THE LIBRARY
OF
THE UNIVERSITY
OF CALIFORNIA
LOS ANGELES
THE FAUNA OF BRITISH INDIA,
CEYLON AND BURMA.
PUBLISHED UNDER THE AUTHORITY OF THE SECRETARY OP
STATE FOR INDIA IN COUNCIL.
EDITED BY A. E. SHIPLEY, Sc.D.Cantab., HON. D.Sc. Princeton,
HON. LL.D. Michigan, F.B.8.
ASSISTED BY GUY A. K. MARSHALL, HON. D.Sc. (Oxon.), F.Z.8.
DIPTERA BEACHYCEKA.
VOL. I.
BY
E. BRUNETTI.
LONDON:
TAYLOR AND FRANCIS, RED LION COURT, FLEET STKEET,
CALCUTTA :
THACKER, SPINK, & CO.
BOMBAY:
THACKER & CO., LIMITED.
May, 1920.
I'RINTKD BY TAYLOR A N' D FUANCIS,
RED LION COURT. Fl.KKT STHKET. LO.VPON.
PREFACE.
OWING to the war, the difficulties of communication, and the
loss of ships coining from the East, there has been an
unusual delay in the issue of the present volume, and it has
unfortunately been found impossible to incorporate in it
certain corrections which the author wished to make.
Further, owing to the difficulties under which publishers and
printers now labour, it was felt by the Editor necessary to
publish Avithout further delay.
Another difficulty which has arisen in the preparation of
the volume is due to the fact that the MS. ran to a far larger
volume than the Indian Government contemplates, and it
was therefore reluctantly decided to publish the following
families : Stratiomyidse, Leptidjr, Nemestrinidfe, Cyrtidee,
Bombyliidce, Therevida?, Scenopinidse, Mydaidse, Empidoe,
Lonchopteridac, and Platype/idse ; leaving the remainder,
i. «., the families Pipunculida?, Syrphidse, Conopidse, and
Oestridse for a second volume. This proceeding is, I gather,
against the wishes of the author, but no other solution of
the difficulty appeared possible.
A.E. SHIPLEY.
1st May, 1920.
SYSTEMATIC INDEX.
Page
Order DIPTEBA 1
Suborder BR ACHY CERA .. 1
Fain. 1. STRATioMYiixas .... 17
Subfam. 1. Pachyyastrinte .... 19
1 Pachvo-nster Mq 20
P
2 Eudmeta Wied
age
41
42
43
48
44
44
45
46
47
47
48
4!)
lit
50
52
52
58
58
55
56
57
57
58
59
(iO
60
61
68
64
65
65
67
67
68
1. maiginHta, Fabr
2. brunnea, Meij.
3 flavida Brun ....
1. longispinus, Brun
4. Epnippiuin, Latr
1. bilineatum, Fabr
5. Negritomyia, Big
1. macnlipennis, Macq. ..
6 Ruba Walk
1 annulipes Brim
21
21
21
22
23
24
24
24
24
25
26
26
28
28
29
29
30
31
32
32
33
33
34
36
37
38
30
40
2. Zabrachia, Coq
1. albipes, Brun
2. annul! femur, Brun
3. Monaoantbomyia, Brun. . .
1. annnndalei, Brun
1 infl.-ta Walk
7. Clitellavia, Mg
1. hen-.inopla. Wied
2. bistriata, Brim
1 felderi Brauer
8. Lasiopa, Brittle
1 . villosa, Fabr.,\&\\ hima-
layensis, Brun
9 Oxyceva Mq
5 Wallacea Dol
1. argentea, Dol
6. Acanthina, Wied
1. sicjnnta, Brim
2. argeiitibirla, Brun
7. Craspedometopon, Kert. . .
1. frontal a, Kert
8 Tin da If 'afk
2. albomicans, Brun
Subfam. 3. Stratifmtyintc ....
1. Cvphoniyia, Wied
1. indica, Brun
1. javana, Macq
9. Evaza, Walk..
1 flavipes Biq
2. Stratiumyia, Geoff.
1. barcai Walk
2. mioropilosa, Brun
3. fulvescens, Brun
3 Odontomyia, Mq
2. bipars, Walk
3 indica Kert
10. Ptilocera, Wied
1. fastuosa, Gerst
11. Strat.iospbecomyia, Brun. .
1 variegata, Brun
1. cyanea, Brun. . . .
2. rufoabdominalis, Brun. .
3. pulcherrima, Brun
4. minuta, Fabr
Subfam. 2. Clitellariinee
I. Hermelia, La.tr
1. Iseta. Meii. .
5. rubrithorax, Macq
6. kashmirensis, Brun. . .
7. dorsoang-ulata, Brun. . .
VI
SYSTEMATIC IKDEX.
Odontornyia (con.). Page
8. soleimis, Walk 69
9. lutatius, Walk 70
10. transversa, Brun 71
11. viridana, Wied. 72 j
2. nigrieornis, Brun. .
3. unicinctus, Bnm. ,
Subfam. 2. Arthroceratinte
Page
. 105
. 106
. 107
Suh
1.
2.
3.
4.
5.
6.
Sul
3.
Su
1.
Fa
Su
1
fam. 4. Sargince ..........
Ptecticus. Lw ....
72
73
75
75
76
77
77
78
79
80
81
81
82
83
84
84
85
85
86
86
87
88
88
89
90
91
92
92
93
93
94
95
96
96
98
98
99
100
104
104
105
1. Desmomyia, Brun
107
107
108
109
110
110
111
112
112
114
115
115
116
117
118
119
120
120
121
121
122
122
123
124
125
126
126
127
128
131
131
132
133
133
134
135
135
136
137
137
138
138
138
139
140
140
141
1. thereviformis, Brun. . .
2. Pararthropeas, Brun
1. thereviformis, Brun. . .
Subfam. 3. Coenomyina
1. Coenomyiodes, Brun
1. edwardsi, Brun
1. longipennis, Wied
3. aurobrunneus. Brun. . .
4 wulpii Brun
5. apicalis, Lw
7. australis, Sch
Subfam 4 Le tin*
8. cingulatus, Lw., var.
ceylouicus, Bnm
Sargus, Fabr
1. gemmifer, Walk
2. metallinus, Fabr
3. mactans, Walk
Chloromyia, Dune
1. sapphirina, Walk
Brachycara, Thorns
1. Atrichops, Verr
1 . calopa, Brun
2. lanopyga, Brun
3. limbata, O.-S.
4. int frmedia, Brun
5. metatarsalis, Brun
/ 6. cincta, Brun
7. caerulescens, Brun
2. Atherix, Mg
1 labiata Biq
1. ventralis, Thorns
1. vitripennis, Dol
3. Suragina, W«lk
1. elegans, Karsch
| 4. Macellopalpus, Big
1. f ul v id us, Brun
2. fascipennis, Brun
5. Leptis, Fabr
1. flaviventris, Wied
2. calopa, Bnm
)fam. 5. Berina
Beris, Lair
1. geniculata, Curt
"1. annul ipes, Brun
Chlorisops, Rond
1. tibialis, My
Allognosta, O.-S.
1. vagans, Lw
2. assamensis, Brun
)fam. 6. Xylomyince
Xylomyia, Rond
l'. flavipes, Dol
2. calopodata, Biy
3. nigricornis, Brun
m. 2. LEPTID^E . .
1. apicipennis, Bnm
2. discoidali^, Brun
3. imicolor, Brun
4. triangulata, Brun
6. Chrysopilus, Macq
1. marmoratus, Brun
2. albopictus, Brun
3. ornatipennis, Brun
4. luctuosus, Brun
5. flavopnnctatus, Brun.. .
6. opalescens, Brun
7. humeral!*, Bnm
8. segmentatus, Brun
9. cochinensis, Bnm
10. birmaneiisis, Brun
12. similis, Brun
13. magnipennis, Brun
14. flavopilosus, Brun
15. yerburyi, Brun
16. albobasalis, Brun
17. insularis, Sch.
bfam. 1. Xylophityince
Rliachicenis, Walk
1. bicolor, Bnm
18. ferrugiDosus, Wied.
SYSTEMATIC INDEX.
Chrysopilus (con.).
19 unicolor Brun
Page
141 3 frmtnliis Vnhv
Page
181
20. atigma, Brun
21. latus, Brun
22. sordidus, Brun
23. alternates, Brun. . . .
Fam. 3. NEMESTRINID.S: . . .
1. Hirmoneura, My
1. annandalei, Licht. . . .
2. basalia, Licht
3. opaca, Licht
. 142
. 143
. 143
. 144
. 144
. 147
. 149
. 149
. 150
4. chrysolampis, Jaeii. .
5. flaviventiis, Dol. . . .
.. 182
183
6. ceuomaus, Rvnd. . . .
7. aurautiaoa Guer.
.. 185
186
8. sphinx, Fabr
9. suft'usipennis, Brun.
10. semifuscata, Brun. .
2. Exoprosopa, Mncq. . . .
1. flavipennis, Brun. .
2. flammea, £run. . . .
3. lar, Fabr.
.. 187
.. 188
.. 188
.. 189
... 192
.. 193
. . . 194
4. cingulata, Licht
5. brunnea, Licht
. 151
. 151
4. pennipes, Wied. . . .
5. latipennis, Brun. . . .
6. lateral is, Brun.
7. annandalei, Brun. .
8. maculiveutris, Brun.
9. stylata, Brun
10. insulata, Walk. . . .
11. puerula, Brun
12. bengalensis, Macq. .
13. brahma, Sch
14. niveiventris, lirun. .
15. vitripennis, Brun. .
16. vitrea, Biy
17. basifascia, Walk. .
18. alex.m, Walk
19. semilucida, Walk. .
20. auviplena, Walk
21 albida Walk
... 197
. . . 198
.. 198
.. 199
.. 200
.. 201
.. 203
.. 205
.. 206
.. 206
.. 207
.. 209
.. 210
.. 211
.. 211
212
.. 212
212
6. ochracea, Licht
7. austeni, Licht
8. coffeata, Licht
2. Trichopsidea, Wesf.w. . . .
1. duhrni, Wand
3. Atriadops, Wand
1. javana, Wied.
. 151
152
. 152
. 153
. 153
. 154
. 355
155
4 Ceylunia, Licht
1. magnifica, Licht
Fain. 4. CYBTIDJE
. 156
. 157
Subfam. 1. Philopotina
1. Philopota, Wied
1 viridaenea. B'run
. 158
. 158
. 159
Subfam. 2. Panopints
1. Pialea, Erichs
1. auripila, Brun
Subfam. 3. Cyrtina
1. Lasia, Wied
1. aurata, Brun
2. Pterodontia, Gray
. 160
. 160
. 161
. 163
. 163
. 164
. 165
. 166
. 167
. 168
169
. 170
171
. 171
. 171
173
176
. 178
180
. 180
3. Spog< stylum, Macq. . . .
1. austeni, Brun
4 Argvramoeba Sch
.. 213
.. 213
214
1 bipunctata, Fabr.. - .
. 217
2 di-tigma, Wied
218
3. obscurifrons, Brun. .
4. limitaris, Brun. . . .
5. intermedia, Brun. .
.. 220
.. 220
.. 221
222
1. kashmirensis, Licht. .
2. dorsolineata, Brun. . . .
3 Oncodes Lair
7 gestroi, Brun
222
8. niveicauda, Brun. .
9. duvaucelii, Macq. . .
10. argentiapicalis, Brun
11 instituta Walk. . . .
.. 224
.. 225
.. 227
227
2. octomaculatus, Brun. .
12. claripennis, Brun. .
13. niveisquamis, Brun.
14. nigrofemorata, Brun.
. . 228
.. 229
.. 230
230
4. angustimarginatus,
5. rufomarginatus, Brun.
Fam. o. BOMBYLIID^;
Subfam. 1. Anthracina
1. Hyperalonia. Rond
1. dives, Walk
2. tristis. Wuln .
5. Lepidanthrax, O.-S
1. compacta, Brun
2. transversa, Brun.. . .
6. Anthrax, Scop
1. fletcheii, Brun
2. guttatipennis, Brun.
3. himalayanus, Brun.
.. 232
.. 233
.. 233
.. 234
.. 236
,. 237
.. 238
SYSTEMATIC INDEX.
Anthrax (con.).
4. afra, Fabr
Page
. 240 8. Phtbiria, Mg
241 1. oT»<>.ilis. Walk.. .
Pago
. . 284
285
6. manifest, Walk. . . .
7. clara, Walk
8. approximata, Brim. .
9 aperta Walk
. 242
. 243
. 244
. 245
. 246
. 247
248
Subfam. 4. Toxophorina ....
1. Toxophora, Mg
1. javana, Wied.
Subfam. 5. Si/stropince
. . 280
. . 287
. . 287
. 289
10. fuscotimbata, Brun. .
11. hottentotta, var. clari
pennis, Brun
1 . Systropus, Wied
. 289
. . 291
. 294
13 dia Wifd . • • .
250
1. ophioneus, Westw. . .
2. edwardsi, Brun
14. baluchianus, Brun
15. albofulva, Walk
16. aureohirta, Brun. . . .
. 250
. 251
. 252
. 253
. 254
3. eunienoides, Westw.
4. flavipleura, Brun. . .
; Fam. 6. THEREVIDJE
. 294
. . 295
. 295
. 297
. 298
18 troflodvta, Fabr
19! iucida/JFaa
. 254
. 255
255
20. limpida, Walk
21 referens Walk
1. kempi, Brun
Subfatn ^ Ltnnatiinee
255
. 256
. 257
259
. 260
260
261
263
2H4
265
267
267
268
270
270
270
271
272
273
273
274
275
276
277
277
278
278
278
279
279
280
281
282
283
2. nigella, Wied. . .
3. flavolineuta, Brun. . .
4. bilineata. Brun
5. cylindrica, Walk. . .
6. indica, Walk
299
. L>99
. 300
. 301
. 301
301
1. Aplioebantus, Lw
1. ceylonicus, Brun. . . .
Subfam. 3. Bombylima
1. Empidideicus, Beck
1. indicia, Brun
8. invaria, Walk
. 302
. 302
. 303
. 304
2 Psilocephala, Zett
1. albina, Wied
2. indica, Big
2. Bombylius, Linn
3. ceylonica, Krob
. 304
2. wulpii, Linn
3. major, Linn
4. erectus, Brun
5. maculatus, Fabr
6. comastes, Brun
4 sequa Walk
305
5 affinis Krob.
306
3 Phycus, Walk
306
1. brunneus, Wied
2. argentiventris, Brun.
3. atripes, Brun
. 307
. 309
. 309
8 vicinus, Brun
Fam. 7. SCENOPINIDJE ....
. 310
. 311
312
9. terminalis, Brun
10. fulvipes, Big
11. propinquu.s, Brun. . . .
'•]. Dischistus Lw.
1 fenestralis, L
Fam. 8. MYDAIDJE
. 314
1. resplendens, Brun. . . .
2. ardens, Walk
1. Mvdas, Fabr
. 316
. 317
. 318
1. socius, Walk
1. ruticornis, Wied. . . . ..
2. carmichaeli, Brun. . . .
3. flavospinosus, Brun. .
4. eupogonatus, Biy. . . .
3. annulata, Brun
2. Leptomydas, Gerst
1. indianus, Brun
. 319
. 320
. 320
1. longirostris, Wulp. . . .
6. Geron, Mg
1. arjjentifrons, Brun
Fam. 9. EMPIDJE
Subfam. 1. Hybotince.
. 321
. 325
7 Usia Latr
1. Synecbes, Walk
1. immaculatus, Brun. .
2. biaroti. Bezzi . .
. 327
. 328
. 329
1. sedophila, Brun
2. marffinata, Bnm.. .
SYSTEMATIC INDEX.
Syueches (con.).
3. inaequalis, Brnn
4. minutus, Brun
5. rusticus, Brun
6. palliditarsis, Brun
7. Iratellus, Brun
8. insicmia Tirun. . . .
Page
330
330
331
331
332
332
333
333
333
334
334
334
336
335
336
338
338
339
340
340
340
341
341
342
342
343
343
343
344
345
3-15
346
34(5
347
349
360
351
352
352
363
353
354
355
356
356
367
357
358
359
S59
360
368
:!()!
]
Subfam. 3. Ofiydromiina
1. Leptopeza, M.aeq
Page
362
362
368
363
364
365
366
366
367
367
368
369
370
371
371
372
373
374
375
37C.
377
377
378
378
379
379
380
381
382
382
383
383
383
384
384
385
385
386
386
387
387
388
339
390
391
392
1. vitripemiis, Brun
Subfain. 4. Hemerodromiinee . .
1. Clinocera, ~M.g . .
2.
8.
4
5.
9. fuscipennis, Brun
Syndyas, Lw
1. fluviatilis, Brun
2. obscura, Brun
2. Microdromia, Big
1. dorsalis, Brun
3. Phyllodromia, Zett
] . flavida, Brun
2. indicata, Brun
4. Dolichocepbala, Mncq.
1. parvicellulata»2?«K:/ ..
Harpamerus, Big
1 . signatus, Big
Acarterus, Lw
1 . pallipes, Bezzi
2. fuscipennis, Brun
Hybos My
1. bisetosus, Bezzi
1. septemnotata, Brun. . .
5. Trichopeza, Rond
1 fusca Brun
2. tenuipes, Brun
3. gagatinus, Big
4. apicis, Brun
5. genie ulatus, Wulp . . . .
6. niger, Brun. . .
Subfam. 5. Tachydromiina; . .
1. Tachydromia, My
1. latifascipennis, Bnm. . .
2. ferruginea, Brun
3. nepalensis, Brun
4. gentilis, Brun
5. orientalis, Brun
1 Riiriups Hnni
8. nitens, Brun
9. flavipalpi8, Brun.
10. nigronitidus, Brun
11. brunnipps, Brun
6. Parahybos, Kert
1. flavipes, Brun
Subfam. 2. Empinte
1. Rbamphomyia, My
1. unifasciata, Brun. . . .
2. bimalayana, Brun. . . .
3. griseonigra, Brnn. . . .
9 Tfrrmis T,imn
2. Tachjpeza, Mg
1. palliditibiae, Brun
2 incisa Brun.
3. Drapetis, My
1. variegata, Brun
2. discoidalis, Bezzi . . . .
3. bre'vior, Brun
4. fascifemorata, Brun. . .
5. rotundicornis, Brun. . .
6. rufipes, Brun
4. Elaphropeza, Mncu
8.
4.
] . amplitarsie, Brun
2. albobalteralis, Brun. . .
3. carbonaria, Brun
4. squaniata, Brun
1 . basalis, Bezzi . .
'2. bicoloripes, Brun
3. fulvitborax, Wulp
4. ferruginea, lirun
5. metatarsata, Bezzi ....
6. variegata, Brun
Fam. 10. LONCHOPTERIDJE . .
1. Lonchoptera, My
5. ceylonica, Bezzi
6. griseonigra, Brun
7. centralis, Brun
8. subcilipes, Brun
9. rostrata, Brun
10. raarginata, Brun
11. elegans, Brun
12. iuconspicua, Brim
Hilara Mg .
1. luU'a. Panz. . . . . .
Fan). 11. PLATYPKZIDJE
1. Platypeza, My
1. argyrogyna, Meij
1. conipacta, Brun
2. bares, Walk
3. rutithorax, lirun. . . .
4. peshawarensis, Brun. .
Howlettia, Brun
1. flavipes, Brun
OEDER DIPTEEA.
SUBORDER BRACHYCERA.
INTKODUCTION.
IN the first volume devoted to the Diptera in the "Fauna of
British India" Series, the two great suborders, ORTHORRHAPHA
and CYCLORRHAPHA, into which the order is now generally divided,
were sufficiently characterised, and a comparison was also given
between the NEMATOCERA and BRACHYCERA, into which the first
suborder is subdivided. The earliest writers divided the Diptera
into NEHOCERA and BRACHTCERA, of subordinal rank based mainly
on palpal and antennal characters, and not on those of the
venation or early stages, this classification finding favour until
towards the close of the last century. Nowadays the old series of
hrachycerous families is split at or near the LO^CHOPTERID^;, all
those from the STRATIOMYIDJE to that family being included in the
ORTHOREHAPHA BRACHYCERA ; the remainder, consisting of the
PLATYPEZID^E, PIPTJNCULID^E, SYRPHID^E, CONOPID.E and CEsTRLoa:,
the enormous family MUSCID^E (sensu latissimo), and the pupiparous
Diptera (the last-named forming in themselves a separate group of
uncertain rank) constituting the CYCLORRHAPHA. One anomalous
family, the PHORID^E, is by some placed next the LONCHOPTERID^E
in the ORTHORRHAPHA BRACHYCERA, and by others next to the
MUSCID.E in the CYCLORRHAPHA.
The formation of groups of higher rank than families has
received the attention of many dipterologists of note, and Verrall
gives an excellent resume of Brauer's (1883) and Osten-Sacken's
(1896) views. Verrall himself admitted that there were several
points in Brauer's tables that were not clear to him and others with
which he could not agree, and it may here be observed that
Brauer's primary divisions or suborders of ORTHORRHAPHA and
CYCLOHRHAPHA have been criticised as lately as 1907 by Prof.
Miall. Verrall, in noting this in a footnote, adds, " I quite
agree with this distinguished savant in considering the two
divisions, ORTHORRHAPHA and CYCLORRHAPHA, at present incom-
prehensible" (Dec. 1908). I find myself in the same predicament,
and, personally, biological affinities notwithstanding, I have an
instinctive leaning towards regarding all the families from the
STRATIOMYIDJE to the MUSCID^E, and the PHORID^E, as more closely
related one to the other than are a certain number of them related
2 BRACHYCERA.
to the NEMATOCERA ; in other words, I hesitate to abandon the
" old-fashioned " classification of NEMATOCERA and BRACHYCERA,
pur et simple, with the PUPIPARA as a third group, either of equal
rank with the other two or as a subordinate section of the
BRACHYCERA.
The NEMATOCERA and BRACHYCERA are certainly natural
divisions, and Osten-Sacken believed that the characters distin-
guishing them were of more fundamental importance than those
separating the ORTHORRHAPHA and CYCLORRHAPHA ; he regarded
them as of equal ordinal rank with the CYCLORRHAPHA.
Williston, on the other hand, considers there is " no distinct
limitation between the NEMOCERA and BRACHYCERA,"* but,
according to Verrall, the shape of the anal cell provides an
infallible character in all those cases in which doubt could possibly
arise. It is quite evident, however, that these two groups have
always been perfectly clear to most dipterologists from the.fact that
no dispute has ever arisen as to the group to which any particular
family belonged ; whilst, on the other hand, authors are by no
means in agreement as to where the ORTHORRHAPHA NEMATOCERA
end and the BRACHYCERA, in the restricted sense, begin. How-
ever, these questions of biological affinities amongst the higher
divisions can only be approached by those possessing an intimate
knowledge of all the families, and it Mould be out of place to
attempt any discussion of the subject in a work restricted to the
consideration of a limited number of families.
For the purposes of the present work it will be sufficient to
give, in abbreviated and slightly modified form, Brauer's divisions
of his CYCLORRHAPHA : —
1. Head not flattened, more or less hemispherical,
placed perpendicularly to longitudinal axis of
body; mouth placed below. Eyes always pre-
sent. Dorsal and ventral segments of abdomen
well developed, the connecting membrane con-
cealed. Imagines living; free, non-parasitic .... 2.
Head flattened, placed in longitudinal axis of body ;
mouth similarly placed, projecting forward. Eyes
present or absent. Dorsal and ventral plates of
abdomen very small or absent, owing to pre-
ponderance of connecting membrane. Imagines
parasitic PCPIPARA.
2. Frontal bladder-seam running round only upper
side of base of antennae, narrowly surrounding
the lunule, the bladder -seam being therefore
restricted, the bladder existing or rudimentary.
Jowls not separated from the superiucumbeiit
cheeks. Anal cell long and pointed 3.
* Manual N. Amer. Dipt. 3rd ed. p. 59.
INTRODUCTION. 3
Frontal bladder-seam semicircular, forming above
the lunule a distinctly margined upright arch,
usually running steeply downwards over middle
of face, usually separated and distinctly sepa-
rating the cheeks and jowls at the sides. Anal
cell generally short and often subequal in length
with 2nd basal cell EUMYID.S:.
3. First posterior cell always closed. Vena spuria
present . SYKPHIDJE.
First posterior cell always open. Vena spuria
absent 4.
4. Arista terminal PLATYPEZID^;.
Arista dorsal PIPUNCULHXS:.
The two latter families were left in the same divisional group
by Brauer, but are easily separated by the character given above.
It must be confessed that the table of groups is difficult to put
to practical use, especially by a beginner. The EUMY.IDJE, unfor-
tunately given a family termination, is a combination of the
CONOPIDJE, (ESTBID^, arid MUSCID^E, all of which are regarded in
the present volume as good and separate families.
The PUPIPARA, it is understood, consist of three distinct
families, HIPPOBOSCID^E, NYCTEBJBIIDJE, and STKEBLID^E.
The superfamilies herein admitted are those recognised by
Verrall in his second published volume on British Diptera, and
the following table of them is wholly drawn from that work : —
Table of Superfamilies.
Three equally well-developed pad-like pulvilli.
Species absolutely eremochaetous (i, e., strong
bristles on head and thorax entirely absent) .... EREMOCHJETA.
Two pad-like pulvilli only, the empodium being
sometimes absent or sometimes represented by a
bristly hair. Species not truly eremochtetous . . 2.
Basal cells long, the anal cell being open or long
and pointed, because the branches of the 5th vein
only gradually diverge, and the upper branch is
usually connected with the discat cell by the
posterior cross-vein, though sometimes (BoMBY-
LIID^E, SCENOPINIDJE, etc.) it forms a part of the
lower margin of the discal cell : wing-venation
usually elaborate. Cephalic bristles on frons and
vertex not strongly developed, though sometimes
visible in ASILIDJE. Often large or very large
species and seldom small, and when small
usually with long pubescence among the stronger
bristles 3.
Anal cell short except in a few EMPID^E (HYBO-
TiN-ff;) and shorter than the preceding basal cell
(when that is present), and the upper basal cell
rather long in the EMPID^: only ; 5th vein
with its lower branch recurrent (except in the
HYBOTINJE) and its upper branch quickly lost
4 BKA.CHYCEKA..
in the 4th vein ; 3rd -vein simple except in a few
EMPIM3 ; posterior cross-vein quite absent ;
posterior cells apparently never more than four,
but usually less ; wing- veins altogether few and
simple; alula obsolete. Face bare or only
slightly pubescent in the MICHOPHONA, but
with peristomal bristles in LONCHOPTERIDJE and
side or genal bristles in PHORID^E, but with
no face beard or pubescence as in ASILIDJE.
Cephalic bristles usually strongly developed.
Species always small and distinctly chseto-
phorous • 5.
3. Aerial species, almost always clothed with dense
furry pubescence. Legs thin and only suitable
for alighting, and consequently armed with little
more than spicules (not true bristles) on the
femora and tibise. Eyes nearly always touching
in the c? (holoptic) and never bulged out because
of a sunken vertex TROMOPTEUA.
Pedestrian species, usually armed with strong bristles
and only occasionally (Laphria, Isopogon, etc.)
clothed with dense coarse pubescence ; or (!)ER-
MATINA) bare of both pubescence and bristles.
Legs strong, usually armed with strong bristles
on at least the tibise and tarsi. Eyes almost
always widely separated in both sexes (dichoptic),
and frequently bulging out because of the deeply
sunken vertex 4.
4. Eremochsetous species in the sense that there are
no strong bristles on any part of the thorax or
scutellum DERMATINA.
Chsetophorous species in the sense that there are
strong and usually numerous bristles on the
thorax, scutellum, and legs, or at least on the
sides of the metauotum, except in the few cases
in which dense coarse pubescence is substituted. ENERGOPODA.
5. Wing-veins (including cross-veins) fairly normal, or
else the 2nd and 3rd veins extended almost to
the wing-tip. Antennae with three (or rarely
two) joints obvious, besides any style or arista. . 0.
Wing-veins extremely abnormal and quile distinct
from those of any other Diptera ( except A spistes
in the BIBIONIDJE) ; anterior veins up to the
3rd vein strong and conspicuous, running into
the basal half of the costa, but the subsequent
veins very faint and incomplete and abnormally
directed ; 4th, oth, and anal veins very difficult
to trace. Palpi porrect, not jointed. Eyes
widely and equally separated in both sexes.
Antenna? apparently composed of one large joint,
which bears a very long apical or dorsal arista.
Hind legs long and the femora flattened HYPOCERA.
Wing rounded at the tip ; venation fairly normal,
with the 2nd vein ending considerably before
the wing-tip, and with at least one apparent
cross-vein well out towards the middle of the
wing, causing the discal cell to be present even if
extended back to the wing-base. Face, rnouth-
margin, and jowls without strong bristles.
Thorax rarely with a soft pubescence. Eyes of
the J sometimes touching or approximated.
Imagines always predaceotis MICROPHONA.
Wings pointed at the tip ; venation abnormal, as
the 2nd and 3rd veins end close together almost
at the wing-tip ; no obvious cross-vein, and the
discal cell absent or reduced to a short middle
basal cell. Face with strong bristles round the
niouth-margin. Thorax with bristles but no
pubescence. Eyes widely and equally separated
in both sexes. Antennae short and porrect, with
a long subdorsal arista. Colour always dull
yellowish, brownish, or greyish, but never green
or metallic ACROPTERA.
Verrall incorporates in the above table the sixteen families of
BRACHTCERA which occur in Britain, as well as the seven super-
families, which accounts for characters of minor value often being
added, but it has seemed better to give his actual diagnoses of the
higher groups in full than to attempt to abbreviate them. One or
two terms in the venation and so on are changed for the purpose
of preserving uniformity with ray previous volume. His table
(pp. 14 to 30) is illustrated with typical figures and is a veritable
epitome of the salient characters of the BRACHYCERA.
Coming to families, there has been less difference of opinion
amongst authors than as regards the higher groups, probably
because the characters are drawn solely from the perfect insects.
The exact rank of the CCENOMYID.E and XYLOPHAGID.E as families
has been questioned, and the affinities of the LONCHOPTERID.S; and
PHORIDJE have given rise to considerable discussion. Notes on
these points will be found under the respective family headings.
I follow Osten-Sacken and Verrall in placing the XYLOMYIN^E
subfamily of XYLOPHAGID.E in the STBATIOMYIDJE, and the remain-
ing subfamily, the XYLOPHAGINJE, in the LEPTID^, to which
family also the COZXOMYIDJE are referred as a subfamily.
Some authors have desired to remove the MYOPINJE subfamily
from the CONOPID.E to the MUSCID^; and also to disband the
CEsTRiD-j; altogether and spread the genera amongst the MusciDjE,
but with these views I cannot concur.
Of two other families the taxonomic rank has been occasionally
disputed, the ACANTHOMERIDVE and APIOCERID.E, but as neither
group occurs in the Orient they need not be discussed here. The
former family contains the most gigantic flies known, so far as
bulk goes, measuring up to 40 millimetres in length and 80 milli-
metres in expanse of the wings, and they are confined to the
Neotropical liegion. They are allied to the STRATIOMYID^E,
LEPTIDVE, and TABANID.*:. The APIOCERID.E are akin to both the
MTDAID^E and ASTLIDJE. Both these groups are now admitted to
be of family rank.
BRAC1IYCEKA.
Table of all Families of Diptera except the NEMATOCEKA.
1. Body not of leathery texture. Head free ;
mouth generally well developed, with a
more or less prominent proboscis. Two
palpi and at least two oral bristles present.
Femora approximate at base. Abdomen
distinctly segmented. Oviparous or larvi-
parous ; in perfect state never parasitic on
warm-blooded creatures 2.
Body of leathery texture. Head placed in a
curved excavation of the thorax or bent
back upon the thorax. Mouth very imper-
fect ; palpi absent. Abdomen indistinctly
segmented. Femora at base remote, through
the broad mesothorax ; claws well deve-
loped. Parasitic during pupal and imaginal
stages in warm-blooded creatures (mammals
and birds) 23.
2. Three distinct pulvilli. Absolutely eremo-
chsetous species (i. e., no strong bristles on
head or thorax as compared with pubes-
cence) 3.
Two pulvilli (sometimes obsolete or absent).*
Species not truly eremoehaetous, and often
with dense furry pubescence, or bristly . . 8.
3. Third antennal joint annulated, Eyes in $
normally contiguous or very approximate
(except in most NKMESTRINIDJE) ; in $
wide apart 4.
Third antennal joint not annulated (except
XYLOPHAGINJE). Eyes in $ generally
contiguous 7.
4. Posterior cells not parallel with hind margin
of wing t- " Diagonal " vein never present.
Eyes in $ normally contiguous 5.
Posterior cells parallel with hind margin of
wing. " Diagonal ' ' vein generally present ;
wings often reticulate. Eyes in <5 gene-
rally separated ." Nemestrinidae,
5. Tibiee with at least middle pair spurred [p. 144.
(indistinctly in ACANTHOMKRIDJE). Veins
not crowded together in anterior part of
wing. Ambient vein present (thin in
ACANTHOMKRIDJE). Scutellum unspined. 6.
Tibiae without spurs (except in XYLOMYINJE
and some BERING). Veins crowded to-
gether anteriorly (except XYLOMYIN^:
and BERINJE). Ambient veiii absent or
* Middle one sometimes shorter in NEMESTRINID.E, but always pad-like.
t Some CYRTID,K form exceptions, but the enormous thoracic squamae,
globular bodies, and general appearance of this family at once distinguish
them.
INTRODUCTION. 7
incomplete. Scutelluru often with two or
four (rarely six or eight) spines on margin. Stratiomyidae,
6. Antennae flagellilbrm. Fourth posterior cell [p. 17.
closed. Wing with marginal space of last
submarginal cell at most twice that of
immediately adjoining cells. Squamae
almost obsolete. Perfect insects never
blood-suckers Acanthomeridae.
Antennae normally sickle-shaped, rarely fla-
gelliform or otherwise formed. Fourth
posterior cell nearly always open. Wing
with marginal space of last submarginal
cell at least three or four times that of
immediately adjoining cells. Squamae
very large. Adult females always blood-
suckers ". Tabanidae.
7. Posterior tibiae always spurred. Body gene-
rally elongate, abdomen never balloon-like.
Eyes in <$ contiguous, in $ wide apart ;
head of normal size and shape. Thoracic
squamae normal. Venation as in TABA-
NIDAE, except that the anal cell is some-
times open Leptidae, p. 100.
All tibiae unspurred or with very short blunt
ones. Body short, rounded ; abdomen
often balloon-like. Eyes in both sexes
contiguous ; head very small, inconspi-
cuous, depressed, nearly all eyes. Thoracic
squamae enormous. Venation eccentric and
complex Cyrtidae, p. 157.
8. Basal cells long, at least one-third the length
of the wing. Anal cell (sometimes called
the third basal) open, or long and pointed. 9.
Basal cells short (except in HYBOTIX^E).
Anal cell shorter than preceding cell, when
that is present. Third vein never forked,
except in some EAIPIDJE 18.
9. Posterior cells parallel with hind margin of
wing. Second longitudinal vein (and some-
times upper branch of third also) ending
in first longitudinal near its tip. Antennae
generally clavate at tip. Eyes in tf dis-
tinctly separated Mydaidae. p. 314.
Posterior cells not parallel with hind margin
of wing. Second longitudinal vein ending
in first only in APIOCKRIDJE and some
ASILIDJE ; in all other families ending in
wing-margin. Antennae not clavate at
tip. Eyes in J contiguous or well sepa-
rated 10.
10. Upper branch of 4th vein ending before
wing-tip 11.
Upper branch of 4th vein ending below
wing-tip 12
11. Venation more complex. Five posterior cells,
the 4th closed. Eyes in tf always wide
8 BRACIIT.CEKA.
apart, but less so than in £ . First longi-
tudinal vein receiving others in it before
its tip. Thorax and scutelluni with strong
bristles arid pubescence ................ Apioceridae.
Venation simple and reduced. Three pos-
terior cells. Eyes in <$ contiguous or
very approximate. First longitudinal vein
never receiving others in it. Species quite
destitute of bristles or pubescence ...... Scenopinidae,
12. Five posterior cells. Chsetophorous species, [p. 310.
generally very pubescent also .......... 13.
Three or four posterior cells. Eremochsetous
species, except BOMBYLIIDJE and some
* 14.
13. Vertex of head deeply sunken between the
eyes, leaving the latter very prominent.
Eyes in <^ always separated ............ Asilidae .
Vertex not sunken, therefore eyes not pro-
minent. Eyes in cT contiguous or closely
approximate .......................... Tnerevidae, p. 295.
14. Third longitudinal vein generally forked.
Posterior cross-vein absent. Eyes in $
normally contiguous in typical groups, but
distinctly separated in ANTHRACIN^: ; con-
tiguous in both sexes in Systropus. Often
densely furry species .................. Bombyliidae, p. 173.
Third longitudinal vein never forked. Pos-
terior cross-vein present. Eyes in tf con-
tiguous or approximate, except rarely; but
in CONOPIDJE wide apart in both sexes.
Normally not densely furry species, except
in some SYRPHID^E .................... 15.
15. Proboscis rigid and geniculate. First pos-
terior cell closed ; antennae with apical
style (CONOPIN^E), or first posterior cell
open ; antennae with dorsal arista
(MYOPIN.S:) .......................... Conopidae.
Proboscis soft. First posterior cell closed or
open. Antennae of very various formation. 16.
16. First posterior cell closed. Vena spuria pre-
sent (sometimes very faint). Eyes in both
sexes of normal size and nature .......... Syrphidae.
First posterior cell open. Vena spuria absent.
Eyes in both sexes very large (especially
in PIPUNCUI.IDJE) ; some of the facets
often enormously enlarged .............. 17.
17. Arista dorsal, 3rd longitudinal vein well
curved ; * apical section of 4th vein curved
downwards; hind tarsi never dilated .... Pipunculidae.
Arista apical, 3rd vein nearly straight ; apical
section of 4th vein curved inwards. Hind
tarsi dilated .......................... Platypezidae,
18. Venation abnormal ; t no obvious cross- vein [p. 390.
* Nearly straight in Chalarus.
t Except in Opetia, a non-Oriental genus.
INTRODUCTION*. 9
well out on disc of wing. Eyes in both
sexes wide apart 19.
Venation fairly normal (reduced in DOLICHO-
PODIDJE) ; at least one obvious cross-vein
well out on disc of wing. Eyes in rf con-
tiguous, approximate, or wide apart 20.
19. Second and third longitudinal veins long,
nearly parallel. Wings Idng, tips pointed ;
venation different in the sexes ; no wing-
less species - Lonchopteridae,
Second and third veins united in one rather [p. 387.
short strong vein. Wings short, tips well
rounded ; venation similar in both sexes ;
some species wingless Phoridae.
20. Discal cell united to 2nd basal cell ; basal
cells very short or obsolete. Third longi-
tudinal vein diverging from a swelling
near base of wing; anterior cross-vein at
same spot ; third vein never forked. Pro-
boscis soft. Arista or stvle dorsal or apical.
Abdomen normally 5- or 6-segmented. . . . Dolichopodidae.
Discal and second basal cells separate (except
in a few EMPID^:). Basal cells generally
long enough to be obvious. Second and
third veins diverging well away from base
of wing ; third vein not forked, except in
some EMPIDJE. Proboscis rigid or soft . . 21.
21. Antennal style present or absent ; arista, if
present, generally dorsal. Proboscis gene-
rally rigid, pointed. Abdomen normally
7-segmented. Second and third longi-
tudinal veins diverging at one-third of the
wing ; third vein often forked Empidae, p. 321.
Antennal style never present ; arista, when
present, nearly always dorsal. Third vein
never forked 22.
22. Proboscis rudimentary ; palpi rudimentary or
absent. In larval stage always parasitic
on mammalia '. CEstridae.
Proboscis and palpi always distinctly pre-
sent. In larval stage rarely parasitic on
mammalia Muscidae (s. hitiss.}.
23. Head folding back on dorsum of thorax.
Wingless species. Parasitic on bats .... Nycteribiidae.
Head not folding back on thorax. Winged
or wingless species. Parasitic on birds and
mammals , 24.
24. Antennae reduced. Wings with distinct
parallel veins ; cross-veins often present.
Claws simple. Nearly all the species
parasitic on bats Streblidae.
Antennae generally more elongate. Wings,
when present, with veins crowded together
anteriorly; cross-veins short and near base
of wing. Claws large. Parasitic on
mammals and birds Hippoboscidae.
10 BEACHYCEEA.
The external characters of the BEACHYCEEA and CYCLOEEHAPHA
exhibit as much, it' not greater variation than those of the
NEMATOCEEA. As the introduction to the previous volume on
Diptera dealt with characters common to 4he order generally,
those pages may be consulted in conjunction with the present
summary, which is confined to the BEACHYCEEA and those families
of the CYCLOEEHAPHA treated of herein.
The eyes in the higher BEACJIYCEEA and higher CYCLOBEHAPHA
are generally contiguous or subcontiguous above in the male, and
such flies are said to be holoptic, the eyes in the female being
distinctly separated hy a comparatively broad Irons, but exceptions
are numerous. Diptera in which the eyes are well separated in
the c? are called dichoptic. The CYETID/E have contiguous eyes
in both sexes (with rare exceptions), and in some cases the eyes
are contiguous both above and below ; the MYDATD^, ASILLIXI;,
DOUCHOPODIM:, LONCHOPTEEIDA-, and COXOPIDJE have the eyes
equally conspicuously separated in both sexes. The lower ANTHO-
MYJNJE and practically all the ACALYPTEATA are also dichoptic.
The eyes may be pubescent or bare, this being sometimes a sexual
character, and iu life they are often green (Hcematopotci) or golden
green (Chrysopihts), and in many species of TABANIDJE, ASILID^E
and some other groups bear brown or purple bands. In many
groups the front facets are from just perceptibly to very con-
siderably larger than the rest in the c? , and of uniform and
smaller size in the 5 , but in the PIPUNCULID.E the front facets,
though enlarged more than usual in the d , are yet more enlarged,
and to an enormous extent, in the 5 . In some PUPIPAEA the eyes
are absent ; in CYETID.E and PIPUNCULID.E they occupy practically
all the head, no other parts except the antenna? being visible in
profile in some species of both families.
Three ocelli are normally present in all the BEACHYCEEA, but
they are occasionally absent (CONOPIN^E).
The lower part of the head, or epistome, exhibits a wide range
of shape and size, from being barely or not at all produced beyond
the eyes in profile either forwards or downwards (as in most
PIPUNCULIDJE) to forming a long strong conical rostrum (Nemotehts
in STEATIOMYIDJE ; Khingia, Volucella, Lycastris in SYEPHIDJE).
The face frequently has a central bump, as in most SrEpHiD^E, or
it may be less prominent, gently curved, or flattened. The mouth-
parts attain their highest development in the TABANID.E, a family
in which the females of all the species have the reputation of
being blood-suckers. The parts of the female Tabanid mouth
consist of the labiuin or lower lip, a pair of maxilla? and of
mandibles, strong piercing organs, and a pair of large, fleshy,
conspicuous, approximately conical maxillary palpi.* The pro-
boscis in the BEACHYCEEA is usually soft, and formed for sucking
the juices of plants or animal matter, but it is not infrequently
of enormous length, longer than the whole body and of extreme
* Compare ' Indian Diptera,' i. p. 18.
INTRODUCTION. 11
slenderness, and in this state either adapted for piercing
the skin (PANGONIA) or for imbibing the juices of flowers
(NEMESTRINID.E, CYRTIILE, and many BOMBYLID^E). Occasionally
it is long, rigid, and geuiculate (CONOPID.E). Biting flies are
distributed throughout many brachycerous and cyclorrbaphous
groups. S>/niphoromyia, in LEPTIDJE, has been definitely proved
to bite man severely on occasions ; the TABANIILE generally confine
their unwelcome attentions to the larger mammals, though one
genus, Hcematopota, widely distributed throughout the world, its
natural home being apparently Africa, is very prone to attack
man. The strong horizontal horny proboscis of the Stomoxys
group of true Muscinids is well known, as are also the tsetse-flies
(Glossina) inhabiting Africa, these being the carriers of sleeping-
sickness. Philtematomyia, a genus of Muscinid flies, possesses a
circlet of teeth-like projections at the tip of the proboscis with
which, even if unable actually to pierce the skin, it can easily
enlarge an abrasion already made. The proboscis in not a few
genera is rudimentary, in some almost obsolete, as in (EsTRiDTE,
the members of which family cause cattle instinctively the greatest
terror on their approach for the purpose of depositing their eggs
on the animals' mouths, nostrils, etc. An isolated case occurs
here and there of a genus with a strong rigid proboscis in a group
in which the organ is normally soft arid fleshy, as Drymeia in the
ANTHOMYINJE. The ASILID^E and EMPIIXE are two other extensive
families generally possessing a rigid horny proboscis, with which
they easily pierce and carry off their prey, consisting of softer
Diptera and other insects. The labella, sometimes called " sucker-
flaps," vary in size, shape, and strength, sometimes being small
and soft, in other cases acting as organs of attrition to rub off
small particles of food.
• The palpi, apart from those of the NEMATOCERA, consist usually
of two joints, in some highly specialized families of one only, these
being the maxillary pair, the labial palpi being considered to be
absent in the Diptera. Though large and prominent in some families
(TABANID^), it is often difficult in others (BOMBYLILU^E, THEUE-
VID^E, ASILULE) to examine them without removing the dense
furry or bristly pubescence characterising these groups. However,
in most of the BRACHYCERA the location of any genus or species
can be readily ascertained without reference to these organs.
They are normally porrect, i. e., directed horizontally forwards,
especially in the blood-sucking Muscids, but frequently project
more or less downwards, and they may be present or absent,
respectively, in genera of considerable affinity, whilst in the
CYRTIDJE they are generally obsolete.
As regards the antennae, a gradual change takes place from the
many-jointed elongate flagellum of the NEMATOCERA to the compact
three-jointed form of the bulk of the remainder of the Diptera,
the 3rd joint often being the only conspicuous one.* In the
* For comparative notes on the antenna; in NEMATOCERA and BRACHTCERA,
see ' Indian Diptera,' i. p. 20.
12 BRACHYCERA.
earlier families of the BRACHYCERA (STRATJ.OMYID.E, TABANIIWE,
some LEPTHLE, etc.) the third joint is usually anuulated, that is,
faintly separated by more or less distinctly impressed lines into
several divisions, generally eight or five, and in this character
considerable resemblance is apparent to the nematocerous families
BIBIONID^E, SiMULiiDJE, and liiiYPHiD^. These annulations in
subsequent familes of BIIACHYCERA gradually disappear in an
obviously single elongate joint, as in many BOMBYLIID^;, THERE-
TID^E, and ASILIDJE, with or without a more or less distinct one-
or two-jointed apical style ; and in later families still, the 3rd
joint is further shortened and may bear either a style or arista.
The arista, the bristle-like hair situated on the upper side of
the 3rd joint, makes its appearance in the very first brachycerous
family, the STRATIOMYID^,* and is present in one or more genera
in probably all the subsequent families, with a few exceptions
(TABANIDjE, ACANTIIOMERIUJE, SCENOPJUSTD^, MYDAIDjK, and APIO-
CEKIUM), and of these, all except the first are groups of quite
limited extent. The arista is at first (STRATIOMYID./E, LEPTID^E)
mainly apical or subapical, afterwards either apical or dorsal in
the same family, and, finally, mainly dorsal (DOLICKOPOUID.E), which
is its normal nature in the remaining brachycerous families and
in the CYCLORRHAPHA. The arista may be quite bare, pubescent to
a varying degree, or conspicuously plumose. In the great majority
of BRACHYOERA the two basal antennal joints are differentiated
from the 3rd in the na.ture of the scapal joints in NEMATOCERA, and
are usually short, bearing short bristles, cylindrical or, especially
the 2nd, more or less cup-shaped ; but in the 3rd joint the most
varied forms exist even in a single family, this being especially the
case in the STRATIOMYJD^E, BOMBYLIIDJE, and SYRPHIDJE. Briefly,
the 3rd joint may be elongate-cylindrical (annulated or not), strap-
shaped, filiform, tapering, conical, tooth-shaped, oblong, rounded,
oval, renit'orm, globular, or of irregular shape. In one aberrant
genus (llhachicerus in LEPTID.E) it is strikingly pectinate, recalling
Pstlliopliora in the NEMATOCERA, whilst in Talnrocera, a genus
of North American Tachinids, the 3rd joint in the female is
of an extraordinarily fantastic shape. In the CYCLORRHAPIIA the
two basal joints are normally as in the BRACIIYCERA, though
the 3rd joint again shows the same extreme range of variety in the
two largest families, SYRPHID.E and MUSCID^E, but remains of
remarkably uniform, though peculiarly characteristic, form in
others (PIPUNCULIDJE, PLATYPEZIDJEJ. In these groups it may
bear a short style or an arista. The 1st and 2nd antennal joints
sometimes depart from their more or less general form, the
former being exceedingly long and cylindrical in Systropus and
conspicuously so in Stmtiomyia, the 2nd showing unusual develop-
ment in certain genera.
Coming to venation, it will be observed that in the BRACHYCERA
* The only nematocerous family in which the arista appears is the OUPIINE-
piiiLiD.E, a group of about hall' a dozen species, and as it consists of several
joints it is perhaps not truly identical.
IXTKODCCTION. 13
several characters adopt nearly the opposite form to that in the
NEMATOCEEA. Of these perhaps the principal is the anal cell,
which is always wide open in the NEMATOCEBA, normally widening
towards its tip, whereas in the BBACHYCEBA, it normally contracts
towards its tip, frequently very strikingly so ; and in a large
number of genera it is completely closed more or less near the
wing-margin, either by the gradual approximation of the 5th and
6th veins or by the lower branch of the former turning down,
frequently at ar sharp angle or well-rounded curve, often in the
latter cases closing the cell some distance from the margin of the
wing. In the higher families of BBACHYCEBA the anal cell is very
elongate, extending generally to the wing-margin, but in sub-
sequent families, notably the EJIPIDJ:, it is considerably shortened
through the almost or quite recurrent ending of the lower branch
of the 5th vein in the anal vein at or before the middle of the
latter, whilst in DOLICHOPODIDJE and LOXCHOPTEBIP^; it is shorter
still. It increases in length again in two out of the three first
families of the CYCLOBBHAPHA, though it is short in one of them
(PLATYPEZIDJE), whilst in the higher Muscids it is always com-
paratively short, and in the lower MUSCIDS normally short or
absent, although conspicuous exceptions occur (some OETALIN^E
and TBYPETIN.E, for example). In the CONOPID^;, one of the two
families* that may be regarded as forming the transition from the
BBACHYCEEA-like CYCLOREIIAPHA to the MUSCIDJE (s. latiss.) it is
long in the CONOPIXJE and short in the MYOPI^E, which tends to
prove the intermediate nature of the COXOPID.!.
The auxiliary and 1st vein are generally obviously present,
separate and simple, but in some groups of ACALYPTEATA they
are frequently coalescent.f The 2nd vein is nearly always very
prominent and simple ; the 3rd, which is nearly always simple in
NEMATOCEBA, is normally forked in the higher families of BEACHY-
CEEA. The 4th and 5th veins in these latter families are generally
both forked, but in the highest three families of the CYCLOREHAPIIA
(PLATYPEZIDJE, PIPUNCULLELE, and SYEPHiDJE) there are normally
only three endings to the two veins, in which cases it is the 5th
vein that is considered forked, its upper branch forming part or
the whole of the hinder side of the discal cell. This cell in the
NBMATOCEEA is confined to the TIPLTLID.E and BHYPHID.E — that is
to say, the two families generally placed next to the BEACHYCEEA,
in which latter the cell is a standard feature, though exceptions
occur (some EMPIIXE), — and it is prominently retained in the
calyptrate Muscids and the bulk of the ACALYPTEATA. The
anterior cross-vein is nearly always present, though rudimentary
in some STBATIOMYID.E ; bufc the posterior cross-vein disappears
after the higher families are passed, if the theory of the forked
5th vein be upheld.J
* The other being tiie CEsriun.E.
t They are then called the 1st vein, following Schiner.
J It may as well be stated here that I am by no means convinced that this
is the correct interpretation.
14 BRACHYCERA.
The brachycerous venation attains its most perfect form perhaps
in the TABANID.E, and it is elaborated most in the ASILIDJE,
NEMESTRINID.E, MiDAio.£, and CYRTiD*! (despite the latter
family also exhibiting some very simple venational types), being
quite eccentric in the last three families, many of the longi-
tudinal veins in the hinder part of the wing running more or
less pai-allel to the hind margin; whilst in certain genera of
NEMESTRINID^ an additional character is the net-like system of
very numerous small cross-veins. An isolated instance of a cell
being subdivided hy numerous cross-veins occurs in Lycastris
(SYRPHID^), in which the subcostal cell is so divided.
After the BRACHYCERA the venation becomes more simplified,
and is, speaking broadly, remarkably uniform in type throughout
the MUSCID.S, in the higher groups of which (Tachinicls, Dexiids,
Sarcophagids, and Muscinids) the 1st posterior cell is quite or
nearly closed by the upturned apical section of the 4th vein,
whilst in the ANTHOMYINJE and the various subfamilies forming
the ACALYPTRATA that cell is broadly open. Numerous minor
differences occur in these groups, but the type of venation
remains the same. Passing on to the PUPIPARA, some genera
may be observed to possess peculiar venation, but they need not
be discussed here.
The general form of the body exhibits the widest diversity.
The head may be variously proportioned to the body, perhaps
attaining its minimum of size in the CYRTIDYE and its maximum
in PIPUXCULID.E, but normally it is approximately of the same
width as the thorax and usually more or less semicircular. The
thorax and abdomen both vary greatly in length and breadth in
relation to one another, and range from the flattened form in some
STRATIOMYIDJE to the humped thorax in many EMPID.T; and to the
globular or balloon-like abdomen in most CYRTID.I, the whole
body in many species of the latter family being practically
spherical. As regards vestiture, a short pubescence over the
greater part of the thorax and abdomen is common to the vast
majority of genera in BRACKYCERA, a few restricted groups being
for all practical purposes absolutely bare (Scenophins). The earlier
families are erernochsetous — that is, possess no distinct bristles of
a chaetotactic nature on the head or thorax ; but such bristles
gradually appear in successive families, as witnessed by the apical
spurs on the tibiae in LEPTID.E, spicules or stick-like spines on the
legs in some BOMBYLIIDJE, and real bristles and spines in that family,
in DOLICHOPODID^, EMPID.E, and THERETID;E, culminating in the
powerful chsetotactic system in most ASILIDJE ; after which these
characters gradually diminish or practically disappear (as in
PIPIINCULIDJE and SYRPHIDJS), reappearing and attaining probably
their greatest development in the order in the TACHININ.E, the
highest group of the MTTSCID^E («. latiss.). The pubescence,
always soft in STRATIOMYIDJE, attains its greatest development
perhaps in BOMBYLIID^:, THEREYID.E, and ASILIDJE ; and true
INTRODUCTION. 15
scales, covering sometimes extensive parts of the body or legs, are
common in many genera of BOMBTLIID.S and TIIEREVID^E.
The vestiture of the legs varies from practical bareness, through
the whole gamut of soft and fine to a long and ragged pubescence,
bristles, spines, scales, and tooth-like serrations (the latter generally
confined to the underside of the femora), all of them often forming
good generic or specific characters. As a rule, aerial species (that
is, those that are frequently on the wing and addicted to hovering)
possess soft pubescence on the body and weak legs devoid of
strong bristles, as they are presumably employed mainly for
alighting ; whereas, on the contrary, pedestrian (which frequently
means predatory) species mostly possess strong bristles or spines,
and powerful legs capable of running swiftly and surely and of
holding their prey when captured, and possessing comparatively
reduced powers of flight. Compare, for instance, a Bombylius
with an average Asilid. A peculiar character called " touch
hairs " is present in certain genera of LEPTID.E, TABANID^;, and
some other widely divergent groups, and this is dealt with under
the genus Leptis.*
The genitalia in the BRACIIYCBRA are usually more or less con-
cealed in most families or are of simple form, but they afford in
the <5 good taxonomic characters in ASILID.I: and DoLicHOi?ODiD^E,
being fully free in the latter family ; but the genitalia are always
more uniform in the $ and of little value lor classification in
that sex.
Life Histories.-^
Although the metamorphoses of the Diptera in general are still
but imperfectly understood, considerable progress in their study
has been made since the older writers (Keaumur, JJe Geer, etc.)
first broke the ice, and the early stages of a comparatively large
number of common species have been worked out. Taking the
order as a whole, the larva may be described as maggot-like,
though extreme forms of shape are far from uncommon. It
generally consists of a head segment and twelve others, of which
the three anterior ones are presumed to represent the thorax
of the perfect insect. The head segment is often so indefinitely
presented that Brauer sometimes uses the expressions " head
capsule" and "jaw capsule." Even when obviously present the
head is sometimes most disproportionately minute or placed below
the anterior end of the body so as to be invisible from above ;
also not infrequently in the flesh-feeding maggots no head can be
detected with certainty. These latter forms are called acephalous,
* These notes on vestiture are compiled chiefly from Verrall.
t These remarks on the early stages are wholly culled from Dr. Sharp's
paper (v. Verrall, p. 31). This article should certainly be studied by nil
interested in the subject. In addition to a preliminary general resume of
brachycerous larvae, it offers a translation of Brauer's classification of the
fumilies in BRACHYCERA. based on larval characters, and, lastly, more detailed
information is given on each family or group of families separately.
1G BRACIIYCERA.
whilst at the other end of the scale, i. e., those larvae with unduly
large heads, the term eucephalous is applied. The eucephalous
kinds are the less common, but they are the normal form in the
nematocerous family CULJCIM:. The head in the larvae of the
majority of brachycerous families exhibits an intermediate type
between the extreme forms. Many kinds of larvse have retractile
heads, the front part being completely withdrawable inside the
following segment or segments,
Kegarding tlie mouth-parts Brauer, roughly speaking, described
the lanse of the NEHATOCERA as having the jaws moving
horizontally, except in cases where the mouth-parts are quite
rudimentary, when the larva is peripneustic and 13-segmented.
He claims for the BRACIIYCERA vertically moving jaws — sucking
and boring groups coming also under this division, the head being
not well developed. Brachycerous lar\ 03 with rudimentary mouth-
parts are amphipneustic or metapneustic, with ten to twelve
obvious segments.
The number of body-segments is frequently by no means easy
to ascertain, Dr. Sharp stating that there is "no morphological
criterion yet discovered by which the segments can be numbered."
In some cases, of course, they are obvious enough, whilst in others
there are what have been termed intercalary segments, apparently
consisting of the undue development of the integument between
the orthodox segments, the larva? of the THEREVID.S: and SCEXO-
PIKIM; being instances in which tlie apparent number of segments
approximates to about twenty. The larva with the smallest
number of segments appears to be that of the nematocerous
family BLEPHAROCERIDJE, in which only five or six are present.
Locomotion is usually effected by means of pseudopods, as
jointed legs are unknown through the whole order of the Diptera,
these pseudopods being transverse swellings on the under (or
sometimes both upper and under) surface of the body, and they
may also be furnished with rows of bristles or small wart-like
projections, placed on a varying number of segments according to
the species.
The spiracles in Dipterous larvae consist of a posterior pair
placed near or at the tip of the body, where there is often some
special development for their protection. Larvae with only one
pair of spiracles, the posterior ones, are called metapneustic ; but
if in addition there is a pair of anterior ones placed a little behind
the head, they are termed amphipneustic: whilst when intermediate
spiracles are also present, making, so to speak, three sets in all,
they are known as peripneustic.
The pupa-case is ordinarily the dried larval skin, the imago
escaping through a longitudinal split occurring between two
transverse splits, normally on the ord and 5th segments re-
spectively.
Further detailed information will be found under the respective
families or genera concerned.
STKATIOMYIIXE 17
Family STRATIOMYID^E.
Head hemispherical or flattened, rather short, usually ahout as
broad as thorax, generally bare. Eyes contiguous in rf (occasion-
ally very narrowly separated, Saryus, etc.), wide apart in $ ; if
hairy in <3 , nearly always bare in $ ; sometimes upper facets
enlarged in tf , all facets equal in $ ; posterior orbits often very
broad aiid conspicuous. Jb'rons seldom prominent, but lower part
of head forming a conspicuous snout in some genera (Nemotelus] ;
three distinct ocelli. Proboscis usually short and fleshy, some-
times rudimentary, occasionally long aud thin (Nemotelus), rarely
conspicuous ; palpi 1- to 3-jointed, iuconspicuous. Antenna?
porrect, approximate at base, showing, perhaps, greater diversity
of form than in any other family of Diptera, 3-jointed ; 3rd joint
annulated, forming apparently several (up to eight) annulations ;
an apical style, often very large, or an apical or subapical arista
often present.
Thorax generally quadrangular or roughly obloug or oval, more
or less pubescent, moderately or considerably arched ; scutellum
semicircular or broadened, unarmed or with 2, 4, 6, or 8 spines,
occasionally produced into a prominent erect bluut coue or spine
(Monaco, nthomy ia, Ceratothyrea). A strong spine on each side of
thorax in some genera (Ephippiwn, Negritomyia).
Abdomen of five to seven, sometimes eighr, segments ; of very
varied shape, globular, much broader than thorax (Pachygastei ,
Acanthina, Cyphomyia) ; broader and shorter than thorax (Culena) ;
quadrate (titratiomyia, Odontomyia) ; oblong or oval, and about as
long as thorax (Evaza, Tinda, aud many others) ; or very elongate,
linear, cylindrical, or flattened (iSargus, Jlermetia, Stratio-
sphecomyia). Genital organs usually inconspicuous.
Legs normally of moderate length and strength, or rather weak ;
destitute of bristles or spines (except for a serration below hind
femora in some XYLQMYIXJE) ; generally bare or with pubescence
mainly confined to the femora ; tibia? unspurred, except in
XYLOMYIN^ and some BEKINJE.
Wings generally with the characteristic venation of the family
(except XYLOMYIN.E and BEHIND) ; anterior veins (subcostal, Isi,
2nd, and 3rd longitudinals) crowded together in fore part of wing
and the posterior veins very faint, often not reaching wing-margin.
In typical genera costa nob reaching wing-tip, ambient vein
absent ; 3rd vein simple or forked, fork occurring at some distance
beyond discal cell, both branches ending in costa before wing-tip ;
discal cell always present, lower side often formed of upper branch
of 5th vein, in which case posterior cross-vein absent ; anterior
cross-vein present, not distally further than middle of discal cell,
joining, except in SAUGINJE, 3rd vein to discal cell. In. CLITEL-
LARIIXJE, 2nd vein apparently anastomosed with 3rd or may be
considered absent, lu SARGIXJE, anterior cross-vein joining
praefurca and discal cell, as 2nd vein originates much later than
c
18 STRATIOHYID^E.
usual. Prsefurca originating in this family about opposite base
of discal cell (except in BEKINTE and XYLOMYIN.-E), whereas in all
other families of BRACHYCERA it originates much sooner. In
BERHNOE and XYLOMYINJE all the veins attain the wing-margin ;*
praefurca quite distinct ; costa nearly or quite reaching wing-tip ;
ambient vein more or less distinct: 4th posterior cell closed in
XYLOMYIN^E. In all subfamilies, membrane of wing always ribbed
or wrinkled ; squama variable in size, thoracic pair sometimes
absent.
The STEATIOMYID^E are invariably bare of bristles or spines
(except the side thoracic spines in a few genera), but often possess
dense soft pubescence extending over the whole body and the
femora. The crowding together of the anterior veins and the
extremely wide variation of form in the antennae (perhaps greater
than in any other familv of Diptera) are their principal characters.
In the BERING and XYLOMYIN^E the venation takes the form of
the other allied families and the antenna? are more uniformly
constructed, elongate or subconical, with distinctly annulated
3rd joint. The family is closely allied to the LEPTID^E and
TABANID^ and to the small family of gigantic flies, the ACAN-
THOMERJD^:, known only from South America.
Life-Mstory. — The larvae show considerable variation, though
easily recognised when once seen, and those of some species are
popularly known as rat-tailed maggots, as are also those of the
ERISTALINYE. They are 12-segmented, including head, herni-
cephalous or conical, very elongate, rather leathery, and more or
less flattened. They live in earth, dung, semi-liquid or decaying
matter, and many are aquatic ; in the latter case the eggs are laid
on plants above the surface of the water. No pseudopods ;
terminal spiracles not visible, placed wholly inside a chamber
which has generally a closed or obscure aperture. Pupa coarctate,
in whicli character this family differs from all other BRACHYCERA,
the imago escaping through transverse slits on the 3rd and 5th
segment and a longitudinal slit between them.
Many species hibernate in North Europe, but very possibly do
not do so in the Tropics. The life-histories of several European
species are well known. The imagines frequent grass and low
herbage in marshy situations and, generally speaking, are sluggish
in habits, though some have a very rapid flight in hot sunshiue.f
Table of Subfamilies.
1. Abdomen of at least seven obvious segments. ... 2.
Abdomen of five, or at most six, segments 3.
2. Prsefurca beginning at about opposite base of
discal cell ; 4th posterior cell wide open .... BERING, p. 89.
Prtefurca beginning considerably before base of [p. 96.
discal cell ; 4th posterior cell closed XYI.OMYINJE,
* Except for an abortive branch of the 4th vein in some BEHIND.
•*• Additional and more detailed information occurs under the various sub-
families or genera concerned.
STRATIOMYIDJE. 19
3. Fourth vein with only two terminations (i.e.,
only two true veinlets issuing from discal cell,
the apparent 3rd veinlet being the upper [p. 19.
branch of the 5th vein) PACHYGASTRINJE,
Fourth vein with three terminations (i. e., three
true veinlets issuing from discal cell); upper
branch of 5th vein either separated from discal
cell by the posterior cross- vein being present,
or in punctiform contact with that cell when
posterior cross-vein is absent 4.
4. Anterior cross-vein joining prrefurca to discal cell. SARGIN^E, p. 72.
Anterior cross-vein joining 3rd vein, as usual, to
discal cell ' 6. [p. 56.
5. Posterior cross-vein present STRATIOMYIN.S:,
Posterior cross-vein absent CLITELLARIINJE,
[p. 38.
The line of demarcation between the two last subfamilies is
very indefinite, and, as more than one author has suggested, they
might well be united into a single subfamily. The posterior cross-
vein is often a variable quantity even in groups in which it is
technically present — for instance, it is absent in Ptecticus australis,
Sch., though present in the other Indian species. Williston sinks
the alleged subfamily HERMETIIX^E in CLITELLARIIK/E and his view
is adopted here.
Subfamily PACHYGASTHIN^.
Head comparatively small ; eyes generally bare ; antennae of
very varied form, branched (Ptilocera), elongate (Tinda, etc.)
filiform (Stratiosphecomyia) , globular (Pachyyaster), or of irregular
shape.
Thorax subquadrate, oval ; scutellum unarmed or with two or
four marginal spines ; sometimes produced upwards or hind wards
into a strong blunt cone. Abdomen 5- or 6-segmented, of very
varied shape ; often exceedingly broad, much broader than thorax,
globular ; sometimes oval or oblong, or much elongated and more
or less cylindrical. Legs comparatively short, slender, smooth.
Wings with 3rd vein simple or forked ; posterior cross-vein absent
through 5th vein forming hinder side of discal cell ; 4th vein with
only two veinlets. Four posterior cells, united on wing-margin.
The essential character of this subfamily is the presence of
only three veins apparently issuing from the discal cell, the lower
one being the upper branch of the 5th vein. Mostly bare
species.
Table of Genera*
[p. 33.
1 . Antennae branched PTILOCERA, "Wied.,
Antennae of various forms, never branched. 2.
For table of Oriental genera, vide Brunetti, Rec. Ind. Mu?. i, p. 89.
c2
20 STRATIOMYIDJK.
2. Eyes pubescent . ACAXTHIXA, Wied.,
Eyes entirely bare 3. [p. 26.
3. Scutellum unspined, or produced conically
upwards 4.
Scutellum 2- or 4-spined, never conically
upturned 8,
4. Abdomen elongate, linear, much longer
than thorax ; antennae conspicuously [Brim., p. 36.
long, filiform STRATIOSPHECOMYIA,
Abdomen very short, thick, rounded ;
antennal 3rd joint rounded or shortly
elongate, never conspicuously so 5. [p. 21.
5. Third vein unforked ZABRACHIA, Coq.,
Third vein forked 6. f p. 20.
6. Scutellum not upturned PA CHYG ASTER, Mg.,
Scutellum distinctly couically upturned . . 7. [p. 24.
7. Antennae filiform, longer than head ACRASPIDEA, Brauer,
Antennal 3rd joint large, round MONACANTHOMYIA,
[Brun.,p. 23.
8. Scutellum with two very short spines .... WALLACEA, Dol.,
Scutellum 4-spined ...." 9. [p. 24.
9. Third antennal joint with elongate style . . TIXDA, Walk., p. 29.
Third antennal joint with pubescent arista. 10.
10. Abdomen barely broader than thorax .... EVAZA, Walk., p. 31.
Abdomen much broader than thorax CRASPEDOMETOPON,
[Kert., p. 28.
Genus PACHYGASTER, My.
Pachygaster, Meigen, 111. Mag. ii, p. 266 (1803).
Tappo, Latreille, Hist. Nat. Crust. Ins. xiv, p. 343 (1804).
Neopachyffcater, Austen, Ent. Month. Mag. xxxvii, p. 245 (1901).
GENOTYPE, Nemotelus atra, Panz. ; by original desiguation.
Head semicircular ; eyes large, contiguous in rf (except in rare
instances), practically bare, posterior orbits broad. Proboscis and
palpi small and inconspicuous. Antennae short, 3rd joint rounded,
with four indistinct anuulations and long subapical pubescent
urista. Thorax large, broadened hindwards, transverse suture
deep, V-shaped, behind which the surface is somewhat inflated on
each side ; scutellum subtriangular, unspined. Abdomen much
broader and barely longer than thorax, subglobular, 5-segmented ;
genitalia small. Legs simple, nearly bare ; femora and metatarsi
long. Winys with 1st and 2nd veins short, the latter ending
barely beyond middle of costa ; 3rd vein normally forked, both
branches ending well before wing-tip ; discal cell large ; 4tli vein
with only two terminations ; posterior cross-vein absent, as 5th
vein forms hinder side of discal cell; anal cell closed far from
wing-border.
lianr/e. Europe, North America, Orient.
Life-history. That of several European species is known. The
larva lives in rotting tree-trunks and probably ft-eds either on
the decaying wood or the frass produced by wood-boring Coleoptera.
1'ACHYGASTER. — J5ABIIACHIA. 21
There is only one Indian species in the restricted genus Pacliy-
g aster, namely, P. annulipes, =p. nov. Macquart's P. rujitarsis*
from Pondicliery, cannot belong to this subfamily, as, according to
the plate, ifc possesses three endings to the 4th longitudinal vein,
apart from the upper branch of the 5th vein.
1. Pachygaster annulipes, sp. nov.
2 . Head : irons between one-third and one-fourth of the head,
shining black ; face, which is conspicuously invaginated, with
greyish dust ; back of head shining black. Antennae with 1st joint
black, the remainder wholly bright yellow, with long arista.
Proboscis dark. Thorax and scutellum shining black, dorsum of
both with very short yellowish pubescence, which does not quite
extend to the anterior margin of the former. Abdomen shining
black, witli sparse short pale yellowish pubescence ; venter black,
with sparser but slightly longer pubescence. Legs yellowish to
tips of tarsi ; a rather broad subapical black or blackish brown
ring on all the femora. Winr/s pale yellowish grey ; stigma
darker yellow ; halteres yellowish.
Length^ 2^ mm.
Described from a single $ from Margherita, Assam.
Type in the Indian Museum.
Genus ZABRACHIA, Coq.
Zabrachia, Coctuillett, Bull. X. Y. State Mus. xlvii, p. 585 (1901).
G-EXOTYPE, Zobraclda polita, Coq. (North America).
Differing from Pachyc/aster in the unforked 3rd vein. Some
degree of latitude must be allowed in my revised Ariew of this
genus, in which the species are chiefly united by the above
primary character. Eyes in d contiguous (minutissima, Zett., a
European species) or narrowly separated (annulifetnur , albipes).
Scutellum not upturned (minutissima), barely upturned and
thickened (albipes), or produced upwards into a short blunt cone
{(tnwulifemur). Third antennal joint transversely elliptical (jpoliUt\
the type-species from New York) or as in. Pad tyy aster.
Range. India, North America.
Z. albipes has wholly pale yellow legs, while Z. annidifemur has
a broad dark band on all the femora.
2. Zabrachia albipes, Brun. (PI. I, figs. 1, 2.)
PacJiyyaster albipes, Erimetti, Ilec. Ind. Mus. i, p. 102, $ (1907).
Pactitjyaster infurcata, de Meijere, Tijd. v. Ent. 1, p. 232, 5 (1907).
$ . Head and front shining black, with a brilliant white streak
on each side of lower part of head ; antennas and proboscis orunge-
* Dipt. Exot. Supp. i, p. 57, pi. vi, fig. 3 (1846).
22 STRATIOMIIPJE.
yellow. Thorax and abdomen shining black, with short, sparse,
silvery grey hair, which is a little thicker and mixed with gold
hairs on dorsutn of thorax ; abdomen uniformly black. Leys pale
yellowish white, tips of tarsi faintly blackish. Wings quite clear ;
veins on anterior part and the stigma pale yellowish ; halteres
white.
Length, 2-2 1 mm.
Described from four 2 9 in the Indian Museum from Calcutta.
My original description was insufficient, and my not noting the
unforked 3rd vein probably prevented de Meijere from recognising
my species in his infurcata, described from Java. The white tomen-
tose stripe on the side of the thorax mentioned by de Meijere is
present, though liable to be not readily distinguished from the
general whitish pubescence of the pleurae.
3. Zabrachia annulifemur, sp. nov.
J $ . Head shining black, bare ; frons at level of antennae about
one-fifth of the head in d1 , in $ one-fourth width of head, sides
parallel except near vertex and towards antennae ; visible only
from above, a pair of moderate-sized oval silvery-white spots
placed lengthwise, nearly contiguous, one above each antenna ;
face with broad white-dusted band contiguous to lower inner eye-
margins. Antennas brownish orange, upper side of 3rd joint
blackish at tip. Thorax, scutellum, and inetanotum shining black,
all with very short whitish pubescence ; scutellum produced
upwards into a short blunt cone. Abdomen shininsr black ; dorsum
and venter with similar pubescence to that of thorax, but less
obvious ; genitalia brownish orange, with a few stiff hairs. Legs
yellowish, front coxae more brownish at base ; all femora with a
broad blackish ring leaving base broadly, and tips narrowly, pale.
Wings colourless, veins and stigma pale yellow ; halteres yellowish,
clubs shining milk-white.
Length, barely 3 mm.
Described from three tf rf in the Pusa collection (including
type sent to British Museum) from Annandale, Simla District,
"on leaves," x. 1911 ; and one $ in the Indian Museum from
Simla, 7000 ft., 9. v.09 (Annandale}.
This species must be near the femorata variety of de Meijere's
P. infurcata, but should be distinct. In P. albipes the
<5 5 . Head with eyes in J with dense yellow pubescence
above and brown pubescence in front, nowhere actually touching
but extremely near each other for a considerable distance, and
separating upwards very gradually, leaving a very narrow elongate
greyish triangle below the large black ocellar triangle which fills
the space from eye to eye ; above this the rather large vertical
space is orange-brown with a few black bristles ; frontal triangle
smooth, shining, bare, orange ; proboscis, palpi, and labella all
bright orange-brown ; antennae with first two joints orange, 8rd
ACA^TIIINA. 27
blackish, microscopically pubescent, sometimes orange at extreme
base with four distinct ammlations and a short two-jointed style ;
occiput considerably concave, dark, with a little grey pubescence.
In $ frons -g- width of head, barely broader at vertex and lower
end ; frontal triangle large, very prominent, shining, bare, bright
orange-brown, as is also the very broad posterior border to the
eyes; a small brown spot on frons just above the suddenly
widened frontal triangle. Thorax dark shining blue, punctate,
with microscopic .black bristles ; in c? a single median stripe of
yellow pubescence, not reaching anterior collar but leaving a shining
black spot bare of pubescence, the stripe extending hindwards to
middle of dorsum ; in $ two very narrow longitudinal median stripes,
rather approximating to one another, formed of minute whitish
pubescence, continued beyond suture a little, where they diverge
and broaden, reaching posterior corners of dorsum ; a moderately
broad transverse stripe of similar pubescence along suture,
narrowest in middle ; a patch of short golden yellow pubescence
on each shoulder and along anterior margin; pleurae as dorsum,
similarly punctate and with a little grey or whitish pubescence,
which is silvery white and thickest above front coxa3, on meso-
pleura just below wing, and on lower part of sternopleura.
Scutellum concolorous, similarly punctate, and with greyish
pubescence ; spines brownish, rather short. Abdomen dark shining
blue, punctate as thorax ; a little grey pubescence about front corners
and sides and denser and longer about centre of hinder part of
4th segment and also about base and centre of 5th ; genitalia
small, brown, two short telescopic cylindrical joints in c? , two
short filaments in $ ; venter deep blue, punctate, with a little
greyish pubescence. Legs : femora black, with a slight blue or
violet tinge and a little whitish-grey hair below ; knees narrowly
dark brown ; tibias black or dark brown, tips very narrowlv a
little paler; tarsi blackish brown, middle pair distinctly paler.
Wings dark brown, stigma and veins darker ; costal cell, discal cell
mainly or wholly, base of 4th posterior cell (sometimes base of 3rd
or even 2nd also), anal and axillary cells (sometimes hinder part
of 2nd basal cell also), clear or nearly so ; halteres brownish yellow.
Length, 8|-9 mm.
Described from six c? d1 and one 2 i11 perfect condition
from Tura, Garo Hills, Assam, 1500 ft., vii. 1917, and above
Tura, 3500 ft., viii. 1917 (Kemp}; a ^ from Sadiya, Assam; a
$ from Karkur Ghat, Nilgiri Hills, 1500 ft., v. 1911 (H. L.
Andrcwes) ; and a second c? from TCohima, Assam, all in the Indian
Museum ; and a specimen from Coorg, S. India (T. B. Fletcher).
This species was given in my first paper on STRATIOMYIDJE as
"auricollis, Bigot," which afterwards proved to be merely a MS.
name.
The gold pubescence on the front of the thorax and that
forming an approximate cross on the thorax in the $ rubs off
very easily.
28 STRATIOMYIDJE.
8. Acanthina argentihirta, Bran. (PL I, fig. 4.)
Acanthina aryenteu, Brimetti, Ifec. Ind. Mus. i, p. 132 (1907) ;
emended in Corrigendum slip of same date to argentihirta
(aryentea preocc. Ost.-Sack. 1880).
3 . Head with eyes extending whole height of head, shortly but
not thickly pubescent, subcontiguous at nearest point of approach,
as the froiis at this point is receding, but attains the surface of
the eyes towards the vertex, which is considerably raised, occupied
by the ocelli ; facets rather large, of uniform si/e ; frons both
above and below nearest approach of eyes, shining white; occiput,
underside of head, and inner orbit of eyes below antennae white.
Antenna? with first two joints black, 3rd reddish brown with
blackish marks ; style thick. Proboscis shorr, yellowish, with a
few hairs. Thorax with dorsum, sides, and .scutellum black,
uniformly covered with short silvery-grey pubescence ; scutellum
with four rather large whitish spines. Abdomen black, with short
silvery-grey pubescence ; venter Mack, with short grey hairs.
Le64, pi. xiv,
figs. 2, 4, Itt (1913).
d $ . Head twice as broad as long ; eyes in <$ contiguous for
greater part of distance from vertex to antennae ; f rons shining
black in J , one-fourth to one-fifth width of head in $ , less
shining, bare ; ocelli on distinct prominence, yellowish ; occiput
black, minutely pubescent. Antennae black, 1st and 2nd joints
subequal, cylindrical, barely broader at the narrowly brown tips,
minutely spinose ; 3rd joint cylindrical, almost filiform, pointed,
with eight annulations. Thorax dull black, with microscopic
pubescence, Avhich in perfect examples is probably blackish or dark
* 2nd and 3rd longitudinal veins respectively,
t 4th longitudinal rein-endings.
AMPSA.LIS. — EPJU1PPIUM. 45
grey ; traces of a little brownish colour behind the wings ; hind
corners of dorsum distinctly so coloured. Scutelhnn of normal
size, shining aeneous or blue-black, with soft (? pale) hairs aud two
very long, powerful, conspicuous, reddish-brown apical spines half
as long as the abdomen, diverging and directed slightly upwards.
Abdomen brownish yellow ; 1st and 2nd segments mainly black,
hind border of latter brownish yellow, 3rd and 4th each with a
rounded or subquadrate blackish spot of considerable size towards
each side but clear of the margins ; these four spots in the $
nearly filling the surface of these segments ; 5th and 6th segments
in both sexes mainly black, but sides and hind margins brownish
yellow. Venter with 1st, 2nd, 4th, and 5th segments mainly black,
remainder brownish yellow ; the black colour a little more ex-
tended in <5 ; whole surface of venter in both sexes with very
short yellowish pubescence. Genitalia in d concealed ; in $ a
narrow cylindrical tube, with a pair of rather long brownish-
yellow hairy filamentous appendages. Legs yellow; hind tibiae
iucrassated on apical two-thirds, black ; hind tarsi longer than
tibia?, the hind metatarsus black for two-thirds of its length and
as long as the remaining joints taken together. Wings distinctly
yellowish, very shining, apical third and hind margin grey ; stigma
black or blackish brown, veins blackish ; halteres yellow.
Length, 10-13 mm.
Described from three <$ J and four $ $ from Darjiling,
1000-3000 ft., v-vi. 1912, and Siiigla, iv. 1913, in fair condition ;
Shillong, 4900 ft., 31. v.l 918(7fao); 5000 ft.,vi-vii. 1918 (Fletcher).
There is nothing material to prevent this species from coming
in Ampsalis. The head is barely as wide as the thorax at its
widest part, and Walker says the scutellum is armed with two
oblique ascending spines, but does not mention their great length.
The only apparent discrepancy is that he says the flagellum is
about twice the length of the scape, whereas in the present species
it is fully three times as long, but this difference would not be
generic.
Genus EPHIPPIUM, Lair.
Epldppium, Latreille, Gen. Crust. Ins. iii, p. 448 (1802-3), and xiv,
p. 341 (1804).
EpAippiomyia, Bezzi, Zeits. Hym. Dipt, ii, p. 191 (1902); Brunetti,
Rec. Ind. Mus. i, p. 115 (1907).
GENOTYPE, Stratiomys ephippium, L. (Europe) ; by original
designation.*
Head comparatively small, as wide as front part of thorax ; face
arched, not projecting in profile, densely pubescent ; palpi long
and thickly pubescent ; eyes densely hairy in both sexes, in 3
* [Latreille did not actually designate n genotype ; in vol. iii he mentions
two species as belonging to his genus, viz., microleon, L., and cpJiiyypium, L. ;
in vol. xiv lie gives the same two species in reverse order. — ED.]
46 STRATIOMYIDjE.
contiguous for a considerable distance, facets of equal size ; in
$ , i'rons broad. Antennae as long as broad, 1st and 2nd joints
subequal, 3rd elongate ; indistinctly annulated, with long pointed
bare apical style. Thorax a little longer than broad, narrower in
front, with a conspicuous spine on each side, just before the base ;
humeri elevated; scutellum higher at base than hind part of
thorax ; with two long thick hairy spines. Abdomen ovate,
flattened, a little longer and broader than thorax and scutellum
together. Legs strong, moderately long ; front coxse large.
Wings strongly rippled, but not ribbed ; traces of posterior cross-
vein, as upper branch of 5th vein is in almost punctiform contact
with discal cell ; discal cell-veinlets distinct ; alulse unusually large ;
alar squamse absent, thoracic pair small.
Range. Europe, Japan, South Asia.
Life-history. That of the European E. thoracicum, Panz., is
known ; a conspicuous but hardly a common species. The larva?
are parasitic in nests of the ant, Formica fulif/inosa. Koser
records finding one in a rotten nut-tree, and thinks they must take
about four years to reach maturity.
The alleged preoccupation of Ephippium by Bolten in Mollusca,
on which Bezzi suggested Ephippiomyia, is very doubtful, accord-
ing to Verrall, and the original name is of too old standing to be
altered.
21. Ephippium bilineatum, F. (PI. I, figs. 15, 16.)
Stratiomys bilineata, Fabricius, Syst. Antl. p. 79(1805).
Clitellaria bivittata, Wiedemann, Zool. Mag. i, p. 3 (1819) ; Auss.
Zweifl. ii, p. 46 (1830).
Ephippium angustum, Macquart, Hist. Nat. Dipt, i, p. 252 (1834).
Rhaphiocera spinithorax, Macquart, Dipt. Exot. Supp. iii, p. 17,
pfi, fig. 7(1848).
Clitellaria tenebrica, Walker, List Dipt. Brit. Mus. iii, p. 522
(1849).
Ephippium spinigerum, Doleschall, Nat. Tijd. Ned. Ind. x, p. 407.
pi. ix, fig. 2(1856).
Negritomyia bilineata, Wulp, Notes Leyd. Mus. vii. p. 95 (1885).
Ephippiomyia bilineatum, Brunetti, Rec. Ind. Mus. i, p. 115 (1907).
c? $. Head: eyes of c? with rather dense brown pubescence,
contiguous for about the same distance as from lowest point of
contact to base of antennae ; vertical triangle rather small, with
blackish-brown hairs ; frons and face black, with dense, white or
whitish, rather long pubescence ; occiput black, margin with
minute snow-white pubescence, which nearly disappears at the
sides and becomes longer on lower part ; proboscis with very large
dark brown labella ; palpi blackish, pubescent. Antenna? with first
two joints black, bristly ; 3rd to extreme tip of style about one and
a half times as long as 1st and 2nd together, dull brown, elongate,
conical, with three distinct broad annulations, bare, after which
it narrows abruptly and bears some distinct hairs and is produced
into a very narrow terminal style. In $ , frons about one-twelfth
EPHIPP1UM.— NEGEITOMYIA. 47
width of head, sides parallel, shining black, with a small patch of
white pubescence in middle. Thorax shining black or blue-black,
closely punctulate ; dorsnm with blackish-brown pubescence and
two moderately wide well-separated stripes from anterior margin
to hind margin, or even indistinctly carried over part of the
scutellum, composed of very short whitish-grey pubescence ;
hutneri rather prominent, shining bright brown, bare ; sides of
thorax shining black, mainly covered with longer or shorter whitish-
grey pubescence. Scutellum concolorous with dorsum, hind mar-
gin with a little grey pubescence ; the spine at each hind corner
strong, long, black, and pubescent. Abdomen shining black or
blue-black, closely punctulate, with dark brown pubescence on
dorsum and longer whitish-grey pubescence at sides and on the
concolorous venter ; a tendency to form a hair-spot towards sides
of each segment, especially on 3rd and 4th ; underside of last
segment bare, shining. Legs black, shortly grey pubescent ; tips
of tarsi brown below. Wings with base narrowly, costal cell,
base of 1st basal cell and the axillary cell, clear ; rest of wing
dark brown, a little paler on hind margin; 5th vein rather more
deeply infuscated ; halteres pale yellow.
Length, 9-14 mm.
Described from several specimens in the Indian Museum :
Tenasserim (Dolierty} ; Semarang, Java, viii. 1909, and Depok,
Java, x. 1907 (Jacobson).
Genus NEGRITOMYIA, Big.
Niyritomyia, Bigot, Bull. Soc. Ent. France, (5) rii, p. Ixxiv (1877).
Negritomyia, Bigot, Ann. Soc. Ent. France, (5) ix, p. 190 (1879).
GENOTYPE, Ephipplum maculipenne, Macq.; by original de-
signation.
Antennae nearly cylindrical, base a little pubescent, 1st and 2nd
segments subequal, 1st much shorter than 3rd, arista apical, about
as long as the whole antenna, densely, shortly, and uniformly
pubescent. Eyes contiguous in tf , distinctly apai-t in $ , densely
pubescent in both sexes. Thorax with a long spine on each side ;
scutellum with two long spines. Wings with four endings to the
4th longitudinal vein, all reaching the border.
Range. Africa, Ceylon, Malay Archipelago, Australia.
22. Negritomyia macnlipennis, Macq.
Ephippium maculipenne, Macquart, Dipt. Exot. Supp. iv, p. 54
(1849), rf.
Negritomyia maculipennis, Brunetti, Rec. Ind. Mus. i, p. 114 (1907),
& vii, p. 451 (1912).
$ . Head black, with a little short whitish pubescence ; eyes
black, shortly pubescent ; frons at vertex about one-fifth the width
of the head, widening at level of antennae to nearly one-third ; an
indistinct whitish mark above each antenna ; palpi black, with
48 STRATIOMYIDjE.
short yellow pubescence ; back of head black ; posterior orbit of
eyes with narrow silvery-white pubescence. Antennae blackish,
inner side of 3rd joint pale; style closely pubescent. Thorax
black, moderately shining, with very short black hairs ; covered
with microscopic grey hairs, except along a broad median line and
on two large spaces towards each side of the dorsnm, i. e., one
behind the shoulder and above each wing ; sides of thorax shining
black, with microscopic grey pubescence below shoulders and some
short snow-white hairs above the fore coxae ; there is also a rather
wide perpendicular side-stripe of similar pubescence above the
middle coxae, joining the dorsal pubescence in front of the wing ;
side-spines short, strong, black. Scutellum black, with short black
hairs and concolorous grey pubescence, and two broadly separated,
strong, short, black spines on hind margin. Abdomen black, with
grey pubescence forming a broad dorsal line on hinder half and a
subtriangular spot towards each side of each segment. Venter
shining black, practically bare. Legs black, with very minute
whitish pubescence ; basal fifth of anterior femora and basal fourth
of hind pair, also major portion (from the base) of .'ill the tarsi,
pale yellowish white. Wings subhyaline ; stigma dark brown,
roughly oval, very distinct ; tip of wing from outer side of discal
cell, thence in a straight line to the costa and extending hind-
wards, until filling the 2nd posterior cell, distinctly brown, the
colour filtering more or less into the 3rd posterior cell ; halteres
with stem yellow, clubs dark brown.
Length, 11 mm.
Kedescribed from a single perfect $ in my collection from
Kandy, Ceylou, viii.1908 (Green) ; Pollibetta, Coorg, x-xi. 1915
(Fletcher). Loew's maculipennis (1856) from South Africa is a
different species.
Genus RUBA, Walk.
Ruba, Walker, Proc. Linn. Soc. Lond. iv, p. 100 (1860).
GENOTYPE, Ruba injlata, Walk. ; by original designation.
Body thick and broad ; head much narrower than thorax ;
proboscis and palpi very short. Antennae nearly as long as
breadth of head ; " 3rd joint broader and longer than the flagellum,
of which the joints are short, compact, and minutely setulose ".*
Thorax a little longer than broad, scutellum unarmed ; abdomen
globose, very much broader and a little longer than thorax. Legs
short. ' Wings moderately broad ; " 1st cubital vein not one-third
of length of 2nd ; four complete externo-medial veins ; subanal
vein curved, joining the anal vein at some distance from the
border ; discal areolet elongated exteriorly, irregularly triangular,
exterior side very short."
Range, India, Celebes, Papua.
* Walker's '' flagellum " is the style.
EUBA. — CLITELLAHIA. 49
23. Ruba inflata, Walk.
Huba inflata, Walker, Proc. Linn. Soc. Lond. iv, p. 101 (1860).
2 . Head : frons and face one-third of head, uniformly wide
except tor a slight contraction slightly above base of antennae ;
oc^lhir triangle very small, black; an oval brown longitudinal
mark on fro us just above the orange antennae, of irregular shape,
continued as a tine line to the ocellar triangle ; whole head
brownish orange; proboscis with very large labella; tips of palpi
rounded, flattened, rather large, deep black;* whole head with
rather short, moderately dense, but not conspicuous, yellow pubes-
cence. Thorax and scutellum all brownish orange, \\ ith rather dense
yellow pubescence; traces of three longitudinal narrow blackish
lines on dorsum. Abdomen brownish orange, with yellow pubes-
cence. Ley* yellowish orange, with yellow pubescence ; apical
hiilt' of hind tibiae black, with black pubescence ; the tiny bristles
at the tips of the tarsi give them a blackish appearance Wings
pale yelloxv, apical half (except extreme tip) rather bright brown,
darker anteriorly ; a quadrate dark brown spot of modera'e si/e
at tip of 1st basal cell, extending to the costa ; anterior margin
of basal half of wing a little deeper yellow.
Length, 9 mm.
Described from a perfect female in the Indian Museum from
Ghumti, Darjiling District, 1000 ft., vii.1911 (Gravely) ; and one
in inferior condition from Kohima, As*am.
Walker described the species from Makassar, in Celebes, and
the only descrepancy is that he says nothing about the black
apical half of the hind tibiae.
Clitellaria, Meigen, Illig. Mag. ii, p. 265 (1803).
Adoxomyia, Kert6sz, Ann. Mus. H
Genus CLITELLARIA., Mg.
. 265 (1803).
ung, v, p. 499 (1907) ; nom. nov.
for Clitellaria, Sch., nee Mg.
GENOTYPE, Clitellaria dahlii, Mg. (Europe) ; designated by
Bezx.i, Wien. ent. Zeit. xvii, p. 75 (1908). f
Head: eyes densely pubescent, contiguous in male, wide apart
in female; proboscis retracted, with moderately broad labella;
palpi 2- jointed, 1st joint thick, kidney-shaped, ^nd narrow, conical;
epistoma slightly arched. Antennae elongate, 1st and 2nd joints
subequal ; 3rd elongate conical, with four annulations, with two-
* The palpi lie along the lower side of the face and the conspicuous black
tips look like two black spots on it, Walker evidently mistaking them Cor such
as he mentions " a black sj ot on each side of the mouth."
t [It seems necessary to point ou' that this designation is entirely at variance
with the generally accepted rules of zoological >;omenclature. Meigen, when
describing Clitellaria. definitely designated ephipptum, F., as the genotype,
and no subsequent action on the part of Meigen himself or anyone elt-e can
alter that fact. The correct designation for the present genus is therefore
Adoxomyia, Kert., and the species described above (p. 46) as Ephippium biliiie-
atum should stand as Ciitellaria biliiieata, F. — ED.]
E
50 STEA.TIOMYIDJE.
jointed apical style, 1st very short, 2nd long. Thorax elliptical,
moderately arched ; scutellum with two spines, often small and
indistinct. Abdomen a little broader than thorax, oval or elliptical,
rather well arched, 5-jointed ; genitalia small. Legs moderately
long and strong. "Wings with normal venation ; 3rd vein distinctly
forked, 4th vein with three endings; posterior cross-vein abseni,
as upper branch of 5th vein forms part of lower side of discal cell.
Range. Europe, Western Asia, North Africa, Orient, North
America.
Life-history. That of the common Indian 0. heminopla, ^\ied.,
has been worked out at the Pusa Institute.
Some confusion has arisen over the name Clitellaria. Verrall
notes that it was established by Meigen in 1803 for ephippium, F.,
and that that author subsequently (1822) enlarged the scope ot
the genus, including in it villosa, F., calva, Mg., and pacifica, Hfg.,
as well as epliippium,T?. Latreille in 1804 erected Epliippium for
the species of that name,* and, as was customary iu those days
in such cases, changed the specific name to thoracicum to avoid
redundancy. Therefore the name Clitellaria must stand for the
remainder of the species placed in it by Meigen or, at the very
least, for one of them. C. pacifica is an Odontomyia and synony-
mous with limbata, Wied., which leaves only villosa and calva, one
of which should be the type of Clitellaria, Mg. Kertesz retains
Meigen's original sense of th*e genus (i. e., for ep7iippium, F.) in
his Kat. Dipt. (1908), but adopts the inadmissible name Potamidu
(Mg. 1800) for it. Meigen added dahlii in 1830 to Clitellaria and
Kertesz has renamed this group of species Adoxomyia. Lasiopa,
Brulle (1832), differs from Clitellaria, Mg., in having an unspined
scutellum, and both villosa and calva have been placed in it.
Therefore Clitellaria, as understood by Meigen, was by this time
reduced to the single species dahlii, and this species is designated
by Bezzi as the genotype. Kertesz's " Potamida, Mg." must con-
tinue to be known as Ephippium, Latr. ; Adoxomyia, Kert., is the
true Clitellaria, Mg., and must retain this latter name; and Lasiopa,
Brulle, is a good genus.
The two Indian species are separated thus : — -
Thorax without definite hair-stripes, but with a
darker median stripe surrounded by four spots
arranged in a square heminopla, Wied.
Thorax with two distinct stripes of short golden-
brown hairs bistriata, Brun.
24. Clitellaria heminopla, Wied. (PI. I, fig. 14.)
Clitellaria heminopla, Wiedemann, Zool. Mag. iii, p. 30 (1819) ; id.,
Ausfl. Zweifl. ii, p. 48 (1830) ; Brunetti, Kec. Ind. Mus. i, p. 120
(1907).
c? £ . Head : eyes in c? practically contiguous for about half
the distance from vertex to frontal triangle; black, upper facets
* [This is not strictly accurate (see footnote on p. 45). — ED.]
CLITELLAEIA. 51
barely larger than lower ones, not clearly demarcated, with
moderately short dense pubescence, which appears grey or brown
according to the point of view; vertex black, with a few long
whitish hairs ; ocelli reddish ; frontal triangle black, with rather
long soft whitish hair ; face and lower part of head black, with
greyish-white pubescence ; proboscis dull yellowish ; palpi reddish
brown; autennal 1st and 2nd joints black, 3rd joint reddish
brown to yellowish, first four anuulations much broader and more
distinct than the rest, which form a narrow blackish cylindrical
tip; a reddish-brown ring round base of antennal prominence.
In $,frons forming | to i width of head, a little narrower at
vertex, with short whitish-grey or brownish-grey pubescence ;
lower part of frons shining, practically bare, except for a bunch
of drooping whitish hairs near each eye-margin above level of
antenna ; the four basal annulatious of 3rd joint rather broader
than in <5 • Thorax black, dorsura wholly covered with whitish-
grey tomentose pubescence ; in perfect specimens, if viewed from
in front, a median stripe surrounded by four spots arranged in a
square can be seen, composed of gold-brown tomentose pubescence ;
•some longer whitish-grey hairs below shoulders. Scutellum large,
with grey and golden-brown admixture of tomentose pubescence ;
spine placed at each hind corner, yellowish brown ; a fringe of
white hairs depends from the hind margin. Abdomen black ; 2nd
segment wholly, 3rd narrowly on hind margin and more broadly
towards sides and along side-margin of segment, 4th broadly
on hind margin (especially towards and along the side-margins),
5th with a large median triangular spot (its base on hind margin
of segment), very shortly but distinctly whitish (occasionally rather
golden) pubescent ; ground-colour of extreme side-margins of
segments brownish orange ; venter blackish, with extremely short
whitish pubescence. Legs : coxae black, brownish orange at tips ;
hind pair brownish orange behind ; femora and tibiae rather shining
black, the former brownish orange at base and narrowly at tips ;
tibiae broadly brownish yellow at base, narrowly brown at tips ;
t'r.nit tarsi blackish, posterior metatarsi yellowish, remaining
joints brown ; all legs with short greyish pubescence, which is
more brownish yellow on tarsi, especially the front pair. Wings
grey, yellowish brown about the centre ; stigma slightly darker
brown ; halteres yellow.
Length, 6^-9 mm.
Described' from a long series of both sexes in the Indian
Museum, Pusa collection and my own, from many parts of India
and Ceylon, where it appears to be generally distributed and
common from March to October on rotten papaya-stems, plan-
tains, and melons.
Mr. Hewlett has taken it " at sex play around dead leaves,
Pusa, 15. iii. 1912."
Larva. This is 11 mm. in length, whitish, oblong, the 4t,h seg-
ment truncate, the three anterior segments much narrower and
apparently withdrawable into the 4th ; 1st segment rather elon-
gate, verv narrow; 4th to 12th segments about equal in length,
E2
52 STJIATIOMYIDJE.
anal segment longer, rounded at tip ; each segment bearing a
transverse row of rather long curved yellow bristles on both upper
and under sides, one or more being also present along the sides.
The surface is covered with minute white particles, apparently
purposely attached to its body. Found in rotten papaya-stems.
25. Clitellaria bistriata, Brun.
Clitellaria bistriata, Brunetti, Eec. Ind. Mus. vii, p. 452 (1912).
3
or practically contiguous in tf , broadly separated in $ . Proboscis
short or lengthened, porrect, with moderately large labella; palpi
with three distinct joints. Thorax elliptical ; scutellum unarmed.
Abdomen broader than thorax, elliptical, arched, five- segmented.
Leys moderately long and stout. Wings with 3rd vein distinctly
forked ; 4th longitudinal vein with four faint veinlets, of which
the 4th originates near or directly from the 2nd basal cell.
Range. Europe, West Asia, North Africa, South Africa, Orient,
New Zealand, South America.
Life-history unknown. The perfect insects occur 011 umbelli-
ferous and other flowers, and often become rather sluggish in their
habits.
26. Lasiopa villosa, /"., var. himalayensis, Brun.
Xe.motelus villosa, Fabricius, Ent. Syst. iv, p. 270 (1794).
Lasiopa villosa, var. himalayensis, Brunetti, Ilec. Ind. Mus. i, p. 117
(1907).
J 5 . Head black, with rather long, shaggy, yellowish-grey
hair ; eyes in J not quite contiguous ; frons in $ one-third width
of head, sides parallel, with rather Jong golden-yellow pubescence ;
face similar, with a rather small elongate spot on each side in $
at level of antennae, contiguous to inner margin of eye ; proboscis
dark brown, not prominent ; underside of head with greyish
pubescence ; antennae blackish brown. Thorax black, with yel-
lowish-grey pubescence, which is shorter in the $ . Abdomen
black: an elongate yellowish spot towards each side* on hind
border of 2nd, 3rd, and 4th segments, all the spots subequal ; 5th
segment with distinct yellow hind margin. Legs blackish brown.
Wings brownish grey ; halteres whitish yellow.
Length, 9-10 mm."
Described from three specimens in my own collection from
Mussoorie, v. 1905 (Brunetti), and others from Kumaon, W.
Himalayas, 6300 ft., ll.v. 1911 (Kemp}-, Nairn Tal, vi. 1893
[Lucknow Museum]; Dungagali, Hazara Dist., 8000 ft,, 21-24.
v. }Q15 (FletcJter).
The only difference from typical European specimens is that
the abdominal yellow spots are not turned up and widened at
their inner ends.
Genus OXYCERA, %.
Oxycera, Meigen, Illig. Mag. ii, p. 265 (1803).
Hypoleon, Dumeril, Expos. Meth. Natur. (1801).
lletcroxycera, -Bigot, Ann. Soc. Ent. France, (3) iv, pp. 6-'> & 85
(1850).
GKKOTYPE, Jfusca hypoleon, L. (Europe) ; by designation of
Curtis, 1833 (as trilineata, F.).
Head with epistoma smooth, a little arched ; eyes from bare in
both sexes to densely pubescent in $ (in which case $ has some
pubescence too), contiguous in J for a considerable distance, the
54 STEATIOMTID^E.
upper facets much larger than the small lower ones and generally
sharply delimited from them ; in $ wide apart, with uniform
facets ; occiput- with lower half considerably puffed out. Pro-
boscis retracted, with broad labella ; palpi rather inconspicuous.
Antennae elongate, first two joints subequal, 3rd spindle-shaped
or peg-shaped, with four annulations and a two-jointed apical or
subapical bristle. Thorax oblong, rather arched, moderately
pubescent; humeri and posterior calli prominent; scutellum with
two spines. Abdomen short and round, very arched, 5-segmented.
Legs simple, short, weak. Wings with normal venation ; 3rd
longitudinal vein forked or simple ; 4th vein with three endings,
the 3rd often weak or absent ; posterior cross-vein absent, upper
branch of 5th vein forming part of lower side of discal cell, the
latter often indistinct : alulae well developed.
Range. Europe, Asia, North America, and one species from
Caffraria.
Life-history. The larva lives in moist mud among Conferva.
The flies frequent leaves and flowers in the sunshine near streams
and damp situations, where they are very active ; on sunless days
they settle on the underside of leaves near the ground. The
metamorphoses of several European species are well known.
These insects are rather small, nearly bare, flies, generally of a
black colour with conspicuous yellow spots.
0. siynata is mainly a black species and 0. albomicons a yellow
one.
27. Oxycera signata, sp. nov.
d . Head : eyes with dense brown pubescence, contiguous for
a considerable distance, leaving vertical and frontal triangles
about equal in size, black ; former with black hairs and three
brownish ocelli on an elevated prominence ; latter bare, shining ;
face black, with a long oval patch of short snow-white pubescence
adjacent to the eye-margin on each side ; no postocular orbit ;
occiput black ; proboscis orange-yellow ; underside of head with
short white hairs. Antennae black, first two joints subequal, 3rd
as long as first two together, with four annulations and subapical
arista. Thorax shining black, with moderately dense brownish-
grey pubescence; a pair of widely separated, pale yellowish,
narrow, longitudinal dorsal stripes, nor quite reaching either
anterior or hind margin of dorsum ; an elongate bright lemon-
yellow spot on each side of dorsum from prothorax up to the
suture, and a large oraiige-yellow one on each posterior angle of
dorsum ; pleurae bkck, with whitish pubescence and a callus-like
long yellow stripe from prothorax (where its end is contiguous to
the stripe on the side of the dorsum immediately above it) to root
of wing. Scutellum bright lemon-yellow, with brown pubescence,
hind margin black ; metanotum shining black, microscopically
pubescent. Abdomen black, moderately shining, with micro-
scopic brown pubescence and a little white hair at sides of 1st
segment, which latter is narrowly yellow at base ; a large subtri-
OXTCEEA. 55
angular spot on 3rd segment on each side-margin reaching from
anterior nearly to posterior margin; a smaller oblong one placed
endways at middle of side-margin of 4th segment ; and a small
semicircular apical one on last (5th) segment ; venter black, with
a little sparse pale pubescence. Legs black, with the following
parts yellow : tips of femora broadly and posterior pairs indis-
tinctly at base, front tibiae broadly at base, middle pair wholly
and hind pair narrowly at base and tip, also posterior metatarsi
and 2nd joint of^hind tarsi ; all legs microscopically pubescent.
Wings colourless ; anterior veins and stigma pale yellowish ;
halteres bright yellow.
Length, 5 mm.
Described from a single tf in the Pusa collection from India,
probably from Mussoorie.
Type presented to the British Museum.
28. Oxycera albomicans, sp. riov.
$ . Head wholly bright sulphur-yellow. Erons and face fully
one-third of head, very slightly narrower at about middle of frons ;
frons rather broader on lower part; eyes bare; upper eye-margins
of frons narrowly black ; a thin line running from the corner of
each eye diagonally to vertex at each side of vertical spot ; an
elongate black spot on vertex, embracing the ocelli, pointed at
anterior end ; an oval or leaf-shaped one of moderate size in
middle of frons and a small elongate, vertically placed one imme-
diately above base of antennae ; tlie latter wholly brownish orange,
of normal size ; arista black ; lower sides of face (viewed from
above) with snow-white shimmer; mouth-region and proboscis
brownish orange ; postocular orbit very broad, lower part with
whitish shimmering tomentum and microscopic pubescence ; occi-
put black. Thorax sulphur-yellow, microscopically punctate, with
very short yellow pubescence; dorsum with three longitudinal black
stripes, the median one slightly the widest and reaching front
margin, the side-stripes just failing to reach it — all three narrow
and united on hind margin ; a small black vitta just above wing-
base. Lower part of thorax, front part of sternopleura and a
contiguous triangular spot on mesopleura, black ; pleurae and
lower part of thorax with a little microscopic white pubescence.
Scutellum yellow, with a little microscopic black pubescence ;
spines rather long, tips a little darker. Abdomen sulphur-yellow,
microscopically punctate, ernarginations of segments very indis-
tinct; two broad black transverse bands, broader still in the
middle of each, lying apparently over the junctions of 2nd and
3rd segments and of 3rd and 4th segments respectively, united
by a short black median stripe ; the anterior transverse band
broadly united to the convex side of a semicircular band, which
has its ends on the anterior corners of the 1st segment, but just
failing to attain the actual margin : hinder transverse band united
by a median black longitudinal stripe to an arched narrower black
56 STBA.TIOMYJ.DjE.
transverse band placed on base of 5th segment, the middle part
encroaching over hind margin of 4th segment. Venter yellowish,
with microscopic pale \ ellow pubescence. Lec/s vellow or brownish
yellow j front tarsi wholly, apical half of middle pair, hind tibiae
(except at tips), and last four joints of hind tarsi dark brown.
Wings colourless; veins yellowish; 3rd vein forked towards tip;
halteres bright yellow.
Length, 4| mm.
Described from two $ $ from Abbottabad, N.W. Frontier
Province, vi. 3916 (Fletcher).
Type sent to the British Museum ; co-type in Pusa collection.
The abdomen might equally well be described as black with the
base narrowly and tip more broadly yellow, and three pairs of
yellow side-spots, the first pair triangular, the 2nd and 3rd pairs
much larger and irregularly oblong.
Subfamily STRATIOMYI1SLE.
Head short, rather flattened in front ; antennae with 3rd joint
elongate, no arista. Thorax subquadrate, often only slightly
arched; scutellum generally with two spines. Abdomen 5 to b'-
segmented, generally subquadrate, not much longer than thorax,
often much flattened and frequently with obvious pale markings.
Legs of moderate length and strength, or somewhat weak. Wings
with 3rd vein generally forked, ending far before wing-tip; 4ih
vein with three terminations or veinlets, seldom reaching v\ ing-
margin, one or more often absent or extremely weak ; posterior
Fig. 2. — Stratiomyia, antenna. Fig. 3. — (
cross-vein present, so that the 5th vein never forms any part of
the discal ceil ; anal cell closed near wing-border ; 5 posterior
cells, united on wing-margin. Wing in repose folded closely
over middle of al'domen, leaving side-margin* exposed; alar
squamae small, thoracic pair large, pubescent.
The principal cha'-acters of this subfamily taken in conjunction
are the subquadrate abdomen of the typical genera (Stratiomyia,
Odontomyia), the elongate antennae devoid of arista, and the
presence of the posterior cross-vein. Larva peripneustic, last
segment more or less long and narrow.
Table of Genera.
1. Third antennal joint with at most six
annulations 2.
Third antennal joint with seven or
eight annulatioiis CYPHOMYIA, Wied., p. 57.
CYFHOMYIA. 57
2. First antennal joint three or more
times longer than 2nd STBATIOMYIA, Geoff., p. 58.
First antennal joint less than three
times as long as 2nd, generally
much less ODONTOMYIA, Mg., p. 61.
The above distinctions are purely arbitrary, but each genus has a
facies peculiarly its own, which the student will soon recognise.
Genus CYPHOMYIA, Wied.
Cypliomyia, Wiedemann, Zool. Mag. i, p. 3.(1819).
GENOTYPE, C. auriftamma, "Wied. (Tropical America); designated
by Brauer, 1882.
Head as broad as thorax, semicircular ; eyes bare (occasionally
just perceptibly hairy), in J closely contiguous for a considerable
distance ; frons in § rather broad, sides approximately parallel ;
postocular orbits in $ broad. Antennae with 1st joint moderately
long, broader at tip; ind cup-shaped, half as long as 1st; 3rd
cylindrical, tapering, of eight subequal annulations, practically
bare. Thorax considerably elongate, much narrowed in front,
moderately arched, anterior corners rather angular; in many
species with three lighter longitudinal stripes. Scutellum rather
large, broader than long, trape/oidal, with two spines that are
longer and more divergent in d than $ . Abdomen subglobular,
thick, about as long as thorax, 5-segmented ; 1st segment very
short, mostly hidden by scutellum. A generic peculiarity is the
presence of hair-spots towards the sides of the last segment,
placed on its anterior margin. Legs rather long, especially the
femora, simple, slender ; tarsi generally longer than tibiae. Wings
fully developed, considerably longer than abdomen and folded over
it when ab rest.
Range. North, Central and South America, South Africa, India,
Sumatra.
29. Cyphomyia indica, sp. nov.
Head wholly bright orange, bare, including the thick postocular
orbits, which are distinctly demarcated from the vertex by a
narrow groove ; frons about one-third width of head, barel)r
broader at vertex ; underside of head with a little short white
pubescence towards the eye-margins ; eves and proboscis black.
Antennae with 1st and 2nd joints dark reddish brown, 3rd black,
•with eight distinct annulations, three times as long as 1st and 2nd
together. Thorax deep black, with rather dense short grey and
brownish-grey pubescence, which is a little longer and more
whitish posteriorly and on the concolorous, rather large, nearly
oblong scutellum ; spines of latter tipped with brown ; pleurae
blackish with a little whitish pubescence. Abdomen much broader
than thorax, nearly circular, brilliantly shining violet-blue, with
very short whitish pubescence; venter similar. Legs black, with
whitish pubescence ; knees barely perceptibly brownish orange ;
58 STRAXIOMYID^E.
more than basal half of front metatarsus, and the hind metatarsus
wholly, and most of the succeeding joints brownish orange;
pubescence of legs black. Winys uniformly rather dark brown, a
little lighter at base and darker about the stigma ; halteres pale
dull orange.
Length, 9 min.
Described from a unique specimen (genital organs missing) in
the Indian Museum from Kalhnpong, 600-4500 ft., Darjiliug
District, iv-v. 1915 (Gravely}.
Genus STRATIOMYIA, Geo/.
Stratiomys, Geofl'roy, Hist. d. Ins. ii, p. 475 (1704). *
Stratiomys, Fabricius, Ent. Syst. iv, p. 262 (1704); Meigen, Illig.
Mag. ii, p. 265 (1803).
Stratiomyia, Macquart, Dipt. Exot. i, pt. 1, p. 179 (1838) ; Brunetti,
Rec. Ind. Mus. i, p. 125 (1907).
Hoplomyia, Zeller, Isis, xi, p. 882 (recte 828) (1842).
Stratiotomi/ia, E. L. Arribalzaga, Bol. Acad. Nat. Cienc. Cordoba,
iv, p. 128(1882).
GENOTYPE, Musca cJiamceleon, L. (Europe) ; by designation of
Latreille, Consid. Gen. (1810).
Head semicircular, face arched, postocular orbits thickened,
especially in $ ; proboscis comparatively small ; palpi small ;
eyes in d practically contiguous, the upper facets often much
larger than lower ones, wide apart in $ ; in both sexes either bare
or hairy, sometimes a sexual character. Antenna approximate at
base, diverging after 1st joint, which is very much longer
than 2nd ; 3rd joint elongate, cylindrical, bare, with live or six
anuulations (occasionally an annotation subdivided). Thorax sub-
quadrate, broader behind, pubescent ; scutellum two-spined.
Abdomen broader than thorax, slightly arched, subquadrate,
shoulders angular; whole doratim sparsely, moderately, or some-
what densely pubescent ; wings folded one above the other at
rest, lying within the side-margins of the abdomen ; genitalia
small. Leys moderately stout, shortly pubescent. Wings mainly
as in Odontomyia; 3rd vein forked, 4th vein terminations (three)
long, curved, nearly reaching wing-margin, the hindmost more or
less parallel with upper branch of 5th vein ; posterior cross- vein
present but sometimes small; anal cell closed near wing-margin;
alar squamae small, thoracic large and pubescent.
llanje. World-wide except Au.-traliaand the Ethiopian Kegion.
Life-history. Metamorphoses of more than one European species
known. Larvae fusil orm, broadest before the middle; known as
* [In this work Geoffroy did not accept the binnry system of nomenclature
upon which all our modern zoological classification is based ; it has therefore
been ruled that all generic names therein proposed by him are just as invalid a»
pre-Linnyean names. Tlie authorship of Stratiomys should thus be attributed
to Fabricius. — Ei>.]
STRAT1OMYIA. 59
" rat-tailed maggots ;" some are aquatic, hanging head downwards
by their tails, probably living on minute organisms ; some live in
mud or moist sand. The pupa floats free. The imagines frequent
Umbelliferae in moist situations.
Table of Species.
1. Hind margin of 3rd and 4th abdominal segments
with uninterrupted yellow band fulvescens, sp. n.
Hind margin oford aud 4th abdominal segments
with a pair of elongate spots, well separated . . 2.
2. Thoracic pubescence moderately thick and long;
scutellum black, except the narrow yellow hind
margin and spines; $ with two conspicuous
yellow po.stvertical spots, sometimes united . . barca, Walk.
Thoracic pubescence extremely short and sparser ;
scutellum all brownish yellow; $ with no
postvei tical yellow spots micropilosa, sp. n.
30. Stratiomyia barca, Walk.
Strut iomys barca, Walker, List Dipt. Brit. Mus. iii, p. 530 (1849).
Statiomyia barca, Brunetti, Rec. Ind. Mus. i, p. 125 (1907).
J $ . Head : eyes contiguous in J for more than half their
height ; shortly and sparsely pubescent ; upper facets distinctly
but not greatly larger than lower ones ; vertical triangle black,
Avith a few yellow hairs ; ocelli large, brownish ; antenna? black ;
irons, face (punctulate), and lower part of head black, with long,
rather shaggy, bright yellow hair ; occiput black, with very sparse
pale pubescence ; hind margin of eyes with a fringe of microscopic
yellow pubescence. In § , frons aud face nearly half width of
head, sides parallel, black ; behind vertex two conspicuous bright
yellow oval spots, sometimes united as a large semicircular spot ;
two very large elongate yellow calli on frons placed diagonally, the
inner ends nearly contiguous, the outer ends touching eyes and
descending below level of antennae ; colour of calli extended to
sides of face broadly ; a broad median black stripe on face, which
is covered with whitish pubescence ; posterior margin of eyes
with merely an extremely short fringe of whitish pubescence ; a
considerable thickening of the orbit on lower half, where it is
yellow. Thorax black, puuctulate, slightly shining; closely clothed
with dense, rather long, brownish-yellow pubescence, which is
much paler on pleurae ; scutellum similar, hind border and spines
brownish yellow. Abdomen black, slightly shining, with an elon-
gate moderate-sized yellow spot on hind border at sides of 2nd
segment, and elongate spots similarly situated on 3rd and 4th
segments ; 5th segment with a elongate yellow spot, often paler
than the others, placed longitudinally, not attaining anterior
margin but extended at lower end for a short distance each way
on hind margin of segment. Pubescence in c? brownish yellow,
dense and fairly long over all the surface ; in $ black, dense but
60 STIUTIOMYID.*;.
very easily overlooked ; a little pale yellowish-white pubescence
about the shoulders, sides, and tip. Venter in d1 with greater
part of centre of each segment and the hind border yellowish,
rest black ; in £ the yellow part is much paler or even whitish,
and of much less extent ; in some specimens almost confined to
hind borders except on the 2nd segment. Legs mainly black ;
knees more or less broadly hrownish yellow ; hind tibiae pale
brown with a more or less distinct, moderately narrow ring at
middle and tip ; all tarsi brownish yellow, a little darker above :it
lips ; pubescence on legs yellow. Wings pale grey : anterior and
basal parts brownish or yellowish brown, limited by discal cell
and anal cell ; halteres pale yellow, occasionally green.
Length, 10-12 mm.
Described from several of each sex in the Indian Museum ;
Jhelum Valley, 5200 ft., and Srinagar, 0000 ft., Kashmir, vii-ix.
1916 (H. T. Pearse). Described originally from China.
Type in the British Museum.
31. Stratiomyia micropilosa, sp. nov.
$ . Very near barca, Walk., differing as follows : —
The large yellow postvertical spot is entirely absent, as are the
two elongate frontal ones, the latter being replaced by a rather
obscure subqu;idrate median chestnut spot just above antennae;
lower posterior orbit of eyes whitish, not yellow. The dull
yellowish pubescence on dorsum of thorax almost microscopic and
much sparser ; scutellum wholly yellowish, with concolorous
tomentum. Abdominal spots brownish orange; side-spots on
2nd segment more triangular, their upper angles only just touch-
ing front corners of segment; those on 3rd segment similar but
slightly smaller; those on 4th segment smaller still, but partaking
more of the nature of a triangular spot than a linear one as in
barca. Tibiae more narrowly pale at base; hind pair similar to
anterior tibiae.
Length, 12 mm.
Described from a single $ in the Indian Museum from Maymyo,
Burma, v. 1910 (//. L. Andmves).
32. Stratiomyia fulvescens, sp. nov.
<3 . Head with eyes shortly but not conspicuously pubescent,
the very narrow inner orbits contiguous for about half the distance
from vertex to frontal triangle ; vertical triangle black, elevated,
with some brownish-yellow hairs ; ocelli dull yellowish ; frontal
triangle small, black \\ith brownish hairs; face and head below
antennae wholly black, with rather thick and long whitish-yrey
pubescence; occiput black, with grey pubescence, no definite
posterior orbit to eyes ; proboscis brownish yellow. Antennae
black, tip of 2nd joint dark brown ; 1st joint about five times as
long as 2nd; 3rd a little longer than first two together. Thorax
STRATIOMYIA. ODONTOMYI A . Gl
\\holly black, dorsuin entirely covered with dense yellowish-grey
pubescence, and sides with the pubescence a little paler. Scu-
tellum black, with yellowish-grey pubescence ; hind margin
broadly brownish yellow ; spines concolorous, with rather long
\vhitish-grey pubescence about their bases. Abdomen moderately
shining black, with rather dense, brownish-yellow pubescence on
dorsuin of 2nd, 3rd, and 4th segments, the pubescence paler on
anterior corners and whitish on 5th and Gth segments. A rather
large triangular orange spot on hind border towards each side of
2nd segment, the inner end of each narrowed to a fine point, the
base of the spot placed on the side-margin but extending only
halfway to the anterior margin ; hind margin of 3rd segment
orange, the colour very narrow in centre of segment, much
broader towards sides but narrowed again just on side-margin ; a
similar orange hind margin to 4th segment and a fair-sized orange
triangular spot on £th, with its base on hind margin ; Gth segment
all orange; the extreme side-margin of all segments orange,
except the anterior half of 2nd segment. Genitalia brownish
orange, small, apparently consisting of a median cylindrical piece
and a small lamella on each side. Venter with 2nd segment
wholly, 3rd broadly, and 4th narrowly on hind margins, brownish
orange. Legs black, with grey pubescence on femora and tibia?,
yellowish on tarsi ; tips of all femora, basal half of hind pair, base
and tips of anterior tibiae, hind tibiae wholly, and all tarsi
orange. Wings pale brownish grey, veins a little darker ; halteres
orange.
Length 12 mm.
Described from a single c? from Taro, Peshawur District, N.W.
India, 16-19. v. 1915 (Fletcher); presented by him to the British
Museum.
Near 8. barca, Walk., but in that species the abdominal orange
(" tawny ") spots are riot united to form a band ; the tibias are
piceous and the legs have tawny pubescence.
Genus ODONTOMYIA, Mg.
Odontomyia, Meijren, Illig. Mag. ii, p. 265 (1803); Brunetti, Eec.
Ind. Mus. i, p. 126 (1907).
Opseoyymnus, A. Costa, II Giamb. Vico Napoli, ii, p. 443 (1857).
Psellidotus, Rcmdani, Arch. p. 1. Zool. Modena, iii, p. 78 (1863).
Eulalia, Kertesz, Kat. Dipt, iii, p. 62 (1908).
GENOTYPE, Musca hydroleon, L. (Europe); by designation of
Westwcod (1840).
Head about as broad as thorax, transverse ; eyes rather flattened,
bare or hairy, contiguous in <$ , upper facets larger than lower
ones, broadly separated in $ ; face prominent, pubescent ; pro-
boscis rather long; palpi small, cylindrical. Antennae with first
two joints short, 1st longer than 2nd ; 3rd much longer, cylin-
drical, tapering, with four to six annulations, last two sometimes
62 STBATIOMYIDJE.
forming an apical style, with or without a minute terminal bristle
(fig. 2). Thorax subquadrate or oblong; scutellum with two
spines. Abdomen o-segmented in cf , 6- or 7-segmented in 5 ,
normally barely broader, sometimes much broader than thorax,
generally nearly bare ; genitalia small. Leys simple. Wings of
normal Stratiomyid type; anterior veins crowded together; 1st
and 2nd veins almost anastomosed, rather short, 3rd vein normally
forked, both branches ending far before wing-tip ; 4th vein with
three endings, of which one or more may be incomplete or absent ;
posterior cross-vein present ; squamae distinct, thoracic pair
pubescent.
Range. World-wide.
Life-history, One or two European species have been bred, and
0. cyanea at Pusa. Larva much as in Stratiomyia, but without
spine- shaped protuberance. The imagines occur in moist situa-
tions, with a partiality for settling on TJmbelliferae.
Talle of Species.
1. Shining metallic-blue species (with a general
resemblance to the Muscid, Chrysomyia
dux} cyanea, sp. n., p. (53.
Never metallic ; always yellow or brownish,
with markings 2.
2. Abdomen as well as thorax conspicuously [p. 64.
pubescent rufoabdominalis, Jirun.,
Abdomen practically bare, normal ; thorax
generally considerably pubescent 3.
3. Thorax with sharply denned spaces of shin-
ing black and reddish brown, and a short
golden tomentose stripe from anterior
margin ; a conspicuous and rather ab- [p. 65.
normal species pulchcrriina, sp. n.,
Thorax dark, with yellowish or grey pubes-
cence ; species of normal appearance .... 4.
4. Some of the femora with distinct broad black
bands 5.
No distinct femornl bands, legs generally
wholly pale 6.
5. All femora with broad black median band ;
hind tibiae with broad apical band minuta, F., p. 65.
Hind femora not banded rubrithorax, Macq.,
C. (a) Abdomen wholly orange ; postocular [p. 07.
orbits in § mo'derately broad lutatius, Walk., p. 70.
(b) Abdomen with distinct dark transverse
bands; postocular orbits in $ mode-
rately broad transversa, sp. n., p. 71 .
(c) Abdomen with dorsal row of spots, more
or less forming a median stripe .... 7.
7. Frons in J black ; facial bump small,
blackish ; postocular orbits in $ mode- [p. 67.
rately broad kashmirensis, sp. n.,
ODONTOJITIA. 63
Frons in tf orange, or if black, then facial
bump more or less orange ; postocular
orbits in $ inappreciable 8. [p. 68.
8. Frons and face all orange in both sexes .... dorsoangiulata, sp. n.,
Frons and face black in both sexes ; 5 with
facial bump brownish orange solennis, Walk., p. 69.
It is impossible to include viridana, Wied. (p. 72), in the table,
but it falls into section G, and my transversa may be identical with it.
33. Odontomyia cyanea, sp. nov.
c? $ . Head in c? with eyes bare, contiguous for nearly half the
distance from vertex to frontal triangle; upper facets rich brown
or violet-brown, lower ones black ; vertex black, with a few black
hairs ; the small frons and the face and lower part of head bright
orange, bare, shining ; antennae orange, 3rd joint from dull
orange-brown to blackish, tip generally darker, twice as long as
first two joints together, with four annulations and small yellowish
apical style ; proboscis black ; eye-margins developed on inner
lower edge of eyes, orange, with a little greyish pubescence ; no
postocular orbits, the facets running out to the extreme margin
above and at sides. In $ , frons one-third of head, orange, bare,
just perceptibly broader at vertex and towards antennas ; latter
rather longer than in c?; no postocular orbits. Thorax deep
blue, brilliantly shining metallic, closely punctulate, with micro-
scopic grey pubescence ; scutellum similar ; pleuras more blackish,
more or less tomentose pubescent, with a little longer grey
pubescence on mesopleura and sternopleura and below shoulders.
In £ four longitudinal, silvery-grey, subequai and equidistant
tomentose stripes, the two middle ones united on anterior margin ;
scutellum also covered with silvery tomentose pubescence. Abdo-
men in <$ deep blue, shining, minutely punctulate ; side-margins
and tip of abdomen apple-green, sometimes only narrowly (but
always continuously) so, but varying in width up to one-third of
the abdomen on each side. In $ as in g , but the green side-
margins always very narrow though continuous, but not including
the tip. which is always blue in this sex, whereas in 3 the green tip
of the abdomen is always continuous with the green side-margins.
Venter wholly apple-green in both sexes. Legs black, a little
silvery pubescence on outer sides of anterior tihise ; front tarsi
(especially the metatarsi) distinctly broader, thicker and shorter
than middle tarsi. Wings quite clear ; veins in anterior part of
wing brownish orange or yellowish ; stigma brownish ; the 3rd of
the faint veinlets issuing from the discal cell practically absent ;
halteres bright apple-green.
Length, 6^-8 mm.
Described from a long series of both sexes, mostly bred at Pusa,
26.iii. 1908, 2.ii. 1909, 3-18. iii. 1909. Types sent to British
Museum ; co-types in Indian Museum and my collection.
64 STRATIOMYIDjE.
34. Odontomyia rufoabdominalis, Brun. (PL I, fig. 24.)
Odontomyia rufoabdominalis, Branetti, Rec. Ind. Mus. ix, p. 205
(1913).
<$ 5. Head black, a little shining: eyes bare, absolutely' con-
tiguous for a short space, leaving a very elongate narrow vertical
triangle ; vertex considerably prominent, with a few black hairs ;
ocelli pale yellowish ; face but little prominent, with short black
and grey hairs ; antennae black, 1st and 2nd joints normal, 1st
black, 2nd dull brownish (3rd missing) ; two small tufts of grey
hairs immediately above antennae, and a trace of grey hairs along
the sides of face. In $ . frons one-third of head, sides parallel,
and with face wholly slightly shining black, with sparse short
black pubescence ; the ground-colour similar to that of the thorax ;
ocelli very distinct, wide apart, brownish yellow, ocellar bump
large, moderately elevated; mouth-opening, proboscis, palpi, and
lower part of head black. Thorax black, with minute white
patches, slightly shining, dorsum and sides with moderately thick,
not long, black and grey hairs, set in minute black sockets, which
give the dorsum a granulated appearance, often two or three
hairs emerging from the same socket ; small patches or bunches of
grey hairs occur around the base of the wing and on the pleurae.
Scutellum clothed like the thorax, with a fringe of rather ragged
grey hair on posterior border and a short blunt spine at each hind
corner. Abdomen: ground-colour of 1st and 2nd segments and
major portion of 3rd, 4th, and 5th black, a little shining ; the sides
of the dorsum, nearly from the shoulders to the extreme tip, and
the hind margins of the 3rd, 4th, and 5th, and the bulk of the 6th
segment all bright reddish orange with concolorous pubescence ;
the black parts with sparse black pubescence ; a small patch of
greyish hairs on hind corners of 2nd segment, and some yellow
hairs extend along the narrow red side-edges of the abdomen,
nearly or quite to the shoulders. A7enter dull reddish orange,
becoming brighter towards tip ; central part of most of the seg-
ments more or less black ; pubescence wholly reddish orange,
j-hort, depressed. Leys black, with a little short greyish hair ;
under and inner sides of tibiae towards tips and about the apical
half of the anterior and the whole of the hind tarsi (the latter
lengthened), orange-yellow, with short golden pubescence. Wings
yellowish grey ; stigmatic region black ; major portion of middle
of wing from anterior to posterior border darker brown, the colour
fading away gradually; halteres buff; thoracic squamae milk-
white, with concolorous soft hairs.
Length, 17 mm.
Described from a J, Darjiling, 1000-3000 ft,, vi. 1912, and
a $, Siugla, Darjiling Dist., 1500 ft.,, v. 1913, in the Indian
Museum.
A very handsome and conspicuous species.
ODONTOMYIA. 65
35. Odontomyia pulcherrima, sp. nov.
5 . Head distinctly broader than thorax; vertex and frons fully
one-third the width of head, the latter slightly wider towards
antennae, with short black hairs ; whole head below antennae
paler yellow, with soft golden hairs ; antennse black, 3rd joint
about 2^ times as long as 1st and 2nd together ; proboscis black ;
occiput yellow, blackish in centre ; posterior orbits of eyes
narrowly yellow ; vertical triangle black, small ; ocelli pale orange.
Thorax with greater part of dorsum down to the suture, except
anterior and side-margins, occupied by a large shining black spot,
bare except for a broad short stripe of golden pubescence, begin-
ning just behind the anterior margin but reaching only halfway
to the suture ; on each side behind the suture (just clear of the
side-margins) a semicircular black spot (the straight side inwards)
and contiguous to the large black spot above the suture, thus
leaving an approximately square space occupying more than half
the surface behind the suture, and reaching the posterior border
of the dorsum with sharply defined sides; this space is reddish
brown, and the large anterior black spot just perceptibly encroaches
on the anterior margin of it. Scutellum large, reddish brown, as
are the spines and the metanotum ; sides of thorax wholly pale
yellow with bright golden-yellow pubescence. Abdomen mode-
rately shining black, bare ; basal half of 1st segment, a large sub-
triangular spot on each side of 2nd extending from anterior to
hind margin, an elongate transverse spot towards each side on
hind margins of 3rd and 4th segments (those on latter rather
smaller), and a long spot narrowed to a point anteriorly on 5th
segment, all light yellow. Venter wholly yellowish. Legs with
coxae yellowish or yellowish brown, with yellow hair ; front legs
black, brown at base of femora ; on apical half of outer side of
tibiae a fan-like row of stiff black hairs ; middle legs black, basal
half of femora yellowish white, a similarly-coloured band on
middle tibiae ; hind legs black, basal half of femora yellowish
white. Pubescence on all legs generally concolorous with ground-
colour, some fairly coarse black pubescence at base of middle
tibia? ; tarsi with golden-brown pubescence below ; pulvilli
brownish orange ; claws orange, blai-k-tipped. Wings pale
yellowish brown; costal cell and extreme base of wing, also
beyond discal cell, pale grey ; stigmatic region blackish brown ;
veins dark brown ; 3rd vein forked very near tip ; halteres pale
dull yellow.
Length, 10 mm.
Described from a unique $ in the Indian Museum from the
Peshoke District, Darjiling, 2000 ft., 26. v.-14. vi. 1916 (Gravely}.
36. Odontomyia minnta, F.
O'lontomyia minuta, Fabricius, Eut. Syst. iv, p. 268 (1794X
Odontomyia ochracea, Brunetti, Kec. Ind. Mus. i, p. 129 (1007).
GO STEATIOMYID^E.
Odontomyia submutica, Brunetti, loc. cit. p. 130 (1907).
Oxycera indica, Brunetti, loc. cit. p. 119 (1907).
? O.pusilla, Eabricius, Ent. Syst. iv, p. 271 (1794).
3 $ . Head with eyes in ^ microscopically pubescent, barely
contiguous at nearest approach, and for a very short distance only,,
gradually separating to vertex ; i'rons shining black, with yellowish
hairs ; ocelli yellowish ; frontal triangle with yello \vish -grey
pubescence ; face and underside of head yellowish, with a little
whitish pubescence ; proboscis large, black ; antennae with 1st
joint distinctly longer than 2nd, orange-yellow, 3rd brownish,
4-annulatecl, tip black ; occiput black. In $ , whole head, in-
cluding the rather broad postocnlar orbits, yellow or orange-
yellow ; a broad shining black band across vertex from eye to eye,
sometimes with the appearance of being formed of three large
contiguous spots ; a pair of large oval shining black spots on frons,
placed transversely and slightly diagonally ; a pair of similar
smaller rounded spots below level of antennae and just below the
larger ones. Thorax black, with very short golden or greyish
pubescence ; sides black, a large pale yellow oblong spot below
shoulders, another in front of wing-bass, the two spots generally
connected above ; a third yellow spot below wing-base. 8cutellum
yellow, base black, spines generally extremely small, occasionally
normal in size. Abdomen of d1 normally yellowish, with a large
rounded or somewhat diamond-shaped spot spread over middle of
first two segments ; a spot on middle of 3rd segment from anterior
to hind margin, narrowed behind ; on anterior margin towards
sides of 2nd segment, a small spot ; a similar larger one an each
side of 3rd segment ; on 4th and 5th segments a broad band on
anterior margin tilling about half the surface, the two bands con-
nected by a short median stripe ; all the above markings black.
In some males the marks are reduced to a dorsal row of small
detached rounded or angular spots ; in one d the abdomen is
wholly yellowish except for a single small black spot on 2nd seg-
ment. In $ a large spot similar to that of the d spread over
middle of 1st and 2nd segments, with a small spot towards each
side in a line with it; sometimes the three spots more or les»
united by a short connecting stripe ; in one $ the spots were re-
placed by a definite uninterrupted broad band ; abroad black band
on anterior margin of 3rd, 4th, and 5th segments nearly filling the
sufraces and sometimes united in the median line by a dorsal
stripe. Venter in both sexes mainly yellowish. Leys brownish
yellow; femora with a broad median dark brown band; hind tibiae
with a similar band on apical half, generally leaving the lip pale ;
tips of tarsi barely darker. Witu/s absolutely colourless, anterior
veins and stigma very pale yellow; discal cell unusually small,
occasionally larger in individuals ; discal cell veinlets almost
invisible ; halteres yellowish.
Length, 4-5 mm.
Described from several specimens of each sex, including types of
ODONTOMYIA. 67
ochrace(^,sulm^ltica,Sind^ndic(l : Siliguri ; liampore, Chaka, United
Provinces, 23-31. ii. 1907 ; Eanchi ; Bareilly ; Calcutta, 21.iii.
J907, 14. vi. 1907, 22. vi. 1907 (Paiva), 23. viii. 1907, 28. ix. 1907.
My indica was described from Chapra, Bengal (Mackenzie}.
The four prominent black spots on the front of tlie head, the
bands on the legs (though these are sometimes much paler than
usual), the very reduced discal cell, and very small scutellar spines
all combine to characterize very distinctly this variable species.
The form with wholly yellow abdomen, except for a single small
black spot at base of 2nd segment, is represented by a . #mtZwith frons almost linear, brilliantly shining violet and
blue, with short soft black hairs ; ocelli placed much below the
vertex, small, pale yellowish brown, ocellar triangle very slightly
elevated; the slight protuberance above the antennae pale whitish
yellow ; face blackish, with black hairs, lower part of head reddish
brown, prominent ; back of head black, with a fringe of white hairs.
Antennae with 1st and 2nd joints black, shining, 3rd joint and
the overlapping inner side of the 2nd joint coffee-brown; arista
black, apical. Thorax dorsally and scutellum brilliantly shining
violet, with rather copious brownish-grey hairs ; thorax below
76 STttATIOMYIDJE.
dorsuin brilliantly shining peacock-green, with short whitish
hairs ; a small brownish-yellow spot on the shoulders is connected
with the wings by a very narrow brownish-yellow ridge delimiting
the dorsal and ventral surfaces. Abdomen brilliant shining violet,
with pale grey hairs ; venter similar. Legs : femora black, with
short white hairs, a little brownish yellow at base ; the knees of
similar colour and the coxae with a mark or two of the same shade ;
tibiae black, more or less yellowish on apical part, the whole surface
with short pale yellow hairs ; tarsi dark, with pale hairs ; under-
side of hind tarsi with short, very dark golden-brown pubescence.
Wings grey ; stigma ill-defined ; this region of the wing brownish ;
veins dark brown ; halteres yellowish,
Length, 14 mm.
Described from one $ in the Indian Museum from Ukhrul,
Manipur, Assam, 6400 ft. (Rev. W. Pcttiyrew).
46. Ptecticus aurobrunneus, sp. nov.
c? . Ifead with vertex and frons down to its narrowest part deep
orange ; former with golden and brown hairs intermixed, latter
with golden hairs only; frontal triangle bright lemon-yellow,
nearly bare ; oeellar triangle black, ocelli with a reddish tinge ;
face, mouth, and proboscis orange-yellow ; antennae orange, arista
black, orange at base ; eyes black ; occiput yellowish, reduced,
owing to the margins of the eyes themselves curving backwards
considerably. Thorax dorsally and scutellum dark chestnut-
brown, with just a suggestion of three darker stripe?, wholly
covered with short, dense, though inconspicuous, golden pu-
bescence ; propleura, pteropletira, metapleura, and metanotum
lighler chestnut-brown ; mesopleura and sternopleura shining
blackish ; whole sides of thorax, even to the mesopleura, with
short dense golden pubescence, though at first sight they appear
to be bare. Abdomen with 2nd to oth segments dark chestnut-
brown, with short dense golden pubescence, which is invisible
when viewed from in front, but from that view a large irregularly
shaped black spot is seen in the centre of each segment, almost
forming a dorsal stripe; 1st segment and tip of abdomen black,
with some yellow or golden hairs ; genitalia and vertex dark
brown, the latter with microscopic golden pubescence, only visible
when viewed from in front. Leys with front coxa3 brownish
orange ; posterior coxae dark brown, all with a little golden hair ;
rest of legs bright orange ; posterior femora black on basal half ;
tips of tarsi a little darker. Whigs rather dark brown ; a small
space in both basal and discal cells slightly paler, and traces of
similar lighter spaces in the centres of some of the other cells;
halteres pale yellowish.
Length, 21 mm.
Described from a unique J in the Indian Museum from
Purambikulam, Cochin State, 1700-3200 ft., 16-24. ix. 1914
(Gravely}.
PTECTICUS. 77
47. Ptecticus wulpii, Erun.
Ptecticus wulpii, Brunetti, Rec. Ind. Mus. i, p. Ill (1907); id.,
op. cit. ix, p. "263 (1913).
Ptecticus apicalis, Wulp (nee Loew), Notes Leyd. Mua. vii. p. 62
(1885).
c? $ . Differing from P. aurifer. Walk., as follows : —
Abdomen normally all orange to tip, 1st segment a little darker,
and often, if viewed from behind, a large blackish spot is seen in
the middle of each-segment, sometimes diamond-shaped, sometimes
irregular in shape, covering nearly all tho surface of each segment ;
genitalia brownish orange, this latter being the best specific dif-
ference of all. Another difference, which seems equally constant,
is in the legs, which are all orange to the extreme tips except for
the moderately dark brown posterior coxae and basal half of the
posterior femora. The black apical part of the wing is of much
less extent, not beginning till some distance beyond the discal
cell ; some specimens have the wing darkened from the tip only
up to midway between the discal cell and the wing-tip.
Same size as aurifer.
Described from two d1 d1 in the Indian Museum, one rf one $
in the Pusa collection : Darjiling District, 1000-3000 ft., v. 1912 ;
Singla, iv.1913; Nilgiri Hills, 3500 ft. (Andreives), 7000 ft.,
v. 1904 (llowson); Panunbikulam, Cochin State, 1700-3200 ft.,
16-24. ix. 1914 (Gravely),
Unless P. aurifer,W&\k., should prove to have fulvous genitalia,
P. ivulpii is easily separated from all the other species in this
group bv that character alone. It presents an extraordinary
resemblance to the Tipulid, Pselliopliora compedita, Wied., and
both have been seen by me from the same locality.
48. Ptecticus apicalis, Lw.
Ptecticus apicalis, Loew, Verh. zool.-bot. Ges. Wien, v, p. 142, pi.,
figs. 3, 4 (1855) ; Brunetti, Rec. Ind. Mus. i, p. 110 (1907).
Sargus hiridus, Walker, Proc. Linn. Soc. Lond. i, p. 8 (1856).
Brownish red-yellow, shining ; 4th abdominal segment with a
very large black spot, the next wholly dull black, the rest deep
black, rather shining; venter much as upper side of abdomen.
Frons very narrow, reddish yellow; lower part whitish yellow;
ocelli placed on a blackish cross-band; frontal triangle whitish
yellow, puffed up. Antennae light reddish yellow, 3rd joint barely
visibly annulated, tip truncate. Legs concolorous with body ;
hind tibiae always darker towards tip ; front and middle tarsi with
last two joints darker, also hind metatarsus. Wings large, reddish
yellow up to beyond discal cell, becoming grey towards hind
margin ; a large blackish-brown spot filling more than apical
third of wing.
STRATIOMYiDJt.
Lemjtli, 15 mm.
Described from Pulo Penang ; in "Westermann's collection,
Vienna Museum.
49. Ptecticas aurifer, Wall:
Sargus aurifer, Walker, List Dipt. Brit. Mus. v, p. 96 (1854).
Ptecticus aurifer, Brunetti, Rec. Ind. Mus. i, p. 110 (1907).
3 2 ' Head with eyes in <$ with their point of nearest approach
distinctly below halfway between vertex and base of antennae,
the space between them perhaps ^ to \ of a millimetre ; thence
upwards frons widening gradually till at vertex it is almost 1 milli-
metre broad ; bright orange with orange pubescence ; frontal
triangle elevated, shining, smooth, lemon-yellow, bare, except for
a smull tuft of orange hairs just above base of antennas ; face very
small, yellowish, as are the very narrow inner lower eye-margins
near the very large mouth-opening ; occiput black, upper margin
with yellow pubescence ; ocelli yellow on a black ocellar triangle ;
proboscis and the very large labella orange-yellow, with a few
hairs. Antenna? orange, tip of upper side with a black mark,
arista black, orange at base. Frons in $ at narrowest part dis-
tinctly broader than in c? , so that it is only about two and a half
times broader at vertex than at frontal triangle. Thorax : sides,
scutellum, and metanotum bright orange, all with a little very
short yellow or orange pubescence, which is a little longer on the
sternopleura, metapleura, and metanotum ; dorsum sometimes with
a little more brown in it and sometimes, especially in the $ ,
traces of three longitudinal brownish stripes can be seen. Ab-
domen normally with first four segments bright orange, remainder
(including genitalia) black, but the orange segmentsof ten have more
or less brown in them, sometimes in the form of large transverse
spots occupying neaily all the segments; sometimes the whole
dorsum is uniformly more brownish ; the first four segments with
very short inconspicuous golden-yellow pubescence, the rest with
almost microscopic greyish pubescence. Yenter mainly orange on
first four segments, remainder black, but 4th, and sometimes 3rd
also, often bear large blackish patches. Legs all orange, with short
yellow pubescence ; last two joints of anterior tarsi brown ; about
apical halt' of hind tibiae brown ; hind tarsi blackish brown,
especially first two joints, underside with rich golden-brown
pubescence. Wings with basal half up to just beyond discal cell
bright orange-yellow, remainder moderately dark brown ; veins a
little deeper orange, veinlets from discal cell dark brown, 3rd only
moderately sinuous ; halteres orange.
Lemjtli , 1 2-20 m m .
Described from a series in the Indian Museum, Sarawak
Museum, and Pusa collections: Mussoorie, 6500 ft. (Bond};
Darjiling, 1000-3000 ft., v-vi. 1912 ; Peshoke Spur, Darjiling
PTECTICUS. 79
Distr., 4000 ft., v-vi. 1916 ; Lebong, Darjiling Distr., 4500 ft.,
2-9. v. 1909 (Hoivlett) • Singla, Darjiling Distr., 1500 ft-., iv. 1913 ;
Siliguri, base of Darjiling Hills, 18-20. vii. 1907 ; Sikkim ; Mar-
gherita, Assam; Nongpoh, Khasi Hills, Assam, 1000-3000 ft.,
vi. 1905 ; also from Matang, Trusan, Klinhang, and Kuching
(all Borneo).
This is the commonest Indian species, apparently.
50. Ptecticus anstralis, Sch.
Ptecticus austraKs, Schiner, Novara Reis., Dipt. p. 65 (1868) ;
Brunetti, Rec. Ind. Mus. i, p. 113 (1907).
<$ $ . Head : eyes in ).
Easily known from all other Indian species by its great size and
splendour. The Indian Museum specimens from Tenasserim
were named by Bigot as his maytiifaus, but. I now feel sure they
are synonymous with Walker's yemmifer, and possibly pubesccns,
Wulp. may be another synonym.
53. Sargus metallimis, F. (PI. I, tigs. 22, 23.)
Sar ) pubescence ;
pletirce concolorous, with shorter pubescence. Abdomen metallic
shining yellowish bronze or cupreous in tf , violet or blue in $ , with
dense though not conspicuous yellow pubescence towards sides in
(5 , the dorsal region often nearly bare ; whole surface in $ with
short white pubescence, a little longer towards sides ; venter
generally dark brown or aeneous, shining, with microscopic
pubescence. Legs pale yellow, with very short pale yellow ( d1 )
cr white ( $ ) pubescence. Wings pale yellowish grey or pale
brownish, stigma a little darker; halteres pale yellowish or
white.
Length, 9-10 mm.
Described from a considerable number of both sexes in the
Indian Museum, Pusa, and Dehra Dun collections and my own.
Common and widely distributed in India in May and July to
October up to 5000 ft. ; also generally distributed in the East,
Rangoon, Singapore (both Srunetti); Borneo, Java, Amboina,
Aru Islands, Shanghai.
Bigot in 1887 described a second species, coming from North
America, under the name pallipes.
54. Sargus mactans, Walk.
Saryus mactans, Walker, Proc. Liun. Soc. Lond. iv, p. 97 (1860) ;
Bruuetti, Reo. Ind. Mus. i, p. 107 (1907) ; de Meijere, Tijd. v. Ent.
liv, p. 263 (1911).
cJ . Head with frons extremely narrow, the sides never parallel,
widening gradually above and below from point of nearest
approach, aeneous yellow or brassy, with yellow pubescence on
vertical and frontal triangles ; ocelli at some distance below
vertex, upper ones contiguous to eye-margins ; face, mouth-
opening, and proboscis yellowish orange, with a little yellowish
hair ; antennas orange ; occiput black, a fringe of whitish hairs on
lower margin. Thorax with dorsum, pleurae, scutelluiu, and
metanotum brilliantly shining cupreous green, with rather dense
soft yellow pubescence, which becomes whitish on anterior margin
of dorsum ; a shoulder-spot, a narrow stripe thence to wing-base,
extending over upper part, of sternopleura, brownish or yellowish.
Abdomen brilliantly shining cupreous, sometimes with a green
tinge, with rather dense yellow pubescence towards and at sides,
G2
84 STKATIOMYID.?;.
the pubescence in the middle of the segments almost microscopic ;
venter blackish, shining, with short soft yellow pubescence ;
genitalia with a violet tinge, rather small, with two terminal,
cylindrical, palp-like lamellae. Legs yellow, coxae more or less
black at base, especially hind pair ; hind tibiae slightly constricted
just beyond middle, with nearly basal half black; hind tarsi with
tips dark ; all legs with minute bright yellow pubescence, which
is longest on hind tibise. Wings uniformly yellowish grey ;
stigma a little darker, brownish yellow ; halteres yellowish.
Length, 9-11 mm.
Described from several specimens in the Indian Museum :
Bhim Tal, Kumaon, 4500 ft., 19-22. ix. 1906 (Annandale) ;
Almora, Kumaon, 5500 ft,, 10-21. ix. 1911 (Paiva)- Shillong,
4900 ft., l.vi. 1918 (.K«o); Katmandu, Nepal. It occurs also in
Ceylon and the Malay Archipelago.
A variety with a black streak on the posterior femora (two
specimens)'was taken at Shillong, 5000 ft. (Fletcher) and 4900 ft,,
30. v. 191 8 (.Rao).
Genus CHLOROMYIA, Dune.
Chloromyia, Duncan, Mag. Zool. Bot. i, p. 164 (1837).
Chrysomyia, Macquart, Hist. Nat. Ins. Dipt, i, p. 262 (1834).
Myochrysa, Rondani, Dipt. Ital. Prod, iv, p. 11 (1861).
Myiochrysa, Verrall, apud Scudder, Nomeu. Zool. p. 204 (1882).
GENOTYPE, Musca formosa, Scop. ; by Verrall's designation
(1909).
Differing from Sargus mainly in the densely pubescent eyes,
which are contiguous for a considerable distance in the d , the
upper facets larger than the lower ; and wide apart with equal-
sized facets in the $ ; 3rd antennal joint longer than in Sargiis :
abdomen rather broader also, especially in the $ ; venation rather
more indistinct,
Range. Europe, South Africa, Asia, North America.
Life-history. The common C. formosa, Scop., of Europe, has
been bred from garden-mould and from Jirassica rufa.
Some confusion has arisen over the generic names Chloi'om>/ia
and Ghrysomyia (under which formosa was known in Europe for
many years), which is concisely cleared up by Verrall (Brit. Flies,
v, p. 1ST). Coquillett claimed designation of the same species as
generic tvpe in 1910, presumably unaware of Verrall's priority.
55. Chloromyia sapphirina, DW/r.
tiaryus sapphirimts, Walker, List Dipt. Brit. Mus. iii, p. 519 (1^49).
" Body bright blue, tinged with purple ; head as broad as
chest ; clypeus white ; eyes dark bronze ; mouth and feelers black ;
chest and abdomen finely punctured, thinly clothed with short,
13RACITYCARA. 85
hoary hairs ; abdomen nearly elliptical, a little broader and longer
than the chest ; legs blue, clothed with short hoary hairs; shanks
and feet piceous ; shanks pale yellow beneath ; wings colourless ;
wing-ribs and veins dark tawny ; poisers pale tawny. Length of
the body 4 lines, of the wings 7 lines.
"East Indies. From Archdeacon Clerk's collection. In the
British Museum."
Though described from the East Indies, it is probably from
East India.
Genus BRACHYCARA. Thorns.
Brachycara, Thomson, Eug. Eesa, p. 460 (1868) (1869).
GENOTYPE, Brucliycara ventralis, Thorns.
Very near Cliloromyia. Head slightly narrower than thorax,
nearly hemispherical ; face not prominent, convex ; mouth-
opening large ; proboscis short, labella broad ; eyes bare.
Antennae short, 3rd joint almost transverse, obconical, a little
shorter than 2nd, six-ringed, last two annulations abruptly
narrower. Thorax with humeri distinct. ; scutellum semicircular,
unspined. Abdomen oval, broader than thorax, 8-segmented ;
last three segments narrower, retracted. Legs moderately strong,
slightly pubescent ; anterior coxae small, subovate, very little
distant from middle pair ; hind pair subtriangular ; femora
clavate fusiform ; tibiae hardly shorter than femora, unspurred ;
tarsi hardly longer than tibiae, 1st joint longer than the remaining
joints together. Wings longer than abdomen.
Range. B. ventraHs is the only species.
56. Brachycara ventralis, Thorns.
Brachycara ventralis, Thomson, Eug. Resa, p. 461 (1869) ; Brunetti,
Eec. Ind. Mus. i, p. 104 (1907).
c? $ . Head shining, with silvery-white pubescence, thicker
below ; frontal triangle shining black, a small subtriangular white
spot contiguous to each eye ; irons with a median furrow ; eyes
in 3 broadly contiguous, bare ; proboscis yellowish white ;
antennae brown, yellow in middle. Thorax and scutellum black,
with depressed pubescence, which is golden brown and almost
scale-like on the humeri ; a narrow line on mesopleurae whitish
yellow. Abdomen yellowish red, a basal and apical blackish- brown
spot ; venter wholly shining yellowish red, with very short
pubescence. Wings subhyaline, veins yellow ; halteres pale
yellow, clubs large. Legs yellowish red, all the tarsi and tips
of hind tibiae blackish brown.
Length, 5 mm.
Cocos-Keeling Islands.
The foregoing is a translation of Thomson's description,
86 STBATIOMYID^B.
Genus PROSOPOCHRYSA, Meij.
Prosopochrysa, de Meijere, Tijd. v. Ent. 1, p. 220 (1907).
GENOTYPE, Chrysochlora vitripennis, Dol. ; by original de-
signation.
Near Microclirysa. Eyes bare, narrowly separated in d , facets
of equal size ; occipital orbit extremely narrow ; frons prominent ;
antennae rather short, 1st and 2nd joints subequal, 3rd as long as
first two together, with four annulations, the 4th the smallest,
pubescent ; arista practically apical. Thorax and abdomen sub-
equally long, latter 5-segmented ; scutellum unarmed. Third
vein simple, 2nd nearly erect ; discal cell 5- sided, only two veinlets
issuing from it ;' the 3rd absent, and 3rd and 4th posterior cells
therefore united.
Range. India, Java.
The absence of the 3rd veinlet from the discal cell might make
the species appear to belong to the PACHYGASTRINVE, but in that
subfamily the fifth posterior cell is in full contact with the discal,
through the posterior cross-vein being absent.
57. Prosopochrysa vitripennis, Dol.
Chri/sochlora vitripennis, Doleschall, Nat. Tijd. Ned. Ind. x, p. 408,
pi. xi, fig. 2 (1856).
Microchrysa albitarsis, Brunetti, Eec. Ind. Mus. viii, p. 156 (1913).
c? $ . Head of tf with upper part of frons shining black or
dark green, very elongate, and narrowed to point of nearest
approach of eyes; below this point broad and prominent,
brilliantly shining blue or violet ; head below antennae metallic
bronze-green, \\ith black or dark brown pubescence; cheeks
Fig. 5. — Prosopochrysa vitripe-nnis, Dol., lateral view of head.
white-haired ; proboscis black ; occipital orbit very narrow, black,
with whitish grey shimmer. Antennae wholly brown, sometimes
1st joint blackish, 2nd and 3rd yellowish; latter with four
annulations ; arista long, practically apical, a little thickened
basally where there are a few hairs on underside. In ? , frons
PROSOPOCH HYSA . — M ICIIOCHBYSA . 87
one-fifth width of head, slightly tinged with blue, brilliant violet
immediately above anteuriae. Thorax punctate, wholly brilliantly
shining metallic blue or blue-green, with very short whitish
pubescence. Abdomen metallic black or dark bronze-green, with
very short sparse whitish pubescence, which is a little longer
round margin and tip ; venter darker, nearly black. Legs black,
with microscopic pubescence ; all the tarsi yellowish white except
Fig. 6. — Prosopochrysa vitripeimis, Pol., wing.
at tips. Wings clear, veins and stigmatic region yellowish ; 3rd
vein simple, straight, ending at some distance before wing- tip ;
halteres apple-green.
Length, 5 mm.
Redescribed from two c? d1 and three $ $ in the Indian
Museum, and some in the Pusa collection.
Genus MICROCHRYSA, Lw.
Microchrt/sa, Loew, Verb. zool.-bot. Ges. Wien, v, p. 146 (1855) ;
Brunetti, Rec. Ind. Mus. i, p. 103 (1907).
Clorisoma, Eondani, Dipt. Ital. Prod, i, p. 168 (1856).
Chlorosia, Rondarii, op. cit. iv,p. 11 (1861).
G-ENOTYPE, Musca polita, L. (Europe) ; by original designation.
Head broader than thorax, face small, slightly pubescent ;
occiput concave in J , posterior eye-orbits distinct in $ ; ocelli
equidistant ; eyes bare, in d1 contiguous, upper facets larger than
lower ones, in $ widely separated, facets equal in size. Antennae
of typical Sargid form, 3rd joint with four annulations, arista
practically apical. Thorax longer than broad, slightly wider
behind, shining metallic, with short dense inconspicuous pu-
bescence. Abdomen very short, rounded, shining ; pubescence
extremely short, though sometimes dense. Legs simple, tibiae
sometimes slightly dilated apically. Wings of normal Sargid type,
terminations of 4th vein less parallel ; thoracic squamae more
normally shaped than in Saryus, pubescent.
Ln all minor characters mainly as in Sargits.
Itange. Europe, Asia, South Africa, North and South America.
Life-history. The common M. polita, L., of Europe, seems to
breed freely in cow-dung and decaying vegetable matter.
M. ca1oj>a differs from M.jlaviventris, in which the abdomen is
imicolorous, by a narrow pale border along the sides and tip,
STRA.TIOMYM>.E.
58. Microchrysa flaviventris, \V\ed.
Sargusflaviventris, Wiedemann, Anal. Ent. p. 31 (1824).
Microchrysa Jlaviventris, Brunetti, Kec. Ind. Mus. i, p. 103 (1907).
Sargus affinis, Wiedemaun, luc. cit. ; Brunetti, loc. eit.
Chrysornyia annulipes, Thomson, Eugen. Resa, p. 461 (1869).
Microchrysa gemma, Bigot, Ann. Soc. Ent. France, (f>) ix, p. 231
(1879) ; Brunetti, Rec. Ind. Mus. i, p. 103 (1907).
c? $ . Head in <$ , vertex black, shining, with a few black hairs ;
ocelli distinct, red ; e3res contiguous for rather more than half the
distance from vertex to frontal triangle ; upper facets bright
brown, much larger than the blackish lower ones ; frons blackish,
with a few short pale hairs ; face brilliantly shining metallic
green, bare ; proboscis and palpi rather bright yellow ; antenna?
pale yellow, with 1st and 2nd joints subequal, 3rd rather longer
than 2nd, with pale yellow arista ; occiput black, very concave ;
no definite postocular orbit. In $ , frons forming one-third of
the head, rather broader at vertex, brilliantly shining metallic
blue-green, carried over the vertex to the blackish occiput and
bearing a little short grey pubescence ; remainder as in J .
Thorax brilliantly shining metallic green, with very sparse pale
yellow pubescence on dorsum and parts of the pleurae ; scutellnm
similarly coloured. Abdomen in c? brownish yellow, varying to a
little lighter or more reddish individually, with microscopic pale
yellow pubescence, 5th segment blackish ; venter yellowish, often
more or less marked with black. In 5 , brilliantly shining green,
with microscopic pale yellow pubescence ; venter generally
brownish, sometimes yellowish or even metallic green. Legs
pale yellow ; hind femora with broad median brown band and
hind tibia? with broad apical band, both bands sometimes much
paler. Wings colourless ; veins and the rather large stigma pale
yellow ; halteres yellow.
Length, 3^-4 mm.
Described" from a considerable number of both sexes in the
Pusa and Indian Museum collections and from other sources from
many parts of India, dated March and from June to October.
Also from Peradeniya, Ceylon, 15. vii. 1910 ; Java (Jacobson) ;
Sibu, Sarawak, 2. vii. 1910 (Beebe). I have taken it myself in
Mussoorie and Darjiling. Common and widely distributed in
India and the East. Van der "Wulp records it from Papua.
M. affinis, Wied., from East India, and M. gemma, Big., from
Ceylon, are probably synonyms.
59. Microchrysa calopa, Bnm.
Microchrysa calopa, Brunetti, Rec. Ind. Mus. vii, p. 453 (1912),
$ . Head with frons broad, bright metallic green on vertex,
changing insensibly to brilliant violet-blue just above antenna),
with very short pale yellow pubescence ; face metallic green ; oral
opening brown ; eyes dark brown, the upper corners rounded off,
UEBINJE. 89
thus widening the vertex considerably : occiput dark ; antennae
pale yellow, tip of 3rd joint, which has a slight reddish tinge,
shortly pubescent ; 1st and 2nd joints with one or two micro-
scopic hairs. Thorax with dorsum brilliant metallic green, with
very short pale yellow pubescence ; venter similarly coloured and
pubescent, sharply delimited from the dorsum by a narrow yellow
side-ridge at the level of the Avings ; posterior calli yellowish.
Scutellum and metanotum brilliant green, the former with pale
yellow hairs and yellow underside. Abdomen dorsally violet, not
intense, shining, and with a bluish tinge in certain lights, with
pale yellow pubescence ; the colour covers all the surface except
a narrow, irregular, yellow border round the edges ; venter yellow,
shortly pubescent. Legs wholly yellow, except a broad brown
ring on distal half of hind femora, and the blackish hind meta-
tarsus. Wings clear, stigmatic area pale yellow ; halteres deep
yellow.
Length, 3 mm.
Described from two $ $ in good condition in the Indian
Museum, from Margherita, Assam, arid Paresnath, Chota Nagpur,
4400 ft., v. 1909 (Dr. Jenkins).
Subfamily BERING.
Head semicircular ; eyes large and prominent, bare or hairy ;
antennae with 3rd joint moderately elongate and more or less
cylindrical, but not conspicuously so, with eight annulations and
no apical style or arista. Thorax roughly oblong; scutellum
generally with four or six marginal spines, occasionally eight, and
in individuals an odd number may occur ; pubescence of thorax
continued over spines. Abdomen with at least seven obvious seg-
ments, longer than thorax, dorsum flattened, sides approximately
parallel. Leys simple, moderately strong; 1st joint of hind tarsi
generally dilated in tf ; spurs on middle tibize in rare cases.*5
Wings not of the normal Stratiomyid type, all the veins, except an
occasionally obviously abortive one, running out to wing-margin :
2nd vein short, beginning opposite base of discal cell, ending near
tip of 1st ; 3rd vein forked, both branches ending before wing-
tip; 4th vein with only two terminations (as in PACHYGASTRIN.E),
as the 5th vein forms the hinder side of the discal cell ; posterior
cross-vein therefore absent ; anal cell closed at some distance
from wing-border ; squamse small.
This subfamily is easily distinguished from all others except
XYLOMTTKYE by the 7-segmented abdomen and by the anterior
veins not being crowded together ; whilst from the XYLOMYIN;E it
is separated by the 2nd vein (prffifurca) originating opposite the
base of the discal cell instead of at some considerable distance
before it, also by the open 4th posterior cell. Their usually
* Acanthomyia. for instance, a European genus.
90 STKATIOMYID.E.
aeneous or metallic colour also renders them conspicuous, and
though in this they resemble the SABGIN-E, the latter are always
easily known by the anterior cross-vein connecting the discal cell
with the praefurca instead of with the 3rd vein. The frequently
dilated hind metatarsus in the cJ is & further character of the
BERING.
The life-histories of a few European species are known, but
not that of Bens. That of Chorisops tibialis has been described
by Handlirsch. The larvae live in the earth and decaying matter.
Table of Genera.
Scutellum unspined AM.OONOSTA, Ost.-Sack., p. 93.
Scutellum spined.
Palpi minute or obsolete ; eyes con-
tiguous in cJ " BERIS, Latr.. p. 90.
Palpi rather long ; eyes not con-
tiguous in rf CHORISOPS, Rond., p. 92.
Genus BERIS, Latr.
lierit, Latreille, Hist. Nat. Crust. Ins. iii, p. 447 (1802).
Oplacantha, Itondani, Arch. Zool. Modena, iii, p. 87 (1863).
He.racantha, Lioy, Atti 1st. Yen. (3) ix, p. 586 (1864).
Octacantha, Lioy, loc. cif.
GENOTYPE, Musca clialybeata, Forst., as Stratiomys 6-dentata,
F. (Europe) ; by original designation.
Head almost semicircular, transverse, as broad as thorax ; face
short, pubescent ; eyes large, contiguous in tf (with rare excep-
tions), wide apart in $ , densely hairy in tf , upper facets larger
than lower, less hairy in $ with all equal facets. Proboscis well
developed, palpi rudimentary. Antennae porrect, nearly as long
as head; two basal joints short, subeqnal, bristlv; 3rd elongate
with eight annulations, the first and last being the longest.
Tlwrax rather arched, closely pubescent in <5 , less so in $ ;
scutellum with four to eight spines bearing pubescence (occasion-
ally an odd number in individuals). Abdomen hardly broader than
thorax, flattened, sides nearly parallel, at least 7-segmented ;
genitalia more developed than in the other subfamilies. Legs
simple, of moderate si/.e ; hind metatarsus dilated in <$ and
elongate in both sexes. Wings longer than abdomen, not of the
typical Stratioinyid nature; anterior veins not crowded together,
all veins distinct and generally reaching wing-margin ; 2nd vein
rather short, enclosing the conspicuous large dark stigma ; 3rd
vein rather widely forked, both branches ending before wing-tip ;
4th vein with only two terminations and occasionally an abortive
third ; posterior cross-vein absent as upper branch of 5th vein
forms hinder side of discal cell ; anal cell closed at some distance
from border of wing. Alar squamae of moderate size ; thoracic
ones undeveloped.
BERIS. 91
Range. Europe, Orient, Australia, North and South America.
Life-history. The European B. chalybeata has been bred from
moss, but the metamorphoses of the genus are not well known.
The perfect insects prefer marehy situations, but are generally
distributed. Col. Yerbury records the males of some British
species as performing a rather wild aerial dance.
The two Indian species of Berts are easily distinguished : —
Legs mainly dark brown yeniculata, Curt.
Legs yellow ; femora and hind tibiaa with brown bands, annulipes, Brun.
60. Beris genicnlata, Curt.
Beris geniculata, Curtis, Brit. Ent. via, p. 337 (1830).
c? $ . Head : frons in <$ rather large, shining black, with a de-
pressed median channel on upper part ; face and lower part of head
shining black ; frons and face with rather dense blackish-brown
pubescence ; vertex wholly occupied by the considerably elevated
ocellar triangle, bearing blackish-brown hairs; proboscis black,
labella orange, palpi black ; eyes contiguous for a considerable dis-
tance, with dense blackish-brown pubescence on front part, less
dense on rest of surface, facets of upper half much larger than
the others ; antenna? black ; 3rd joint not quite twice as long as
the 1st and 2nd put together,* gr-eyish on inner side about
the middle, tapering to tip after about 3rd or 4th annulation,
extreme tip with two or three minute bristles. In $ , frons nearly
one-third of head, shining black, as is the face, both with short
pale pubescence ; eyes with very short pubescence. Thorax and
scutellum metallic shining green or blue-green, the former dis-
tinctly punctate except towards hind margin; scutellum impunc-
tate ; both with rather dense blackish pubescence, which is less
dense on scutellum ; the latter with eight shining green spines
(normally in European specimens only six). In the $ , pubescence
on dorsum of thorax pale and short, rather depreised. Abdomen
very dark brown, nearly black, with rather long pubescence
towards sides, which is brownish in tf , pale yellowish and much
shorter in $ ; venter shining brown ; genitalia rather small. Legs
very dark brown, tips of femora narrowly orange ; base of tibia?
orange to about one-third on anterior legs and to a quarter on hind
pair ; anterior metatarsi pale, rest of tarsi black, hind metatarsi
pale, considerably and equally dilated throughout their length
except at base and tips, longer than four remaining joints
together, the latter black. Wings brown, costal cell a little paler ;
stigma large, dark brown, oval ; in $ wings clearer ; halteres
brownish orange.
Length, 4—5 mm.
* A slight error as to the length of the 3rd autennal joint seems to have
crept into Yen-all's usually scrupulously correct writings, as he contradicts
himself in the tahle ot species and the description. The discrepancy may not
be important.
02 8TRATIOMTIDJ5.
Described from a J and two $ $ in the Indian Museum taken
by me at Darjiling, 6900ft., 2.x. 1908.
Whether this species is the true B. geniculata of Curtis is not,
quite certain, as though it agrees in practically all other points,
the difference in the width of the frons in the $ is important :
less than one-fourth the width of the head in true yeniculata and
nearly one-third in the present form. My specimens are also
distinctly a size smaller, 4-5 mm. against about 6. B. genieulafa,
Curt., is recorded from Central and Southern Europe.
61. Beris annulipes, Brun.
Beris annulipes, Brunetti, Itec. Ind. Mus. vii, p. 455 (1912).
$ . Head with frons nearly one-fourth the width of the head :
brilliantly shining, dark blue, uniformly wide ; occiput shining
black, with a little grey hair on posterior ocular orbits ; eyes very
shortly but distinctly pubescent ; proboscis reddish yellow. An-
tennae with first two joints black, with bristly hairs ; 3rd joint
yellowish on basal, black on apical half ; a little grev hair about
the base of the antennae. Thorax brilliantly shining violet-blue,
with yellow pubescence on dorsum ; scutellum brilliantly shining
peacock (greenish) blue, with six strong equidistant shining darker
green spines on hind margin. Abdomen violet, moderately shining,
sides with brownish yellow pubescence ; venter darker, with short
yellow hairs. Legs yellow ; tips of anterior femora with an indis-
tinct brown band; hind femora and tibiae with u distinct broad
brown apical band on each ; tarsi marked extensively with brown.
Wings grey, upper part (except costal cell), as far hindwards as
to include the anterior basal and first posterior cells, brown ;
halteres yellow.
Length, 6J mm.
Described from one example taken by me at Darjiling, 7000 ft.,
27. v. 1910 ; in the Indian Museum.
Genus CHORISOPS, Uond.
CJiorisops, Rondani, Dipt. Ital. Prod, i, p. 173 (1856).
Clilontopn, Brauer, Denks. Ak. Wien, xliv, p. 72 (1882).
GENOTYPE, Beris tibialis, Mg. ; by original designation.
Differing from Beris by the palpi being elongate, as long as
proboscis, and the eyes in the d distinctly though rather narrowly
separated. Eyes bare in both sexes. Scutellum with four yellowish
spines. Genitalia in c? very protruding. Hind tibife incrassaled
after the base, especially in d1 . Wings with 3rd vein more shortly
forked ; 4th vein with an abortive 3rd termination ; lower branch
of 5th vein much curved, closing anal cell far before the wing-
border ; stigma very distinct.
Range. Europe, India, Chile.
CHOBI80PS. ALLOGNOSTA. 93
Life-history. The metamorphoses of C. tibialis have been well
described by Handlirsch.*
02. Chorisops tibialis, %.
Beris tibialis, Meigen, Syst. Beschr. ii, p. 3, pi. xii, fig. 8 (1820).
Chorisops tibialis, Brunetti, Rec. Ind. Mus. vii, p. 456 (1912).
Actina hyaUventris, A. Costa, II Giamb. Vico Napoli, ii, p. 455 (1857).
d1 $ . Head : irons shining in tf , blue-black or dark green, one-
eighth width of head at vertex, narrowing to half that width on
lower half; in $ about one-fifth of head at vertex, slightly narrow-
ing downwards, greenish black; vertex with black hairs, frons
with pale pubescence ; face shining white ; proboscis dull orange,
palpi yellowish, porrect, distinct; eyes bare, facets all equal.
Antennas rather longer than head (longer still in $ ), brownish
black, pale below towards base, variable ; 3rd joint 8-annulated.
Thorax and scutellum brilliant shining green, finely punctate,
with fine yellow pubescence ; pleurae with pale pubescence, middle
part of mesopleura bare ; scutellar spines yellow, with a long apical
hair (besides other hairs); humeri yellowish in $. Abdomen
black, moderately shining, more or Jess yellowish on the disc
(especially in the $ ), including generally the greater part of the
3rd and 4th segments ; pubescence short and dark ; pale yellow at
sides down to 4th segment ; venter more yellowish than dorsum ;
genitalia very distinct, and protruding, connected with last abdo-
minal segment by a short neck. Legs yellow to dull orange : tarsi
blackish from tip of 1st joint ; hind femora brownish yellow,
variable ; hind tibiae mainly blackish brown, considerably incras-
sated after the base ; hind tarsi rather short, 1st joint incrassate,
as long as remaining joints together ; pubescence of legs pale
yellow ; 5 with legs paler than in the d . Wings with a faint
brown suffusion ; stigma large, dark brown, sometimes paler in
2 ; halteres yellow, with large knobs.
Length, 6 mm.
The above description is abridged from Verrall.t
Binsar, Kuinaon Distr., W. Himalayas, 28, v. 1912 (Dr. A. D.
Itnms).
Genus ALLOGNOSTA, 0. S.
Alloynosta, Osteu-Sacken, Berl. Ent. Zeits. xxvii, p. 297 (1883).
Metoponia, Loew (nee Macq.), Dipt.-Faun. Siidafr. i, p. 1 (1860).
GENOTYPE, Beris fuscitarsis, Say ; by designation of Coquillett,
1910.
cJ $ . Head transverse, rather flattened, as broad as thorax ;
eyes contiguous in d* ; frons in $ rather prominent, convex (in
assar.iensis), between one-third and one-fourth width of head.
Antennas set at middle of head in profile ; scapal joints subequal,
* Yerh.iool.-bot. Ges. Wieu, xxxiii, p. 243 (1883).
t British Flies, Stratiomyidte, &c., v, p. 214 (1909).
94 STRATIOMTID.S:.
cylindrical, 3rd with eight annulations, 1st the largest, about as
large as 2nd scapal joint, remainder annular, slightly narrowing
in width towards tip. Proboscis and palpi short, latter not extend-
ing beyond epistonia. Thorax subquadrate, corners rather angular,
rather broader behind. Scutellum fully semicircular, convex,
unarmed. Abdomen longer and slightly wider than thorax, ovate,
with tip rather pointed, 8-segmented ; geuitalia small. Leys simple,
moderately long and strong. Wings with normal Berid venation ;
2nd vein rather sharply upturned towards tip ; anterior branch of
3rd vein nearly erect ; only three endings to 4th vein ; discal cell
large, hexagonal ; anterior cross-vein just beyond base, posterior
cross-vein at about the middle ; anal cell large, closed a long way
before border.
Ranye. Europe, India, America.
Ho previous description of the genus appears to have been
given. Loew placed certain new species in what he took to be
Metoponia, Macq., but Osteii-Sacken in pointing out his mistake
merely renamed Metoponia, Lw., as AUognosta, without character-
ising it, as he had no specimens available of the genus at the
time. *
In A. vayans the legs are all black except at the knees; in
A. assamensis the anterior femora and middle tibiae are yellow.
63. Allognosta vagans, Lw. (PI. I, figs. 25, 26.)
Metoponia vayans, Loew, Besch. Europ. Dipt, iii, p. 71 (1873).
Alloynosta inermis, Brunetti, Rec. Ind. Mus. vii, p. 455 (]912).
J . Head with vertex very small, occupied by the three ocelli ;
eyes with uniform, very small facets, bare, contiguous; frons an
equilateral triangle, moderately large, blackish seen from above,
but with grey reflections viewed from other directions ; occiput
blackish. Antennae black, normal, as in Beris ; the 3rd joint
with seven annular impressions, appearing as if composed of eight
closely compressed joints. Palpi and proboscis black; the under-
side of the large labella yellowish. Thorax &\\A scutellum blackish,
with a slight aeneous tinge, microscopically roughened, and with
microscopic yellow hairs ; the anterior margin and shoulders with
;i very little short yellowish hair, which extends also over the
pleurae to some extent. Scutellum unspined, with microscopic
yellow hairs. Abdomen dull black, barely shining, the sides with
very short yellow hairs; venter similar, with very minute yellow
hairs. Genitalia consisting of a pair of blackish, fairly thick
claspers, each with a small narrow appendage ; there being also
an upper pair of yellow, hairy, linger-like appendages. Leys
black, the knees distinctly but not broadly yellowish brown ;
pulvilli pale yellowish. Wings blackish brown ; stigma large,
well defined, brown ; halteres black.
* As to the supposed identity of Metoponia, Macq.. aiid Metoponia, Lw., see
Bruiietti, Rec. Ind. Mus. vii, p. 455.
AlLOGKO&TA. 95
Length, 44 mm.
Described" from a unique r? in the Indian Museum taken by
me at Darjiling, 7000 ft., 29. v. 1910. I found the species not
uncommon at Hankow, China, 22-26. iv. 1906.
64. Allognosta assamensis, sp. nov.
$ . Head as broad as thorax ; frons very retreating, slightly
swollen on upper corners, where it projects above eyes in profile,
forming half width of head, black, shining, parallel-sided, closely
punctulate, and with almost microscopic brownish pubescence ;
lower part smooth, vertical triangle small and low; ocelli incon-
spicuous ; face as broad as frons, sides parallel, shining black, bare ;
two large, dull grey tomentose spots placed diagonally just below
base of antennae, their upper corners nearly touching; proboscis
brownish black, palpi black, rather long ; occiput black; eyes bare,
small, only a little over half height of head ; posterior orbits on
upper half only, rapidly diminishing and disappearing at about
middle of eye. Antennae placed below middle of head in profile ;
first two joints subequal, cylindrical, pale yellow; 3rd with eight
anuulations, 1st annulation broadest and longest; basal part of
fiagelluin brownish yellow, gradually becoming blackish at tip.
Thorax subquadrate, barely wider behind, front corners rather
angular, hind corners cut away by the very distinct brown posterior
calli ; dorsum thickly punctnlate, with very short soft yellowish-
brown pubescence; the moderate-sized semicircular scutellumand
the pleura similarly clothed, black ; a broad yellowish-white stripe
on upper part of latter from anterior margin to wing-base; meta-
notuin small, dull brown. Abdomen 8-segmented, practically
bare ; 1st and 2nd segments almost coalescent, 6th distinctly
narrower than 5th, and the 7th and 8th narrower than 6th ; all
blackish brown, or even slightly violet-tinged; middle half of 1st,
2nd, and 3rd yellowish (the pale part probably variable in extent);
venter mainly yellowish, but blackish at sides and tip. Legs mainly
yellow; apical three-fourths of front tibiae and all front tarsi, last
three joints of middle tarsi, apical half of hind femora, hind tibiae
(except narrowly at base), and last two joints of hind tarsi black.
Wings pale smoke-grey, a very little darker at tip ; stigma large,
black, conspicuous, clearly demarcated and filling all marginal cell
except extreme tip, which is slightly yellowish, as is also the
costa for a very short distance at this point : 1st submarginal cell
practically clear, except at the very narrow base and extreme tip,
thus in striking contrast to the dark contiguous stigma ; extreme
base of wing clear; 5th and 6th veins yellow, remtu'nder dark
brown ; halteres with stem yellowish white, clubs large, oval,
black.
Length barely 5 mm.
Described from a single $ in the Indian Museum from above
Tura, Garo Hills, Assam, 3500-3900 ft., viii. 1917 (Kemp).
96 STRATIOMYIDJ:.
Subfamily XYLOMYIN^.
Head with eyes separated iu both sexex ; face depressed in
middle ; palpi elongate and upturned ; antennal 3rd joint elongate,
with eight annulations and generally a minute apical bristle.
Thorax practically bare, subquadrate ; scutellum unspined. Abdo-
men of 7 to 9 segments, elongate, longer than thorax, with nearly
parallel sides ; genitalia of d conspicuous. Legs moderately long
and stout, bare or nearly so, hind femora often incrassate or
spinose below ; posterior tibiae with two spurs. Wings not of
the normal Stratiomyid type, all the veins reaching wing-margin,
the anterior veins not crowded together ; 2nd vein (praefurca)
originating some distance before base of discal cell, the latter
much more elongate than in any other subfamily ; costal vein
complete, ambient vein present for at least some further distance ;
3rd vein with an elongate fork, lower branch ending at about
wing-tip ; 4th vein with three terminations ; 4th posterior cell
closed near wing-border, a special character of this subfamily ;
five posterior cells ; posterior cross- vein normally present, but in
some species the contact of upper branch of 5th vein with discal
cell is practically punctiform, in which case cross-vein absent ;
alar squama? very small, thoracic squamae absent.
The XYLOMYIN^ resemble the BERINJG more than any other sub-
family, and the principal distinguishing characters are given under
BEHIND. Further characters of the XYLOMYI^^E are the depressed
face, the much more elongate discal cell, the unspined scutellum,
the non-metallic or aeneous nature of the species, and the undilated
metatarsus in the art than at vertex, black, with a little bright yellow pubescence
and some whitish pubescence just above antennae ; face black,
with a little whitish pubescence, more conspicuous near eye-
margius ; occiput black, practically bare ; antennae and palpi
wholly orange-yellow. Thorax black, dorsum and pleura3 with
very short yellow pubescence, minutely punctulate ; humeri and a
ridge thence to wing-base, also the scutellum bright yellow.
Abdomen yellow, punctulate like thorax ; a large dark blackish-
brown spot on each segment, leaving only the sides and hind
margin of each narrowly pale ; genitalia very small ; venter much
as dorsum, but the yellow of gi-eater extent. Legs yellow, extreme
tip of underside of hind femora very dark brown. Winys nearly
clear ; venation normal, veins brown ; halteres yellow.
Length, 4 mm.
Described from several specimens in the Pusa collection from
Chapra, Beiigal (Mackenzie).
66. Xylomyia calopodata, Mg.
Subula calopodata, Bigot, Ann. Soc. Eut. France, (5) ix, p. 195
(1879).
Xylomyia calopodata, Brunetti, Rec. Ind. Mus. i, p. 87 (1907).
Head with frons above antennae one-fourth width of head, a
little narrower at vertex, black, with grey or yellowish grey
pubescence ; ocellar triangle slightly elevated, rather large, black ;
ocelli yellowish ; face apparently dark, with a little whitish
E1 scence ; labella very large, dark reddish brown ; palpi very
and prominent, sharply upturned, hiding all the face, bright
rt' ; occiput dark, grey-dusted, with a friuge of whitish
pubescence, which dies away on upper margin and is longest on
lower part ; antennae dark brown, underside, especially on basal
half, yellowish. Thorax black, slightly shining, closely punctulate,
with minute black bristles and very short yellowish-grey pubes-
cence, which also covers the greater part of the pleurae ; humeri
prominent, bright yellow, with a ridge thence to wing-base ;
scutellum yellowish ; metanotum black. Abdomen black, slightly
shining : closely punctulate, with minute black bristles and soft
short yellow pubescence ; hind margin of each segment except 1st
narrowly but distinctly yellow, bare and shining ; venter similar ;
genitalia rather small. Legs bright yellow, with microscopic pale
pubescence, tips of femora (and for a rather greater distance on
underside) also very narrowly at base, tips of hind tibiae broadly,
anterior tarsi on apical half, and hind tarsi entirely black. Wings
XTLOMTIA. 99
clear, highly iridescent ; venation normal, veins dark brown ;
halteres yellow.
Lenyth, 9 mm.
Described from a single specimen in the Indian Museum, from
Xaini Tal, W. Himalayas, 6000 ft., 5. vi. 1909 ; a second speci-
men from Shillong, 5000 ft. (Fletcher}.
67. Xylomyia nigricornis, sp. nov.
$ . Head with irons gradually widening from vertex to aboVe
antennae, where it is barely one-fifth the width of the head, wholly
covered with yellowish pubescence, except immediately above
antennas, where the pubescence is yellowish white ; vertex dark
grey tomentose, with golden-yellow pubescence ; ocelli not very
conspicuous ; face with nearly parallel sides, fractionally wider
than lower part of frons, black, with greyish or yellowish grey
pubescence : inner margin of eye slightly cut away a little above
antenna?, at a spot where there is a peculiar and conspicuous
shining, black, smooth, elongate, triangular space devoid of facets ;
antennae blackish brown, a little pale about inner and under side
ot 2nd scapal and 1st flagellar joints ; proboscis orange-brown;
palpi bright lemon-yellow, very large and erect, nearly hiding the
face ; occiput grey-tomentose, with some golden-yellow hairs on
margin, and longer ones just behind vertex. Thorax black, thickly
punctulate, with black bristles ; seen from the front only there
are three longitudinal stripes of almost microscopic golden-yellow
pubescence ; humeri conspicuous, bright yellow, a thin dull yellow
line thence to wing-base ; hind corners of dorsum margined with
brown ; pleurae black, closely punctulate, with black bristles and
very short whitish pubescence ; scutellum bright lemon-yellow,
witli lemon-yellow bristles, the base narrowly black ; metanotum
black. Abdomen distinctly 7-segmented, 1st and 2nd scarcely
separable except by the fringe of hairs on hind border of 1st;
1st segment orange-yellow, hind border broadly black : 2nd orange-
yellow, with a large round black spot in middle just clear of the
margins, hind margin vellovvish white ; rest of segments black,
hind margin of 3rd to 6th inclusive yellowish white, clearly demar-
cated ; sides of very small 7th segment yellowish ; all the black
parts of abdomen closely punctulate, with minute black bristles,
the yellow parts with short pale yellow hairs ; genitalia very small,
a pair of small brownish-yellow lamella? ; a little pale, rather long
pubescence at sides of 1st segment, and shorter similar pubescence
on sides of whole abdomen ; venter black, 2nd segment and hind
margins of remainder yellowish. Leys mainly yellow ; hind cox*
(except a patch on hinder side), hind trochanters, underside of
hind femora narrowly, inner and outer sides of same towards base
(the colour almost united on upper side), also extreme tips, black ;
anterior tarsi brownish black ; hind tibiae and tarsi black, except
upper side of former at base. Wings uniformly yellowish grev ;
100 LEPTIDJE.
stigma small, brownish ; 4th posterior cell closed distinctly but
shortly before wing-border ; halteres brownish yellow.
Length, 11 mm.
Described from a unique perfect $ in the Indian Museum, from
above Tura, Garo Hills, Assam, 3900 ft., viii. 1917 (Kemp).
Family LEPTID^E.
Head semicircular, short, rather flattened in front, as broad as
or narrower than thorax ; frons at vertex nearly level with the
eyes ; face short, " usually with a socketed middle part (epistoma)
on which there is usually a rounded raised tubercle " ( Verrall) ;
cheeks more conspicuous in some genera than others. Eyes bare
(except in one non-Indian genus Coznomyia and in Coenomyiodcs,
gen. nov.) ; contiguous or subcontiguous in <3 , distinctly, some-
times widely separated in the $ ; in a few cases the frons of
about equal width in both sexes ; upper facets in d often much
larger than lower ones ; the eyes in life in some species brilliant
green ; three ocelli. Proboscis generally robust, sometimes very
large, though short ; labella prominent ; palpi moderately long,
slender. Antenna? very variable; in the typical LEPTID^ with
3rd joint simple, short-conical, round, or renJform, with a bristle
or style which is usually terminal, but in some genera is dorsal ; *
third joint flagelliform and annulated in XYLOPHAGIX^E. Thorax
a little longer than the head, roughly oval or oblong, with rounded
corners, generally widest at the middle ; scutellum moderately
large, rounded or semicircular ; metanotum larger and more con-
spicuous in XYLOPHAGIN.E than in most LEPTIN^. Abdomen
7-segmented, elongate or conical, longest and nearly cylindrical in
some d XYLOPHAGIN.E. Leys rather long and slender, often
minutely bristly on the tibiae or below the femora ; front coxa)
rather elongate ; fore tibiae with or without a single spur, middle
tibiae always with two spurs, hind tibiae always with one or two
spurs, exceptions being rare ; pulvilli three, empoclium pad-like.
Wings glabrous, but minutely pubescent under high microscopic
power, somewhat broad, with more or less rounded tips. Auxiliary
and 1st longitudinal vein somewhat approximate, always ending
beyond middle of wing ; '2nd longitudinal beginning at from one-
third to one-half the length of the wing, ending not far beyond
tip of 1st, often dipped towards tip to embrace the generally
obvious stigma ; 3rd vein begins a little beyond the 2nd, (almost
always) practically in a line with the base of the discal cell ;
always forked, the fork varying in length f and in width of
* Williston figures the antenna* of Atherix with the arista absolutely basal ;
Verrall evidently claims the most distant point horizontally as the true tip
and considers the 3rd joint produced downwards. Most authors, I believe,
regard as the tip the extremity of its greatest length, as I do here ; it can be
accepted either way, but misunderstandings may arise.
t It is long and narrow in 8tfmphonrmyia,sund short and wide open in
Hilarimorphu, neither of these genera being Oriental.
LEPTI1XE. 101
opening, but always embracing the wing-tip.* Discal cell always
present, except in Hilarimorpha (non-Oriental) ; anterior cross-
vein near basal end of discal cell, always before its middle ;
4th longitudinal with anterior branch forked, lower branch
simple ; posterior cross-vein nearly opposite anterior cross-vein ;
5th longitudinal forked at about its middle, the upper branch more
or less curved or angled at the point where it meets the posterior
cross-vein ; 6th vein complete. Two submarginal cells, 5 posterior
cells; anal cell open or closed; alulae of moderate size, generally
well developed, absent in XYLOPHAGINJE ; alar squamae rather
small, thoracic squamae absent, " but the frenum distinct and
developing a rattier broadened membrane near the angle ; "
halteres distinct, stem rather long.
The whole body in LEPTID^E is usually clothed with soft
pubescence, microscopically so on the often apparently bare legs.
There are no bristles of a chffitotactic nature, though a few short
ones or bristly hairs often occur about the thorax, wing-roots, and
metapleurae.
Life-history. The metamorphoses of many European species
are more or less known.f Larva amphipneustic, with parchment-
like skin, generally living under bark, in moss, or earthy matter,
predaceous on Coleopterous larvae and other soft-wood insects,
though some are aquatic. Twelve-segmented, including head,
which is exserted though small, long, pointed, the mouth-parts
very small ; the three thoracic segments with strongly chitinized
dorsal areas, the prothoracic segment also hardened beneath ; last
segment with a strongly chitinized area on hinder part, with two
strong hooks, this area also bearing two spiracles. Abdominal
segments often with transverse swellings on lower surface, loco-
motion being effected by means of these, which take the form of
roughened intercalated spaces between the segments, though
Brauer affirms that true pseudopods exist in Aiherix.
The perfect insects are found usually in open woods and the
meadow-lands in their vicinity, or in marshy and soft places,
being fond of resting on tree-trunks.
With the exception of one or two North- American biting
species of Symphoromyia, which bite like TABANID^E, the LEPTIDJE
are harmless to mankind. J
The LEPTIDJE are rather closely allied to both the STRATIOMYIDTE
and the TABANID^E, but the former are easily known by the
crowding of the venation on the front part of the wing (except
in BERING) and by the shape of the discal cell, which is always
small, and seldom much longer than wide (except in Xylomyia), at
* In one or two European species of Xylophagus, the lower branch of the
3rd vein may in individual specimens occasionally end barely above the wing-tip.
t Atherix ibis, Xylophagus ater, Leptis tringaria, L. scolopacea, etc.
I [Five species of blood-sucking Leptids have been described by White and
Ferguson from Australia and Tasmania, all belonging to the genus Spaniopsis,
White ; Trichopalpus obscurus, Phil., has been noted as a greedy blood-sucker in
Chile, and Leptis scolopacea, L., and L. strigosa, Mg., have been recorded as
biting man in France. — ED.]
most about one and a half times. In LEPTID^E the discal cell is
distinctly oblong and always more than twice as long as wide.
The only two genera about which doubt might arise would be
Xylophagus and Xylomyia* The TABANID^E, though possessing
the same venation as the LEPTIDTE, may at once be recognized
by the characteristic antennae and their more robust structure
throughout.
Table of Subfamilies.
1. Third antennal joint flagelliform, more or less
elongate, simulated, without arista 2.
Third antennal joint not flagelliform, short, or
but little elongated, rounded, conical, or
reniform, never distinctly annulated ; arista
always present 4.
2. All tibiae with spurs , 3.
Front tibiae without spurs ARTHROCERATIN^E.
3. The 5th posterior cell in broad contact with
discal cell ; face with rounded socketed
epistoma, margined by broad side-cheeks ;
eyes in c? contiguous ; scutellum with two
short blunt spines COENOMYTX.S:.
The 5th posterior cell not touching discal cell ;
face flat, epistoma not socketed or margined
by broad side-cheeks ; eyes in c? well sepa-
rated; scutellum unarmed XYLOPHAGIN^E.
4. Eyes in $ wide apart ; 3rd vein widely forked
at much beyond half its length ; face not at
all socketed VERMII.EONINA:.
Eyes in <$ contiguous ; 3rd vein less widely
forked before, or at latest at, half its length;
face with rounded socketed epistoma LEPTIN^;.
In few families of Diptera is there such diversity of opinion
with regard to the subfamilies as in the present one. Prof. Kertesx,
in his ' Catalogue of Diptera,' admits none at all, but ranks
XYLOPHAGIDJE (including Xylomyia and CCENOMYIJ)^;) as separate
families. Verrall recognises the four last subfamilies presented
here with CHHYSOPILIN^E as a fifth ; Williston divides the LEPTIDJ-
into three (dealing with North-American species only), XYLO-
PHAGINJE (including Ccmomyia], ARTHROCERATIN;E (including
Xylomyia), and LEPTIN.E. Beling has shown the affinity between
Ccenomyia and Xylophagus in their larval stages.
Verrall divides the XYLOPHAQIN^E and VERMILEONIN^E, taken
together, from the C.
Third antennal joint round, oval, or pointed ;
arista alwaj's apical 9.
(i. Arista apical; palpi wholly bare, much en- [p. 122.
larged, spatulate M
Arista dorsal or subdorsal ; palpi slender,
always pubescent, generally bisinuate, more
conical and pointed than in Leptis and
Chrysopilus 7.
7. Eyes in d separated by about the width of
the ocellar tubercle, or actually contiguous
for a short distance ; in $ the frons three
times as wide as the ocellar tubercle .... 8,
104 LEPTIDJE.
Eyes in tf contiguous for a considerable
distance ; frons in $ only as broad as [p. 1 til .
ocellar tubercle .". SURAGINA, Walk.,
8. Frons and side-margins of face (c? $) dis-
tinctly hairy ATHERIX, Mg., p. 120.
Frons in tf and side-margins of face (c? $) [?• 112.
bare ATRICHOPS, Verr.,
9. Anal cell open ; hind tibiae with two spurs . . LEPTIS, F., p. 124.
Anal cell closed ; hind tibiae with one spur [p. 128.
only CHRYSOPILUS, Macq.,
Subfamily XYLOPHAGIN^].
Face flat, no socketed epistome margined by broad side-cheeks.
Palpi long, curved upwards ; the 3rd antenna! joint flagelliform,
generally divided by annulations into seven parts, but in Eliaclii-
cerus into many more. Eyes in c? distinctly separated, in $
rather wider apart. Body and legs almost bare. Scutellum
smaller and metanotum more conspicuous than in normal LEP-
TID^E. Abdomen considerably elongated or cylindrical ; the male
genitalia rather complex, those of the $ long and telescopic. All
tibiae spurred. Wings with auxiliary and 1st veins long, 2nd
shorter than in LEPTINJE, 3rd with fork rather short, widely open,
lower branch ending above or at wing-tip; anterior cross-vein
very short, situated towards base of discal cell ; posterior cross-
vein exceedingly short or absent, anal cell normally closed at wing-
border ; axillary vein indistinct or absent ; alulae absent.
Life-history. " Larva amphipneustic, allied to those of C(ENO-
MYINJE and LETTING ; carnivorous ; living under the bark of dead
trees and preying upon other (probably dead) larvae occurring
there, especially those of wood-boring Coleoptera, or upon the
debris and detritus left by those larvae" (Verrall). Birch, alder,
and pine are the trees mostly favoured. The imagos are usually
found about their breeding-places in woods.
Only one genus of this subfamily (Rhachicerus) occurs in India.
Xylophagus, Xylomyia, and one or two other genera were for
many years regarded as a separate family, but the resemblance
between them is more apparent than real ; and^ Osten-Sacken,
who very closely criticised Brauer's paper on the characters of the
NOTACANTHA (Berl. ent. Zeit. xxvi, p. 363), showed satisfactorily
that they (Xylophagus and Xylomyia) cannot both come in the
same ultimate subdivision.
Genus RHACHICERUS, Walk.
Rhachicerm, Walker, List Dipt. Brit. Mus. v, Supp. i, p. 103 (1854).
? Rhyphomorpha, Walker, Proc. Linn. Soc. Lond. v, p. 275 (1861).
Antidoxion, Snellen v. Vollenhoven, Versl. Meded. Kongl. Akad.
van Wetensch. Afd. Natuurk. xv, p. 1 (1863).
GENOTYPE, Ehachicerus fulvicollis, Walk.; the original species.
" Body slender, nearly linear, cylindrical. Head transverse,
nearly as broad as thorax, vertex narrow. Eyes large, with small
RHACHICERTJS. 105
facets, deeply iiotched and including a smooth space on each side
by the base of the antennae. Ocelli approximate. Proboscis not
prominent ; palpi short. Antennae with 34 or 35 joints, tapering
slightly, much more than half the length of the body, deeply
serrated on each side; joints transverse, very short, cyathiform,
petiolated. Thorax elongate -quad rate ; abdomen pubescent,
8-segmented, hardly tapering from base to tip, about twice as
long as thorax. Legs rather slender, moderately long ; hind
tibiae with very minute spurs ; ungues and onychia very small.
Wings narrow."
Range. North and Central America, Cnbn, India, Malay Archi-
pelago, and one species in Europe.
Life-history unknown.
Table of Species.
Thorax orange nigricornis, sp. n., p. 105.
Thorax black.
Dorsum of first three abdominal segments
mainly yellow bicolor, Brim., p. 105.
Dorsum of abdomen all black, except a
transverse yellow band on 2nd segment, unicincta, sp. n., p. 10G.
68. Rhachicerus bicolor, Brun.
Bhachicerus bicolor, Brunetti, Rec. Ind. Mus. vii, p. 461 (1912).
$ . Head with frons nearly one-third width of head, shining
black, bare ; three small yellow ocelli at vertex ; eyes blackish,
bare ; proboscis bright yellow, palpi of similar colour, very small ;
a whitish triangular spot on each side between the eyes and the
base of the antennae. Antennal 1st and 2nd joints black, 2nd
with a circlet of black and yellow bristles around its tip ; tlagellar
1st joint much wider than the rest, which are blackish brown, the
comb-like prolongations on the underside bright yellow. (The
antennas being incomplete, the exact number of joints cannot be
given.) Thorax and scutellum wholly shining black, practically
bare ; humeri conspicuous, bright yellow. Abdomen black, dorsum
of first three segments mainly yellow, the borders being black, the
4th segment being more or less yellow in the centre ; venter
principally yellow on the first three segments, remainder blackish.
Legs bright yellow, hind coxae black. Wings with venation as in
Xylomyia, grev, with a hardly perceptible darkening from the
costa partly across the wing, and again towards the tip ; halteres
bright yellow.
Length, 8| mm.
Described from one $ in good condition in my collection from
Kandy, v. 1909 (E. E. Green).
69. Rhachicerus nigricornis, sp. nov. (PI. II, fig. 1.)
cJ . Head black ; frons at vertex less than one-fourth width of
head, widening to one-third at level of antennae, with some
whitish dust around base of antennae ; scape of latter yellow, with
short black bristles, flagellum black for 23 joints, with the comb-
106 I.KPTID.E.
like prolongation of each yellowish white, last seven joints
(making with scape 32 in all) with their comb wholly yellowish
white ; proboscis orange-yellow. Thorax and abdomen orange ; a
large black spot on pleurae in front of wing-base and another just
behind wing-base; sternopleura shining black; abdomen with
'2nd segment much the longest ; hind border of each narrowly
black, less distinctly so on 5th and 6th segments. Legs orange-
yellow ; hind coxae shining black ; hind femora and tibiae with
basal two-thirds of each black ; hind tarsi all black. Wings grey,
subcostal and marginal cells blackish ; halteres yellow.
Length, 9 mm.
Described from a unique d in t.he Indian Museum from
Peshoke Spur, Darjiling, 2000 ft., vi. 1916 (L. 0. Hartless).
70. Rhachicerus unicinctus, sp. nov. (PI. II, fig. 2.)
$ . Head with frons brilliantly shining black, longitudinally
furrowed, at vertex one-third of head, slightly wider just above
level of antennae ; face dull black seen from below, but at least
upper part brilliantly white seen from above, narrowing towards
mouth ; proboscis short, thick, bulbous, orange, as are the up-
turned cylindrical palpi, slightly thickening towards tips ; eyes
brown; occiput moderately shining black, with a little short
brown pubescence towards margins. First two joints of antennae
blackish above, orange below ; 3rd with upper side and tips of
rays black, the intermediate part of rays orange-yellow ; 20 or
21 joints present in flagellum, according to whether the small
terminal prolongation is part of the last rayed joint or not: the
flattened irregularly-shaped elevation on which the antennae are
placed lemon-vellow. Thorax shining black, dorsum finely punc-
tate, with extremely short yellowish pubescence ; humeri and
mesopleurae dull shining yellowish ; scutellum like thoracic
dorsum. Abdomen slightly shining black; 1st segment very
short; 2nd longest, with a yellowish transverse band of moderate
width, of which the hind border lies just in front of middle of
segment and is punctuated by a row of small indentations, almost
giving the appearance of two segments; 3rd and 4th segments
barely narrower than 2nd ; 5th rapidly narrowing from base to
tip ; 6th, 7th, and 8th extremely narrow and very elongate, having
a telescopic appearance, the last one possibly forming two segments
(making nine in all) in addition to the two pale yellow elongate
lamellae ; whole abdomen with sparse short yellowish-white
pubescence; venter black, 2nd segment mainly dull yellowish.
Legs mainly orange-yellow ; trochanters, about basal halt of
femora, and hind tibiae wholly black. Wings yellowish grey,
vitreous, stigmatic region a little yellowish brown, a slight infus-
c.'ition at wing-tip ; halteres yellowish.
Length, 8 mm.
Described from one $ in the Indian Museum from Peshoko,
Darjiling, 1000 ft., 26, v.-M, vi. 1916 (Gravely),
DESMOMYIA. 107
Subfamily ARTHROCERATIN^E.
Auteunae obviously elongate, 3rd joint flagelliform, with distinct
annulations, arista absent. Front tibiae without apical spurs.
Only two genera occur in India, and they may be distinguished
as follows : —
First autennal joint as long as 2nd and 3rd
together ; 4th posterior cell open DKSMOMYIA, Brim.
Third antennal joint much more than double
as long as 2nd and 3rd together ; 4th pos- [gen. nov.
terior cell closed . . PARARTHROPKAS,
Genus DESMOMYIA, Brun.
Desmomyia, Brunetti, Rec. Ind. Mus. vii, p. 462 (1912).
GENOTYPE, Desmomyia thereviformis, Brun. ; the original species.
Apparently near, but quite distinct in several characters from
Arthroceras* Five posterior cells, 4th without any trace of
contraction towards wing-border; anal cell closed just before wing-
border ; remainder of venation in accordance with Williston's
figure of the wing of Arthroceras. f Antennae elongate, 1st joint
as long as 2nd and 3rd together, cylindrical, 2nd broadly annular,
3rd elongate with an apical style-like prolongation. Eyes con-
tiguous for a considerable distance, bare ; three ocelli. Proboscis
rather short, stout, cylindrical; palpi elongate, narrow. Face
with two dividing furrows below antennae and rather bulbous
cheeks, as in Arthroceras. General appearance of whole insect
that of a Thereva, with soft pubescence on thorax and abdomen,
the latter with seven distinct segments. Legs slender, like those
of -Thereva • fore tibiae unarmed, posterior tibiae with two distinct
pale spurs each.
71. Desmomyia there viformis, Brun.
Desmomyia thfreriformis, Brunetti, Rec. Ind. Mus. vii, p. 462 (1912).
c? . Head with vertex small, dark ; occiput of a lighter grey
than the dark blackish-grey face; proboscis and palpi blackish
grey ; the whole head with stiff black hairs ; eyes bare. Antennae
dark grey; 1st joint elongate, cylindrical; 2nd broadly annular,
half the length of the 1st; the 3rd joint is best described as
diamond-shaped, with rounded angles, the greatest breadth being
beyond the middle ; the tip produced into an elongate narrow
style with blunted tip, the style as long as the whole antenna.
Tnorax dark grey above, with three separated, moderately broad,
blackish stripes, the outer ones with a tendency to an interruption
in front of the middle; sides and scutellum moderately dark grey,
* The description of this genus is not accessible in India.
t ' North-American Diptera' (3rd ed.), p. 161, fig. 18.
1 08 LEPTIDvE
with soft pale yellowish-white pubescence. Abdomen 7-segmented,
2nd the broadest, dark blackish grey with soft pale yellowish-
white pubescence ; genitalia small, inconspicuous. Legs black ;
knees and femora more or less brownish yellow at base, the
colour on the fore pair extending beyond the middle ; hind tibiao
widened towards tips, hind metatarsus distinctly incrassate ; the
whole of the legs shortly pubescent; anterior tibiae unarmed,
posterior tibiae with two distinct yellowish spines. Wings as in
Leptis, except that the anal cell is closed just before the border,
pale grey ; stigma blackish, very elongate ; two indistinct pale
blackish suffusions, the first, from the origin of the 2nd vein,
embracing the anterior cross-vein, the base of the discal cell, and
the posterior cross-vein ; the second runs from the fork of the
3rd vein to the hinder side of the discal cell, along its outer side ;
costal cell rather broad, the costa curving outwards slightly;
halteres brownish yellow.
Length, 5-6 mm.
Described from two d" c? in the Indian Museum from Badrinath,
Garhwal District, 10,200 ft., 27. v. 1910 (Dr. A. D. Imms).
Genus PARARTHROPEAS, nov.
GENOTYPE, Pararihropeas thereviformis, sp. nov.
Head transverse, as broad as thorax ; eyes bare, with very small
facets of uniform size ( § ) ; proboscis rather short, thick ; palpi
as long as proboscis, cylindrical, straight, bare, porrect ; frons
narrow, with parallel sides ; face broader, with nearly parallel
sides, not projecting beyond eye-margins ; occiput flattened, or
slightly concave ; antennae longer than breadth of head, cylindrical,
barely appreciably narrower towards tip ; 1st and 2nd joints short,
bristly ; 2nd shorter than 1st, as broad as long ; 3rd divided
by eight impressed lines into nice annulations of equal length,
except the 1st, which is considerably the longest; last joint
bluntly pointed, without style. Thorax oblong, humeri promi-
nent, nearly bare, with no obvious bristles, minutely pubescent ;
scutellum transverse, unarmed. Abdomen subcylindrical, about
twice as long as thorax, slightly flattened, with nearly parallel
sides for some distance, gradually narrowing to tip, 7-segmented,
the last segment very short. Legs of moderate size and strength ;
hind femora considerably incrassate, with a row of small blunt
spines on underside ; posterior tibiae with two spurs, front tibiae
apparently unarmed; tarsi as long as tibiae, front pair distinctly
longer ; claws distinct, three pulvilli. Wings with auxiliary vein
ending at about middle of costa, 1st longitudinal ending a little
beyond ; 2nd vein beginning at about middle of wing, ending a
little beyond 1st vein ; 3rd vein beginning shortly after 2nd,
forked at half its length, the branches gradually diverging, the
lower branch ending at wing-tip ; anterior cross-vein near base of
discal cell, which is twice as long as broad ; 5 posterior cells,
1st gradually widening from base to tip, 2nd and 3rd truncate at
£ARABTHROPEAS. 109
base, 4th closed distinctly before wing-margin ; posterior cross-
vein present ; anal cell closed.
c? unknown.
Range. Assam.
This genus differs from Arthropeas, L\v., by (1) the 1st flagellar
joint not being differentiated from the others, except that it is
longer, and the whole antennae boing practically cylindrical
throughout instead of gradually narrowing to tip ; (2) the abdo-
men being much narrower and longer relatively to the thorax ;
(3) the 4th posterior cell being distinctly closed before the border.;
and (4) the hind femora being distinctly incrassated and bearing
a row of small blunt spines on the underside.
Williston, in his table of genera of North-American LEPTID.E,
emphasizes the absence of contact of the 5th posterior cell with
the discal cell, but in Loew's figure of the genotype such contact
is only punctiform. These cells may be more separated in the
other two North-American species ; they are distinctly apart in
Pararthropeas.
72. Pararthropeas thereviformis, sp. nov. (PI. II, figs. 5, 6.)
$ . Head as broad as thorax ; eyes bare, black, facets small, of
uniform size ; frons about one-seventh of the head, almost flush
with eyes, blackish, with short pale pubescence, which is a little
thicker just above antennae ; face not projecting beyond eyes,
flattened, with nearly parallel sides, dark grey, practically bare ;
occiput blackish, bare, except for some short whitish pubescence
behind lower margin of eyes. Proboscis short, thick, not pro-
jecting beyond base of antennae, brown, bearing a few hairs ; palpi
as long as proboscis, bare, brownish yellow. Antennae longer
than breadth of head, cylindrical ; 1st and 2nd joints black,
bristly, 2nd shorter than 1st, as broad as long ; 3rd bare, divided
by seven impressed lines into eight anuulations, of which the 1st
is the longest, tip bluntly pointed; 3rd joint black, except the
brownish-yelloxv 1st annulation and base of 2nd. Thorax oblong;
dorsutn black, rather flattened, with microscopic, depressed, grey-
ish pubescence, which is a little more obvious in front of the
scutellum ; the latter black, with centre and hind margin yellowish,
sparsely and shortly pubescent ; humeri prominent, yellowish
brown ; pleurae black, with very short grey pubescence. Abdomen
subcylindrical, nearly Phycus-like, 7-segmeuted, the first six
subequal, the last very short; black, with minute grey pubescence,
hind margins of segments yellow ; genitalia ending in two small
cylindrical yellowish pieces, which are all that is externally
visible. Legs Therevid-like, comparatively stout and strong ; coxae
black, with minute grey pubescence; fore femora brownish yellow,
with upper side and distal half of lower sides black ; middle pair
black, except narrowly at base and tips ; hind pair black, with a
broad yellow band just before middle ; tibiae black, except that
anterior pairs are very narrowly brownish yellow at base and tips ;
front tarsi distinctly longer than tibiae, posterior pairs subequal to
110 LEPTIDJ«.
tibiae ; all tarsi black, with reddish-brown pubescence on underside
of hind pair; all legs with minute grey pubescence; two obvious
spines oil middle and hind tibiae, arid it is just possible that a small
one may be present on fore pair just .be fore the slightly narrowed
tips, though apparently it is absent. Wings pale grey, veins
dark brown ; halteres brownish yellow.
Length, 10 mm.
Described from a perfect unique $ in the Pusa collection from
Khasi Hills, 1000-3000 ft., iii. 1907.
Type sent to British Museum.
Subfamily CCENOMYIN^E.
Head', antennal 3rd joint nagelliform and annulated, with sty li-
forui tip and no arista. Palpi long, thin, pointed, porrect, 2-jointed.
Eyes hairy or bare, contiguous in cf • Thorax and abdomen
robust, shortly pubescent or bare ; meso- and metapleurae with
tufts of pubescence ; scutellum with two short blunt spines or
unarmed ; genitalia small. Legs comparatively short and strong ;
fore tibiae with one spur, posterior tibiae with two spurs. Basal
wing-cells long ; discal cell present ; two submarginal cells, the
2nd long, with bell-mouthed tip ; five posterior cells, 5th in broad
contact with discal cell.
The CffixoMYis^E and XYLOPHAGIN^E are easily recognised by
the annulated 3rd antennal joint, and the latter are distinguishable
from the former by the four characters previously given in the
table of genera of LEPTIDJE.
Only a few genera and species of CffiNOMYiN^E are known
throughout the world.
Life-history. That of Ccenomyict ferruginea, Scop., a widely
distributed European and North -American species, is known.
The larva is amphipneustic, cylindrical, 12-segtnented ; the head
conical, brown, chitinized ; the rest of the body with chitinized
spots and bands. Pupa free. The species breeds in decaying
poplars and at the roots of trees, especially beeches ; the larvae are
carnivorous and voracious ; their total development may last over
a year.
The adult insects are rather sluggish, frequenting woods.
Genus CCENOMYIODES, gen. iiov.
GENOTYPE, Ccenomyiodes edtvardsi, sp. nov. ;, the only species
known.
Head comparatively small, narrower than thorax ; placed some-
what low down ; occiput flattened ; epistoma separated from
cheeks by an impressed hue ; eyes hairy, separated by moderately
broad frons in $ ( <$ unknown) ; antennae 3-jointed, the 3rd
flagellate, with eight annulations, ending in stylate form, arista
absent ; palpi long, thin, labella broad ; proboscis rather short.
C&NOMYIODES. Ill
Thorax broad, robust ; abdomen similar, rather longer, pointed ;
both shortly pubescent or nearly bare ; scutellum with t\vo blunt
apical spines ; genitalia small. Leys as rioted previously. Winys
as noted above, in rest covering abdomen, posterior cross-vein
absent ; alulae well developed, alar squama small, thoracic squama
very narrow.
c? unknown.
Range. Assam.
This genus differs from Ccenomyia, Latr., by the broad frons in
the 5 , by the anal "cell being closed distinctly before the wing-,
margin, and by the 4th posterior cell also being closed at some
distance from the margin, rounded apically, and sending no veinlet
to the margin.
73. Ccenomyiod.es edwardsi, sp. nov. (PI. II, tigs. 3, 4.)
$ . Head chrome-yellow, a little darker on frons, which bears
a transverse streak just above antennae, not quite reaching eye-
margins ; ocellar triangle black, also a stripe from it passing over
vertex and part of occiput ; mouth-opening pale yellowish ; head
bare, except for some hardly visible pale hairs on occiput, about
the eye-margins. Antennae with 1st and 2nd joints yellow, with
some very short black bristles, giving the joints a partly blackish
appearance ; 3rd joint divided by impressed lines into eight
annulations, brownish yellow, with a black streak on upper side.
Thorax mainly chrome-yellow, although the yellow and black
parts are about equal in extent ; a broad black median stripe
from anterior margin, where it is slightly dilated, to hind margin,
narrowing shortly after passing the suture ; a black stripe on
each side of the median one, leaving a narrow intermediate space,
extending forwards barely to the humeri, and narrowing gradually
behind the suture but reaching hind margin, on which all three
stripes are joined ; sides of thorax yellow, underside black, with
a broad black stripe running from just behind the fore coxae,
curving forwards, narrowing to a point a little below the humerus ;
a similar stripe joins the underside of the thorax with the wing-
base, spreading out hindwards to the hind coxae and upwards
narrowly to the scut ell um, which latter is yellow, the metanotum
being black; dorsum covered with microscopic yellow pubescence,
with a little on parts of the pleurae. Abdomen with first four
segments bright chrome-yellow, with a fairly wide black dorsal
stripe with irregular edges, dilated on hind borders of 1st and
2nd segments, occupying two-thirds of the width of 3rd and 4th
segments ; rest of segments reddish brown, the black dorsal stripe
carried over all but the last one, dilated on hind borders ; 8th
segment canary-yellow on apical half ; a small curved yellow
mark towards each side of 6th segment ; a email indistinct yellow
spot on 7th segment on each side of the middle; venter
reddish brown, a small yellow mark extending over 6th and
7th segments ; genitalia yellowish brown, with two rather short
112
projecting lamellae. Legs practically bare, bright chrome-yellow,
middle coxae obscured ; a short blackish-brown streak at tip of
hind side of middle femora ; hind legs with tip of coxae, extreme
base of femora, and a broad apical band on femora and tibiae black.
Winys yellow, veins brownish black ; halteres bright yellow.
Length, 20 mm.
Described from a single § in the British Museum from the
Khasi Hills, Assam.
Subfamily LEPTIN^E.
Face with a rounded socketed epistoma ; eyes bare, generally
contiguous in the $ , facets of uniform size ; proboscis often
swollen, conspicuous, though never very long; palpi distinct,
moderately long, porrect, or pendant. Antennae with 3rd joint
simple, rounded, oval, or reniform, always with a long apical,
subapical, or subclorsal arista. Abdomen elongate-conical; genitalia
of moderate size. Legs slender ; tibiae with rows of minute
bristles, middle pair with two spurs at tip, hind tibiae with one or
two spurs.* Alulae well developed ; alar squamae of moderate
size, thoracic pair absent.
The genera have already been tabulated on p. 103.
Genus ATRICHOPS, Verr.
Atrichops, Verrall, Brit. Flies, v, p. 291 (1909).
GENOTYPE, Aiherix crassipes, Mg. (Europe) ; by original de-
signation.
Head semicircular, a little narrower than thorax, more rounded
and less tilted forwards than in Atherix • face with a quite bare,
rounded, socketed epistoma ; cheeks large, bare in both sexes ;
t'rous in c? nearly or quite bare, slightly narrowing from vertex to
antennae ; in £ with a little sparse pubescence ; occiput promi-
nent, especially on lower part in c? ; eyes bare in both sexes,
contiguous or practically so in <5 , facets uniform in size ; the
eyes in $ separated distinctly. Proboscis prominent, thick, with
long oval hibella ; palpi two-jointed, prominent, slightly drooping.
Antennae rather short, the space above their base bare; 1st joint
cup-shaped, 2nd more or IPSS rounded, rather short, both bristlv ;
•3rd kidney-shaped, with a dorsal arista.f Thorax slightly arched,
with close short pubescence ; pleurae partly pubescent ; scutellum
* The apparent disagreement here with Verrall, who states that two spurs
are found on all the posterior tibiae, is due to his ranking Chrysopila as a
separate subfamily, in which genus the hind tibise possess only a single spur.
t Verrall speaks of it as "terminal, which from the shape of the 3rd joint
(deepened downwards) appears to be dorsal." This author regards the tip of
the 3rd joint as its furthest point in a straight line, in which case the arista is
terminal, but I am regarding its tip as the extremity of its greatest length, in
which case the arista is dorsal.
ATRICHOPS. 113
normally also pubescent ; metanotum mainly concealed, with bare
sides. Abdomen oblong, narrower than thorax and barely double
as long ; male geuitalia prominent, in $ the tip of the abdomen
pointed. Leys with front pair, and especially front tarsi, longer
and thinner than in Leptis ; front tibia? shorter than, middle pair
equal to, and hind tibiae rather longer than, the corresponding
tarsi ; front coxse rather long ; front tibiae unspurred ; middle
tibiae with two spurs, of which the front one is considerably the
longer; hind tibiae with two nearly equal spurs; the "touch
hairs " beneath the front tarsi obvious. Wings with the auxiliary;
1st and 2nd longitudinal veins ending at or a little beyond middle ;
2nd starting well before the discal cell ; 3rd beginning opposite
base of discal cell, forked at a more or less acute angle rather
soon after quitting anterior cross-vein, the branches gradually
diverging and embracing the wing-tip ; discal cell elongate ; upper
branch of 4th longitudinal vein forked at tip of discal cell ; lower
branch simple ; posterior cross-vein proximad of anterior cross-
vein ; 5th vein forked just before posterior cross-vein, the branches
separating widely, lower branch always curved downwards, meeting
6th vein at or just before wing-border, closing anal cell.
Range. Europe, India.
Life-history unknown.
The genus Atrichops was unfortunately wholly overlooked by
me in describing the species herein referred to it, which were
placed in Aiherix at the time of description. Atrichops is quite
well founded, the species having the cheeks distinctly bare. Set
up by Verrall for Atherix crassipes, Mg., a European species, I
know of no others belonging to it except those I have myself
recorded from the East.
Table of Species.
1. Abdomen with distinct reddish or yellow
markings 2.
Abdomen practically all black, at most
a bluish-grey baud at base or 1st seg-
ment pale yellow 5.
2. Hind femora without distinct band near
tip 3.
Hind femora brownish yellow, with a
broad brown baud before the tip 4.
3. Basal half of abdomen tawny ; wings with
distinct wide median band calojw, Brun., p. 114.
Abdomen mainly black, but reddish tawny
towards sides at base ; wings pale brown,
with indistinct darker band in middle. . . . lanopyga, Brun., p. 115.
4. Sides of thorax yellowish grey posteriorly,
without white stripe ; middle femora
Hack only at base and tip, hind fe-
mora black at t>ase and tip in addition
to broad median baud limbata, O.-S., p. 115.
114 LEPTIDjE.
Sides of thorax all black, with oblique
whitish stripe; middle femora with
basal half black and apical half yellow, [p. 110.
Avith a broad mediau white band intermedia, Brun.,
£. Hind metatarsus not thickened ; apical
half of wing infuscated 6.
Hind metatarsus distinctly thickened;
wing pale, with approximately three [p. 117.
darker bands .metatarsalis, Bruu.,
6. Apical half of wing without narrow pale
band in the infuscated portion ; 4th pos-
terior cell nearly closed ; length, 10 mm. . . ctncta, Brun., p. 118.
Apical half of wing with such a pale band ;
4th posterior cell widely open ; length, [p. 119.
6 mm ccerukscens, Brun.,
74. Atrichops calopa, Brun. (PL II, tig. 7.)
Atherix calopa. Brunetti, Rec. Ind. Mus. ii, p. 420, pi. xii, fig. 2
(1909).
3 $ . Head with vertex black ; frons blackish, with bluish grey
shimmer; proboscis and palpi blackish, the latter hairy below;
antennae brownish yellow, with a trace of bluish grev dust on
upper side of two basal joints ; occiput blackish grey, with a thick
fringe of long whitish-grey hair, which is absent on the upper
part ; lower facets of eye barely smaller than upper ones. Thorax
dark brownish black, moderately shining ; humeri sbining brown ;
sides of thorax brown, dusted with light bluish grey ; scutellum
dark brown, with erect brown hairs. Abdomen with first four
segments mainly yellow, but dark brown on the upper side of 1st.
at the base, and a narrow dark brown dorsal line, with a narrow
line at each side of the abdomen, these three lines extending over
the four segments ; posterior border of 4th segment dark brown,
the colour extending forward in the centre, till it joins the dorsal
stripe ; remainder of abdomen dark shining brown, tbe posterior
border of each segment bearing a little bluish grey dust ; venter
yellowish ; the whole abdomen with moderately long hairs of
similar colours to that of the respective parts on which they are
placed ; genitalia moderately large, dark brown, shining, with
black hairs. Legs with very short black hairs ; coxae dark brownish,
with bluish grey shimmer ; femora yellow, basal third in fore
pair, basal half in middle pair, and basal two-thirds in hind pair
black ; tibiae yellow, knees brown ; basal half of tarsi yellow,
apical half black. Wings very pale grey, apical half a little
darker ; discal cell three times as long as broad ; no distinct
stigma ; a brownish irregular streak from the centre of the fore
border, reaching to about halfway across the wing; halteres
yellow, knobs black ; tegulae blackish with yellow edges.
Length, 8-10 mm.
Described from three <3 3 in fairly good condition, in the
Indian Museum collection. The type is from Sadiya, Assam, and
the others from Tenasserim.
ATEICHOPS. 1 15
75. Atrichops lanopyga, Brun. (PI. II, fig. 8.)
Atherix lanopyga, Brunetti, Rec. Ind. Mus. ii, p. 420, pi. xii, fipr. 3
(1909).
5 . Head with rather more than the upper half of frons velvet-
•black, remainder of frons, the vertex, and face below antennae
blackish, with bluish grey reflections; antennae with two basal
joints brown (third missing) ; eye-facets of uniform size ; proboscis
and palpi blackish, the latter pubescent above and below ; occiput
blackish grey, with a continuous fringe of short grey haiiv
Thorax black, with whitish grey reflections at the sides and on
the humeri ; scutellum black, with black hairs, posterior border
brown. Abdomen mainly black; 1st segment dark brown, the
-colour extending as a wide dorsal stripe across the 2nd ; sides of
first t\vo segments occupied by a large, light yellow, translucid
spot ; 3rd segment velvet-black, with a bicoloured spot at the
sides of the posterior border, the anterior half of this spot being
orange and the posterior half grey, the orange-colour extending
along the sides of the abdomen in a streak, until it reaches the
large translucent side-spots in front of it ; remaining segments
black, whitish grey posteriorly, the sides of 6th and 7th being
light orange-brown ; anal process black ; dorsum of abdomen with
yellowish hair, which at the tip becomes brown ; venter yellow,
with soft yellow hair, the middle segments with some irregular
black marks. Legs : coxae brown, with bluish white shimmer, and
silky white hair ; femora yellow, base of fore pair arid basal half
of middle and hind pairs light brown ; tibiae brownish yellow,
hind pair darker ; tarsi brownish black ; the femora have a little
pale yellow hair, which is brown on the tibiae, bright golden brown
on hind pair, and black on the tarsi. Wings pale grey, with a
faint brown streak from the elongated brown stigma reaching
nearly to the posterior border ; a small elongated clear spot in the
middle of the discal and of the fifth posterior cells ; discal cell
3| times as long as broad; halteres yellow, knobs black; tegulae
pale yellow, transparent.
Length, 10 mm.
Described from one 6" in good condition in the Indian Museum
-collection from Sadiya, Assam.
76. Atrichops ? limbata, O.-S. (PI. II, fig. 10.)
Atherix limbata, Osten-Sacken,Berl. ent. Zeits. xxvi, p. 100 (1882) ;
Brunetti, Kec. laid. Mus. ii, p. 418, pi. xii, tig. 4 (1909).
$ . Head : vertex cinereous grey, with a few blackish hairs,
upper halt' of frons dull coal-black, lower part bluish grey with
•(seen from above) a central small black spot, lower part of face
blackish grey, proboscis brown ; antennae blackish grey with a
few hairs at base, 3rd joint tawny, arista long ; eyes blackish ;
•occiput bluish grey, with a fringe of light hairs, which are longest
i2
116 LEPXIDJ3.
behind the sides of the eyes ; palpi black, hairy below. Thorax
aeneous black above, the colour not attaining anterior margin
except as a wide stripe in the centre, but extending to both
wings and to the scutellum ; humeral calli tawny, with a whitish-
grey tomentose spot, contiguous to each, on the anterior border ;
a bluish grey spot in front of each wing, placed almost on the
dorsum ; sides of thorax yellowish grey posteriorly ; mesopleur8&
bluish grey ; scutellum yellow, base blackish ; metanotum blackish
grey. Abdomen brownish yellow, with a black dorsal stripe on
first three segments, which spreads out over the greater part of
the 4th and 5th segments ; tip of abdomen yellow, sides with a
blackish line and a fringe of black hairs mixed with some paler
ones; venter yellowish; some short pale hairs over the dorsal
surface of the abdomen. Legs : fore coxae yellowish with bluish-
grey reflections, posterior coxae blackish, all of them black at th&
junction with the femora, which are yellow, with the tips narrowly
black, the middle pair having a very small black streak on the
underside near the base, and the hind pair a wide black ring in
the middle ; remainder of legs black, but middle tibiae dark
brownish yellow. Wings pale grey, rather strongly iridescent,
with a rather dark brown middle stripe from the centre of the
costa, narrowing posteriorly and reaching hind margin of wing at
tip of anal cell ; distal part of wing dark grey down to the fifth
posterior cell, the centre of which is pale grey (reaching to th&
border), but a rather wide pale grey space remains between the
dark brown stripe and the distal dark grey part, this clearer part
ceasing at the fifth posterior cell; fifth longitudinal vein dark
brown suffused ; upper transverse vein placed just before one-
third of the discal cell ; halteres yellowish, knob black.
LenytJi, 8 mm.
I describe under the above specific name a $ specimen (now in
the Indian Museum collection) captured from the base of the-
Dawna Hills, Arnherst District, Lower Burma, 1. iii. 1908 (Annan-
dale), which I at first thought was a new species. It differs from
limbata, O.-S., by the presence of the clearer space in the dark
distal part of the wing and by the hind femoral band being in the
middle, not near the tip, other minor differences hardly being^
specific. My augmented description may enable others to identify
it definitely either as limbata or as new.
77. Atrichops intermedia, Snm. (PI. II, figs. 11, 12.)
Atherix intermedia, Brunetti, Rec. Ind. Mus. iii, p. 211 (1909).
c? . This species closely resembles both limbata, O.-S., and
calopa, Bruu., and takes an intermediate position between the
two.
From limbata it is distinguished by an all-black thorax, with a
little whitish shimmer on the humeral parts, round the edges of
the dorsum, and in front of the insertion of the wings ; also by
an oblique whitish stripe on the sides of the thorax, which
otherwise are wholly black, as is also the scutellum. The legs are-
ATKICIIOPS. 117
black ; the fore femora (except base and tip), the apical half of
the middle femora, the middle tibiae wholly, and the hind femora
are yellow, but the latter bear a broad deep black band, occupying
about the middle half of their length. The wing has a large, very
distinct, blackish-brown stigma, whilst the cross-band on the wing
is much darker. The abdomen is differently coloured, closely
resembling that of calopa, except that the posterior half of the
4th and 5th segments is occupied by a whitish band, which is
invisible from certain points of view ; the basal half of the 5th,
and the whole of the 6th, segment is black.
Prom calopa the broad band on the hind femora easily separates
it, this character being consistent in the ten specimens examined.
The wing-marks also distinguish it, owing to the prominent
stigma and the much more distinct apical part.
Length, 8| mm.
Described from ten c? c? in excellent condition taken by Dr.
Annandale, 16. iv. 1909, on Paresnath Hill, West Bengal, 2000 ft.,
on rocks at the edge of a small jungle-stream, where it was
present in large numbers ; Bojiaghat, 12. vi. 1915.
This is evidently a good species, the ten specimens agreeing
exactly in all particulars, whilst the three examples of my calopa
are also consistent ; limbata of Osteu-Sacken, however, according
to him, appears to be variable. The present species is best
described as possessing the abdomen of calopa with the wing of
limbata, with the addition of an all-black thorax and a large black
•stigma.
78. Atrichops metatarsalis, Brun. (PI. II, fig. 14.)
Atherix metatarsalis, Brunetti, Rec. Tnd. Mus. ii, p. 422, pi. xii,
figs. 5, 6 (1909).
dark brown, normal, first joint hairy above only, second joint
hairy above and below ; face below antennae ash-grey ; palpi and
proboscis cinereous grey, with scattered hairs, the former with
some strong black hairs at tip. Thorax blackish, with traces of
close cinereous grey pubescence, which in perfect examples pro-
bably covers all the dors urn and sides ; with also some scattered
longer hairs over the whole thorax. Scutellum blackish, with
scattered yellowish grey hairs and reddish-brown posterior border
carrying a fringe of black and grey hairs. Abdomen shining dark
blackish brown, shoulders prominent, forming tawny-brown calli
with grey hair ; underside of 1st segment continued in front as a
yellow scaly process extending downwards till it reaches the hind
coxaa; 1st and 2nd segments occupied by a pale bluish-grey
transverse band, narrowly interrupted in the middle, extending
over the sides to a considerable extent, and clothed with pale
concolorous hair which is continued along the lower sides of the
whole abdomen to the tip ; dorsum clothed with short black
pubescence which, beginning with the 3rd segment, extends round
the sides until it meets the line of grey side-hairs ; a very narrow
bluish-grey border to 4th and 5th segments, the extreme edge of
5th and 6th segments being orange-brown. Leys blackish brown,.
ATRICIIOPS. 119
minutely pubescent ; apical half of all femora and the whole of
the middle tibiae reddish tawny : some whitish-grey hairs below
femora ; hind tibiae shortly pubescent, with reddish-bronze reflec-
tions. Wings pale grey, apical half pale brownish; no distinct
stigma, but a rather broad ill-defined brown band runs from the
stigmatic portion of the costa towards the posterior border, but
fades away before reaching it ; discal cell three times as long as
broad ; anal cell closed just before the border ; by a strong con-
vergence of the veins the fourth posterior cell (termin, O.-S.) is.
almost closed, thus (if a stable specific character) separating this
species from all other Oriental species in this genus ; halteres
yellowish, knobs black, rather large ; tegulae cinereous grey.
Length, 10 mm.
Described from one £ from Margherita and two $ $ from
the base of the Dawna Hills, Amherst District, Lower Burma,
1. iii. 1908, all in the Indian Museum (Annandale) ; Bojiaghat,
12. vi. 1915 ; Ivathgodam, 16. vi. 1915.
80. Atrichops caerulescens, Brun. (PL II, fig. 13.)
Atherix c&rulescens, Brunetti, Kec. Ind. Mus. vii, p. 463, pi. xxxvii,
fig. 3 (1912).
$ . Head blackish, mainly dusted with bluish grey; frons, from
the ocelli downwards for three-fourths of the distance to the
antennae, shining black, bare, in which colour is inset a deep dull
velvet-black triangle, its base on the lower margin of the shining
part : antennae black, dusted with bluish grey, 3rd joint dull
reddish yellow : mouth-parts dusted with bluish grey. Thorax
blackish, with a shining bluish-grey reflection ; two not very obvious
whitish dorsal stripes, and there are greyish reflections on the
shoulders, posterior corners, and elsewhere, viewed in certain lights;
a few soft black hairs on dorsum ; sides, pleura?, and scutelium
concolorous, a little yellowish behind the thoracic stigma. Abdomen
blackish, barely shining, with short sparse greyish pubescence;
1st segment dusted with bluish grey, 5th and 6th with light
bluish-grey hind margins; venter mainly blackish. Legs black,
minutely pubescent, coxae dusted with bluish grey. Wings very
pule «rey; across the centre a reversed triangular brown spot
reaching from the costa to the hind margin, where its apex meets
the tip of the closed anal cell, the colour filling most of the discal
cell and crossing the tip of the upper basal cell ; following the
discal margin of this brown triangle (at a distance leaving a
moderately wide intermediate clear band) is the proximal margin
of a second large brown band, which fills thence the whole of the
discal part of the wing ; a brown streak in the middle of each
basal cell; stigma dark brown, filling distal half of cell; halteres
yellow, clubs black.
Length, 6 mm.
Described from a perfect $ in the Indian Museum from
Kurseong, 5000 ft., 6. ix. 1909 (Annandale).
120 LEPTID.E.
Genus ATHERIX, Mg.
Atherix, Meigen, Illig. Mag. ii, p. 271 (1803).
Ibisia, Rondani, Dipt. Ital. Prod, i, p. 154 (1856).
Peleckoidocertt, Bigot (p.p.}, Ann. Soc. Ent. France, (2) ii, p. 91
(1856).
GENOTYPE, Atherix ibis, F. ; by general acceptance.*
This genus differs from Atrichops, Verr., chiefly in the pubes-
cent frons and face. In A. ibis, a well-known European species,
both these parts bear dense long pubescence, whereas in Atrichops
they are bare.
Verrall, in describing Atrichops, adds a number of minor charac-
ters which, though illustrative of the type-species (crassipes, Mg.),
may possibly not occur in all the species that it may be advisable
to refer to the genus.
The characters therefore allotted in this work to Atrichops may
be considered to apply to Atherix also.
Only one species is Indian, labiata, Big.f
Range. World-wide.
Life-history. It has been said that the larva has tracheal gills,
whilst Brauer states that it possesses true pseudopods instead of
the roughened intercalated areas generally present in this family.
The $ $ of A. ibis, a European species, cluster on a bough and
lay their eggs all together in a mass, dying on the cluster, the
larvae when hatched falling into the water.
81. Atherix labiata, Big.
Atherix labiata, Bigot, Bull. Soc. Zool. France, xii, p. 117 (1887) ;
Brunetti, Rec. Ind. Mus. iii, p. 212 (1909).
$ . " Proboscis at least as long as height of head ; labella
distinctly enlarged ; palpi bare, a little dilated, dark brown : frons
black, with a few black hairs, face whitish-tomentose. Thorax
and scutellum moderately shining black, the former with grey
reflections, tomentose on anterior part, sides grey-tomentose.
Coxae black; halteres chestnut, with tips blackish. Abdomen
shortly pubescent, grey at the sides ; 1st segment black at base
and middle, with a half band of grey dust on each side ; 2nd
segment dull tawny, with a triangular basal black spot, and at the
middle of each side a brownish transverse half-band ; 3rd segment
with a transverse reddish line in middle. Legs all black, except
base of hind femora which is tinged with pale yellow. Wings
nearly clear at base, posterior half brownish grey, with three
* Coquillett adopts Rhagio diadema, F., as genotype, but Verrall successfully
establishes ibis as such (British Flies, v, p. 285).
t My thanks are due to Mr. Collin for the information that labiata, Big., is
truly an Atherix after an examination of the type, otherwise, as all the species
that have come before me belong to Atrichops, I should have been tempted to
include labiata also in that genus.
SURAGINA. 121
broad contiguous hyaline spots ; stigma oblong, brown ; veins with
brownish int'uscation ; anal cell closed before the border." (Bigot.}.
Ceylon.
Type in Bigot's collection.*
Genus SURAGINA, Walk.
Suragina, Walker, Proc. Linn. Soc. Lond.'iv, p. ] 10 (1860).
GENOTYPE, Suragina illucens, Walk. ; the original species.
$ . "Body linear, moderately broad. Head almost as broad as
the thorax; vertex and front of equal breadth. Proboscis porrect,
compressed, a little shorter than the breadth of the head. Palpi
lanceolate, contiguous to the proboscis. Antennae very short ;
3rd joint round ; arista slender, bare, longer than the antenna;.
Thorax a little narrower in front. Abdomen somewhat flat, less
than twice the length of the thorax, obtuse at the tip. Legs bare,
unarmed, rather long and slender. Wings moderately long and
broad ; radial vein slightly curved ; forks of the cubital vein a
little longer than the preceding part ; 3rd externo-medial vein
inclined beyond the discal areolet towards the 4th, which is
straight ; subanal and anal veins united close to the border ;
discal areolet nearly six times longer than broad, its fore side
hardly angular." ( Walker.)
Range. Ceylon, Celebes, Gilolo.
It is necessary to reproduce Walker's own description of this
genus unaltered, since no example has been available for re-
description.
Osten-Sacken drew attention t to the fact that, in setting up
Suragina, Walker omitted to note its close affinity to Atlierix,
and observes that that author had labelled as a Suragina a species
which Osteu-iSacken himself subsequently described as Atherix
limbata (loc. cit.). Yet Walker certainly knew of Atlierix, a not
uncommon European, even British, genus.
Karsch admits the great affinity between the two genera, but
seems to have successfully differentiated them.J
Only three species are known.
82. Suragina elegans, Karsch.
Suragina elegans, Karsch, Berl. ent. Zeits. xxviii, p. 174 (1884).
m the rather
clearer middle parts ol most of the cells . . 4.
4. Prongs of upper branch of 4th longitudinal
vein issuing from discal cell well sepa-
rated ; grou .d-c<>lour of hind border of
wing of same shade as remainder of wing;
stigma brown, rather ill-defined; length, [p. 131.
3 mm albopictus, Brun.,
Prongs of upper branch of 4th longitudinal
vein issuing from discal cell united or
nearly so ; hind part of wing nearly clear,
except for the infuscatious along the veins ;
stigma nearly blcick, more shai'ply defined ; [p. 132.
length, 4 mm ornatipennis, Brun.,
5. The long dark brown stigma uninter- [p. 133.
rupted luctuosus, Brun.,
Stigma interrupted by an elongate yellow [p. 133.
spot Jlavopunct(itus,Brnn.,*
6. Wii.g with six bluish opalescent spots ; 2nd
posterior cell much longer than 3rd opalescens, sp.n.,p. 134.
Wing without traces of opalescent spots;
2i,d posterior cell subequal to 3rd, at least
in ad my species 7.
7. Mainly black species, at least the thorax . . 8.
Mainly yellowish or brownish species, at
least the thorax almost nlways so 18.
8. Wings with a di-tinct suffusion in addition
to stigma ; (wings pale brown in humeralis
only) 9.
Winers unmarked except for the stigma .... 13. [p. 135.
9. Wings uniformly pale brown humeralis, Brun.,
Wing- never uniformly pale brown ; prac-
tically clear, w;th more or less distinct
suffusions 10.
10. Wing-tip clear, a distinct brownish suffu-
sion from stigma to beyond middle of [p. 13?.
wing segmentatus, Brun.,
Wing-tip always distinctly infuscated .... 11.
K
130 LKPTIDjfi.
11. Wing-tip blacldsh from near stigma up to
and including 3rd posterior cell 12.
Wing-tip suffusion consisting of infuscation [ p. 136.
of both branches of 3rd vein cochinensis, sp. n.,
12. Second and third antenual joints and palpi
black ; abdomen all black ; wing-tip suf-
fusion stronger gravelyi, sp. n., p. 137.
Second and third antennal joints and palpi
yellow ; abdomen yellowish at base ; wing-
tip suffusion less distinct similis, sp. n., p. 138.
13. Antennae black 14.
Antennae yellow 16. [p. 138.
14. Stigma large, dark brown, well-defined. . . . maynipennis, Brun.,
Stigma brownish yellow, diffused or ill-
defined 15.
15. Femora all yellow, except the narrowly
black base ; stigma pale brownish yellow, [p. 138.
ill-defined flavopilosus, sp. n.,
Femora all yellow, except the black basal
half of anterior pairs; stisrma yellowish [p. 137.
brown, suffused . . 1tirmanensisy sp. n.,
16. Legs yellow yerbunji, sp. n., p. 139.
At least basal half of anterior femora [p. 140.
black albobasatis, Brun.,
17. Stigma pale yellow; two shining white
spots above antennae insularts, Sch., p. 140.
Stigma conspicuous, dark, but not always
with well-defined limits 19.
18. Wings distinctly yellow 20.
Wings grey 21.
19. Lower eye-facets barely smaller than upper
ones; tibiae more or less darkened soon [p. 141.
after the base ; length, 8-11 mm ferruginosus, Wied.,
Lower eye-facets very much smaller than
upper ones; tibiae practically all yellowish ;
length, 5| mm unicolor, Brun., p. 141.
20. Third antennal joint black 22.
Third antennal joint yellow 23.
21. Basal half of abdomen yellowish, except for
a dorsal dark stripe and dark hind margins ;
rest of abdomen black ; wings practically
clear, except for a small dark brown
stigma and faint transverse suffusion to-
wards tip; narrow, graceful, nearly linear
species ; anal cell closed at some distance
before border stigma, Brun., p. 142.
Abdomen dark shining brown, hind margins
of 2nd, 3rd, and 4th segments broadly
yellow ; wing distinctly grey, almost yel-
low-tinged ; large brown stigma ; broader,
comparatively shorter species; abdominal
segments after 4th suddenly contracted ;
anal cell closed just before border latus. sp. n., p. 143.
22. First antennal joint black; legs darker brown. sordidus,sp.i\.,p. 143.
Antennae wholly yellow ; legs yellow, tibiae
and tarsi a little darker a#ema mm.
Described originally by Lichtwardt from a type J in his col-
lection, which, purchased from a dealer, bore the label " Ecuador,
iii. 1899." This author subsequently received from the Indian
Museum for examination the c? specimen from which the above
description is drawn up — from the Dawna Hills, Burma, 2000-
3000 ft., 3. iii. 1908 (Annandcde).
Type in Herr Lichtwardt's collection.
114. Hirmoneura opaca, Liclit. (PI. II, fig. 22.)
Hinnoncura o))aca, Lichtwardt, Deut. Ent. Zeits. p. 643, $ (1909) ;
id., Rec. Ind. Mus. ix, p. 334,
(1909) ; id., Rec. Ind. Mus. ix, p. 333, $ (1913).
<$ 2 . A small grey species with abdominal bands. Head
with eyes thickly pubescent, especially in •$ , narrowly separated
in c? ; frons scarcely wider in $ , bearing sparse black hair,
around the antennal base with thicker yellowish-grey hair ;
antennae, palpi, and legs all yellowish red,* tarsi hardly darker.
Thorax blackish, wifh grey pubescence, which at the sides is longer
and more yellowish. Abdomen with basal part of 1st segment
dark brownish, apical part pale slate-grey; rest of segments grey,
with thin yellow pubescence and brown margins ; in the tf the
rows of abdominal pubescence are more erect and prominent,
forming in both sexes on the side-margins almost bunches of
yellow and black hairs intermixed. Wings grey, light brown
anteriorly ; halteres blackish brown.
Length, 11-12 mm.
Described originally from the $ sex only from India; the tf
subsequently described from two specimens from Phagu, Simla
Hills, 9000 ft., 12. v. 1909.
Type ( $ ) in the British Museum, from the Saunders collection ;
of the c? c? referred to, one is in the Indian Museum.
The above description is mainly Lichtwardt's, to which I have
added a note or two from the Indian Museum specimen.
116. Hirmoneura brunnea, Licht.
Hirmoneura brunnea, Lichtwardt, Deut. Ent. Zeits. p. 645, £
(1909).
$ . Near H. coffeala, possibly a seasonal form of it.
In this species the wings are proportionately much shorter, being
only 11 mm. long to a body of 11 mm., plus ovipositor of 2 mm. to
3 mm. Apart from the difference in the proportions, the charac-
ters are mainly as in II. coffeata ; the pubescence on the sides of
the thorax and at the base of the abdomen is yellow, not pale
brown ; legs all yellowish red; wings rather duller in colour, and
the fore border darker.
Described by Lichtwardt from a single $ in the British Museum
from Trincomali, Ceylon, 3. iv. 1892 (Col. Yerbury).
117. Hirmoneura ochracea, Licht.
Hirmoneura ochracea, Lichtwaidt, Deut. Ent. Zeits. p. 643, <^
(1909).
c? . Very near H. austeni. Body pale brown, with short velvety
yellow pubescence, which is longer behind the scutellum and at
the base of the abdomen. Eyes with distinctly yellow hairs,
* According to Lichtwardt ; I call the legs bright yellow, as in H. annandalei.
152 NEMESTBINID^E.
closely contiguous, so that only a very small frons remains, which
latter is clothed with yellow ochre and dark brown hairs inter-
mixed. Antennae brownish ; eye-margins bent, reaching to base
of antennae. Legs wholly yellow ochre. Wings yellowish grey,
fore border yellow ochre.
Length, 18mm.
Described by Lichtwardt from a single 3 from the Tau Plateau,
Burma, 4000 ft., ii. 1890 (Col. C. T. Bingluam).
Type in the British Museum.
118. Hirmoneura austeni, Licht.
Hirmoneura austeni, Lichtwardt, Deut. Ent. Zeits. p. G43, rf (1909).
c? . A large robust species. Body deep dark brown, evenly
clothed with long, erect, woolly, pale brown pubescence, which is
thicker on the sides of the thorax and abdomen, where in certain
lights it has a kind of bronze tinge. Head semicircular; eyes
closely hairy ; frons broad, forming one-sixth the width of the
head with dense deep black pubescence ; epistome a little broader,
black, with concolorous pubescence ; a few light, brown hairs only
about the mouth-opening and around the short proboscis ; antennae
and palpi black ; the latter curving upwards, lying close along the
eye-margins and reaching to base of antennae. Lower surface of
head and sternum with yellowish hair; venter with yellowish
pubescence ; the extreme base of abdomen writh a narrow band of
deep black hairs. Legs stout, reddish yellow: anterior femora
brownish, with dense pale pubescence, which makes them appear
even thicker. Wings brown, anteriorly darker.
Length, 18-20 mm.
Described by Lichtwardt from five 3 <3 in the British Museum
from the Khasi Hills, Assam.
119. Hirmoneura coffeata, Licld.
Hirmoneura coffeata, Lichtwardt, Deut. Ent. Zeits. p. 644, $ (1909).
? . A large species with coffee-hrown body, rather bare, with
some pale brown pubescence only on sides of thorax between the
shoulder and wing-base ; underside of thorax with patches of
pale hairs above the front coxae and under the wing-base ; every-
where else, including the venter, only very thinly pubescent. The
pubescence on frons rather longer, and towards vertex it is
blackish brown, at base of abdomen a narrow band of all black
hairs. Wing surface uniformly coffee-brown, with bronze and
cinnamon-brown reflections ; alulae very reduced so that the wing
is almost clavate; legs paler brown than the body.
Length, 16mm. H- ovipositor 4mm.; wing, 24mm., breadth of
wing, 5| mm.
Described by Lichtwardt from a single $ from Eambodde,
Ceylon (Nietner).
Type in the Berlin Museum.
THICHOPSIDEA. 153
Genus TRICHOPSIDEA, Westw.
Trichoptidea, Westwood, Trans. Ent. Soc. Lond. ii, p. 151, pi. xiv
(183G) ; Maequart, Dipt. Exot. ii, pt. 1, p. 397, pi. iii, fig. 1
(1840);* Wandolleck, Ent. Nach. xxiii, p. 250 (1897).
GENOTYPE, Trichopsidea cestracea, Westw. ; by original desig-
nation.
Of the normal Nemestrinid type ; the species smaller than
those of Atriadopz, with exceptionally long wings, allied to
Triclioplitlialma and Meyistorliynclius. Three ocelli on a small
conical tubercle ; epistome (Estrid-like, mouth-parts very reduced
and indistinct, mouth itself very rudimentary ; abdominal seg-
ments puffed up ; genitalia in male clubbed ; $ unknown. First
and Just tarsal joints each as long as the other three together.
Auxiliary and 1st longitudinal veins apparently united until
beyond middle of wing, the former then turning up to costa rather
suddenly ; upper branch of 3rd vein turned up suddenly, meeting
2nd vein about its middle; both branches of 4th vein apparently
united at tip of discal cell, upper branch eitlier single or, if forked,
the lower prong may consist also of the tips of both the lower
branch of the 4th vein and upper branch of the 5th. If this
reading of the veins be correct there is a somewhat long posterior
cross-vein. Anal cell open, axillary vein and squamae absent.
llanye. Except dolirni from the Andamans and Sumatra, the only
other known species (cestracea, Westwood) is from Australasia.
120. Trichopsidea dohrni, Wand.
Trichopsidea dohrni, Wandolleck, Ent. Xacli. xxiii, p. 251, figs. 7, 8
(1897) ; Liclitwardt, Deut. Ent. Zeits. p. 648 (1909).
3 . Ground-colour reddish brown, with not very reddish-
yellow hairs. Head rather broader than thoi-ax, shining golden
brown ; eyes contiguous below vertex, upper facets much larger
than lower ones; three yellowish-brown ocelli on prominent
vertical tubercle, with black and reddish-brown hairs : epistome
(Estrid-like, with thick reddish-brown pubescence ; antennae
3- jointed, 3rd joint rod-shaped, with undulating outline; at tip
with some very short fine pellucid hairs ; mouth-opening indistinct.
Thorax brown, with fine velvet-like tomentum, showing several
darker and lighter indistinct longitudinal stripes ; in front and at
sides with long thick reddish-yellow pubescence ; scutellum pale
brown, anterior margin darker ; squama3 absent ; halteres greenish
yellow. Wings pale yellowish brown, with six clear-cut hyaline
spots, which in certain lights show a pearly lustre; 1st longi-
tudinal vein present ; anal vein absent. Abdomen yellow, with
velvet tomentum; 1st segment pale yellowish, 2nd with a pale
•* Macquart's figure shows the broad elongale-conical labrum ; the longer,
concave, round-tipped labium ; and the long filamentous organs, all as
described by Westwood, who assumed the latter to be "analogous to palpi"
and stated that the mouth was rudimentary and concealed. Wandolleck also
describes the mouth as rudimentary.
354 NEMESTRINID-S.
band interrupted in middle and with two pale side-spots ; these
spots occur on each succeeding segment also, and are visible, with
the interrupted band, through absence of tomentum ; venter pale
yellowish red, with a darker mark at sides of each segment. Legs
with coxa?, trochanters, and femora of both anterior pairs yellowish
red, with moderately long concolorous pubescence ; tibiae fawn-
yellow with reddish-brown tips ; hind femora long, thin basally,
thence distinctly clubbed; the thin part fawn-coloured, the
clubbed part reddish yellow ; last tarsal joint and pulvilli reddish
yellow, the other joints fawn-colour with darker tips ; claws
dark brown.
Length, 10mm ; wing, 11'4 mm.
Described by Wandolleck originally from a type tf from
Sumatra, and Lichtwardt subsequently recorded another 3 from
Boss I., Andaman Is., v. 1904 (Col. C. T. Bingham}.
Type in Herr Dohrn's collection at Stettin ; the second speci-
men in the British Museum.
Genus ATRIADOPS, Wand.
Atriadops, Wandolleck, Ent. Xach. xxiii, p. 245 (1897).
Colax, Wiedemann (nee Hiibner, 181G), Analec. Ent. p. 18, pi. i,
fig. 8 (1824); id., Auss. Zweifl. ii, p. 260, pi. ix, fig. 11 (1830) ;
Macquart, Dipt. Exot. ii. pt. 1, p. 34, pi. iii, fig. 2 (1840);
Westwood, Cab. Orient. Entom. p. 38, pi. xviii, fig. 5 (1847) -r
Walker, Ins. Saund., Dipt. pi. v, fig. 4 (1850).
GENOTYPE, Colaxjavana, Wd., or C. macula, "Wd.*
Moderate-sized species with peculiar tomentose-like pubescence
and dark-coloured and spotted wings. Epistome (Estrid-like ; eyes
in cf subcontiguous for a short space ; ocelli absent ; proboscis
not. obvious, if projecting from mouth-opening, hidden in the long
hair of the face ; antennae 3-jointed, much as in Hirmoneura.
Thorax subquadrate, rather broader than long ; abdominal seg-
ments somewhat puckered-up just before the hind margins, forming
a blunt ridge ; genitalia in c? short, tip with a broader base, in £
longer with sabre-shaped lamella. Hind femora slightly thicker ;
first and last tarsal joints each as long as remaining three together.
Wings with auxiliary vein short, ending free at about opposite
origin of 4th vein ; 1st longitudinal and greater portion of vein
" diagonal " considerably thickened ; 3rd vein forked ; 4th vein
forked, anastomosed as usual for some distance with 3rd ; lower
prong of upper branch not always reaching wing-margin ; f anal
cell open, axillary vein present; thoracic squama? well developed ;
two submarginal and four posterior cells, 1st sometimes divided
* Both originally described together on the same page. I cannot per-
sonally decide and have seen no species set up as genotype.
f It does not do so in the one example of A. javana that I have seen, but
Waiidolleck figures it running to the wing-margin, with also a short cross-vein
between it and the vein in front (Ent. Nacb. xxiii, p. 246, fig. 1, 1897).
AT1UADOPS. — CEYLONIA. 155
by a cross-vein, 3rd closed ; discal cell sometimes narrowly
open ; * anal cell open.
Range. The very few species known come from China, Brazil,
Ceylon, Sumatra, Java, and South and West Africa.
121. Atriadops javana, Wied.
Colax javana, Wiedemann, Analec. Ent. p. 18 (1824) ; id., Auss.
Zweifl. ii, p. 261 (1830).
Atriadops javanaj Walker, Ins. Saund., Dipt. pi. v, fig. 4 (1850) ;
Wandolleck, Eut. Nach. xxiii, p. 246, fig. 1 (1897).
c? . Head with eyes contiguous for a short space ; frons pale
brownish grey, with greyish-white hairs ; antennae pale brownish
yellow. Thorax (?) blackish (discoloured, probably brownish in life),
sides with longer whitish-grey hairs. Abdomen brighter brown,
each segment towards hind border drawn up slightly, forming a
slight ridge ; anterior half of each segmeiit with fine golden-grey
pubescence, which towards sides becomes longer, thicker, and
whiter, and laterally on the hind margin of each segment is some
long black pubescence. Wings dark brown, the darker parts
blackish, hind margin rather broadly greyish, except at extreme
wing-tip, at tips of lower prong of upper branch of 4th vein, tips
of both branches of 5th vein, two or three intermediate spots
situated rather close together, and at tips of anal and axillary
veins, at each of which is a small blackish-brown spot ; similar
dark spots occur on the distal side of the diagonal veins, at the
juncture of the veins, and larger ones at the forking of both
4th and 5th veins ; at the distal end of the 2nd basal cell is an
ill-defined clearer space, and a similar subtriangular one extending
from the 3rd vein just before it meets the 4th, which, inclining
distally and diagonally, nearly reaches the costa. Thoracic
squamse considerably developed, with long greyish fringe.
Length, 8 mm.
Ee- described from a single c? in the Indian Museum from
Trincomali, Ceylon, 10. ix. 1909 (Col. Yerbury}, presented by
Herr Lichtwardt, the species having been originally described
from Java.
Genus CEYLONIA, Licht.
Ceylonia, Lichtwardt, Deut. Ent. Zeits. p. 646 (1909).
GENOTYPE, Ceylonia magnified. Licht. ; by original designation.
Body robust, linear. Proboscis hidden by the long hair of the
face; palpi small, thin; "j" epistome pointed, as in Trichopsidea,
* In Wandolleck's figure of A. macula, Wd. (/. c. p. 247, fig. 2), a Brazilian
species, the discal cell is prolonged narrowly to the wing-margin, remaining
open through the failure of the lower branch of the 4th vein to meet the lower
prong of the upper branch. The species seems abnormal, as the 2nd longi-
tudinal rein is either absent or the 3rd vein must be unforked. Also in the
basal angle of the last posterior cell is a very small quadrate additional cell.
t I follow Lichtwardt in sfating that the face has long hair, but I have not
noted this fact in describing my nivea. In any case in my type of that species
the palpi are perfectly obvious.
156 NEMESTRINID^.
Dicrotrypana, and Symmictus. • Eyes in J contiguous for some
distance, upper facets larger than lower ones ; vertex very small,
tliree ocelli on small prominent tubercle. Antennae short, of normal
Nemestrinid type ; 1st joint cylindrical, 2ud half; as long as 1st,
3rd globular, with apical bristle longer than the whole antenna.
Wings with auxiliary and 1st longitudinal veins verv long, straight,
close together, ending at a little distance before wing-tip ;
2nd vein beginning at half the wing's length, straight ; 3rd begin-
ning immediately afterwards, projected diagonally downwards,
anastomosing as usual with 4th vein, forking shortly beyond;
4th vein with upper branch forked, the branches of 3rd and
4th veins all parallel with 2nd vein and with hind margin of
wing; all ending above wing-tip, except the lowest, which ends
exactly there ; lower branch joining upper one just before forking
of latter; 5th vein rather peculiarly forked, so that the shape of
the last posterior cell at base is that of a slightly curved cone ;
posterior cross- vein absent; anal cell open, axillary vein long.
Two submarginal and five posterior cells, 4th closed.
122. Ceylonia magnifica, Licht. (PI. II, fig. 23.)
Ciylonia magnifica, Lichtwardt, Deut. Ent. Zeits. p. 646, rf (1909).
Atriadops n'ivea, Brimetti, Rec. Ind.fMus. vii, p. 477, tf (1912).
J . Head blackish grey ; palpi thin, small, of same colour ; eyes
brown, very large, contiguous for half the distance from the very
small vertex (bearing tliree small ocelli) to base of antenna?, the
lirst two joints of which are brown (3rd joint missing); underside
of head with some grey hairs. Thorax thickly clothed with pale
yellowish-grey hairs, which become white on the sides. Abdomen
thickly clothed with pale yellowish-white hairs, which, seen in
front, appear silvery white ; venter dark grey, with grey hairs.
Legs dark reddish brown. Winys clear, narrowlv dark brown at
base; also a narrow, irregular but clearly-cut, stripe across the
middle from the costa nearlv to the hind border; a second wider
similar band towards tip, which reaches the hind margin, spreading
over both sides of the "diagonal" vein, and connected along the
costa with a small apical spot, and proximally less distinctly with
the median band, leaving a well-marked clear oblong spot between
the 1st and 2nd longitudinal veins ; a small brown spot on wing-
margin at tip of 5th longitudinal vein and a larger one at tip
of 6th.
Length, 12 mm.
The species was described originally by Lichtwardt from a single
J from Pundaluoya, Ceylon (Green}, in the British Museum.
At the time of describing my nivea I had overlooked this author's
paper on Oriental NEMESTRIXID.T:, and the description given
here is that of the type of my species from a single S in mv
collection from Haldumulla, Ceylon (Green").
157
Family CYRTID^E.
Head very small, subglobular, almost wholly consisting of the
enormous eyes, which tire contiguous in both sexes ; vertex very
small ; frons generally very small. Proboscis varying from very
thin and elongate (MS in some BOMBYLIID.E) to very short or
obsolete; palpi generally obsolete. Antennae approximate at
base, placed in some genera well above, in others well below the
middle line of the -head, 3-jointed, variable in shape, 1st joint
sometimes minute.
Thorax greatly humped ; prothorax sometimes (PHILOPOTIN^E,
non-Oriental) enormously developed ; pubescence moderately
thick, sometimes furry ; scutellum lai-ge, without bristles ; rneta-
uotum generally concealed by it.
Abdomen very broad and gibbous, often quite globular and
pellucid, normally 5-segmented ; pubescence variable ; genitalia
inconspicuous.
Legs simple, comparatively short, rarely with a short blunt spur
on middle tibiae, or with one developed and one less developed spin-
on posterior tibiae, or a circlet of very short spines on front tibiae :
3 pulvilli; empodium as pad-like as outer pulvilli; claws long.
The entire body is wholly devoid of spines or strong bristles.
Wings diverging and deflexed when at rest, longer in § than in
J ; venation very characteristic, but a general interpretation of
its varied forms would be out of place here, as only four genera
are Oriental. Ambient vein at times quite absent, even above
the wing-tip ; auxiliary and 1st veins long, 2nd vein often absent;
praefurca beginning about opposite the base of the discal cell ;
anterior cross-vein near base of discal cell, sometimes very short
or absent; 3rd longitudinal vein generally forked ; posterior cross-
vein present or absent, 4th vein with manifold ramifications ; 5th
vein generally forked; anal vein present, sometimes weak and, with
the axillary vein, possibly absent. Technically two submarginal
cells and from three (sometimes apparently only two, owing to
the dividing vein being obsolete) to five posterior cells. The
venation in this family is, perhaps, not thoroughly understood
yet.* Thoracic squamae enormously developed and very promi-
nent, wrinkled, and generally pubescent, their size being one of
the chief characters of the family.
Life-history. The metamorphoses of several species are knownf,
the larvae being parasitic in the egg-cocoons or the abdomens of
spiders. The larva is amphiueustic (though Konig describes that
of Oncodes as metapneustic), short, thick, 12-segmented, with very
* See Verrall, Brit. Flies, v, p. 447 et seq.
t At-tomclla linden it, Ericlis., Brauer, Verh. zool.-hot. Ges. Wien, xix, p. 737,
pi. xiii, figs. l-(i (1869), and in Denks. Ak. \Viss. Wien. xlvii, pi. v, figs. 89-92
(1883). Oncodes pallipes, Lntr., Menge, Schrift. Danzig. Natur. Ges. i, p. 37
(18C>6); Oncodes fumatiis, Erichs., Brauer, Verb, ssool.-bot. Ges. Wien, xix,
p. 737 (186'J). Henops brit,
List, xx, p. 100, pi. x(1888).
158 CYIlTIDjE.
small head and small mouth-parts ; the posterior spiracles accom-
panied by peculiar large plates. Pupa free, with a median longi-
tudinal row of spines on dorsum of thorax. Lundbeck writes
of one species : — " The larva was lying in the abdomen of a Cteniza
ariana with the terminal spiracles in one of the lungs of the
Cteniza. Before pupation the larva left the now dead Arachnid
and transformed in the nest." Stein and Konig state that the
eggs of some species are deposited on branches of trees, also that
the young larvte possess considerable leaping powers.
The globular shape of the CIBTIDJE with the very large
squamae and characteristic venation stamp the members of this
family with a facies peculiarly their own, except for a few short,
hump-backed BOMBYLIID^, but from these the three (not two)
pulvilli and the venation at once separate them.
This family is by some authors called the ACUOCEIUD^, but the
name is " absolutely contradictory to most of the genera "
(Verrall). The term CYRTLD.E was used as far back as 1841, the
genus Gyrtus antedates Acrocera, and the family name is sup-
ported by Verrall, Osten-Sacken, and others.*
The range of the family is world-wide, but only about two
hundred species are known.
Table of Subfamilies.
Prothoracic plates enormously developed, united
dorsally, forming a. large shield between meso-
notum and neck , PHiLOPOTiN^E,p.l58.
Prothoracic plates not conspicuously developed.
Third antennal joint long and strap-shaped ;
no trace of style or apical bristle PANOPIN^:, p. 160.
Third antennal joint generally short ; apical
bristle or stiff hairs always present .... CYETIN^:, p. 163.
There are different views as to the location of some of the
genera, as to their synonymy and their limits. I follow Verrall
in keeping Pterodontia in CYRTIN^E.
Subfamily PHILOPOTIN^l.
The principal character is the enormous development of the
prothoracic plates. Only one species is Oriental.
Genus PHILOPOTA, Wied.
Philopota, "Wiedemann, Ausser. Zweifl. ii, p. 17 (1830).
GENOTYPE, Philopota conica, Wied. (Brazil).
Head much puffed out behind eyes; three ocelli t; eyes con-
* The family has been called by various other names by different authors;
INFLATE, for instance, and ONCODID*, Kertesz even retaining this latter name
to the present day.
t According to Wiedemann ; the lower ocellus in my species is not discernible,
but it is impossible to state definitely that it is not present.
PHILOPOTA. 159
tiguous, pubescent ; frons slightly prominent on lower part ;
antennae elongate, porrect, approximate and bulbiform at base,
situated immediately above base of proboscis, 3rd joint with apical
bristle. Proboscis very long and thin, projected below the body
to a considerable distance. Thorax with prothoracic lobes enor-
mously developed, prominent, their inner angles touching one
another. Abdomen conical (conica, Wied., type) or rounded
(viridcenea). Legs of moderate length and size, of the normal
Cyrtid type. Wings with auxiliary and 1st longitudinal veins
long, nearly parallel, prsefurca beginning just before middle of
wing; 2nd longitudinal vein long, ending much above wing-tip;
3rd vein simple, dying away a long distance from border of wring ;
anterior cross-vein present ; 4th vein simple, nearly straight,
almost attaining border; 5th vein simple, twice bent towards tip;
anal vein long, reaching wing-border.*
Range. South Europe, Asia Minor, India, Japan, Mexico, and
South America.
The above is an attempt at an augmented description of the
genus, as Wiedemann's was so short.t When additional species
are known a further description may be given.
123. Philopota viridsenea, sp. nov.
Head much extended behind the eyes, the surface shining aeneous
green, closely punctured, covered with rather dense, moderately
short, vellow pubescence ; eyes touching from just below the
small vertex bearing the ocelli,]; and with moderate! v short, not,
dense, dark brown pubescence, facets of uniform size; frons of
moderate size, flush with eyes at apex, but rather prominent on
lower part, dark shining blackish brown, more or less yellow on
lower margin; antennae elongate oval, dark shining brown, with
long, yellow, translucent stvle; cheeks blackish grev ; the large,
smooth, oval, shining black base of the proboscis beginning just
below the antenna? ; the rest of the proboscis bright yellow, bnre,
in length reaching under the body to tip of abdomen, the tip
widely bifid. Thorax with the entire surface of dorsum and sides,
and of scutellum, shining aeneous green, closely punctured, covered
with very short yellow pubescence. Prothoracic lobes very large
and prominent, subt ri angular, their inner angles contiguous, the
upper angles tipped with yellow. Abdomen aeneous green, closely
punctured, with very short yellow pubescence, which on the last
two segments and towards sides of previous segment is changed
to greyish ; extreme side-margins of most of the segments
* Verrall suggests a different reading of the veins, calling them in order,
auxiliary, 1st and 3rd longitudinals, and 4th vein (forked, the anterior cross-
vein being therefore absent).
t " Antenna; porrect, approximate, small, inserted above base of proboscis,
bulbiform, bristles apical. Proboscis elongate, deflexed before thorax. Eyes
contiguous, hairy. Three ocelli. Abdomen conical."
} Only the two upper ones are obvious; there should be three.
160
narrowly brownish yellow ; venter dull aeneous, with short whitish
pubescence ; geuitalia concealed, somewhat yellowish. Legs black-
ish with short grey pubescence ; femora black, extreme base and
nearly the apical halt' yellow ; rest of legs wholly yellow ; ail legs
with very short inconspicuous pale pubescence ; the three yellow
pulvilli and the black claws distinct. Wings yellowish grey,
considerably iridescent and rippled ; veins in anterior part of
wing black and distinct, the remainder yellowish and less distinct.
Thoracic squamae enormous, grey, with short greyish pubescence
on disc arid edge, sufficiently transparent for the brownish-yellow
halteres concealed below them to be seen.
Described from a perfect unique from Dungagali, 8000 ft.,
Hazara District, 21-22. v. 1915 (T. B. Fletcher).
Type sent to British Museum.
Subfamily PANOPIN^E.
Little can be added to the characters given in the table. The
basal and discal cells are generally well defined.
Genus PIALEA, EricJis.
Pialea, Erichson, Entomographica, i, p. 160 (1840).
GENOTYPE, Pialea lomata, Erichs. ; by original designation.
Body of the normal Cyrtid type. Head very small, mostly
occupied by the densely pubescent, contiguous eyes ; occiput
flattened, three ocelli ; proboscis very short, no obvious palpi.
Antennae deflected, placed on the underside of a distinct tubercle
in the middle of face between the eyes ; 1st and 2nd joints short,
bristly, 3rd very large, elongate, leaf-shaped with obtuse tip, much
longer than two basal joints together. Thoracic squamae enor-
mously developed. Abdomen thick, oblong, conical, longer than
thorax, densely pubescent, 6-segmented. Legs of normal Cyrtid
type, tibiae ending in a small blunt tooth-like projection on inner
aide. Wings of moderate size, slightly elongate ; costa ending at
tip of lower branch of 3rd vein, immediately before wing-tip ;
auxiliary vein ends at from two-thirds of wing (aurifrila) to nearly
end of wing (lutescens) ; 1st longitudinal vein very long; 2nd
begins at one-fourth of the wing, straight or gently curved towards
tip, ending just beyond 1st; 3rd begins very near origin of 2nd,
widely forked towards tip, nearly straight, with lower branch
ending below wing-tip (lutescens), or bent upwards, both branches
ending before wing-tip (aurtpt&s); anterior cross-vein very near
origin of 3rd vein, at basal end of the elongate discal cell; 4th
vein with apparently both branches forked, upper prong of upper
branch closing the 1st posterior cell and giving the appearance of
another discal cell in front of and parallel with the true one. In
lutescens upper prong of lower branch throwing off a veiulet
PIALEA. 161
closing discal cell (or it may be regarded as a discal cross-vein) ;
in auripila, if such veinlet is a discal cross-vein, the 4th vein's
lower branch is not forked, joining upper branch of 5th a little
before margin of wing. Posterior cross-vein short; 5th vein
forked, lower branch closing anal cell ; axillary vein present (at
least in auripila). Three submarginal cells ; three or four posterior
cells (first and last closed).*
Range. The only other known species (P. lomata) is from
Brazil. " .
Life-history unknown.
124. Pialea auripila, Brun. (PL II, figs. 24, 25.)
Pialea auripila, Brunetti, Rec. lud. Mus. vii, p. 472 (1912).
cJ . Head about half the full height of the thorax ; eyes closely
touching from the extremely small vertex to the base of the
antennae, rather densely covered with long brownish-yellow hair;
vertical triangle with three ocelli, slightly elevated; occiput much
flattened, with brownish-yellow hairs ; proboscis very short, sub-
conical, blackish, with some yellow hairs at tip; palpi not obvious ;
frontal triangle black, very small, projecting over basal joints of
antennae, which are short, a little broader at tip, blackish, with
black bristles; the third joint of the shape of an elongated leaf,
flattened, with obtuse tips, three times as long as the two basal
joints together, yellowish, with dark brown lower margins.
Tliorax higli, robust, nearly quadrate; ground-colour dark shining
brown, closely covered with rather long thick bright golden-yellow
hair. Scutellum and metanotum concolorous in ground-colour and
hair with the dorsum ; sides of thorax dull yellowish, with long
thick golden-yellow hair; a little brownish hair is intermixed here
and there with the yellow hair of the thorax. Abdomen robust,
thickened, longer than thorax, oblongo-conical, tip pointed.
Brownish yellow; 1st segment, very short; the dorsum of each
segment, mainly occupied by a large semi-circular dark brown
spot at base, and extending in the middle of the segment to the
hind margin, leaving the sides of the segments mainly yellow;
5th and 6th segments all black, the latter very small. Venter
blackish brown, practically bare, emargination of segments
narrowly yellow. The whole upper side of the abdomen with
bright golden-yellow hair, which is brownish towards the tip.
Legs with coxae half as long as femora, yellowish, with pale yellow
hairs. Tibiae (which are a little longer than the femora) and tarsi
black, with blackish-brown hairs ; underside of tibiae brownish
yellow, and ernargi nations of the tarsal joints below narrowly
yellowish ; tibiae a little broadened at tips, where they end in a
small blunt tooth-like projection on the outer side. Tarsi a little
longer than tibiae, base of metatarsus (which is equal in length to
* My reading of the venation in my description of P. auripila was slightly
faulty, the present interpretation being more probably correct.
If
162 CYBTIOS:.
the rest of the tarsus) pale yellow. Winys barely as long as
abdomen, rather broad, tip rounded, pale grey ; costal vein very
distinct, ending at tip of lower branch of 3rd vein, immediately
before the extreme tip of the wing; auxiliary (subcostal) and 1st
longitudinal veins thick, the former ending at about two-thirds
of the wing, the latter a little beyond it, both nearly straight,
distinctly separate (though approximate) from just beyond the
humeral cross-vein ; the 2nd and 3rd longitudinals with the
anterior cross-vein have the appearance of springing from a
common stem (or else the 3rd longitudinal and the anterior cross-
vein spring simultaneously from the 2nd vein at a little beyond its
base) ; 2nd vein nearly straight, but curved upward at its tip, to
thecosta; 3rd vein nearly straight, and in a straight line with
the praefurca, forking widely towards its tip, and becoming
suddenly much thinner, both branches ending in the costa
distinctly before the wing-tip; anterior cross-vein short, near
base of discal cell; 4th longitudinal vein forking just before
anterior cross-vein ; upper branch nearly straight for more than
half its length, thence with a slight curve, forked, the upper
Fig. 10.--Pialea auripila. Brim., Fig. ll.—Pialca auripila, Brun.,
antenna. wing.
prong erect, closing 1st posterior cell, lower prong running to
wing-margin ; lower branch of 4th vein also comparatively
straight, forked so widely towards its tip that the prongs are
nearly in the same straight line with one another, upper one
closing discal cell, lower one joining upper branch of 5th vein
towards its tip, closing 3rd posterior cell. Posterior cross-vein
very short, placed at about middle of discal cell ; 5th vein
thickened, forked near its junction with posterior cross-vein ;
lower branch closing anal cell. First basal cell rather large and
broad, bounded distally by the anterior cross-vein ; 2nd basal
cell much longer and narrower, bounded distally by the posterior
cross-vein; discal cell elongate, about 3^ times as long as its
greatest breadth. The 1st submarginal cell nearly as long as the
marginal and narrowed at wing-margin; 2nd submarginal cell
small, subtriangular. The 1st posterior cell embraces the wing-
tip, but the upper prong of the upper branch of the 4th vein
divides the cell at two-thirds of its length; 2nd posterior cell
large, of irregular shape; 3rd closed, much wider distally; 4th
subtriangular, formed by the branches of the 5th vein ; anal vein
slightly curved ; axillary vein short, axillary cell large. Alulae
LAS1A. 163
very small ; squamae pale yellowish, the upper ones small, the
lower ones very large and pubescent. Halteres yellowish.
Length, 12 mm.
Described from a single perfect <$ in the Indian Museum from
Kurseong, 6000 ft., 26. viii. 1909 (D'Alreu).
This species is undoubtedly a Pialea, though it offers a slight
variation from the venation of the type species,* and the shape
of the abdomen is different. In my remark after my original
description of auripiki. about Westwood's figure of the antennae,
I overlooked his statement in his description of the tubercle on
the face and the fact that the antennae are placed on the under-
side of it. This fact further convinces me that my species is
correctly placed in Pialea.
Subfamily CYRTIN.E.
Prothorax not forming a dorsal shield in front of mesonotum.
Third anteunal joint normally short, with apical arista or hair-
like rays.
Three genera are Oriental, and are easily distinguished as
follows : —
An additional cell in wing above the discal and
about as long LASIA, Wd., p. 163.
No such cell.
Costa with an angular projection at tip of 1st
longitudinal vein; antenual tip with three fp. 165.
stiff' hairs, or a style bearing three such hairs. PTERODONTIA, Gray,
Costa without such projection ; antennae ending [p. 168.
in a long arista ONCODKS, Latr.,
Genus LASIA, Wiedj
Lasia, Wiedemann, Analec. Entom. p. 11 (1824); id., Ausser. Zweitl.
i, p. 329 (1828).
Mesophysa, Macquavt, Dipt. Exot. i, pt. 2, p. 166 (1838).
Feriexutemma,aiyot) Ann. Soc. Ent. France, (3) iv, p. 65 (1856).
Verticistemma, Bigot, op. cit. vii, p. 200 (1859).
GENOTYPE, Lasia splendens, "Wied. (Brazil).
Body of the usual Cyrtid type. Head with eyes bare, con-
tiguous in both sexes, facets very small, of uniform, size ; no
ocelli ; proboscis from a little shorter to a little longer than the
body, narrow, cylindrical ; palpi apparently absent or obsolete.
Antennae 3-jointed, porrect, approximate at base, diverging; 1st
joint cylindrical, short ; 2nd broader than 1st, wider towards the
* P. lomata, Erichs. (as P. lutescens, Westw., sp. nov.) Trans. Ent. Soc.
Lond. (1876) pi. vi. This is the only other known species, and is from Brazil.
t Kertesz in his catalogue ranks Ijasia as synonymous with Panops, Lam.
(Ann. Mus. d'Hist. Nat. iii, p. 263, 1804). Nearly all the species have been
described under Lasia, and, as Verrall accepts it without comment, I follow
him. It is a moot point if Lasia should belong to the PANOPIN*; or OYETINJK,
M2
164 CYRTIDjE.
bluntly truncate tip; 3rd elongate lance-shaped, bare, shining,
produced into a long style, which is longer than the entire
antenna. Tlioracc and abdomen densely pubescent. Leys of
moderate length, simple. Wings with auxiliary vein ending
about middle of eosta ; 1st longitudinal ending towards wing-tip,
nearly straight; 2nd starting soon after humeral cross- vein,
nearly parallel with 1st, both slightly upturned at tips; 3rd
beginning immediately beyond base of 2nd, descending in a
straight ine diagonally across the wing, forking abruptly beyond
its middle, the two branches curving upwards, practically parallel,
ending above wing-tip; 2nd stibuiarginal cell truncate at base;
anterior cross-vein just beyond origin of 3rd vein, very near base
of discal cell, short ; upper branch of 4th longitudinal vein
forked at tip of discal cell, at which point a short transverse vein
connects with the forking of the 3rd vein; lower branch of 4th
vein simple, suddenly recurrent towards tip, ending jointly with
upper branch of 5th vein at about middle of hind border of wing ;
di^cal cross-vein long, closing discal cell ; 5th vein forked at half
its length ; upper branch meeting 4th vein at middle of discal
cell ; lower branch closing anal cell well before margin ; posterior
cross-vein absent, axillary vein foreshortened. Two submarginal
cells ; five posterior cells, 1st divided by a transverse vein,* 4th
closed. Squama) very large.
llniige. Europe, India, Australia, North and South America.
Life-Tit 'story apparently unknown.
The present description is built up mainly on L. aurata, as the
genus does not appear to have been diagnosed since Wiedemann
founded it. Some of the characters herein given may require
modification, as no other species is available for comparison.
125. Lasia aurata, sp. nov.
Send with vertical triangle black, roughened, elevated, with
long chrome-yellow, rather drooping, bristly hairs; eyes black,
facets very small, of uniform size, with no trace of any transverse
channels ncross the discs; frontal triangle elevated, black, shining,
bare. Antennal 1st and 2nd joints bright yellow, short; 2nd
* In his notes on the genus Lasia (Brit. Flies, v, p. 450) Verrall distinctly
states that the extra cell "is a second portion of the upper basal cell and not a
basal part of the 1st posterior cell," and he compares the venation with that
cf the NEMESTRINID./E. but I venture to think that the anterior cross-vein (also
recognised by Verrall as such) cannot be pluced in the biisal cell, but must,
when present, invariably divide the 1st basal from the 1st posterior cell. It
seems to me that the short, cross-vein connecting the 3rd and 4th veins in
Lasia and one or two other genera in this family (Eulonchus, P(ernpe.nts,
etc.), represents the point at which in the NEMESTRTNID.B the 3rd and 4th veins
anastomose, which in that family occurs in contact with the discal cell and
not distinctly beyond it as in CYKTID*. Even if in NEMESTRINID/E this
usually punctiform contact is replaced by a short cross-vein, it should be
regarded as identical with the same extra cross-vein in CYETID^. and
not as the anterior cross-vein, which would, therefore, be wholly absent in
NKMESTKIKID*.
LA.SIA. — PTEIIODONTIA. 165
subcylindrical, narrower at base, tip almost rounded; 3rd shining
chestnut-brown, base yellowish, lance-shaped, the tip drawn out
into a long concolorous style, which is about as long as the whole
antenna; 2nd joint with yellow hairs. Face, except immediately
below antennae, deeply sunken between the eyes, shining bla«-k,
bare, narrower on upper part; occiput grey, covered along the
margin apparently with long bristly bright yellow scale-like
pubescence.* Thorax moderately shining black on disc, with
apparently a slight aeneous tinge, the dark colour extending
forward in a broad b'and to anterior margin ; rest of dorsum
bright yellow, including shoulders, humeri, and mesopleurae ;
ground-colour of scutellum brownish yellow, that of remainder
of the pleurae grey or yellowish grey ; whole thoracic dorsum,
scutellum, and pleurae covered with dense long bright yellow
bristly scale-like pubescence, which becomes a little paler over-
shoulders and lower part of sides of thorax, and more orange over
scutellum and on hinder part of thoracic dorsum. Abdomen
yell.nvish brown, lighter on 1st segment and deeper on 3rd and
4th; a'l segments with a well-defined lemon-yellow hind border
of moderate width, broadest and least well-defined on 1st, and 5th
segments, on which latter it occupies more than half the surface.
"Whole dorsal surface covered with moderately short light brown
pubescence, and towards tip some microscopic pale yellow depressed
hairs. Venter mainly yellowish, with a blackish median band on
hinder part and some transverse black marks on 3rd and 4th and
following segments ; genitalia inconspicuous, hidden. Legs * holly
lemon-yellow with short soft concolorous pubescence. Wings
practically clear to base, distinctly ribbed; veins brown ; thoracic
hquuuae pain yellow, with long bright yellow fringe.
Described from a single specimen in the British Museum,
labelled simply "Ind."
Genus PTERODONTIA, Gray.
Pterodont.ia, Gray in Griffith's Anira. Kingd. xv, p. 779 (1832).
Plerethntin, Macquart, Hist. Nat. Dipt, ii, p. 514 (1835).
GBNOTYPE, Pterodmtia ftnvipcs, Gray ; by original designation.
Body of the normal Cyrtid type, pubescent. Head extremely
small, nearly wholly occupied by the eyes, which are very large,
contiguous in both sexes, densely pubescent; three ocelli; mouth-
opening more or less concealed; proboscis short. Antennae
situated at bottom of h -ad, 3-jointed; 1st cylindrical, 2nd short,
3rd conical, with (according to Macquart) a long bare style, but
in P. orsurn of abdomen without white scale-
spots 10.
9. Larger species, 14 mm. ; aatennal style [p. 200.
normal maculiventris, sp. n.,
Smaller species, 8 mm. ; antennal style as
long as 3rd joint stylata, sp. u., p. 201.
10. Clear part of wing with several isolated dark
spots insulata, Walk., p. 203.
Clear part of wing without any isolated
dark spots puerula, sp. n., p. 205.
11. Scutellum and antennae black; all the
abdominal segments with a band of yel- [p. 206.
lowish white pubescence benyalensis, Macq.,
Scutellum and antennae reddish yellow or
brown ; the 2nd abdominal segment with
basal band of whitish pubescence* .... brahma, Sch., p. 206.
12. Length 20 mm niveiventris, Brun.,
Length 7-10 mm 13. [p. 207.
13. Antennae reddish brown ; pubescence on
anterior margin of thorax whitish ; scu-
tellum all black; wing wholly clear;
venter with small snow-white scales at [p. 209.
bases of segments , vitripennis, Brun.,
Antennae black ; thorax with bright yellow
thick scale-like pubescence; scutellum
* After Schiuer; E. bengalensis and brahma may possibly be synonymous.
192 BOJIBYLIIDJE.
reddish brown ; wing clear, costal and
subcostal cells pale yellowish grey;
venter with soft snow-white hairs in
middle vit>'ea> BiS-» P- 21°-
Antenna} black, ferruginous at base;
"thorax with tawny hairs"; scutellum
and thorax ferruginous ; dorsum of abdo-
men with fringe of pale yellow hairs on
fore border of each segment, and " more
completely clothed on the underside " ;
wings a little greyish, slightly tawny at [p. 211.
base and on fore border basifascia, Walk.,
143. Exoprosopa flavipennis, Bnm.
Exoprosopaflampennis, Brunetti, Rec. Ind. Mus. ii, p. 465 (1909).
2 . Head with frons and ocelli as in flammea ; frons orange,
with close short golden-orange hairs ; mouth-border yellow :
proboscis brown, shorter than the head; eyes reddish brown,
bare, with small, uniform facets ; back of head reddish orange,
bare, considerably prominent, with a narrow fringe of very short
bright yellow hairs on the margin of the cavity behind. Antennae
orange; first two joints sub-cylindrical, wider at tip, with yellow
hairs ; 2nd shorter than 1st ; 3rd joint twice as long as first two
together, bare, the upper side quite straight, the underside rapidly
narrowing from base to tip, a very distinct cylindrical style at
tip. Thorax ferruginous orange, with black dorsum and blackish-
grey underside ; a fringe of long yellow hair-like scales on
anterior margin ; a bunch of fiery orange-red bristly scale-like
hairs on shoulders, and shorter similar ones on the humeral calli ;
the dorsum clothed with moderately close, short orange hairs ;
several long fiery orange-red bristles, directed backwards on the
posterior calli ; underside nearly bare, a few blackish hairs here
and there. Scutellum orange-red, covered with short, similarly
coloured pubescence ; posterior margin with a horizontal row of
concolorous strong bristles. Abdomen sub-conical ; ground-colour
black, apparently with short, close, black pubescence. A bunch
of elongate orange-red scales at the shoulders of the 1st segment ;
sides of 2nd segment dull reddish ; venter black, unmarked ;
genital apparatus in one specimen, orange, subspherical, slightly
protruding, and apparently twisted to the left. Leys with coxa?
and femora dark brown ; anterior tibiae brownish yellow, hind
pair black ; all the knees pale ; tarsi black ; fore legs practically
bare, posterior ones minutely pubescent and with distinct black
bristles, which are stronger on the hind pair; posterior tarsi
closely pubescent, with black bristles .on underside. Wings with
first posterior cell closed some distance before the border ; rather
more than the proximal half bright orange-yellow, apical part and
a narrow border along the posterior margin almost to the base,
quite clear; an intermediate dark brown, ill-defined cross-band,
commencing at the upper edge of the marginal cell opposite tip
EXOPEOSOPA. 193
of auxiliary rein, the width of the baud being about one-third of
the length of the 1st submargiual cell, across the middle of which
it passes ; continuing across the middle of the closed portion of
the 1st posterior cell, the distal half of the discal cell, and the
bases of the 2nd and 3rd posterior cells, after which it is narrowed
to a streak and turning, extends towards the base of the wing,
separating the yellow part from the clear margin. Alulae with
fringe of dirty grey elongate scales ; alar squamae reddish orange,
with a short dense fringe of concolorous elongate scales.
Length, 14-18 mm.
Described from two specimens from Pusa, Bengal. One (type)
in the Pusa collection, taken 19. iv. 1907; the other in the Indian
Museum collection, taken 25. v. 1906, now partly greasy.
E. flammea, Brun., and flavipennis, Brun., are distinguished
from all the other Eastern Exoprosopa known to me by the 1st
posterior cell being closed at some distance before the border, the
4th longitudinal vein meeting the 3rd before the origin of the
anterior branch of the latter (\i\flammea\ or just below it (in
flavipennis). They therefore belong to the group for which
llondani established the genus Argyrospyla (emended by Verrall
to Argyrospila), but which the late Baron Osten-Sacken says
(Biol. Cent.- Am., Dipt, i, p. 78) cannot be sustained, this character
in various species showing all stages between a closed and open
cell, sometimes even in the same species.
144. Exoprosopa flammea, Brun.
Exoprosopa Jiammea, Brunetti, Rec. Ind. Mus. ii, p. 466 (1909).
$ . Head with entire frons and face orange-yellow, covered
with short golden-yellow hairs ; lemon-yellow round the mouth,
•\yjth short bright yellow hairs; frons at level of antenna} one-
third the width of the head, narrowing towards vertex: ocelli
small, close together in front of the vertex; antennae missing,
except 1st joint which is reddish ; proboscis brown, shorter than
length of head ; eyes dark reddish brown, facets very small,
uniform ; occiput brick-red, rather prominent, with sparse short
golden-yellow hairs. Thorax cinereous ; humeral and posterior
calli orange-tawny, the latter bearing five or six concolorous
spines pointing backwards; the anterior border and shoulders
are covered with thick long fiery reddish-yellow bristle-like scales,
which are also abundant below the shoulders and behind the
wings ; the dorsum (slightly denuded) is evidently lightly clothed
with short orange-yellow hairs ; underside of thorax cinereous,
with a moderate amount of orange-yellow hair. Scutellutn red-
dish brown, the posterior margin bearing a row of concolorous
strong bristles directed backwards ; dorsum with sparse yellowish
hairs. Abdomen sub-conical ; ground-colour chiefly reddish brown ;
2nd segment with a narrow anterior and rather wider posterior
baud connected by a thin dorsal stripe ; a rather wide irregular,
o
194 BOMBYLIID^.
not well-defined black transverse band on 3rd segment, which is-
repeated less distinctly on the following t\vo or three segments.
The extreme posterior border of all the segments is reddish
brown, and the whole dorsum is covered lightly with short,
bright orange hairs, which become brightest, longest, and fiery red
at the abdominal tip ; a bunch of fiery-orange elongate scales at
sides of 1st segment. Venter cinereous, covered with short orange
hairs; posterior borders of all the segments orange-yellow.
Legs with coxa3, femora, and most of the tibiae reddish orange ;
the tibiae towards the tips (especially the hind pair) and all the
tarsi black ; the middle femora have a few black short spines
below, the hind pair a row of stronger ones ; the posterior tibiae
are beset with short black bristles (longer on the hind pair) and
have a circlet of strong black spines at the tip, and the whole
legs, especially the tibiae and tarsi, are minutely but densely
spiuose. Wings clear ; first posterior cell closed at some distance
before the border, the 4th longitudinal vein joining the 3rd half-
way between fork of latter^ and the wing-border. Two broad
dark brown bands ; base of wing bright orange-yellow, the colour
extending across the wing from the costa to (and including) the
alula, and reaching distally to just within the two basal cells ; the
costal cell orange-yellowish, also a spot on and over the discal
cross-vein, and another small spot in the centre of the upper
basal cell. The first brown band begins in the upper basal cell,
which it fills, extending posteriorly, filling the 2nd basal cell
and basal fourth of discal cell, thence narrowing to the hind
border of the wing, leaving about the distal fourth of both the
anal and axillary cells clear ; the second band begins approximately
on the costa, blending with the orange colour of the costal cell*
fills three-fourths of the 1st submarginal and 1st posterior cells>
thence narrowing somewhat, it fills nearly the distal half of the
discal cell and terminates distinctly deal of the posterior margin
of wing, entering the 2nd and 3rd posterior cells ; a small round
quite clear spot is in the extreme upper angle of the lower basal
cell. A few very short stiff black bristles at the base of the
costa, and some short close yellowish-orange hairs on the margin
of the alulae and the orange tegulae ; halteres yellow.
Length, 18 inm.
Described from a single 5 (now headless) in the Indian Museum
collection, taken at Pusa, Bengal, 17. iv. 1907 ; other specimens
are from Pusa, l.v. 1911, at light (Hoivlett) ; Pusa, 3.iv. 3914;
Trincomali, Ceylon, 25. vi. 1891 (Col. Yerbun/).
This species closely resembles E. Zar, K, in the shape of the
wing-markings.
145. Exoprosopa lar, F. (PL III, fig. 3.)
Bibio lar, Fabricius, Sp. Ins. ii, p. 414 (1781).
Anthrax lar, Wiedenmnn, Auss. Zweifl. i, p. 268 (1828).
Anthrax collaris, Wiedemann, Auss. Zweifl. i, p. 271 (1828)
Walker, List Dipt. Brit. Mus. ii, p. 247, 2 (1849).
EXOPSOSOPA. 195
Litorhynchm collaris, Macquart, Dipt. Exot. ii, 1, p. 80 (1840).
Anthrax ruyicollis, Saunders, Tr. Eiit. Soc. Lond iii n 59 iil v
tig. 5 (1841).
Exoprosopa binotata, Macquart, Dipt. Exot. Supp. 5, p. 89 (1855)
Exoprosvpa collaris, Brunetti, Rec. Ind. Mus. ii, p. 444 (1909).
c? . Head with f rons distinctly narrower at vertex than above
antennae, where it equals one-third the width of the head; dark
nut-brown, with a slight pink or orange tint, covered with short
black pubescence, and a few minute elongate pale brownish -yellow
scales or scale-like hairs on lower part; face of similar colour to
frons, with black pubescence and similar scales ; proboscis dark,
withdrawn ; anteunal 1st and 2nd joints ferruginous, 3rd black •
occiput yellowish brown, with minute black bristles, which con-
tinue over the vertex nearly to the ocelli; and with whitish
scales forming a band of irregular width contiguous to the eve-
margins. Thorax black, with sparse black pubescence, covered
with very short depressed yellowish (almost pinkish) brown
scale-like hairs ; anterior margin with a thick fringe of very long
narrow brownish-yellow (varying to bright ferruginous) hair-like
scales; a bunch of similar scales below the humeri, on prothorax,
and below posterior corners of dorsum : coarse hairs of similar
colour cover the mesopleurae, with some black hairs intermixed
below wing base; sides of thorax dull orange-brown; humeral
calli reddish brown, with^numerous black bristles ; posterior calli
reddish brown, covered with pubescence like that of the thorax,
with several long strong spines projected hindwards. Scutellutn
uniformly dark reddish brown, with similar pubescence to the
thoracic dorsum ; hind margin with a row of spines. Abdomen
black, the sides often more or less reddish ; posterior margin of
1st segment narrowly, 2nd and 3rd segments from base to tip
(covering one-third of the surface on each side) dull but distinct
reddish brown ; the whole dorsum with short depressed blac-k
pubescence ; a fan-like bunch of rather dirty white very elongate
scales at sides of 1st segment ; a patch of long yellowish-white
scales towards each side of 3rd segment, contiguous to the
margin, and 6th segment (and possibly 7th also) covered with
similar scales ; the sides of the abdomen from 2nd segment to tip
with coarse black bristly hair. Venter moderately dark brown,
barely tinged with vellowish, but with paler hind margins to the
segments and a few soft yellowish hairs ; genitalia rather largo,
ferruginous. Legs reddish brown, metatarsi more or less darker,
rest of tarsi black ; fore legs, apart from the usual minute
pubescence, with rows of very short bristles on tibiae ; middle
femora with at least two rather strong bristles of unequal length
on inner side towards tip ; hind femora with a row of stronger
bristles below, posterior tibiffi with the usual rows of small
bristles; tarsi with short stiff black pubescence below. Wings
best described as pale grey with the basal third dark brown,
filling the anal and axillary cells except their tips, the colour
continued along the costa nearly to the tip of the 1st longitudinal
c>2
196 BOMBYLIID.E.
vein and extending bindwards to the upper side of the discal cell,
thence spreading out and embracing the apical third of that cell,
the base of: the 2nd posterior, and just entering the 3rd posterior,
the colour sharply delimited distally by a line drawn from its
costal limit to where it ends posteriorly in the 3rd posterior cell ;
.a small clear spot at upper corner of 2nd basal cell, with a small
yellowish spot immediately above it, and a slight pale yellowish
tinge about the middle of the 1st posterior cell ; the brown
colour is only very slightly variable in its extent. Alulze and
alar squamae with a fringe of dirty brown scales ; halteres black.
Length, 14-16 mm.
Bedescribed mainly from a 3 in good condition in the Indian
Museum from Bara Banki, United Provinces, 14. x. 1910, and
others iu less good condition from Kangra Vallev or Sikkim
(Dudgeon] ; Allahabad, i. 1910 (Chatterjee) ; Bandhara^ Cent. Prov.,
India, l.xii.1912 (Imms); Sahibganj, Bengal; Bellary, South India,
10. viii.-ll. ix. 1913 ; Surat, Bombay, 11. xi. 1911 (Fletcher);
Bangalore ; Trivandrum ; Mahagany, Ceylon, 15. ii. 1891 ; Nila-
velli, Ueylon, 30. v. 1891 (Col. Yerbury).
Types in Fabricius's, Westermaun's and Wiedemann's collections.
It must be remembered that the foregoing description is chiefly
from an individual specimen. Wiedemaun noted the close affinity
between collaris and lar, claiming for the former the more fox-red
collar, the absence of white tomentum at the tip of the abdomen,
the absence of white pubescence towards the sides at the base of
the abdomen, the wholly black ground-colour of the body, the
black-haired abdomen, and the shortening of the second trans-
verse black band (so to speak) on the wing, so that it does not
reach the hind border.
Most of these characters are more or less variable, especially
the distance to which the more distal of the two extensions of the
costal dark border reaches, and the reddish-brown colour of the
sides of the abdomen. Saunders's figure of his rugicollis shows
the limits of variation in three characters, the ferruginous collar,
the shortened wing-band, and the all-black abdomen. Wiede-
mann's characters of the absence of white vestiture at the base
and tip of the abdomen are negative ones, and it may reasonably
be supposed that his specimen was somewhat worn.*
Two specimens in the British Museum from Mahagany, Ceylon
{Col. Yerbury), appear to represent an undescribed species allied
to E. lar, but are not in sufficiently good condition to describe
properly.
One of the specimens bears a label : " ? E. binotata, Macq."
This hitter I concluded t to be synonymous with collaris, Wied.,J
beinu; further convinced by a specimen of the latter in the Indian
* As the nrst few examples of this species seen by me answered fully to
Wiedemann's collaris, a certain number of specimens were identified by me as
such and returned to correspondents. These should bear the name of lar.
t Eec. Ind. Mus. ii, p. 444.
\ E. collaris, Wied., is now regarded by me as synonymous with lar, F.
EXOPEOSOPA. 197
Museum, labelled " E. bimaculata, Macq." in Bigot's handwriting.
Bigot evidently wrote bimaculata in mistake for binotata, but the
two specimens described above are certainly distinct from lar. In
the event of binotata, Macq., being a good species, they may be
representatives of it.
146. Exoprosopa pennipes, Wied.
AntJtra.r pennipes. Wiedemann, Dipt. Exot. i, p. 129 (1821) : id..
Auss. Zweifl. i, p. 272 (1828).
Exoprosiipa pennipes, Macquart, Dipt. Exot. ii, 1, p. 49 (1840);
Bruuetti, Kec. Ind. Mus. ii. p. 443 (1909).
Hyperalonia pennipes, Rondairi, Ann. Mus. den. vii, p. 452 (1875).
Head with frons more than twice as wide above antennae (where
it equals one-third the width of the head) as at vert ex, blackish, with
black hairs on upper part, and below with black hairs mixed with a
number of short silvery scales of rather greater width ; face black,
with black bristly hairs and small silver scales ; proboscis yellowish
brown ; antennae black, possibly sometimes reddish brown at base ;
occiput black, with small silvery scales at least near the eye-
margins, probably a considerable part of the surface thus covered
in perfect specimens. Thorax black ; dorsum with a dark indigo-
blue tinge, caused bv the whole surface being covered in perfect
specimens with brilliant small subquadrate blue scales, longi-
tudinally striated, black pubescent ; anterior margin with long
narrow black scales ; bunches of similar scales or bristly hairs
below shoulders, on mesopleurae and metapleurae, and below hind
corners of dorsum ; apparently some black bristly or coarse hairs
around the wing-base. Scutellum black, covered with scales like
those of the thorax, pubescent, dull dark brown towards hind
margin, which bears a row of spines. Abdomen black, covered
with blue-green or green scales, with black pubescence; some
long, thin, very dark brown scales in a bunch at sides of 1st seg-
ment ; venter dark blackish indigo, with a little black pubescence
and no trace of scales. Legs black; fore tibiae and metatarsi
yellowish brown ; middle tibiae very dark brown ; hind femora
towards tips and hind tibiae from base to tip with rows of dense
long dark brown narrow scales. (No distinct bristles are visible
on the legs, but this may be due to the bad preservation of the
specimens.) Wings deep brown, violet-tinged, fore border about
the middle a little yellowish, also just a little paler about the
cross-veins ; tip of wing quite clear, limited by tip of 2nd longi-
tudinal vein, and hindwards on the wing-border by the proximal
side of the 2nd posterior cell.* Alulae and alar squamae fringed
with long narrow dark brown scales ; halteres blackish brown.
Length, 11-18 mm.
* In the second specimen (from Karachi) the wing is pale along the hind
margin of the 3rd posterior cell.
198 BOMBYLIID-S.
Eedescribed mainly from a specimen (? ), and two <5 tf in the
British Museum (Khasi Hills and Cherrapunji, Assam).
148. Exoprosopa lateralis, Brun. (PI. Ill, fig. 7.)
Exoprosopa lateralis, Brunetti, Rec. Ind. Mus. ii, p. 467, pi. xii,
tig. 17 (1909).
$ . Head with frons and face dark olive-brown, becoming more
or less orange towards the lower part of the latter, both covered
, with black pubescence and (except the upper part of the frons)
with small pinkish-yellow scales ; proboscis dark brown, palpi
EXOPBOSOPA. 199
yellow; antennae brown, 1st and 2nd joints bristly; occiput dark
grey, with yellowish scales around eye-margins. Thorax black, with
long bright ferruginous scales or bristly hairs on anterior margin,
below shoulders, and on posterior calli ; dorsum (denuded) evi-
dently more or less covered with very short yellow scale-like hairs
closely applied, which towards the "sides and hind margin are
thinner and longer ; lower part of thorax bare, with ferruginous
marks.* Abdomen elliptical ; black, the sides more or less broadly
brick-red ; this coloration is very variable in quantity, as in one
specimen it only moderately narrowly borders the abdomen, and in
another one it extends so far inwards as to reduce the black part
almost to a dorsal row of spots. Bunches of ferruginous elongate
scales at each side of 1st and 2nd segments, and a small tuft of
stiff black hairs at the sides of each segment ; dorsum thinly covered
with short tawny yellow scales and black hairs ; last segment red-
dish, black at the middle of the base ; Tenter brownish brick-red,
with sparse yellow hair-like scales. Leys (mainly missing) tawny
brown : fore coxae with rather long yellow hairs ; hind femora and
tibiae with short black spines. IVinys dark brown, tip and
posterior border nearly to the base irregularly and indistinctly
clear, the clear part extending into the discal cell ; traces of a
small round clear spot in upper corner of 2nd basal cell : alulae
and alar squama? with dirty brown elongate scales.
Length, 14-18 mm.
Described from three £ ? in the Indian Museum from Calcutta ;
and Talewaddi, N. Kauara Distr., 3-10. x. 1916 (Kemp).
149. Exoprosopa annandalei, Brun. (PI. Ill, tig. 8.)
Kvoprosopa annandalei, Brunetti, Rec. Ind. Mus. ii, p. 469, pi. xii,
tig. 20 (1909).
$ . Head with vertex, frons, and face blackish, with short sparse
yellowish hairs, and some stiff bristly black hairs on frons, which
is considerably narrowed on the vertex, where the ocellar triangle
is very small ; mouth-opening yellowish, proboscis dark brown ;
palpi black, short, filamentous, with a single row of hairs ; antennae
yellowish, first joint hairy, third rather elongated, with moderately
long style ; back of head dark grey ; eyes dark brown, with some
yellowish hairs, which are whitish behind the eyes. Thorax
blackish ; tawny-yellow hairs rather thickly placed on fore border,
and on the sides, where there is a pale tawny spot on the pleurae ;
dorsum nearly bare (? denuded). Scutellum light brown, posterior
border with a row of black spiny bristles and short sparse
yellowish hairs ; metanotum bidden. Abdomen ovate, chestnut-
brown (ground-colour), with a row of irregularly oval dorsal
black spots at the base of each segment, and with traces of a
narrow black line almost on the posterior borders of some of the
segments ; a sparse band (interrupted in the middle) of short
* Possibly due to discoloration.
200
whitish hairs on the anterior part, and a similar band of blackish
hairs on the posterior part, of each segment ; the second segment
possessing a basal row of short sparse tawny hairs and a bunch oi
longer white hairs on each side at the base ; the short whitish hairs
on the dorsum may possibly extend over the greater part of the
surface, and appear more like scales than hairs. Venter chestnut-
brown, with irregular black marks and with short white scaly hairs
generally distributed over it. Legs tawny brown, fore coxae similar,
posterior coxa3 blackish brown ; hind femora with a few hairs
below, posterior tibise moderately covered with short bristles ;
tarsi blackish brown, minutely pubescent below. Wings pale grey,
with a dark brown oblique baso-costal band, which has an indenta-
tion on the lower side : the brown colour extends along the costa
to the tip of the first longitudinal vein, its outer edge extending
posteriorly to just behind the second posterior cell, the colour thus
filling slightly more than half the first longitudinal and discal cells,
and extending in an irregularly straight line from the latter cell
to the base of the wing, filling half the anal cell ; round brown
spots, all of equal size, are placed at the tip of the second longi-
tudinal vein, at the base of the fork of the third longitudinal and
at the tip of the veinlets dividing the second, third, and fourth
posterior cells; on the upper corner (adjoining the base of the
discal cell) of the lower basal cell a small bluish opalescent spot ;
at extreme base of wings and along the stronger veins a distinct
tawny colour ; halteres yellowish white.
Length, 9-12 mm.
Described from the two original females in the Indian Museum
from Moulmein, Lower Burma, 6. iii. 1908 (Annandale).
150. Exoprosopa maculiventris, sp. nov.
J . Head wholly blackish grey ; frons about half as wide again
at level of antennae as at vertex, with fine black hairs, intermixed
with short yellow ones on lower part ; face with sparse short
yellow hairs and on lower part with rather short black hairs ;
antennae and mouth-parts black ; the considerably puffed-out
occiput with a few short yellow hairs, and on hied border a
narrow fringe of them. Tliorax and scutellum black, with mode-
rately short black pubescence, which is longest on the scutellum ;
anterior border narrowly with short yellow hairs, and a con-
spicuous bunch of long bright yellow pubescence below each
shoulder ; some yellow hairs along the sides below the wings and
a large bunch below the posterior angles of the dorsum ; the
posterior calli themselves having a few much shorter and paler
yellow hairs on the hinder side. Abdomen black, surface of seg-
ments (except 1st) mainly covered with small closely-set black
scales, and bearing long fine black hairs, except at base and hind
margins ; 1st segment dark blackish grey, unsealed, with short
yellow hairs on dorsum and a bunch of long bright yellow hairs
EXOPEOSOPA. 201
towards and over each side ; on the 2nd segment towards each
anterior corner is a circular spot of yellowish- white scales ; this
pair of spots is duplicated on the 3rd and 4th segments, the scales
forming them being less yellow on the 3rd segment and nearly
white on the 4th ; oih segment bearing no white spots, but 6th and
7th each with a similar pair of white scale-spots ; sides of 2nd
segment with long yellow hairs anteriorly (as in 1st segment) and
black hairs posteriorly ; sides of remaining segments with thick
black pubescence. Venter black, with small closely impressed
black scales and thin black pubescence ; sexual organs too con-
cealed for observation. Leg* wholly black, with ordinary pubes-
cence ; tibia? with obvious closely impressed small black scales and
two distinct rows of bristles on hind pair, with numerous others
more or less arranged in longitudinal rows ; hind femora with a
little soft black hair and a row of distinct, fairly long bristles on
underside. Wings with about anterior half deep blackish brown,
the colour filling the marginal cell except the tip, thence running
parallel to the hind margin of the wing, filling half the 1st
posterior cell ; thence retreating, filling barely half the discal cell ;
thence extending to the hind angle of the wing, filling half the
anal and axillary cells. An oval spot at tip of 2nd vein, joining
the costa and also the extremity of the costal darkening ; a spot
at fork of 3rd vein, another over the cross-vein joining the 2nd
vein, and another over the posterior cross-vein ; all the spots
concolorous and contiguous to the main expanse of colour. Alar
squamae black, with short fringe of bright yellow hair; halteres
black.
Length, 14 mm. ; expanse, 34 mm.
Described from a single perfect specimen in the Indian Museum
from Kousauie, Garb wal State, 6075 ft.,22. vii.1914 (Col. Tyiler).
The wing-markings are very nearly similar to those of E. annan-
dalei, figured in Rec. Ind. Mus. ii, pi. xii, fig. 20, except that in
this latter species the colour is of a much lighter brown.
151. Exoprosopa stylata, sp. nov.
tS • Head with f rons at vertex about two-thirds as wide as at level
of antenna?, where it is less than one-third the width of the head ;
covered with dull brownish tomentnn or dust with a tinge of yellow,
sparsely beset with stiff black hairs ; lower part of f rons with
depressed black scales, which in certain lights exhibit a silvery
lustre; face distinctly produced into a blunt cone, covered thickly
with similar black scales (shining silvery) and long bristly hairs
as on frons ; upper edge of mouth-opening furnished with very
long, thin, dark brown scales ; mouth-opening and proboscis brown,
latter extending forward as far as tip of antennas. Antenna?
black, 3rd joint elongate conical, as long as 1st and 2nd together,
with a style as long as the joint itself, bearing a pale elongate thick
bristle at, tip. Thorax black, faintly shining ; at least hinder part
with small impressed black scales (which possibly cover the entire
202 BOMBTLIID^I.
surface) : dorsum also with stiff black pubescence of moderate
length ; anterior margin with a thick fringe of very elongate
bright yellow scales projecting forwards, forming a sort of dense
collar ; shoulders entirely covered with a dense bunch of very
elongate black scales, which extend (diminishing in strength) over
the mesopleura, and continue as short stiff hairs over part of the
sternopleura ; pteropleura bare, shining, but the metapleura with
a thick bunch of black scales on its posterior part ; sternopleura
with a slight greyish tinge on lower part. Scutellum black, with
minute adpressed black scales covering its entire surface, and a
row of curved black bristles on hind margin. Abdomen black,
faintly shining, entirely covered with minute adpressed black
scales, which, if viewed at a low angle from in front, show a silvery
sheen ; and with a row of black bristles on hind margins of at least
the last two or three segments, the hind margins of which bear a
row of rather short but obvious dark brown scales ; the outer two-
tliirds of 1st segment with a dense covering of very elongate,
bright chrome-yellow scales, the actual sides with elongate black
scales ; sides of other segments also with elongate very narrow
dark bro\vu scales, and a few other quite broad dark brown ones,
intermixed posteriorly with black bristly hairs ; six spots coin-
posed of scales, which, viewed from in front, appear silvery, and
from behind dull milk-white, are placed as follows : one towards
each side of the dorsum on 3rd segment, one at base in centre of
4th, one towards each side of 6th, and an elongate one at base of
7th extending over the greater part of the segment. Venter
covered uniformly with minute brownish-black scales that appear
silvery when viewed at a low angle either from in front or behind ;
a row of bristles on hind margin of each segment. Genitalia
rather large, apparently consisting of a small sub-globular reddish-
piece and a greyish cone-shaped piece of larger size, pubescent.
Legs black, coxse with bristly hairs ; femora closely covered with
black scales that appear silvery in certain lights ; tibiae apparently
reddish brown, but in certain lights the scales shine silvery; tarsi
brown, Wings pale grey, iridescent ; a brown baso-costal band,
limited distally by tip of 1st longitudinal vein, the hinder margin
of the band nearly a straight line thence, through the anterior
cross-vein and extreme base of 4th posterior cell, and thence to
base of wing, encroaching narrowly on anal cell; discal cell
nearly clear, a darker brown spot jointly over tip of praef urea and
anterior cross-vein ; another over base 'of discal cell and forking
of 5th vein, a small one at base of 3rd posterior cell.
Length, 8 mm.
Described from a unique specimen from Hadagalla, Ceylon
(T. Bainbrigge FletcJier).
Type in the British Museum, presented by Mr. Fletcher.
The conspicuous silvery-white spots on the abdomen placed
like the " 5 " of a playing card, in addition to the exceptionally
long antennal style, render this species easily recognisable.
EXOPEOSOPA. 203
152. Exoprosopa insulata, Walk. (PI. Ill, fig. 4.)
Anthrax (s. g. Litorhi/nchus) insulata, Walker, Ins. Saund., Dipt.
pt. iii, p. 11-2 (1852)'.
, Brunetti, Rec. Ind. Mas. ii, p. 440 (1909).
Head with frons about half as wide again above antennae, where
it forms one-third the width of the head, as at vertex ; dull olive-
brown (in well-preserved specimens, otherwise blackish), with black
pubescence and minute elongate pinkish-yellow scales or scale-like
hairs ; face concolorous, but yellowish about the mouth-opening,
with similar hairs and small scales ; proboscis black, projecting
some distance beyond tip of antennae ; antennas black ; occiput
dark grey, with very small yellow scales in the neighbourhood of
the eye-margins, hinder edge with a fringe of short yellow hairs.
Thorax dark olive-grey or olive-brown (in perfect specimens),
with short black pubescence and very minute elongate yellow
scales, which perhaps are sparser about the centre of the dorsum ;
anterior margin with long brownish-yellow scales, which are also
fairly abundant from below the shoulders to below the wing-base,
and, again, below the posterior corners of the dorsum. Lower
part of prothorax, the mesojileurge, pteropleurae, and sternopleurse
with long black bristly hairs; scutellum concolorous with thoracic
dorsum, with small yellow scales and a few black hairs, arid a row
of rather sinuous thin black hairs on posterior margin. Abdomen
rather dull black, slightly shining: 1st segment with a bunch
towards each side of comparatively short, very erect, elongate,
brownish-yellow scales, and a narrow band of such erect scales
extends across the segment on the anterior half ; 2nd segment
with broad band on anterior half of small elongate closely-applied
yellow scales, the posterior half being covered with black scales of
similar nature ; 4th and following segments with similar black
scales on anterior part and yellow scales on hinder part ; 4th seg-
ment with a pTitch of silvery-white scales towards each side, and
7th segment mainly covered with such scales ; sides of abdomen
with long brown scales and long black hairs ; venter black, with a
little black pubescence and covered with minute yellow scales.
Legs black ; tibiae more or less dark brown ; femora with small
closely-applied pale scales, posterior femora and tibiae with the
usual bristles. \Vin(/s pale grey, with a broad oblique dark brown
or blackish-brown band extending on the costa to tip of 1st longi-
tudinal vein, and distinctly beyond bifurcation of 2nd and 3rd
veins, and anterior cross-vein, embracing basal third of discal cell
and crossing anal and axillary cells about their middles ; several
blackish spots are present, variable in number and size, four on
the clear part of the wing being always present, placed respec-
tively (l)near tip of 2nd vein, (2) at fork of 3rd vein, (3) on
2nd "vein at the junction of the veinlet joining this vein to the
•3rd, and (4) over outer side of discal cell, generally elongate and
sometimes reduced to two separate spots ; on the brown part of
the wing, the spots being much darker in colour, there are
generally one over the anterior cross-vein, one at the fork of the
204 BOMBTLIID^.
4th vein, and at the base of both the 3rd and 4th posterior cells ;
adventitious spots freely occur about the tips of the 2nd and 3rd
longitudinal veins, and occasionally in other parts, and the 3rd
posterior cell is often more or less bisected longitudinally by an
additional incomplete veinlet. The 1st posterior cell is not infre-
quently closed just before the border. Htilteres black.
Length, 7-10 mm.
Redescribed from several specimens, mostly in inferior condition,
in the Indian Museum from Soondrijal, Chitlong, Nagorkote,
Pharpiug, and Khumdhik, iii. 1909 — all in Nepal ; Dehra Dun, xi.
1907 [Lt.-Col. Wyville Thomson'] ; Nairn Tal District, 4. iv. 1910 ;
Mussoorie, ix. 1906 ; Simla, x. 1911 ; Allahabad, v. 1911 and xii.
1910 (Chatterjee); Bandhara, Central Provinces, l.xii.1912 (Imms};
Mahagany, Ceylon, 17. i. 1912 (Col. Yerbury); Castle Eock, K".
Kanara District, 11-26. x. 1916 (Kemp); Madhupur, Bengal,
16. x. 1909 (Paiva); Pipera, Gondo District, United Provinces,
9. iii. 1909; Ootacamund, S. India, 24-31. xii. 1913 (Fletcher) ;
lower ranges, N. Khasi Hills, 1878 (Chennell).
There seems no reasonable doubt of this being Walker's
insulata, described originally from the " East Indies." It is
common through Nepal and extends to the Indian plains.
Walker says the sides of the abdomen are fringed alternately
with black and yellow hairs, but this does not appear to be the
case, though possibly in perfect specimens the white scales on
the 4th and 7th segments may overhang the margin sufficiently
to produce this effect. His description also of the wing-spots
does not quite agree with mine, but these markings are admittedly
variable.
A specimen in the Indian Museum from Dehra Dun labelled
" E. S-notdta, Big., sp. nov/' is certainly identical, and is
apparently merely a MS. name ; a second specimen with precisely
similar markings is from Calcutta.
A further specimen from " Thingaunyinaung to Myawadi,"
Lower Burma, 900 ft,, 24-26.xi. 1911 (Gravely), has tha spot* in
the clear part of the wing all so enlarged and blurred that hardly
any grey surface remains.
Some notes of mine on the variation of the wing-markings
appeared recently (Eec. Ind. Mus. ii, p. 446).
In comparing the descriptions of insulata, Walk., bengalensis,
Macq., and brahma, Sch., it will be seen that they are somewhat
similar. The latter I have not seen, but a number of specimens of
both insulata and bengalensis in the Indian Museum (both iden-
tified by me, but, I think, without possibility of doubt) prove that
at least these two are quite good and distinct species. E. insulata
is known by the black or deep blackish-brown colour of the baso-
costal band, which, though not actually clearly defined, stands
put if a number of specimens be glanced at collectively ; whereas
in bengalensis the colour is brown, without any trace of' black, and
there is no apparent line of demarcation between the anterior
half and the posterior half, the latter being only a little lighter.
EXOPROSOPA. 205
The spots in insulate stand out distinctly, especially those nearest
the wing-border ; they are black or blackish brown, and some of
them are placed elsewhere than on the cross-veins. The "spots"
in bengalensis are small, moderately light brown infuscations, and
occur only on the cross-veins, although the veius themselves are
often lightly infuscated for considerable distances. E. brahma
has distinct spots, apart from infuscations of cross-veins, and
should therefore in this character approximate to insulata, but
Schiner distinctly describes the anterior part of the wing as only
smoke-brown. Again, in brahma the scutellum and antennae are
reddish yellow or brownish, whereas in both Insulata and bengalensis
these parts are black. The presumption is that all three are good
species.
153. Exoprosopa puerula, sp. nov. (PI. Ill, fig. 9.)
Head with frons distinctly broader at level of antennae, where it
is one-third the width of the head, than at vertex ; both frons and
face blackish, former more or less cinereous grey towards vertex,
latter orange yellowish around the mouth ; the lower half, or rather
more, of the frons and the major part of the face covered with
closely-applied yellowish scales and a little fine black pubescence ;
some yellow hairs about the lower part of the face ; proboscis
blackish ; antennae blackish grey ; occiput blackish, with a con-
siderable amount of dirty white scales, closely applied. Thorax
blackish, thickly covered with very thin yellowish hair-like scales ;
anterior border with a fringe of long brownish-yellow scales ;
some long bristly black hairs on lower part of prothorax ; pleurae
with bristly scale-like brownish-yellow hairs ; scutellum black,
about the apical half reddish brown, the whole dorsum (apparently)
covered with short yellowish scales like those of the thorax.
Bristles of thorax normal. Abdomen blackish, dull ; hind border
of segments rather narrowly brownish yellow, the colour spreading
out considerably towards sides of 2nd and 3rd segments ; the
whole dorsum covered with black hairs and closely-applied small
brownish-yellow scales, which towards sides of 3rd segment are
whitish ; a bunch of elongate pale brownish-yellow scales at sides
of 2nd segment and a similar bunch of shorter small black scales
at sides of 3rd segment. Venter yellowish, marked here and there
with black, and mainly covered with small yellow scales. Legs
black ; femora and tibiae with yellowish scales and the usual bristly
hairs ; tarsi black. Wings clear, the dark oblique band running
diagonally from tip of auxiliary vein, crossing the centres of the
anal and axillary cells ; a loop extending from the middle of the
1st posterior cell, along the outer side of the discal cell ; the colour
also encroaches well into the base of the 4th posterior cell ; a pale
yellowish spot around the anterior cross-vein and base of 4th
posterior cell, a semi-hyaline small spot about the base of the
discal cell; halteres brown ; clubs dull yellow.
Z/ength. 4|-6 mm.
Described from two specimens in the Indian Museum from
Madras (Capt. Cragg).
206 BOMBTLIIDJB.
154. Exoprosopa bengalensis, Macq. (PI. Ill, fig. 5.)
Exoprosopa bengalensis, Macquart, Dipt. Exot. ii, p. 49, pi. xviiir
fig. 4 (1840) • Brunetti, Rec. Ind. Mug. ii, p. 445 (1909).
c? $ . Head with frons at vertex half as wide as at level of
antennae, where it is barely one-third the width of the head ; dull
olive-brown or cinereous, the amount of yellowish tinge variable ;
whole frons and face with black pubescence in moderate amount,
also (except towards vertex of former) covered with short narrow
yellowish scales, closely applied to the surface ; mouth-border
brownish yellow, with black hairs ; proboscis blackish brown,
labella narrow ; palpi black, thin, with fine black hairs ; antennal
1st and 2nd joints brownish yellow, with black bristles, 3rd joint
black ; occiput blackish, with yellowish scales around eye-margins.
Thorax brownish grev, dorsum covered with short coarse yellowish
scale-like hairs ; anterior margin with elongate orange-yellow
scales, thickly placed, and bunches of similar or slightly paler
scales below the shoulders, on the mesopleurse, and around and
behind the wings ; black spines on humeral and posterior calli ;
scutellurn blackish, hind margin more or less reddish brown,
covered with short yellowish hairs and with a row of black
bristles on posterior margin. Abdomen blackish (?) or reddish
brown, wholly covered with very small elongate yellowish scales
and black bristly hairs ; a bunch of elongate yellowish scales at
sides of 1st segment ; venter blackish or reddish brown, with a
few long yellow hairs ; hind margins of segments more or less
pale, with some yellowish scales. Legs blackish ; femora and
tibiae with rather thickly -placed, closely-applied, yellow scales,
which extend over the upper side of the metatarsi, and apparently
occasionally to the remaining joint also; the whole of the legs
with black bristles, the tarsi with black pubescence. Winr/s
yellowish grey; a slight infuscation over practically all those
parts of the veins that, lie transversely; the veins themselves
more or less darkened basally ; halteres yellowish, knobs a little
darker; alulae and alar squamae with a dirty brown and yellowish
fringe of scales respectively.
Length, 10-12 mm.
Described from several examples in bad condition in the Indian
Museum, labelled "? India"; Hadagalla, Ceylon, ix, x. 1911 ;
Trincoraali, 14. i. 1891, and Mahagany, Ceylon, 15. ii. 1891 (Col.
Yerbury).
The genitalia are quite hidden ; the sex of the specimens is
therefore uncertain, and no difference in the width of the frons
in any of them is apparent.
Type in the Paris Museum.
155. Exoprosopa brahma, Scli. (PI. Ill, fig. o.)
Exoprosopa bralima, Schiuer, Reise ' Novara,' Dipt. p. 199 (1868).
"Brown ; venter and scutellurn reddish yellow, latter at base
blackish, \\hole ground-colour of body can be seen to be pale
EXOPROSOPA. 207
brown through the dense pale pubescence. Thorax in front and
at sides with longer golden-yellow hairs ; in front of and around
wing-base with black bristles. Abdomen at the base with thick
and long golden-yellow hair; 2nd segment on fore border with a
whitish- yellow cross-band. Head reddish brown, epistome paler,
frons darker; pubescence short, bright golden yellow; frons
black on hinder part. Antennas brown, 3rd joint very elongate,
gradually tapering and produced into the style. Legs reddish
yellow, knees rather darker ; tarsi brown ; femora with shining
scales ; all bristles black ; pulvilli rudimentary. Wings sooty
brown, darker on front margin and more reddish, with eight
black spots : two at base of outer marginal (cubital) cell,.
two at base of 2nd posterior cell, one each at bases of 3rd and
4th posterior cells, one on outer side of discal cell, and the last
one suffusing the anterior cross-vein. Cross-vein joining upper
branch of cubital vein to subcostal vein, with a recurrent
appendix ; only three submarginal cells. Length 4 lines. Two
specimens ; Ceylon."
The above is Schiner's description ; he compares it with
E. alexon, Walk., and punctulata, Macq.
Type presumably in the Vienna Museum.
156. Exoprosopa niveiventris, Brim.
Exoprosopa niveiventris, Brun. Rec. Ind. Mus. iii, p. 214 (1909).
2 . Head : froiis at widest (level of antennae) one-third the
width of the head, narrowing at vertex to half this width ; dark
brown, covered with close, light brownish-orange scales, which
become lighter coloured below the antenna ; the whole frons
covered with short black hairs ; ocelli small, black, close together,
placed on a small tubercle ; mouth-border yellow, proboscis dark
brown. Antennae black; 1st and Und joints thickly beset
above and at the sides with strong black bristles ; 3rd joint
rather long, absolutely bare, with a very distinct style, which
itself bears a microscopic but distinct apical joint. From the base
of the antennae obliquely to the edge of the eye there is on each
side of the frons a narrow groove, quite destitute of scales or
pubescence. Back of head broad behind the eye, with minute
black pubescence, and covered towards the sides with soft
yellowish-white scales, which may perhaps extend over nearly all
the back of the head in some specimens; a fringe of short, bright
orange-yellow hairs encircles the back part of the head, meeting
the dense pubescence of the thorax. Thorax dull black, with
some short black hairs towards anterior border, and in front of
the scutelluni ; front part of thorax (not extending to the dorsum
proper) covered densely with narrow, elongate, bright yellow
scales, which also extend to the sides of the thorax, and form fan-
shaped bunches below each shoulder and behind each wing;.
these scales become almost thick pubescence at the sides of the
dorsum, being especially thick above each wing. The true dorsum-
208 BOMBTLIID.E.
is very sparsely covered with very short yellowish scales, which
probably even in perfect specimens never wholly cover the surface;
scales at sides of thorax more yellow than orange. Posterior calli
prominent, ferruginous, covered with irregular small black bristles,
with some long yellowish-white scaly pubescence on the outer
side, and there are four powerful long black spines placed close
together and directed horizontally backwards. Scutellum ferru-
ginous brown, with an irregular double row of black spines on
posterior margin, and short yellow scales round the dorsum, which
is quite bare in the middle, both of scales and hairs. Abdomen
black, covered with microscopic black scales ; thick, scaly, whitish
pubescence at the sides of the 1st segment; at the base of the
'2nd segment (which is the widest) a wide band of scaly yellowish-
white pubescence ; the remaining segments also bear similar bands
at their bases of more whitish scaly pubescence, this being
shortest on the 5th segment ; on the posterior borders of the 1st,
6th, and 7th segments a row of rather strong black bristles,
directed backwards. Genital organ distinct, black, encircled
towards the posterior part and at the sides by a rather thick
fringe of orange hairs ; the tip itself more greyish, and bearing
two vertical rows (containing seven in each) of rather long,
roseate scales, like the teeth of a comb. Venter black, nearly
covered, except towards the tip, with close, nearly snow-white,
scaly pubescence, the tip with black bristles. Legs black ; coxae
with yellowish-white scales and distinct black bristles; femora
with similar scales and short black hairs, the middle pair with
two rows of short, distinct spines on underside ; the hind pair
with a similar row ; fore tibiae bare, middle tibiae beset somewhat
irregularly with four rows (one on each side) of conspicuous but
moderately short bristles ; hind tibiae closely beset with elongate
dirty white scales, lying close to the surface, and intermixed with
strong, moderately short, black bristles, and with an irregular circlet
of black spines at tip, most of them on the underside ; tarsi minutely
but closely pubescent, with a row of minute spines below, being
longest on hind pair. Wings pale grey, costal cell and the basal
parts of all the veins ferruginous brown ; costal vein apparently
bare, but viewed microscopically, it presents a very closely-placed
row of minute black spines, which are conspicuous on the extreme
base of the costa, where they are larger and are intermixed with
clo?e-lying, dark greyish scales ; at the extreme base of the costa
is a very powerful, thick, black, curved spine. Tegulaa bright
orange, with a fringe of elongated scales, which are black on
outer part of the edge and dirty white towards the base; alula
of wing grey, with a fringe on posterior border of grey scales,
which continue for a short distance along the posterior margin of
the wing ; halteres with stem brownish, club very pale creamy
orange.
Length, 13-20 mm.
Described from two $ 2 in the Indian Museum, taken by
Dr. Annandale at the base of Paresnath Hill, West Bengal,
EXOPEOSOPA. 209
16. iv. 1909 (1000 ft.) ; South Coorg, S. India, 15-26. v. 1914
(Fletcher) ; Trincomali, 15. ix. 1890 ; and Mabagany, Ceylon,
8. ii. 1891 (Col Yerbury).
A conspicuous species, resembling a Ilyperalonia of my fourth
group (Kec. Ind. Mus. ii, p. 443).
A single a
distinct yellow colour to the legs. Wings clear, costal and
subcostal cells and extreme base of wing yellowish brown ;
halteres pale yellow.
Length, 5—7 mm.
Described from a few specimens in the Indian Museum and
Pusa collections from Allahabad, 26. iii. 1906 (type d ), 10. iv. 1906
(type $ ), 14. x. 1905 (Hoivlett).
Type d and $ sent to British Museum.
151. Lepidanthrax transversa, sp. nov.
? $ . General characters as in L. compacta. Head : frons
gradually widening from vertex to level of antennae, where it is
rather narrower than in compacta ; covered with small yellowish
depressed scales and erect black pubescence ; face covered with
dense elongate pale yellowish scales, no black bristles; 3rd
antennal joint short onion-shaped, black, with long black style ;
proboscis lying along mouth-opening, but apparently long enough
to project as far as in compacta if pointed forwards ; occiput
black, with small white scales around eye-margins and yellowish
scales in the middle. Thorax as in compacta. Abdomen with a
broad basal band of small depressed yellowish scales on each
segment except the 1st ; rest of dorsal surface with depressed
dull black scales; posterior margins of 5th and 6th segments
234
with black bristles ; latter also with a fringe of yellowish scales,
which become white at their tips ; 7th segment covered with
yellowish scales and black bristles, hind border with a dense
fringe of long snow-white scales ; venter black, wholly covered
with small pale yellowish scales. Legs black ; femora and tibiae
covered with yellowish scales, middle tibiae brownish yellow-
tarsi black, joints distinct. Wings quite clear, vitreous, subcostal
cell brownish yellow ; base of 3rd posterior cell more sinuous
than in cornpacta ; halteres chalk-white.
Length, 9 mm.
One specimen (? $ ) in the British Museum from Dehra Dun,
Western Himalayas, 1907 (Lt.-Col. Wyville Thomson).
Genus ANTHRAX, Scop.
Anthrax, Scopoli, Ent. Cam. p. 358 (1763) ; Brunetti, Rec. Ind.
Mus. ii, pp. 451, 455 (1909).
Villa, Lioy, Atti Istit. Veneto, (3) ix, p. 732 (1864).
Hemipenthes. Loew, Bevl. Ent. Zeits. xiii, p. 28, note (1869).
Dipalta, Osten-Sacken, Bull. U.S. Geol. Surv. iii, p. 236 (1877).
Chrysanthrax, Osten-Sacken, Biol. Cent.-Amer., Dipt, i, p. 121
(1886).
Isopenthes, Osten-Sacken, op. cit. pp. 80, 96.
Pcecilanthrax, Osten-Sacken, op. cit. p. 119.
Stonyx, Osten-Sacken, op. cit. pp. 80, 94.
Thyridanthrax, Osten-Sacken, op. cit. p. 123.
Hyalanthrax, Osten-Sacken, op. cit. p. 134 (1877).
Aspiloptera, Kiinckel, Bull. Sci. France et Belg. xxxix, p. 145, note
(1905).
? Paranthrax, Bigot, Bull. Soc. Ent, France, (5) vi, p. Ixvi (1876).
GENOTYPE, Anthrax morio, F. ; by Rondani's designation.
Head semicircular, inflated behind, but slightly inarched at the
middle, as broad as or a little broader than thorax ; face very
short, sometimes conically projecting or gently convex ; proboscis
short, withdrawn into the large mouth-opening, labella rather
broad ; palpi small, thin, cylindrical, apparently one-jointed ; three
ocelli ; eyes in both sexes wide apart ; frons rather narrower in
c? , an indentation at middle of hind margin in both sexes.
Antennae porrect, short, wide apart at base ; 1st joint short,
cylindrical; 2nd cup-shaped; 3rd varying from elongate conical
to flattened onion-shaped, with an apical microscopic bristle,
never with an apical pencil of hairs. Thorax quadrate, with
rounded angles, broadest behind, with indistinct preesutural but
distinct postalar bristles. Scutellum broad, short, bare of bristles ;
metanotum concealed. Abdomen flat, oblong, longer than thorax,
7-segmented. Genitalia small, mainly concealed, placed asym-
metrically ; ovipositor with a circlet of short blunt spines. Legs
moderately long, hind pair lengthened and often bearing con-
spicuous scaly pubescence ; all femora and tibiae with short
scaly pubescence in some species ; fore tibiae sometimes quite
bare, sometimes with small spicules; posterior tibiae with
ASTHBAX. 235
an apical circlet of spines ; fore tarsi variable in shape and
pubescence ; front claws varying in size ; pulvilli very small or
absent, occasionally well developed. Wings Avhen at rest half
open. Venation as in the other Antbracine genera; only two
submarginal cells ; 3rd vein sometimes with a short appendix at
its fork ; 2nd vein not infrequently with a recurrent veinlet at
its rectangular bend; both 2nd vein and upper branch of 3rd
often sinuous and parallel towards their tips. A prealar hook
near base of wing, generally hidden by the pubescence.
Ifange. World-wide ; a very extensive genus.
Life-history. Larva amphipneustic, 13-segmented, nearly cylin-
drical, a little flattened below, tapering at each end. Several
European and North American species have been bred from
Lepidopterous larva?, and one from the egg-cases of a locust,
others being known to live in the cocoons of Hymenoptera. "In
cases where the Anthrax larva lives in other Iarva3 these latter
pupate before they are destroyed, and the Anthrax larva lies in
the pupa of its host " (Lundbeck). The images occur in sandy
or dry districts, bare open spots and pathways ; their flight is
short but swift.
Several subgenera of Anthrax were suggested by Osten-Sacken,
founded primarily on the wing-markings : Thyridanthrax for
species with brown wings with more or less hyaline spots; Anthrax
(s. sir.) for those with clear wings, except for a broad baso-costal
dark baud ; Hyalanthrax for those with practically wholly clear
wings or at most an inf uscated costal margin ; and one or two others.
It was urged that these characters were supported by others, and
also by the fact that the species of each group were parasitic on
a totally different group, or even order, of insects. As, however,
intermediate forms are sure to occur, and the subgenera have not
been adopted by subsequent authors, they are herein ignored. So
far as the Indian species go, fletcheri and guttatipennis belong
to a group with wholly dark wings; afra, semifuscata, and
himalayanus would fall in Anthrax (s. s.) ; whilst all the other*
belong to Hyalanthrax.
Table of Species.
1. Anal cell closed clttusa, Brun., p. 253.
Anal cell open 2.
2. Wings nearly wholly dark brown .... 3.
Wings always with an appreciable
clear part 4.
3. No clearer spots in any of the wing-
cells fletcheri, sp. n., p. 23G.
Clearer spots on basal half of wing . . guttatipennis, sp. n., p. 237.
4. A distinct oblique baso-costal wing-
band 5.
No such distinct band ; costa generally
narrowly, dark, light brown, or yel-
lowish, the colour not extending
behind anterior cross-vein (nor-
mallv) ; or wings practically clear . . 7.
236 BOMBYLIIDJE.
•5. Hind margin of \ving-baud nearly
straight C.
Hind margin of wing-band with deep
indentations himalayanus, Brim., p. 238.
6. Wing-band filling less than g of the
wing; hind margin clear cut afra, F., p. 240.
Wing-band filling fully half the wing ;
hind margin indistinct semifuscata, sp. n., p. 241.
7. Legs mainly yellow 8.
Legs all black 9.
8. Femora with a considerable part black ;
wings practically clear albofulva, Walk., p. 251.
Femora all yellowish ; basal half of
wings brownish aureohirta, Brun., p. 252.
9. Abdomen uniformly densely pubescent,
without obvious transverse bands of
scales 10.
Abdomen with transverse bands of
yellowish, grey, or white scales ;
pubescence sparser II.
10. Wing with deep brownish-black costa ;
larger species, 14-16 mm manifesto, Walk., p. 242.
Wing less extensively brown ; smaller
species, 10A-11 mm dara, Walk., p. 243.
11. Pubescence on sides of abdomen
wholly snow-white, except for few
bunches of scales on each side near
tip aperta, Walk., p. 245.
Pubescence on sides of abdomen never
as wholly white as in aperta,
generally yellowish grey or black
and white, and not arranged as in
aperta 12.
12. Ground-colour of abdomen or tomen-
tum (apparently) reddish brown. . . . dia, Wied., p. 250.
Ground-colour of abdomen black 13.
13. Costal cell yellowish baluchiattus, sp. n., p. 250.
Chottentotta vav. clanpennis,
\ [Brun., p. 247.
Costal cell lighter or darker brown*. .•( paniscus, Rossi, p. 248.
| fuscolimbata, Brun., p. 246.
^ approximate, Brun., p. 244.
Four species that appear to require confirmation as Indian
are not included in this table, but descriptions are appended
(pp. 254, 255).
182. Anthrax fletcheri, sp. nov.
c? . Head black ; frons at level of antennae between one-third
and one-fourth width of head, less than half this width at vertex ;
frons and face (apparently) covered with bright chrome-yellow
* These species are too closely allied to be satisfactorily differentiated in
tabular form.
ANTHRAX. 237
small impressed scales, with the addition of black hairs on the
former : antennae all black ; apical end of proboscis brown ;
occiput (apparently) with minute yellow impressed scales and
very short concolorous pubescence ; a fringe of such pubescence
on hind margin of occiput. Thorax black, with small bright
chrome-yellow impressed scales on dorsum and with bright
concolorous long scaly pubescence on anterior and side margins
and on pleura, where it is a little paler below shoulders and
wings. Scutellum black (denuded), apparently with small yellow
scales as on thorax and a row of black bristles on hind margin.
Abdomen black, surface apparently covered with minute black
impressed scales, which are longest on the 2nd segment ; an upper
clothing of bright yellow small impressed scales ; sides of first
four segments with bunches of concolorous elongate scales ; sides
of oth and most of the sides of the 6th with elongate black scales,
hind part of sides of 6th with bright yellow elongate scales; 7th
segment with elongate black scales on hind margin. Venter
black, with minute black scales ; some fine yellow hairs at base
and small yellow scales on hind margins of most of the segments.
Legs black ; coxa3 with some fine yellow hairs ; femora and tibiae
with traces of small yellow scales (mostly denuded) and the usual
black bristles ; middle femora with some pale fine hairs on inner
side. Wings wholly dark brown, the anterior part still darker ;
anterior cross-vein distinctly before middle of discal cell ; anal
cell narrowly open ; halteres creamy yellow.
Length, 12 mm.
Described from a unique $ from Mysore, 4400 ft., 25. iii.-
29. v., sent for examination by Mr. T. Bainbrigge Fletcher, and
presented by him to the British Museum.
183. Anthrax guttatipennis, sp. nov.
Head : f rons at level of antenna? one-third width of the head,
diminishing to one-third as wide at vertex, black, covered
(apparently), as is the face also> with bright chrome-yellow
elongate scaly pubescence, intermixed (at least on frons) with
long black hairs ; antenriaB black, 1st joint bristly, 3rd short
onion-shaped, with long style ; occiput black, with minute greyish-
white scales on eye-margins. Thorax black (denuded), with
bright chrome-yellow scaly pubescence forming a collar and
towards sides of dorsum, covering shoulders and extending over
pleura; scutellum black, with minute bright yellow scales on
posterior margin and some concolorous bristles, with which are
mixed some long black ones. Abdomen black ; 1st segment with
a little ragged bright yellow bristly pubescence; all segments
except 4th* with broad basal bands of depressed small bright
yellow scales, the bands narrowest at the centre of each segment ;
rest of dorsal surface apparently covered with depressed small
Possibly an individual aberration.
238 BOMBYLIID^E.
dull black scales ; a few long black bristles on hind margins
of at least last two segments ; sides of first four segments with
long shaggy bright yellow scaly pubescence ; venter black, basal
halt' covered with small depressed bright yellow scales and some-
what longer concolorous pubescence, and apparently two or more
narrow transverse rows of similar scales on apical half. Leys
black ; coxae with long yellowish hairs and black bristles at tip,
directed backwards ; femora, at least on hinder side, with minute
pale yellowish adpressed scales and a few black bristles below ;
tibiae with some yellow scales on hinder side, rest of surface with
closely-impressed, very dark brown scales and the usual bristles ;
tarsi 'black. Wings dark brown, darker anteriorly ; centre of
both basal cells, discal cell, and 3rd arid 4th posterior cells
narrowly pale : a small elongate pale streak in marginal cell just
before bifurcation of praefurca, and another towards tip of that
cell ; two similar streaks in 1st submarginal cell ; halteres
yellowish.
Length, 13 mm.
Described from a unique specimen (sex uncertain) in the British
Museum from Trincomali Hot Wells, Ceylon, 15. x. 1891
(Yerbury).
184. Anthrax himalayanns, Brun.
Anthrax liimalayanus. Brunetti, Rec. Irid. Mus. ii, p. 222, figs. 1, 2
(1909).
Anthrax maura, Brunetti, op. cit. ii, p. 451 (1909).
S $ • Head black : frons in J and £ fully or over one-third
width of the head at level of antennae, between one-third and one-
fourth this width at vertex in J and about two-fifths in $ ; frons
and face with prominent long black pubescence and minute
elongate yellow scales, which latter are rather longer on face, where
there are also a few yellow hairs ; antennae black, 3rd joint short
onion-shaped, witli a long style ; proboscis black, withdrawn ;
occiput black, with minute yellowish-white scales. Thorax black,
with rather long, fine, very dark brown hairs; the surface apparently
sparsely covered with minute depressed elongate yellow scales ;
anterior margin with a row or fringe of long brownish-yellow
scales ; side margins with long, pale yellow, almost scale-like
pubescence. Sides of thorax light grey, with elongate brownish-
yellow scales and coucolorous pubescence on lower part ; an elon-
gate bunch of long black scales below shoulders and a fringe of
similar scales in the centre of the lower anterior margin, imme-
diately below bead, with elongate brownish-yellow scales on each
side. Scutellum black, with minute orange-yellow depressed
scales on dorsum and with fine black hairs; a row of black
bristles on hind margin. Abdomen black ; 1st segment with
some long yellowish-grey scaly pubescence towards sides, which
stretches across the anterior margin also, posterior margin with a
ANTHRAX. 239
narrow row of minute orange-yellow (varying to yellowish-grey)
scales ; surface of 2nd and 3rd segments, posterior half of 4th, hind
margins, and more or less of the clorsum of 5th, 6th, and 7th with
minute reddish-yellow depressed scales ; anterior half of 4th seg-
ment occupied by a transverse baud of whitish scales ; hind margins
towards the sides of 6tli and 7th segments with similarly coloured
scales. Whole surface of abdomen with fine long black pubescence,
there being apparently no row of bristles on hind margins of any
of the segments ; sides of 1st, anterior lialf of 2nd, and all the
4th with white scaly pubescence ; hind half of 2nd and all the
3rd, 5th, and 6th with black scaly pubescence. Venter rather light
frey, with minute yellow scale-like depressed hairs and long tine
lack pubescence, with longer more bristly black hairs towards
tip; genitalia blackish, tip brown. Legs dark brown: fore coxae
with a few fine yellow hairs, hind pair with black hairs : femora
with minute depressed yellowish-white scales ; tibise with similar
but apparently less numerous scales ; some fine black hairs on
underside of femora ; femora and tibiae with the usual rows of
bristles ; tarsi blackish. Wings very pale grey, with a broad
deep brown baso-costal band occupying about half the surface ;
the colour extending along all the costa to tip of upper branch
of 3rd vein ; posteriorly the colour filling axillary and anal cells
except their tips, nearly half the 4th posterior cell, less than a
third of the 3rd posterior cell, and basal half of discal cell ; a
downward extension (blob) stretching across the 1st posterior cell
and filling half of the 2nd posterior cell, the colour limited by
the outer side of the discal cell ; the colour also fills the basal
half of the 1st submarginal cell and the whole of the marginal
cell, except a small space in a line with the proximal limit of the
clear half of the marginal cell ; a small oval sub-hyaline spot in
upper corner of 1st basal cell, with a narrow longitudinal (not
transverse, as I erroneously noted in my first description of the
species) streak above it.*
Length, 7-11 mm.
Described from three J rf and three $ $ in the Indian Museum,
Kufri to Phagu, 18. v. 1916 ; Phagn, Simla Distr., 9000 ft,,
12. v. 1909, type rf ; Simla, 7000 ft,, 16. v. 1909, type $ (both
Annandale) ; two rf <5 , Naini Tal, v-vi. 1893 (Lucknow Museum );
two $ $ , Simla, 16. v. 1909 and 9. v. 1910, on the flowers of the
white stonecrop (Annandale) ; Khasi Hills, Assam (Brit. Mus.).
This species is very close to A. maura, L., of Europe, and was
referred to under that name in my first paper on this family (I. c.).
I have already detailed the differences between the two species
(llec. Ind. Mus. iii, p. 222).
* A more rough-and-ready description of the wing would be: antero-
proxiuial half dark brown, tlie colour cutting the 4th posterior and discal
cells about their middles ; a large blob from the dark part at two-thirds the
wing-length reaching nearly to the hind margin, a similar smaller blob falling
the curve in the 2nd longitudinal vein, and a minute 3rd blob between the
2ud one and tip of upper branch of 3rd longitudinal vein.
240 BOMBYLIIDyE.
185. Anthrax afra, F.
Anthrax afra, Fabricius, Ent. Syst. iv, p. 258 (1794) ; Schiner,
Fauna Austr. i, p. 50 (1862) ; Brunetti, llec. Ind. Mus. ii, p. 451
(1909;.
Anthrax fimbriatus, Meigen,Klass. i, p. 205 (1804) ; id., Syst, Beach.
ii, p. 154, pi. xvii, %. 13 (1820).
cJ $ . Head : frons one-third as wide at vertex as at level of
antennae, where it forms one-third the width of the head, dark
olive-brown or blackish, with black hairs and small elongate
yellow scales ; ocelli pale, ocellar tubercle quite clear of eye-
margins; face clothed as frons ; eyes dark brown ; antennae black,
2nd joint often dull red, 1st joint with long, 2nd with short black
bristles ; proboscis marked with blackish brown ; occiput black,
with very small yellow scales and minute white scaly pubescence
in centre and around upper half of margin. Thorax from dark
olive-brown to blackish, more or less covered on upper surface
with very small yellow flat-lying scales ; towards the sides with
from pale yellow to brownish-yellow bristly pubescence, which
on anterior margin changes to very narrow elongate yellow
bristle-like scales, a large fan-shaped bunch of which (of deeper
yellow) occurs below the shoulders, extending nearly to the wing-
base ; on the shoulders and extending posteriorly for a short
distance the scales are often yellowish white and nearly silky
in nature ; on the posterior calli a large fan-shaped bunch of
bright yellow- elongate bristle-like scales, with some whitish
ones in front of them. Sides of thorax with some black hairs
and apparently a few yellow hairs around wing-base ; scutellum
with short flat-lying yellow scales and a row of long black bristles
towards hind margin. Thorax with usual macrochsetae. Abdomen
black, 1st segment with small yellow and white scales inter-
mixed ; anterior half of 3rd and 6th segments with small white
scales ; most of the 2nd segment and a band on hinder part of
3rd to 6th segments, yellow-scaled ; rest of dorsal surface with
black scales ; the 7th segment bears white and black scales, and
there is a large fan-shaped bunch of very elongate milk-white
scales on anterior corners of abdomen. Genitalia with a circlet of
reddish-yellow spines ; venter mainly with black scales and a few
long pale hairs, a narrow row of small yellow scales on hind
borders of segments. Leys black or dark brown ; femora with
small, dark-brown, closely-applied scales and others that reflect
a greyish lustre in certain lights ; fore femora with a few stiff,
moderately long hairs below, hind pair with a row of moderate-
sized bristles below ; anterior tibiae with short pubescence, hind
pair with dark brown and greyish-lustred scales and rows of
small black bristles ; tarsi blackish. Wings clear ; a dark brown
oblique band on anterior margin extending nearly to tip of costal
cell and defined posteriorly by a line drawn, with slight angu-
lations at the veins, from that spot diagonally to the base, cutting
the anal cell at one-third of its length ; halteres yellow.
Length, 4g-7 mm.
ANTHRAX. 241
Described from a good number of specimens in the Indian
Museum, Pusa, my own, and other collections. A widely distri-
buted species in India : Dehra Dun, iv. and ix. 1904 (Chatterjee) ;
Anamalai Hills, 4000 ft., 24. i. 1912 (Fletcher) • Allahabad, 24-29.
iii. 1906, 6 and 19. x. 1905 (Hoivlett) ; Satara Distr., 3200 ft.,
Bombay, 23. iv. 1912 (Gravely); Puri, 24-28. i. 1911 (Annandale,
Gravely), 29.ii.1908 and 2.iii. 1908 (Paiva) ; Gonda Distr., United
Prov., 26. ii. 1907, 9. iii. 1909 ; Madhupur, Bengal, 14. x. 1909 ;
Pusa, common, ii, iii, iv, vi, xi : Pipera, Gonda Distr., United
Prov., 9.iii. 1909 (Hocl/art) ; Muttra, 22. iv. 1905 ; Meerut, 25. iv.
1905; Ferozepore, 28. iv. 1905 (all Bnmetti) ; Guindy, Madras
(Capt. Cragg) ; Dawna Hills, Burma, 2000-3000 ft., 2-3. iii. 1908
(Annandale) ; Trincomali, Ceylon, 21.viii. 1890 (Col. Yerbury)-y
Port Blair, Andaman Is., 15. ii. to 15. iii. 1915 (Kemp).
186. Anthrax semifuscata, sp. nov.
cJ 5 . Head : frous at level of antennae less than one-third width
of the head, at vertex less than half this width, blackish grey,
with minute bright yellow scaly pubescence, which latter also
covers the face ; antennas black, 3rd joint short, with long style;
occiput blackish grey, a fringe of small snow-white scales on
lateral eye-margins. Thorax black (denuded), hind corners of
dorsum dull reddish brown; in good specimens the dorsum,
anterior margin, and sides of thorax are probably covered with
bright chrome-yellow scaly pubescence, which extends to the
scutellum also. Abdomen black ; 1st segment with a band of
depressed bright yellow scales ; 2nd, ,3rd, 4th, and 6th segments
each with a basal band of similar scales, broadening towards and
at the sides, where it is augmented by concolorous long scaly
pubescence covering all the sides of the 2nd, 3rd, and 4th
segments ; 5th segment apparently devoid of such a band ; dorsal
surface of abdomen not covered by yellow scales, furnished with
small depressed black scales; venter black, basal half with
depressed bright yellow scales, concolorous scales about the tip.
Legs black ; femora with whitish scales on hinder side (those on
fore pair appear more yellowish); tibiae with some yellowish-
white scales ; tarsi black. Wings pale grey, basal and anterior
parts moderately dark brown, the colour fading away distally,
its approximate limit running from tip of 1st vein, just escaping
fork of 3rd vein, filling discal cell and bases of 3rd and 4th
posterior cells and filling anal and auxiliary cells; halteres
pale yellow.
Length, 7—8 mm.
Described from three specimens in indifferent condition in the
British Museum from Trincomali, Ceylon, 2. x. 1890 (type a ),
9 and 15. x. 1890 (Col Yerbury).
Although not one of the three specimens is in good condition,
the species is a quite a well-marked one by the extent of inf uscation
of the wing. A noticeable point is the variability in the exact
position of the bifurcation of the prsefurca.
187. Anthrax manifesta, Walk.
Anthrax manifesta, Walker, Ins. Saund., Dipt. pt. 3, p. 178 (1852).
c? $ . Head: frontal sexual differences as in A. aperta ; frons
at level of antenna forming one-third the width of the head,
black, with dense short stiff black hairs, which become denser
round base of antenna) and are continued along the middle of the
face ; frons also with very small recumbent hair-like bright
yellow scales ; face, except just below antennae, covered with
bright chrome-yellow or orange-yellow elongated scales, inter-
mixed with yellow hairs. Antennae black, 1st joint broader at
tip, with long black bristles on outer and inner sides and yellow
hairs below ; 2nd and 3rd joints normal ; proboscis blackish ;
occiput black, upper margin with very small whitish scales,
centre with short yellowish pubescence. * Thorax black, dorsum
with soft dark brown hairs, and some small yellow scales about
middle of hind margin, opposite which are some dead-black thin
scales on the scutellum. Anterior border, shoulders, and sides of
thorax with long dense chrome-yellow or orange bristly, almost
pcale-like pubescence, which becomes paler or sometimes even
whitish on underside; fan-shaped brushes of this pubescence
occur as in A. aperta. Abdomen black ; base of first four seg-
ments with small short flat-lying bright yellow scales, thickest on
2nd and 4th segments ; 5th and remaining segments with longer
similar scales; the rest of the dorsal surface with small dead-
black flat-lying scales and sparse long black or brown hairs. Sides
of abdomen with dense scale-like bristly pubescence or very thin
real scales, this vestiture varying from deep orange to yellowish
in different specimens, and being, generally speaking, deeper
coloured on anterior half and paler, sometimes whitish, on hinder
half of abdomen ; sides of 5th and 6th segments with elongate
dark brown scales and longer dark brown or blackish thick bristly
pubescence. Venter black, covered with small bright yellow
scales and sparse fine yellow hairs, both scales and hairs 'longer
and denser towards tip ; hind margin of last segment with elongate
dark brown scales. Legs scaled as in A. aperta ; femora with soft
yellow hairs below; hind tibiae with numerous dark brown or
black elongate scales, intermixed with black spines and some
smaller yellow scales, which latter appear to occur more or less
on the middle tibiae also and to cover most of the femora. Wings
pale grey; costal and subcostal cells dark brown, the colour ex-
tending in diminishing intensity into the marginal and filling the
1st basal cell ; halteres dull yellowish, variable ; squamae brownish,
with a fringe of elongate orange or yellowish scales.
Lcnf/th, 14-16 mm.
Reclescribed from two rf c? and three $ $ from Mussoorie
ix. 1906; Jhajjur, Kashmir, 22.xi.1911; Simla, 26. v. 1914 (Capt
Evans) ; Lower Eanges, Khasi Hills, 1878 (Chennell).
Type in the British Museum.
ANTHHAX. 243
188. Anthrax clara, Walk.
Anthrax cltira, Walker, Ins. Saund., Dipt. pt. 3, p. 179 (1852) ;
Brunetti, Rec. lud. Mus. ii, p. 452 (1909).
c? $ . Head: frons in d six times, in $ three times as wide at
level of antennae (where it forms nearly one-third the width of the
head) as at vertex, black, and, as is also the face, covered thickly with
dense erect stiff black pubescence, amidst which numerous small
bright yellow depressed scales are visible, more numerous on the
face, where they are intermixed with yellow hairs; antennae black,
3rd joint moderately elongate, with rather long style ; mouth-
parts reddish brown or blackish ; hind margin of eyes with a narrow
fringe of small white impressed scales, which perhaps may extend
over most of the occiput. Thorax black, dorsum covered with
minute bright yellow impressed scales, either placed on top of
(forming an upper layer) or intermixed with similar dull black
scales;* fine long black hairs are scattered irregularly over the
dorsum ; margins of dorsum with thick bright yellow scaly
pubescence, a few black bristles at sides in front of wings and
three or four brownish-yellow spines on posterior corners, directed
backwards. Scutellum like thoracic dorsum ; hind margin with a
row of long yellow hairs and (apparently) some yellow bristles
and some elongate black scales. Abdomen : integument between
thorax and abdomen whitish grey ; dorsum of abdomen black,
mainly (or entirely ?) covered with very small impressed black
scales, and apparently the greater part covered in good specimens
with small yellow scales ; margins of dorsum. and sides of abdomen
with dense long fine scaly pubescence, the dorsum covered more
thinly, though, when viewed at a low angle from behind that part,
it appears almost as thickly covered as the sides ; this pubescence is
rather darker yellow in the $ ; on hinder part of dorsum in $
the pubescence is black. Hind margin of 6th segment narrowly,
and basal three-fourths of 7th, with yellow scales ; remainder of
7th with black scales ; sides of 5th and 6th segments with thick
bristly black pubescence ; ground-colour of side margins of first
three' segments in £ yellowish; venter black, with minute black
and yellow impressed scales and some fine yellow pubescence.
Legs black ; femora with closely-placed small black scales, which
in certain lights have a greyish tinge ; small yellow scales also
present; tibiae and tarsi with small black scales and bristles.
\Vinys practically clear; costal and subcostal cells dark brown,
the colour extending irregularly over praefurca and also at extreme
base proximadto basal cells; anterior cross-vein just before middle
( 3 ) or at first 3rd ( 2 ) of the discal cell ; halteres pale yellow.
Length, 10£-llmm.
Eedescribed from a rf from Majkhali, Western Himalayas,
* The dorsum of the c? specimen is mainly denuded, that of the £ partly
so, but probably a double covering of scales is present in perfect specimens.
B 2
244 BOMBYLIID^.
6000ft., 4.x. 1907, and a ? from Naini Tal, 6300ft, 11. v. 1911
(Kemp), both in rather inferior condition but with all the charac-
ters fairly obvious, and both in the Indian Museum, as are also
two others— one named by Bigot from Dehra Dun, the other
from the Garhwal Distr., 21. x. 1907. These two specimens have
been compared by Mr. P. W. Edwards with the types in the
British Museum, where the species also figures from Dehra Dun,
xi. 1907 (Lt.-Col. Wyville Thomson), and the Khasi Hills, Assam.
Type in the British Museum.
189. Anthrax approximata, Brun.
Anthrax approximata, Brunetti, Rec. Ind. Mus. xiii, p. 74 (1917).
$ . This species is very close to A. clara, "Walk., yet certainly
distinct. The scales on the frons and face vary from yellow to
snow-white, in one specimen only being yellow on both parts,
in others yellow on frons and white on face, those of the latter
colour encroaching to some extent on lower part of frons ; in one
example snow-white on practically the whole of both frons and
face ; sometimes some yellow scales around the mouth-parts amidst
the white ones. Abdomen not uniformly pubescent, as is pro-
bably the case in clara, hut with distinct transverse bands of
yellow or whitish scales at base of segments, those on the 2nd and
4th being widest, extending on the former segment nearly to the
middle, and on the latter much further, especially towards the
sides ; hind borders of 6th segment with conspicuous snow-white
scales and a large bunch of longer ones on about the hinder half
of each side; sides of 5th segment and anterior half or more of
6th with long very dark blackish-brown scales ; dorsum of abdo-
men, except for the transverse bands, with black scales, wholly
covered with soft fine pubescence, and with a row of black bristles
on hind borders of 5th, 6th, and 7th segments, becoming stronger
on each successive segment ; a row of fine yellow hairs before the
hind border on first four segments and a few fine yellow hairs on
dorsum of basal segments. Costal cell clear or faintly obscured ;
subcostal cell dark brown, the colour not extending further hind-
wards ; anterior cross- vein a little before middle of discal cell,
the exact position apparently rather variable. In all else as in
A. clara.
Described from five rf 3 in the Indian Museum : Simla, 7000-
8000ft., 28. v. 1914, type (Cap*. Evans), and 16. v. 1909 ; Siliguri,
base of Darjiling Hills, 28. iii. 1910 ; Dawna Hills. Tenasserim,
2000-3000 ft., 2-3. iii. 1908 ; Kawkareik, Amherst Distr., Tenas-
serim, 5. iii. 1908,* and 23. iii. 1910 (Annandale). In the British
Museum from Khasi Hills, 1878 (Chennell); Trincomali, Ceylon,
12.x. 1890, 12. xi. 1890 (Col. Yerbury); Nilaveli, Cevlon,
19. vii. 1891 (Col. Yerbury).
* Erroneously quoted as 5. ii. 1908 in Eec. Ind. Mus. ii, p. 474.
ANTHBAX. 245
The two specimens alluded to by me in my first paper on
BOMBYLIIDJE as allied to A. dam, Walk., are amongst those now
referred to approximata.
190. Anthrax aperta, Walk.
Anthrax aperta, Walker, Ins. Saund., Dipt. pt. 3, p. 180 (1852).
3 . Head : f rons in tf sufficiently narrow at vertex for the
small ocellar tubercle to touch the eye on each side, in $ broad
enough for this tubercle to be quite clear of the eyes ; in both
sexes frons broadening to base of antennae, where it forms one-
third of the head, black, covered wholly with black bristles and on
the lower part with elongate black or dark brown scales and some
shorter yellow or yellowish-white scales intermixed ; face in c?
wholly and thickly covered with elongate bright yellow scales,
with which are intermixed long black hairs, and in the median line
from base of antennae to mouth-opening a stripe of dark brown
scales : in $ the face-scales mostly whitish, those in the median
line yellowish ; eye-facets small, uniform ; antennas black, 1st and
2nd joints bristly, 3rd onion-shaped, produced into a long bare
style. Thorax black, covered with moderately short blackish-brown
hairs ; anterior margin with a thick fringe of reddish-tawny
bristly hairs, which extend to a considerable portion of the under-
side. Sides of thorax with long whitish-grey scale-like pubescence,
forming fan-shaped bunches about the humeral and posterior calli,
near the wing-base and on the pleurae ; this pubescence varies
from nearly white to dirty grey and yellowish. Along the hind
border of the mesonotum there 'is always, in the narrow depression
occurring just before the extreme margin, a row of very small flat-
lying bright yellow scales.* Scutellum black, with some soft
black hairs and (probably) with whitish pubescence.f Abdomen
black ; at base of 2nd segment a moderately wide band of very
small flat-lying white scales; a similar but narrower band of
yellow scales at base of 3rd segment and a row of yellowish or
whitish (or apparently both intermixed) at base of remaining
segments; rest of surface covered with small flat-lying black
scales ; entire dorsum covered with rather long fine dark brown
hairs, which along the basal bands of pale scales appear to be
replaced by whitish ones. Sides of abdomen thickly clothed
on first two segments with long bushy scale-like whitish hairs ;
on 3rd segment with long very dark brown scales and hairs, which
extend a little over the sides of the 2nd segment; on 4th segment
with elongate white scales, connected with the basal cross-baud
* A few of these generally remain even in badly-denuded specimens, but
there seems no reason to infer that they extend beyond the narrow space
t This is conjecture, not one of the specimens before me bearing any
pubescence.
246 BOMBYLILD.S:.
of whitish scales (which, after those on the 2nd segment, are the
most conspicuous of these transverse scale-bands) ; on 5th and
6th segments with elongate dark brown scales and hairs ; and on
the 7th segment with white scales. This vestiture of the sides
of the abdomen is quite distinct from the dorsal, and of much finer
pubescence. Genitalia dark brown or blackish, with some line
black hairs. Yenter black, with flat-lying dull black scales and
sparse black hairs; hind margins of segments with a more or less
broad border of whitish or yellowish scales. Legs black ; coxae
with long yellowish-grey hairs ; femora with closely-placed very
dark brown scales and a further clothing on underside of yellowish-
white scales, also with some long fine hairs ; tibiae with very short
bristles, hind pair with short but distinct very dark brown scales.
Wings quite clear ; base of costa with thick dark brown scales,
over which lies a covering of elongate silvery snow-white scales;
costal cell dark brown up to humeral cross-vein ; subcostal cell
wholly brown, the brown colour also extending very narrowly
across the wing at the extreme base ; halteres dull pale yellow.
Length, 8-14 mm.
Eeclescribed from several specimens fi-om both Mussoorie,
ix. 1906, and Simla, 16. v. 1909 (Annandale) ; Simla, x. 1911 ;
Dehra Dun, xi. 1907 (Lt.-Col Wt/ville Thomson"); Sukli, Dawna
Hills, 2100ft., 22-29. xi. 1911 (Gravely); Khasi Hills. Found
on Sedum and the common marguerite.
Type in the British Museum.
This species is fully redescribed here from four c? c? and five
$ § , mostly in tolerably good condition, and this must super-
sede my former description, drawn up from a single specimen.
191. Anthrax fuscolimbata,
Anthrax fuscolimbata, Brunetti, Kec. lud. Mus. xiii, p. 75 (1917).
$. Head: frons one-third width of head at level of antennae,
less than half this width at vertex, covered with yellowish
impressed scales and black pubescence, the scales near the antennae
sometimes becoming gradually whitish ; on face all snow-white in
type, and yellowish white and yellowish respectively in the other
two specimens ; antennae black, 1st joint with long black bristles,
2nd with a ring of shorter bristles, 3rd elongate conical, tapering
to a rather long style; proboscis dull yellowish or obscure;
occiput with a border of snow-white minute scaly pubescence
behind eyes, and some yellow scales on upper part. 'Thorax black,
more or less covered with small impressed yellow scales ; anterior
and side margins, shoulders, and pleurae covered with dense elon-
gate brownish-yellow scales, which are paler on the mesopleura
and sternopleura or sometimes altogether paler; a rather short
fine sparse black pubescence on dorsum, sometimes remaining after
nearly all the yellow scales have been worn off. Scutellum black,
with small yellow impressed scales ; a few fine yellow hairs and a
ANTHKA.X. 247
row of black bristles on hiud margin. Abdomen black, with trans-
verse bauds of small yellowish, yellowish-grey, or whitish scales,
broadest on 2nd and 4th segments, where, especially towards sides,
they reach up to .and beyond middle of segment ; a narrow basal
band on 3rd segment and a narrow band on hind margins of 5th
and 6th ; a large bunch of elongate brownish-yellow scales towards
and on sides of 1st, 2nd, and 3rd segments ; dorsum of 1st with
sparse concolorous fine hairs. Sides of 3rd and 4th segments
with elongate brownish-yellow scaly pubescence, which is paler
on the latter ; sides of 5th and 6th segments with numerous
elongate dark brown scales and long black bristles ; posterior part
of side of 6th segment with a conspicuous bunch of elongate
snow-white scales ; 7th segment with black scales only ; all the
dorsal surface of abdomen, not occupied by pale scales, covered
with minute impressed black scales ; sparse fine pale hairs on
about basal half of dorsal surface, replaced on posterior half by
black ones ; hind margins of segments with a row of black bristles.
Venter with a broad transverse band of yellowish scales beyond
middle ; remaining segments with white scales ; whole surface of
venter plentifully covered with long yellowish or yellowish-grey
hairs, with fine black hairs towards tip. Genitalia brownish
yellow, with a circlet of blunt reddish-brown spines and black
pubescence. Leys black, coxae with rather long grey or whitish
pubescence; femora and tibiae with small white or yellowish-grey
scales and rows of black bristles. Winys nearly clear, the brown
suffusion on anterior part in type-specimen limited to subcostal
cell ; in 2nd and 3rd specimens extending from costa up to but
not encroaching on discal cell, dying away towards tip; anterior
cross-vein just before middle of discal cell ; halteres cream-
yellow.
Described from three paratype $ ? in the Indian Museum,
neither one being in sufficiently good condition to regard as the
ultimate type; Simla, 7000-8000 ft., 26. v. 1914 (Copt Evans);
Mussoorie, 6500 ft. (Bond) ; Guindy, Madras (Capt. Patton).
192. Anthrax hottentotta, //., var. claripennis, Brun.
Anthrax hottentotta var. claripennis, Brunetti, Rec. Ind. Mus. ii,
p. 452 (1909).
$ . Head : f rons half as wide at vertex as at level of antennae,
where it is one-third of the head, black, covered (at least on lower
part) with minute yellow impressed scales, and wholly covered
with stiff black pubescence ; face black, wholly covered with dense
elongate yellowish-white scales ; antennae black, 3rd joint con-
siderably elongate ; proboscis obscure, withdrawn ; occiput black,
apparently covered with minute scales, which are yellow behind
the vertex and white near the lower eye-margins. Thorax black,
dorsum (apparently) covered with minute elongate yellow scale-
like depressed pubescence ; anterior margin, sides of dorsum, and
the shoulders with dense elongate upright brownish-yellow scales;
248 BOMBYLIIJXE.
sides with similar scales of a yellowish-white colour or concolorous
long hairs. Scutellum black, hind border (probably whole surface,
more or less) with minute impressed yellow scales (no bristles
present on hind margin, but these have probably been broken off).
Abdomen black, base of 2nd, 3rd, and 4th segments with broad
transverse bands of minute pale yellow impressed scales, the band
on 2nd segment the broadest ; 5th, 6th, and 7th segments nearly
wholly covered with yellowish-white scales, which are longest on
the hind margins ; 7th segment with a thick fringe on hind
margin of very elongate yellowish-white scales ; surface of abdo-
men which is not occupied by yellowish scales, covered with
similar minute black scales. Sides of 1st segment with elongate
yellowish-grey scaly pubescence, which extends as long fine con-
colorous hairs across the dorsum ; sides of 2nd segment with scaly
pubescence as on 1st, this pubescence being continued along
sides of rest of abdomen, but in gradually diminishing quantity.
Tenter obscure, with minute snow-white impressed scales and long
yellowish hairs on basal part, and with white hairs towards tips ;
genitalia brownish yellow. Leys black ; coxae with rather long
yellowish or grey pubescence ; femora and tibiae with minute
yellow and whitish impressed scales, often intermixed ; the white
ones predominating at base of middle femora and below hind
femora. Wings entirely clear, hyaline ; subcostal cell yellowish
brown ; bifurcation of 2nd and 3rd veins exactly in a line with
anterior cross-vein, which latter is just before middle of discal
cell ; halteres pale creamy yellow.
Length, 12% mm.
Described from the unique type in the Indian Museum from
Bhura, in the plains of the Naini Tal District, Western Hima-
layas, 14-17. iv. 1907.
The differences distinguishing this variety from typical A. hot-
tentotta of Europe are the clear wings, the wholly black-haired
frons, the whitish scales on the face, and minor characters.
193. Anthrax paniscus, Rossi.
BiUo paniscus, Rossi, Fauna Etrus. ii, p. 256 (1790).
Anthrax bimaculatus, Macquart, Suites a Buff, i, p. 403 (1834).
Anthrax cingulatus, Meigen, Syst. Besch. ii, p. 145 (1820).
Anthrax hottentottus, Walker, List Dipt, Brit. Mas. ii, p. 259 (1849).
with dark rufous-tawny, nearly black, pubescence, narrowing in
width towards wing-base ; lower part of pleurae with a narrow
band of whitish pubescence, below which the pubescence is black.
Scutellar pubescence concolorous with that of dorsum. Abdomen
black, with long dense brownish-orange pubescence ; sides of 3rd
segment with black pubescence, which is broader anteriorly,
extending across the venter ; isolated long black hairs on hind
margins of 2nd, 3rd, and 4th segments ; 4th and remaining seg-
ments with numerous black hairs, scattered irregularly. Venter
with whitish pubescence, crossed by the transverse band of black
hairs on 3rd segment, and with a similar median longitudinal
band. Legs dull orange ; coxae and trochanters black ; fore
femora (except narrowly at tip), basal half of middle pair, and
basal third (less on upper side) of hind femora black ; tarsi
blackish, but pale on first two joints for a greater or less distance ;
fore coxae with long dense brownish-white pubescence, middle
pair with less dense and darker pubescence, hind pair with blackish
pubescence ; femora with long but not dense blackish pubescence
on about basal half, also with shorter greyish pubescence merging
into adpressed almost scaly pubescence; hind femora with a row
of about eight long black bristles on basal half below, and smaller
ones on apical half ; fore tibiae with three posterior pairs with
four rows of 12 to 20 small black bristles in each row ; fore tibia*
with three, middle pair with five, hind pair with a circlet of small
black spurs ; tarsi minutely bristly ; pulvilli dirty whitish, em-
podium reduced to a minute spine ; claws rather long, black.
Wings clear, with an oblique baso-costal band covering the whole
of the base, and extending distally in gradually but irregularly
diminishing width, ending at about tip of 2nd vein. Roughly
speaking, the hind border of the darkened part emits four short
but not conspicuous extensions with sloping sides respectively
over the alula, over upper branch of 5th vein, up to about middle
of 1st posterior cell, and just before tip of 2nd vein ; alar squamae
brown, margin darker, with long fringe; halteres small, orange,
knob black.
$ . Mainly as the c? . Frons at widest part about half width
of head, narrower above, with long thin yellow hairs and short
adpressed scaly pubescence, which latter is absent on the centre
part and for a transverse space above antennae ; upper part of
irons with short brownish pubescence and long black hairs. The
stripe of dark hair across the lower part of pleurae paler than in . First and 2nd antennal joints brownish
yellow; femora and tibia? wholly bright
brownish yellow; abdominal spines [p. 277.
yellow JlavosptROtut) sp. n.,
Antenna? all black; femora brownish. . . . eupogonatus, Big., p. 278.
* This is so in many species ; 1 cannot vouch personally for the constancy
of the character.
270 ISOMBYLlID.i:.
The distinctions between these alleged species are not at all
clear. I have not seen eui>oyonatus, which may be the same as
socius, the words "legs reddish, femora brownish" being applicable
to a specimen with the femora more brown than black ; whilst
the rest of the description would apply quite well to AValker's
species. My nivalis differs from sowts only in the nearly white
pubescence, and may represent only an individual variation . There
may, therefore, be only two species, sociits and jf?auo«p*no«tt«, with
the addition of an undescribed one in bad condition in the British
Museum.
218. Systcechus socius, Wall:
Bombylius tocius, Walker, Ins. Saund., Dipt. pt. 3, p. 201 (1852).
Systcechus socius, Bruuetti, IJec. Jnd. Mus. ii, p. 458 (1909).
cT 2 • Head blackish grey ; eyes in <5 distinctly but narrowly
separated at nearest point of approximation ; vertex with
numerous short stiff hairs; ocelli reddish; frons with long
black bristly pubescence ; face in tf with a few long whitish
scales, more or less depressed, amidst numerous long dark brown
bristles ; in $ , with a considerable quantity of long bristle-like
whitish scales extending from eye to eye; mouth-opening with
dense long blackish-brown scale-like bristles, some whitish pubes-
cence around lower part of mouth-opening, which is itself whitish ;
lower part of head whitish grey, with dense elongate very thin
white scaly pubescence ; antennae black, 1st joint with long dark
brown bristles ; probo-cis black, as long as the head and thorax
(not scutellum) together; back of head with very dense long
bristle-like scales, varying from pale to bright yellow ; lower eye-
margins with whitish scales. Thorax black ; dorsum, pleurae, and
scutellum wholly covered with dense elongate scales forming a
close thick pubescence, varying from nearly whitish to deep
yellow, almost orange, that below the shoulders being generally
deeper in shade than that on the thoracic dorsum ; scutellum with
hind part always more or less reddish brown, sometimes practically
entirely so ; a row of yellow spines on hind margin. Abdomen
black; densely covered \\ith similar bristle-like pubescence to
that of the thorax ; a row of black spines near hind border of
each segment from the 3rd onwards ; venter black, with long
pale yellow hair-like scale." ; base of some of the middle segments
often reddish brown. Legs : femora black, with minute whitish
scales ; tibiae paler or darker brown, sometimes blackish at tips,
especially the hind pair; tarsi black, underside pale brown.
\V-ings pale grey; base, costal cell, and the veins on basal half
brownish ; halteres cream-yellow.
Length, 7-11 mm.
Redescribed from several of each sex in the British and Indian
Museums, the Pusa and my own collections.
This species extends from the Himalayas to Ceylon: Kashmir,
1915 (II. T. Pease); Dehra Dun, xi." 1907 (Li. -Col. ll'i/ville
81-STGSOHUS. 277
Thomson], iv. 3914, common (Ghatterjee) ; Theog, Simla Hills,
14. v. 1909; Ivumaon, 18 & 27. v. 19 12; Bhowali, 5700 ft.,
iii. 1910 (CluMerjee) ; Runjit Valley, Sikkim, v. 1894 (Lt.-CoL
C. T. JBingJtamf: Kangra Valley, 4500 ft., x. 1899 (Dudgeon);
Ooimbatore, 30. v. 1912 ; Hadagalli, Ceylon, x. & xi. 1911.
Type in the British Museum.
The colour of the pubescence is so variable that I am inclined
to think my nivalis may be an extreme form. The proboscis in
nivalis is distinctly longer than from the tip of the frons to the
tip of the scutellurri.
Mr. T. B. Fletcher finds that the larvae prey on the eggs of
the Deccan grasshopper.
219. Systcechus nivalis, B,-un.
Sytt&dau nivalis, Brunetti, Rec. Ind. Mus. vii, p. 472 (1912).
d1 . Head with frons and face blackish grey, darker above, both
thickly covered with long black bristly hairs. Antennas rather
long, black ; 1st and 3rd joints subequal, each abqut three times
iis long as the 2nd ; first two joints cylindrical, 3rd much more
slender on apical half. Proboscis Arery long, labium reaching to
middle of abdomen (actually bent backwards to this distance
below the body in the rype), labrum shorter; palpi extremely
short and slender ; underside of head dark grey, with whitisli
hairs. Thorax and abdomen closely covered with pale yellowish
grey, almost white, bristly pubescence, which is more yellowish
on anterior part of former and whitish on pleurae and apical part
of abdomen ; underside of thorax and abdomen with whitish hair.
Legs black ; tibi* from the base, for the major part, brownish
yellow. Wings pale grey, costal cell a little darker grey ; base of
wing a little yellowish.
Length, 9 mm.
Described from a unique cf in the Indian Museum from
Airadeo, Kumaon Distr., Western Himalayas, 31. v. 3912 (Imms).
Possibly a variety of sodas, AValk.
220. Systcechus flavospinosus, sp. nov.
$ . This species differs from S. sodus only as follows :— Face
wholly covered with long dense silky yellowish scales, no black
bristles ; 1st and 2nd antennal joints brownish yellow, 1st with
some long pale hairs ; 3rd rather shorter and broader basally than
in sodas; scutellum almost entirely b rick -red ; spines on hind
margins of abdominal segments yellow, not black ; femora and
tibiee wholly bright brownish yellow, tarsi gradually darkening to
brown at tips.
Length, 7 mm.
One $ in the British Museum from the South Shan Status,
Upper Burma, 4000 ft., xi. 1899 (U.-Col. C. T. Bingham).
278 BOMBTLlIDyE.
221. Systceclms eupogonatus, Big.
Systcechus eupogonatus, Bigot, Ann. Soc. Ent. France, Ixi, p. 365
(1892).
" d1 . Antennae black, 1st joint covered above with long blackish
hair, and below grey ; face dull vellow, with grey hairs above, hairs
on lower part long, black, and thick. Frons black, with long black
hairs. All the body covered with thick pale yellow pubescence.
Scutellurn black, reddish at tip; halteres tawny, knobs whitish.
Legs reddish, femora brownish, with grey tomenturu. Wings
nearly clear.
"§. Frons grey, with grev pubescence; proboscis a little
longer than head and thorax together. India." (Bigot.)
Length, 10 mm.
Types in the Bigot collection.
Genus ANASTOECHUS, O.-S.
Anastccchtis, Osten-Sacken, 'Western Diptera,' p. 251 (1877).
GENOTYPE, Bombylius nitidulus, F. (as barbattts, O.-S.) (Europe);
by original designation.
This genus differs from Bomlylius by both basal cells being of
equal length. The dense furry pubescence of the underside of the
head distinguishes it from SystoKclms (which also has the basal
cells subequal in length), but not from Bombylius.
llanye. Europe, Asia, North Africa, North America, and India.
Only one Oriental species has been recorded.
222. Anastcechus longirostris,
Anasta-chus longirostris, Wulp, Notes Levd. Mus. vii,p. 85 (1885).
$ . " Small, black ; pubescence of head and body white, of frons
brown ; abdomen with several black bristles ; proboscis longer
than head and thorax ; legs pale yellow ; wings grey, base and
costa yellowish ; 5| mm.
" Near the American A. barbalus, O.-S. The thorax and
abdomen more slender; head broader than thorax, with dense
long white hairs on face and dark brown or blackish on frons.
Antennas black ; terminal stvle of latter as long as the pyriform
3rd joint ; end of style a little thickened. Proboscis black, as
long as two-thirds of body. Thorax and abdomen thickly clothed
with long yellowish hairs, with u. white reflection, amidst which
are some long black bristles on the sides of the abdomen. Legs
and halteres pale rufous. Wings greyish, yellowish at base
and costa; at the beginning of the ccsta is "a comb of black-
tipped yellowish hairs. Venation as in A. barbatus. One $
from the Himalayas (Felder)" (van der Wulp.}
This species is unknown to me.
279
Genus GERON, Mg.
Geran, Meigen, Syst. Besch. ii, p. 223 (1820).
GENOTYPE, Bombylius hybridus, Mg. (Europe) ; bv original
designation.
Head semicircular, narrower than thorax and set low down on
it ; epistome short ; mouth-opening large ; proboscis prominent,
horny, curved, with narrow labella ; eyes in c? contiguous, widely
.separated in $ . Antennae porrect, approximate at base, longer
than head ; 1st joint long, cylindrical ; 2nd very short ; 3rd as
long as 1st, spindle-shaped. Thorax oval, considerably arched,
projecting forward ; thinly pilose or with microscopic tomentum ;
scutellum triangular. Abdomen extremely short, conical, tip
pointed, thinly pilose. Legs slender, minutely pubescent. Winys
with 3rd longitudinal vein forked ; anterior cross-vein beyond
middle of discalcell, erect ; two submarginal cells ; three posterior
cells ; anal cell nearly reaching border of wing; anal vein attaining
the border.
Range. World-wide.
Life-history unknown.
The two Indian species are thus distinguished : —
Legs pale brown ; tibiae paler ; length 2£-4 mm. argentifrons, Brim.
Legs pale tawny; coxse, tips of femora, and
hind tarsi black ; length 5 mm albescens, Brun.
223. Geron argentifrons, Brun.
Geron argentifrons, Brunetti, Rec. Ind. Mus. ii, p. 482 (1909).
d1 . Head with eyes contiguous from the very reduced raised
vertex to antennae, facets of moderate and uniform size ; vertex
with several long black hairs of variable lengths ; ocelli well
separated, pale brown ; frons covered with dull yellowish oblong
scales, lying flat on surface, and with a second clothing of silvery-
white scales ; face bluish white, with silvery reflections seen from
above, as are also the lower eye-orbits ; some long black hairs on
face ; upper side of mouth-opening yellowish ; occiput black,
with long brownish-grey liairs. Antennas black, tip of 1st and
2nd joints sometimes very narrowly whitish, 1st joint about double
as long as 2nd ; 3rd distinctly longer than first two together ;
basal joints occasionally pale. Thorax black, with long brownish-
grey or yellowish-grey pubescence, through which can be seen
a lower clothing of very short yellow scale-like hairs ; pleurae
bluish grey, with whitish-grey pubescence. Abdomen clorsally like
thorax, sides with numerous silvery-white scale-like hairs ;
apparently the whole dorsum normally covered with long hairs,
which maybe thicker and longer at the sides ; venter like dorsum,
except that the long hairs are a tittle shorter. Legs normally dark
brown or blackish ; tibiae always paler, varying in shade from
yellowish to moderately dark reddish brown ; femora apparently
280 BO11BYL1IJL*.
(and probably coxae also) covered with small silvery-white scales :
tibiae with small yellowish scales, short black spines and a circlet
of them at tips, which are a little darker ; tarsi black, with usual
pubescence, metatarsi partly or almost wholly pale. Wings quite
clear, iridescent; veins pale yellow; squamae pale yellow, with
short hairs ; halteres large and prominent, clubs egg-shaped, from
nearly white to bright yellow.
$ . Frons at vertex forming one-fourth the width of the head,
covered with bluish-grey dust and with very small yellow or
yellowish-white scales ; a cluster of silvery -white scales around
base of antennae ; face as in d1 , except that the long hairs are
white, not black. Underside of first two antennal joints pale.
Posterior femora and apical part of fore pair, also all the tibia?,
pale yellowish.
Length, 2|-4 mm.
Redescribed from type c? from Lahore, 9. v. 1903 (Annandalt),
and three other <5 c? , also three $ $ , all (except type J ) from
Pusa, Bengal (Hewlett, etc.), where it is common, March to May.
Also from Alinora, Kumaon Distr., 5500 ft., v-vi. 1911 (Paiva) ;
Parel, Bombay Presid., 26. xi. 1909. In the British Museum
from four places in Ceylon, Trincomali, x. 1890, and Nilavelli,
31.viii.1890 (Col. Yerbury); Pundaluoya, ii. 1898 (Green);
Madulsima, 19. v. 1908 (T. E. Fletcher).
This species is parasitic on caterpillars found under the bark of
the sissoo-tree (Dalbergia sissoo) ; it also occurs on the pipal.
Types in the Indian Museum.
224. Geron albescens, Brun.
Geron albescens, Brunetti, Rec. Ind. Mus. iii, p. 229 (1909).
<5 . Head black ; eyes contiguous almost to antennae, frontal
triangle extremely small, upper facets slightly larger than lower
ones ; vertex elevated, small, with a few short black hairs ; ocelli
Fig. 17.— Geron albescens, Brun.; lateral view of head.
red; mouth black, grey bordered; face small, squarish, black,
with some snowy-white scales on each side of base of antennae ;
GERON.— USIA. 281
proboscis black ; antennae black, relative length of joints as in
argent ifrons, tips of first two joints narrowly pale, with greyish
hairs ; occiput black, with long yellow hairs above and greyish
ones below ; hind margin of eyes with small white scales. Thorax
velvet-black, dorsum covered with bright yellowish hairs, which
are greyish on the grey shoulders ; underside ash-grey, with long
grey hairs and some grey scales ; scutellum black, yellow-haired.
Abdomen black, with minute bright yellow fine scales and hairs ;
hind margins of segments with long deflexed grey hairs ; whole
dorsum with long thin greyish pubescence, which is denser at the
sides ; venter wholly covered with small silvery-white scales and
long grey hairs. Legs with coxae and fore femora black, former
with grey hairs ; posterior femora and all tibiae pale tawny ; tarsi
black, middle metatarsi more or less pale at base ; legs almost
Fig. 18. — Geron albescens, Brun., wing.
bare except for a few short hairs below femora and the usual
minute tibial bristles. Wings quite clear, veins pale yellowish on
basal half; halteres yellowish, knobs bright pale yellow.
Length, 5-8 mm.
Redescribed from the original type from Balugaon, Puri Distr.,
Orissa, 14. xii. 1908 (Gaunter), and a second J from Tambi,
Satara Distr.; Bombay, 2100 ft., 22-26. iv. 1912 (Gravely), both
in the Indian Museum.
^.-The only definite character by which this species differs from
argentifrons seems to be the pale posterior femora in the J and
the larger size.
Genus USIA, Latr.
r/«Vi, Latreille, Hist. Nat. Crust. Ins. xiv, p. 300 (1804).
Volucella, Fabricius .(nee Geoff.), Ent. Syst. iv, p. 412 (1794); id.,
Syst. Antl. p. 114 (1805) ; Meigen, Klass. i, p. 194 (1804).
GENOTYPE, Volucella florea, F. (Europe) ; by Rondani's desig-
nation.
Rather small bare or nearly bare flies, sometimes of metallic
ground-colour, sometimes with yellow and black markings. Head
hemispherical, small, set rather low on the thorax; epistome
short, proboscis prominent, at the base membranous, apical part
horny with extremely narrow labella ; palpi short, united to the
basal part of the membranous portion, which is entirely with-
drawable within the mouth-border. Antennas short, porrecl,
hardly longer than head, approximate at base ; the first two joints
subequal ; 3rd elongate, spindle-shaped above, blunt, with spines
282
BOMBi'LllD-E.
and a short apical style. Eyes in both sexes normally distant,
but sometimes contiguous in d" . Thorax oval or nearly rounded,
arched ; scutellmn short and broad. Abdomen of seven segments,
short oval, broader than thorax. Legs rather stout, bare, without
spicules. Wings in rest, halt' open ; venation as in Geron.
Range. Europe, Asia, North Africa.
Life-history unknown.
The two Indian species are distinguished as follows : —
Thorax with two or three elongate light grey
spots on anterior margin stdophila, Brun.
Thorax with anterior margin broadly light
grey, produced posteriorly as four short
stripes maryiiuita, Brun.
225. Usia sedophila, Brun.
Usia sedophila, Brunetti, Rec. lud. Mus. iii, p. 227 (1909).
cJ. Head: vertex prominent, black, shining, with a few erect
long hairs ; ocelli distinct, whitish ; eyes black, bare, contiguous
for more than half the distance from vertex to antennae ; facets in
front and above distinctly larger than those behind and below,
but no distinct dividing line. Antennae black, bare, except for an
occasional hair ; first two joints short ; 3rd somewhat flattened,
but thick and slightly curved, elongate, and with a peculiar notch
towards the tip on upper side, in which is a single hair. Proboscis
about twice the length of the head, black, cylindrical, stiff, straight,
bent upwards at tip, bare, except for a few minute hairs at tip ;
face dark greyish, with some stiff black hairs on each cheek, and
Fig. 19. — Usia sedophila, Brun., wing.
a long elevated shining black callus between the median line and
the eye-margins ; back of head black, not produced behind margins
of eyes, with long sparse black hairs, continued on underside of
head. Thorax dull black, velvety on dorsum, with some black and
pale hairs towards the sides, and three whitish-grey elongate spots
on anterior border, of which the middle one is often absent, and
the outer ones are occasionally indistinct or absent also; lower
part of thorax more or less dark grey ; scutellum black, with
some erect light tawny hairs. Abdomen dull black, conical ; 1st
segment narrow, wholly black ; the remaining six broader, but of
diminishing length and breadth to the tip, and with a rather
bright yellow narrow posterior border to each segment ; dorsum
USIA. 283
comparatively bare, but the sides of the abdomen with pale yellow
hairs. Genitalia small, inconspicuous, conical, blackish. Venter
blackish, posterior margins of segments whitish grey. Leys
uniformly black; very shortly pubescent ; femora with some soft
hairs; hind tibiae with short bristles. IVinr/s nearly clear ; sub-
costal cell pale yellow ; venation normal ; halteres dark brown or
blackish, knobs large.
$. Head: eyes separated by a wide frons equalling one-third
width of head ; upper half of frons, as also whole back of head,
greyish yellow, the latter with black hairs as in c? ; upper half of
frons with a row of hairs near margin of eyes; vertical triangle
distinct, dark , two narrow fuscous vertical lines from the vertical
triangle to an irregular transverse dark streak dividing the upper
from the lower half of the frons, the latter part being more ash-
grey and unmarked ; lower part of the head greyish ; proboscis and
mouth larger than in J,thus narrowing the cheeks considerably.
Thorax yellowish grey, with a median pair of narrow fuscous
lines, with a broader outer stripe, more or less divided into three
spots, the anterior one small and conical, the other two more
elongate ; also an indistinct central line between the median pair
of narrow ones, and a small spot above the insertion of the wings;
sides of thorax cinereous grey ; scutellum yellowish grey, with
some yellow hairs as in tf . Abdomen black, bare ; posterior
margin of segments rather broadly yellow ; sides with a little less
hair than in rf ; venter blackish ; ovipositor pale yellow, in-
conspicuous. Leys and wings as in c? .
Length, 2f-3| mm.
Described from several of both sexes taken by Dr. Annaudale
at Simla, 16. v. 1909, where it was common on ' Sedum roguUtuni
(white stonecrop) ; the species was not seen anywhere except on
this plant. Phagu, 18-21. v. 1916 (Annandale).
Types in the Indian Museum.
226. Usia marginata, Brun.
Usia marginata, Brunetti, Eec. Ind. Mus. iii, p. 228 (1909).
rf. Head with eyes contiguous for a shorter space than in
sedophila, black, bare ; upper facets larger than lower, and more
sharply defined from one another than in sedophila ; frons ash-
grey, small, with a few black hairs ; vertex prominent, with some
black hairs, ocelli distinct : front margins of eyes and cheeks
with long black hair, which is considerably thicker on the under-
side, and which extends over the whole of the back of the head.
Antennae as in sedophila, but the third joint with a pale ring at
the base, giving the appearance of a minute basal joint, and the
tip bears on its upper side a minute style-like process, just beyond
the subapical depression ; 1st joint with some bristly black hairs
2nd with one or two minute hairs, 3rd bare. Thorax (lull
black, almost velvety, covered lightly on dorsutn, and more thickly
284 UOMBi'MIDJE.
in front and at sides with long black hairs ; anterior margin of
thorax broadly whitish grey, which in the centre is produced
posteriorly into four narrow stripes, of which the outer pair just
reach the suture, the inner pair being very slightly shorter ; sides
of thorax black, with long black hairs, and the whitish-grey colour
of the anterior margin of dorsum extends downwards somewhat
below the shoulders. Abdomen dull black, posterior margins of
segments narrowly yellowish, with the colour continued over the
sides and across the venter ; a row of grey hairs on posterior
border of segments, and the sides of the abdomen with rather
thick whitish-grey hairs ; venter concolorous. Leas black, minutely
pubescent ; femora with rather long soft black hairs on upper and
lower sides ; hind tibi«3 very shortly bristly. Wings nearly clear ;
subcostal cell very pale yellow ; halteres pale brown, clubs chalk-
white.
Length, 4 mm. »
Described from one J taken by Dr. Annandale at Simla,
10. v. 1909, in company with the previous species. A second
specimen with the same data; both in the Indian Museum.
Genus PHTHIRIA, Mrj.
Phthiiia, Meigeii, Illig. Mag. ii, p. 268 (1803).
Megapaipw, Macquart, Hist. Nat. Ins., Dipt, i, p. 394 (1834).
Dasypalpus, Macquart, Dipt. Exot. ii, pt. 1, p. 112 (1840).
Cyclorhynchus, Macquart, Dipt. Exot. ii, pt. ], p. 114 (1840) preocc.
Phtyria, Rondani, Arch. Zool. Modena, hi, p. 05 (1863).
PcBfilognathia, Juennicke, Abh. Senck. Nat. Ges. vi, p. 3/30 (1863).
GENOTYPE, Bombylius pulicariua, Mikan (Europe) ; by original
designation.
Head in d1 nearly ' semicircular, in $ nearly circular, not
broader than high viewed from in front, about as wide as thorax ;
face broad but short down the middle, because the large and
rather wide mouth-opening extends up between the wide side
cheeks ; frons in $ broad, depressed in the middle ; three ocelli ;
eyes in tf contiguous or subcontiguous, in 5 wide apart. Pro-
boscis long, thin, almost bristle-like, directed forwards in a slight
arch, labella very small ; palpi rather short and thin. Antenna?
porrect, shorter than head, approximate at base; 1st and 2nd
joints short, subequal in length ; 3rd elongate, peg-shaped, Avith a
terminal style (sometimes indistinct). Thorax oval, longer than
broad, slightly arched, in d1 with fairly long erect dense pubes-
cence, without bristles; in 5 with only short depressed pubescence;
pleura? with only slight pubescence, except for a rather dense tuft
on the mesopleurae ; metapleurae bare ; scutellum large, semi-
circular. Abdomen 7-segmented, conical in <5 , ovate and rather
flattened in $ ; pubesceuce as on the thorax in 5 . Leys long
and thin, entirely without bristles except for some tiny spicules on
the hind tibia3. Wings longer than abdomen ; 1 st basal cell
pimiiaiA. 285
considerably longer than 2nd, 3rd vein forked, the 2nd sub-
margiual cell elengate ; anterior cross-vein at about middle of
discal cell ; anal cell closed just before \viug-border.
Range. World-wide.
Life-history unknown.
The adults occur on bare patches of ground among low-growing
flowers, and on Composite.
227. Phthiria gracilis, Wall:
Phthina gracilis, Walker, Ins. Saund., Dipt. pt. 3, p. 194 (1852).
<5 $ . Head in c? b)ack ; vertex very small, dark grey, with
three or four short black hairs; ocelli dark ruby-red, shining;
eyes closely contiguous for nearly all the distance to the small
protruding triangular dark grey frons, which bears a few black
hairs; mouth-opening extremely large, extending from immediately
below base of antennae to lower part of head, black; face and
under part of head black, both with long black hairs : antennae
black ; 1st and 2nd joints as long as wide, subglobular, with two
or three short black hairs each, 3rd joint narrower, flattened,
about two and a half times as long as first two together, with
microscopic depressed pubescence and a short style-like prolonga-
tion at tip on upper side; proboscis as long as whole body, black,
slender, gently curved ; palpi half as long as head, porrect, black.
Head in $ bright orange-yellow ; frons at vertex about half as
Fig. 20.— Phthiria gracilis, Walk., wing.
wide as at level of antenna3, where it forms nearly one-third the
width of the head ; a broad black stripe from vertex to above
antennae, sometimes wholly interrupted in middle ; ocelli distinct ;
sides of face between antenna? and eyes blackish ; occiput black,
broadly orange behind eye-margins, the colour not extending
behind the vertex. Thora.r in d1 dark blackish brown, with a
little short dark brown pubescenbe, which is longer towards the
sides of thedorsum; scutellum concolorous. In 5 greenish grey,
lower margin of dorsum rather broadly though somewhat irregu-
larly pale yellowish ; scutellum yellow, with a small blackish spot
at base. Abdomen in <5 black, with long black hairs, hind margins
of basal segments more or less narrowly yellow. In $ yellowish,
with short, brown, not very obvious pubescence ; base of segments
286 BOMBYLIIDJE.
more or less broadly blackish, the colour confined to the middle of
the disc on the basal ones, but extending to nearly or quite the
full width in the others. Venter and genitalia in both sexes
black, with some soft hairs. Legsm , and usually adpressed on basal segments. Genitalia not
prominent, but of'ten visible ; ovipositor with a circlet of strong
black spines. Legs moderately long, slender; some black bristles
on coxaB and anterior femora ; tibiae with rows of small bristles and
a circlet of spines at tip. Wings with normal Therevid venation ;
4th posterior cell generally closed or narrowly open.
Range. World-wide, but apparently better represented in
Europe and North America.
Life-history. Partly known ; larvae very active, occurring in
damp earth, under stones, and in similar places ; possibly they
feed, at least occasionally, on Lepidopterous larvae.
Only four species are known for certain from India ; these may
be separated as follows : —
Antemue mainly bright brownish yellow. . kempi, sp. n., p. 298.
Antennae wholly black.
Legs all blackish niyella, Wied., p. 299.
298 THEREVIDJE.
Tibiae brownish yellow.
Posterior margins of abdominal seg-
ments narrowly and uniformly
yellow jlauolineata, Brim., p. 299.
Posterior margins of abdominal seg-
ments ash-grey, much broader
towards sides bilineata, Bruu., p. 300.
Four other species are probably from India, but require con-
firmation.
233. Thereva kempi, sp. nov.
<5 . Head with eyes contiguous for about one-third of the distance
from vertex to antennae, with a dull greenish tinge ; vertex
yellow-dusted, with a few black hairs ; ocelli shining black ; frons
yellow-dusted, rather closely punctured, with long black bristly
hairs on upper part, continuing down the sides as far as level of
antennae ; rest of frons and whole of face and cheeks covered with
dense bright chrome-yellow bristly hairs. Antennae orange ;
upper part of 3rd joint, except at base, blackish, as is also the
style ; 1st joint with conspicuous long black bristles ; 2nd with
an incomplete circular band of smaller black spines ; 3rd with a
little whitish pubescence and a few small bristles at base above.
Proboscis orange ; occiput yellowish, on upper half with short
yellow pubescence and a row of strong black bristles along upper
margin ; lower part behind eyes whitish, with long black hairs.
Thorax black, covered with rather dull yellow tomentum and, at
least towards sides of dorsum, with short bright yellow somewhat
coarse pubescence ; the whole dorsum covered with long fine black
hairs, whicli are densest in the centre. Scutellum yellow-dusted,
darker in centre and at base, with yellow pubescence with a con-
siderable number of long black hairs intermixed ; hind margin
with four long bristles. Pleurae light grey, closely covered with
bright yellow pubescence, which is mainly replaced on the sterno-
pleura by white hairs. Abdomen with 1st segment dark grey, with
sparse soft black hairs ; rest of segments deep black, more or less
shining at base, with a well-marked bright golden-yellow hind
margin to each, the colour extending forwards towards the sides
as far as the middle of each segment or beyond ; whole dorsum
covered with dense black or yellow pubescence according to the
ground-colour. Venter light grey, appearing darker in certain
lights, with distinct pale yellowish-white hind margins to
segments ; the whole surface with sparse soft bright yellow hairs.
Legs : coxae light grey, with rather long yellow pubescence ;
femora black, anterior pairs with similar pubescence ; hind pair
nearly bare, but with a row of seven or eight black bristles below;
tibiae brownish yellow, blackish at tips, with the usual bristles
and microscopic pubescence; tarsi dirty brownish yellow, tip of
1st joint, apical third of 2nd, base of 3rd and 4th narrowly, and
all 5th black. Winrjs pale grey, highly iridescent ; 4th posterior
cell closed slightly before the border ; stigma dark brown ; halteres
blackish.
Length, 12 mm.
T1ILREVA. 299
Described from a perfect unique d in the Indian Museum from
Shillong, Assam, 5500 to 6400 it., 29.viii-5.ix.1915 (Kemp}.
A. very beautiful species, unlike any other Indian form; it must
bear considerable resemblance to T. fuJva, Mg., of Europe.
234. Thereva nigella, Wied.
Thereva niyella, Wiedemann, Auss. Zweifl. i, p. 23:2 (1828).
$. "Antennae blackish, in certain ligbts whitish. Epistome,
beard, and moustache white ; f rons and back of head with a white
shimmer. The rather rubbed thorax shows faint traces of stripes ;
the side margins broadly whitish. Abdomen with some white
hairs on the side margins, the segments hardly whitish. Wings
quite clear ; halteres brown, with white knobs. Legs blackish,
with short white hairs. Type in Westermann's collection.
Tranquebar. Length 4g lines." (Wiedemunn.}
235. Thereva flavolineata, Brun.
Thereva jlavolineata, Brunetti, Eec. Ind. Mus. vii, p. 480 (1912).
Head: frons small, black, with black hairs; ocelli red; eye-
facets larger in front ; face grey, with long stiff black hairs above
the antennas and with long greyish hairs below them ; antennae
black, microscopically grey-dusted; oral orifice reddish yellow; back
of head dead grey, with greyish hairs ; vertex and upper border of
eyes with black hairs. Thorax dark brown, with two well-separated,
moderately wide, chrome-yellow stripes ; dorsum covered with
mixed brown and yellow hairs, the former mainly on the darker
parts ; sides of thorax blackish grey in front of wings, ashy grey
on lower part, with shaggy dirty white hairs in front of and below
shoulders, and in front of wing-base; some yellow hairs above
insertion of wings. A row of three bristles between the humerus
and transverse suture, two supra-alar bristles (others possibly
broken off). Scutellum yellowioh, with a brownish basal triangle,
dorsum with brown and grey hairs ; two preapical well-separated
bristles. Abdomen blackish grey, posterior margins of segments
very narrowly yellow, but more widely so on 3rd and 4th : 1st
segment wholly, 2nd, 3rd, and 4th wholly (except posteriorly at
the sides), 5th at base, black; dorsum of abdomen with black
hairs, which are thickest and longest on the black parts; sides
with moderately long whitish-yellow hairs ; posterior margins
of segments with very short, similarly coloured hairs. Venter
bluish grey, posterior margins of segments more broadly yellow,
especially on 3rd and 4th segments ; the whole surface with spit
white hairs. Genitalia distinct, shining brown, with brownish
hairs on upper, and whitish hairs on lower parts. Legs (hind pair
wanting): coxas and femora blackish grey, the former with long
greyish-white hairs in front; the latter with grey hairs below on
proximal half, blacker hairs on discal half ; tibia? reddish yellow,
300 THEREVIDjE.
blackish at tip, microscopically pubescent, with numerous rather
short spines ; tarsi darker. Wings nearly clear, veins brownish
yellow ; tegulae dirty white ; halteres brown and yellow.
Described from a d in the Indian Museum from Shi Hong.
A second specimen, also from Shillong, 4900 ft., 10-12. x. 1914
(Kemp), is identical, except that the abdominal segments have no
pale sides, as in the type, but the hind borders of the 2nd segment
and onwards are very narrowly yellowish towards each side, and
bear thereon some rather long yellowish hairs.
236. Thereva bilineata, Brun.
Thereva bilineata, Brunetti, Rec. Ind. Mus. xiii, p. 78 (1917).
§ . Head with frons at vertex distinctly more than one-third
width of head, at level of antennae over half the head ; its upper
two-thirds yellowish brown, the rest (with face) ash-grey ; whole
frons with scattered black hairs of moderate length ; face wholly
covered with long pendent whitish-grey hairs, extending over
cheeks and lower part of occiput ; some black hairs below or around
mouth ; proboscis reddish brown ; antennae black, first two joints
with strong black bristles, 3rd as long as 1st ; vertical triangle barely
raised ; a row of rather strong bristles along upper occipital margin,
a second row a short distance behind the first. Thorax dark
brown, nearly black, with two distinct, yellowish-white, well-
separated dorsal stripes from anterior to hind margin and minute
sparse yellowish pubescence. A row of five powerful spines
behind humerus, extending towards wing-base; tw^o supra-alar
(postsutural), one post-alar, a large and lesser one on each side
margin of scutellum and a pair of apical scutellar decussate nearly
perpendicular ones. Pleurse ash-grev, with long whitish-grey
pubescence. Scutellum ash-grey, with a large brown spot at base ;
hind margin with soft yellow hairs as well as the bristles. Abdomen
black ; 1st segment grey, with yellow pubescence on hind margin
and whitish hairs at sides ; rest of segments with distinct ash-grey
hind borders, which are narrowest in middle, bearing some yellow
or yellowish-white pubescence; sides of 2nd and 3rd segments
with whitish hairs; rest of dorsuin and at sides with short black
pubescence; 7th segment shining chestnut-brown; genitalia the
same, marked with yellow, fairly prominent; venter black, all
segments except 1st with well-defined bare hind margins. Leys:
femora black, with depressed white pubescence and a short row of
bristles in middle of underside ; tibiae brownish yellow, darker at
tips ; fore tibiae with three (inner pair wanting), posterior pairs
with four rows of strong spines ; tarsi brownish, yellow or dark
brown, joints paler at base, shortly pubescent. Wings pale grey,
normal ; halteres black.
Length, 11 1 mm.
Described from a unique $ in the Indian Museum from Theog.
Simla Distr., 8000 ft., l.v. 1907 (Annandale).
This species bears some resemblance to my fotvolineala.
THEREVA. 301
237. Thereva cylindrica, Walk.
Thereva cylindrica, Walker, List Dipt. Brit.Mus. i, p. 224 (1848).
<3 . " Body long and narrow ; head white, hoary behind, with
a few black bristles along the hind borders of the eyes, clothed
beneath with white hairs ; eyes green and red ; feelers black; 1st
joint clothed with black hairs; 2nd joint and base of the 3rd
tawny ; mouth piceous ; chest hoary, clothed with hoary down, and
with a few black bristles on each side, where there is a white
stripe, and it has also a broad yellowish-brown stripe in the
middle ; breast white and clothed with white hairs ; abdomen
brown, covered with silvery glance, yellowish brown towards the
tip, where it is clothed with dark tawny hairs ; hind borders of
tli3 segments pale tawny and adorned with fringes of white hairs ;
legs grey ; hips white ; wings slightly grey, tawny along the fore
border ; brands brown and narrow; veins piceous, slightly bordered
with dark tawny ; poisers piceous, with dark tawny knobs.
" Length of the body 5 lines, of the wings 8 lines. East Indies."
(Walker.)
Though described from the " East Indies," the type in the
British Museum bears the simple label " Ind.'"', which is considered
to signify India proper.
238. Thereva indica, Walk.
Thereva indica, Walker, Ins. Saund., Dipt. pt. 3, p. 159 (1852).
$. "Body black; head shining above, tinged with grey and
clothed with hoary hairs beneath ; sucker and feelers black ; breast
tinged with grey; abdomen spindle-shaped, very much longer
than the chest ; hind borders of the segments and underside
tinged with grey ; legs tawny, beset with a few black bristles ;
claws black ; wings grey ; brands brown ; wing-ribs pitchy ; veins
black, pitchy at the base ; poisers pitchy.
" Length of the body 5 lines ; of the wings 9 lines. East
India." (Walker.}
239. Thereva persequa, Walk.
Thereva persequa, Walker, Ins. Saund., Dipt. pt. 3, p. 158 (1852).
2 . " Body black, head very shining above, silvery white in
front and beneath, where it is clothed with white hairs ; sucker
black; palpi tinged with grey, thickly clothed with white hairs;
chest grev, dull, adorned on each side with a pair of hoary stripes,
a pitchy tinge between each pair ; sides and scutcheon armed with
black bristles ; breast hoary, clotlied with white hairs ; abdomen
black, shining, clothed with very short black hairs, slightly
decreasing in breadth from the base to the tip, near twice the
length of the chest ; hind borders of the 1st and 2nd segments
fringed with short white hairs ; tip armed with a circlet of black
spines ; legs ferruginous ; hips black, tinged with hoary, clothed
with white hairs ; shanks beset with black bristles ; feet black,
pitchy at the base; foot-cushions pitchy: wings grey, slightly
tinged with brown along the borders of the veins; poisers tawny,
their knobs black, with a hoary tinge.
" Length of the body 3^ lines, of the wings 5 lines. East India."
(Walker.)
240. Thereva invaria, Walk.
Thereva invaria, Walker, Ins. Saund., Dipt. pt. 3, p. 159 (1852).
2 . " Body black, clothed with black hairs ; head clothed
beneath with white hairs ; sucker pitchy, clothed with tawny
hairs ; breast and underside of abdomen grey, the former partly
clothed with whitish hairs ; abdomen spindle-shaped, very much
longer than the chest ; hind borders of the segments lurid ; a large
tuft of white hairs on each side of the base ; legs black, clothed
with short black hairs, beset with a few black bristles ; foot-
cushions pitchy; wings slightly grey; wing-ribs pitchy; veins
black, strongly marked, pitchy towards the base ; brands brown ;%
poisers pitchy.
" Length of the body 5 lines, of the wings 9 lines. East India."
(Walker.)
Genus PSILOCEPHALA, Zctt.
Psilocephala, Zetterstedt, Ins. Lapp. p. 625, note (1838) ; id., Dipt.
Scand. i, p. 19 (1838) (Psilocehala, lapsus).
GENOTYPE, Bibio imberbis, Fin. (Sweden) ; by Coquillett's
designation (1910).*
This genus differs from Thereva in the bare face ; and from a
nearly allied European genus, Dialineura, which may probably
occur in India, it is known by the 1st antennal joint being no
broader than the 2nd and not more bristlv than usual. The
alleged character that the 1st joint is shorter than the 2nd and
3rd combined, as compared with its being longer than the 2nd and
3rd together in Dialineura, seems an uncertain one.
The species are by no means almost bare, as stated by Verrall.
Range. Europe, Asia, North, Central, and South America,
North and South Africa, Madagascar, Sokotra.
Life-history. Probably similar to that of Thereva.
Five species are definitely known from India or Ceylon, with
one additional species, sequens, Walk., requiring confirmation.
These five are difficult to tabulate, from the general similarity of
the descriptions, although when placed side by side the species
may appear quite distinct. The colour of the legs, though
apparently an unreliable character, is perhaps as constant a one
as any, but the present table is, at best, only a preliminary one.f
* It would have been better to have selected the well-known European
urdea, F., as genotype.
t It is advisable in this family to compile a separate table for each sex, but
tho females are known of only three out of the five species.
PSILOCEPHALA.. 303
Table of Species.
1. Femora dark brown ; all tibiae and tarsi
pale yellowish brown ; (thorax pre-
sumably uiTstriped) ulbina, Wied., p. 303.
Femora black or blackish, with or with-
out white or silvery shimmer -2.
2. Front tibiae blackish brown, or at least
distinctly darker than posterior pairs. 3.
Alat mo" Sr£y) U±U:ly.Pale: JT«> W.K P. 305,
3. Thorax dull black, with two grey ' (^ms> Krob'>* P- 306«
stripes ; emarginations of abdominal
segments reddish or yellowish indica, Big., p. 304.
Thorax dark grey, one median darker
stripe ; abdominal emarginations not
coloured ; eyes absolutely contiguous, ceylonica, Krb'b., p. 304.
241. Psilocephala albina, Wied.
Thereva ulbina, Wiedemann, Zool. Mag. i, pt. 3. p. 3 (1819) ; id.,
Auss. Zweih1. i, p. 233 (1828).
Psilocephala albma, Krbber, Eut, Mitteil. Berl. i, p. 252 (1912).
J . " Head with silvery-white toraentum, extreme margins of
frontal triangle appearing dark brown ; in certain lights can be
seen two dark parallel stripes, fading away downwards towards
the antennae; antenna) blackish brown, sparsely beset with black
bristles; palpi and proboscis long, with snow-white hairs; back
of head with whitish-grey tomentum and white hairs ; orbital
bristles black. Thorax dull grey, with yellow and grey pubescence ;
scutellurn like thorax, with black bristles ; sides of thorax blackish,
grey-dusted and with grey hairs. Abdomen black, dull, with
somewhat strong silvery glistening and long whitish hairs, which
in certain lights, especially on the last segment, appear yellow ;
2nd to 4th segments with yellowish hind margins. Venter
blackish, with grey pubescence, with yellowish hind margins to
segments ; genital lamellae reddish yellow, with black and whitish-
yellow hairs. Legs: coxae blackish, white-haired; femora dark
brown, covered with white pubescence ; tibiae and tarsi pale
yellowish brown, tips darkened. Winys hvaline, veins weak,
yellowish, 4th posterior cell closed ; halteres blackish brown, tips
of clubs whitish. Length, 9 mm. Type c? from Java, in the
Vienna Museum.
5 . " Frons narrow, dull blackish brown ; epistome shining
silvery white ; two dull black spots near each other above the
antennae, outlined with white ; antennae and back of head as in J .
Thorax and scutellum black, with very sparse yellowish and grey
hairs ; sides of thorax blackish, with silvery-white tomentum and
white hairs. Abdomen dull blackish brown ; each segment bears
an ill-defined triangular silver-white spot on each side on the hind
* These two species are almost certain to be identical. No characters of
differential value can be selected from Krober's description of his unique type.
304 TIIEIIEYIM;.
border, not meeting on the disc ; according to the light the
abdomen above appears deep black or dark brown ; 2nd segment
with a distinct silky-white margin. Ground-colour of venter
black, with a silvery sheen; the 1st segment white-haired on eacli
side; the last one short, with black raised hairs. Femora blackish
brown, with silvery-white hairs : tibiae and tarsi pale yellowish
brown, front tarsi darkened. Wings as in c? ; 4th posterior cell
closed. Length, 10 mm. Type $ from East India, in Vienna
Museum." (JLrober.)
This species is introduced to the British Indian fauna on the
strength of the $ recorded by Krober from East India.
242. Psilocephala indica, Big.
Psilocephala indica, Bigot, Ann. Soc. Ent. France, (6) ix, p. 326
(1889).
6 . " Antennte blackish, the two first segments furnished with^
some black hairs ; proboscis and palpi blackish, these latter with
white hair ; face black, covered with whitish tomentum ; beard
white ; thorax dull black, as well as scutellum, two grey stripes on
dorsum, sides of dorsum and sides of thorax with white tomentum ;
halteres brown, pale at tips ; abdomen dull black, with venter and
sides with white tomentum, the emarginatious of the segments
reddish ; femora black, tibia? pale tawny with black tips ; tarsi
brown, metatarsi pale tawny at base ; wings hyaline.
$ . " Resembling the male ; frons black, whitish below, ab-
dominal segmentations paler.
"India. Three tf 6 , two $ $ . Length, 10 mm." (Bigot.}
243. Psilocephala ceylonica, Krrib.
Psilocephala ceylonica, Krober, Eut. Mitteil. Berl. i, p. 251 (1912).
d1 . " With sparse snow-white hairs throughout. Eyes abso-
lutely contiguous, rather large, shining green ; epistome and frons
snow-white, with silvery shimmer ; extreme margins of frontal
triangle velvet-black ; 1st antenual joint black, 2nd and 3rd dark
reddish brown, style black, bristles black. Thorax dull dark grey,
witli a blackish-brown median stripe, which is narrowly bordered
with whitish grey ; the short erect hairs are blackish, bristles
black; scutellum dark grey, blackish brown in centre; sides of
thorax silver-grey. Abdominal ground-colour deep velvet-black,
appearing wholly black or glistening silvery white according to
the direction of the light ; posterior margins of segments indis-
tinctly whitish ; genitalia small, reddish ; venter like dorsnm,
borders of segments distinct. Femora black, with silvery-white
pubescence ; tibia? and tarsi brownish yellow, front tibia? and tarsi
blackish brown, tips of posterior pairs darker. Wings with a
brown tinge, rather deeply yellowish brown on anterior margin ;
veins very strong, slightly infuscated ; 4th posterior cell closed in
one wing and open in the other ; halteres blackish brown, clubs paler.
Length, 12 mm. Colombo. Type in Hungarian Museum." (Krober.)
PSILOCEPHALA. 305
244. Psilocephala sequa, Walk. (PI. III. figs. 17-20.)
Thereva sequa, Walker, Ins. Saund., Dipt. pt. 1, p. 157 (1852).
<5 $ . Bead in c? with eyes contiguous for about half the distance
from fro us to vertex, leaving an elongate light yellowish-grey
vertex bearing a few brown hairs, with the front ocellus at some
distance from the other two ; ocelli red ; frons at level of antennae
nearly one-third of the head, and with face wholly covered with
snow-white tomentum, with here and there a little short con-
colorous soft pubescence ; upper corner of frons with a few black
stiff hairs. Proboscis short, thick, dark brown, extending as far
as tij) of 1st antennal joint, with moderately long soft white hairs.
Anteunal 1st joint blackish, white-dusted, with numerous strong
black bristles; 2nd and 3rd joints dull red-brown, with a little
greyish dust ; 2nd nearly cylindrical, with a few short bristles ;
3rd elongate, bare. Occiput projecting only slightly behind eyes,
covered with hoary tomentum, a little white pubescence, and some
strong black bristles. In $ , frons at vertex broader than in from Mandalay, ]2. iii. 1908, and Moulmein, 28. ii.
1908 (Annandale), all in the Indian Museum.
247. Phycus argentiventris, sp. nov.
3 . ITead: eyes contiguous for a considerable distance ; vertical
triangle bluish grey ; frons and face bluish grey, rather shining;
:intennal prominence black ; antenna? dull brownish yellow, 1st
and 2nd joints pubescent ; vertical margin and behind upper part
of eyes with a row of well-separated black bristles ; occiput
ash-grey, rather shining in certain lights ; lower part wilh soft
white hairs. Thorax as in brunneus, the median stripe with a
bluish tinge. Abdomen comparatively flattened above, covered
with bluish-grey tomentum, which (seen from in frontj glistens
silvery white ; some sparse white pubescence at sides, longer
towards base ; hind border of 3rd segment with a narrow yellowish
band, rather inconspicuous ; traces of a similar band on 4th seg-
ment; genitalia large, conspicuous, bright orange with a little
black pubescence ; venter black, hind margin of at least 2nd, 3rd,
and 4th segments with a distinct yellowish band. Legs : femora
and posterior tibiae blackish ; fore tibia? and tarsi dull yellowish
brown ; posterior tarsi pale yellowish, blackish at tip and at tips
of joints. Wings as in brunneus, unmarked; halteres blackish
brown.
Length, 7 mm.
Described from a single specimen from Unt-hogaon, Naini Tal
Distr., 13.iv.1910, in the Indian Museum. Specimens subsequently
seen in the British Museum from Periyakulam, Ceylon, 22, 23. ii.
1891 (Yerbury).
A very distinct species, characterised by the contiguous eyes and
silvery abdomen.
248. Phycus atripes, Brun., nom. nov.
Phycus nigripes, Bruuetti (nee Kriiber, 1912), Bee. Lid. Mus. vii,
p. 480 (1912).
Allied to brunneus, but quite distinct owing to the 3rdantennal
joint being evidently longer than the 1st and the wholly black
legs. Lower half of frons wholly shimmering silvery white,
without the large shining jet-black spot immediately above the
antenna? which is so conspicuous in both those forms ; 3rd
antennal joint cylindrical, much narrower than in the other
species. Median dorsal stripe faint ; abdominal bands very nar-
row, milk-white, only the one on the 2nd segment really distinct.
Legs wholly black, including coxa?, which have a little bluish-
white shimmer. Wings clear, with a distinct blackish suffusion
towards tip, extending from the costa over about the distal half
of the 2nd longitudinal vein, reaching irregularly to the hind
310
margin, just encroaching on the discal cell, and with a small
irregular clear space in each of the two submarginal cells ;
halteres with knobs wholly black. From the narrow frons and
the appearance of the abdomen I conclude the single specimen
is a cT .
Length, 10 mm.
Kurseong, 27. hi. 1910 (Annandale).
Type in the Indian Museum.
Family SCENOPINID.E.
Head nearly semicircular ; frons and face nearly flush with the
eyes ; three ocelli ; epistome short, retreating ; proboscis very
short, labella broad; palpi short, cylindrical, or slightly clavate.
Eyes bare, generally contiguous in d, wide apart in $; upper
facets larger than lower ones in th veins to the wing-margin may be regarded as the continua-
tion of either one of them or of the two united.
310
axillary vein indistinct, perhaps sometimes absent. Alar squamae
small, nearly bare of fringe, but larger and with coarse fringe in
Mydas ; thoracic squamae absent.
Life-history mainly unknown, except that the larvae are said to
live in decaying wood and to resemble those of the ASILID.E.
Williston states that the larva of a North American species
(Mydas fulvipes, Walsh) preys upon Coleopterous larvae, and notes
the pupa of M. clavatus, Drury. The perfect insects have been
accused of being predatory, but there seems to be little decisive
evidence on the matter. An Egyptian species is said to be
predatory on other insects, catching them in the air ; and another
species is recorded as parasitic on the larvae of Longicorn beetles
The MTDAIDJB occur chiefly in the New World and Australia,
only about 15 species being Palaearctic and six Oriental, of
which four are Indian.
Their peculiar venation differentiates them easily from all other
groups, except the NEMESTIUNIDJE and APIOCERID^E. These
latter are a small non- Oriental group that has been at various
times associated with the BOMBYLIIDJE, THEEEVID^, and ASILID^E.
There is some resemblance in the venation of the MYDAID^E to
that of the NEMESTIUNID.E, but the bodily structure and general
facies of the latter are different.
The presence also in the NEMESTHIXIDJE of what has been
conveniently though incorrectly termed the "diagonal vein" will
also immediately separate that family from the present one.
Verrall has carefully analysed the systematic position of the
present family (Brit. Flies, v, p. 605), and in the same volume
(p. 606) has discussed the question of spelling, several authors
having adopted Midas instead of Mydas. The exact origin of the
generic name is not clear, and he supports Mydas, with the family
name MYDAIDJE, and these terms are also adopted in the Kertesz
Catalogue and in Willistou's Manual.
Table of Genera.
Proboscis short; labella terminal, triangular, compressed ;
hind tibise generally Avith a spur; anal segment in
$ without a wreath of spines ................. . MYDAS, F.
Proboscis long ; hind tibiae without spur ; anal segment [Gerst,
in $ with a wreath of spines .................... LEPTOMYDAS,
Genus MYDAS, F.
Mydas, Fabricius, Ent. Syst. iv, p. 252 (1794) ; Macquart, Hist. Nat.
Dipt, i, p. 273 (1834) ; ]d., Dipt. Exot. i, pt. 2, p. 10 (1838).
GENOTYPE, Musca clavata, Drury; by Latreille's designation
(1810).
Head broad, short transverse, as broad as thorax, neck short ;
MTDAS. 317
frons and vertex sunken, ocelli indistinct or removed far from
vertex ; proboscis only slightly prominent,* labella terminal, tri-
angular, compressed; palpi one-jointed; antenna porrect, at
base approximate, clubbed or knobbed at tip, three-jointed, the
first two moderately short, the 3rd elongate, with annular im-
pressions. Thorax subquadrate, with sparse, sometimes thicker
pubescence on dorsum and coarse pubescence towards the sides ;
pleurae with more pubescence in some species than others ;
humeri prominent; scutellum oblong, small, unspined, meta-
notum very large. Abdomen elongate, 7-segmented, with light
pubescence only, the last segment in the $ without a circlet of
spines ; genitalia of moderate size in c? . Legs strong and stout ;
hind femora slightly thickened, generally serrate or spinose on
underside ; hind tibiae generally with a spur. Wings : auxiliary,
1st, 2nd, and 3rd longitudinal veins as in the family diagnosis,
both branches of the latter vein ending in the 1st; the 4th vein's
upper branch ends in the costa distinctly before the wing-tip, its
lower branch bends downward and joins the upper branch of
the 5th vein. As for the small vein forming the outer side of
the discal cell, it may be regarded as a lower prong of the upper
branch of the 4th vein or as a discal cross-vein. Eemaining
veins in accordance with the family diagnosis.
Range. S. Europe, "N". Africa, India, Malay Archipelago, North
and South America, and Cuba.
Life-history practically unknown.
Table of Species.
Third autennal joint bright red ruficornis. Wied.
Third antennal joint black.
Abdomen wholly black carmiahaeli, Brun.
Abdomen with pale hind margins to segments . annulata, Bran.
250. Mydas ruficornis, Wied.
Mi/das riificornis, Wiedemann, Analec. Entom. p. 20 (1824) ; id.,
Auss. Zweifl. i, p. 242 (1828) ; id., Monog. Gen. Midas, p. 40,
pi. lii, fig. 11.
" Head black ; 1st and 2nd antennal joints black, 3rd fiery red-
yellow ; epistome on both sides [presumably above and below]
with yellowish, hairs ; a white hair spot on the side of the
:mteana3 ; occiput whitish. Thorax brownish honey-yellow, be-
coming blackish behind. Abdomen almost metallic black, rather
sparsely punctured. Wings brown, with violet-blue reflections ;
halteres brown. Legs black ; hind tibiae and base of tarsi dark
red-brown ; hind femora below with three or four rather long
spines." ( Wiedemann.)
Length, 18 mm.
Tranquebar.
Type in the Vienna Museum.
* Scbiner adds " with four bristles."
313 MYDAIDJE.
251. Mydas carmichaeli, Brim.
My das carmichaeli, Branetti, Rec. Incl. Mas. ix, p. 266 (1913).
cf § . Head black, irons slightly wider than one-third of the
head, somewhat wrinkled, with a moderate amount of blackish-
brown pubescence ; a small tuft of greyish hairs on each side of
the mouth-opening. Proboscis rather less than the height of the
head, dark brown ; palpi slender, black, reaching as far forward
as the vertical line drawn through the antennal prominence, which
latter is small and black. Antennae normal, black, bare, about as
long as from the vertex to the tip of the proboscis. Occiput
black, shortly pubescent; a little grey tomentum behind the eye-
border. Thorax wholly dull velvet-black, with very short
sparse black pubescence, which is a little longer below the wings.
Fig. 23. — Mydas carmichaeli, Brun., <$ .
Abdomen shining black, with a slight indigo-blue tinge and very
short pubescence ; in <5 subcylindrical, as long as head and thorax
together and barely as wide; in § a little broader and longer and
much more bulky. The tf genitalia consist of a hollow triangular
piece, hairy above, with a deep keel, a pair of dark reddish-brown,
nearly bare, moderately long, narrow, finger-like claspers (only
one joint being visible, the organs being somewhat withdrawn),
and a curved ventral plate, with black pubescence on its lower
side. The $ genitalia appear to consist of two telescopic cylin-
drical shells, the inner one ending in two small (probably) retractile
lamellae. Legt wholly black ; coxae with a little short black
pubescence ; hind femora with two rows of spines on underside,
an inner one of about a dozen and an outer one of a few only.
Hind tibiae with a row of reddish spines on underside (not at all
conspicuous), and 5 or 6 such spines at the tip on the underside
only ; fore tibise with some inconspicuous spines at tip, which
appear to be only the terminal ones of a row on the underside
nearly hidden by the thick though short black pubescence which
clothes all the tibiae ; a row of spines, including the apical Ones, is
.-ipparently present on the middle tibiae also, though certainly
MYDAS. 319
much less conspicuous. Claws distinct, pulvilli pale brownish-
yellow. Wings dark blackish brown, a little paler towards tip
and hind margin, and with a strong violet tinge. Halteres and
squamae black, the latter with a narrow fringe of dark brown
hairs.
Length, <$ 25 inm., $ 27 mm.
Described from a single pair, the <3 from Darjiling, v. 1912, the
$ from Singla, iv. 1913.
252. Mydas annulata, Brun.
C'ephalocera annulata, Brunetti, Rec. Ind. Mus. vii, p. 513 (1912).
c? . Head : frons rather more than one-third width of the head,
with a small swelling on each side of vertex, contiguous to the
corners of the eyes ; proboscis short. The whole head moderately
shining black, with rather thick and long whitish-grey pubescence
over frons and face except lower part of the latter ; occiput black,
with whitish-grey hairs, which are longer on lower part of head.
Antennae all black, long ; 2nd joint very short, 3rd joint with an
impressed constriction at some distance before its tip; beyond the
constriction the joint is slightly thickened ; style long, nearly as
long as rest of antennae, black, with an impressed constriction at
nearly one-third of its length, the style thickening upically except
at extreme tip, where it is slightly but obviously narrower.
Thorax black, a grey tomentose stripe at each side of the dorsum,
a short one from each shoulder, narrowing rapidly and not reach-
ing centre of dorsum ; the middle of the posterior margin also
grey tomentose ; the whole dorsum covered with tine grey pubes-
cence, which is a little longer beiow the shoulders and on the side
stripes. Scutellum broad, extremely short, shining black, bare ;
sides of thorax moderately shining black, with a little grey
pubescence on the pleurae. Abdomen shining black; posterior
margins of all segments (indistinctly on the 1st segment) with a
narrow, but clearly-defined, pale whitish-yellow border; whole
surface of abdomen with depressed pubescence, which is dark
brown or blackish on basal halt' (or thereabouts) of each segment
and pale grey on apical half. Venter similar, the yellowish-white
bands 'passing over the sides of the segments and continued
narrowly across the ventral surface. Geuitalia comparatively
small, black ; a dorsal plate, a pair of moderately long finger-like
claspers, and some yellowish-brown internal organs. Legs black ;
shortly and normally grey pubescent ; base of hind tibiae and tarsi
slightly brownish yellow. Wings pale grey ; venation in exact
agreement with Macquart's figure of Cephalocera nigra (Dipt.
Exot. i, pt. 2, pi. i, fig. 3) ; halteres black.
Length, 15 mm.
Described from two 6 6 in the Indian Museum from Bhoira-
kund, Darrang District, Assam-Bhutan Frontier, 18-22. x. 1912
(Kemp).
320 MTDAIIME.
I described this species under CepJialocera, as Bigot's MS. name
for it was adopted, but, as the proboscis is short and labella are
present, it must be referred to Mi/das.
Genus LEPTOMYDAS, Gerst.
Leptomydas, Gerstaecker, Stett. Ent. Zeit. xxix, p. 81 (1868).
GENOTYPE uncertain ; * Mydas lusitanicus, Wied., according to
Coquillett (1910).
This genus differs from Mydaa in the long proboscis, in the
absence of a spur to the hind tibiae, and the presence of a wreath
of spines at the tip of the abdomen in the $ . From Ceplialocera
it differs by the presence of the labella.
Range. South Europe, Egypt, Arabia, Sudan, India, East and
South Africa, North and Central America. %
253. Leptomydas indianus, Bnm. (PI. IV, fig. 1.)
Leptomidas indianus, Brunetti, Eec. Ind. Mus. vii, p. 478, fig. 12
(1912).
$ . Head much wider than thorax ; frons black, shining, with
two narrow diagonal grooves on upper part ; vertex consisting of
two dark brown shining oval calli placed end to end, with a small
recessed space between them ; face rather dark grey, smooth ;
antennae missing, except the short, .cylindrical, dark brown,
slightly pubescent first joint, placed on a slight transverse ridge
running from eye to eye ; eyes prominent, black, bare, shining ;
proboscis and oral opening dark brown, shining, former very
short ; back of head grey ; the whole frons, face, back, and under-
side of head with moderately thick (but not dense) grey hairs.
Thorax dull black, with a little dirty grey tomentum towards the
sides and posterior part, and on the humeral calli, which bear a
small bunch of coucolorous hairs ; similar hairs on posterior part
of dorsum, in front of and below the wings ; sides of thorax more
or less shining black, mostly bare; prothorax and neck with
silvery-grey tomentum ; scutellum and metanotum blackish,
•with silvery-grey tomentum. Abdomen cylindrical, slightly nar-
rowed in middle, very dark blackish brown, shining; posterior
borders of segments pale yellow, most distinct on 2nd to 5th seg-
ments ; the bands on the 2nd, 3rd, and 4th segments passing
unbroken over the sides and across the venter; 4th segment light
reddish brown ; whole abdomen minutely pubescent ; 1st and 2nd
* There is a little uncertainty here, as in the Kertesz Katalog lusitanicus is
referred to Mydas again, although Leptomydas is acknowledged as a valid
genus. The next species guen by Gerstaecker is cincfus, Macq., which Kertesz
puts in Syllcyomt/das. Gerstaecker's third species is rujipcs, Wied., which
Kertesz retains in Leptomydas. Therefore, if lusitanicus is really a Mydas, the
type of Leptotomydas may be rufipcs, Wied.
LEPTOMYDAS. 321
segments with grey hairs at sides. Genitalia small, partly with-
drawn, dark shining brown; ultimate portion with a row of light
brown scale-like spines on each side. Legs simple, no traces of
bristles or longer hairs, except two apical bristles at the tips of the
anterior tibiae (the hind tibiae and tarsi are missing) ; dark brown,
with minute grey pubescence, which is longer on the coxae and
short and thick on the tibiae ; tarsal joints with black tips.
Wings uniformly pale yellow, veins darker yellow, costa black ;
border of wing without a single hair ; 3rd longitudinal vein with
a minute appendix at base of upper fork; alulae concolorous, bare;
tegulae very small, bright yellow with a very narrow fringe; halteres
blackish, stem brown.
LenytJi, 16 mm.
Described from a single $ in the Indian Museum from Sadiyar
Assam.
The type is a $ , as noted on Bigot's MS. label, though in
describing it 1 stated it was probably a c? . I placed indianus
in Leptomydas because Bigot's MS. label bore that name, but an
examination of the various genera shows that in this genus the
proboscis should be long. In indianus it only projects from
the mouth-opening for a short distance, the tips of the distinct
labella reaching as far forward as the 2nd antennal joint. The
only other genus to which it could be referred seems to be
Ectyphus, Gerst., in which the proboscis is described as rudi-
mentary, barely projecting from the oral orifice. But the epistome
in Ectyphus is long and vertical, whereas in indianus the lower
part of the head does not descend below the level of the eyes.
The proboscis in the type projects upwards at an angle ; if directed
forwards, it would, of course, appear comparatively longer : Ger-
staecker's figure shows a proboscis reaching to half the length of
the antennae. However, the tip of the abdomen bears the circlet
of spines, which is one character that differentiates Leptomydas
from Mydas, so in the circumstances the species is left here-
temporarily.
Family EMPLOYE.*
Comparatively small-sized flies, generally bare or nearly so
except for a few bristles, of sombre colours ; t found chiefly in
woods and shady situations, many species frequenting the neigh-
bourhood of water. In the tropics they occur mostly in localities
of some elevation. Systematically much more variety of struc-
ture is present than in the adjacent families of ASILID^K and
DOLICIIOPODID^E.
* So.ne authors adopt the name EMPIDID*:. The late G. H. Verrall saj»
(Brit. Flies, v, p. 28): " Dr. A. W. Verrall is of opinion that the word Lmptdidce
is pedantic, and instances Euelpida."
t The only exceptions are a few metallic species from the tropical region*
of the New World.
322 EMPIDJE.
Head comparatively small, sometimes very small, more or less
globular, generally bulging out posteriorly ; as wide as, or narrower
than the thorax ; joined to the thorax by a narrow neck. Eyes
large, often forming nearly the whole of the head, generally bare,
contiguous in both sexes or in the c? only, or separated in both
sexes ; in contiguous eyes the upper facets nearly always larger
than the lower ones ; in the middle of the inner eye-margin is an
incision of greater or less size, said to be always present and
perceptible ; three ocelli. Proboscis of various structure ; short
and thickened, of more moderate size both in thickness and length ;
or, as is frequently the case, horny, adapted for piercing, directed
forwards ; or elongate, in some species as long as the whole body,
and directed obliquely forwards, downwards, or backwards. Occa-
sionally the proboscis is curved. Mouth-opening generally small.
Palpi generally 1 or 2, rarely 3, jointed ; of various structure,
size, and length ; bare or bristly. Antennae porrect, approximate
at base, sometimes 2-jointed, generally of three unquestionable
joints, of which the 3rd may taper into, or bear, an arista, which
is always bare except when styliforrn. In some genera a pro-
minent 1- or 2-jointed style is present, and some authors have
described such antennae as being 4- or 5-jointed. Even in certain
genera in which they are regarded as 3-jointed, the 3rd joint
(in these cases nearly always elongate conical or pear-shaped) is
apparently of a different texture from the basal two, partaking
more of the nature of a style. In the present work the latter
genera are regarded as having a 3-jointed antenna, attention being
called to the structure of the 3rd joint. In the genera with so-
called 4- or 5-jointed antennae the 4th and 5th joints are regarded
as representing a style. Head with a few weak bristles on vertex,
frons, or occiput.
Thorax of moderate size, often greatly humped, generally sub-
quadrate, the prothorax sometimes produced forwards. Scutellum
small, semicircular, or gently curved, with two or more bristles ;
a few bristles generally occur on the margin of the thoracic
dorsum.
Abdomen of 7 or 8 segments,* bare or pubescent ; generally in
the <$ elongate, conical, or cylindrical, in some genera short,
rounded, or ovate ; io the 5 sometimes broader at the base, but
always more pointed at tip. Genitalia in <5 of various structure,
sometimes complex and prominent ; in £ usually in the form of
a slender conical ovipositor with two terminal styliform lamellae.
Abdomen never with conspicuous bristles.
Legs exhibiting various forms of structure. In some groups
they are comparatively normal, that is to say, of moderate length
and thickness ; but in others the coxae may be greatly lengthened,
the femora (either pair singly or the whole of them) being often
conspicuously incrassated, and in some genera furnished with
spines below. The tibiae and metatarsi are sometimes enlarged,
whilst marked differences in sexual ornamentation not infrequently
* Melander says 5 to 7.
EMPID.E. 323
occur in the legs in certain genera — Erupis and EhampJiomyia, for
instance, — in which in some species those of the
4. Femora and tibiae pennate sgwtnata, Brim., *
Femora and tibiae not pennate 5. fp. 352.
5. Auxiliary vein and 1st longitudinal veins [p] 350..
"nited ccylonica, Bezzi,
Auxiliary and 1st longitudinal veins separate, carbwiaria, Brun.,
6. Anterior cross-vein between J- and middle of !~p. 351.
discal cell 7.
Anterior cross-vein distinctly before £ of discal
cell 11.
7. Posterior femora and tibiae pennate 8.
Posterior femora and tibiae not pennate .... 10.
8. Hind tibiae shortly pennate on basal half
only, in front and behind; proboscis 1'
times height of head, not reaching base of
abdomen ; anterior cross-vein at middle of [p. 353.
discal cell centralis, f Brun.,
Hind tibiae with a long fringe for practically
their whole length, in front and behind . . 9.
9. Hind metatarsus pennate ; proboscis more than
twice height of head, reachingbeyondba.se
of abdomen ; anterior cross-vein between [p. 355,
£ and middle of discal cell rostrata, Brun.,
Hind metatarsus not pennate; proboscis 1^
times height of head, not reaching base of
abdomen; anterior cross- vein just before Pp. 354.
middle of discal cell . . ' . sulcilipes, Brun.,
10. Dorsum of thorax with three black stripes ;
legs dark brown ; anterior cross-vein at £ [p. 353.
of discal cell ; abdomen uniformly grey . . yriseonigra, Brun.,
Dorsum of thorax with four black" stripes ;
legs bright orange : anterior cross-vein just
before middle of discal cell ; hind margins
of 2nd, 3rd, and 4th abdominal segments [p. 356.
black marf/inata, Brun.,
11. Legs pale yellow eleyam, Brun, p. 356.
Legs dark brown inconspictia, Brun.,
[p. 357.
282. Empis amplitarsis, sp. nov.
c? . Head with vertex moderately large, blackish ; two long^
ocellar bristles and some smaller ones ; upper eye-facets bright
brown, smaller lower ones coffee-brown ; antennae black, 3rd
joint as long as 1st and 2nd together, subconical, with elongated
tip and long style ; proboscis brownish yellow, about twice as
long as vertical height of head ; palpi rather small, upturned,
yellowish ; occiput dark grey, with the usual bristles. Thorav
moderately shining blackish, with grev-dusted reflections in certain
lights. Abdomen moderately shining dark brown, with short sparse
pubescence, which is longer at the sides. Legs uniformly mode-
rately dark brown, except the extraordinarily developed front
tarsi, which are a little darker ; these latter are fully as long a»
the rest of the front leg from base of coxa to tip of tibia ; 1st joint
* May be the $ of carbonaria, Brun.
t May be the ? of griseoniyra, Brun.
350
nearly as long as tibia, flattened, broad at tip ; 2nd joint rather
more than half as long as first, narrower, cylindrical, thickened ;
3rd joint nearly as long as 1st, much incrassated, broader and
much longer than 2nd and broader at tip than at base ; 4th joint
about as long as 3rd, narrower, thin, considerably arched, so that
the underside forms a large cavity ; 5th joint shorter than 4th,
moderately incrassated, bluntly conical ; claws and pulvilli normal.
Pubescence on coxa3 and femora moderately long and dense, a little
thicker on hind femora, with a few longer hairs on outer side ;
front tibia3 with a row of short hairs on inner side, a row of longer
ones on outer side, and a row of very long ones on upper side ;
middle tibiae moderately pubescent, with two long bristles on outer
side (one at middle, one preapical) ; hind tibiae covered with rather
dense, moderately long pubescence and longer hairs, and with a row
of quite long hairs on hinder side; anterior tarsi with 1st joint
nearly bare except for a few hairs towards tip, remaining joints
Fig. 29. — Empis amplitarsis, sp. DOT., front leg of male.
with long dense hairs on upper and outer sides, a few short hairs
on inner and lower sides, the 4th also bearing a denser bunch on
upper side near tip. Wings pale grey ; venation approximately
normal, anterior cross-vein at \ discal cell ; halteres moderately
dark brown.
Described from a single <3 from the Pusa collection from
Belgaum, Bombay Presidency, 2500 ft., 11. viii. 1910 (Fletcher],
presented to the British Museum.
A label attached by Mr. T. Bainbrigge Fletcher says : " Fore-
legs are stretched out in front and function as antenna?; they
are not used for locomotion." The extraordinary development
of the front tarsi easily distinguishes this species from all other
Indian EMPID.E, yet this character is the only one in which it
differs from typical species of Empis.
283. Empis albohalteralis, sp. nov.
<3 . Head with vertex blackish, frons and face moderately
dark grey; first two antennal joints yellowish, 3rd black, base
narrowly pale, style rather long ; proboscis black, reaching (bent
under body) to middle of abdomen ; palpi pale yellowish, curved
upwards, rather long, with a few black bristles ; eyes with large
upper facets bright brown, lower small ones coffee-brown; occiput
ash-grey, a marginal row of small black bristles and a more central
EJIPIS. 351
row of stronger ones. Thorax shining black, closely and entirely
covered with cinereous dust, in which three narrow slightly
yellowish stripes of the usual nature can be faintly traced; scu-
tellum and metanotum with dark grey dust ; sides of thorax
concolorous with dorsum ; metapleural hairs yellow. Abdomen
blackish, rather shining, with sparse pubescence; hind margin of
segments indistinctly pale ; venter yellowish ; genitalia shining
yellowish brown. Legs: coxae dull orange, more or less grey-
dusted on basal part, a little pale pubescence in front ; remainder
of legs orange, hind tibiae more brownish towards tips ; anterior
tarsi blackish brown towards tips, hind metatarsi much enlarged
and lengthened, remaining hind tarsal joints black. Wings pale
grey, venation normal; anterior cross-vein just before middle of
discal cell ; halteres conspicuously chalk-white.
Length, 5 mm.
Described from one d1 in the Indian Museum from Zairat,
Baluchistan, 8000-9000 ft., viii. 1914 (Major Wilson).
Easily recognised from all other species by the conspicuous
white halteres.
284. Empis carbonaria, Brun.
Empis carbonaria, Brunetti, Rec. Ind. Mus. ix, p. 27 (1913).
c? . "Wholly and uniformly coal-black, moderately shining.
Eyes separated by a narrow frons. Antennal 3rd joint distinctly
longer than the first two together, and its distal half rather
suddenly narrow ; style thick, half as long as the joint. Proboscis
nearly twice the height of the head, labium bifid at the tip.
Fig. 30.— Empis carbonaria, Brun.
Head, thorax, and abdomen covered with long, soft, but not
copious pubescence. Legs dark brown, with rather long soft black
hairs ; fore femora almost bare ; fore metatarsus distinctly incras-
sated, cylindrical, lengthened, as long as remaining tarsal joints
together ; hind metatarsus as long as next three joints together,
352 EMMDJE.
slightly thickened ; posterior tibiae and tarsi with a row of well-
separated long bristly hairs. Genitalia small, rounded, not
conspicuous. Wings clear; anteiinal cross-vein at one-fourth
of the discal cell ; 4rh vein shortened ; stigma slightly brownish,
small ; halteres black.
Length, 4 mm.
Described from two d J in the Indian Museum and one in
Mr. E. E. Green's collection from Horton Plains, Ceylon, v. 1911
(Green).
Type in the Indian Museum.
285. Empis squamata, Bmn. (PI. IV, fig. 11.)
Empis squamata, Brunetti, Rec. Ind. Mus. ix, p. 27 (1913).
5 . Wholly coal-black. Proboscis three times as long as the
head. Antennae normal. Thorax witii soft black hairs. Abdomen
nearly bare, roughened, shining; venter similar; ovipositor small,
elongate, normal. Legs with a dense row of long, closely placed,
dark brown scales on underside of fore femora, on upper- and
undersides of posterior femora, front and hind sides of fore and
hind tibiae, outer and inner sides of middle tibiae, and on
upper sides of all metatarsi ; a single long bristly hair on outer
and inner sides, projecting somewhat backwards towards tips of
all tibiae and of first four tarsal joints, these hairs very much
longer on hind legs ; the unsealed parts of the legs bear a little
black pubescence. Wings pale brown ; 4th vein abbreviated ;
stigma black ; halteres black.
Length, 2|-3 mm.
Described from several 2 $ collected by Mr. E. E. Green ou
the Horton Plains, Ceylon, May 1911.
Type and other specimens in the Indian Museum.
Probably the $ of carbonaria, having been taken so often by
Mr. Green in company with that species. However, it must be
distinct from ceylonica, Bezzi, with which at first I thought it
might be synonymous, as that species lias no conspicuous brown
scales on the legs, and has the auxiliary and 1st longitudinal veins
anastomosed. E. squamata has a considerable resemblance to
jacobsoni, Meij., described from Java, but. in the latter species the
upper side of the front femora bears long brown scales like the
posterior pairs.
286. Empis ceylonica, Bezzi.
Empis ceylonica, Bezzi, Ann. Mus. Hung, ii, p. 343 (1904).
$. A shining black species, distinguished by the broadened
brownish wings, with basal third hyaline ; pubescence black, but
below the setae the body is bare ; proboscis black, as long as head
and thorax together; frons shining black, bare, head-bristles
rather long. Antennae short, black, shortly pubescent at base;
3rd joint with upper edge straight and excised below, ending in a
EMPIS. 353
rather long style. Thorax shining black and hare, with the
pleurae greyish : 2-3 long hlack metapleural bristles and two
moderately long presutural ones ; squamae yellowish, with black
fringe. Abdomen brilliantly shining black, with a few black hairs;
venter with longer hairs on margins of segments ; ovipositor long,
slender, with elongate lamellae. Wings broadened, axillary angle
developed ; 1st longitudinal vein rather long, united to auxiliary
vein ; 3rd vein bent downwards somewhat towards tip so that it
ends at tip of wing, its upper branch nearly perpendicular ; discal
cell large, apical end truncate ; 4th vein rather long, ending at
some distance from the margin ; 6th vein shortened. Legs brown,
rather long, especially hind pair, pubescent and with bristles ;
middle tibiae with two long bristles above ; hind tibiae with 6 or 7
long ones in two rows ; hind metatarsus very elongate, as long as
remaining joints together, with three or four long bristles above ;
remaining joints with a single bristle at tip of each.
Length, 3 mm. ; wing, 4 mm.
A unique female from Pattipola, Ceylon, 2000 metres,
22. ii. ]902.
Type in the Hungarian Museum.
287. Empis griseonigra, Brun.
Empis yriseonigra, Brunetti, Uec. Lid. Mus. ix, p. 29 (1913).
c? . Head with occiput blackish ; eyes absolutely contiguous
above antennae ; face dark grey ; antennae black ; palpi black ;
proboscis one-fourth longer than the height of the head, dark
brownish yellow, black on underside. Thorax brownish grey,
with three moderately wide brown dorsal stripes, the median one
attaining the anterior margin, the outer ones only extending as
far as the shoulders. Some short bristly hairs on shoulders, soft
black pubescence over the dorsum generally, and stiffer hairs and
bristles distributed as in the other species. Sides of thorax a
little darker grey, pleurae light grey. Abdomen blackish, with
short sparse black pubescence ; whitish pubescence at sides of the
first t\vp segments ; genitalia blackish. Legs dark brown, with
black pubescence; knees reddish brown, with a trace of this
colour over some of the other joints. Wings pale grey ; stigma
imperceptible ;' anterior cross-vein at about two-fifths discal cell;
halteres bright yellow.
Length, 5 mm.
Described from one c? in the Indian Museum from Mundali,
Jaunsar division, Dehra Dun, 9000 ft., 12. v. 1910 (A. D. Imm»).
288. Empis centralis, Brun.
Empis centralis, Brunetti, Rec. Ind. Mus. ix, p. 26 (1913).
$ . Head : occiput black, with black hairs above, and some
white hairs below ; frons and face proportioned as in K. rostrata
2 A
354
but blackish, as are also the antennae, but the face, viewed from
below, shows dark grey reflections ; palpi bright brownish yellow ;
proboscis one and a half times the height of the head, the upper
side brownish yellow. Thorax light brownish grey, with soft
black pubescence, and bristly hairs laterally aud towards the hind
margin; two narrow median well-separated brown stripes from
anterior margin extending three-fourths of the distance to the
scutellum ; a lateral broader stripe on each side, practically formed
of three irregularly-shaped elongate spots (one on the shoulder,
one in front of and one behind the pre-sutural depression), which
are contiguous, the hind one attaining the posterior margin of
the dorsum. Some bristles laterally and towards hind margin
of mesoriotum, also four on the scutellum. Sides of thorax
and metanotum light grey, dorsum of scutellum brownish grey.
Abdomen darker grey, with short sparse black hairs ; posterior
margins of segments more or less black ; genitalia elongate, con-
colorous, much as in E. rostrata ; venter dark grey. Legs : cdxse
dark grey, remainder of legs brownish yellow, extreme tips of
femora blackish, and tips of tarsal joints more broadly blackish.
All the legs with black pubescence ; posterior femora with blackish
bristly hair above ; hind femora with scales ; posterior tibia? with
blackish-brown scales on front and hind sides, as in E. rostrata,
but much less strong and appai-ently confined to the basal half of
the hind pair; on the hind side of the hind tibia? a row of moder-
ately long, well-separated bristles. Wings very pale grey, nearly
clear; anterior cross-vein directly over the middle of the discal
cell ; halteres bright brownish yellow.
Length, 5 mm.
Described from one § in the Indian Museum from Mundali,
Jaunsar division, Dehra Dun, 12. v. 1910.
It seems likely that this will prove to be the female of E.
griseonic/ra.
289. Empis subcilipes, Emu.
Empis subcilipes, Bruuetti, Rec. Ind. Mus. ix, p. 24 (1913).
? . Head ash-grey ; frons about one-fourth the width of the
head ; some short stiff black hairs behind the eyes, and some soft
white hairs on lower part of occiput ; antennae reddish brown,
rather long, black-streaked on upper side and with a whitish
shimmer on the inner side in certain lights; antennal style of
moderate length ; proboscis nearly three times the height of the
head, reddish brown, with the underside black. Thorax moderately
dark ash-grey ; a broad brownish-grey median stripe from the
anterior margin and a less distinct much shorter stripe on
each side, not extending to the shoulders. Surface of dorsum
with short setae more or less irregularly placed, but a distinct
double row in the centre and a number towards each shoulder ;
some stiff bristles above and in front of each wing and four on
EMPIS. 355
posterior margin of scutelluin, which latter with the metanotum
are both light mouse-grey ; sides dark grey. Abdomen rather
dark grey ; 1st segment with a little black and grey pubescence ;
remainder of segments with scattered, very short pale hairs ;
venter tawny ; genitalia very dark brown. Legs brownish yellow,
coxae a little bluish grey on basal half ; femora more or less
blackish in middle and on upper side generally ; tarsi mainly
black. Upper and lower sides of four posterior femora with thick
but short black ciliated hairs ; rest of legs finely black pubescent.
Winys pale grey, veins brownish yellow on basal part, blackish
distally. Anterior cross-vein just before middle of discal cell ;
stigma imperceptible ; halteres brownish yellow.
Length, 5^ mm.
Described from a single 5 in the Indian Museum from Mundali,
Jaunsar division, Dehra Dun Distr., 9000 ft., 12. v. 1910.
290. Empis rostrata, Brun. (PI. IV, figs. 9, 10.)
Empis rostrata, Brunetti, Kec. Ind. Mus. ix, p. '25 (1913).
£ . Head with occiput grey, with a few bristly black hairs above ;
frons and face concolorous grey, the former comparatively wide,
with parallel edges, the latter much wider ; antennae black, 2nd
joint very distinctly bright brownish yellow ; palpi bright brownish
yellow, short ; proboscis two and three quarter times height of
head ; reaching beyond base of abdomen, brownish yellow, a little
blackish in middle on underside ; a few whitish hairs on underside
of head. Thorax : domain (somewhat discoloured in the centre)
light brownish grey, with a narrow median darker brown stripe
and a wider one on each side not quite extended to anterior
margin of thorax ; and laterally two brownish spots, one in front
Fig. 31. — Empis rostrata, Brim., wing.
of and one behind the presutural depression; sides, scutellum,
and metanotum concolorous, or a little more nearly ash-grey.
(Thorax denuded of bristles.) Abdomen light grey, brownish at
base of each segment, the colour broader in the centre ; genitalia
slender, cylindrical, terminated by two thin finger-like lamellae,
ooncoloroiis; venter light grey, unmarked. Leys between light
brown and tawny ; coxae a little grey-dusted and with a little soft
whitish hairs ; tarsi more or less blackish on upper side ; extreme
tips of femora very narrowly black; posterior femora with blackish-
brown scales above and below ; posterior tibiae with blackish-brown
scales, much thicker and larger on hind pair, on which they occur
356 EMPID.E.
below the metatarsi also, and are even continued in a less pro-
nounced scale-like form on the underside of the remaining tarsal
joints ; the legs generally with minute stiff black hairs. Wings
brownish yellow ; anterior cross-vein at two-fifths of discal cell ;
stigma practically imperceptible ; halteres brownish yellow.
Le&giht 7 mm.
Described from a single $ in the Indian Museum from Theog,
Simla Distr., 8000 ft., 1. v. 1907 (Annandale).
291. Empis marginata, Brun.
Pachymeria marginata, Brunetti, Rec. Ind. Mus. xiii, p. 79 (1917).
$ . Head : frons ash-grey, with parallel sides ; ocellar triangle
large, black ; palpi bright orange ; antennas all black, 3rd joint
considerably broadened at base, style joints distinct ; occiput
bluish grey, with bristles and stiff hairs. Thorax moderately dark
grey, with a slight yellowish tinge ; two dark median well-separated
narrow stripes from anterior margin to well beyond middle ; an
outer stripe on each side, abbreviated, enlarged iuto an irregularly
shaped and rather indefinite elongated large spot behind the
suture ; dorsum covered with short black hairs, except on the
stripes ; pleura3 concolorous, with black hairs ; a bunch of bristly
hairs on each humerus ; two macrochaetae just in front of wing-
base, and a fringe of long stiff black hairs on each side of the
metanotum ; scutellum concolorous, with four marginal bristles.
Abdomen moderately dark ash-grey ; hind margins of segments
shining black, the colour extending forwards in centre nearly or
quite to anterior margins ; 5th segment wholly grey ; 6th and 7th
grey, shining black basally ; venter grey ; ovipositor black. Legs :
coxae grey, tips brownish yellow ; femora in no way incrassatedy
brownish orange, a brown streak on upper side of fore pair, and
traces of such on middle pair ; tibiae and tarsi brownish yellow,
tips of joints of latter black. Femora with soft black pubescence,
longest on underside of posterior pairs ; hind pair with a row of
eight or more bristles ; tibiae with soft black pubescence and
longitudinal rows of bristles ; tarsi pubescent. Wings very
pale yellowish grey, unmarked ; stigma absent ; anterior cross-vein
just before middle of discal cell; halteres brownish yellow.
Length, 5 mm. (to tip of ovipositor).
Described from a single $ in the Indian Museum, taken between
Kugri and Phagu, Simla Distr., 21. v. 1906 (Annandale and Kemp).
Very like the European femorata, F., which might quite likely
occur in the Western Himalayas, but differing radically in t be-
hind femora showing no trace of incrassatiou.
292. Empis elegans, Brun.
Empis eleyans, Brunetti, Rec. Ind. Mus. ix, p. 26 (1913).
5. Head black; antennae black ; palpi large, yellow ; proboscis
twice the height of the head, bJack or blackish brown. Thorax
EMPIS. — HILARA. 357
shining black, including sides, scutellum, and metariotum ; wholly
unmarked ; a little soft whitish hair ; distinct bristles apparently
absent ; a few stiff pale hairs towards hind margin and on scu-
tellum. Abdomen darker brown, nearly bare ; genitalia slender,
normal, elongate, concolorous. Legs brownish yellow; hind coxaa,
tips of tibia?, and all the tarsi blackish. Femora quite bare ; tibise
with only a few short stiff hairs towards tips ; tarsi pubescent.
Wings clear, highly iridescent; anterior cross-vein just before
middle of discal cell ; stigma barely perceptible ; halteres brownish
yellow.
Length, 3 mm.
Described from two £ § in the Indian Museum from Mundali,
12. v. 1910, taken in company with E. griseonigra
293. Empis incouspicua, Brun.
Empis inconspicua, Brunetti, Rec. Ind. Mus. ix, p. 28 (1913).
c? . Head blackish ; antennae black ; palpi a little yellowish ;
proboscis twice the height of the head, yellow above, black below.
Thorax of an intermediate shade between light and dark grey,
with comparatively long and rather stiff hairs ; four long stiff
hairs on scutellum, which, with sides of thorax and metanotum,
are concolorous, but lighter grey just below shoulders. Abdomen
concolorous, rather broad and flat, with parallel sides and soft
black pubescence. Genitalia blackish, with reddish-brown bisinuate
terminal hooks; venter blackish. Leys uniformly dark brown,
pubescent ; middle tibiae with four long stiff hairs on outer side ;
hind tibiae with long hairs on hinder side. Wings pale grey;
anterior cross-vein a little before | of discal cell ; stigma brownish,
ill-defined, much elongated, and reaching the costal margin;
halteres yellow.
Length, 2| mm.
Described from two 3 in the Indian Museum from Lucknow,
17. i. 1908 (type) and 9. ii. 1908.
Genus HILARA, Mg.
Hilara, Meigen, Syst. Besch. iii, p. 1 (1822).
GENOTYPE, Empis maura (Europe) ; by Curtis's designation.
Allied to both Empis and llhamphomyia.
Head in some species nearly as broad as the thorax, shorter than
high. Eyes nearly always separated by a narrow irons in both
sexes, but in some species contiguous in the male, in which case
the upper facets are larger than the lower ones. Antennae
3-jointed, with distinct 2-joiuted style ; 1st and 2nd joints short,
about as long as broad, bristly ; 3rd elongate-conical, bare, slightly
compressed ; 1st joint of style very short, 2nd elongate, cylin-
drical. Proboscis about as 'long as height of head or a little
shorter ; palpi one-jointed, slightly curving upwards. Tliorar
rectangular, moderately arched. There are normally two strong
straight bristles on upper side of prothorax ; disc of thorax with
weak (lorso-central and acrosticlial hristles ; margin of dorsum
generally with a humeral, post-humeral, one to three notopleural,
one supra-alar, and a postalar bristle ; scutellum generally with
four bristles ; metapleura bare. Abdomen slender, of 8 segments,
cylindrical in male, with complex and rather uniform genitalia ;
broader and more pointed apically in female. Legs of normal
length and form ; fore metatarsus in the male nearly always more
or less thickened ; in the female the hind tibiae not infrequently
slightly compressed and curved ; occasionally the hind femora in
both sexes, or in male only, obviously thickened and suddenly
narrowed at apex. Legs moderately pubescent, characteristic
bristles often present ; tibia3 generally with apical spurs. Wings
as in Empis, except that the auxiliary vein distinctly reaches wing-
margin ; fork of 3rd vein perhaps less widely open ; anal vein
weaker, generally failing to attain wing-margin ; axillary lobe
well developed.
Range. World-wide.
Life-history. Beling has described the early stages of several
European species. They appear to resemble the ordinary
Dipterous larvae and pupae, the former being amphipneustic, of
twelve segments, including the head ; living in clamp earth or
amongst decaying leaves. The larva hibernates as in Empis and
Rliampliomyia. The perfect insects are general! v distributed,
more commonly occurring in woods and fields arid near water.
They prey on small Diptera or other soft insects, and have often
been captured in copula still carrying their prey, which is enveloped
in a little web, said to be spun from the mouth of the male.
Only four Indian species are described. Walker's bares is
unknown to me and may possibly be identical with my compacta.
Table of Species.
Thorax blackish or greyish.
Body brown bares, Walk., p. 359.
Body blackish ; fore metatarsi relatively
smaller compacta, Brim., p. 358.
Body blackish grey ; fore metatarsi re-
latively larger peshawarensis, sp. n., p. 360.
Thorax bright light yellowish brown nifithorax, Brun., p. 359.
294. Hilara compacta, Brun.
Hilara compacta, Brunetti, Rec. Ind. Mus. ix, p. 30 (1913).
c? . Head blackish ; frons rather narrow ; antennae with greyish
reflections viewed from certain angles. Proboscis about as long
as height of head, shining black above; lahium, which is shining
yellowish, much longer than the hypopharynx and sharply
pointed ; palpi large, nearly as long as proboscis, brownish,
JIILABA.
359
yellow, pubescent. Thorax wholly dark grey, including sides
scutellum, and metanotum ; a little black pubescence, and some
stronger hairs on dorsum laterally. Abdomen blackish, pubescent,
sometimes pale below at base ; genitalia a little broader than the
abdomen. Legs pale yellowish brown, pubescent; tibia with a
row of longer stiff hairs on outer side ; front metatarsi much
Fig. 32. — Hilara compacta, Brim., wing.
thickened, as wide as the tibiae, and longer than the remaining
fore tarsal joints ; posterior metatarsi normal, not so long as the
remaining tarsal joints. Wings pale grey; stigma elongate, pale
brownish, ill-defined ; halteres dirty brownish yellow.
Length, barely 2£ mm.
Described from three (J d1 in the Indian Museum from Simla,
16. v. 1909 (type) and 9. v. 1909 (Annandale).
295. Hilara bares, Walk.
Hilara bares, Walker, List Dipt. Brit. Mus. iii, p. 491 (1849).
" Body brown, clothed with short black hairs ; eyes dark red ;
feelers and mouth black ; breast hoary ; legs tawny, clothed with
short black hairs ; feet brown : wings light brown, darker for
one-third of the length along the fore-border ; wing-ribs and veins
piceous; poisers brown. Length of the body, ]| line; of the
wings, 3 lines.
" East India. Presented by Capt. Parry."
Type in the British Museum.
296. Hilara rufithorax, Brun.
Empimorpha rufithorax; Brunetti, Ilec. Ind. Mus. ix, p. 30 (1913).
S . Head : occiput, f rons, which is moderately wide with
parallel linear edges, and face all black, with a few black bristles,
the latter with a pair of long bristly hairs; vertex with three
distinct large reddish ocelli, with a pair of strong divaricate
bristles between the upper ones. Antenna? yellow ; 3rd joint
elongate onion-shaped, with a long closely pubescent black arista,
ending in a small bare black pointed bristle ; the iirst two antennal
joints with some short bristles near their tips. Proboscis about
as long as the height of the head, the basal half yellowish, the
apical half blackish ; palpi very long, longer than proboscis, some-
what broadened and flattened at the tips, with a few yellowish
hairs. Thorax bright light yellowish brown, with three black
dorsal stripes, of which the median one is of moderate width, the
360 EMPTDjE.
outer ones are broad enough to extend over the lateral margins,
and all three attain the anterior margin of the dorsum ; a few
bristly hairs about the roots of the wings and much smaller ones
on the dorsal stripes ; sides a little more yellowish than the
dorsum, with which latter the scutellum and metanotum are con-
colorous, the scutellum bearing a few stiff black hairs on the hind
margin ; the metanotum a little dusky in the centre. Abdomen
dark blackish brown, with black pubescence ; genital organs dark
grey, with a little pubescence ; venter similar to dorsal side of
abdomen. Legs: coxae and femora yellowish ; tibiae dark brownish
yellow, tarsi dark brown ; front metatarsi considerably widened ;
all the le»s shortly pubescent ; tibiae with txvo rows of five or six
long stiff hairs. Wings pale brown ; stigma vague, very elongate,
a little darker brown ; halteres yellowish brown.
Length i nearly 4 mm. .
Described from a single <5 in the Indian Museum taken by me
at Darjiling, 29. v. 1910, on the hillside.
297. Hilara peshawarensis, sp. nov.
c? ?. Whole insect blackish grey. Head: proboscis shining
black, as long as height of head, with stiff hairs ; antennae robust,
microscopically grey pubescent; palpi rather long, with moderately
long bristly hairs. Thorax with almost microscopic yellowish-grey
tomentum, which is more conspicuous on hinder part and on
scutellum, the latter with four long marginal bristles. Abdomen
with the segments more or less microscopically grey-dusted (in
good specimens) ; some grey pubescence with longer bristly hairs
on hind margins of segments ; genitalia knob-like, rather large,
concolorous, pubescent. Legs chestnut-brown, varying a little in
colour, pubescent ; fore metatarsus in $ verv large, oval, much
larger proportionately than in //. compacta ; pubescence on front
tibiae becoming longer towards tip ; knees yellowish ; hind tibiae
in $ considerably, though not conspicuously, thickened except at
base. Wings pale grey, iridescent, stigmatic region brownish ;
stigma darker brown, elongate, with ill-defined edges.
Length, 3 mm.
Described from a series of both sexes from Peshawar, N.W.
India, 16. hi. 1913 (Hoivlett).
Type rf and $ sent to British Museum, cotype c? £ in Indian
Museum and my collection.
Genus HOWLETTIA, Brun.
Hoiclettia, Erunetti, Rec. Ind. Mus. ix, p. 23 (1913).
GENOTYPE, Hoivlettia favipes, Brun.
A genus of anomalous position, allied to the EMPIRE, OCTDEO-
MIIN.E, and TACHYDHOMIIXJE (Platypalpus).
$ . Head with eyes separated above by a moderately broad
frons and only very uarrowlv separated below antennae. Antenna)
HOWLETTIA. 361
of three distinct joints; first two short, subequal; 3rd shortly
conical, with a long terminal arista. Proboscis about equal to
height of head, perpendicular ; palpi halt' as long as proboscis or
rather longer, quite broad. Thorax normally arched. Abdomen
rather longer than thorax, somewhat compressed. Legs rather
long and slender ; fore coxae slightly longer than posterior pairs ;
middle femora somewhat incrassated ; all tarsi longer than tibiae.
Wings with auxiliary vein faint and incomplete; 1st longitudinal
vein rather further from costa than usual, ending beyond middle
of wing ; 2nd vein beginning at one-fourth of wing, ending
towards tip of costa ; 3rd vein beginning at middle of wing ;
anterior cross-vein just beyond origin of 3rd vein, which latter
ends at about wing-tip ; 4th vein nearly straight, ending just
below wing-tip ; 5th nearly straight ; 6th (anal) very long, com-
plete to wing- border. Posterior cross-vein meeting 4th vein at
spot where anterior cross-vein meets it, but not in the same
straight line ; anal cross-vein nearly perpendicular to anal vein ;
2nd basal cell slightly longer than 1st, apex pointed; anal cell
barely half as long as 2nd basal ; discal cell absent,
c? unknown.
Range. Only the type-species known.
The genus is named after Mr. F. M. Howlett, of the Agricul-
tural Research Institute at Pusa.
298. Howlettia flavipes, Brim.
Hoivlettiajlavipes, Brunetti, Rec. Ind. Mtis. ix, p. 23 (1913).
$ . Head blackish ; proboscis dark brown ; palpi brownish
yellow ; ocellar triangle rather prominent. Ihorax shining
black, with very short whitish pubescence towards margins of
dorsuin ; sides of thorax and dorsum of scutellum slightly grey-
dusted ; a few weak prealar bristly hairs. Abdomen black, a little
shining; venter brownish yellow, with a little very short whitish
pubescence. Legs brownish yellow, minutely pubescent, middle
femora a little incrassated, with a row of very short bristles on
underside ; tarsi blackish. Wings clear, iridescent ; venation in
accordance with generic description ; halteres dull brownish
yellow.
Length, 2| mrn.
Described from a single $ in the Pusa collection, now sent to
the British Museum, from Mussoorie, ix. 1906 (Hawlett).
This genus is left in the subfamily EMPIN^ for the present, but
it is distinctly anomalous, and does not exactly fit into any sub-
family. The distinctly though only moderately incrassated middle
femora suggest the TACHYDROMIIN.E, from which the very distinct
anal cell and complete anal vein combined separate it. The
absence of the discal cell makes it abnormal in EM PIN-*:. It has
affinities with the OCYDROMIIX^: in the less distinct anal angle of
the wing, though it is by no means cuneiform, also in the anal
cross-vein being perpendicular to the anal vein.
302
Subfamily OCYDROMIIN^.
Head : eyes in male contiguous, in female contiguous or not,
rarely narrowly separated in both sexes. Proboscis generally
shorter than head, if long, projecting directly or obliquely for-
wards ; palpi one-jointed. Antennae 3-jointed, with terminal
arista or 2-joiuted style, which latter may be aristiform.
Thorax more or less humped; metapleura bare or with short
hairs. Wings with axillary lobe well developed (except in
Leptopeza) ; 3rd longitudinal vein simple ; 4th simple or forked.
Discal cell present ; three or four posterior cells ; basal cells sub-
equal, anal cell not longer than 2nd basal. Lower branch of 5th
longitudinal vein meeting anal vein parallel with, perpendicular
to, or at an angle of 45° to hind margin of wing, making anal cell
truncate at tip or the anterior margin of it the longer ; anal vein
of varying length ; squamae very small or absent. »
This subfamily comprises smaller and more delicate species,
often of pale colour, of which the short proboscis is a principal
feature.
Genus LEPTOPEZA,
Lemtopcza, Macquart, Soc. Sci. Lille, p. 143 (1827) (emend.}.
GENOTYPE, Ocydromiaflavipest Mg. (Europe).
Head almost as broad a.s thorax ; eyes occupying whole front
of head, contiguous in both sexes, rarely otherwise ; * facets of
uniform size. Antennae 3-jointed ; first two short, bristly ; 3rd,
though narrowed slightly at base, conical, bearing a long rather
thickened bare arista joined to tip of 3rd by a minute joint.
Epistome very narrow, -the eyes reaching the mouth-opening ;
proboscis Aery short. Thorax rectangular, strongly arched, with
minute discal bristles ; on the margin generally a notopleural,
postalar, and sometimes a supra-alar bristle, with other small
bristles or bristly hairs; scutellum with two or more marginal
bristles or stiff hairs ; metapleura with short fine hairs. Abdomen
slender, 8th segment small or hidden. Genitalia in male with
asymmetrical parts ;, in female forming a long, compressed, sword-
shaped ovipositor. Leys rather long and slender, hind pair
longest ; hind tibia? sometimes slightly thickened towards apex ;
sol't pubescence and various bristles on the legs. Wings with
auxiliary vein abbreviated, not reaching margin ; 3rd vein simple ;
anterior fork of upper branch of -4th vein indistinct or absent;
lower branch of eth vein strongly recurrent, parallel with hind
margin of wing ; anal vein shortened.
Range. Europe, India, ]STorth Africa, Cape Horn, North
America, Australia.
Life-history unknown.
* A narrow frons above in at least one European species (sph-enopiera), yet
the eyes contiguous below antenna-.
LEPTOPEZA. 363
299. Leptopeza'vitripennis, Bnm. (PI. IV, figs. 12, 13.)
Leptopeza vitripennis, Brunetti, Eec. Ind. Mus. ix, p. 31 (1913).
c? $ . Head blackish ; proboscis horizontal, as long as length of
bead ; shining black ; palpi short, porrect, black ; antennae with
elongate conical 3rd joint, arista long, black. Thorax very highly
arched, shining black in middle of dorsum, with short whftish
hairs laterally. Abdomen shining black, with short whitish
pubescence; venter similar. Legs rather pale pitch-brown, a
little pubescent. Wings very clear and iridescent ; halteres black.
Length, 2 mm.
Described from a type d1 from Oncha Gaon, Naina Tal Distr.,
1-6. v. 1907, and a type $ and a second $ from Eaugoon, 24. ii.
1908 (Annandale).
Types in the Indian Museum.
Subfamily IIEMERODROMIIN^E.
Head with eyes separated in both sexes, frons sometimes nar-
rower in d1 ; eyes sometimes contiguous below; when this is so,
frontal facets larger than the others ; eyes sometimes pubescent.
Proboscis short, not longer than height of head ; palpi 1- or
(indistinctly) 2-jointed. Antenna? 3-jointed, with, in some genera,
a 2-joiuted style which may be aristiform ; one or two excep-
tions occur. Thorax longer than broad, but slightly arched ;
metapleura bare, with short hairs or a few bristles. 'Leys with
fore coxae always lengthened, sometimes about as long as femora,
always longer than posterior coxae. Fore femora often incrassate
and fore legs often raptorial. Wings cuneiform at base, the
axillary lobe being greatly reduced or absent ; 3rd longitudinal
vein simple or forked ; 4th vein forked ; lower branch of 5th vein
may be distinctly recurrent, or placed perpendicularly to hind
margin of wing, or absent. Four posterior cells ; discal cell
present or absent ; basal cells more or less subequal ; anal cell
longer or shorter than 2nd basal cell, or absent; anal vein of
varying length or absent ; squarme absent.
The cuneiform wings and much elongated fore coxae will easily
distinguish members of this subfamily. Many of the yellowish
forms of EMPLOY belong to it.
Table of Genera.
1. Third longitudinal vein forked ........ 2. Tp- 367.
Third longitudinal vein simple ........ PHYLT.ODJIOMIA, /«'tt..
2. A cross-vein between 2nd and 3rd veins. . Doi.iCHOCEi'HALA.Macq.,
No such cross-vein, .................. 3. I p. 3/ 0.
3. Discal cell absent .................... MICKODIIOMIA, Big.,
Discal cell present .................... 4. [p. 366.
4. Proboscis thick, shorter than height of
hea(l ............................ CMNOCEHA, Mg., p. 364.
Proboscis thin, about ns long as height [p. 371.
of head . ................... TRICHOPKZA, Rond.,
364
Genus CLINOCERA, Mg.
Clinocera, Meigen, lllig. Mag. ii, p. 271 (1803).
Bydmdromia, Macquart, Hist. Nat. Dipt, ii, p. 658 (1835).
Paramesia, Macquart (nee Steph. 1829), loc. tit. p. 656 (1835).
GENOTYPE, Clinocera nigra, Mg.
Head usually broader than long, nearly or quite as long as
thorax ; epistome more or less elongated downwards ; eyes well
separated in both sexes, shortly and densely pubescent, facets of
uniform size. Proboscis very short and thick; palpi 1-jointed
or indistinctly 2-jointed, short, slightly clavate at tip. Antennae
inserted above middle of head, 3-jointecl ; 1st joint cup-shaped,
2nd subglobular, bristly, both short ; 3rd onion-shaped, about as
long as first two together, with a thick styliform pubescent arista
articulated to tip of 3rd joint by a microscopic joint. Thorat a
little longer than broad, rectangular, moderately arched. A
humeral (occasionally more than one), a post-humeral, one or two
notopleural, a supra-alar, and postalar bristles present ; scutelltim
with two long apical bristles, rarely more ; metapleura with some
hairs. Abdomen elongate, slender, the complex male genitalia
concealing the reduced 7th and 8th segments ; in female abdomen
more pointed. Leys long and slender, fore coxae not greatly
elongated ; front femora often thickened at base and with bristles
below ; legs shortly pubescent. Wings somewhat lengthened ;
auxiliary vein rather short but reaching margin ; 1st longitudinal
vein ending at about middle of wing ; 2nd vein sometimes wavy —
in some species with a sudden loop downwards enclosing the
stigma; 3rd vein forked; discal cell present, varying in length,
discal end pointed or truncate ; lower branch of 5th vein strongly
recurrent, parallel with hind margin of wing ; anal cell with distal
end bluntly pointed or rounded ; anal vein abbreviated. Alula
absent ; alar squamula very small.
llanye. Europe, Siberia, India, North America.
Life-history unknown ; probably aquatic in larval state.
This genus has a facies distinguishing it easily from other
EMPIDJE, and with some resemblance to some water-haunting
DoLicnopomD^. The elongate body and proportionately longer
legs, the pubescent eyes and rather long wings, the usually
greyish colour with lighter parts, and the not infrequently spotted
wings, give them a characteristic appearance. All are found in
the neighbourhood of water, running streams, or stagnant pools,
on the surface of which most species appear capable of running
with ease.
The European species of Clinocera may be fairly satisfactorily
grouped into several subgeuera, mostly set up by Mik; but as
only two species are yet known from the Orient, of which at least
one (fluviatilis) comes in Clinocera, s. sir., these subdivisions may
be ignored here.
CLINOCERA. 365
The two Indian species differ as follows : —
Brownish-grey species, with dark brownish-grey legs ;
length, 3 mm flmiatilis, Brtm.
Blackish-grey thorax, dark brown abdomen, brownish-
yellow legs ; length, 2 mm obscura, Brun.
800. Clinocera fluviatilis, Brun.
Clinocera fluviatilis, Brunetti, Rec. Ind. Mus. ix, p. 34 (1913).
c? . Head brownish grey, with isolated black bristles on the
occiput and a few on the frons; frontal tubercle bearing two
divergent bristles. Antennae black ; 1st and 2nd joints subequal ;
3rd onion-shaped, with moderately long, nearly bare arista. Pro-
boscis and palpi blackish, both short. Thorax very dark brownish
grey, nearly black, a whitish-grey stripe on the sides from just
below the shoulder to as far as below the ends of the transverse
Fig. 33. — Clinocera fluviatilis, Brun., and antenna.
suture ; ventral part of thorax also whitish grey. Dorsum bearing
several isolated bristles (many of these are broken off), of which
one is present on the shoulder, and one towards, but not at the
hinder corner ; scutellum with two long bristles. Abdomen dark
grey, normally pubescent. Genitalia large, consisting of a ventral
dark grey plate, a large concave lower piece, and a pair of claspers
above, these latter constituted of a large oval basal concolorous
joint and an elongate brownish-yellow horny bifid second joint :
the whole of the genitalia a little pubescent. Legs long and
slender, uniformly dark brownish grey, practically bare. Wings
very pale grey. The 3rd longitudinal vein forked beyond three-
fourths of its length, the branches approximate to one another
and parallel ; anterior cross-vein before middle of discal cell, and
at about one-third the length of the wing; branches of upper
branch of 4th longitudinal vein issuing close together from discal
cell, thence gradually diverging ; the discal cell much longer on
its anterior border than its posterior one; lower branch of 4th
vein nearly straight, rather more strongly developed ; 2nd basal
cell short, lying against the middle third of the upper basal cell ;
366
anal cell barely longer than 2nd basal; 6th vein very short.
Halteres : stein yellow, clubs dark grey.
Length, 3 ram. ; wing, 4 mm.
Described from a single specimen from Bhowali, Kumaon Distr.,
12. vi. 1910 (Imms), on stones on banks of stream.
A specimen which probably belongs to this species, but showing
a discrepancy in the shape of the discal cell, is amongst the
Diptera collected by Mr. Kemp on the Abor Expedition.
301. Clinocera obscura, Bnm.
Clinocera obscura, Brunetti, llec. Ind. Mus. ix, p. 34 (1913).
cJ . Head blackish grey, some stiff hairs on occiput and frons,
which latter is very broad. Proboscis short, half the height of
the head, shining black, with a much swollen apical part (this
latter in one specimen partly brownish yellow) ; palpi very small,
brownish yellow. Antenna! 3rd joint elongate oval, with long
black arista. Thorax brownish grey, including scutellum ; sides
and metauotum bluish ash-grey ; some dorso-central and lateral
bristly hairs. Abdomen dark brown, softly pubescent. Genitalia
extremely long and conspicuous, blackish, with a few pale hairs,
egg-shaped, convex, bent under the venter, the narrower end
pointing towards the base of the abdomen ; posteriorly (i. e. from
the broader end) project some yellowish filamentous appendages.
Legs brownish yellow ; coxae ash-grey, especially the posterior
pairs ; tarsi a little darker. Fore femora with a few soft pale
longer hairs; remainder of legs microscopically pubescent. Wings
very pale grey, wholly unmarked, highly iridescent; halteres
brownish yellow.
Length, 2 mm.
Described from two rf c? in the Indian Museum from Simla,
10. v. 1909 (Annandale),
Genus MICRODROMIA, Big.
Mia-odromya, Bigot, Ann. Soc. Ent. France, (3) v, p. 503 (1857)
(emend.).
GENOITPE, Tachydromia oratorio, Fin. (Europe).
This genus differs from Clinocera by the fore coxse being nearly
as long as the femora, the latter being considerably thickened.
Head elongate, ovate ; eyes more elongate than in Clinocera and
sometimes contiguous below the antennae ; arista un jointed ;
proboscis about as long as height of head. Thorax with the major
bristles apparently variable ; metapleura bare. Legs with fore
coxae from rather shorter than to nearly as long as femora ;
fore femora much thickened, with two rows of short spines
below, and a row of long bristles on each side ; fore tibia? curved
at base, with a row of short spines on inner side and a long apical
spine ; posterior tibiae without apical spur. Winys with subcostal
vein short; no discal or anal cell ; 3rd vein forked; lower branch
MICEODROMIA. — PHYLLODUOMIA. 367
of 5tli longitudinal vein absent or abortive; anal vein generally
absent ; no stigma.
Range. Europe, Siberia, India, Java, North America, Chile.
Life-history. The metamorphoses are known of pnec'itoria,
Eln., a European species of Jfemerodromia, of which Microdromia
is generally regarded as a subgenus, and Brocher has described
and figured them.* The larva lives in mud in streamlets, the
pupa being also aquatic.
The perfect insects frequent the borders of woods and lakes, or
humid places, generally on low herbage and in grass.
302. Microdromia dorsalis, Brun.
Chelipoda dorsalis, Brunetti, Rec. Ind. Mus. ix, p. 33 (1913).
c? $ . Head blackish ; proboscis pale yellowish, very stout at
the base, the upper piece prolonged into a long hard shining
rostrum ; antennas wholly pale yellowish white. Thorax brownish
yellow ; dorsuin black or dark bro\vn, the colour extending more
or less over the sides ; scutellum and metanotum concolorous with
dorsum. Abdomen brownish yellow ; the dorsum of each segment
black, thus forming a continuous black stripe from base to tip,
but the hind borders of some of the segments escape the black
colour ; venter brownish yellow. Geniralia in /"<
melanocepliala, F., as type of Phyllodromia, the same species which Coquillett
chose as type of Chelipoda in 1910.
368 EMPID.E.
antennae, with facets of uniform size ; proboscis short, thick,
vertical ; palpi one-jointed. Antennae inserted a little below the
middle, 3-jointed ; basal joints short; 3rd elongate, pointed, with
long terminal arista, attached by means of a minute joint. Thorax
rectangular, moderately arched ; prothorax forming a short neck.
A few uniserial dorso-central bristles ; a notopleural and supra-
alar bristle present ; scutellum with two bristles ; metapleura
with a few bristles. Abdomen with last two segments in male
reduced, hidden by the genitalia, which latter are rather large ;
abdomen in female more pointed, genitalia normal. Legs as in
Clielipoda • fore coxae nearly as long as fore femora. Wings in
venation and shape as in Chelipoda; discal cell present; anal
angle nob developed.
Range. Europe, India, N. America, Argentina.
Life-history unknown. The images frequent grass and damp
meadows.
Phyllodromia, Zstt., differs from Chelipoda, Macq., practically
only in the presence of the discal cell, and is ranked as a sub-
genus of the latter in the Kertes/ Catalogue ; but if the character
is constant one way or the other in all tlie species, there seems no
reason why both should not be admitted to generic rank. My
flavida was placed in Chelipoda on the broader view of the genus,
but must now be transferred to Phyllodromia, with my indica,
the latter having been described under Litanomyia, which is now
considered synonymous with Zetterstedt's genus.
P. flavida differs from indica by the thorax being mainly
brownish yellow as opposed to blackish grey.
303. Phyllodromia flavida, Bnm. (PI. IV, figs. 14, 15.)
Chelipoda flavida, Brunetti, Rec. Ind. Mus. ix, p. 32 (1913).
d1 $ . Head yellowish ; three ocelli on the black vertical triangle,
which also carries two divaricate long bristles : frons moderately
broad; occiput with four long curved bristles placed equidis-
tantly ; seen from behind, the occiput has a greyish tinge, as has
sometimes the frons also; eyes black, rounded, almost contiguous
immediately below the antennae for a short space, the face
broadening below ; the latter, seen from above, appears silvery
white ; proboscis very short, half the height of the head, conical,
sharply pointed, yellowish, the upper part (hypopharyux) narrow,
hard, shining. .First two antennal joints yellowish, with some
short bristles at the tips of each ; 3rd joint dark br.mvn, elongate
onion-shaped, with long brown microscopically pubescent arista.
Thorax very variable, generally brownish yellow, with more or less
blackish colour, either in the form of a dorsal stripe, a lateral ir-
regular and somewhat indistinct border, or the posterior part of the
dorsum more or less blackish; sides generally lighter, in some speci-
mens attaining a pinkish-fawn colour. Abdomen variable, brownish
yellow, more or less blackish in part or wholly ; venter brownish
PHYLLODROMIA. 369
yellow ; genitalia small, concolorous. Tip of abdomen in c?
stump-like; in ? tapering, the two terminal lamella? pale
yellowish. Legs yellowish or pale yellow ; front coxae narrow,
cylindrical, nearly as long as the femora, the latter slightly
brownish at tips and considerably iucrassated, with two yellow
Fig. 34. — Phyllodromia flavida, Brun.
bristles near the base on the underside and a row there also of
microscopic spines ; posterior coxae and femora of normal size,
the former a little brownish, as are also the tips of the tarsi.
Wings very pale grey, veins black ; halteres brownish yellow.
Length, 2 mm.
Described from several specimens in the Indian Museum from
Paresnath, 4000-4300 ft., Chota Nagpur, 9-13. iv. 1909 (Annan-
dale), including type 3 (9. iv. 1909) ; Dawna Hills, 2000-3000 ft.,
2.iii. 1908, type $ (Annandale) ; Darjiling, 28. v. 1909 (Brunctti).
The Pai'esnath specimens were taken on stones, where they
were in considerable numbers hidden in undergrowth at the edge
of a spring.
304. Phyllodromia indica, Brun. (PI. IV, figs. 16-18.)
Litanomyia indica, Brunetti, Rec. Ind. Mus. ix, p. 36 (1913).
ydro-
122.
dontia), 167.
mi a), 376.
fulvipes (Bombylius),
Drapetis, 380.
duvaucdii (Anthrax),
ferrugincus ( Platypalpus),
376.
271.
fulvithorax (Elaphro-
225.
ferruginosns (Chryso-
peza), 385.
duvaucelii(Argyran.iceba),
225.
piliis), 141.
ferrugiiwsus (Leptis}, 141.
fusca (Trichopeza), 372.
fuacipenuis (Acarterus),
Jimbriatus (Anthrax), 240.
335.
llainmea (Exoprosopa),
fuscirjennis (Synechos),
ed wards! (Coonomyiodes),
193.
323.
111.
flaveol us (Macellopalpus),
f uscitnrsis (Boris), 93.
edwardsi (Systropus),
294.
122.
flavida (Chelipoda), 3(58.
fuscolimbata (Anlhrax),
246.
Elaphropeza, 384.
flavida (Eudnieta), 43.
fuscus (Oncodes), 171.
398
ALPHABETICAL INDEX.
Gabaza, 24.
Hyalanthrax, 234.
insulator (Exoprosopa),
gagatina (Hybos), 339.
'gagatinus (Hybos), 339.
gemma (Microchrysa), 88.
"gemmifer (Sargus), 81.
geniata (Ampsalis), 44.
genieulata (Beris), 91.
gcniculata (Hybos), 340.
geniculatus (Hybos), 340.
gentilis (Argyramoeba),
230.
hyalina (Anthrax), 254.
hyaliventris (Actina), 93.
Hybos, 336.
HYBOTXN.E, 325.
Jiybridus (Bombylius),
279.
Hydrodromia, 364.
hydroleon (Musca), 61.
Hyperalonia, 178.
Hypoleon, 53.
203.
inter media (Atherix), 116.
intermedia (Atrichops),
116.
intermedia (Argyra-
moeba), 221.
invaria (Thereva), 302.
iridipennis (Parahybos),
343.
irrorata (Tachvdromia),
qentilis (Platypalpus),
377.
hypoleon (Musca), 53.
Hypselura, 311.
370.
Isopenthes, 234.
gentilis (Tachydromia),
377.
Hyrmoneura, 147.
Hyrmophlttba, 147.
Geron, 279.
gestroi (Argyramoeba),
ibis (Atherix), 120.
javana (Atriadops), 155.
javana (Beris), 30._
javana (Colax), 155.
gibbosa (Musca), 168.
(/laucesfens (Platypeza),
392.
Ibisia, 120.
illuceus (Musca), 39.
illucens (Suragina), 121.
javana (Tinda), 30.
javana (Toxophora), 287.
gracilis (Phthiria), 285.
imberbis (Bibio), 302.
gravelyi (Chrysopilus),
immaculata ( Odonto-
kashmirensis (Odon-
137.
myia), 67.
tomyia), 67.
griseonigra (Empis), 353.
ariseonigra (Blmmpho-
immaculatus (Syneches),
328.
kashmirensis (Ptero-
dontia), 166.
myia), 346.
inasqualis (Svneches),
keinpi (Thereva), 298.
grossipes (Musca), 336.
330.
guttatipeunis (Anthrax),
237.
incisa (Tachypeza), 379.
inclsiis(Platypalpus), 379.
labiata (Atherix), 120.
inconspicua (Empis),
]seta (Hermetia), 40.
357.
la,nopyya (Atherix), 115.
Harpamerus, 334.
inclianus (Leptomydas),
lanopyga (Atrichops),
Heliomyia, 128.
320.
115.
HEMERODKOMIIN.E, 363.
indica (Biast.es), 29.
lar (Anthrax), 194.
hetniuopla (Clitellaria),
indica (Cyphoinyia), 57.
lar (Bibio), 194.
50.
indica (Evaza), 33.
lar (Exoprosopa), 194.
Hemipenthes, 234.
indica (Lita.nomyia), 369.
Lasia, 163.
Henops, 168.
iniica (Oxycera), 66.
Lasiopa, 52.
Hermetia, 39.
indica (Phyllodroinia),
lateralis (Exoprosopa),
Hermoheura, 147.
389.
198.
Heteralonia, 189.
indica (Psilocephala),
latifascipennis (Tachy-
Heteroxycera, 53.
304.
dromia), 375.
Hexacantha, 90.
indica (Thereva), 301.
latipennis (Exoprosopa),
Hilnra, 357.
indica ( Tinda), 30.
198.
himalayana (Eharapho-
indicus (Biastes), 30.
latus (Chrysopilus), 143.
myia), 346.
iudious (Empidideicus),
Lepidanthrax, 232.
himalayanus (Anthrax),
260.
LEl'TID.E, 100.
238.
Inei-mia, 52.
LEPTIN.E, 112.
Hinnoneura, 147.
inermis (Allognosta), 94.
Leptis, 124, 128.
Holoclera, 345.
Inermyia, 52.
Leptomydas, 320.
Hoplomyia, 58.
inflata (Euba), 49.
Leptopeza, 362.
hottentotta (Argyra-
infurcata (Pachy '(/aster),
Leptosceles, 370.
maeba), 228.
'21.
limbata (Atherix), 115.
hottentotta var clari-
insignis (Syneches), 322.
limbata (Atrichops), 115.
pennis (Anthrax), 247.
institnta (Anthrax), 227.
limbiventris ( Toxocera),
hottentottus (Anthrax),
instituta (Argyramoeba),
42.
248.
227.
liraitaris (Argyramoeba),
Howlettia, 360.
insularis (Chrysopilus),
220.
humeralis (Chrysopilus),
140.
limpida (Anthrax), 255.
135.
insulata (Anthrax), 203.
Litanomyia, 367.
ALPHABETICAL INDEX.
399
JMorhynchus, 189.
lomata (Pialea), 160.
LOMATIIN^E, 255.
Louehoptera, 388.
LONCHOPTEKID.E, 387.
Lonch&pteryx, 388.
longicornis (Brachy-
stonia), 371.
longicornis (Brevios), 377.
longipennis (Ptecticus),
75.
longipennis (Sargus), 75.
longirostris (Anastce-
clius), 278.
longispinus (Auipsalis),
44.
lucens (Anthrax), 254.
lucida (Anthrax), 254.
luctuosus (Chrysopilus),
133.
luridus (Sargus), 77.
lusitanicus (Mydas), 320.
lutntius (Odontoinyia),
70.
lutatius (Stratiomys), 70.
lutea (Lonchoptera), 389.
Macellopalpus, 122.
niacilenfcus (Systropus),
289.
Macroceromys, 96.
Macrostomus, 345.
mactans (Sargus), 83.
maculipenne (Ephip-
pium), 47.
maculipennis (Negrito-
myiii ), 47.
maculiventris (Exopro-
sopa), 200.
maculatus (Asilus), 287.
raaculatus (Bombylius),
267.
magnifica (Ceylonia),
156.
magnifaus (Sargus), 81.
magnipennis (Chrvso-
pilus), 138.
major (Bombylius), 261,
265.
manit'esta (Anthrax),
242.
marginata (Empis), 356.
marginata (Eudmeta), 42.
marginata(Hermetia), 42.
marqinata (Pacliymeria),
356.
marginata (Usia), 283.
marmoratus (Chryso-
pilus), 131.
matira (Anthrax), 238.
maura (Ernpis), 357.
niyeicauda (Argyra-
Megapalptu, 284.
moeba), 224.
Mesophysa, 163.
niveisquamis (Argyra-
inetallinus (Sargus), 82.
mo3ba), 229.
metatarsalis (Atherix),
niveiventris (Exopro-
117.
sopa), 207.
metatarsalis (Atrichops),
nubila (Tachydroinia),
117.
378.
metatarsata (Elaphro-
peza), 386.
obesa (Clitellaria), 26.
Meta.fon.ia, 93.
obscura (Clinocera), 366.
Microchrysa, 87,
Microdromia, 366.
obscura (Hirmoneura),
Microdromya, 366.
micropilosa (Stratio-
obscura. (Platypeza), 392.
obscurifrons (Argyra-
myia), 60.
mocba), 220.
minimus (Bombylius),
ochracea (Hirmoneura),
273.
151.
minuta (Odontomyia),
ochracea ( Odontmnyia),§5.
65.
Octacuntha,, 90.
minutus (Svneches), 330.
modifera(Tinda),SO.
octomaculatus (Oncodes),
170.
Monacanthomyia, 23.
OCYDKOMIIN^E, 3(52.
morio (Anthrax), 234.
Odontomyia, 61.
morio (Stomoxys), 178.
renoinaus (Hyperalonial,
MYDAID.E, 314.
185.
Mydas, 316.
Myiochrysa., 84.
cestracea (Trichopsidea),
153.
Myochrysa, 81.
Oncodes, 1(58.
mystaceum (Spogosty-
opaca (Hirmoneura), 150.
lum), 213.
opaca (Syndyas) 333.
opalescens (Chrysopilue),
134.
Negritomyia, 47.
ophioneus (Systropus),
NKMESTKINID.E, 144.
291.
Neopachygaster, 20.
Oplacantha, 90.
nepalensis (Tachy-
Opseogymnus, 61.
droinia), 377.
oratorio (Tachydroinia),
Nerna, 31.
366.
nigella (Thereva), 299.
orientalis (Bombylius),
niger (Hybos), 340.
263.
nigra (Clinocera), 364.
orientalis (Platypalpus),
niqricaudus ( Si/stropus),
378.
291.
orientalis (Plati/peza),
nigricornis(Rhachicerus),
392.
105.
orientalis (Tacbydromia),
nigricornis (Xylomyia),
378.
99.
ornatipennis (Chryso-
ni gripes (Phycus), 309.
pilus), 132.
Sigritomya, 47.
Oxycera, 53.
nigrofemorata (Argyra-
mceba), 230.
Pachygaster, 20.
nigrofemorata (Pachy-
PACHYGASTRIN.E, 19.
gaster), 25.
Pachymei-ia, 347.
nigronitidus (Hybos),
Pachymerina, 347.
342.
palliflitarsis (Syneches),
nitens (Hybos), 341.
331.
nitidulus (Bombylius),
pallid it ibi
8. ,, lanopyga, Brun., 3 ; p. 115.
9. ,, cincta, Brun., wing ; p. 118.
10. „ Uimbata, O.-S., wing; p. 115.
11. „ intermedia, Brun., lateral view of head;
12. „ „ wing. [p. 116.
13. ,, ccerulescens, Brun., wing; p. 119.
14. ,, metatarsalis, Brun., wing; p. 117.
15. Macellopalpus fulvidus, Brun., lateral view of head ;
[p. 122.
16. Leptis apicipennis, Brun., wing ; p. 125.
17. Clirysopilus marmoratus, Brun., wing; p. 131.
18- ,, albopictus, Brun., wing ; p. 131.
19. ,, ornatipennis, Brun., wing; p. 132.
20. „ unicolor, Brun., Aving ; p. 141.
21. „ stigma, Brun., wing; p. 142.
22. Hirmoneura opaca, Licht., wing ; p. 150.
23. Ceylonia magnified, Licht., wing ; p. 156.
24. Pialea auripila, Brun., ^ ; p. 161.
25. „ „ lateral view of head.
26. Pterodontia dorsolineata, Brun., $ ; p. 167.
27. „ „ antenna.
28. Oncodes octomaculatus, Brun., tf 5 p. 170.
29. „ arista.
PLATE II.
INDIAN LEPTID/E, NEMESTRINID/E AND CYRTID/E.
EXPLANATION OF PLATE III.
Fig. 1. Hyperalonia suffusipennis, Brun., wing ; p. 188.
2. ,, semifuscata, Brun., wing ; p. 188.
3. Exoprosopa lar, F., wing ; p. 194.
4. ,, insulata, Walk., wing ; p. 203.
5. „ bengalensis, Macq., wing ; p. 206.
6. „ brahma, Sch., wing ; p. 206.
7. ,, lateralis, Brun., wing ; p. 198.
8. ,, annandalei, Brun., wing ; p. 199.
9. „ puerula, Brun., wing ; p. 205.
10. Argyramoeba distigma, "Wied., wing; p. 218.
11. ,, obscurifrons, Brun., wing ; p. 220.
12. ,, gestroi, Brun., wing; p. 222.
13. ,, duvaucelii, Macq., wing; p. 225.
14. ,, argentiapicalis, Brun., wing ; p. 227
15. ,, gentilis, Brun., wing ; p. 230.
16. Toxophora javana, Wied. ; p. 287.
17. PsilocepTiala sequa, Walk., d ; p. 305.
18. ,, „ head in profile.
19. ,, ,, antenna.
20. „ „ abdomen, $.
21. Phycus britnneus, Wied. ; p. 307.
22. ,, ,, head in profile.
23. „ „ antenna.
PLATE III.
INDIAN BOMBYLIID/E AND THEREVID^E.
EXPLANATION OF PLATE IV.
Fig. 1. Leptomydas indianus, Big., wing ; p. 320.
2. Mydas annulata, Brun., wing ; p. 319.
3. Syneclies bigoti, Bezzi, wing ; p. 329.
4. „ palliditarsis, Brun., head in profile ; p. 331.
5. Acarterus fuscipennis, Brun., head in profile; p. 335.
6. ffybos yagatinus, Big., hind leg; p. 339.
7. „ tenures, Brun., antenna ; p. 338.
8. „ „ hind leg.
9. Empis rostrata, Brun., middle leg ; p. 355.
10. „ „ hind leg.
11. „ squamata, Brun., $, hind leg ; p. 352.
12. Leptopeza vitripennis, Brun., wing : p. 363.
13. ,, ,, antenna.
14. Phyllodromia flavida, Brun., wing j p. 368.
15. „ „ antenna.
16. ,, indica, Brun., head in profile ; p. 369.
17. „ ,, wing.
18. ,, ,, antenna.
19. Dolichocepliala 7-notata, Brun., head in profile; p. 371.
20. ,, ,, wing.
21. „ „ antenna.
22. Tacliydromia nepalensis, Brun., antenna ; p. 377.
23. Drapetis brevior, Brun., c? ; p. 382.
24. „ „ antenna.
25. „ rotund icornis, Brun., antenna; p. 383.
INDIAN MYDAID/E AND EMPID/E.
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