2) Se ee AE eh eater oy? he i pn fe o ALBERT R. MANN LIBRARY New York STATE COLLEGES OF AGRICULTURE AND HOME ECONOMICS AT CORNELL UNIVERSITY Ferns: British & foreign.The history, or Cornell University Library The original of this book is in the Cornell University Library. There are no known copyright restrictions in the United States on the use of the text. http://www. archive.org/details/cu31924001743032 FERNS: BRITISH & FOREIGN. THEIR HISTORY, ORGANOGRAPHY, CLASSIFICATION, AND ENUMERATION. WITH A TREATISE-ON THEIR CULTIVATION, ETC. ETC. BY JOHN SMITH, A.L.S. ACAD. CHS. NAT. CUE, B0C., aND EX-CURATOR OF THE ROYAL BOTANIC GARDENS, KEW, LONDON: ROBERT HARDWICKE, 192, PICCADILLY, CONTENTS. PREFACE.,......... en re sian eamERtS oq 1® I. History oF tHE Inrropuction or Exoric Ferns ... 1 II. OreanocraPHy, EXPLANATION OF THE TERMS USED 1n Descrisine FErns ...... Ginx thicesesses wiseaen saviavaceve AA ‘TI. CLASSIFICATION OF FERNS EXPLAINED .......0005 Gace 56 {V. Geyeric CHARACTER AND ENUMERATION OF CULTI- VATED SPECIES ........ sjaaisolvarnaed Susana Fengte Pemettenawed 73 V. Erymotoey, tor Mranine or THE GENERIC Nawés... 293 VI. Tue CuttivaTion oF FErRyS..... I. Pretiminary Remarks. II. SpectaL on Pot CULTIVATION ..........ceeeeeeee .. 808 TIT. Navturat CULTIVATION ......cccececeeeeeeeueeee sarhOae IV. Warpian Case CULTIVATION ......... sisscneat .. 354 V. PROPAGATION ....... weeiaats nadaenneWaeerGindeaeste waney OOF VII. List or AvTHors anD Books QUOTED IN THIS WORK: =tavanestsesienibeaessmadaniesopamihivsaiecemnimaeeinag Ble VIII. Generat InpEx or GENERA, SPECIES, AND Syn- ONYMES ....... sec teeee ee eeeneeneeaaes aaltleman@eaaitein ouiectnine OO IX. INDEX OF SPECIAL 'PERMS ..cecssseececeeeseeeeseeseseeees 411 =, > J Re 4 WF ”, Fitch, det, &. Ww Rugte. EXPLANATION OF THE PLATE. ORGANOGRAPHY, page 44. Annulate Sporangia— Fic. 1. Sporangium with a vertical ring, mag. 100 diameters. 2. + horizontal ring, mag. 100 diameters. 3. apical ring, mag. 100 diameters. (a) Spores of sacle highly magnified, 200 and 300 diameters. Lxannulate— Fic. 4. Sporangia (two) free, opening by a vertical slit, mag. 25 diameters. 5. Sporangia united (synangium), Gpautiig by pores, mag. 7 diameters. 6. Sporangia united (synangium), opening by slits, mag. 9 diameters. , (a) Spores of each highly magnified, 300 diameters. Indusia— Fig. 7; Indusium peltate orbicular, slightly magnified. 8. 7‘ lateral reniform, ditto. 9. ” calyciform, ditto. 10. 9 linear, interiorly attached, _— ditto. 11. ” valvate, ditto. 12, universal, ditto. PREFACE. EARLY ten years have now elapsed since my N “Catalogue of Cultivated Ferns” was pub- lished. During that period many new and fine species have been introduced to the gardens of this country. I have therefore been induced to draw up the follow- ing enumeration, including in it all the species that have come under my notice in a living state, either by the evidence of plants in the Kew collection or by specimens from living plants sent me from various sources. Thinking it might be interesting to Fern growers, T have drawn up a brief history of the introduction of Exotic Ferns from the earliest records to the present time. In order to assist students in the study of Ferns, an explanation is given of the principal organs and terms used in describing them, as well as remarks on their classification. I have also given an alpha- betical list of the generic names, accompanied with the derivation of each name. Great complaints are frequently made, and not with- x PREFACE, out good reason, of the many names given to the same plant; to assist in correcting this, I have drawn up a complete Index of the species and synonymes noticed in this work ; therefore, by turning to the page referred to in the Index, the name will be ‘found either with a number before it showing it to be the adopted name, or following the same as a synonyme, or what it has been and is still called by some writers and cultivators. Although this Index contains nearly three thousand names, yet, it must be understood, I have only taken up the synonymes that I consider most essential ; those who desire to enter more fully into this subject, may consult the “ Index Filicum” of Mr. T. Moore— a work which, when completed, will be a treasure to pteridologists. With regard to the wood-cuts, I have to explain that about half of the number were not specially pre- pared for this work; these, in general, represent a portion of the fronds of their natural size,"in some cases without fructification, the venation only being given. The drawings for the other half have been made principally from specimens in my herbarium or from living plants. In many of them a magnified portion of the frond is given, showing the character of the genus more distinctly. As a companion to the scientific enumeration, I have given a treatise on their cultivation, which occu- pies a considerable space in this book, and it is hoped will be of service to the Fern-growing public, PREFACE. xi In drawing up this treatise I have been greatly assisted by Mr. Henry Prestoe and Mr. William Gower (both recently foremen in the Fern department at Kew), two highly scientific and nature-observing practical cultivators. In conclusion, I have to thank Dr. Berthold Seemann and R. Heward, Esq., for looking over the proofs as they passed through the press. Several causes have occurred to delay the publication of this work; the principal one being a partial failure of my sight, incapacitating me from much use of the pen and examination of new garden Ferns, which I trust will be accepted as my excuse for any errors or omissions that may be found in the following pages. J. SMITH, Ex-Curator, Royal Botanic Gardens, Kew. Kew, May, 1866, FERNS: BRITISH AND FOREIGN. HISTORY OF INTRODUCTION OF EXOTIC FERNS. HOUGH Ferns now occupy a conspicuous place in our gardens, and are in high favour with cultivators, it is only in comparatively recent times that they have been brought into notice. During the last - century certain classes of plants came into fashion, and after-a season of popularity again fell into disrepute. Thus: Tulips were once the rage. At the time of the establishing of the several pro- vincial Botanic Gardens, all of which were founded upon a strictly botanical footing, though many of them have now, to a greater or less extent, degenerated ‘into places of amusement, the plants in greatest demand were those of our New Holland and Cape colonies, ’ principally the Heaths, Proteas, Aloes, and their kindred. In after-years dealers obtained large prices for Cactuses; but, with the exception of a few of the easily-grown and most showy kinds, these are now scarcely saleable. Still more recently the magni- ficent-flowering Orchids were promoted to the first place in our gardens; and though these may still be said to maintain their position, the expense attending B 2 FERNS: BRITISH AND FOREIGN. their cultivation is so great that they are for the most part confined to the gardens of the wealthy. Ferns, on the contrary, may, as a general rule, be grown in a comparatively inexpensive manner. The discovery made by Mr. N. B. Ward, that these. plants can be grown to great perfection in small ornamental closed cases (now well known as “ Wardian Cases”’), suitable not only for the drawing-rooms of the wealthy, but for humbler dwellings, renders it pos- sible for amateurs to indulge their love of Ferns without going to the expense of erecting hothouses and em- ploying a staff of gardeners; and it is to be hoped that this will be the means of retaining them in favour and spreading them still wider. The enumeration in the following pages shows that at the present time above nine hundred exotic species of Ferns are cultivated in the various public and private gardens in this country; and of these by far the greater number have been introduced during the last quarter of a century. A very large, indeed almost a complete, collection of them may at present be seen in the Royal Botanic Garden at Kew, where, during forty years, I devoted attention to their cultivation, and to the study of their generic and specific distinctions, using every endeavour in my power, assisted by the extensive influence of the eminent Director, Sir W. J. Hooker, to introduce new species, both by raising them from spores taken from my herbarium, and through correspon- dence with persons residing in our Colonies and the Superintendents of Botanic and other gardens on the Continent. Being well acquainted with the latter branch of the subject,—the introduction of new HISTORY OF INTRODUCTION OF EXOTIC FERNS. 3 species,—I have thought that a few pages devoted to the history of these plants in its connection with our gardens might prove acceptable. My own personal knowledge dates from about the year 1822; and for information respecting those in cultivation previous to that time, I have taken for my guide the most im- portant and most reliable of the garden catalogues. The first work upon which dependence c can be placed for the early-introduced species is the so-called second edition of the “ Hortus Kewensis,” published by the younger Aiton, in 1813; I say the so-called, for there were in reality two previous editions—one by Dr. John Hill, in 1768, and a second by William Aiton, in 1789. In the first of these only ten exotic species are recorded as being in the garden at Kew, which then belonged to the Princess of Wales, but the dates of their introduction are not given; and in the second, that of the elder Aiton, thirty-four, all of which have the name of their supposed introducer attached to them, and the date at which they were introduced. Ihere give preference to the last edition, as I know, frorh personal intercourse with the com- pilers, that great care was exercised in its prepara- tion, particularly with regard to the rectification of dates, and I shall take it as the basis of my remarks. ‘But, in the first place, it is necessary to say a few. words in defence of that work. It has been asserted that a considerable number of. the plants enumerated in the “ Hortus Kewensis” never existed in a living state, either in the garden at Kew or elsewhere in this country. This statement, however, is merely supposition, and rests upon questionable authority. In the early days of Kew Gardens, large B 2 4; FERNS: BRITISH AND FOREIGN, sums of money were devoted to the payment of bota- nical collectors, and great exertions were made, under the patronage of Sir Joseph Banks, to stimulate the introduction of new and rare plants, by inducing the commanders of ships of war and East-Indiamen to take an interest in the subject. By these means a very large number of plants were actually introduced into the country in a living state ; but the Department under whose charge the garden then was, took no steps to provide proper accommodation ; and this, together with the very frequent change of foremen, led, as a natural consequence, to the death of the | plants. Taking, then, the third “ Hortus Kewensis,” and excluding our indigenous species, I find that the total number of “Garden Ferns”? introduced previously to the year 1813 amounts to as many as eighty-three. The merit of being the first- introducer of these plants belongs to Mr. John Tradescant* the younger, who in early life made a voyage to Virginia; and I find it recorded in Parkinson’s ‘“ Theatrum Botanicum,” published in 1640, that upon his réturn from that country in 1628 he brought with him, amongst other rare plants, the Cystopteris bulbifera and Adiantum pedatum. These, therefore, must be regarded as the nucleus of our present large collections. At first the progress seems to have been exceedingly slow, for between the time of Tradescant and the close of the seventeenth century, only five additional species were introduced ; viz., Asplenium rhizophyllum and Onoclea * John Tradescant had a Botanic Garden and Museum at Lambeth. HISTORY OF INTRODUCTION OF EXOTIC FERNS. 5 sensibilis from North America, the former in 1680 and the latter in 1699, in which year also Adiantum reni- forme and Davallia Canariensts were brought from Madeira, while the fifth, Blechnwm australe, was, according to Plukenet, who figured it in the second part of his “ Phytographia,” cultivated in the King’s garden at Hampton Court as early as the year 1671, when his work was published, the garden there containing a considerable collection of rare plants. During the next forty-two years no additions appear to have been made, excepting the Phlebodium aureum, which was introduced by Lord Petre at some time prior to 1742, the date of his death, the precise year being unknown. Progressing onwards, Ido not find any more recorded until the year 1769 ; but between that year and the commencement of the following century sixty-eight species were added to the eight already existing in our gardens. Out of this large number, no less than thirty-seven were brought home by Rear-Admiral Bligh, in H.M.S. Providence, on his return, in 1793, from his second voyage, undertaken for the purpose of introducing the Bread-fruit and other useful trees into our West Indian colonies. And, finally, during the first few years of the present century, up to 1813, the date of the publication of the “Hortus Kewensis,” seven others were introduced. . Asummary of the foregoing shows that upwards of one-half of the Ferns known at the last-mentioned date were West Indian species, forty-four having been received at various times from those islands, the majority through Bligh’s expedition. North America and Madeira, with the neighbouring islands, stand 6 FERNS: BRITISH AND FOREIGN. next as the largest contributors, fifteen having been introduced from the former and twelve from the latter country ; while of the remaining sixteen, four appear: to have come from the East Indies, four from the Cape of Good Hope,-three from New Holland, and: one from St. Helena,—making in all eighty-three species. The next catalogue of garden plants worthy of notice is the “Hortus Suburbanus Londinensis,” published in 1818, only five years after the “ Hortus Kewensis,” by Mr. Sweet, the Superintendent of the then celebrated nursery of Mr. Colville, at Chelsea. In it I find an enumeration of one hundred and eight exotic ferns; but this work, like the similar more im- portant “Hortus Britannicus,” brought out by the in- defatigable Loudon in 1830, and which contains no less than three hundred and thirty exotic Ferns, includes” not only a considerable proportion of bad species, but also a large number that did not really exist in British gardens, many having been entered without authentic evidence, and others added upon the mere expectation that they might ‘shortly be introduced,— expectations which, in many cases, have not been realized to this day. No reliance can therefore be placed upon either of these works, and I cannot accept them as authorities. During the latter part of the eighteenth century and the commencement of the nineteenth, the only pri- vate individuals who turned their attention, with any amount of energy, to the introduction of new and rare plants, were the long- and far-famed nurserymen at Hackney, the Messrs. Loddiges; and to them we owe the greater part, if not the whole, of the HISTORY OF INTRODUCTION OF EXOTIC FERNS. 7 Ferns existing at that period in British gardens, and not included in the “Hortus Kewensis.” Speaking from my personal recollection of the im- portant collection of plants in the Hackney Nursery, as it existed in the year 1825, I think it contained about a hundred good species of exotic Ferns; but I can obtain no earlier catalogue than one published in 1849, in which two hundred and fifty-one are enumerated, In the year 1822 I found the collection of Ferns at Kew extremely poor, especially as regards Tropical species, very many of those introduced in previous years having been lost, and very few new ones added. Any person who remembers the hothouses in existence forty years ago, will have but little difficulty in accounting for the falling off of the Fern collection. In those days hot-water pipes were unknown, and the houses were exclusively heated by means of brick flues, too often imperfectly constructed, and the excessively dry and ungenial atmosphere thus induced was quite unsuited for the good cultivation or even for the mere preservation of these moisture- loving plants. Nearly all the North American species enumerated in the “‘ Hortus Kewensis” were growing very finely in a north border, and most of the Madeira species were also in existence; but, including these and the few added since 18138, I cannot estimate the entire Kew collection of exotic Ferns at that period at more than forty species. Between 1813 and 1846, when my first Catalogue of the Ferns at Kew appeared, no reliable list was published in this country. Several, however, were brought out by Continental botanists, which 8 FERNS: BRITISH AND’ FOREIGN. are of sufficient importance to be worthy of a passing notice. The first of these in point of date is the “ Enumeratio Plantarum Horti Regii Botanici Bero- linensis,” by Wildenow, published in 1809, with a Supplement by Schlechtendal, bringing it down to 1813. We are thus enabled to compare the num- bers in the Kew and Berlin gardens at the same period; which were eighty-three in the former, and thirty in the latter, including eleven not known at Kew. During the succeeding nine years more atten- tion appears to have been given to Ferns at the Berlin garden; for Link, in his first ‘‘ Enumeratio,” in 1822, describes ninety-one exotic species, which is more than double the number then existing at Kew. After this the increase in number was still more rapid; for in the second edition of Link’s “ Enumeratio,” published in 1883, no less than two hundred and thirty-nine are described; and in the third, in 1841, two hundred and fifty-eight, exclusive of varieties. By this time, however, the collection at Kew had received large additions, both through importations of living plants and by raising from spores. In 1845 it was so. extensive that I was induced to draw up a classified enumeration, which was pub- lished as an appendix to the Botanical Magazine for 1846. The number of exotic species there enume- rated is three hundred and forty-eight, and I do not think many were to be found in other gardens in this country which were not at Kew, so that the Kew list may be taken as a guide to the number then in British gardens generally. Four years later, Kunze, of Leipzig, contributed to HISTORY OF INTRODUCTION OF EXOTIC FERNS. 9 the 28rd-volume of the “Linnea” an alphabetical index of the Ferns cultivated in European gardens, and in this the large number of eight hundred and forty-three exotic species, are enumerated; but the authorities upon which a very considerable portion of these were inserted cannot be relied upon, many names having been taken from such catalogues as those of Sweet, Loudon, &., and I am therefore obliged to con- clude that the number given by Kunze as living in Europe in 1850 is greatly exaggerated. This con- clusion, too, is confirmed by the fact that in 1857, after I had by correspondence become acquainted with the collections in the principal gardens on the Conti- nent, and after that at Kew had obtained most of their novelties by means of exchange, I could, in my “ Catalogue of Cultivated Ferns,’ enumerate only five hundred and sixty exotic species as known in British gardens. Since the last-mentioned year, the con- stantly increasing demand for Ferns consequent upon their wider spread cultivation, has greatly stimulated the introduction of new ones, and our collections have increased at the rate of about fifty species a year. It now remains to say a few words regarding the means by which these plants have. been obtained, and the persons who have been most active in introducing them, taking the Kew collection as a foundation. Firstly, with reference to the paid collectors employed in various parts of the world, directly or indirectly in the service of the Royal Botanic Garden, Kew, and to whom that garden is indebted for additions to its Fern collection. It would appear that so long back as the year 1775 Mr. Francis Masson, one of the earliest, if 10 FERNS. BRITISH AND FOREIGN. not the earliest, collector sent out from Kew, and who succeeded in introducing large numbers of Cape Proteacee and LHricacece, sent home several Ferns from the Cape of Good Hope and Madeira. This col- lector proceeded to the Cape in 1774, and came home by way of Madeira about the year 1778, returning again in 1736, and remaining at the Cape during the nine following years. arly in the present century Mr. George Caley, who was originally a horse-doctor, residing near Birmingham, but acquired a love for plants through collecting herbs, was sent out by Sir Joseph Banks to New South Wales, and to him we owe Platycerium alcicorne, Doodia aspera, and Davallia pyxidata, the first introductions from Aus- tralia, received about the year 1808. The next col- lectors to whom the garden was indebted for Ferns, are the Messrs. Allan Cunningham and James Bowie. They left Kew in 1814, on a botanical expedition to Brazil, where they remained exploring the country and sending home large collections till 1816, when the former proceeded to New South Wales, and the latter to the Cape of Good Hope. No living Ferns appear to have resulted from the Brazilian ex- pedition ; but several Australian species and one or two from Norfolk Island were received from Mr. Cunningham, and two or three from Mr. Bowie from the Cape. Several other collectors were employed in the ser- vice of these gardens, when under the Directorship of Mr. Aiton, such as Messrs. Barclay and Armstrong ; but I can trace no Ferns to them, nor, with certainty, to David Lockhart, a gardener from Kew, who accom- panied the ill-fated expedition of Captain Tuckey up HISTORY OF INTRODUCTION OF EXOTIC FERNS. 11 the Congo, in 1816, though I believe it possible that some of the first-known Western-African species are attributable to him. Lockhart was one of the few who did not fall a victim to the deadly climate of that country. After lying a long time in the hospital at Bahia, he returned to England, and shortly afterwards received the appointment of Superintendent of the Botanic Garden in Trinidad, where he died in 1845, after a service of a quarter of a century in that island, during which period he sent many fine plants to Kew. The principal collectors employed directly or indi- rectly in the service of Kew during the Directorship of Sir William Hooker, and to whom the garden is in- debted for any additions to its Fern collection, are Messrs. Purdie, Seemann, Milne, Barter, and Mann. The first of these, Mr. William Purdie, was engaged as collector in 1848, in which year he was despatched to Jamaica, and after spending several months in the exploration of that island, and forwarding many valuable plants. to Kew, he proceeded to Santa Martha, and was employed for more than two years in various parts of New Granada. To him we are indebted not only for numerous showy flowering plants, but for a considerable number of our garden Ferns. Upon the death of Mr. Thomas Edmonston,* Mr. (now Dr.) Berthold Seemann was appointed his suc- cessor, as botanist to H.M.S. Herald, and joined that vessel at Panama, in January, 1847, remaining with her until her return to England in June, 1851. Previously * Mr. Edmonston was shot by the accidental discharge of a rifle, at Sua Bay, Ecuador, January 24, 1846. 12 FERNS : BRITISH AND FOREIGN, to joining the Herald, he explored several parts of Panama and Veraguas, and while on board the Herald visited various parts of the western opast of America, between Lima on the south and California on the north, occasionally making long journeys inland, particularly in Peru, Ecuador, and Mexico. He likewise accompanied the Herald in her three voyages to the Arctic regions in search of the ill-fated Sir John Franklin; visited twice the Sand- wich Islands; and returned home by way of Hong- Kong, Singapore, and the Cape of Good Hope. Not- withstanding the few facilities afforded by surveying expeditions for collecting living plants, Dr. Seemann succeeded in introducing some interesting ones to - our gardens; and among Ferns we may mention two very remarkable ones, Deparia prolifera from the Sandwich Islands, and Dictyoxiphium Panamense from Panama. On the Herald being recommissioned in 1852, Mr. William Milne was appointed assistant-botanist, for the special purpose of collecting plants for the Royal Gardens. During the six years he was attached to the Herald he visited New Caledonia, the Fiji, and other Polynesian islands, as well as many other places in the southern hemisphere. The Admiralty having, early in 1857, determined deosunnding out a second exploring expedition up the Niger, under the command of Dr. Baikie, R.N., Mr. Charles Barter, a zealous and intelligent young gardener, was appointed to accompany him in the capacity of botanist, and to collect plants for Kew. The officers of the expedition proceeded by way of Sierra Leone, and joined the Duyspring, a small HISTORY OF INTRODUCTION OF EXOTIC FERNS. 13 vessel fitted for river navigation, at Fernando Po; but, unfortunately, after they had penetrated up the river as far as Nupe, their ship was wrecked, and they were obliged to form a camp on the banks. Poor Barter, ~ however, fell a victim to the deadly climate in July, 1859, Asa collector he was indefatigable in the dis- charge of his duties, and discovered a large number of undescribed plants, including several new genera, one of which has been dedicated to his memory under the name of Barteria, But, owing to the mishaps attending this expedition, and the difficulties of transit, no living plants were received from the interior of the country, but several Ferns were trans- mitted from Sierra Leone and Fernando Po.* No sooner had the news of Barter’s death arrived, than Mr. Gustav Mann, undeterred by his fate, volunteered to fill the vacancy, and- was appointed accordingly. Being unable, from want of means of communication, to join Dr. Baikie, he was employed for three years in exploring the island of Fernando Po and the African coast, in the neighbourhood of the Camaroons and Gaboon rivers. In spite of the difficulties and dangers attending the under- taking, he made several ascents of the lofty Clarence Peak of Fernando Po and of the Camaroons moun- tain on the African mainland. On the former, at an elevation of 5,000 feet, he found a fine new Cyathea, forming groups, with stems rising to a height of 30 feet. We are likewise indebted to him * Since the above was written, news has reached this country of the death of Dr. Baikie at Sierra Leone, in January, 1865, whilst on his homeward voyage, - 14 FERNS: BRITISH AND FOREIGN. for many other rare and interesting living Ferns, and fine sets of dried specimens. Besides contributions from special collectors, the Fern collection at Kew has been largely increased through the instrumentality of the officers of our numerous Colonial Botanic Gardens, and likewise by many private individuals residing in foreign countries ; and as the exertions of these gentlemen are worthy of being recorded, I give a few particulars respecting them, adopting a geographical arrangement for the sake of brevity. The first Colonial garden from which, so far as I am aware, Ferns were received at Kew, was that of Seylon, Mr. Alexander Moon, the Director, having, in 1824, sent home a collection of plants, amongst which was Niphobolus costatus. But the first persou who forwarded any considerable number from that island was the lamented Mr. George Gardner, well known to botanists as a botanical traveller in Brazil, who was Director of the Ceylon garden between 1844 and his death in 1848. This gentleman was succeeded by Mr. (now Dr.) G. H. K. Thwaites, the present able Director, and to him also the Kew collection is greatly indebted for a large number of rare and beautiful species; such as NSehizoccena sinuata, Asplenium radiatum, Actinostachys radiata, Helminthostachys Zeylanica, and many others. Though Continental India is extremely rich in Ferns, it has, singularly enough, contributed very few to our gardens, no persons in that country having devoted themselves specially to the subject; indeed most of those received thence have been accidentally imported along with Orchids, including the half-dozen species HISTORY OF INTRODUCTION OF EXOTIC FERNS, 15 recorded in my first Enumeration as coming from Dr. Wallich, the only Superintendent of the Calcutta garden who has the credit of having introduced any. Indeed, with the exception of those from Ceylon, Kew has received very few Ferns from Asia and. the adjacent islands, most of those now in cultivation having been introduced by nurserymen or through Continental gardens. Two or three were brought from Hong-Kong, in 1850, by Mr. J. C. Braine, including one which proved to be a new genus, and to this I gave the name of Brainea in honour of its introducer. Another tropical ee in the Kastern hemisphere, whence large additions have been made to the Fern collection at Kew, is the Mauritius. The Botanic Garden in that island has long enjoyed the reputation of possessing a fine set of plants; but until the year 1852, when. it came under the able management of the present Director, Mr. James Duncan, very little correspondence was kept up with the gardens of Europe. Mr. Duncan has, at considerable risk, ransacked the forests of the island in quest principally of Ferns, and has been very successful in transmitting living plants to this country, enriching our gardens with many fine species. The “Synopsis Filicum Capensis” of Pappe and Rawson shows that the Fern Flora of Southern Africa is extremely rich; but up to the present time we possess scarcely a dozen Cape species in our gardens, and most of them have been raised from spores. Alsophila Capensis and Lomaria Capensis were intro- duced in 1845 by Mr. Charles Zeyher, as also were Lastrea athmantica and Cyathea Dreget from Natal by Mr. J. Plant. 16 FERNS: BRITISH AND FOREIGN, Progressing westwards we come to the two strangely isolated islands in the Southern Atlantic, St. Helena and Ascension. From the former of these we have As- plenium compressum, introduced by Mr. Thomas Fraser in 1825, and Asplenium reclinatum, brought home by Dr. J. D. Hooker on his return from Sir John Ross’s Antarctic expedition in 1844; together with Lomaria alpina and L. Magellanica from the Falkland Islands ; while from Ascension Mr, Wren sent numerous fine plants of Maratiia purpurescens in 1848. From Australia several individuals: have been con- tributors. Grammitis Australis was received from the Sydney garden in 1833, when under the direction of Mr. Richard Cunningham; and Mr. Charles Moore, the present Director of that garden, has also intro- duced several, including Trichiocarpa Mooret, from New Caledonia, while to Mr. Bidwill we owe the curious Platycerium grande. But some of the most beautiful of the Australian Ferns, such as the Gilei- chentas, were transmitted to this country by Mr. Walter Hill, the able Director of the Botanic Garden of Brisbane, in the rapidly rising colony of Queensland, who obtained them during his stay in Sydney in 1850. Two species of Gileicheniacee were, however, previously known in our gardens,—the Glei- chenia nuicrophylla and G. flabellata, both of which, together with several other Ferns, were sent from Tasmania, in 1845, by Mr. Ronald Gunn. About the year 1841 or 1842; some very fine Ferns, including two Tree-Ferns, the Dicksonia squarrosa and Cyathea medullaris, were brought from New Zealand, where they had been collected by Mr. J. Edgerly, a gardener, who had proceeded to that country on HISTORY OF INTRODUCTION OF EXOTIC FERNS. 17 speculation, and who was the first to introduce tho beautiful Veronica speciosa. Others have also been sent from New Zealand by the Rev. William Colenso, and by the late Dr. Sinclair, R.N., the beautiful little Trichomanes reniforme being one of those due to the latter gentleman. Turning next to the West Indies, we commence with the island of Jamaica, whence more Ferns have been received at Kew than from any other part of the Western hemisphere. The person to whose energy and perseverance this is mainly due is Mr. Nathaniel Wilson, the Island botanist and Director of the Botanic Garden. He has been a resident in the island for upwards of twenty years, and during that time has thoroughly explored the Blue Mountains and other districts rich in Ferns, liberally forwarding to Kew the results of his numerous journeys. Among his earliest contributions was the beautiful Tree-fern, Cyathea arborea, which, though recorded in the “‘ Hortus Kewensis” as having been brought home by Admiral Bligh in 1798, had long been lost to our gardens. Within the last few years he has succeeded, after many failures, in transmitting numerous species of Trichomanes and Hymenophyllum, which now form so conspicuous a feature in the present rich collection. The other contributors from the same island are, in 1851, Mr. George Manson, and in 1854 and following years, W. T. March, Hsq., the latter gentleman sending several arborescent species as well as Hymenophyllece, and others. Numerous fine species, including several Oyatheas and Alsophilas, were received in 1855 and 1856 from the French island of Martinique, where they had been Cc 18 FERNS: BRITISH AND FOREIGN. collected by the Director of the Garden, M. Belanger. A considerable number of rare species were likewise sent from Dominica in 1853, and several following years, by Dr. Imray; such as Hemitelia Imrayana, Neurocallis prestantissima, Elaphoglossum undulatum, and the true EH. longifolium, &c. While from Trinidad we are indebted to the Island botanist, the late Dr. Criiger, for Schizca elegans, Saccoloma elegans, Amphidesmium rostratum, Hymenostachys diversifrons, Trichomanes pennatum, and other equally rare species. His suc- cessor, Mr. Henry Prestoe, who previous to his appoint- ment last year was foreman of the collection of Ferns at Kew, has already transmitted a large collection of rare Ferns, and in the finest condition, proving that they had been collected and put up by one who perfectly understood their nature. The situation he now holds will enable him to be of great service in transmitting new plants to this country. Comparatively few, considering the richness of its Fern Flora, have been received from tropical America; Dr. Gardner and J. Wetherall, Esq., in Brazil; H. Cadogan Rothery, Esq., in Guiana; Mrs. Colonel McDonald, in Honduras ; and Mr. Wagener, in Vene- zuela, being the principal contributors. From the above it will be seen that a large number of Ferns have been introduced through the agency of the garden at Kew; but, besides these, a very con- siderable number.are due to the exertions of some of our leading nurserymen, who, in consequence of the great demand for, and the large prices realized by, the finer and rarer species, have imported them, either direct from their native countries, or from the Con- tinent, where, as will be presently noticed, a good HISTORY OF INTRODUCTION OF EXOTIC FERNS. 19 many species not previously known in the gardens of this country have been introduced. I have already alluded to the Messrs. Loddiges, of ‘Hackney, as having at an early period turned their attention to Ferns, and as being the earliest to form , a collection of them. But the only nurserymen whose names are recorded in the second edition of the “Hortus Kewensis” are the old-established firm of Messrs. Lee & Kennedy, of Hammersmith, who are stated to have introduced Polypodiwm asplenifolium and Asplenium monanthemum in 1790: in later times the Messrs. Lee have imported several from New Zealand. Other New Zealand species have been brought into notice by Mr. Standish, of Bagshot, they having been collected in New Zealand by Mr. J. Watson, now a nurseryman at St. Alban’s, and who still continues to import. Several sent from Japan by Mr. Fortune have likewise been sent out from Mr. Standish’s nursery. To the Messrs. Low & Sons, of the Clapton nursery, we are indebted for some rare Bornean and Malayan species, collected by Mr. Hugh Low, jun., and amongst others for the remarkable Arthropteris obliterata, called Lindscea Lowii in the gardens, and the little curious Leucostegia parvula; but more especially many rare species of Hymenophyllum and Trichomanes, as well as the rare Thyrsopteris elegans, collected by Mr. Thomas Bridges in Juan Fernandez. But to the Messrs. Veitch & Sons, of Exeter and Chelsea, among nurserymen, must be assigned the credit of having introduced the greatest number of these plants, the collectors employed by them in Chili and other parts of the American continent, in India, the Malayan continent c2 20 FERNS: BRITISH AND FOREIGN. and islands, and in Japan, having sent home numerous fine species, while through other sources they have obtained many additions from Australia, New Zea- land, and other countries. Messrs. Rollisson, of Tooting, have likewise succeeded in enriching our collections with a considerable number, received principally from Mr. John Henshall, their collector in Java and the neighbouring islands. Mr. Robert Sim, of Foot’s Cray, has an extensive collection, which he increases by importations, and is very successful in raising plants from spores; and the Messrs. Back- house & Son, of York, the principal nurserymen cultivators of exotic Ferns in the provinces, have introduced a good many fine species of Trichomanes, and others from Chili and the West Indies. | —— —j 2 => | ——————F3 — | = = = = — ——| i =e ——— s —— == = =e z= — es Genus 109.—Portion of barren and fertile pinna, naturel size. No. Le AN ENUMERATION OF CULTIVATED FERNS. 209 flexuose, climbing to an indefinite height ; pinnse 1-2 feet long, adherent. Veins forked; venules combined by a transverse marginal vein, and in the fertile also near their base by a transverse, continuous, sporangiferous receptacle, forming a linear costal sorus. Indusiwm revolute, vaulted, cylindrical, sporangiferous along its base. 1. S. volubile, J. Sm. in Hook. Gen. Pi. #. 93. Blechnum volubile, Kawlf.; Kunze, Anal. ¢ 13; Hook. Gard. Ferns,t.15. Blechnum scandens, Bory, 4 in Dup. Voy. t. 86.—Tropical America, Triss IX.—_ASPLENIE A. Sori oblong or linear, oblique to the midrib or axis of vena- tion, Furnished with a plane or vaulted lateral indusiwm. 110. ASPLENIUM, Linn. Vernation fasciculate, erect or decumbent, rarely uniserial sarmentose. Fronds varying from simple-entire to decompound and from a few inches to 2-6 feet high, generally smooth Veins rayed, forked, or pinnate; venules free, sporangiferous cn the superior side., Sori simple, oblong, or linear, Indusiuin plane or vaulted. § 1. AspLenium verum.—Indusiwm piane, 1. Lancewm group. Vernation wniserial, sarmentum slender. Fronds distant, linear-lanceolate, 1-14 foot long. ae siinple, anti or opposite binate. (Triblemma.) 1. A. lanceum, Thunb. Ie, Plant. Jap. Dec, 11, t. 18. A. subsi- nuatum, Hook. et Grev. Ic. Fil, t. 27. Diplazium Jan- ceum, Presi. Scolopendrium dubium, Don,—Inuia, China, Japan, 2. Serratum group. Vernation fascieulate, erect. Fronds simple, broad, elliptical, or lanceolate, 1-2 feet long. (Phyllitis.) 2, A. sorratum, Linn. (Plum. Fil. t. 124); Schk. Fil, t. 64; Hook. Fil. Exot. t. 70.—Tropical America, P 210 FERNS: BRITISH AND FOREIGN. 3. A.erenulatum, Presi. A. Nidus, Radd. Fil. Bras. t. 58 (non Linn.). A. Brasiliense, Hort. (non Sw.) ; Lowe’s Ferns, 5, t.14 B—Tropical America. “ A. sinuatum, Beawv. Fl. d’Oware, 2, t.79; Hook. Fil. Exot. t, 16.— West Tropical Africa. Genus 110.—Fertile pinna, underside. No. 69. 5. A. stipitatum, J. Sm. Neotopteris stipitata, J. Sm. Cat. Cult, Ferns (1857). Asplenium squamulatum, var. 3 Smithii, Hook. Sp. Fil. p. 83.—East Indies. AN ENUMERATION OF CULTIVATED FERNS. 211 3. Palmatum group. Vernation decumbent, subfasciculate. Fronds palmately lobed. 6. A. Hemionitis, Linn.; Hook. Bot. Mag. t. 4911. A. pal- matum, Lam.; Schk. Fil. t. 66; Lodd. Bot. Cab. 868 ; Lowe’s Ferns, 5, t. 6, (3 var. cristata, Gard. Chron. (Jany. 1865), Garden Sport.——South Europe, North Africa, Madeira. 4, Trichomanes group. Vernation fasciculate, generally becoming cespitose. Fronds pinnate, rarely pinnatifid only, linear, 3-12 inches long; pinne short, often dimidiate, with the costa excentric. (Asplenium verum.) a. Fronds pimnatifid. 7. A. alternans, Wall.; Hook. Gard. Ferns, t. 88. A. Dal- housize, Hook. Ic. Pl. t. 105.—Hast Indies. b. Fronds pinnate. 8. A. Trichomanes, Linn.; Bolt. Fil. t. 18; Eng. Bot. t.576; Sowerby’s Ferns, t. 30; Lindl. and Moore’s Brit, Ferns, t. 39; Lowe’s Ferns, 5, ¢. 22. A. anceps, Soland. ; Hook. et Grev. Ic. Fil. t. 195. A. melano- caulon, Willd, Var. incisum, Moore; Lindl. and Moore’s Brit. Ferns, t. 89 D, EH; Schk. Fil. t. 74, f. Var. cristatum, Moore; Lindl. and Moore's Brit. Ferns, t. 39 H. Var. depauperatum, Wollast. ; Lindl. and Moore’s Brit. Ferns, t.39 C. Var. multifidum, Moore; Lindl. and Moore’s Brit. Ferns, t. 89 G. Var. ramosum, Moore; Lindl. and Moore’s Brit. Ferns, t. 39 F. Europe, Madeira, South Africa, North India, Australia, North America. P2 212 FERNS: BRITISH AND FOREIGN. 9. A. ebeneum, Ait.; Lodd. Bot. Cab. t.5; Lowe's Frvvs, 5, #.2. A. polypodioides, Sw.; Schk. Fil. t.73.—North America. 10. A. monantbemum, Linn.; Sm. Ic. ined. t. 73; Lodd. Bot. Cab. t.1700; Metten. Fil. Hort. Lips. t. 9, f. 7,8; Lowe’s Ferns, 5, t.1 A—Tropical and Sub-tropical America, South Africa, Madeira. 11. A. Petrarchee, De Cand.; Hook. et Grev. Ic. Fil. t. 152; Lowe’s Ferns, t. 5, 38 A, and var. lata, 38 B—South of Europe. 12. A, formosum, Willd.; Hook. Fil. Exot. t. 16; Lowe’s Ferns, 5, t.43 B. A. subalatum, Hook. et Arn. Beech Voy. t. 71.—Tropical America. 13. A. Brasiliense, Radd. Fil. Bras. t. 51,f.1. A. dimidiatum, Lowe’s Ferns, 5, t. 13 A.—Tropical America. 14, A. tenellum, Roxb. in Beat. St. Helena. A. reclinatum, Moore et Houlst.; Lowe’s Ferns, 5, t=. 138 B; J. Sm. Cat. (1857). A. erectum,—var. proliferam, Hook. Fil. Eeot. t. 72. . alata, Sm.; Hook. et Grev, Ic. Fil. t, 18; Huok. Gen. Fil. t.7.—West Indies and Tropical America. 2. D. nodosa, Sm.; Schk. Fil. t.152; Hook. et Grev. Ic. Fil. t.51. Asplenium nodosum, Linn. (Plum. Fil. t. 108). —West Indies and Tropical America. 270 FERNS: BRITISH AND FOREIGN. 154. KAULFUSSIA, Blume. Vernation uniserial, contiguous, decumbent; sarmentum thick, fleshy ; each frond rising from between two short, fleshy, stipulaform appendages. Fronds broad, pinnately-trifoliate, with : : o> _ the two lower pinnz sometimes Z{| bipartite, 1-2 feet high, long, stipate, pale underneath, and- furnished with numerous con- cave dots. Veins costeform, parallel; venules compound ana- stomosing, with free veinlets terminating within the areoles. Synangia sessile, compital, super- ficial, distant, orbicular, concave- hemispherical, each consisting of 10-20 sporangia, laterally con- : : nate in a circular series, and Genes iigitly enlarged. No.1...” Opening by a slit on their interior side. 1. K. esculifolia, Blume; Hook. et Grev. Ic. Fil. ¢. 229; Hook. Gen. Fil. t. 59 A—Java and Philippine Islands. ! Oxprr II.—_OPHIOGLOSSACE. Flowerless plants consisting of straight evolved fronds, which produce from their disk or base a rachiform, simple or com- pound paniculate spike, of unilocular, sessile, connate, homo- geneous cases (sporangia), opening by a vertical slit in two valves, containing reproducing spores. 155. OPHIOGLOSSUM, Linn. Fronds erect or pendulous, with dissimilar fertile and sterile segments, or rarely the whole frond rachiform, Sterile frond or segments foliaceous, simple, entire, palmately lobed or dicho- tomously branched. Veins reticulated. Fertile segments spike- like, simple or rarely forked. Sporangia connate, in two rows, AN ENUMERATION OF CULTIVATED FERNS. 271 forming a distichous, synangeous spike, each cell opening horizontally in two equal valves. Genus 155.—Plant natural size ; fertile spike enlarged. No.3. Fronds erect, the sterile segments ovate or linear (Terrestrial). (Euophioglossum.) 1. O. Lusitanicum, Linn.; Hook. et Grev. Ic. Fil. t. 80; Lindl. and Moore’s Brit. Ferns, t. 51 C; Sowerby’s Ferns, t. 47. Ophioglossum vulgatum, var. angusti- folium, Hook. Brit. Ferns, t. 47—South of Europe. 2.0. pedunculosum, Desv.; Kumze, Fil. t, 29, f. 2— North America. 272 FERNS: BRITISH AND FOREIGN. 3. O. vulgatum, Linn. ; Schh. Fil. t. 153; Hook. Gen. Pil. 4.59 B; Eng. Bot. t. 108; Lindl. and Moore's Brit. Ferns, t. 51 B; Hook. Brit. Ferns, t. 46; Sowerby’s Ferns, t. 46.— Temperate Zone of the Northern Hemisphere. 4, O. reticulatum, Linn. ; Hook. et Grev. Ic. Fil. t.20 (Plum. Fil. t. 164).—Tropics. ** Fronds pendulous; the sterile segments ribbon-formed, usually dichotomously branched (Hpiphytal). (Ophioderma.) 5. O. pendulum, Linn.; Hook. et Grev. Ic. Fil. t.19; Hook. Gard. Ferns, t. 33.—Tropics of Hastern Hemisphere. 6. O.fureatum, J. Sm. Ophioglossum pendulum ; /3 furcatum, Presi, Tent. Pterid. Supp. p. 56.—Queensland. *** Fronds palmate, pendulous. (Cheiroglossa.) 7. O. palmatum, Linn. (Plum. Fil. t. 163); Hook. Ic, Pl. t.4. —West Indies, New Granada, Peru, Brazil, Mauritius. 156. HELMINTHOSTACHYS, Kaulf. Fronds erect, with dissimilar sterile and fertile segments. Sterile segments foliaceous, digitate, pedate. Veins forked; Genus 156.—Portion of sterile pinna, natura! size ; ditto enlarged; fertile spike, natural size. No. 1. AN ENUMERATION OF CULTIVATED FERNS. 273 venules free. Fertile segment w rachiform spike. Sporangia subglobose, sessile, in pedicellate crested whorls (rarely distinct), ‘ forming a clustered simple spike, opening vertically by a slit on the exterior side. 1. H. Zeylanica, Hook. Gen. Fil. t.47 B; Hook. 2nd Cent. Ferns, t. 94; Hook. Gard. Ferns, t. 28. Osmunda Zeylanica, Linn. Helminthostachys dulcis, Kaulf. —Ceylon, Malay, Molucca, Philippine, and other islands, 157. BOTRYCHIUM, Linn. Fronds erect, with dissimilar sterile and fertile segments. Sterile segments foliaceous, deltoid, bi-tripinnatifidly decom- Genus 157.—Portion of barren and fertile segmeuts, natural size; fertile enlarged. 2 274 - FERNS: BRITISH AND FOREIGN. pound, rarely pinnate. Veins forked; venules free. Fertile segments rachiform, compound paniculate. Sporangia distinct, in two unilateral rows, opening vertically in two equal valves. 1. B. simplex, Hitchcock ; Hook. et Grev. Ic. Fil. t. 82.— North America. 2. B. Lunaria, Sw. ; Schk. Fil. t.154; Hook. Gen. Fil. t.47 A ; Lindl. and Moore’s Brit. Ferns, t. 51 A; Hook. Brit. Ferns, t. 48; Sowerby’s Ferns, ¢. 45. Osmunda Lunaria, Linn. Eng. Bot. t. 318;— rataceum. Botrychium rutaceum, Sw.; Schk. Fil. t. 155 B— Temperate Zone of the Northern Hemisphere. 3. B. lunarioides, Sw. Botrypus lunarioides, Miche. ;— PB obliquum, A, Gray. Botrychium obliquum, Muhi. B. lunarioides, Schk. Fil. t=. 157;— y dissectum, A, Gray. Botrychium dissectum, Spr.; Schk. Fil. #.158.— North America, 4. B. Virginicum, Willd. Osmunda Virginica, Linn. Botry- chium Virginianum, Sw.; Schk. Fil, t, 156; Hook, Gard. Ferns, t. 29.—Temperate Zone of the Northern Hemisphere, Tropical America, East Indies, and Ceylon, Orvrz IV—LYCOPODIACE:. Flowerless moss or fern-like plants, consisting of firm, erect, creeping or pendulous, simple or branched, often flagelliform stems, furnished with acerose, rusciform or jungermannia-like, sessile leaves, which are generally imbricate, and often disti- chous and of two kinds, bearing in their axis, or on contracted terminal spikes, 1—3-celled reniform or globose sessile spore- cases (sporangia), of one or of two forms; one called Antheri- dangia, containing numerous spores ; the other Oophoridangia, containing 1-3-8 large spores. 158. PSILOTUM, Sw. Stems dichotomously forked, compressed or angular, rigid, erect or slender, pendulous; leaves obsolete or small, bract- . AN ENUMERATION OF CULTIVATED FERNS. 275 like. Sporangia subglobose, trilobed, vertically trivalved, solitary in the axis of the bract-like leaves. , Genus 158.—Portion of fertile frond, natural size; spore-case enlarged. No.1. 1. P. triquetrum, Sw.; Schk. Fil. t. 165 6; Hook. Gen. Fil. t. 87; Lodd. Cab. t. 1916. Lycopodium nudum, Linn. — Tropics and sub-tropical regions of both hemispheres. 159. LYCOPODIUM, Linn. Stems rigid, erect or slender, flagelliform, pendulous or creeping ; leaves acerose, subulate or rusciform, distant or imbricate. Sporangia reniform, vertically bivalved, solitary in . the axis of the leaves, or in terminal, rarely lateral, contracted, ament-like spikes. Spores numerous, small, uniform (Antheri- dangia only ?). TQ 276 FERNS: BRITISH AND FOREIGN. * Stems creeping. (Epigeous.) 1. L. clavatum, Linn.; Schk. Fil. t.162; Eng. Bot. t. 224.— Temperate Zone of the Northern Hemisphere, Britain. 2. L. annotinum, Linn.; Schk. Fil. t.160; Eng. Bot. ¢. 239, — Temperate Zone of the Northern Hemisphere, Britain. 3. L. inundatum, Linn.; Schk. Fil. t.162; Eng. Bot. t.1727. —Temperate Zone of the Northern Hemisphere, Britain. Genus 159.— Portion of plant, natural size; spore-case enlarged. - ** Stems erect. (Epigeous.) 4. L. Selago, Linn.; Schk. Fil. t. 159; Eng. Bot. t. 233.— Europe, Britain. 5. L. alpinum, Linn.; Schk. Fil. 1.161; Eng, Bot. ¢.234.— Temperate Zone of the Northern Hemisphere, Britain. 6. L.densum, Labil. Nov. Toll. 2, ¢. 251, f. 1—Australia. AN ENUMERATION OF CULTIVATED FERNS. 277 7. L. fastigiatum, R. Br—New Zealand. 8. L. complanatum, Linn. (Plum. Fil. ¢. 165, f. B); Schk. Ful. ¢. 163.—Temperate Zone of the Northern Hemi- sphere. 9. L. dendroideum, Micha. ; Willd. Sp. Pl. 5, p. 21.—North America. 10. L. cernuum, Linn. (Rheed. Mal. t. 2, ¢. 39); Burm. Fl. Zey: t. 66 (Plum. Fil. t. 155, f. A).—Tropics; very general. *ke Stems pendulous. (Epiphytal.) 1l. L. Phlegmaria, Linn. (Rheed. Mal. 12, ¢.14).—Tropics of the Eastern Hemisphere. : 12. L. verticilatum, Linn.; Willd. Sp. Pl. 5, p. 48.— Mauritius. 18. L. ulicifolium, Vent.; Willd. Sp. Pl. 5, p. 27.—India. 14, L. Hookeri, Wall. ; Hook. et Grev. Ic. Fil. t. 165.—India. 15. L. taxifolium, Sw.; Willd. Sp. Pl.5, p,48.—West Indies. 160. SELAGINELLA, Spring. Fern-like plants. Stems of frondules creeping, sub-ascending or erect, scandent, of undefined extension (surculose), or rising Genus 160.—Portion of plant, and the two kinds of spore-cases enlarged. singly from an underground elongating stolon (cawlescent), or in a fasciculate manner from a simple erect centre (roswlate) ; 278° FERNS: BRITISH AND FOREIGN. leaves generally imbricate, distichous, jungermannia-like, of two sizes; the smaller stipuleform. Sporangia reniform, vertically bivalved, sessile in the axis of the leaves, or on con- tracted terminal spikes. Spores of two forms: in some, sporangia (Antheridangia) small and numerous; in others (Oophoridangia), 3-4, large. A. Plant leafy on all sides, 1. 8. spinulosa, Spring. Lycopodium selaginoides, Linn. ; Schk., Fil. t.165; Eng. Bot. t.1148.—Europe, Britain. 2. 8. uliginosa, Lab. Nov. Holl. 2, p. 104, t. 251, f. 2; Willd. Sp. Pl. 5, p.32.—Australia and Tasmania. B. Leaves distichous. “§ 1. Plants surculose. + Stems decumbent, creeping. 3. S. Apus, Spring. Monogr. p.75. S. densa, Hort. Lycopo- dium Brasiliense, Radd. Fil. Bras. 82, ¢. 1—North America and Brazil. 4, S. Ludoviciana, A. Braun. Revise, No. 5.—Louisiana. 5. §. Helvetica, Link ; Spring. Monogr. 2, p. 88. Lycopodium Helveticum, Linn.—Europe, Switzerland. 6. S. denticulata, Link; Spring. Monogr. 2, p. 82. Lyco- podium denticulatum, Linn.—Central Europe. 7. S. delicatissima, A. Brawn. Revise, No.8, 8, microphylla, Spring. —Columbia. 8.8. serpens, Spring. Monogr. 2, p. 102. Lycopodium serpens, Desv. §. mutabalis, Hort. S. variabilis, Hort. S. Jamaicensis, Hort.— Jamaica. 9. S. uncinata, Spring. Monogr.2, p. 109. 8. cesia, Hort— China. 10. S. hortensis, Metten. Fil. Hort. Lips.p. 128. §. denticu- lata, Hort.—South of Europe. 11. S. sarmentosa, A. Braun. S. patula, Spring. S. apo- thecia, Hort.— West Indies. 12, 8. Poppigiana, Spring. ; Hook. Fil. Exot. t.56.—Tropical erica. 13. 14, 16. 16. i, 18. 19, 20. 21. 22, 23. AN ENUMERATION OF CULTIVATED FERNS. 279 8. stenophylla, 4. Braun. Revise, No. 35. 8. micro phylla, Hort.—Mexico. tt Stems sub-erect. Y S. Martensii, Spring. Monogr. 2, p. 129. Lycopodium stoloniferum, Link. Lycopodium Brasiliense, Hort— Mexico. S. Breynii, Spring. Monogr. 2,p. 119. 8S. Panamensis, Hort. 8. Peppigiana, Hort——Guiana. 8. Galeottii, Spring. Monogr. 2, p. 220. Lycopodium stoloniferum, Mart, e Gal. S. Sehottii, Hort— Mexico, S. sulcata, Spring. Monogr. 2, p. 214. Lycopodium sulcatum, Desv.—Columbia. 8, atroviridis, Spring. Monogr. 2, p. 124. Lycopodium atvogidiae, Wall. ; Hook. et Grev. Ic. Fil. t. 839.—East Indies. S. inequalifolia, Spring. Monogr. 2,p.148. Lycopodium ineequalifolium, Hook. et Grev.—Hast Indies and Java, 8. ciliata, A. Braun. Revise, No.14. Lycopodium ciliatum, Willd.—Tropical America. 8. Griffithii, Spring. ; Veitch. Cat. (1861).—Borneo., ttt Stems scandent. S. levigata, Spring. Monogr. 2, p. 1387. Lycopodium levigatum, Willd. Lycopodium Willdenovii, Desv. Selaginella ceesia, var. arborea, Hort. S&S. altissima, Klot.—Kast Indies. § 2. Plants caulescent. S. caulescens, Spring. Monogr. 2, p. 158. Lycopodium caulescens, Wail. var. minor, Veitch. Cat. (1861). — East Indies. . §. erythropus, Spring. Monogr. 2, p. 156. Lycopodium erythropus, Mart, Ic. Sel. Pl. Crypt. t. 20, f. 3.— Tropical America. . S. viticulosa, Klot.; Spring. Monogr. 2, p. 186. — Columbia. . 8. flabellata, Spring. Monogr. 2, p. 174. Lycopodium flabellatum, Zinn.—Columbia and Peru. 280 FERNS: BRITISH AND FOREIGN. 27, &. filicina, Spring. Monogr. 2, p.189. §.dichrous, Hort. —Columbia and Peru. 28, S. Africana, A. Braun. Revise, No. 23. S. Vogelii, Spring. —Fernando Po. 29. S. pubescens, Spring. Monogr. 2, p.173. Lycopodium pubescens, Wall. S. Willdenovii, Hort—Hast Indies. 30. S. Lyallii, Spring. Monogr. 2, p. 168. Lycopodium Lyallii, Hook. et Grev—Madagascar. 31. 8. Lobbii, Hort. ; A. Braun. Revise, App. No. 26; Veitch. Cat. (1861).—Borneo. 32. 8. Wallichii, Hort.; Veitch. Cat. (1861).—Penang. § 3. Plants rosulate. 33. 8. cuspidata, Link; Spring. Monogr. 2, p. 66. S. pal- lescens, Klot. S. circinalis, Hort—Tropical America. 34. S. convoluta, Spring. Monogr. 2, p. 69. Lycopodium convolutum, Walk. and Arnott. Lycopodium para- doxa, Hort.—Tropical America. 35. S. involvens, Spring. Monogr. 2, p. 63. ‘Lycopodium involvens, Si.—India. 36. S. lepidophylla, Spring. Monogr. 2, p. 27. Lycopodium lepidophyllum, Hook. et Grev.—Mexico. Onver V—MARSILEACE ZB, Plants growing in, or floating in water (rarely not in water), very varicus in character, consisting of grass or trefoil-like leaves, or branched with imbricated leaves, bearing sporangia (conceptacles*) at their roots, or base, or on the footstalks, or on small branchlets on the under side, which are 1-4, or many- ceited. 161. MARSILEA, Linn. Plants with quadrifid (trefoil-like) leaves on along footstalk, from a rhizome growing under water. Sporangia (conceptacles) borne on the rhizome at the base of the footstalk, or pedicellate * A hollow case containing spore-cases; a compound sporangium AN ENUMERATION OF CULTIVATED FERNS. 281 on the footstalk, 2-valved, containing numerous obovate recep- tacles in two longitudiual series, bearing cellular vesicles of Genus 161.—Plant with spore-cates, natural size. No. 1. two kinds — 1. (Antheridangia), containing numerous small spores; 2. (Oophoridangia), containing a single large spore. 1. M, quadrifolia, Linn.; Willd. Sp. Pl. 5, p. 588; Schk. Crypt. t. 173.— Germany. 2. M. macropus, Hook. Ic. Pl. t. 909; Gard. Ferns, t. 63; Seemann, Journ. Bot. 1, p. 6. (The Nardoo plant of Australian explorers.)— Australia. 282 FERNS: BRITISH AND FOREIGN. 162. PILULARIA, Linn. Plants with filiform leaves, from a creeping rhizome under water. Sporangia (conceptacles) radical (at the base of the leaves on the rhizome), globose, coriaceous, 2—4-celled, 2-4-valved, each Genus 162.—Plant, natural size. No.1. cell containing different kinds of bodies :—1. (Antheridangia), consisting of vesicles containing many minute granular spores ; 2. (Oophoridangia), each containing a single large spore. 1. P. globulifera, Linn.; Willd. Sp. Pl. 5, p. 535; Bolt. Fil. t. 40; Schk. Crypt. t.173; Hng. Bot, ¢, 521. — Europe and Britain. 163. ISOETES, Linn. Plants with awl-shaped leaves, from a thick creeping rhizome under water or out of water. Sporangia (conceptacles) borne in the AN ENUMERATION OF CULTIVATED FERNS. 283 axis of the leaves at their base, globose, 1-celled, traversed by thread-like receptacles of two kinds: —1. (Antheridangia), containing numerous small spores; 2. (Oophoridangia), con- atining large 4-sided spores. Genus 163.—Plant, uatural size. No.1. 1. L. lacustris, Linn.; Willd. Sp. Pl. 5, p. 584; Bolt, Fil, t.41; Schk. Crypt. t.173; Eng. Bot. 1084.— Europe, Britain, 284 ADDENDA. —+eee A CONSIDERABLE time having elapsed since the pre- ceding Enumeration was prepared and sent to the press, has enabled me to note a number of recently introduced, and a few omitted older species, which I now enumerate in the form of addenda. N.B.—Those marked thus + after the name of the country are entered on the authority of Mr. T. Moore’s notices of them, in the Proceedings of the Royal Horticultural Society and the Gardeners’ Chronicle newspaper ; not having myself seen them alive or obtained specimens. 38. LOMARIOPSIS. 2*, L. fraxinea, J. Sm. Lomana fraxinea, Willd. Acro- stichum (Lomariopsis) sorbifolium, Hook. Sp. Fil. 5, p. 241 (non J. Sm.).—Mauritius. Oxss.—This is a very distinct plant from the West Indian type of L. sorbifolia. 50. GYMNOGRAMMA. § 7. Ampelogramma, J. Sm. Vernation sarmentose, according to Hook. ; fronds indefinite ; rachis flexuose ; pinne bi-tripinnate, refracted ; pinnules small, cuneiform. 18. G. flexuosa, Desv.; Hook. Sp. Fil. 5, p. 192. G. retro- fracta, Hook. et Grev. Bot. Misc. 8, ¢, 112.—Tropical America, AN ENUMERATION OF CULTIVATED FERNS. 285 62, DICTYOPTERIS. 2. D. macrodonta, Presl, Tein. Pterid.; J. Sin. Gen. Fil. Philipp. ; Hook. Journ. Bot. 3, p. 396. Polypodium macrodon, Reinw. in Herb. J. Sm. Aspidium difforme, Blume, accord. to Reinw. in Herb. J. Sm. Polypo- dium confluens, Wall.—East Indies, Malayan Archi- pelago, Fiji. Oss.—In Herbariums and books there is great confusion in the synonymy of what, according to Garden plants, seem to be two distinct species ; — viz., Dictyopteris wregularis and D. macrodon of Presl; judging from Herbarium specimens, they are difficult to be recognized as distinct ; but living plants show the first to have erect vernation, the other decumbent. The Kew Collection is indebted for this species, as also the beautiful Mécrolepia platyphylla, and others, to Mr. Robert Kennedy, Florist and Fern-dealer in Covent Garden. 3. D. Cameroonianis, J. Sm. Polypodium (Dictyopteris) Cameroonianum, Hook. Sp. Fil. 5, p.104. Dictyopteris varians, Moore, in Gard. Chron. (1864).— Tropical West Africa. Oxzs.—A plant of this Fern was sent some time ago from Old Calabar to the Royal Botanic Gardengdiginburgh, and I am indebted for a specimen of it to Mr. James McNab, the Curator of that truly scientific garden. 63. MENISCIUM. 3*, M. angustifolium, Willd.; Hook. Sp. Fil. 5, p. 164.— Tropical America, 65. NEPHRODIUM. 4*, N. cyatheoides, Kaulf.; Hook. Sp. Fil. 4, 4.241. Poly- stichum Dubreuillianum, Gaud. in Freyc. Voy. Bot. Crypt. t. 9 —Sandwich Islands. Ozs.—It is but right to observe that this remarkable species, as also the Sandwich Island Ferns in this addenda, with the previously entered Phegopteris wridentata, and the remarkable 286 FERNS : BRITISH AND, FOREIGN. Colysis Spectrum, also the beautiful Cibotium Menziesit, were sent to the Royal Gardens, Kew, from the Sandwich Islands by Dr. Hillebrand, in 1863. 68. CYRTOMIUM. 1*, C. Fortunei, J. Sm. Fronds 1-14 foot long, pinnate ; pinne lanceolate, faleate, acuminate, 2-3 inches long, 1 inch wide, entire, the base oblique, the inferior rounded, the superior truncate and subauriculate. Costa ebenous, upper surface dull, unreflecting. Sort numerous.—Japan. Oxs.—In Herbaria, specimens of this cannot readily be dis- tinguished from C. falcatum ; but on seeing living plants stand- ing side by side, the difference is evident ; C. Fortunei being a smaller-growing plant, thinner in texture, and not lucid and reflecting light, as C. falcatum. 70. ASPIDIUM. 3*, A. polymorphum, Wall. ; Hook. Sp. Fil. 4, p. 54 (ewelud, syn.).—India, Ceylon. 3**, A. Barteri, J. Sm. Vernation erect. Fronds pinnate, 1-2 feet high; pinnz linear-lanceolate, 6-7 inches long, 1-13 broad; the lower pair sometimes binate, or auriculated on the lower margin. Sori regularly biserial between the primary veins. Indusiwm small, fugaceous. Aspidium polymorphum, Wall. ac- cording to Hook. Sp. Fil. 4, p. 54.—West Africa, Fernando Po. Oxzs.—This is quite distinct from the preceding species. In A. Barteri the sort are perfectly serial, whereas in A. poly- morphum the sort are scattered. ‘ 74, POLYSTICHUM. 6*. P. lepidocaulon, J.8m. Aspidium lepidocaulon, Hook. Sp. Fil. 4, t. 217—Japan. AN ENUMERATION OF CULTIVATED FERNS. 287 Oxzs.—On receiving this plant, it accorded so well with the Indian specimen of P. obliguum, which led to that name being inserted at p. 149, but the plant grew out of its imported form ; and if the Indian plant (which has not yet been introduced) should assume the character of this by cultivation, then this name must become a synonym. The introduction of this species is due to Mr. Richard Oldham, the last special collector of plants sent out from Kew, who, after remaining three years in Japan, visited Formosa, where his health failed. He returned to Amoy, and there died in November, 1864. 10*. P. coneavum, Moore, Proc. Hort. Soc. 11, 377. Lastrea Standishii, Hort.—Japan.t 10**, P. ordinatum, Fée ; Moore, Proc. Hort. Soc. 11, 367.— Tropical America.+ 75, LASTREA. 29*, L. spectabilis, J. Sm. in Enum. Fil. Philipp. Hook. Journ. Bot. 8 (1841). Aspidium spectabile, Blume.—Philip- pine and Malayan Islands, India. 41*, L. sparsa, Moore. Nephrodium sparsum, Don. As- pidium purpurascens, Blume, according to Hook. Sp. Fil. 8, p. 188, t. 262.—Throughout India, Java, and Ceylon. 41**, L. latifrons, J. Sm. Nephrodium (Lastrea) Jatifrons, Hook. Sp. Fil. 4, p. 138.—Sandwich Islands. 41***, I, membranifolia, Presl, Pterid. Nephrodium mem- branifolium, Presl, Relig. Haenk. t. 6, f.3; Hook. Sp. Fil. 4, ¢, 26.—India, Ceylon. 79. NEPHROLEPIS. 3*, N. falciformis, J. Sm. Fronds suberect, linear, pinnate, 14-2 feet in length; pinne: numerous; the sterile (lower ones) elliptical, obtuse, base truncate; the superior ones fertile, lanceolate, falcate, acute, sub- deflexed, 13 inch long by 3 inch wide; base truncate, 288 FERNS : BRITISH AND FOREIGN. subauriculate on the upper margin. ‘Sort uniserial, antimarginal. Indusiwm suborbicular—Borneo. Oxs.—The affinity of this species is with N. tuberosa 3 but seeing the two plants together, the difference is readily seen. 79a. ISOLOMA, J. Sm. Vernation fasciculate, erect, stoloniferous. Fronds linear, 1-2 feet long, pinnate; pinnew oblong, elliptical or lanceolate, falcate, coriaceous, the base truncate and subauriculated; petiole short, articulated with the rachis. Veins forked; sterile Genus 79a.—Portion of sterile frond and a fertile pinna, natura ».. itto enlarged. No. 1. venules free, the fertile transversely combined at the margin, forming a continuous receptacle. Indusiwm linear, interiorly attached, plane, equal with and conniving with the margin, forming with it a vertical exteriorly open groove, containing the sporangia. 1. I, lanuginosa, J. Sm. in Lond. Journ. Bot. 1, p. 420. Lindsea lanuginosa, Wall.; Hook. Sp. Fil. 1, t. 69 B. —Singapore, Penang, New Guinea, and Seychelles Islands, AN ENUMERATION OF CULTIVATED FERNS. 289 Oss.—This remarkable Fern has lately been added to the Kew collection, having been sent from the Mauritius Botanic Garden by Mr. John Horn.* In general habit and appearance this Fern is the prototype of Nephrolepis; but is distinguished by the receptacles being laterally confluent, forming a linear marginal sorus, similar to that of Lindscea, from which it is, however, quite distinct in habit. 81, PHEGOPTERIS. 2*, P. plumosa, J. Sm. Asplenium Felix-foomina, var. plu- mosum, Moore, Nat. Print. Ferns, oct. ed. p. 56; Lowe’s New Ferns, t.14,—Yorkshire (Mr. Stansfield). Ons.—A few years ago three plants of this Fern were found . wild in Yorkshire. It was soon alterwards described and figured in the works above quoted, under the name of Asplenium Felix- fomina,var.plumosum ; but upon what grounds it was referred tc Asplenie I cannot explain, as all the specimens I have examined of ithave small, punctiform, naked sori, perfectly characteristic of the genus Phegopteris, with which it also agrees in habit. This leaves me no other alternative than to consider it a species of that genus, and consequently a new British species. In doing so, the question arises.as to whether it represents an ancient species not before noticed, or the modern result arising from the power of nature to generate new forms, in accordance with the Darwinian Theory of creation of species. It is, how- ever, to be observed that in abnormal or difformed states of Aspleniwm and Scolopendrium, the sori are depauperated, in some instances having no vestige of an indusium; but such is not the case with this plant. The fronds are perfect in every respect, and if Herbarium specimens had been received from some furcisu country, no Pteridologist, on seeing the naked sori, would retur it to Asplenive. 93, ADIANTUM, 35*. A. Gheisbeghtii, Backhouse, Cat. (P) A. tenerum, var. (J. Sm.)—Tropical America.t+ * Who left Kew in 1861 to be assis aut to Mr. Duncan, See page 16 U 290 FERNS : BRITISH AND FOREIGN. 39%. A. colpodes, Moore, Gard. Chron. (1865).—Ecuador.t 44, A. tinctum, Moore, Proc. Hort. Soc. 11, 369.—Tropical America.f 96. PTERIS. 20%. P. straminea, Metten. P. crispa, Hort. (non Linn.).— Chili. 97. LITOBROCHIA. 14*, L. areolata, Moore. Pteris areolata, Lowe’s New Ferns t. 57.—India.t 102. LOMARIA. 2%, L. rigida, J. Sm. Vernation fasciculate, erect, becoming ceespitose. Fronds (the sterile) lanceolate, 8-10 inches long, 13-2 inches broad, erect, rigid, pinnatifid to the rachis; sinus acute; segments alternate, contiguous, lanceolate, falcate, finely serrulate; the lower ones decreasing in size and obtuse. Veins evident. Fertile fronds as broad and rigid as the sterile; segments be- coming involute, densely sporangiferous.— Chatham Islands. 12%, L. Germanii, Hook. Sp. Fil. 8, 1. 152. LL. crenulata, Hort.—Chili, 110. ASPLENIUM. 33%. A. Kaulfussii, Schlecht, Adwm. (29 in obs.). A. protensum, Kaulf. (non Schrad.).—Sandwich Islands. 19*. A. tenerum, Forst.; Schk. Fil. t. 69.—Islands of the ‘Pacific, Ceylon. 92", A. nigripes, DMetten. ; Hook. Sp. Fil. 8, p. 222.—Ceylon. AN ENUMERATION OF CULIIVATED FERNS. 291 127. DICKSONTIA. 6. D. Youngii, Moore, Proc. Hort. Soc.—New South Wales.+ 128. CIBOTIUM. 4, C. regale, Linden; Moore, Gard. Chron. (1864), 414. ~ Mexico.+ CYATHEA. 5*, C. princeps, J. Sm. Cibotium princeps, Linden, Cat.— Mexico. Ozs.—Plants of this Fern were sent to this country under the name of Cibotiwm princeps ; but a plant in Messrs. Lee’s nursery having produced fructification during 1865 proves it to be a Cyathea. Abstract of the number of species, and date of intro- duction :— Exotic species at Kew in 1822... 14. eee oer 40 Do. My Enumeration of Kew Ferns, 1846 ... 355 Do. My Catalogue of Cultivated Ferns, 1857... 559 This Enumeration, Exotic and British ... 1028 Do. Fern Allies, Exotic and British ... 1. .. 56 To.lin 1865... .., ... 1084 In closing this enumeration, I deem it necessary to state that a few species recently introduced to the Kew collection have failed to become esta- blished, after their names were entered on the list; the principal uf which are: Dicranoglossum furcatum, Aconiopteris nervosa, Pleocnemia Leuziana, Adian- tum lunulatum, Lomaria Fraseri, Asplenium lanceum, A. radiatum, Antigramma repanda, Polystichwm anomalum, Loxsoma Ounninghamit, Kaulfussia escu- lefolia. Also the following, entered from my cata- u 2 292 FERNS : BRITISH AND FOREIGN. logue of 1857, were not inthe Kew collection in 1864, viz., Gymnogramma rutcefolia, Hemionites pedata, Notholena lanuginosa, Myriopteris vestita, Onychium auratum, Blechnum triangulare, Nephrolepis undulata, Lindsea guianensis, Schizoloma ensifolia, Thyrsopteris elegans, Botrychium virginicum. Several special causes which lead ‘to the loss of species have already been noticed. I did not intend to advert to them again; but the recent death of Sir W. J. Hooker * necessitates me to repeat what I have stated at pages 42 and 43, where, in speaking of my having resigned the charge of the Kew collec- tion, I said that “happily it remains under the direction of Sir W. J. Hooker.’ These words will now apply to Dr. Hooker, the present Director, whose name is sufficient in itself to sustain the scientific reputation of Kew. Let us hope that the general collection of living plants of the Botanic Garden, now famous for more than a century,t will not suffer by the modern taste for showy flowers, and what is now fashionably called “ foliage plants.” * Died August 12th, 1865. + Number of species at Kew in 1768 .. .. 8,400 Ditto 1786 .. .. 5,500 Ditto 1813 .. 9,800 Since the latter date no general catalogue has been pub- lished, and no public record kept of the plants introduced or lost. 293 ETYMOLOGY. —— ot ANY cultivators and plant amateurs, not versed in scientific literature, complain of the (to them) difficulty to pronounce names given. to plants by botanists. This complaint is, however, more imaginary than real. They have only to recollect that time and use has made them perfectly familiar with such names as Pelargonium, Geranium, Hydran- gea, Calceolaria, Rhododendron,Chrysanthemum, Mesem- bryanthemum, Elscholizia, Fuchsia, and many others now fluently spoken as if they were original words of their mother-tongue. Others say, these long-sounding names would be more readily reconciled to them. if the meaning and bearing upon the plant were ex- plained. To meet this desire, I have drawn up the following table, showing the derivation of the genera of Ferns characterized in the preceding pages. As a general rule, it may be admitted that names of plants are derived from three principal sources. The first, and greater number, being generally a com- pound of two Greek words. The second, names of persons, with the addition of a Latin termination. The third source is various; such as the adoption of local aboriginal names,names from Heathen Mythology, and 294 FERNS: BRITISH AND FOREIGN. often words of doubtful origin, and unmeaning appli- cation to the plants, have been given as generic names by even the most learned botanists. N.B.—For all names ending and beginning with Pteris, see Pteris. Aconiopteris, akon, a point, and pteris; the veins forming sharp angles close to the margin of the frond in the typical species. Acrostichum, akros, highest, stichos, order ; the fructification at the top of the fronds. Adiantum, adiantos, dry; when plunged in water comes out dry. Alsophila, alsos, grove, phileo, to love; grows in groves and shady places. Amphidesmium, amphi, around, or on both sides, desmos, a band; relating to the position of the sori to one another. Anapeltis, ano, without, peltis, shield; the sori naked, in oppo- sition to Pleopeltis. Anchistea, agnati, kindred, intermediate between Woodwardia and Doodia. Anemia, aneimon, naked; naked spikes of fructification. Anemidictyon, veins reticulated ; not free as in Anemia. Anetium, probably from the Greek anaitios, guiltless. This is perhaps meant to indicate that A. citrifolium had been put wrongfully to Acrostichum. Angiopteris, angio, open, pteris; the open sporangia. Antigramma, anti, opposite, gramma, a line; the linear sori being in pairs opposite to each other. Autrophyum, antrum, a den or hollow; the receptacles of sporangia being in a grove or channel. Arthropteris, arthron, joint, pteris; the frond and pinnss being articulate. Aspidium, aspidos; the indusium being like a buckler or shield. Asplenium, a, privative, splen, the spleen; medical qualities. Balantium, balantion, a purse or bag; the form of the indu- sium. Blechnum, one of the Greek names for a fern. Botrychium, botrys, bunch or raceme; the fructification like a bunch of grapes. ETYMOLOGY. 295 Brainea, in honour of C. Braine, Esq., who introduced the first plant. Callipteris, kallos, beautiful, and pteris, beautiful fern. Campyloneurum, kamptos, arched, newron, a nerve; the veins forming arches. Ceratopteris, keras, keratos, a horn, pieris; horned fern. , Ceterach, a name given by Persian physicians. Cheilanthes, chedlos, lip or margin, anthos, flower; the fructifi- cation on the margin. . Cibotium, kibotos, a casket or coffer; the form of the indusium. Cincinalis, meaning unknown. Colysis, kolysis, separation, a species separated from other genera. Coniogramma, konis, dust, gramma, a line; imperfect sporangia, but not applicable in perfect specimens. Cryptogramma, kiyptos, hidden, gramma, a line or writing ; the lines of fructitication being hidden. Cyathea, kyathos, a cup; the form of indusium. Cyclodium, kyklos, circle; the form of indusium. ‘Cyclopeltis, kyklos, circular, peléis, shield; the form of indu- sium, ‘Cystopteris, kystos, a bladder; the inflated indusium, Danza, in. honour of Pierre Martin Dana, a Piedmont botanist. Davallia, in honour.of Edmond Davall, a Swiss botanist. Dicksonia, in honour of James Dickson, a British crypto- gamist. Dicranoglossum, dikranos, forked, glossa, tongue; the fronds forked and pendulous. . Dictymia, diktyon, a net; the netted venation. ' Dictyogramma, diktyon, gramma, a line; the sori reticulated. Dictyopteris, diktyon, pteris; the veins reticulated. Dictyoxiphium, diktyon, wiphion, sword; the veins reticulated, and the form of the frond. Didymochlena, didynos, two or double, chlena, a cloak; the indusium being double. Diplazium, diplazo, to be double; two indusia on the same receptacle. Doodia, in honour of Samuel Doody, a London apothecary, and _ British cryptogamist. 296 FERNS: BRITISH AND FOREIGN. Doryopteris, dory, spear or halbert, pteris ; form of the fronds. Drymoglossum, drymos, wood, glossa, tongue; the fronds like tongues on trees. Drynaria, dryads; the sterile fronds being like oak leaves, which tree was sacred to the Dryads. Dryomenis, dryos, dryads (as above), meniskos, a crescent ; shape of the sori. Egenolfia, in honour of Christian Egenolph, a German author of a book on herbs. ; ; Elaphoglossum, elaphos, a stag, glossa, tongue; the fronds being like the stag’s tongue. Eupodium, eu, good, pous, podos, a foot; the sorus having a foot-stalk. Fadyenia, in honour of Dr. M‘Fadyen, an eminent physician in Jamaica. Feea, in honour of Mons. A. L. A. Fée, Professor of Botany at Strasburg, a celebrated writer on ferns. Gleichenia, in honour of Baron P. F.von Gleichen, a German botanist. Goniophlebium, gonia, angle, phlebes, veins; the veins meeting, forming angles. Goniopteris, gonia, angle, pteris ; the veins meeting, forming angles, Grammitis, gramma, a line; the sori being linear. Gymnogramma, gymnos, naked, gramma, Bite the sori on the veins in lines, and naked. Gymnopteris, gymnos, naked, pteris; the fructification naked. Haplopteris, haploos, simple, pteris ; simple venation. Helminthosiachys, helminthos, worm, stachys, a spike; the fructification in compact spikes, worm-like. Hemidictyum, hemi, half, diktyon, a net ; the outer portion only ‘of the veins being reticulated. Hemionites, a name given by ancient botanists to a plant now called Aspleniwm hemionites, but retained as a generic name fora West Indian fern by Linnzeus. Humata, humatus, humid; in opposition to Adiantum. Hymenodium, hymen, membrane; the character of the frond. Hymenolepis, hymen, membrane, lepis, a scale; membraneous scales covering the fructification. ETYMOLOGY. : 297 Hymenophyllum, hymen, membrane, phyllon, leaf; “ Filmy- leaf Fern.” Hymenostachys, hymen, membrane, stachys, a spike; the firuc- tification on spikes. Hypoderris, hypo, under, derma, skin; the indusium under the sori, like a thin skin. Typolepis, hypo, under, lepis, a scale; the sporangia under the indusium. Isoetes, isos, equal, etes, the year; remaining the same through- out the year. Isoloma, isos, equal, loma, a border or margin; the indusium and margin equal. Kaulfussia, in honour of D. G. F. Kaulfuss, of Halle, a celee brated writer on ferns. Lastrea, in compliment to Chev. de Lastre, a French nobleman. Lepicistis, lepis, scale, kistis, a cyst or cell; the sori being im- mersed in scales, which form a cyst. Leptochilus, leptos, slender, cheilos, lip ; narrow indusium. Leptogramma, leptos, slender, gramma, a line; short linear sori. Leucostegia, leukos, white, stegos,a cover; the indusium being pale-coloured, approaching white. Lindsza, in honour of Dr. Lindsay, of Jamaica, a writer on the germination of mosses and ferns. Lithobrochia, lithos, a stone, brocha, spots; the areoles of the reticulated veins like pavement. Llavea, named by Lagasca after a Mexican injcanen Lomaria, loma, a fringe or border; relating to the indusium, Lomariopsis, lomaria, opsis, like; like Lomaria, Lonchitis, logche, a lance; form of pinnz. Lopholepis, lophos, a crest, lepis, scale ; the sori being furnished with a tuft of slender scales. Lophosoria, lophos, crest or tuft; sori farnished with hairs like a tuft. Lorinseria, in honour of Gustave Lorinser, a Bohemian physician. Loxsoma, loxos, oblique, soma, band; the oblique broad ring of the sporangium. Lycopodium, lykos, wolf, pous ; foot-resemblance, Lygodium, lygodes, flexible ; climbing plant. 298 FERNS: BRITISO AND. FOREIGN. Marattia, in honour of J. F. Maratti, of Tuscany, who wrote on ferns. Marsilea, in honour of Count L. F. Marsigli, of Bologna. Meniscium, meniskos, crescent ; shape of the fructification. Mesochleena, mesos, middle, chlena, a cloak ; attachment of the indnsium. Microlepia, mikos, small, lepis, scale; the indusium small. Microsorum, mikros, small; small sori. Microstaphyla, mikros, small, staphyle, a bunch; the fertile frond being like a little bunch. Mohria, in honour of D. D. Mohr, a German botanist. Myriopteris, myrios, myriad, pteris; the frond being divided into a number of small parts, Neottopteris, nevitia, nest, pteris; bird’s-nest fern. Nephrodium, nephros, kidney ; form of the indusium. Nephrolepis, xephros, kidney, lepis, scale; the indusium being kidney-shaped and scale-like. Neurocallis, newon, a nerve, kallos, beautiful; the venation. Neurodium, neuron, nerve; the venation. Niphobolus, niphes, of snow, bolos, a large pill; the under side of the frond densely covered with white scales, snow-like and the round sori. Niphopsis, niphos, of snow, opsis, like; like niphobolus. Notholana, nothos, spurious, chlena, cloak; the imperfect indusium. Ochropteris, ochros, pale; pale fern. Odontosoria, odontos, of a tooth, sori; the sori-like teeth. Oleandra, resemblance to Nerium Oleander; the Oleander. Olfersia, in honour of Professor Olfers, a celebrated astro- nomer. Onoclea, onos, a vessel, kleio, to enclose; the rolled-up fertile segments of the frond. Onychium, onychion,.a little nail; resemblance to the fertile segments of the fronds. Ophioglossom, ophios, of a serpent, glossa, tengues ; the spikes of fructification. Osmunda, Osmunder, one of the names of Thor, a Celtic divinity. Paragramma, para, near to, gramma, a line; the sori in short lines parallel with and close to the margin. Pellaa, pellos, dark-coloured; the dusky colour of the fronds. Phegopteris, phegos, beech ; beech fern, by some called sun fern. ETYMOLOGY. 299 Phlebodium, phiebes, veins, odous, a tooth; the joining of the veins in the areoles being like teeth. Phymatodes, plymata, tubercles ; the impressed sori having -the appearance of tubercles on the upper side of the frond. Pilularia, pilula, a pill; the form of the spore-cases. Platycerium, platys, broad, keras, horn; the fronds divided in broad segments like stags’ hors. (The stags’-horn fern.) Platyloma, platys, broad, loma, border ; oe broad sori close to the margin. Pleocnemia, pleos, full, knemia, rays; full of rays; the vena- tion. Pleopeltis, pleos full, peltis, shield; the sori being furnished with numerous round scales. Pleuridium, pleura, side (ribs); the primary veins being coste- form, like ribs. Peecilopteris, pelilos, spotted, pteris; the appearance of the venation. Polybotrya, polys, many, botrys, bunch ; hs fructification being in bunch-like racemes. Polypodium, polys, many, pous, foot; polypus; the rhizome, ' when destitute of the fronds, having the appearance of some kind of sea-polypus. Polystichum, olys, many, stichos, order; not specially applicable to the genus Polystichum, as now defined. Psilotum, psilos, naked; destitute of leaves. Psomiocarpa, psomion, a small pellet, karpa, fruit; the sporangia being in small round patches, in spikes. Pteris, pteryx, wing; the scientific name given to the plant known by the names of Brake, Bracken, and Fern; on account of the supposed likeness of the branching of its fronds to wings. This, being the commonest ot all Ferns, has become the type “of the whole race; hence Pteris means Fern, all generic names of ferns ending in pteris; such as Ceratopteris means horned Fern, Diotyopteris netted Fern; also, if at the beginning of a word, as Pteridography, a book or writing on Ferns ; Pieridologist, a studier and writer on Ferns; Péeridophilist, a lover of Ferns. Rhipidopteris, rhipis, fan; shape of the fronds like a fan. 300 FERNS: BRITISH AND FOREIGN. Saccoloma, sakkos, a bag or sack, loma, margin; the union of the indusium with the margin, forming a hood or cyst. Salpichlena, salpi, pipe, chlena, a cloak; the indusium being like a pipe. Schellolepis, schello, skeleton, lepis, scale; the character of the scale surrounding the sori. Schizoloma, schizo, I cut, loma, a border or margin; the sori in the form of a slit on the margin of the frond. Scolopendrium, skolopendra, a centipede; the appearance of fructification on the under side of the frond. Selaginella, the diminutive of selago (Lycopodiwm Selago). Selliguea, in memory of M. Selligue, a French optician. _ Sitibolium,* sitos, food, bolos, a large pill; the appearance of fructification. Soromanes, soros, heap, mania, fancy; the sporangia in fanciful and irregular heaps, ; Stenochlena, stenos, narrow, chlena, cloak; narrow indusium. Stenosemia, stenos, narrow, sema, standard; the fertile frond with its narrow segments being like a flag. Struthiopteris, struthios, ostrich ; the fronds being like the feathers of an ostrich. Thyrsopteris, thyrsos, bunch or raceme, pteris ; the fructification in racemose bunches. Todea, in honour of Henry Julius Tode, of Mecklenburg, an experienced mycologist. -Trichocarpa, thria, trichos, a hair, karpa, fruit ; the fructification borne on a hair-like stalk. Trichomanes, thriz, hair, manos, soft; the delicate nature of the fronds, : Trichopteris, thrix, hair, pteris; the sori being furnished with hairs. Vittaria, vitta, riband; shape of narrow fronds. Woodsia, in honour of Joseph Woods, a celebrated British botanist (died 1864), Woodwardia, in honour of Thomas Jenkinson Woodward, an English botanist. Xiphopteris, wiphos, sword, pteris ; form of the frond. * For Sitilobium, at page 236 and elsewhere in this work, read Sitidolium. 301 CULTIVATION, —1o+— 1. Preztiminary Remarks. AVING given an account of how the Exotic Ferns of the preceding enumeration have been introduced to this country, I shall now proceed to state the best means for growing and preserving them in our collections; and as the successful eultivation of Ferns depends much upon a knowledge of the con- ditions under which they grow in their native country, I begin with a few observations on that point. "Ferns have already been spoken of as favourites with the plant-loving public; but it is not simply on heir merits as pretty and interesting objects that they claim attention. Geology reveals to us that Coal— that source of our domestic comforts and national greatness—is formed chiefly of Ferns, which at some remote period grew upon the earth. However, this is not the place to discuss the views and speculative theories regarding the thermal and gaseous condition of the earth and atmosphere, under which Ferns then flourished, or to reason on the manner by which they were converted into coal. I may simply remark that fossil remains show, that contemporaneous with Ferns grew plants of remarkable character, quite 302 FERNS: BRITISH AND FOREIGN. distinct from those of the present era. Not so the Ferns, for their beautiful and well-preserved remains show that their fronds were of various sizes and forms, in every way analogous to the present race; having free and anastomosing venation, round and linear fructification, and, in some instances, almost identical with species now living. At present, Ferns rank amongst the widest spread of all the orders of the vegetable kingdom, being found in more or less number in all climates, between the most northern and southern limits of vegetable life, and at elevations ranging from the sea-level to 14-15,000 feet within the tropics, their number in any localities being generally in proportion to the degree of atmospheric moisture in conjunction with elevation, the latter applying specially to the interior of continents. Comparatively few spczies are found in open, grassy, thinly-wooded countries, whether it be the plain or mountain-slope ; such districts are often in full possession of the most gregarious and abundant of all Ferns, the common Brake (Pteris aquilina), which, under slightly different forms, and in some countries accompanied by different species of Gileichenia, occupy vast tracts of the earth’s surface. In hot and moist plains, in valleys of great extent, the number of different species are few; even in the valley of the Amazon, teeming as it does with vegetable life, the number of Ferns found by Dr. Spruce after he left the coast Flora, at Para, in his journey of 2,000 miles, were very few. They became moré numerous on attaining an elevation of 1,500 feet, and in one locality, at a higher elevation, he found 250 species in a diameter of fifty miles. Another extensive tract with but few Ferns is the CULTIVATION. 303 dry zone of Northern Africa, and few have as yet been recorded from the interior of Australia; while, on the contrary, on elevated coast-ridges and islands, they form a large proportion of the entire Flora. They are the most numerous, both in regard to genera and species, in the tropical regions, where, too, a greater number of individual plants are to be found than in temperate regions.* On reviewing the above, it may readily be supposed that the varied influences under which Exotic Ferns naturally grow, necessitate various kinds of treatment for cultivating and preserving them in this country ; but such is not the case. As might be expected, our native representatives (although only forty-two in number) have received special attention. We have Fern Tourists in plenty; almost every spot of our country, where are conditions congenial to their development, has become more or less familiar to those who take an interest in their study—in the damp shady glen, the wildest wood, banks of streams, mountain-sides, whether it be in their sheltered easy slopes, or in the almost inaccessible craggy steeps. Indeed, so diligently have the Fern explorings been carried on, that numerous curious and beautiful varieties, highly prized in our gardens, have been found. But, however laudable and agreeable Fern- growing may be, yet it is to be regretted that it leads to the extinction of some of our rarest native species. Even the more common are becoming scarce in locali- ties within easy reach; great quantities being yearly * Want of space prevents me entering more fully into the history of geographical distribution. a0 FERNS: BRITISH AND FOREIGN. consigned to the London markets. Since Ferns have become so popular, those who patronize horticulture, from the members of the upper classes who construct large hothouses, to the humbler patron, or the lard- working mechanic who prides himself on his posses- sion of a Wardian case, are desirous to obtain such information as will enable them to cultivate them successfully, either in the Conservatory, Wardian case, or Rockwork in the open air. To supply this, several books on the subject have been published. Indeed, if an example- may be taken, besides the extraordinary degree of familiarity so rapidly attained in the knowledge of Ferns by all classes, the constant increase of collections gives sufficient evidence. Nevertheless, in publications which, through their cheapness, are within the reach of every Fern lover, frequent complaints are met with, tending to show that the result is not always satisfactory to the culti- vator, especially among amateurs and persons of small means. These complaints are mainly attri- butable to the author not familiarizing his readers sufficiently with the habits of Ferns, the conditions under which they luxuriate in their native localities, and the necessity of imitating those conditions as far as practicable. It is, however, gratifying to observe that, in the progress of horticulture, the knowledge of the natural conditions of plants is more and more sought afte, and appreciated with very good results, forming a lively contrast with the old times, when the Trichomanes radicans could only with difficulty be kept alive even by the most eminent horticulturists ; a fact observed by the originator and promoter of the Wardian case system, under which this shade and CULTIVATION. 305 moisture-loving Fern stands pre-eminent,-even sur- passing in size and luxuriance those in their native retreats. A familiar example of opposite nature to the preceding is our native species Asplenium septen- trionale, confined to northern localities, generally growing on rocks and insinuating its delicate roots in the deep crevices and among the débris of irregular shelving and prominences, not receiving any apparent injury from the rigour of winter or the scorching heat of summer. Although such is its hardy nature, it nevertheless does not flourish when taken from its native rocks and brought under artificial cultivation. What has been stated regarding the shade and moisture-loving Trichomanes, and the rock-loving Asplenia, may be viewed as the extremes in the nature of those Ferns that do not readily conform to ordinary cultivation; but, on the other hand, the greater number are not particular in their choice of place of growth, either in a wild state or under cultivation. Examples may be cited of species conforming to the most untoward and varying influences; for instance, the common hart’s-tongue Fern (Seolopendriumvulgare) maintains itself in situations of the most opposite kind, having great predilection for the works of man, whether elevated in the air or sunk below the surface of the earth; such as stone, brick, or turf walls, embankments, hedge-banks and road-sides, pits, quarries, or deep open wells. In either place it multi- plies freely, conforming itself to the various atmo- spheric changes ta which those situations are liable. Such being the case, it is not surprising to find it assume different sizes and forms; in dry places it is only a few inches in height, while in open wells, such x 306 FERNS: BRITISH AND FOREIGN. as may be seen in some of the nurseries and market- gardens near London, it produces fronds 2-3 feet in length. The Asplenium Trichomanes, A. Ruta-muraria, Ceterach oficinarum, and Polypodium vulgare, are also wall and tombstone-lovers, and may be called our domestic Ferns. Another remarkable example of a Fern making itself at home under extremes of temperature and moisture, is Pteris longifolia, a species having a wide range throughout the tropical and sub-tropical regions of both hemispheres. On the island of Ischia (Bay of Naples) it is found luxuriating within the influence of the hot vapours rising out of the cavities left by extinct volcanoes, growing in soft muddy soil at a temperature ranging from 140° to 160°. In our hothouses its spores vegetate abundantly upon all moist surfaces, and in the crevices of brick walls. Plants of it are nearly always to be found over the openings of hot-water tanks, and it has been seen in crevices of the walls outstde hothouses, or even under iron gratings, where it could receive but little light, and where the temperature was often near the freezing-point. In the dry air of the Cactus-house plants of it have produced fronds from 2-8 feet in length. In general the fronds of Ferns remain Iong in a perfect state; the exceptions to this rule are com- paratively few, and these are chiefly supplied by the natives of climates alternating with seasons of heat and cold. But as many species are wanderers and conform to th: effects of various clmates, it is no wonder to see some of our native Ferns assuming the evergreen habit of their -foreign allies, when CULTIVATION, 307 grown in a temperate house ; of which Polystichwm aculeatum, Lastrea dilatata, and L. Filia-mas may be cited as examples, as also Asplenium marinum. In 1820 I found plants of it, having fronds from 2-4 inches in length, growing in a cave facing the German Ocean, on the east coast of Scotland ; of these, two plants have been grown at Kew from that time; of late years, one in the Temperate and the other in the Tropical House. These became fine cespitose plants, with fronds varying from 1-1} foot in length; the greatest length being attained by the plant in the Tropical House, even assuming the character of a species native of the West Indies and Tropical America, and quite unlike the original plant; thus showing that although at home in the cold, sunless cave, it can well appreciate a better fed and warmer abode. Exceptions to this rule are some alpine species, Ferns in that respect being analogous to our ill success in growing many alpine flowering plants. This is no doubt partly owing to the difference of atmospheric density, and the varying influences of temperature and moisture common to the sea-level of this climate. The species of cold climates truly deciduous, produce their fronds from an underground creeping sarmentum, of which Péeris aquilina, Sitolobium punctilobium, Onoclea sensibilis, Anchistea virginica, Lorinseria areo- lata, Leucostegia immersa, and Phegopteris aurita are the principal examples. On the other hand, instances of tropical species periodically losing their fronds without any apparent cause, such as by undue excess of heat and moisture, are Phymatodes oxyloba, Pleuridium palmatum, P. venustum, Drynaria propinqua, several x2 308 FERNS: BRITISH AND FOREIGN. species of Davallia, and a few others. Their rhizomes remain perfectly naked during the winter, and seem to be constituted for a season of dry rest. These obversations are sufficient to show how certain species will grow under the most opposite conditions, and that a knowledge of them materially assists our efforts of cultivation, which I shall pro- ceed to treat of under two heads,—special and natural, II. Specrar on Pot Corrtrvatton. ITH the exception of those species sufficiently hardy to bear the cold of this climate, the whole family of Ferns can be artificially and most luxuriantly grown, under but two different scales of temperature, and with as little difference in other respects. There can be no better instance than that afforded by the immense collection nowat Kew,* where, in the Tropical Fern-house, a great many species from various climates are placed under an average tempera- ture of 60° to 70°. In this houset are species from nearly all the tropical and sub-tropical countries of the world, and, being in one compartment, they are all alike subject to the same amount of atmo- spheric moisture as of heat. However, in respect to moisture supplied to their roots, the amount is varied according to the nature and require- ments of the plant. The same remarks apply to the Temperate House, of course with a reduction * May 1864, + Length 130 feet, width 34 feet. CULTIVATION. 809 of moisture as well as temperature, the latter ranging trom 40° to 50° in winter, that being the only period of the year when artificial heat is required. In this house* are growing, in the greatest luxuriance, species from Australia, New Zealand, China, Japan, North India, elevated portions of Ceylon, South Africa, North and South America, and other elevated regions within the tropics. The whole collection is grown in common deep or shallow pots, pans, and tubs; the latter, however, are only used for the large species of Tree Ferns and Angiopterts ; and though such a system is the least natural in appearance, yet it is most convenient and found generally consistent with the prevailing fashion for in-door horticulture; this mode of cultivation, being mostly adopted, merits our first consideration. In the tropical and sub-tropical localities, where heat and moisture are abundant, by the manner in which they grow and the various positions they occupy, soil is of but little importance, except for affording their roots the means of obtaining permanency of position. This is evident by the remains of native soil adhering to the roots of imported plants, showing that Tree and other large Ferns in some places grow in stiff adhesive, red clay: To imitate this soil in pot culture is attended with no success, and it is remarkable to see how quickly the roots of newly-imported plants take to the fine loam and peat in which they are potted. In cultivation, how- ever, the great beneficial influence of the natural atmosphere is not obtained, and the soil is therefore of great importance, necessitating caution in its * Length 82 feet, width 13 feet. 310 FERNS: BRITISH AND FOREIGN. selection. Taking a view of the large number of Ferns now in cultivation, their requirements as regards soil, and mode of growth, we easily recognize two classes—terrestrial and epiphytal. This division is important. For the first class, by far the most ex- tensive, it is necessary to use the soil of a finer or coarser consistence, according to the delicate or stronger character of the plant. In it there is a more varied character of vernation, by which their terrestrial character is indicated and very easily recog- nized. Natural affinity of course is not taken into consideration. However, with tolerable accuracy in this respect, the genera Phegopteris, Lastrea, Ne- phrodium, Adiantum, Asplenium, and Sitolobium give examples of the chief variety of vernation indicative of the terrestrial nature ; viz.,erect, ceespitose, decumbent or sub-hypogeous, or, as in Nephrodium unitum, and N. pteroides, which, although with a true sub-scandent sarmentum, they, however, prefer the firm soil of the terrestrial group, a compost of two-thirds peat and one loam, with abundance of sand, according to the size of the plants for which it is to be used. In potting Ferns, an over-depth of soil should be avoided, as well as a great depth of drainage. The one promotes stagnancy, the other gives an oppor- tunity for the most vigorous roots to descend among its particles, and oftentimes they become too dry when the soil is apparently moist enough, and the tips of the foliage are in consequence injured, and the plant disfigured. This applies more strictly. to species of small and delicate structure, such as Asplenium firmum, A. dentatum, A. decussatum, &c.,and for such the shallow pot is best adapted, the width of CULTIVATION, all which being considerably greater than the depth, gives a good extent of surface, and renders a deep drainage unnecessary. Such a kind of pot, of course, is most suitable for all species of decumbent or creeping habit, and those producing rapidly a great number of offsets. The best time for repotting (or shifting, as it is termed) Ferns, more especially those of the terrestrial division, is at the end of February, or during March, as soon after that time they commence growth; but any time of the year, except winter, will suit them. The operation of potting is a matter of little difficulty, nor are the necessary utensils or materials very varied— a firm potting-bench, several different sizes broad- pointed firming-sticks, and a garden trowel. It is also necessary to have a few extra pots of different sizes, of both the deep upright and shallow flat kinds, also a quantity of material for drainage, which may consist of broken soft brick or hard knobs of old mortar; but for general purposes, potsherds, broken into different sizes, are most commonly used, and well known by the name of “crocks.” The removal of the plant, with its ball of soil and roots, is readily effected by spreading the left hand over the surface of the soil, allowing the plant, when not too large, to come between the fingers; then, by inverting the plant and pot, and giving the rim of the pot a gentle but sharp tap with the right hand on the edge of the board ; if the roots are in a healthy state, the ball will slip easily out of the pot. In all cases, athorough drainage is indispensable, and to ensure this, the draining material should be of two sizes, the larger size at the .botteom, with a concave large crock over 312 FERNS: BRITISH AND FOREIGN. the hole of the pot, the concave side downwards: there should be two, three, or four holes at the side, quite at the base, made in all pots larger than eight inches in diameter, the number of holes of course according to the size of the pot. This is the more essential when the pots are placed on smooth benches, as of stone or slate, the capillary attraction between the pot and bench retaining the superfluous water, and causing much stagnancy. After properly arrang- ing the drainage, a small quantity of fibry material should be placed on it to keep the new soil from mingling with it. No plant should be repotted in a wet state, nor should it be allowed to become dry enough to cause it to flag, as this will not only give an immediate and severe check, but in giving water after it is potted, it will percolate the new soil (which should always be used in a moderately moist state) without entering the dry ball; if, from bad drainage or exhaustion of the soil, it has become impure, so much of it and of the old inactive roots should be removed. In such cases, as small a pot as possible should be used, which will admit the pre- served roots freely, being carefully laid out by the fingers amid the new soil; for if in too large a pot,. the soil becomes sour before any vigorous roots have entered and promoted drainage, as they are tardy to perform their functions vigorously after having been necessarily so much disturbed. However, when in a healthy and vigorous state, such space should be given it that will admit the fingers or both hands freely by its sides (according to the size of the plant) in inserting it in the new pot, where it should be so placed that its crown, if of the CULTIVATION. 313 erect vernation, will be nearly level with the rim. The soil should be carefully introduced, and made firm round the sides by pressure of the firming-stick and a few thumps of the pot (if easily handled) on the potting-bench, if not, the stick must suffice ; a space of about half an inch, more or less, according to the size of the pot, should be allowed at the surface to contain as much water as would moisten the whole ball. The size of the plants will depend upon the amount of space and'the number of species in the collection. For an amateur’s collection in a small house, very fair specimens may be grown in pots from eight to twelve inches in diameter, after the shift into the largest-sized - pot, and with good management the plant will not require any repotting for two or three years. By that time the running and czspitose kinds, such as Adiantum and Gleichenia, will have become exhausted in the centre; the ball will, therefore, require division by passing a sharp knife through it, taking care not to injure the young growths next the sides of the pot; the most healthy portions to be selected for repotting, to become the new representative plant of the collec- tion; and, if proper care is taken, the fronds will suffer but little injury. In operations of this kind, some gardeners entirely shake out the soil, cut away the whole of the fronds and roots, in order to make the plant, they say, come up strong. This may not do much injury to certain plants, such as bulbs, tubers, fuchsias, and such-like plants that rest in winter; but for Ferns it is a great mistake; it so weakens the plant that it takes a year or more to be worth looking at, and, indeed, some never recover. In Adiantum 314 FERNS: BRITISH AND FORKIGN. and other genera, the roots are naturally of a dark, or even quite black colour, and it is known to have been considered by some cultivators, not well versed in the nature of Ferns, as dead, and accordingly theyare totally removed, even although with evidence of the plant being healthy and vigorous. The healthy state of the plant is readily explained, for on examining the nume- rous points of the black roots, each will be seen to be of a pale colour, which are the active feeders of the plant; and, in shifting, great care should be taken not to bruise or injure them. The same directions apply to the gigantic Tree Ferns and species of Angiopteris. The soil for them, however, should be of a coarser nature when of a size to require the largest-sized pots or tubs; in no case is it desirable to give more than three inches extra space at one shift, even in the largest pot or tub. In all pots above one foot in diameter, the drainage should be elevated in the centre. When the shift is large, an inverted pot should be placed in the new pot, placing a layer of drainage round it, the height of the pot to be such that the crocks of the old ball (which must not be taken out) rest on the top of the new drainage or pot. By this the outer circle of bottom roots are not pressed together, or crushed by the weight of the ball, which is sure to be the case when the ball is set on a level surface, it also keeps the ball to its proper height. This mode of drainage has been the constant practice at Kew with all large shifts, not only for Ferns but Proteacee and all fine fibrous-rooted plants, the nature of which is to extend outwards and downwards through the new soil, ultimately forming a web of roots against the sides of the pot. In the ordinary CULTIVATION, 315 practice of potting, it is a rule with many gardeners _ to remove all the old drainage crocks, the centre thus becomes a mass of soil, which in time becomes com- pact, inert, and useless, liable to become stagnant, and in time causing the plant to sicken. This is avoided by preparing and retaining the old drainage. This mode of drainage is, however, only necessary for plants required to be permanent in botanical collec- tions. Its utility in prolonging life is verified by some yet existing plants of Proteacee at Kew, some of which are forty, and others above sixty years of age. What has here been stated may be considered as a general rule for the greater number of the Fern family, such as are usually found in ordinary collec- tions of a hundred species, more or less. But in this extensive family there are many possessing some pecu- liar nature, and a few examples of these require to be specially noticed. The group which has been termed Epiphytal, consists of the genera Davallia, Goniophlebium, Drynaria, Phymatodes, Pleuridium, Platycerium, Lo- mariopsis, Polybotrya, and others of like habit, charac- terized by a true and highly-developed rhizome or sarmentum, generally creeping, and adhering by their fine fibrous roots to the surface on which they grow. Their positions in their natural homes are generally on more or less perpendicular surfaces of moist and shaded places of rocks or ordinary. soil, where decomposing vegetable matter abounds as a surfacing. They are often found on trees, and with many species of Polybotrya and Lomariopsis this seems to be their true position, and often by their vigorous growth, the trees are clothed with them in 316 FERNS: BRITISH AND FOREIGN. the manner of our common ivy. It will be under- stood by this that an open loose soil is essential for the whole group; good fibry peat is therefore all that can be desired, and on account of their creeping habit, as great an extent of its surface as possible is. necessary to be gained in potting. To this end the shallow pan is indispensable; by an ordinary deep pot being inverted in the pan and covered with a layer of peat, of a thickness according to the size of the pan or pot used (at a proportion of two inches thickness for a pan of one foot diameter) carried up over the inverted pot in a conical form, the whole pressed firmly together. The rhizomes are fixed to its surface by pegs, and they will soon attach them- selves by their own roots. The height of the cone will be according to the character of the plants; for Pleopeltis, Anapeltis, Niphobolus, &c., a cone of from 6-10 inches high, with a base from 1 to 1} foot in width, will make handsome specimens. The genus Oleandra comes under Epiphytal, the rhizomes elon- gating rapidly, and in their natural places adhering to trees and moist rocks. 0. neriformis is, however, truly terrestrial, producing roots from its woody erect rhizome after ascending a few inches from the’ soil.. The other species in cultivation, however, cannot be well suited by the conical mode, or being trained on straight sticks. An open netted wire cylinder, about three feet high, and about six inches diameter, answers the purpose exceedingly well. The cylinder is filled with peat, and by fixing the rhizomes round the bottom of the cylinder, the roots will soon adhere to the peat through the meshes, which being kept moist, a rapid growth is stimulated, and the cylinder soon CULTIVATION. 817 becomes furnished with beautiful projecting fronds. Ifa greater height is desired, another cylinder may be added to the first. This mode is equally useful for Stenochlena, Polybotrya, Lomariopsis, &c. In.respect to species in which the rhizomes elongate slowly, the upper part of the cylinder is bare and unsightly for a “ time. This may be obviated by having a short cylinder and heighten it by adding short lengths when required. The genus Elaphoglossum is generally epiphytal, and as some of its larger growing species, such as H. calle- folium and FE. latifolium, being decumbent cespitose in vernation, it is not necessary to have the soil raised much above the rim of the pot, which should be of the flat kind, and three inches depth of soil is quite sufficient. Many smaller species of this genus are most lovely objects, and some of them are not yet in cultivation. Of the whole Fern Family the Platycerium may be considered the most grand, beautiful, and extraordinary; and it is thoroughly typical of this epiphytal group. ‘Its natural position of growth is sometimes on moist rocks, but usually on the trunks and larger branches of trees. The spores becoming lodged there, germinate, and, sending out spongy fibrils, a little plant, like a circular disk, adhering to the tree, analogous to a foliaceous lichen, is formed, each succeeding disk (frond) becoming larger and overlapping the preceding one. In time the older ones loose their vitality, and by this mode of growth envelop, or nearly so, that portion of the tree whereon they grow in a dense, thick, spongy mass, among which the roots insinuate themselves and receive nourishment. As equivalent to this, in cultivation blocks of wood are mostly used ; but they are objectionable from their 318 FERNS: BRITISH AND FOREIGN. constant liability to breed fungi and harbour insects. On that account pots are preferable, not of the ordi- nary shape, but with less difference in the diameter at the base than at the top, and with a wide opening at the side, extending about a third of its diameter from the rim down to within a sixth of its depth from the base (this will leave the pot in much the same form asascoop). . Thorough drainage should be given, and a material of very fibry peat will suit. The crown of the plant should be placed about the centre of the side opening from where the fronds will be produced. In time, the sterile fronds will spread in all directions, but mostly upwards, and quickly obscure the pot, and the fertile ones will hang looselydownwards. Their position in the house should be elevated, and, if possible, against a wall or partition. Platyceriwm alcicorne increasing: rapidly by offsets, requires a’ considerable extent of surface ; consequently rough sandy peat, arranged in a conical manner on a shallow pot, to which the plants will soon attach themselves, is most suitable, and which, if fancy leads, may be suspended from the roof of the house by a-strong wire. For the large-growing species of Drynaria, Gonio- ahlebium, and Phlebodium, it is not necessary to raisé the soil much above the level of the pot; their fleshy rhizomes soon reach the margin, to which they cling, and if standing on a moist surface, or near water, they creep down the sides of the pot. In Davallia pyxidata, D..ornata, and others, the rhizomes are what may be termed aérial, rising con- siderably above the soil, often extending to a distance beyond the edge of the pot, and, as they in these cases do not produce roots, the rhizomes in time CULTIVATION. 319 (three or four years) become weak, producing small fronds ; it therefore becomes necessary to remove the older parts and re-pot the younger portions in fresh soil. Very interesting and natural examples may also be had by placing common cylindrical red chimney- pots or drain-pipes in a pan wider by two inches than the base of the cylinder,—this space to be filled with soil and planted; the rhizomes will soon become attached to the surface of the cylinder and in time will cover the whole, forming a handsome pillar of fronds. In order to maintain a proper. degree of moisture, a pan of water may be fixed inside the cylinder, and if a piece of woollen cloth is placed in the water, in contact with the cylinder, a constant moisture will be kept up congenial to the plants. The top of the cylinder is also useful for placing on it some of the pendulous species; for instance, Phymatodes geminata, Goniophlebium dissimile, G. neriifulium, Campyloneuron angustifolium, Elaphoglos- sum Herminiert. In Scheilolepis subauriculata and S. verrucosa the fronds are long and pendulous; this necessitates the plants being placed in an elevated position, such as suspended from the roof of the house in shallow wire baskets, the inside of which should have a lining of sphagnum moss, which assists to retain moisture, and also gives a clean and neat appearance. Plants of these two species thus treated have at Kew produced fronds 12 feet in length, hanging down in a very graceful manner. There are several other Epiphytes of special interest, such as the species of Vittiria and Haplopteris, which hang down from trees like 320 FERNS: BRITISH AND FOREIGN. bunches of grass, as also the remarkable Ophioglossum pendulum, which may be likened to ribands or bands hanging loose and waving with the wind, often many feet in length. These plants succeed in a small quantity of soil, firmly fixed in pots, with pieces of soft stone or potsherds, and the pot hung against a shady wall or pillar. For this purpose the pot should have a flat back, with the front rim lower than the back, so as to allow the fronds to hang quite free of the pot. Neottopteris Australasica, and a few Aspleniums nearly allied, such as A. sinuatum and A. crenulatum, of precisely the same mode of -growth, are of erect fasciculate vernation. Their roots being of peculiar mossy and delicate nature, they are not adapted for deep insinuation of stiff soil, but are rather what may be termed aérial. Two-thirds of their mass is produced above the surface of the soil. Substantial but open material is therefore required, of very rough, fibry peat, and porous, broken bricks, or soft sandstone, in equal parts; very little pot- room is necessary; a shallow pot of 18 inches diameter, with such material, will support a plant of two dozen fronds, and none less than 8 feet 6 inches long and 8 inches broad, with a stem a foot high, and as much through, principally composed of its mossy roots forming a spongy mass. As an instance of the long life under regular treatment may be cited the original plant of Neottopteris Australasica, which was imported in 1825, and is now (1864) a magnificent plant, in perfect health, having received but few shifts the whole of the forty years. This is, however, far surpassed in size by the mag. CULTIVATION. 321 nificent Neotiopteris muscefolia, which is described by a Penang correspondent in the following words :—“ TI saw two fine specimens of the Birds’-nest Fern ; each had between forty and fifty perfect green leaves; the average length of the leaves was six feet, and from one foot to fourteen inches across in the broadest part. They were growing on each side of a doorway; when I was walking up to them I thought they were American Aloes.” The remarks that have been previously made respecting the nature of the rock-loving Asplenie are equally applicable to others that inhabit rocky places, such as certain species of the genera Notholena, -Myriopteris, Cheilunthes, &c.; some are rocky-coast plants, others are mountain, being found in elevated situations within or near the tropics, where they are subject to the heaviest showers and most powerful sun, their surfaces. being generally furnished with beautiful scales or woolly covering, which resist in a great measure the action of the sun. They all seem very impatient of moisture under cultivation, especially in winter. A material composed of finely-broken and mixed sandstone, bricks, old mortar, and a small quantity of sandy loam, suits them, placed in a position of the coolest shade, with abundance of moisture in summer, and in winter very little water, just sufficient to keep the soil slightly moist; a comparatively dry atmosphere and as much light as may be obtainable at that dull season, with a temperature not below 36°. As a general rule, the above may be considered appli- cable to all small-growing rock species of temperate regions, including the natives of this country. The species of the genus Gymnoyramma, especially those ¥ 322 FERNS: BRITISH AND FOREIGN. with farinose fronds, commonly called Gold and Silver Ferns, are also very susceptible of moisture; they.’ should never be syringed, or water allowed to fall on their fronds, as the: farina, being loose, is distarbed by the water, and running down, gives the appearance as if the plants were smeared with dust. They, however, differ from the preceding, requiring more light, and the temperature of the Tropical House. The species — of Gymnogramma vary very much in habit, as regards size and circumscription of the fronds, G. trifoliata having fronds from three to. four feet high, while in _ G. cherophylla and G. leptophy lla they are fragile, and average from two to six inches in length. These two species are peculiar in being, with the exception. of Ceratopteris thalietroides, the only known truly annual Ferns. G. cherophylla grows freely ; its spores vege- tate abundantly throughout the house, often as a weed. G. leptophylla is, however, not so free in its growth. When its fronds decay,’ the pot should be covered with a piece of glass, and put in a dry place until the proper season arrives in spring, when the application of moisture will cause the latent’ spores to vegetate. G. flewuosa differs from, the whole of the genus in having prelonging fronds, which climb in a rambling manner over bushes, like those of Lygo- dium, noticed in another page: it will probably here- after form the type of a distinct genus. The beautiful genus Lindswa, of which no less than sixty species are described in Hooker’s “ Species Filicum,” are, with few exceptions, natives of the tropics of both hemi- spheres; in my Catalogue of 1857, only two species are recorded as being cultivated; but, within these few years, the number has thigronaed to fourteen, the greater CULTIVATION. 823 part having been introduccd from Guiana by Mr. Backhouse, of York, and on account of their slow increase, plants of them are yet very rare: they are natives of generally the lower regions of tropical vege- tation, growing in open places amongst herbage of small plants and grass, or on the skirts of woods, and sometimes under the shade of trees, the soil being very poor, sometimes almost nothing but sand or stony débris, in which their sarmentums are partially hypogeous, the soil and air never being much below the temperature of 80°, and although almost daily subject to the influence of tropical thunder-showers, yet, on account of the nature of the’soil, the surface is never over saturated. Finding that they do not flourish underthe medium temperature of a Tropical Fern House, _it therefore is necessary that a special part of the house, on the principle of a Wardian Case, should be adapted, so that a moist air of 80° may be steadily maintained, and the plants occasionally sprinkled overhead, taking care that no superfluity remain in the soil, which should be no more than moist. Like Lindscea, the curious and interesting genus -Schizea does not readily conform to cultivation ; plants of Schizcea elegans have often been freely im- -ported from Trinidad, and although tried in various ways in high and moderate temperatures, it cannot be said they have yet become established. Under the Wardian Case, the native imported fronds remain for a considerable time fresh, and sometimes new fronds show themselves, but fail to come to maturity. Ina letter lately received from Mr. Prestoe, in Trinidad, be informs me that the Schizwa elegans grows in solitary patches in loamy soil, covered with three or. iour x2 324 FERNS : BRITISH AND FOREIGN. inches of leaf soil, in company with Adiuntum and other Ferns, as well as much Cyperacece, the whole forming a dense undergrowth in woods. From this it appears there is no great peculiarity from other Ferns as to its place of growth, and that our ill success arises in consequence of the nature of the plant, being difficult to re-establish after being taken from its native soil, of which there are many instances; the common Pteris aquilina being a familiar example. The numerous and beautiful species of Hymeno- phyile, called Filmy Ferns, merit particular attention. They are natives of both tropical and temperate regions, and where they abound are generally found in shady moist woods, clothing the lower parts of the trunks of trees, especially Tree Ferns, or on dripping rocks, or surface-soil of the deepest ravines, rarely in exposed situations. In order to surround them with a moist atmosphere, adaptations must be resorted to, such as moveable cases upon the Wardian principle, hereafter to be described, and which are essentially necessary for the cultivation of this delicate tribe. Such, indeed, is their delicacy, that if once allowed to become dry on the surface of their fronds for any length of time, a rusty and shrivelled appearance will soon follow. Having, as already stated, succeeded in cultivating Trichomanes radicans, it becomes obvious that other species could also be made subservient to cultivation ; of which there is ample evidence to be seen at Kew, where there are forty cases* of different sizes, arranged on a stone shelf on the north side of the large Fern House, in such a position that they * May, 1864, CULTIVATION. 825 receive but little sun in winter, and are densely shaded. in summer, all filled with patches of fine fronds, of a number of different species, varying from the delicate hair-like Trichomanes tricvideum, not more than two inches high, tothe robust 7. anceps and T. radicans. They are grown in square shallow pans and boxes, well drained in the ordinary way, and having about two inches of peat soil mixed with nearly half its bulk of sand and small broken potsherds; but soft sand- stone is best. For the creeping sorts the soil should be raised in the form of a mound, and for those that have long-extending sarmentums, if soft stone cannot be had, it is desirable to invert a pan or common deep pot, covering it with a layer of soil, as already ex- plained, to which the plant will cling, and soon form a green hillock: junks of wood answer the purpose; but im a moist, close, and warm atmosphere, fungi and insects breed, and in a short time the wood decays, causing unnecessary disturbance of the whole mass of the plant. The singular genus Lygodium, and its ally Lygo- dictyon, grow naturally in firm soils, generally amongst trees and bushes, their wiry, flexile, climbing fronds growing over and involving everything within their reach in the most intricate complexity. In most Ferns, the whole of the divisions of the fronds are formed in the nascent or bud state, and are unfolded as the fronds elongate ; when the whole of the deve- loped parts are unfolded, the frond ceases further ex- tension. This is, however, not the case in Lygodium, Salpichlena, Pellea flecuosa, Gymnogramma flexuosa, Odontosoria aculeata, and a few others, the fronds of which are of indefinite extension, their apices con- 326 FERNS : BRITISH AND FOREIGN. tinuing to grow and produce lateral pinne, in every way analogous to the development of branches and leaves in woody plants. The climbing and rambling nature of these plants necessitates some kind of sup- port when cultivated in pots, which may consist of open wire trellises of # cylindrical form, varying in height from three to six feet or more, which, if fixed to a pot ten to twelve inches in diameter, the plants will, with careful training, become handsome speci-. mens ; they are also well adapted for covering trellis against wall or pillars. The fronds of the much admired genus Gleichenia are also indefinite in extension, and some of the smaller species,when growing amongst bushes, assume a climbing habit; but in G. furcata, G. dichotoma, G. Cunninghami, and G. flabelluta, the fronds are rigid and erect, produced from a more or less stout or slender sarmentum, which is either superficial or creeps a little below the surface of the soil; therefore shallow pans or boxes are best suited for these plants ; and with attention specimens of considerable size may be attained, as, for instance, at Kew a plant of G. flabellata measured twelve to thirteen feet in circumference and four and a half feet high, consisting of a thicket of fine fan-like fronds. i It may be expected that in such an extensive family some species would be found bearing the appellation of aquatics, but such is not the ease; for although many species grow in wet places, such as Osmunda regalis and Acrostichum aureum, both of which love water, but also flourish even in dry places, the only Fern really entitled to be called a water Fern being Ceratopteris thalictroides, and which is aiso singular CULTIVATION, ‘ ‘327 __in another respect—in being one of the few Ferns that are only annual. It is widely dispersed throughout the tropics, growing in wet places, often flooded; its sterile, viviparous fronds floating on or below the- surface of the water, as may be yearly seen in the Victoria Lily tanks at Kew. Being annual, care must be taken to preserve spores, which in the spring should be sown in a shallow pan of loamy soil made wet like mud, kept moist ; and when the plants are of sufficient size, the pan may be either filled with water, or be placed about an inch deep ina tank. Acrostichum aureum is mentioned as growing in wet places; for instarce, in Jamaica and other of the West-India Tslands it is described as taking the place of the Euro- pean Typha latifolia, attaining the height of seven or eight feet. By imitating its natural condition, placing it in a pan of water, or ina tank, specimens have been grown at Kew to the height of six feet; but although it enjoys and luxuriates in water, it also grows in dry places, its height then not exceeding one to two feet, and often with simple fronds. _ The delicately beautiful Selaginellas, on account of the large share of notice they receive and meet from the admirers of the Cryptogamic family, deserve here an especial notice. Originally the species were included under Lycopediwm ; but have been separated on account of differing in.the character of their spore-cases. The species are numerous, and their habit and free growth mark them as a very distinct _ group from true Lycopodium. With a knowledge of a few species, two very distinct modes of growth are easily understood; the upright or climbing ones, such as 8. Africana and S._ filicina, are exanple, and the - 828 FERNS: BRITISH AND FOREIGN. decumbent or creeping species, such as S. serpens, S. uncinata, S. Galiottii, &c., are examples. The latter delight in a light soil, composed of fibry peat and well-decomposed leaf-soil, free from pieces of wood (as it generates fungi), with a little sand inter- mixed, and require a depth of only about two inches in ordinary round pans, with ample drainage. Tf, however, good specimens are desirable, with an arrangement on a bench or low shelf, square pans about twice their height at the back as in front, arein every way preferable, and as at Kew, arranged alter- nately with the cases of Hymenophylle, with which they harmonize. The former-mentioned species, on account of their robust habit and strong rooting cha- racter, require the shallow pot, and a similar soil as for the last, but with a small proportion of light loam. The magnificent Selaginella levigata, if supported by wires against a wall, becomes, in a remarkably short. time, the loveliest object to be found in a Fernery. The species of the Rosulate section differ from the rest of the genus in having erect, fasciculate vernation, the frondules rising from a central developing axis, and, as in S. lepidophylla, spreading out nearly horizontal, and overlapping one another, forming a beautiful green rosette, about six inches in diameter ; when dry, they turn upwards and inwards in an involute manner, the whole mass of the plant thus forming a firm ball, which, on becoming moist, again expands; and even perfectly dead plants, when expanded, seem as if alive. In that respect being analogous to that of the insig- nificant cruciferous plant called the Rose of Jericho (Anastatica hierochuntica): hence the balls of this Selaginella are frequently met with in curiosity-shops CULTIVATION, 29 folie) under the above name. This hygrometric property seems to indicate that this and its allied species are subjected to occasional drought in their native localities. But although they may frequently undergo this pro- cess in nature with impunity, it is, however, not desirous to be too often repeated on plants artificially cultivated. Shallow pots, four to five inches in diameter, best suit this section; and being firm holders, it is necessary to insert pieces of soft sand- stone or potsherds amongst the shallow soil, so as to fix the plant firmly in the pot. The extremely delicate nature of the tissue of these little plants neces- sitates that a position the shadiest and moistest in a Fernery should be assigned to them. 8S. Willdenovii, S. filicina, and others are termed deciduous from their fronds decaying altogether in the early spring; new fronds, however, soon begin to grow. The species of true Lycopodium are also numerous, some being terrestrial and others epiphytal, hanging from the trees like various-sized cord, one to four feet in length. Of the terrestrial, five are natives of thiscountry; they, as well as several beautiful species that have from time to time been introduced from North America, refuse to become domesticated under ordinary treat- ment; to which must be added the wide-spread and beautiful tropical species L. cernuum. Hqual ill success attends the epiphytal species ; for although frequently imported, they are yet but poorly represented in our collections ; they adhere to trees by producing many fibrous roots, which ultimately become a large spongy mass. The extra height of roof required for the growing of good examples of Tree Ferns excludes them from many 330 FERNS: BRITISH AND FOREIGN. amateur collections. This has also been the case at Kew for the last twenty years, especially as regards Tropical Tree Ferns, many fine plants succumbing to the make-shifts that of necessity had to be resorted to after they had attained a certain height ; but by beginning with young plants, they may be grown fora number of years in houses of the usual average height of ten to twelve feet, as also the large fronded tree- like Lastrea villosa, Intobrochia podophylla, Asplenium striatum, Hemidictyon marginatum, and many others of like habit. The latter, at Kew, in a 20-inch pot, produced beautiful fronds, seven feet in height, and which might, with encouragement, soon be made to produce them equal to those of native growth—fourteen feet. But in order to get rid of the inconvenient and unsightly look of large pots and tubs, it is best to adopt for these plants the system of natural cultivation explained further on. In the “ Species Filicum” about one hundred and twenty species of Tree Ferns are described; but, according to Mr. Moore’s ‘“TInudex Filicum,”’ the number amounts to nearly two hundred. They are widely distributed, chiefly within the tropics. They love shade and, solitude, and are generally found at elevations of from three thousand to five thousand feet in the humid regions. In the southérn hemisphere they, however, extend much beyond the tropics, their southern limits being New Zealand, Norfolk Island, New South Wales, and Tasmania, where they grow at a lower elevation than within the tropics. On Mount Wellington, in the latter island, Dicksonia Antarctica is found in the greatest abundance, at an elevation of from one thousand five hundred to two thousand CULTIVATION. 831 . feet, attaining the height of from eighteen to twenty feet, growing only in damp. places, generally gullies, where the sun rarely penetrates, and where they are sometimes covered with snow, as represented in the vignette on the title-page. This suggests that, with proper selection of situations, they might live in the open air in the south and west of England, as also the mild climate of Argyleshire, where shaded ravines and gullies may be found similar to those of Mount Wellington. With few exceptions, Tree Ferns readily conform to cultivation. Alsophila capensis, A. excelsa, A. Australis, Oyathea deulbata, C. medullaris, Dicksonia antarctica, and D. squarrosa, grow freely in the Temperate House ; the lofty Alsophila glauca, the beautiful tessellated stem of Cyathea arborea, with its crown of fine fronds, and the broad shining fronds of Hemitelia horrida, assume a grand appearance in the Tropical House. In general, the stems of Tree Ferns are of sufficient size to warrant the name of trees; but in many the thickness is more apparent than real, the diameter of the woody centre being often only a few inches, but in many cases covered with successive productions of out-growing aérial roots, which become hard and wiry, and by their interlacing, form a com- pact mass; the points of these roots are, however, the active feeders, and if a layer of fresh soil is occasion-. ally placed round the base of the stem, their growth will be promoted, and vigour given to the plant. The most fastidious Tree Fern to cultivate is Dicksonia arborescens, anative of St. Helena. It was first intro- duced to this country in 1786, and many times since, but refuses to become established, either in a tropical 332 FERNS: DRITISI. AND FOREICN. or temperate house. This species, as well as the whole of the species of true Dicksoni@, are characterized by the base of the stipes being clothed with a more or less coating of beautiful, articulated silky hairs; in some species of Cibotium, the quantity is so great, that in the Sandwich Islands it is collected, and ship-loads of it sent to California and Australia for stuffing cushions, beds, &c. Another remarkable species of this alliance is Cibotium Barometz, a native of China and other parts of Hastern Asia; its fronds attain the height of twelve to fourteen feet (even in this country) ; they rise from a thick decumbent caudex, which is densely covered with silky hairs, as above described ; lying on the ground, it has the appearance of a woolly- clad animal. The stories told about it to early travellers led them to describe it as an animal with flesh and blood, but fixed to one position, from which it never moves ; hence the story of the now fabulous Barometz, or Vegetable Lamb. This plant is of easy cultivation ; if placed on soil slightly raised, a few years’ growth will produce very good specimens of the ‘ Lamb.” The rare Schizocena sinwata is an exception to the general rule that characterizes Tree Ferns, the stem being slender, not exceeding an inch in thickness, attaining the height of three or four feet, and bearing a fascicle of simple fronds. This remarkable Fern is a native of Ceylon; it grows in shady places in woods where a degree of coolness prevails. A few years ago plants of it were received at Kew, and it was found necessary to place them in a large Wardian Case, in which they flourished. Although the plants belonging to the order Muarat- tiacee are ranked with Ferns, they nevertheless differ CULTIVATION, 333 much in habit, chiefly as regards the nature of the roots, which are thickened and fleshy, and generally pene- trating deeply into firm soil. In their general character they present much resemblance to the roots of Cycadee, Cyclanthce,and other allied endogens; also in venation the remarkable genus Stangeriaconnects them through Dancea with Cycadee ; and their compound fronds are represented in the same family through the new genus Bowenia. The species of Marattia and Angiopteris grow freely in a loamy soil, requiring plenty of water. Some species of Angiopteris require much space, for although the caudex does not rise much above the ground, yet a plant at Kew produced fronds that reached the height of twelve feet, spreading outwards, forming a diameter of thirty-four feet. The species of Danea and Kaulfussii- are, however, not such free growers; they require a moist and high temperature. T have now gone through a few of the principal genera: requiring special kinds of treatment ; the next consider- ation is good management as regards temperature, watering, airing, and keeping free from msects. Much depends on the size and nature of the house ; the larger it is, the less fluctuations of temperature take place, especially in a house of the ordiuary construction of glass roof and sides, whether lean-to or span. The maintenance of a proper condition of the air in plant- houses depends much on the nature of the interior fittings. White or polished, reflecting and radiating surfaces, should be avoided as much as possible, such ag iron pillars, rafters, spandrels, polished slate or smooth stone, iron or stone floors, &c. These kinds of matcrial are often introduced in superabundance, for 334 FERNS: BRITISH AND FOREIGN. the purpose of giving consequence and dignity to the house, thus laying the foundation for conditions un- favourable to plants. The shelves orbenches may consist of dark-coloured porous stone or rough slate, with an edging of smooth slate two inches deep, so as to form a shallow trough, which should be ‘filled with dark- coloured sand or fine-sifted coal-ashes, the whole pressed down, so as to form a firm smooth surface, on which the pots are to be arranged. By this means a degree of moisture will be retained more congenial to the plants than when standing on stone or slate. An edging of about three inches of Selaginella hortensis or S. denticulata gives a neat appearance, and if allowed to run between the pots, the whole becomes more con- genial than a naked surface. White sand or broken quartz is frequently used, but it soon becomes dirty, and has a harsh and dry appearance. In the arrangement of the plants some degree of order must be observed, so that every specimen plant of a species should be seen to advantage, and if the collection is extensive, and the house has different aspects as regards light and shade, then it is desirable to arrange the plants according to their habits and requirements; thus, at Kew, the division Hremobrya occupy the whole of the south side of the house; this division being less sensitive to the effects of the sun or deficiency of water than the more extensive division Desmobrya, which, on account of their thin texture, very quickly suffer from dryness or too much exposure to the sun; therefore the north side of the house is best for them. i The natural direction of the fronds is the most pleasing, whether uprigut, pendulous, or spreading. CULTIVATION. 335 In the latter case, sometimes their own weight makes them fall lower than is convenient; it therefore becomes necessary to support them by inconspicuous stakes. Care must, however, be taken not to raise the fronds above their natural position, and to avoid making them have a stiff formal appearance. In “Theories of Horticulture,” it is said that abundance of lght and a free circulation of air are indispensable for the cultivation of plants under glass. In my long experience, I have found, as a rule an abundance of light and a too free circulation of air prejudicial to the good cultivation of plants in general. With regard to Ferns, they love a quiet, dull atmo- sphere ; the light afforded by a roof glazed with the ordinary transparent glass, admits fully double the amouut of light necessary during eight months of the ‘year. Sufficient means are, however, necessary for admitting air, in order to keep down the temperature in summer; but care must be observed in admitting it; if not, the increased ventilation in lowering the temperature will cause a rapid dispersion of the essen- tial moist atmosphere at a time when most required. Shading must then be resorted to, this being the chief purpose which it serves, besides protecting the plant’s foliage from the too powerful rays of the sun. It may i consist of canvas blinds on rollers; two moderately thick ones are much better than one very thick, espe- cially when Trichomanes and Hymenophyllum are grown; one should be permanent during the height of tlie summer, the other to roll over it on occasions of full sunshine. Although canvas is here spoken of as best for shading, and is extensively used, still it is trouble- ‘some, and in the end expensive, which makes it de- 336 FERNS: BRITISH AND FOREIGN. sirable to resort to some other method; such as by using several kinds of dulled or rough glass. In former years, the fruit and plant-houses at Kew were glazed with a very dark-green glass called Stourbridge-green, and which was patronized by the late Mr. Aiton. Fine crops of fruit were produced under it, also the tropical plants in the Botanic Garden flourished without the aid of canvus or shade ° of any kind. Not many years ago, solitary squares of this glass might be seen in the roofs of the old hot-hcuses, which strongly contrasted with the modern clear glass. My experience with this glass led me to recommend green glass for the Palm House, which was adopted ; but the modern-made green tint does not appear to be so fixed a colour as that of the old Stourbridge-green. The Palm House in the nursery of the late Messrs. Loddiges, at Hackney (now things of the past), affords another example of successful plant-growing without the aid of canvas or other moveable shading material. On the late Mr. George Loddiges being consulted respecting the glazing of the Kew Palm House, he was asked if they shaded theirs; his reply was, “Oh, no; our thick rafters and sash-frames, with sooty glass, just afford the amount of light necessary for the plants.” Under this roof, in an atmosphere of stillness and gloom, Palms, Ferns, Orchids, and numerous other tropical plants, grew in the greatest luxuriance, which, with the proverbial solitude of the place, and when viewed from the elevated platform, - gave the idea of a ravine in a tropical forest. To judge by the above examples of Kew and Hack- ney, it appears that fruits and plants were successfully CULTIVATION. 8o7 grown under roofs very different in appearance from the elegant clear glass roofs of the present day, which, nevertheless, necessitates some kind of invention for temporary shading in summer; but it is to be hoped this will be superseded by further improvements. Double glazing is now spoken of as answering the twofold purpose of saving shading and fuel; but as there appear to be different opinions on its merits, further experience is therefore desirable before it can be safely recommended. With regard to watering, it is difficult to explain in words, or laydown arule, what constitutes the extremes of dryand wet soil ina flower-pot, feeling is perhaps the best guide; if, on taking a pinch of soil between the finger and thumb, the particles will not adhere, then it is too dry ; if it adheres, and falls like a dry wafer, then the plant is not in immediate want of water; but, on the contrary, should the soil stick to the fingers like a wet wafer, then the whole is in a bad condition, and should the plant be weakly, it will get worse if allowed to remain in the sodden soil; for the correctness of this test care must be taken not to be deceived by the surface- soil, for it may be either dry or wet, and the contrary lower down. Another test of the state of soil is to give the side of the pot a smart tap with a hard piece of wood; should the sound be hollow, with a little tone, then the ball is dry.; if, on the contrary, there is only the dull sound of the tap, without any tone, then the ball is moist and tight in the pot. To those in the habit of watering the same plants for a length- ened time, this test is of service, but it must not altogether be depended upon. The quality of the water has great effect on the health of the plants; PA 338 FERNS : BRITISH AND FOREICN. It should always be soft, or, if hard, should be aerated: - ‘in open tanks; rain-water is best, but if collected from the roofs of hothouses, care must be taken that it does not become impure by the decomposition of the paint and putty, which is of frequent occurrence, and in time it so impregnates the water as to cover the foliage of the plants, when often syringed, with a thin film, that gives them a dull look; care should also be taken that the temperature of the water should not be less than 50°, especially in winter. Syringing is made an operation of great importance in guide- books on cultivation, and is readily accepted by the unskilled amateur’ as necessary, and being amusing, is often carried out in the extreme, to the injury of the plants. An amateur remarked not long ago, “Now we have got such a nice lady’s syringe, it is quite a delight to use it’? Another remarked, “I every day regularly water, syringe, and sponge my pet Adiantums, but with all my attention they are getting of a brown colour, surely your plants (at Kew) must have great attention to keep them so green.” The lady syringer is told that many of the plants before her have been in the same pots for several years, and that: they get water when dry, and are never syringed or sponged, unless for removing insects when they appear, From what has now been stated, an amateur’s successful cultivation of Ferns simply depends on proper soil and potting, careful watering, placing in a quiet, moist atmosphere, keeping down temperature in summer by shading, and to avoid currents of dry air, and in winter to maintain the proper temperature, according to the nature of the plants, by means of CULTIVATION. 399 hot-water pipes. For the latter purpose, in small houses, temporary stoves of gas and charcoal are used, often to the utter destruction of the plants. To assist in maintaining a moist atmosphere besides the usual moist surfaces of the pots, it is desirable to have water-troughs on the top of the hot-water pipes, as also to place pans of water on or near the pipes; also in hot weather, the paths and other evaporating surfaces should be wetted every morning and evening. An occasional syringing may be given, which should be done when the out-door air is charged with vapour; wetting plants when the air is dry causes a sudden evaporation, and a reduction of temperature on the surface of the frond is the con- sequence. This is especially the case with broad, smooth, glossy species, such as the genera Phymatodes, Neottopteris, Asplenium, Diplazium, &c. ; occasionally the whole of some fronds or part of a frond becomes black. In order to raise vapour, a practice prevails with many cultivators to throw water over the hot-water pipes (or flues) ; there is no objection to this, pro- viding the air out of doors is warm, but in the winter season itis often done in the evening, in order to counteract the dryness of the air, caused by the extra heat of the pipes required at that season ; the house then becomes filled with hot vapour, which coming in contact with the glass of the roof condenses and falls in a shower of cold drops on the plant; at the same time the temperature of the house rapidly falls, thus causing black fronds, spots in orchids, and such-like complaints. Ferns, like other plants, are liable to be more or less infested with insects; the soft membraneous kinds Z2° 340 FERNS : BRITISH AND FOREIGN. being the most subject, while, on the contrary, the smooth-fronded kinds, such as the whole of the division of Hremobrya, Elaphoglossum, &c., are, it may be said, quite exempt. The mealy bug, three kinds of scale, and thrips, are the chief pests, and should be carefully watched. The oblong brown scale is generally the most common, and very quickly overruns the under side of the frond, forming lines along the midribs. Many kinds of fluid mix- tures, powders, and other nostrums, have from time to time been advertised for their destruction, but it too often happens what kills insects also injures the plant, and makes it look unsightly. Fumigation with tobacco is the grand preventive against the breeding of plant-insects; very slight fumigation destroys the winged male insect of all the Coccos family, and if frequently repeated, the whole in time will become extinct. When the mealy bug gets a head, it is readily got rid of by syringing; not so the scale, it requires to be loosened with a soft brush, or blunt-pointed peg, which, if the weather permit, should be done out of doors, and the plant afterwards syringed, so as to clear away all loose scale aud eggs. Scale insecis are adherers and suckers only ; not so the less conspicuous insect the thrip, which is a small, slender, black, shining insect, with large prominent eyes, and quick in its movements, herdmg in groups, generally on the under side of the frond, voraciously feeding on the cuticle, which they soon destroy, permanent injury being often done without any indication of their presence till too late; they must, therefore, be carefully watched, and, when seen, at once destroyed by pinching with the fingers, CULTIVATION. 841 and syringing the plant. They are generally found on species of Adiantum, Pellea, Platyloma, Doryo- pteris, Hemionites cordifolia, and other genera having fronds of a smooth, chestnut-brown colour. Within the last twenty years a small, white-winged insect, like a midge, has made its appearance (supposed to have been first introduced with imported plants to Kew). They congregate on the under side of the fronds, and, — when the plant is moved, dart off like a flock of white pigeons. At first it was supposed to be harm- less; but such is not the case, as it has been found to feed on the cuticle like thrips; but they are not so easily caught. Repeated tobacco fumigation destroys it.* Red spider seldom attack Ferns; when such is the case, it is a sure sign that the air of the house is too dry. Few plant-houses are exempt from the well-known cockroach, which when once introduced is one of the greatest of pests. They increase and multiply most rapidly, and are most voracious feeders, eating and gnawing the fronds of Ferns, old and young, often completely in one night spoiling fine plants; there- fore war in all its forms must be waged against them. They are night marauders, hiding and breed- ing in crevices of masonry in dark places, generally near the furnaces, or hottest parts of the heating-pipes, and are even found lurking amongst the drainage, in large pots, from which they issue at dusk to commence their ravages. Many expedients are resorted to for their destruction, such as attacking them in their day * This insect has been described and figured in the Gardeners’ Chronicle, of 1856, by Mr. Westwood, under the name of Aleyrodes vaporariorum. : 342 FERNS: BRITISH AND FOREIGN. abode, by the appliance of boiling-water, sulphur fumes, or exploding gunpowder: when such agents can be used, they deal with them quickly and whole- sale. Various kinds of traps are used, which, with poison, will, if daily attended to, completely extirpate them. But it must be borne in mind that, although the whole, old and young, may be got rid of in the course of a fortnight by poison— the effect of which is greatly increased by the living eating the poisoned dead,—eggs are however left, which will soon pro- duce a new generation that must not be allowed to arrive at maturity. Under the ordinary varying atmosphere of hot- houses, insects seem not to be affected, for if their extirpation is not attended to, they will be found in more or less abundance all the year. Not so the sooty mildew,* a fungus covering the upper surface of the leaves of plants with a black, sooty coat, and for their sudden appearance, like that of the grape mildew, the potato disease, and other sporadic plagues, no satisfactory causes have as yet been assigned: The pest now under consideration may be called one of these plagues ; in some years it is not seen, while in another it soon overruns and quickly covers Ferns, and other plants, in hothouses, The broad-fronded species of Aspidiwm, Meniscium, Gonio- pteris, Anyiopteris, &c., are very subject to its attacks. Books on mycology name and describe these pests, but not how to prevent them; and books on horti- culture instruct how to get rid of them; the principle of which seems to be dusting with sulphur, washing, * Fumago foliorwm, Fries. CULTIVATION, 343 and syringing, the latter being the only remedy for the black mildew. This concludes my observations on pot-cultivation, and in order to save repetition, it must be understood that the process as regards soil, watering, airing, &c., is equally applicable to natural cultivation, which I now proceed to describe. TI. Narorat Cunrivarion. HE natural cultivation of Ferns consists in growing 7 them without the aid of garden-pots. This is accomplished by placing them, as far as artificial appliances will permit, under conditions and influences analogous to that of their native wilds; for that purpose uneven irregular surfaces are best adapted, whether natural or artificially formed; and as Ferns are generally called rock-plants, which many truly are, therefore, in order to imitate rocks, the surface on which they are grown is made irregular and covered with rude stones, on or between which the Ferns are planted; hence the term Rockery is applied to this kind of cultivation. For all species of Ferns, either hardy or tender, this system of culture, which admits of their being planted out, is far preferable, to whatever extent it may be desirable to practise it, whether in the sheltered nook, in the open air, or in stoves or greenhouses, either partially or entirely devoted to it. For by ib is the finest health imparted to the plants. The rockery bank which once occupied the Temperate Fern House at Kew, together with the fine fronds of tropical species grown on rockery in the Palm House, afforded excelient 844 FERNS: BRITISH AND FOREIGN. examples of the superiority of this mode of growing plants over that of pot culture; by proper arrange- ment from ordinary level positions, their delicate hue and elegant form of outline may be seen to much greater advantage, as the upper surfaces of their fronds are generally presented to the eye more fully. And where this system is largely carried out, if a raised platform or footway be erected considerably above them, the advantage is still greater. The great beneficial influence this system has on the plant is mainly consequent on the more uniformly moist at- mosphere, so congenial to all Ferns, which can be successfully maintained from the great extent of the more natural evaporating surface of the soil, and ma- terial, with which the rockery is composed ; and as they are allowed to establish themselves at freedom in it, there is a similar uniformity of temperature at the root, as well as of moisture. As is well known, in large masses of earthy and other solid material, its temperature does not fluctuate with that of the house, at least, as in the case with ordinary-sized pots of soil, where the absorption and radiation of the heat takes place so much more rapidly. It may further be men- tioned, that, in a house entirely devoted to the cul- tivation of Ferns under this system, its evaporating surface does not wholly consist of the principal mass of - material composing the rockwork ; there is the large extent of wall-surface, which, if properly constructed with rough and absorbent material, is continually giving off its moisture, besides, from that reason, affording an excellent opportunity for Ferns of more or less scandent habit to assume their native character in luxuriant profusion, by growing on it as on natural CULTIVATION. 845 rocks. As regards the design of any structure in- tended to be wholly or partially devoted to the culti- vation of Ferns, it is of course a good deal a matter of taste and convenience, as the system is available in almost any kind of structure, large or small, but varying in the degree of economy in its construction. To this end, the first consideration should be to construct in a manner that will insure the necessary amount of shade and moisture and (when required) heat, with as little auxiliary assistance as possible. Therefore if a lean-to house be adopted, the aspect should be west or north-west, with the back consisting entirely of wall. Should a span-roofed structure, how- ever, be preferred, which indeed for all purposes is much the best (and which will be here treated. of as a general example), its aspect should be north and south, by which a shadier and consequently a moister side of the house will be obtained, suitable for Ferns of more or less delicate structure, such as the genera Trichomanes and Hymenophyllum. A sunk house, in the manner of a ditch or railway-cutting, is most suitable, and will give ample height for the growth of Tree Ferns, without the necessity of having high side walls, and thus exposing a lessened portion of the house’s surface to the action of climatic vicissitude; consequently a warm moist atmosphere may be main- tained with but little assistance from artificial heat. As a source of heat the hot-water apparatus is to be preferred, on account of its economic utility, and where practicable the pipes should be hidden as much as possible with the rockery material. Perkins’s sys- tem of heating is perhaps the best for a house of this kind; it consists of coils of small pipe, which can be 815 FERNS! BRITISH AND FOREIGN. arranged in any part of the house, and are readily detached and again fixed at another point when found necessary. The first deposit for the rockery should be rough and somewhat absorbent, such as irregular brickbats, masses of stone, loosely disposed to admit of a ready dissemination of the heat amongst its parts, and then follow with the material proper. Apertures for the escape of the heat should be con- structed at frequent intervals, drain-pipes being very snitable for this purpose, which, by the mass being kept properly moist, will allow the heated air to circulate in a congenial steamy condition throughout the house. In the design and arrangement of the principal mass of material composing the rockwork, it is, as in the design of the house, a good deal a matter of taste and convenience as regards size; whether in repre- senting in miniature a rugged mountain side, or deep glen, with its clear pool, reflecting the elegant forms of the Ferns growing near it; or may be a miniature valley, with undulating surfaces gradually rising on either side, with projecting rocks. There are nu- merous kinds of material more or less suitable for constructing a rockery either indoors or out, and but a few localities where some of these may not be easily procured. The chief desideratum is to have.such that is of porous consistence, and generally with rough and ragged surfaces. It is useless to be particular in the selection of various kinds of stone on account of their fine quality, either in texture or colour ; for where Ferns properly thrive all such will be speedily obscured by their luxuriant growth; consequently any brick- field affords very good material that would contribute largely in the general structure, in the way of con- CULTIVATION. 347 glomerated masses of brick, or bricks artificially amassed with Portland cement to form miniature rocks, rugged cones, rough pillars, or perpeudicular surfaces. However, when procurable, the absorbent androughly quarried soft stone should enter largely into the composition of the rockery, as it is pre-eminently useful, not only in contributing so largely to the natural appearance of the whole, but, when the more delicate members of the Fern family of creeping habit are grown, it is invaluable in affording a moist surface for their tender rhizomes to cling to. It is seldom that wood of any kind is found of much use, as it is so soon reduced by decomposition, and fungi produced im contaminating profusion. The soil to be used in the rockery-house should be selected and applied according to the principle already given in pot- culture. The style of the Fern-rockery having been decided on and executed accordingly, the next step will be to arrange the plants about it; and to carry this out satisfactorily is not the least difficult part of the whole undertaking, as it is not only essential to provide positions for each individual plant, but such that will prevent any injurious interference of its neighbour, either by fohage or root. Should the roof require support (and if not, should it be consistent with elegance and regularity), pillars composed of rough porous stone or brickwork may be constructed, and, if admissible, united above as archings, which, when clothed with the luxuriant foliage of scandent and pendulous Ferns, intermixed with the broad foliage of creeping Aroidew, growing from their crevices and recesses, would greatly enhance the beauty of the whole, and give a most accurate and 348 FERNS: BRITISH AND FOREIGN. durable example of the manner in which many spccics of Ferns and Aroidee clothe the trunks of trees in tropical forests; also several species of epiphytal woody plants may be introduced with good effect, such as Tanecium parasiticum, Marcgravia umbellata, Norantea coccinea, Hoyas, and other creepers not ‘subject to be infested with insects. The species of Phymatodes, Pleopeltis, Davallia, and several allied genera, as also Polybotrya and Stenochlena,will luxuriate over the moist surfaces and form a compact mass; and, if planted near the wall, direct their rhizomes upwards, and the plants will quickly assume a most elegant appearance. The grand and beautiful Schellolepis verrucosa, and its allies, should be placed in elevated positions, as their fronds are long and pendulous. Elevated positions will also be found for the species of Platycerium, the grand Drynaria Heraclea,and D. coro- nans: the latter is remarkable in its thick rhizome, progressing in a circular direction round trees, which, with the erect, broad, rigid fronds, gives the idea of a coronet or crown; this circular direction appears to be normal. A fine plant at Kew, in the course of eight years, grown in a shallow pot, formed a circle more than a foot in diameter. For that portion of the terrestrial group composed of stronger growing spe- cies, such as Phegopterideee, Pieridew, Aspleniee, &c., for which firmer soil is necessary, the ordinary surfaces of the rockery offer most suitable positions, arranged there in a manner as their sizes indicate, so as to be in harmony with those of smaller size around them. It is perhaps essential to note that the Gymnogramma, Cheilanthes, and their allies, require a place in the Fernery, the lightest and best drained; therefore CULTIVATION. 849 the higher part of the rockery is best suited for them. In such a Fernery as this now before the mind’s eye, convenient positions, in the form of dripping crypts and recesses, may be provided for the delicately beau- tiful and easily grown Trichomanes and Hymenophyllum, without the aid of glass cases that are necessarily used, as already explained, under pot-culture. Where Tree Ferns would be grown, the proper situation for them is the lowest part of the house, in the valley, where the greatest head-room would be obtained, and most constant shade and moisture about their stems, which will encourage the essential growth of the aerial out-grown roots. The path in the valley may be straight or winding between the Tree Ferns, the intervening spaces to consist of raised mounds or lillocks, to be planted with the smaller kinds of Tree Ferns and other large-fronded Ferns. The different species of Selaginella should occupy the intervening spaces between the plants, which will add greatly to the natural appearance of the whole and assist in pro- moting a genial atmosphere. To whatever extent a house of this nature may be carried for tropical Ferns, the same must be allowed for the extra-tropical kinds, and being sunk, the necessary amount of heating power required will be just sufficient to keep out frost. Keeping the house shut during the winter season, so as to maintain a quiet, still atmosphere, greatly assists in repelling frost, indeed for such houses ventilation is only necessary to keep down high temperature during summer; free admission is, however, beneficial when the external air is still and moist, when even the tropical] division may be freely left open during the nights of 350 FERNS: BRITISH AND FOREIGN. summer, but taking care to prevent currents of dry air. I have now stated the chief points to be observed in forming and arranging an indoor natural Fernery, the principles of which are capable of application to houses of the ordinary construction and usual average size, or to any extent that means will allow; even to realize the grand idea of the celebrated Loudon, who, more than thirty years ago, speaking in favour of span- roofs, said, “There would be no difficulty in covering ten acres of Kew Gardens with glass by a series of span-roofs.” Although since then much has been done at Kew, by the erection of lofty plant-houses, still the area covered falls far short of ten acres; but as natural cultivation is now patronized, and with the improving age and desire for novelty, let us hope that the time is not far distant when Kew may have at least one acre converted into a straight or windine Fern-valley covered with glass, the highest part not to exceed thirty or thirty-five feet above the centre of the valley, that being sufficient height for such Trec Ferns as may perchance withstand the vicissitudes of thirty years careful treatment. In a house of this kind, furnished with means of maintaining a temperature of 60 in the coldest weather, situations in it would be found suitable to the good growth of not only Fvrns and Aroidece, but also Lromeliacee, Orchids, and other kinds of plants of an epiphytal nature. For an open-air Fernery a sheltered situation should be chosen, and if possible within the influence of shelter and shade, but not under trees ; a sloping bank, a natural or artificial hollow, such as an old gravel-pit or sunk fence, may be turned to good account. The CULTIVATION, 851 number of species of exotic Ferns sufficiently hardy to bear the cold of our winters is very limited, much less than is generally believed, the chief of them being natives of North America, which, with the British species, amount to about 80 in all; therefore the extent of space required for a simple collection of ‘hardy Ferns is small ; an area of 60 feet by 10 will be ample, and which may be either on the side of a slope, or between two slopes with a path in the centre and open at one end only; or it may be a raised oblong mound with a path all around it. If not confined to a strictly scientific collection, and space to be had, an interesting natural glen may be formed by introducing other plants conspicuous for their large fohage. It has been already stated, that although our small- growing native and several exotic species, such as the rock Asplenie, Woodsia, &c., endure the extremes of heat and cold in their native localities, they neverthe- less suffer by full exposure under artificial cultivation ; it is therefore necessary to plant them on the rockery, so that they can readily be protected in winter; an inverted flower-pot, placed over each plant and covered with leaves, or some loose material analogous to the covering of snow of their native country, is a good protection against being injured by excess of moisture or severe cold. Where the rockery is to be carried out on a large scale, then eaves and chasms may readily be constructed, and with a natural or artificial head of water they can be kept moist by the constant drop- ping and spreading spray; thus making appropriate positions for the natural growth of Adiantum Cappilus, Hymenophyllec, &c. But when a full collection of the many varieties of Scvlopendrium, Asplenium, Lomaria, 852 FERNS: BRITISH AND FORZIGN. &c., are grown, then a pit, or frame with glazed sashes, is the most convenient mode of keeping them. The width of the pit should not exceed 5 feet, 3 feet or more high at the back, to slope to about a foot at front, its length regulated according to the number of plants. They may be either grown in pots, arranged to slope to the front, or a bank may be formed with small rockery stones, neatly arranged with the plants between them, and if the mason work for the frame or pit is contrived to be hidden, the whole will then be in character with the general rockery. By this an interesting little bank may be formed, and by proper arrangement of the plants the variety lover can at a glance see the difference that marks one favourite from another. The glass pro- tection will stimulate the plants into early growth, and finer fronds will be made, but care must be taken not to expose the young fronds to the harsh drying winds of spring, or late frosts, which often do great injury to out of door Ferns; by midsummer, the glass sashes may be entirely removed, due attention being paid to watering and shading. If the soil and subsoil of the situation chosen for a hardy Fernery is stiff and retentive of moisture, and the neighbouring trees, walls, &c., are clothed with Musct and Iichens, it is favourable for Ferns without the aid of much rockery; but if the soil is light and sandy, then rockery is most essential for retaining moisture, not only for Ferns, but also all kinds of small herbaceous plants. Formerly there were great masses of rockery in the Botanic Garden at Kew, but modern iuste has swept the whole away, which, with the removal of trees, walls, and old shrubberies, has CULTIVATION. 853 caused the garden to become drier, as is manifested by the disappearance of above nearly forty species of Mosses, Hepatice, and Lichens; the effects of this, with other causes, have been hurtful to the hardy Fern col- lections. In forming a new rockery, after the plants have become once established, as few changes should take place as possible, either with the plants or stones on which Musci should be encouraged to grow; and in order to give a decorative effect, Sedwms, Semper- vivums, Sazifrages, &c., may be placed on the pro- jections and shelvings of the stones, the whole bordered by a line of spring-flowering plants, such as Iberis sempervirens, Alyssum sawatile, Arabis alpina, and ‘other plants of that nature. It may be also mentioned that the vicinity of water is a favourable position for a Fernery; where such does not already exist, an artificial piece of water should be made in conjunction with the rockery, varying in size from a small basin to any extent: it may be circular, oblong, straight, or winding, with irregular projections. In ordinary cases its width need not exceed ten to twelve feet; its margin should con- sist of a bog or border, not less than three feet wide, having a slope to the edge of the water. This border will afford the means of growing a great many curious sub-aquatic and bog plants, and the projections will be excellent positions for the different species of Osmunda ; water-lilies occupying the centre. The want of such a rockery and aquarium has been- much felt at Kew; for with all the great cost for lakes, ponds, and water-works, yet no arrangement has been made for growing a scientific collection of bog and aquatic plants, either hardy or tropical. 24 854 -FERNS: BRITISH AND FOREIGN. TV. Cuttrvarion or Ferns 1x Warp’s Caszs. T is now thirty years since I was invited by Dr. Ward to visit him at his house in Wellclose Square, for the purpose of seeing plants growing in cases and glass jars, so closed as to be considered air-tight. Knowing, as I did, the common practice of growing plants under hand and bell-glasses, I vherefore could not appreciate what I had gone to see until I was made aware that the plant-loving residents of such smoky and soot-falling districts of the metro- polis, as that of Wellclose Square, could grow rare and delicate plants equal to those at Kew. An account of this: method of growing plants appeared in the Companion to the Botanical Magazine for 1836, and in April, 1838, the celebrated philosopher Mr. Faraday delivered a lecture at the Royal Institution on the subject, which may be considered as the advent and introduction of Wardian cases, under which a large portion, and decidedly the most beautiful of the Fern family, are now successfully cultivated in the sitting-rooms of the town-confined lovers of natural objects. In 1842 Dr. Ward published a small work on the subject, giving a history and details of manage- ment, which renders it unnecessary for me to say more regarding the early history of Ward’s cases. The principle on which the system is founded, consists simply in shutting up air in glass cases, in such a manner that it is not readily influenced by changes of the external atmosphere. The case also contains several inches depth of moist earth, that gives off CULTIVATION. 355 moisture to the absorbent, undisturbed air above it, which becomes more or less saturated, as dependent upon changes of temperature, and thus becomes proper for the growth of plants. The case may be constructed of any shape or size, according to taste or means; it can be square or round, an octagon or hexagon; the roof may be a dome, span, or sloping, but by all means avoid a flat one; and be the shape whatever it may, the design should inevery point be neat, and not of such an ornamental description as to be more attractive than the plants, A very good, interesting collection may be grown in one 3 ft. long by 14 ft. in width, and 24 fect high; it will have the best appearance if the sides are con- structed with single squares of glass; but if divided, it must not be into more than three, as many divisions spoil the effect; the ends should be made to open, to enable any dressing or removing of old fronds to be done conveniently, and a small piece of perforated zinc should be inserted in the apex of the roof at each end, which will assist in preventing the almost universal complaint, that the plants cannot be seen for condensed water on the inside of the glass; this is caused by the variation of temperature. If the case stands in the sun or becomes warm inside during the day, and retains the heat, whilst the temperature of the room may fall considerably during the night, the cold air, acting upon the glass, con- denses: the warmer vapour inside and obscures the plants. By the introduction of the above-named remedy the temperature is more equally balanced, and the plants are always to be clearly seen. Should the air outside become very hot and dry, it will be 242 306 FERNS: PRITISH AND FOREIGN. advisable to close the ventilators for a short time during the day. The glass case should be entirely in- dependent of the soil-box, but to fit the inside, resting in a groove or rabbet. The box should be about 6 in. deep, and may be made of zinc, brass, or wood lined with gutta-percha or zinc; but metal of any kind in contact with the earth or air in which plants are grown is not genial to either their roots or foliage. I have always found the plants succeed best in a neatly-made wooden box lined with pitch, having a small tap or cock in one corner of the bottom, for letting away any excess of water; but this will not be necessary if proper attention is observed in supplying the plants with a sufficient amount of water at one time, which, in consequence of little or no evaporation taking place, will be seldom required. This knowledge can only be gained by practice ; many amateurs’ failures with Ward’s cases being caused mostly by giving too copious waterings at certain stated intervals. The height of the stand must be regulated accord- ing to whether the plants are to be viewed in a sitting or standing position; for the former the ordinary height of a table is a sufficient guide, and for the latter afew inches higher, so that the plants can be seen through the side glass rather than from the top. In preparing the box for the plants, about one inch of its depth should be filled with sand or other drainage material, such as is already explained in pot-culture—but in Ward’s cases this is only necessary as a precaution against an over-supply of water,—the rest filled with soil, which should be good fibry peat and silver sand, intermixed with pieces of ba CULTIVATION. 857 sandstone broken small, or (if peat cannot be pro- cured) good light loam. Having proceeded so far, two systems of planting the case present themselves. ‘The first plan is to plant in the soil, which should be raised in the centre; the number of plants will depend on the size of the box, and care must be taken not to plant them in too crowded a manner, the distance apart depending on the size and nature of the plants, but in no case should they be closer than six inches. The second method is to have the - plants established in 4 to 6-inch pots, plunging them in the soil sufficiently deep to hide the rims. of the pots. The last system has one decided advantage, namely, should a plant die or does not succeed, it can be removed and replaced without disturbing its neighbour. It must be understood that the above mode of planting relates to plants with erect verna- tion only, but those with creeping rhizomes, that form cespitose tufts, require to be planted each on a separately raised hillock, of which, if the case is large, there may be a series, and for the creeping Hymeno- phylla lumps of porous stone are very suitable; fine patches may be obtained in this way, not only of Hymenophylla, but also of the smaller species of Niphobolus, Anapeltis, the beautiful Humata pinnati- fida, Davallia pentaphylla, and others of like habit. After the Ferns are planted, some small-growing Selaginella, such as S. apus, may be pricked in over the surface of the soil, in pieces a few inches apart, -which will soon cover it, and give to the whole a neat and finished appearance, and also materially assist in maintaining the proper state of moisture in the air of the case and about the plants. The planting bein 2 358 FERNS: BRITISH AND FOREIGN. now completed, the soil must be brought to a uni- form state of moisture; this is best accomplished by two or three moderate waterings at intervals of half an hour, and when thoroughly drained, the case may be closed and placed in position. To make cases, ornamental pieces of marble, shells, &c., are often introduced ; but this should not be tolerated, as they: do not harmonize with the occupants of such a structure. For Hymenophylla it is necessary they should be sprinkled overhead occasionally ; to enable this to be done, the tops of all small cases should be moveable, and in large ones a hinged pane is required. A sponge, or small thumb-pots filled with water and placed out of sight, will greatly assist in keeping a moist atmosphere, which is so essential for the health of these filmy-leaved plants. If the cases stand in a room where a fire is regularly kept in the winter, a great many tropical species may be grown, and in situations where they stand exposed, such as balcony windows, halls, &c., care must be taken, as winter approaches, that the soil does not become frozen, such being very detrimental to even the hardiest Ferns. “Hot bricks and bottles filled with hot water have been resorted to as a preventive against frost, as also for maintaining a proper temperature for tropical species; but unless the bottom of the case is constructed for that purpose, and arrangements made for this mode of heating, to be strictly and regularly attended to, it had better be dispensed with. Should the case be exposed to the direct rays of the sun during the summer, it must be shaded, and care taken that the temperature inside does not CULTIVATION. become too high; it should not exceed 70°. 859 This will be much assisted by placing the shading material at some distance from the case. Subjoined is a list of species suitable for cases of ordinary dimensions ; it must, however, be understood that a Ward’s case may consist of a bell-glass, containing a ‘single plant, up to a shut-in window, or area covered with glass, where larger-fronded species may be grown. Humata heteropliylla. Davallia bullata. pentaphylla. Canariensis. Leucostegia hirsuta. — cherophylla. pulchella. Polypodium pectinatum. Schkuhrii. Lepicystis sepulta. squamata. rhagadiolepis. Goniophlebium appendicu- latum. Lopholepis piloselloides. Anapeltis stigmatica. Pleopeltis nuda. Niphopsis angustatus. Drymoglossum piloselloides. Hymenolepis spicata. Phymatodes longipes. glauca. . Pleuridium juglandifolium. venustum. Selliguea caudiformis. Niphobolus Lingua. Drynaria propinqua. Elaphoglossum piloselloides. vestitum. Hymenodium crinitum, Rhipidopteris peltata. Microstaphyla bifurcata. Psomiocarpa apiifolia. Stenosemia aurita. Gymuopteris quercifolia. Platycerium alcicorne. Xiphopteris serrulata. Gymnogramma tomentosa. Calomelanos, —— Martensii. chrysophylla. Peruviana, var. argyro- phylla. pulchella. Llavea cordifolia, Hemionitis palmata. cordifolia. 7 Dictyoxiphium Panamense, Ceratopteris thalictroides. Meniscium simplex. Goniopteris asplenioides. Nephrodium molle. Cyrtomium faleatum, Fadyenia prolifera. Aspidium Pica. Hypoderris Brownii. Trichiocarpa Muorii. Polystichum mucronatum, Lonchitis. —— triangulum. anomalum, denticulatum, 360 Lastrea elegans. concinna. —— strigosa. — vestita. — podophylla. —— erythrosorum, —— Mexicana. —— emula. glabella. — Shepherdi. —— hirta. sancta. hispida. deparioides. Woodsia hyperborea. polystichoides. mollis. Arthropteris albo-punctata. Nephrolepis pectinata. Notholena brachypus. trichomanoides. Marantee. sinuata. sulphurea. Myriopteris lendigera. myriophylla. elegans. tomentosa. vestita. hirta. Cheilanthes viscosa. fragrans. tenuifolia. microphylla. — Alabamensis. —-wmultiida. . - —— argentea. —— farinosa. -—-— pulveracea. capensis. radiata. pedata. FERNS : BRITISH AND FOREIGN. Cincinalis nivea. flavens. pulchella. Pcllea geraniifvlia. ——— intramarginalis. hastata, atropurpurea, Calomelanos. ternifolia. eordata. Platyloma Brownii. Adiantum reniforme. caudatum, lucidum. macrophyllum. — villosum. ° pulverulentum, ‘rigidum. fovearum. curvatum. affine. formosum. hispidulum, setulosum. — fulvum. tenerum, sulphureum. —— Chilense. Capillus. AXthiopicum. Onychium Japonicum. auratum. Pteris Cretica, lineata. longifolia. -—— crenata. heterophylla. semipinnata. argyrea. tricolor. scaberula. Litobrochia denticulata. var. “uar. albo- Litobrochia leptophylla. CULTIVATION, 861 Asplenium viviparium, Doryopteris sagittifolia, viride. pedata. fontanum. collina. flabellifolinm. Blechnum Lanceola. —— pinnatifidum. cognatum. —— macilentum. occidentale. ——- fragrans. longifolium. Diplazium Zeylanicum. campylotis. Antigramma rhizophylla. Doodia blechnoides. Odontosoria tenuifolia. caudata. Microlepia cristata. Lomaria Patersoni. Gleichenia dicarpa. lanceolata. rupestris. L’Herminieri. alpina. Asplenium Hemionitis. speluncer. alternans, — Belangeri. Hymenophyllum species. formosum. Trichomanes species. Brasiliense. Féea spicata. tenellum. Lygodium palmatum. erectum. Anemia tomentosa. marinum. fulva. firmum. adiantifolia. obtusatum. Anemidictyon Phyllitidis. brachypteron. Mohria thurifraga. Todea hymenophylloides. V. Propagation or Ferns. YFVHERE is considerable variety in the way in which _ Ferns reproduce themselves, and we only aid nature when we attempt to increase any particular. species; such is usually effected, and certainly most numerously, by sowing their spores or seeds, as com- monly called, which are contained in cases, as already explained at page 51; they consist of atomic particles, which, under favourable conditions as regards light, 362 FERNS: BRITISH AND FOREIGN. can be seen discharging from the spore-cases like puffs of smoke; therefore Fern seed is said to be in- visible, and to have the power of rendering persons invisible. “I have the receipt of Fern seed and walk invisible.”* Be that as it may, each particle is by the aid of the microscope seen to have definite forms vary- ing in different genera, being globose, oval, or angu- lar, smooth, ‘plain, striated, or echinate, presenting very beautiful microscopic objects. Although these spores (see plate I.) are so small, they nevertheless are endowed with an extraordinary power of, retaining their vitality ; and being easily wafted by currents of air, readily account for the wide geographical range of many species of Ferns. When naturally or artificially placed under favourable conditions, the spore vegetates by expanding in the form of a simple oblong cell, from which other cells are successively produced, ultimately forming a thin green membrane, called the Prothallium, which lies nearly flat, and by the aid of fine spongioles attaches itself to the surface, and when arrived at full size it is of a reniform cordate shape, or sometimes bilobed, or obcordate, varying from about { to } inch in diameter, having much the appearance of a small foliaceous lichen. But before proceeding to describe the method of sowing and rearing, it is important that the seed sower should be made acquainted with the remarkable discovery made, in 1848, by Count Leszezyc Suminski, that Ferns... possessed urgans analogous to stamens and pistils of flowering plants, and that these were produced on the Prothallium, In the progress of growth peculiar cells * Shakespeare. CULTIVATION. 363 are formed on its under. surface, of two kinds, one called Antheridia, and the other Archegonia ; the first (of which there are generally between 30 and 40) con- taining round cells, called sperm cells, which contain vermicular spermatozoids that issue from the cells when arrived at maturity. The cells of the Archegonia differ considerably in their structure from the Anthe- ridia; they contain an embryonal sac, which at a certain period protrudes from the cell, and with which the moving spermatozoids are said to come in contact, and thus produce fertilization ; soon after this the embryo sac is changed, a bud is formed, and ultimately a young Fern Plant is gradually developed, and in time the Prothallium decays. This singular phenomenon excited considerable interest, and has been carefully inves- tigated by the late Professor Henfrey and other eminent microscopists.* Although the above is only an outline of the prin- cipal points of this curious subject, yet it will be suffi- cient to draw the attention of Fern seed sowers to the subject, and materially assist them in understand- ing, and arriving at the cause of the many anomalies in the good and ill success attending the raising of Ferns from seed. Much has, however, yet to be investigated before any rule can be arrived at ; such as why the Desmobrya division does, asa general rule, pro- duce plants from seed in the greatest abundance, even to some species becoming the weeds of the hothouse ; while, on the other hand, those of the division Hremo- brya are comparatively few, and may be considered the exception and barrenness of this rule. The differ- * Henfrey, in Linnean Transactions, vol. xxi, 1853. 864 FERNS: BRITISH AND FORKIGN. ence in the fertility of the spores of these two divi- sions is veryremarkable; this is,it must be remembered, as found with the plants under cultivation, and as with many true Ferns we find, from experience, the spores of the latter seldom germinate in less than eighteen months after having beensown. On the contrary, in the Desmobrya group, many require but two or three weeks, and occasionally certain species of Gymmno- gramma and Cheilanthes will germinate in as many days. From the irregularity, however, in this respect it is in a great measure governed by the state of, and conditions which surround, the seed; from repeated trials with spores of Brainea insignis, Prothallia were produced in forty-eight hours. Consequent on this irregularity, ib is impossible to state what time of the year is most suitable for sowing to ensure success, especially for imported spores, which should be sown as soon as received, provided a proper condition can be given them. Therefore extra attention must be paid when it unfortunately occurs that the young Ferns are in their Prothalliwm stage during the winter season, to keep them from damping and the growth of conferva and musci. When it is desirable to increase any particular species by spores, some precaution is neces- sary to secure them in a proper ripe condition, and without spores of other species growing near being mixed with them. The fructified frond should be taken as soon as the spore-cases on the plant begin to open, ~and placed between sheets of paper in a moderately dry place for a few days, at the end of which time there will be escaped spores in abundance. These should be immediately sown in pots prepared in the following manner ;—four to six-inch pots are generally the most CULTIVATION. 865 convenient size; they should be half filled with good drainage material; with the smallest particles at top, when another fourth of depth should be occupied by fine soil, half sand and peat, one quarter loam, with a sprinkling of finely-broken sandstone or soft brick slightly pressed down on the top; it should -then be watered and time allowed for the whole to become uniformly moist ; then the spores to be very thinly dis- tributed over it, the whole covered with a bell-glass or a piece of glass same size as pot, to be placed on its rim, allowing a space of about one inch between it and the surface of the soil. In order to keep the whole moderately and constantly moist, the pots should be placed in pans of water of half an inch depth, care being taken not to allow the soil to become over saturated ; and whenever any copious condensation takes place on the glass, it should be carefully sponged off. As the spores germinate, and the Prothallia become crowded, so as to touch each other, they should be im- mediately thinned, and if it is desirable to save the thinnings they can be removed in little clumps on the particles of brick or sandstone to other pots prepared as for spores. It is not, however, with all our care in sowing different species in separate pots, that the species sown come up in the pot in which it was sown: plants of it may be found in other pots, or in different parts of the house on moist surfaces. This is easily ac- counted for, as the least motion of the air carries away the spores while in the action of sowing, and indeed all superfluity of spores may be with profit distributed over the whole huuse, the moist walls often affording abundance of young plants. It also often happens that a good crop of Pteris aquilina is the result, its spores 366 FERNS: BRITISH AND FOREIGN. being no doubt latent in the soil used for filling the pots. To avoid this it is advisable to bake the soil before using, and to bring it again to its proper moist state by the use of water that has been boiled. It is said that spores retain their vitality for a number of years; in my experience I have no direct proof of this; but several remarkable. instances of plants making their appearance without the spores of the species having been sown, or even an Herbarium specimen having been seen in this country. In the instance of Lomaria Patersoni, a species originally discovered in Tasmania, which spontaneously made its appearance at Kew in 1830, only one specimen was at that time said to be in the possession of Mr. Brown, at the British Museum, which I never saw; and Allan Cunningham informed me that he never found the plant, and was very much surprised when he saw it growing at Kew. This in time gave specimens to many Herbaria, and living plants to botanic gardens. A similar instance was that of Doodia blechnoides, which made its appearance at Kew in 1835. Other instances might be quoted, such as the appearance-of Asplenium stipitatum,of which two plants spontaneously made their appearance about twenty years ago, and I at first supposed they had originated from the spores from a specimen in my Herbarium of a Luzon plant named by me Neottopteris stipitata; but in time it became evident that the two plants were quite distinct from it, and, like the Zomaria and Doodia, I had never seen native specimens. By what means the spores that produced these plants came to Kew it isimpossible to say. In 1829 I found a plant of Ceterach officinarum growing in a crevice of masonry on one of the CULTIVATION, 867 towers of the New Palace at Kew (since taken down). As this Fern is not found wild near London, it would be useless to speculate where this solitary spore came from ; it seemed however to have found a proper nidus in the crevice, enabling it to germinate and resist all untoward influences, to pass through the Prothallium state and become a plant. Polypodium vulgare and Asplenium Ruta-muraria may be considered our domestic Ferns ; for many years a plant of Polypodiwm vulgare grew on the brick wall separating Hyde Park from Kensington Gardens, and there it remained till the wall was taken down. These few instances of isolated appearances of Ferns readily explain the wide geographical distribution of some species over the surface of the earth. It has been shown that the spores of many species germinate quickly and abundantly, and become fully developed Prothallia, yet it often happens that no plant: bud is formed, and in time the Prothalli« decay; the cause of this has always been supposed to be undue moisture or some atmospheric action not sensible to us, as this has always occurred in certain species of special interest, such as Brainea insignis, the spores of which, as already stated, germinate readily ; yet we have not succeeded in obtaining young plants, not even one Prothallium being seen to makea plant bud. Without special microscopical examination of the Prothallia we can only speculate on the proba- bility, that, as in flowering plants, the whole of the spores of some Ferns (such as Braineu) are unisexual or may even be entirely destitute of both Antheridia and Archegoma, in either case deficient of the elements necessary for the production of a plant bud. £68 FERNS: BRITISH AND FOREIGN. I have at page 65 mentioned that a few intermediate forms of the genus Cy ymnogram: ma had been raised from spores, which are considered sports by some and by others hybrids ; the latter can only be admitted on the supposition of two Prothallia of two different species growing so contiguous to each other that the sperma- tozoids of one Prothallium have the power of passing and fertilizing the Archegonia of the other, and thus produce a hybrid, as in flowering plants. Another point of some practical importance is, that in general only a single plant bud is formed on each Prothallium, This may be supposed to be owing to the vital function of the Prothalliwm not being able to support more, in that respect analogous to only one ovulum being fertilized in ovaries of many flowering plants. Admitting that, then, how are we to explain that on removing the plant bud a new bud is formed; and even as many as eight to ten have been obtained from Prothallia of Hymenodium crinitum, each of which, by proper care, becomes aplant. Then again experiments have shown, that by dividing the Prothallium from the base upwards with a sharp instrument into two or even four parts, each part produces a plant bud. Seeing this, it is reasonable to infer that Prothallia have the power of producing plant buds, analogous to leaves of Begonias and other plants ; but whether such is the case, or each bud is the result of the action of spermatozoids on latent Archegonia, is not known. Then again we have the remarkable instance of the great profusion of plants produced by the spores of all farinose Ferns, such as species of Gymnogramma, Cheilanthes, Nothvlcena, Cincinalis, §c., and also of the smooth ebenous genera Pellea, Llatyloma, Doryo- CULTIVATION. 369 pteris, and Adiantum, while comparatively only a few of the smooth-fronded species of the division Hreme- brya produce plants from spores. This subject yet requires much experimental investigation before satisfactory reasons can be assigned for what is here stated. The majority of Ferns that do not increase by spores, often, however, readily do so by other means, such as by offsets, and viviparous buds, or bulbils produced on the upper surface, on the apex of the fronds, or in the axils of the segments, which, when placed under favourable circumstances, become plants. Ferns of czespitose vernation will occasionally produce buds or crowns laterally on the old caudex, which may be readily separated for propagation with a sharp knife; when the vernation consists of a creep- ing rhizomé, such may be cut in pieces of whatever length desirable, with a bud or growing point in each piece, and, as with the separated lateral crowns, should be placed in as small a pot as convenient, with soil suitable to their kind (selected according to the tule already given for establishing plants), and the whole subjected to an extra close atmosphere till thoroughly established. Up to the present time at- tempts to propagate Ferns by separated portions de- void of any previously joined bud have proved fruitless ; although by some a solitary instance in Scolopen- driwm vulgare is considered sufficient evidence to the contrary, as portions of the base of its fronds, if separated, inserted in soil, and kept close and moist by the aid of a bell-glass, will readily strike; so also with some of the abnormal forms of the same genus, if portions of the margins of their fronds are treated in 28 370° FERNS : BRITISH AND FOREIGN. the same manner. Upon close examination, however, . previously formed embryo buds are observable and, considering, too, its close affinity with the Aspleniums, , the most proliferous. (in the formation of bulbils) of. all Ferns, this cannot be wondered at. To propagate Ferns by the buds produced on their foliage is most. easy. As soon as the bulbil plants have attained a size to be handled conveniently, they should be carefully- taken off and pricked out in pots filled with mode- rately fine soil, and kept covered with a bell-glass till thoroughly rooted, when they. may be potted off in single pots, as required. Species with long, slender, hard sarmentum, such. as Gleichenia, do not root readily when separated ; in- deed, large plants have been entirely destroyed by too » free division of their sarmentum; to prevent this, layers are resorted to, which is accomplished by fixing pro- . longing sarmentums over small pots filled with soil, which, when well rooted, can be. separated with safety, and without injuring the specimen plant. Again, in regard to the division Mremobrya, they are not only remarkable in the sterility of their spores, but also in| not producing viviparous buds; however, the readiness - with which small portions of their rhizomes form plants, and the already described tenacity of life,seem to make them independent or to render less need of: perfect spores or bulbils. In concluding this treatise, I deem it necessary to explain, that, in consequence of the woodcuts occupying more space than was calculated for, and in order to keep the book within a limited size, it has become necessary to considerably reduce the. original manuscript on Cultivation. It is, however, CULTIVATION, $71 hoped that what is now given in the preceding pages will be sufficient to show the nature of Ferns, and the methods adopted for propagating and préserving them in the collections of this country. NOTE. In the preceding article on Cultivation it has been shown that all Ferns are capable of being cultivated in this country under one of three conditions as regards temperature, deter- mined by the nature of the native climate of the different species, which, as regards the species enumerated in the preceding catalogue, I classify as follows :— 1st. Hardy, in the open air. North and Central Europe, including Great Britain and Ireland, North America, North Asia. 2nd. Temperate House. South Europe, Madeira, South Africa, North India, China, Japan, Australia, South of the Tropics, Tasmania, Norfolk Island, New Zealand, Chili, Mexico. A few species of those countries are hardy,—such are marked .H. after the name of the country; some others improve by a higher temperature,—such are marked -Tr. after the name of the country. 8rd. Tropical House. West Tropical Africa, including St. Helena and Ascension, Mauritius, Ceylon, India, Malacca, the 2B2 3872 FERNS : BRITISH AND FOREIGN. F Malayan Archipelago, including Penang, Singapore, Java, and the Philippines, Borneo, the Polynesian and other Islands of the Pacific within or near the Tropics, Sandwich Islands, Tropical America, inclu- ding Venezuela, New Granada, Panama, Peru, Guiana, Brazil, West Indies. Some species from high altitudes within these countries will thrive in the Temperate House,—such are marked T., after the name of the country. 373 A LIST OF AUTHORS AND BOOKS QUOTED IN THIS WORK. N.B.— When no special book is quoted after an author’s name, his writings on feens are to be found in botanical and other scien- tific journals of their time, and which are too numerous to notice in this work,—Those with an asterisk are living authors. *Agardh. J. C. Agardh, Professor of Botany at Stockholm ; author of a “Monograph on the genus Pteris.” At. W.&W.T. Aiton, father and son, Directors of the Royal Botanic Gardens, Kew, from 1760 to 1841 ; authors of 1st and Qnd edition of “ Hortus Kewensis,” 1793 and 1813. A. Rich. A. Richard, a French botanist ; “ Voyage de l’Astrolabe (Botanique).” *Arn. Walker Arnott, Professor of Botany at the University, Glasgow (see Hook. et Arn.). *Bab. C. 0. Babington, Professor of Botany at the University, Cambridge ; an eminent British botanist. *Backhouse. James Backhouse & Son, nurserymen, York; importers and cultivators of Ferns. Bauer. Francis Bauer, a celebrated botanical painter (see Hook. et Bauer). Beauv. Palisot de Beauvois, a French botanist ; author of “Flore d@Oware ;” figures and descriptions of plants in the Bight of Benin, 1810. : Bernh. J. J. Bernhardi, Professor of Botany at Erfurt. Blume. ©. L. Blume, Director of the Botanic Garden, Batavia ; “Enum. Plant. Jav.,” 1830; “FL Jav.,” figures and descrip- tions of the plants of Java. , Bolt. ‘J. Bolton, an English botanist ; “ Fil. Brit.” figures of British Ferns, 1790. 874 FERNS: BRITISH AND FOREIGN. Bory, J. B.@. Bory de St. Vincent, a French traveller and botanist. *Brack. W. D. Brackenridge, Botanist to the United States’ exploring expedition ; Descriptions and figures of the Ferns of the expedition. Braun. A. Braun, Professor of Botany, Berlin ; “ Monograph on Selaginella.” Br. B. Braun, the most celebrated of botanists ; “ Prodromus Flore Nove Hollandie,” 1810; “Observations on Ferns in Wallich’s Plante Asiaticee Rariores ;” Horsfield’s “ Plante Jave.” Br. P. Browne, author of a “ History of Jamaica,” 1756. Brongn. A. Brongniart, a French: botanist. Burm. J. Burmann, a Dutch botanist, and writer on plants of India, Ceylon, &e. Carm. Captain D. Carmichael, a Scotch cryptogamic botanist. Cav. A.J. Cavanilles, Professor of Botany, Madrid. *Colenso. Rev. W. Colenso, a New Zealand botanist. Col. A. Colla, a collector and namer of Chilian ferns. Cunn. A. Cunningham, a celebrated botanist and traveller in Brazil and New South Wales from 1815 to.1830. Dec. Aug. Decandolle, a celebrated French systematic botanist. Desf. M. Desfontaines, a French botanist and traveller in Barbary ; “Flora Atlantica.” Desv. N. A. Desvauc, an eminent French botanist; author of several papers on Ferns, from 1808 to 1814. De Vries. G. H. De Vriese, a Professor of Botany, Leyden ; “Monograph on the genus Angiopteris.” Dick. J. Dickson, an English cryptogamic botanist. Don. D. Don, Professor of Botany, King’s College, London ; “Prodromus Flore Nepalensis.” — - Dry. Jonas Dryander, librarian to Sir Joseph Banks ; a writer on Ferns in the “ Linnzan Transactions.” *Fiat. D.C. Eaton, an American botanist. Ehrhart. F. Ehrhart, a German botanist. Endl. Endlicher, Professor of ‘Botany, Vienna; “ Prodromus Flore ae Eng. Bot. English Botany, “Figures and descriptions of British plants,” by-Sir J. E. Smith a Jas. Scmertie Eschw. F. L. Eschweiler, a German botanist. *Fée.