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DECAPOD CRUSTACEANS OF THE
NORTHWEST COAST OF NORTH AMERICA

BY MARY J. RATHBUN

CONTENTS.

ANMOdUuCtHON’. . Ses setae Nera tee etait Gino are BRck Geo) Vlo-~ &. we alg abacateiars al 2
ere halle ted ATES tu eve & ctrettaee: ora a ahe inlays ws) Wee, «a trose oh aes foueirm, 08h, Gre 4
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PRISCA CODY LAD LOT eS hd, track <a aielans ae weadie a “owes ae niece are Syd <imxeieass 8

SyysbemI atic, MSCUSSION Ol speCleS ice veo cau ssn aeutes sos as ea ose 19g

INTRODUCTION.

THIS report includes primarily the crabs and shrimps collected
by the Harriman Expedition. In order to name the shrimps it
became necessary to overhaul the large collection of undeter-
mined material of that group which has accumulated in the U. S.
National Museum. This was derived from several years’ work
of the A/éatross, and from the investigations of Dr. W. H. Dall
and others, and embraces the entire coast from Arctic Alaska to
southern California. The results of this study are given below,
short descriptions of the new species having already been published
in the Proceedings of the National Museum (XXIV, pp. 885-905,
May, 1902). More than that, the accompanying list of species is
designed to serve as a check-list of the Decapoda’ known to in-
habit the region indicated, together with their distribution.

The collection in the National Museum affords exceptional

1The Anomura of the Museum collection, here listed, have been determined by Dr.

J. E. Benedict. The Alpheide are to be reported upon by Dr. H. Coutiére of the Mu-
seum at Paris; therefore the data given herein are quoted largely from Dr. Holmes.

(3)

4 RATHBUN

opportunities for studying the Pacific fauna, as the material is in
some cases remarkably abundant. So rich in crabs and shrimps
were some parts of the sea bottom explored by the A/dbatross
that only a portion of each dredge haul was preserved for study,
many a ‘peck’ or ‘quart’ of these animals being thrown over-
board by the ship’s naturalists.

Our knowledge of the Decapod fauna of the Northwest is, for
the most part, of recent growth. More than two fifths of the
species enumerated below have been described within the last
twelve years, while during that time our acquaintance with the
species known to Brandt, Stimpson, Lockington, and other pio-
neers in this field has been greatly extended.

In 1900 Dr. S. J. Holmes published a very useful ‘Synopsis
of California Stalk-Eyed Crustacea,’ * with descriptions of genera
and species, and it has been thought unnecessary to repeat cita-
tions for the species which he gives. An effort has been made to
figure all the little-known species.

In addition to the general collection of Decapods made by Dr.
William E. Ritter of the Harriman Expedition, those obtained by
Dr. W. R. Coe and Professor Trevor Kincaid have been placed at
my service.

One new species, a Beteus, is notable as the first Alpheid
reported from as high a latitude as Sitka.

A remarkable case of dimorphism is here recorded for the first
time: the cosmopolitan species Processa canaliculata Leach, more
commonly known by the later name WVzka edulis Risso, is found
to possess in certain cases two chelate feet of the first pair instead
of the characteristic asymmetrical disposition of a chelate foot on
one side and a simple one on the other.

General features.—The Decapod fauna of the North Pacific is
rich in individuals, if not in species or in variations of form.

In the mass of material examined (about 50,000 specimens)
certain types were found to be largely in excess, namely, the Pan-
dalid, Hippolytid, and Crangonid shrimps, the Pagurids or hermit-
crabs, the Maioids or true spider-crabs, and the Lithodids or
Anomuran spider-crabs.

In point of numbers the Pandalids take the lead. The most

1Occas. Papers Calif. Acad. Sci., vil, pp. 1-262, pls. I-IV, 1900.

DECAPODS 5

abundant species are Pandalus borealis and P. montagui, both
boreal forms which descend from the Arctic regions into the At-
lantic as well as the Pacific, but in the latter find conditions most
favorable to their increase. The form of P. montaguz inhabiting
the Pacific exhibits modifications which entitle it to recognition
as a distinct subspecies.

In number of species the genus Sfzrontocaris of the family
Hippolytide is unsurpassed. Like Pandalus, it is primarily a
boreal genus, and is common to the Atlantic and the Pacific. In
the Pacific it is represented by 51 species, exhibiting great diver-
sity inform. Several are identical with Atlantic species.

Quite as conspicuous in the North Pacific fauna are the Cran-
gonidz. These occur in great numbers, and exhibit 32 different
forms, for the most part restricted to the Pacific.

The Paguridz, or hermit-crabs, occur in vast numbers, and some
of the species appear to have local centers of distribution. Each
of these species attains its maximum development, both as to size
and numbers, in a particular area, while elsewhere it is stunted
and infrequent.

Several of the spider-crabs (Maiide) abound in shallow as well
as in deeper water, and Oregonia, Chorilia, the two Chionecetes,
and the two species of Hyas form no inconsiderable part of every
haul of the dredge.

Less abundant are the Lithodide or Anomuran spider-crabs,
which include the giant Decapods of the region.

The crowding of crustacean life in certain localities is especially
favorable to parasitism. Bopyrids (of a few species only) are of
frequent occurrence on many species of shrimps'; Rhizocephalids
are less common, while worm parasites have been noticed in sev-
eral instances embedded underneath the carapace of Spzrvontocarts.

The Decapods contribute without doubt a large proportion of
the food of fishes, several species having been taken from their
stomachs; but our knowledge on this subject is very fragmentary.

Many species are caught for the market, to be used for the
table or for bait. The list of these furnished in ‘The Fisheries
and Fishery Industries of the United States,’ Washington, 1884,

1Miss Harriet Richardson will publish a report on the Bopyridz of the Museum
collection in the Proceedings of the U. S. National Museum during the next year.

6 RATHBUN

is the only one so far published, but it is necessarily incom-
plete.

Geographic distribution.—The following points are brought out
in the accompanying table of distribution:

That Arctic species often continue southward through Bering
Strait along the west coast of Bering Sea to Okhotsk Sea and the
Kurile Islands.

That some of these species may also stretch along the Alaska
shores southward, occasionally to Puget Sound or even farther
south.

That the winter line of floating ice in Bering Sea determines
the northern limit of many species. This line extends approxi-
mately from the neighborhood of Nunivak Island westward just
north of the Pribilof and Commander Islands to the Kamchatkan
shore.

While many species range continuously from this line south-
ward to California, others indicate a division of that stretch of
coast-line into several faunz. So far as the Crustacea are con-
cerned, the vicinity of Kadiak appears to be a boundary between
subregions. Aleutian species, however, are often found out of
their normal region, in the cold glacier-fed bays and sounds of
southeastern Alaska.

The Straits of Fuca and Puget Sound also form a partial boun-
dary between species, partial because, while nineteen species have
Puget Sound for a southern limit, and nine species find here their
northern limit, seventy others run uninterruptedly north and south
of this point.

The vicinity of Monterey Bay, California, is a more striking
barrier to species than those above mentioned, the crustacean
fauna south of that promontory being strongly Mexican or Lower
Californian in character.

In exceptional cases, as in Philyra pisum and Cancer amphiv-
tus, a Japanese species is found to occur in approximately the
same latitude on the American coast, without obvious connection
by way of Alaska.

As is to be expected, the inhabitants of the deeper waters of
Bering Sea (below 500 fathoms) are likely to extend much
farther south in the North Pacific Ocean than the shoal-water

DECAPODS 7.

species which follow the coast-line. The occurrence of some
abyssal forms, as Hymenodora glacialis and H. frontalis, in the
deep pocket (1569 fathoms) east of Prince of Wales Island is
worthy of note.

The following is a list of the genera occurring below the 500-
fathom line. Those whose range extends below 1000 fathoms
are printed in italics:

Pasiphea Pagurus (2 species)
Parapasiphz Parapagurus
Acanthephyra Pristopus
fymenodora Leptolithodes
Pandalopsis (1 species) Lithodes
Spirontocaris (5 species) Munida
Crangon (3 species) Munidopsis
Sergestes Chorilia
Benthesicymus Chionecetes
Gennadas Hyas
Calastacus

Dr. Dall’ divides his Oregonian fauna, which stretches from the
end of the Aleutian chain to Point Conception, California, into
three subfaunal areas, with divisions at Yakutat Bay or Mount St.
Elias, Alaska, and at Cape Mendocino, California.

Professor Nutting,” on the other hand, reasoning from his study
of the Hydroids, emphasizes the effectiveness of Puget Sound as
a faunal barrier and the continuity of the area between that point
and the end of the Aleutian chain.’

It is highly probable that future research will make it possible
to subdivide the coast into several small areas. The accompany-
ing table shows the intricate overlapping of species.

1The Fur Seals and Fur-Seal Islands of the North Pacific Ocean, Pt. III, 539-546,
1899.

2Proc. Wash. Acad. Sci., 111, 161, IgOT.

3 Professor Nutting had not consulted Dr. Dall’s report above cited, in which state-
ments made twenty-three years earlier are considerably modified.

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DECAPODS

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DECAPODS 19

SYSTEMATIC DISCUSSION OF SPECIES.
Suborder MACRURA.
Family PA SIPH AIDA.
Genus Pasiphea Savigny.

KEY TO THE SPECIES OF PASIPHAA.

A. Carapace carinated throughout its length.
B. Telson truncate, not forked or notched .... . . magna.
B’. Telson forked or notched.
C. Branchiostegal spine over the angle of the antero-lateral

sinus . . ional a POceiCd:
Co Branchiostegal spine farther forward, near or on anterior
margin.
D. Telson longer than sixth segment . . . . emarginata.
D’. Telson shorter than sixth segment.
H. Abdomen-carmate . 30 a oo we Sa an a Prtaceps:
E’. Abdomen not carinate . <6 © Cortestana.
‘, Carapace not carinated in its posterior Pale Mee 28 dos ole ayjinis.

PASIPHAZA MAGNA Faxon?

Pasipheaa magna FAXON, Bull. Mus. Comp. Zool., XXIV, 209, 1893; Mem.
Mus. Comp. Zool., XvII, 176, pl. XLV, figs. 2-2c, 1895. Gulf of Panama,
458 fathoms.
Distribution.—From off Point Arena, California, to Gulf of Panama;
265 to 552 fathoms.
Dredged by the A/Jatross at the following localities:

Off Point Arena, California, 455 fathoms, station 3348.

Off the Farallones, California, 552 fathoms, station 3162.

Off San Francisco, California, 276 fathoms, station 3479.

Off Pigeon Point, ‘California, 296 fathoms, station 3112.

Off Monterey Bay, California, 418-456 fathoms, stations 3126, 3127.
Off Point Sur, California, 328 fathoms, station 3186.

The identification of these specimens with P. magna is doubtful, but
as I have not had the opportunity of comparing them with the type,
they are placed here provisionally. Though from the same depth of
water, the integument is much firmer in our examples than in the type,
the carapace is deeper and its dorsal outline less arched, the branchial
ridge is straighter throughout the posterior half of its course, the palm in

the first pair of chelee is longer than the fingers. _
The largest specimen, an ovigerous female (SSS
from station 3348, is 155 mm. long. The tel-

Fic. 1. Pasitthea magna. Tel-
son is a little longer than the sixth segment of son (x 13). Station 3348.

20 RATHBUN

the abdomen, and reaches about to the end of the inner uropod; it is
broadly channeled, extremity truncate or slightly convex, and armed with
about 13 slender bristles. Eyes brown in alcohol. In the first pair of
legs the palm is about one third longer than the fingers; in the second
pair the right merus has 14 spines, the left 13.

PASIPHAZA PACIFICA Rathbun.

Pasiphea pacifica RATHBUN, Proc. U. S. Nat. Mus., XXIV, 905, 1902.

The carapace is a little less than half the length of the abdomen,
measured on the median line; it is laterally compressed and rises into a
well-marked ridge extending nearly to the posterior margin and terminat-
ing anteriorly in the median tooth; the ridge is rounded except on

Fic. 2. Pasipheapacifica. @ (X14). Station 3186.

the forward part of the gastric area, where it forms a thin, sharp keel
terminating in a sharp pointed tooth, which is not nearly so advanced as
the anterior margin. ‘This tooth is variable, being more or less inclined
upward; its terminal portion is usually slender, its anterior margin con-
cave. Median tooth of frontal margin narrow, rounded; infra-orbital
and antennal angles rounded and ill-defined; antero-lateral angle
rounded and slightly obtuse. Behind this angle the antero-lateral
margin runs almost directly backward for a short distance, then turns
abruptly downward, forming an obtuse angle, and joins obliquely the
infero-lateral margin. Above this sinus there is a sharp spine on a level
with the middle of the basal segment of the antenna. There is a well-
defined blunt ridge on each side of the carapace, running from the

DECAPODS 21

hepatic area backward over the branchial region nearly to the posterior
border of the carapace.

Eyes of moderate size, subglobular on somewhat smaller cylindrical
stalks, the cornez of a dark brownish color.

The flattened antennular spine does not reach the end of the basal
segment; both of the flagella exceed the carapace in length. The second
antennz exceed in length the whole body; there is an acute spine on
the peduncle below the articulation of the scale; the scale is long-oval,
armed with a spine at the distal end, and overreaches the antennular
peduncle by a third of its length.

The third maxillipeds extend as far forward as the end of the antennal
scale.

The first pair of legs, when extended forward, surpass the third
maxillipeds by the length of the dactylus; the merus is either unarmed,
or may have one, two, or even three spines; the basal part of the
propodal segment is half again as long as the dactylus. The second
pair of legs are longer than the first by about half the length of the
fingers, and have longer and more slender chele; the second segment
is armed with a small spine at the distal end of its inferior margin. The
third pair reaches forward a little beyond the proximal end of the chela
of the preceding pair; except for the first two segments, they are thread-
like; all the segments are naked and unarmed; the merus is about one
half the length of the whole appendage, and more than twice as long as
the propodus. The fourth pair of legs reaches scarcely to the middle
of the merus of the second pair; the lower margin of the dactylus,
propodus, and distal half of the carpus is fringed with bristles. The
fifth pair, when extended, reach to the middle of the merus of the
second pair; the dactylus is elongate-oval, the margin of its distal half
fringed with long sete.

The second to sixth abdominal segments inclusive are carinated on
the median line. The sixth segment is from two thirds to three fourths
longer than the fifth; it is marked on either side by a
curved longitudinal ridge. Telson four fifths as long C-
as sixth segment; dorsal surface channeled; extremity ee eae oc eay
with a deep notch, bordered by spinules, which in- Station 3455.
crease in size and are continued to the posterior outer angle of each
lobe; the outer spinule is considerably longer than the others and is
very often broken off.

Dimensions.—Length of female, 64.8 mm.; length of carapace, 20.3
mm.

Distribution.—This species occurs sparingly from Unalaska and the

a

22 RATHBUN

Gulf of Alaska southward to the Gulf of California, in 53 to 399 fathoms
(one specimen from 13 fathoms, if the label be correct). The following
are the specific localities of the A/datross represented :

North of Unalaska, 399 fathoms, station 3329.

Portlock Bank, Alaska, 230 fathoms, station 2858.

Off Queen Charlotte Sound, British Columbia, 204 fathoms, station 2861.
Queen Charlotte Sound, British Columbia, 238 fathoms, station 2862.
Gulf of Georgia, British Columbia, 67 fathoms, station 2863.

Strait of Fuca, 135-152 fathoms, stations 3449, 3454-3450.

Puget Sound, 82 fathoms, station 3067.

Off Tahwhit Head, Washington, 178 fathoms, station 3076.

Off Pigeon Point, California, 296 fathoms, station 3112.

Monterey Bay, California, 13-382 fathoms, stations 3134, 3202, 3669.
Off Point Sur, California, 316, 328 fathoms, stations 3188, 3186 (type).
Off San Simeon Bay, California, 53 fathoms, station 3191.

Off San Luis Obispo Bay, California, 252 fathoms, station 3195.

Off Point Conception, California, 278, 284 fathoms, stations 3198, 2892.
Santa Barbara Channel, California, 233-280 fathoms, stations 2840, 2960,

3199, 3200, 3201.
Off San Miguel Island, California, 376 fathoms, station 2896.

Off Santa Cruz Island, California, 266, 269 fathoms, stations 2947, 2948.

Off San Diego, California, 339 fathoms, station 2925.

Gulf of California, northwest of Tiburon Island, 145 fathoms, station
3015. ity : eee
Our species differs from the others with carinated carapace in having

the branchiostegal spine situated above the angle of the antero-lateral

sinus.

PASIPHEA EMARGINATA Rathbun.

Pasiphaa emarginata RATHBUN, Proc. U. S. Nat. Mus., XXIV, 905, 1902.

I have proposed the name P. faxoni for P. acutifrons Faxon, not Bate.
There are in the National Museum specimens which I believe to be P.
acutifrons Bate, from the west coast of Patagonia, 194 fathoms, station
2784, Albatross. FP. faxoni differs from them in the broader median lobe
of the front; in the antero-lateral sinus being an obtuse angle, while in
P. acutifrons it is a right angle or less; in the sixth abdominal segment
compressed, blunt above, scarcely carinate, in P. acutifrons sharply cari-
nate; in the telson much shorter than the sixth segment, in P. acutifrons
very little shorter; in its shallower notch, which is broader than long.

P. emarginata differs from P. faxoni as follows:

The median carina is less sharp and less prominent; the gastric tooth
smaller; the carapace proportionally higher in its posterior portion, the
branchial ridge more sinuous; the carapace relatively longer, being equal
to the first five segments and half of the sixth segment of the abdomen;

DECAPODS 23

the telson is longer than the sixth segment, and at the posterior end is
notched with a very shallow V-shaped
sinus, so shallow that the extremity can
hardly be called ‘ forked.’
Dimensions.— Length of female 81
mm., of carapace 33.3 mm. ;

Distribution.— Santa Barbara Chan-
nel, California, 265 to 322 fathoms (===
(Albatross stations 2903, 2904, 2960, Fic. 4. ae emarginata. 9. Station
3200, and 3201). Gulf of California, ees oie a Hes 8) 5
off Concepcion Bay, Lower California, 857 fathoms, station 3009 (type

locality).

PASIPHA‘A PRINCEPS Smith.

Pasiphaé princeps SMITH, Rept. U. S. Commr. Fish and Fisheries for 1882,
p. 381, pl. v, fig. 2 (1884); of. cit. for 1885, p. 682 (1886).
Pasiphaeia princeps F AXON, Mem. Mus. Comp. Zool., XVIII, 175, 1895.

One large female, 167 mm. long, was dredged by the A/éatross off Sea
Lion Rock, Washington, in 859 fathoms, station 3075; and one small
specimen about 47 mm. long, north of Unalaska, in 399 fathoms, station
3329:

I have at hand only one specimen determined by ProfessorSmith. It was
taken south of Marthas Vineyard in 538 fathoms, at station 2546. It
measures 150 mm. long. It differs from the type in having the carina
of the posterior two thirds of the carapace well marked though blunt, in
the gastric tooth projecting well beyond the frontal margin, the anterior
(or inferior) margin of this spine contiguous with the dorsal surface of
the carapace in advance of the spine, and in having the merus of the first
and second pairs of feet armed with numerous spines (instead of the first
pair unarmed and the second pair few-spined). The antero-lateral sinus
is rectangular, as in the figure of the type.

In five smaller specimens from off Cape Romain, South Carolina, 353
fathoms, station 2626, and two from off Marthas Vineyard, 349 fathoms,
station 1093, the gastric tooth projects only a little beyond the frontal
margin, and the antero-lateral sinus has margins oblique to each other ;
otherwise as in the preceding.

In the specimen from off Ecuador described by Faxon (/oc. cit.) the
merus of the first pair of feet is unarmed.

The Washington specimen agrees quite closely with that from station
2546, excepting that the median carina of the carapace is sharper along
its posterior two thirds. The small individual from Bering Sea may

24 RATHBUN

prove to be distinct; the gastric spine is shaped as in the figured type,
but a little narrower and more ascending; the merus of the first pair
of feet has one or two spines, of the second pair many spines.

Distribution.— North Atlantic, from lat. 39° 56’ 00” N., long. 69° 45’
oo. W;, to lat. 22° 27’ 30 N., long. 77°20! 30" \W ..-349 to 1342
fathoms. North Pacific: Bering Sea, 399 fathoms; Washington, 859
fathoms. Ecuador, 1132 fathoms.

PASIPHEA CORTEZIANA Rathbun.

Pasiphea corteziana RATHBUN, Proc. U.S. Nat. Mus., XXIv, 905, 1902.
Carapace equal to the first five segments and part of the sixth segment
of the abdomen. Gastric tooth dentiform, continued back in a very blunt
carina; tip acute, upper margin nearly horizontal, anterior margin in-
clined slightly forward and upward, not as produced as the anterior

margin of the carapace. Median
Cee Sos tid mG lobe of front triangular, obtuse.

ee Branchiostegal spine small, inserted
Sy ee | near the antero-lateral angle just be-
ow hind the anterior margin. Antero-
“lateral sinus with sides forming an

SS obtuse angle.
25 2 ey The eye-stalks are only slightly

RMN od Sea ed enlarged distally, the cornee reddish

Bice renee |) 8. Telson (Xx 23). brown. ‘The flattened antennular
scale falls short of the end of the

first peduncular segment. End of antennal scale more lobiform than in
LP. pacifica. The merus of the first pair of legs is armed with 3 or 4
spines; of the second pair, with many spines. Distal end of second
segment of second pair ending ina small spine; fingers longer than palm.

Abdomen without median carina. Telson very nearly as long as the
sixth segment, and forked with a deep sinus.

For the rest, almost as in P. pacifica.

Dimensions.— Length of male 64 mm., of carapace 23.8 mm.

Lype locality.—Near Cortez Bank, Calif., in lat. 32° 44’ 00” N., long.
119° 32’ oo” W., 776 fathoms (AZbatross station 3627).

PASIPHAAA AFFINIS Rathbun.

Pasiphea affinis RATHBUN, Proc. U. S. Nat. Mus., XXIV, 905, 1902.
Very closely related to P. corteziana.
Carapace equal to the first four segments and half of the fifth seg-

DECAPODS a5

ment of the abdomen; not carinated behind gastric tooth; outline much
as in the preceding, as are also the eyes and antennz. Second to sixth
abdominal segments inclusive carinate; telson very nearly as long as the
sixth segment, less deeply notched than in Fesiphea corteziana.

The side view strongly resembles that of F pers

——

P. corteziana, from which the carinated ab- *j;=————_———-_>
ey
domen and shorter carapace without median

carina at once distinguish it. Co
Dimensions. —Lengthof male67mm.,length b
Fic. 6. Pasiphea affinis. Sta-
of carapace 22 mm. tion 2919. a. Telson (x 33).

Type locality.—Near Cortez Bank, Calif., Hand of second pair (x2).
lat. 32° 17/ oo N., long. 119° 17/ 00” W., 984 fathoms (Albatross
station 2919).
Genus Parapasiphz Smith.
PARAPASIPH SERRATA Rathbun.

Parapasiphe serrata RATHBUN, Proc. U. S. Nat. Mus., XXIV, 904, 1902.
Carapace and rostrum as long as the first five abdominal somites.
The median carina extends backward to the posterior fifth of the cara-

Fic. 7. Parapasiphe serrata. (xX about 14). Station 2919.

pace or the cervical groove, and forward along the high thin rostrum
which reaches half way along the eye-stalks; the carina is a little concave
at the middle of the carapace and anteriorly slopes downward, and is
armed with 16 small teeth or spines, two of which are on the rostrum and
one is terminal; below this spine the rostrum is a compressed, finely
crenulate lobe. Orbital and antennal angles blunt. There is a small
spine just behind the orbital angle, and another still farther back in a

26 RATHBUN

line with the acicle. The branchiostegal sinus is rectangular, the angle
rounded. From the post-antennal spine a ridge runs nearly straight
back, with one interruption, almost to the posterior margin. A groove
runs from just in front of the middle of the posterior margin vertically
half way down the carapace.

Eyes light brown, almost colorless in alcohol, nearly as wide as their
stalks. The antennular scale, which is foliaceous and has a sharp spine
at the upper extremity, is as long as the basal joint of the peduncle.
Antennal scale nearly half as long as carapace proper, elongate-oval,
with a stout midrib, and its outer border thickened and ending in a
slender terminal spine.

The external maxillipeds, which are stouter than any of the last three
pairs of legs, reach almost to the tip of the antennal scale; all their joints
are setose; the antepenultimate segment is broadened in its middle por-
tion, tapering at each end.

The first two pairs of legs are similar in form, the second pair the
longer and about half the length of the body. In the first pair, the pos-
terior border of the merus and palm are spinous, and the distal angles of
the wrist are produced; the fingers equal the palm in length. In the
second pair, the posterior border of the basis, ischium, merus, and palm is
spinous; the angles of the wrist are more strongly produced; the fingers
are nearly as long as the palm. In each case the hand forms about
two fifths of the entire length. The filiform third pair (tip broken off)
are equal in length to the carapace exclusive of the rostrum. The fourth
and fifth pairs have the terminal joint narrow-oval; the fifth pair are the
longer and two thirds the length of carapace.

The first three abdominal somites are non-carinate, the fourth is pos-
teriorly produced in a thin compressed spine which is carinate and has a
slight notch at its base, visible in profile; the sixth has a longitudinal
groove on either side. The telson is a little longer than the sixth somite,
shorter than the inner uropod, and much shorter than the outer one; it
is truncate at the tip.

Dimensions.—In an egg-laden female the length of the carapace and
rostrum is 23.6 mm., the rostrum 1.8 mm., the abdomen 42 mm.

Dype locality.— Off Cortez Bank, Calif., 984 fathoms (4/datross station
2919), one female.

This species is akin to P. gilesii Wood-Mason, from the Indian Ocean,
which also has a serrated carina running the whole length of the cara-
pace, but the character of the rostrum alone separates it distinctly from
that species. In P. gi/esii, the rostrum is a fine, acute, upcurved spine,
extending about a third of its length beyond the eyes.

DECAPODS 27

Family OPLOPHORID:.
Genus Acanthephyra A. Milne Edwards.

ACANTHEPHYRA CURTIROSTRIS Wood-Mason.

Acanthephyra curtirostris WOOD-MASON, Ann. Mag. Nat. Hist. (6), vil, 195,
1891; IX, 364, fig. 5, 1892. Ill. Zool. Investigator, Crustacea, pl. 11,
fig. 4, 1892.—F AXON, Mem. Mus. Comp. Zool., XVIII, 164, pl. XLIII,
figs. 2-5, 1895.— ALCOCK, Desc. Cat. Indian Deep-Sea Crust., 81, 1901.
Distribution.— Arabian Sea, 937-1043 fathoms; Bay of Bengal, 364—
913 fathoms; Andaman Sea, 922 fathoms (Alcock). From off San Diego,
California, to off Gulf of Panama, 458-2232 fathoms. Off San Diego,
623 fathoms, one male of the typical form (4/éatross station 2929);
Gulf of California, 857 and roos fathoms (Aébatross stations 3009
and 3010); see also Faxon, oc. cit.

Genus Hymenodora Sars.

KEY TO THE SPECIES OF HYMENODORA,

A. Rostrum very short, not reaching beyond the first
antennular segment .. . nice) needs.
A’. Rostrum reaching end of antennular peduncle int aia PRONLAIES.

HYMENODORA GLACIALIS (Buchholz).

paeree Sis ital BUCHHOLZ, Zweite Deutsche Nordpolarfahrt, 11, 279, pl. 1,
ae aan pare G. O. SARS, Arch. for Math. og Naturvid., 11, 241,
1877; Norske-Nordhavs-Exped., Crustacea, I, 37, 275, pl. Iv, 1885.—
SMITH, Proc. U. S. Nat. Mus., VII, 501, 1885 (part); Rept. U. S. Fish
Commr. for 1885, 678, pl. xv, figs. 3, 10, pl. XVI, fig. 5, 1886.
Distribution.— Between Norway and Greenland, lat. 63° to 80° N..,
452 to 1862 fathoms (Norwegian North Atlantic Expedition); Faroe
Channel (Avight Errant), East coast of North America (U. S. Fish
Commission): Lat. 42° 48/ 00” N., long. 50° 55/ 30” W., 826 fathoms (sta-
tion 2428) ; lat. 40° 03/ 30” N., long. 679 27/15” W., 1149 fathoms (station
2535); lat. 38° 19/ 26” N., long. 68° 20! 20” W., 2369 fathoms (Smith) ;
lat. 37° 12! 20” N., long. 69° 36/00” W., 2949 fathoms (Smith). Bering
Sea, south of Pribilof Islands, 1401 fathoms (station 3604), 19. East of
Prince of Wales Island, Alaska, 1569 fathoms (station 2859),2 4. Gulf of
California, 857 fathoms (station 3009); go5 and 1218 fathoms (Faxon).
Gulf of Panama, 1832 fathoms (Faxon). Off Ecuador, 1740 fathoms
(Faxon).

28 RATHBUN

HYMENODORA FRONTALIS Rathbun.

fHymenodora frontalis Rathbun, Proc. U. S. Nat. Mus., XXIv, 904, 1902.

Integument very thin, but firmer than in ZH. glacialis; covered with
fine wrinkles or rugose lines. Carapace and rostrum more than half as
long as abdomen; median carina extending almost or quite to the mid-
dle of the carapace, and advanced in a rostrum which is unusually long
for the genus, being from two fifths to one half as long as the remainder
of the carapace, and reaching the end or a little beyond the end of the
antennular peduncle. The rostrum is a slender, sharp-pointed spine;
distal half slightly curved upward and usually unarmed, basal half armed

Fic. 8. Hymenodora frontalis. % (X about 2). Station 3308.

with from 3 to 6 small spines above, two or three of which are beyond the
line of the orbit. Occasionally there is a single spine on the distal half.
Orbit deep; a tiny spine tips the suborbital lobe. Antennal spine strong ;
from it a sharp carina runs back subparallel to the side margin quite to
the posterior margin. Above this the branchial region bears a blunt
ridge, chiefly longitudinal, but bent down at either end; a deep groove
extends obliquely downward and backward from the orbital sinus nearly
to the lower carina.

The eyes reach to the middle of the basal segment of the antennule ;
they are without pigment, and bear a slender, blunt spine close up to the
cornea on the inner margin. The antennular peduncles are short, the
segments diminishing successively in length; basal scale narrow-oval, as
seen from above; upper flagellum very thick at base. Antennal scale
lanceolate, outer margin nearly straight, tipped with a small spine;
peduncle very stout, attaining the end of the first antennular segment.

DECAPODS 29

The outer maxillipeds do not reach the end of the antennal scale; the
first pair of feet reach to the middle of the scale; the second pair extend
not quite so far, and are more slender; the third pair reach nearly to or
beyond the end of the scale; the fifth pair to the end of the antennal
peduncle. The meral and propodal joints of the last three pairs are
furnished with fine sete.

Abdomen devoid of a median carina or median spines; the sixth seg-
ment is two and a half times as long as the fifth, and between three
fourths and four fifths as long as the telson. The telson is about as long
as the outer branch, longer than the inner branch of the swimmeret; it
is armed with 7 to g spinules on either side and 4 long spines at the
end. Eggs few and very large, the greatest diameter equaling the length
of the fifth abdominal segment.

Dimensions.—Male, length of carapace and rostrum 19 mm., of
rostrum 6.5 mm., of abdomen 32.5 mm.

Distribution.—¥rom Bering Sea to off Monterey Bay, California, and
Kamchatka; 322 to 1771 fathoms.

Taken at the following stations of the A/satross :

North of Rat Islands, Aleutians, 850 fathoms, station 3784.

Southwest of Pribilof Islands, 1625, 1771 fathoms, stations 3308, 3603.

North of Islands of Four Mountains, 1033 fathoms, station 3307.

South of Pribilof Islands, 1401 fathoms, station 3604.

West of Unalaska (type locality), 322 fathoms, station 3327.

North of Unalaska, 399 fathoms, station 3329.

Southeast of Chirikof Island, 695 fathoms, station 3340.

East of Prince of Wales Island, 1569 fathoms, station 2859.

Off Queen Charlotte Islands, British Columbia, 1588 fathoms, station
3342.

Off Destruction Island, Washington, 516 fathoms, station 3343.

Off Sea Lion Rock, Washington, 636-877 fathoms, stations 3070, 3071,
3°74; 3975.

Off Monterey Bay, California, 418 fathoms, station 3127.

Off Copper Island, Kamchatka, 1567 fathoms, station 3783.
Relations.—The rostrum of this species approaches that of 4. rostrata

Bate, but is considerably longer; the pleon is like that of HZ glacialis

(Buchholz), 1. glauca Bate, and 4. mollicutis Bate in being without median

carina and spines.
Family PALZMONIDA.
Genus Palemon Fabricius.

PALAMON RITTERI Holmes.

In this species either one or two of the seven or eight dorsal teeth are
situated behind the rostrum, ‘Three ventral teeth.

30 RATHBUN

Distribution.—From San Diego, California (type locality), to Gulf of
California. Bay of St. Elena, Ecuador (Nobili). Specimens are in the Na-
tional Museum from San Bartolomé Bay, Magdalena Bay, and La Paz
Harbor, Lower California, and Carmen Island, in the Gulf of California;
all of these were collected by the A/batross.

PALAZZMONETES KADIAKENSIS Rathbun.

Palemonetes kadiakensis RATHBUN, Proc. U.S. Nat. Mus., XXIV, 903, 1902.

Rostrum about as long as the carapace; it may be a little longer ora
little shorter; nearly horizontal, very slightly upcurved, dorsally 6- or

sooo,
os ==,

==

—— SS
H | ~
Nee ph hl oI teh see
ae S

a See ———a)) Le

Re “SSeS Ys

= tS

SSS

<7 / 7 (4

Fic. 9. Palemonetes kadiakensis. 2 (X 22). Kadiak.

7-serrate, one tooth behind the orbit, ventrally 2- or 3-serrate. Sub-
orbital angle rounded, a spine just below it, also an antennal spine;
antero-lateral angle rounded.

Eye wider than the stalk, light-colored in alcohol, a small black ocellus
above near its margin. Antennular peduncle reaches about four fifths
the length of antennal scale, basal joint the longest, having an outer scale
which projects along one third of the second joint, and is tipped at its
outer distal angle with a small spine. The outer filament of the outer
flagellum is over half as long as the body; the inner filament is very
short, equaling in length the last segment of the peduncle; inner flagel-
lum about three fourths as long as outer.

DECAPODS 3I

Antennal scale as long or nearly as long as the carapace, oblong, the
blade projecting considerably beyond the scale; preceding segment
armed with a small spine at the outer base of the scale; peduncle not
quite reaching end of first antennular segment; flagellum one and a third
times the length of the body.

Outer maxillipeds very slender, reaching when extended only a small
bit beyond the antennal peduncle. Feet of first pair reaching nearly to
end of scale, carpus longer than merus, enlarged distally, and more than
twice as long as propodus, fingers subequal in length to palm and covered
with hair. Feet of second pair reaching beyond scale by half the length
of propodus; carpus one and a half times merus, propodus three fourths
length of carpus and no wider than in the first pair; fingers shorter than
palm. The third to fifth pairs of feet increase successively in length by
about half the length of the dactyli; the fourth pair reaches just to the
end of the scale; dactyli slender and contained about three times in
their propodi, which are sparingly spinulous.

Sixth abdominal segment twice as long as fifth, and nearly as long as
the telson. Swimmerets longer than the telson, the outer branch longer
than the inner.

Dimensions.—Length of 2 39 mm., length of carapace and rostrum
15 mm., of rostrum 7 mm,

Type locality.—Kadiak Island, Alaska, under stones at low water;
William J. Fisher, collector.

The abdomen is much longer than in P. vulgaris Say of the Atlantic
coast of North America, the sixth segment being one third again as long
as in P. vulgaris with equal carapace; the rostral teeth are fewer, the
acicle larger, the chelipeds of the second pair more slender. From P.
varians Leach it differs in its longer rostrum, shorter feet of the second
pair, in which the relative length of the segments also is quite different.

Genus Urocaris Stimpson.
UROCARIS INFRASPINIS Rathbun.

Urocaris infraspinis RATHBUN, Proc. U.S. Nat. Mus., XXIV, 903, 1902.
Closely allied to U. longicaudata Stimpson of the West Indian region.
Carapace and rostrum equal in length to the first three segments and

half of the fourth segment of the abdomen. Rostrum not reaching

end of second antennular segment, convex above, armed with 5 to 7

teeth above, 1 or 2 small teeth below near tip, tip acuminate; behind the

rostrum a median gastric spine. Suborbital angle blunt. Antennal and
hepatic spines of good size. Eyes two thirds as long as first antennular

32 RATHBUN

segment, a minute black ocellus above and toward the outside, and be-
yond the limit of the cornea.

Fic. 10. Uvrocaris infraspinis.. % (X 33). , Locality unknown. a. Side. 6, Dorsal
view of anterior portion.

Antennular peduncle nearly as long as carapace, exclusive of rostrum ;
first segment broad, its thin outer margin armed with a spine on the basal
portion and another at the extremity; the inner angle of the lateral ex-
pansion is advanced beyond the articulation of the second joint; second
and third segments subequal and together nearly equal to the first. The
acicle is oblong, extremity oblique, produced at inner angle away beyond
outer spine, which is in line with end of antennular peduncle. The an-
tennal peduncle scarcely reaches end of first antennular segment; fla-
gellum as long as body.

The outer maxillipeds extend to the middle of the last joint of the an-
tennal peduncle. The first pair of feet reach the spine of the acicle;
merus, carpus, and propodus subequal, palm and fingers subequal. The
second pair of feet are as long as the distance from the end of the ros-
trum to the posterior end of the first abdominal somite; the ischium,
merus, and carpus subequal, palm a little shorter, swollen; fingers nearly
aslongas palm. Last three pairs of feet reaching about to end of acicle ;
dactyli short, with a slender spine on the under side.

Abdomen geniculated at the third segment. Posterior margin of all
the segments truncate. Sixth segment shorter than carapace (rostrum
excluded), less than twice as long as fifth segment, and a little longer than
the telson. Telson with two pairs of lateral spines, the outer uropods
much longer than the telson, the inner uropods intermediate in length
between the outer pair and the telson.

Dimensions.—Length of carapace and rostrum of ovigerous @ 6 mm.,
of rostrum 2.5 mm., of abdomen rs mm.

DECAPODS 33

Type locality.—Gulf of California, in Concepcion Bay, Lower Calif.,
two @ (Albatross).

Distribution.—Also taken at San Diego Bay, California, 3 fathoms
(Albatross station 3567); Gulf of California, off San Josef Island, Lower
California, 8 fathoms, station 3006; Guaymas, Mexico, inner harbor
(P. L. Jouy, collector). One lot without label was associated with Crangon
nigromaculata Lockington.

Relations.—This species, while having the general appearance of U.
longicaudata, is markedly different: U. /ongicaudata has no antennal spine
on the carapace; UV. infraspinis has a distinct ocellus outside the cornea,
U. longicaudata has not; in U. infraspinis the palm and fingers of the
first pair are subequal; in U. Jongicaudata the palm is longer than the
fingers; in U. infraspinis the carpus of the second pair is subequal to
the merus, the palm is a little shorter, and the fingers still shorter; in U.
longicaudata the carpus, palm, and fingers are subequal to one another
and shorter than the merus; in our species the sixth abdominal somite
is shorter than the carapace, and less than twice as long as the fifth; in
U. longicaudata the sixth segment is as long as the carapace (rostrum
excluded) and twice as long as the fifth.

Family PONTONIIDA.

Genus Pontonia Latreille.

PONTONIA CALIFORNIENSIS Rathbun.

Pontonia californiensis RATHBUN, Proc. U.S. Nat. Mus., XXIV, 902, 1902.
The carapace and rostrum exceed in length the first six segments of

the abdomen. Rostrum more than one third the length of the re-

mainder of the carapace, very narrow throughout, deflexed, reaching to

_ the middle of the second segment of the antennular peduncle. A tooth

on the anterior margin just above the base of the antenna. ‘The eyes
reach to the middle of the first antennular segment. The lateral expan-
sion of this segment is broad behind and narrow in front, terminating in
a short spine at the anterior angle; second segment one and a half times
as long as third, both together half as long as first. Scale about two fifths
as long as carapace, scarcely exceeding the antennular peduncle, the
spine of the scale about as produced as the blade. Antennal peduncle
reaching to end of scale.

Feet of first pair extending beyond the scale by the length of the
propodus and nearly the whole of the carpus; merus and carpus subequal
and each twice the ischium; propodus a little shorter than carpus; palm

34 RATHBUN

and fingers subequal. The right foot of the second pair is missing; the
left has a short, stout merus, carpus cup-shaped, with an inner distal
tubercle, and a tubercle near the middle of the distal
margin; palm and fingers subequal in length, palm
two thirds as broad as long, inner edge marginate,
fingers gaping, prehensile edges denticulate, fringed
with long hair. Dac-
tyli of last three pairs
of feet with a sub-
terminal and a ter-
minal spine.

The telson is twice
\ ‘. as long as the pre-

J ceding segment, and
has two pairs of

) long lateral appressed
~" spines inserted on

the anterior half.
The outer uropod is as long as the tel-
son, the inner uropod a little longer.

Dimensions.—Length of carapace
and rostrum 6.7 mm., of rostrum 1.8
mm., of abdomen g mm.

Type locality. — Off Santa Cruz Is-
land, Calif., 30 fathoms (Adbatross
station 2945), one female.

Fic. 11. Pontonia californiensis (X 49) This is the only Fontonia described

ation 2945.
from the west coast of North America,
the P. margarita of Smith being a Conchodytes.

Genus Periclimenes Costa.
PERICLIMENES TENUIPES (Holmes).

Anchista tenuipes HOLMES, Occas. Papers Calif. Acad. Sci., VII, 216, 1900.
Not P. tenuipes Borra-
daile, which may be
called P. borradailei,
nom. nov.

Distribution.— From San-
ta Catalina Island, Califor-
nia (Holmes), to Gulf of
California. The Albatross

7 Fic. 12. Pevriclimenes Meneies (x 4). Concepcion Bay.
collected specimens’ at a. Carapace. 4. Foot of second pair.

DECAPODS 38

Concepcion Bay, Lower California, and at stations 2824 and 282s in the
Gulf of California, 7 and 8 fathoms.

They agree with Dr. Holmes’s description, except that the fingers of
the second pair of feet are only a little over half as long as the palm and
the postero-lateral angle of the sixth abdominal segment is subacute.

Family PANDALIDA,
Genus Pandalus Leach.
KEY TO THE SPECIES OF PANDALUS.

A. Third segment of abdomen in part compressed and carinated, the
carina forming a more or less well defined lobe or spine in front
of the posterior margin.

B. Third and fourth segments of abdomen armed with a median spine

on posterior margin. . . . borealis.

B’. Third and fourth segments of abdomen. without median spine on
posterior margin.

C. Rostrum unarmed on distal half of superior margin . goniurus.

C’. Rostrum with spines on distal half of superior margin. jordani.

A’, Third segment of abdomen not compressed and carinated, and with-

out a median lobe or spine in front of posterior margin.
B. Dorsal spines not reaching behind middle of carapace.
C. Sixth abdominal segment more than twice as long as wide.
D. Carapace and abdomen covered with short transverse rugose
Imes... wo ue) ww) we a POD LOCEIAES:
D’. Carapace and abdomen smooth - « « . montagui tridens.
C’. Sixth abdominal segment less than twice as long as
Wide ox: 4 - « « » platyceros.
B’. Dorsal spines extending behind middle of carapace.
C. Dorsal spines more than 15 (17-21). . . . . .« Aypsinotus.
C’. Dorsal spines less than 15.
D. Rostrum one and a half or more than one and a half times as
long as carapace. a = SUrneyt.
D’. Rostrum less than one and a 1 half times 4 as long as Carapace.
E. Antennal scale very narrow, the terminal half of the blade
narrower than the adjacent thickened portion stenolepis.
E’, Antennal scale of moderate width, the terminal half of the
blade not narrower than the adjacent thickened por-
TOMEI Sine: < grea tha et are es ean bath gO AMES

PANDALUS BOREALIS Kréyer.

Pandalus borealis KROYER, Naturh. Tidsskrift, 11, 254, 1838; (2) 1, 461,
1845 ; in Gaimard’s Voyage en Scandinavie, en Laponie, etc., pl. VI, fig. 2.
STIMPSON, Jour. Boston Soc. Nat. Hist.,v1, 501 [61],1857; Ann. Lyc. Nat.
Hist. N. Y., X, 128, 1871.—SMITH, Trans. Conn. Acad. Arts Sci., v, 86,
1879. —BIRULA, Ann. Mus. Zool. Acad. Impér, Sci. St. Pétersbourg,

1897, p. 420 [16]; 1899, pp. 22 [3], 28 [9].

36 RATHBUN

Surface of body smooth and naked. Rostrum about one and three
fourths times as long as the carapace, slightly arched above the eyes,
terminal half slightly ascending; slender, armed with 12 to 16 teeth above
(including 3 or 4 on the carapace), of which all but the one, two, or three
most anterior are movable, and 6 to g distant immovable teeth below,
diminishing in size anteriorly; tip bifid, upper tooth the smaller. The
posterior of the dorsal spines lies between the middle and the anterior
third of the carapace; the anterior is situated at least as far forward as
the distal third of the rostrum.

Antennal spine strong, pterygostomian slender, feeble. Eyes large,
pyriform.

Peduncle of antennule reaching to the middle of antennal scale;
second and third joints subequal; outer flagellum twice as long as cara-
pace, its basal third thickened; inner flagellum longer than outer. An-
tennal scale seven eighths as long as carapace, the blade obliquely
rounded at the end and exceeding the spine.

Antennal peduncle extending to end of second joint of antennular
peduncle; flagellum longer than the body. Outer maxillipeds reaching
to distal fourth or nearly to end of scale. First pair of feet shorter, but
overreaching middle of scale. Of the second pair of feet, the right is
one and a half or one and two thirds times the length of carapace; its
carpus is divided into 25 to 27 segments: the left is slenderer and one
fourth or one third again as long as the right; its carpus has about 58
segments. The third, fourth, and fifth diminish slightly in length in the
order named, the third pair overreaching the scale by at least the length
of its dactyl. The dactyli are slender, and are contained in the propodi
from three to three and a half times.

The terminal portion of the third segment of the abdomen is com-
pressed and carinated; a sharp spine or lobe pointing backward is situ-
ated at the posterior third of the segment; posterior margin produced
backward in an acute lobe terminating in a small spine. A similar spine
terminates the truncate fourth segment. Sixth segment elongate, about
twice as long as fifth; seventh armed with 7 to 10 aculei on each side,
and 4 terminal, of which the submedian pair are slenderer and about
half as long as the outer pair.

Dimensions.—Ovigerous female, length 13.5 cm., length of carapace
and rostrum 64 mm., of rostrum 41 mm.

Females carrying eggs were taken August 5th and 6th in Bering Sea
(stations 3528, 3530).

Distribution.— Circumpolar. Bering Sea and North Pacific southward
on the American coast to Columbia River; 29% to 350 fathoms.

DECAPODS 37

Okhotsk Sea (Brandt). On Atlantic coast of North America from
Greenland southward to Massachusetts Bay, 40 to 160 fathoms. Scan-
dinavia.
Juneau, Alaska, 50 fathoms (Harriman Expedition).
Taken by the A/datross at the following localities:
Bering Sea, southwest of St. Matthew Island, 59-77 fathoms, stations
3539 3532.
Bering Sea, off Pribilof Islands, 39-184 fathoms, stations 3309, 3439-
3442, 3482-3489, 3491-3497, 3500, 3511, 3523, 3524, 3526-3528,

3533-3536, 3538, 3539) 3544, 3553-3556, 3559-3561, 3602, 3605,
3606, 3609, 3610.

Between Bristol Bay and Pribilof Islands, 2914-36 fathoms, stations 3252,
3253, 3306.

North of Rat Islands, 270 fathoms, station 3785.

North of Umnak Island, 49 fathoms, station 3537.

North of Unalaska, 61-350 fathoms, stations 3226 (1 quart rejected), 3316
(309 fathoms, 1 specimen), 3317, 3318, 3331 (350 fathoms, 1 speci-
men).

Chernofski Harbor, Unalaska, 109 fathoms, station 3324.

Pumicestone Bay, Unalaska, 54 fathoms, station 3321.

Iliuliukk Harbor, Unalaska, 50-93 fathoms, stations 3310, 3311, 3313;
3314 (1 quart rejected), 3334-3336.

Off Akutan Island, 56-91 fathoms, stations 2841, 2842, 3548, 3549.

Off south entrance to Akutan Pass, 45 fathoms, station 2843.

Off Aektok Island, 54 fathoms, station 2844.

Northwest of Unimak Island, 41-121 fathoms, stations 3224 (% peck
discarded), 3225 (2 bushels rejected), 3257 (1 peck rejected), 3258,
3259, 3263 (2 quarts rejected).

Unimak Pass, 34 fathoms, station 3220.

North of Unimak Island, 49 fathoms, stations 3256, 3542.

Davidson Bank, 42 fathoms, station 2845.

South of Unimak Island, 61 fathoms, station 3216.

Off Kudobin Islands, 36-53 fathoms, stations 3278, 3279 (1 pint rejected),
3280,-3202, 3263.

Shumagins, 48-110 fathoms, stations 2847, 2848 (abundant), 2849, 2852.

South of Trinity Islands, 67 fathoms, station 3341.

Off Sitkalidak Island, 60 to 69 fathoms, stations 2854, 2855.

Gulf of Georgia, British Columbia, 67 fathoms, station 2863.

Strait of Fuca, 40-152 fathoms, stations 3445, 3446, 3451, 3453, 3455)
3458-3462, 3464, 3465, 3596, 3597-

Washington Sound, Strait of Fuca, 48 fathoms, station 2864 (very numer-
ous).

Puget Sound, 82 fathoms, station 3067.

Admiralty Inlet, Puget Sound, 40 fathoms, station 2865.

Off Cape Flattery, Washington, 38 fathoms, station 2872.

Off Columbia River, 68 fathoms, station 2882.

Station 3675 (locality not given), 110 fathoms.

Unalaska (Brandt),

38 RATHBUN

PANDALUS GONIURUS Stimpson.
Plate 1, fig. 3.

Pandalus goniurus STIMPSON, Proc. Acad. Nat. Sci. Phila., x11, 36 [105],

eee MuRDOCH, Proc. U. S. Nat. Mus., VII, 519, 1884; Marine
Invertebrates, in Rept. International Exped. to Pt. Barrow, Alaska, 141,
pl. 1, figs. 2-2c, 1885. RATHBUN, The Fur Seals and Fur-Seal Islands
of the North Pacific Ocean, Pt. 111, 557, 1899.

Allied to P. dorealis. Rostrum one and a half times as long as cara-
pace, terminal half ascending, posterior half horizontal; armed above
with 8 to 9 movable spines, including 3 on carapace; the anterior spine
behind middle of rostrum, the posterior spine in front of middle of
carapace; lower limb deeper in front of eye than in P. dorealis, and
armed with 6 or 7 immovable spines; tip bifid, upper tooth smaller.

Antennular flagella shorter than in P. dorealis, the inner and longer
one being one and a half times as long as carapace. The spine at the
antero-lateral angle of the antennal scale extends nearly to the end of
the blade. Antennal peduncle falls a little short of end of second joint
of antennular peduncle; flagellum nearly as long as body.

Carpus of right foot of second pair divided into 18 to 20 articles; the
left foot nearly one and a half times the right and with 51 to 54 articles.

The third segment of the abdomen possesses a lobe as in P. borealis,
but the lobe is blunt, not sharp; the median line in front of the lobe is
nearly horizontal; the posterior margin is only slightly produced back-
wards at its middle; and it, as well as the fourth segment, is devoid of a
median spine. ‘The sixth segment is a little shorter than in P. dorealis,
seventh segment bears 5 to 6 lateral spinules.

Otherwise as in P. dorealis.

Sometimes occurs with P. Jorealis, though an inhabitant of shallower
water. They are easily distinguished by the presence of spines on the
anterior half of the top of the rostrum in P. dorealis and the absence of
the same in P. goniurus; and by the blunt rather than sharp lobe on the
third abdominal segment in the last-named species, as well as the
absence of median spines from the posterior margin of the third and
fourth segments.

Dimensions.—Large female: Length go mm., length of carapace and
rostrum 41.5 mm., of rostrum 25.2 mm.

Distribution —Ranges from the Arctic coast of Alaska southward to
Okhotsk Sea on the one side and Puget Sound on the other, in 3 to 100
fathoms. Its occurrence below so fathoms is exceptional.

Taken by the Albatross at the following localities;

DECAPODS 39

Bering Sea, off St. Matthew Island and Pribilof Islands, 21-59 fathoms,
stations 3439, 3513, 3514, 3523, 3524, 3531, 3536, 3611.

Off Nunivak Island, 17-24 fathoms, stations 3516, 3517.

Off Cape Newenham, 17 fathoms, station 3247 (1 gallon rejected).

Off Hagemeister Island, 414-174 fathoms, stations 3243, 3244, 3246.

Kulukak Bay, 11-14% fathoms, stations 3240-3242 (% peck rejected).

Off Kulukak Bay, 15-17 fathoms, stations 3300, 3301 (2 quarts rejected).

Bristol Bay, 5-18 fathoms, stations 3233, 3234, 3236, 3238.

Off Bristol Bay, 2514-36 fathoms, stations 3251-3253, 3302 (1 quart
rejected), 3303, 3306, 3501.

Off Cape Menshikof, 24 fathoms, station 3296.

Off Cape Strogonof, 26-32 fathoms, stations 3291-3294.

Herendeen Bay. lame

Off Cape Seniavin, 35-37 fathoms, stations 3285, 3286.

Off Kudobin Islands, 36-39 fathoms, stations 3281, 3283 (1 pint
rejected).

Off Cape Leontovich, 18-22 fathoms, stations 3275, 3276.

Off Amak Island, 19-39 fathoms, stations 3273, 3274.

Off Unimak Island, 32—34 fathoms, stations 3256, 3259, 3267.

Pumicestone Bay, Unalaska, 54 fathoms, station 3321.

Gulf of Georgia, British Columbia, 67 fathoms, station 2863.

Strait of Fuca, 48-53 fathoms, stations 3460, 3465.

Washington Sound, Strait of Fuca, 48 fathoms, station 2864.

Bellingham Bay, Washington, 11 fathoms, station 3612.

Admiralty Inlet, Puget Sound, 40 fathoms, station 2865.

Off Cape Nalacheff, Kamchatka, 39-42 fathoms, station 3781.

Avacha Bay, Kamchatka, 16 fathoms, station 3642.

Southeast coast of Kamchatka, g6—100 fathoms, stations 3643, 3644.

Off Robben Island, Okhotsk Sea, 18-28 fathoms, stations 3646, 3647,
3649, 3650.
Collected by W. H. Dall:

Off Cape Sabine, 13 fathoms.

Hagemeister Strait, 8-15 fathoms.

Iliuliuk Harbor and Captains Harbor, Unalaska, 3-15 fathoms.

Shores of Amaknak Island.

Popof Strait, Shumagin Islands.

Chugachik Bay, Cook Inlet, 20-60 fathoms.

Other specimens in the National Museum are from:

Ten miles west of Point Franklin, Alaska, 13% fathoms (Point Barrow
Expedition, types of P. dapfifer Murdoch).

Bering Strait (Dr. Robert White).

Lat. 66° 12’ N., long. 168° 54’ W. (Lieutenant G. M. Stoney, U.S. N.).

Lat. 63° 50’ N., long. 167° 21/ W., 17 fathoms (Lieutenant G. M.
Stoney, U.S. N.).

Off mouth Yukon, 3% fathoms (E. W. Nelson).

Bering Island (N. Grebnitzky); some specimens found in stomach of
Gadus macrocephalus.

Petropavlovsk, Kamchatka (L. Stejneger).

Avacha Bay (Stimpson).

4O RATHBUN

PANDALUS JORDANI Rathbun.
Plate 1, fig. (3:

Pandalus jordani RATHBUN, Proc. U. S. Nat. Mus., XXIV, 900, 1902.

Surface smooth and shining. Rostrum about one and three fourths
times as long as carapace, slightly arched above the eyes, terminal two
thirds ascending; slender, armed with 14 to 17 spines above, includ-
ing 4 on the carapace; distal three immovable, others movable; closer to-
gether above the eye; 7 to 10 immovable spines below, extending nearer
to tip than superior spines; tip acuminate; the posterior of the dorsal
spines lies between the middle and the anterior third of the carapace.

Antennal spine strong; pterygostomian small, slender. Eyes large,
pyriform.

Peduncle of antennule reaching to middle of antennal scale; second
and third joints subequal; flagella subequal, one and a half times as long
as carapace, thickened portion of outer flagellum extending half its length
beyond the antennal scale. Scale three fourths to four fifths as long as
carapace, extremity of blade broadly rounded, and equally produced with
the spine; antennal peduncle reaching to end of second segment of an-
tennular peduncle; flagellum one fourth longer than body.

Outer maxillipeds slender, not reaching end of acicle. First pair of
feet a little shorter; of the second pair, the right foot extends about to
end of scale, the left may be half again as long; the nght carpus has 19g
to 22 segments, the left 58 to 63 segments. The last three pairs of feet
diminish a little in length successively, the third pair exceeding the scale
slightly; their dactyli are elongate, and are contained from two and a
half to two and four fifths times in their propodi.

The third segment of the abdomen is compressed and its posterior
half carinated, the carina interrupted by a slight lobe at the posterior third
of the segment; this lobe is of varying size, at no time strong, and hav-
ing a tendency to disappear in large specimens, where it may occur only
as a slight unevenness in the curve seen in profile; posterior margin well
produced backward, rounded, unarmed; fourth segment with posterior
margin truncate, entire; sixth about twice as long as fifth and three
fourths as long as seventh, which has ro to 13 lateral aculei on each
side.

Females bearing eggs taken in January and February off southern
California.

Dimensions.—Ovigerous female, length 124.5 mm., length of cara-
pace and rostrum 59.5 mm., of rostrum 38 mm.

DECAPODS 41

Distribution.—From Unalaska to southern California, 35 to 178
fathoms :
Monterey and San Pedro, California (D. S. Jordan, 1880),

Taken by the 4/éatross at the following localities:

Iliuliuk Harbor, Unalaska, 85, 68 fathoms, station 3311 (1 specimen),
station 3313 (1 specimen).

Gulf of Georgia, British Columbia, 67 fathoms, station 2863 (1
specimen).

Strait of Fuca, 98-152 fathoms, stations 3447-3456, 3458, 3459, 3461.

Off Strait of Fuca, 142 fathoms, station 3457.

Flattery Bank, Washington, 77 fathoms, station 3673.

Off Tahwhit Head, Washington, 178 fathoms, station 3076.

Near Flattery Rocks, Washington, 171 fathoms, station 2866.

Off Grays Harbor, Washington, 48-58 fathoms, stations 2870, 3046
(abundant), 3047 (abundant), 3048.

Off Columbia River, 55-68 fathoms, stations 2882 (abundant), 3066.

Off Tillamook Rock, Oregon, 46 fathoms, station 3064.

Off False Tillamook, Oregon, 62 fathoms, station 3090.

Off Siletz Bay, Oregon, 77 fathoms, station 3059.

Off Heceta Bank, Oregon, 93 fathoms, station 3080.

Off Point Arena, California, 51 fathoms, station 3251.

Off Bodega Head, California, 167 fathoms, station 3170.

Off Tomales Point, California, 57 fathoms, station 3175.

Off Drake Bay, California, 35 fathoms, station 3155.

Monterey Bay, California, 56 and 68 fathoms, stations 3666, 3671.

San Luis Obispo Bay, California, 77 fathoms, station 3197.

Off Santa Cruz Island, California, 155 fathoms, station 2949 (type
locality).

Southwest of San Nicholas Island, California, 158 fathoms, station 2898.

Distinguished from /. dorealis, which has a similar rostrum, by the
reduction of the abdominal lobe or hump, and the absence of posterior
median spines on third and fourth segments.

PANDALUS MONTAGUI TRIDENS Rathbun.
Plate 11, fig: 2:

? Pandalus annulicornis RICHTERS, Abh. Senck. Natur. Gesell., XIII, 405,
1884. Bering Sea, N. of Akutan Pass, 70 fath.; Plover Bay.

Pandalus montagui RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. III, 557, 1899.

Pandalus montagui tridens RATHBUN, Proc. U. S. Nat. Mus., XXIV, gol,
1902.

Distinguished from the three foregoing species by its non-carinated ab-
domen devoid of a hump. Rostrum slender, from one and a half to one
and four fifths times the length of carapace. Dorsal spines ro to 12 in

42 RATHBUN

number, all movable, and situated, 3 to 5 on the carapace, and the re-
mainder on the basal half of the rostrum. Posterior spine just in front
of middle of carapace. Inferior spines 6 or 7, rigid. Distal two thirds
of rostrum ascending; tip trifid.

Outer margin of acicle slightly arcuate, blade truncate at tip, spine
stout, reaching to or beyond the end of blade.

The outer maxillipeds may fall short of or exceed the scale. Carpus
of right foot of second pair divided into 20 to 28 segments, of left foot
into about 74 segments.

Dactyli of last three pairs short, being contained from five and a half
to seven and a half times in their propodi. Last pair reaching end of
acicle or beyond.

No median spines on abdomen; third segment produced moderately
backward at the middle, margin broadly rounded at that point. Lateral
aculei of telson 5 to 7.

Dimensions. —Female, station 2842, length 104 mm., length of cara-
pace and rostrum 48.5 mm., of rostrum 30.2 mm.

The largest specimen on the Pacific coast was taken off Point Arena,
California, in 239 fathoms. It measures 110 mm. long, carapace and
rostrum 53 mm., rostrum 34 mm.

Distribution.—From Bering Sea to Point Arena, California, 3-351
fathoms.

Taken by the Adsatross at the following localities:

Off Pribilof Islands, 25-184 fathoms, stations 3482-3491, 3494, 3496,

3497, 359, 3504; 3505, 3536, 3540, 3544, 3552, 3554, 3558-3561,
3602, 3605, 3611, 3637.

Off Rat Islands, Aleutian Islands, 55 fathoms, station 3599.

Off Unalaska, 59-351 fathoms, stations 3236, 3315, 3317) 3319) 333

331.

Pumicestone Bay, Unalaska, 35-54 fathoms, stations 3321, 3322 (1 quart
rejected at each station),

Tliuliuk Harbor, Unalaska, 85 fathoms, station 3311.

Off Akutan Island, 36-91 fathoms, stations 2842 (type locality, abun-
dant), 3546, 3548.

Off south entrance to Akutan Pass, 45 fathoms, station 2843.

Northwest of Unimak Island, 43-70 fathoms, stations 3258, 3262.

Unimak Pass, 34-56 fathoms, stations 3220, 3222, 3223.

Northeast of Unimak Island, 38 fathoms, station 326s.

Davidson Bank, 42-43 fathoms, stations 2845, 3215.

South of Unimak Island, 42-61 fathoms, stations 3216, 3217.

Southwest and south of Sannak Islands, 38-44 fathoms, stations 2846,
3213, 3214 (abundant).

North of Amak Island, 39 fathoms, station 3273.

Off Kudobin Islands, 36-53 fathoms, stations 3279, 3281, 3282.

DECAPODS 43

Shumagins, 21-69 fathoms, stations 2847, 2849-2852.

Off Shumagin Bank, 138 fathoms, station 3339 (1 peck rejected).

Off the Trinity Islands, 67-159 fathoms, stations 2853, 3341.

Portlock Bank, 51-230 fathoms, stations 2856 (abundant), 2857, 2858.
Off Queen Charlotte Sound, British Columbia, 204 fathoms, station 2861.
Near Flattery Rocks, Washington, 171 fathoms, station 2866.

Off Point Arena, California, 239 fathoms, station 3349.

Small specimens were collected by Dr. Dall at Bay of Islands, Adak,

Nazan Bay, Atka, and Iliuliuk Harbor and Captains Harbor, Una-.

laska, in 3 to 80 fathoms.

Puget Sound (T. Kincaid).
Berg Bay, Glacier Bay (Harriman Expedition).
Juneau, 20 fathoms (Harriman Expedition).

I have separated this form from P. montagui Leach of the North
Atlantic on account of its somewhat longer rostrum, which varies from one
and a half to one and two thirds times the carapace, the dorsal spines
terminating behind the middle of the rostrum, while in typical P. monta-
gui the rostrum is from one and two fifths to one and a half times the
carapace, and its dorsal spines reach to or in front of its middle. InP
montagui the tip is bifid; in the subspecies usually trifid.

PANDALUS LEPTOCERUS Smith.

Pandalus leptocerus SMITH, Proc. U. S. Nat. Mus., 111, 437, 1881; Bull.
Mus. Comp. Zool., x, 58, 1882; Rept. U. S.-Commr. of Fish, and
Fisheries for 1882, 367 [23], pl. V, fig. 1, 1884.—A. MILNE EDWARDS,
Recueil Planches Expéd. Travailleur, pl. XXII, 1883.

An elongate species. Surface of carapace and abdomen roughened
with short and irregular transverse, punctate ridges, which give rise to
very short, bristle-like hairs.

Rostrum from about one and a third to nearly twice as long as the
rest of the carapace, and curved very slightly upward; armed above
with 11 to 13 movable spines, of which one is near the tip and usually
only two on the carapace proper, while a considerable space back of
the terminal spine is unarmed; below 6 to 8 immovable spines. The
posterior dorsal spine is considerably in front of the middle.

Outer maxillipeds provided with an exopod; the endopod falls short of
the end of the antennal scale, and the first pair of feet are correspond-
ingly shorter than the maxillipeds. Right leg of second pair reaches
about to end of first pair; carpus has 5 segments, proximal half undi-
vided, and followed by 3 subequal segments, each about as long as broad,
together equal to terminal segment; chela about half as long as carpus.
The left leg of second pair has 52 to 64 segments in the carpus; the

44 RATHBUN

merus and distal end of ischium are also faintly segmented. The last
three pairs of legs are very slender, and have slender, nearly cylindrical
dactyli, which have only a few small spinules beneath near the base, and
are contained between two and three times in their propodi.

Abdomen more slender than in P. montagui tridens. Otherwise as in
that species.

Dimensions. —Female (off Cape Ann, Massachusetts), length 98 mm.,
carapace and rostrum 43.5 mm., rostrum 27.4 mm.

Distribution.—Very common on the Atlantic coast of America from
Nova Scotia to Chesapeake Bay in 15 to 321 fathoms.

One specimen only has been taken in the Pacific, by the A/batross, off
Shumagin Bank, Alaska, 138 fathoms, station 3339.

PANDALUS PLATYCEROS Brandt.

Pandalus platyceros BRANDT, in Middendorff’s Reise in den aussersten Norden
und Osten Sibiriens, Band 11, Zool., Theil I, 123, 1851.—STIMPSON,
Jour. Boston Soc. Nat. Hist., v1, 501 [61], 1857.

Pandalus pubescentulus DANA, Crust. U. S. Expl. Exped., 1, 568, 1852;
pl. XXXVI, fig. 8, 1855.—STIMPSON, Jour. Boston Soc. Nat. Hist., v1,
501 [61], 1857.—KINGSLEY, Bull. Essex Inst., x, 63, 1878.—SMITH,
Rept. Prog. Geol. Survey Canada, 1878-79, B, p. 214.—HOLMES,
Occas. Papers Calif. Acad. Sci., VII, 210, 1900.

Body stout. Carapace covered with a dense, short pubescence.
Rostrum one and a half to one and two thirds times the carapace, pro-
vided with a broad entire laminar crest on each side. Median crest
arising half-way back on the carapace, armed with 14 to 17 spines ex-
tending to middle of rostrum, the anterior 1 to 5 fixed, the rest movable;
usually a solitary spine not far behind the acute tip; lower limb armed
with 7 or 8 fixed spines, diminishing gradually in size; the basal tooth
very large. Posterior part of rostrum deflexed, anterior half or two
thirds ascending, tip above level of carapace. Antennal spine very
strong; pterygostomian small, but well marked.

Eyes large, pyriform; cornea in alcohol light greenish; ocellus black.

Antennular peduncle reaching two fifths the length of acicle, second
and third joints subequal; outer flagellum one half longer than carapace,
its basal half thickened; inner flagellum a little longer; outer basal scale
reaching nearly to end of first joint.

Antennal scale four fifths to seven eighths as long as carapace, oblong,
extremity of blade subtruncate, slightly exceeded by the spine; peduncle
reaching nearly to the middle of the third segment of antennular
peduncle; flagellum stout at base, equaling or exceeding length of body.

Maxillipeds stout, reaching nearly or about to end of antennal scale;
first pair of feet to middle of terminal joint of maxillipeds. Right leg of

DECAPODS 45

second pair reaching to end of maxillipeds, carpus 8- to g-jointed, the
first joint as long as the next 5 or 6, which are subequal, and twice as
long as the last joint; propodus about half as long as carpus; fingers
two thirds as long as palm. Left leg of second pair two fifths again as
long as right; carpus divided into 27 or 28 joints, the first and last joints
about twice as long as the adjoining segment; intervening segments
varying a little in size, but those of the distal half a little larger; pro-
podus as long as last three segments of carpus; fingers three fourths as
long as palm. Third to fifth legs stout, the third reaching about the
length of its dactyl beyond the acicle, the fifth reaching to middle of
acicle; dactyli contained from 4 to 6 times in their propodi.

Abdomen slightly more than twice as long as carapace, smooth, non-
carinate; third segment very slightly produced backward in the middle;
sixth segment short and stout, one and a half times as long as wide;
seventh one and a half times as long as sixth, with six spinules on each
side.

Dimensions. —Ovigerous female (station 3129), length 214 mm.,
length of carapace and rostrum 113 mm., of rostrum 68 mm.

Distribution.— Unalaska (Brandt) to off San Diego, California. Strait
of Fuca at Dungeness, Washington (Dana). The specimens in the
National Museum from north of the Strait of Fuca were found in shal-
low water, while in the Strait and along the coast of California the spe-
cies occurs in considerable depths, from 48 to 266 fathoms.

Taken at the following localities by the 4/datross :

Klinkwan, Prince of Wales Island, Alaska.

Karta Bay, Alaska.

Metlakatla, Annette Island, Alaska.

Mary Island, Alaska.

Otter Bay, Pender Island, British Columbia.

Port Angeles, Washington.

Strait of Fuca, 100-151 fathoms, stations 3445, 3449-3451, 3458.
Washington Sound, Strait of Fuca, 48 fathoms, station 2864.

Off Bodega Head, California, 167 fathoms, station 3170.

Off Farallone Islands, California, 191 fathoms, station 3161.

Off Point Ano Nuevo, 203 fathoms, station 3208.

Off Monterey Bay, 65-204 fathoms, stations 3125, 3129.

Off Point Carmel, 162 fathoms, station 3183.

Off Cape San Martin, 218 fathoms, station 318g.

Off Point Conception, 145 fathoms, station 2893.

Off Santa Cruz Island, 150-266 fathoms, stations 2946, 2948, 2949.
Off San Diego, 124 fathoms, station 2935.

Killisnoo, Alaska (Northern Trading Co.).

Lituya Bay, in stomach of halibut (W. H. Dall).
Hood Canal, near Union, Washington, 20 fathoms (R, W, Doane).

46 RATHBUN

PANDALUS HYPSINOTUS Brandt.
Plate 1, fig. 5.
Pandalus hypsinotus BRANDT, in Middendorft’s Reise in den dussersten Norden
und Osten Sibiriens, 11, Zool., I, 125, 1851. Unalaska.

Body stout. Rostrum a little (one sixth or one seventh) longer than
the carapace, which is strongly arched dorsally. The median crest
reaches nearly to the posterior margin; the superior median spines are
movable, occupy the anterior three fifths of the carapace, and extend an
equal distance on the rostrum, in all numbering 17 to 22. The distal
portion is strongly ascending and unarmed, except at the tip, which is
oblique and armed with three immovable spines; the lower margin is
furnished with 7 to g fixed spines, the posterior ones very large and
hooked; lateral carinz of moderate prominence. Antennal spine of
moderate size, acuminate; pterygostomian spine small.

Eyes pyriform, nearly as broad as long; cornea in alcohol bluish;
ocellus darker.

Antennular peduncle reaching not quite to the middle of the acicle;
second joint a little longer than the third. Outer flagellum two thirds as
long as carapace, the posterior five sixths thickened; inner flagellum
about one fourth longer than outer; basal scale lobiform, not reaching
end of eye.

Antennal scale about four fifths as long as carapace, oblong, extremity
obliquely subtruncate, spine very slightly longer. Peduncle reaching to
end of second segment of antennular peduncle; flagellum nearly as long
as the body.

Maxillipeds in large specimens reaching a little in advance of the acicle;
first pair of feet nearly to end of acicle. Right leg of second pair reach-
ing about to the end of the rostrum, carpus with 19 or 20 segments, the
first and last of which are about equal to the three adjoining segments;
propodus as long as the last four segments of the carpus, fingers two
thirds the length of palm. Left leg of second pair half again as long as
right; carpus composed of about 60 segments, the first and last subequal
to the three adjoining segments; propodus a little longer than the last
three segments of the carpus; fingers nearly as long as palm. The merus
and the ischium of the second pair are also more or less segmented. The
third pair of legs extends beyond the acicle by the length of the dacty-
lus and half or more of the propodus; the fifth pair reaches beyond the
middle of the acicle. Dactyli of last three pairs, in the female, contained
from two and one fourth to four times in their propodi. In the male the
propodi of the third and fourth pairs are a little shorter, are recurved (in
the third pair strongly so), and are narrowed at the extremity.

DECAPODS 47

Abdomen two and a third times as long as carapace, smooth, non-
carinate, third segment very slightly produced backward in the middle;
sixth segment twice as long as wide; seventh one and a half times as
long as sixth, with six spinules on each side.

Dimensions.—Ovigerous female (Herendeen Bay), length 133 mm.,
length of carapace and rostrum 61 mm., of rostrum 31 mm.

Distribution.—From Bering Sea to Strait of Fuca and Kurile
Islands, 3 to 20 fathoms.

Unalaska (Brandt).

Specimens have been examined from the following localities:

Cape Etolin, Nunivak, 8 fathoms; Hagemeister Island, 8 to 15
fathoms; Nazan Bay, Atka, 10 to 16 fathoms; Unalaska, 3 to 6 fath-
oms; Iliuliukk Harbor, Unalaska, 3 to 10 fathoms; Port Levashef,
Unalaska; Popof Strait, Shumagins, 6 fathoms; Shahafka Cove, Ka-
diak; Chugachik Bay, Cook Inlet, between 20 and 60 fathoms; Sitka
Harbor, 15 fathoms (W. H. Dall).

St. Michael, Norton Sound (E. W. Nelson).

Herendeen Bay, Alaska, fine specimens, abundant (A/batross).

Kadiak, dredged (W. R. Coe, Harriman Expedition), “ very bright col-
ored.”’

Port Angeles, Washington (A/batross).

Portage Bay, Alaska (Lieutenant H. E. Nichols, U.S. N.).

Bering Island (L. Stejneger and N. Grebnitzky).

Petropavlovsk, Kamchatka (L. Stejneger).

Rakovaya Bay, Avacha Bay, Kamchatka (A/batross).

Off Iturup Island, Kurile Islands, 18 fathoms (A/éa/ross station 3653).

PANDALUS DAN Stimpson.

Pandalus dane STIMPSON, Proc. Boston Soc. Nat. Hist., vi, 87, 1857;
Jour. Boston Soc. Nat. Hist., VI, 502 [62], pl. XxI, figs. 6, 7, 1857.
Opposite Fort Townsend in Puget Sound.

Pandalus franciscorum KINGSLEY, Proc. Acad. Nat. Sci. Phila., 1878, 94.
San Francisco.

A rather stout species. Surface finely pitted. Rostrum a little
longer than the carapace (from one fifth to one eighth longer), distal
two thirds directed obliquely upward. Median dorsal spines 10 to 12,
movable, about half of them on the carapace, the posterior spine a little
behind the middle; terminal half of rostrum unarmed above, apex
trifid, lower margin 6- to 8-spined, the spines diminishing anteriorly ;
lateral carinze moderately prominent. Antennal spine acuminate;
pterygostomian spine small, but well marked.

Antennular peduncle reaching about half the length of acicle, last two
joints subequal; outer flagellum about seven tenths length of carapace,

48 RATHBUN

basal three fourths thickened; inner flagellum one fourth or one third
longer than outer; basal scale reaching only to middle of first segment,
its distal outer angle almost a right angle.

Antennal scale from three fourths to five sixths as long as carapace,
tapering to the extremity, the laminar part of which is rounded, and much
exceeded by the spine; peduncle reaching to end of second segment of
antennular peduncle; flagellum equaling or exceeding the length of the
body.

Maxillipeds reaching nearly to end of acicle; first pair of feet to mid-
dle of terminal joint of maxilliped. The right leg of the second pair may
slightly exceed or fall short of the tip of the rostrum; merus slightly
annulated; carpus with from 18 to 21 joints, those of the proximal half
very indistinctly marked, the last joint as long as the two preceding; pro-
podus the length of the last four carpal segments, palm longer than digits.
Left leg of second pair one third or nearly half again as long as right
leg; merus and distal portion of ischium faintly annulated; about 60
carpal segments, hand and fingers much as in the right leg. Third leg
of female reaching beyond the rostrum by one third or one fourth the
length of the propodus. Fourth leg reaches one third or one half the
length of the propodus of the third pair, while the fifth may reach beyond
the middle of the
} propodus of the
fourth. Dactyli
of the last three pairs contained about
four times in their propodi. The third
and fourth pairs of legs of the male differ
from those of the female in having
shorter propodi, that of the third scarcely
overreaching the rostrum; the propodus
of the third is also recurved instead of
ates. ee ee Liane as straight and narrow at the extremity,
Ane IES Ob y forming an oblique margin against which
the dactylus folds; the dactyli of the last three pairs are contained about
three times in their respective propodi.

Abdomen two and a half times as long as carapace, smooth, non-car-
inate, third segment very slightly produced backward in the middle;
sixth segment one and two thirds as long as wide; seventh one and a half
times as long as sixth, and with six spinules on each side.

Dimensions.—Adult female (San Francisco market), length 110 mm.,
length of carapace and rostrum 50 mm., of rostrum 26.5 mm.

Distribution.—¥rom Sitka, Alaska, to San Francisco, California.

DECAPODS 49

Specimens have been examined from:

Sitka, Alaska, 10 fathoms (Harriman Expedition).

Ward Cove, Revillagigedo Island, Alaska (Dr. T. H. Streets, U. S. N.).
Off Cape Beale, Vancouver Island, 24 fathoms (A/batross, station 2881).
Otter Bay, Pender Island, British Columbia (A/da/ross).

Strait of Fuca, 40-97 fathoms (Aédasross, stations 3443, 3462, 3464,

3495).
Admiralty Inlet, Puget Sound, 40 fathoms (4/éatross, station 2865).
Puget Sound (T. Kincaid).
Port Angeles and Port Townsend, Washington (Adbatross).
Port Orchard, Washington (O. B. Johnson).
San Francisco, California (D. S. Jordan).
San Francisco market (R. E. C. Stearns).

PANDALUS STENOLEPIS Rathbun.
Plate 11, fig. 4.
Pandalus stenolepis RATHBUN, Proc. U.S. Nat. Mus., XXIV, gol, 1902.

Differs from 2. dane:

(r) In the stouter eyes, the transverse diameter being as great as the
axial diameter.

(2) Inthe more slenderantennal scale. This |
scale has a concave outer margin, and the distal —
half of the blade is very narrow and filiform, ,_ Eee en nes
narrower than the adjacent thickened portion. Acicle (X 2). Station 3464.

(3) The first pair of feet is longer, reaching almost to the extremity of
the maxillipeds.

(4) The third, fourth, and fifth pairs of feet in the male, while shorter
than in the female, as in P. dave, do not differ in shape from those of
the female, as in the older species.

(5) In the tip of the rostrum, usually 2-spined, though occasional
specimens have 3 spines.

Dimensions. —Adult female (station 3464), length 80 mm., length of
carapace and rostrum 37.2 mm., of rostrum 21 mm.

Distribution.—From the Aleutian Islands to Oregon, 27 to 125 fath-
oms. ‘Taken at the following stations:

Pumicestone Bay, Unalaska, 59 fathoms, station 3319.

Tliuliuk Harbor, Unalaska, 85 fathoms, station 3311.

Off Akutan Island, Alaska, 36 fathoms, station 3546.

Unimak Pass, Alaska, 34 fathoms, station 3220.

Northwest of Unimak Island, Alaska, 43 fathoms, station 3262.
Southwest of Sannak Islands, Alaska, 41 fathoms, station 3213.

Off Cape Beale, Vancouver Island, 34 fathoms, station 2879.

Off Cape Flattery, Washington, 27-40 fathoms, stations 2873, 2874.

50 RATHBUN

Strait of Fuca, 37-125 fathoms, stations 3443, 3445, 3451, 3452, 3458,

3459, 3461, 3462, 3464 (type locality), 3593.
Off Alsea River, Oregon, 42 fathoms, station 3085.

Off Heceta Bank, Oregon, 68 fathoms, station 3078.
Heceta Bank, Oregon, 42-50 fathoms, stations 2886, 2887, 2889.
Granite Cove, Port Althorp, Alaska (W. H. Dall).

PANDALUS GURNEYI Stimpson.
Plate 11, fig. 6.

Pandalus gurneyi STIMPSON, Ann. Lyc. Nat. Hist. N. Y., x, 128, 1871. Mon-
terey, Calif.

The species which I take to be P. gurneyi is very near P. dane.
Our adult specimens are smaller than P. dane , rostrum longer, being
from one and a half to one and two thirds longer than the carapace.
Median dorsal spines 8 or g, rather distant, ventral spines 9 or 10. An-
tennular peduncle only two fifths as long as acicle; the flagella are sub-
equal and one and a half times as long as carapace, the thickened
portion of the outer one being a little over half the entire length; the
basal scale is rounded, not angled.

Antennal scale as long as carapace, flagellum exceeding length of
body.

Maxillipeds reaching three fourths the length of antennal scale, first
pair of feet reaching nearly as far. Right leg of second pair extends to
tip of acicle, carpal segments 17; left leg one third longer than right,
carpal segments about 45.

As the last three pairs of feet are about the same length as in P. dane,
it follows that in the female the third pair does not reach as far beyond
the rostrum as in that species, or fails to reach the end of the
rostrum.

In the male the last three pairs of legs are shorter than in the female,
> — and the propodus of the third pair is slightly re-
ie curved and narrowed at the extremity, forming a
margin against which the dactylus closes; this is, if
I mistake not, the condition which Stimpson de-
scribes as ‘ subcheliform.’

Sixth segment of abdomen one and a half times
as long as wide; seventh less than one and a half
times as long as sixth, 5 spinules on each side.
Otherwise as in P. dane.

Dimensions.— Ovigerous female (station 2961), length 77.5 mm.,
length of carapace and rostrum 38.5 mm., of rostrum 25 mm.

Fic. 15. Pandalus
gurneyt. Third leg
of 6 (X 2). Station
3130.

DECAPODS BT

Distribution.—Southern California, 9 to 55 fathoms (rare). Taken at
the following stations by the Albatross :

Monterey Bay, g fathoms, station 3130.

Off Santa Barbara, 21 fathoms, station 2961.

Off San Miguel Island, 55 fathoms, station 2959.

Off Santa Cruz Island, 30 fathoms, stations 2944, 2945.
Off Santa Rosa Island, 52 fathoms, station 2956.

Genus Pandalopsis Bate.

KEY TO THE SPECIES OF PANDALOPSIS.

A. Rostrum with spines on distal half of upper margin . . . dispar.
A’. Rostrum without spines on distal half of upper margin.

B. Dorsal spines not continued to middle of carapace. . . ampla.

B’, Dorsal spines continued to middle of carapace.

C. Palm of chela more than one and a half times as long as fingers,

Rostrum less than twice as long as carapace proper . aleutica,

C’, Palm of chela less than one and a half times as long as

fingers. Rostrum more than twice as long as carapace

DIODE eet sarong Pe. Payne Fen, Maha et wets SOM CEPOSLI TS.

PANDALOPSIS AMPLA Bate.

Pandalopsis amplus BATE, Challenger Rept., XXIV, 671, pl. CXV, fig. 3, 1888.
Pandalopsis ampla FAXON, Mem. Mus. Comp. Zool., XVIII, 155, 1895.

Surface remotely punctate. Rostrum one and a third to one and nine
tenths times as long as the rest of the carapace (in small specimens it may
be two and a half times the carapace), slightly ascending, slightly arched
above the eyes. Median crest occupying the anterior two thirds of the
carapace, armed with 7 to 13 movable spines, of which 3 to 5 are on the
carapace and in front of the middle, the anterior spine considerably
behind the middle of the rostrum. There is also a subterminal fixed
spine; and occasionally two subterminal spines—both above, or one
above and one below. Greater part of the rostrum unarmed above.
Lower margin armed with 13 to 16 slender spines, larger toward the
base of the rostrum. Suborbital spine slender, prominent. Pterygos-
tomian spine short and slender.

Eyes very stout, ocellus rudimentary. The peduncle of the anten-
nula extends to the middle or nearly to the middle of the antennal scale ;
the second segment is twice as long as the third; inner flagellum at least
half as long as the body; outer flagellum one fourth longer than inner,
thicker at the base, but gradually tapering. The peduncle of the an-
tenna does not reach quite to the end of the second segment of the
antennular peduncle; flagellum one and a third times the length of the

52 RATHBUN

body. Scale four fifths to eight ninths as long as carapace; blade
broadly rounded at the tip and exceeded by the spine.

The maxilliped falls considerably short of the tip of antennal scale,
but its penultimate segment reaches to or beyond the end of the anten-
nular peduncle.

The ischium of the first pair of feet is dilated in a thin, broad laminar
inferior projection, which is anteriorly lobiform; the terminal segment
overlaps a little that of the maxilliped. The feet of the second pair are
equal; the carpus has 20 to 24 segments; the right and left carpi may
or may not have an equal number of segments; the chela is as long as
the five adjacent segments of the carpus. The third, fourth, and fifth
pairs of feet are slender and similar, and armed with slender spines;
while they diminish regularly in length from the third to the fifth, their
propodi increase in length proportionally; the dactyli are short and con-
tained from five to six times in their respective propodi. The third pair
overreaches the acicle by the length of the dactylus and at least half the
propodus.

The third segment of the abdomen is moderately produced backward
in the middle, forming a lobe in the posterior margin; the sixth segment
is nearly two and a half times as long as wide, and about two thirds as
long as the seventh; this has 5 to 8 spinules on each side, of which the
anterior is just in front of the middle.

Dimensions.— Length 164 mm., length of carapace and rostrum 85.7
mm., of rostrum 57.5 mm.

Distribution.—¥rom Washington to Mexico; off Monte Video; 309-
984 fathoms:

Off Monte Video, 600 fathoms (Challenger, type locality).

Off Acapulco, 660 fathoms, station 3418, and near Tres Marias Islands,
Mexico, 676 fathoms, station 3424 (A/batross, Faxon).

Off Cortez Bank, California, 984 and 776 fathoms, stations 2919 (one
young) and 3627.

Off San Diego, 822 and 623 fathoms, stations 2983 (abundant) and
2929.

Off Sea Lion Rock, Washington, 877 and 859 fathoms, stations 3074
and 3075 (abundant),

PANDALOPSIS ALEUTICA Rathbun.
Plater, wise en:

Pandalopsis aleutica RATHBUN, Proc. U. S. Nat. Mus., XXIV, 901, 1902.

Surface of carapace covered with a short and soft pubescence. Bran-
chial region traversed by a curved longitudinal ridge. Rostrum one and

DECAPODS 53

one seventh to one and a half times the length of the rest of the cara-
pace, ascending. Anterior two thirds of carapace furnished with a
median crest. Median spines 8 to 13, four to six of which are behind
the orbits, the insertion of the posterior spine being at the middle of
the carapace; the anterior spine is behind the middle of the rostrum;
this is exclusive of a subterminal spine. Inferior spines 8 to 12.

The peduncle of the antennula extends a little past the middle of the
antennal scale; the second segment one and a half times as long as
third; outer flagellum two thirds as long as body, inner flagellum shorter.

Peduncle of antenna reaching to end of second antennular segment ;
scale three fourths to four fifths as long as carapace, broader behind than
in P. ampla, flagellum one and a half times the length of the body.

The outer maxillipeds extend either to the tip or nearly to the tip of
antennal scale; the first pair of pereiopods overreach the penultimate
segment of the maxilliped; second pair subequal, extending beyond
acicle, carpal segments 18 to 21, chela equal in length to
five and a half or six of the adjoining segments of the = —

. Fic. 16. Pandalopsis
carpus; palm more than one and a half times as long as aleutica. Chela of 9
fingers. There is very little difference in the length of aa
the third, fourth, and fifth pairs of feet; the third pair reaches beyond
the acicle by the length of the dactylus and half the propodus; the dactyli
are contained about five and a half times in their propodi.

The pleon is much like that of P. amp/a; the pleura of the second seg-
ment are narrower than in P. amp/a, the sixth segment is shorter, only
twice as long as wide.

For the rest, as in P. amfpla,

Dimensions.—Adult female, length 132 mm., length of carapace and
rostrum 61.5 mm., of rostrum 33 mm.

Distribution.—Over one hundred specimens were taken by the 4/Ja-
tross at station 3480, off Seguam, Aleutian Islands, 283 fathoms (type
locality). North of Rat Islands, Aleutians, 270 fathoms, station 3785.

Affinities.— Besides its resemblance to P. ampla, this species is also
similar to P. damelligera (Brandt), from Kamchatka, but differs in the
fewer dorsal spines, in the longer feet of the first pair, and in the nar-
rower pleura of the second segment of the pleon.

PANDALOPSIS LONGIROSTRIS Rathbun.

Pandalopsis longirostris RATHBUN, Proc. U. S. Nat. Mus., XXIV, 902, 1902.
Male,—Surface smooth. Rostrum a little more than twice as long as
the carapace proper, strongly ascending, and continued backward in a

54 RATHBUN

carina behind the middle of the carapace, armed with 9 movable spines,
four of which are on the carapace (the hinder one inserted very slightly
behind the middle) and
five on the base of the
rostrum; 1 subterminal
immovable spine ; lower
margin armed with 11

Z c= = -~_ immovable spines.
ee) _——_ Antennular peduncle
5 f extending to middle of

Fic. 17. Pandalopsts longirostris. Station 3316. a. Side of
carapace (natural size). 6. Acicle (x 2). c. Chela (x 34). scale. Scale as long as
¢ cee carapace. Antennal pe-
duncle reaching to middle of second antennular segment. Maxillipeds
reaching almost to end of scale, very stout and hairy. The first pair
of pereiopods overlaps the basal fourth of the last joint of the maxilli-
peds; second pair extending beyond acicle by length of chela, carpus of
21 joints, chela equal to the seven adjoining segments, fingers almost
as long as palm; third pereiopods extending beyond the acicle by the
length of the dactylus and two thirds of the propodus.

Sixth segment of abdomen two and a half times as long as wide;
telson broader than in P. aleutica.

Dimensions. —Male, length 112 mm., carapace and rostrum 59 mm.,
rostrum 41 mm.

Distribution.—Off liuliuk Harbor, Unalaska, 309 fathoms, station
3316, Albatross, 2 males (one without rostrum).

PANDALOPSIS DISPAR Rathbun.
Plate, fig. 2.

Pandalopsis dispar RATHBUN, Proc. U. S. Nat. Mus., XXIV, 902, Ig02.

Surface very finely and closely punctate. Rostrum two to two and a
half times the length of the rest of the carapace, arched over the eyes,
the remainder slightly ascending. Median crest occupying two thirds
the length of the carapace, posterior spine at the anterior third, spines 16
to 21, three or four of which are on the carapace, spines closely placed
on the arch, distant on the remainder of the rostrum. Inferior spines 9
to 15, extremity bifid or sometimes trifid. Antennal spine long and
slender; pterygostomian spine minute.

Eyes very large, with a very small but distinct ocellus outside the
corneal area.

Antennal peduncle reaching two fifths the length of antennal scale;

DECAPODS 55

third segment but little shorter than second; outer basal scale a narrow
lobe; outer flagellum one third longer than the body; inner flagellum
one half as long as the outer. Antennal scale a little shorter than the
carapace, tapering distally; extremity of blade obliquely rounded and
exceeding the spine; peduncle reaching end of second segment of anten-
nular peduncle; flagellum one and a half times length of body.

The outer maxillipeds reach to the distal fifth or sixth of the antennal
scale. The first pair of feet overlap the proximal third of the terminal
joint of the maxilliped. Second pair subequal, overreaching the acicle a
little; carpus composed of from 26 to 33 segments. Third to fifth pairs
of feet nearly equal, the third reaching beyond the acicle by length of
dactylus and one half or more of propodus, their propodi increasing in
length from the third to the fifth pair, while the dactyli increase in reverse
order; so that while the dactylus of the third pair may be contained three
and a half or four times in the corresponding propodus, the dactylus of
the fifth pair is contained seven or eight times in its propodus; the spines
of the meral and carpal segments are replaced by bristles on the propodi;
dactyli subentire, although there may be a few small very appressed
spinules at the base of the concave surface.

Abdomen two and four fifths to three times as long as the carapace
(exclusive of rostrum); third segment slightly compressed, its posterior
margin produced in a short subacute lobe. Sixth segment nearly three
times as long as wide and three fourths as long as seventh; seventh with
five to seven spinules on each side.

Outer branch of tail-fan about as long as the telson; inner branch
much shorter.

Dimensions.— Length of female 181 mm., length of carapace and
rostrum roo.5 mm., length of rostrum 73.5 mm. Length of female (sta-
tion 3675) 211.5mm. |

Distinctive characters— The spines distributed along the whole upper
margin of the rostrum, the unequal dactyli of the third to fifth pairs of
feet, and the inequality of the branches of the swimming-fan easily dis-
tinguish this species from the preceding.

Distribution.—From Bering Sea to Washington, 53 to 351 fathoms,
at the following stations of the Albatross :

Bering Sea, west of Pribilof Islands, 184 fathoms, station 3489.
Chernofski Harbor, Unalaska, 109 fathoms, station 3324 (type locality).
North of Unalaska, 351, 350 fathoms, stations 3330, 3331.

Between Unga and Nagai Islands, Shumagins, 110 fathoms, station 2848.
Off the Trinity Islands, 159 fathoms, station 2853.

Clarence Strait, Alaska, 322 fathoms, station 3077.

Queen Charlotte Sound, British Columbia, 238 fathoms, station 2862.

56 RATHBUN

Off Strait of Fuca, 142 fathoms, station 3457.
Strait of Fuca, 53-136 fathoms, stations 3446-3449, 3451, 3452, 3456,

3458-3460, 3596, 3597-
Puget Sound, 82-135 fathoms, stations 3067, 3068.

Off Tahwhit Head, Washington, 178 fathoms, station 3076.
Stations 3675, 3676 (locality not given), 110 and 122 fathoms,

Family 27PPOL VTIDZ.
Genus Hippolyte Leach.

HIPPOLYTE CALIFORNIENSIS Holmes.

Distribution.—From Sitka, Alaska, to San Diego, California. Speci-
mens are in the National Museum from Sitka, 10 fathoms (Harriman
Expedition, one specimen); Barclay Sound, British Columbia (A/Zatross,
one specimen); Puget Sound (T. Kincaid, several specimens); Bodega
Bay, California (determined by S. J. Holmes); San Diego, California
(Albatross).

The Sitka specimen, about 24 mm. long, has on the rostrum 3 teeth
above on basal half, below 3 near the middle and 1 near the tip. The
example from Barclay Sound has 2 teeth above on basal half, and 1 near
the tip, below 2 near the middle and 1 near the tip. In a lot of eleven
specimens from Puget Sound, the dorsal teeth are usually 2 (in one
case 3) on basal half of rostrum, and 1 or none near tip. In the few
individuals from San Diego the rostral teeth are typical.

Genus Hippolysmata Stimpson.

HIPPOLYSMATA CALIFORNICA Stimpson.

Distribution.—¥rom Santa Barbara to San Diego, California.
Genus Spirontocaris Bate.
(Includes Heptacarpus Holmes.)

KEY TO THE SPECIES OF SPIRONTOCARIS.

A. One or more supraorbital spines present.

B. Rostrum subcircular. Median spines of carapace compound or
formed by short transverse rows of small ae 3 OF 4 supra-
orbital spines. SEG AA een - . prionota,

B’. Rostrum not subcircular, usually elongate. Median spines of
carapace simple, or not formed by transverse rows of small
spines. Not more than 2 supraorbital spines.

C. First to third abdominal segments laterally acute or spinous.
D. First and second abdominal segments armed with slender
spines. Rostrum narrow ..... . greniandica.

DECAPODS BY

D’. First and second abdominal segments laterally acute. Ros-
tramp dee pris, xe . . lamellicornis.
C’. First to third abdominal segments laterally rounded, not acute.
D. Two supraorbital spines.
E. Rostrum longer than the remainder of the carapace and
with a long slender tip . . . . bispinosa.
E’. Rostrum shorter than the remainder of the carapace.
F. Rostrum not reaching end of antennular peduncle
truncata,
F’. Rostrum reaching beyond end of antennular peduncle.
G. Antennular scale not reaching middle of second seg-
ment of peduncle.
H. Dorsal spines all in front of middle of carapace

Sica.
H’. Dorsal spines continued posterior to middle of
carapace ..% : . snyderi.

G’, Antennular scale reaching beyond middle of second
segment of peduncle.
H. Dorsal spines not reaching to posterior third of

carapace.

J. Upper limb of rostrum gradually diminishing in
width from the orbit to the tip . . phippsit.

J’. Upper limb of rostrum arcuate, widest in the
middle.

K. Rostrum extending only to end of antennular
peduncle, deep, usually bifid at tip ochotensis.
K’. Rostrum extending nearly to end of antennal
scale, less deep, tip acute . . . . dallt.
H’. Dorsal spines continued to posterior third of cara-
pace.
J. Midrib of rostrum terminating in a spine which
projects well beyond lower limb of rostrum.

K. Eyes large, pyriform. Spine of antennal scale

extending beyond lamellar portion
hivjeborgit,
K’. Eyes of moderate size, subcylindrical. Spine
of antennal scale not reaching beyond lam-
ellar portion <.. a. « . 6. murdochi.
J’. Midrib of rostrum terminating in a spine which
does not project beyond lower limb of rostrum.
K. Upper margin of carapace and rostrum in 9
convex. Third abdominal segment in ¢@ in
profile not projecting behind fourth segment
arcuata.
K’. Upper margin of carapace and of rostrum in 9
separated by a depression. Third abdominal
segment in ¢ in profile projecting in a horn-
like process behind fourth segment . spina.

D’. One supraorbital spine.

58 RATHBUN

E. Rostrum less than twice as long asthe eye washingtoniana.
E’. Rostrum twice or more than twice as long as the eye.
F. Antennular scale shorter than first segment of peduncle
vicina.
F’. Antennular scale longer than first segment of peduncle.
G. Fifth segment of abdomen not armed with a spine on
either side. - affinis.
G’. Fifth segment of abdomen ‘armed with a spine on
either side.
H. Rostrum longer than the rest of the carapace
unalaskensis.
H’. Rostrum no longer than the rest of the carapace
polaris.
A‘. No supraorbital spine or spines.
B. Rostrum about as long as or longer than the rest of the carapace.
C. Third, fourth, and fifth segments of abdomen carinated, the carina
on each segment terminating in a sharp spine . . darbata.
C’. Third, fourth, and fifth segments not all carinated nor ending in
a sharp spine.
D. Terminal half (at least) of rostrum devoid of spines above.
E. Third abdominal segment with an angle or hump, in profile,
toward the posterior end.
F. More than 3 dorsal teeth, one or more in front of eyes.
G. Sixth abdominal segment less than twice as long as
wide. Rostrum deep, one fourth as deep as long
carinala,
G’. Sixth abdominal segment more than twice as long as
wide. Rostrum more slender.
H. Maxilliped with epipod. Scale at base of antennula
extending beyond first segment. . . . flexa.
H’. Maxilliped without epipod. Scale at base of an-
tennula not reaching poate or only slightly

beyond, first segment . . sp seraciis:

F’. Three dorsal teeth, none in front of eyes oF Nios REZELCHS.

E’. Third abdominal segment smoothly rounded, without angle
or hump.

F. One or more superior rostral spines in front of eyes.
G. No pterygostomian spine . . . . « « « stylus.
Gi A pterygostomian spine oun ct m Qmabiles:
F’. No superior rostral spine in front of eye sims Saori.
D’. Terminal half of rostrum with spines (in part at least).
E. Terminal third of rostrum unarmed above and below
biunguis.
E’. Terminal third of rostrum not entirely unarmed.
F. Sixth abdominal segment longer than seventh . decora.
F’, Sixth abdominal segment shorter than seventh.
G. Maxilliped exceeding acicle.
H. Rostrum straight above. Epipods on maxillipeds
and first two thoracic feet. . . . paludicola.

DECAPODS 59

H’. Rostrum concave above. ey aa on maxillipeds

and first footonly. . . « . > + MOSsert,
G’. Maxilliped not exceeding acicle.

H. Third abdominal segment with a lobe (in profile),
which is in the form of a hook in the @, of a
hump in the @. 8 to 12 dorsal spines

gaimardit belcheri.

H’, Third segment without a lobe (in profile) in the @.

J. Rostrum with more than 8 spines above . Jayi.
J’. Rostrum with 8 or fewer than 8 spines above.
K. Fourth abdominal segment with a spine on
Gleherside: Yr 2.5 » + 4 Suckleyt.
K’. Fourth abdominal segment without spine.
L. Sixth segment barely twice as long as high.
M. Maxillipeds reaching just beyond middle
of acicle® .- . . . camtschatica.
M’. Maxillipeds reaching nearly to end of
ACICLE a ie . . Rincatdi.
L’. Sixth segment more ‘than twice as long as
Hight <5" “qieet - . vownsendi.
B’. Rostrum shorter than the rest of the carapace.

C. Upper and lower limbs of rostrum deep and with convex margins.
D. Upper limb of rostrum deepest above the eye . . macilenta.
D’. Upper limb narrow above the eye . . . macrophthalma.

C’. Upper and lower limbs of rostrum not both convex.

D. Rostrum elongate, reaching beyond the middle of the an-
tennal scale.
FE. Upper margin of rostrum concave.

F. Fingers of first pair of chelipeds less than one third as
long as palm. No pterygostomian spine. No spine
on fourth segment of abdomen. . . brachydactyla,

F’. Fingers of first pair of chelipeds about half as long as
palm. <A pterygostomian spine or spinule. A spine
or spinule on fourth segment of abdomen mazxillipes.

E’. Upper margin of rostrum straight.
F. Antennal peduncle reaching end of antennular peduncle

picta.
F’, Antennal peduncle reaching end of second segment of an-
tennular peduncle, guetta = SPLCHETISES.

D’. Rostrum short, not reaching beyond the middle of the an-
tennal scale,
E. Rostrum not reaching as far as the cornea . . . faylori.
E’. Rostrum reaching as far as or beyond the cornea.
F. Rostrum not reaching the second segment of antennular

peduncle.
G. Rostrum with superior margin strongly convex over
the eyes, extremity straight. . . . . . avina.

G’. Rostrum with superior margin not strongly convex
over the eyes, but nearly straight.

60 RATHBUN

H. Maxilliped with exopod :). .. . « pusiola,
H’. Maxilliped without exopod.
J. Antennal scale about same length as, or shorter

than, telson .. . « brevirostris.

J’. Antennal scale longer than telson . . palpator.

F’. Rostrum reaching second segment of antennular pe-
duncle.

G. Rostrum straight . . . ee «| Meramani:

G’. Rostrum arched over the eye.

H. Fourth segment of abdomen with lateral spinule.
Maxillipeds reaching, or nearly reaching, end of
acicle : : cristata.

H’. Fourth segment of abdomen unarmed. Maxillipeds
reaching only to distal third of acicle . stoneyi.

SPECIES OF SPIRONTOCARIS, ARRANGED ACCORDING TO THEIR
EXOPODS AND EPIPODS.

A. One or more supraorbital spines present.
B. Third maxilliped with exopod and epipod.
C. First thoracic foot with epipod. . . , 2) ee See.
C’. First and second thoracic feet with epipods : bispinosa, snyderi.

C’, First, second, and third thoracic feet with epipods
prionota, lamellicornis, spina, arcuata, murdocht,
truncata?, liljeborgit, phippsit, ochotensis, dal.
B’. Third maxilliped without exopod but with epipod.
C. First thoracic foot with epipod . . = » icing:
C’. First and second thoracic feet with epipods polaris, unalaskensis.
C”. First, second, and third thoracic feet with epipods
greniandica, washingtoniana.
B’. Exopods'and epipods unknown 9. 4s. <<. « « «Gents.
A’. No supraorbital spine or spines.
B. Third maxilliped with exopod and epipod.
C. Thoracic feet without epipods darbata, biunguis, macrophthalma.
C’. First thoracic foot with epipod
Jabricii (usually), suckleyt (usually).
C”. First and second thoracic feet with epipods
Jabricii (sometimes), gaimardi bel-
chert, townsendi, suckleyi (sometimes).
C’’. First, second, and third thoracic feet with epipods
pusiola, herdmani, stoneyi, avina, macilenta.
B’. Third maxilliped without exopod but with epipod.

C. Thoracic feet without epipods
decora, tridens, stylus, maxilhipes,

brachydactyla, camtschatica, Rincaidi.
C’. First thoracic foot with epipod . . . . moseri, sitchensis.
C”. First and second thoracic feet with epipods
fiexa, picta, paludicola.
C’, First, second, and third thoracic feet with epipods
carinata, palpator, brevirostris, taylori, cristata.

DECAPODS 6I

B”’, Third maxilliped without acca or ened Thoracic feet
without epipods . . % 2 oi RCIUS.
B’’, Exopods and epipods unknown Ogre. Tan a: OMQOIUS. faye,

SPIRONTOCARIS PRIONOTA (Stimpson).

Hippolyte prionota STIMPSON, Proc. Acad. Nat. Sci. Phila., 153, 1864.—
KINGSLEY, Bull. Essex Inst., XIV, 127, pl. II, fig. 9, 1883.—SHARP,
Proc. Acad. Nat. Sci. Phila., 117,,.1993:

Spirontocaris prionota WALKER, Trans. Liverpool Biol. Soc., XII, 277, 1898.

Spirontocaris prionata HOLMES, Occas. Papers Calif. Acad. Sci., VII, 206,
1900.

To Kingsley’s figure one must add 2 or 3 supraorbital spines arranged
in a longitudinal series. While 3 seems to be the normal number, the
anterior one is often reduced and sometimes wanting.

Distribution.—From Bering Sea to Monterey, California:

Dutch Harbor, Unalaska, and Kadiak (Harriman Expedition, W. R. Coe).

Collected by W. H. Dall at Cape Etolin, Nunivak Island, 8 fathoms;
Captains Harbor, Unalaska, 9 and 80 fathoms; Coal Harbor, Unga,
8-g fathoms; Belkofski Bay, 15-25 fathoms; Chiniak Bay, Kadiak ;
Chugachik Bay, Cook Inlet, 20-60 fathoms.

Dredged by the A/atross in Bristol Bay, Alaska, 744 and 10% fathoms,
stations 3232 and 3233; off North Head, Akutan Island, 72 fathoms,
station 2842; Unimak Pass, 34 fathoms, station 3220; North of Bird
Island, Shumagins, 21 fathoms, station 2850; Admiralty Inlet, Puget
Sound, 40 fathoms, station 2865.

Puget Sound (T. Kincaid).

Pacific Grove, California (J. O. Snyder).

Bering Island, 2-12 fathoms, some in stomach of Gadus macrocephalus
(N. Grebnitzky).

Puget Sound (Stimpson); Marmot Isles, Alaska, 45 fathoms (Sharp) ;
Monterey, California (Holmes).

SPIRONTOCARIS GRAGNLANDICA (J. C. Fabricius).

Astacus Grenlandicus J. C. FABRICIUS, Syst. Entom., 416, 1775; Entom.
syst. auct. et emend., II, 484, 1793 (gre@nlandicus).

Cancer aculeatus O. FABRICIUS, Fauna Greenlandica, 239, 1780.

Alpheus Aculeatus SABINE, in Supplement to Appendix of Parry’s [First]
Voyage, p. CCXXXVII, pl. II, figs. g and Io, 1824.

Hippolyte aculeata J. C. Ross, in John Ross, Appendix to Narrative of a
Second Voyage in Search of a North-West Passage, IJ, p. ]xxxili, 1835.

Hippolite armata OWEN, in Zoology of Capt. Beechey’s Voyage [of the
Blossom], Crustacea, p. 88, pl. XXvII, fig 2, 1839 (@).

fippolite cornuta OWEN, op. cit., p. 89, pl. XXVIII, fig. 2,°1839 (2).

Hippolyte grenlandica MiERS, Ann. Mag. Nat. Hist. (4), xx, 62° (22),
1877.—SMITH, Trans. Conn. Acad. Arts Sci., Vv, 85, pl. X, fig. 251879.

Spirontocaris grenlandica WALKER, Trans. Liv erpool Broly. Soc., x11, 270;
1898.—HOLMES, Occas. Papers Calif. Acad. Sci., VII, 236, 1900.

The number of spines on the abdomen is the same in the two sexes,

62 RATHBUN

but there is some individual variation. The usual number of spines is
one on the second and sixth segments and two on each of the other
segments.

Distribution.— Arctic coast of America; Bering Sea to Puget Sound;
Kamchatka; Okhotsk Sea; Atlantic coast of America from Greenland
to Narragansett Bay, Rhode Island, 1-72 fathoms:

Berg Bay, Glacier Bay, Alaska (Harriman Expedition).

Dredged by the A/datross from off Cape Menshikof, Bering Sea, to the
Aleutian Islands and Portlock Bank, Alaska, in 16 to 72 fathoms; at
the unusual depth of 283 fathoms, off Seguam, Aleutians, a lot of
large specimens, the largest 9 measuring 77 mm., the largest ¢ 67 mm.;
Strait of Fuca and Puget Sound, 40-48 fathoms; off Kamchatka,
12 and 42 fathoms, stations 3780 and 3781; Avacha Bay, Kamchatka,
16 fathoms, station 3642; Rakovaya Bay, Avacha Bay; off Robben
Island, Okhotsk Sea, 10 fathoms, station 3645.

Various localities from Kadiak westward along the Aleutian Islands, in
3-25 fathoms (W. H. Dall).

Puget Sound (T. Kincaid).

Avacha Bay and Petropavlovsk, Kamchatka (L. Stejneger).

Bering Island, among Laminaria, and Solovarennaja Bay, Petropavlovsk,
Kamchatka, 10-12 fathoms (N. Grebnitzky).

Arctic Ocean (Stimpson).

SPIRONTOCARIS LAMELLICORNIS (Dana).

Hippolyte lamellicornis DANA, Proc. Acad. Nat. Sci. Phila., 1852, 24; Crust.
U. S. Expl. Exped., 1, 567, 1852, pl. XxxvI, fig. 6, 1855.—STIMPSON,
Jour. Boston Soc. Nat. Hist., v1, 498, 1857.—KINGSLEY, Bull. Essex
Inst., .X, 62, 1878.

Spirontocaris lamellicornis WALKER, Trans. Liverpool Biol. Soc., XII, 277,
1898.—HOLMES, Occas. Papers Calif. Acad. Sci., VII, 208, 1g00.

This species, though resembling strongly S. spina, is at once separated
by the acuteness of the lateral margins of the first to third abdominal
segments, which in S. spina are broadly

rounded. The rostrum is about three

fourths as long as the rest of the cara-

pace, and does not exceed the antennal

scale, although it may exceed the blade

Fic. 18. Spirontocaris lamellicornis. Side of the scale. The spine of the scale ex-

of carapace of ¢ (x 18). Station 3046. tends beyond the blade to a distance
equaling or exceeding the distal width of the blade. The dactyli of the
last three pairs of feet are longer than in S. sina, those of the last pair
being half the length of their propodi; they are armed with spinules so
minute that the segment appears entire in comparison with S. spina.
Dimensions.— 2, length 63 mm., carapace 23 mm.

DECAPODS 63

Distribution.—This species occurs sparingly from Unalaska to Point
Arena, California; 9-77 fathoms.

It has been taken by W. H. Dall at Captains Harbor, Unalaska, g
fathoms; Port Etches, 12-18 fathoms; Sitka Harbor, 15 fathoms. By the
Albatross at Gulf of Georgia, British Columbia, 67 fathoms, station 2863;
Strait of Fuca, 53 fathoms, station 3460; Bellingham Bay, Washington,
11 fathoms, station 3612; off Destruction Island, Washington, 32 fathoms,
station 2869; off Grays Harbor, 48-58 fathoms, stations 2870, 3046—
3048; off Columbia River, 68 fathoms, station 2882; off Oregon, 38-77
fathoms, stations 3057-3059; off Point Arena, California, 51 fathoms,
station 3351.

SPIRONTOCARIS SPINA! (Sowerby).
Plate 111, fig. 5.

Cancer spinus SOWERBY, British Miscellany, 47, pl. XXIII, 1805 (deste Steb-
bing).

Alpheus spinus LEACH, Edinb. Encyc., vu, 431, 1814; Philadelphia reprint,
VII, 271.

Alpheus Spinus LEACH, Trans. Linn. Soc. London, XI, 347, 1815.

Hippolyte Sowerbai LEACH, Malac. Pod. Brit., pl. XXXIX, 1817.

Hippolite sowerbei J. C. Ross, in John Ross, Appendix to Narrative of a
Second Voyage in Search of a North-West Passage, I, p. Ixxxiii, pl. B,
fig, 2, 1835.

Hippolyte spinus WHITE, List Crust. Brit. Mus., 76, 1847.—BELL, Hist.
Brit. Crust., 284, 1853.—SMITH, Trans. Conn. Acad. Arts Sci., v, 68,
1879.

Hippolyte spina STIMPSON, Proc. Acad. Nat. Sci. Phila., XII, 34 (103), 1860;
Ann, Lyc. Nat Hist. N. Y., X,.126, 1371,

Spirontocaris spinus BATE, Challenger Report, XXIV, 596, pls. CVI and Cvq,
1888 (part).—RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. 111, 556, 1899 (part).

There is considerable variation, in this species, in the height of the
carapace in proportion to its length; in the eyes, which may be widely
pyriform or smaller and subcylindrical; in the
length of the outer maxillipeds (in none of the aks
Pacific specimens do the maxillipeds reach the
end of the acicle, while in many Atlantic ones
they do); in the carination and the length of the | Fic.19. Spirontocaris spina.

; . F Side of carapace of ? (Xx 12).
spine of the third abdominal segment. Station 2842. .

1In regard to spina vs. spinus, Stimpson says (Ann. Lyc. Nat. Hist. N. Y.,
x, 126, 1871): ‘‘ Sowerby, by the name he gave to this species, doubtless had refer-
ence to a spine, or the backbone: in Latin sfiza, not sfinus. Spinus is not an adjec-
tive, and means only the sloe-tree, which could scarcely have been intended.”” While
spinus has another signification, from orfvoc, the name of a small bird, yet it was
without doubt used by Sowerby to call attention to the spine or spines of the animal.

64 RATHBUN

Ross (loc. cit.) probably represents the % of S. sina, but the shape of
the rostrum is inaccurate; both upper and lower lamine should have
more convex margins.

Distribution.—Circumpolar. Arctic Alaska, Bering Strait, Bering Sea,
Aleutian Islands, and Alaska Peninsula to Lituya Bay, 5-91 fathoms;
Atlantic Ocean southward to Massachusetts Bay, 5-90 fathoms;
northern Europe:

Taken by the Albatross at the following stations:

Off the Pribilof Islands, 41-62 fathoms, stations 3439, 3442, 3482, 3483,

3485, 3505, 3540, 3544, 3554, 3560, 3561.

Off Bristol Bay, 30 fathoms, station 3302.

Off Cape Strogonof, 30 fathoms, station 3293.

Off Kudobin Islands, 36 fathoms, station 3280.

Off Akutan Island, 56-91 fathoms, stations 2841, 2842, 3548.
Pumicestone Bay, Unalaska, 35 fathoms, station 3322.

Northwest of Unimak Island, 81 fathoms, station 3257.

South of Unimak Island, 42 fathoms, station 3217.

Between Bird and Nagai Islands, Shumagins, 35 fathoms, station 2851.

Collected by Dr. Dall at:

Bering Strait, 12 fathoms.

Bay of Islands, Adak, 9-16 fathoms.

Nazan Bay, Atka, 10-16 fathoms.

Captains Harbor, Unalaska, 9-80 fathoms.

Port Levashef, Unalaska.

Lituya Bay, 6-g fathoms.

Off Point Hope, Arctic Alaska, 25 fathoms (U.S. R. S. Corwin).
Lorenz Bay and Plover Bay (Richters).

SPIRONTOCARIS ARCUATA Rathbun.
Platevinm, sigsea.

Spirontocaris spinus RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. III, 556, 1899 (part).
Spirontocaris arcuata RATHBUN, Proc. U. S. Nat. Mus., xxiv, 893, 1902.

On the Pacific coast, associated in part with S. spina, is a very closely
allied species, easily mistaken for S. sfzza, but apparently distinct. In the
peers adult g , the upper line of carapace and ros-
Sa SS trum in profile forms a single curve, without
\, the sinus shown in 5S. sfiza, the rostrum is on

} a higher level than in that species, the rostral

spines larger and more distinctly marked; the
ieee apie of eo Ge), “Sx posterior lobe of the third abdominal segment
ter shorter and broader; the sixth segment con-
siderably shorter, being less than one and a half times as long as wide,

DECAPODS 65

while in S. sdiva it is more than one and a half times as long as wide;
the maxillipeds reach to the tip of the antennal scale; the dactyli of the
third to fifth pairs of feet are shorter than in SS. sdiza, that of the last
pair being contained at least three times in the propodus, while in SS. spina
the dactylus of the last pair is contained only two and a half times in its
propodus.

The males of this species and of .S. sAina are more difficult of deter-
mination. ‘The rostrum has the midrib more strongly curved upward, the
upper limb deeper and with more convex superior outline than in SS. spina.
The same differences in the abdomen exist as in the females of the two
species, but in a lesser degree.

Dimensions.— 2, length 53.8 mm., length of carapace and rostrum 20
mm., of rostrum 8 mm.

Distribution.—¥rom Pribilof Islands to Strait of Fuca; 3-350 fathoms.
A more southerly range than that of S. spina.

flarriman Expedition—Kadiak, 5 fathoms (W. R. Coe); Berg Bay,
Glacier Bay; Juneau, 20 fathoms.

Collected by the Albatross :

Off Pribilof Islands, 48-150 fathoms, stations 3484, 3486, 3500, 3561,
602,

of Cape Menshikof, 24 fathoms, station 3296.

Off Cape Strogonof, 26 fathoms, station 3291.

Northeast of Unimak Island, 24 fathoms, station 3266.

Northwest of Unimak Island, 43 fathoms, station 3262.

Unimak Pass, 34 fathoms, station 3220.

Off North Head, Akutan Island, 72 fathoms, station 2842.

Off south entrance to Akutan Pass, 45 fathoms, station 2843.

Off Aektok Island, 54 fathoms, station 2844.

Davidson Bank, 42 fathoms, station 2845.

North of Unalaska, 350 fathoms, station 3331.

Iliuliuk Harbor, 93 fathoms, station 3335.

Off Makushin Bay, Unalaska, 61 fathoms, station 3318.

Southwest of Sannak Islands, 41 fathoms, station 3213.

15 miles south of Sannak Islands, 44 fathoms, station 2846.

Off Shumagin Bank, 138 fathoms, station 3339.

Off Falmouth Harbor, Shumagins, 48 fathoms, station 2847.

Shumagins, 21-58 fathoms, stations 2850-2852.

Off Trinity Islands, 159 fathoms, station 2853.

Admiralty Inlet, Puget Sound, 40 fathoms, station 2865.

Washington Sound, Strait of Fuca, 48 fathoms, station 2864 (type
locality).

Strait of Fuca, 40-152 fathoms, stations 3443-3446, 3451, 3454, 3455,

3458, 3459, 3461, 3462, 3464, 3465, 3596.

Dr. W. H. Dall has collected the species at 16 localities along the
Aleutian Islands and eastward to Port Etches in 3-80 fathoms,

66 RATHBUN

SPIRONTOCARIS MURDOCHI Rathbun.

Plate 11, fig. 6.

Hippolyte spinus MURDOCH, Report Internat. Polar Exped. to Point Barrow,
Alaska, 140, 1885. Not Cancer sfpinus Sowerby.
Spirontocaris murdochi RATHBUN, Proc. U. S. Nat. Mus., XXIV, 893, 1902.

Very near S. “deborgi and S. spina.

Female stout. Rostrum three fourths as long as carapace, not reach-
ing tip of antennal scale. Carapace cristate, except on posterior fourth,

and furnished with 3 or 4 large teeth. Midrib

LA of rostrum nearly horizontal ; tip acute ; upper

limb with convex upper margin, armed with 20

or 22 small irregular teeth; lower limb, also with

oi aaa Re eee me- convex margin, deeper than the upper, and fur-

(X 18). Station 3650. nished with 1 to 4 small teeth near the extremity.

Two supraorbital spines, one antennal, and one pterygostomian spine,
all well developed.

Eyes of moderate size, slightly pyriform, the width of antennular
peduncle and half its scale. Antennular peduncle extending three fifths
the length of the antennal scale; second segment twice as long as third;
basal scale reaching nearly to end of second segment; thickened portion
of outer flagellum reaching about to tip of antennal scale. Antennal
peduncle reaching slightly beyond the first segment of the antennular
peduncle; scale narrow-ovate, about as long as the rostrum, its spine
reaching about as far forward as the blade.

Outer maxillipeds rather broad and reaching nearly to tip of acicle.
First pair of feet extending a little beyond antennal peduncle, the second
pair beyond acicle, fifth pair nearly to tip of maxilliped; dactyli of last
three pairs long and slender, that of last pair half as long as propodus.

Abdomen not carinate; third segment produced well backward;
fourth provided with a lateral spine; sixth three fourths as wide as long,
and about three fifths as long as the seventh, which has 3 pairs of lateral
spinules.

Male.—The male differs from the female in the same manner as in S.
“iljeborgii; it is smaller and slenderer than the female, rostrum and anten-
nules longer, dorsal teeth much reduced.

Dimensions.— 9, length 46.6 mm., length of carapace and rostrum 17
mm., of rostrum 7.5 mm.

Distribution.—Arctic coast of Alaska; Kamchatka; Okhotsk Sea.
Rare. Taken at the following localities:

Lat. 71° 02/ 00’ N., long. 157° 46/ oo” W., 19 fathoms, 1 4 (U.S. R. S.

Corwin).

DECAPODS 67

to miles west of Point Franklin, 1314 fathoms, 3 ¢ (Point Barrow Expe-
dition).

Off ae Sabine, 12 fathoms, 1 2 (W. H. Dall).

15 miles off Cape Krusenstern, 14 fathoms, 1 9 (W. H. Dall).

Rakovaya Bay, Avacha Bay, Kamchatka, 1 2 (Adbatross).

Off Robben Island, east coast of Sakhalin, Okhotsk Sea, 28 fathoms,

I ovigerous 9 , type (A/batross station 3650).

The species differs from S. “/jeborgii in its lower carapace, shorter (in
the ) and less ascending rostrum, smaller eyes, shorter spine of antennal
scale (in S. “deborgii the spine extends considerably beyond the blade),
longer dactyli of third to fifth pairs of feet, longer abdomen, and stouter
sixth segment. From S. spina it differs in the lower carapace and less
ascending rostrum, in the lesser advancement of the lower limb of the
rostrum, smaller eyes, shorter spine of scale, shorter and broader lobe of
third abdominal segment, which is not carinated, longer dactyli of third
to fifth pairs of feet (in S. sfzvza the dactylus of the fifth pair is only one
third length of propodus).

SPIRONTOCARIS TRUNCATA Rathbun.
Spirontocaris truncata RATHBUN, Proc. U. S. Nat. Mus., XXIV, 894, 1902.

Allied to S. spina. Median carina extending nearly to the posterior
margin of the carapace, convex, and armed with 4 spines. Rostrum sub-
triangular, widest at the extremity, about three fifths
as long as the carapace, reaching end of second
antennular segment, midrib curving upward as in
S. spina, Upper margin with 1 spine at its middle,
extremity with 7 spines, 2 above and 4 below the
midrib. Supraorbital spines 2, equal, large, one ete le ee
behind the other but scarcely above it, well sepa- (*°# Station 286.
rated. Anterior margin with 2 spines below the orbit as in S. spina.

Antennular acicle reaching beyond the anterior margin of the second
segment but not so far as the slender spine at the outer angle of that
margin; a small spine at the middle of the anterior margin of the third
segment. Antennal acicle tapering, narrow at the end, reaching one
third the length of the thick antennular flagellum. Peduncle reaching
beyond first segment of antennular peduncle; flagellum two thirds as
long as body.

Maxillipeds reaching end of antennal scale; first pair of feet to end of
antennal peduncle. The propodi of the third, fourth, and fifth pairs are
three times as long as their dactyli.

Margins of first to third abdominal segments entire; of fourth to sixth

Ss =
—_
= J

68 RATHBUN

segments with a spine at the lower postero-lateral angle; sixth segment
less than twice as long as fifth, and only a trifle longer than wide. Tel-
son with four pairs of lateral spinules.

Dimensions.— 4, length 14 mm.

Type locality.—Heceta Bank, Oregon, 50 fathoms (AJ/datross station
2886); one specimen.

Aside from the peculiar form of the rostrum, this species differs
notably from 5S. sfiva in its subequal and more widely separated supra-
orbital spines. Its habitat is also farther south than the known range of
S. spina.

SPIRONTOCARIS LILJEBORGII (Danielssen).

Hippolyte Lifjeborgii DANIELSSEN, Nyt Mag. for Naturvid., XI, 5, 1859
(brief description; Vol. XI appeared in 1861, but the separate contain-
ing Danielssen’s article was issued in 1859).—DANIELSSEN and BOECK,
Nyt Mag. for Naturvid., x1x, 196 [8], plate, figs. 15-20, 1872.

Hippolyte securifrons NORMAN, Rept. Brit. Assoc. Adv. Sci. for 1861, XXXI,
151 (1862); Trans. Tyneside Naturalists’ Field Club, v, 267, pl. XII,
figs. 1-7, 1863.—SMITH, Trans. Conn. Acad. Arts Sci., v, 69, 1879.

Distribution.—East of Point Franklin, Arctic Alaska (lat. 71° 02/ 00”
N., long. 157° 46 oo” W.), 19 fathoms, August 24, 1884, one small male
(U. S.R.S. Corwin). Known from the Atlantic coast of America, from
off Nova Scotia to Delaware Bay, 27-452 fathoms; northern Europe.

SPIRONTOCARIS BISPINOSA Holmes.

Spirontocaris bispinosa HOLMES, Occas. Papers Calif. Acad. Sci., VII, 207,
1900. Puget Sound.

Dimensions.—Ovigerous ? , length
ae td SEED (approximate) 59.5 mm., length of
carapace and rostrum 25.2 mm., of
rostrum 14.8 mm.
Fic. 23. Sfzrontocaris bispinosa. Side of cara. Distribution. — From Strait of
OE AN ees Fuca to off San Diego, California,
13-211 fathoms. Taken by the Albatross at:

Strait of Fuca, 116 and 98 fathoms, stations 3447, 3448.

Puget Sound, 82 fathoms, station 3067.

Off Tahwhit Head, Washington, 178 fathoms, station 3076.

Off Tillamook Rock, Oregon, 29 fathoms, station 2882.

Off Bodega Head, California, 167 and 62 fathoms, stations 3170, 3172.
Off Farallone Islands, California, 191 fathoms, station 3161.

Monterey Bay, 13-68 fathoms, stations 3132, 3134, 3666.

Off Lobos Rocks, 77 fathoms, station 3184.

Off San Simeon Bay, 211 and 160 fathoms, stations 3191, 3193.

Off Esteros Bay, 92 fathoms, station 3194.

DECAPODS 69

Southwest of San Nicholas Island, 158 fathoms, station 2898.
Off San Diego, 124 fathoms, station 2935.

SPIRONTOCARIS SNYDERI Rathbun.
Spirontocaris snydert RATHBUN, Proc. U. S. Nat. Mus., XxIv, 894, 1902.

This species bears a curious resemblance to 5S. dispinosa, in general

terms it is a dispinosa with the slender spine of the rostrum broken off;
but a closer examination shows other and more radical differences. The
specimens, among which are several egg-bearing fe-
males, are all smaller than S. dispinosa, but this is not ‘
absolute proof that the species is smaller, because the
females of S. dispinosa are fertile when rather small.
The median spines of the carapace proper are 3 OF gupnit idee? Caan
4, instead of 2, as in S. dispinosa, and are larger; ° * (78) Station 2886.
the last spine is at the posterior two fifths of the carapace. Between
these spines and the small ones on the rostrum there is a considerable
space. The rostral teeth are 5 or 6 above, 3 or 4 below. The rostrum
is a little shorter than the carapace and is much the shape of that of
S. bispinosa, excepting that the midrib is more rapidly ascending and
ends in a short point in front of the laminz, and the upper lamina is
shallower and of rather even depth throughout its length.

The antennal peduncle reaches only about a third the length of the
scale, in .S. dispinosa half the length of the scale. The first pair of
chelipeds reach the end of the antennal peduncle; in S. dispinosa they
are shorter. The dactyls of the third, fourth, and fifth pairs of feet,
which are very long in SS. dispinosa (about half as long as the propodi),
are in S. szyderi a third, or less than a third, as long as their propodi.

Dimensions.—Ovigerous @, length (approximate) 28 mm., length of
carapace and rostrum g.4 mm., of rostrum 4.4 mm.

Distribution.—From Puget Sound to Lower California, as follows:

Puget Sound (T. Kincaid).

Heceta Bank, Oregon, 50 fathoms (A/batross station 2886).

Monterey Bay, California, type locality (J. O. Snyder).

Near Lobos Rocks, 77 fathoms (A/ébatross station 3184).

Off Santa Catalina Island, 47 fathoms (A/éarross station 3663).
Southern California (W. H. Dall).

Off Cerros Island, Lower California, 44 fathoms (A/batross station 2838).

SPIRONTOCARIS SICA Rathbun.
Spirontocaris sica RATHBUN, Proc. U. S. Nat. Mus., XXIV, 894, 1902.

Allied to S. dispinosa, but inhabiting deeper water on the coast of
California. Differs as follows: The teeth of the dorsal carina begin

70 RATHBUN

considerably in front of the middle of the carapace (exclusive of ros-

trum); in S. dispinosa they begin at or a little posterior to the middle.

Rostrum shorter; the upper and lower

laminz are narrower and extend nearer

the tip; there are g to 14 superior teeth

Pe Reiter nmr gran Ws WE (in S. dispinosa 10 to 12), of which two

pace of ¢ (X 1§). Station 3200. are on the carapace; 3 to 5 inferior teeth

on the lamina (the same in 5S. dsfinosa), but none on the slender termi-
nal portion.

The second segment of the antennular peduncle is longer than in S.
bispinosa, being three times as long as the third segment, while in S.
bispinosa it is twice as long. The outer maxillipeds are longer, reaching
beyond the antennal scale; in \S. d¢spinosa they do not reach the end of
the scale. The thoracic feet are longer and slenderer, the first pair over-
reaching the antennular peduncle; in S. dzspinosa they fall short of the
peduncle. The last pair exceed the antennal scale by the length of
the dactylus and one half the propodus; in S. dispinosa, by the length
of the dactylus only.

Dimensions.— 2, length 58 mm., length of carapace and rostrum 24.8
mm., of rostrum 12.8 mm.

Type locality. — Santa Barbara Channel, California, 265 fathoms
(Albatross station 3200).

Distribution. —Off the coast of California, from Point Arena to San
Diego, in 211-464 fathoms, at 19 stations of the Albatross.

SPIRONTOCARIS PHIPPSII (Kroyer).

Hippolyte Phippsii KROYER, Naturh. Tidssk., 111, 575, 1841 (4); K. Danske
Vidensk. Selsk. Afhand., IX, 314, pl. III, figs. 64-68, 1842 ( ¢ ).

Hippolyte turgida KROYER, Naturh. Tidssk., 111, 575, 1841 (2); K. Danske
Vidensk. Selsk. Afhand., 1x, 308, pl. II, figs. 57, 58, and pl. III, figs.

59-63, 1842 ().
Hippolyte vibrans STIMPSON, Ann. Lyc. Nat. Hist. N. Y., X, 125, 1871 (¢).

Female. —Posterior spine of dorsal crest at about the middle of the
carapace; 7 to 12 dorsal spines, which diminish on the rostrum; 3 or 4
oncarapace. Midrib of rostrum slightly inclined upward, nearly straight ;
upper limb diminishing gradually in depth toward the tip; lower limb
deeper than the upper, triangular, 4 to 7 spines on distal half; tip acute,
reaching a little beyond the peduncle of the antennula. On the anterior
portion of the carapace are 2 supraorbital spines, one above and a
little behind the other, the lower smaller; an antennal spine; and a well-
developed pterygostomian spine.

The antennular scale reaches the end, or nearly the end, of the second

DECAPODS "I

segment of the peduncle; the eye extends to the second segment, which,
as well as the third, is very short. The antennal peduncle falls short of
the middle of the scale; scale short and broad, subequal in length to the
rostrum.

The maxillipeds exceed the antennal scale a little. The palm of the
first pair of feet overlaps the last segment of the maxillipeds. The last
three pairs of feet are rather stout, and the last pair exceeds the antennal
peduncle a little.

Abdomen conspicuously punctate.

Male.—The rostrum is horizontal or deflexed, and very slender, and
the spines are small and appressed. Peduncles of antennule more
elongate than in the @, eyes not reaching end of first segment, basal
scale extending to middle of second segment.

Dimensions. — Ovigerous 2: length (approximate) 37.5 mm., length
of carapace and rostrum 13 mm., of rostrum 5.3 mm.

Distribution.— Arctic Alaska to the Shumagins; 6-72 fathoms. Cir-
cumpolar. Atlantic coast of America southward to Cape Cod; 8-125
fathoms. Northern Europe.

In the collections of the U. S. National Museum are specimens ob-
tained by Dr. W. H. Dall at various localities along the Aleutian Islands
eastward to the Shumagins, in 6—30 fathoms; in Bering Strait, 13-17
fathoms; and in Plover Bay, Siberia, 1o—25 fathoms.

Taken by the Albatross at the following stations:

Pumicestone Bay, 35 fathoms, station 3322.

Off North Head, Akutan Island, 72 fathoms, station 2842.

Unimak Pass, 34 fathoms, station 3220.

North of Bird Island, Shumagins, 21 fathoms, station 2850.

ro miles west of Point Franklin, Alaska, 1334 fathoms (Point Barrow
Expedition).

Lat. 71° 02/ oo” N., long. 157° 46/ 00” W., 19 fathoms (U. S. R. S.
Corwin).

Off Point Hope, Alaska, 25 fathoms (U.S. R. S. Corwin).

SPIRONTOCARIS OCHOTENSIS (Brandt).
Lippolyte ochotensis BRANDT, Middendorff’s Reise Sibir., Band 11, Zool.,
Theil ¥, Krebse, p. 120, pl. Vv, ‘fig. 17, 1$51:

Dorsal crest arising at about the middle of the carapace; 3 large
spines on the carapace. Rostrum reaching to end of antennular
peduncle; midrib ascending in its terminal portion; upper lamina con-
vex above, tapering forward and backward, armed with about 6 or 7
small spines distant from those on the carapace; tip usually bifid; lower
lamina deep, subtriangular, armed with 4 or 5 small spines on the anterior

72 RATHBUN

half. Two spines above the eye, the posterior one above and much
larger than the anterior.

Outer spine of antennula reaching the end of the second segment or

a little beyond it; flagella reaching only a little

SS beyond antennal scale. Scale broad, about two

thirds as long as carapace. The maxillipeds

reach a little beyond the antennal scale. The

dent Side of can h'@ Palms of the first pair of feet extend to the last

Crab) Pore Molle: segment of the maxillipeds. The last 3 pairs of

feet are stoutish, the last pair scarcely reaching the end of the anten-
nal peduncle.

The telson is a little more than one and a half times as long as the
sixth segment, and is armed with 4 pairs of lateral spinules.

In the males the carapace is flatter, the rostrum more horizontal, less
upturned.

Dimensions.— 3 , length 40 mm., length of carapace and rostrum 13.5
mm., of rostrum 5.6 mm.

Distribution. — Bering Sea to Sitka; Kamchatka; Okhotsk Sea;
Japan.

Taken by W. H. Dall at 14 stations from Nunivak southward and
along the Aleutian Islands and Alaska Peninsula to Sitka, 5-20 fathoms.
Off Bristol Bay, 74%{-1434 fathoms (Addatross stations 3232, 3233,

236).
of ine Bay, 15 fathoms (A/batross station 3300).
Lat. 63° 50’ oo” N., long. 167° 21/ oo” W., 17 fathoms (Lieutenant

George M. Stoney, U. S. N.).

Sitka (Harriman Expedition).

Bering Island (L. Stejneger).

Bering Island, in stomach of Gadus macrocephalus (N. Grebnitzky).
Petropavlovsk, Kamchatka, 10-12 fathoms (N. Grebnitzky).
Okhotsk Sea (Brandt).

Hakodate Bay, Japan (Stimpson).

SPIRONTOCARIS DALLI Rathbun.

Spirontocaris dalli RATHBUN, Proc. U. S. Nat. Mus., XXIV, 894, 1902.

Female.—Allied to S. ochotensis, from which it is separated by few
but well-marked characters. The rostrum is longer than in SS. ochotensis,
reaching nearly to the end of the antennal scale; the midrib is straighter,
less sinuous, and terminates in a single sharp spine; the teeth on the ros-
trum are 6-8 above, 3-4 below. The carapace is lower. The last 3
pairs of thoracic feet are longer and more slender. The sixth segment
of the abdomen is also longer than in S. ochotensis, being more than one
and a half times as long as the fifth segment.

DECAPODS 73

Male.—As compared with the female, the teeth of the median carina
and rostrum are much smaller and more appressed, sometimes obsoles-
cent, the rostrum less deep and more horizontal; the antennular acicle
does not reach the end of the second segment of ——*
the peduncle; the spine of the antennal scale falls x
short of the end of the blade.

Dimensions. — 2 , length (approximate) 38 mm., =~
length of carapace and rostrum 13.6 mm., of ros- 7)
trum 6 mm. °

Distribution.— Arctic Alaska, Aleutian Islands pig a7 spivontocaris dallé
to Sitka. (Xx 22). Attu.

: P a. Side of carapace of 8.

Although this species appears to be not rare 4. Side of carapace of ¢.
in Alaska, it has been collected almost exclusively by Dr. Dall, who ob-
tained it at 17 stations along the Aleutian Islands and eastward to Port
Etches, 6-20 fathoms; off Cape Sabine, 13 fathoms; and 15 miles off
Cape Krusenstern, 14 fathoms.

Sitka, 2 specimens (Harriman Expedition, W. R. Coe, collector).

SPIRONTOCARIS POLARIS (Sabine).

Alpheus polaris SABINE, Supplement to Appendix of Parry’s [First] Voyage,
p- ccxxxviii, pl. 11, figs. 5-8, 1824.

flippolite borealis JAMES C. Ross, in John Ross, Appendix to Narrative of
Second Voyage in Search of a North-West Passage, p. Ixxxiv, pl. B, fig.
3, 1835 (2).

Lippolite Sr JAMES C. Ross, of. ci?., p. Ixxxv (9 ).

fTippolyte St. Pauli BRANDT, Middendorff’s Reise Sibir., Band 11, Zool.,
Theil 1, Krebse, p. 118, pl. v, fig. 19, 1851.

fHTippolyte cultellata NORMAN, Rept. Brit. Assoc. Adv. Sci., 1866, p. 200
(1867) (= H. polaris, teste Norman, Museum Normanianum, II, Crus-
tacea, p. 8, 1886).

LfTippolyte polaris SMITH, Trans. Conn. Acad. Arts Sci., v, p. 80, pl. XI, figs.
I-4, 1879.

fTippolyte Amazo PFEFFER, Jahrb. Hamburg. Wiss. Anst., III, 46, plate,
figs. 7a, 7b, 1886.

fTletairus gaimardti BATE, Challenger Rept., Zool., XXIV, 611, pl. CIX, fig.
2, 1888 (not Hippolyte gaimardii Milne Edwards).

Fletatrus tenuis BATE, of. cit., 613, pl. CIX, fig. 3.

fTetairus debilis BATE, op cit., 615, pl. CIX, fig. 4.

Professor Smith (/oc. cit.) sets forth the variations in the sexes, in the
number of rostral spines, and in the aculei on the telson. He states that
“the disappearance of the dorsal teeth of the rostrum is evidently a
character peculiar to, but not characteristic of, the adult male.” In the
series of specimens from the North Pacific and Bering Sea this is not
the case, as the majority of the females from those localities are devoid
of superior rostral teeth. This series also exhibits other variations. The
body and thoracic feet are usually stouter, the antennal scale shorter and

74 RATHBUN

broader, the antennular scale, on the contrary, commonly longer than in
Atlantic forms, in the ? often reaching the end of the peduncle, and in
the ¢ sometimes the end of the second segment. ‘There is, however, a
lack of constancy in these characters which prevents the separation of
the stout form as a distinct species; it was described by Brandt as H. S¢.
faul. Again, the rostrum may be only as long as the peduncle of the
antennula, or reach nearly to the end of the antennal scale; it may be
of greater or less depth and acuteness. From the variations at a single
locality, Bate (/oc. cit.) described three species for which he constructed
the genus He/airus.

Distribution.—Circumpolar. Atlantic coast of North America south-
ward to Cape Cod, 10-218 fathoms. Northern Europe. Bering Sea,
Okhotsk Sea, Aleutian Islands eastward to Kadiak, to a depth of 283
fathoms :

Taken by the Albatross :

Off Pribilof Islands, 25 fathoms, station 3558.
Off Rat Islands, Aleutians, 55 fathoms, station 3599.
Off Seguam, Aleutians, 283 fathoms, station 3480.
Pumicestone Bay, 35 fathoms, station 3322.
Northwest of Unimak Island, 41 and 43 fathoms, stations 3259, 3262.
Northeast of Cape Leontovich, 22 fathoms, station 3275.
Okhotsk Sea, 10 fathoms, station 3645.
Collected by Dr. W. H. Dall at:

Plover Bay, East Siberia, 10-25 fathoms.

Near Indian Point, Bering Strait, 17 fathoms.

Various localities along the Aleutian Islands eastward to Kadiak, 3-80
fathoms.

St. Paul Island, Pribilofs (William Palmer).
Bering Island (L. Stejneger).
Bering Island, among Laminaria (N. Grebnitzky).

SPIRONTOCARIS UNALASKENSIS Rathbun.
Spirontocaris unalaskensis RATHBUN, Proc. U. S. Nat. Mus., XXIV, 895,
1902.

Female.—Body rather robust. Carapace carinated in its anterior half;
posterior median spine situated at the anterior third of the carapace.
Rostrum longer than the carapace or the antennal scale, nearly horizontal,

but slightly concave above, dor-
sal spines 4 or 5, equally spaced,
2 on the carapace and 2 or 3 at
base of rostrum; no upper limb
on distal portion, lower limb

Fic. 28. Sfzrontocaris laskensis. Sid .
Pete (ke) Beatin gaa arP#* shallow, convex, armed with three

DECAPODS Vis

spines in front of middle, tip acuminate, sometimes minutely bifid. An-
terior margin of carapace armed with 3 spines, one supraorbital, one
antennal, and one smaller, pterygostomian. Eyes large, pyriform.

First two segments of antennules with an antero-external spine, third
segment less than half as long as the second and armed with a spine at
the middle of its anterior upper margin; the scale at outer base over-
reaches the first segment. Outer flagellum reaching but little beyond the
rostrum ; inner flagellum twice as long.

Antennal peduncle extending nearly to the end of the second segment
of the antennular peduncle. Scale oblong, tapering, shorter than the
carapace, tipped with a small spine which is exceeded by the blade.
Flagellum at least two thirds as long as the body.

Tip of maxilliped intermediate between rostrum and antennal scale.
Carpal joint of first pair of feet reaching end of antepenult segment of
maxilliped. Second and fifth pairs of feet reaching about to end of
rostrum.

Abdomen ecarinate, third segment very prominent, its hinder portion
extending well behind the line of the fourth segment. The sixth segment
is about one and two thirds as long as the fifth, and seven ninths as long
as the telson, which is armed with 5 or 6 spinules on each side.

Dimensions. —A female, soft shell, measures about 79 mm. in length;
a smaller female is 60 mm. long, carapace and rostrum 26.5 mm., rostrum
14.5 mm.

Distribution.— North of Unalaska, 277-351 fathoms, A/éazross stations
3315, 3330, 3331 (type locality).

Variations.—In the male the rostrum is slenderer and more sinuous,
the antennal flagella considerably longer. In small specimens the rostrum
is relatively shorter.

SPIRONTOCARIS VICINA Rathbun.

Spirontocaris vicina RATHBUN, Proc. U. S. Nat. Mus., xxIv, 895, 1902.

Allied to S. unalaskensis.

Female.— Posterior dorsal spine at anterior third of carapace. Ros-
trum longer than the carapace or antennal
scale, concave above, dorsal spines 7 or 8, of
which two are on the carapace, the anterior
not very near the tip, inferior spines 4, tip Tie a eee ee
broken off in our specimens. Anterior carapace of ¢ (x 13). Station 3316.
margin 3-spined, the pterygostomian spine the smallest. Eyes large,
pyriform.

The antennular peduncles reach nearly to the spine of the antennal

76 RATHBUN

scale; the second segment is about three times as long as the third; the
basal scale falls considerably short of the second joint of the peduncle.
The antennal peduncle does not reach the middle of the second segment
of the antennular peduncle; scale similar to that of S. uzalaskensis.

Abdomen like that of the preceding species. ‘The telson is incomplete
in all the females.

Male.—In the single male, the rostrum is a little shorter than the cara-
pace and does not reach the end of the antennular peduncle; it is also
straighter than in the female, and is armed with 6 spines above and 3
below.

Dimensions.—A female with rostrum and telson broken measures 39
mm. from the orbit to the end of the sixth abdominal segment. <A female
about the same size, in very bad shape, is laden with eggs, each about 3
mm. in its greatest diameter. Male, length 50 mm., length of carapace
and rostrum 18.5 mm., of rostrum 8.5 mm.

Distribution.—North of Unalaska, 309-406 fathoms, stations 3316

(type locality), 3330, 3332.

SPIRONTOCARIS WASHINGTONIANA Rathbun.

Spirontocaris washingtoniana RATHBUN, Proc. U.S. Nat. Mus., xxIv, 895,
1902.

Female.—Carapace stout, carinated in its anterior half; posterior
median spine situated at the anterior fourth of the carapace. Rostrum
slender, half as long as the carapace, reaching just to the end of the first

segment of the antennules, nearly horizontal,
slightly sinuous; dorsal spines 4, two of which
are on the carapace, anterior two fifths of rostrum
unarmed above; 3 teeth below on anterior third,
Fic. 30. Sfirontocaris wash. One tooth close to the tip; except for the teeth,
ingtontana. Side of carapace of 3 °
@ (X 2}). Station 3071. the rostrum is scarcely limbed above or below.
Anterior margin armed with 3 spines, one supraorbital, one antennal, one
smaller at the antero-lateral angle.

Antennular peduncles reaching nearly to the base of the spine of the
antennal scale; first segment with a large bifurcate spine on the anterior
margin toward the outer angle; second segment with a long simple
spine in the same position; third segment with a small dorsal spine at
the middle of its anterior margin; the first segment is about twice as long
as the second, the second three times as long as the third; the flagella
are subequal in length, and half again as long as the peduncle; scale
about two thirds as long as first segment. The antennal scale is two
thirds as long as the carapace, oblong-lanceolate, extremity of blade

DECAPODS "7

oblique and exceeding the spine; the peduncle attains the middle of the
scale; flagellum about as long as the body.

The maxilliped extends beyond the antennal scale by half the length
of its last segment; the first pair of feet just reaches the end of the
antennal scale, the second pair extends a little beyond the maxilliped,
the fifth pair not quite so far as the maxilliped. Second to fifth pairs
very slender.

The abdomen is narrow compared with the carapace; the posterior
margin of the third segment is moderately produced; the sixth segment
is one and a half times as long as the fifth, and a little more than one
half as long as the seventh; this last has 5 or 6 spinules on either side.

Dimensions.— 9, length 39 mm., length of carapace and rostrum 14
mm., of rostrum 5 mm. A mutilated specimen of the same size is
ovigerous.

Lyfe locality.— Off Sea Lion Rock, Washington, 685 fathoms (Ad/batross
station 3071).

SPIRONTOCARIS AFFINIS (Owen).

Hippolite affinis OWEN, in Zoology of Capt. Beechey’s Voyage [of the
Blossom], Crustacea, p. 90, pl. XXVI, fig. 4 (rostrum).

Locality.— Monterey, California (Owen).

SPIRONTOCARIS GRACILIS (Stimpson).

Hippolyte gracilis STIMPSON, Proc. Acad. Nat. Sci. Phila., 1864, 155.
Heptacarpus tenuissimus HOLMES, Occas. Papers Calif. Acad. Sci., VII,
203, 1900.
Heptacarpus? gracilis HOLMES, of. cit., p. 205.
Distribution. —From the Shumagins, Alaska, to Monterey, California:
North of Bird Island, Shumagins, 21 fathoms (A/batross station 2850).

Off Cape Beale, Vancouver
Island, British Columbia,
34 fathoms (Adbarross sta-
tion 2879).

Off Cape Flattery, Washing- ;
ton, 40 fathoms (Albatross Big. 3%, Sooners ere
station 2873).

Off Drake Bay, California, 35 fathoms (4/batross sta-
tion 3155).

Off Point Reyes, California, 47 fathoms (A/batross
station 3166).

Monterey Bay, California (J. O. Snyder).

Puget Sound (T. Kincaid, 1895).

Puget Sound, deep water (Stimpson); Monterey
(Holmes).

78 RATHBUN

On the few specimens examined the superior teeth are 4; the inferior
vary from 4 to 8.

Stimpson says that the thick flagellum of the antennulz extends to the
end of the rostrum; this is probably true of the 4. All the specimens in
hand are females, in which the flagellum extends only about two thirds
the length of the rostrum.

Dimensions.— Approximate length of 9 with eggs 48.4 mm., length of
carapace and rostrum 18 mm., of rostrum 10.5 mm.

SPIRONTOCARIS FLEXA Rathbun.

Spirontocaris camtschatica RATHBUN, The Fur Seals and Fur-Seal Islands
of the North Pacific Ocean, Pt. III, 557, 1899. Not S. camtschatica
(Stimpson).

Spirontocaris flexa RATHBUN, Proc. U. S. Nat. Mus., XxIv, 896, 1902.

Female.—Slender. Carapace crested on its anterior third; rostrum
longer than the carapace, armed with 4 or 5 teeth above, including 1 or

2 on the carapace; anterior tooth near middle of rostrum; anterior half

of rostrum not limbed above. Rostrum very slender, nearly horizontal, a

Fic. 32. Spirontocaris flexa. @. Station 2850. a. Side of carapace and abdo-
men (X 2). 8. Dorsal view of anterior portion (X 3).

little arched above the eye; lower limb narrow and armed with from 5 to
8 spines; tip reaching nearly to or a little beyond the end of the antennal
scale. An antennal and a pterygostomian spine, the latter well developed.

Antennular peduncle reaching one third the length of the antennal
scale; basal spine to a little beyond first segment; thickened portion of
outer flagellum reaching to about distal third of antennal scale; inner
flagellum to end of scale. Spine of antennal scale not nearly as advanced
as the membranous portion. Peduncle of antenna advanced as far as
the second antennular segment; flagellum nearly as long as body.

DECAPODS 79

Maxillipeds produced to about middle of antennal scale; first pair of
feet to end of antennal peduncle; second pair to a little beyond the
maxillipeds; the last pair of feet to the end of the maxillipeds.

The abdomen is strongly geniculated or bent at a right angle at the
third segment, which is posteriorly produced and compressed, forming
a smooth, rounded carina. The first to fourth segments are without
spines or spinules; the sixth is more than twice as long as wide, but
shorter than the seventh, which is narrow and has 4 spinules on each
side.

Male.—The thickened portion of the outer antennular flagellum ex-
tends nearly to the end of the antennal scale.

This species is very near S. gracilis, from which it differs in the longer
basal scale of the antennula, in the lower position of the pterygostomian
spine, in the presence of epipods on the maxilliped and first two pairs of
Tek,

Dimensions.— 9, length 54 mm., of carapace and rostrum 20 mm., of
rostrum 11.5 mm.

Distribution.—¥From Bering Sea to Drake Bay, California:

Bering Island, one specimen from stomach of Gadus macrocephalus (N.
Grebnitzky).
Alaska, probably Bering Sea (U.S. R. S. Corwin).
Nazan Bay, Atka (W. H. Dall).
Albatross stations:
Off Pribilof Islands, 20 fathoms, station 3438.
Off Rat Islands, Aleutians, 55 fathoms, station 3599.
Jliuliuk Harbor, Unalaska, 93 fathoms, station 3335.
Off North Head, Akutan Island, 72 fathoms, station 2842.
Off south entrance to Akutan Pass, 45 fathoms, station 2843.
Unimak Pass, 34 and 50 fathoms, stations 3220, 3222.
Northwest of Unimak Island, 43 fathoms, station 3262.
South of Unimak Island, 42 fathoms, station 3217.
15 miles south of Sannak Islands, 44 fathoms, station 2846.
Shumagins, 21-58 fathoms, stations 2850 (type locality), 2851, 2852.
Drake Bay, California, 35 fathoms, station 3155.

SPIRONTOCARIS AMABILIS (Lenz).

Hippolyte amabilis LENZ, Zool. Jahrb., Syst., XIV, 432, pl. XXXII, figs. 2, 3,
1gOl.

Bare Island, British Columbia (type locality).

SPIRONTOCARIS DECORA Rathbun.
Spirontocaris decora RATHBUN, Proc. U.S. Nat. Mus., XXIv, 896, 1902.
Female.—Carapace crested on its anterior half; rostrum longer than
the carapace, armed with 4 or 5 teeth above, of which one or two are on

80 RATHBUN

the carapace, the anterior tooth in front of the middle of the rostrum.
Rostrum not deep, but less slender than in SS. gracilis and S. flexa, a lit-
tle concave
above, but:
nearly hori-
zontal ; lower
limb narrow-
ing distally
from near
the posterior end, provided with 6 to 8 teeth; tip
acuminate, reaching nearly to or beyond the an-
tennal scale. Antennal and pterygostomian spines
present, both well developed.

Antennular peduncle between one third and one
half the length of the antennal scale; spine at base
reaching about to end of first segment. Thick-
ened part of outer flagellum not reaching end of
antennal scale; inner flagellum twice as long as
outer. Outer margin of antennal scale as long as the carapace; pedun-
cle extending as far as the end of the second antennular segment. Max-
illipeds extending beyond middle of antennal scale.

The posterior margin of the third abdominal segment is produced
backward in the middle, and the segment itself is posteriorly prominent
but not laterally pinched or carinated as in allied species. The sixth
segment is more than twice as long as wide, and a little longer than
the seventh, which has 5 to 7 pairs of lateral spinules, and is shorter
than the uropods.

Male.—The thickened part of the outer flagellum of the antennula
extends to the end, or nearly to the end, of the antennal scale; the inner
flagellum is correspondingly long.

Dimensions.—Approximate length of @ 47 mm., length of carapace
and rostrum 16.4 mm., of rostrum g mm.

Distribution.—From Strait of Fuca to San Diego, California; 50 to 171
fathoms. Dredged by the A/satross at the following stations:

Spirontocarts decora.

Fic. 33.
Station 2946.

@ (X 3).

Strait of Fuca, 152, 114 fathoms, stations 3454, 3461.

Near Flattery Rocks, Washington, 171 fathoms, station 2866.

Heceta Bank, Oregon, 50 fathoms, station 2886.

Off Point Conception, California, 145 fathoms, station 2893.

Off Santa Cruz Island, California, 150 fathoms, station 2946 (type local-
ity).

Off Santa Rosa Island, California, 52 fathoms, station 2956.

Off San Diego, California, 124 fathoms, station 2935.

DECAPODS SI

SPIRONTOCARIS TRIDENS Rathbun.
Spirontocaris tridens RATHBUN, Proc. U. S. Nat. Mus., xxIv, 896, 1902.

Allied to S. gracilis, S. flexa, and S. decora, but more robust than
these.

Female.—The median crest occupies the anterior third of the carapace ;
superior spines or teeth 3, one on the carapace, one (the largest and
highest) over the base of the eye-stalk and nearer to the anterior than to
the posterior
tooth. In front
of the teeth the
rostrum has_al-
most no upper
limb and is gently
ascending and
slightly curved;
midrib rather stout
and acute at tip;
the lower limb —==

diminishes eae Fic. 34. Spitvontocaris tridens. 2. Station 2865. a@. Side of carapace and
riorly and isarmed abdomen (xX 13). 4. Dorsal view of anterior portion (x 3).

with 3 to 6 teeth, one near the tip. Rostrum longer than the carapace
or antennal scale. Anterior margin of carapace furnished with a strong
antennal and a very small pterygostomian spine.

Antennular peduncle extending two fifths the length of the antennal
scale; second and third segments short and subequal; basal scale with a
spine which extends nearly to the end of the second segment. The
thickened portion of the outer flagellum reaches nearly to the end of
the antennal scale; inner flagellum twice as long. The antennal pedun-
cle extends a little beyond the end of the second antennular segment ;
the flagellum is nearly as long as the body; the scale, measured along
its outer margin, is a little shorter than the carapace.

The maxillipeds reach two thirds the length of the antennal scale;
the first pair of feet barely to the end of the antennal peduncle. The tip
of the second pair when extended lies between the end of the maxilliped
and that of the antennal scale; the tip of the last pair lies between the
end of the maxilliped and the antennal peduncle.

The abdomen has the third segment well produced backward, and its
posterior portion carinated ; the carina is defined on either side by a deep
groove like the imprint of a nail. The sixth segment is more than twice
as long as high, and about as long as the seventh; this last is a little

82 RATHBUN

shorter than the uropods and is armed with three pairs of lateral
spinules.

Male.—The thickened portion of the outer antennular flagellum reaches
to the end or a little beyond the end of the antennal scale.

Dimensions.— @ , approximate length 61 mm., length of carapace and
rostrum 22 mm., of rostrum 12.5 mm.

Distribution. —¥From the Aleutian Islands to Washington. ‘Taken by
the Albatross at the following stations:

Unimak Pass, 34 fathoms, station 3220.

Northeast of Amak Island, 19 fathoms, station 3274.

Northeast of Cape Leontovich, 22 fathoms, station 3275.

North of Bird Island, Shumagins, 21 fathoms, station 2850.

Off Cape Flattery, Washington, 40 fathoms, station 2873.

Strait of Fuca, 48 fathoms, stations 2864, 3465.

Admiralty Inlet, Puget Sound, 40 fathoms, station 2865 (type locality).

A single specimen was taken by Dr. Dall at Port Levashef, Unalaska,
and another at Granite Cove, Port Althorp, Chichagof Island.

Variations. —Three teeth on the dorsal carina is the rule in this
species; in more than two hundred specimens handled, there was one in
which the anterior of the three teeth was absent, another in which the
posterior tooth was absent, and two cases in which there were 4 teeth
instead of 3.

Distinguished from nearly related species by the fewer dorsal teeth
and by the groove on either side of the third segment of the abdomen.

SPIRONTOCARIS BARBATA Rathbun.

Spirontocaris barbata RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, III, 556, 1899.

Anterior two thirds of carapace carinated. Rostrum from one third
to one fourth longer than the carapace, and also a little longer than the
antennal scale; nearly straight and horizontal; upper limb very narrow,
widening a trifle in front of eye, then diminishing to the tip, and armed
with 5 to 7 spines, one of which is on the carapace; anterior tooth at
about anterior third of rostrum; lower limb broader, gradually dimin-
ishing forward from a point in front of the eye, armed with about 10 to
20 teeth and denticles, of which the posterior three or four are of fair
size, the remainder very small and decreasing toward the tip, where they
are almost indistinguishable; tip acute. Antennal and pterygostomian
spines well and equally developed, the latter unusually high in position.

Antennular peduncle reaching two fifths the length of the antennal
scale; outer basal spine reaching to about the end of the first segment;
second segment one and a half times as long as third; outer flagellum

DECAPODS 83

not attaining end of scale, a little longer in male than in female; inner
one exceeding scale. Scale oblong, nearly as long as carapace, laminar
portion considerably
exceeding spine; pe-
duncle extending to
the middle of the third
joint of the antennu-
lar peduncle; flagel-
lum nearly as long as
body.

The maxillipeds ex-
tend to middle of an-
tennal scale; first pair
of feet nearly to end
of antennal peduncle ;
third, fourth, and fifth
pairs of nearly equal Fic. 35. Spzrontocaris barbata. 9%. Station 3318. a. Side of cara-
length, the third pair pace and abdomen (X 13). 8. Dorsal view of anterior portion (x 23),
reaching spine on antennal scale; second pair a little shorter.

The abdomen is sharply carinated from the middle of the third to the
end of the fifth segment; the third to sixth segments terminate in a
slender median spine, by which this species differs from all others; third
segment with a subterminal hump visible in profile. The fourth segment is
provided with a lateral spine, the seventh with 3 pairs of lateral spinules.

Dimensions.— 9, length 74.5 mm., length of carapace and rostrum
29.6 mm., of carapace 12 mm.

Distribution.—¥rom Pribilof Islands to Strait of Fuca; 45-277
fathoms (Adbatross). | Occurs usually in small numbers; most abundant
in Iiuliuk Harbor.

Taken by the A/atross at the following stations:

Off Pribilof Islands, 56-184 fathoms, stations 3484, 3487, 3488, 3489,

3491, 3492, 3494, 3495, 3497 (type locality), 3500, 3609.
Off Makushin Bay, Unalaska, 61 fathoms, station 3318.
Off Iliuliuk Harbor, Unalaska, 277 fathoms, station 3315.
Iliuliuk Harbor, Unalaska, 45-93 fathoms, stations 3311-3314, 3334-3336.
Off Akutan Island, 51~72 fathoms, stations 2841, 2842, 3547.
Off Aektok Island, 54 fathoms, station 2844.
Unimak Pass, 50-56 fathoms, stations 3222, 3223.
South of Unimak Island, 61 fathoms, station 3216.
Shumagins, 48-58 fathoms, stations 2847, 2852.
Off Shumagin Bank, 138 fathoms, station 3339.
Off the Trinity Islands, 159 and 61 fathoms, stations 2853, 3341.
Strait of Fuca, 123 and 114 fathoms, stations 3459, 3461.

al
CU |)

84 RATHBUN

SPIRONTOCARIS CARINATA (Holmes).
Heptacarpus carinatus HOLMES, Occas. Papers Calif. Acad. Sci., VII, 202,
pl. 111, fig. 60, 1900.
Distribution.— From Monterey Bay, shallow water (Holmes), to Point
Loma, California (A/batross, one specimen).
The eyes of this species are very slender. The antennular peduncles
extend about one third the length of acicle.

SPIRONTOCARIS STYLUS (Stimpson).

Hippolyte stylus STIMPSON, Proc. Acad. Nat. Sci. Phila., XvI, 154, 1864

(June).

Hippolyte esquimaltiana BATE, Proc. Zool. Soc. London, 1864 (read De-
cember 13), 666.

Hyppolyte esquimaltianus BATE, in Lord’s The Naturalist in Vancouver
Island and British Columbia, 11, 278, 1866.

A rather slender species. Rostrum about one third longer than cara-
pace, exceeding acicle, slender, somewhat styliform, horizontal and
straight, armed above with
4 or 5 teeth near its base
(one on the carapace in-
cluded) and 5 or 6 teeth
below. An antennal, but
no pterygostomian spine.

Eyes small. Antennu-
lar peduncle reaching
about two fifths the length
of the acicle; second and
third segments very short ;

Fic. 36. Spirontocaris stylus. ¢. Barclay Sound. a. Side outer flagellum scarcely
of carapace and abdomen (X 2). 4. Dorsal view of anterior por- 5
tion (X 3). reaching the terminal
fourth of the acicle; inner flagellum exceeding acicle; basal scale not
reaching beyond first segment. Antennal peduncle extending to end of
second segment of antennular peduncle; acicle (measured along outer
margin) shorter than carapace.

Maxillipeds short, slightly exceeding antennal peduncle; first pair of
feet stout, not exceeding eyes; second pair reaching to middle of acicle;
fifth pair scarcely to end of maxilliped.

The abdomen is strongly geniculated, though the third segment is
gently rounded in profile; fourth segment unarmed; sixth segment not
quite twice as long as fifth; telson with 3 pairs of side spinules.

I think there is little doubt that Bate’s 1%. esguimadtiana belongs here.

DECAPODS 85

Dimensions.— 3, length 44.5 mm., length of carapace and rostrum 19
mm., of rostrum 11.1 mm.

Distribution.— Barclay Sound and Sucia Island, British Columbia (4//a-
tross); Esquimalt Harbor, British Columbia (Bate) ; Strait of Fuca (Stimp-
son); Port Orchard, Puget Sound (U.S. Nat. Mus.); Puget Sound (T.
Kincaid).

SPIRONTOCARIS FABRICII (Kroyer).

Hippolyte Fabricit KROYER, Naturh. Tidsskrift, 111, 571, 1841; K. Danske
Vidensk. Selsk. Afhand., IX, 277, pl. I, figs. 12-20, 1842.—SMITH,
Trans. Conn. Acad. Arts Sci., V, 63, 1879.

Distribution.— Arctic coast of Alaska southward through Bering Sea
to Siberia and Alaska Peninsula (Cook Inlet); Atlantic coast of
America from Greenland southward to Massachusetts Bay. ? Europe
(Kingsley). Depth, low water to 80 fathoms.

Taken by the A/batross at the following stations:

Off Cape Newenham, 17 fathoms, station 3247.

Southwest of Hagemeister Island, 1744 fathoms, station 3246.
Kululak Bay, 14 fathoms, station 3241.

Off Kululak Bay, 17 fathoms, station 3301.

Off Bristol Bay, 30-33 fathoms, stations 3302-3303.

Off Cape Menshikof, 24 fathoms, station 3296.

Off Cape Strogonof, 26-32 fathoms, stations 3291-3294.

Off Cape Seniavin, 37 and 30 fathoms, stations 3286, 3287.
Northeast of Cape Leontovich, 18 fathoms, station 3276.
Northeast of Amak Island, 19 fathoms, station 3274.
Shumagins, 21 and 35 fathoms, stations 2850, 2851.
Southwest of Sannak Islands, 38 fathoms, station 3214,

Off Rat Islands, 34 fathoms, station 3598.

Off Robben Island, Okhotsk Sea, 10-25 fathoms, stations 3645, 3649.
Unalaska, beach.

Collected by W. H. Dall:

Off Cape Sabine, 13 fathoms.

Near Indian Point, Bering Strait, 17 fathoms.

Plover Bay, Siberia, 10-25 fathoms.

Various localities in the Aleutian Islands and eastward to Chugachik
Bay, Cook Inlet; low water to 80 fathoms.

Also collected:

Ten miles west of Point Franklin, Alaska, 1334 fathoms (Point Barrow
Expedition).

Bering Sea, lat. 63° 50! 00” N., long. 167° 21/ 00” W., 17 fathoms (Lieu-
tenant George M. Stoney, U.S. N.).

Bering Island, two specimens in stomach of Gadus macrocephalus; one
specimen at 12 fathoms (N. Grebnitzky).

Solovarennaja Bay, Kamchatka, 10—12 fathoms, alge (N. Grebnitzky).

Kadiak (Harriman Expedition, W. R. Coe).

86 RATHBUN

SPIRONTOCARIS GAIMARDII BELCHERI (Bell).

Plate 111, figs. 3, 3a.

Hippolyte belcheri BELL, in Appendix to Belcher’s Last of the Arctic Voyages
in Search of Sir John Franklin, 11, 402, pl. XXXIV, fig. 1, 1855.

2? Hippolyte dayi LOCKINGTON, Bull. Essex Inst., X, 161, 1878 (not HZ. dayz
Owen).

Hippolyte gaimardit MURDOCH, Rept. Exped. Point Barrow, p. 140, 1885

ee gibba RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. III, 556, 1899.

Hippolyte gibba BIRULA, Ann. Mus. Zool. Acad. Impér. Sci. St. Pétersbourg,
VII, 428 [11], text fig. 1, 1899 [1900] (probably not H. gzbba Kroyer).

Female.—Robust. Rostrum longer than the carapace, nearly hori-
zontal posteriorly, anterior half ascending, armed above with 8 to 12
teeth (including 2 to 4 on the carapace), below with 3 to 5 teeth; ros-
trum tapering gradually to an acuminate tip, which exceeds the antennal
scale. A strong antennal and a well-developed pterygostomian spine.

Eyes rather large, pyriform. Scale at base of antennules reaches one
third the length of the second segment of the peduncle. Peduncle half
as long as antennal scale; second segment twice as long as third;
thickened portion of outer flagellum not reaching the tip of antennal
scale, but sometimes as far as the acicular spine. This scale is shorter
than the carapace; blade arcuate at extremity and exceeding the spine.
The peduncle reaches about two fifths the length of the scale; the
maxilliped to the distal third of the same; the first pair of feet to
the end of the antennal peduncle; the second pair to a little beyond
the antennal scale; the fifth pair overreaches the maxillipeds.

The fourth and fifth segments of the abdomen are each armed with
a postero-lateral spine. The third segment is laterally compressed in its
posterior part, which has an angular median lobe, seen in profile, a little
in front of the posterior margin; on very large females this lobe becomes
hook-shaped (as on one 80 mm. long). The sixth segment is about two
and a half times as long as high, and shorter than the seventh, which is
armed with 5 to 8 spines on each side. The outer uropods are longer
than, the inner ones shorter than, the telson.

Male.—More slender than the female. Rostrum more nearly straight
and horizontal, 8 or g teeth above. Thickened portion of outer anten-
nular flagellum exceeding acicle. Lobe on third segment of abdomen
more prominent and hooked.

Young.—In those one inch long and smaller the abdomen shows no
median lobe on third segment, though it is slightly compressed. The
antennular scale does not reach beyond the first segment of the peduncle.

DECAPODS 87

Typical form.—In the typical S. gaimardii (Milne Edwards), of which
Hippolyte gibba Kroyer is probably the male form, there is in the female
no lobe on the third abdominal segment (in a side view), although the
segment is somewhat compressed; the rostrum is distinctly shorter than
the antennal scale, scarcely if at all longer than the carapace, and is
armed with fewer spines. In both sexes the scale is longer and wider
in the typical form, being about four fifths as long as the carapace.

Distribution.— S. gaimardii belchert is found from the Arctic coast of
Alaska and Siberia southward to Sitka, taking the place of typical
S. gaimardii; it occurs also in Kara Sea (Dijmphna Exped.), Nova
Zembla (Birula), and at Nakvak, Labrador (L. M. Turner).

Lat. 70° 15/10 N., long. 162° 55’ 00’ W., 16 fathoms; lat. 719 02!
oo” N., long. 157° 46/ 00’ W., 19 fathoms; and off Point Hope, Alaska,
25 fathoms (U.S. R. S. Corwin).

Ten miles west of Point Franklin, 13% fathoms (Point Barrow Exped.).

Bering Strait and Bering Sea in lat. 66° 12’ oo” N., long. 168° 54!
oo” W., 30 fathoms; lat. 64° 12’ oo” N., long. 162° 52’ oo! W., 17
fathoms; lat. 62° 15’ oo’ N., long. 167° 48’ 00” W., 20% fathoms
(Lieutenant George M. Stoney, U. S. N.).

Plover Bay, Siberia, 10 to 25 fathoms; Cape Lisburne; off Cape Sa-
bine, 13 fathoms; Bering Strait, 13 fathoms; Sitka Harbor, zg fathoms
(W. H. Dall).

Off mouth of Yukon, 31% fathoms (E. W. Nelson).

Off St. Matthew Island, 37 fathoms, station 3519; off Pribilof Islands,
50 to 52 fathoms (A//azross stations 3527 and 3611).

That the de/cheri form cannot be considered a species is shown by the
existence of specimens which unite the characters with those of S. gai-
mardi. In three females from Henley Harbor, Labrador, the abdomen is
without a lobe, but the rostrum is longer than the rest of the carapace,
and just as long as the acicle; the acicle is long as in typical S. gaz-
maradit.

SPIRONTOCARIS TOWNSENDI Rathbun.

Spirontocaris gaimardii RATHBUN, The Fur Seals and Fur-Seal Islands of
the North Pacific Ocean, Pt. III, 556, 1899 (part).
Spirontocaris townsendi RATHBUN, Proc. U. S. Nat. Mus., XXIV, 897, 1902.

On the Pacific coast there is a large group of species of the gaimardii
type, which can be distinguished only by the closest observation. Of
these species, .S. fownsendi resembles S. gaimardii and S. gaimardii belcheri
in having the outer maxilliped provided with an exopod and epipod, and
the first two feet also with epipods. S. éowmsendi continues the range of

83 RATHBUN

S. gaimardi belcheri southward. <As typical S. gaimardii does not
occur on the Pacific coast, I will compare the new form with S. gaz-
mardi belcheri, and indicate the characters by which it differs from the
latter.

Female.—In S. townsendi the rostrum reaches almost or quite to the
end of the acicle, and is armed above with 5 to 7 (2 on the carapace)
spines, below with
3 to 6 spines; the
lower limb is deeper
than in S. gaimardii
belcheri, more as in
typical S. gaimar-
ait, but the rostrum
is nearly straight.
The pterygostomi-
an spine is very
small. The scale
at the outer base
of the antennula
reaches to the end
or nearly to the end

Fic. 37. Spirontocaris townsendi. ¢%. Station 2865. a. Side (x 23). of the second seg-

4. Dorsal view of anterior portion (x 3). ment; the second

segment is scarcely longer than the third. The antennal scale is almost

as long as the carapace. The maxillipeds reach to the distal fourth of

the antennal scale; the fifth pair of feet do not attain the end of the
maxilliped.

The third abdominal segment is smoothly rounded, without lobe or
angle, in a profile view; posterior margin produced moderately backward
at the middle. The fourth segment is devoid of a lateral spine; the sixth
is about twice as long as high; the telson is provided with 3 or 4 lateral
spinules on each side.

Male.—The males, which are fewer in number in the collection than
the females, appear to be smaller and a little more slender; otherwise,
save for their longer antennule, they present no differences from the
females.

Dimensions.—Female, approximate length 60.5 mm., of carapace and
rostrum 20 mm., of rostrum 10.6 mm.

Distribution.— Ranges from the Pribilof Islands to Puget Sound, and
from 21 to 114 fathoms. One exception to this bathymetrical range is
238 fathoms in Queen Charlotte Sound, British Columbia (4/batross

DECAPODS 89

station 2862); here only one specimen was taken, of which the rostrum
and tail are missing; the characters of the remaining part coincide with
those of typical specimens.
Other stations at which this species was collected by the A/Jatross are:
Off Pribilof Islands, 25-60 fathoms, stations 3484, 3496, 3558.
Off Rat Islands, Aleutians, 55 fathoms, station 3599.
Unalaska, 35-85 fathoms, stations 3311, 3319, 3321, 3322.
Off North Head, Akutan Island, 56 and 72 fathoms, stations 2841, 2842.
Off south entrance to Akutan Pass, 45 fathoms, station 2843.
Unimak Pass, 34-56 fathoms, stations 3220, 3222, 3223.
Northwest of Unimak Island, 43 fathoms, station 3262.
Southwest of Sannak Islands, 41 fathoms, station 3213.
15 miles south of Sannak Islands, 44 fathoms, station 2846.
Off Kudobin Islands, 36-41 fathoms, stations 3279, 3281.
Shumagins, 48 and 21 fathoms, stations 2847, 2850.
Off Cape Strogonof, 26 fathoms, station 3291.
Portlock Bank, 51-68 fathoms, stations 2856, 2857.
Admiralty Inlet, Puget Sound, 40 fathoms, station 2865.
Washington Sound, Straits of Fuca, 48 fathoms, station 2864 (type
locality).
Strait of Fuca, 48-114 fathoms, stations 3443, 3445, 3461, 3462, 3465.

Granite Cove, Port Althorp, Chichagof Island, Alaska (W. H. Dall).
Puget Sound (T. Kincaid).

SPIRONTOCARIS SUCKLEYI (Stimpson).

Hippolyte suckleyt STIMPSON, Proc. Acad. Nat. Sci. Phila., 1864, 154; LENZ,
Zool. Jahrb., Syst., XIV, 432, pl. XXXII, fig. 1, 1901.

Spirontocaris gatmardit RATHBUN, The Fur Seals and Fur-Seal Islands of
the North Pacific Ocean, Pt. 111, 556, 1899 (part).

Female.—The nearest approach in the Pacific to the S. gaimardi of
the Atlantic. Associated with S. zownsendi, with which it may easily be
confounded; distinguished from it as follows:

The rostrum is rather more concave above instead of nearly straight ;
it reaches end of, or may exceed, antennal scale.

The antennular scale does not reach the end of the second segment of
the peduncle.

The antennular peduncle reaches middle of antennal scale (length
measured along outer margin). The second segment is appreciably
longer than the third.

The antennal scale, measured on its outer margin, is much shorter than
the carapace.

The outer maxillipeds overlap the distal fourth of the antennal scale,
and may reach the end of the scale.

The second pereiopod is commonly destitute of an epipod; sometimes

90 RATHBUN

it is furnished with a rudimentary epipod, or with a fully developed one.
The last pair of pereiopods reach the end of the antennal scale.
The fourth abdominal segment is provided with a spine on either side

Fic. 38. Spivontocaris suckleyi. & (X 23). Station 3334. @. Side. 4. Dorsal
view of anterior portion.

below. The sixth segment is a little shorter and broader than in S.
townsend.

Male.—As in S. townsendi, the male resembles the female, except for
being smaller, more slender, and with longer antennules.

SS. suckleyt differs from S. gaimardii in the fewer dorsal spines, 5 or 6
against 7 or 8 in S. gaimardii,; in the larger eyes; in the antennular
peduncle of the female reaching middle of antennal scale; in the absence
of hook or hump on the third abdominal segment in the male; in the
slightly shorter sixth segment; and in the absence usually of an epipod
from the second pereiopod.

Variations.—In about three hundred specimens examined only one
lacks the spines on the fourth abdominal segment; in another specimen
the spine is present on one side, absent on the other.

Dimensions.—Length of @ (station 2864) approximately 79 mm.,
length of carapace and rostrum 28.7 mm., of rostrum 16 mm.

Distribution.—From the Arctic coast of Alaska southward to Wash-
ington; 6 to 165 fathoms.

Taken by the Albatross at the following stations:

Off the Pribilof Islands, 20-65 fathoms, stations 3438, 3439, 3483, 3494;

3496, 3504, 3535, 3538, 3552, 3560, 3561, 3611, 3637.
Bristol Bay, 19 fathoms, station 3237.

DECAPODS gI

Off Bristol Bay, 33 fathoms, stations 3303, 3306.
North of Alaska Peninsula, 26-47 fathoms, stations 3278, 3285, 3292,

3297-
Off Unimak Island, 24-43 fathoms, stations 3259, 3262, 3265, 3266.
Off Akutan Island, 72 fathoms, station 2842.
Unalaska, 19-165 fathoms, stations 3310, 3317, 3319, 3322) 3333-3330:
Shumagins, 21-69 fathoms, stations 2847, 2849-2851.
Davidson Bank, 43 fathoms, station 3215.
Off Sitkalidak Island, 60 fathoms, station 2854.
Off Grays Harbor, Washington, 50 fathoms, station 3047.
Strait of Fuca, 48-152 fathoms, stations 2864, 3443, 3445, 3446, 3451,
3454, 3460-3462, 3596.

Cape Smyth, Alaska (Point Barrow Expedition).

Captains Harbor, Unalaska, 80 fathoms (W. H. Dall).

Bay of Islands, Adak, 9-16 fathoms (W. H. Dall).

Lituya Bay, 6-9 fathoms (W. H. Dall).

Berg Bay, Glacier Bay, and Juneau, 20 fathoms (Harriman Expedition).
Puget Sound (Stimpson); Bare Island, British Columbia (Lenz).

SPIRONTOCARIS MOSERI Rathbun.

Spirontocaris gaimardii RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. III, 556, 1899 (part).
Spirontocaris moseri RATHBUN, Proc. U. S. Nat. Mus., XXIV, 897, 1902.

Allied to S. gaimardit.

Female.—The rostrum may be about as long as, or slightly exceed, the
carapace; it may reach the end of the antennal scale or fall short of
that point ; it is gently
ascending and armed
with 6 to 8 teethabove
(2 on carapace) and
4 to 7 below; upper
limb very narrow;
lower limb very broad
in front of the eye, tapering rapidly to the tip, which is
abruptly acute. The antennal spine is strong, the ptery-
gostomian small but well marked.

Eyes large, pyriform. ‘The antennular peduncle reaches
to the middle or to the distal third of the antennal scale;
the second segment is a little longer than the third; the
thickened portion of its outer flagellum overreaches the
scale; the spine of the antennular acicle reaches to the middle of the
second segment or about to the end of the first segment. The antennal
peduncle is very stout and nearly as long as the antennular; the scale is
narrow-ovate, its outer margin about five sevenths as long as the cara-

Fic. 39. Sfirontocaris moseri. & (X 2).

Station 3480.

92 RATHBUN

pace; the flagellum may equal the length of the body. The maxil-
lipeds exceed the scale by one fourth or more of the length of the last
segment. The first pair of feet extended overreach the antennular
peduncle; the second pair reach the tip of the maxillipeds; the fifth pair
extend to or beyond the end of the antennal scale. The dactyli of the
third, fourth, and fifth pairs are short and very stout, and armed with
strong spines.

The third abdominal segment in profile is smoothly rounded; poste-
riorly it is strongly produced over the fourth; the fourth has a spine on
either side; the sixth is twice as long as the fifth; the seventh is longer
than the sixth and bears 4 or 5 spines on either side.

Male.—The males are smaller and their antennulz a little longer than
in the females.

Dimensions. —Female, length 57 mm., length of carapace and rostrum
22.7 mm., of rostrum 11.5 mm.

Distribution. —Bering Sea southward and eastward to Washington,
60 to 516 fathoms, at the following stations of the Albatross :

Off Pribilof Islands, 81 and 276 fathoms, stations 3602, 3608.

North of Rat Islands, 270 fathoms, station 3785.

Off Seguam, Aleutians, 283 fathoms, station 3480 (type locality).

North of Unalaska, 309-406 fathoms, stations 3316, 3330-3332.

Off North Head, Akutan Island, 72 fathoms, station 2842.

Off Trinity Islands, 159 fathoms, station 2853.

Off Sitkalidak Island, 60 fathoms, station 2854.

Off Queen Charlotte Sound, British Columbia, 204 fathoms, station 2861.
Off Destruction Island, Washington, 516 fathoms, station 3343.

Variations.—Specimens from the two southernmost localities show a
tendency to differ from the types; the lower limb of the rostrum is nar-
rower, the rostrum shorter and more ascending, the antennular peduncle
longer and its basal scale shorter than in the types. A larger series
might serve to differentiate the southern from the northern form, but at
present they are combined.

SPIRONTOCARIS MAXILLIPES Rathbun.

Spirontocaris maxillipes RATHBUN, Proc. U.S. Nat. Mus., XxIv, 898, 1902.

Very near S. moseri, but distinguished as follows: The rostrum is
shorter, only about four fifths as long as the carapace, and has usually
fewer teeth, 5 to 8above, 2 to6 below. The antennular peduncle reaches
two thirds or more of the length of the antennal scale; the second seg-
ment is a little longer than in .S. moseri. The thickened portion of the
outer flagellum overreaches the scale by about half its length. The spine

of the antennular acicle scarcely reaches the middle of the second seg-

DECAPODS 93

ment, or may not extend beyond the first segment. The antennal peduncle
does not reach beyond the second segment of the antennular; the scale
is shorter, about two
thirds the length of the
carapace ; more than one
half of the last segment
of the outer maxilliped
extends beyond the scale.
The feet are without epi-
pods; the first pair reach
the end of the scale or
extend beyond it.

The spine of the

fourth abdominal somite Fic. 40. Spzrontocaris maxillipes. %. Station 3480. a. Side (X 2).
is reduced to an almost 4, Dorsal view of anterior portion (X 2). c. Chela of first pair (x 4).

imperceptible spinule; the sixth somite is shorter than in \S. moseri, being
only one and a half times the fifth; the side spinules of the telson are
commonly 3 in number, occasionally 4.

In this, as well as in the preceding species, the fingers of the first chelz
are a little over half as long as the palm.

Dimensions.— Female, length 48.9 mm., length of carapace and ros-
trum 18.4 mm., of rostrum 8.6 mm.

Distribution.— Aleutian Islands, 283 to 625 fathoms, at the following
Albatross stations:

Off Seguam, 283 fathoms, station 3480 (type locality).
North of Unalaska, 351 and 350 fathoms, stations 3330, 3331.
Off Shumagin Bank, 625 fathoms, station 3338.

SPIRONTOCARIS BRACHYDACTYLA Rathbun.
Spirontocaris brachydactyla RATHBUN, Proc. U. S. Nat. Mus., xxIv, 898,
1902.

This species is closely allied to S. moseri and S. maxillifes, but is dis-
tinguished by the short fingers of the first chelipeds, which are not more
than a third as long as the palm.

There are two specimens in the collection. The rostrum is distinctly
shorter than the carapace, and, in the one specimen in which it is perfect,
is armed with 6 teeth above (one on the carapace) and 3 below. No
pterygostomian spine. Antennular peduncles reaching a little over half
the length of the antennal scale, second segment very little longer than
third; antennular scale reaching slightly beyond the first segment in the
larger specimen, shorter than first segment in the smaller specimen; thick-

94 RATHBUN

ened portion of outer flagellum extending nearly half its length beyond
the scale. Antennal peduncle just as long as antennular. Half the last
segment of the maxilliped lies be-
yond the scale. The first pair of
feet just reach the tip of the scale ;
the fingers are unusually short,
as above mentioned. No spine
on fourth segment of abdomen;
sixth segment twice as long as
fifth; telson broken.

Dimensions.—Length of ovig-
erous female, exclusive of ros-
trum and telson, 24.8 mm., length of carapace 7.7 mm.

Distribution. —Southern California, deep water. The type, an ovig-
erous female, was dredged off Santa Cruz Island, in 266 fathoms (A/ba-
tross station 2948). A much smaller female comes from off San Diego,
417 fathoms (A/datross station 2928).

Fic. 41. Spirontocaris brachy-
dactyla. @. Station 2928. a@. Side
(X 3). &. Chela of first pair (Xx 5).

SPIRONTOCARIS CAMTSCHATICA (Stimpson).

Hippolyte camtschatica STIMPSON, Proc. Acad. Nat. Sci. Phila., XII, p. 33,
[102], 1860.

Belongs to the gaimardii group.

Female.—Rostrum a little longer than the carapace, shorter than
antennal scale, nearly straight (or slightly concave above), horizontal,
armed with 4 or 5 teeth above, the ante-
rior tooth at about the distal third, 1 or
2 teeth on carapace; upper limb very
narrow, disappearing anteriorly; lower
limb deepest a little in front of the eye, : :

Ms : i 1G. 42. Spirontocaris camtschatica. &.
and tapering distally, armed with 4 to 6 — Side of carapace (x 2). Petropavlovsk.
teeth; extremity acute. A strong antennal, a very small pterygostomian
spine.

The antennular peduncle reaches about two fifths the length of the
antennal scale; second and third segments very short, the second a little
the longer; the thickened portion of the outer flagellum extends to the
terminal third of the antennal scale; the inner flagellum reaches to the
end of the scale. The spine at the outer base of the antennules reaches
to, or nearly to, the distal margin of the second segment. The outer
margin of the antennal scale measures nearly as long as, just as long as,
or a little longer than, the carapace; its spine falls considerably short of
the laminar portion; the peduncle extends to the end of the second

DECAPODS 95

segment of the antennular peduncle.. The outer maxillipeds reach a little
beyond the middle of the antennal scale; the first pair of feet to the end
of the antennal peduncle; the second pair midway between the maxilli-
peds and the tip of the scale; the fifth pair to the end of the maxillipeds,

The third segment of the abdomen seen in profile is evenly rounded ;
it is laterally slightly compressed; fourth segment without spines; the
sixth is less than twice as long as the fifth, and also shorter than the
seventh, which has 4 or 5 spines on either side.

Male.—Smaller and more slender. Rostrum longer, sometimes ex-
ceeding the scale a trifle, curving upward in terminal half, armed with 5
to 7 teeth above, 5 to 9 below. The pterygostomian spine may be
obsolete. The thickened portion of the outer flagellum extends to the
terminal fifth of the antennal scale.

Dimensions. — é, approximate length 50 mm., length of carapace and
rostrum 18.5 mm., of rostrum 10.2mm. @, approximate length 34 mm.,
of carapace and rostrum 16 mm., of rostrum 9.5 mm.

Distribution.—From Arctic Alaska southward to Kamchatka and
Aleutian Islands, and eastward to Port Etches.

Taken by Dr. Dall in small numbers at Cape Lisburne, 5—7 fathoms ;
Cape Etolin, Nunivak, 8 fathoms; Hagemeister Strait, 8-15 fathoms;
Chichagof Harbor, Attu, 5~7 fathoms; Kiska Harbor, g-12 fathoms;
Constantine Harbor, Amchitka, 6-10 fathoms; Bay of Islands, Adak,
9-16 fathoms; off station reef, Iliuliuk Harbor, Unalaska, 3 fathoms;
Belkofski Bay, 15-25 fathoms; Anchorage, Big Koniuji Island, Shu-
magins, 6-20 fathoms; Shahafka Cove, Kadiak; Chugachik Bay,
Cook Inlet, 20-60 fathoms; Port Etches, 5-8 fathoms.

Bristol Bay, 1434 fathoms (A/éarross station 3236).

Kadiak, 5 fathoms (Harriman Expedition, W. R. Coe).

Bering Island (L. Stejneger).

Bering Island, in stomach of Gadus macrocephalus (N. Grebnitzky).

Rakovaya Bay, Avacha Bay, Kamchatka (L. Stejneger).

Solovarennaja Bay, Petropavlovsk, Kamchatka, 10-12 fathoms, alge
(N. Grebnitzky).

SPIRONTOCARIS KINCAIDI Rathbun.

Spirontocaris kincaidi RATHBUN, Proc. U.S. Nat. Mus., XxXIv, 899, 1902.

Near S. camtschatica.

Female.—Carapace short in relation to the abdomen. Rostrum one
third longer than the carapace, not reaching tip of antennal scale,
slightly concave above, horizontal, armed above with 5 (occasionally 6)
teeth, two of which are on the carapace, the anterior tooth a little before
the middle of the rostrum; tip acute; lower limb, as in all the gaimardiz

96 RATHBUN

group, tapering from a point in front of the eye to the tip, armed with
s teeth, one subterminal. Antennal spine strong, pterygostomian very
small.

Antennular peduncle reaching about two fifths the length of the
antennal scale, second and third segments very short and subequal, outer
scale reaching distal
margin of second seg-
ment, outer flagellum
falling short of the tip
of antennal scale, inner
flagellum exceeding it.
Antennal peduncle al-
most as long as anten-
nular; scale tapering, measured on outer margin exceeding
the carapace; its spine long, but still not so advanced as
the blade. Outer maxillipeds not quite reaching tip of ros-
trum; first pair of feet to middle of antennal scale; second pair ex-
tending just beyond maxilliped.

The posterior margin of the third abdominal segment is produced
backward in a strong lobe; the abdomen is strongly bent at this point;
the fourth segment is spineless; the sixth is less than twice as long as
the fifth, and a little shorter than the telson, which is armed with four
pairs of lateral spines.

Dimensions.—Ovigerous @, length (approximate) 36.5 mm., length
of carapace and rostrum 13.6 mm., of rostrum 7.7 mm.

Distribution.—From Washington to Santa Cruz, California:

Puget Sound (T. Kincaid).

Off Flattery Rocks, Washington, 40 fathoms (Addatross station 2873),1 2.

Off Santa Cruz, California, 21 fathoms (A/éatross station 3124), 1 Q (type).

Strait of Fuca, 37 fathoms (A/batross station 3593), 1 egg-bearing ¢,
placed here with doubt. The rostrum and telson are broken, the

antennule are a little longer than in the type, their peduncles reach
nearly to the middle of the acicle.

Fic. 43. Spivontocaris kincaidt. @ (X 23).
Station 3124.

SPIRONTOCARIS LAYI (Owen).

Hippolite Layt OWEN, in Zoology of Captain Beechey’s Voyage [in the Béos-
som], Crustacea, p. 90, pl. XXVII, fig. 3, 1839.

Distribution.— Monterey, California (Owen) ; Esquimalt Harbor,
Vancouver Island (Bate).

Owen’s description is as follows:

“Hip. rostro acuminato, supra multi-serrato, ante medium subtus
quadri-serrato. Long. corp. unc. 2%. Color ruber.” According to

DECAPODS 97

Owen, the rostrum is longer and narrower than in .S. afinis, extending to
the extremity of the superior antennze; the dental formula was the
same, 4°, in the 3 specimens preserved. According to Owen’s figure,
the antennular peduncle extends about to the middle of the rostrum, the
second and third segments are subequal, the thickened portion of the
outer flagellum is shorter than the peduncle. The rostrum curves strongly
upward in its anterior half.

SPIRONTOCARIS BIUNGUIS Rathbun.
Spirontocaris biunguis RATHBUN, Proc. U. S. Nat. Mus., XXIV, 899, 1902.

This species of the gaimardii group is an inhabitant of deeper water,
and is distinguished by its larger eyes, longer rostrum, subchelate dactyli,
and by having an exopod and an epipod on the outer maxilliped, while
the pereiopods are destitute of epipods.

Carapace carinated on its anterior half or two thirds. Rostrum from one
and a fourth to one and two thirds times as long as remainder of cara-
pace, its poste-
rior half or third
gradually as- ee ee
cending, the re- ~< -
mainder more See
abruptly $0; WD- _mesemeersSS=EOER) A
per limb narrow, = aa
the widest point eee
in. front of the Fic. 44. Sfpirontocaris biunguis. Station 3324. @. Side.
eyes lower limb 3 ciyits of ous of age 3 pairs oflege (Xm),
a little wider than
the upper; terminal third a slender sharp spine, not limbed ;
dorsal spines 5 to 7 (two on carapace), ventral spines 4 to 7.
Antennal spine strong; pterygostomian spine or tooth very small.

Eyes very large and oblique, pyriform, the cornea covering the greater
part of the peduncle. Antennular peduncle extending a little past the
middle of the antennal scale; second joint three times as long as third;
outer scale reaching to end of first segment; thickened portion of outer
flagellum extending in the female one third, in the male two thirds of its
length beyond antennal scale. This last is four fifths as long as carapace ;
antennal peduncle reaching end of second segment of antennular
peduncle; flagellum longer than the body.

The tips of the maxillipeds are half way between the antennal peduncle
and the tip of the scale. The first pair of feet attain the middle of the
last joint of the antennal peduncle; the palm is less than twice as long as

98 RATHBUN

the fingers. The second pair exceed the acicle. The last 3 pairs are very
slender and about the same length, the tip of the last pair reaching the
terminal third of the rostrum; the extremity of the propodi is fringed with
bristles ; the dactyli are slender; the proximal half of their concave mar-
gin is furnished with 5 or 6 spinules so closely appressed that to the
naked eye the dactylus appears unarmed; the distal spinule is long,
and with the slender tip of the dactylus gives the appearance of a minute
chela.

The abdomen is non-carinated; the fourth segment is unarmed; the
sixth segment is one and a half times the fifth; the seventh is a little
longer than the sixth and armed with 5 or 6 spinules on each side.

Dimensions.—Length of egg-laden 9 83 mm., length of carapace and
rostrum 37.6 mm., of rostrum 23.2 mm.

Distribution.—¥rom Bering Sea (off Pribilof Islands) to Oregon, 109—
987 fathoms.

Dredged by the Albatross :

Off Pribilof Islands, 184-688 fathoms, stations 3489, 3501, 3502, 3608.

North of Aleutian Islands (178° 31/ 00” E.), 850 fathoms, station 3784.

North of Rat Islands, 270 fathoms, station 3785.

North of Unalaska, 225-987 fathoms, stations 3227, 3316, 3329-3332,
3607.

West of Unalaska, 576 fathoms, station 3326.

Chernofski Harbor, Unalaska, 109-284 fathoms, stations 3324, 3325.

Off Shumagin Bank, 625 fathoms, station 3338.

Southeast of Chirikof Island, Alaska, 695 fathoms, station 3340.

Off Cape St. James, Queen Charlotte Islands, British Columbia, 876
fathoms, station 2860 (type locality).

Off Sea Lion Rock, Washington, 636-877 fathoms, stations 3069-3071,
3°74, 3975.

Off Cape Elizabeth, Washington, 831 fathoms, station 3344.

Off Grays Harbor, Washington, 559 fathoms, station 2871.

Off Tillamook Bay, Oregon, 786 fathoms, station 3346.

SPIRONTOCARIS PALPATOR (Owen).

Hippolyte palpator OWEN, in Zoology of Captain Beechey’s Voyage [in the
Blossom), Crustacea, p. 89, pl. XXVIII, fig. 3, 1839.

Heptacarpus palpator HOLMES, Occas. Papers Calif. Acad. Sci., vil, 196,
pl. 111, figs. 48, 49, 1900, and synonymy.

Distribution.—San Francisco Bay, California (Stimpson), to Magda-
lena Bay, Lower California (Lockington). Monterey (Owen). The
specimen from Kadiak referred by Brandt to this species is probably S.
brevirostris, as stated by Dr. Holmes.

Specimens in the National Museum are from Pacific Grove, California
(John C. Brown); San Diego (H. Hemphill, C. R. Orcutt); San Diego

DECAPODS 99

County (C. R. Orcutt); San Diego Bay, 5-634 fathoms (Addatross sta-
tions 3575-3578, 3582).

SPIRONTOCARIS BREVIROSTRIS (Dana).

Hippolyte brevirostris DANA, Proc. Acad. Nat. Sci. Phila., 1852, 24; Crust.
U. S. Expl. Exped., 1, 566, 1852, pl. XXXVI, fig. 5, 1855.—STIMPSON,
Jour. Boston Soc. Nat. Hist., VI, 500, 1857.

Spirontocaris brevirostris WWALKER, Trans. Liverpool Biol. Soc., x11, 276,
1898.

Heptacarpus brevirostris HOLMES, Occas. Papers Calif. Acad. Sci., v1, 198,
pl. Ill, figs. 50, 51, 1900, and synonymy.

Distribution.— Attu, Aleutian Islands, to San Francisco Bay, California.

Nine localities between Attu and Sitka, low water to 25 fathoms (W. H.
Dall).

Unalaska (S. Applegate).

Aleutian Islands (L. M. Turner).

Kadiak (W. J. Fisher).

Tongass, Alaska (H. E. Nichols).

Vakutat; Kadiak, 5 fathoms; Sitka (Harriman Expedition).

Otter Bay, Pender Island, Seymore Narrows, British Columbia; Sucia
Island, British Columbia; Port Angeles, Washington (A/datross).

Fort Rupert, British Columbia (Harlan I. Smith).

Strait of Fuca.

Puget Sound (T. Kincaid).

Cape Mendocino, California (University of California).

San Francisco Bay (H. Hemphill).

Previously recorded from Kadiak (Brandt, as pa/pfafor); Vancouver

Island (Smith); Strait of Fuca, near Dungeness (Dana); Humboldt
County, California (Holmes); San Francisco Bay (Stimpson, Holmes).

SPIRONTOCARIS PUSIOLA (Kroyer).

Hippolyte pusiola KROYER, Naturh. Tidssk., 11, 576, 1841; K. Danske
Vidensk. Selsk. Afhand., Ix, 319, pl. III, figs. 69-73, 1842.—SMITH,
Trans. Conn. Acad. Arts Sci., v, 77, pl. 1X, figs. 4-7 (telson), 1879.

Distribution.—Bering Island, Aleutian Islands, and Alaska Peninsula
eastward to Chichagof Island, 5-159 fathoms. Gulf of St. Lawrence to

Long Island Sound, low water to 105 fathoms. Greenland? Iceland.
Northern Europe.

Collected by the Albatross :

Off North Head, Akutan Island, 56-72 fathoms, stations 2841, 2842.
Davidson Bank, 42 fathoms, station 2845.

Northeast of Amak Island, 19 fathoms, station 3274.

North of Bird Island, Shumagins, 21 fathoms, station 2850.

Off Trinity Islands, 159 fathoms, station 2853.

100 RATHBUN

By W. H. Dall at:

Port Levashef, Unalaska, 20-30 fathoms.

Tliuliuk Harbor, Unalaska, 10 fathoms.

Captains Harbor, Unalaska, outside of ridge, 25-75 fathoms.
Off Round Island, Coal Harbor, Unga, 8-9 fathoms.
Anchorage, Big Koniuji Island, Shumagins, 6-20 fathoms.
Popof Strait, near reef, 6 fathoms.

Shahafka Cove, Kadiak, 12-14 fathoms.

Chugachik Bay, Cook Inlet, 20-60 fathoms.

Port Etches, 5-8 fathoms.

Granite Cove, Port Althorp, Chichagof Island.

Bering Island (L. Stejneger).
Bering Island, among Laminaria (N. Grebnitzky).

The males grow to be as large and robust as the females; they have
the antennular peduncle and outer maxilliped much longer than in the
female; in old males the peduncle is about four fifths as long as the an-
tennal scale, first segment extending half its length beyond the eyes, outer
basal scale only two thirds as long as first segment, second segment twice
as long as third. Antennal scale, measured on outer margin, two thirds
as long as carapace; peduncle overreaching slightly the first segment of
the antennular peduncle. The penult segment of the outer maxilliped
slightly overreaches the antennal peduncle; the terminal segment is very
long, as long as the carapace.

Dimensions.— A male measures 25 mm. long, carapace and rostrum
8.3 mm., rostrum 2.5 mm.

SPIRONTOCARIS HERDMANI Walker.

Spirontocaris herdmanit WALKER, Trans, Liverpool Biol. Soc., XII, 277, pl.
XVI, fig. 2, 1898.

Small. Carapace high. Rostrum two thirds length of carapace
proper, reaching nearly or quite to end of, or a little beyond, first antennu-
lar segment, straight, deflexed; upper margin armed with 3 to 6 teeth, one
of which is on the carapace, anterior tooth remote from the tip, which
is pointed; lower margin with one or two teeth near the tip. Antennal
and small pterygostomian spine present. Antennular peduncle reaching
a little past middle of antennal scale; second and third joints very short,
the second a little the longer; basal scale reaching to end of first seg-
ment; thickened part of outer flagellum extending beyond acicle.
Acicle nearly as long as carapace; antennal peduncle nearly as long as
antennular. Outer maxilliped extending beyond scale by half length of
last segment. First pair of feet exceeding scale a little. Fourth seg-
ment of abdomen provided with a small lateral spine; sixth segment short,
very little longer than wide; telson with three spinules on each side.

DECAPODS IOI

Walker says of this species that “the rostrum reaches a little beyond
the end of the peduncle of the inner antennz,” but this may be an error,
as in the figure the rostrum is represented not much longer than the eye.

Dimensions.— Length of female with ova 30 mm. (Walker). Length
of female with ova 16.2 mm., length of carapace and rostrum 5.2 mm.,
of rostrum 2.1 mm.

Distribution.—Sitka, Alaska, 10 fathoms, June 15, one @ with ova
(Harriman Expedition). Puget Sound, 1895, 3 specimens (T. Kincaid),
Puget Sound (Walker).

SPIRONTOCARIS TAYLORI (Stimpson).

fippolyte taylori STIMPSON, Jour. Boston Soc. Nat. Hist., v1, 500, 1857.
Heptacarpus Taylori HOLMES, Occas. Papers Calif. Acad. Sci., VII, 199, pl.
Ill, figs. 52, 53, 1900, and synonymy.

Distribution.—San Francisco Bay, California, to Magdalena Bay,
Lower California. Specimens in the National Museum from Monterey
(D. S. Jordan, 1880, and J. O. Snyder, 1895), San Pedro (Adbatross),
Point Loma (A/datross), San Diego (Rosa Smith).

One of the three females collected by Dr. Jordan has an abnormal
rostrum, longer and more slender than in typical specimens, still not
reaching the cornea; in other respects the individual agrees with its
associates.

SPIRONTOCARIS PICTA (Stimpson).
Hippolyte picta STIMPSON, Ann. Lyc. Nat. Hist. N. Y., X, 125, 1871.
Fleptacarpus pictus HOLMES, Occas. Papers Calif. Acad. Sci., v1, 200, pl. IIT,
figs. 54 and 55, 1900, and synonymy.

Distribution.— Monterey Bay to San Diego, California. Specimens are
in the National Museum from Monterey Bay (A/batross), Pacific Grove
(H. Heath, John C. Brown), Catalina Harbor (W. H. Dall), Lajolla
(Albatross), San Diego (various collectors).

SPIRONTOCARIS PALUDICOLA (Holmes).

Heptacarpus paludicola HOLMES, Occas. Papers Calif. Acad. Sci., VII, 201,
ple 111, figs, 56; 57, 1900.

Distribution.—Humboldt Bay (Holmes) to San Diego, California.
Specimens are in the National Museum from San Diego (A/¢atross), and
from either Monterey or Catalina Harbor (Dall).

Since this paper went to press, a Spzrontocaris from Pearson, Wash-
ington, has been received, through R. W. Doane, which seems to be un-
described. It has the coloration of S. Aicfa, a rostrum similar to that of
S. paludicola, while the epipods are as in S. sétchensis,

102 RATHBUN

SPIRONTOCARIS SITCHENSIS (Brandt).

Hippolyte sitchensis BRANDT, Middendorff’s Reise Sibir., 11, Zool., Theil 1, 116,
pl. v, fig. 18, 1851.—?? STIMPSON, Jour. Boston Soc. Nat. Hist., VI,

499, 1857 (Monterey).
2? Hippolyte sitchensis STIMPSON, Ann. Lyc. Nat. Hist. N. Y., X, 125, 1871.

This species resembles most .S. pa/udicola of California. It differs,
however, in its shorter rostrum, which is two thirds or four fifths the
length of the carapace; in the shorter antennular scale, only slightly
exceeding the first segment of the peduncle; in the shape of the antennal
scale, which is broadly arcuate at the end, the blade exceeding the spine ;
in lacking an epipod on the second pereiopod.

Distribution.—¥rom Sitka, Alaska, southward to Puget Sound:

Sitka (Brandt); Cape Fox, 39 (W. R. Coe, Harriman Expedition) ;
Tongass, July 16, 1885, 3 9 (Lieutenant H. E. Nichols, U.S. N.); Ref-
uge Cove, Port Chatham, 1 9 (W. H. Dall); Puget Sound (Calman).

SPIRONTOCARIS CRISTATA (Stimpson).

Hippolyte cristata STIMPSON, Proc. Acad. Nat. Sci. Phila., 1860, 33 (not
HT, cristatus De Haan, 1849). 1

Spirontocaris cristata WALKER, Trans. Liverpool Biol. Soc., XII, 277, 1898.
Heptacarpus cristatus HOLMES, Occas. Papers Calif. Acad. Sci., VII, 202,
pl. 111, figs. 58, 59, 1900.

Distribution.—¥rom Sitka, Alaska, to San Diego, California:

Sitka, 10 fathoms (Harriman Expedition); Puget Sound (Walker, also
collected by T. Kincaid).

San Pablo Bay, taken in Chinese shrimp-nets (4/batross).

Monterey Bay, 33 fathoms (4/¢atross station 3132); Monterey (J. O.
Snyder).

San Diego Bay 3-634 fathoms (A/tarross stations 3563, 3567, 3575-
3578).

San Diego, 10 fathoms (H. Hemphill); south of San Diego Bay, 22
fathoms (A/éatross station 3679).

Southern California (W. H. Dall).

I have a little doubt about the single speci-
<I) men from Sitka, as it is much mutilated; it is
efi not, however, referable to any other species.

os ae The male differs peculiarly from the female.

; It has the customary points of difference: It is

Ob iG gh gfeerontocaris crate smaller, more slender, the rostrum is more slen-

aed a der and slightly overlaps the second antennular

1] have considered that De Haan’s use of this name on plate xLv did not invalidate

it for Stimpson’s species, because De Haan in his Errata (p. 244), as well as in the

explanatory text (p. 194), corrected the name of his species to Sicyonia cristata, the
correction appearing, so far as I am able to ascertain, simultaneously with the plate.

DECAPODS 103

segment instead of reaching to the end of it; but the most striking char-
acter is the unusual length of the fingers, which exceed the palm in
length and are rather slender, giving the chele a Pulemon-like appear-
ance; in the 9, on the contrary, the fingers are considerably shorter
than the palm, as in most species of the genus.

SPIRONTOCARIS STONEYI Rathbun.

Spirontocaris stoneyi RATHBUN, Proc. U. S. Nat. Mus., XXIV, 899, 1902.

Near 5S. cristata; a small, slender species. Rostrum about half as
long as the carapace and reaching a little beyond first segment of anten-
nular peduncle ; upper margin
convex and armed with 7 to
g teeth (1 or 2 of which are on
carapace), except near the tip,
where it is straight and un-
armed; tip pointed; midrib
nearly straight, lower margin
with a tooth near the tip. A strong antennal, a short but
slender pterygostomian, spine.

Antennular peduncle reaching to middle of antennal scale;
second segment one and a half times as long as third; outer
scale not reaching end of first segment; thickened portion
of outer flagellum reaching about to end of antennal scale.
Scale as long as carapace; spine reaching as far as blade;
peduncle reaching a little beyond first segment of anten-
nular peduncle. Maxillipeds very slender, reaching to the
distal third of the antennal scale; the first pair of feet to a little beyond
the antennal peduncle, the palm widening distally and only a little
longer than the fingers; second pair of feet about reaching tip of scale.

Fourth abdominal somite unarmed; sixth long, over 1% times as long
as fifth, and as long as telson, which has 3 pairs of side spines.

Dimensions.— Female, length 20 mm., length of carapace and rostrum
5.5 mm., of rostrum 1.8 mm.

Type locality. —Bering Sea, lat. 629 15’ 00” N., long. 167° 48/ 00” W.,
20% fathoms (Lieutenant George M. Stoney, U.S. N., June, 1884).

Fic. 46. Spirontocaris stoneyé
(xX 5). Bering Sea.

SPIRONTOCARIS AVINA Rathbun.

Spirontocaris avina RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, III, 557, 1899.

Carapace with anterior half carinated; rostrum half or less than half
the length of the carapace, reaching to end of first antennular segment ;

104 RATHBUN

strongly arched over the eye, and armed with 12 to 14 small teeth; the
posterior 3 or 4 behind the orbit, the posterior 2 distinctly separated from
the others; the hindermost is much depressed, obsolescent; the distal
third of the rostrum is
straight, either horizon-
tal or slightly deflexed,
acute, unarmed above,
furnished with 1 or 2
teeth below; lower limb
very narrow. Antennal
spine well developed,
pterygostomian very
small.

Eyes large, pyriform.

Fic. 47. ee avina. Station 2853. _a@. Side of carapace Antennular peduncle

xX 3). 4. Dorsal view of anterior portion (x 4). extending two thirds
the length of the antennal scale, second segment three times as long as
third, outer basal scale suberect and not reaching end of first segment;
flagella subequal in length; thickened part of outer one extending half
its length beyond antennal scale. Outer margin of antennal scale a
little more than two thirds as long as carapace, scale tapering, distal
margin very oblique, peduncle extending quite to middle of scale and
nearly to end of second segment of antennular peduncle.

The outer maxillipeds extend beyond the scale by nearly half the
length of their last segment. First pair of feet reaching half-way
between antennal peduncle and end of scale; second pair exceeding
maxilliped by the length of the propodus and the last joint of the carpus ;
third to fifth pairs nearly equal, the fifth pair overreaching the second
pair; dactyli very slender and unarmed.

Abdomen smooth; fourth segment furnished with a lateral spine;
sixth segment a little less than twice as long as fifth, and shorter than
seventh, which bears 3 spinules on each side.

Male.—Smaller, more slender, and has slightly longer antennule than
the female.

Dimensions. —Female, length 35 mm., length of carapace and rostrum
12 mm., of rostrum 3.5 mm.

Distribution.—From Pribilof Islands to Oregon, 41 to 351 fathoms.
Taken exclusively by the A/batross, at the following localities:

Off Pribilof Islands, 60-184 fathoms, stations 3484, 3485, 3489, 3494,
3605, 3609.

North of Unalaska, 165-351 fathoms, stations 3315, 3317, 333° (type
locality), 3331.

DECAPODS 105

Tiuliuk Harbor, Unalaska, 58-68 fathoms, stations 3310, 3313.

Off Akutan Island, 72-91 fathoms, stations 2842, 3548.

Northwest of Unimak Island, 81-85 fathoms, stations 3225, 3257.

Off Kudobin Islands, 41 fathoms, station 3279.

Between Unga and Nagai Islands, Shumagins, 110 fathoms, station
2848.

Off the Trinity Islands, 159 fathoms, station 2853.

Gulf of Georgia, British Columbia, 67 fathoms, station 2863.

Strait of Fuca, 53-100 fathoms, stations 3443, 3445, 3446, 3460, 3596,
3597-

Near Flattery Rocks, Washington, 171 fathoms, station 2866.

Off Heceta Bank, Oregon, 277 fathoms, station 28go.

SPIRONTOCARIS MACILENTA (Kréyer).

Hippolyte macilenta KROYER, Naturh. Tidssk., 111, 574, 1841; K. Danske
Vidensk. Selsk., Afh., IX, 305, pl. II, figs. 55, 56, 1842.—-SMITH, Trans.
Conn. Acad. Arts Sci., Vv, 71, 1879.

Spirontocaris macilenta RATHBUN, The Fur Seals and Fur-Seal Islands of
the North Pacific Ocean, Pt. III, 557, 1899.

Distribution.—Arctic and Atlantic coasts of North America, as far
south as Halifax, Nova Scotia; Bering Sea; Kamchatka; Okhotsk
Sea; 16 to 100 fathoms.

The following localities on the Alaskan and Asiatic coasts are repre-
sented in the Museum collection:

Arctic Ocean (U.S. R. S. Corwin).

Bering Strait (Dr. Robert White).

Lat. 66° 30! 25” to 66° 43’ oo” N., long. 167° 14! 10” to 168° 08! 15”
W., 19-30 fathoms (U. S. R. S. Corwin).

Bering Sea, between Matthew Island, Pribilof Islands, and Bristol Bay;
2914-39 fathoms (A/datross stations 3252, 3306, 3507, 3511, 3513;
3518, 3519).

Off Kamchatka, 39-42 fathoms (A/batross station 3781).

Avacha Bay, Kamchatka, 16 fathoms (A/datross station 3642).

Southeast coast of Kamchatka, 100 and 96 fathoms (A/¢atross stations

3643, 3644).
Off Robben Island, Okhotsk Sea, 25 fathoms (4/éatross station 3649).

SPIRONTOCARIS MACROPHTHALMA Rathbun.
Spirontocaris macrophthalna RATHBUN, Proc, U. S. Nat. Mus., XxIVv,
goo, 1902.

Allied to S. macilenta and S. bispinosa.

Female, — Carapace carinated in its anterior half, posterior spine at the
anterior fifth; upper line of carapace and midrib of rostrum nearly hori-
zontal, the rostrum gently ascending in its anterior portion; rostrum
about three fourths or five sixths as long as the carapace, upper and
lower limbs with convex margins, and widest at about the middle of their

106 RATHBUN

length; upper margin armed with ro to 14 teeth, including 2 or 3 on the
carapace; lower margin with 1 to 3 spines; tip acute. A strong antennal

Fic. 48. Sptrontocaris macrophthalma. Station 3076. a. Side view, $ (X 2).
6, Dorsal view of anterior portion, @ (X 2%).

spine on the anterior margin, no supraorbital nor pterygostomian spine;
antero-lateral angle sharp.

Eyes large, pyriform; corneze extending on the inner side almost to
the base of the peduncle. Eyes reaching two thirds the length of the
first antennular segment; this segment is twice as long as the second,
and the second three times as long as the third; the second and third
are each provided with a terminal spine; the peduncle does not reach
the end of the rostrum; the thickened part of the outer flagellum exceeds
by half its length the antennal scale; inner flagellum half again as long
as outer; the basal scale does not quite reach the end of the first seg-
ment; it has a strong lobe on its proximal outer margin. The peduncle
of the antennz extends to the end of the second segment of the anten-
nulz and the middle of the scale; the flagellum is one and a third times
the length of the body. The blade of the scale exceeds the spine con-
siderably, and is most advanced toward its inner margin.

The outer maxillipeds are slender and reach midway between the end
of the antennal peduncle and the end of the scale. The first pair of feet
reach just beyond the base of the scale; the second pair just beyond the
end of the scale; of the carpal joints, the second is greater than the first,
the first plus the second equals the third, the fourth equals the seventh,
the fifth is greater than the sixth, the fifth plus the sixth exceeds a little
the seventh. The last three pairs of feet are long, fragile, and unarmed;

DECAPODS 107

the last pair may extend beyond the antennal scale by the length of the
dactylus and two thirds of the propodus.

The first four segments of the abdomen are unarmed; the third pro-
duced moderately backward in the middle; fifth with a spine at the
infero-posterior angles; sixth with a spine at the infero-posterior and
supero-posterior angles; sixth nearly twice as long as fifth; telson about
as long as sixth segment, with 5 or 6 pairs of lateral spinules; extremity
with a pair of long median spines, a longer pair of submedian spines, and
a pair of short outer spines.

Male.—The males exhibit the usual differences from the females in
being more slender, in the longer antennular flagella, and in the abdominal
appendages.

Dimensions.—Female, length 62.6 mm., length of carapace and
rostrum 24 mm., of rostrum 11 mm.

Distribution.— From the north coast of Unalaska eastward and south-
ward to Point Sur, California, 178 to 636 fathoms. Taken by the
Albatross at the following stations:

North of Unalaska, 399 and 350 fathoms, stations 3329, 3331.

Off Davidson Bank, 280 fathoms, station 3337.

South of Sannak Islands, 483 fathoms, station 3210.

Clarence Strait, 322 fathoms, station 3077.

Off Queen Charlotte Sound, British Columbia, 204 fathoms, station
2861.

Off Tahwhit Head, Washington, 178 fathoms, station 3076 (type locality).

Off Destruction Island, Washington, 516 fathoms, station 3343.

Off Sea Lion Rock, Washington, 636 and 477 fathoms, stations 3070,

3°73:
Off Cascade Head, Oregon, 345 fathoms, station 3347.
Off Heceta Bank, Oregon, 277 fathoms, station 2890.
Off Point Arena, California, 455 and 239 fathoms, stations 3348, 3349.
South of Farallone Islands, California, 391 and 217 fathoms, stations

3104, 3105.
Off Pigeon Point, 296 fathoms, station 3112.
Off Monterey Bay, 418-581 fathoms, stations 3126, 3127, 3670.
Monterey Bay, 278 fathoms, station 3669.
Off Point Sur, 298 fathoms, station 3187.

A ffinities.— Resembles the preceding species, S. macilenta, but differs
notably in the much larger and more reniform eyes, the longer rostrum,
of which the upper limb is narrowed at its base and the lower limb is
less deep, in the larger dorsal spines and the shorter scale. From S.
bispinosa at once distinguished by the absence of a supraorbital spine,
of the long, slender process of the rostrum and of the spine at the middle
of the carapace. The sixth segment of the abdomen is longer and more
slender than in S. dispinosa.

108 RATHBUN

Family ALPHEIDA,
Genus Alpheus Fabricius.

ALPHEUS DENTIPES Guérin.

Alpheus dentipes GUERIN, Exp. Scient. Morée, part. Zool., 39, pl. XXVII,
fig. 3, 1832.
Alpheus clamator HOLMES, Occas. Papers Calif. Acad. Sci., vil, 182, 1900,
and synonymy.
Distribution.—Farallone Islands, California, to San Bartolomé Bay,
Lower California. Mediterranean; Cape Verde Islands; Bermudas;

Porto Rico; Cuba; Key West.

ALPHEUS BELLIMANUS Lockington.
Distribution,—From Monterey to San Diego, California. Chile (Cou-
tiére).
ALPHEUS BARBARA Lockington.
fTabitat,—Santa Barbara, California.
According to Coutiére, probably the same as 4. macrocheles (Hail-

stone).
ALPHEUS CALIFORNIENSIS Holmes.

Habitat.—San Pedro, California.

ALPHEUS EQUIDACTYLUS Lockington.

Distribution.— Monterey to Santa Barbara, California.
According to Coutiére, this is Apheopsis trispinosus (Stimpson).

Genus Betzus Dana.

BETAZUS HARFORDI Kingsley.
Distribution.—Point Arena to Catalina Island, California.

BETAUS LONGIDACTYLUS Lockington.
Distribution.—San Pedro to San Diego, California.

BETAUS HARRIMANI sp. nov.

female, with eggs.—Carapace compressed, upper line nearly hori-
zontal; frontal margin nearly straight, with a broad, shallow, almost im-
perceptible median sinus, and a deeper sinus just outside the eye. Basal
scale of the antennules broad at base (the breadth of each equaling the
interspace), narrowing distally, and somewhat scythe-shaped, curving
slightly inward toward the extremity, which reaches the distal third of

DECAPODS 109

the second joint; second joint about one and a half times as long as the
third; inner flagellum about one fourth longer than the carapace; outer
flagellum about two thirds the length of the inner
and becoming much smaller at the middle, the distal
half very slender.

Antenne with a strong spine below the base of
the acicle; peduncle slightly exceeding that of the
antenna; scale with a broad terminal spine, which
reaches the end of the antennular peduncle, and ex-
ceeds the blade, from which it is separated by a short | =} <3 |S ¢
narrow slit; flagellum nearly twice the length of the |

cos F : ‘ IG. 49. Beteus har-
carapace. Maxillipeds reaching a little past the mid- vimanz.\" a. Dorsal view

of anterior portion (x 4).

dle of the terminal segment of the antennal peduncle. 4. Carpus and chela of sec-
. aa ond pair (X 53). c. Left

Chelipeds subequal, similar, of very moderate size; chela of first pair (x 22).
merus with rounded angles, the distal end widened, length about two and
a half times breadth, outer surface with a broad oblique sulcus, a trans-
verse groove at the supero-distal angle, behind which the upper margin
ends abruptly, but does not terminate in a spine; surface granulate,
lower surface granulo-spinulous. Carpus small, rounded. Hand narrow-
oblong, somewhat compressed, finely scabrous, margins rounded, the
length not more than one half greater than that of the preceding joints
combined. Fingers fitting close together, the pollex narrowly conical
and twice as wide at base as the dactylus, which is subcylindrical; the
tips are sharp and cross each other; the prehensile edges are subentire,
very finely denticulate, that of the pollex convex except at the tip.

The first joint of the carpus of the second pair of feet is about as long
as the three following combined; the third and fourth are equal, the
second a little longer than either; the fifth twice as long as the fourth,
and slightly shorter than the palm, which is subequal to the fingers.

The propodi of the remaining feet have a row of six or seven spines
beneath; the dactyli are over a third the length of the propodi and very
acuminate.

The postero-lateral angles of the fourth and fifth abdominal segments
are rounded, of the sixth blunt. The distal end of the peduncle of the
uropods bears a pair of spines; a spine near the outer angle of the
uropods.

Color.—Light green.

Dimensions.—Female, length of body to tip of telson 26.6 mm.;
length of carapace 10 mm.; length of cheliped 17.5 mm.; length of palm
6 mm.; of dactyl 4.6 mm.

Dype locality.—Sitka (W. R. Coe, Harriman Expedition); one female

IIO RATHBUN

with ova (U. S. National Museum, No. 25,692). This species represents
a more northern latitude than any Alpheid hitherto described.

Affinities. —The species differs from &. longidactylus Lockington in
its small hands, non-gaping fingers devoid of teeth, in the relative lengths
of the carpal segments of the second pair of legs, in the shorter antennular
spine, in the unequal peduncular joints of the antennule, as well as in
some minor respects. }

Family ZYSIZA 7/DZ.
Genus Processa Leach.
PROCESSA CANALICULATA Leach.

Processa canaliculata LEACH, Mal. Podoph. Brit., pl. XLI, and correspond-
ing text, July 1, 1815.—RATHBUN, Bull. U. S. Fish Comm. for 1900,
vol. Il, p. 104, 1901, and synonymy.

Two specimens of unusual interest were taken at San Diego, Califor-
nia, by D. S. Jordan, in 1880. They are about 22 mm. long, and differ
from typical specimens in having the left foot of the first pair similar to
the right, or chelate. One specimen is a female, and has both chelipeds
present. The other is so mutilated that the sex is indeterminable; it has
a left cheliped, the right is missing. This form might perhaps be deemed
a distinct species or genus were it not that among a lot of specimens
from Cedar Keys, Florida, both forms occur. From this locality they
are small (12 to 15 mm. long), and five specimens are bichelate, while
four have only a right cheliped, the left foot being simple, as in typical P.
canaliculata. These two forms from the same locality present no other
appreciable difference.

Aside from this remarkable dimorphism in the left first foot, the species
is a most variable one. The rostrum may be half as long or nearly as
long as the eye. The eyes, while always of good size, are not uniform, in
some cases larger and more reniform, with the cornea extending on the
outer side almost back to the carapace. The second joint of the anten-
nulz varies from one and a fourth to twice the length of the third joint.
The antennal scale may be a little more than half as long as the carapace
(rostrum excluded) or even two thirds as long as carapace; it may be just
as long as the antennular peduncle, or distinctly longer. Of the speci-
mens examined, those from the west coast of Mexico and Panama Bay
have the largest eyes; they agree fairly well with the description and
figure of Bate’s P. processa from Amboina, 15 fathoms.

1 The type of B. harrimani has been compared with Dr. Holmes’s description of 2B.

longidactylus (Occas. Papers Calif. Acad. Sci., VII, 190, 1900), specimens of that
species not being at hand.

DECAPODS III

To the localities recorded by me in 1901 (/oc. ci¢.) may be added the

following:

Cedar Keys, Florida, in seaweed, between tides, two forms as described
above (H. Hemphill).

Trinidad (Adbatross).

San Diego, California; feet of first pair chelate (D. S. Jordan).

Off Abreojos Point, Lower California, 48 fathoms (A/éa/ross station
2834).

Gulf of California, 29-71 fathoms (A/batross stations 2998, 3011, 3014).

Panama Bay, 5114 fathoms (4/atross station 2805).

Range.—Europe; Madeira; Bermudas; from North Carolina to
Trinidad, including Gulf of Mexico and West Indies; from San Diego,
California, to Panama Bay; Japan; Amboina. Shallow water to 111
fathoms.

Family CRANGONIDZ.
Genus Crangon Fabricius.
KEY TO THE SPECIES OF CRANGON.

A. Gastric region not depressed below the general level of the carapace.
B. Carapace with 1 or more median spines.
C. Carapace with 1 median spine.
D. Sixth segment of abdomen with 2 prominent longitudinal
Carine ey SS Ae, tet RY Ge Ge vik Mes SOA,
D’. Sixth segment of abdomen not carinated.
E. Sixth segment of abdomen sulcate beneath.

F. Manus of first pair of feet very slender, four or more
times as long as broad. Fifth abdominal segment
with a posterior spine at the supero-lateral angles.

G. Hand of male four and a half times as long as wide
Jranciscorum.
G’. Hand of male five and a half times as long as wide
Jranciscorum angustimana.
F’. Manus of first pair of feet not slender, less than three
and a half times as long as wide.
G. Fifth segment of abdomen not carinate.
H. Blade of antennal scale having its anterior margin
more advanced at its inner than its outer angle.
Sixth segment of abdomen with a large circular
spot on either side of its posterior end
nigromaculata,
H’. Blade of antennal scale having its anterior margin
retreating toward the inner angle. Sixth seg-
ment of abdomen without a large circular spot
seplemspinosa.
G’. Fifth segment of abdomen furnished with a superior
median carina.
H, Antero-internal angle of scale advanced, reaching

I12 RATHBUN

nearly as far as, or farther than, the spine.
Hands a little over twice as long as wide

nigricauda.

H’. Antero-internal angle of scale is not produced, and

the spine reaches much beyond the blade.

Hands two and a half to three times as long as

wide.
J. Scale shorter than the Stee exclusive of ros-
trum =. . : = 4 .« Qlaskenses.
J’. ‘scale -as long as the carapace, exclusive of
rostrum . . . . . alaskensis elongata.

E’. Sixth segment of abdomen convex, not sulcate, beneath.

FE. Hands very stout, only two and a fourth times as long
as wide; antepenult segment of outer maxillipeds
greatly dilated . 2 = Walla.

F’, Hands elongate, about three and a half times as long as
wide; antepenult segment of outer maxillipeds not
ditited as My pee ke coe Me nag aa Teer

C’. Carapace with 2 median spines.
D. Both median spines in front of the middle of the carapace.
E. Rostrum slightly seat has rounded at the tip. Surface
pubescent’ <°'": . . communis.
FE’. Rostrum ascending at an angle of 48 degrees, tip pointed.
Surface naked.

B., Eyes otsmoderatesizes sy jena) jouw simone) Meme caters

Fi, Eyes very largess once es aie enue Oy SSOP,

D’. Posterior of the median spines situated at about the middle

of the carapace ". 7 21. » gio” 6) at, SSPIUCAIRCQED.

Bl. Carapace without a median gastric spine aie ae SH Lerostres.

A’. Gastric region depressed below the general level of the carapace.
B. Second lateral carina of the carapace (counting from the middle)
unarmed.
C. Rostrum not spiniform, extremity obtuse.
D. First to fourth abdominal segments, inclusive, smooth.
E. Anterior median spine not advanced so far as the line of

the: orbitsi <<. 4s ; . 2.) «6Mmuntiia.
E’. Anterior median spine projecting in front of the line of the
orbits. ait hoop Uo ehe ie aciet. eu RNCGLELLG Tse

D’. First to fourth abdominal segments more or less carinated.
E. First to third segments of the pleon armed laterally with

2 spines each: <2) sivas 6 os ue Some te SSP UPOSESSeaa.

E’. First to third segments of pleon unarmed . . variadilts.

C’. Rostrum spiniform, sharp . . .. .. . © Sfimirostris.
B’. Second lateral carina of the carapace armed with a spine a little
behind the superior lateral spine . . . . . « « munitella.

CRANGON NIGRICAUDA Stimpson.

Crangon nigricauda STIMPSON, Proc. Calif. Acad. Sci., I, 97, 1856; Jour.
Boston Soc, Nat. Hist., VI, 496, pl. XXII, fig. 6, 1857.

DECAPODS 113

Crangon nigricauda HOLMES, Occas. Papers Calif. Acad. Sci., v1, 170, pl.
II, fig. 31, 1900, but not all refs. to synonymy.

Carapace only a little more than one third the length of the abdomen,
furnished with a single median gastric spine, and a hepatic spine, in a
transverse line. Rostrum rather short, grooved, rounded at the tip.
Suborbital and antennal spines present, the latter the larger.

The process on the outer side of the base of the antennules is broad,
irregularly ovate, and about reaches the tip of the first joint; flagella
longer than the peduncle, the inner one, and sometimes the outer one,
exceeding the scale. Antenne as long as, or longer than, the body,
acicle about two thirds the length of the carapace, the antero-internal
angle rounded and produced, reaching nearly as far forward as the spine
at the outer angle (sometimes farther than the spine). The outer maxil-
lipeds reach about as far forward as the antennal scale.

The first pair of feet may extend as far forward as the maxillipeds, or
be a little shorter; merus with a spine near the middle of
the lower side; hand oblong, a little more than twice as (Ue.
long as wide, with the edges subparallel; the margin
against which the finger closes is regularly convex and more mies Ca
nearly transverse than longitudinal; spinous pollex directed Bay, Calit eee
obliquely forward. A spine on the sternum between the bases of the
third pair of feet.

Antepenultimate segment of abdomen distinctly carinated in the mid-
dle; the sixth segment has a tendency to become carinated, and has a
shallow median groove. Telson grooved above, subacute at tip, and ex-
ceeding the sixth plus one half of the fifth. An acute tooth on the sixth
segment on either side of the base of the telson, and a spine at the
postero-inferior angles, between which there is a median ventral spine
curved backward; the posterior three fourths or more of the ventral sur-
face of this segment is marked by a groove.

Color in life, dark gray with a blackish tail (Holmes).

Dimensions.—Length of female (Catalogue No. 3052) from tip of
rostrum to tip of telson 68 mm., length of carapace 17 mm., of antennal
scale 10.5 mm.

Distribution.— According to Holmes (who includes here C. alaskensis),
this species ranges from Alaska to Lower California. I have seen speci-
mens ranging only from Comox, British Columbia, southward, as fol-
lows:

Comox, British Columbia (4/atross).

Neah Bay, Washington, in drag seine (AdZatross).
Puget Sound (T. Kincaid).

114 RATHBUN

Bodega Bay, California (University of California).

San Francisco Bay, California (W. H. Dall; Livingston Stone).

San Francisco Bay, off San Mateo, 3 fathoms, dredged (A/batross).

San Francisco Bay, on South Belmont oyster beds, seine (A/batross).

San Francisco Bay, in San Pablo Bay, in Chinese shrimp nets (A/batross).

San Francisco Bay, off San Bruno Point, in Chinese shrimp nets (4/-
batross).

San Francisco market (R. E. C. Stearns).

Monterey (D. S. Jordan).

Catalina Island (Albatross).

San Diego (H. Hemphill; D. S. Jordan; A/atross).

San Diego Bay, 534-12 fathoms (A/atross stations 3574, 3581, 3582,
3621).

Station 3674 of Albatross (locality not given), 31 fathoms.

San Geronimo Island, Lower California, 7 fathoms (A. W. Anthony).

Mogi, Japan (Dr. F. C. Dale, U. S. S. Falos, June 18, 1881); one speci-
men without chelipeds, which otherwise corresponds closely with the
American form; probably, however, distinct from C. afinis De Haan
and C. propinguus Stimpson.

Monterey (Owen, as C. vulgaris).

CRANGON NIGROMACULATA Lockington.

Distribution.—From northern California to Lower
(Cae. California, 3-33 fathoms.
Specimens in the National Museum range from Gulf of

Fic. 51. Crangon i 4 A
nigromaculata. Chelathe Farallones to San Diego, California; also found at
of 2 (x 23). Station ji .
3132. Turtle Bay, Lower California.

CRANGON ALASKENSIS Lockington.

Crangon alaskensis LOCKINGTON, Proc. Calif. Acad. Sci., vil, 34 [7], 1876

(1877). Mutiny Bay, Alaska.

Differs from C. nigricauda as follows:. The antennal scale is a little
longer and much narrower distally; the end of the blade is rounded and
not produced at the antero-internal angle; the spine is
much longer, extending considerably beyond the blade.

r The first pair of feet are shorter, reaching
Ce only to the middle of the terminal segment A
of the maxillipeds; the hands are more ,
Fic. 52. Cran- °
gon alaskensis. Slender and elongate, being from two and a
Chela of ¢ (Xx 33). i A Fic. 53. Crangon
Station 3652. half to three times as long as wide; their alaskensis. 9. Station
: omagte ° ° 3243. @. Dorsal view
distal margin is more oblique, forming an angle of about of rostral region (x
° : : 23). 5. Acicle (X 2).
forty-five degrees with the lengthwise margins.
Dimensions.—Length of large egg-bearing female from Kulukak Bay,

DECAPODS 115

Alaska (station 3243), 77-2 mm.; length of carapace 20.1 mm.; length
of antennal scale 14 mm.

Distribution.— East part of Bering Sea and along the Aleutian Islands,
in 3%-41 fathoms, at 38 stations of the Albatross.

Also taken at the following localities:

Off Kurile Islands, 14 fathoms (A/datross station 3652).

Gulf of Georgia, British Columbia, 67 fathoms (A/éatross station 2863).

Sucia Island, British Columbia (4/éatross).

Bellingham Bay, Washington, 11 fathoms (A/batross station 3612).

Strait of Fuca, 81 fathoms (A/éatross station 3596).

Port Townsend, Washington, rostrum narrower than typical (A/batross).

Puget Sound (T. Kincaid).

From Sitka northward to Port Clarence and westward along the Aleutian
Islands to Attu, 5-20 fathoms, at sixteen localities (W. H. Dall).

CRANGON ALASKENSIS ELONGATA Rathbun.

Crangon alaskensis elongata RATHBUN, Proc. U. S. Nat. Mus., XxIv, 888,
1902.

The typical C. alaskensis described above passes by insensible grada-
tions into a form occurring off the coast of California and Oregon, which
differs from it as follows: The rostrum is longer and
narrower. The outer flagellum of the antennules falls
considerably short of the antennal scale. The scale is
much longer than in the typical form, being equal to the A
length of the carapace exclusive of the rostrum. The
fourth segment of the abdomen shows signs of carination, ="

: : Fic. 5 Crangon
The telson is flattened above, but scarcely grooved; its petee nee e.
tip is more acute. Anterior part of cara-

Dimensions.—Length of ovigerous female from tip of ae a al
rostrum to tip of telson 55.7 mm.; length of carapace 13.5 mm.; length
of antennal scale 11.6 mm.

Type locality.— Off Santa Barbara, California, 29 fathoms, stations 2970,
2971 (Albatross).

Distribution.—This species occurs in abundance off the west coast of
the United States. It has been taken by the A/sazross at thirty stations
in 9-278 fathoms, off California, Oregon, Washington, and British Colum-
bia, and in Strait of Fuca and Puget Sound. The southern limit is off
Wilmington, California. It has also been collected in Monterey Harbor
by Dr. W. H. Dall, and at Pacific Grove by Mr. J. O. Snyder.

Variations.—The more northern of these specimens (those off the coast
of Washington more especially) have the scale not quite so long as the
types, but still equal to about four fifths the length of the carapace; the

116 RATHBUN

rostrum, too, is often intermediate between typical a/askensis and typical
alaskenstis elongata.

CRANGON SEPTEMSPINOSA Say.

Crangon septemspinosus SAY, Jour. Acad. Nat. Sci. Phila., 1, 246, 1818.
Crangon crangon and Crangon vulgaris of authors (part).

Associated sometimes with C. a/askensis, which it closely resembles ;
it can be recognized at once by the rounded (non-carinated) abdominal
segments. In Alaskan specimens the antennal scale is a little more than

two thirds the length of the carapace; it is equal to or
| = = exceeds the distance from the gastric spine to the pos-

terior margin; the extremity of the blade is obliquely
ee Boe subtruncate, slightly rounded, the antero-internal angle
(Xx 2). Station 323° retreating; the spine surpasses the blade to a distance
exceeding the anterior width of the blade. The hands are variable in
length, being from three to three and one half times as long as wide;
the obliquity of the terminal margin is as in C. alaskensis. All the ab-
dominal segments are free from carine; the sixth and seventh have a
slight median sulcus.

There is considerable variability in Atlantic specimens of this species,
in the length both of the scale and of the hands; I cannot see that our
Alaskan specimens differ essentially from those of Chesapeake Bay.

Dimensions. — Female, station 3230, length from tip of rostrum to tip
of telson 60.4 mm., length of carapace 15.6 mm., length of antennal
scale 11 mm.

Color.—It is probable that in life the species could readily be detected
by the color markings. In alcohol the speckles of the dorsal surface
are more uniform and closer than in C. alaskensis, and the telson and
uropods are darker than in C. alaskensis from the same dredge haul.

Distribution.—This species is less abundant along the Alaskan coast
than C. alaskensis,; it extends from the Arctic coast of Alaska at
Eschscholtz Bay southward along the eastern shore of Bering Sea to the
Shumagins; the separate localities are as follows:

Chamisso Harbor, Eschscholtz Bay, 5-8 fathoms (W. H. Dall).

Port Clarence, in the seine, with C. alaskensis (W. H. Dall).

St. Michael, Norton Sound (L. M. Turner; E. W. Nelson).

Head of Norton Sound, 5 fathoms (Point Barrow Expedition).

Off mouth of Yukon, 3% fathoms (E. W. Nelson).

Cape Etolin, Nunivak Island, anchorage, 8 fathoms, stony (W. H. Dall).
Hagemeister Strait, 8-15 fathoms, gravel (W. H. Dall).

Bristol Bay (C. L. McKay).

Bristol Bay, 314 fathoms, with C. adaskensis (Albatross station 3230).

DECAPODS 117

Bailey Harbor, Alaska Peninsula (4/batross).
Coal Harbor, Unga Island, Shumagins, 3 fathoms (W. H. Dall).

Also occurs along the eastern coast of North America, from East
Florida (Say) northward.

It becomes necessary to separate the common Cvangon of the Atlantic
and Alaskan coasts of North America from the form found in Europe.
These two species have usually been combined under the name Crangon
crangon (Linneus) or C. vu/garis Fabricius.

In Crangon crangon (of Europe) the antennal scale is wider at its
distal end, and that margin, although convex, trends forward toward the
inner angle. The spine is shorter than the distal width of the blade.

In Crangon septemspinosa Say (of America) the scale is narrower at its
distal end, and that margin, although convex, does not trend forward
toward the inner angle, but rather slopes backward. ‘The spine is as
long as, or longer than, the distal width of the blade.

CRANGON ALBA Holmes.

Crangon alba HOLMES, Occas. Papers Calif. Acad. Sci., VII, 174, 1900.
Monterey, dredged.

A short stout species. Carapace about two fifths as long as abdomen.
Rostrum broader than in C. nigricauda. The first segment of the anten-
nule exceeds the process on the outer side of its base;
inner flagellum exceeding the antennal scale; outer ~~
flagellum shorter than the scale. Blade of scale with
very oblique inner margin, the tip scarcely wider than
the adjacent portion of the spine; spine extending .

considerably beyond the blade. Scale «
about three fourths the length of carapace. Fic. 6 _ Crazgon
alba. @. San Diego.

1 eee Maxillipeds reaching the end of the ie oie eae
alba. Chela of ¢ blade, the antepenult segment much ex- (*?)-
it ata panded. The first pair of feet reach to the middle of
the last joint of the maxillipeds; the hand is very stout, wider than in
C. nigricauda, the length being two and one fourth times the width
measured from the inner base of the immovable spine; the anterior
margin is more longitudinal than transverse.

The segments of the abdomen are smoothly rounded; the sixth is
rounded beneath, not grooved; the seventh is not sulcate above.

Dimensions.—An ovigerous female (station 3679) measures 48 mm.
long, carapace 12.5 mm., scale 9.7 mm.

Distribution. —From Vancouver Island, British Columbia, to San Diego,
California, to a depth of 47 fathoms, at the following localities:

—

118 RATHBUN

Off Cape Beale, Vancouver Island, 34 fathoms, stations 2879, 2880.
Off Cape Flattery, Washington, 40 fathoms, station 2873.
Gulf of the Farallones, 29 fathoms, station 3100; 47 fathoms, station

3157-
Off Lobos Rocks, California, 41 fathoms, station 3185.
South of San Diego Bay, 22 fathoms, station 3679.
Off Cortez Bank, California, 47 fathoms, station 2922.
San Diego, depth not given (A/batross).
Pacific Grove, California (J. O. Snyder).

CRANGON HOLMESI Rathbun.

Crangon holmest RATHBUN, Proc. U.S. Nat. Mus., xxiv, 888, 1902.

A small species allied to C. aa Holmes in not having a groove on the
under side of the sixth abdominal segment.
Carapace a third as long as the abdomen; rostrum narrow-triangular,
short, with a rounded tip. Antennule as in C.a/da. Antenne nearly as
long as the body; scale either as long or nearly as
g long as the carapace exclusive of the rostrum;
‘ _ blade very narrow at the extremity; spine much
exceeding it. Maxillipeds exceeding the scale a
= little; antepenult segment not dilated. The first
Fi, 3h, Crane alms pair of feet reach to ee middle of the terminal
view of rostral zegion (8), Joint of the maxillipeds ; the hand is elongate, about
8. Acicle(X 4). ¢. Chela(x 8). three times as long as wide; the anterior margin,
against which the dactylus folds, is more longitudinal than transverse.
The third pair of legs is a very little longer than the second; they are
equally slender. The segments of the abdomen are smoothly rounded
above; the sixth without a ventral sulcus; the seventh is a little longer
than the sixth, or about as long as the antennal scale, and is exceeded by
the uropods.
Dimensions.—Ovigerous female, length 23 mm., length of carapace
5-3 mm., of antennal scale 4.2 mm.
Distribution. —Off Wilmington, California, 27 fathoms (A/batross sta-
tion 2939), type locality, 4 females.
Catalina Harbor, California, 30-40 fathoms, sandy mud; 1 female
(Wo. Dall):
Northwest of Cerros Island, Lower California, 58 fathoms (4/batross
station 2983), 4 females.

CRANGON STYLIROSTRIS Holmes.

Crangon stylirostris HOLMES, Occas. Papers Calif. Acad. Sci., VII, 174, pl.
II, figs. 33-35, 1900.

DECAPODS 11g

Rostrum long, narrow, grooved above, tapering to a narrow-acute tip
which is curved strongly downward and much compressed laterally. There
is no median gastric spine; but with a lens one can discern a scabrous
granule in the spot where the spine usually occurs. The antennular pe-
duncle has shorter segments than in C. alba, to which
this species is nearly related ; outer branch of flagellum
as long as, or longer than, the blade of the antennal
scale. Antenne shorter than the body. Scale shaped _ Fic. 59. Crangon sty-
as in C. aléa, but much shorter, being only a little over Gunceriaeaa 2
half as long as the carapace. Maxillipeds with the antepenult segment
outwardly dilated as in C.a/ba. The hands are slightly widened distally,
shorter and broader than in C. a/ba, the length being barely twice the
width; the anterior margin is more transverse than longitudinal. Second
pair of feet more slender and a little longer than the third.

Abdominal segments smoothly rounded above, not carinated; sixth
segment not sulcated below; seventh segment a very little longer than
the sixth. The uropods extend considerably beyond the telson. Each
segment in the male bears a median spine on the ventral side.

Dimensions.—Length of female 54 mm., of carapace 15.1 mm., of
acicle 9.1 mm.

Distribution.—Santa Cruz, California, in seine, April 12, 1897 (Alba-
tross); Trinidad, Humboldt Co., California (Holmes); Chirikof Island,
Alaska, anchorage, 9-14 fathoms, sand (W. H. Dall).

CRANGON DALLI Rathbun.
Crangon dalli RATHBUN, Proc. U.S. Nat. Mus., xxIv, 889, 1902.

Rostrum rather long and narrow, sides parallel for a portion of their
length, edges upturned. Second antennular segment about one and a
half times as long as the third; inner flagellum exceed-
ing the acicle; outer flagellum considerably shorter.
Acicle about seven tenths as long as the carapace; blade
obliquely subtruncate, inner angle rounded, receding ; spine
extending beyond the blade to no greater extent than the
anterior width of the blade. The first pair of feet reach
only to the middle of the terminal joint of the maxillipeds ;
the hands widen considerably from the proximal to the
distal end; the distal margin is transversely oblique; the

width of the palm is contained in its greatest length about
Fic. 60. Crangon 2 ~
dalli. @. Station two and two thirds times,

3287. a. Dorsal view

of rostral region(x The first to fifth segments of the abdomen are smooth;
3)._ 8. Acicle (x 2
¢.

Chela (x4). the sixth has two blunt but prominent longitudinal carine,

120 RATHBUN

with a deep intervening sulcus; the seventh segment has a shallow
median sulcus.

Dimensions.—Length of ovigerous female from tip of rostrum to tip
of telson 61.5 mm., length of carapace 16.2 mm., of antennal scale
11.5 mm.

Type locality.—Bering Sea, off Cape Seniavin, Alaska, 30 fathoms
(Albatross station 3287).

Distribution. —Bering Sea and Aleutian Islands to Sitka; Kamchatka,
Okhotsk Sea, and Kurile Islands; in 444-61 fathoms. Taken by the
Albatross at 70 stations, by W. H. Dall at 15 stations; by Lieutenant
G. M. Stoney, U. S. N., at 7 stations, by C. L. McKay at Bristol Bay.
Occurs in large numbers in Bering Sea.

Bellingham Bay, Washington, 11 fathoms, 200 specimens (Adbatross

station 3612).

Bering Island, stomach of Gadus macrocephalus, 1 specimen (N. Greb-
nitzky).

ae Bay, Avacha Bay, Kamchatka, 1 specimen (AJbatross).

Off Robben Island, Okhotsk Sea, 18 fathoms, 1 specimen (AJbatross

station 3646).

Off Kurile Islands, 14 fathoms, 1 specimen (A/batross station 3652).

Affinities. —This species is analogous to C. a//mani Kinahan of the
North Atlantic, but in that species the rostrum is shorter and narrower,
the antennal scale is shorter and the end of its blade more oblique, the
hand is longer and its anterior margin is more longitudinal than trans-
verse.

In the form of the carapace C. dali resembles C. alaskensis and C.
septemspinosa, with which it occurs, but is readily separated from them
by the prominent carine of the sixth abdominal segment and the
shape of the hands, which in those species are oblong, but in C.
dali widen distally.

CRANGON FRANCISCORUM Stimpson.

Crangon franciscorum STIMPSON, Proc. Cal. Acad. Sci., 1, 97, 1856; Jour.
Boston Soc. Nat. Hist., VI, 495, pl. XXII, fig. 5, 1857.— HOLMES, Oc-
cas. Papers Cal. Acad. Sci., VII, 172, 1900, and synonymy.

Carapace about three eighths as long as abdomen. Rostrum short,
triangular, tip broadly rounded. Antennal flagella very unequal, the
inner one longer than the antennal scale and greatly exceeding it, the
outer one less than half as long as the scale. Scale about three fourths
as long as the carapace, the blade broad and slightly rounded at the
extremity, spine exceeding it, flagellum nearly as long as, or longer than,
the body. Maxillipeds not attaining the end of the scale; first pair of

DECAPODS 12!

feet reaching the end of the maxillipeds; hand
very long and narrow, the length from four to At
four and a half times the width, longer in male

than in female, inflated near the base, outer margin
concave for the most part, inner convex, dactylus
when flexed almost longitudinal.

Abdominal segments smoothly rounded; fifth co

° ° : . = San Franciscomarket. 6, Chela
with a spine on either side of the posterior ex- of ¢. Station 2528.

tremity at the supero-lateral angles. Sixth segment as in C. nigricauda,

telson sharp.

Dimensions.—Length of male 77 mm., length of carapace 20.5 mm.,

of antennal scale 15 mm.

Distribution.—I have examined specimens from

Thorn Arm, southeastern Alaska, in drag seine (4/batross).

Promise Island, southeastern Alaska (A/datross).

Loring, Revillagigedo Islands, southeastern Alaska (A/batross).

Strait of Fuca (D. S. Jordan).

Puget Sound (D. S. Jordan).

Mouth Tenmile Creek, 10 miles south of Florence, Oregon (S. E. Meek,
U. S. Fish Commission).

Off Tillamook Rock, Oregon, 29 fathoms (A/éa/ross station 2883).

Off Yaquina Head, Oregon, 28 fathoms (4/atross station 3055).

San Francisco Bay (A/batross): on South Belmont Oyster Beds, with
seine; off San Mateo, 3 fathoms, with boat dredge; in San Pablo
Bay, in Chinese shrimp nets; San Pablo Bay, 12 fathoms, station
3097; and off San Bruno Point, in Chinese shrimp nets.

San Francisco Bay (various collectors).

San Francisco market (R. E. C. Stearns).

Pacific Grove, California (Stanford University).

San Diego, California (H. Hemphill).

Previously known also from Shoalwater Bay, Washington, and To-
males Bay, California.
This form is represented in deeper waters by a form which I have called

b

Fic. 61. Crangon francis-
rum (X 23). a. Chela of &.

CRANGON FRANCISCORUM ANGUSTIMANA Rathbun.

Crangon franciscorum angustimana RATHBUN, Proc. U.S. Nat. Mus., XxIv,
889, 1902.

Its chief difference lies in the shape of the hands,

which are narrower than in C. franciscorum, the differ-

ence being less in the female; in the
Fic. 62. Crangon fran- E
Chace s ese ly male the length is about five and a Sa
ela o x24). Station 2 és .
362. : half times the width, the dactylus is

ae : Fic. 63 C
more longitudinally placed, and the pollex is more trans- francisconum anguste

Z - Chel 5
verse and nearer the proximal end of the hand. Cee nr

122 RATHBUN

Dimensions. —Length of female 70.6 mm., of carapace 18.9 mm., of
antennal scale 13 mm. Length of hand of male 9.5 mm., width 1.8 mm.
Length of hand of male of C. franciscorum 11 mm., width 2.6 mm.

Type locality —Offt Chuck-a-nuts Island, Bellingham Bay, Washington,
11 fathoms (A/da/ross station 3612).

Distribution. — British Columbia to Oregon as follows:

Gulf of Georgia, British Columbia, 67 fathoms (A/satross station 2863).
Washington Sound, Strait of Fuca, 48 fathoms (A/datross station 2864).
Strait of Fuca, 53-67 fathoms (A/datross stations 3460, 3597).
Admiralty Inlet, Puget Sound, 40 fathoms (A/bazross station 2865).
Seattle, Washington, taken at the surface by electric light (A/batross).
Off Columbia River, 27 fathoms (A/éatross station 3065).
Astoria, Oregon (Aug. C. Kinney).
Off Tillamook Rock, Oregon, 23-28 fathoms (A/¢atross stations 3060,
3061).
It will be noticed that the two forms meet off the coast of Oregon,
where the differences are less sharply drawn.

CRANGON INTERMEDIA Stimpson.

Crangon intermedia STIMPSON, Proc. Acad. Nat. Sci. Phila., x11, 25 [94 of
separate], 1860. In Bering Sea near Cape Chepoonski, 40 fathoms.
Crangon tenuifrons KINGSLEY, Bull. Essex Inst., XIv, 128, pl. I, fig. Io,

1882. Marmot Island, Kadiak.

Carapace about two fifths as long as the abdomen; furnished with a me-
dian carina having 2 spines, a stout one about half way between the orbit
and the posterior margin, and a smaller one a little behind the rostrum.
Rostrum convex in profile, having a longitudinal sulcus and a rounded tip.

On the anterior margin there is a blunt spine at the outer angle of the
orbit and a sharp spine at the antero-lateral angle. On a line with the
orbital spine there is a smooth carina; below the carina a sharp spine
well forward and terminating a short carina; higher up a flattened blunt
spine or tubercle, situated about half way between the two median spines.

The antennal scale is short and broad, about half as long as the cara-
pace; the spine scarcely exceeds the blade. The outer maxillipeds ex-
ceed the antennal scale by half the length, or more, of their terminal
segment. ‘The first pair of feet overreach the scale slightly; the merus
has a short spine at the middle of its inner margin; the manus is about
three times as long as wide; the pollex arises at about the distal third;
the dactylus when flexed is slightly more longitudinal than transverse.
The sternum has a lobate median crest, which in the male has a tendency
to become spiniform.

The first five segments of the abdomen are distinctly carinated; the
sixth segment has two prominent carine.

DECAPODS 123

Dimensions.—Length of female (A/satross station 3504) from tip of
rostrum to tip of telson 45 mm.; length of carapace 12 mm.; length of
antennal scale 6 mm.

Distribution.— Bering Sea and Aleutian Islands, 21-g1 fathoms, at
31 stations of the Albarross; also off Vancouver Island, at 1 station in
24 fathoms; off coast of Kamchatka, 39-100 fathoms, at stations 3643
and 3781; usually in small numbers. Also taken by Dr. W. H. Dall
at 6 stations between Adak and Kadiak, in 9-60 fathoms. Bering Island
(N. Grebnitzky).

CRANGON COMMUNIS Rathbun.

Crangon communis RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. III, 556, 1899.

Carapace about a third as long as the abdomen; furnished with a
median carina on the anterior half, which is armed with two spines, both
in front of the middle; the posterior of these spines is the larger. The
rostrum is slender, tapering, ascending, hav-
ing a median sulcus, tip rounded, not extend- () (S)
ing beyond the eyes. A lateral spine nearly ,
in line with the anterior of the median spines ; '

two large spines on the anterior margin of the A

carapace, one of which is at the outer angle i es

of the orbit, and the other at the antero-lateral = c d
angle. Fic. 64. Crangon communis. @.

Station 3611. @. Dorsal view of ante-

The first segment of the antennular peduncle rior part of carapace (x 2). 4. Side
: view (x 2). c. Acicle (xX 2). @.
extends beyond the eyes to a distance exceed- Chela (x 23).
ing the length of the last two segments; the second segment is about
twice as long as the third. The antennal scale is elongate, about two
thirds the length of the carapace; the spine is slender and exceeds the
blade. The outer maxillipeds are very slender and extend slightly be-
yond the scale. The first pair of feet may not quite reach the scale or
may extend beyond it; the merus has an outer distal spine; the manus
is over three times as long as its average width; the immovable digit is
long and slender, and the dactylus when flexed is obliquely transverse.
The sternum has a series of median tubercles.

The abdomen is smoothly sculptured; there is a blunt median carina
on the third to fifth segments; the first and second segments have a
transverse sulcus; the third and fourth segments have each two trans-
verse sulci, which are connected either side of the median carina; the
sixth segment is furnished with two prominent longitudinal carine, a
median sulcus, and a sulcus on the outer side of each carina; the seventh

124 RATHBUN

segment has a deep median sulcus. The abdominal sulci are filled with
a thin short pubescence easily rubbed off. The sixth segment is about
one and a third times as long as the fifth.

Dimensions.— Female (station 3441), length from tip of rostrum to
tip of telson 64 mm., length of carapace 16 mm., length of antennal
scale 10.5 mm.

Harriman Expedition.—Juneau, 20 fathoms.

Distribution.—From Bering Sea to San Diego, California (including
Puget Sound and Strait of Fuca), 20-309 fathoms; taken at 142
stations by the A/vatross, often in large numbers. Less abundant at
southern localities. Off southeastern coast of Kamchatka, g6-100
fathoms, at two A/satross stations.

Two specimens were received from the Hopkins Laboratory at Pacific
Grove; they were probably taken in deep water.

One specimen was collected at Port Etches, Alaska, in 12-18 fathoms,
by W. H. Dall.

Variations.—The rostrum in normal individuals does not extend be-
yond the eyes; in many cases, however, it is evident that the rostrum
has been broken off or injured and later renewed, the new rostrum having
a narrow, elongated, and ascending tip exceeding the eyes, but of variable
length. In other respects these abnormal forms do not differ from the

types.
CRANGON RESIMA Rathbun.

Crangon resima RATHBUN, Proc. U. S. Nat. Mus., xxIv, 889, 1902.

Near C. communis. Carapace and rostrum a little more than one
third the length of the abdomen; a median carina on the middle third
of the carapace, armed with two spines, not

Sel= far apart, the posterior larger and more ele-
vated and in front of the middle. A sharp

hepatic spine is in a transverse line with the
anterior median spine and terminates a short

but prominent carina. Rostrum long, narrow,
pointed, advanced beyond the eyes, ascend-

a ) ing at an angle of about forty-five degrees,
Fic. 65. Crangon resima. &- slightly curved, and prolonged downward in

Station 2935. @. Dorsal view of a

Side view of game (XC ap) 2). ba thin compressed plate which appears spatu-

(X 38). d. Chela (X 4). late in a side view. The development of this

plate is dependent on age; specimens 20 mm. long show no evidence of it.
The second segment of the antennular peduncle is three times as long

as the third; the scale on the outer side at the base is very thin and

DECAPODS Bas

rhomboidal, tipped with a short spine; the flagella are not very unequal
in length, and exceed the antennal scale, but not the hair which fringes
it. The antennal flagellum is longer than the body; the scale is four
fifths of the carapace exclusive of the rostrum; its extremity is rather
broad and slightly rounded, and is surpassed by the outer spine.

The maxillipeds are slender and reach beyond the scale; the first
pair of feet extend to the middle of the last joint of the maxillipeds ;
they are similar to those of C. communis, the hands are shorter, three
times as long as broad; their anterior margin, and also the dactylus when
flexed, are more longitudinal than in C. communis.

The second and third pairs of feet are equally slender; the second
pair is shorter than the first; the third pair is considerably longer than
the first.

The first and second segments of the abdomen have a thickened band
along the posterior margin, in front of which there is a slight transverse
depression; the fifth segment has an obscure median carina with a short
depression on either side at the anterior end; for the rest, the first five
segments are smooth; the sixth has two prominent carine and a corre-
sponding median depression; the telson is slender, and has a slight
median sulcus; it exceeds the uropods.

Dimensions.—Length of ovigerous female (station 2935) from tip of
rostrum to tip of telson 48.3 mm., length of carapace 12.2 mm., length
of antennal scale 8.2 mm. Even females only 22.2 mm. long are laden
with eggs.

Type locality.—Off— San Diego, California, 124 fathoms (A/batross sta-
tion 2935).

Distribution.—From Monterey Bay, California, to San Domingo
Point, Lower California, station 3043; 44-266 fathoms, at 17 stations of
the Albatross, usually in small quantities.

CRANGON ABYSSORUM Rathbun.
Crangon abyssorum RATHBUN, Proc. U. S. Nat. Mus., XXIV, 890, I9g02.

Integument very thin, membranaceous.

Carapace and rostrum about two fifths the length of the abdomen.
‘Carapace with three carine, extending nearly its whole length; median
armed with 2 spines, one at the anterior third of the carapace (ros-
trum not included), the other minute, in front of the first. This small
spine is often broken off. Rostrum linear, flattened above, acute, as-
cending at an angle of about thirty degrees with the carapace, and
slightly curved, a little higher than wide, the lower part laterally com-
pressed, in length about one fourth the remainder of the carapace. The

126 RATHBUN

lateral carina extends backward from the orbital sinus. There is a strong
hepatic spine at the end of a short carina; also a spine at the outer angle
of the orbit and an antennal spine.

Eyes very large, hemispherical, their inner faces flat and contiguous,
the carinz covering nearly all the outer face. Antennules rather slender;
the scale at base is small, falcate, and
acuminate, the second segment about one
and a half times as long as the third; the
flagella both exceed the antennal scale,
the inner is only a little longer than the
outer. The antennz have a slender spine
at the outer base of the scale, which is
narrow-oblong and about three fourths as

EEG AC Honton heen unel Oe: long as the carapace (rostrum excluded) ;
Seaton 363 o Howol wen, of anterior the blade is rounded at the end, and
and dorsal spines. c. Acicle. d@. Chela. the spine projects beyond it; the peduncle
reaches to the distal third of the scale, its last joint is very slender, the
flagellum is about as long as the body.

The maxillipeds are slender and exceed the scale by nearly half the
length of the terminal segment. ‘The first pair of feet do not reach the
end of the scale. The merus has a small superior terminal spine. The
manus widens a little distally ; its length is three times its width measured
at the inner base of the spine; the dactylus is more longitudinal than
transverse. The second and third pairs of feet are equally filiform and
delicate; the second reaches to the middle or beyond the middle of the
palm of the first; the third reaches beyond the antennal scale; the fourth
pair also exceeds the scale; the fifth pair is a little shorter. In the male
the sternum has a median carina with a small spine on each segment.

The fifth segment of the abdomen has a low blunt carina; the sixth
has two prominent dorsal carinz and a low lateral carina on each side
near the dorsal, but extending only half the length of the segment; the
telson has a median furrow on its anterior fourth; it exceeds the uropods.

Dimensions.—Length of female 61 mm., of carapace and rostrum 17
mm., of carapace exclusive of rostrum 13.5 mm., of antennal scale 10.3
mm.

Type locality.— Bering Sea, southwest of Pribilof Islands, 1771 fath-
oms (A/batross station 3603).

Distribution.—A deep-water species extending from Bering Sea to the
southern extremity of California, in 685-1771 fathoms. Taken by the
Albatross at the following localities:

Bering Sea, south of Pribilof Islands, 1401 fathoms, station 3604.

DECAPODS 127

Bering Sea, southwest of Pribilof Islands, 1625 fathoms, station 3308.

Bering Sea, north of Rat Islands, 850 fathoms, station 3784.

Bering Sea, north of Unalaska, 987 fathoms, station 3607.

East of Prince of Wales Island, Alaska, 1569 fathoms, station 2859.

North of Islands of Four Mountains, Alaska, 1033 fathoms, station 3307.

Off Queen Charlotte Islands, British Columbia, 876 and 1588 fathoms,
stations 2860, 3342.

Off Sea Lion Rock, Washington, 685 to 877 fathoms, stations 3069,

3971, 3974, 3975:-
Off Cape Elizabeth, Washington, 831 fathoms, station 3344.
Off Tillamook Bay, Oregon, 786 fathoms, station 3346.
Off Cortez Bank, California, 984 fathoms, station 2919.

This species in general appearance much resembles a Lontophilus.

CRANGON MUNITA Dana.
Crangon munitus DANA, Crust. U. S. Expl. Exped., I, 536, 1852; pl.

MEK, fis--5, 195'5.

Female.—Body short and thick. Carapace less than two fifths the
length of the abdomen; the anterior and middle portion of the carapace
is depressed and pubescent; this depressed area is defined by a curved
line extending backward from the antero-lateral angle nearly to the pos-
terior margin on the median line, and is crossed in part by nine carine.
The median carina is the length of the depression and is armed with two
spines, one at two fifths
the distance from the
posterior margin, the
other at an equal dis-
tance from the first. First
lateral carina short, ter-
minating in a spine half
way between the two }

. . Fic. 67. Crangon munita. &(X 2). Station 2881. a. Dorsal
median spines. Second view of carapace and abdomen. 4. Side view of carapace.
lateral carina as long as the dorsal depression, and unarmed except for the
spine at the anterior end, which marks the outer angle of the orbit. Third
lateral carina short, terminating in a spine half way between the superior
lateral spine and the anterior median spine. Fourth lateral carina form-
ing the boundary of the depression and terminating in a spine at the
antero-lateral angle of the carapace. Rostrum short, reaching to a
line between the tips of the orbital and the antero-lateral spines; nearly
horizontal, medially sulcate, tip rounded.

The scale at the base of the antennules is broad, has a straight outer
margin, and is tipped with a small spine. The first and second segments
of the peduncle have a blunt spine at the outer distal angle, that on the

128 RATHBUN

second segment reaching the distal end of the third segment; the flagella
are unequal, both exceed the antennal scale. The antennal peduncle
reaches nearly to the end of the scale; the scale is short and broad, the
blade is rounded at the end, strongly advanced at the antero-internal
angle, and greatly exceeds the spine; the flagellum is one and a half
times the length of the carapace.

The maxillipeds surpass the antennal scale by nearly the whole length
of the last segment. The first pair of feet are shorter than the maxillipeds
but longer than the scale. Hands oblong, narrowing a little distally, or
presenting a slight constriction at the base of the spine; three times as
long as wide; anterior margin slightly longitudinal; spine short. The
second and third pairs of feet are equally long, and shorter than the first
pair; the fourth is shorter than the third, the fifth still shorter.

The first to fourth segments of the abdomen are dorsally smooth; the
fifth has a faint blunt median carina; the sixth is about one and a third
times the length of the fifth and has four blunt carinz, two of which are
submedian ; the lateral carinee converge posteriorly ; the posterior superior
angles are armed with a strong spine, the inferior with a small spine; the
telson has a deep median sulcus; the ridges on either side of it are armed
with two pairs of spinules; the tip is triangular, pointed.

The male differs from the female as follows: The spine at the antero-
lateral angle is longer and more slender, and is sometimes directed out-
ward. ‘The median carina is strong even to the posterior margin of the
carapace, and its spines are higher. The maxillipeds are shorter, exceed-
ing the scale by only half the length of the terminal segment. The
carine of the fifth and sixth abdominal segments are stronger. The
ventral surface of the thorax has a thin, strongly projecting carina, cut
into four large saw-teeth. Each segment of the abdomen, except the
last, has a median ventral spine.

Dimensions.—Male (station 2881), length of body 32.6 mm., of cara-
pace 9.1 mm., of antennal scale 4.2 mm. Female with eggs, length of
body 36.2 mm., of carapace ro mm., of scale 4 mm.

Distribution:

Port Etches, Alaska, 12-18 fathoms (W. H. Dall). __

Off Cape Beale, Vancouver Island, British Columbia, 24-34 fathoms
(Albatross stations 2879-2881).

Puget Sound (Dana, Walker; also collected by T. Kincaid).

Off Point Conception, California, 31 fathoms (A/bazross station 2908).

Off San Miguel Island, California, 53 fathoms (4//azross station 2894).
Dana describes the abdomen as smooth; the carina of the fifth seg-

ment is very indistinct in the female, while Dana in his figure indicates

two carinz on the sixth segment.

DECAPODS 129

CRANGON ACCLIVIS Rathbun.
Crangon acclivis RATHBUN, Proc. U.S, Nat. Mus., xxiv, 890, 1902.

Allied to C. munita. Lower lateral spine of carapace farther back;
anterior median spine much larger and farther forward; it projects well

in front of the posterior line of the orbits.
The rostrum is narrower, and ascending at
an angle of about forty-five degrees. The c
d
scale 3 mm.
Lype locality.— Ot Santa Cruz Island, Cal- Fic. 68. Crangon acclivis. a. Dorsal
iioraia, 260 fathoms (A/baross station Yew ot corapece. $x 4). Station 2946,

Dimensions.— Length of male 24.8 mm.,

anterior margin of the hand is more longi-
length of carapace 7.4 mm., of antennal
b. Side view (X 34). c. Acicle. 2 (x 8).

tudinal than in C. munifa. The divisions of

the ventral carina of the thorax in the male

are much more spiniform in this species.

2948), I male. d. Chela. @ (X 8).

Distribution.— Off the Trinity Islands, Alaska, 159 fathoms (A/batross
station 2853); Santa Catalina Island, California, 80 fathoms (A/batross
station 3664); southwest of San Nicholas Island, California, 158 fathoms
(Albatross station 2898). One specimen taken at each locality.

In the foregoing species the first four segments of the abdomen are
smooth. We now come to a group also resembling C. munita in general
appearance, but having the abdomen in front of the fifth segment more or
less carinated.

CRANGON VARIABILIS Rathbun.
Crangon variabilis RATHBUN, Proc. U. S. Nat. Mus., XxIv, 890, I902.

The carapace is much like that of C. munifa, but is nearly half as long
as the abdomen. The rostrum is short, convex from behind forward
and the tip rounded and
thickened. The anten-
nal scale has a concave
outer margin and the

spine equals or extends
beyond the blade. The

V 5 . chelipeds are stouter; the
e hands vary from two and
a half to three times as

id

¢ °
Fic. 69. Crangon variabilis, @. Station 2842. a. Dorsal long as wide. The sec-

view of carapace and abdomen (xX 2). 4. Side view of carapace 5
(X 2). ¢. Acicle (x 4). @. Chela (<4). ond, third, fourth, and

130 RATHBUN

fifth, and sometimes the first, segments of the abdomen are carinated,
and (except on the first two) the carina usually high, laterally com-
pressed, and blunt. The lateral spinules of the telson are considerably
behind the middle of the segment. The median spines or teeth of the
ventral side of the thorax in the male are laminar and dentiform; the
anterior one has an acuminate, the others an acute, tip.

Dimensions.— Female, length of body 32.2 mm., of carapace 9.2 mm.,
of hand 5.6 mm., width of hand 2mm. Male, length of body 24.6 mm.,
of carapace 7 mm., of hand 4.1 mm., width of hand 1.5 mm.

Dype locality.—Off North Head, Akutan Island, Alaska, 72 fathoms
(Albatross station 2842).

Distribution.—Bering Sea, Alaska Peninsula, and Aleutian Islands;
California; 50-695 fathoms. Taken by the A/batross at the following
stations :

Bering Sea, off the Pribilof Islands, 81-184 fathoms, stations 3486, 3488,

3489, 3500, 3602.

Off Rat Islands, 55 fathoms, station 3599.

North of Unalaska, 225 fathoms, station 3227.

Off Akutan Island, 91 fathoms, station 3548.

Northwest of Unimak Island, 121 fathoms, station 3224.
Unimak Pass, 50-56 fathoms, stations 3222, 3223.

Off Davidson Bank, 280 fathoms, station 3337.

South of Sannak Islands, 483 fathoms, station 3210.

Off Shumagin Bank, 625 and 138 fathoms, stations 3338, 3339.
Southeast of Chirikof Island, 695 fathoms, station 3340.

Off the Trinity Islands, 159 fathoms, station 2853.

Southwest of San Nicholas Island, California, 158 fathoms, station 2898.

Variations—There is more variation in this species than is usual in
this genus, perhaps owing to the great bathymetrical range.

The rostrum is often much thickened and elevated, the median sulcus
being almost obliterated.

The hand is not of uniform proportion. It is a little longer in the male
than in the female, and varies in specimens of the same sex from differ-
ent localities. ‘The median abdominal carina is in general less strong on
specimens from deeper water.

These differences are not constant enough to warrant the division of
this species.

CRANGON SPINOSISSIMA Rathbun.

Crangon spinosissima RATHBUN, Proc. U.S. Nat. Mus., XXIV, 891, 1902.
The relation of this species to C. variadilis is similar to that which C.

acclivis bears to C. munita. The lower lateral spine of the carapace is

more nearly in line with the superior lateral spine; the anterior median

DECAPODS 131

spine is larger and farther forward, advancing to a line in front of the
rear line of the orbits; the antero-lateral spine is longer and slenderer and
bent more outward; the rostrum is narrower, longer, and acute, and as-
cends at an angle equal to that of the spine directly behind it; the spine

SATs,

Fic. 70. Crangon © saa retin g (x 2). Station 3351. @. Dorsal view
of carapace. 4. Side view.

of the antennal scale is much shorter than the blade; the hand is longer
and narrower, and its anterior margin more longitudinal ; the spines of the
ventral surface of thorax and abdomen are long and slender. This species
furthermore differs from those hitherto described in having the segments
of the pleon armed laterally with spines, the first three segments with two
spines each, the next three with one spine each at the postero-lateral angle ;
the sixth segment also with a slender spine at the postero-superior angle.

Dimensions.— Female, length of body 36 mm., of carapace 10.5 mm.
Male, length of body 34.6 mm., of carapace ro mm. Length of hand
loose in bottle 5 mm., width 1.4 mm.

Lype locality.—Off Point Arena, California, 51 fathoms (A/batross sta-
tion 3351).

Distribution.— Off Oregon and California, 51-96 fathoms:

Off False Tillamook, Oregon, 62 fathoms, station 3090.
Off Point Conception, California, 96 fathoms, station 2906,

CRANGON SPINIROSTRIS Rathbun.

Crangon spinirostris RATHBUN, Proc. U. S. Nat. Mus., xxiv, 891, 1902.

Also allied to C. munita and its associates, but differing from them all
in its long spiniform rostrum.
The position of the spines of the carapace is Sa;
similar to that in C. munifa. The rostrum and the “Sy ~~ aut
spine behind are long, slender, and suberect, the
rostrum the longer. The orbital spine is slender
and ascending; the antero-lateral spine is directed «4
upward and outward. The spines of the first two
antennular segments are slender; that of the anten-

; : Fic. 71. C ind
nal scale is longer than usual, exceeding the blade. yospas.7" 9 (CoS "shin

a. Dorsal view of

The hand is about three times as long as wide. Of datapace. 8. Side view.
the projecting plates on the ventral surface of the thorax in the male, the

132 RATHBUN

anterior has an acuminate, the other a spiniform, tip. The third to fifth
segments of the abdomen are carinated; the anterior pair of lateral
spinules of the telson are two fifths the distance from the tip.

Dimensions. — Female, length of body 35.4 mm., length of carapace
10.5 mm., of scale 4.4 mm., of hand 5.2 mm., width of hand 1.8 mm.
Male, length of body 33 mm., length of carapace 10.3 mm., of scale 3.9 mm.

Distribution. — Bering Sea and off the Aleutian Islands, 276-625
fathoms, as follows:

North of Rat Islands, 270 fathoms (4/éatross station 3785).

South of Pribilof Islands, 276 fathoms (A/batross station 3608).

Off Tliuliuk Harbor, Unalaska, 309 fathoms (A/dazross station 3316).
North of Unalaska, 350-406 fathoms, stations 3329 (type locality) to

3332.
Off Shumagin Bank, 625 fathoms, station 3338.

CRANGON MUNITELLA Walker.

Crangon munitellus WALKER, Trans. Liverpool Biol. Soc., XII, 275, pl. XvI,
fig. 1, 1898. Puget Sound.—HOLMEs, Occas. Papers Calif. Acad. Sci.,
VII, 176, 1900.

This species is related to C. munitus, but differs from it and all its allies
in having the second lateral carina from the middle armed with a spine a
little behind the superior lateral spine. The distance between the two
median spines is equal to the distance between the posterior of these
spines and the posterior margin. The antero-lateral angle of the cara-
pace is armed with a spine. The end of the blade of the antennal scale
is so produced at the inner angle that it is more advanced than the spine
at the outer angle. The first four abdominal segments are smooth; the
fifth has a carina faintly indicated; the sixth has a broad, smooth carina
set off by a depression on either side; the seventh has a slight median
depression. The hands of the first pair of feet are two and a half times
as long as wide, oblong, swollen, the anterior margin more longitudinal
than transverse, spine very long.

Dimensions.—Female with eggs, length 25 mm., length of carapace
7 mm.

Distribution.—Puget Sound (Walker, Calman; also collected by T.
Kincaid); Pacific Grove, California (J. O. Snyder); Catalina Harbor,
California, 30-40 fathoms, sandy mud (W. H. Dall).

Genus Sclerocrangon Sars.

KEY TO THE SPECIES OF SCLEROCRANGON.

A. A long suberect spine situated on the rostrum; four spines on median
Carina: a es) a A ee Ge) ie ce a Ne Sze

DECAPODS 133

A’, No long suberect spine situated on the rostrum; fewer than four spines
on median carina.
B. Carapace elongate, about one fifth longer than wide; blade of an-

tennal scale considerably overreaching the spine. . . doreas.
B’. Carapace nearly as long as wide; blade of antennal scale not over-
reaching the spine. . . + + © + © + © 6 © » alata,

SCLEROCRANGON BOREAS (Phipps).

Cancer boreas PuiPpPs, Voyage North Pole, 190, pl. XII, fig. 1, 1774.

Crangon Boreas SABINE, in Suppl. to Appendix of Capt. Parry’s [First] Voyage,
p. ccxxxv, 1824.

Sclerocrangon boreas SARS, Den Norske Nordhavs-Expedition, XIv, Zoologi,
Crust., Pt. 1, p. 15, 1885.—HOLMES, Occas. Papers Calif. Acad. Sci. Vit,
177, 1900, and synonymy.

Carapace one fifth longer than wide; three equidistant spines on the
dorsal carina; a tuberculated carina on the branchial regions armed with
an anterior spine and one or two obscure teeth. Rostrum triangular,
acute; tip horizontal. Antero-lateral angles armed with a spine project-
ing forward or a little outward. Blade of antennal scale much produced
beyond the spine. Hand twice or more than twice as long as wide; distal
margin obliquely transverse; spine obliquely longitudinal. The merus has
an upper distal spine, the carpus a lower distal spine.

Abdomen sculptured, the anterior segment with a median dorsal carina
which forms a forward projecting tooth on the first segment; the sixth
segment with a double carina, with a sulcus between the ridges; also on
each side of the posterior margin a supero-lateral spine. Pleura of the
abdominal segments with only one spine each. Telson sulcate; two pairs
of lateral spinules on the terminal half.

Dimensions.—Male (Alaska), length 110 mm., length of carapace 32
mm., width of carapace at branchial spine 26.4 mm., length of hand
13.2 mm., width of hand at inner base of spine 6 mm.

Harriman Expedition,—Kadiak, 5 fathoms (W. R. Coe); Berg Bay,
Glacier Bay.

Distribution,— Arctic Europe and America; Atlantic coast of America
southward to Cape Cod; Alaska (as far south as Killisnoo) ; ‘ California’
(Ross, Owen); Siberia; Kamchatka (Ross). Depth, 1-44 fathoms.

Numerous specimens were taken by the 4/atross in Bristol Bay and
vicinity in 31%{/-17 fathoms; by the U. S. R. S. Corwin in lat. 669 12’
oo” N., long. 168° 54/ oo” W., 30 fathoms, lat. 65° 49’ 15’ N., long.
169° 04! 30 W., 26 fathoms, and lat. 71° 02/ oo” N., long. 157° 46/
oo” W., 19 fathoms.

Killisnoo, Alaska (Northwestern Trading Company).
Plover Bay, Siberia, 10-25 fathoms (W. H. Dall).

134 RATHBUN

Cape Etolin, Nunivak Island, 8 fathoms (W. H. Dall).
Various localities in the Aleutians and Alaska Peninsula, 6-28 fathoms

(W. H. Dall).

SCLEROCRANGON ALATA Rathbun.
Plate 11, fig. 2.

Sclerocrangon alata RATHBUN, Proc. U. S. Nat. Mus., xxIv, 891, 1902.

Allied to S. doreas. Carapace only slightly broader than long; surface
very uneven, sculptured and pitted. Median carina armed with a small
spine just behind
the orbits, a still
smaller one at
the middle of the
carapace, and a
tubercle near the
posterior mar-
gin. Rostrum
with an acute

Fic. 72. Sclerocrangon alata. % (X 2). Station 2865. @. Dorsal view of upturned and
carapace and abdomen. 4. Side view ef carapace. curved tip. A

spine at the outer angle of the orbit. Antero-lateral angles broadly
alate. Branchial carina having a strong anterior spine; posterior half
protuberant and deeply pitted, as is also the surface below the crest.
The spine of the antennal scale extends as far as or a little farther than
the blade.

The first pair of feet have a strong upper distal spine on the merus, a
large outer dentiform expansion on the carpus, and a
very short stout hand less than twice as long as broad, « 5
the anterior margin nearly transverse and the spine nearly
longitudinal.

The abdomen is sculptured. ot eee

The median carina of the first five segments is very eae eee ee
blunt, and on the third segment especially prominent. % Acicle. 4. Chela.
The pleura are without spines. The lateral spinules (one or two pairs)
on the telson are situated at about its middle.

Dimensions.— Male, length 38 mm., length of carapace 11 mm., width
of carapace at branchial spine 10.7 mm., length of hand 4.4 mm., width
of hand 2.5 mm. A female 25.7 mm. long is laden with ova.

Distribution, —¥From Bering Sea to Puget Sound, 6-91 fathoms.
Usually not more than one or two specimens in a haul.

Taken at the following localities by the 4/atross:

DECAPODS 135

Bering Sea, off Akutan Island, 36 and gr fathoms, stations 3546 and
48.

N Be eck of Unimak Island, 43 fathoms, station 3262.

Off south entrance to Akutan Pass, 45 fathoms, station 2843.

Southwest of Sannak Islands, 41 fathoms, station 3213.

Shumagins, 21 and 35 fathoms, stations 2850 and 2851.

Strait of Fuca, 37 fathoms, station 3593.

Admiralty Inlet, Puget Sound, 40 fathoms, station 2865 (type locality).

By Dr. Wa, Be Dallat:

Constantine Harbor, Amchitka, 6-10 fathoms.
Bay of Islands, Adak, 9-16 fathoms.
Captains Harbor and Iliuliuk Harbor, Unalaska, g-15 fathoms.
Belkofski Bay, 15-25 fathoms.
Coal Harbor, Unga, 8-9 fathoms.
Popof Strait, Shumagins, 6 fathoms.
Chiniak Bay, Kadiak.
By N. Grebnitzky, at Bering Island, from stomach of Gadus macro-

cephalus.

SCLEROCRANGON SHARPI (Ortmann).
Plate 11, figs; 1, ta.

Paracrangon echinatus SHARP (non Dana), Proc. Acad. Nat. Sci. Phila., 1893,
126.
Crangon (Sclerocrangon) sharpi ORTMANN, Proc. Acad. Nat. Sci. Phila., 1895,

178.

Carapace about as broad as long, with three keels, the median one with
four strong spines, the first longest and placed on the upper margin of the
rostrum, the second nearly as long and situated immediately behind the
base of the rostrum ; the first, third, and fourth spines are directed obliquely
upward and forward, the second directly upward or a little backward.
The rostrum is short, not reaching the end of the first antennular seg-
ment, sharp and curved upward. ‘The lateral keels are formed by four
spines; the foremost, on the anterior margin of the carapace near the
base of the antenna, is the largest, and is directed obliquely outward and
forward, more than half as long as carapace; the three others are smaller
but sharp. There is a sharp spine at the outer angle of the orbit and
another on the outer side of the antenna at the base of the scale.

Merus of first pair of feet with a strong superior terminal spine; car-
pus with an outer, an inferior, and a smaller inner spine; hand more
than twice as long as broad; anterior margin nearly transverse; spine
short.

The abdomen is sculptured, first to sixth segments with a median keel,
that of the third arched and produced somewhat posteriorly, that on the
third to sixth segments with a median furrow, that of the sixth terminat-

136 RATHBUN

ing in two spines. Two other spines are placed at the posterior margin
of this segment, one on each side. Fifth segment furnished with a sharp
spine on the posterior margin near the median line; also a small spine
on the lateral surface. Lateral faces of the first five segments sculptured
by two irregular transverse furrows, sixth segment laterally with an
indistinct longitudinal ridge. Pleura of first to fourth segments tri-
angular, inferior angles blunt, without spines. Sternal crest low, dentate,
terminating anteriorly in a spine.

Dimensions. —Female, length 36.7 mm., length of carapace 11.5 mm.,
width of carapace between tips of antero-lateral spines 10.4 mm.

Distribution.— Bering Sea, Aleutian Islands, and Alaska Peninsula, 35—
150 fathoms. Taken by the Albatross at the following stations:

Off Pribilof Islands, 54 and 150 fathoms, stations 3552, 3486.

Pumicestone Bay, Unalaska, and vicinity, 35 and 59 fathoms, stations
3322, 3319-

Off North Head, Akutan Island, 56 and 72 fathoms, stations 2841, 2842.

Off south entrance to Akutan Pass, 45 fathoms, station 2843.

Northwest of Unimak Island, 43 fathoms, station 3262.

Portlock Bank, 51 fathoms, station 2857.

Marmot Isle, Kadiak, 45 fathoms (Sharp).
Genus Nectocrangon Brandt.

KEY TO THE SPECIES OF NECTOCRANGON.

A. Carapace with only two median spines behind the anterior margin.
B. Carine of sixth segment of abdomen not terminating in a small

sharp tooth or spine. . . ; : : lar.
B’. Carine of sixth segment terminating in a small sharp tooth or
spine.

C. All the abdominal segments carinated.

D. Eyes with low tubercle. Spines of carapace appressed; those
on anterior margin short. Hand ene about five times
aslongas wide .. <% « 16, weentara,

D’. Eyes with elongated sharp ‘tubercle. Spines of carapace sub-

erect; those on anterior margin long, slender. Hand
shorter, only three and a half to four times as long as wide
ovifer.
C’, The first and second abdominal segments not carinated
californiensis.
A’, Carapace with three or four median spines behind the anterior mar-
gin.
B. Carinz of sixth segment of abdomen terminating posteriorly in a
small spine. . . «| +e 6 1) @laSkenses:
B’. Carine of sixth segment of abdomen not terminating posteriorly
in a small spine.
C. First to fourth segments of abdomen carinated . . . crassa.
C’. First to fourth segments of abdomen not carinated. . “evior.

DECAPODS 137

NECTOCRANGON LAR (Owen).

Crangon lar OWEN, in Zoology of Capt. Beechey’s Voyage [of the Blossom],
Crustacea, p. 88, pl. XXVIII, fig. 1, 1839.
Argis lar KROYER, Naturh. Tidssk., Iv, 255, pl. v, figs. 45-62, 1842.
Nectocrangon lar STIMPSON, Proc. Acad. Nat. Sci. Phila., 25, 1860.—ORT-
MANN, Proc. Acad. Nat. Sci. Phila., 181, 1895.
Female.—Carapace about two fifths or one third as long as the abdo-
men; two spines on the median line, the distance between them being
greater than the distance from the anterior spine to the anterior margin,

and about equal to or less than the distance from the posterior spine to

Fic. 74. Mectocrangon lar. a. Dorsal view of carapace andabdomen. ¢ (nat. size). Station 3642.
5, Side view of carapace and part of abdomen. @ (nat. size). Station 3642. ¢. Side view of abdomen.
8 (X 14). Station 3779.

the posterior margin. Sometimes there is a tubercle behind the rostral
spine. A low blunt irregular carina runs from the orbit nearly to the
posterior margin; below this a spine on either side; a short antennal
spine on the anterior margin. Blade of antennal scale exceeding the spine.

Hands of first pair of feet from three and a half to four times as long
as the width of the palm. The anterior margin forms an angle of about
forty-five degrees with the side margins. First to fifth abdominal seg-
ments furnished with a sharp median carina; sixth seg-
ment with two sharp caring, which toward the posterior
end become lower and may disappear altogether in
front of the posterior margin. Pleura of first four
segments with rounded entire angles; fifth and sixth ¢ e

5 ; Fic. 75. Nectocran-

segments with an acute tooth or small spine at the gox dar (x 18). a. Aci-
postero-inferior angles; sixth segment with a supero- Chela, Reco ems
lateral spine on each side of the posterior margin. eo geet
Seventh segment with two blunt carinz armed with three pairs of spin-
ules on the terminal half.

Dimensions.— Female, length 79.2 mm., length of carapace 21.5 mm.

Male.—The male differs markedly from the female. It is much smaller.
The eyes are considerably larger. The anterior margin of the hand is

138 RATHBUN

more transverse, making the palm appear longer. The abdominal
carinz are higher and thinner, especially on the third segment; the in-
fero-posterior angle of the fourth segment may have a small spine. The
abdomen is more elongate, notably the sixth segment.

Dimensions.— Large male, length 55.7 mm., length of carapace 13.6 mm.

Distribution.— Arctic coast of Alaska and Siberia southward to Sitka
and Kurile Islands; Greenland (Kroyer). Beach to 47 fathoms.

In the U. S. National Museum this species is represented by numer-
ous specimens ranging from the Arctic coast of Alaska and Siberia south-
ward via Bering Strait to Bering Sea, lat. 56° 12’ o0// N. Within these
limits it is very abundant and was taken at 21 stations of the A/éatross,
at 8 stations by Lieutenant G. M. Stoney, U.S. N., at 7 stations by W. H.
Dall, at 2 stations by the U.S. R. S. Corwin, off the mouth of the Yukon
by E. W. Nelson, and at Cape Smith by the Point Barrow Expedition.

Also taken at the following localities:

Shahafka Cove, Kadiak, 12-14 fathoms (W. H. Dall).

Sitka Harbor, 15 fathoms (W. H. Dall).

Avacha Bay, Kamchatka (L. Stejneger and Alvatross; also recorded by
Stimpson).

Rakovaya Bay, Avacha Bay (Adbatross).

Off Kamchatka, 12-13 fathoms (A/batross stations 3776, 3777, 3779;
3780).

Off Iturup Island, Kurile Islands, 14-18 fathoms (A/datross stations 3652,

3653).
Off Robben Island, Okhotsk Sea, 18-20 fathoms, stations 3646, 3647.

NECTOCRANGON DENTATA Rathbun.

Nectocrangon lar SMITH, Trans. Conn. Acad. Arts Sci., v, 61, 1879 (part);
not JV. /ar(Owen).—RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. 111, 556, 1899 (part)—-HOLMES, Occas. Papers
Calif. Acad. Sci., vil, 178, 1900 (part)—-ORTMANN, Proc. Acad. Nat. Sci.
Phila., 1901, 164 (part).

Nectocrangon dentata RATHBUN, Proc. U. S. Nat. Mus., XXIV, 892, 1902.

Fic. 76. Nectocrangon dentata. @ (X 14). Station 3216. a. Dorsal view of
carapace and abdomen. 4. Side view of carapace and part of abdomen.

DECAPODS 139

The typical Wectocrangon Jar is primarily an Arctic species, and is re-
placed in the Aleutian Islands and the southern part of Bering Sea by a
species which is very closely allied, and has been heretofore
united with JV. Zar. It differs chiefly in the carinz of the sixth
abdominal segment terminating posteriorly in a small sharp
tooth or spine. The hand is more elongate, being about five p
times or more than five times as long as its width across the palm. “

Dimensions.— Female, length 73 mm., length of carapace Neri de

° dentata. @
19 mm., length of hand ro mm., width of palm 2.1 mm. (< 13). Sta-

Type locality.—Off Sitkalidak Island, Alaska, 69 fathoms Acie” %
(Albatross station 2855). a

Distribution.— Bering Sea southward to Sitka and southeast coast of
Kamchatka; Atlantic coast of North America from Greenland to Nova
Scotia, 6 to 96 fathoms.

From Bering Sea, in lat. 59° 55’ 00” N., southward to Aleutian Islands
and Alaska Peninsula, at 65 stations of the 4d/éatross, 21-93 fathoms.

Southeast coast of Kamchatka, 96 fathoms (A/batross).
Aleutian Islands eastward to Sitka Harbor, 6-80 fathoms (W. H. Dall).
Plover Bay, Siberia, 10-25 fathoms (W. H. Dall).

It will be noticed that the habitats of V. Jar and JV. dentata overlap in
Bering Sea, that each is occasionally found at the extreme limit of the
other’s range (e.g., 4V. dar at Kadiak and Vancouver Island, and JV.
dentata at Plover Bay), and that 1V. dentata extends into deeper water.

The species recorded from the North Atlantic by Professor S. I. Smith,
under the name J. Zar, is, I think, identical with V. dentata. The same
form was collected by the Princeton Expedition at Greenland; specimens
from Granville Bay are in the National Museum.

NECTOCRANGON OVIFER Rathbun.

Nectocrangon lar RATHBUN, The Fur Seals and Fur-Seal Islands of the North
Pacific Ocean, Pt. III, 556, 1899 (part).
Nectocrangon ovifer RATHBUN, Proc. U. S. Nat. Mus., XXIV, 892, 1902.

In the deeper waters of Bering Sea there lives another species of JVec/o-
crangon closely allied to V. dar and XW. dentata, Like them, it bears two
median spines on the carapace; the median crest is, however, higher,
and the spines more ascending; the three spines of the anterior margin
above the eyes are longer and more deeply separated from each other.
The tubercle on the anterior surface of the eye is more prominent and
acute; the eyes themselves are of greater size. The spine of the antennal
scale extends away beyond the blade, much more so than the allied species.
The pleon is relatively shorter than in V. Jar and NV. dentafa,; the median
carina in the female is higher; the carine of the sixth segment end in a

140 RATHBUN

small spine or tooth, as in JV. dentata. The median carina of the male is
higher than in the female, but there is less difference than in the above

Fic. 78. Mectocrangon ovifer. § (nat. size). Station 3225. a@. Dorsal view of
carapace and abdomen. 46. Side view of carapace and part of abdomen.

species. The hands are from three and a half to four times as long as
their width across the palm, like those of VV. far. The eggs are larger
than those of 2V. Zar or WV. dentata, having a diameter of

¢ 6 about 1.8 mm. as against 1.2 in WV. denfata.
| Dimensions.—Female bearing eggs, length 60 mm.,
length of carapace 17.4 mm. Male, length 35.5 mm.,

Fic. 79. Necto. length of carapace g mm.

o ni a oe Type locality.—Off the Trinity Islands, Alaska, 159
te etka” fathoms (Albatross station 2853).

Distribution.— Bering Sea, from 57° 39/ oo” north latitude to Aleutian
Islands and Alaska Peninsula as far as Kadiak, 56-368 fathoms, at 26
stations of the A/atross.

The three foregoing species are not to be separated at a glance, but
the examination of hundreds of specimens of each shows that the differ-

ences here mentioned are constant.

NECTOCRANGON CALIFORNIENSIS Rathbun.

Nectocrangon californiensis RATHBUN, Proc. U.S. Nat. Mus., XXIV, 892, 1902.

Allied to ZV. ovifer. Represented only by small specimens which differ
from specimens of JV. ovifer of equal size. The anterior margin outside
the orbital fissure is fur- 9
nished with two spines
close together, instead of 3
one in the allied species.
The eyes are smaller than in WV. ovifer,
but have a prominent tubercle. The
spine of the antennal scale extends only
slightly beyond the blade. The hands ) "Shon séeq “a Dorsal view of cata
are from three and a half to four times) Pare s57ebdomens 2) Size view.
as long as wide; they have the spine situated nearer the proximal end

DECAPODS 141

than in JV. ovifer, and the anterior margin (against which the dactylus
fits) in consequence more longitudinal.
The first and second abdominal segments are without a 2

&
carina; that on the third and fourth segments is rather feebly
developed.
Dimensions.—Male, length 31 mm., length of carapace 5, 9, voc.

tocrangon calie
8.6 mm. JSorniensts (X

Dj pe locality.—Ofk Santa Catalina Island, California, 80 3)... Sates
fathoms (A/éatross station 3664). 6. Chela.

Additional localities.—Off Santa Cruz Island, California, 155 fathoms,
station 2949; south of Santa Catalina Island, 59 fathoms, station 366s

(specimen 39 mm. long).

NECTOCRANGON ALASKENSIS Kingsley.

Nectocrangon alaskensis KINGSLEY, Bull. Essex Inst., XIv, 128, 1882.
Female.—Three equidistant spines on the median carina, and between

these and the spine on the anterior margin there is usually a small spine.

Lateral carina scarcely traceable. Antennal spine large. Spine of an-

Fic. 82. WNectocrangon alaskensis. ¢ (X_14). a. Dorsal view of carapace and
abdomen. Station 2869. 6. Side view. Station 2868.

tennal scale overreaching the blade. Hands of first pair of feet about
four and a half times as long as wide, anterior margin longitudinally
oblique. Sternal crest in the male armed with slender spines. First to
fourth abdominal segments smoothly rounded; fifth having a

’ sharp median crest ending posteriorly in a sharp spine, and a

spine at the postero-inferior angles ; sixth segment with two sharp

crests terminating in small spines; the posterior margin has

Fic. 83. also a large supero-lateral spine, and a small infero-lateral

Nectocran-

gon alasken- 1
ie 8 (% spine.

12). Station — )faJe,—Smaller than the female. The spine at the antero-
2869. a. Aci-

cle. 6. Chela. Jateral angle is much longer than in the female, considerably
overlapping the antennal scale. The scale is a little longer also. The

a

142 RATHBUN

first to fourth segments of the abdomen have a median carina, usually
sharp and high on the first three segments but not extending the full
length of each segment.

Dimensions.— Female, length 66.5 mm., length of carapace 17.7 mm.
Male, length 44.3 mm., length of carapace 11.2 mm.

Distribution.— From Bering Sea, near the Pribilof Islands, to Oregon,
including Puget Sound, 24-121 fathoms. Taken by the Albatross at 38
stations.

Also collected by W. H. Dall at the following localities:

Iliuliuk, Unalaska, 3-6 fathoms; Captains Bay, Unalaska; Shahafka
Cove, Kadiak.

NECTOCRANGON CRASSA Rathbun.

Nectocrangon crassa RATHBUN, The Fur Seals and Fur-Seal Islands of the
North Pacific Ocean, Pt. Il], 556, 1899.

Female.—Body stout.

Carapace more than one third the length of the abdomen; three
strong equidistant spines on the median carina; between the first of these
spines and the
spine on the ante- «
rior margin there
is a small tooth
or spinule. Lat-
eral carina well marked. Antennal
spine not reaching the antennal scale.

Spine of scale extending well beyond

the blade Fic. 84. Nectocrangon crassa. 2 (X13). Sta-
: tion 3245. @. Dorsal view of carapace and ab-

The hand of the first pair of feet is domen. 4. Side view.
a little more than three times as long as wide; its anterior margin is longi-
tudinally oblique; the spine is short and stout. Sternal crest of male
armed with spines.

Abdomen slightly sculptured; first to fifth segments furnished with a

(\ (| blunt median carina; sixth and seventh segments having
a

two carinz, those of the sixth segment converging and
disappearing toward the posterior margin; this margin has
Fic. 85. Mec- a large lobe at the supero-lateral angles; the fourth, fifth,

tocrangon crassa.

2K, St and sixth segments have a small tooth or spine at infero-
cle. “4. Chela. posterior angles.

Male.—Smaller and more slender than female; abdominal carine
stronger.

Dimensions.— Female, length 48.5 mm., length of carapace 13 mm.

DECAPODS 143

Distribution.—Bering Sea and Aleutian Islands, 11% to 55 fathoms
at 24 stations of the A/datross, Aleutian Islands to Sitka, 5 to 28 fathoms,
at 10 localities (W. H. Dall); Alaska (U.S. R. S. Corwin), one specimen ;
Bering Island, some specimens taken from the stomach of Gadus macro-
cephalus (N. Grebnitzky).

NECTOCRANGON LEVIOR Rathbun.

Nectocrangon levior RATHBUN, Proc. U.S. Nat. Mus., XXIv, 892, 1902.

This species is very close to WV. crassa. The first four abdominal seg-
ments are non-carinate, except for a short feeble elevation on the first;
the first segment has a
median tubercle; and -
the pleura of all the
segments have shallow
depressions suggestive
of the sculpturing of JV. crassa, the carine
of the fifth and sixth segments are less
clearly marked than in &. crassa. qiiaece: “ieerneneoredceed oni

Dimensions.— Female, length Ag ata ae, ae. poe 3
length of carapace 13.1 mm.

Distribution.—This species is more rare than WV. crassa and extends
farther south. One female was taken in Admiralty Inlet, Puget Sound,

40 fathoms, station 2865, by the A/batross, and serves as the
(| 1" type. Another female was dredged off Cape Beale, Vancou-

6

ver Island, in 34 fathoms, station 2880. One female from

Fic. 87; Nee- Portlock Bank, 68 fathoms, station 2856. Six females and
tocrangon levi- one male come from Davidson Bank, south of the Aleutian
Station "2865. Islands, 42 fathoms, station 2845. One or two specimens
Chela. “were taken by Dr. Dall at each of the following localities :

Port Levashef, Unalaska, 20-30 fathoms.
Semidi Islands, 12-28 fathoms.
West side Middleton Island, 10-12 fathoms.

Genus Paracrangon Dana.

PARACRANGON ECHINATA Dana.

Paracrangon echinatus DANA, ree Acad. Nat. Sei... Phila;, -1852; p.:203
Crust: OU; S.. Expl. Exped., 1, p. 538, 1652, pl. XXXII, fig.'6, 1955.
— HOLMES, Occas. Papers Calf Acad. Sci., vil, 176, pl. J, figs. 36 and
37, 1900, and synonymy.

Carapace furnished with a median carina unequally four-toothed.
Rostrum long, obliquely erect; posterior margin with one tooth near the

144 RATHBUN

middle, anterior margin with one tooth near the tip, and at base a long
curved spine. Sides of carapace carinated in such a way as to form
irregular quadrangular spaces, with spines at most of the angles; four or
five of these spines form an irregular lateral carina, the anterior spine
being much the longest, about four fifths the length of the rostrum. On
the anterior margin of the carapace there is a spine at the outer angle of
the orbit and another at the antero-lateral angle. The antennular
peduncle is two thirds as long as the carapace; the second segment is
more than twice as long as the third; the basal scale is very short and
ovate; the outer flagellum is very much stouter and twice as long as the
inner. ‘The antennal scale is as long as the antennular peduncle but not
so long as the antennal peduncle; the laminar portion projects consider-
ably beyond the short, stout outer spine; the outer margin is slightly
concave. The flagellum when bent back reaches the sixth abdominal
segment. The maxillipeds reach beyond the antennal peduncle by half
the length of the dactylus. The first pair of feet are about as long as the
maxillipeds; the upper margin of the merus terminates distally in a
tooth; the hands are narrow-elongate, about four times as long as wide;
the digital spine is very long and slender and directed in an obliquely
longitudinal direction; the anterior margin is more longitudinal than
transverse. The third pair of feet (it is to be borne in mind that the
second pair are absent) are very slender and barely reach to the end of the
first pair; the fourth and fifth pairs are of about equal length, and stretch
a little beyond the first pair.

The abdomen behind the second segment is carinate, the carina of
the third segment being especially high; on the sixth and seventh seg-
ments it is medially sulcate. The first to fifth segments are deeply
sculptured with transverse grooves, and their inferior margins are
pointed, those of the fourth and fifth segments being sharply so. In the
male the pleura are much more spiniform than in the female. The
fourth segment has on each side of its posterior margin a blunt tooth;
the fifth segment has a corresponding spine. The sixth segment has on
each side two lateral and one posterior spine. The telson is armed with
two small spines at the proximal end, and three pairs of lateral spinules
on the terminal half. The outer uropod is shorter than the inner, is
obliquely truncated at the end, and has a broad blunt tooth at the
postero-lateral angle. The median ventral line of the pleon is armed
with very long spines.

Dimenstons.— Female with eggs, length of body measured from tip of
spine at anterior base of rostrum 47.4 mm., length of carapace from
same point 16 mm., length of carapace from posterior line of orbit 12 mm.

DECAPODS 145

Distribution.—Port Etches, Alaska, to Puget Sound; 1 Japan. Port
Etches, Alaska, 12 to 18 fathoms (W. H. Dall), one specimen. Dredged
by the Addazross in the Straits of Fuca in 40 to 48 fathoms at stations
2864, 3464, and 3465, and in Puget Sound in 40 fathoms, station 2865.

Puget Sound (T. Kincaid).

Puget Sound, Oregon, dredged (Dana).

McLaughlin Bay, Campbell Island, 10-30 fathoms (Whiteaves).
Vancouver Island (Smith).

North of Japan, lat. 44° 27’ oo” N., long. 141° 22/ oo” E. (Miers).

Family SERGE STIDA.
Genus Sergestes Milne Edwards.
KEY TO THE SPECIES OF SERGESTES.

A. First joint of antennular peduncle only slightly longer than third joint.
Rostrum spiniform. Cornea larger than eye-stalk. . atlanticus.

A’. First joint of antennular peduncle considerably longer than third
joint. Rostrum lobiform. Cornea no larger than eye-stalk,

Sp. indet,

SERGESTES ATLANTICUS Milne Edwards.

Sergestes atlanticus MILNE EDWARDS, Ann. Sci. Nat., xIx, 349, 1830; Hist.
Nat. Crust., 1, 428, 1837.—HANSEN, Proc. Zool. Soc. London, 1896, 949
and 951.

Sergestes frisii KROYER, Kongel. Danske Vidensk. Selsk. Skr., 5 Rekke,
naturvidensk. mathem. Afd., Iv, 235, pl. 1, fig. 1, 1859.

Sergestes pacificus STIMPSON, Proc. Acad. Nat. Sci. Phila., xu, 45 [114],
1860.

Distribution. — North and South Atlantic, North and South Pacific, and
Indian Oceans:

North Atlantic: Northward to 42° (Hansen); various localities, from
off Chesapeake Bay, 2425 fathoms, southward (Bate); Sargasso Sea, 500
to 700 meters (Ortmann).

South Atlantic: Off Monte Video, 600 fathoms (Bate); South Equa-
torial Stream (Ortmann).

North Pacific: Lat. 27%° N., long. 138° E. (Stimpson); China Sea
(Hansen); off Japan, 345 fathoms (Bate). Specimens were taken by the
Albatross, as follows:

Off Destruction Island, Washington, 516 fathoms, station 3343.
Off Sea Lion Rock, Washington, 477-636 fathoms, stations 3070, 3072,
3°73-

1 The locality ‘‘ California” attributed to Miers is probably an error, as he appa-
rently considered Puget Sound as on the Californian coast, and the specimens in the

British Museum which were received from the Smithsonian Institution were undoubt-
edly from Puget Sound. (See Proc. Zool. Soc. London, 1879, 52.)

146 RATHBUN

Off Cascade Head, Oregon, 345 fathoms, station 3347.

Off Point Arena, California, 239 fathoms, station 3349.

Off Farallone Islands, 552 fathoms, station 3162.

South of Farallone Islands, 217 fathoms, station 3105.

Off Monterey Bay, 418 fathoms, station 3127.

Off San Luis Obispo Bay, 252 fathoms, station 3195.

Off Point Conception, 233 and 284 fathoms, stations 2891, 2892.
Santa Barbara Channel, 322 and 314 fathoms, stations 2903, 2904.
Gulf of California, northwest of Tiburon Island, 145 fathoms, station 3015.

South Pacific: Lat. 15°S., long. 109° 20’ E. (Hansen); off Fijis, 315
fathoms (Bate); south of Australia, 2150 fathoms (Bate).

Indian Ocean (Hansen).

Size. —The largest specimen examined (station 3195) measures 52 mm. ;
the average length is about 4o mm. There is no indication on any of
the labels that the specimens came from the surface or from any inter-
mediate depth.

Common at surface, where no specimen exceeds 30 mm. (Hansen).
At 600, 2150, and 345 fathoms, specimens measure 38, 43, and 50 mm.
respectively (Bate).

SERGESTES sp. indet.

A single mutilated female Sergestes, without maxillipeds or trunk-legs,
was dredged by the A/datross in 417 fathoms off San Diego, California,
station 2928. It resembles S. mollis Smith. Body stout; length about 38
mm. The rostrum and anterior outline are as in that species. ‘There is
a minute hepatic spine. A strong ridge runs from the anterior margin at
the outer edge of the eye-stalk backward to the posterior margin; the
front part of the ridge is higher up than in S. mo/lis ; at its middle the
ridge gives off a weaker branch directed obliquely downward and back-
ward, and then backward to the posterior margin. Gastro-cardiac groove
deep; cervical groove present. Eyes a little shorter than in S. molits,
reaching 24 the length of the first antennular segment; cornea brown.
Antennular peduncle rather stout, first joint the longest, 124 times the
second joint; second and third subequal; notch in outer side of first seg-
ment very slight. Antennal scales broken off. Abdomen asin S. mo/iis.
Telson broken off. No hairs visible on outer margin of outer uropod.

Family PENAIDAZ.
Genus Penzus Fabricius.
PENZUS BREVIROSTRIS Kingsley.

Peneus brevirostris KINGSLEY, Proc. Acad. Nat. Sci. Phila., 1878, 98.

Penaeus californiensis HOLMES, Occas. Papers Calif. Acad. Sci., Vi, 218, pl. Iv,
figs. 64-69, Igo00.

Peneus brevirostris RATHBUN, Proc. Wash. Acad. Sci., Iv, 287, 1902.

DECAPODS 147

Distribution.—San Francisco Bay, California, to Panama; Galapagos
Islands. To a depth of 5114 fathoms.

Taken by the A/atross at the following localities: Magdalena Bay,
Lower California; off Santa Margarita Island, Lower California, 47
fathoms, station 3039; Gulf of California, 914-33 fathoms, stations 2823,
3013, 3022, 3025, 3031, 3037; also at Guaymas; Concepcion Bay,
mouth of Rio Mulege; Algodones Lagoon; and La Paz Harbor.
Panama; Panama Bay, 29%-51%4 fathoms, stations 2795, 2799, 2804,
2805.

; Genus Benthesicymus Bate.
BENTHESICYMUS TANNERI Faxon.

Benthesicymus tanneri FAXON, Bull. Mus. Comp. Zool., XXIV, 215, 1893;
Mem. Mus. Comp. Zool., XVIII, 205, pl. H, 1895.

Distribution.— From off San Diego, California, to Ecuador; Galapagos
Islands. 331-1322 fathoms.

The Albatross has collected this species at the following localities,
besides those cited by Faxon:

Off San Diego, 822 and 623 fathoms, stations 2923, 2929.
Gulf of California, 857 and 1005 fathoms, stations 3009, 3010.
Off Cape San Francisco, Ecuador, 401 and 741 fathoms, stations 2792,

2793-
Galapagos Islands, 634 and 392 fathoms, stations 2808, 2818.

Genus Gennadas Bate.
GENNADAS BOREALIS Rathbun.

Gennadas borealis RATHBUN, Proc. U.S. Nat. Mus., XxIv, 887, 1902.

Submembranous, glabrous. Rostrum reaching at least half-way along
the eye-stalk, sometimes to the cornea, armed with a single tooth, carina
very distinct nearly to the
posterior border of the cara-
pace, but sharpest in front of
the cervical groove. There
is a sharp marginal spine at
the angle of the antero-
lateral sinus.

Eyes light brown, globu- ; ns

ae Fic. 88. Gennadas borealis. 9. Station 3783. a. Side

lar, having” a speck of black view of carapace (X 2). 6. Thelycum (x 5).
pigment near their base on the outer margin of the stalk; tubercle large
and acute. Antennular flagella broken, the upper one thick at base.
The antennal scale extends beyond the antennular peduncle by about the
length of the last segment of the peduncle. Antennal flagellum as long

é

148 RATHBUN

as body. The dactylus of the external maxillipeds is subspatulate, fully
twice as long as broad. The chelz of the first pair of feet are narrow
and elongate, as in the succeeding pairs.

Exopodites of first abdominal appendages not so long as the carapace;
between the bases of these appendages is a sharp spine, equally devel-
oped in both sexes. The thelycum consists of a horizontal convex sub-
triangular plate or tubercle, placed between the third pair of legs,
followed by two transverse plates between the fourth and fifth pairs. The

anterior of these two plates is subquadrilateral,
narrowest in front; the posterior one is somewhat
fan-shaped, narrow behind, its anterior margin
rounding and with a blunt median point. The
b andricum or petasma consists of a pair of small
leaves not in contact, each of which is attached at
Fic. 89. Genxnadas borealis. , F ake 5 P
8. Station 3783. @. Petasma Its proximal end. At extremities of distal margin
(x 63). 4. Foot of first pair i :
(x4). are two lobes or teeth, the outer one of which is
curved. ‘The inner portion is partially folded to form an irregular longi-
tudinal plait. The sixth abdominal somite is carinate; it is more than
twice as long as the fifth. The telson has a small lateral spine at its
posterior fourth, two spines at the tip. The caudal swimmerets are
broken in all our specimens.

Dimensions.—Male, length of carapace and rostrum 13.6 mm., length
of abdomen on middle line 29 mm. Female, length of carapace and
rostrum 18 mm., length of abdomen on middle line 37 mm.

Distribution.—North of Rat Islands, Aleutians, 850 fathoms (A/da-
tross station 3784), 24; off Copper Island, Kamchatka, 1567 fathoms,
station 3783 (type locality), 13,29.

Very near G. parvus Bate, but differs in the longer rostrum, the
presence of a lateral spine on the telson, the greater length of the anten-
nal scale, the elongated chele of the first pair of feet, and the different
form of the thelycum and andricum.

A specimen of Gennadas was dredged by the Albatross off San Diego
in 417 fathoms, station 2928, but it is so badly mutilated that it is im-
possible to say whether it is the same species as the above or not.

Family PALINURID.
Genus Panulirus White.
PANULIRUS INTERRUPTUS (Randall).

Distribution.—Southern California to Mexico. In the National Mu-
seum are specimens ranging from Santa Barbara, California, to Rosalia
Bay, Lower California.

DECAPODS 149

Family 4X/7/D2.
Genus Axius Leach.
AXIUS SPINULICAUDA Rathbun.
Axius spinulicauda RATHBUN, Proc. U.S. Nat. Mus., xxiv, 886, 1go2.

Carapace (rostrum included) measured in the middle line, as long as
the first 5 abdominal somites, cervical groove deep, branchial groove
indistinct. Gastric region traversed by 5 carinz, all of which fade out
before reaching the cervical groove; the median extends along the basal
third of the rostrum and is armed with 4 spines just behind the line of

Fic. 90. A-xius spinulicauda. @ (X2). Station 3172. a. Carapace and
antenne. 6. Swimming-fan.

the orbits; the posterior of these spines is broken off in the unique type;
the outer carine are a continuation of the side margins of the rostrum ;
they, as well as the shorter intermediate carine, are unarmed. The sur-
face is covered with rather distant scabrous granules. In front of the
narrow median posterior lobe, the surface is compressed or pinched to
form a short smooth ridge. The rostrum reaches the middle of the
second joint of the antennular peduncle, is slightly deflexed, longitudinally
channeled, sharp-pointed, armed with 5 spines on one side and 6 on the
other.

Eyes black, of the same diameter as the stalk, not reaching the middle
of the basal antennular segment. The basal expansion of this segment
has convex side margins, an antero-external spine, and reaches just beyond
the middle of the segment; the second segment is a little longer than
the third, and the two together are shorter than the first. The entire pe-

150 RATHBUN

duncle extends to the middle of the penultimate joint of the antennal
peduncle. The outer flagellum is almost as long as the carapace and
rostrum; the inner is a fifth longer than the outer. The scaphocerite
projects as far as the middle of the second antennular segment, while
the stylocerite projects to the end of the penultimate joint of the anten-
nal peduncle; this joint is nearly 3 times as long as the last joint. The
flagellum is broken off, but the part remaining reaches beyond the anten-
nular flagella.

The outer maxillipeds reach the end of the antennal peduncle. The
first pair of chelipeds are missing. The second pair are rather narrow;
the merus has 3 spines on its lower margin; the carpus is half as long as
the merus and three fourths as long as the propodus; palm and fingers
subequal ; lower margin of merus, carpus, and propodus fringed with long
hair. ‘The third to fifth pairs of feet are slender; the lower margin of the
distal half of the propodi is fringed with long hair; the dactyli are
slender, those of the third and fourth pairs contained 3 times in their
propodi, those of the fifth pair contained 4 times in their propodi.

The abdomen is smooth above, but the pleura are sculptured and
pubescent; the telson is elongate, subquadrilateral, lateral margins
spinulous, dorsal surface with a few spines; caudal swimmerets about as
long and as broad as the telson; inner one with two outer marginal
spines, carina spinulous; outer branch with serrate outer margin, outer
carina with 1 or 2 spinules, obliquely transverse suture bordered by slender
spinules.

Dimensions.—In the unique type, a female, the length of the carapace
and rostrum along the median line is 19.6 mm., length of rostrum 3.5
mm., of abdomen 31.5 mm.

Lype locality.—Off Bodega Head, California, 62 fathoms (Albatross
station 3172).

From the other species on the west coast of North America, viz., Axius
crista-galli Faxon and A. acutifrons (Bate), both of which occur off Pana-
ma, A. sfinulicauda is readily distinguished by the five gastric carinze and
the black eyes.

Genus Calastacus Faxon.

KEY TO THE SPECIES OF CALASTACUS.

A. Carapace granulate; no spines behind those at base of rostrum,
investigatoris.
A’. Carapace not granulate; 5 rows of spines behind the rostrum,
guingueseriatus.

DECAPODS 151

CALASTACUS INVESTIGATORIS Anderson.

Calastacus investigatoris ANDERSON, Jour. Asiatic Soc. Bengal, LXV, Pt. II, 97,
1896. _ Illustrations of the Zoology of the Investigator, Crustacea, Pt.
Iv, pl. XXvV, fig. 1, 1896.—ALCOCK, Descriptive Catal. Indian Deep-Sea
Crust. Dec. Macr. and Anom. in Indian Mus., 191, Igor.

Distribution.—Taken by the Albatross, south of Sannak Islands,
Alaska, 483 fathoms, station 3210, 1 Q ; off Cascade Head, Oregon, 345
fathoms, station 3347, 1 ; off San Diego, California, 417 fathoms, sta-
tion 2928, 29.

Type locality.—Arabian Sea, off the coast of Sind, 947 fathoms
(Lnvestigator).

The antennal spines of one Californian specimen agree with Alcock’s
description, while in the Alaskan and Oregonian examples the scapho-
cerite and stylocerite are about equally produced; in the second
Californian specimen the stylocerite is broken off. There is also variability
in the spination of the upper margin of the merus of the large chelipeds;
in one individual (from California) the whole margin is armed with spines
which are proximally reduced in size; in the other three individuals ex-
amined only the distal half of the margin is armed, as in the types.

CALASTACUS QUINQUESERIATUS Rathbun.
Calastacus quingueseriatus RATHBUN, Proc. U. S. Nat. Mus., xxiv, 887,
1902.

Carapace (rostrum included) measured in the middle line, as long as
the first 5 abdominal somites ; its surface is pitted, especially on the infero-
lateral portions, and a few feeble sete spring from the pits; cervical
groove deep, branchial grooves indistinct. The rostrum reaches to the
end of the second joint of the antennular peduncle and is tipped with a
spine; its lateral margins are armed with 3 to 7 spines, and the pro-
longations of these margins extend two thirds the length of the gastric
region, are armed with 5 or 6 spines, and form a horseshoe on the cara-
pace which is open behind and outlined in front by a groove a little
posterior to the orbit. The median carina extends from the middle of the
rostrum to the rear of the horseshoe and is from 2 to 6 (usually 3) spined
atits middle. Between the median carina and the sides of the horseshoe
is another row of 3 to 5 spines. The eyes are colorless and form a
globular tip to the short eye-stalk.

Expanded basal portion of the first joint of the antennular peduncle
narrow-oval, bearing 1 or 2 marginal spinules near the tip. Stylo-
cerite of antennal peduncle reaching to distal third of the penultimate
joint; scaphocerite not reaching middle of the stylocerite. The external

152 RATHBUN

maxillipeds reach to the end of the antennal peduncle. The chelipeds
are unequal, the longer one in the male nearly as long as the body;
setose ; its inner surface and also the outer surface of carpus and manus
covered with sharp granules or short spines; upper margin spinose, also
lower inner margin of merus; lower outer margin of merus outlined with
very short blunt spines and
a single longer distal spine ;
lower margins of propo-
dus armed with dentiform
granules. Margins of pro-
podus sub-parallel. Fin-
gers shorter than palm in
, adult males, just as long
“as palm in smaller males
and in females; usually
slightly gaping at base;
occludent edges finely and
irregularly dentate. The
stouter cheliped may be
longer or shorter than the
slenderer one. In the fe-
male the chelipeds are two
thirds as long as the body.
The second pair of feet are
a little stouter and shorter
than the last three pairs;
the first four joints have a
few spines on lower mar-
gin ; thecarpus a fewspines
Fic. 91. Calastacus guingueseriatus. § (X about 3). toward the distal end ; mays
Stshon\srge: gins long-setose. Meral
joints of third and fourth pairs of legs with an infero-distal spine; last
three pairs more or less setose, especially so on the dactylus and distal
end of the propodus.

Abdomen almost smooth; there are a few sete on the sides and on
the swimming-fan. The pleura behind the first, which is subacute and
armed with a spine, are rounded and decrease in length from the second
to the fifth, the anterior margin of the third, fourth, and fifth, and lower
margin of sixth, being armed with a small spine. Telson subquadrangular,
broadly rounded behind, a median groove, a few lateral and one median
marginal spine, 2 larger dorsal spines, Inner branch of caudal swim-

DECAPODS 153

meret with a longitudinal 4- to 6-spined carina and a postero-lateral spine.
Outer branch with a transverse row of small spines near the posterior
end, and a few lateral marginal spinules also near the posterior end.
Dimensions.— Male, length of carapace and rostrum, measured in the
middle line, 28 mm., length of abdomen 41 mm.
Distribution.—Southern California, 160-388 fathoms, at the following
stations of the Albatross:

Off Point Sur, 298 fathoms, station 3187.

Off San Simeon Bay, 160 fathoms, station 3193.

Off San Luis Obispo Bay, 252, 200 fathoms, stations 3195, 3196 (type lo-
cality).

Off Point Conception, 233-284 fathoms, stations 2891, 2892, 3198.

Santa Barbara Channel, 205-280 fathoms, stations 2909, 3199-3201.

Off Anacapa Island, 388 fathoms, station 2979.

Allied to C. felix Alcock and Anderson, from which it is easily dis-
tinguished by the more numerous spines of the carapace, the longer
penultimate segment of the antennular peduncle, the squarer telson, etc.

Family CAZLZJANASSIDA2.
Genus Upogebia Leach.
UPOGEBIA PUGETTENSIS (Dana).

Harriman Expedition.—Cape Fox, Alaska (W. R. Coe).
Distribution.—¥rom southeastern Alaska to San Quentin Bay, Lower
California (Lockington).
Localities worthy of note represented in the National Museum are:
Southeastern Alaska: Kasaan Bay, Prince of Wales Island; Union Bay,
Cleveland Peninsula; Thorn Arm.
British Columbia: Nanaimo; Comox; Departure Bay; Sucia Island;
Otter Bay, Pender Island.
California: Mouth of Tia Juana River, San Diego County.

Genus Callianassa Leach.
KEY TO THE SPECIES OF CALLIANASSA.

A. Front with median tooth either obscure or not prominent. Eyes
pigmented.
B. Eye-stalks with acute and divergent extremities.

C. Median prominence of front rounded; cornea at middle of eye-
stalk; large cheliped of male very broad, the carpus very
little longer than broad, but considerably longer than palm,

californiensis.

C’, Median prominence of front subacute; cornea just behind

154 RATHBUN

middle of eye-stalk; large cheliped of male elongate, the

carpus twice, or nearly twice, as long as broad . . dongimana.

B’. Eye-stalks oblong, their inner extremities tuberculiform, and not
diverging ; cornea in front of middle of eye-stalk ; carpus of large
cheliped of male very little longer than broad, and very little
longer, sometimes even shorter, than palm. . . . . affnis.

A’. Front with a sharp and prominent median tooth.

B. Eyes not pigmented, oblong; large cheliped of male with carpus
much shorter than wide . . . . . . . © goniophthalma.

B’. Eyes pigmented, tips acute and divergent, cornea behind the mid-
dle of the stalk; large cheliped of male with carpus a little
longer than twidey 7). jen ie, sean io ist acu ne Ua oneness:

CALLIANASSA CALIFORNIENSIS Dana.
Distribution. —From Mutiny Bay, Alaska (Lockington), to mouth of
Tia Juana River, San Diego County, California (Dr. E. A. Mearns, U.S. A.).
Besides ‘published localities, there are in the National Museum speci-
mens from Departure Bay, Vancouver Island, British Columbia, and San
Diego, California.

CALLIANASSA LONGIMANA Stimpson.

Distribution. — From Vancouver Island, British Columbia (Bate), to San
Quentin Bay, Lower California (Lockington).

CALLIANASSA GIGAS Dana.
Distribution.— Puget Sound (Dana, Calman). Gulf of the Farallones,
California, 21 fathoms (A/darross station 3150), one fragmentary specimen
showing the large cheliped and portion of abdomen.

CALLIANASSA AFFINIS Holmes.
Distribution.— From Catalina Harbor to San Diego, California.

CALLIANASSA GONIOPHTHALMA Rathbun.
Plate viii.
Callianassa goniophthalma RATHBUN, Proc. U.S. Nat. Mus., XXIv, 886, 1902.
Carapace contained in the abdomen two and a third times. Rostrum
triangular, acuminate, reaching barely one third the length of eyes. Lat-
eral projections of the front shallow blunt teeth. Behind and parallel to
the front there is a sulcus which is continued back, defining the raised
area of the gastricregion. No median carina, but a slight, blunt elevation
near the posterior margin, corresponding to the strong tooth of C. cecigena.

DECAPODS 155

Eyes reaching nearly to end of first antennular segment, oblong, more
than twice as long as wide; sides subparallel; antero-external angle
rounded; antero-internal angle produced in a tuberculiform tooth, these
teeth being slightly divergent from each other; eyes without pigment.
First segment of antennular peduncle longest; second segment stouter,
and a little shorter, than the third; flagella subequal and less than twice
the length of peduncle. The antennal peduncle overreaches the antennu-
lar by the length of the last and one third of the preceding segment; the
penultimate segment is twice as long as the antepenultimate and one and
a half times as long as the ultimate; the flagellum is nearly as long as the
body. The ischium and merus of the outer maxillipeds, while wider than
the succeeding joints, are not very broad; the ischium has a lobiform
opercular dilatation on its inner or lower margin; dactylus almost circular.

The chelipeds are very unequal; the larger one is equal to the cara-
pace and first four abdominal somites combined; the ischium has a few
minute spinules at the proximal end of its lower edge; the merus may
have a small spine at the proximal end of its lower margin, or it may (as
in the largest specimen) be devoid of armature; the carpus is nearly
twice as deep as it is long and one half as long as the palm; its lower
angle is rounded; the palm is a little longer than high, and subequal in
length to the dactylus; its distal lower quarter has a few tubercles
irregularly disposed, from which proceed bunches of hair. The lower
margin of the ischium, merus, and carpus, and both upper and lower
margins of the manus are carinated. The dactylus is longer than the
pollex; the upper surface is very broad and has a rounded carina on the
inside and the outside; the outer one is tuberculate near the base of the
finger; the outer margin of the cutting surface has a broad tuberculated
tooth near the palm, and a lobe near the tip; the inner margin of the
cutting surface is tuberculate along its distal half; the pollex has a stout
tooth on the upper edge. The fingers gape widely in the male, not at
all in the female, and are beset with bunches of long hair.

The smaller cheliped is one third as wide as the larger and reaches
only to the distal third of the palm of the larger. The ischium and
merus are similar to those of the larger cheliped; the carpus is longer
than high, lower angle subacute. The palm is a little shorter than the
carpus, longer than wide. Dactylus slender, nearly twice as long as the
palm and longer than the pollex, which is armed with a single sharp
tooth at its proximal third; the fingers do not gape, as they overlap far
from the extremities.

The second, third, and fourth pairs of legs are stout and compressed,
and increase slightly in length in the order named; the fifth pair is as

156 RATHBUN

long as the third. Those of the second pair are a little longer than the
carapace, and end in a short, broad compressed hand, the edges and
outer surface of which are setose, as are also the lower border of the
merus and the upper and distal borders of the carpus. The dactyli of
the remaining pairs are almost hidden in sete.

The first of the abdominal somites is the shortest, and is one half the
length of the second, which is the longest. The lateral margins of the
first are rounded and upturned. The second is half as long as the cara-
pace; its angles are rounded, the anterior pair being carinated; the third
and fourth somites have rounded margins, the posterior angles being
setose; the fifth somite has dentiform posterior angles; the sixth has a
lateral spine in front of the middle, pointing down and back; this somite
is a little longer than wide, exclusive of the spines. The telson is nearly
as long as the sixth somite; its sides are subparallel, notched at the
anterior third; posterior angles rounded; the caudal swimmerets are
broad and rounded, the outer much the larger. The first abdominal
appendage in the @ is slender and forked, the outer branch filiform, the
inner very short, truncate, and fringed with long hair. The second to
fifth pairs are very broad, foliaceous; in the second, the outer branch is
very much larger than the inner; in the third, fourth, and fifth the two
branches are subequal; in each case the inner edge of the endopodite
bears near its middle a small styliform appendage. In the ¢ the ter-
minal joint of the first abdominal appendage is somewhat scythe-shaped,
with a lobe at the inner base of the scythe. The third to fifth pairs are
similar to those of the @? ; the second pair is much smaller, branches
subequal, inner styliform appendage larger and fringed with hair toward
the extremity.

Dimensions.—Male, station 3198, length of carapace 30.5 mm., of
abdomen 67.5 mm.; female, station 3077, length of carapace 22.2 mm.,
of abdomen 52 mm.

Distribution.—Clarence Strait, Alaska, 322 fathoms, station 3077, 42,
2 @; off Point Conception, California, 278 fathoms, station 3198 (type
locality), 1 4 (Albatross).

Closely allied to C. cecigena Alcock and Anderson, but differs in the
shorter rostrum, in the absence of spines on the second to fifth somites
of the abdomen, in the squarer telson. The shape of the first pair of
chelipeds is the same as in C. ce@cigena, but the carpus of the larger one
is without a spine, the palm is not serrated on its lower margin, its outer
surface is furnished with tubercles on the distal lower quarter, to which
also the hairs are restricted; the pollex is shorter in our species and its
tooth is nearer the middle.

DECAPODS 157

Suborder ANOMURA.
Family PAGURID.Z.

Genus Pagurus Fabricius.

PAGURUS ALASKENSIS (Benedict).
Pagurus alaskensis Benedict, Proc. U. S. Nat. Mus., XxIII, 456, text fig., 1901,
and synonymy.
Distribution.— Bering Sea (Siberian and Alaskan coasts) to Oregon;
beach to 136 fathoms.

PAGURUS ALEUTICUS (Benedict).
Pagurus aleuticus Benedict, Proc. U. S. Nat. Mus., XXIII, 460, text fig., Igor,
and synonymy.

Color.—Specimens in formalin showed the spines orange-red or crimson-
red, eye-stalks and antennal scales reddish.

Harriman Expedition.— Juneau, 50 fathoms.

Distribution.— From Bering Sea (latitude of Pribilof Islands) to Oregon ;
8 to 238 fathoms.

PAGURUS OCHOTENSIS Brandt.

Pagurus ochotensis Benedict, Proc. U.S. Nat. Mus., XXIII, 463, text fig., IgoI,
and synonymy.

Distribution. Unalaska to San Diego, California; 6 to 80 fathoms.
Okhotsk Sea (Brandt). Japan (Stimpson).

PAGURUS TRIGONOCHEIRUS (Stimpson).

Distribution.— From the Arctic coast of Alaska and Siberia southward
through Bering Sea to the Aleutian Islands and Kamchatka; 3 to 100
fathoms.

PAGURUS CAPILLATUS (Benedict).
Plate iv, fig. 3.

Harriman Expedition.—Sitka, 10 fathoms.

Distribution. — Arctic Ocean southward through Bering Straits to Kam-
chatka and California (lat. 36° 55’ N.); 3 to 240 fathoms.

PAGURUS BRANDTI (Benedict).
Plate tv, fig. 4.
Distribution.—Bering Sea (latitude of Pribilof Islands) southward to
Oregon; g to 121 fathoms.

158 RATHBUN

PAGURUS DALLI (Benedict).
Plate tv, fig. 1.

Color.—In formalin, legs banded with yellowish-red, the red coming
at the articulations, the light band in the middle of the segments.

Harriman Expedition.—Dutch Harbor, Unalaska (W. R. Coe); Ka-
diak; Berg Bay, Glacier Bay, 20 fathoms; Juneau, 20 fathoms.

Distribution.— Bering Sea (latitude of Nunivak) to Oregon; low
water to 151 fathoms.

PAGURUS RATHBUNI (Benedict),
Plate rv, fig. 2.

Distribution.—Bering Sea; east coast to Kamchatka; Okhotsk Sea,
near Robben Island; 10 to 100 fathoms.

PAGURUS TANNERI (Benedict).
Plate tv, fig. 7.

Distribution. —¥rom Mliuliuk Harbor, Unalaska, to off San Simeon Bay,
California (lat."35° 35’ N.); 50 to 559 fathoms.

PAGURUS CONFRAGOSUS (Benedict).
Plate tv, fig. 8.

Harriman Expedition.—Juneau, 50 fathoms.
Distribution.—¥rom Bering Sea (latitude of Pribilof Islands) to mouth
of Columbia River, Oregon; 41 to 238 fathoms.

PAGURUS CORNUTUS (Benedict).
Plate v, fig. 3.
Eupagurus cornutus Benedict, Proc. U. S. Nat. Mus., Xv, 12, 1892.

Distribution.— Bering Sea (lat. 55° 38’ N.) to off Queen Charlotte
Sound, British Columbia; g1 to 406 fathoms.

PAGURUS TOWNSENDI (Benedict).
Plate tv, fig. 5.
Eupagurus townsendi Benedict, Proc. U. S. Nat. Mus., Xv, 13, 1892.

Distribution.— From Pribilof Islands to Unalaska and Shumagin Bank;
284 to 625 fathoms.

DECAPODS 159

PAGURUS BERINGANUS (Benedict).
Plate v, fig. 5.

Eupagurus beringanus Benedict, Proc. U. S. Nat. Mus., Xv, 17, 1902.

Harriman Expedition.—Orca, Prince William Sound (W. R. Coe);
Yakutat Bay (T. Kincaid); Sitka, 10 fathoms; Sitka Harbor.

Distribution.— Bering Sea (latitude of Nunivak) southward along the
Aleutian Islands and coast of Alaska to Monterey, California; 5 to 19
fathoms.

I have included . xewcombet (Benedict) in P. deringanus, as it seems to
be scarcely distinct. The species varies in the sharpness of the tubercles
or spines of the chelipeds.

PAGURUS UNDOSUS (Benedict).
Plate tv, fie: 6.

Eupagurus undosus Benedict, Proc. U. S. Nat. Mus., xv, 18, 1892.
Distribution, — Bering Sea, in vicinity of Pribilof Islands; Bering Island;
Kamchatka; off Robben Island, Okhotsk Sea; 10 to 35 fathoms.

PAGURUS KENNERLYI (Stimpson).
Plate v, fig. 4.

Harriman Expedition.—Victoria, British Columbia (W. R. Coe).
Distribution.—Aleutian Islands to Washington; 9-97 fathoms.

PAGURUS SETOSUS (Benedict).
Plate v, fig. 1.

Eupagurus setosus BENEDICT, Proc. U. S. Nat. Mus., Xv, 19, 1892.

Color.—In formalin, the legs are banded with crimson-red, hair dirt
color. -

Harriman Expedition.—Kadiak; Juneau, 50 fathoms. Not known
previously north of Sitka.

Distribution.—F¥rom Kadiak, Alaska, to Santa Catalina Island, Cali-
fornia; 50-266 fathoms,

PAGURUS HIRSUTIUSCULUS (Dana).

Color.—In formalin, body and legs pinkish-red. Segments of second
and third pairs of feet, white at distal ends; dactyli striped longitudinally
with pinkish and white.

160 RATHBUN

Harriman Expedition.—Orca; Yakutat; Juneau and Farragut Bay;
Sitka, 10 fathoms; Sitka Harbor; Wrangell (T. Kincaid). Also taken
by Dr. Coe at Dutch Harbor, Unalaska; Kadiak; Virgin Bay; Hot
Springs; Tongass Village.

Distribution.—St. Paul Island, Pribilofs (T. Kincaid); Aleutian
Islands to southern California (Holmes). Specimens are in the National
Museum from as far south as Wilmington, California. East Cape,
Siberia; Bering Island; Copper Island, Kamchatka; Japan (Stimpson).
Low tide to 17 fathoms.

PAGURUS SAMUELIS (Stimpson).
Plate v, fig. 7.

Distribution.—Sitka, Alaska, to San Diego, California. Japan (Stimp-
son, Ortmann; doubted by Holmes).

PAGURUS MIDDENDORFFII Brandt.

Distribution.—Sitka (Brandt); Puget Sound (Calman); Bering Island
(L. Stejneger) ; Castries-Bai, Siberia (Ortmann); Okhotsk Sea (Brandt) ;
Japan (Stimpson).

PAGURUS MINIMUS Holmes.

Distribution.— Off San Diego, California, 30 fathoms (Holmes).

PAGURUS GRANOSIMANUS (Stimpson).
Plate v, fig. 8.

Color.—Deep red, with white tubercles on the chele, and darker red
spines on the carpus and merus of the chelipeds. Ambulatory legs red,
with small white spots; dactyli with a broad white band at middle.

Harriman Expedition.—Sitka Harbor. Collected by Dr. Coe at
Yakutat; Hot Springs; Sitka; Cape Fox; Tongass Village.

Distribution.—Unalaska to Ensenada, Lower California. Beach to 15
fathoms.

PAGURUS HEMPHILLII (Benedict).
Plate v, fig. 9.

Distribution. —Humboldt County to Monterey, California.

PAGURUS TENUIMANUS (Dana).

Color.—In formalin, claws yellowish-red, spines lighter. Dactyli and
propodi of second and third pairs of legs spotted with darker red; carpal
joints with two broad longitudinal stripes of red on upper side.

DECAPODS 161

Harriman Expedition.—Kadiak; Sitka, 10 fathoms; Sitka Harbor.
Distribution.—¥rom Aleutian Islands and Alaska Peninsula to Strait
of Fuca and Puget Sound. Beach.to 123 fathoms.

PAGURUS MUNITUS (Benedict).
Plate: ¥, ig,.-2.

Color.—In formalin, yellowish red. Second and third pairs of legs of
the same color, with white spots which tend to coalesce.

flarriman LExpedition.—Juneau, 50 fathoms (an extension of the
range).

Distribution.— Bering Sea (lat. 56° 56’ N.) southward and eastward
to Portlock Bank and Juneau, Alaska; 20 to 138 fathoms.

PAGURUS GILLI (Benedict).
Plate v; fig. ro.

Eupagurus gilli BENEDICT, Proc. U. S. Nat. Mus., Xv, 20, 1892.

Color.—In formalin, orange-red. Ambulatory legs more of an orange-
vermilion, as are also the bases of the antenne. Pits on legs whitish.

FHlarriman Expedition.—Dutch Harbor, Unalaska; Kadiak; Orca,
Prince William Sound, shore on rocks (W. R. Coe); Sitka.

Distribution.—Bering Island, Aleutian Islands; Alaska Peninsula
southward to Revillagigedo Island, southeastern Alaska. Low water to
60 fathoms.

PAGURUS CALIFORNIENSIS (Benedict).
Distribution. — From Santa Catalina Island, California, to Gulf of Cali-
fornia; 8 to 58 fathoms.
PAGURUS SPLENDESCENS Owen.

fTarriman Expedition.—Juneau, 50 fathoms.

Distribution.—From Arctic coast of Alaska (at Point Barrow) west-
ward through Bering Sea to Kamchatka and Washington. Below low
water to 225 fathoms.

Genus Paguristes Dana.
PAGURISTES TURGIDUS (Stimpson).

Distribution.— Vancouver Island, British Columbia, to Santa Catalina
Island, California.

PAGURISTES PARVUS Holmes.

Distribution.—Whites Point near San Pedro, California (Holmes).

162 RATHBUN

PAGURISTES BAKERI Holmes.

Distribution.—San Diego, California (Holmes).

Genus Holopagurus Holmes.
HOLOPAGURUS PILOSUS Holmes.
Distribution.— Off San Diego, California, 25 fathoms (Holmes).

Genus Parapagurus Smith.
PARAPAGURUS MERTENSII (Brandt).
Plate v, fig. 6.

Distribution.—I\slands of Kadiak and Atka; Nootka Sound, west
coast of Vancouver Island; North California; Kamchatka (Brandt).
In the National Museum are specimens from the coast of California
between Lobos Rocks (lat. 36° 26’ 40’ N.) and southeast of San Nicholas
Island (lat. 33° 00! 30” N.), depth 77 to 266 fathoms. The depth and
locality quoted by Holmes for the specimen examined by him are
probably erroneous.

PARAPAGURUS 5p. indet.
Distribution.—From southeast of Chirikof Island, Alaska, to off San
Diego, California; 414 to 877 fathoms.
Family Z7J7HODIDZ.
Genus Hapalogaster Brandt.
HAPALOGASTER CAVICAUDA Stimpson.
Distribution.— California: Cape Mendocino to Monterey. Common
under rocks at low tide.
HAPALOGASTER DENTATA (de Haan).
Lomis dentata DE HAAN, Fauna Japon., 219, pl. XLVIII, fig. 2, and pl. Q,
1849.
Hapalogaster dentatus STIMPSON, Proc. Acad. Nat. Sci. Phila., x, 232, 1858.
Distribution.—“ Colonie rosso-americane” (Schalfeew); Japan (de
Haan, Stimpson).
HAPALOGASTER MERTENSII Brandt.

Flarriman Expedition.—Dutch Harbor, Unalaska (W. R. Coe); Fox
Island (T. Kincaid); Yakutat and Sitka (W. R. Coe).

DECAPODS 163

Distribution.—From Atka, Aleutians, eastward and southward to
Puget Sound; to a depth of 19 fathoms.

HAPALOGASTER GREBNITZKII Schalfeew.

Harriman Expedition.— Dutch Harbor, Unalaska (W. R. Coe); Popof
Island; Orca (T. Kincaid); Sitka (W. R. Coe), extending the known
range.

Distribution.—¥rom Bering Sea off the Pribilofs and Cape Newenham
southward along the Aleutian Islands to Sitka, Alaska; Bering Island.

Genus Dermaturus Brandt.
DERMATURUS MANDTII Brandt.

Harriman Expedition.— Dutch Harbor, Unalaska; Popof Island; Sitka.
Distribution.—Pribilof Islands; Aleutian Islands and Alaska Peninsula
to Sitka; Bering Island; 6 to 40 fathoms.

Genus dignathus Benedict.
CEDIGNATHUS INERMIS (Stimpson).

Hapalogaster inermis STIMPSON, Ann. Lyc. Nat. Hist. N. Y., vi, 243, 1860.

Hapalogaster Brandti SCHALFEEW, Mel. biol., XII, 330, 1892, figs. 1 and 5b,
of the Bull. Acad. imp. sci. St. Pétersbourg, XXXv, 336, 1892.

Cdignathus gilli BENEDICT, Proc. U. S. Nat. Mus., Xv, 487, 1894.

G@dignathus inermis HOLMES, Occas. Papers Calif. Acad. Sci., VII, 119, 1900,
and synonymy.

Gdignathus Brandti HOLMES, of. cit., 118, and synonymy.

I think there is little doubt that Schalfeew’s species is identical with
Stimpson’s. That the number of squame on the carapace is variable
is evidenced in the small series I have examined. Stimpson’s type was
smaller than any examples in the National Museum.

Distribution.— Unalaska to Pacific Grove, California.

Genus Acantholithodes Holmes.
ACANTHOLITHODES HISPIDUS (Stimpson).

Distribution.—Vancouver Island, British Columbia, to Monterey, Cali-
fornia. To a depth of at least 16 fathoms.

Genus Placetron Schalfeew.
PLACETRON WOSNESSENSKII Schalfeew.
Plate vi, fig. x.

Placetron Wosnessenskii SCHALFEEW, Mel. biol., XIII, 333, 1892, fig. 6, of the
Bull. Acad. imp. sci. St. Pétersbourg, XXXV, 339, 1892.
Lepeopus forcipatus BENEDICT, Proc. U.S. Nat. Mus., xvu1, 488, 1894.

164 RATHBUN

Distribution.— Aleutian Islands to British Columbia: Unalaska; west
of Amaknak Island, 60 fathoms; northwest of Unimak Island, 43
fathoms (Addatross station 3262); Kadiak (Schalfeew); Parry Passage,
Graham Island, British Columbia.

Genus Phyllolithodes Brandt.
PHYLLOLITHODES PAPILLOSUS Brandt.
Harriman Expedition.—Dutch Harbor, Unalaska. Hitherto not
known west of Kadiak.

Distribution.—Unalaska to Monterey, California. To a depth of at
least 16 fathoms.

PHYLLOLITHODES BICORNIS (Bate).
Distribution.— Esquimalt Harbor, British Columbia, 10 fathoms (Bate).
Perhaps not distinct from P. papillosus.

Genus Cryptolithodes Brandt.

CRYPTOLITHODES TYPICUS Brandt.
Distribution.—Belkofski Bay, Alaska, to Monterey, California. Be-

tween high and low tide marks.
CRYPTOLITHODES SITCHENSIS Brandt.
Distribution. —From Sitka, Alaska, to Pacific Grove, California.

CRYPTOLITHODES BREVIFRONS Miers.
Distribution. —Vancouver Island, British Columbia.
Perhaps not a valid species.

Genus Lopholithodes Brandt.

LOPHOLITHODES MANDTII Brandt.
Distribution.—From Sitka, Alaska, to Monterey, California.

LOPHOLITHODES FORAMINATUS (Stimpson).
Distribution.—¥rom Victoria, British Columbia, to near San Francisco
and the Farallones, California.

Genus Rhinolithodes Brandt.

RHINOLITHODES WOSNESSENSKII Brandt.

Rhinolithodes wossnesenskit BRANDT, Bull. Phys.-Math. Acad. Impér. Sci. St.
Petersbourg, VII, 174, 1849 (typo. error for wosnessenskit).
pgs Wosnessenskit NEWCOMBE, Bull. Nat. Hist. Soc. Brit. Col., 1893,
29. pl.10t,

DECAPODS 165

Distribution.—Kadiak ; Port Etches, 12 to 18 fathoms; Sitka; Crescent
City, California.
Genus Pristopus Benedict.

PRISTOPUS VERRILLI Benedict.

Pristopus verrilli BENEDICT, Proc. U. S. Nat. Mus., XviI, 486, 1894.

Distribution.—Off Pribilof Islands, 688 fathoms, station 3501; off
Sea Lion Rock, Washington, 859 fathoms, station 3075; off Cortez
Bank, California, 984 fathoms, station 2919 (A/batross).

Genus Leptolithodes Benedict.
LEPTOLITHODES MULTISPINA Benedict.

Leptolithodes multispinus BENEDICT, Proc. U.S. Nat. Mus., xvi, 484, 1894.

Distribution.—Off Shumagin Bank, Alaska, 625 fathoms, station
3338; Off Queen Charlotte Islands, British Columbia, 876 fathoms,
station 2860; off Sea Lion Rock, Washington, 685 fathoms, station 3071 ;
off San Diego, California, 822 fathoms, station 2923 (Advatross).

Genus Lithodes Latreille.
LITHODES RATHBUNI Benedict.

Distribution.— Off San Simeon Bay, California, 211 fathoms.

LITHODES CALIFORNIENSIS Benedict.
Distribution.— Off Santa Cruz Island, California, 155 fathoms.

LITHODES BREVIPES Milne Edwards and Lucas.

Lithodes brevipes MILNE EDWARDS and Lucas, Arch. Mus. Hist. Nat. Paris,
II, 465, pls. XXIV-XXVII, 1841.—BENEDICT, Proc. U. S. Nat. Mus., XVII,
484, 1894.

Lithodes camtschaticus RICHTERS, Abh. Senck. Natur. Ges., Frankfurt, XIII,
404, plate, figs. 9, 10, 1884.

Distribution.—Bering Island; St. Paul Island, Pribilofs; off the
Pribilofs, 25 to 47 fathoms; Unalaska; Kamchatka; Okhotsk Sea.

LITHODES CAMTSCHATICUS (Tilesius).

Maja camtschatica TILEstus, Mém. Acad. Impér. Sci. St. Pétersbourg, v, 1812,
p- 336, pls. v and vi (1815).
ee camptschensis LATREILLE, Cuvier’s Régne Animal, 2nd ed., IV, p.
5, 1829.
Lithodes spinosissimus BRANDT, Bull. Phys.-Math. Acad. Impér. Sci. St.
Pétersbourg, VII, 172, 1849 (young).—STIMPSON, Boston Jour. Nat. Hist.,
VI, 478,.1857.

166 RATHBUN

Harriman Expedition.—Kadiak; Orca. Young specimens only.
Distribution. — Bering Sea (from Norton Sound) to Aleutian Islands to
Orca, Prince William Sound, 5 to 49 fathoms. Kamchatka; Okhotsk Sea;
Japan.
LITHODES AEQUISPINA Benedict.
Lithodes equispinus BENEDICT, Proc. U.S. Nat. Mus., XviI, 481, 1894.

Distribution. — Bering Sea (off the Pribilof Islands and north of Un-
alaska), 150 to 406 fathoms.

LITHODES COUESI Benedict.

Lithodes couest BENEDICT, Proc. U. S. Nat. Mus., xvil, 481, 1894.

Distribution.—North of Unalaska and off Shumagin Bank, 399 and
625 fathoms.

Family GALA THEID.
Genus Pleuroncodes Stimpson.
PLEURONCODES PLANIPES Stimpson.

Distribution.—From go miles southwest of San Francisco, California,
to 150 miles southwest of Cape St. Lucas, Lower California.

Genus Galathea Fabricius.
GALATHEA CALIFORNIENSIS Benedict.

Galathea californiensis BENEDICT, Proc. U. S. Nat. Mus., XXvVI, 247, fig. 1,
1902.

Distribution.— Off Santa Cruz Island, California, 150 fathoms.

Genus Munida Leach.
MUNIDA QUADRISPINA Benedict.

Distribution.— From Sitka, Alaska, to San Diego, California, 50 to 559
fathoms.
Probably the species designated by Owen as MZ. gregaria.

Genus Munidopsis Whiteaves.
MUNIDOPSIS HYSTRIX Faxon.

Munidopsis hystrix F AXON, Bull. Mus. Comp. Zool., XXIV, 183, 1893; Mem.
Mus. Comp. Zool., XVIII, 89, pl. XIX, fig. 1, 1895.

Distribution.—¥rom off Anacapa Island, California, to off Acapulco,
Mexico; 364 to 680 fathoms.

DECAPODS 167

MUNIDOPSIS BERINGANA Benedict.

Munidopsis beringana BENEDICT, Proc. U. S. Nat. Mus., XXVI, 279, fig. 23,
1902.

Distribution.— Bering Sea, southwest of Pribilof Islands, 1771 fathoms.

MUNIDOPSIS VERRILLI Benedict.

Munidopsis verrilli BENEDICT, Proc. U. S. Nat. Mus., XXVI, 291, fig. 34,
1902.

Distribution.— California: Off Cortez Bank and off San Diego, 984
and 822 fathoms.

MUNIDOPSIS QUADRATA Faxon.

Munidopsis gquadrata FAXON, Mem. Mus. Comp. Zool., XVIII, 97, pl. XXIII,
fig. I, 1895, and synonymy.

Distribution.—From off Destruction Island, Washington, to Tres
Marias Islands, Mexico. Usually between 339 and 859 fathoms; also
at 47 and 60 fathoms, off Wilmington, California, and Los Coronados
Islands, respectively.

MUNIDOPSIS ASPERA (Henderson).

Elasmonotus asper HENDERSON, Ann. Mag. Nat. Hist. (5), XvI, 416, 1885;
Rept. Challenger Anomura, 163, pl. XIX, fig. 4, 1888.

Munidopsis aspera F AXON, Bull. Mus. Comp. Zool., xxiv, 188, 1893; Mem.
Mus. Comp. Zool., XVIII, 96, 1895.

Distribution.—From north of San Clemente Island, California, to
Straits of Magellan; Galapagos; 69 to 782 fathoms. Brazil, 1500 fathoms
(Henderson).

Family AZBUNEIDZ.

Genus Blepharipoda Randall.
BLEPHARIPODA OCCIDENTALIS Randall.

Blepharipoda occidentalis RANDALL, Jour. Acad. Nat. Sci. Phila., VIII, 131,
pl. vI, 1839.

Distribution. —From San Francisco, California (D. S. Jordan), to San
Quentin Bay and Rosalia Bay, Lower California.

Genus Lepidopa Stimpson.
LEPIDOPA MYOPS Stimpson.

Distribution.—From San Diego, California, to Cape St. Lucas, Lower
California.

168 RATHBUN

Family H/PPIDA.
Genus Emerita Gronovius, Benedict.
EMERITA ANALOGA (Stimpson).

Hippa analoga STIMPSON, Proc. Boston Soc. Nat. Hist., v1, 85, 1857. For
synonymy, see Holmes.

Distribution.— According to Holmes, this species extends from Oregon
to Panama. The collections in the National Museum range from
Drake Bay, California, to San Bartolomé Bay, Lower California, and
from Peru to Chile. The intermediate region (from Lower California to
Panama) is represented by £. emerita (Fabricius), which occurs also on
the Atlantic side of tropical America.

Family PORCELLANID.
Genus Petrolisthes Stimpson.
PETROLISTHES CINCTIPES (Randall).
Distribution. —From Vancouver Island, British Columbia, to the Gulf
of California.
PETROLISTHES ERIOMERUS Stimpson.
Distribution.—From Humboldt Bay to Pacific Grove, California.

Genus Pachycheles Stimpson.

PACHYCHELES RUDIS Stimpson.
Plate vi, fig. 6.

Distribution.—British Columbia to Santa Catalina Island, California ;
Lower California (Lockington).

PACHYCHELES PUBESCENS Holmes.

Distribution.— California, from Humboldt County to the Farallones.

Suborder BRACHYURA.
Family DORIPPID:.
Genus Clythrocerus A. Milne Edwards and Bouvier.

CLYTHROCERUS PLANUS (Rathbun).
Plate 1x, fig. 4.

Cyclodorippe plana RATHBUN, Amer. Nat., XXXIV, 519, 1900.

DECAPODS 169

Carapace subcircular, a little broader than long; dorsally flat, finely
granulate, the granules larger toward the outer margin; gastric and
cardiac-intestinal regions bounded by deep grooves. ‘The front is occu-
pied by two triangular lobes, each tipped with a blunt tooth, and sepa-
rated from each other by a broad V-shaped sinus, which is prolonged on
the dorsal surface by a broad, shallow depression continued to the gastric
region; outer margin of each lobe slightly concave. Outer orbital tooth
narrow, blunt, well marked. A little in front of the middle of the lateral
margin is a tooth somewhat larger than the orbital, and directed forward
and slightly outward. At one third the distance from the orbital to this
branchial tooth is a much smaller triangular tooth directed outward.
The antennule are hidden under the carapace. ‘The basal segment of
the antennz is moderately enlarged. The anterior end of the buccal
cavern and of the merus of the outer maxillipeds project slightly in front
of the median sinus of the front.

Chelipeds equal, rather short, stout, coarsely scabrous-granulate. Merus
stout, unarmed. Carpus broader than long, having a shallow, plate-like
blunt tooth along its outer surface, a short blunt tooth at inner angle.
The palm is about as broad as long, bears a stout blunt spine or tooth
on the outer side at the articulation of the carpus, and a lower, less
conspicuous tooth at the articulation of the fingers; two feebly marked
carine, one connecting the two teeth, the other lower down. The digits
are longer than the upper margin of the palm, bent down, not gaping,
the pollex much stouter than the dactylus, edges denticulate; the pollex
has a superior longitudinal groove; its inner superior margin is subacute
and continuous with that of the palm. Second pair of feet about twice
as long as the carapace, dactylus longer than propodus. ‘Third pair
exceeding the second by half the length of the dactylus. Last two pairs
of feet missing.

Color.—Carapace speckled with small black spots, in alcohol.

Dimensions.— Length of carapace of male, to end of horns, 3.4 mm.,
width 3.7 mm. Length of female 2.8 mm., width 3 mm.

Type locality.—Southern California, either at Catalina Harbor or
Monterey (probably the former); W. H. Dall; 14, 19 (Cat. No.
14,256).

Additional specimen.—A female corresponding to the above was
dredged in 1863 at Catalina Island by Dr. J. G. Cooper. With it 1s a
male (Plate 1x, fig. 5) a little larger and as long as wide (4.3 mm.
long to end of horns and 4.2 mm. wide), which differs notably in the
following particulars: The frontal lobes are prolonged in cylindrical
blunt spines; the upper orbital margin has an inner rectangular sinus,

170 RATHBUN

while the fissure next the outer tooth is larger than in the typical form;
the first tooth of the lateral margin is nearer the posterior tooth than it is
to the orbital tooth; the surface is more uneven, and more coarsely
granulate. The legs are absent. I think the differences are not those of
age, and indicate a distinct species, to which, however, I hesitate to give
a name on account of the poor condition of the specimen.

Family ZEUCOSHDA.
Genus Randallia Stimpson.
RANDALLIA ORNATA (Randall).

Distribution.—From Mendocino County, California, to Magdalena Bay,
Lower California; 534 to 51 fathoms.

Varies greatly in the prominence of the granules on the carapace.

RANDALLIA BULLIGERA Rathbun.
Distribution.— Off San Diego, California, 30 fathoms, and Magdalena
Bay, Lower California, 12 fathoms.
Genus Philyra Leach.
PHILYRA PISUM de Haan.
Distribution.— Puget Sound; Japan; Korea; Philippine Islands.

Only one specimen of this species has been taken in Puget Sound, as
noted by Calman.}

Family CAZAPPIDA.
Genus Platymera Milne Edwards.
PLATYMERA GAUDICHAUDII Milne Edwards.

Distribution.—From the Farallones, California, to Panama; Chile;
26 to 218 fathoms.

Family PARTHENOPID.?
Genus Heterocrypta Stimpson.
HETEROCRYPTA OCCIDENTALIS (Dana).

Distribution.—From Gulf of the Farallones to San Diego, California,
and Los Coronados Islands; 13 to 36 fathoms.
1Ann. N. Y. Acad. Sci., x1, 262, 1898.

2 Leiolambrus punctatissimus (Owen) is not known to occur north of latitude
27° 45’ N.

DECAPODS 171

Family JZ47/DA.
Genus Podochela Stimpson.
PODOCHELA HEMPHILLII (Lockington).
Plate:x;. fig. 2;

Distribution, —¥From San Luis Obispo, California, to Gulf of California.
Recorded from 8 to 47 fathoms, and perhaps occurs in shallower water.

Genus Anasimus A. Milne Edwards.
ANASIMUS ROSTRATUS Rathbun.
Plate x, fig. 4.

Distribution.—Southern California (either at Catalina Harbor or
Monterey) ; northwest of Cerros Island, Lower California, 58 fathoms.

Genus Erileptus Rathbun.
ERILEPTUS SPINOSUS Rathbun.
Plate x,. fig. 3,

Distribution.—Off San Diego, California, 36 fathoms; also southern
California (either at Catalina Harbor or Monterey). (W. H. Dall.)

Genus Inachoides Milne Edwards and Lucas.
INACHOIDES MAGDALENENSIS Rathbun.
Distribution.—From Wilmington, California, to Gulf of California;
3% to 66 fathoms.
Genus Oregonia Dana.

OREGONIA GRACILIS Dana.

Harriman Expedition.—Dutch Harbor, Unalaska; Juneau; Sitka.
Distribution.—From Bering Sea at Nunivak and Bering Island, to
northern California; 5 to 135 fathoms.

OREGONIA BIFURCA Rathbun.
Plate vi, fig. 5.

Oregonia bifurca RATHBUN, Proc. U. S. Nat. Mus., XxIv, 885, 1902.

Carapace about three fourths as wide as long (rostrum included) ;
width between tips of postorbital spines two thirds of the branchial width.
Branchio-hepatic constriction very slight. Surface of carapace covered
with tubercles from which proceed long yellow hairs, curved at the tips,
making a tolerably thick coating. Rostrum not divided quite to the base ;

172 RATHBUN

horns slender, divergent, their outer margins almost longitudinal, their
length along the inner margin about one eighth of the entire length of
the carapace. Supraocular eave extending laterally not quite so far as
in O. gracilis, Postocular spine narrow, as in O. gracilis, and directed
a little more forward than in that species. The interantennular spine is
curved a little more forward than in O. gracilis. Eyes a little stouter than
in that species. Basal segment of antenna armed along its outer margin
with one stout blunt subterminal spine and three small spines, one of which
is at the anterior angle, and along its inner margin with one or two
tubercles. Outer maxillipeds asin O. gracilis. Lower surface pubescent.

Chelipeds of the female of moderate size, one and a half times as long
as carapace, subequal, shaped much as in O. gracilis. Inner surface of
merus bordered by short blunt spines. Propodus slightly longer than
merus, fingers and palm subequal, margins of palm subparallel, fingers
fitting close together, denticulate. Ambulatory legs slender, diminishing
gradually in length from the first to the fourth pair, the third pair being
the length of the female cheliped. The dactyli are contained about one
and a third times in their propodi. Chelipeds and ambulatory legs covered
with long yellow hair, mixed with shorter and finer hair.

Dimensions.—Length of carapace and rostrum 26.5 mm., width of
carapace 19.6 mm., width between tips of postorbital spines 13 mm.,
length of horn measured along the inner edge 3.4 mm., distance between
tips of horns 3.5 mm.

Type locality.—North of Rat Islands, Aleutian Chain, 270 fathoms
(Albatross station 3785), one adult female.

This species, by reason of the character of its orbits, maxillipeds, and
trunk-legs, falls into the genus Oregonia, but differs from O. gracilis in its
spreading horns, greater anterior breadth, different ornamentation of the
basal antennal segment, etc.

Genus Dasygyius Rathbun.
DASYGYIUS TUBERCULATUS (Lockington).
Platex.fos.44.5 3a:

Distribution.—From Alamitos Bay, Cali-
fornia, to Panama Bay, including Gulf of
California; 4 to 33 fathoms.

A fine series of specimens of this species
was collected in San Diego Bay, 6 fathoms,
by the A/satross (station 3620); the largest
Fic. 92. Dasygyius tuberculatus. §. Male measures 23 mm. long and 17.2 mm.

Station 3030. @. Maxilliped, much en- ;
larged. 4. Side view of carapace (x 13). wide.

DECAPODS 173

Genus Epialtus Milne Edwards.
EPIALTUS PRODUCTUS Randall.

Distribution. —From British Columbia! to Lower California.

EPIALTUS NUTTALLII Randall.

Distribution.—From Santa Barbara, California, to Ballenas Bay, Lower
California.

EPIALTUS BITUBERCULATUS Milne Edwards.

Epialtus bituberculatus MILNE EDWARDS, Hist. Nat. Crust., 1, 345, pl. Xv,
fig. 11, 1834.—RATHBUN, Bull. U. S. Fish Comm., 1900, vol. 2, p. 60
(1901), and synonymy.

Epialtus bituberculatus ? var. bermudensis VERRILL, Trans. Conn. Acad. Sci.,
KE. FOr), 1; fey. Py TOOT.

Distribution.—Southern California (either Catalina Harbor or Monte-
rey) to Chile; Indian River, Florida, to Rio de Janeiro; Bermudas.

Genus Mimulus Stimpson.
MIMULUS FOLIATUS Stimpson.

Distribution.—From Unalaska to Mazatlan, Mexico (A. Milne Ed-
wards).

Genus Pugettia Dana.
PUGETTIA GRACILIS Dana.

Harriman Expedition.—Dutch Harbor, Unalaska (W. R. Coe); Kadiak
(Coe); Fox Island (Kincaid); Virgin Bay (Coe); Orca; Yakutat; Sitka,
to fathoms; Tongass Village (Coe).

Distribution.— From Aleutian Islands to southern California (Holmes) ;
low tide to 40 fathoms.

PUGETTIA RICHII Dana.

Distribution.— British Columbia to San Diego, California; low tide.

PUGETTIA DALLI Rathbun.
Plate 11, figs. 1, 1a.

Distribution.—From San Pedro, California, to San Geronimo Island,
Lower California; 614-30 fathoms.

1The specimen recorded from Kiska Harbor, Alaska, was wrongly determined ; it
was overgrown with sponge and proved to be Pugettia gracilis.

174 RATHBUN

Genus Chorilia Dana.

CHORILIA LONGIPES Dana.

Distribution.—From 57° north latitude, off Kadiak, Alaska, to 32°
north latitude, off San Diego, California; 27-603 fathoms.

Genus Chionecetes Kroyer.

CHIONCECETES OPILIO (0. Fabricius).

Harriman Expedition.—Yakutat, in cod’s mouth; Juneau, 20-50
fathoms.

Distribution.—From British Columbia northward to Bering Sea, thence
eastward to Greenland, thence southward to Casco Bay, Maine; Kam-
chatka; Okhotsk Sea (off Robben Island); 3-206 fathoms.

CHIONC@CETES TANNERI Rathbun.

Distribution.—From Bering Sea to off San Diego, California (lat. 32°
17/ N.); 29-1625 fathoms.

Genus Hyas Leach.

HYAS COARCTATUS Leach.

Distribution.—Circumpolar; Bering Sea; Kamchatka; Okhotsk Sea,
as far south as Robben Island; Atlantic coast of America southward to
off Cape Hatteras, North Carolina; northern Europe; 7-906 fathoms.

HYAS LYRATUS Dana.

Harriman Expedition.—Berg Bay, Glacier Bay; Juneau, 50 fathoms.
Distribution.— From Bering Sea (off Kululak Bay and at the extreme
end of the Aleutian Islands) to Puget Sound; 5-110 fathoms.

Genus Pelia Bell.
PELIA TUMIDA (Lockington).
Distribution.—Santa Catalina Island, California, to Magdalena Bay,
Lower California.
PELIA PACIFICA Milne Edwards.

Distribution.—From Santa Catalina Island, California, to Bay of
Panama; 10-40 fathoms,

DECAPODS 175

Genus Loxorhynchus Stimpson.
LOXORHYNCHUS GRANDIS Stimpson.

Distribution.— From the Farallones and San Francisco to San Diego,
California; 614-68 fathoms.

LOXORHYNCHUS CRISPATUS Stimpson.

Distribution.—From the Farallones to San Diego, California; 4-53
fathoms.
Genus Scyra Dana.

SCYRA ACUTIFRONS Dana.

Distribution.—¥rom Kadiak, Alaska, to San Diego, California. Low
tide to 40 fathoms.

Genus Rhodia Bell.
RHODIA PARVIFRONS (Randall).

ferbstia parvifrons RANDALL, Jour. Acad. Nat. Sci. Phila., VIII, 107, 1839.
— HOLMES, Occas. Papers Calif. Acad. Sci., VII, 38, 1900.

flerbstia (Herbstiella) camptacantha HOLMES, of. cit., p. 37, and synonymy.

Rhodia parvifrons RATHBUN, Amer. Nat., XXXIV, 511, 1900.

Distribution.—From San Pedro, California, to Acapulco, Mexico;
30-40 fathoms.}
Family CANCRIDZ.
Genus Cancer Linnzus.
CANCER PRODUCTUS Randall.

Harriman Expedition.—¥ox Island (T. Kincaid).
Distribution.—From Kadiak, Alaska, to Magdalena Bay, Lower Cali-
fornia.

CANCER AMPHICETUS Rathbun.
Plate vi, fig. 3.

Trichocarcinus dentatus M1ERS, Proc. Zool. Soc. London, 1879, p. 34; not
Cancer dentatus BELL, 1835; nor C. dentatws HERBST, 1785.
Cancer amphietus RATHBUN, Proc. U. S. Nat. Mus., XXI, 582, 1898.

Distribution.—From San Diego Bay, California, to Magdalena Bay,
Lower California; Japan; Korea; 1114-169 fathoms.
Size.—Specimens from Magdalena Bay, in the Museum of Compara-

1 Libinia emarginata Leach, included in Holmes’s list on the authority of Randall, I
have omitted, because so many errors have been discovered in the localities cited by
Randall that the existence of this species on the west coast of America needs verifica-
tion.

176 RATHBUN

tive Zoology, measure as follows: Male, length 24.2 mm., width 33.4
mm.; female, length 18.7 mm., width 25 mm.

CANCER ANTENNARIUS Stimpson.

Distribution.—Queen Charlotte Sound, British Columbia, to Magda-
lena Bay, Lower California. Shallow water to 7 fathoms.

CANCER GIBBOSULUS (de Haan).

Corystes (Trichocera) gibbosula DE HAAN, Fauna Japon., 45, pl. 1, fig. 4, pl.
XMM hie: 35.5635.

Trichocarcinus gibbosulus MIERS, Proc. Zool. Soc. London, 1879, p. 34.

Cancer gibbosulus RATHBUN, Proc. U.S. Nat. Mus., XXI, 581, 1898.

Distribution.—From Granite Cove, Port Althorp, Alaska, to San
Geronimo Island, Lower California; Japan. Shallow water to 40
fathoms.

CANCER ANTHONYI Rathbun.
Plate v1, fig. 2.
Cancer anthonyi RATHBUN, Proc. Biol. Soc. Wash., XI, 111, 1897; Amer.
Nat., XXXIV, 134, 1900.

Distribution.—Long Beach, California (H. N. Lowe); Playa Maria
Bay, Lower California.

Size.—Among the specimens taken by Mr. Lowe from the wharf at
Long Beach is a male somewhat larger than the type, measuring 52.1
mm. long and 77.5 mm. wide.

CANCER JORDANI Rathbun.
Plate vi, fig. 4.

Cancer jordanit RATHBUN, Amer. Nat., XXXIV, 133, 1900.

A small species allied to C. magister and C. antennarius.

Female.— Length of carapace one and a fourth times width. Surface
hairy and covered with small crowded scabrous granules. Regions indi-
cated by narrow and shallow depressions. Teeth of front and lateral
border not thickened as in C. antennarius ; inner orbital tooth very slightly
produced ; margin of upper orbital tooth almost transverse, scarcely denti-
form. Lateral teeth separated to their bases, tips spiniform and curved
forward; second, fourth, sixth, and eighth teeth smaller than the others
(the outer orbital tooth reckoned as the first); ninth tooth scarcely more
prominent than eighth. No postero-lateral tooth. Basal antennal tooth
and the adjoining tooth of the orbital margin acute. Movable part of
antenne half as long as carapace. The outer maxillipeds overlap con-

DECAPODS 177

siderably the basal joint of the antennze; merus longer than wide. The
palms of the chelipeds have 2 superior and 5 external carine fringed
with hair. On the prehensile edges of the fingers the dark color runs
from the proximal end of the fingers nearly to the tip, but on the outer
edges the dark color begins near the middle of the fingers.

Dimensions.—Female, length 15.5 mm., width 19.5 mm., length of
movable part of antenna 7.5 mm., fronto-orbital width 9g mm. A male
is about 20 mm. long, but too damaged for measurement.

Distribution.—Monterey Bay, California (Harold Heath), one under
rocks between low tide mark and mean tide, and three from the sub-
umbrella space of a large violet-and-white jellyfish (type locality, Cat.
No. 22,868). Young specimens were taken by John C, Brown at Pacific
Grove, and by H. N. Lowe at San Pedro, California.

Relations.—This species may have been confounded with the young
of better known species. The young of C. magister has a nearly naked
carapace, the ninth tooth of the lateral margin is produced sideways, the
antenne are less than twice as wide as front, the merus of the maxilli-
peds is no broader than long, the carine of the upper and outer surfaces
of the hand are 6 instead of 7.

The carapace of young C. antennarius is also nearly naked (Dr. Holmes
may have had in his hand C. gibdosulus when he described the carapace
of young C. antennarius as thickly covered with hair); the teeth are all
much thickened, the orbital teeth prominent, the two underneath being
rounded, not sharp; the maxillipeds reach only to the base of the anten-
nal segment, the merus not broader than long; the dark color on the
fingers is more extensive than in C. jordani.

Cancer gibbosulus has a much more uneven carapace, strongly marked
and rounded orbital teeth, a postero-lateral tooth, ninth tooth more
prominent than eighth. Carapace hairy, as in C. jordani.

In C. anthonyi the antero-lateral teeth are low and broad, not alter-
nately large and small, the first 6 having blunt angles; the inner supra-
orbital tooth is well developed.

CANCER MAGISTER Dana.

Harriman Expedition.—Berg Bay, Glacier Bay.
Distribution.— Unalaska to Magdalena Bay, Lower California. Low
water to 50 fathoms.

CANCER GRACILIS Dana.

Distribution.—Kasaan Bay, Prince of Wales Island, Alaska, to Playa
Maria Bay, Lower California. Shallow water to 56 fathoms.

178 RATHBUN

CANCER OREGONENSIS (Dana).
Plate vii, fig. 2.

Trichocera oregonensis DANA, Proc. Acad. Nat. Sci. Phila., v1, 86, 1852;
Crust. U.S. Expl: Exped., 1,299, 1852, pl. XVIII, fig. 5, 1855.

Platycarcinus recurvidens BATE, Proc. Zool. Soc. London, 1864, 663.

Trichocarcinus oregonensis MIERS, Proc. Zool. Soc. London, 1879, 34.—
HOLMES, Occas. Papers Calif. Acad. Sci., VII, 54, 1900, and synonymy.

Trichocarcinus recurvidens WALKER, Trans. Liverpool Biol. Soc., XII, 271,
pl. Xv, figs. 1-1b, 1898.

Trichocarcinus Walkeri HOLMES, of. Cit., 53.

Harriman Expedition.—¥ox Island (Kincaid); Virgin Bay (Coe);
Orca; Yakutat; Sitka.

Distribution.—Aleutian Islands to Lower California (Holmes). Low
water to 238 fathoms.

I am unable to separate from typical C. oregonensis the form described
by Walker under the specific name vecurvidens, and later by Holmes as
Trichocarcinus walkeri, because the species displays more variation and
intergradation than was known to those authors. Material from 43
localities has been examined. In a lot of specimens collected by Dr.
Dall at Coal Harbor, Unga Island, Alaska, are not only typical ovegonensis
and typical wa/keri, but some with sculpturing and teeth intermediate
between the two; others more deeply areolated than in typical qwadke77,
but with teeth as narrow and well separated as in the other extreme of
oregonensis. In a lot of fairly typical oregonensis from the Strait of Fuca
(D. S. Jordan) is one in which many of the granules of the areolz
are prolonged into conical spines. Still another variation is shown
by a single male (plate vu, fig. 1) from station 3274 (Albatross),
northeast of Amak Island, Alaska, 19 fathoms, where the lateral teeth
are broader and more overlapping than in the specimen figured by
Walker (/oc. cit.); the granulate areole are small, round, and raspberry-
like, the interspaces much smoother than usual.

Size.—The largest specimen measured is a female from Puget Sound
(O. B. Johnson), 36.5 mm. long, 47.1 mm. wide. a

Cancer oregonensis might well be set apart generically on account of
the antero-lateral margins not being sharply marked off from the postero-
lateral, in which respect it differs from the other species of Cancer, in-
cluding C. gibbosulus and C. amphietus. The name Z7ichocarcinus is
not, however, available for the species ovegonensis, as the type of Zricho-
carcinus Miers, substituted for Zrichocera de Haan, preoccupied, is Cancer
gibbosulus (de Haan). This species I consider a true Cancer (see p.176) ;
therefore Z7ichocarcinus is a synonym of Cancer.

DECAPODS 179

Genus Telmessus White.
TELMESSUS CHEIRAGONUS (Tilesius).

Cancer cheiragonus TILESIUS, Mém. Acad. Impér. Sci. St. Pétersbourg, v, 347,
pl. vil, fig. I, 1815.

Telmessus cheiragonus BENEDICT, Proc. U. S. Nat. Mus., xv, 224, pls. XXv
and XXVI, figs. 2, 3, and 4, 1892.

Telemessus chetragonus HOLMES, Occas. Papers Calif. Acad. Sci., VII, 69, 1900,
and synonymy.

Harriman Expedition.—Popof Island, Shumagins; Fox Island, Prince
William Sound (Kincaid).

Distribution.—Northeastern Siberia; Kamchatka; Kurile Islands;
Bering Sea to California (Holmes). Low water to 20 fathoms.

Note on specimens from the type locality.—Since the publication of Dr.
Benedict’s ‘Corystoid Crabs of the Genera Telmessus and Erimacrus,’
Dr. Leonhard Stejneger has collected specimens of Zée/messus at Avacha
Bay, Kamchatka, the type locality of Cancer cheiragonus Tilesius. These
specimens are identical with 7. cheiragonus, as figured by Benedict;
otherwise the exaggerated lateral spines and the aculeate front of Tile-
sius’s figure would lead one to suppose that his was a different species.

Genus Erimacrus Benedict.
ERIMACRUS ISENBECKII (Brandt).

Platycorystes (Podacanthus) isenbeckii BRANDT, Bull. Phys.-Math. Acad,
Impér. Sci. St. Pétersbourg, vil, 180, 1848; Middendorff’s Sibirische
Reise, Bd. 11, Theil 1, p. 83, 1851.

Cheiragonus isenbeckii BRANDT, Middendorff’s Sibirische Reise, Bd. 11, Theil

Wpsta7,. Lost.

Platycorystes tsenbeckii RICHTERS, Abh. Senck. Natur. Gesell., Frankfurt,
X11, Heft 4, p. 402, plate, figs. 1 and 2, 1884.

Erimacrus tsenbeckii BENEDICT, Proc. U. S. Nat. Mus., XV, 229, pl. XXVI,
figs. 5,6, and 7, pl. XXVII, 1892.

Distribution.—Bering Sea, Aleutian Islands, and Alaska Peninsula.
Shallow water to 85 fathoms.

Family PORTUNIDZ.
Genus Portunus Fabricius.

PORTUNUS XANTUSII Stimpson.

Distribution. —From Puget Sound to Chile (A. Milne Edwards). Ex-
cept for a single specimen taken by the A/batross in Puget Sound,
this species is not known north of San Pedro, California.

180 RATHBUN

Genus Callinectes Stimpson.
CALLINECTES BELLICOSUS (Stimpson).

Distribution.— Point Loma, California, to Gulf of California.

Family P7Z UM NIDA
Genus Cycloxanthops Rathbun.

CYCLOXANTHOPS NOVEMDENTATUS (Lockington).
Plate vi, fig. 10.

Distribution.—San Pedro, California, to Guadalupe Island and San
Martin Island, Lower California.

CYCLOXANTHOPS RUGOSUS Holmes.

Distribution.—San Diego, California.

Genus Lophopanopeus Rathbun.
LOPHOPANOPEUS BELLUS (Stimpson).

as bella STIMPSON, Ann. Lyc. Nat. Hist. N. Y., VII, 204, pl. v, fig. 2,

hip eds bellus A. MILNE EDWARDS, Crust. Rég. Mex., 257, pl. XLVI,
fig. 4, 1879 (part).—HOLMES, Occas. Papers Calif. Acad. Sci., VII, 60,
pl. I, fig. 3, 1900, and synonymy.

Lophopanopeus bellus RATHBUN, Bull. Lab. Nat. Hist. State Univ. Iowa, Iv,
272, 1898.

Harriman Expedition.—Prince William Sound, at Fox Island (Kin-
caid) and Virgin Bay (Coe). This is the most northern locality known
for this species.

Distribution.— Prince William Sound; Kasaan Bay, Prince of Wales
Island, Alaska (U.S. Nat. Mus.); Vancouver Island, British Columbia
to Monterey, California (type locality of Xanthodes hemphillit Locking-
ton). Lower California (A. Milne Edwards).

The figure given by A. Milne Edwards resembles Z. de//us more than
it does any of the other species here described; the locality given is
Lower California.

There are on the Pacific coast six closely allied species of Lophopano-
peus ; some of these have been confused with LZ. de//us and L. leucomanus ;
therefore the published localities from which I have not examined speci-
mens are accepted with a reservation.

1 I have concluded to omit Daiva americana Stimpson from the list until it is known
that the species really occurs in California in contradistinction to Lower California.

DECAPODS 181

The characters most useful in determining the species are given below:

The carapace is roughened antero-laterally, but not conspicuously so.

The frontal lobes are slightly sinuous and oblique, the outer tooth
faintly indicated.

The anterior margin of the third antero-lateral tooth (or tooth N of
Dana) is in line with the truncate margin of the second tooth or the one
coalesced with the outer orbital tooth.

The carpus of the chelipeds is slightly roughened.

The hand is smooth, without a tooth on the upper margin; the proxi-
mal end of the inner margin of the upper surface is, however, well
marked.

The dark color of the fingers does not run back on the palm.

The carpal joints of all the ambulatory legs are slightly bilobed on the
anterior or upper margin.

The propodal joints have a convex anterior margin without lobes.

Dimensions.— Male, length 22.5 mm., width 33.8 mm., proportion of
length to width 1:1.5, fronto-orbital width 16 mm., proportion of fronto-
orbital width to width of carapace 1: 2.11.

LOPHOPANOPEUS FRONTALIS Rathbun.
Plate vu, fig. 8.

Distribution.—San Diego, California. The locality ‘Monterey’ cited
for this species is an error.

The carapace is almost smooth.

The frontal lobes are markedly oblique and sinuous, the middle lobes
very convex.

The third antero-lateral tooth is more produced than the second tooth.

The carpus of the chelipeds is smoother than in Z. dellus.

The hand is smooth, with a large tooth projecting inward on the inner
side of the upper margin and extending half the length of the upper
margin.

The dark color of the fingers runs back on the palm; in this respect
differing from all other allied species.

The carpal joints of all the ambulatory legs are slightly bilobed on
the anterior or upper margin,

The propodal joints have a convex anterior margin, but are narrower
than in Z. del/us.

Dimensions.—Male, length 17.2 mm., width 23.7 mm., proportion of
length to width 1 : 1.32, fronto-orbital width 12.5 mm., proportion of fronto-
orbital width to width of carapace 1: 1.9.

182 RATHBUN

LOPHOPANOPEUS HEATHII Rathbun.
Plate vu, fig. 9.

Lophopanopeus heathiti RATHBUN, Amer. Nat., XXXIV, 137, 1900.
Lophoxanthus leucomanus HOLMES, Occas. Papers Calif. Acad. Sci., VII, 61,
1900 (part).

Distribution.—Specimens are in the National Museum from Monterey
Bay, California, under stones, mean and low tide mark (H. Heath;
types, Cat. No. 22,870); Monterey Bay (J. O. Snyder) ; Catalina Island.
Others also from Monterey and Catalina Island, and belonging to the
Museum of the California Academy of Sciences, have been examined.

The carapace is smooth except on the outer half of the hepatic region,
where it is slightly roughened.

The margin of the frontal lobes is slightly oblique, the outer tooth
inconspicuous.

The anterior margin of the third antero-lateral tooth is in line with the
second tooth.

The carpus of the chelipeds is smooth, or nearly so.

The hand is smooth, one tooth on inner side of upper margin.

The dark color of the fingers does not run back on the palm.

The carpal joints of the ambulatory legs have the anterior or upper
margin slightly bilobed.

The propodi have slightly convex anterior margins.

Dimensions.—Male, length 12.2 mm., width 16.5 mm., proportion of
length to width 1 : 1.35, fronto-orbital width 11.6 mm., proportion of fronto-
orbital width to width of carapace 1:1.42.

While the antero-lateral teeth resemble those of Z. del/us, the smooth
surface and the tooth on the manus ally it closely to Z. frontalis.

LOPHOPANOPEUS LEUCOMANUS (Lockington).

Xanthodes leucomanus LOCKINGTON, Proc. Calif. Acad. Sci., VII, 32, 1877 (not
X. leucomanus, op. Cit., p. 100).

Lophoxanthus bellus A. MILNE EDWARDS, Crust. Rég. Mex., 257, 1879 (part).

Lophopanopeus leucomanus RATHBUN, Bull. Lab. Nat. Hist. State Univ. Iowa,
IV, 272, 1898.

Lophoxanthus leucomanus HOLMES, Occas. Papers Calif. Acad. Sci., VII, 61,
pl. I, fig. 4, 1900 (part).

Distribution.—¥From Monterey to San Diego, California: San Pedro,
Catalina Island, and San Diego (National Museum); Monterey, Santa
Rosa Island, and San Diego (Lockington). Types not extant (Holmes).

The anterior half of the carapace is deeply rugose, the roughness on

the hepatic region not divided from that on the two adjacent teeth.

DECAPODS 183

The frontal lobes are strongly oblique, except the outer tooth, which
is not strongly marked.

The third antero-lateral tooth is more produced than the second
tooth.

The carpus of the chelipeds is covered with numerous small irregular
pits separated by reticulating lines.

The upper portion of the hand is pitted and reticulated like the carpus,
and has two or more irregular inward-pointing teeth on upper margin,
the proximal the larger.

The color of the fingers does not run back on the palm.

The carpal joints of the ambulatory legs have on the anterior or upper
margin a very thin crest, strongly bilobed.

The propodal joints are wide, and have a prominent lobe at the prox-
imal end.

Dimensions.—Male, length 8.7 mm., width 11.6 mm., proportion of
length to width 1: 1.33; fronto-orbital width 7.1 mm., proportion of
fronto-orbital width to width of carapace 1: 1.63.

LOPHOPANOPEUS LOCKINGTONI Rathbun.
Plate vu, fig. 7.

Xanthodes leucomanus LOCKINGTON, Proc. Calif. Acad. Sci., vII, 100, 1877
(not X. leucomanus, op. cit., p. 32).

Lophoxanthus bellus A. MILNE EDWARDS, Crust. Rég. Mex., 257, 1879 (part).

Lophoxanthus leucomanus HOLMES, Occas. Papers Calif. Acad. Sci., vil, 61,
1900 (part).

Lophopanopeus lockingtoni RATHBUN, Amer. Nat., XXXIV, 137, 1900.

Distribution.—San Diego Bay, California, to Gulf of California. Speci-
mens examined from San Diego Bay, 414 fathoms (A/batross, type local-
ity); San Diego (H. Hemphill); San Diego (Cal. Acad. Sci.); Gulf of
California at La Paz, 3 fathoms, Mulege Bay, and Port Escondido (speci-
mens described by Lockington, of. cit., p. 100, not p. 32).

The anterior half of the carapace is irregularly roughened ; the obliquely
oval granulated patch on the hepatic region is separated from the gran-
ules on the adjacent teeth.

The frontal lobes are slightly oblique, but the outer tooth is well
marked.

The third? antero-lateral tooth is more produced than the second
tooth.

The carpus of the chelipeds is crossed by a few thickened ridges
incompletely reticulating, anterior margin having a thick granulated ridge
distinctly separated by a deep sulcus from the rest of the carpus; a large
granulated tubercle near articulation with hand.

184 RATHBUN

The upper margin of the hand has two longitudinal ridges, the outer
straight, the inner curved and ending proximally in a tooth; outer sur-
face of hand granulate, the granules tending to form in longitudinal and
transverse lines, and becoming fainter toward the fingers.

The dark color of the fingers does not run back on the palm.

The carpal joints of the ambulatory legs are distinctly but not mark-
edly bilobed on the anterior or upper margin.

The propodal joints have a slightly convex anterior margin.

Dimensions. —Male, length 9.6 mm., width 13.3 mm., proportion of
length to width 1: 1.39; fronto-orbital width 7.9 mm., proportion of
fronto-orbital width to width of carapace 1: 1.68.

LOPHOPANOPEUS DIEGENSIS Rathbun.
Plate Ix, . fie. 3:

Lophopanopeus diegensis RATHBUN, Amer. Nat., XXXIV, 137, 1900.

Distribution.—Off Point Conception, California, 31 fathoms (Albatross
station 2908); San Diego, ro fathoms (H. Hemphill), one male, type
(Cat. No. 4281).

The carapace has a few granulate lines on the antero-lateral regions.

The frontal lobes are slightly oblique and sinuous, the outer tooth
inconspicuous. The front is deflexed so that its true edge does not show
in the view figured.

The second antero-lateral tooth, though small, is dentiform and not
merged with the first or orbital. The third, fourth, and fifth teeth are
prominent, carinate, and subacute.

The carpus of the chelipeds has about 35 unequal tubercles, irregularly
placed, some of them forming lines.

The hand is bicarinate above, the inner carina having a small promi-
nence at the proximal end; a few tubercles on upper part of inner sur-
face; upper and proximal portion of outer surface rough with fine granu-
lated ruge.

The dark color of the fingers does not run back on the palm.

The carpal joints of all the ambulatory legs have on the anterior
margin two prominent naked truncate tubercles, by which this species
may at once be recognized.

The propodal joints of the last two pairs have a smaller similar lobe at
the proximal end of the anterior or upper margin.

Dimensions.—Male, length 7.9 mm., width 11.4 mm., proportion of
length to width 1: 1.44; fronto-orbital width 6.6 mm., proportion of
fronto-orbital width to width of carapace 1: 1.37.

DECAPODS 185

Genus Xanthias Rathbun.

XANTHIAS TAYLORI (Stimpson).

Distribution—Monterey, California, to Magdalena Bay, Lower Cali-
fornia. Beach to 55 fathoms.

Size.— Unusually large individuals have been collected at Pacific
Grove, California, by John C. Brown. An egg-laden female measures
24.6 mm. long on the median line of the carapace, and 42 mm. wide.

XANTHIAS LATIMANUS (Lockington).
Distribution.—San Diego, California (Lockington).
Species not seen since Lockington’s record.

Genus Pilumnus Leach.
PILUMNUS SPINOHIRSUTUS Lockington.
Plate vit, fig.-2;
Distribution.—San Diego, California, and Gulf of California; 10-22
fathoms. Bay of St. Elena, Ecuador (Nobili).
Genus Heteractza Lockington.

HETERACTAA LUNATA (Milne Edwards and Lucas).
Pilumnus lunatus MILNE EDWARDS and Lucas, D’Orbigny’s Voy. l’Amér.

Mérid., Crust., 20, 1843, pl. IX, fig. 2, 1847.
Heteractaa lunata KINGSLEY, Proc. Acad, Nat. Sci. Phila., 1879, 396 (1880).

Distribution.—San Diego, California (Faxon) to Chile.
Family P7-MNOTHERIDA.

Genus Pinnotheres Latreille.
PINNOTHERES NUDUS Holmes.

Distribution. — Monterey and Santa Cruz, California.

PINNOTHERES PUGETTENSIS Holmes.

Distribution, — Puget Sound, in branchial cavity of a Halocynthia.

Genus Raphonotus Rathbun
(= FABIA Dana).
Raphonotus RATHBUN, Proc. Biol. Soc. Washington, XI, 166, 1897.

186 RATHBUN

RAPHONOTUS SUBQUADRATUS (Dana).

Fabia subguadrata HOLMES, Occas. Papers Calif. Acad. Sci., vil, 87, 1900
(part).

Distribution.—Alaska to Monterey, California. Specimens are in the
National Museum from: South entrance to Akutan Pass, Alaska, 45
fathoms (AJbatross station 2843); Monterey, in folds of Lucapina crenulata
(Dr. C. A. Canfield); Pacific Grove, from mantle cavity of Atlus edulis
(John C. Brown).

I believe that the specimens recorded from farther south than Mon-
terey Bay are a distinct species from 2. subguadratus. Those seen and
described by Dr. Holmes from San Pedro as Fabia subguadrata are the
species which follows, 2. owe.

RAPHONOTUS LOWEI Rathbun.

Fabia subguadrata HOLMES, Occas. Papers Calif. Acad. Sci., vil, 87, 1900
Py ase lowei RATHBUN, Amer. Nat., XXXIV, 590, 1900.

General appearance like that of 2. sudguadratus, differs as follows:

In &. Jowei the turned-down front has no trace of a transverse groove
and is naked; in &. suéguadratus the turned-down front has a shallow
transverse sulcus which is covered with pubescence.

In &. /owei the last joint of the maxilliped does not
reach the end of the penultimate joint, as it does in 2.
subquadratus.

In &. lowei the palm of the chela does not widen dis-
tally ; its margins are subparallel or a little convex; its
lower surface has only one line of hair, which is con-
/ tinued to the end of the pollex; in A. subguadratus the
Fic. 93. Raphonotus palm widens a little distally, as figured by Dana, and

lowet. 9%. San Pedro

Bava Ree ae is furnished with two rows of hair below, the inner
b. Chela (X 33). one of which is continued to the end of the pollex.

In &. Jowei the fingers are less arched and less gaping than in #. sud-
guadratus,

The carapace is also a little wider in our species.

Dimensions.—Female, length of carapace 1o mm., width 12.5 mm.,
length of merus of third ambulatory leg 5 mm.

Distribution.—San Pedro to San Diego, California: San Pedro Bay,
in siphon of Pholas pacifica, collected by H. N. Lowe, for whom the
species is named (type locality); San Pedro, in shell of Zapes (Holmes,
as Fabia subguadrata); San Diego (H. Hemphill), one of the specimens

a

DECAPODS 187

examined is from Luchydesma crassatelloides. Recorded by Lockington
from the same locality, commensal, and collector, under the name of
Fabia subquadrata,

Genus Pinnixa White.
PINNIXA OCCIDENTALIS Rathbun.
Plate vu, fig. 4; plate 1x, figs. 6, 6a.

Distribution.—Unalaska to Humboldt County, California (Holmes).
Shallow water to 238 fathoms.

The specimen recorded by me in 1893, from San Diego, is P. Uittoralis
Holmes.

Harriman Expedition.—Juneau, 20 fathoms; Hot Springs; Sitka;
Wrangell, in burrows of Achiurus (T. Kincaid); Cape Fox.

This species shows remarkable variation in the length and breadth of
the ambulatory legs, especially of the third pair, as is shown in the
accompanying table:

Sex. Station. Length of | | Widthof | "SPE ambu | of third ambuc
Seances Usaha cara paeS latory leg. latory leg.
fot 2862 7.5 mm 15.8 mm. 8.7mm 2.8mm
g Wrangell 12 Bi 2.1
2002 7:3 14.5 7 2.6
2 Wrangell Uf 14.2 5.5 Sor

I was at first led to believe that the Wrangell specimens represented a
distinct species, but a reéxamination of the entire collection of P occi-
dentalis shows intermediate forms. It may be added that station 2862,
which yielded the longest and slenderest legs, is also the greatest depth
(238 fathoms) from which the species is known.

Small specimens often exhibit an almost complete absence of the car-
diac ridge.

PINNIXA CALIFORNIENSIS Rathbun.
Plate vit, fig. 3.

Distribution.—¥From off Point Ano Nuevo, California, to Magdalena
Bay, Lower California; 13 to 51 fathoms.

PINNIXA TUBICOLA Holmes.

Distribution.—From Puget Sound to San Diego, California. Taken
by the Albatross off Point Conception, California, 31 fathoms, station
2908; in calcareous tube of worm, Trinidad, California (Holmes).

188 RATHBUN

PINNIXA LITTORALIS Holmes.
Distribution. —Bodega Bay to San Diego, California.

PINNIXA LONGIPES (Lockington).

Distribution.—Tomales Bay to San Pedro, California. In tube of
annelid (C/ymenella).
PINNIXA FABA (Dana).

Distribution.—From Sitka, Alaska, to San Pedro, California. In shells
of large bivalve mollusks, and in the cloaca of a large holothurian,
Liosoma arenata Stimpson.

Genus Scleroplax Rathbun.
SCLEROPLAX GRANULATA Rathbun.
Plate vit, fig. 5.

Distribution.—From Bodega Bay, California, to Ensenada, Lower

California.
Genus Parapinnixa Holmes.

PARAPINNIXA AFFINIS Holmes,

Distribution. — Deadman Island, San Pedro, California.

Genus Cryptophrys Rathbun.
CRYPTOPHRYS CONCHARUM Rathbun.
Plate vu, fig. 6.
aus Distribution.—From Puget Sound to San Diego,
California. From mantle of Mya arenaria Lin-

Fic. 94. Cryptophryscon- neeus, Cardita borealis Conrad, and Mytilus edulis
charum. Outer maxillipeds

(Xx 8). Pacific Grove, Calif. Linnezus.

Genus Opisthopus Rathbun.
OPISTHOPUS TRANSVERSUS Rathbun.

Plate 1x; fig. 2.
Distribution.— From Monterey to San Diego, California. |
In Lucapina crenulata Sowerby, siphon of Pholas, and (\

mantle cavity of Afptilus edulis. The suture dividing the /”

ischium from the merus of the maxillipeds is obsolescent ; rope ae aa
while in most specimens it is deep on the ventral surface, {Sin *3¢ pe
in none is it evident on the upper surface. Packers

DECAPODS 189

Family GRAPSIDA.
Genus Pachygrapsus Randall.
PACHYGRAPSUS CRASSIPES Randall.

Distribution.—¥rom Oregon to Gulf of California; Japan; Korea.
The locality Sandwich Islands, given by Randall, is too doubtful to be
relied upon.

PACHYGRAPSUS TRANSVERSUS Gibbes.

Distribution —¥From California to Peru; Galapagos Islands; Oriental
region; West Africa; Bermudas; Bahamas and Florida Keys to Rio
de Janeiro.

Genus Hemigrapsus Dana.

HEMIGRAPSUS NUDUS (Dana).

Brachynotus nudus HOLMES, Occas. Papers Calif. Acad. Sci., vit, 81, 1900,
and synonymy.

femigrapsus nudus RATHBUN, Amer. Nat., XXXIV, 587, 1900.

Harriman Expedition.—Sitka, 10 fathoms; Metlakatla, Alaska;
Grenville Channel, British Columbia.

Distribution.—From Sitka to Gulf of California.

HEMIGRAPSUS OREGONENSIS (Dana).

Brachynotus oregonensis HOLMES, of. cit., 82, and synonymy.
Hemigrapsus oregonensis RATHBUN, @oc. cit.

Harriman Expedition.—Fox Island, Prince William Sound (T. Kin-
caid); Yakutat; Sitka, 10 fathoms; Metlakatla; Grenville Channel,
British Columbia.

Distribution. — From Prince William Sound, Alaska, to Gulf of Cali-
fornia. Fox Island is farther north than the species has been collected
hitherto.

Genus Grapsodius Holmes.

GRAPSODIUS EXIMIUS Holmes.

Distribution, —San Diego, California.

Genus Planes Leach.
PLANES MINUTUS (Linnzus).

Distribution.—Temperate and tropical seas. Specimens are in the
National Museum from lat. 41° N., long. 141° W., at sea west of Hum-
boldt Bay, California (W. H. Dall, 1871); California (D. S. Jordan,

190 RATHBUN

1880); Point Loma, California; and off San Diego (A/Jatross sta-
tion 2928).
Family OCYPODIDA.
Genus Ocypode Fabricius.
OCYPODE GAUDICHAUDII Milne Edwards and Lucas.

Distribution.—The normal range of this species is from Nicaragua
(Gulf of Fonseca) to Chile and Galapagos Islands; Honolulu (Cano).
Three young specimens from San Pablo, California, are in the Museum
of Comparative Zoology, determined by Dr. Faxon.

Genus Uca Leach.
UCA CRENULATA (Lockington), 1877.

(= Gelasimus gracilis RATHBUN, 1893 = G. macrodactylus BOUVIER,
1895,not Milne Edwards and Lucas = U. stenodactyla ORTMANN, 1897,
part (specimen from San Diego), not Milne Edwards and Lucas.)

Distribution. —From San Diego, California, to Gulf of California.
Genus Speocarcinus Stimpson.
SPEOCARCINUS CALIFORNIENSIS (Lockington).
Plate 1x, fig. x.

Distribution.—San Pedro and Alamitos Bay, California.

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PLATE I.

Fic. 1. Pandalopsis aleutica.
9, X 3%. Albatross station 3480.
2. Pandalopsis dispar.
9, X 3. Albatross station 3675.
3. Landalus goniurus.
Q, nat. size. Albatross station 3293.

(192)

Fic.

Ve

2.

PLATE II,

Pugettia dalli.
9, X13. Southern California.

. Pugettia dalli.

g, X14. Southern California.
Pandalus montagui tridens.

9, X2. Albatross station 2842.
Pandalus jordani.

9, X#. Albatross station 2949.
Pandalus stenolepis.

9, X2. Albatross station 3464.
Pandalus hypsinotus.

&, X 2. Herendeen Bay, Alaska.
Pandalus gurneyt.

9, Xx. Albatross station 3130.

(194)

‘16

‘37

re

na

Fic.

PLATE III.

Sclerocrangon sharpt.

dé, dorsal view, nat. size. Albatross station 2842.
Sclerocrangon sharpi.

é, profile. nat. size. Adbatross station 2842.
Sclerocrangon alata.

é, dorsal view, nat. size. Albatross station 2865.
Spirontocaris gaimardit belcheri.

gf, nat. size. Adlbatross station 3519.
Spirontocaris gaimardii belcheri.

Q, nat. size. Albatross station 3519.
Spirontocaris arcuata.

@, nat. size. AJldatross station 2864.
Spirontocaris spina.

@, nat. size. Albatross station 2842.
Spirontocaris murdochi.

Q, nat. size. Albatross station 3650.

(196)

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PLATE IV.

Fic. 1. Pagurus dalli.
&, X14. Albatross station 3233.
2. Pagurus rathbuni.
o, X43. Albatross station 3519 (chelz foreshortened).
3. Pagurus capillatus.
&, X#. St. Michael, Alaska.
4. Pagurus brandti.
&, X# Albatross station 2847.
5. Lagurus townsendi.
S$, X23. Albatross station 3502 (chelz foreshortened).
6. Pagurus undosus.
&, Xx. Albatross station 3638.
4. Pagurus tanneri.
3, X#. Albatross station 3016 (chelz foreshortened).
8. Pagurus confragosus.
3g; X23. Cat. No. 19,228 (chelz foreshortened).

(198)

Gi Areiee VOLES 2x

DECAPOD CRUSTACEANS

HELIOTYPE CO., BOSTON.

PLATE

FIG.

10.

PLATE:V.

Pagurus setosus.
3d, X%- Albatross station 3449.
Pagurus munitus.
@, X2. Albatross station 2843.
Pagurus cornutus.
9, X%. Albatross station 3608.
Pagurus kennerlyt,
g, in sponge (Suderites), X 2. Albatross station 3265.
Pagurus beringanus.
&, X%- Albatross station 3234.
Parapagurus mertensis,
&, Xz. Albatross station 2948 (larger chela foreshortened).
Pagurus samuelis.
3, X 7%. Pacific Grove.
Pagurus granosimanus.
&, X34; Pacific Grove.
Pagurus hemphillit.
dé, X7%- Monterey (chelz foreshortened),
Pagurus gilli.
é, X #2. Dutch Harbor.

(200)

WW

Fic.

PLATE, Vi.

Placetron wosnessenskii.

Q. Type of Lepeopus forcipatus, X ya
Cancer anthonyi.

dé, Xz. Playa Maria Bay.
Cancer amphietus.

&, nat. size. Albatross station 3656.
Cancer jordani.

Q, nat. size. Monterey Bay.
Oregonia bifurca.

©, type, X x5:
Lachycheles rudis.

df, nat. size. Pacific Grove.

(202)

uw

a

Fis,

10.

PLATE VIl-.

Cancer oregonensis.
&, nat. size. Albatross station 3274.

Pilumnus spinohirsutus,

@, nat. size. Albatross station 3026.
Pinnixa californiensis.

Q, nat. size. Albatross station 3133.
Pinnixa occidentalis.

&, nat. size. Albatross station 2862.
Scleroplax granulata.

9, x 1%. Ensenada.
Cryptophrys concharum.

é, X 1%. Neah Bay, Washington,
Lophopanopeus lockingtoni.

@, nat. size. AJldatross station 3591.
Lophopanopeus frontalis.

&, nat. size. Southern California,
Lophopanopeus heathii.

g, nat. size. Monterey Bay.
Cycloxanthops novemdentatus.

fg, nat. size. Guadalupe [sland,

(204)

PLATE VIII.

Callianassa goniophthalma.
é, nat. size. Albatross station 3198.

(206)

PLATE VIII

H.A.E. VOL.X

DECAPODS

Fic.

PLATE IX.

Speocarcinus californiensis.

&, X14. Alamitos Bay.
Opisthopus transversus.

9, X 1}. Pacific Grove.
Lophopanopeus diegensts.

d, type, X 14.
Clythrocerus planus.

&, X6. Southern California.
Clythrocerus, sp.

Carapace of $, X 6, Catalina Island.
Pinnixa occidentalis.

&, X14. Wrangell.

6a. Pinnixa occidentalis.

Chela of ¢, X 14. Wrangell.

(208)

LAE VOL :X

DECAPODS

Fic.

PLATE X.

1. Lyrileptus spinosus.
8, X24. Albatross station 2934.
2. LPodochela hemphiltit.
é, X&. Albatross station 2831.
3. Dasygyius tuberculatus.
g, nat. size. <AJléatross station 3030.
3a. Dasygyius tuberculatus.
d, ventral view, nat. size. <A/dbatross station 3030.
4. Anasimus rostratus.
Q, X 34. Albatross station 2983.

(210)

H.A.E. VOL.X

DECAPODS

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