ne 4 inte ee f rte a ERR R. B. HINMAN COLLECTION PROFESSOR OF ANIMAL HUSBANDRY 4 1921-1943 New York © State College of Agriculture At Cornell University Ithaca, N. Y. aize; its history, cultivation, handling MAIZE ITS HISTORY, CULTIVATION, HANDLING, AND USES Photograph by Fred Coop of Pretoria GENERAL THE RicGHt HonourasBLe Lourts Borna, P.C. , Prime Minister and Minister for Agriculture, Union of South Africa. MAIZE ITS HISTORY, CULTIVATION, HANDLING, AND USES PITH SPECLAL REFERENCE TO SOUTH .4FRIC4A A TEXT-BOOK FOR FARMERS, STUDENTS OF AGRICULTURE, AND TEACHERS OF NATURE STUDY BY JOSEPH BURTT-DAVY, F.L.S., F.R.G.S. GOVERNMENT AGROSTOLOGIST AND BOTANIST, DEPARTMENT OF AGRICULTURE ) UNION OF SOUTH AFRICA WITH FRONTISPIECE AND 245 ILLUSTRATIONS LIONGMANS, GREEN AND CO. 39 PATERNOSTER ROW, LONDON NEW YORK, BOMBAY AND CALCUTTA I9I4 TO GENERAL THE Ricut Hon. LOUIS BOTHA, P.C. PRIME MINISTER AND MINISTER FOR AGRICULTURE OF THE UNION OF SOUTH AFRICA IN APPRECIATION OF HIS EFFORTS TO DEVELOP THE MAIZE INDUSTRY THIS VOLUME IS INSCRIBED No other plant we grow will produce 3,172 lbs. of digestible food on one acre of land at so little expense. No other cereal crop yields the farmer so large a return for his labour as Indian Corn. It is the king of the cereals. —Drrector C. S. Plums. vi PREFACE. THE materials for this book have been collected during a period of three or four years, but the book itself has been written during a Term, and part of the Long Vacation of 1913, spent at the School of Agriculture, Cambridge, where the author has been studying in- heritance of characters. He has endeavoured to adapt it to the needs of (1) the farmer; (2) students in the Schools and Colleges of Agriculture ; (3) teachers in the country schools who are endeavouring to interest their pupils in Nature Study. He hopes, also, that it will interest others concerned with the maize industry in its various branches, e.g. commerce, manufactures, and the supply of agricultural implements, machinery, and fertilizers. It is a difficult task to meet such diverse needs, and the result is necessarily open to criticism. The actual time available for its preparation has been too short, but if publication had not been completed before the author’s return to South Africa, it would have been postponed indefinitely, and in the present stage of development of the local maize industry, there seemed to be a need for a book of this character. The author is indebted to the following, among other gentlemen, for valuable assistance or contributions. The information on milling has been supplied by Mr. W. H. Horsfall of Aliwal North, and the chapter on the construction of silos by Mr. A. Morrison Hay, of the Public Works Department, Pretoria. For the VIL viii PREFACE chapter on maize and maize products as stock food, the author has drawn largely upon Feeds and Feeding, by Professor W. A. Henry, from whom he once enjoyed the privilege of a valuable course of lectures. Much of the information on insect-pests has been furnished by his friend, Mr. C. W. Mally, Government Entomo- logist, Cape Town, whose work in investigating and fighting the pests of the maize crop, while stationed at Grahamstown, is well known and highly spoken of by Eastern Province farmers. Much of the infor- mation on the use of maize-harvesting machinery in America, has been taken from a bulletin specially dealing with the subject by Mr. Zintheo, of the U.S. Department of Agriculture. Much valuable informa- tion has been obtained from the writings of Professors T. F. Hunt, C. G. Hopkins, Bateson, and Punnett, Dr. E. M. East, and Dr. G. H. Shull. To many kind friends and correspondents the author is indebted for the native names in use in different parts of South Africa; and to Mr. R. T. A. Innes, Director of the Union Observatory, Johannesburg, for much valuable information on climatology. The author’s warmest thanks are due to his sister- in-law, Miss Florence Bolton, A.B. (Stanford), and to his wife, for patient and careful revision of manuscripts and proofs, without which it would have been impossible for him to have prepared the book for publication in the limited time at his disposal. He is also indebted to Mr. H. R. Mallett of Cambridge, for the preparation and revision of the Index at the last moment, and to Miss Pate of Cambridge and her staff, for their care and accuracy in copying the tables and bibliographical list, and typing the manuscript, and also for reading some of the proofs. The author desires to express his thanks to those who have supplied photographs and blocks, or who PREFACE ix have given permission for the reproduction of illustra- tions from other books and publications; where the latter have been used credit has been given at the foot of the illustration. Several drawings were specially prepared by Mrs. Burtt-Davy, and many of the illus- trations have been reproduced from the 7ransvaal and Union Agricultural Journals, by the courteous permis- sion of the Editor, Dr. Wm. Macdonald. Grateful acknowledgment is made to those friends at Cambridge and elsewhere, especially to Professor R. H. Biffen, F.R.S., and Professor R. H. Punnett, for valuable suggestions and assistance, particularly in connection with the inheritance of characters; and to Mr. J. D. Anderson, M.A., and Dr. Nicholson, for assistance and information with regard to Hindoo and Persian names. Thanks are also offered to the Com- mittees, Members and Secretaries of the Liverpool, Mark Lane and Baltic Corn Exchanges, for their courtesy in obtaining and supplying information, and particularly to Mr. Broomhall, Editor of George Lroom- hall’s Corn Trade News, Mr. A. Grenville Turner of the Liverpool Corn Exchange, and Mr. H. M. Cole- brook of the Baltic and Mark Lane Exchanges, for their valuable assistance. JOSEPH BURTT-DAVY. ScHOOL OF AGRICULTURE, CAMBRIDGE, 28 August, 1913. PREFACE - 4 " a = = a > = CONTENTS OF CHAPTERS. ConTENTS - = - = = f z : e : = “ Ss List oF ILLUSTRATIONS - - # x . - 2 = 5 4 List oF TABLES : _ 7 # = # 5 2 = : CHAPTER I, II. Ill. IV, V. VI VII. VIII. IX Xx XI. XII. XIII. XIV. XV. XVI. XVII. IMPORTANCE AND History - < 3 2 = . S CuirimaTic REQUIREMENTS - - - - : - - - GEOGRAPHICAL DISTRIBUTION - * # . - 7 i BoTanicaAL CHARACTERS - 3 - 4 2 : a - INHERITANCE OF CHARACTERS AND IMPROVEMENT BY BREEDING - JUDGING AND SELECTION FOR EXHIBITION — - - - - . VARIETIES AND BREEDS - - - - . : # 2 : SOILS AND Wanvaus gta # 2 . z + - p TILLAGE, PLANTING,. AND CULTIVATION - * - - - - DISEASES AND PESTS OF THE MAIZE Crop - - - - - HARVESTING AND STORAGE, AND PESTS OF STORED GRAIN - - THE Conmmnas IN Maize GRAIN - - - - - - - THE MILuinc, MILL-PRopUCTS, AND CHEMICAL COMPOSITION OF Maize GRAIN - - - - - - 7 2 5 : Maize GRAIN aS Foop - - - - - - - - - THE PRESERVATION AND USE oF MalizE STOVER, Hay AND SILAGE, FoR Stock Foop - : = 2 5 _ CONSTRUCTION OF MODERN SILOS - = = = 2 = f Uses or Maize PRODUCTS IN THE ARTS AND MANUFACTURES - BIBLIOGRAPHY - - fe 2 « ¥ = ‘ = S INDEX - - a = im s = _ ‘ - " xiii XXIX XXXVil CONTENTS, CHAPTER I. IMPORTANCE AND HISTORY. Importance. pect ION —{—1. Importance of the maize crop. ' 2. What the American farmer thinks of it 3. Maize is the leading product of America 4. Amount and value of the United States crop 5. American maize is not grown for export 6. Maize is a white man’s crop 7. Maize is the staple crop of South Africa 8. Future possibilities of development in South Africa g. Relative importance of the worid’s maize and wheat crops =p History. —frio. Origin of maize “11. History 12. Introduction into Europe 13. Introduction into Africa 14. Introduction into Asia 15. Meaning and history of the botanical n name 16. The name Maize : 17. The word Corn 18. The word Mielie 1g. Other vernacular names CHAPTER IL. = oer = CLIMATIC REQUIREMENTS. . Climate : : . Factors which limit distribution z . Altitude . Temperature . Night temperature . Frost ‘ Hail . Soil temperature . . Moisture requirements . Rainfall . Sunshine : . Influence of climate upon vegetative characters and time of maturity : . Acclimatization ; . Influence of climate upon varieties E . Influence of climate upon chemical composition , Xili > o m ON ANNUM & XIV CONTENTS CHAPTER III. GEOGRAPHICAL DISTRIBUTION. SECTION F PAGE Be eek Geographical distribution . : : ; i . : - 46 \ 36. Distribution in the United States ; ‘ } : : ‘ . 48 37. The Sab-arid Zone. 2 : ‘ . : . i - 49 38. The Rocky Mountains Zone ; ; é ; ; : a - 49 39. The Great Basin ‘ : ‘ ‘ : . ; ; é . 49 40. The Pacific Slope , ‘ ‘ ‘ _ . 4 : ’ 2 49 4. The Atlantic States . . 7 , F ‘ i : : : 49 42. Canada : : ' . ; ‘ : : ' : : 50 43. Mexico : : : . : : : 50 44. Central America and the WwW ‘est Indies . : ; , : ; 50 45. Tropical South America. ‘ é y ’ * ‘ ‘ , SI 46. Argentina ‘ ‘ ; : ‘i - ‘ 51 47. Possible increase in the Argentine crop , ‘ ‘ F ‘ ~ §S 48. Europe : ‘ é , ‘: : - ; " : F » 52 4g. Asia. . : , : : ; ‘ R : : ‘ . 53 50. Australasia . : : ; F : , . ‘ i z » 55 51. North Africa ; é ‘ ‘ ‘ F : ‘ : : s) “56 2. Tropical Africa. (See also addendum at end of chapter) . ‘ = §0 53. South Africa ‘ ‘ ‘ : ; ¢ F : : « x $7 54. Orange Free State : : ‘ ‘ ‘ : : ‘ , - 58 55. Transvaal. 3 ‘ ; - x » 58 56. Relative yields of Transvaal Districts | : . : . ‘ » $9 57. Natal . ‘ . ; q ‘ é : A : 5 60 58. Cape Province . e % ‘ ‘ : F H 5 ; « 64 CHAPTER IV. BOTANICAL CHARACTERS. 59. Botanical relationship . ; Z : x . 7 : : » 65 \ 60. Description. é . : : ‘ : ‘ : ' ‘ » 65 — 61. Plant structure. : : ; : . : . . a> | 367. \ 62. Theseed . d : : : . . : «= 169 63. The embryo and endosperm : ; : : : : . a. 296 64. Germination ‘ : 4 ; : * i : i . ee 65. The maize seedling. i : ‘ ‘ 4 ‘ : : 2 66. The root and its functions . 3 ‘ : : : : i eee 67. The stem and its functions . : : : F : : : & OF 68. Sucker-shoots . ‘ ‘ , , $ i ‘ » 38 69. The leaf and its functions. ‘(See also addendum at end of chapter) . 80 7». The inflorescence 3 , f : 5 . , ’ ‘ & 85 71. Barren plants. 2 : E : : § ‘ : ; er 189 72. Flowering period F ‘ : ¥ , ‘ ; ‘ » go 73. The spikelet : : : : : : : : p 2) 495 74. The pollen and its vitality : : ; 2 : : F : e399 75. The young ear. : : . : 3 F ' : : . IO 76. Thesilk — . 4 : : : : : : : ; . - og mand Rey Pe Pollination . : : : : : ; , ‘ 3 : . 105 |—y8. Fertilization : : ; ; : ‘ : : : ‘ . 105 79. Dichogamy . i i i F . 107 80. Form tor describing ce maize plant i in ithe field - F . r » Tog 81. The shank . : : . E . : : ; : : - 10g 82. The husk. ‘ ‘ : z : P : . : : . Ito 83. The mature ear . F , ‘ i ‘ ‘ , c p . IE 8;. The cob : ‘ A ‘, fi . é . r « It3 85. Number of rows of grain. ; 3 : : i ‘ : Paar a $6. Twisted rows. : : : ‘i : 4 : : : . IIS CONTENTS SECTION 87. 88. 8g. go. gI. 92. 93- 94- 95: 96. Number of grains per ear Proportion of grain to ear Form for describing the ear The grain The hull The aleurone layer The endosperm Form for describing the grain Tubular glands in the embryo Apogamy - is CHAPTER V. INHERITANCE OF CHARACTERS AND IMPROVEMENT BY BREEDING. Necessity for Improvement. The object of breeding . The necessity for improvement of crops . Need for increase in the yield per acre . The cause of poor yields . Importance of a perfect stand . Importance of increasing the size of the ears . Average weight of grain per ear . : . Need for increase in the weight of grain per ear. . Percentage by weight of grain and cob . Effect of depth of grain on yield . : . Increasing yield by increasing the number of rows at ‘the butt and tip . Effect of width of sulci on yield . . Effect of shape of grain on yield . . Effect of number of rows . Effect of diameter of cobs . . Need for earlier-ripening breeds . . Drought resistance . Disease resistance A . Loss from weak stalks, shanks, or ¢ cobs . Necessity for the production of is seed . Other desirable points . Necessity for development of new breeds Inheritance of Characters. . Fluctuations : . Characters may be inherited . Importance of a as al of the laws governing the transmission of characters . Inheritance of characters in maize follows Mendelian Law . Reproduction and transmission of characters . Mechanism of transmission . The zygote . The homozygote . The heterozygote . Unit-characters . . Allelomorphic pairs of unit characters . . Dominant and recessive allelomorphs . Interaction of unit-characters . Repulsion and coupling of characters. . Xenia. . Splashed purple colour of the aleurone layer . Gametic segregation : 5 XV PAGE 116 116 118 119 122 122 123 123 124 124 126 127 128 130 131 135 137 137 148 149 I5I I51 I51 152 154 154 156 156 157 158 158 159 160 160 160 I6r 162 163 163 163 164 164 170 r72 173 174 175 179 180 xvi CONTENTS SECTION 136. 137. £38. 139. 140. Iq4I. £42. 143. 144. 145. 146. a7. 148. 14g. 150. I5I. 152. 153- 154. I55- 156. 157. 158. 159. 160. 161. 162. The reason for segregation in mathematical bigpor ion? Monohybrid ratios : : Dihybrid ratios Trihybrid ratios . Inheritance of colour . Yellow endosperm White starchy endosperm Inheritance of characters which affect ‘the growing plant Pericarp colour . : , Somatic variation in pericarp colour Silk colour . 2 Red cob-colour . Glume colour. Development of “ pods” Inheritance of ligule and auricles Physical condition of the starch . Size characters Inheritance of height of plants Inheritance of abnormal dwarfness Inheritance of length of ears Inheritance of size and weight of grain Inheritance of row numbers Four-rowed ears . Inheritance of fasciated and lobed ears Inheritance of laterally-branched ears Striped leaves Difficulties encountered in studying inheritance in maize Methods of Plant Breeding. . A few general principles . Methods of plant breeding . . Selection of parents ; . Effect of inbreeding . Improvement in yield by use e of first- generation c crosses . Fundamental points of seed selection . : . Correlation of characters . Desirable stalks . . Desirable leaves . . Desira’sle ears . Desirable cobs . Desirable grains . . Fancy points F . Methods of selection . Importance of care in selection . Field selection of parent ears . Seed-room selection of ears . Character of the grain . Selection by continuous performance- record . Method of propagation : ; . The breeding plot . Devices to prevent or detect cross- s-pollination . Production of new types by artificial cross- pollination . Reciprocal crosses ; : . Method of cross- pollinating 3 ; ‘ . Collecting the pollen . : e : . . . . Covering the silks ‘ < ‘ A . . . . The F, plants. < - ‘ : é . . The F, plants : - . Improvement by breeding i is s slow at first ese: Wed rel cel : z : . ‘ ‘ : : + 300 247. Ladysmith . 4 7 * : ; : 5 : + 301 248. Natal White Horsetooth _ : F 7: ° “ P + 303 249. Eureka : ‘ : 3 ‘ . é - F + 304 250. Chester County . i : 3 : . ‘ : : : + 304 251. Yellow Hogan . eae: : : 2 2 : Bees - 306 252. Golden Beauty . . wwe 06 253. Yellow Horsetooth . : : : : : : ‘ : - 306 254. Reid Yellow Dent : 3 : : é ‘ : : : - 309 255. Minnesota Early . ‘ ‘ s : : i : , : + 309 256. Star Leaming . : : : : . : : : : + 31r 257. Golden Eagle. ‘ ‘ : . . 312 258. Principal American breeds of flint maize 6 ‘ ‘ : 5 - 314 259. Principal South African flint breeds. ‘ : ‘ é : - 314 260. Cango, white. : : A : : » 315 261. Thoroughbred, Rural . : : d : : : : : - 316 262. Cango, yellow . : B : : : ‘ e . 3 - 317 263. Wills Gehu ‘ 5 rf : i : F i F 5 « 317 264. North Dakota. ‘ : : : ‘ ‘ ‘ A : * 317 265. Botman, white . . z : , , n , : ° - 320 266. Botman, yellow . 5 é 3 s 3 : : ‘ : + 820 267. New England 8-row . F : : . : . : : » 320 268. Burlington Hybrid ; . : jr . , ‘ 5 i « “$22 269. Gillespie Yellow . ; : ‘ ‘ : F : : Z - 322 270. Indian Pearl ‘ F : § 2 322 271. Principal breeds of soft maize or flour corn ; ; : . + 322 272. Brazilian flourcorn . ; R ‘ + 9323 273. Principal breeds of sugar maize grown in America : : : » 323 274. Sugar breeds introduced into South Africa . : : : ‘ + 323 275. Clark Favourite . ‘ s : x . 5 ‘ . F - 324 276. Arcadia sugar-maize . ; é . ‘ . é . 2 ROA 277. Claret sugar ‘ , i ‘ . ‘ ‘ ‘ ‘ ; » 324 278. Union sugar : 2 : ‘ . 6 4 a . : - 326 279. Golden sugar é 3 i ‘ z . 4 : > : » 926 280. Pop-corn . : . : iS g : : ‘ 5 . 326 281. Special-purpose sorts a 5 : : _ : 2 ‘ i - 326 282. Silage breeds . : ; : on B27 283. Classes best suited for ‘cultivation i in South “Africa : : Z « 328 284. Relative length of growing season of different breecs . ‘* e « 396 285. Breeds suitable for the High-veld ; 331 286. Breeds suitable for the Maize-belt of the Transvaal and Orange Free State. 332 287. Breeds suitable for the Maize- belt of the “ Midlands ” east of the Drakensberg . : : 3 F - 332 288. Breeds suitable for the Coast- belt : A : : ‘ . 332 28g. Breeds suitable for the semi-arid western region 5 e . » 333 290. Breeds suitable for the upper Bush-veld .. ; : is : 2333 291. Breeds grown in Rhodesia . . 2 S i : - 333 292. Relative yields of breeds in the Transvaal : 2 : 5 : & 2333 293. Relative yields of breeds in Natal : . : E : + 339 294. Third season’s results, Cedara, Natal . : » 344 295. Relative weight of grain per bushel of different breeds : ‘ - 344 CHAPTER VIII. SOILS AND MANURES. 296. The soil . . . 5 - 346 297. Chemical elements of the soil required by plants p : - 347 298. Soil moisture. . : : : : : + 347 CONTENTS SECTION 299. 300. 301. 302. 303. 304. 305. 306. 307. 308, 309. 310. 311. 312. 313. 314. 315. 316. 317. 318. 319. 320. 321. 322. 323. 324. 325. 3206. 327. 328. 329. 330. 331. 332. 333- 334- 335- 336. 337- 338. 339. 340. 341. 342. 343. 344. 345. 346. 347- 348. 349. 350. 351. 352. 353- 354. Conservation of moisture by Bunge Dry-land farming : Irrigation . Available plant- food Recuperative power of soils Character of South African soils . Soils suitable for maize-growing . New v. old lands 3 Effect of tillage Effect of continuous cropping Maintaining the crop- ee power of the soil Summer fallowing é Rotation of crops Organic matter Use of leguminose green- manure crops Rotations with maize in other countries Some Transvaal rotations The functions of manures Manurial requirements of the maize crop Does the use of fertilizers pay? d Cost of fertilizers in the interior provinces . Residual value of manures . Stable and kraal manure : Artificial manures or commercial fertilizers Method of applying fertilizers Influence of season on efficacy of fertilizers Use of lime : : Indication of need of lime Kinds of lime Preparation of the lime Method of applying lime Phosphatic manures Superphosphate alone Bone-meal alone. Superphosphate and bone- meal mixed Basic slag alone . 3 Nitrate of soda alone . ‘ Superphosphate and nitrate of soda Manganese compounds Potassium . CHAPTER IX. TILLAGE, PLANTING, AND CULTIVATION. Time of ploughing Depth of ploughing Different soils require different treatment Preparation after a a Time of planting : Listing Use of planters Check-rowing Distance of planting ‘ Distance tests in the Transvaal . Distance tests in Natal 5 Distances tried in the United States . Planting distance for silage or fodder maize : Effect of thickness of planting on composition of the fodder Number of plants to an acre of ground at different distances Amount of seed planted per acre 392 393 393 393 395 XX CONTENTS SECTION 355+ 356. 357+ 358. 359- 360. 361. 362. 363. 364. 365. 366. 367. 368. 369. 379. 371. 372. 373- 374- 375- 376. 377- 378. 379. 380. 381. 382. 383. 384. 385. 386. 387. 388. 389. 390. 391. 392. 393- 394- 395- 396. 397- 398. 399. 400. Depth of planting : Planting behind the plough Planting before ploughing . After-cultivation . 4 Implements for weeding Power CHAPTER X. DISEASES AND PESTS OF THE MAIZE CROP. Plant Diseases. Brown rust of maize . Red rust of maize. ‘‘ White rust” or “ blight” Maize smut or “‘ brand” Leaf scorch or maize blight Ear-rots of maize Dothiorella . : Burrill’s bacterial disease of dent and : sugar maize Stewart's corn wilt Yellow foliage Chlorosis Physiological effect of drought Weeds. Weeds : Parasitic weeds . Non-parasitic weeds Perennial weeds . Annual weeds Volunteer maize . How weeds spread Plant less maize and produce more Cultivation . Effect of clean cultivation of the maize © crop Harrowing . . ; Fallowing : Rotation of crops The best time to kill weeds | ; ‘ Weed seeds do not all germinate at once Is-ona, witch-weed, or rooibloem Remedies for is-ona Early planting Manuring Clear the land of is-ona seed already there . Buy seed-maize from clean farms Animal Pests of the Maize Crop. The Chacma baboon . Monkeys Hares. The reed- rat The porcupine or vyster- vark Birds . - d ‘ Insect Pests. Insect pests of the maize crop PAGE 395 396 396 396 400 402 405 408 409 CONTENTS Xxl SECTION PAGE 401. Methods of combating insect pests of the maize crop . ; 5 . 436 402. Cutworms, Agrotis spp. . : : ‘ é ‘ i + 437 403. Remedies for cutworms ; ; ; : : : ‘ ; » 437 404. The maize stalk-borer : : : : : ‘ : : . 438 405. Life-history of the stalk-borer . : : ‘ é 3 , - 440 406. Parasites of the stalk-borer . : ; ‘ : . 441 407. Burning the stalks to destroy the stalk-borer , ‘ d ‘ - 441 408. Ploughing-under the stalks : : : : : - 442 409. Early and late planting to avoid stalk- borer. : : : : - 443 410. Trap-crops for stalk-borer . : ‘ : . 443 411. Ensiling and shredding maize as a remedy for stalk- borer . ‘ » 443 412. The striped beard-grub or ear-worm, Heliothis armiger Hubner - 444 413. Remedies for the striped beard- em : : : : : : » 444 414. The maize cricket : ; : : : : : : » 445 415. Locusts 4 F ‘ . ’ ¥ » 446 416. The tok-tokje, Psammodes Reichei S. ‘ é ; ; 4 - 449 417. Remedies for the tok-tokje . : F 3 : é . : . 449 418. Plant-lice . ‘ . : : : “ : i : . 450 41g. Rose-chafers ‘ : ; ; : : . : . ; . 450 CHAPTER XI. HARVESTING AND STORAGE, AND THE PESTS OF STORED MAIZE. 420. Maize harvesting . : : Z ; - 451 421. Best condition of the crop for harvesting : ‘ : : - 452 422. The best stage of growth for both grain and stover . > , » 455 423. The best stage of growth for fodder. : : : ‘ ‘ . 456 424. The best stage of Bee for ensiling . : : : : é - 456 425. Frosted maize . 7 . 458 426. Composition of the maize plant at different stages of maturity . 458 427. Composition of maize fodder at different stages of growth. 459 428. Comparative digestibility of maize fodder and silage at different stages of maturity : - 459 429. Feeding value of maize fodder at different stages of growth : . 460 430. Pulling , , : F : , ‘ ‘ é : . . 460 431. Topping. F : : ‘ ; y : . 461 432. Methods of harvesting for grain i : : : : : . 46 433- Husking by hand from the standing stalks ‘ : : : : . 461 434. Cost of hand-picking in the United States . : ; ; ? . 465 435. Cutting maize by hand ‘ , . 462 436. Does it pay to use machinery for harvesting the maize e crop ares . 463 437. Sled harvesters . : j ; : : ; : : . 464 438. Mechanical harvesters 5 . 4 F F . i . 464 439. The cost and efficiency of harvesters : : ; : : ‘ . 466 440. The maize binder : : “ ; ‘ j . 466 441. Estimated cost of using a maize binder : : ‘ : F . 468 442. The maize stubble cutter. : : 4 : : ‘ 4 . 469 443. Draught of maize binders . i : ‘i : 3 5 i . 470 444. Shocking maize . : : 5 : : ; : : F . 470 445. The maize shocker. : 3 é : ‘ , : : . 471 446. A maize shock loader . : . ‘ f i: : » 472 447. Husking shocked maize by hand ‘ 4 ‘ : i : . 472 448. Maize pickers. : : : 2 : ‘ - 473 449. Cost and efficiency oft maize , pickers : : , ‘ ; ; 7470 450. Hand-husking in America . ‘ : : : F f ‘ 4-470) 451. Combined husker and shredder . : ‘ ‘ : : . » 477 452. Combined husker and sheller_. . : : ; 3 ‘ . 480 453. Machines for shelling husked maize . ; F é ‘ F . 480 Xxli CONTENTS SECTION 454. 455- 456. 457- 458. 459- 460. 461. 462. 463. 464. 465. 466. 467. 468. 469. 470. 471. 472. 473- 474- 475: 476. 477- 478. 479. 480. 481. 482. 483. 484. 485. 486. 487. 488. 489. 490. 491. 492. 493- 494. 495- 496. 497. 498. 499. 500. 501. 502. 503. 504. Importance of drying-out the eran Loss of weight in drying Variation in moisture-content is not ‘identical with ‘loss « or gain in weight due to change of moisture-content Storage in the husk Storage of husked maize Storage of shelled grain Kaffir method of storage Need for public maize stores or silos it in South Africa . Yield of grain from a given measure of ears Country damage . Pests of Stored Grain. Losses accruing from storage of grain Insects injurious to stored grain . Weevils Angoumois grain- ‘moth Remedies for insect pests Rats and mice in maize stores CHAPTER XII. THE COMMERCE IN MAIZE GRAIN. Time of arrival of the South African crop Local markets : The mines trade . Consumption on the Kimberley Mines Cape stock-farmers The native trade Local prices . Classes of maize called for in the local trade Comparative local prices of maize classes Transvaal maize imports Rapid increase in production Importance of the export trade Oversea markets . European consumption Possibility of developing trade with Canada Egypt as a possible market . : India . Australia Prices in European markets Prices on the English market, 1880 to 1908 The high London prices of 1907-8 : Early export prices for South African maize Changing prices per quarter to prices per muid . Changing prices per 1,000 kilos to prices per muid Market reports Prices affected by the world’s supply and demand Some factors which control prices in the world’s maize market . The world’s supply of maize Early attempts at an export maize trade from South Africa Natal Government enterprise Reduction in freight rates Government control of export Effect of good prices in stimulating trade Cause of abnormal prices Natal shipments, 1907. PAGE CONTENTS SECTION 505. 506. 507. 508. 509. 510. SII. 512. 513. 514. 515. 516. 517. 518. 519. 520. 521. 522. 523. 524. 525. 526. 527. 528. 529. 530. 531. 532. 533- 534- 535. 536. 537- 538. 539. 540. 541. 542. 543. 544. 545. 546. 547. 548. 549. 550. 551. 552. 553. 554. 555. 556. 557. 558. 559. 560, 561. 562. 563. Transvaal and Orange Free State shipments Some difficulties encountered Inter-colonial conferences . F Pretoria conference, 7 and 8 January, ‘1903. Establishment of a clearing house at Durban Bloemfontein conference, 18 and 19 aan IgIo Annual maize committee ; Government facilities for export . Government regulations and railage rates American railage rates Amounts and sources of supply of South African 1 maize : exported, 1go6-12 ‘ Details of exports, 1gII and 1912 Monthly exports : Destination of maize exported South African ports of export Varieties and classes of maize exported Grading at the ports . . : Effect of grading at the ports Description of grades . Grader’s certificate Weevily maize Removal of eS maize at ports Re-bagging Marking grades on bags Uniformity in practice of handling and storing at wharves Bag handling of grain ‘ Quality of grain bags . Bulk handling. Time saved by bulk handling Saving in cost by bulk handling . Accumulation and storage at inland centres Transit silos and elevators . Payment to farmers Trucking in bags and in bulk Storage at ports of export : Electric belt-conveyors for bagged grain Wharf-shed storage charges Construction and capacity of elevators Cost of erection of elevators Firms of elevator engineers . Elevator systems . The working of elevators Elevator charges . Elevator certificates Qualifications for managership of local elevators. Heating of grain in the elevator . ‘ Heating caused by moisture Loss of weight due to heating Degree of dryness required for export . Conditioning wet maize : : Ocean freight Shipping Tonnage Importing ports of Europe | 2 : Bulk handling at ports of import British elevators . New silo and grain- handling plant at Immingham Dock New silo for the Manchester =hip Canal Co. . Continental elevators . XXIV CONTENTS SECTION PAGE 564. Canadian elevators . ‘ 5 ‘ . ‘ : . OLE 565. United States terminal elevators ‘ ‘ ; P . ; * “OLE 566. Argentine elevators . : » O42 567. Silos and grain-handling plant at Puerto Galvan, Argentina E . 613 568. Private ownership of elevators . i , , , te 569. Railway ownership and control of elevators : : ‘ ; . 614 570. Co-operative elevators i j ‘ ‘ 2 ‘ ‘ . 615 571. The European market ‘ ; i ; : : : : . 615 572. Sale by sample . : 5 ; : F : ; ; : : 617 573. Import duty : : : : » 617 574. Classes of maize required by the European market. ‘ ; . 618 575. Differences in market value of maize grades F i , ; . 619 576. Number of South African classes ‘ r : 3 i . 619 577. Standards of weight and measurement ri ‘ , : . 620 578. Relative weight and bulk of South African maize r : Z . 622 579. International trade in maize ‘ i 2 é A F i % 622 580. United States exports ‘ , ‘ . , x 7 : a. 023 581. American maize grades : . ; : F 2 62 582. Kiln-drying American maize for export ‘ ‘ . . ‘ « 027: CHAPTER XIII. THE MILLING, MILL-PRODUCTS, AND CHEMICAL COMPOSITION OF MAIZE GRAIN. The Milling and Mill-products of Maize. 583. Native methods of grinding ‘ : : : : : ; . 629 584. Modern milling methods. : : : : : ; é + 102 585. Condition for milling . : : : ‘ , : ‘ é - 630 586. Mill-products of maize s ‘ : : ‘ f : ; . 630 587. Hominy-mill products : : : : : : ; : . 631 588. Best sorts of maize for milling . : : 5 : . 633 58g. Construction of modern milling machinery . : F : ; . 634 590. Cleaning and i aes : ; ‘ ‘ ‘ A , - 636 591. Grinding . Zi : 5 . ? : ; : : + 637 592. The millstone process . : : : ‘ ‘ ‘ : : 3 637 593. The roller-mill process i : : ‘ 3 : é ‘ . 640 594. Loss in ae ‘ : ‘ ‘ . e ‘5 ; : « O42 595. Samp . : . ‘ ; : ; ; ; : ‘ . 642 596. Flaked maize. : : : d : : : : ; - 643 Chemical Composition. 597. Importance of a knowledge of the chemical composition. i . 643 598. The important chemical constituents of foodstuffs : : : - 645 599. Chemical composition of the whole ae : : : . : . 646 600. Protein : : : : : - 646 6or. Protein obtainable from roo Ibs. of maize grain : : : : . 647 602. The proteids of maize ia : : A A : : a: . 648 603. Zein . : : E : : : : . 649 604. Ether extract or “fat” ; ; : : : : : : . 650 605. re : ; ‘ : . ‘ : ; : : - 651 606. Ash. ‘ ; ‘ ‘ ‘ ‘i A z ‘ i . 651 607. Water : ‘ 3 : S F 3 4. O52 608. Physical composition of the grain : : ; - 653 609. Mechanical separation of the different parts for analysis : : . 654 610. Relative proportions of the parts of the grain. : : . 656 611. Chemical composition of the cs ea of the € grain 3 ; . 656 612. The tip cap : é F f : - 658 CONTENTS SECTION 613. 614. 615. 616. 617. 618. 619. 620. 621. 622. 623. 624. 625. 626. 627. 628. 629. 630. 631. 632. 633. 634. 635. 636. 637. 638. 639. 640. 641. 642. 643. 644. 645. 646. 647. 648. 649. 650. 651. 652. 653. 654. 655. 656. 657. 658. 659. 660. 661. The hull The embryo The endosperm The horny gluten The horny starch The white starchy parts Chemical composition of different varieties and breeds of South African maize : Chemical composition of different varieties of North American maize Composition of maize grain grown in different localities Relative feeding-value of maize, wheat, and other cereals . Composition of maize by-products. Composition of maize bran compared with wheat bran Digestibility of maize products : Actual amounts of protein, etc., obtainable from each part of the grain CHAPTER XIV. MAIZE GRAIN AS FOOD. The uses of maize grain For Human Food. Maize the staple foodstuff of the American aborigines Maize adopted as the staple food of the African races . Use of maize in tropical Africa, in 1795. An important article of diet of the American people Probable increase in demand among the white races . Advantages of maize as an article of food : Injurious effect of unsound maize Pellagra Variety of maize ‘preparations available Maize meal, corn meal or mielie meal Corn-flour, Oswego, Maizena, etc. Maize starch F Samp, hominy, and cerealine Stamped mielies . Whole or crushed maize as a ‘ cereal food” # : Corn-flakes, silver-flakes, corn-crisp, fanko, etc. . Tortillas and enchilladas Maize as a “‘ green” vegetable To keep maize on the cob Dried maize for winter use . Maize as a sweetmeat Glucose Maize stalks as a source of sugar and syrup Corn oil : Maize vinegar Maize juice Fermentation products ‘of the grain Beer . : A ‘ ‘ Whisky and gin . Coffee substitute . For Stock Food. Maize grain for stock food . Grain and pasturage Feeding maize on the cob Frequency of feeding grain. 673 673 675 676 677 678 678 680 680 684 686 688 688 688 688 689 689 690 690 691 601 692 692 693 696 696 697 698 700 700 700 Jol 702 704 795 XXV1 CONTENTS SECTION PAGE 662. Preparation of grain for feeding . : : : . . : » 705 663. Dry v. soaked maize . : ; F : ‘ : : 3 . 706 664. Maize-and-cob meal . ‘ : . a i f 5 4 . 706 665. Maize-cob charcoal. : : : : ? : : F . 708 666. Maize meal. ‘ . ‘ ‘ ‘ ‘i ‘ + 709 667. Maize meal for pigs. ; : ‘ : ‘ : : : » 710 668. Maize meal for lambs . z : 6 i ‘ - 710 669. Mill and factory by- products for feeding : : : : : + 7it 670. Maize bran . r . . : ‘i x Fok 671. Gluten feed. p 3 : g ‘ A 7 . A ‘: yee s 672. Gluten meal. ‘ : : ‘ s ” . : . 712 673. Maize ‘‘germ” . 4 i ‘ i " 2 5 ‘ A 2 713 674. Corn-oil cake ; , a ¥ : : . ® » 9I4 675. Germ meal or corn-oil meal ; 4 y ¥ : , . rahe: 676. Distillers’ grains . ‘ f : : : 3 . ‘ : . 714 677. Distillery slop. P ‘ ; : : f 4 , » 955 678. Brewers’ grains . : ‘ ‘ i 4 a ‘ ‘ : ye RLS, 679. ‘*New corn product” . P ‘ : ; : ‘ ‘ . . 715 680. Cerealine-feed . : % ; ‘ s : x é . 715 681. ‘*Hominy-chop” and ‘ hominy-feed ” : . : é 5 G15 682. Maize for dairy cows . : : : : ‘ z “ . = LO! 683. Maize rations for dairy cows i : , . j 4 » GI7 684. Maize for fattening cattle . ; . rs , : j « 9ig 685. Maize for work-oxen . F . Fi : , é ‘ , ~ yer 686. Maize for horses . . F * ‘ : r < ‘ é 5 (720 687. Maize for sheep . : . , ‘ F . . ‘ < « G24 688. Maize for pigs. : é : : : : ‘ . 726 689. Maize for ostriches and poultry , ; 3 : : : ‘ - 729 690. Manurial value of foodstuffs é é ‘ ‘ < s ; . 730 CHAPTER XV. THE PRESERVATION AND USE OF MAIZE STOVER, HAY AND SILAGE, FOR STOCK FOOD. 691. Loss of stock from lack of winter food 5 ‘ : . : a. 932 692. The remedy ; : : A - 2. 9733 693. The feeding-value of an acre e of m maize . : ‘ : ; * 734 694. Yield of dry fodder. ; : : é : . 736 695. Yield of green maize forage ‘and silage ; , : 3 : . 736 696. Food value of weeds . : 2 2739. 697. Forms in which maize can be preserved for stock food s 2 a, 937 698. Relative composition of maize stover, fodder, silage, and grain . 2 (937 699. Relative digestibility of maize fodder, stover, and silage. . 738 700. Amount of digestible matter in different parts of the maize plant . 739 7o1. Loss of weight and of feeding- value and other changes due to curing . 739 702. Losses in the silo : P : ; é ‘ : : . - 739 703. Maize stover ; ‘ ‘ . : d ‘ . z - 740 704. Stover for dairy cows . i F r : : ‘A ‘ : « F4E 705. Stover for sheep . i . i : Z 6 G42 706. Maize fodder or ‘“ shocked-corn’ lenge : : é . - 743 707. Maize fodder for dairy cows : : : : - A ‘ - 744 708. Maize fodder for bullocks : : : : : : : . 744 70g. Composition of dry maize fodder F : y : < - 745 710. Relative value of fodder from different varieties : : . » 745 711. Moisture-content of maize fodder and stover : , 5 é - 746 712. Green maize forage. . . 746 713. Relative value of green maize forage from different varieties 2 - 746 714. Maize silage : . < ‘i : : 5 < : P + 747 CONTENTS SECTION 715. 716. 717. 718. 719. 720. 721. 722. 723. 724- 725. 726, 727. 728. 729. 730. 731. 732. 733- 734: 735+ 736. 737- 738. 739- 740. at. 742. 7424. 743- 744: 745- 746. 747- 748. 749: 750. 751. 752. 753+ 754 755: 756. 757- 758. 759- 760. Maize for silage may be ee late Uses of silage : Silage for dairy cows . Silage for bullocks Maize silage v. Timothy hay Maize silage v. roots - Comparative farm value of maize ‘grown for silage and for grain. Cost of silage production Amount of silage required for feeding . The feeding of silage . : Silage feeding-table Importance of a ‘“‘ balanced”’ ration “ Nutritive ratios” of some foodstuffs Mixtures to increase the feeding-value of maize silage Kinds of silage : : : Composition of maize silage Changes in the protein due to ensiling Moisture-content of maize silage . Composition of maize silage compared with that of green maize forage Popular objections to silage : : Best breeds of maize for silage Planting-distance for silage or fodder-maize Best condition of the maize crop for harvesting . Methods of ensiling : : The addition of salt The modern silo . The stack silo The pit silo . The American cornstalk disease CHAPTER XVI. CONSTRUCTION OF MODERN SILOS. Historical Form . Size Capacity Position Materials m Reinforced concrete Floor . Walls . Plaster Roof . Doors . Chute . CHAPTER XVII. USES OF MAIZE PRODUCTS IN THE ARTS AND MANUFACTURES. Importance of maize products for manufacturing purposes . Starch : : ; ; Dextrine Corn-oil : Rubber filler XXVIl PAGE 749 749 749 750 75° 751 751 751 752 752 753 753 755 770 772 772 773 774 775, 776 778 778 778 780 780 781 782 783 785 785 735 XXVIII CONTENTS SECTION 761. 762. 763. 764. 765. 766. 767. 768. 769. 770. 771. 772. Glycerine . : De-natured alcoho ; : Gas for illumination and heating . Maize charcoal Paper material Cellulose : : ; Other uses for maize husks Other uses for maize cobs Maize chaff , “Zea” or maize-silk . ‘‘Maidis Ustilago ” Maize-straw for thatching PAGE 786 786 792 793 793 798 799 801 801 801 802 802 FIG. on NAN LIST OF ILLUSTRATIONS BY CHAPTERS. General the Right Hon. Louis Botha, P.C. oar) by Fred Coop of Pretoria) . : ‘ Frontispiece CHAPTER IL PAGE . A field of maize on the demonstration farms of Messrs. John Fowler & Co. (Leeds), Ltd., at Vereeniging. 2 . In a maize field in the Transvaal Bush- veld (Tzaneen, Zoutpansberg District) ‘ 3 . Fields of maize, Government Experiment Farm, Potchefstroom, Trans- vaal. (Photograph by Fred Coop of Pretoria) : ‘ 7. . Maize fields at Vereeniging, on the High-veld of the Transvaal . 8 CHAPTER II. . Effect of hail on the leaves of the maize plant . 3 , . a. 134 . Effect of hail on the stem and leaves of the maize plant. 35 Grains which have been fertilized but yet sa filled with starch, pro- bably owing to drought : : - 38 Grains cracked from exposure to unfavourate weather conditions 38 . Grains cracked from exposure to unfavourable weather conditions . 39 CHAPTER IV. . Teosinte, Euchlena mexicana L., a near relative of the maize plant, and the only species with which it is known to hybridize. = 106 . Maize plants inthe Transvaal. : : , z 3 OF . Flowering plants of maize (Zea Mays ba, eee by Dr. W. Macdonald) . : 68 . Plant cells, as seen under a * high- power microscope, showing strands of protoplasm, nucleus, nucleolus, etc. (From Sir F. Darwin’s Elements of Botany) . i : 3 - 69 . Transverse section through a leaf (of hellebore), showing tissues and cells. (From Sir F. Darwin’s Elements of eo) F ‘ . 69 . Embryo and endosperm of maize ; : - 70 . Germinating maize grains . : : : : : . : s OE . Maize grains planted upside down : : F f ‘ : e 93 . Maize seedlings at two stages of growth . : 3 : : - 74 . Part of a transverse section of maize stem. eS Cavers’ Practical Botany) ‘ : : é » ¥6 . Transverse section of vascular bundle of maize. (From do.) P - 76 . Part of a radial longitudinal section of maize stem. (From do.) =) 7. . Part of a tangential longitudinal section of maize stem. en do.) . 77 . Base of young maize plant . 5 > - 79 . Bisexual tassels of sucker-shoots, bearing small round grains b . 80 . Leaf sheath and base of blade. . : : . : : ey BOT . Leaf sheath : : : : - . 82 . Three stomata with surrounding epidermic cells. (From Sir F. Darwin’s Elements of Botany) . : ; : z ; , . . 83 . Tassel of Odessa maize 5 ‘ * < " 3 : : . 84 XX1X LIST OF ILLUSTRATIONS . Young ear showing silks ready for pollination . Young ears showing silks at different stages of development . Young ear with fully developed silk : . Young ear with husks removed . Bisexual tassel of a sucker-shoot . . Bisexual tassel (bearing small ears) of a sucker- shoot . . . Bisexual tassel on main stem . Bisexual ear . Bisexual ear . Silks injured by larvae . . Blind ear, on which the silks have failed to “develop . Tassel of sugar maize . . . Spikelets of the maize plant. (After De Vries) . Pollen grains of maize : . Effect of complete or partial lack of pollination , . Effect of partial pollination . 4 . Young ear showing homology of ‘husks : . Young ear showing ovaries and styles (‘‘silks”’) . . Two-lobed ear 3 . Bifid ear . Branched ear of Hickory King . Branched ear of Ladysmith . Pod maize (Zea seas var. tunicata) . The style or silk . ‘ . Diagram showing course of pollen tube through style to ovule. (After drawing by C. S. Ridgway, in Duggar’s Southern Field Crops) . The embryo-sac in maize at time of fertilization. (After drawing by F. E. Lloyd in Duggar’s Southern Field Crops, by permission of The Macmillan Company) . Four secondary ears developed from the nodes on the shank of a single ear . Secondary ear developing from a node of the shank . Maize plants developing two ears : é : : . Left-hand twist of rows i é . : : : . . Right-hand twist of rows * 5 : . . Enlarged section pee hull of maize grain. ‘(From Passmore and Webber) . Variation in shape of maize ‘grains. (From U.S. Department of Agri- culture Year-book) CHAPTER V. . ‘Nubbins” of Hickory King ; one cause of poor yields . Desirable and undesirable types of Hickory King . Increasing yield by increasing depth of ous . A desirable type for selection . . Increasing yield by increasing number of rows at the butt . . Result of breeding for reduction of sulci . Variation in shape and size of grain in the same breed . Shortening the growing season . Result of weak stalks . . A heterozygous F, ear . Segregation of characters after crossing P, and F, ears . Segregation of characters in F, ears : . Segregation of characters in the F; (seed) generation : all-black ears . . Segregation of characters in the F, (seed) aa black-and-white ears . Segregation of characters in the F; (seed) generation : "all-white « ears . . Part starchy, part wrinkled grain. . Xenia in colour 106. 107. 108. Tog. IIo. III. rig. 113. IIq. II5. 116. 117. 118. IIQ. 120. rai. LIST OF ILLUSTRATIONS . Diagram to illustrate segregation of characters . Somatic variation in pericarp colour 2 . Inheritance of size characters : t plant generation . Inheritance of size characters : F, plant generation . Inheritance of row numbers 3 . Inheritance of row numbers . Fasciated and lobed ears . Laterally branched ear . A, Desirable shape of grain; B, Device for standardizing grains. ‘ Selecting seed maize: the final selection , ; i . Selecting the best ears from the bulk plot a z 5 : . A new breed of maize in process of development. (Courtesy of the South African Railways Publicity Department) CHAPTER VI.' - Exhibit of the Division of Botany, Transvaal Department of Agri- culture, at the First South African Maize and Citrus Show, Johannesburg, 1910 . Maize Exhibits at the First South African Maize and Citrus Show, Johannesburg, 1910 . American students learning to judge m maize. CHAPTER VIL. AG Enea King, unusually a ear, but sulci too wide; B, Lady- smith . Hickory King ; A, defective tip; 'B, a ‘good average ear . Variation in ears of Hickory King . Ladysmith Hickory; a promising but unfixed cross-bred grown at Ladysmith, Natal . . 12-row Hickory or ‘* Hickory Horsetooth” . Salisbury White 100. Io. 102. 103. 104. 105. Noodsberg Horsetooth; a promising, unfixed cross-bred Mercer : é é F : : Virginia Horsetooth : ; ‘ é : : : . 5 A, grains of Mercer; B, surplus crop after filling Mr. Mercer’s barn 10-row Hickory or “ Louisiana” Two prize ears of 10-row Hickory, at ‘the First South African Maize Show. A, grown by Reynoids Bros.; B, grown oie Hutchinson and Shaw (Val Station, Transvaal) : : : : Iowa Silver-mine, a prize ear Two prize ears of Iowa Silver-mine, at the First South African Maize Show. A, grown by W. A. McLaren (Vereeniging) B, grown by M. Geerdts (Boksburg, See) A é é Boone County : Variation in types of Ladysmith Natal White Horsetooth ; short type . Natal White Horsetooth x Hickory King « A, Eureka; B, Chestey County . A, Golden Beauty ; B, Yellow Hogan Yellow Horsetooth or German Yellow Reid Yellow Dent Minnesota Early . Star Leaming : Golden Eagle, as grown in Rhodesia . Cango, North Dakota, and Wills Gehu . A, New England 8-row ; B, Improved Yellow Botman : é A, Gillespie yellow flint (Red: ek ewe Oe W. ee Zand- spruit, Transvaal . : XXXII LIST OF ILLUSTRATIONS FIG. 122. 123. 124. 125. 126. 127. 128. 129. 130. 131. 132. 133. 134. 135. 136. 137. 138. 139. 140. I4I. 142. 143. 144. 145. 146. 147- 148. 149. 150. I5I. 152. 153. 154. 155. 156. 157. 158. PAGE Brazilian flour corn; grown by F. le Roux, Volksrust, Transvaal; prize ear at the First South African Maize Show, rg1o : + 321 Arcadia Sugar-maise ; improvement by See . : . + 325 Pop-corn, Zea Mays var. pre@cox : : 3 : : - 326 CHAPTER VIII. Effect of basic slag on the maize crop 368 Effect of growing maize without manure ‘(on plot adjacent to that shown in Fig. 127) : : : - 369 Effect of superphosphate and nitrate of soda mixed. . : + 370 Effect of manganese compounds on maize . : : : 3 + 371 Effect of sulphate of potassium on maize . ¢ : : z - 372 ‘CHAPTER IX. Primitive method of preparing land for maize, in Zululand . ‘ 2 295 Deep ploughing by steam; 12 furrows at a time by direct traction . 375 The “ Fowler”’ direct traction engine (for oil or coal) . 3 : 3370 Disk cultivating ; double-engine system. ‘ ‘ ‘ é 2 378 Harrowing by steam; double-engine system : * - 378 Zigzag harrow. (Courtesy of Messrs. Malcomess & Co., Ltd.) . - 379 Riding cultivator. (Courtesy of do.) . : F : : 5 - 379 Riding disk cultivator. (Courtesy of do.) . : ; : : - 380 Spring-tooth harrows. (Courtesy of do.) . : ‘ : : . 350 Harrowing the young maize plants, Vereeniging 381 Part of the gang of fifty-two ‘‘Champion”’ planters at work on Messrs. John Fowler & Co.’s demonstration farms at Vereeniging . » 3382 Fowler’s steam planter , & : , ‘ ‘ 3 : » 383 Combined lister and planter : : : : : : . 356 A, a home-made hand planter for maize. ae ee The Book of Corn) . : j = 387 B, home-made marker for planting 1 maize ’ by hand. (After Myrick, The Book of Corn) x : : . ; ‘ ; « 3% Maize planter, ‘‘ Moline Champion ” : ‘ : ‘ ; A . 388 Cultivating maize lands in Kaffraria . len Steam- cultivating maize plants 36 inches high, with spring- ‘tooth cultivators doing 6 rows at a time. (Courtesy of Messrs. John Fowler & Co., Ltd.) . i : ‘ ‘ 3 ‘ a . 398 ““New Western” cultivator. (Courtesy of Messrs. Malcomess & Co., Ltd.) . ‘ . . ‘ ‘ “ < ‘ : , - 399 Anti-clog weeder . “ ¥ ¢ A . , - 400 ‘Captain Kidd”’ cultivator. (Courtesy of Messrs. Malcomess & Co., Ltd.) . 5 : 3 . 400 “ Single Dutchman ” cultivator. "(Courtesy of do. >) : : : . 401 Adjustable cultivator. (Courtesy of do.) . : : , . 401 Adjustable weeders. (Courtesy of do.) ‘ 402 Steam-ploughing and subsoil packing the land at Vereeniging ; double-engine system. (Courtesy of Messrs. John Fowler & Co., Ltd.) . : z 403 “Lucky Jim” weeder. An implement for cleaning the growing crop 404 “Red King” cultivator. Another implement for cleaning the grow- ing crop : ; : : ‘ : ‘ 3 . : - 404 CHAPTER X. Brown rust of maize . 5 ; : z < : + 407 Maize smut or brand on the. tassel : : : : : - 410 Maize smut or brand on the ear . FIGe 159. 160. 161. 162. 163. 164. 165. 166. 167. 168. 169. 170. 171. 172. 173. 174. 175. 176. 177. 178. 179. 180. 181. 182. 183. 184. 185. 186. 187. 188. 189. Igo. IgI. 192. 193. 194. 195. LIST OF ILLUSTRATIONS XXXi11 A and B, dry-rot of maize, Diplodia Zea— A. ear showing white hyphze , B, grains showing small black fruiting bodies of the fungus i Rooibloem, Striga lutea Lour. Rooibloem, Striga lutea Lour. (Drawn from ‘plate by Mr. Claude Fuller, in first report of Government Entomologist of Natal) Chacma baboon, Charopithecus porcarius. ee Transvaal Agricul- tural Fournal) ; ? : . Jumping hare or spring- -haas, Pedetes caffer. (Photograph from specimen in Cambridge University Museum of Zoology) Maize ear damaged by small birds : : 5 : : : Maize stalk-borer, Sesamia fusca Hampson. (From coloured plate by McManus, illustrating article by C. W. sila in ac a Mala tural Fournal) Striped beard-grub pupa in sheath at base of ear. . . Locusts. (Photograph by Exton, Pietersburg, Transvaal) . CHAPTER XI. Maize crop ready for harvest. (Courtesy of Publicity Department, S. African Railways) Harvested ears of maize carried to the headland ready for shelling Cane knife used for cutting maize , A device for cutting maize in the field. (After “Myrick, The Book of Corn) . 3 ; a F Another Mctaeeh for cutting maize in the field. (From Transvaal Agricultural Fournal) . : ‘ F McCormick maize binder. (Courtesy of Messrs. Malcomess & Co., Ltd.) : ‘ Maize binder at work in America Shocking maize in America Maize picker at work in America A : : . : ; : Deering combined husker and shredder. (Courtesy of Messrs. Malcomess & Co., Ltd.) Marshall & Son’s, sheller. (Courtesy of Messrs. D. E. Hochly & & Co, Steam traction sheller . Convertible hand or power sheller, suitable for small crops Native method of storing maize in trees, Swaziland ‘ Native method of storing maize in trees, Swellendam District Maize hock, Waterberg District . : Maize hock, Bechuanaland . ; Method of storing maize, Government Experiment ‘Farm, Potchef- stroom . Re-weighing and shipping stored maize, Vereeniging, Messrs. “John Fowler & Co.'s store Stacks of shelled maize at Vereeniging ready for market. * (Courtesy of Messrs. John Fowler & Co., Leeds, Ltd. ys . Drying maize, Marico District, 1904 Native method of storing maize, Zoutpansberg . 3 3 : Basuto ‘“ sesco” of woven grass, for storing grain. (Courtesy of the Director, MacGregor Memorial Museum, Kimberley) 3 Effect of angoumois grain-moth on maize ears CHAPTER XII. Grading maize for shipment, Durban . Re-bagging maize from small dealers, Vereeniging Granary and elevator . ‘ 3 F Spencer’s improved system of granary floor spouts. (Courtesy of Messrs, Spencer & Co., Ltd., Melksham) A : . c PAGE 413 413 423 425 428 432 434 439 445 447 454 462 463 465 465 467 468 470 474 477 478 479 480 483 484 485 485 486 487 487 489 490 491 495 569 578 579 580 XXXIV LIST OF ILLUSTRATIONS FIG, 196. 197. 198. 199. 200. 201. 202. 203. 204. 205. 206. 207. 208. 209. 210, 2ii. 212. 229. 230. 231% 232. 233. Automatic weighing and bagging off, from warehouse. (Courtesy of Messrs. W. & T. Avery, Ltd., Birmingham) . : : Loading trucks, Vereeniging ‘ : : : : . Grain silos at Puerto Galvan, Argentina ney of Messrs. Henry Simon, Ltd.) . ; ‘ . . Loading s.s. Dunluce Castle with maize, by cranes, at Durban. (Courtesy of Union-Castle S.S. Co.) : Grain conveyor-belts : 2 : f . Central granary, Milwall Docks, London. (Courtesy of Messrs. Goldstiick, Hainzé & Co.) . Steamer being discharged by travelling elevator, Victoria Docks, London. (Courtesy of Messrs. Spencer & Co., Ltd., Melksham) . Steamer being discharged by floating elevator, Surrey Commercial Dock, London. (Courtesy of Messrs. Goldstiick, Hainzé & Bey : A, drying wet maize, Durban wharves, 1909 : . B. maize dried and re-graded, ready for shipment The Hess grain-drier for conditioning grain Vertical section through Hess grain-drier : : . 5 i Granary and barge elevator on the Thames. (Courtesy of Messrs. Spencer & Co., Ltd., Melksham) . i ‘ . ‘ , Diagram of floating pneumatic elevator. (Courtesy of Messrs. Henry Simon, Ltd., Manchester) Diagram showing general arrangement of barge elevator and ‘quto- matic weigher. (Courtesy of do. ) Grain warehouses, London, fitted with Avery scales. (Courtesy of Messrs. W. & T. Avery, Ltd., Birmingham) Travelling ship elevator. (Courtesy of Messrs. Henry Simon, “Ltd., Manchester) ; Terminal grain silos, Bahia Blanca, Argentina. ‘(Courtesy ‘ot Messrs. Spencer & Co., Ltd. , Melksham) . . Interior of the Baltic Exchange, London. (Courtesy of the Secretary) CHAPTER XIII. . Zigzag separator. (Courtesy of Messrs. Samuelson, Banbury) . . Diagram of maize meal plant. (Courtesy of do.) . Dreadnought” grinder. (Courtesy of Messrs. W. S. Barron & Son, Gloucester) . Diagram of ‘“ Dreadnought ” grinder. (Courtesy of do.) ‘ : ‘ . Centrifugal dressing machine. (Courtesy of Messrs. Samuelson, Banbury) . Posser. (Courtesy of do.) - , : . . . Three-pair high roller mill for grinding maize. (Courtesy of do.) . Improved degerminator. (Courtesy of do.) : : 5 ‘ . Hominy separator. (Courtesy of do.) . Diagram of samp plant. (Courtesy of do.) . Diagram of flaked-maize plant. (Courtesy of do. ys . Physical composition of low-protein maize grain. (After Hopkins ; from Bulletin of Illinois State Agricultural Experiment Station) . . Physical composition of high-protein maize grain. (After do.) . Enlarged longitudinal section of maize grain. (From Hunt, The Cereals in America, Orange Judd Co.) . Hull, endosperm, and embryo. aca Myrick, The Book ied Corn, Orange Judd Co.) . CHAPTER XIV. Shangaan kraal, Zoutpansberg Zulus eating maize-meal pap Native women grinding maize Modjajie women stamping mielies 3 Zulu women carrying kaffir beer for a “ Beet. drink 3 PAGE 581 582 583 585 586 588 590 592 596 596 597 598 600 603 605 607 608 612 616 636 637 638 639 639 640 641 642 643 644 644 649 653 655 659 675 676 687 689 699 FIG. 234. 235- 236. 237. 238. 239. 240. 241. 242. 243. 244. 245. LIST OF ILLUSTRATIONS XXXV CHAPTER XV. PAGE Cattle feeding on standing maize stover, Transvaal . 5 5 + 740 Shredding stover in the United States 742 Shredded stover on an American farm 743 Filling silage-pit, Vereeniging. (Courtesy of Messrs. "John Fowler & Co., Leeds, Ltd.) . : A z 2 YGE Making stack- silage, Standerton District, Transvaal . 2 : - 764 Stack-silo, Springbok Flats, Transvaal : 2 - . 765 Cutting silage and filling pit, Burttholm, Vereeniging . ; - 766 CHAPTER XVI. Filling round silo in the United States r - fe ; F = (992 Filling twin tub-silos, Australia . , ‘ ‘ e af - 773 Filling square stone silo, Irene, ‘Transvaal ‘ s " 978 Plan of silo. (Courtesy of Mr. A. Morrison Hay) : 3 : - 779 CHAPTER XVII. Mats, etc., made of maize husks. (Courtesy of Ladies’ Home Fournal) : é . : e : 4 Z . : . 800 LIS] “OF “CABLES BY CHAPTERS. CHAPTER IL. I. Comparative table of mean monthly temperatures XVI. XVII. XVIII. XIX, XX, XXI. XXII. XXIII. XXIV. XXV. XXVI. XXVII. XXVIII. . Mean temperatures of the cereal belt of Argentina: November, December, January, and February . Mean monthly temperatures, Transvaal . Mean minimum temperatures at Government Observatory, Johannesburg . Mean minimum temperatures at Vereeniging . Mean minimum temperatures at Bethal . Rainfall of South Africa CHAPTER III. . The world’s maize crop of 1906 . Statistics of the maize-surplus States . Acreage under maize in India for ten years (1897- 1907) . Acreage and yields of maize in the Indian Provinces . Areas and yields of Transvaal Districts . Maize production of Natal, 1906-7 . Area and yield of Natal Districts . Maize production of Cape Colony, 1906- 7. CHAPTER IV. Variation in time of flowering Correlation of flowering and ripening period CHAPTER V. Summary of percentage stands of maize Analysis of yields of 134 ears of Hickory King Analysis of yields of 100 ears of Natal White Horsetooth Analysis of yields of 123 ears of Ladysmith Analysis of yields of 150 ears of Iowa Silver-mine and 10 ears of Chester County Analysis of yields of r10 ears of Champion White Pearl (a breed closely allied to Iowa Silver-mine) Analysis of yields of 170 ears of Yellow Horsetooth . é Analysis of yields of 352 ears of Yellow Hogan Analysis of yields of 200 ears of Golden King Analysis of yields of 370 ears of Wisconsin , Analysis of yields of 100 ears of Skinners Court 10 . XXXVii 93 94 134 138 139 139 140 I4I I4I 142 143 144 145 XXXVIiI TABLE XXIX. XXX. XXXI. XXXII. XXXIII. XXXIV. XXXV. XXXVI. XXXVIT. XXXVIII. XXXIX. XL. XLI. XLII. XLIII. XLIV. XLV. XLVI. XLVII. XLVIII. XLIX. L. LI. LII. LIII. LIV. LV. LVI. LVII. LVIII. LIX. LX. LXI. LXII. LIST OF TABLES Comparative weights and yields of grain of 1,684 ears of Transvaal-grown maize. Yield per acre from a 75 Hee cent stand, at various average weights per ear . A ‘ 2 ‘ é Percentage of grain to ear Variation in composition and yield of different ears selected for breeding : Distribution of gametes ina ‘a dihybrid ; Analysis of a dihybrid cross between white wrinkled and red starchy maize x . “ The distribution of gametes ina ‘trihybrid . Classified summary of Table XXXV . Summary of Table XXXVI Frequency distribution of heights of maize plants in a "cross (after East and Hayes) Frequency distribution of lengths of ears in corn (after East and Hayes) Row numbers in a family of Arcadia “Sugay-maize Inheritance of row numbers in cross-bred maize Inheritance of rows in a maize cross Performance-record of a breeding plot CHAPTER VI. For conversion of centimetres to inches, in measuring circum- ference of ears 5 CHAPTER VII. Summary of Potchefstroom breed tests Relative yields of maize breeds, Government Experiment Farm, Potchefstroom, from 1904-11 : Rainfall at Government ees Farm, Potchefstroom, 1906-7 to Ig0g-10 Relative yields of maize ‘breeds at Government Experiment Farm, Cedara, Natal, 1904-5. : Results of maize breed tests, Cedara, Natal, 1905- “6. Weights per bushel of South African shelled maize CHAPTER IX. Effect of treatment of soil on yield Effect of time of planting on yield Results of distance tests on yield of Hickory King Results of distance tests on yield of Iowa Silver-mine Results of distance tests in Natal * Number of plants per acre when sown ‘at certain distances, allowing one plant per “hill” . Effect of depth of planting on germination CHAPTER XI. Influence of maturity on yield Influence of maturity on composition . Influence of maturity on composition of fodder . Shrinkage in weight of maize stored on the cob Relative weight of sound and country damaged maize PAGE 146 147 148 159 183 184 185 186 186 194 198 201-2 204 205 226 262 334 335 338 340 342 345 377 384 390 390 392 394 395 457 458 459 482 493 LIST OF TABLES XXXIX CHAPTER XII. TABLE LXIII. Maize consumed by the De Beers Consolidated Mines, Ltd., during the twelve months ending 31 Dec., rgr2. LXIV. Mielie meal consumed in the compounds of the De Beers Con- solidated Mines, Ltd., Igi2 LXV. Variation in maize prices in South African markets - LXVI. Comparative local prices for different classes of maize LXVII. Natal production, import and re-export of maize and maize pro- ducts, 1904-6. : : LXVIII. Transvaal maize imports, 1907- 8. LXIX. Average yearly prices of American and La ‘Plata maize in London : LXX. Prices of South African maize in Europe LXXI. Comparative prices of South African maize in Europe LXXII. To change prices per quarter to prices per muid LXXIII. To change prices per 1,000 kilogrammes to prices per muid LXXIV. Amounts and sources of supply of South African maize, 1906-12 LXXV. Export of maize and maize meal, 1g11-12 : ‘ ; LXXVI. Graded maize exported ex each port, rg11 LXXVII. Monthly maize exports from Durban, 1907-8 LXXVIII. Destination of maize exported, 1908 LXXIX. Destination of maize exported, 1911 LXXX. Amounts of maize exported through the several ports in 1912 , LXXXI. Varieties and classes of maize exported from South Africa, rg11 LXXXII. Maize rejected by graders, 1g11r : LXXXIII. Number and capacity of United States terminal elevators LXXXIV. Table for reduction of bushels to muids . : : LXXXV. World's exports of maize in bushels LXXXVI. World’s imports of maize in bushels LXXXVII. Moisture-content of American grades CHAPTER XIII. LXXXVIII. Composition of parts of the grea as oe ed a the mill and by hand LXXXIX. Average composition of 1 maize ‘grain . XC. Distribution of protein in parts of the maize grain : XCI. Protein in roo lbs. of maize : XCII. Distribution of protein in roo Ibs. of maize XCIII. Distribution of fat in maize : XCIV. Distribution of carbohydrates in maize XCV. Distribution of ash in maize XCVI. Relative proportions of the parts of the maize ‘grain XCVII. Chemical composition of hull, embryo and endosperm XCVIII. Chemical composition of the physical parts of the grain. ‘ XCIX. Percentage distribution of the chemical components of the grain x : : * ‘ C. Percentage composition of the tip cap CI.- Percentage composition of the hull. CII. Percentage composition of the embryo CIII. Percentage composition of horny gluten CIV. Percentage composition of horny starch , CV. Percentage composition of white starch . CVI. Chemical composition of different varieties and breeds CVII. Protein-content of eighteen samples of Transvaal maize CVIII. Chemical a ea of different varieties of North American maize 4 : ‘ CIX. Composition of grain grown in different localities . CX. Chemical composition of the different cereals of the world PAGE 500 500 502 5093 504 505 512 514 515 516 517 558 560 562 563 564 565 566 567 579 611 62t 623 624 627 632 xl TABLE CXI. CXII. CXIII. CXIV. CXV. CXVI. CXVII. CXVIII. CXIX, CXX. CXXI. CXXII. CXXII1. CXXIV. CXXV. CXXVI. CXXVII. CXXVIII. CXXIX. CXXX. CXXXI. CXXXII. LIST OF TABLES Composition of some maize by-products . Comparison of maize and wheat products Composition of maize bran and wheat bran Digestible nutrients in maize products . Actual weight of protein, carbohydrates, fats and ash in x00 Ibs. of grain, and its distribution in each of the several parts of the grain ; CHAPTER XIV. Total digestible nutrients in 100 Ibs. of several cereals Comparison of the food-value of wheat bread and maize bread Sugar in maize juice at different stages of plant growth . Relative feeding-value of gluten meal and cotton-seed meal Theoretical rations for steers of 1,000 lbs. ; Fertilizing constituents of 1,000 Ibs. of certain maize products CHAPTER XV. Yield per acre of dry maize fodder . Yield per acre of green maize forage Relative composition of maize silage, fodder, stover, and grain Relative digestibility of maize fodder, stover, and silage Composition of the different parts of the dry maize fodder Relative value of fodder from different varieties Silage feeding table : . Nutritive ratios for different animals Nutritive ratios of different foodstuffs Composition of maize silage compared with that of green maize forage . . . i P CHAPTER XVI. Capacity of silos PAGE 668 669 669 670 672 678 679 693 713 720 731 736 736 738 738 745 745 754 75 755 760 774 CHAPTER I. IMPORTANCE AND HISTORY. Heer, of one grain of maiz, a reed doth spring That thrice a year five hundred grains doth bring. —SYLVESTER, translation of Du Bartas' Divine Weekes and Workes, i. 3. All around the happy village stood the maize-fields. —Hiawatha. Importance. 1. Importance of the Maize Crop.—Maize is one of the staple Cra food crops of the world ; the quantity produced is greater than that of any other sata and climatic conditions alone limit its more widespread cultivation. In those countries adapted to its production it is more extensively grown than any other grain. The total world’s crop reaches the extraordinary figure of 1,085,700,000 muids (3,875,927,000 bushels, or 1084 million Colonial tons). Of this more than 75 per cent is pro- duced in the United States, where the acreage is about double, and the total production about four times that of wheat. The whole of civilized Africa produces, at present, only about one per cent of the world’s supply. The following figures show the comparative world’s crop of the leading cereals for a single year :— Maize : : : ; 3,875.927,000 bushels. Oats . é ‘ : ‘ 3,532,470,000 i Wheat : : : : 3,428,998,000 * Rice . , 7 ‘ ‘ 3,203,782,000 ne Rye. . - ss T44325395,000 Barley ‘ x ; F 1,293,916,000 ee 16,767,488,000 + The domestic consumption of maize in the United States is at the rate of 7°14 muids (25°5 bushels) per capita of popu- I 2 MAIZE CHAP. lation, which is said (/7uzz, 1) to be the heaviest rate of I. consumption of any cereal by any people in the world ; it is nearly twice as much, according to population, as the con- sumption in Europe of all the other cereals. Maize is one of the easiest crops to grow, standing more rough usage than perhaps any other ; a favourite Kaffir method of planting is to scatter the seed broadcast over the unbroken veld and then plough the ground ; even with this crude treat- ment crops of 1} to 2 muids! of grain per acre are obtained. Fic. 1.—A field of maize on the Demonstration Farms of Messrs. John Fowler & Co. (Leeds), Ltd., at Vereeniging, Traansvaal. Davis (2), writing of maize and wheat in Argentina, points out that not only are they the two cereals requiring the least amount of capital, expense, and labour for their production, but that there is always an assured and immediate demand for them in the market. The farm value of maize must not be calculated solely on the yield of grain, important as that is, for its total yield per acre of vegetable matter is larger than that of almost any other 'r muid = 200 Ibs. avoirdupois. IMPORTANCE AND HISTORY 3 farm crop. Maize produces a large quantity of ‘“stover” (the CHAP. stalk and leaves after the ears have been removed), which is of | * considerable use for winter-feeding stock if properly harvested ; this should be taken into account in estimating the relative value of the crop. The husks, stalks, and cobs are also used for a number of manufacturing purposes, which are mentioned Fic. 2.—In a maize field in the Transvaal Bush-veld (Tzaneen, Zoutpans- berg District). in detail in a later chapter. As silage material, maize is one of the very best crops that can be grown, both on account of its heavier yield per acre and also because of the succulence and physical character of the plant, which render it peculiarly suitable to the process of ensiling. We still hear South African farmers say that maize is a I * 4 MAIZE CHAP. Kaffir crop, and that maize-growing does not pay the more I. ambitious white farmer. We hope to show in the following pages that, except where abnormal economic or unfavourable climatic conditions prevail, this is not the case when the crop is grown properly. 2. What the American Farmer Thinks of [t.—In view of the fact that the United States produces 820,000,000 muids of maize per annum—three-quarters of the world’s crop—and that this is not grown with cheap “native” labour, it may be well to look for a moment at the attitude of the American farmer towards the maize crop. In the United States it is a common saying that “Corn is King”. “Corn” in America is maize. The American farmer has earned the reputation of being a shrewd business man who does not conservatively stick to a crop whether it continues to pay or not, because his fore- fathers grew it; if anything, he is inclined to change too rapidly, and to “scrap” anything which he considers unprofit- able. If maize did not pay he would soon drop it; but we find on the contrary that he has 108,750,000 acres under maize although, in 1906, his maize crop paid him only 5s. 11d. per muid, and in 1896 the farm price was only 3s. 13d. per muid. 3. Marze is the Leading Product of America—Mr. T. N. Carver (1), Professor of Economics in Harvard University, boldly states, and then proceeds to demonstrate, that maize is the leading product of the United States of America and maize-growing its leading industry. Not only is it grown more extensively than any other cereal, but the maize crop of the United States considerably exceeds in value that of wheat and cotton combined. No other American product or group of products equals it in value. In 1899 the value was greater by about £5,000,000 (five million pounds sterling) than that of all the products of the great iron and steel industry. It is the staple grain crop in most of the States of the Union, and its culture maintains a larger number of American people than any other industry. We hear much about the American wheat crop, but comparison of American crops of wheat and maize shows that where the total value to the farmer of a crop of wheat is $9:07 (£1 17s. 10d.) per acre, the maize crop is worth $14°56 (£3 os. 8d.); or, if we add the value of the straw, stover, or fodder, the relative values are : IMPORTANCE AND HISTORY 5 wheat, $16°26 (43 7s. 9d.) and maize, g50°72 (45 IIs. 4d.)/ per acre. / 4. Amount and Value of the United States Crop.—Tl hie total production for all the North American States (not i:n- cluding Canada and Mexico) amounts to 2,927,416,000 (two billion, nine hundred and twenty-seven million) bushels, eind the farm value, at a shade above 6s. 6d. per muid, is cover £267,000,000. About 984 per cent of this immense crojp is wanted for domestic consumption and only the surplus 14; per cent is exported ; 14 per cent of such a crop is no small lace however, for it amounts to over 12,000,000 (twelve mil lion) muids. ) The acre value of the maize crop in the United /States varied in one year from 18s. 8d. on the poorer soils of / South Carolina, to 44 9s. 4d. in Rhode Island, where more irftensive agriculture is practised, and from £1 13s. 4d. to £2/ Los. in the Corn-belt, and that was a year when the farm /price of maize was only 35-7 cents per bushel (5s. 3d. per muifd). 5. American Matse ts not Grown for Export——Bly far the largest part of the maize corn produced in the /Corn-belt never leaves the farm on which it is grown, except ifa the form of a second product. Nearly every maize-growjer finds it more profitable to turn the major part of his crofp into beef or pork before it is sold. It is customary to bu/y up three- year-old steers, or flocks of sheep, raised on {che Western stock ranges, to fatten on maize corn and stover, through the winter; in the spring they are “finished off” ohn maize corn and green pasturage, and are ready for sale jon the stock markets of Chicago and other Western cities) in June and July. Hogs are raised on the spare milk of the farm, and when older are allowed to follow the steers and pick out the un- digested grain front the droppings, and are /finally fattened off on maize and sent to market. A certain anpount of the corn is eaten green as a boiled vegetable; som¢ is ground into “corn-meal” for domestic use; only a small a/mount of grain is left to sell for manufacture or export. It h/as been well said that maize is and always will be the King /of Crops, and the greatest of all cattle feeds. 6. Maize is a White Man's Crop.—Mlaize is essentially a white man’s crop, and Prof. Carver (1 doubts whether it CHAP. I. CHAP. 6 MAIZ “could be grown at all, as it is grown in the Corn-belt, if dependence had to be placed upon negro labour”, The labour employed in that part of the country is entirely white, earning about £5 per month and board the year round, Yet tliat maize pays under these conditions is evident on all sides ; it pays because it is a crop which can be handled almost en- tirely by machinery ; because the soil is in good tilth ; because the’ crop is kept clean; and, last but not least, because the farmier uses well-bred seed. It is an instructive fact that in the American Maize-belt the enormous aggregate of the crop is made up of the pro- ducts, of a large number of small or moderate-sized farms, running from 80 to 300 acres in size, and worked mainly by the owners themselves or by tenants who pay cash rent. The reason for this, Prof. Carver (1) concludes, is that maize- growing requires a higher class of farming than any of the other staple crops, and cannot be so successfully carried on with hired labour alone. It requires such close and conscientious attention that it is doubtful if large farms, where the work is done by hired labour, can ever compete successfully with the smaller farms where the owner or renter does the work himself, or at least has it done under his immediate care and attention. | With increased size of farm (as in the Western States), there is noticeable a general decline in the intensity of cultivati¢n and consequent yield per acre, for good culti- vation is essential to a good maize crop. 7. Maize ts the Staple Crop of South Africa,—Maize is not only the staple food crop of the South African Kaffir, it has become an important item in the diet of the white people; but more than this, it has also become the staple cash crop of the South African farmer. In one of the writer's first Reports to the Director of the Transvaal Department of Agriculture, he stated that “Maize is a crop eminently suited to the Trans- vaal; every farmer grows it to a limited extent, and a vast quantity could be produced if he knew how to dispose of it. By the application of capital and the use of proper machinery, the maize crop can be made extremely profitable.” Further observation and study not only confirm this view, but show that perhaps no \country in the world is better suited to maize-growing on ‘a large scale than South Africa; it has an IMPORTANCE AND SIIISTORY 7 ample average rainfall, at the right season of the year, and CHAP. phenomenally favourable winter weather for the natural pro- duction of the quality of grain most suitable for shipment, In fact the climatic conditions of a large part of the Orange Free State, Transvaal, Natal, Rhodesia, Basutoland, Swaziland, and the Transkei are all that could be desired for maize- growing. 8. future Possibilities of Development in South Africa.— European corn brokers have recently referred to South Africa as the future maize granary of Europe. Maize will always be Fic. 3.—Fields of maize, Government Experiment Farm, Potchefstroom, Transvaal. (Photograph by Fred Coop of Pretoria.) the staple cash crop of South Africa. As its value for stock food becomes better appreciated, the local demand will increase, and in this connection Earl Grey’s recent prophecy! of a coming shortage in the world’s beef supply is suggestive. At the present time the country has only begun to show that it is possible to produce good maize. The traveller is impressed with the enormous areas of fertile land, suitable for growing maize, which are at present untouched by the plough, virgin sod like the American prairies. So far the average yield has been low; but it has been clearly demonstrated that, by means , 1 At the “South African Dinner,” 1913. 8 MAIZE cuaP. of good farming and good management, it can be trebled and I. even quadrupled. The present low yields are considered to just about cover expenses of production; if the yield were more than doubled, therefore, maize-growing should pay, provided prices hold good and cost of export remains low. The exact acreage under maize in South Africa is not known, but it is grown on practically every occupied farm in the Transvaal Province. Many farmers are growing 200 to 1,000 acres each, and at least three have 6,000 acres under crop to maize, A good deal is also grown by Kaffirs, for their own use, both on native locations and on rented farms. The farms average about 5,000 acres each, but the area planted to Fic. 4.—Five miles of maize fields at Vereeniging, on the High-veld of the Transvaal. maize is often not more than 5 to IO acres per tarm, and sometimes less, There are 11,679 registered farms in the Transvaal, of which about one-half are occupied; allowing an average of 10 acres per farm, the total area in maize (outside of Native Locations) would be only 60,000 acres, and an average yield of four muids per acre gives, roughly, but 240,000 muids. Only a very limited area of the Transvaal seems unsuited to the production of maize, but if we take into consideration only the farms at present occupied, andallow 250 acres of maize to every 1,000 acres of land, by raising the average production to merely 5 muids (172 bushels) per acre, the IMPORTANCE AND HISTORY 9 Transvaal alone ought to produce without difficulty 35,000,000 muids, Owing to the dryness of the winter over the greater part of South Africa, the farmer is able to continue harvesting and shelling in the field up to the very day when he starts planting the new crop; in this respect he has an enormous advantage over the American farmer. The percentage of grain which is damaged by the weather is exceedingly small. The moisture content of the grain exported is some 5 per cent lower than that of the American-grown article. South Africa has, and is likely to have for many years, an excellent local market for a large part of her crop. Because she owns her own railroads she can carry her surplus to the coast at cost. With these great advantages in her favour, South Africa has good reason for optimism as to the future of her maize industry. There is good ground for the prophecy that South Africa is to become the maize granary of Europe. 9. Relative Importance of the World's Matse and \Vheat Crops.—There is a popular idea that wheat is a more profit- able crop than maize. This is due to the fact that in the inland provinces of South Africa the price of wheat is more than double that of maize, and the former yields a heavier crop than the latter. But if the wheat acreage were much increased the price would fall; present prices are quite abnormal, the average farm price in the United States during the last ten years was only 77 cents per bushel or ros. 8d. per muid ; when South Africa produces enough for local needs her wheat prices will probably fall to those prevailing in the States. The difference in average yield is due partly to the fact that wheat is at present grown on the best alluvial lands, and, in the Transvaal at least, practically all of it under irrigation, while much of the maize crop is produced on newly broken veld, and all of it as a dry-land crop. When the maize lands are in better “ heart,” the average yield will probably be doubled, and even now the best maize crops are nearly double those of wheat. History. 10. Origin of Maise.—Researches into the history and geographical distribution of the maize plant show clearly that it originated in America. CHAP. 1. CHAP. 10 MAIZE Darwin (2) considered that maize “is undoubtedly of American origin”. Humboldt (1) observes that maize was found by the Euro- pean discoverers of the New World from the south of Chile north to Pennsylvania. Prescott (1) adds that he might have given its known range as to the St. Lawrence as “our Puritan fathers found it in abundance on the New England coast wherever they landed”; he cites as his authorities Morton, New England's Memorial, page 68 (Boston, 1826), and Gookin, Massachusetts Historical Collections, chapter Ul. Prescott (2) states that maize was “the great agricultural staple of both the northern and southern divisions of the American Continent ; and which, after its exportation to the Old World, spread so rapidly there, as to suggest the idea of its being indigenous to it”. . . . “The misnomer of d/¢ de Tur- gute shows the popular error. Yet the rapidity of its diffusion through Europe and Asia, after the discovery of America, is of itself sufficient to show that it could not have been indi- genous to the Old World, and have so long remained gener- ally unknown there.” Alphonse de Candolle, the famous Swiss botanist and historian, who made a special study of the origin and history of cultivated plants, came to the conclusion, as long ago as 1855, that “ maize is of American origin, and has only been introduced into the Old World since the discovery of the New. I consider these two assertions as positive.” Twenty-seven years later he reiterated this view, and added: “The proofs of American origin have since been reinforced. Yet attempts have been made to prove the contrary, and as the French name, d/¢ de Turgure, gives currency to an error, it is as well to resume the discussion with new data. . . . From all these facts we conclude that maize is not a native of the Old World. It became rapidly diffused in it after the discovery of America, and this very rapidity completes the proof that, had it existed anywhere in Asia or Africa, it would have played an important part in agriculture for thousands of years” (De Candolle, 1). He concludes that circumstantial evidence points to New Granada as the original home of the plant, and suggests that the Chibchas, who occupied the table-land of Bogota at the 1 Cf. Prescott (2) as quoted above. IMPORTANCE AND HISTOR V 11 time of the Spanish conquest, and considered themselves aboriginal, may have been the first to possess and cultivate maize. Some later botanists are inclined to consider, how- ever, that it is of Mexican origin. The maize plant is not known to exist in a truly wild state, i.e. reproducing itself spontaneously from self-sown seed ; no plant has been found which can be looked upon as the true parent form, unchanged by cultivation. Some botanists are inclined to think that maize is a descendant of the Teosinte plant of Mexico, Euchlena mexicana, with which it can be hybridized ; or that the two had a common prototype (Easy, 5). 11. Azstory—Maize has been cultivated by the inhabi- tants of North, Central, and South America since prehistoric times. The early American explorers found the Indians cultivating it; Columbus, writing to Ferdinand and Isabella of Spain, mentions maize fields eighteen miles in length. Hochelaga, which later became the city of Montreal, was situate in the midst of large maize fields when Cartier visited it in 1535 (De Candolle, 1). Hakluyt (1) quaintly and eraphically describes the new cereal as ‘a corne called maiz, in bignesse of a pease, the eare whereof is much like a teasell” Maize-grain has been found in the Inca cemetery at Angon, Peru, which is nearly contemporary with the discovery of America (De Candolle, 1). It was, even in those days, a staple crop from the valley of the La Plata to that of the Mississippi. Investigations show that it was grown by the Chibchas of New Granada, the Mayas of Central America, the Nahuas, and their successors the Toltecs and Aztecs of Mexico, and by the Incas in Peru. De Candolle concludes that though these civilizations date at earliest from the beginning of the Christian Era, the cultivation of maize was doubtless still earlier. After an exhaustive survey of the philological proofs of its origin, Harshberger (1) concludes: (1) that maize was intro- duced into the region now comprised in the United States, from two sources—from the tribes of Northern Mexico and the Caribs in the West India Islands; (2) that the Pueblos and Northern Mexican tribes derived maize from Central Mexico ; (3) that tribal connections existed between the North and South American continents, and that an interchange of products was CHAP. CHAP. I, 12 MAIZE carried on by way of the Isthmus of Panama; (4) that the wild tribes living along the Andean system and in the El-Gran- Chaco and elsewhere used Peruvian loan-words for maize ; (5) that South American words for maize extended throughout the Greater and Lesser Antilles and Florida, and that the Arawak word for Indian corn, adopted by Christopher Columbus, was used by tribes of that stock in the impenetrable and luxuriant Brazilian forests. On the shores of the island of San Lorenzo, Peru, Darwin (1) found “the head of a stalk of Indian corn,” embedded among shells and sea-drifted rubbish with some bits of cotton thread and plaited rush similar to those obtained in ancient Peruvian huacas or burial mounds. The stratum in which this was found had been elevated to a height of 85 feet above sea-level, and was itself overlaid by other strata, containing shells, etc., and having a thickness of over 85 feet, indicating great antiquity. 12. Lntroduction into Europe,—De Candolle (1) finds that maize was unknown in Europe at the time of the Roman Empire. Fée (1) states that from the year A.D. 1500 maize had been sent from America to Seville for cultivation. From Spain it was introduced into France and Italy, Turkey and other parts of Eastern Europe. 13. /utroduction into Africa.—The wide distribution and extensive use and cultivation of maize on the African conti- nent have led many to suppose that the plant was indigenous, or at least in use from time immemorial among the aboriginal peoples. But this was not so. Burchell (1), who visited what is now British Bechuanaland in 1812, makes no mention of maize in his account of the crops cultivated by the Bechuanas, though he notes its use and cultivation in Griqualand West, at the Mission Station at “Klaarwater” and at “Jan Bloems Kraal” in the Asbestos Mountains, in 1811. It was there grown for poultry food, but he also notes that “ the half-ripe heads, when boiled, made a very agreeable and wholesome dish”. He notes that it was planted in the first week in October and came into flower before the middle of December. Visiting Burchell’s old camp at Litakoon, Bechuanaland, in 1912, the writer met a native who remembered Moffat, and the introduction of maize into Bechuanaland by the mission- aries ; now it is one of their staple crops. IMPORTANCE AND HISTORY 13 In the tenth century, according to Abn Zeyd Hassan and “Suleiman the Merchant,” the Zeng peopies, the progenitors of the eastern branch of the Bantu-speaking races now south ‘of the Zambesi, who were then located north of that river, in the country round Zanzibar, grew szzllet, which was their chief food (Tooke, 1). Giant millet or kaffir corn (Sorghum vulgare) appears to have been the former staple food-stuff of the natives of much of temperate South Africa. North- wards in the Bush-veld this was supplemented, and in places perhaps replaced by pearl millet or m’nyouti (Pennisetum Spicatum), and still farther north, within the tropics, rapoko (Eleusine Coracana) was largely used. Writing on the bearing of Bantu philology on early Bantu life, the Rev. Father Norton (1) states :— “We find a tradition that sweet-reed and millet, or kafir- corn, were given to the first human couple; maize, on the other hand, was introduced in historic times by the Portuguese to the Becoana, to eke out their scanty list of cane, pumpkin, beans, melon. Our old centenarian told me that mealies ap- peared in Modderpoort district together with the missionaries.” Mr. Allister M. Miller of Mbabane, Swaziland, who pro- bably knows as much, if not more, about Swaziland than any other man living, writes that from inquiries he made many years ago, he is of opinion that maize was introduced into Swaziland about the time the Hlamini clan, the conquerors of that territory, crossed from Tongaland, say the end of the eighteenth century. They do not call it by their word for “food,” mabela, but by the Zulu words m’lungu, meaning “ white man,” or m’bila, the Zulu name for maize. Maize would easily be carried from the north shores of the Mediterranean to the ports of Northern Africa, and probably * reached the latter from Spain and Italy, with which countries there was much commerce in those days. In 1623 Caspar Bauhin referred to the occurrence of pod-maize (Zea Mays L. var. ¢untcata St. Hil.) in Ethiopia under the name of manigette. But its introduction into other parts of the African conti- nent is traceable to the Portuguese, who were great voyagers in the sixteenth century ; they had colonies in Brazil and in the CHAP. 14 MAIZE cHap. East Indies and had established settlements on the African I. coasts in 1450. It is instructive to note that in Angola maize was at one time known by the name “blé portugais” (Portu- guese wheat), which suggests its source of introduction. They may have introduced it to West Africa with the object of fur- nishing food on the voyage across the Atlantic for the slaves whom they took from Africa to work their Brazilian planta- tions; one of the Brazilian names of the plant—vlho de Guine—suggests this. Moodie (Records) mentions the follow- ing interesting fact under the date 1658: “ As the season for sowing Dutch grain is past, he recommended that each farmer should sow a good quantity of mily, or Turkey wheat brought [to South Africa] from Guinea by the Hassalt”. Its introduction into South Africa may have taken place before the establishment of the Dutch Colony in 1650. Portuguese vessels calling at the Cape for water on the voyage to and from their East African and East Indian Settlements, probably left maize-grain with the Cape Colonists (or even earlier, with the natives) in trade for water, meat, and other commodities. This is suggested by the Afrikaans word for maize, ‘‘mielie” being undoubtedly a corruption of the Portu- cuese word w/o, meaning grain. We know how easily new plants of economic value spread among the native tribes of Africa, as witness tobacco, peanuts, rice, jatropha, colocasia, etc. Once the culture of maize was established at several points on the African continent, eg., the Mediterranean shore, the Guinea coast, Cape Town, and Mozambique, the native population would soon distribute it throughout the Continent. 14. [ntroduction info Asia.—To the Portuguese voyagers, also, is probably due the early and rapid introduction of maize into India, China, Cochin, and other parts of the far East Indies, soon after the establishment of their East Indian settlements by Vasco de Gama at the beginning of the six- teenth century. Mendoza mentions (1585) among the plants observed by him in China, “the plant called maiz, which constitutes the principal food of the Indians in Mexico” (Watt, 1). Another route of introduction into Asia appears to have been by way of Turkey (and possibly also south Russia), Arabia, or Persia. The exotic character of the plant IMPORTANCE AND HISTORY 15 is indicated by the absence of characteristic Asiatic names, the names now in use often combining an indication of the route of introduction with the vernacular for the particular corn of the country. A recent bibliographer (Lacy, 1) considers that though it is no longer a disputed question that maize is of American origin, ‘the possibility of its having been known in the East before the discovery of America by Columbus is by no means closed”. She revives the alleged mention of rows (a name treated as synonymous with é/é de Turquie or maize) by one of two Persian historians of the fifteenth century, of which Bonafous, in his monumental work on maize, admits that the translation “if it is exact, would leave no doubt that maize was known in the Old World before the discovery of the New”. Bonafous finally dismissed the reference, having failed to trace it back, but Miss Lacy points out that this may have been due to an incorrect citation to Mirkond (1433-98) in- stead of to Khondemir (1475-1534), grandson of Mirkond, who wrote “at almost the same time as Mirkond ... and whose best known work, the ‘Khelassé-al-Akhbar,’ is very nearly identical in subject-matter with Mirkond's //¢story of the IVorld, the ‘ Rauzet-al-safa’”. Quoting Mirkond, or perhaps Khondemir, Herbelot, to whom Bonafous refers as “ Le célebre orientaliste d’ Herbelot,’ states that Rous, from whom Russia has taken its name, the eighth son of Japhet, son of Noah, sowed in all the islands of the River Volga, which empties into the Caspian Sea, “/e bled which we call de Turguze, and which the Turks still call to-day, in their language, by the name of rows and boulgar”’. [In this connection, however, it is instructive to note that the modern Persian names for maize are ghendum, gandunti-mak- kah (i.e. Mecca corn), and faldah.' The mere fact (if it should be proven a fact) that the name d/¢ de Turquie was used in one place for rows (whatever that may have been) and in an- other for sazze is no indication that the two were one and the same thing; for example, the word corm means wheat in England, maize in America, and the grain most commonly 1The word haldah, according to Meninski (Lex. Arab., Pers., Turk. 1780), was a name for “ frumentum sarracenicum,” i.e. Sarracens’ corn, also pointing to its western origin. CHAP. CHAP. I. 16 MAIZE used for food in several other countries, e.g. Sorghum vulgare, kaffir corn or Egyptian corn; the “corn” which Joseph's brethren went down into Egypt to buy was certainly not Indian corn or maize. De Candolle (1) suggests that the rapidity of the recent distribution of maize in Europe, Asia, and Africa completes the proof, if further proof were needed, that had it previously existed in Asia or Africa, maize would have played an impor- tant part in agriculture for thousands of years, instead of being but a comparatively modern culture. As recently as 1832 maize was grown in India only as an ornamental plant in gardens, not asa regular farm crop for grain. _15. Meaning and History of the Botanical Name.— The botanical name of the maize plant is Zea Mays. The generic name Zea is derived from the Greek Zesa or Zea, a sort of grain mentioned by Homer (Odyssey, 4, 41, 604) as used for feeding horses. The Zeca of the Greeks was certainly not the maize plant, which was unknown to them; but when, in 1753, Linnaus was renaming all the then known plants in accordance with his new system of binominal nomenclature, he used many of the classical names of the ancients, often taking the risk of applying an old name toa new plant, where the former was appropriate or pleasing. Linnaeus might, however, have adopted the generic name ‘ Mays,” already published by Tournefort in 1719, and followed in 1729 by Micheli who spelt it Mayz. The specific name AZays was used by the earlier botanical writers, Matthiole (1570), Dodoens (1583), and Camerarius (1588) as being the name under which the plant was intro- duced from America. From their writings it was adopted by Linneus. According to Prescott (1), Hernandez (1) derives the name maize from the Haytian word ma-h7z ; this was the name used for it by the Haytians when Columbus visited the island in 1492. 16. The Name Matze-—Maize is an Arawak word, met with in many forms in South America and the West Indies, e.g. mahiz, marisi, marichi, mariky, mazy, maysi, etc. This name followed the introduction of the grain throughout Europe, and was adopted into many of the European languages, being variously spelled maiz, maize, mais, mays, mayz, or mayze. IMPORTANCE AND HISTORY 17 The word mazze, therefore, dating back to the introduction of the crop into the Old World, having been adopted into many languages, as detailed below, and forming part of the botanical name of the plant, has the highest claim to recogni- tion as its universal vernacular name. The following are the different forms in which it is in use in different parts of the world :— Maiz (Portugal, Spain, Italy, Germany, Peru, Brazil, Uru- guay, and Argentina); mais (Germany and Denmark); mais (France); mays (Holland and Belgium); masé (Northern Africa), Maheende (Central Africa), Mahindi (Northern Africa), and Mihindi (Suahili), ie. Indian corn or maize of India ; mahiz (Hayti); marichi (Guiana) ; maysi (Cuba, Jamaica, and the Bahamas) ; maize (British Empire and the United States). In English literature the word appears in the following forms: maith (doubtless the phonetic spelling of the Spanish pronunciation of szahzs), maix, maise, Maiz, maize, maizium, mays, mayis, maijs, mayz, mayze, maes, maez, maze, mass. The ¢ isa comparatively modern suffix, which is not found in some English writers of the sixteenth and early seventeenth centuries, soon after the introduction of the grain into Europe. Du Bartas (1544-90), in his Divine Weekes and Iorkes (Sylvester’s translation), writes :— Heer, of one grain of maiz, a reed doth spring That thrice a year five hundred grains doth bring. Hakluyt (Voy., 1600) and Dampier (lVoy., 1676) spell the word azz, and Bacon, also, apparently throughout his writings, for we find it so in his Matural History, § 49; Sylva (1626); Med. Rem. Wks. (1626), and the earlier edi- tions of Essay 33 “of Plantations,” though in some modern editions the e has been added (perhaps in a pedantic effort to correct a “typographical error”). Bazley’s Dictionary (17th ed., 1757) gives only the spelling mazse, as does Murray’s ; the Century Dictionary has — “ maize, formerly also maiz, mais, mayz, mays”; while the Stavdard Dictionary gives both maiz and maize, the former on the authority of the Philological Society. Although mazze is practically the only form in which the word occurs in modern English literature, there are several 2 CHAP. CHAP. I. 18 MAIZE good reasons for dropping the e and adopting the spelling MALS -— (1). Simplified spelling is a demand of the age, which should be complied with where there is good reason and authority. (2). The e is unnecessary from the phonetic standpoint. (3). The etymology of the word does not appear to provide for it. (4). The form mays may, perhaps, have better claims to adoption from the etymological point of view, and is part of the botanical name of the plant, but is not in such general use in continental languages, and would be less easily adopted. (5). Mazz is the form in use in Germany, Spain, Portugal, Italy, Peru, Brazil, Uruguay, and Argentina, as already noted. (6). The authorities already cited seem ample. It remains for the literary and scientific public to decide whether the suggested change is sound, and whether it should be generally adopted. 17. Lhe Word Corn.—Corn as used in American literature designates maize. The Saxon word “corn,” Teutonic orn (whence Afrikaans “ koren”), is the general term for any cereal, and is applied in any country to that cereal most extensively used there for human food; in England the words “corn” and the “corn trade” as generally used, refer to wheat, while in America they usually mean maize. Recent American writers on cereals are endeavouring to introduce the word “ maize” as a substitute for “corn,” as the South African Department of Agriculture is endeavouring to use it in place of ‘‘ mielie ”. The term corn or grain has been applied to maize in the following countries: United States and Canada (corn, Indian corn); Great Britain (Indian corn); Sweden (korn); Holland (Turks koren); Belgium (Turkisch koorn); Germany (Turk- ischer kérn); Greece (Arabosite, i.e. Arabian corn); Formosa (fanmeh, i.e. foreign corn); Russia (Tureskorichljeb, i.e, Turkish corn); Japan (nan bamthbi, i.e. foreign corn); Italy (grano Turco, grano d’India, frumentum sarracenicum, grano Siciliano) ; Persia (gandumi-makkah, i.e. Mecca corn); Shan- gaan of N. Transvaal (mabele, i.e. kaffir corn). 18. The Word Mielie.—Mielie (often misspelled mealie) is the usual South African name, but is not known in IMPORTANCE AND AISTORYV 19 any other part of the world where maize is grown.’ It is derived from the Portuguese word mz/ho, from Latin I/?/éum, the name for millet, a grain at one time much used for food by the inhabitants of Southern Europe. In Portuguese-speaking countries, milho is the term for any cereal used for human food. The name mielie was doubtless an early Africander- Dutch corruption of the word milho, as used by the Portuguese sailors who first left maize at the Cape on their way to the East Indies ; it is significant that in Angola maize should have been known under the name of 6&/¢ portugais. We may, therefore, look upon the word mze/ze as a colloquial form of the word milho, meaning any kind of grain used for food, rather than the name of the particular grain to which it has been applied locally and in modern times. It would be better to use the now universal word maiz or maize as the connection with the English word mea/ is remote. The forms in which the word milho are applied to maize are: milho grande (Portugal, Brazil); milho d’India (Portugal); milho da India, milho de Guine (Brazil); mielie and mealie (South Africa). Moodie (Records, p. 137, 1841) spells the word mily, and Damberger (/vravels, p. 71, 1801) as melzs. 19. Other Vernacular Names.—In other countries where millet was a staple cereal before the introduction of maize, the local word for millet was often made use of in naming the new cereal, thus: gros millet des Indes (France); durah-shami (Arabic), dourah de Syrie (Egypt), dourah being the Arabic name of millet; bari-joar (Panjab), bari-jowar (Oudh), Mak- ka-jari (India), joar or juari being the general name in India for the great millet (Sorghum vulgare) and Makkai = Mecca (Watt), i.e. Mecca-joar or Mecca millet, indicating the route of introduction; jade sorgho (China); yuh-kau-liang (China), kao-liang being a Chinese name for Sorghum vulgare ; mashela bahry, i.e. millet from the sea (Abyssinia). On the other hand, where wheat was used more extensively, the local word for wheat was adopted in coining a name for the new cereal, thus we find: blé de Turquie or blé Turquet, blé de I’Inde, blé des Incas, blé de Guine, blé d'Afrique, blé d’Astrakan (France); blé de Rome (Vosges), blé de Barbarie 1 The Dutch word for millet is gierst, for maize it is mays and mais, and for Indian meal or corn meal, mais meel. % 2 CHAP. CHAP. I. 20 MAIZE (Provence) ; blé d’Espagne (Pyrenees) ; blé portugais (Angola) ; Turkischer weizen (Germany); Turkische waitte (Groningen) ; Turkse tarwe (Holland); misr-bogdag, i.e. Egyptian wheat (Turkey); trigo de Indias, trigo de Turkina (Spain); Turkish hvede (Sweden). Several other names, included in the following geographical list, are in use in various parts of the world :— Abyssinia; mashela bahry. Africa (North): masé, mahindi. Africa (Tropical) : bekkolo (Galla), maheende, gafuli nosri, simsin (Darfour), kasoli (Uganda); matawe (Chikaundi of Kasemba Dist., N.W. Rhodesia) ; tjibakwe (Mashuna) ; umum- bu (Matabele); mafluera (Kimwani, a coast dialect of Suahili; mafluera applies to both plant and grain) ; in Suahili language the fruit and plant are known as mwhzndi, the grain alone as mahindi (dim. vehind?, i.e. hind? = maize, singular la-, pl. = ma-—), and the ear as guzez ; in the Makau language it is called nakuo, in all stages of growth, and in Nyasaland chimanga ; in Angola it is known as mazza manputa or blé portugais, Africa (South) : maize (English); mielie (Afrikaans), angli- cized into mealie ; poone (Basutoland); lefeela, plural sefeela (Transvaal Basutos) ; sepeéla (Mapochs); shifake (Shangaan) ; shibakwe (Mashona) ; ma’ghea (phonetically ma’hea), plural, or le’ghea, one (Sapidi of Sekkukuniland) ; ‘’m-umbu (Mata- bele) ; semaka and monidi, plural mabidi (Bechuana): 'm-lungu (Swazi; in Zulu ’m-lungu means “white man”; its adoption by the Swazies may perhaps signify ‘“white-man’s corn”. The Swazies now also use the word ’m-lungu for white man, in place of their own word ’m-lumbi); 'm-bila or ’m-beela (Zulu; this word is also used by the Swazies, in deference to the practice of the white man) ; sinjembani (Zulu for the dark red grains of the type of maize grown by natives in the early days). The growing maize plant is called by the Transvaal Basutos le’tlaaka, while the Shangaans, according to Rev. E. Creux, call it mabele, which is the recognized name for kaffir corn among most South African tribes; mihindi (Suahili). In the Transkei the following names are used, according to Archdeacon Woodrooffe, for many years Anglican missionary to the Transkeian natives (J7a//y, 5): um-bona, the common Xosa term for both the maize plant and the “ fruit” ; utiya is IMPORTANCE AND HISTORY oa used by some of the Kaffirs and is synonymous ; izikweba- sombona = green mielies, i.e. izikweba = ears, and sombona = of maize; um-pa = the cob after the corn has been stripped ; in-tshatshoba = the flower of the maize plant; isi-gezenge = maize bread made from green maize ; um-kupa = maize- bread made from dry maize. Arabia: durah-shami. Argentina: maiz, pinsingallo (a pod-maize). Australia: maize, Indian corn. Austria: \kukurutz. Bahamas : maysi. Belgium: mays, Turkisch koorn. Bohemia: \ukurice. Brasil: maiz, milho de Guine, milho grande, milho da India, zabemo, avati or abati. Special breeds have particular names, e.g. milho dourado and milho catete are flints, milho pipoca and milho perola are pop-corns. Burmah : pyoung-boo. Canada; corn, maize, Indian corn. Ceylon: muwa. Chile: maiz (Spanish); cua (Indian). China; jade sorgho, yuh-kau-liang ;_ yii-shu-shu. Costa Rica: kup (Boruca) ; ep, ip (Terrabo) ; ain (Guatuso). Cuba: maysi. Denmark : mais. Egypt: dourah shammy, dura shami or dourah chamy, dourah de Syrie. Ethiopia: manigette (a pod-maize). fyi: sila-ni-papalegi. Formosa: fanmeh. France: mais, blé de Turquie, blé de I’Inde, blé de Guinée, blé de Rome (Vosges); blé de Barbarie (Provence); blé d’Espagne (Pyrenees); blé des Incas; blé d'Afrique; ble Turquet; blé d’Astrakan; gros millet des Indes; mais quarantain, mais nain and mais a poulet are pop-corns; mais d’été is an early flint. Germany : mais, maiz, Turkischer korn, Turkischer weizen, gemeiner mais. Great Britain: maize, Indian corn; Turkey corn (obsolete). Greece: arabosite. CHAP. CHAP. I. 22 MAIZE Guatemala; aima (Xinca Indian). Guiana: marichi. Flaytt: mahiz. Holland: mays, Turks koren, Turksche tarwe, Turkse tarwe; Turkische waitte (Groningen). Hungary: kukoricza (Magyar). India: the numerous Indian dialects produce a number of separate names for the varicus crops grown. The name for maize in most general use—in one or other of its forms—ap- pears to be makai (Hindustani; Bihar); makkai (Panjab) ; makka! (Tamil); makka-janar (Bengal) ; makka-sholam (Ma- dras); mukka-bhuta (Hindustani) ; elsewhere makkajari, mok- ka jonna, maki pyaungbt; an alternative name in Bihar is makatya. Where the joar, Sorghum vulgare, has been the staple cereal, its local name has been applied to maize, often with a qualifying adjective, thus: makka-janar (Bengal); bari-jowar (Oudh) ; bari-joar (Panjab) ; janara (Hindustani); janera® (in the west); junora (Patna); and makkajari, mokka jonna, junri and junala elsewhere. Other names in use in India are: kukri (Panjab) ; goom- dhan* (Assam) ; zonalu (Telugu) ; cholam (Madras) ; djagoung, mungari, and chhale. Sir George Grierson (1) gives the following words applied in India to different, parts of the maize plant :— Stalks: dhattha to the west ; thathera to the north gener- ally; thathero (south Bhagalpur); dant or danti elsewhere. The stalks are used for fodder under the name makai ke dant (Gaya). The Broken Stalk: lather (the north-west and in west Tirhut); nighas or nighesa (east Tirhut); no special name for this has been noted south of the Ganges. !'The Madras Manual of Administration, Vol. III, s.v. cholam, derives makka from mecca, saying that mecca means the west generally. According to Sir George Grierson makka is generally said to be derived from Skr. markataka, but the derivation from Mecca may or may not be true. 2 Sir George Grierson gives a possible derivation of janér as Skr. yavanala, and says that Platts derives junhar, another name for joar, from Skr. jivana-dhara. ‘Dhan (Skr. dhanya) is the Assam word for rice. Mr. J. D. Anderson of Cambridge says that in Assam, where rice is a staple crop, the word dhan is used in the sense of corn, and that goom means hidden or secret, and so “strange” or “foreign,” goomdhan meaning fureign corn. IMPORTANCE AND ATSTORY 23 x, Tassel: dhanbal or dhanahra. Silk: bhtia (to the west); ghua (south-west Shahabad) ; san (Champaran and Gaya); monchh (Patna); moccha (South Munger); moch or mocha (Tirhut and South Bhagalpur) ; it is also called kesi. Young Ear (when the grains begin to form but are not yet fit for eating): sancha. Unripe Ear: duddha (to the west generally); dodha (Shahabad) ; khichcha or aju (Tirhut); dudhghottu (Gaya) ; duddha makai (Saran and Patna); dudhbhoro (South Bhagal- pur); dant kamra (South Munger). Ripe Ear and fit for eating: bhutta ! or bal. Roasted Ear: horha (generally); orha (to the east). Dead-ripe Ears (grain hard and unfit for eating): pakthail. Blind Ear (ie. with no grain on it): bhorah or bhorha (north of the Ganges). Ear with few Grains : pachgotiya. Cob (after the grains are shelled off): lenrha (generally) ; lenruri (Shahabad); nerha (east Tirhut); baluri (an optional name in Patna and south-east Tirhut); balri (an optional name in south-east Tirhut and south Munger); khukhuri? or khonkhri (south-west Shahabad) ; haddi (south Bhagalpur). Shelled Grain: gota or got. Husk: balkhoiya or bokla (generally); khotya (to the west); khoincha (east Tirhut); pataura (South Munger) ; pocho (South Bhagalpur); in Champaran another name is kalchoiya, and in south-west Tirhut balko or kosa. Italy: maice, maiz; grano Turco or Granturco; grano d'India, grano Siciliano ; melliga or melgone (Lombardy). Jamaica: maysi. Japan: nan bamthbi, sjo-kuso, too-kibbi, tomoro-koski, or tomorokoshi. 1Sir George Watt (1) says that the word bhutta (Bengal and Bihar; Hin- dustani but, mukka-bhuta, elsewhere maki pyaungbu) may possibly be derived from bhukta or butta, to eat. Sir George Grierson says that it may be derived from Skr. bhrsta, roasted, which is suggestive in view of the method of cooking the young ears. 2The word kukri is used in the Panjab for some form of maize; and in south-west Shahabad khukhuri or khonkhri is the word for the maize cob after the grains have been shelled off. This suggests a connection with the Slav word kukuru (Turkey); kukurusa (Roumania); kukoricza (Magyar); kukurutz (Austria); kukurice (Bohemia). CHAP. I. 24 MAIZE Malaysia: djagoeng, jarung, cholam. Aaya: ixim. Mexico: mais, maize (Spanish); cintli=ear, olote = grain (Aztec). Paraguay : bisingallo (Guarany Indians). Persia: ghendum ; gandumi-makkah ; haldah. Peru: maiz (Spanish); sara or zara (Quichua Indian). Portugal ; maiz, milho da Indias, milho grande. Roumania : kuku-rusa. Russta: Tureskorichljeb. Siam: hacpot. Spain: el maiz, trigo de Turkina, trigo de Indias, zaras. Sweden - Turkish hvede, korn. Turkey » misr-bogdag, kukuru. United States: corn, Indian corn, maize. Uruguay : maiz. According to De Candolle (1), there is neither Sanskrit nor Hebrew name for maize. There is no Greek or Latin name because the plant was unknown to the Greeks and Romans. CHAPTER II. CLIMATIC REQUIREMENTS. The term climate, in its broadest sense, implies all the changes in the atmosphere which sensibly affect one’s physical condition.—HumBo.tpr. 20. Climate-—The profits from the cultivation of the soil, and in relation to these the preference given to certain crops, depend perhaps more on the extent of the market for such products than on the quality of the soil and the climatic condition within the geographical area of agricultural pro- duction (Davis, 2). Hann and Ward (1) define c/7maze as meaning the sum total of the meteorological phenomena that characterize the average condition of the atmosphere at any one place on the earth’s surface. That which we call weather is only one phase in the succession of phenomena whose complete cycle, recurring with greater or less uniformity every year, constitutes the climate of any locality. 21. Factors which Limit Distribution—The known facts of distribution of maize as a crop lead us to inquire what factors limit that distribution. These have an important bearing on the question of the world’s future supply, and more particularly on the problem of South Africa’s future share in the world’s trade. ; Maize seems peculiarly sensitive to climatic variations, and these furnish the principal limiting factor of distribution of the crop. The variations referred to include temperature, sunshine, amount and incidence of rainfall, and length of growing season. Geographic features and the character of the soil are also important. Only when these several factors are suitably combined does the culture of maize become commer- cially successful; the absence of any one of them may limit production on a large scale. CHAP. If; CHAP. Il. 26 MAIZE It is important that we should understand the relation of these several factors to the maize crop. Speaking broadly, the most favourable conditions are long humid summers, hot days and warm nights, comparatively heavy, intermittent rains, with abundance of clear, sunshiny weather between. Hot, arid climates, hot and continuously humid and shady regions, and arid cool-temperate climates with short summer season, are, generally speaking, unfavourable to the commercial pro- duction of maize. Open plains or plateaus are therefore more suitable than forest country. 22. Altitude—Altitude affects the growth of crops indi- rectly as it influences length of season, temperature, precipita- tion of moisture, depth, and richness of soil, etc. Evidence collected by Harshberger (1) suggests that the maize plant came originally from tropical table-lands at a considerable altitude, probably above 4,500 feet. Increase in elevation is accompanied by decrease in tem- perature and a steady shortening of the summer season, until, at very high altitudes, alpine conditions prevail; long before this point has been reached, however, it has ceased to be possible to produce maize. Davis (1) finds that in Argentina the decrease in temperature due to altitude is not a constant factor but varies according to the season of the year and the dryness of the air. The shortening of the season with in- creasing altitude has an immediate effect upon the crop in that the earliest autumn frosts, not falling regularly at the same date, are apt to kill the plants before the grain is ripe for harvest. The range of altitude within which a maize crop can be successfully grown largely depends on latitude; the nearer the Equator the higher the altitude, within certain limits, and the farther from the Equator the lower must be the altitude. Humboldt records vast maize fields on the Mexican plateau (between the 15th and 30th parallels) at 8,680 feet. Near Lake Titicaca, Peru, at about the 16th degree of S. latitude, maize is grown successfully at 10,000 feet. In the Indian Panjab, between the 30th and 35th parallels, it is more extensively grown in the hill country at 7,000 feet and over, than in the valleys, where it is largely replaced by rice. In Baluchistan it is grown as a regular crop at 5,000 to 9,000 CLIMATIC REQUIREMENTS 27 feet altitude. In the United States the major part of the crop CHAP. is produced between the 35th and 45th parallels, and 82 per cent of it at an altitude between 500 and 1,500 feet; the proportion grown above 1,500 feet is only 4:4 per cent. Harshberger (1) points out, however, that this is partly due to the absence of large areas of tillable land at an elevation of 2,000 feet, for very fine maize crops are raised in North Carolina at 4,000 feet, between the 34th and 37th parallels. 23. Temperature —Careful investigations carried out in the United States fail to show any direct relation between actual temperature and yield of the maize crop. Maize is a tropical plant, susceptible to frost in all stages of its growth; but being an annual, it can be grown as a summer grain crop in warm-temperate climates, and as a fodder crop (not for grain) even in cool-temperate areas such as the south of England. Dwarf, early-maturing sorts have been known to ripen seed in the south of England and even in Norway as far north as 63° 13’ (Wueller, 1). Martyn (1) states! that maize was cultivated in England in 1562, but that the seeds ‘‘rarely ripen in England... . Mr. Miller thinks that maize might be cultivated in England to advantage. But it can scarcely be expected to be grown here for the grain, except in favourable seasons and warm soils and situations, Yet as a fodder it might be of considerable service, if it were cut when just opening into ear, and given fresh to the cattle every day” (Martyn, \.c.). Of var. y Zea vulgaris, Mill. (Déct., n. 3), he says, “This ripens its grain perfectly well in England in as little time as Barley”. But ‘Maize is seldom cultivated in England for use” (Martyn, |.c.). Eighty-eight per cent of the American crop of 1897 was grown between July isotherms 70° and 80° Fahr. (Srewer, 1). The Argentine crop is grown with a mean January tempera- ture of 75°78”, while that of the Transvaal Maize-belt is under 70° Fahr. The actual highest yields of the United States have been obtained between July isotherms 75° and 80° Fahr. (Harshberger, 1). The average temperature of the maize-belt of Argentina is given as :— 1 On the authority of Turner’s Herbal, Part U1, fol. 58 n. CHAP. II. 28 MAIZE 1856-1875 : ‘ : ‘ , : 62°9° 1876-1896 : : : : : : 61'5° 1897-1900. é : ; ; . 63°1? The maximum temperature is said seldom to exceed 95° Fahr., though it seems much higher owing to the excessive humidity. The maize zone lies between summer isotherms 71°6° and 788° Fahr., and annual isotherms 59° Fahr. and 68° Fahr. Tas_e I, COMPARATIVE TABLE OF MEAN MONTHLY TEMPERATURES. | Febru- Mean, 4 | Place | Feet Years. ber. | ber. January ary. Months. careers |—— Fee Fare |e aa | |— ae eee | Transvaal— | | Vereeniging . | 4700 1903-10 67°97 | 69°63 | 70°34 | 69°24 | 69°29 Bethal . y 5580 1903-10 | 62°83 64°75 | 65°96 | 65°18 | 64°68 Pretoria | | | | | (Arcadia) . | 4500 1903-10 | 69°18 | 70°78 | 71°54 | 69°64 | 70°28 Natal— | | | | | Cedara 4 | sees } 68°2 | 69°6 | 72°0 70°75 | 7O'l4 Rhodesia— | | | Bulawayo aH 4470 | 1898-1902 71'8 qr | 7r0 | 689 | 70°70 Bulawayo. | 1908-9 7o6 | 724 | 706 | 684 | 70°50 Salisbury .| 4810 | 1898-1902 | 69°6 | 68:2 | 69'4 | 67°6 | 68-70 United States of | | | America—! | | Georgia. 13 years 72°t | 73°6 79'6 | 7S | aro2 lowa « =| 18 years | 59'8 69°3, | 73°6 | 71°6 68°3 Argentina— | | | | | Mean of 13] | stations. | — 69°12 | 73°54 | 75°78 | 74°59 | 73°2 Bahia Blanca | | aes } 65°37 | 7o°6r | 73°81 | 73°89 70°42 Goya . P | 1876-1g00 | 2°85 | 77°38 | 78:22 | 7745; | 7O%47 Paraguay— | | | Mean of 2 sta- | | | | } tions . : | 1892-1900 | 76°55 | 80°53 | 80°60 | 79°99 | 79°42 S. Europe—* | | Vienna . | 636 | 100 years 59°18 | 65°84 | 68:90 | 67°46 | 65°34 ' The heaviest maize yields of the United States have been grown between July (= January) isotherms 75° and 80° Fahr. ? Various periods from 1855 to 1goo. “May to August inclusive. In the Transvaal it is generally considered that the Lower Bush-veld (below 2,000 feet), though hotter, cannot compete with the High-veld in the production of maize; if this is actually the case it is probably due largely to the character of CLIMATIC REQUIREMENTS 29 the soil, as there is a narrow strip of poor soil running through CHAP. the Lower Bush-veld from north to south, and good maize | crops are raised on either side of this belt at the same altitude. It may also be due in part to deficiency or irregularity of rain- fall, and greater evaporation. Taste II. MEAN TEMPERATURES OF THE CEREAL BELT OF ARGENTINA. November, December, Fanuary, and February. | | November, | December. | January. February. | Station. | Period. ———| ae ) Sie | G& | Fab] G | Fate, | coe | Fabr.| C. | Fahr. | Sener | Peewee geen a EE ae Buenos Aires. | 1856-1900 | 19°90 | 67°82 | 22°43 | 72°37 | 23°66 | 74°59 | 23°06 | 73°51 4 (| 1860-83 19, Red ‘ x ees ‘B81 | 20° aed | Bahia Blanca | 1897-1900 | 5 18°54 | 65°37 | 21°45 | 70°61 | 23°23 | 73°81 | 16 | 71°39 Tandil . . | 1876-82 17°37 | 63°26 | 19°30 | 66°74 | 21°20 | 70°16 | 2I°II | 70°00 Viedma . .| 1876-82 17°95 | 64°32 | 20°13 | 69°85 | 23°36 | 74°05 | 21°13 | 70°04 Rosario . ‘ 1891-1900 | 20°78 | 69°40 | 23°73 | 74°71 | 24°70 76°46 | 24°33 | 75°79 Céres. . | 1896-1900 | 23°15 | 73°67 | 25°68 | 78:22 26°61 | 79°91 | 26°83 | 80°30 5 (| 1896-9 \, ‘ 24°08 | 76°06 | 24°8 6: Parana. .{ rBrgeBa [SACO | 7844 | 24-20 | 75°56) 24°98 | 70%9 ae 5 | 76°73 Concepcion del | | ae Uruguay .| 1894-9 | 2rro | 69°98 | 23°94 | 75°09 | 24°44 | 75°00 | 24°28 17 Hernandarias.| 1877-92 22°82 | 73°08 | 25°11 | 77°20 26°23 79°23 | 25°33 77°59 Goya. . | 1876-1900 | 22°69 | 72°85 25°21 | ange 25°68 | 78°22; 25°25 77°45 Cordoba . . | 1873-1900 | 20°87 | 69°56 | 22°97 | 73°34 | 23°44 | 74°19 2274 72°93 Rio Cuarto . | 1881-1900 | 20°98 | 69°76 | 23°36 | 74°05 | 23°96 | 75°22 | 23°07 73°52 San Luis =| 2874-9 | 19°96 22°25 | 72°05) 24°70 | 76°46 | 23°39 | 74°10 268°02 | 299°76 316°19 307°53 Mean of 1 | ; ; stations | 20°62 | 69°12 | 23°08 | 73°54 | 24°32 | 75°78 | 23°66 | 74°59 Paraguay. Asuncion | 1892-1900 | 24°78 | 27°09 | 80°80 | 27°20 | 80°96 | 26°95 | 80°51 Itacurubi_ del | ; : ae ee : Rosario ‘ | 1892-9 | 24°72 | 26°83 | 80°30 | 26°80 | 80°25 | 26°38 | 79°50 | | | | | 49°50 | : 54°00 53°33 | 49°50 | 53°92 54 BS 3" y | | 24°75 | 76°55 | 26°96 | 80°53 | 27°00 | 80°6 | 26°66 | 79°99 In the warmer coast-region of Natal, though there ts less maize grown, the yields per acre are heavier than on the es lands; but this is probably due to increased fertility os the soil, and to longer growing season, rather than to actual in- crease in temperature. CHAP. Il. 30 MAIZE Tasve III. MEAN MONTHLY TEMPERATURES, TRANSVAAL. Bethal (5,580 feet). SS ————— | Year 1903-4. | 1904-5. | 1905-6. | 1906-7. 1907-8. | 1908-9. | 1909-10. Average. —-— Eker | ee Nov. = 63°9 | 63°93 | 616 | 623 | 63:3 | 62°6 62°83 Dec. | — 624 | 66:4 | 64°2 64°7 | 65°4 | 654 | 64°75 | Jan. — | 660 | 69°8 | 66:1 | 67:0 | 62°9 | 64°0 | 65°96 | Feb. | — | 64°5 | 64°6 | 658 | 66-4 | 648 | 650 | 65718 | | ie | 258°72 | boot i ae 84°08 | : Vereeniging (4,700 teet). Nov. 68:2 69°7 68:9 | 66°8 | 664 | 67°6 | 68-2 67°97 Dec. | 73°4 65'9 va aes 638°6 63°38 70"4 69°2 69°63 | Jan. | 716 | yor | 74'1 | 69°8 Joo | 68"4 63-4 | 70°34 | Feb. 705 69°4 68°6 | 6g'0 74 67°6 68-2 69°24 | | | | 277718 | 69°29 | Pretoria (Arcadia) (4,500 feet). Nov. | 686 | 711 | 69°6 | 67°1 68°5 69°8 | 69°6 | 69°18 Dec. | 707 | 69'0 929 | 69°6 | 69°9 | 73°2 | 70°4 70°78 Jan. 714 722 | 74°38 | 7V0 | 722 | 706 ~~ 70°0 71°54 Feb. | 69°3 707 | 706 | 709 | 75°9 | 69°0 | 70'L 69°64 | | | | | 28114 | | | | | | | | 70°28 The above tables show the mean monthly temperatures of the four growing months for maize, November to February inclusive. In the case of the Transvaal stations the means are for the seven years 1903 to 1910 inclusive (/zzes, 1), and are obtained by halving the sum of the mean daily maximum and the mean daily minimum. The Rhodesian figures are those given by Mr. Hutchins (1), and the Georgia figures are the means for thirteen years as furnished by Director Redding (1); the latter are for the corresponding summer months of May to August inclusive. The lowa figures are from Bowman and Crossley (1). 24. Night Temperature—Some writers (Darwin, 2; Harsh- berger, 1) conclude that the maize crop does not flourish where the nights are ‘cool,’ no matter how favourable the other CLIMATIC REQUIREMENTS 31 33 conditions. The term “cool” is relative, and may be mis- leading, for in those parts of the Transvaal where the maize crop thrives the summer nights are invariably cool. The director of the Union Observatory gives the following figures showing the mean daily temperature at 6 a.m. (the coolest hour of the night) at the Government Observatory, Johannes- burg. TaBLe IV.! MEAN MINIMUM TEMPERATURES AT GOVERNMENT OBSERVATORY, JOHANNESBURG. September, 1904—March, 1909. | Mean Daily Minimum Temperature, 6 a.m. Mean, 5 | Month. |e z eee =~ =a | Seasons, | Sept., 1904- | 1904-5. 1905-6. 1906-7. 1907-8. 1908-9. Dec., 1908. | | as | ese | | | September ‘ 45°8 48°8 49°8 488 532 49°3 October . : 51°9 54500.) Sr | srr | 51°7 52°1 | November - | 57°4 54°55 | 5370 542 | 55°6 549 December. | 5472 57°5 | 55°72 | 55°9 57°8 56°r | | January . - | 56°8 59°8 gre | SG 57°7 57°6 | February . Fi 56°6 561 57°38 57°7 56°r 56°8 | | March | 53°5 535 | 566 | om oe 54°6 | | 7 Months’ mean 53°7 55°0 | 54°4 54°0 55°3 | 54°5 | Annual mean 50°52 514 | §0°5 50°7 516 | 50°9 | | | | At Buenos Aires the mean minimum summer temperature (December to February) at 5 a.m. is 63°5° Fahr. lAbstracted from Reports of Transvaal Department of Meteorology (Innes, 1). 2.N.B.—The mean daily minimum is 1°7° lower than the mean tempera- ture at 6 a.m. The following tables, taken from the publications of the late Transvaal Department of Meteorology, Johannesburg (Innes, 1) show the mean daily minimum temperature over a series of years at Vereeniging (4,700 feet altitude), and Bethal (5,580 feet altitude), two of the most important maize centres of the Transvaal. Comparing the two tables we find that for the 880 feet difference in altitude we have an average differ- ence in the annual mean of 1:°2°, and in the 7 months’ mean of 2°7°, CHAP. Om 32 MAIZE CHAP. TaBLe V. MEAN MINIMUM TEMPERATURES AT VEREENIGING. September, 1904—March, 1909. | Mean Daily Minimum Temperature. | | Mean,5 | | Month. ie as Fee | Seasons. 1904-5 1905-6 | 1906-7. | 1907-8. 1908-9. a a ee ir Se EE |e ceed | Se Meet eRe Seen | ent oe We etree ene | | | j; September © | 434 4553 | 44°2 43°6 46°7 44°6 | October . : . | 50°7 51:0 6} «(49°3 50°0 401 49°8 November : .| 55°6 55°8 | 53° 52°3 543 | 544 | December : .| 53°3 58-6 | 54°4 56°7 56°35 | 55°9 January . ‘ «|, 25782 61°0 59°0 551 599 | 584 February . : - | §8°3 56°7 58°5 59°5 583 | 582 | March. : =| 529 2°5 545 | 526 542 | 53°3 7 Months’ mean + | 53°0 54°4 | 53°4 | 52°8 540 | 535 _ Annual mean . el) 452 45°06 | 45°2 | 44°0 45°38 | 452 | | | TaBLeE VI. MEAN MINIMUM TEMPERATURES AT BETHAL. September, 1905—March, 1909. SE ee Mean Daily Minimum Temperature. | Ties One eae re ea Dn ee VLC ALL Month. | | Seasons. | 1904-5. 1905-6. 1906-7. 1907-8. 1908-9. | | | September , . ae 41 42°3 Aas 45°5 | 42°8 October ‘| ar? 47°0 40°4 46°6 46°9 | November | 5I4 50°3 50°2 51°'7 | 50°9 December | 54°58 530 53°6 540 | 538 | January 57°4 56°7 54°6 28 | 55°4 February . : | eee 53°2 55°7 53°58 556 | 54°6 March . ‘ sf oe 49°2 52°5 50°5 52°7 | 51°2 | 7 Months’ mean 50°7 51°r 50°2 51°3 | 50°8 | Annual mean 43°9 43°8 43°1 44°90 | 43°79 i 25. Frost.—Late winter frosts have little effect on the South African maize crop, as they usually come at a time when there has not been enough rain to start the crop; but when they fall as late as the middle of October they may cause some damage. Early winter frosts are more dangerous ;! 1On the Transvaal High-veld killing frosts are sometimes experienced as early as 28 March. CLIMATIC REQUIREMENTS 32 for on the high plateau of the Transvaal, at 5,000 to 6,000 feet elevation, the season is usually too short for late-maturing breeds of maize, and almost every year a proportion of the crop of medium-late sorts—such as Hickory King—is seriously injured. This is due to the sudden though temporary fall of temperature which frequently precedes the advent of the real winter by two or three weeks, when the frost is often suffi- ciently severe to injure the unripe grain. A remedy can be found in autumn tillage and earlier planting to bring the crop sufficiently forward to miss the frost; but too early planting often results in loss from cut-worms. When the South African maize crop is once ripe, frost does not injure it, and it can be left standing in the field to dry out, through the winter, without fear of injury. As there is considerable difference in the time required for the maturity of different breeds, the earlier-maturing sorts should be used at the higher altitudes ; some of these yield rather less than the longer-growing sorts, and farmers are reluctant to drop the latter, even though they entail greater risk. At lower altitudes, as along the coast of Natal, the Ubombo Range in Swaziland, and the adjacent portion of the Transvaal, the season between frosts is so long that two successive crops of maize-grain can be matured in the same year. Temperature appears to have no direct effect upon yield per acre, but it does influence the maturing of the grain, and often in this way affects the yield of marketable grain, especi- ally at higher altitudes and in the southernmost of the maize- growing districts of South Africa. In Argentina frost is apt to cause a considerable loss of crop. Where the crop has been thrown late, from one cause or another, the stalks may be cut just before the time when frosts are expected, and “stooked” in the field. This does not interfere with the proper filling out and ripening of the grain if the crop is not cut before the grain has begun to harden. Not only does this method enable farmers on the High-veld to save their crop from injury by frost, but it results in a saving of some 50 per cent of the feeding value of the “stover”. At the suggestion of the writer this method was tried by several High-veld farmers in the Transvaal, during the very backward season of 1909-10, with excellent results. 3 CHAP. Il. CHAP. Il. 34 MAIZE 26. Hatli—No country in the world has such a_ perfect climate that the farmer is entirely free from worry, what- ever his crop may be. On the whole the climate of South Africa is probably as nearly perfect as any; but it is not without drawbacks. Perhaps the chief of these is hail. The worst hailstorms usually fall in the months of November and December; on 16 November, 1909, hailstones weighing 44 oz. were reported from Germiston, and considerable damage was done to crops in the districts of Bethal, Er- melo, Standerton, Heidelberg, Marico, . Rustenburg, Wakker- stroom, and the Witwatersrand. At this time of year the maize plant is still comparatively small; so long as it is not in tassel, a new crop of leaves may be produced and, though somewhat retarded in develop- ment, the crop usually recovers. Hailstorms coming in Janu- ary are most likely to damage the maize crop. A storm of hail in the Glencoe District of Natal, on 21 January, 1908, cut the maize crop to the ground and injured it beyond recovery ; but it is said to have been the heaviest hailstorm known in that Fic, 5.—Effect of hail on the leaves distriet tbe forty ‘Aosae Most af the mabeplaat. damage is done when the main stem of the plant is broken by the hail, causing the development of sucker shoots, which do not bear good grain. More frequently the injury is restricted to stripping the foliage into ribbons, sometimes leaving only the midrib (see Figs. 5 and 6) and sheath of the leaf to function in photo- synthesis (169). Ifthe storm occurs before the tassel appears, the plant may be able to throw out additional leaves, by which photosynthesis can be carried on; but if it has reached the stage shown in the above figure, no fresh leaves can form on CLIMATIC REQUIREMENTS 38 35 the stem; there must then be some loss in weight of grain, CHAP inasmuch as all the starch must first be elaborated in thie leat Mt. before it can be deposited in the grain (1 69). It is a fortunate feature of the South African hailstorm that it is usually confined to a comparatively narrow strip, so that not all of any one farm is damaged, and as a rule he Fic, 6.—Effect of hail on the stem and leaves of the maize plant. same farm is rarely visited by hail two years in succession. There are, however, ‘“hail-belts,” in which hailstorms appear to recur almost yearly. Apparently there is no reliable preventive for hailstorms , but insurance policies against damage from hail may be effected. * 3 CHAP. Il. 36 MAIZE 27. Soil Temperature—The temperature of the soil has much to do with the successful growth of maize and other tropical crops. In cold soils germination and subsequent growth are retarded. Experiments conducted at Pretoria show that whereas maize planted in September and early October usually requires eight days to appear above ground, that planted at the end of December or early January will sometimes appear in three days; in each case the seed was well watered daily, so that lack of moisture was not the cause of retarded germination, which may therefore be attributed to low temperature of the soil. Maize lands should be well drained, for wet soils are usually cold soils. It is largely on this account that maize germinates badly, and its growth, also, is retarded in water-logged soils. When water stands for any time on the maize lands, the foliage becomes yellow and the plants remain stunted. In cycles of droughty weather there is a tendency to plant the crops in low-lying ground which retains a certain amount of moisture ; but when normal seasons return the crops in these lands suffer. It is better to apply the principles of dry-farming (i.e. good tillage) to the soil (see chap. IX.), in order to con- serve the moisture, than to use undrained land in anticipation of possible drought. 28. Motsture Regutrements.— Maize is, on the whole, a drought-resistant plant, but some breeds suffer more from drought than others, and these should not be chosen for regions where the average rainfall of the period from December to February inclusive is too light for the ordinary breeds. King (1) has found that in Wisconsin the maize plant abstracts from the surrounding soil 270 lbs. of water for each pound of dry matter grown, which is equal to a rainfall of 2-4 inches for each ton, or only about half the amount required (in Wisconsin) by oats and clover. But the maize plant requires a consider- able amount of water at certain stages of growth. In Illinois the growth of maize during one week in July has been found equal to 1,300 lbs. of dry matter per acre, which would require 1°5 inches of rainfall, according to King’s experiments. Ata time of such rapid growth the plant is apt to suffer from drought unless the soil is in the best physical condition (Afunt, 1). CLIMATIC REQUIREMENTS 39 An American writer points out that the curling of the leaves of the maize plant in July (equivalent to January in South Africa) is a bad omen to maize-growers in the drier districts. The time of the formation of the ear (January and February) is the critical period in the life of the plant, and lack of moisture at this time means curtailment of yield (Bowman and Crossley, 1). 29. Rainfall_—Though no direct relation exists between actual temperature and yield, rainfall, on the contrary, has a very direct bearing upon yield. At the Illinois Agricultural Experiment Station a rainfall of 13 inches during the five growing months, produced 1,792 lbs. (practically 9 muids) of dry maize-grain per acre. The following year, with 22°5 inches during the same period, the yield (without fertilizer) was 5,264 Ibs. (over 26 muids) per acre. The mean temperature was more favourable the first season than the second. The results indicate that the increase of 17 muids per acre was due to the additional 9°5 inches of rain, an average of 1°9 inches per month. In the Corn-belt of the United States the most favourable condition is found to be a series of comparatively heavy rains during the growing season, but at considerable intervals, with clear sunshiny weather between, and followed by a warm, dry, ripening period. Ifthe rainfall equals 11°5 to 12°0 inches in the three summer months corresponding to December, January, and February in the Southern Hemisphere, this should be adequate; of this, 4°5 to 5:0 inches should fall in January, when the ears are growing most rapidly. The aver- age rainfall for these three months for thirty-nine stations through the Maize-belt of South Africa is 11°92 inches, while the average for January is 4°35, or 4°6 if we omit the three driest localities, which are really outside the Maize-belt. Heavy rainfalls and cloudy weather during the planting season (corresponding to October and November in South Africa and South America) are in North America found to decrease the yield. In South Africa a wet October and November prevents proper weeding and encourages early growth of weeds which withdraw moisture and plant-food from the young maize plants, thereby reducing the yield. But excess of moisture, as already explained (1 27), is injurious. CHAP. Il. 35 MAIZE CHAP. The average monthly rainfall for the four growing months of the maize crop, as recorded during thirteen years at the Fic. 7.—Grains which have been fer- Fic. 8.—Grains cracked from ex- tilized but not fully filled with posure to unfavourable weather starch, probably owing to drought. conditions. Georgia (U.S.A.) State Agricultural Experiment Station, was as follows (Redding, 1) :— South African Georgia. Equivalent. Rainfall. May November 2°QI in. June December 4°37 July January 5°52 12°80 August February 6:06 CLIMATIC REQUIREMENTS 39 The rainfall of the Argentine Maize-belt ranges from 31°53 CHAP. inches to 39°4 inches, but u. this is divided with fair uniformity between summer and winter. March is the rainiest month, and April is about as wet as February, which is unfortunate for the drying of the grain; this is one of the greatest draw- backs to maize-growing in Argentina. The winter is wet and frosty, which makes it difficult to get the grain into merchantable condi- tion, and to store it satis- factorily; the consequent percentage of loss in Argen- tine cargoes during the ocean voyage is heavy. Even when the greatest care is exercised, the un- favourable climatic condi- tions are likely to handicap maize-growing in Argen- tina. The following table (VII), prepared with the courteous assistance of the director and staff of the Government Observatory, Johannesburg, shows that the summer rainfall con- ditions throughout a great part of South Africa are eminently suited to maize production. Certain areas must be excepted, however, such as the Cape Peninsula Fic. 9.—Grains cracked from exposure to and the adjacent areas unfavourable weather conditions. 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Tl. 42 MAIZE which have chiefly winter rains; the areas of very low rain- fall such as the Karroo; areas where there is a deficiency in the spring rains, or where the intervals between rains are too great, as in parts of the south-western Transvaal and western Orange Free State; and the higher mountain ranges where, though the rainfall is ample, the growing season is too short. 30. Sunshine—The maize plant is especially suited to the treeless grass-steppes of upland plateaux, and also thrives in efen “bush” country. But it does not seem at home in humid, shady, tropical forests. Sagot (1) shows that maize does not thrive in the warm, damp climates where manioc (Manthot spp.) is grown, and De Candolle (1) supplements this observation by pointing out that forests are generally un- favourable to the production of axy annual plants. The latter view seems in harmony with the general geo- graphical distribution of cereal crops in the tropics. Major Whitlock (1) observes that guinea corn (Sorghum vulgare var. ), an annual plant, is the staple cereal of the natives on the plains of Nigeria, at about 1,400 feet above sea-level, from Lake Chad almost to the foot of the watershed plateau between the Benue and the Cross Rivers. South of this plateau, however, where the country is clothed with forest, no more guinea corn is seen, the natives subsisting entirely on yams and plantains. In parts of Uganda, also, bananas are more extensively grown for food than any cereal. This is probably due, in the case of maize, to lack of sunshine. It is noticeable in South Africa that in cloudy seasons, like that of 1909-10, when there was nearly twice as much cloud as usual during the months of January and February, the maize crop is light. In continuous wet weather, pollination appears to be retarded; if the wet weather alternates with warm sunshine at short intervals, pollination can take place readily ; nature has provided that the silks shall be receptive for a considerable period (sometimes as much as fourteen days if pollen is not applied earlier), while the pollen supply may last for two to three weeks through a natural irregularity in time of flowering of different individuals ; in one plant, alone, pollen continues to fall for about four consecutive days. 31. Lnfluence of Climate upon Vegetative Characters and Time of Maturity.—Careful study of the influence of climate CLIMATIC REQUIREMENTS Fi ioes upon habit of growth is needed. The same breed appears to differ in size and in time of maturity at different altitudes and latitudes. Hunt (1) concludes that in the United States, as a general rule, a breed becomes one day later for each ten miles south or north of a given latitude, if the altitude is the same. This means that a variety which ripens two weeks before a killing frost in a given locality would only barely ripen if taken 140 miles away from the equator at the same altitude, the date of the first killing frost remaining the same. He advocates that, in introducing new seed it should prefer- ably be obtained from about the same latitude. Similarity of latitude may be a sufficient guide on a vast and nearly level plain like that of the Ohio valley; but where the topography varies as it does in South Africa, other factors than latitude and altitude influence the climate, and we doubt whether (with the present lack of knowledge of these controlling factors) South African farmers can make much practical use of the suggestion. A breed which matures in 90 days in Australia may take roo or 110 days in the Transvaal. It is very noticeable that the same breed takes a longer or shorter time to mature in different years, according to the “ season”; in a time of drought, growth is checked and flowering and fruiting are hastened, while in a rainy season, growth is continued much longer, Even within the Transvaal, and during the same season, the same breed, grown from the same lot of seed, is reported as having varied considerably in time of maturing in different districts. This is due partly, no doubt, to the con- dition of the soil as regards moisture at time of planting; in an air-dry soil such as is found over large areas during a great part of the spring, the seed does not germinate as quickly as ina moist soil. Allowance must also be made for the personal equation, different observers holding different views as to when the ear could be considered “ mature” or safe from frost ; lack of purity in strains of the same breed may also be a factor. 32. Acclimatization.—It is said that when a recognized breed has been grown for some time under diverse climatic conditions, it not only changes considerably in stature and “time of maturity, but that these habits become more or less fixed so that, when taken back to the old conditions, the plant does not at once respond to the change. In this way different CHAP. IT, CHAP. II. 44 MAIZE strains of the same breed are supposed to be developed ; they are said to become adapted to different conditions. The amount of such change, if it does occur, must be limited, however; within these limits it could be made use of in the acclimatization of new breeds, but it would not enable us to take a very late maturing sort suddenly from a tropical climate to a much higher altitude and colder latitude, and acclimatize it successfully ; this would have to be done gradually and by intermediate steps, and with some breeds might not be suc- cessful even then. Little is known, at present, of the actual effect of climate upon the maize crop. If the facts are such as have been indicated, farmers would do well to make use of them, or at least to keep them in mind when purchasing seed-maize. It appears clear that seed-maize from one climate takes some time to become acclimatized to another, and in the United States it has not been found desirable to take seed- maize from the rich alluvial plains of the Mississippi to the poorer soils of Virginia. For these reasons it is not desirable to buy bulk seed from hot, humid regions, at low altitudes, for cultivation in cooler and drier conditions at high altitudes. Nor is it desirable to obtain seed from deep, fertile soils for growth on thin, poor soils; one of the chief reasons that Hickory King has become such a favourite in South Africa is its ability to thrive on relatively poor soils and with rough treatment. But the converse may perhaps also be true, that it is not desirable to obtain bulk seed-maize from colder and drier climates and poorer soils for growth in hotter and more humid climates and on richer soils, because the quicker maturing habit will have been formed and the plant will not be able immediately to take advantage of the longer growing season and greater amount of plant food, and the resulting crop may be less than would have been the case with a breed already acclimatized to those conditions. 33. Influence of Climate upon Varieties.—The origin of the different varieties of maize is unknown, through lack of historical records, but it seems probable that there is some relation between climate and existing varieties. It is notice- able that those breeds grown in the most northerly parts of the United States and in Canada are mainly flints, while CLIMATIC REQUIREMENTS 45 in the Southern States dents are grown almost exclusively. Dents generally yield more heavily than flints, but take longer to mature; flints mature earlier, but as the yield is lighter they are grown only where others cannot mature owing to shortness of season. It is said that a northern flint variety, after having been grown for several years in Illinois, changed under the influence of climate into a dent; but before concluding that this change was caused by change of climate it should be known whether any precautions had been taken against cross-pollination, for if crossing had taken place between the flint and a dent, the heterozygous grains might retain the flint appearance and would later produce dent grains, which, if selfed, would breed true. Sugar maize is said to be rarely grown in the Southern States, but whether this is due to the climatic conditions being unfavourable for the production of sugar maize, or whether it may be due to the taste of the people, is not clear. 34. Influence of Climate upon Chemical Composition.—As the result of thirty-five analyses of dent maize grown in the Northern States, and forty-nine from the Southern, Hunt (1) concludes that there is no material difference in composition in maize grown in different parts of the country,. over a very wide range of soils and climates. Analyses of Transvaal samples made by the Division of Chemistry of the Union Department of Agriculture tend to confirm this view (see chap. XIII). CHAP. Il. CHAP. ITI. CHAPTER III. GEOGRAPHICAL DISTRIBUTION. To the wise man all the world’s a soil.— BEN Jonson. 35. Geographical Distribution.—In the preceding chapter we have seen that maize requires a hot, sunny climate; it thrives best between the 4oth parallels of latitude. Early ripening breeds are grown for grain in warm-temperate regions as far north as the 48th parallel in the northern hemisphere, and for fodder, still farther north, in the cool-temperate zone. The value of maize as a cereal crop for man and his domestic animals has led to its world-wide distribution in the brief space of time since the discovery of America. Although, as already pointed out (1 10), maize was probably first grown in New Granada as a cultural crop, and has only been known in Europe since the beginning of the sixteenth century, to-day three times as much is produced in Europe as on the whole of the South American continent. The countries mentioned in Table VIII are the leading producers of maize-grain; the figures given are for the year 1906, which was a record year for maize production in the northern hemisphere; they are taken from the Year Books of the United States Department of Agriculture (U.S.D.A. 7)! and other official publications, and are stated in United States standard bushels of 56 lbs. In the case of Mexico and the African continent, the figures are only approximate, through lack of precise data. Maize is also grown, but to a lesser extent, in the Province of Quebec (Canada), Central America, the West Indies, Brazil, Paraguay, Bolivia, Chile, Peru, Nyassaland, Uganda, British 1The U.S.D.A. Year Book omits entirely the production of British India, which is larger than that of Mexico and Canada combined. 46 TVasBLeE VIII. THE WORLD’S MAIZE CROP OF 1906. Bushels. Servia Bulgaria Spain Portugal France Asia— British India Africa— Egypt : . F Natal (and Zululand) Orange Free State Transvaal . Cape Colony Rhodesia . Algeria Australasia— New South Wales Queensland Victoria New Zealand Western Australia Sudan (Anglo- Egyptian) Bushels. Bushels. North America— United States 2,927,416,000 . Mexico 70,000,000 Canada (Ontario) 23,989,000 South America— Argentina . 194,912,000 Uruguay 3,226,000 Chile 846,000 Europe— Austria- Hungary— Austria . 18,177,000 Hungary 162,925,000 Croatia-Slavonia 20,470,000 Bosnia- Herzegovina 8,900,000 210,472,000 Roumania. 130,546,000 Italy . 93,007,000 Russian Empire— Russia proper, including Bessarabia and South- ern Russia . 59,320,000 Northern Caucasia 11,181,000 70,501,000 27,786,000 27,780,000 18,714,000 15,000,000 14,581,000 30,000,000 3,845,000! Not stated. Not stated. 3,200, 0007 Not stated. 544,000 300,000 5,714,000 2,233,000 661,000 653,000 1,000 3,021,405,000 195,954,000 3, 220,389,000 608,387,000 107,318,000 37,359,000 ,262,000 3,983,245,000 1The rgo1 crop is given by Harrison (1) as 1,351,045 muids (4,825,160 bushels). 2In the year 1894-5 the total crop of Cape Colony was given as 920,369 muids (3,287,032 bushels), (Wallace, 1). CHAP. LIL CHAP. Ill. 48 MAIZE East Africa, Madagascar, Mesopotamia, Ceylon, China, Japan, the Malay Archipelago, and New Caledonia. 36. Distribution in the United States.—The United States has 108,750,000 acres under maize and produces 75 per cent of the world’s crop, but though maize is grown to a greater or less extent in most of the States of the Union, 58 per cent of the crop is produced in the comparatively small region com- prising the seven central States of Iowa, Illinois, Nebraska, Kansas, Missouri, Indiana, and Ohio. These are known as the “Corn-surplus States,” because they are practically the only States which grow more than is required for their own consumption. Their combined area is only about 268,000,000 acres, or 114 per cent of the total area of the United States, and some 25,000,000 acres less than the area of the Union of South Africa excluding native territories. Only 18 per cent of the land of these seven corn-surplus States is planted to maize, but it produces 481,614,384 muids, or 58 per cent of the total crop of the country. The area of the Transvaal is approxi- mately 71,000,000 acres; if only 18 per cent were under crop to maize, and if the average yield were only 5 muids per acre (only half the average of the Corn-belt), the Transvaal would be producing the respectable crop of 64,000,000 muids of maize. Tasie IX. STATISTICS OF THE MAIZE-SURPLUS STATES. Squ A lant Yield i 06 ae « are Average plante jield in 1906. viel Corn-surplus States. Miles, lo Maize, Bichele per ree Bushels. Iowa . 7 . ‘: 56,025 9,450,000 373,275,000 390°5 Illinois é 5 * 56,650 9,616,886 347,169,585 36°1 Nebraska ‘ 4 * 77,510 7,325,000 249,782,500 34°1 Missouri. “i ° 69,415 7,075,000 228,522,500 32°3 Kansas : . E 82,080 6,750,000 195,075,000 28°9 Indiana ‘ : 3 36,350 4,643,782 183,893,767 39°6 Ohio . 5 : ‘ 41,060 3,325,000 141,645,000 42°6 419,090 | 48,185,668 | 1,719,363,352 36°15 The maize zone of the United States may, for practical purposes of competition with other parts of the world, be GEOGRAPIIICAL DISTRIBUTION 49 considered as lying east of the tooth meridian. It occupies the rich, alluvial bottom lands of the rivers Missouri, Missis- sippi, Ohio, and their tributaries. West of this there are, roughly speaking, four zones of vegetation, none of which is likely to become a maize producer of importance, owing to the climatic and other conditions described below. A_ recent American authority has stated that this country has now reached a point where increased acreage will play a minor réle in the future in the increased production of this great cereal (Bowman and Crossley, 1). 37. The Sub-arid Zone.—The western portion of the States of South Dakota, Nebraska, Kansas, Oklahoma, and Texas, lying, approximately, west of the 1ooth meridian, is a sub- arid zone of prairie, nearly 200 miles wide, where, without irrigation, good crops grow only one or two years out of five. The eastern border of this zone roughly corresponds with the 2,000 feet contour, where the country begins to rise from the river basin towards the Rocky Mountains. 38. The Rocky Mountains Zone.—West of the sub-arid zone lie the Rocky Mountains, comprising pastoral and forest areas. 39. The Great Basin.—Between the Rocky Mountains and the Sierra Nevada of California, lies the Great Basin, at one time known as the ‘‘Great American Desert”. Though the rainfall is scanty, this region is scarcely a desert in the ordinary sense of the word; it is largely covered with sage- brush and sparse grass furnishing grazing for stock. Crops are only grown where irrigation can be applied, and irrigated land grows lucerne more profitably than maize. 40. The Pacific Slope-—In the northern portion there is ample rain, but, as it falls principally in winter, the region is not well suited to maize-growing on a large scale. The irrigated lands can be more profitably planted to lucerne and fruit than to maize. A certain amount of sugar maize is grown for canning and eating fresh as “ green corn”. 41. The Atlantic States—The North Atlantic States of Maine, New Hampshire, Massachusetts, and the New England States, have too short a growing season to produce large crops of maize. Sugar maize is extensively grown, however, for canning purposes, even in localities too far north to permit of the ripening of the grain. In New Hampshire maize is grown 4 CHAP. III. CHAP. ITI. 50 MAIZE in greenhouses for the very early market (Rave, 1). Virginia, North Carolina, and Georgia are the only ones of the Atlantic States which produce any quantity of maize grain. The im- mediate coast region of the South Atlantic States is not much of a maize zone, partly on account of the character of the soil. 42. Canada.—The Canadian climate, speaking broadly, is better suited to wheat than to maize. But a number of the earlier-maturing breeds are grown and ripen grain in the eastern portion of Ontario Province, and the southern part of the Provinces of Montreal and Quebec. Ontario has about 332,000 acres, and Quebec 33,000 under maize. 43. Mexico.—As is the case in most of the Latin-American countries, the agricultural resources of Mexico are by no means well developed. The topography and consequent climatic conditions stand in the way, but a good deal more could be done by the development of irrigation, The north-western States of Lower California, Sonora, and Chihuahua are very dry, the average (twenty-two years) rainfall in the case of Lower California being only 10°5 inches. In most parts of the country maize and beans (frijoles) form the staple article of diet. The principal maize areas are south of the States of Sinaloa, Durango, Nuevo Leon, and Tamaulipas. In the drier States, such as Lower California, Coahuila, Durango, Nuevo Leon, and Yucatan, maize is mainly grown under irrigation, yet even in some of these, especially Sinaloa and Chihuahua, the maize area is considerable and is in- creasing. In others the crop might be doubled by irrigation. The strip of coast land is largely devoted to special tropical crops. The Maize-belt lies between this and the dry interior, at an altitude of 3,000 to 6,000 feet, where the climate re- sembles that of Southern Italy. Maize is the chief product of the States of Aguas Calientes and Colima. The estimated yields per acre range from 44 to 14 muids (15 to 50 bushels). In some parts two crops a year are produced. In some States American capital, machinery, and enterprise are being applied with promising results to maize- and cattle-growing. 44. Central America and the West Indies.—Maize is grown to a limited extent throughout Central America, but though used as an article of diet, it does not form an important article of commerce, other tropical products paying better. GEOGRAPHICAL DISTRIBUTION 51 45. Tropical South America.—Maize is grown to a greater or less extent in all of the tropical South American countries, but statistics of production seem to be non-existent in most cases, Chile is reported to have had 63,100 acres under crop in 1908, and Uruguay 524,200 acres in 1907. Peru and Bolivia are also known to produce for their own consumption. In Brazil maize is grown in the more open parts, two crops a year being produced in some places. Flint breeds form prac- tically the only variety grown, and the yield per acre is said to be higher than in the United States. The dense tropical forests which clothe the river valleys are unsuited to the pro- duction of maize, and there manioc largely takes its place. 46. Argentina,—Argentina is the most serious competitor with South Africa for the maize trade of the world, and the only country that she has to fear at the present time. In the season 1908-9 Argentina had 8,342,559 acres under maize, and her crop was 49,590,000 muids, or an average of nearly 6 muids per acre; on some farms 14 to 17 muids (50 to 60 bushels) are obtained. The planting season extends from the middle of August , to the middle of January, but the safest time is considered to be from the middle of September to the middle of December. Early planting gives the best yield when the season is favour- able. The crop is drilled, not check-rowed; it is harrowed when the plants are 2 or 3 inches high, and hilled up by machinery when 12 inches high. Ninety per cent of the crop consists of a small flint type, much appreciated on the London and Liverpool markets; Hzckory King and Queen have also been tried. There is a tendency to harvest before the crop is mature, in order to get the grain to the coast before the heavy rains begin ({ 29). In some seasons locusts play serious havoc with the crop. The Provinces of Buenos Aires and Santa Fé are the largest producers. 47. Possible Increase in the Argentine Crop.—Fifty per cent of the total crop is exported, but it is likely that local consumption will increase owing to the enormous develop- ment of the meat-packing trade. However, with increased local demand, there will certainly be an increase in acreage under crop. During the five years 1905 to 1909 inclusive, the average annual increase in area planted to maize amounted 4* CHAP. III. 52 MAIZE CHAP. to 78 per cent, the production 28 per cent and the export 33 III. per cent. Only about an eighth part (say 12 per cent) of the area suited to the production of the four crops, wheat, maize, linseed, and oats, is at present under cultivation. Of this, one-half is devoted to wheat and one-quarter to maize. Eight times the present maize acreage would be 66,000,000 acres, which at 6 muids per acre gives a potential crop of 396,000,000 muids. Unless the population in the country increases, this means a possible surplus of 198,000,000 muids for export. The importing countries consumed, in the year 1908-9, only about 49,689,180 muids. If the present rate of increase in Argen- tine maize exports is maintained, viz. 33 per cent increase in four years, it will take twenty-eight or perhaps thirty years for the export to reach the maximum indicated above. By that time local consumption in Argentina will probably have increased greatly, as it has done in the United States. Out of a crop of 850,000,000 muids (more than double the potential crop of Argentina) the United States exports only about I°5 per cent or 8,500,000 muids. If the States continue to export I per cent, and the Argentine export falls to 1 per cent of her potential crop, the two countries will only export about 12,500,000, or one-quarter of the present world’s imports, instead of ¢hree-quarters, as at present. This will leave ample opportunity for South Africa to supply 25,000,000 to 30,000,000 muids, while the Danube and South Russia can make up the balance. This estimate makes no allowance for increase in consumption by importing countries, which is steadily growing. Under these circumstances there seems to be plenty of room for the expansion of the South African maize trade. 48. Europe.—In the ten years from 1880 to 1890 the pro- duction of maize in the Austro-Hungarian Empire is said to have increased by 40 per cent, but during the last few years the maize acreage and production have been fairly uniform, and no great increase in future is to be expected. Most of the crop is grown on the rich, alluvial plain soils of the Danube, Dnieper, and Dniester Rivers, Roumania, Hungary, and Bessarabia together furnishing over half of the European crop. This is largely exported from Odessa, Galatz, and Fiume; grain forms three-quarters of the export trade of Galatz. GEOGRAPHICAL DISTRIBUTION 53 In Roumania maize is the staple crop, and the staple food-stuff of the people; there are local distilleries which produce whisky from the maize. Italy is the next largest producer with about 15 per cent of the European crop. The largest and richest. agricultural area is the basin of the River Po, including the plains of Lombardy, Venetia, and Emilia. Part of the crop is shipped from Genoa and part from Venice; at the latter port the grain is stored in air-tight silos to await shipment. 49. Asia.—Very little information is obtainable as to the culture of maize among Asiatic peoples. Maize is grown in India, Ceylon, Persia, China, Japan, and the Malay Archi- pelago, but few statistics are accessible except in the case of India. The value of maize as a cereal crop is strikingly emphasized by its distribution in the latter country ; though doubtless first introduced into the Portuguese Settlements of the East Indies early in the sixteenth century, the conservatism of the Indian peoples naturally stood in the way of its adoption as a regular crop. As recently as 1832, Roxburgh observed TABLE X. ACREAGE UNDER MAIZE IN INDIA FOR THE DECADE 1897-8 TO 1906-7, BOTH INCLUSIVE! | British India. Native States. | Total. | ~ as oe ie ao Acres. Acres. Acres. | 1897-1898. : : ‘ 6,414,732 | | 1898-1899. : : ; 6,144,240 | | 1899-1900. a ‘ é 5,195,472 | 1g00-IgoT. : ‘ : 5,849,533 | IQOI-1g02. ‘ rl : 6,198,063 | 1902-1903. i i : 6,331,816 306,346 | 6,638,162 1903-1904. . . . 6,135,511 269,268 6,404,779 1904-1905. ; : : 5,961,487 250,285 | 6,211,772 1905-1906. “ ‘ ‘ 51790,543 221,687 | 6,012,230 Ig06-1g07. , ‘ ‘ 6,171,751 302,350 6,494,101 | Average . ‘ 6,019,514 269,987 | 6,352,209 The distribution of the crop through the several Provinces is instructive ; the highest yields per acre are obtained in the North-west Frontier Provinces; the lowest (as far as we have figures) in Bengal; but Bengal has by far the largest acreage. 'Indian Government Publications, 1. CHAP, Til. CHAP. Ill. 54 MAIZE TasBLe XI. ACREAGE AND YIELDS OF MAIZE IN THE INDIAN PROVINCES. . Acreage rs Hetimated Province. | Planted. Yield per Acre. cae Bengal . i ‘ 1,802,400 820 Ibs. 7,389,840 Agra : . + 1,454:497 1050 5, 7,636,109 | hee », irrigated | ; 850 ,, dry i x Faaab : : 1,195,849 pee 5, dry & ieee | Bpauegs 1007 ,, mean Oudh wl, 710,938 | 3050 ,, 35732,424 gee - noes ] re 1] 2 J 745 ” Ty 1 | N.W. Frontier f 390,529 | 342 1 eng Se irvigated | | 2,556,012 I309 ,, mean Bombay . F f 157,079 850 ,, estimated 667,585 Central Provinces . 135,852 850 ,, is 577,371 Upper Burma . 106,700 850 ,, + 453475 Madras . P ‘ 104,913 850 ,, < 445,380 | ( ad rm Cie aa ee J 960 ,, dry E Bo Ajmer-Merwara =| 64,003 | sig, diy & irrigated | 563,226 880 ,, mean Lower Burma. : 21,879 850 ,, estimated 2,985 Eastern Bengal e 21,120 850 ,, - 89,760 Berar : . : 2,116 850 ,, 7H 8,993 Sind F : : 1,394 850 ,, ‘a 5,924 Central India . : 1,269 850 ,, ae 55393 Assam. ‘ ‘ 1,213 850 _,, ‘8 55155 Coorg ‘ * Bee PU oReeno dacesees ids Native States . : 302,350 850 ,, a 1,284,987 | 6,474,101 31,536,218 1 Where the yield per acre varies between the irrigated and dry crops, and the crops fully irrigated and partly dry, the mean of these figures has been taken in calculating the yield. Where no yield per acre is given, the estimate has been made on the basis of the yield of the dry-land crop in the Panjab, which is probably a little on the low side. that maize was “cultivated in different parts of India in gardens, and only as an ornament, but nowhere on the con- tinent of India as an object of cultivation on a large scale” (Roxburgh, 1). In course of time native prejudice gave way before the unanswerable demands of hunger and a rapidly increasing population; Church (1) observes that in 1886 there were already in India 2,250,000 acres under maize. In another twelve vears the area under crop had increased to GEOGRAPHICAL DISTRIBUTION 55 6,500,000 acres. Since then it has averaged about 6,352,000 acres per annum, with very little variation above or below, from year to year, indicating that it has probably reached its limit of geographical and economic distribution. Maize has now become a staple article of food in India, especially among the hill peoples. The average yield per acre ranges from 714 lbs. to 1,841 lbs. (i.e. from less than 4 muids to over 9 muids, the latter yield being obtained only under irrigation) ; the average production for all the States and Provinces is slightly over 5 muids per acre. Maize is grown more exten- sively in the hill country than on the plains, where it is largely replaced by rice; in the Panjab it is grown at about 7,000 feet altitude. In Baluchistan a dwarf breed is grown successfully at between 5,000 and 9,000 feet altitude, where it forms a staple food of the people (J/weller, 1). It seems hardly probable that India, with its dense popu- lation and increasingly intensive agriculture, will become a maize-exporting country; it is more likely that she will be an importer, in exchange for some of the more costly articles of commerce which she produces in abundance. 50. Australasia,—The total Australian maize area is only about 385,000 acres, and the crop about 3,000,000 muids, of which New South Wales contributes roughly one-half and Queensland over one-third. The coastal belt appears to be best suited to maize cultivation; two crops may be grown each year on the low coast lands of South Queensland. The climatic conditions of the interior of the continent appear to be generally too dry for maize-growing on a large scale, and the irrigated lands are too valuable for lucerne and dairying, to be devoted to maize-grain growing. Only a small quantity is produced in New Zealand. In New Caledonia maize is the principal cereal grown; it is used for feeding horses (replacing barley and oats), work-oxen, pigs, and poultry, but not for human food ( /eanney, 1). From the point of view of competition in the European trade, Australasia is not likely to be a serious competitor with South Africa, owing to the much greater distance from market, and the limited area available for maize production. Development of the Australian meat trade may lead to a larger consumption of maize, and to a steady import, in CHAP. Ill. CHAP. Ill. 56 MAIZE which case South Africa would be the nearest producing country.! 51. North Africa.—Egypt is the largest producer of maize in North Africa, having nearly 2,000,000 acres under crop, and producing some 8,500,000 muids. Maize is a staple article of diet, and a certain amount is imported annually. The Egyptian Sudan produces only a small quantity, an average of 84,000 muids per annum for the three years to 1908 inclusive. In Algeria, owing to lack of summer rains, maize occupies but a very limited area, almost confined to the Province of Oran (Azviere and Lecog, 1), and the annual production is only about 125,000 muids, 52. Tropical Africa.—Excellent maize is grown in parts of Rhodesia, and this Colony is likely to become a very large producer. A small export trade has been started (valued at about 430,000 in 1912), through the port of Beira. The development of cattle-ranching on a large scale is likely to lead to greater local consumption; 460,000 worth of maize was imported from the Union of South Africa in 1912. In Nyassaland a small quantity of maize is grown and there appears to be scope for development, but it is probable that more intensive crops such as coffee and cotton will prove more profitable for that Colony. On the uplands of British East Africa and the Uganda Protectorate maize-growing is on the increase, and it is possible that this region will become a competitor with South Africa for the export trade. The Suez Canal charges may, perhaps, more than offset the shorter sea- distance to Europe, but there seems to be a good opportunity for developing the trade with India, Ceylon, and China. Comparatively little maize is produced in the remainder of tropical Africa, partly owing to the general lack of agricultural development, partly to the fact, already alluded to, that maize does not thrive in tropical forest country, and partly, also, to the depredations of the elephant and wart-hog. In Italian Somaliland, German East Africa, Madagascar, the Astove atolls, the Cosmoledo Islands, and Portuguese East Africa, small quantities are produced, but the coast conditions do not seem favourable to maize production on a large scale, and other tropical crops generally pay better. 'Since this was written Australia has imported 242,000 muids of South African maize in one season. GEOGRAPHICAL DISTRIBUTION 57 Maize seems to be almost unknown to many of the native tribes of equatorial Africa, probably for the reasons before given (11 21 and 30). It is grown by the Baambas to the west of Mt. Ruwenzori, and by the Unyoros of Uganda, but “bolu” (Eleusine Coracana) is said to be the favourite cereal of the latter people. In the Acholi country, millets appear to take the place of maize, except at a small Nubian settlement near the Nyama River, where a red kind of maize is also grown. A small quantity of maize is grown by the Bahoru at Katonia in the Ankole country east of Mt. Ruwenzori (Dawe, 1). Major Bright (1) states that maize is grown in large quantities in the plain around Kasenyi, on the west shore of Lake Albert, at about 2,170 feet elevation, where they are not troubled by elephants and wart-hogs. In Nigeria maize does not appear to have yet supplanted the native cereal, guinea corn (Sorghum vulgare var.) (IVhitlock, 1). In Portuguese West Africa a limited quantity of maize is pro- duced, and in the Congo country it is used in the preparation of a beverage. In 1795 Mungo Park (7rave/s) found the natives at Pisania, on the Gambia, cultivating maize in considerable quantities. The French Sahara is too dry for maize-growing ; water is plentiful in the country about Lake Chad, but the heat is intense and the rainfall very scant; during the rainy season (July to October inclusive), the mean fall is about 5:2 inches, while in the very wet year of 1908 it only reached 7°8 inches. In this region millet seems to be the staple cereal (77¢ho, 1). 53. South Africa—As a field for maize-growing, the Union of South Africa takes front rank, and for the farmer with energy and enterprise there awaits a rich reward in con- nection with this industry. A young, vigorous, and steadily (if slowly) increasing population provides an expanding local consumption, and the world’s markets—owing to the excel- lent lines of communication linking South Africa with the older countries—lie within easy distance. The climate of a large part of South Africa is peculiarly well suited to the easy production of enormous quantities of maize of exceptionally good quality, especially for manufac- turing purposes. The rainfall is ample if the soil is cultivated properly. The possible planting season lasts for two months, CHAP. Ill. CHAP. III. 58 MAIZE as compared with a maximum limit of eighteen days in some of the maize-growing States of North America. The moisture contained in maize exported from South Africa is some 4 per cent lower than that of the American-grown article, which minimizes danger of damage in transport, and puts a premium on South African grain for manufacturing purposes. Farmers in Argentina find their climate unfavourable to the proper conditioning of the crop for export (@ 29). Not all of South Africa is equally well suited to the pro- duction of maize. She has her Maize-belt just as the United States has her Corn-belt. The hot coastal zone and the dry Karroo and Kalahari regions are not well suited to maize- growing. The maize zone may be roughly defined as the country lying east of the 26th meridian, ie., a line drawn between Algoa Bay, Bedford, Cathcart, Queenstown, Aliwal North, Wepener, Bloemfontein, and thence north to Lichten- burg and Zeerust. From this area the coast belt below 1,000 feet altitude, and the mountain region above 6,000 feet, should be excluded. 54. Orange Free State.—The Orange Free State, together with the adjoining native territory of Basutoland, is by far the largest producer and exporter of maize of any of the four Provinces of the South African Union. But of the total area of the Province less than 2} per cent is planted to this crop, and in the best producing Districts only 54 per cent. The largest acreage and best crops are found in the north-eastern Districts, where the rainfall is about 11 inches during the three growing months of December, January, and February. A con- siderable part of the crop is grown by natives, and the average yield is estimated at only 3 muids, or 11 bushels, per acre. 55. ZLvansvaal.—Maize is grown more or less in every District and on practically every farm, but the principal Dis- tricts, in approximate order of production, are: Bethal, Heidelberg, Potchefstroom, Pretoria, Standerton, Ermelo, Middelburg, and Lichtenburg. Most of the Transvaal maize is produced on the High-veld, because the population is greater and more land is under cultivation. A good deal is also raised in the Upper and Lower Bush-veld, but chiefly by natives. The altitudinal range of the crop is from 600 feet at Komatie-poort, to over 6,000 feet in the Steenkampsberg and GEOGRAPHICAL DISTRIBUTION 59 Drakensberg ; but the major part comes from the plateau be- tween 4,000 and 5,500 feet. The south-western Districts are less suited to the production of the ordinary types of maize owing to low rainfall, short growing season, and often shallow soil. But it is probable that in time breeds will be developed especially suited to the climate and soil of that part of the country. 560. Relative Vtelds of Transvaal Districts.—The relative productiveness of a District cannot be determined from the actual number of bags of grain produced by it, for the simple reason that the areas of the different Districts are so enor- mously disproportionate; as an example we need only com- pare that of Bethal (384,000 morgen) with the Zoutpansberg, which is nineteen times its size (7,256,400 morgen’). The best method of comparison of relative productiveness is to reduce the yield to the average of some unit common to all, e.g. an acre, morgen, or square mile. The writer has therefore reduced the maize yields of the Transvaal to the average per square mile, which is the most convenient unit to use in the present state of agricultural de- velopment. In the following table (XII) the Districts are arranged in order of productiveness. If we considered only the total production of a District, we should have to give Lichtenburg first place with 191,405 bags, whereas in yield per square mile of veld she comes only sixth on the list. The value of closer settlement and consequent improve- ment in cultivation of the soil are clearly brought out by the fact that the Witwatersrand goldfields produce more maize per square mile than any other District of the Transvaal. The Witwatersrand, comprising the Magisterial Districts of Johannesburg, Germiston, and Boksburg, is the most thickly populated area in the Transvaal; it covers only 556 square miles, a large part of which is occupied by mines, mine dumps, towns, and villages; yet in 1909 it produced 68,400 muids of maize, or 122°8 per square mile, almost doubling the yield of Bethal, the next largest producing District. , The soils of the Witwatersrand are not as suitable for maize culture as those of many other parts of the country. 1] morgen = 2°1165402 acres. CHAP, Ill. 60 MAIZE TABLE XII. AREAS AND YIELDS OF TRANSVAAL DISTRICTS. | | Average in Muids per Square Mile. | Area.1 District. Square Miles. Morgen. 2 Muids. Pee | 1908-9. 1909-10. # blag «spony St | Johannesburg. | 18 4°18 | 55,692 | ‘Total Wit- | 556°78 | 168,358 68,399°5 | 122°8 | -122°8 watersrand (excluding | | Krugersdorp Bethal a a I,270°04 334,035 93,5550 | 93°3 | 170°0 | Heidelberg : 2,351°O1 711,081 140,496°0 59°7 102°9 Standerton . 2,003 62 | 605,855 119,062°0 50°4 go's Potschefstroom . | 4,904°15 1,482,918 | 190,653°3 38°8 58°7 Lichtenburg ‘ 4,478°81 1,354,304 IQI,405'0 42°97 35°0 Wolmaransstad . 2,0061'69 623,414 51,929'0 25°1 29° Middelburg : 5,028°98 1,520,664 | 106,796°5 213) 371 Ermelo ss : 3,003°43 908,178 56,7510 189 36°0 Wakkerstroom . | 2,197'76 664,559 39,123°5 17°38 20°9 Krugersdorp : | 1,174°67 355,197 19,800°5 16°8 25°5 Piet Retief. | 1,615°93 | 488,625 25,100°0 15'5 31°5 Pretoria 6,641°54 | 2,008,272 73,486°5 Ilo 376 Carolina . vl 2,095°72 | 633,704 14,231°0 6:3 27°6 Bloemhof . | 3,703°53 965,661 19,754'0 62 79 Marico ; 3,636°89 1,099,724 18,423°75 570 Sr Lydenburg | 10,176°84 3,077,276 43,239°5 4i2 | 6-9 Waterberg . : 15,625'°77 | 4,724,925 64,242°0 4°I 69 Zoutpansberg. 23,997°78 7,256,455 65,3680 247, 16°1 Rustenburg ‘ 95739°73 2,942,381 25,017°5 2°6 O°4 Barberton . ° 4,679°57 I,415,010 9,900°5 21 37 | eer Total i 110,425°84 | 3313701596 | 1,437,834°9 | 12°9 | 42°0 1 Figures of area kindly furnished by the Surveyor-general of the Transvaal; statistics by the statician of the Department of Agriculture (Foubert, 1). 2 morgen = 2°1165402 acres. %Estimated only. The actual crop was only about one-half of the expected, owing to a remarkably unfavourable season. 57. Natal.—Maize is produced in all parts of Natal, but the Midland Districts (2,000 to 3,000 feet altitude) are generally conceded to be the best for maize-growing. The coast-belt is better suited to sugar cane and citrus fruits than to maize, owing to the too rapid and luxuriant growth of weeds and con- sequent cost of cleaning the land; but the average yield per acre is 1} muids higher on the coast than ‘ up-country” LABLE A111. MAIZE PRODUCTION, NATAL, 1906-7. nate ne Acteabe | Total Yield per eae Magisterial Divisions. index Mase. | ae eae : Nata. | Coast — | Lower Umzimkulu. . ; 1,012 | 5,064 50 Alexandra . : ? : 4,180 | 27,250 68 ay ‘ : 7 Fe 782 4,576 59 Inanda \ : Indwedwe f A ‘ 4 1,631 10,417. 6°5 Lower Tugela \ a . Mapumulo— f° ; g 748 41939 oe as = Totals. ‘ 8,353 52,246 Average 6°1 Midlands— Impendhle . Z 3 : 1,089 | 5,968 5°90 | Alfred : ‘ F ; 1,291 6,142 49 | Ixopo a 75319 35,670 49. Richmond . ‘ . é 9,882 59,230 6:0 Umgeni and Camperdown . 16,087 73,488 45 New Hanover . ‘ : 8,908 49,113 56 Lion’s River : . : 4,983 22,398 4°7 Umvoti 2 : ; : 10,132 34,287 34 Krantzkop . A ; ; 2,242 3,837 40 Totals. : 61,933 295,133 Average 4°9 | Uplands— : Underberg . ‘ ‘ 1,098 4,980 4°4 Polela 5 ‘ ‘ : 92 4,290 46 Bergville . : : : 3,229 14,758 4°5 Estcourt. : : : 8,763 48,377 5°5 Weenen . 4 ‘ 1,018 4,963 49 Klip River . ‘ ‘ : 6,493 25,317 39 Umsinga . ; ‘ : 992 4,670 48 Dundee. : . : 8,372 34,190 4° Newcastle . ; , : 9,012 37,468 4° Vryheid | 6 N’Gotshef 1722 Sa i Utrecht. : : ; 1,934 7,896 ee Paulpietersburg . ‘ : 1,650 6,218 3°9 Babanango : ; , 7388 2,596 3°3 Totals. : 40,001 | 202,683 Average 4°3 ZULULAND, | Coast— | Eshowe : ; : dirs | ves ue Mtunzini . 3 : : 474 2,070 4 4 Lower Umfolosi z : 53 265 Bye) Hlabisa ie ves ae Ubombo Totals . 527 2,335 Average 4°7 Inland— Nqutu) | 29 476 470 Nkandhla J ‘ es Emtonjaneni_. d : 907 2,919 Bi Mahlabatini ve : Ndwandwe Ingwavuma eee Totals. : 936 3,035 Average 3°6 Grand Totals for the Province | 117,750 555432 Average 4°7 CHAP. III. 62 MAIZE Districts favourable for wattle-growing do not seem to be well suited to maize ; the rainfall and lack of sunshine are perhaps too great, or it may be that maize is not equally profitable and is therefore neglected where wattle is grown. Although the Uplands produce a considerable quantity (only 90,000 muids less than the Midlands), they seem better adapted to stock-raising than to agriculture. Camperdown and Richmond Districts are considered the TABLE XIV. MAGISTERIAL DISTRICTS OF NATAL ARRANGED ACCORDING TO PRODUCTION. Yield per Division. Acres. Square Miles. Square Mile in Muids. Upper Umkomanzi— Richmond . . , - 332,800 520 IIq4'o Camperdown . : : F 235,520 368 : Umgeni . 4 ‘ A r 233,600 ca p02 New Hanover 3 : 3 332,800 520 04°4 Alexandra 5 A % é 428,800 670 40°7 Umvoti . r : ‘ : 550,400 860 39°9 Ixopo.. ‘ . F . 624,640 976 365. Lion’s River . . 4 f 403,200 630 35°5 Dundee . . 7 . é 605,440 946 35°4 Newcastle : : ‘ : 698,880 1092 34°3 Estcourt . é 4 : . 1,164,800 1820 26'5 Inanda 8 : Indwedwe f ° ‘ : ‘ 203,520 443 235 Klip River F % 4 Z 920,960 1439 17°6 Krantz Kop. . z 3 366,720 573 I5"4 Bergville : . i ‘ 704,000 II00 13°4 Alfred. : : 4 . 364,800 570 10'8 Paulpietersburg ‘ ‘ . 395,520 618 roo Lower Umzimkulu . : i 341,760 534 9°5 Umlazi . E . é . 320,000 500 QL Impendhle . E : ; 448,000 700 8°5 Weenen . ‘ . 400,000 625 8-0 Polela and Underberg 2 ‘ 774,400 1210 76 Umsinga ‘ i ‘ 394,240 616 75 Lower Tugela. : : : 289,920 453) 6:8 Mapumulo_. , : z 172,800 270f M’Tunzini. F : : 239,360 374 5°5 Emtonjaneni . : < ‘ 414,720 648 4°5 Utrecht . F 2 F 1,310,720 2048 3°83 Vryheid, N’Gotsche, and Babanango . 2 : P 2,729,600 4265 22 Lower Umfolosi . r . 656,000 1025 2 Nqutu_ . : . F ‘ 400,000 625) i Nkhandla 2 ‘ : : 487,680 762 f 08 GEOGRAPHICAL DISTRIBUTION 63 best part of the Maize-belt of Natal ; the rainfall at Mander- ston is about 30 inches, One of the pioneer maize-growers of this part of Natal, famous for his large-sized H7ckory King, has only 220 acres under maize, but has harvested as much as 3,581 muids in one season, or an average of 16:28 muids per acre, while 10 acres averaged 22 muids; it has been his am- bition to get a crop of 4,000 muids from his 220 acres. He started maize-growing twenty-five years ago, and the first sea- son only produced 5 muids per acre. Good farming and the regular use of bone-meal have brought up the producing power of the land from a non-paying to a profitable yield. In Table XIII the acreage given for the various Magisterial Divisions excludes the Boroughs of Ladysmith, Newcastle, and Dundee, and the townships of Greytown, Verulam, Utrecht, and Vryheid. Those divisions of Zulu- land for which there are no returns have a few hundred acres put under maize by Europeans, but altogether for local con- sumption (Harvey, 1). 58. Cape Province.—The Cape Province produces less maize in proportion to total area than any of the other Provinces. The lack of summer rains in the south-western portion, the old “ Western Province,” renders that part of the country but poorly adapted to maize culture. In the Karroo and North-western Districts of the Province the total rainfall is too low to produce good crops of maize. In some of the Eastern Districts, on the other hand, especially the Transkei, the climate is favourable to the production of excellent maize crops. In these Districts, however, there does not appear to have been the same rapid increase in production that is notice- able in some other parts of South Africa ; in fact, in the twelve years from 1895 to 1907 the production fell in the District of Victoria East from 28,000 muids to nearly one-third, while in King William’s Town District it fell off 15 percent. In other Districts, however, the production increased, and in two or three cases it doubled and trebled. CHAP. Ill. CHAP. HI. 64 MAIZE TABLE XV. MAIZE PRODUCTION, CAPE COLONY, 1906-7. | Estimated Yield. Muids. e Area in | Average Districts. oe Square |—-— | sduare ; Miles: 1894-5. lise 0 1906-7. | “Mile. —— | Komgha . —. —.| 342,400 | $35 13,651 38,722 2"4 | East London : .| 428,800 670 20,480 40,556 60°5 | King William’s Town . 807,040 1261 si ia 71,705 56°9 Stutterheim . . i 428,800 67 | 33,547 50°0 | Bathurst : ¥ . 373,760 584 ‘a 23,653 49°0 | Stockenstroom 7 200,320 313 11,674 | 13, 0364 | 41°7 | Alexandria. r : 615,040 g6r 39,329 40°9 | Victoria East : : 241,920 378 28,260 10,740 | 2874 | Fort Beaufort c ; 545,920 853 13,722 17,314 | 209 | Cathcart : : i 634,240 ggI Ess 18,276 | 184 | Knysna ‘ . fe 621,600 815 10,624 13'0 Aliwal North 5 4 613,760 959 ae 11,645 | ror |Queenstown. . — .| 1,232,000 | 1925 17,315 22,990 II'9 | Humansdorp : . | 1,235,200 1930 9,317 19,7774 | 10°22 | Albany . és : . | 1,066,240 | 1666 be 14,967 go | Wodehouse . : . | 1,254,400 1g60 ae II,009 5°6 | Mafeking. : - | 2,345,600 | 3665 aan 12,870 35 | Herschel | ae 693 32,880 wes ae | | Glen Grey | 825 27,860 | Bedford | 1200 13,277 | Oudtshoorn . 1630 11,228 | | Peddie . P 619 9,704 | Somerset East * | | 2900 8,576 | Graaf Reinet -| | 2651 7,859 | | | | 415,780 | ! | Addendum,—The possibilities of Rhodesia have been re- ferred to in 1 52. Average yields of ten (and even twenty) bags of maize, year after year, are reported. It is stated that about one-third of the whole of the Mazoe Valley country could be planted to maize and that in the Kafue Valley of N.W. Rhodesia there are vast stretches of level alluvial soils, suitable for maize, recalling the level plains of the North American Corn-belt. The total area of Rhodesia is approxi- mately 250,000,000 acres ; if only one-fiftieth part were planted to maize, and yielded on the average only 7 muids per acre, the country would be producing 35,000,000 muids. CHAPTER IV. BOTANICAL CHARACTERS. Day by day did Hiawatha Go to wait and watch beside it Till at last a small green feather From the earth shot slowly upwards, Then another and another, And before the summer ended, Stood the maize in all its beauty, With its shining robes about it And its long, soft, yellow tresses. And in rapture, Hiawatha Cried aloud, “It is Mondamin! Yes, the friend of man, Mondamin!”’ —Hiawatha. Tis sweet... to... scent the breathing maize at setting day. —COLLINS. 59. Botanical Relationship.—Maize belongs to the group cHap. of Monocotyledons, and the family Gramineew, or Grasses ; it is the type of the tribe Maydee to which the genera Euchlena, Coix, and Tripsacum also belong. Of these, Euchlena (Fig. 10) is its nearest relative, and the only one with which it is known to hybridize. Some botanists are inclined to consider Euchlzna as the prototype of Zea, for the latter is not known in a truly wild state. Montgomery considers that Zea and Euchlana may have had a common origin, and that in the process of evolution the pistillate spikes in teosinte were probably developed from the lateral branches of a tassel-like structure, while maize was developed from the central spike (cf. Fig. 40) (Bowman and Crossley, 1). 60. Descraption.—The maize plant (Figs. 11 and 12)isa tall, annual, moncecious grass, with stout, erect, solid stem, and broad leaves; the staminate flowers form a terminal panicle ; the pistillate flowers are arranged in a densely-crowded spike, 65 5 IV. 66 MAIZE CHAP. IV. be) v < S s © Fic. ro.—Teosinte, Euchlena mexicana, a near relative of the maize plant, and the only species with which it is known to hybridize. BOTANICAL CHARACTERS 67 the ‘ear,’ terminating a short lateral branch and closely CHAP. enveloped in leaf-sheaths called the husk; the long styles, !Y exserted in anthesis, form the sz/k or beard. 6t. Plant Structure—All plants are living organisms, which feed and breathe in order to grow and multiply their kind. Their food-material consists of water, several of the Fic. 11.—Maize plants in the Transvaal. chemical substances of which the soil is composed, and carbon which is obtained from the air. Plants are built up of a vast number of ceé/s of different forms (Fig. 13); the cell is a microscopic sac usually consisting of a cell-wall surrounding a jelly-like mass called protoplasm (Fig. 13C). The cell-wall is a colourless membrane composed 5 * CHAP. IV. 68 MAIZE of cellulose ; as it surrounds each cell, cellulose forms a large part of the substance of the higher plants. Protoplasm is the Zving substance of the plant, and comprises various minute, differentiated bodies, some of which (‘he chloroplasts) contain a green colouring matter (chdorophyll), and are present Fic. 12.—Flowering plants of maize (Zea Mays). A, Tassel. B, Leaf- blade. C, Leaf-sheath (the stem is entirely surrounded by the sheath). D, Ear surrounded by husks. E, Silk exposed for some days. F, New silk just appear- ing (Photograph by D. W. Macdonald). in such enormous quantities that they cause the whole plant- surface to appear green. In the more highly-developed forms of plant life the cells are not massed together promiscuously, but are associated in groups forming bands, plates, or cylindrical masses called BOTANICAL CHARACTERS 69 tassues (Fig. 14). The tissues of such plants become differ- entiated into groups, forming organs which have different duties to perform, Of these organs the most important are the root, stem, and leaf which are connected with nutrition and growth, and the jfower which contains the reproductive organs of the plant. The life-cycle of the maize plant begins with the fertilization of the egg-cell in the ovule by the male germ cell, as described in § 78, by which a new plant-being is brought into existence. The new cell, formed by the union Fic. 13. Fic. 14. Fic. 13.—Plant cells, as seen under a high-power microscope, showing strands of protoplasm, nucleus, nucleolus, etc. A, B, young cells; C, an older cell, from the developing maize root; D, cell from the hair of Tradescantia; E, parenchymatous cell from the cortex of Ranunculus, (From Sir F, Darwin’s Elements of Botany, Cambridge University Press.) Fic. 14.—Transverse section through a leaf (of hellebore), showing tissues and cells. From above downwards are seen the upper epidermis, the palisade cells, the spongy tissue (in which a vascular bundle is seen), the lower epidermis (in which is shown a single stoma opening into a large intercellular space). Note that the chloroplasts are arranged along the cell walls, especially in the palisade tissue (A). (From Sir F. Darwin’s Elements of Botany, Cambridge University Press.) of the two germ-cells, develops by cell-growth and repeated cell-division (somatic division) into a seed. 62. Lhe Seed.—The seed consists of an embryo plant and a mass of reserve food-material known as endosperm, wrapped in two protective seed-coats, the outer or ¢esfa and the inner or fegmen. In the case of maize and other grasses, the seed is further surrounded by the perzcarp or envelope of the grain CHAP. IV. CHAP. IV. 46 MAIZE or fruit, which in many kinds of plants encloses more than one seed, but in grasses and some other plants only one. We may therefore define the seed as a miniature living plant and its food-supply, wrapped in a protective envelope. The pro- duction of seed is a provision of nature to enable a living plant to remain dormant during a period when climatic con- ditions, as, for example, an intensely cold winter or a long, dry summer, are unfavourable to its existence as an active organism. 63. The Embryo and Endosperm.—The embryo (Fig. 15B) is the vital part of a seed; it is a living, though dormant, plant-in-embryo. The embryo comprises all the essential Fig, 15.—Embryo and endosperm of maize. A, Section through maize grain showing relative position of embryo and endosperm. a, hull; 6, aleurone layer; c, horny endosperm; d, white starchy endosperm; e¢, plumule; /f, radicle ; g, scutellum. B, Embryo removed. C, Germinating embryo. organs of growth, i.e. the vadzcle which develops into the root of the plant; the cotyledon or seed-leaf; and the plumule or young shoot, which develops into stem and leaves. In maize and other grasses the growing embryo absorbs the endosperm through a special organ, the scztel/um (so named from its shield-like shape), which in maize can readily be seen with a low-power microscope. The embryo of the maize-grain lies to one side of the endosperm (Fig. 15A); as the grain stands on the ear, the embryo is on the upper side of the grain, ie. facing toward the tip of the ear, its position being indicated by an oval depression in the grain. BOTANICAL CHARACTERS TL The endosperm (Fig. 15¢) consists of a store of prepared CHAP. food-material which the growing embryo absorbs for use in 1: the formation of new cells and tissues, in developing a root- system with which to absorb food-materials directly from the soil, and a leaf-system capable of photosynthesis (4 65). The seeds of some plants (e.g. the castor-bean and the Fic. 16.—Germinating maize grains, showing developing shoot, primary root, root-hairs and adventitious roots. ({1 66.) lucerne) contain no endosperm, but store in the cotyledons a supply of food-material for the use of the growing seedling ; such seeds are sometimes spoken of as ex-albuminous, in contradistinction to the albuminous or endosperm-bearing seeds, 64. Germination.—The commencement of growth in a fe CHAP. AV. res MAIZE hitherto dormant seed is known as germination (Figs. 15¢ and 16). A seed will not germinate until it comes under the influ- ence of favourable conditions, and may lie dormant for many years until such conditions supervene; these conditions vary with different sorts of plants, but all include: (1) moisture ; (2) heat ; and (3) sufficient air for the growing plantlet to breathe. The seed of the maize plant will retain its vitality for two or even three years; but after the first year there is a marked decrease in vitality, and after two years maize is considered practically useless as seed ; this is independent of any question of injury by weevil or grain moth, and may, perhaps, be con- nected in some way with the presence of a considerable quan- tity of oil in the embryo; this oil readily turns rancid at high temperatures. Experiments conducted at the Botanical Experiment Station of the Department of Agriculture, Pretoria, show that it requires from three and a half to eleven days after sowing the maize-seed for the seedling to appear above the ground. The difference in time of germination is largely influenced by the warmth of the soil; but it is evident that temperature is not the sole controlling factor, and that associated with it is the degree of moisture of the soil. Depth of planting also affects germination, deep planting tending to delay it; very shallow planting (i.e. less than 2 inches) has in South Africa a similar effect, perhaps because the surface soil is more rapidly affected by drought. The germinating embryo depends on the endosperm for its supply of food-material until it has developed a root- and leaf-system ({ 63). In germination the radicle grows downwards or earthwards, and is therefore said to be geotropic; the plumule upwards or away from the earth, and it is therefore called apogeotropic. If the seed be turned upside down so that the radicle is forced to commence growth upwards and the plumule to grow downwards, they quickly bend round until they have regained their normal positions; this is shown in Fig. 17, 65. The Maize Seedling —The seedling stage (Fig. 18) of the maize plant is in many respects the most critical in its ex- istence. While it is small it is more seriously affected by the depredations of insect pests (chap. X.), and is more sensitive to fluctuations in the weather conditions. It is obvious that BOTANICAL CHARACTERS 93 if it has only one or two leaves, and these are eaten off CHAP. by a caterpillar, the plant will suffer more than when it has me six leaves, some of which will be able to function and repair the damage done while the others are being devoured. If the root-system only penetrates to a depth of 2 or 3 inches, the young plant is more likely to suffer from a temporary drought than if its roots are tapping. a supply of soil moisture at a greater depth. It is highly desirable, therefore, that the seedling should be encouraged to get beyond this critical Fic. 17.—Maize-grains planted upside down, showing geotropism of the root and apogeotropism of the shoot. stage as quickly as possible; this is one important reason for the use of artificial fertilizers in South Africa (chap. VIIL). After the seedling has developed a root- and leaf-system of its own (Fig. 18), growth is rapid if suitable conditions prevail. Conditions favourable to rapid growth include warmth and moisture of the soil, sunshine, and a suitable tilth of soil to prevent water-logging. At the Geneva (New York, U.S.A.) Experiment Station, growth of the maize plant has been measured and found to range from 3 to 184 inches per week; CHAP. IV. 714 MAIZE a growth of 5 inches has been recorded on one day under specially favourable conditions in Iowa, and in Illinois an increase of growth equal to 1,300 Ibs. of dry matter per acre was recorded in a single week. 66. The Root and Its I'unctions.—The root is the part of the plant which grows downward into the soil for the purpose Fic, 18,—Maize seedlings at two stages of growth, the youngest on the left. of anchorage and absorption of food materials; it does not bear leaves nor reproductive organs. The water held in the soil dissolves the carbonates, nitrates, and other salts which occur in the soil. Dilute solutions of these are drawn in through the minute rvot-hazrs (Fig. 16) on the younger roots, and are carried up into the plant, where they are chemically BOTANICAL CHARACTERS 48 changed into the various compounds on which the plant lives and grows. The plant gets most of the elementary constitu- ents of its food in this way, Maize is a surface-rooting plant. The majority of its permanent roots usually start at about 1 inch below the sur- face of the soil, regardless of the depth of planting (Auwt, 1), The general tendency of the roots is to spread horizon- tally, near the surface, for 1 or 2 feet all round, and then to turn abruptly downward. As a rule the horizontal roots occur within 4 inches of the surface. Hunt (1) found that in young plants one to six weeks old, by far the largest part of the root-system occurred at a depth of 2 to 4 inches from the surface. He concludes that the distribution of the roots is probably dependent more upon a proper supply of oxygen and water than upon temperature of the soil. Maize roots have been measured 8 feet in length (not depth). They have been traced to a depth of 4 feet and slightly over, but as a rule most of the root-mass occurs within the first 2 feet of soil, The following measurements have been recorded :-— Height of plant 4 inch, root 8 inches long. 44 5 3 inches, root 13 inches long oF 5 5 inches, root 11 to 24 inches long. The joints (zodes) at the lower portion of the stem, above the surface of the soil, are often provided with roots, few or many in number, called adventitious or “ brace-roots” (Fig. 23), some of which grow downward till they reach the soil, and then appear to assist in anchoring the plant. These ad- ventitious roots are more plentiful in some of the less improved Tropical American breeds (e.g. Cusco, Mevrican, etc.) than in many of the more highly-bred North American sorts. In some breeds they occur at a considerable distance up the stem ; it seems probable that in such cases the plants are accustomed to grow on river banks, subject to floods which carry a deep deposit of river mud, and into which the “brace-roots” may penetrate. From the above description of the root-system we draw the following conclusions: /77s7, that the maize plant being a sur- face feeder requires that its food supply shall be within a short CHAP. IV. 16 MAIZE ake distance of the surface ; second, that deep cultivation while the Fic. 19.—Part of a transverse section of a maize stem, showing mass of pith, with scattered vascular bundles, more numerous near the stem-wall. (From Cavers’ Practical Bolany, W. B. Clive.) plants are growing is likely to prune off the surface-roots and thus reduce the absorptive power of the plant; ¢hzrd, that cee PROTOPHLOEM SIEVE TUBES COMPANION CELLS cee LOGOS LARGE PITTED os VESSELS SMALL PITTED VESSELS SPIRAL AND = ANNULAR VESSELS XYLEM PARENCHYMA rae LYSIGENOUS CAVITY SCLERENCHYMA Fic. 20.—Transverse section of vascular bundle of maize. (From Cavers’ Practical Botany, W. B. Clive.) surface-rooting weeds interfere with the supply of moisture and plant-food required by the maize crop. BOTANICAL CHARACTERS a3, 67. The Stem and Its Functions.—The stem grows upward CHAP. and bears the leaves and inflorescences (Hig, 12). Tt ishuiltap °% of a series of lengths or ¢zternodes connected by joints or A B Cc Fic. 21.—Part of a radial longitudinal section of stem of maize, showing one of the vascular bundles. A and G, parenchyma of the ground tissue; B and F, sclerenchyma ; C, phloem; D, pitted vessels of the xylem; E, spiral and annular vessels. (From Cavers’ Practical Botany, W. B. Clive.) nodes. In many grasses it is hollow, but in the maize plant it is filled with pct, which consists largely of cellulose. The hard outer portion or stem-wall contains numbers of fibres called vascular bundles (Fig. 19), through the woody part Fic. 22.—Part of a tangential longitudinal section of stem of maize, show- ing one of the vascular bundles. Aand E, parenchyma (ground tissue); B and D, sclerenchyma; C, xylem—note the large pitted vessel on either side, and the small pitted vessels in the middle. (From Cavers’ Practical Botany, W. B. Clive.) (xylem) of which the solution of water and food-material ab- sorbed by the roots travels upward to the leaves. As in other grasses the stem-wall is well supplied with silica, which aids CHAP. IV. 78 MAIZE in maintaining it in an erect position. Because they differ from the stems of other families of plants, the stems of grasses are usually spoken of as cus. The stem or culm of the maize plant is extremely variable in height, ranging from 14 to 30 feet in different breeds and in different climates ; the usual range in South Africa is from about 5 to 14 feet. At the Botanical Experiment Station, Pretoria, in 1907-8, the maximum height! was 6 feet 6 inches (2 plants), and the minimum 3 feet, while the average of 313 plants was 5 feet 4 inches. The circumference of an average stem ranges from about 3 to 44 inches between the first and second nodes, in a dent or flint maize. The internodes are channelled on alternate sides, next the leaf blade, and on the side where the branch or ear may occur, The histology of the maize stem has been studied by Dr. Cavers (1), whose illustrations of the tissues are reproduced here (Figs. 19 to 22), by kind permission. Figs, 19 and 20 show transverse sections through one of the lower internodes, and 21 and 22 longitudinal sections. The function of the stem is to carry food materials from the roots to the leaves, and then to other parts of the plant, and to raise the leaves and inflorescences into the air and light, to facilitate photosynthesis (4 69) and pollination (4 73). 68. Sucker-shoots—The maize plant often produces sucker- shoots (Fig. 23), especially when planted thinly; these are branches which arise from the lower nodes near or below the surface of the soil. Some breeds have a much greater tendency to sucker than others; the tendency appears to be specially strong in cross-bred plants. These suckers often produce . tassels bearing both male and female flowers (Fig. 24), which sometimes develop small, round grains. As a rule suckers do not bear proper ears; they are, therefore, undesirable in crops grown for grain, because they take food and moisture from the soil and give no return except the fodder. Some breeds bear several branches from nodes higher up the stem, but as a rule the stems are unbranched except for the suckers and the shanks of the ears, for the shanks are in reality only short branches. 1 Of a cross between Iowa Silver-mine and Wisconsin white dent. 79 BOTANICAL CHARACTERS CHAP. IV. ‘s}OOYs-1ay9NS puk $}OOI SNONHuUaape Sulmoys Gueyd azieur Sunod yo aseg—Ez “org CHAP. IV. 80 MAIZE 69. The Leaf and Its Functions —The leaves are borne on the stem around which the sheath (Figs. 12c and 25), or lower portion of the leaf, is closely wrapped ; the broad upper portion which spreads away from the stem is called the d/ade (Figs. 12B and 26c). At the junction of sheath and blade there is a pro- jection which clasps the stem, and which is called the /eule (Figs. 25 and 26). To a certain extent the ligule of the maize plant checks the entrance of water and soil, which might start decay between the sheath and the stem. Fic. 24.—Tassels of sucker-shoots, bearing small round grains as well as the normal male spikelets. The leaf blade of the maize plant is long, broad, and flat, with wavy margins; the mid-rib is broad and strong (Fig. 5). The surface is usually more or less rough with scattered, adpressed white hairs (Fig. 25), which may be short or longer. The cuticle is found by Wager (1) to form a thin layer on both surfaces, with practically no difference in thick- ness in the different breeds. In the dent variety of maize the number of leaves on a stem usually varies from 12 to 18, but as the lower leaves die off before maturity, it may happen that only about 12 function BOTANICAL CHARACTERS 81 at one time; 15 is usually considered a desirable number. The average number for 313 plants’ at the Botanical Experi- ment Station, Pretoria, in 1907-8 was 10°3, the maximum being 14 (on 3 plants only) and the minimum 6 (on 2 plants only). Leaf measurements were made of these 313 plants, the sixth from the base was selected as the most fully de- Fic. 25.—Leaf sheath and base of blade, showing ligules, and hairs on blade. veloped ; the maximum width was 5:25 inches, the minimum 2°5 inches, and the average 3°8 inches. The total external leaf-surface of a single maize plant has been measured, at the Missouri State Experiment Station, and found to equal 24 square feet. An acre often carries over 1Of a cross between Iowa Silver-mine and Wisconsin white dent. 6 CHAP. EN: 82 MAIZE CHAP. 8,500 maize plants, which, with 24 feet of leaf-surface each, would yield a total of 204,000 square feet, or 4°68 times the area of the soil covered by the crop. IV. B, A sheath sually long auricles. C, Blade removed, showing ligule at A, Sheath wrapped round stem, showing unu removed and opened, showing shorter auricles, with ligule between them. Fic. 26.—Leaf sheath. base (stem end). The following method of measuring the leaf-surface of maize has been recommended by some of the Agricultural Experiment Stations in the United States :— Select an average-sized leaf. Take the width at 3 inches from the ligule, BOTANICAL CHARACTERS 53 Take the width at 6 inches from the tip of the leaf. Take the average of these two measurements and multiply by the length of leaf between the two points of measurement. Add the area of the isosceles triangle formed by the 6 inches left at the tip. Multiply by 2 for the two surfaces. Multiply by the number of leaves on the plant. The leaves absorb air into their tissues through microscopic openings called stomata (Figs. 14 and 27). At the suggestion of the writer, Mr. H. A. Wager (1), of the Transvaal University College, Pretoria, kindly undertook to determine the number of stomata on the maize leaf. He found that on the under surface they varied from 75 to 126 per square millimetre, and on the upper surface from 60 to 97. Carbon-dioxide (CO,) is one of the gases of which the air is composed, and consists of the two chemical elements carbon and oxygen. When the air comes in contact with the chloroplasts ({1 61) in the leaf- cells, in the presence of light and moderate warmth, the carbon-dioxide is decomposed, and some of the oxygen is given off into the air. The carbon is retained, and, com- Fic. 27.—Three stomata with sur- bining with the water and_ rounding epidermic cells (E). G, G, chemical substances obtained furl of some, Tom Si F from the soil through the roots, bridge University Press). ; various complex organic com- pounds are formed. This chemical action takes place chiefly during the day-time, and only in the presence of light, and is therefore called photosynthesis. The new compounds are used in the building up of tissues required for the increasing growth of the plant. All of that enormous quantity of starch required to fill out the endosperm of the maize grain must first be chemically formed in the leaf before it is carried to the grain on the ear, where it is finally deposited. The importance of the leaf in the life-history of the plant is thus evident; 7¢ 7s a chemical laboratory tn which the various elements of plant-food are separ- ated out from the compounds in which they originally occur, and are re-united into such forms as can be made use of by the plant. 6 * CHAP. IV. CHAP. IV. 84 MAIZE Maize plants poor in leaf-surface, through lack of food or water, or from insect-injury, damage by hail, or undue short- ness of growing season, cannot manufacture and store as much starch, and therefore fail to develop as much weight of grain as those with a proper proportion of leaf. It does not neces- sarily follow, however, that the larger the leaf-surface the Fic. 28.—Tassel of Odessa maize. greater the amount of seed produced. Beyond a certain potnt (which perhaps varies in different varieties or breeds), the amount of seed produced appears to decrease in inverse ratio to the increase in leaf-surface. After the food-materials are used up, the surplus water in which they were dissolved and carried is transpired or given BOTANICAL CHARACTERS 85 off from the leaf-surface, and is replaced by a fresh supply CHAP drawn up from the roots. Frank (1) notes that a single 1! maize plant can pass off into the atmosphere 31 Ibs. weight of water in 147 days of growth. This is less than the amount lost by some other plants, e.g. :— The maize plant can lose 31 Ibs. in 147 days. The hemp . 60 ,, in 140 The sunflower __,, 147 ,, in 140 ” ” The rate of transpiration is greater in light than in dark- ness. Inthe maize plant it has been found to vary | in the following ratio :— In darkness . * 4Q7. | \ In diffused light . 114 \\ In direct sunshine 785 | There is thus a stream aa of water constantly pas- \\\ sing away from the soil \\\ HI into the air, through the } \ : | / / leaves of the plant; it soon drains the soil dry \\\ \ \ We, unless replaced by rain : \\ | Hl orirrigation. It has been \\ found that 500 tons of \\ RS) water (containing salts ae AW | in solution) is required ~~ S\X_ to form one ton of dry ~ . ges S¥ matter (Bowman and ee ey." Crossley, 1). \\ HN The leaves are ar- AN \ wt ranged dzstechously, i.e. in two opposite rows, but there is no uni- formity as to the direc- tion in which the rows Fic. 29.—Young ear showing silks ready point. for first pollination; the husks have developed 70. The Inflorescence. leaf blades, showing the homology of husk and . leaf sheath. —The maize plant is monecious, i.e. it bears the reproductive organs in separate flowers on the same plant (Fig. 12). The staminate or male CHAP. IV. 86 MAIZE flowers are usually borne by themselves in the terminal inflores- cence called the tassel (Fig. 28). The pésél/late or female flowers are usually borne on one or more lateral inflorescences (Fig. 12b) called ears, at the ends of short branches arising from the nodes of the stem; the young ear is characterized by its long beard-like styles called sz/ks (Figs. 29, 30, 31, and 32). Moncecism is not always complete, i.e. the separation of male and female flowers into different inflorescences on different Fic. 30.—Young ears showing silks at different stages of development. parts of the same plant is not constant; bisexual tassels are frequently met with on sucker-shoots (in the dent breeds at least) (Figs. 33 and 34); in some cases also, the tassel of the main stem bears the two kinds of flowers (Fig. 35); this is said to be especially the case in pod maize, a breed rarely seen in South Africa. Bisexual ears are occasionally, though rarely, met with (Figs. 36 and 37). Where mixed inflorescences occur, stamens and pistils may be found in the same spikelet, BOTANICAL CHARACTERS 87 and the writer has even found hermaphrodite florets (i.e. with stamens and pistils in the same floret), but in those cases one or other organ was abortive. Moncecism in maize facilitates cross-pollination and hybri- dization, though the only other species with which maize can hybridize, as far as we are aware, is the Mexican Teosinte, Fic. 31.—Young ear with fully developed silk. Euchlena mexicana (Fig. 10), of which hybrids have been ob- tained by Harshberger. 71. Barren Plants.—One of the causes of low yield of grain is the occurrence of barren plants, i.e. plants which though bearing tassels produce no ears, or ears without grain. Cases are reported from America of 60 per cent of barren CHAP. IV. 88 MAIZE CHAP. stems in a crop of maize. This subject has been much dis- IV. cussed by maize breeders, but it is still a moot point whether the tendency to produce barren stems is an inherited char- acter. Some writers, e.g. Hunt (1), maintain that ‘if it were an hereditary characteristic the fact that the stalks are barren would tend to eliminate them”. If barren stems were adso- Fic. 33.—Bisexual tassel of a sucker-shoot. Fic. 32.—Young ear with husks removed to show silks. Note that the silks at the tip of the ear are not yet fully grown. futely barren, in the strict sense of the word, this would be true, but the fact that they produce tassels and pollen lends colour to the view that they may tend to reproduce their kind. The percentage of barren stems on a given soil is said 89 BOTANICAL CHARACTERS CHAP. 2 ‘aures Jo youeig ‘gq *pasiejua *yooys-sJayons *w9}8 UTUI UO Jasse} [eNxasIq—'SE ‘917 jo (srea pews Surreaq) jesse} yenxasig—PE “ory LZ CHAP. IV. go MAIZE (Hunt, 1) to vary with the thickness of planting and the season, and barrenness does not seem to be a variety char- acteristic, but rather the result of environment. The subject needs further investigation as a possible cause of low yields in South Africa. Sometimes the maize plant bears ears which are barren Fic. 36.—Bisexual ear. Fic. 37.—Bisexual ear. owing either to the destruction of the silk by the larve of certain moths (Fig. 38) or to failure to produce silks even when the ovules are developed (Fig. 39). 72, Flowering Period.—Considerable difference is notice- able in the time taken by different breeds to reach the flower- ing stage; it usually requires nine to twelve weeks, This is BOTANICAL CHARACTERS gI an important point for the farmer. A breed which répens off CHAP. too quickly after flowering will not have time to form and !Y- store enough starch to produce a heavy crop of grain; but if the shortening of the time of growth takes place between germination and flowering, it may, perhaps, not have such an effect on the yield. The shortest period between germination and flowering noted by the writer has been sixty-four days. Variation in this respect may be due in part to difference in Fic, 38.—Silks injured by larvae. amount and time of rainfall. If the grain is planted early in the season germination and growth are retarded because the temperature of the soil is not sufficiently high. In South Africa, drought in December and January sometimes checks vegetative growth and hastens flowering. But soil-moisture and temperature will not account for all of the differences noticed. Within the same breed the flowering period of individuals varies greatly ; the extremes noticed are fourteen and twenty- CHAP. IV. 92 MAIZE one days. This is partly due to variation in the character or composition of the soil within a few feet of surface area; partly, perhaps, to variation in vigour between the individual plants; and perhaps largely to the occurrence of different Fic. 39.—Blind ear, on which the silks have failed to develop. strains in an ordinary commercial breed; a well-bred strain may be expected to show greater uniformity in time of flowering. Yet there seems to be a definite relation between the time of flowering and the time of ripening. BOTANICAL CHARACTERS 93 TABLE XVI, VARIATION IN TIME OF FLOWERING. al | First Flowers. | Last Flowers. | Breed. Date Sown. | = om 7 | Appeared. Days. Appeared. Days Later. | | ee ee | | Arcadia Sugar . . | 20 Aug., tga g Nov. 81 | Dec. 2! 23 Louisiana . E . | 27 Sept., 1910 | 8 Dec. 73 Dec. 25 17 | Black Sugar . ; ra | 30 Nov. 64 | | Hickory King . . | 15 Oct., gE | 30 Dec. | 77 | | | Yellow Dent a ie 27 Dec. 74 | King of the Earliest? . g Dec., 1907 | 11 Feb. 64 White-cap Dent” | ” | - | 64 | | Yellow Hogan 2 | | me | 64 | | Wealth of Nations? . | 8 12 Feb. 65 | | Hawkesbury Champion” rf | ee | 65 Hundred-day Bristol? | fy | 14 Feb. 67 | Skinner’s Court ro. . | 3 Oct., 1907 17 Dec. | 75 | 1 Ears were ready for boiling on rr December, and these were not from the earliest-Alowering plants. 2 Grown without irrigation ; Skinner's Court 10 was irrigated. An investigation conducted at the Botanical Experiment Station, Pretoria, in which careful records were obtained from thirty-eight different breeds or strains of maize (see Table XVIJ), resulted as follows :— Aggregate time taken from appearance above ground to period of flowering . . 2,125 days. Aggregate time taken from date ee open nee above ground to ripening of seed. : : ‘ . 4,256 days. From the figures in Table XVII we conclude that the period of flowering occurs, roughly speaking, half-way between the first appearance of the plant above ground and the ripen- ing of the seed. Hence a farmer should be able to foretell approximately the date at which he may reasonably expect his seed to be ripe and safe from frost. It is possible, how- ever, that results obtained in one district may not be repeated in another, and it is therefore desirable to obtain records of the results obtained by farmers in different parts of the country. CHAP, IV. 94 MAIZE TaBLe XVII. CORRELATION OF FLOWERING AND RIPENING PERIOD. Corrected for Date of Days after Sowing. Appearance above the | Ground. Breed. | " ess Flowering. Ripening. Flowering. Ripening. Brewer Yellow Dent . ‘ 61 112 53, 104 » noo : 63 115 57 10g Cuban Giant , y , 61 | 110 53 102 " é ; ‘ 61 II5 54 108 Snowflake . 3 : : 60 IIo 52 102 4 A . . : 58 10g 52 103 King Philip (flint) . - | 67 To8 59 100 o Ni 2 : 59 113 54 108 Queen of the Prairie . ‘ 61 IIo | 53 102 a mo -| 59 130] 853 107 Early Mastodon . : ; 63 122 54 113 ae s « > 62 125 57 120 Sanford ‘ ‘ : 60 113 52 105 ” . : : : a2 m5 53 10g Compton Early . . - | 59 | 108 51 100 ” : : - | 59 113 53 107 Improved Early Canada Pll 56 I05 50 99 Wills Gehu (flint) : : 54 87 49 82 Wills Dakota (flint). , 54 87 49 82 Champion White Pearl , 67 II7 59 109 King of the Earliest . : 66 115 58 107 a ae ; if 58 113 2 107 Yellow Horsetooth . ¥ 72 123 63 11g Chester County . : ; 67 128 60 121 4s P F é 58 II5 53 110 Eureka . : F ; 65 128 59 122 New England 8-row . : 67 128 60 121 Yellow Botman . : a 65 125 60 120 Hickory King. : | 66 | 128 60 122 Boone County. : st 65 | 128 60 123 4 r . sil 64 128 | 59 123 Pride of the North. ; 58 113 2 107 Sheepstooth : 67 128 61 122 Thoroughbred w.f. 62 12 57 120 Wisconsin w.d. | 62 128 | 57 123 Golden Dent | 65 | 128 59 122 Iowa Silver-mine : | 205 128 59 122 White Flint . y «| 67 128 61 122 White Cap y.d. . . : 59 IIs 53 109 Tuscarora . F : : 59 113 53 107 Red Cob Fodder . ‘ é 75 | 131 ‘ | 2,555 | 4,808 2,223 4420 Deduct Wills Gehu and | Wills Dakota 108 174 98 164 25447 | 4,634 2,125 4256 BOTANICAL CHARACTERS 95 The only exceptions are the two earliest flint breeds, and here the figures would seem to indicate that earliness is pro- duced by the hastening of the period between flowering and ripening. 73. The Spikelet. — The tassel consists of numerous branches (Fig. 40) bearing more or less dstéchous rows of staminate spkelets (Fig. 40A) which are MY arranged in pairs, fe one pedicellate Mi (stalked), the other We : : ny; sessile (without a mi stalk). Each spike- Pi let consists of a pe pair of protective, ia $ sheath-like bracts, ia 5 (i called glumes or “empty glumes” "ya MM (Fig. 41d), which SX y enclose two florets. Each floret consists of a pair of thin bracts, the lower or outer of which is called the valve (called also “ flower- ing glume” by some botanists), and the upper or inner the patea. Enclosed within the valve and palea of each floret, when in bud, are three stamens (Figs. 40 and 41); there are thus six stamens to each spikelet. When the flowers open, the stamens are exrserted (pushed out). Each stamen is composed of a long and narrow anther hanging freely at the end of a fine, slender thread, the fi/ament. Each anther consists of two sacs, attached side by side, and having an opening, a fore, at the lower end, for the emission of the minute, dust-like, QS WY, NN f fF Fic. 40.—Tassel of sugar maize, with male spikelet and stamens enlarged CHAP. IV. CHAP. IV. Fic. 41.—Spikelets of the maize plant (much enlarged). A, male spike- let. B, female spikelet. a, anther; b, pore; c, filament; d and e, glumes; /, ovule. (After De Vries, from models by Brendel, Berlin.) Fic. 42,—Pollen grains of maize (much enlarged). A, normal shape of living ., grains; B, shrunken, ‘‘dead” grains; C, dead grains moistened and allowed to dry again. BOTANICAL CHARACTERS 97 yellowish follen (Fig. 42), which is the medium of sexual re- CHAT. production. At the base of the anthers are minute organs, 'Y- called /odzcules, which at the time of flowering become turgid and press open the valve and palea, allowing the stamens to protrude and scatter their pollen. Naf A Fic. 43.—Effect of complete or partial lack of pollination. A, Barren ear due to protection from access of pollen. B, Two ovules were fertilized, probably by pollen which caught on the husks before the bag was placed on the ear. Both glumes and anthers vary in colour from pale green, through cream, salmon, etc., to deep magenta. Often the glumes are striped longitudinally with magenta or pink. 74. The Pollen and Its Vitality.—Pollen is the fine cream- coloured or golden dust which may be seen flying in clouds from the tassels when they are shaken on a still morning. It 7 CHAP. IV. 98 MAIZE has a peculiar, heavy, sweet, lasting odour. Pollen is essential to the fertilization of the pistillate flowers and consequently to the development of the grain; without it no grain can be formed. This may easily be demonstrated by tying a clean paper bag tightly over a maize ear, before the silks develop, ee C3 ¢g tens F es Cras. iv Begg me on a paces v= Fic. 44.—Effect of partial pollination. A, Pollen was applied artificially > after most of the silks had ceased to be receptive. b, Imperfect pollination, possibly due to temporary injury to silks by insects. and leaving it so covered until after the silks have dried up (Figs. 43 and 44). Pollen “dust” is composed of an enormous number of roundish grains (Fig. 42). Each individual grain is a separate cell, consisting of a cell-wall of usually two layers or coats, surrounding a mass! of protoplasm; within the cell are two > BOTANICAL CHARACTERS 99 smaller bodies called cle’, one being known as the vegetative nucleus and the other as the generative nucleus. The writer has counted 636 spikelets on a tassel of sugar maize, containing in all 3,816 stamens, while the more robust “field corns” (i.e. dent and flint varieties) may carry about 200 stamens. The pollen grains produced in one anther Fic. 45.—Young ear showing homology of husks. have been counted and found to average about 2,500 each. At this proportion the tassel of sugar maize referred to would contain 9,500,000 grains of pollen. It has been estimated that a tassel of field maize will produce 18,000,000 grains, and especially vigorous plants from 30,000,000 to 60,000,000. An average ear of sugar maize produces from 250 to 350 grains of corn; Afvckory King about 400; some of the more ie CHAP. IV. MAIZE 100 CHAP. IV. (“silks”). B, Ovaries and silks enlarged. Fic. 46.—A, Young ear showing ovaries and styles BOTANICAL CHARACTERS 1ot productive breeds from 1,100 to 2,800. Allowing for a pro- duction of 1,000 ovules ({75) requiring pollination, and a minimum of 6,000,000 grains of pollen available, we have 6,000 grains of pollen for every grain of corn. As only one is actually required, there is abundance to spare for the bees and chafers, and for waste. The amount of waste must be enormous, yet we often find large numbers of cars incompletely 1h oe ae e 0@ 4}; 6: aco <6) Fic. 47.—Two-lobed ear. Fic, 48.—Bifid ear. fertilized, probably through having been receptive at a time when little pollen was available in their vicinity. Maize pollen retains its vitality for seven or eight days in the Eastern United States ; in the dry climate of South Africa it keeps well for three days, but after five days most of it is no longer viable. 75. The Young Ear.—The ear is situate at the end of a much shortened branch, which develops leaf sheaths forming the Ausk of the ear (Fig. 45). It is composed of a more or CHAP. io2 MAIZE CHAP. less cylindrical or tapering core, the cob, bearing from 4 to 48 IV, rows of immature grains or carpels (Fig. 46). Though the maize ear has a solid core, it is in reality made up of two or more connate, two-rowed sfzkes which have grown together, or failed to separate, during their early de- Fic. 49.—Branched ear of Hickory Fic. 50.—Branched ear of King. Ladysmith, velopment. Each spike bears at the nodes two two-flowered spikelets, as on the tassel, but the lower floret of each spikelet is abortive, leaving only a pair of carpels to develop at each node; this accounts for the uniformity in the development of rows of grain in pairs. Lobed and branched ears are _fre- quently met with (Figs. 47, 48, 49, and 50). BOTANICAL CHARACTERS 103 The carpel consists of a roundish body called the ovary, and a long, slender, soft thread called the s¢y/e or “ silk” (Fig. 46). Each ovary contains a minute egg, called the ove, which, on fertilization by the nucleus of the pollen-grain, develops into a seed, The cob does not complete its growth lengthwise before the first silks are ready for pol- lination. If the growth of the plant is checked through lack of sufficient moisture or plant- food, or inclemency of the season, the tip of the ear fails to grow out, and the cob, in- stead of being almost cylindri- cal, may remain tapering in shape through lack of develop- ment. In “pod maize” (variety tunicata) the glumes are large, completely enclosing the ovary and persisting around the ripe grain (Fig. 51). In the varieties usually under cultivation, how- ever, the glumes, valve, and palea (173) of the female flower cease to function, and are reduced to small rudiments around the base of the carpel. The position of the ear on the stem varies greatly in in- dividual plants according to the particular node from which the shank (81) is developed, and according to the length of the several internodes below it. Some positions are more desir- able than others (see chap. V.). Fic, 51.—Pod maize (Zea Mays var. tunicata). CHAP. IV. CHAP. IV. io4 MAIZE 76. The Silk.—The style or “silk” is a long, terete, hollow tube, bifid at the end (Fig. 524) and it contains a viscous sub- stance. When receptive, the silk is exserted from the apex of the ear to receive the pollen, and may then become 6 to 12 or more inches in length. After pollination the silk dries up, but persists. If pollination is prevented or is incomplete, the silk continues to grow to an unusual length, and remains green much longer than otherwise. Under a lens the silk is seen to be covered with short, branched hairs, on which the pollen grains are caught. Fic. 52.—The style or silk (much magnified). A, Showing bifid end, and pollen grains caught among the hairs. B, Hairs magnified, showing cells and nuclei. The silks do not all mature at the same time; those at the base of the ear develop first (Fig. 32); then those from a little higher up, and finally those from the tip. It takes about a week for all the silks on an ear to mature (Figs. 30 and 31). This progressive development appears to be one of nature’s ways of ensuring the pollination of at least some of the ovules. It sometimes happens that there is not sufficient pollen avail- able at the appearance of the earliest or latest silks, which results in the production of incompletely filled ears or ‘nub- BOTANICAL CHARACTERS 105 bins”; but such a shortage of pollen is not likely to occur CHAP. throughout the whole period of development of the silks. Ns If the silk be injured, the proper fertilization of the ovule may be prevented, with consequent reduction in yield of seed. 77. Pollination.— Unless a pollen-grain reaches the style no seed will be formed (Fig. 43). The maize plant is anemophilous, i.e. wind-pollinated. The pollen is very light and is carried for long distances by the wind; there is thus danger of cross-pollination if two sorts of maize are planted near to each other—4oo yards is considered a safe distance though some writers recommend half a mile. Because it is so light, and easily carried by a breath of air, very little pollen usually reaches the silks of the same plant which produces it; it is carried by the least puff of wind to the plants beyond. The writer has observed cases in which the pollen fell directly on to the silks below, on a very still evening; but such cases appear to be rare in nature; and as a rule the arrangement of the leaves is such that they partially protect the silks from such a contingency. The maize tassel is much visited by bees, which collect the heavy-scented pollen for food for their young larvae. But the bees do not visit the silks, and are not, therefore, direct agents in pollination, and the amount of pollen produced is so great that the little taken by the bees is not likely to have any effect on the crop of grain. In South Africa at least three sorts of beetle also visit the tassels, to feed on the pollen (see chap. X.), without affecting the yield. An experiment was conducted at the Botanical Experiment Station, Pretoria, to determine the receptivity of silks to pollen at different stages of development, with the following result: Application of pollen on the first day of appearance of the silk resulted in the fertilization of only 14 ovules, situate toward the base of the ear. Of the ear pollinated the second day, about half of the ovules were fertilized, the upper part of the ear being bare of grains. The ear pollinated on the third day was three-fourths covered with grain, the extreme butt and the tip having no grains. From this we conclude that if pollen is not available by the third day, the earliest silks to develop may not be fertilized. 78. Fertiltzation.—When a pollen grain is caught among CHAP. IV. 106 MAIZE the stigmatic hairs of the silk or s¢y/e, it begins to grow. The pollen grain takes up moisture from the stigma and begins to swell; a tube, called the pollen tube, pushes out from one side of the pollen grain, penetrates the silk, and grows down its whole length till it reaches the ovary at the base (Fig. 53). Fic. 53.—Diagram showing course of pollen tube through style to ovule. A, Section near outer end of style, showing pollen grain (a) and pollen tube (g). B, Section through base of silk and through ovule (e). (After drawing by C. S. Ridgway, in Duggar’s Southern Field Crops, New York, The Macmillan Co.) Into this tube the contents of the pollen grain, including the nuclei, pass. During growth the vegetative nucleus becomes gradually disorganized and is lost in the protoplasm. The generative nucleus, however, has divided and formed two nuclei. On reaching the ovary, the pollen-tube enters the embryo-sac, and discharges its two nuclei; one of these fuses BOTANICAL CHARACTERS 107 with the nucleus of the egg-cell to form the embryo (Fig. 54); fertilization is then accomplished. The other male nucleus fuses with the two fused female polar-nuclei; from these the endosperm is developed. Cell-growth and cell-division then commence, and are continued until the mature seed js developed. The chromosomes of maize are small and difficult to study, and scarcely anything is known of their behaviour during the maturation division (Zas¢, 6), 79. Dichogamy.—According to Kerner (1 ), Most moncecious plants, including maize, are profo- gynous, i.e. the female flowers are receptive before the pollen of the p same plant is shed, thus necessitat- ing cross-pollination. If protogyny were complete, the very first plant in a field, which developed female flowers, would, in consequence, fail to propagate its kind, unless accidentally pollinated from a still earlier plant in a neighbouring field ; but there would always be one plant in a district that was Fic. 54.—The embryo-sac in earliest of all, and which would maize at the time of fertilization. < ; Pt., pollen tube which has just dis- therefore fail to develop grain. Sie che tin wale gues This habit would act to some g , egg-cell which, after union with extent as a check to any natural one of the male nuclei, forms the embryo; end., nucleus of the en- ; dosperm, with which the second maturity. male nucleus may unite. (After If all the plants in a field drawing by F. E. Lloyd in Dug- flowered on exactly the same cee ees Sores day, and all were completely protogynous, there would be no pollination except from other and earlier-planted fields, and those of the first planting would always fail to develop grain. But this is not the case. As already pointed out (| 72), there is a great difference in time of flowering with individuals in the same field, due to many causes, e.g. individual characteristics, difference in depth of planting, variation in soil fertility, soil texture and soil moist- ure, etc. The flowering period in any one field or plot may thus extend over ten days or three weeks. end. tendency to increased earliness in y CHAP. IV. CHAP. 1V. 108 MAIZE Protandry is the opposite of protogyny, i.e. the anthers shed all or part of their pollen before the female organs of the same plant are receptive. The very earliest pollen shed will, in such a case, be entirely wasted unless there is a protogyn- ous plant in the vicinity which is receptive at the same time. Plants of Black Sugar-maise in Pretoria, which matured pollen on 14 December, 1907, did not have any receptive stig- mas till the 16th, 17th, and even later. In some instances the tassels had shed a// their pollen before any silk appeared. As far as the writer has been able to determine, from observation of Transvaal maize fields, protandry is the rule in South African maize. And Shamel (1) states that in America “in most varieties the pollen matures before the silks”. On this account well-filled butts are more frequent than well-filled tips. At the Botanical Experiment Station, Pretoria, no case of protogyny was observed among seventeen plants under obser- vation for time of relative maturity of pollen and silk. See : =i Number of Relative Time of Appearance of Silk. Plants. 3 days later than the pollen 3 2 2 2 oy ” ” 2 Is ” ” ” 4 Same day as the pollen 8 In another experiment, however, with another variety of maize (a yellow flint, IV7e//s Gehu), every one of twenty-six plants under observation was protogynous. Plants of Arcadia Sugar-matze which developed silks on g November, 1910, had no anthers exserted until the 11th, while others had no anthers till the 13th. In some cases the tassels appear long before the anthers; the writer has a note of one vigorous plant of Loweszana Hickory on which the tassel first appeared on 25 November, but there were no anthers until 8 December, i.e. thirteen days later, the silks appearing on the same day. It seems probable that protogyny is a breed characteristic. It appears to be constant in Arcadia Sugar-matze and in IWells Gehu (yellow flint), while protandry is the rule in BOTANICAL CHARACTERS 109 Black Sugar, Hickory King, Louisiana Hickory, and many other dent breeds. This matter is more fully discussed else- where (Burtt-Davy, 25). 80. Form for Describing the Maize Plant in the Field.— The following is a convenient form for use in recording the vegetative characters of selected plants in the field or the breeding plot. It can be printed off on to record cards of uniform size for subsequent filing. By marking off the char- acter which is present, much time is saved. which would other- wise be spent in writing out the cards. For instance in (0), if the plant is in silk at the time of taking the notes, put a mark, thus /, over the word “silking,” or, if it is ripe, then over the word “ripe”. More space must be allowed for the writing than is here indicated :— (a) Namerot Breed). iieioi tick cain sarwes ada nemeeanteseses Date of record......... (BD): WIRES: BTOWT ecg. ce cece oms cuvadaigus nenaiwane aatanirene Date of planting......... (c) Maturity of plant; silking; roasting ear; partly dented or glazed; fully dented or glazed; nearly ripe; ripe. (d) Height of stem: average of ten plants......... feet caikis inches. (e) Stem; straight; medium; zigzag. (f) Stem circumference at middle of internode between second and third node from ground......... inches. g) Stem circumference at middle of internode below main ear......... inches. h) Number of ears on roo stems......... i) Number of barren stems in too plants......... J) Position of ear; in middle of stem; above the middle; below the middle. k) Direction of ear; pointing upwards: horizontal; pointing downwards. 1) Length of shank; distance from node to base of ear, avcrage of ten plants........ inches. (m) Husks; abundant; medium; scanty. (x) Husks; tight ; medium; loose. (0) Number of leaves; average of ten plants......... ( ( (#) ( (7 ( ( p) Width of leaf-blades on ten plants: maximum......... inches; minimum en inches; average.........inches. (q) Length of leaf-blades on ten plants: maximum......... inches; minimum oasis inches; average.....,...inches. (ry) Length of tassel; average of ten plants......... inches, (s) Number of branches of tassel; average of ten plants......... (er Additional Notesy co.cc daradewiecavs ncqsssasdienen danlanweintnasasarac 3 arenes anew 81. develops. It is a much-reduced lateral branch, arising from a node on the main stem, and, like the stem, having nodes and internodes, the latter much shortened. Each node of the branch gives rise to a leaf, and in some cases to one or more distinct secondary ears (Fig. 55); this may produce silks, but CHAP. IV. CHAP. IV. ILO MAIZE appears rarely to develop grain; occasionally, however, fully developed secondary ears are produced (Fig. 56). A variation of one to twelve inches in length of shank has been noted. Very long shanks are undesirable because they allow the ear to hang too far away from the stem, which often results in its breaking off prematurely. Fic. 55.—Four secondary ears developed from the nodes on the shank of a single ear. The diameter of the shank also varies ; it may be ‘‘/arge,” i.e. nearly the diameter of the cob; “ medvuz2,” i.e. half the diameter of the cob; or “ swa//,” i.e. one-third the diameter of the cob. 82. The Husk.—Vhe husk of the ear consists of the leaf- sheaths which arise from the nodes of the shank (Fig. 39); BOTANICAL CHARACTERS Itt their homology is clearly indicated by the fact that often a diminutive leaf-blade, varying greatly in length, is developed at the end of each sheath of the husk (Fig. 45). Sometimes the husks are tightly wrapped around the ear; in other cases they are loose and baggy, giving a deceptive appearance of size to what may be only a small or medium ear. In some cases the husks entirely cover the ear, and even extend a long way beyond it; in other cases they may be so short that the tip of the ear protrudes beyond them; the latter is an undesirable char- acter, as it allows birds and insects to damage a good deal of the grain at the tip of the ear (Fig. 164). 83. The Mature Ear.— The number of ears which a maize plant can bear varies considerably with the variety and breed, and also varies according to the richness or poverty of the soil and length of growing season. The tendency to produce more than one fully developed ear on a plant (Fig. 57) may be seen in any maize field. But it is not the usual thing, in South Africa at least, for more than one good ear (on a dent breed) to develop fully and to mature grain. Some varieties naturally develop more ears; eg. cases of ten or more ears on a flint breed have been reported, and one plant has been known to produce twenty-three ears; pop- corn has been known to produce twelve to nineteen ears; and six to fourteen have been reported for dent maize. But Hunt (1) points out that in the United States, dent breeds Fic. 56.—Secondary ear developing from a node of the shank. CHAP. IV. CHAP. IV. 112 MAIZE produce but one ear under ordinary conditions of culture; ‘no two-eared dent breed has ever been produced which has become extensively grown or widely popular”. It seems reason- able that it should be easier for a plant to develop one large ear than two or more small ones, and it is better from the farmer's point of view, for it costs less to harvest one good Fic. 57.-—Maize plants developing two ears. ear than to pick two small ones from each plant. The mature ear consists of a central pithy core, called the cod (Fig. 43) on which the grains are borne. The ear varies in length from 4 an inch to 16 inches, but 4 to 9 inches is a usual range in early and medium-maturing sorts, and 9 to 14 inches in the later sorts, such as white Horsetooth. Occasionally we have an BOTANICAL CHARACTERS Lr3 early maturing sort which develops a long ear, e.g. Chester County. The circumference of a mature ear at 2 inches from the butt should average 3 inches in an ear 4 inches long, or 74 inches in one 10 inches long. The weight at harvest may vary from 3 to 18 ounces or more, but in time it may lose in drying from one-half of 1 per cent up to 35 per cent. In shape the ear may be cylzndrical (of uniform circum- ference) throughout its entire length, or more or less ¢aperzng. The dutt or base varies in shape and size. In a normal ear it should be of the same diameter, and have the same number of rows of grain as the main part of the ear, but this is often not the case. If the entire end of the cod is exposed, with the butt-grains at right angles to the axis of the cob, the butt is described as even. It may be rounded at the end and show the marks of the tightly-clasping husks on the grain, when it is called compressed. If there is a greater space be- tween the rows at the butt than on the rest of the ear, it is open. It may be expanded by additional rows of grain, or enlarged without having any extra rows. The cavity formed by a rounded butt may be shallow and broad, of moderate depth and diameter, or deep and of small diameter. The apex of the ear is called the #. The tip may be en- tirely covered with grains ; it is then described as fi//ed¢. The tip grains may be scattered or in rows, or the tip may be bare through exposure, from lack of pollination or of adequate covering by the husks, or from ravages of ear-worm or birds, or through drought or lack of plant-food. Ifa central grain projects from a filled tip it is called capped. In shape the tip may be rounded or flattened. The spaces between rows are called su/cr, The smaller the number of rows, the greater the tendency to width of sulci. But if the grain is well shaped, the sulci will be narrow even when the rows are few, 84. The Cob.—The cob (f 83) varies greatly in shape and circumference. If the latter is over 44 inches it is de- scribed as “J/arge,” if from 34 to 44 inches as “medium,” and if 34 inches or under as “ smad/’’. The cob increases in length during the growing season of the plant. In colour, the chaffy 8 CHAP. IV. CHAP. IV. 114 MAIZE glumes on the cob may be blood-red or white. Forms inter- mediate in colour occur, but this may be the result of cross- pollination. Asa rule, colour of cob is a fixed characteristic of a breed, eg. in true Wickory King it is always white. White-grained breeds should have white cobs; coloured chaff discolours the mill-products. 85. Number of Rows of Grain.—The rows of grain on a cob vary in number from four to about forty-eight, but as a rule they range from eight to twenty in the breeds grown in South Africa. We have met with four-row and six-row ears, but these appear to be due to some abnormal condition, which has retarded the development of some of the rows. Odd numbers of rows occur very rarely ; such irregularity is prob- ably due to injury of one of the spikes of carpels during the stage of development. It often happens that a pair of rows fails to develop fully, both rows stopping short without reach- ing the apex ; this may, perhaps, occur to only one of a pair of rows (Fig. 83B), Sometimes one or two rows on one side of an ear fail to develop through lack of pollination (Fig. 62), probably owing to the silks having been unable to protrude properly. Ears are sometimes found in which the grains are so scattered that the number of rows cannot be traced (Fig. 384A); sometimes this breaking up of the rows occurs through- out the whole ear ; and sometimes it is confined to one part of the ear, most commonly to the butt. Each breed has its characteristic number of rows, eg. Hickory King has 8, Loutstana Hickory 10, Hickory Horse- tooth 12, Arcadia Sugar-maise 12, Black Mexican 8. In some breeds, however, the number of rows is not yet definitely fixed, eg. Jowa Silver-mine, as grown in South Africa, has 14, 16, or 18. This is, perhaps, due to crossing with other breeds or with different strains of the same breed. In some cases the percentage of a given number of rows is found to be distinctly higher than in others, as shown by the following cases, which suggest crossing :— Fars, Rows. Per cent. Breed. I4 24°00 Iowa Silver-mine. I4 32°58 Ladysmith. 16 57°00 Iowa Silver-mine. oF BOTANICAL CHARACTERS TIS Rows. Kars, Ty-row, 10-TOW. 15-row, Breed. Examined. Percent. Percent. Per cent. Iowa Silver-mine — 24 57 1g Ladysmith 89 a2°5 45 22°5 Increase in number of rows is often accompanied by a cor- responding decrease in the breadth of the individual grains. But this is not always the case; much depends on the thick- ness of the cob; the grains on a 16-row ear are sometimes broader than those on a 12-row ear of the same breed, if the cob of the former is thick while that of the latter is thin. Increase in number of rows means a larger number of grains, but it does not necessarily follow that it is accompanied by increase in amount of grain produced. In Golden Ning (yellow dent), the best I4-row ears gave more vrain than the best 12-row, but some of the 10-row also gave more than the 12-row. In Vellow Fforsetooth (yellow dent), the ten best I4-row ears also gave more grain than the corresponding [2-row ears, but the ten poorest ears of the latter gave more than the ten poorest 14-row ears. In Yellow Hogan (yellow dent), the seven best 14-row ears gave more grain than the seven best 16-row ears (Lurtt-Davy, 18). The same number of rows does not give equally good returns in all breeds alike. Each breed seems to have an optimum number, which gives the best results in weight, shape, and size of grain, above or below which deterioration commences. In some breeds the rows occur in distinct pairs; the rows are then described as a7strchous. 86. Lwisted Rows.—The occurrence of a twist in the rows, either to left or right (Figs. 58 and 59), is a common feature. It is often confined to the upper portion of the ear, but may start from near the base. It is an undesirable character, as it tends to the development of irregular grains. This twisting appears to be in some way associated with the development of the number of rows. Out of a number of cases examined the following figures were obtained (Burtt-Davy, 18) :— Row numbers. Left twist. Right twist. Ears examined. I4 17 = 59°6 per cent I2 = 41°4 per cent 29 Id I2 = 60°0 a 8 = 40°0 eA 20 16 Rows very straight, no twist. 29 = 59°3 per cent 20 = 40°7 per cent 49 CHAP. IV. CHAP. IV. 116 MAIZE 87. Number of Grains per Ear.—The number of grains per ear varies to some extent with soil, seasonal rainfall, etc., but apart from this, there appears to be a definite relation be- tween the number of grains and the breed. H7ckory King in the Transvaal ranges from about 350 to 400; /owa Szlver- mine runs from 800 to 1,100. Burton (1) reports a case in itis re Obs eeIV TES Ary ye avae OECEO ecoS® oeont =e 2 2 2 a > 2 pe] 2 ~J 2 ance 6d S88 et gogecdsee Se ac dang (saa ue CLO6e g) G6GOGGEOGUG On00enggEsEd a oudaG ages Gao TeVadddad v v et ik f A186 oeol6 eae Ws ¥ $ Fic. 58.—Left-hand twist of rows. Aliwal North Division, Cape Colony, 5. Africa, of an ear bearing 2,828 grains ; this was one of eight ears from the same plant, but the name of the breed was not stated. 88. Proportion of Grain to Ear.\—Vhe proportion of grain to ear is exceedingly variable, not alone as between breeds and under different conditions of growth, but also in the same ‘ie. percentage by weight of grain and cob. BOTANICAL CHARACTERS 117 breed grown under similar conditions. In the United States it is said that 86 or 87 per cent of grain per ear may be considered a fair proportion. In Transvaal-grown samples examined, the percentage has usually been much lower, aver- aging only 82°13 per cent. Fic. 59.—Right-hand twist of rows. There does not appear to be any connection between “igh yield of grain per ear and percentage of grain on the ear, The heaviest yielding breed sometimes gives the lowest per- centage of grain, while the one giving the highest percentage of grain may give a relatively low yield; but it does not follow that the breed giving the highest percentage of grain gives actually the lowest yield CHAP. IV. CHAP. IV. iT8 MAIZE American growers lay much stress on high percentage of grain to ear, because a great deal of maize is there sold on the cob, and buyers prefer strains which, when shelled, will bag up well. The growers admit, however, that the /rofor- tion of grain to cob ts of less importance than the actual weight of grain per ear. In South Africa, where maize is sold, entirely, off the cob, the question of percentage does not appear to be of great importance. 89. Form for Describing the Ear.—The following is a useful form for recording the characters of typical ears in order to determine from year to year whether any change is taking place or whether they are remaining true to type. Owing to the difficulty of keeping specimen ears for any length of time, a written record is desirable. For definition of terms, see preceding paragraphs under each head. This form can also be used to advantage by students in agriculture. In this case each student should have two or more ears of each of the five varieties of maize, or of five different breeds of the same variety. Ten ears of a given variety or breed are none too many for a thorough study. ars of other varieties or breeds, showing the characters here mentioned, should be shown for the guidance of students. The character present may be marked by a ,/ across the word. Name? Varlety.cvcssiscsaneaenece Bread riche ain ssanpeeacsmiaiaataanignton vasaserales Date of: (a) Record......... (6) Sowing......... (G)) RAVES tiie assess sence (a) Colour of grain: white; yellow; golden; red; purple; blue; or black. (b) Colour of cob: white; light red; deep red. (c) Surface: smooth; medium ; rough; very rough. (d) Sulci: absent; apparent; narrow; distinct; very distinct. (c) Pairs of rows: distichous; not distichous. (f) Number of rows: at $ length from butt......... ; from tipi. cicsce: (gz) Direction of rows: oeraehes right twist; left twist; irregular. (h) Grains: very loose; loose; firm. (‘) Grains: regular; mosaic-like; uneven. a Grains: upright; sloping; imbricated. (k) Ear: cylindrical; cylindraceous; slowly tapering; tapering. /) Butt: even; shallow rounded; moderately rounded; deeply rounded. m) Butt: depressed; compressed; depressed-rounded; depressed-com- pressed; enlarged; expanded; open. (x) Tip: sides of cob exposed; end exposed; end covered; capped. (0) Juncture of shank with ear: large; medium; small. ( ( (p) Extreme length of ear; maximum......... inches; minimum......... inches ; average of ten......... inches. (7) Circumference of ear at 2 inches from butt: maximum......... inches ; minimum,........ inches; average of ten.....,... inches, BOTANICAL CHARACTERS 119g (v) Circumference of ear at 2 inches from tip: maximum......... inches; CHAP minimum......... inches; average of ten......... inches. IV. : (s) Weight of ear: maximum......... 0z.; minimum......... oz.; average of CEN wicaanies O4. (t) Weight of cob: maximum......... 02.3 Minimums... +.. oz.; average of tet eaters oz (uw) Percentage of grain: maximum......... ; minimum......... ; average of ten......0.. (v) Circumference of cob at 2 inches from butt:......... inches. (w) Ratio of circumference of cob to that of ear:......... 90. The Gratn.—The grain of maize and other grasses is often spoken of as a seed, but it is in reality more than a seed, it is a whole fruz¢t. It is a peculiar form of fruit, for the peri- carp or outer covering of the fruit is completely united with the seed-coat. This special form of fruit is called a caryopsts, The caryopsis of the maize plant comprises the following parts:—the /ad//, which is the combined peri- carp and seed-coat; lying immediately under the hull is a layer of cells known as the a/ewrone layer (Figs. 15A and 60). The hull and aleurone layer together enclose the main mass of the grain. This consists of two distinct parts, the endosperm and the embryo (Fig. 15). The grains are arranged with the embryo side towards the tip of the Fic. 60.—Enlarged section through hull ear; when the ear hangs of maize grain. f, pericarp; ft, testa or down, the embryo is then seed coat; 1, perisperm ; a, aleurone fi layer; s, endosperm. (From Passmore and on the under side of the Webber.) grain. The grain may be firm on the cob or movable. Movable grains may, sometimes, be due to the ear being not fully mature when gathered, or to lack of adequate moisture at the time of ripening off. The grain may be set on at right angles to the surface of the cob, or it may slant towards the tip. me f, MAIZS: 120 (‘yoog-av9,X ainynousy jo juawziedag *S*~Q Wor) ‘sures aziew jo advys ul uONeIeA— ‘Ig “D1 BOTANICAL CHARACTERS 121 The shape of the grain varies greatly (Fig. 61). In most breeds it is flattened and more or less wedge-shaped, with an indented apex (Dent maize); in one variety it is spheroidal or conical (Flint maize); in Pop maize it is distinctly and sharply beaked; in Sugar maize, when dry, the grain is much wrinkled owing to absence of starchy endosperm. Depth and Breadth of Grain. The depth (or length) of the grain varies greatly, and is said to be a quite constant character in different breeds. Breadth and thickness of grain, on the other hand, are not considered so constant. Each of the five cultural varieties of maize (Pop, Flint, Dent, Flour, and Sugar) contains three well-defined sub-types, based on the relative breadth and depth of the grain. Thus :— Group A—Grain broader than deep. Group B—Grain as broad as deep. Group C—Grain much deeper than broad. According to Sturtevant (2) these depend on climatic con- ditions; the A group grows in short-season climates, the C group in long-season climates, while the B group is inter- mediate. A climate suitable for the C group will, naturally, also suit the other two groups, but they may not prove as profitable, and would therefore be less desirable. Deep grain appears, therefore, to indicate a long growing season, and vice versa; if this is constant, one should be careful not to select seed-maize having too deep a grain to suit the particular altitude or rainfall. Shape of Grain. (a) If broad above, tapering to a slender base with straight sides, the grain is described as straight cuneate. (b) If the same general shape, but with rounded edges, it is curved cuneate. (c) Broad above, narrower below, connected by straight lines, truncate cuneate. (d) Long and uniformly narrow above, only tapering to a more or less broad base, shoe-peg form. (e) Short, and as broad at base as at summit, rectangular. (f) Slightly rounded at corners, both above and below, round-cornered, CHAP. IV. CHAP. IV. 122 MAIZE Apex of Grain. (a) Roof-shaped at one edge, i.e. convex at one and flat at the other. (6) Shingled, i.e. overlapping like shingles on a roof. (c) Flat or square, corvers not rounded at summit. (d) Rounded, corners rounded at summit. (e) Rostrate or beaked, with long, sharp, tapering projection. (f) Mucronate or pointed, with small, sharp point at summit from embryo side. (g) Dented (only in dent breeds), with an indentation of varying size and form. Indentation of Grain. (a) Round dimple, rounded or cup-shaped and quite smooth. (6) Long dimple dented, i.e. longer than broad, and quite smooth. (c) Creased, i.e. edges pressed towards each other, leaving a smill space between and the edges parallel. (d) Pinched, the edges pinched closely together and projecting upward and forward. (e) Rough, with any rough, jagged, or beaked projection from the summit. (f) Bridged, with a fold across the centre. (g) Crumpled, or wrinkled, as in sweet maize. ol. The Hull.—The hull or outer covering of the ripe maize grain (Figs. 15 and 60) is hard and shiny. It comprises the pericarp of the fruit together with the Zesta or seed-coat (with which it is united), and the pertsperm, a layer of tissue beneath the testa and surrounding the endosperm (Fig. 15). Of these three tissues the pericarp forms the larger part of the hull of the ripened grain. The bull can be easily removed from the aleurone layer for study, by soaking in hot water for about fifteen minutes. 92. The Aleurone Layer—Lying immediately beneath the hull, between the perisperm and endosperm, is a tissue com- posed of a single row of comparatively large cells, rather regular and rectangular in transverse or cross section (Fig. 60). This tissue is called the aleurone layer ; it comprises 8 to 14 per cent of the grain. Webber (2) has shown that the blue, purple, and black colour of the soft flour and sugar varieties of maize lies in the aleurone layer. In yellow maize, however, the yellow colour- ing matter is not confined to the aleurone layer, but penetrates the endosperm. In some of the red-grained breeds of maize, and in the red-striped Cusco, the red colouring matter is con- fined to the pericarp, which accounts for the fact that red maize is sometimes produced from white seed. BOTANICAL CHARACTERS 123 93. The Endosperm.—This is the mass of tissue lying below the aleurone layer, but above and partly surrounding the embryo. It comprises about 73 per cent of the whole grain. Its function is that of a reserve store of elaborated plant-food for the use of the young seed-plant before it is able to absorb food materials from the soil or to elaborate them in the leaf. As seen in section, the endosperm shows a variation from translucence to opaque snowy whiteness. Hopkins (3) reports a difference of 2 per cent more protein in the corneous than in the white endosperm ; Hunt (1) questions whether this may not be due to lack of complete separation from the aleurone iayer in the samples analysed. The latter author points out that there is no material difference in structure noticeable under the microscope, which has led to the suggestion, not positively proven, that the difference between them is a difference in density analogous to the difference between snow and ice. The relative proportion and arrangement of the translucent or corneous and the white endosperm have been used in part to differentiate between the five varieties of maize. 94. Form for Describing the Grain.—This may be used to advantage for the same purposes, and in the same manner, as the form for describing the ear. For the use of students in agricultural botany, twenty-five to thirty grains should be given of each of the five varieties, or of five breeds of one variety. For determining the points in (2) to (f) inclusive, a number of grains should be soaked in hot water for thirty minutes, or in cold water for twenty-four hours. For measuring the grains, a sheet of cross-ruled paper can be used to advantage for marking off distances, which can then be measured accurately by a scale divided to 32nds of an inch, or preferably to millimetres. Natnes: Varrety cs sicesioaasedieues Bre 6d spy srs aionmsvui ins seamen nwanarsenaaaysntennh Date of (a) record......... ; (b) sowing......... ; (c) harvest...... .- (a) Weight: ten average grains in duplicate (a)...... .. sD ace ceay (b) Length: ten average grains in duplicate (a)......... ra (2) oneness (c) Width: ten average grains in duplicate (a)......... © UB) iasoaienys (d) Thickness: ten average grains in duplicate (a)......... S(O cagicetaris (ce) Ratio of width to length: divide length of ten grains by width of ten grains (d)......... a (DNicemanraxs (/) Ratio of thickness to width: divide width of ten grains by thickness of ten grains (a)......... So (D)iseencase CHAP. IV. CHAP. IV. 124 MAIZE (g) Shape: flat; spheroidal; conical. (h) Shape (side view): straight cuneate ; rounded cuneate; curved cuneate ; truncate cuneate; shoepeg ; rectangular; round cornered. (1) Apex: roof-shaped; shingled; rostrate; mucronate; rounded; flat ; dented. (7) Indentation: round dimple; long dimple; creased; pinched; rough ; bridged ;_ wrinkled. (k) Colour: white; cream; yellow; golden; red; blue; purple; black ; striped; mottled; mosaic. (1) Place of colour: endosperm ; aleurone layer ; hull. (m) Character of endosperm: corneous; partly corneous; farinaceous ; sugary. (x) Proportion of corneous endosperm (in dent variety): large; medium ; small. (0) Embryo size: large; medium ; small. (p) Sketch of longitudinal cross-section: show arrangement to scale, of embryo, and of corneous and white endosperm. 95. Tubular Glands in the Embryo.—Dr. C. Stuart Gager (1) describes the occurrence of true glands of the tubular and sub-racemose type in the tissue of the scutellum, formed by invaginations of the glandular epithelium of the latter. The significance of these glands, as in harmony with the theory that the scutellar epithelium is principally an organ of secre- tion, is indicated by the author. 96. Afogamy.—Collins (4) has described a case in which the staminate flowers were replaced by young leafy and root- forming maize plants. Addendum.—Since this chapter went to press the writer has seen a paper by Prof. Emerson (1) of Nebraska, in which he describes and illustrates a family of maize in which the ligule and auricle were absent. He refers to a similar con- dition in oats reported by Nilsson-Ehle (1) and Hurst (1). Prof. Emerson notes that in the case of non-auriculate leaves, the sheath and the lower part of the blade are rolled some- what closely about the stalk, and that the leaf as a whole assumes an upright position nearly parallel with the stalk, the tip of the leaf curving away gently if the blade is long and the mid-rib sufficiently flexible. “Whether the abs- ence of the ligule proper is disadvantageous to the plant is somewhat questionable. . . . My own observations on liguleless corn are to the effect that the inside of the sheath is more often discoloured, as if from incipient decay, than is the case when a well-developed ligule is present. In no case, however, have I found decay of the stalk or leaf sufficient to BOTANICAL CHARACTERS 125 be of any material injury to the plant—not even during the past summer when the plants were grown near Boston, where they were exposed to moister conditions than is usually the case in Nebraska.” He concludes that it is the absence of the auricle that makes the leaves stand so erect. “It is the triangular shape of the auricle that makes possible the immediate flattening out of the leaf blade at the termination of the cylindrical sheath, and that allows the blade to bend abruptly away from the stalk while the sheath still clasps it.” CHAP. IV. CHAP. CHAPTER V. INHERITANCE OF CHARACTERS AND IMPROVEMENT BY BREEDING. Tis often seen adoption strives with nature, and choice breeds a native slip to us from foreign seeds. —SHAKESPEARE. And he gave it for his opinion, that whoever could make two ears of corn . to grow upon a spot of ground where only one grew before, would deserve better of mankind, and do more essential service to his country, than the whole trace of politicians put together. —Gulliver’s Travels. Necessity for [mprovement. 97. The Olject of “ Breeding”.—Plant breeding is the application to crops of the principles applied in improving breeds of live stock. No breed of domesticated animals or plants is perfect in all respects; each one has its good and its weak points. There are therefore two primary objects which the breeder keeps in view in order to produce satisfactory results: (a) macnfenance of quality and type, by the elimination of the unfit and untrue to type; (0) zwprovement of the type by the substitution of desirable for undesirable characters. To allow the poor types in a herd or crop to propagate their kind always results in race deterioration. To allow only the strongest and best to mate and propagate means, on the other hand, race maintenance and also, within certain limits, race improvement. “In the herd of cattle to destroy the strongest bulls, the fairest cows, the most promising calves, is to allow those not strong, nor fair nor promising, to become the parents of the coming herd. Under this influence the herd will deteriorate, although the individuals of the inferior herd are no worse than their own actual parents. Such a process is called race- 126 INHERITANCE—IMPROVEMENT BY BREEDING 127 degeneration, and it is the only race-degeneration known in the history of cattle or men. The scrawny, lean, infertile herd is the natural offspring of the same type of parents. On the other hand, if we sell or destroy the rough, lean, or feeble calves, we shall have a herd descended from the best. . . . “In selective breeding with any domesticated animal or plant, it is possible, with a little attention, to produce wonder- ful changes for the better. Almost anything may be accom- plished with time and patience. To select for posterity those individuals which best meet our need or please our fancy, and to destroy those with unfavourable qualities, is the function of artificial _ selection, Add to this the occasional crossing of unlike forms to promote new and desirable variations, and we have the whole secret of selective breeding. This process Youatt calls the ‘magician’s wand’ by which man may sum- mon up and bring into existence any form of animal or plant useful to him or pleasing to his fancy” (/ordan, 2). To accomplish the best results possible from breeding, it is necessary (@) to understand something about the manner in which characters are inherited from the parent; (0) to be well acquainted with the characteristics and variability of the plants or animals with which one is dealing; and (c) to understand clearly what characters are required by the market, or are in other ways desirable or undesirable. 98. The Necessity for Improvement of Crops.—South Africa produces good maize and has established a good name in the maize market, thanks to the assistance rendered by the several Governments. There is no difficulty in selling the maize of South Africa; the difficulty is to supply the demand. There is even danger that, unless the output is increased, she may lose the market that she has gained, for Europe requires a steady and dependable supply; irregularity of supply tends to dis- credit the crop with the merchant, and reacts unfavourably on the producer. South Africa must produce more if she is to become the “maize granary of Europe”. Increased production depends upon three things: (1) more intensive cultivation of the area now under crop; (2) a larger farming population to increase that area; (3) increase of the yield per acre by scientific maize- breeding. CHAP. V. CHAP. 128 MAIZE Intensive cultivation means greater profit; if we can pro- duce 600 muid bags of maize from 50 acres, che profit per bag is much greater than if we get only 300 bags from the same acreage, for the rent of the land and first costs of ploughing, harrowing, and cultivating are the same in each case. Through breeding we may further increase the produc- tion by developing drought-resistant or early-maturing sorts adapted to regions now outside the Maize-belkt. With low yields, maize-growing ceases to be profitable when prices also are low; but if we double and treble the yield per acre, prices may fall much lower and still yield a good profit. The remedy for low prices lies not in restrictive fiscal legislation, but in reducing the costs of production by increas- ing the yield per acre and improving the quality of the crop. 99. Need for Increase tn the Yield per Acre.—QNiartley states that good farmers in the United States frequently grow from 75 to 100 bushels of maize per acre. An American bushel of maize on the cob weighs 70 lbs.; 75 bushels or too bushels would, therefore, be equivalent to 26 or 35 muid bags per acre. An American bushel of shelled maize must weigh (by statute) 56 lIbs., which would be equivalent to 21 to 28 muid bags per acre. Maximum yields are, of course, higher. A certain Pennsylvania farmer has been known to harvest no less than 100 bushels (23 muids) per acre during twelve successive years (excepting only two seasons), and in 1902 his average yield over 90 acres was 130 bushels or 36°4 muids. The Transvaal records are pretty good, for, at Tzaneen in the Zoutpansberg District, the Department of Agriculture has produced 354 muids (127 bushels of shelled maize) per acre, of dust7 Colossal yellow dent, and at the Government Experiment Farm, Potchefstroom, 35 muids (125 bushels) of a white dent. There seems reason to expect, therefore, that given proper treatment of the crop, and with heavy-yielding varieties and well-bred seed, good farmers in the Transvaal will raise an average of 20 muids (71 bushels) per acre ; it is an ideal worth working toward! c » ies) os Aabb + aAbb = 2 Aabb aaBB = 1 aB aaBb + aabB = 2 aBb aabb = I ab Thus there are nine cases in which both dominants (A and B) meet; three contain the dominant A, without B; three contain B without A, and one carries the two recessives @ and 6, without either dominant. This gives the ratio :— Oly Se 3 GB ss a8 or 56:25 : 18°75 : 18°75 : 6°25 per cent, CHAP. 184 MAIZE When this ratio is met with in a heterozygous family, we conclude that we are dealing with a adzhybrid. The following example of a dihybrid cross (ear 1,157) between a white wrinkled and a red starchy maize is taken at random from among the writer’s records :— TABLE XXXIV. ANALYSIS OF A DIHYBRID CROSS BETWEEN WHITE WRINKLED AND RED STARCHY MAIZE. | Starchy. | Wrinkled. | | | 5 ——_———| Total. | | Red. | White. | Red. | White. ee | | | | 15 ee || oe ee ar | | 16 6 | 10 | = 33 | | 18 3 | 8 | 3 2 | |, Z 9 i, Oe ees 31 16 8 | 4 | 4 2 , 79 4 4 | 4 31 19 4 i SO i 3 2 Ti: 9 ass ib 2 30 14 7 | | 1 29 | 18 5 | 6 hy 8 31 oe) 4 | zy | 8 30 | 17 Fs 4+ 3 31 | | | | Total . : | 202 | 93 | <9: | 26 | 373 | Average per row. : Sil SN 6 | 6 lee 31 | Pereent . é ; .| 54°16 | 19°57 19°30 | 6°97 — Mendelian expectation | 56°25 | 18°75 | 18°75 | 6°25 _— 139. Trthybrid Rattos——Crosses are met with in which a third factor, which we may designate as C, has an influence on the second factor B, so that when the two meet they produce colour, but when ether is absent, the appearance is the same as though wez¢her was present. In such cases we get the ratio 48:9:7. By growing on the seven recessives we find that they are not homozygous, but consist of 3 B, 3 C and 1 abc, the latter only (ie. #y of all) a pure recessive; the actual ratio 1s therefore :— 4d: Gas nO As ok or 75: 14°0625 :9°375 : 15625 per cent. When three allelomorphic pairs (e.g. Aa, Bb, and Cc) enter into the cross, there are sixty-four possible combinations of these characters. In such a cross the results will be:-— INHERITANCE—IMPROVEMENT BY BREEDING ABC 27 ABe 9 — = 30 AB Abc 3 AbC 9 — = 12 Ab = 45 with A aBC 9 —= 9BC aBe 3 abC 3 —= 6BorC=15 , BC,B,orC abc I Ey abe 64 64 185 The distribution of the gametes is shown in Tables XXXV, XXXVI and XXXVII following. TaBLE XXXV. DISTRIBUTION OF THE GAMETES IN A TRIHYBRID CROSS. ABC | ABC | ABC | ABC | ABC | ABC | ABC | ABC | ABc | AbC | Abe | aBC | aBc | abC ABc | ABc | Ape || ee | ABc | ABc | ABc | ABC | ABc | AbC | Abc aBC | aBc abC See ae, fal —|—| (ae == | AbC | AbC AbC AbC | AbC | AbC | AbC ABC ABc AbC Abc | aBC aBc | abC | | | | | i | assent ner ee a ee ee eT Abc Abc Abc Abc | Abc Abc | Abc ABC | ABc | AbC Abc aBC | aBc | abC aBC | aBC | aBC | aBC | aBC | aBC abc ABC | ABc AbC Abc | aBC aBc abC : aie | | aBc | aBe aBc aBc aBe | aBc | aBe | ABC | ABc | AbC | Abe | aBC | aBe | abC (iene : | abC | abC abC abC abC abC abC ABC | ABc AbC Abe aBC — aBe abC pee ee ee ee abe =| abe abe | abc abe | abe abc ABC | ABc AbC | Abc aBC | aBc abC | | The above “chess-board” is classified in Table XXXVI following :— CHAP. 186 MAIZE TaBLE XXXVI. CLASSIFIED SUMMARY OF TABLE XXXV. AABBCC. AABBCc + cC. AABBcc . AABbCC. | AABbCc + cC AABbcc AADbbCC . AAbbCc + cC . AAbbce AaBBCC. AaBBCc + cC. AaBBcec AaBbCC . | AaBbCc + cC AaBbcc . | AabbCC AabbCc + cC Aabbcc aaBBCC . aaBBCc+cC . aaBBcc aaBbCC . aaBbCc + cC aaBbce aabbCC aabbCc + cC aabbcc . + bB lip I I 2 2 I I tee oe aes ay cas ) ie = | 20 — | ie | 24 Tt bop aA 3 a | Ir I | ge Se | 2 = J I= ees ae I I I 2 Ps I I I — a = Pag cee BNR HN H ARDY | ww hes] tate 3 4 to (2 Aoi eae [S| neve oe DaED PE eee A TABLE XXXVII. SUMMARY OF TABLE XXXVI. Per Cent. ABC 27 = 42°1875 AB 9 = 14'062 AC 9 = 14°0625 A 3= 4°6875 BC 9 B 3 = 46875 Cc 3 = 476875 abe T= 1°5625 48 = 75'0 9 = 14°0625 7 = TO'9375 Per Cent. 1°5625 3125 1°5625 16 IZ | 32 . }10 IZ | INHERITANCE—IMPROVEMENT BY BREEDING 187 It is conceivable that there might be other interactions of CHAP factors in a trihybrid combination of this character, which would _ Y- give quite different Mendelian ratios; for instance if A were only visible in the presence of C we should have a ratio of— 36 AC: 21 B:3 A:3 C:1 abe of which A and C might conceivably give the same appear- ance as abc, giving an apparent ratio of 36:21:7. Bateson gives the following scheme by which the number of types, and the ratios in which each will appear, is given for any number of pairs of factors, one factor of each pair being dominant and the other recessive. 4=341 TO= (31)? S37 +343 +0S 9+ 34342 64 = (3 + 18 = 3% 4+ 33° +33 41527 + 27+ 941 250 = (3: aL)? = 34 gest Org? tae a ll 8r + 27 + 27 + 27 + 27 TQ Oe Oba Oi FOr +3+34+ 343 +1. So in general yt = 3" ae eat tae ea n times ee eee ec oe dn (uu — 1) times ne ea oa ean oe 4n (a — 1) (2 — 2) times. + etc, 140. Inheritance of Colour.—There are several different colours in the maize grain, and also in the vegetative part of the growing plant, which behave as separate allelomorphic pairs, transmissible independently. In a few cases coupling of allelomorphs appears to take place. Owing to the different behaviour of these several colours it will be convenient to dis- cuss them separately. 141. Vellow Endosperm.—Yellow colour of the endosperm and its absence behave as an allelomorphic pair. Shull, East and Hayes, and the present writer have found two kinds of yellow each of which behaves with its opposite as an independent allelomorphic pair. One of these (the darker yellow) gives the Mendelian ratio 1: 2:1; the other (the paler) gives the ratio g: 3:3:1. In some cases the latter is so faint as to be easily overlooked ina poor light, and has been mistaken for a ‘‘dominant white”. If non-yellows breed true CHAP. 188 MAIZE in the F, generation, they are usually considered pure (Aas¢ and Hayes, 1), but if the absence of yellow is due to the absence of a factor on which the appearance of colour depends, they may, even though heterozygous, continue to breed white until crossed with another white carrying the complementary factor, when yellow will appear. The relative amount of soft and horny starch contained in the seed is one cause of varying in- tensity of the yellow colour in F, crosses between yellow and white. East and Hayes appear to have found six shades of yellow in the progeny of a cross between yellow and white maize, for they observe that “in the case of the two yellow colours in the maize endosperm, the intensity of the yellow decreases in the following order :— Ya Ya Yo: Ya Y,y1 Y2 Y. or Y, Y, Y1 y Y, ¥i or Y, Yo Yi yi or Yy Yo Y1 1 Yo Yo [i-e. pure white ?]”’ The present writer has found six shades of yellow in the F, seed generation, and ten shades in the F, seed generation after crossing with white. The two yellows already referred to are involved in the production of these shades, and there is a distinct break in the gamut of tints between what appears to be the palest of No. 1, and the darkest of No. 2. 142. ‘‘ White Starchy”’ Endosperm.—This was shown to be a Mendelian dominant by Correns in t901, and confirmed by Lock in 1904, and later by the writer and by East and Hayes working simultaneously though unknown to each other. East and Hayes found that dominance was complete; in no case was there the slightest difference between the homozygous and the heterozygous seeds in either outward appearance or in the character of the starch cells when examined microscopically. 143. /nherttance of Characters which Affect the Growing Plant.—TVhe characters which we have been discussing are those which affect the endosperm, and are therefore visible in the daughter seeds of the ear that has been crossed, i.e. the first new generation. There are other characters, however, affecting the growing plant, which do not show in the seed, These will be discussed seriatim. INHERITANCE—IMPROVEMENT BY BREEDING 189 144. Pertcarp Colour.—The colour of the maize-grain is sometimes found in, and confined to, the pericarp; in such cases this is readily demonstrated by soaking the grain in water until the pericarp can be peeled off without removing the aleurone layer. The red striped Cusco flour corn is a case in point ; we also meet with sporadic cases of red ears appearing unexpectedly in crops of white breeds, which, on examination, prove to be cases of red pericarp colour. The pericarp of the grain belongs to the parental, and not to the filial, generation; it is part of the female plant parent. The pollen grain therefore does not produce xenia in the peri- carp. Thus if a white breed is crossed with pollen of a breed with red pericarp colour, the resulting grain (F, seed genera- tion) will not show the red colour, though this will appear in the second (F, seed) generation. The present writer has found four distinct red pericarp colours, and East and Hayes describe five, which they call R,, R,, Rs, Ry, and R,. R,.—An ordinary dark red pop-corn ; glumes of male florets sometimes reddish, but neither cobs nor silks red. Crossed with white it gave 75 R and 22 W ears in the F, generation, the reds being all dark and the whites showing no trace of colour. The writer has met with a red pop, answering to this description, in South Africa. R,.—A dark mosaic red, occurring as irregular red stripes radiating from the point where the silk was attached. A similar red occurs in South African maize. R,.—A dirty red colour, more abundant at the base of the seed and almost wanting at the summit; it appears to be coupled with red silks. The dye occurs in small amounts. “Tt is almost certain that this red forms an allelomorphic pair with its absence, that is entirely independent of R,, R,,and R,.” R,.—A rose-red, which “ develops only in the presence of light, hence the ears with thick husks show the colour but faintly. When the husks are stripped away and the ear matures in full sunlight, the colour appears over the entire ear as a bright rose-red.”. Red was not present in other parts of the plants grown. A rose-red answering this description occurs also in South Africa, where it threatens to give some trouble to growers. CHAP. 1g0 MAIZE CHAP. R,.—A rose-red resembling R,, but occurring in lesser careful examination. amounts, and on thick-husked ears only detected by very It behaves with its absence as a sepa- rate allelomorphic pair to R,. RK,.—Is a dark red pericarp colour of sugar maize, from an jear of unknown parentage, found by the writer in South lic. 80.—Somatic variation in pericarp colour. Africa. The combined red colour and wrinkled endosperm tend to give the impression of dried raisins. 145. Somatic Variation in Peri- carp Colour.—Cases not infrequently occur in which an ear in a crop of white develops red pericarp on one side, or part of one side, of the ear, and white, or white striped with red, on the other side or part of that side (Fig. 80). Such cases have been attributed by East and Hayes to bud variations similar to those which occur in perennial garden plants, and also, though less fre- quently noticed, in annuals. In such cases the plant due to produce a red ear varies somatically so that one part of the ear becomes red and the rest white or striped. In a case which they record, this variation was transmitted by the seeds. 146. Szlk-colour. — Maize silks vary from almost colourless, through cream and green, to dark red. Some- times the style itself is red, some- times only the hairs on the style. Sometimes the colour of the silk is coupled with that of the pericarp, and perhaps also with colour in the glumes. But redness of the silk occurs commonly when the cob and pericarp are not coloured. According to East and Hayes, study of the transmission of this colour char- acter is obscured by the action of the bag over the ear to be hand-pollinated, which prevents the full development of the INHERITANCE—/IMPROVEMENT BY BREEDING 191 red colour by shutting out the light, so that it is difficult to CHAP. tell whether the F, silks which are selfed are full red, or only Me red-haired. 147. Red Cob-colour.—ixperiments conducted by the writer (and it appears, simultaneously by East and Hayes (1)), show that red cob-colour is dominant to white cob-colour, and behaves asasimple monohybrid. These authors suggest that “it is not beyond probability” that dihybrid reds may be found in an extensive series of crosses. Cob-colour is not necessarily coupled with red endosperm colour, for red cob-colour is not infrequently found in a white-grained breed, e.g. Hickory King, which normally produces white cobs. Many yellow-grained breeds have red cobs, but white-cobbed ears are often found amongst them; other yellow-grained breeds normally have white cobs, but red-cobbed ears are occasionally found amongst them. East and Hayes record a case in which perfect coup- ling occurred between red cobs and red pericarp colour. 148. Glume Colour—The glumes often carry red colour, and vary considerably in the amount of redness present ; some- times it is confined to very narrow or broader streaks along the nerves. Glume colour appears to be correlated with colour in other parts of the plant, for East and Hayes state that they have not yet found a plant which has red glumes and yet shows no red colour in other parts of the plant, though one has been found that is pure for red glumes and yet shows no red in other parts with the exception of the silks. 149. Development of “ Pods”.—Pod maize (Fig. 51) when crossed with breeds free from pods, behaves as a simple Men- delian monohybrid. The podded character is dominant, and appears in the F, generation in the proportion of 3 to 1; the extracted recessives breed absolutely true (Zas¢ and Hayes, 1). 150. Inheritance of Ligule and Auricles—Emerson (1) finds that absence of ligule and auricle behaves as a recessive to presence of these organs, and that they are transmitted as a single hereditary character. Four liguleless plants, crossed with unrelated normal-leaved plants, produced 103 F, individ- uals, all with normal leaves. Twelve of the latter were selfed, and produced 748 F, plants, of which 572 had normal leaves and 176 had no ligules, or practically a 3:1 ratio. Counting together all families in which both types of leaf occurred, he CHAP. 192 MAIZE obtained a total of 672 normal-leaved and 221 liguleless- leaved plants ; to have given as nearly a 3:1 ratio as possible the respective numbers would only have to be changed to 670 and 223. 151. Physical Condition of the Starch.—Presence and ab- sence of starch, as we have already seen, behaves as a Men- delian allelomorphic pair; the presence of starch is a filial character, and shows as xenia in individual seeds. East and Hayes find, however, that the physical condition of the starch behaves as a plant character affecting the whole ear :— “The characters which give the flint or the dent appear- ance to maize are transmitted as plant characters to the entire ear and not as endosperm characters to the seed. They conform to the essential feature of Mendelism by showing segregation ; and they are due to the action of more than one transmissible character. The question remains, can any or all of these characters be named? Our experience suggests that the proportion of corneous starch to soft starch depends partially upon size and shape of the pericarp, and upon the number of rows per ear... . There is also some relation between -the size of the plants and the amount of soft starch in their seeds. . . . Relationship between the physical char- acter of the starch and shape of pericarp is much more intimate than it is between the former and size characters. . . . The shape of the pericarp depends somewhat on the number of rows, as the greater this number the more the seeds are crowded together and thus lengthened. . . . These relation- ships may simply be correlations and not direct causes of the proportion of corneous starch to soft starch that exists in various strains of corn, But even if they were directly concerned, they could not account for the large number of differences in varieties, for none of the correlations are suffi- ciently high. Many other characters, the exact nature of which is unknown, must be concerned in the matter. The simplest interpretation . . . seems to be the interaction of independent allelomorphic pairs, of the nature reported by Nillson-Ehle (1) and East (4) in earlier papers.” 152. Stse Characters—Among size characters are in- cluded: height of plant, number and length of internodes, thickness of stem, breadth and length of leaf, length of sheath, length of ear, number of rows per ear, thickness of cob, size of seed, etc. INHERITANCE—IMPROVEMENT BY BREEDING 1 93 In the maize plant the factors which interact to cause the transmissible differences in the size of the organs are very complex, as has been pointed out by East and Hayes, who refer to the consequent difficulty of working out in detail their inheritance. “Tt is perfectly obvious to one familiar with the maize plant that it is almost impossible to work out in detail the inherit- ance of the complex factors that interact to cause the trans- missible differences in the size of the organs. “That size characters are complex in themselves is shown by the numerous varieties grown commercially. They each vary from their own means, but different variety means in height are found all the way from two and one half to fourteen feet, with but little actual difference between the most similar strains. Further to complicate matters, all size characters respond to environmental stimuli, and these non-inherited fluctuations obscure the analysis of pedigree cultures in a still greater degree. “For these reasons we do not attempt to analyse our results further than to say that they do show segregation tn every case. And segregation ts held to be the tmportant and essential feature of Mendelism. Therefore we belteve that size characters Men- delise. . . . But in size characters dominance is probably very incomplete or absent. . . .” “ Several genes for the same character may extst in the germ cells of one organism, the number being limited possibly by the number of chromosomes. The limited number of cases, thus far found, presumably is due to the fact that few size characters have been investigated, for nowhere would these phenomena be so likely to occur as in quantitative characters. . . . Several independent allelomorphic pairs may produce the same somatic character.” “A heterozygous combination presumably produces half the effect of a homozygous combination. Then as dominance becomes less and less evident the Mendelian classes vary more and more from the formula (3 + 1)” and approach the normal curve of error ($ + $)”. When there is no dominance and open fertilization, a state is reached in which the curve of variation simulates the fluctuation curve, with the difference that the gradations are heritable. The heritable variations are al- ways more or less obscured, however, by the ever present fluctuation,” 13 CHAP. MAIZE 194 *SASS¥[D [BIJUI AY} puNose yuasaauOd A[BU0NS |]V 6or | es ae eee | aoa G Qzz | - I z |h |g |6 joz 6tg abs = -— |€ |€ lee |r€ of - a gz | - e4=|-=|—= we cc fe Be He en esha eae oor = any jes {tee [pane chr | I € {2 bx |rx jrz lo Keaneaers ~ es eas be ae 1eI0L 16 gg | Sg zg | oSt. F Sg.c1 fof. Fez.or | C16. F tz.1g | -|- lc b is 6SS. F z0.c1 gtr. 4 6S.6 1zg. + gt.64 Sehr |r SLo. + PEEL LoS. + gl.1r | cog. FEE.BS |= Zz |g jor |€ gg. $SL.S1 | ggh. gl.ci Sg9. Foo1g | — zc |r |r |g eos. ¥ 89-8 61S. F 12.8 obZ. F £S.F6 - Lor |r oS 6Lz. + 10.$ fea. 3 £05 oot. # gr.tor | £ g itr ig +1 zh. = 16.6 | tof. 6F.9 | gzb. F 72.99 | ay ea allel pee: gor|So1/zo1 66 ia) a's Vv | V(SAAVH GNV LSVA YALAV) SSOUD V NI SLNW1d AZIVN JO SLHDIGH AO NOLLNETALSIC AONANOANA CHAP. V. | | “syue]d JO SJYSIaz] JOF sayIUy ul seajuag sse]D qucar ae *s1¥9 JUDIAJIP 9914} WOIF UMOID « "S189 JUIIAYIP OM} WOIJ UMOID 5 ‘19 ]UDIIYIP AAY WOIF UMOID ¢ ‘pajyunoo sjuxjd jo Jaquunu pue painsvaul aiaM SaWa1yXY 5 ‘uosvas aus ul apis Xq apis uavord squid oy samsy Surars suoinqiysiq _ €xbr \rzlLz 66 76 | Lr F€ cL og gg ifs Er gf 6c SE Ex £19 |S |r | ger L11/S6| PF | gz |gtig |r | Pa Sier= or 19 |Z | | QS SS 2S 6b ob St ‘THIAXXX S14v or 11/6 lgt bz Lze| 11 \1z 418 lis le x S) rig jg joz |Szizz|€rj6 | jz1]) b+ x ¢) g |S lo ler lg |zr{S jo le lrlze |= x S) org €xr xr £ or gt jez sale G4 x ¢) 9/6 \6 \€ je |s }-J-|-|-| - ,- x $) frie le jr [—)—t—-jy-l-}-) - J- Z| Se Sate ys ig) jorjeniea| 41 11 ire alsa lesen ella eee aes i 96 £606 [Lg tg 1g gt [sclee 69 99 £9 1 INHERITANCE—IMPROVEMENT BY BREEDING 195 ‘In considering experiments in the inheritance of size characters in maize, we must remember that fluctuations are present, and that often many genotypes are present in one parent.” 153. Inheritance of Hetght of Plants.—The results of East and Hayes’s investigations, as far as described in their paper (1), show segregation from the lowest class range of the shorter parent to the highest class range of the taller parent, but they do not consider these segregates as pure types, and “their behaviour in further generations is still problematical”. In every case the comparative size of the coefficient of varia- tion was at least 50 per cent higher in the F, generation than in the F, generation (see Table XXXVIII). The F, generation is not inter- mediate between the two parents, but is nearly as high as the taller parent. This fact, they point out, is not to be regarded as in any way connected with dominance, but is Fic. 81a.—Inheritance of size characters. due to the increased Hickory King, 8 tog feet high. (Type used to produce cross shown in Fig. 82 and grown vigour of the maize plan 2 ae pleat the same season.) which comes from cross- ing, as pointed out by East (3) in a previous paper. An entirely different case has come under the observation of the writer, in the F, progeny of a cross between AHzckory King (Fig. 814) and IVzlls Gehu (Fig. 818). The relative height is shown by the walking-stick which stands alongside, at the same relative distance from the camera as the stem of the plant, in each case. In this case the F, plants, Figs. 824 and B, show the dwarf habit of the IW7//s Gehu parent, and little, if any, increase 13 * CHAP, Fic. 818.—Inheritance of size characters. Wills Gehu, 4 to 44 feet high. (Type used to produce cross shown in Fig. 82 and grown the same season.) Fic. 82a,.—F, plant progeny of cross between types shown in Fig. 81; plants 44 to 5 feet high. INHERITANCE—IMPROVEMENT BY BREEDING 197 in vigour due to crossing was noticeable, such as was met with by East and Hayes. The same result was obtained with the reciprocal cross, showing that in this case the short habit is dominant. 154. laherttance of Abnormal Dwarfness.— Abnormally dwarf forms, 14 to 3 feet high, sometimes appear, possibly as examples of “reversion” or of “mutation”. East and Hayes Fic, 828.—F, plant progeny of cross between types shown in Fig. 81; plants 44 to 5 feet high. find that segregation takes place in their inheritance, in the proportion of three normal to less than one abnormal, but the number of cases studied was perhaps inadequate to determine whether it was a case of Mendelian dominance. 155. Inheritance of Length of Ears.—Ear length does not show the increased vigour, according to East and Hayes, due to heterozygosis, that is seen in the heights of the plants. They conclude that there is scarcely a doubt that the greatly CHAP. MAIZE 198 “S189 9914} WL] UMOID 5 “S189 OM} WOIZ UMOID *sI¥9 JAY WO UMOID ; ‘or6r ul uoNeiouad *y ‘6061 UL UAOIS UONLsaUAs | put og ‘ON $(1) SadvH puL seq WoL | x | || | ] l l bel of.c0 | Leo. F 1g.z | ger. F g.z1 | 1 | z | I zr jae |-or.)|) er.) te.) 2a | Ee | €o| Gb] £3 | S z | | | €1t.F thdr| €S0. F Sz.z | odo. F621 | —| —| x | 6 lor cz 6€ 89 | 99 €L | Lr gz 6x ort | Sige Hae | dears Beal geece tee a an teen eed el | ie Sex] og | oS | ez |S | + | gel. = gb.cr| ggo. F 1Sa | rer. Fut | — eee Beet es | + 6 | dr | tr | ex G1 ||-E Vata | | | | | | | r€6.F €r.11| ggo. F LZg.r | ri. FQ.gr | z ! | or | Sx | gz | Sx zi |air|e& |—|—]— costes | ee | €gl. 4 dz.cr! 10. F 1g. elo. ¥ 9.9 | — cee pe et eos vee =) ae ae Se Se nee | | | | a | =| | foment | | | | | | | pte | er ee a ae or | Sx | Fr | €x ut ie Ol ea. "AD a's ¥ dee Cole: ed seul ee ee eee eee eS ae | “sieq jo syIsuayT IOj “WD Ul samqUad Sse[D “(FS x 09) SSOND NI SUVA AO SHLONAT AO NOILAGINLSIG AONANOANA CHAP. V. UXIXXK @TA¥ LT INHERITANCE —TMPROVIE MENT BY BREEDING 199 increased variability in F, (see Table XNNXIX) is the direct CHAP. result of segregation. Me 156. Inheritance of Size and Weight of Grain.—East and Hayes conclude that segregation occurs in the progeny of ears heterozygous for size and weight of grain. B A Fic. 83.—Inheritance of row numbers. A, ro-row Hickory (Louisiana). B, Hickory with 10 rows below and 8 above. 157. Inheritance of Row Numbers.—The number of rows of grain in an ear varies according to the breed. In some’ breeds it is more definitely fixed than in others, e.g. Hzckory King; many of the flint breeds are normally 8-rowed and rarely exceed that number when pure-bred; in these cases an ear carrying more than 8 rows is considered untrue to type, CHAP. 200 MAIZE but it is doubtful whether any commercial maize crop grown breeds entirely true in regard to row numbers. That this is traceable, in part at least, to fluctuating variability, seems to be indicated by the following cases of irregularity :— Not infrequently ears are met with in which two or four rows cease before reaching the tip (Fig. $38). That such cases may be dependent on nutrition, as affected by variation in the character of the season, is suggested by the fact that in the season 1910-11 the writer found ears bearing two more rows on the upper (tip) portion of the ear than on the lower portion; the weather was dry during the early part of the season, which tended to check development, but was wet during the latter part of the season. The possible connection between the season and such cases requires further investigation before we can definitely connect them as cause and effect. In some cases, as shown in Fig. 84B, it is clear that a pair of rows has been dropped; this is unusual, however. More frequently one row of each of two pairs has stopped (Fig. 83B). Many cases have come under the writer’s notice, in which two ears borne on the same plant produce different row numbers; in thirty-three plants of Arcadia Sugar-maize each bearing two fully-matured ears on one culm, twelve plants bore an equal number of rows on each ear; fourteen plants had more rows on the lower ear than on the upper; on seven plants the largest number was borne on the upper ear; in most of these cases there were but two extra rows, but ina single case there were four more. The actual results obtained are shown in Table XL. INHERITANCE —IMPROVEMENT BY TARLE XL. BREEDING 201 ROW NUMBERS IN A FAMILY OF ARCADIA SUGAR-MAIZE., Plants with Two Ears. Plant No. wotlbiet Rows Surticrer news : | 8 12) 2 | Io aie) 3 | 10 14 4 Io T2 5 12 12 6 10 8 a | pas) 8 8 10 12 9 12 12 10 38 To II 8 12 12 12 12 13 12 12 14 Io 12 15 | a4 L2 16 12 12 17 2 12 18 12 Io 19 12 14 20 | Io 12 ai | 8 10 22 Io Io 23 Io 8 24 12 Io 25 8 | 12 26 12 | 12 27 r2 To 28 12 ace) 29 Io ane) 30 8 10 31 Io 12 2 3 ears upper middle 8 | 8 | no grain 33 Io 12 | | | | | Numb f T Of | Class. Upper Ear. | Lower Ear. es ae of Buys in Sy each Class. - (eh tte aes As many rows in lower as “| ae oe | ‘) | a in upper {70 | 1 | 3) ies ts a ee 1) J | (12 T4 T) : : (exe) | 14 I) | es in lower than in (10 | ay 6) T4 (8 12 3) | (8 Io 3) Fewer rows in lower than { (12 0 4) \ in upper (10 8 3) yf 202 MATIZS¢ Tas_e XL (continued). SUMMARY. Upper Ear. | Lower Ear. | Number of Plants. | ( 8 I | 8 se) 3 | { 12 | | ( 8 3 | To 3 | Io +3 6 } ( 4 I | { 8 fe) | 10 4 ee 12 3 | ( I4 I = — Total 33 The total number of ears producing any given number of rows was as follows :— Rows. | Ears. | 8 II 10 2 T2 | 30 14 | Nees — | — | Total 66 A few ears are also met with in which the grains are scattered promiscuously, in mosaic fashion, over the ear (Fig. 84A), so that the number of rows can only be determined by cross-sectioning the cob. Experiments conducted by the writer show that the cross between an 8-row and an 18-row breed results in the produc- tion of an intermediate form in the F, generation. Of thirty- nine cross-bred F, ears examined, only two produced the same number of rows as either parent, while nearly 75 per cent produced either 12 or 14 rows, ie. more than one, and less than the other, parent; as regards these two row numbers, the cross and the reciprocal cross produced nearly the same INHERITANCE —IMPROVEMENT BY BREEDING 203 proportions, the 18-row ? x 8-row @ giving 71°5 per cent, CHAP. and the 8-row ? x 18-row ¢ 76 per cent. The cross was made between No. 904, an 8-rowed white dent, and No. g05, an 18-rowed yellow dent, both pedigree ears as regards colour and row numbers, which bred true when selfed (Aurtz- Davy, 27). The results obtained are given in Table XLI. Fic. 84.—Undesirable types for seed. A, Mosaic arrangement of grains. B, Failure to develop two pairs of rows: a 4-rowed ear. Eighty-nine ears from a commercial crop of Ladysinzth, studied by the writer, gave the following figures :— Row Classes. : 2 I4 16 18 Number of Ears ; : 29 40 20 Percentages. é . 32°58 44°94 22". CHAP. V. 204 MATIZ Ie TaBLe XLI. INHERITANCE OF ROW NUMBERS IN CROSS-BRED MAIZE. 1063 ex go5 (18-row 2) x gog (8-row 3) Yellow Grain with Light Yellow Caps. 1064 ,, do. do. Yellow Grain with White Caps. 1066 ,, go4 (8-row 2) x g05 (18-row GC) Do. 1067 ,, do. do. Do. 1068 ,, do. do. Do. Separating the two crosses, we have the following figures :— 18-row 2 x 8 row Jd Rows. | Parent Ear No. SS — ee 7 — a Total Lars. 8 10 ui) 14 | 16 | eae ve9 ee ee = any 1063 0 I | # > || 2 I 8 1064 oO 2 | | fo) fo) 6 Total . ; 0 al 8 2 7 14 Per Gent) -. fe) 214 57°1 I4°3 72 = 100 es = Meet begs cn | ec 8) ee 8-row 2 x 18-row rf Rows. | | Parent Ear No. 5 SS aie ars —— | Total Ears. | | 8 10 | 12 | T4 16 | | se lecse ent | | 1066 I 2 3 I oO 7 | 1067 oO 2 4 fe) 9 T1068 ng fo) | 3 5 fe) 9 | | Total . 2 4 ro 9 fe) 25 | | Per Cent . 8 16 | 40 36 oO 100 | | = : : : <= Summary. | Rows. | | | Parent Ear No. ae ii ye ee oh Ota Bary, 8 10 | 12 14 16 18 x 8 o (| 3 8 2 I | 14 8 x 18 2 | 4 1%) 9 oO | 1 25 | Total . : 2 Caen hf eres Tt || | | | 39 | Per Cent . BPE | wr | 462 28°2 | 2°6 | 100 —— | | INHERITANCE—IMPROVEMENT BY BREEDING 205 East and Hayes find that two distinct kinds of irregularity of row numbers occur: one a physiological fluctuation which is not inherited, and one a definitely inherited character, or possibly a set of characters, “The non-inherited fluctuations are always present, while the inherited irregularity may be present or absent. The latter kind has been isolated in several varieties [breeds], the most con- spicuous being the Country Gentleman [sweet] corn. . . . Since the inherited irregularity can only be distinguished from the fluctuation by breeding, and then with difficulty owing to the obscuring effect of the latter, it is difficult to come to any conclusion regarding the method of its transmission when dealing with mixed strains. It could undoubtedly be deter- mined by careful work with a cross of which Country Gentle- man formed one of the parents. We have not made such a cross, but observations of large commercial cultures of Country Gentleman \ead us to believe that irregularity is a Mendelian dominant, although it may not act as a simple monohybrid. ... The one fact that stands out clearly is that 7f the per- centage of irregular ears increases much over 4 per cent in a commercial progeny row culture, the whole culture must be dis- carded to eliminate the undesirable ‘ blood’.” The results obtained by them, to the F, generation, are shown in Table XLII following :— TaBLE XLII. INHERITANCE OF ROWS IN A MAIZE CROSS. Row Classes. Rows No. Gen. of aa ] 7 a Fes —— Parents. | | ( 8 | 10 | 12] 14 | 16] 18 | 20 | 22 | 24 mee |e sell eee ed ea beer | i No. 8dent . ae 12 —= | 34) §4)) 36) 22] 2 | No. 54 sugar LP 8 89 | 25 7 | | No. 8 x 54. Fi 12 I 6) 14 | | 8 x 54) -—1 F, 12 g | 22!| 16] 1 ries Hy) or |) on) g]-a6| a) | ese Pa welled | | (i as -1i1-2.|F, 8 20 | 38 | 50 | | (eke ' -1-2a|F, 10 61 | 48 | 54 | | (Sen Eset be 10 32] 75 | 15 | (, )-1r-3a]|F; 8 § | 20 | 2 I | (Se Diese Serge | as; ore — | 33 |158| 26 | 3 | ( , )-1-6.]F, 12 4|36 |1og| 8] 2 | ( 4 )-—1z-—10 | F, 8 Very irregular, mostly 8-rowed | (Gj) = 73) | B 10 96 | 43 8 | | | | | | | CHAP. V. CHAP. 206 MAIZE 158. /vur-rowed Ears.—The writer has occasionally found in both flint and dent breeds, ears with only 4 or 6 rows (Fig. 848). The occasional presence at the base of the ears of the beginnings of additional rows, suggests that they are cases of failure to develop certain pairs of rows. East and Hayes associate the condition with the tendency commonly met with in ears of 8-row flint breeds, to split at the base into 2- rowed sections. Their investigations indicate that the tendency to the abnormality is dominant to the normal condition. 159. Inheritance of Fasciated and Lobed Ears,—F lattening, Fic. 85.—A, Fasciated ear. B, Lobed ear. fasciation, and lobing of the tip of the ear (Fig. 85.\ and Bb) are not infrequently met with, and flattening and fasciation are more common in some races than in others; in the Transvaal flattening is common in strains of Chester County. East and Hayes have studied the inheritance of this ab- normality and find that it is a dominant character, alternatively inherited, and that it is difficult to tell the pure normal ears by inspection, but that they appear to breed true when isolated. 160. Inheritance of Laterally Branched Ears.—\aterally branched ears (Fig. 86) are occasionally met with. Attempts INHERIT ANCE—IMPROVEMENT BV BREEDING 207 made by East and Hayes and by the writer to study the in- heritance of the tendency have not been very successful. But East and Hayes conclude that the character is transmitted and does segregate, for both normals and abnormals are pro- duced in the F, generation. Such ears are undesirable, especially as they produce irregular grain, and they should not be used for seed purposes. 161. Striped Leaves.— Races of maize occur, as, for example, Zea Mays var. japonica, in which the leaves are green, with white stripes (they are deficient in chloro- phyll), but which breed true, the striping being appar- ently homozygous. But East and Hayes record cases in which the striping indicates a hetero- zygous condition; the dominant form in this case is fully green. Plants with- out chlorophyll died, when only a few inches high, from lack of the power of assimilation ; these were considered by the authors as “probably homozygous recessives ”. 162. Difficulties En- countered in Studying In- heritance tn Matze-—The investigation of the inheri- Fic. 86.—Laterally branched ear. tance of characters in the maize plant is not as simple a problem as might at first appear. (az) The amount of pollen produced is so great that it is continually present during the flowering period, in the air of the maize field; it adheres to the clothes and hands of the breeder, or to the leaves and stems of the plant, and is easily CHAP. CHAP. V. 208 MAIZE transmitted from them to the silks of plants which are to be kept pure. In spite of all possible precautions, seeds of un- known paternal ancestry do creep into the cultures. With tassels bagged three days before any pollen was ripe, it was found that stray pollen was already present, and though old, a certain percentage was viable, although East and Hayes con- clude that the possible error from this source would be only one to about 10,000. But in the bagging of the silks there is also a chance of enclosing foreign pollen; the same investi- gators have found that about one ear in five would have one or more grains so crossed, even when the greatest care was taken. They conclude that in this work there is a_ possibility of an experimental error of five or six seeds out of the 200 to 800 produced on an ear; this is to be considered as a maximum and not the probable error, the latter being less than one seed per ear. As they well observe, however, “the determination of a probable error in a mass of data is not sufficient in genetic work; an actual error, in which a single seed of unknown paternity becomes the ancestor of a pedigreed line, is sufficient to upset all inductions drawn from the data”. (0) The small size of the chromosomes makes them diffi- cult to study. (c) And, finally, maize seed is rather delicate; when pro- perly matured and dried it remains in fairly good condition for only three seasons ; seed older than this is almost worthless, and there is even a possibility of the results from second year seed being distorted. All these factors add to the difficulty of carrying out in- vestigations. Methods of Plant Breeding. A man should be very careful in the selection of his parents. —HEINE. 163. 4 Few General Principles—In plant breeding it is necessary to carefully decide upon an ideal and to work steadily and persistently toward it. We must remember that it takes several generations of the plant to acquire and fix a desired character, and that any deviation from the original aim INHERITANCE—IMPROVEMENT BY BREEDING 209 may involve us in complications difficult to unravel. With our present knowledge of genetics, the safest course to follow is to work step by step, building up the new type, one character ata time, rather than to attempt to add two or three characters at once. Promiscuous or aimless crossing, and crossing which is not followed by rigorous selection, is worse than useless, for it spoils an established breed only to produce a mongrel race. Vacillation in breeding is equally unproductive; success is then mere chance, and we work like men lost on the veld, wandering sometimes forwards, sometimes back on our tracks. In order to breed intelligently and to good purpose, it is necessary not only to know what we want, but also how to attain it, which involves a close and thorough study of each breed. In the case of maize, our ideal should include not only the colour and shape of the grain and ear, but also the average yield of grain from each ear, and the average stand of plants per acre. Briefly, we may say that there are three things essential to the development of pedigree stock, whether of animals or plants: (1) start with the best stock you can get ; (2) propa- gate only the best (which implies also the elimination of the unfit) ; (3) improve by crossing, when you know how to ob- tain and fix the desired character. 164. Methods of Plant Breeding. ene methods employed in the breeding of plants are much the same as those used with domestic animals—horses, cattle, sheep, pigs, or poultry. The fundamental point, after the determination of the desired type, is the continuous mating (i.e. without interruption) of those parents, both male and female, which most nearly approach that type. In the breeding of plants three principal courses are followed: (1) selection, (2) cross-fertilization, and (3) hybridi- zation. Inbreeding necessarily follows any one of these three methods. Selection may be roughly defined as the choice of suitable parents for the production of a strain of the desired type. They may belong to the same or to different breeds. Selec- tion implies that there is a choice of characters to select from. It is necessary to resort to selection and inbreeding to 14 CHAP. 210 MAIZE: obtain pure strains from mixed ones, and to propagate pure strains when one has them. By rigid selection we avoid the production or propagation of new forms ; by cross-breeding we encourage it. By cross-fertilization we mean crossing plants of different “varieties,” breeds, or races of the same species as, for instance, two kinds of maize. This will be discussed more fully a little farther on. The strict definition of the term /ybrzd7zation implies breeding from two parent plants belonging to different species or genera as, for instance, wheat and rye. It is of little or no practical importance in the improvement of farm crops, and need not be further discussed here. 165. Selection of Parents—The first step in breeding is to secure well-bred stock. This does not always mean the im- portation of fresh strains, unless the latter offer decided im- provement over the old; if the breeds already acclimatized in the country are satisfactory as regards breed characteristics, it is sometimes better to use them as a basis for improvement than to rely entirely on something the adaptabttity of whitch to local conditions has not been proven. As Hartley (5) tersely puts it: “Under extremely difficult conditions of growth, teosinte and the buffalo will thrive better than im- proved types”. But local strains are often so mongrel (ie. heterozygous) in character that it would take years of patient toil, and much expense, to breed them pure; in such cases we must rely on the importation of fresh stocks, Whichever course we follow, we must select the parents best fitted to produce the desired type of offspring. Sheep farmers, who start with a highly-bred stud flock of acclimatized animals, do not take long to build upa large flock if they are good sheep farmers. But the best of them find it comparatively slow work to grade up a mixed flock. So it is with maize. If we can start with pure, high-bred, acclimatized seed, we gain enormously in time, for all that is then necessary is to maintain the purity, and continue the improvement already started. But at the present time it is impossible to meet the demand for pure high-bred seed, acclimatized to each maize- growing district in South Africa. In the selection of parents care is taken that both are as INITERITANCE—IMPROVEMENT BY BREEDING 211 true to type as possible. The usual result of mating like with like, is to produce like, provided always that the strains are pure, the last point is essential. There are exceptions to this rule, due to the interaction of other factors (% 131) which we need not discuss at the moment as they do not affect the principle; we refer to such cases as the crossing of two white-grained breeds of maize, which sometimes results in the production of purple grain; and the crossing of two dwarfs, which in some instances results in the production of tall plants. It is equally important to remember that by crossing the unlike we usually produce unlike, at least in the second generation. Selection can carry us to a certain point, but no farther; it isolates characters and eliminates the unfit, but it cannot add characters which are not there. This is accomplished by cross- breeding. ‘‘Any permanent improvement that is made by selection is merely the separation of one of the extreme biotypes. When an extreme line is entirely separated, however, selection of zts extreme fluctuations causes no change (or at least no permanent change) of type, because there is almost complete regression to the mode of its line” (Zasv, 2). 166, Effect of Inbreeding.—Strict inbreeding gradually leads to the isolation of the homozygous type, as was pointed out by Mendel, and as is demonstrated in the following diagram, for which the writer is indebted to Professor Punnett :— DR (Heterozygous Parent, inbred) Des af | ; - ae F, 4 DD selfed 38 DR selfed 4 RR selfed F, 64DD 32DD 16DD 32DR 16RR 32RR 64RR This gives, in the F, generation, 112 DD (pure extracted dominants), 112 RR (pure extracted recessives), and only 32 DR (heterozygotes). This scheme supposes that each plant of the progeny produces but four offspring and that chances 14* CHAP. CHAP. Ne 212 MAIZE are equal for the development of progeny of homozygous and heterozygous parents. But as Shull (1), East (2), and others have pointed out, in maize “self-fertilization, or even inbreeding between much wider than individual limits, results in deterioration”. Again: “ Although a study of the injurious effects of self-fertilization was not the aim of the investigation, it was immediately ap- parent in the smaller, weaker stalks, fewer and smaller ears, and the much greater susceptibility to the attacks of the corn- smut (Uszlago Maydis). The results were almost as marked when the chosen parents were above the average quality, as when they were below it, which in itself refutes the idea that the injurious effect is due to the accumulation of deficiencies possessed by the chosen parents” (S/z//, 1). “Inbreeding in maize gives the same effect as lack of nutrients, while cross-breeding gives the opposite effect. There is retardation or acceleration of cell division, respectively. It is an established fact, although the cause is unknown, that crosses between nearly related types are more vigorous than either of the types alone” (Zasv, 2). Collins (2) calls attention to the fact, however, that while it is fully recognized that isolating the pure strains or biotypes will very greatly reduce their vigour and yield, yet by making a combination of the proper strains, so isolated, it is believed that the degree of fertility of the cross will reach that of the most productive plants in the original mixed strain, and that an increase of the total yield can be obtained in this way. He also quotes an experiment of Dr. Shull, in which two self- fertilized strains which were separated from a common stock in 1904, and continuously self-fertilized since that time, were reciprocally crossed in 1907. In 1908 the yields of these re- ciprocal crosses were compared with each other, with the self- fertilized plants, and with cross-bred stocks of the original breed. The yield from the cross-pollinated seed was 30 per cent greater than that from the self-pollinated ear, and 2 per cent greater than the average of the original cross-bred stock. By this means it is found possible to isolate a homozygous type, the individuals of which, when mated, are as vigorous and productive as the original mongrel heterozygote, but with- out its objectionable features. INHERITANCE—IMPROVEMENT BY BREEDING 213 Inbreeding, then, has no permanently injurious effect on the breed. 167. Improvement in Vield by Use of First-generation Crosses.—The facts mentioned in the last paragraph indicate the possibility of utilizing the added vigour gained by crossing. The suggestion of making practical use of this fact was made by Prof. W. J. Beal, as long ago as 1876, but no advantage appears to have been taken of the idea until quite recently, when it was again brought forward by Shull (1), East (2), and Collins (2). Increases of 51 per cent over the normal crop have recently been obtained in this way in the United States. In the case of maize the beneficial effect is noticeable mainly in the first season, and is said to disappear gradually. In the case of wheat, Professor Biffen finds it applies only in the F, generation. The principle, therefore, involves a new cross each year; this fact accounts for the loss of vigour and pro- ductiveness which new breeds often show when grown on a commercial scale. Difficulties in the use of first-generation crosses in farm practice have been met with, but will doubtless be overcome in time. 168. Fundamental Points of Seed Selection —The funda- mental points in seed-maize selection are those which affect yield and quality ; briefly they include :— 1. Depth of grain ; Shape of grain ; Thickness of grain ; Narrowness of sulci ; 5. Length of ear. The following points are of lesser importance, but should not be neglected :— 6. Shape of ear ; 7. Straightness of row; 8. Regularity of grain in the row. g. Covering and regularity of the butt ; 10. Covering and regularity of the tip ; 11. Thickness of the cob. 169. Correlation of Characters.—It is well known among breeders both of plants and animals that certain characters in an individual plant are more or less related to each other, and fw iS) CHAP. V. 214 MAIZE are inherited together. When one of such characters is present in an individual, another character is almost certain to be present which is correlated with it. These correlations may be of several kinds; Webber (3) divides them into four groups, which he has termed Environmental, Morphological, Physio- logical, and Coherital. By environmental correlation he means to indicate relation to physical conditions or environment, such as to soils of vary- ing degrees of fertility. Such correlations include increase in number of grains with increase in height of culm, etc. They “are merely the expression for equality or conformity to con- dition of luxuriance. Strictly speaking, these are not corre- lated characters, and their consideration is of little or no value to the breeder.” He defines morphologica! correlations as those cases where a variation in one character is the primary cause for variation in another character, e.g. where the relationship between the characters is similar to that which exists between size of germ and oil-content of maize-grain. Physiological correlations include such cases as the reduc- tion in yield of fruit and seed in inverse ratio to excess of leaf-production, as in some races of tobacco, or of wood as in the case of certain fruit-trees. Coherital correlations include ‘those characters which are not related to each other in any direct or causal sense, but which are inherited as single unit-characters”. Such cases include the naked grain of certain races of oat correlated with large number of flowers in a spikelet. It is of practical importance to the breeder to understand the correlations of the characters with which he deals. It is essential to a proper selection of parent plants that he should not only pick out those bearing good ears, but that he should also study the habit of growth of the plant, its stem, leaf, and flowers, for these have an important influence on the produc- tion of good grain, and their precise relationships should be accurately determined and defined. “As yield is the character of paramount importance, and as this character can now be determined only by laborious field tests, it is of the utmost importance that careful considera- tion be given to plant characters that may be correlated to INHERITANCE—IMPROVEMENT BY BREEDING 215 yield. Discussions along this line have been almost wholly CHAP. confined to characters of the ear. A careful tabulation of yields as compared with other ear characters, covering six years’ work with four varieties, embracing in all more than 1,000 ear- to-row tests of production, indicates that no visible characters of apparently good seed-ears are indicative of high-yielding power. It is reasonable to expect, however, that a careful study of the entire plant in connection with its environment will reveal such characters” (Hartley, 5). plant, and corresponds to the individual animal, the form and size of which are so carefully selected by stock-breeders. For a grain-maize (i.e. apart from the question of ensilage), a desirable stalk should have no suckers or off-shoots, should have well-developed roots, be thick at the base and gradually taper to the top, and bear a good ear; this should be a little below the middle point of the stem to reduce the danger of . blowing down ina strong wind. For the same reason the stalk should not be too high; even in the Low-veld of South Africa it is doubtful whether a height of more than 8 feet is desirable. 171. Destrable Leaves—To produce the large amount of starch which is stored in a full ear, a large leaf surface is necessary ; 14 to 16 blades is a good number, and, on well- grown plants, the blade of the middle leaf should be from 4 inches to 6 inches across. 172. Destrable Ears.—The shank of the ear should not be more than 4 inches or 5 inches long ; individual plants pro- duce shanks of g inches to 12 inches, which is an undesirable character. An ear of cylindrical shape, well rounded at each end, gives the largest percentage of grain to cob; its grains are also more uniform in shape. The number of rows should be uniform and typical of the breed (8, 10, 12, etc.); the rows should be straight and with little space between; the grains should fit together compactly and be firm on the cob, and should be uniform in shape and length on all parts of the ear. It is sometimes suggested that it would be desirable to save seed from plants bearing two or three ears, in order to develop a more productive race. Experience shows, however, that with most breeds it is preferable to grow one good ear CHAP. V. 216 MAIZE on a plant rather than two medium or poor ones , few plants seem able to develop two really good ears, and much energy is wasted in the attempt, which might better be devoted to the production of one good ear per plant. The weight of the husked ear can be taken as a fairly good guide to the relative yield of grain. In some breeds, however, the heaviest ears do not a/ways give the greatest weight of grain, though such exceptions seem rare. Certain ears of Wisconsin x Iowa Silver-mine, weighing 10°95 ozs., gave 8°64 ozs. of grain, while those of Yellow Hogan, weighing only 10°80 ozs., gave 9°20 ozs. of grain, the difference in percentage of grain to ear being 5°81 per cent in favour of the Vedlow logan. Other similar cases have come under the writer's observation. It is quite possible, however, that such differences are not constant. American experiments show that the pro- duction of a large number of well-bred but medium-sized ears is more profitable than the attempt to produce abnormally large ears. In endeavouring to improve the yield by breeding, we may start with the moderate aim of a 75 per cent stand and an average of 8 ozs. of grain per ear. If this is attained it will give 6,534 plants (planted 40 x 18 inches), and 16 muids of shelled grain per acre. Some local breeds show strong tendency for the sheath of the ear not to cover the end of the cob. This is a bad de- fect, and should be bred out. It leaves the uppermost ovules and silks exposed to weather and insects, with resulting loss of grain. The most desirable character of a seed-ear is its power to reproduce abundantly a good quality of ears, but this can only be finally determined by comparative growing tests. 173. Desirable Cobs.—“ Selecting for small cob results in reducing the size of the ear, and it is also an easy matter to reduce the size of the cob to such an extent that the pressure of the kernels causes the ear to break” (Hartley, 5). But it is important that the cob should not be too thick, or it will not dry out quickly, when it is apt to discolour the grain. 174. Destrable Grains.—VYo again quote Hartley (5): ‘Length is a very desirable character for the grains of maize INHERITANCE—IMPROVEMENT BY BREEDING 217 to possess, as it is by increased length in proportion to the diameter of cob that the percentage of grain is increased. Soft, chaffy grains, though long, or grains with prolonged chaffy caps, are not desired. It is much better to select for increased length of kernel than to select for small cobs.” The most desirable shape of grain, he adds, is that of a wedge hav- ing straight sides and edges. This shape admits of the grains fitting together so compactly that little or no space is wasted. “The germ, the most nutritious portion, and the portion in which is located the embryo plant, should be large, smooth, and firm.” This American ideal excludes such broad types of grain as the broad Natal Hrckory King, and yet Hickory King is con- Fic. 87.—A, Desirable shape of grain. B, Device for standardizing grains. sidered one of the best selling types on the local South African market. But it does not necessarily follow that Weckory King is the most profitable type to grow. We need further informa- tion on several points in this connection before we can form a definite opinion as to the breed that will pay best. //fwe can get the best price, combined with good yield, from the broad Hickory King, that may be the best type for us to grow. But if we get, say, Ios. a muid for Hzckory King, yielding 12 muids per acre, and only 9s. per muid for some other variety yielding 15 muids per acre, it will pay better to grow the gs. variety, for it will sell for 15s. per acre (or £7 IOs. per ten acres) more than the Hickory King. 175. Fancy Points.— There are some points made use of in judging maize at shows which are good in their way, but which CHAP. CHAP. V. 218 MAIZE are not known to be of practical value in the selection of parent ears for breeding purposes; these include such fancy points as well-covered tips, perfectly straight rows, very thin cobs, etc. On this question we may think over the words of Dr. Hopkins of the Illinois State Agricultural Experiment Station :— “There is some danger of corn breeders making too much of what might be called fancy points in selecting seed ears. We would learn the facts which are facts and not base our selections too much upon mere ideas and opinions. For example, it is not known that ears whose tips are well filled and capped with kernels are the best seed ears. Indeed it is not improbable that the selection of such seed ears will cause the production of shorter ears and a reduced yield per acre. It is true that the percentage of shelled corn from a given ear is the greater, the greater the proportion of corn to the cob, but our interest in that percentage is very slight compared to that of yield per acre, and perhaps for the greatest possible yield of shelled corn per acre it requires that the ears shall have good-sized cobs. Possibly the corn which shall ultimately surpass all others for yield per acre will have tapering and not cylindrical ears. These are some of the points regarding which men have some ideas and opinions, but as yet we have no definite facts and we shall need several years more to obtain absolute knowledge regarding some of these points. Let us base our selections of seed-corn first upon known facts and performance-records, and secondly upon what one may call his ‘type’ of corn.” 176. Methods of Selection,—The attempt to practise plant breeding without sufficient knowledge of either the science or the practice has led to disappointment and failure in many cases. One mistake has been the buying of prize bags of shelled seed at agricultural shows, irrespective of the pedigree of the seed. Now prize bags of shelled maize generally con- tain the largest grain the farmer is able to find; the largest grain is often borne on the smallest ears and therefore does not represent high crop-producing power; the largest grain does not always produce good ears and good yields. Prize maize may have been sifted from bulk grain, shelled in the field, and often consists of the tailings which pass over the riddles ; it is sometimes produced by a very indifferent crop! Again, too much reliance has been placed upon seed-maize INHERITANCE—IMPROVEMENT BY BREEDING 219 taken from the biggest ears from the bulk crop. Big ears are more likely to reproduce their kind than big seed, and this is better than no selection ; but the farmer soon finds that it does not bring him beyond a certain point. This is because the ears in the ordinary bulk field have been cross-bred with inferior strains, In an ordinary commercial field of maize the pro- portion of good, typical ears is very small. Not long ago the writer went through a South African field of what looked like a good maize crop, running probably ten bags to the acre. In the course of about an hour he could find only two plants, in the best parts of the field of 15 acres, which could be con- sidered ideal for seed purposes. And though this was much superior to the ordinary crop of the country, he could not find 100 plants in an acre that were worth picking for seed. An acre carried over 8,000 plants, and at this rate there would be 80 poor plants to every good one. As the maize plant is usually not self-pollinated, but depends on cross pollination, it is probable that every plant in that field was crossed with pollen from one or more of the many poor plants with which it was surrounded. Deterioration in the quality of the seed produced must inevitably have followed. Such deterioration might not show in the ear produced that year, but it would show in the following crop. Deterioration is constantly taking place where breeding is not practised. It is not only big ears that produce heavy yields. A big ear is better than a little one, but big ears often produce small, light grains; medium ears with deep heavy grains usually produce the heaviest yield per acre. In practical plant breeding, three processes are usually followed in the selection of maize for the breeding plot :— (a) Field-selection of original mother plants ; (6) Selection of ears in the seed-store ; (c) Selection by continuous performance-record in the breeding plot. 177. Linportance of Care in Selectton.—Selection should not be done carelessly, nor be left to the ignorant. Hand picking by Kaffirs may be better than no selection, but it will not carry forward the work of improvement. One _ season’s careless handling of the seed crop may undo all the good which has been accomplished in three or four years. The man who is CHAP. CHAP. 220 MAIZE selecting must know thoroughly what points to select, and this knowledge depends on an intimate acquaintance with the laws governing the transmission of characters. The day of empiri- cal selection has passed; it can no longer be left in the hands of the ordinary farm labourer. 178. Field Selection of Parent Ears.—\f we select only from among harvested ears, we cannot tell whether the mother plant was vigorous or weakly, tall or short, leafy or sparsely leaved, subject to rust or rust-resistant, or whether a particular ear has grown at the right place on the stem, or has had a desirable shank. All these points and many others have a definite bearing on the future yield of the crop to be grown, for they are correlated with characters directly concerning yield. Field selection is obviously most important. Good ears cannot come from poor plants any more than good wool from poor sheep! And the one is as likely to pro- pagate its kind as the other. A sheep witha poor constitution would not be used in a good stud flock. No more should the grain from a maize plant with a poor constitution be used for seed purposes, for it will not produce a heavy crop. It is not sufficient to select all the largest ears at harvest to be used for the seed plot. The large ear does not always come from a plant desirable in other respects. In breeding for wool, a sheep farmer does not base his selection of his stud sheep solely on the amount and quality of the wool. A shapely body and robust constitution also take an important place in the list of characters which make up a desirable stud sheep. Plants like animals are living things with varying degrees of vigour. As with live stock, it is important that we make a similar study of desirable points in selecting our parent seed plants. In order to produce good crops we must begin with the mother plant in the field, and that plant must be vigorous, must have plenty of leaf surface, produce large ears, and possess other qualities correlated with the characters which our standard demands. Having in mind the standard of stalk, leaf, ear, and shank, it is necessary to select 100 to 500 plants which come as close as possible to that type ; these should be marked conspicuously so that they will be found at harvest time. A field of from 15 acres to 30 acres should be chosen for this purpose. The INHERITANCE—IMPROVEMENT BY BREEDING 221 time when the selection is made will depend partly on the CHAP, object sought; if this be early maturity it will be desirable to go through the field when the frst ¢assels and silks appear, marking all the earliest plants, provided they are desirable from other points of view. It will also be desirable to repeat the process when the first plants begin to ripen, because it does not seem to be the case that the earliest plants to mature are always the earliest to flower; this point needs further in- vestigation, however. For ordinary selection for yield and quality the best time is probably when the ears are well de- veloped, and before the leaves have turned brown; at this stage the breadth and colour of the leaf can be observed to good advantage. By systematically walking through a field, row by row, and tying labels on the desirable plants, it does not take long to mark off 500. The principal points to be observed in field- selection are :— General vigour of the plant. Leafiness of plant and width of leaf. Size of ear. Straightness and strength of stem. Stem broad at base, tapering gradually. Ear borne about middle of stem. Shank of ear short. Husks compact and firm on ear. . Apex of ear well covered with the husk. 10. Freedom from rust and smut. NOOO ae Where early maturity is desired, as on the extreme High- veld, this can also be taken into account. Each selected plant should be marked with a conspicuous label which will not be lost sight of when the leaves turn brown at harvest. For this purpose sized cloth labels prove least satisfactory ; they quickly blacken, lose the “size” and become indistinguishable, losing entirely the figures written on them. Our best results have been obtained with ordinary brown paper parcel-labels, numbered with ordinary black or blue pencil. But even these are too much like the dry maize leaves and husks in colour to be easily seen at harvest ; and when a label is tied near the tassel away from the leaves, CHAP. 222 MAIZE both tassel and label will disappear in many cases by the breaking off of the tops of the culms. At harvest all the marked plants that can be found are cut by hand and removed before the rest of the crop is harvested. The ears. are hand-husked and stored until they can receive personal attention. By that time they should have dried out thoroughly, so that reliable comparative tests of weight can be made. The ears should be allowed to ripen well on the stalks; the stalks of the selected plants might be harvested and shocked by themselves to avoid delaying the rest of the harvest. When thoroughly dry, careful selection of the ears must be made, only forty or fifty of those which come closest to the ideal .being retained for the breeding plot. These should be weighed separately, and a record kept of the total weight and of the weight of shelled grain from each. 179. Seed-room Selection of Ears.—The 500 ears selected in the field are weighed in bulk and then laid out on benches in the seed-room. All small, distorted, or otherwise undesir- able ears are at once discarded, their total weight being taken, as a check. Field selection cannot be so perfect that none but desirable ears will be harvested. In a test case of five breeds selected in the field, the following proportions were retained as suitable for the breeding plot :-— Per Cent. Yellow Hogan . : : : : . : 80 Yellow Horsetooth . . : : ; ? 65 Hickory King . 3 i ; : ; 5 61 Golden King . . 4 2 é * . 59 Lady smith . ; : : é ‘ : 56 It is doubtful whether in ordinary selection of large ears at husking it would be possible to obtain even 10 per cent of desirable ears from an ordinary crop. And then one would not have the advantage of knowing that they came from robust and otherwise desirable parents. Of the 300 or so ears left, a more critical study is made. One hundred of the best are reserved for the centre of the breeding plot and the remainder shelled off at once to be used for the end rows. After some practice the selection of the best 100 ears can easily be made by eye. When increased yield per acre is the INHERITANCE—IMPROVEMENT BY BREEDING 223 primary consideration, the points on which selection is made CHAP. can be reduced to the following :— “ Size and weight of ears; Depth of grain ; Closeness of rows (i.e. narrow sulci) ; Regularity of rows ; Regularity of grain ; Shape of ear, and character of tips and butts ; Yield of grain per ear. NOUR WYN Fic. 88.—Selecting seed-maize: the final selection. All these points have a direct bearing on the yield per acre. The final selection consists in classifying the picked 100 ears into groups of ten each, according to depth of grain and size and weight of ear, etc. (Fig. 88). The weights of each group of ten are then taken; they are arranged in a row, with the twenty best ears in the centre, the next best next, and so on, the poorest of the 100 occupying the two ends of the row. In the final selection too much reliance must not be placed on weight or sizeof ear. The writer has frequently found that CHAP. V. 224 MAIZE the smaller of two ears gives the greater weight of grain. This is due to the fact that there is an immense difference in the depth of the grain on different ears of the same variety. Other things being equal, the greater the length of the individual grains the greater the yield per acre. It is bulk of grain that is sought, not size of ear; the latter is important only as it aids to produce the former. It is true that the ear, as well as the vegetative characters of the plant, give an indication of its capacity; but it is the individual grain which carries the embryo plant, and as the grain is, so will its progeny be, except always for such changes as may be brought about by the influence of environment or cross-pollination. The ear must be taken into account only in connection with the grain that it bears. 180. Character of the Gratn—Therefore, after the sub- division by size, weight, and other external characters, it is necessary to examine carefully the character of the grains of each ear, and to re-arrange the ears in accordance therewith ; it is surprising to find how great a range of variation occurs in respect to the size and shape of the grain within the limits of one breed (see Fig. 68). This comparison is best accomplished by taking six grains from each ear, two from a point about one-third from the tip, two from the same distance from the butt, and two from the centre. These six grains are laid on the table at the foot of the ear from which they are taken. Comparison is then made of the grains from all the ears, especially in regard to uniformity, length, shape, thickness, and size of embryo. It is important to consider uniformity of grain, for if—as is often the case—the grains on the upper part of the ear are shorter than those on the lower part, the weight of grain must obviously be less than if they were longer. After careful study of the grains, the ears must be arranged accord- ingly, even at the expense of size and weight; in some cases a compromise may be made with advantage, but this can only be done effectively by persons having a thorough knowledge of the subject. In the case of the ten best ears of each breed, full notes are taken of the length, circumference, and character of each ear; samples of the grain from each are retained for reference the following season, in order to determine whether the char- NHERITANCE—IMPROVEMENT BY BREEDING 225 acters for which the ear was selected are being transmitted or whether they are only of a temporary nature. If any one or more ears show fluctuating variability as regards such char- acters, the progeny of those ears can be discarded next season. The object of placing the very best ears in the centre, and the worst of the 100 on the outside, is that the same sequence may be preserved in the breeding plot. By this means, and as the grain from each ear will be used to plant only one row of the plot, the grain from all of the best ears will be removed as far as possible from danger of pollination by plants de- rived from poorer ears. In this way the tendency to deteriora- tion, through cross-pollination with poor plants, is reduced to a minimum. After this the ears are shelled by means of a hand-sheller ; the cobs of each ten are weighed separately ; the weight of cob deducted from the weight of ear previously taken gives the weight of grain. Being in groups of ten the average weight of individuals in a group can be determined at a glance. Each ear is shelled into a paper bag (half-pound bags with folding flap have been found satisfactory) and num- bered with a consecutive number from 1 to 100, preserving carefully the same sequence as that of the final selection. The packets of each breed are then placed in separate boxes, the highest number at the bottom, and carefully labelled ready to be taken to the breeding-plot at planting time. This completes the work of selection for the season. 181. Selection by Continuous Performance-record. — This consists in the strict and continuous selection of parent ears from among the best progeny of the best plants which have year after year given the best performance-record in the direction desired (i.e. yield, early maturity, drought resistance, etc.). An idea of the method practised is given in Table XLIII following, which shows the sort of record kept each year of the performance of certain strains grown in the breed- ing plot of the Illinois State Experiment Station. Selection for the following year would be made of those ears which give the best record in this analysis. It is noticeable that the nine best ears! are those in the centre of the table, which come from the centre of the breeding plot, and that the best result of all 1 As regards protein-content, for which the ears were selected. 15 CHAP. CHAP. Ve 226 MAIZE is obtained from the centre row (No. 10). .This indicates inheritance of good characters, for in the breeding plot the best ear was planted in the centre of the row, the two next best on either side, and so on, to prevent crossing of the best with the poorer plants. TaBLE XLIII. PERFORMANCE-RECORD OF BREEDING PLOT, 1gor. ILLINOIS EXPERIMENT STATION. (Breeding for High Protein.) | Relative Weight | | | Field Row of Ear Cornin | No. of Ears per | Weight per Acre Protein Namber. Crop (Average of | Acre (on an |(on an Average of in 200 Plants per Average of 8,000). | 1 lb. per Ear). | Seed-ear. Row). | | | | | | Lbs. | No. Muids. Per Cent I ; ‘ gi‘o 7,280 | 182 12°06 2 ; al 86°0 6,880 17°2 | 12°17 4 AI 98°5 | 7,880 19°7 | 12°19 4 aH 99°5 | 7,960 | 19°9 | 1226 5 ‘ 77°0 | 6,160 15"4 12°31 6 I18'0 | 9,440 23°6 | 12°40 . 7 I16'0 ; 5250 23°2 12°66 4 8 | 54°5 4,360 10°9 12°83 | & 9 107'0 5,560 | 21°4 1290 | = 10 103'0 8,240 | 20°6 15°78 | # 11 870 6960 17"4 12°93 | | 12 127°5 | 10,200 25°5 12°94 | 8 | 13 1130 Q,040 22°6 12°72! | 14 123°5 | 9,880 24°7 r2"45 J ; 15 103°5 | 8,280 20'7 12°32 16 92°0 | 7,360 18°4 12°31 17 85°5 6,840 I7°I | 12°23 18 II7'0 | 9,360 23°4 | 12°18 19 140°5 11,240 28°1 12°07 20 97°0 7,760 - 19°4 12°06 ee ee eee | See ae | Average IOL'g | 8,148 20°37 12°59 182. Method of Propagation.—Difficulty has been experi- enced by growers in propagating the seed-maize after it has been selected, and some have been so discouraged by the results of cross-pollination, that they have given it up. It should be remembered that it takes three years, at least, before material improvement can be expected. The difficulty is largely due to the small amount of seed available as the result of the first selection. The following method has been found satisfactory. INHERITANCE—IMPROVEMENT BY BREEDING 227 First Year's Selection —Let us suppose that only twenty CHAP. good ears, coming up to our ideal, have been selected this Me Fic. 89.—Selecting the best ears from the bulk plot. season; what are we to do to prevent this progeny being crossed by plants in the same field grown from unselected seed? The best plan is to plant the seed in a square of, say, 15 * CHAP. V. 228 MAIZE 70 yards by 70 yards (an English acre), at one corner of the field where the bulk of the same breed is grown. This should be, preferably, on the side nearest the homestead or road, so that the plot may be watched. This is called the breeding plot (1 183). The rest of the field is planted with the best of the seed which is not considered suitable for the breeding plot. Second Year's Selection.—The three outer rows of the breed- ing plot (which have been most exposed to crossing with the plants in the bulk field) are harvested with the bulk crop. The rest of the breeding plot is harvested separately, and any nubbins or undesirable ears are rigidly discarded. From the remainder the twenty best ears are again selected and planted in the breeding plot. The remainder are used for what is known as the propagation plot, which may vary in size from five to ten acres or more, according to the amount of seed available. The propagation plot is planted around the breed- ing plot, and forms a buffer between it and the bulk plot. The best ears selected (Fig. 89) from the bulk plot are used to plant the current season’s bulk plot. Third Vear's Selection. At harvest time the three outer rows of the breeding plot are harvested with the propagation plot, and the three outer rows of the propagation plot are thrown in with the bulk crop. The remainder of the breeding plot is harvested separately, and any nubbins and undesirable ears are discarded; from the balance the twenty best ears are again selected for the next season’s breeding plot, and the remainder are used to plant the propagation plot. The remainder of the propagation plot is harvested sepa- rately ; nubbins and objectionable ears are discarded, and the best of the balance is used to plant the bulk fields. Thus in the third and subsequent years the bulk fields are supplied entirely from twice selected seed from the propagation plot, while the propagation plot is itself supplied from the breeding plot. In the fourth and subsequent years the work is continued as in the third year; it must always be maintained,-for though there is a maximum beyond which “selection” alone cannot carry one, deterioration takes place with remarkable rapidity when selection ceases. INHERITANCE —IMPROVEMENT BY BREEDING 239 The fundamental principle of the method described above is the rigid elimination of undesirable types, which may appear owing to reversion, and the mating together in the breeding and propagation plots of the most desirable types. 183. The Breeding Plot should be so selected that the soil will be typical of that on which the main crop will be grown. The same preparation should be given as for the main crop, no extra care or fertilizer being used. The object is to find out which plants will give the best results under normal con- ditions; if they do well, then they may be expected to do better on well-fertilized soils. It is absolutely necessary that the plot should be isolated at least 400 yards from any other sort, or from strains of the same sort, flowering at the same time. Any stray plants from previous plantings must be carefully rogued out before they have a chance to tassel. The rows should be 200 or more yards long and of exactly the same length. The seed should be planted on the principle of one row to each ear, After removing the tips and butts from the selected ears, the rest of the grain should be planted inasingle row. It is better to drill the seed than to check- row it, as it is difficult to isolate suckers from main stalks when more than one plant occupies a place. Each row should be numbered consecutively and labelled with a stake at the end. With the seed left over from the rows three or four border rows may be planted all round the breeding plot to protect the plot to some extent from depredation. 184. Devices to Prevent or Detect Cross-pollinatton.—As a means of minimizing the amount of cross-pollination between breeds of maize grown near to each other on similar soils, the following devices, among others, have been resorted to at the Botanical Experiment Station, Pretoria; none of them, how- ever, has been entirely successful. Planting one week and two weeks apart was tried. Several of the breeds tested were new to us, and their relative time of flowering in that climate was not known, so that some of the later-sown’ flowered earlier than those planted before them. The danger of cross-pollination was minimized by bagging and hand-pollination, but the number of plants of each that could be treated this way was small, and the amount of seed CHAP. V. CHAP. V. 236 MAIZE saved was therefore limited. It was also found that in some cases in the same breed there was a difference of nearly two weeks in the time of flowering. This experiment demon- strated clearly that close planting, even allowing two weeks between plantings, cannot be relied upon to prevent cross- pollination, By planting first an early-maturing breed, and two weeks later a late-maturing breed, a certain degree of immunity may be secured. But there would be little practical advantage in this, except where it is necessary to plant several breeds in close proximity, for the farmer usually desires to plant all his late- maturing maize first, and afterwards that which takes less time. The use of rows of sorghum and kaffir-corn to separate the different sorts has proved unsatisfactory on the Transvaal High-veld, the maize having come into flower before the sor- ghum was tall enough to afford any protection. Detasselling is perhaps the most satisfactory method of deel- ing with the problem. But there is a certain amount of fluctuat- ing variability as regards time of flowering, and variability will be still more pronounced where the strain is heterozygous for this character. In practice this means that detasselling must be repeated (in the same row) two or perhaps three times within a week or ten days, and that great vigilance must be exercised to detect individual plants which flower later than others, and prevent pollen scattering from their earliest anthers, Covering the plots with cheese-cloth was tried at the Government Experiment Farm, Potchefstroom, as a means of isolating the breeding plots, but was not found satisfactory. 185. Production of New Types by Artificial Cross-pollination. —The popular idea of cross-breeding maize is that by crossing two sorts the result will be a hybrid combining the characters of the two parents. This may be the case with the first ear obtained by the cross, but it is not always the case in the pro- geny of the first generation. Experience shows, moreover, that even where it does occur, the second generation from the cross produces a great deal of variation, more particularly with moncecious plants like maize, and unless scientifically guided efforts are made, this variation will continue from generation to generation for an indefinite period. We have already dis- cussed the reason for this mixture of characters, and how it may be avoided or made use of. INHERITANCE—IMPROVEMENT BY BREEDING 231 Cross-breds of unknown pedigree are difficult to deal with, CHAP. and it may take years before they yield any desirable progeny. They should, therefore, be avoided, and the work of improve- ment should be started with well-bred seed. Where it is desired to add new characters to a breed which does not already possess them, cross-breeding must be resorted to. The actual process of crossing is easy, but to isolate and fix the desired type is an entirely different problem. Crossing produces such varied heterozygous combinations that endless confusion results, and it requires knowledge of the laws of in- heritance, and infinite time and patience, to produce order out of chaos. Therefore cross-breeding should only be practised or permitted where the effect of crossing ts understood, the object sought ts well known, and the method well planned. 186. Reciprocal Crosses—Where it is desired to transmit a definite unit-character from one breed to another, it appears to be immaterial which breed furnishes the male and which the female parent; the results in the F, generation are usually the same in either case. 187. Method of Cross-pollinating.—Cross-pollination is a comparatively simple matter. The silks of the plant to be pollinated must be carefully protected from the access of any stray pollen (Fig. 90); and the pollen of the male plant must be carefully collected so that it will not be mixed with stray pollen of other plants in the neighbourhood. The pollen is then shaken on to the silks and the latter are again covered up until all danger from stray pollen is over. It should be remembered that the pollen is light and easily carried by the wind ; when the field or plot is in tassel, the air may be charged with pollen grains, so that difficulty is experienced in pre- venting contamination of the silks with stray pollen. 188. Collecting the Pollen—The tassel should be covered with a paper bag, an ordinary 2 lbs. grocer’s bag, of zw but tough brown paper, is found satisfactory. This is tied tightly round the stem below the lowest branches of the tassels. The bag should not be placed on the tassels until the first anthers appear on the terminal branch of the tassel, otherwise the anthers are apt not to develop properly. In the Transvaal, much of the pollen is found to lose its vitality after the third day. MAIZE Fic. 90.—A new breed of maize in process of development. (Courtesy of the South African Railways Publicity Department.) INHERITANCE—1IMPROVEMENT BY BREEDING 233 The tassels may appear before the silks, and sometimes even shed all their pollen before any silk appears (4 79). The appearance of the tassels is by no means uniform in individuals of the same variety. This is an important point in connection with the work of detasselling, for it makes it neces- sary to go through the breeding plot three or four times, at different dates, to effectually prevent self-pollination. 189. Covering the Sitks.—This is done by means of paper bags similar to those used for covering the tassel (1 188). The ear should be covered just before the silks first appear, to pre- vent contamination with stray pollen. The ear may be left for four days or a week before the pollen is applied, in order to allow all the silks to develop. For pollination, the bag is removed and a good dose of pollen shaken on to the silk, care being taken that it reaches all the silks. The bag is then re- placed quickly to avoid contamination with stray pollen. The first attempts at hand-pollination are not always satisfactory, but excellent ears may be obtained as a result of skill gained by experience and practice. The silks may appear either before or after the tassels. Not all the silks mature at the same time; those from the ovules lowest on the cob appear first. Sometimes fresh silks continue to appear over a period of seven days. A single hand-pollination, effected when the silks first appear, is therefore inadequate; nor is it sufficient to repeat it on two successive days; this results in the lower half of the cob being well-filled while the apex remains undeveloped. Three pollinations, at intervals of three days between each, generally prove the most effectual, but by this means greater risk of contamination is incurred. In some cases, and in the same breed, the silks appear be- fore the tassels. When such an individual happens to be the earliest to flower in a field, it may fail to develop seed; this tendency is therefore unlikely to be propagated to any great extent. 190. The F, Plants.—Three or four hundred grains will be obtained from a single successful crossing. If the parents differ in colour of endosperm, it will sometimes be possible to tell which grains have been crossed and which selfed acciden- tally, and the latter can be discarded. But F, seed should be CHAP. CHAP. Vi 234 MAIZE planted in an isolated breeding plot and each resulting plant should be carefully selfed. By this means it will be possible to detect the results of accidental crossing with stray pollen, and to prevent its spread to the other plants of the cross. The ears produced by these plants will show segregation in the seed, if the characters involved in the cross are seed characters, and from them selection of the desired grains can be made. 191. The F, Plants—The F, plants must also be grown by themselves, and selfed. If the desired character is recessive, it will be possible to isolate it and commence propagation. But as our knowledge of the individual unit-characters is at present imperfect, it is desirable to grow the plants on for another generation, and self them, in order to eliminate any undesirable character which may not have appeared. If the F, generation breeds true, the new type may be considered fixed and we may proceed to propagate. 192. [improvement by Breeding ts Slow at First—At best, improvement by breeding is a tedious process, and the man who is not prepared to be patient, methodical, and persistent, should not undertake it. The writer has known men who started out well, with no little expenditure of time and money, but who, seeing no visible results, gave it up in disgust after the first year. As has been said, visible results cannot be ob- tained during the first few seasons. It is probable that there is not a pure pedigree commercial crop of maize in South Africa to-day, and very few elsewhere, for maize is a remark- ably heterozygous mixture. Before we can hope to make definite progress, we must purify the strains we wish to im- prove. Hitherto time has been largely taken up with trying out breeds suited to different parts of the country. Now that we have formed definite ideas on this subject, we can devote our attention to their zzprovement by breeding. Addendum.— Arrangements have been made by the Agvv- cultural Supply Association, Limited, P.O. Box 1148, Johannes- burg, to supply pure-bred seed of heavy-yielding strains of maize especially suited to the High-veld of the Transvaal and Orange Free State. CHAPTER VI, JUDGING AND SELECTION FOR EXHIBITION. Send forth the best ye breed. —Kip.ina. 193. The Object of Exhibiting at Agricultural Shows.—The agricultural show does not exist solely nor primarily for the purpose of winning and awarding prizes. Unfortunately there are too many people who exhibit merely for the sake of prize- winning, having in view either the value of the prize itself, or the advertising of their seeds and other farm products, Those who take all the prizes year after year discourage others who have not equal facilities for preparing special exhibits, but whose work is, nevertheless, worthy of a prize; therefore the number of prizes which can be drawn by any one ex- hibitor in any section should be limited. The main object of the agricultural show should be educa- tional: the farmer should be able to learn from the exhibits (1) the need for, and the means of, improving his own crops, (2) the relative merits of new breeds, and (3) where to obtain good seed. He may not realize the need for improving his methods and seed until he sees that other farmers’ results are better than his own, and an agricultural show should be the best place for him to see this, We agree with the American writer who says that exhibits arranged with respect to the ready comparison of typical samples of different breeds, offer one of the most effective methods of diffusing knowledge with regard to the character- istics of different breeds. The main points enumerated in this chapter are already familiar to maize judges of experience, but are given here for reference, Owing to the short time usually allotted to judg- 235 CHAP. VI. MAIZE 236 ‘o161 ‘sinqsauueyof ‘moys snizig pue aZIe UBILYY YINOG IsIly OY} 3 ‘ainzNoIIsy Jo JusWIedag [eeasuLIy ‘AuRJOg JO UOISIAIG ay} JO WqQIyXY— ‘16 ‘OILY JUDGING AND SELECTION 237 ing at local shows, certain of the details can be taken into consideration only where competition is very close, 194. Rules Governing Maize Exhibits.—The following rules are based on experience gained at leading South African shows :— (1), Each entry must be accompanied by a certificate giving as nearly as possible the date of planting and date of harvesting of the crop and name of the district in which it was grown. These certificates must not be seen by the judge till after the judging. (2). No exhibit may be entered in more than one class. (3). An exhibitor may receive only one prize in any one class, (4). An exhibitor is barred from exhibiting in more than three classes in any one of Sections I to HI inclusive. This allows each exhibitor to show an early, medium or main-crop, and a late breed in each section. (At some American shows an exhibitor may enter only in ¢kree classes in all.) (5). Every exhibitor may enter for all classes in Section IV (special prizes), but may only take ¢wo przzes in this section ; should he obtain more awards he will have the option of choosing which two prizes he will take. All awards will count as points in the aggregate for the Grand Championship. (6). Where there is but one entry in a class a prize shall be awarded only if the judge considers the exhibit deserving of recognition. (7). In such a case the judge shall decide whether a first, second, or third prize shall be awarded. (8). Grand Championship —The Grand Championship Prize will be awarded for the highest number of points obtained by any exhibitor. Points will be given as follows: Com- mended, } point; highly commended, 4 point; third prize, I point ; second prize, 2 points; first prize, 3 points; champion- ship of first prizes in a section, 4 points extra. The last- named provision is made to prevent mere number of entries from scoring over quality of exhibit. An exhibitor A who makes three entries in each of the Sections I to III inclusive, and who obtains six first prizes, would score 18 points ; another exhibitor B who enters in only one class in each of the three sections may obtain the first prize and the championship in CHAP. VI. 238 MAIZE CHAP. each section; unless the championship counted for more VI. than 3 points, B would score no more points than A, al- though the quality of his exhibit was superior as evidenced by his taking three championships. The aim of agricultural shows should be to encourage guality rather than number of exhibits from any one exhibitor. (9). In the event of a tie, the judge must decide as to the general relative merits of the two tieing exhibits and award the championship to the one which in his opinion is the best. If taken in an absolutely mathematical sense, the counting of points may result in an injustice to the best exhibit. It is obvious that ifan exhibit wins prizes against severe competi- tion, it is more worthy of a championship than one which has no competition. (10). The exhibits must have been harvested during the twelve months immediately preceding the show. (11). Exhibits must not, be treated unfairly by removing poor, cross-bred, injured, or otherwise undesirable grains and replacing them by good ones. Any unfair or tricky occur- rences bar the exhibitor from all entries and all privileges of the show. One, or not more than three, grains may be removed by the exhibitor from one or more sides of each ear, in order to decide whether the ear is fit for exhibition. (12). ‘‘Grooming” of the ears in such a manner as to allow of their best possible presentation, is strongly recommended ; e.g. shanks of ears should be neatly removed with a pocket knife, and loose silks should be carefully taken off. (13). Exhibits must be delivered to the stewards of the pro- duce section two clear days before the opening of the show. They must be carefully labelled both zvzs¢de and outside of the bag or box, for it often happens that the outside label is torn off in transit, and the owner is then traced with difficulty. The inside label should bear the name and address of ex- hibitor, the date of forwarding, and the section and class in which the entry is made. It is best to tie this on to at least one of the ears, or inside the mouth of the sack of shelled grain. (14). All exhibits are subject to necessary handling by the judge, but remain the property of the exhibitor, and may be secured by him immediately after. the show is declared closed and the awards have been made. JUDGING AND SELECTION 239 (15). Professional maize breeders, seed dealers, or expert judges will not be allowed to compete except in classes specially arranged for them. (16). A bag of shelled maize shall weigh 203 lbs. gross. (17). A bag of ears must be contained in a full muid maize sack, and must weigh about 100 lbs. (18). An ear of maize is a cob with the grain still attached, but with the husks removed. (19). In classes in which the breed is not specified, each entry must be conspicuously labelled with the name of the breed, or the entry will be disqualified, and the name of the breed, as given by the exhibitor, should be printed in the show catalogue. 195. The Prise-list.—It is important that the growing of recognized standard breeds of maize be encouraged ; the offer- ing of prizes for specific named breeds is doing much to per- manently improve the maize industry of South Africa. The custom of allowing one recognized breed to compete with another in the same class (except for a championship) should not be allowed, except in certain special cases indicated farther on in this chapter. 196. Classtfication.—The proper classification of exhibits is essential to the educational value of a show, and to suc- cessful judging. In the classification of maize two main points should receive consideration :— (1) The exhibition of ears selected for seed, by which the would-be buyer can determine by whom and where the best seed maize is grown in his particular dis- trict, province, or country. -In this section there are usually two subsections, (2) the ten-ear and (6) the single-ear competitions. It is sometimes argued that this section has no value to the prac- tical farmer because “anyone can grow ten good ears in his back garden”. This, however, is not the case. To produce ten really good exhibition ears from a small plot is almost impossible, owing to the much greater danger of imperfect pollination, attacks by insect pests, etc. The ten-ear and single-ear competitions are essentially educational ; through them a farmer learns what to select for seed. CHAP. VI. CHAP. VI. 240 MAIZE (2) The exhibition of the commercial article—the maize- grain—by which the merchant and manufacturer are enabled to learn where and by whom are grown the best qualities for their particular classes of trade. Two distinct score-cards are required for the judging of these sections. 197. Sections. —The following sections are found suitable for South African shows :— Section I.—Shelled maize for market or export. One muid (203 lbs. gross) of shelled maize, each bag to be ac- companied by one full bag (to weigh about 100 lbs.) of ears from the same crop; these ears to be taken into consideration by the judge in making the awards. Section I].—Seed-maize; ten ears selected for the breed- ing plot. Section IfI.—Best single breeding ear. Section 1V.—Special prizes. As far as possible all of these sections should be repre- sented in every prize list, in districts where maize is a staple crop. 198. Classes—The following is a list of classes suitable for maize shows, It is usually only the central shows which are able to offer such a complete list as is here given; very few district shows need include all of the classes, for only a few of the breeds named are grown in any one district. SECTION I.—SEED-MAIZE: TEN EARS SELECTED FOR THE BREEDING PLOT. Entrance fee, 5s. per class. Prizes (in each class) : rst, £2; 2nd, £1 ; 3rd, tos. Cham- pionship (of all classes in this section), £3 »y ADs Dent Breeds (white). Class. 1. Hickory King (8-row). 10-row Azckory or Louisiana. 12-row Hickory or Hickory Horsetooth. Salisbury White, Masoe, or Brindette. Mercer. mk wind JUDGING AND SELECTION 241 Class. 6, Towa Silver-mine. 7. Boone County. 8. Ladysmith or Champton White Peart. 9. Natal White Horsetooth. 10. Any other white dent breed. (In this class entries ‘must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) Dent Breeds (yellow). 11. Eureka. 12. Yellow Hagan. 13. Chester County. 14. Reid. 15. Vellow Horsetooth or German Yellow. 16. Golden Beauty. 17. Minnesota Early. 18. Golden Eagle. 19. Any other yellow dent breed. (In this class entries must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified. ) Flint Breeds (white). 20. Any white flint breed. (In this class entries sust be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) Flint Breeds (yellow). 21. Yellow Botan. 22. Yellow Cango. 23. Wills Gehu. 24. Any other yellow flint breed. (In this class entries 26. must be labelled with the name of the breed or the exhibit will be disqualified. ) Flour corn or Bread-mielies. Brazilian Flour corn or South African Bread-mtelve. Sugar maize. Any breed of sugar maize. (In this class entries must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified. ) 16 CHAP. -ViI-L 242 MAIZE CHAP. Pop-corn. VI. Class. 27. Any breed of pop-corn. (In this class entries st be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) SECTION IJ.—BEST SINGLE BREEDING EAR. Entrance fee, 2s. 6d. per class. Prizes (in each class): ist, £1; 2nd, toa; 3rd, 5s. ; Champion Ear (the best ear of all the first prize ears), £2. Dent Breeds (white). 28. Hickory King (8-row). 29. 10-row Aickory. 30. 12-row Hickory or Hickory Horsetooth. 31. Salisbury White. 32. Mercer. 33. Lowa Silver-mine. 34. Boone County. 35. Ladysmith or Champion White Pearl. 36. Natal White Horsetooth. 37. Any other white dent breed. (In this class entries must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) Dent Breeds (yellow). 38. Eureka. 39. Yellow Hogan. 40. Chester County. 41. Reid. 2. Yellow Horsetooth or German Yellow. 43. Golden Beauty. 44. Minnesota Early. 45. Golden Eagle. 46. Any other yellow dent breed. (In this class entries must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified. ) JUDGING AND SELECTION 24 Ww Flint Breeds (white), Class. 47. Any white flint breed. (In this class entries aust be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) Flint Breeds (yellow). 48. Yellow Botman. 49. Yellow Cango. 50. Wells Gehu. 51. Any other yellow flint breed. (In this class entries must be labelled with the name of the breed, or the exhibit will be disqualified. ) Flour corn or Bread-mielies. 52. Brazilian Flour corn or South African Bread-mielie. Sugar maize. 53. Any breed of sugar maize. (In this class entries szs¢ be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) Pop-corn. 54. Any breed of pop-corn. (In this class entries must be conspicuously labelled with the name of the breed, or the exhibit will be disqualified.) SECTION III.—SNELLED MAIZE FOR MARKET OR EXPORT. One muid (203 lbs. gross) of shelled maize; each entry must include one full bag (to weigh about 100 Ibs.) of ears from the same crop. These ears will be considered in judging the sample. Entrance fee, 5s. per class. Prizes (in each class): Ist, £3; 2nd, £2; 3rd, 41 ; cham- pionship (of all classes in this section), £5. White “Flat”. 55. Hickory King. 56. Natal White Horsetooth. 57. Any other flat white dent breed. 16 * CHAP. VI. CHAP. VI. 244 MAIZE Yellow “ Flat”. Class. 58. Any flat yellow dent breed. White ‘“ Round”. 59. Any round white breed. Yellow “ Round”. 60. Any round yellow breed. SECTION IV.—SPECIAL PRIZES. Entrance fee, 20s. Every exhibitor will be allowed to enter for all classes in this section, but may take only two prizes; should he get more awards in this section, he will have the option of choosing which two prizes he will take. All awards will count as points in the aggregate for the Grand Championship Prize. 61. Five muids of shelled “Choice White Flat” maize, suitable for export (any breed), with one bag of ears of seed-maize of the same breed, each bag to weigh about 100 lbs. Each entry must be accompanied by a certificate signed in the presence of the Field Cornet or the Resident Justice of the Peace, stating that the ex- hibitor has produced at least 500 muids of the same breed of maize on dry lands, during the past season, and that this particular exhibit was grown without irrigation. First prize : ; . £10 Second prize . : BS Third prize. . B63 62. Five muids of shelled “Choice Yellow Flat” maize, suitable for stock food or for export (any breed), with one bag of ears of seed-maize of the same breed, each bag to weigh about 100 lbs. Each entry must be accompanied by a certificate signed in the presence of the Field Cornet or the Resident Justice of the Peace, stating that the ex- hibitor has produced at least 500 muids of the same JUDGING AND SELECTION 245 Class. breed of maize on dry lands, during the past season, and that this particular exhibit was grown without irrigation, First prize , : » 610 Second prize . : » 2S Third prize. : S 63 63. Five muids ‘ Choice Yellow Round” maize, suitable for export (any breed), with one bag of ears of seed- maize of the same breed, each bag to weigh about 100 lbs. Each entry must be accompanied by a certificate signed in the presence of the Field Cornet or the Resident Justice of the Peace, stating that the ex- hibitor has produced at least 500 muids of the same breed of maize on dry lands, during the past season, and that this particular exhibit was grown without irrigation. First prize. . #10 Second prize . : » BS Third prize. : 63 64. Five bags of ears of Heckory King seed-maize (grain on the cob). Each bag to weigh about 100 Ibs. First prize. : ~ Blo Second prize . ; . 85° Third prize. : : $3 65. Five bags of white seed-maize on the cob, of any one breed (except Hickory King). Each bag to weigh about 100 Ibs. First prize ; : . ALS Second prize . ; . £10 Third Prize. 5 : L5 66. Five bags of yellow seed-maize on the cob, of any one breed. Each bag to weigh about 100 Ibs, First prize : ; ¢ 3605 Second prize. : . £10 Third prize. : BS 199. Champitonships.—Championships are a great stimulus to keen competition, if properly arranged and managed. But CHAP. VI. CHAP. VI. 246 MAIZE at some local shows in South Africa the whole purport of a championship seems to have been misunderstood, and the mis- take has been made of calling for separate entries for the championship, thus turning it into a distinct class, which is uncalled for and undesirable. The aim and object of the championship is to determine which is the best exhibit in any class in the whole show; experience proves that if separate entries are called for, for the championship, the end aimed at is defeated, for the majority of exhibitors will not make separate entries, nor pay two fees, on the chance of securing the championship. 200. Principles of Judging.—Although each Province and District specializes in particular breeds of maize, and though the ears produced in each may differ in size, etc., the principles underlying maize judging are the same for all conditions, and these must be clearly understood in order to judge successfully. It is not merely a question as to which is the best exhibit in its class, on a particular show, but whether the exhibit com- pares favourably with a definite standard. This standard should be the one recognized by authorities as embodying all of the qualifications of the best maize. Therefore the judge must be thoroughly familiar with the points on which maize is judged, and with the standards which have been set for each breed. A printed “standard of perfection” is a useful guide ; such a standard cannot be final, but will grow or be modified from year to year as the various breeds are improved or altered. A well-arranged score card is of great assistance in main- taining a judicial balance. But the judge should bear in mind that there are no absolute rules which can be reduced to writing by which maize samples can be properly judged, in- dependent of that intuitive perception of good and bad points which in a good judge accompanies experience. The score card may easily be abused if it is used in a strictly mathematical sense, for there are certain points which cannot be reduced to precise figures, and which will be neglected in the effort to do so. If the score card is slavishly followed, a wrong decision will result. On this account the use of the score card is sometimes condemned, though it is usually not the score card but the lack of comprehension on JUDGING AND SELECTION 247 the part of the judge which is to blame. The judgment of the CHAP. person who is comparing the exhibit must enter into the score, Y" and experience should guide him in marking each point on the card. There are some men who are born judges, who can intuitively judge by eye without the aid of a score card ; there are a few who even claim to find the score card a hindrance, because their perception and summarizing of points go together so quickly ; but we believe such men are rare. There are so many points in an exhibit, that one may easily place undue value on those which can be seen at a glance, such as length of ear, uniformity, narrow sulci, good colour and good tips, to the neglect of such important points as shape and depth of grain, circumference and shape of ear, yield per ear, percentage of grain to cob, and the like. To the average judge of maize, the score card is of great assistance in just such cases. 201. Methods of Judging.—TVhe aim in judging is to de- termine which is the best exhibit, by careful comparison with a uniform standard scale of points. It is desirable to keep the separate scores of different exhibits side by side for comparison, and the most convenient way to do this is to have a printed score card of uniform size and shape. In scoring any point, it is useful to have the exhibits which have already been judged also laid out side by side, so that at any time a comparison of scores already made may be easily and quickly noted. Where there are many entries and the scoring is at all close, it is well to refer from time to time to the score previously given for the same point in the other exhibits. Unless this is done, it is difficult for the judge to keep clearly in mind the exact “cut” made on a point in previous cases. The stewards, judge, and assistants should use care not to injure the ears, which should be handled as little as possible, and should not be broken. If damaged by falling, the loss of grain will affect the percentage of yield. Good light is necessary in order to detect poor colour of grain and cross-bred grains. A convenient table is essential to good judging; it should be of such a height that the judge can see both tips and butts without handling the ears, and so that he need not stoop unduly to examine them. The judging table should be of such size and so arranged that 248 MAIZE Fic, 92.—Maize Exhibits at the First South African Maize and Citrus Show, Johannesburg, Tgto0. JUDGING AND SELECTION 249 the ten ears of each exhibit can be laid out side by side for comparison. An exhibit of ten ears is desirable, as it facilitates scoring of points and rapid calculation of averages. Rapidity of movement is essential to success in judging a large number of entries; ten minutes should be long enough for any one ex- hibit of ten ears, except for determination of percentage of grain to cob and total yield of grain. To judge a sample accurately in ten minutes means that all unnecessary moves must be omitted. The eye must be trained to judge accurately at first sight. After the other points have been determined in any one sample, a competent assistant should follow to shell off and weigh up the ears to determine percentage of grain to cob. For this purpose the five alternate ears of the ten are weighed and shelled, the weight of shelled grain is taken, and the percentage determined. 202. Judging Matze for Seed.—The aim in judging seed- maize is to determine which is the best sample for seed pur- poses. The dest seed-maize is that which will produce the heaviest yield per acre of grain, of the best quality for feeding or for manufacture. Such a type will, obviously, be the most profitable to grow. The competing exhibits are carefully compared with the standard recognized as embodying all of the qualifications of the best seed-maize. These qualifications include points (1) which ensure good yield, eg. size, uniformity, and shape of ears, straightness of rows, well-filled butts and tips, shape of grain, yield of grain per ear, per- centage of grain to cob, etc. ; (2) which ensure a perfect “stand”’ or crop in the field, e.g. uniformity in shape of grain, size of embryo, percentage and vigour of germination ; (3) which ensure good condition for consumption or export, e.g. maturity, soundness, dryness, etc. ; (4) trueness to type and breed characteristics in shape, colour, etc. ; (5) the value of the sample for feeding or manufacturing purposes as evidenced by the comparative percentage of protein, oil, starch, etc. CHAP. VI. CHAP. VI. 250 MAIZE Shamel points out that these standards have been de- veloped and arranged by experienced growers, breeders, and judges to such a degree that a sample which comes up to these standards has been found (1) to give the best yield; (2) to have the greatest degree of vitality; (3) to be the most profitable seed to grow, and consequently (4) to command the highest price as seed. It is recognized, however, that not enough is yet known about the correlation of characters to say that the standards are perfect. 203. Desirable Characters for Breeding FEars.—A casual glance at an ordinary harvested crop of maize ears conveys but little idea of the degree of variation among them. It is surprisingly difficult to find ten uniform ears in a heap of many thousands from an ordinary crop. Much remains to be done in the thorough scientific study of the maize plant to find out which visible characters are associated (“correlated ’’) with the invisible characters to which we owe yield and quality. That such visible characters exist is well known among stock-breeders ; an experienced dairyman buying a milch cow looks for one with a long, thin tail, prominent ‘‘milk”’ veins, good udder, and with a certain type of body and head. It is not probable that a thin tail has any direct connection with the supply of milk, but experience shows that a thin-tailed cow is usually a better milker than one with a thick coarse tail. So with plants; there are visible cha- racters which may be correlated with the invisible. Experience shows that certain characters of the ears are in certain breeds associated with heavy yields. A study of these correlations has led to the framing of score cards for judging. 204. South African Score Card for Seed-maize-—The following score card has been successfully used by the Depart- ment of Agriculture of the Union of South Africa. It has been carefully prepared by comparing and testing the various score cards in use in the United States. It differs from any one of them in that greater stress is laid on weight of grain per ear than on proportion of grain to cob; it is the yield of grain that is important, irrespective of the amount of cob. JUDGING AND SELECTION Score CarD FOR JupGING Maize Ears. Nook Ex hibttiecsdcyccianecuicqecetee Name and Address of Exhibitor Disqualify any exhibit of white maize which has a red cob. If any exhibit is conspicuously deficient in one or more of the eight last. named points, those particular points should be taken into consideration first. SCoRE. 1. Length of ears. (Measure and compare with standard of perfection; add the total sum of the deficiency, and for each inch cut 1 point) 3 ; : e 2. Sulci, i.e. space between rows. (Cut 1 for sulci 4 inch wide or over, °75 for 41, to 43 *5 for ,') to at) a 3. Shape of grain. (This depends partly” on the breed; it is usually desirable for the grain to be wedge-shaped, and even in Hickory King it should be longer than broad; take particular note of the shoulders at top and bottom of the grain. Allow ‘5 for every ear having well-shaped grain). 4. Length of grain. (This must depend on the breed standard; a well-grown Hickory King may be }} inch long; and a good Yellow Horsetooth, Eureka, Reid, or Iowa Stlver-mine should reach ? inch; cut 1 for every ear having short grain) 5. Uniformity of grain. (Place one grain from each ear side by side ; cut 1 for every grain which is not uniform with the majority) 6. Yield of grain. (Carefully shell and weigh the. grain from half the ears in each exhibit; take the average weight per ear, and for each $ oz. below standard cut I point) If exhibits score at all closely on the above six points, the following eight additional points should be taken into consideration :— 7. Trueness to type or breed characteristics. (Cut 5 for each ear not coming up to standard in this particular) 8. Shape of ear and straightness of rows . g. Uniformity of exhibit 10. Covering of butts . 11. Covering of tips : : : d ‘ : 12. Colour of grain. (Cut for variation in shade or oe 13. Size of embryo 14. Market condition (i.e. dryness and soundness of ear and grain and firmness of grain on the cob; the grains should be free from decay and should be well filled, not shrivelled nor chaffy) . N.B.—Some breeds, e.g. Ladysmith, Boone County, and Jowa Silver- -mine, have naturally rough grain; no cut should be made for roughness unless it is clearly due to lack of condition; cut 1 for every ear out of condition. Points. “Possible. Io 10 Io 10 MAnnnnwn Io Award. ; CHAP. VI. 252 MAIZE 205. Length of Ear.—Standards at best are but approxi- mate, and especially is this true in regard to length of ear. Change of altitude and latitude affect development, so that it is necessary to vary the standard for length of ear of the same breed, according to the part of the world (e.g. America or South Africa) or even according to the part of the same country (e.g. the Transvaal or Natal), in which it is grow- ing. Seven and a half to 84 inches is the usual length for ears of Hickory King in the United States. We have been able to grow ears of this breed 114 inches long, and it may be necessary to fix the South African standard higher than that of the States. The difference is perhaps due to crossing and subsequent selection over a period of years. Variation of season also affects length of ear, so that it will not do to take the measurements of a single season as a guide in setting the . standard. But variation of season need not affect the use of the standard in judging. In an unfavourable season it may happen that no exhibit comes up to standard length; then all exhibits lose alike on this point, and, at the most, length cf ear only affects the score by 10 per cent. Very long ears are usually produced only when the season is long and particularly favourable, for long ears appear to require a relatively long season for full development. It is probable that the majority of the longest ears in a crop were produced on late-maturing plants, therefore we may reasonably expect that if in seed selection we pick the very longest ears we may be developing a late-maturing strain. This may be counteracted, to some extent, by selecting from among the long ears those that have the most perfect tips. In measuring ears take the full measurement from extreme butt to tip. This can be done best by the use of the foot-rule held in both hands, one end even with the butt the other end over the tip of the ear. Add together the deficiency and excess of length of each of the ten ears, as compared with the standard, and cut one point for each inch so obtained. 206. Sulct or Spaces between Rows.—A wide space between rows means waste of space that should be filled by the grains, and therefore means loss of grain. lhere are two places at which waste space must be looked for— JUDGING AND SELECTION 253 (1) the sude¢ or spaces between the rows of grain on the surface of the ear ; (2) the space between the tips of the grain, especially noticeable with sharply-pointed grains. The sulci are generally widest in broad-, shallow- and smooth-grained ears, and in those breeds having fewest rows. Cut 1 point for sulci } inch wide or over; -75 for 7s to } inch, and °5 for z's to 7's of an inch. But judgment must be guided by experience in this matter. The space at the tips can be judged fairly well by the shape of the grain, but in close judging it is well to shell off a space 4 inches long and 4 rows wide, on the five ears that are to be shelled for determination of yield of grain. By examining the exposed ends and sides of the rows it is easy to determine the degree of loss of space; ‘25 may be cut for each ear show- ing too much loss of space. Reduction of width of sulci is well illustrated in Fig. 67 of Doyle Hickory King. 207. Shape of Grain. — Generally speaking, the wedge- shaped grain is the best type to breed to, because it neces- sarily furnishes the greatest amount of grain for the same size of ear. The shape of the grain is influenced to some extent by the number of rows, for we find that ears bearing eighteen to twenty-four rows usually have wedge-shaped grain, while those with less than eighteen rows are apt to have broader, rectangu- lar, or round-edged grains. This is not constant, however, for 10-row Azckory has more or less wedge-shaped grains. The question of the most desirable shape for each breed is largely a matter of experience. In South Africa the types have not yet been definitely fixed. The edges of the wedge should not be curved but straight. The proportion of starch is much higher in a ¢#zck grain than a thin one, and the proportion of bran and waste lower, which appeals to the miller and merchant ; therefore thick grains are more desirable than thin ones. By thick grains we mean thick in the direction of the main axis of the ear. The tip of the grain should be thick, plump, and not sharply pointed. Grain with a thin tip has a relatively low oil- and protein-content, and usually a lower vitality. CHAP. Vi. CHAP. VI. 254 MAIZE In scoring, 5 is allowed for each ear having well-shaped grain. 208. Length of Grain.—The longer the grain, the greater the yield, but the shape varies with the breed, and the length should be in good proportion to the width. Breeders classify by shape of grain, having three groups of breeds :— (2) Grain broader than deep. (6) Grain as broad as deep. (¢) Grain deeper than broad. Scoring should be based on the characteristic shape of the particular breed being judged, which implies a good knowledge of the different breeds: e.g. Hzckory King is judged as Heckory King, and should not be cut because the grain is broad in pro- portion to its length; but even in Afzckory Azng there is great variation in length of grain, and preference should be given to the exhibit having the longest grain, if true to breed characteristics. 209. Uniformity of Grain.—Take two grains from every ear at about one-third of the distance from the butt and place them on the table in front of the ear, with the tip of the grain pointing to the ear. The shape of the grain will vary with the breed (see Fig. 61), and its shape should be true to the characteristic of that breed. Whatever the shape and size of grain in the breed, the grains should be uniform on all parts of the ear, not only in shape and size, but also (in dent breeds) in the character of the dent and smoothness or roughness of the grain end. Thus on smooth ears all the grains should be smooth, and on rough ears all should be rough. Roughness of grain is not necessarily objectionable, for as a general rule we find that a smooth grain is usually shallow, while a more or less rough grain is usually deep; but roughness may be too pronounced, and an extremely rough ear is difficult to handle and makes husking a slower and more expensive process. 210. Yreld of Grain per Ear.—Weigh together the five alternate ears of the exhibit, shell them carefully so that none of the grain is lost, and weigh the grain; calculate the average weight per ear by dividing by five, and for each half ounce be- low standard cut 1 point. The percentage of grain to cob can then be calculated; this will depend partly on the condition of the ears, for ears JUDGING AND SELECTION 255 that are thoroughly dry yield a greater percentage than those which are still wet. Ifthe yield of grain per ear is obtained, there appears to be little—if any—advantage in scoring for percentage ; it is the total weight of grain which we wish to increase, and whether it is borne on a thin or a comparatively thick cob is immaterial. Experience shows that very thin cobs do not give such heavy yields of grain as comparatively thick ones. 211. Trueness to Type and Breed Characteristics.—In live- stock breeding the desirability of keeping to uniform types is universally recognized. It is equally important in plant- breeding, but the principle is less rigorously applied because the types of farm crops are not so definitely fixed. Stock- breeding on systematic lines has been practised for many generations ; maize-breeding on definite lines is much younger, and in some cases the types change as the breeds improve. But even in maize, standards have been set by Breeders’ As- sociations, for the older and well-established breeds, and where this is the case it is desirable to adhere to them. Variations of one sort or another are met with, more or less frequently, in most breeds of animals and plants; these are culled out by the careful breeder. If any such variation is likely to prove beneficial, the individuals showing it may be isolated and inter-bred in order to “fix” the new character, and by this means new strains, races, or breeds may be pro- duced. Lut tt 7s not desirable to start new breeds unless their distinctive characters are clearly worth having ; there is far too much naming and propagating of novelties based on trivial or unstable characters. Such “new creations” often lead to bitter disappointment on the part of the grower; the farmer would be wise to await the verdict of a competent Breeders’ Association before spending time and money over untried novelties, and the Show Committee should not include them in the prize list without good authority. Selection to type is necessary to the preservation of the characteristics of the breed, and also to the development of that uniformity which is essential to the production of the best merchantable article. With maize it is not always easy to furnish a written description of the breed characteristics, which could be recognized by anyone unfamiliar with them; in judging this point experience is the best guide. CHAP. Vi. CHAP. VI. 256 MAIZE In the case of breeds the characteristics of which are not yet definitely fixed, the grower must choose for himself since there is no established standard to guide him. Among the ears in his crop which show variation in characters there is usually one type which is better than the others. 212. Shape of Ears.—The shape of the ear affects the yield, quality, and uniformity of the grain. The object in view is the selection of the best shape of ear to produce the largest possible yield of shelled grain, and to ensure proper maturity under prevailing climatic conditions. The cylindrical ear is, on the whole, the best for these purposes, but some breeds are characterized by a more or less tapering ear, and where a tapering ear is a characteristic of the breed it should not be treated as a defect, nor should the exhibit be “cut” on that account. If the grower does not like a tapering ear he can discard that particular breed in favour of one with a more cylindrical ear, or he can begin to develop from the old breed a new one which will meet his particular requirements. But little is yet known as to the actual relative merits of the different ear-shapes. The principal objection to a tapering ear is that the grains in the upper portion are usually much smaller than those on the rest of the ear, and an uneven sample is the result. It is sometimes found that two rows run only part way up the ear, or in other words are “lost” (Fig. 83); this is a defect, as it means loss of grain. It is difficult to define the varying degree of tapering in different breeds, and here, again, experience rather than written rules must be the judge’s guide. Some breeds being grown in South Africa at the present time, e.g. Chester County, show a tendency to flattening or lateral compression of the ear which sometimes develops into a fasciation (Fig. 854) of either the whole, or the upper portion, of the ear, and sometimes to a lobing or division of the apex into fingers (Fig. 858). These features are undesirable, and all tendency towards them should be bred out by discarding ears which show flattening at the tip. A quarter point is allowed for each well-shaped ear in the exhibit. 213. Straightness of Rows.—Straightness of rows may be less important than size or shape of ear and depth of grain, JUDGING AND SELECTION 267 but aithough it may appear at first sight to be merely a “fancy point” (1 175), it has a bearing upon the yield and quality of the grain. An ear with twisted (Figs. 58 and 59) or irregular (Fig. 84) rows cannot carry as much grain of uniform quality as one of the same size but with straight, regular rows. Moreover, the tendency to twisted rows seems to be cumulative, and may develop into complete loss of rows, which tends to a reduction of yield and an unevenness of grain. At the same time it has been observed that some other- wise well-bred strains develop a marked tendency towards slight twist in the row, and an exhibit should not be scored too heavily on this account, if it has depth of grain and other de- sirable characters. 214. Uniformity of Exhibit.—Uniformity of exhibit refers to uniformity in appearance, shape, size, colour, indentation, smoothness, etc., but not to the Azad of shape or colour. The shape and size may be poor, but if the ears are alike they must be given full marks for uniformity ; the poor shape and size will be scored down when those particular points are dealt with. In scoring for uniformity remove those ears which are distinctly different from their fellows; half a point is allowed for each of the remaining uniform ears. 215. Butts of Ears.—To some extent the shape and cover- ing of the butt are breed characteristics, and in such cases allowance may be made for this fact. With most breeds, however, it is desirable that the rows of grain should be carried well over the butt (Fig. 664) leaving only a narrow opening through which the shank passes to the cob. If the rows of grain end abruptly on a level with the end of the cob (Fig. 66B) the ear will not yield as much (other things being equal) as if they are well carried over. A szwo//en butt is an undesirable character, for large, poorly filled butts usually have unduly large and thick cobs; these dry out slowly, and delay harvest ; in frosty parts of the country this results in damage to the grain. The shank should be medium in size, for large, coarse shanks break off with difficulty and delay the work of harvesting and shelling where the latter is done by hand. But the shank should not be too small (Fig. 70) or the ear will break off in the wind before the main crop is ready to harvest. Exhibitors at a show should trim out the shanks 17 CHAP. VI. CHAP. VI. 268 MAIZE with a penknife before sending in their exhibits, for if the shanks are left on, the appearance of the exhibit is spoiled and the judge is apt to mark down accordingly. Well-filled butts are more frequently met with than well- filled tips, because the silks from the butt appear first and remain in a receptive condition until sufficient pollen is avail- able for fertilization. The silks from the tip of the ear appear last, and it not infrequently happens that all the pollen has been shed before they appear. In most South African breeds the pollen is mature some days before the silks become receptive. The following scores are allowed for butts :— (1) For butts having the grains swelled out around the shank in a regular manner, leaving a concave de- pression, allow full marks. (2) Grains swelling out but not in a regular manner: cut ‘I. (3) Grains not swelling out beyond cob but regular in size : cut ‘2. (4) End of cob covered, but grains flat, shallow, and irre- gular: cut °3. (5) For poorly covered butt: cut °5. 216. Thickness of Cob.—The cob is merely the support which carries the grains, and the larger the cob—other things being equal—the greater the number of grains that can be carried upon it. A careful study of this point shows that the best yielding ears have thicker cobs than those which give poor yields, or, in other words, that thin cobs generally result in poor yields. Buta very thick cob should be avoided, because it requires too long to dry out thoroughly and is more difficult to shell off in a hand-sheller. 217. Tips of Ears.—There should not be any projection of bare cob beyond the uppermost grains on the ear, because this indicates lack of pollination through irregularity in flowering or other defects in the parent. The tip of a well-bred ear should be regularly covered with uniformly-sized grains. The percentage of such well-covered tips will vary with the season, and in some seasons it may be difficult to find any so covered ; but if the scoring is uniform in this particular, all exhibitors will be affected alike and no injustice will be done. It is usually found that short ears are better filled than long JUDGING AND SELECTION 259 ones, and, other things being equal, it is preferable to select long ears which are not so well covered than uniformly short ears which are well covered. In judging, however, length of ear is dealt with independently and must not affect the scoring for covering of the tips. In a well-covered tip the grains should continue in straight rows up to the very end, and not be scattered irregularly; for irregular grains the tips should be scored down according to degree of irregularity. For every exposed or badly covered tip 1 inch long, a cut of °5 may be made, while less is taken off for shorter exposed tips. 218. Colour of Grain.—Yellow grains on a white ear in- dicate crossing, whether the yellowness is dark or pale; this means either :— (1) that the crop has been grown too near to a yellow breed ; or (2) that the seed used was not quite pure, containing some (perhaps only a few) yellow grains or white grains carrying a yellow “ factor”; or (3) that grains from a crop of yellow previously grown on the same ground have produced volunteer plants which have caused the crossing. In the first case the yellow grains are usually most plenti- ful near either the tip or the butt, owing to the fact that the volunteer plants, or the neighbouring field of yellows, came into flower at the beginning or close of the flowering period of the white breed. The effect of crossing a yellow breed with white pollen is not always as clearly marked as in the case of yellow on white ; in some cases the whiteness on the yellow is quite imperceptible, in other cases it shows in the form of a white cap on the yellow grain. Some pure breeds have a normally white-capped grain, e.g. White-cap Dent and Bristol roo-Day ; in such cases a white cap does not necessarily indicate crossing. For one or two yellow grains on a white ear, or white grains on a yellow ear, a “cut” of -25 points is made; for three or four such grains, ‘5 ; for five or six, ‘75 ; for seven or more, cut I point. Missing grains are considered as having been crossed, for the judge has no means of telling that they were not removed by the exhibitor to prevent a “cut” for crossing. In practice rye CHAP. Vi. CHAP. VI. 260 MAIZE cut ‘I of a point for each missing grain, except for those grains (three only) (194, § 11) which have been removed by the exhibitor from near the centre of each ear to determine whether the ears were suitable for seed or exhibition. Richness of colour is a point in favour of yellow maize, but some breeds are naturally paler than others, e.g. Golden King as compared with Yellow Hogan. Dullness of colour may be due to age, to damage in drying, or to harvesting before the grain was sufficiently dry. 219. Size of Embryo.—A small, poorly developed embryo gives a poor germination and a weak seedling, which is less capable of withstanding drought, insect attacks, etc. Most of the oil of the maize-grain is contained in the embryo, so that the larger the embryo, the higher the oil-content. 220, Market Condition.—By market condition is meant the best condition for marketing purposes. Condition includes dryness, firmness of grain on the cob, soundness, maturity, and freedom from injury or disease. Maturity is deter- mined by the filling out of the grain. Chaffy ends usually indicate lack of maturity, but some breeds naturally have rough ends even when the grain is thoroughly mature, and a good knowledge of breeds is essential, to avoid mistakes in this respect. Loose grain usually indicates lack of maturity, but here again knowledge of breeds is necessary, because some of them naturally have a loose grain. By twisting the ear sharply in the hand (but not sufficiently to break it) it is easy to determine whether it is mature or not; if it remains rigid, it is generally ripe and dry, but if it yields to the twist it generally means that the cob is still moist; a cut of half a point,(5) is made for each ear not in condition. At agricul- tural shows held early in the season it is difficult to obtain thoroughly dry ears, and allowance must be made accordingly. In the case of seed-maize the vitality of the sample is of very great importance ; this can be determined by means of a germination test, but at an agricultural show there is not time to make such a test, which takes five days. To determine the viability of a sample of seed-maize, three grains are taken from each ear, one from near the butt, one near the tip, and one near the centre. These will germinate between wet blotting paper, or preferably on a plate of pure damp sand; JUDGING AND SELECTION 2601 the grains are planted with the point downward and barely covered with the sand. They are arranged and marked in such a way that the particular ear from which any set of three was taken can always be determined, so that ears with a poor germination may be discarded. A small plate or saucer or a sheet of glass are used to cover the germinating grain to check evaporation, and if the test is made in cold weather it should be carried on in a warm room. Daily examinations should be made and a note taken of the sets which take long- ‘est to germinate. A germination of 97 per cent in five days is the usual standard. 221. Colour of Cob—White maize should have a white cob. Yellow breeds of maize usually have red cobs. Some breeds of yellow, however, always have white cobs; this is particularly the case with most of the yellow flint breeds (Cango, Botman, New England 8-row,etc.)and with Golden King, Hawkesbury Champion, Yellow Horsetooth, and German Yellow. A red cob in a white breed or a white cob in a yellow breed, excepting in the cases noted above, is an indication of careless selection, and the exhibit is disqualified accordingly. 222. Circumference of Ears.—An exceptionally thin ear _ usually indicates shallow grain, and an abnormally thick ear indicates an unusually thick cob; both will have been scored down when considering depth of grain, uniformity of exhibit and yield of grain per ear, while the lack of proportion will have received further consideration in scoring for length of ear. Where competition is keen it is customary to take into consideration the circumference of the ears as compared with their length. The standard is approximately as 74 inches to 10 inches of length, or 8 inches to 12 inches, but this varies to some extent with the breed. As in the case of length of ear, the excess or deficiency of each ear as compared with the standard, are added together, but the cut made for each inch so obtained is only -25. A mechanic’s small steel tape, divided into millimetres, is applied at a point about one- third the distance from the butt. With a little practice these measurements can be taken with ease and some degree of rapidity, but it is slow work at best, and as it is of minor value it is usually omitted unless necessitated, as was said before, by close competition, CHAP. VI. CHAP. VI. 262 MAIZE It is best to measure the circumference in centimetres. The average can be converted into inches by the following table :— TABLE XLIV. FOR CONVERSION OF CENTIMETRES TO INCHES, IN MEASURING CIRCUMFERENCE OF EARS. Centimetres. Inches, Centimetres. | Inches. — SS ——— ine as Ir‘o 4°32 17'0 5°7 11'5 4°51 17°5 | 6'9 12'0 4°71 17°78 7 oO 12°5 4°92 18'0 | rae) 12°7 5:0 18°5 73 13°0 51 1g'0 | 75 13°5 5°3 19°5 TREN 14°0 5°5 20°0 | 78 14°5 Si 20°32 | 8-0 I5'0 5°9 20°5 | 8:08 15°24 6-0 2I°O | 8°26 15°5 61 21'5 | 8-46 16'0 63 22°0 8°66 16°5 65 | 223. Standards of Perfection.—Standardization of breeds is essential to good judging ; where there are as many breeds as there are in maize it is impossible to carry their several measure- ments in mind without great risk of error. All the leading American breeds have been standardized. The word standard, as here used, is not intended to imply finality; probably no one of the recognized breeds is yet perfect or thoroughly fixed; as improvement takes place, standards gradually change. For the newer breeds, standards have yet to be es- tablished. The following provisional South African standards are given for the guidance of growers and exhibitors. Weight of ear refers to well-matured ears weighed! between July and October; weight per bushel varies with locality; the large, fine-looking Natal-grown //ickory King weighs less per bushel than the smaller Transvaal-grown grain. 1 These weights exceed the figures for weight of grain per ear plus weight of cob, because they are taken from the best single ears available, whereas the weight of ear here given is the average of a number of ears. JUDGING AND SELECTION PROVISIONAL STANDARD OF | | | Hickory King. Iowa Silver-mine. Ear: | | Shape .| Partly — cylin- | drical Length . gin. Circumference | 6°5 in. Rows [8 | Arrangement . | Distinct Sulci Medium to wide Butt Even Tip Regular rows of grain | Shank | Small | Weight . | 10 Oz. Cob: | Size Very small | Colour White | Weight . 1°45 OZ. Grain: Condition —. | Firm upright | Colour | Pearl white | | Apex |Smooth _ to| roughish Form of dent . | Crease Shape Broader than deep | Per cent to ear | 87 Number per ear | 400 | Weight per ear | 8°75 Weight per | bushel : Ibs. | 53 to 62 Ibs. Cylindrical to in. een, I4 or 16 Pairs Narrow Moderately | rounded Regular rows of grain Small 17 02. Small White 1°86 oz. Firm upright Cream white Very rough Pinched Medium wedge go 800 to 1,100 IO OZ. 57 to 64 lbs. Boone County. PERFECTION.—DENT BREEDS. Natal White Horsetooth. 263 Ladysmith. CHAP, | — Cylindrical ro in. 7°5 in. 16-22 Pairs | Medium Moderately | rounded compressed Regular rows of grain Medium 17°5 OZ. Medium White Firm upright Cream white Rough Pinched |Medium | wedge | 86 1,000 to 1,100 | = | | Slowly taper-| Slowly taper ing 12 in. 8°5 in. 14-18 Pairs Medium Even to| shallow rounded Regular rows | of grain Very large 22 OZ. Very large White 3°84 Oz. Firm upright Pearl white Smooth | Crease | Broad and shallow but | thick 78 759 12°5 OZ. 62 | Ing 1o'5 in. | 8°75 in, I4-20 Pairs Narrow Even rounded Small 18°5 Oz. Medium White 1°75 OZ. Firm upright Pearl white Very rough Pinched Deep wedge | 88 800 10°25 OZ. 574 to 62 to | shallow 224. Judging Shelled Maize and the Accompanying Ears.— Both quality and condition are taken into consideration in In a close competition it is impossible judging shelled maize. to give a just judgment without reference to thoroughly presentative ears from the crop. In the classes for commercial (shelled) maize there has been a good deal of divergence of opinion and practice as to whether tip and butt grains should be included or not, and owing to the loose wording of many prize-lists the decision of the judge has been a matter of bitter controversy. re- In 264 MAIZE CHAP. some cases the best entries have been disqualified on this *h account. Unless the prize-list clearly states that tip and butt grains are not to be removed, the exhibitor is entitled to remove them, and should do so. A good judge does not study the tip and butt grains, and it only makes it more difficult to determine the relative merits of the bulk of the grain (which is that from the centre of the ear) if the tips and butts are left PROVISIONAL STANDARD OF PERFECTION.—DENT BREEDS. scab Hickory: 10-row Hickory Yellow Yellow H. eats ieee’ or Louisiana. Horsetooth. SO eT OB ans Eureka. har: Shape Partly cylindri- | Partly cylin- | Slowly taper- | Slowly taper- | Slowly taper- cal drical ing ing ing Length . 8°50 in. 8°5 in. Io in. gin. II in. Circumference | 6’g in. 6°9 in. 7°75 in. 7°25 in. 7°25 in. Rows a2 10 14 I2 or 14 16 or 18 Arrangement . | Pairs Distinct Distinct Pairs Pairs Sulci Medium Medium Medium Medium Narrow Butt Even to shal-| Even Even, some- | Evento shal-| Shallow low rounded times ex- low round- rounded, panded ed, slightly enlarged enlarged Tip Regular rows | Regular rows | Regular rows | Regular rows | Regular rows of grains of grains of grains of grains of grains Shank Medium Small Large Small Large Weight . 12 OZ. 13°5 OZ. 16°5 Oz. 14 Oz. 17°5 OZ. Cob: Size . | Small Small Large Small Large Colour | White White White Red Deep red Weight . _ _ 2°25 OZ. 1°85 oz. — Grain : Condition Firm upright Firm upright | Firm upright | Firm upright | Firm upright Colour Pearl White Pearl white | Yellow with | Orange yel- | Deep yellow light cap low Apex Smooth Slightly | Smooth Medium |Medium rough smooth or smooth : smooth Form of dent. | Crease Crease Dimple Crease Crease Shape Medium wedge |Medium | Broad shal-|Medium |Medium wedge low wedge wedge wedge Per cent to ear — — 82 86 = Number per ear. + | 500 450 625 650 goo Weight per ear — —_ TO"75 Oz, 9°25 02. — Weight per bushel : Ibs. | 63 5534-56 _— 594-64 62 in. JUDGING AND SELECTION Nothing is gained by leaving them in, and much precious time is saved if they are removed by the exhibitor; if he does 265 CHAP, VI. not do it the judge or stewards must do it for him (when judging the sample), and they have more important work. Tf the show committee considers that it is a fairer competi- tion to have the tip and butt grains left in, she fact should be clearly stated in the prise-list. It does not appear that any PROVISIONAL STANDARD OF PERFECTION.—DENT BREEDS. Chester County. Leaming. Ear: | Shape | Length . : | Circumference | Rows. : | Arrangement . Sulci Butt Tip Shank Weight . Cob: Size Colour Weight . Grain: Condition Colour Apex Form of dent . Shape Per cent to ear Number per ear. : Weight per ear Weight per bushel :; lbs. Slowly taper- | Tapering ing Io in. 8 in. 7 in. 7 in. 16 or 18 | 16-24 Pairs or not Pairs | Narrow | Medium Well rounded |Shallow rounded, compressed, expanded Irregular rows | Irregular rows of grains of grains Small Medium 13 OZ. | 13 Oz. Small | Medium Deep red | Deep red 1°54 OZ. | Firm upright | Firm upright Deep yellow | Deep yellow with lighter cap Smooth or me- , Rough dium smooth Dimple | Crease Medium wedge) Medium wedge | 85°25 | — 1,000 | 1,100 9 07. | — 57-634 59-66 Reid. Golden King. | Golden Eagle. | | Slowly taper- | ing | ro in, | 7 in. | 18-24 | Pairs | Narrow (,5 or Jess) Deeply rounded, compressed Regular rows of grains Small 12°5 Oz. Medium Deep red Firm upright Light yellow Medium smooth Dimple Long wedge, shoulder square 88 | Slowly taper- | Slowly taper- 10-14 Distinct Medium Even to shallow rounded Regular rows of grains Large 14 OZ. Large White 2'4 OZ. Firm upright Dull yellow Smooth Dimple Broad and shallow 83°25 600 II*5 oz, | ing g in. 7 in. 16-20 Distinct Medium Moderately rounded, compressed Regular rows of grains Small Small Deep red Loose upright Deep yellow Very rough Broad wedge go 266 MAIZE advantage is gained by this form of competition, and it It must also be New England 8-row (Flint). Partly cylin- drical 1r‘75 in. 5°5 in. |8 Pairs Medium Even, usually expanded Regular rows ot grains Large 10 OZ. Medium White Firm upright Orange yel- low Smooth Flat sides, rounded above Brazilian Flour- corn (Flour). | Tapering g in, 6°5 In. 14 Distinct Narrow Even Regular rows of grains Medium g OZ. | Large White Firm upright | Milk white Smooth Flat sides, rounded above 680 CHAP. ] . e ay VI. certai nly offers the temptation to exhibitors to remove at least some of their tip and butt grains. borne in mind that with modern shelling machinery, some of the tip and butt grains can be removed in the process of shelling. Is a farmer to be penalized for using such machinery ? PROVISIONAL STANDARD OF PERFECTION.—DENT, FLOUR, AND FLINT BREEDS. | | Golden Beauty | Yellow Cango | White Cango (Dent). (Flint). | (Flint). = | | Ear Shape Slowly taper- | Cylindrical | Tapering Ing Length . 9°75 in. 11 in, | g in. Circumference | 7 in. 6°5 in. | 5°5 in Rows | 12 12 12 Arrangement . | Pairs Pairs at the | Pairs at the | butt butt , Sulci Medium | Medium Medium | Butt : | Even Even de-| Even,slightly | pressed | enlarged Tip Regular rows | Regular rows | Regular rows | | of grains of grains of grains Shank 3 | Medium | Small Large Weight . 13°5 OZ. | 12 oz. | 8 oz. | | | Cob: | | | Size . | Medium | Medium | Medium Colour Deep red | White White Weight . _ — = Grain: | Condition Firm upright | Firm upright ) Firm upright Colour Deep yellow) Orange yel- | Dirty white paler cap low Apex. . | Smooth | Smooth | Smooth Form of dent. | Crease _— | = Shape Broad wedge, | Flat sides, | Flat sides, rounded) rounded rounded corners | above above Per cent to ear _ | —_— — | Number _ per ear. - | 550 | 600 | 600 Weight per ear | _— = | = | Weight per | | bushel : Ibs. | a 62-67 | 60-68 JUDGING AND SELECTION 267 Score CarRD FOR JUDGING SHELLED Maize. Score. N.B.—Some judges find it more convenient to allow only Points. one point instead of five for each item on the score card ; it makes no difference so long as either method is used consistently throughout. Possible. | Award. Quality of Grain :— | | 1. Length Shape . Thickness . Uniformity (in size, | shape, and thickness) . Colour (trueness to type) . Shade and uniformity of colour . Colour of cob . . Weight per bushel . g. Chemical composition Condition of Grain :— | 10. Dryness . . Sweetness ; | 12. Soundness (freedom from decay) | 13. Plumpness (grain should be well filled not shriveled | | OI AUP W NHN eee Bea me momen moi mes! monn nor chaffy ‘ 14. Cleanness and freedom from rubbish 15. Brightness Ears Accompanying Shelled ‘Maize: — 16, Length : : 5 17. Sulci (space between n rows) : : F 3 5 18. Trueness to type . ; ; : F 5 5 5 monn 19. Shape and straightness of rows 20. Firmness of grain on the cob . Notes. Quality of Grain.—Quality refers to thickness, shape, size, uniformity, and colour of grain, weight per unit measure, and colour of chaff. Length of Grain.—A deep grain gives a larger percent- age of grain to cob than a shallow one, CHAP. VI. 268 MAIZE Shape of Grain.—A rather narrow, wedge-shaped grain gives a larger percentage of embryo to endosperm than a very broad shallow grain, and for some classes of trade the embryo is of more value than the starchy endosperm. But wedge-shaped grains should not have narrow tips. Thickness of Grain.—A thick grain contains more starch in proportion to “ hull” or “bran” than a thin one, and is there- fore preferred for certain classes of manufacture. Uniformity in thickness, in shape, and in depth of grain, improve the quality ; these can be secured by good breeding. Purity of Colour.—White grain must be at least 98 per cent white, and yellow grain 95 per cent yellow ; all else is classed as “mixed”. This applies to bulk shelled grain only. Shade.—White grain should be pure white, free from black tips and brown blotches; cut for brownish tinge ac- quired in sun-drying. Yellow grain should be clear, deep yellow, and uniform in colour. Colour of Cob.—White grain should not have red tips, which spoil the colour for certain manufacturing purposes ; yellow grain should have white tips in Golden King, Austin Colossal, German Yellow, Yellow Horsetooth, and the standard yellow flint breeds. Weight per Unit Measure.—This is usually given in standard bushels. The American Standard in all but two States is 56 Ibs. of shelled grain. Although in South Africa grain is not sold by measure, it is desirable to take the weights per unit of measure, as samples vary greatly, and the weight gives some indication of quality and chemical composition, for the richer the grain in protein the heavier it usually is. Chemical Composition.—The character of the endosperm can be determined to some extent by holding the grain to the light and by cutting it longitudinally parallel with the broad axis; the larger the amount: of translucent horny starch the richer in protein. Inasmuch as most of the oil-content of the maize-grain occurs in the embryo a large embryo usually in- dicates a high oil-content. Dryness.—Dry grain should not (on the High-veld) con- tain more than 12 per cent moisture. A parcel containing not more than 12 per cent will travel safely from South Africa to Europe. JUDGING AND SELECTION 269 Sweetness.—Sweet grain is free from mustiness or other objectionable smell. Soundness.—Sound grain is free from decay, or the ravages of insects and Dzflodva injury (Fig. 159). Plumpness.—Plump grain is well filled, not shrivelled nor chaffy. Cleanness.— This refers to freedom from bits of cob, chaff, and all extraneous matter. Brightness.—A prime choice parcel of maize should be bright and shiny. Some breeds, e.g. Golden King, lack the lustre of others. Grain which has been harvested wet, and then dried out, often loses its brightness, and a dull sample (from whatever cause) is assumed to be due to harvesting when wet. Condition of Shelled Grain refers to soundness, plumpness, sweetness, dryness, cleanness, and brightness. Soundness and plumpness are considered the primary points in studying condition ; sweetness comes third, dryness fourth—for a sweet sample, but not quite dry, may dry out, but a dry sample that is musty will never get quite sweet again. For grader’s requirements, see chapter NII. Twenty-five points may be reserved for the ears accompany- ing a sample of shelled maize. The points to be considered in this connection are those which particularly affect uniformity and the quality of the grain, i.e. trueness to type, shape, space between rows, straightness of rows and regularity of grain, and firmness of grain on the cob. Length.—Length of ear affects the yield; other things being equal, the longest ears should have the preference pro- vided they no not exceed the standard of length for the breed. Space between Rows.— Wide space between rows is space wasted, and usually implies badly shaped grain. Trueness to Type.— Unless the ears are true to type the sample will not be uniform. Shape of Ears.—The more cylindrical the ear the more uniform the grain. Allowance must be made for breed char- acteristics in this respect, for the ears of Leamzng, Chester County, and some other breeds are naturally tapering. Firmness of Grain on the Cob.—If grain is loose on the cob it may mean that it is not as plump and well filled as CHAP. VI. to judge maize. ing a o a a n ee S 3) , =I ar 7] S 8 (3) ‘a E << Sel i>) 8 jem JUDGING AND) SELECTION 2471 possible. Observation of this point is the quickest and surest way to detect this defect. But the point is comparative only, for in some breeds the grain is always more or less loose ; however, this does not appear to apply to breeds grown in South Africa. Straightness of Rows and Regularity of Grain.—Unless the rows are straight and the grain is regular in the rows it will not be uniform. 225. Judges Computing Sheet.—The following form has been found of great assistance in reducing the time required for judging exhibits. It was first designed for use with ex- hibits of ten ears, but by increasing the number of points from fourteen to twenty and leaving out the figures for possible award, it has been adapted for use with either ears or shelled grain :— CHAP. VI. 272 MAIZE CHAP. Form For Usb In JupGinGc Maize By Pornts. DOE Hc rccesentniicniseircinsoay, AA Mecass Show....... Class. Points. N Possible | | Be Award. ¥ 2 |) 3 4 C0 ON OUFWHNH II | Yield of Grain per Ear ; | Weight of Grain per Bushel . | | Average Length | of Ear. | | AWARDS. | NOTEs. Ist Prize. and Prize. | 3rd Prize. | Highly Commended. Commended. | Special Prize. | 226. Useful Form of Judge's Card.—It is a great con- venience to the judge, and is conducive to greater rapidity and accuracy in the granting of awards, if a convenient form of judge's card is used. The variety of judge’s cards and note- JUDGING AND SELECTION 273 books is almost as great as the number of shows held; it includes plain notebooks, printed triplicating books, and printed cards. The card of which a facsimile (reduced in size) is given below, is one.of the best for convenience and rapidity of handling. The actual measurements of the card are 10 X 6 inches. SCCEONy | ere as eet aa AGRICULTURAL SOCIETY. Class No. JupGe’s Carp. Judge's Name: iii, iawccaun dinguavenqasiiany nemnecessmndnudeasahanmanmiascquliaeuean ne Class No. JupGE’s REMARKS. AWARDS. ESt. Prize: Noi, masociasiavessaads 3rd Prize No........... H. Commended No.. ais Commended No. ............... | Ch aM P10 Nisgsssiies edeorer eaaies | i} | and Prize No... Viaeiiiedseienaueis oo Oe | stewards Dat@hisieasandssovecntimetssandan I % Pc =e ein is 2 Re cy DM eng * zi Fs Gage? Cees E> ye Ly Ee iiah ime ee fe. oma ar hg E> lin Oe Ss } iy Pp p A fy ae » , A 63 i ey ah ) 2 ga ee p f % 3 : iG n rt 4 j oy 4 om i D ke “5 p) . , -o pas Te n “Y aa B Fic. 112.—A, Eureka; B, Chester County. 20 CHAP. VII. 306 VWAIZE deep, narrow wedge-shaped, thick, round dimple-dented, smooth or medium smooth, deep yellow with lighter cap. Introduced by the writer in 1905 from Peter Henderson, New York, U.S.A., and now widely distributed. Valued for its early maturity, drought resistance, and heavy yield. Gave the heaviest yield at Potchefstroom in 1910-11, and the second highest average for six years. Is proving suited to the Up- lands of Natal, which have hitherto been considered outside the Maize-belt, or only suited to flint breeds. The small size of the grain is considered objectionable by some local buyers, but the grain is appreciated for stock- food on the European markets. 251. Vellow Hogan.—Fig.113b. Class: late yellow dent ; rows, 12 or 14; length, 9 inches; circumference at 2 inches from butt, 7} inches, from tip, 6 inches; grain, medium wedge- shaped, crease-dented, medium smooth or smooth, rich orange yellow. Introduced by the writer in 1904 from P. L. C. Shepheard and Son, Sydney, N.S.W. At Potchefstroom Yellow Hogan beat Zureka for yield in 1905-6 and 1906-7, and came a close second in 1910-11; it stands third in the average for seven years. 252. Golden Beauty.—Fig. 1134. Class: late yellow dent ; rows, 12; length, 12 inches; circumference at 2 inches from butt, 64 inches, from tip, 5+ inches; grain, deep, broad wedge- shaped rounded corners, crease-dented, smooth to medium smooth, rich orange-yellow. Introduced by the writer, in 1904, from Burpee, Phila- delphia, U.S.A., but did not prove satisfactory in the Trans- vaal, and was subsequently discarded. Separately introduced into Natal where it has proved more satisfactory, and is grown in the Richmond District. A handsome ear, producing a fine, well-coloured grain. According to Sawer, Golden Beauty-takes 140 days to mature in Natal, proves a vigorous grower, is very drought and wind resistant and does not sucker; ‘‘a good general purpose breed, but rather slow for the High-veld””. 253. Yellow Horsetooth —Fig. 114. Class: medium-late yellow dent; rows, 14; length, 9 to 10 inches; circumference at 2 inches from butt, 74 to 8 inches ; from tip, 6 inches; grain, 3°7 EDS y VARIETIES AND BRE CHAP. Vil. $ Bey voonnmanNMNDNNONM nnnanonno9nn9099 Peer Iey, iy TT IOP q oat | DBA NN 010000000009 0000 g0003i onge UU stab Ona rOCen Saraett B, Yellow Hogan. 5 , g Fic. 113.—A, Golden Beauty ; 20* 308 MAIZE CHAP. thick, shallow wedge, with shallow crease-dent; cob, large, VII. \ SSE os 60804! 4 ee” 68 ete 60te bgt gepgeece eeeret Ggr8 * a ee We = Pat ¥¥Uw QF A ; 6 acgrggt eb sdtuyaiy ¥ ‘ a ee a Se tiie -—_ =< & a ee =e 2 = a = = Ss ae a ap Gea 0, reer |) Fic. 114.—Yellow Horsetooth or German Yellow. white. Synonyms: Ger- man Yellow Horsetooth ; Natal Vellow Horsetooth ; Bishop. There are two types in South Africa, known respectively as German Yellow (Fig. 114) ‘and Natal Yellow, Horsetooth ; the former is apt to be earlier in maturing and more drought resistant than the latter, but both vary greatly in these re- spects, according to the part of the country in which they have become acclimatized, and_ the character of the other strains which have en- tered into their composi- tion, for both are very impure. In some cases the type grown under this name is a mongrel flint, segregating each year into flints and dents, and in others it is a par- tial dent which segregates into dents and flints. We are not aware that any distinctive and pure strains have been iso- lated. If planted early, Ve/- low Horsetooth does well on the High-veld, where it proves a good yielder both in grain and fodder. VARIETIES AND BREEDS 309 The grain, when true to the dent type, is valued on the London market, both for manufacturing purposes and for the preparation of flakes. 254. Reid Yellow Dent.—Fig. 115. Class: medium-early yellow dent ; rows, 18 or 20; length, 9-10 inches ; circumference, 7 inches; grain, long wedge, dimple-dented, medium smooth (the rougher grains usually have greatest depth). An early maturing and comparatively drought resistant type, giving good yields. Introduced by the writer from the United States, in 1909; gave excellent results at the Botanical Experiment Station, Pretoria, in 1909-10, IQIO-11, and 1911-12; now being tested at Potchefstroom, where the returns have so far been good. Retd Yellow Dent was originated in 1846 by Mr. James L. Reid, of Tazewell County, Illinois, U.S.A., as a cross be- tween ‘‘ Gordon Hopkins” of Brown County, Ohio, and Lizéle Yellow. Its characteristics of shape, length and circumference of ear, filling out at tips and butts, size and shape of cob, and shape and indentation of grain, are said to have been strongly fixed by careful and intelligent selection, and are uniformly reproduced, and it is considered one of the most improved breeds of maize grown in the United States. The following are characteristics as recorded by Sturtevant and Myrick :— Ear, 93 inches long, 6{ inches (6’9 inches) circumference, 24 inches diameter, slowly tapering ; rows, 18-20 in distinct pairs ; sulci narrow, scarcely well defined ; butt deeply rounded, very compressed, with diverging grains. Grains very firm on the cob, upright, broadly truncate-cuneate, tapering to a point by straight lines, ,°, inch broad, $ inch deep ; long dimple-dented, smooth ; tip grains conical ; colour horn-orange with yellow cap. Cob red. Shank medium to small. Season reported as 118 days in Indiana. According to Sturtevant this appears from the description to be the same as Queen of the Field from Iowa. 255. Minnesota Early.—Fig. 116. Class: medium-early yellow dent; rows, 16 or 18; length, 94 inches ; grain, medium wedge, fairly deep, dimple-dented, medium smooth. An early-maturing yellow dent introduced by the writer in 1909 from the United States; it gave promising results at the Botanical Experiment Station, Pretoria, in 1909-10, and. is being tested further, CHAP. VIl. 310. MAIZE CHAP. - VII. oe A B Fic. 115.—Two types of Reid Yellow Dent, showing improvement by selec- tion, B is the improved type. VARIETIES AND BREEDS 311 256. Star Leaming.— Fig. 117. Class: medium - early yellow dent ; rows, 16 to 20; length, 10 inches ; circumference, AEA ERSTE SCAT AULA AS & f suoUHehoaralL ea ta DALE { ; { } 4 Fic. 117.—Star Leaning. Fic, 116.—Minnesota Early. CHAP. VIL. CHAP. VII. 312 MAIZE 7 inches; grain, medium wedge, crease-dented, rough. Intro- duced by the writer in 1904 from P. L. C. Shepheard and Son, Sydney, N.S.W. Proved particularly useful on the eastern High-veld. Leaming is one of the most extensively grown breeds of maize in the United States, and is said to have the most uni- form characteristics of any yellow maize grown. It there proves adaptable, by selection, to widely different conditions of soil and climate. It was originated in 1826 from a common yellow sort growing on the bottom lands of the Little Miami River, Hamilton County, Ohio, by Mr. J. S. Leaming, and was selected by him towards a standard type for a period ot fifty- six years. It is supposed to be the type from which many breeds of yellow maize have been developed, as most of the yellow breeds show some of its characteristics, and many of them can be actually traced back to Leamzng. The general characteristics of the Leamzng group are: Ear tapering, 9,", inches long, 7 inches circumference; butt rounding or moderately rounding, more or less compressed, with tendency to expand; rows in distinct pairs but mixed at tip, 16 to 24, with a tendency to reduction about the middle of the ear; sulci medium. Gvazns generally firm (sometimes loose) and mostly upright ; generally wedge-shape with square-cut summits and nearly straight edges, long dimple to pinch-dented, horn-orange with yellow cap. Cod red, medium. Sax’ medium to large. Sturtevant recognized five strains of Leamzng, viz.: (1) Early Leaming (Illinois); (2) /uproved Leaming , (3) Leaming Yellow (Ohio); (4) Messourd Leaming ; (§) Star Leaming (Ohio). Of these Early Leaming, Improved Leaming, and Star Leaming have been tested at the Government Experiment Farm at Potchefstroom, but have now been discarded in favour of better yielding breeds. They were also distributed among farmers on the High-veld, where they have done well in some cases. 257. Golden Eagle.—Fig. 118. Class: late yellow dent; rows, 16 to 20; length, 10 to 11 inches ; circumference, 7 inches ; grain, broad wedge, dimple-dented, rough to very rough (or medium smooth, Fig. 118k). Introduced from the United States, where it was originated by Mr. H. B, Perry of HMlinois in 1871, CHAP. VII. 313 VARIETIES AND BREEDS HPLCE HOHE cachh , Salisbury. Fic. 118.—Golden Eagle, as grown in Rhodesia; ears courteously supplied by the Department of Agriculture CHAP. VII. 314 MAIZE In South Africa the cultivation of this breed appears to be mainly confined to Rhodesia, and even there (we are informed) the tendency is to discard yellows in favour of whites for export purposes. Reported (Sawer, 1) as a strong, gross feeder, requiring heavy and fertile soils. 258. Principal American Breeds of Flint Matze.—The following twenty-one breeds of flint maize are recommended, principally for grain production, by the Agricultural Experi- ment Stations of the States mentioned after their names, as reported by Hunt (1):—! zr. Angel of Midnight (r): Utah or1. Milliken’s Prize (y): Vermont. (tested ten years), Vermont, 12. *North Dakota (w): North Dakota~ Wyoming. (No. 148); Utah (tested seven 2. *Canada yellow (y): Nevada (did years). not ripen grain). 13. Orange County (w): Vermont. 3. Canada 12-rowed (y): Vermont. 14. Rideout corn (y): Wyoming. 4. Early Demand (0): Vermont. 15. “Sanford (w): New Hampshire, 5. French Squaw No. 32 (w): North Vermont. Dakota. 16. Smut-nose (w): South Dakota. 6. *Gehv, No. 123 (y): North Dakota. 17. Squaw corn (w): Utah (tested 7. Golden Dewdrop (y}: Utah (tested seven years). , seven years). 18. *Thoroughbred (w): Vermont. 8. *King Philip (y): Ottawa, Kansas, ~19. Waushakum (y): Vermont. Oregon, South Dakota, Utah 20. White flint (w): Utah (tested ten - (tested nine years), Wisconsin. years). g. *Longfellow (y): Ottawa, Vermont. ~21. Yellow Flint Corn (y): South to. Long Yellow flint (y): Utah (tested Carolina (on thin upland soil). ten years). 259. Principal South African Flint Breeds. —Flints appear to have been grown in South Africa long before the introduc- tion of dents, and still persist in the Native Territories, and in districts of poor rainfall devoted mainly to stock-raising. The breeds principally grown were: So¢nzan (both white and yellow) ; Cango (both white and yellow); Repatriation (yellow) ; Bushian (yellow); and “Kaffir mieltes” (mixed white, red, blue, and yellow). The three first-named are now so mixed as often to be indistinguishable, except that difference in time of maturity is associated with the strains grown in one part of the country as compared with those of another. “ Bushman” is rarely met with. The“ Kaffir mielie” is probably a descendant of the old Squaw or “native” corn still grown in Canada, anda similar type has been received from Italian Somaliland. The natives are said to have a predilection for parti-coloured ears, ' Red = (r); orange = (0); yellow = (y); white = (w). Those marked with an asterisk have been tried in South Africa. VARIETIES AND BREEDS 315 The Transvaal Department of Agriculture introduced and tested a number of the improved American and other sorts, some of which have been discarded as unsatisfactory, while others having proved superior to the older sorts already grown, have been widely distributed, and are now mixed with the old types. Among the best of these are New England 8-row (y) and Rural Thoroughbred (w). Wills Gehu (y) and North Dakota (w), introduced in 1909, matured in eighty- seven days at the Botanical Experiment Station, Pretoria, and gave a very fair yield for such a short season; a fresh con- signment of seed was obtained and widely distributed among farmers for the season of 1910-11. The early-maturing flints of South Europe, such as C7n- guantino, Odessa, and Bessarabia, while maturing quickly (some- times in forty to fifty days) give such poor yields as to be unsuited to the broad type of agriculture necessarily in vogue ina large part of South Africa. They seem better suited to the small farm areas of the south of Europe. Another draw- back is their habit of bearing ears low down on the stem, which renders them particularly liable to injury by vermin and to damage by torrential summer rains. These breeds might be useful to plant late, where the regular crop has been destroyed by hail or locusts; but the difficulty would then be to obtain a sufficient quantity of seed at the moment when it is needed, as no one cares to grow for seed those breeds for which there will only be a demand in a bad season. The following have also been tested :-— Vellow Flints—Compton Early, Canada Early (8-rowed), Vilmorin Early, Vilmorin Early (long-eared), Henderson Large Yellow, Ninety-day, Longfellow, Improved King Philip, Harris Golden, Cinquantino, Odessa, La Plata, Argentina, and Shep- pard Yellow Flint. Red Flints.—Indian Pearl; the ears often carry white, red, and blue grains mixed. White Flints—Egyptian, Somali, and Western Beauty have been tried and discarded. Burlington Hybrid is grown to a very limited extent. 260. Cango, white.—Class: medium white flint; ears, 9 to 10 inches long ; 5§ inches circumference at tip, 13 inches diame- ter; slowly tapering ; butt even; rows distinctly distichous CHAP. VII. CHAP. VIL. 316 MAIZE below, 8 to 10. Grazu rather large, 6 lines broad, 44 deep ; roundish at apex, flattish at sides (sometimes classed as a “flat” in the trade), brownish white, embryo large. Shank medium. Coé thin, white. Medium early; roth in average yield at Potchefstroom ; useful breed for late districts. Svea medium short ; tillers freely. Extensively grown throughout the country. Supposed to take its name from the Cango Dis- trict, Oudtshoorn Division, Cape Province, whence it is said to have been brought north. A form of white flint with larger ears than the Botman, and with grains “ flatly rounded on top,” is described by Sturtevant as being grown at Rio Claro, on the uplands of Brazil, under the name A7/c/ho catete ; it was distributed by American seedsmen between 1881 and 1884 under the names “ Homdny,” ‘White Pearl” (not Champion White Pearl), and “ Large White Flint”. There is no demand for this class of maize on the London or Liverpool markets. Cango and Botman are probably the types introduced by the Portuguese, from their Brazilian settlements, into the East Indies and China, and dropped at Mossamedes, Cape Town, and on the East Coast ex route. An African-grown ear of white flint maize, received probably from the late Prof. MacOwan, Cape Town, about the year 1884, was classified by Sturtevant as differing only in colour from Chinese samples exhibited at the Centennial Exposition, and from the sz7/ho dourado grown at Rio Claro, in the Province of Goyaz, on the uplands of Brazil; Rio Claro is on the trade route from Rio de Janeiro to Matto Grosso and Bolivia. Colour we now know to be a most inconstant character, due to crossing ; there are both white and yellow Canxgos and white and yellow Botmans. 261. Thoroughbred, Rural—Ear, 11 to 12 inches long, 2 inches diameter, depressed, very open, strongly distichous, often expanded at butt; rows, 8. Gvrain 632 lines broad, scant 44 lines deep, rounded, dingy white. Shank large. Cob medium, white. Early to medium early. Szes medium short ; growth fair. Introduced by the writer in 1905 from Thorburn, New York, under the name “ Thoroughbred White Flint,’ but it ap- pears to be the same as the standard breed known as Rura/ Thoroughbred. Widely distributed and fayourably reported VARIETIES AND BREEDS S17 on from the Heidelberg, Standerton, Bethal, and Carolina Districts. The “blood” of this strain has entered into the composition of much of the “ ll’Azfe Cango” now grown, from which it is no longer kept distinct, and the name of 7 horough- bred has been dropped. 262. Cango, yellow.—Fig. 119A. Class: medium-late yellow flint ; ear, 84 to 9 inches long; circumference at butt, # to 64 inches, at tip, 54 to 5,', inches; diameter, 14 to 14 inches ; cylindrical; butt even, slightly compressed ; rows, 10 or 12. Gratn medium, 4% to 5 lines wide, 4} to 5 lines deep, shallowly rounded above, flat on sides, colour golden yellow. Shank small. Cob thin, white. Stew medium-tall, robust, leafy, tillering ; good drought-resister, well acclimatized. Medium late. Valued for maize hay, and at one time exten- sively grown in small plots. 263. Wells Gehu.—Fig. t1gc. Class: early yellow flint ; ears, 6 to 74 inches long; circumference at butt, 4% to 5 inches, at tip, 32 to 4% inches; diameter, 1} to 14 inches; almost cylindrical to slowly tapering; butt even; rows, 12, 10, or 8. Gran 5 lines broad, 4 lines deep, shallow rounded above, flat on the sides (intermediate between ‘flat’? and ‘‘round”’), bright yellow. Shank large. Cob white. De- scribed from ears imported by the writer in 1910 from the breeders, Oscar H. Will & Co., Bismarck, North Dakota. Stem short; inclined to stool; early maturing, ripened at the Botanical Experiment Station, Pretoria, in eighty-two days. Has given excellent results in British Bechuanaland and the semi-arid regions of the S.W. Transvaal and western Orange Free State. 264. North Dakota —Fig. 1198. Class: early white flint ; ears, 7 to 8 inches long; circumference at butt, 4} to 5 inches, at tip, 44 to 44 inches; diameter, 1} to 14 inches ; cylindrical to tapering; butt even; rows, 12 (rarely 8). Grau small, roundish, 4 lines broad and deep, shallowly rounded above, flattish on sides (intermediate between round and flat), greyish white. Shank large. Cob white. Described from ears im- ported by the writer in 1910 from the breeders, Oscar H, Will & Co., Bismarck, North Dakota, U.S.A. Sen short, inclined to stool. Early-maturing; ripe at Skinner’s Court, Pretoria, in eighty-seven days, in 1910. CHAP. VII. 318 MAIZE Fic. 119.—Three flint breeds grown in South Africa. A, Yellow Cango ; B, North Dakota; C, Wills Gehu. 319 AND BREEDS VARIETIES CHAP, Vil. : PPeR dP ELD SESPLSSDSELINE STS eee B, Improved Yellow Botman. Fic, 120.—A, New England 8-row Yellow Flint ; 320 MAIZE 265. Botman, white.—Class: medium-early white flint ; ears, 83 to g inches long; 54 to 6% inches circumference at butt, §$ to 5} inches at tip; 14 inches diameter ; tapering to cylindraceous, even and slightly enlarged at butt; rows, 12, 14, or 16. Grain rather small, 44 lines broad and deep, rounded above, flattish on sides (classed as “round” in the trade), white. Shank variable. Cod thin, white. Sze medium height, tillering, leafy below. Early maturing, earlier than white Cango but less robust. One of the principal sorts for- merly grown in small patches in the drier parts of the Western Transvaal and adjacent Bechuanaland, as far west as Kuruman. After seven years’ trial at Potchefstroom it remains, as usual, at the bottom of the list as regards yield, a striking demonstra- tion of the fact that new types developed by sound scientific breeding are superior to some of the older established and longer acclimatized breeds. 266. Lotman, yellow.—Fig. 1208. Class: medium-early yellow flint; ears, 8 to 84 inches long; circumference, 54 inches at butt, 42 inches at tip; diameter, 14 inches; slowly tapering, even at butt; rows, 12 or 14. Grazu 44 lines broad, 3% lines deep, rounded above, light yellow, said to be softer than that of Vew England 8-row and therefore preferred by the older residents of South Africa. Shank small. Cod slender, often found to be white, but the typical yellow Lotman is said to have a red cob. Considered less drought-resistant than New England 8-row. Tillers freely. One of the principal sorts formerly grown by the smaller producers in the Transvaal and Orange Free State. Synonym: 7vansvaal Yellow. 267. New England 8-row.—Fig. 120A. Class: medium- early yellow flint; ear, 9 to 12 inches long; circumference at butt, 5,', inches, at tip, 44 inches; diameter, 14 inches ; a little enlarged at butt, but rarely depressed or expanded ; rows, 8. Grain hard, 6 lines broad, 44 to 7 lines deep, rounded above, flat on sides (sometimes classed in the trade as “flat”), golden yellow. Shank large. Cod thin, white. Medium-early ; s¢emz medium height ; plant stools freely ; said to stand drought better than yellow Lotman,; one of the best of the flint breeds. Introduced by the writer in 1904 from Burpee, Philadelphia (No. 758:04). Subsequently grown and distributed by the Government Experiment Farm, Potchef- stroom, and by Messrs. John Fowler & Co., at Vereeniging. VARIETIES AND BREEDS 321 268. Burlington Hybrid.—Class : white flint; ear, 84 to CHAP, 9 inches long; circumference, 64 inches at butt, 52 inches at VII. t ‘ i Fic. 121.—Gillespie Yellow Flint. Fic. 122.—Brazilian Flour Corn, 21 CHAP. VII. 32/2 MAIZE tip; diameter, 14 inches; slowly tapering ; butt even, enlarged and open; rows, 10. Gvazz 6 lines broad, 5 lines deep, thick, rounded above, flattish on sides, dusky white, embryo large. Shank large. Cob rather thick, white. Stems, 5 feet, not tillering, ears borne low; maturing fairly early. Ears from W. Gillespie, Rietpoort, Zandspruit, Transvaal, 1909, who grew it from seed obtained in Natal. Grown in Natal for some years; catalogued by Kirchoff, Howie, etc. ; said to make a good table maize. Has been described as a hybrid between sugar and flint maize, but in South Africa it shows no sign of sugariness, 269. Gillespie Vellow.—Fig. 121. Class: medium yellow flint; rows, 12; length, 10% to 11 inches; circumference at 2 inches from butt, 54 inches, from tip, 4% inches; cob red; grain reddish. Selection towards a lighter coloured grain would be desirable from a commercial point of view. A red-cobbed segregate obtained by Mr. W. Gillespie, Rietpoort, Zaudspruit, Transvaal, and exhibited at the Johan- nesburg Maize Show, 1910. 270. Indian Pearl,—Class: parti-coloured flint. Ears long. Cob thin, white. Gratz small, roundish, of high feeding quality, mixed dark red, purple, blue or pearly white on the same or on separate ears. Medium early; yield good to medium. Stem medium height, tillering freely, leafy, good for ensiling. Introduced by the writer in 1903 from Vilmorin, Paris (No. 639°03). Grown for some years at the Government Experi- ment Farm at Potchefstroom, and distributed, but now dis- carded. The strain has entered into the composition of other breeds, and occasionally crops out as a “reversion”; probably the source of the red colour in Claret Sugar. Probably one of the earliest forms introduced into the Old World, and of common parentage with the “Sguaw” or “ Native” maize grown by the Indians of the Northern United States and Canada. 271. The Principal Breeds of Soft Matse or Flour Corn.— Owing to their poor keeping quality in districts subject to weevil and grain moth, and their consequent unsuitability for export, the flour corns are now but little grown in countries of large maize production. There are, therefore, but few com- mercial breeds. VARIETIES AND BREEDS 323 The old South African Bread-mielie differs but little, if at all, from the Brastlian Flour Corn of America. It is stated ! that before the Boer War, there were two sorts of flour corn grown in the Transvaal: (1) the Kaffir Bread-mielve (known in Zulu as whlansa-gazaan), having a short stem, small ears, and white grain; and (2) the true Bread-mze/ie, with taller stem, larger ears, and dirty-white grain with red tips; this latter type is said to be most nearly approached by the sorts of bread mielie recently exhibited at local shows in the Transvaal. A white flour corn was exhibited at the Johannesburg Maize Show, by Mr. Glass of Grahamstown, Cape Province, under the name G/ass’s Early Flour Corn. 272. Brazilian Flour Corn.—Fig. 122. Class: white soft maize; rows, 14; length, 9 inches; circumference, 64 inches ; grain rounded above, with slight dent, and flat sides; cob large, white. Probably the original type from which the old South African Bread-mielie was derived. Introduced by the writer in 1903 from the United States; chiefly grown for local consumption, especially on High-veld farms. Mr, F. le Roux of Oudehouts- kloof, YVolksrust, Transvaal, often exhibits it at local shows. 273. Principal Breeds of Sugar Matse Grown in Amertca. —The following breeds are mentioned by Hunt (1) as having been recommended by three or more of the State Experiment Stations. Early Sorts —Cory, Marblehead, Crosby, Chicago Market, Early Landreth. Medium Sorts.—Squantum, Maule XX, Stabler Early. Late.—Ne-plus-ultra, Stowell Evergreen, Country Gentle- man. 274. Sugar Breeds introduced into South Africa.—Several breeds have been introduced into South Africa by seedsmen and by the Transvaal Department of Agriculture. These include: Black Mexican (7.D.A.), Crosby Early (7.D.A.), White Cory (Clark), Landreth (7.D.A.), Stowell Evergreen (C/ark, Howie), Country Gentleman (Clark, Howze), Golden Bantam (7.D.4A.), New Cory (Clark), Cory Early (fowze), Marblehead (Howze). Poor germination has generally been experienced with imported seed of sugar maize. To Mr. James Clark, seeds- le.g. by Mr. C. J. Morgan, at the ots. Show, 3 March, 1g10. 21 CHAP. VI. CHAP. VII. 324 MAIZE man and florist of Pretoria, is due the credit of what was pro- bably the first attempt to produce a distinctly South African breed of sugar maize. In one of his early catalogues (un- dated, but from internal evidence issued later than 27 Septem- ber, 1904), he offers “ Clark’s Favourite” as a South African production. 275. Clark Favourtte—Mr. Clark issued the following account in the catalogue referred to above :— ‘« Szweet Corn: Clark’s Favourite (Early Sugar Corn).— For years I have been working to get a cross with our Bread Mealie ; this lam pleased to say is a true cross with the Early Cory and Bread Corn. Clark’s Early Sugar Corn is not only the earliest, but has a good-sized ear, white cob, very tender, sweet and nutritious ; it also keeps a long time in good condi- tion before hardening. In all-round good qualities it is the finest of all Sweet Corns; it grows to a medium height, and the ears are formed low and two or three on each stock. This is undoubtedly the most important Vegetable Novelty of this year [1904-5]. Stock very limited.” 276, Arcadia Sugar-maize.—Fig. 123. In 1906 the writer obtained a few white sugar grains from a cross between Black Mexican’ and a Transvaal white flint; in the season of 1907-8 these were propagated and bred true, producing ears of white sugar maize, without any admixture of black. The new breed has since been improved, and was distributed in 1911. It produces two good ears on a stalk ; the ears are 12-rowed ; planted on 20 August, I911, it flowered on 9 Nov- ember ; ears were ready for boiling on 9 December, ie. in I10 days ; it was ready to harvest for seed on 4 January, or 137 days (Burtt-Davy, 8). Earlier-maturing strains producing larger ears have since been produced (Fig. 1238). 277. Claret Sugar.—Developed from a few grains selected by the writer from a Black Mexican cross grown in his garden in Arcadia, Pretoria. A dwarf, early-maturing sort, with pale claret coloured grains. Planted 8 November, flowered 6 January, ready for eating 6 February, i.e. ninety days. It seems likely that the red colour (an aleurone colour) was ob- ' Brought to South Africa in 1903 by Miss Florence Bolton, from the farm of Mr. B. Hayward ot Pescadero, California, U.S.A. DS 7 TIES AND BRI} y VARTE CHAP. Meartcan and lack ) ) tained by an accidental cross between / Indian Peart. VIL. Breeds true. 2 | i | é mt Li Nie, pryt oie | | se { Wee ty Pn me A ie a g | blob A, original type ; Improvement by breeding. B, improved type. Ze. Fic. 123.—Arcadia Sugar-mai: 290 320 MAIZE CHAP, 278. Union Sugar.—Developed from a few grains obtained VII. Fic. 124.—Pop-corn ; a hetero- zygous F, (seed generation) ear (yellow and white). by the writer from a cross between an unnamed red sort and Arcadia Sugar, grown at the Bo- tanical Experiment Station. The grains were obtained from the red ear, and are blood-red, and very deep (4 inch), giving an excellent bite. Probably contains some Indian Pearl strain. Breeds true. 279. Golden Sugar.— Developed from a few grains selected by the writer from a multiple cross grown in his garden in Arcadia, Pretoria. Grains shallow, but broad. Breeds true. 280. Pop-corn.—Fig. 124. Pop- corn is at present but little grown in South Africa, though the demand is increasing. Several breeds have been introduced from time to time from the United States and South Europe. 281. Spectal-purpose Sorts.—It is an axiom in agriculture that breeds adapted to particular classes of trade generally command a better price than ordinary general-purpose sorts, provided they are well grown, and that their production is not overdone. The manufacture of corn-flour, corn-flakes, silver-flakes, grits, semola, hominy, samp, and other special food preparations, calls for particular qualities of grain, which the climate of South Africa seems better fitted to produce than that of most parts of the world. But good ‘condition” (due to climate) is not the only requisite of the manufacturer, and it is VARIETIES AND BREEDS 327 necessary to produce for him the Zy/e of grain called for by the market. One of the first requirements is a large percentage of endosperm in proportion to “bran” or “hull”; to secure this we should aim to produce a ¢hzck (not necessarily broad), long grain, instead of the very thin, flat type which is being grown so much to-day. . There is a large demand in Europe for a small, ‘ round,” flint maize suitable for feeding poultry, pheasants, etc. This commands from 6d. to Is. 6d. per muid more than ordinary yellow flint maize. South African Cango and New England 8-row are too large for this trade, and are being classed as “Flats” instead of rounds by buyers for Europe. If South African farmers can grow the smaller type of maize, it may prove worth while as a catch-crop, provided the yield is satis- factory. 282. Srlage Breeds.—I\n growing crops for silage, it is an object to get as large a yield of forage as possible from an acre of ground. For this purpose tall, leafy sorts which tend to sucker freely are preferred. Useful sorts are ‘‘ Red-cob En- silage,” “Indian Pearl,’ and “ Sweet Fodder-corn” But it is difficult to obtain seed of breeds specially suited to silage production, as they are not regularly grown for seed. South African farmers do not care to grow their own seed of special silage maize, because of the danger of crossing with their grain crops. Most farmers, therefore, prefer to grow for silage the same sorts which are grown for grain, planting them closer in order to get the requisite yield. In America we find that it is mainly those States which are not grain producers which grow special sorts for silage; in the ‘ Corn-belt” the ordinary grain sorts are used, but dents are always preferred to flints because of the greater yield. There is a tendency to use as silage-crops, on the High- veld of South Africa, breeds which are grown as grain crops only at lower altitudes, such as Nadal IVhite Horsetooth and Yellow Horsetooth. The reason is that their season of growth being longer than the upland sorts, they continue to grow later in the season, and thus may produce a greater yield of green-stuff per acre than those breeds grown mainly for grain. But the percentage of ears to stalk and leaves is lower, and the percentage of water to dry matter is apt to be higher, requiring CHAP. VII. CHAP, VII. 325 MAIZE the handling and storing of a larger mass and weight of green- stuff in proportion to amount of dry matter and feeding value. It is not desirable to cut silage too green, and it is therefore doubtful whether it is desirable to use very Jate-maturing sorts even for this purpose. 283. Classes Best Sutted for Cultivation in South Africa.— There is not, and probably never will be, any one breed or variety of maize which can be said, without reserve, to be the best for general cultivation throughout the Union, or in any one Province. Maize is sensitive to changes in climate or soil, and a breed which proves suitable in one district is not equally suited to all, owing to the great variations in altitude, temperature, moisture, and soil between different parts of the country. Speaking generally, the dent breeds are the best for the main crop, as they usually give the highest yields, and are in greatest demand. The flint breeds are most suitable for localities where the rainfall is limited and the growing season short, or for planting after the last date suitable for dents, to increase the acreage under crop. On the other hand, the flint breeds are generally richer in protein than most of the dents, and are therefore more nutritious for stock-food; but on account of their lower yield they cannot be recommended for the main crop where dents can be grown satisfactorily. A certain amount of flint maize is useful on every farm, however. Yellow flints are more suitable than white, for the yellows have a better flavour, and are preferred for stock-food, though the yellow colour does not necessarily indicate higher feeding value. The oversea demand for white flints seems to be nil, while for yellow flints it is unlimited. Hickory King is now more widely grown in South Africa than any other breed of dent maize. It is in special demand for the mines’ trade, though it does not appear to command a higher price than any other well-grown breed of white dent, unless a glut in the market gives the buyer a choice. For the export trade, well-grown Heckory King has met with a favour- able reception on the European markets, especially among manufacturers of grits, flakes, breakfast foods, etc., and among distillers and brewers, The higher price already obtained indicates that ifthe trade is carefully fostered by rigid grading VARIETIES AND PREEDS 324 PD ae and the exclusion from the ‘No, 1 white flat” grade of all but the very best, there is a possibility of securing increased demand and still better prices for the best Mickory Ning. Ladysmith and lowa Silver-mine sell readily in Europe as “ No. 2 flat white,” and choice parcels as ‘' No. 1”. For the grade known as No. 2, or fair average quality (‘F.A.Q.”), yellow dents are in demand for stock-feeding purposes, but yellow flints may be of equal market value if of good quality. For the supply of the present markets, the course for the South African farmer to follow is clearly to produce :— (a) A very “choice” (No. 1) grade of Azckory King, in order to increase the promising, though at present very limited, export trade in this class of maize. (6) A good quality of white dent maize of the No. 2 grade, such as Hickory Horsetooth, 10-row Hickory, Ladysmith, Boone County, Mercer, or Towa Stlver- mine, for use on the mines and for export. (c) A No. 2 quality of yellow dent maize, to be used for stock food locally and in Europe. (¢) A limited amount of yellow flint (where dents cannot be grown profitably) for feeding his own. stock, and for export when the price warrants. In the present condition of the market, farmers will pro- bably find it better to limit their export trade to the first, second and third classes indicated above, though it is not improbable that the export demand for yellow flint. will increase. With the exception of choice Hickory King for export, and yellow maize for local stock food, it does not appear to matter what particular dreed is grown so long as it meets the above general requirements. From the farmer’s point of view, there- fore, the question resolves itself into finding out which breed gives the best yield of maize of good quality under the con- ditions of climate and soil of his particular farm, and this de- pends largely on (1) the time taken by a breed to mature, and (2) length of growing season, which latter depends in turn on (a) altitude, (4) amount and (c) incidence of rainfall. CHAP. Vil. CHAP. Vil. 330 MTAIZL: 284. Relative Length of Growing Season.—The time taken for the crop to mature varies greatly from year to year, accord- ing to the fluctuations of the seasons; in a dry, warm, sunny season the crop takes a shorter time than in a cold, wet, rela- tively cloudy season. Therefore no definite time limit for the ripening of the grain can be assigned to any variety or breed of maize; the time fluctuates with the season. Prof. Morrow, of the Illinois Agricultural Experiment Station, notes the fol- lowing differences in the ripening of Burr IVhite Dent :— 1888 in 135 days. 1889 in 144 and 156 days. 18go in 130 days. 1892 and 1893 in 127 days. Prof. Burrill, of the same experiment station, notes the following difference in time of reaching edible maturity in Crosby Early sweet maize :— 1888 in 62 to 64 days. 1889 in 83 to 85 days. 1890 in 79 days. The season does not affect all breeds in equal degree, but there is a relative proportion between their times of maturity which can be used for classificatory purposes and as a guide to the adaptability of the different breeds to climatic conditions in different parts of the country. The relative ripening period of the different breeds may be roughly classified as follows, the time referring to the period between appearance above ground and the time when the plant is safe from injury from early frost :— Very Late i : ‘ : . 150 days and over. Late ‘ : , : : . 40 to 150 days. Medium Late. i é : = 25 a Rae. Medium Early i ‘ i i. THO EAS ys Early. : ‘ F 4 e 05. LTO” 35 Very Early. ‘ . é s 985 4 05> 3s The relative time of maturity for different breeds is shown in the following list, but the reader should understand clearly that such a list is only approximate, and that the relative posi- tion of different breeds varies in different districts and in different seasons. Asa rough guide, however, such a list has its uses. VARIETIES AND BREEDS 331 Very Lates (150 days and over), for early planting :— Natal White Horsetooth (White dent) Ladysmith (White dent). Salisbury White (White dent). Yellow Horsetooth (Yellow dent). Lates (140 to 150 days), for early planting :— Hickory King (White dent) ; in parts of the Transvaal and Natal. Hickory Horsetooth (White dent). Yellow Hogan (Yellow dent). Golden Beauty (Yellow dent). Golden King (Yellow dent). Brazilian Flour Corn (White soft). Mercer (White dent). Boone County (White dent) ; in Natal. Medium Lates (125 to 140 days), for main crop :— Hickory King (White dent). to-Rowed Hickory or ‘‘ Louisiana” (White dent). Yellow Cango (Yellow flint). German Yellow (Yellow dent). Eureka (Yellow flint). Boone County (White dent); in Transvaal. Golden Eagle (Yellow dent). Medium Earlies (110 to 125 days), for main crop :— Iowa Silver-mine (White dent) ; in Natal. Reid (Yellow dent). Star Leaming (Yellow dent). White Botman (White flint). White Cango (White flint). New England 8-row (Yellow flint). Earlies (95 to 110 days), for late planting :— Wills Gehu (Yellow flint, 85 days). North Dakota (White flint, 85 days). Chester County (Yellow dent). Thoroughbred (White flint). Transvaal Bread-mielie (Flour corn). Minnesota Early (Yellow dent). Iowa Silver-mine (White dent) ; in Transvaal. 285. Breeds Suitable for the High-veld.—On the extreme High-veld (5,000-6,000 feet) it is now generally recognized that Flickory King is too risky for the main crop owing to danger from early frost; many thousands of bags are annually lost from this cause. Where this is the case farmers would be well advised to discontinue growing it, except for very ea7/y planting, and to choose a sort which will supply one of the three other classes of merchantable grain mentioned in 7 283. /owa Sz/ver- mine is said to make as good a “ mielie meal” as Hickory King, CHAP. VII. 332 MAIZS: for use on the mines. For the export trade, or for stock food, earlier maturing yellow dents, such as Chester County, Reid, or Minnesota Early, cau generally be sown where the season is too short for Hickory King. The two early flints already referred to, Wills Gehu and North Dakota, prove particularly valuable for late planting. For Early Planting —Hickory King (w.d.) ; Hickory Horsetooth and 10-row Hickory (w.d.); Eureka (y.d.). For Matin Crop.—lowa Silver-mine (w.d.) ; Chester County (y.d.); Reid (y.d.). For Late Planting —Wills Gehu (y.f.). 286. Breeds Suitable for the Maise-belt of the Transvaal and Orange Free State—For the Maize-belt proper, lying (roughly) between 4,000 and 5,000 feet, the following seem most suitable :— For Early Planting —Ladysmith (w.d.); Yellow Horse- tooth (y.d.); Yellow Hogan (y.d.). For Main Crop.—Hickory King (w.d.); Hickory Horse- tooth and 10-row Hickory (w.d.); Eureka (y.d.). For Late Planting.—lowa Silver-mine (w.d.) ; Chester County (y.d.); Reid (y.d.); Minnesota Early (y.d.). For Very Late Planting, or as a catch-crop.—Wills Gehu (y.f.). 287. Breeds Suitable for the Maisze-belt of the “ Midlands” East of the Drakensberg.—F¥or the lower-lying country east of the Drakensberg, 2,000 to 3,500 feet elevation, later-maturing breeds can be grown owing to the longer growing-season and larger rainfall. For Early Planting.—Natal White Horsetooth (w.d.). For Main Crop.—Hickory King (w.d.) ; Ladysmith (w.d.) ; Boone County (w.d.); Mercer (w.d.) ; Golden Beauty (y.d.) ; Yellow Horsetooth (y.d.). Yellow Hogan (y.d.) is worth trial, the quality of the grain being superior to that of either of the other two yellow dents named. For Late Planting.— Eureka (y.d.) is worth trial. 288. Breeds Suitable for the Coast-belt,—TVhis part of the _ country is suitable for the cultivation of late-maturing sorts when the incidence of the rainfall gives a long growing-season. Boone County (w.d.), Ladysmith (w.d.), and “ Yellow Dent” do well. Hickory King and Golden Beauty are also grown. VARIETIES AND BREEDS 333 At the Government Experiment Station, Stanger, the three following breeds stood out prominently in the breed tests of 1907 :— Yield per Acre. Ladysmith . : : : : 5,100 Ibs. = 25°5 muids. Hickory King , . : $ 4,260: 5; Ser. , Golden Beauty. ‘ : : 3,950 ,, = 19°75 4, 289. Breeds Suitable for the Semi-arid \WVestern Regton.— The region west of a line drawn between Bloemfontein and Lichtenburg is with small exception too dry for any but short- season breeds. Avtiman (flint) is the breed which has been grown most extensively but it yields poorly; a white Botman is grown successfully at Grootfontein and Blikfontein on the Kaap Plateau. German Yellow (y.d.) is favourably reported on from parts of the Orange Free State, as fairly drought- resistant, early, and a good yielder. AHechory Azug is only safe in exceptionally favourable seasons and when planted early : 10-row Heckory (w.d.) is more suitable. /owa Stlver-mine and Chester County have given excellent results in the Wolmarans- stad District. [7s Gehu (y.d.) and Wells Dakota (w.d.) should be especially valuable here. 290. Breeds Suitable for the Upper Bush-veld.—There is an enormous area of territory in the Northern Transvaal, west of the Drakensberg, which is at present but thinly settled by white people. It is a good cattle country and grows excellent maize in ordinary seasons. But the rains fall late and the growing-season is somewhat short. On the Springbok Flats, Hickory King (w.d.) has given good results, but Hckory FHlorsetooth is more promising. /owa Szlver-mine (w.d.) should do well and Chester County (y.d.) is worth trial. 201. Breeds Grown in Rhodesia.— Hickory King, Salisbury White, and Golden Eagle appear to be favourites, and do excellently in those parts of the country suited to maize growing. 292. Relative Vields of Different Breeds in the Transvaal. —The relative yields obtained in any one district do not necessarily apply, except in a very general way, to districts belonging to a different crop-zone or to localities having a CHAP. VIL. different soil. Nor will the returns obtained during one or . two years be a safe criterion as to the relative merits of differ- CHAP. VII. 334 WAIZE ent breeds. Seasonal conditions affect the various breeds differently, and as it requires a ten-year average of the various factors to determine the character of the climate, so also a ten-year average is desirable for determining accurately the relative yields of different breeds. But some breeds can be discarded after two seasons, and many before the expiry of ten years, so even without the full ten-year record a table of relative yields has its value. Breed tests have been conducted on the Government Ex- periment Farm, Potchefstroom, since 1905-6, and the results are summarized in Tables XLV and XLVI. TABLE XLV. SUMMARY OF POTCHEFSTROOM BREED TESTS. | Average Yield in Muids for 3 Years (1906-7 | 5 Years (1906-7 6 or 7 Years. | | to 1908-9). to IgI0-11). eS Se eee 1. Eureka : a 25°79 24°I 21°75 (6) 2, Chester County. ere) LeaeT? BAHT 2115 (6) 3. Yellow Hogan . : | 23°27 213 20°59 (7) | 4. Hickory Horsetooth . | 23°00 20°65 20°00 (6) 5. Natal White Horse. | ae Pe : noe | 23°33 22°10 19°73 i | 6. Hickory King =| 25°14 21°9 19°53 (7 | 7. New England 8-row . | 20°82 Ig'T — 8. Iowa Silver-mine! . | 21°70 20°4 18°76 (6) g. Champion White Pearl | 21°50 20°0 18°72 (6) | 10. White Cango . | 23°50 20°3 17°36 (7) | tr. Yellow Cango 21°58 20°1 17°24 (7) | 12. White Botman . 18'6 16°4 14°73 (7) The following brief notes giving some idea of the method followed in conducting these tests are taken from the Reports of Mr. Holm (1) and Mr. Bell (1) :— Definite comparative experiments with maize were com- menced on this farm in the season 1904-5. Breeds.—The crops were grown on land which had, during the previous year, produced mangels or potatoes, which received from 8 tons to Io tons dung per acre, and about 300 lbs. of a ' The low position held by this breed at Potchefstroom is not maintained in some other parts of the country; it does not like low-lying, damp ground, and does better in localities with less soil-moisture. 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VI. 336 MAIZE phosphatic manure, but no manure was applied directly to the maize crop. The first comparative” trials were put down in the season 1904-5. More breeds were procured for the 1905-6 planting, but as they largely consisted of imported seed it would not be safe to take the results of the first year’s growth as a basis of comparison. In this report, the comparative records are there- fore dated from the season 1906-7. During that year and the previous two years the results, however, served a useful purpose in the determination of breeds which could be discarded. The Seed—Most of the breeds were procured by the Division of Botany from America, Canada, and Australia, but attention was also paid to local sorts from the best seed procurable, It was soon seen that the results obtained in the first two years of experiment could not be regarded as strictly comparative, since the seed of most of the breeds was not acclimatized. This is a further reason for only dating these records from the year 1906-7. The Cultivations.—As a general rule the land was deeply ploughed once after the potatoes or mangels were cleared. Such cultivations as “cultivating,” “harrowing,” and “ rolling” were performed to procure a satisfactory tilth prior to planting. Immediately after planting the land was harrowed ; subsequent cultivations generally consisted of one or two harrowings, one hand-hoeing, and one horse-hoeing. In the year 1906-7, the breeds were planted with the ordinary “2-row planter,” and a special series of plates was used to plant grains of such great diversity, equidistant, but it was found that this did not give as great accuracy as could be desired for experiment. Consequently, in succeeding years, the planting for this breed test was done by hand, the grains dropped in hills 3 feet apart in each direction, three to each hill. In 1909-10 the trials were conducted on land consisting of a brown loam in a very good state of fertility. The actual piece of land devoted to the experiment was of course selected for its uniformity, and to ensure results most favourable for comparison, two rows of each breed of maize were planted across the full length of the field, breeds of VARIETIES AND BREEDS 337 similar robustness and general character of growth being placed next to each other. In previous years the trials had been conducted on check- row principles, the seed being sown in hills 3 feet apart in each direction; but in this year that system was discarded and the seed planted in continuous rows 3 feet apart. “The field was watched throughout the season, and any blanks in the ‘plant’ due to borer, faulty germination, or other such accidental cause, carefully noted. Such blanks, of course, affect the yield, and as they are the result of accident, and not a characteristic of particular breeds, allowance must be made for them in comparative experiments. These allowances, and the fact that the most even stand in a field, even if not the best part, is certainly not the worst, together make the cal- culated yield per acre come out at a higher figure than is actually obtained over large areas. This, however, though giving a somewhat inflated appearance to the yield generally, does not affect the relative positions of the different breeds.” In harvesting, two stretches, each 22 yards in length, were selected out of the full length of the rows at places where the stand seemed to be the most even for all breeds. The ears from both rows of each breed were harvested from these two selected areas, shelled, the grain weighed, and the result calculated out to weight per acre, after the necessary additions for blanks were made. This makes a total of four chains length harvested for each breed. The character of the season should be taken into account in comparing the yields. In 1905-6 two very severe droughts were experienced during the growing period, viz., in January and February, and it is probable that the medium-early breeds were most affected, as these droughts took place at a critical stage in their growth. Late breeds also suffered to some extent, but the early breeds did not appear to have been much affected. The year 1906-7 was ideal for securing heavy yields. A good rainfall took place throughout the growing period, and at no stage of its growth did the crop suffer either from “ drought” or a supersaturated soil. The year 1907-8 was too dry in February, a critical stage in the crop's development, while in 1908-9, though a sufficient 22 CHAP. VIL. CHAP. Vil. 338 MAIZE total rainfall fell during growth, the crop suffered during December from drought, and, later, from a water-logged soil and absence of sunshine. In the season 1909-10 the crops suffered from drought in the early stages, were flooded out in December, recovered somewhat, and produced a fair crop. In the last season (1910-11), owing to heavy early rains, the young plants made rapid growth and got an excellent start. Then came the drought, which reached, and continued at, its worst at the time when the majority of the breeds were at the most critical stage of growth, ie. during the flowering period. Owing to this, in many cases imperfect pollination took place, which resulted in an abnormal proportion of small, badly-filled ears. Rain fell after this, but to the end of the season the fall was considerably below the average and quite insufficient to produce the best results. The rainfall for the period 1906-7 to 1909-10 is given in Table XLVII following :— TasLeE XLVII. RAINFALL AT GOVERNMENT EXPERIMENT FARM, POTCHEF- STROOM, 1906-7 TO Ig0g-Io. 1907-8. 1908-9. 1909-10. ] | | | 1906-7. | } 910-11. | | | | | | Pa [ee = ae Sai g | | Inches. | ¢ eu Inches. | ee | Inches. | ¢ 2 | Inches. ae | Inches.| ¢ @| E fee OE cial aeee (Siet 22) ee | | | | July . : Fo ee ee eee Fae oy Mr | 030) 2) = | — August ; — _ —_ — “98 4 | 0°31 2) — —_ September | ‘ro | 3 | 2°04 | 6 | 167 5 | 0702 I | 0°88 I October | 2°03 9 | 148 | ro | 1°73 7 | 0°39 4 | 6°00 | 13 | November | 3°93 | To | 3°96 | Io | 2°64 9 281 | 10 | 162 Meer | December } 350] Q| 4Ir | 12] 2°59 | 7 \10°26 | 14 | 4'13 | 33 | January | 640 | 15 | 4°21 | 12 | 7:00 | 17 | 3°45 | 12 | 1°46 | x0 February . | 894 | Io | 2°95 | 16) 4°72 | 13 | 1°77 | Ir | 3°33 | ro March - | 2°00 | IO | 3°39 | 12 | 2°93 7 | 2°49 5 | I'gt | It April : - | 5°25 | 1x 02 Ij} ‘80 | 4] org | 3 | 2°53 | 9 | May | £39] “7 | 706 t | E2r 6 | om5 3 | 2°68 | g | June | 02 I 03 tr) — | =| 006) 2 | o'or I | | Total Rainfall . | 33°56 | — |22°25 | — | 269r | — | 22°21 | — | 24°35 ies | Number of Days | — | 85 | =) Bro) | 83 | — |69) — | &4 | Ye eee ae | | | | These climatic conditions are reflected in the yields. In each year all the breeds were safe before frost occurred. VARIETIES AND BREEDS 339 293. Relative Vields of Breeds in Natal.—TVhe following report on breed tests carried on at Cedara, Natal, is furnished by Mr. Sawer (1) :— “In 1904-5 the variety tests were mace in Block 1C of the variety section. The rows were 35 feet long and 3 feet apart, and there were two rows of each variety. Ten tons of farm- yard manure were ploughed in, and a mixture of 200 lbs. sulphate of ammonia, 400 lbs. superphosphate, and 100 lbs. potash chloride was spread broadcast and harrowed in. The same lots of seed from America, France, and England were used as in the previous year; also several lots from Dammann and Co., of San Giovanni a Teduccio, near Naples, Italy, and some two or three other lots. In all cases the seed was planted too late for the best results, the Brazilian and the Virginian Dent on 28 and 29 December, and all the others on 19 December, three weeks later than desirable. Some of the yields were good, but in all probability the general average would have been higher had the seed been planted earlier. The peculiarity of the 1904-5 season, to produce much stalk but comparatively little grain, was shown throughout. The following are the results in detail, arranged in order of grain yield” :— 22 CHAP. VII. 340 MAIZE Tas_e XLVIII. CHAP. VII. RELATIVE YIELDS OF MAIZE BREEDS AT GOVERNMENT EXPERIMENT FARM, CEDARA, NATAL, 1904-5. | | ee Yield. | | No. Variety. | Source. Height | | | Feet. | Grain. Statlete| |. J | | x| Hickory King (L.) . | Trelawny Adams | 73 | 3,538 | 6,128 | | 2} Boone County (L.) . | Henderson (grown | 7 2,917 | 10,417 | by W.Pepworth) | | 3| White Flint Henderson, New 6 1,719 | 6,771 York | | Hig Yellow Hogan Hawkesbury Exp. 7 | 1,614 | 6,406 | | Farm, N.S. Wales | | | | 5| Adams Early Kilminster, Durban) 4 = | 1,510 2,838 | | 6] Early Butler Henderson 6 1,432 | II,041 | | 7| Longfellow i | 6 | 1,354 | 6,328 | | 8| Improved Leaming 46 6 1,237 | 6,341 | g| Improved Early Horsetooth is | by} 1,224 | 8,346 | | t0 | Early Yellow Long Eared . | Vilmorin, Paris 6 1,175 1,680 | | rr | Early Mastodon . Henderson 7 1,172 | 12,578 | | 12 Sugar Corn, Evergreen Late Vilmorin 5 1,167 5,768 | 13 | Early } Minnesota . | Dammann, Italy 5 1,167 | 3,633 | 14 | Golden Beauty . . | Henderson esr: 1,146 | 10,286 | 15 | King Philip . at Z | 6 1,133 5,300 | 16] Rural Thorawslbred . < | 6% | 1,120 9,895 | | r7| Large Yellow Flint : rs | 6 1,015 | 6,419 | 18} Southern Horsetooth | te I) Ce I,OIl5 | 4,297 | | xg | Queen of the Prairie . . | 63 | 975 | 7,526 | | 20 | Late White Horsetooth | Vilmorin | 6 gir | 10,338 | 21 | Six Weeks, or Quarantino . | Dammann Ih oa gil 1,953 | | 22| Crosby Early Sweet Corn . 43 | 34 898 | 6,706 | 23 | White Cap Yellow Henderson 6 885 | 4,948 24 | Extra Early Huron i I) 36 7941 4,414 | 25 | White Early Pyrenean | Vilmorin | 53 791 | 1,706 | 26 | Sweet Fodder . . | Hend.rson eee 755 | 2,905 27 King Philip Early White Vilmorin | 5 743 | 2,070 28 | Compton Early . Henderson | 6 72 4,674 | 29 | Henderson Eureka e aan 716 | 11,783 30 | White Pyrenean Sutton, England 4 703 2,200 31 | Pearl White ‘ Dammann 53 664 | 2,473 2| Extra Early Szekely . Vilmorin | 4 664 | 1,068 33 | Stowell Evergreen Kilminster, Durban) 5 651 6,627 34 | Curagua Dammann h-6 612 | 4,805 | 35 | Sugar Corn, Early Dwarf . | Vilmorin | 44 586 | 1,549 36 | Indian Corn Yellow ‘ 53 567 1,693 37 | Ambra Dammann 3 560 1,719 38 | Seven Weeks, or t Cinquan- tino . c ” 4 547 2,317 39} Early Dwarf Sugar Cane . | Sutton 3 531 2,578 40| Kansas King (in the ey Dammann 43 521 | 6,016 4t | Hickory King | Henderson ] 508 9,778 2| Extra Early Yellow Sutton 4 504 | 1,354 | 43 | Brazilian (L.) C. Harding, near Estcourt 6 456 | 3,398 | L. = local seed | VARIETIES AND BREEDS TaBLE XLVIII (continued). RELATIVE YIELDS OF MAIZE BREEDS AT GOVERNMENT EXPERIMENT FARM, CEDARA, NATAL, 1904-5. Variety. Late Bicolor Pearl a Early Small Yellow Au- xonne é Evergreen Sweet 5 Extra Early Small Yellow Mastodon Improved . Moor Early Concord . King Philip, Early Brown. Iowa Silver-mine Japanese Striped Perry Hybrid Virginia Dent Cuzco White Mammoth . Nanerotollo Egyptian Cuzco Red : Very Early August Adams Extra Early Cory Early Red Cob . Iowa Gold-mine Longfellow Yellow Leaming Yellow Average of 60 Source. Vilmorin a Henderson Vilmorin Dammann ” Vilmorin Dammann Sutton Dammann Henderson Vilmorin Dammann ” ” Vilmorin ” Dammann ” 2 ” ” weig * The ears were all destroyed by hed 200 Ibs. per acre. crows. 34i | Anes | Yield | age = Height Feet. | Grain. | Stalk, etc. al — | 5 | 443 | 12,174 4 | 420 1,146 44) 377 | 45353 3 | 266 846 7 182 963 | 3 | 156 gil I) sds ely A382 625 3. | 133 664 | | 38 | 20¢ | tabd | |) 3 | 104 443 | 6 39 | 9,127 | 5 26 1,380 | 5 20 304 | I 20 | 54 13 ae 5 a 3,125 2 * 2,083 | | - |Failed | | | | eR res 768 | 4,496 The grain would perhaps have CHAP, VI. CHAP. VII. 34 MAIZE TABLE XLIX. RESULTS OF MAIZE BREED TESTS, CEDARA, NATAL, WKH a) aI DN WwWwWwnhh WnH OW ww ns Www NOD Ww [o's) w Ko} Variety. Hickory King Virginia White Dent 1 I Early Mastodon Boone County White. Yellow Hogan Mastodon Improved . Queen of the Prairie . Zulu Red, Zululand Mealie Golden Beauty . Late White Horsetooth | Extra Early Huron Horsetooth and Hick- ory King, Greytown Hickory King Improved Early Horse- | tooth Virginia White Dent 2 Burlington Hybrid, Ingogo . : White Cap Yellow Japanese Striped Southern Horsetooth . Henderson Eureka Improved Leaming Iowa Silver-mine =| Large Yellow Flint Horsetooth or Cura- gua, N.S Rural Thoroughbred . Early Butler Curagua King Philip Brazilian White Flint Evergreen Sweet Compton Early . : King Philip Early Brown : % Longfellow ‘ Kansas King (in the Husk) Mammoth . Small Yellow S. Ameri- can, N.S. King Philip Early Brown, N.S. . 4 Late Bicolor Pearl, NGSa ee ae eet Source. Trelawny Adams Henderson » through W. Pepworth Hawkesbury Ex. Farm, N.S.W. Dammann Henderson Rawlins Henderson Vilmorin Henderson Thresh Henderson ” ” Panzera Henderson Sutton Henderson i ” Dammann | Henderson Vilmorin Henderson ” Dammann Henderson C. Harding Henderson ” Vilmorin Henderson Dammann Natal Dept. Agricult. Vilmorin 1905-6. ey i al Yield. lAverage oe een ee | SD AteTOF Height Harvest- Feet | Grain as ing: | i eee) oe) (ence 5 7 |5,990| 9,869 | 30 May 7 | 5,220/18,385],, ,, 7 4,844 | 5575.5: as, Fo 7 iia 8,242] ,, 4, | 74 | 4279 | 9,247) 4, 45 8 | 4206 | nara }3i4 4, | 8 | Bde | or ae | eee 8 4,075 | 18,789 | 30, | 9 |4oro} 6745/14 ., | 64 | 3,906 | 15,729] 30 ,, | Se | 3037) S073 | Ta | 7h | 3,633| 8,507/30 ,, | 7 | 5,482] ,, 5 | | | 7% |3515| 5.4601, ,, 6 |3,4Ir] 4,662),, ,, | | sil 7 3,398 4,453 | Bh 8 | 3,372 | 5182 | ta 45 4 | 3,268| 4,844] 30 , 7% | 3,255 | 12,696] ,, 4 | 383 | 39255 5560/14 ,, | G- || B3s2TGl 25776 a yes 7 | 3,190 | 6,107 | 12 ,, | 7 | 3,060) 3,515] 3 | 63 | 3,008 | 11,042 30 | 5 | See | PeeD Te ess || a 2,956 | 4,779 | 3 Ta | 2,903) 7274. (124, | 6§ | 2,786] 4.375] 3 5, | 6 | 2,760 | 7,865 | 30 ,, 64 2,760 | 3,763} 3. ,, 64 | 2,760] 3,620} 12 ,, 6} | 2,708} 3,685] 3 4, | | 6 2,682 | 6,797 ie Raut | 63 | 2,630] 5,651] 3 May| | 7. | 2,408) 5,52r/14 ,, 62 | 2,356| 427r| 2 ,, | 5% | 2,356] 4,219] 30 4, | 6 2,253 | 4,948 | 12 April 5 | 2,161 | 5,654 | 30 May | | | | | | | | VARIETIES AND BREEDS 343 TaBLE XLIX (continued). RESULT OF MAIZE BREED TESTS, CEDARA, NATAL, 1905-6. | | , | | | Yield. | | Average £4 oe Dateiof. No. | Variety. | Source. Height Harvest- | Feet. Giain: ae ing. | | | 40| Boone County White, | N.S. . | Henderson 64 | 2,148] 5,248 | 30 May 41 | Indian Corn, Large Yellow Vilmorin 54 | 2,122) 3,893 | 12 April 42 | Perry Hybrid | Dammann y) ,096 | 2,773| 2 May 43 | Sweet Fodder Henderson 6 2,083 | 3,125} 3 4, 44 | Late Bicolor Pearl Vilmorin 54 | 2,051) 5,937}30 ,, 45 | Moor Early Concord. | Dammann 5+ | 2,005] 2,304] 2 ,, 46 | White Early Pyrenean, | I). Ns. é | 6 I,QI2 | 3,711 | 12 April 47 | Crosby Early Sweet | | Corn . «| Dammann | 54 | 1,784| 2,565| 3 May) 48 King Philip Early | | White, N.S. . | Vilmorin lj, 2:6 1,706 | 4,167 | 12 April 49 | Sugar Corn, Evergreen, Late, N.S. =A 52 | 1,680) 4,844 | 2 May 50 | Pearl White Dammann 5% | 1,667) 5,768) 30 ,, 51| Early Yellow, ne Eared | Vilmorin 53 | 1,653) 2,044 | 12 April 52| Early Minnesota Dammann 5% | 1,627] 2,395) 2 May 53 | Large Yellow, N.S. . | Vilmorin 54 | 1,615 | 3,503 | 12 April 54 | White Early Pyrenean | Fn 54 | 1,615] 2,825!1,, 4, 55| King Philip Early White x 6 7,602 | 3,006|,, 56 | Sugar Corn, Everoteen| Late ‘ re 6 1,588 | 3,633| 2 May 57 | Sugar Corn, Early | | Dwarf, N.S. | Pe | 42 | 1,536] 2,109 | 12 April 58 | Adams Early : | Kilminster, Dur- | : | ban 5 1475 | 21457] yo 59 | Stowell Evergreen. | a i 6 AZ4 |) 22261 be. 55 60 | Early Yellow, Long | Eared, N.S. Vilmorin 429i! 35807) ay ae ‘I 61 | Ambra Dammann 53 1406 | 1,693] ,, 4, 2| Egyptian . ‘ i F 64 5380 |} 5,912) 2 May 63| Seven Weeks, or Cinquantino a 6 1,380 | 2,773 | 12 April 64) Extra Early Dwarf | Sugar Corn Vilmorin | 43 5380'| 2,526] ,, 45 65 | Six Weeks, or Quaran- | tino : . | Dammann | 53 g828'| TOOT) as 4 66 | White Pyrenean | Sutton | 43 3315 | 2,330 | 31 Mar. 67. Early Szekely . | Vilmorin 6 | 13,146) 2,317/26 ,, | 68 | Early Small Yellow | ; Auxonne . Pe 54 | 1,016] 3,368|21 ,, 69 | Austin Yellow Corn . Ferguson 6 989 | 2,486 | 30 May | 70 | Extra Early Yellow Henderson 44 g89 | 1,015 | 3x Mar 71 | Extra Early Szekely . | Vilmorin 6 976 | 4,447] 21 ,, 72| Extra Early Small | Yellow - af 53 926 | 2,279 35 23 | Early Small Auxonne re 6 871 | 3,385 rf 74 | Early Dwarf Sugar 5 Corn x . | Sutton 38 599| 2,669| 31 ,, 75 | Very Early August Vilmorin 13 13 218/17 4, CHAP. Vil. CHAP. VI. 344 ; MAIZE 294. Third Season’s Results, Cedara, Natal.— In 1905-6 tests were carried out on the same grounds as in the previous season, but as there were more varieties to be tested the ground had to be extended into Block 1D. This block in the previous season had been manured with 1o tons per acre of farmyard manure, and 200 lbs. superphosphates and 50 lbs. potash chloride. The whole ground was manured again in 1905-6 with 10 tons per acre of farmyard manure ploughed in, and 415 Ibs. per acre superphosphate applied in the drills with the seed. The seed used was from the previous season’s variety plots in all cases except those marked ‘n.s.’ which means new stock or newly imported seed, or with the words Zululand, Greytown, or Ingogo. The rows were, as before, 35 feet long and 3 feet apart, and the plants: were 18 inches apart in the rows, There were two rows of each variety. The seeds were all planted on g December, about ten days later than they should have been, but the 1905-6 season was favourable to late sowing, so that no great harm was done by the delay. Some very fine yields were obtained : the average of seventy-five varieties was at the rate of 2,424 lbs. grain and 5;085 Ibs, stalks, etc., to the acre, and the highest yield was only 10 lbs. short of 30 muids of grain per acre, or actually 106} bushels.”’ Table XLIX gives the results in detail. 295. Relative Weight of Grain per Bushel of Different Breeds.—The standard weight of a bushel measure of maize, in the United States, is 56 lbs. But there is great difference between the relative weight per bushel of the different breeds. The grain which weighs heaviest in the bag or bushel measure does not necessarily give the heaviest crop, in fact the re- verse is generally the case; the flint breeds usually weigh heavier than the dents, but we generally find that they give fewer muids per acre of ground. The weight per bushel varies with the degree of dryness, so that to make a reliable comparison between them, it is necessary to take all the weights at the same time. It would be more accurate to take them in conjunction witha moisture- test. The comparative weights per bushel of some leading South African breeds, taken at the Johannesburg Maize Show, July, 910, are given in Table L. VARIETIES AND BREEDS 345 TaBLe L. WEIGHTS PER BUSHEL OF SOUTH AFRICAN SHELLED MAIZE. Breed. | Where Grown. mS Pet : | | ee | | Lbs Hickory King . ; : : é . | Natal 53 ” yt . : . : : ” SE: 1o-row Hickory (Louisiana). j . | Transvaal 554 m2 oy ” . . =| ” 553 Iowa Silver-mine . . ‘ ; x | ‘8 58 ns ey ‘ , : : a ” 57 Ladysmith ‘ . ; F ' . | Natal 574 Yellow Hogan ‘ ; . : . | Transvaal 5904 ” ” . . = . » | ” 60 Chester County. 3 : : el 3 | 60 * ; : ; ; : ” 57 | Leaming : : : : ‘ 5 59 | Yellow Cango. P : . ; . ‘ 62 | New England 8-row ‘ : : ' ia 62 Gillespie Yellow ; . ; ‘ 3 * 62 White Cango . ; : : : Hs 60 Glass Early Flour Corn . : : . Cape Province 55 Brazilian Flour Corn é ‘ : . Transvaal 524 18/r0404 Average for 18 samples ‘ : ; P| 57°8 Lightest of all = Brazilian Flour Corn : | 523 Heaviest Flint = Cango, New England and Gillespie Yellow 62 Lightest Flint = White Cango . : 60 Heaviest Dent = Yellow Hogan and Chester County . : 60 Lightest Dent = Hickory King .. : : : 5 53 American Standard Weight : i : : . -| 56 CHAP. VIII. CHAPTER VIIL SOILS AND MANURES. The fundamental secret of continued success in farming is the maintenance of soil fertility—Mr. Runciman, President of the Board of Agriculture. Maize requires a better quality of land, and a higher grade of farming, than any other of the great staple crops. —Prof. T. N. Carver. By dung we are limited to the quantity of it we can procure, which in most places is too scanty. But by tillage we can enlarge our field of subterranean pas- ture without limitation. JETHRO TULL. 296. The Soil—Soil is the medium in which plants grow and from which they draw the chemical substances used in the processes of growth. Soils are produced by the weathering and decomposition of rocks. They vary in texture, and are described as stony, sandy, loamy, or clayey; a loam is inter- mediate between sand and clay. Soils vary in chemical composition according to the nature of the rocks from which they have been derived, and may thus vary in the amount of plant-food which they contain. Ninety to 95 per cent of most of the fertile soils consist of the following substances: phosphoric acid, potash, lime, soda, magnesia, iron oxide, sulphuric acid, chlorine, silica, and alumina, which, how- ever, do not usually exist in a free state. The remaining ro or 5 per cent is made up of /Azmus or decayed vegetable matter containing nitrogen. Sometimes a fertile soi] has only 2 or 3 per cent of humus, while in other cases it may contain 25 and even 50 per cent. Nitrogen is also contained in rain-water, in varying amounts, and is further added to the soil by the action of nitrifying bacteria living on the roots of leguminose plants (41 311). Both texture and chemical composition of the soil have an important bearing on plant growth. 346 SOILS AND MANURES 347 297. Chemical Elements of the Soil Required by Plants.— All of the above-mentioned chemical substances of the soil occur likewise in plants, with the exception of alumina (oxide of aluminium), and this is always present in good soils, so that it may be said that all are requisite to plant growth. Some are always present in such abundance that there is no danger of their becoming exhausted. The supply of others, how- ever, especially phosphoric acid, potash, lime, and nitrogen, is frequently insufficient for the production of good crops, and the deficiency has to be supplied by the farmer. Of these, nitrogen is the one which is most expensive to replace. All of the mineral substances found in the ash of plants must come from the soil; plants cannot get them in any other way. The carbon used in the structure of plant tissue is taken from the air. The hydrogen and oxygen come from the water in the soil. 298. Sol Moztsture.—The chemical substances in the soil can only be made use of by the plant when they are dissolved in water, and water is capable of dissolving from the soil all the substances that it contains which enter into the food of plants; this explains why moisture is necessary to plant life. Dilute solutions of these substances are drawn in through the minute hairs which clothe the ends of the youngest rootlets, and are carried up into the plant, where they undergo a chemical change into the various compounds on which the plant feeds. Soil has the power of absorbing and retaining water that passes through it, and also of drawing up water from below ; the latter is known as capillary action, and is similar to the action by which the oil is ‘‘drawn” up into a lamp-wick. Good soils will frequently absorb and hold one-half or more of their own weight of water; some, much more, and those containing most humus will hold most water. Soils also absorb a small amount of moisture from the air. Even when a soil seems perfectly dry it still contains considerable moisture. Soils ex- posed to the direct rays of the sun or to drying wind give up much of their moisture by evaporation, and as the surface dries water begins to ascend from the lower strata by capillary action. When the soil is protected by a “ mulch,” evaporation is checked. CHAP, VIII. CHAP. VIII. 348 MAIZE 299. Conservation of Moisture by Tillage.—South African maize-growers in the drier districts have often experienced loss of their crops, and have had to plant twice or thrice because the stands have “burned off” in a long drought following a good spring rain. Such loss may often be prevented—or greatly reduced—by good tillage. After a rain or after irrigation the surface of the soil is packed tight and a “crust” is formed. Through this crust capillary action is set up, and, as the water evaporates from the surface, more is drawn up from below, until the soil is dried out to a considerable depth. But if that crust is finely broken up, capillary action cannot take place ; the fine soil on top forms a “mulch,” evaporation is checked, and the soil moisture is left for the young plants instead of being drawn into the air. 300. Dry-land Farming.—By the conservation of soil-mois- ture, through the adoption of better tillage, it is probable that maize-growing may be extended considerably beyond the pre- sent western limits of the South African Maize-belt ; just how far, has yet to be determined, but the practice of dry-farming methods would undoubtedly add greatly to the area at present under crop. Jethro Tull (1) says :— “The well hoed earth, being open, receives and retains the dews; the benign solar influence is sufficient to put them in motion, but not to exhale them from thence. The hoe pre- vents the [growth of] turf, which would otherwise by its blades or roots intercept and return back the dews into the atmos- phere, with the assistance of a moderate heat. So that this husbandry [i.e. dry-farming] secures Luserne from the injury of a wet summer, and also causes the rain-water to sink down more speedily, and disperse its riches all the way of its passage ; otherwise the water would be more apt to stand on the surface, chill the earth, and keep off the sun and air from drying it: for, when the surface is dry and open, Luserne will bear a very great degree of heat, or grow with a mean one.” 301. /rrigation.—Irrigation, also, might extend the area now planted to maize. But irrigated land is too valuable to be devoted to this crop except in the vicinity of good markets, where early ‘green mielies,” for table use, command a suffi- ciently high price, or where climatic conditions do not permit maize to be grown otherwise; and then only if the cost of importation exceeds the local value of the crop. SOILS AND MANURES 349 302. Available Plant-food.—Some of the phosphoric acid, potash, etc., present in the soil is in a state of chemical com- bination in which the plant cannot make use of it for food, until a certain amount of “ weathering” and decay has taken place through the action of moisture and air. Estimations of the fotal guantities of such salts present in the soil are there- fore of comparatively little value, alone, as indications of its actual fertility or of its manurial requirements, without a know- ledge of the amounts available as plant food at any one time. 303. Recuperative Power of Sotls.—Soils are possessed of great recuperative power, and if the conditions are favourable the renewal of the available plant-food may take place with considerable rapidity. Under the action of moisture and air a process of ‘‘ weathering” is constantly taking place, and the salts are thereby rendered soluble. This is one reason why summer fallowing often proves so beneficial. Even if the avatlable salts had been quite exhausted before fallowing was resorted to, the soil would not long remain unproductive; the available plant-food would soon be restored by the action of moisture and air on the mineral matter. 304. Character of South African Soils.—As a general rule South African soils are not rich in the total amount of salts required by plants, as compared with those of many parts of the world. Yet the peculiar fact remains, as pointed out by Ingle (1), that Zxruriant crops are yielded by soils which, on analysis, appear to be extremely deficient in plant-food. This is partly explained by the favouring influences of abundant sun- shine and high temperature. In tropical and sub-tropical countries the processes of soil renewal appear to go on more rapidly than in other climates. Soils poor in available salts may, under these conditions, give the plants actually greater nutriment than soils containing a considerably larger percent- age of salts under conditions less favourable (/ng/e, 1). 308. Sorls Suitable for Maize-growing.—Maize as a sur- face-rooting plant is quickly affected by change of climate (Burtt-Davy, 16). It is also sensitive to variations of soil— perhaps more so than other cereal crops. To succeed well it requires a good deal of moisture, but standing water or water-logged soils are injurious; to secure the best results the soil should be moist, but well drained. It should also be of CHAP. VIII. CHAP. VIII. 35° MAIZE good depth, for shallow soils require manuring sooner than deep ones. A friable soil which neither bakes nor cracks much in dry weather is desirable. The black “turf” soils of parts of South Africa are often rich, but in seasons of drought are apt to dry out too much, or in wet years to become water-logged. Red clay soils, also, give good crops in some districts. Some of the more sandy soils bear two or three crops and are then exhausted, or become so loose with cultivation that they blow away from the roots of the young maize plants. There are some soils along the Drakensberg range of mountains which Prof. Watt (IVa¢t, R. D., 1) has found to contain so much of the ferrous iron compounds that maize and kaffir corn do not grow more than a few inches in height even in favourable seasons. Loamy soils, whether red or grey, with some admixture of sand, are among the best all- round soils for maize. In the Transvaal such are found largely in the Heidelberg, Standerton, Bethal, Ermelo, and Lichtenburg Districts, and on that large stretch of country known as the Springbok Flats. Deep, loamy, alluvial river- bottom soils, such as are found along parts of the Vaal River, Kaffir Spruit, the Crocodile, Hex, Marico, and other Transvaal streams, are admirably suited to maize-growing. The rich soils found in pockets along the foot of the eastern slopes of the Drakensberg produce some of the finest crops in the country, but only a short distance out on the plains beyond, there occurs a strip of ashy grey soil which seems unsuited to maize or almost any other crop. 306. New v. Old Lands.—A common practice in South Africa is to abandon maize lands after the third year, either because they are supposedly “ worn out,” or on account of weeds. New lands usually give poor maize crops. Experi- ence at the Government Experiment Farm at Potchefstroom, the Government Stud Farm at Standerton, and the demon- stration farms of Messrs. John Fowler & Co. at Vereeniging, all in the Transvaal, shows that the best crops may be ob- tained in the fourth and fifth year of continuous cultivation of the soil. As the land becomes well opened up to air and water, chemical changes take place in the soil which liberate the plant-food or make it available to the plant. There are a few places in the Transvaal where the soil does not SOILS AND MANURES 351 seem able to stand cropping with maize for more than three CHAP. years, but in most cases the abandonment of the land at the V!!- third year means giving it up just when it should be pro- ducing the very best crops. At Vereeniging an average of 18 muids per English acre has been obtained over a field of 32 acres, without manure, on steam-ploughed land; this was the sixth crop of maize from the land, five having been in succession. On new lands the Vereeniging crop has been as low as 24 muids per acre. 307. Effect of Tillage.—“ Tillage is manure” is an oft- quoted saying attributed to Jethro Tull, but sometimes mis- understood. Prof. Morrow states that proper tillage of the soil increases its productive power; the ability of a soil to produce crops is often as directly increased by tillage as by the application of manures (JZorrow and Hunt, 1). Tillage is described by Jethro Tull (1) as “ breaking and dividing the ground by spade, plough, hoe, or other instru- ments which divide by a sort of altition (or contusion), as dung does by fermentation. . . . Tillage (as well as dung) is bene- ficial to all sorts of land. . . . The finer the land is made by tillage, the richer will it become, and the more plants it will maintain.” But it should not be concluded from this that good tillage makes it unnecessary to manure, for manure adds to the soil, while tillage only makes available what is already there. Fora few years after the first breaking of the veld, land may continue to improve and yield better crops, under good tillage, but after that, deterioration begins. The following are the principal reasons why cultivation makes soils more productive :— (1) Stirring and pulverizing a hard, compact soil enables the roots of plants to penetrate more easily and reach a larger quantity of the salts which are to be converted into plant-food. (2) It opens the soil to the weathering effect of air and water, which increases the supply of available plant-food. (3) With very fine, loose soils tillage (and rolling) may make them more compact, increasing the capillary action. (4) Surface cultivation, which keeps the surface soil loose and dry, forms a mulch, which checks evaporation. (5) Tillage kills weeds, which otherwise rob the soil of food and water. CHAP. VIII. MAIZE ioe) mn nN 308. Effect of Continuous Cropping.—It is well known that the crop-producing power of the soil is reduced to a point below that of profitable cultivation by continuous cropping with maize or any other cereal, even where the soils are rich in the constituents of plant-food. As the average soils of South Africa are not rich in these substances (1 304) they will the sooner become exhausted, unless steps are taken to renovate them. There has been a tendency in parts of South Africa to crop continuously over a long series of years, until the soil has become “sick”? or ‘‘worn out”. Continuous cropping means the annual removal of a certain amount of plant-food from the soil, without replacing any. Neither the greatest quantity, nor the best quality, can be produced by growing the same crop year after year on the same soil. Fortunately there are still large tracts of unbroken veld, but they cannot be drawn upon indefinitely ; the time is rapidly approaching when there will be no more raw veld to be broken to the plough. When this time comes South Africa will be compelled to resort to some means of restoring the fertility of the soil. This has been the experience in every farming country, until the lesson was learned and a more normal practice established. Where the soils are naturally poor in plant-food farmers should not wait until they are “ worn out” before adopting a better method of treatment; it is easier and cheaper to main- tain and add to what is already there than to undertake to renovate an already impoverished soil. 309. Maintaining the Crop-producing Power of the Sotl.— Continuous cropping with a heavy-yielding crop like maize, will inevitably result in exhaustion of the soil unless steps are taken to maintain its crop-producing power. Quoting Mr. Runciman,! the President of the British Board of Agriculture: “The fundamental secret of continued success in farming is the maintenance of soil fertility. This is where England excels. Her system of land tenure is often criticized, but it is a significant circumstance that it is associated with 'In a speech at the Government dinner given in honour of the American Commission on Agricultural Credit and Co-operation, July, 1913, as reported in The Field, Vol. CXXII, p. 128, 19 July, 1913. SOILS AND MANURES 353 the best and most enduring methods of husbandry known in any country.” There is no need for South African farmers to become pessimistic on the question of soil exhaustion, if they will study the example of England in contrast with that of the United States; in the latter we find, according to Hopkins (6), thousands of acres of land practically ruined from an agri- cultural point of view, after but 200 years of farming; while on the other hand we learn from Mr. Runciman that “ ¢he older England grows the richer become the average soils ; cases of impoverishment are few and far between”. The English tenant-farmer is compelled, under the terms of his lease, to restore to the soil what he takes from it, and nothing ends a tenancy more speedily than evidences of exhaustive farming. If proper steps are taken to maintain the crop-producing power of the soil, maize does not prove an exhaustive crop. Hunt (1) makes the following points—that (1) the amount of soil elements removed is small in proportion to the amount of foodstuff produced; (2) large quantities of organic matter are produced which when fed to live-stock make large quantities of organic manure to return to the soil; (3) the intercultural tillage required by the maize crop is beneficial to the soil. Hopkins (5, p. 200) says that to return the maximum amount of organic matter to the land requires that the manure shall be applied to the soil before losses occur by fermentation and decay. ‘In ordinary farm practice more or less loss of organic matter is almost certain to occur unless the manure is applied to the soil within a day or two after it is produced.” English farm practice is changing in accord with this view, largely owing, no doubt, to the effect of the dairy regulations which require that the manure shall be removed daily from the immediate vicinity of the milking sheds; dairy farmers now find it convenient to cart it direct to the land, and the results appear to be entirely satisfactory. The best means of maintaining the crop-producing power of the soil on a maize farm, as at present demonstrated, are :— (1) The use of stable manure and kraal manure wher- ever available. The available amount can be increased by the 23 CHAP. VIII. CHAP. VIII. 354 MAIZE adoption of the best methods of preserving the fodder and stover of the crop, and conserving the manure. (2) The ploughing-in of green-manure crops. (3) The suitable rotation of crops in connection with stock- raising. (4) The use of suitable artificial manures. 310. Summer Fallowing.—Where large areas are under crop on any one farm (as in many parts of the western United States and parts of South America and South Africa) it is not practicable to give the same amount of cultivation as would be possible on a farm of smaller area. It needs too large an investment of capi- tal in machinery and draught animals, not to mention the diffi- culty of getting labour. But without cultivation weeds soon get hold of the land, smother the crop, and greatly reduce the yield. Under such circumstances summer fallowing may be resorted to with advantage. By summer fallowing we mean leaving a portion of the land without crop during the summer season, so that it may be cleaned of weeds. Crops of young weeds are allowed to grow, and are then ploughed under as ‘‘green manure,” or harrowed off before they get too large to be pulled out or old enough to scatter seed. The latter point is of great importance, for there is much truth in the old pro- verb that ‘‘one year’s seeding makes seven years’ weeding ” Summer fallowing has been decried alike by practical far- mers and writers on agriculture, because no immediate return is obtained from the land for a whole season. Another objection offered is that much of the plant-food may be leached out of the soil and carried away in the drainage water, if the lands have a steep slope and the rains are heavy. Experiments conducted by Mr. W. A. McLaren at Vereeniging, Transvaal, have shown, however, that summer fallowing was followed by an ¢ucrease of 11 muids (2,200 Ibs.) per acre in the maize yield, without the use of fertilizers. At one of the American State Experiment Stations the yield of wheat from a field cropped only in alternate years, during a period of ten years, was greater than from a field cropped every year during the same period, thus five crops gave a heavier yield than ten. Somewhat similar results were recorded by Lawes and Gilbert at Rothamstead, England. If the total yield obtained by cropping in alternate years SOILS AND MANURES 356 is equal to that obtained by cropping each year over the same period, the cost of production is reduced and the profit con- sequently increased. Unless the cost of fallowing and rent of the land (or its equivalent in interest) are heavier than the cost of production of a crop without fallow, fallowing will thus pay for itself while at the same time it cleans the land. Not only so, but the cost of cultivation is lessened by plough- ing under two or more crops of weeds on the fallowed land before they have had a chance to seed. Where farms are large and land is cheap, there need be no loss of revenue, if each year only one-third or one-quarter of the land is kept in fallow and the rest under crop. For example, if 1,000 acres of arable land is all that a South African farmer can maintain each year, owing to lack of either capital or labour, or both, he might have 200 or 300 acres of it under fallow each year. During the comparatively slack season, from the end of December to the end of March, he could usually employ his draught animals and “boys” to cultivate these fallow lands. By this means he would save much of the time and expense which would otherwise have to be devoted to cleaning the crop during the following growing season. 311. Rotation of Crops for Fertility Conservation.—One of the cheapest and most profitable methods of resting the land after it has once reached good cropping condition ({ 306) is'to adopt a system of change or “ rotation” in which some other crop than maize is grown every third or, even, second year. Some farmers grow potatoes the third year with the aid of commercial fertilizers, but for unmanured land the best kind of rotation for maize is a leguminose crop, such as cowpeas, kaffir beans, velvet beans, soybeans, peas, or peanuts ({ 313). This rotation crop may be cut for hay or silage, or, better still, ploughed into the ground at the beginning of winter. Practising rotation of crops is one of the best methods of checking the wearing-out of the land (Aurtt-Davy, 15). The principal advantages of rotation are :— (1) That as some crops require more of one kind of plant- food than others, an intermediate crop can be grown between two crops of a kind without interfering with the general fertility of the soil, and still allow time for the chemical changes which replace a certain proportion of the available salts required for 23 * CHAP. VIII. 356 MAIZE CHAP. the following crop. For example, a crop of wheat withdraws VIII. large amounts of phosphoric acid from the soil, but takes re- latively small quantities of lime and potash, while a crop of beans requires a great deal of potash and relatively less phosphoric acid. (2) Some crops are surface feeders while others root more deeply, drawing their food from the lower layers of the soil. By alternation of deeper with shallower rooted crops the avail- able food supply is utilized to better advantage and made to last longer. (3) Certain crops replace in the soil certain ingredients which have been removed by other crops, for instance such crops as lucerne and peanuts give back nitrogen. (4) Rotation of crops helps to clean the ground of ‘“vol- unteer” plants from the preceding crop, which, in the case of wheat, oats, and especially of maize, cause so much mixing or crossing of seed, with resulting loss to the farmer. (5) As different crops require different treatment of the soil, a change enables the farmer more easily to clean the land of such weeds as are particularly injurious to a given crop. In the rotation the farmer can use crops which are known as “ cleaning” crops. The general results of rotation may be summarized as :— (a) The production of crops of greater vigour and better yield. When crops of the same kind are grown continuously on the same land the crop becomes less vigorous and, con- sequently, more susceptible to attack by insect and fungous pests. Rotation disturbs the “balance of nature” in such a way that the pest dies from lack of its normal food or goes away to search for it elsewhere. (6) The reduction of the manure bill. (c) A principle of rotation is the division of the land into two to four portions, no two of which bear the same crop in the same season. A well-arranged rotation reduces the labour required at any one time, inasmuch as not all crops require to be planted, cultivated, or harvested at once. The principal points to observe in planning a rotation are :— (1) Have at least one leguminose crop in the rotation. (2) Have at least one cultivated or “cleaning crop,” or, in its place, a ‘‘smother crop” for weeds. SOILS AND MANURES 387 (3) Alternate shallow-rooting crops with deep-rooting crops. (4) Where there is danger of loss of plant-food from leach- ing of the soil owing to the slope of the land, plan to have a growing crop on the land all the summer. (5) Bare summer fallow can be used to advantage if the weeds are allowed to grow during the rainy season and are ploughed in defore they seed. (6) Do not rotate cereals with cereals. (7) Plan the rotation so as to have about the same amount of forage, hay, and roots each year. (8) Unless it is thoroughly rotted, so that the weed-seeds which it contains are killed, apply the stable manure to the root crop (if one is used in the rotation) or to a rank-growing crop like maize. 312. Organic Matter.—By organic matter is meant matter composed of substances that are or have been living organisms, in contradistinction to the inorganic matter derived directly from rocks, metals, etc. Organic manures include farmyard manure and humus, which are valuable sources of plant-food, especially nitrogen. Humus and organic matter are not synonymous, for humus includes only that part of the organic matter which has passed the most active stage of decomposition and completely lost the physical structure of the materials from which it is made; it has thus become, as a rule, thoroughly incorporated with the soil mass (//opkzns, 5). Beside returning plant-food to the soil, organic matter im- proves its mechanical condition; when it is in the proper state it may materially modify the water-content. Soil which has been manured with stable manure is usually moister than un- manured ground; this may be for some or all of four reasons :— (1) It may absorb more rain-water ; (2) It may draw up more water from below, by capillary action ; (3) It may lose less water from the surface by evaporation ; or (4) It may lose less water by drainage. Briefly, then, organic matter improves the texture of the soil, adds to its moisture-retaining power, and furnishes nitrogen. CHAP. VIII. CHAP. VIII. 358 MAIZE South African soils are frequently deficient in humus, and where this is the case organic matter must be added if good crops are to be obtained. This may be done by manuring with farmyard manure (% 309), or by ploughing into the soil some ‘‘green-manure ” crop. 313. Use of Leguiminose Green-manure Crops.—On sandy soils a leguminose crop, such as soybeans, velvet beans, cowpeas, or kaffir beans, proves very beneficial to the maize crop following. A greater yield of maize may be obtained on a poor, sandy soil from the use of a crop of this character, with the addition of a phosphatic fertilizer, than would be secured in two years where maize is grown continuously (IVat¢z, KD), 2). To the South African farmer a particularly important feat- ure of crop rotation is the possibility it furnishes of adding humus (¢ 312) to the soil by ploughing in a growing crop, usually a legume, which at the same time adds one of the most expensive elements of plant-food, namely nitrogen, to the soil, The leguminose crop in the rotation ({ 311 and 314) is often treated in this way, but a crop of any kind of green weeds may also be ploughed in to advantage, where humus only is wanted, and this may be done when the land is in summer fallow. Hopkins (5, p. 199) definitely states that the most zimport- ant, and least appreciated, method of maintaining or increasing the supply of organic matter in the soil is dy the use of green manures and crop residues. A ton of clover ploughed under will add nearly ¢hree ¢2mes as much organic matter to the soil as can possibly be recovered in the manure if the clover is fed ; but with maize only one-tenth of the dry matter of the crop is found in the manure. 314. Rotations with Matse in other Countries.—A look at a few rotations practised elsewhere may be instructive. One in use in the Northern United States is :— First year : : . Wheat or rye. Second year. ‘ . Clover or grass. Third year. : . Maize with farmyard manure, and with winter rye sown at the last weeding to furnish late pasture and winter feed. Fourth year. ‘ . Oats. SOILS AND MANURES 389 In the State of Rhode Island two different rotations with maize have been practised on light and worn-out lands :— (1) A four-year course . Maize, potatoes, rye, clover. (2) A five-year course . Maize, potatoes, rye, grass and clover for two years. In the State of Delaware the following rotation has been practised to advantage :— First year é , . Maize, followed by crimson clover. Second year . - . Cowpeas, followed by winter oats. Third year. : . Red or crimson clover. In Louisiana, where the climatic conditions are more nearly like those of the warmer parts of South Africa, the rotation recommended by the State Agricultural Experiment Station is :— First year i é . Maize. Second year . é . Oats, followed by cowpeas. Third year. : . Cotton, Another rotation practised in parts of the United States is -— First year ‘ Wheat. Second and third years . Clover and pasture (or hay) grass. Fourth year. : . Maize (manured with farmyard manure). In the Maize-belt of Illinois a twenty-year test was made with maize after maize, as compared with maize in a six-course rotation, viz., oats one year, clover three years, maize two years. The average increase the first year after clover was 5 muids 120 lbs., and the second year 4 muids 51 lbs. (Ant, 1). 315. Some Transvaal Rotations—No systematic plan of rotation has yet been adopted in the Transvaal. On the light sandy loams of some of the potato farms in the Standerton District it is customary to grow maize for two years in suc- cession after potatoes. From 600 to 800 Ibs. per acre of commercial fertilizer is applied to the potato crop; the two maize crops which follow use the residue of the manure not required by the potatoes, and give crops varying from 20 down to 15 muids per acre. On these soils, however, it has been found that after six years’ cropping a change is required, to add humus to the soil. Farmers are, therefore, conducting experiments to include green-manure crops in the rotation ; CHAP. VIII. 360 MAIZE cHap. the kaffir bean, cowpea, and soybean have given the most VIII. promising results. In the Standerton District ploughing in of teff for green manure has been tried.!- The rapid growth of this grass makes it possible to get it in as a catch crop where other crops might not be practicable. At the Botanical Experiment Station, Pretoria, experiments conducted to determine the effect of green-manure crops on subsequent crops of maize, wheat, and sunflowers all gave marked results in favour of the green-manured plots. The Division of Tobacco and Cotton of the South African Department of Agriculture is conducting a series of rotation experiments at Rustenburg, Tzaneen, Barberton, and Piet Retief, in the Transvaal. These experiments are designed to determine the best rotation for the improvement of worn- out tobacco lands. The experiments include the following rotation :— First year. ¥ : . Tobacco. Second year. : . Cotton, Third year : : . A leguminose crop such as velvet beans, cow- peas, peanuts, or soybeans. Fourth year. 5 . Maize. It is intended to add to this a small cereal winter crop such as wheat, barley, or oats, on irrigated lands, to come between the tobacco and cotton, or between the legumes and maize crops. The experiments are not yet complete. 316. The functions of manures are twofold, restorative and additive; to maintain fertility and to increase it. Restorative manures are merely intended to replace in the soil those ele- ments of plant-food which have been taken out of it by cropping. Restorative manures should be of a “complete” or genera] character, i.e. must contain all the fertilizing ingredients. Restoration is chiefly effected by farmyard manure. Billings, F..S.; Peters, 4. T., and Avery, S.; Peters, A, T.+ and Mayo, N. 5. 49 CHAP. XV. CHAP. XVI. CHAPTER XVI. THE CONSTRUCTION OF MODERN SILOS. Probably the most important change that has been made in the handling of the maize plant in the last quarter of a century is the practice of putting the unripened plant, cut into small pieces by a feed-cutter, into a receptacle with air-tight sides and bottom, called a silo.—Prof. T. F. Hunt. The information contained in the following chapter was prepared by Mr. A. Morrison Hay, of the Public Works Department, Union of South Africa, and appeared originally in Farmers’ Bulletin 59 of the Transvaal Department of Agri- culture; it was subsequently revised and published in the Agricultural Fournal of the Union of South Africa (Hay, 1). Mr. Hay has kindly given permission to reproduce it here, with certain alterations and additions which he has himself suggested. The present writer has omitted Mr. Hay’s paragraph describing silage, as the information has already been given in chapter xv. 743. Hestortcal—Silos, or chambers for the storage and preservation of food, have been in use in one form or other in various countries from very early times. At the time of Pliny, in France, Spain and other parts of Europe, grain was pre- served in trenches, dug in the ground; he mentions in certain of his writings that ‘‘the best plan of preserving grain is to lay it up in trenches dug in a dry soil, called siri, as they do in Cappadocia, Thracia, Spain, and in Africa”. This method of preserving grain was not confined to the East, as at the time of the discovery of America by Columbus the natives were in the habit of storing grain in pits, and certain tribes continue the practice to the present time. The ancient Egyptians, as we learn from Scripture, stored sufficient grain in the seven years of plenty to serve themselves and other nations during the seven years of famine that followed. The Egyptian silos were evidently of a more improved and permanent nature than the rude trenches above referred to, as Wilkinson in his work, ‘The Ancient Egyptians,” states that “some of the rooms in which they housed the grain appear to have had vaulted roofs. These were filled through an aperture near the top, to which 770 THE CONSTRUCTION OF MODERN SILOS 771 the men ascended by steps, and the grain, when wanted, was taken out from a door at the base.”’ It is important to note that the early silos were used prin- cipally for the preservation of corn and other dried cereals, for indefinite periods, and there are instances of corn having been preserved in good condition for the long period of 200 years. On the other hand, the silos of the present day are used more particularly for the storage of green fodder, to provide food for cattle during the winter months, or for a year at most. Food thus preserved may, however, be kept indefinitely, provided that air is not allowed to enter the silo. For this reason it is important that the sides and floor should be perfectly air-tight, and it is also essential that the sides be smooth and vertical, so that the silage may settle uniformly and compactly, leaving no vacant spaces for the accumulation of air. The word “ silo,” taken from the Greek ‘“siros”—a pit for holding grain—is the name now applied to any air-tight chamber formed for the preservation either of dried grain or green food. It may be simply a trench or pit dug in the ground in any dry position, into which the silage is filled and weighted down with planks, earth, or other material. More commonly it is a structure of wood, metal, brick, stone, or other building material, or a combination of these, built en- tirely above ground or extending down only a few feet, and standing either by itself or forming a part of the other farm buildings. The origin of structural silos on the modern plan is of very recent date, but the progress of siloing has been so rapid and successf{ul—in America, at any rate—that in the short period of thirty years, over a hundred thousand silos were built in the United States alone, and the number is steadily increasing. Bey 744. Form.—In form the silo may be built either square, rectangular, octagonal, or circular on plan. If forming part of a scheme of buildings its shape would probably be decided by the position it occupied and the space available, but if standing alone, either form could be adopted at will. The round silo (Fig. 241) is more favourable to the even and compact settling of the silage owing to the absence of corners and, consequently, more favourable to its perfect preservation. On the other hand, it does not fit in with other buildings so readily and with- 49° CHAP. XVI. CHAP. XVI. 92 MAIZE out loss of space, as a square or rectangular silo, and for this reason the latter are likely to be more frequently built. The same remarks would also apply to octagonal silos. If standing alone, however, no objection can be raised either to the round or the octagonal form of silo, and as the proper preservation of the silage is the main object to be attained, one or other of these Fic, 241.—Filling round silo in the United States. forms should be adopted in preference to the others. The main objection to square or rectangular silos is the presence of corners, which prevent the silage from settling uniformly and compactly, thus causing waste of space and creating un- desirable accumulations of air. This defect may be remedied to a considerable extent by having the corners well rounded. Of the two, the square silo is perhaps the better form, and probably somewhat cheaper than a rectangular one of equal capacity, as the wall space is rather less, 745. Sise—-Where a large quantity of silage is required it is advisable to have it stored in two or more moderately-sized CHE CONSTRUCTION OF MODERN SILOS 173 silos in preference to one very large one. With very large CHAP. silos too much surface of silage is exposed while feeding ; and, *Y! if the height is increased to modify the surface atmensene beyond a reasonable limit, excessive labour is involved in the working. With rectangular silos the difficulty can be obvi- ated by having one or more partitions dividing the silo into two or more square, or nearly square, compartments. The height above ground should not exceed 20 to 25 feet, and the depth under ground should not be more than 5 or 6 feet the limit from which a man can conveniently lift the forage. ) Whatever form is adopted for the silo, it should always be Fic. 242.—Filling twin tub-silos, Australia. built with a greater capacity than would be necessary it it could be filled at once to the top with good silage, as con- siderable allowance has to be made for waste from settling and from spoiled silage. Even if material is added a second or third time there will still be a certain amount of waste space to be allowed for. One-fifth is considered a low estimate for loss of space and waste silage. 746. Capacity.—As a basis on which to ascertain the size of silo necessary to hold food for a certain number of cattle for a certain period of time, 40 lbs. may be taken as the average amount of silage required to feed one animal per day, and 40 lbs. may also be taken as the average weight of a cubic foot of silage. Therefore, if the number of animals is CHAP. XVI. 174 MAIZE multiplied by the number of days during which they have to be fed, the result will equal the number of cubic feet of silage space required. Supposing thirty animals have to be fed for six months, or 184 days, the total amount of space required would be 30 x 184 = 5,520 cubic feet. To find the number of cubic feet in a square or rectangular silo multiply the length in feet by the width, and then by the height, and the result will give the total number of cubic feet. Thus, if a square silo measures 14 feet each way on plan and 25 feet in height, the cubical contents would be 14 feet x 14 feet x 25 feet = 4,900 feet. In the case of a round silo, multiply the square of the diameter in feet by the height, and then by ‘7854. For example, if a round silo measures 14 feet in diameter and 25 feet in height, the cubical contents would be 14 feet x 14 feet x 25 feet x -7854 = 3,848 feet approximately. The following table (No. CXXXII) shows the capacity of silos of different dimensions and the acreage of maize re- quired to fill them, on the basis of a crop of 15 tons (Colonial) to the acre :— TaBLE CXXXII. CAPACITY OF SILOS. Acres of Crop necessary Capacity in Tons. (15 Tons per Acre). | Dimensions. ie | 28 | | 20' | 3 12' x 20’ | 30 | 3 | T2! x 24! 49 3% | | 12’ x 28/ 60 | 4 | 14’ x 22’ 61 44 | | 14’ x 24’ 67 45 | 14’ x 28" 83 | 53 | | 14’ x 30° 93 6 16’ x 24’ 87 62 | 16’ x 26’ 97 | 7 16’ x 30’ I1g | 8 18’ x 30' 151 | To} | x 124 18’ x 36’ 189 | 4 | 747. Position.—As regards position, the silo should be placed as near as possible to the centre of feeding, to minimize the labour of carrying food to the various mangers. In round or octagonal farm buildings, such as are common in America, THE CONSTRUCTION OF MODERN SILOS 7175 the silo usually takes the same shape as the main building, cHap. and is placed in the centre, where it occupies a convenient *V!. position from which to feed the various animals stalled around the building. A circular silo may be situated at a corner of the main building, where it not only proves a useful adjunct but also forms a very pleasing feature. A convenient arrange- ment is to have the doors of the silo opening into a passage- way, or into the feed store or mixing room, which usually occupies a central position, A saving in the cost of erection Fic. 243.—Filling square stone-silo, Irene, Transvaal. can often be effected by utilizing one or more of the walls of the main building in the construction of the silo. 748. Materials —The material to be used in the construc- tion of a silo would probably be decided by local conditions. That most readily obtained in the locality in which the silo is to be built would naturally be chosen, provided it fulfils the conditions required for a good silo, Metal has been tried in some countries, but has not been found satisfactory, on ee of the initial cost and the readiness with whee it gies se : corroding action of the silage juices. In America wood has CHAP. XVI. 776 MAIZE hitherto been very largely used in the construction of silos, on account of the abundance of timber in that country and the comparative ease and cheapness of erection, and wood is con- sidered one of the best materials for the preservation of silage. It is not likely, however, to be much used in South Africa, on account of its scarcity and want of durability. For durability, either reinforced concrete, stone, or brick, or a combination of these materials, is to be recommended. Reinforced concrete is probably the best material, especially for circular silos, but requires more skilled labour in the con- struction, and is consequently more costly. Stone or brick structures require less skill in building, and as either one or other of these materials can usually be readily obtained throughout South Africa, they are likely to enter more largely into the construction of silos. Whatever material is used—whether concrete, stone, brick or wood—the foundation should always be of some material that is not subject to early decay, more particularly if it extends for some depth below the surface of the ground. In a dry soil a good quality of brick or stone built with lime mortar is sufficient, but if there is a tendency to dampness, either cement concrete or stone, built with cement mortar and plastered on the outside with any damp-proof composition, forms the best foundation. It is important that the founda- tion should go down to a solid bottom, and it should extend a few inches above the highest point of the ground, and be covered on top with a damp-proof course to prevent moisture from rising to the structure above. 749. Reinforced Concrete-—The methods of building with either brick or stone are too well known to everybody to require any explanation, but concrete construction being less familiar, a short description of the materials used and methods adopted in the erection of reinforced concrete silos may be useful. The walls, only, need to be described, as the foundations, floor, roof, doors and other parts can be practically the same as for brick or stone silos. The concrete should be composed of one part best Portland cement, two parts clean sharp sand and four parts stone broken to such a size as will pass through a ring of ¢ inch diameter, all thoroughly mixed together to the proper consistency with clean water. The reinforcement should con- THEA CONSTRUCTION OF MODERN SILOS 777 sist of round iron rods arranged both vertically and horizontally and embedded in the concrete near the outer surface of the wall. The vertical rods should be { inch in diameter, spaced from 12 inches to 15 inches apart and extending from founda- tion to top of wall. The horizontal rods should be 4 inch in diameter, spaced from 9 inches to 12 inches apart. They should be in as long lengths as possible, hooked together at ends so that each row forms a continuous band round the silo. The horizontal and vertical rods should be securely wired together at all points of intersection to form a rigid network of iron. There are of course other methods of reinforcement, but the above with a 6-inch thickness of concrete is simple and sufficient for any silo up to say 18 feet in diameter and 30 feet in height. To erect the walls it is necessary to have moulds or forms curved to the same radius as the silo, one convex on face placed inside the wall, and one concave on face placed out- side the wall, the two 6 inches apart from each other with the reinforcing rods between.