department of Comayagua, 1,350 meters, Yuncker, Dawson
& Youse 6136 (type in herb. Univ. Illinois; duplicate in Herb. Field
Mus.).

Quercus achoteana var. sublanosa Trelease, var. nov.—
Ramuli et praesertim folia subtus subpersistente ferrugineo-tomen-
tosa; folia obovata vel subpandurate elongata apiculato-crenata
9-18 cm. longa 5-10 cm. lata obtusa, basi cordulata. — A small tree,
5 meters tall, the trunk 10 cm. in diameter.

HONDURAS : Thicket on plain, Siguatepeque, Department of Coma-
yagua, 1,000-1,100 meters, Yuncker, Dawson & Youse 5961 (type
in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

FLORA OF HONDURAS 357

Quercus amissaeloba Trelease, sp. nov. — Erythrobalanos-Per-
seaefoliae(l). Ramuli graciusculi 2-3 mm. crassi striati, primo sub-
grisei serius fuscescentes et punctis minutis albis conspersi; folia
persistentia chartacea praesertim subtus lucida, late obovata et 6.5
cm. longa, 4 cm. lata, vel suboblonga et 9-12 cm. longa 3.5-5 cm.
lata, obtusa, paullo crispata, Integra vel interdum apice bilobulata,
basi subacuta vel subcordulata, petiolo vix 5 mm. longo; inflores-
centia? — A small, glabrous tree 6 meters tall, the trunk 10-15 cm.
in diameter.

HONDURAS: In ravine near El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6379 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Quercus Dawsoni Trelease, sp. nov. — Erythrobalanos-Acuti-
$oliae(1}. Ramuli graciles 2-3 mm. crassi vix striati transiente
aliquanto velutini ; gemmae juveniles subglobosae glabratae; stipulae
caducae; folia decidua oblanceolata acuminata, ad petiolum gracilem
longiattenuata, 15-17 cm. longa 5 cm. lata, in fere toto margine
cuspidato-serrata, cito glabrescentia et lucida, nervis 10-12x2,
petiolo 1-2 cm. longo; inflorescentia?— A small tree, 3 meters tall,
the trunk 7-8 cm. in diameter.

HONDURAS: In ravine near El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson & Youse
6378 (type in herb. Univ. Illinois).

Quercus perseaefolia(?) Liebm. var. achoteana Trelease,
var. nov. — Erythrobalanos-Perseaefoliae. Glabra; ramuli 2-3 mm.
crassi grisei inconspicue lenticellati ; gemmae rotundatae vel ovoideae
2-3 mm. longae 2 mm. crassae; folia persistentia suboblonga utrinque
subacuta vel obtusa et basi aliquanto inaequaliter obtusa, 7-15 cm.
longa 3-4 cm. lata, integra chartacea subcrispata paullo lucida,
petiolo vix 5 mm. longo; inflorescentia? — A small tree, 4 meters tall,
the trunk 5-7 cm. in diameter.

HONDURAS: In thicket west of El Achote, above Siguatepeque,
Department of Comayagua, 1,300 meters, Yuncker, Dawson & Youse
6203 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Quercus pertessellata Trelease, sp. nov. — Erythrobalanos-
SyntheticaeC*} . Glabra; ramuli graciles 2-3 mm. crassi striati, primo
rubescentes serius grisei vel ochracei, lenticellis inconspicuis; folia
decidua sublanceolato-oblonga acuminata, basi acuta, 15-16 cm.
longa 4-5 cm. lata crasse papyracea, supra obscure viridia, utrinque
cancellate venosa, nervis ca. 12x2 arcuato-conjunctis, petiolo

358 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

5-10 mm. longo gracili ; inflorescentia? — A small tree, 3 meters tall,
the trunk 5-7 cm. in diameter.

HONDURAS: In ravine near El Achote, above Siguatepeque,
Department of Comayagua, 1,350 meters, Yuncker, Dawson &
Youse 6376 (type in herb. Univ. Illinois).

Quercus tenuiaristata Trelease, sp. nov. — Erythrobalanos-
Tristes(t). Ramulis transiente subvelutinis gracilibus 2-4 mm.
crassis striatis subgriseis vix lenticellatis exceptis glabra; gemmae
ovoideae glabratae; folia decidua oblongo-lanceolata vel elliptico-
lanceolata apice aristata, basi obtusa ad late rotundata, grosse ser-
rata, nervis 12-15 dentibus graciliter setiferis terminatis, petiolo vix
5 mm. longo; inflorescentia? — A small tree, 3 meters tall, the trunk
5-7 cm. in diameter.

HONDURAS: In thicket west of El Achote, above Siguatepeque,
Department of Comayagua, 1,400 meters, Yuncker, Dawson & Youse
6204 (type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Quercus trichodonta Trelease, sp. nov. — Erythrobalanos-Acuti-
foliae(l). Ramulis novellis pallido-puberulis exceptis glabra; ramuli
3-5 mm. crassi vix striati vel lenticellati ; folia decidua lanceolata e
medio utrinque angustata, integra, nervis 15 et ultra delicate ari-
statis, vel supra medium grosse crenata dentibus graciliter aristatis,
subtus pallidiora, nervis albis, petiolo crasso vix 5 mm. longo; inflores-
centia?— A small tree, 6 meters high, the trunk 5-7 cm. in diameter.

HONDURAS: In ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6116
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Quercus Yousei Trelease, sp. nov. — Leucobalanos-Corrugatae(*!).
Ramuli pallidi glabri graciles; gemmae subglobosae glabratae;
stipulae 2 mm. longae deciduae striatae; folia decidua prope nodos
dense inserta lanceolata- obtuse subacuminata, basi cordulata, petiolo
brevissimo, supra medium grosse obtuseque crenato-serrata, 14-15
cm. longa 3.5 cm. lata, nervis ca. 12-15x2; inflorescentia? — A small
tree, 4 meters tall, the trunk 5-6 cm. in diameter.

HONDURAS: Ravine at El Achote, above Siguatepeque, Depart-
ment of Comayagua, 1,350 meters, Yuncker, Dawson & Youse 6297
(type in herb. Univ. Illinois; duplicate in Herb. Field Mus.).

Quercus Yunckeri Trelease, sp. nov. — Leucobalanos-Circi-
natae(l}. Ramuli crassiusculi 4-5 mm. crassi striati ferrugineo-
villosi; folia ut videtur ad nodos anni prioris fasciculata late oblanceo-

FLORA OF HONDURAS 359

lata acute acuminata, versus basin Integra longicuneata, supra
medium subgrosse mucronato-serrata, 22 cm. longa 9 cm. lata, primo
subtus laxe tomentosa, nervis 15x2 non arcuato-conjunctis, petiolo
fere nullo; fructus? — A small tree, 6 meters tall, the trunk 5-7 cm.
in diameter.

HONDURAS: In dense forest near the summit of the range above
El Achote, above Siguatepeque, Department of Comayagua, 1,800
meters, Yuncker, Dawson & Youse 6180 (type in herb. Univ. Illinois;
duplicate in Herb. Field Mus.).

ULMACEAE

CHAETOPTELEA Liebm.

Chaetoptelea mexicana Liebm. A tree about 6 meters tall
and 15 cm. in diameter. In a marsh near El Achote.

MORACEAE
CECROPIA L.

Cecropia mexicana Hemsl. A tree about 7.5 meters tall and
10-12 cm. in diameter. In a wet ravine near El Achote.

DORSTENIA L.
Dorstenia Contrajerva L. River bank.

FICUS L.

Ficus costaricana (Liebm.) Miq. A tree about 9 meters tall
and 1-1.5 meters in diameter. In a wet ravine near El Achote. This
tree has not been recorded for Honduras.

Ficus glabrata HBK. A fine, broad-topped tree about 15
meters tall and 60 cm. in diameter. River bank.

Ficus involuta (Liebm.) Miq. A tree about 4-5 meters tall and
30 cm. in diameter. River bank.

Ficvis padifolia HBK. A tree about 8 meters tall and 60 cm.
in diameter. River bank.

Ficus radula Willd. A tree about 9 meters tall and 60 cm. in
diameter. River bank.

Ficus velutina Willd. A tree about 6 meters tall and 10 cm.
in diameter. In a wet ravine near El Achote.

MORUS L.

Morus multicaulis Perr. A small plantation of this species was
found near Siguatepeque.

360 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

TROPHIS L.

Trophis chorizantha Standl. A tree about 7.5 meters tall and
10 cm. in diameter. Near the summit of the range above El Achote.
This tree, known formerly only from Honduras, has been collected
recently in the region of El General, Costa Rica.

URTICACEAE
BOEHMERIA Jacq.

Boehmeria caudata Sw. Flowers greenish white. River bank
about 6 km. west of Siguatepeque.

MYRIOCARPA Benth.

Myriocarpa yzabalensis (D. Sm.) Killip. A tree about 4.5
meters tall. In a wet ravine near El Achote.

LORANTHACEAE
PHORADENDRON Nutt.

Phoradendron commutatum Trel. On a tree near Potrerillos.

Phoradendron piperoides (HBK.) Trel. On a tree near the
shore of Lake Yojoa.

Phoradendron yojoanum Trelease, sp. nov.— Aequitoriales-
Annulatae. Plus minusve pseudodichotoma, cataphyllis omnibus
basalibus, subconspicue nodosa, internodiis breviusculis 3-5- cm.
longis, infra nodos ancipitibus, alibi in sectione rhombeis vel tere-
tibus; par cataphyllorum 1 basale, cataphyllis annulatis; folia plus
minusve falcate oblanceolata obtusa, basi cuneato-contracta vix
petiolata, 5-6 cm. longa 1-2 cm. lata, opaca subpapillata basinervia,
nervis apicem versus evanescentibus; spicae breves vix 2 cm. longae,
articulis ca. 4 clavatis, floribus luteis 4-seriatim insertis, pedunculo
brevi; baccae ut dicitur albae.

HONDURAS: Paral, Lake Yojoa, Dept. Corte"s, at 630 meters, T. G.
Yuncker 4879 (type in Herb. Univ. Illinois), 4872.

Phoradendron Yunckeri Trelease, sp. nov. — Aequitoriales-
Heydeanae. Aliquanto pseudodichotoma glabra, internodiis brevi-
usculis 3-4 cm. longis primo infra nodos ancipiti-dilatatis cito tereti-
bus; par cataphyllorum 1 basale in nodo infimo tantum insertum,
cataphyllis patentibus; folia oblonga vel elliptica aliquanto falcata
obtusa, basi cuneatim in petiolum 10 mm. longum angustata, firma
opaca pinnatinervia, costa percurrente, 5-7 cm. longa 2.5-3 cm. lata;
spicae solitariae longiusculae 40-50 mm. longae, articulis ca. 4

FLORA OF HONDURAS 361

gracilibus subfusiformibus, floribus 4+2-seriatim insertis, pedunculo
vix 5 mm. longo; fructus juvenilis globosus, sepalis arete inflexis.
HONDURAS: Near Paral, Dept. Cortes, 630 meters, T.G. Yuncker
5097 (type in herb. Univ. Illinois).

STRUTHANTHUS Mart.

Struthanthus cassythoides Millsp. On an oak tree near El
Achote.

Struthanthus densiflorus (Benth.) Standl. On pine. Hills
east of Siguatepeque.

Struthanthus orbicularis (HBK.) Blume. On a tree about
12 km. west of Tela.

POLYGONACEAE
COCCOLOBA L.

Coccoloba cardiophylla Standl. A small tree about 4.5 meters
tall and 10 cm. in diameter. On plains. The species has been known
previously only from the Yucatan Peninsula, and was scarcely to be
expected in central Honduras.

POLYGONUM L.
Polygonum hispidum HBK. Flowers white. In a marsh.

Polygonum punctatum Ell. In a marsh near the shore of Lake
Yojoa.

NYGTAGINACEAE
BOERHAAVIA L.
Boerhaavia erecta L. In waste places near Potrerillos.

MIRABILIS L.

Mirabilis Jalapa L. Flowers pink or white. Escaped along
bank of river.

PHYTOLACCACEAE

PHYTOLACCA L.

Phytolacca Meziana H. Walt. An herb. Perianth greenish to
somewhat pink. Sprawling over shrubs and on the ground in the
forest near the summit of the range above El Achote.

Phytolacca rivinoides Kunth & Bouche". Stems 12-14 mm.
thick at the base and somewhat woody; racemes up to 60 cm. long;
flowers pinkish white; berries purple. Sprawling over shrubs in
forest near the summit of the range above El Achote.

362 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

CARYOPHYLLACEAE
ARENARIA L.

Arenaria lanuginosa (Michx.) Rohrb. A prostrate herb with
white flowers. River bank.

DRYMARIA Willd.

Drymaria villosa Schl. & Cham. A low herb. A very common
weed, in waste places.

RANUNCULACEAE
THALICTRUM L.

Thalictrum hondurense Standl., sp. nov. — Herba perennis ca.
metralis, caule crassiusculo tereti superne dense glanduloso-puberulo
inferne fere glabro, internodiis elongatis; folia magna, petiolo 2-2.5
cm. longo ad vel fere ad apicem late vaginato, vagina nervosa glabra;
foliola numerosa paullo supra basin peltata membranacea ambitu
rotundata vel obovato-rotundata, interdum ovato-rotundata, ple-
rumque 2-2.5 cm. longa, longipetiolulata, plerumque plus minusve
triloba et vulgo grosse paucicrenata, supra sparse glandulosa, subtus
ubique glanduloso-scaberula, pallida, venis valde elevatis laxe reti-
culatis; flores dioeci, feminei laxe paniculati, panicula magna laxa,
floribus ad 2.5 cm. longe pedicellatis; achaenia vulgo 2-3 sessilia
glanduloso-puberula valde obliqua et curva, immatura fere 5 mm.
longa, basi acuta, apice acuta et subito in stylum filiforme usque 6
mm. longum contracta, tenuiter costata. — Honduras: On rocky
hillside near Siguatepeque, Dept. Comayagua, July 11, 1936, T. G.
Yuncker, R. F. Dawson & H. R. Youse 5796 (type in Herb. Field
Mus.).

"Plants more or less viscid and with a pungent odor; roots a
rich yellow color." Closely related to T. guatemalense C. DC. &
Rose, which is noteworthy for the much longer and denser pubescence
found on all parts of the plant.

MENISPERMACEAE
CISSAMPELOS L.

Cissampelos Pareira L. A woody vine. Climbing over shrubs
in a thicket near Potrerillos.

Cissampelos tropaeolifolia DC. A woody vine. Climbing
over shrubs. River bank.

FLORA OF HONDURAS 363

ANNONACEAE
GUATTERIA Ruiz & Pavon

Guatteria grandiflora Bonn. Sm. A tree 6-7.5 meters tall
and 10-12 cm. in diameter. Perianth yellow-buff tinged with pink.
In a wet ravine near the summit of the range above El Achote.

SAPRANTHUS Seem.

Sapranthus microcarpus (D. Sm.) Fries. A shrub about
2 meters tall. Corolla dull, dark red; flowers fragrant. River bank.

MONIMIACEAE
SIPARUNA Aubl.

Siparuna nicaraguensis Hemsl. A small tree about 4.5
meters tall and 3-5 cm. in diameter; fruit dark red, fleshy. Bottom
of a ravine near El Achote; also in a thicket near the shore of Lake
Yojoa.

LAURACEAE

LITSEA Lam.

Litsea glaucescens HBK. A tree about 2.5-3 meters tall and
3-5 cm. in diameter. In a wet ravine near El Achote. New to the
Honduran flora.

NECTANDRA Roland

Nectandra sanguinea Rottb. A shrub 3.5-4.5 meters tall.
In a thicket west of El Achote.

Nectandra surinamensis Mez. A tree about 9 meters tall and
15-20 cm. in diameter; corolla white; flowers fragrant. In a ravine
about 6 km. west of Siguatepeque.

PHOEBE Nees

Phoebe helicterifolia Mez. A tree about 7 meters tall and 15
cm. in diameter. River bank.

Phoebe mexicana Meissn. A tree about 9 meters tall and 45 cm.
in diameter River bank.

GRASSULACEAE
BRYOPHYLLUM Salisb.

Bryophyllum pinnatum (L.) Kurz. An herb about 1 meter
tall; flowers reddish. River bank.

364 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

HAMAMELIDACEAE
DISTYLIUM Sieb. & Zucc.

Distylium guatemalense Radlk. ex Harms, Notizbl. Bot.
Gart. Berlin 11 : 716. /. 13. 1933.— A tree about 9 meters high and 20-
25 cm. in diameter; in wet ravine near El Achote, hills above the
plains of Siguatepeque, 1,350 meters, Yuncker, Dawson & Youse
6377. This tree is the only American representative of the genus
Distylium, represented otherwise by about nine species in tropical
and subtropical southeastern Asia. Its occurrence in Guatemala
(where first collected by Tuerckheim in 1907, near Coban, at 1,350
meters, No. II. 1613} and Honduras is, therefore, a matter of more
than passing interest from the standpoint of distribution of the
Hamamelidaceae. The Tuerckheim material is in flower, and at
that stage the plant at first glance gives little indication of its relation-
ship with this group, at least to one not especially familiar with the
family. A specimen of the Tuerckheim number sent me some years
ago from the Delessert Herbarium I failed to place, but when Dr.
Harms's description and excellent illustration were published in 1933,
I immediately recalled the specimen that I had been unable to name.

The Honduras collection is clearly of the same species, and bears
fruits, which were not available to Radlkofer and Harms. These
in appearance remind one at once of those of witch hazel (Ham-
amelis). They are very hard and woody, blackish and somewhat
roughened on the outer surface, which is densely stellate-tomentose,
nearly 1.5 cm.; no seeds are present on the evidently old capsules
with the specimen. A remarkable character much in evidence in the
Honduran material but not mentioned in the original description is
the fact that some but not all the leaves are conspicuously peltate
at the base, the blade being extended for about 2 mm. below the
insertion of the petiole. Paul C. Standley.

LIQUIDAMBAR L.

Liquidambar Styraciflua L. A tree about 6 meters tall and
15-20 cm. in diameter (some specimens seen were much larger).
Near El Achote.

ROSACEAE
PHOTINIA Lindl.

Photinia microcarpa Standl. A tree about 4.5 meters tall and
8-10 cm. in diameter; fruit red. In a wet ravine near El Achote.
The tree occurs also in the mountains of British Honduras.

FLORA OF HONDURAS 365

RUBUS L.

Rubus adenotrichos Schlecht. A shrub about 2 meters tall.
Edge of a ravine near El Achote.

Rubus miser Liebm. A shrub about 2.5 meters tall; flowers
pink or white. In a thicket near margin of forest at the summit of
the range above El Achote.

CONNARACEAE
CONNARUS L.

Connarus Lambertii (DC.) Britton. A shrub about 2.5 meters
tall; pods red. In a marsh west of Tela.

KRAMERIACEAE

Krameria cuspidata Presl. A shrub about 1 meter high;
flowers light maroon. On a rocky hillside.

LEGUMINOSAE
ACACIA Willd.

Acacia Farnesiana (L.) Willd. A small tree about 3 meters
tall ; flowers yellow. Alluvial soil, river bottoms.

Acacia Milleriana Standl. A tree 5-6 meters tall and 30 cm.
in diameter. Frequent on the plains.

Acacia salvadorensis (Britt. & Rose) Standl. A shrub about
1.5 meters tall; corolla greenish, becoming reddish toward the base;
stamens pure white. In an open ravine, under pines, near El Achote.

CALLIANDRA Benth.

Calliandra arborea Standl., sp. nov. — PI. XII. Arbor 4-9-
metralis, trunco 5-15 cm. diam., ramulis ochraceis subteretibus
solemniter elevato-lenticellatis glabris dense foliatis; stipulae parvae
striatae; folia glabra 1-2 cm. longe petiolata, pinnis unijugis, foliolis
6; foliola sessilia coriacea valde obliqua, semiovata, paris terminalis
4-11 cm. longa 1.5-4 cm. lata, apicem versus longe anguste acu-
minata, apice ipso obtuso, basi valde obliqua semiobtusa, lucida,
supra griseo-viridia, nervis prominulis, venulis manifestis vix pro-
minulis, subtus fere concoloria, basi triplinervia, nervis 2 centralibus
fere ad apicem prolongatis gracilibus prominentibus, venis prominulis
laxe reticulatis; capitula axillaria solitaria dense multiflora staminibus
exclusis fere 1.5 cm. diam., pedunculis 1.5-2 cm. longis glabris vel
sparse villosulis; calyx tubuloso-campanulatus 1.5-2.5 mm. longus

366 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

sparse minute puberulus, dentibus brevissimis obtusis; corolla 1 cm.
longa angusta glabra vel sparse minutissime puberula, lobis acutis ca.
2 mm. longis; stamina 3-3.5 cm. longa. — Honduras: In thicket near
El Rincon, about 10 miles west of Siguatepeque, Dept. Comayagua,
1,400-1,500 meters, July 24, 1936, T. G. Yuncker, R. F. Dawson &
H. R. Youse 6047 (type in Herb. Field Mus.); corolla reddish, the
filaments red on the outermost half. Side of ravine near El Achote,
hills above the plains of Siguatepeque, 1,350 meters, Yuncker, Dawson
& Youse 5912; round-topped. Open mountain forest, Jacaieto,
Dept. Comayagua, 600 meters, August, 1933, J. B. Edwards 637;
flowers red and white.

Calliandra arborea is noteworthy for the large size of the plant,
herbs and shrubs being much more common in the genus than trees,
even small ones. It is most closely related to C. centralis (Britt. &
Rose) Standl., also Honduran, which has smaller, very obtuse leaflets
and usually a noticeable amount of pubescence on various parts of
the plant. C. centralis (Anneslia centralis Britt. & Rose, N. Amer.
Fl. 23: 52. 1928) is based upon C. Thieme 5214 from Santa Barbara,
Honduras, a sheet of which is in the Herbarium of Field Museum.
The original description of this species is so badly and incorrectly
drawn that I suspected the type must be different from the Field
Museum specimen. The type was kindly lent from the U. S. National
Museum, and was found to agree exactly with the specimen in
Field Museum. It is probable, therefore, that all specimens of this
number in other herbaria are conspecific, although upon comparing
them with Britton and Rose's description, one would be convinced
that they represented a distinct species.

Calliandra centralis (Britt. & Rose) Standl. A small tree,
about 4.5 meters tall. Stamens scarlet. Growing on a river bank.

Calliandra leucothrix Standl., sp. nov. — Frutex 1.5 m. altus,
ramis crassiusculis teretibus fusco-brunneis dense pilis brevibus
albidis patentibus hirsutis, sat dense foliatis; stipulae deciduae 4 mm.
longae hispidulae; folia mediocria breviter petiolata ad 16 cm. longa,
pinnis ca. 13-jugis; foliola parva ca. 25-juga crasse membranacea
anguste oblonga 4-6 mm. longa 1.5 mm. lata obtusa vel acutiuscula,
basi valde obliqua, 1-nervia, glabra sed ad margines dense pilis longis
albis adpressis ciliata, nervis lateralibus subtus prominulis mani-
festis; capitula pauciflora breviter pedunculata racemum brevem
terminalem efformantia, pedunculis ad 1 cm. longis subadpresso-
hispidulis, bracteis ad basin capituli insertis parvis vix 2 mm. longis,

FLORA OF HONDURAS 367

floribus crasse fere 2 mm. longe pedicellatis; calyx ca. 2 mm. longus
atque 4 mm. latus remote triangulari-dentatus, extus pilis albidis
adpresso-hispidulus; corolla 1 cm. longa extus dense albo-strigosa,
lobis tubo aequilongis oblongo-lanceolatis acuminatis erectis; fila-
menta purpurea ca. 6 cm. longa. — Honduras: In pine forest near
El Achote, hills above the plains of Siguatepeque, Dept. Comayagua,
1,350 meters, July 26, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 6128 (type in Herb. Field Mus.).

Belonging to the group Houstonianae of the genus Anneslia, as
treated by Britton and Rose in the North American Flora, and
apparently closest to their Anneslia pueblana.

Calliandra Yunckeri Standl., sp. nov. — Frutex 2-metralis,
ramis crassis teretibus ochraceis densiuscule lenticellatis, foliis
inflorescentiisque e ramis abbreviatis crassis nascentibus; stipulae
persistentes imbricatae; folia parva, petiolo 6-12 mm. longo sparse
longipiloso, rhachi gracili ad 1.5 cm. longa sparse pilosa; pinnae 2-3-
jugae 1.5-3.5 cm. longae, rhachi sparse pilosa, foliolis 9-16-jugis
oblongis plerumque 6-7 mm. longis atque 2.5-3 mm. latis acutius-
culis vel obtusis, basi valde obliquis, glabris sed dense pilis longis
albis ciliatis, subtus paullo pallidioribis costa excentrica prominente,
nervis lateralibus paucis prominulis irregularibus; capitula axillaria
pauca ca. 2.5 cm. longe pedunculata dense subpauciflora; calyx
tubuloso-campanulatus fere 3 mm. longus dentibus brevissimis
pilosulis exceptis glaber; corolla 6 mm. longa lobis brevibus late
ovato-triangularibus sparse minute puberulis exceptis glabra;
stamina ca. 3 cm. longa.— Honduras: In thicket, plain near Sigua-
tepeque, Dept. Comayagua, 1,050 meters, July 4, 1936, T. G. Yun-
cker, R.F. Dawson & H. R. Youse 5622 (type in Herb. Field Mus.).

CALOPOGONIUM Desv.

Calopogonium coeruleum Benth. A woody vine; flowers
purple. River bank.

CANAVALIA HBK.

Canavalia villosa Benth. A herbaceous vine with showy,
blue-lavender flowers. Thicket near El Achote.

CASSIA L.

Cassia laevigata Willd. A shrub about 2 meters tall; flowers
yellow. Ravine near El Achote.

368 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

Cassia serpens L. A low, prostrate herb with yellow flowers.
In wet, rocky soil in pine forest.

Cassia spectabilis DC. A spreading, round-headed tree about
7-8 meters tall; flowers yellow. River bank.

COLOGANIA Kunth

Cologania procumbens Humb. & Bonpl. Flowers blue. In
wet soil, pine forest.

DALBERGIA L. f.

Dalbergia cubilquitzensis (D. Sm.) Pittier. A tree about 6
meters tall. In forest near the summit of the range above El Achote.

DESMODIUM Desv.

Desmodiuni axillare (Sw.) DC. Corolla red; a weed, in open
places about Lancetilla valley.

Desmodium cajanifolium (HBK.) DC. Flowers lavender.
River bank.

Desmodium frutescens (Aubl.) Schindl. Plants 30-60 cm.
tall, becoming woody below; flowers lavender. In wet soil, pine
forest.

Desmodium luteolum Standl., sp. nov. — PI. XIII. Herba ut
videtur erecta 60 cm. alta vel ultra, caule crasso tereti striato pallide
viridi densissime pilis longis patentibus laxis albidis vel fulvis piloso,
internodiis brevibus; stipulae longius persistentes ca. 12 mm. longae
e basi late subcordato-ovata abrupte anguste caudatae brunneae
striatae densissime pilosae; folia modica longipetiolata, petiolo
gracili 6-7.5 cm. longo dense patenti-piloso, rhachi ca. 1.5 cm. longa,
stipellis lineari-filiformibus ca. 5 mm. longis attenuatis, petiolulis
crassis ca. 3 mm. longis dense pilosis; foliola 3 deltoideo-rotundata,
ovato-rotundata vel fere orbicularia 4.5-6 cm. longa 4-5.5 cm. lata,
apice late rotundata, basi subtruncata vel late rotundata et emargi-
nata, crasse membranacea, supra in sicco luteo-viridia dense pilis
laxis subadpressis albidis longipilosa et papillosa, subtus paullo
pallidiora ubique similiter sed densius pilosa, nervis venisque pro-
minentibus teneris, venis subarcte reticulatis; racemi simplices
longipedunulati subdense multiflori ca. 30 .cm. longi densissime vis-
cido-pilosi, bracteis caducis, pedicellis gracilibus inaequalibus usque
18 mm. longis adscendentibus dense viscido-pilosis; calyx ca. 8 mm.
longus sparse longipilosus profunde lobatus, lobis lanceolatis longe
subulato-attenuatis; corolla ca. 11 mm. longa, petalis glabris; legu-

FLORA OF HONDURAS 369

men non visum. — Honduras: On rocky hillsides above Siguatepeque,
Dept. Comayagua, 1,200 meters, July 11, 1936, T. G. Yuncker, R. F.
Dawson & H. R. Youse 5793 (type in Herb. Field Mus.).
"Flowers greenish yellow."

DIALIUM L.

Dial in in guianense (Aubl.) Steud. A tree 7.5 meters tall. In
wet soil at Puerto Arturo, 12 km. west of Tela.

ERIOSEMA Desv.

Eriosema diffusum (HBK.) Don. Flowers pinkish yellow. On
a rocky hillside in pine forest.

ERYTHRINA L.

Erythrina rubrinervia HBK. A tree 6-8 meters tall; flowers
scarlet. Rather frequent along river banks.

GLIRICIDIA HBK.

Gliricidia meistophylla (Donn. Sm.) Pittier. A small tree
about 2 meters tall. Ravine near El Achote. The species is new to
the flora of Honduras.

INDIGOFERA L.

Indigofera suffruticosa Mill. A shrub about 3 meters tall;
flowers pinkish. In wet soil, waste places on plains.

INGA Scop.

Inga cobanensis Pittier. A tree about 10 meters tall; pods
brown, woolly. In a wet ravine near El Achote.

MIMOSA L.

Mimosa albida Humb. & Bonpl. A shrub about 1.5 meters
tall; corolla pinkish brown; stamens white or pink. In a ravine in
pine forest near El Achote.

Mimosa pigra L. A shrub about 2.5 meters tall. In a marsh.

Mimosa pudica L. Flowers pink. Abundant in old meadows
and waste areas.

PARKINSONIA L.

Parkinsonia aculeata L. A tree about 6 meters tall. Planted
near Siguatepeque.

370 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

PAROSELA Cav.

Parosela sericea (Lag.) Rose. In open pine forest above El
Achote.

PHASEOLUS L.

Phaseolus anisotrichus Scheele. Flowers orange. Rocky
hillside.

PISCIDIA L.

Piscidia grandifolia (Donn. Sm.) I. M. Johnston. A tree
about 6 meters tall. Rather frequent along river banks.

PITHECOLOBIUM Mart.

Pithecolobium arboreum (L.) Urban. A small tree about
3.5 meters tall. In forest near the summit of the range above El
Achote.

RHYNCHOSIA Lour.

Rhynchosia pinetorum Standl., sp. nov. — Herbacea scandens,
caule gracillimo subtereti pilis paten tibus viscidis lutescentibus piloso,
internodiis plerumque foliis longioribus; stipulae persistentes lineari-
oblongae acutae costatae ca. 4 mm. longae breviter pilosae; folia
parva 6-13 mm. longe petiolata, petiolo gracili viscido-piloso, rhachi
ad 6 mm. longa; foliola 3 triangularia vel ovato-deltoidea 1.5-2.5 cm.
longa 1-1.5 cm. lata sensim attenuato-acuminata, basi late rotundata,
crasse membranacea, supra in sicco viridia papillosa et dense molliter
pilosa, subtus fere concoloria, minute glanduloso-punctata, dense
molliter pilosa, nervis conspicuis elevatis; racemi axillares laxe 3-6-
flori ad 3.5 cm. longe pedunculati, rhachi dense viscido-pilosa,
bracteis brunnescentibus tenuibus caducis ovato-oblongis, floribus
remotis, pedicellis ca. 2 mm. longis; calyx viscido-pilosus 5-6 mm.
longus, tubo late campanulato basi obtuso vel rotundato, lobis tubo
paullo longioribus, lobo infimo aliis longiore, omnibus lineari-lanceo-
latis longiattenuatis enerviis; corolla 8 mm. longa, vexillo extus
apicem versus breviter piloso; legumen immaturum oblongum 17
mm. longum 5 mm. latum sessile dense pilis longis et brevibus inter-
mixtis plus minusve viscidis pilosum, apice obtusum vel subrotunda-
tum et subito in stylum ad 9 mm. longum contractum; semina 2.—
Honduras: On rocky hillside above Siguatepeque, Dept. Comayagua,
1,200 meters, July 11, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 5791 (type in Herb. Field Mus.).

VIGNA Savi

Vigna repens (L.) Kuntze. A trailing vine with yellow flowers.
Subalpine bog near El Achote.

FLORA OF HONDURAS 371

ZORNIA Gmel.

Zornia diphylla (L.) Pers. A low, prostrate herb with yellowish
flowers. In wet areas.

OXALIDACEAE
OXALIS L.

Oxalis latifolia HBK. Flowers lavender or violet. Common in
the pine forest belt above the plains.

Oxalis Neaei DC. Flowers yellow. River bank.

RUTACEAE
ZANTHOXYLUM L.

Zanthoxylum limoniodorum Standl., sp. nov. — Frutex ut
dicitur scandens, fere omnino ut videtur glaber, ramulis crassis
teretibus fuscescentibus atque rimosis, internodiis plus minusve elon-
gatis, sparse aculeis recurvis basi latis vix 4 mm. longis armatis; folia
breviter petiolata ca. 13 cm. longa, rhachi supra canaliculata exalata
ut petiolo aculeis numerosis recurvis brunneis ca. 3 mm. longis
armata; foliola ca. 19 sessilia vel brevissime petiolulata oblongo-
ovalia vel terminale ovato-ovale, 2-3.5 cm. longa 1-1.6 cm. lata,
apice obtusa vel rotundata et breviter retusa, basi obtusa vel rotun-
data, coriacea, fere ad basin grosse glanduloso-crenata, supra in
sicco olivacea sublucida, costa impressa, nervis obsoletis, subtus
lutescentia vel brunnescentia, costa gracili elevata interdum prope
basin aculeolata, nervis lateralibus obscuris; inflorescentia ut videtur
axillaris pyramidali-paniculata folio brevior multiflora, ramis puberu-
lis, floribus breviter crasse pedicellatis; folliculus ut videtur 1 tantum
globosus sessilis basi et apice rotundatus ca. 5 mm. diam. — Hon-
duras: Climbing in trees in a ravine near El Achote, hills above the
plains of Siguatepeque, Dept. Comayagua, 1,350 meters, July 19,
1936, T.G. Yuncker, R.F. Dawson & H. R. Youse5974 (type in Herb.
Field Mus.).

"Fruit with a strong lemon odor." Noteworthy for the small,
blunt, coriaceous, coarsely crenate leaflets, and for the abundant
prickles of the leaves. The material is in such advanced fruiting
condition that the flower details can not be determined.

Zanthoxylum microcarpum Griseb. A small tree, about 4.5
meters tall and 8-10 cm. in diameter, flowers greenish white. River
bank.

372 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

SIMARUBACEAE
PICRAMNIA Sw.

Picramnia andicola Tulasne. A tree about 4.5 meters tall and
5-8 cm. in diameter. In a wet ravine near El Achote. A species
new for Honduras.

Picramnia locuples Standl., sp. nov. — Frutex vel arbuscula
2-4.5 m. alta, trunco ad 5 cm. diam., ramulis gracilibus teretibus
brunnescentibus dense pilis brevibus sordidis patentibus vel adscen-
dentibus pilosis, internodiis plerumque elongatis; folia breviter
petiolata 7-16 cm. longa, rhachi gracili dense pilosula; foliola ca. 19
parva membranacea oblique ovata ad lanceolato-oblonga 1.5-3.5 cm.
longa 8-12 mm. lata obtusa ad acuminata, apice ipso obtuso, basi
oblique rotundata vel obtusa, supra in sicco fusco-viridia ad costam
pilosula aliter glabra, nervis obsoletis, subtus paullo pallidiora tantum
ad costam gracilem prominentem incurvo-pilosula, ciliata, nervis
prominulis paucis; inflorescentia feminea racemosa simplex gracili ter
longipedunculata, racemis laxe multifloris ad 8 cm. longis, rhachi
dense breviter pilosa, pedicellis patentibus plerumque 8-10 mm.
longis glabris; sepala decidua; fructus immaturus obovoideus glaber
apice rotundatus 4 mm. longus. — Honduras: In forest near the
summit of the ridge above El Achote, above the plains of Siguate-
peque, Dept. Comayagua, 1,800 meters, July 21, 1936, T.G. Yuncker,
R. F. Dawson & H. R. Youse 6006 (type in Herb. Field Mus.). In
wet ravine near El Rincon, about 10 miles west of Siguatepeque,
1,400 meters, Yuncker, Dawson & Youse 6063.

Fruit bright red. Well marked among Central American species
of Picramnia by the small and usually numerous leaflets.

MELIACEAE

MELIA L.

Melia Azedarach L. A small tree about 3 meters tall and 7-8
cm. in diameter. Naturalized on a river bank.

TRICHILIA L.

Trichilia havanensis Jacq. A tree 7-8 meters tall and 45 cm. in
diameter. Frequent. River bank.

MALPIGHIACEAE
BUNCHOSIA Rich.

Bunchosia pilosa HBK. A shrub about 2 meters tall; corolla
yellow. Bottom of a wet ravine near El Rincon ; also near El Achote.

FLORA OF HONDURAS 373

BYRSONIMA Rich.

Byrsonima crassifolia (L.) DC. A small tree about 3 meters
tall and 7-8 cm. in diameter; flowers orange. On a rocky hillside in
pine forest belt; also on sand dunes near the beach west of Tela.

MALPIGHIA L.

Malpighia glabra L. A tree 7.5-9 meters tall and 10-12 cm. in
diameter; petals pink. In wet ravine near El Achote.

STIGMAPHYLLON Juss.

Stigmaphyllon ellipticum (HBK.) Juss. A woody vine,
climbing over shrubs in a thicket. River bank.

Stigmaphyllon puberum Juss. A woody vine; corolla yellow
and orange. Climbing in a thicket between Tela and Lancetilla
Valley.

POLYGALACEAE

POLYGALA L.

Polygala glochidiata HBK. Flowers white. River bank.
Polygala incarnata L. Corolla purplish. In pine forest.

Polygala longicaulis HBK. Flowers pink. In an old, wet
pasture.

DICHAPETALACEAE
DICHAPETALUM Thouars

Dichapetalum Donnell-Smithii Engler. A shrub or small
tree 4.5 meters tall. In a thicket near the beach west of Tela.

EUPHORBIACEAE

ACALYPHA L.
Acalypha alopecuroides Jacq. A weedy plant in waste places.

Acalypha leptopoda Muell. Arg. A small, slender tree about
3-4 meters tall. Thicket. River bank.

Acalypha macrostachya Jacq. A tree about 3.5 meters tall
and 2.5-5 cm. in diameter. On a ridge about 6 km. north of Si-
guatepeque.

Acalypha porphyrantha Standl. A shrub 0.5-1 meter tall.
River bank.

374 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

CROTON L.

Croton ciliato-glandulosus Ortega. A shrub about 1 meter
tall; flowers white. River bank.

Croton repens Schlecht. A shrub about 50-60 cm. tall or less;
flowers white. Frequent in thickets.

EUPHORBIA L.
Euphorbia graminea Jacq. A weedy herb. River bank.

JATROPHA L.

Jatropha Curcas L. A small tree about 3 meters tall and 15 cm.
in diameter. River bank.

ANACARDIACEAE

RHUS L.

Rhus terebinthifolia Schl. & Cham. A tree about 3.5 meters
tall and 5 cm. in diameter. In a wet ravine near El Achote.

AQUIFOLIACEAE

ILEX L.

Ilex hondurensis Standl., sp. nov. — PI. XIV. Arbor 4.5-7.5 m.
alta, trunco 7.5-20 cm. diam., omnino glabra, ramulis subteretibus
fuscescentibus lenticellatis rimosis, novellis subangulatis fuscis,
internodiis brevibus; folia modica breviter petiolata rigide coriacea,
petiolo crasso 2-4 mm. longo; lamina elliptica, ovalis vel rotundato-
ovalis 5-6.5 cm. longa 3-4.5 cm. lata, apice late rotundata vel obtusa,
basi rotundata vel obtusa, ubique supra basin remote obscure ad-
presso-crenata, interdum fere integra, supra in sicco sublucida, costa
prominente, nervis obscuris non elevatis, subtus fere concolor, costa
crassiuscula prominula, nervis lateralibus utroque latere ca. 6 angulo
lato adscendentibus fere rectis vix prominulis remote a margine
irregulari ter arcuato-junctis, venulis fere obsoletis; flores feminei
axillares fasciculati pauci (ca. 3 in axilla), pedicellis gracilibus ca. 4
mm. longis; calyx subadpressus 2 mm. latus brevissime obscure 5-
lobus; fructus immaturus globosus 2.5 mm. longus stigmate magno
terminates. — Honduras: Edge of ravine near El Achote, hills above
the plains of Siguatepeque, Dept. Comayagua, August 3, 1936, T. G.
Yuncker, R. F. Dawson & H. R. Youse 6293 (type in Herb. Field
Mus.). In forest near the summit of the ridge above El Achote,
1,800 meters, July 28, 1936, Yuncker, Dawson & Youse 6179. Near
El Achote, 1,350 meters, Yuncker, Dawson & Youse 5838.

FLORA OF HONDURAS 375

Related to the Mexican /. tolucana Hemsl., in which the leaves
are acute at the base and merely obtuse at the apex.

CELASTRACEAE
RHACOMA L.

Rhacoma eucymosa (Loes. & Pitt.) Standl. A shrub. In
dense forest, hills above Lancetilla Valley.

STAPHYLEACEAE
TURPINIA Vent.

Turpinia paniculata Vent. A tree about 7.5 meters tall and
10-12 cm. in diameter. Ravine near El Achote.

SAPINDACEAE
CARDIOSPERMUM L.

Cardiospermum microcarpum HBK. A vine climbing in a
thicket. River bank.

VITACEAE
CISSUS L.

Cissus cacuminis Standl., sp. nov. — Frutex scandens, ramis
crassis subteretibus ferrugineis lenticellis magnis elevatis notatis
glabris vel sparse pilosis rimosis, cortice cito exfoliante, internodiis
plus minusve elongatis; folia mediocria longipetiolata in sicco mem-
branacea et fragilia, simplicia, petiolo gracili 1.3-4.5 cm. longo sparse
pilis longis laxis albidis piloso vel glabrato; lamina ovata vel late
ovata 4-9.5 cm. longa 3-5 cm. lata sensim acuminata, basi truncata
vel saepius breviter late cordata, ubique remote vel dense serrata,
serraturis angustis subulato-apiculatis adpressis, supra in sicco
viridis primo sparse villosa cito glabrata, costa nervisque prominenti-
bus, subtus concolor primo sat dense pilis longis pallidis vel brun-
nescentibus laxis villosa cito glabrata, basi 5-nervia, costa gracili
prominente supra basin nervos laterales ca. 3 emittente, venulis
obviis sed non elevatis; cymae oppositifoliae gracili ter pedunculatae
laxe multiflorae ca. 3 cm. latae et aequilongae, ramis sat dense villosis,
floribus lutescentibus parvis umbellatis, pedicellis gracilibus Z-4 mm.
longis adpresso-pilosis; calyx patelliformis vix ultra 1 mm. latus
undulatus; petala paullo ultra 1 mm. longa. — Honduras: Climbing
in trees in wet ravine near El Achote, hills above the plains of
Siguatepeque, Dept. Comayagua, 1,350 meters, August 5, 1936, T.G.
Yuncker, R. F. Dawson & H. R. Youse 6330 (type in Herb. Field

376 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

Mus.)- In forest near the summit of the range above El Achote,
in cloud zone, 1,800 meters, Yuncker, Dawson & Youse 6005.

Although doubtless related to the common C. sicyoides L., the
present plant is obviously different in general appearance as well as
in some of the minor details, and I have no doubt that it represents
a well marked species.

Cissus Martiniana Woodson & Seibert. Climbing over shrubs
in forest near the summit of the range above El Achote. A recently
described species, the type from Panama, occurring also in Costa
Rica.

Cissus rhombifolia Vahl. Scrambling over shrubs in a wet
ravine near El Rincon.

Cissus salutaris HBK. Climbing in a thicket on a rocky
hillside.

VITIS L.

Vitis tiliifolia Humb. & Bonpl. A woody vine climbing over
shrubs in a wet ravine near El Achote.

MALVACEAE
MALVAVISCUS Cav.

Malvaviscus arboreus Cav. A shrub about 3.5 meters tall;
corolla scarlet. In a wet ravine near the summit of the range above
El Achote.

SIDA L.

Sida ciliaris L. A low shrub; flowers salmon. On plains.
Sida linifolia Juss. A low, half-woody shrub. In a thicket.

ACTINIDIACEAE
SAURAUIA Willd.

Saurauia leucocarpa Schlecht. A small tree about 4.5 meters
tall and 5 cm. in diameter; fruit succulent, with scattered, small,
brown seeds in a viscid pulp. In forest near the summit of the range
above El Achote. An addition to the known trees of Honduras.

Saurauia villosa DC. A tree 6-7 meters tall and 15 cm. in
diameter; flowers white, fragrant. In forest near the summit of the
range above El Achote.

FLORA OF HONDURAS 377

OCHNACEAE
SAUVAGESIA L.

Sauvagesia tenella Lam. An herb; flowers pink, delicate. In a
wet pasture.

MARCGRAVIACEAE
MARCGRAVIA L.

Marcgravia rectiflora Triana & Planch.? Climbing on a tree on
stream bank, bottom of a ravine near El Achote.

SOUROUBEA Aubl.

Souroubea guianensis Aubl. A woody vine; flowers and
pedicels orange or reddish. In a ravine near El Achote; also on a
tree near the shore of Lake Yojoa.

THEACEAE
EURYA Thunb.

Eurya guatemalensis Bonn. Sm. A small tree about 4.5
meters tall and 8 cm. in diameter; flowers white, fleshy. In forest
near the summit of the range above El Achote. The species is new
to the Honduran flora.

TERNSTROEMIA L. f.

Ternstroemia Tepezapote Cham. & Schl. A tree 6-7 meters
tall and 15-20 cm. in diameter; leaves leathery, dark green, shiny.
River bank.

GUTTIFERAE
CLUSIA L.

Clusia Salvinii Bonn. Sm. A tree 7-8 meters tall and 30 cm.
in diameter. The fruit opens in a characteristically stellate form.
In a wet ravine near El Achote.

HYPERICUM L.

Hypericum uliginosum HBK. Flowers orange-yellow. In a
subalpine bog near El Achote.

VISMIA Vand.

Vismia guianensis (Aubl.) Pers. A tree about 9 meters tall and
15 cm. in diameter. In pine-oak association, El Achote. Not
recorded previously for Honduras.

378 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

VIOLACEAE
HYBANTHUS Jacq.

Hybanthus oppositifolius (L.) Taub. An herb with some-
what purplish flowers. River bank.

FLACOURTIACEAE
OLMEDIELLA Baill.

Olmediella Betschleriana (Goepp.) Loes. A shrub or small
tree about 3.5 meters tall. In thicket near El Achote. For notes
regarding this interesting plant see Trop. Woods 32: 17. 1932.

XYLOSMA Forst.

Xylosma Hemsleyana Standl. A shrub about 1 meter tall.
In a dry ravine east of Siguatepeque.

PASSIFLORACEAE
PASSIFLORA L.

Passiflora ciliata Dryander. Climbing over shrubs in a thicket,
edge of forest, Lancetilla Valley.

Passiflora foetida L. var. gossypiifolia (Desv.) Mast. Climb-
ing in a thicket, river bank.

BEGONIACEAE
BEGONIA L.

Begonia tovarensis Klotzsch. Petals pale pink; stems light red.
Subalpine bog near El Achote; also in forest, Lake Yojoa.

Begonia Yunckeri Standl., sp. nov.— PI. XV. Herba ut videtur
erecta caulescens ramosa gracilis, ramis sparse pilis longis laxis
villosis vel fere glabris, internodiis elongatis; stipulae hyalinae
caducae ad 1.5 cm. longae albidae nervosae apice obtusae vel rotun-
datae glabrae; folia modica longissime petiolata in sicco membrana-
cea, petiolo gracillimo 2.5-8.5 cm. longo sparse laxe villoso vel fere
glabro; lamina ambitu oblongo-ovata vel lanceolato-oblonga 6-13
cm. longa 2-5.5 cm. lata, petiolo lateraliter inserto, longissime
anguste attenuato-acuminata, basi insigniter obliqua, altero latere
semiobtusa, lobo postico late rotundato vel angulato, plus minusve
breviter lobata, in toto margine inaequaliter dentata, dentibus
breviter vel longe mucronatis et interdum villoso-ciliatis, supra in
sicco viridis sublucida sparse pilis brevibus patentibus pallidis villosa
vel glabrata, subtus ad nervos dense villosa, aliter glabra, basi vulgo

FLORA OF HONDURAS 379

7-nervia; flores monoeci in umbellas axillares 3-4-floras longe graci-
liter pedunculatas dispositi, pedicellis gracillimis ad 8 mm. vel in
statu fructifero ad 15 mm. longis sparse breviter villosulis vel glandu-
loso-pilosulis; sepala exteriora floris masculi ovato-rotundata 6 mm.
longa et fere aequilata alba glabra apice rotundata basi subcordata;
capsula (perfecta non visa) 6 mm. longa atque 4 mm. lata basi obtusa,
angulis angustissime alatis, alis vix 1 mm. longis vel obsoletis. —
Honduras: On forest floor, forest near the summit of the range above
El Achote, in cloud zone, above the plains of Siguatepeque, Dept.
Comayagua, 1,800 meters, July 28, 1936, T. G. Yuncker, R. F. Dawson
& H. R. Youse 6148 (type in Herb. Field Mus.).

"Corolla white. Stems up to 1 cm. in diameter."

CACTACEAE
CEREUS L.

Cereus hondurensis Schum. Forming large patches on rocks
near the shore about 8 km. east of Tela.

EPIPHYLLUM Haw.

Epiphyllum caudatum (Vaupel) Britt. & Rose. On a tree at
edge of clearing in Lancetilla Valley.

Epiphyllum strictum (Lem.) Britt. & Rose? On a tree near
shore of Lake Yojoa; flowers white, 5-8 cm. long. Also on fallen log
in a deep, moist ravine near El Rincon.

RHIPSALIS Gaertn.

Rhipsalis Cassutha Gaertn. Forming large clumps on a tree
near the shore of Lake Yojoa. Corolla white, fruit pinkish.

LYTHRACEAE
CUPHEA Adans.

Cuphea carthagenensis (Jacq.) Macbr. An herb; flowers
magenta. In a wet meadow.

Cuphea Hookeriana Walp. A low shrub less than 1 meter tall ;
flowers white to purple, attractive. In a ravine near El Achote.

Cuphea utriculosa Koehne. A low, sprawling shrub about
30-50 cm. tall; flowers violet. Frequent in mud along river bank.

MYRTACEAE
CALYPTRANTHES Sw.

Calyptranthes hondurensis Standl. A small tree or shrub up

380 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

to 3^4 meters tall. In a thicket near El Achote. The species is
known only from this region.

EUGENIA L.

Eugenia apodantha Standl., sp. nov. — Arbuscula 4.5 m. alta,
trunco 7 cm. diam., ut videtur dense ramosa, ramulis crassiusculis
teretibus ochraceis rimosis, cortice exfoliante, novellis angulatis
brunnescentibus dense pilosis, internodiis brevibus; folia modica fere
sessilia coriacea, petiolo crasso ca. 2 mm. longo piloso; lamina oblonga
vel elliptico-oblonga 7-9 cm. longa 3-3.5 cm. lata acuta, apice ipso
obtuso, basi anguste rotundata vel obtusa, supra in sicco griseo-
viridis dense minute puncticulata, sparse vel subdense pilis longi-
usculis laxis pilosa vel glabrata, costa impressa, nervis obsoletis,
subtus paullo pallidior densiuscule pilis lutescentibus sublucidis
pilosa, costa gracillima prominente, nervis lateralibus utroque latere
ca. 10 tenerrimis obscuris vix prominulis fere rectis angulo latiusculo
divergentibus prope marginem in nervum collectivum fere regularem
conjunctis, venis obsoletis; flores axillares solitarii sessiles; fructus
immaturus semiglobosus ca. 4 mm. longus et 5 mm. latus basi rotun-
datus densissime pilis brevibus patentibus ochraceis pilosus; sepala
persistentia 4 oblongo-ovata obtusa 4-5 mm. longa patentia utrinque
dense breviter pilosa; stylus gracilis glaber 8 mm. longus.- — River
bank, plain near Siguatepeque, Dept. Comayagua, 1,050 meters,
July 9, 1936, T. G. Yuncker, R. F. Dawson & H. R. Youse 5766 (type
in Herb. Field Mus.). Also referable here is a sterile specimen:
Thicket along river bank near Siguatepeque, a tree of 4.5 meters,
February, 1928, Standley 56057. It is not improbable that both
collections are from the same individual.

The crushed leaves are said to be aromatic. Because of its soli-
tary, quite sessile flowers this can scarcely be confused with any
other species of Eugenia found in Central America.

Eugenia axillaris (Sw.) DC. A tree 3.5-4.5 meters tall and 10
cm. in diameter; flowers white. River bank.

Eugenia Doubledayi Standl. A small tree about 6 meters tall
and 8-10 cm. in diameter; flowers white. River bank. The species
is known only from the region of Siguatepeque.

Eugenia vincentina Krug & Urban. A small tree 4-5 meters
tall and 15 cm. in diameter. River bank.

Eugenia Yunckeri Standl., sp. nov. — Frutex 2.5 m. altus,
ramulis gracilibus teretibus brunnescentibus vel fuscis, novellis

FLORA OF HONDURAS 381

ferrugineis densissime velutine patenti-pilosis, internodiis brevibus
vel elongatis; folia modica elliptico-oblonga vel elliptica 7-14 cm.
longa 3-7 cm. lata sensim vel subabrupte acuminata vel acuta, apice
ipso obtuso, interdum apice fere obtusa et tantum obtuse apicata,
basi acutisuscula vel obtusa, plus minusve lucida, sparse molliter
pilosa, serius glabrata, nervis venisque non elevatis, subtus paullo
pallidior, ubique sat dense pilis patentibus pallidis velutino-pilosa,
costa gracili elevata, nervis lateralibus utroque latere ca. 9 angulo
latiusculo adscendentibus rectis vel paullo arcuatis remote a margine
irregulariter arcuato-conjunctis, venis vix obviis, ubique dense sub-
grosse punctata, margine interdum anguste incurvo; flores in axillis
fasciculati pauci fere sessiles, pedicellis vix ad 2 mm. longis crassis
pilosis; hypanthium hemisphaericum ca. 3 mm. latum dense molliter
pilosum; sepala 4 rotundata 2.5-3 mm. longa apice late rotundata
ciliata, extus breviter patenti-pilosa, intus dense breviter sericea.—
Honduras: In thicket near trail west of El Achote, hills above the
plains of Siguatepeque, Dept. Comayagua, 1,300 meters, July 28,
1936, T.G. Yuncker, R.F. Dawson & H. R. Youse 6178 (type in Herb.
Field Mus.). Also (sterile material), El Achote, open mountain
forest, a tree 12 meters high, October, 1932, J. B. Edwards 499.

Similar toE. Banghamii Standl., a frequent species of the Panama
Canal Zone.

PSIDIUM L.

Psidium Guajava L. A shrub about 3 meters tall; flowers
white. On a rocky hillside.

Psidium molle Bertol. A shrub 2-3 meters tall; flowers white.
River bank.

Psidium Oerstedeanum Berg. A low shrub with white flowers.
On a rocky hillside.

MELASTOMACEAE
ARTHROSTEMMA Ruiz & Pavon

Arthrostemma fragile Lindl. Corolla pink. Side of a ravine
near El Achote.

BLAKEA L.

Blakea aeruginosa Standl., sp. nov. — Arbor 7.5-9 m. alta,
trunco 12-15 cm. diam., ramulis crassis obtuse angulatis densissime
pilis run's setoso-pilosis vel fere tomentosis, internodiis valde abbre-
viatis; folia majuscula longipetiolata coriacea, petiolo crassiusculo
usque 4.5 cm. longo supra anguste canaliculate densissime ferru-

382 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

gineo-setuloso; lamina elliptico-oblonga 11-16 cm. longa 5-8 cm. lata
abrupte breviter caudatrf-acuminata, acumine obtuso vix ultra 8 mm.
longo, basi anguste rotundata, fere Integra, supra in sicco griseo-
viridis glabra lucida, nervis solemniter impressis, subtus primo sat
dense breviter femigineo-setulosa vel serius glabrata, triplinervia
vel quintuplinervia (nervis exterioribus multo tenerioribus et mar-
ginibus saepe recurvis fere occultis), nervis 3 mediis crassis elevatis
apice junctis, nervis lateralibus numerosis approximatis rectis angulo
paullo minore quam recto divergentibus prominulis; flores axillares
solitarii, pedunculo crasso dense ferrugineo-setuloso 1.5-2 cm. longo;
bracteae 4 fere distinctae lanceolatae vel anguste lanceolatae,
exteriores ca. 2 cm. longae longissime attenuatae, extus densissime
pilis ferrugineis hispidulis setulosae, intus fere glabrae sed apicem
versus stellato-puberulae; calycis tubus hemisphaericus ca. 8 mm.
longus et aequilatus fere glaber, limbo (lobis inclusis) usque 14 mm.
longo extus ferrugineo-setuloso, lobis lanceolato-oblongis ca. 8 mm.
longis obtusis vel acutiusculis. — Honduras: In forest near the summit
of the ridge above El Achote, in cloud zone, above the plains of
Siguatepeque, Dept. Comayagua, 1,850 meters, August 1, 1936, T. G.
Yuncker, R. F. Dawson & H. R. Youse 6267 (type in Herb. Field
Mus.).

The corolla, which I have not seen, is described as pink. Among
Central American species this is conspicuous because of the abundant,
rusty pubescence found on almost all parts of the plant.

CENTRADENIA G. Don

Centradenia Bernoulli! Cario. A shrub 60-75 cm. tall. On
rocks in a ravine near El Achote.

CLIDEMIA D. Don

Clidemia dentata Don. A shrub about 3 meters tall, spreading.
In forest near the summit of the range above El Achote.

Clidemia hirta (L.) Don. A shrub nearly 2 meters tall; flowers
white. River bank.

Clidemia rubra (Aubl.) Mart. A shrub about 1 meter tall.
Thicket.

CONOSTEGIA D. Don

Conostegia subhirsuta DC. A tree 7-8 meters tall and 7-10
cm. in diameter; flowers white, aromatic. Side of a ravine near El
Achote.

FLORA OF HONDURAS 383

Conostegia xalapensis (Bonpl.) Don. A shrub 2-3 meters tall;
flowers pinkish; fruit smooth, dark purple. Common on the plains
and in the pine forest.

MICONIA Ruiz & Pavon

Miconia albicans Triana. A shrub 1-2 meters tall; fruit red.
Common in thickets on the plains and in the pine belt.

Miconia globulifera Naud. A small tree about 3 meters tall
and 3-5 cm. in diameter. Bottom of a ravine near El Achote. New
for the flora of Honduras.

Miconia humilis Cogn. A shrub about 2 meters tall. Rim of a
ravine near El Achote.

Miconia longifolia DC. A small tree 4.5-6 meters tall and 5-8
cm. in diameter; fruit purplish. In a thicket in a wet ravine near El
Rincon. Not recorded previously for the flora of Honduras.

Miconia Schlechtendalii Cogn. A low shrub 0.5-1 meter tall.
Thicket.

TIBOUCHINA Aubl.

Tibouchina longifolia (Vahl) Baill. A shrub 60-90 cm. tall;
petals white. In pine forest near El Achote.

ONAGRACEAE
JUSSIAEA L.
Jussiaea linifolia Vahl. In a marsh.

Jussiaea peruviana L. A small tree 3.5-4 meters tall. In a bog
near El Achote.

Jussiaea repens L. In a marsh.

Jussiaea suffruticosa L. A weedy plant, common in moist,
waste places.

ARALIACEAE

ARALIA L.

Aralia humilis Cav. A shrub about 2 meters tall. On a rocky
hillside. The species is new for Honduras.

GILIBERTIA Ruiz & Pavon

Gilibertia leptopoda Donn. Sm. A tree about 6 meters tall
and 8-10 cm. in diameter. In a wet ravine near El Achote. Not
reported previously from Honduras.

384 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

OREOPANAX Dene. & Planch.

Oreopanax Liebmannii March. A tree about 15 meters tall
and 60 cm. in diameter. In dense forest near the summit of the range
above El Achote. The tree is new for the Honduran flora.

Oreopanax peltatum Linden. A small tree about 4.5 meters
tall. Ravine near El Achote. Reported previously from the same
region under the name 0. Salvinii Hemsl.

Oreopanax xalapense (HBK.) Dene. & Planch. A tree about
3.5 meters tall and 5 cm. in diameter. In forest near the summit of
the range above El Achote. An addition to the known trees of
Honduras.

UMBELLIFERAE
ERYNGIUM L.

Eryngium caricinum Standl. In an old, wet pasture. The
species is known only from Siguatepeque.

HYDROCOTYLE L.

Hydrocotyle umbellata L. Corolla greenish white. In a sub-
alpine bog near El Achote; also in a low, wet area about 12 km. west
of Tela.

SPANANTHE Jacq.

Spananthe paniculata Jacq. Corolla yellow. River bank,
about 6 km. north of Siguatepeque.

CLETHRACEAE
CLETHRA L.

Clethra hondurensis Britton. A tree about 6 meters tall and
10 cm. in diameter; flowers greenish white. Rocky hillside.

PYROLACEAE
CHIMAPHILA Pursh

Chimaphila dasystemma Torr. Flowers white with green
pistils, attractive. In partial shade, hillside near El Achote. This
species and the following one are new to the flora of Honduras.

MONOTROPA L.

Monotropa coccinea Zucc. Entire plant bright red. In humus
of forest floor near the summit of the range above El Achote.

FLORA OF HONDURAS 385

ERICACEAE
ARBUTUS Adans.

Arbutus xalapensis HBK. A tree about 4.5 meters tall and
5-8 cm. in diameter; stems and petioles red. In pine forest near
El Achote.

BEFARIA Mutis

Befaria mexicana Benth. A small tree about 4.5 meters tall
and 5-7 cm. in diameter; inflorescence viscid; flowers white, fragrant.
In a ravine near El Achote. An addition to the known trees of the
country.

SPHYROSPERMUM Poepp. & Endl.

Sphyrospermum ma jus Griseb. Fruit light purple, fleshy.
Climbing on a tree in forest near the summit of the range above El
Achote. The species is new for Honduras.

THIBAUDIA Ruiz & Pavon

Thibaudia costaricensis Hoer. A tree 15-18 meters tall and
about 60 cm. in diameter; fruit purplish black. In dense forest at
the summit of the range above El Achote.

MYRSINACEAE
ARDISIA Sw.

Ardisia compressa HBK. A small tree about 4.5 meters tall
and 8 cm. in diameter; flowers white; fruit black. Side of a ravine
near El Achote.

PARATHESIS Hook. f.

Parathesis Donnell-Smithii Mez. A small tree about 4.5
meters tall and 10-12 cm. in diameter; flowers, pedicels, etc. flesh-
colored. Side of a ravine near El Achote. New for the Honduran flora.

SAPOTACEAE
CALOCARPUM Pierre

Calocarpum mammosum (L.) Pierre. A tree about 15 meters
tall and 60 cm. in diameter. River bank.

CHRYSOPHYLLUM L.

Chrysophyllum oliviforme L. A small tree 7-8 meters tall
and 45 cm. in diameter; leaves bright green above, brown-silky
beneath. River bank.

386 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

STYRACACEAE
STYRAX L.

Styrax argenteus Presl. A tree 9-10 meters tall and 50 cm. in
diameter; flowers waxy- white with yellow stamens. River bank.

Styrax glabrescens Benth. A tree 6-7.5 meters tall and 12-15
cm. in diameter; fruit with a hard seed and thin pericarp. In forest
near the summit of the range above El Achote. The species is new
for Honduras.

OLEACEAE
FRAXINUS L.

Fraxinus hondurensis Standl., sp. nov. — Arbor 18-metralis,
trunco 60 cm. diam., corona depressa lata, ramulis teretibus cras-
siusculis lenticellis paucis pallidis parvis conspersis, novellis dense
minute patenti-pilosulis; folia modica graciliter 4-5.5 cm. longe
petiolata, petiolo exalato minute pilosulo, rhachi gracili exalata vel
obscure marginata minute pilosula usque 7 cm. longa; foliola 5,
lateralia sessilia, vel inferiora petiolulata, terminale longe graciliter
petiolulatum, lanceolata vel oblongo-lanceolata 4.5-12.5 cm. longa
et ultra 1.5-4.5 cm. lata, plerumque longissime angustissime attenu-
ato-acuminata, terminali basi longiacuminato vel acuminato, laterali-
bus basi saepe valide obliquis et acutis vel acuminatis, in sicco laete
viridia, supra lucida, sparse minute puberula, costa ut nervis venisque
prominulis, subtus fere concoloria, subsparse breviter patenti-
pilosula, costa gracili elevata, nervis lateralibus utroque latere ca. 9
angulo acuto adscendentibus, venulis prominulis arete reticulatis,
foliolis remote inaequaliter serrulatis vel fere integris. — Honduras:
In grove near river bank, plains near Siguatepeque, Dept. Comaya-
gua, 1,050 meters, July 23, 1936, T. G. Yuncker, R. F. Dawson &
H. R. Youse 6038 (type in Herb. Field Mus.).

"Bark furrowed. A fine, spreading tree." The genus is new to
Honduras and probably to Central America, this being the southern-
most occurrence known for it at the present time in America. The
material is, unfortunately, sterile, but in leaf characters it appears
to be so definitely different from any of the Mexican species that I do
not hesitate to describe it as new.

LOGANIACEAE

SPIGELIA L.

Spigelia Humboldtiana Cham. & Schlecht. An herb with
white flowers. In shade, river bank.

FLORA OF HONDURAS 387

GENTIANACEAE
LEIPHAIMOS Schlecht. & Cham.

Leiphaimos parasiticus Schlecht. & Cham. A saprophytic
herb, pale red below, tawny toward the top. In forest near the
summit of the range above El Achote.

MENYANTHACEAE
LIMNANTHEMUM Gmel.

Limnanthemum Humboldtianum (HBK.) Griseb. Growing
in water. Corolla white. Marsh near Siguatepeque.

APOCYNACEAE
MANDEVILLA Lindl.

Mandevilla tubiflora (Mart. & Gal.) Woodson. Climbing in a
thicket near El Rincon. Corolla greenish yellow.

ASCLEPIADACEAE

ASCLEPIAS L.

Asclepias curassavica L. A common weed in waste places
about Siguatepeque.

Asclepias guatemalensis Donn. Sm. Corolla yellowish green,
horns light violet. Rocky hillside.

METASTELMA R. Br.

Metastelma pedunculare Dene.? Sprawling over a fallen log
in forest near the summit of the range above El Achote.

VINCETOXICUM Walt.

Vincetoxicum porphyranthum Standl., sp. nov.— Volubilis,
gracilis, caule viridi dense minute puberulo atque pilis longis rigidis
patentibus fulvis hirsute, internodiis foliis longioribus; folia inter
minora petiolata membranacea, petiolo gracili 10-17 mm. longo
dense longihirsuto; lamina oblongo-ovata 3.5-6 cm. longa 1.5-2.5 cm.
lata longe anguste attenuato-acuminata, basi anguste rotundata
et breviter cordata, sinu lato aperto, utrinque praesertim ad nervos
venasque pilis longissimis patentibus hirsuta, supra in sicco viridis,
venis nervisque prominulis, subtus paullo pallidior, costa gracili
prominente, nervis lateralibus utroque latere 3-4 tenerrimis angulo
angusto adscendentibus, venulis inconspicuis laxe reticulatis; um-
bellae axillares solitariae laxe pauciflorae, pedunculo vix ultra 5 mm.
longo puberulo et hirsuto, pedicellis plerumque 6-7 mm. longis dense

388 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

viscido-puberulis et sparse hirsutis, bracteis filiformibus viridibus
ca. 5 mm. longis; calyx adpressus extus sparse hirsutus purpureus
5-6 mm. longus fere ad basin 5-lobus, lobis anguste lanceolato-
oblongis anguste longiattenuatis; corolla in sicco fusco-purpurea 7-8
mm. longa subrotata, extus sparse puberula et hispidula, ad medium
5-loba, lobis triangulari-ovatis acutiusculis, intus sparse puberulis et
sparse breviter hirsutis vel interdum fere glabris. — Honduras: Climb-
ing in thicket near river, Siguatepeque, Dept. Comayagua, 1,050
meters, July 5, 1936, T.G. Yuncker, R.F. Dawson & H. R. Youse5676
(type in Herb. Field Mus.).

Vincetoxicum trichoneuron Standl., sp. nov. — Scandens,
caule gracillimo viridi subtereti puberulo et ad sulcos dense brunneo-
piloso, internodiis foliis multo longioribus; folia modica petiolata
membranacea, petiolo 1-2 cm. longo crassiusculo densissime brunneo-
piloso; lamina oblonga 7.5-12 cm. longa 3-4 cm. lata apicem versus
paullo angustata longiacuminata, basi late rotundata vel subtruncata,
supra in sicco viridis, ad nervos venasque adpresso-pilosa, costa
nervisque manifestis sed vix plus quam prominulis, subtus multo
pallidior, molliter pilis brevibus subpatentibus brunnescentibus
pilosa, ad costam densissime pilis longis brunneis rigidulis pilosa,
costa gracili elevata, nervis lateralibus utroque latere ca. 6 angulo
latiusculo adscendentibus irregularibus, venis manifestis conspicuis
laxe reticulatis; umbellae axillares solitariae 1-3-florae, pedunculo
crasso vix 5 mm. longo, pedicellis usque 1 cm. longis breviter pilosulis
et sparse longipilosis; calyx viridis 7-8 mm. longus extus breviter
pilosulus fere ad basin 5-lobus, lobis oblongo-lanceolatis longiacumi-
natis adpressis; corolla olivacea rotata fere 2 cm. lata extus minute
breviter pilosula, intus glabra, ad medium 5-loba, lobis orbicularibus
apice late rotundatis, intus nervis 5 longitrorsis rectis notatis et
arete reticulato-venosis. — Honduras: Twining over shrubs in pine-
oak forest near El Achote, hills above the plains of Siguatepeque,
Dept. Comayagua, 1,350 meters, July 30, 1936, T. G. Yuncker, R. F.
Dawson & H. R. Youse 6211 (type in Herb. Field Mus.).

The type specimen bears a single opened corolla that unfortu-
nately is incomplete.

CONVOLVULACEAE
CUSCUTA L.

Cuscuta strobilacea Liebm. Abundant on various herbaceous
and woody hosts along river banks.

FLORA OF HONDURAS 389

IPOMOEA L.

Ipomoea cathartica Poir. An herbaceous vine; flowers large,
purplish. Climbing in a thicket. River bank.

Ipomoea digitata L. Corolla reddish. Climbing in a thicket
near the beach west of Tela.

Ipomoea muricata Cav. Flowers pinkish; roots tuberous. On
a rocky hillside.

Ipomoea silvicola House. Corolla purplish. Climbing in a
thicket. River bank about 6 km. north of Siguatepeque.

BORAGINACEAE
HELIOTROPIUM L.

Heliotropium angiospermum Murr. Corolla white. In
waste places near Potrerillos.

Heliotropium indicum L. A common weedy herb about 1
meter tall ; flowers lavender. In waste places.

TOURNEFORTIA L.

Tournefortia glabra L. A shrub; fruit white. River bank
about 6 km. west of Siguatepeque.

Tournefortia hirsutissima L. A scandent shrub. River bank
about 5 km. northwest of Siguatepeque.

Tournefortia volubilis L. A shrub about 3 meters tall. River
bank.

VERBENACEAE
CALLICARPA L.

Callicarpa acuminata HBK. A shrub about 2.5 meters tall;
corolla white. In a thicket near Potrerillos.

CITHAREXYLUM L.

Citharexylum Mocinni D. Don. A shrub 1.5-2 meters high.
Wet ravine near El Achote. The species is new for Honduras.

LANTANA L.

Lantana Camara L. A shrub about 3 meters tall; flowers
orange. In a thicket near the shore of Lake Yojoa.

Lantana involucrata L. A shrub about 1 meter tall; flowers
pink to lavender. Frequent on the plains.

390 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

LIPPIA L.
Lippia dulcis Trev. Flowers white. River bank.

Lippia Kellermanii Greenm. A tree about 6 meters tall and
7-10 cm. in diameter; corolla pale yellow. River bank about 6 km.
west of Siguatepeque.

VERBENA L.
Verbena Carolina L. A weedy plant in waste places.

Verbena littoralis HBK. A weedy herb about a meter tall;
flowers lavender. Common in waste places.

LABIATAE
HYPTIS Jacq.

Hyptis asperifolia Standl. A shrub 1.5-2 meters tall. Side
of a ravine near El Achote. The species is known only from this
region.

SALVIA L.

Salvia siguatepequensis Standl. A shrub about 2 meters tall;
flowers scarlet. Side of a ravine near El Achote. Known only from
the region of Siguatepeque.

STACHYS L.

Stachys guatemalensis Epling. A decumbent herb with
rather long stems. River bank.

SOLANACEAE

CAPSICUM L.

Capsicum macrophyllum (HBK.) Standl. A shrub or small
tree about 4.5 meters tall; flowers pale yellow; berries orange.
Ravine near El Achote.

CESTRUM L.

Cestrum nocturnum L. var. mexicanum Schulz. A shrub
1.5 meters tall. Near El Achote.

CYPHOMANDRA Sendtn.

Cyphomandra heterophylla Bonn. Sm. A shrub or small tree
about 3.5 meters tall. In a wet ravine near El Achote. A synonym
of this is C. mollicella Standl.

LYCIANTHES Hassler

Lycianthes sideroxyloides (Schlecht.) Bitter. A scandent
shrub. In a thicket near the summit of the range above El Achote.

FLORA OF HONDURAS 391

MERINTHOPODIUM Bonn. Sm.

Merinthopodium neuranthum (Hemsl.) Donn. Sm. Corolla
green without, purplish within. A small shrub, on a tree trunk in
forest near Lancetilla Valley.

PHYSALIS L.

Physalis ingrata Standl., sp. nov. — Herba perennis erecta(?)
ramosa, ramis viridibus gracilibus pilis tenuibus albidis brevibus
paten tibus pilosis, internodiis brevibus vel elongatis; folia inter
minora longipetiolata herbacea, petiolo gracili 0.8-2 cm. longo
dense breviter albido-villosulo; lamina ovata vel late ovata 2-5 cm.
longa 1.5-2.5 cm. lata acuta vel subacuta, rarius attenuata, basi plus
minusve inaequali late rotundata ad obtusa, supra sparse breviter
villosula, subtus viridis densius villosula, margine integro vel paullo
undulato; flores axillares solitarii cernui (?), pedicellis fere filiformibus
usque 1.5 cm. longis dense villosulis; calyx florifer 8 mm. longus
campanulatus viridis dense breviter villosulus, lobis tubo aequilongis
lanceolate- triangularibus attenuatis; corolla lutea ca. 12 mm. longa
extus sparse puberula, fauce maculis 5 magnis atropurpureis notata;
calyx fructifer anguste ovoideus basi ut videtur non depressus 1.5
cm. longus vel ultra viridis subdense breviter villosulus, lobis tubo
accrescente multo brevioribus. — Honduras: On rocky hillsides
above Siguatepeque, Dept. Comayagua, 1,200 meters, July 14, 1936,
T. G. Yuncker, R. F. Dawson & H. R. Youse 5865 (type in Herb.
Field Mus.). Collected also in the same region, July 11, No. 5794-

SARACHA Ruiz & Pavon

Saracha procumbens (Cav.) Ruiz & Pav. An herb, about 1
meter tall ; flowers greenish white. River bank.

SOLANUM L.

Solatium aculeatissimum Jacq. A shrub about 1.5 meters
tall; flowers greenish white. In a wet ravine near El Achote.

Solanum bicolor Mill. A shrub about 1 meter tall. In a
thicket, margin of the forest at the summit of the range above El
Achote.

Solanum erythrotrichum Fernald. A somewhat scandent
shrub; flowers greenish white to purplish. Frequent on the plains
and to the summit of the range above El Achote.

Solanum guanicense Urban? In an old pasture.

392 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

Solanum laurifolium Mill. A shrub about 1.5 meters tall;
flowers purplish. In a thicket on the plains.

Solanum madrense Fernald. A shrub about 2 meters tall;
flowers white. Ravine, El Achote.

Solanum nigrum L. In a thicket, river bank.

Solanum nudum L. A shrub about 2.5 meters tall; flowers
white. In a thicket near El Rincon.

Solanum Seaforthianum Andr. A climbing, semi-woody vine
with very attractive, purple flowers. Cultivated in gardens but
collected as an escape on a river bank.

Solanum umbellatum Mill. A shrub 3-4 meters tall; flowers
white. River bank.

SCROPHULARIACEAE

BACOPA Aubl.
Bacopa rotundifolia (Michx.) Wettst. In a swamp.

BUCHNERA L.

Buchnera pusilla HBK. An herb; flowers violet. In an old,
wet pasture.

ESCOBEDIA Ruiz & Pavon

Escobedia laevis Cham. & Schlecht. Herb; flowers white. In
wet grassland. The genus is new for Honduras.

ILYSANTHES Raf.

Ilysanthes inaequalis (Walt.) Pennell. Corolla blue at the
base, becoming white toward the edges. In a subalpine bog near
El Achote.

LENTIBULARIACEAE

UTRICULARIA L.
Utricularia mixta Barnh.? In a marsh.

Utricularia pusilla Vahl. Corolla golden yellow. In a marshy,
alpine meadow near El Achote.

BIGNONIACEAE
AMPHILOPHIUM Kunth

Amphilopium pilosum Standl., sp. nov. — Frutex scandens,
ramis quadrangulatis densissime pilis patentibus fulvis pilosis,
internodiis elongatis; cirrhi pilosi apice breviter trifidi; folia 3.5-4.5

FLORA OF HONDURAS 393

cm. longe petiolata, petiolulis 3-4 cm. longis dense longipilosis;
foliola 2 membranacea rotundato-ovata vel late ovato-elliptica 8-12
cm. longa 5.5-9 cm. lata abrupte breviter acuminata, basi breviter
cordata vel truncata, supra viridia sublucida sparse vel dense pilis
simplicibus pilosa, subtus fere concoloria molliter, ad nervos densius,
patenti-pilosa, basi 5-7-nervia, nervis pallidis prominentibus, venulis
laxe reticulatis, inter venas dense minute glanduloso-puncticulata;
flores purpurascentes in cymas trifloras dispositi, cymis racemosis
pedunculatis, paniculis angustis usque 25 cm. longis angustis, ramis
densissime fulvopilosis, flore centrali cymae breviter crasse pedi-
cellate, lateralibus longipedicellatis; calyx pro genere typicus extus
dense pilosus ca. 8 mm. longus, lobis late rotundatis intus minute
puberulis; corolla 2.5 cm. longa extus glabra. — Honduras: Climbing
in thicket along river bank on plains near Siguatepeque, Dept.
Comayagua, 1,050 meters, July 25, 1936, T.G. Yuncker, R.F. Dawson
& H. R. Youse 6683 (type in Herb. Field Mus.). Also, from the
same region, July 8, No. 5788.

Two other species of the genus found in Central America are
conspicuously unlike the present plant in having fine, stellate pubes-
cence.

GESNERIACEAE
ACHIMENES P. Br.

Achimenes grandiflora (Schiede) DC. Corolla purple; stems
red. Rocky hillside.

Achimenes longiflora DC. Flowers attractive, blue. Rocky
hillside.

BESLERIA L.

Besleria laxiflora Benth. A shrub 1-2 meters tall; corolla
scarlet. In a wet ravine near El Achote.

COLUMNEA L.

Columnea pilosissima Standl.? Creeping on a rotten tree
trunk. Near the shore of Lake Yojoa.

Columnea rubricaulis Standl., sp. nov. — Frutex subscandens
epiphyticus ramosus, ramis crassis ferrugineo-rubris subteretibus,
novellis dense foliatis pilis longis adpressis albis pilosis, internodiis
brevissimis; folia inter minora in sicco subcoriacea, petiolo crasso
vix ultra 3 mm. longo adpresso-piloso ; lamina lineari-oblonga 3.5-
5.5 cm. longa ca. 8 mm. lata apicem angustum versus sensim atte-

394 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

nuata, basi paullo obliqua obtusa vel subacuta, supra in sicco cinereo-
viridis opaca glabra, costa impressa, nervis obsoletis, subtus pallidior,
sparse pilis brevibus albidis adpressis pilosa vel fere glabra, ad costam
crassiusculam pilis longioribus albidis et ferrugineis intermixtis
adpresso-pilosa, nervis obsoletis; flores axillares, pedicellis ut videtur
rubris ad 2 cm. longis dense pilis rubescentibus longis adpressis
pilosis; calyx ca. 1 cm. longus, sepalis lanceolato-ovatis vel lanceolatis
longe anguste attenuato-acuminatis integris vel subundulatis rubro
tinctis sparse adpresse pilosis; corolla non visa. — Honduras: Climbing
on a tree in forest near the summit of the range above El Achote, in
the cloud zone, above the plains of Siguatepeque, Dept. Comayagua,
1,850 meters, August 1, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 6255 (type in Herb. Field Mus.).

CORYTHOLOMA Dene.

Corytholoma Warscewiczii (Bouche" & Hanst.) Standl. Corolla
orange. Rocky hillside.

ACANTHACEAE
DYSCHORISTE Nees
Dyschoriste oaxacensis Kobuski. In an old, wet meadow.

PLANTAGINACEAE

PLANTAGO L.
Plantago Galeottiana Dene. In a clearing above El Achote.

RUBIACEAE
ANISOMERIS Presl

Anisomeris protracta (Bartl.) Standl. A shrub or small tree up
to 6 meters tall ; flowers white or pale yellow. River bank.

Anisomeris rudis Standl., sp. nov. — Arbor 9-metralis, trunco
12-15 cm. diam., ramis gracilibus subteretibus fusco-ferrugineis,
novellis dense pilis patentibus vel adscendentibus lucidis ochra-
ceis pilosis, internodiis brevibus; stipulae deciduae lineari-subulatae
erectae pilosulae ; folia modica breviter petiolata crasse membranacea,
petiolo crassiusculo 8-10 mm. longo dense paten ti-piloso; lamina
ovato-oblonga vel anguste elliptico-oblonga 7-11.5 cm. longa 2.5-5
cm. lata longe sensim anguste acuminata, basi acuta vel acuminata
et saepe plus minusve decurrens, supra in sicco laete viridis sub-
lucida sparse breviter pilis albidis hirsuta, inter venas dense trans-
verso-lineolata, subtus concolor, densiuscule praesertim ad nervos

FLORA OF HONDURAS 395

patenti-pilosula, ad axillas dense albido-barbata, costa gracili
prominente, nervis lateralibus utroque latere 6-9 obliquis prominen-
tibus gracillimis, nervulis subimpressis; inflorescentiae axillares
solitariae capitatae vel cymulosae graciliter 1.5-2.5 cm. longe pedun-
culatae dense pauciflorae, pedunculo dense patenti-piloso, floribus
sessilibus vel brevissime pedicellatis; hypanthium claviforme vix
ultra 2 mm. longum densissime patenti-pilosulum, calycis lobis
linearibus vel oblongo-linearibus attenuatis extus pilosulis intus
glabris 1.5-3 mm. longis subpatentibus. — Honduras: In forest near
the summit of the ridge above El Achote, in cloud zone, above the
plains of Siguatepeque, 1,800 meters, August 1, 1936, T. G. Yuncker,
R. F. Dawson & H. R. Youse 6253 (type in Herb. Field Mus.).

The most closely related species is Anisomeris protracta (Bartl.)
Standl., a frequent species in Honduras, which differs in having
closely appressed pubescence and very short, blunt calyx lobes.

BORRERIA Meyer

Borreria fruticosa Standl. A shrub about 60 cm. tall; corolla
white. In pine forest near El Achote. Known only from this region.

CEPHAELIS Sw.

Cephaelis axillaris Sw. A shrub about 1 meter tall; flowers
white. In forest near the summit of the range above El Achote.

Cephaelis elata Sw. A small tree 4.5-6 meters tall and 5-8 cm.
in diameter; bracts red. In a deep, wet ravine near El Rincon.
New for the flora of Honduras.

CHIOCOCCA P. Br.

Chiococca alba (L.) Hitchc. A somewhat scandent shrub;
corolla golden yellow. In a thicket near Lake Yojoa.

Chiococca phaenostemon Schlecht. A tree about 4.5 meters
tall and 5-8 cm. in diameter; perianth green. In a wet ravine near
El Achote.

COCCOCYPSELUM P. Br.

Coccocypselum hirsutum Bartl. Flowers and fruit blue.
Creeping in wet soil on a rocky hillside.

CRUSEA Cham. & Schlecht.

Crusea calocephala DC. Flowers purple. In wet soil in pine
forest.

396 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

DECLIEUXIA HBK.

Declieuxia fruticosa (Willd.) Kuntze, var. mexicana (DC.)
Standl. Corolla white. In pine forest near El Achote.

DIODIA L.

Diodia brasiliensis Spreng. var. angulata (Benth.) Standl.
A shrub about 30 cm. tall; flowers white. Between rocks on a river
bank.

GONZALAGUNIA Ruiz & Pavon

Gonzalagunia panamensis (Cav.) Schum. A shrub about 1.5
meters tall; flowers white. Near the shore of Lake Yojoa.

HAMELIA Jacq.

Hamelia patens Jacq. A shrub 3-4 meters tall; flowers orange
to red. River bank.

HOFFMANNIA Sw.

Hoffmannia angustifolia Standl. Climbing in trees at the
bottom of a ravine near El Achote.

Hoffmannia gesnerioides (Oerst.) Kuntze. A low, sprawling
shrub; fruit scarlet, pulpy. On forest floor near El Rincon. A rare
species, new to Honduras, known also from Nicaragua.

Hoffmannia nebulosa Standl., sp. nov. — Frutex metralis
sparse ramosus, ramis gracilibus teretibus viridescentibus vel sub-
ochraceis, internodiis brevibus vel elongatis, novellis sparse vel
densiuscule villosulis; stipulae parvae deciduae; folia modica petio-
lata membranacea, petiolo gracili vulgo 1-3 cm. longo dense breviter
villosulo; lamina obovato-oblonga 10-16 cm. longa 4-6.5 cm. lata
abrupte breviter cuspidato-acuminata, basin versus sensim angus-
tata, basi ipsa acuta, supra in sicco laete viridis sparse vel ad nervos
dense villosula cito glabrata, subtus paullo pallidior, ad costam
nervosque densiuscule pilis longioribus villosa, aliter sparse brevis-
sime villosula vel fere glabra, costa crassiuscula prominente, nervis
lateralibus utroque latere ca. 1 angulo lato saepe fere recto adscen-
dentibus arcuatis; flores in axillis fasciculati pauci, pedicellis fructi-
feris vix ultra 2 mm. longis villosulis; baccae juvenes oblongae 5 mm.
longae paullo ultra 2 mm. latae glabrae basi obtusae; calyx ad apicem
baccae persistens 0.6 mm. longus breviter acute dentatus.- — Hon-
duras: In forest near the summit of the range above El Achote, in
the cloud zone, above the plains of Siguatepeque, Dept. Comayagua,
1,850 meters, August 1, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 6252 (type in Herb. Field Mus.).

FLORA OF HONDURAS 397

"Flowers green," but no corollas, at least, are present on the two
sheets of type material. Like most species of Hoffmannia, this
exhibits no outstanding characters, unless in the sessile, few-flowered
inflorescence. It differs in several respects from all the too numerous
species that have been described from Central America.

PALICOUREA Aubl.

Palicourea Galeottiana Martens. A tree about 6 meters tall
and 7-8 cm. in diameter; branches of the inflorescence golden brown.
Ravine near El Achote.

PSYCHOTRIA L.

Psycho tria crebrinervia Standl. A shrub about 2 meters tall.
Side of a ravine near El Achote.

Psychotria erythrocarpa Schlecht. A shrub about 1.5 meters
tall. In a ravine near El Achote.

Psychotria involucrata Sw. A shrub about 1 meter tall;
flowers greenish; bracts red. In a thicket near the shore of Lake
Yojoa.

Psychotria microdon (DC.) Urban. A shrub, somewhat
scandent; flowers aromatic. In a thicket near the beach west of Tela.

Psychotria patens Sw. A shrub about 2 meters tall. In a
thicket, ravine near El Rincon.

Psychotria trichotoma Mart. & Gal. A shrub about 1.5
meters tall. Bottom of a ravine near El Achote.

Psychotria uliginosa Sw. A shrub about 1 meter tall; corolla
pinkish; fruit scarlet. In a deep, wet ravine near El Rincon.

Psychotria Yunckeri Standl., sp. nov. — Subgen. Mapouria.
Arbuscula 3-4.5 m. alta, trunco 2.5-5 cm. diam., omnino glabra,
ramis -crassis subteretibus fusco-ferrugineis interdum striatis, inter-
nodiis brevibus; stipulae ut videtur primo connatae et calyptri-
formes caducae ferrugineae oblongo-lineares ad 3.3 cm. longae 3-3.5
mm. latae attenuato-acutae; folia majuscula longiuscule petiolata
firme membranacea, petiolo crassiusculo 1.5-3 cm. longo; lamina
elliptica vel oblongo-elliptica 10-20 cm. longa 4.5-7.5 cm. lata acuta
vel acuminata, basin versus attenuata et interdum subabrupte
acuminata atque decurrens, supra in sicco brunneo-fuscescens, costa
nervisque conspicuis sed vix prominulis, subtus fere concolor, costa
gracili elevata, nervis lateralibus utroque latere ca. 15 angulo latius-

398 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

culo adscendentibus parum curvis prope marginem arcuato-con-
junctis prominulis, venulis obsoletis; inflorescentia terminalis erecta
cymoso-paniculata dense vel sublaxe multiflora crasse 1-2 cm. longe
pedunculata vel interdum sessilis 4-6.5 cm. longa et saepe aequilata,
ramis basalibus 3 vel numerosis adscendentibus crassis, bracteis
parvis caducis, floribus plerumque umbellatis breviter crasse pedi-
cellatis vel sessilibus, pedicellis vix 2 mm. longis; hypanthium obo-
voideum vix ad 1.5 mm. longum, calyce ca. 1 mm. alto 2-2.5 mm.
lato truncate vel obscure remote brevissime denticulate; corolla
viridi-alba 5 mm. longa extus glabra fauce albo-barbata, in alabastro
apice late obtusa, tubo crasso cylindraceo, lobis ovato-oblongis
obtusis aequilongis vel paullo brevioribus; pedicelli fructiferi usque
6 mm. longi; fructus globoso-obovoideus 7 mm. longus, basi obtusus,
apice late rotundatus, 5 mm. diam., pyrenis 2 dorso grosse obtuse
6-costatis, facie commisurali anguste leviter canaliculatus. — Hon-
duras: In forest near the summit of the range above El Achote, in
cloud zone, above the plains of Siguatepeque, Dept. Comayagua,
1,850 meters, July 21, 1936, T. G. Yuncker, R. F. Dawson & H. R.
Youse 6013 (type in Herb. Field Mus.); also No. 6143 from the same
locality. In ravine near El Achote, 1,350 meters, No. 6118.

RANDIA L.

Randia aculeata L. A shrub about 2.5 meters tall; flowers
white. Near El Achote.

RONDELETIA L.

Rondeletia amoena (Planch.) Hemsl. A tree about 4.5 meters
tall. Side of a ravine near El Achote.

Rondeletia buddleoides Benth. A tree about 4.5 meters tall
and 10 cm. in diameter. In a dry ravine east of Siguatepeque.

Rondeletia nebulosa Standl. A small tree about 4.5 meters
tall and 8-10 cm. in diameter. In forest near the summit of the
range above El Achote. The species is confined, so far as known, to
Honduras.

Rondeletia strigosa (Benth.) Hemsl. A shrub. Edge of a
ravine near El Achote.

CAPRIFOLIACEAE

VIBURNUM L.

Viburnum discolor Benth. A tree about 7.5 meters tall and
15 cm. in diameter; flowers white. Side of a ravine near El Achote.

FLORA OF HONDURAS 399

VALERIANACEAE

VALERIANA L.

Valeriana scorpioides DC. An herb; flowers white. Rocky
hillside.

CUCURBITACEAE
LAGENARIA Ser.

Lagenaria siceraria (Mol.) Standl. Flowers white. Climbing
in a thicket on river bank about 6 km. north of Siguatepeque.
Escaped from cultivation?

LOBELIACEAE
LOBELIA L.

Lobelia splendens Willd. In a subalpine bog near El Achote.
Corolla scarlet.

COMPOSITAE
ACHYROCLINE Less.

Achyrocline Yunckeri Blake, sp. nov. — PI. XVI. Annua
paullum erecto-ramosa ca. 6 dm. alta; caulis canescenti-tomentosus;
folia ovata v. elliptico-ovata acuta basi cuneata petiolata non
decurrentia integra triplinervia supra cinerascentia v. aetate viri-
descentia tenuiter tomentosa subtus canescenti-tomentosa; capitula
5-flora sessilia glomerato-paniculata; fl. fem. 4, hermaph. 1; involucri
scariosi margaritacei phyllaria late ovalia v. oblonga obtusa saepius
apiculata.

Erect, leafy annual; stem subterete, striate, pithy, 3 mm. thick
below; leaves alternate, the internodes 0.6-2.5 cm. long; petioles
slender, essentially unmargined, canescent-tomentose, 3-6 mm. long;
blades ovate, elliptic-ovate, or sometimes elliptic, 3.8-6.5 cm. long,
1.8-2.4 cm. wide, acutish or the lower ones sometimes obtuse, dark-
callous-apiculate, above usually grayish or in age greenish, rather
thinly tomentose and beneath the tomentum puberulous with short,
thick, several-celled hairs (from which the long hairs of the tomentum
arise), not glandular, beneath densely and persistently canescent-
tomentose, papery, pliable, erectish to spreading or in age marcescent
and reflexed, the upper gradually reduced to lanceolate or elliptic
bracts 1.8-3 cm. long; panicle terminal, flat, with about 3 principal
branches, many-headed, 7 cm. wide, pubescent like the stem, the
bracts subulate, 5 mm. long; heads sessile, glomerulate, oblong-ovoid
(moistened), about 4.2 mm. long, 2 mm. thick; involucre about 4-
seriate, strongly graduate, 4.2 mm. high, scarious, pearly white,

400 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

the phyllaries about 9-11, the outermost broadly oval, 2.8-3 mm.
long, 1.5-1.8 mm. wide, obtuse or rounded, sometimes apiculate,
the others oblong, obtuse to sub truncate, abruptly apiculate, all with
greenish, stipitate-glandular, and more or less tomentose midline
below; pistillate corollas nearly equaling the style, whitish, filiform,
3 mm. long, minutely 5-toothed, puberulent at the apex with clavate,
subglandular hairs; disk corolla apparently brownish at the apex,
3.5 mm. long (tube 1 mm., throat slender, 2 mm., teeth triangular,
acute, puberulous with clavellate hairs, 0.5 mm. long); achenes
oblong, light-colored, glabrous, 0.5 mm. long; pappus deciduous, of
barely serrulate, white bristles, 3 mm. long.

HONDURAS: On a wet cliff near El Achote, hills above the plains
of Siguatapeque, Dept. Comayagua, alt. 1,350 meters, July 15, 1936,
T.G. Yuncker, R.F. Dawson & H. R. Youse5872 (type No. 1,637,728,
U. S. Nat. Herb.).

The closest relative of this species, Achyrocline deflexa Robins.
& Greenm., of Oaxaca, has longer and relatively narrower, lanceolate
or oblanceolate leaves with the petiole less abruptly demarked from
the blade and margined or winged to the base, and the phyllaries, vary-
ing from obtuse to acute, are indistinctly or not at all apiculate.

AGERATUM L.
Ageratum conyzoides L. A common weed in waste places.

ASTER L.
Aster bullatus Klatt. Rays white. River bank.

BACCHARIS L.

Baccharis glutinosa Pers. A small tree about 6 meters tall
and 7-8 cm. in diameter. On a river sandbar.

Baccharis trinervis (Lam.) Pers. A shrub about 2 meters tall.
Thicket on plains.

BIDENS L.
Bidens pilosa L. var. radiata Sch. Bip. A very common weed.

CHAPTALIA Vent.

Chaptalia nutans (L.) Polak. Rays purplish or white. In
wet, rocky soil in pine forest.

COSMOS Cav.
Cosmos crithmifolius HBK. Rays pink. River bank.

FLORA OF HONDURAS 401

ERIGERON L.

Erigeron jamaicensis L. In a moist pasture.
Erigeron spathulatus Vahl. Rays white. River bank.

EUPATORIUM L.

Eupatorium ivaefolium L. Corolla lavender. In an old
pasture.

Eupatorium ligustrinum DC. A shrub about 2.5 meters tall.
Edge of a ravine near El Achote.

Eupatorium macrocephalum Less. In an old pasture.

Eupatorium morifolium Mill. A tree about 6 meters tall and
10 cm. in diameter. In a wet ravine near El Achote.

GALEANA Llave
Galeana pratensis (HBK.) Rydb. A common weed.

HIERACIUM L.

Hieracium hondurense Blake. Rays yellow. In wet soil,
in pine forest.

MELAMPODIUM L.
Melampodium brachyglossum Donn. Sm. A common weed.

MIKANIA Willd.

Mikania Tonduzii Robinson. Sprawling over shrubs in a wet
ravine near El Achote.

ORTHOPAPPUS Gleason

Orthopappus angustifolius (Sw.) Gleason. Corollas pale
lavender. In an old pasture.

PEGTIS L.
Pectis sp. Rays yellow. In short grass of an old pasture.

PEREZIA Lag.
Perezia nudicaulis Gray. Flowers white. On a rocky hillside.

POLYMNIA L.

Polymnia maculata Cav. Stems 2.5-3.5 meters tall; leaves up
to 45 cm. long; corollas yellow. About 6 km. north of Siguatepeque.

Polymnia oaxacana Sch. Bip. Stems about 1.2 meters tall;
rays orange-yellow. River bank.

402 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

SENECIO L.

Senecio arborescens Steetz. A small tree about 4.5 meters tall
and 8 cm. in diameter; leaves fleshy. Side of a ravine near El Achote.

SONCHUS L.

Sonchus oleraceus L. Side of a trail about 6 km. west of
Siguatepeque.

TAGETES L.

Tagetes filifolia Lag. Rays white, shading to yellow in the
throat. In pine forest near El Achote.

Tagetes Schiedeana Less. Rays orange; very aromatic. In
wet soil, border of a banana plantation.

TRIDAX L.

Tridax procumbens L. Rays buff. In wet soil near Potre-
rillos; also on a rocky hillside near Siguatepeque.

VERBESINA L.

Verbesina vicina Blake, sp. nov. — PI. XVII. Arbor parva, ramis
strigillosis glabratis; folia alterna majuscula obovata breviter acumi-
nata basi longe cuneata petiolata crenato-mucronulata pergamentacea
utrinque viridia supra asperule hirsutula pilis basi ima lepidotis, sub-
tus non dense hirsutula; capitula minuscula aurea radiata cymoso-
paniculata vix numerosa; radii ca. 5 mm. longi; achaenia latiuscule
alata.

"Small tree about 4.5 meters tall and 5-7.5 cm. in diameter";
branches usually densely strigillose or subappressed-hirsutulous, in
age glabrescent or glabrate, subterete, wrinkled-striatulate, wingless,
the younger greenish, the older gray-barked, 3-4 mm. thick; petioles
rather slender, flattish above, rounded beneath, strigillose or erectish-
hispidulous especially above, 1-1.8 cm. long, not auriculate or de-
current; blades obovate, (7-) 11.5-23 cm. long, (3-) 4-7.5 cm. wide,
usually rather abruptly short-pointed with an acute, callous-tipped
apex, rarely obtuse, crenate-mucronulate above the entire, cuneate
base (teeth 8-12 pairs, 5-20 mm. apart, the mucro callous, about 0.4
mm. high), above deep green, somewhat shining, roughish, evenly
but scarcely densely hirsutulous with subappressed or antrorse hairs
with lepidote but scarcely or not tuberculate base, densely hirsutu-
lous on the costa, in age glabrescent and inconspicuously lepidote-
maculate, beneath scarcely lighter green, evenly but not densely

FLORA OF HONDURAS 403

hirsutulous or substrigillose with inconspicuously lepidote-based
hairs, feather- veined, the costa prominent beneath, the chief lateral
veins 8-10 pairs, prominulous beneath, curved-ascending; cymes or
cymose panicles axillary, subterminal, much surpassed by the leaves,
about 3-10-headed, 1.5-3.5 cm. wide, convex, densely erect- or
ascending-pilosulous, the peduncle about 1.5 cm. long, the pedicels
slender, 5-17 mm. long, the bracts minute; heads about 1 cm. wide;
disk campanulate, 8-10 mm. high, 5-8 mm. thick (as pressed);
involucre 3-4-seriate, 4.5-6 mm. high, appressed, the phyllaries
rather few, very unequal, graduate, the outermost minute, ovate,
the others oblong or ovate-oblong, 0.8-1.2 mm. wide, with pale,
somewhat indurate, about 2-ribbed base and longer or shorter,,
greenish tip, all obtuse, loosely and not densely pilosulous and short-
ciliate; rays 4, yellow, pistillate, the tube spreading-pilosulous, 1.8
mm. long, the lamina oval, 3-dentate, 9-nerved, 5.5 mm. long, 3.3
mm. wide; disk flowers not very numerous, their corollas yellow,
hirsutulous on the tube and base of throat, otherwise glabrous, 4.7-
5.2 mm. long (tube 0.8-1 mm., throat cylindric, 3.5-3.7 mm., teeth
4 or 5, ovate, papillose-margined, obtuse, 0.5-0.7 mm. long); pales
rather broad, several-vittate, sparsely pilosulous, thinner and green-
ish toward the obtuse, erect or very slightly spreading tip, about 5
mm. long; disk achenes cuneate-obovate, 4.8 mm. long, 3.2 mm. wide
(including the wings), the body narrowly obovate, blackish, sparsely
hirsute above with partly tuberculate-based hairs, 4.5 mm. long, 1.8
mm. wide, the thin, whitish wings (probably not fully mature)
dentate-ciliate, 0.6-0.8 mm. wide; awns 2, unequal, linear-lanceolate,
acuminate, sparsely hirsutulous-ciliolate, somewhat divergent, 2.2-
3.2 mm. long, adnate dorsally to the wings at the base.

HONDURAS: In a ravine, bank of a stream near El Achote, hills
above the plains of Siguatepeque, Dept. Comayagua, alt. 1,350
meters, July 17, 1936, T. G. Yuncker, R. F. Dawson & H. R. Youse
5907 (type No. 1,637,745, U. S. Nat. Herb.).

In range and characters this species is intermediate between V.
pleistocephala (Bonn. Sm.) Robinson, of Guatemala, and V. Oer-
stediana Benth., of Costa Rica. The inflorescence of the specimens
examined (including those retained by Dr. Yuncker) is probably
dwarfed and not representative of the species in its full development.
In V. pleistocephala the leaves are only about half the size of those
of V. vicina, usually sharply serrate or serrulate, and extremely
rough above with strongly tuberculate-based but scarcely lepidote
hairs. In V. Oerstediana the leaves are smooth to the touch above

404 FIELD MUSEUM OF NATURAL HISTORY — BOTANY, VOL. XVII

and there usually glabrous (except along the puberulent costa), or
sometimes sparsely pilosulous.

Verbesina Oerstediana Benth. (in Oerst. Vidensk. Medd. 1852: 96.
1852) was recorded as one of the most characteristic plants on the
southern slopes of Mount Irazu, Costa Rica, at 8,000-9,500 feet
altitude. The type at Kew is labeled as from 9,000 feet; a photo-
graph and fragments from this sheet are in the U. S. National Her-
barium, which also possesses a sheet collected by Oersted bearing
essentially the same data and presumably belonging to the type
collection. Bentham described the branches as well as the panicle
as villous; Robinson & Greenman (Proc. Amer. Acad. 34: 558. 1899),
who had examined only Pittier 11583, from Irazu (the altitude given
on the specimen of this number in the U. S. National Herbarium is
2,900-3,000 meters; Robinson and Greenman give the altitude as
2,000-2,500 meters [about 6,500-8,200 feet], which may be due to
an error in labeling) called them lanate. In the specimen of Pittier
11583 in the U. S. National Herbarium the branches and petioles are
villous-lanate, and the panicle correspondingly pubescent; the
pubescence on the Oersted specimen, although much weathered, has
been essentially the same. The leaves were described by Bentham
in his diagnosis as pilosulous above, loosely villous beneath, in his
fuller description as scabriusculous above, at length nearly glabrate.
The leaves are essentially the same in all three specimens above
cited — at first evenly but not densely pilosulous above with hairs
that are not at all or only very slightly tuberculate-thickened at the
base, and smooth to the touch, in age glabrate or glabrescent above.

Eight other collections from Costa Rica that have been referred
to V. Oerstediana are in the U. S. National Herbarium, all collected
in the provinces of San Jos4 and Cartago by Paul C. Standley, and
all, with a single exception, from altitudes of 1,500-1,900 meters.
In all these plants the pubescence of the branches and panicle is much
less conspicuous, shorter, and usually sparse, varying from strigillose
to spreading-pilosulous. They appear to represent a distinct variety
of lower altitudes, which may be called

Verbesina Oerstediana var. glabrior Blake, var. nov. — Rami
novelli et paniculae sparse v. dense strigillosa v. strigosa, vel inter-
dum pilosula pilis adscendentibus v. subpatentibus; folia supra
glabra v. subglabra, subtus sparse v. sparsissime pubescentia.

COSTA RICA: Province of San Jose": Moist woods, Cerro de Piedra
Blanca, above Escasu, Jan., 1924, Standley 32^89; moist forest,

FLORA OF HONDURAS 405

between Aserri and Tarbaca, alt. 1,600-1,900 meters, Feb., 1924,
Standley 34142; open slope, Cerro de Las Vueltas, alt. 2,700-3,000
meters, Dec., 1925-Jan., 1926, Standley & Valeria 43504; oak forest,
vicinity of Santa Maria de Dota, alt. 1,500-1,800 meters, Dec.,
1925, Standley 41619 (type No. 1,252,372, U. S. Nat. Herb.); Dec.,
1925-Jan., 1926, Standley & Valeria 43448.— Province of Cartago:
Wet forest, Cerro de La Carpintera, alt. 1,500-1,850 meters, Feb.,
1924, Standley 35715, 35728; wet forest, La Estrella, March, 1924,
Slandley 39429.

VERNONIA Schreb.

Vernonia melanocarpa (Gleason) Blake. A shrub about 3
meters tall. In a thicket on river bank.

WEDELIA Jacq.

Wedelia parviceps Blake. Stems more or less woody at the
base; rays yellow. In a thicket on river bank. .

ZEXMENIA Llave

Zexmenia frutescens (Mill.) Blake, form. A shrub 1.5-2
meters tall. In a thicket on the plains.

Zexmenia perymenioides Blake, sp. nov. — PI. XVIII. Arbor
parva; rami tenues hispiduli et strigillosi etiam patenti-hispidi ; folia
elliptico-lanceolata acuminata basi acute cuneata petiolata serrulata
triplinervia supra aspere hispida et hispidula, pilis basi tuberculatis,
subtus pallidiora hirsuta et hirsutula; capitula 3 terminalia pedun-
culata foliis breviora radiata aurantiaca, pedunculis strigillosis;
involucri 3-seriati paullum gradati phyllaria exteriora oblonga
strigosa apice herbaceo ovato acuto squarroso, interiora angustiora
scariosa; achaenia immarginata tuberculato-hispidula; pappi arista
1 tenuis achenio paullo brevior, squamellae ca. 2-3 saepius ovatae ca.
1 mm. longae.

"A small tree about 15-20 feet (4.5-6 meters) tall and 3 inches
(7.5 cm.) in diameter"; branch hexagonal, 2 mm. thick, sparsely
branched, greenish, rather densely hispidulous with spreading or
reflexed hairs and in places strigillose, also rather densely hispid with
wide-spreading, slightly tuberculate-based hairs about 1 mm. long,
the pubescence especially dense at the nodes; petioles slender,
essentially naked, pubescent like the branch, 3-9 mm. long; blades
6-8.5 cm. long, 1.8-2.7 cm. wide, rather shortly falcate-acuminate
with an acute, callous-tipped apex, serrulate above the subentire,
cuneate base, firm-papery, above deep green, not shining, evenly

406 FIELD MUSEUM OF NATURAL HISTORY— BOTANY, VOL. XVII

but scarcely densely hispid and hispidulous with tuberculate-based,
antrorse hairs, beneath lighter green, on the veins densely and rather
softly hirsute with spreading to subappressed hairs, on the surface
evenly hirsutulous with spreading to subappressed hairs, triplinerved
3-10 mm. above the base, in age with the smaller veins impressed
above, prominulous-reticulate beneath; peduncles 3.5 cm. long or
less, very slender, densely strigillose, 1 -headed; heads 2.5 cm. wide
(as pressed); disk campanulate, about 12 mm. high, 8 mm. thick
(as pressed); involucre campanulate, impressed at the base, 3-4-
seriate, slightly graduated, 10 mm. high, the phyllaries rather few,
those of the 2 outer series together 6-8, oblong or oblong-obovate,
1.5-2.8 mm. wide, with indurate (in the inner greenish-centered,
scarious-margined) base and shorter, ovate or lance-ovate, usually
acute or acuminate, rarely obtusish, squarrose, herbaceous tip,
densely strigillose on their exposed surface below, more sparsely so
and indistinctly ciliate on the herbaceous tip, those of the inner
series linear-oblong, obtuse, scarious, erose and usually ustulate at
the tip, appressed, strigillose along the midline; rays 8, fertile, the
tube very sparsely subglandular-puberulous above, 3.3 mm. long,
the lamina oval, tridenticulate, 12-14 mm. long, about 8 mm. wide,
short-hirsute on the back on the 2 chief nerves and finely hispidulous
at the apex, about 17-nerved; disk corollas golden yellow, glabrous
except for the sparsely hispidulous, papillose-margined teeth, 8.7
mm. (tube 3 mm., throat slender-funnelform, 4.5 mm., teeth ovate,
1.2 mm. long) ; pales narrow, scarious, acuminate, often with a lateral
tooth on each side below the apex, ciliolate toward the tip, sparsely
hispidulous on the midline above, about 11 mm. long; ray achenes
narrowly cuneate, 5.5 mm. long, 1.7 mm. wide, trigonous, keeled on
the inner face, somewhat rounded and 1-3-nerved on the outer,
wingless and marginless, blackish brown, evenly but not densely
tuberculate-hispidulous, not obviously contracted at the apex, their
pappus of 3-4 lanceolate (rarely linear) to ovate or obovate, mostly
acuminate, hispidulous-ciliolate squamellae or paleae, 0.8-1.5 mm.
long, that on the inner angle sometimes aristiform and 3.5 mm. long;
disk achenes narrowly cuneate, 5-5.3 mm. long, 1.2-1.5 mm. wide,
compressed-quadrangular, normally with 1 nerve between each pair
of angles, marginless and wingless, slightly rounded at the apex but
not at all necked, blackish brown, sparsely tuberculate-hispidulous,
their pappus rather persistent, of 1 narrowly lanceolate, long-
acuminate, hispidulous awn about 4.5 mm. long (on the inner angle),

FLORA OF HONDURAS 407

and 2-3 slender-subulate to ovate, acute, hispidulous-ciliolate, free
squamellae 0.5-1.3 mm. long, or the latter lacking.

HONDURAS: In a wet ravine near El Achote, hills above the plains
of Siguatepeque, Dept. Comayagua, alt. 1,350 meters, July 26,
1936, T. G. Yuncker, R. F. Dawson & H. R. Youse 6099 (type No.
1,637,729, U. S. Nat. Herb.).

This species is strikingly suggestive of Perymenium in all features
except the achene and pappus. Its closest ally is clearly Z. aurantiaca
Klatt, of Costa Rica. In that species, of which two sheets of the
type collection (Tonduz 9836 and its equivalent, J. D. Smith 7071)
are before me, the stem and branches are closely strigillose without
any spreading hairs, the leaves are ovate and much broader (only
twice as wide as long), and the heads are somewhat thicker and on
longer peduncles. In essential characters of achene and pappus the
two species are very similar, although no achene examined has the
pappus squamellae as large as on some of the achenes of Z. pery-
menioides. W. W. Jones (Proc. Amer. Acad. 41: 161. 1905 ["A
revision of the genus Zexmenia," Contr. Gray Herb. 30: 161])
thought that Z. aurantiaca might not be a Zexmenia, and suggested
that the pappus somewhat resembled that of Perymenium. The
species, however, is certainly a Zexmenia, although it was probably
improperly placed in the subgenus Auchenocarpa by Jones. His
description of the pappus as of "very fragile plumose awns, generally
only one as long as the body, and several unequal bristle-like squa-
mellae," although based on Klatt's type, gives a very misleading
notion of the pappus actually present in this species. The ray
achenes are 3-angled, those of the disk 4-angled (this feature reversed
in his description), tuberculate-hispidulous, and very closely similar
to those of Z. perymenioides above described. The pappus in the ray
achene consists of a very slender, fragile, hispidulous awn 5 mm. long,
slightly dilated at the base, on the inner angle, a similar but much
shorter awn (1.8 mm. long) on each of the outer angles, and about 5-6
lanceolate, hispidulous squamellae 0.5-1 mm. long, the whole
slightly connate at base. The disk achene bears one very slender,
fragile, hispidulous awn 5 mm. long, sometimes paleaceous-dilated
at the base (probably representing connation with 2 squamellae),
and about 6-12 mostly lanceolate, hispid-ciliate squamellae, the
latter shortly but distinctly connate at the base.

THE LIBRARY OF THE

FE3191938

UNIVERSITY OF ILLINOIS

Field Museum of Natural History

Botany, Vol. XVII, Plate II

a
VIEW OF SIGUATEPEQUE, HONDURAS

HAND-SAWING PINE LOGS INTO BOARDS FOR USE IN LOCAL CONSTRUCTION

Field Museum of Natural History

Botany, Vol. XVII, Plate III

LYCOPODIUM ARCANUM MAXON
Type specimen; natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate IV

PLANTS OF THE REPUBLIC OF HONDURAS. C. A.

ANTHURIUM BASIATTENUATUM STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate V

A

L ANTS OF THE REPUBLIC OF HONDURAS, C. A.

ANTHURIUM LUCENS STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate VI

r

PLANTS OF THE REPUBLIC OF HONDURAS. C. A.

n ini. peduncle lie tit

.

ANTHURIUM YUNCKERI STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate VII

THECOPHYLLUM MONTANUM L. B. SMITH
Type specimen; one-third natural size; details of flower natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate VIII

PLANTS OF THE REPUBLIC OF HONDURAS, C. A.

•

Epiphyte. Bracts a flesh pint:. .
green, t, lr?a recurved. In r«vlne n

TILLANDSIA GLOSSOPHYLLA L. B. SMITH
Type specimen; one-third natural size; details of flower natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate IX

TILLANDSIA LAMPROPODA L. B. SMITH
Type specimen; one-third natural size; details of flower natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate X

PLANTS OF THE REPUBLIC OF HONDURAS. C. A

TILLANDSIA YUNCKERI L. B. SMITH
Type specimen; one-third natural size; details of flower natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XI

PIPER FAVICULIFERUM TRELEASE
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XII

•
.

.

CALLIANDRA ARBOREA STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XIII

H57K73

PLANTS OF THE REPUBLIC OF HONDURAS, C. A

WjL.'

N° =rm Da* J ' ~~

' , , T/ll/36

^U/^>W^!Ct•"-n^.--tu-/«*^^Lm ^P^,^ '

•--• W~nl.h-y.no,. On r

DESMODIUM LUTEOLUM STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII. Plate XIV

PLANTS OF THE REPUBLIC OF HONDURAS. C. A.

ILEX HONDURENSIS STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XV

BEGONIA YUNCKERI STANDLEY
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XVI

'

fjW^
^ *>

ACHYROCLINE YUNCKERI BLAKE
Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XVII

PLANTS OF THE REPUBLIC OF HONDURAS. C.

n

|
e"T1N«'»* ', . .. ~J~ ' »

LABOBATOl..
.««., , APPARATUS

HiTilililili

•'•aa, bank of ••-

.
i"*."b'; ' "rd *•' ln'

VERBESIKA VICINA BLAKE

Type specimen; one-third natural size

Field Museum of Natural History

Botany, Vol. XVII, Plate XVIII

ZEXMENIA PERYMENIOIDES BLAKE
Type specimen; one-third natural size

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UNIVERSITY OF ILLINOIS URBANA