J/ 3

SOUTHERN CALIFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden, Claremont, CA 91711

CROSSOSOMA Vol . 13, No. 1
Managing Editor: Allan Schoenherr

February, 1987

Tecate Cypress in Orange County

by

Connie Spenger

1318 E. Glenwood Avenue

Fullerton, CA 92631

Tecate cypress is a rare species, existing as a relict in
isolated stands in upper and lower California. During previous
glaciations, it was more widely distributed, but with the coming of
a warmer and dryer climate, the cypress retreated to its present
locations .

Recently renamed as a subspecies of the Guadalupe Island Cypress,
( Cupressus guadalupensis spp. forbesii ) Tecate cypress' classification
is controversial. Some botanists prefer the old name, forbesii .

There are four stands of Tecate cypress in California, the
northernmost is near Sierra Peak, Orange County. Three populations
occur in San Diego County on Otay, Tecate, and Guatay mountains.

The remaining 11 stands are in Baja California, Mexico, including
some large ones and a few as small as only a dozen or so trees in an
arroyo. Tecate cypress occurs in conjunction with chaparral. In
Mexico, closed-cone pines are also found with the cypress.

Tecate cypress is a small tree with bright green leaves and
reddish or mahogany-colored bark, which exfoliates in thin curls.

Cones remain in place for the life of the tree. Tecate cypress trees
are conical when young, but those older than about 80 years develop a
rounded, spreading form which has a picturesque appearance.

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The physical features of Tecate cypress are such that a stand
depends on fire for 90% of its reproduction. Tecate cypress' thin,

curling bark lifts the fire upward (Dunn, 1984) . There the fierce
heat melts the cones' tight resin seal, which evaporates in a puff

of smoke, while the cones char, insulating the seeds within from
high temperatures. After a fire, cones slowly open to release Lheir
seeds. As cones are produced only after the tenth year of life, and
optimum seed production begins at about age 35, fires occurring at
frequent intervals are a threat. In fact, half the Tecate Peak stand
has been lost in this century due to too-frequent man-caused fires
(Dunn, 1984) .

The northernmost stand contains both the oldest and the largest
known Tecate cypress trees. The oldest are found on the Fremont
Canyon ridgeline at about 2,300 feet elevation, and the largest is
on the floor of Coal Canyon at about 750 feet. The stand occurs on
the 65-million-year-old sedimentary substrates of the Silverado
formation.

Other rare plants occur in and near the northernmost Tecate
cypress. These include:

1. Heart-leaved pitcher sage ( Lepechinia cardiophylla ) . This
is a large shrub with striking white flowers. It is a co-dominant
with the cypress. Heart-leaved pitcher sage is at its most abundant
location in the northern Santa Ana Mountains with Tecate cypress.

2. Fire-follower, Braunton's milk-vetch ( Astragalus brauntonii )
appeared in two locations after the 1982 Gypsum Canyon burn. One of
these populations had last been seen in the 1930 's; the other was a
new discovery. Of A^ brauntonii 's 13 recorded locations, about half
have not been seen since the 1940 's or even earlier, and at least
three are probably extirpated. None of the 13 known Braunton's
milk-vetch populations occurs on Forest Service land.

3. Many-stemmed live-forever ( Dudleya multicaulis ) occurs in
three places in and near the northermost Tecate cypress. Two small
populations are on thin, rocky soil, and one is on the west side of
a sloping meadow, scattered throughout the needlegrass ( Stipa spp.) .

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4 .

San Diego reed-grass ( Calamagrostis densa ) , occurs at its
northernmost location with the Tecate cypress. All of these rare
plants are candidates for Federal Endangered listing.

Four years after the 1982 Gypsum Canyon fire, young Tecate
cypress trees in the northernmost stand have grown past knee height.
This stand, covering 1,000 acres in Coal, Gypsum and Fremont canyons,
Orange County, remains in jeopardy.

A large "For Sale" sign, visible from the Riverside Freeway,
marks the entrance to the Coal Canyon property. This site, owned
by the Interpace Corporation of New Jersey, contains one-half the
area of the northernmost Tecate cypress and other rare plants in and
near the stand. The remainder of this rare plant community is owned
by the Irvine Company, which owns one-sixth of Orange County. None
of the Tecate cypress stands is on Forest Service land.

Orange County agencies are now preparing an Environmental Impact
Report ( EIR) for a major landfill in Coal or Gypsum canyon. Various
proposals for a jail, housing tracts, fuelbreak, post-fire grass
reseeding, roads and arterial highways also threaten the stand.

There is a ray of hope for the northernmost Tecate cypress and
its associated rare species. After Coal Canyon Shooting Sports lost
its option on the Coal Canyon property in early 1983, efforts were
intensified to contact public agencies as well as private conservation
foundations in order to save the northernmost stand, as either a state
or private nature preserve. As a result, the northernmost Tecate
cypress stand has been placed on the California Department of Parks
and Recreation's acquisition list. Private conservation organizations

could also be interested in the stand. In addition, a threatening
section 68 of a proposed Eastern Transportation Corridor was removed

from county plans.

You can help by writing to:

Senator William Campbell

1661 Hanover Road

City of Industry, CA 91748 #

encouraging him to work for the preservation of the Orange County
Tecate cypress and associated rare plants as a State Nature Preserve.

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Tecate cypres?

( Cupressus quadalupensis forbesii )

You can also help by being informed on the issues. A free slide
show about the northernmost Tecate cypress plant community for
botanical societies or community groups is available by calling Connie
Spenger (714) 879-3471 or Gordon Ruser (714) 541-0944.

SELECTED REFERENCES

Barbour, Michael G., and Majors. Terrestrial Vegetati on of C alifornia . _
Ch. 9, "The Closed-cone Pines and Cypresses. Wiley Inter science ,
New York, NY. 1977.

Dunn, Anthony T. A Preliminary Report on the Ecology and Ma nagement
of the Tecate Cypress in San Diego County. November, 1984.
(Unpublished) .

Wolf, Carl B. The New World Cypresses, El Aliso 1:1-444. 1948.

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A LIST OF HIGH ELEVATION ANGIOSPERMS AND THEIR

PHENOLOGY IN THE SAN GABRIEL MOUNTAINS,
LOS ANGELES COUNTY, CALIFORNIA

by

Wayne E. Sawyer
38 W. Sierra Madre Blvd., #8
Sierra Madre, CA 91024
(818) 355-4017

Introduction

In a small area between 7000 and 8400 feet (2300-2700 m) , on
a north facing slope in the San Gabriel Mountains, is an ideal location
for studying plant phenology of high elevation plants. The area is
accessible by trail, and within a short distance are five distinct
plant communities. Montane meadows are found at Lily and Little
Jimmy Springs. Most of the surrounding forest is moist yellow pine
forest characterized by Jeffrey pine, Pinus jef frey i; sugar pine,

Pinus lambertiana ; and white fir, Abies concolor . On dry slopes is
scrub vegetation dominated by chinquapin, Chrosolepis sempervirens ,
a community known as montane chaparral. On ridge tops is a forest
of lodgepole pine, Pinus murrayana and limber pine, Pinus f lexilis .

In exposed sites along the ridge is a wind blown edaphic community
of widely spaced low growing plants on coarse well-drained soil.

This is an edaphically controlled fell field.

Study Site

A study site was established immediately south of Highway 2,

2.1 miles east of the junction of Highway 2 and Route 39 in the San
Gabriel Mountains of southern California (see Figure 1) . It was
surveyed sixteen times during the spring and summer of 1980 to
identify all species of flowering plants. Plant specimens were
collected, pressed, and identified. These specimens are presently
on deposit at the herbarium of the Rancho Santa Ana Botanic Garden,
Claremont, California. Common insect visitors were collected and
later identified by Dr. Roy Snelling of the Los Angeles County
Museum of Natural History, Los Angeles, California.

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Flowering Phenology and Visitation

Plant populations accessible by hiking trails were observed
at least twice a week between May 2, 1981 and September 11, 1981.
Flowering status and floral visitors of each species were noted. A
subjective method was used to evaluate flowering status. Generally,
a species was considered to be flowering if? (1) at least a few
individuals each bore many open flowers, or (2) many individuals
each bore a few open flowers. Flowering phenology data are
summarized in Table 1.

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Table 1. Summary of flowering phenology data. The X's note on which days a
species was in bloom.

Date

May Jun Jul Aug Sep

Species 8 15 22 29 5 12 19 26 3 10 17 24 31 7 14 21 28 « 11

Alliuim burlewii ---X X-- - - - - - - - - - - -

Allium monticola - - - - - -- X X--- - - - - -

var . monticola

Aquilegia formosa - - - - - - X X XXXXX - - - - - -

var. truncata

Arabis platysperma - - - - - -- X X X X X - - - - -

Arabis repanda - - - - - - - - XX--- - - - - -

Arctostaphylos parryana XXX- - - - - - -- -- - - - - -

Argemone munita - - - - - - - - - X X X X X X X X X X

ssp. rotundata

Brickellia californica - - - - - - - - - -- -- -XXX X

Brickellia microphylla - -- - - - - - - -- -- - - X X X X

Calochortus invenustus - - - - - - - X X X X - - - -

Calyptridium monospermum - - -

Castilleja winiata - - -

Caulanthus amplexicaulus - -

Ceanothus cordulatus - - -

Cercocarpus ledifolius XXX
Chaenactis 6antolinoides - - -

Chrysolepis sempevirens -

- - X X - -- --

--- - XXXXX

XXXX - -- --

XXX - - -- --

- - X X XX---

--- - - -- XX

Chrysothamnus nauseosus ---- ____

var. bernardinua

Clematis liqustlcifolia ----

var. liqusticifolia

Collinaia torreyi ---X XXX-

var . wr iqhtii

Cryptantha muricata - - -XXX

var. denticulate

XXX

X

XXXX

XXXX

XXX-

X

X

X

X

Cycladenia humilis
var. venusta

X X

Delphinium glaucum

Oodecatheon reddens

Draba corrugata
var. corrugata

Draba stenoloba
var. nana

Epilobium brevistylum

Epilobium oregonense

Erigeron brewer i
var. jac i ^teus

Erigeron foliosus
var. stenophyllus

Er iodictyon tnchocalyx
var. tnchocalyx

XXX X X - -

XXXX XX

XXXX - -

-XXX - -

XXX XXX

X XXXXX - -- - - -

---XX XXXX - -

XXX XXXX X -

X XXX

7 -

Table 1. (cont.)

May

Jun

Jul

Aug Sep

7 14 21 28 4 11

Species

8 IS 22 29 5 12 19 26 3 10 17 24 31

Er iogonum davidsoni i

Er iogonum nudum
vat . pauci f lorum

Er iogonum pa r i s h i i

XXX XXXX XX

XXX XXXX XX

-XX XXXX x -

Er iogonum saxatile

XXX X X - - - -

Er iogonum umbel latum
var. minus

Eriogonum wrightii
var . subscaposum

XXXX - - - - - -

- - X X XXXX XX

Fntillaria pinetorum

Galium bifolium

Gayophytum sp.

Gilia ochroleuca
ssp. vivida

Gnaphaliua palustre

Habenaria leucostachys

Habenana sparslf lora

Helenium bigelovii

Heuchera elegans

Holodiscus microphyllus
var. microphyllus

Leptodactylon pungens
ssp. pulchrif lorum

Lewis l a nevadensis

Lewisia rediviva
var . minor

Lilium parryi

Lithophragaa tenellum

Lotus argophyllus
var. decorus

Lupinus elatus

Mimulus cardinalus

Mimulua f loribundus
var. f loribundus

Mimulus moschatus

Mimulus pilosus

Mimulus rubellus

Monardella cine tea

Oraonana vestita

Osmorhiza chi lensis

Oxytheca parishu
var. parishii

Pediculatis semibarbata

Penstemon bridgesi i

Penstemon grinnellii
ssp. grinnellii

Phacelia austrowontana

Phacelia curvipos
var. curvipes

Phacelia longipes

x x - -

-XXX
- - X X

X XXXX

- X

- X

X X

- - - - - XXX

- - - - - - X X

- - - - XXX-

X

X

X

X

-XX

X

- - - - X

x - -

-XX

- - X X

- - X

X

X

X

- - - - - - X X

---X XXX-

- - - - - - X X

- - - - - XXX

x--- - - - - - - -

xxxxx XXXX - -

x - - - - - - - - * ■

---XX XXXX XX
XXXXX - -- - - -

xxxxx - - - - - -

-XXXX XXXX - -

XX--- - - - - --

- XXX- - - - - - -

XX--- - - - - - -

-XXXX - - - - - -

XX--- - -- - - -

XX--- - -- - - -

xxxxx X-- - - -

XXXXX XXXX - -

- - X X X XXXX XX

xxxxx

- - X X X

- XXX-

- - X X X

X x - -

XXXX

XXXX

xx-- - - -- - - -

xxxxx XXXX XX

xxxxx X---

XX--- - -- -

X X

8 -

Table 1 . (cone.)

May

Jun

Jul

Aug Sep

7 14 21 28 4 11

Species

Polygonum aviculare

Potentilla alandulosa
ssp. nevaaensis

Potentilla qlandulosa
ssp. ref lexa

Pterospora andromedea

Pyrola picta

Rhamnus californica
ssp. cuspidate

Pibes c-.reum

Pibes nevadense

Pibes roezlii

Pubus leucodermis
var. bernardinus

15 22 29 5 12 19 26 3 10 17 24 31

XXX XXXX XX

- X X X XX

- X X X

- - - - - - - - - - X X X XXX- - -

- - - - - - - - -xxxx - - - - -

--- - - - X X XX-- - - - - - - -

-XXX XXX- - -- -- - - - - - -

-XXX XXX- - -- -- - - - - - -

-XXX XXX- - -- -- - - - - - -

--- - - - X X XX-- - - - - -

Salix lasiolepis XXX - - -- - - -- -- - - - -

var. lasiolepTs

Salsola iberica --- - - - - - - - X XXXX

Sambucus caerulea - - - - - - - X X X X X X X - - - -

Sar codes sanguines - - - - - - X X - -- -- - - - - -

Senecio ionophyllus

Silene parishii
var. latifolia

Silene verecunda
var . platyota

Sisyrinchium eastwoodiae

Sailacina stellata

XXX

XXX XXX-

XXX

X

X - - -

XX XXXXX XXXX XX

XX XXXXX XXX - - -

Solidaqo californica - - - - - - - - - -- -- XXXX XX

Symphoricarpos parishii - - - - - - - - X X X - - - - - - -

Tetradymia canescens - - - - - - - - - - - - X XXX- -

Tri folium monanthua - - - - - XXX XXXXX X-- - - -

var. grantianua

Veratrum calif ornicum - - - - - - - - XXXX- - -- _ -

Viola purpurea - - - - -XX- - -- -- - - - - -

ssp. xerophyta

Zauschneria californica - - - - - - - - - XXXX XXXX XX

Discussion

All but 5 of the 100 species believed to be biotically-
pollinated are listed. No phenology data were collected for
Calyptridium parryi var. parryi as no individuals of this species
were observed in 1981. Flowering phenology of Coral lorhiza maculata ,
Miroulus johnstonii , Penstemon caesius , and Sisymbrium altissimum was
not followed because these species were already in fruit when first
observed during the summer of 1981. Failure to find these species
before they had completed flowering was due to their rarity in the
study site.

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Most species had a relatively long flowering period. Only 21
percent had a flowering period of three weeks or less and the average
flowering period was 5.4 weeks.

Just one species had a disjunct flowering period. Purple-
flowered Mimulus rubellus , a small annual, was in flower from 6/5 to
6/12. A yellow-f lowered race of this same species (Thorne, personal
communication) was bloom from 7/10 to 7/24.

ANNOUNCEMENTS

Annual Southern California Botanist*' Symposium

Tha Board of Directors and Officers were able to begin preparations
for the 13th annual one-day symposium almost a year ahead. The
theme was suggested on the questionnaires collected at the 1986
symposium and was approved at the December meeting of the Board.

The theme and title for the 1987 symposium will be "Botanical
Discoveries and Exploration in Southern Cal i f orni a" . Taxonomists,
plant physiologists, ecologists, gardens, museums, etc. of Southern
California have made many exciting discoveries and advanced the
understanding and appreciation of vegetation communities and flora.
Many principles of plant science have been developed based on theories
originally conceived in Southern California.

Some of the suggested topics for individual speakers were:

— the anecdotes of a famous botanist who explored this region
throughout his/her career as told by that botanist

— the biography of a famous botanist who explored this region
as told by a historian

— the history of botanical exploration in southern California

— an overview of the scientific discoveries made in this area

— the current status of botanical research (who, where, what)
supported in this region

— a history of the creation of a particular botanic garden, e.g..
Rancho Santa Ana Botanic Garden, and its future plans

— the development of scientific inquiry on and knowledge about
a prevalent plant community type in Southern California

— the history of the rare plant conservation effort in this region
and its current status

— the history of the development of a theory or principle and the
contribution made by Southern California scientists

Some names were suggested for speakers; however, we would like to
get the input of the SCB members. If you have a speaker you would
like to hear or another i nterpretat ion of the 19S7 theme, please
contact anyone of the Officers or Board of Directors or call Mona
Myatt at BIB 302-1466 (weekdays) or BIB 447-075S (evenings and
weekends) before March 15, 1987.

The symposium will be held at the same place as last year, i.e. ,
the Heritage Room of the University Center at the University of
California, Irvine. The date is Saturday, November 14, 1987. The
cost is always reasonable. There will be free refreshments and
book and biota-themed clothes for sale. The 1986 symposium was
very informative and enjoyable. Set aside November 14 and tell your
friends and botanical associates. Hope to see you there.

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Fourth Annual Environmental Legislative Symposium. Jan. 31, Feb. 1,
1987. The Planning and Conservation League will hold their 4th annual
symposium on Saturday, January 31 and February 1, 1987. Attendees
will be informed of major issues that will be before the legislature
in 1987. There will be workshops on today's leading environmental
issues. For more information call: Anne Hedges, Symposium Coordinator,

(916) 444-8726 or write to: PCL Symposium, 909 12th St., #203,

Sacramento, CA 95814.

Friends of the Big Bear Valley Preserve . Bald eagles over-winter at
Big Bear Lake. Throughout winter months, eagle tours are conducted
nearly every weekend. On Sunday, February 8, the Orange County Chapter
of CNPS has arranged for a tour. Reservations are required and the
fee is $5.00 unless you are a member of the "Friends." Arrangements
also can be made to visit the Nature Conservancy's North Baldwin Lake
Preserve where rare and endangered species occupy an interesting
habitat, a pavement plain. For information about the CNPS trip, call
Nancy Kenyon (714) 786-3160. For information about the "Friends,"
call Lauren Pollock (714) 585-9266.

Help Needed for a "Weeding Party" at Irvine Park. Recent rains have
encouraged growth of the new plants at the Irvine Park revegetation
site. There are also lots of healthy weeds. A party to remove
pesky weeds has been scheduled for Saturday , February 1^ at 8 :00 a.m.

For information call Gus or Frieda (714) 974-1639.

Third California Islands Symposium. March 2-6, 1987. The Southern
California Academy of Sciences, the Santa Barbara Museum of Natural
History and the Santa Barbara Botanic Garden are hosting this conference
focusing exclusively on research on the islands off the coasts of
California and Baja California, Mexico. The meeting will be held in
Santa Barbara. Pre-registration is $40 until January 15, 1987; regular
registration is $55. Registration forms are available from California
Islands Symposium, Santa Barbara Museum of Natural History, 2559 Puesta
del Sol Road, Santa Barbara, CA 93105.

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Symposium on the Scientific Value of the Desert. April 5, 1987.

The symposium, to be held at the Manchester Conference Center of the
University of San Diego, will be hosted by the Anza-Borrego Foundation,
to celebrate its 20th anniversary, and the Environmental Studies
Program of the University of San Diego. Papers will deal with
anthropology, archaeology, biology, ecology, geology, paleontology,
and wildlife management. Applications and abstracts of proposed
papers should be sent by January 1, 1987 to Dr. James Richard,

P. O. Box 777, Borrego Springs, CA 92004. The $15.00 registration/
luncheon fee will be collected at the symposium. For more information,
call Dr. James Rickard, (619) 767-3776, or Dr. Richard Phillips
(619) 260-4600, ext. 4451.

Second Native Plant Revegetation Symposium. April 15-18,

Hanalei Hotel, San Diego. Riparian Systems will be featured in a
special section of the Second Native Plant Revegetation Symposium,
co-sponsored by the Southern California Chapter of the Wildlife
Society (WS), the San Diego Chapter of CNPS, and the Riparian
Revegetation Work Study Group ( RRWSG) . The goal is to exchange
information on current research, and new or refined techniques and
projects using native plants in revegetation and landscaping. There
will also be a poster and commercial exhibit. For more information,
write to: Native Plant Revegetation Symposium, 3808 Rosecrans St.,

#373, San Diego, CA 92110; or call John Rieger (TWS) , (619) 237-6754;
Bobbie Steele (CNPS), (619) 175-3732; or Anne Sands (RRWSG) ,( 415) 381-2629 .

Mary D eDecker Symposium. From Thursday, April 30 through Sunday,

May 3. The University of California White Mountain Research Station
with the California Native Plant Society and its Bristlecone Chapter
is co-sponsoring a symposium and field trip honoring Mary DeDecker.

The topic of the symposium is the flora and plant biology of the
eastern Sierra, Owens Valley, White-Inyo Mountains, and western Basin
and Range Provinces. The field trip to the Eureka Dunes on Saturday,

May 2, will be led by Mary DeDecker and others; a BBQ cookout will
follow. Registration for the conference is $10 per person, and
dormitory facilities are available for $1.50 per night. Motels are
available in Bishop. Those who wish to attend all or part of the
symposium or to present a paper should contact Bart O'Brien,

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(408) 741-9534 evenings, (415) 941-7705 days. Registration and
abstracts should be sent to Dr. Clarence A. Hall, Jr., White Mountain
Research Station, 6713 Geology Building, University of California,

Los Angeles, CA 90024; phone; (213) 825-2093.

Third Annual Southwestern Botanical Systematics Symposium: Advances
in Plant Systematics and Ecology. Rancho Santa Ana Botanic Garden
is presenting the third in its series of annual symposia aimed at
systematic botanists in the southwestern United States and Mexico.

This year*s roster of speakers will present original research papers
on a wide range of subjects concerning plant systematics and ecology.

An informal social for all participants will be held in the home
demonstration garden on Friday evening. May 22 . Invited papers
will be presented on Saturday, May 23, in the auditorium of the
main administration building. Library and herbarium facilities will
be available during the meeting, and low-cost housing will be
available locally. Detailed information on housing will be sent
with confirmation of registration. Please register early, as space
is limited to 100. For information or reservations contact:

Rancho Santa Ana Botanic Garden, Botanical Systematics Symposium,

1500 North College Avenue, Claremont, CA 91711? phone: (714) 625-8767.
Fund for Environmental Defense (F.E.D.). A new legal defense fund,
patterned after the Sierra Club Legal Defense Fund, has been established
for Orange County. F.E.D. has been established for the purpose of
providing a legal defense fund to which environmental activist groups
can apply for assistance in maintaining or increasing the quality of
life in Orange County. Specific objectives are to advise and assist
in litigation proceedings by environmental groups that seek to create,
maintain or increase the aesthetics of their surroundings, such as
(but not limited to) residential areas, parks, greenbelts or other
open space areas. Memberships ranging from $25.00 to $1000.00 are
available. For more information write to: Fund for Environmental

Defense, P. O. Box 9118, Fountain Valley, CA 92708 or call Charlotte
Clark (714) 871-8000, Ext. 263.

******

-13-

FIELD TRIPS

January 31 ( Saturday ) . Myco logical Forays. The Los Angeles
Mycological Society in conjunction with their Mycological Faire (see
below) is hosting several field trips in Los Angeles and Orange
Counties. Two mushroom collecting trips are planned for Orange
County:

1. Rancho Mission Viejo. Follow Ortega Highway east about
5 miles from 1-5. Meet at the yellow gate on Cristianitas Road.

This road intersects Ortega Highway on the south before you reach
Casper's Park.

2. Blue Jay Campground. Take Ortega Highway east from 1-5.

Meet at the first turnoff to the Blue Jay Campground.

For more details, including meeting times, call (213) 292-1900.

February 1 ( Sunday ) . Mycological Faire . The Mycological Society
(see above) with host identification classes, cooking demonstrations,
and lectures about edible and poisonous mushrooms. Meetings will
be held at the Orange County Natural History Center, 2627 Vista Del
Oro, Newport Beach. For details, including times, call Dave Bramlet
(714) 549-0647.

February 22 ( Sunday ) . Rare Plants of the Salton Basin. 8:00 a.m.
John Stewart of the Living Desert Reserve will lead this Riverside
CNPS field trip. The trip will search for Pi taxis adenophora ,

D. californicum , Machaeranthera capoda , M. orcuttii and Pilostyles
thurberi . Take I-10E past Palm Springs to the Ramona/Bob Hope Dr.
exit; take Bob Hope Drive south to Palm Desert Hwy. 111. Go 2-3 miles
to Portola Ave. and turn right on Toitoia. Go about 1-1/2 miles to
the Living Desert Reserve. We will meet in the parking lot. Call
Andy Sanders (714) 787-3601 for details.

February 28 ( Saturday ) . 9:30 a.m . Rancho Mission Viejo.

Walt Wright will lead this mycological foray to search for mushrooms
in oak woodland, native grassland, and coastal sage scrub. Take
Ortega Highway east from 1-5 for about 5 miles to Cristianitos Road.
Meet at the yellow gate. For details call Walt Wright (714) 529-4134.

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March 8^ ( Sunday ) . 9:00 a.m. San Joaquin Hills . Fred Roberts

of U. C. I. will lead this trip to examine unusual coastal sage
scrub near Laguna Beach. Plants to look for include crown-beard,
Verbesina dissita ; adder ' s-tongue fern, Ophioglossum californicum;
shooting star, Dodecatheon clevelandii , and other interesting early-
blooming species. Take Pacific Coast Highway south through Laguna
Beach. Meet at parking lot on inland side of the highway across from
the Treasure Island Trailer Park. For more details call Dave Bramlet
(714) 549-0647.

March 14, 15 (Saturday-Sunday) . 9:00 a.m. Northern Baja California .

Dr. Thorne of the Rancho Santa Ana Botanical Garden will lead this
trip down Mex. 3 to Punta Bunda. We will see Tecate cypress, Englemann
oak and many other interesting plant species. We may also go south
of Maneadero to look for Fremontodendron mexicana . Meet in Tecate
(USA) at the shopping center parking xo c, just Lc^ore the border.
Bring camping gear, pesos, Mexican auto insurance and a birth
certificate or passport. For more information, please send an SASE
to David Bramlet, 1691 Mesa Dr., A-2, Santa Ana, CA 92707 or call
at (714) 549-0647.

March 21, 22 (Saturday-Sunday). 10:00 a.m. Joshua Tree National
Monument. Andy Sanders, Rare Plant Coordinator of R/SB Chap, and
Curator of U. C. Riverside Herbarium will lead this trip in southern
California desert region to enjoy magnificent spring display. Meet
10 a.m. Sat. at Cottonwood Springs Visitors' Center. Take 1-10,

26 miles east of Indio, go north 7 miles on Cottonwood Canyon Road.
Camp overnight at Sheep Pass Group Camp; Bring water. Motels
available at Twenty-Nine Palms; Budget Motel, (719)367-2833; Gardens
Motel (Best Western) , (619) 367-9141; and others. Trip ends 3 p.m.
Sunday. Reserve deadline : 8 March. Camping fee, $5 per vehicle
(additional $5 if arriving Friday) . Check payable to CNPS. Enclose
SASE; send to Lucien Haas, 638 Swathmore, Pacific Palisades 90272;
(213) 454-2361. Alternate: Andy Sanders, 422 Campus View, Riverside,
Ca 92507; (714) 684-0448.

- 15 -

March 28 (Saturday). 8:00 a.m. to dusk. Pines to Palms Trip.

Geoff Smith will lead this trip to examine desert, chaparral and
mountain habitats. Meet at the S. E. corner of the Ramona Expressway
and 215 south of Riverside. Caravan from Hemet through the San
Jacinto Mountains on Hwy 74 with stops at plant communities discussed
in Geoff's recent Crossosoma article (April, 1986). Bring a sack
lunch and comfortable walking shoes. For further information contact
Geoff at (714) 526-6963 (evenings); (714)929-5248 (evenings) or
(714)871-8000, ext. 371.

April 4 (Saturday). 9; 00 a.m. Harford Springs Park. Steve Boyd of
the Rancho Santa Ana Botanical Garden will lead this trip to see the
unique flora of the Gavilan Hills. We will find excellent stands of
Juniper Woodland and will look for Munz's Onion, Paysen's fclanthus ,

F allbrook spinef lower and many other disjunct species. Take the 91
freeway east to the 15 and go south. Take the turnoff at Cajalco Rd.
and go east to Gavilan Rd. At Gavilan go south to Ida Leona Rd.

(1st left past the trailer park) and meet at the top of the rise on
Ida Leona. Bring a sack lunch and water.

April 25, 26 (Friday - Sunday ) . Desert Studies Center at Soda Springs
(Zzyzx) . Trip limited to 25 individuals. A cost of approximately
$8 per person for use of the facility will be charged. Reservations
or information about the trip can be obtained from Alan Romspert by
calling (714) 773-2428. We will see a variety of desert wildflowers
and shrubs in bloom. This looks like it may be an excellent year
for a spectacular wildf lower display in the Mojave Desert.

May 2 (Saturday). 9:00 a.m. Chiquito Basin. Fred Roberts of the UCI
Systematic Museum will lead this trip to the unique deer grass portero
found at Chiquito basin in the Santa Ana mountains. We will see large
areas of chaparral, oak woodland and native grassland. This area
also contains the largest population of the San Miguel Savory
( Sature ja chandleri ) , which should be in flower. Bring a lunch, sun
protection, water and a hand lens. We will meet at the interseqtion
of the Ortega Highway (74) and Forest Service road #6S05 that leads
to the conservation camp. For more information call Dave Bramlet
(714)549-0647.

- 16 -

May 10 (Sunday). 10:00 a.m. Pine Canyon. Karlin Marsh will lead
this trip into the Santa Ana Mountains. Pine Canyon is a north-facing
slope tributary of Silverado Canyon. The area is a highly shaded
mixed hardwood, conifer forest with many ferns and wildf lowers. Meet
at 10:00 a.m. at the end of Silverado Canyon Road at the Forest gate.
Parking is very limited so please carpool. A ranger may let us
through to park inside the gate. If not and the lot is full, then
park down the canyon 1/4 mile, near the bridge. For more information,
call Dave or Karlin at (714)649-0247.

SOUTHERN CALIFORNIA BOTANISTS SPRING CAN OE TRIP

Once again we will ba gathering at the Colorado Ri var for a spring
canoa trip. Tha trip will ba hald on tha weekend of April 11 and 12.
This year, however, wa will not ba returning to tha Topock Gorge
section of tha River, but will ba canoeing on a different stretch of
the river south of Blythe and north of Yuma. It is also possible that
this trip will be extended to three days.

All arrangements are not yet complete and the layout of the area
requires either a complex car shuttle or the use of a bus to return
us all to our starting point. The cost per person will hopefully
still remain under $50 even for a 3-day trip. Total costs and the
logistics of various put-in and take-out points on the river as well
as camping locations are currently being confirmed.

If you are interested and would like to reserve your canoe (there will
probably be a limit of 11 canoes, unless response is ex traordi nary ) ,
call Mona Myatt at (018) 302-1466 weekdays or (810) 447-0755 weekends
and evenings. You will be sent a complete package of instructions
and a list of participants and things to bring. This trip will be a
f irst-to-cal 1-gets-conf ir med -reserve ti on s type of arrangement. Also
you will be expected to pay in full to hold your reservations. The
trip has been very popular and a lot of fun, so signup as soon as
possible. Reservations (not payment) will be needed be fore
March 2, 1987.

This area of the Colorado River and desert holds many interesting
biologic, scenic and historic sites. As part of your instruction
package you will be sent information on places to stop at traveling
to and from the River. You might even want to set aside a day or
two for additional exploration of this area.

Hope to see some of the veterans from canoe trips of past years as well
as lots of other 6CB members, maybe some of you who have never been
on any of our botanical trips. This trip has it all — adventure,
scenery, wildlife, history, neat plants, sunny skies, and good
companionship. Be there! !!!!!!!!!!!!!

P.S. Mona Myatt is organizing a Whitewater trip in Canyonlands or
the Grand Canyon as well as dog-sledding in Canada. If these trips
tickle your fancy, call at the above numbers.

- 17 -

The following article on George Engelmann is reprinted from the
Spring, 1986 (Vol. 2, No. 3) issue of the Tenaja Telegraph. This is
the journal of the Santa Rosa Plateau Preserve where groves of
Engelmann oak, Quercus engelmannii are a featured attraction.

ROOT OF THE ENGELMANN OAK
By Isabel P. Ludlum

While on docent station along Adobe Trail last September during
Santa Rosa Plateau Preserve’s hosting of the California Chapter of
TNC , I was asked numerous times not only about the oak named
Engelmann, but about the man himself. My knowledge was sketchy, but
through subsequent researching I have made the acquaintance of an
interesting and important pioneer botanist. For those of you who are
unacquainted, let me introduce George Engelmann, M. D.

Dr. Engelmann was born February 2, 1809 in Frankfort-on-the-Main ,
Germany and died February 4, 1884 in St. Louis, Missouri. He was the
eldest of thirteen children born of parents who were educators. Even
in his teens he showed an interest in botany and exhibited a scientific
mind. He studied at Heidelberg and Berlin Universities and took the
M. D. degree at the University of Wurtzberg in 1831. The following
year he went to Paris for medical and scientific studies and counted
as his friends there Louis Agassiz and Alexander Braun.

When his uncles became interested in land investments in the
Mississippi Valley, young Dr. Englemann became their willing agent,
sailing to the United States in 1832. Botanical explorations, usually
by horseback through Southern Illinois, Missouri and Arkansas, consumed
much of his time and limited monies.

By 1835 he was impoverished and at that time established himself
in the practice of medicine in St. Louis, laying the foundation for
the successes in his profession. In four years' time he was
prosperous enough to return to Germany and marry his fiancee, a cousin.

During the latter part of his life he was able to explore his
adopted country and in his early seventies made a long journey through
the forests of the pacific State?, where he finally observed "in the
state of nature" plants he had studied and described. Summing up his
seventy-five years of accomplishments. Dr. Engelmann, a recognized
authority on botany at home and abroad, frequently took extended
periods of time out for his avocation — botanical investigations?

- 18 -

he maintained an herbarium and botanical library adjoining his office;
he was among the first to use quinine in treating malaria; he was
much occupied with meterology, and he was the organizer of the St.
Louis Academy of Science. He also made important contributions in
the study of grapevines, the cactus family and the cuscutaceae
(dodder) , but outstanding are his various papers on the American oaks
and the conifers.

Because of his research and authorship, his name is associated
with the Buffalo-grass of the plains, the Engelmann spruce ( Picea
engelmannii ) seen all over the Rockies and in spots in the Cascades,
and with our own Engelmann Oak ( Quercus engelmannii ) .

BOARD OF DIRECTORS

c OUTHERN CALIFORNIA

OFFICERS

Geoff B. Smith (President)

28209 Vista del Valle
Hemet, CA 92344
w(714) 871-8000, ext 371
h (714) 929-5248 or
(714) 526-6963

David Bramlet (First Vice-President)
1691 Mesa Drive, Apt. A-2
Santa Ana, CA 92707
w ( 714 ) 855-02222
h ( 714 ) 549-0674

.Leo Song (Second Vice-President)
Dept, of Biological Science
Calif. State University
Fullerton, CA 92634
w(714) 773-2766
h(714) 528-6316

Alan P. Romspert (Treasurer)

605 N. Pomona Ave.

Fullerton, CA 92632
w(714) 773-2428
h(714) 870-0946

Marybeth Roehm (Secretary)

6300 Stephens Ranch Road
La Verne, CA 91750
w —
h —

Mona Myatt (Past President)

6421 N. Golden West Ave
Temple City, CA 91780
w ( 818) 302-1466
h (818) 447-0755

Allan Schoenherr (Editor,

(Crossosoma)

Biological Sciences, Fullerton
College

321 Eo Chapman Ave.

Fullerton, CA 92634
w(714) 871-8000, ext. 263
h (714) 494-0675

BOTANISTS - 1987

DIRECTORS

Katie Barrows
53277 Diaz
La Quinta, CA 92253
w —

h (619) 564-2413

Coleen Cory
301 N. Baldwin Avenue
Arcadia, CA 91006
w( 818) 446-8251
h —

Jon E. Keeley
Dept, of Biology
Occidental College
Los Angeles, CA 90041
w ( 213) 259-2898
h —

Karen Kirtland
285 Tangelo
Irvine, CA 91714
w(714) 640-6363
h —

Kathleen Pahl
240 Loma St.

Long Beach, CA 90803
w ( 818) 302-4519
h (213) 439-7301

Barry Prigge

Herbarium & Botanic Garden
Univ. of Calif., Los Angeles
Los Angeles, CA 90024
w (213) 413-1117
h —

Sherry Schmidt
P. O. Box 771
Silverado, CA 92676
w (714) 594-5611, ext. 764
h (714) 649-2283

Rick Strom
538 Brookshire
Orange, CA 92667
w —

h (71 4) 637-8070

ne maintained an Herbarium and botanical library adjoining his office;
he was among the first to use quinine in treating malaria; he was
much occupied with meterology, and he was the organizer of the St.
Louis Academy of Science. He also made important contributions in
the study of grapevines, the cactus family and the cuscutaceae
(dodder) , but outstanding are his various papers on the American oaks
and the conifers.

Because of his research and authorship, his name is associated
with the Buffalo-grass of the plains, the Engelmann spruce (Picea
engelmannii ) seen all over the Rockies and in spots in the Cascades,
and with our own Engelmann Oak (Quercus engelmannii ) .

BOARD OF DIRECTORS

c OUTHERN CALIFORNIA

OFFICERS

Geoff B. Smith (President)

28209 Vista del Valle
Hemet, CA 92344
w(714) 871-8000, ext 371
h(714) 929-5248 or
(714) 526-6963

David Bramlet (First Vice-President)
1691 Mesa Drive, Apt. A-2
Santa Ana, CA 92707
w(714) 855-02222
h (714) 549-0674

.Leo Song (Second Vice-President)
Dept, of Biological Science
Calif. State University
Fullerton, CA 92634
w(714) 773-2766
h (714 ) 528-6316

Alan P. Romspert (Treasurer)

605 N. Pomona Ave.

Fullerton, CA 92632
w (714) 773-2428
h(714) 870-0946

Marybeth Roehm (Secretary)

6300 Stephens Ranch Road
La Verne, CA 91750
w —
h —

Mona Myatt (Past President)

6421 N. Golden West Ave
Temple City, CA 91780
w(818) 302-1466
h ( 818) 447-0755

Allan Schoenherr (Editor,

(Crossosoma)

Biological Sciences, Fullerton
College

321 E. Chapman Ave.

Fullerton, CA 92634
w(714) 871-8000, ext. 263
h (714) 494-0675

BOTANISTS - 1987

DIRECTORS

Katie Barrows
53277 Diaz
La Quinta, CA 92253
w —

h (619) 564-2413

Coleen Cory
301 N. Baldwin Avenue
Arcadia, CA 91006
w (818) 446-8251
h —

Jon E. Keeley
Dept, of Biology
Occidental College
Los Angeles, CA 90041
w(213) 259-2898
h —

Karen Kirtland
285 Tange lo
Irvine, CA 91714
w ( 714 ) 640-6363
h —

Kathleen Pahl
240 Loma St.

Long Beach, CA 90803
w (818) 302-4519
h (213) 439-7301

Barry Prigge

Herbarium & Botanic Garden
Univ. of Calif., Los Angeles
Los Angeles, CA 90024
w ( 213) 413-1117
h —

Sherry Schmidt
P. 0. Box 771
Silverado, CA 92676
w (714) 594-5611, ext. 764
h (714) 649-2283

Rick Strom
538 Brookshire
Orange, CA 92667
w — •

h f 71 41 637-8020

SOUTHERN
CALIFORNIA

13TH ANNUAL SYMPOSIUM BOTANISTS

COSPONSORED BY THE DEPT. OF ECOLOGY
AND EVOLUTIONARY BIOLOGY AND PETER
BOWLER, DIRECTOR OF THE COOPERATIVE
OUTDOOR PROGRAM IN THE STUDENT
ACTIVITIES OFFICE AT UC, IRVINE

BOTANICAL DISCOVERIES
AND EXPLORATION IN
SOUTHERN CALIFORNIA

date: SATURDAY, NOVEMBER 14, 1987

time: 8:45 AM TO 4:00 PM

place: HERITAGE ROOM, UNIVERSITY
CENTER: U.C., IRVINE

IF YOU HAVE A SUGGESTION FOR ONE OR MORE OF THE FIVE SPEAKERS,
PLEASE CONTACT ONE OF THE S.C.B. OFFICERS OR DIRECTORS.

VOUFfc

- 20 -

Two revised floras from the
Southern California Botanists

A FLORA OF THE SANTA ROSA PLATEAU ,
SOUTHERN CALIFORNIA. By Earl W.
Lathrop and Robert F. Thorne. 39
pages; paperback; coab binding;

FLORA OF THE SANTA MONICA
MOUNTAINS, CALIFORNIA.

By Peter H. Raven, Henry J.
Thompson, and Barry A. Prigge.
179 pagea; paperback; sayth
aevn binding; $10.50

Please send :

Price

copies o

f A FLORA OF THE SANTA ROSA PLATEAU $ $7.00

copies of FLORA OF THE SANTA MONICA MOUNTAINS • $10.50

Return to: So. Calif Botanists
Dept, of Biology
Calif. State University
Fullerton, CA 92634

Total

Price includes tax, handling,
and postage.

Make check or money order payable to: Southern California Botanists

- 21 -

SOUTHERN CALIFORNIA BOTANISTS
MEMBERSHIP/TREASURER SUMMARY REPORT
1986

Members^ following statement summarizes the monetary transactions for 1986. We closed
the 1986 year with 338 members, down from the approximately 400 members in 1985,
Financially we were in better shape at the end of 1986 in comparison to 1985, but we
still need emphasis on the recruitment of new members. Please pass on the enclosed
membership form to someone who might be interested in our organization.

CHECKING ACCOUNT

BALANCE 12-31-85

$162.96

RECEIPTS

DISPERSEMENTS

Interest

76.17

Mailing

253.67

Membership

3,023.00

Printing

1,338.81

Booksales

137.43

Postage

129.20

Plant Sale

1,722.75

Symposium

955.19

Symposium

689.00

Plant Sale

1,207.10

Donations

74.00

Student Grants

300,00

Field Trips

2.22

Miscellaneous

169.34

Miscellaneous

219.58

TOTAL +$5,944.15

TOTAL

$4,353.31

BALANCE 12-31-86

MONEY

MARKET ACCOUNT

$1,753.80

BALANCE 12-31-85

$6,368.19

RECEIPTS

DISPERSEMENTS

Interest

Santa Rosa
Plateau Flora

268.45

405.34

Printing
(Santa Monica

At! 8 ih2 5 ra)

Santa Monica
Mountain Flora

1,084.04

San Diego Co.
Flora

294.50

Calendars

176.70

Shipping &
Handling

44.78

Petty Cash

35.75

Tax

56.81

TOTAL

+$2,366.37

TOTAL

-$4,186.45

BALANCE 12-31-86 $4,548.11

Future Field Trips. In upcoming issues of CROSSOSOMA look for details
on the followings

/

May 23-25. Clark Mountain
June 6. San Mateo Marsh

June 20. Tetracoccus Ridge, Panamint Mountains.

- 22 -

SOUTHERN <_ A l I FOR Nl A BOTANISTS

Rancho Santa Ana Botanic Garden
1500 North College Avenue
Claremont. CA *>|7II

The purpose of the SOUTHERN CALIFORNIA BOTANISTS Is the study,
preservation in«. conservation of the native plants of California; and
the education of the public to tne value of the native and its

habitats. It is a non-profit association forced in 1927.

Membership benefits include:

Various field trips throughout the state led by competent

field botanists and biologists.

A yearly plant sale featuring native California species.

An annual symposium on various aspects of the California

vegetat ion .

The SCB Journal, CROSSOSOMA .

Discounts on botanical and natural history books.

Dues are for a calendar year. New members Joining from May through
September, please deduct $1.00 from your dues. Those Joining in October
through December are credited with the following year's dues.

Membership categories are:

/ / Individual* $ 8*00

/ / Croup or organization $15.00

^Includes membership for entire family.

Name

Address
City

Phon e_j } _

In addition. I want to give S_ help «wpP° rt SCB -

Make check payable to: SOUTHERN CALIFORNIA BOTANISTS.

Mall check and form to:

Alan P. Romspert
Southern California Botanists
Department of Biological Science
California State University, Fullerton
Fullerton, CA 92634

CROSSOSOMA is published bimonthly (February, April, June,
August, October, and December) by Southern California Botanists,
a non-profit association.

Zip Code

/ / New Member

/ / Renewal

Date

- 23 -

SCB COMING EVENTS ( DETAILS WITHIN )

January 31
February 1
February 22
February 28
March 8
March 14, 15
March 21, 22
March 28
April 4
April 11, 12
April 25,26
May 2
May 10

Mycological Forays

Mycological Faire

Rare plants of the Salton Basin

Rancho Mission Viejo

San Joaquin Hills

Northern Baja California

Joshua Tree National Monument

Pines to Palms

Harford Springs Park

Colorado River canoe trip

Soda Springs (Zzyzx)

Chiquito Basin

Pine Canyon, Santa Ana Mountains

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CROSSOSOMA

DBRAPV

SOUTHERN CALIFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden, Claremont, CA 91711

MAY 26 m

CROSSOSOMA Vol . 13, No. 2
Managing Editor: Allan Schoenherr

ApfMl t'v^87 ^

iQTANICAL GARDjE$

ADDITIONAL NOMENCLATURAL CHANGES AND NOTES
IN A FLORA OF SOUTHERN CALIFORNIA, PA. Munz, 1974

Fred M. Roberts, Jr.

Museum of Systematic Biology
University of California, Irvine

In a previous paper (Roberts, 1986) I presented a summary of
revisions and additions to the flora of southern California nomen-
clature occur ing since the publication of Munz (1974). Continued
investigation has revealed further changes in the nomenclature. The
results of these investigations are presented below.

As in Roberts (op. cit.), new additions are listed only if they
clarify existing taxonomic problems. Each entry is listed in order as
they appear in Munz (1974), followed by the newly accepted or proposed
name and finally the source of the revision.

ASPIDACEAE

Athyrium alpestre (Hoppe) Ry lands var. cyclosorum (Ledeb.) Moore *

Athyrium filix-femina (L.) Roth. ssp. cyclosorum (Rupr.) Chr.
Recognized by Kartesz & Kartesz, 1980, and Lellinger, 1985.
Apparently originates with F.S. Liew, Numerical Taxonimic Studies
of North American Lady Ferns and their Allies, Taiwania 17: 190-
221, 1972.

ASPIDACEAE (continued)

Athyrium filix-feminia (L.) Roth var. californicum Butters, iricl. in
Athyrium f il ix-femina (L.) Roth, ssp. cyclosorum (Rupr.) C. Chr.
SEE reference under Athyrium alpestre.

Athyrium filix-feminia var. sitchense Rupr. included in:

Athyrium fil ix-femina (L.) Roth, ssp. cyclosorum (Rupr.) C. Chr.
SEE reference under Athyrium alpestre.

PTERIDACEAE

Chei lanthes parishii Davenp.

OMIT from Flora, a hybrid. SEE A.R. Smith, Madrono 22: 376-
378, 1974a.

Notholaena sinuata (Sw.) Kaulf. var. cochi sens is (Goodd.) Weath.
Notholaena cochisensis Goodd.

Recognized by Kartesz & Kartesz, 1980 and supported by Lei linger,
1985.

OJPRESSACEAE

Cupressus stephensonii C.B. Wolf 3

Cupressua arizonica Greene ssp. stephensonii (C.B. Wolf)

Beauchamp

R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of
San Diego County, California. Phytologica 59: 437-440, 1986.
(supercedes listing in Roberts, Crosossoma 12: 3, 1986.)

APIACEAE

Caucalis microcarpa H. & A.

Kartesz & Kartesz, 1980 recognizes this taxon under the name
Yabea microcarpa (Hook. & Arn.) K.-Pol. I have not seen the
reference for this change but its usage is becoming more popular.
Beauchamp (1986) recognizes this name and it is presently being
applied at the Rancho Santa Ana Botanic Gardens.

ASTERACEAE

Haplopappus acradenius (Greene) Blake ssp. eremophilus (Greene) Hall =
Isocoma acradenia Greene ssp. eremophila (Greene) Beauchamp
R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of
San Diego County, California. Phytologica 59: 437-440, 1986. (see
also remarks in Roberts, Crosossoma 12: 3, 1986.)

- 2 -

ASTERACEAE (continued)

Haplopappus venetus (H.B.K.) Blake ssp. furfuraceus (Greene) Hall =
Isocoma veneta (H.B.K.) Greene var. furfuraceus (Greene)

Beauchamp

SEE reference under Haplopappus acradenius above. Interestingly,
Beauchamp makes no mention of Isocoma veneta var. decumbens
Jeps., Man. Fl. Pi. Calif.: 1029, 1925 under synonyms.

Haplopappus venetus ssp. oxyphyl lus (Greene) Hall =

Isocoma veneta var. oxyphylla (Greene) Beauchamp
SEE reference under Haplopappus acradenius.

Machaeranthera arida Turner & Horne =

Machaeranthera coulter! A. Gray var. arida (Turner & Horne)
Turner

B.L. Turner, Reduction of Machaeranthera arida to Varietal Status
under M. coulter i (Asteraceae-Astereae) . Phytologica 61: 143-146.
1986.

Perezia microcephala (DC.) Gray *

Acourtia microcephala DC.

Reveal, J.L. & R.M. King, Re-establishment of Acourtia D. Don
(Asteraceae). Phytologica 27: 228-232, 1973.

Pulicaria hispanica (Boiss.) Boiss. ■

Pulicarla paludosa Link, in Schrader

Flora Europea 4: 137, 1976. (recognized by Kartesz & Kartesz,
1980) .

BORAGINACEAE

Plagiobothrys dalifornicus (Gray) Greene var. californicus. *

Plagiobothrys collinus (Phil.) Jtn. var. californicus (Gray)
Higgins

Author omission in Roberts, Crossosoma 12: 7, 1986.

BRASS ICACEAE

The following changes are listed in Kartesz & Kartesz (1980) and
originate with Flora Europea.

Eruca sativa Hill. =

Eruca vesicaria (L.) Cav.

Flora Europea 1: 340, 1964.

- 3 -

BRASSICACEAE (continued)

Dr aba verna L. =

Erophila verna (L.) Chevall.
Flora Europea 1: 312, 1964.

CAMPANULACEAE

Downingia concolor Greene var. brevoir McVaugh. -

Downingia concolor ssp. brevoir (McVaugh) Beauchamp

R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of

San Diego County, California. Phytologica 59: 437-440, 1986.

CARYOPHYLLACEAE

Arenar ia macrophylla Hook. is recognized as Moehr ingia
macrophylla (Hook.) Fenzl in Kartesz & Kartesz (1980). The genus
Moehringia is recognized in Flora Europea 1:123-125, 1964. Add-
itional information concerning the status of the genus is des-
cribed in J. McNeill, Rhodora 82: 495-501, 1980. Moehringia has
not been adopted by southern California authors.

Cerastium vulgatum L. *

Cerastium fontanum Baumg. ssp. triviale (Link.) Jalas
Flora Europea 1:142, 1964.

Silene cucubalus Wibel. *

Silene vulgaris (Moench) Gar eke
Flora Europea 1:168, 1964.

COASTRACEAE

Euonymus occidental is Nutt, ex Torr. var. parishii (Trel.) Jeps. =
Euonymus occidental ia ssp. parishii (Trel.) Beauchamp
R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of
San Diego County, California. Phytologica 59: 437-440, 1986.

CHQJOPODIACEAE

Salsola iberica Sennen and Pau. =

Salsola australis R. Br.

Originates with work of T.C. Fuller (Calif. Dept. Agriculture
retired) showing that Salsola iberica Sennen and Pau. is not
available for this species. I have seen a personal communication
dated 1985 from Fuller describing the situation but no published
paper. Lathrop & Thorne (1985) recognize this name.

EUPHOFBIACEAE

Euphorbia polycar pa Benth. =

Chamaesyce polycarpa (Benth.) Mi lisp.

Author omission in Roberts, Crossosoma 12: 10, 1986.

6

FABACEAE

Cercidium vs. Parkinsonia.

See reference in Roberts, Crossosoma 12: 11, 1986. Or. C.E. Jones
et al. (California State University, Fullerton) have found add-
itional support for including American species of Cercidium under
Parkinsonia as interpreted by Isely (1975). This evidence will be
presented elsewhere later this year by Jones et al.

Thermopsis macrophylla H. & A. var. semota Jeps. =

Thermopsis macrophylla ssp. semota (Jeps.) Beauchamp

R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of

San Diego County, California. Phytologica 59: 437-440, 1986.

LDMUmfiCEAE

Limmanthes gracilis Howell var. parishii (Jeps.) C.T. Mason s
Limnanthes gracilis ssp. parishii (Jeps.) Beauchamp
R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of
San Diego County, California. Phytologica 59: 437-440, 1986.

POLEMONIACEAE

Langloisia matthewsii (Gray) Greene =

Loeseliastrum matthewsii (Gray) Timbrook

Timbrook, S., Segregation of Loeseliastrum from Langloisia (Pol-
emoniaceae). Madrono 33: 157-174. 1986.

Langloisia punctata (Cov.) Goodd. *

Langloisia setosissima (T. & G.) Greene ssp. punctata (Cov.)
Timbrook

SEE reference under Langloisia mattewsii.

Langloisia schott ii (Torr.) Greene =

Loeseliastrum schott ii (Torr.) Timbrook
SEE reference under Langloisia mattewsii.

Langloisia setosissima (T. & G.) Greene =

Loeseliastrum setosissima ssp. setosissima.

SEE Langloisia punctata .

- 5 -

POLYGONACEAE

Chorizanthe staticoides Benth. ssp. chrysacantha (Goodm.) Munz.

This is the correct form. C. staticoides var. compacta (Goodm.)
Reveal (see Roberts, Crossosoma 12: 13, 1986) has not been valid-
ly published and is unavailable.

RHAttOCEAE

Ceanothus greggii A. Gray var. perplexans (Trel.) Jeps. =

Ceanothus greggii ssp. perplexans (Trel.) Beauchamp

R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of

San Diego County, California. Phytologica 59: 437-440, 1986.

RQSACEAE

Rubus glaucifolius Kell. var. gander i (Bailey) Munz =

Rubus glaucifolius ssp. gander i (Bailey) Beauchamp

R.M. Beauchamp, Taxonomic Rearrangements proposed for a Flora of

San Diego County, California. Phytologica 59: 437-440, 1986.

CYPERACEAE

Scirpus robustus Pursh

Scirpus maritimus was erroneously stated as being introduced from
Europe in Roberts, Crossosoma 12: 14, 1986. S. maritimus occurs
in both Europe and North America naturally. S . maritimus is part
of a widely occurring geographic complex that has been variously
segregated and combined. Of the named segregates, definitive
characters are often ambigous at best. Recent workers have found*
that the relationship between these plants is best presented
under one name. Munz, however, oversimplified the problem by
placing plants fitting S. maritimus under the name S. robustus.
According to Wayne Ferren (UC Santa Barbara, pers. com.) , S.
robustus and S. maritimus are clearly separate but related spe-
cies. Both S. maritimus and S. robustus occur in southern
California.

JUNCACEAE

J uncus kelloggii Engelm. (S. Calif, refs.)

Listed as Juncus tiehmii Errter in Lathrop & Thorne, 1985. This
is later corrected to Juncus luciensis Errter in a list of errata
supplied with their publication.

- 6 -

POACEAE

Agropyron

I have seen no additional support for changing local Agropyron to
Elytrigia (SEE Roberts, Crossosoma 12: 15, 1986).

Elymus

I have seen no additional support for changing local Elymus to
Leymus (SEE Roberts, Crossosoma 12: 15, 1986).

Eragrostis cilianensis (All.) E. Mosher *

Eragrostis cilianensis (All.) Janchen

Perry, G. & J. McNeill, The Nomenclature of Eragrostis cilianen-
sis (Poaceae) and the Contribution of Bellardi to Allioni's Flora
Pedemontana. Taxon 35: 696-701, 1986.

Eragrostis mexicana (Homem.) Link. *

Eragrostis mexicana (Homem.) Link. ssp. mexicana.

S.D. Koch, Eragrostis mexicana, Eragrostis neomexicana and
Eragrostis virescens, the Resolution of a Taxonomic Problem.
Phytologica 58: 377-381, 1985.

Eragrostis neomexicana Vasey =

Eragrostis mexicana ssp. mexicana.

SEE reference under Eragrostis mexicana.

Eragrostis orcuttiana Vasey *

Eragrostis mexicana ssp. virescens (Presl.) S.D. Koch
SEE reference under Eragrostis mexicana.

Festuca confusa Piper 3

Vulpia microstachys (Nutt.) Munro ex Benth var. confusa (Piper)
Lonard & Gould.

Author correction in Roberts, Crossosoma 12: 15, 1986.

Festuca pacifica Piper *

VUlpia microstachys var. pauci flora (Beal) Lonard & Gould
Lonard, R.I. & F.W. Gould, The North American Species of Vulpia
(Gramineae). Madrono 22: 217-280, 1974. (SEE also F. reflexa) .

Festuca reflexa Buckl. =

Vulpia microstachys var. pauci flora.

SEE reference under Festuca pacifica . Correction in Roberts,
Crossosoma 12: 16, 1986.

- 7 -

I thank the following people for pointing out papers, problems or
clarifying southern California nomenclature used in this and my pre-
vious paper: Wayne R. Ferren (UC, Santa Barbara), Steve Junack (Santa
Barbara Botanic Garden), Dr. C.E. Jones (CSU Fullerton), Karlin G.
Marsh and Dr. Robert Thorne (Rancho Santa Ana Botanic Garden). I also
thank Robert L. Allen for proof reading this paper and Gordon A. Marsh
for his support and encouragement in attempting to keep up with the
nomenclature .

Additional References cited above:

Beauchamp, R.M. A Flora of San Diego County, California, Sweetwater
River Publications, 198 6.

Kartesz, J.T. & R. Kartesz, A Synonymized Checklist of the Vascular
Flora of the United States, Canada and Greenland, University of
North Carolina Press, 1980.

Lathrop, E.W. & R.F. Thorne, A. Flora of the Santa Rosa Plateau, South-
ern California. S. Cal. Bot. Spec. Publication 1: 1-39, 1985.

Lei linger, D.B. A Field Manual of the Ferns & Fern Allies of the
United States and Canada, Smithsonian Institution Press, 1985.
Roberts, F.M., Nomenclatural Changes in A Flora of Southern
California, P.A. Munz, 1974. Crossosoma 12: 1-16, February 1986.

PLANT COLLECTING

The Plant Conservation Roundtable's Conservation Guidelines
for native plant collecting apply to collection of U. S. native
plants for use as herbarium specimens, research material for study,
for teaching, or for experimental horticulture. If you would like
a copy of the guidelines, write to CNPS, San Diego Chapter,

P. 0. Box 1390, San Diego, CA 92112. Remember, you need a permit
to collect on public lands, and you must have the permission of the
owner to collect on private land.

LITERATURE AVAILABLE

Vascular Plants of Extreme Southwestern San Diego County is
a 36 page xerox and annotated compilation of the floristic areas
(1) South Coast, (2) Otay Mesa, (3) Mount Miguel and Jamul,

(4) Otay Mountain, and (5) Lyons Valley and Dulzura. Botanical
and common names are listed for 1,384 taxa in 114 families. This
informative piece of Work by Duffie Clemons is on sale at the
meetings of the San Diego Chapter of CNPS for $3.00 + tax, or by
mail for $4.00 (includes tax and postage). Send check made out to
CNPS to CNPS, c/o San Diego Natural History Museum, P. O. Box 1390,
San Diego, CA 92112.

AUTOBIOGRAPHICAL SKETCH BY COLEEN CORY

Aloha. I have just moved back to the mainland from a three
year stay in Hawaii. As a graduate student at CSUF, I was working
on the pollination biology of endemic species of Hawaiian birds and
flowers. Thesis work had me traveling back and forth to the islands
for three years, and, after graduating in 1984, I stayed there to
become the botanist/horticulturist at Waimea Arboretum and Botanical
Garden on the north shore of Oahu. Waimea Arboretum, an 1800 acre
garden, associated with Waimea Falls Park, specialized in growing
and propagating endangered tropical plant species from all over the
world, including those from Hawaii. We worked closely with private,
state, federal, and international agencies and groups dealing with
conservation. In addition to coordinating the endangered species
projects (inventories, collecting trips, propagation, etc.), I
assisted botanists from Honolulu's Bishop Museum in gathering
specimens of cultivated plants (both native and introduced) for the
revision/update of the classic Mary Neal book In Gardens of Hawaii .

But economic factors (a 22% higher cost of living and what
seemed like a 22% lower wage scale) , plus a bad case of "rock fever”
(like cabin fever, especially to us native Californians) , and a
climate that was just too wet (80% humidity most of the time. . .
even my books went moldy) , prompted me to move back to California
and accept the job of plant propagator at the Los Angeles State and
County Arboretum. Now my main task is to relearn as many California
native plants as possible. (It is amazing how much can be forgotten.)
The SCB field trips should provide an excellent chance to do so, and
I look forward to meeting many of you on these outings.

ANNOUNCEMENTS

TROPICAL RAINFORESTS OF MEXICO CONSERVATION TALK. APRIL 2, 7:30 p.m.
FULLERTON HIGH SCHOOL LITTLE THEATER. The Orange County Group of the
Sierra Club and Fullerton Arboretum are co-sponsoring the presentation
of a talk entitled, "Save the Tropical Rainforests of Mexico: A Race

Against Time." Guido Rahr, of the Nature Conservancy, and Ramon Perez
Gil Salcido, renowned biologist and Deputy Director of the Institute
of Natural History in Chiapas, Mexico will give a slide show and
lecture on the impact these two organizations are having on
rainforest conservation.

.9.

April 4 (Saturday). Southprn California Botanist? Spring Plant
and Book Sale. 3:00 A.M. to 3:00 P .M. This year’s plant and
book sale will be held in the visitor oarkinq lot at Rancho Santa
Ana Botanic Garden. Featured will be a wide selection of native
and drought tolerant exotic plants for use in water-conserving land-
scapes. Take 1-10 east to Indian Hill Blvd. Go north to Foothill
Blvd. Go east on Foothill to College Avenue. The address is 1500
N. College Avenue in Claremont. For additional information call
Geoff Smith at (714) 871-8000. Fxt. 371 or (714) 929-5248 during
evenings .

Symposium on the Scientific Value of the Desert. April 5, 1987,

The symposium to be held at the Manchester Conference Center of the
University of San Diego, will be hosted by the Anza-Borrego Foundation
to celebrate its 20th anniversary, and the Environmental Studies
Program of the University of San Diego. Papers will deal with
anthropology, archaeology, biology, ecology, geology, paleontology,
and wildlife management. Applications and abstracts of proposed
papers should be sent by January 1, 1987 to Dr. James Richard,

P. 0. Box 777, Borrego Springs, CA 92004. The $15.00 registration/
luncheon fee will be collected at the symposium. For more information
call Dr. James Rickard, (619) 767-3776, or Dr. Richard Phillips
(619) 260-4600, ext. 4451.

Second Native Plant Revegetation Symposium . April 15-18,*

Hanalei Hotel, San Diego. Riparian Systems will be featured in a
special section of the Second Native Plant Revegetation Symposium,
co-sponsored by the Southern California Chapter of the Wildlife
Society (TWS) , the San Diego Chapter of CNPS , and the Riparian
Revegetation Work Study Group ( RRWSG) . The goal is to exchange
information on current research, and new or refined techniques and
projects using native plants in revegetation and landscaping. There
will also be a poster and commercial exhibit. For more information,
write to: Native Plant Revegetation Symposium, 3808 Rosecrans St.,

#373, San Diego, CA 92110; or call John Rieger (WS). , (619) 237-6754;
robbie Steele (CNPS), (619) 175-3732; or Anne Sands (RRWSG) ,( 415) 381-2

- 10 -

SOUTHERN CALIFORNIA BOTANISTS

r-lant Lovers of Southern California . The journal of the Southern
California Botanists, CROSSOSOFA, provides an ideal means by which
you can nublish anything of botanical interest to southern Califorians.
Do you have a favorite field trips, gradening hints, or some pre-
liminary data that you'd libe to have in print? Submit it to:

Dr. Allan Schoenherr

Division of Biological Sciences, Fullerton College
321 E. Chapman Ave .

Fullerton, CA 92634

- 11 -

APRIL 11 , 12 ( SATURDAY-S UNDAY ) . NATIVE PLANT SOCIETY DESERT
RAMADA. This is a special display that probably will be set up
in Blair Valley, Anza Borrego area. The goal is to provide a point
where people can stop and see the desert in full bloom. Meet at
Dudley Bakery in Santa Isabel at 9:00 a.m. For more information
contact Duffie Clemons at (619)277-3719.

RESTORING SELF-SUFFICIENT VEGETATION . (University of San Diego
Seminar) . WED., APRIL 15, 1:00-4:30 p.m. This seminar is designed
to accompany the Second Native Plant Revegetation Symposium.
Registration fee is $55.00. Practical emphasis will be placed upon
California terrestrial vegetation, but the principles discussed
will be applicable to the vegetation aspects of most comparable
biological systems. Many specific subjects will be covered, such
as nutrient requirements, symbiotic relationships, (mycorrhizae and
nitrogen-fixation), plant-soil-water relationships, and plant
adaptation to stress. Basic restoration of many vegetation types,
including chaparral, soft chaparral, perennial grassland, riparian,
and woodland plant communities and their modification for aesthetic
enhancement, fire hazard reduction, and other purposes, will be
discussed.

OPEN HOUSE AT RANCHO SANTA ANA BOTANI C GARDEN . SATURDAY , APRIL 15
FROM 10:00 a.m. to 4: 00 p.m. The theme of the open house is
"CELEBRATING SIXTY YEARS." Admission is free. The day will provide
a potpourri of special indoor and outdoor activities for all ages
including a variety of lectures, interpretative displays, book and
plant sales, interesting food booths, entertainment, door prizes
and educational activities for children and adults. Special guided
tours will also be available throughput the day.

Open House visitors will also have the opportunity to behind-
the-scenes to see areas of the Garden not usually open to the public.
The Garden's impressive dried plant collection of nearly one million
specimens, the Library and the nursery, where horticultural research
takes place, will all be open.

The Rancho Santa Ana Botanic Garden is located at 1500 N. College
Ave., east of Indian Hill and Foothill Blvd. in Claremont. Call
714-625-8767 for information.

- 12 -

I

Mary D eDecker Symposium. From Thursday, April 30 through Sunday,

May 3. The University of California White Mountain Research Station
with the California Native Plant Society and its Bristlecone Chapter
is co-sponsoring a symposium and field trip honoring Mary DeDecker.
The topic of the symposium is the flora and plant biology of the
eastern Sierra, Owens Valley, White-Inyo Mountains, and western Basin
and Range Provinces. The field trip to the. Eureka Dunes on Saturday,
May 2, will be led by Mary DeDecker and others? a BBQ cookout will
follow. Registration for the conference is $10 per person, and
dormitory facilities are available for $1.50 per night. Motels are
available in Bishop. Those who wish to attend all or part of the

symposium or to present a paper should contact Bart O'Brien,

(408) 741-9534 evenings, (415) 941-7705 days. Registration and

abstracts should be sent to Dr. Clarence A. Hall, Jr., White Mountain
Research Station, 6713 Geology Building, University of California,

Los Angeles, CA 90024; phone: (213) 825-2093.

SOUTHERN CALIFORNIA ACADEMY OF SCIENCES ANNUAL MEETING . MAY 8, 9 .

This year's meetings will be held at California State University,

Los Angeles. Watch for further details.

Third Annual Southwestern Botanical Systematics Symposium: Advances
in Plant Systematics and Ecology. Rancho Santa Ana Botanic Garden
is presenting the third in its series of annual symposia aimed at
systematic botanists in the southwestern United States and Mexico.
This year's roster of speakers will present original research papers
on a wide range of subjects concerning plant systematics and ecology.
An informal social for all participants will be held in the home
demonstration garden on Friday evening. May 22 . Invited papers
will be presented on Saturday, May 23, in the auditorium of the
main administration building. Library and herbarium facilities will
be available during the meeting, and low-cost housing will be
available locally. Detailed information on housing will be sent
with confirmation of registration. Please register early, as space
is limited to 100. For information or reservations contact:

Rancho Santa Ana Botanic Garden, Botanical Systematics Symposium,

1500 North College Avenue, Claremont, CA 91711; phone: (714) 6*5-8767.

-13-

Jepson Manual Revision

A revision of the classic MANUAL OF THE FLOWERING PLANTS OF
CALIFORNIA by W. L. Jepson is currently underway. It stands to
benefit from Environmental License Plate Funds. In the draft
budget for 1987-88, Item 3600, Department of Fish and Game, includes
$300,000 for support of the Jepson Manual Project. Unfortunately,
we must convince the legislature to retain this item in the
Governor's budget. Write to your assemblyman and senator today
urging its retention.

Fund for Environmental Defense (F.E.D.). A new legal defense fund,
patterned after the Sierra Club Legal Defense Fund, has been established
for Orange County. F.E.D. has been established for the purpose of
providing a legal defense fund to which environmental activist groups
can apply for assistance in maintaining or increasing the quality of
life in Orange County. Specific objectives are to advise and assist
in litigation proceedings by environmental groups that seek to create,
maintain or increase the aesthetics of their surroundings, such as
(but not limited to) residential areas, parks, greenbelts or other
open space areas. Memberships ranging from $25.00 to $1000.00 are
available. For more information write to: Fund for Environmental

Defense, P. O. Box 9118, Fountain Valley, CA 92708 or call Charlotte
Clark (714) 871-8000, Ext. 263.

******

FIELD TRIPS

March 28 (Saturday). 8:00 a.m. to dusk. Pines to Palms Trip.

Geoff Smith will lead this trip to examine desert, chaparral and
mountain habitats. Meet at the S. E. corner of the Ramona Expressway
and 215 south of Riverside. Caravan from Hemet through the San
Jacinto Mountains on Hwy 74 with stops at plant communities discussed
in Geoff's recent Crossosoma article (April, 1986). Bring a sack
lunch and comfortable walking shoes. For further information contact
Geoff at (714) 526-6963 (evenings); (714)929-5248 (evenings) or
(714)871-8000, ext. 371.

- 14 -

ril 5 (Sunday). 9:00 a.m. Harford Springs Park. Steve Boyd of
uie Rancho Santa Ana Botanical Garden will lead this trip to see the
unique flora of the Gavilan Hills. We will find excellent stands of
Juniper Woodland and will look for Munz's Onion, Paysen's fclanthus,

F allbrook spinef lower and many other disjunct species. Take the 91
freeway east to the 15 and go south. Take the turnoff at Cajalco Rd.
and go east to Gavilan Rd. At Gavilan go south to Ida Leona Rd.

(1st left past the trailer park) and meet at the top of the rise on
Ida Leona. Bring a sack lunch and water.

oril 10. 11, 12 ( Friday- Sunday ) . Soring Canoe Trip . Mona Myatt
will lead this float trio along the Colorado River from south of
Blythe to north of Yuma. T'his area of the river and desert holds
many interesting biologic, scenic, and historic sites. For details
of costs, itinerary, and logistics call Mona at (818) 302-1466 during
ueekdays or (818) 447-1466 on weekends or evenings.

April 25, 26 (Friday - Sunday ) . Desert Studies Center at Soda Springs
(Zzyzx) . Trip limited to 25 individuals. A cost of approximately
$8 per person for use of the facility will be charged. Reservations
or information about the trip can be obtained from Alan Romspert by
calling (714) 773-2428. We will see a variety of desert wildf lowers
and shrubs in bloom. This looks like it may be an excellent year
for a spectacular wildf lower display in the Mojave Desert.

May 9(Saturday). 9:00 a.m. Chiquito Basin. Fred Roberts of the UCI
Systematic Museum will lead this trip to the unique deer grass portero
found at Chiquito basin in the Santa Ana mountains. We will see large
areas of chaparral, oak woodland and native grassland. This area
also contains the largest population of the San Miguel Savory
( Satureja chandleri ) , which should be in flower. Bring a lunch, sun
protection, water and a hand lens. We will meet at the intersection
of the Ortega Highway (74) and Forest Service road #6S05 that leads
to the conservation camp. For more information call Dave Bramlet
(714) 549-0647.

- 15 -

May 10 (Sunday). 10:00 a.m. Pine Canyon. Karlin Marsh will lead
this trip into the Santa Ana Mountains. Pine Canyon is a north-facing
slope tributary of Silverado Canyon. The area is a highly shaded
mixed hardwood, conifer forest with many ferns and wildf lowers. Meet
at 10:00 a.m. at the end of Silverado Canyon Road at the Forest gate.
Parking is very limited so please carpool. A ranger may let us
through to park inside the gate. If not and the lot is full, then
park down the canyon 1/4 mile, near the bridge. For more information,
call Dave or Karlin at (714) 649-024.7.

MAY 16-17 (SATURDAY -SUNDAY) . 9:30 a.m. CLARK MOUNTAIN . Barry

Prigge will lead this trip to examine the blackbrush scrub, pinyon-
juniper woodlands and the white firs located on this unique mountain
"island” in the eastern mojave desert. If time allows, we may climb
further into the mountain to look for limestone endemics, especially
Ivesia jaegeri . Bring camping gear, water, and Thorne and Prigge' s
flora of the eastern Mojave. Be prepared for cold and hot weather.
Meet in the parking lot of the gas station (formerly Stuckey's) at
the Cima Road offramp of Interstate 15.

JUNE 6 (SATURDAY). SAN MATEO MARSH . Meet at 9:30 a.m. at the
Lifeguard HQ building at top of Basilone Road offramp on 1-5, about
1 mile south of San Clemente. Celia Kutcher will lead us through
the varied habitats of a fresh water marsh that is now almost
unique in Southern California. Walk through the marsh in the a.m.,
eat lunch on the beach, look at Coastal Strand vegetation in the
afternoon on the way back to the cars. Bring lunch, water, sturdy
walking shoes; and be prepared for mosquitos, poison oak, and some
bushwhacking.

JUNE 13 - POTENTIAL FIELD TRIP TO LAGUNA MOUNTAINS , SAN DIEGO CO.

For details call Dave Bramlett at (714)549-0674.

JUNE 20-21 . ( SATURDAY -SUNDAY ) . TETRACOCCUS RIDGE , PAN AMI NT MOUNTAINS .

People interested in participating on the field trip should try to
meet at the Thorndike Campground up Wildrose Canyon in the Panamint
Mountains (see AAA map of Death Valley) on Friday night or before

- 16 -

FAUAMINT MOUNTAINS (CONTINUED)

0800 hours, Saturday morning. We will leave the campground at 0800
on Saturday morning and drive to Aguereberry Point (See AAA map of
Death Valley) . From Aguereberry Point it is a semi-arduous hike
down and up to Tetracoccus Ridge, a Dolomite (limestone) outcropping.
This outcropping provides a substrate for many endemics, most on the
CNPS Rare and Endangered Plant Inventory. In the past we have seen:
Bear Poppy - Arctomecon merriamii
Jointed Buckwheat - Eriogonum intrafractum
Death Valley Monkey flower - Mimulus rupicola
Limestone Penstemon - Penstemon calcareus
Ripley's Gilia - Gilia ripleyi

Holly-leaved Tetracoccus - Tetracoccus ilicifolius
Naked-stemmed Daisy - Enceliopsis nudicaulis
Curled-leaved Mountain Mahogany - Cercocarpus intricatus
Panamint Milk-vetch - Astragalus panamintensis
Blazing Star - Mentzelia oreophila
Limestone Phacelia - Phacelia p. perityloides
Sunday will be spent botanizing around Thorndike Camp (Water Canyon) ,
Mahogany Flats or the trail to Telescope Peak if time permits. It
can get cool in the evenings and there will be insects so come
prepared. For additional information contact Alan Romspert
( 714 ) 773 - 2428 .

FUTURE FIELD TRIP. CONE-BEAP.ING TREES OF THE SOUTHERN SIERRA NEVADA
Some time in mid-July, Allan Schoenherr will lead this motor-campinq
tcur. The trip begin? at Freeman Grove, one of the southernmost
groves of old growth sierra redwoods (Giant Sequoia), that is located
on the plateau west of the Kern River. The route will proceed down
to the Kern River, then over the eastern divide at Sherman Pass to
camp in a picturesque forest-setting west of Kennedy Meadows. The
route will continue through Kennedy Meadows and down the spectacular
east side of the Sierra Nevada. We should see at least 13 species
of coniferous trees, including 7 species of pines. While the
emphasis will be on conifers, many species of mountain wildf lowers
should be in bloom. For details, call Allan at (714) 871-8000

A

weekdays or at (714) 494-0675 evenings.

- 17 -

SOUTHERN

COSPONSORED BY THE DEPT. OF ECOLOGY

AND EVOLUTIONARY BIOLOGY AND PETER n i t TnA^tlT A

BOWLER, DIRECTOR OF THE COOPERATIUE PAT | Lt| UfJ | A
OUTDOOR PROGRAM IN THE STUDENT !.k}.l rUAIllfl

ACTIUITIES OFFICE AT UC, IRVINE VI BUI IV JIM All

13TH ANNUAL SYMPOSIUM BOTANISTS

BOTANICAL DISCOVERIES
AND EXPLORATION IN
SOUTHERN CALIFORNIA

date: SATURDAY, NOVEMBER 14, 1987

time: 8:45 AM TO 4:68 PM

place: HERITAGE ROOM, UNIVERSITV
CENTER; U.C., IRVINE

IF YOU HAVE A SUGGESTION FOR ONE OR MORE OF THE FIVE SPEAKERS,
PLEASE CONTACT ONE OF THE S.C.B. OFFICERS OR DIRECTORS.

r— ■ JRt K V O O « c ^ F- E M 3> Ft

- 18 -

SOUTHERN LA 1 IFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden
• 500 Nrvrih College Avenue
Claremont. GA Ql 7 1 1

The purpose of the SOUTHERN CALIFORNIA BOTANISTS is the study,
preservation and conservation of the native plants of California; and
the education of the public to the value of the native flora and its
habitats. It is a non-profit association foraied in 1927.

Membership benefits Include:

Various field trips throughout the state led by competent

field botanists and biologists.

A yearly plant sale featuring native California species.

An annual symposium on various aspects of the California

vegetat ion .

The SCB Journal , CROSSOSOMA .

Discounts on botanical and natural history books.

Dues are for a calendar year. New members Joining from May through
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CROSSOSOMA

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Rancho Santa Ana Botanic Garden, Claremont, CA 91711

CROSSOSOMA Vol. 13, No. 3
Managing Editor: Allan Schoenherr

June, 1987

CRASSULACEAE IN SOUTHERN CALIFORNIA DESERTS

Kei M. Nakai

Mildred E. Mathias Botanical Gardens and Herbarium
University of California, Los Angeles
405 Hilgard Ave.

Los Angeles, California 90024

INTRODUCTION

Succulent plants are often associated with desert habitats;
however, the succulent family Crassulaceae is best represented on the
cismontane (coastal) side of southern California's mountains. Of the 70
taxa currently described, only nine, 13% of the family, are found on the
transmontane (desert) side of the mountains. Apparently because of
narrow environmental requirements and tolerances most of the desert
forms are restricted in their distribution. For this reason, the
presence of these taxa can be used as indicators of interesting
microhabitats. I have prepared keys for the Crassulaceae of the desert
portions of California. Included also are a brief description
and discussion of each taxon. The geographic area covered by this
key includes the part of California that lies east of the Sierra
Nevada, Transverse and Peninsular ranges.

KEY TO THE GENERA

A. Plants annual; stamens equalling the number of sepals. 1. Crassula
AA. Plants perennial; stamens twice the number of sepals.

B. Flowering stems borne laterally from a basal

rosette of leaves 2. Dudleya

BB. Flowering stems borne terminally 3. Sedum

1. CRASSULA L.

1. Crassula erecta (H. & A.) Berger subsp. eremica (Jeps.) Wiggins
[ Crassula connata (R. & P.) Berger var. eremica (Jeps.) Bywater &
Wickens, Crassula connata var. erectoides Bywater & Wickens). Small,
much branched annual, 2-6 cm tall; leaves minute, opposite; pedicels to
3 mm long; flowers in axillary bracteate clusters; sepals 4.

- 1 -

Crassula erecta subsp. eremica is found in the Colorado or northern
Sonoran Desert, mostly in open areas where they often occur in large
masses. Bywater and Wickens (1983) transf erred this taxon to a variety
of Crassula connata and extended its geographic range from coastal
central and southern California into Baja California and central
Arizona. However, employing a scanning electron microscope, they
studied microcharacteristics of seeds from herbarium specimens. I
cannot accept their findings until living specimens have been studied.

2. DUDLEYA Britt. 6 Rose

A. Petals connate for. more than a third of their length.

B. Rosette 1 to 4 dm in diameter? leaves densely white

pulverulent? flowers red •

3. D. pulverulenta subsp. arizonica

BB. Rosette usually less than 1 dm in diameter? leaves
glaucous, not densely white pulverulent? flowers pale

yellow, often marked with red striations 1. D. abramsii

AA. Petals connate for less than a third of their length.

C. Pedicels 0.5 to 6 mm long.

D. Petals pale yellow, often with red lines along the

midribs, 2 to 3 mm wide, connate for 1.5 to 4.5 mm.
1. D. abramsii

DD. Petals red, 3-5 mm wide, connate for 1 to 2 mm

2. D. lanceolata

CC. Pedicels 5 to 20 mm long.

E. Rosette leaves 5 to 30 cm long, 1 to 4 cm wide?
petals red, connate 1 to 2 ran; n» 34. Stoddard and

Ord Mountains 2. D. lanceolata

EE. Rosette leaves 3 to 15 cm long, 0.5 to 2.5 cm wide;
petals yellow, occasionally tinted with red, to pale
yellow; n - 17, 68, 85. Panaraint Mountains and the
higher ranges in the Mojave Desert 4. D. saxosa

1. D. abramsii Rose (Pig. 1). Plants simple or cespitosely branched?
rosettes usually 2 to 7 cm in diameter and consisting of 10 to 20
oblong-lanceolate to oblanceolate glaucous leaves that are 0.5 to 6 cm
long and 5 to 15 mm wide; floral stem 5 to 15 cm tall; inflorescence
consisting of 2 to 3 simple to bifurcate branches; petals pale yellow,
often with red striations along the midribs, connate for 1.5 to 4.5 mm;
epipetalous filaments 2.5 to 6 mm long, adnate 1 to 3 mm; antisepalous
filaments 4.5 to 7 mm long, adnate 2 to 4 mm. Plowering from April to
June (July).

Dudleys abramsii is a polytypic species consisting of four subspecies,
three of which are found in the desert. Dudleys abramsii subsp.
abramsii is found in southern San Diego County and northwestern Baja

- 2 -

Figs. 1-5. — 1. Dadleya abramsii. — 2. D. lanceolafca. — 3. D.

pulverulenta. — 4. D. sazosa subsp. aloides. — 5. Sedum niveum.

- 3 -

California between 800 to 2000 m. in elevation, Dudleya abramsii subsp.
affinis in. ed. and subsp. calcicola (Bartel & Shevock) in. ed. are
primarily montane taxa, however, they do marginally enter the desert;
subsp. affinis near Cushenbury Springs on the north slopes of the San
Bernardino Mountains [Deburg 2608 (RSA), Nakai 1147 (CAS, LA), Parish
1629 (POM) ] ; subsp. calcicola in Jawbone Canyon, southeast slopes of the
Piute Mountains, Kern County [Moran & J.M. Zavinovich 24196 (SD)1.

Key to the subspecies

A. Rosette leaves oblong to oblong-lanceolate; plants with few to
many branches.

B. Inflorescence of 2 to 3 mostly simple branches; petals

usually with red striations subsp. abramsii

BB. Inflorescence of 2 to 3 simple to usually bifurcate

branches; petals with or without red striations

subsp. calcicola

AA. Rosette leaves elliptic to oblanceolate; plants usually

unbranched •••••••••••••••••••••••••••••••••...•••»• subsp . affinis

2. Dudleya lanceolata (Nutt.) Britt. & Rose (Fig. 2). Plants simple to
few branched; rosette consisting of 10 to 25 green to (in our area)
glaucous lanceolate to oblong- lanceolate leaves that are 5 to 20 cm
long and 1 to 4 cm wide; floral stem 2 to 7.5 dm tall; inflorescence of
2 to 4 branches that bifurcate once or twice; pedicels mostly 3 to 6 mm
long, rarely to 12 mm long; petals (in our area) red. n =* 34. Near
Stoddard Well to Ord Mountains, San Bernardino County. Flowering from
April to June.

This ubiquitous California and northwestern Baja California species is
primarily found among chaparral and riparian woodland vegetation.
Dudleya lanceolata is known in the western Mojave Desert from Stoddard
Well to the Ord Mountains at an elevation of 1000 to 1250 m. It was
first collected by P. Hutchison (U.C. Bot. Gard. 57.305-1) at the pass
between Lucerne Valley and Stoddard Well, on east facing slope in rock
clefts, elevation 4000 ft. (1220 m) and identified as D* cymosa (Lem.)
Britt. & Rose. After cultivating plants from the same and other
localities and counting their chromosome number, the dudleyas were
clearly not D. cymosa, which is a small diploid montane species, nor D.
saxosa subsp. aloides, a taxon one would expect in this area (see
description below). With respect to its location in this new habitat,
D. lanceolata could be considered the most adaptable taxon within the
genus .

3. Dudleya pulverulenta (Nutt.) Britt. & Rose subsp. arizonica (Rose)
Moran (Fig. 3). Plants unbranched; rosette 5 to 30 cm in diameter and
consisting of 10 to 35 more or less mealy-pulverulent leaves that are
oblong to oblong-obovate, 3 to 15 cm long, 1 to 5 cm wide; floral stem
1.5 to 6 dm tall; petals red to reddish yellow. Flowering from April to
June.

- 4 -

This is the most widespread taxon in the genus , found from central
Arizona and southern California in the United States, Sonora and central
Baja California in Mexico. It occurs on granitic, volcanic or calcareous
substrates, occasionally under shrubs or among cacti. Munz (1974) and
Munz and Keck (1959) have maintained this taxon as a species, however,
Moran (1943) presents a convincing argument for maintaining this taxon
as a subspecies of D. pulverulenta, and a recent study of the
epicuticular wax of Dudleya using the scanning electron microscope
(McCabe 1985) seems also to support that argument.

4. Dudleya sazosa (M.E. Jones) Britt. & Rose (Pig. 4). Plants
unbranched to several cespitose branches; rosette 3 to 15 cm in
diameter and consisting of 10 to 25 green to mostly glaucous
oblong to oblong-lanceolate leaves that are 3 to 15 cm long and
0.5 to 2.5 cm wide; floral stem often reddish, 5 to 40 cm tall;
pedicels often as long or longer than the corolla; petals yellow to
greenish-yellow, 9 to 20 mm long, connate 1.5 to 4 mm. n * 17 in
subsp. aloldes (Rose) Moran, n ■ 68 or 85 in subsp. sazosa. Flowers
from April to June.

Dudleya sazosa is a polytypic interior species with three comparatively
distinct subspecies, two of which are found in California. Dudleya
sazosa subsp. sazosa is a high polyploid taxon, n * 68 or 85 (octoploid
or decaploid, Uhl & Moran 1953), and is found in the Panamint Mountains
between 1100 to 2200 m in elevation. The plants are comparatively
small, individual rosettes are 5 to 10 ctn in diameter, floral stems 5 to
20 cm long. Petals are a bright yellow often tinged with red. It is
often on north facing slopes on granite or limestone. Dudleya sazosa
subsp. aloldes occurs over a larger area, from the higher ranges of the
eastern Mojave Desert to the desert slopes of the Peninsular Ranges of
southern California and the Sierra Juarez in Baja California (Wiggins
1980). Distribution occurs primarily on granites from 250 to 1675 m in
elevation. It is larger than subsp. sazosa, rosettes 5 to 15 cm in
diameter, floral stems 10 to 40 cm long. Petals are greenish-yellow to
yellow, rarely tinged with red. It is also a diploid, n * 17 (Uhl &
Moran 1953).

3. SEDUM L.

1. Sedua nlveua A. Davids. (Pig. 5). Plants prostrate with thick,
fleshy stems 1 to 3 dm long; leaves oblong-obovate, rather fleshy, 5 to
6 mm long; flowers solitary or in small cymes; petals white, often
tinged pink. Flowers from June to August.

Sedua nlveua is primarily a montane species, found at an elevation
between 2050 to 2900 m. Ito only known "desert" occurence is in the New
York Mountains of the eastern Mojave Desert at an elevation of 2050 to
2200 m (Henrickson & Prigge 1975). This species has widely disjunct

r 5 —

populations , occuring in the New York Mountains , the San Bernardino
Mountains (Holcomb Valley , Sugar loaf Mountain , Dollar Lake), and the
Santa Rosa Mountains in California, and Observatory Peak, Sierra San
Pedro Martir in Baja California. It grows on granite, quartzite, and,
in Holcomb Valley, limestone.

ACKNOWLEDGMENT

I would like to thank Barry Prigge and Stephen McCabe for their review

and constructive criticism of an earlier manuscript.

REFERENCES

Bywater, Marie and G. E. Wickens. 1983. New World species of the
genus Crassala. Kew Bull. 39:699-728.

Henrickson, J. and B. Prigge. 1975. White fir in the mountains of
eastern Mojave Desert of California. Madrofio 23:164-168.

McCabe, S. 1985. Epicuticular wax patterns as biosystematic characters
in Dudleys (Crassulaceae) . MJV. Thesis. San Francisco State
University, San Francisco, CA.

Moran, R.V. 1943. Dudleys pulverulenta. Des. PI. Life 15:86-74.

Munz, PJV. 1974. A flora of Southern California. Univ. Calif. Press,
Berkeley.

Munz, P -A., and D.D. Keck. 1959. A California Flora. Univ. Calif.
Press, Berkeley.

Uhl, C.H., and R.V. Moran. 1953. Cytotaxonomy of Dudleya and
Hasseanthus. Amer. J. Bot. 40:492-501.

Wiggins, I. 1980. Flora of Baja California. Stanford University
Press.

AUTOBIOGRAPHY - Jon E. Keeley, Occidental College
Chaparral covered canyons are among my earliest recollections.

At 5 years of age my family moved to the rural community of Bonita
in San Diego County and took me with them. Much of my time was spent
exploring the wilderness around me, often constructing "forts" under
aborescent Laurel-leaf Sumacs. Some say early childhood experiences
are critical in shaping a person's life - I'm a true believer; a
modern day chaparral "fire" ecologist who recalls making campfires
in our forts beneath sumac canopies. That you are reading this today
attests to the fact that none ever escaped, although wildfires were
common enough in our neighborhood. During my childhood, however.

- 6 -

chaparral was just brush, as birds were my true interest; identifying
them, collecting nests, raising barn owls, repairing injured raptors
and taxodermy (not necessarily in that order) .

Although growing up with a strong natural history bent, I never
focused my interests until enrolled as a junior in an ecology class
at San Diego State University (then SDS College and briefly CSUSD) .

For a class project I compared the vegetation types growing on north
and south-facing slopes of San Miguel Mountain. At the outset I
was convinced (incorrectly) that they would be the same, but it was
an excuse to be outdoors. This project forced me to learn some of
the flora, something which so intrigued me I avidly collected the
rest of the year; consequently getting a D in chemistry and F in
another biology course. By the time I actually took a plant taxonomy
course I already had a reasonable size herbarium and knew several
hundred species. So impressive was I, that the girl. Sterling,

sitting across from me in class later married me.

My first ecology course was also crucial in that on a field trip

to Borrego Desert I decided upon my future profession. It was clear
to me that ecology professors "had it made," spending weekends in the
field and getting paid for it.

After a master's thesis on post fire recovery of chaparral at SDSU,
my wife and I moved to the University of Georgia for our Ph.D.'s.

After my first Georgia summer at 95° and 95% relative humidity, plus
bugs, my research steered away from field work and I spent the remainder
of my time in the lab studying the effects of flooding on alcoholic
fermentation in tree roots. These research interests in fires and
floods later prompted one seminar introduction as "a disaster ecologist."

Since graduating I have been on the faculty at OXY and am
currently the Biology Department Chair. My longstanding interest in
chaparral has not diminished, and currently Sterling and I are
studying the mechanism behind how charred wood chemically stimulates
the germination of many postfire annuals. Since returning to California
I have also become intrigued with vernal pools, sparked in large part
by the discovery that some of the submerged aquatic species possess
CAM photosynthesis, previously considered solely as an adaptation
to xeric environments. This find has proven to be productive as I
have spent a year and a half on a Guggenheim Fellowship traveling to
South America for further research in this area.

- 7 -

BAJA CALIFORNIA FLORAS

Dr. Robert Thorne of Rancho Santa Ana Botanic Garden is
composing a flora for the northern portion of Baja California.

As a product of that effort he has compiled a series of valuable
checklists for selected localities in Baja California. The following
list of vascular plants found in Guadalupe Canyon is the first of
what will be a series of such lists to be published by CROSSOSOMA.

VASCULAR FLORA OF THE LOWER PORTION OF CA&ON GUADALUPE,
SIERRA DE JUAREZ, BAJA CALIFORNIA, MEXICO

By

Robert F . Thorne
Rancho Santa Ana Botanic Garden
Clarement, California

This list of species is for plants mostly collected or observed
in the lower canyon or below its mouth on two field trips sponsored
by Southern California Botanists on February 18-20, 1984, and
March 21-23, 1986. The mouth of Guadalupe Canyon is ca. 250 m in
elevation and near 32° 10* N - 115° 46' W; at about 600 m the
coordinates are ca. 32° 08' N - 115° 48* W, the flora having been
surveyed only between 200 and ca. 600 m. * — presumbably introduced
species and ? = immature specimens probably correctly determined.
Acanthaceae

Jueticia call formica (Benth.) D. Gibson ( Beleperone calvformca
Benth. )

Adiantaceae

Notholaena parryi D. C. Eaton
Agavaceae

Agave deserti Engelm. ssp. deserti
Amaranthaceae

Amaranthus powellii S. Wats.

Apocynaceae (incl. Asclepiadoideae)

Aeclepiae albicans S. Wats.

Matelea parvi folia (Torr.) Woods

Sarcostemma cyanchotdes Decne. ssp. hartwegii (Vail) R. Holm
Arecaceae

Brahea armata S. Wats.

Washingtonia filifera (Linden) Wendl.)

Asteraceae

Ambrosia dumoea (A. Gray) Payne
Ambrosia flexuosa (A. Gray) Payne
Ambrosia ilici folia (a. Gray) Payne
Artemisia dracunculus l.

Baccharis salici folia (R. & P.) Pers. (B. glutinosa (R. & P«)
Pers. )

Baccharis sarothroides A. Gray
Bebbia juncea (Benth) . Greene
Chaenactis carphoclinia A. Gray
Cirsium neomexicanum A. Gray

- 8 -

Asteraceae (Cont.)

*Conyza canadensis (L.) Cronq.

Dyaaodia porophylloide8 A. Gray
*Eclipta proatrata (L.) L. (

Enoelia farinoaa A. Gray ex Torr.

Gnaphalium caneseena DC (

^Gnaphalium luteo-album L.

Gnaphalium Nutt.

Gutierreaia sarothrae (Pursh) Britt. & Rusby
Haplopappue propinquua Blake
Hymenoolea aalaola T. & G. var.

Monoptilon bellioides (A. Gray) Hall
Palafoxia arida Turner & Morris
Parity le emoryi Torr. in Emory

Pleuroooronia pluriaeta (A. Gray) King & Robins.

Porophyllum graaile Benth.

Seneoio flavus (Decne.) Sch.-Bip. in Webb & Berth. (S.

mohavan8ia A. Gray)

*Saneoio vulgaris L.

Solidago oalifornioa Nutt.

Solidago con finis A. Gray
*Sonohus oleraoeua L.

Stephanomeria pauoi flora (Torr.) A. Nels.

Sty Iodine micropoidea A. Gray

Teaaaria aericea (Nutt.) Shinners ( Pluahea aerioea (Nutt.)

Cov. )

Trichoptilium inoieum (A. Gray) A. Gray
Trixie oalifornioa Kell. var. oalifornioa
Viguiera pariahii Greene

*Xanthium atrumarium L. var. oanadense (Mill.) T. & G.
Bignoniaceae

Chilopaia line aria (D. Don) Sweet ssp. arouata (Fosb.) Henricks.
Boraginaceae

Amainkia intermedia F. & M.

Cryptantha anguati folia (Torr.) Greene
Cryptantha barbigera (A. Gray) Greene
Cryptantha maritime (Greene) Greene

Cryptantha miorantha (Torr.) Jtn. ssp. lepida (A. Gray) Mathew
& Raven

Cryptantha murioata (H. 6 A.) Nels. and Macbr. (incl. var.

jone8ii (A. Gray) Jtn.)

Cryptantha pte roeary a (Torr.) Greene
Cryptantha raeemoaa (S. Wats.) Greene
Peotooarya setosa A. Gray
Brassicaceae

Deeourainia pinnata (Walt.) Britt.

Lepidium laaiooarpum Nutt, ex T. & G.

Lepidium virginioum L.

Lyrooarpa ooulteri Hook. & Harv. var. palmeri (S. Wats.) Roll.
Sisymbrium irio L.

Sisymbrium orientals L.

• Burseraceae

Burs era miarophylla A. Gray
Cactaceae

Ferooaotue aoanthoidee (Lemaire) Britt. & Rose
Mammillaria dioioa K. Bdg.

Opuntia eohinocarpa Engelra. & Bigel.

Opuntia phaeaoantha Engelm. var. disoata (Griff.) L. Bens &
Walkingt.

Camp anul ace ae

Nemaoladua glanduliferua Jeps. (incl. var. disoata McVaugh)
Caryophyllaceae

Aohyronyohia oooperi T. & G.

Chenopodi ace ae

Atriplex polyoarpa (Torr.) S. Wats.

Chenopodium fremontii S. Wats.

*Chenopodium murale L.

Crassulaceae

Dudleya abramsii Rose
Cucurbitaceae

Cuourbita palmata S. Wats.

- 9 -

Cyperaceae

Cyperua laevigatus L.

Cyperus odoratua L.

Elecharia geniculata (L.) R. & S.

Scirpus americanus Pers. (S. olneyi A. Gray)

Datiscaceae

Dati8ca glomerata (Presl.) Baill.

Dracaenaceae

Nolina bigelovii (Torr.) S. Wats.

Ephedraceae

Ephedra calif ornica S. Wats.

Euphorbiaceae

Argythamnia lanceolata (Benth.) Muell.-Arg. ( Ditaxis lanceolata
(Benth.) Pax & K. Hoffm.

Chamae8yce melanadenia (Torr.) Millsp. (Euphorbia melandenia
Torr.)

Chamae8yce pediculifera (Engelm.) Rose & Standi. (Euphorbia
pediculifera Engelm.)

Chamae8yce polycarpa (Benth.) Millsp. var. hirtella (Boiss.)

Millsp. ( Euphoriba polycarpa Benth. var. hirtella Boiss)
Chamae8yce aetiloba (Engelm. ex Torr.) Millsp. in Parish
(Euphorbia aetiloba Engelm.)

Croton calif omicua Muell.-Arg. var. calif ornicua
Euphorbia eriantha Benth.

Fabaceae

Accacia greggii A. Gray ssp. arizonica (Isely) ined.

Amorpha fruticoaa L. ssp. occidentalia (Abrams) ined.

Aatragalua palmeri A. Gray

Caeealpinia virgata E. M. Fischer ( Hoffmanseggia microphylla
Torr.

Calliandra eriophylla Benth.

Cercidium microphyllum (Torr.) Rose & Jtn.

Dale a mo Ilia Benth.

Lotu8 argophyllua (A. Gray Greene ssp. argophyllua

Lotua argyraeu8 Greene

Lotua rigidua (Benth.) Greene

Lotua atrigoaua (Nutt, in T. & G.) Greene

Lupinua concinnua Agardh. ssp. orcuttii (S. Wats.) D. Dunn.
Lupinu8 ex cubitus Jones ssp. auatromontanua (Heller) Beauchamp
Lupinua apareiflorue Benth. ssp. apareiflorue
Marina orcuttii (S. Wats.) Bameby ( Dalea orcuttii S. Wats.)
Marina parryi (T. & G) Barneby ( Dalea parryi T. & G.)

*Melilotue albue Medic.

*Me li lotus indicua (L.) All.

Olneya teaota A. Gray
*Pithecellobium dulce (Roxb.) Benth.

Proaopie glanduloaa Torr. ssp. torreyana (L. Bens.) E. Murray
Paorothamnua apinoaua (A. Gray) Bameby ( Dalea 8pino8a A. Gray)
Seebania exaltata (Raf . ) Cory)

Fouquieriaceae

Fourguieria splendens Engelm. ssp.

Gentianaceae

Euatoma exaltatum (L.) Griseb.

Geraniaceae

*Erodium oicutarium (L.) jL'Her.

Hydrophyllaceae

Eriodictyon trichocalyx Heller ssp. lanatum (Brand) Munz
Eucrypta ohryaanthemifolia (Benth.) Greene var. bipinnatifida
(Torr.) Const.

Nama demies urn A. Gray
Phacelia af finis A. Gray
Phacelia distana Benth.

Phacelia pedicellate A. Gray
Pahacelia euf fruteacens (Parry) Const.

Juncanceae

Juncua acutua L. ssp. leopoldii Snogerup
Juncua bufoniua L. var. bufonius
Juncua microphyllua Cov.

Juncua mexicanua Willd. ex R. & S. F.

Juncua xiphioidea E. Mey.

- 10 -

Krameriaceae

Krameria grayi Rose & Painter
Lamiaceae

Hyptis emoryi Torr.

*Marrubium vulgare L.

Loasaceae

Eucnide rupestris (Baill.) Thoraps. & Ernst

Mentzelia hirautissima S. Wats. var. stenophylla (Urban & Gilg)

Jtn.

Mentzelia veatohiana Kell.

Petalonyx linearis Greene
Petalonyx thurberi A. Gray ssp. thurberi
Lythraceae

iythrum calif brnicum T. & G.

Malpighiaceae

Januaia graailia A. Gray
Malvaceae

Abutilon incanwn (Link) Sweet ssp. pringlei (Hochr.) Fleger &

Lowe

Abutilon palmeri A. Gray (incl. A , aurantiacum S. Wats.)

Hibiscus denudatus Benth.

Horsfordia newberryi (S. Wats.) A. Gray
*Malva parvi flora L.

Spaeralcea ambigua A. Gray ssp. rosacea (M. & J.) Kearney
Nyctaginaceae

Abrongia villosa S, Wats. var.

Allionia incamata L.

Boerhavia diffusa L. [B . ooccinea Mill.)

Mirabilia bigelovii A. Gray
Onagraceae

Camiaaonia calif omica (Nutt.) ex T. & G.) Raven
Camisaonia oardiophylla (Torr.) Raven
Camissonia chamaenerioidea (A. Gray) Raven
Camiaaonia claviformia (Torr. & Frem. ) Raven ssp.

(Munz) Raven

Camiaaonia hirtella (Greene) Raven

Epilobium canum (Greene) Raven ssp. lati folium (Hook.)

Raven ( Zauechneria calif omica Presl. ssp;. lati folia (Hook.) Keck)
Oenothera calif omica (S. Wats.) S. Wats.

Oenothera elata H.B.K. ssp ,Hirsutiasima (A. Gray ex S. Wats.)

Dietrich (0. hookeri T. & G. ssp. griaea (Bartlett) Munz
Orchidaceae

Epivaatia gigantea Dougl. ex Hook.

Paparevaceae

Argemone aubintegri folia G. P. Ownbey
Eschscholzia minuti flora S. Wats.

Eschacholzia pariahii Greene
Plantaginaceae

Plantago ere eta Morris ssp. e recta
Plantago ovata Forsk. (P. faetigiata Morris)

Poaceae

Andropogon glomeratus (Walt.) B.S.P.

Ari8tida adscenaionis L.

Ariatida purpurea Nut. var. (Nees) A. & N. Holmgren (4.

glauca (Nees) Walp.)

*Avena fatua L.

*Bromus rubene L.

*Cynodon dactylon (L.) Pers.

Diplachne uninervia (Presl.) Parodi ( Leptochloa uninervia
(Presl) Hitchc. 6 Chase)

Echinochloa colonum (L.) Link
Erioneuron pulchellum (H.B.K.) Tateoka
Hilaria rigida (Thurb.) Benth. ex Scribn.

Muhlenbergia rigene (Benth.) Hitchc.

Phragmitea australis (C av.) Trin. ex Steud.

*Polypogon monapelieneie (L.) Desf.

*Schismua barbatua (L.) Thell.

*Vulpia myuroa (L. ) K. C. Gmelin var. hirauta Hack. ( Festuca
megalura Nutt.

Vulpia octo flora (Walt.) Rydb. ( Festuca octo flora Walt.)

- 11 -

Poleraoniaceae

Eriastrum denai folium (Benth.) Mason ssp. elongatum Benth.)

Mason

Gilia 8tellata Heller

Ipomop8i8 tenui folia (A. Gray) V. Grant
Linanthua joneaii (A. Gray) Greene
Polygonaceae

Choriaanthe brevioornu Torr. ssp. brevioornu

Eriogonum faaoioulatum Benth. ssp. poll folium (Benth.) S.

Stokes

Erigonum inflatum Torr. & Frem. ssp. inflatum
Eriogonum orouttianum S. Wats.

Eriogronum parishii S. Wats. ?

Eriogonum reni forme Torr. & Frem.

Eriogonum triohopes Torr.

Rumex aalioifoliua Weinm. ?

Rubiaceae

Galium atellatum Kell. ssp. eremioum (Hilend & Howell) Ehrend.
SAlicaceae

Populue fremontii S. Wats. var.

Salix exigua Nutt.

Salix gooddingii Ball var. variabilie Ball
Sapotaceae

Pouteria leuoophylla (S. Wats.) Cronq. ( Sideroxylon leucophyllum
S. Wats.

Scrophulariaceae

Caetilleja folioloea H. & A. ?

Caetilleja atenantha A. Gray

Mimulua guttatua Fisch. ex DC ssp. guttatua

Mimulua piloaue (Benth.) S. Wats.

Orobanohe oooperia (A. Gray) Heller
Penatemon alevelandii A. Gray ssp. olevelandii
Stemodia duranti folia (L.) Sw.

Selaginellaceae

Selaginella eremophila Maxon
Simmondsiaceae (not Buxaceae)

Simmondsia oheinenaia (Link) C. K. Schneid.

Solanaceae

Datura dieoolor Bernh.

Lyoium anderaonii A. Gray
Lyoium fremontii A. Gray ssp. fremontii
JUiootiana attenuata Torr. ex S. Wats, in King
Niaotiana trigonophylla Dunal in A. DC.

Phyealia oraaai folia Benth. var. oraaei folia
Solanum douglaeii Dunal in DC.

Sterculiaceae

Ayenia oompaota Rose
Tamaricaceae

*Tamarix ramoaaiaima
Thelypteridaceae

Thelypteria puberula (Baker) Morton var. aonorenaia A. Reid
Smith

Typhaceae

Typha domingenaia Pers.

Urticaceae

Parietaria heapera Hinton ssp. (P. floridana Auct.)

Urtica dioioa L. ssp. holoaerioea (Nutt.) Thorne
((/. holoaerioea Nutt.)

Viscaceae

Phoradendron oalifornioum Nutt.

Zygophyllaceae

Fagonia laevia Standi.

Larrea divarioata Cav. ssp. tridentata (Sesse & Moc. ex DC.)

Felger & Lowe. ( L . tridentata (Sesse & Moc. ex DC.) Cov.)

Robert F . Thorne
14 July 1986

- 12 -

Announcements from the May Board of Directors Meeting
I. Fall Symposium .

We are delighted to announce the speakers who will participate In the
13th Annual Symposium at U.C. Irvine on Saturday, November 14, 1987, entitled:
"Botanical Discoveries and Explorations 1 ^ Southern California "

The purpose of this symposium Is to emphasize the diversity of the
regional Southern California Flora and the impact this region has on botanical
sciences — the richness of this flora has Intrigued and attracted botanists
and researchers for over 200 years. Special focus will be placed on: A. sig-
nificant botanical explorations and Investigations which have greatly expanded
our knowledge of desert, clsmontane and island plant associations, B. the
vital role of our regional botanic gardens in supporting various research
programs on the native taxa, and C. current strategies designed to promote
public awareness of the esthetic values and potential usage of native plants
In our Increasingly urbanized environment.

The speakers will Include Dr. Mildred E. Mathias (Professor Emeritus,
Dept, of Biology, UCLA) who will talk about the fascinating evolution of
early botanic explorations and investigations which occurred In Southern
California; Dr. Tom Ellas (Director, Rancho Santa Ana Botanic Garden) will
speak on the Impact and expanding Importance our regional botanic gardens
have towards promoting public awareness of native species and botanical
research; Dr. Ralph Phil brick (Director, Santa Barbara Botanic Garden) will
emphasize the significance and history of the California Islands on current
theories of Island biogeography; Dr. Jon Keeley (Chairman, Dept, of Biology,
Occidental College) will cover various dynamics dealing with the wealth of
research In chaparral ecology; and Lloyd Smith (former Professor, Dept, of
Biology, Orange Coast College) will give Insights and personal accounts of
the Invaluable contributions made by Edmund C. Jaeger towards our knowledge
and appreciation of the unique beauty and botanic richness of southern
California deserts.

Additional Information regarding times of presentations, parking
regulations, etc. will be forthcoming In the August Issue of Crossosoma.

Geoff Smith, President
Allan Schoenherr, Editor

-13-

II. Spring Plant Sale : The April Plant Sale was very well -received, with S.C.B.
obtaining a net income of $1200 for combined sales and new membership!

A special thank you of appreciation goes to the following people for having
made the sale so successful:

Suppliers of plant material — Don Coughlin (Mockingbird Nursery, RiversideJ,
Mike Evans and Jeff Bohn (Tree of Life Nursery, San Juan Capistrano) and
Ray Walsh (Wildwood Nursery, La Verne)

Rancho Santa Ana Botanic Garden : Tom Elias (Director) and Linda Aberbom
(Propagation) for assistance in facilities availability
Publicity : Kathleen Pahl

Sale Set-up and Operation : Kathleen Pahl, Barry Prigge, Dave Bramlet,

Robert Thorne, Marvin Chesebro, Rick Strom, Shirley Mohr and Judy Smith
Sale Takedown: A1 Romspert, Barry Prigge and Judy Smith

COSPONSORED BY TM1 DEPT. OF ECOLOGY
AND EUOLUT1 ONARY BIOLOGY AND PETER
BOWLER, DIRECTOR OF THE COOPERATIVE
OUTDOOR PROGRAM IN THE STUDENT
ACTIVITIES OFFICE AT UC. IRVINE

13TH ANNUAL SYMPOSIUM

SOUTHERN i

CALIFORNIA

BOTANISTS

BOTANICAL DISCOVERIES
AND EXPLORATION IN
SOUTHERN CALIFORNIA

DATE: SATURDAY, NOUEMBER 14, 1987

time: 8:45 AM TO 4:00 PM ™7. ma

PLACE: HERITAGE ROOM, UNIVERSITY
CENTER; U.C., IRVINE

- 14 -

ANNOUNCEMENTS

SANTA MONICA MOUNTAINS WILDFLOWER WALK. SATURDAY, JUNE 13, 1987.

Why drive miles when wildf lowers abound locally? California
Native Plant Society naturalists lead 3-5 mile walks in different
locations based on best blooms. This is the sixth and last in a
series of biweekly walks cosponsored with the Sierra Club. Bring
water, lunch, stout walking shoes, hat, and camera. Meet at the
Federal Building in Westwood at 8:45 a.m. or at Taft High School
in Woodland Hills at 9:00 a.m. Walks usually end around 2 p.m.

For more information, call Jo Kitz, (818) 348-5910, or Linda
Hardie-Scott, (213) 479-1942.

WILDFLCWER TOURS OF THE BIG BEAR VALLEY PRESERVES, BIG BEAR LAKE ,
CALIFORNIA .

Big Bear Valley boasts the highest floral endemism for an area
of its size in California. Its Ice Age pebble plains, subalpine
meadows and limestone ridges provide habitats for 27 rare plant
species, 14 of which are restricted in the world to within the
Big Bear Ranger District and 2 to within the San Bernardino
Mountains. The Friends of Big Bear Valley Preserve have undertaken
an ambitious Spring and Summer schedule of fieldtrips to visit most
of the 15 preserve areas. We will see everything from the splendidly
beautiful and fragrant lemon lilies to the obscure little eye-strain
monkeyf lower — the tiniest monkeyf lower in the world I Tours are
by reservation only, $5 per person for non-Friends members. Proceeds
of the guided tour programs go directly toward fencing and protection
of the rare plant areas from off-road vehicles. Send $5 and a S.A.S.E.
to Friends of the Big Bear Valley Preserve, P. O. Box 1418, Sugar loaf
t CA 92386 for a copy of the Preserve Newsletter and Calendar, or call

Dave Hanson for reservation information at (818) 357-3542.

NATIVE PLANT SOCIETY ANNUAL PLANT SALE. SATURDAY, OCTOBER 17, 1987.

San Diego Chapter, CNPS, will hold its Annual Plant Sale in
Patio A of Balboa Park's Casa del Prado. There will be a fine
selection of California native plant materials for sale, as well as
plant posters, CNPS shirts, books, and pamphlets. People will be
available to answer questions concerning propagation and use of
native plants. For additional information, call Bobbie Steele
(619) 275-3732.

- 15 -

FIELD TRIPS

JUNE 6 ( SATURDAY ) . SAN MATEO MARSH . Meet at 9:30 a.m. at the
Lifeguard HQ building at top of Basilone Road offramp on 1-5, about
1 mile south of San Clemente. Celia Kutcher will lead us through
the varied habitats of a fresh water marsh that is now almost
unique in Southern California. Walk through the marsh in the a.m.,
eat lunch on the beach, look at Coastal Strand vegetation in the
afternoon on the way back to the cars. Bring lunch, water, sturdy
walking shoes; and be prepared for mosquitos, poison oak, and some
bushwhacking.

June 13 (Saturday) . Dry Lakes Ridge. The Channel Islands Chapter
of CNPS will host this trip. Dave Magney, Botanist, will lead a
strenuous walk in an area which is the subject of his flora,
published through UCSB Herbarium. Features disjunct stand of Pinus
ponderosa , recently burned chaparral, Boschniakia strobilacea (ground
cone) . Meet 8:30 a.m. at Ojai Safeway, jet. of Hwys 150 and 33.

Bring lunch, water, good boots. Ends in p.m. Contact: Dave Magney,
509 Palomar, Ojai, CA 93023, (805)646-7420; or Ron Wilkinson
(805)643-5007.

June 13-14 (Saturday -Sunday ) . Laguna Mountain . San Diego County,
Santa Monica Mountains and San Diego Chapters of CNPS will host this
field trip. Erik Jonsson, Botanist and member of San Diego Chapter
will lead a driving/walking tour of mountain flora. Meet 10 a.m.
Saturday at entrance to Laguna Camgoround, Cleveland National Forest,
Sunrise Hwy 18 mi. south of Julian. Those who wish may camp at
Laguna Camground, or if available may stay at Sierra Club facility,
reached via a dirt road through a gate off Sunrise Hwy. opposite
entrance to Laguna Campground. Ends 2 p.m. Sunday. Contact: Geoff

Burleigh, 602 N. Brand Blvd. , San Fernando, CA 91340, (818)361-1015;
or Eric Jonsson, 5148 Elnera Place, San Diego, CA 92117, (619)565-0548
JUNE 20-21 . ( SATURDAY -SUNDAY ) . TETRACOCCUS RIDGE , PAN AMI NT MOUNTAINS
People interested in participating on the field trip should try to
meet at the Thorndike Campground up Wildrose Canyon in the Panamint
Mountains (see AAA map of Death Valley) on Friday night or before

- 16 -

PANAMINT MOUNTAINS (CONTINUED)

0800 hours, Saturday morning. We will leave the campground at 0800
on Saturday morning and drive to Aguereberry Point (See AAA map of
Death Valley) . From Aguereberry Point it is a semi-arduous hike

and up to Tetracoccus Ridge, a Dolomite (limestone) outcropping,
outcropping provides a substrate for many endemics, most on the
Rare and Endangered Plant Inventory. In the past we have seen:
Bear Poppy - Arctomecon merriamii
Jointed Buckwheat - Eriogonum intrafractum
Death Valley Monkeyflower - Mimulus rupicola
Limestone Penstemon - Penstemon calcareus
Ripley's Gilia - Gilia ripleyi

Holly- leaved Tetracoccus - Tetracoccus ilicifolius
Naked- stemmed Daisy - Enceliopsis nudicaulis
Curled- leaved Mountain Mahogany - Cercocarpus intricatus
Panamint Mi lk- vetch - Astragalus panamintensis
Blazing Star - Mentzelia oreophila
Limestone Phacelia - Phacelia p_. perityloides
Sunday will be spent botanizing around Thorndike Camp (Water Canyon) ,
Mahogany Flats or the trail to Telescope Peak if time permits. It
can get cool in the evenings and there will be insects so come
prepared. For additional information contact Alan Romspert
(714)773-2428.

July 7-12 (Tuesday -Sunday) . Sierra Nevada-Stanislaus National Forest .
Calaveras and Alpine Counties and Santa Monica Mountains Chapters ,
CNPS, will tour an area of magnificent scenery and great diversity,
from 4500' to 9000' elevation. Plan to drive to Big Meadow Campground
on Tuesday, 4 full days to explore, return home Sunday. Expect to
include Calaveras Big Trees State Park, Bear Valley, Lake Alpine,
Mosquito Lake, Ebbetts Pass, Highland Lakes; all are accessible by
auto and short walks. Meet 7 p.m. Tuesday at Big Meadow Campground,
(USFS - fee req.) 20 mi. northeast of Arnold on SR-4 (Ebbetts Pass
Hwy) , about 85 mi. east of Stockton. Bring camping gear. Ends
10 a.m. Sunday. For information, write (please enclose (SASE) : Geoff
Burleigh, 602 N. Brand Blvd. , San Fernando, CA 91340, (818)361-1015.

down

This

CNPS

- 17 -

July 11 (Saturday) . Jenks Lake, San Bernardino Mountains . Andy Sanders
will lead this field trip in the forest near Barton Flats. Meet at
9:30 AM at the entrance to the Barton Flats Campground. For more
information call Dave Bramlett at (714) 855-0222.

July 18-19 (Saturday-Sunday) . White Mountains. Bristlecone Pine
and San Diego Chapters of CNPS will hosts weekend trip to see the
famous Bristlecone Pines. Camp in a primitive campground, so plan
to bring everything you need, and plenty of water . LIMIT: 30 persons,
reserve early. For reservations and information, call Maribeth
Kottman (619)445-6235, days.

July 26 (Sunday) . Newport Back Bay . Dave Bramlett will lead this
popular trip to see salt marsh vegetation at one of California's
few remaining estuaries. Interesting plants include salt marsh bird's
beak, Cordylanthus maritimus , and Yerba Mansa, Anemopsis califomica .
Meet at 9:00 AM at the intersection of San Joaquin Road and Back Bay
Drive. For details call Dave at (714) 855-0222.

August 22-23 ( Saturdav-Sunday) . Round Valley, San Jacinto State Park .
Mike Hamilton will lead this tour of the forest on Mount San Jacinto.
Meet at the station for the tramway at 10:00 AM. The fare is $11.00.
Lodging is available at the James Reserve. There is also a trip to
Garner Valley planned. For more details call Dave Bramlett at (714)
855-0222.

Plant Lovers of Southern California . The journal of the Southern
California Botanists, CROSSOSOMA, provides an ideal means by which
you can publish anything of botanical interest to southern Califorians.
Do you have a favorite field trips, gradening hints, or some pre-
liminary data that you.'d like to have in print? Submit it to:

Dr. Allan Schoenherr

Division of Biological Sciences, Fullerton College
321 E. Chapman Ave.

Fullerton, CA 92634

- 18 -

southern California botanists

Rancho Santa Ana Botanic Carden
l}00 North College Avrmit
Claremont. CA 9l?tl

The purpose of the SOUTHERN CALIFORNIA BOTANISTS Is the study,
preservation and conservation of the native plants of California; and
the education of the public to the value of the native flora and Its
habitats. It is a non-profit association forced in 1927.

Membership benefits include:

Various field trips throughout the state led by competent
field botanists aad biologists.

A yearly plant sale featuring native California species.

An annual symposium on various aspects of the California
vegetation .

The SCB Journal. CROSSOSOMA.

Discounts on botanical and natural history books.

Dues are for a calendar year. Nee members Joining from May through
September, please deduct $1.00 from your dues. Those Joining in October
through December are credited with the following year's dues.

Membership categories are:

/ / Individual* $ a.00

/ / Croup or organization $15.00

*Includes membership for entire family

/

/

New Member

/

Renewal

Date

Name

Address _____ - --

City Zip Code

Phon ej )

In addition. I want to give $ to help support SCB.

Make check payable to: SOUTHERN CALIFORNIA BOTANISTS.

Mall check and form to:

Alan P. Romspert
Southern California Botanists
Department of Biological Science
California State University. Fullerton
Fullerton. CA 92634

CROSSOSOMA (ISSN 0891-9100) Is published bimonthly (February, April, June, August,
October, and December) by Southern California Botanists, a California non-profit
corporation. Back issues of CROSSOSOMA are available for $2.00 an Issue (plus 25*
postage) or $8.00 a volume (plus $1.00 postage). Send a check with your request to
Alan P. Romspert, Treasurer, at the above address. Manuscripts submitted for
publication should be addressed to Dr. Allan A. Schoenherr, Editor of CROSSOSOMA,
Division of Biological Sciences, Fullerton College, Fullerton, CA 92634.

- 19 —

SCB COMING EVENTS (DETAILS WITHIN )

June 6

San Mateo Marsh

June 13

Dry Lakes Ridge

June 13,

14

Laguna Mountains

June 20,

21

Panamlnt Mountains

July 7-12

Sierra Nevada

July 11

San Bernardino Mountains

July 18,

19

White Mountains

July 26

Newport Back Bay

August 22,23

San Jacinto Mountains

November

14

13th. Annual Symposium

111

DBF? ARY

UUN 1 5 1987 g

I'V . , v |

BOTANICAL garden

xo /

■ /?7^
vJ3

#4-

CROSSOSOMA

southern cal iforni a botanists

Rancho Santa Ana Botanic Garden, Claremont, CA 91 71 1

CR0SS0S0MA Vol . 13, No. 4 August, 1987

Managing Editor: Allan Scboenherr

A BRILL NOTE ON

THE POLLINATION ECOLOGY OF TWO PALMS:

ONE FLORIDIAN AND ONE CALIFORNIAN

Scott Zona

Rancho Santa Ana Botanic Garden
1500 North College Avenue
| Claremont, CA 91711

Recent tield studies on the pollination ecology of palms
have revealed a variety of previously unsuspected pollination
mechanisms. Traditionally, palms were thought to be primarily
anemophilous (wind-pollinated) (Henderson, 1986). However,
studies in several genera have shown a great diversity of biotic
pollen vectors, including bees (Brown, 1976), flies (Schmid,

1970), beetles (Henderson, 1984), and bats (Beach, 1986). This
study investigated the pollination ecology of two species of fan-
palms, one tloridian and one Californian.

The genus Sabal (Palmae: Coryphoi deae) consists of
approximately twenty species that occur in and around the
Caribbean Basin, including two species from Sonora, Chihuahua,
Sinaloa, and Nayarit, Mexico (Bailey, 1944): S. rosei (Cook)

Becc. and S. uresana Trelease. A fossil Sabal is known from
Miocene and Pliocene deposits in Southern California (Axelrod,
1975) .

Sabal etonia Swingle ex Nash, a Florida endemic, is found in
xeric habitats on the Central Florida Ridge and the Atlantic

<

LIBRARY

AUG 31 «87

N cvv tORK

BOTANICAL GARDE®

coastal Ridge (Zona 6 Judd, 1986). It is commonly tound in
association with xerophytic pines ( P i nus clausa (Chapm. ex

Lnglm.) Vasey ex Sarg. or P. ell lott 1 1 Englm. var. densa Little &
Doorman) and several species of xerophytic oaks (viz., Que reus
gem inata Small, 0* my rt i tol ia Vv i 1 Id., and 0- inopi na Ashe). This
study, however, was conducted in an area in Palm beach County
along the Atlantic Coastal Ridge where the pine canopy is absent.

Sabal eton la blossoms trom late May through July. The
t lowers are small (ca. 6 mm across), white, fragrant, and
hermaphroditic. They tunction for only one day. Odor production
occurs trom approximately 03:00 and ceases between 18:20 and
20:00 (Eastern Standard Time equivalent). Nectar is produced
trom approximately 05:00 to 14:20. Flowers are slightly
protandrous, with tirst anther dehiscence observed at 07:20.
Stigmas become receptive around 09:50. The principle pollinators
are bees (Hymenoptera : various families) (Table 1), but syrphid
tlies (Diptera: Syrphidae) may also tunction as pollen vectors.

A detailed decription ot phenology and pollination will appear
elsewhere (Zona, in press).

Table 1. bees (Hymenoptera) visiting Saba 1 etonia and bearing s.
eton la pollen in Palm Beach County, Florida (from Zona, in press).

Collet idae

Col letes mand ibular is F . Smith
Ant hophor idae

Xy locopa micans Le pe 1 e t i e r
Ap l dae

Ap i s mei 1 l ter a L.

Megach 1 1 idae

Megach i le albitarsis Cresson
M . xylocopoides F . Smith
Coe 1 loxys aster is Crawford
Ha 1 ict idae

A^ apostemon splendens Lepelet ier
Auqoc hi props is metal 1 ica (Fabricius)

2

Brown (1976) reported that a very closely related species,
Sabal palmetto (Walt.) Lodd. ex J. A. 6 J. H. Schultes, is
protogynous. While protogyny may represent a specific difference
between S. palmetto and S. etonia , it appears more likely that
protandry is characteristic of both species. Brown suggested
that pollination occurs when early morning nectar-foragers brush
stigmas with residual pollen accumulated on the previous day.

The honeybee ( Apis mell i tera ) , one of the primary pollinators
named by Brown, sometimes exhibits a division of labor among
workers in that some workers collect pollen and others collect
nectar (tree, 1977). If this were true, then pollen loads of
nec ta r- f orag i ng honeybees would be, in fact, accidental pollen
loads, and the loads might be reduced further after a night in
the hive. Pollination would thus seem an uncertain proposition
tor a protogynous S. palmetto . Bees (including the honeybee)
foraging for pollen on protandrous S. etonia carry conspicuous
pollen loads and readily contact the stigma while crawling over
the flowers. The bees do not appear to collect nectar from S.
etonia . Protandry is more typical for bee-pollinated palms
(Henderson, 1986), and as the floral morphologies of S. palmetto
and S. etonia are nearly identical, the exceptional condition of
s. palmetto warrants reinvestigation.

Washingtonia (Palmae: Coryphoideae) is a genus of two
species. The familiar W. t i 1 i fera (Linden) H. Wendl. is native
to Southern California, Arizona, and Northern Baja California.

The commonly cultivated W. robust a H. Wendl. is found farther
south in Baja California. Only preliminary observations of the
floral biology of Washingtonia t il ifera have been made at Rancho
Santa Ana Botanic Garden. The flowers are white, fragrant, and
hermaphroditic. The sepals are connate, tubiform, ca. 5 mm long.
The petals are ca. 6 mm long and are reflexed for half their
length. The stamens and gynoecium are approximately 6 mm long.

The pollen is sticky, and sampling recovered no air-borne
pollen even when sampling devices (glycerine jelly-coated glass
slides) were only a tew centimeters away from the flowers.
Anatomical investigations revealed the presence of septal

3

nectaries at the base ot the gynoecium. The phenology ot W.
t 1 1 i tera is not known.

At the Botanic Garden, tlowers ot W. t i 1 i tera are actively
visited by non-native pollen-gathering honeybees ( Apis mel 1 i tera )
and other Hymenoptera. Curiously# temale Anna's hummingbirds,
Calypte anna # were observed toraging on W. t i 1 i tera in August
when tew other nectar-producing tlowers are available in the
botanic Carden. The significance ot hummingbirds as "beak tip"
pollinators is not known.

Neither Sabal etonia nor Wash i ngtonia f i 1 i f era are wind
pollinated; both appear to be primarily bee pollinated with many,
but not all, ot the characteristics of the "bee pollination
syndrome" described by Henderson (1986). These characteristics
include protandry, tew stamens, early morning stamen dehiscence,
nectar production, and fragrance.

Acknowledgements

I am grateful to Roy R. Snell ing, Los Angeles County Museum
ot Natural History, for providing bee identifications. This work
benetitted trom discussion and criticism provided by Andrew
Henderson and Kay Tomlinson. A portion ot this study was funded
in part by a grant from Southern California Botanists.

Literature Cited

Axelrod, D. I. 1975. Evolution and biogeogrphy ot Madrean-

Tethyan scierophyll vegetation. Ann. Missouri bot. Card.

62; 28U-334.

bailey, L. n. 1944. Revision ot the palmettoes. Gentes Herb.
6(7); 365-459.

beach, J. H. 1986. Pollination biology of spadices and spicate
inflorescences in Cyclanthaceae and Palmae. American Jour.
Bot. 73(5); 615, 616 (abstract),
brown, K. E. 1976. Ecological studies ot the cabbage palm,

Saba 1 palmetto . Principes 20(1); 3-10.
free, J. b. 1977. The Social Organization ot Honeybees. Edward
Arnolu, London. 68 pp.

4

Henderson, A.

1984.

Observations on pollination of Chryosoph 1 la

a_lbida. Principes 28(3): 12U-126.

Henderson, A. 1986. A review of palm pollination studies. Bot.
Kev. 52(3): 221-259.

Schmid, R. 1970. Notes on the reproductive biology of

Asterogyne mart iana (Palmae). II. Pollination by syrphid
tlies. Principes 14(2): 39-49.

Zona, S. In press. Phenology and pollination biology ot

Sabal e ton ia (Palmae) in southeastern Florida. Principes.

Zona, S. & W. S. Judd. 1986. Saba 1 e ton ia (Palmae):

systematics, distribution, ecology, and comparisons to other
Florida scrub endemics. Sida 11(4): 417-427.

BAJA CALIFORNIA FLORAS

Or. Robert Thorne of Rancho Santa Ana Botanic Garden is composing
a flora for the northern portion of Baja California. As a product of
that effort he has compiled a series of valuable checklists for
selected localities in Baja California. The following list of vas-
cular plants is the second of a series that will be published in
CR0SS0S0MA

VASCULAR PLANTS OF PASO SAN MAT IAS AND VALLE DE TRINIDAD,

BAJA CALIFORNIA, MEXICO
by

Robert F. Thorne
Rancho Santa Ana Botanic Garden
Claremont, California

List of vascular plant species observed or collected in or above
the Canon and Canada de San Matias between the Sierras de Juarez
and San Pedro Martir, mostly from the head of the pass to the
Ejido San Matias, with additional Sonoran Desert (Lower Colorado
Desert) species (T) found farther west in the Valle de Trinidad
in the Sonoran mixed desert scrub and microphyll woodland. The
elevation at the head of the pass alonq Hwy 3 is ca . 900 m near
31° 19’ N, 115* 30' W; the elevation at the foot of the pass is
ca. 700 m near 31° 18’ N, 115 25' W. Species found only lower

in the pass and not yet collected farther west are included in
brackets. Species believed to be introduced are preceded by an
asterisk. Vouchers collected by R. F. Thorne and by R. V. Moran
are mostly filed at RSA, SD, MEXU, and BCMEX.

ACANTHACEAE

Jueticia cali forn i ca (Benth.) D. Gibson (Be leperone cal i form' s i

Benth .

ADIANTACEAE

A >li tin turn capi 1 1 us- veneri s L. - T

(’he i l ant hen so vi l lei Maxon

Che i l an t he a desert i Mickel ( Not holaena aalifornica D. C. Eat.)
Nothalaena sinuata (lag.) Kaulf.

Felluea mucronata (D. C. Eat.) D. C. Eat. ssp. mucronata
Pi tyrogramm triangularis (Kaulf.) Maxon ssp. maxonii (Weath.)
ined .

AGAVACEAE

Agave deaerti Engelm. ssp. pringlei (Engelm. ex Trel.) Gentry
yut?.?a schidigera Roe z 1 ex Ortlqies
ALLIACEAE

Allium haematochi ton S. Wats.

AMARANTH ACE AE

Amaranthua fimbriatua (Torr.) Benth. ex S. Wats.

Amaranthua palmeri S. Wats.

APOCYNANCEAE (incl. Asclepi adoideae )

Mate lea paroi folia (Torr.) Woods.)

Carcoatemma cynanchoides Dene. ssp. hartvegii (Vail) R. Holm
ANACARDI ACE AE

Rhus trilobata Nutt, ex T. ft G. var. ai mplici folia (Greene)
Barkley

Rhus ouata S. Wats. - T.

ARALI ACEAE (incl. Apioideae)

Gpermo Lepi a echinatus (Nutt.) Heller
ASTERACEAE

Ambrosia campho rata (Greene) Payne - T
Ambrosia con ferti flora DC.

Ambrosia Jumoaa (A. Gray) Payne
Ambrosia flexuosa (A. Gray) Payne
Ambrosia pumila (Nutt.) A. Gray
Ani so coma acaulis T. ft G.

Artemisia ludoriciana Nutt. ssp. albula (Woot . ) Keck

Haocharis brachyphylla A. Gray - T

Hailey a p lent rad iata Harv. ft Gray

Hebbia Junes a (Benth.) Greene - T

Bricks Ilia frutescene A. Gray

Calycoseri a parry i A. Gray

Chaenactie ,jl abri uscula DC. var. tenufolia (Nutt.) Hall
Ciraium occidentals (Nutt.) Jeps.

Pyssodia porophyl loide a A. Gray

Fncelia farinoea A. Gray ex Torr. var. farinosa
t'rigeron di oe rgens T. ft G. - T
t'riophyllum vail ace i A. Gray
Filago aalifornica Nutt. - T
Geraea viacida (A. Gray) Blake

Gutierrezia aarothra, (Pursh) Britt. ft Rusby - T
Haplopappua linear i fo l i us DC
Haplopappua p ropinquua Blake
Haplopappua acabrellua Greene

He lianthua pe tiolaria Nutt. ssp. fallax Heiser - T
He te ro theca grandi flora Nutt. - T
Hy me hoc lea monogyra T. ft G. ex A. Gray - T
Hymenoclea salaola T. ft G. ex A. Gray ssp. salsola
iayia glanduloea (Hook.) Hook, ft Arn. ssp. glandules a
Lepi dospart urn aquamatum (A. Gray) A. Gray

Machaeranthera asteroi des (Torr.) Greene ssp. lagunensi s (Keck)
ined. ( M . tephrodes (A. Gray) Greene)

Machaeranthera juncea (Greene) Hartman (Haplopappua junceus
Greene)

Malacothrix g lab rat a (A. Gray) A. Gray
Malacothrix a tebbinaii Davis ft Raven

Microaeria lindleyi (DC.) A. Gray. [M. lineari fo lia (Nutt.)

Sch. Bip.)

Palafoxia arida Turner ft Morris ssp. arida
Pectis pappoaa Harv. ft Gray ex. A. Gray
Porophyl lum gracile Benth.

Rafinesquia neomexicana A. Gray
Cenecio lemmonii A. Gray
l ephanome ri a pauci flora (Torr.) A. Nels.

!• t y locline micropoidea A. Gray

Trixis aalifornica Kell. var. aalifornica

Vi gui era parishii Greene

BIGNONI ACEAE .

Chilopsis linearis (D. Don) Sweet ssp. arcuata (Fosb.) Henricks.

b

BORAGINACEAE

Amsinckia intermedia F. i M.

Crypt ant-ha angusti folia (Torr.) Greene
I'ryptantha barbi gera (A. Gray) Gr-ene
i’ryptantha decipiens (Jones) Heller
Cryptantha maritima (Greene) Greene

Crypantha micrantha (Torr.) Jtn. ssp. lepida (A. Gray) Mathew
& Raven

Cryptantha pterocarya (Torr.) Greene [incl. var. cyclop tera
(Greene) Macbr.)

Harpagone l la palmeri A. Gray - T

Pe a ternary a linearis (R. fc P.) DC ssp. ferocula (Jtn.) Thorne
Fectocarya penicillata (H. 6 A.) A. DC. - T
Fectocarya recurvata Jtn.

Tiqui lia plicata (Torr.) A. Richards. - T
BRASSICACEAE

Arabia perennane S. Wats.

•Hrasaiaa tournefortii Gouan
[Caulanthus cooperi (S. Wats.) Pays.]

Caulanthus last ophyl lum (H. 6 A.) Pays.

Pescurainia pinnata (Walt.) Britt, ssp. halictorum (Ckll.)

Detl.

Di thy re a cali fornica Harv.

Draba cunei folia Nutt, ex T. & G. var. i ntegri folia S. Wats.
Lepidium lae iocarpum Nutt, ex T. & G. var. lasiocarpum
Lyrocarpa coulteri Hook. & Harv. var. palmeri (S. Wats.) Roll.
* 0 i aymbri urn irio L.

The ly podium wrighti i A. Gray

Thysanocarpus curvipes Hook. var. elegans (F. i M.) Robins, in
A. Gray
BURSERACEAE

[Burs era miorophylla A. Gray)

C ACT ACE AE

Echinoce reua enge Imannii (Parry) Ruempler var. engelmanni i
Fe rocactus acanthodee (Lemaire) Britt, fc Rose
bophocereus schottii (Englem.) Britt, fc Rose var. schottii
Mammillaria lioica K. Bdg.

Opuntia acanthocarpa Engelm. & Bigel. in Englem.

Opuntia bi gelovii Engelm. var. bigelovii
Opuntia chlorotica Engelm. & Bigelo in Engelm.

Opuntia eohinocarpa Engelm. fc Bigel. var. echinocarpa
Opuntia prolifera Engelm.

[Opuntia teaajo Engelm. ex Coulter)

Fachy ce re ue pringlei (S. Wats.) Britt. & Rose
CAMPANULACEAE

[Nemalcadua glanduli fe rue Jeps. (incl. var. orientahs McVaugh]
Nemacladus pinnatifidus Greene - T
CARYOPH Y LLACE AE

Arenaria douglaeii (Fenzl ex T. 4 G.) Mattfeld - T
CHENOPODIACEAE

Atriplex caneocena (Pursh) Murr. ssp. canescen3
Atriplex polycarpa (Torr.) S. Wats.

Sal 8o la australis R. Br. (S. kali L. var. tenui folia Tausch)
CONVOLVULACEAE

Evovulus alsinoide8 L. var. aaapulcensis (Willd.) v. Ostrstr.
CRASSULACEAE

Dudleya ari zoni ca Rose

Dudleya pauci flora Rose in Britt. & Rose
CUCURBITACEAE

Cucurbita palmata S. Wats.

CUPRESSACEAE

Juniperua cali fornica Carr.

DRACAENACEAE

Nolina bigelovii (Torr.) S. Wat.

EPHEDRACEAE

Ephedra cali fornica S. Wats.

EUPHORBI ACEAE

Acalypha cal i fornica Benth.

Argythiimnia lanceolate (Benth.) Muell.-Arg. [Pi taxis lanceolat i
Pax. & K. Hoffm.)

Argy thamni a neomexicana Muell.-Arg. [Di taxis neomexicana
(Muell.-Arg.) Heller]

7

Bernards a incana Morton

Chamaeeyce albomarginati, (T. i G.) Small (buphorbic

albomarqinata T. fc G. ) . j„.;.iTnrr

Chamaesyce melanadenia (Torr . ) Millsp. ( Euphorbia m» lanadeni artozr ■

Chamaesyce micromera (Boiss.) Woot. i Standi. (Euphorbia

mi crone ra Boise.) . , . . . ln

Chamaesyce polycarpa (Benth.) Millsp. in Parish yar hirtella
(Boiss. ) Millsp. ( Euphorbia polycarpa var. hirtella Boiss.)
Chamaesyce eerpylli folia (Pers. ) Small. {Euphorbia
eervulli folia Pers.) . „ „ _ .

Chamaesyce eetiloba (Enqelm. ex Torr.) Millsp. in Parish

(Euvhorbia setiloba Eng®!**** Torr.) .

Chamaesyce tomentuloea (S. Wats.) Millsp. (Euphorbia tomentuloea

S. Wats.

Croton cali fornioue Muell.-Arg.
Euphorbia eriantha Benth.

StilH npia lineari folia S
FABACEAE

Wats.

Acacia greggii A. Gray ssp. arizonica (Isely) ined.

As t ragalus acuti roetri e S. Wats.

Astragalus gambelianus Sheld. - T
Astragalus palmeri A. Gray
Calliandra eriophylla Benth.

Cercidium microphyllum (Torr.) Rose fc Jtn.

Palea bicolor Humb. fc Bonpl . in Willd. ssp. seemannii.

ined. {Dalea eeemannii S. Wats. ssp. eeemanni i )

Dalea mollis Benth. ssp.
lotus argyraeue (Greene) Greene - T
lotus humietratus Greene - T
lotus ri jidus (Benth.) Greene

lotus sa suginosus Greene ssp. brevi vexi Hue (Ottley) Munz
Lotus BC'<:>arius (Nutt, in T. «. G.) Ottley ssp. brevialatus

(S. Wats.)

(Ottley) Munz . , t

Lotus strigosus (Nutt, in T. f. G.) Greene var. hirtella
(Greene) Ottley

[Lotus strigosus (Nutt, in T. & G.) Greene var. tomentellue
(Greene Isely)

Lupinus concinnus Agardh ssp. orcuttii (S. Wats.) D. Dunn
Lupinus sparei florus Benth. ssp. sparsi florus

Marina orcuttii (S. Wats.) Barneby. f Dalea orcuttii S. Wats.)
Marina parrot (T. t G.) Barneby. ( Dalea parryi T. & G.)

[Olneya teeota A. Gray]

*Fark insonia aculcata L. - T

Proeopis glandulosa ’Torr. ssp. torreyana (L. Bens.) E. Murray
Psorothamnus echottii (Torr.) Barneby ( Dalea echottii Torr.)
Senna coveeii (A. Gray) Irwin i Barneby ( Cassia covesir A. Gray)
FAGACEAE

Quercus turbinella Greene ssp. turbinella
F0UQUIEF1ACEAE

Fouquieria splendens Engelm. ssp. splendens
GERANI ACEAE

*Erodium cicutarium (L.) L'Her.

GROSSU LARI ACEAE

Ribes querce torum Greene
HYDRO PH YLLACEAE

Emmenanthe penduli flora Benth.

Eriodictyon anguetifolium Nutt. - T

Eucrypta chrysanthemi f olia (Benth.) Greene var. bipinnati fida
(Torr.) Const.

Eucrypta micrantha (Torr.) Heller
I Mama demiesum A. Gray

A lama hiepidum A. Gray var. revolutum Jeps. - T
Phacelia affinis A. Gray
Phacelia cryptantha Greene
Phacelia dietane Benth.

Phacelia pedicellata A. Gray
Phacelia euf f rutescens (Parry) Const.

KRAMERI ACEAE

Kramer ia grayi Rose fc Painter
Krameria parvifolia Benth. var.

LAMI ACEAE

l Hyptis emoryi Torr.)

Salvia apiana Jeps. var. apiana

Salvia columbariae Benth. ssp. columbariae

8

LOASACEAE

Mentzelia albicaulis (Dougl . ex Hook.) T. & G.
Petalonyx thurberi A. Gray ssp. thurberi - T

MALPIGHIACEAE

Janus ia gracilis A. Gray

incanum (Link.) Sweet ssp. pringlei (Hochr.) Felger 4
Lowe (4. pringlei Hochr.)!

Heri ssantia crispa (L.) Bnzicky
Hibiscus coul teri Harv. ex A. Gray
Hibiscus denudatus Benth.

[Hors fordid neuberryi (S. Wats.) A. Grayl K^arnev

Jphaeralcea ambigua A. Gray ssp. rosacea (Munz fc Jtn.) Kearney

MARTYNIACEAE _

Proboscidea parvi flora (Woot.) Woot. f* Standi. - T

MOLLUGINACEAE

*Hollugo cerviana (L.) Ser .

NYCTAGINACEAE

Abronia villose S. Wats. var. vtllosa
Allionia incarnuta L.

Boerhavia coulteri (Hook, f.) S. Wats.

Boevhaoia diffusa L. (fl. coccinea Mill.)

Boerhaoia intermedia Jones

Boerhavia spicata Choisy

Commi carpus scandens (L.) Standi.

Mirabilio bigelovii A. Gray (incl. vars. aspera and retrorsa)
OLEACEAE

Fo rest i e ra neomexicana A. Gray
Menodora scoparia Engelm. ex A. Gray

ONAGRACEAE

Camisaonia californica (Nutt, ex T. i G.) Raven
Cam i a son i a chamaene rioide s (A. Gray) Raven
Camissona pallida (Abrams) Raven ssp. pallida
Camio sonia strigulosa (F. 4. M.) Raven

OXALI DACEAE

i >xa li s albicans H.B.K. ssp .cali form ca (Abrams) Eiten
PAPAVERACEAE

Fschsaholzia minutiflora S. Wats.

PINACEAE

Finus monophylla Torr. 6 Englem.

PLANT AG 1 NACE AE

Vlantago patagonica Jacq. var. spinulosa (Dene.) A. Gray
{Plant ago purshii R. i S. var. oblonga (Morris) Shinners]

POACEAE

Aristida adscensionis L.

[Aristida californica Thurb.J
Aristida g lab rat a (Vasey) Hitchc.

Aristida urightii Nash var.

(A. pari8hii Hitchc.) . ... .

Bothrioehloa barbinodis (Lag.) Herter ( Andropogon barbinodis Lag.)
Bouteloua aristidoides (H.B.K.) Griseb.

Bouteloua barbata Lag.

Bouteloue curtipendula (Michx.) Torr. in Marcy
[Bouteloua trifida Thurb. in S. Wats.)

Brachiaria arizonica (Scribn. & Merr.) S. T. Blake
*Bromu8 rubens L. .

Digitaria californica (Benth.) Henr. [Trichachne californica

(Benth.) Henr.l

*Eragrostis cilianenais (All.) Mosher
Erioneuron pulchellum (H.B.K.) Tateoka [Tridens pulchellus
(H.B.K.) Hitchc. |

He teropogon contortus (L.) Beauv. ex R. i S .

MeLica frutescens Scribn.

Muhlenbe rgi a microsperma (DC.) Kunth
Muhlenbe rgi a porteri Scribn. ex Beal
Pan i cum hirtioaule Presl .

*Echismus barbatus (L.) Thell.

Setaria leucopila (Scribn. & Merr.) K. Schumann
3 porobolus contractus Hitchc.

Cporobolus crypt an dr us (Torr.) A. Gray
t i pa parishii Vasey
Ctipa speciosa Trin. u Rupr .

Tridens muticus (Torr.) Nash in Small
*Vul-pia bromoi lea (L.) S. F. Gray [Festuca dertonens is (All.)

Asch . & Graebn . |

Vulpia octo flora (Walt.) Rydb . [Festuca octo flora Walt)

9

ssp. brevicornu
var. f imbri at a

POLEMON I ACEAE

Eriaotrum eremicum (Jeps.) Mason
uili > i diegenaia (Munz) A. 4 V. Grant
it ilia ate l lata Heller

Ipomopaia tenui folia (A. Gray) V. Grant
Linanthue floribundus (A. Gray) Greene ex Milkn ssp.

R. Patterson

Linanthue joneai i (A. Gray) Greene
POLYGALACEAE

Vo l yga l a macradeni a A. Gray
POLYGON ACEAE

Chorizanthe brevi cornu Torr .

Chorizanthe fimbriata Nutt.

Eriogonum davidsonii Greene
Eriogonum elongatum Benth. . .

Eriogonum fasciculatum Benth. ssp. poll folium (Benth.) S.

Stokes

Eriogonum nudum Dougl . ex Benth. ssp. pauciflorum (S. Wats.)

Munz

Eriogonum piloaum S. Stokes
Eriogonum thurberi Torr.

Eriogonum x jrightii Torr. ex Benth. ssp. membranaceum S. Stokes
Oxy theca trilobata A. Gray
Pteroategia drumarioides F. 4 M. - T
Humex hymenosepa l us Torr.

PORTULACACEAE

('alyptridium monandrum Nutt, in T. 4 G.

*l*o rt ulaca oleracea L.

Po rtulaca parvula A. Gray

RANUNCULACEAE L , . , t . .

Delphinium pariahii A. Gray ssp. subgloboaum (Wigq.) Lewis 4

Epl ing
RHAMN ACEAE

Zizyphus obtuoi folia (Hook.) A. Gray ssp. aaneaacn a (A. Gray)
ined. [Condal iops i 6 lycioidea (A. Gray) Suesseng. var.
caneacena (A. Gray) Suesseng)

ROSACEAE

Prunus / aaciculata (Torr.) A. Gray
Prunua fremontii S. Wats. — T
RUTACEAE

Thamnosma montana Torr. 4 Frem.

SALICACEAE

Haliz exigua Nutt. - T

Salix laevigata Bebb var. araquipa (Jeps.) Ball
S C ROPH U L A R I AC E AE

Caetil leja folioloea H. 4 A. - T . , , .

Keckiella antirrhinoidee (Benth.) Straw ssp. microphylla (A.
Gray) Straw

Mimulue fremontii (Benth.) A. Gray
renatemon cent ranthi fo li ua Benth.

Penatemon clevelandii A. Gray ssp. Cleveland it
Penstemon eximiua Keck

Penatemon thurberi Torr. ssp. thurberi
SELAGINELLACEAE

He l agine lla eremophila Maxon
S IMMONDSI ACEAE „ „ . . .

immonda ia chineneia (Link) C. K. Schneid.

SOLANACEAE

bycium anderaonii A. Gray

Lycium pallidum Miers - T

Niaoti ana trigonophylla Dunal in A. DC

Physalia aouti folia (Miers.) Sandwith

Phyaalie crascifolia Benth. var. infundibularis I. M. Jtn.

*S ’olanum americanum Mill. (5. nodiflorum Jacq.)
liol anum douglaaii Dunal in DC.

STERCU LI ACEAE

Ayenia compact** Rose

Fremont odendron cali fornicum (Torr.) Cov.

VERBENACEAE

Verbena abramaii Moldenke

boUanum (Seem.) Eichler ssp. (Torr.) Wiens

Fhoradenlron cali fornicum Nutt.

ZYGOPHYLLACEAE „ . .

Kallatroemia California (S. Wats.) Vail
bar re a divaricate Cav. ssp. tridantata (Sesse i Moc. e
Felger 4 Lowe [L. tridentata (Sesse 4 Moc ex DC) Cov. I
( Vir.cainoa geniculate (Kell.) Greene var. geniculata \

ANNOUNCEMENTS

I. Fall Symposium , at U.C. Irvine on Saturday, November 14, 1987
" Botanical Discoveries and Explorations in Southern California "

The purpose of this symposium Is to emphasize the diversity of the
regional Southern California Flora and the impact this region has on botanical
sciences -- the richness of this flora has Intrigued and attracted botanists
and researchers for over 200 years. Special focus will be placed on: A. sig-
nificant botanical explorations and Investigations which have greatly expanded
our knowledge of desert, cismontane and island plant associations, B. the
vital role of our regional botanic gardens in supporting various research
programs on the native taxa, and C. current strategies designed to promote
public awareness of the esthetic values and potential usage of native plants
in our increasingly urbanized environment.

The speakers will include Dr. Mildred E. Mathias (Professor Emeritus,
Dept, of Biology, UCLA) who will talk about the fascinating evolution of
early botanic explorations and investigations which occurred In Southern
California; Dr. Tom Ellas (Director, Rancho Santa Ana Botanic Garden) will
speak on the impact and expanding importance our regional botanic gardens
have towards promoting public awareness of native species and botanical
research; Dr. Ralph Philbrick (Director, Santa Barbara Botanic Garden) will
emphasize the significance and history of the California Islands on current
theories of island biogeography; Dr. Jon Keeley (Chairman, Dept, of Biology,
Occidental College) will cover various dynamics dealing with the wealth of
research in chaparral ecology; and Lloyd Smith (former Professor, Dept, of
Biology, Orange Coast College) will give insights and personal accounts of
the invaluable contributions made by Edmund C. Jaeger towards our knowledge
and appreciation of the unique beauty and botanic richness of southern
California deserts.

NATIVE PLANT SOCIETY ANNUAL PLANT SALE. SATURDAY , OCTOBER 17, 1987^
San Diego Chapter, CNPS, will hold its Annual Plant Sale in
Patio A of Balboa Park's Casa del Prado. There will be a fine
selection of California native plant materials for sale, as well as
plant posters, CNPS shirts, books, and pamphlets. People will be
available to answer questions concerning propagation and use of
native plants. For additional information, call Bobbie Steele
(619) 275-3732.

1 1

field trips

August 22-23 ( Saturdav-Sunday ) . Round Valley, San Jacinto State Park .
Mike Hamilton will lead this tour of the forest on Mount San Jacinto.
Meet at the station for the tramway at 10:00 AM. The fare is $11.00.
Lodqing is* available at the James Reserve. There is also a trio to
(lamer valley olanned. For more detail* call Dave Bramlntt at (714)
855-0222.

August 29-30 (Saturday-Sunday ) . CONE-B f.aring trees of THE
SOUTHERN SIERRA NEVADA . Allan Schoenherr will lead this motor-camoinq
tour. The trip begin? at Freeman Grove, one of the southernmost
grove? of old qrowth sierra redwood? (Giant Sequoia), that i? located
on the Dlateau west of the Kern River. The route will Droceed down
to the Kern River, then over the “astern divide at Sherman Pas? to
camo in a picturesque forest-set t inq west of Kennedy Meadows. The
route will continue throuqh Kennedy Meadows and down the spectacular
east side of the Sierra Nevada. We should see at least 13 soecies
of coniferous trees, including 7 specie* of pine?. While the
emphasis will be on conifers, many species of mountain wild flowers
'••hould be in bloom. For details, call Allan at ( 714) 992- 71 29

weekdays or at (714) 494-0675 evenings.

Plant Lovers of Southern California . The journal of the Southern
California Botanists, CROSSOSOMA, provides an ideal means by which
yon can publish anything of botanical interest to southern Califotian
Do you have a favorite field trips, gradeninq hints, or some pre-
liminary data that you'd like to have in print? Submit it fro:

Dr. Allan Schoenherr

Division of Biological Sciences, Fullerton College
321 E. Chaoman Ave.

Fullerton, CA 92634

1 2

cosponsored by the dept oe ecology

iiND KUOillTlONARV BIOLOW AND PETEK
BOWLER, DIRECTOR OF THE COOPERATIUE
OUTDOOR PROGRAM IN THE STUDENT
ACTIVITIES OFFICE AT IIC.IKVINE

SOUTHERN
CALIFORNIA

13TH ANNUAL SYMPOSIUM BOTANISTS

BOTANICAL DISCOVERIES
AND EXPLORATION IN
SOUTHERN CALIFORNIA

DATE: SATURDAY, NOVEMBER 14, 1987

time: 8:45 AM TO 4:08 PM “7...

PLACE : HERITAGE ROOM, UNIVERSITY
CENTER; U.C., IRVINE

r

^ '■* "T'f

Parking

v

University Center

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\

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,i «?y , * - - ^

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1 3

Two revised floras from the
Southern California Botanists

A FLORA OF TBE SANTA ROSA PLATEAU,
SOUTHERN CALIFORNIA. By Earl W.
Latbrop and Robert F. Tboroe. 39
pages; paperback; coab binding;

$7.00

FLORA OF THE 8ANTA HONICA
MOUNTAINS, CALIFORNIA.

By Peter H. Raven, Henry J.
Thoapaon, and Barry A. Prigge.
179 pagea; paperback; earth
eeao binding; $10.50

Pleaae aend :

copies of A FLORA OF THE SANTA ROSA PLATEAU # $7.00

copies of FLORA OF THE SANTA MONICA MOUNTAINS « $10.50

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14

SOUTHERN f A I I FOR N I A BOTANISTS

Rirxho S*ni« Ana Boiinx Gtrden
* SOO Nnnh Collect A venue
Cli'cmoni. CA

The purpose of the SOUTHERN CALIFORNIA BOTANISTS l* the study,
preservation and conservation of the native pleats of California, and
the education of the public to the value of the native flora and Its
habitats It Is a non-profit association formed In IMF

Membership benefits Include

Various field trips throughout the state Isd by competent

field botanists and biologists.

A yearly plant sale featuring native California species

An annual symposium on various aspects of the California

vegetat Ion

The SCB Journal. CROSSOSOMA

Discounts on botanical and natural history books

Dues are for a calendar year Ne» members Joining from May through
September, please deduct tl 00 from your dues. Ttoose Joining in October
through December are credited *lth the following year's dues

Membership categories are:

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/

Individual*

$ 8.00

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/

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$15.00

/

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Renewel

•Includes membership for entire family.

Oate

Name

Address

City ’ Zip Code

Phone J

In addition, I want to give $_ to help support SCB.

Ueke check payable to SOUTHERN CALIFORNIA BOTANISTS
Mall check and form to

Alan P. Romspert
Southern California Botanists
Department of Biological Science
California State University, Fullerton
Fullerton. CA 926)4

CROSSOSOMA (ISSN 0891-9100) is published bimonthly (February, April, June, August,
October, and December) by Southern California Botanists, a California non-profit
corporation. Back issues of CROSSOSOMA are available for $2.00 an issue (plus 25c
postage) or $8.00 a volume (plus $1.00 postage). Send a check with your request to
Alan P. Romspert, Treasurer, at the above address. Manuscripts submitted for
publication should be addressed to Or. Allan A. Schoenherr, Editor of CROSSOSOMA,
Division of Biological Sciences, Fullerton College, Fullerton, CA 92634.

15

SCB COMING EVENTS (DETAILS WITHIN)

August 22 ,
August 29 ,
November 1

23

Newport

BacV.

Bay

30

Southern

S i ei

rra Conifers

13th. An

nua 1

Sympos i um

CROSSOSOMA

SOUTHERN CALIFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden, Claremont, CA 91711

o

CROSSOSOMA Vol . 13, No. 5 (PROGRAM ISSUE)
Managing Editor: Allan Schoenherr

October, 1987

SOUTHERN

COSPONSORED BY THE DEPT. OF ECOLOGY
AND EVOLUTIONARY BIOLOGY AND PETEfi
BOWLER, DIRECTOR OF THE COOPERATIVE
OUTDOOR PROGRAM IN THE STUDENT
ACTIVITIES OFFICE AT UC, IRVINE

CALIFORNIA

ACTIVITIES OFFICE AT UC, IRVINE VIIUX i VJIIIi

13TH ANNUAL SYfIPOSIUtL BOTftNISTS

BOTANICAL DISCOVERIES
AND EXPLORATION IN
SOUTHERN CALIFORNIA

DATE: SATURDAY, NOUEMBER 14, 1987

time: 8:45 AM TO 4:00 PM

REGISTRATION
BEGINS AT 8:00 AM

PLACE: HERITAGE ROOM, UNIVERSITY
CENTER; U.C., IRVINE

COST: $ 7.00 CURRENT S.C.B. MEMBER

$15.00 WITH RENEWAL OF MEMBERSHIP

610.00

NON-MEMBER

UNIVERSITY OF CALIFORNIA

PROGRAM SCHEDULE

8:00 Registration; coffee

Heritage Room, U. C. I.

9:00 Program Introduction. Geoff Smith, SCB President.

9:10 Dr. Mildred E. Mathias , UCLA, Los Angeles, California 90024:
"The Fascinating History of the Early Botanical Explorations
and Investigations in Southern California."

Early botanical explorations in western North America
were concentrated primarily in the northwest, extending into
northern California as far south as Monterey Bay. Southern
California was neglected except for the vicinity of the
several ports of call by the early sailing vessels: Santa
Barbara, Ventura, San Pedro, and San Diego.

Vegetation in southern California was briefly described
by the Franciscan friars in the last half of the 18th century
as they journeyed north from Baja California to establish
the series of missions but no collections exist to document
this flora. Except for a rare specimen obtained when a
ship stopped at one of the mission ports or an intrepid
collector made his way overland south from Monterey to Santa
Barbara southern California was neglected and its flora
recorded primarily from the immediate coastal areas of the
harbors and the missions. It was not until gold was
discovered and California became a state that botanical
explorations began in the south. These were usually carried
on as part of the boundary and railroad surveys. A few
collections were made by surgeons and others stationed at
the army forts in the Owens Valley and at Fort Tejon. These
early collections give us a limited documentation of the
native flora since it had been modified during the
previous century by European plant introductions and the
ranching and timbering activities of the settlers.

The last half-century had seen an explosive growth in
botany in southern California with the post-war boom of
new educational facilities, the improvement of roads
providing ready access to many once remote sites, and in
recent years the interest in finding and recording the

- 3 -

occurrence of rare and endangered species. The publication
of general floras for the region and the many check-lists
and florulas for local areas has increased popular interest.
Readily available documentation of the flora has provided
a base for ecological and biogeographic studies. This will
lead to a better knowledge of one of the more complex and
interesting floras of North America.

10:00 Dr. Tom Elias: "The Role of the Rancho Santa Ana Botanic

Garden in California Botany."

The Rancho Santa Ana Botanic Garden, since its
inception 60 years ago, has played a major role in the
study and conservation of the California flora. The
Garden began in Orange County on the original Rancho
Santa Ana and then moved in 1951 to its present
location in Claremont. This move and affiliation with
the Claremont Colleges and Claremont Graduate School
provided the setting for a graduate program in botany.

The herbarium and library have grown steadily in size
and importance to the systematic community. The living
collection is devoted to the native California flora
and now includes many taxa which are rare and endangered
in their natural habitat.

The Garden has embarked on an ambitious long-range
plan to strengthen its activities in systematic and
evolutionary botany, the graduate program in botany and
programs relating to the conservation of the California
flora. Major changes to the grounds and living
collections will result in the construction of a new
entrance, a visitors' and education center, several new
theme gardens, and a complete renovation of our plant
communities display. This, coupled with an expanded
public education program, will allow us to better
address community and regional needs concerning the plants
and aspects of the natural history of California.

- 4 -

10:50

Break.

11:10 Dr. Ralph Philbrick , 29 San Marcos Trout Club, Santa

Barbara, California 93105. "Botanical Lessons from the
California Islands."

Some aspects of the vegetation of the California
offshore islands have changed since the time of Greene and
the Brandegees and so have some aspects of access and
technique for the usual island botanical study. Twentieth
century botanists have improved transportation systems
and field facilities, which help support resident and
part-time resident botanists closer to their work, thus
contributing, for example, to the rediscovery of Arabis
hoffmannii and a detailed knowledge of the factors
influencing the distribution of Lyonothamnus f loribundus
asplenifolius . Biosystematic techniques involving controlled
crossing, cytology, and electrophoretic studies are being
applied, with significant results, to at least a few
problem groups such as island Malacothrix hybrids and
endemic taxa. During the 1960's and 1970' s many plants
that are restricted to one or more of these islands were
documented as being extremely abundant, at least in portions
of the islands where they occur. Such plants include taxa
of Eriogonum, Dudleya , Astragalus , Cryptantha , Castille ja ,
Haplopappus , Hemizonia, and Malacothrix . The abundance of
other plants is now rapidly increasing with a reduction in
herbivore predation. Feral burros have been removed from
San Miguel Island, rabbits from Santa Barbara Island, and
sheep from much of Santa Cruz Island. Vegetative cover is
increasing on San Miguel. Coreopsis gigantea and Dudleya
traskiae are expanding their populations on Santa Barbara;
and pines, bunch grasses, and endemic shrubs are recolonizing
new areas on Santa Cruz.

The favorable and open insular habitat and appropriate
plant adaptations have been present for centuries and more.
The habitat is being enhanced on at least some of the islands
by appropriate management, and the resulting revegetation

- 5 -

Perhaps in the future it will not be necessary to collect
only on Prince Island, off San Miguel, to find Elymus
condensatus "Canyon Prince;" and perhaps the bizarre flora
of Isla Zapato will return to the adjacent Guadalupe
Island. The current crop of island botanists should be
able to establish the effect of vegetation changes on the
rate of hybridization and the perpetuation of hybrid products,
by native plants has been established as a predominant trend.

12:00 Lunch.

1:40 Dr. Jon E. Keeley: "A Geneology of Chaparral Research:

The Family Shrub."

Contributions to the understanding of chaparral have
come from different fields and numerous institutions. In
the early part of this century most studies related to the
systematic and evolutionary relationships of both herbaceous
and shrubby members of this community. Starting in the
1950's there was a shift towards investigations into the
fire ecology and demography of chaparral, and this interest
has accelerated in recent years. In the 1970's the study
of shrub structure and function and comparative studies
with other mediterranean systems was an important focus.

.Of course behind all of this there are faces, and an attempt
will be made to call attention to the large number of
scientists who have contributed. Unlike a genelogical
family tree, which has a clear basis for illustrating
relationships, this chaparral family shrub must be more
loosely interpreted. Where clear academic lineages are
evident, these will be illustrated. In other areas, such
as systematic botany, individuals have made significant
contributions, rather than influencing large numbers of
students. Many of these were colorful characters and will
be illustrated as they reflect on the personality of
persons attracted to this ecosystem. One theme that is seen
repeatedly in this family shrub of chaparral biologists,

- 6 -

is that this ecosystem has attracted expertise from a
diversity of backgrounds and other areas of interest. Some
of these persons have made major contributions beyond
chaparral, e.g., Joseph Grinnell, animal ecologist noted
for his coining the term "ecological niche;" Theodosius
Dobzhansky , emminent population geneticist attracted to the
study of manzanita hybrids; Frits Went, discoverer of the
plant hormone auxin who started some early studies on
postfire seed germination. Many major contributors to
chaparral research also have had a history of contributions
to other systems, e.g., the arctic tundra, tropics, vernal
pools, salt marshes, deserts, etc. Chaparral is an
ecological and evolutionary system which undoubtedly will
continue to attract attention and we are far from thoroughly
understanding its mysteries.

2:30 Break.

2:40 Lloyd Smith : "Accounts of the Contributions of a Desert

Botanical Explorer, Edmund C. Jaeger, to our Knowledge and
Appreciation of the Beauty of Southern California Deserts."

EDMUND CARROLL JAEGER - January 28, 1887 to August 2,
1983, age 96. Born in Loup City, Nebraska; family moved to
Riverside, California in 1906. Degree from Occidental
College; 2 years' graduate study at University of California
honorary doctorates from Occidental College and University
of California, Berkeley.

Taught 1 year in Palm Springs in 1915; friends with
Smeaton Chase (the desert writer) and Carl Eytel (the
desert artist) .

Became head of zoology department at Riverside City
College from 1922-1952; retired after 30 years of teaching.

First book: "The Mountain Trees of Southern California"
which he illustrated. Then wrote "Denizens of the Desert,"
"Denizens of the Mountains," "The California Deserts,"
"Desert Wildf lowers , " "The North American Deserts,"

- 7 -

"Sourcebook of Biological Names and Terms," "Sourcebook of
Medical Terms," "Biologist's Handbook of Pronunciations,"
"Desert Wildlife," and "A Naturalist's Death Valley," plus
111 monthly articles for Desert Magazine , as well as articles
in the National Geographic , Condor , Journal of Mammology ,

Auk , and Pacific Discovery.

Accomplishments: First report on the plants and animals

of the Spring (Charleston) Mountains of Nevada; first
botanical collection in the Clark Mountains (4 new species
in one day!); discovered Munz Cholla 10 years before it was
named; discovered new Joshua Tree in southern Nevada
subsequently named " jaergeriana ; " explored and collected
plants for identification all over the Mojave and Colorado
Deserts, filling in many distribution gaps; first to
describe a hibernating bird (the Desert Poorwill) in 1946
in the Chuckawalla Mountains of California (observed 4
winters, to 1949) .

Dr. Jaeger helped establish Joshua Tree National
Monument by showing desert bighorns to Horace Albright,
Secretary of the Interior.

Honors: 160 acre site of the Poorwill location in

1964 became the Edmund C. Jaeger Nature Sanctuary by Nature
Conservancy; Loma Linda established a Nature Reserve in
his name in the San Bernardino Mountains.

Started palavers in 1943 to interest young people in
the desert by camping-out, with lecture and guided walks;
well over 50 of those were held, with 100-200 present at
each. Many of these went on into careers of biology,
including David Keck of the New York Botanic Gardens.

Transportation: by burro named Nettie in Palm Springs;

1937 Chevrolet sedan called "Whichwhat - the Golden Scorpion,"
and finally a jeep station wagon in which he slept when afield.

For his wildf lower book he made 857 pen-and-ink drawings,
all but 8 from life in the field; for his sourcebooks of
names he wrote literally hundreds of letters to find
derivations.

- 8 -

When young, two books influenced him: John Muir's
"Thousand Mile Walk to the Gulf: and Robert Louis Stevenson's
"Travels with a Donkey."

Dr. Jaeger gave scholarships to over a hundred students,
usually anonymously and established a scholarship fund at
Loma Linda University.

Tribute by Ed Ainsworth of the Los Angeles Times:
"Nature's most precious gift to a man is to endow him with
the faculty of translating the world around him so his
fellows have a better understanding of the meaning of life.
Among those thus richly blessed is one who has rightly earned
the designation "naturalist." He is Edmund C. Jaeger who
has just been aptly described as "Philosopher Emeritus of
the Lands of Meager Rains."

30 Closing Remarks.

ANNOUNCEMENTS
TRAVELING FINE ART SERIES

Julie Ann Schneider is a scientific illustrator who works at
Rio Hondo College. She is developing a traveling fine art series
about rare, threatened, and endangered plants and habitats. She
is seeking input on special areas, habitats, or species that may be
known to SCB members. If the area, habitat, or species meets her
criteria she will do a watercolor painting to be included in her
series. For more information contact Julie at work (213)692-0921,
Ext. 230 or at home (213)698-6059. You also may contact her by
writing to P. O. Box 5321, Whittier, CA 90607-5321.

NATURAL HISTORY OF ORANGE COUNTY
This symposium on October 10, 1987 will run from 9:00 a.m. to
4:00 p.m. at the Heritage Room of the University Center at U. C.
Irvine. It is sponsored by the Natural History Foundation of
Orange County and the U. C. I. Student Activities Office. Topics
cover the past, present, and future of Orange County. Some of the
topics include Plant Communities of Orange County, Control of
Artichoke Thistle, and Current Research on Mountain Lions. For
more details contact U. C. I. (714)856-5000 or the Natural History

Museum (714)640-7120.

- 9 -

NATIVE PLANT SOCIETY ANNUAL PLANT SALE

On Saturday , October 17, 1987, the San Diego Chapter, CNPS ,
will hold its annual plant sale in Patio A of Balboa Park's Casa
del Prado. There will be a fine selection of California native
plant materials for sale, as well as posters, CNPS shirts, books
and pamphlets. People will be available to answer questions about
propagation and use of native plants. For additional information
call Bobbie Steele (619)275-3732.

TROUBLED WATERS: THE FUTURE OF THE COAST
A variety of environmental organizations is sponsoring this
national conference on offshore drilling issues. Legislators,
politicians, environmentalists, industrialists, and reporters are
scheduled participants. The conference is to be held Sunday
evening, November 8, and all day Monday, November 9, at the
Sheraton Miramar Hotel, 101 Wilshire Blvd., in Santa Monica. For
more information call (415)485-5933.

ENDANGERED AND SENSITIVE SPECIES OF THE SAN JOAQUIN VALLEY
California State College, Bakersfield is hosting this conference
on the biology and management of endangered species in the San
Joaquin Valley. It will be held at the Bakersfield campus on
December 10 and 11, 1987. The purpose of the conference is to
provide a forum for exchange of information among governmental,
academic, conservation, and industry groups on the ecology and
conservation of biota of the San Joaquin Valley. For more
information contact Ted Murphy, Department of Biology, California
State College, Bakersfield, CA 93311.

WILDLIFE , COASTAL , AND PARKS INITIATIVE
This initiative, endorsed by developers and environmentalists
alike, will place a bond act for $776 million on the June 1988
ballot. If passed by California voters, it will provide money for
the purchase of land for parks and open space. If you can circulate
a petition, write to Californians for Parks and Wildlife, 909 12th St.,
Suite 203, Sacramento, CA 95814 or contact the local Sierra Club.

- 10 -

CALENDAR OF EVENTS

October 31, November 1. ( Saturday-Sunday ) . CNPS Desert Weekend .

Erik Jonsson of the San Diego Chapter of the California Native
Plant Society will lead this field trip to the Colorado Desert.

(In a future issue of CROSSOSOMA there will be an article on
interesting floristic finds of this area co-authored by Erik Jonsson
and Duffie Clemons) . The meeting place for this field trip will be
at the "Liquor Barn parking lot" in El Cajon at 8:00 A.M. If you
plan to go, please call Erik in advance at (619)565-0548.

November 14 ( Saturday ) . Botanical Discoveries and Exploration
in Southern California . This, the SCB Thirteenth Annual Symposium,
will be held at the Heritage Room on the U.C.I. campus. Details
are in this issue of CROSSOSOMA. For more information contact
Geoff Smith at (714)992-7381 or Peter Bowler at (714)856-5181.

November 26-29 ( Thursday-Sunday ) . Baja California . Walt Wright
•would like to take a group to Bahia de Los Angeles on the Sea of
Cortez. For details of this trip scheduled for Thanksgiving weekend
call Walt at (714)529-4134.

December 20 ( Sunday ) . Cabrillo Marine Museum . Walt Wright
will lead this trip to visit the museum and nearby tide pools to
observe intertidal algae. Meet at the museum at 11:00 A.M. Low tide
is at 3:00 P.M. Take the San Diego Freeway (405) toward San Pedro.
Take the Harbor Freeway (110) south to 9th St. Go east to Pacific,
then south to Stephan M. White Blvd., than east to the Museum. For
more information call Walt at (714)529-4134.

Plant Lovers of Southern California . The journal of the Southern
California Botanists, CROSSOSOMA, provides an ideal means by which
you can publish anything of botanical interest to southern Califorians.
Do you have a favorite field trips, gradening hints, or some pre-
liminary data that you'd like to have in print? Submit it to:

Dr. Allan Schoenherr

Division of Biological Sciences, Fullerton College
321 E. Chapman Ave .

Fullerton, CA 92634

- 11 -

October

November

November

December

SCB COMING EVENTS (Details Within)

31, November 1

CNPS Desert Weekend

14

Thirteenth Annual Symposium

26-29

Baja California

20

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CROSSOSOMA

SOUTHERN CALIFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden, Claremont, CA 91711

CROSSOSOMA Vol . 13, No. 6

December, 1987

Managing Editor: Allan Schoenherr

Effects of Dyer's Greenwold, Cytisus linifolius (Fabaceae)

Introduction

The spread of escaped exotic weeds can ruin the integrity of
native California plant communities. Cytisus linifolius (L.)
Lam. and C_^ monspessulanus L. (Fabaceae) are proving to be such
weeds on Santa Catalina Island.

In many areas of California, these two species, respectively
referred to as Dyer's Greenwold and French Broom, have become
invasive exotic pests that seem to outcompete native and endemic
plants (McClintock, 1985). Chemical or mechanical methods of
control are not feasible due to watershed problems and habitat
(steep slopes or streambeds). In order to control the spread of
Cytisus on Santa Catalina Island, a baseline study on the natural
history of Cytisus must first be undertaken before considering or
implementing a biological control.

The purpose of this study is to (1) gather pertinent
information for future management use in biological control of C.
linifolius on Santa Catalina Island and (2) test the hypothesis
that C^ li ni folius* has an effect on native and endemic
vegetation. Although the Santa Catalina Island endemics are not
state or federally listed, their habitats are similarly
threatened by feral animals in addition to the invasive Cytisus

on the native vegetation of Santa Catalina Island

Lisa Wolfbrandt Mastro
California State University, Long Beach
Department of Biology

species .

Cytisus linif olius , a European species, has seemingly
adapted well to Santa Catalina Island's Mediterranean climate.
Its potential for spreading and quickly dominating disturbed and
burned areas has led to a concern for controlling the species.

Biological control of weeds may involve the introduction of
natural enemies, the manipulation of the environment to increase
the impact of natural enemies, and the release of natural enemies
against natural pests. To apply biological control, information
such as taxonomic position, total present distribution, biology,
ecology and economic importance of the target species is needed
as well as on the biology and impact of natural enemies on
potential hosts (National Academy of Sciences, 1968).

Taxonomy

Although commonly known under the genus Cytisus and listed
as such in the flora for Santa Catalina Island (Thorne, 1967),
these two species are also referred to as Teline monspessulana
(L.) Koch and T^ linif olia (L.) Webb and Berth. (Gibbs and
Dingwall, 1971) as well as Genista monspessulana and G_^ linifolia
( Polhill , 1976).

According to an unpublished list compiled by H. M.
Butterfield, C^_ monspessulanus has been in California since at
least 1871, when it was listed in a catalog for Belleview Nursery
in Oakland (McClintock, pers. comm.). Cytisus linifolius was
recorded as naturalized on Santa Catalina Island in 1938
(McClintock, pers. comm.) and is currently the more predominant
of the two species.

Distribution

On Santa Catalina Island, Cytisus linifolius is abundant in
three major canyons surrounding Avalon (Avalon Canyon, Descanso
Canyon and Hamilton Canyon) and along Pebbly Beach Road. A small
population occurs in one canyon in the interior of the island
(Cottonwood Canyon). Island endemic plants potentially
threatened include Crossosoma calif ornicum (Crossosomataceae) ,
Lotus argophyllus ssp. ornithopus (Fabaceae), Eriogonum giganteum
ssp. £igan t eum ( Polygonaceae ) , and Rhamnus Pilil olia
(Rhamnaceae ) . Two additional species are currently listed on the

2

State of California List of Rare and Endangered Species. The
Santa Cruz Island bird's foot trefoil, Lotus argophyllus ssp.
niveus is listed as endangered and the Santa Barbara Island
buckwheat, Eriogonum giganteum var. compactum is listed as rare.

A third plant, the San Clemente Island buckwheat, Eriogonum
giganteum var. formosum, is not on the state list but is still
considered endangered (California Native Plant Society, 1984).

Methods

To test the effects of C^ linif olius on native plant
communities on Santa Catalina Island, sixteen plots were
established throughout areas where C^_ linifolius occurs. Of the
sixteen plots, ten contained C^_ linifolius and six did not.
Over a period of one and a half years, plots were monitored for
the number of individuals per species. There were no signir leant
differences in number of individuals per species per plot between
counts for 1985 and 1986 (p > .05, paired t-test) (Figure 1).

e e 5

3 3 o>

Figure 1. Mean number of individuals per plot for plots contain-
ing Cytisus linifolius . Eriogonum giganteum ssp. giganteum is a
Santa Catalina Island endemic, and Rhamnus pirif olia is a Channel
Island endemic. All other native species were lumped together.
There are no significant differences (p < .05, paired t-test) for
number of individuals for each species for 1985 and 1986 (n * 10
5m x 5m plots).

*One plot did contain 524 individuals of C^ linifolius .

3

One plot containing linif olius had a high number of

individuals (526) which skews the mean. Because there was no
significant difference in years, the mean of both years was used
to compare plots with li ni f ol iu s to plots without

linif olius (Figure 2). Using a One-Way ANOVA there were no
significant differences (p > .05) for the mean number of individ-
uals per plot for plots with and without linif olius .

linifolius for 1985 and 1986. Eriogonum qiqanteum ssp. giganteum
is a Santa Catalina Island endemic. Rhamnus pirif o l ia and
Crossosoma calif ornicum are Channel Island endemics. All other
native species were lumped together. There are no significant
differences (p < .05, 1-Way ANOVA) for number of individuals for
each species for plots with and without C^ linifolius (n = 16 5m
x 5m plots ).

Discussion

Based on my limited data, C^ linifolius does not appear to
be outcompeting the endemic or native vegetation on Santa
Catalina Island. However, C^_ linifolius does appear to
demonstrate the potential for competition as exhibited in burned
areas (Figure 3). Current studies are underway to determine the
effects of uprooting mature C^ linifolius plants, thereby
creating a disturbed habitat that may be more suitable for seed
germination. Although evidence of competition requires much more
than a one year study, a careful monitoring of the situation on
Santa Catalina Island may be wise.

4

1 6 0

tf. eeS

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Figure 3. Mean number of individuals per plot that have
germinated or crown-sprouted in plots established after a July 2,
1985 fire (n = 2 5m x 5m plots). Eriogonum qiganteum ssp.
giganteum is a Santa Catalina Island endemic; RhamnuS pirifolia
and Lotus ar^oph^l lus ssp. orni thopus are Channel Island
endemics. These two plots appeared to be an adequate
representation of post fire recovery as the fire burned less than
five acres.

Until the last few months, C^_ linif olius has been the host

of very few insects, the most notable being Ure^ijDhita
reversalis , the Genista moth. Prior to September 1986, the moth

had only been observed in two locations on the island. Recent

surges in population have shown a much more widespread
distribution of the insect. This moth, a North American species,
is capable of extensive damage to the plants. Although not
currently found on native vegetation, further studies will be
conducted to analyze the influence of the insect on C .
linif olius .

ACKNOWLEDGMENTS

I thank Dr. E. L. Sleeper for advice during this study and
E. Mastro for reviewing the manuscript. I also thank Mr. Terry
Martin and the Santa Catalina Island Conservancy for their
support. This study has been partially funded by Southern
California Botanists, California Native Plant Society, El Dorado
Audubon Society, and Sigmi Xi, The Scientific Research Society.

5

LITERATURE CITED

California Native Plant Society. 1984. Inventory of rare and
endangered vascular plants of California. Special
Publication No. 1 (3rd edition) J. P. Smith, (ed.) Eureka,
CA. 174 p.

Gibbs, P. E. and I. Dingwall. 1971. A revision of the genus
Teline. Boletim da Sociedade Broteriana XLV(2 Serie):
269-316.

McClintock, E. 1985. Escaped exotic weeds of California.
Fremontia 12(4) :3-6.

National Academy of Sciences. 1968. Principles of Plant and
Animal Pest Control. Volume 2 Weed Control. Publication
1597. Washington D. C. 471 p.

Polhill, R. M. 1976. Genisteae (Adans.) Benth. and related

tribes (Leguminosae ) . In V. H. Heywood (ed.), Bot. Syst.
1:143-368. Academic Press Inc. London.

Thorne, Robert F. 1967. A flora of Santa Catalina Island.

Aliso. 6(3): 1-77 .

BAJA CALIFORNIA FLORAS

Dr. Robert Thorne of Rancho Santa Ana Botanic Garden is composing
a flora for the northern portion of Baja California. As a product of
that effort he has compiled a series of valuable checklists for
selected localities in Baja California. The following list of vas-
cular plants is the third of a series that will be published in
CR0SS0S0MA

VASCULAR PLANTS OF THE PARQUE NACIONAL DE CONSTITUCION DE 1857
SIERRA DE JUAREZ, BAJA CALIFORNIA, MEXICO

By

Robert F. Thorne
Rancho Santa Ana Botanic Garden
Claremont, California

A list of the species is for plants collected in the Parque
Nacioal de Constitucion de 1857, especially in and about Laguna
Hanson (Laguna Juarez) , with an elevation at the lake surface of
ca. 1610 m at ca. 32° 03' N - 115° 55' W, on decomposed granitic
soil in open forest dominated by Pinus jef freyi or in montane
chaparral with scattered Pinus quadrifolia and Pinus coulteri among
the sierritas to the west and north of the lake. A few other species
of interest collected to the north, east, and south of the lake are

6

also included. The list is based primarily on collections made by
Robert F. Thorne and associates from the Rancho Santa Ana Botanic
Garden, Claremont, California on various trips in 1983 to 1985,
usually in May or June. Late summer and fall botanical surveying
trips are badly needed to complete the list. A few additional
species not found on our surveying trips are included from collections
made by others, especially by Reid V. Moran and Ira Wiggins, deposited
at RSA, SD, and DS . The Thorne et al. specimens are mostly deposited
at RSA, SD, MEXU , and BCMEX. A preceding * in the list denotes
species believed to be introduced in the area.

Adiantaceae

Cheilanthes couvillei Maxon

Pellaea mucronata (D. C. Eat.) D. C. Eat.

Pi ty ro gramma triangularis (Kaulf .) Maxon var. triangularis
Fteridium aquilinum (L. ) Kuhn var. lanuginosum (Borg.) Fern.
Aizoaceae

Sesuvium vevruoosum Raf.

Alismataceae

Alisma triviale Pursh.

Alliaceae

Allium fimbriatum S. Wats var. parryi (S. Wats.) Ownbey & Aase
Diehelost-emma pulchellum (Salisb. ) Heller var. pulchellum >

Amaranthaceae

Amaranthus albus L.

Amaranthus californicus (Moq.) S. Wats.

Amaranthus powellii S. Wats.

Anacardaceae

Rhus trilobata Nutt. ex. T. & G. var. quinata
Apocynaceae (incl. Asclepiadoideae)

Apocunum oannabinum L. var. glaberrimum A. DC.

Asclepias eriocarpa Benth.

Aspidiaceae

Woodsia plummerae Lemmon
Asteraceae

Achillea millefolium L. var. californica (Poll.) Jeps.

Agoseris he terophylla (Nutt.) Greene

Ambrosia psilostachya DC ssp. californica (Rydb.) ined.

Anisocoma acaulis T. & G.

Artemisia dracunculus L.

Artemisia ludoviciana Nutt. ssp. albula (Woot.) Keck
Artemisia tridentata Nutt ssp. tridentata
Aster frondosus (Nutt.) T. & G.

Aster occidentalis (Nutt.) T. & G. var. delectabilis

(Hall) Ferris

Brickellia californica (T. & G.) A. Gray var. californica
Chaenactis parishii A. Gray
Chaetopappa aurea (Nutt.) Keck

Chry sothamnus nauseosus (Pall.) Britt, ssp. consimilis
(Greene) Hall & Clem.

Cirsium call fornicum A. Gray ssp. bernardinum (Greene) ined.
Cirsium tioganum (Cong.) Petr. (C. foliosum (Hook.) DC.)

Conyza canadensis (L.) Cronq.

Corethrogyne filagini folia (Hook. & Arn.) Nutt. var.
glomerata Hall
* Cosmos bipinnatus Cav.

Erigeron divergens T. & G. var. cinereum A. Gray
Erigeron foliosus Nutt.

Eriophy Hum confertiflorum (DC) A. Gray

Eriophyllum wallacei A. Gray ssp. australe (Rydb.) Wigg.

7

osa

Asteraceae (Cont.)

Gnaphalium bioolor Bioletti
Gnaphalium luteo-album L.

Gnaphalium microcepahlum Nutt.

Gnaphalium palustre Nutt.

Gutierrezia sarothrae (Pursh) Britt. & Rusby
Haplopappus ouneatus A. Gray
Helianthus californicus DC.

Mulsea mexicana Rydb.

*Hypoehoeris glabra L.

Hymenopappus filifolius var. lugens (Greene) Jeps.

Lasthenia oalifornioa DC ex Lindl. (L. chrysostoma

(Fisch. & Mey.) Ornduff)

Layia glandulosa (Hook.) Hook. & Arn. ssp. glandulo f
Madia elegans d« Don ex Lindl. ssp. elegans
Madia exigua (Smith) A. Gray

Machaeranthera asteroides (Torr.) Greene spp. lagunenszs
(Keck) ined. (M. tephrodes (A. Gray) Greene)

Microseris lindleyi (DC.) A. Gray ( M . linearifolia
(Nutt.) Sch. Bip.)

Solidago oalifornioa Nutt.

*Sonohus asper (L.) Hill. . ,

Stephanomeria virgata Benth. ssp. pleurocarpa (Greene) Gottlieb
* Taraxacum laevigatum (Willd.) DC.

Wyethia oPata T. & G. in Emory

Berberidaceae , , . . ..

Mahonia higginsae (Munz) Ahrendt. ( Berberzs hzggznsae Munz)

Boraainanceae ^

" Crypt antha micrantha (Torr.) Jtn. ssp. lepzda (A. Gray) Mathew
& Raven ( Eremooarya micrantha (Torr.) Gr-ene var. lepzda
(A. Gray) Macbr.)

Cryptantha murioata (Hook. & Arn.) Nels. & Macbr.

H elio tr opium ourassavicum L. ssp. oculatum (Heller) Thorne
Tlagiobothrys bracteatus (Howell) Jtn. ( Allocarya bracteata

Howell)

Plagiobothrys californicus (A. Gray) Greene
Plagiobothrys undulatus (Piper) Jtn. ( Alloc
Brassicaceae

Arab is sparsi flora Nutt, in T. & G.

Barbarea orthoceras Ledeb.

Descurainia pinnata (Walt.) Britt.

Erysimum oapitatum (Dougl.) Greene

Lepidium virginioum L. var. pubescens (Greene) Thell.
Lesquerella kingii S. Wats. ssp. latifolia (Nels.)

Roll. & Shaw

Borippa tenerrima Greene
Strep tanthus oampestris S. Wats.

Callitrichaceae

Cal litriche longipedunculata Morong
Callitriche verna L.

Caprifoliaceae , ,

Lonicera subspicata Hook. & Arn. var. denudata Rehd.

Lonicera subspicata var. johnstonii Keck

Sambucus mexicana Presl ex DC. _ ...

Symphori cargos parishii Rydb. (S’, oreophilus of Wiggins)
Caryophyllaceae

Arenaria oonfusa Rydb.

Silene antirrhina L.

Silene verecunda S. Wats. ssp.

Hitchc. & Maguire
Ceratophy 1 laceae

Ceratophy Hum demersum L.

Chenopodiaceae

Chenopodium rubrum L.

Monolepis nuttalliana (Schult.) Greene
Crassulaceae

Crassula oonnata (R. & P.) Berger in Engl. -

var. oonnata ( Tillaea ereota Hook. & Arn.)

Dudleya abramsii Rose

jreene

( Allocarya unaulata Piper)

platyota (S. Wats.)

& Prantl

8

Cyperaceae

Carex globosa Boott

Carex praegracilis W. Boott

Carex subfusoa W. Boott

Cyperus squavvosus L. ( C . aristatus Rottb. var. inflexus (Muhl.) Boeckl.
Eleoohavis aoicularis (L. ) R. & S.

Eleoohavis bella (Piper) Svenson

Eleo charts macros tachya Britt, ex Small

Eleo charts parishii Britt.

Eleoohavis pavvula <R. & S.) Link var. colovadensis
(Britt.) Beetle

Eleoohavis pauci flora (Lightf.) Link ssp. bevnardina
(Munz Jtn.) ined.

Soivpus aoutus Muhl. ex Bigel. ssp. occidentalis (Beetle) ined.
Datiscaceae

Datisca glomevata (Presl.) Baill.

Dracaenaceae

Nolina palmevi S. Wats. ssp. palmevi
Elatinaceae

Elatine califovnioa A. Gray ssp. oalifovnica
Ericaceae

Arotostaphylos peninsulavi s Wells
Arc tost aphylos pungens HBK
Euphorbiaceae

Chamaesyee sevpylli folia (Pers.) Small (Euphovbia
Eerpylli folia Pers.)

Euphovbia palmevi Engelm.

Fabaceae

Amovpha califovnioa Nutt.

Astragalus circumdatus Greene
Astragalus didymocavpus Hook. & Arn.

Astragalus douglasii (T. & G. ) A. Gray var. pavishii
(A. Gray) M. E. Jones

Astragalus filipes Torr. ^

Astragalus palmevi A. Gray var. palmevi

Lathy vus laetiflovous Greene ssp. alefeldii (White) Brads.

Lotus argyvaeus (Greene) Greene
Lotus hamatus Greene
Lotus nevadens is Greene
Lotus oblongi folius (Benth.) Greene

Lotus puvshianus (Benth.) Clem. & Clem. ssp. puvshianus
Lotus vigidus (Benth.) Greene

Lupinus 'bicolor Lindl. ssp. micvophyllus (S. Wats.) D. Dunn
Lupinus concinnus Agardh ssp. optatus (C. P. Sm. ) D. Dunn
Lupinus excubitus M. E. Jones ssp. austvomontanus

(Heller) Beauchamp

Lupinus fovmosus Greene ssp. proximus (Heller) Conrad
Lupinus lati folius Agardh ssp. wigginsii (C. P. Sm. )

Kenney & Dunn

Psoralea califovnioa S. Wats.

Psoralea orbicularis Lindl.

Psoralea vigida Parish

Tri folium amplectens T. & G.

Tri folium microcephalum Pursh
Tri folium tridentatum Lindl.

Trifolium variegatum Nutt, in T. & G.

Tri folium wormskjoldii Lehm.

Fagaceae

Quercu8 ahrysolepis Liebm.

Quercus dunnii Kell.

Quercus peninsularis Trel.

Quercus turbinella Greene ssp. turbinella

Quercus uislizenii A. DC. ssp. frutescens (Engelm.) ined.

Garrvaceae

Garrya veatchii Kell.

Gentianaceae

Frasera parvyi Torr.

Geraniaceae

*Erodium cicutarium (L.) L'Her. ex Ait.

Geranium atropurpureum Heller
Grossulariaceae

Ribes indecorum Eastw.

9

Hydrophyllaceae

Eriodictyon trichocaZyx Heller ssp. Zanatum (Brand) Munz
(E. Zanatum (Brand) Abrams)

Hesperochiron caZifornicus (Benth.) S. Wats.

PhaceZia imbricata Greene ssp. patuZa (Brand) Heckard
PhaceZia suf frutescens (Parry) Const.

Iridaceae

Sisyrinchium beZZum S. Wats.

Juncaceae

Juncus bryoides F. J. Hermann

Juncus bufonius L. var. haZophiZus Buch. & Fern.

Juncus macrophyZZus Coville
Juncus mexicanus Willd.

Juncus sphaerocarpus Nees

Juncus tiehmii Ertter (J. Kelloggii of Wiggins)

Juncaginaceae

LiZaea sciZZoides (Poir.) Haum.

Lamiaceae

Mentha arvenis L. var. viZZosa (Benth.) S. R. Stewart
Monarde Z la ZanceoZata A. Gray var. micvocephaZa A. Gray
Salvia pachyphyZZa Epling

Stachys rigidus Nutt. ssp. quercetorum (Heller) Epling
Trichostema austromontanum Lewis ssp. austromontanum
Lemnaceae

Lemna minuscuZa Herter (£. minima Phil.)

Lentibulariaceae

UtricuZaria vuZgaris L.

Liliaceae

CaZochortus spZendens Dougl. ex Benth.

Linaceae

Linum Zewisii Pursh.

Loasaceae

MentzeZia montana (A. Davids.) A. Davids.

Malvaceae

SidaZcea maZvaefZora (DC.) A. Gray ex Benth. ssp. sparszfoZva
C. L. Hitchc.

Nyctaginaceae

MirabiZis pumiZa Standi.

Onagraceae

BoisduvaZia densifZova (Lindl.) S. Wats.

Camissonia hirteZZa (Greene) Raven
Camissonia striguZosa (F. & M. ) Raven

Epilobium canum (Greene) Raven ssp. ZatifoZvum (Hook.) Raven

{Zauschneria caZifornica Pressl. ssp. Zat%foZza (Hook.) KecK)
Gayophytum diffusum T. & G. ssp. parvifZorum Lewis &

Szweykowski

Oenotheva caZifornica (S. Wats.) S. Wats.

Papaveraceae

PZatystemon caZifornicils Benth.

Pinaceae

Pinus coutteri D. Don
Pinus jeffreyi Grev. & Balf.

Pinus monophyZZa Torr . & Frem.

Pinus quadri foZia Pari.

Plantaginaceae

*PZantago major l.

Poaceae

Agropyron trachycauZum (Link) Malte
Agrostis scabra Willd.

Agrostis semiverticiZZatus (Forsk.) C. Chr .

Aristida schiediana Trin. & Rupr. (incl A. orcuttiana Vasey)
Bouteloua curtipenduZa (Michx.) Torr. var. caespitosa
Gould & Kapadia
Bromus carinatus Hook. & Arn.

*Bromus diandrus Roth ( B . rigidus Roth)

*Bromus rubens l.

*Bromus tectorum l.

Deschampsia danthonioides (Trin.) Munroe ex Benth.

DistichZis spicata (L.) Greene ssp. stricta (Torr.) Thorne
EZymus ZongifoZius (j. G. Sm. ) Gould ( Sitanion hystrix
(Nutt . ) J . G. Sm. )

10

Poaceae (Cont . )

^Eragrostis cilianensis (All.) E. Mosher
Eragrostis intermedia Hitchc.

Eragrostis pectinacea (Michx.) Nees
*Hordeum glaucum Steud.

*Hordeum leporinum Link.

Melica imperfecta Trin.

Muhlenbergia asperifolia (Nees & Mey.) Parodi
Muhlenbergia emersleyi Vasev
Muhlenbergia fragi-lis Swallen
Muhlenbergia rigens (Benth.) Hitchc.

Muhlenbergia wrightii Vasey

Panicum hirticaule Presl (P. distichum auct.)

Paspalum paspaloides (Michx.) Scribn. ( Paspalum
distichum auct.)

*Poa annua L.

Poa f endleriana (Steud.) Vasey

Poa secunda Presl (P. scabrella (Thurb. ) Benth. ex Vasey)
*Polypogon monspeliensis (L.) Desf.

Scribneria bolanderi (Thurb.) Hack.

Sporobolus cryptandrus (Torr.) A. Gray
Stipa coronata Thurb. in S. Wats.

Vulpia microstachys (Nutt.) Benth. var. pauciflora (Beal)

Lonard & Gould ( Festuca pacifica Piper, F . reflexa Buckl . )

( Festuca octo flora Walt.)

austromontanum
sapphirinum

Vulpia myuros (L.) K. C. Gmel. var. hirsute Hack.

(Festuca megalura Nutt.)

Vulpia octoflora (Walt.) Rydb.

Po lemon iaceae

Eriastrum densifolium (Benth.) Mason ssp
(Craig) Mason

Eriastrum sapphirinum (Eastv/.) Mason ssp
Gilia diegensi s (Munz) A. & V. Grant
Gilia mexicana A. & V. Grant
Ipomopsis tenui folia (A. Gray) V. Grant
Linanthus bellus (A. Gray) Greene

Linanthus floribundus (A. Gray) Greene ssp. floribundus
Linanthus J amauensis Moran
Linanthus lemmonii (A. Gray) Greene
Linanthus melingii (Wigg.) Grant
Microsteris gracilis (Dougl* ex Hook.) Greene
Navarretia intertexta (Benth.) Hook
Phlox aus tromontana Coville
Polygonaceae

Eriogonum davidsonii Greene
Eriogonum foliosum S. Wats.

Eriogonum has tatum Wigg.

Eriogonum parishii S. Wats

Eriogonum wrightii Torr. ex Benth. ssp. membranaceum
S. Stokes ex Jeps.

Polygonum amphibium L. var. stipulaceum Coleman
* Polygonum arenas trum Bor.

* Runes. : crispus L.

Rumex maritimus L. var. fueginus (Phil.) Dusen
(R. fueginus Phil.)

Rumex salici folius Weinm. var. denticularis Torr.

(R. calif ornicus Rech.f.)

Polypodiaceae

Polypodium calif ornicum Kaulf.

Portulacaceae

Claytonia perfoliate Donn ssp. perfoliata
Claytonia perfoliata ssp. viridis (A. Davids.) Fellows
Lewisia brachy calyx Engelm. ex A. Gray
Montia fontana L. ssp. amporitana Sennen
*Portulaca oleracea L.

Primulaceae

Anagallis minima (L.) Krause ( Centunculus minimus L. )

Ranunculaceae

Aquilegia formosa Fisch. in DC. var. hypolasia (Greene) Munz
Clematis pauciflora Nutt, in T. & G.

Myosurus aristatus Benth. ex Hook.

Myosurus minimus L.

Ranunculus aquatilis L. var. capillaceus (Thuill.) DC.
Ranunculus cymbalaria Pursh ssp. saximontanus (Fern.) Thorne

11

Rhamnaceae

Ceanothus greggii A. Gray ssp. perplexans (Trel.) Beauchamp
Ceano thus leucodermis Greene

Rhamnus californica Esch. ssp. tomentella (Benth. ) C. B. Wolf
Rhamnus ilici folia Kell.

Rosaceae

Adeno stoma fasciculatum Hook. & Arn. var. / as oioulatum
Adenoatoma sparsi folium Torr. in Emory
Amelanchier utahensis Koehne
Cercocarpus be tuloi.de 8 Nutt, ex T. & G.

Horkelia bolanderi A. Gray ssp. ssp. clevelandii (Greene) Keck
Fotentilla biennis Greene

Fotentilla glandulosa Lindl. ssp. reflexa (Greene) Keck
Fotentilla saxosa Lemmon ex Greene
Prunus ilici folia (Nutt.) Walp. ssp. ilicifolia
Prunus virginiana L. ssp. demissa (Nutt.) E. Murra
Rosa californica Cham. & Sch.

Vauquelinia californica (Torr.) Sarg.

Rubiaceae

Galium andrewsii A. Gray ssp. andrewsii

Galium angusti folium Nutt, ex T. & G. ssp. angusti folium
Galium wrightii A. Gray
Salicaceae

Salix exigua Nutt.

Salix laevigata Bebb var. laevigata
Salix lasiolepis Benth. var. lasiolepis

Saururaceae

Anemopsis californica (Nutt.) Hook. & Arn.

Sax if ragaceae

Reuchera leptomeria Greene var. peninsularis Rosend. ,

Butt . , & Lak.

Scrophulariaceae (incl. Orobanchoideae)

Castilleja foliolosa Hook. & Arn. (toward C. lanata A. Gray)
Cas title ja miniata Dougl. ex Hook. var. oblongif olia
(A. Gray) Munz

Castilleja stenantha A. Gray

Cordylanthus filifolius Nutt. ssp. involutus (Wigg. ) Heckard
Cordylanthus nevinii A. Gray

Keckiella ternata (Torr. ex A. Gray) Straw ( Penstemon
ternatus Torr.)

Limo sella acaulis Sesse & Moc.

Linaria canadenis (L.) Dum.-Cours. ssp. texana
(Scheele) Pennell
Mimulus diffusus Grant
Mimulus florihundus Dougl. ex Lindl.

Mimulus guttatus Fisch. ex DC. ssp. guttatus

Mimulus nasutus Greene

Mimulus pilosus (Benth.) S. Wats.

Orobanche californica C. & S. ssp. feudgei (Munz) Heckard
Orobanche fasciculata Nutt.

Orthocarpus attenuatus A. Gray
Penstemon bridgesii A. Gray
Penstemon centranthi folius Benth.

Penstemon clevelandii A. Gray ssp. clevelandii
Veronica peregrina L. ssp. xalapensis (HBK . ) Pennell
Solanaceae

Nicotiana attenuate Torr. ex S. Wats in King
Petunia parvi flora Juss.

Solanum xanti A. Gray
Sterculiaceae

Fremontodendron cali fornicum (Torr. ) Cov.

Typhaceae (incl. Sparganioideae)

Sparganium eurycarpum Engelm. (5. greenei Morong)

Urticaceae

Urtica dioica L. ssp. holosericea (Nutt.) Thorne
Verbenaceae

Verbena orcuttiana Perry
Violaceae

Viola purpurea Kell ssp. purpurea

12

Viscaceae

Arceuthobi um campy lopodum Engelm. (on Pinus jeffreyi J
Avceuthobium divaricatum Engelm. (on Pinus quadrifolia)
Phovadendron vittosum (Nutt, in T. & G.) Nutt. ssp.
Zannichelliaceae

Zannichellia palustris L.

A Commentary on the 13th Annual Symposium

If a symposium is considered "successful" in terms of quality
of presentation by the speakers, then the November Sumposium held
at U.C. Irvine was outstanding! Each speaker in turn was thoroughly
prepared and gave an excellent presentation on his/her topic (refer
to the October issue of Crossosoma for the speaker's abstracts).

Many anecdotes were given to illustrate the rich and colorful
history of those botanists who have significantly contributed to
our current wealth of botanical collections and research on the
Southern California flora. Additionally, advance publicity of the
Symposium was well organized (thanks to Kathleen Pahl, Katie Barrows,
and Allan Schoenherr of the S.C.B. Board of Directors), facility
accommodations were excellent (thanks to Peter Bowler of U.C. I. and
Alan Romspert of Cal. State Fullerton), and the program registration/
parking was handled efficiently (thanks to Sherry Schmidt, Coleen
Cory, Judy Smith, Diana Cosand, and others) . A special thank you
goes to Bruce Trotter for his contributing the fascinating display
of photos, journals, and camping memorabilia of the famous Edmund
Jaeger "palavers."

An Additional Comment on the Symposium .

In light of the reputation of the speakers and their obvious
dedication to present a high-quality symposium, and with respect
to the well-prepared advance publicity, it was disappointing to have
mediocre attendance at the program. Of particular concern to me and
the Board of Directors is that virtually no students or faculty of
botany departments at our regional colleges and universities were in
attendance. If the faculty members of these institutions show an
apparent apathy and non-interest in supporting botanical symposia/
conferences, their students will likewise not become involved. This
is regrettable, becuase it is through our educational programs that
recognition of the importance of the natural sciences to our highly
urbanized society is best fostered.

Geoff Smith, SCB President
13

FIELD TRIPS

December 20 ( Sunday ) . Cabrillo Marine Museum . Walt Wright
will lead this trip to visit the museum and nearby tide pools to
observe intertidal algae. Meet at the museum at 11:00 A.M. Low tide
is at 3:00 P.M. Take the San Diego Freeway (405) toward San Pedro.
Take the Harbor Freeway (110) south to 9th St. Go east to Pacific,
then south to Stephan M. White Blvd., than east to the Museum. For
more information call Walt at (714)529-4134.

January 16 (Saturday) . Torrey Pines State Park. This trip will
examine some of the unique plants found in the reserve. We will
look for Shaw's agave, warty stem ceanothus and western dichondra.

We will also see some of the unique desert elements in this flora
including jojoba and Mojave yucca. Take 1-5 (South) to Carmel
Valley Road, West to Torrey Pines Road to the park entrance. Go
up the hill and we will meet at the Visitor's Center at 9:00 a.m.

For details, contact Dave Bramlet at (714)549-0647.

February 13 (Saturday) . Fungus Foray. Starr Ranch . Walt Wright
will lead this foray potentially held at Starr Ranch. We will
look for mushrooms and examine the oak woodlands and grasslands
at the Ranch. Take 1-5 South to Ortega Hwy (74) and go east. We
will meet at the Caspers Parking lot at 9:00 a.m. (immediately west
of the entrance). If you have a high clearance vehicle, please
bring it along. Be prepared to pay up to $10.00 in entrance fees.

For details, contact Dave Bramlet at (714)549-0647.

February 21 (Sunday) . Laguna Beach . Fred Roberts will lead this
trip to examine some of the unique plants including Verbisina dissita ,
Laguna Beach dudleya, multi-stemmed dudleya and western dichondra
found along the coast near Laguna Beach. To reach the meeting site
take 1-405 S to 133 (Laguna Beach Freeway) and go south on PCH to
Aliso Beach (just south of the Aliso Golf Course) and go right on
Aliso Way. We will meet at 9:00 a.m. in the parking lot. For
details, contact Dave Bramlet at (714)549-0647.

February 27 (Saturday) . Thousand Palms Reserve . Katie Barrows will
take us on this tour of a California fan palm oasis. The reserve
is run by the Nature Conservancy. To reach the reserve take the
I -10 to Rawan Road and go East 6 miles to Palm Cyn. Rd. Then go left
to the Thousand Palms Reserve entrance. Meet at 9:00 a.m. For
details, contact Dave Bramlet at (714)549-0647.

14

Charter Boat Trip to Guadalupe and Cedros Islands of Coastal Baja
California , Mexico March 26 through April 1, 1988 . Several botanists,
ornithologists and naturalists have expressed an interest in a charter
boat trip to Guadalupe, Cedros and San Martin Islands during the
Easter break of 1988. The primary purpose of the trip is the
observation of the unique flora and fauna of these islands and
surrounding waters. Much of the time will be spent hiking and exploring
the interesting biology and geology of the islands. There will also
be abundant marine mammal and sea bird observations in route and on
the islands. Plans are to have prominent botanists from Mexico and
the U. S. on the trip and persons knowledgeable of marine mammals,
birds and island biogeography.

This is a non-profit cooperative charter that will cost
approximately $950 per person for the 7 day trip. All meals and
lodging on the vessel are included. Departure from San Diego will
be Saturday, March 26 and return on Friday, April 1. Plans are to
spend 3 days in the vicinity of Guadalupe Island with onshore hikes
to visit the Monterey Pine and Guadalupe Cypress forest and other
botanical and wildlife interests. Hopefully we will be able to gain
permission to spend one night on the island in the cypress forest
found at the higher elevations. A day will be spent on Cedros Island
and a half day on San Martin if time is available.

More information is available by contacting Margie Stinson at
(619)726-2228, Thomas Oberbauer at (619^225-8397; work 565-3041) or
writing to Biological Adventures, Fisherman's Landing, 2838 Garrison,

San Diego, CA 92106. Make checks payable to Pacific Sportf ishers .

The trip is sure to be fun and educational. Don't miss out.

TENTATIVE DATES FOR FUTURE EVENTS.

March

5

Camp Pendelton

March

12

Guadalupe Canyon,

Baja Calif.

March

19

Santa Ana River

March

20

Plant Sale at Rancho Santa Ana

April

10

Pines to Palms Highway

May 15

Spring Mountains,

Nevada

15

ANNOUNCEMENTS

Natural History Museum of Orange County . The Natural History Museum
of Orange County offers a variety of programs and nature walks. The
next walk on Saturday, December 5, 8:30 a.m. , will be led by Dr.

Phil Brylski of U. C. Irvine. Call the museum (714)640-7120.

ENDANGERED AND SENSITIVE SPECIES OF THE SAN JOAQUIN VALLEY
California State College, Bakersfield is hosting this conference
on the biology and management of endangered species in the San
Joaquin Valley. It will be held at the Bakersfield campus on
December 10 and 11, 1987. The purpose of the conference is to
provide a forum for exchange of information among governmental,
academic, conservation, and industry groups on the ecology and
conservation of biota of the San Joaquin Valley. For more
information contact Ted Murphy, Department of Biology, California
State College, Bakersfield, CA 93311.

Orange County F. E. D. Valentine Dance. The Orange County Fund for
Environmental Defense is hosting a Valentine Masquerade Dance on
Saturday, February 13. The F. E. D. is currently trying to preserve
open space in Caspers Park. A legal appeal will be heard soon.
Donations are tax deductible. Pre-registration for the dance is $8.00
It costs $10.00 at the door. Mail pre-registration check to F.E.D.,

P. 0. Box 9118, Fountain Valley, CA 92708. Enclose SASE for
confirmation. For more details, call Charlotte Clark at (714)963-1430

WILDLIFE , COASTAL , AND PARKS INITIATIVE
This initiative, endorsed by developers and environmentalists
alike, will place a bond act for $776 million on the June 1988
ballot. If passed by California voters, it will provide money for
the purchase of land for parks and open space. If you can circulate
a petition, write to Californians for Parks and Wildlife, 909 12th St.
Suite 203, Sacramento, CA 95814 or contact the local Sierra Club.

16

Restoring the Earth Conference. The first national gathering to
consider the restoration of all natural resource types and the
redesign of urban areas will be held on January 13—16 at the University
of California, Berkeley. The conference is organized by the Restoring
the Earth project of The Tides Foundation, San Francisco, and cospon-
sored both by the College of Natural Resources and the Center for
Environmental Design Research of the University of California,

Berkeley. It will bring experts in natural resource restoration and
management together with a broad selection of academic, government,
industry, foundation, labor, public health, and environmental
representatives. Participants will help create new solutions to the
nation's environmental problems, through restoration of damaged
resources .

Topics to be covered include restoration of coastal ecosystems
and estuaries; rivers and lakes; streams and fisheries; rangelands,
prairies, mined lands, forests and wildlife; atmosphere and climate;
dry lands and agricultural lands; urban environmental planning; and
control of toxic wastes. Formal refereed papers will be presented
at scientific and technical sessions. Non- technical sessions will
include accounts of restoration successes and discussions of policy
issues, legislation, litigation, trends, and resource conflict
resolution. The program also includes keynote panels, plenary
sessions, workshops, films, and exhibits.

Amateur a n d Professi onal Botanists . The journal of the Southern
California Botanists, CR0SS0S0MA, provides an ideal means by which
you can publish things of botanical interest to southern Californians,
If you have a favorite field trip, gardening hints, or some pre-
liminary data that you'd like to have in print submit your manuscript

to: Dr. Allan Schoenherr

Division of Biological Sciences, Fullerton Callege
321 E. Chapman Avenue
Fullerton, CA 92634

17

Two revised floras from the
Southern California Botanists

A FLORA OF THE SANTA ROSA PLATEAU,
SOUTHERN CALIFORNIA. By Earl W.
Lathrop and Robert F. Thorne. 39
pages; paperback; comb binding;

FLORA OF THE SANTA MONICA
MOUNTAINS, CALIFORNIA.

By Peter H. Raven, Henry J.
Thompson, and Barry A. Prigge.
179 pages; paperback; smyth
sewn binding; $10.50

Please send:

Price

copies o

f A FLORA OF THE SANTA ROSA PLATEAU 0 $7.00

copies of FLORA OF THE SANTA MONICA MOUNTAINS £ $10.50

Return to: So. Calif Botanists
Dept, of Biology
Calif. State University
Fullerton, CA 92634

Total

Price includes tax, handling,
and postage.

Make check or money order payable to: Southern California Botaniste

18

SOUTHERN CALIFORNIA BOTANISTS

Rancho Santa Ana Botanic Garden
1500 North College Avenue
Claremont, CA 91711

The purpose of the SOUTHERN CALIFORNIA BOTANISTS Is the study,
preservation and conservation of the native plants of California; and
the education of the public to the value of the native flora and its
habitats. It is a non-profit association formed in 1927.

Membership benefits include

Various field trips throughout the 6tate led by competent
field botanists and biologists.

A yearly plant sale featuring native California species.

An annual symposium on various aspects of the California
vegetat ion .

The SCB Journal. CROSSOSOMA .

Discounts on botanical and natural history books.

Membership categories are:

/

/

Individual*

$ 8.00

/

/

New Member

/

/

Croup or organization

$15.00

/

/

Renewal

^Includes membership for entire family.

Date

Name

Address . .

City Zip Code

Phone_( J

In addition, I want to give $ to help support SCB.

Make check payable to: SOUTHERN CALIFORNIA BOTANISTS.

Mail check and form to:

Alan P. Romspert
Southern California Botanists
Department of Biological Science
California State University, Fullerton
Fullerton, CA 92634

CROSSOSOMA (ISSN 0891-9100) is published bimonthly (February, April, June, August,
October, and December) by Southern California Botanists, a California non-profit
corporation. Back issues of CROSSOSOMA are available for $2.00 an issue (plus 25<t
postage) or $8.00 a volume (plus $1.00 postage). Send a check with your request to
Alan P. Romspert, Treasurer, at -the above address. Manuscripts submitted for
publication should be addressed to Dr. Allan A. Schoenherr, Editor of CROSSOSOMA,
Division of Biological Sciences, Fullerton College, Fullerton, CA 92634.

19

till 11 II lutilililtiliiilili

SCB COMING EVENTS (DETAILS WITHIN)

.December

20

Cabri 1 1 o Museum

January

16

Torrey Pines State Park

February

13

Fungus Foray to Starr Ranch

Februa ry

21

Laguna Beach

February

27

Thousand Palms Oasis

March 5

Camp Pendleton

March 12

, 13

Guadalupe Canyon, Baja California

March 19

Santa Ana River

March 20

Annual Plant Sale

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