2)
Se ee AE eh eater oy?
he i pn fe
o
ALBERT R. MANN
LIBRARY
New York STATE COLLEGES
OF
AGRICULTURE AND HOME ECONOMICS
AT
CORNELL UNIVERSITY
Ferns: British & foreign.The history, or
Cornell University
Library
The original of this book is in
the Cornell University Library.
There are no known copyright restrictions in
the United States on the use of the text.
http://www. archive.org/details/cu31924001743032
FERNS: BRITISH & FOREIGN.
THEIR HISTORY, ORGANOGRAPHY, CLASSIFICATION,
AND ENUMERATION.
WITH
A TREATISE-ON THEIR CULTIVATION,
ETC. ETC.
BY JOHN SMITH, A.L.S.
ACAD. CHS. NAT. CUE, B0C.,
aND EX-CURATOR OF THE ROYAL BOTANIC GARDENS, KEW,
LONDON:
ROBERT HARDWICKE, 192, PICCADILLY,
CONTENTS.
PREFACE.,......... en re sian eamERtS oq 1®
I. History oF tHE Inrropuction or Exoric Ferns ... 1
II. OreanocraPHy, EXPLANATION OF THE TERMS USED
1n Descrisine FErns ...... Ginx thicesesses wiseaen saviavaceve AA
‘TI. CLASSIFICATION OF FERNS EXPLAINED .......0005 Gace 56
{V. Geyeric CHARACTER AND ENUMERATION OF CULTI-
VATED SPECIES ........ sjaaisolvarnaed Susana Fengte Pemettenawed 73
V. Erymotoey, tor Mranine or THE GENERIC Nawés... 293
VI. Tue CuttivaTion oF FErRyS.....
I. Pretiminary Remarks.
II. SpectaL on Pot CULTIVATION ..........ceeeeeeee .. 808
TIT. Navturat CULTIVATION ......cccececeeeeeeeueeee sarhOae
IV. Warpian Case CULTIVATION ......... sisscneat .. 354
V. PROPAGATION ....... weeiaats nadaenneWaeerGindeaeste waney OOF
VII. List or AvTHors anD Books QUOTED IN THIS
WORK: =tavanestsesienibeaessmadaniesopamihivsaiecemnimaeeinag Ble
VIII. Generat InpEx or GENERA, SPECIES, AND Syn-
ONYMES ....... sec teeee ee eeeneeneeaaes aaltleman@eaaitein ouiectnine OO
IX. INDEX OF SPECIAL 'PERMS ..cecssseececeeeseeeeseeseseeees 411
=,
> J
Re 4
WF
”, Fitch, det,
&.
Ww Rugte.
EXPLANATION OF THE PLATE.
ORGANOGRAPHY, page 44.
Annulate Sporangia—
Fic. 1. Sporangium with a vertical ring, mag. 100 diameters.
2. + horizontal ring, mag. 100 diameters.
3. apical ring, mag. 100 diameters.
(a) Spores of sacle highly magnified, 200 and 300 diameters.
Lxannulate—
Fic. 4. Sporangia (two) free, opening by a vertical slit,
mag. 25 diameters.
5. Sporangia united (synangium), Gpautiig by pores, mag. 7
diameters.
6. Sporangia united (synangium), opening by slits, mag. 9
diameters. ,
(a) Spores of each highly magnified, 300 diameters.
Indusia—
Fig. 7; Indusium peltate orbicular, slightly magnified.
8. 7‘ lateral reniform, ditto.
9. ” calyciform, ditto.
10. 9 linear, interiorly attached, _— ditto.
11. ” valvate, ditto.
12, universal, ditto.
PREFACE.
EARLY ten years have now elapsed since my
N “Catalogue of Cultivated Ferns” was pub-
lished. During that period many new and fine species
have been introduced to the gardens of this country.
I have therefore been induced to draw up the follow-
ing enumeration, including in it all the species that
have come under my notice in a living state, either by
the evidence of plants in the Kew collection or by
specimens from living plants sent me from various
sources.
Thinking it might be interesting to Fern growers,
T have drawn up a brief history of the introduction
of Exotic Ferns from the earliest records to the present
time.
In order to assist students in the study of Ferns,
an explanation is given of the principal organs and
terms used in describing them, as well as remarks
on their classification. I have also given an alpha-
betical list of the generic names, accompanied with
the derivation of each name.
Great complaints are frequently made, and not with-
x PREFACE,
out good reason, of the many names given to the same
plant; to assist in correcting this, I have drawn up a
complete Index of the species and synonymes noticed
in this work ; therefore, by turning to the page referred
to in the Index, the name will be ‘found either with a
number before it showing it to be the adopted name, or
following the same as a synonyme, or what it has been
and is still called by some writers and cultivators.
Although this Index contains nearly three thousand
names, yet, it must be understood, I have only taken
up the synonymes that I consider most essential ;
those who desire to enter more fully into this subject,
may consult the “ Index Filicum” of Mr. T. Moore—
a work which, when completed, will be a treasure to
pteridologists.
With regard to the wood-cuts, I have to explain
that about half of the number were not specially pre-
pared for this work; these, in general, represent a
portion of the fronds of their natural size,"in some
cases without fructification, the venation only being
given. The drawings for the other half have been made
principally from specimens in my herbarium or from
living plants. In many of them a magnified portion
of the frond is given, showing the character of the
genus more distinctly.
As a companion to the scientific enumeration, I
have given a treatise on their cultivation, which occu-
pies a considerable space in this book, and it is hoped
will be of service to the Fern-growing public,
PREFACE. xi
In drawing up this treatise I have been greatly
assisted by Mr. Henry Prestoe and Mr. William Gower
(both recently foremen in the Fern department at
Kew), two highly scientific and nature-observing
practical cultivators.
In conclusion, I have to thank Dr. Berthold Seemann
and R. Heward, Esq., for looking over the proofs as
they passed through the press. Several causes have
occurred to delay the publication of this work; the
principal one being a partial failure of my sight,
incapacitating me from much use of the pen and
examination of new garden Ferns, which I trust will
be accepted as my excuse for any errors or omissions
that may be found in the following pages.
J. SMITH,
Ex-Curator, Royal Botanic Gardens,
Kew.
Kew, May, 1866,
FERNS: BRITISH AND FOREIGN.
HISTORY OF INTRODUCTION OF EXOTIC FERNS.
HOUGH Ferns now occupy a conspicuous place
in our gardens, and are in high favour with
cultivators, it is only in comparatively recent times
that they have been brought into notice. During the
last - century certain classes of plants came into
fashion, and after-a season of popularity again fell
into disrepute. Thus: Tulips were once the rage.
At the time of the establishing of the several pro-
vincial Botanic Gardens, all of which were founded
upon a strictly botanical footing, though many of them
have now, to a greater or less extent, degenerated
‘into places of amusement, the plants in greatest demand
were those of our New Holland and Cape colonies,
’ principally the Heaths, Proteas, Aloes, and their
kindred. In after-years dealers obtained large prices
for Cactuses; but, with the exception of a few of
the easily-grown and most showy kinds, these are
now scarcely saleable. Still more recently the magni-
ficent-flowering Orchids were promoted to the first
place in our gardens; and though these may still be
said to maintain their position, the expense attending
B
2 FERNS: BRITISH AND FOREIGN.
their cultivation is so great that they are for the
most part confined to the gardens of the wealthy.
Ferns, on the contrary, may, as a general rule,
be grown in a comparatively inexpensive manner.
The discovery made by Mr. N. B. Ward, that these.
plants can be grown to great perfection in small
ornamental closed cases (now well known as “ Wardian
Cases”’), suitable not only for the drawing-rooms of
the wealthy, but for humbler dwellings, renders it pos-
sible for amateurs to indulge their love of Ferns without
going to the expense of erecting hothouses and em-
ploying a staff of gardeners; and it is to be hoped
that this will be the means of retaining them in favour
and spreading them still wider.
The enumeration in the following pages shows that
at the present time above nine hundred exotic species of
Ferns are cultivated in the various public and private
gardens in this country; and of these by far the
greater number have been introduced during the last
quarter of a century. A very large, indeed almost
a complete, collection of them may at present be
seen in the Royal Botanic Garden at Kew, where,
during forty years, I devoted attention to their
cultivation, and to the study of their generic and
specific distinctions, using every endeavour in my
power, assisted by the extensive influence of the
eminent Director, Sir W. J. Hooker, to introduce
new species, both by raising them from spores
taken from my herbarium, and through correspon-
dence with persons residing in our Colonies and the
Superintendents of Botanic and other gardens on the
Continent. Being well acquainted with the latter
branch of the subject,—the introduction of new
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 3
species,—I have thought that a few pages devoted to
the history of these plants in its connection with our
gardens might prove acceptable. My own personal
knowledge dates from about the year 1822; and for
information respecting those in cultivation previous
to that time, I have taken for my guide the most im-
portant and most reliable of the garden catalogues.
The first work upon which dependence c can be placed
for the early-introduced species is the so-called second
edition of the “ Hortus Kewensis,” published by the
younger Aiton, in 1813; I say the so-called, for there
were in reality two previous editions—one by Dr. John
Hill, in 1768, and a second by William Aiton, in
1789. In the first of these only ten exotic species
are recorded as being in the garden at Kew, which
then belonged to the Princess of Wales, but the
dates of their introduction are not given; and in
the second, that of the elder Aiton, thirty-four, all
of which have the name of their supposed introducer
attached to them, and the date at which they were
introduced. Ihere give preference to the last edition,
as I know, frorh personal intercourse with the com-
pilers, that great care was exercised in its prepara-
tion, particularly with regard to the rectification of
dates, and I shall take it as the basis of my remarks.
‘But, in the first place, it is necessary to say a few.
words in defence of that work. It has been
asserted that a considerable number of. the plants
enumerated in the “ Hortus Kewensis” never existed
in a living state, either in the garden at Kew or
elsewhere in this country. This statement, however,
is merely supposition, and rests upon questionable
authority. In the early days of Kew Gardens, large
B 2
4; FERNS: BRITISH AND FOREIGN,
sums of money were devoted to the payment of bota-
nical collectors, and great exertions were made, under
the patronage of Sir Joseph Banks, to stimulate the
introduction of new and rare plants, by inducing the
commanders of ships of war and East-Indiamen to
take an interest in the subject. By these means a
very large number of plants were actually introduced
into the country in a living state ; but the Department
under whose charge the garden then was, took no
steps to provide proper accommodation ; and this,
together with the very frequent change of foremen,
led, as a natural consequence, to the death of the |
plants.
Taking, then, the third “ Hortus Kewensis,” and
excluding our indigenous species, I find that the total
number of “Garden Ferns”? introduced previously to
the year 1813 amounts to as many as eighty-three. The
merit of being the first- introducer of these plants
belongs to Mr. John Tradescant* the younger, who in
early life made a voyage to Virginia; and I find it
recorded in Parkinson’s ‘“ Theatrum Botanicum,”
published in 1640, that upon his réturn from that
country in 1628 he brought with him, amongst other
rare plants, the Cystopteris bulbifera and Adiantum
pedatum. These, therefore, must be regarded as the
nucleus of our present large collections. At first
the progress seems to have been exceedingly slow, for
between the time of Tradescant and the close of the
seventeenth century, only five additional species were
introduced ; viz., Asplenium rhizophyllum and Onoclea
* John Tradescant had a Botanic Garden and Museum at
Lambeth.
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 5
sensibilis from North America, the former in 1680 and
the latter in 1699, in which year also Adiantum reni-
forme and Davallia Canariensts were brought from
Madeira, while the fifth, Blechnwm australe, was,
according to Plukenet, who figured it in the second
part of his “ Phytographia,” cultivated in the King’s
garden at Hampton Court as early as the year 1671,
when his work was published, the garden there
containing a considerable collection of rare plants.
During the next forty-two years no additions appear
to have been made, excepting the Phlebodium
aureum, which was introduced by Lord Petre at
some time prior to 1742, the date of his death, the
precise year being unknown. Progressing onwards,
Ido not find any more recorded until the year 1769 ;
but between that year and the commencement of
the following century sixty-eight species were added
to the eight already existing in our gardens. Out of
this large number, no less than thirty-seven were
brought home by Rear-Admiral Bligh, in H.M.S.
Providence, on his return, in 1793, from his second
voyage, undertaken for the purpose of introducing
the Bread-fruit and other useful trees into our West
Indian colonies. And, finally, during the first few
years of the present century, up to 1813, the date of
the publication of the “Hortus Kewensis,” seven
others were introduced.
. Asummary of the foregoing shows that upwards
of one-half of the Ferns known at the last-mentioned
date were West Indian species, forty-four having been
received at various times from those islands, the
majority through Bligh’s expedition. North America
and Madeira, with the neighbouring islands, stand
6 FERNS: BRITISH AND FOREIGN.
next as the largest contributors, fifteen having been
introduced from the former and twelve from the latter
country ; while of the remaining sixteen, four appear:
to have come from the East Indies, four from the
Cape of Good Hope,-three from New Holland, and:
one from St. Helena,—making in all eighty-three
species.
The next catalogue of garden plants worthy of
notice is the “Hortus Suburbanus Londinensis,”
published in 1818, only five years after the “ Hortus
Kewensis,” by Mr. Sweet, the Superintendent of the
then celebrated nursery of Mr. Colville, at Chelsea.
In it I find an enumeration of one hundred and eight
exotic ferns; but this work, like the similar more im-
portant “Hortus Britannicus,” brought out by the in-
defatigable Loudon in 1830, and which contains no less
than three hundred and thirty exotic Ferns, includes”
not only a considerable proportion of bad species,
but also a large number that did not really exist
in British gardens, many having been entered without
authentic evidence, and others added upon the mere
expectation that they might ‘shortly be introduced,—
expectations which, in many cases, have not been
realized to this day. No reliance can therefore be
placed upon either of these works, and I cannot
accept them as authorities.
During the latter part of the eighteenth century and
the commencement of the nineteenth, the only pri-
vate individuals who turned their attention, with any
amount of energy, to the introduction of new and
rare plants, were the long- and far-famed nurserymen
at Hackney, the Messrs. Loddiges; and to them
we owe the greater part, if not the whole, of the
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 7
Ferns existing at that period in British gardens,
and not included in the “Hortus Kewensis.”
Speaking from my personal recollection of the im-
portant collection of plants in the Hackney Nursery,
as it existed in the year 1825, I think it contained
about a hundred good species of exotic Ferns; but I
can obtain no earlier catalogue than one published
in 1849, in which two hundred and fifty-one are
enumerated,
In the year 1822 I found the collection of Ferns at
Kew extremely poor, especially as regards Tropical
species, very many of those introduced in previous
years having been lost, and very few new ones added.
Any person who remembers the hothouses in existence
forty years ago, will have but little difficulty in
accounting for the falling off of the Fern collection.
In those days hot-water pipes were unknown, and
the houses were exclusively heated by means of
brick flues, too often imperfectly constructed, and
the excessively dry and ungenial atmosphere thus
induced was quite unsuited for the good cultivation
or even for the mere preservation of these moisture-
loving plants. Nearly all the North American species
enumerated in the “‘ Hortus Kewensis” were growing
very finely in a north border, and most of the Madeira
species were also in existence; but, including these
and the few added since 18138, I cannot estimate the
entire Kew collection of exotic Ferns at that period
at more than forty species.
Between 1813 and 1846, when my first Catalogue
of the Ferns at Kew appeared, no reliable list
was published in this country. Several, however,
were brought out by Continental botanists, which
8 FERNS: BRITISH AND’ FOREIGN.
are of sufficient importance to be worthy of a passing
notice. The first of these in point of date is the
“ Enumeratio Plantarum Horti Regii Botanici Bero-
linensis,” by Wildenow, published in 1809, with a
Supplement by Schlechtendal, bringing it down to
1813. We are thus enabled to compare the num-
bers in the Kew and Berlin gardens at the same
period; which were eighty-three in the former, and
thirty in the latter, including eleven not known at
Kew. During the succeeding nine years more atten-
tion appears to have been given to Ferns at the
Berlin garden; for Link, in his first ‘‘ Enumeratio,”
in 1822, describes ninety-one exotic species, which
is more than double the number then existing at
Kew. After this the increase in number was still
more rapid; for in the second edition of Link’s
“ Enumeratio,” published in 1883, no less than two
hundred and thirty-nine are described; and in the
third, in 1841, two hundred and fifty-eight, exclusive
of varieties.
By this time, however, the collection at Kew had
received large additions, both through importations
of living plants and by raising from spores. In
1845 it was so. extensive that I was induced to
draw up a classified enumeration, which was pub-
lished as an appendix to the Botanical Magazine for
1846. The number of exotic species there enume-
rated is three hundred and forty-eight, and I do not
think many were to be found in other gardens in
this country which were not at Kew, so that the Kew
list may be taken as a guide to the number then
in British gardens generally.
Four years later, Kunze, of Leipzig, contributed to
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 9
the 28rd-volume of the “Linnea” an alphabetical index
of the Ferns cultivated in European gardens, and in
this the large number of eight hundred and forty-three
exotic species, are enumerated; but the authorities
upon which a very considerable portion of these were
inserted cannot be relied upon, many names having
been taken from such catalogues as those of Sweet,
Loudon, &., and I am therefore obliged to con-
clude that the number given by Kunze as living in
Europe in 1850 is greatly exaggerated. This con-
clusion, too, is confirmed by the fact that in 1857,
after I had by correspondence become acquainted with
the collections in the principal gardens on the Conti-
nent, and after that at Kew had obtained most of their
novelties by means of exchange, I could, in my
“ Catalogue of Cultivated Ferns,’ enumerate only five
hundred and sixty exotic species as known in British
gardens. Since the last-mentioned year, the con-
stantly increasing demand for Ferns consequent upon
their wider spread cultivation, has greatly stimulated
the introduction of new ones, and our collections
have increased at the rate of about fifty species a
year.
It now remains to say a few words regarding the
means by which these plants have. been obtained, and
the persons who have been most active in introducing
them, taking the Kew collection as a foundation.
Firstly, with reference to the paid collectors employed
in various parts of the world, directly or indirectly in
the service of the Royal Botanic Garden, Kew, and to
whom that garden is indebted for additions to its Fern
collection. It would appear that so long back as the
year 1775 Mr. Francis Masson, one of the earliest, if
10 FERNS. BRITISH AND FOREIGN.
not the earliest, collector sent out from Kew, and
who succeeded in introducing large numbers of Cape
Proteacee and LHricacece, sent home several Ferns
from the Cape of Good Hope and Madeira. This col-
lector proceeded to the Cape in 1774, and came home
by way of Madeira about the year 1778, returning
again in 1736, and remaining at the Cape during the
nine following years. arly in the present century
Mr. George Caley, who was originally a horse-doctor,
residing near Birmingham, but acquired a love for
plants through collecting herbs, was sent out by
Sir Joseph Banks to New South Wales, and to him
we owe Platycerium alcicorne, Doodia aspera, and
Davallia pyxidata, the first introductions from Aus-
tralia, received about the year 1808. The next col-
lectors to whom the garden was indebted for Ferns,
are the Messrs. Allan Cunningham and James
Bowie. They left Kew in 1814, on a botanical
expedition to Brazil, where they remained exploring
the country and sending home large collections till
1816, when the former proceeded to New South Wales,
and the latter to the Cape of Good Hope. No living
Ferns appear to have resulted from the Brazilian ex-
pedition ; but several Australian species and one or
two from Norfolk Island were received from Mr.
Cunningham, and two or three from Mr. Bowie from
the Cape.
Several other collectors were employed in the ser-
vice of these gardens, when under the Directorship of
Mr. Aiton, such as Messrs. Barclay and Armstrong ;
but I can trace no Ferns to them, nor, with certainty,
to David Lockhart, a gardener from Kew, who accom-
panied the ill-fated expedition of Captain Tuckey up
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 11
the Congo, in 1816, though I believe it possible that
some of the first-known Western-African species are
attributable to him. Lockhart was one of the few
who did not fall a victim to the deadly climate of that
country. After lying a long time in the hospital at
Bahia, he returned to England, and shortly afterwards
received the appointment of Superintendent of the
Botanic Garden in Trinidad, where he died in 1845,
after a service of a quarter of a century in that island,
during which period he sent many fine plants to Kew.
The principal collectors employed directly or indi-
rectly in the service of Kew during the Directorship of
Sir William Hooker, and to whom the garden is in-
debted for any additions to its Fern collection, are
Messrs. Purdie, Seemann, Milne, Barter, and Mann.
The first of these, Mr. William Purdie, was engaged as
collector in 1848, in which year he was despatched to
Jamaica, and after spending several months in the
exploration of that island, and forwarding many
valuable plants. to Kew, he proceeded to Santa
Martha, and was employed for more than two
years in various parts of New Granada. To him we
are indebted not only for numerous showy flowering
plants, but for a considerable number of our garden
Ferns.
Upon the death of Mr. Thomas Edmonston,* Mr.
(now Dr.) Berthold Seemann was appointed his suc-
cessor, as botanist to H.M.S. Herald, and joined that
vessel at Panama, in January, 1847, remaining with her
until her return to England in June, 1851. Previously
* Mr. Edmonston was shot by the accidental discharge of a
rifle, at Sua Bay, Ecuador, January 24, 1846.
12 FERNS : BRITISH AND FOREIGN,
to joining the Herald, he explored several parts of
Panama and Veraguas, and while on board the
Herald visited various parts of the western opast of
America, between Lima on the south and California
on the north, occasionally making long journeys
inland, particularly in Peru, Ecuador, and Mexico.
He likewise accompanied the Herald in her three
voyages to the Arctic regions in search of the
ill-fated Sir John Franklin; visited twice the Sand-
wich Islands; and returned home by way of Hong-
Kong, Singapore, and the Cape of Good Hope. Not-
withstanding the few facilities afforded by surveying
expeditions for collecting living plants, Dr. Seemann
succeeded in introducing some interesting ones to -
our gardens; and among Ferns we may mention two
very remarkable ones, Deparia prolifera from the
Sandwich Islands, and Dictyoxiphium Panamense from
Panama.
On the Herald being recommissioned in 1852, Mr.
William Milne was appointed assistant-botanist, for
the special purpose of collecting plants for the Royal
Gardens. During the six years he was attached to
the Herald he visited New Caledonia, the Fiji, and
other Polynesian islands, as well as many other places
in the southern hemisphere.
The Admiralty having, early in 1857, determined
deosunnding out a second exploring expedition up
the Niger, under the command of Dr. Baikie, R.N.,
Mr. Charles Barter, a zealous and intelligent young
gardener, was appointed to accompany him in the
capacity of botanist, and to collect plants for Kew.
The officers of the expedition proceeded by way
of Sierra Leone, and joined the Duyspring, a small
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 13
vessel fitted for river navigation, at Fernando Po; but,
unfortunately, after they had penetrated up the river
as far as Nupe, their ship was wrecked, and they
were obliged to form a camp on the banks. Poor Barter,
~ however, fell a victim to the deadly climate in July,
1859, Asa collector he was indefatigable in the dis-
charge of his duties, and discovered a large number of
undescribed plants, including several new genera,
one of which has been dedicated to his memory under
the name of Barteria, But, owing to the mishaps
attending this expedition, and the difficulties of
transit, no living plants were received from the
interior of the country, but several Ferns were trans-
mitted from Sierra Leone and Fernando Po.*
No sooner had the news of Barter’s death arrived,
than Mr. Gustav Mann, undeterred by his fate,
volunteered to fill the vacancy, and- was appointed
accordingly. Being unable, from want of means of
communication, to join Dr. Baikie, he was employed
for three years in exploring the island of Fernando
Po and the African coast, in the neighbourhood
of the Camaroons and Gaboon rivers. In spite
of the difficulties and dangers attending the under-
taking, he made several ascents of the lofty Clarence
Peak of Fernando Po and of the Camaroons moun-
tain on the African mainland. On the former, at
an elevation of 5,000 feet, he found a fine new
Cyathea, forming groups, with stems rising to a
height of 30 feet. We are likewise indebted to him
* Since the above was written, news has reached this country
of the death of Dr. Baikie at Sierra Leone, in January, 1865, whilst
on his homeward voyage, -
14 FERNS: BRITISH AND FOREIGN.
for many other rare and interesting living Ferns, and
fine sets of dried specimens.
Besides contributions from special collectors, the
Fern collection at Kew has been largely increased
through the instrumentality of the officers of our
numerous Colonial Botanic Gardens, and likewise by
many private individuals residing in foreign countries ;
and as the exertions of these gentlemen are worthy of
being recorded, I give a few particulars respecting
them, adopting a geographical arrangement for the
sake of brevity.
The first Colonial garden from which, so far as I
am aware, Ferns were received at Kew, was that of
Seylon, Mr. Alexander Moon, the Director, having,
in 1824, sent home a collection of plants, amongst
which was Niphobolus costatus. But the first persou
who forwarded any considerable number from that
island was the lamented Mr. George Gardner, well
known to botanists as a botanical traveller in Brazil,
who was Director of the Ceylon garden between 1844
and his death in 1848. This gentleman was succeeded
by Mr. (now Dr.) G. H. K. Thwaites, the present able
Director, and to him also the Kew collection is greatly
indebted for a large number of rare and beautiful
species; such as NSehizoccena sinuata, Asplenium
radiatum, Actinostachys radiata, Helminthostachys
Zeylanica, and many others.
Though Continental India is extremely rich in Ferns,
it has, singularly enough, contributed very few to our
gardens, no persons in that country having devoted
themselves specially to the subject; indeed most of
those received thence have been accidentally imported
along with Orchids, including the half-dozen species
HISTORY OF INTRODUCTION OF EXOTIC FERNS, 15
recorded in my first Enumeration as coming from
Dr. Wallich, the only Superintendent of the Calcutta
garden who has the credit of having introduced any.
Indeed, with the exception of those from Ceylon, Kew
has received very few Ferns from Asia and. the
adjacent islands, most of those now in cultivation
having been introduced by nurserymen or through
Continental gardens. Two or three were brought
from Hong-Kong, in 1850, by Mr. J. C. Braine,
including one which proved to be a new genus, and
to this I gave the name of Brainea in honour of its
introducer.
Another tropical ee in the Kastern hemisphere,
whence large additions have been made to the Fern
collection at Kew, is the Mauritius. The Botanic
Garden in that island has long enjoyed the reputation
of possessing a fine set of plants; but until the year
1852, when. it came under the able management
of the present Director, Mr. James Duncan, very
little correspondence was kept up with the gardens
of Europe. Mr. Duncan has, at considerable risk,
ransacked the forests of the island in quest principally
of Ferns, and has been very successful in transmitting
living plants to this country, enriching our gardens
with many fine species.
The “Synopsis Filicum Capensis” of Pappe and
Rawson shows that the Fern Flora of Southern Africa
is extremely rich; but up to the present time we
possess scarcely a dozen Cape species in our gardens,
and most of them have been raised from spores.
Alsophila Capensis and Lomaria Capensis were intro-
duced in 1845 by Mr. Charles Zeyher, as also were
Lastrea athmantica and Cyathea Dreget from Natal
by Mr. J. Plant.
16 FERNS: BRITISH AND FOREIGN,
Progressing westwards we come to the two strangely
isolated islands in the Southern Atlantic, St. Helena
and Ascension. From the former of these we have As-
plenium compressum, introduced by Mr. Thomas Fraser
in 1825, and Asplenium reclinatum, brought home by
Dr. J. D. Hooker on his return from Sir John Ross’s
Antarctic expedition in 1844; together with Lomaria
alpina and L. Magellanica from the Falkland Islands ;
while from Ascension Mr, Wren sent numerous fine
plants of Maratiia purpurescens in 1848.
From Australia several individuals: have been con-
tributors. Grammitis Australis was received from the
Sydney garden in 1833, when under the direction of
Mr. Richard Cunningham; and Mr. Charles Moore,
the present Director of that garden, has also intro-
duced several, including Trichiocarpa Mooret, from
New Caledonia, while to Mr. Bidwill we owe the
curious Platycerium grande. But some of the most
beautiful of the Australian Ferns, such as the Gilei-
chentas, were transmitted to this country by Mr.
Walter Hill, the able Director of the Botanic
Garden of Brisbane, in the rapidly rising colony of
Queensland, who obtained them during his stay in
Sydney in 1850. Two species of Gileicheniacee were,
however, previously known in our gardens,—the Glei-
chenia nuicrophylla and G. flabellata, both of which,
together with several other Ferns, were sent from
Tasmania, in 1845, by Mr. Ronald Gunn.
About the year 1841 or 1842; some very fine Ferns,
including two Tree-Ferns, the Dicksonia squarrosa and
Cyathea medullaris, were brought from New Zealand,
where they had been collected by Mr. J. Edgerly, a
gardener, who had proceeded to that country on
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 17
speculation, and who was the first to introduce tho
beautiful Veronica speciosa. Others have also been
sent from New Zealand by the Rev. William Colenso,
and by the late Dr. Sinclair, R.N., the beautiful little
Trichomanes reniforme being one of those due to the
latter gentleman.
Turning next to the West Indies, we commence
with the island of Jamaica, whence more Ferns have
been received at Kew than from any other part of
the Western hemisphere. The person to whose
energy and perseverance this is mainly due is Mr.
Nathaniel Wilson, the Island botanist and Director
of the Botanic Garden. He has been a resident
in the island for upwards of twenty years, and
during that time has thoroughly explored the Blue
Mountains and other districts rich in Ferns, liberally
forwarding to Kew the results of his numerous
journeys. Among his earliest contributions was the
beautiful Tree-fern, Cyathea arborea, which, though
recorded in the “‘ Hortus Kewensis” as having been
brought home by Admiral Bligh in 1798, had long
been lost to our gardens. Within the last few years
he has succeeded, after many failures, in transmitting
numerous species of Trichomanes and Hymenophyllum,
which now form so conspicuous a feature in the
present rich collection. The other contributors from
the same island are, in 1851, Mr. George Manson,
and in 1854 and following years, W. T. March, Hsq.,
the latter gentleman sending several arborescent
species as well as Hymenophyllece, and others.
Numerous fine species, including several Oyatheas
and Alsophilas, were received in 1855 and 1856 from
the French island of Martinique, where they had been
Cc
18 FERNS: BRITISH AND FOREIGN.
collected by the Director of the Garden, M. Belanger.
A considerable number of rare species were likewise
sent from Dominica in 1853, and several following years,
by Dr. Imray; such as Hemitelia Imrayana, Neurocallis
prestantissima, Elaphoglossum undulatum, and the
true EH. longifolium, &c. While from Trinidad we are
indebted to the Island botanist, the late Dr. Criiger,
for Schizca elegans, Saccoloma elegans, Amphidesmium
rostratum, Hymenostachys diversifrons, Trichomanes
pennatum, and other equally rare species. His suc-
cessor, Mr. Henry Prestoe, who previous to his appoint-
ment last year was foreman of the collection of Ferns
at Kew, has already transmitted a large collection of
rare Ferns, and in the finest condition, proving that
they had been collected and put up by one who
perfectly understood their nature. The situation he
now holds will enable him to be of great service in
transmitting new plants to this country.
Comparatively few, considering the richness of its
Fern Flora, have been received from tropical America;
Dr. Gardner and J. Wetherall, Esq., in Brazil; H.
Cadogan Rothery, Esq., in Guiana; Mrs. Colonel
McDonald, in Honduras ; and Mr. Wagener, in Vene-
zuela, being the principal contributors.
From the above it will be seen that a large number
of Ferns have been introduced through the agency of
the garden at Kew; but, besides these, a very con-
siderable number.are due to the exertions of some of
our leading nurserymen, who, in consequence of the
great demand for, and the large prices realized by, the
finer and rarer species, have imported them, either
direct from their native countries, or from the Con-
tinent, where, as will be presently noticed, a good
HISTORY OF INTRODUCTION OF EXOTIC FERNS. 19
many species not previously known in the gardens
of this country have been introduced.
I have already alluded to the Messrs. Loddiges, of
‘Hackney, as having at an early period turned their
attention to Ferns, and as being the earliest to form ,
a collection of them. But the only nurserymen whose
names are recorded in the second edition of the
“Hortus Kewensis” are the old-established firm of
Messrs. Lee & Kennedy, of Hammersmith, who are
stated to have introduced Polypodiwm asplenifolium
and Asplenium monanthemum in 1790: in later times
the Messrs. Lee have imported several from New
Zealand. Other New Zealand species have been
brought into notice by Mr. Standish, of Bagshot,
they having been collected in New Zealand by Mr.
J. Watson, now a nurseryman at St. Alban’s, and
who still continues to import. Several sent from
Japan by Mr. Fortune have likewise been sent out
from Mr. Standish’s nursery. To the Messrs. Low
& Sons, of the Clapton nursery, we are indebted for
some rare Bornean and Malayan species, collected by
Mr. Hugh Low, jun., and amongst others for the
remarkable Arthropteris obliterata, called Lindscea
Lowii in the gardens, and the little curious Leucostegia
parvula; but more especially many rare species of
Hymenophyllum and Trichomanes, as well as the rare
Thyrsopteris elegans, collected by Mr. Thomas Bridges
in Juan Fernandez. But to the Messrs. Veitch &
Sons, of Exeter and Chelsea, among nurserymen,
must be assigned the credit of having introduced
the greatest number of these plants, the collectors
employed by them in Chili and other parts of the
American continent, in India, the Malayan continent
c2
20 FERNS: BRITISH AND FOREIGN.
and islands, and in Japan, having sent home numerous
fine species, while through other sources they have
obtained many additions from Australia, New Zea-
land, and other countries. Messrs. Rollisson, of
Tooting, have likewise succeeded in enriching our
collections with a considerable number, received
principally from Mr. John Henshall, their collector in
Java and the neighbouring islands. Mr. Robert Sim,
of Foot’s Cray, has an extensive collection, which
he increases by importations, and is very successful
in raising plants from spores; and the Messrs. Back-
house & Son, of York, the principal nurserymen
cultivators of exotic Ferns in the provinces, have
introduced a good many fine species of Trichomanes,
and others from Chili and the West Indies. |
—— —j
2 => |
——————F3 — |
= = =
= — ——|
i =e ———
s —— ==
= =e
z= —
es
Genus 109.—Portion of barren and fertile pinna, naturel size. No. Le
AN ENUMERATION OF CULTIVATED FERNS. 209
flexuose, climbing to an indefinite height ; pinnse 1-2 feet long,
adherent. Veins forked; venules combined by a transverse
marginal vein, and in the fertile also near their base by a
transverse, continuous, sporangiferous receptacle, forming a
linear costal sorus. Indusiwm revolute, vaulted, cylindrical,
sporangiferous along its base.
1. S. volubile, J. Sm. in Hook. Gen. Pi. #. 93. Blechnum
volubile, Kawlf.; Kunze, Anal. ¢ 13; Hook. Gard.
Ferns,t.15. Blechnum scandens, Bory, 4 in Dup. Voy.
t. 86.—Tropical America,
Triss IX.—_ASPLENIE A.
Sori oblong or linear, oblique to the midrib or axis of vena-
tion, Furnished with a plane or vaulted lateral indusiwm.
110. ASPLENIUM, Linn.
Vernation fasciculate, erect or decumbent, rarely uniserial
sarmentose. Fronds varying from simple-entire to decompound
and from a few inches to 2-6 feet high, generally smooth
Veins rayed, forked, or pinnate; venules free, sporangiferous cn
the superior side., Sori simple, oblong, or linear, Indusiuin
plane or vaulted.
§ 1. AspLenium verum.—Indusiwm piane,
1. Lancewm group.
Vernation wniserial, sarmentum slender. Fronds distant,
linear-lanceolate, 1-14 foot long. ae siinple, anti or opposite
binate. (Triblemma.)
1. A. lanceum, Thunb. Ie, Plant. Jap. Dec, 11, t. 18. A. subsi-
nuatum, Hook. et Grev. Ic. Fil, t. 27. Diplazium Jan-
ceum, Presi. Scolopendrium dubium, Don,—Inuia,
China, Japan,
2. Serratum group.
Vernation fascieulate, erect. Fronds simple, broad, elliptical,
or lanceolate, 1-2 feet long. (Phyllitis.)
2, A. sorratum, Linn. (Plum. Fil. t. 124); Schk. Fil, t. 64;
Hook. Fil. Exot. t. 70.—Tropical America,
P
210 FERNS: BRITISH AND FOREIGN.
3. A.erenulatum, Presi. A. Nidus, Radd. Fil. Bras. t. 58
(non Linn.). A. Brasiliense, Hort. (non Sw.) ; Lowe’s
Ferns, 5, t.14 B—Tropical America.
“ A. sinuatum, Beawv. Fl. d’Oware, 2, t.79; Hook. Fil. Exot.
t, 16.— West Tropical Africa.
Genus 110.—Fertile pinna, underside. No. 69.
5. A. stipitatum, J. Sm. Neotopteris stipitata, J. Sm. Cat.
Cult, Ferns (1857). Asplenium squamulatum, var. 3
Smithii, Hook. Sp. Fil. p. 83.—East Indies.
AN ENUMERATION OF CULTIVATED FERNS. 211
3. Palmatum group.
Vernation decumbent, subfasciculate. Fronds palmately lobed.
6. A. Hemionitis, Linn.; Hook. Bot. Mag. t. 4911. A. pal-
matum, Lam.; Schk. Fil. t. 66; Lodd. Bot. Cab. 868 ;
Lowe’s Ferns, 5, t. 6, (3 var. cristata, Gard. Chron.
(Jany. 1865), Garden Sport.——South Europe, North
Africa, Madeira.
4, Trichomanes group.
Vernation fasciculate, generally becoming cespitose. Fronds
pinnate, rarely pinnatifid only, linear, 3-12 inches long; pinne
short, often dimidiate, with the costa excentric. (Asplenium
verum.)
a. Fronds pimnatifid.
7. A. alternans, Wall.; Hook. Gard. Ferns, t. 88. A. Dal-
housize, Hook. Ic. Pl. t. 105.—Hast Indies.
b. Fronds pinnate.
8. A. Trichomanes, Linn.; Bolt. Fil. t. 18; Eng. Bot. t.576;
Sowerby’s Ferns, t. 30; Lindl. and Moore’s Brit,
Ferns, t. 39; Lowe’s Ferns, 5, ¢. 22. A. anceps,
Soland. ; Hook. et Grev. Ic. Fil. t. 195. A. melano-
caulon, Willd,
Var. incisum, Moore; Lindl. and Moore’s Brit.
Ferns, t. 89 D, EH; Schk. Fil. t. 74, f.
Var. cristatum, Moore; Lindl. and Moore's Brit.
Ferns, t. 39 H.
Var. depauperatum, Wollast. ; Lindl. and Moore’s
Brit. Ferns, t.39 C.
Var. multifidum, Moore; Lindl. and Moore’s Brit.
Ferns, t. 89 G.
Var. ramosum, Moore; Lindl. and Moore’s Brit.
Ferns, t. 39 F.
Europe, Madeira, South Africa, North India, Australia,
North America.
P2
212 FERNS: BRITISH AND FOREIGN.
9. A. ebeneum, Ait.; Lodd. Bot. Cab. t.5; Lowe's Frvvs, 5,
#.2. A. polypodioides, Sw.; Schk. Fil. t.73.—North
America.
10. A. monantbemum, Linn.; Sm. Ic. ined. t. 73; Lodd.
Bot. Cab. t.1700; Metten. Fil. Hort. Lips. t. 9, f. 7,8;
Lowe’s Ferns, 5, t.1 A—Tropical and Sub-tropical
America, South Africa, Madeira.
11. A. Petrarchee, De Cand.; Hook. et Grev. Ic. Fil. t. 152;
Lowe’s Ferns, t. 5, 38 A, and var. lata, 38 B—South
of Europe.
12. A, formosum, Willd.; Hook. Fil. Exot. t. 16; Lowe’s
Ferns, 5, t.43 B. A. subalatum, Hook. et Arn. Beech
Voy. t. 71.—Tropical America.
13. A. Brasiliense, Radd. Fil. Bras. t. 51,f.1. A. dimidiatum,
Lowe’s Ferns, 5, t. 13 A.—Tropical America.
14, A. tenellum, Roxb. in Beat. St. Helena. A. reclinatum,
Moore et Houlst.; Lowe’s Ferns, 5, t=. 138 B; J. Sm.
Cat. (1857). A. erectum,—var. proliferam, Hook. Fil.
Eeot. t. 72. . alata, Sm.; Hook. et Grev, Ic. Fil. t, 18; Huok. Gen. Fil.
t.7.—West Indies and Tropical America.
2. D. nodosa, Sm.; Schk. Fil. t.152; Hook. et Grev. Ic. Fil.
t.51. Asplenium nodosum, Linn. (Plum. Fil. t. 108).
—West Indies and Tropical America.
270 FERNS: BRITISH AND FOREIGN.
154. KAULFUSSIA, Blume.
Vernation uniserial, contiguous, decumbent; sarmentum
thick, fleshy ; each frond rising from between two short, fleshy,
stipulaform appendages. Fronds broad, pinnately-trifoliate, with
: : o> _ the two lower pinnz sometimes
Z{| bipartite, 1-2 feet high, long,
stipate, pale underneath, and-
furnished with numerous con-
cave dots. Veins costeform,
parallel; venules compound ana-
stomosing, with free veinlets
terminating within the areoles.
Synangia sessile, compital, super-
ficial, distant, orbicular, concave-
hemispherical, each consisting of
10-20 sporangia, laterally con-
: : nate in a circular series, and
Genes iigitly enlarged. No.1...” Opening by a slit on their interior
side.
1. K. esculifolia, Blume; Hook. et Grev. Ic. Fil. ¢. 229;
Hook. Gen. Fil. t. 59 A—Java and Philippine
Islands. !
Oxprr II.—_OPHIOGLOSSACE.
Flowerless plants consisting of straight evolved fronds, which
produce from their disk or base a rachiform, simple or com-
pound paniculate spike, of unilocular, sessile, connate, homo-
geneous cases (sporangia), opening by a vertical slit in two
valves, containing reproducing spores.
155. OPHIOGLOSSUM, Linn.
Fronds erect or pendulous, with dissimilar fertile and sterile
segments, or rarely the whole frond rachiform, Sterile frond or
segments foliaceous, simple, entire, palmately lobed or dicho-
tomously branched. Veins reticulated. Fertile segments spike-
like, simple or rarely forked. Sporangia connate, in two rows,
AN ENUMERATION OF CULTIVATED FERNS. 271
forming a distichous, synangeous spike, each cell opening
horizontally in two equal valves.
Genus 155.—Plant natural size ; fertile spike enlarged. No.3.
Fronds erect, the sterile segments ovate or linear (Terrestrial).
(Euophioglossum.)
1. O. Lusitanicum, Linn.; Hook. et Grev. Ic. Fil. t. 80;
Lindl. and Moore’s Brit. Ferns, t. 51 C; Sowerby’s
Ferns, t. 47. Ophioglossum vulgatum, var. angusti-
folium, Hook. Brit. Ferns, t. 47—South of Europe.
2.0. pedunculosum, Desv.; Kumze, Fil. t, 29, f. 2—
North America.
272 FERNS: BRITISH AND FOREIGN.
3. O. vulgatum, Linn. ; Schh. Fil. t. 153; Hook. Gen. Pil.
4.59 B; Eng. Bot. t. 108; Lindl. and Moore's Brit.
Ferns, t. 51 B; Hook. Brit. Ferns, t. 46; Sowerby’s
Ferns, t. 46.— Temperate Zone of the Northern
Hemisphere.
4, O. reticulatum, Linn. ; Hook. et Grev. Ic. Fil. t.20 (Plum.
Fil. t. 164).—Tropics.
** Fronds pendulous; the sterile segments ribbon-formed,
usually dichotomously branched (Hpiphytal). (Ophioderma.)
5. O. pendulum, Linn.; Hook. et Grev. Ic. Fil. t.19; Hook.
Gard. Ferns, t. 33.—Tropics of Hastern Hemisphere.
6. O.fureatum, J. Sm. Ophioglossum pendulum ; /3 furcatum,
Presi, Tent. Pterid. Supp. p. 56.—Queensland.
*** Fronds palmate, pendulous. (Cheiroglossa.)
7. O. palmatum, Linn. (Plum. Fil. t. 163); Hook. Ic, Pl. t.4.
—West Indies, New Granada, Peru, Brazil, Mauritius.
156. HELMINTHOSTACHYS, Kaulf.
Fronds erect, with dissimilar sterile and fertile segments.
Sterile segments foliaceous, digitate, pedate. Veins forked;
Genus 156.—Portion of sterile pinna, natura! size ; ditto enlarged;
fertile spike, natural size. No. 1.
AN ENUMERATION OF CULTIVATED FERNS. 273
venules free. Fertile segment w rachiform spike. Sporangia
subglobose, sessile, in pedicellate crested whorls (rarely distinct),
‘ forming a clustered simple spike, opening vertically by a slit on
the exterior side.
1. H. Zeylanica, Hook. Gen. Fil. t.47 B; Hook. 2nd Cent.
Ferns, t. 94; Hook. Gard. Ferns, t. 28. Osmunda
Zeylanica, Linn. Helminthostachys dulcis, Kaulf.
—Ceylon, Malay, Molucca, Philippine, and other
islands,
157. BOTRYCHIUM, Linn.
Fronds erect, with dissimilar sterile and fertile segments.
Sterile segments foliaceous, deltoid, bi-tripinnatifidly decom-
Genus 157.—Portion of barren and fertile segmeuts, natural size; fertile
enlarged.
2
274 - FERNS: BRITISH AND FOREIGN.
pound, rarely pinnate. Veins forked; venules free. Fertile
segments rachiform, compound paniculate. Sporangia distinct,
in two unilateral rows, opening vertically in two equal valves.
1. B. simplex, Hitchcock ; Hook. et Grev. Ic. Fil. t. 82.—
North America.
2. B. Lunaria, Sw. ; Schk. Fil. t.154; Hook. Gen. Fil. t.47 A ;
Lindl. and Moore’s Brit. Ferns, t. 51 A; Hook. Brit.
Ferns, t. 48; Sowerby’s Ferns, ¢. 45. Osmunda
Lunaria, Linn. Eng. Bot. t. 318;— rataceum.
Botrychium rutaceum, Sw.; Schk. Fil. t. 155 B—
Temperate Zone of the Northern Hemisphere.
3. B. lunarioides, Sw. Botrypus lunarioides, Miche. ;—
PB obliquum, A, Gray. Botrychium obliquum, Muhi.
B. lunarioides, Schk. Fil. t=. 157;— y dissectum,
A, Gray. Botrychium dissectum, Spr.; Schk. Fil.
#.158.— North America,
4. B. Virginicum, Willd. Osmunda Virginica, Linn. Botry-
chium Virginianum, Sw.; Schk. Fil, t, 156; Hook,
Gard. Ferns, t. 29.—Temperate Zone of the Northern
Hemisphere, Tropical America, East Indies, and
Ceylon,
Orvrz IV—LYCOPODIACE:.
Flowerless moss or fern-like plants, consisting of firm, erect,
creeping or pendulous, simple or branched, often flagelliform
stems, furnished with acerose, rusciform or jungermannia-like,
sessile leaves, which are generally imbricate, and often disti-
chous and of two kinds, bearing in their axis, or on contracted
terminal spikes, 1—3-celled reniform or globose sessile spore-
cases (sporangia), of one or of two forms; one called Antheri-
dangia, containing numerous spores ; the other Oophoridangia,
containing 1-3-8 large spores.
158. PSILOTUM, Sw.
Stems dichotomously forked, compressed or angular, rigid,
erect or slender, pendulous; leaves obsolete or small, bract- .
AN ENUMERATION OF CULTIVATED FERNS. 275
like. Sporangia subglobose, trilobed, vertically trivalved,
solitary in the axis of the bract-like leaves. ,
Genus 158.—Portion of fertile frond, natural size; spore-case
enlarged. No.1.
1. P. triquetrum, Sw.; Schk. Fil. t. 165 6; Hook. Gen. Fil.
t. 87; Lodd. Cab. t. 1916. Lycopodium nudum,
Linn. — Tropics and sub-tropical regions of both
hemispheres.
159. LYCOPODIUM, Linn.
Stems rigid, erect or slender, flagelliform, pendulous or
creeping ; leaves acerose, subulate or rusciform, distant or
imbricate. Sporangia reniform, vertically bivalved, solitary in .
the axis of the leaves, or in terminal, rarely lateral, contracted,
ament-like spikes. Spores numerous, small, uniform (Antheri-
dangia only ?).
TQ
276 FERNS: BRITISH AND FOREIGN.
* Stems creeping. (Epigeous.)
1. L. clavatum, Linn.; Schk. Fil. t.162; Eng. Bot. t. 224.—
Temperate Zone of the Northern Hemisphere, Britain.
2. L. annotinum, Linn.; Schk. Fil. t.160; Eng. Bot. ¢. 239,
— Temperate Zone of the Northern Hemisphere,
Britain.
3. L. inundatum, Linn.; Schk. Fil. t.162; Eng. Bot. t.1727.
—Temperate Zone of the Northern Hemisphere,
Britain.
Genus 159.— Portion of plant, natural size; spore-case enlarged. -
** Stems erect. (Epigeous.)
4. L. Selago, Linn.; Schk. Fil. t. 159; Eng. Bot. t. 233.—
Europe, Britain.
5. L. alpinum, Linn.; Schk. Fil. 1.161; Eng, Bot. ¢.234.—
Temperate Zone of the Northern Hemisphere,
Britain.
6. L.densum, Labil. Nov. Toll. 2, ¢. 251, f. 1—Australia.
AN ENUMERATION OF CULTIVATED FERNS. 277
7. L. fastigiatum, R. Br—New Zealand.
8. L. complanatum, Linn. (Plum. Fil. ¢. 165, f. B); Schk.
Ful. ¢. 163.—Temperate Zone of the Northern Hemi-
sphere.
9. L. dendroideum, Micha. ; Willd. Sp. Pl. 5, p. 21.—North
America.
10. L. cernuum, Linn. (Rheed. Mal. t. 2, ¢. 39); Burm. Fl.
Zey: t. 66 (Plum. Fil. t. 155, f. A).—Tropics; very
general.
*ke Stems pendulous. (Epiphytal.)
1l. L. Phlegmaria, Linn. (Rheed. Mal. 12, ¢.14).—Tropics of
the Eastern Hemisphere. :
12. L. verticilatum, Linn.; Willd. Sp. Pl. 5, p. 48.—
Mauritius.
18. L. ulicifolium, Vent.; Willd. Sp. Pl. 5, p. 27.—India.
14, L. Hookeri, Wall. ; Hook. et Grev. Ic. Fil. t. 165.—India.
15. L. taxifolium, Sw.; Willd. Sp. Pl.5, p,48.—West Indies.
160. SELAGINELLA, Spring.
Fern-like plants. Stems of frondules creeping, sub-ascending
or erect, scandent, of undefined extension (surculose), or rising
Genus 160.—Portion of plant, and the two kinds of spore-cases enlarged.
singly from an underground elongating stolon (cawlescent), or
in a fasciculate manner from a simple erect centre (roswlate) ;
278° FERNS: BRITISH AND FOREIGN.
leaves generally imbricate, distichous, jungermannia-like, of
two sizes; the smaller stipuleform. Sporangia reniform,
vertically bivalved, sessile in the axis of the leaves, or on con-
tracted terminal spikes. Spores of two forms: in some,
sporangia (Antheridangia) small and numerous; in others
(Oophoridangia), 3-4, large.
A. Plant leafy on all sides,
1. 8. spinulosa, Spring. Lycopodium selaginoides, Linn. ;
Schk., Fil. t.165; Eng. Bot. t.1148.—Europe, Britain.
2. 8. uliginosa, Lab. Nov. Holl. 2, p. 104, t. 251, f. 2; Willd.
Sp. Pl. 5, p.32.—Australia and Tasmania.
B. Leaves distichous.
Ҥ 1. Plants surculose.
+ Stems decumbent, creeping.
3. S. Apus, Spring. Monogr. p.75. S. densa, Hort. Lycopo-
dium Brasiliense, Radd. Fil. Bras. 82, ¢. 1—North
America and Brazil.
4, S. Ludoviciana, A. Braun. Revise, No. 5.—Louisiana.
5. §. Helvetica, Link ; Spring. Monogr. 2, p. 88. Lycopodium
Helveticum, Linn.—Europe, Switzerland.
6. S. denticulata, Link; Spring. Monogr. 2, p. 82. Lyco-
podium denticulatum, Linn.—Central Europe.
7. S. delicatissima, A. Brawn. Revise, No.8, 8, microphylla,
Spring. —Columbia.
8.8. serpens, Spring. Monogr. 2, p. 102. Lycopodium
serpens, Desv. §. mutabalis, Hort. S. variabilis,
Hort. S. Jamaicensis, Hort.— Jamaica.
9. S. uncinata, Spring. Monogr.2, p. 109. 8. cesia, Hort—
China.
10. S. hortensis, Metten. Fil. Hort. Lips.p. 128. §. denticu-
lata, Hort.—South of Europe.
11. S. sarmentosa, A. Braun. S. patula, Spring. S. apo-
thecia, Hort.— West Indies.
12, 8. Poppigiana, Spring. ; Hook. Fil. Exot. t.56.—Tropical
erica.
13.
14,
16.
16.
i,
18.
19,
20.
21.
22,
23.
AN ENUMERATION OF CULTIVATED FERNS. 279
8. stenophylla, 4. Braun. Revise, No. 35. 8. micro
phylla, Hort.—Mexico.
tt Stems sub-erect. Y
S. Martensii, Spring. Monogr. 2, p. 129. Lycopodium
stoloniferum, Link. Lycopodium Brasiliense, Hort—
Mexico.
S. Breynii, Spring. Monogr. 2,p. 119. 8S. Panamensis,
Hort. 8. Peppigiana, Hort——Guiana.
8. Galeottii, Spring. Monogr. 2, p. 220. Lycopodium
stoloniferum, Mart, e Gal. S. Sehottii, Hort—
Mexico,
S. sulcata, Spring. Monogr. 2, p. 214. Lycopodium
sulcatum, Desv.—Columbia.
8, atroviridis, Spring. Monogr. 2, p. 124. Lycopodium
atvogidiae, Wall. ; Hook. et Grev. Ic. Fil. t. 839.—East
Indies.
S. inequalifolia, Spring. Monogr. 2,p.148. Lycopodium
ineequalifolium, Hook. et Grev.—Hast Indies and
Java,
8. ciliata, A. Braun. Revise, No.14. Lycopodium ciliatum,
Willd.—Tropical America.
8. Griffithii, Spring. ; Veitch. Cat. (1861).—Borneo.,
ttt Stems scandent.
S. levigata, Spring. Monogr. 2, p. 1387. Lycopodium
levigatum, Willd. Lycopodium Willdenovii, Desv.
Selaginella ceesia, var. arborea, Hort. S&S. altissima,
Klot.—Kast Indies.
§ 2. Plants caulescent.
S. caulescens, Spring. Monogr. 2, p. 158. Lycopodium
caulescens, Wail. var. minor, Veitch. Cat. (1861). —
East Indies.
. §. erythropus, Spring. Monogr. 2, p. 156. Lycopodium
erythropus, Mart, Ic. Sel. Pl. Crypt. t. 20, f. 3.—
Tropical America.
. S. viticulosa, Klot.; Spring. Monogr. 2, p. 186. —
Columbia.
. 8. flabellata, Spring. Monogr. 2, p. 174. Lycopodium
flabellatum, Zinn.—Columbia and Peru.
280 FERNS: BRITISH AND FOREIGN.
27, &. filicina, Spring. Monogr. 2, p.189. §.dichrous, Hort.
—Columbia and Peru.
28, S. Africana, A. Braun. Revise, No. 23. S. Vogelii, Spring.
—Fernando Po.
29. S. pubescens, Spring. Monogr. 2, p.173. Lycopodium
pubescens, Wall. S. Willdenovii, Hort—Hast Indies.
30. S. Lyallii, Spring. Monogr. 2, p. 168. Lycopodium
Lyallii, Hook. et Grev—Madagascar.
31. 8. Lobbii, Hort. ; A. Braun. Revise, App. No. 26; Veitch.
Cat. (1861).—Borneo.
32. 8. Wallichii, Hort.; Veitch. Cat. (1861).—Penang.
§ 3. Plants rosulate.
33. 8. cuspidata, Link; Spring. Monogr. 2, p. 66. S. pal-
lescens, Klot. S. circinalis, Hort—Tropical America.
34. S. convoluta, Spring. Monogr. 2, p. 69. Lycopodium
convolutum, Walk. and Arnott. Lycopodium para-
doxa, Hort.—Tropical America.
35. S. involvens, Spring. Monogr. 2, p. 63. ‘Lycopodium
involvens, Si.—India.
36. S. lepidophylla, Spring. Monogr. 2, p. 27. Lycopodium
lepidophyllum, Hook. et Grev.—Mexico.
Onver V—MARSILEACE ZB,
Plants growing in, or floating in water (rarely not in water),
very varicus in character, consisting of grass or trefoil-like
leaves, or branched with imbricated leaves, bearing sporangia
(conceptacles*) at their roots, or base, or on the footstalks, or on
small branchlets on the under side, which are 1-4, or many-
ceited.
161. MARSILEA, Linn.
Plants with quadrifid (trefoil-like) leaves on along footstalk,
from a rhizome growing under water. Sporangia (conceptacles)
borne on the rhizome at the base of the footstalk, or pedicellate
* A hollow case containing spore-cases; a compound sporangium
AN ENUMERATION OF CULTIVATED FERNS. 281
on the footstalk, 2-valved, containing numerous obovate recep-
tacles in two longitudiual series, bearing cellular vesicles of
Genus 161.—Plant with spore-cates, natural size. No. 1.
two kinds — 1. (Antheridangia), containing numerous small
spores; 2. (Oophoridangia), containing a single large spore.
1. M, quadrifolia, Linn.; Willd. Sp. Pl. 5, p. 588; Schk.
Crypt. t. 173.— Germany.
2. M. macropus, Hook. Ic. Pl. t. 909; Gard. Ferns, t. 63;
Seemann, Journ. Bot. 1, p. 6. (The Nardoo plant of
Australian explorers.)— Australia.
282 FERNS: BRITISH AND FOREIGN.
162. PILULARIA, Linn.
Plants with filiform leaves, from a creeping rhizome under
water. Sporangia (conceptacles) radical (at the base of the leaves
on the rhizome), globose, coriaceous, 2—4-celled, 2-4-valved, each
Genus 162.—Plant, natural size. No.1.
cell containing different kinds of bodies :—1. (Antheridangia),
consisting of vesicles containing many minute granular spores ;
2. (Oophoridangia), each containing a single large spore.
1. P. globulifera, Linn.; Willd. Sp. Pl. 5, p. 535; Bolt.
Fil. t. 40; Schk. Crypt. t.173; Hng. Bot, ¢, 521. —
Europe and Britain.
163. ISOETES, Linn.
Plants with awl-shaped leaves, from a thick creeping rhizome
under water or out of water. Sporangia (conceptacles) borne in the
AN ENUMERATION OF CULTIVATED FERNS. 283
axis of the leaves at their base, globose, 1-celled, traversed by
thread-like receptacles of two kinds: —1. (Antheridangia),
containing numerous small spores; 2. (Oophoridangia), con-
atining large 4-sided spores.
Genus 163.—Plant, uatural size. No.1.
1. L. lacustris, Linn.; Willd. Sp. Pl. 5, p. 584; Bolt, Fil,
t.41; Schk. Crypt. t.173; Eng. Bot. 1084.— Europe,
Britain,
284
ADDENDA.
—+eee
A CONSIDERABLE time having elapsed since the pre-
ceding Enumeration was prepared and sent to the
press, has enabled me to note a number of recently
introduced, and a few omitted older species, which I
now enumerate in the form of addenda.
N.B.—Those marked thus + after the name of the country
are entered on the authority of Mr. T. Moore’s notices of them,
in the Proceedings of the Royal Horticultural Society and the
Gardeners’ Chronicle newspaper ; not having myself seen them
alive or obtained specimens.
38. LOMARIOPSIS.
2*, L. fraxinea, J. Sm. Lomana fraxinea, Willd. Acro-
stichum (Lomariopsis) sorbifolium, Hook. Sp. Fil. 5,
p. 241 (non J. Sm.).—Mauritius.
Oxss.—This is a very distinct plant from the West Indian
type of L. sorbifolia.
50. GYMNOGRAMMA.
§ 7. Ampelogramma, J. Sm. Vernation sarmentose, according
to Hook. ; fronds indefinite ; rachis flexuose ; pinne bi-tripinnate,
refracted ; pinnules small, cuneiform.
18. G. flexuosa, Desv.; Hook. Sp. Fil. 5, p. 192. G. retro-
fracta, Hook. et Grev. Bot. Misc. 8, ¢, 112.—Tropical
America,
AN ENUMERATION OF CULTIVATED FERNS. 285
62, DICTYOPTERIS.
2. D. macrodonta, Presl, Tein. Pterid.; J. Sin. Gen. Fil.
Philipp. ; Hook. Journ. Bot. 3, p. 396. Polypodium
macrodon, Reinw. in Herb. J. Sm. Aspidium difforme,
Blume, accord. to Reinw. in Herb. J. Sm. Polypo-
dium confluens, Wall.—East Indies, Malayan Archi-
pelago, Fiji.
Oss.—In Herbariums and books there is great confusion
in the synonymy of what, according to Garden plants, seem
to be two distinct species ; — viz., Dictyopteris wregularis and
D. macrodon of Presl; judging from Herbarium specimens, they
are difficult to be recognized as distinct ; but living plants show
the first to have erect vernation, the other decumbent.
The Kew Collection is indebted for this species, as also the
beautiful Mécrolepia platyphylla, and others, to Mr. Robert
Kennedy, Florist and Fern-dealer in Covent Garden.
3. D. Cameroonianis, J. Sm. Polypodium (Dictyopteris)
Cameroonianum, Hook. Sp. Fil. 5, p.104. Dictyopteris
varians, Moore, in Gard. Chron. (1864).— Tropical
West Africa.
Oxzs.—A plant of this Fern was sent some time ago from Old
Calabar to the Royal Botanic Gardengdiginburgh, and I am
indebted for a specimen of it to Mr. James McNab, the
Curator of that truly scientific garden.
63. MENISCIUM.
3*, M. angustifolium, Willd.; Hook. Sp. Fil. 5, p. 164.—
Tropical America,
65. NEPHRODIUM.
4*, N. cyatheoides, Kaulf.; Hook. Sp. Fil. 4, 4.241. Poly-
stichum Dubreuillianum, Gaud. in Freyc. Voy. Bot.
Crypt. t. 9 —Sandwich Islands.
Ozs.—It is but right to observe that this remarkable species,
as also the Sandwich Island Ferns in this addenda, with the
previously entered Phegopteris wridentata, and the remarkable
286 FERNS : BRITISH AND, FOREIGN.
Colysis Spectrum, also the beautiful Cibotium Menziesit, were
sent to the Royal Gardens, Kew, from the Sandwich Islands by
Dr. Hillebrand, in 1863.
68. CYRTOMIUM.
1*, C. Fortunei, J. Sm. Fronds 1-14 foot long, pinnate ;
pinne lanceolate, faleate, acuminate, 2-3 inches long,
1 inch wide, entire, the base oblique, the inferior
rounded, the superior truncate and subauriculate.
Costa ebenous, upper surface dull, unreflecting. Sort
numerous.—Japan.
Oxs.—In Herbaria, specimens of this cannot readily be dis-
tinguished from C. falcatum ; but on seeing living plants stand-
ing side by side, the difference is evident ; C. Fortunei being a
smaller-growing plant, thinner in texture, and not lucid and
reflecting light, as C. falcatum.
70. ASPIDIUM.
3*, A. polymorphum, Wall. ; Hook. Sp. Fil. 4, p. 54 (ewelud,
syn.).—India, Ceylon.
3**, A. Barteri, J. Sm. Vernation erect. Fronds pinnate,
1-2 feet high; pinnz linear-lanceolate, 6-7 inches
long, 1-13 broad; the lower pair sometimes binate,
or auriculated on the lower margin. Sori regularly
biserial between the primary veins. Indusiwm small,
fugaceous. Aspidium polymorphum, Wall. ac-
cording to Hook. Sp. Fil. 4, p. 54.—West Africa,
Fernando Po.
Oxzs.—This is quite distinct from the preceding species.
In A. Barteri the sort are perfectly serial, whereas in A. poly-
morphum the sort are scattered.
‘
74, POLYSTICHUM.
6*. P. lepidocaulon, J.8m. Aspidium lepidocaulon, Hook.
Sp. Fil. 4, t. 217—Japan.
AN ENUMERATION OF CULTIVATED FERNS. 287
Oxzs.—On receiving this plant, it accorded so well with the
Indian specimen of P. obliguum, which led to that name being
inserted at p. 149, but the plant grew out of its imported form ;
and if the Indian plant (which has not yet been introduced)
should assume the character of this by cultivation, then this
name must become a synonym.
The introduction of this species is due to Mr. Richard Oldham,
the last special collector of plants sent out from Kew, who,
after remaining three years in Japan, visited Formosa, where
his health failed. He returned to Amoy, and there died in
November, 1864.
10*. P. coneavum, Moore, Proc. Hort. Soc. 11, 377. Lastrea
Standishii, Hort.—Japan.t
10**, P. ordinatum, Fée ; Moore, Proc. Hort. Soc. 11, 367.—
Tropical America.+
75, LASTREA.
29*, L. spectabilis, J. Sm. in Enum. Fil. Philipp. Hook. Journ.
Bot. 8 (1841). Aspidium spectabile, Blume.—Philip-
pine and Malayan Islands, India.
41*, L. sparsa, Moore. Nephrodium sparsum, Don. As-
pidium purpurascens, Blume, according to Hook. Sp.
Fil. 8, p. 188, t. 262.—Throughout India, Java, and
Ceylon.
41**, L. latifrons, J. Sm. Nephrodium (Lastrea) Jatifrons,
Hook. Sp. Fil. 4, p. 138.—Sandwich Islands.
41***, I, membranifolia, Presl, Pterid. Nephrodium mem-
branifolium, Presl, Relig. Haenk. t. 6, f.3; Hook. Sp.
Fil. 4, ¢, 26.—India, Ceylon.
79. NEPHROLEPIS.
3*, N. falciformis, J. Sm. Fronds suberect, linear, pinnate,
14-2 feet in length; pinne: numerous; the sterile
(lower ones) elliptical, obtuse, base truncate; the
superior ones fertile, lanceolate, falcate, acute, sub-
deflexed, 13 inch long by 3 inch wide; base truncate,
288 FERNS : BRITISH AND FOREIGN.
subauriculate on the upper margin. ‘Sort uniserial,
antimarginal. Indusiwm suborbicular—Borneo.
Oxs.—The affinity of this species is with N. tuberosa 3 but
seeing the two plants together, the difference is readily seen.
79a. ISOLOMA, J. Sm.
Vernation fasciculate, erect, stoloniferous. Fronds linear,
1-2 feet long, pinnate; pinnew oblong, elliptical or lanceolate,
falcate, coriaceous, the base truncate and subauriculated;
petiole short, articulated with the rachis. Veins forked; sterile
Genus 79a.—Portion of sterile frond and a fertile pinna, natura »..
itto enlarged. No. 1.
venules free, the fertile transversely combined at the margin,
forming a continuous receptacle. Indusiwm linear, interiorly
attached, plane, equal with and conniving with the margin,
forming with it a vertical exteriorly open groove, containing
the sporangia.
1. I, lanuginosa, J. Sm. in Lond. Journ. Bot. 1, p. 420.
Lindsea lanuginosa, Wall.; Hook. Sp. Fil. 1, t. 69 B.
—Singapore, Penang, New Guinea, and Seychelles
Islands,
AN ENUMERATION OF CULTIVATED FERNS. 289
Oss.—This remarkable Fern has lately been added to the Kew
collection, having been sent from the Mauritius Botanic Garden
by Mr. John Horn.* In general habit and appearance this Fern
is the prototype of Nephrolepis; but is distinguished by the
receptacles being laterally confluent, forming a linear marginal
sorus, similar to that of Lindscea, from which it is, however,
quite distinct in habit.
81, PHEGOPTERIS.
2*, P. plumosa, J. Sm. Asplenium Felix-foomina, var. plu-
mosum, Moore, Nat. Print. Ferns, oct. ed. p. 56;
Lowe’s New Ferns, t.14,—Yorkshire (Mr. Stansfield).
Ons.—A few years ago three plants of this Fern were found
. wild in Yorkshire. It was soon alterwards described and figured
in the works above quoted, under the name of Asplenium Felix-
fomina,var.plumosum ; but upon what grounds it was referred tc
Asplenie I cannot explain, as all the specimens I have examined
of ithave small, punctiform, naked sori, perfectly characteristic
of the genus Phegopteris, with which it also agrees in habit.
This leaves me no other alternative than to consider it a species
of that genus, and consequently a new British species. In
doing so, the question arises.as to whether it represents an
ancient species not before noticed, or the modern result arising
from the power of nature to generate new forms, in accordance
with the Darwinian Theory of creation of species. It is, how-
ever, to be observed that in abnormal or difformed states of
Aspleniwm and Scolopendrium, the sori are depauperated, in
some instances having no vestige of an indusium; but such is
not the case with this plant. The fronds are perfect in every
respect, and if Herbarium specimens had been received from
some furcisu country, no Pteridologist, on seeing the naked sori,
would retur it to Asplenive.
93, ADIANTUM,
35*. A. Gheisbeghtii, Backhouse, Cat. (P) A. tenerum, var.
(J. Sm.)—Tropical America.t+
* Who left Kew in 1861 to be assis aut to Mr. Duncan, See page 16
U
290 FERNS : BRITISH AND FOREIGN.
39%. A. colpodes, Moore, Gard. Chron. (1865).—Ecuador.t
44, A. tinctum, Moore, Proc. Hort. Soc. 11, 369.—Tropical
America.f
96. PTERIS.
20%. P. straminea, Metten. P. crispa, Hort. (non Linn.).—
Chili.
97. LITOBROCHIA.
14*, L. areolata, Moore. Pteris areolata, Lowe’s New Ferns
t. 57.—India.t
102. LOMARIA.
2%, L. rigida, J. Sm. Vernation fasciculate, erect, becoming
ceespitose. Fronds (the sterile) lanceolate, 8-10 inches
long, 13-2 inches broad, erect, rigid, pinnatifid to the
rachis; sinus acute; segments alternate, contiguous,
lanceolate, falcate, finely serrulate; the lower ones
decreasing in size and obtuse. Veins evident. Fertile
fronds as broad and rigid as the sterile; segments be-
coming involute, densely sporangiferous.— Chatham
Islands.
12%, L. Germanii, Hook. Sp. Fil. 8, 1. 152. LL. crenulata,
Hort.—Chili,
110. ASPLENIUM.
33%. A. Kaulfussii, Schlecht, Adwm. (29 in obs.). A. protensum,
Kaulf. (non Schrad.).—Sandwich Islands.
19*. A. tenerum, Forst.; Schk. Fil. t. 69.—Islands of the
‘Pacific, Ceylon.
92", A. nigripes, DMetten. ; Hook. Sp. Fil. 8, p. 222.—Ceylon.
AN ENUMERATION OF CULIIVATED FERNS. 291
127. DICKSONTIA.
6. D. Youngii, Moore, Proc. Hort. Soc.—New South Wales.+
128. CIBOTIUM.
4, C. regale, Linden; Moore, Gard. Chron. (1864), 414. ~
Mexico.+
CYATHEA.
5*, C. princeps, J. Sm. Cibotium princeps, Linden, Cat.—
Mexico.
Ozs.—Plants of this Fern were sent to this country under the
name of Cibotiwm princeps ; but a plant in Messrs. Lee’s nursery
having produced fructification during 1865 proves it to be a
Cyathea.
Abstract of the number of species, and date of intro-
duction :—
Exotic species at Kew in 1822... 14. eee oer 40
Do. My Enumeration of Kew Ferns, 1846 ... 355
Do. My Catalogue of Cultivated Ferns, 1857... 559
This Enumeration, Exotic and British ... 1028
Do. Fern Allies, Exotic and
British ... 1. .. 56
To.lin 1865... .., ... 1084
In closing this enumeration, I deem it necessary
to state that a few species recently introduced to
the Kew collection have failed to become esta-
blished, after their names were entered on the list;
the principal uf which are: Dicranoglossum furcatum,
Aconiopteris nervosa, Pleocnemia Leuziana, Adian-
tum lunulatum, Lomaria Fraseri, Asplenium lanceum,
A. radiatum, Antigramma repanda, Polystichwm
anomalum, Loxsoma Ounninghamit, Kaulfussia escu-
lefolia. Also the following, entered from my cata-
u 2
292 FERNS : BRITISH AND FOREIGN.
logue of 1857, were not inthe Kew collection in 1864,
viz., Gymnogramma rutcefolia, Hemionites pedata,
Notholena lanuginosa, Myriopteris vestita, Onychium
auratum, Blechnum triangulare, Nephrolepis undulata,
Lindsea guianensis, Schizoloma ensifolia, Thyrsopteris
elegans, Botrychium virginicum.
Several special causes which lead ‘to the loss of
species have already been noticed. I did not intend
to advert to them again; but the recent death of
Sir W. J. Hooker * necessitates me to repeat what I
have stated at pages 42 and 43, where, in speaking
of my having resigned the charge of the Kew collec-
tion, I said that “happily it remains under the
direction of Sir W. J. Hooker.’ These words will
now apply to Dr. Hooker, the present Director, whose
name is sufficient in itself to sustain the scientific
reputation of Kew. Let us hope that the general
collection of living plants of the Botanic Garden, now
famous for more than a century,t will not suffer by
the modern taste for showy flowers, and what is
now fashionably called “ foliage plants.”
* Died August 12th, 1865.
+ Number of species at Kew in 1768 .. .. 8,400
Ditto 1786 .. .. 5,500
Ditto 1813 .. 9,800
Since the latter date no general catalogue has been pub-
lished, and no public record kept of the plants introduced
or lost.
293
ETYMOLOGY.
—— ot
ANY cultivators and plant amateurs, not versed
in scientific literature, complain of the (to
them) difficulty to pronounce names given. to plants
by botanists. This complaint is, however, more
imaginary than real. They have only to recollect
that time and use has made them perfectly familiar
with such names as Pelargonium, Geranium, Hydran-
gea, Calceolaria, Rhododendron,Chrysanthemum, Mesem-
bryanthemum, Elscholizia, Fuchsia, and many others
now fluently spoken as if they were original words of
their mother-tongue. Others say, these long-sounding
names would be more readily reconciled to them. if
the meaning and bearing upon the plant were ex-
plained. To meet this desire, I have drawn up the
following table, showing the derivation of the genera
of Ferns characterized in the preceding pages.
As a general rule, it may be admitted that names
of plants are derived from three principal sources.
The first, and greater number, being generally a com-
pound of two Greek words. The second, names of
persons, with the addition of a Latin termination. The
third source is various; such as the adoption of local
aboriginal names,names from Heathen Mythology, and
294 FERNS: BRITISH AND FOREIGN.
often words of doubtful origin, and unmeaning appli-
cation to the plants, have been given as generic
names by even the most learned botanists.
N.B.—For all names ending and beginning with
Pteris, see Pteris.
Aconiopteris, akon, a point, and pteris; the veins forming sharp
angles close to the margin of the frond in the typical
species.
Acrostichum, akros, highest, stichos, order ; the fructification at
the top of the fronds.
Adiantum, adiantos, dry; when plunged in water comes out
dry.
Alsophila, alsos, grove, phileo, to love; grows in groves and
shady places.
Amphidesmium, amphi, around, or on both sides, desmos, a
band; relating to the position of the sori to one another.
Anapeltis, ano, without, peltis, shield; the sori naked, in oppo-
sition to Pleopeltis.
Anchistea, agnati, kindred, intermediate between Woodwardia
and Doodia.
Anemia, aneimon, naked; naked spikes of fructification.
Anemidictyon, veins reticulated ; not free as in Anemia.
Anetium, probably from the Greek anaitios, guiltless. This is
perhaps meant to indicate that A. citrifolium had been put
wrongfully to Acrostichum.
Angiopteris, angio, open, pteris; the open sporangia.
Antigramma, anti, opposite, gramma, a line; the linear sori
being in pairs opposite to each other.
Autrophyum, antrum, a den or hollow; the receptacles of
sporangia being in a grove or channel.
Arthropteris, arthron, joint, pteris; the frond and pinnss being
articulate.
Aspidium, aspidos; the indusium being like a buckler or
shield.
Asplenium, a, privative, splen, the spleen; medical qualities.
Balantium, balantion, a purse or bag; the form of the indu-
sium.
Blechnum, one of the Greek names for a fern.
Botrychium, botrys, bunch or raceme; the fructification like a
bunch of grapes.
ETYMOLOGY. 295
Brainea, in honour of C. Braine, Esq., who introduced the first
plant.
Callipteris, kallos, beautiful, and pteris, beautiful fern.
Campyloneurum, kamptos, arched, newron, a nerve; the veins
forming arches.
Ceratopteris, keras, keratos, a horn, pieris; horned fern. ,
Ceterach, a name given by Persian physicians.
Cheilanthes, chedlos, lip or margin, anthos, flower; the fructifi-
cation on the margin. .
Cibotium, kibotos, a casket or coffer; the form of the
indusium.
Cincinalis, meaning unknown.
Colysis, kolysis, separation, a species separated from other genera.
Coniogramma, konis, dust, gramma, a line; imperfect sporangia,
but not applicable in perfect specimens.
Cryptogramma, kiyptos, hidden, gramma, a line or writing ; the
lines of fructitication being hidden.
Cyathea, kyathos, a cup; the form of indusium.
Cyclodium, kyklos, circle; the form of indusium.
‘Cyclopeltis, kyklos, circular, peléis, shield; the form of indu-
sium,
‘Cystopteris, kystos, a bladder; the inflated indusium,
Danza, in. honour of Pierre Martin Dana, a Piedmont
botanist.
Davallia, in honour.of Edmond Davall, a Swiss botanist.
Dicksonia, in honour of James Dickson, a British crypto-
gamist.
Dicranoglossum, dikranos, forked, glossa, tongue; the fronds
forked and pendulous.
. Dictymia, diktyon, a net; the netted venation.
' Dictyogramma, diktyon, gramma, a line; the sori reticulated.
Dictyopteris, diktyon, pteris; the veins reticulated.
Dictyoxiphium, diktyon, wiphion, sword; the veins reticulated,
and the form of the frond.
Didymochlena, didynos, two or double, chlena, a cloak; the
indusium being double.
Diplazium, diplazo, to be double; two indusia on the same
receptacle.
Doodia, in honour of Samuel Doody, a London apothecary, and
_ British cryptogamist.
296 FERNS: BRITISH AND FOREIGN.
Doryopteris, dory, spear or halbert, pteris ; form of the fronds.
Drymoglossum, drymos, wood, glossa, tongue; the fronds like
tongues on trees.
Drynaria, dryads; the sterile fronds being like oak leaves,
which tree was sacred to the Dryads.
Dryomenis, dryos, dryads (as above), meniskos, a crescent ; shape
of the sori.
Egenolfia, in honour of Christian Egenolph, a German author
of a book on herbs. ; ;
Elaphoglossum, elaphos, a stag, glossa, tongue; the fronds
being like the stag’s tongue.
Eupodium, eu, good, pous, podos, a foot; the sorus having a
foot-stalk.
Fadyenia, in honour of Dr. M‘Fadyen, an eminent physician
in Jamaica.
Feea, in honour of Mons. A. L. A. Fée, Professor of Botany at
Strasburg, a celebrated writer on ferns.
Gleichenia, in honour of Baron P. F.von Gleichen, a German
botanist.
Goniophlebium, gonia, angle, phlebes, veins; the veins meeting,
forming angles.
Goniopteris, gonia, angle, pteris ; the veins meeting, forming
angles,
Grammitis, gramma, a line; the sori being linear.
Gymnogramma, gymnos, naked, gramma, Bite the sori on the
veins in lines, and naked.
Gymnopteris, gymnos, naked, pteris; the fructification naked.
Haplopteris, haploos, simple, pteris ; simple venation.
Helminthosiachys, helminthos, worm, stachys, a spike; the
fructification in compact spikes, worm-like.
Hemidictyum, hemi, half, diktyon, a net ; the outer portion only
‘of the veins being reticulated.
Hemionites, a name given by ancient botanists to a plant now
called Aspleniwm hemionites, but retained as a generic name
fora West Indian fern by Linnzeus.
Humata, humatus, humid; in opposition to Adiantum.
Hymenodium, hymen, membrane; the character of the frond.
Hymenolepis, hymen, membrane, lepis, a scale; membraneous
scales covering the fructification.
ETYMOLOGY. : 297
Hymenophyllum, hymen, membrane, phyllon, leaf; “ Filmy-
leaf Fern.”
Hymenostachys, hymen, membrane, stachys, a spike; the firuc-
tification on spikes.
Hypoderris, hypo, under, derma, skin; the indusium under the
sori, like a thin skin.
Typolepis, hypo, under, lepis, a scale; the sporangia under
the indusium.
Isoetes, isos, equal, etes, the year; remaining the same through-
out the year.
Isoloma, isos, equal, loma, a border or margin; the indusium
and margin equal.
Kaulfussia, in honour of D. G. F. Kaulfuss, of Halle, a celee
brated writer on ferns.
Lastrea, in compliment to Chev. de Lastre, a French nobleman.
Lepicistis, lepis, scale, kistis, a cyst or cell; the sori being im-
mersed in scales, which form a cyst.
Leptochilus, leptos, slender, cheilos, lip ; narrow indusium.
Leptogramma, leptos, slender, gramma, a line; short linear sori.
Leucostegia, leukos, white, stegos,a cover; the indusium being
pale-coloured, approaching white.
Lindsza, in honour of Dr. Lindsay, of Jamaica, a writer on
the germination of mosses and ferns.
Lithobrochia, lithos, a stone, brocha, spots; the areoles of the
reticulated veins like pavement.
Llavea, named by Lagasca after a Mexican injcanen
Lomaria, loma, a fringe or border; relating to the indusium,
Lomariopsis, lomaria, opsis, like; like Lomaria,
Lonchitis, logche, a lance; form of pinnz.
Lopholepis, lophos, a crest, lepis, scale ; the sori being furnished
with a tuft of slender scales.
Lophosoria, lophos, crest or tuft; sori farnished with hairs like
a tuft.
Lorinseria, in honour of Gustave Lorinser, a Bohemian
physician.
Loxsoma, loxos, oblique, soma, band; the oblique broad ring of
the sporangium.
Lycopodium, lykos, wolf, pous ; foot-resemblance,
Lygodium, lygodes, flexible ; climbing plant.
298 FERNS: BRITISO AND. FOREIGN.
Marattia, in honour of J. F. Maratti, of Tuscany, who wrote
on ferns.
Marsilea, in honour of Count L. F. Marsigli, of Bologna.
Meniscium, meniskos, crescent ; shape of the fructification.
Mesochleena, mesos, middle, chlena, a cloak ; attachment of the
indnsium.
Microlepia, mikos, small, lepis, scale; the indusium small.
Microsorum, mikros, small; small sori.
Microstaphyla, mikros, small, staphyle, a bunch; the fertile
frond being like a little bunch.
Mohria, in honour of D. D. Mohr, a German botanist.
Myriopteris, myrios, myriad, pteris; the frond being divided
into a number of small parts,
Neottopteris, nevitia, nest, pteris; bird’s-nest fern.
Nephrodium, nephros, kidney ; form of the indusium.
Nephrolepis, xephros, kidney, lepis, scale; the indusium being
kidney-shaped and scale-like.
Neurocallis, newon, a nerve, kallos, beautiful; the venation.
Neurodium, neuron, nerve; the venation.
Niphobolus, niphes, of snow, bolos, a large pill; the under side of
the frond densely covered with white scales, snow-like and
the round sori.
Niphopsis, niphos, of snow, opsis, like; like niphobolus.
Notholana, nothos, spurious, chlena, cloak; the imperfect
indusium.
Ochropteris, ochros, pale; pale fern.
Odontosoria, odontos, of a tooth, sori; the sori-like teeth.
Oleandra, resemblance to Nerium Oleander; the Oleander.
Olfersia, in honour of Professor Olfers, a celebrated astro-
nomer.
Onoclea, onos, a vessel, kleio, to enclose; the rolled-up
fertile segments of the frond.
Onychium, onychion,.a little nail; resemblance to the fertile
segments of the fronds.
Ophioglossom, ophios, of a serpent, glossa, tengues ; the spikes
of fructification.
Osmunda, Osmunder, one of the names of Thor, a Celtic
divinity.
Paragramma, para, near to, gramma, a line; the sori in short
lines parallel with and close to the margin.
Pellaa, pellos, dark-coloured; the dusky colour of the fronds.
Phegopteris, phegos, beech ; beech fern, by some called sun fern.
ETYMOLOGY. 299
Phlebodium, phiebes, veins, odous, a tooth; the joining of the
veins in the areoles being like teeth.
Phymatodes, plymata, tubercles ; the impressed sori having
-the appearance of tubercles on the upper side of the
frond.
Pilularia, pilula, a pill; the form of the spore-cases.
Platycerium, platys, broad, keras, horn; the fronds divided in
broad segments like stags’ hors. (The stags’-horn
fern.)
Platyloma, platys, broad, loma, border ; oe broad sori close
to the margin.
Pleocnemia, pleos, full, knemia, rays; full of rays; the vena-
tion.
Pleopeltis, pleos full, peltis, shield; the sori being furnished with
numerous round scales.
Pleuridium, pleura, side (ribs); the primary veins being coste-
form, like ribs.
Peecilopteris, pelilos, spotted, pteris; the appearance of the
venation.
Polybotrya, polys, many, botrys, bunch ; hs fructification being
in bunch-like racemes.
Polypodium, polys, many, pous, foot; polypus; the rhizome,
' when destitute of the fronds, having the appearance of
some kind of sea-polypus.
Polystichum, olys, many, stichos, order; not specially
applicable to the genus Polystichum, as now defined.
Psilotum, psilos, naked; destitute of leaves.
Psomiocarpa, psomion, a small pellet, karpa, fruit; the
sporangia being in small round patches, in spikes.
Pteris, pteryx, wing; the scientific name given to the plant
known by the names of Brake, Bracken, and Fern;
on account of the supposed likeness of the branching
of its fronds to wings. This, being the commonest
ot all Ferns, has become the type “of the whole race;
hence Pteris means Fern, all generic names of ferns
ending in pteris; such as Ceratopteris means horned
Fern, Diotyopteris netted Fern; also, if at the beginning
of a word, as Pteridography, a book or writing on Ferns ;
Pieridologist, a studier and writer on Ferns; Péeridophilist,
a lover of Ferns.
Rhipidopteris, rhipis, fan; shape of the fronds like a fan.
300 FERNS: BRITISH AND FOREIGN.
Saccoloma, sakkos, a bag or sack, loma, margin; the union of
the indusium with the margin, forming a hood or cyst.
Salpichlena, salpi, pipe, chlena, a cloak; the indusium being
like a pipe.
Schellolepis, schello, skeleton, lepis, scale; the character of the
scale surrounding the sori.
Schizoloma, schizo, I cut, loma, a border or margin; the sori in
the form of a slit on the margin of the frond.
Scolopendrium, skolopendra, a centipede; the appearance of
fructification on the under side of the frond.
Selaginella, the diminutive of selago (Lycopodiwm Selago).
Selliguea, in memory of M. Selligue, a French optician. _
Sitibolium,* sitos, food, bolos, a large pill; the appearance of
fructification.
Soromanes, soros, heap, mania, fancy; the sporangia in fanciful
and irregular heaps, ;
Stenochlena, stenos, narrow, chlena, cloak; narrow indusium.
Stenosemia, stenos, narrow, sema, standard; the fertile frond
with its narrow segments being like a flag.
Struthiopteris, struthios, ostrich ; the fronds being like the
feathers of an ostrich.
Thyrsopteris, thyrsos, bunch or raceme, pteris ; the fructification
in racemose bunches.
Todea, in honour of Henry Julius Tode, of Mecklenburg, an
experienced mycologist.
-Trichocarpa, thria, trichos, a hair, karpa, fruit ; the fructification
borne on a hair-like stalk.
Trichomanes, thriz, hair, manos, soft; the delicate nature of the
fronds, :
Trichopteris, thrix, hair, pteris; the sori being furnished with
hairs.
Vittaria, vitta, riband; shape of narrow fronds.
Woodsia, in honour of Joseph Woods, a celebrated British
botanist (died 1864),
Woodwardia, in honour of Thomas Jenkinson Woodward, an
English botanist.
Xiphopteris, wiphos, sword, pteris ; form of the frond.
* For Sitilobium, at page 236 and elsewhere in this work, read Sitidolium.
301
CULTIVATION,
—1o+—
1. Preztiminary Remarks.
AVING given an account of how the Exotic
Ferns of the preceding enumeration have been
introduced to this country, I shall now proceed to state
the best means for growing and preserving them in
our collections; and as the successful eultivation of
Ferns depends much upon a knowledge of the con-
ditions under which they grow in their native country,
I begin with a few observations on that point.
"Ferns have already been spoken of as favourites
with the plant-loving public; but it is not simply on
heir merits as pretty and interesting objects that they
claim attention. Geology reveals to us that Coal—
that source of our domestic comforts and national
greatness—is formed chiefly of Ferns, which at some
remote period grew upon the earth. However, this is
not the place to discuss the views and speculative
theories regarding the thermal and gaseous condition
of the earth and atmosphere, under which Ferns then
flourished, or to reason on the manner by which they
were converted into coal. I may simply remark that
fossil remains show, that contemporaneous with
Ferns grew plants of remarkable character, quite
302 FERNS: BRITISH AND FOREIGN.
distinct from those of the present era. Not so the
Ferns, for their beautiful and well-preserved remains
show that their fronds were of various sizes and forms,
in every way analogous to the present race; having
free and anastomosing venation, round and linear
fructification, and, in some instances, almost identical
with species now living. At present, Ferns rank
amongst the widest spread of all the orders of the
vegetable kingdom, being found in more or less
number in all climates, between the most northern and
southern limits of vegetable life, and at elevations
ranging from the sea-level to 14-15,000 feet within the
tropics, their number in any localities being generally
in proportion to the degree of atmospheric moisture in
conjunction with elevation, the latter applying specially
to the interior of continents. Comparatively few
spczies are found in open, grassy, thinly-wooded
countries, whether it be the plain or mountain-slope ;
such districts are often in full possession of the most
gregarious and abundant of all Ferns, the common
Brake (Pteris aquilina), which, under slightly different
forms, and in some countries accompanied by different
species of Gileichenia, occupy vast tracts of the earth’s
surface. In hot and moist plains, in valleys of great
extent, the number of different species are few; even
in the valley of the Amazon, teeming as it does with
vegetable life, the number of Ferns found by Dr.
Spruce after he left the coast Flora, at Para, in his
journey of 2,000 miles, were very few. They became
moré numerous on attaining an elevation of 1,500
feet, and in one locality, at a higher elevation, he
found 250 species in a diameter of fifty miles.
Another extensive tract with but few Ferns is the
CULTIVATION. 303
dry zone of Northern Africa, and few have as yet
been recorded from the interior of Australia; while,
on the contrary, on elevated coast-ridges and islands,
they form a large proportion of the entire Flora.
They are the most numerous, both in regard to genera
and species, in the tropical regions, where, too, a
greater number of individual plants are to be found
than in temperate regions.*
On reviewing the above, it may readily be supposed
that the varied influences under which Exotic Ferns
naturally grow, necessitate various kinds of treatment
for cultivating and preserving them in this country ;
but such is not the case. As might be expected,
our native representatives (although only forty-two
in number) have received special attention. We have
Fern Tourists in plenty; almost every spot of our
country, where are conditions congenial to their
development, has become more or less familiar to
those who take an interest in their study—in the
damp shady glen, the wildest wood, banks of streams,
mountain-sides, whether it be in their sheltered easy
slopes, or in the almost inaccessible craggy steeps.
Indeed, so diligently have the Fern explorings been
carried on, that numerous curious and beautiful
varieties, highly prized in our gardens, have been
found. But, however laudable and agreeable Fern-
growing may be, yet it is to be regretted that it leads
to the extinction of some of our rarest native species.
Even the more common are becoming scarce in locali-
ties within easy reach; great quantities being yearly
* Want of space prevents me entering more fully into the
history of geographical distribution.
a0 FERNS: BRITISH AND FOREIGN.
consigned to the London markets. Since Ferns have
become so popular, those who patronize horticulture,
from the members of the upper classes who construct
large hothouses, to the humbler patron, or the lard-
working mechanic who prides himself on his posses-
sion of a Wardian case, are desirous to obtain such
information as will enable them to cultivate them
successfully, either in the Conservatory, Wardian
case, or Rockwork in the open air. To supply this,
several books on the subject have been published.
Indeed, if an example- may be taken, besides the
extraordinary degree of familiarity so rapidly attained
in the knowledge of Ferns by all classes, the constant
increase of collections gives sufficient evidence.
Nevertheless, in publications which, through their
cheapness, are within the reach of every Fern lover,
frequent complaints are met with, tending to show
that the result is not always satisfactory to the culti-
vator, especially among amateurs and persons of
small means. These complaints are mainly attri-
butable to the author not familiarizing his readers
sufficiently with the habits of Ferns, the conditions
under which they luxuriate in their native localities,
and the necessity of imitating those conditions as far
as practicable. It is, however, gratifying to observe
that, in the progress of horticulture, the knowledge of
the natural conditions of plants is more and more
sought afte, and appreciated with very good results,
forming a lively contrast with the old times, when the
Trichomanes radicans could only with difficulty be
kept alive even by the most eminent horticulturists ;
a fact observed by the originator and promoter of the
Wardian case system, under which this shade and
CULTIVATION. 305
moisture-loving Fern stands pre-eminent,-even sur-
passing in size and luxuriance those in their native
retreats. A familiar example of opposite nature to
the preceding is our native species Asplenium septen-
trionale, confined to northern localities, generally
growing on rocks and insinuating its delicate roots in
the deep crevices and among the débris of irregular
shelving and prominences, not receiving any apparent
injury from the rigour of winter or the scorching
heat of summer. Although such is its hardy nature,
it nevertheless does not flourish when taken from its
native rocks and brought under artificial cultivation.
What has been stated regarding the shade and
moisture-loving Trichomanes, and the rock-loving
Asplenia, may be viewed as the extremes in the nature
of those Ferns that do not readily conform to ordinary
cultivation; but, on the other hand, the greater
number are not particular in their choice of place of
growth, either in a wild state or under cultivation.
Examples may be cited of species conforming to the
most untoward and varying influences; for instance,
the common hart’s-tongue Fern (Seolopendriumvulgare)
maintains itself in situations of the most opposite
kind, having great predilection for the works of man,
whether elevated in the air or sunk below the surface
of the earth; such as stone, brick, or turf walls,
embankments, hedge-banks and road-sides, pits,
quarries, or deep open wells. In either place it multi-
plies freely, conforming itself to the various atmo-
spheric changes ta which those situations are liable.
Such being the case, it is not surprising to find it
assume different sizes and forms; in dry places it is
only a few inches in height, while in open wells, such
x
306 FERNS: BRITISH AND FOREIGN.
as may be seen in some of the nurseries and market-
gardens near London, it produces fronds 2-3 feet in
length. The Asplenium Trichomanes, A. Ruta-muraria,
Ceterach oficinarum, and Polypodium vulgare, are also
wall and tombstone-lovers, and may be called our
domestic Ferns.
Another remarkable example of a Fern making
itself at home under extremes of temperature and
moisture, is Pteris longifolia, a species having a wide
range throughout the tropical and sub-tropical regions
of both hemispheres. On the island of Ischia (Bay
of Naples) it is found luxuriating within the influence
of the hot vapours rising out of the cavities left by
extinct volcanoes, growing in soft muddy soil at a
temperature ranging from 140° to 160°. In our
hothouses its spores vegetate abundantly upon all
moist surfaces, and in the crevices of brick walls.
Plants of it are nearly always to be found over the
openings of hot-water tanks, and it has been seen
in crevices of the walls outstde hothouses, or even
under iron gratings, where it could receive but little
light, and where the temperature was often near the
freezing-point. In the dry air of the Cactus-house
plants of it have produced fronds from 2-8 feet in
length.
In general the fronds of Ferns remain Iong in a
perfect state; the exceptions to this rule are com-
paratively few, and these are chiefly supplied by the
natives of climates alternating with seasons of heat
and cold. But as many species are wanderers and
conform to th: effects of various clmates, it is no
wonder to see some of our native Ferns assuming
the evergreen habit of their -foreign allies, when
CULTIVATION, 307
grown in a temperate house ; of which Polystichwm
aculeatum, Lastrea dilatata, and L. Filia-mas may be
cited as examples, as also Asplenium marinum. In
1820 I found plants of it, having fronds from 2-4
inches in length, growing in a cave facing the German
Ocean, on the east coast of Scotland ; of these, two
plants have been grown at Kew from that time; of
late years, one in the Temperate and the other in the
Tropical House. These became fine cespitose plants,
with fronds varying from 1-1} foot in length; the
greatest length being attained by the plant in the
Tropical House, even assuming the character of a
species native of the West Indies and Tropical
America, and quite unlike the original plant; thus
showing that although at home in the cold, sunless
cave, it can well appreciate a better fed and warmer
abode. Exceptions to this rule are some alpine
species, Ferns in that respect being analogous to our
ill success in growing many alpine flowering plants.
This is no doubt partly owing to the difference of
atmospheric density, and the varying influences of
temperature and moisture common to the sea-level of
this climate.
The species of cold climates truly deciduous, produce
their fronds from an underground creeping sarmentum,
of which Péeris aquilina, Sitolobium punctilobium,
Onoclea sensibilis, Anchistea virginica, Lorinseria areo-
lata, Leucostegia immersa, and Phegopteris aurita are
the principal examples. On the other hand, instances
of tropical species periodically losing their fronds
without any apparent cause, such as by undue excess of
heat and moisture, are Phymatodes oxyloba, Pleuridium
palmatum, P. venustum, Drynaria propinqua, several
x2
308 FERNS: BRITISH AND FOREIGN.
species of Davallia, and a few others. Their rhizomes
remain perfectly naked during the winter, and seem
to be constituted for a season of dry rest.
These obversations are sufficient to show how
certain species will grow under the most opposite
conditions, and that a knowledge of them materially
assists our efforts of cultivation, which I shall pro-
ceed to treat of under two heads,—special and
natural,
II. Specrar on Pot Corrtrvatton.
ITH the exception of those species sufficiently
hardy to bear the cold of this climate, the
whole family of Ferns can be artificially and most
luxuriantly grown, under but two different scales of
temperature, and with as little difference in other
respects. There can be no better instance than that
afforded by the immense collection nowat Kew,* where,
in the Tropical Fern-house, a great many species from
various climates are placed under an average tempera-
ture of 60° to 70°. In this houset are species from
nearly all the tropical and sub-tropical countries
of the world, and, being in one compartment, they
are all alike subject to the same amount of atmo-
spheric moisture as of heat. However, in respect
to moisture supplied to their roots, the amount
is varied according to the nature and require-
ments of the plant. The same remarks apply to
the Temperate House, of course with a reduction
* May 1864, + Length 130 feet, width 34 feet.
CULTIVATION. 809
of moisture as well as temperature, the latter ranging
trom 40° to 50° in winter, that being the only period
of the year when artificial heat is required. In this
house* are growing, in the greatest luxuriance, species
from Australia, New Zealand, China, Japan, North
India, elevated portions of Ceylon, South Africa,
North and South America, and other elevated regions
within the tropics. The whole collection is grown in
common deep or shallow pots, pans, and tubs; the
latter, however, are only used for the large species
of Tree Ferns and Angiopterts ; and though such a
system is the least natural in appearance, yet it is
most convenient and found generally consistent with
the prevailing fashion for in-door horticulture; this
mode of cultivation, being mostly adopted, merits our
first consideration. In the tropical and sub-tropical
localities, where heat and moisture are abundant, by
the manner in which they grow and the various
positions they occupy, soil is of but little importance,
except for affording their roots the means of obtaining
permanency of position. This is evident by the remains
of native soil adhering to the roots of imported plants,
showing that Tree and other large Ferns in some
places grow in stiff adhesive, red clay: To imitate
this soil in pot culture is attended with no success,
and it is remarkable to see how quickly the roots of
newly-imported plants take to the fine loam and
peat in which they are potted. In cultivation, how-
ever, the great beneficial influence of the natural
atmosphere is not obtained, and the soil is therefore
of great importance, necessitating caution in its
* Length 82 feet, width 13 feet.
310 FERNS: BRITISH AND FOREIGN.
selection. Taking a view of the large number of
Ferns now in cultivation, their requirements as regards
soil, and mode of growth, we easily recognize two
classes—terrestrial and epiphytal. This division is
important. For the first class, by far the most ex-
tensive, it is necessary to use the soil of a finer or
coarser consistence, according to the delicate or
stronger character of the plant. In it there is a
more varied character of vernation, by which their
terrestrial character is indicated and very easily recog-
nized. Natural affinity of course is not taken into
consideration. However, with tolerable accuracy in
this respect, the genera Phegopteris, Lastrea, Ne-
phrodium, Adiantum, Asplenium, and Sitolobium give
examples of the chief variety of vernation indicative of
the terrestrial nature ; viz.,erect, ceespitose, decumbent
or sub-hypogeous, or, as in Nephrodium unitum, and
N. pteroides, which, although with a true sub-scandent
sarmentum, they, however, prefer the firm soil of the
terrestrial group, a compost of two-thirds peat and
one loam, with abundance of sand, according to the
size of the plants for which it is to be used.
In potting Ferns, an over-depth of soil should be
avoided, as well as a great depth of drainage. The
one promotes stagnancy, the other gives an oppor-
tunity for the most vigorous roots to descend
among its particles, and oftentimes they become too
dry when the soil is apparently moist enough, and the
tips of the foliage are in consequence injured, and
the plant disfigured. This applies more strictly.
to species of small and delicate structure, such as
Asplenium firmum, A. dentatum, A. decussatum, &c.,and
for such the shallow pot is best adapted, the width of
CULTIVATION, all
which being considerably greater than the depth,
gives a good extent of surface, and renders a deep
drainage unnecessary. Such a kind of pot, of course,
is most suitable for all species of decumbent or
creeping habit, and those producing rapidly a great
number of offsets.
The best time for repotting (or shifting, as it is
termed) Ferns, more especially those of the terrestrial
division, is at the end of February, or during March,
as soon after that time they commence growth; but
any time of the year, except winter, will suit them.
The operation of potting is a matter of little difficulty,
nor are the necessary utensils or materials very varied—
a firm potting-bench, several different sizes broad-
pointed firming-sticks, and a garden trowel. It is
also necessary to have a few extra pots of different sizes,
of both the deep upright and shallow flat kinds, also
a quantity of material for drainage, which may consist
of broken soft brick or hard knobs of old mortar; but
for general purposes, potsherds, broken into different
sizes, are most commonly used, and well known by
the name of “crocks.” The removal of the plant,
with its ball of soil and roots, is readily effected by
spreading the left hand over the surface of the soil,
allowing the plant, when not too large, to come
between the fingers; then, by inverting the plant and
pot, and giving the rim of the pot a gentle but sharp
tap with the right hand on the edge of the board ; if
the roots are in a healthy state, the ball will slip
easily out of the pot. In all cases, athorough drainage
is indispensable, and to ensure this, the draining
material should be of two sizes, the larger size
at the .botteom, with a concave large crock over
312 FERNS: BRITISH AND FOREIGN.
the hole of the pot, the concave side downwards:
there should be two, three, or four holes at the side,
quite at the base, made in all pots larger than eight
inches in diameter, the number of holes of course
according to the size of the pot. This is the more
essential when the pots are placed on smooth benches,
as of stone or slate, the capillary attraction between
the pot and bench retaining the superfluous water,
and causing much stagnancy. After properly arrang-
ing the drainage, a small quantity of fibry material
should be placed on it to keep the new soil from
mingling with it. No plant should be repotted in
a wet state, nor should it be allowed to become dry
enough to cause it to flag, as this will not only
give an immediate and severe check, but in giving
water after it is potted, it will percolate the new soil
(which should always be used in a moderately moist
state) without entering the dry ball; if, from bad
drainage or exhaustion of the soil, it has become
impure, so much of it and of the old inactive roots
should be removed. In such cases, as small a pot as
possible should be used, which will admit the pre-
served roots freely, being carefully laid out by the
fingers amid the new soil; for if in too large a pot,.
the soil becomes sour before any vigorous roots
have entered and promoted drainage, as they are
tardy to perform their functions vigorously after
having been necessarily so much disturbed. However,
when in a healthy and vigorous state, such space
should be given it that will admit the fingers or
both hands freely by its sides (according to the size
of the plant) in inserting it in the new pot, where
it should be so placed that its crown, if of the
CULTIVATION. 313
erect vernation, will be nearly level with the rim. The
soil should be carefully introduced, and made firm
round the sides by pressure of the firming-stick and a
few thumps of the pot (if easily handled) on the
potting-bench, if not, the stick must suffice ; a space
of about half an inch, more or less, according to the
size of the pot, should be allowed at the surface to
contain as much water as would moisten the whole
ball.
The size of the plants will depend upon the amount
of space and'the number of species in the collection.
For an amateur’s collection in a small house, very fair
specimens may be grown in pots from eight to twelve
inches in diameter, after the shift into the largest-sized -
pot, and with good management the plant will not
require any repotting for two or three years. By
that time the running and czspitose kinds, such as
Adiantum and Gleichenia, will have become exhausted
in the centre; the ball will, therefore, require division
by passing a sharp knife through it, taking care not
to injure the young growths next the sides of the pot;
the most healthy portions to be selected for repotting,
to become the new representative plant of the collec-
tion; and, if proper care is taken, the fronds will
suffer but little injury. In operations of this kind,
some gardeners entirely shake out the soil, cut away
the whole of the fronds and roots, in order to make
the plant, they say, come up strong. This may not do
much injury to certain plants, such as bulbs, tubers,
fuchsias, and such-like plants that rest in winter; but
for Ferns it is a great mistake; it so weakens the
plant that it takes a year or more to be worth looking
at, and, indeed, some never recover. In Adiantum
314 FERNS: BRITISH AND FORKIGN.
and other genera, the roots are naturally of a dark, or
even quite black colour, and it is known to have been
considered by some cultivators, not well versed in the
nature of Ferns, as dead, and accordingly theyare totally
removed, even although with evidence of the plant
being healthy and vigorous. The healthy state of the
plant is readily explained, for on examining the nume-
rous points of the black roots, each will be seen to be
of a pale colour, which are the active feeders of the
plant; and, in shifting, great care should be taken not
to bruise or injure them.
The same directions apply to the gigantic Tree
Ferns and species of Angiopteris. The soil for them,
however, should be of a coarser nature when of a size
to require the largest-sized pots or tubs; in no case
is it desirable to give more than three inches extra
space at one shift, even in the largest pot or tub. In all
pots above one foot in diameter, the drainage should
be elevated in the centre. When the shift is large, an
inverted pot should be placed in the new pot, placing
a layer of drainage round it, the height of the pot to
be such that the crocks of the old ball (which must
not be taken out) rest on the top of the new drainage
or pot. By this the outer circle of bottom roots are
not pressed together, or crushed by the weight of the
ball, which is sure to be the case when the ball is set
on a level surface, it also keeps the ball to its proper
height. This mode of drainage has been the constant
practice at Kew with all large shifts, not only for Ferns
but Proteacee and all fine fibrous-rooted plants, the
nature of which is to extend outwards and downwards
through the new soil, ultimately forming a web of
roots against the sides of the pot. In the ordinary
CULTIVATION, 315
practice of potting, it is a rule with many gardeners _
to remove all the old drainage crocks, the centre thus
becomes a mass of soil, which in time becomes com-
pact, inert, and useless, liable to become stagnant,
and in time causing the plant to sicken. This is
avoided by preparing and retaining the old drainage.
This mode of drainage is, however, only necessary for
plants required to be permanent in botanical collec-
tions. Its utility in prolonging life is verified by some
yet existing plants of Proteacee at Kew, some of
which are forty, and others above sixty years of age.
What has here been stated may be considered as a
general rule for the greater number of the Fern
family, such as are usually found in ordinary collec-
tions of a hundred species, more or less. But in this
extensive family there are many possessing some pecu-
liar nature, and a few examples of these require to be
specially noticed.
The group which has been termed Epiphytal,
consists of the genera Davallia, Goniophlebium,
Drynaria, Phymatodes, Pleuridium, Platycerium, Lo-
mariopsis, Polybotrya, and others of like habit, charac-
terized by a true and highly-developed rhizome or
sarmentum, generally creeping, and adhering by
their fine fibrous roots to the surface on which they
grow. Their positions in their natural homes are
generally on more or less perpendicular surfaces of
moist and shaded places of rocks or ordinary. soil,
where decomposing vegetable matter abounds as a
surfacing. They are often found on trees, and with
many species of Polybotrya and Lomariopsis this
seems to be their true position, and often by their
vigorous growth, the trees are clothed with them in
316 FERNS: BRITISH AND FOREIGN.
the manner of our common ivy. It will be under-
stood by this that an open loose soil is essential for
the whole group; good fibry peat is therefore all
that can be desired, and on account of their creeping
habit, as great an extent of its surface as possible is.
necessary to be gained in potting. To this end the
shallow pan is indispensable; by an ordinary deep
pot being inverted in the pan and covered with a
layer of peat, of a thickness according to the size of
the pan or pot used (at a proportion of two inches
thickness for a pan of one foot diameter) carried up
over the inverted pot in a conical form, the whole
pressed firmly together. The rhizomes are fixed to
its surface by pegs, and they will soon attach them-
selves by their own roots. The height of the cone
will be according to the character of the plants; for
Pleopeltis, Anapeltis, Niphobolus, &c., a cone of from
6-10 inches high, with a base from 1 to 1} foot in
width, will make handsome specimens. The genus
Oleandra comes under Epiphytal, the rhizomes elon-
gating rapidly, and in their natural places adhering
to trees and moist rocks. 0. neriformis is, however,
truly terrestrial, producing roots from its woody erect
rhizome after ascending a few inches from the’ soil..
The other species in cultivation, however, cannot be
well suited by the conical mode, or being trained on
straight sticks. An open netted wire cylinder, about
three feet high, and about six inches diameter, answers
the purpose exceedingly well. The cylinder is filled
with peat, and by fixing the rhizomes round the
bottom of the cylinder, the roots will soon adhere to
the peat through the meshes, which being kept moist,
a rapid growth is stimulated, and the cylinder soon
CULTIVATION. 817
becomes furnished with beautiful projecting fronds.
Ifa greater height is desired, another cylinder may
be added to the first. This mode is equally useful for
Stenochlena, Polybotrya, Lomariopsis, &c. In.respect
to species in which the rhizomes elongate slowly, the
upper part of the cylinder is bare and unsightly for a
“ time. This may be obviated by having a short cylinder
and heighten it by adding short lengths when required.
The genus Elaphoglossum is generally epiphytal, and
as some of its larger growing species, such as H. calle-
folium and FE. latifolium, being decumbent cespitose
in vernation, it is not necessary to have the soil raised
much above the rim of the pot, which should be of
the flat kind, and three inches depth of soil is quite
sufficient. Many smaller species of this genus are
most lovely objects, and some of them are not yet in
cultivation. Of the whole Fern Family the Platycerium
may be considered the most grand, beautiful, and
extraordinary; and it is thoroughly typical of this
epiphytal group. ‘Its natural position of growth is
sometimes on moist rocks, but usually on the trunks
and larger branches of trees. The spores becoming
lodged there, germinate, and, sending out spongy
fibrils, a little plant, like a circular disk, adhering to
the tree, analogous to a foliaceous lichen, is formed,
each succeeding disk (frond) becoming larger and
overlapping the preceding one. In time the older
ones loose their vitality, and by this mode of growth
envelop, or nearly so, that portion of the tree whereon
they grow in a dense, thick, spongy mass, among which
the roots insinuate themselves and receive nourishment.
As equivalent to this, in cultivation blocks of wood
are mostly used ; but they are objectionable from their
318 FERNS: BRITISH AND FOREIGN.
constant liability to breed fungi and harbour insects.
On that account pots are preferable, not of the ordi-
nary shape, but with less difference in the diameter at
the base than at the top, and with a wide opening at
the side, extending about a third of its diameter from
the rim down to within a sixth of its depth from the
base (this will leave the pot in much the same form
asascoop). . Thorough drainage should be given, and
a material of very fibry peat will suit. The crown of
the plant should be placed about the centre of the side
opening from where the fronds will be produced. In
time, the sterile fronds will spread in all directions, but
mostly upwards, and quickly obscure the pot, and the
fertile ones will hang looselydownwards. Their position
in the house should be elevated, and, if possible, against
a wall or partition. Platyceriwm alcicorne increasing:
rapidly by offsets, requires a’ considerable extent of
surface ; consequently rough sandy peat, arranged in a
conical manner on a shallow pot, to which the plants
will soon attach themselves, is most suitable, and
which, if fancy leads, may be suspended from the roof
of the house by a-strong wire.
For the large-growing species of Drynaria, Gonio-
ahlebium, and Phlebodium, it is not necessary to raisé
the soil much above the level of the pot; their
fleshy rhizomes soon reach the margin, to which
they cling, and if standing on a moist surface, or
near water, they creep down the sides of the pot.
In Davallia pyxidata, D..ornata, and others, the
rhizomes are what may be termed aérial, rising con-
siderably above the soil, often extending to a distance
beyond the edge of the pot, and, as they in these
cases do not produce roots, the rhizomes in time
CULTIVATION. 319
(three or four years) become weak, producing small
fronds ; it therefore becomes necessary to remove the
older parts and re-pot the younger portions in fresh
soil.
Very interesting and natural examples may also
be had by placing common cylindrical red chimney-
pots or drain-pipes in a pan wider by two inches than
the base of the cylinder,—this space to be filled with
soil and planted; the rhizomes will soon become
attached to the surface of the cylinder and in time
will cover the whole, forming a handsome pillar of
fronds. In order to maintain a proper. degree of
moisture, a pan of water may be fixed inside the
cylinder, and if a piece of woollen cloth is placed in
the water, in contact with the cylinder, a constant
moisture will be kept up congenial to the plants.
The top of the cylinder is also useful for placing
on it some of the pendulous species; for instance,
Phymatodes geminata, Goniophlebium dissimile, G.
neriifulium, Campyloneuron angustifolium, Elaphoglos-
sum Herminiert.
In Scheilolepis subauriculata and S. verrucosa the
fronds are long and pendulous; this necessitates the
plants being placed in an elevated position, such as
suspended from the roof of the house in shallow wire
baskets, the inside of which should have a lining of
sphagnum moss, which assists to retain moisture, and
also gives a clean and neat appearance. Plants of
these two species thus treated have at Kew produced
fronds 12 feet in length, hanging down in a very
graceful manner. There are several other Epiphytes
of special interest, such as the species of Vittiria
and Haplopteris, which hang down from trees like
320 FERNS: BRITISH AND FOREIGN.
bunches of grass, as also the remarkable Ophioglossum
pendulum, which may be likened to ribands or bands
hanging loose and waving with the wind, often many
feet in length. These plants succeed in a small
quantity of soil, firmly fixed in pots, with pieces of
soft stone or potsherds, and the pot hung against a
shady wall or pillar. For this purpose the pot should
have a flat back, with the front rim lower than the
back, so as to allow the fronds to hang quite free of
the pot.
Neottopteris Australasica, and a few Aspleniums
nearly allied, such as A. sinuatum and A. crenulatum,
of precisely the same mode of -growth, are of erect
fasciculate vernation. Their roots being of peculiar
mossy and delicate nature, they are not adapted
for deep insinuation of stiff soil, but are rather
what may be termed aérial. Two-thirds of their
mass is produced above the surface of the soil.
Substantial but open material is therefore required,
of very rough, fibry peat, and porous, broken bricks,
or soft sandstone, in equal parts; very little pot-
room is necessary; a shallow pot of 18 inches
diameter, with such material, will support a plant
of two dozen fronds, and none less than 8 feet
6 inches long and 8 inches broad, with a stem a foot
high, and as much through, principally composed of
its mossy roots forming a spongy mass. As an
instance of the long life under regular treatment may
be cited the original plant of Neottopteris Australasica,
which was imported in 1825, and is now (1864) a
magnificent plant, in perfect health, having received
but few shifts the whole of the forty years.
This is, however, far surpassed in size by the mag.
CULTIVATION. 321
nificent Neotiopteris muscefolia, which is described by a
Penang correspondent in the following words :—“ TI
saw two fine specimens of the Birds’-nest Fern ; each
had between forty and fifty perfect green leaves; the
average length of the leaves was six feet, and from
one foot to fourteen inches across in the broadest part.
They were growing on each side of a doorway; when
I was walking up to them I thought they were American
Aloes.”
The remarks that have been previously made
respecting the nature of the rock-loving Asplenie are
equally applicable to others that inhabit rocky places,
such as certain species of the genera Notholena,
-Myriopteris, Cheilunthes, &c.; some are rocky-coast
plants, others are mountain, being found in elevated
situations within or near the tropics, where they are
subject to the heaviest showers and most powerful
sun, their surfaces. being generally furnished with
beautiful scales or woolly covering, which resist in a
great measure the action of the sun. They all seem
very impatient of moisture under cultivation, especially
in winter. A material composed of finely-broken and
mixed sandstone, bricks, old mortar, and a small
quantity of sandy loam, suits them, placed in a position
of the coolest shade, with abundance of moisture in
summer, and in winter very little water, just sufficient
to keep the soil slightly moist; a comparatively dry
atmosphere and as much light as may be obtainable
at that dull season, with a temperature not below 36°.
As a general rule, the above may be considered appli-
cable to all small-growing rock species of temperate
regions, including the natives of this country. The
species of the genus Gymnoyramma, especially those
¥
322 FERNS: BRITISH AND FOREIGN.
with farinose fronds, commonly called Gold and Silver
Ferns, are also very susceptible of moisture; they.’
should never be syringed, or water allowed to fall on
their fronds, as the: farina, being loose, is distarbed by
the water, and running down, gives the appearance as
if the plants were smeared with dust. They, however,
differ from the preceding, requiring more light, and
the temperature of the Tropical House. The species —
of Gymnogramma vary very much in habit, as regards
size and circumscription of the fronds, G. trifoliata
having fronds from three to. four feet high, while in _
G. cherophylla and G. leptophy lla they are fragile, and
average from two to six inches in length. These two
species are peculiar in being, with the exception. of
Ceratopteris thalietroides, the only known truly annual
Ferns. G. cherophylla grows freely ; its spores vege-
tate abundantly throughout the house, often as a
weed. G. leptophylla is, however, not so free in its
growth. When its fronds decay,’ the pot should be
covered with a piece of glass, and put in a dry place
until the proper season arrives in spring, when the
application of moisture will cause the latent’ spores to
vegetate. G. flewuosa differs from, the whole of the
genus in having prelonging fronds, which climb in a
rambling manner over bushes, like those of Lygo-
dium, noticed in another page: it will probably here-
after form the type of a distinct genus. The beautiful
genus Lindswa, of which no less than sixty species are
described in Hooker’s “ Species Filicum,” are, with
few exceptions, natives of the tropics of both hemi-
spheres; in my Catalogue of 1857, only two species are
recorded as being cultivated; but, within these few
years, the number has thigronaed to fourteen, the greater
CULTIVATION. 823
part having been introduccd from Guiana by Mr.
Backhouse, of York, and on account of their slow
increase, plants of them are yet very rare: they are
natives of generally the lower regions of tropical vege-
tation, growing in open places amongst herbage of
small plants and grass, or on the skirts of woods, and
sometimes under the shade of trees, the soil being
very poor, sometimes almost nothing but sand or
stony débris, in which their sarmentums are partially
hypogeous, the soil and air never being much below
the temperature of 80°, and although almost daily
subject to the influence of tropical thunder-showers,
yet, on account of the nature of the’soil, the surface is
never over saturated. Finding that they do not flourish
underthe medium temperature of a Tropical Fern House,
_it therefore is necessary that a special part of the house,
on the principle of a Wardian Case, should be adapted,
so that a moist air of 80° may be steadily maintained,
and the plants occasionally sprinkled overhead, taking
care that no superfluity remain in the soil, which
should be no more than moist.
Like Lindscea, the curious and interesting genus
-Schizea does not readily conform to cultivation ;
plants of Schizcea elegans have often been freely im-
-ported from Trinidad, and although tried in various
ways in high and moderate temperatures, it cannot be
said they have yet become established. Under the
Wardian Case, the native imported fronds remain for
a considerable time fresh, and sometimes new fronds
show themselves, but fail to come to maturity. Ina
letter lately received from Mr. Prestoe, in Trinidad, be
informs me that the Schizwa elegans grows in solitary
patches in loamy soil, covered with three or. iour
x2
324 FERNS : BRITISH AND FOREIGN.
inches of leaf soil, in company with Adiuntum and
other Ferns, as well as much Cyperacece, the whole
forming a dense undergrowth in woods. From this it
appears there is no great peculiarity from other Ferns
as to its place of growth, and that our ill success
arises in consequence of the nature of the plant, being
difficult to re-establish after being taken from its
native soil, of which there are many instances; the
common Pteris aquilina being a familiar example.
The numerous and beautiful species of Hymeno-
phyile, called Filmy Ferns, merit particular attention.
They are natives of both tropical and temperate
regions, and where they abound are generally found
in shady moist woods, clothing the lower parts of the
trunks of trees, especially Tree Ferns, or on dripping
rocks, or surface-soil of the deepest ravines, rarely in
exposed situations. In order to surround them with
a moist atmosphere, adaptations must be resorted to,
such as moveable cases upon the Wardian principle,
hereafter to be described, and which are essentially
necessary for the cultivation of this delicate tribe.
Such, indeed, is their delicacy, that if once allowed to
become dry on the surface of their fronds for any
length of time, a rusty and shrivelled appearance will
soon follow. Having, as already stated, succeeded in
cultivating Trichomanes radicans, it becomes obvious
that other species could also be made subservient to
cultivation ; of which there is ample evidence to be
seen at Kew, where there are forty cases* of different
sizes, arranged on a stone shelf on the north side of
the large Fern House, in such a position that they
* May, 1864,
CULTIVATION. 825
receive but little sun in winter, and are densely shaded.
in summer, all filled with patches of fine fronds, of a
number of different species, varying from the delicate
hair-like Trichomanes tricvideum, not more than two
inches high, tothe robust 7. anceps and T. radicans.
They are grown in square shallow pans and boxes,
well drained in the ordinary way, and having about
two inches of peat soil mixed with nearly half its bulk
of sand and small broken potsherds; but soft sand-
stone is best. For the creeping sorts the soil should
be raised in the form of a mound, and for those that
have long-extending sarmentums, if soft stone cannot
be had, it is desirable to invert a pan or common deep
pot, covering it with a layer of soil, as already ex-
plained, to which the plant will cling, and soon form a
green hillock: junks of wood answer the purpose;
but im a moist, close, and warm atmosphere, fungi
and insects breed, and in a short time the wood
decays, causing unnecessary disturbance of the whole
mass of the plant.
The singular genus Lygodium, and its ally Lygo-
dictyon, grow naturally in firm soils, generally amongst
trees and bushes, their wiry, flexile, climbing fronds
growing over and involving everything within their
reach in the most intricate complexity. In most
Ferns, the whole of the divisions of the fronds are
formed in the nascent or bud state, and are unfolded
as the fronds elongate ; when the whole of the deve-
loped parts are unfolded, the frond ceases further ex-
tension. This is, however, not the case in Lygodium,
Salpichlena, Pellea flecuosa, Gymnogramma flexuosa,
Odontosoria aculeata, and a few others, the fronds of
which are of indefinite extension, their apices con-
326 FERNS : BRITISH AND FOREIGN.
tinuing to grow and produce lateral pinne, in every
way analogous to the development of branches and
leaves in woody plants. The climbing and rambling
nature of these plants necessitates some kind of sup-
port when cultivated in pots, which may consist of
open wire trellises of # cylindrical form, varying in
height from three to six feet or more, which, if fixed
to a pot ten to twelve inches in diameter, the plants
will, with careful training, become handsome speci-.
mens ; they are also well adapted for covering trellis
against wall or pillars.
The fronds of the much admired genus Gleichenia
are also indefinite in extension, and some of the
smaller species,when growing amongst bushes, assume
a climbing habit; but in G. furcata, G. dichotoma, G.
Cunninghami, and G. flabelluta, the fronds are rigid
and erect, produced from a more or less stout or slender
sarmentum, which is either superficial or creeps a
little below the surface of the soil; therefore shallow
pans or boxes are best suited for these plants ; and
with attention specimens of considerable size may be
attained, as, for instance, at Kew a plant of G. flabellata
measured twelve to thirteen feet in circumference and
four and a half feet high, consisting of a thicket of
fine fan-like fronds. i
It may be expected that in such an extensive family
some species would be found bearing the appellation
of aquatics, but such is not the ease; for although
many species grow in wet places, such as Osmunda
regalis and Acrostichum aureum, both of which love
water, but also flourish even in dry places, the only
Fern really entitled to be called a water Fern being
Ceratopteris thalictroides, and which is aiso singular
CULTIVATION, ‘ ‘327
__in another respect—in being one of the few Ferns that
are only annual. It is widely dispersed throughout the
tropics, growing in wet places, often flooded; its
sterile, viviparous fronds floating on or below the-
surface of the water, as may be yearly seen in the
Victoria Lily tanks at Kew. Being annual, care must
be taken to preserve spores, which in the spring
should be sown in a shallow pan of loamy soil made
wet like mud, kept moist ; and when the plants are of
sufficient size, the pan may be either filled with water,
or be placed about an inch deep ina tank. Acrostichum
aureum is mentioned as growing in wet places; for
instarce, in Jamaica and other of the West-India
Tslands it is described as taking the place of the Euro-
pean Typha latifolia, attaining the height of seven or
eight feet. By imitating its natural condition, placing
it in a pan of water, or ina tank, specimens have been
grown at Kew to the height of six feet; but although
it enjoys and luxuriates in water, it also grows in dry
places, its height then not exceeding one to two feet,
and often with simple fronds.
_ The delicately beautiful Selaginellas, on account of
the large share of notice they receive and meet from
the admirers of the Cryptogamic family, deserve
here an especial notice. Originally the species were
included under Lycopediwm ; but have been separated
on account of differing in.the character of their
spore-cases. The species are numerous, and their
habit and free growth mark them as a very distinct
_ group from true Lycopodium. With a knowledge of
a few species, two very distinct modes of growth are
easily understood; the upright or climbing ones, such
as 8. Africana and S._ filicina, are exanple, and the
- 828 FERNS: BRITISH AND FOREIGN.
decumbent or creeping species, such as S. serpens,
S. uncinata, S. Galiottii, &c., are examples. The
latter delight in a light soil, composed of fibry peat
and well-decomposed leaf-soil, free from pieces of
wood (as it generates fungi), with a little sand inter-
mixed, and require a depth of only about two inches
in ordinary round pans, with ample drainage. Tf,
however, good specimens are desirable, with an
arrangement on a bench or low shelf, square pans
about twice their height at the back as in front, arein
every way preferable, and as at Kew, arranged alter-
nately with the cases of Hymenophylle, with which
they harmonize. The former-mentioned species, on
account of their robust habit and strong rooting cha-
racter, require the shallow pot, and a similar soil as for
the last, but with a small proportion of light loam.
The magnificent Selaginella levigata, if supported by
wires against a wall, becomes, in a remarkably short.
time, the loveliest object to be found in a Fernery.
The species of the Rosulate section differ from the rest
of the genus in having erect, fasciculate vernation, the
frondules rising from a central developing axis, and,
as in S. lepidophylla, spreading out nearly horizontal,
and overlapping one another, forming a beautiful
green rosette, about six inches in diameter ; when dry,
they turn upwards and inwards in an involute manner,
the whole mass of the plant thus forming a firm ball,
which, on becoming moist, again expands; and even
perfectly dead plants, when expanded, seem as if alive.
In that respect being analogous to that of the insig-
nificant cruciferous plant called the Rose of Jericho
(Anastatica hierochuntica): hence the balls of this
Selaginella are frequently met with in curiosity-shops
CULTIVATION, 29
folie)
under the above name. This hygrometric property
seems to indicate that this and its allied species are
subjected to occasional drought in their native localities.
But although they may frequently undergo this pro-
cess in nature with impunity, it is, however, not
desirous to be too often repeated on plants artificially
cultivated. Shallow pots, four to five inches in
diameter, best suit this section; and being firm
holders, it is necessary to insert pieces of soft sand-
stone or potsherds amongst the shallow soil, so as
to fix the plant firmly in the pot. The extremely
delicate nature of the tissue of these little plants neces-
sitates that a position the shadiest and moistest in a
Fernery should be assigned to them. 8S. Willdenovii,
S. filicina, and others are termed deciduous from their
fronds decaying altogether in the early spring; new
fronds, however, soon begin to grow.
The species of true Lycopodium are also numerous,
some being terrestrial and others epiphytal, hanging
from the trees like various-sized cord, one to four feet in
length. Of the terrestrial, five are natives of thiscountry;
they, as well as several beautiful species that have from
time to time been introduced from North America,
refuse to become domesticated under ordinary treat-
ment; to which must be added the wide-spread and
beautiful tropical species L. cernuum. Hqual ill success
attends the epiphytal species ; for although frequently
imported, they are yet but poorly represented in our
collections ; they adhere to trees by producing many
fibrous roots, which ultimately become a large spongy
mass.
The extra height of roof required for the growing of
good examples of Tree Ferns excludes them from many
330 FERNS: BRITISH AND FOREIGN.
amateur collections. This has also been the case at
Kew for the last twenty years, especially as regards
Tropical Tree Ferns, many fine plants succumbing to
the make-shifts that of necessity had to be resorted
to after they had attained a certain height ; but by
beginning with young plants, they may be grown fora
number of years in houses of the usual average height
of ten to twelve feet, as also the large fronded tree-
like Lastrea villosa, Intobrochia podophylla, Asplenium
striatum, Hemidictyon marginatum, and many others
of like habit. The latter, at Kew, in a 20-inch pot,
produced beautiful fronds, seven feet in height, and
which might, with encouragement, soon be made to
produce them equal to those of native growth—fourteen
feet. But in order to get rid of the inconvenient and
unsightly look of large pots and tubs, it is best to
adopt for these plants the system of natural cultivation
explained further on.
In the “ Species Filicum” about one hundred and
twenty species of Tree Ferns are described; but,
according to Mr. Moore’s ‘“TInudex Filicum,”’ the
number amounts to nearly two hundred. They are
widely distributed, chiefly within the tropics. They
love shade and, solitude, and are generally found at
elevations of from three thousand to five thousand feet
in the humid regions. In the southérn hemisphere
they, however, extend much beyond the tropics, their
southern limits being New Zealand, Norfolk Island,
New South Wales, and Tasmania, where they grow at
a lower elevation than within the tropics. On Mount
Wellington, in the latter island, Dicksonia Antarctica
is found in the greatest abundance, at an elevation
of from one thousand five hundred to two thousand
CULTIVATION. 831 .
feet, attaining the height of from eighteen to twenty
feet, growing only in damp. places, generally gullies,
where the sun rarely penetrates, and where they are
sometimes covered with snow, as represented in the
vignette on the title-page. This suggests that, with
proper selection of situations, they might live in the
open air in the south and west of England, as also the
mild climate of Argyleshire, where shaded ravines
and gullies may be found similar to those of Mount
Wellington.
With few exceptions, Tree Ferns readily conform to
cultivation. Alsophila capensis, A. excelsa, A. Australis,
Oyathea deulbata, C. medullaris, Dicksonia antarctica,
and D. squarrosa, grow freely in the Temperate House ;
the lofty Alsophila glauca, the beautiful tessellated
stem of Cyathea arborea, with its crown of fine
fronds, and the broad shining fronds of Hemitelia
horrida, assume a grand appearance in the Tropical
House. In general, the stems of Tree Ferns are of
sufficient size to warrant the name of trees; but in
many the thickness is more apparent than real, the
diameter of the woody centre being often only a few
inches, but in many cases covered with successive
productions of out-growing aérial roots, which become
hard and wiry, and by their interlacing, form a com-
pact mass; the points of these roots are, however, the
active feeders, and if a layer of fresh soil is occasion-.
ally placed round the base of the stem, their growth
will be promoted, and vigour given to the plant. The
most fastidious Tree Fern to cultivate is Dicksonia
arborescens, anative of St. Helena. It was first intro-
duced to this country in 1786, and many times since,
but refuses to become established, either in a tropical
332 FERNS: DRITISI. AND FOREICN.
or temperate house. This species, as well as the whole
of the species of true Dicksoni@, are characterized by
the base of the stipes being clothed with a more or
less coating of beautiful, articulated silky hairs; in
some species of Cibotium, the quantity is so great, that
in the Sandwich Islands it is collected, and ship-loads
of it sent to California and Australia for stuffing
cushions, beds, &c. Another remarkable species of
this alliance is Cibotium Barometz, a native of China
and other parts of Hastern Asia; its fronds attain the
height of twelve to fourteen feet (even in this country) ;
they rise from a thick decumbent caudex, which is
densely covered with silky hairs, as above described ;
lying on the ground, it has the appearance of a woolly-
clad animal. The stories told about it to early travellers
led them to describe it as an animal with flesh and
blood, but fixed to one position, from which it never
moves ; hence the story of the now fabulous Barometz,
or Vegetable Lamb. This plant is of easy cultivation ;
if placed on soil slightly raised, a few years’ growth
will produce very good specimens of the ‘ Lamb.”
The rare Schizocena sinwata is an exception to the
general rule that characterizes Tree Ferns, the stem
being slender, not exceeding an inch in thickness,
attaining the height of three or four feet, and bearing
a fascicle of simple fronds. This remarkable Fern is a
native of Ceylon; it grows in shady places in woods
where a degree of coolness prevails. A few years ago
plants of it were received at Kew, and it was found
necessary to place them in a large Wardian Case, in
which they flourished.
Although the plants belonging to the order Muarat-
tiacee are ranked with Ferns, they nevertheless differ
CULTIVATION, 333
much in habit, chiefly as regards the nature of the roots,
which are thickened and fleshy, and generally pene-
trating deeply into firm soil. In their general character
they present much resemblance to the roots of
Cycadee, Cyclanthce,and other allied endogens; also in
venation the remarkable genus Stangeriaconnects them
through Dancea with Cycadee ; and their compound
fronds are represented in the same family through the
new genus Bowenia. The species of Marattia and
Angiopteris grow freely in a loamy soil, requiring
plenty of water. Some species of Angiopteris require
much space, for although the caudex does not rise
much above the ground, yet a plant at Kew produced
fronds that reached the height of twelve feet, spreading
outwards, forming a diameter of thirty-four feet. The
species of Danea and Kaulfussii- are, however, not
such free growers; they require a moist and high
temperature.
T have now gone through a few of the principal genera:
requiring special kinds of treatment ; the next consider-
ation is good management as regards temperature,
watering, airing, and keeping free from msects. Much
depends on the size and nature of the house ; the larger
it is, the less fluctuations of temperature take place,
especially in a house of the ordiuary construction of
glass roof and sides, whether lean-to or span. The
maintenance of a proper condition of the air in plant-
houses depends much on the nature of the interior
fittings. White or polished, reflecting and radiating
surfaces, should be avoided as much as possible, such
ag iron pillars, rafters, spandrels, polished slate or
smooth stone, iron or stone floors, &c. These kinds of
matcrial are often introduced in superabundance, for
334 FERNS: BRITISH AND FOREIGN.
the purpose of giving consequence and dignity to the
house, thus laying the foundation for conditions un-
favourable to plants. The shelves orbenches may consist
of dark-coloured porous stone or rough slate, with an
edging of smooth slate two inches deep, so as to form a
shallow trough, which should be ‘filled with dark-
coloured sand or fine-sifted coal-ashes, the whole
pressed down, so as to form a firm smooth surface, on
which the pots are to be arranged. By this means a
degree of moisture will be retained more congenial to
the plants than when standing on stone or slate. An
edging of about three inches of Selaginella hortensis or
S. denticulata gives a neat appearance, and if allowed
to run between the pots, the whole becomes more con-
genial than a naked surface. White sand or broken
quartz is frequently used, but it soon becomes dirty,
and has a harsh and dry appearance.
In the arrangement of the plants some degree of
order must be observed, so that every specimen plant
of a species should be seen to advantage, and if the
collection is extensive, and the house has different
aspects as regards light and shade, then it is desirable
to arrange the plants according to their habits and
requirements; thus, at Kew, the division Hremobrya
occupy the whole of the south side of the house; this
division being less sensitive to the effects of the sun
or deficiency of water than the more extensive division
Desmobrya, which, on account of their thin texture,
very quickly suffer from dryness or too much exposure
to the sun; therefore the north side of the house is
best for them. i
The natural direction of the fronds is the most
pleasing, whether uprigut, pendulous, or spreading.
CULTIVATION. 335
In the latter case, sometimes their own weight makes
them fall lower than is convenient; it therefore
becomes necessary to support them by inconspicuous
stakes. Care must, however, be taken not to raise the
fronds above their natural position, and to avoid making
them have a stiff formal appearance.
In “Theories of Horticulture,” it is said that
abundance of lght and a free circulation of air are
indispensable for the cultivation of plants under glass.
In my long experience, I have found, as a rule an
abundance of light and a too free circulation of air
prejudicial to the good cultivation of plants in general.
With regard to Ferns, they love a quiet, dull atmo-
sphere ; the light afforded by a roof glazed with the
ordinary transparent glass, admits fully double the
amouut of light necessary during eight months of the
‘year. Sufficient means are, however, necessary for
admitting air, in order to keep down the temperature
in summer; but care must be observed in admitting
it; if not, the increased ventilation in lowering the
temperature will cause a rapid dispersion of the essen-
tial moist atmosphere at a time when most required.
Shading must then be resorted to, this being the chief
purpose which it serves, besides protecting the plant’s
foliage from the too powerful rays of the sun. It may
i consist of canvas blinds on rollers; two moderately
thick ones are much better than one very thick, espe-
cially when Trichomanes and Hymenophyllum are
grown; one should be permanent during the height of
tlie summer, the other to roll over it on occasions of full
sunshine. Although canvas is here spoken of as best
for shading, and is extensively used, still it is trouble-
‘some, and in the end expensive, which makes it de-
336 FERNS: BRITISH AND FOREIGN.
sirable to resort to some other method; such as by
using several kinds of dulled or rough glass.
In former years, the fruit and plant-houses at Kew
were glazed with a very dark-green glass called
Stourbridge-green, and which was patronized by the
late Mr. Aiton. Fine crops of fruit were produced
under it, also the tropical plants in the Botanic
Garden flourished without the aid of canvus or shade °
of any kind. Not many years ago, solitary squares
of this glass might be seen in the roofs of the
old hot-hcuses, which strongly contrasted with the
modern clear glass. My experience with this glass led
me to recommend green glass for the Palm House,
which was adopted ; but the modern-made green tint
does not appear to be so fixed a colour as that of the
old Stourbridge-green.
The Palm House in the nursery of the late Messrs.
Loddiges, at Hackney (now things of the past), affords
another example of successful plant-growing without
the aid of canvas or other moveable shading material.
On the late Mr. George Loddiges being consulted
respecting the glazing of the Kew Palm House,
he was asked if they shaded theirs; his reply was,
“Oh, no; our thick rafters and sash-frames, with
sooty glass, just afford the amount of light necessary
for the plants.” Under this roof, in an atmosphere
of stillness and gloom, Palms, Ferns, Orchids, and
numerous other tropical plants, grew in the greatest
luxuriance, which, with the proverbial solitude of the
place, and when viewed from the elevated platform, -
gave the idea of a ravine in a tropical forest.
To judge by the above examples of Kew and Hack-
ney, it appears that fruits and plants were successfully
CULTIVATION. 8o7
grown under roofs very different in appearance from
the elegant clear glass roofs of the present day,
which, nevertheless, necessitates some kind of invention
for temporary shading in summer; but it is to be
hoped this will be superseded by further improvements.
Double glazing is now spoken of as answering the
twofold purpose of saving shading and fuel; but as
there appear to be different opinions on its merits,
further experience is therefore desirable before it can
be safely recommended.
With regard to watering, it is difficult to explain in
words, or laydown arule, what constitutes the extremes
of dryand wet soil ina flower-pot, feeling is perhaps the
best guide; if, on taking a pinch of soil between the
finger and thumb, the particles will not adhere, then it
is too dry ; if it adheres, and falls like a dry wafer, then
the plant is not in immediate want of water; but, on the
contrary, should the soil stick to the fingers like a wet
wafer, then the whole is in a bad condition, and should
the plant be weakly, it will get worse if allowed to
remain in the sodden soil; for the correctness of this test
care must be taken not to be deceived by the surface-
soil, for it may be either dry or wet, and the contrary
lower down. Another test of the state of soil is to
give the side of the pot a smart tap with a hard piece
of wood; should the sound be hollow, with a little
tone, then the ball is dry.; if, on the contrary, there
is only the dull sound of the tap, without any tone,
then the ball is moist and tight in the pot. To those
in the habit of watering the same plants for a length-
ened time, this test is of service, but it must not
altogether be depended upon. The quality of the
water has great effect on the health of the plants;
PA
338 FERNS : BRITISH AND FOREICN.
It should always be soft, or, if hard, should be aerated: -
‘in open tanks; rain-water is best, but if collected
from the roofs of hothouses, care must be taken that it
does not become impure by the decomposition of the
paint and putty, which is of frequent occurrence, and
in time it so impregnates the water as to cover the
foliage of the plants, when often syringed, with a thin
film, that gives them a dull look; care should also
be taken that the temperature of the water should not
be less than 50°, especially in winter. Syringing is
made an operation of great importance in guide-
books on cultivation, and is readily accepted by the
unskilled amateur’ as necessary, and being amusing,
is often carried out in the extreme, to the injury of
the plants. An amateur remarked not long ago,
“Now we have got such a nice lady’s syringe, it is
quite a delight to use it’? Another remarked, “I
every day regularly water, syringe, and sponge my pet
Adiantums, but with all my attention they are getting
of a brown colour, surely your plants (at Kew) must
have great attention to keep them so green.” The
lady syringer is told that many of the plants before her
have been in the same pots for several years, and that:
they get water when dry, and are never syringed
or sponged, unless for removing insects when they
appear,
From what has now been stated, an amateur’s
successful cultivation of Ferns simply depends on
proper soil and potting, careful watering, placing in a
quiet, moist atmosphere, keeping down temperature in
summer by shading, and to avoid currents of dry air,
and in winter to maintain the proper temperature,
according to the nature of the plants, by means of
CULTIVATION. 399
hot-water pipes. For the latter purpose, in small
houses, temporary stoves of gas and charcoal are used,
often to the utter destruction of the plants.
To assist in maintaining a moist atmosphere besides
the usual moist surfaces of the pots, it is desirable
to have water-troughs on the top of the hot-water
pipes, as also to place pans of water on or near the
pipes; also in hot weather, the paths and other
evaporating surfaces should be wetted every morning
and evening. An occasional syringing may be given,
which should be done when the out-door air is charged
with vapour; wetting plants when the air is dry
causes a sudden evaporation, and a reduction of
temperature on the surface of the frond is the con-
sequence. This is especially the case with broad,
smooth, glossy species, such as the genera Phymatodes,
Neottopteris, Asplenium, Diplazium, &c. ; occasionally
the whole of some fronds or part of a frond becomes
black. In order to raise vapour, a practice prevails
with many cultivators to throw water over the hot-water
pipes (or flues) ; there is no objection to this, pro-
viding the air out of doors is warm, but in the winter
season itis often done in the evening, in order to
counteract the dryness of the air, caused by the extra
heat of the pipes required at that season ; the house then
becomes filled with hot vapour, which coming in
contact with the glass of the roof condenses and falls
in a shower of cold drops on the plant; at the same
time the temperature of the house rapidly falls, thus
causing black fronds, spots in orchids, and such-like
complaints.
Ferns, like other plants, are liable to be more or
less infested with insects; the soft membraneous kinds
Z2°
340 FERNS : BRITISH AND FOREIGN.
being the most subject, while, on the contrary, the
smooth-fronded kinds, such as the whole of the
division of Hremobrya, Elaphoglossum, &c., are, it
may be said, quite exempt. The mealy bug, three
kinds of scale, and thrips, are the chief pests, and
should be carefully watched. The oblong brown
scale is generally the most common, and very quickly
overruns the under side of the frond, forming lines
along the midribs. Many kinds of fluid mix-
tures, powders, and other nostrums, have from time
to time been advertised for their destruction, but it
too often happens what kills insects also injures the
plant, and makes it look unsightly. Fumigation
with tobacco is the grand preventive against the
breeding of plant-insects; very slight fumigation
destroys the winged male insect of all the Coccos
family, and if frequently repeated, the whole in time
will become extinct. When the mealy bug gets a
head, it is readily got rid of by syringing; not so
the scale, it requires to be loosened with a soft
brush, or blunt-pointed peg, which, if the weather
permit, should be done out of doors, and the plant
afterwards syringed, so as to clear away all loose scale
aud eggs. Scale insecis are adherers and suckers
only ; not so the less conspicuous insect the thrip,
which is a small, slender, black, shining insect, with
large prominent eyes, and quick in its movements,
herdmg in groups, generally on the under side of
the frond, voraciously feeding on the cuticle, which
they soon destroy, permanent injury being often done
without any indication of their presence till too late;
they must, therefore, be carefully watched, and, when
seen, at once destroyed by pinching with the fingers,
CULTIVATION. 841
and syringing the plant. They are generally found
on species of Adiantum, Pellea, Platyloma, Doryo-
pteris, Hemionites cordifolia, and other genera having
fronds of a smooth, chestnut-brown colour. Within
the last twenty years a small, white-winged insect, like
a midge, has made its appearance (supposed to have
been first introduced with imported plants to Kew).
They congregate on the under side of the fronds, and, —
when the plant is moved, dart off like a flock of
white pigeons. At first it was supposed to be harm-
less; but such is not the case, as it has been found
to feed on the cuticle like thrips; but they are not so
easily caught. Repeated tobacco fumigation destroys
it.* Red spider seldom attack Ferns; when such is
the case, it is a sure sign that the air of the house
is too dry.
Few plant-houses are exempt from the well-known
cockroach, which when once introduced is one of the
greatest of pests. They increase and multiply most
rapidly, and are most voracious feeders, eating and
gnawing the fronds of Ferns, old and young, often
completely in one night spoiling fine plants; there-
fore war in all its forms must be waged against
them. They are night marauders, hiding and breed-
ing in crevices of masonry in dark places, generally
near the furnaces, or hottest parts of the heating-pipes,
and are even found lurking amongst the drainage, in
large pots, from which they issue at dusk to commence
their ravages. Many expedients are resorted to for
their destruction, such as attacking them in their day
* This insect has been described and figured in the
Gardeners’ Chronicle, of 1856, by Mr. Westwood, under the
name of Aleyrodes vaporariorum. :
342 FERNS: BRITISH AND FOREIGN.
abode, by the appliance of boiling-water, sulphur
fumes, or exploding gunpowder: when such agents
can be used, they deal with them quickly and whole-
sale. Various kinds of traps are used, which, with
poison, will, if daily attended to, completely extirpate
them. But it must be borne in mind that, although the
whole, old and young, may be got rid of in the course
of a fortnight by poison— the effect of which is
greatly increased by the living eating the poisoned
dead,—eggs are however left, which will soon pro-
duce a new generation that must not be allowed to
arrive at maturity.
Under the ordinary varying atmosphere of hot-
houses, insects seem not to be affected, for if their
extirpation is not attended to, they will be found in
more or less abundance all the year. Not so the
sooty mildew,* a fungus covering the upper surface
of the leaves of plants with a black, sooty coat, and
for their sudden appearance, like that of the grape
mildew, the potato disease, and other sporadic
plagues, no satisfactory causes have as yet been
assigned: The pest now under consideration may be
called one of these plagues ; in some years it is not
seen, while in another it soon overruns and quickly
covers Ferns, and other plants, in hothouses, The
broad-fronded species of Aspidiwm, Meniscium, Gonio-
pteris, Anyiopteris, &c., are very subject to its attacks.
Books on mycology name and describe these pests,
but not how to prevent them; and books on horti-
culture instruct how to get rid of them; the principle
of which seems to be dusting with sulphur, washing,
* Fumago foliorwm, Fries.
CULTIVATION, 343
and syringing, the latter being the only remedy for
the black mildew.
This concludes my observations on pot-cultivation,
and in order to save repetition, it must be understood
that the process as regards soil, watering, airing, &c.,
is equally applicable to natural cultivation, which I
now proceed to describe.
TI. Narorat Cunrivarion.
HE natural cultivation of Ferns consists in growing
7 them without the aid of garden-pots. This is
accomplished by placing them, as far as artificial
appliances will permit, under conditions and influences
analogous to that of their native wilds; for that
purpose uneven irregular surfaces are best adapted,
whether natural or artificially formed; and as Ferns
are generally called rock-plants, which many truly are,
therefore, in order to imitate rocks, the surface on
which they are grown is made irregular and covered
with rude stones, on or between which the Ferns are
planted; hence the term Rockery is applied to this
kind of cultivation.
For all species of Ferns, either hardy or tender,
this system of culture, which admits of their being
planted out, is far preferable, to whatever extent it
may be desirable to practise it, whether in the sheltered
nook, in the open air, or in stoves or greenhouses,
either partially or entirely devoted to it. For by ib is
the finest health imparted to the plants. The rockery
bank which once occupied the Temperate Fern House
at Kew, together with the fine fronds of tropical species
grown on rockery in the Palm House, afforded excelient
844 FERNS: BRITISH AND FOREIGN.
examples of the superiority of this mode of growing
plants over that of pot culture; by proper arrange-
ment from ordinary level positions, their delicate hue
and elegant form of outline may be seen to much
greater advantage, as the upper surfaces of their
fronds are generally presented to the eye more fully.
And where this system is largely carried out, if a
raised platform or footway be erected considerably
above them, the advantage is still greater. The great
beneficial influence this system has on the plant is
mainly consequent on the more uniformly moist at-
mosphere, so congenial to all Ferns, which can be
successfully maintained from the great extent of the
more natural evaporating surface of the soil, and ma-
terial, with which the rockery is composed ; and as they
are allowed to establish themselves at freedom in it,
there is a similar uniformity of temperature at the
root, as well as of moisture. As is well known, in
large masses of earthy and other solid material, its
temperature does not fluctuate with that of the house,
at least, as in the case with ordinary-sized pots of soil,
where the absorption and radiation of the heat takes
place so much more rapidly. It may further be men-
tioned, that, in a house entirely devoted to the cul-
tivation of Ferns under this system, its evaporating
surface does not wholly consist of the principal mass of -
material composing the rockwork ; there is the large
extent of wall-surface, which, if properly constructed
with rough and absorbent material, is continually
giving off its moisture, besides, from that reason,
affording an excellent opportunity for Ferns of more
or less scandent habit to assume their native character
in luxuriant profusion, by growing on it as on natural
CULTIVATION. 845
rocks. As regards the design of any structure in-
tended to be wholly or partially devoted to the culti-
vation of Ferns, it is of course a good deal a matter
of taste and convenience, as the system is available
in almost any kind of structure, large or small, but
varying in the degree of economy in its construction.
To this end, the first consideration should be to
construct in a manner that will insure the necessary
amount of shade and moisture and (when required)
heat, with as little auxiliary assistance as possible.
Therefore if a lean-to house be adopted, the aspect
should be west or north-west, with the back consisting
entirely of wall. Should a span-roofed structure, how-
ever, be preferred, which indeed for all purposes is
much the best (and which will be here treated. of as
a general example), its aspect should be north and
south, by which a shadier and consequently a moister
side of the house will be obtained, suitable for Ferns
of more or less delicate structure, such as the genera
Trichomanes and Hymenophyllum. A sunk house, in
the manner of a ditch or railway-cutting, is most
suitable, and will give ample height for the growth
of Tree Ferns, without the necessity of having high
side walls, and thus exposing a lessened portion of
the house’s surface to the action of climatic vicissitude;
consequently a warm moist atmosphere may be main-
tained with but little assistance from artificial heat.
As a source of heat the hot-water apparatus is to be
preferred, on account of its economic utility, and
where practicable the pipes should be hidden as much
as possible with the rockery material. Perkins’s sys-
tem of heating is perhaps the best for a house of this
kind; it consists of coils of small pipe, which can be
815 FERNS! BRITISH AND FOREIGN.
arranged in any part of the house, and are readily
detached and again fixed at another point when found
necessary. The first deposit for the rockery should
be rough and somewhat absorbent, such as irregular
brickbats, masses of stone, loosely disposed to admit
of a ready dissemination of the heat amongst its
parts, and then follow with the material proper.
Apertures for the escape of the heat should be con-
structed at frequent intervals, drain-pipes being very
snitable for this purpose, which, by the mass being kept
properly moist, will allow the heated air to circulate
in a congenial steamy condition throughout the house.
In the design and arrangement of the principal
mass of material composing the rockwork, it is, as in
the design of the house, a good deal a matter of taste
and convenience as regards size; whether in repre-
senting in miniature a rugged mountain side, or deep
glen, with its clear pool, reflecting the elegant forms
of the Ferns growing near it; or may be a miniature
valley, with undulating surfaces gradually rising on
either side, with projecting rocks. There are nu-
merous kinds of material more or less suitable for
constructing a rockery either indoors or out, and but
a few localities where some of these may not be easily
procured. The chief desideratum is to have.such that
is of porous consistence, and generally with rough
and ragged surfaces. It is useless to be particular in
the selection of various kinds of stone on account of
their fine quality, either in texture or colour ; for where
Ferns properly thrive all such will be speedily obscured
by their luxuriant growth; consequently any brick-
field affords very good material that would contribute
largely in the general structure, in the way of con-
CULTIVATION. 347
glomerated masses of brick, or bricks artificially
amassed with Portland cement to form miniature
rocks, rugged cones, rough pillars, or perpeudicular
surfaces. However, when procurable, the absorbent
androughly quarried soft stone should enter largely into
the composition of the rockery, as it is pre-eminently
useful, not only in contributing so largely to the
natural appearance of the whole, but, when the more
delicate members of the Fern family of creeping habit
are grown, it is invaluable in affording a moist surface
for their tender rhizomes to cling to. It is seldom
that wood of any kind is found of much use, as it is
so soon reduced by decomposition, and fungi produced
im contaminating profusion. The soil to be used in
the rockery-house should be selected and applied
according to the principle already given in pot-
culture. The style of the Fern-rockery having been
decided on and executed accordingly, the next step
will be to arrange the plants about it; and to carry this
out satisfactorily is not the least difficult part of the
whole undertaking, as it is not only essential to provide
positions for each individual plant, but such that
will prevent any injurious interference of its neighbour,
either by fohage or root.
Should the roof require support (and if not, should
it be consistent with elegance and regularity), pillars
composed of rough porous stone or brickwork may
be constructed, and, if admissible, united above as
archings, which, when clothed with the luxuriant
foliage of scandent and pendulous Ferns, intermixed
with the broad foliage of creeping Aroidew, growing
from their crevices and recesses, would greatly enhance
the beauty of the whole, and give a most accurate and
348 FERNS: BRITISH AND FOREIGN.
durable example of the manner in which many spccics
of Ferns and Aroidee clothe the trunks of trees in
tropical forests; also several species of epiphytal
woody plants may be introduced with good effect,
such as Tanecium parasiticum, Marcgravia umbellata,
Norantea coccinea, Hoyas, and other creepers not
‘subject to be infested with insects. The species of
Phymatodes, Pleopeltis, Davallia, and several allied
genera, as also Polybotrya and Stenochlena,will luxuriate
over the moist surfaces and form a compact mass; and,
if planted near the wall, direct their rhizomes upwards,
and the plants will quickly assume a most elegant
appearance. The grand and beautiful Schellolepis
verrucosa, and its allies, should be placed in elevated
positions, as their fronds are long and pendulous.
Elevated positions will also be found for the species of
Platycerium, the grand Drynaria Heraclea,and D. coro-
nans: the latter is remarkable in its thick rhizome,
progressing in a circular direction round trees, which,
with the erect, broad, rigid fronds, gives the idea of a
coronet or crown; this circular direction appears to
be normal. A fine plant at Kew, in the course of eight
years, grown in a shallow pot, formed a circle more
than a foot in diameter. For that portion of the
terrestrial group composed of stronger growing spe-
cies, such as Phegopterideee, Pieridew, Aspleniee, &c.,
for which firmer soil is necessary, the ordinary surfaces
of the rockery offer most suitable positions, arranged
there in a manner as their sizes indicate, so as to be
in harmony with those of smaller size around them.
It is perhaps essential to note that the Gymnogramma,
Cheilanthes, and their allies, require a place in the
Fernery, the lightest and best drained; therefore
CULTIVATION. 849
the higher part of the rockery is best suited for them.
In such a Fernery as this now before the mind’s eye,
convenient positions, in the form of dripping crypts
and recesses, may be provided for the delicately beau-
tiful and easily grown Trichomanes and Hymenophyllum,
without the aid of glass cases that are necessarily
used, as already explained, under pot-culture.
Where Tree Ferns would be grown, the proper
situation for them is the lowest part of the house, in
the valley, where the greatest head-room would be
obtained, and most constant shade and moisture about
their stems, which will encourage the essential growth
of the aerial out-grown roots. The path in the valley
may be straight or winding between the Tree Ferns,
the intervening spaces to consist of raised mounds or
lillocks, to be planted with the smaller kinds of Tree
Ferns and other large-fronded Ferns. The different
species of Selaginella should occupy the intervening
spaces between the plants, which will add greatly to
the natural appearance of the whole and assist in pro-
moting a genial atmosphere.
To whatever extent a house of this nature may be
carried for tropical Ferns, the same must be allowed
for the extra-tropical kinds, and being sunk, the
necessary amount of heating power required will be
just sufficient to keep out frost. Keeping the house
shut during the winter season, so as to maintain a
quiet, still atmosphere, greatly assists in repelling
frost, indeed for such houses ventilation is only
necessary to keep down high temperature during
summer; free admission is, however, beneficial when the
external air is still and moist, when even the tropical]
division may be freely left open during the nights of
350 FERNS: BRITISH AND FOREIGN.
summer, but taking care to prevent currents of dry air.
I have now stated the chief points to be observed
in forming and arranging an indoor natural Fernery,
the principles of which are capable of application to
houses of the ordinary construction and usual average
size, or to any extent that means will allow; even to
realize the grand idea of the celebrated Loudon, who,
more than thirty years ago, speaking in favour of span-
roofs, said, “There would be no difficulty in covering
ten acres of Kew Gardens with glass by a series of
span-roofs.” Although since then much has been
done at Kew, by the erection of lofty plant-houses,
still the area covered falls far short of ten acres; but
as natural cultivation is now patronized, and with
the improving age and desire for novelty, let us hope
that the time is not far distant when Kew may have
at least one acre converted into a straight or windine
Fern-valley covered with glass, the highest part not
to exceed thirty or thirty-five feet above the centre of
the valley, that being sufficient height for such Trec
Ferns as may perchance withstand the vicissitudes of
thirty years careful treatment.
In a house of this kind, furnished with means of
maintaining a temperature of 60 in the coldest weather,
situations in it would be found suitable to the good
growth of not only Fvrns and Aroidece, but also
Lromeliacee, Orchids, and other kinds of plants of an
epiphytal nature.
For an open-air Fernery a sheltered situation should
be chosen, and if possible within the influence of
shelter and shade, but not under trees ; a sloping bank,
a natural or artificial hollow, such as an old gravel-pit
or sunk fence, may be turned to good account. The
CULTIVATION, 851
number of species of exotic Ferns sufficiently hardy
to bear the cold of our winters is very limited, much
less than is generally believed, the chief of them
being natives of North America, which, with the
British species, amount to about 80 in all; therefore
the extent of space required for a simple collection of
‘hardy Ferns is small ; an area of 60 feet by 10 will be
ample, and which may be either on the side of a slope,
or between two slopes with a path in the centre and
open at one end only; or it may be a raised oblong
mound with a path all around it. If not confined to
a strictly scientific collection, and space to be had, an
interesting natural glen may be formed by introducing
other plants conspicuous for their large fohage.
It has been already stated, that although our small-
growing native and several exotic species, such as the
rock Asplenie, Woodsia, &c., endure the extremes of
heat and cold in their native localities, they neverthe-
less suffer by full exposure under artificial cultivation ;
it is therefore necessary to plant them on the rockery,
so that they can readily be protected in winter; an
inverted flower-pot, placed over each plant and covered
with leaves, or some loose material analogous to the
covering of snow of their native country, is a good
protection against being injured by excess of moisture
or severe cold. Where the rockery is to be carried
out on a large scale, then eaves and chasms may readily
be constructed, and with a natural or artificial head of
water they can be kept moist by the constant drop-
ping and spreading spray; thus making appropriate
positions for the natural growth of Adiantum Cappilus,
Hymenophyllec, &c. But when a full collection of the
many varieties of Scvlopendrium, Asplenium, Lomaria,
852 FERNS: BRITISH AND FORZIGN.
&c., are grown, then a pit, or frame with glazed sashes,
is the most convenient mode of keeping them. The
width of the pit should not exceed 5 feet, 3 feet or
more high at the back, to slope to about a foot at
front, its length regulated according to the number
of plants. They may be either grown in pots,
arranged to slope to the front, or a bank may be
formed with small rockery stones, neatly arranged
with the plants between them, and if the mason work
for the frame or pit is contrived to be hidden, the
whole will then be in character with the general
rockery. By this an interesting little bank may be
formed, and by proper arrangement of the plants the
variety lover can at a glance see the difference that
marks one favourite from another. The glass pro-
tection will stimulate the plants into early growth, and
finer fronds will be made, but care must be taken not
to expose the young fronds to the harsh drying winds
of spring, or late frosts, which often do great injury
to out of door Ferns; by midsummer, the glass sashes
may be entirely removed, due attention being paid to
watering and shading.
If the soil and subsoil of the situation chosen for a
hardy Fernery is stiff and retentive of moisture, and
the neighbouring trees, walls, &c., are clothed with
Musct and Iichens, it is favourable for Ferns without
the aid of much rockery; but if the soil is light and
sandy, then rockery is most essential for retaining
moisture, not only for Ferns, but also all kinds of
small herbaceous plants. Formerly there were great
masses of rockery in the Botanic Garden at Kew, but
modern iuste has swept the whole away, which, with
the removal of trees, walls, and old shrubberies, has
CULTIVATION. 853
caused the garden to become drier, as is manifested
by the disappearance of above nearly forty species of
Mosses, Hepatice, and Lichens; the effects of this, with
other causes, have been hurtful to the hardy Fern col-
lections. In forming a new rockery, after the plants
have become once established, as few changes should
take place as possible, either with the plants or stones
on which Musci should be encouraged to grow; and
in order to give a decorative effect, Sedwms, Semper-
vivums, Sazifrages, &c., may be placed on the pro-
jections and shelvings of the stones, the whole bordered
by a line of spring-flowering plants, such as Iberis
sempervirens, Alyssum sawatile, Arabis alpina, and
‘other plants of that nature.
It may be also mentioned that the vicinity of water
is a favourable position for a Fernery; where such
does not already exist, an artificial piece of water
should be made in conjunction with the rockery,
varying in size from a small basin to any extent: it
may be circular, oblong, straight, or winding, with
irregular projections. In ordinary cases its width need
not exceed ten to twelve feet; its margin should con-
sist of a bog or border, not less than three feet wide,
having a slope to the edge of the water. This border
will afford the means of growing a great many curious
sub-aquatic and bog plants, and the projections will
be excellent positions for the different species of
Osmunda ; water-lilies occupying the centre.
The want of such a rockery and aquarium has been-
much felt at Kew; for with all the great cost for
lakes, ponds, and water-works, yet no arrangement
has been made for growing a scientific collection of
bog and aquatic plants, either hardy or tropical.
24
854 -FERNS: BRITISH AND FOREIGN.
TV. Cuttrvarion or Ferns 1x Warp’s Caszs.
T is now thirty years since I was invited by Dr.
Ward to visit him at his house in Wellclose
Square, for the purpose of seeing plants growing
in cases and glass jars, so closed as to be considered
air-tight. Knowing, as I did, the common practice
of growing plants under hand and bell-glasses, I
vherefore could not appreciate what I had gone to see
until I was made aware that the plant-loving residents
of such smoky and soot-falling districts of the metro-
polis, as that of Wellclose Square, could grow rare
and delicate plants equal to those at Kew. An
account of this: method of growing plants appeared in
the Companion to the Botanical Magazine for 1836, and
in April, 1838, the celebrated philosopher Mr. Faraday
delivered a lecture at the Royal Institution on the
subject, which may be considered as the advent and
introduction of Wardian cases, under which a large
portion, and decidedly the most beautiful of the
Fern family, are now successfully cultivated in the
sitting-rooms of the town-confined lovers of natural
objects. In 1842 Dr. Ward published a small work
on the subject, giving a history and details of manage-
ment, which renders it unnecessary for me to say more
regarding the early history of Ward’s cases. The
principle on which the system is founded, consists
simply in shutting up air in glass cases, in such a
manner that it is not readily influenced by changes
of the external atmosphere. The case also contains
several inches depth of moist earth, that gives off
CULTIVATION. 355
moisture to the absorbent, undisturbed air above it,
which becomes more or less saturated, as dependent
upon changes of temperature, and thus becomes
proper for the growth of plants.
The case may be constructed of any shape or size,
according to taste or means; it can be square or
round, an octagon or hexagon; the roof may be a dome,
span, or sloping, but by all means avoid a flat one;
and be the shape whatever it may, the design should
inevery point be neat, and not of such an ornamental
description as to be more attractive than the plants,
A very good, interesting collection may be grown
in one 3 ft. long by 14 ft. in width, and 24 fect high;
it will have the best appearance if the sides are con-
structed with single squares of glass; but if divided,
it must not be into more than three, as many
divisions spoil the effect; the ends should be made
to open, to enable any dressing or removing of old
fronds to be done conveniently, and a small piece of
perforated zinc should be inserted in the apex of the
roof at each end, which will assist in preventing the
almost universal complaint, that the plants cannot be
seen for condensed water on the inside of the glass;
this is caused by the variation of temperature. If
the case stands in the sun or becomes warm inside
during the day, and retains the heat, whilst the
temperature of the room may fall considerably during
the night, the cold air, acting upon the glass, con-
denses: the warmer vapour inside and obscures the
plants. By the introduction of the above-named
remedy the temperature is more equally balanced,
and the plants are always to be clearly seen. Should
the air outside become very hot and dry, it will be
242
306 FERNS: PRITISH AND FOREIGN.
advisable to close the ventilators for a short time
during the day. The glass case should be entirely in-
dependent of the soil-box, but to fit the inside, resting
in a groove or rabbet. The box should be about 6 in.
deep, and may be made of zinc, brass, or wood lined
with gutta-percha or zinc; but metal of any kind in
contact with the earth or air in which plants are
grown is not genial to either their roots or foliage.
I have always found the plants succeed best in a
neatly-made wooden box lined with pitch, having
a small tap or cock in one corner of the bottom, for
letting away any excess of water; but this will not
be necessary if proper attention is observed in
supplying the plants with a sufficient amount of
water at one time, which, in consequence of little or
no evaporation taking place, will be seldom required.
This knowledge can only be gained by practice ;
many amateurs’ failures with Ward’s cases being
caused mostly by giving too copious waterings at
certain stated intervals.
The height of the stand must be regulated accord-
ing to whether the plants are to be viewed in a
sitting or standing position; for the former the
ordinary height of a table is a sufficient guide, and
for the latter afew inches higher, so that the plants
can be seen through the side glass rather than from
the top. In preparing the box for the plants, about
one inch of its depth should be filled with sand or
other drainage material, such as is already explained
in pot-culture—but in Ward’s cases this is only
necessary as a precaution against an over-supply of
water,—the rest filled with soil, which should be good
fibry peat and silver sand, intermixed with pieces of
ba
CULTIVATION. 857
sandstone broken small, or (if peat cannot be pro-
cured) good light loam. Having proceeded so far,
two systems of planting the case present themselves.
‘The first plan is to plant in the soil, which should be
raised in the centre; the number of plants will
depend on the size of the box, and care must be
taken not to plant them in too crowded a manner,
the distance apart depending on the size and nature
of the plants, but in no case should they be closer
than six inches. The second method is to have the
- plants established in 4 to 6-inch pots, plunging them
in the soil sufficiently deep to hide the rims. of the
pots. The last system has one decided advantage,
namely, should a plant die or does not succeed, it can
be removed and replaced without disturbing its
neighbour. It must be understood that the above
mode of planting relates to plants with erect verna-
tion only, but those with creeping rhizomes, that form
cespitose tufts, require to be planted each on a
separately raised hillock, of which, if the case is large,
there may be a series, and for the creeping Hymeno-
phylla lumps of porous stone are very suitable; fine
patches may be obtained in this way, not only of
Hymenophylla, but also of the smaller species of
Niphobolus, Anapeltis, the beautiful Humata pinnati-
fida, Davallia pentaphylla, and others of like habit.
After the Ferns are planted, some small-growing
Selaginella, such as S. apus, may be pricked in over
the surface of the soil, in pieces a few inches apart,
-which will soon cover it, and give to the whole a neat
and finished appearance, and also materially assist
in maintaining the proper state of moisture in the air
of the case and about the plants. The planting bein 2
358 FERNS: BRITISH AND FOREIGN.
now completed, the soil must be brought to a uni-
form state of moisture; this is best accomplished by
two or three moderate waterings at intervals of half
an hour, and when thoroughly drained, the case may
be closed and placed in position. To make cases,
ornamental pieces of marble, shells, &c., are often
introduced ; but this should not be tolerated, as they:
do not harmonize with the occupants of such a
structure.
For Hymenophylla it is necessary they should be
sprinkled overhead occasionally ; to enable this to be
done, the tops of all small cases should be moveable,
and in large ones a hinged pane is required. A
sponge, or small thumb-pots filled with water and
placed out of sight, will greatly assist in keeping a
moist atmosphere, which is so essential for the health
of these filmy-leaved plants. If the cases stand in a
room where a fire is regularly kept in the winter, a
great many tropical species may be grown, and in
situations where they stand exposed, such as balcony
windows, halls, &c., care must be taken, as winter
approaches, that the soil does not become frozen,
such being very detrimental to even the hardiest
Ferns. “Hot bricks and bottles filled with hot
water have been resorted to as a preventive against
frost, as also for maintaining a proper temperature
for tropical species; but unless the bottom of the
case is constructed for that purpose, and arrangements
made for this mode of heating, to be strictly and
regularly attended to, it had better be dispensed with.
Should the case be exposed to the direct rays of the
sun during the summer, it must be shaded, and
care taken that the temperature inside does not
CULTIVATION.
become too high; it should not exceed 70°.
859
This
will be much assisted by placing the shading material
at some distance from the case.
Subjoined is a list of
species suitable for cases of ordinary dimensions ; it
must, however, be understood that a Ward’s case may
consist of a bell-glass, containing a ‘single plant, up
to a shut-in window, or area covered with glass, where
larger-fronded species may be grown.
Humata heteropliylla.
Davallia bullata.
pentaphylla.
Canariensis.
Leucostegia hirsuta.
— cherophylla.
pulchella.
Polypodium pectinatum.
Schkuhrii.
Lepicystis sepulta.
squamata.
rhagadiolepis.
Goniophlebium appendicu-
latum.
Lopholepis piloselloides.
Anapeltis stigmatica.
Pleopeltis nuda.
Niphopsis angustatus.
Drymoglossum piloselloides.
Hymenolepis spicata.
Phymatodes longipes.
glauca. .
Pleuridium juglandifolium.
venustum.
Selliguea caudiformis.
Niphobolus Lingua.
Drynaria propinqua.
Elaphoglossum piloselloides.
vestitum.
Hymenodium crinitum,
Rhipidopteris peltata.
Microstaphyla bifurcata.
Psomiocarpa apiifolia.
Stenosemia aurita.
Gymuopteris quercifolia.
Platycerium alcicorne.
Xiphopteris serrulata.
Gymnogramma tomentosa.
Calomelanos,
—— Martensii.
chrysophylla.
Peruviana, var. argyro-
phylla.
pulchella.
Llavea cordifolia,
Hemionitis palmata.
cordifolia. 7
Dictyoxiphium Panamense,
Ceratopteris thalictroides.
Meniscium simplex.
Goniopteris asplenioides.
Nephrodium molle.
Cyrtomium faleatum,
Fadyenia prolifera.
Aspidium Pica.
Hypoderris Brownii.
Trichiocarpa Muorii.
Polystichum mucronatum,
Lonchitis.
—— triangulum.
anomalum,
denticulatum,
360
Lastrea elegans.
concinna.
—— strigosa.
— vestita.
— podophylla.
—— erythrosorum,
—— Mexicana.
—— emula.
glabella.
— Shepherdi.
—— hirta.
sancta.
hispida.
deparioides.
Woodsia hyperborea.
polystichoides.
mollis.
Arthropteris albo-punctata.
Nephrolepis pectinata.
Notholena brachypus.
trichomanoides.
Marantee.
sinuata.
sulphurea.
Myriopteris lendigera.
myriophylla.
elegans.
tomentosa.
vestita.
hirta.
Cheilanthes viscosa.
fragrans.
tenuifolia.
microphylla.
— Alabamensis.
—-wmultiida. . -
—— argentea.
—— farinosa.
-—-— pulveracea.
capensis.
radiata.
pedata.
FERNS : BRITISH
AND FOREIGN.
Cincinalis nivea.
flavens.
pulchella.
Pcllea geraniifvlia.
——— intramarginalis.
hastata,
atropurpurea,
Calomelanos.
ternifolia.
eordata.
Platyloma Brownii.
Adiantum reniforme.
caudatum,
lucidum.
macrophyllum.
— villosum. °
pulverulentum,
‘rigidum.
fovearum.
curvatum.
affine.
formosum.
hispidulum,
setulosum.
— fulvum.
tenerum,
sulphureum.
—— Chilense.
Capillus.
AXthiopicum.
Onychium Japonicum.
auratum.
Pteris Cretica,
lineata.
longifolia.
-—— crenata.
heterophylla.
semipinnata.
argyrea.
tricolor.
scaberula.
Litobrochia denticulata.
var.
“uar.
albo-
Litobrochia leptophylla.
CULTIVATION, 861
Asplenium viviparium,
Doryopteris sagittifolia, viride.
pedata. fontanum.
collina. flabellifolinm.
Blechnum Lanceola. —— pinnatifidum.
cognatum. —— macilentum.
occidentale. ——- fragrans.
longifolium. Diplazium Zeylanicum.
campylotis. Antigramma rhizophylla.
Doodia blechnoides. Odontosoria tenuifolia.
caudata. Microlepia cristata.
Lomaria Patersoni. Gleichenia dicarpa.
lanceolata. rupestris.
L’Herminieri. alpina.
Asplenium Hemionitis. speluncer.
alternans,
— Belangeri.
Hymenophyllum species.
formosum. Trichomanes species.
Brasiliense. Féea spicata.
tenellum. Lygodium palmatum.
erectum. Anemia tomentosa.
marinum. fulva.
firmum. adiantifolia.
obtusatum. Anemidictyon Phyllitidis.
brachypteron. Mohria thurifraga.
Todea hymenophylloides.
V. Propagation or Ferns.
YFVHERE is considerable variety in the way in which
_ Ferns reproduce themselves, and we only aid
nature when we attempt to increase any particular.
species; such is usually effected, and certainly most
numerously, by sowing their spores or seeds, as com-
monly called, which are contained in cases, as already
explained at page 51; they consist of atomic particles,
which, under favourable conditions as regards light,
362 FERNS: BRITISH AND FOREIGN.
can be seen discharging from the spore-cases like
puffs of smoke; therefore Fern seed is said to be in-
visible, and to have the power of rendering persons
invisible. “I have the receipt of Fern seed and walk
invisible.”* Be that as it may, each particle is by the
aid of the microscope seen to have definite forms vary-
ing in different genera, being globose, oval, or angu-
lar, smooth, ‘plain, striated, or echinate, presenting
very beautiful microscopic objects. Although these
spores (see plate I.) are so small, they nevertheless are
endowed with an extraordinary power of, retaining
their vitality ; and being easily wafted by currents
of air, readily account for the wide geographical
range of many species of Ferns. When naturally or
artificially placed under favourable conditions, the
spore vegetates by expanding in the form of a simple
oblong cell, from which other cells are successively
produced, ultimately forming a thin green membrane,
called the Prothallium, which lies nearly flat, and by
the aid of fine spongioles attaches itself to the surface,
and when arrived at full size it is of a reniform
cordate shape, or sometimes bilobed, or obcordate,
varying from about { to } inch in diameter, having
much the appearance of a small foliaceous lichen. But
before proceeding to describe the method of sowing
and rearing, it is important that the seed sower should
be made acquainted with the remarkable discovery
made, in 1848, by Count Leszezyc Suminski, that Ferns...
possessed urgans analogous to stamens and pistils of
flowering plants, and that these were produced on the
Prothallium, In the progress of growth peculiar cells
* Shakespeare.
CULTIVATION. 363
are formed on its under. surface, of two kinds, one
called Antheridia, and the other Archegonia ; the first
(of which there are generally between 30 and 40) con-
taining round cells, called sperm cells, which contain
vermicular spermatozoids that issue from the cells
when arrived at maturity. The cells of the Archegonia
differ considerably in their structure from the Anthe-
ridia; they contain an embryonal sac, which at a certain
period protrudes from the cell, and with which the
moving spermatozoids are said to come in contact, and
thus produce fertilization ; soon after this the embryo
sac is changed, a bud is formed, and ultimately a young
Fern Plant is gradually developed, and in time the
Prothallium decays. This singular phenomenon excited
considerable interest, and has been carefully inves-
tigated by the late Professor Henfrey and other
eminent microscopists.*
Although the above is only an outline of the prin-
cipal points of this curious subject, yet it will be suffi-
cient to draw the attention of Fern seed sowers to the
subject, and materially assist them in understand-
ing, and arriving at the cause of the many anomalies
in the good and ill success attending the raising of
Ferns from seed. Much has, however, yet to be
investigated before any rule can be arrived at ; such as
why the Desmobrya division does, asa general rule, pro-
duce plants from seed in the greatest abundance, even
to some species becoming the weeds of the hothouse ;
while, on the other hand, those of the division Hremo-
brya are comparatively few, and may be considered
the exception and barrenness of this rule. The differ-
* Henfrey, in Linnean Transactions, vol. xxi, 1853.
864 FERNS: BRITISH AND FORKIGN.
ence in the fertility of the spores of these two divi-
sions is veryremarkable; this is,it must be remembered,
as found with the plants under cultivation, and as with
many true Ferns we find, from experience, the spores
of the latter seldom germinate in less than eighteen
months after having beensown. On the contrary, in
the Desmobrya group, many require but two or three
weeks, and occasionally certain species of Gymmno-
gramma and Cheilanthes will germinate in as many
days. From the irregularity, however, in this respect
it is in a great measure governed by the state of, and
conditions which surround, the seed; from repeated
trials with spores of Brainea insignis, Prothallia were
produced in forty-eight hours. Consequent on this
irregularity, ib is impossible to state what time of the
year is most suitable for sowing to ensure success,
especially for imported spores, which should be sown
as soon as received, provided a proper condition can
be given them. Therefore extra attention must be
paid when it unfortunately occurs that the young
Ferns are in their Prothalliwm stage during the winter
season, to keep them from damping and the growth of
conferva and musci. When it is desirable to increase any
particular species by spores, some precaution is neces-
sary to secure them in a proper ripe condition, and
without spores of other species growing near being
mixed with them. The fructified frond should be taken
as soon as the spore-cases on the plant begin to open,
~and placed between sheets of paper in a moderately
dry place for a few days, at the end of which time there
will be escaped spores in abundance. These should
be immediately sown in pots prepared in the following
manner ;—four to six-inch pots are generally the most
CULTIVATION. 865
convenient size; they should be half filled with good
drainage material; with the smallest particles at top,
when another fourth of depth should be occupied by
fine soil, half sand and peat, one quarter loam, with a
sprinkling of finely-broken sandstone or soft brick
slightly pressed down on the top; it should -then be
watered and time allowed for the whole to become
uniformly moist ; then the spores to be very thinly dis-
tributed over it, the whole covered with a bell-glass
or a piece of glass same size as pot, to be placed on
its rim, allowing a space of about one inch between it
and the surface of the soil. In order to keep the whole
moderately and constantly moist, the pots should be
placed in pans of water of half an inch depth, care being
taken not to allow the soil to become over saturated ;
and whenever any copious condensation takes place
on the glass, it should be carefully sponged off. As
the spores germinate, and the Prothallia become
crowded, so as to touch each other, they should be im-
mediately thinned, and if it is desirable to save the
thinnings they can be removed in little clumps on the
particles of brick or sandstone to other pots prepared
as for spores. It is not, however, with all our care in
sowing different species in separate pots, that the
species sown come up in the pot in which it was sown:
plants of it may be found in other pots, or in different
parts of the house on moist surfaces. This is easily ac-
counted for, as the least motion of the air carries away
the spores while in the action of sowing, and indeed
all superfluity of spores may be with profit distributed
over the whole huuse, the moist walls often affording
abundance of young plants. It also often happens that
a good crop of Pteris aquilina is the result, its spores
366 FERNS: BRITISH AND FOREIGN.
being no doubt latent in the soil used for filling the
pots. To avoid this it is advisable to bake the soil
before using, and to bring it again to its proper moist
state by the use of water that has been boiled.
It is said that spores retain their vitality for a
number of years; in my experience I have no direct
proof of this; but several remarkable. instances of
plants making their appearance without the spores of
the species having been sown, or even an Herbarium
specimen having been seen in this country. In the
instance of Lomaria Patersoni, a species originally
discovered in Tasmania, which spontaneously made
its appearance at Kew in 1830, only one specimen
was at that time said to be in the possession of Mr.
Brown, at the British Museum, which I never saw; and
Allan Cunningham informed me that he never found
the plant, and was very much surprised when he saw
it growing at Kew. This in time gave specimens to
many Herbaria, and living plants to botanic gardens.
A similar instance was that of Doodia blechnoides,
which made its appearance at Kew in 1835. Other
instances might be quoted, such as the appearance-of
Asplenium stipitatum,of which two plants spontaneously
made their appearance about twenty years ago, and I
at first supposed they had originated from the spores
from a specimen in my Herbarium of a Luzon plant
named by me Neottopteris stipitata; but in time it
became evident that the two plants were quite distinct
from it, and, like the Zomaria and Doodia, I had never
seen native specimens. By what means the spores that
produced these plants came to Kew it isimpossible to
say. In 1829 I found a plant of Ceterach officinarum
growing in a crevice of masonry on one of the
CULTIVATION, 867
towers of the New Palace at Kew (since taken down).
As this Fern is not found wild near London, it would
be useless to speculate where this solitary spore came
from ; it seemed however to have found a proper nidus
in the crevice, enabling it to germinate and resist all
untoward influences, to pass through the Prothallium
state and become a plant.
Polypodium vulgare and Asplenium Ruta-muraria
may be considered our domestic Ferns ; for many years
a plant of Polypodiwm vulgare grew on the brick wall
separating Hyde Park from Kensington Gardens, and
there it remained till the wall was taken down. These
few instances of isolated appearances of Ferns readily
explain the wide geographical distribution of some
species over the surface of the earth.
It has been shown that the spores of many species
germinate quickly and abundantly, and become fully
developed Prothallia, yet it often happens that no
plant: bud is formed, and in time the Prothalli«
decay; the cause of this has always been supposed to
be undue moisture or some atmospheric action not
sensible to us, as this has always occurred in certain
species of special interest, such as Brainea insignis,
the spores of which, as already stated, germinate
readily ; yet we have not succeeded in obtaining young
plants, not even one Prothallium being seen to makea
plant bud. Without special microscopical examination
of the Prothallia we can only speculate on the proba-
bility, that, as in flowering plants, the whole of the
spores of some Ferns (such as Braineu) are unisexual or
may even be entirely destitute of both Antheridia and
Archegoma, in either case deficient of the elements
necessary for the production of a plant bud.
£68 FERNS: BRITISH AND FOREIGN.
I have at page 65 mentioned that a few intermediate
forms of the genus Cy ymnogram: ma had been raised from
spores, which are considered sports by some and by
others hybrids ; the latter can only be admitted on the
supposition of two Prothallia of two different species
growing so contiguous to each other that the sperma-
tozoids of one Prothallium have the power of passing
and fertilizing the Archegonia of the other, and thus
produce a hybrid, as in flowering plants. Another
point of some practical importance is, that in
general only a single plant bud is formed on each
Prothallium, This may be supposed to be owing to
the vital function of the Prothalliwm not being able to
support more, in that respect analogous to only one
ovulum being fertilized in ovaries of many flowering
plants. Admitting that, then, how are we to explain that
on removing the plant bud a new bud is formed; and
even as many as eight to ten have been obtained from
Prothallia of Hymenodium crinitum, each of which, by
proper care, becomes aplant. Then again experiments
have shown, that by dividing the Prothallium from the
base upwards with a sharp instrument into two or
even four parts, each part produces a plant bud.
Seeing this, it is reasonable to infer that Prothallia
have the power of producing plant buds, analogous to
leaves of Begonias and other plants ; but whether such
is the case, or each bud is the result of the action
of spermatozoids on latent Archegonia, is not known.
Then again we have the remarkable instance of the
great profusion of plants produced by the spores of
all farinose Ferns, such as species of Gymnogramma,
Cheilanthes, Nothvlcena, Cincinalis, §c., and also of the
smooth ebenous genera Pellea, Llatyloma, Doryo-
CULTIVATION. 369
pteris, and Adiantum, while comparatively only a few
of the smooth-fronded species of the division Hreme-
brya produce plants from spores. This subject yet
requires much experimental investigation before
satisfactory reasons can be assigned for what is here
stated.
The majority of Ferns that do not increase by
spores, often, however, readily do so by other means,
such as by offsets, and viviparous buds, or bulbils
produced on the upper surface, on the apex of the
fronds, or in the axils of the segments, which, when
placed under favourable circumstances, become plants.
Ferns of czespitose vernation will occasionally produce
buds or crowns laterally on the old caudex, which
may be readily separated for propagation with a
sharp knife; when the vernation consists of a creep-
ing rhizomé, such may be cut in pieces of whatever
length desirable, with a bud or growing point in each
piece, and, as with the separated lateral crowns,
should be placed in as small a pot as convenient, with
soil suitable to their kind (selected according to the
tule already given for establishing plants), and the
whole subjected to an extra close atmosphere till
thoroughly established. Up to the present time at-
tempts to propagate Ferns by separated portions de-
void of any previously joined bud have proved fruitless ;
although by some a solitary instance in Scolopen-
driwm vulgare is considered sufficient evidence to the
contrary, as portions of the base of its fronds, if
separated, inserted in soil, and kept close and moist
by the aid of a bell-glass, will readily strike; so also
with some of the abnormal forms of the same genus, if
portions of the margins of their fronds are treated in
28
370° FERNS : BRITISH AND FOREIGN.
the same manner. Upon close examination, however, .
previously formed embryo buds are observable and,
considering, too, its close affinity with the Aspleniums, ,
the most proliferous. (in the formation of bulbils) of.
all Ferns, this cannot be wondered at. To propagate
Ferns by the buds produced on their foliage is most.
easy. As soon as the bulbil plants have attained a size
to be handled conveniently, they should be carefully-
taken off and pricked out in pots filled with mode-
rately fine soil, and kept covered with a bell-glass till
thoroughly rooted, when they. may be potted off in
single pots, as required.
Species with long, slender, hard sarmentum, such.
as Gleichenia, do not root readily when separated ; in-
deed, large plants have been entirely destroyed by too »
free division of their sarmentum; to prevent this, layers
are resorted to, which is accomplished by fixing pro- .
longing sarmentums over small pots filled with soil,
which, when well rooted, can be. separated with safety,
and without injuring the specimen plant. Again, in
regard to the division Mremobrya, they are not only
remarkable in the sterility of their spores, but also in|
not producing viviparous buds; however, the readiness -
with which small portions of their rhizomes form
plants, and the already described tenacity of life,seem
to make them independent or to render less need of:
perfect spores or bulbils.
In concluding this treatise, I deem it necessary
to explain, that, in consequence of the woodcuts
occupying more space than was calculated for, and
in order to keep the book within a limited size,
it has become necessary to considerably reduce the.
original manuscript on Cultivation. It is, however,
CULTIVATION, $71
hoped that what is now given in the preceding pages
will be sufficient to show the nature of Ferns, and the
methods adopted for propagating and préserving
them in the collections of this country.
NOTE.
In the preceding article on Cultivation it has been shown
that all Ferns are capable of being cultivated in this country
under one of three conditions as regards temperature, deter-
mined by the nature of the native climate of the different
species, which, as regards the species enumerated in the
preceding catalogue, I classify as follows :—
1st. Hardy, in the open air.
North and Central Europe, including Great Britain
and Ireland, North America, North Asia.
2nd. Temperate House.
South Europe, Madeira, South Africa, North
India, China, Japan, Australia, South of the Tropics,
Tasmania, Norfolk Island, New Zealand, Chili,
Mexico.
A few species of those countries are hardy,—such are marked
.H. after the name of the country; some others improve by a
higher temperature,—such are marked -Tr. after the name of
the country.
8rd. Tropical House.
West Tropical Africa, including St. Helena and
Ascension, Mauritius, Ceylon, India, Malacca, the
2B2
3872 FERNS : BRITISH AND FOREIGN. F
Malayan Archipelago, including Penang, Singapore,
Java, and the Philippines, Borneo, the Polynesian
and other Islands of the Pacific within or near the
Tropics, Sandwich Islands, Tropical America, inclu-
ding Venezuela, New Granada, Panama, Peru,
Guiana, Brazil, West Indies.
Some species from high altitudes within these countries will
thrive in the Temperate House,—such are marked T., after
the name of the country.
373
A LIST OF AUTHORS AND BOOKS
QUOTED IN THIS WORK.
N.B.— When no special book is quoted after an author’s name,
his writings on feens are to be found in botanical and other scien-
tific journals of their time, and which are too numerous to notice
in this work,—Those with an asterisk are living authors.
*Agardh. J. C. Agardh, Professor of Botany at Stockholm ; author
of a “Monograph on the genus Pteris.”
At. W.&W.T. Aiton, father and son, Directors of the Royal
Botanic Gardens, Kew, from 1760 to 1841 ; authors of 1st and
Qnd edition of “ Hortus Kewensis,” 1793 and 1813.
A. Rich. A. Richard, a French botanist ; “ Voyage de l’Astrolabe
(Botanique).”
*Arn. Walker Arnott, Professor of Botany at the University,
Glasgow (see Hook. et Arn.).
*Bab. C. 0. Babington, Professor of Botany at the University,
Cambridge ; an eminent British botanist.
*Backhouse. James Backhouse & Son, nurserymen, York;
importers and cultivators of Ferns.
Bauer. Francis Bauer, a celebrated botanical painter (see Hook.
et Bauer).
Beauv. Palisot de Beauvois, a French botanist ; author of “Flore
d@Oware ;” figures and descriptions of plants in the Bight of
Benin, 1810. :
Bernh. J. J. Bernhardi, Professor of Botany at Erfurt.
Blume. ©. L. Blume, Director of the Botanic Garden, Batavia ;
“Enum. Plant. Jav.,” 1830; “FL Jav.,” figures and descrip-
tions of the plants of Java. ,
Bolt. ‘J. Bolton, an English botanist ; “ Fil. Brit.” figures of
British Ferns, 1790.
874 FERNS: BRITISH AND FOREIGN.
Bory, J. B.@. Bory de St. Vincent, a French traveller and botanist.
*Brack. W. D. Brackenridge, Botanist to the United States’
exploring expedition ; Descriptions and figures of the Ferns of
the expedition.
Braun. A. Braun, Professor of Botany, Berlin ; “ Monograph on
Selaginella.”
Br. B. Braun, the most celebrated of botanists ; “ Prodromus
Flore Nove Hollandie,” 1810; “Observations on Ferns in
Wallich’s Plante Asiaticee Rariores ;” Horsfield’s “ Plante
Jave.”
Br. P. Browne, author of a “ History of Jamaica,” 1756.
Brongn. A. Brongniart, a French: botanist.
Burm. J. Burmann, a Dutch botanist, and writer on plants of
India, Ceylon, &e.
Carm. Captain D. Carmichael, a Scotch cryptogamic botanist.
Cav. A.J. Cavanilles, Professor of Botany, Madrid.
*Colenso. Rev. W. Colenso, a New Zealand botanist.
Col. A. Colla, a collector and namer of Chilian ferns.
Cunn. A. Cunningham, a celebrated botanist and traveller in
Brazil and New South Wales from 1815 to.1830.
Dec. Aug. Decandolle, a celebrated French systematic botanist.
Desf. M. Desfontaines, a French botanist and traveller in Barbary ;
“Flora Atlantica.”
Desv. N. A. Desvauc, an eminent French botanist; author of
several papers on Ferns, from 1808 to 1814.
De Vries. G. H. De Vriese, a Professor of Botany, Leyden ;
“Monograph on the genus Angiopteris.”
Dick. J. Dickson, an English cryptogamic botanist.
Don. D. Don, Professor of Botany, King’s College, London ;
“Prodromus Flore Nepalensis.” — -
Dry. Jonas Dryander, librarian to Sir Joseph Banks ; a writer on
Ferns in the “ Linnzan Transactions.”
*Fiat. D.C. Eaton, an American botanist.
Ehrhart. F. Ehrhart, a German botanist.
Endl. Endlicher, Professor of ‘Botany, Vienna; “ Prodromus
Flore ae
Eng. Bot. English Botany, “Figures and descriptions of British
plants,” by-Sir J. E. Smith a Jas. Scmertie
Eschw. F. L. Eschweiler, a German botanist.
*Fée.