(^i-iojf' FROM THE LIBRARY OF Dr. CHARLES VOSE BEMIS, MEDFORD, MASS. THE GIFT OF Dr. NORMAN FITCH CHANDLER, TO THE BOSTON MEDICAL LIBRARY, 1907. Digitized by the Internet Arcinive in 2011 witii funding from Open Knowledge Commons and Harvard Medical School http://www.archive.org/details/courseofmedicals01burd COURSE OF MEDICAL STUDIES: CONTAINING A COMPARATIVE VIEW OF THE ANATOMICAL STRUCTURE OF MAN AND OF ANIMALS ; A HISTORY OF DISEASES ; AND AJJ ACCOUNT OF THE KNOWLEDGE HITHERTO ACaiTISED WiTSJ REGARD TO THE REGULAR ACTION OF THE DIFFERENT ORGANS. A WORK CHIEFLY DESIGNED FOR TEE USE OF MEDICAL STUDENTS. BY J. BURDIN, M.D. TRANSLATED FROM THE FRENCH. IN THREE VOLUMES, VOL. I. LONDON: PRINTED rOE CUTHELJ, AND MARTIN, MIDDLE-ROW* HOLBORKJ BY H. WilXS, CHANCERY.LWfE, 1803. ^ ' ■?*, ? \ 5 J\J^ PRELIMINARY DISCOURSE, In this work I have endeavoured to exhibit a general view of the knowledge acquired in regard-to the organi- zation of man. My principal obje£l In undertaking this labour was to unite different branches of fcience, defigned to illuftrate each other, in fuch a manner as to enable every intel- ligent perfon to become acquainted with fubje6ls, which, confidering their importance, ought to be mojre gene- rally cultivated. It is indeed aftonifhing, at a period when attempts are made to comprehend in public in- ftruiStion all the ufeful fcie^ces, that we do not find among them one which gives to man a knowledge of his organic fl:ru£\ure ; which makes known to him the phsenomena exhibited by the regular adtion of his dif- ferent parts ; and which points out to him the general caufes and natural progrefs of his numerous derange- ments. A methodical ftudy of the phainomena of the organi- zation is alone capable of enabling man to avoid what may be hurtful to him, and to acquire thofe habits which are moft conducive to his prefervation. For want of this knowledge fome abandon themfelves with fecu- rity to irregularities, the dangerous confequences of which they cannot fprefee ; others, with a view to ame- liorate or to preferve their health, fubje^l themfelves to an auftere regimen ; deprive themfelves of thofe varied and agreeable enjoyments which are necelTary to com- YOL. I. a plete 11 PRELIMINARY DISCOURSE. plete life ; and at length, by rniftaken care, weaken that good conlthution which they are defirous to main- tain. Thofe who remain ignorant in regard to the nature of their orp:;ani£ation expofe themfelves alfo, when it experiences any derangements^ to the mercy of every thing ai : un^ them; and it is not uncommon to find perf ns of great talents, who in fuch cafes believe in the greatefr abfurdines, and thus give importance to the rncit d2.,geroiis errors. To this may be adced, that the only means of banifh- ing quackery, againft which mankind declaim fo much, is to propagate the knowledge acquired in regard to the phyfical nature of man. Thofe, indeed, who have cor- real: ideas of the phsenomena of their organization will not readily become dupes to advice dictated by ignorant zeal or by forward felfifhnefs ; whereas the rich and the ill-informed are always more expofed to deception, as the honeft and enlightened phyfician has too little influence to be able to convince them of their danger, and to pro- tect them againft the ignorance and effrontery of thofe who endeavour to miflead them. If the ftudy of the organization, notwithftanding fo many motives of intereft, is not fo generally cultivated as it ought to be, and if it has not yet been made a part of the ufual courfe of education, this deficiency mud be afcribed to the detached manner in which the different branches it embraces have been treated ; as they have not yet been united into a proper fyftem, worthy of being claffed among works- on the exacl fciences. But other caufes alfo have contributed to retard the ' progrefs of this ufeful fcience. It may be obferved, in the firft place, that a ftudy which is fo difficult, and which, PRELIMINARY DISCOURSE. Ill which, befides great talents, requires a mind entirely divefted of prejudice, could not make a confpicuous figure among the objects of human knowledge, till at a very advanced period of its improvement. It may be re- marked, alfo, that few cultivate this fcience merely with a view of fearching for new dlfcoverles ; and this, per- haps, is owing to the difgufi: which mufl be overcome before people can acquire fufficlent refolution to extend the field of their obfervation to the fick and the dying. It Is well known, alfo, that young men, who devote themfelves to the ftudy of medicine, are in general obliged to apply too foon to pradlice ; which does not allow them fufficient leifure for difficult refearches and profound meditations. In the laft place, thofe who have divided among them- felves the different branches which relate to the phyfical nature of man, do not fufficiently comm.unlcate to each other the refult of their experience and of their obfer- vatlons. A phyfician who begins pra6lice with a certain degree, of knowledge, is always (truck with the confufion exhi- bited to him by the fcience of medicine. At firfl:, he afcribes it to the difficulty of the fludy; but if he con- tinues to be a calm obferver, and to believe nothing implicitly, he foOn perceives that thofe who cultivate the different branches of which it is compofed have fo little intercourfe, that, though very ex.tenfive knowledge has been obtained, the refult of their individual labours is, for the moft part, loft to the fcience in general ; and that a great number even, without knowing it, are divided in their opinions in regard to the p: Incipal points on which it is founded. Hence it happens, that the difllnifl; fludy of the different parts of the hiftory of the a 2 organi- IV PRELIMINARY DISCOURSE. organization, inftead of promoting, ferves rather to re- tard its progrefsj becaufe thofc by whom it is cuhivated do not fet out from common and fundamental princi- ples, to which all their particular obfervations might be referred. When a phyfician, therefore, begins to praflife, he is neceflarily flruck with the . number of popular errors which he every where finds among perfons who are always defirous to explain, in their own manner, the different ph«nomena of health and difeafe ; which indi- cates, at leaft, how anxious they are to obtain informa- tion refpe£ling every thing that relates to the organiza- tion. Unfortunately, moft phyficians feem, at firft, to participate in thefe falfe notions, that they may not too openly attack received opinions : by thefe means they gradually become familiarized with them ; and at length adopt them entirely as inconteftable truths. Thus, men who in confequence of their fituation ought to contribute moft towards the difFufion of ufeful knowledge, only give importance to fatal errors, which being eftablifhed by time and the force of habit, muft be difficult to be era- dicated, and can be deftroyed only by their united eftorts. Before I enter into any detail in regard to the plan of the work, the motives of which I have here explained, it will be proper to make a few general obfervations on organized bodies. Organized bodies have been diyid.ed into vegetables and animals ; but they are far from being all naturally fufceptible of arrangement under thefe two divifions. There is a very numerous ferles of beings evidently or- ganized, with the intimate ftruAure of which we are ab- folutely unacquainted, either on account of their ex- treme- PRELIMINARY DISCOURSB, Y treme mlnutenefs, or becaufe their tranfparency deprives us of the means of obferving their organs, by making them appear to us under the form of a homogeneous vifcofity. When thefe beings develop themfelves without changing their place, or by exhibiting a fort of ramifi- cation, they are clafled among vegetables : of this kind are the byfli, lichens, fuci, ulvjc, confervse, and- tre- mellas. On the other hand, when they move, or feem to have a diftincl alimentary canal, like the volvox, mo- nades, fponges, lithophites, alcyons, &c., they form a part of the clafs of animals. There are alfo feveral fpecies which are placed alternately in thefe two clafles, according to the ' different obfervations which have been made. It appears, that it is among thefe beings, fo little known, that the divifion between animals and vege- tables takes its origin, or where the two clafies are confounded. The numerous clafs of animals exhibit a natural divi- fion by the exiftence or non-exiftence of the vertebral column. The difference between vertebral and in-ver- tebral animals is fo great, that Daubenton was of opi- nion that the former ought not to retain the name of animals, if it be given to the latter. Vertebral aairnals comprehend fifties, reptiles, birds, and the order of the mammalia, at the head of which man is placed. The knowledge acquired in regard to the organiza- tipn of man, which I have endeavoured to unite in this work, comprehends : — ift. Anatomical refearches ; 2d. A hiftory of difeafes ; 3d. An examination of the vital a 3 fun£lion$. Vi PRELIMINARY DISCOURSE. funftions. Thefe three branches of knowledge, which compofe the, philofophy of man, naturally divide the Work into three parts. Firjl Tart. The anatomical defcription of man, as well as that of a great number of animals, which forms the fubje£l of the firft part, has been given with great minutenefs, and fcrupulous exa£lnefs, by celebrated ana- tomifts : but as a complete and connected view of the different fyftems of organs in man, compared with the fame fyftems of organs in other animals, is not to be found in any work, the refuk was, that only one part of the pi£lui*e was exhibited j and therefore it was im- poffible to form a proper idea of the whole. The lec- tures of Cuvier, from which I have endeavoured to de- rive as much advantage as poffible, and which were col- lected and in part publifhed by Dumeril, will form in this new line a complete work, the conclufion of which is looked for with Impatience. The firft difHculty which occurs, in defcriblng the fyftems of organs, arifes from the choice of the order proper to be adopted. As all the fyftems of organs difcharge their • fun6lions almoft fimultaneoufly, a fuc- ceflive defcription of them is always attended with this difadvantage, that it feparates cbjeCIs which ought to remain united. It was therefore neceffliry that the order adopted ihould, as far as poffible, remedy this inconve- nience. The whole of the parts of the body is compofed of fix principal fyftems: I ft. The fyftem of the bones and mufples. 2d, The cerebral and nervous fyftem. - 3d. The fyftem of the fenfes. 4th, PRELIMINARY DISCOURSE. Vir 4th. The fyftem of digeftion. 5th. The fyftem of circulation and refpiratlon. . 6th. The fyftem of generation. The defcription of each of thefe fyftems, in man, is preceded by= a curfory view of the correfponding or- gans in the different clailes of animals. In the fyfleni of the bones :will be feen the variety of forms exhibfted by the flseleton.^ from that confifting of one ieries of fimilar vertebras, of which the head feems to be only one more complex, to that which is compofed of a vertebral column, a head, a thorax^ and limbs," . The.t di^pofition of the mufcles varies according to the parts to whicli they are attached. The defcription of tne bones, and of the mufcles. Is followed by tables, the arrangement of which is fuch, that, while they prefcnt a nomenclature of tiie different parts, they exhibit their pofu'ion and their ufe. The fyftem of the bones, and that of the mufcles, conftitute the fouadation of the human body. They comprehe.id tte greater pajt of its material fubftance ; and the other organs exhibit a difpohtion which is ne- cefldtily fubordinate to them,. The cerebral fyftem is compofed; id. Of the ence- phalic organ, united to iis rachidian prolongation. 2d. Of the ganglions of fhe trifplanclmic. T'hefe two principal organs, analogous ones to v/hich are found in all vertebral animals, give rife to the nerves that diftri- bute the principle of motion and fenfation to the differ- ent parts. It is obferveu, that the nerves proceeding from the trifplanchnic are diftributed, in particular, to the vefTels which form the different tiffoes, and to the organs not fubje£l to the empire of the will j and that a 4 they Viii PRELIMINARY DISCOURSE. they thus maintain that animal life only which each poiiit of the organization enjoys ; while thofe which proceed from the encephalon and its prolongation furnifli to the principal fyftems of organs fecondary nerves, which communicate to them relative life, by caufing them to perform the fundlion for which they are proper. It is to be remarked, that in all animals the organs dif- tributed to the organs of the fenfes, and to fome other determinate parts, are always furnilhed by analogous pairs. The defcription of the cerebral and nervous fyftems in man is followed by a fynoptic table of the nerves, and an analytical view of the cerebral organ, v/ith a reca- pitulation of the various parts which compofe it, and of which the ufe is unknown. The organs of the fenfes are merely particular termi- nations of fome nerves, difpofed in fuch a manner as to perceive the fligbteft. impreflions, and to vary the forms of touching. An examination of thefe organs in the different clafles of animals, beginning at the fimpleft, and proceeding to the moft complex, affords the only mean of difcovering the ufe of the different parts which -enter into their compofition. Thus, it is feen that among the numerous pieces of which the ear is formed, the one moft important and indifpenfably neceflary for that organ is the membranous capfule, containing a vifcous fluid, amidfl which the auditory nerve expands j becaufe this capfule is found in all animals, and in fome it exifts alone. In the defcription of the digeftive fyftem, either In animals in general, or in man in particular, the pro- grefs of the aliments has been followed. It is curious to examine the gallric fyftem in different animals, from thofe PRELIMINARY DISCOURSE. IX thofe In which it confiils of a fmgle bag with one aperture, to thofe in which it is compofed of a mouth armed with teeth, with a tongue, with falivary glands* &c. J of an cefophagus, which leads to one, two, three, and even four ftomachs ; of inteftines of different lengths and fizes, provided with ccecal appendices, more or lefs numerous, and receiving fluids from a liver and a pancreas. The refult of digeftion is the chyle : this fluid, ab- forbed by particular veffels, proceeds into the veins; which affords a natural tranfition from digeftion to cir- culation. la the fyftem of circulation, the courfe of the blood has been followed, by giving a fucceflive defcription of the organs which it traverfes. This fyftem, in animals, exhibits a ftill more aftonilhing variety of forms than the digeftive fyftem. How many particular arrange- ments are obferved between the mammalia, whofe blood is warm and red, and the worm, which feems to have only one dorfal velTel, with branches ramified to every part of the organization, and a few external apertures which afford an entrance to the atmofpheric air ! , In man, the arterial blood, which Is of a bright red colour, and which has the temperature of 40 degrees of the centigrade thermometer (104^ Fahr.), when it proceeds from the aorta is diftributed by an order of velTels (the arteries) to every part of the organization. During its progrefs, it is continually freeing itfelf from the excefs of its heat by the excretion of the cutaneous organ ; and from its excefs of aqueous, faline, &c, parts, by urinary fecretion. The blood, after fupplying every part with the materials of nutrition, returns by two ordert X PRELIMINARY DISCOURSE. orders of veffels (the veins and the lymphatics), and tra- verfes organs which tend to make it refume the quahties of arterial blood. For the lymph, thel'e organs are : the numerous lymphatic glands i and for a part of the venous blood, the hepatic fyftem. The lymph mixes with the ■'■enous blood, and the veins unite into two large trunics, v\hich pour the blood into the right fide of the heart ; whence the whole of it pafles into the pulmonary organ : it is difLributed to every part of that organ, and auring the acl of refpira- tion refumes its former qualities of arterial blood : it then proceeds to the left fide of the heart, whence it pafles int'o the aorta, to ferve for a new circulation. In the defcription of the different parts of the fyftem of circulation, by thus following the pro2,refs of the fluids, the reader is bettei enabled to comprehend the organic phsenomena, and in fome meafure to divine the lavi's by which life is maintained. Having defcribed the organs which the blood tra- verfeSj and fhown the refults of the chemical analyfis of that fluid, an accurate defcription is given of the difbribudSn of the arterie?, followed by a fynoptical table of thefe vefl^els. This article is terminated by the veins and lymphaticc-j and in explainins; them the progrefs of the fluids is frill followed, as far as poflible, by taking the veflels at their capillary origin, and tracing out their union into ramufculi, rami, branches, and trunks. In explaining the lymphatics, the glands are defcribed in the order in which thefe veflels proceed thither. This method feems to be the moft convenient. In the laft place, this firft part is terminated by a de- fcription of the fyftem of generation, with a curious de-. tail of the parts of which it is compofed indifferent animals. 7 It PRELIMINARY DISCOURSE. XI It is there feen that if all organized beings are conneded by feme relations, it is in particular by the organs of ge- neration ; for an analogous ftrudlure of pans is found in all thofe the generating organs of which are known. This firft divifion concludes with a curfory examina- tion of the general ftruclure of the tiffaes, of which the different fyllems of organs are compofed. It is ob- ferved that all thefe fyflems (anatomical fyllems of or- gans) confifl of a certain number of fimilar parts. It is of very great importance to examine carefully their gene- ral difpofition, as thefe parts perform analogous func- tions, and as they experience derangements of the fame nature. This laft circumftance tends to throw great light on the hiftory of difeafes, which is treated of in the fecond part. In defcribing the different derangements to which the higher orders of fociety are at prefent fubjeft, it appears on the firft viev/ impoffible to clafs, in a proper manner, all thefe diforders fo exceedingly various : however, by the help of a few general confiderations, refpe£lin;g the whole of their mod important phcenomena, it will foon be found that they may all be arranged under a few prin- cipal heads, from which the particular cafes may be na- turally deduced. Difeafes confifl in an alteration of the parts, or in a derangement of their regular action. Th^fe which ex- hibit a manifeft alteration in the intimate flrudlure of the organs compofe i\iQ phleg?naji« \r, gcms^ &c.) require only, for their expanfion, to be feparated from their mother, and to be placed under certain favour- able clrcuradances [JJiti). In others, thefe fmall bodies (feeds^ova^ germs) are not completely developed until fome fpecific irritant has imprefTcd on them, by its conta6l, the firft movement of Individual life. Reprodu£lion, confidercd in tnis general manner, becomes a phjenome- non which can be reconciled m.uch better with the prin- ciples of philofophy, than that of the transformation of the aliments into a cnymous pulp. The lad pait concludes with a view of the knowledge acquired in regard to the vital i"an6lions. In this part, particul.T attention is paid to the development of the infelle£lual faculties. By obferving the ilov/ progrefs of this faculty, during the fa-ft years of life, and at the different periods of civilization, the fuccefiive connection of thefe different phaenomena can be followed. But if we negle6l to examine the earliefb exertioi^s of this func- tion ; if we attempt to account for the phsenomena ex- hibited by the human mind when it has attained to the hi^heil degree of inteliecSlual vigour, without having previoufly obferved, with care and attention, the deve- lopment of ti\Q different fenfations, and the origin of the racionai faculties, we {hall find nothing but an im- menfe abyfs, into which it will appear impoiTible to pene- trate,-and man will feera condemned to remain a flranger to the nature of that facult)' v.^hich he daily employs for making new discoveries. When it is conGdered that mtn who had no knowledge of the organic ilru£lure were the nrfl who attempted to PRELIMINARY DISCOURSE. XXV to explain the phcenomena of tlie mod aftonifhlng func^ tion of the organizatioiij the capital errors which occur in their works will not excite fo much furprife as the great difcoveries which they made. It may readily be conceived that the only proper me- thou to be piirfiKd in order to explain, in a fatisfa£lory manner, the fun£lion which gives rife to the intelietS^ual ■ faculties, is to acquire a thorough knowledge of the or« ganic flimSlure, and of the natural progrefs of the func- tions under the different circumftances of health and of difeafe. If in the fearch after truth vye (Dught to proceed from thin . known to thofe which are unknown, it will be neceflary to iludy the phyjical nature of man before we attempt to make refearches in regard to ^cvhat is beyond the province of natural philofophy, and which forms the. obje£l of metaphy/ics. CONTENTS CONTENTS OP THE FIRST VOLUME. System of the bones ' . ... Page 1; Of the trunk , . . . ib. Vertebral column in animals . , 2 Vertebral column in mai^ . . . . 5 Ribs and sternum in animals .... 10 Hibs and sternum in man .... 12 Of the head ...... 14 Cranium in animals ...... . ib. Cranium in man . 16 ii'ace in animals ..... o . 20 Face in man . . . ... : 23 Articulation of the head in animals . . 29 Articulation of the head in man . 30 Of the limbs in a-nimaes .... 33 Thoracic limbs of animals .... . ib. Shoulder in animals ...... . 33 Shoulder in man ,,..... 34 Tlie arm ...,.., 35 The fore arm • 37 The carpuS;, metacarpus, and phalanges in animals . 39 The hand of man . ..'.,. . 42 General disposition of the thoracic limbs . 45 Of the pelvian limbs in animals . . 47 Of the hip bone in animals . . . ib. Of the hip bone in man ..... , 43 Of the thigh ....... . 51 Of the leg . 52 Last CONTENTS OF THE FIRST VOLUME. XXV Xast part of llie pelvian limb in animals . - Page 55 Of die Jbot in man , , , . , . . * 56 General disposition of the skeletori . . . , GO Differences exhibited by the female skeleton - ' . 65 Development^ physical stiuctui-e_, and cliemical analysis of the boues ' . . , . ' . 6S Table of the bones . , . . ^2 MUSCULAR SYSTEM . . . . 7^ General disposition of the muscles in animals . . 76 Muscles in man . . . • , ib. Mtiscles which produce motion in tlie vertebral column 77 Muscles which give motion to the breast in animals . 79 Muscles which give motion to the breast in man . 80 Muscles which move the head In animals . .83 Muscles ^^'hich move the head in man . . 84, Muscles which move the face . . .87 Muscles of the limbs in animals . . Q2 Muscles \yhich move the shoulder on the breast . §4 Muscles which move the arm on the shoulder . q6 Muscles which move the fore arm on the arm . Qg Muscles which make the radius turn on the os cubitus . 101 Muscles which move tlie wrist on I he fore arm . 103 Muscles which produce motion in the fingers . , 107 Muscles which move the tliigh on the pelvis . .113 Muscles which move the leg on the thigh . .118 Muscles which move the foot on the leg . .132 Muscles which give motion to the toes . .127 Physical structure and chemical analysis of tlie muscles 132 Table of the muscles .... 134 CEREBRAL AND NERVOUS SYSTEM . .141 In animals without vertebrae '. . . . ib. In vertebral animals . , , , .142 Of the encephalon and encephalic nerves' . .143 Cei'ebral and rachidian prolongation with its nerves . 149 The ^ STV^ilt - CONTENTS OF The trisplanchnic Page 150 Cerebral and nervous system in man . ib. Encephalon . . ib. Meninx and uieningine . 153 Cerebellum . 164 Encephalic nervas . 167 First pair . ib. Second pair . . l68 Third pair . 169 Fourth pair . ib. Fifth pair . 170 Sixth pair . 177 Seventh pair . . 178 Eighth pair . ISO Nintli pair . ]Si Tenth pair . 182 Eleventh pair , 1S4 Twelfth pair . . 1S5 Vertebral nerves . ib. The cervical pairs . 186 Trachelo-cutaneous plexus . 187 Diaphragmatic nerve . 189 Brachial plexus . ib. Nerves of the thoracic limb . IfiO Dorsal pairs . 196 I ambar pairs . , 198 Sacral paii's / 200 Hypogastric plexus . ib. Crural plexus . 201 NeiTcs of the pelvian limb . ib. Lumbar plexus ^ • . . ib. Sacral plexiis . 203 Trisplanchnic ' . 208 CcHliac ganglion .211 Cueliac plexus . 212 Physical structiii'e and chemical analysis of the nei-ves . 213 Table THE FIRST VOLUME. XXIS Table of thf cerebral system Table of the nerves SYSTEM OF THE SENSES . 233 Of vision . ib. Cornea . 234 Aqueous hnmonr . ib. Ciystalline humour* . ib. Vitreous humour .' 235 H}'aIoid membrane . ib. The retina ■ . , . ^ . . 236 Sclerotica . ■ . . ib. The choroid membrane . 237 The orbits . 240 Muscles of the eye . 241, Eye-lids . ib. -Eyes in man . .244 Of hearing . 252 Organs of hearing in animals . ib. Organs of hearing in man . 257 Membranous labyrinth . ib. Cochlea . 258 Semi-circular canals . ib. Vestibulum . 259 Osseous labyrinth . ib. Tympanic cavity . ib. Small bones of the ear^ and mu 3cles attached to them . 2Sl Membrana tympani . 262 The concha . . - . 203 .Ligaments and muscles of the e ar ,. . . 264 Of smelling , .266 Olfactory organs in animals . ib. Olfactory organs in man . 271 Ethmoidal nerve , ib. Nasal fossae . 272 Superior and middle tm-binated bones , . 2/3 - Inferior Page 216 , 221 SXX CONTENTS OP inferior turbinated bone . : . Page 274 Cartilages of the nose^ and mtiscles attached to them . ib^ Sinuses . , ^ . i . ib. Etlimoidal cells . „ , i . TJS SYSTEM OF DIGESTION ... 270 System of digestion in animals i . • ib. Jaws , ... . .277 Teeth , , * . i i 279* The tongue ^ • . . * • 284 The stomach , i i i ' i i 285 Ventriculus or paunch .... 288 Honeycomb^ bonnet^ or king's hood . . . 289 Omasum or manyplies . , . , ib. Abomasum . ^ : . . . 29O Kumination , , l i. i i29i System of digestion in man . . . • 297 The teeth ..,.♦. ib. Inside of the mouth . . . . • 300 The tongue . 0 . . « • 301 0& hyoides . . . . . • 302 Muscles which move the tongue and the jaw i . 303 Velum palati . i , . ^ » 305 Salivaiy glands . ... • 307 The pharynx ...."• 307 Glottis and epiglottis . . . • 308 CEsophagus . . , . . • 309 The stomach . . . " . .310 Intestines . • . • - .311 Small intestines . . ' . . .312 Muscles of the anus . . . . .313 The liver . . : . • .314 The bile . . . . •. -310 Chemical analysis of it . * » .317 Bilious concretions . . . • .318 The spleen . . . , . . ib. The THE FIRST VOLUME. XXXI The pancreas . . . , s Page 319 ''The peritonaeum . . ^ . ib. SYSTEM OF CIRCULATION ' . ^ 321 Circulation in plants and zoophites . ib. In crustaceous animals, &C, . 32a In fishes . , '. . . 323 In reptiles . . • . . 324 In the mammalia and birds . 325 Urinaey systsm . . . . 327 Hepatic system ; 323 ReSI'IRATION . . 330 Tracheae .... . ib. Branchiae . , « . . 331 Lungs .... . 332 Apparatus of circulation in man . 33s Stracture and disposition of the arteries . 337 The skin . . .• . . .239 Dermis .... . ib. Mucous tissue . ib. Epidermis . . . 340 Super-renal glands . . . . 341 Kidneys .... . a>. Ureters \, . • , 342 The bladder • - . . . 343 Urine .... . 344 Chemical analysis of it s . 345 Urinar}^ calculi . S48 The veins .... . 351 Lymphatic vessels . 352 The liver .... . 355 Sinus of the venae cavse . 359 Pulmonaiy ventricle . 3(50 The lungs . . . , . 361 The larynx .... . 362 -Muscles of the larynx . :j63 Trachea XXXVl CONTENTS OF THE FIRST VOLUME. Trachea Bronchiae - . Sinus of the pulmonary veuia Aortic ventricle The pleura . ' The tJiymus . The pericardiiuii • The blood The lymph . Chemical analysis of the blood Distribution of tiie arteries The aorta . . Cardiac arteries Cephalic artery Anterior cerebral artery Brachial artery Axillary artery Humeral artery Radial artery- Cubital artery Thoracic aorta Abdominal aorta Gastric artery Hepatic ariery Splenic artery Pelvian artery Crural artery Taele of the heart and arteries Page 365 366 367 ib. 308 36^ ib, 370 371 ib.. 373 ib. ib. ib. 379 383 387 389 391 393 395 39 39s ib. 399 405 408 4lS A COURSE OF MEDICAL STUDIES. SYSTEM OF THE BONES, OF THE TRUNK. I, The organs of fmall animals are not fupport- cd by olleous parts ; bones in general are found only in the large animals ; and the fkeleton which they afford feems always to have been formed ac- cording to an uniform plan. The Ikeleton determines the general form of the animal ; it contains fome organs, and fupports others ; it affords points to which a great many are attached, and ferves for various kinds of motion. Animals which have a fkeleton are called ver- ielral animals \ becaufe in the Ikeleton the verte- bral column is a conflant and central part^ little fubjedl to variation, yoL. lo , B The a ' ORGANIC STRUCTtTRE. The Ikeleton of fome animals Confifts only of the vertebral column and the head ; but in the greater number thei'e are alfo limbs. 2,. Vertebral column in animals. The vertebral column, rachisj or fpine, is conftituted of a feries of longer or fhorter bones^ variable in extent, called vertehrce* The number of the vertebrae varies between i6 or 20 to more than 2,00 in fifhes, and in fome ferpents it exceeds 300. The vertebra are each compofed of one efiential part^ which is thehody of the bone ; the vertebra are placed one above the other, and in this manner form a column, which ferves to fupport the trunk, and as a ftay to the head. Vertebrae, of the limpleft kind, confift IbmC' times merely of the body. The more complex have alfo a groove or an arch, and protuber- ances on four faces. Each vertebra therefore has tv/o faces for joining to the two contiguous ver- tebrae ; the reft of its furface, whatever be its formj always prefents four faces: a fpinal face, fpna dorfi ; a praefpinal or antifpinal \ and two lateral ones. The fpinal face has fometimes a notch like that in the fpine of olTeous iifl:es ; but for the moil part it is furnifbed with a cavity which forms a hole. The SYSTEM OF THE BONES, 3 The jundlion of the notches produces a groove, that of the cavities a canal. The fpinal face exhibits alfo, in general, a pro- 'tuberance caUed the fpinal apophyfts : it is placed above the cavity ; ancrwhen there is a groove the latter is cut out on the apophyfis. The fpinal apophylis of the vertebrae varies both in its form and its dire61ion ; it is more or lets elongated, round or flattened ; terminates in a point or edge, and ftands at right angles, or pro- ceeds in an oblique dire(5lion. The prse fpinal face is feldom furnidied with apophyfes ; they are obferved however in the ver- tebra of the rattle-fnake, and in the tails of fifhes. The lateral faces fometimes have only apophyfes,, called tranfverfe, which fupport the ribs, as is the cafe in fiflies ; but other animals have two other articular or lateral apophyfes^ one above, the other below the tranfverfe apophyfis, and articulated with the adjacent vertebrae correfponding to them. On the (ides of the vertebral canal there are holes ox foramina, which penetrate to its cavity. Striking differences are obferved in the verte- brae in regard to their conne(?l:ion. In fifhes they join only by their fubftance ; in other animals they are articulated not only by their fubflance, but by their articular apophyfes. B 2 The 4 ORGANIC STRUCTURE. The articular faces of the bodies of the verte- brae have fometimes conical cavities, as in fifhes : the jun61ion then takes place by the edges of thefe cones ; and the rachis or fpine forms a ferics of elliptical cavities, filled with a white firm and elaftic or cartilaginous fubftance ; but for the moft part the vertebrag are articulated by their plain furfaces. The furfaces of the articular apophyfcs exhibit the fame difpofition. The difpolition of the articular faces of the ver- tebrae, and that of their apophyfcs, always deter- mine the quantity of motion they can perform by the means of mufcles J this motion may be ex- ceedingly free when the vertebrae are articulated only by their bodies, and when their apophyfcs do not mutually confine each other, as is the cafe in iifhes : by a contrary difpolition they may be reduced to perfc6l immobility. The rachis or fpine of fome fifhes con fi (Is of a long feries of vertebrse, which exhibit no diffe- rence ; in ferpents, a great part of the vertebrae are joined to ribs, but thofe towards the tail are not : the former are called dorfal, the latter cau- dal vertebra. The greater part of fifhes have alfo vertebrae united to ribs, and which are called dorfal ; but the caudal vertebrae, towards the belly, are ail fur- niQied with apophyfcs : fifhes have fome like- wife without ribs, and which exhibit no ventral apophyfcs : SYSTEM OF THE BONES. 5 apophyfes : thefe vertebrse, when they are above thofe of the tail, are called lumbar^ and when im- mediately below the head, cervical. In animals provided with limbs a part of the vertebrae receive the ribs ; another part are fuf- tained by the pelvis, and the reft remain free above and below the //^or^^ and the pelvis : this difpoii- tion allows us to diftinguifh the vertebrse into cer- vical, dorjal, lumbar, pehian, and caudal. The cervical vertebrse are between the head and the thorax ; the dorfal fuflain the ribs ; the lum" har lie between the thorax and the pelvis ; the pelvian are affixed to the pelvis, and the caudal come after the pelvis. The cervical vertebrae, in the mammalia *, are in number always feven : in the other claffes they vary. 3. Vertebral column in man. Man has feven cervical vertebrae, twelve dorfal, five lum- bar, five pelvian united and forming a bone called the os Jacrum, and three or four caudal alfo united, and forming a fmall bone called the coccyx, 4. Vertehra. In general, each vertebra pre- fents a body, on the fpinal face of which is a ring ; * Except the three-toed floth, which has nine. B 3 the 6 ORGANIC STRUCTURE. the whole of thefe rings forms the vertehrdl canal. Before the fplnal arch of each ring is a fp'inal afO" fhyfis % on each fide of the body or ring of the vertebras is a tranfverje apophjjis, and above and below this apophyfis there is another articular one. The articular apophyfes unite with the corre- fponding ones of the neighbouring vertebras. In the cervical and dorfal vertebra the facets of the fuperior articular apophyfes are turned backwards ; thofe of the inferior forwards ; while in the lumbar vertebrce the fuperior facets are turned inwards_, and the inferior ones outwards. Between the articular apophyfes of one vertebra and that adjoining to it is a hole which communi- cates with the vertebral canal (24 -vertebral fora- mina). In the cervical region the upper part of the bodies of the v^ertebrae is indented and inclined forwards: in the reft this part is plane and hori- zontal. The firft cervical vertebra, called the atlas, is a fimple ring, v/hich on its upper part and both fides prefents two facets for its articulation with the head, and two others below unite it with the fecond. The internal part of the arch of this firft vertebra exhibits alfo an articular facet, which fupports a round and elongated tubercle (procejfus odontoides) which rifes above the body of the fe- cond SYSTEM OF THE BONES. J cond vertebra. The latter is diftlnguifhed by the -name of ^A'o/V/^i". The free motion of the head on the neck arlfes from the peculiar difpofition of thefe firll: two vertebra?. The cervical vertebrae are the fmalleft : their bifid or forked fpinal apophyfes are turned a little downwards, and their tranfverfe apophyfes are perforated. In the dorfal vertebra the fpinal apophyfes, which are longer^ are turned downwards ; the tranfverfe apophyfes, inclined backwards, exhibit demi-facetSj which when united form complete fa- cets deflined to receive the ends of the ribs ; the ift, theiith, and the 12th, exhibit a complete facet. Thefe facets are turned downwards in the upper vertebrae, and upwards in the lower. T^e lumbar vertebrze are the largeft ; their fpi- nal apophyfes are exceedingly llrong, and turned backwards. The different regions of the fpinal and pracfpfnal face of the rachis or fpine are alfo dillinguithed by peculiar names. Thus the region of the neck behind is called the cervical, and before the trachelian ; that of the back behind, \\\c dorfal, and before, the prcedorfal\ that of the loins behind, the lumhari and before, the prcsJumhar. The five pelvian vertebras united into oiie piece B 4 form S ORGANIC STRUCTURE. form the os facrum j this bone is flat, triangular, and concave before ; its broad and thick fummit is united by its body and articular apophyfes to the laft vertebras of the back ; it is articulated on the tides with the bone of the pelvis, and on the lower part it receives the coccyx. The OS facrum exhibits on the fore part the bo- dies of the vertebrae of which it is formed, and behind and on the fides confufed tubercles, which correfpond to the fpinal, tranfverfe and articular apophyfes. This bone contains alfo the vertebral canal ; the foramina which terminate there are placed before and behind. The laft three or four vertebrae, united into one, form a fmall bone called the coccyx^ which termi- nates in a point, and is articulated with the os facrum, on which it can be bent backwards. In this bone the form of the vertebras which compofe it can fcarcely be diftinguifhed. Between the articular furfaces of thebodies of the vertebrae is found a white, thick, firm and elaftic fubftance, the conliftence of which decreafcs from the circumference to the centre ; this fubftance, which is of a fibrous ftru(?l:ure, adheres ftrongly to the two contiguous vertebrae, and is called the I'l' gamentO' cartilaginous. The bodies of the vertebrae are covered, both before and behind, with a tiffiie of white compact fibres, exceedingly folid, which proceeds along the whole SYSTEM OF THE BONES. ^ whole rachisor fpine, and which iscaJled the com^ mon vertebral ligament, pqfterior and anterior. On the poilerior part of the fpinal canal, and between the articular apophyfes, there are exceed- ingly ftrong ligaments called the infermeJIar (yd- low ligaments). The vertebrae are faftened alfo behind by a liga- ment (the Juferffinal), which proceeds along the fpinal apophyfes, and by ligaments (inier^ fpinal) which extend from one fpinal apophy (is to the other. Each articular apophylis has alio a ftrong capjular ligament^ which is united to the neighbouring vertebra*. The means by which the lail: lumbar vertebra is united to the os facrum are the fame as thole by which the vertebra are united \ and the coccyx is fixed to the os facrum by an articular capfule and ligaments placed before and behind. The vertebral column is flightly bent, in fuch a manner that the region of the neck before is con- vex, that of the back concave, the region of the loins convex, and that of the os facrum concave. Though each vertebra can perform on the other * All the articular furfaces of thcfe bones, which form moveable articulations, are covered with a ftrong, Imootb, and mpift cartilage, and the portions of the bone articulated arc always wrapped up in a fort of membrane which furrounds the articulated furfaces in the form of a capfule, and keeps them firm. This arrangement being always the fame, we (hall diitin- gulih it In future by the name of the ariuular ea^uh. only 30 ORGANIC STRUCTURE. only very confined movements, the whole ofthefe movements is complete in the whole of the fjJine; find it is well known how far thofe who exhibit feats of tumbling can carry the pliability of this part. It is by means of the lumbar vertebrae in particu- lar, that the greatefi: movements of the trunk arc performed, becaufe they are not impeded in their motion, like that of the back, by being united to ribs. The body of the vertebras confifts of a fpongy fubftance ; that of the apophyfes is of a more com- pact texture. 5. The ribs and sternum in animals. In almofi: all animals the ribs are fuftained by a greater or lefs number of vertebra. Thefe bones, however, are wanting in many cartilaginous fifhes ; in fome of the ofleous, and in frogs. The number of the ribs, which varies from 12, to 23 on each fide in the mammalia^ is conliderably increafed in ferpents and fome fidies, which have almofl: as many as they have vertebrae. In the falamander, and feveral genera of fiOies, the ribs are very fhort ; in fome of the mammalia they are exceedingly long. In the herring they are as fmall as needles ; and in the two toed ant- eater (myrmecophaga d'ldaByla) fo long that they cover each other like tiles. In general they are fingle ; in feveral fiOies however they are forked, fo that two proceed fometimes from the fame vertebra. The SYSTEM OP THE EONES. H The ribs arefeldom flraight, being for the mod part bent almofl in a femicirciilar form j in the mammalia their configuration determines the form of the thorax, and in iiflies and ferpents that of the belly and abdomen. One of the extremities of the ribs always rells on the vertebra ; the other either remains ful^ pended in the foft parts, or is united with a carti^ lage, or refts on a bone called the fiernum*. The name o^ Jlernal vertebras is given to thofe which on one fide adhere to the vertebrae, and on the other to the fternum ; and that of ajicrnal to thofe which do not reach the fternum. In mammalia the ribs which end at the fternum are towards the head : birds, in general, exhibit a contrary difpofition ; fometimes, however, they have afternal ribs towards the head, and towards the pelvis. In animals which have no ribsf the fternum is wanting; ferpents and almoU all fillies have ribs, without having a Iternum. In mammalia the fternum is long, of confidera- ble thicknefs, broader or narrov/er, ofl^ous, fome- times cartilaginous, and, for the moft part, confifts of feveral pieces. In birds it is exceedingly broad as well as thin, and prcfents a very falicnt ridge, particularly in thofe of a rapid flight. In fome reptiles it afiiimes very Angular forms, is extremely fmall, and in fifties is always wanting. * In the crocodile the ventral ribs reft on the as puhh, ■\ Frogs excepted. 6. Ribs ■I 12 ORGANIC STRUCTURE. 6. Ribs in man. In man the ribs are In num-* ber 1 2 on each fide ; thefe bones are long, bent irregularly in a femicircular form ; are flat, and, as it were, tvvifted backwards. At the pofterior extremity they prefent a double facet, feparated by a falient line ; this facet is articulated between the lateral parts of two verte- brae. At the diftance of an inch from this extre- mity, the ribs have behind another facet, which is applied to the tranfverfe apophylis of the lowed vertebra. Thefe two articular facets are feparated by a narrow part or ftri6lure called the neck of the rib. The ribs then proceed backwards and outwards, after which they feem to bend in order to proceed forwards, and exhibit there a falient line nearer the vertebrae in the upper ribs : angle of the ribs. The anterior extremity is terminated by a car- tilage which unites with the flernum ; the flrft feven ribs (Jiernal ribs) proceed in this manner to the fterrium, by means of an Intermediate carti- lage ; but the cartilages which terminate the lad five (ajiernal ribs), reft one' upon the other, and are fupported by the cartilage of the feventh rib. The firft rib, which is fhortcr as well as broader, has its external face turned upwards ; the follow- ing ones become longer, and their external faces are directed forwards : in the laft place, the lower ones become fhorter, and obferve the fame di- redion. The SYSTEM OP THE BONES. Ig The articulation of the ribs with the bodies of the vertebrae is fecured by an articular capfule, an anterior ligament, a cojio-tranfverfe ligament, and an interior ligament. The tuberotities of the firll ten ribs are faftencd to the tranfverfe apophyfes of the correfponding vertebrse, by a capfule and a tranfverfe ligament, 7. The fiernum. The llernum in man is a flat bone, broad at the top, and terminating at the bottom in a thin cartilage of greater or lefs extent. This bone often confifts of two pieces, which in the courfe of time coalefce and adhere together ; on the fides it has facets proper for receiving the cartilages, which terminate the ribs ; thefe carti- lages are retained on the fides of the fternum by an articular capfule, an anterior and 2^ poflerior ligament. The fternum is exceedingly fpongy ; it is co- vered both without and within by a very ftrong feriofteum. The fternum, the ribs, and the dorfal vertebras, form the capacity of the thorax. The thorax has the form of a very irregular cone, with its bafe truncated downwards and its fummit upwards ; it is flat before, broad and rounded on the fides ; flat and concave behind. OF OF THE HEAD. The bead, with refpe^l to its form^ is that part of animals moft liable to variation. It confifts of the cranium and face. 8. Cranium of animals. The cranium, or cephalic cavity, contains the brain. The differences obferved in the form of the head of various animals confift chiefly in the greater or lefs contraction of the fize ef the cra- nium, and the greater or lefs elongation of the face. The extent of the cranium, in regard to the elongation of the face, may be determined on the furface pre fen ted by a vertical fe6lion of the head : it is obferved, in the different clalies of the Hiamraalia, that the cranium occupies on this fur- face an extent either greater or fmaller than or equal to that of the face, without including the lower jaw. Among the Europeans the extent of the cranium is nearly quadruple that of the face; in the Negro, the extent of the cranium remaining the fame, that of the face is increaled by a fifth, 6 In tSYSTEM OF THE BONES. 1^ In the different fpecies of apes, the area of the face goes on increaling ; in the mandril (fima maimon), and the greater part of carnivorous animals, it is nearly equal to that of the fkull ; in various fpe- cies of graminivorous animals it continues to in- creafe to fuch a degree, that in the horfe it is aimofl four times as large as that of the cranium. The cranium aife(5ts different forms ; but for the moil part the fammit of it is oval or rounded, and its bafe an irregular plane. The bones of the cranium do not feem io be numerous in fifhes ; in thefe animals they are united by plane fur faces, coal efce at an early period, and leave fev^ traces of their jun6lion. In birds thefe bones, vi^hich are fitted into each other by futures, coalefce alfo at an early period, and the traces of their articulations difappear vi^ith age. But in the greater part of the mammalia, and in man, thefe ofleous pieces are very difi:in6l, and at moft are eight in number, viz. before, the frontal bone ; on the fummit and at the fides, the two parietal bones ; lov/er, and on the fides, the two temporal bones ; behind, the occipital ; and at the bafe, the fphenoid, and the ethmoid. Of thefe bones the frontal, fphenoid, and ethmoid, are com- mon to the cranium and face. Thefe bones, in the various genera of animals, differ both in regard to their form and relative iize; and it may thence be eafily conceived in how \ i6 ORGANIC STRUCTURE* how many ways the configuration of the craniUM may be varied. The cavity of the cranium in the mammalia 19 filled with the brain ; but in reptiles and fifhes this organ occupies only a very fmall part of it* 9. Cranium in Man. In man the craniunni is round, well expanded, and prefents a cephalic cavity comparatively larger than that of other ani- tnals : it confifts of eight bones. 10. The fpbenoij, osJpbenoiJes. The fphenoid is a bone of a very lingular form ; it is long, and pafles through the middle of the bafe of the era* nium and the breadth of the face ; it exhibits a great number of prolongations, by means of which it is articulated with all the bones of the craniumj and a great part of thofe of the face** On account of its poiition and numerous con- nexions, it has been confidered as the key of the * This bone has on each of its fides two unequal prolonga- tions, called the greater and kfs ala, or wings j at the bottom two apophyfes, called the pt. rygoides, which form behind ^fojfa^ called the pfery^i^y id. The m ddle of this bone is the body of it, the upper part of which, traniverlally deprefled, is known under the name of the fella tuc'ica, and is furrounded by the anterior and pqjierior dmoid apophyfes. This bone is perforated with holes, which afford a palfage to the velfels and nerves j viz. the optic foramina, the fphenoidal fiffures or rima, the upper and lower maxillary foramina, and tvvo fpinous foramina. In the thicknefs of the body the fphenoid tiffurcs are contained. 8 bones SYSTEM OP THE BONtS, If bones of the head. This bone is feen atthebafe and on the (ides of the cranium ; it concurs to form the cavities of the eyes (brhits) and thofe of the noftrils. i I. Frontal Ip&ne^ Os frontis. The interior part of the cranium, or the forehead, is formed by the OS fro7itis. This bone forms alfo the upper part of the face; it is curved and rounded at the fum- rriit, and before and on the fides it is furnifhed with two procefles called the frontal, and in the middle, lOwer down, with two other procefles called the nafal. The lower part of it is bent back in a horizontal direction: it is indented in the middle to receive the os ethmoides, and its fides are arched to form the upper part of the cavities that contain the eyes. This bone is articulated below with a part of the bones of the face ; behind and at the bottom with the OS fphenoides, and behind and at the top with the two following bones. 12. Parietal hones. The fummit and fides of the cranium are formed by the junction of the two parietal bones. Thefe bones are convex o\\ the outfide, quadrilateral and ferrated at the edges; they articulate with each other above, with the frontal bone before, with the occipital behind, and at the bottom with the temporal bones and the fphenoid. 13. Occipital bone^ Os hajilare. The back of the VOL. I. c head 18 ORGANIC STRUCTURE. head and a part of the bale of the cranium are formed by the occipital bone. This bone is con- vex on the outfide, and is (haped fomewhat like a lozenge ; two of its edges arc fitted by a future into thofe of the parietal bones ; it prefents on the outfide behind a protuberance called the oar- pi/al; on the (ides two falient lines called the occipital arch, and within two grooves, corre- fponding to thefe two lines. Below is the large 6ccij)ilal foramen, which forms a continuation with the vertebral canal; on the fides and within this hole there are two tuherofities, with which the head is articulated on the vertebral column : in the lad place, that part of the bone before this hole is called the fuh-occipital. This bone has feveral holes sJfo for the pafllige of the nerves and veilels : it is articulated before with the fpbenofd ; at the bottom with the rachis or Ipine, and on the lides with the following bones. 14. Temporal hones, Ojja temporaTm. Below and on the fides the cranium is formed by the two ieu'cporal hones. Thefe bones are of a very irregu- lar figure ; they prefent on thj| outfide a flat, rounded, and very thin part, which may be com- })ared to the fhell of an oyfter. This part gives- birth on tlie outfide to an apophyfis^ called the zygomatic : it is long and thin ; proceeds forwards,, receding from the bone, and unites with the apo- phyfis of another bone to form an arch. 5 Below SYSTEM OP tHE BONES, \g Below the root of this apophyfis Is the glenoid cavity, which receives the condyle of the jaw- bone J beyond this cavity is the external orifice of the meatus auditorius ; and ftill further back is a large tiiberofity called the apophyfis maf- toideiis. Below the auricular foramen is the apophyfis Jiyloides, behind which is obferved the jugular foffa. The temporal bone prefents, in the infidc, a thick part, very unequal and exceedingly hard, called the petreous (petrea). It is in the thickncfs of this part that the organs of hearing are con- tained. The temporal bone exhibits a great num- ber of holes, the principal of which are : below, \k\Q Jiylo-maftoid \ the carotid canal, and the gut- tural conduit of the ear ; in the infide, the interior auditory foramen. This bone is articulated before with the fphe- noid ; at the top with the parietal bones, and be- hind with the occipital. 15. All thefe bones of the cranium are fitted into each other by futures, or are united by fmooth furfaces. Their edges are generally cut obliquely in different dire(?tions : the fame bone, therefore, has a part of its edge cut into a bevel on its ex- ternal face, while in another part it is cut into a bevel on the internal face. This peculiar difpofi- tion exhibits a method of articulation as flrong as it is wonderful. > ^^ c 3 Thofe 10 . ORGANIC STRUCTURE, Thofe ferrated marks produced by the jiincllonr of the bones of the cranium, and which are called futures, become effaced fometimes with age. The bones of the cranium exhibit, on the out- ilde, particularly behind and on the fides, depref- fions or afperities, that indicate theinfertion of the nnifcles by which thefe parts were covered. In different points of the bone of the cranium there are holes of greater or lefs fize, which pene- trate to the iBtide of it^ and which afford a paffagc to the veflels and nerves. The upper part of the cavity of the cranium- exhibits furrows of greater or lefs depth, which extend in a ramrfied form. : thefe furrows corre- fpond to veffels which reited directly againft the lides of this cavity. The bafe of the cranium prefents, in the infide, eminences and excavations very diftindl: the ca- vities fcrve to lod_Q;e portions of the cerebral organ. The lower part of the cephalic cavity, the bafe of the cranium and the fides of the head contain holes, iiffures, and indentations, which afFord a paffagc to the nerves., arteries, and veins'^., The * The inferior fide of the cavity of the cranium, proceeding from the fore part backward', prefents the foramen civcwn, the holes with which the cribriform plate of the os ethmoides is per- forated, the optic foramina, the fphenoid fiflures, the up.per maxiilas-y or large round foiamina, the lower or oval maxillary forajr.ine. SYSTEM OP THE BONES. 21 The bones of the cranium are covered through- out their whole extent, both internally and exter- nally, by a membrane which firongly adheres to them, and is called the pericranium. i6. The pace in animals. It is of import- ance to examine the face in the different clafles of animals, in regard to its greater or lefs elon- gation. The form and elongation of the face are always determined by the gre>ater or lefs prolongation of the jav/-bones. It is obferved, in general, that the perfedlibility of the oriranization of animals is in the inverfe foramina, the fpinous or fmall round foramina, the internal orifice of the carotid canal, the polterior foramina laqcra, titie anterior foramina lacera, the interior auditory foramina, the pof- terior foramina lacera, the anterior condyloid foramina, (ihe pofterior condyloid, and the maftoid foramina when they exift,) the grand occipital foramen. The bafe of the cranium prefenlsj the anterior palatine fora- mina, the pofterior palatine foramina, the pollerior aperture of the nafal foffas, the potterior aperture of the palatine and pterygoid cbnduits, the anterior foramina lacera, the carotid canal, the pofterior fora-mina lacera, the anterior and pofterior condyloid foramina, and the grand occipital foramen. The lateral parts of the head prefent: the external orifice of the auricular conduit, the ftylo-maftoid foramina, the fpinous foramen, the foramen ovale or lower max'.lhry foramen, and tie zygomatic fofla, in which are found : the fpheno-maxillary rirr,a or fiffure, the orifice of the fub-orbitar, palatine, pterygoid, fpheno-palatine, and pterygo-palatine conduits. c 3 ratio 22 ORGANIC STRUCTURE, ratio of the fmall fize of the cranium, and of the great elongation of the jaw-bones. Man, of all animals, has the largeft cranium and the fhorteft face. To determine the iize of the facial angle, Cam- per raifes above the plane of the bafe of the cra- nium a line, which, proceeding from the bottom of the upper incifive teeth, is continued in an in- clined diredlion to the moft prominent- part of the forehead. The plane of the bafe of the cranium pafles through the anterior edge of the upper part of the noftrils, and through the auditory foramina. The facial angle, therefore, in Euro- pean heads, in general, is equal to eighty degrees ; in thofe of the Moguls to feventy-five ; in thofe of the negroes to feventy ; in the orang-outang to fixty-five ; and in carnivorous and graminivorous animals it decreafes fo much, that in the horfe it does not exceed twenty-three degrees. This meafurement, however, can be applied with any degree of exa6lnefs only to the human fpecies and the quadrumana* . Other animals ofteq prefent fuch fingular configurations of the head, and fuch projedlions of the forehead, that it is not poffible to meafure their facial angle : befidcs, it is of little importance to obfervc the fize of the * Quadrumana — anltnjtls with four bands ; a name given to the fecond order of the mavimalia in Blumenbach's Handhucb dex Naturgefchichte, The fpecies included under this denomination are : Simia, Papio, Cercopithecus, Lemur. In the older works Ihis order were called Pitted. Trans. headi System of the bones, 23 head, except in regard to the volume of brain it is fappofed to contain : in reptiles and fifhes the brain occapies oply a fmall portion of the cavity of the head. The Greeks feem to have been fentible that the beauty of the head confifts, in an eflential manner, in a ftraight dirediori of the face. In the ftatues of the gods or heroes whom they were defirous of ranking among their deities, it is feen that they increafed beyond the hmits of nature this angle, which they fometimes carried to nearly a hundred degrees, as if they wifhed by thefe means to give them a form of countenance as far diftant as poi""- iible from that of the common race of men. It is obferved, indeed, that the ftraight form of the face generally pleafes even in animals ; and, on the other hand, that elongation of the muzzle and depreffion of the cranium give them always a ftupid and ferocious air. 17. Face tn man. In man the face has fuch fi diredlion that the eyes and mouth look diredtly forwards : the bones which compofe it ^vc in number fifteen, befides the thirty-two |:eeth. 18. Bones op the nqse. The bones of the nofe are fmall, obloftg, and quadrilateral ; they coalefce, and thus form the upper part of the nofe; they reft laterally on two prolongations of the upper jaw-bone ; at the top they are indented c 4 into '14 ORGANIC STRUCTURE. ■ into the frontal bone and the os ethnoidesx their lower edges being iKt(t are united with the carti- lage of the nofe. 19. Lachrymal bones. On the interior fides qf the orbits there are two fmall thin bones, called the lachrymal bones. Their exterior face exhibits a portion of ihe fo/fa lachrymaUs, which has a com- munication with the noftrils. Thefe bones are articulated : at the top, with the frontal bone; before and below wiih the upper jaw-bone, and behind with the os ethmoides and the lower cornets. 20. Zygomatic hones. On the external and lower fide of the orbits there are two fmail bones, called Zygomatic hones, of a very irreg'ular form, flat, and fomewhat convex on the outfid.e. Their circumference cpnfifts of four indented fides, terminated by pretty long angles, called apophyjes. Thefe bones are articulated : by the upper apopbyfis, with the os frontis and os fphenoides ; before and at the bottom with the upper jaw- bone. The pofterior apophyfis, being longer, is united with that which, as already faid, rifcs above the temporal bone, and forms with it an arch., called \}i\& zv somatic. 21. Super-maxillary hones. At the middle and anterior part of the face are the two bones of the upper jaw, or iho, fuper-maxillary bones: thefe bones are each very irregular, and prefent, in thejr thicknefs. SYSTEM OF THE BONES. 25 thicknefs, a cavity called the fmus maxillaris, which has a communication with the noftrils. The fuper-maxillary bone is connedled with all the bones of the face, and with a part of thofe of the cranium. It is articulated : in the inlide and at the bottom with its fellow, and two other bones ; before and at the top it has an afcending apophy- lis, called the orbito-frontal, which refis on the frontal, the lachrymal and the ethmoid bone, and which fupports the bones of the nofe : at the bot- tom and on the 'outfide of this apophyfis is the an- terior orifice of the fub-orbitai conduit : on the outfide it is united to the zygoma and the os fphe- noides : before and below it receives the teeth. Thefe bones conftitute a great part of the face and palate ; they contribute to form the arch of the nofe, the fides of the noftrils and thofe of the orbits ; they receive the upper teeth, and ferve for raaftication. 22. Bones of the palate. Behind the fuper- maxillary bones, and at the bottom of the noftrils, are the palatine bones, which extend as far as the orbits, Thefo bones are of a very fingular form, which is necefiary that they may adapt themfelve?, in a great part, to the flill more lingular form of the 05 fpbenoides. Thcfe two bones unite behind the fuper-maxil- lary bones : they are articulated with the os fphe- no'ideSy the os ethmoides, and the lower cornets (turbinated bones j), and with an ofl^eous plate, 23. The '26 ORGANIC STRUCTURE. 23. The OS ethmoldes. In the infide, and at the fummit of the noflrils, under the indentation of the frontal bone, is the os ethno'ides, of a cubic and elongated form, exceedingly light, and con- flfling in the inlide of very thin plates^ pierced with holes, which affume different directions, and form a great many cells communicating with each other. The upper face of this bone, wJiich is of a more compa(5l nature than its interior plates, is pierced with about forty holes : it correfponds on the in- iide to the bafis of the cranium, and in the middle prefents a ridge. Its compact lateral faces concur to form the internal fides of the orbits. The other faces of this bone are open, and correfpond to the fummit of the noftrils. This bone is articulated : at the top with the frontal bone ; before, with thofe of the nofe, the fuper-maxillary bones and the lachrymal ; behind, with the OS fphenoides^ z.nd. the palatine bones; at the bottom and in the middle with an olieous plate. 24. Lower cornets. TurVinated hones. Be- neath, and on the fides of the os eihmoides, are two bones, formed of plates twifted into the form of cornets : thefe bones feem to be an appendix of the OS ethmoides, and are united to it. Thele cor- nets are articulated alfo with the fuper- maxillary bones, the lachrymals, and the palatine bones. 2^, The vomer. Between thefe two cornets, below SYSTEM OF THE BONES. ^27 below the os ethmoides, and in the middle of it, is an ofleous plate, which divides the nafal fofTse into two noftrilsj and is called the vomer. This thin and long bone receives in a groove, on its anterior and fuperior edge, a cartilage, which completes with it the partition of the noftrils. It refts at the bottom, between the two fuper-maxillary and the palatine bones, and is joined at the upper part, in the bottom of the noftrils, to the middle of the OS JphenoideSy by a large farrow. 2.6. Jaw-bone. The lower jaw-bone, oj- ;;w.v/7- lare, is formed of a bone turned round on its plane face, in the form of'a horfe fhoe : the upper edge of the arc contains fixteen cavities for receiving the teeth ; the lower edge has a projection to- wards the chin. Behind and in the middle of this bone is a fmall ridge, called the genian ; on the (ides of which are two falient lines, known by the name of the mylean, which terminate at the orifice of the dental canal. The extremities of the maxillary bone are bent up and backwards, fo as to make an angle of forty- iive degrees, and terminate in a flattened trans^ vcrfe tuberofity, called the condyle, which is re- ceived in the articular cavities of the temporal bones. Before this tuberofity is a large apophyfis, called the coronoid, feparated from the condyle by ^ deep indentation. The 28 ORGANIC STRUCTURE. The articulation of this bone with the temporal isfecuredbya lateral external ligament, a lateral internal, a tylo-maxillary ligament, and an articu- lar capful e. Mammalia, iifhes, and many reptiles, have jaw- bones armed with teeth. {See that part of the article on Digejiion.'j J* 27. The bones of the face, as well as thofe of the cranium, arc conne6led by means of fmootb furfaces or indentations, and exhibit the fame pe- culiarities. Thefe bones have alfo depreffions or afpcrities in thofe places to which the mufcles were attached, and holes through which the vef- fels and nerves were conveyed to the interior parts. The aggregate of the bones of the face, toge- ther with thofe of the cranium, feem fitted in a fpecial manner, by the formation of the cavities of the ear, the orbital foflae, the nafal foffe, and thofe of the mouth, for receiving the organs of hearing, of fmell, and oftafle. The aggregate of the bones of the cranium and face exhibit alfo externally fome remarkable parts, to which it is of importance that particular names fhopld be ailigned, in order that they may be diftinguiOied Vv-ith greater facility. Thqs the cranium has on each tide a large depreflion called the temporal fojja, the lower end of which termi- nates near the zygomatic arch : below this arch is another SYSTEM OP TI-iE BONES, 2g' another cavity of lefs breadth, but much deeper, known by the name of the zygomatic fojf a *. 28. Articulatioi<3" of the head. The head, in diff^^rent animals, is articulated on the .vertebral column by one or two condyles. In birds, it is articulated behind by a tingle condyle, which allows it to perform fuch extenfive motion, that thefe animals can turn their beaks diredly behind them : it is well known that they often fleep with their heads refting in this manner between their wings. A part of the oviparous quadrupeds have only one occipital condyle. Ser- pents have a condyle with three facets, difpofed in the form of a trefoil leaf, which admits of few movements. Moil fifhes have only one condyle> '* If the face be examined from the top downwards, we ob- ferve the holes of the eye-brows and the orbital foITse. In the orbital foffae are found the ocular holes, the fphenoidal and fpheno-maxillary fiffures, the internal orbital foramina, and the orifice of the nafal canal. Below and between the orbital foflae the face prefents the aperture of the nafal foflae. In thefe foffae are feen the holes ©f the cribriform plate of the os ethmoides, the apertures of the fphenoidal finuses and of the pofterior ethmoidal cellules ; the aperture common to the anterior ethmoidal cellules and the frontal finuses j thofe of the maxillary finuses 5 the fpheno-palatine foramina; the orifices of the anterior palatine conduit. On the fides of the nofe, the face exhibits alfo a fofla called the canine., in which is feen the fub-orbital foramen ; and below, the face exhibits the dental arches and the orifices of the-, dtntal canals. articulated 3d ORGAlSriC STRUCTUE.Ec articulated by means of cartilages, and which pof- feOes very little mobility. The ray and fhark have two condyles, the movements of which- are very confined. In the lafi: place, the mammalia are furnifhed with two condyles, more or lefs moveable. In man, the head is articulated on the vertebral column, by means of two tubercles or condyles, placed on the fides of the large occipital foramen. Thefe condyles are fitted into two articular cavi- ties fituated on the interior fides of the arch of the iirfi: vertebra. This difpofition of the articular faces permits the head to perform, with eafe, every motion of flexion and extenfion, backwards and forwards; and eafy movements of flexion and rotation on the fides. But the movements of rotation performed by the head are owing, in a great meafure, to thofe performed in the fame diredion by the firfi ver- tebra on the fecond ; and the different motions which the vertebrae perform on each other contri- bute greatly alfo to thofe of the head. The articulation of the head is fecured by an anterhr liganmit ^ a fojier'ior ligammt, and an articit' lar capjide. The peculiar articulation of the fecond vertebra on the lirfi-^ by means of the odontoid apophyfis of the latter on the anterior arch of the former, is fecureti SYSTEM OF THE BONES. 31 fecLired by a tranfuerfe ligament, two lateral Tiga" merits, an accejjory ligament^ and a capfide. The middle of the head is not placed exadily on the vertebral column, and Camper has obferved that the part which is before the large occipital foramen is to that behind in the ratio of eisrht to o (even ; fo that the head, when not kept back, has a flight tendency to fall forwards. In the vertical pofition of the head, man alone has the mouth and eyes looking diredlly forwards. We have already feen that the trunk and head confift of bones, fome of v/hich have always their fellow, while others are conftantly fingle. The fingle bones are always fymmetric, that is to fay, can be divided into two iimilar parts : thefe are necefTarily placed in the centre. Bones which have, fellows are always placed one on the right and the other on the left, and never have a fym* metric form ; fo that in the ikeleton one fide Jias a perfed refemblance to the other. OF 32 OF THE LIMBS. 29* Limbs in animals. The fkeleton of fer- peots and fifhes confifts of the trunk and the head only ; but that of oviparous quadrupeds, birds, and the mammaJia, is provided with four limbs, which feem always to be formed according to an uniform plan. The fins of fifhes, properly fo called, are parts' which fupply the place of limbs; but they catinot be confidered as fuch, becaufe their ftru6lure has nothing in common with that of limbs. Cetaceous animals, which are not fifhes but amphibious mammalia, have no fins : their place is fupplied by limbs, which difcharge the fame functions. The limbs of all animals are conftantly fixed, fome to the breafi: or thorax, and others to the pel-vis; on which account they are diilinguifhed into thoracic and pelvian. Thoracic Limbs, 30. Thoracic limbs of animals. The thoracic limbs of animals are thefhoulder, the arm, the fore-arm and the hand j the pelvian are the haunch, the thigh, the leg, and the foot. Th^ SYSTEM OP THE BONES. 33 The other limbs vary in the different orders of animals, with refpet^t to their form, their length, their covering, and in particular their termination. This variety determines the ufe for which they are proper. 31. Shoulder in animals. The fhoiilder confifts fometimes of one bone called the fcapula {omopJata), as is the cafe in the mammalia with folid hoofs. It is often formed of two bones, ih^ fcapula and the clavicle ; as in the qnadrumana, fome car- nivorous animals, and the greater part of oviparous quadrupeds. In birds it confifts of three bones, viz. the fcapula, the clavicle, and the tbrk. The fhoulder has always a cavity which receives the head of the bone of the arm. This cavity, for the moft part, is contained in the fcapula; fometimes it is formed both in the fcapula and clavicle i and in birds the three bones of the fhoul- der concur towards its formation. The fcapula is generally a broad, flat bone, iituated at the dorfal part of the ribs ; but its figure greatly varies in different animals : in the mole it is exceedingly long; in the frog and the toad it confifls of two pieces. The clavicle, for the moft part, is a long thick bone, refting on one fide on the top of the fler- num, and on the other againft the fcapula, which it VOL. I. D fixes 34 ORGANIC STRUCTUEE.' fixes and keeps back. All animals provided vvitfe this bone can extend their thoracic limbs forwards, with more or lefs facility to perform great motions. Animals which have no clavicle employ their thoracic limbs only for walking. Some carnivo- rous animals, fuch as dogs and cats, &c. have the clavicular bones fufpended in the flefh, and alto- gether ufelefs. The forked bone in birds has the form of the letter V ; it ferves, together with the clavicle, to keep the fcapula firm, and to prevent it from ad- vancing too far forwards: birds v^hich have this bone exceedingly flrong fly^ in general, with great rapidity. The fhouldcr of the tortcife confifts of three bones, analogous to thofe of birds : thefe bones coalefce, and form only one offeous piece. The (houlder, in man, confifts of the fcapula and the clavicle. 32. The fcapula. The fcapula is a flat, thin, triangular bone, placed at the lop of the back be- hind, having one of its angles elongated and di- rected downwards. Its external face has, at the upper part, a pro- jecting apophytis, running acrofs it, which is pro- longed outwards by a large tuberofity flattened from the top downwards, and on the fide of which the clavicle refl;s. This apophyfis is called the acromion : SYSTEM OP THE BONES. 35 Acromion: its upper edge is dlftinguifhed by the name o( fdper-acro??itan, and the lower by that of fub-acromlan. The external angle of the fcapula, which is truncated and fwelled up, prefents a facet, to re- ceive the head of the bone of the arm : above and within this facet is a very ftrong apophyiis bent forwards, and known by the name of the coracoid. The fcapula is fecured by mufcles. 33. The clavicle is a long, round bone, bent into the form of an J: it is articulated on one part into the top and fides of the fternum, and on the other with the fide of the acromion apophyfis. The acromian extremity is united to the acro- mion by an articular capfule, and two Tigammis (the rhomboid and conoid) which proceed from the coracoid apophyfis. The fternal extremity is fixed to the fternum and the cartilage of the firfl rib, by an articular capfule, a cojlo- clavicular and a flrong inter-clavi^ cular ligament. This articulation has alfo an inter-articular ligament. It is feen by this difpofition that the fhoulder is articulated with the trunk only by the extremity of the clavicle, which is united to the fternum ; and that the fcapnla can perform great movements by gliding on the thorax. 34. The arm. The arm, in all animals_, con- n z fills 3d ORGANIC STRUCTUHE* fifrs of only one bone, called the humerus^ thb bone varies with relped to its proportion ; it is ex- ceedingly fliort in cetaceous animals, and verj long in bats. 35. The HUMES.US. The iiumeras of maa has at its upper extremity a fmooth hemirphere, turned inwards and upwards *, by means of which it is articulated with the fcapula. Around this hemifphere is a flight depreffion^ which forms its neck ; and on the back part of it there are two tuh'ei'ofjks, one larger than the other ; the largeft is dillinguifhed by the name of the im" chitery and the fmaller by that of the t7-oc]mi-: they are feparated by a groove. The articular facet of the fcapula, which is narrow and almoU plane, is incrufled in a thick cartilage turned up at the edges, fo as to form a cavity proper for receiving the hemifpheric facet of the humerus : thefe bones are maintained in their proper portions by an ar- ticular eapfule, which embraces the neck of the humerus, from the ofleous and cartilaginous edge of the cavity of the fcapula. The arm can perform on the (houlder very free: movements, in every dirediioii. * In defcribing the bones of the thoracic Kmbs of man, it i» fuppofed, in regard to the refpedlive pofition of the different parts, that the limbs are extended along the body with the palm of the hand turned forwards. The SYSTEM OF THE BONES. QJ The humerus becomes broader at the lower part : it is plane behind, convex before, and turned a little outwa4-ds : it is terminated on the fides by two tuberofities : the interior, which is larger and does not defcend fo low, is called the epitjochlea ; the other is known by the flame of the eplcmdyJe, At the extremity of the bone there are two fmooth round furfaces, flightly hollowed into two grooves, Separated from each other by a projecting line : the interior moves around the bone, from the fore- part backwards, in the form of a pulley, trochlea ; and is terminated before and behind by a cavity i the exterior is placed at the lower part before, and is called the condyle. 36. The pore-arm. The fore-arm in fome animals, fuch as the bat, confifts of only one bone ; in ruminating animals it appears to be formed of two bones which have coalefced : in the greater part of aflimals, and in man, it confifts of two di- ftin^l bones, called the cubitus or uhia and the radius. 37. Cuhitiis. The bone of the elbow, which is flender and of a triangular fhape, becomes larger at the upper part, where it has a deep articular indentation of a femi- circular form, turned for- wards and inwards, called the Jygmoid cavity. The humeral extremity of this bone has on the back part an eminence^ which forms the elbow or ole^ D 3 cranon: 3S ORGANIC STBl'CTURE. cranon.' before, it has a larger one called the coro-' noid, and within a fmall articular y^<:^^ called the {m^WJygmoid cavity : this articulation is fecured by an miterior ligament, a fojierior ligament, two la- teral one^, and a capfule. The bone of the elbow is terminated below by a fmall plane articular facet, on the interior tide of which is an apophyfis called Xhejlyloid. 38. The radius. The radius at the upper end has a fmall articular facet, fomewhat concave, the fmooth internal edge of which is articulated with the fmall fygmoid cavity of the cubitus. Below the head it is much fmaller, fo that this part forms its neck. This bone becomes broader towards the lower end, and is terminated by an articular facet, lengthened in a crofs dire(51:ion, and fome- what concave. On the exterior fide of this bone is a Jlyloid apophyfis^ and on the interior a fmall articular facet, vvhich correfponds with that of the cuVitus. * The fore-arm is articulated on the arm by its two bones, which are articulated alio with each other above and below. The fygmoid cavity of the cubitus embraces the articular facet of the humerus, which is fhaped like a pulley, and gliding on it caufes the fore-arm to execute great movements on the arm as if by a hinge. The olecranon placing itfelf in the ca- vity which is behind the pulley of the humerus, limits SYSTEM OF THE BONES. 3Q limits the motion of extenfion ; and the coronoid eminence by refti^ng againfi: the cavity, which is tefbre this pulley^ fixes the movements of flexion. The articular facet of the radius reds againfi the rounded furface on the exterior fide of tlie pulley of the humerus, and glides on that furface, accompanying the cubitus in its motions. In its articulation with the humerus, the radius can turn on itfelf. This rotary motion takes place at the top againfi: th^ external edge of the cubitus, and at the inferior end around that bone. The bones which form the articulation of the cubitus are fecured by an articular capfulc, and two lateral ligaments. The interfiiice between the two bones of the fore-arm is filled up by a very flrong interojfeous ligament. The two bones of the fore-arm are kept together alfo above by an annular ligament, and below by a capfule or triangular ligament. 39. The laH part of the thoracic limb has re- ceived different names*, according to its par- ticular difpofition; but in all anim.als it confifis of the ^-^r^z^j-, the metacarpus^ and ihe phalanges. 40. Carpus. The carpus is compofed of fmall fhort bones of an irregular figure, the num- ber of which is variable. In birds it is formed of two bones in a row. The number of the bones of the * It has been called hand, paw, hoof, paficrn, biftard wing, fin, &c. p 4 carpus 40 ORGANIC STRUCTURE. carpus varies in the mammalia from 5 to 10, which are always difpofed in two rows. In oviparous quadrupeds, moft of the batrachians have the carpus formed of eight bones in three rows. 41. Metacarpus. The metacarpus conlifts of longifh bones, the number of which varies from I to 5 ; but in general it is equal to that of the fingers. In ruminating animals and the foIij)eda the meta- carpus confifts of a very long bone, which conlli- tutes the cannon bone. This bone feems to add to the Hmbs of thefe animals one part more, which in them is generally taken for the fore-leg. The metacarpus of birds is formed of a bone with two branches united. In cetaceous animals the bones of the metacarpus are united and flat- tened. 42. The phalanges. The phalanges are tliofe longifh bones which form the fingers. Sometimes they have the appearance only of imperfe6t rudiments ; the perfect phalanges form fingers, which remain concealed under the fkin^ or are covered by lax membranes, or are en- veloped in a .carneous matter, or form externally Separate and diftincSt fingers. Animals have nevef more than five fingers, each of which confifts, at moft, of three phalanges; and the thumb has never more than two. The folipeda have only one perfe(£l finger, which is known by the name of the paftern, coronet or coffin- SYSTEM OF THE BONES. 41- coffin-bone. Ruminating animals have two fin- gers, fupported by the bone of their metacarpus which forms the cannon bone. The three-toed lloth has three fingers. Several animals with folid hoofs have four fingers, as the fwine, the tapir, and the hippopotamos. In the laft place, five fin- gers are obferved in mofl carnivorous animals, in apes, and in man. In bats, the extremity of the laft phalanges, which are very long, terminates in points without claws*: it is fometimes enveloped in a corneous fubftance, as in ruminating animals and the folipeda : it ter- minates mfxed claws in dogs, and retra£lile claws in cats: in apes and in man it is enveloped by the fkin, and covered with a fiat nail. Birds have a large finger with two phalanges, and a pointed bone which fupplies the place of a thumb : thefe two fingers, as well as the bone of the metacarpus which fupports them, ferve for ^?earing feathers f . The thumb is formed of one or two phalanges; compared with the other fingers it is generally ihorter : in carnivorous animals it is of the fame length, and in the phocae longer : in apes, and in man, the tliumb is free, and can be oppofed to * Except the thumb, which remains fhort and nail-like. -|- The thumb bears the baftard feathers, and the large finger ai well as the metacarpus bears the primary feathers. the 42 ORGANIC STRUCTURE. the other fingers ; this difpolition forms the cha» radier of the hand. ^ Thefe different conftitutions of the extremity of the thoracic limb enable animals to employ it differently ; and their habits^ in this refpedi, are always the refult of the peculiarities obferved in the organization of this part. 43. The hand of man. In man the hand is compofed of the carpus^ the metacarpus, and the fingers. 44. The Carpus. The carpus of man is form- ed of eight bones'^, difpofed in two rows : the two of the firll row towards the thumb are re- ceived in the articular facet of the bone of the ra- dius ; the third is united to the bone of the cubi- tus; and the fourth rolls on the interior fide of the third. The bones of the fecond row are ar- ticulated above with thofe of the firf}-, viz. the two towards the thumb with that which is above ; the third with the fecond and third ; and the fourth with the third : all the bones of the carpus are articulated alfo with each other on the fides. The bones of the carpus being articulated on * Each of thefe bones Is uiftinguidied by ^ particular name. Thofe of the tiirt row counting from the thumb are; ift, os fcaphoidee ; zd^ os femi lunave ; 3d, os cuneiforme; 4th, os pififorme. Thofe of the fecond row, taken in the fame direc- tion, are ; ift, OS trapezium; 2d, os trapezoides j 36, os mag- num j 4thj OS unciforme. thofe SYSTEM OF THE BONES. 43 thofe of the fore-arm, muft follow the movements which the latter perfonn on each other. Thus the bone of the radius is articulated above with the humerus, and can turn on itfelf by gliding on the fide of the i}'gmoid cavity: below, this bone of the radius is articulated on the exterior fide of the fraall head of the os cubitus, around which it turns- by moving forwards, and the radius crofles before the bone of the cubitus, in the form of a laltire, defcribing below an arc, the fize of which. is determined by the breadth of the bone in that place. In this motion of the radius the carpus, which on the one fide is articulated with it by its firft two bones, and which on the other is united to the bone of the cubitus by its third, is carried forwards and inwards by the radius ; and in this movement it turns on the extremity of the os cubitus as on a pivot. It is thus that the move- ments o^ pronation and fupinaf ion are performed. The articulation of the three bones of the car- pus with the two bones o( the fore-arm is enve- loped by an articular capfule ; this articulation contains a triangular cartilage, and is fixed by an anterior ligament, a fojierior 3lU(\. two lateral I^ga^ ments. The fecond row of bones of the carpus is alfb fixed to thofe of the fecond by an articular cap- fule ; and the bones of the carpus are conneded 44 ORGANIC STRUCTURE, by ligamentous fibres exceeding!}! varied^ wbicb extend from the one to the other. 45, Metacarpus. The five bones of the me- tacarpus in inan are long, and fomewhat convex behind; on the upper extremity they have a facet for their articulation with the bones of the carpus j and on the fides there are other fmall facets for uniting them to each other : their inferior extre- mity is terminated by a round head^ which is re- ceived by the phalanges. The three firft bones of the metacarpus are ar- ticulated with the three hril of the fecond row of the carpus ; the laft two are articulated with the fourth. The articulation of the bones of the metacar- pus with the bones of the carpus is fecured by an articular capfule ; and by ligaments which proceed- ing from the fecond row of the bones of the carpus are fixed at the bafe of the bones of the metacarpus: thefe ligaments are numerous, exceedingly varied, and placed before, behind, and on the fides. 46. Fingers. The five' fingers of man are each formed of three phalanges, except the thumb^ which has only two. The firft phalanges are longer, and at the upper end have articular facets which receive the heads of the bones of the metacarpus ; and at the lower end round procefles which are received by the facets of the following phalanges (phahng'mes). In the SYSTEM OF THE BONES. 4$ the laft place, thofe which terminate the fingers (hhaJangettes) are very fmall ; they have alfo at the fuperior end articular facets, which receive the heads of the phalangines, and are terminated by a tubercle which fupports the nail. The metacarpal articulation of the thumb has always, towards its palmary face, two fmall round bones called ojfa fefijnoidea. Others are fome- times found towards the articulations of the other phalanges. The articulation of each phalanx is lecured by an articular capfule and lateral liga- ments. Independently of the movements which the hand is fufceptible of performing by following thofe of the radius, it can alfo bend forward's, backwards, and towards the tides. The bones of the carpus, as well as thofe of the metacarpus, perform on each other only very light and gentle movements; but the aggregate of thefe move- ments produces a very ftriking effe61:, and greatly ^ contributes to give the hand pliability and mobi- lity. The four fingers, being feparated and free, can perform feparate movements of flexion, in ths articulation of the phalanges, and lateral move- ments on the bones of the metacarpus. The two phalanges of the thumb, and tlie cor- refponding bone of the metacarpus, form a fepa- rate finger, v/hich -is articulated diredlly on the carpus ; 40, ORGANIC STRUCTURE. carpus ; moves freely in every direclion^ and cnti be oppofed to the other fingers. The thumb of man is freer, and proportionally larger, than that of apes; its metacarpal bones are alfo lefs concave than thofe ofthefe animals; fo that man, v»'ith rerpe6l to the bones of the hand, exhi- bits an arrangement more advantageous than any other animal. , 47. The thoracic limbs are inferior in flrength to the pelvian ; they are alfo fhorter : their great mobility on the fhoulder and with the fhoulder 1 their eafy flexibility in the articulation of the cu- bituSj but in particular that variety and pliability of motion of which the fore-arm and the hand are fufceptiblcj form one of the great advantages which man enjoys over oth<^r animals. Some relation feems to have been remarked between theincreafe of the thoracic limbs and the expanfion of the bones of the cranium. It has been obferved that children, in whom the bones of the cranium do not afTume their ufual develop- ment, becaufe the brain is in part wanting face- phali), have their arms exceedingly long, while thofe in whom the bones of the cranium, in con- fequence of water collected in the iniide of itj afliime a great expanfion, (hydrocc^halij) have the arms very fhort. It may be remarked alfo that, of all the jnanima- SYSTEM OF THE BONES, 47 //i7, man has the largeft head and the fhorteft limbs. This ilrudure would be very unfavour- able for walking on all four. Of the Pelvlan Lirnbs. 48. Pelvian limbs op animal?. All animals which have thoracic limbs are furnifhed alfo with pelvian limbs : we muft however except cetaceous animalsj in which thefe parts are wanting. The pelvian limbs, in general, have a flruc- tyre analogous to that of the thoracic limbs : thus a bone has a cavity proper for receiving the bone of the thigh, as the fcapula receives the humerus : the thigh is formed of only one bone, in the fame manner as the arm ; the leg like the fore- arm confifts of two bones ; and the foot, in the fame manner as the hand, is formed of three analo- gous bones. Nctv^ithilanding this great limila- rity between the thoracic and the pelvian limbs, each of their parts exhibits differences {o (Irik-? ing, that it is hardly poflible to confound them ; and therefore it is of importance that they fhould be examined feparately. 49. The hip. The upper part of the pel- vian limb is formed of a broad bone placed at the bottom, and on the fide of the vertebral co- lumn : it is called the coxal bone, and forms the hip. The hip bones are direded in- the portion of a curve 4a ORGANIC STRUCTURE. a curve towards the belly, and unite at a certam diilance from the vertebral column. The excavation formed by the two coxal boneSj and that part of the vertebral column to which they are attached, has bee ndiftinguifhed by the appellation ot pelvis. The hip bones are generally articulated with- out motion on the vertebral column : in tortoifeSj however, they are moveable. In birds the lum- bar vertebras, the os facrum, and the coxal bones form only one. Animals with pouches, fuch as the didelphk^ have before the pelvis (pubis), two articulated and moveable bones, called tha marjup'ial hones : to thefe bones are attached particular mufcles, which fupport a bag or pouch containing the teats. The coxal bones vary much in their form iii different animals; which gives to the pelvis various configurations^ and produces ftriking changes in the mode in which the thigh is articulated on thefe bones. 50. The hip in man. The hip bon^ in man, which is very irregular, forms chiefly the haunch : it is iDroad behind, and at the top ; but becomes narrower, excavated, and indented at the bottom and before. At an early age this bone confifls of three parts, which have been diflingulGied by names accoi'ding to their polition. The broad portion, 5 which SYSTEM OF THE BO^-ES. 4g which is behind and on the fides, is called the ilium ; that part below the firft is named the if- chium', and that entirely before is \hQ pubis. The iliac portion of the coxal bone exhibits be- hind and within a large articular furface, with which this bone is united to the facriim ; it then advances by a large plane portion which juts out at the top and on the fides ; the upper edge of this bone is broad and rounded ; it defcends be- hindj and at the end has a large tuberofity : be- fore, this edge terminates in a right angle, at the bottom of which is a tuberofity. The ijchlal portion is below and before the ilial ; in the middle and on the exterior fide it has a deep cavity called the cotyloid^ which receives the head of the thigh-bone ; this cavity, which turns outwards, downwards and a little forwards, is not completely formed by this part of the hip bone: the other two contribute alfo to its forma- tion. At the bottom of this cavity is a fmall fofi[a, and on its interior edge a deep indentation. The ifchial portion has behind two notches fe- parated by a tubercle ; it is terminated below by a large tuberofity, on which the human fig-ure refls when it fits ; in the infide and before it has a notch, which being united to another belonging to the pubis forms the /uh-fuhia?i hole, which in the male is oval, and in the female triangular. The pubic portion of the hip bone is entirely VOL. I. E before i 50 ORGANIC. STRUCTURE. before ; it is terminated by a narrow edge farnidied with a cartilage, by means of which this bone is applied to the cartilage of the correfponding part. This articulation is called the JymphyfiS of the piVis. Above their junction, thefe portions of bone have a prominence, aiid below an arch (of Xht pubis) ^ The fpace circumfcribed by the broad hollaw part of the ihum conftitutes the large pelvis ; the other fortned below by the ifehium., the pubis, the OS faqrum, and the coccyxj and which is nar- rower, conftitutes the lejfer pelvis: the name of ufper Jlralt has been given to the fomewhat nar- row edge which feparates thefe two cavities, and that of lo%vei:Jk^i4iJjQ_^Q fpace comprehended be- iweenytfre •uiberblitiesjtj\he ifchium, the arch of the cfu^is and the coccyx^ 'hefe^EHiisl'behk/d haye notches filled up in part W exceedii3^l3L-ftr0r)wligaments, which ferve ta fix tnbfeJboft^s th-fe^s facrum, and to the lafl of the lumbar vertebras : they are called the ileo^ lumbar^ the large awd fniall facro-fciatic\\ga.m&n\^j and ihefacro-iliac ligaments. The articular portions of the hip bones, which form the f}'mphylis of the pubis, are covered be- fore with a cartilage of greater or Icfs thickncfs; They are kept together before by a bunch of ligamentous fibres, and below by a triangular li- gament. SYSTEM OF THE BONES. 51 In confequence of the vertical pofition of man, the pelvian cavity is turned upwards ; and it can receive and fupport a part of his abdominal vifcera: this advantageous difpolition cannot take place in animals, and particularly quadrupeds, the aper- ture of whofe pelvis is turned dire6lly forwards ; but the region of the pubis, which in thefe is very narrow and forwards, is lower, and often very broad in man, and fupports a part of the vifcera. 51. The thigh. The thigh always conlifts of one bone, named the fe?nur ; which, in gene- ral, has nearly the fame fhape, but varies a great deal in its proportion. In ruminating animals and the folipedait is fo fhort, that the limb which it forms, being in part concealed by the fiefh, is not commonly taken for the thigh : this bone is ftill fhorter in the phoc£e. 52. Femur. In man, the thigh, the length of which determines that of the limb, is nearly cylindrical : it has, however, behind a rough line called the femoral, which indicates the place where the mufcles are attached, and is continued along two thirds of the bone at the upper part. This bone has at its fummit an articular head, which is turned inwaids, and a little upwards^ and which is fupported by a neck: this head has a fmall depreffion in the middle. The femiK- direcflly above, is terminated by a large tuberolity, E 2 - which 52 ORGANIC STRUCTURE* which projects a little outwards, and which is di- ftinguiflned by the name o{ trochanter. On its in- terior fide behind it has a cavity. Behind thefe parts, but lower down, and a little within, is an- other fmaller tubcrofity, called the trocbant'in. The femur is terminated below by two large articular facets, rounded from the tore part back- wards in the form of the portion of a wheel, and called condyles: of thefe condyles the exterior is fmaller, anil does not defcend fo low as the inte- rior : they are feparated by a groove, which ter- minates behind at a dcpreffion called the fojfa ^o^l'itea : from behind the condyles proceed two lines, which afcend obliquely, and unite at the fe- moral line : before thefe condyles the rotula is placed. The femur is articulated by its Bead with the cotyloid cavity of the hip bone, and can perform On it motions in every direclion. Thefe bones are retained in their pofition by a fi:rong articular capfule, and an inter-articular {round') ligament. 53. Leg. The leg fometimes is formed of only one bone, called the tibia, as in ruminating animals. In others, fuch as the dog, rat, fwine, and birds, it is compofed of the tibia and another bone or portion of a bone, which coalefces with it for a greater or lefs extent. In a great many of the mammalia the leg is formed of two diftindl and feparate bones, viz. the tibia, and another much SYSTEM OP THE EONES. 5g much flenderer called the perone or find a : thefe two bones are articulated below with thofe of the foot ; above, the tibia only is joined to the bone of the thigh, and the perone remains united on the fide of the tibia. In the laft place, in the greater part of oviparous quadrupeds, the two bones of the leg are articulated at the top with the bone of the thigh, and below wirh thofe of the foot. Before the articulation of the thigh with the leg, there is generally a fmall 'flat bone called the rotula. In man, the thigh is formed of the tibia and a dx^xm& perone, which is articulated above only with the tibia : the knee is furniOied alfo with a rotula. 54. Tibia. The tibia, throughout almoft its whole length, afFeds a triangular form : the an- terior edge (pratibial) is very falient ; it is termi- nated at the top by a tubercle, to which a ftrong ligament is attached. This portion of the tibia be- comes much larger in that part, and is diftin- guifhed by the name of the head : below this head is a plane furface, on which there are two articu- lar faces, flightly excavated, which receive the con- dyles of the femur, and are feparated from before backwards by a double elongated tubercle. The lides of the head of the tibia are difiinguiflied by the name of condyles. A little behind the head of the tibia, under the exterior condyle, is a fmall ftrticujar facet, which receives ihe perone, E 3 The 54 ORGANIC STRUCTURE. The tibia is fmaller below, and terminates in b concave articular furface of greater extent in a crofs dire6lion, and which refts on a bone of the foot. On the interior fide of this furface is a large tuberofity, prolonged below, and which forms the interior malleolus : its exterior fide, flightly notched, has an articular facet, which re- ".eives the perone. The poflerior face of the tibia J difiinguifhed by the name of the pophtean. ^^, Perofte. The perone, which is very flender,, is fituated at the exterior fide of the tibia, and a little behind it. At the upper extremity it has a head, on the fide of which is an articular facet refling agalnfi that bone. At the lower end the perone defcends below the tibia: it makes a pro- je(9ion outwards, which forms the exterior malleo- lus, and on the infide has an articular facet for its jun6lion with the tibia, and the bone of the foot which receives it, ^6. Rohda. The rotula is a fmall flat round bone, having behind it a double facet, which cor- refponds before the femur. The articulation of the knee is fecured by an articular caffuU and two later al\\^2im^Vi\'&\ before by a firong ligament, which belongs to the rotula, and behind by two crofs ligaments: this articula- tion has alfo two inter-articular cartilages of 2ifemi~ lunar form. The SYSTEM OP THE BONES. 55 The perone is fixed to the tibia by an articular capfule, and an anterior and pofterior ligament. The fpace left between the two bones of the leg, which is larger above, is filled by a ligament attached on the edges of thefe bones. The leg can perform only very extenfive move- ments of ilexion backwards. 57. Last part of the pelvian limb. The lafl: part of the pelvian limb, like that of the thoracic limb, has received different names ac- cording to its mode of termination ; but it is alwavs formed of a tarfus^ a inetatarfus, and toes. The bones of the tarfus vary very much both with refpe61: to their form and number. The tar- fus in birds confifts of only one bone ; in the three-toed floth it confifts of three; in ruminating animals, of from five to fix, and very often of feven. The bones of the tarfus are in general thick and (hort : in fome animals, however, they are very long. The bones of the leg are articulated, in general, with only one bone of the tarfus ; but in reptiles they reft on two bones of that part *. The bones of the metatarfus, in animals, have the fame o-eneral arrangement as thofe of the meta- carpus ; and the toes, which are articulated to one or the other of thefe parts, exhibit alfo the fame mode of termination. * The tibia refts on the aftragalus, and the perone on the cal- caneum. £ 4 The 56 ORGANIC STRUCTURE. The pelvian limb has only five toes * ; each toe confifts at moll of four phalanges j-, and the large toe has only two ;|:, The folipeda have only one toe on the hind-foot, and one finger to the fore- foot. Ruminating animals have two on each foot, articu* lated on their cannon bone. Some animals, fuch as the tapir and the rhinoceros, have three toes, and the fwine has four. The greater part have five. The frog has fix. The toes are often pa- rallel, as in carnivorous animals: in the quadru- mana and pedimana the great toe is a thumbs which can be*Oppofed to the other toes. In birds which have three toes they are parallel : thofe fiir- nifbed with four have one or two turned back- wards, or all the four turned forwards. The number of the phalanges in thefe animals is in- creafed by one for each toe, proceeding from the great toe. The large toe is fhorter than the reft in the quadrumana and pedimana, all the toes of which are very long : in fome of the carnivorous animals it is nearly equal to the refi : in man it is the largcfl of all the toes, 58. Foot of man. The foot of man is formed of the tarfus, the metatarfus, and the toes. Tarfus. The tarfus of man confifis of {QVtn. bones. That on whiph the two bones of the leg * The frog has fix. •f" In the toe before the lafli of the lizard there are five, J In that of the chameleon there are three. SYSTEM OP THE BONES, 57 Sfe articulated is called the afiragahis \ it is placed on another larger bone which forms the heel, and which is called the calcaneiun z a third, fituated before the afiragahs, is diftinguilhed by the name of the fcaphoid bone (cs fcaphouies) ox navicular bone (os naviculai-e). The other four, placed in a row, receive the bones of the mctatarfus : the firft three, counting from the great toe. arc knowa by the name of the cuneiform bones, and ihe fourth by that of the cuboid. 59. j4firagalus. The aflragalus has at the top an articular facet, rounded from before backwards, in the form of the portion of a pulley ; it is re- ceived by the articular cavity of the tibia. On the fides of this facet, the afiragalus receives with- out, the extremity of the pcrone, which forms the exterior malleolus, and within the tuberoiity of the tibia, which produces the interior mal- leolus. The aftra2;alus has alfo articular facets for its union below with the calcaneum, and before with the fcaphoid and even the cuboid bones. 60. Calcaneum. The calcaneum, which is larger than the preceding bone, is lengthened from the fore part backv.'ards ; and has above and before a deep indentation or notch which receives the af- tragalus, and diredly before another facet for its junclion with the cuboid bone. This bone is lengthened behind by a large tuberofity, which forius 58 ORGANIC STUUCTURE. - forms the projedion of the heel, and below it there are two tubercles. 6i. Scaphoid hone. The fcaphoid bone is arti- culated behind with the aftragalus ; on the out- lide with the cuboid bone, and before with the three cuneiform bones. 62. Cuhoid bone. The cuboid is united behind to the calcaneum and the afiragalus; in the in- lide to the fcaphoid bone and the third cuneiform bone; before it is articulated with the two 1 aft bones of the metatarfus. 63. Cuneiform hones. The three cuneiform bones are articulated behind with the fcaphoid bone ; on the outfide with the cuboid, and before with the four firft of the metatarfus ; fo that the firft cuneiform bone touches the two iirfl; bones of the metatarfus ; the fecond touches only the fe- cond J the third touches the fecond, the third, and the fourth. Thefe bones are articulated with each other on the fides. It is to be obferved that the three cuneiform bones have really the form of a wedge ; that their difpofition with the other bones of the tarfus is fuch, that this part forms a move- able arch marked by the inftep, and that, in pro- greffion, all the bones of the tarfus do not reft on the ground. 64. The articulation of the \tg with the foot is Secured by a capfule, an interior lateral ligament, and three exterior lateral ligaments,. The SYSTEM OP THE BONES. 5Q The five bones of the metatarfLis, which are nearJy of the fame length, have behind articular facets for their union with the mhoidRnd the three cuneiform bones; and before fmall heads, which are received bv the excavated facets of the firft pha- langes : thefe bones are articulated alfo with each other on the fides by their pofterior extremity. The articabtion of the bones of the tarfus with thofe of the metatarfus, and that of all thefe bones with each other, is fecured by ligaments analogous to thofe which fix the bones of the carpus and metacarpus i except the firft bone of the metatar- fi.is, which is not feparated like the firft of the me- tacarpus. The phalanges of the toes have the fame mode of articulation and the fame means of junction as thofe of the fingers, and are equal to them in number. The five toes are parallel ; and the ' large toe is longer than the reft, which go on decreafing in fizc to the laft. 6^. The ftrucfture and arrangement of the bones of the foot are fuch, that they conftitute an arch more elevated and more extenfive in the in- fide than without: the foot refts behind on the ex- tremity of the calcaneum, which is dire6led a little outwards; it refts before on the anterior extremity of the bones of the metatarfus, and on the toes ; and in the middle of its exterior edge the foot refts on go ORGA^^IC STRUCTURE. on the pofterior head of the laft bone of the meta- tarfus. It Is to be remarked, that the aftragalus forms in the infide the fummit of the arch, and that the leg, fupported by this bone, correfponds with this arch in the place where the bones by which it is formed do not touch the ground. 66. Befides the bones already mentioned, there are always two fmall round bones below the meta- tarfian articulation of the large toe : there are fmall bones alfo fometimes at the articulations of the phalanges : thefe bones are called ihQfefamoid (ojjli fefamoidea). 67. The bones of the limbs have alfo fome pe- culiarities common to the bones of the trunk and the head : thus, for example, they are all covered with a periofteum ; frequent afperities, which mark the places where the mufcles adhered, are alfo ob- ferved in them, and they have holes by which they are pierced throughout their whole thicknefs. The long bones have alfo in the infide a cavity, larger in the middle, which goes on increafing to the ex- tremities, where it terminates in offeous fibres that? crofs each other in different dire6lions, and form a kind of tiet-ivork. The cavities of the long bones are lined alfo with an interior periofteum. 68. The flvcleton, examined in general, e^^ 6 . hibits SYSl'F.M OF THE BONES. 6l hiblts an aflemblage of pieces equally pliable and ilrong. The vertebral column confifls of 24 very fliort bones, light, exceedingly ftrong, articulated in fe- veral points, and their articular furfaces are covered with a thick cartilage highly elallic^ and fufcepti- ble of yielding to every purpofe. All thefe bones are retained in their portion by ligaments fiiffi- ciently ftrong and numerous to give that column the requifite degree of tlrength. The aggregate of the vertebrae afFedis different curves, which can be increafed or diminifhed according to circum- llances. The head refts on the vertebral column almofl by its centre ; and the two firfi: vertebrse have a pe- culiar difpofition, which allows the greateft move- ments. The difpofition of the bones of the cranium, in the form of an arch, is exceedingly well calculated to (ecu re the brain from the a61ion of all external bodies ; and the vertebral canal protedls the pro- longation of the brain which it receives. The vertebral column fcrves alfo to fupport the ribs, the aggregate of which, together with the flernum, forms a conical cavity ftrong and exceed- ingly flexible, which envelops the heart and the lungs. The vertebral column rcfts on the os facrum : this bone with the two hip bones forms a cavity, the 52- OUSANIC STRUCTURg* the different pieces of which, united ftrongly by cartilages * and ligaments, are however fufceptible, when neceflary, of very linking movements. The cotyloid cavity is fituated in an oblique diredlion on the exterior fide of the hip bones : the femur is bent forwards j its head and neck have an oblique dire(5lion, and its interior condyle is longer : the knee is flightly inclined inwards, and the thigh is always ready to bend upon the leg : the tibia is fomewhat curved, and the foot inclined outwards, and forms an arch. This general conflru6tion, which every where exhibits curves, angles, and folding levers, unites agility to ftrength, and feems to be well adapted for performing, without any fhock, thofe move- ments communicated to the trunk by fiamping the feet againft the ground. 69. When a man walks fail, flamps with his foot, or falls in an upright pofiure, there is pro- duced, in this adlion, a quantity of motion which mufi: neceflanly be employed, and produce an effect ; and if by habit this efFecl occaiions no dif- order, the motion muft be diftributed by a peculiar difpofition to every part, without producing any fliock. When the foot indeed flrikes the ground, the * Thefe cartilages fwell up during the laft months of preg- nancy; which gives more amplitude to the pelvis, and difpofes the bones of which it is formed to move with greater facility. 7 motioiv SYSTEM OF THE BONES. ^ 53 motion produced is communicated to the places which begin to bear : thus the heel, the anterior extremity of the bones of the metatarfas, and the exterior edge of the foot, receive the firfl impref- lion. The arch formed by the foot on the interior iide gradually links down ; the inter-articular car- tilages yield, and the motion then propagated along the tibia immediately increafes its curva- ture. The motion, when it reaches the knee, is not communicated in a ftraight line to the thigh : it divides itfelf on the fides, pafles from the condyles of the tibia to thofe of the femur, and the femilu- nar cartilages, interpofed between thefe parts, im- mediately yield : on the other hand, the direc- tion of the knee inwards and fomewhat forwards dilpofes it to give way, and it then bends. The motion being then continued along the femur immediately increafes its curvature, and when it reaches its neck forces it to bend. The bead of the femur does not bear diredlly under the pelvis, but reds obliquely in the cotyloid ca- vity; fo that it acls upwards and backwards on the edge of that cavity, compreffing the articular cartilages ; and at that moment the pelvis bends on the thigh as the thigh bent on the leg. It is to be obfervcd, tltat, in this aiiion of the parts, the articular capfules, the ligaments and the mufcles. 64 ORGANIC STRUCTURE;. mufcleSj which farround the articulations, eafily give way, and recover their former ftate. The motion communicated to the bottom of the cotyloid cavities tends to carry the hip bones upwards and inwards. To obey this impreiiion, the curvature of thefe bones decreafes momen- taneoufly ; they move before in the articulation of the f}^mphyfis, and behind they glide on the fa- crum. This bone, fliaped like a wedge, pene- trates between the hip bones, tends to feparate them, and oppofes the firft impulfe that tended to bring them together : in this adtion the inter-arti- cular cartilages and the ligaments yield more or lefs. In the laft place, if any motion remains, it is communicated obliquely to the vertebral column by the top of the facrum, and continuing along that part it increafes its different curvatures; de- pretles its inter-articular cartilages ; and caufes the vertebrce to glide on their different furfaces. The motion, when it reaches the cervical region, communicates itfelf to the head, which is there fufpended, and as it were in equilibrium ; and as it has a tendency to fall forwards it bends on the neck. From this arrangement, it may be feen that the fum of the'motion communicated to the feet mufl be very great before there can reach the vertebral column SYSTEM OF THE BONES. 65 eolamn and the head a quantity capable of pro- ducing derangement in the parts it contains; which indeed happens Very rarely, 70. Female Jkeleton. The female Ikeleton is in general fhorter than the male ; but the trunk is comparatively lengthened, fo that the half of the body, which in man correfponds to the pubis, is above that part in the female. The bones which compofe the female Ikeleton are flenderer, weaker, fmoother, rounder and whiter : the eminences, afperities, cavities, and holes_, are neither fo ftrong nor fo ftriking. The female fkeleton, compared to that of the male, exhibits alfo very apparent local differences. The vertebral column is comparatively longer, find the capacity of the breaft fhorter i fo that the lumbar region is the moll extenfiVe. The greater elongation of the trunk feems to be connected with* a greater thicknefs of the bodies of the lum- bar vertebrae, and particularly of their inter-articu- lar cartilages. The alternate curvatures of the vertebral column feem a little lefs ftriking in the female ; but the whole trunk affedls a fort of obliquity ; the facral region is thrown backwards ; the bread is carried more forwards, and the head requires to be kept more ere6l. This difpofitlon, which is owing to the greater capacity of the pelvis, and the more VOL. I, F forward 66 ORGANIC STRUCTURE. forward pofition of the cotyloid cavities, is ftili increafed in the fiat'e of pregnancy, when the projeclion of the abdomen forces the breafl and. head to be thrown backwards, in order that the equihbrium may be preferved. The tranfverfe apophyfes of the dorfal vertebrae are thrown more backwards; the pofterior arch of the ribs is more convex, and the vertebral grooves are more excavated. The ribs are flenderer as well as fmoother ; their convexity behind is much greater, and the cartilages of the falfe ribs are more elongated. Of the two pieces which compofe the fternmn, the fuperior is generally thicker and longer. The inferior is much fhorter in the female, but in fuch a proportion that the whole fternum is fhorter : the point of it is thrown a little forwards, while in man it is turned direclly down- wards. In confeqaence of the peculiar ftruclure of the dorfal vertebrae, of the ribs and the fternum, the thorax, in the female, has lefs tranfverfe breadth ; is lefs flattened before, and is more convex be- hind ; it is alfo wider below, and more rounded in its circumference. The clavicle, in the female, is longer, and al- moft ftraight, fo that the flTOulders are thrown more backwards, and project lefs on the fides of the thorax. To this peculiar difpofition of the fcapula^ SYSTEM OF THE BONES, Qy) fcapula, thrown farther back, is afcrlbed the dif- ;' ficulty which women experience in 'performing i great movements with the arms, and in projedl- ing bodies, which they cannot do with the fame '- force and eafe as men. The head, in the fctnale, is fmoother and more ' rounded; and the face i'^.fhorter. The pelvis', in general, is broader and (halloxver in the female ;- the iliac portions of the hip bones are thrown more back; the pubis is tranfverfely broader, arid has lefs height : the pubian arch is wider, and the fab pubian hole is tri.mgular. The ' aperture of the pelvis is turned more upwards j the diftance of the tuberofities of the ifchium is great- * er, and the curvature of the os facrum is more ap- parent. The cotyloid cavities are more forward^ lefs oblique, and of lefs depth. The neck of the femur has a more horizontal dire6lion ; the interior condyle of that bone is : larger, more convex, and dcfcends louver. It follows from the peculiar conformation of the pelvis, and that of the femur in women, that the haunches are broader; that the facral region pro- jecSts more pofteriorly ; which determines the in- clination of the trunk, an'd the projection forwards of the thorax. The cotyloid cavities are at a , greater diftance, and the knees are larger, and ap- - F 2 preach 08 ORGANIC STRUCTURE. proach each other more j which forces the feet i(3 turn outwards. This peculiar difpofition determinev'3 the grand lateral movements of the pelvis^ which women perform in walking to tranfport alternately the weight of the body to the two femora, and thus prcferve the centre of gravity. Dupuytren has obfervcd, that till the age ol* puberty the female pelvis diifers very little from that of the male; that both have the fame trian- gular form, and the antero-pofterior diameters nearly equal : but at that period the female pelvis acquires a rapid expanfion ; it becomes oval ; its tranfverfe diameter has lefs extent, and it foon aflumes all its diilinguifhing chara6ters. The general difpofition of the bread and pelvis, in the male and female, is fuch, that if two paral- lel lines be continued upwards and downwards, from the fides of the thorax, it is obferved in the male that a part of the fhoulders is cut. by thefe lines ; while in the female thefe lines interfedl a part of the bones of the pelvis. The breaft in the male is in fome meafure fquare ; in the female it is conical. 71. During the firft: days of conception, the germ is a gelatino-albuminous fluid, in the mid- dle of which fibres or filaments gradually appear. This SYSTEM OF THE BONES. 'g'^ This ftkte continues till the 20th day, at which period there is developed in different points of the embryo a concrete, white, elaftic fubflance, called cartilage ; and towards the 30th or 40th day the foetus begins to fexhibit in various parts of the carti- lages fome fpecks of more confidence, which are the firft development of the bones. : It is remarked that orie cartilage only is formed for the whole of the bones that are to rfem^in united ; fuch as the cranium, the vertebral co- lumn, and the pelvis; and that a partidiilar one is formed for each of the bones which are to remain fepafate alnd moveable. Thfe part which at that period forms the cavity of the cranium is a fort of membrane. The bones continue to expand during the t?m6 of geftation : the firft points of offification' appear in the bones of the fhoulder and thofe of the cra- nium ; they next appear in the ribs, the bones of the arm and fore-arm; then in the vertebrae and the pelvis : in the laft place, they are fecn in the bones of the thigh and leg, in thofe of the meta^ carpus, the metatarfus, the fingers and the toes. The bones of the carpus, thofe of the tarfus, the calcaneum excepted, and the rotula, remain cartilaginous till after birth. The long bones explmd by three points of ofll-. fication ; one for the centre, and two for the ex- tremities : the large flat bones have their point of F 3 offificatioa 70 .-.-QRCANIC STRUCTURE. oflification in lh,e cenlre, : This poinf furnifhes longi.tudinal or radiated; fibres, which advance, raeel, and crofs each other to form one iblid piece. _ , ■;; ; ' . , ,. . At th;^ period of Wrth the fl^eleton of the foetus jexh.ibits- my rernarkable ■ pece1iaritit;s • the/ whole of it is fti!!, in a grca,^jn:veafu;r?, cartilaginons.' .Tlie ver^earal 0011101 n is quite ftraight, and; its p,ppphyrx reticular \A^K\^ \ in a' word, they compo(e fblid* bodies more or lefs elafiic, which feem to conlifl of laminae laid above each other. 74. If a bone be placed in an acid, fuch as the muriatic, diluted with water, it becomes foft, and F 4 lofes '^'l OaGANIC STRUCTURE. Jofes about the half of its weight. Carbonic aci4 gas is difengaged from the liquor, and there is depoiited at the bottom a fediment, the weight of which is equal to that lotl: by the bone : this de- poiit is muriate and phofphate of lime : what re- mains of the bone exhibits a fibrous, vafcular and gelatinous charadler. If a bone be placed in a folution of cauilic al-s kali (potafh or foda), it lofes half its weight in the fame manner; it remains folid, but becomes brit- tle and friable: the refiduum is found to be a fort of foapj and the earthy part is phofphate and carbonate of lime. In the laft place, if a bone be calcined in an operi fire, it flill lofes half its weight, and what remains is phofphate and carbonate of lime. If it be required to afcertain, by a morp accu- rate analyfis, the quantity of folid matters which enter into the compoiition of bones, the following method may be employed : calcine the bones till they are white, and, having pulverized them, fuh- je6l them to the adtion of the nitric acid. This acid will diflblve the lime and the phofphate of lime : then pour ammonia into the folution, and the phofphate of lime will be precipitated : fepa- rate this fait by filtration, and weigh it after it has been thoroughly dried. The filtrated liquor ilill contains lime, which muft be precipitated by «3ommon potaOi : this lime is united tq carbonic 6 acidj TABLE OF THE BONES WHICH FORM THE SKELETON. ( 1st. OF THE TRUNK, wAicA comprehends^ 2d. or THE HEAD, ivhicA comprehends. 5d. OF THE LIMBS 1st. THE VERTEBRAL COLUMN, formed by 2d. THE THORAX, foryntd by 1st. THE CRANIUM, formed by 2d. THE FACE, t formed by ^ ■•{ 1st. THE THORACIC, formed by 2d. THE PELVIAN, formed by VOL. I. — To front page 72. 24 VERTEBRA, VIZ. 7 ccrvical (the 1st Atloid, the 2d Axoid), 12 dorsal, 5 lumbar, the sacrum and the coccyx. 12 RIBS on each side (7 sternal, 5 asternal), the sternum and the 12 dorsal vertebras. The sphenoid, the frotital, the parietals, the occipital, the temporals, and the ethmoid. The frontal, the sphenoid, the bones of the nose, the lachrymals, the zygomas, the super-maxillary, the pala- tine bones, the ethmoid, and its turbinated bones, the maxillary j 32 teeth, 8 of which are incisors, 4 angular, and 20 molar. 1st. THE SHOULDER formed by 2d. THE ARM, composed of Sd. THE FORE-ARM composed of ■I 4th. THE HAND, formed by 1st. THE HAUNCH, composed of 2d. THE THIGH, composed of 3d. THE LEG, formed by 4th. THE FOOT, formed by ■{ } } { 1st. The clavicle ; 2d. The scapula. The humerus. 1st. The cubitus. 2d. The radius. 1st. THE CARPUS, composed of 2d. THE METACARPUS, composed of Sd. THE FINGERS, each composed of The hip-bone. The Femur. 1st. The Tibia. 2d. The Perone. 3d. The Rotula. 1st. THE TARSUS, composed of ' 8 bones in two rows, which are counted from th«. thumb, and which are called : , X Ti (1st. The scaphoid, 2d. the similunar, 3d. 1st Row. { , ./ , , \ i the cuneiform, 4th. the pisiform. 2d Row ^ ^^^' ^^^^ trapezium, 2d. the trapezoid, Sd, i OS magnum, 4th. the unciform. 5 bones of the metacarpus. 3 phalanges (2 in the thumb), which are counted from the metacarpus ; and which are called Phalanges, Phalangines, Phalangettes. The two hip-bones, with the sacrum and coccyx, form the pelvis. 1 f 7 bones : the astragulus, the calcaneum, the scaphoid, i^ the 3 cuneiform bones, and the cuboid, 2d. THE METATARSUS, C , ^ < 5 bones of the metatarsus. composed oj i 3d. THE TOES, r 3 phalanges (2 in the great toe), vf\i\ch. ^ve counted each composed of ■< from the metacarpus, and which are called Pha- (^ langes, Phalangines, Phalangettes. [ 74 ] MUSCULAR SYSTEM. 76. Our organs are moved by means of mufcles. The greater part of the carneous mafs, the mea- gre and fibrous portion of the fltfli of animals, conflitutes the mufcles. They are compofed of fibres in bundles. The fibres of the mufcles are fubdivided in an Indefinite manner. ' Each fibre, and each bundle of fibreSj is enveloped by a cellular m.embranej which is a tiffue of veflels. Each mufcle alfo is furrounded by a cellular membrane of greater thicknefs. The mufcles are continued, and generally termi- nate in a fmooth, white, fhining fubfiance, of a very compa6l fibrous texture^ round or flat, and always of lefs fize than the body of the mufcle. That portion of a mufcle which terminates in a cord is called a tendon-, and that which fpreads itfelf into a broad flat furface is diftingulfhed by the name of aponeurofts. It is generally by thefe tendinous or aponeurotic parts that the mufcles are fixed to the bones. The gelatinous, albuminous, and fat fubfiance, contained in the cells o{ this fi/fue, produces fullnefs of the mufcles. • The cellular tifiue, interpofed between the fibres of MUSCULAR SYSTEM. 75 of the miifcles, varies both in regard to its nature and quantity in different animals, in their diffe- rent parts, and according to the different circum- liances of health or difeafe. In the mammalia the mufclesareof a darker or lighter red colour; in the gallinaceous they are paler, and in reptiles and fifhes almoft white. The mufcular fibres have the property of con- tradling, of becnming Ihorter, of carrying with them the moveable points to which they are fixed, and of producing, in this manner, various move- ments. The mufcles of reptiles and of fifhes, which have white fibres, contrac!:! with much greater force and velocity than thofe with red fibres of other animals, Mufcles are never found but in parts which perform very flriking movements ; and they are ftronger and of greater length, according as thefe ipovements are greater and more energeiic. The movements performed by our different parts are always a refult of the form of the bones, of the peculiar difpofition of their articular furfaces, of the different points by which the mufcles are afiixed to thefe bones, and of the dire(Sfion of the mufcular fibres. Conk-quently a knowledge of the ratio of the articular furtaces of the bones of any part, and of the nature of the motions which it performs, may always lerve to deter anne, in a general 76 ORGANIC STRUCTURE. general manner, the fize, form, and dired^ioij-'of" the mufcles of that part. But as the mufcles do not alvvaj's prodace their a6iion in a ftraight line; as they often a6l by points of reflection ; to have a fufficiently corre«5l idea of them, it is neceffary to know minutely their pecuhar difpofition in man. 77. In the different animals the pofition of the mufcles, which are fixed to the vertebral column, varies according to the motions which are per- formed by that part. Thus the mufcles are op- pofite to the fpinal and praefpinal face of the ver- tebraSj when the vertebral column bends forwards or is thrown back, as in the mammalia, birds, and reptiles. In animals which perform only lateral movements with the fpine, fuch as fiflies, thefe mufcles are fituated on the fides of the rachis or fpine. In animals which make the vertebral co- lumn, or one of its parts, move in all directions, as is the cafe with the bodies of ferpents, the neck and tail of many of the mammalia, mufcles are found on the four faces of that part. In thofe parts of the vertebral column which perform no motion, mufcles are wanting : this is the cafe with the fpinal face of filhes, the dorfal region of birds, sr.d the praefpinal face of the dorfal and lumbar vertebrce of moft animals. 78. Muscles in man. In man, the mufcles MUSCULAR system;, 77 affixed to the vertebral column, in order to make it move, are placed chiefly on the fpinal face. 79. Mufcles which produce motion in the verte" hral column. A very thick mufcle, which extends from the os facrum to the atloid vertebra, runs along the fpinal face of the rachis. This raufcle is tingle inferiorly, and is fixed to the polterior part of the os facrum, and the edges of the os ileum. When it reaches the lumbar region, it divides itfelf into three parts : the firft is inferted in the tranfverfe apophyfes of the lumbar Vertebra?, at the angle of the ribs, and in the tranfverfe apo- phyfes of the laft cervical vertebras : the dorso— TRACHELIAN ov facro-lumhar portion ; the fecond part adheres to the tranfverfe apophyfes of the lumbar and dorfal vertebrae, as well as to the in- ferior edge of the laft eight ribs : the costo- TRACHELrAN or long dorfal fart ; this portion comprehends another, which extends from the tranfverfe apophyfes of the laft five vertebrse oi the neck to the firil four or five of the back : accejfory of the long dorfal or great tranfverfe muf cle of the neck; the third portion is inferted in the tranfverfe articular and fpinal apophyfes of all the vertebras : the lumbo-cervical portion of tranfverfe fpinal *. * The particular name of axoido-atlo'tdian haf5 been given to that which proceeds from the tranfverfe apophyfis of the axoid to 'tbc fpinal apophyfis of the atloid. » This 7S ORGANIC STRUCTURE, " This mufciC, in contracting, bends the vertebral column backwards, or keeps it in a ftate of recti- tude : THE SACK.OSPINIAN*. So. The fpaces comprehended between the fpi- nal apophyfes of the vertebrae of the neck are fill- ed by miifcles which are fixed to thefe apophifes. They keep the neck ftraight, and draw it back- wards. Interspinian of the NiiCK, fniall fpi- nal of the ?ieck. 8 1. In the cervical and Inmbar recrions, the mufclcs are fixed from one tranfverfe apophylis to the other. When thofe on one fide a6t by them- felves, they bend the vertebral column towards the fame fide ; w4icn they ad fimullaneoufiy they keep it in a fi:ate ofre6fi[ude. Inter-transver- SIAN OF THE NECK AND LOINS. '■ 82. Amafcle which proceeds before the bodies of the firfi: three dorfal veftebrse, along the cervi- cal vertebrae, is inferted in the anterior tubercle of * In defcrlbing the mufcles we fliall retain the nomenclature of Chaufller. This methodical nomenclature of the mufcles is founded on the points to which they are attached. 1 hus the name of every muCcle is compofed in generdl of two words, which indicate the'e two piints : the firft indicates the point which is the mult fixed, and the fecond that which is moft mpveable ; fo that the name is a fort of concife defcription. We Ihail adopt alfo, for the names of the muflles, the unitorm ter- mination in ien^ which Diimeril has given to them. [The Engliih tranOator has changed thefe terramalionb into ian.'\ the MUSCULAR SYSTEM. 79 the atlas.; it . bends the neck forwards. Pr^- DORSO-ATLOiDiAN ; Jofig jimfch of the necTt» 83. To the body of the laft vertebra of the neck is fixed a thin mufcle which is frequently want- ing. This mufcle defcends in an oblique direc- tion on the fide of the lumbar vertebrae, paiTes be- fore the ilium, and is inferted by a flat tendon in the ilio-pubic eminence. It tends to bend the fpineon the pelvis. PuBio-PRiELUMBJAN, ihefmall pfoas, 84. From above the upper and pofierior edge of the ilium rifes a mulcle, which is fixed to the tranfverfe apophyfes of the firfi: four vertebrae of the loins, and to the lafi: rib. It ferves to bend, the vertebral column on the pelvis. Ilio-trans- .VERSIAW, fquare of the loins. 85. Two fmall mufcles extend from the fpine of the ifchium to the fides of the coccyx : thefe mufcles tend to carry that fmall bone backwards. Ischio-coccygian, idem. 86. M'lfdes which give motion to the hreajt. The breafi: is moved by means of numerous muf- cles fixed between the ribs, or which extend fi'om the ribs to the vertebrae ; from the ribs to the Her- num, and even from the ribs and the fternum to the bones of the pelvis. The mufcles which extend from the breafi to the pelvis form in a great part the fides of the lower belly. In $0 ORGANIC STRUCTUlRf. In Ibme clafTes of animals the breafl and tovnet belly form only one cavity, as is the cafe in birds i Itnt in the greater number, thefe two cavities are fe- paratedby a broad, thin mufcle called iht diaphragm. In the different animals^ all thefe mufcles ex- perience variations analogous to the different con^ figurations of the fkeleton ; they are in a great meafure wanting in thofe which have no breaft properly called. 87. hi 7nan, the interval between the ribs Is filled up by two layers of mufcular fibres, which adhere to the edges of the ribs, and are dire6led obliquely in a contrary diredlion. The inter- COSTiAN, inter'' cojiajy exterior and interior.. 88. To the tranfverfe apophyfes of the laft five Tertebrss of the neck is fixed a mufcle, which is inferted in the middle pofierior part of the three firft ribs, by as many digitations. The tra- CB.'Ei.o-cosT I Ai^jfcalene. 89. Twelve mufcles proceed from the tranf- verfe apophyfes of the laft vertebra of the neck, and from the iirft eleven of the back, to the angle of the ribs. Transverso-costian, elevators of the rihs. 90. A mufcle extends from the fpinal apophyfes of the lafl two vertebra? of the neck, and the firft two of the back, to the angle of the firft five ribs (the firft excepted). Dorso-costian, fmall in- dented pofteiior, fuperior. All thefe mufcles tend, in an elTential manner, to elevate the ribs. 91. To MUSCULAR SYSTEM.' 81 91. To the fpinal apopbyfes of the laft two ver- tebras of the back, and the firft three of the loins, is fixed a mufcle which extends to the lafl four ribs, and tends to deprefs them. LjUMEo-costian, fmail indented pojierior inferior. 92, A mufcle of a triangulsr form proceeds from the interior furface of the fternum, and ex- tends to the lafl: five fl:ernal ribs : it contributes alfo to deprefs the ribs. Sterno-costian, /ri- angidar cf the Jiemum. g2' The thorax is feparated from the abdo- men by a large odd mufcle, which forms its dia" phragm. This broad, thin mufcle, tendinous in the mid- dle and flefhy on the edges, is fixed to the lower part of the fternum, the cartilaginous contour of the ribs, the tranfverfe apophyfes of the lafl: verte- bra of the back, the firfl; of the loins ; and to the lateral parts of the bodies of the firfl: two or tliree lumbar vertebrae, by two flefhy pillars. This muf- cle, convex on the fide towards the breaft, is pierced with three holes : the one on the right af^ fords a paflage to a large vein (vena cava); the other enters the pillars, and gives paflage to a large artery {^the ventral aorta)^ to ihe canal of the chy'e and lymph, and to the azygos vein ; the third receives the alimentary canal and a nerve (eighth pair). It retains the name of diaphragm. This mufcle^ becoming flat, by its continued and fucceilive con- voL. I. G tradions. 82 ORGANIC STRUCTURE, traftions, increafes the capacity of the thorax, and thus contributes to infpiration. The mufcles here defcribed produce the move-' ments performed by the thorax during refpiratioD. 94.. From the ilium and the pubis proceeds a large mufcle which rifes in an oblique dircciionj and is fixed to the lad eight ribs by as many digita- tions. This mufcle unites and is confounded with that of the oppofite lide by a very flrong aponeu- rotic expaniion, which forms longitudinally on the middle of the abdomen a line called the median. Thcfe mufcles in contrac^insr tend to lower the O breaft, and to diminifli the capacity of the lower belly. 1 1,10-puEio-cosTiAN, exterior obVique or large ohhque. 95. Thcfe mufcles cover two others which pro- ceed alfo from the ridges of the ilium to the carti- lages of the aflernal ribs j they are fixed behind to the fpinal apophyfes of the laft two vertebra3 of the loins, and to the os facrum ; they are united be- fore by a double aponeurotic expanfion along the median line^ and have the fame acSlion as the prece- ding. Ilio-costian, interior oblique QxJniallolUque. 96. Between the aponeurotic leaves of the two preceding mufcles, there are two mufcular bands which extend from above the pubis to the -fi-cr- num ; to its appendix, and to the cartilages of the laft iternal ribs. The aponejLirotic leaves which receive thefe j; mufcles ■■H? MUSCULAR SYSTEM. 83 rnufcles adhere to them in fome points, and pro- duce on them three or four tranTverfal tendinous lines. They bend the bread on the pelvis. Pu- :bio-st ETi'^iAi^ f Jrraight of thd lozver helly. 97. A mufcle, fixed by a long aponeurotic ex- panfion to the tranfverfe apophyfes of the iirll three lumbar vertebras, proceeds, in a crofs direc- tion, to the median line. It adheres alfo to the cartilaginous edge of the thorax, the n'dgc of the OS ileum, the crural arch, and the pubis. Thefe mufcles comprefs the fides of the abdomen. LuMBO-AB DOMINI AN, tranfverje of the lower hlly, 98. From above the pubis rife alfo two fmall mufcles, which terminate in a point below the na- vel. PuBio-suB-UMBiLiAN, pyramidal. 99. Mufcles which move the head. The difJDO- fition of the mufcles, which ferve to move the head, varies according to the manner in which the head is articulated with the neck. Thcfe mufcles, in animals which have the neck long, fuch as the inofl part of birds, or thofe which have no thorax, as ferpcnts, are fixed to a point in the neck : in the mammalia they are fixed to the neck, the breafl, and the fhoulders. In quadrupeds, whofe head being often \&vy large is fufpended before the vertebral column, G 2 and 84 ORGANIC STRUCTURE. and has need of being continually fupported, ihefe mufcles mufl be very ftrong *. loo. In many the mufcles which move the head have their fixed points of adhefion in the neck, the breail^^ and fhoulders : they are fituated before, behind, and on the fides. The rhufcles which have a lateral pofition move the head when they a6l fingly ; when they a6l limultaneoufly they direct it forwards or back- wards, or keep it fixed : the latter are the mofl numerous. I or. To the fummit of the fternum and a part of the clavicle are fixed two portions of a mufcle, which unite, and afcend obliquely outwards -and backwards, in order to be inferted by a broad flat tendon at the bafe of the mafi:oid apophyfis. The aflion of one of thefe mufcles makes the head turn on one fide : when they a6l together they bend it forwards. Sterno-mastoidtan, idem. 102. A mufcle is fixed before to the tubercles of the tranfverfe apophyfes of the lafl; five verte- brae of the neck, by the fame number of fmall bands, which unite in afcending. This mufcle is joined to its fellow before the firfl vertebra, and ** The head of thefe animals js retained chiefly by a verj broad and thick ligament^ which is attached to the Ipinal apo- phyfes of the back and neck, and is fixed to the occiput \ it is known by the name of the cervical ligament. both MUSCULAR SYSTEM. 83 both are inferted before the large occipital fora- men. They raife the head when inclined back- wards, and bend it forwards. The large tra- CHELO-suB-occiPiTiAN, anterior large Jiraight , 103. Behind this miifcle is another very fmall one, fixed before and to the tide of the atloid, which afcends within and is inferted before the large occipital foramen. It has the fame a6lion as the preceding. Small trachelo-sub-occi- PiTiAN, anterior fmall Jtra'ight, 104. Another fmall mufcle, fixed before the tranfverfe apophyfes of the atloid, is inferted near the maftoid apophylis. It a<5ls like the preceding. Atloido-mastoidian, lateral fmall Jlraight. 105. A mufcle is fixed behind along the fpinal apophyfes of the firft five vertebrae of the back, the lafl vertebra of the neck, and to the cervical liga- ment ; it then rifes, becomes broader, and is in- ferted in the tranfverle apophyfes of the laft two vertebrae of the neck {cervical fortiori), and the mafloid apophyfes {majloidian portion) ; it carries the head backwards. Cervico-mastoidian, fjjlenius of the head. 1 06. To the firft four vertebrae of the back and thofe of the neck, the firft excepted, are fixed the fame number of fmall tendinous bands, which unite in afcending. This mulcle proceeds back- wards, and is inferted by a flat tendon towards the middle of the occipital arch clofe to its neighbour, G 3 from S6 ORGANIC STRUCTURE. from which it is feparated only by the cervical li- gament. It keeps up the head, and pulls it back- wards. Trachelo-occipitian, large complexus and d'lgajlric of the neck. 107. To the tranfverfe apophyfes of the firfl vertebra of the back, and the laft four of the neck, are fixed the fame number of fmall tendinous and flefhy bands, which unite in afcending. The muf- cle which they form is inferred by aflat tendon be- hind the maftoid apophyfis. It has the fame adlion as the preceding. TRACHELo-MASTOiDiAN,y»?(2// complexus or lateral majlo'idian, 108. To the tranfverfe apophylis of the firft vertebra of the neck, is fixed behind, by a thick tendon, a mufcle which becomes much broader, afcends obliquely behind and on the fide, and is inferted in the lower part of the occipital arch near the mafioid apophyfis. Thefe mufcles elevate the head. Atloido-sub-mastoidian, y/z^mor oblique or fmall ohliqiie. 109. A mufcle, which grows broader as it rifes, and is inferted on the fides of the occipital arch, below the preceding mufcle, is fixed to the fpinal apophyfis of the fecond vertebra of the neck : it elevates the head. Axoido-cccipitian, ^o/^mor large fir aight. no. In the fpace comprehended between the preceding mufcles is a fmaller one, which proceeds from the pofterior tubercle of the atloid, expands as MUSCULAR SYSTEM. 87 as it rlfes, and is inferted on the fides of the occi- pital arch, behind the preceding : it has the fame a6lion. Atloido-occipitian, pojierior fmall Jlraight. III. There are fome mufcles alfo which pro- ceed from the head to the fcapula ;. but^ as they a^ more powerfully on the fhoulder, we (hall in- troduce them in another place. All the mufcles here defcribed fcarcely ever atfl; iinglj, but always in concurrence with a greater or lefs number of others ; and it is the indefinitely varied combination which may take place between all thefe portions of force, that gives rife to the pliability and variety of motion of which the parts of our bodies are fufccptible. IIS. Mufcles which move the face. In all ani- mals the face is overfpread with mufcles which fur- round the eyes, the nofe, and the mouth; and thefe mufcles, covered by a flefhy panicle, have no other a6^ion than that of enlarging, contract- ing, or flmtting the apertures of thefe different organs. But in man, thefe mufcles compofe a niovcablc phyfiognomy, the features of which become the living expreffion of the various fenfations he ex- periences. 113. The head of man is covered in the middle by a thin aponeuiotie mufcle, which is fixed alone; 04. the gS ' ORGANIC STRUCTURE, the occipital arch, and, proceeding towards the forehead, extends to the eye brows and the con- vexity of the nofe. This mufcle corrugates the Ikin of the forehead, and elevates the eye brows, OcciPiro-FRONTiAN, e^lcranius and fyramidal of the nofe. 114. Two fmall mufcles, inferted in the nafal projedions of the os frontis, lofe themfelves in the fubftance of the eye-brow. They elevate and corrugate the eye-brows, Froistto-supercilian^ fu^erciliar. 115. A fmall mufcle ilTues from the bottom of the arch of thp orbit, and expands over the upper edge of the tarfal cartilage of the upper eye-lid. Itraifes it when Ihut. ORiUTo-pALPEBRALiANa ^levator of the upper eye-lid. 116. The eyes are furrounded by a mufcle, the femi- elliptical fibres of which feem to itfue from the interior angle of the orbits where they adhere, in order to proceed to the exterior angle : by contra6ting they fhut the eye-lids. Palpebra- iiAN, orhicular of the eye-lids, 117. A fmall mufcle, fixed to the maxillary bone, proceeds in a tranfverfe diredlion on the carti!ao;e of the nofe : it ferves to dilate the nof- trils. SuPER-MAXiLi.o-NASiANj tranfverfe of the nofe. I J 8. Below the alveoli of the upper incifive teeth is a fmall mufcle, fixed to the alae of the nofe. MUSCULAR SYSTEM, Sg nofe, which it tends to deprefs. Alveolo-na- siAN, myrtiform, lip. From the zygomatic arch arife two muf. cles, which defcend obliquely towards the angle of the lips : by contracting they carry the lips up- fvards and backwards. The frnaller of thefe two mufcles is placed abo^e the other. Zygomatic-labian^ greater and LESS, great 2iX\dfmaJl zygomatic. 1 20 Three fmall mufcles are fixed to different points of the fub-maxillary bone : the largejl arifes on that bone towards the interior angle of the eyes J proceeds to the upper lip, and leaves Ibme fibres on the fides of the nofe : the middle one is inferted on that bone, below the orbitar cavity, and expands equally over the upper lip ? the lejfer is fixed above the alveolar edge near the angular tooth, and proceeds to the angle of the lips. I hefe mufcles ferve to elevate different points of the upper lip, and even the noflrils, Super MAXiLLO-LABiAN, gheater, mean, and LESS ; I ft, Elevator of the ala naft and of the upper Up ; 2d, Incifive or proper elevator of the upper lip , 3d, Canine^ or elevator of the angle of the lips. 121. From the part of the maxillary bone which correfponds to the chin, arifcs an odd mufcle of a fquare form, which proceeds through the fub» ftance of the lower lip. It depreffes that part.. MEKT0-LA3iAN,y^z/^r^ of the chin* 122. At 00 ORGANIC STRUCTLTRS, 122. At the bottom of the lower jaw, and on the fides, arifes a mufcle, which expands as it afcends, and proceeds to the angle of the Hps. It depreffes that part. Maxillo- labian, triatigular, or deprejfor of the angle of the lips. 123. At each fide of the mouth Is a thick mufcle, which is fixed at the top and bottom to the alveolar fides of the two jaws, and behind to the inter-maxillary ligament : all the fibres of this ;nufcle take a crofs direction, and expand towards the angle of the lips : by contra6t:ing, thefe mufcles fwell out the cheeks. They were called buccinators^ becaufe they are thofe princi- pally put in aclion when a trumpet is founded. BUCCO-LABIAN. 124. Around the lips is an odd mufcle, the femi-oval fibres of which crofs each other towards the angles 1 they concur to form the thicknefs of the lips : by contra6ling they leflen the aperture of the mouth. Labian, orbicular of the lips ^ or fniall incifive of the upper and lower lip. 125. All thefe mufcles of the face are covered, and their int-ervals are filled up with a greafy tifiiie, exceedingly fine, and of a nature altogether peculiar : the fibres of thefe mufcles crofs each other in many points; and this difpofition increafes the relation of thefe parts, and gives them a great degree of mobility. 126. A broad thick mufcle occupies the whole temporal MUSCULAR SYSTEM". gl temporal fofTa and a part of the zygomatic. This mufcle, which is covered within by a ftrong apo- neurofis, is formed interiorly of two layers of fibres which unite below in a thick tendon: this tendon embraces the coronoid apophyfis of the maxillary bone, below the zygomatic arch. This mufcle tends ftrongly to elevate the lower jaw. Temporo-maxillian, crotaphite or tenipo7-al» 127. At the bottom, and on the fides of the cheek, is a thick mufcle, which is fixed below the zygomatic arch, to the os zygoma, and to that part of the fuper-maxillary bone united to it. This mufcle defcends along the branch of the maxillary bone, and is inferted in the exterior and inferior fide of the angle of the jaw. It has the famea61ion as the preceding. Zygomato-maxil- LiAN, majfeter. 128. In the cavity of the pterygoid apophyfis of the fphenoid is fixed a mufcle which defcends outwardly, becomes thick, and is inferted in tlie interior fide of the angle of the maxillary bone. It elevates the jaw, and tends to draw it back- wards. Great ptef.ygo-maxillian, great or mterior pterygoidian, 1 29. The pterygoid apophyfis receives alfo on its exterior fide a fmall mufcle, which proceeds hori- zontally before the condyle of the maxillary bone, to which it is fixed as well as to the articular cap- fule. This mufcle tends to draw the condyle of the gl ORGANIC STRUCTUSE. the jaw forwards. Small pterygo-maxillian^ /mall or exterior pterygoidian. 130. A mufcle extends from the maftoid apo- phyfis to the middle of the interior face of the jaw ; it defcends by a flefhy portion towards the angle of that bone, where it prefents a tendon^ which continues with another flefhy portion in- ferted in the genian apophyfes. This digaliric mufcle tends to deprefs the jaw. Mastoido-genian, digaflric. 131. Beneath the fkin, between the breaft and the chin, is a thin mufcle which originates below by a few fibres under the clavicle, and the acro- mion : thefe fibres approach each other, and rife nearly to the chin, where they crofs the fibres of the oppofite mufcle : the action of this mufcle is not fenfible. This mufcle is much ftronger and more ex- tended in animals ; it fpreads itfelf over the face, Stod forms the carneous panicle, which, as already faid, covers the mufcles of that part. It ftrongly corrugates the fldn of the neck, Thoraco- FAciAN, cut emeus. 132. Muscles OF THE limbs. The limbs of animals have different ufes, and perform various motions ; the rcfult not only of the configuration of the bones which form them, but of the peculiar difpofition alfo of the mufcles fixed to them. ThuSj in apes, it is not the peculiar fl:ru6lure MUSCULAR SYSTEM, gS of their bones that prevents them from Handing perfectly erd6l, with the leg extended^ and the fole of the foot entirely refting on the ground, but the peculiar difpoiition of the mufcles, which keep the .thighs half bent on the pelvis, and the heel raifed up. The cafe is the fame with other animals ; they all perform movements, which are the refult of the peculiar flrudture of the bones and mufcles of which their limbs are formed. The thoracic limbs of birds, covered with fea- thers, form win'gs proper for flying; their pelvian limbs ferve them for walking; and in fome they are furnifhed with membranes, and anfwer the purpofe of fins. ^ . Thofe of oviparous quadrupeds ferve for pro- greffion. Thofe of amphibious and cetaceous animals, being fhort and flat, are fitted for natation. Thofe of bats, covered with broad membranes, perform the fame office as wings. In the greater part of quadrupeds, however, they ferve only for walk- ing. In the quadrumana, the thoracic limbs ferve for walking and grafping. In fome genera of animals the limbs have alfo peculiar ufes: by carnivorous animals they are em- ployed for catching and tearing their prey ; by others they are employed for climbing trees, dig- ging in the earth, &c. In man the thoracic limbs ferve for apprchen- 8 lion. g4 ORGANIC STRUCTURJS, lion, and the pelvian for walking ere6l : he is the only animal who does fo. l^horacic Llmhs. 133. Mufcles which move the fooulder on the Ireajt. In man, the mufcles which give mvOtion to the fhoulder are fix on each fide. A mufcle of a trapezoidal form proceeds on the one hand along the fpinal apophyfes of the verte- brae of the back, from the feventh of the neck to the cervical ligament, and as far as the occiput ; and on the other extends along the upper edge of the fpine of the fcapula, and is fixed to its acro- mion apophyfis, and the humeral extremity of the clavicle. The fuperior part of this mufcle, the fibres of which defcend obliquely before, elevates the fhoulder ; the middle portion, which has tranfverfb fibres, draws it backwards; the lower part, the fibres of which afcend by proceeding forwards, tend ta lower the fhoulder. The total adion of this mufcle is to pull the fhoulder back- wards ; and when it remains fixed it moves the head in the fame direction, by being conne61ed with the occiput. This mufcle is united to its fel- low along the vertebrae, and both together repre- fent a fort of lozenge. Dorso-super-acromian^ trapezius, - 134. Below this mutcle is a fmaller one, nearly of a rhomboidal form, which defcends obliquely from MUSCULAK. SYSTEM. 05 from the fpinal apopbyfes of the lafl; two or three cervical vertebrae, and the iirfl: three or four ef the lumbar, and proceeds along the pofterior edge of the fcapula below the fpine of that bone. This mufcle elevates the bafc of the fcapula by drawing it backwards; and thus lowers the ante- rior angle of the flioulder. Dorso-scapulian, v rhomboid. 135. Higher up than the preceding mufcle is another, which is inferted in the tranfverfe apo- phyfes of the firft four vertebrse of the neck, and defcends obliquely towards the pofterior angle of the fcapula to which it is fixed, as well as to the portion of the pofterior Q,dgQ which is above the fpine of that bone. This mufcle raifes the fhoulder, as the preceding one does, by caufing it to make a kind of movement like that of a lever, which de- prefles the humeral angle. Traghelo-scapu- LiAN, angular. 136. To the anterior part of the four ribs which follow the firft, are fixed, near to their cartilages, the fame number of mufcular digita- tions, which afcend obliquely behind, and unite in a flat tendon inferted in the coracoid apophyfis* This mufcle carries the fhoulder forwards, by depreffing its humeral angle. Costo-coracoi- DiAN, anterior ferrate J ox fmall 'perioral. 137. Above and between the firft eight or nine ribs is fixed, by the fame number of digitations, a large g§ ORGANIC STRUCTURE* large mufcle, which becomes narrower, proceeds backwards, palling under the feapula, and is in- ferted in the poftcrior edge of that bond. All the portions of this mufcle tend to draw the fhoulder ftrongly forwards, at the fame time that its upper part raifes and its lower deprefies it, CosTo-sc AP uLi AN, largcjer rated. 138. A fmall mufcle extends obliquely from the anterior edge of the firft rib to the middle of the clavicle ; lituated between thefe two bones, it appears that it ought to fix the claviclco Costo- c LAV I AN, Jub-clav'iar. 139. A {imple indication of thefe different mufcular forces, and of the combinations which can be formed of them, ferves to acQOunt for the variety of motions of which the (houlderis fufcep- tible. It is to be remarked that the fFioulder has this peculiarity, that the bone of which it is ef- fentially formed is in a manner free, and fuf- pended amidft mufcles which fuftain it, and give it motion ; and that it is on this bone that the arm is articulated. In the different actions of the mufcles of the fhoulder we mull always keep in mind that the fcapula is confined by the clavicle^ which regulates its progrefs, and often occalions very compound motions on the principle of the ^ lever. 140. Mufcles which 7nove the arm on the JhouU der» MUSCULAR SYSTEM. gj df.r. The mufcles which move the arm on the fhoulder are fixed to the bones of the fhoulder^ and of the breaft ; to thofe of the vertebral co- lumn, and even to thofe of the pelvis. 141. To the fternal portion of the clavicle, the llernum, the firfl feven or eight ribs and their car- tilages, is fixed a broad mufcle, the digitated por- tions of which extend in a tranfverfe diredlioa over the bread. This mufcle, while it proceeds towards the arm, becomes narrower; the inferior fibres ar« folded back behind, and at the top on the fupe- rior ones, and terminate in a broad, flat tendon which is inferted on the fuperior quarter of the humerus, along the exterior edge ; and groove of that bone. This mufcle, which forms the ante- rior edge of the cavity of the arm-pit, fervcs to pull the arm f^rongly forwards, and towards the fides of the breaft, by making it turn on itfelf from without inwards. Sterno-humerian,^;-^^/ peroral, J 42. Above the fhoulder is a thick mufcle, fixed before the humeral extremity of the clavicle, to the acromion, and along a part of the lower border of the ridge of the fcapulum. The fibres of this mufcle pafs above the articulation of the arm with the Qiouldcr ; the anterior and pofterior fibres are folded back under the middle fibres, and they all terminate in a tendon which is YOL. I. H inferted QQ ORGANIC STRUCTURE. inferted in the rough line at the bottom of the groove of the humerus. This mufcle elevates ftrongly the arm. Sub-acromio-humerian, dellold. 143. From the coracoid apophyfis arifes a fmall mufcle, which is fixed- within the humerus to- wards its middle part. It tends to pull the arm forwards, by brino'ins; it near to the trunk.. Co- :. A G o H u M E R ! A N , CO f a CO -bra ch'iaU 144. The face of the fcapula, which is above its ridge, ferves to lodge a mufcle, the carneous fibres of which terminate anteriorly in a tendon, which pafies above the jun61ion of the acromion with the clavicle, and defccnds to fix itfelf to the trochiter. This mufcle, fituated below the fub- acromio-huinerian , concurs towards the fame aClion. Superior super-scapulo-trochiterian, fu- per-Jp'inal, 145. Another very broad mufcle, fixed to the ridge of the hip bone and the os lacrum, afcends along the fpinal apophyfes of the lumbar vertebrae and the laft feven of the back: it extends over the laft four ribs by as many digitations. AH the fibres of this mufcle approach each other, and proceed towards the arm, where tbey terminate in a flat tendon, which is turned round on itfelf, and inferted at the bottom of the trochin, on the interior edge of the groove. This mufcle enve- lops the trunk behind: it forms at the top the poflerior MUSCULAR SYSTEM. gQ pofterior edge of the cavity of the arm -pit t by contra6ling it tends to pull the arm downwards and backwards, making it turn inwardly on its axis. LuMBD-HUMERiAN, greaf dorjal or very large of the hack. 146. To the inferior angle of the fcapula is fixed a mufcle, which afcends obliquely before, and terminates in a flat tendon : this tendon pafles behind that of the preceding mufcle, around which it turns, and is inferted on the interior edge of the humerus below its head. The action of this mufcle is analogous to that of the preceding. ScApuLO-HuMERiATsr, large round. 147. To the face of the fcapula, fituated below its ridge, is fixed another mufcle, the fibres of which terminate before and at the top in a tendon that pafl^s over the exterior fide of the head of the humerus, and proceeds to infert itfelf in the trochiter. This mufcle tends to make the head of the bone turn outwards. Inferior super-sca- PULO-TROCHiTERiAN, fiihfp'mal, an d fmaJl round, 148. The interior or coftal face of the fcapula receives, in its whole extent, a thick mufole, which produces before a large tendon that pafles over the interior fide of the head of the humerus, and is fixed to the trochin. This mufcle tends to make the head of the humerus turn inwards. SuB-scAPULO-TROCHiNiAisr, fub-fca^ular, 149. Mufcles '■juhl.ch move the fore-ar??i on the H 2 arrtu 100 ORGANIC STRUCTURE. arm. The fore-arm is moved on the arm back- wards and forwards : two irtufcles bend it, and one extends it. 150. One of the bending mufcles is compofed of two carneous portions united below. At the top, the interior portion of this mufcle is fixed by a tendon to the coracoid apopbyfis ; it then de- fcends to about the lower third of the humerus* The exterior portion exhibits a larger tendon, which furrounds^ at the top and within, the edge of the glenoidal cavity of the fcapula : it pafles above the head of the humerus in the infide of the articulation^ and, after contra6ling itfelf^ iiTues from it to lodge in the groove of that bone, along which it defcends. Thefe two portions unite into one mufcle, which foon becomes narrower, and gives birth to a round tendon : this tendon pafles before the articulation, and is infer ted in a tuber- cle, which is below the neck, and at the interior part of the radius. This mufcle bends the fore- arm on the arm, and by making the radius turn inwards and outwards it contributes to the motion oi fuphiations Scapulo-radian, biceps. 151. Before the bone of the arm, at its middle and exterior part, is another thick mufcle which defcends towards the bending of the arm, paffing below the preceding mufcle : it terminates below in a tendon, which is inferted in the coronoid apophyfis of the cubitus: it bends the fore-arm on the arm. Humero-cubitian^ interior hracUal, 7 ' Thefe MUSCULAR SYSTEM. 10! Thefe two flexor muicles are covered with a very thin aponeurotic membrane. 152. Behind the humerus is a broad,, thick mufcle, which adheres at the top by three feparate portions : one exterior and fomev'/hat anterior is fixed below the trochiter : the middle one, which is larger, is fixed by a flat tendon to the anterior edge of the fcapula, below its glenoidal cavity : the interior and pofterior, which is fliorter, is in- ferted behind, below the head of the humerus. Thefe three portions unite towards the middle of that bone, embrace it, and adhere to it nearly as far as the articulation, where this mufcle changes into a tendon, which pafl^s behind the articulation, and is fixed to the olecranon. This mufcle alone extends the fore-arm on the arm. Scapulo-ole- CRANiAN, brachial triceps. 153. On the exterior flde of the olecranon is a very fmall mufcle, which is fixed to the epicon- ' dyle, and proceeds obliquely over the exterior fide of the OS cubitus, in a fmall part of its extents The a(ftion of this mufcle is very weak. Epicon- DYLO-cuBiTiAN, fmall uncomus, 154. Mufcles which make the radius turn on ihs- OS cubitus. Four mufcles make the radius turn on the OS cubitus ; two of them by pulling it in- wards, and two by drawing it outwards. 155. Before, and at the bottom of the epi- H 3 trochlea. 102 ORGANIC STRUCTURE, trochlea, is fixed a mufcle, which defcends ob- liquely without, to infert itfelf before in the mid- dle of the radius. Epitrochleo-radian, round pronator, 156. Before the two bones of the fore-arm and below, is placed, in a deep fituation, a thin broad and carneous mufcle, which extends tranfverfely from the interior edge of the cubitus to the exte- rior edge of the radius. Cubito-radian, fquar& j)ronator, Thefe two mufcles make the radius bone turn before the cubitus, and thus produce the motion of 'pronation, 157. The bottom of the epicondyle gives birth to a mufcle which adheres alfo td the neighbour- ing articular ligaments : it proceeds outwards and inwards, around the head and neck of the radius, which it embraces, and defcends to infert itfelf along the interior edge of the fuperior quarter of that bone : it draws the radius outwards, Epi- CONDYLO-RADiAN, Jhort fuphiator. 158. To the exterior and inferior edge of the humerus, and before its epicondyle, is fixed a flender mufcle, which defcends along the con- vexity of the radius, and is- iinferted on the outfide, in the inferior extremity of that bone. This mufcle, which evidently tends to pull the radius outwards when it is before the os cubitus, C^n alfo drciw it inwards when thefe two bones are MUSCULAR SYSTEM. ' 103 are parallel. HumerO'Super-radian, longfupi- nator. Thefe two mufcles pull the radius outwards when it has been drawn before the cubitus : they thus produce the motion o^ fupination. The movements of pronation and fupinatlori are produced, in an eflential manner, by the a61ion of the radius ; but the os cubitus, and even the humerus, yield to thefe motions in a fenfible manner. 159. Mufcles which . move the wr'i/i on the fore- arm. Four mufcles move the wrift on the fore- arm ; two by bending, and two by extending it. On the interior fide of the fore-arm is a long llender mufcle, which is fixed at the top to the humerus behind the epi trochlea, clofe to the ole- cranon : it defcends along the os cubitus, and terminates below in a tendon inferted before the laft bone of the firft row of the carpus (ffiformej, Epitrochlo-carpian, interior cubital. 160. Before the epitrochlea is fixed another mufcle, which defcends obliquely on the fore-part of the arm; rejoins the radius, and accompanies that bone two thirds towards its lower part. This mufcle is terminated by a long tendon, which glides in the groove of the firfb bone of the carpus. (unciforme), and is inferted in the .fecond bone of the metacarpus. Epitrochlometacarpian, interior radial, H 4 The 104 ORGANIC STRUCTURE. The action of thcfc two mufcles tends, in an efTential manner, to bend the wrill on the fore- arm. i6i. On the exterior fide of the arm is a thick mufcle, which proceeds above from the epicon- dyle ; defcends a little inwards, and is fixed along the exterior edge of the cubitus, to which it itrongly adheres. This mufcle terminates in a tendon which pafi^es inferiorly behind the cubitus, and is inferted on the exterior fide of the fuperior extremity of the laft bone of the metacarpus. The action of this mufcle tends to pull the wrift backwards, by inclining it on the cubitus. Cu- BiTO-supER-METACARPiAN, exteHor cubital. 162. On the exterior fide of the arm there are alfo two mufcles, fixed fuperiorly to the ei)icon- dyle, which defcend together along the exterior edge of the radius, and become tendinous to- wards the middle of that bone. Thefe two ten- dons proceed together towards the lower part of the radius, and in that place feparate : one of them is inferted without on the fummit of the fecond bone of the metacarpus, and the other on the fummit of the third. This double mufcle ex- tends the wrift backwards, by inclining it on the radius. Epicondylo-super-metacarpian, j^r^ andjecond exterior radial. When thefe three mufcles a6l together they extend the wrift dire(?Hy backwards. S63. Of the four mufcles here dcfcribcd (the laft MUSCULAR SYSTEM. 105 laft two being confidered as one), it is feen that the two fituated before have a tendency to bend the wrift on the fore-arnij and that the two fitu- ated behind have a tendency to pull it back : but when the one before, and that behind, fituated on the exterior edge, adl together, they bend the wrift towards that fide : the cafe is the fame with the other two, fituated on the interior edge. The varied adlion of thefe four mufcles is ftill combined with that of the mufcles which ferve for pronation and fupination. The variety of the motions that can refult from the combination of thefe different forces may give fome idea of the great mobility of which the wrift is fufceptible. 164. The mufcles of the arm are covered with a thin aponeurofis, which envelops the arm beneath the fkin : it is confounded above with the tendons of the mufcles, and terminates below towards the articulation of the elbow. The mufcles of the fore-arm are alfb covered with an aponeurofis, which is fixed fuperiorly around the articulation of the elbow, and chiefly to the epitro- chlea. This aponeurofis is very ftrongon the interior fide of the fore-arm, where it furnifhes the mem- branous prolongations which penetrate between the fcapulo-radian and the humero-cubitian mulcles, and fupplies them with points of adhefion. It is thin on the exterior fide of the fore-arm, as well as to- wards the wrift, where it entirely difappears, This aponeurofis. 105 ORGANIC STRUCTURE. aponeurofis, by enveloping the mufcles, confines them, fapports them, and thus favours their a61ion. The mufcles, which ferve for the motion of the fingers, and which extend from the arm and fore- arm to the hand, have tendons which pafs over the circumference of the wrift. Thefe tendons are lirongly retained in that place by an aponeurotic annular ligament, broad, and in the form of a bracelet, which prevents them from feparating. The tendons of thefe mufcles are alfo received in that place in fheaths, or tendinous grooves^ along which they glide as far as the lower point, by which they are fixed: of thefe flieaths fome are common to feveral tendons, and others are peculiar to particular ones. The mufcles of the infide of the hand are co- vered with a very ftrong a^oneurofts, which lies immediately beneath the flcin. It feems to arife, from the interior part of the annular ligament, and expands on the inlide of the hand as far as the fingers. The interior face of this palmar apoiieurofis. feems alfo to give birth to membranous prolonga- tions, which form partitions along the bones of the metacarpus, and thus feparate the tendons , that pafs over thefe parts. 165. The palmar aponeurofis feems to con- tinue itfelf fuperiorly along v/ith the tendon of a 9 long MUSCULAR SYSTEM. 107 long and flender mufcle,- which ftill remains to be defcribed. It is fixed above to the epitrochlea : it is carneous for a fmall fpacc, but foon becomes tendinous : the flender and fiat tendon of this mufcle defcends along the interior edge of the fore-arm, and pafles over the interior face of the annular ligamentj where it adheres by fome ten- dinous fibres : it then expands, and feems to lofe itfelf, and to be confounded with the palmar aponeurolis. This mufcle can contribute to the flexion of the wriil, and feems to be capable of flretching the palmar aponeurofis. Epitrochlo- PALMiAN, pahnaris longus^ Jle?ider cubital. i66. Beneath the fkin of the palm of the hand is obferved alfo a fmall mufcle, exceedingly thin, the tranfverfe fibres of which feem to lofe them- felves under the palmar aponeurofis, and in the thicknefs of the fkin : its allien mufl be very weak. Palmo-cutian, ■pahnaris cutaneus. > 1 57. Mufcles 'which produce motion in the fingers. The mufcles which make the fingers move are very numerous : fome of then ferve for bending, and others for extending them ; and there are fome which make them move on all fides. Some of thefe mufcles are common to feveral fingers ; others are peculiar to one : fome of them alfo are very long, and fixed to the fore-arm; while others, much fhort- cr, are fixed to the carpus and metacarpus. 168. Along 108 ORGANIC STRUCTURE. 1 68. Along the interior part of the fore-arm is a mufcle fixed fuperiorly, by a very thick carneous part, to the epitrochlea, the interior fide of the cubitus, and the fore- part of the radius. This mufcle defcends till towards the middle of that bone, where it feparates into four difiin6l portions, which terminate inferiorly in the fame number of" thin, flat tendons. Thefe four tendons pafs to- gether before the wrift, and then feparate to pro- ceed to the laft four fingers. They glide before the bones of the metacarpus and the firfi: pha- langes, where thefe fmall tendons are perforated ; they then unite, and are fixed along the interior face of the fecond phalanges. This mufcle ferves to bend the fecond phalanges on the firfi, the firtgers on the hand, and the wrifi: on the fore- arm. Common epitrochlo-phalangian, j/«^- lime or perforated flexor. 169. Below this mufcle is another analogous, to it, thin above and thick in the middle. It is fixed along the interior edge of the cubitus, and divides itfelf into four portions, which terminate in as many tendons. Thefe tendons proceed below thofe of the preceding mufcle, as far as the place where the latter are perforated, traverfe the fiffures of them, and are conneded to the in- terior edge of the laft phalanges. This mufcle, which has the fame adion as the preceding, can bend alfo the third phalanges^ to which it is fixed. COMMOlf MUSCULAR SYSliJEM. 10^ Common cubito-phalangettian", profound ot perforating flexor, 170. Towards the fuperlor extremities of the bones of the metacarpus, between the two com- irion flexors of the fingers, are fixed four fmall rnufcles, which proceeding from the tendons of the profound flexor, are inferted along the inte- rior edge of the firfl: phalanges of the fingers. They are auxiliaries to the two mufcles between which they are placed. Palmo-phalangian, Itmibrlcales . 171. The interval comprehended between the bones of the metacarpus is filled, in the infide, by three fmall mufcles, which proceed from the carpus to the firfl phalanges. They contribute to bend the fingers. Interior metacarpo-super- PHALANGiAN, inferior OX hit ei'ior inter-ojfeous, lyi. Along the exterior face of the fore-arm Is a mufcle fix.ed to the epicondyle : it grows larger below the point where it is fixed ; and when it reaches the middle of the radius, ife divides itfelf into four portions, each of which covers its neigh- bour, and becomes tendinous. Thefe four ten- dons pafs behind the carpus, and feparate from it to proceed to the convex face of the third pha- langes of the fingers. This mufcle extends the four fingers which follow the thumb. Common- fiPICONDYLO-SUPER-PHALANGETTIAN, COmMon extenfor^ 173. The no ORGANIC STRUCTURE. 173. The interval between the bones of th6 metacarpus is filled up without, by fovir fmall mufcles, which proceed from the carpus to the phalanges. They contribute to the extenfion of the fingers. Exterior metacarpo-super- PHALANGIAN, fupeuor Or exterior inter-offeous. 174. Before the upper third of the radius, and along the interior edge of that bone, is fixed a thin mufcle, which terminates below in a flat ten- don. This tendon paffes to the exterior edge of the carpus, and proceeds to the laft phalanx of the thumb. This mufcle bends the two phalanges of the thumb on the bone of the metacarpus, and even that bone on the carpus. Radio-pha- lANGETTIAN OF THE THUMB, long fieXor of the tJjumh, 175. A thick mufcle is fixed to the bafeof the firft bones of the carpus, palles before the firft, and even the fecond bone of the metacarpus, and envelops the firft phalanx of the thumb. It tends to bend the thumb. Carpo-phalangian op the THUMB, JJjort flexor of the thumb, 176. To the top of the cubitus, on the outfide and for a part of its length, is fixed a mufcle, the tendon of which pafles behind the firfl: bone of the metacarplis, and is inferted in the firft pha- lanx of the thumb. The adlion of this mufcle is to extend the thumb. Cubito-super-pha- lanqiaN, long extenfor of the thumh. ]77. Another MUSCULAR SYSTEM. Ill 177. Another mufcle, much fhorter and thin- ner, proceeds in the fame dire6lion, and has the fame a61ion as the preceding : its tendon termi- nates at the fecond phalanx of the thumb. Cu- BITO-SUPER-PHALAXGIAN OP THE THUMB, Jhort extenjor of the thumb. 178. Along the interior fide of the third bone of the metacarpus is inferted a mufcle, which pro- ceeds in a tranfverfe diredion, and is fixed to the cubital edge of the firft phalanx of the thumb. li brings the thumb towards the other fingers, by pulling it forwards. Metacarpo-phalangian, adduSlor of the thumb. 179. To the exterior face of the cubitus is fixed fuperiorly a mufcle, which extends between the two bones of the fore-arm, and terminates inferiorly in a tendon, often divided, which is implanted in the fummit of the firft bone of the metacarpus, towards its radial fide. This mufcle removes the thumb from the other fingers. Cu- BITO-SUPER-METACARP.IAN OF THE THUMB, long abduBor of the thumb, 180. A fmall mufcle is inferted before the two firft bones of the \.\ In apes the two preceding mufcles are fixed to the tibia, much lower down than in man, and keep the leg in a continual flate of flexion, which prevents walking erecl. ao6. On the outfide of the thigh behind, is a mufcle divided inferiorly into two portions : one longer than the other is fixed to the bottom of the tuberofity of the ifchium by a broad tendon, which defccnds obliquely behind; becomes broader and unites towards the middle of the thigh with the fhort carneons portion : the latter is inferted in the middle of the femoral ridge; continues along this ridge, and that which proceeds to the exterior condyle. This mufcle terminates in a ftrong tendon, which paffes behind the exterior fide of the knee, and is inferted in the head of the perone. It bends the leg. Ischio-peronian", J amoral bleeps. 207. Behind the articulation of the knee is a fhort mufcie, deeply fituated between the bottom of the exterior condyle of the femur, behind the articular MUS'CULAR SYSTEM. 121 articular eapfule, and the pofteiior and fuperior part of the tibia. It tends alio to bend the leg. PoPLiTO-TiBiAN, fopUlean. ao8. To the anterior angle of the ridge of the ilium is fixed a thin and very long mufcle, which defcends before the thigh ; pafles on the interior iide of the knee, and is inferted in the upper part of the tibia, on the interior fide of its ridge, be- low the tubcrofity of that bone. This mufcle bends the leg on the thigh by pulling it inwards, as if to make it crofs the other ; it makes the thigh and leg alfo turn outwards when the mem- ber is extended. iLio-PR^-tTiBiAN, (tayhr mnj- fle^) fartorms. 2,09. A thin long mufcle attached before the ifchium and the pubis, by a broad flat tendon, defcends ■along the interior part of the thigh, and terminates in a round flender tendon, which pafles behind the condyles of the femur and the tibia. This mufcle then tarns round on the inte- rior fide of the knee, and is inlcrted in the top of the ridge of the tibia, near its tuberofity. It bends the leg. PuBio-PRi®T7BiA]si, Jhnder or Jtraight piierior. i\Q, All the mufcles of the thigh are enveloped and maintained in their pofitions, by a very ftrong fort of aponeurotic covering, which extends over the pelvis before the pubis, the ifchium, the an- terior . 122 ORGANIC STRUCTURE. ierior ansfle of the ilium, the exterior face of that bone, that of the facriim and the coccyx, and over the articulation of the head of the femur. This femoral aponeurojis is very ftrong on the ex- terior fide of the thigh ; adheres behind along the femoral ridge, and defcends on the knee, which it 'envelops on all fides. It fends out prolongations betvv^een the mufeles, and feparates them without adhering to them. It ftrongly fecures the mufeles which it envelops, furnidies them with a firm point of fupport during their contra6tion, and in this manner greatly contributes to favour their adlion. This aponeiiroiis exhibits at its upper part two difdn6l laminse, between which is lodged a parti« colar mufcle. This mufcle is fixed towards the top of the anterior angle of the ilium, and de- fcends between the two laminasof the aponeurofis as far as the middle of the thigh. The principal a61ion of this mufcle is to ftretch the aponeurofis, and thus to fuftain with more force- the mufeles of the thigh. Aponeurotic ilio-femorian, mufcle of the fafcia Jala, and aponeurofis of the fafcia lata. 211. Mufeles ivhich move the foot on the leg. The mufeles which make the foot move on the leg extend, and bend it^ or pull it to either fide. The feet are extended by the mufeles which form the calf of the leg. Two of thefe mufeles Q are MUSCULAR SYSTEM. 123 are placed clofe to each other : the interior, which is thicker and longer, is fixed behind the interior condyle of the femur ; and the exterior behind the exterior condyle of that bone : both of them become very thick along the upper half of the leg ; they then decreafe, are confounded, and terminate in an exceedingly ftrong and thick flat tendon which is inferted behind the calcaneum. Their adlion is to extend ftrongly the foot on the leg. Bi-FEMORO-CALCANiAN, gemellus^ gafirocnemian, ^12. Below thefe two mufcles is another^ al(b very thick, which is fixed at the top, behind the head of the perone, and below that of the tibia. This mufcle becomes very broad, and defcends behind along the tibia, to which it adheres in a fmall part of its extent : its inferior tendon is con- founded with that of the preceding mufcle, and both have the fame a6^ion. TiBio-CALCANiAN',y&/^«j'. Thefe mufcles, the gemellus, and the foleus, are thofe which exercife the moft eflential a6lion in progreflion, running and jumping. By extend- ing the foot, thefe mufcles raife the heel and ele- vate the body, which for a moment refis only on the point of the foot. The carneous fwelling of thefe mufcles, which conflitutes the calf of the leg, is obferved only in man : this firu61:ure, by increafmg the flrength of the leg, adds greatly to the beauty of its form. ^13. There 124 ORGANIC STRUCTURE. 213. There is alfo another xtvy Jlender mufcle which is attached on the fide of the exterior con- dyle of the femur ; it is carneous in its upper fourth, then changes into a fmall tendon which defcends between the two preceding mufcles, and is confounded with them below on the interior edge of their common tendon. It exercifes the fame a6lion. Femoro-calcanian, t'lhial or Jlender plantarh. In apes, the tendon of this mufcle does not ad- here to the calcaneum, but pafTes below, and conti- nues with the aponeurofis plantaris,as the epitrochh^ falmaris continues with the aponeurofis pajmaris in man ; fo that thefe animals cannot place their heel on the ground without bearing on this tendon, which prevents them from refting the whole fole of their foot on the ground: a condition neccflary for walking on two (qqU 214. Below and behind the upper articulation of the bones of the leg is attached a mufcle which defcends along thefe bones, and adheres to them : iris terminated inferiorly by a tendon which pailes on the interior fide of the malleolus, and is in- •ferted on the interior fide of the os fcaphoides and of the firft OS cunciforme. This mufcle extends the foot, by pulling it inwards, Tibio-sue- TAR SI AN, tihialis poficus, 215, On the fide of the tibia^ below its exte- 6 rior MUaCULAR SYSTEM. 125 rior condyle^ i$ attached a mufcle which proceeds on the exterior fide of the perone, and adheres to it for three-fourths of its length at the upper ex- tremity. This mufcle is bulky in the middle, and terminates below in a tendon, which pafles behind the exterior malleolus, glides on the fide of the calcaneum, then proceeds below the os cuboides, and pafles obliquely below the foot to infert itfelf under the tarfian extremity of the firft bone of the metatarfus. This mufcle extends the foot and pulls it outwards. Peron^o-sub-tarsian, long ^eronaus, a 1 6. Before the leg, and a little on the outfide, is a mufcle attached fnperiorly along two-thirds of the exterior edge of the ridge of the tibia. This mufcle terminates inferiorly in a tendon, which proceeding obliquely on the infide, pafies before the articula- tion of the foot, and is inferted on the interior fide of the firfl os cuneiforme. It bends the foot by pulling it gently outwards. Tibio-su3-tar.- giAN, anterior tibial. 2,17. Before and on the exterior fide of the lower two-thirds of the perone is fixed a mufcle, the lower tendon of which pafl!es behind the ex- terior malleolus in a groove, common to it with the preceding mufcle. This tendon proceeds on the exterior fide of the calcaneum, before and along the exterior edge of the os cuboides, and is inferted 126 ORGANIC STRUCTURE. inferted in the tuberofity which ferves as the tarfian extremity of the fifth bone of the meta- tarfus. This mufcle bends the foot by drawing it outwards. Great peron^o-super-tarsian, middle ^eronceus. 2x8. Before the lower half of the perone is at- tached a mufcle, thin at the top and thicker in the middle : the tendon in which it terminates de- fcends before the convexity of the foot, patles in a groove common to it with another mufcle, and proceeds on the outfide to infert itfelf along the exterior edge of the fifth bone of the metatarfus. The a61ion of this mufcle is the fame as that of the preceding. Small peron^o-super-tarsian ; Jmall peronaus. 2.1C). The mufcles of the leg are enveloped in an aponeurofis, very ftrong before, but much thinner behind, and particularly below, where it difappears. This tibial aponeurofis is attached fu- periorly around the knee, where it feems to be con- founded with that of the thigh : its anterior face throws out membranous prolongations, which are interpofed between the mufcles of the anterior part of the leg, and adhere to it. The tendons of all the mufcles, which proceed from the leg to the foot, are fecured by two annular ligaments, finiilar to that which, as already faid, furrounds the wrift in the form of a bracelet. One of MUSCULAR SYSTEM. 127 of thefe ligaments is at the bottom of the leg^ and the other above the foot : they fecure the ten- dons, which they cover and confine in their places. Below the foot is an aponeurofis, limilar to the aponeurofis pahnaris. It begins under the calca- neum, where it is very thick, and proceeds expand- ing itfeh^ towards the anterior extremity of the bones of the metatarfus. This aponeuroiis plan- taris is furnifhed with feveral prolongations, which penetrate between the mufcles, fnpply them with numerous points of adhefion, and are fixed on the iide of the articulations of the bones of the me- tatarfus with the firfl phalanges of the toes. 220. Mufcles which give motion to the foes. The mufcles of the toes, like thofe of the fingers, are exceedingly numerous : they bend or extend the toes, or make them move tov^^ards the fides. They are common to feveral' toes, or are peculiar to fome : in a word, they are affixed to the bones of the leg, or to thofe of the foot. 221. A mufcle, attached behind and towards the upper part of the tibia, is changed towards the lower part of that bone into a tendon, which pafTes in a groove behind the interior malleolus : it defcends below the calcaneum, and proceeds' outwards beneath the middle of the foot : in that place it receives a flefhy expanfion, which arifes from the calcaneum (acceffbry or fquare portion): this 128 ORGANIC STRUCTURE* this tendon then divides itfelf into four fmall flat tendinous bands, which proceed towards the heads of the metatarfuS; where they enter a groove common to tour other perforated tendons, through which they pafs, to proceed to the bottom of the third phalanges of the lad four toes. It bends or deprelles the toes. . Common, tibio-phalan- G E T T s A N 5 common long flexor of the toes. 222. Below the large tuberofity of the calca- neum is fixed a mufcle^ which proceeds before and divides itfelf into four tendons : the extre- mities of thefe tendons are cleft to afford a paf- iz .... The same. By bending the breast on the pehis, and forming the sides of the abdomen. 8. Ilio-pubio-co£tian. . . . ~. . Exterior oblique, or great oblique. 9. llio-costian, .... . . Interior oblique, or small oblique, 10. Pubio-sternian. . ..... Right of the lower bell}. 11. Lumbo-abdominal. ...*.. Transverse of tht lovjer belly. 12. I'ubio-sub-umbUical. . . , - . Pyramidal, MUSCLES TABLE OP THE MUSCLES, 135 JMUSCLES WHICH MAKE THE HEAD MOVE ON THE NECK, Bi/ bending it fortvards, and toxoards the sides. NEW NAMES. OLD NAMES. 1. Sterno-mastoldian. ..„,,- The Same, 2i GreaE-trachelosub-occipitian Grr'at straight, anterior, 3. Small-crachelo-sub-occipician. , . . Small straight, anterior. 4. Atloido-mastoidian. . . • . . Small straig/i:, lateral. By lehding it backwards, and totvards the sides. 5. Cervico-rtiastoidian. . * . . Splenius of the head. C. Trachelo-occipitian. ..... Great complex-us , and digastric of the tieek. '7. Trachelo-mastoidian. Small complexus, or lateral moitoiiian, 8. Atloido-sub-mastoidian. ..... Superior obliqHe, or small oblique. 9. Axoido-occipitian. ... ... Great straight, pcstericr. 1*?. AtioidO'OCCiiTitiaii. S7n.-dl straight y posterior. MUSCLES WHICH MOVB THE FACE, By co7'rugating the forehead, and raising the eye-broxvs and eye-lids, 1. Occipito-fiontian. . . . . i . Epicranian and pyramidal of tht r.ose, i, Fronto-supercilian. . . . . . . Superciliar, 3. Orbito-palpeljralian Elexulian. ... ... Grtat indented, S. Costo-clavian Suli-clavian. MUSCLES WHICH MOVE THE ARM ON THE SHOULDER, By pulling foi'xcards. 1. Sternn-humcrian. . , ', . . . Great pectoral. By pulling it upwards. C. ?ub-acromio-humerian. .- . . . . DelfAJ. 3. foraco-hiiiTierian. - , Coraca-brachhI. 4. Supei-ic.ipulo-tiOLhiiei ian, superior. . . Super-spimu!. By pulling it bachxoards. -.h. Lumho-humerian. . . .... Great dorsal, or very large cf the back, 6, Scapiilo-humerian Great round. By ?nahing it ticrn outxcards. ". ^upcr-scjpulo-trocliiterian, interior. . . Subspinous and imall round. ■ By making it turn inwards. ^. Sub scapu!o-tiochinian. • . ... Sub-scapular, MUSCLES WHICH MOVE THE FORE-ARM ON THE ARM, By bending it. 1. Scapiilo-raili-n; *..... Biceps, 2. lliiTnero-cul)itian. ... . . • Brachial, interior. By extending if. 'i. ^^capul.i-^ilccrnnifln, ...... Brachial triceps, ^ r.pi(.ondylo-cu!)irian. ..,.,. Small anconeus. J *- Tilt: u hur rauiclcs v.hich lower the taw tie attached tp the os hyoidesj a deicriptipn of them wilA he fouM uniii! till hcnii S}Stem of Di.iesli.n. M5)SCLES TABLE OP THE MUSCLES. 1-37 MUSCLES WHICH MAKE THE OS RADIUS TVRN ON THE OS CUBITUS, JBj/ carn/ing itforxcardsfor pronation. NEW NAMES. OLD NAMES. 4.' Epitrochlo-radun Round fronatcr. 3. Cubito-radian Square pronator, Bj/ pulling it outwards for supination. 3. Epicondylo-radian. > Short supinator, 4. Humero-super-ratiian. ..... Long supinator. MUSCLES WHICH MOVE THE CARPUS OR WRIST ON THE PORE-ARM, Si/ bending it. -J. Epitrochlo-carpian Cubital, inferhr, a. Epitrochlo-metacaipian Raiial inKrhr. By pitUing it backtcards. "}. Cubito-super-metacarpian. , . . . Cubital, rxter'ior, 4. Epicondylo-iuper-metacirpian 1st and 2ri. . Radial, rxterhr Yst and id. Brachial, anti-brachial and palmar aponeurosis ; annular ligament , muscles xvhich seem to act on t/ie palmar aponeurosis. 5. Epitrochlo-palmar • . Slfnder cubital, long palmar, i, Pklmo-cutian • • . Cutaneous palmar. MUSCLES WHICH CONTRIBUTE TO THE MOTION OF THE FINGI'.RS, By bending the last four fingers. \. Common epitrochlo-phalangian, . . . Sublime or prrf orated firxor . 2. Common cubito-phalangettian. , . . profound or perforating fiexor, 3. Falmo-phatangian. .... Lumbricaf. 4. MeUcarpo-phalangian^ interior. . » . Interosseous interior or inferior. By extending the last four fingers. 5. Epicondylo-sapcr-phalangettian, common. . Common extensor of the fingers. §. Metacarpo-phalangiaHj exterior. . . . interosseous, superior or ixierior. By bending the thumb, T. EaHio-phalanjettian of the thumb. . . . Lcnz flixor of Ihe thumb. 8. Cirpn-phalangian of the thumb. , . . Short fiexor of the thumb. By extending the thumb. 9. Cubito-!upcr-pha!angePian of the thumb. . Lmg extensor of the thumb, iO. Cubito-jupcr-phalangian of tiie thumb. . . short extensor of the thumb. By bringing the thumb nearer the other fingers. :■.. M-tacarpophalans'ar. 01 ihi ;h-ijnib. . . Adductor cf the thurai. By^ 138 • ORGANIC STRUCTURE. Bj/ removing the thumh from the other fingers. NEW NAMES. OLD NAMES. 12. Cubito-super-metacai-pian of the tHumb. , tons abductcr of the thumli. 13. Caipo-super-phalangian of the thumb. . lihort abductor of the tJiumh, Br/ carrT/'ing the thumb before the other fin f^ers. 14. Carpo-metacarpian of the thumb. . , -Metacarpianof the thumh ^ or oppossr^' By extending the index finger. 15. Cubito-saper-phalangeftian of the index. . Proper extensor of the index, JBy bending the little finger. 16. Carpo-super-phaiangettian of the little finger. Short flexor of the little fift^er, Bi/ extending the little finger. n. Epicondvlo-super-phalangcltian of the little 7 > Proper extensor of thelittlejitiger. finger. ....... y By removing the little finger from the rest. 38. Carpo-phalangian of the little finger. . . Abukctor of the little finger, ly. Carpo-metacarpian of the little finger. ... Opjtoser of the little finger. MUSCLES WHICH MAKE THE THIGH MOVE ON THE PELVIS, JBy 'pulling it hacktoards and outx:ards. 1. Sacro-femorian Gluteus maximus. 2. Greater ilio-trochanterian. .... Gluteus mediiis. 3. Lesser ilio-trochancerian. .... Gluteus minimus. By making it turn outxvards. 4. Sacro-trochanterian. ..... Pyramidal. 5. Iscliio-trochanterian. ..... Gem-ini. 6. Sub-pubio-trochanterian, interior. . j . Interior obturator. 7. Sub-pubio-trochanterian, exterior. . . Exterior obturator. S. Ischio-sub-trochanterian. . . . ^ square of the thighs. ■ By bending if forwards. 9, Pr^elumbo-trochantinian. .... Psoas. 10. Ilio-trochantiiiian. ..... Iliac, 11. tjuper-pubio-feuiorian. ..... Pectinalis. By pulling it inwards, 12. FublO", sub-pubic-, and Ischio-femorian. . . Three adductors, or triceps adductor. MUSCLES WHICH MOVE THE LEG ON THE THIGH, By extending it. " I, Ilio-rotullan. . ; Straight or slender anterior. 2. Triferaoro-rotuliaa. Vast interior, vast m terior^ and crural or triceps crural. By TABLE OP THE MUSCLES. 13§ Bt/ he7iding it. f.T;W NAMES. OLD KAMES, 5, Ischio-sulj-tibian. . ..... Semi-rtiembranous. A, Ischio-pi^etibian Seml-tendmsus. 5. Ischio-peronean • • B'.ceps. €. Poplito-tibian, . . - ... Popiitian. Bi/ bending it and pulling it intvards. ■7. Ilio-praetLb^2n. ; ,. Sartorius. ». Pubio-oraetibian. Slender intirhr. FEMORAL APONEUROSIS AND MUSCLE WHICH ACTS ON IT, Bi/ contracting it. 1. Aponeurotic-ilio-feniorJan Miuck of the fascia Utj. MUSCLES WHICH MAKE THE FOOT MOVE ON THE LEG, By extending it. 1. Bi-femoro-calcanean Gemellus, gastrocnemius. "i. Tibio-calcanean Soleus. 3. Femoro-cakanean Tibialis cr slenier plantaris. By exteiiding it and pulling it in'wards. A. Tibio-sub-tarsiar. Tihidlis posticus. By extending it and pulling it out~.vards. 5. Pcroneo-sub-tarsUii. . . , '. . Lon^ peroneas. By bending it. f:. Tibio-sub-ursian Tibialis amicus. By bending it and pulling it ouiti-ards. 7 Greater peroneo-supcr-tarsian. . . . Middle peroneus, • I.ess pcroneo-snper-tarsian. .... Less peromus. Tibial and plantar aponeurosis and annular ligaments. MUSCLES WHICH SERVE TO MOVE THE TOES, By bending the last four toes. 1. Tibio-phaUngeftian, common. . . . Long common flexor. 2. Cilcaneo-sub-phalangettian, common. . . Short common fiexor, 3. Mctatarso-inter-phalangian, infenor. . , Infrrior interosseous. 4. Planti-sub-phalaniian Ltmbrical. By straightening the last four' toes. 5. Peronco-supei-phalangettian, common. . Long common extensor of the toes. 6. Calcaneo-super-phalangettian, common. . Pedious, orshort common extensor cf the tees. 1 Macatarso-inur-phaUngianj superior. , . Superior interosseous Bi/ 140 ORCANIC STUUCTUREi* By hringmg the articulations of the bones of the metatarsus closer to the Jirst phalahgesy in the form of an arch. NEW NAMES. OLt) NAMES. 8. Trjnsverse-metatarso-phalangian. , . . Transverse cf the toes. By trending the great toe. 9. Pcronco-sub-phalangettian of the great toe. . L!:ng flexor ofthr ^rtat toe. 10, T»rso-sub-phalansian of the sreat toe. . . Short flfxor of the great toe^ By straightening the great tocj It. Peronco-jTiijcr-phalangettian. of the great toe. Extensor oftht great toe. By bringing the great toe nearer to the rest. 12. Metataiso-sub-phalangian of the great toe. . Adductor of thf%reat toe. By removing the great foe from the rest, ■13. Cilcaneo-sub-phllangian of the gieat toe. . AMuctor of the great toe. By bending the little toe. 14. Tarso-sub-fhalans'ian of the little toe. . . Short flexor of the little' toe. By removing the little toe from the rcsi^ ij. Cilcaneo-sub-phalanjian of the littk toe. Abiuctirof the VttU tu> CEREBRAL [ I'll 1 CEREBRAL AND NERVOUS SYSTEM. S3 7" The cerebral and nervous fyjtem in animals. The cerebral and nervous l}'(lem in zoophytes, has the fame appearance as the other parts ; for the gelatinous and homogeneous confidence of the fubilance of thefe animals prevents their dif- ferent organs from being diftinguifhed. It can- not, however, be fuppofed that this fyflem is wanting in animals, which have the fenfes of touching and fmell exceedingly delicate, and which are fenfible to the impreffions of the light, heat, and moifture : it is probable that it is equally difleminated throughout their whole fubftance ; and as thefe animals can be divided into feveral parts, fufceptible of increafe and life, this opinion feems thence to acquire a further degree of ftrcngth. The cerebral fyflem begins to be obferved in worms, the nereids and amphinoma j it confifts of a long cord divided by knots ; in the leech the knots are more perceptible, and appear to be ganglions. Thefe ganglions are very diftindl in the aculeated aphrodite (aphrodita acuJeata), where theyatTume an arrangement which is found among infedls. In the mollufca acephaJuy fuch as the oyfler, the cerebral fyftem confifts of two ganglions U'nitcd by two nervous cords. 142 ORGANIC STRUCTURE. In the gafteropoda, fuch as the fnail and the flug, this fyftem confifts of a cerebral ring around the cEfophagiis, and a greater or lefs number of ganglions, fcattered throughout the different or- gans. Thefe parts have a communication with each other by means of a cord, and give birth to the nerves. In the mollujca cephalopoda^ fuch as the cuttle- £fh (fip'id officinalis and/epia o^opiisj, the cartilage of the, head contains a peculiar cavity, in v;hich is lodged the cerebral collar, and this cartilage is pierced with holes to afford a paffiige to the nerves. Infe6ls and the crujlacea have a brain, confiiiing of two lobes, from which proceed nerves for the eyes, the antenns and the mandibles : this brain then furnifhes two nervous cords, which furround the Gsfophagus, and proceed to a feries of gang- lions, more or lefs numerous, united by a cord, and giving birth to the nerves which are diflri- buted to the other parts. 238. In vertebral animals^ the cerebral and ner- vous fyitem is more com.plex ; and notwithftand-' ing the great variety of configuration which mull occur among animals fo different as tithes, rep- tiles, birds, and the mammalia, this fyflem feems to be eftablifhed in them all, according to a uniform^ plan. It always confifts, ift, of an - organ contained in the cavity of the cranium r encephaJon ; CEREBRAL AND NERVOUS SYSTEM. 143 encephaJon; 2d, of a prolongation lodged in the hole or cavity of the vertebral column : the cere-' hral or rachidian prolongation ; '^di, of a nervous cord^ or a feries of ganglions, united by a nervous cord, fituated interiorly on the fides of the ver- tebral column : trifphnchmc. Ganglions alfo are found in various parts. Nerves diflributed to all the organs proceed from thefe different fources. 2,39. Eifcephalon. The encephalic fyllem, and its prolongation, are inclofed in a double mem« brane ; tiie exterior, which is thicker, is called the meninx of dura mater .-the interior (men'mghie) confifts of two membranes, the exterior one of which is called the arachnoid, and the interior the pia-mater. The encephalon always exhibits the two lobes of the brain, the layers of the ocular nerves, the cerebellum, and the commencement of the cere- bral prolongation. Thefe parts are very different-; in the various elatlesj but they almoft always give birth to twelve pairs of nerves which proceed from the cranium, to dillribute themfelves to the fame number of organs. The fiVft pair proceed to the noftrils, by tra- verting the os ethmoides. The ethno'idql. The fecond pair to the eye, where they form the retina. The ocular. The third pair to the mufcles of the eye. The common cculo-mufculdr. The 144 ORGANIC STRUCTURE. The fourth pair to the large oblique mufcle of the eye. The interior oculo-mujcular. The fifth pair divide themfelves into three branches, which are diftributed to different parts of the face. The trifacial. The fixth pair proceed to the right exterior mufcle of the eye. The exterior ocido-mnjcidar. The feventh pair proceed to the face. The facial. The eighth pair, to the labyrinth of the ear. The lahyrinthic. The ninth pair, to the pharynx and mufcles of the tongue. The fharyngo-glojfian. The tenth pair, to the organs of refpiration and digellion. The pneumo-m/iric , The eleventh pair, to the tongue and its muf- cles. The hyo-gloffian. The twelfth pair arife from the rachidian pro- longation, and afcend into the cavity of the cra- nium, from which they are diftributed to the cer- vical and dorfal regions. The trachelo-dorfal. In fifhes, birds, and reptiles, the different parts of the ehcephalon are more or lefs di(lin6l and feparated ; fo that thefe parts often touch each other only in one point : in the mammalia they approach each other, and arc in fome mcafure contbunded. \vi quadrupeds, the brain, which is flat and fliort, leaves the cerebellum uncovered behind. In CEREBRAL AND NERVOUS SYSTEM. 145 In apes it aflumes more thicknefs, and is conti- nued backwards on the cerebellum. This arrange- ment is more apparent in man. In fifhes, reptiles, and birds, the furface of the brain is fmooth ; in the mammalia it exhibits cir- cumvolutions. In man and the dolphin thefe cir- cumvolutions are numerous and deep : in apes and carnivorous animals they are lefs vifible, and in the rodentia fcarccly appear. In fiflie?, the encephalon is very fmall ; it oc- cupies only a part of the cavity of the cranium, and is compofed of a double feries of tubercles, united in a point. Two of thefe tubercles repre- fent the lehes of the brain j two others form the ocular layers ; that which conftitutcs the cere- bellum is comparatively very large ; in the laft place, - there is one which produces the con;j- mencement of the cerebral prolongation. The encephalon of fifhes-has behind the cere- bellum various other fmall lobes, which give birth to fome pairs of nerves, not found in the other clafl^s. The encephalon of reptiles varies greatly in the different genera of this clafs ; but it is ahvays formed of diftindl tubercles for the lobes of the brain, the ocular layers, the cerebellum, and the cerebral prolongation. In birds the encephalon eonfids of fix tuber- cles : two of thefe form the lobes of the brain ; VOL, X. L two 146 ORGANIC STRUCTURE. two others the ocular layers : one conftitutes the cerebellum, and the laft the cerebral prolon- gation. In the mammalia thefe parts approach each - other, To that the lobes of the brain are almoft united ; the ocular layers are confounded with the brain ; but the cerebellum remains difliniil. Each lobe of the brain has in the middle a cavity : 'ventricle. The ventricles of the brain affiime various configurations, and the fides of them have feveral prominences : that found in all vertebral animals is a protuberance marked vi'ith ftrias ahernately gray and v^'hite in the infide, among the mammalia j and with fome whiter lines, in the other claffes : corpora Jiriata^Jiriated bodies. Thefe bodies are very large in birds, and conftitute almoft alone the whole lobe of the brain. ' The upper part of the lobes of the brain, w^hich is continued with the ftriated bodies, and forms the hemlfpheres, is thin in fithes, reptiles, and birds : in the mammalia it alTumes more thick- oefs, and in man its thicknefs is confiderable. Thefe ventricles have in the iniide alfo feveral prominences, which in different ciailes of animals are wanting, either in v/hole or in part ; and which we Oiall examine minutely in man. In fifhes, reptiles, and birds, the layers of the ocular nerves are diftincl, feparated, and exhibit ventricles k • CEREBRAL AND NERVOUS SYSTEM. 147 ventricles in the infide : in the mammalia tbacy are found in the fubfiance of the lobes of the brain, and have no ventricles. Below and between the ocular layers is a third ventricle. The cerebellum in all animals is a diftinc^ part : in fifhes it is round and very large ; in birds it is flatted in a tranfverfe direction. In the mamma- lia there are two lateral lobes and a middle lobe. In mauj the middle lobe^ which is very fmall, is concealed by the other two. In fifhes,, reptiles, and birds, the furface of the cerebellum is fmooth ; in all the mammalia it is marked with tranfverfe and parallel furrows. The cerebellum is united to the brain by two tranfverfe peduncles ; and the latter proceed to two longitu- dinal peduncles, by means of which the brain is continued with its cerebral prolongation. The commencement of the cerebral prolonga- tion is generally marked by a diftincl tubercle, of a different form. Between the cerebellum and the rachidian pro- . longation is 2l fourth ventricle, in which is always obferved a thin medullary lamina, called impro- perly a valvula. All the ventricles have a communication with each other by foramina, the fides of which are contiguous. In birds, the anterior ventricles are fhut on the interior fide by a thin and radiated partition. The conduit which forms a communication bc^ L 2 tween 148 ORGANIC STRUCTURE. tween the third and fourth ventricle is called the aquedu^ of Silv'ius. The ventricles of the ocular layers have a communication, by this aquedu(5l, with the other ventricles. Before and behind the third ventricle is always found a tranfverfe cord : the two comnujfurae, the anterior of which is generally longer and thin- ner, the^o/^mor larger and ftiorter. Before the third ventricle is a funnel-like exca- vation, which is direded downwards, and conti- nues with a tube that adheres to a tubercle. This tube and tubercle are diflinguifhed by the naqie of the pituitary gland. In all vertebral animals is found a tubercle, C2Mtdiht pineal gJa7id. In the mammalia arc obferved fome parts which are not found in the other claiTes. In the bottom of the interval, which feparates the two lobes of the brain, is a large band, the extremities of which are folded downwards: nie^ foJohe, Below, is found a triangular protuberance, arch with three pillars : two of its angles are prolonged backwards and downwards j the third is tbrmed of two cords, united, which proceed forwards and downwards. This trigone is continued inferiorly and anteriorly with a thin partition, called the feptiim lucidnm. This partition, which feparates the two ventricles of the brain, is formed of two 6 contiguous CEREBRAL AND NERVOUS SYSTEM. 14g contiguous laminae ; the triangular fpace left be- tween them is known by the name of the j^ftb 'ventricle. In a part of the ventricles of the brain is found an elongated protuberance, which in quadrupeds is larger than in man^ cornna ammonis (ram's horns). x Above the aqueduct of Silvius there are four tubercles [tubercula quadrigemina)^ which are alfo larger in quadrupeds than in man. The two an- terior {nates), which are higher, are much larger in graminivorous than in carnivorous animals: on the other hand, the two pofterior {tejles) are much more voluminous in the carnivorous than in the graminivorous. Fiihes and birds have alfo four tubercles ; but they are placed before and above the ocular layers. In apes and in man, the ventricles of the brain are continued backwards, with a cavity {digitalis) which lodges a protuberance {thefpiir), 240. Cerebral or rachidian prolongation. The cerebral prolongation gives birth to as many pairs of nerves as there are inter-vertebral fpaccs. Each rachidian nerve arifes from two roots, fepa- rated by a thin membrane ; thefe two roots pafs through the meninx, and proceed to a ganglion which gives birth to two nervous branches. L 3 The 150 ORGANIC STRUCTURE. The rachldian or vertebral pairs are diilin- guifhed into cervical, dorfaJ, limihar, pelvian, and caudal; and are ditiributed to different parts of the trunk. The laft cervical and the firft dorfal pairs unite to form a plexus, which furnifhes the nerves of the thoracic member. The lad lumbar and the firft pelvian form the plexus^ which furnifhes the nerves of the pelvian member. '1^1, Trlfplanchnic. All vertebral animals, on the fides of the prasfpinal face of their vertebral column, have a nervous cord in the fame manner as fifhes ; or a feries of ganglions united by a ner- vous cord, in the fame manner as the mammalia and birds. This lyilem has a communication, by filaments, with all the vertebral and with fome of the encephalic pairs : it furnifhes branches which are difiribuled to the veflels of the three large cavities. 242. Cerebral aiid nervous Jyjlem of man. In man, the cerebral and nervous fyflem is compofed of the encephalic organ, the cerebral prolongation, the trifplanchnics and various ganglions and nerves which proceed from thefe different parts. Encephalon. The encephalic organ compre- hends the brain, the cerebellum, the mefence- phalon, and the commencement of the cerebral prolongation. It has the form of the cavity of the cranium. M CEREBRAL AND NERVOUS SYSTEM. 15 i cranium in which it is contained ; it is foft, and its gravity is to that of water as 13 lo is to 1000, It forms the thirtieth part of the weight of the body in adults, the feventh or eighth part at the period of birth, and nearly one hah*" of its total weight during the firfl months of conception. Its furface exhibits a number of deep finuous anfrac- tuofities over which the fanguine vefTels are fpread. The blood which arrives at this oro;an has been efli mated at nearly the feventh or eighth part of that which ifTues from the aorta. The cerebral arteries are exceedingly fiexuous, and thei;; tides are very thin : at the bafe of the cranium they form an anaflomotic circle ; they then expand over the whole furface of the brain, divide and fubdi- vide themfelves into communicating branches, and terminate jn very delicate ramifications, which penetrate into the fubftance of the brain, and in a great meafure difappear from the fight. The blood returns by veins without valves ex- ceedingly flexuous, which prefent frequent anafto- mofes, and do not follow the dire6lion of the ar- teries ; they form diftincSt branches, which, inftead of uniting to conftitute trunks, pour the bJoDd obliquely, in the inverfe diredtion of its courfe, into linufes or membranous ducts. Though we do not know what takes place be- tween tbefe two orders of circulation ; though the encephalic organ prefents itfclf under a pulpy and L 4 homo- 152 ' ORGANIC STRUCTURE, homogeneous appearance, and though microfcopic obfervation (hows nothing in it but a colledlion of very fine globules, every thing announces the moft perfedl organization^ and the moft import- ant fecretion. In the interior, the encephalic organ exhibits a ilru61ure altogether peculiar; the fides of the ca- vities are lined with a thin membrane ; there are found alfo protuberances, fmall bands^ cords, ftriae of different forms, colours and dcniities ; holes, fif- fares and conduits, which have a communication with each other ; and the whole of thefe forms, exceedingly numerous, exhibits a confiant regu- larity. In the lafl place, this encephalic organ and its rachidian prolongation are covered by a double membranous tunic, and enclofed in flrong ofleous cavities ; they give birth to numerous pairs of nerves, which proceed to every part of the body ; thefe nerves divide themfelvesinto branches, which are fubdivided into very delicate ramifications, that fpread in all the organs, penetrate into every part of them, where they terminate in different ways, and carry thither the principle of a6fion and of life. As we are unacquainted with the particular ufe of the different parts which enter into the compo- fition of the encephalic organ, the defcription of them is merely graphic. 243. The CEREBRAL AND NERVOUS SYSTEM, 1^3 2,43. The double membrane which envelops this organ {men'mx and men'mg'me) form folds which are interpofed between its lobes. The 7nemnx {jdura mater) is a thick, femi-tranf- parent membrane, over which the meningian ar- teries are fpread : it lines the interior face of the cranium, to which it ferves as periofteum and ilrongly adheres to it, efpecially along its futures and at its bafe. Its interior face covers the me-? ningine, without adhering to it. The meningine (pa mater) confifls of two lami- nae, clofely united ; the interior of thefe laminae furnifhes different folds, of which there are three principal ones. One of them extends from the ethmoidal ridge to the occipital protuberance, and forms a partition between the tw^o lobes of the brain : 7nedian feptum ; falx of the brain. Another is placed below the pofterior part of the brain, which it feparates from the cerebellum ; it extends from the occipital protuberance to the fuperior edge of the apophyfis petrofa; it is pierced before with a hole for the mefencephalon : trarifverfe feptim of the cereheJliim j tentorium of th^ cerehellum. A third extends from the occipital protuberance to the occipital hole, and feparates the two lobes of the cerebellum * : median fepum of the cerehel- lum {falx of the cerehellum). The * The meninx furnifhes alfo two foIJs, which extend from the 354 ORGANIC STRUCTURE. The meninx is prolonged through all the holes of the bafe of the cranium, and accompanies the parts which ifllie from them. It furnifhes chiefly a meningianjljeath to the rachis': this (heath adheres along the vertebral canal, envelops the rachidian prolongMion, and furrounds the origin of the nerveswhich proceed from it. The meninx in different parts of its thicknefs has difierent conduits, often of a triangular form {/mils'), vv'hich receive the blood of the veins, and pour it into the gulf of the jugular. Thefe linufes are very numerous : but there are reckoned to be four principal ones : one of them extends from the ethmoidal ridge to the occipital protu- berance, along the upper edge of the median fep- tum of the brain: median finus [fuperio?' longitu^ d'maljlnus). It there divides itfelf into two lateral fmufes hollowed out in the thicknefs of the adhe- ring edge of the tranfverfe feptum of the cerebel- lum, which extend as far as the occipito-petrean hiatus, and proceed to the jugular foflae. Thefe Unufes exhibit in the interior fome tranfverfe fraena, and a few fmall granulous bodies. At the bafe of the med'an feptum of the brain, and near the tranfverfe feptum of the cerebellum, is a clmridlan fmus (Jiraightfmus), which tranfmits into the lateral finufes, the blood it receives from the choroidian veins, and from ihe inferior the anterior cIlMoid apophyfes, to the pofterlor, and two others which ruix along the fmall alss.of the fphenoid. longitudinal CEREBRAL AND NERVOUS SYSTEM. 155 longitudinal finus. The latter proceeds along the median feptum of the cerebellum*. The meningine is a thin tranfparetvt membrane^ fituated beneath the meninx, and which envelops the encephalicorgan and its prolongation ; it is formed of two laminae, between which are di- iperfed a great number of veflels. The exterior lamina {arachnoid) covers the ex- terior face of that organ ; it accompanies, to the place where they iiTue, the nerves which proceed from it; and then bends back on the interior face of the meninx, in the fame manner as the ferous membranes-]-. This lamina has feveral prolonga- tions : one of them penetrates into the lateral ventricles, and covers the different prominences, * The meninx forms alfo feveral fmall finufes, fomc of which run along the pofterior edge of the median feptum of the cere- bellum. Occipital Jinufes ; others are placed along the upper edge of the petrous apophyfis : fjipsrior petrous finufes. Be- low the preceding, there are two others which extend along the articulation of the petrous apophyfis with the occipital : Inferior fetrousfimifes : they empty themfelves direftly into the gulf of the jugular. hfmus is fituated tranfverfely on the bafiliar apophyfis, and communi-cates with the two preceding ones. Two ca'venious fniifes are lodged in the carotid canals, and open into the petrous fimfes. In the laft place XMioJmufes., one before and the other behind the fella turcica, communicate with the cavernous, and form a circle (coronarj) around that part. f See Traitil des Membranes, du C. Richat. obferved 166 ORGANIC STRUCTt^RE. obferved m them. Another covers, vvith thd meninx, the rachidian prolongation; accompa- nies to the place where they iiTue the nerves which proceed from it, and bends back on the interior face of the meninx. The interior lamina of the ineninglne ip'ia mater) produces prolonga- tions which penetrate into all the anfradtuofities of the brain. It accompanies the meninx in its different paflages, and produces along the verte- bral canal indented expanfions, which are inter-" pofed between the anterior and pollerior bundles of the nerves o( the rachis. The furface of the brain exhibits numerous flex- nous and deep anfradliiofities, and is divided Xoa- gitudinally, at the fummit, into two lobes, fepa- rated by the median feptum. Each of thefe lobes may be divided inferiorly into three lobules. The firil reils on the orbitar arches; the fecond is lodged in the temporal fofiae ; it is feparated from the flrfl by a deep fcilTure: the third reiis on the cerebellum, from which it is feparated by the tranfverfe partition of the meninx. When the two lobes of the brain are feparated, there is feen a large white band, which unites thefe two bodies near their bafe. C. Chauifier has given it the name oi mejohlie* : (^corpus callofum). The * We ftiall provlfionally adopt the more rational language which C. Chauflier ha$ fubllitutcd for the whimfzcal and ridicu- lous ■M CEREBRAL AND NERVOUS SYSTEM. J s;- The fuperior face of the melblobe is covered by the fharp edge of the longitudinal -feptum^ and by fanguine vefleis ; it exhibits in the middle a fort of future, and on the fides tranfverfe ftriaa. The pofterior extremity of the mefolobe, which is broader than the ant(Jrior, forms a thick, round roll. The mefolobe occupies nearly the middle half of the fpace comprehended between the fore- head and the occiput ; it is, however, nearer to the former. It continues on the fides with the whiie fuhjiance (medullary) of the lobes of the brain. This part forms in the middle of the lobes an oval centre, furrounded by the gray fubflance (cortical). Below the mefolobe are the two ventricles of the brain (lateral ventricles). Thefe cavities, the fides of which are contiguous, defcribe in their paflfage two oval furrows, interrupted before, placed back to back internally, and feparated from each other by a thin lamina : median fejytum of the ventricles^ {Jeptum lucidum). Thefe furrows, taken towards their anterior and lous nances employed by the antients to denote the different parts of the encephalic organ. But, it muft be confefled, that zootomy will never be in poffeffion of a nomenclature completely fatisfac- tory, and fufceptible of being generally adopted, until intelligent anatomifts employ themfelves, as the modern chemifts have done, to reform the language of their fciencej and until, after adopting a method of nomenclature, they fhall have given to the different parts names fuited not only to the organs of man, but alfo to the fimilar or analogous parts in animals, fo as to conncft by lan- guage two bri^ncheg of natural hiftory, which ought never to be feparated. interipF 158 ORGANIC STRUCTURE. interior extremity, proceed backwards till they ap- proach near to the pofterior lobes, where they are refledled outwards, and return in the thicknefs of the middle lobes, directing themfelves forwards and downwards. The ventricles of the brain are broad at their entrance, and terminate in a point behind; in their paiTage they defcribe the portion of a curve, th6 convexity of which is turned outwards. The median feptum is formed of two very thin laminse : thefe laminae, which are not contiguous^ leave between them a vacuity, in which a ferous matter is fometimes colled ed. Ventricle ofthefep'* lum, {^th ventricle). The fides of the ventricles of the brain have pro- tuberances and cavities, which are fitted to each other, fo as to leave no vacuity between them ; they are lined with a very fine ferous membrane, which is a coritinuation of the meningine, and which fe- cretes a lymphatic dew ; when this lymph is col- letHed, it diflends the fides of the ventricles, and forms of them real cavities. Along the bafe of the ventricles proceeds a membrano-vafcular^/^^z/j. of a reddiOi colour, free and floating at the top, which adheres by its inte- rior face to the ferous membrane that covers the ventricles. This body feem.s to be formed by the union of the arteries which proceed from the mid- dle lobar, and the veins which convey the blood into the choroidian finus : choroid plexus. Q Below CEREBRAL AND NERVOUS SYSTEM. 15^ Below the median feptum and the choroid plexus, and at the inferior fide of the ventricles, is a body of a white fabftance, which has the form of a right-angled triangle, cerehral trigone, (arch with three pillars). The anterior angle terminates in two cords, which defcend between the layers of the ocular nerves. The two pofterior angles turn backwards and outwards, and terminate in a point in the exterior and inferior part of the ventricles. The inferior face, which is flightly excavated, ex- hibits a triangular fpace, on which are obferved lon- gitudinal ftrise in the form of a Jyre. On the fides of the ventricles there are two broad, round eminences, which approach each other before, and terminate gradually in a point behind. The outfide of them is formed of a grayifh fubftance, interfperfed in the interior with white ilrias, which proceed from a centre of the fame , colour, pyriform eminences, (ftriated bodies). Towards the middle of the ventricles of the brain, below and behind the pyriform eminences, are two whitifh ovoid protuberances, covered by the choroid plexus, and the cerebral trigone, layers of the ocular nerves, (layers of the optic nerves). ■ They are confounded on the outfide with the lobes of the brain ; and the exterior part of them coniifts of a white ftratum, which mixes itfelf with the gray fubftance in the centre. The pyriform eminences, and the layers of the ocular l60 ORGANIC STRUCTURE. ocular nerves, are feparated, on each fide, by a fur- row, which lodges a fmall grayifh femi-circular band. Thefc fmall bands are thicker and nearer each other before than behind. Small bands of the fyriform eminences, (fmall femicircular bands). In that part of the ventricles, which is turned pofteriorly outwards and downwards, are two whitifh, oblong, cylindric protuberances, thin and narrow behind : their anterior extremity, which cor- refponds to the termination of the ventricles, is broader and thicker, and exhibits three or four light tubercles, feparated by furrows : cylyndro'id p-otu- herances, cornua Ammonis, ranis horns. On the interior fide of the cylindroid protube- rances, at the termination of the poflerior pillars of the trigone, is obferved a grayifh lamina, indented and free towards its exterior edge. The bordered or fringed bodies, (corpora fimbriata). The exte- rior fide of thefe protuberances exhibits an emi- nence, broader behind than before, which corre- fponds to an anfra6luofity of the inferior part of the brain. The accejfories of the cylindroid frotube- ranees, (accefibries of the cornua Ammonis). Above and behind the cylindroid protuberances are two cavities, in the form of a bag, hollowed out in the fubftance of the pofterior lobes of the brain, appendices of the ventricles, (digital cavities). In thele cavities is obferved an unciform enmence, broad before, terminating in a point behind, and bent CEREBRAL AND NERVOUS SYSTEM. i5l bent inwards, (the f pur) which is formed by a pro- minence of a gray fubftance, cov&red by a white flratum. The fides of the ventricles are lined with a very fine ferous membrane, overfpread with arte- ries and veins. The veins unite to thofe which proceed from the choroid plexus, and form in each ventricle a choroldiah vein (vein of Galen), which approaches that on the oppolite fide, be- tween the layers of the ocular nerves. Thefe veins proceed backwards, and anaftornofe in a connmon fhort trunk, which enters the anterior extremity of the right finus. The lavers of the ocular nerves leave between them an interval in the form of a fifiure, the third ventricle. This interval is bounded at the top by the cerebral trigone, and at the bottom by the mefencephalon. The ocular layers are united at the bottom by a foft grayifh tubercle ; before this tubercle is obfervcd an elongated aperture : anterior aperttire (vulva) ; and behind a round aperture, pojierior aperture (anus) : thefe apertures cdmrnunicate with the lateral finufes, and arc cbvered fuperiorly by the cerebral trigone. The lateral finufes are bounded at the bottom by the rhefencephalon. Before the anterior aperture is a whitifh cord, which extends profoundly, in a tranfverfe direc- tion, into the fubftance of the middle ventricles VOL. I. M of iu2 0RGA^3IC STRUCTURE. of the brain, and turns in the form of a bow « anterior coimni[jura . Below the anterior commiffara is a funnel- Qiaped excavation ; the bottom of this excavation is continued with a folid grayifh peduncle, which terminates at a round tubercle, refting on the body of the os {^hcvio'ides-. juperf2he7ioidal -peduncle and appendix (pituitary peduncle and gland). Below and behind the pofterior aperture of the third ventricle is afecond whitifh cord, larger and (liorter than the anterior, which extends tranfverfely in the ftrata of the ocular nerves : pGJlerior commijfura. . Behind the pofterlor commifllira is a fmall grayifli conical body, of the lize of a pea, foftilh, and often containing .gravel in the infide : tlje conarium (pineal gland). The bafe of the cona- rium, which is turned forwards, feems to adhere to two very fine white cords, which pafs over "the pofterior commifllira, and arc continued on the interior fides of the ftrata of the ocular nerves: pedwicJes of the conarium (the kidneys). The conarium refts above the four vt'hite tuber- cles: tubercles uf the mefencephalon (tubercula quadrigemina). The two fuperior and anterior of thefe tubercles, which are oval and larger (nates), are continued before with the pofierior commifiLira. The two inferior ones, which are round and fmallcr (teftes), are continued with the fuperior CEREBRAL AND NERVOUS SYSTEM, l63 i\iTp&v\oY prolongations of the peduncles of the cere- bellum, and with a vvhitith lamina, rounded at the top and covered with horizontal Arise: medullary lamhia of the cerebellum (valvule of VieutTens), Below the polterior commifTura and the tuber- cles of the mefencephalon is a canal, which pro- ceeds downwards : intermediate canal of the ven- tricles (aqueduct of Sylvius). It proceeds be- tween the fides of a fourth cavity : fourth ven- tricle. This ventricle is fituated below and between the peduncles of the cerebellum. It exhibits be- fore, a longitudinal groove and two lateral ones, which terminate obliquely in a point on the firfl-, and refemble thfe cut part of a pen (calamus fcriptorius). Behind, this ventricle is clofed by the medullary lamina of the cerebellum ; the bafe of it correfponds to the cerebral prolongation, and at its fummit is found the aperture which con- ducts to the third ventricle. The third and fourth ventricles contain a fmall choroid plexus; their fides are contiguous, and lined Vv^ith a feroiis membrayie, overfpread with fanguine veflcls. This membrane continually pours forth a lymphatic dew, which is fome- times found in a pretty large quantity, and whiah, in this cafe, diftends the fides of thefe cavities. In the lafi: place, the brain exhibits below and in the middle two large peduncles (prolongations M 2 of 104 ORGAN-IC STRUCTURE. of the brain, crura of the brain, crura of the me- dulla oblongata) j thefe peduncles arife from the inferior part of the pyriform eminences. They feparate at their origin, and are placed behind the fuperfphenoidal fhank j they unite with thofe of the cerebellum, and concur together to the for- mation of the mefencephalon. Before thefe pe- duncles, there are two round tubercles tituated clofe to each other, which refult from the bifid termination of the anterior pillar of the trigone : pyriform tubercles (mammillary eminences). 244. The cerebellum is lodged in the occipital (o^dd, which determine its form : it is fituated un- der the pollerior lobes of the brain, from which it is feparated by the tranfve^fe feptum ; its volume . is nearly the eighth or ninth part of that of the cerebrum. It is divided pofteriorly, into two lobes, by a fcifTure, which lodges the medimi feptum, and on its furface has deep tranfverfe furrows, which interfe(5t each other at an acute angle. The furface of the cerebellum exhibits alio a pretty large number of mammillary protube- rances ; in general, there arc five on the upper face, ten on the inferior, and two or three behind. Between the lobes of the brain before is a pro- tuberance, placed behind the quadrijumelli tuber- cles : 7yiedlan protuberance of the cerebellum (fuperior vermicular eminence) ; at the bottom and in the middle CEREBRAL AND NERVOUS SYSTEM, 105 middle is found the median lohde (inferior ver- micular eminence). The number of laminae produced by thefe an- .fra(5luolities, is from 25 to 30 on each face. Be- tween thefe laminae are found others itnaller and thinner. The laminas of the cerebellum are formed of a gray fubftance, and ramifications of a white fub- ftance; the white ramifications of feveral laminag unite and form branches. Thefe branches pro- ceed to a white nucleus, placed in the middle of each lobe of the cerebelkim (arbor vita). The middle and anterior part of the cerebellum furnilbes two large cords or peduncles^ which arife in the white centre of its two lobes. Thefe pe- duncles become larger, and each divides itfcif into three btmdles : one afcends before, and unites with the inferior tubercles of the mefencephalon ; the other proceeds to the lateral parts of the bulb of the cerebral prolongation ; the third, which is much larger, concurs towards the formation of the mefencephalon : fuperior, middJe^ and inferior prolongations of the cerehelhm. The peduncles of the cerebellum are formed of a very white fabflance, which towards the centre contains a yellowilli. gray fubflance. They unite with the peduncles of the brain to form the ?nefen- cephahn (annular protuberdnce — pojis Varolii). 245. The mefencephalon is found below the M 3 brain ]66 OPvGANIC STRUCTUREo brain and before the cerebellum; its fuperior face, inclined backwards, correfponds to the third ven- tricle, and the intermediate canal of the ventri- cles ; its inferior face, which is rounded, refi;s on the balillary eminence of the occipital bone. On its middle part is obferved a longitudinal furrow or raphe, which lodges the trunk of the mefen- cephalic artery. Its anterior convex part is con- tinued with the peduncles of the brain ; its pofterior concave part correfponds on the fides to the pe- duncles of the cerebellum ; and in the middle gives birth to the cerebral prolongation. The exterior part of its fubllance confifts of white fibres, which lofe themfelves in the gray fubftance of the interior part. It. exhibits tranfverfe or hori- zontal fibres, interfedled at right angles by two bun- dles of a whitifh fubllance ; one on the right, the other on the left. Thefe bundles traverfe the mefen- cephalon, expand and proceed before to the pyriform bodies, and behind to the cerebral prolongation. 246. The commencement of the cerebral or rachidian prolongation is feparated from the me- fencephalon by a tranfverfe furrow; its- form is pyramidal and quadrilateral ; its anterior face, in- clined downwards, has in the middle a longitu- dinal furrow, on the fides of which are obferved the pyramidal eminences, Thefe eminences are feparated by a longitudinal furrow from two other em'mcnccs called the OJivana'ru ' \ ' 7 The CEREBRAL AND NERVOUS SYSTEM. ] 6/ The diameter of the rachidian p-oVongation (medulla elongata) is to that of the brain as i to 7 ; it has the form of ihe cavity which contains it. At its entrance into that cavity it is large, and decreafes towards the fourth cervical vert&bra, but increafes towards the following ones. It then decreafes along the dorfal vertebrae, and again becomes larger towards the laft of that region. This prolongation terminates, towards the iirti: Inmbar vertebra, in two tubercles, fituated before the other, and feparated by a groove : they give birth to a bundle of nerves which occupies the reft of the vertebral canal. The cerebral prolongation is of a fofter con- iiftence than the brain ; its fubftance is white, And in the infide has a grayifh furrow. It gives birth to thirty pair of nerves, which ifTue through the vertebral holes. 247. Encephalic nerves. The brain, at its bafe, gives birth to twelve pair of nerves, which ifTue through different holes of the bafe of the cranium. 248. Thtfrjt pair arife from the inferior and anterior part of the brain, by three filaments ; two of which coriie from the interlobar fciffure ; the third, interior, ifilies before and below the pyriform eminences. Thefe filaments unite into a thick triangular cord, which foon becomes fmallcrt this nerve advances under a furrow of M 4 the l6g ORGANIC STRUCTURE. the anterior lobe oX the brain, and approaches that of the oppofite lide ; it becomes larger and terminates on the fides of the ethmoidal lidge, in a round tubercle^ e'xceedingly foft, femi-tranfpa- rent and grayifh. This tubercle gives birth to from 20 to 24 very delicate filarnents^ which traverfe the holes of the cribriform plate of the os ethmoides, de- fcend along very fmall canals, and proceed to the pafal foiisq. A part of thefe nervous filaments are diilributed within, on the feptum, and with- out, on the fuperior and middle cornets: they then become fo flender and pulpy, that they efcape the fight. Ethmoidal nerve, olfa6Iofy. Xhis nerve, in moll quadrupeds, is hollow. 249. The fecond fair arife from the pofterior and- inferior of the ocular firata. The cords of which they are compofcd feparate by bending around the peduncles of the brain, and approach each other before the anterior com- ipiflura ; they then again feparate ; proceed ob- liquely towards the ocular holes, and penetrate into the orbit enveloped in a ftieatb, furnifhed by Ihe meninx. The ocular nerve paffes between the tendons of the four ftraight mufcles; when it arrives behind the lobe of the eye, it pierces the fclerotica and the choroid, becoming fmaller, and then expands to form the retina : Ocular nerve, opiic. • 250. TIk, CEREBRAL AND NERVOUS SYSTEM. igp 250. The third pah' arife from an excavation found between the peduncles of the brain, near the mefencephalon, and ilTue in feveral filaments, which unite into a cord. This nervous cord di- verges from that on the oppofite lide, advances behind the pofterior clinoid apophyfes, pafles through the fphenoidal fiffure, and penetrates into the orbitar cavity, where it feparates into two branches : one of tbcfe, called the fuperior, dif- tributes itfelf by feveral filaments to the fuperior flraight mufcle, and proceeds to the orbito-palpe- bralian ; the other branch, which is larger, divides itfelf into three fmaller branches : the firll^ or inferior, which is the largeft, diftributes itfelf to the interior ftraight mufcle, palling under the ocu- lar > nerve; the fecond, or exterior, which is longer, firft furnifhes a ramus to the orbitar gan- glion, and then diftributes itfelf to the fmall oblique mufcle ; the third proceeds to the inferior right mufcle : CaMMON oculo-muscular, common mover. 251. The fourth pair arife by two or three fmall filaments, clofe to each other, between the peduncle of the cerebellum, and the inferior tu- bercles of the mefencephalon ; turn round on the fides of the mefencephalon ; advance along the cavernous finus, in a canal formed by the meninx, and penetrate into the orbit by the fphenoidal riifure. This nerve pafi[es under the common oculo- 170 ORGAJNIG STRUCTURE, oculo-mufcnlar, proceeds towards the" upper and interior tide of the orbit, and diflributes itfelf to the large oblique mufcle: Interior oculo-mus« cuLARj pathetic. 252. The fifth 'pair arife from the inferior and anterior part of the peduncles of the cerebellum, near the mefencephalon, in two portions: one, behind, coniifts of about thirty filaments united ; the other, before, which is fmaller, confiils offe- veral lax filaments. This nerve pierces the meninx towards the petrous apophyfis, and form.s there a plexus, which gives three branches. The tri- facial, the trigemn'i. Of thefe three branches one penetrates into the orbit by the fphenoidal fiffure : the orhlto -frontal ; the other proceeds to the face by the large round foramen ; the fuh-maxillary : the third diftributes itfelf to the lower jaw : the maxillary'. A, The OKBiTO'FRO'S'TA'L, ophthahnic of inilis, on entering the orbit divides itfelf into three rami. a. One proceeds over the periofieum of the arch of the orbit; iffues through the faper-orbital foramen, and diftributes itfelf to the eye- lid and the fore-head : the palpehro-frontal (frontal or fu- percilian). In its palTage, it divides itfelf into two rami : the interior' rainus fends out a filament which uni'es to another of the nafal ; it ifibes from the orbit above the cartilaginous pulley f/r^f^'i J .'.and feparates into a great number of foiall CEREBRAL AND NERVOUS SYSTEM. I7I fmall branches^ which are diftribnted to the fronto-fupercilian mufcle, the occipito-frontian, the integuments, and the bulbs of the cilia. The exterior ramus, which is larger, follows the diredlion of the mufcle that raifes the eye- lid i ifllies through the fuper-orbitar foramen, and divides itfelf into a great number of filaments, diftributed to the eye-brows, the fore- head, and even to the fummit of the head : a filament unites to a branch of the facial in the fubflance of the eye-lid. h. The other ramus proceeds along the exterior fide of the orbit, diftributes itfelf to the lachry- mal gland and the upper eye-lid : lachrymal. In its paffage it throws out a filament which ifTues again through the fpheno-maxillary fifTure, and communicates with another of the fuperior maxil- lary ; a fecond filament pafles through a hole of the OS jugale ,and anaflomofes with another of the facial ; the trunk then traverfes the gland, and lofes itfelf on the conjund^ive membrane. c. The lad, the nafo-palpehral (nafal), throws out a filament on its entrance into the orbit: this filament unites to another which comes from the fuperior cervical ganglion of the trifplanchnic^ and both proceed to the orhitar ga7iglio7i. The nafo-palpebral nerve then proceeds towards the interior fide of the orbit, and divides itfelf into two rami. The fitO paffes through the anterior orbitar hole, 172 ORGANIC STRUCTURE. hole, and penetrates into the cranium^ traverfing the cribriform plate of the os ethmoides ; it then re-enters the nafal foflae, by another hole of that bone ; gives fome filaments to the anterior ethmoi- dal and frontal cells f defcends behind the- bones of the nofe along their cartilages, and expands on the lobe of the nofe. The fecond proceeds towards the interior angle of the orbit, where it unites to a filament of the frontal ; it then paffes under the cartilaginous pulley of the great oblique mufcle, diflributes it- lejf to the neighbouring parts, and terminates at the eye-lids : palpehral. The ORBiTAR GANGLION, ophthalmic or lenticu- lar, is very fmall ; it is found on the exterior fide of the ocular nerve, near its entrance into the orbit. It receives the two filaments already mentioned and another very fhort one tranfmitted to it by the common oculo-mufcular : there proceed from it ten or twelve filaments, which furround the optic nerve ; traverfe the fclerotica in different parts, creep over the choroid, and expand on the irisi irici?2> B. The super-maxillary, fuperior maxiU lary, ifTues from the cranium, through thefuperior maxillary tbramen. On ifiuing from that hole, it throws out a ramus which penetrates into the or- bit, by the fpheno-maxillary filTure; the orhitar : it then proceeds towards the exterior fide of that cavity, and divides itielf into two filaments : one of CEREBRAL AND NERVOUS SYSTEM. IJg of thefe unites to the filament of the lachrymal, which traverfes the os jugalis ; the other pierces the orbitar portion of that bone, and penetrates into the temporal fofla, where it unites with a fila- ment of the inferior maxillary. The fuper-maxillary nerve penetrates into the fumrait of the zygomatic fofla ; and furnifhes two filaments, which proceed to \he-fphenoidal ganglion. It throws out alfo feveral others which proceed behind the maxillary fin us : fome penetrate into the tuberofity of that bone, and terminate in the den tar cavities of the lalt molar teeth {pojierior dentary), the relt are diftributed to the gums and the buccinator mufcle. The fuper-maxillary then enters the fuper^ orlniar canal, along which it gives out a ramus {^in- terior dentary) that divides itfelf into feveral fila- ments. Thefe filaments pierce the alveolar edge, and diftribute themfelves to the roots of the other teeth : one of thefe filaments communicates with another of the pofterior dentary. The fuper-maxillary nerve then ifTues through the fub-orbitar foramen, and divides itfelf into a great number of rami, which have- a communica- tion with each other and with filaments of the facial ; their aggregate forms a fort of plexus, the ramifications of which are diflributed to the muf-* cles of the nofe, the lips and the cheek. The SPHENOIDAL GANGLION, fpheno-pahtbie, is fituated at the fummit of the zygomatic fofla ; 5 , it 174 ORGANIC STRUCTURE. it receives, independently of the two filaments furnifhed to it by the fuper-maxillary, a ramus formed by the union of a filament of the facial and another of the upper cervical ganglion. The firft filament (the fuperior ramus of the vidiari) ifTuesirom the injlex canal of the os tem- pofis, through the hiatus of that bone, efcapes from the cranium through the anterior foramen lacerum, and unites itfelf to the fecond filament ; the latter (the inferior branch of the vidian) arifes from the fuperior cervical ganglion of the great iympathetic, afcends along the carotid canal, and its union with the firft produces the pterygoidian (vidian) nerve ; this nerve traverfes the canal of the fame name, and proceeds behind the Jphenoi- dal ganglia?!. The fphenoidal ganglion furniflies feveral rami or filaments. a. The largeft of thefe proceeds towards the pofterior palatine canal, which it penetrates, after throwing out one or two filaments that pafs through the holes in the fuberofity of the palatine bone ; thefe filaments diftribute themfelves to the guttural conduit of the ear, to the velum palatiy and to the uvula. The gulturo-^alatine (palatine). This large ramus then traverfes the pofterior pala- tine canal, and on ifTiiing from it divides itfelf into feveral filaments, which are diftributcd on the palatin^e arch : the fuperior palatine. h. The other filaments, furniflied by the fphe- noidal CEREBRAL AND NERVOUS SYSTEM. I7S noidal ganglion, arife fram the interior pari of it ; penetrate into the nafal foilaSj through the fpheno- palatine foramen, and dillribute thcmfelves on the exterior fide of that cavity : The ethmoidal (fphe- no-palatine). One of thefe filaments defcends in the infide on the partition of the nofe, to the bot- tom of the entrance of the noftrils, where it pafles into the anterior palatine foramen, unites to that on the oppofite fide, and forms a bulb or ganglion, which correfponds to the palatine arch behind the dentes incifores : the nafo-palatine. c. The MAXILLARY, inferior maxillary, is the largeft branch of the trifacial : it ifi^lies from the cranium by the inferior maxillary foramen, pene- trates into the zygomatic fofi^a, and fends out a great number of rami, which form two bundles. The firfi: or fuperior one fiarnifiies : a. A ramus which defcends below the exterior pterygoidian, pafi^es before the condyle of the jaw, and expands in the zygomato-maxillian mufcle : the fuh- zygomatic. h. Two filaments, which afcend in the temporal fofia, difi:ribute themfelves to the mufcle which covers it, and unite with a filament of the orbitar : the iem'poro-mufcular . c, A ramus which defcends between the pterygo- maxiliian mufcles, gives fome filaments to the fmall pterygo-maxillian, as well as to the tempore- maxillian \ pafies behind the coronoid apophyfis of the l'^6 ORGAWIC STRUCTURE. the jaw, expands into ieveral filaments, in the bucco-labian mufcle, communicating with the fa-' cial and the fub-orbitar : tBe hueco-lahian, d. In the laft place a dehcate and profound ramus which defcends before, and didributcs itfelf to the grand pterygo-maxilhan mufcle : theperygO"' mujcular. The fecond or lower bundle furniihes : a, A ramus which defcends behind the condyle oi the jaw, fends out a filament to the auditory con-* duitj and another which communicates with the facial at the place where it ifTues from the flylo- iDaftoidian foramen. This ramus is then refledled before the ear, and divides itfelf into two filaments^ which are diftributed to the temporo-maxillian mufcle, accompanying the fupcr-facial temporal artery, and communicating with filaments of the facial : the cutaneous temporal. h. A ramus which defcends obliquely behind the ramus of the lower jaw, pafles under the fiib- maxillary gland on the mylo-hyoidian mufcle, pro- ceeds to the fides of the tongue, and lofes jtfelf at the point of it : the I'mgual. In its pailage, this ramus receives the tympanic ramus arifing from the facial : it then throws out two or, three filaments which form a plexus or ganglion, whence proceed other filaments which are diftributed to the fub-maxiilary gland and fur- round .the fub-lingual artery: the fub -maxillary y In CEREBRAL AND NERVOUS SYSTEM. 177 In the laft place, it furniflies fome filaments which are diftributed to the fab-lingual gland, and others which proceed to the membrane of the, mouth, and communicate with filaments of the hyo-gloiiian mufcle. The lingual then feparates into two rami ; one of which fends out filaments that proceed to the mufcles of the tongue ; the other furnifhes fome which are ramified on the fides and at the tip of it, and tvhich, by their termination, form the papiliss of that organ. c, A third ramus, which arifes from a trunk, common to the preceding, the maxillo-dentary^ defcends behind the condyle of the jaw, and pro- ceeds towards the maxillary canal. Before it en- ters this canal it gives out a filament which is dif- tributed to the maxillary gland, the mylo- and genio-hyoidian mufcles {the rnylo-hyoidian) : this ramus then runs through the dentary canal, and in its pafi^ge fends out filaments, which proceed to the dentary cavities of the large and of the firft of the fm^Il dentes molares {dentary). When it comes oppofite to the hole of the chin it furnifhes a filament which follows the diredlion of the den- tary canal, and is diftributed to other teeth: it then ifl'ues through that hole, is refledled upwards, and divides into feveral filaments, which expand to the mufcles of the lower lip and the cheek. 253, ThQ fixth pair arife in feveral united fJa- VOL. I, N ments ly.S ORGANIC STRUCTURE. men ts from the furrow which feparates the mef- encephalon from the cerebral prolongation. This nerve is fiat, advances towards the point of the petrous apophylis, and pierces the meninx to pro- ceed into the cavernous finufes, where it receives two filaments^ which embrace the anterior cere- bral artery : thefe filaments, which arife from the fuperior cervical ganglion of the tri-fplanchnic, form a. plexus around that artery ; it then pene- trates into the orbit by the fphenoidal fiffure, and diftributes itfelf to the exterior right mufcle of the eye. Exterior oculo-muscular, exterior motor, 254. The /eventh pair arife from the triangular foilet which is found between the mefencephalon and the exterior fide of the bafe of the rachidian prolongation, near the olivarian eminence : it pro- ceeds outwards with the labyrinthic nerve, then , bends forwards and proceeds to the auditory fora- men. This nerve foon feparates from the laby- rinthic to penetrate into the fpiroid canal of the temporal : it paffes through its different windings and gives feveral rami, which are diftributed to the face. The facial, the hard portioii of the feventh pair. a. One enters the cranium by the hiatus of the temporal bone, iffues through the anterior foramen lacerum, and unites to a ramus which afcends ^ from the fuperior cervical ganglion of the tri- Iplanchnic CEREBRAL AND NERVOtJS SYSTEM. l^Q fplanchnic by the carotidian canal : it then pro- ceeds to the fphenoidal ganglion ; the pUrygoidian. h. Others very fmall proceed to the mufcles of the fmall bones of the tympanum. c. A third, which is very delicate, penetrates into the tympanic cavity, pafies between the incus and the malleus, applies itfelf to the membrane of the tympanum^ iffiies from that_ cavity by the glenoidal fcifllire, and forms a communication with the lingual nerve of the maxillary: tlfe tym- fame: (cord of the tympanum.) d. The facial nerve ifllies from the fpiroid canal by the ftylo-maftoidian foramen, and immediately produces feveral rami : one afcends before the raaftoid apophyfiSj and diftributes itfelf behind the ear, the pqfierior auricular \ others defcend be- low the fterno-mafioidian mufcle, and proceed to the tracbelo-occipitian and the ftylo-hyoidian muf- cles : the fuh-majtoidmyi, and Jiylo~hyoid'ian : the former communicates with the pharyngo-gloffian ■mufcle. The facial nerve then defcends in the fubftancc of the parotid gland, where it divides itfelf into two branches. , e. The firit proceeds upwards, palfes before the articulation of the jaw^ and divides itfelf into feveral rami^ fome of which are diflributed on the temples: the fuper-orlntar. Others are diftributed belov/ the eye and lofe themfelvcs in the orbicular N 2. and 180 ORGANIC STRUCTURE. and zygomato-Iabian mufcles: the fuh-orhttar ^ the third proceed acrofs on the cheek, and expand into a great number of filaments on the mufcles of that part and on thofe of the lips, the lahiah thefe rami are anafiomofed with the preceding and with an orbitar ramus of the maxilla^y. f. The fecond branch defcends behind the angle of the jaw, and divides itfelf into fevera! rami: fome proceed before the zygomato-maxillian jnufcle, the mentonmary and divide into a great number of filaments which are diftributed to the mufcles of the chin : they communicate with the filaments of the fuperior branch and rami of the buccal. Others defcend under the chin : thefub- mentonniari and expand on the cutaneous mufcle into a great number of filaments, which communi- cate with fome arifing from the firfi: cervical pairs. 255. The eighth pair are placed behind the preceding, to which they are connected by fome filaments : the root of them is turned back on the pofterior edge of the median protuberance of the cerebellum : it is confounded behind with that of the oppolite fide. This nerve accompanies the facial to the interior auditory foramen, which it abandons towards the orifice of the fpiroid canal, and penetrates into the vefiibulumj where it divides itfelf into twobranches ; one proceeds forwards and expands in the form of fi br Lifh i[] the ramps of the cochlea ; the othef proceedv*^ CEREBRAL AND NERVOUS SYSTEM. 181 proceeds backwards and divides itfelf into three rami, which are diftributed in the veftibulum and in the interior of the femi-circular canals. The LABYRiNTHic ^'E'B.YE, fofl portion of the feventb fair. 256. The ninth pair of nerves arife in three or four filamenis, from the peduncle of the cerebral prolongation, near the olivarian eminence. This nerve iflues from the cranium by the pofterior foramen lacerum, and immediately receives a fila- ment from the facial, and another larger from the pneumo-gaftric. It defcends along the ftylo-glof- iian m'ulcle, throws out filaments which follow the divifions of the maxillo-facial artery, and which concur to the formation of the plexus furrounding the divifions of that artery. This nerve gives out a great number of filaments, which diflribute them- felves to the mufcles of the pharynx, and its in- terior membrane, the pharyngian. Some unite to others arifing from the pneumo-gaftric, the handle of the hyoidian and the tri-fplanchnic, and form a plexus on the fides of the pharynx. The nerve of the ninth pair then defcends be- tween the fiylo-pharyngian and ftylo-gloffian mufcles, gives filaments to them as well as to the tonlillas, and is lofi: by diftributing itfelf to the mufcles which are at the root of the tongue. The pharyngo-glossian, glosso-pharyn- GiAN, hranch of the eighth pair, , N 3- 257. The ' 182' - ORGANIC STRUCTURE. 2-5 7. The tenth pair arife from the cerebral prolongation by ten or twelve filaments^ which ifTue immediately behind thofe of the preceding nerve. Thefe two nerves pafs together through the pofterior foramen lacerum, where they are fe^ parated by a membranous produftion. This nerve defcends before the neck, and pene- trates into the thorax, where it gives out rami to the lungs. It then iffues from that cavity with the oefophagus, enters the abdomen, and diftri-, bates itfelf to the flomach and liver. The PNBUMO-GASTRic, eighth pair, wandering pair. Qn its iiTuing from the cranium, this nerve far- nidies filaments which communicate with ^the pharyngo-gloffian, the hyo-glollian, and the tri- fplanchnic ; it then defcends before the neck, and gives out a large ramus, which pafies behind the anterior cerebral artery, and which proceeds to the larynx, where it divides itfelf into two rami: one of thefe is diftributed to the mufcles of the pharynx and to the oefophagusi the pharytigian : the other, which is larger, penetrates into the larynx, enters the os hyoides and the thyroid cartilage, and divides itfelf into feveral filaments, which are diftributed to the mufcles and the iiiv terior membrane of that organ : the laryyigian. The pneumo-gaftric nerve then defcends on the exterior fide of the anterior cerebral artery; furniflies one or two filarnents to the cardiac plexiiSj C,EREBRAL AND NERVOUS SYSTEM. 183 plexus, and penetrates Into the breaft. It pafTcs before the fub-claviar artery, and on the left before the aorta, where it give§ a large recurrent ramus ; the tracheal. This tracheal ramus proceeds in- wards, is refle6ied on the fub-claviar artery or aorta, and furnifhes feveral filaments to the cardiac plexus: it then afcends again on the fides of the tracheal artery, gives numerous filaments to it as well as to the oefophagus, the oefophagmi ; and at length reaches the larynx, into v/hich it penetrates between the cricoid and the thyroid cartilages, vs'here it terminates in feveral filaments, which are diflribufed to the mufcles of that organ, forming a communication with thofe of the laryngian nerve. This nerve furnifhes alfo a very great number of filaments, which concur to the formation of the pulmonary plexus. The two pneumo-gaftric nerves delccnd along the oefophagus, one before (that on the left) and the other behind (that on the right). They reci- procally throw out a great number of filaments, which form a plexus around that canal. When they reach the abdomen, the one hefore divides itfelf into feveral filaments, vv^hich follow the fmall curvature of the fiomach, and expand on its dia- phragmatic face. Some of thefe filaments proceed to the liver, and concur towards the formation of the hepatic plexus. The one hehind divides itfelf ^Ifo into numerous filaments, fome of which are N ^ diftributed 184 OR; and meningine). The nerves have the appearance of whitifh cords^ compofed of fibres in himdles ; thefe fibres are furrounded by a firatum of cellular tiffiie, which dividing them from each other, ferves to unite them. This cellular tiffue forms alfo a com- rnon covering around the nervous cords. Not- withftanding this fibrous appearance, their compo- lition feems to be analogous to that of the brain. Each fibre indeed of a nerve feems to be formed of a pulpy fubftanccj contained in a membranous vagina or fheath : the pulpy fubflance of the nerves continues with that of the cerebral organ, and theic membranous fheath (neurolemma.) with the menin- gine. Reil has proved this mode of f^rudure, by plunging the nerves either into dilute nitric acid, or into an alkaline ley (of potafh or foda). In the. firll cafe, the nervous covering becomes diffblved^ and the pulpy fubftance remains concrete. In the. iecond cafe, the pulpy matter is dilTolved, and the jnembranous tube remains empty. The nerves are diilributed to the different parts ■ by CEREBRAL AND NERVOUS SYSTEM. 215 by the mere reparation of their numerous fmall fibres ; it does not appear that thefe fmall fibres ever divide themfelves info ramufculi. At their termination, the nerves feem to aban- don their membranous covering-, and to lofe them- felves in the fubftance of the different organs, by an expanfion of their pulpy fubftance. Towards the cerebral organ, the nerves ahb leave their covering, which is continued with the meningine, and their pulpy fubftance is confound- ed with that of the brain, the mefencephalon, or rachidian prolongation. Scemering fays, he traced the encephalic nerves, and particularly thofe of the organs of the fenfes, as far as the lides of the ventricles of the brain ; from which he concludes that the fluid contained in the interior of the ventricles mull be confidered as the centre of fenfation. From various chemical experiments made on the pulpy fubftance of the cerebral organ, it refults, that this fubftance is almoft entirely a fort of albu- men in a ftate of femi-concretion. P 4. TABLE TABLE OF THE CEREBRAL SYSTEM. Ithe cerebral system eontists of the encephalic organ^ the cere^ hrai prolongation^ the tri-fplanchnic and fome ganglhns. Tl^e encephalic organ comprehends the cerebrum, the cerebellum, the mesencephalon, ah^ the CQmmeiicem.ent of the cerebral pro- longation. It has the form of the cavity of the cranium in -which it is contained; its surface exhibits numerous winding and deep anfrac- tuositfe?, over which the sanguine vessels are spread; it is inclosed m a double covering-, the exterior, -which is very strong', is called NEW NAMES. OLD NAMES. Themeninx. . • . . ... Sura mater, and the interior, much thinner, .... ( Arachnaii. Meninsme. > . . - 'J J Via mater. The meninx is formed of two laminae closely united ; the exte- rior adheres to the cranium, and serves it as a periosteum; the interior is prolonged between the divisions of the cerebrum, and forms different folds, cH which there are three principal ones : One of them is between the two lobes of the cerebrum : ', Median septum of the cerebrum. . . ■ . Falxofthe certbrum. The other is placed between the cerebrum and the cerebellum : Transverse septum of the cerebellum. .' , ^ jTertionum of ths cerebellum. , The third is interposed between the two lobes of the cerebellum : Median septum of the cerebellum. , 1 . Falx of the cerebellum. The rneninx is prolonged irx the cerebral canal ; adheres to its sides ; serves them as a periosteum, and surrounds the vertebral nerves at their origin : it is reflected also into the different holes of the cranium, and confounded with the neighbouring periosteum. The rneninx exhibits, in different parts of its substance, sinuses , which receive the blood of the veins, and pour it into the jugular j of these sinuses, which are vesy nunjerous, there are four principal ones : Oni5 TABLE OP THE CEREBRAL STSTSM. fUJ One of them runs along the superior edge of the median septum of the brain : NEW NAMES. - OLtl If AMEA The median sinus. . . . , . Ssperkr tons'itui'mal sinus. It is continued with The lateral sinuses. . . , , . The savie. The third is at the base of the medito septum of the train : . Choitidiajvsinus. ..... Straight sinus. And the fourth runs along the longitudinal septum of the cere- "bellum: Inferior longitudinal sinus *. . . ■ . Th! same. The meningiue is formed of two veiy thin lamins, one of "which envelops only the exterior surface of the encephalon : " The arachnzil. The otlier is prolonged throughout all its anfractuosities : Pia mater. ■' The encephalon is composed, in the interior part, of a tuJiite sub- stance, covered by a thick stratum of a gray substance : ■White substance. ..... MeiuiUxry substance, Cray substance. . , . . Cortical sUbstanie. Tlie brain is divided longitudinally, on the summit, into two lobes ; and each of these is divided into three lobules : the antenor, middle, and posterior. The first two are divided inferiorly by a transverse scissure : Inter-Iobar scissure Scissure of Sylvius. Between the lobes of the brain is found a broad white band : Meiolobe. . . ... . . Corpus calksum. Below these are two ventricles, the sides of which are contiguous: Ventricles of the brain. lateral ventticles. They form two ovoid furrows, incomplete before, placed back to back within, and separated by a thin lamina: Median septum of the ventricles. . . . Septum lucidum. * The others arc thp occipital sinuses; the superior petrous sinuses; the inferior f«(tous sinuses ; the lltniTeHe sinus ; the cavei nous sinus; the coronary sinaiei. This 0,1$ ORGANIC STRUCTURE, This lamina is formed of hvo contiguous thin leaves, the sepa* ration of which can produce a triangular cavity : ySEVf NAMES. OLD Iv'AMES. Ventricle of the septum. .... Tlfth ventricle. The ventricles of the brain communicate, behind, with another cavity, the sides of which are contiguous: Appendix of the ventricics. . . . , Digital cavities. Along the base of the ventricles of the brain runs a membranouSjg vascular, and reddish plexus. riexus choroicles. . . . '. . Tm same. Below the septum, at the base of the ventricles, is a part of ^ triangular form : Cerebral trigone. . . > « . . Fornix zcith tiires pillars. On the sides of the ventricles are two oblong grayish bodies, placed back to back ; Pyriform eminences. . . . , .' . Striates boil's. Towards the middle of the ventricles of the bi-ain, below and be- hind the pyriform eminences, are two whitish ovoid protuberances, united by a small, soft, and grayish cord : Strata of'the ocular nerves, , ., . Strafaof the optic nemes. The pyriform eminences, and the strata of the ocular nerves, are separated by a furrow, which lodges a small grayish band : Small band of the pyriform eminences. . . Small semi-circular band. In the exterior part of the ventricles of the brain there are two protuberances, of the form of these cavities : Sylindroid protuberances. . . . Cornua Ammonis,, On the interior side of these eminences is a grayish serrated lamina : Fringed body. On the exterior side are two other protuberances: Accessories of the cylindroid protuberancp^'. . Accessories of the cornua Ammonis, In (he appendix of the lateral ventricles is observed I?]'^ unciform eminence. . o . . The spur. Between the strata of the optic nerves is found the trunk of 'Ihe choiridian vein. : . . ... Vein oj Galen, The strata of the ocular nerves leave between thern -m inter v;ii_ inihe form of a fissure : Third vcn'.ric:e, ■ , , , . , 'Chi 'arie, TiiJ5- TABLE OP THE CEREBRAL SYSTEM. 210 This ventricle eontains a small choroid plexus ; it presents before NEW-NAMES. OLD NAMES. It; anterior aperture Vulva, It exhibits at the top and behind Its posterior aperture. . , , . , Anus, Before the anterior aperture is a long, -whitish cord, which ex- tends across it, in the form of an arc : Anterior commissura. • ,' , , • ° T!:e same. Below this commissura is a funnel-formed excavation : Infandihulum. The bottom of this excavation is continued with a grayish cord, adhering to a round tubercle, which rests on the body of the sphenoid : Super-splienoidal peilur.cle and appendix. . . Fit-dUcry peduncle ani glan.l. Below and behind the posterior aperture of the third ventricie is another thick and short transverse cord: JPosteriov commissura. , , > , , The same. Behind the posterior commissura is a small conical and grayish body ; Conariom. . .<•..•. Pineal gland. It seems to adhere to two very fine cords, which pass over the posterior commissura, and proceed on the ocular strata : pedanculiofthe conatiam. .... Pedunculi of tk jiinea! gland. The conarium rests on the mesencephalon, below four white tubercles : Tubercles of the mesencephalon. • . . 5 ^■'»"-«''' J''''*'^""'''"'. ^ Testes et nates. The two inferior are continued with a whitish lamina, round at the top and striated: JrteduCary lamina of the ceretiellum. ... ValvulaofVieusens,' Below the posterior commissura is The intermediate canal of the ventricl;s. . , Aqueduct of Sylvius, This canal proceeds to The fourth ventricle, The same, yfhich is situated between the pedunculi of the cerebellum and the cerebral prolongation. This ventricle contains a small choroid plexus : it exhibits behind a Ion- ^2© ORGANIC STJ5.UCTURE* a longitudinal and two lateral grooves, •which terminate in a point on the former : ^-EV/ NAMES. OLT) NAMES. Calavius scriptorius^ The contiguous sides of ths ventricles of the brain are lined with a serous membrane, which is a continuation of the meningine; this membrane continually secretes a Ij^mphatic moisture, which lubricates its surface. In the last place, the brain presents below, and in its middle, C Prohngathn'of tht brain, crura (■/ the }rra:a, anm Itstwopedancuh. > '^ ,,i , „ ,, - (_ Ff the vieduUa oblongata* Before the junction of the pedunculi are found The pisiform tubercles Mammillary eminences. The cerebellum presents on its surface mammillarj' eminences, "and transverse and lamellar anfractuosities, ^vhich intersect each other at an acute angle. The centre of this organ is formed of a •white substance, which is prolonged and ramified between the gray substance and its laminas : Arisf vitis. Between the lobes of the cerebellum appears before* TliB median pvotubeiMUce. . . . . Superior vermicular eminence,. Below and in the middle of it. The median lobule. ... . Inferi'.r vermicular etninencf. From the white centre of each lobe arise before. The pedunculi of the cerebellum, ... Trolcngaticnff the cerebellum, \ "vvliich are divided into three bundles : superior, middle, and inferior proUmgatiyn], The pedunculi of the cerebellum unite to those of the brain to form The mesencephalon. . « „ . . . S ■^^"'^''''>'- P^f^l^""-"^- C. Pons Varolii. The mesencephalon is continued with the commencement of The cerebral prolongation, ..... Medulla oblongata, from which it is separated by a transverse furrow. The commencement of the cerebral prolongation is of a pyrami- dal form ; it exhibits before and in its middle a longitudinal fur- row, on the sides of which are observed The TABLE OP THE CEREBRAL SYSTEM. 221 NEW NAMES. OLD NAI.IES. The oliyarian eminences. ..... The same. Close to these eminences is another furrow, which separates them from Itic pyramidal eminences. ...... The same. The cerebral or rachidian prolongation has the form of the ver- tebral canal in which it is contained ; it terminates towards the first lumbar vertebra in two tubercles, and a loose tufted nervous bundle ; Sondlc of Che lumbar and sactaj Acrvcs. . . Cauda equina. It exhibits before and behind a longitudinal groove 5 it is whit^ ©fa soft consistence, and in the middle has a grayish furrow. The encephalon gives birth to twelve pairs of nerves, and th'3 £/;rebral prolongation to thirty. TABLE TABLE OF THE NERVES, All fhe nerves originate from the encephalic organ, therachi- dian prolongation, the tri-splanchnic, and some ganglions. The encephalic organ gives birth, at its base, to twelve distinct pairs of nerves; the rachidian prolongation to thirtj', and the tri- splanchnic to an indeterminate number. EN-CEPHALIC NERVES. NEEVE WHICH PROCKEDS ON THE SIDES OF THE NASAL CAVITIES. I Pair : arise by three filaments, tv^of Avhich come from iti interlobular scissure, and the third from the striated bodies : NEW NAMES. . OLD NAMES. Ethmoidal. ... . .... Olfactory. NERVE WHICH PROCEEDS TO THE EYE, AND FORMS THE RETINA. II Pair : derive their origin from the posterior tubercles of the ocular strata: NERVES V/HICH PROCEED INTO THE ORBIT, Distriduting themselves to almost all the muscles of the eye, III Pair : arise by several filaments from the excavation between the peduncuii of the brain, near the mesencephalon, Common_oculo*mu!cular. ..... Common motor. Distributing thcmsehes to the great oblique muscle of the eye. IV Pair: arise by two or three small filaments close to each other, between the peduncuii of the cerebellum, and the inferior tubercles of the mesencephalon. Jntcrior oCulo-muscuUr. . „ . . . Pathetic, NERVE WHICH PROCEfiDS TO THREE PARTS OF'THE FACJ:. V Pair: it issues from before the peduncuh of the cerebellunij, Hear the mesencephalon : , tiSscia!. . . i . o . .' . Tii^emersi, TABLE OF THE NERVES. 223 This nerve divides itself intothree branches : The first distributes itself to the eyes and the forehead, NEW NAMES. " OLD NAMES. A. Orbito-ffontal. ophthalmic cf Willis. It divides itself iiito three rami z 1. The paljicbro-frontal, ..... Frontal or iuperjliar, which gives a palpebralian ramus, that, expands in ciliar rarauscuil : 2. The lachrymal. ... ... Th-' same» S. The naso-palpebral -Vajji. The latter gives a filament to the orbitar ganglion — a ramus to the nasal — eth?noidal filaments — naso-lobar filaments — a palpebral ramus. GANGLION SITUATED NEAR THE INSERTION OF THE OCULAR - NERVE INTO THE EYE. Orbiter janjUon ; - . Ophthalmic cr lenticular ganglisru It receives A filament from the naso-palpebral — a filament from the superior cervical ganglion — a filament from the common oculo-muscular. It furnishes Ten or twelve filaments which proceed to the eye, and distribute themselves behind the iris : "** The irian. . . . . .... The r/'jr. The second ramus of the trifacial distributes itself to the middle part of the face : B. The super-maxiUarv Supiriar maxillary. . It gives, . First, An orbitar ramus, from which proceeds ajugaland a tem- poral filament ; second, one or two filaments to the sphenoidal ganglion; third, anterior and posterior den tar filaments; fourth, the sub-orbitar nerve. GANGLION SITUATED AT THE SUMMIT Of THE ZYGOMATIC yossA: Sphenoidal ganslion. . .,•... Sphino-palatine g.ini'.ion.. It 7'eceives It gives Two filament? ffom the super-maxillary, a ramus I _ .. , . . •„ i. " I Two fitamints if the tuptnor maxilla'}, the formed by a pte.yBO.Jian filament of the tri- ^ ftery^oU^ano, viiioniranch, •lili/..:tn;c, and a pJetysoJdian raiaus of the facial, J It |24 ORGANIC STRUCTURS. I Jt gives Nerves which proceed to the guttural region of the phavjnx and t® the palate: NEW NAMES. OLD NAMtS.. ■ ■ ■ ' ~ f PO^^-'"!!- ■ J To the nasal fossa^ : The gutturo-palatine Ther superior palatine The ethmoidal. .. .... . '. . Spheno-fdlatiKt. Naso-palatine. -<.#«. . . Jhi iame^ The third branch of the trifaeial distributes itself to the inferior jaw : C. The maxillary. , . • . .■ . Inferior maxillary. It gives two bundles : The superior one furnishes Isr, The siib-zygomatic; 2(1, the temporo muscu- ist. The massftrriji; 11, the UmporD-propuiUf ZS^ fcsr; 3d, Uiu bucco-labial ; 4th, the ptcrygo-m-js- the iu^'cal; itH, the ft/rygoHian. ealar.' The inferior gives lit, Tht cutaneous temporal ; 2d, the lingual, from lit, The superficial temporal ; 2d, the lingual ; 3i, iht which proceed tlie sub-maxillary ; 3d, the maxil- inferior dentary. io-dentar,. from which proceed the mylo-hyoi- uliaii and the dcntarj'. ..... NERVE WHICH PROCEEDS INTO THE ORBIT, A7td dlstrihutes itself to the inferior right tnuscle. VI Pair: it arises between the mesencephalon and the cerebral prolongation, in several filaments united : Exterior oculo-muscular. . . ' , • . Exterior motor. It reeeives Two filaments, which united to two others of the snperior cervical ganglion, form a plexus around the divisions of the anterior cere- bral artery. I NERVE WHICH DISTRIBUTES ITSELF TO Tllr. FACI, Passing through the splrold eanal of the temporal Lone, VII Pair : it issues from the triangular facet which is found be- tween the mesencephalon and the exterior side of the base of the jachidiaix prolongation : ThcUcial, . . ... ... Hard portion of the seventh pair. It TABLE OF THE NERVES. 225 It gives ffEW NAMES. ■ OLD NAMES. 1st, A pter>'soidian filament; 2d, filaments to the 1st, Superior ramus of the vidian ; small bones of the tympanum ; 3d, a t> mpanic Qd, The same ; ram:.-5, posterior auricular filaments, a sub- 3i, Cord of the tympanum ; temporal, milar ani mastoidian ramus, a stylo-hyoidian ramus, buccal rami, super-orbitar, labia], mentonniar and sub-men- .tonniar rami. NERVE WHICH PROCEEDS INTO THE EAR,, Distributing itself in the labyrinth: VIII Pair : it arises from the sides of the fourth ventricle, be- hind the seventh pair, with which it is connected by some filaments : The labyrinthic ■ Soft part of the sez'enth pair. NERVE WHICH PROCEEDS TO THE TONGUE, Distributing itself to the phari/nx, and to the muscles at the root of the tongue ; IX Pair: it originates in three or four filaments from the pe- dunculus of the rachidian prolongation, near the olivarian emi- nence : The pharynso-slossian Thegksso-pharyns.ianorbratichofth!eiihthfa:r, It gives The pharynsian NERVE WHICH PROCEEDS TO THE LUNGS AND THE STOMACH, Distributing itself along the necJc, in the thorax and abdomen : X Pair : it comes from the base of the rachidian prolongation, in ten or twelve filaments : The pneumo-gastric. .... . . Eiihth pair or jar vaptm. It furnishes On the neck, pharyngian rami, a laryngian ramus, filaments to th<; cardiac plexus, a tracheal ramus. The recurrent. To the ihorax, filaments for the pulmonary plexus; In the abdomen, rami to the stomach. . . The stomachic anterior and fOiterijr, NERVE WHICH PROCEEDS TO THE NECK AND THE TONGUE, DistriLuting itself to the muscles of the tongue, and to the greater part of those attached to the os hyoidesi XI Pair : it arises towards the middle of the furrow placed be- tween the pyramidal and olivarian eminences : Hyo-Blo:sian. . . , . . , . Ninth^air, ^teat h/pogbssus, lisgua!. VOL. I. Q If 226 ORGANIC STRUCTURE. It ftirnishes A branch for the plexuous arch of the hyoidians — a hyo-thyroidian ramus — filaments to the muscles of the tonuiue. KERVE WHICH DISTRIBUTES ITSELF TO THE STERNO-M AST 01- DIAN AND THE DORSO-SUPER-ACROMIAN : Xll Pair : it arises towards the middle of the trarJielian region, and the sides of the rachidian prolongation, by several distinct fila- ments, -which unite into one trunk : this trunk enters into the cra- nium through the occipital hole, and issues through the posterior foramen lacerum : NEW NAMES. OLD NAMES. Traclielo-dor.-al. .... . . . Sfinal, accessory of iVillii- VERTEBRAL NERVES. These arise from the cerebral prolongation, by a great number of filaments, united into two bundles, the posterior of which is larger than the anterior ; both proceed to a ganglion, from which one trunk of nerves then issues ; this nerve soon divides itself into two branches, which issue through the foramina of the rachis. The posterior branches proceed to the spinal face of tlie rachis, and dis- tribute themselves to the muscles and the neighbouring integu- ments. The anterior branches, soon after they issue from the vertebral foramina, unite with each other by filaments of communi- ■cation, and all of them send one or more filaments to the tri- splanchnic. NERVES which PROCEED TO THE NECK. The eight cervical pairs. Distributing themselves to the 77iuscles and the integume>its of the spinal Jace of the neck : The posterior branches. Which distribute theinsehes behind the head and the neck : The anterior branches. These furnish 1st, The nerves which form the trachelo-cutanecus plexus, and which are : the branches of the second, third, and fourth cervical pairs. 1 This TABLE OP THE NERVES„ 227 This plexus furnishes, 1st, mentonniar, zygomato-auricular, and occipito-auricnlar branches ; 2J, sternal, acromian and scapu- lar branches. 2d, The rami which form the diaphragmatic nerve, and "vvhich -are : the anterior branches of the fourth, fifth, and sometimes the sixth cervical pairs. 3d, The nerves which form the Irachial plexus, and which are ; the anterior branches of the last four cervical pairs, and of the first lumbar. The nerves which proceed to the thoracic member arise from NEW NAMES. OLD NAMES. The brachial plexus. . . i ... The same. This plexus divides itself into nine branches or bundles. 1st. nerve which proceeds to the shoulder. Distributing itself to the siiper-scapido-trochiterian, and dorso-super- acromian muscles : The fuper-scapular The same. 2C. NERVES WHICH PROCEED TO THE STERNAL FACE OF THE THORAX, Distributing themsehes in three filaments ^ Isf, to the sterno-hume- rian ; 2d, to the costo-coracoidian ; 3d, on the side of the thorax : Tlic scerno-thoracic. ...... Thoracic. 3d. nerves which PROCEED TO THE SHOULDER AND THE ARM, Di.<;triduting themselves hy three filaments : 1st, to the costo-scapic- It'atz; 2d, to the hunho-humerian and the siih-scapiilo-trochinian ; 3d, to the scnpulo-humerian, and the inferior super-scapulo- trochiferian tnuscles : The sub-scapular. ^- . . . . . The sam. '4TH. NERVE WHICH PROCEEDS AROUND THE SCAPULAE. ARTICULATION OF THE HUMERUS, Distributing itself : 1st, to the sub-scapulo-trochinian muscle ; <2d, to the sub-acromio-humerian ; Sd, to the radial edge of the ar?n : ILe :i-3pulo-tiumciian. . . • , . Axillary cr circumflex. 5TH. NERVE WHICH PROCEEDS TO THE ARM ANt» FORE-AP.M, Distributing itself superficial!!/ to the interior side of the arm, alon.g the radial edge of the fore-arm as far as the thumb * Die radio-cutarieoui. ^ MuKulo-eutrnisiui. <^ 2 6th. nep.ve •228 ORGANIC STRUCTURE. 6th, nerve which proceeds to the arm and fore-arm. Distributing itself superficially/ along the cubital edge of the arm and fore-arm, as far as the little finger ; NEW NAMES. OLD NAMES. The cubito-cut2neoii5. ..... Intiriar cutaneous. 7th. nerve which proceeds to the arm, the fORE-ARM, AND HAND, Distributing itself a7^ound the humei'us, behind, along the radial edge of the fore-arm y on -the conxexity of the liand, and to the first three fingers : The radio-digital. . . . . - . Radial, 8th. nerve which proceeds to the arm, the FORE-ARMy AND THE HAND, Distributing itself to the interior side of the arm, behind the articu- lation of the elbovj ; along the cubital edge of the fore-arm, and to the ixvo last fingers : The cubito-dijital Cubital. 9TH. NERVE WHICH PROCEEDS TO THE ARM, THE FORE-ARM, AND THE HAND, Distributing itself to the interior of the arm, before the articulation of the elboxo ; on the middle of the palmar face of the fore-arm f and to the fingers : Digital median, , . , . . . . Mf.Har, NERVES WHICH PROCEED TO THE DORSAL REGION, AND TO THE sides of the THORAX AND ABDOMEN : The twelve dorsal pairs. Which distribute the?nselves to the integuments of the back: The posterior branches. Which pass bettveen the ribs, and distribute themselves before the thorax, to the muscles and integuments of the abdomen : The anterior branches. NERVES WHICH PROCEED TO THE LUMBAR REGION, TO THE SIDES OF THE ABDOMEN, AND INTO THE PELVIS : The five lumbar pairs. Which distribute themselves to the muscles of the lumbar region and to the integuments of the nates i The posterior branches. Which TABLE OF THE NERVES. 22g Which distribnte themselves to the sides of the abdomen and the pelvis, and furnish some to the crural plexus : The anterior branches. NERVE WHICH TRAVERSES THE PELVIS, AND PROCEEDS TO THE REGION OF THE GROIN : . The anterior branch of the first lumbar pair. It gives, 1st, A ramus wb.ich proceeds to the muscles of the abdomen and to the pubis, suh-pubian ; 2d, a ramus to tlie integuments of the groin; 3d, a ramus which follows the contour of the ridge of the ilium, and proceeds to the scrotum, along the testicular cord. NERVES WHICH ISSUE FROM THE OS SACRUM: The five sacral pairs. Which distribute tliemsehes to the nates and to the anus : The posterior branches. Which supply the hypogastric and crural plexus, and distribute them- selves to the coccijx and anus : The anterior branches. NERVES WHICH FORM THE HYPOGASTRIC PLEXUS: The anterior branches of the fourth and fifth sacral pairs, and filaments of the tri-splanchnic. The7/fu?mish The pelvian, which are distiibuted to the bladder, the rectum, and the genital parts contained in the pelvis. NERVES WHICH, FORM THE CRURAL PLEXUS: The anterior branches of the last four lumbar pairs, and of the first four sacral. The nerves distributed to the pelvian memljers arise fom the crural plexus, which divides itself into two other plexus : 1st, the lumbar plexus ; 2d, the sacral plexus : tliey each furnish four prin- cipal branches to the pelvian member. y 3 NT." VLS 230 ORGANIC STRUCTURE. KERVES ARISING FROM THE LUMBAR PLEXUS: 1st, Nerve which proceed'^ to the thigh, Passing near the anterior tubercle of the ilium, and sending out ram i^ jications to the exterior part of the thigh and the leg : NEW NAMES. OLD NAMES. The interior cutaneous. . , ... Crural. 2d. Nerves which distribute themselves in three, four, or five filaments to the glands, the integuments of the groin, and to the genital parts : The insuinal. . , , . , ; . Crural, Sd. Nerve which proceeds to the thigh, the leg, and the fqot. Descending behind the pralumho-trochantinian muscle, passing through the crural arch, and distributing itself to the integuments of the anterior par't of the thigh, and by a long ramus (tibio-cuta- iieous) to the muscles and integuments of the leg and the foot : T!ic fcnoro-proEtibial. ..... Crural. 4th. Nerve which proceeds to the interior part of the thigh, passing through the sub-pubian hole, and distributing itself to the 7nuscles and integuments of the anterior and interior part of the thigh : The sub-pubio-femoral. ..... Obturator. NERVES AiflSING FROM THE SACRAL PLEXUS : 1st. Ner\-e which proceeds to the penis and the clitoris, Passing through the ischiatic notch, and distributing itself to the perinawn, to the penis, and the ciit oris : Ischic-pejiiarij ischio-clitoriun. . . . . Pullzal, 2d. Nerve which proceeds to the nates, Distributing itself to the sacro-femoral muscle, the ilio-troehanterian fnuscles, and to the integuments of the thigh : The fes.*o branches: one (the }^tTom.2L'c) furnishes ; \sf, the peroneo- cutaneous ; 2d, the pi'atibio-digital and the fratihio-sub-plantar. The other (the tibial) gives; 1st, the digital peroniar ; 2d, the tlbio- plantar. On the anterior and lateral parts of the bodies of the vertebrae, from the base of the cranium to the sacrum, is a series of ganglions united by a nervous cord : The ui-splanchnic. . .' Ths great sympathetic or intercostal nerve. The ganglions of the cervical region are : 1st. A superior, very large and fusiform. 2d. A middle one, very small, and often even a simple plexus. Sd. Aninferior, round, and situated before the neck of the first rib The ganglions of the other regions are disposed in such a man- ner, that they correspond nearly to the sides of each inter-verte- bral space. The nerves which communicate with the tri-splanchnic are : Some encephalic pairs, and the thirty rachidian : they each send to it, one, two, three, four, and even five filaments. The nerves furnished by the tri-splanchnic, either from its gang- lions or the cord which unites them, are distributed chiefly to the arteries of the organs contained in the three large cavities. In their distribution these nerves form, or contribute to form, a great number of plexus ; such as, 1st. The plexus, which surrounds the trunk of the anterior ce- rebral artery, in its passage along the inflex canal of the temporal. 2d. The plexus cardiaci. From the fifth to the eleventh intercostal space the tri-splanchnic gives five or six filaments, which unite and form the great splanch- nic : this nerve penetrates into the abdomen, and proceeds to. an assemblage of ganglions, situated on the aorta and the pillars of the diaphragm : Ottliic ganglion Seni-tanar, q4 The 232 ORGANIC STRUCTURE. The numerous filaments which proceed from the coeliac ganglion form, on the aorta, the divisions of the opisto-gastric artery, and on the superior mesenteric artery, a considerable plexus. NEW NAMES. OLD NAMES. Cctliac plexus. . . ..... Solar. The filaments which proceed from this plexus are distributed on the trunks of the principal arteries, surround them, and form on them the s:astr'ic, hepatic, splenic, mesenteric, &c. plexus, and ac- company them in all their distributions. Towards the eleventh and twelfth dorsal vertebrff!, the tri- splanchnic gives also one or two filaments, which constitute the small splanchnic : this nerve penetrates into the abdomen, unites to others of the coeliac plexus, and forms a plexus around the renal and spermatic artery. The tri,-splan'.hDic penetrates into the pelvis, ^i^d unites on the sacrum with that on the opposite side. SYSTEM [ 233 3 SYSTEM OF THE SENSES. OF VISION. 27 ^* The organ of fight confifts efientially of a very thin membnane, placed at the bottom of the eye. This membrane has the property of per- ceiving the adiion of the light refle(?led from ex- ternal bodies^ and refracfted on it by the tranfpa- rent liquids of the eye. The eyes feem to be wanting in zoophltes, feveral of the articulated worms, in fome of the larvae of infe6ts, and in the acephala mollufca. All vertebral animals have two eyes, fituafed in the cavities of the bones of the face*. They are compofed of two or three tranfparent fluids of dif- ferent denfities, placed the one before the other, each of which is contained by a thin pellicle ; and they are covered at the back by the fine mem- brane which perceives the contadi of the light. This apparatus is contained in a common, thick, double covering, for the mod part blackifh in the infide, having in it an aperture fufccptible of contraction, before which is placed a membra- nous and tranfparent fegment of a fphere. *■ The eyes of infe6ls are very numerous, and have a peculiar ftrudture, which cannot be known on account of their fmallnefs. This 234 ORGANIC STRUCTURE. This organized dioptrical apparatus is nearly round in all animals j but its anterior part, more or lefs truncated, is covered by a tranfparent Ipherical fegment called the cornea, 1^1. The cornea is nearly round; but its tranf- verfe diameter is often fomewhat longer : it is com- pofed of laminae cemented together, and is thicker in the middle than at the edges. In fidies, the cetacea, and the greater part of reptiles, it is very much flattened; fo that the eye of thefe animals is often a hemifphere. In the terreftrial mam- malia the cornea is a fhort fegment of a fphere, fmaller than that of the eye ; and this organ is nearly round. The cornea of birds is a narrow bemifphere, and the eye has a greater degree of projection. Behind the cornea, and before the eye, the mammalia and birds have an aqueous humour, fome- what faline, the denfity of which is nearly equal to that of water : it feems to be contained in a very thin pellicle. This aqueous humour is often wanting in fifhes % the fmall quantity of it found fometimes in the eye of thefe animals is always vifcous : it does not exift in the fepias. 2 73- CryJialUne humour. Behind the aqueous humour, in the mamraaha and birds, and behind the cornea of aquatic aninjals, is a thick, diapha- nous fubftance, heavier than water, and enveloped^ 9 without SYSTEM OP THE SENSES. 235. without adbefion, in a very fine membranous capfule : the cryjtalline humour. In fifties, the cryflalh'ne humour is nearly round ; in the mammalia it has a lenticular form ; it is more flattened in birds, and ftill more fo in man. In all thefe animals this body is formed of two fpherical fegments, the pofierior of which is fmaller. Its convexity is in the inverfe ratio of that of the cornea. The cryf^alline is harder, the greater its convexity ; and its confiftence decreafes from the centre to the circumference. '1"}^. Vitreous humour . The cry ftalline is lodged in a depreflion prefented to it by another body of lefs deniity, having the confifi:ence of the white of an t^gi which occupies the greater part of the bottom of the eye, and is enveloped by a thin diaphanous membrane. This fluid is called the vitreous humour. From the tranfparent membrane (memhrana hyaloideajy which envelops the vitreous humour, proceed internally a great number of thin lamina}, which crofs each other in ditFerent directions, and form lenticular cells containing the vifcous fluid. The denfity of this fluid, which conftitutes the vitreous humour, is greater than that of the aqueous humour, and lefs than that of the crydallinf. This fubHance hardens and becomes opake by heat and by alcohol. The anterior furface of the vitreous humour is feparated into two lamincC. one of which paffcs behind 236 ORGANIC STRUCTURE, behind the cryflalline, and the other is refle6^cd on its fides. 275. The retina. The vitreous humour is en- veloped, behind and on the fides, by an exceed- ingly fine membrane, of little confiftency, which receives the impreffion of the rays of light. It is only an expanfion of the optic nerve; and is di- ftinguifhed by the name of the retina. The retina expands difTerent ways on the vi- treous humour ; but in all animals it terminates around the aperture of the cornea, and does not pafs before the cryftailine. The aque^s humour, the fubflance of the cryftailine, and the vifcous fluid contained in the cells of the vitreous body, are fecreted by the fiir- faces of the membranous capfules which contain them. Thefe three tranfparent bodies are contained in a common double covering. 276. Sclerotica. The exterior tunic, called the fclerotica, is foft, thin, and of a clofe texture in the greater part of the mammalia ; and the eye has that round form which all fluids contained in a flexible membrane naturally tend to afRime ; but among animxals, in which the configuration of the eye is different, the peculiar form of that organ depends on the greater thicknefs or firength of this tunic, or on its being fupported by fmall bones. This external tunic of the eye is hard and very ' 6 thick SYSTEM OP THE SENSES. 237 thick in fbme aquatic animals, fuch as the whale, and particularly the flurgeon, where it is thicker than the reft of the eye : in other fiflies it is thin, but has the confiftence of a cartilage. In birds this tunic is thin, and is divided be- fore, into two fmall leaves ; between thefe leaves are received fmall ofleous laminae which lap over each other, and form a circle or oval, proper for fuftaining it. The fclerotica has always in the fore-part. a large aperture, fhut by the cornea, v^'ith which it is united: behind, it affords a pafThge to the optic nerve by a hole or canal.. 277. The choroid memhrane. The interior face of the iBierotica is lined by another thin tunic, called the choroid, united to it by a lax cellular tiffue : before, it adheres with greater force around the aperture which covers the cornea, where there is a whitifh vifcous and foft ring, called the ciliary Tigamenf. The choroid membrane is then turned forwards, • and feems to conftitute two folds : one of them, exceedingly thin, is folded back and difpofed in the form of radii (the ciliary p-ocefs) ; it palTes before the vitreous humour, in which it is deprefled into a hollow, and terminates on the edges of the cryftalline humour. This arrangement does not take place in fiflies : the other fold, called the 238 ORGANIC STRUCTURE, uvea, proceeds more forward, and forms an ex- panfion pierced with a hole, known by the name of the ^uj)il : it is covered before with a fibrous membrane which adheres to it clofely, and whieh prefents ftrise of different colours : the iris*-. The iris has the property of dilating and con- trading, for the purpofe of diminifliing or enlar- ging the pupil. In the flate of dilatation, the pupil is generally round ; but when the iris dilates, this aperture, in contra6ling, often changes its form. In man^ apes, many carnivorous animals, and birds, it retains its round form ; in cats it aflumes that of a lozenge, which goes on always con trading, and at length becomes a vertical line. In the horfe, the ox, and the whale, it afFeds the fame form, but in a tranfverfe diredion. The pupil of fome animals is fafceptible alfo of afTuming various peculiar configurations, by the expanfion of the iris. The ray and the torpedo have at the top of the h'is a membrane, folded back, which is fufccpti- bie of falling down, fo as to fhut the pupil by expanding itfelf like a curtain or blind. This peculiarity has been remarked in thefe animals only. * In moft of the mammalia the iris has a brown colour j and" in birds it is ornamented with the moft beautiful tints. It SYSTEM OF THE SENSES, 23^ It has been obferved, that the pupil in the foetus of feveral animals is fhut by a thin membrane {the fuplJIary), which is deftroyed and difappears Ibme time before birth. All this part of the choroid, which forms the uvea and the iris, is generally plane, and fome- times a little convex 5 between it and the cornea the aqueous humour is lodged. The choroid, which is much thinner than the Sclerotica, is formed o( a beautiful vafcular tifTue : in the mammalia a part of it is often coloured. In fifhes with fixed branchiae it is compofed of two diftindl laminae, between which is a foft glan- dulous cylindric body, very often of a beautiful red colour. The choroid is generally covered with a blackifh, mucous fubilance, often very thin, which does not prevent from being feen through it the beautiful colours with which in fome animals * it is orna- mented ; it covers alfo the interior face of the ciliary procefs of the uvea. This blackifh, mucous fubftance, which is found in almoft all animals, is wanting in the Albinos, among whom a ftate of whitcncfs is always a difeafe. * In the ox this part is of a beautiful golden green changing to ikyrblue ; in the horfe, the goat, the cervine antelope, and the ftag^ it is a filvery blue changing to violet ; in the fheep, of a pale golden green, and fometimes blueifh ; in the lion, the cat, and the dolphin^ it is of a pale golden yellow ; In the dog, the wolf^ and the badger, it is pure white,, bordered with blue. 278. TU 240 ORGANIC STRUCTURE. 2,78. The orhlts. In all animals the eyes are placed in cavities of greater or lefs depth, called the orbitar foffe, or orbits. The orbits of fifhes are not very diflindl, being confounded externally with the temporal fofTse; the inferior edge of them is formed of one ofleous piece, or a feries of fmall bones, which, for the moft part, are five in number. In birds they conlift in a depreffion interrupted at the bottom. In reptiles the exterior edge of the orbits is formed, in whole or in part, by an offeous branch, behind which the temporal foffe are confounded with the orbitar. In quadrupeds 'is obferved alfo an afcending branch, which forms, on the outfide, a complete border to the orbits ; but behind this border the exterior fide is wanting, and the orbitar and the temporal foffie are united. In man and apes thefe orbits are complete. In apes, and in man, the edges of the orbits are formed by the frontal, the fuper-maxillary, and the jugal bones. The fides are formed by the fron- tal, the ethmoid, the lacrymal, the palatine, the fuper-maxillary, the jugal, and the fphenoid : the cavity which they compofe is a cone, the apex or point of which is turned backwards. The orbitar foffe fometimes are feparated from each other only by a thin lamina, which is often in part wanting, as is the cafe in fifhes; but, in general, they are feparated by the noflriis. Thefe SYSTEM OF THE SENSES* 241 Thefe foflae, in moft animals^ are turned more or lefs towards the fides; in fome reptiles and fe- veral fifhes* they are turned upwards, and in the quadrumana and man they look almoft dire6lly forwards. The eyes reft in the orbits on a fmall ciifhion of - very fine fat, which permits them to glide with great facility; this cufliion is thicker in the mam- malia, the orbits of which are deeper. In fiflies it is formed of a gelatinous fubftance, enveloped by a lax cellular tifTue ; but in the ray and fhark the eyes are articulated on a cartilaginous pedun- cle, v/hich is itfelf articulated in the bottom of the orbit. 279. Mufcjes vf the eye. The eye, in man, apes, birds, and fifhes, is moved by fix mufcles ; in moft of the mammalia by feven. Thefe mufcles are fixed to the bottom of the orbit, and are at- tached before on the contour of the fclerotica. Of thefe mufcles, four 2ive.Jiraighty and placed on the four fides of the orbit ; the two others have au oNlque dire61ion. 280. Eye-Jids. In fifbes, ferpents, and fome other reptiles, the eye-lids are wanting; lizards and the moon-fifh have one of a circular form ; in the mammalia there are two. Birds and feveral reptiles have three; in thefe animals the third eye- * The pleurone6les have the two orbits on the fame fide of the head 5 but only one of them is perfeft. VOL. I. R lid^ 242 ORGANIC STRUCTURE. lid, 7nemhrana niBltans, is found towards the in- terior angle ; it is thin, and extends over the eye like a curtain, by a I'^'ufcular arrangement altoge- ther peculiar: this tine eye-lid permits birds to fee through it, and to look at the fun. The mammalia, towards the interior angle of the eye, inftead of the third eye-lid of birds, are furnifhed with a membranous fold, which in man and apes is very fmall, and has the form of a cref- cent. In the other mammalia it is larger, femi- lunar, and porous ; and in part of its extent has the appearance of a cartilaginous lamina. The eye-lids feem to be merely a prolongation of the fkin : the interior face of them is reflected on the eye, and forms the white of it, called the membrana conjunB'iva. The eye-lids are clofed by the a6lion of a muf- cle, called the ^alpehralian, the fibres of which are difpofed in the form of radii: in birds they have alfo in their thicknefs a membrane which lines the infide of the orbit. In moft of the mammalia, the edge of the eye- lids is terminated by a cartilaginous circle, the tarfus, furnifhed with hair, eye-lajbes. In thefe animals the upper eye-lid is the largeft^ and, by falling down, covers the greater part of the eye : it is raifed by a particular mufcle called the^^^/vor orhtO'palfiehralian,^ which is attached in the bot- tom of the arch of the orbit. The SYSTEM OF THE SENSES. 113 The eye lids of birds have, in general, a con- trary difpofition : the lower, which is the largeft, is Jined by an ovoid, cartilaginous membrane, and by rifing up covers a great part of the eye : it is pulled down by a particular mufcle, which is attached to the bottom of the orbit. The eyes of animals which live in the air are continually moiftened by a fluid (tears) fecreted from a particular gland, called the lacrymal. This gland is fituated at the top, and often around the' orbit ; it is of a white or grayifli colour, granulated, and often divided into feveral follicles, from which proceed very fine fmall du6ls. having their aper- tures near the interior edge of the eye-lids. The liquid which diftills from them, being dif- fufed over the eye, ferves to moiften it; and the fuperfluous part, proceeding towards the interior angle of the orbit, is in general abforbed by two fmall tubercles, having each at the fummit a filtu- lous hole, fiin6ia lacry mafia : thefe holes are con- tinued in two fmall du6ls, which proceed to a common excavation, Jaccus lacrymalis^ and termi- nate at the nofe in a particular du61 called the najal. In fome fpecies, fuch as hares and rabbits, the tears trickle down by a finuous and cartilaginous furrow, which ends in a common bag furnifhed with a valvula. In the mammalia the edge of the eye-lids is furniflied alfo with a great number of glandulous R 2 , follicles^ 244 ORGANIC STRUCTURE. follicles^ which fecrete an un(5tuous humoor calfed the gum of the eyes. Birds^ and fome of the mam- malia, towards the interior angle of the orbit, have a fmall gland of a different formj which fecretes a thick humour, yellowifh in birds, and whitifh in the mammalia. This gland, called that of Har- deruSj pours the produ6l of its fecretion through an aperture fituated behind the third eye-lid, or its reprefentative : it is the largeft of thofe of birds. The mammalia have alfo, towards the nafal angle, a fmall reddifh body confifting of feveral glandulous follicles, caruncula lacrymaliSi which fecretes a thick and whitifh humour,' 281. Inmariy the eyes are placed in complete orbits, which have the form of a cone irregularly flattened ; the bafe forms the entrance, and the apex is turned iny/ards and backwards. At the bottom of the upper iide is the ocular foramen, and below and within this hole is the fphenoidal Jiffura. Along the exterior and inferior angle is the fpheno-maxUJary filTure, before which is the fub-orbitar canal : towards the interior angle is obferved alfo Xhtfojfa lacrymalis. The orbitar fofla prefents alfo fmall holes, which afford a paffage to veiTels and nerves, and an ex- cavation which lodges a gland. ^ The eye has a round form, Ibmewhat deprefled at the top, at the bottom^ and on the fides. The SYSTEM OP THfi SENSES. 24& The cornea is the fegmcnt of a fmaller fphere than that of the eye; it is tranfparent, cartilagi- nous, and unites with the fclerotica by an edge cut externally into a bevel. The aqueous humour, which is diaphanous, and nearly of the fame denfity as water, is lodged be- tween the cornea and the iris ; and a fmall quan- tity of it is contained even between the iris and the cryftalline. It appears to be fecreted and con- tained by an exceedingly fine membrane. The cryftalline is lodged behind the aqueous humour, in a depreffion of the vitreous body. It is lenticular, and more convex behind ; it is from four to five lines in diameter, and fomewhat more than two lines in thicknefs ; it is rounder in the foetus, and with age becomes flattened, yellow, and muddy. Its denfityjncreafes from the centre to the circumference; it appears to be formed of fibrous ftrata, placed one above the other : the capfule, by which it is enveloped without adhering to it, is moiflened by a humour called that of Mor' ^agni, which is rarer before. The vitreous humour fills up behind, the greater part of the eye; it occupies l.\. of its whole axis ; while the cryflalline occupies only -'-, and the aqueous humour .^^r its denfity is greater than that of water. The hyaloid membran': which retains it, gives birth to a great many expanfions; and thefe bj R 3 crofling 246 ORGAtnc STRUCTURE, crolling fonn regular cells which communicate with each other. The hyaloid membrane is divided before into two laminas, one of which pafles behind the cry- flalline humour; the other proceeds before, under the ciliary procelles, which imprint on it blackifh Uriae, and is confounded with the anterior part of the capfule of the cryfiialline. The circular fpace comprehended between thefe two laminae forms, on the edge of the cryftalline humour, a triangular plaited canal, which is rendered apparent by forcing into it fome bubbles of air. The retina, which is an expanfion of the optic nerve, is a membrane of a white a(h colour, and of a foftifh confiftence : it feems to be fibrous and vafcular without, and villous within.. Soemering difcovered in the potterior part of the retina a very fmall hole, the edges of which are plaited, and of s. jellow colour: this hole, which has been found in the ape, has not been obferved in other animals. , The retina, after enveloping the whpole vitreous humour, continues before, and terminates behind the ciliary procefles. The humours of the eye are contained in two membranes. ifl. The fclerotica, which is thick, white, and of a clofe texture ; it ha^ before a large aperture^ dofed by the cornea with which it is united. Behind^ SYSTEM 0-P THE SENSES. 247 Behind, it affords a paiiage to the ocalar nerve, and is traverfed in an oblique dire6lion, through - out the reft of its extent, by vefiels and nerves. id, The choroid, which covers the fclerotica internally : it appears under the form of a delicate membrane of little confillence, compofed of an exceedingly fine and beautiful vafcular reticula- tion. Its exterior face, which is reddifh, adheres to the fclerotica by a loofe cellular tiffue, and by vellels and nerves: the interior face, which is vil- lous, is covered with ablackifh coating j it is ap- plied to the retina, and affords a paffage to the ocular nerve. ^efore, the choroid is confounded with the fclerotica around the corneaj where the ciliary li- gament is found, under the form of a thick fpongy and whitifh ring. Below this ring arife the ciliary procefles, which appear to be a continuation of the choroid. They expand before the vitreous humour, and terminate in a- ferrated border around the cryftalline. The ciliary proceffes are compofed of a radiated membrane, fmooth before, and plaited behind ; the projeclionsof the folds are covered with a blackilh fubftance, like the choroid, and are depreffed into a hollow before the vitreous humour. This black- \ii\ fubilance, or varnifh, remains on the vitreous humour when the ciliary procelfes are removed, R 4 and ''i4S ORGANIC STRUCTXyRE, and produces a radiated ring formed of black UricC. Before the ciliary ligament is the commence- ment of the uveous membrane, which feems to be a continuation of the choroid ; it proceeds dire6liy towards the axis of the eye, before the ciliary pro- cefles, and produces the aperture of the pupil. Behind, this membrane is compofed of fibres, which converge from the ciliary ring towards the pupil ; it is covered by a blackifh varnifh, in the fame manner as the choroid. Before, it is covered by numerous fmall fibres, difpofed in the manner of rays, and which form the iris. Thefe fmall fibres are fufceptible of elon- gation and contradlion, for the purpofe of con- tra6ting or enlarging the aperture of the pupil : this contraction, in all probability, is produced by a fpontaneous afflux of blood towards that part. In man, the colour of the iris varies. The eyes may be covered by both the eye-lids: the upper, which is larger, is a continuation of the eye-brows. The eye brows reft on the fuperciliary arch ; they are formed by the fuperciliary mufcles, and by the fkin, which in that part is covered with hair: thefe mufcles deprefs the eye-brows by bring- ing them nearer to each other. The eye lids are formed by the fkin and the pal- pebral ian SYSTEM OF THE SENSES. • 24^ pebralian mufcle; the upper one has alfo a proper elevating mufcle. When the eye-lids are broaght together, the upper one falls down as four, and the lower one riles as one. The free edges of the eye-lids are furnifhed with a cartilage {tarjus), retained by a thin liga- ment, the palpebral. The tarfian cartilage has a great many farrows, which ferve to lodge glandu- lous follicles, called the glands of Mclhomius : they are of a yellowifh white colour, and fecrete the gumaiy matter. This thick, undluous humour iflues through a great number of pores called the ciliary pores. Towards the interior commifTuraj behind the membrana nicStltans, is a fmall, red- difh, conical, villous tubercle, compofed for the moft part of feven fmall follicles, cemented two and two, with an odd one called the canmcula lacrymalis. The edges of the eye-lids are furnifhed with <6ne or two rows of eye-la(Iies. The fliin, which forms the eye-lids, feems to be refle6ted interiorly, and then to be continued be- fore on the lobe of the eye, to form the conjunc- tive membrane, which appears to be continued as far as the cornea. The conjun61ive membrane is formed of a clofe, fmooth tiflue, tranfparent on the cornea : it conti- nually fecretes a ferous matter, which moiftens the eye. Towards the interior commifTura of the eye- lids ^50 ' ORGANIC STRUCTUKE. lids is a fold in the form of a crefcent, which con-« ftitutes a fort of membrana niditans. The lacrymal gland, which is oblong and of a whitifh gray colour, is fituated in the upper part of the orbitj towards its exterior angle, and is ap- plied on the lobe of the eye. The excretory du6ls of this gland open on the interior face of the up- per eye-lid near its edge, where there are fix or feven fmall apertures, which pour the tears upon the eye, Thefe tears are difFufed over the whole furface of the eye by the a6lion of the orbit oal- pebralian mufcle, which always brings them back towards the interior commiflura. The free edge of each of the eye-lids has, to- wards its interior angle, a small hole, the projed:- ing mouth of which is cartilaginous. When the eye-lids are ihut thefe two holes correfpond ; they are continued by two duels, which proceed to the face us lacrymal is. This membranous bag is lodged in the lacry- mal fofla, and is continued inferiorly with the nafal canal. The eye is moved by fix mufcles, four of which are Jiratght and two oblique. The four ftraight mufcles are attached to the bottom of the orbit, at the circumference of the ocular hole; they then proceed forwards, and each terminates in a broad tendinous expanfion on the fclerotica, embracing the contour of the globe of the eye before. Of SYSTEM OF THE SENSES. 251 Of thefe four mufcles the fuperior elevates the eye ; the inferior deprefles it ; the interior pulls it inwards, and the exterior outwards. Of the two oblique ones, the fuperior, which is larger, is at- tached to the bottom of the eye, on the interior edge of the ocular hole, and changes to a tendon, which is refie61ed on a cartilaginous lamina fixed at the interior angle of the os frontalis. This ten- don receives in that part a Iheath, and then pro- ceeds outwards, paflTing over the fuperior right mufcle, and is fixed at the top and behind to the exterior fide of the eye, which it caufes to turn in- wards and downvv^ards. The inferior oblique mufcle, which is fmaller, is attached to the bafe of the orbit, and to the bot- tom of the lacrymal canal ; it proceeds backwards in an oblique direction, paffing under the depref- for, and fixes itfelf to the pofterior and exterior part of the eye, which it pulls upwards and outwards. OF OF HEARING. 282. The organ of hearing is obferved in fome of the mollufca, fome infeds, and in all vertebral animals. It confifts chiefly of a membranous cap- fbile, containing a vifcous fluid, in the middle of which the auditory nerve is expanded. This capfular apparatus is found in the ear of all animals : in crabs, and the fepias, it exifls alone. In fifhes, the capfule, for the mod part, is di- vided by leveral contra61:ions ; and contains one, two, or three fmall bones of different denfities^ fufpended in the middle of the vifcous fubftance. In thefe animals are obferved three bent or femi- circular veflels, the extremities of which proceed into the interior of the capfule. In the greater part of fifhes the whole of this . apparatus is contained in the common cavity of the cranium, which has depreflions fitted for receiving it, and fome furrows which ferve to lodge a part of the femi-circular vefTels. In fifhes with fixed branchiae, this apparatus is contained in the thicknefs of the bones of the cranium, and the capfule communicates with a- eonduit, which traverfes the cranium behind. The aperture of this conduit is covered, on the out- fide. SYSTEM OF THE SENSES. 253 - lide, by a thin membrane called the ^nmibrana tym^ani, and by the Ikin, The fpace comprehended between this membrane and the capfule is called the barrel or tympanum. In fome reptiles the barrel is covered only by the Ikin: in the greater part of them it is furnifhed with a membrana tympani ; and in all, the fala- manders excepted, this cavity communicates with the capfule by an aperture covered with an ofTeous plate, and with the mouth by a particular conduit called the guttural conduit, Eujlachian tube. In birds, the femi-circular veflels have not a direct communication with the membranous cap- fule, but with an intermediate cavity, known by the name of the vejiibulum. In thefe animals, the capfule exhibits alfo a conical prolongation, beat and divided by a membranous partition into two cells, which communicate with each other: one of thefe proceeds into the vejiibulum, and the other into the barrel or tympanum, by an aperture co- vered with a very fine membrane. In the mammalia, the organ Vv'ith two cells is formed of two fcalce, divided by a membranous or ofTeous partition : thefe fcala3 make feveral turns in a fpiral form, fo as to reprefent the fhell of a fnail, and on this account are called the cochlea. The membranous apparatus of the cochlear the femi-circular canals, aud the vejiibulum, conilitute the MEMBRANOUS LABYRINTH. In 254 ORGANIC STRUCTURE, In fifhes with fixed branchice, this lahyrinih is enveloped by an ofleous ftratum of the fame form, very wide, and in the middle of which thefe parts are free, and as it were fufpended. In moft reptiles, in birds, and in the mammalia, it is en- veloped by a thin ofleousftratum, which embraces it clofely, and feems to have been moulded on the form of its parts : this folid covering conftitutes the ojfeous lahyrinih. In young mammalia, the ofieous labyrinth may be difengaged from the body of the bone, which only furrounds it ; but in adults it is confounded with it, and feems hollowed out in its fubltance. The veflibulum is pierced with Xqsqw apertures, viz. five for the three femi-circular canals, becaufe the two extremities of one of thefe canals unite before they reach the veftibulum ; the fixth com- municates with one of the fcalae of the organ isjtth f"iVo cells, or of the cochlea ; the feventh opens into the barrel or tympanum, and is co- vered by a fmall bone. \n moil of the mammalia, the cochlea makes two turns and a half in a fpiral form ; in cetaceous animals it is fhort and plane, and makes only one turn and a half; in the Guinea pig, the cabiai, and porcupine, it is of a turreted form, and makes three turns and a half. The barrel or tympanum, which is very large in birdSj is extended in the thicknefs of the bones of the cranium. In the mammalia, it is for SYSTEM OF THE SENSES, 255 the mofl part fingle, but fometimes divided^ as m the s:enus of the cat, and of the civet-cat. In moft reptiles, in birds, and in the mamma- lia, the barrel has four apertures : one is that of the veftibulum, which is covered by an ofleous plate ; the fecond is that of the cochlea, which is clofed by a membrane ; the third is the hole or canal that communicates with the mouth ; and the fourth, which is larger, condufis to the out- iide, and is fhut by the membrana tympani. AH thefe apertures are of different iizes, and alTume different forms and diredlions, in the different kinds of animals. In fifhes with free branchiae, and in falaman- ders, the membrana tympani, and alfo the barrel, are wanting. This membrane, which in fifhes with fixed branchise, and in fome reptiles, is on a level with the head, is covered only by the fkin, which in that part is often thinner. Birds, and particularly the mammalia, have the membrana tympani placed at a greater depth in the thicknefs of the bones: in thefe animals it is fixed on an offeous frame, fufceptible at a young age of being feparated from the body of the bone: this frame, for the mofl part, is incomplete. The membrana tympani affects alfo different forms : in fpme reptiles it is plane ; in birds it is convex on the outfide, and in the mammalia it projed^s inwards. The 256 ORGANIC STRUCTURE. The ofleous plate which covers the aperture of the vefiibulam is for the moll: part continued in a handle, or feries of fmall bones, one of which is applied to the membrana tyrapani, to which it ad», Beres. Salamanders, on the aperture of the veftibulunij bave only a fmall cartilaginous piece covered by the flefti. In ferpents and the camelion the handle of fmall bones traverfes the f\e(h, and re(is againfl: the bone which fupports the lower jaw. In all thefe animals the membrana tympani is wanting. Birds and fome reptiles have only one fmall bone I in the frog and the toad there are two, and . in all the mammalia there are four. Thefe four fmall bones are : the malleus, incus, Jlapes, and os orVtculare, The plate of the Jiapes covers the aperture of the veflibulum ; and the handle of the malleus adheres to the membrana tympani. Thefe fmall bones are moved by one or more fmall mufcles, which are often fcarcely percepti- ble. Birds have one, and the mammalia three i in the latter, two are attached to the malleus, and one to the ftapes. Fitbes and reptiles have no exterior auditory conduit. Birds have one, which is very fhort : the exterior orifice is furrounded and covered by a (tvj very Q line SYSTEM OF THE SENSES. 257 fine feathers^ difpofed in a manner altogether pe- culiar. In the mammalia, the auditory conduit is of greater extent, and afFecis various diredlions : it is in part ofieous, and is generally terminated on the outlide by a cartilaginous espanfion, concha^ of dif- ferent forms and lizes, and more or lefs moveable by means of particular mufcles. Cetaceous animals have for auditory conduit a very fmall, twifted, cartilaginous canal, which is lodged in the thicknefs of the fat. In feveral of the mammalia the concha is wanting. 2,83. In man, the organ of hearing confifls alfo of a membranous labyrinth contained in an ofleous, one, a barrel or tympanum, an auditory conduit, and a concha. The membranous labyrinth is compofed of a capfule containing a vifcous fluid, in which is found the labyrinthic nerve : this foft and pulpy- nerve expands into a reticulatioif in the different excavationsof that part. The membranous labyrinth has the fame form as the offeous one in which it is contained. It confifts of a cochlea, a veftibulum, and three femi- circular canals. All this apparatus, contained vn the petrous portion of the temporal bone, feems, in the foetus, to be feparated from the reft of the bone; but in the courfe of time it becomes con- '^01.-. X, s founded 258 ORGANIC STRUCTURE. founded with it, and appears to be hollowed out in the fabftanceof it. The cochlea is placed over and correfponds with the carotid canal ; its cavity prefents a double fpiralj which makes two turns and a half. The bafe of the centre of the fpiral correfponds nearly with the acouftic foramen ; it is pierced with fe- veral holes to afford a paffage to the filaments of the labyrinthic nerve, and its fummit drre6^ed out- wards, and upwards terminates in a fort of funnel. The partition which feparates the two fcalse of the cochlea rs half ofleous and half membranous s the ofleous part is towards the axis. Of the two fcal^ of the cochlea, the interior,, ■which is larger and fhorter, is terminated at the bafe by a round aperture that proceeds to the barrel or tympanum : foramen of the cochlea. The exterior one, which is narrower and Ion- ger^ ends at the veftibulum : thefe two fcalse communicate with each other at the fummit of the cochlea. The three femi-circular canals, fituated b-ehind the cochlea, are feparated from that part by the cavity of the veftibulum. They form three arcs^ the two extremities of which are widened and pro- ceed into the veflibulum. The three lemi-circu- lar canals give only five apertures in the vefiibu- lar cavity, becaufe two of them^ are confounded together on entering it, 6 The SYSTEM OF THE SENSES. 250 The veftibulum forms an intermediate cavity between the cochlea and the three femi-circular canals. It is of a round form, and prefents in the infide : the five apertures of the femi-circular ca- nals ; the orifice of the exterior fcala of the coch- lea; an oval hole which enters the tympanum; and a fmall triangular aperture which penetrates into the cranium. This aperture is terminated by a flit towards the middle of the fuperior edge of . the apophyfis petrofa, where it condu(5ls to a re- ceptacle formed by the two laminae of the meninx, and which contains a ferous matter : aquedu5l of the vejlibulum or of cotunnl. Near the aperture of the interior fcala of the cochlea is another very fmall one^ which proceeds to the interior of the cranium, opening in the middle of the pofterior edge of the apophyfis pe- trofa : aquedii6l of the cochlea. The offeous labyrinth communicates with the tympanic cavity by the aperture of the cochlea and that of the veflibulum. The tympanic cavity correfponds with the ex- terior fide of the labyrinth ; it is round, contains a feries of four fmall bones, and exhibits feveral apertures. Of the apertures which enter the tympanic ca- vity, one of them, of a round form, communicates with it at the bottom of the interior fcala of the cochlea; the other, which is oval, opens into the s 2 veflibulum: 260 ORGANIC STRUCTURE, veftibulum ; it is covered by the bafe of the ftapes, Thefe two apertures are feparated by a round emi- tience called the promonlury. The tympanic cavity exhibits alfo before, an aperture divided at its entrance by an ofleous plate fhaped like the mouth of a fpoon : this aperture affords a paffage, above the plate, to the interior mufcle of the malleus, and below to the guttural conduit of the ear. This conduit, which Is very narrow, ifTues from- the temporal above the inflex canal of that bone; It becomes cartilaginous, and forms a fort oiEuJia^ clnan tube, which proceeds to the bottom of the mouth behind the nafal fofTae. This conduit, which ellablifhes a communication between the back part of the mouth and the tympanic cavity^ is lined with a mucous membrane. Behind, the tympanic cavity prefents a triangu- lar aperture, which proceeds into the malloidiaii cells ; thefe cells are not very apparent in infancy, but expand with age. At the top and behind, it exhibits a fort of pyramid, having at its bafe an aperture which communicates with the fpiroid canal of the temporal bone, and affords a paffage to the tympanic nerve. At the bottom it has a imall flit through which pafs one of the mufcles of the malleus and the tympanic nerve, glenoidal fjpire. At the lower part it has feveral fmali holes for the pafTage of the fanguine vefTels.. The SYSTEM OF THE SENSES. 2'6l The four fmall bones, contained in the cavity of the tympanum, are the ftapes, the os orbiculare, the incus and the malleus. Th&Jlapes has exadlly the form indicated by its name ; the bafe of it is applied to the oval aperture of the veltibulum, where it is retained by an ex- panfion of the periofteum. On the fummit it has a fmall cavity which receives the os orbiculare. This bone^ which is exceedingly fmall, is found between the ftapes and the long branch of the incus. The incus confifis of a body and two branches ; the body has an elliptic cavity, v^hich receives the head of the malleus. The malleus has the form of a club ; its head, turned upwards and backwards, is articulated with the body of the incus ; its neck has a long flender apophyfis, and its handle another fhorter one, which is turned outwards and upwards. The extremity of the handle of the malleus is fixed to- wards the middle part of the membrane of the tympanum. Thefe bones are moved by three mufcles, two of which belong to the malleus, and the third to the flapes. Of the two mufcles of the malleus, the inferior one is attached to the apophyfis placed below the neck of that bone, and proceeds in the canal of the guttural conduit of the ear : it feems as if intend- S3 ed 262 ■ ORGANIC STRUCTURE. ed to pull the malleus inwards and forwards, and to ftretch the tympanic membrane. The anterior, which is lefs apparent, is attached to the long apo- phyfis of the neck of the malleus; iflues through the glcnoidal fifTure, and is attached to the exterior fide of the guttural conduit : it feems intended to O pull the malleus outwards and forwards, and to relax the tympanic membrane. The mujcle of the Jiapes is the fmalleft % it is attached, on one fide, to the fummit of the pyramid, and on the other to the neck of the ftapes. Its adion feems to be t6 pull the ftapes outwards, and to firetch the membrane which retains the bafe of that bone around the aperture of the veftibulum. The four fmall bones, by their union, form a chain between the aperture of the veftibulum and the tympanic membrane. This membrane forms the exterior fide of the tympanic cavity. It correfponds, without, to the bottom of the auditory conduit, and feparates the interior from the exterior part of the ear. In the foetus, it adheres around an ofleous ring incom- plete at the top. This ring, in adults, is con- founded with the body of the bone, and is con- verted into an ofleous conduit. The merabrana tympani has an oblique direc- tion ; it proje61:s outwards, and in the middle ad- heres to the extremity of the malleus ; it is thin and SYSTEM OP THE SENSES. 263 and tranfparent, and Teems fiifceptible of being feparated into four thin leaves. On the exterior fide of the membrana tympani there is found, in adults, an ofleous conduit of eight or ten lines, flightly bent downwards. This conduit, which is broad at the bottom, has after- wards a fort of contraclion, and becomes wider on the outiide. The edges of the exterior orifice of this oileous conduit are unequal, and have attached to them a cartilaginous cornet, which becomes uider^ and forms a cavity : concha. The exterior edges of the concha are produced by a cartilaginous thick roll, called the aritheUx, This roll is fingle towards the lower part of the ear, and at the upper is divided into two grooves, which leave between them a fmall fofla, called the navicular. From the upper part of the concha arifes another cartilaginous roll, which proceeding upwards, and then backwards, marks out the contour of the ear: helix. It is terminated, at the bottom, by a foft flexible appendix. The interval between the two rolls is marked by a groove. Before theconchais a triangular eminence, which riles above the middle of its cavity, tragus ; oppo- fite to this eminence is another, towards the lower part of the anthelix : it is feparated from the for- mer by a deep groove, antitragus. The whole dif- s 4 pofitiou 264 ORGANIC STRUCTURE, polition of the exterior ear is fuch, that the fono- rous radii, which fall on thefe different parts, are reflected towards the auditory conduit. The exierior ear, which is formed of a cartila- ginous fubftance covered by the fkin, is retained by three ligamentous expanfions, and exhibits the elements of feveral mufc)es. Of three ligaments which proceed towards the cartilage of the ear, the upper one arifes from the exterior aponeurofis of the temporo-maxillian mufclci {heanterior^ from thebafeof the zygomatic apophyiis; and the pq/ierior, from the bafe of the maftoidian apophyiis. Of the mufcles of the ear, three are fituated around the exterior part of it, and could move it if they were more prominent : one of them is at the top, the other below, and the third behind. Exterior mufcles. The reff, fiill lefs apparent, are fituated on the interior part, and do not extend beyond it. In^ ter'ior mufcles. 2,84. The exterior mufcles. The fuperior ex- tends from the aponeuroiis of the occipito-fron- tian, to the navicular fofia, temporo-auricidmn : it would tend to raife the cartilage of the ear. The anterior extends from the aponeurofis of the occipito-frontian, above the zygomatic apophyfis, as far as the anterior part of the helix : the zygO" 7natQ aurkuliiin: it would draw the ear forwards. The SYSTEM OF THE SENSES. 265 The. pojierior, which is the moft apparent, varies both in its form and pofition, and is often divided : it extends, in general, from the maftoid apophytis to the pofterior part of the concha, the majioido- ^uricuTian ; it would tend to pull the ear back- wards. In fome perfons this mufcic has been feen to perform very ftriking movements. The mufcles called interior confift of fome pale flefhy fibres, fcarcely fen(ible, diffufed over the helix, the tragus, the anti-tragus, and the concha ; the great and ihQf?nall heJifian, the tra- gian, the antl-tragian, and the tranfverfian. The Ikin of the interior part of the concha, and that of- the exterior auditory conduit, contain a great number of glandulous follicles, which fecrete a thick humour of a yellowifli colour, and limilar to wax, called the cerumen. The entrance of the auditory conduit is co- vered alfo with very fine hairs, proper for inter- cepting fmall bodies which might be introduced into that cavity. OF [ 266 ] OF SMELLING. 285. The organ of frnelling confifts chiefly of a membrane called the olfa&ory^ which lines the folds of the nafal fofFae, and which has the pro- perty of perceiving the contadl of certain mole- culse of matter. The nafal foffe ferve always for affording a palTage to the air, in the refpiration of animals with lungs ; in fifhes thefe foflse are hollowed oat only in the thicknefs of the fnout. The organ of Imell feems to be ftronger, ac- cording as the olfadory membrane is of greater extent. The nafal fofTae of the mammalia are formed by the OS ethmoides, its anfra6tuofities and its tur- binated portions, the vomer, the bones of the nofe, the fuper-maxillary bone, the inter- maxillary, the fpbenoid, and the palatine. The nafrd fotiie arc of greater or lefs extent, being fometimes broad and fometimes narrow : the entrance of them is generally turned forwards; in cetaceous animals, and birds, they are turned upwards. Some fifhes, fuch as the toad-fifh, have their noftrils fupported by pedunculi, The SYSTEM OF THE SENSES. 267 Thefize of the noflrils is much increafed by the numerous tvvifted folds, communicating with each other, of the ethmoid ; and of its turbinated por- tions, which have a fpiral form. Thefe turbinated bones, in mod of the mammalia, are very large, and turned round in a very lingular manner: in birds they are, for the mod part, cartilaginous, and large, efpeciall}^ in birds of prey. Reptiles have fome membranous laminae, which enlarge the furfaces of their nafal fofTse. In fifhes, thefe furfaces are enlarged by mem- branous prolongations, which are indefinitely fub- divided in wonderful order. In the mammalia, the nafal folTse have a com- munication with cavities formed in the thicknefs of the frontal bone, the maxillary and the fphe- iveid ; thefe cavities, which communicate with the nafal fofTee by narrow apertures, are diftin- guifhed by the names of: the frontal, maxillary^ and fphmoidal finuft's, according to the bones in which they are formed. The frontal -finufes are wanting in fome of the rodentia, fome edentia, and feveral other animals; they are very fmall or are wanting in apes ; in man, and in many of the mammalia, they aipe very ilriking ; and in fome carnivorous animals, fuch as the dog, wolf, and fox, and particularly in fome ruminating anim,als, as the ox, the goat and the fhcep, they are of great extent : they arc very large 0,58 ORGANIC STRUCTURE. large alfo in the fvvine, and exceedingly large in the elephant. The maxillary finnfes are generally very fmall in carnivorous animals, in moft of the rodentia and edentia. In ruminating animals^ and in feveral more of the mammalia, they are very large. Th^Jphenoidal fnmfes are wanting in moft ani- mals, and particularly in thofe which have the body of the fphenoid very much flattened. They are fmall in the fwine and the hippopotamus, and very large in the elephant. Cetaceous animals have no finufes. In birds the cavities of the bones of the cranium have a communication with the ears, and not with the noftrils. The nafal cavities and their numerous folds are covered by a periofleum, like all the other bones. But this periofleum is lined with a fungous, thick and very red membrane, on which is expanded a beautiful vafcular reticulation ; it is furniflied with glandulous follicles, which fecrete a peculiar kind of mucous matter, and adheres very ftrongly to the periofleum. The membrane v,'hich covers the infide of the finufes, and ferves them as a periofleum, is very thin : it is fmooth, and feems to be of a texture different from that which lines the noflrils. The fummit of the ethmoid bone, which cor- refponds to the bafe of the cranium, in all the mammalia. SYSTEM OF THE SENSES. 260 mammalia^ cetaceous animals excepted, is pierced with a great number of fmall holes : in other ani- mals it has only one hole or canal. Thefe boles afford a paffiige to branches of the ethmoidal nerve, which expand over the whole furface of the iiafal membrane. The number and iize of thefe holes feem to be proportioned to the power of fraelling : in car- nivorous animals they are very numerous. To the fore- part of the bones, which form the anterior edges of the nafal cavities, and the offeous plate which fcparates thefe cavities, are attached cartilages, which are prolonged in order to form the edges of the noftrils. Thefe cartilaginous edges are of different fizes, and aflume various configurations in different animals; in the greater part of the marnmalia they are moved by fome par- ticular mufclcs, fixed on the cheeks, and which tend to dilate or to contrail the entrance of the nafal cavities, and to move, in different dire(?fions, the projecting muzzle of fome fpecies, fuch as the fvvine's fnout. ^ In the elephant, the noftrils affe6l a configuration altogether peculiar; they are continued in the form of a very elongated cone, which conftitutes the trunk. This canal, which is broader at the root, and divided by a partition, has two fpiral turns to- wards its bate, and communicates with the ofleous noftrils by means of a valve, which can be raifed. The '270 ORGANIC STRUCTURE, The trunk of the elephant is lined with an apo- neurotic membrane, pierced with holes through which oozes a vifcous liquid. The fides are formed of two layers of fibres^ one of which proceeds, in a radiated form, from the interior aponeorofis to- wards another aponearolis beneath the Ikin : thefe fibres, by contracting, render thinner and extend the fides of the trunk, without diminifhing its ca- vity ; the other layer conlifts of fhort longitudinal fibres, placed quite around it, and proper for (hort- ening the trunk partially. Moft reptiles have fome fmall mufcles for di- lating or contracting the entrance of the noftrils. In birds, the aperture of the noftrils is furrounded only by a roll of the fkin. In fithes, they are nar- rower at the bottom, and fufceptible of fome movement. In cetaceous animals, the noftrils exhibit no pro- je(9:ing folds in the infide, and have no communi- cation with the finafes; they are Imed with a thin membrane of a clofe and fmooth texture, the ftru6iure of which has no refemblance to that with ■which thefe cavities are covered in other animals t the ethmoid has no aperture, and there is no olfadlory nerve ; fo that it is highly probable that the noftrils in thefe animals cannot ferve as the organ of fmelling. Their peculiar ftru6lure renders them proper for another purpofe^ they are divided, .at their ofileons entrance^ by a valve which can SYSTEM OF THE SENSES. ^/l be raifed ; the inferior parr, or that which cor- relponds to the pharynx, is provided with a very llrong mufcular apparatus, and in the fuperior or external part is a cavity, the contrad:ible fides of which terminate outwardly in the narrow aperture of the nofi:rils. Thefe animals, by means of this arrangement, can throw up' the water, which they have in their mouth, to a very great height, through the noitrils. In the guttural conduit of the ear of cetaceous animals is obferved a pretty wide hole, terminating in a large cavity, lined with a foft, mucous, and blackifb membrane, which has a communication with the frontal llnus. The nerves of this cavity, as well as thofe of the noftrils, proceed from a pair, (the fifth) which, in all the mammalia, fend out others to the noilrils ; fo that it is very probable that this peculiar apparatus ferves as the organ of fmelling to thefe animals. 286. In man, the olfaclory membrane is of a moderate lize, and of a rofe colour; is thick, of a villous texture, and contains a great number of mucous follicles, which fecrete the mucus o( the noftrils. This membrane receives a g-reat number of fanguine vefTels, which arife chiefly from the fpheno-palatine and ethmoidal arteries. The ethmoidal nerve, when it arrives near the ethmoid bone, becomes thicker; alTumes a pulpy conlifteQce, 271 ORGANIC STRUCTURE, confiflencCj and penetrates to the noftrils by a great number of fmall holes or ofleous tubes, ex- hibited by that bone. This nerve feems to be exclu- iively appropriated to the fenfe of fmelling'. The fifth pair furnifli, in the nofe, feveral nerves which feem to be deflincd only for the life of that organ. The tiafal foflk have a communication with the frontal, maxillary, fphenoidal,and ethmoidal iinus- es ; and the membrane which lines thefe iinufes is of a ftru6inre different from that which lines the infide of the nofirils : it has a finer texture, is fmoother and tranfparent : it adheres weakly to the bones, and ferves th'em as a periofteum. • The parts on which the olfactory membrane ex- pands conftitute the nafal fofTae. Before and at the top, they are formed by the bones of the nofe ; on. the fides, by the fuper-maxillaryj lacrymal, and ethmoid bones, and by theinferior turbinated bones; behind by the fphenoid ; at the bottom by the fuper- maxillary bones, and thofe of the palate. They are divided alfo longitudinally in the middle by the vomer, and the perpendicular plate of the eth- moid, and before by the cartilage by which thefe bones are continued : this plate is very often warped. The nafal {bf[k are not very apparent in infancy^ and do not attain to their complete fize till the period of full growth ; at which time they are nearly three inches in extent from before back- wards. SYSTEM OF THE SENSES. 273 wardsj and two inches from the top to the bot- tom. The lateral fides of the nafal foffe approach each other at the top, and diverge at the bottom. The narrovv fummit of the noftrils exhibits an arch of three planes. The anterior, which is in- clined forwards, conftitutes the arch of the nofe ; the middle one, which is horizontal, exhibits the orifice of the ethmoidal holes ; and the poflerior correfponds to the aperture of the fphenoidal linufes. On the lateral fides of the nafal foffe there are thrte turbinated bones, two of which belong to the ethmoid bone : the inferior one, for the moil part, is free. The fuperior turVinated hone adheres to the arch of the nafal foffie, and is feparated from the middle one by a large groove^ ihc fuperior meatus. This groove exhibits before, the aperture of the pofterior ethmoidal cells j and behind, the fpheno- palatin foramen. The middle turVinated hone, larger than the pre- ceding, fwells out before, and terminates in a point behind ; it is feparated from the lower one by a broad groove, called the middle meatus. Be- fore, this groove exhibits the common aperture of the ethmoidal cells, and of the frontal finufes; and in its middle part an aperture, which pro- ceeds into the maxillary finufes. VOL. I, T The 274 ORGANIC STRUCTURE. The inferior iurhinated lone, lefs twlded and wider than the other two, has below it a large furrow, called the inferior meatus, which exhibits before the aperture of the nafal canal. To the bones, which conftitute the anterior aperture of the noftrils, are affixed two cartilages, which form the arch and alas of thenofe ; and an- other which produces, before, the middle portion. Thefe cartilages are covered on the outfide by the Ikin, and on the infide by the olfa61ory membrane : in that part, this membrane is thinner and lefs villous than in the reft of its extent, and it exhi- bits there fome very fine hairs. The cartilages of the nofe are moved by the occi- pito-frontian, fuper-maxilIo-nafian,the great fuper- maxillo-labian, and the alveolo-nafian mufcles, the ufes of which are common to the other parts. The pofterior aperture of the nafal foflae corre- fponds to the back part of the mouth ; and, at the moment of deglutition, is clofed by the velum palati. The finufes expand with age, and give more extent to the face. The two frontal finules arc divided, in the middle, by a longitudinal partition for the moft part warped. Their extent varies a great deal j fometimes they occupy the whole frontal region, and fom.etimes they are fcarcely apparent. The two fphenoidal finufes are hollowed out in the SYSTEM OP THE SENSES. 275 the body of the fphenoid, the whole extent of which they foinetimes occupy, and they open be- hind the fuperior turbinated bone. The two maxillary finufes are the largefl: ; they occupy the whole thicknefs of the fuper-maxiJIary bone; they afte6l a triangular form, and open into the middle meatus. ■ The four ethmoidal cells correfpond to the top and fides of the nafal foflas, and have a com- munication with each other. The two exterior open into the middle meatus, with 'the frontal finus; the two pofterior proceed into the fuperior meatus. T 2, • SYSTEM [ 276 ] SYSTEM OF DIGESTION. 287. iHEfyJiem of dig eft ion in animals. The fy- ftem of digeftion confifts chiefly of a membranous bag, the fides of which fecrete a particular juice. The aliment, introduced into this bag, is there transformed into a kind of pulp, which contains the nutritive juice. This juice is diflributed to the different parts of the body, where it becomes mixed with the blood, and the reliduum is difcharged. Some animals have merely an alimentary bag with one aperture. In the greater number, the apparatus of digeftion confiils of one canal with an aperture for entrance, and another for efcape. This apparatus, however, is rarely fo fimple : it is rendered complex by feveral dilatations, and confifls of different pieces. In fome animals the alimentary canal exhibits fmall dilatations ; but in others thefe dilatations are of confiderable lize, and conflitute ftomachs. When animals have a difiin6l ftomach, the name of (sfophagus is given to that part of the alimen- tary canal which proceeds into it ; and that of inteftine to the one which proceeds from it. The juice of the alimentary canal appears to be the only one which in feme animals ferves fordigef- tion % SYSTEM OP DIGESTION. 277 tion ; but in the greater nnmber this canal alfo re- ceives the liquors fecreted by the liv^er and by the pancreas. The entrance into the oefophagus is fometimes a fimple aperture ; but, for the moll part, this en- trance is a mouth, formed of jaws, and furnithed with teeth, a tongue, falivary glands, &c. 2,88. Jaws are found in crufiaceous animals, in- fers, and in feveral of the mollufca; but in thefe animals they exhibit a peculiar ftru6^ure, which has very little relation to that of the fame parts in the mammalia. In fome zoophytes, fuch as the fea hedge-hog and fea-nettle, there are tbund alfo an organ of maftication, and teeth moveable by the means of mufcles. All vertebral animals are provided with two jaws * : in birds, fiOies, and ferpcnts, they are both moveable. In fome reptiles, as the crocodile and the lizard, and in all the mammalia, the lower one alone is moveable. In the mammalia, the lower jaw exhibits a con- dyle, which is articulated in a cavity of the tem- poral bone. In fifhes, reptiles, and birds, the jaw and man- dible are each furnifhed with a cavity ; and thefe two cavities receive an intermediate bone (fqiiarc hone), which ferves as the means of union, and fupplies the place of a condyle. * Crabs have four, five, and fometlrnes ten pairs of jaws > T 3 In 27S ORGANIC STRUCTURE. In all vertebral animals, the jaws move vertically and horizontally ; in infeds they move tranfver- fally. In ferpentSj the jaws, which are moveable from above downwards, can be feparated alfo in a lateral dire6lion, which produces an aperture capable of receiving an obje6l of prey.larger than their whole body. Some fifhes have jaws which, without any move- ment of the head, -can be carried fuddenly for- wards to catch their prey ; as is the cafe in the ray, the (hark, and fome reptiles. In the mammalia, the condyles are of greater of lefs breadth and flattened ; in ruminating animals, their tranfverfe diameter is greateft ; in the roden- tia it is greateft from before backwards ; and in mat), the ape, &c. it is greateft in an oblique di- reftion. In animals which do not mafticate, the condyles are not very apparent ; in the large carni- vorous animals they are very ftrong, and are lodged in deep cavities, in which they are inclofed and retained by a roll and an ofleous capfule. The mufcles which move the jaws vary with refpedl to their number in the different clafies of animals. In the mammalia, thefe parts are elevated by four mufcles^, and deprefted by a greater number. The former are : the crotaphite, the mafleter, and the two pterygoidian ; the latter are attached to the OS hyoides, the bafe of the cranium, and to the fternum. SYSTEM OP DIGESTION. 2/9 flernum. Of tiiefe, the digaftric of man and of apes is monogaftric in carnivorous animals. In moll of the carnivorous animals, fuch as the hyena and the tiger, the crotaphite mufcles extend as far as the fummit of the cranium, vi^here they are fixed to a ilrong ofleoLis ridge. The mafleter, which is very large, is attached to. a prominent and extenfive zygomatic arch. Thefe two mufcles, in thefe ani- mals, move the jaws from above downwards; and in graminivorous make them perform lateral move- ments. Quadrupeds have lips which ferve to clofe the mouth, and to retain the aliments j in man, they contribute alfo to fpeech and to the adion of the phyfiognomy. 289. Teeth. Fifhes, reptiles, and the mam- malia, have jaws provided with teeth, implanted in the bodies of the maxillary bones, or retained only in the gums. Fifhes have teeth not only in the jaws, but alfo in the palate, the throat, on the bones which bear the branchiae and the tongue, and even on the tongue itfelf. If this difpofilion is favourable to maftication, it cannot be fo for tafting. In thefe animals the teeth, which afFe6l different forms, are fupported by an offeous tubercle covered with a membrane. When this membrane is deftroyed, the tooth and tubercle drop out, and the place of both is afterwards fupplied by others, T 4 In '280 ORGANIC STRUCTURE* In the mammalia, the teeth are of three kinds : the fharp ones in front are called incifors; na* turalifls always give the name of incifors to thofe implanted in the inter-maxillary bone ; others cut into feveral faces correfpond to the angles of the lips, and are the angular teeth ; and the large flat ones, placed backwards and on the fides of the mouth, are the grinders on molar teeth. In fome animals, the teeth are fo large, and of {o lingular a conformation, that they cannot be diltinguillied by that name: fuch are the lufks of the narval and the hippopotamus ; thofe which proceed from the mouth of the wild boar, and thofe which fall back even on the head of the barbyroufla. The grinders exift in all animals furnifhed with teeth. The prefence of the reft is not conftant: grinders only are found in the edentia ; incifor teeth are wanting in thefloth,and angular teeth in the rodentia. Ruminating animals have incifive teeth only in the lower jaw. The three kinds are obferved in the folipeda, feveral of the pachyder- mata, the carnivorous, the quadrumana, and in man. In the lad place> the ant-eaters are defti- tute of them. Several animals, fuch as the folipeda, have a part of their jaws without teeth, interdentium. It is in this part that the bit of the horfe is placed. In the hare, the fox, and often in children, the incifors SYSTEM OP DIGESTION. 2S f incifors are ferrated ; in bats, thefe itidentations are fometimes fo deep, that they refembie the teeth of a comb ; in the rodentia they arc long and curved ; the didelphis marotarfas has the angular teeth cfhorter than the reft ; in moft apes the angular teeth begin to be lengthened, and in the large carnivorous animals they are ^o long that they crofs each other, and are received in the indenta- tions which they mutually prefent. The large furface of the grinders has various configurations ; in carnivorous' animals it is co- vered with points in one or more rows ; in the pachydermata it is tuberculous, and in the large graminivorous exhibits falient lines : thefe lines in the rumirtantia form feveral crefcerits, and in the folipeda a double feftoon. Carnivorous animals, the quadrumana and man bave the teeth formed of an ofleous fubftance, co- vered on the outfide by a coating of enamel. In the folipeda and ruminantia the teeth confift of an ofleous fubftance, enveloped in the membranous follicle which originally contained them. The bodies of the teeth are covered with an enamel of a golden yellow colour : the membranous follicle oflifies, and forms around the teeth a very hard cortical covering. In the elephant and the hippopotamus, when young, they are compojed of vertical laminae, placed on each other without adhefion. Thefe ofTeous 28'2 ORGANIC STRUCTURE. olTeous laminas, cut unequally at the rummit, foon unite by means of an en am el ly juice, which infi- nuatcs itfelf between them, and they then form a folid tooth compofed of plates of enamel and oileous plates. «» By the efFeft of maflication, the edge of the ofleous planes fooin becomes worn, and in the courfe of a tew years the furface of the teeth is covered with falient and unequal lines of enamel. This difpofition, analogous to that of millllones, is exceedingly proper for maftication- In the orySierope the teeth have the porofity of reeds : they are foft, and can be eafily cut. The name of root is given to that part of the teeth which is inferted in the jaw; the part with- out is called the crown; and the ftridure which often feparates thefe two parts is called the neck. The fingle teeth are hollow during the tirfl: years of their growth ; the cavity lined with a thin membrane, over which the yeflels and nerves are fpread, is filled with a gelatinous fubftance, and becomes partly clofed up with age. The part firft produced is the body of the tooth; it foon fliows itfelf without, and begins to be worn before the root has attained to its full growth. When it has ceafed to grow, the capacity of the alveolus decreafes, and is gradually filled upj the root is deflroyed in the fame proportion, and the tooth thus worn at both ends becomes very (hort 6 at SYSTEM OF DIGESTIONT. 2S3 at an advanced age: it then turns loofe in the focket, and at laft drops out. In the ruminantia and folipeda the body of the tooth is fometinnes completely worn out, and the roots are feparated. In thefe animals the decay of the teeth is much more rapid than in the car- nivorous ; in man, this decay takes place in a very ftriking manner. The wearing out of the teeth, which goes on in a conftant and regular manner, ferves to indicate with precifion the age of the fo~ lipeda and ruminantia. In a great number of animals, a part of the teeth drop out, and their place is fupplied by others. Some fidies, fuch as the fhark, have their teeth inferted in the flefh, and fufceptible of being re- newed four or five times : thefe new teeth are con- cealed behind thofe the places of which they are intended to fupply. The fame difpofition is ob- ferved in fome reptiles, as may be feen by the venomous dart of the viper. In the mammalia, a part of the teeth drop out at a determinate period j the germs of thofe which fucceed are placed behind the firft. In the cro- codile, the teeth are conical and hollov^, and re- ceive in their cavities thofe which are to fucceed them. The roots of the teeth which drop out are fpeedily deftroyed ; they become loofe in the fockcts. 284 ORGANIC STRUCTURE, fockets, and are forced out by thofe which aflame their place. This fecond dentition, ferves to indi- cate the age of the animals. 2CO. The to7io-ue is a mufcle which affedls dif- ferent forms : in fome of the mammalia it is broad and fiiort ; in feveral birds it is long and narrow % in moft ferpents and feveral lizards it is forked. In mofi: of the mammalia the tongue is covered by a mucous membrane, furnifhed with nervous papillae proper for perceiving the impreffion of favours. In feveral reptiles, it is covered by a fcaly fubftance, and in ibme fifhes it is furnifhed with teeth. The tongue ferves for maftication, for deglutition, for the voice, and for fpeech. It is fixed to the os hyoides by mufcles, and the point of it is gene- rally directed forwards : in the batrachians, how- ever, it is turned backwards. To the os hyoides, which is free and fufpended in the back part of the throat, the mufcles of the pharynx and of the jaw are affixed. It appears, for the moft part, under the form of a portion of a cartilaginous Cylinder ; but in feveral animals this form varies. The tongue, in general, poflefies great mobi- lity ; in molt animals it can be conveyed beyond the mouth, and by reptiles in a very remarkable manner. In the green wood-pecker, the tongue, which is contained in a membranous fheath, and armed SYSTEM OF DIGESTION. 285 at the point with hooks, can be extended beyond the bill of the animal to a greater length thsn that of its body : it is lengthened and drawn back by a very curious organic difpofitlon. 291. In fifhes the mouth communicates with the oefophagus, and with the aperture of thebran- chi^i in animals with lungs, it proceeds to the cefophagus, the pharynx, and the noftrils. In thefe animals, which perform real madica- j tion, the infide of the mouth is moiftened not only by the fecretion of the mucous membrane, which lines that cavity, but alfo by the faliva fecreted from feveral glands. In carnivorous animals, the faliva under certain circumftances afiumes a very venomous quality. This property is found in the produdlion of a gland of the viper, and in an analogous liquor of fome infedls. The oefophagus is a mufculo-membranous^ ca- nal, lined with a miiccas membrane: its length varies as that of the neck ; it proceeds to the tio- mach. 292. The Jiomach is a dilatation of the ali- mentary canal. In very ravenous fifhes, and moft ferpents, this dilatation fcarcely exifts. In the carnivorous mammalia the ftomach is tingle, and not very capacious ; in the graminivorous it is' larger ; in birds it is double, and often triple; in the 2S6 ORGANIC STIIUCTURE. the ruminantia it is quadruple ; in cetaceous ani- mals there are five in fucceffion. The flomachs of infecls have a great analogy with thofeof red-blooded animals. In butterflies the ftomach is merely a dilatation fcarcely lenlible, or does not exift at all ; in other inre6ls the flo- raach is fingle, double, or multiple. In thofe which feed on the juice of flowers, fuch as bees, ihe fingle ftomach is large ; in infects which fuck blood, or feed on animal matters, it is of lefs ex- tent. Double fl;omachs belong, in particular, to carnivorous infecfls, fuch as the coleoptera of that genus : in thefe infecls, the firfl: flomach is inuf- cular, like the gizzard of birds, and the fecond membranous : in fome there is alfo a fort of crop. Multiple ftomachs are found in infeds which ex- ercife a kind of rumination, as is the cafe with locufls. The numerous clafs of infedis, with refpedl to this part, exhibit an organization exceedingly cu- rious and complex j but we muft confine ourfelves chiefly to red-blooded animals. The flomach of carnivorous, and particularly ot herbivorous animals, which do not ruminate, ex- hibits alfo one or more dilatations ; but for the mofl part there are two. In herbivorous animals thefe dilatations are very large ; in the pojyphaga they are lefs, and in the carnivorous fmall. In the SYSTEM OF DIGESTION. 287 the laH^, the ftomach appears to be only an intefti- nal dilatation, which has little obliquity ; it is in general wrinkled, and can be confiderably dii- tended to receive an ample repaft. The fides of the ftomach are for the moft part of a mufculo-membranous nature ; and the inte- rior part of them is always covered with a mucous membrane. In birds, which have two and for the moft part three ftomachs, the firft, called the crop^ is a membranous bag, in which feeds remain and be- come foft ; the fecond, ventnculus fuccenturiatus, is thicker, but of lefs fize ; it is furnifhed with a great many mucous glands which fecrete abun- dance of digeftive juice. It was the juice of this ftomach which Spallanzani employed in his inge- nious experiments on the gaftric juice. The third, ov gizzard y is formed of two very thick mufcles ; the cavity of it is not larger than the ventricle, but the thicknefs of it makes it appear much more voluminous on the outfide. The two mufcles of the gizzard exhibit, on two faces, a very ftrong round tendon, from which the mufcular fibres ex- tend in a diverging form. The cavily of this ftomach is lined with a very thick mucous mem- brane, which may be eafily feparated, and which is fafceptible of regeneration. The gizzard pof»- feftes a very ftrong contrad^ile force : its adlion feems to fupply the place of maftication. Granivorous 2BS ORGANIC STRtCTUKE. Granivorous birds, in general, fvvallow fmal! pebbles, and the Itrong and varied con traction of the gizzard on the feeds, mixed with thefe hard bodies, effedts in this manner their trituration bv repeated fri61:ion. In the flomach of the oilrich and the caflbwary federal pounds of pebbles are often found intermixed with bits of giafs, iron, &c. In carnivorous birds the gizzard is much thinner : in thofe vi'hich feed on -fifla or worms there is Icarcely any. Thefe birds have no crop ; and the ventricle which is their principal ftomach is gene- rally very large. The gaftrlc apparatus of the mollufca cephalo- poda and gafteropoda has fome analogy with that of birds. They have a gizzard preceded by a crop. The ftomach of the volutas as well as of crabs is furnifhed with two olleous pieces proper for trituration. In the bivalva, fuch as the mytilus, the alimen- tary canal traverfes the liver, and is dilated in its thicknefs to form a ftomach. Some of thefe ani- mals, fuch as the oyfter, have a fecond ftomach. Ruminating animals have four ftomachs, the aggregate of which exhibits a very remarkable fyflem of digeftion. The firftj veniriculus, called paunch by the vul- gar, is exceedingly large, and occupies a great part of the abdominal capacity ; it exhibits infe- riorly a longiludinal re-entering fold, which di- vides SYSTEM OF DIGESTION. ^Sg vides It into two principal cavities; its interior furface is furnifhed with large projedling conical glands. The fecond flomach, the honeycomb. Ion-- net, or hng^s hood, is round and much fmaller ; it is feparated from the former by a membranous partition in the form of a horfe-flioe, above which thefe two Itomachs have a communication with each other. The inlide of the lonnet exhibits deep cells of different forms, analogous to the cells of a bee-hive. The cefophagus communicates with thefe two ftomachs towards their upper part, where it forms an inverted gutter, the thick edges of which can approach each other to form adu6t. This gutter, continued with the cefophagus, conlifts of two folds, which advance in a parallel diredion as far as the third ftomach, where they interfedl each other at an acute angle, and prevent the entrance of coarfe aliment. In this part the gutter exhibits furrows, which are continued with the membranous laminae that form the third fto- mach : omafum or many plies. The omafum, of an ovoid form, and fome- wbat larger than the honHet, is divided, in the di- redlion of its two orifices, by a great many mem- brajious partitions, which produce furrows deep and narrow, particularly towards their middle. The fuperior furrows are deeper and much more crooked than the inferior ; the inlide of thefe fur- voL, I, u rows '2gQ ORGANIC STRUCTURE. rows is farnifhecl with rugous glands covered with papillas. The orifices of this ftomach are mucb lower than its body : the anterior, on the left^ corrcfponds to the horinet, and the poftcrior, on the right, to the ahomajum or fourth ftomach. The omafum proceeds into the fuperior part of the laft ftomach, the abomajuin^ which is much larger than the preceding two, but lefs voluminous than the paunch : it is fituatcd under the omafum, between the bonnet and the paunch : it is fhaped like the bag of a bag- pipe. The aperture which forms a communication be- tween the omafum and the abomafum is furnifbed with a valvula which prevents the return of the aliments. The interior part of this cavity is lined with a thic\ mucous membrane, which forms fe- veral longitudinal folds. This ftomach is conti- nued with the duodenum by a pyloric orifice higher than the preceding. The aliments coarfcly ehewed at firft are formed into balls, and conveyed fpeedily into the bottom of the left cavity of the paunch, from which they proceed to the bottom of the cavity on the right, and place themfelvcs immediately below the cefo- phagian aperture, in the order of their entrance. Thefe aliments become foft and penetrated with juices. When the animal has taken a proper quantity, and is in a ftate of reft and of good health, it has the faculty of making them afcend SYSTEM OF DIGESTIOIN, ^Ql into the mouthy where they are again mafticated : for this purpofe a portion of the grafs contained in the paunch palles into the cefophagus; the part which is moil mafticated falls into the bonnet, and the coarfeft penetrated with juices is reduced to a ball, and returns to the mouth. When the latter phaenomenon takes place, it is obferved that the animal extends its neck, makes a flrong infpi- ration, and by the firaultaneous acSlion of the fides of the paunch, of the abdomen, the diaphragm, and the cefophagus, it caufes the ball to re-afcend. with rapidity along the oefophagian canal. When it reaches the mouth it is chewed again much more llowly, and with great care ; and the animal feems to find in thefe alimentary balls, penetrated with juices, a favour which renders the maftication of them agreeable. Rumination, therefore, is produced only by the firft flomach. The alimentary balls, triturated and reduced to a fine pulp, re-defcend along the cefophagus, and pafs into the gutter ; the coarfeft fall into the bonnet, the reft remain between the furrows of the omafum, and the moft liquid pro-? ceed dire6lly into the abomafum. The adion of the bonnet prepares the aliments for being received into the omafum, which foftens them, and renders them fufficiently liquid to pafs into the abomafum, where real digeftion is per- fijrmed. u a When '202 ORGAMIC STRUCTURi, When ruminating animals drink, or take very liquid aliment, and during ladlation, thefe fluid fubftances are conveyed through the oefophagus into the gutter, the edges of which unite to form a du6t, and they proceed dire6lly into the omafum, from which they pafs into the abomafum. The paunch of the camel exhibits a peculiar dilatation, furniO^ied with falient lamincB, like the omafum, and in which it can preferve, for a very long time, a large quantity of water. 293. The intejiines. That part of the alimen- tary canal, which is below the llomach, varies both with refpedl to its length and to its dia- meter. In animals, the length of the inteftines is in the inverfe ratio of the nutritive quality of the ali- ments on which they feed. In fome very ravenous fifhes, fuch as the ray, the inteftine is fhorter than the body. In the carnivorous mammalia, it is twice or thrice that length ; in' the herbivorous mammalia with a An- gle ftomach, it is equal to ten or twelve times that of the body. The capacity of the abdomen, in graminivorous animals, is for this reafon much greater ; while in the carnivorous the contrary is the cafe. In reptiles, the inteftines are of the fame width throughout their whole extent ; in birds, the width goes on decreafing towards the ret'rlum. 8 h\ SYSTEM OF DIGESTION, ^QS In the mammalia, the inteftlnes are flender in the firft part of the alimentary canal, and large in the latter. The large inteflines often exhibit flefhy circular bands, [which produce conftri6lions5 and make them appear as if fwelled up. The inteftinal canal, in different parts of its extent, has one or more lateral appendices en cuU de-fac : c cecum. In reptiles the caecum is wanting. The mam- malia have one ; birds have two ; and fifhes have often a very great number. The coecum of the mam'Tialia is placed at the commencement of the large in'eftines ; in carni- vorous animals it is very fhort ; in the gramini- vorous it is much larger ; and in the rodentia, it has often more extent than the ftomach. In man and fome apes, the caecum, which is fhort, has at its extremity a fmall vermiform ap- pendix. Birds have often two coecums on the fides of the reftum. Some fifhes have no caecum ; while others have a prodigious number, placed in general near the pylorus : they are thick and glandular, and feem to be organs which fecrete a liquor proper for di- geftion. The inteftine terminates, for the moft part, in an aperture furnifhed with a fphiniSter. u 3 The 2Q4 ORGANIC STRUCTURE. The anus is wanting in the zoophytes, which have only one aperture for the ahmentary canal : It is commonly placed at the extremity of the trunk or the tail j but in the mollufca it is found on the fides of the neck. The wholeof the alimentary canal is lined with a mucous membrane, which fecretes a digeftive juice. The inteftines receive alfo, fordigeftion, the bile fecreted by the liver, and the liquor produced by the pancreas. 294. The liver exifts in all vertebral animalsj, and in all the mollufca; in infers it appears under the form of a vafcular reticulation. The fize of the liver appears, in general, to be in the inverfe ratio of that of the organ of refpi- ration^ In all animals the liver occupies a great part of the abdomen ; the venous blood it receives is that which returns from almofi: all the abdominal vif- cera ; the veins of all thefe vifcera unite into a very large trunk (tl)e vena porta;) y which proceeds into the liver. The moft important ufe of the hepatic appara- tus is, not to fecrete a fmall quantity of bile, but to, receive a great part 'of the venous blood, and to make it undergo peculiar changes ; fo that the hiftory of the liver belongs rather to circulation than to digeftion. However, as the bile is a liquor SYSTEM OP DIGESTION. 205 of great importance to digeftion, the organ which fecretes it is here worthy of attention. The hver exhibits five orders of vefTels : ill, the large arterious vein, the Juh-hepattc or ve7^a forta^ which conveys its blood to the liver ; 2d, a fmall artery, the hepatic ; 3d, the large vein which conveys back the blood, the Ju'per-hepat'ic-\ 4th, the biliary vefTels ; 5th, the lymphatic vefiels. The canals which carry the bile from every part of the liver unite into one or more hepatic dadls, which open into the intefi:inal canal near the pylo- rus, and very rarely into the flomach. A part of the bile often proceeds to and remains in a bladder annexed to the liver. In the mollufca there is no bladder ; it is want- ing alfo in fome of the mammalia, fuch as the flag and horfe ; in fifhes it is found fometimes in the infide of the hepatic organ. The bile is carried to the bladder two ways: ifi^, it is conveyed dire6lly from the liver by peculiar du6ls, the hepato-cyjiic, as is the cafe in birds ; id, the hepatic canal has a communication with a peculiar cyfi'ic du61, along which a part of the bile flows back to proceed to the bladder, as is the cafe in the mammalia. In the firft cafe, the bile proceeds direclly from the bladder to the intedine by the ducIi/T choledo'^ chus ; in the fecond, it re-defcends from the cyftiq du6t to pafs into the duBus choledochus. V 4 295. The 296 ORGANIC STRUCTURE. 295. Tht fpleen is found in all red-blooded ani- mals ; the blood which returns from it proceeds to the liver : no other ufe of it is known than that of funiifhing all its blood to the hepatic ap- paratus. 296. The pancreas is a glandular body which fecretes a liquor analogous to the faliva ; this liquor proceeds into the inteftine by a particular dud)", which opens near the dudus choledochus . The pancreas is found in almoft all animals which have a liver ; it is wanting, however, in the mollufca, and in all fithes which have a great number of coecums. Birds have two or three pancreases, which pour their liquor into the inteftine by peculiar du6ls. 297. reritonaum. The gaftric and other organs contained in the abdomen are enve;loped by a dou- ble membrane, called the peritonaeum. In the mammalia, the peritonceum is of a vafcular texture, lax, tranfparent and greaty : in fifhes, it is of a clofe texture and opake, has a brilliant metallic appearance^ and is covered with a black varnifh. The exterior membrane of the peritonaeum lines the tides of the lower belly ; the interior feems to be prolonged different ways to envelop each organ^ and to retain it in a determinate. The SYSTEM OF DIGESTION. 2p7 The portion of the peritonaeum which covers the inteftines, forms, after it has enveloped them, a loofe and fmooth raembrane, which keeps them fixed to the vertebral column : the mefentery, ■ The periton^euni produces alfo a large mem- branous fold, which covers like an apron the ante- rior part of the inteftines: the eppJoon (omentum). In man, the epiploon defcends as far as the abdomen, and on the bladder i in apes, which have an open inguinal ring, it falls back towards the pelvis, and forms a fecond covering. Animals fubjedl to winter-lleep have ieveral epiploons, which become charged with fat : this fat feems to ferve for their nutrition during their ftate of torpor. Birds have no epiploon ; their large membra- nous and aerian bags become charged with fat. 298. System op digestion in man. In man, the fyftem of digeftion holds a mean plac«, in feveral refpe61:s, betv/een that of carnivorous and that of graminivorous animals. The fides of the mouth are formed, above and before, by the maxillary bones, and behind by the palatine bones, the pterygoid apophyfes, and the guttural fofla j below, by the under jaw and the teeth. 299, The teeth. Man has two incifor teeth, one angular {^cufpdatus), two fmall and three large molar teeth, on each lide of each jaw. In 2(38 ORGANIC STRUCTUHE, In the embryo, the rudiments of the teeth are- fcarcely apparent ; they are nothing then hot a mucilage confounded with the other parts ; and do not expand till towards the fourth month of geilation : at that period they prefent themfelves i]nder the form of a follicle or membranous bag, traverfed by a very great number of veffels, and iilled with a gelatinous mucus. The follicles of the incifors firil appear, then thofe of the angular teeth, and in the laft place the great and fmall molares. Thefe follicles, which are of an ovoid form, adherent to the slveoli, are at firft reddifli ; they then become v/hite, affume a cartilaginous conliftence, and ex- pand by a point of offification. The furface which correfponds to the body of the tooth begins to be covered with a thin cortical coating of enamel. Towards the fixth or eighth month after birth, the lower incifor teeth and the fuperior pafs through the gums, and fhovv themfelves without i the angular teeth appear towards the end of the firil year, and' the fmall molares at the end of the fecond. Behind the follicles of thefe firfl twenty teeth there are others of a fimilar kind, which gradually increafe, and about the lixth or feventh year * aflume the place of the firft. At that period the roots of the firft teeth, in a great meafure deftroyed^ * All thefe periods are fubje ceeds downwards and backwards. The pharynx exhibits before, in the middle of its paffage, and below the os hyoides, the entrance of a canal which conducts to the lungs : the larynx. The aperture of the larynx, glottis^ is fhut by a fort of femi-circular, cartilaginous, elaflic valve (epiglottis), affixed before towards the root of the tongue, and which proceeds obliquely upwards. The epiglottis is deprefled towards the aperture of the^/o//ij-, by the pallage of aliment, the falling back of the tongue, and the elevation of the larynx at the time of deglutition. The pharynx defcends behind the larynx, and continues with the oefophagus ; it is lined by a fmooth membrane, furnifhed with a great number of glandulous follicles. Its fides are formed by different layers of mufcular fibres. The longitu- dinal fibres are attached fuperiorly to different points of the bafe of the cranium ; in the back part, before the large occipital foramen ; on the fides, to the fiyloid and pterygoid apophyfes 5 and before, towards the middle of the interior face of the jaw. Below, the mufcular fibres are affixed to the OS hyoides and to the cartilage of the larynx : thyroid. Several layers of thefe fibres are tranfverfe and obhque ; they crofs each other behind by a more compadl SYSTEM OF DIGESTION. 309 compadl tifllie, which forms a median line. This difpofition allows the pharynx to be fhortened, and to be contradied in every point. To the ftyloid apophylis is affixed a round muf- cle, which defcends and expands over the fides of the pharynx : it contributes to raife that part. Stylo-pharyngian. The pharynx rells againft the trachelian region of the rachis, on the mufcles which cover it; and is feparated from it only by a lax cellular tiflue, which is never greafy. The pharynx becomes narrower at its lower part, which correfponds to about the iixth vertebra of the neck ; it then continues in a fingle canal, which proceeds to the ftomach : the cefophagus. 322. The cefophagus is lined with a mucous membrane^ analogous to that of the pharynx : its lides are formed by two layers of pale mufcular fibres : the exterior, which is thicker, has its fibres longitudinal : thofe of the interior layer are thin- ner, and difpofed in a circular or fpiral form : they can fhorten or contra(5l that cavity. The cefophagus defcends a little to the left, be- fore the bodies of the laft cervical vertebrae; it then penetrates into the breafi, and inclines gently to the right, along the bodies of the dorfal vertebrae ; it then traverfes the diaphragm behind, and ter- minates in the abdomen by a wide aperture, the, pefo'^hagian, which is continued with the Aomach. X ^ 323. The 310 ORGANIC STRUCTURE. 323. The JiQmach, fomewhat wide at firft, is contained in the left hypochondre : it is confider- ably dilated, and is folded back on itfeif, fornning a large convexity : th^ greater curvature : it then proceeds tranfverfally to the right into the epi- gaftric region/ where it becomes narrower, and exhibits a fmall convexity: the hjfer curvature; after which it terminates in an aperture inclined down- wards and to the right. Pylorus. The ftomach, therefore, exhibits the form of a cone folded back on itfeif, and fituated in a tranf- verfe, oblique direction below the diaphragm. It has two apertures. One of them, the cefophag'ian^ is dire6led upwards, forwards, and to the left : the other, the, pyloric, which is fmaller, is turned down- wards, backwards, and to the right. It therefore exhibits two curvatures, from the one orifice to the other : one of thefe, vyhich is fmall, is turned up- wards and backwards; the other, which is larger, is turned downwards and forwards. The two aper-r tures of the flomach are very near each other in the foetus, which renders it, in fome meafure, fpherical. In the flate of plenitude it moves up- wards: its large curvature is entirely before, and its lets behind. The ftomach is every where covered by i^ferous membrane, furniQied by the peritonaeum. The part of this membrane which covers the gafiric organ before, is united to that Vvhich covers it be- hind. SYSTEM OP DIGESTION-. '311 hind, towards the lefler and greater curvatures, where thefe membranes are applied one to the other, and form avafcular, plaited and adipofeexpanfion. The fold whieh is towards the fmall curvature (lejfer epploon) comes from the portion of the peritoneum which covers the liver: that formed towards the large curvature (the great epiploon) \^ very extenfive ; it defcends before the abdomen, and covers a great part of the inteftines. When riie ftoniach is diftended by the prefence of aliment, the laminse of which thefe folds are formed towards the curvatures of the llomach fe- parate, and permit that organ to aflume great am- plitude. Below the peritonaea! membrane, the fides of the ftomach exhibit two layers of mufcular fibres : the exterior extend from the one aperture to the other ; thofe which are below afTume a circular and tranfverfe diredlion. In t^e laft place, the ftomach exhibits interiorly a thick mucous membrane, full of villofities, which give it a velvety appearance ; it has in the infide folds which difappear in the ftate of dilatation. The fides of the ftomach become much thicker towards the pyloric aperture, where the interior membrane forms a thick membranous roll, which contains mufcular fibres in the form of a ring : this roll produces a fort oi^ pyloric valvula. ^^4. The intejlines. The fi:omach is then con- X 4 tinuecl 312 ORGANIC STRUCTURE. tinued with the inteftinal canal. This canal is twifled, folded back on itfelf, and is narrow at the top '.fmall iniejiines : towards the anal extremity it is larger : the large 'miefrmes : its length is about fix or feven times that of the body. The firft part of the intefline, tlie duodenum, which is continued with the ftomach, is fufceptible of great dilatation ; it defcribes different curves, and is in part unprovided with a peritonseal mem- brane. It has one and often two fmall apertures at the diftance of fome inches below the pylorus : this portion proceeds backwards, downwards, and to the right ; then afcends, and continues with a long portion of the inteftines : \he jejunum and the ileum. Thefe inteftines form a great number of circum- volutions, and occupy the whole anterior, middle, and lateral part of the abdomen. They;//*^// inteftines are continued with the large inteftines, firft by a fhort thick portion called the coecum\ it is fituated in the right iliac region, and at its entrance is furnifhed with a circular membra- nous valvula, which oppofes the return of the fascal matters : this portion of inteftine has a lateral ap- pendix, the cavity of which, called the ccecal, termi- nates like the finder of a glove : the coecum. This fhort portion of inteftine is continued with one much longer called the colon: the latter is of the faiijc fize, and is twifted around all the fmtill guts ; it rifes from the right iliac region, traverfes the SYSTEM OF DIGESTION. 3]3 the abdomen, below the Horn ach, defcends along the left fide, is refleded on the praelumbo-trochan- tinian mufcle, and is continued with the laft part of the inteftinal canal. The latter, called the reBum, is large, fhort, and upright, and terminates by a mufcle, the annular fibres of which clofe exa6tly its aperture. Interior fphin6ler of the anus. A thin mufcle is placed in the cellular tifTue, which lines the anus : it is inferted in the coccyx, by one of its extremities, and at the other is con- founded with the bulbo-urethral or cavernous muf- cle : by its action it tends to clofe the anus. Coc- CYGIO-ANIAN, exterior fphm£ler. A large thin mufcle forms the bottom of the pelvian cavity : it is affixed at the top, behind the pubis, to the upper part of the fub-pubian hole, and the fpine of the ifchium ; and below to the fides of the coccyx : it then unites with that on the oppo- f-iit fide, by an aponeurotic line, and embraces the lateral part«. It elevates the anus. Sub-pubio- CoccYGiAN, elevator of the anus. The flru6lure of the fides of the intefl:inal canal is analogous to that of the fides of the fiomach. The inteftines are covered by the ferous membrane, which en- velops all the vifcera of the abdomen : peritonaeum. This peritoneal membrane, after having enve- loped the intefiinal canal, is refleded backwards, and forms a double wrinkled, vafcular, adipofe ex- panfion of the breadth of the hand, and much fhorter 3'!4 ORGANIC- STRUCrURE. Shorter than that canal : it is affixed before the bodies of the dorfal and lumbar vertebras, and thus fecures the inteftines. A part of this mem- branous expanfion retains all the fmall intef- tines in a bundle in the middle of the abdomen t mejentery : the other part confines the large intef- tines around the former : mefo-colon. The double membrane of the mefentery is fufceptible of being feparated near the inteftines, when the latter ac- quire more amplitude. Below the ferous membrane of the inteftines is a- double layer of very thin mufcular fibres: the exterior longitudinal ones are difpofed in feparate bands; the interior ones, which are circular, are alio very thin. The longitudinal layers, which are little fufceptible of exteniion, confine the fides of the canal, and produce thofe gibbolities which are obferved in the large intefiines. Theoutlide of the alimentary canal is furnifhed %vith a mucous membrane, analogous to that of the ftomach. At the entrance of the duodenum is obferx'ed one or two fmall apertures, v^hich pour into the inteftine the produ6lof the fecretion of the liver and of the pancreas. 325. The liver is a glandulous organ of a eon- fiderable fize ; it occupies a great part of the ab- domen,. aod- is fituated in the right hypochondre, the epigaflric region, and a part of the left hypo- chondre, below the diaphragm^ &;c. This organ- receives SYSTEM OF DIGESTICm. 315 receives the venous blood, vv^hich comes from the ' principal vifcera of the abdomen. The blood by traverlingthis gland undergoes important changes; it is freed from thofe greafy and albuminous mo- leculsg which it contained in excefs, : thefe fub- ftances, by their combination with a fmall quantity of foda, form the bile, which proceeds from every point of the liver by a peculiar order of veflels. If it be confidered that the liver is the heaviefi: of all the organs, and that the great quantity of venous blood which it continually receive'?, under- goes in this organ the moft important changes, it may be readily conceived that a minute defcrip- tion of its vafcular ftru6lure belongs rather to the fyftem of circulation than to that of digeftion. We ihall, therefore, introduce here only what relates to the excretion of the fmall quantity of bile pro- duced by the materials taken from the large quan- tity of blood which pafles through the liver. The bile is excreted in all points from the hepatic organ by ramifications, rami, and branches^ which decreafing in number, and increafing in calibre, unite together, and iffne from its tranfverfe furrow by two or three trunks, which terminate in one du6l called the hematic. The hepatic du6l, after proceeding a certain; way to the left, communicates with another du6l, the cyjiic, which is refledled upwards and to the \e% at a very acute angle, and proceeds to the gall- bladder. 3l6 ORGANIC STRUCTUK.E. bladder. Thefe two da6ls form, by their union, a larger du(5):, called the du£fus choledochus, which proceeds to the left, and opens into the duodenum. The cyjilc dud, which is an inch in length, is terminated by a dilatation that conflitutes the gall- bladder. This pyriform veficle is lodged in a depreffion of the gaftric face of the right lobe of the liver ; it touches the colon, the commencement of the duo- denum, and the pancreas. The exterior tunic of it is formed by a prolonga- tion of the peritoucBum, in every part of it which does not adhere to the liver ; the interior fide is villous, interfperfed with folds, and exhibits all the chara61ers of the mucous membranes. The du^us choledochus is a continuation of the hepatic and cyftic du61sj its length is about three iinger-breadths ; it defcends, on the left, before the fub-hepatic vein; is inferted obliquely between the tunics of the duodenum \ often receives in that part the pancreatic dudl ; and at length pierces the villous tunic of the intefline, two finger^ breadths below the pylorus. The (Irudure of the hepatic, cyftic, and cha- ledocb du(9:s is nearly the fame as that of the gall-bladder. 326. The I'ik, which proceeds from all parts of the liver, runs along the hepatic du6V, as far as the dudus choledochus j the la,tter pours it into the SYSTEM or DIGESTION. 317 the duodenum : but when it does not flow into the duodenum, it may flow back by the cyilic dud, and be depofited in the gall-bladder, where it acquires more confiflence, and aflumes a darker colour. The bile is a liquid of agreenifh yellow colour, fat, and foft to the touch, vifcous, exceedingly bitter, mifcible with water like foap, and foluble in oils, alcohol, and ether. The chemical corapofition ot' the bile may be afcertained by the following mode of analyfis : Pour dilute fulphuric acid over this animal liquor, and expofe it to heat. You muft then obferve whether precipitation takes place, and whether a ialine fubftance remains in the liquor : the preci- pitate muft be Separated by filtration. Ths liquor, when properly evaporated, will give cryftals of fulphate of foda. Then treat the preci- pitate with alcohol, which will diflblve a part of it : the other is to be feparated by the filter. If the alcohol be evaporated, a pure oily matter will be depofited : the other part of the precipitate, which remains on the filter, is albumen. Hence it refults that bile is eflentially formed of albumen, oil, and foda, in a kmd of faponaceous fiate. Difi^erent falts alfo are found in the bile^ in fifiall quantity, and particularly phofphate of foda. The bile very often produces concretions in the gall-bladder. 327. Bi- 3lg dRdAKiC STRUCTURE. 327. Bilious coricretidns are fometimes very nu- merous; they afFeci, for the moft part, a tetrae- dral pyramidal form, and, when there are feveral;, become polyedra by fri6tion. They are brown, green, or yellow; exceedingly light, and burn^ emitting a thick fmoke. They affe6l three modes of compofition. Some are irregular, and confifl: of grains united around a nucleus : others are angular, and produced by concentric ftrata of different denfities, and often very compad. The lait kind are ovoid, exceed- ingly hard, covered by a white coating, are brilliant, and appear to be^ cryftallized : this kind are infolu- bte in alcohol ; they are formed of adipocera ; the reft are of the nature of bile. 328. Thefplem. The blood which proceeds to the liver by the fub-hepatic vein comes in part from the fpleen ; this vifcus, which has an ovoid form, is of a blueifh red colour. It is placed lengthwife in the left hypochondre, and is co- vered by the peritonaeum, the folds of which fix it to the diaphragm, the flomach, and the mefo- eolon. The fpleen is convex on the outfide, and fome- what concave within, v*?here it exhibits a longitu- dinal fciflure filled with fat. The artery which proceeds to this organ (the left branch of the fub- gaftric or coeliac trunk) is very large, twifled, and penetrates into its fubftance near the fciflure; The SYSTEM OF DIGESTIOS?. 3Jg The veDOus trunk, which returns from that organ, IS united to the mefenteric vein ; and both form the fub-hcpatic vein^ vena porLe^ which proceeds to the liver. No other ufe is yet known of the fpleen, than that of conveying all its venous blood to the hepatic organ, 329. The pancreas is an oblong gland, very inuch analogous to the falivary glands. It is placed, in a tranfverfe diree'-lion, in the left hy- pochondre, below the ftomach, between the liver and the fpleen, on the pofterior aperture of the two membranes of the mefo-colon, which cover it before: behind it is enveloped only by cellular tiffue. The pancreas is of a pale rofe colour, and ap- pears to be formed of lobes enveloped by a cellular membrane. Thefe lobes are an aflcmblage of lobules and glands indeltinitely divided, and are alfo covered by a ftratum of cellular membrane. Acrofs the middle of the pancreas runs a du(5t, on the fides of which fome exceedingly fine fecreting duels terminate. The pancreatic duel proceeds to the right, and opens into the ductus choledo- chus, or very near it, in the duodenum. The fluid fecreted by this gland appears to be very analogous to faliva. 30. The peritoneum, v/hich, as already faid, envelops the whole vifcera of the abdomen, and the 320 ORGANIC STRUCTURE, the fides of that cavity, forms a common covering* Its ftru^lure is fuch, that anatomifls compare it ta a membranous bag every where fhut. The cavity of this bag forms the fpace comprehended between the inteftines and the fides of the abdomen. This cavity, the fides of which are contiguous, contains nothing but ferous matter. The exterior face of this bag lines before, and laterally, the interior fides of the abdomen ; and behind it envelops, in its numerous folds, the vif- cera contained in that cavity. The ligaments which fix the vifcera of the ab- domen to the neighbouring parts, are, in a great meafure, only folds of the peritonaeum. SYSTEM JM [ 321 ] SYSTEM OF CIRCULATION. :3:gi. In circulation, the blood, proceeding fron> a certain point, is conveyed by one order of veilels to all the parts of the body, and diftributes to them the materials neceflary for the different changes which are there continually taking place : it is then conveyed back to the point from which it proceeded, by one or two orders of veffels. r; In this courfe, the blood paiTes either in whole or in part through peculiar organs, which tend jn an eflential manner to maintain it in a pro- per flate of temperature and of liquidity, or to free it from the reiiduums of organization ,with which it is charged, or in the laft place to reft ore to.it thofe e]ffential materials which it has loft. All thefe organs neceflarily compofe a part of the iyftem of circulation. 332. Circulation. In plants and zoophytes no diftin(5lion is remarked in the order of circula- tion, or in the veftels : the fluids feem to proceed to the different parts by a kind of fudlion or abforp- tion. Infe^ls have a long dorfal tranfparent vcffel endowed with periftaltic motion : the relation of this veflel to the other parts is not known : be- voL. I. Y fides. 322 ORGANIC STRUCTURE, fides, the tranlparency of all the organs of thefc animals renders it impofiible to afcertain their in- timate llrudure, and we are ftill unacquainted with the nature of their circulation. There are . obferved, however, in thefe animals organs of fe- eretion for the bile, feminal liquor, &c. "which fcem to be formed of floating vafcular packets. In the worm and leech, a dorfal veflel which performs the office of a heart is obferved. In cruilaceous animals, fuch as crabs, the mo- noculus or water-flea, &c. the heart receives the blood which returns from the branchiae, and dif- tributes it to every part of the body ; but it is never feen to return to the branchiae. In the bivalves, the blood proceeds into an au* ricle, which pours it into a ventricle, whence it iflues by an aorta. Mufcles exhibit a very Angular organization i they have two very large tranfparent auricles, and the inteftinal canal pafles through the heart. In the aplyfia, the blood, which comes from the branchiae, unites in a vein the fides of which be^ come arterial. This vcflel then performs the office of an aorta. Slugs have on each fide of the body two con- tradlible apertures, which communicate with two pulmonary cavities, over which the veins are fpread. Thefe cavities terminate in an inteftinal canal I the blood, after traverfing the pulmonary organ^ SYSTEM OF CIRCULATION. 323 organ, unites in a vein which proceeds into an auricle; whence it palles into a ventricle farnifhed with valvulas ; and from this ventricle it iflues by an aorta, which diftributes it to all the parts of the animal. In the fcpia, a large vein divides itfelf into two branches, which proceed to two auricles farniflied with a valvula. Thefe two auricles are feparated, and furnifh an artery which is diftributed to each branchia. The blood comes from the branchiae by two veins, which unite in a common trunk ; and this trunk proceeds to a folitary ventricle, from which the blood iffues through an arterial aorta. 333. In the different clafles of animals here mentioned, the blood appears under the form of a ferous liquid, more or lefs whitifli, and feldom red. It feems to return from the different parts in the fame manner in which it was conveyed ihither ; that is to fay, by one order of velTels. But in the other clades of animals, which we are about to examine, the blood is red ; it is diflri- buted to all the parts by one vafcular fyflem (the arteries), and returns by two orders of veflels : one of them (the veins) brings the greater part of the blood to a red flate ; the other, much lefs ap- parent (the lymphatic veflels), colle<51s the ferous refiduums, and the produ6l of digcftion which it pours into the veins. In tithes, two ven^ cavag pour the blood into Y 2 an 324 ORGAKIC STRUCTURE, an auricle, whence it paiVes into a ventricle fur- niilied with two femilunar valvule© (one at its entrance and the other at the place of exit). The blood iiTaes from' this ventricle through an artery which divides itfelf into four branches on each fide. Thefe arteries are diftributed to the branchiae. Each branchial artery afcends along each lamina of the branchi^: when it arrives towards the middle of them it divides itfelf into two rami, one on the right and the other on the left : each of thefe rami then divides into two others^ one of which defcends, and the other ateends, the whole length of the lamina. The laft branches furnifli a great number of very fhort ones, which proceed acrois and cover the whole furface of the lamina. The blood returns by veins, which in uniting follow the fame order as the arteries do in dividing. The union of thefe veins forms a large dorfal veflel, which performs the office of an aorta. In reptiles, the lungs receive only a part of the blood. In animals of this clafs fubjedt to meta- morpholis, fuch as frogs, fyrens, and falamandersj two venae cava? pour the blood into an auricle furniilied with valvulse, whisnce it ifTues by an ar- tery furniflied alfo with a valvula. This artery, which divi^les itfelf, conveys the greater part of the blood to a ventricle; the other part proceeds to the pulmonary organ, from which it returns by the veins to the fame ventricle ; and it then ifTues from that cavity todiftribute itfclf to all the organs, 111 SYSTEM OP CIRCULATIOJT. ^ 325 In the ftate of tadpole (without paws and lungs) thefe animals have on each fide of the neck a branchia in the form of a plume, which receives the blood; but as thefe animals increafe in growth the branchiae become obltru(51cd, wither, and drop off along with the tail: at the fame time the lungs expand, and pulmonary refpiration takes place. The tortoife has tw9 auricles and two ventricles, which communicate with each other. Three venas cavas pour the blood into one auricle^ furnifhed vj^ith valvulse, whence it pafles into a ventricle to be diftributed to the pulmonary organ : it then returns by veins to an auricle, and pafles into the fecond ventricle- This fecond ventricle has a communication with the former, by an aperture clofed by a valvula. This aperture correfponds to the arterial aorta, and the arterial blood proceeds direclly thither without mixing much with the venous blood, though thefe two forts of blood are together in the fame cavity. In the mammalia and birds, the blood proceeds to an auricle which pours it in:o a ventricle fur- nifhed with valvuloe. From this ventricle it paffes through the lungs, and returns by veins which convey it to a fecond auricle, whence it pafles intoa fecond ventricle provided v.*ith valvuls : it ifflies from this cavity by the aorta, which diftri- butes it to the different parts. In ihc fiate of fcstus, the mammalia and birds ^ y 3 exhibit 326 ORGANIC STRUCTUkE. exhibit another order of circulation. The blood poured into the pulmonary auricle pafles, in part, into the aortic auricle, by an aperture furniflied with a vajvula {foramen ovale). The remainder of the blood flows into the pulmonary ventricle, whence it iffues by the artery of the fame name. At that place a part of the blood patTes through a du6t, which eilablifhes a communication between that artery and the aorta (arterial du6l) ; the reft traverfes the lungs, and returns by the veins. The latter pafles through the aortic auricle and ventri- cle, and mixes in the aorta vi'ith that which pro- ceeds diredlly thither through the arterial duel. The circulation in the foetus of the mammalia and of birds refembles that of reptiles, as only a part of the blood pafles through the lungs. In cetaceous and amphibious animals, the aper- ture which forms a communication between the two auricles, clofes more flowly ; but at length it always becomes obftrudted, as in the mammalia and birds. 334. It is obferved in all clafles of animals, and even in plants, that the nutritive fluid, or the blood, has a temperature peculiar to itfelf. In the greater part of animals this temperature is not much different from that of the medium in which they habitually live ; but it is conftant, and does not directly follow the variations of thefe me- diums. Iq SYS.TEM OF CIRCULATlOJt* 327 In animals with lungs^ through which the whole of the blood pafles^ fuch as the mammalia and birds, the temperature of the blood is much higher than that of the atmofphere of temperate climates. This temperature, which is about 40 degrees of the centigrade thermometer, varies only a degree or two in the moft oppolite latitudes. 325' The principal changes which the blood ex- periences, during its circulation, take place while it palies into the kidneys, the liver, and in parti- cular through the organs of refpiration, 2^6. Urinary system. The blood is freed from its excefs of aqueous parts, and from different faline fubflances, by means of the kidneys. All red-blooded animals have kidneys ; white- blooded have nothing analogous. The kidneys are nearly of the fame relative lize in all animals ; they are however more volumi- nous in birds. The renal fyflem, in general, confifts of one large gland on each fide. The urine fecreted from every point of this gland ifTues from it by fmall duels, which proceed to a common canal, called the ureter, as in fiflies, reptiles, and birds. In mofl of the mammalia, the urine, which pro- ceeds from every point of the kidneys, oozes through fmall funnel-like capfules : feveral of thefe capfules unite, and proceed into fmall com- Y 4 nion 32S ORGANIC STRUCTURE, nion refervoirs or bafons, from which the urine af- terwards pafles into the ureter. The kidney, fometimes, is compofed of feveral fmall glands united without refervoirs, each hav ing its particular ureter ; as is the cafe in the bear^ otter, and cetaceous animals. The ureters either convey the urine directly out- wards, or pour it into a bladder; from which it after- wards iiiiies by a fecond canal, called the urethra. Birds have no bladder : the ureters proceed into a cavity, common to the excrements and the eggs, cloaca ; and iffue through the anus *. A bladder is found in fifhes, in reptiles, and m- the mammalia. In fiflies it is fituated behind the redlum ; in reptiles it is placed before, and is fometimes double, as in frogs : in thefe two clafles of animals the urine proceeds to the rectum. In all the mammalia the bladder is placed be- fore the re6lum ; and the canal of the urethra palles through the parts of generation. 337. Hepatic system. The blood, on its pallage into" the liver, is freed from feveral fub- iiances, and in particular from fat and albuminous matters which it contains in excefs. The liver feems to exift in infetts under the form of a vafcular reticulation : it is found in the * In the oftrlch, the ureter proceeds direftly outwards, near the aperture of the fphuifter ani, and behind the cloaca. . , moliufca SYSTEM OP CIRCULATION. S^Q mollufca and in all vertebral animals. Its fize in general is in the inverfe ratio of that of the organ of refpiration ; which has induced fome to believe that thefe two organs have analogous fundlions, and that they can, in fome meafure, fupply the place of each other in their adlion on the blood. The liver^ in general, is exceedingly large in the mollufca; v*ry voluminous and oily in fifhes, but fmaller in the mammalia and birds. It is ob- ferved in the foetus of the laft two clafles of ani- mals, that the liver, comparatively fpeaking, is of a conliderable fize, and that the fize decreafes after birth in proportion as the organ of refpiration is expanded. The liver, in general, is divided, to a greater or lefs depth, into two or more lobes. The venous blood, proceeding into the hepatic organ, is that which returns from the principal ab^ dominal vifcera by veins that unite into a large trunk, Xh^fith-Jpepaik vein or ve7ia porta:,, On its padage through all the abdominal vifcera, which in general are covered with fat, the blood becomes charged with adipofe and albuminous m.atters, of which it is in part deprived in the liver. The bile, which refults from this particular excretion, be- comes afterwards one of the mod powerful agents of digeftion. The blood, when it has thus palTed through the liver, and been freed from a part of the fubilances 5 combined 330 ORGANIC STRUCTURE. combined with it in excefs, returns by numerous veins, which unite and proceed into the vena cava« 338. System op respiration. It is in the organ of refpiration, in particular, that the blood undergoes the mod remarkable changes ; it is there that it is at lafl freed from the fubftances foreign to its compolition, and acquires the prin- ciples it has loft by diftributing itfelf to the dif- ferent organs. It is in this organ that the blood pafles completely from the venous to the arterial ftate, and acquires all the qualities neceffary to fit it for a new circulation. The fyftem of refpiration brings the blood con- flan tly into relation with the circumambient fluid. There are three forts of refpiratory organs : the tracheae, branchiae, and lungs. 339. Tracheae. In the cortical part of vegeta- bles, but dill better in the ligneous, are ob- ferved fmall thin laminae, twifted into a fpiral form, and lengthened out like a fpiral fpring. Thefe trachese may be ealily feen on half breaking by flexion a young twig. It has been fuppofed that the circumvolutions of thefe laminae muft form a hollow cylinder, and that they conftitute air- veflcls : this hypothelis is not fupported by proofs. Tracheae, or elaftic tranfparent filaments, dif- pofed in a fpiral form and fufceptible of being un- rolled. SYSTEM OF CIRCULATION". 331 rolled, are obferved alfo in different parts of in- fe61s. In feveral of thefe animals, fuch as flies, bee?, the cock-chafer, and beetle, are obferved two holes or wide cells, which end at two refervoirs: the air which fills thefe refervoirs pafTes into the trachese. In thefe animals, the tracheae are fubdivided, and diftribute themfelves to every part of the body ; they have a brilliant filvery appearance, and often produce a wonderful efFedt on their different organs. 340. Branchiae. Branchia; are found in the tad- poles of reptiles fufccptible of metamorphofis, aquatic mollufca, and fifhes. Reptiles in the fiate of tadpole have on each fide branchiae in the form of a plume : in propor- tion as the animal expands and acquires perfection, thefe branchiae become obftruc^ed, wither, fall off with the tail, and their place is fupplied by the lungs. All cruftaceous animals, from the coleoptera to crabs, have branchiae in the form of laminae. Aphrodite worms have branchiae in the form of laminae or of a plume ; other infed^s have tubes proper for inhaling the air. In the acephalous mollufcaj the branch icE are of different forms ; they are fituated either around an aperture, which ferves as a mOuth, as in the bivalves. 332 OHGANIC STRUCTURE. bivalves, or around the anus as in the doris, or around the body as in the aplyfias, he. In the gafteropoda, ilugs, and fnails, the branchi^ are lituated in a large cavity below the neck. In the cephalopoda, thel^ranchiaSi which confift of laminaSj are placed in a cavity on the lides of the belly. In filhes, the branchiaa are compoled of laniiniSj the number of v^'hich, in general, is four on each lide. The laminse are formed of long narrow leaves placed at the fide of each other, like the teeth of a comb^ and are united by a thin membrane to the half of their height. Each lamina is compofed of cartilaginous pedicles united fide by lide, for three fourths of their length. The laminae of the branchiae are fupported by s fmall hollow bone, on which refts the tongue, and which is articulated at the bafis of the cranium. 341. Lungs. In fome reptiles the lungs confill: only of a large membranous cavity, the inferior face of which is covered by veflels, as is the cafe in the filamander*. But, in general, the lungs of * The cameleon has only a large plaited pulmonary cavhyj, ■fthlch It can Iwell up and diftend to fuch a degree, that it confi- derably Increa'.es the volume of Its body : it then becomes tranf- pare:;:, and aiiumes different tints. A fjrailar conformatloa is obkrved in the marbled lizard.- reptiles- SYSTEM OP CIRCULATION, 333 reptiles are a fpongy body, the cells of which are almoft always vifible to the naked eye. Theie Iimgs are inclofed in a Ibrt of pleura, and float in the common cavity of the trunk, which has no diaphragm. The a?r penetrates into their lungs by a larynx, without an epiglottis. The aperture of the glottis is narrow in ferpents, and broad in frogs aud toads. The males of the latter have membra- nous bags, which become filled with air when they emit cries. The mammalia have on each fide of the breaft a lung formed of a very fine fpongy tifTue. The fanguine veflels overfpread and expand over the- fides of the fmall cells of this tifTue i and the air penetrates into the interior part of them by numerous ramifications of branchial du6ls. Each lung is enveloped and remains free in a membrane (pleura), which adheres to the fides of the thorax. The two pleuras produce between them a fpace which lodges the heart, the cefophagus^ the aorta and the thymus. The air penetrates to the lungs by two bron- chial du<5is, which are ramified in theinfide: thefe two dud.s are the d4vifion of the tracheal artery. The tracheal artery and the bronchise are formed of cartilaginous rings, incomplete behind, and united by a membrane. This trachea is con- tinued with the larynx, which is a cartilaginoi^s {dilatation. The 334 ORGANIC STRUCTURE. The larynx communicates with the lower part of the pharynx, by an aperture (glotlh) covered with a cartilaginous lamina {epiglottis). This la- mina falls down on the glottis, during the paffagc of the aliments ; it ferves alfo to make the air vibrate when it ifllies with force from the lungs, and is the principal caufe of the founds produced by animals. The larynx, in all the mammalia, is formed of iive cartilages, viz. the thyroid, the cricoid, the two arithenoids, and the epiglottis. But in fome animals of this order it exhibits very remarkable peculiarities, calculated to produce a variety of founds *. In the mammalia, the larynx is generally moved by four or five pairs of mufcles, proper for lengthening or fhortening the trachea ; and for di- lating or contracting the trachea and the glottis. In birds, the lungs are much more extenfivej -tbcy are not inclofed in a pleura^ but adhere to * In the orang-cutang the ventricles of the glottis are pierced •with an aperture, which terminates in two membranous bags. Mandrils have a membranous bag, the aperture of which is at the root of the epiglottis. In the howling ape, the os hyoidcs is Ihaped like an offeous box of the fize of the fift. The afs, at the. extremity of each ventricle, has a hole correfponding to a particular bag. The larynx of the fwine exhibits in its length a deep finus. The air expired with force enters into thefe dif- ferent cavities, and produces different founds. The horfe, near the thyroid cartilage, has a triangular membrane, placed in a tranfver- fal direftioHj which by quivering is capable of producing neighing. tlie SYSTEM OF CIRCULATION. 335 the fides of the thorax by a loofe cellular tifTue.' Thefe lungs communicate with feveral membra- nous bags, fituated chiefly in the abdomen, which fcem to be formed by a fold of the peritonaeum. They are generally fix in number, placed longitu- dinally between the jnteftines and the abdominal vifcera: three alfb are found in the breaft, and two under the axillae. The lungs have a communication with thefe bags by particular apertures ; and the bags them- felves are continued with other cavities of lefs fize, which are in the thighs, the wings, the fubfl:ance of the bones of the cranium, in the cavities of the long bones, and even in that of the tubes of the feathers. In birds, the larynx is compofed of fix or four ofl^ous pieces : the principal one is analogous to the cricoid cartilage of the mammalia ; but they have none which correfpond to the thyroid and arythenoid cartilages : they have alfo no epiglottis. The only fundlion of this larynx is to open and fhut the aperture of the glottis. The edges of its aperture are furnifhed with cartilaginous points proper for preventing the entrance of the ali- ments. The tracheal artery is formed of complete rings: inferiorly the two apertures of its bronchial bifur- cation exhibit, on their fides, a projedling mem- brane, which in part clofes them. This moveable membrane performs the office of an epiglottis ; it 9 caufes 336 ORGANIC STRUCTURE.' caufes the air to vibrate in its paflage, and in ihli place the found is formed : this apparatus conilir- tutes the inferior hirynx of birds. The rings of the bronchiee are incomplete^ and vary both in their iize and figure. The inferior larynx of birds^ in many of thefe animals, has no proper mufclcs : fcveral of the latter are furnifhed Vv'ith ofieous or membranous dilatations. Some birds have a mufcle which is inferted in feveral of the half-rings of the bronchia. Others have three pairs of mufcles, as is. the cafe with perroquets : in linging-birds and in feveral others there are five pairs. All thefe mufcles are difpofed in fuch a manner as to produce indefinite, varintions in the dimenfions of the larynx and of the bronchia?. 34.2. In man, the apparatus of circulation ex- hibits an auricle, which receives the blood return- ing from all parts by the veins and the lymphatic vefiels. This auricle pours it into a ventricle, which tranfmits the whole of it to the pulmonary organ. The blood then returns to another auricle and a ventricle, from which it is expelled through the aorta, to be diftributed to every point of the organization by means of the arteries. In this double and fucceffive circulation, through all parts of the body, and through al! thofe of the lungs, it is obferved that the blood un- dergoes very remarkable chai-ges from feveral of the organs. Duiiog SYSTEM OF CIRCltLATION. 337 JDiiring its arterial circulation, it frees itfelf from its excefs of heat by the aiTiftance of the ficin i and from its excefs of aqueous parts, by tneans of the kidneys. On its return through the veins, it is freed from albuminous and adipofe fubftaiices in the liver. On its paffage in the lungs, it gives off carbonic acid gas, and takes up oxygen gas. In the lad place, the ferous matter, which returns by the lymphatic veiTels, and the produ61 of digeftion con- veyed by the chyliferous vefTelSj experience eflen- tial changes on their pafiage through the lympha- tic glands. The fyllem of circulation is thus compofed, not only of cavities or vefiels which ferve fat convey- ing the blood to every part of the body, but alfo of organs, v^hich, during itspaflage, make it under- go changes of great importance to the order of the organic functions* In our defcription of this complex fyftem of organs, we fhall therefore fol- low the progrefs we have indicated in this fhort view of the parts of which it is compofed. 343. The arterial fydem exhibits a ramified expanfion, of which the aorta is the trunk. The fucceffive and indefinite ramifications which feem to proceed from this trunk, are divided in fuch a manner, that there is no part of the organization to which they do not penetrate, and to which they do not convey the repairing fluid they contain. The arterial fyftem; as it divides, increafes in capacity; VOL. I. z and 338 ORGANIC STRUCTURE, and the blood, which finds itfelf more at freedom, muft flow with lefs rapidity as it pafTes from the large arteries into the fmall ones. The arteries exhibit frequent anaftomofes, and are ex©eedingly flexuous, efpecially in the parts fufceptible of extenfionj fuch as the vifcera of the abdomen. The (ides of the arteries are of a clofe texture, compact, elafticj and little fufceptible of extenfion. Their thicknefs is comparatively greater in the fmall arteries than in the large ones. They feem to confifi: of three tunics, the exterior one of which is cellular, foft, and very loofe : the middle one is thicker, and conlifts of a clofe yellowifh tiflue, lit- tle fufceptible of extenfion : the interior one is only a very thin and exceedingly fmooth mem- brane. The blood contained in the arteries is fpumous, florid, and of a bright red colour ; it appears to be fomewhat warmer and liMiter than the venous o blood, and has not the brown tint of the latter. » The arterieSj after dividing and fubdividing themfelves into exceedingly fine and delicate ra- mufculi, which proceed to every part of the or- gans, at lad completely difappear to the eye^ even when afSfled by the befl microfcope, fo that their manner of termination is totally unknown. 344. During the arterial circulation, the excels of the heat of the blood is carried off by the fecre- 6 tion SYSTEM OP CIRCULATIOH. 33g tion of the Hun, reduced into vapour at the furface of the body : its excefs of aqueous parts is expelled, together with feveral faline fubftances, by the uri- nary fecretion. When the blood and all the parts which it moiftens have too great a quantity of free caloric, arifing either from a high temperature of the at- molphere, or from too violent exercife of the muf- cular fyftem^ or from fome peculiar morbific ex- citement, the cutaneous organ enters into a6iion and fecretes abundance of perfpirable matter, the evaporation of which at the furface of the body lowers the, temperature, and always maintains it at about the fame degree. This abundant per- fpiration frees the blood alfo from a quantity of aqueous matters, more or lefs conliderable ; it carries off alfo a little albumen, and feveral faline fubftances. The fkin properly fo called {dermis, cutis vera) confifts of a thick, clofe and whitifh tiffue ; it ap- pears to be produced by the lymphatic and fan- guine vefTels eroding each oth^r, and by a large quantity of nerves. The nerves expand at the furface of the dermis, in very fine papillae, which feem to conflitute the organ of touching. The dermis adheres to the parts which it covers by a loofe cellular tiffue. Above the dermis is found a vifcous, foftifli, and fomewhat thick fubfiance, called the mucous tif- z 2 fue 340 ORGANIC STRUCTURE, fue {refc miicofum, corpus reticularey mucus malpighu anus)^ This fubftancc is of different colours among different nations^ and is the caufe of- the diverfity of colour in the Ikin, The mucous tifTue is covered by a thin mem- brane, dry and tranfparent, which conflitutes the epidermis or cuticle. This membrane forms a covering, in fome meafure inorganic, proper for feeuring the parts immediately beneath it from the impreflion of external bodies. The Ikin exhibits, in its fubftance, fmallglands^, which fecrete an un^luous humour, with which the whole furface of the Ikin is, as it were, varnifh- ed : thefe glands are more abundamt in the par-ts furnifhed with hair. In the lafl place, the Ikin contains the bulbs or radicles of the hairs and briftles, with which thefe different parts are co- vered. This kind of cutaneous vegetation takes place with much more energy in animals, whofe bodies are habitually naked ; and in them this part of tjie cutaneous organ, whether compofed of fcales, feathers, wool, or hair, fupplies them with a thick covering, and becomes to them a very im- portant apparatus of excretion. 345. When the blood, by the digeflion of too large a quantity of drink, Jias received an excefs of aqueous parts, it is freed from this excefs by urinary fecretion. At the fame time the kidneys fecrete different faline fubflances, and particularly -^ matter SYSTEM OF CIRCULATrON. 341 a matter (ziree) the prefence of which characterizes the urine of man. The -kidneys are fituated one on each fide in the region of the loins. Above them, and behind the peritonaeum, is a ghndnlous hody, fhaped like a flattened hemifphere, of a yeilowifh brown co- lour, but larger and redder in infancy. This fuper-renal gla?2d receives a number of vef- fels and nerves ; it is enveloped by fat, and appears to be compofed of lobules covered by cellular tif- , fue : in the infide it exhibits a capfule or cavity of the fame form as the gland, and which at the bot- tom has a fmall fold in the form of a ridge. The fides of this cavity are applied to each other; they have the appearance of mucous membranes, and fecrete a yeilowifh liquid. The ufe of this organ is unknown. The kidneys are of a reddifli brown colour; they have pretty nearly the form of a kidney-bean : they are fmooth in adults; unequal and rough in in- fants. They are placed length wife on the edges of thei vertebral column, and correfpond to the extent of the bodies of the laft two dorfal vertebrse, and the' firft two lumbar; They are covered before by the peritonaeum, and reft behind againfl the fquare of the loins. The right kidney is lituated below the liver, and the left below the fpleen. The con- cave edge correfponds inwardly, and receives the veffels and the nerves, Z3 ■ Ths 342 ©RGANIC STRUCTURE. The kidney is immediately enveloped by a fmooth, compa(5i: membrane, which adheres clofely to it, and which is covered by a thick ftratum of an adipofe tiffue. It receives a very large artery (the renal), which comes from the abdominal aorta. The orifice of the two renal arteries is fuch, that it can admit the eighth part of the blood which iflues from the aorta. The kidney, when cleft lengthwife, exhibits a firfl: exterior ftratum (the cortical) of a dark red colour, about two lines in thicknefs, confifting of a compact tiflue. From this tiflbe arife, in dif- ferent points, fibres or fmall tubes, of a paler red, which converge internally in fmall bundles, and terminate in the form of papilLe : the number of thefe papiilse varies between twelve and eighteen. The exterior compafl tiftue is continued betweeri the bundles of thefe- fibres or tubes. Each of thefe papillge opens into a membranous and grealy capfule in the form of a funnel. Thefe funnels unite into feveral trunks, which end at a common refervoir or fmall bafon. It appears that the urine is fecreted in the com- pa6l tifTue o^ the furface of the kidney, that it flows through the tubulated bodies, oozes from the pa- pillae, and is poured into the particular capfules which proceed to the bafon. Towards the centre of the interior edge of the kidney, the bafon opens into a very large membra- nous du61: [ureter), which defcends behind the pe- ritonaeum SYSTEM OF CIRCULATION. 343 ritonseum along the^ lumbo-trocbantinian mufcle, and proceeds into the leiier pelvis below, behind, and on the fides of the bladder. The ureter is formed of a white, compa^l, and llrong membrane, the interior fides of which have the appearance of mucous membranes. The bladder, which is capable of containing a little more than a quart, has a round form, fome- what flattened at the bottom;, and terminates at the fummit in a blunt point : in the female it is fomewhat flattened from before backwards ; and in infants is more lengthened from the top down- wards. In the male it is fituated in the lefler pel- vis, between the pubis and the redium ; and in the female between the pubis and the matrix : it is covered at the top and behind by the peritonseum, the expanfion of which fupplies it, below and be- hind, with fome ligaments, by which it is fixed to the neighbouring parts. The bladder has, at the top, a ligarrient formed of three cords, which proceed to the navel, raifing up the peritonaeum ; two of thefe cords are produced by the obftru(?led umbilical arteries ; the lafi: by a du6l which, in the foetus, proceeded from the blad- der to the umbilical cord (urachus). This hollow organ ftill adheres anteriorly to the poflerior part of the pubis, by a coinpa6l cellular tJ.flue in the form of a ligament, z 4 The 344 ' OR.GANIC STRUCTURE. The fides of the bladder exhibit outwardly feve- ra.1 layers of mufcular fibres^ which crofs each other, and proceed in all diredlions ; thefe fibres are covered behind by the peritonaeum, and in the reft of their extent by a cellular tifliie. The ureters proceed below, behind, and on the fides of the bladder ; they are difperfed in the fubftance of its fides, and proceed to the interior part of it. The interior furface of the bladder is formed of a thick, villous membrane, furnifhed with a great number of folds. Below and before, this organ exhibits an aperture {neck), the fides of which are very thick ; behind this aperture, the bottom of the bladder has a triangular fpace (trigone) of a clofer tifi:ae, and exceedingly irritable. The in- terior angle of this trigone is continued in the lower part of the aperture of the neck, by a fmall round tubercle: the other two correfpond to the apertures of the ureters. The exterior aperture of the blad/ier is conti- nued by a very long du<5l, the urethra^ which in man runs alono- the vard : in the female it is fhort- er, and terminates at the furamit of the vagina. 346. The urine varies very much according to circumftances, in regard to its quantity, its colour, and its fmell. In general, it is thicker the more flowly it flows, and the longer it has remained in the bladder : that voided immediately after meals, and SYSTEM OF CIRCULAT10>I» SAS snd that, in particular, which refults from an ex- cefs of alcoholifed- beverage, is exceedingly clear^ and contains fcarcely any thing but water. The human urine has been carefully analyfed by the modern chemifts. We (hall here give a (hort view of the principal experiments by which its compolition has been afcertained. This animal liquor always contains uree ; 711'icacid; an animal matter infolublein alcohol; ^vtt, phofpho^ ric acid; phofphates o( lime, oijoda^ oi ammonia, and o( magnejia; muriates o( foda and of ammonia : the Julpbates ofpotajb and of Joda are alfo fometimes found in it. The urine of children always contains benzoic acid. The pretence of the firft ten fubftanccs is afcertained by the following means : The prefence of free fhofphoric acid is afcertained by pouring urine into tindure of turnfole. If the phofphafe of lime be feparated by ammo- nia, this alcali will faturate the free phofphoric acid, and the phofphate of lime will be preci- pitated. By evaporating, to the confidence of iyrup^ urine treated by ammonia and then filtered, and afterwards combining it with alcohol, the uree and the muriate of ammonia will be diflblved. The mu- riate of ammonia may be decompofed by barytes, and it is then expofed to heat to expel the ammo- nia. If it be again treated with alcohol, and if the matter be evaporated to drynefs, nothing will •be diffolved but the uree. Ammonia may be ftill poured 346 ORGANIC STRUCTURE. poarecl over the refiduurr), and after the liquor has heen filtered it may be concentrated by evapora- tion. If muriatic or any other acid be then poured into it, the wic acid will be precipitated in white flakes, which, when thrown on the filter, affume the form of. teal es. Diflblve in water the refiduum of the former ex- periment, and after filtrating the liquor fubje6t it to evaporation. You will then obtain, by cryf- tallization, muriate ^nAphofphate o(foda, combined, in whole or in part, with phofphate of ammonia. Tbefe falts may be obtained alfo by evaporating new urine : in proportion as the evaporation is effected, the liquor becomes turbid by the preci- pitation of X\\Q phofphate of lime, becaufe there is extricated ammonia, which faturates the free phofphoric acid that held this fait in folution. If the liquor be then filtered, and the evaporation be continued, the different falts will cryfiiallize : the ammoniacal fait \n cuh\c2\ cvy^dXs, on account' of the uree with which it is combined, and the ma- rine fait in o<5laedra for tHe fame reafon ; the phofphate of foda, which is combined in wiiole or in part with the phofphate of ammonia, cryftal- lizes in rhombuses. The prefence of the ammoniaco-magnefian phofphate may be afcertained two ways : ift, if the urine be left at reft, {]\\?,falt will very often be obferved in white cryftalline lamina;' ad, if potafn be poured into the urine, phofphate of lime an4 SYSTEM OF CIRCULATION. 34/ and o^magmfia will be precipitated ; by treating this precipitate with acetous acid, acetite of magnefia will be formed j and if this acetite be decompofed h-^ potafh, the magmfta will be feparated. The animal matter^ which confiantly exifls in urine^ is infoluble in alcohol : the nature of it has not yet been properly afcertained. If urine contains alfo ihtfulphates of foda and of putajh, they may be obtained in the fame manner by cryftallization. In the lalt place, if it contains lenzoic acid, as is the cafe with that of children, it raufi: be evaporated almoft to the confiftence of l)Tup ; if it be then introduced into a retort, with a fufficient quantity of weak muriatic acid, and be expofed to diftillation, the benzoic acid will pafs over into the receiver. Urine contains alfo fometimes gelatin , albumen, oxalate of lime and of ammonia; bat all thcie fabftances are found only very rarely. In fome difeafes the urine contains a great deal of gelatin : the prefence of it may be afcertained by the infoluble precipitate which it forms with tannin. What characierizes the urine of man, in an ef- fential manner, is the prefence of uree and of the uric acid. Thefe fubflances are found only in this liquid. Urec produces, in a great meafure, all the phsenomena which urine exhibits; it cryf- tallizes in lamina: like the muriate of barytes -, it combines 34S ORGANIC STRUCTURE. combines with the nitric acid^ and forms an infolu- ble compound. When a folution of uree in water is boiled, there is difengaged a very large quantity of ammonia, which arifes from the decompolitioii of that fubllance. Urine, by the precipitation of the fubftances it holds in folution, is fufceptible of forming calcu- lous concretions. Urinary calculi are found in the kidneys, along the ureters, in the bladder, and even in the canal of the urethra. The introdu61ion of a foreign body into the bladder often determines the formation of a cal- culus, to which it ferves as a nucleus. The different human calculi are formed of the following fubftances: phofphate of lime, ammoniaco- viagnefian phofphate, uric ac'idy urate of ammonia^ oxalate of lime, carhonate of lime, and flex : thefe matters are pure or mixed, and almofS: always dif- pofed in ftrata. Calculi ofphofphate of lime, of ammoniaco-mag- i)elianphofphate,andofcarbonateof lime, are white; thofe of phofphate of lime are never cryftallized, and form a magma with fulphuric acid ; thofe of ammoniaco-magnefian phofphate are alwaj^s cryf- tallized in brilliant laminse, and diflblve com- pletely in fulphuric acid ; thofe of carbonate of lime cryftallize generally in clofe filaments ; they diflblve entirely in the nitric and muriatic acids,with which they efFervefce : the laft Ikind are very rare. Calculi SYSTEM OF CIRCULATIOX. 34g Calculi of urate of amrnonia are of a yellowilli gray colour; ibey diliblve completely in potafh, with a difengagement of ammonia ; thofe formed by pure uric acid are more or lefs yellow, and dif- folve alfo in potalli, but without a difengagement of ammonia. Calculi of oxalate of lime are always more or lefs black and tuberculous; and hence they have been called muraux, on account of. their fuppofcd re- femblance to a mulberry : by calcination they give quick-lime, or carbonate of lime. Calculi which exhibit filex affecl alfo the mul- berry form J they are foluble in acids, give no lime, and are not altered by calcination. Silex has never been found but in two calculi, and even then it was covered, firft by oxalate of lime, and then by uric acid. Calculi formed only of phofphate of lime, or ammoniaco-magnelian phofphate, are rarely found; thefe two falts are almoft always mixed. Calculi of pure uric acid are very often found; and fome of urate of ammonia equally pure ; but the uric acid, for the moft part, is accompanied by urate of ammonia. It is not uncommon to find calculi of pure ox- alate of lime; but this fait in general ferves as a nucleus to the other calculi. When a calculus is compofed of phofphate of lime, ammoniaco-magnelian phofphate, uric acid, urate 350 ORGANIC STRUCTtJJlSi urate of ammonia, and oxalate of linne, thefe mat- ters are almoft always difpofed in the following order : oxalate of lime in the centre, next urate of ammonia, then uric acid, and in the laft place phofphate of lime mixed with, and fometimes co-» vered by, am moniaco-magnefian phofphate, Thefe refults of the analyfis of urine and human calculi are a very ihort extract from the mofl in- genious feries of experiments ever made on ani- mal fubftances. This labour, for which -we arc indebted to Fourcroy, Vauquelin, and Thenar, while it conveys ufeful information to the medical pra6lilioner in regard to the nature of urinary calculi, will furnifh hints for difcovering the pro- per means to prevent the > ravage occafioned by calculous afFe61ions. None however but phyli- cians who are good chemifts will be really deli- rous to take advantage of thefe means, or can pro- perly employ them. 347. The blood, after having diftributed itfelf to all the organs by means of numerous arterial rami- fications, and their indefinite divifions, and after having every where conveyed the materials necef- fary for the different fecretions, and for eifedling the continual changes which take place in every part of the organization, returns by two orders of vefTels, the veins and the lymphatics. The veins bring back the greater part of thebloody and in particular that which has undergone the leafl: change. SYSTEM 0? CIRCULATIOX% 351 change, and which Hill retains its red colour. The means of the communication which exifts between the extremities of the arteries and the firft radicles of the veins cannot be obferved ; they are con- cealed from us on account of their infinite mi- nutenefs,' and have hitherto eluded our microfco- pic refearches. I-t is known, however, that under certain circumftances the blood feems to pafs di- redlly from the arteries into the veins, without undergoing any remarkable changes. The fides of the veins are much thinner than thofe of the arteries. Their fi:ru6lure, examined in the large trunks, is compofed of three tunics, the middle one of which appears to be vafcular. The venous veflfels are alfo more numerous, larger, and more extenfible, than the arterial : it is efi.i- mated that the capacity of the veins is to that of the arteries as nine to four. The interior of the veins exhibits membranous folds in the form of valves; thefe valvulous folds, which oppofe the return of the blood, are for the moft part difpofed in pairs, and are not found in the fmall veins nor in the large trunks : the veins which convey the blood from the vifcera contained in the large cavities, and thofe which are fituated at a confi- derable depth between the mufcles, are alfo un- provided with them. The veins in their progrefs follow a dire6lion contrary to that of the arteries, and are not very flexuous. Their anaftomofes are alfo 052 ORGANIC STRUCTURE, alfo lefs frequent : it is to be obferved, ttiat m uniting they decreafe in fize ; the blood, which is then more confined, mull circulate with greater velocity. The veins which bring back the blood from the inferior limbs and from all the parts of the abdomen unite into a large venous trunk, which afcends to- wards the heart (vena cava afcendens). Thofe which bring back the blood from the head, from the fuperior limbs, and from the thorax, unite alfo into a large trunk, which defcends towards that organ (vena cava defcendens.) Thefe two large veins open into a finus, auricle, with the fmall vein which conveys back the blood from the beart. 348. The lymphatic vefiels are exceedingly thin and tranfparent; their fides feem to be formed of two laminse, which, notwithllanding their thin- nefs, are remarkably "ftrong. Their radiculae arife at the furface of the membranes, and open into all the cellules or cavities by pores or villolities, which feem to poffefs a power of fu6lion. The lymphatics abforb the aqueous fluids difperfed throughout all the tilTues, and the ferous re(i- duums produced by the changes which are con- tinually efFedled in every part of the organization, Thefe vefTels are highly contractile, and exhibit frequent anaftomofes ; in general they accompany the veins ; like them, are difpofed-in two ftrata, one SYStEM OF CIRCULATION. 353 one fuperficial and the other profound, and are furnifhed with valves which prevent the return of their fluid. The lymphatic vefTels are exceedingly nume- rous: by their inextricable interfe6lions they pro- duce plexus, and feem even to form the whole tifllie of the white organs. The lymphatics^ after a certain paflage, meet with fmall glands, which they enter. The ferous fluids and the chyle which they carry thither are there aflimilatedj and, having undergone a fort of digeflion^ iffue from thefe fmall glands by larger and lefs numerous veflTels. The lymphatics con- tinue to advance towards the thorax, traverfing the glands which they meet with ; and the lymph in thefe glandulous organs acquires new chara6iers of animalization. The lymphatic glands are fmall and very nume- rous; they vary in their form and conflftence; they are obferved more evidently towards the bend of the large articulations, in the abdomen, the neck, and around the organs of fecretion. In proportion as the lymphatics unite, and the rami form branches, their abfolute capacity de- creafes, the fluid is more confined, and mufl cir- culate with more rapidity. The refult of the fucceffive union of all the lymphatics is two trunks, which difcharge them- felves into a vein fltuated under the clavicle. Of VOL. I, 2 A thefe 354 -ORGANIC STRUCTURE, tbefe two trunks, the largePf^ the mOil exten-' live, and fometimes the only one apparent, arlies in the abdomenj on the left fide of the vertebral column, towards the middle of the lumbar region, where it exhibits fometimes a fort of dilatation (refervoir of the chyle), and receives the lymphatic velTelsofthe inferior limbs, thofe of the organs contained in the pelvian and abdominal cavities, and thofe of the fides of thefe cavities. This lym- phatic trunk traverfes the diaphragm with the aorta, and penetrates into the thorax (thoracic du<51:) : it afcends on the fides of the bodies of the vertebrae, in the fubftance of the pofterior medi«. aflinum ; and when it reaches the cervical region it bends to the left, and opens into the fub-clavian vein of that fide. This thoracic dud receives, in different points of its extent, the lymphatics which proceed from the right fide of the head, the neck, and the bread, and frcm the thoracic limb of that lide. When the left thoracic du61 exifts alone, it receives the lymphatics of the right fide. The right lymphatic trunk is obferved on the fide of the laft vertebra of the neck. This fmall trunk (the brachio-cephalic) receives the lympha- tics which return from the right fide of the head, the neck, and. the breafi, and thofe of the right thoracic limb : it opens into the fub-clavian vein of that fide. 349. The arterial bicod, in diftributing itfelf to SYSTEM OP eiRCULATlON. 355 tfevery part, and traverfing every point of the orga- nization, undergoes a fort of alteration : it lofes ibme fubftances eflential to its compofition, and becomes charged with feveral foreign matters. It is in this flate that it returns by the veins and the lymphatic veflels. The lymphatics^ in their progrefs, traverfe the numerous lymphatic glands already mentioned. The lymph and the chyle experience in thefe glands a peculiar affimilation, and undergo im- portant changes, before they unite with the venous blood. The venous blood, in its partial paflage through the liver, begins to be diveftcd of its foreign fub- ftances, and is completely freed from them in the lungs : it there acquires thofe materials which were taken from it, and refumes the qualities of arterial blood, to ferve for a new circulation. We fhall here defcribe the different organs through which the venous blood pafles before it penetrates to the aorta. 350. The veins which bring back the blood from the principal .vifcera of the abdomen unite into a large trunk which proceeds into the liver. The liver is a gland of conliderable fize, and of a dark red colour, lituated in the right hypochon- dre, the epigaftric region, and a part of the left hypochondre ; below the diaphragm, to which it is contiguous fuperiorly, and above the ftomach 2, A a « aad 356 ORGANIC STRUCTURE* and the inteftines : it is more voluminous towards the right tide j is fmooth and convex at the top and before, concave and unequal at the bottom and behind. At the lower part and before it exhibits a fharp edge, in the middle of which is a flight depreffion that divides it into two lobes, of which that on the right is larger. This gland has alfo a fmall lobe at the bottom and behind (lobulus Spigelii). The liver is covered throughout its whole ex- tent by the peritonaeum : this membrane is refledled from diiFcrent points of its furface, and its double fold forms feveral ligaments, by which it is fixed to the neighbouring parts. One, which is very large^ arifes from above the middle of the convexity of the liver, ^nd proceeds to the navel and the centre of the diaphragm (fufpenfor ligament). It receives before,, the umbilical vein, v/hich is ob{lru61ed after birtb, and becomes ligamentous. The peritonaeum pro- duces on the iides two expaniions, which form two lateral ligaments. It adheres alfo behind the dia- phragm by the cellular tifTue (coronary ligament.) The hepatic organ receives its blood from a fmall artery (right branch of the cogliac) and a very large vein (fub-hepatic or vena portse). The blood of the fmall artery feems deflined for the particu- lar nourifhment of the liver i the ligature of this veffel in animals does not interrupt the fecretion of the bile. SYSTEM OP CIRCULATION, 357 Tbefub-hepatic vein or vena porta is formed by two trunks (the fplenic and merenteric)^ which bring back the blood from the ftomach, the epiploon^ Jhe inteftines, the mefentery, the pan- creas, and the fpleen. This large vein proceeds into the liver towards the middle of its depreffion, tranfverje fc'ijfure, which is obferved on its concave face, where it forms a large finus, and then dif- tributes itfelf to the right and the left ; it then divides and fubdividesitfelf indefinitely in the fub- ftance of that vifcus. The divilions of this vein accompany thofe of the artery, the nerves of the liver, and the duds which bring back the bile. The fub-hepatic vein is enveloped from its origin by a prolongation of the peritonseum ; this prolongation accompanies the three orders of vef- lels, as well as the nerves, and forms of th<2m a fort of common capfule (^capfida GUjfonii), from which proceed very fine membranous partitions interpofed between thefe parts. The fub-hepatic vein, with its divifions, dilcharges the office of an artery to the liver, fince it furnifhes it with almofl all the blood it receives : it is obferved that the fides of* this vein are thick, and that it has no valves in the infide. The nerves of the liver (branches of the pneu- mogaflric and trifplanchnic) unite and form a plexus (the hepatic), which accompanies the bili^ a A 3 ^ry 358 ORGANIC STRUCTUKE. ary da^ls, and the veffels which carry the blood thither. The blood is diftributed throughout the whole fubftance of the hepatic organ, where it frees it- felf from the albuminous and oily matters, which it contains in excefs. The combination of thefe fubftances with a fmall quantity of foda forms a faponaceous liquor, which excretes from every point of the liver, through particular du(^s, and is conveyed into the duodenum, where it becomes one of the mofl powerful agents of digefdon. The venous blood, after being freed from the materials of the bile, returns by veins and lympha- tic veflels. The veins (fub-hepatic) bring back the blood from every part of the liver, purfuing a direcStion perpendicular to the veflels which carry it thi- ther : their fides are thin, and without valves. From the union of their numerous rami arife three branches, which proceed from the three lobes of that gland : thefe three veins form a trunk, which proceeds into the vena cava, The lymphatic veflels of the liver are very nu-» merous in the fubflance of that vifcus, and parti-^ cularly at its furface. Thofe of the gaftric flde unite into fome trunks, which proceed into glands fituated near the hepatic dudls : thofe of the diaphragmatic iide unite along the fufpenfor ligament SYSTEM OP CIRCULATION. 35g ligament of the liver, traverfe the diaphragm, and proceed into the large chyliferous du6l (the thoracic). 351. The venous blood, after undergoing fome changes on its partial pafTage through the liver, and after receiving all the fluids brought by the lymphatic and chyliferous veflels, unites into two large veins, which proceed into a flnus called ihe^mis of the vena cav£ (pulmonary or right auricle.) This linus is lituated at the upper part of the heart, on the right, and fomewhat behind ; it is fhaped like the fegment of a fpbere, and exhibits a fmall appendix. Its interior furface is furrowed by carneous pil- lars, which adhere to its fides. At the top and before is the oripics of the fu- perior vena cava ; that of the inferior vena cava is lituated below and behind ; it is larger than the former, and furnifhed with a valve fhaped like a crefcent, which prevents the blood from flowing back : in general this valve is incomplete, and feems to be in part torn or pierced : it is more ap- parent in the foetus, and dircfls ail the blood of the inferior vena cava towards the foramen ovale. Before this valve is the aperture of the vein of the heart (the coronary), v;hich is alfo furnifhed with a valvular fold. The fin us of the venas cavae correfponds wilhin to that of the pulmonary veins, from u-hich it is 2 A 4 f^paratcd 360 ORGANIC STRUCTUREo feparated by a carneous partition: on this parti- tion is obferved a fmall oval fofTa, which in the foetus is a hole of the fame form ; the greateft di- ameter of this hole is four or five lines, and it is furnifhed with a femilunar valv6, which correfponds to the linus of the pulmonary veins. After birth^ this valve is prefled clofeiy to the aperture by the blood which arrives in greater abundance in that linus from the lungs ; it adheres to it, and the foramen ovale is foon completely obftru61ed : it has however been found open at an advanced age. The finus of the vense cavae communicates with ihe pulmonary ventricle by a round aperture. Its tendinous circumference is furnifhed with a valve^ produced by a membrane cut into three tri- angular portions of unequal fize, which termi- nate in a point (valvulas tricufpides, triglochines). To thefe membranous portions are attached, by their free edge, tendinous or carneous filaments, which proceed from the fides of the ventricle. The a6lion of this valve is to prevent the blood from flowing back into the auricle during the contrac- tion of the ventricle. The pulmonary ventricle has the form of a folid triangle I its interior face exhibits a great number of carneous pillars, adherent or free, and of dif- ferent fizes. The greater part of thefe pillars are dire6]ed from the bafe to the apex, and others crofs the latter in an oblique diredion. The SYSTEM OP CIRCULATION. 3<)I The pulmonary orifice of the ventricle Is fur- Bitlied in its circumference with three femilu- nar valves (figmoid). Thefe valves, when dilated, have been con^pared to three pigeon-bafkets the convex faces of which touch each other: they ex- a6lly clofe the aperture, and prevent the reflux of the blood from the pulmonary artery into the ven- tricle. The pulmonary artery, after a fhort pailagc, di- vides itfelf into two branches, which proceed to the two lungs. In the foetus, thele two branches are very fmall ; bat the pulmonary artery has a dire6l communication with the aorta by a canal. The blood which arrives from the ven- tricle, being able to penetrate only in fmall quantity into the lungs comprefled and not yet dilated by refpiration, proceeds dire61Iy into the aorta by this arterial canal\ but, after birth, the pulmonary arteries expand ; the whole blood palles into them ; and the arterial canal, which flovvly be-.^ comes obftrudted, firft towards the aorta, is changed into a ligament during the firft years of life. 352. The LUNGS are placed in the two cavi- ties of the breaft, the form of which they nearly retain ; they are convex and fmooth on the out- fide, and excavated towards the heart, which they embrace. They are broad ; are flat and have fharp edges below, and are narrower and rounded at the top and behind. The 36*2 ORGANIC STRUCTURE. The right lung is compofed of three lobes, and the left of two. They are of a dark red colour ia the fcstus; of a purple red in infancy ; become of an afh white colour in adults, and are inter- fperfed with blaeifh fpots, which with age increafe in number and inteniity. In the foetus, the lungs are clofe, compa6l, and heavier than water; but, when they have been diftended by air during refpiration, they retain a great part of that air after death : they continue light, fpongy, and float in water. The cellules of the lungs (the bronchial) are ex- ceedingly fmall : the lad membranous ramifica- tions of the bronchia? terminate in theinfide of them. The bronchial ramifications of the two lungs arife from two trunks, produced by the tracheal artery at its termination : the fuperior aperture of the tracheal artery is the larynx. The larynx is a cartilaginous dilatation fltuated at the bottom of the pharynx, and before the cefo- phagus. The principal cartilage which enters into, the formation of it islituated below theos hyoides; it has been compared to a buckler, on which ac- count it is called the thyroid. It is broad and round before, and in the middle exhibits a vertical line : it is incomplete behind, and its lateral faces ter- minate at the top and bottom in angles or horns^ which are prolonged backwards: the fuperior ones are thinner and longer. fj- Below SYSTEM OP CIRCULATIOJT. 303 Below the thyroid cartilage is another annular cartilage (the cricoid), narrow before and broad behind, which is articulated with the inferior horns of the thyroid. Above the poilerior edge of the cricoid cartilage, in the fpace left by the thyroid behind, there are two fmall triangular cartilages, twifted fomewhat round, fo as to reprefent the portion of a funnel (arytsenoid) : the broad bafes of them reft upon and are articulated with the cricoid ; their fummit, which is thin and narrow, is dire^ed inwards. In the laft place, the larynx above its fuperlor aperture (glottis) exhibits a cartilage (epiglottis), which is thin, broad and round ; its bafe is united before to the os hyoides, and on the fides to the thyroid and arytaenoid cartilages, by lax liga- ments ; its fummit falls down behind on the aper- ture of the glottis, during the paiTage of the ali- ments. At the entrance of that aperture it forms a fort of pipe, which makes the air vibrate on its paflage, and produces found. All thefe cartilages are united by ligaments, which concur to form a very flexible cavity : thefe ligaments produce, by their folds in the interior of the larynx, two deep excavations {lateral ven- tricles). The cartilaginous pieces which form the larynx are moved by means of feveral pairs of fmall muf- fles. One extends from the fuperior edge of the cricoid 3^4 ORGANIC STRUCTURE. cricoid cartilage to the inferior edge of the thyroid, on the lateral parts of thefe cartilages : erico-thy- ro'ulian. Another is inferted in the pofterior part of the cricoid cartilage, and is fixed at the bafe of the arytenoid cartilages : crko'arytamid'mn, ' From above the lides of the cricoid cartilasfe. and behind the thyroid, arifes a mufcle which is inferted before the arytenoid cartilages : lateral crlco-arytisnoidian. From the interior face of the thyroid cartilage arifes a fmall mufcle, which proceeds backwards and upwards, towards the arytsenoid cartilage: thyro-arytaenoidian. In the laft place, a fmall lingle mufcle covers the two arytaenoidal cartilages behind : arytatioidian. The combined a6lion of thefe mufcles can lengthen or fhorten, dilate or contra6l the capacity of the larynx, and produce a variation iri the aper^ ture of the glottis. Befidcs thefe fmall mufcles, which tend to move the different pieces of the larynx, there are two others^ which can lower or raife the whole of it, Of thefe mufcles, one is inferted in the interior face of the firft piece of thx fternum, below the fiylo-hyoidian, and proceeds to the oblique line obferved on the thyroid cartilage. This mufcle tends to lower the larynx : Jlerno-thyroidian^ Another, of a fmaller lize, is attached on the - 6 . thyroid SYSTEM bf CIRdUtATION. $Qq thyroid cartilage, above the preceding, and pro- ceeds to the OS hyoides. This mufcle can raife the larynx, when the os hyoides is fixed : hyo-thy" . roidian» The larynx which is lined with a liiucdus mem- brane is furrounded by feveral glands : the prin- cipal of thefe glands, which is broad and fiar, is fituated before the thyroid cartilage, by the name of which it is diftinguifhed j before the cricoid and the upper part of the trachea. The particular ufe of the produ6l of its fecretion is not known. A feries of fmall glands is obferved alfo before the arytssnoid cartilages, and a fmall glandulous body on the epiglottis. The larynx is continued inferiorly with the tra- chea, which defcends before the oefophaguSj to the height of the fecond dorfal vertebra. This conduit is compofed of cartilaginous rings united by looie intermediate ligaments t their number varies between fixteen and twenty. They are incomplete behind, and the vacuity is filled up by a longitudinal ligament* The firudture of the trachea is fuch, that it3 length and diameter can be eafily varied : the cavity of it is lined, like the larynx, by a mucous membrane furnifhed with fmall glands. Inferiorly, the trachea is divided into two trunks, which are the commencement of the bronchia s the 366 ORGAHIC STRUCTURE, the (horter one proceeds to the right long, and the oiher to the left. < In penetraiing into the lungs, the bronchiae are divided into rami, and labdividcd into ramifica- tions, exceedingly delicate in every point- of that organ. . The texture of the bronchis is the fame as that of the trachea ; but in proportion as the diameter of thefe conduits decreafes, the rings of them be-? come irregular, and at length difappear. The bFoncb.ial conduits are then membranous, and ter- " niinate in the numerous cells of the lungs. Thefe cells have not a dired communication with each other, but only by the cpnduits of the bronchise. , , Ip the interior part of the lungs, along the bronchial ramifications, are obferved a g'reat num- ber of fmall glands, fituated chiefly at the angles of their divifions (bronchial glands.) 353. The two divifions of the puhnonary artery, at their entrance into the lungs, fubdivide thera- felves into three parts on the right, and into two on the left. Thefe arteries aceompany the bron- chia?., and foKovv them by fubdividing in the fame manner. Their lafl ra.mufculi penetrate into the bronchial cells, expand over the fides of them, and there anaftomofe dire611y with the capillary extra- pi", ties of the veins. ■ • The SYSTEM OF CIRCULATION-. 36f The pulmonary veins follow alfo the divlfions of the bronchise, and bring back the blood from every part of the lungs : the ramufculi unite to for;:i rami ; the rami to form branches ; and the branches produce large veins for each lung, which pour the blood into the fin us : Jinus of tbe pulmonary veins (left auricle or aorta). 354. This linns (auricle) is ftoaller than the preceding ; it is fituated at the pofterior and left part of the heart, Tn the ftate of dilatation it af- fe(5ls a cubic form, with an indented prolongation at the top and on the outfide : auricular appendix* The interior face of it is fmooth : it exhibits, be- hind, the orifice of the four pulmonary veins; in the infide, it corrcfponds to the iinus of the vens cavae, and in the foetus exhibits the foramen ovale : at the top and on the outfide is found the cavity of the appendix. Before, the lin'us communicates with the aortic ventricle by an aperture, the cir- cumference of which is tendinous. This aperture is furnifhed behind with a membranous valve, cut into two fharp-pointed portions {valvules mi- trales"), to which are attached tendinous and flefhy fibres inferted in the fides of the ventricle. This valve prevents the blood from flowing back whea the ventricle contrads. The left or aortic ventricle is fituated before the finus of that fide ; and on the left fide of the pul- monary ventricle it is fmaller, and of a form ana- logous 36s ORGANIC ST^UCtU'RE^ logous to that of the latter ; but its fides are much thicker : its interior face is furnifhed with earneous pillars, like the right ventricle. At the top and on the right is found the orifice of the aortic artery, which is furnifhed with a triple valve (figmoid)j fimilar to that which is found at the entrance of the pulmonary artery. 2SS' ^^^ heart and the lungs are contained in the cavity of the thorax. The interior fides of this cavity are lined with a membrane, which is refleft- ed on each lung, and forms a particular covering to them, called the j)Ieura. To have a proper idea of the difpofitioh of this membrane, it will be necefi^ary to follow it in its different folds. The pleura, after covering the fides of the thorax and the diaphragm, is reflected, before, along the middle of the llernum, with that of the oppofite fide, and forms the two laminae of the anterior mediaftinum, between which the heart is lodged. When it arrives behind that organ, and before the pulmonary veflels, it is reflected on it felf, and covers the interior face of the lungs, and then their exterior face. When it reaches the bafe of that organ, it is united to that of theoppofitefide,and again feparates to produce the pofterior mediaflinum. The pleura then continues on the lateral parts of the vertebral column with that which lines the fides of the thorax. Ill SYSTEM OF CIRCULATION. SGQ In the anterior mediaftinum are found the heart and the thymus gJand; in the pofterior, the aorta, the cefophagus, the thoracic canal, and the prse- lumbo-rhoracic vein. 356. The thymus, of which we have not yet Ipoken (in the calf it is diftinguiflied by the name o^fweetbread), is a voluminous gland, which in the foetus is foft and reddifh ; in children it decreafes and becomes yellow ; with age it continues to lofe fome of its bulk, aiTumes more confiftence, and becomes of a dark yellow colour ; in old age it difappears almoft entirely. In children it fecretes a milky humour, the excretory du6ts of which are not yet known. We are ftill ignorant with re- fpe6l to the ufe of this organ. 357. The pleura feems to be formed of only one membrane ; it adheres by a cellular tiffue to the fides of the thorax, and to the furface of the lungs, which it covers. Its interior furface is applied to itfelf without adhefion, by the contadl of the lungs with the fides of the thorax. This furface is fmooth, and fecretes abundance of ferous matter, with which it is continually lubricated. 358. The heart is enveloped alfo by a peculiar membrane, called the pericardium, thick, and of a compa6l texture. It is formed of two folds, the exterior of which, after enveloping thfe heart, is continued on the large veflels : it adheres at the. VOL. I. as bottom 370 ORGANIC STRUCTURE. bottom to the diaphragtrij and in the reft of its ex- tent to the mediaftinum. The interior fold, when it reaches the large vefTels, is refle(5led on the heart, and is thus placed in conta6l with itfelf : its fmooth furface fecretes abundance of ferous matter, which continually moillens the heart and pericar- dium. 359. The blood is a red vifcous fluid of a fa- line favour. On the firft view it appears to-be compofed of a limpid fluid, having floating in it red moleculas, refpe61ing the nature of which a great deal has been written. The red globules of the blood have been feen under a lenticular form, and pierced with a hole in the middle, &c. It has been judged that they are fcarcely the eight hun- dredth part of a line in diameter. The temperature of the blood is nearly forty degreesof the centigrade thermometer (104° Fahr.). That contained in the arteries is of a bright red colour, and appears to be warmer than that of the veins. The blood when newly drawn from a vein ex- hales an aqueous, odorous, and putrefcible vapour; that of the veins feems to difengage azote, and that of the arteries oxygen. Chemical analyfis of the blood (hows that it con fills of three principal fubftances, Jihrous mat- ter ^ albumerif and colouring matter. There are found SrSTEM OP CIRCULATION. 371 found in it alfo different falts which are pure foda, phofphate of lime, phofphates of foda and of am- monia, and muriates of foda and of ammonia. To analyfe the blood, it muft firft be left at refl; it then feparates into two partSj one folid, called the crajjame?itum) or cake ; and the other liquid^ called thejenim or ly?nph. The craffamentum is more or lefs red ; it con- tains fibrous matter, colouring matter, and albumen . If a pound of craiTamentum, wrapped up in apiece of linen, be fufpended under the cock of a ciftern, fo as to be fubje6ted to a continual ftream of water, it is obferved that the water gradually carries off the colouring matter, and a part of the albumen ; land at the end of twelve or fifteen hours nothing remains but the white fibrous matter. To feparate the colouring matter from the albu- men, which is found mixed with the former in the water, the water mufi; be boiled ; the albumen then coagulates, and carries with it the colouring mat- ter. This matter is then obtained by calcination^ mixed only with a fmall quantity of the phofphate of lime. The colouring matter, after this procefs, gives only phofphate of iron with excefs of oxide. The waters employed for wafhing lofe their colour by boiling; they retain only a yellowifh white tinge, and hold nothing in folution but a very fmall quantity of the foluble falts which^ as already faid, are found in blood. 2 B 2 The 372 ORGANIC STRyCTURE. The ferum, which is of a yellowifh white coioufy h vifcous and perfedlly limpid ; it turns fyrup of violets green, in confequence of the cauflic foda which it contains. When nitrate of filver is poured into ferum, it produces an abundant flaky precipi- tate, which difTolves only in part in pure nitric acid : this proves that the precipitate is partly owing to muriate of filver which has been formed. Serum, when mixed with lime water or a folublc calcareous fait, fuffers to be precipitated phofphate of lime. In the laft place, ferum, when heated, forms itfelf into a mafs by the concretion of the albumen. Serum then is compofed of water holding in fb- lution : albumen, foda, phofphates of foda and of ammonia, muriates of foda and of ammonia. This liquor is perfe611y analogous to the white of an egg. Though the lymph has not yet been well analyfed, it is however known that it has a great analogy, to the blood, and none to milk : it feparates fponta- neoufly into a ferous part and a fmall curd : the latter contains fibrous matter. 360. The blood, in returning froiu every part of the lungs, is collected by four veins which pour it into a finus \finus of the pulmonary veins (left or aortic auricle). From this finus it pafies into the left ventricle^ aiid is propelled thence into the aorta. Thff SYSTEM OP CIRCULATION. 373 The aorta in its pafTage furnifhes a very great number of arterial branches, the fubdivilions of which are diftributed to every part. On its ifTuing from the heart, it gives out, on (he right and left, the fmall cardiac artery (coro- nary)^ which is reflected on that organ, and is dif- tributed to the ventricle and finus belonging to its fide. Thefe two arteries penetrate the whole fubflance of the heart, and anaftomofe in feveral places. At the fummit of its curve, the aorta furniOies two large branches on each lide : it furnifhes them dire6lly from the left fide ; but from the right tide gives out in general only one large branch, which is divided. Of thefe two branches, one is diftributed to the liead, the cephalic ^ and the other to the thoracic member : the hrachial. 361. The cephalic {primitive carotid artery) afcends obliquely without, on the tides of the tra- cheal artery, and at the height of the larynx is di- vided into two branches. Of thefe two branches, one is diftributed to the neck, the face, and the exterior part of the cra- nium, the maxillo-facial 'y the other, traverfing the temporal bone, proceeds to the brain : anterior cerebral. 362. The maxillo-facial {exterior carotid ^rtery) afcends on the fides of the neck as far as the neck of the condyle of the maxillary 2 B 3 bone-. 374 ORGANIC STRUCTURE. bone ; and in its paflage farniflies eight principal branches. The firfb arifes near the origin of the maxillo- facial, and defcends, within and before, towards the fuinmit of the thyroid gland, on which it ex- pands : a ramus penetrates the larynx, the laryn- pan, between the os hyoides and the cricoid carti- lage, another proceeds into the interval which fe- parates the cricoid and thyroid cartilages. This artery, in its paflage, diilributes itfelf to the neigh- bouring parts, and exhibits frequent anaftomofes: the Juper'ior thyroidian *. The fecond ifllies above the preceding ; it pro- ceeds inwards, towards the os hyoides, paiTes be- tween the hyo-gloffian and the genio-gloflian mufcles, diftributing itfelf to the mufcles attached to that bone ; where it divides : one part paffes between the genio-gloffian and the mylo-hyoi- dian mufcles, and proceeds to the root of the tongue, where it advances in a ferpentine direc- tion, below and on the lides of that organ, as far as the point of it. The other is directed towards the back of the tongue, and expands at the root of it : the Tmgual. This artery furniflies the Jub-Tin- gual, i\\Q /up er -Tmgual, and a hyo'idian ramus. The third is obferved below the lingual. It is final!, afcends on the lides, and a little behind the * We fliall not mention the numerous varieties obferved in the origin and diftribution of the arteries, we Ihall exhibit only the moft conftant ftate, pharynx^ SYSTEM OF CIRCULATION, 3^5 pharynx, and proceeds to the pharynx and the gut- tural conduit of the ear, diflributing itfelf to the neighbouring parts: a ramus penetrates into the craniunrij through the pofterior foramen lacerum, and conltitutes the poflerior meningian : inferior pharyngian. The fourth arifes above the lingual. It afcends along the fuperior part of the pharynx, and pafles over the maxillary gland, near the angle of the jaw. In its palTage, it furnillies the mufculo-pa- latine, the tonfillary, and the fub-maxillary. It proceeds before, along the interior face of that bone, is then refle6led outwards, afcends on its ex- terior face, near the anterior edge of the zygomato- maxillian, and proceeds to the commifTura of the lips, after giving out the inferior labial. In that place this artery farnithes two rami, which expand in the thicknefs of the lips ; it then afcends along the cheek, on the fides of the nofe, at the famnjit of which it terminates, anaftomoling with the fub- orbitar : the labial. The labial, in this long paflage, furnifhes the mujculo'^alatine, ihQ fub-maxillary ^ the iotifillary, the inferior labial jihe, two labial coronary, and fome rami. The fifth iffucs from the maxillo-facial, almoft oppofitc to the lingual. It afcends obliquely be- hind, under the fterno-, cervicO', and trachelo-maf- toidian mufcles -, pafles between the maftoid apO' phylis and the tranfverfe apophyfis of the atlas, 2 B 4 and 376 ORGANIC STRUCTURE. and afcends under the integuments as far as the occiput, where it expands in numerous rami : the occipital. The fixtb, which is very fmall, arifes in the fub-^ fiance of the parotid gland. It proceeds back- wards, above that gland, as far as the maftoid apo- phylis, where it furnifhes a ramus, which pene- trates into the ear by the Itylor-mafloidian foramen (aquedu6l of Fallopius), and diftributes itfelf in the inlide of that organ. This artery then divides on the fides of the maftoid apophyfis : a branch proceeds before the ear, and another on the occi- put : pojierior auricular. This artery gives the Jiylo' majio'idiariy the tympa?iic, and fome mufcular and cutaneous rami. The maxillo-facial, at its termination behind the neck of the condyle of the jaw, divides into two branches, which are the laft furnifhed by this artery. The feventh continues in the dire61ion of the maxillo-facial. At its origin it gives out a branch, which proceeds, before, on the parotid gland, the zygomato-maxillian, the falival du6l of Steno, and as far as the palpebralian mufcle. The body of this artery then continues under the parotid gland, pafles betvreen the meatus auditorius, where it throws out a ramus to the anterior part of the ear and the condyle of the jaw : it then af-? cends behind the zygomatic arch^, above the tem- poro-i SYSTEM OF CIRCULATION. ^Jf poro-maxilHan mufcle, in the fabflance of vv^hich it fends out a branch, and terminates in two rami, one of which expands on the forehead, and the other on the fides of the head : the temporal. This artery furnifhes the anterior auricular, the Jub-%ygomatic, and iho. fuperjicial temporals. In the laft place, the eighth rifes towards the middle of the branch of the jaw-bone. It afcends along that bone, and the fmall pterygo-maxillian, and proceeds to the fummit of the zygomatic fofia, towards the fpheno-maxillary fifiTure : the maxiUo- huccal. In its pafTage, this artery furnifhes feveral branches : a. The firft proceeds within, under the fmall pterygo-maxillian mufcle, and penetrates into the cranium through the fpheno-fpinous foramen, where it gives out fome ramufculi, and divides into two rami, which expand on the meninx, and are lodged in the furrows obferved on the fides of the interior part of the cranium : middle meningiaii. h. Another defcends anteriorly between the great pterygo-maxillian and the branch of the jaw ; and proceeds in the dentary canal : the maxillo-den- tary. Before it enters this canal, this artery furnifhes a ramus to the mylo-hyoidian and the membrane of the mouth. It then pailes through the dentary canal, giving out ramufculi which enter the alveoli of 378 . ORGANIC STRUCTURE. of the dentes molares. When it reaches the hole of the chin, it furnifhes another fmall twig, which penetrates into ihe alveoli of the canine and inci- for teeth: it then itTues from that hole, and diltri- butes itfelf to the mufcles of the lower lip, anafto- mofing with the labial. c. A branch pailes under the fmall pterygo- maxillian mufcle, to which it diftributes itfelf, as well as to the great pterygo-maxillian : the pery- zotdian. d. Two branches are diftributed chiefly in the bottom of the temporal foflae : the profound tem- porals. One arifes below the zygomatic arch, palles before the fmall pterygo-maxillian mufcle, and gives out rami, which expand on the temporo- inaxillian. The other proceeds in the furrows of the temporal foffa, below the temporo-maxillian. It generally gives out a ramus which pafles before the condyle of the jaw, and is diftributed to the zygomato-maxillian • the majfeter'ine. Thefe ar- teries anaftomofe with the fuperficial temporals. e. The maxillo-buccal, before it penetrates to the bottom of the zygomatic foffa, furniflies alto a branch which defcends anteriorly on the maxillary bone: y«/>ifr-m^^i//,2;_)'. It furnifhos ramufculi which penetrate into the maxillary iinufes, and to the roots of the upper dentes mo- lares: it then lofes itfelf in the bucco-labian muf- cle. ifM SYSTEM OF CIRCULATION. 3*g cle, the membrane of the mouth, the neighbouring integuments, and anadomofes with the labial and the fiib-orbitar, f. The maxillo-buccal, when it reaches the bot- tom of the zygomatic fofla^ gives out a branch, which penetrates into the fab-orbitar canal: tljs fub-orbitar. Before it ifTues from this canal, it fur- nifhes a ramus, which pafies through the dentary canal, and diftributes itfelf to the canine tooth and the incifbrs. It then ifTues through the inferior orbitar hole, and diftributes itfelf to the mufcles of the nofe and the lips, prcfenting frequent anafto- mofes. In its paflage it fends out ramufculi, which penetrate into the orbit and the maxillary (inus. g. It then fends out two rami, one of which pafles through the pterygoidian canal, and the other through the pterygo-palatine foramen. Both thefe rami are diftributed to the fummit of the pharynx and the guttural conduit of the ear : fuperior pharyngian. b. In the laft place, the maxillo-buccal furnifhes two rami, one of which defcends along the pofte- rior palatine canal, and diflributes itfelf to the palate : the other pafles through the fpheno-pala- tine foramen, penetrates into the nafal foilse, and divides into feveral ramufculi, which are diftri- buted on the fides of that cavity : ■palatine and ^terygo-falathie. 363. The anterior, cerebral, mUrlor caro- tid, 380 ORGANIC STRUCTURE. tid^ Is larger in infancy than the maxillo-facial : in adult age it becomes equal to it. This artery afcends on the fides of the larynx and of the vertebral canal ; it proceeds as far as the bafe of the cranium, making feveral infledlions, without giving any rami, and penetrates i^ito the cranium through the winding canal near the pe- trous portion of the os temporale. When it reaches that cavity, it pafles on the fide of the body of the OS fphenoides ; is then reflected upwards, turns round in the notch of the anterior cliqoid apophyfis, and traverfes the meninx. The anterior cerebral, immediately after traver- iing "this membrane, gives out a branch, which enters the optic hole, and is diflributed to the orbit: the orhitar. The ophthalmic artery penetrates into the orbit, below and on the exterior fide of the ocular nerve, around which it turns ; it pafi!es below the fuperior mufcle of the eye, and proceeds towards the in- terior fide of that cavity. In this fhort pafiage it furnifhes feveral branches, namely : a. The lacrymal, which advances along the exterior fide of the orbit, is diftributed to the ex- terior mufcle of the eye, the orbito-palpebralian, the lacrymal gland, and the upper eye- lid. A ramus even traverfes the os jogale, and expands on the cheek. 6 ■ h. The SYSTEM OF CIRCULA.T10K". 3S1 h. The/uh-irian branches, which pierce the fcle-^ rotica in different places : they proceed between that membrane and the choroid, as far as the ciliar ring fligamentum clUare), where they are fub-divided into a great number of capillary vef- fels, which form anaflomotic circles, exceedingly delicate, around the large and fmall circumference of the iris. c. The two ethmoidal branches, which advance inwards, pafs over the lobe of the eye, penetrate into the two interior orbitar holes, enter the cra- nium, and are refied^ed in the ethmoidal cells, through the crebriform plate of the os ethmoides. d. The central of the retina, which penetrates into the optic nerve before it enters the orbit. This artery advances into the centre of that nerve, as far as the retina, on which it expands. e. The nmfcular, which gives out two rami : one of thefe, which is larger and conftant, is diftri- buted to the inferior and lateral mufcles of the eye ; the other proceeds into the fuperior, the int^-> rior, and the great oblique mufcles of the eye. f. The two falpeh-alian: thefe are dire(5ted to- wards the interior angle of the orbit, from which they are diftributed to both eyelids, furnifhing ra- raufculi to the neighbouring parts. g. In the laft place, the orbitar furniflies alfo fome rami, which are diliributed to the eye- brows, the forehead (the frontal), and to the nofe (the 382 ORGANIC STRUCTO'RE. (the nafal), and terminates by anafl-omofing witli the labial. The anterior cerebral artery, after having pierced the dura mater, and furnifhed theorbitar, proceeds backwards under the brain, and then gives out a connnunicating branch, which anaflomofes with the pofterior cerebral. This fmall and pretty long branch furnifhes, in its paflage, ramufculi to the furrounding parts, and then fends out a ramus which is diflributed to the choroid plexus. The anterior cerebral artery is then divided into two branches. The fmaller proceeds before : the anterior lobar-, the other proceeds on the lide, be- tween the anterior and middle lobes of the brain : nuddle lobar. The anterior lobar is refledled forwards, below the mefolobe, and by a fhort tranfverfe branch conimunicates near its origin with its fellow. This artery, when it reaches the anterior part of the mefolobe, afcends before the brain, and is refle6ted on its farface. In this whole pailage it furnifhes numerous ramifications, which expand on the meningine, between all the anfradiuofities of the brain, from which they penetrate into the interior parts of that organ. The middle lobar takes a lateral dire(5lion below the brain. It pafles between its anterior and mid- dle lobes,. and divides into feverai branches, which continue to advance between thefe parts, furnifh- ing SYSTEM OF CIRCULATION. 3S3 ing numerous rami to the choroid plexus, the meningine, and the neighbouring parts of the brain. Thefe branches are then fubdivided, and expand over the meningine, in the numerous folds which it forms on the lateral parts of the encepha- tic organ, and lofe themfelves in the interior of that organ. 364. The brachial artery extends from the aorta to the bending of the arm. To facilitate the defcription of this artery, its ex- tent is divided into three parts : ift, that which is under the clavicle (fub-clavian) ; 2d, that which is under the arm-pit (axillary) ; 3d, that which correfponds to the arm (humeral). ^6^. The sue-clavian furniflies, in general, lix branches : The ift proceeds to the brain, the ^ojierior cere- hral (vertebral) ; it rifes along the neck, accom- panying the inferior thyroidian, then enters the canal produced by the holes of the tranfverfe apo- phyfes of the vertebra of the neck, and proceeds as far as the great occipital foramen, making feveral inflexions, efpecially at the top. In this paflage^ it fends out feveral rami to the vertebral canal and to the neighbouring mufcles. When it enters the cranium, it proceeds ante- riorly on the fides of the rachidian prolongation, and gives out a great number of fmall rami, one of which is diftributed to the pofterior part of the meninx: 384 ORGANIC STllUCTtJlt,E* meninx : the occiplto-nienmg'mn : two others de- fcend, one before and the other behind that pro* longation. The anterior extends as far as its extremity ; the pofterior terminates towards the lumbar region : anterior ami fojierior median of the rachis (anterior and poiterior fpinals). The pofterior cerebral artery then fends out a large ramus, which is diftributed on the inferior face of the cerebellum : inferior great cerehelhus. The two pofterior cerebral arteries then unite and form a trunk, which advances over the whole length of the mefencephalon : the me/encephalic (bafiliar) ; this trunk furnifhestwo fmall rami, one behind, inferior fmall cerehellous ; and the other before, near its termination : fmall fuperior cere- hellous, Thev diftribute themfelves to the bafe of the mefencephalon and of the cerebellum. The mefencephalic trunk then divides into two branches, the pojlerior lobar ^ which proceed on the iides. Each firft furnifhes a communicating rartlus, which proceeds to the anterior cerebral artery and fome other fmall rami. This artery then divides, and expands over the pofterior lobes of the brain, diftributing itfelf in its numerous anfradluolities. It is to be obfervcd, thac the brain receives a large quantity of blood, fince the calibre of the four cerebral trunks has been eflimated at a lixth part of that of the aorta. This blood circulates in thefe veffels, forming at the bafe of the cranium an SYSTEM OF CIK-CULATION. 3S5 an anaftomotic circle, in fuch a manner that it would be poffible to tie any one of the cerebral arteries without the circulation of the blood in the brain being interrupted. The fecond defcends into the breaft, proceeds over the cartilages of the ribSj near the fternum, between the pleura and the inter-coftian muf- cles. Between each inter- cotl:al fpace it gives out two rami, which traverle the inter-coftian muf- cles, thofe above the breaft, and lofe themfelves in the mammella?. It dillributes itfelf alfo to the thymus gland, themediaftinum and the diaphragm. This artery then defcends towards the bottom of the breaft, furnifhes a ramus to the itcrnal appen- dix, and fome others which are loft in the fub- fiance of the mufcles of the abdomen, analto- moling with the ep'igajiric and ih^fub-Jienial (inte- rior mammillary). This artery furnifhes medlajiinei, Juh-mammillary ^ inter-cojial, 2iudf up 67' -diaphragmatic rami, and often a thymic and a bronchial ramus. The third arifes oppolite to the fub-fternal, and afcends before and on the fides of the neck, far- nifliing a ramus to the mufcles of that part: af- cending cervical. It then patfes under the cephalic, proceeds to the thyroid gland, and penetrates its fubftance. It there anaftomofes with rami of that of the oppofite fide, and of the fuperior thy- roidian. This artery diftributes itfelf alfo to the VOL. I. a c oefophagus^ 360 ORGANIC STRUCTURE, cefophagu?, the larynx, and as far as the bottorti of the pharynx: inferior thyroidian. The fourth arifeSj for the moll part, on the ex- terior fide of the thyroidian. It proceeds upwards on the lides of the neck, pailes between the tranf- verfe apophyfis of the laft cervical vertebra, and that of the firll: of the back, and afcends obliquely towards the occiput, furnifhing rami to the muf- cles over which it pafles : Irachelo-cervical (pofte- fior cervical.) The fifth, trachelo-fcapular, often forms two branches at its origin : one of thefe proceeds out- wards and downwards, under the clavicle, and gives out feveral rami to the neighbouring parts. When it reaches the fuperior edge of tfce fcapula, it pafles under the fuperior fuper-fcapulo- trochiterian, gHdes under the acromion apophyfis^ and lofes itfelf in the inferior fuper-fcapulo-tro- chiterian mufcle '. fuperior fcapular. The other, which' is larger, takes an oblique direc* tion upwards and outwards, under the fterno-maf- toidian mufcle, where it, gives out a ramus which Jofes itfelf on the fides of the neck. It then divides into two branches : the firft proceeds upwards, and diftributes itfelf between the dorfo-fuper-acromian and the trachclo-fcapulian mufcle : the fecond pro- reeds towards the fuperior e,^gG of the fcapula, paffing under the dorfo-fcapulian mufcle ; it then defcends SYSTEM OF CIRCULATIOTST. 38f ^iefcends along the pofterior edge of that bone, and divides itfelf, near its inferior angle, into feve- ral rami, which lofe themfelves in the fub-fcapulo- trochinian, the dorfo-fcapulian, and the cofto-fca-» pulian mufcles: the tranfverfe cervical. The lixth ifTues from below the exterior extre- mity of the fub-clavian. It dcfcends before the neck of the firft rib, often of the fecond, and lomeUmes of the third, and furnifhes feveral rami* Some of thefe are dire6led backwards, pafs over the fides of the vertebrae, and lofe themfelves ia the mufcles of the neck and the back. The reft proceed outwards, along the inferior edge of the ribs, betweeo the inter-coftal mufcles : fome rami are diftributed to the oefophagus and the bron- chiae : fuperior inter-cqfiaL 366. The axillary is a continuation of the fub-clavian. It proceeds outwards, forming an arc in the hollow of the arm-pit : it extends from the firft rib to the point where the large dorfal mufcle is attached. In this pafTage it furnifhes five principal branches, the diftribution of which is pretty con- ftant ; but they vary a great deal with refpe6l to their origin, as feveral of thefe branches often arife from a common trunk. The firft arifes before and in the middle of the axillary. It then immediately divides itfelf int<^ 2 c 2 feveral 388 ORGANIC STRUCTURE. feveral rami, which proceed to and expand be- tween the flerno-hiimerian and the cofto-coracoi-' dian mufcles : Jierno-thoracic (fuperior thoracic.) The fecond iffiies clofe to the preceding. It defcends below the fterno-humerian mufcle, and divides into feveral rami, which are diftributed to the coHo-fcapulian mufcle, the inter-coftians^ and the mammilla: cojio-lhoracic (inferior thoracic). The third arifes near the preceding ones. It pafles over the cofto-coracoidian mufcle, and fur- nifhes feveral rami, which are loft in the fterno- humerian and cofto-clavian mufcles; it then conti- nues between the fterno-humerian and the fub- acromio-humerian mufcles, and in this paflage fur- nifhes a ramus, which expands on the articulation of the humerus with the fcapula : the fufer-fca- pular (acromial). The fourth defcends polteriorly, below the point where the fub-fcapulo-trochinian mufcle is attached, and furnifhes fome fmall rami to that mufcle, and to the cellular tiffue of the arm-pit i it then divides into two branches : the inferior branch, which is fmaller, inferior fcapular, follows the inferior edge of the fub-fcapulo-trochinian mufcle, and lofes itfelf in the cofto-fcapulianj the lumbo-hurnerian, and the fcapulo-humerian. The other, which is ftronger, exterior fcapular , is dire6led backwards, between the fcapulo-hume- rian and the inferior fuper-fcapulo-trochiterian mufcles. SYSTEM OF CIRCULATION. SgQ mufcles, and divides into two rami. One of thefe raoii pafles over the anterior edge of the fcapula, and lofes itfelf in the inferior fuper-fcapulo-trochi- terian mufcJe, and another fmaller is diftributed to the fub-fcapulo-trochinian : fuh-fcapular (com- mon fcapular). The fifth ilTues from below the head of the nu- merus. It is often divided, at its origin, into two branches : one, the anterior circumflex, pailes under the coraco-humerian, and the fliort portion of the fcapulo-olecranian mufcle, and expands be- fore and around the articulation of the fhoulder. The other, which is much larger,' the pofteiior circumflex, palles behind the humerus, between the fcapulo-humerian mufcle and the inferior fuper- fcapulo-trochiterian, below the fub-acromio-hu- merian ; gives out a great number of rami, and lofes itfelf in turning round before and below the head of that bone -. fcapulo-humeral. 367. The humeral artery is a continua- tion of the axillary ; it extends from the middle of the hollow of the arm-pit, as far as the middle of the bending of the arm. In this paflage it fur- ni(hes above, towards the upper third of the hu- merus, a very large branch, the exterior coUateraJ, which proceeds outwards, betvveen the humero- cubitian and the fcapulo-olecranian mufcle, and divides itfelf into two branches. One of thefe branches diftributes itfelf in the fubftance of the a c 3 fcapulo" ^gO ORGANIC STRUCTURE. fcapulo-olecranian ; the other, which is ftrongef, turns round behind the middle of the humerus^ and defcends along the exterior fide of the arm, between the humero-fuper- radian, and the exte- rior portion of the fcapulo-olecranian : it terminates around the articulation, anaftomofing with the radial recurrent. Below this collateral branch arifes another, the inierior collateral, which defcends along the inte- rior iide of the arm, before the fcapulo-olecranian mufcle, and as far as the articulation. The humeral artery then continues before the arm, along the interior edge of the fcapulo-ole- cranian mufcle, and before the humero-cubitian, giving out to them frequent rami : the mujculat of the arm. In the laft place, the humeral artery gives out near the articulation of the elbow two xami, one of which proceeding towards the epicondyle dif- tributes itfelf to the neighbouring meifcleSj and anaftomofes with the interior collateral and the cubital recurrent. The other proceeds towards the epitrochlea, and is diftfibuted in the fame manner as the interior : collaterals of the articula- tion of the elhow. ^ At the bending of the arm the humeral artery divides into two branches, one of which follows the radial edge of the fore-arm, and the other its <^hital edge. 368. The SYSTEM OF CIRCULATION. 3gf 368. The radial artery extends along the anterior and exterior face of the fore- arm, as far as the extremity of a part of the fingers. Near its origin it furnifhes a branch, which is refledled ■ towards the epicondyle, recurrent of the epicondyk^ and is diftributed to the mufcles attached to that part, anaftomofing with the exterior collateral of the arm. In its paflage along the exterior edge of the fore-arm, the radial artery fnrniflies feveral rami to the neighbouring mufcles : inufcular of the fore- arm. Near the articulation of the wrift, this artery divides into two branches, after giving out a fmall ramus before that articulation. One of thefe branches, the radio-palmaris, pafles before the an- nular ligament of the carpus, gives out a ramus to the mufcles of the thumb, and proceeds to- wards the palm of the hand. It there furniilies an artery, which anailomofes with the cubital arch, and then terminates in two rami: one of thefe lofes itfelf on the radial fide of the thumb; the other defcends into the interval between the thumb and the index finger, and expands along the correfponding edges of thefe two fingers. The other branch, furniOied by the radial artery at its termination, proceeds to the exterior fide of the articulation, the radio-fuper-palmaris ; it gives out a ramus which proceeds in a tranfverfe direc- tion on the carpus, dorfal of the carpus, and an-^ a c 4 other ORGANIC STEUCTURg. ther which proceeds to the radial fide of tbe thumb. The radial artery, wLen it reaches the interval between the two firft bones of the metacarpus, feparates into three branches. One of thefe branches glides in the fubftance of the metacarpo- phalangian mufcle of the thumb ; pafTes along the tirft bone of the metacarpus, and divides into two branches, which are diftributed on the fides of the thumb. The fecmd pafles over the firft dorfal inter-ofTeous mufcle, along tbe radial edge of the fecond bone of the metacarpus. The thirds which is much larger, advances between the carpjan ex- tremities of the two firft bones of the metacarpus ? it turns round in the palm of the hand, and unites itfelf to a branch of the cubital to form the pro- found palmary arch. This arch furniihes, if!:, rami which are reflc6led before the carpus, to the ligaments of that part and to the mufcles of the thumb ; ad, four rami which pafs before the in-^ ter-oiieous mufcles, to vi^hich they diftribute them- felves as well as to the lumbricals, and terminate towards the heads of the bones of the metacar' pus, anaftomofing with the digital arteries ; 3d, three rami, which pierce tbe lecond, third, an4 fourth dorfal intei'-offeous mufcles ; diftribute themfelves to the fuper-palmary face, and each j:iivide into two ramufculi, which lofe thenrifelve^ on the tides of the laft four fingers. 0^6.^, The SYSTEM OF CIRCULATION. 3g3 369. The cubital artery, which is larger than the radial, defcends before, along the cubital edge of the fore arm, as far as the fingers. Near jts origin it gives out two branches, one of which pafles before the epitrochlea, diftributes itfelf to the neighbouring- parts, and anaftomofes with the col- laterals of the cubitus. The other, which is larger, pafles behind the epitrochlea, fends out fe- veral rami to the neighbouring mufcles, and anafto- mofes with the inter-mufcular of the arm : th& recurrent of the epitrochlea. The cubital then penetrates under the mufcles attached to the epitrochlea, and furnifhes two branches. One of thefe branches palTes between the inter-ofleous ligament and the mufcles by which it is covered : the anterior inter-ojjeous : it furnifhes a ramus which glides between the two layers of the mufcles, and fome others which are diftributed to the mufcles and to the periofteum, Inferiorly, this artery traverfes the inter-offeous ligament, and terminates on the convexity of the hand, where it anaftomofes with the dorfal of the carpus. The other, the pojierior inter -ojfeous, traverfes at the top the inter-ofleous ligament, and gives out a branch, the olecranian recurrent, which is refle6led between the olecranon and the epicondyle, in the fubftance of the epicondylo-cubitian mufcle. It diilributes itfelf to the neighbouring mufcles, and anaflomofcs 3Q4 ORGANIC STRUCTURE, anaftomofes with the exterior collateral artery. Thepofterior inter-ofleous then continues between the two layers of the mufcles of the fuper-palmary face of the fore-arm, to which it fends out a great number of rami, and terminates near the vvrilt, anaflomofing with the anterior inter-offeous. After fumifhing thcfe rami, the cubital artery pe- netrates, at the top, under the common epitrochlo- phalanginian mufcle, and ifTues towards the mid- dle of the fore-arm, from which it then proceeds as far as the wrift : the cuhko-fuper-palmaris. At this place it gives out feveral rami, one of whicli, diHributes itfelf over the hand, anaftomofing with the dorfal of the carpus. Another tranfverfe branch palTes before the articulation, and anafio- mofes with a limilar one furnifhed by the radial artery. A third branch pafTes under the tendons of the common epitrochlo-phalanginian mufcle^ and of the common cubito-phalangettian, and concurs to the formation cf the profound palmary arch. In the ]aft place, the cubital artery defcends into the palm of the hand, the cuh'ito-p.ahnar'is, under the palmary sponeurofis, and forms a tranf- verfe arch from which arife four principal rami, fometimes five, and even fix. The firft paffes over the mufcles of the palmary face of the little finger, and ternlinates along its cubital edge ; the fecond proceeds in the interval between the fourth . and SYSTEM OF ClRtULATIOK, SQS and fifth bones of the metacarpus, and feparateS^ into two rami, which terminate along the oppolite edges of thefe two fingers ; the third and fourth proceed, in like manner, to the fummi^ of the other two intervals between the following fin- gers, and lofe themfelves along their oppofite edges. When the cubital furnifhes one or two rami more, they terminate alfo on the fides of the index finger and the thumb. The fuperficial palmary arch is then formed entirely by the cubital, but for the moft part the latter rami -proceed from the radial. The two rami, which proceed on the fides of each finger, anaflomofe with each other towards the nail-like extremity. 370. The aorta, after furnifhing, towards its curvature, the cephalic and brachial arteries^ is re- iledled dire6i:ly downwards, and defcends, a little to the left, before the bodies of the dorfal verte- brae, between the lamina of the pofterior medi- aflinum on the right fide, and a little behind the oefophagus. It advances as far as the lower part of the brcafi:, traverfes the diaphragm between its two pillars, and penetrates into the abdomen, where it advances as far as the fourth lumbar vertebra. The. ^g§ ORGANIC STIltJCTUREe, The iborack aorta^ during its pailage, furniilies feveral fmall branches. I ft. It gives out one at the top, which expands on the pofterior part of the pericardium : ^ofterlor fsncardian. ad. It furniflies, at the top and on each fide, a branch which follows the diftribution of the bron- chia? in the rnterior of the lungs : tie hronchiaL The right branch, which in general is the largeft, arifes often from the trunk of the iirft inter-coftal : it turns round on the cefophagus, gives out to it fome rami, and proceeds to the origin of the bron- ©hiae, where it feparates into feveral rami, which sre diftriboted on the aerian canals, and "penetrate with them into th@ interior of the lungs. The left, which is often double, gives out mmi to the csfophagus and the pericardiumj and like the right diftributes itfeif in the lungs. 3d. Before the aorta are obferved three or four iiiiali branches^ diftributed along the csfophagus, and. which fend out fome rami to the pofterior mediailinum : the cefophagian. 4tb. The aorta fends out two or three rami^ which expand on the pofterior mediaftinum : the pofierlor mediqfi'me, 5th. Behind, and on the fides of the aorta, there arife from nine to twelve branches, which proceed on the lateral parts of the bodies of the vertebras, and are continued between the ribs; the inter- c.ojial. Thefe SYSTEM OF eiRCULATiOl& StSTEM OP eiRC0LATION'. 401 branch then continues towards the coeeum, and furnifhes a ramus to its appendix : the ccecal. From the convexity of the fuperior mefenteric artery arife, on the left, a great number of rami (from twelve to fifteen). Each of thefe feparates into two, which proceed in the folds of the mefen- tery, as far as the jejunum and the ileum. Alt thefe rami anailomofe vs^ith the neighbouring ones, and form arches, whence arife other rami which unite with their collaterals to form fmaller arches: this arrangement is repeated four and even five times. When thefe arches reach the inteftines, they give birth to two right layers of ramufculi^, which expand in the. fubftance of the fides of the inteftines, where they form a vafcular reticulation, which covers the whole furface of them. All the veflels diftributed on the inteftines ex- hibit, in general, a fimilar arrangement. 4th. The abdominal aorta gives out inferlorly, dole to its bifurcation, an artery which is diftri- buted to the mefo-colon and to the mefo-redlum : inferior mefenteric. This artery, near its origin, fends out a trunk which foon divides into two branches : ihefirji proceeds towards the tranfverfe portion of the colon, and furnifhes two rami, one of which afcends before the kidney, and proceeds towards the tranfverfe portion of the colon, where it terminates by uniting with a branch of the fu- perior mefenteric, and forms there a remarkable VOL. J. 2D anafiomofis: 402 ORGANIC STRUCTURE. anaftomofis : that of IlioJan, The other follows the dire6lion of the defcending portion of the colon, and terminates by anaftomofing inferiorly with the neighbouring rami : left great colic (fiipe- rior and middle colic). The/econdj which is not fo large, proceeds to- wards the defcending portion of the colon, and divides itfelf into two rami which anaftomofe with their collaterals : thefmaU left colic. A third is dif- tributed to thecommencement of thefigmoid flexure of the colon, and anaftomofes with its neighbours. In the lafl place, the inferior mefenteric artery, after giving out fome rami to the iliac part of the colon, defcends on the right, penetrates into the lefler pelvis, diftributes itfelf behind the re^lum, and anailomofes with feveral of the neighbouring rami : artery of the reBum. 5th. The abdominal aorta, immediately after furnifhing the fuperior mefenteric, gives out, on each fide, a branch which divides into feveral rami. Some of thefe proceed to the pillars of the dia- phragm, and others to the fuper-renal capfules, and the neighbouring adipofe tifllie : the fu-per- r^«<2/ (middle capfular). 6th. Below the latter arife one, two, and fome- times three, on each fide of the aorta, which pro- ceed tranfverfally towards the kidneys, ths renal, furnifhing fome ramufculi to the fuper-renal cap- fules, the neighbouring adipofe tifTue, and the ureter. SYSTEM OF CIRdULATION.' 40*3 ureter. When thefe branches reach the kidneys^ they divide into feveral rami, which lofe themlelves in the finuofities of that vilcus. 7th. Below the renal rami there arlfe, in-generalj on each fide of the aorta, two Jong flender arteries, which proceed downwards in an obHque dire(?tion, along the pfoas, giving out fome fmall rami to the neighbouring parts. Each then iflues from the abdomen, through the inguinal ring, defcends along the tefticular cord, and proceeds to the epi- didymis and the tefticles: the teji'icuJar (fpermatic). In the female, this artery penetrates into the lefler pelvis, and diftributes itfelf to the ovariurh, giving out fome rami to the broad ligaments, the conduits of the ovaria, and the lateral parts of the matrix: artery of the ovarium, Sth. The lateral and pofterlor parts of the ab- dominal aorta give birth, on each fide, to four or five arteries, which proceed tranfverfely on the bodies of the firft four of the lumbar vertebrse. Thefe arteries furnifh fome ramufculi to the neigh- bouring parts, and then divide into two rami. Some of thefe proceed backwards between the tranfverfe apophyfes, give out rami which pene- trate into their rachidian canal, and lofe themfelves in the mufcles of the back. The reft pafs behind the praelumbo-trochantinian mufcles, fend out rami to the pillars of the diaphragm, and to the ilio-tranueifian; are refle6ied on the mufcles of a D 2 th« ORGAl^IC STRUCTURE. the abdomen, and terminate by anaftomofing with rami of the laft inter-coflals, of the fub-ftcrnal and fub-pubian : the lumhar. 9th. Behind the abdominal aorta, and near its bifurcation, is an odd artery, which defcends on the middle of the body of the laft lumbar vertebra, before the middle part of the facrum, and as far as the coccyx : middle facral. In this palTage it furnifhes on each fide a ramus, which proceeds in a crofs dired^ion on the body of the laft verte- bra, and anaftomofes with the neighbouring parts. It then continues and gives out feveral rami on the fides of the facrum. 372. The abdominal aorta, when it reaches to- wards the fourth lumbar vertebra, divides into two branches : pelvi-crurales (iliac). Thefe two branches proceed downwards, and on the fides, near to the articulation of the facrum with the bones of the ilium, where they again bifurcate, and farnifh two other branches: one of thefe penetrates into the pelvis, the pelvian ; and the other pro- ceeds to the abdominal limb : the crural. In the foetus, the peiyi- crural feparates alfo into two branches, one of which proceeds to the abdo- minal limb, and the other^ which is much larger, penetrates into the pelvis. The latter gives birth to two branches furniOied in general by the pel- vian, but in that cafe they are only very delicate rami % the body of the artery runs along the fides SYSTEM OF CIRCULATION. 405 iides of the bladder, and then proceeds to the um- bilicus. ^ 373. The PELViAN ARTERY penetrates into the lefler pelvis, and gives birth to eight principal branches, or arterial bundles, which extend over its fides, and are diftribut^d to the different organs it contains, ill. At the top and behind, a branch which af- cernds on the fide, and divides itfelf into two rami ; the firfi: proceeds on the lateral parts of the bodies of the laft lumbar vertebrae, and is diftributed on the praelumbo- and iliaco-trochantinian mufcles, and enters into the rachidian canal. The fecond afcends outwardly, and diilributes itlelf in the iliac fofla : U'mco-mufcidar. 2d. Several branches which proceed on the fides of the facrum, furnifh rami to that bone, and others which proceed, through the facral holes, into the vertebral canal : lateral facral, 3d. An artery which proceeds before, and on the fides of the pelvis, as far as the interior fub- pubio-trochantinian mufcle: the fuh-^ubio-femoraL In this paflage it fends fome rami to the prae- jumbo- and, ilio-trochantinian mufcles, pafi!es through the fub-pubian hole, traverfing the inte- rior (ub-pubio-trochantinian, and gives two rami, one on the right and the other on the left, which diftribute themfelves upwards and inwards to the muf2les of th^ thigh. 4th, Before, an artery very large in the foetus, 203 and 400 ORGANIC STRUCTURE. and in a great part obliterated in adults. In the latter^ it furnifhes feme rami, which are diflributed on the fides of the bladder, to the matrix, the va- gina, &c.: the umViUcal. 5th. Above the latter, fome branches which proceed to the bladder, the re6lum and the uterus. One of thefe branches pafles between the bladder and the .redlum, and diftributes itfelf to thefe or- gans, to the proftate gland and the feminal veifels. In the'female, this artery gives out a confiderable branch which penetrates the broad ligament, runs along the fides of the matrix, and the Fallopian tube, proceeds to the ovarium, and anaftomofes with the artery of that part. Another ramus is diflributed to the vagina : the veftco-projiatic (ute- rine and vaginal veficals). 6th. The pelvian artery, near its termination, gives out a large branch, which diftributes itfelf to the mufcles of the nates, fejfiere (pofierior iliac), Thisbranch furnifhes, in the lefi^r pelvis, fome rami totheredium, and to the facro-trochanterian muf- cle : it then iflTues from this cavity^ pafi^ing by the fummit of the ifchiatic notch, and foon fepafates into feveral branches, which expand between the great and the lefler trochanterian and the facro- femorian mufcles. 7th. Below the preceding arifes another artery, which gives fome rami to the re6tum, the bladder, and the uterus. It then iffues from the pelvis gbove the facro trochanterian mufck, towards the lower SYSTEM OF eiRCULATlON. 40/ lower part of the ifchiatic notch, and divides itfelf into fcveral rami, which expand under the facro- femorian mufcle, towards the coccyx, the tubero- fity of the ifchium, and to the pofterior mufcles of the thigh : the ifchiatic. 8th. In the laft place, the pelvian artery gives a branch, which arifes generally from a trunk com- mon to the preceding. It iflues from the pelvis, between the pubio-flib-umbilian and the ifchio- coccygian mufcles, advances on the interior face of the ifchiatic tuberolity, and terminates at the organs of generation .- the fuh-^elvian (interior pudical). In the pelvis, it furnifhes rami to the bladder, and often a large ramus to the middle part of the re6lum. On the outiide of this cavity, it gives rarhi to the facro-trochanterian and the facro-femorian mufcles ; it then furnifhes two branches : one of thefc, which is fuperficial, pafles over the tube- rolity of the ifchium, and proceeds to the fcrotum : the perinaaJ : the other penetrates to agreater depth between the tuberofity of the ifchium and the canal of the urethra, and gives rami to the mufcles of the anus and of the perinasum : ifchio-ferimeal (tranfverfe of the perinaeum). When it reaches the lower part of the fymphyfis of the pubis, this branch divides itfelf into two other rami, one of which pafles under the root of the cavernous body, proceeds over the back of the penis, and termi- 2 D 4 nates 408 ORGANIC STRUCTURE, nates in the fubftance of the glans : the other lofes itfelf in the cavernous body. In the female, the perinaeal artery lofcs itfelf in the large labia ; and the profound one proceeds to the clitoris : the ifehio-clitorian, 374. The crural artery arifes before the jun6i:ion of the facrum with the hip-bone; it pro- ceeds towards the ilium, palles under the crural arch, defcends firll on the interior lide of the fe- mur, and then behind that bone, and terminates at the ham, where it feparates into two branches^ which are diftributed to the leg and to the foot. The crural artery, in order to facilitate the de- fcription of it, is divided into three portions: one correfponds to the pelvis, near the iliac fofTse (the iliac) ; another correfponds to the groin (the in- guinal); and the third extends along the thigh (the femoral). The ILIAC PORTION farniflies interiorly a. branch which proceeds on the pubis, Juh-pubum (epigaftric), and is reiled^ed under the fides of the abdomen, chiefly along the fierno-pubian, forming feveral anaftomofes. It gives out a ramus, which accompanies the tefticular cord as far as the tefli- cles, and terminates below the flernum, anafto- mofing with the fub-fternal. The other artery, furnifhed by the iliac portion, afcends interiorly as far as the anterior angle of the ilium, turns round on the edge of that bone, and SYSTEM OF CIRCULATION. 4Qg ihnd expands in feveral rami in the mufcles of the abdomen, and in the ilio-trochantinian, producing fome anaftomofes : circuvijiex of the Ilium (anterior iliac). The INGUINAL PORTION fumiihes exteriorly fome branches, which are ramified in the inguinal glands, the mufcles and neighbouring integu? ments : the inguinal (tegumentous of the abdo-^ men.) Interiorly, this portion gives out two or three fmall branches, which are diflributed to the region of the pubis, to the fcrotum (thefcrotalj, or to the vulva, the vulvar (exterior pudical). The FEMORAL PORTION givcs to the upper part of the thigh a large branch, which penetrates be- tween the triceps addudlor and the tri-femoro- rotulian mufcle, and is diftributed behind the thigh : the inter-mujcular or profound of the thigh. This artery furnifhes, near its origin, two large rami, one of which proceeds inwards between the fub-pubio-femorian and the tendon of the pras- lumbo-trochantinian mufcle, and terminates in fe- veral ramufculi, which are diftributed to the muf- cles of the fuperior and interior part of the thigh, around the articulation, and to the exterior parts of generation : the fuh-trochantinian (the interior circumflex). The other defcends exteriorlvj pafTcs between the ilio-pra;tibian and the ilio-rqtulian mufcles, and 410 ORGANIC STRUCTURE. and divides itfelf into two rami, the firft of which is refle6led around the articulation, and lofes itfelf in the mufcles of the pofterior part of the thigh : the fecond defcends before the thigh, and expands between the ilio- and the trifemoro-rotulian : the Jub-trochantenan (exterior circumflex). The profound artery then defcends, and gives out a great number of rami, which are diftributed interiorly and pofterioily to the mufcles of the thigh : it furnifhes alfo a ramufculus, which peme- trates into the thigh. The femoral artery^ after fending out the pro- found one, defcends below the ilio-prstibian muf- cle, furni(hing in its paflage a great number of rami to the neighbouring mufcles : the mufcular. Towards the lower third of the thigh, it traverfes the tendon of the third addu6lor, proceeds to the pofterior part of the thigh, and defcends along the ham behind the articulation, where it afTumes the name of the poplitean. In this fhort pafiage it furnifhes four branches, two on each fide, the one above the other : the poplitean articular (the fupe- rlor and inferior articular). Of the two fupcrior, one (the interior) afcends within, pafTes above the interior condyle of the femur, and diflributes itfelf to the articulation of the knee. In this paflage, it gives rami to the neigh- bouring mufcles, to the interior lateral ligament, the articular capfule, and upon the rotula. The other SYSTEM OP CIRCULATION. 41! other (the exterior)^ which is more profound, af- ccnds on the exterior cond}']e of the femur, and diilributes itfelf to the articulation, fending out rami, which terminate, like the preceding, at the iimilar parts of the opposite lide. Below thefe branches, the poplitean artery pro- halangettian mufcle, and forms a curvature. The concavity of this curvature furnifhes fome rami to the articulation of the bones of the tarfus. From its convexity three rami proceed^ which advance towards the bones of the metatarfus : when they approach the heads of thefe bones, they divide themfelves into two ramufculi which expand on the SYSTEM OF CIRCULATION. 413 the fides of the toes. The other, which is larger, Droceeds on the tarfus, as far as the os cuboides, where it gives out a branch which pafles over the firfl inter-ofleous mufcle, and advances towards the anterior extremity of the firft bone of the meta- tarfuSj where it divides itielf into two ramufculi, which lofe themfelves on the fides of the firft and fecond toes. The trunk of the anterior tibial penetrates be- tween the heads of the firfl: and fecond bone of the Ti\&\2iXzx{x\.'^,fuper~metaiarfian, and divides itfelf into two branches : one of thefe branches anaftomofes with the exterior plantaris^ and concurs towards the formation of the plantar arch : the other dil^ tributes itfelf under the great toe. 376. The posterior tibial, at its origin, fends out a very large branch, which defcends be- hind the perone : the •peroneal. This artery is co- vered by the tibio-calcanian,and the peronso-fub-, phalangettian mufcle of the great toe ; it fends out a great number of rami, which are diftriljuted to ^he mufcles of the poflerior and exterior part of the leg. The peronfeal then defcends between the extremities of the tibia and of the perone, and di- vides itfelf into two branches, one of which pafies under the calcanian tendon, diftributes itfelf around the articulation, and terminates on the ex- terior fide of the foot by feveral aaafloraofes. The other 414 ORGANIC STRUCTURE. other pierces the inter-offeous ligament, defcendls before the articulation of the tibia and the perone, and terminates in feveral rami, which proceed over the foot, anaftomoting with the fuper-tarfian and the exterior malleolar. The pofl:erior tibial then defcends behind the tibia, under the tibio-calcanian mufcle, as far as the lower part of the leg, where it becomes fuper- ficial. In its paflage, it fends out a great number of ra- mi to the mufcles of the pofterior part of the leg, and a ramus which penetrates into the tibia : the medullary of the tibia. When it reaches the inte- rior fide of the calcaneum, it divides itfelf into two branches : one of thefe branches follows the interior edge of the foot, the interior ■plantaris ; fends out rami to the mufcles of the great toe, and terminates on its interior fide, anaftomofing with fuper-meta- tarfian rami. The other, the exterior plantaris. which is larger, penetrates between the mufcles of the fole of the foot, to which it diflributes itfelf. When it approaches the fifth bone of the metatar- fus, it is refledted inwards, and forms a curvature, the convexity of which is turned forwards. This curvature fends out fome rami, which proceed backwards, and diftribute themfelves in the muf~ cles, on the bones of the metatarfus. It furnifhes three rami which afcend through the metatarfo- inter- « SYSTEM OP CIRCULATION. 415 inter-phalangian mufcles. It then fends out four larger rami, in the interval of the bones of the metatarfus ; thefe rami, when they arrive between the heads of thefe bones, divide into two ramuf- culi, which expand on the fides of the toes. All the rami which proceed on the tides of the toes anaftomofe at their extremities. TABLE / [ 41§ 1 TABLE OF THE HEART AND ARTERIES. S77. The blood, which returns from every part of the body, is conveyed to the heart by two large veins, the vexa cava; these veins pour it into a sinus, sitms of the 'vena cava, (right or pulmonary au- ricle,) through two orifices, one at the top, the other at the bottom : the latter is furnished with a valve, or portion of valve, in the form of a crescent. From this sinus the blood passes into the right or pulmonary ven- tricle, through a circular aperture furnished with a tricuspid valve. It issues from this ventricle by an orifice provided with a triple semi- lunar valve (sigmoid), and proceeds into the pulmonary artery. The pulmonary artery, which contains venous blood, divides it- self, after a short passage, into two trunks which proceed to the two lungs. The right trunk divides into three branches, and the left into two, each of which is divided and subdivided indefinitely, and. terminates in exceedingly fine ramusculi, which proceed ,on the sides of the last bronchial cells, into which the air penetrates during respiration. These arteriolae, it is probable, anastomose with the veins which bring the blood from every part of the lungs. The small veins unite to form venous rami, which produce branches, and the union of branches forms two large trunlts for each lung. The four pulmonary veins contain attcrial blood j they proceed into a sinus, sinus of the pulmonarj/ veins (left or aortic auricle), which is smaller than the preceding, and the sides of which are thicker. From this sinus the blood passes into the left or aortic ventricle, through a round aperture furnished with a two-pointed valve (val- vula mitralis). This valve is smaller and thicker than the preced- ing j it drives the blood into the aorta, whence it is distributed to all the parts, by means of numerous arteries which are divided and sub- It furnishes the laryngian and several rami. £d. One proceeds to the tongue and to its different muscles^ The lingual. . . . . . The same. It gives the sub-lingual and a hyoidian ramus. Sd. One furnished to the pharynx and the velum palati, Inferior pharynsian. . .... the same. It sends out the posterior meningian. 4th. One is distributed to the maxillary gland, ^to the jaws, t» the commissurcE of the lips, and as far as the tip of the nose, , The labial. ■ . , . , E.vJcrfsr maxiltan'. It TABLE OP THE HEART AND ARTERIES. 4I9 It o-ives out, the muscuh-palatine, the sub-maxillary, the tonsillar}/, the inferior labial, the two coronary labial, and soine rami. 5th. A branch which proceeds to the occiput, NEW NAMES. OLD NAMES. The occipital • The same. - It furnishes muscular and cutaneous rami. 6th. One expands behind the ear, within that organ and on the occiput, Poiterior auricular. ... . The same. It gives out the stylo-mastoidian, the tympanic, some muscidar and cutaneous rami. 7 th. One distributes itself on the sides of the head to the parotid gland, before the ear, to the cheek, and to the integuments of the upper part of the head, The temporal Tfe samt. It gives the anterior auricular, the sub-zygomatic (trSinsvevse of the face), and the superjicial temporals. 8th. One proceeds to the meninx, to the teeth, to the muscles of the jaw, to the cheek, to the pharynx, to the velum palati, and to the nasal fossae, The maxillo-buccaL . . . . ■ . Interior maxUlaryi It gives the middle ?nenijigian, the maxiilo-dentary (inferior deo- tary), the pterygoidian, the profound temporals, the zygomato-max- illary (ihasseterine), the super-maxillary (buccal), the sub-orbitar, the superior pharyngian, the palatine, and the spheno-palatine. B. The ANTERIOR CEREBRAL penetrates into the cranium, and at its entrance gives a branch which proceeds in the orbit, distri- buting itself to the lacrymal gland, to the eye and to its muscles, to the eye-lids and in the nose, Theorbitary. .... . Ophthalmic, Which gives the lacrymal, the sub-irian (ciliary), Me ethmoidals, the central of the retina, the muscular, the palpebral, the frontal, and the nasal. The anterior cerebral is afterwards distributed in the anterior and middle lobes of the brain : It furnishes. The communicitiBj, . . . . The same, 2 E 9 Which 420 ORGANIC STRUCTURE. Which gives sotne rami to the choroid plexus, choroidian, NEW NAMES. OLD NAMES. The sntcrior lobai". . • - : • Artery cf the corpus callosum. The middle lobar. III. The BRACHIAL TRUNK, v/hich in its passage along the thoracic liiT:b is dhnded into sub-claviar, axillarv\ and humeral. A. The suB-CLAViA" furnishes six branches : lst> One is distiibiited to the posterior part of the brain, to the cerebcUuui, and to the rachidian prolongation, The posterior cercbial. . .... The vertebral. It ftcr'tiishes. The occipito-meningian. .... Pcsterlor ^>entn^'-:in. The anterior snd potteiior median of the rachis . A-nierKr an', po^terhr si :nal, Tlici .feri.^r ^arge ceiebeilous. . . I'lferior ',f tk- ctnlnl.m. The raeencephalic. . • . . Th' bi i'ltjry, Thr pos'erior lobar Tiir posterior or in/ert-.r cf the cerebrum. Thp 'tife'ior sTiail cerebeliou:. . ". . . Inferior c,,' the cerebnium. Th= siljclior iCrcieUou:. . . . Superior cf the cerebellum. 5;d, Cne Is dj' tritutcd be^o^v the stcnum, to the thymus, to the medi"iciiui!r., tc the d;?,phrLgm, and to the parietes of the abdoilien. The sub-sternil. . ... Int-rl-r j.-.in.viUlcry, It i^ives '7i7ediastiiv', siib-inaniini.la,'',', inier-costal, super-dia- phragniaiic (superior di?-phr^.2■lr.a^ic) rnmi, 'V'^ii-^^fur.ihf! a ''hijmic and Ircnchial rnrnus. ,'' ' ~. . ; :c;.cj- .; CO he thj'roid ^Liua ^na to the muscles of the iic- :, Iiifeiior thyroidiaii The snme. It furnishes the ascending cervical and several ravti, 4th. One proceeds on the sides of the neck, The trachelo-cervical. . . . . . Puterhr profoufj c-rv:ca!. 5th. One is distributed to the muscles of the lateral parts of the neck and of the shoulder, Trachelo-scapular. . .... Tramversecei'vkctl. It furnishes. The superior scnpnLiry. 6th. One proceeds beneath the first rib, and gives some rami to the oesophagus, Superior inter-costa!. . . ... The same, B. The TABLE OP THE HEART AND ARTERIES. 421 B. The AXILLARY furnishes five principal branches : 1st. One branch is distributed to the pectoral muscles, / NEW NAMES. OLD NAMES. Tilt sterr.o-thoracic. . . . . . The sv^^yioi thor.ick. 2d. Another to the costo-scapulian and the inter-costian muscles, and to the breast, The costo-(horacic. • . . . . Thf in fir hr thoracic. 3d. One proceeds on the summit of the shoulder. The supcr-scapularj'. . . " . . . Acrcmial. 4th. One proceeds to the muscles of the interior side of the scapula. Sub-scapular. . . . . . . . Common scapular. 5th. One is distributed around the articulation of the shoulder with the arm, The scapulo-humera!. . . . : . Anterior and^fjstirior circumpx. C. The HUMERAL ARTEnY (brachial) furnishes three principal branches before it divides at the bend of the arm : 1st. One branch penetrates profoundl}' into the muscles oi the arm. The inter-muscular or profound. It furnishts. The interior and exterior collateral. 2d- Ssveral rami are distributed to the muscles of the anterior part of the arm. Muscular of the arm. Sd. Two rami proceed on the sides of tii? articulation of the elbow ; Collaterals of the articulation of the elbow. The humeral artery, when it reaches the bead of the arm, di- vides into two branches. The radial and the cubitaL A. The RADIAL furnishes four principal branches : 1st. One which is reflected towards the epicondyle. Recurrent of the epicondyle Radial recurrent, 2d. Rami which proceed to the radial side of the fore-arm, Ths mujcularolthefoie-arm. , , , . 2 E 3 3d. A branch 422 ORGAKIC STRUCTURE, Sd. A branch which is distributed in the palm of the hand to the last fingerSj and which concui-s towards the formation of the super- ficial paimar arch, NEW NAMES. OLD NAMES. The radio-palmar, . . . . . The palmar raiial. 4tb. A branch which descends on the back of the hand, sends out a ramus to its pahnar surface, and concurs towards the formation of the profound palmar arch, Kadio-super- palmar. ..-.-. Dorsal Radial. B. The CUBITAL furnishes four principal branches : 1st. Two are reflected towards the epitrochlea where they are distributed, Eecunent of the epitrochlea. .... Anterior arid posterior cubital recurrent, 2d. Two are directed on the faces of the inter-osseous ligament. Anterior inter-osseous. The intcr-osscous, I Posterior inter-osseous. The posterior inter-osseous furnishes a branch which is reflected towards the olecranon, Olecranian recurrent. ..... Posterior radial recurrent. Sd. One is distributed to the back of the hand, Cubito super-palmar. .... . Cubital dorsal. 4th. One proceeds in the palm of the hand to the first three fin- gers, and forms in a great measure the superficial palmar arch, Tlie cubiio-palmar The palmar. THE AORTA GIVES IN THE THORAX: I'st. A branch, which is distributed to the posterior part of the pericardium, Posterior pericardian. 2d. Two arteries, which proceed to the bronchiae and are rami- fied with them in the lungs. The bronchic. ... ... Bronchial. Sd. Some rami, which are distributed to the cesophagus. The teophagian The samp. 4th. Some small arteries, which expand on the posterior part of the media^iinum, Posterior mediastirie. ... . . The somt. ^ 5th. From TABLE OF THE HEART AND ARTERIES, 423 Slh. From nine to twelve branches, which divide between the heads of the ribs into two rami, one of which is distributed to tiie muscles of the back and the other to the inter-costal muscles, Aew names. ^e inter-coscal. . . « OLD NAMES. • Inferior or aortic intercostal!. THE AORTA FURNISHES IN THE ABDOMEN, I. Two branches, which are distributed to the two sides of the ab- dominal surface of the diaphragm, and give some rami to the super- renal capsules and to the liver, ^ub-diaphragmatic. . . . • . Inferior diaphragmatic. II. A large trunk situated beneath the stomach, THE OFISTRO-GASTRIC Cdliac. Which divides into three branches i 1st. One is distributed to the stomach. The gastric. - . . - Stomachic cormtrj. It furnishes sometimes The left lobar of the li-!er. . . . Li/t gastro-hepatic 2d. The other proceeds to the liver. The hepatic. It gives rami . To the pyloruSt The jastro-pyloric . . . , Pyloric. To the pancreas and the duodenum^ The pdncrtatico-dmdenal. To the stomach and the epiploon, Rijht gastro-epiploic Jhe same. To the gall bladder , The cystic, 3d. The last proceeds to the spleen. The splenic. ..... The same. And gives branches To the pancreas. The spleno-pancreatic. To the stomach. The sf Icna-eaiiric . -» <> » . < Short vaiclH ,424 OJKJANIC STRUCTUHE. To the left side of the great epiploon, KEW NAMES. OJ.D i'AMES. LcfE saicro-epiplcic. III. A large branch, v^^hich is distributed to the mesentery of the ^mall intestines, to these intestines and to the coecum. The superior mesenteric. . . , , The same. It gives rami To the pancreas, 'Ihe pancreatic. To the duodenum. The duodenal. To tht nieso-colon, The meso-colic. To the right colon. The right colic. To the coecum, the ilium, and the colon. The ilip-colic. ^ To the co?cum. The cffical. IV. A branch, which arises near the bifurcation of the aorta, and is distributed to the mesentery and the large intestines, The inferior meienteric. . . . . The sam -. It furnishes, The left great colic. . . . . ) Superhr left colic. 2, Mi We left cUic. The small left eolic f Ai tery of rlic rectum. . . . . . ^mfi^'O'-m colic. V. Two branches, which are distributed to the super-renal cap- sules and to the cellular tissue of the kidney. The super-renal. . . , Middle capsular. VI. Two large branches, which proceed to the kidneys. The renal. t jlie same. They furnish super-renal, adipose, and ureteric rami, VII. Two small arteries, which proceed to the testicles or to the ovarium. The testicular. . . , . Thssftymatlc. Ai tery oi the ovarium. VIII. Four ' TABLE 'OP THE HEART AND ARTERIES. 425 VIII. Four or five arteries, -which are distributed to the muscles of the loins and to the parietes of the abdomen, NEW NAMES. OLD NAMES. The lumbar !« ■same. IX. An odd branch, which arises betwiftn the two divisions of the aorta and descends on the sacrum. The middle sacral. THE AORTA AT ITS TERMINATION DIVIDES INTO TVl'O TRUNKS: THE TELVI-CRURAL. . The primitivi iliac. The pelvi-crural artery, which arises at the height of the fourth lumbar vetebra, descends outwardly, and after a short passage di- vides into two branches 3 one of which is distributed in the pelvis. THE PELVIAN. . Interior or hypogastric iliac. The other issues from the pelvis through the crural arch, and de- scends along the thigh, TIIE CRURAL The same. A. The PELVIAN divides into eight principal branches or ar- terial bundles : 1st. A branch is distributed in the muscules of the iliac fossa. The iliaco-musailar Ilio-lumhar. 2d. Several proceed on the sides of the sacrum, The lateral sacral Tiic same. 3d. One issues from the pelvis through the sub-pubian holf, and is distributed to the three adductors. The iub-pubio-femoral. ..... O^Jturr.trix. 4th. Another, considerable in the foetus and almost obliterated in adults, is distributed to the bladder, to the utei us, and to the vulvo- uterine eonduit. The umbilical. . - The santf. 5th. One proceeds to the bladder, to the prostate gland, ^ to thd rectum, to the uterus, and to the vaginal conduit. The vesico-pvostatic. Vter'ine and vaginal vesical. 6th. Brancli 426 ORGANIC STRUCTURE^ " 6th. Branches which proceed to the muscles of the nates, NEW NAMES. OLD NAMES. The fessieres - • ^ posterior iliac, 7th. A branch passes under the sacro-feniorlan muscle, acconi" panics the sciatic nerve, and loses itself in the muscles of tfae pos- terior part of the thigh, - The ischiatit:> Sth. The last is distributed to the organs of generation, Ilie sub-pelviin^ ..... The interior puiicaL The latter furnisheSj the perineal, . ... ... Transverse of the perinautn. The ischio-penian Dorsal oftheyari. Theischio-clicorian. . . ... Artery oj tht clitoris . B. The CRURAL divides into three portions: the iliac, inguinal, and femoral. I. The ILIAC PORTION furnishes two branches, one of which is- reflected on the sides of the abdomen, and loses itself in the muscles. The supcr-pubian. . ^ - ^ . Spgajtriic^ The other follows the direction of the ridge of the ilium, and loses itself in the muscles of the abdomen, , circumflex of the ilium. Anteritr iliac. II. The INGUINAL PORTION fumishes branches, which are ra- mified to the glands of the groin and to the neighbouring cellular tissue. The inguinal. . Tegumentous of the abdomen. And some branches, which proceed to the scrotum or to the vulva. The scrotal. ... The vulvar. .1 Exterior fuiical. III. The FEMORAL PORTION gives: lEt. A large branch, which is distributed profoundly in the mus- cles of the thigh, Ifitcr-muEcalar or profound. . ■ • . Prof lund artery. The TABLE OF THE HEART AND ARTERIES. 427 The latter , furnishes from its upper part two branches, one of •which proceeds inwards and turns round on the articulation of the femur, ' KEWNAMESf. OLB NAMES. Sub-lrochantinian._. . . - . . Inttrhr circumfia: The other loses itself on the exterior side of the articulation, Sub-trochantinian. ..... Exterior circumflex. 2d. Branches along the thigh. The muscular. 3d. Four branches, two on each side, and one above the other, ■which are distributed to the articulation of the knee. Popliteal articular, . , , . . . Superior and inferior articular. The femoral, when it reaches the ham, divides itself into tvvo branches : The anterior and posterior tibial. a. The anterior tibial divides itself into four branches ; 1st. One small one is reflected towards the knee, 'Recurrent of the knee. 9d. Two proceed towards the sides of the articulation of the foot. The malleolar. . . . . Sd. A branch is directed on the tarsus, and distributes itself to the muscles and bones of that part, The super-tarsian. .... Arttry of the tarsus. 4th. The last forms a curvature on the convexity of the foot, The auper-metatarsian. .... Artery of the metatarsus. I. The POSTERIOR TIBIAL gives first two branches: 1st. One descends behind the perone, and is distributed io the muscles of the posterior part of the leg. The peroneal. . ... thtferoneal. 9d. The other, which is very small, pcnenetrates into the medullary canal of the tibia through its nourishing foramen, Mrd'illiry of tli« tibia, . , » ^'ckriihlnt ^tficsiiia. o At 428 ORGANIC STRUCTURE, At the lower part of the leg the posterior tibial divides itself into two branches : 1st. One follows the interior edge of the foot and terminates on the great toe, NEW NAMES. OLD NAMES. Interior plantar. .... The same. The other penetrates below the sole of the foot and forms the plantar arch, from which rami proceed to the muscles of the sole of the foot and to the toes, txteiior plantar. . ... The same. END OP THE FIRST VOLUME, W.i;ks and Taylor, Vsmtea, Chaatet^-Lar.e. v.^fcJt*V?»*2f-; '^^^S€¥^i^K^;j^'" :*5^. mf\